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Circular No. 278 


The Commercial Storage of 


Fruits, Vegetables, and 


Florists’ Stocks 


DEAN H. ROSE, Senior Physiologist e R. C. WRIGHT, Physiologist 
T. M. WHITEMAN, Associate Horticulturist 


Division of Fruit and Vegétable Crops and Diseases 


Bureau of Plant Industry, Soils, and Agricultural Engineering 


Agricultural Research Administration 


For sale by the Superintendent of Documents, Washington 25,D.C.- - - = 2+ = = = Price 20 cents 


UNITED STATES DEPARTMENT OF AGRICULTURE 


WASHINGTON, D. C., JULY 1949 


goers ATA 
: OTL ER ARTI 
i ® 
t 


Circular No. 278 
September 1933 * Washington, D.C. 


Revised November 1941 ° Slightly revised July 1949 
UNITED STATES DEPARTMENT OF AGRICULTURE 


The Commercial Storage of Fruits, 
Vegetables, and Florists’ Stocks 


By Dean H. Ross, senior physiologist, R. C. WRIGHT, physiologist, and T. M. 
WHITEMAN, associate horticulturist, Division of Fruit and Vegetable Crops and 
Diseases, Bureau of Plant Industry, Soils, and Agricultural Engineering, 
Agricultural Research Administration * 


CONTENTS 

Page | Fruits and nuts—Continued Page 
lnttroductiomee 2 8 eee 2 igs s(inesin) see So 2 ele. 21 
Factors involved in cold storage_ 3 Grapeiritit oe ee es 21 
Temperature of storage rooms 3 Grapesed tev ila eae Feeds fal is 22 
Humidity of storage rooms _ _- 5 JESINcK) OVS) Ym Le PRE Wie NRSC OR eer 23 
Evolution of heat by com- TSWNE Sime ioe tee ae oy es Oe 24 
MOOG Vee Meee soe ees. Y 6 Logan blackberries ____------ 24 
Effect of cold storage on subse- Olives. @iresh) 2VeRI es Horas ts 24 
quent behavior of fruits and Orangesse cts Wee Tes 24 
vegetablesaei eo ne ae se 12 JERSPY Gl aVeSpsMeel ay Ss 2 Dy ips Sn ona tl es 25 
Sea Glin Geers Me renee hee 13 REARS RSE Saks als ea i kein ea ey 25 
Wading Merais “CULE is Ti Pas 13 Pineapplese: 24's Ul tare Gal 
Choice of storage conditions__-__ 13| Plums (including prunes) _-__-- 27 
Bnuicseancenitsyye Stel. 2 ek Eee 14 Quince says See eee ee ee Te 28 
PATO DLE Si ae ets care ih le Ss a ee 14 VASP Derries : 420 ess Cais sete 28 
TeV OVS(COUS es Se  cl eeeaes IVA Strawberries ce eee o Leeman 28 
Wvocados:!£ Libis, Cult it 17 Dried irultses. 22 se eae om 28 
Bananas _____-------------- 18| Frozen fruits and vegetables_- 29 
Bldiekberriés fat. s.r v8). 19 Nut 29 

Cleric sits he spar Giese oe a 1 ee a Le ee 
Care ete ea Ton ceetablester ses) 2 === 22-2. 30 
Cranberricsue i. Ss rie 19 AS WATAg USS. jee” 22 Sanaa 31 
Date sw nts bee Ee eS DO Beans: Pls sae eet Si ee es 32 
Dewhermiegi 2 te) oe tee 2 ee 21 BOGGS eee ere ie Sir) ate 682 1 32 


*The writers not only have included results of their own investigations but 
also have drawn freely on published and unpublished results of other staff 
members, whose assistance and cooperation have done much to make possible this 
publication in its present form. Acknowledgment for unpublished information 
furnished on refrigeration or commodities indicated is due the following present 
or former members of this Division: W. R. Barger, dates and dried fruits; C. O. 
Bratley, limes and pineapples; Fisk Gerhardt, A. L. Ryall, and W. H. English, 
pears; C. W. Mann, lemons; W. T. Pentzer, refrigeration, garlic, olives, pears, 
and peppers ; Edwin Smith, refrigeration, apricots, and pears; Morris Lieberman, 
lima beans, cabbage, celery, and cucumbers; M. H. Haller, spinach; E. B. Lambert, 
mushrooms. Acknowledgment is aiso due J. N. Kelley, Fruit Dispatch Co., for 
information on bananas. Special acknowledgment is made to D. F. Fisher, of 
this Division, for valuable suggestions and criticisms throughout the preparation 
of this and earlier editions of the circular. 


829965—49——1 1 


2 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


Page Page 
Vegetables—Continued Vegetables—Continued 

Broccoli (Italian, or sprout- Pease (Green) S22 ee ee ener 41 
ING) = eee he eee 33 Peppers =: >a See 41 
Brusselsssprouts_- = 2s sees 33 Potatoes: 2. 2s 2 eat SF 41 
Galbuges ts0220 425.2" 4 See 33 Pumpkins and squashes-__---- 43 
Carrotseyye (er 22 Jos ee 34 Radishes (winter) _..._.------ 44 
Camlilgwerss 222 Ss =e errs 34 Rhubarbsss = 2 ees 44 
Celermac: 2458 2s eee 35 Rutabagas st... 2s See 5 44 
Gelerysc 25-3 Si eee ees 35'|: **Salsifiye se 5 ee fa eee eS 44 
Corn (eneem) hes ceases 36 Spinach: Ss aee ae. eee a4 
Cucumbers ent oe eer 36 Squashes acess ewe eee 44 
Hooplants es eee 37 Sweetpotatoes= == se 44 
Endive, or escarole_____-____- 37 Pomatoes ss. Se See ae 45 
Garlies(dry ia See 37 (PUTRI S ey a ye ee 45 
Horseradish #2 2 a By Frozen vegetables______.---- 46 

Jerusalem-artichokes.._ - -__-- 37 | Cut flowers, florists’ greens, rhi- 

Kohitahi == 38 zomes, tubers, corms, and 
Leeks (green) <2 2 = eee 38 bulbs 46 

Hhettuce= 2s Ge Se a ee 38 Cut fi Setgoeiesse 
MicloTish 352 ree ss eee ee 38 aes owers eR eed er 48 
Mushrooms (cultivated) wee 39| Florists’ greens______-------- 92 
Owes Si Sia ae 29 Rhizomes, tubers, and corms _ 53 
Onions and onion sets____-__- 391) Bulbssse SA eo ees 54 
Parsnips= sau eee eee se en 40; Literature cited === 222 —="* eae 54 

aR RS ARR 
INTRODUCTION 


The purpose of this circular is to present a series of brief sum- 
maries of the essential average storage requirements of most of the 
more important varieties of fresh fruits, vegetables, cut flowers, and 
certain other perishable commodities that enter the market on a 
commercial scale. Many details are of necessity omitted, as the work 
is intended primarily for general practical reference. The conditions 
given should not be considered as absolute or final, but rather as the 
safe limitations under which the various products can ordinarily be 
stored. Detailed information on the handling and storage of some 
of the commodities discussed is available elsewhere in the form of 
bulletins or textbooks; for many of them only general information 
exists. 

Fresh fruits, vegetables, cut flowers, etc., intended for storage 
should be as free as possible from skin breaks, bruises, and decay. 
They should be neither immature nor overmature, because in either 
case it may be difficult to keep them from too rapid deterioration, 
and even if they do not change appreciably in storage, they will not 
be commercially desirable when removed. The proper degree of 
maturity in each case can usually be determined by consulting the 
various sections of this circular, or the publications listed in Liter- 
ature Cited (p. 54), or on the basis of previous experience. There is 
a seasonal variation, however, in the storage quality of certain prod- 
ucts, particularly fruits; hence care must be taken not to assume too 
much from one year’s experience concerning the probable behavior of 
a given commodity grown the next year. 

Decay and other deterioration in storage is too broad a subject to 
be discussed in detail in this publication. It is discussed very gener- 
ally in connection with various fruits and vegetables covered by this 


. 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 3 


circular and in greater detail in other publications (8, 14, 22, 38, 44, 
58, 72, 77, 78, 79, 80, 82, 83, 84, 85, 86, 88, 95, 98, 109, 110, 112) ? 


FACTORS INVOLVED IN COLD STORAGE 


Recommendations for the best conditions for the storage of fresh 
fruits, vegetables, and cut flowers are subject to change from time to 
time as more definite information is gained in the handling of these 
commodities. The conditions and requirements given in this circular 
are derived from the best commercial practice at the present time 
and from scientific experimentation. The term “cold storage” as 
used in this circular refers to storage where temperatures in the ap- 
proximate range of 25° to 45° F. are maintained by mechanical means 
or by the use of ice and salt; “freezer storage” to temperatures rang- 
ing from about 15° to —20°, usually close to zero; “cool storage” to 
temperatures ranging from about 50° to 65°, or possibly 70°; “un- 
refrigerated storage” to temperatures that are allowed to fluctuate with 
outside temperature; and “common storage” to storage in which the 
temperature best suited to the commodity is obtained as nearly as 
possible by insulation and ventilation only. 

All of the temperature requirements are given in degrees Fahrenheit 
and represent the average air temperatures that should be maintained. 
The humidities are relative and are expressed in percentage of satura- 
tion; for example, when it is stated that a certain humidity should be 
85 percent, this means that the air should be at approximately 85 per- 
cent of complete saturation with water vapor at the recommended 
temperature. Allowance has been made for the facts (1) that the 
temperature of fruits and vegetables in transit is usually higher than 
the recommended storage temperature, (2) that, except during winter 
weather, ripening or other changes are likely to go on more rapidly 
during a given transit. period than during the same length of time 
in cold storage, and (3) that when the commodities arrive at destina- 
tion, even if they had been in storage at shipping point, they are likely 
to be somewhat more mature than if they had remained in storage the 
whole time. 


‘TEMPERATURE OF STORAGE RooMs 


If the best results are to be obtained in the cold storage of the prod- 
ucts discussed herein, it is highly important that the temperature in 
storage rooms be held fairly constant. Variations of 2° or 8° F. 
above or below the desired temperature are in most cases too large. 
They can usually be avoided if the storage rooms are well insulated 
throughout and have adequate coil surface and if the spread be- 
tween the temperature of the refrigerant and that of the room to be 
refrigerated is kept small. For example, in a room where 45° is 
the desired temperature, using ammonia evaporating at about 26° 
in the coils, fluctuations of =114° may occur in the air temperature 
in the room; whereas at 32°, with ammonia also at 26°, fluctuations 
in the room temperature are usually less than +14°. However, 


* Italic numbers in parentheses refer to Literature Cited, p. 54. 


4 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


the spread between the temperature of the refrigerant and that of 
the room is more important in maintaining humidity than in pre- 
venting fluctuations in the room temperature. This point is dis- 
cussed in more detail on page 5. Storage rooms should be equipped 
either with reliable, accurate thermostats or with means for manual. 
control which are given frequent personal attention by someone 
charged with that duty. Even when reliable automatic controls are 
used, they should be checked periodically by some responsible 
person. 

In commercial cold-storage rooms thermometers are usually placed 
at a height of about 5 feet, sometimes slightly lower, for conven- 
lence in reading. It is important, however, to take temperatures 
frequently at the floor and the ceiling also, and at any other places 
where they might be expected to be undesirably high or low. In 
providing recommended temperatures for given products, considera- 
tion should be given to differences between the temperatures of the 
air at the position of the thermometer and at different places around 
the packages of the stored product, and also to differences between 
air and commodity temperatures. Often the packages are piled too 
closely together or distribution of refrigeration is inadequate to reach 
all parts of the piles of the stored commodity, and it is not unusual 
under such conditions to have commodities remain for several days 
or even weeks at temperatures several degrees higher than those in- 
dicated by the thermometer. ‘This condition can be detected by open- 
ing the pile and taking commodity temperatures and can be cor- 
rected by wider spacing of packages and the use of portable fans 
and bafiles for the direction of air currents to the centers of the 
piles. The installation of such equipment will also be useful in pre- 
venting dead-air pockets that are hard to cool, especially in rooms 
cooled by direct-expansion coils or by brine coils and without means 
for air circulation. . 

The importance of maintaining fairly constant temperatures in 
cold-storage rooms lies in the effect of such control, or the lack of it, 
on the keeping quality of stored commodities. Most varieties of 
apples keep best and longest if held constantly at 30° to 32° F.; the 
best temperature for Bartlett pears is between 29° and 31°. If the 
air temperature where either of these fruits is stored rises 2° or 3° 
above the upper limit mentioned, there is danger of increased decay 
and undue ripening, the danger being greater the longer the period 
during which the temperature is above 32°. For example, 3 or 4 
days at 35° usually would have little or no effect, partly because of 
a slower rise in the temperature of the fruit than in that of the air; 
‘but 10 days at this temperature would probably shorten the life of 
the fruit by about a week and possibly result in more decay. On the 
other hand, if the temperature goes a degree or two below 29°, 
there is a chance that freezing will occur. Celery and cabbage al- 
‘lowed to remain too warm in storage may show yellowing and decay; 
potatoes are likely to begin to sprout if the temperature is too high 
and usually become undesirably sweet if it is too low. Other com- 
modities undergo these or other kinds of deterioration if the tem- 
perature variations throughout long storage periods exceed the limits 
given for them in this circular. In addition, there is always the 
possibility that fluctuations in temperature will cause condensation 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS  § ~- 


of moisture on stored products, which in itself is undesirable because 
it favors the growth of mold and the development of decay. 

Control of temperature is usually easier in large rooms than in 
small ones if both are filled to capacity. This is because of the “‘fly- 
wheel” effect produced by the larger mass of material, including 
both the commodity and the building material. Refrigeration is thus 
stored up, so to speak, and the temperature changes occur more slowly. 
For this reason small storage rooms generally will require closer 
attention than large ones. 


Humiupity oF STORAGE Rooms 


The relative humidity of the air in storage rooms has a direct 
relation to the keeping quality of the products held in them. With 
only a few exceptions (see especially onions), the most desirable con- 
dition is that in which the humidity of the air in the storage room 
is kept at approximately the moisture content of the produce stored 
therein. If it is too low, wilting is hkely to occur in most fruits, 
vegetables, cut flowers, etc.; if it is too high, it favors the develop- 
ment of decay, especially in rooms where there is considerable varia- 
tion in temperature. The exact control of humidity is rather diffi- 
cult, however, and in the past was not often attempted in commercial- 
storage warehouses. However, at the present time new plants are 
being designed and old ones changed over in such a way that a foun- 
dation for humidity control in cold storage is being developed. 
Sufficient refrigeration, whether coils or brine spray, is provided so 
that the room can be operated with only a small difference in tem- 
perature between the refrigerant and the air of the room, thus rais- 
ing the dew point of the air and its relative humidity. This permits 
the maintaining of high humidity. For low humidity the tempera- 
ture of the refrigerant is lowered. To build up humidity quickly 
in cold-storage rooms some operators evaporate water from insulated 
open-top tanks equipped with insert electrical heaters. Others use 
a fan to beat water into very fine particles and to blow this spray 
into the room. ‘These humidifiers are sometimes made automatic, 
connecting with humidistats. However, if the coil temperature is 
too low, that is, if the spread between the temperature of the refrig- 
erant and that of the room is too great, the effect of these various 
measures will be short-lived. The relative humidity will soon become 
undesirably low, and harmful drying out of the stored product will 
occur. 

When warm_products are placed in a cold room the spread between 
commodity temperature and refrigeration temperature is wide at 
first. Such a condition hastens cooling and shortens the time during 
which undesirable changes can occur in the commodity. As the 
spread is reduced, the cooling proceeds more and more slowly. 
Eventually, if the temperature of the refrigerant is properly adjusted 
and there is adequate refrigerated surface, the desired temperature 
can be reached without subjecting the commodity to excessive desic- 
cation. If it could be successfully managed under commercial con- 
ditions, a temporary reduction in refrigerant temperature below what 
is desired or safe for long-time operation would be extremely helpful 
in shortening the initial cooling period, 


6 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


Operators of certain large cold-storage houses have found that 
when a room is filled with apples or celery, for example, the relative 
humidity soon becomes constant at a fairly definite level, which for- 
tunately is about the optimum for the particular commodity con- 
cerned. The same result has been observed with other commercially 
important fruits and vegetables and probably is obtained in many 
of the larger well-constructed cold-storage warehouses of the country. 
This is probably due to the fact that when a storage room is nearly 
full and there is comparatively little air space left the pull on the 
moisture in the produce is less, and it is easier to maintain the desired 
humidity. In these cases, however, it must be assumed that the spread 
between coil temperature and air temperature is kept narrow. 

For most fruits that are stored commercially the statement is true 
that a relative humidity of 80 to 90 percent gives the best results. 
Exceptions are discussed at various places in the text. For leafy 
vegetables and root crops the relative humidity should be about 90 
to 95 percent; for other vegetables, except as noted in the text, 85 to 
90 percent. If it seems necessary to increase the relative humidity 
in rooms used for common, or air-cooled, storage, this can best be done 
by sprinkling the floor occasionally. Earth floors are more desirable 
in air-cooled storages than floors of concrete because they are more 
easily kept damp. An increase in air circulation calls for an increase 
in relative humidity if wilting of the stored commodity is to be 
avoided. Allen and Pentzer (2). found that doubling the rate of air 
movement increased moisture loss by about one-third and was equiva- 
lent to about a 5-percent drop in relative humidity. The drying effect 
of increased rate of air movement is particularly marked if the hu- 
midity of the air is lower than the moisture content of the commodity. 


EvoLuTIOoN oF Heat py CommopitTy 


In any consideration of the storage of fresh fruits and vegetables, 
cut flowers, etc., it should be remembered that these commodities are 
alive and that by virtue of that fact they carry on within themselves 
many of the processes characteristic of all living things. Unless the 
relative humidity is high, they give off moisture to the surrounding 
air and most of them, in time, become shriveled or wilted, even at 
32° F. The enzymes, or ferments, they contain act on various sub- 
stances in their tissues and gradually bring about changes in color, 
texture, and chemical composition which ripen the commodity and 
may result in serious deterioration or even complete break-down. 
The most important of these changes are produced by respiration, 
the process in which the oxygen of the air is combined with the carbon 
of the plant tissues, occurring chiefly in sugars, to form various de- 
composition products and eventually carbon dioxide and water. -Dur- 
ing this process energy is released in the form of heat, the amount 
of which varies with the commodity and increases as the temperature 
increases, up to about 100°. This heat is always a part of the re- 
frigeration load which must be considered in handling fruits, vege- 
tables, and cut flowers in cold-storage rooms or refrigerator cars. 
The approximate rate of evolution of heat by various commodities is 
given in table 1. 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS STOCKS if 


Taste 1.—Approximate rate of evolution of heat by certain fresh fruits 
and vegetables when stored at the temperatures indicated 


Heat evolved per | Litera- 
C dit Temper-| ton of fruits or ture 
CO UN ature | vegetables per 24 | refer- 
hours ¢ ence 
oars Bates 
adbte | 32 1, 500 
Yellow Dnansparent 22. 2-4 ue ab 2 oom (°) 
70 12) 380 
32 700 to 800 
a Ouraeas laren 2 Cire cpr ae er etn Weare eee 40 820 to 840 
60 2,610 to 3,470 || os, 
32 300 to 320 
AYU OUSISEE la pe ir eae ty aad uae 40 590 to 600 
2 60 2, 270 to 2, 350 
ananas: 54 3. 300 
Cike ein = ape ee ee eee { 68 8 360 
Rania ge a A 68 9,240 |f (62) 
Jet OSes SE salut ti Aa erg er ere 68 8, 360 
Beans, lima: ae Fe aan 
IRGTHG| CCY0)| ess Hes oe Raa ese (6 crea Ae 60 21, 990 (°) 
70 29, 220 
32 3. 160 
Variety, unknown0gs Ss. 3 as se oe Ap sek re 
70 37, 120 
Beans, snap: ti > eae 
Stringless flat-podded_____._...____- 60 39° 090 (°) 
70 45, 370 
39 5, 500 
BlackpVielembines 2220 shin ee oo oll ab rin Be 
70 52, 950 
32 6, 160 
StiringlesstGreen Pod! 9221. fk =. 1) ap ni oe @ 
80 49, 590 
32 2) 650 
IBeetsn (Cope dass van ae Ne ahs 40 4, 060 (2) 
60 7, 240 
32 7, 450 
Broccoli (variety unknown)____________ an aS! a €2) 
70 47, 340 
32 1, 200 
Cabbage (Globejus (en 8) las 1 a NETO 
70 6, 120 
32 1, 320 
Cantaloupsy (Turlock) s-20 2 1 Ve 40 1, 960 
60 8, 500 (2) 
32 2, 130 
DALEOES | CCOPIEE) ees tee so Meee ts eg ah PI Be 40 3, 470 
60 8, 080 
32 1, 620 
Celery (New-York white): _-2_@2_2__2_ a a aan (¢) 
70 14, 150 


See footnotes at end of table, p. 9. 


§ CIRCULAR 278, U. 8. DEPARTMENT OF AGRICULTURE 


Taste 1.—A pproximate rate of evolution of heat by certain fresh fruits 
and vegetables when stored at the temperatures indicated—Con. 


Commodity 


Gherriess(Sour)22 2 a Ea ee ae 


Cranberries: 
Harly Black 2—_= == — pe cette ao AEN So 


Cucumbers (Producer and Wanchula) __- 


Grapelrul the ene ee sg 


Grapes: 
Concord mies Ao 2 eevarbiee ie ee 


Cornichon and Flame Tokay _-__-_--__- 


Sultaainae se eke a Pe. 


Lemons? Giurekae. 2 225 _ a) ta ee 


Wet hice sb eee ee AF Be Rae eS oN = Se are ote 
Mushroomsi(cultivated)-< == 222 


Onions (VellowsGlobe) 2.2522 Sees se 


Oranges?) BRR ek tt ye I | 


Peachest#at i ts es tee ee 


Pears (Bartletweo* =e eee ee 


See footnotes at end of table, p. 9. 


SS 8 a LLL SSS S566 SSS 


Temper- 


ature 


Heat evolved per 
ton of fruits or 
vegetables per 24 
hours ¢ 


Bote us 
1, 320 to 1, 760 
11, 000 to 13, 200 
600 
870 
1, 800 
720 
970 
1, 650 
1, 690 
2, 550 
10, 460 
460 
1, 070 
2, 770 
4,180 
602 
1, 170 
3, 487 
8, 481 
660 to 1, 100 
2, 200 to 2, 640 
5, 500 to 6, 600 
430 
1, 050 
1, 690 
350 
850 
1, 810 
300 
740 
1, 570 
580 
810 
2, 970 
6, 200 
1320 
15, 990 
45, 980 
6, 160 
22, 000 
58, 000 
660 to 1, 100 
1, 760 to 1, 980 
3, 080 to 4, 180 
690 to 900 
1, 400 
5, 000 
8, 000 
850 to 1, 370 
1, 440 to 2, 030 
7, 260 to 9, 310 
17, 930 to 22, 460 


660 to 880 | 
8, 800 to 13, 200 | 


Litera- 
ture 
refer- 
ence 


— “= 

Co Cm Lo NS 
PS Coys & Mm 
Ww Ww 


= 
tS 

Co 
Neb 


Cm) 
i} 
? 


— 

Co 
tS 
4 


— 
- & 
Ve 


a 

S's) 
PSS 
=’ 


—= 
Or 
Co 

SS 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 9 


Taste 1.—A pproximate rate of evolution of heat by certain fresh fruits 
and vegetables when stored at the temperatures indicated—Con. 


Heat evolved per | Litera- 
i Temper-| ton of fruits or ture 
Commodity ature | vegetables per 24 |  refe- 
hours ¢ ence 
Sea Jaxmtiavyr 
32 8, 160 
Peas: 40 13, 220 
TnmpOrONe Gee AO Ge. see Sp eee SS oe Se 60 39, 250 
80 75, 500 
32 8, 360 
: 40 16, 020 
Gat ONEN A fy A Se ee NS Le | 60 44, 510 
80 82, 920 (4) 
{ 32 2, 720 
ED PETS: oo eee pores tee a oe Some 40 4, 700 
(ok 260 8, 470 
32 440 to 880 
Potatoes @nrishyCobbler) 2.27 = 2s 40 1, 100 to 1, 760 
70 2, 200 to 3, pao 
: 36 4, 400 to 6, 
Raspberries - - - - --------------------- { 60 | 15, 400 to 17, 600 \ Cy) 
] 32 4, 240 
Spinach: 40 7 850 
Bloomsdale*Savoy£s Lest) sia a 50 17, 940 
60 38, 000 (") 
32 4, 860 
Stes HAG 11, 210 
Vili CIM AVON ok ya 6 2 eae. hey Te | 50 20. 640 
60 36, 920 
32 2, 730 to 3, 800 
Sirawbertes 2) 22 74 babe ieee a 1 Senne oN (34) 
80 | 37, 220 to 46, 440 
Sweet corn (Golden Bantam cross; not ti e aoe 
husked) ---~----------------------- Goole 38, 410 
80 61, 950 
Sweetpotatoes (Nancy Hall): | 32 2, 440 
INTOER CUT CCS ee aee ts BENIN AAS RR ASIY ON aie 8 40 3, 300 
[Pee 60 6, 300 
32 1, 190 
OO [=a A eee om eee ie 28 ges eee | 40. 1, 710 
60 4, 280 (2) 
Tomatoes: 32 580 
Niaturevoneen tt B= iyi Ge Fiver so ork 40 1, 070 
60 6, 230 
32 1, 020 
J XD) OB) ek eat eal te a ep Ma gee 40 1, 260 
60 5, 640 
32 1, 940 
Munmipse (copped) =... a5 fe ne ee 40 2, 150 
60 5, 280 


2 The figures in this column were obtained (1) by assuming that the heat lib- 
erated by respiration is produced by the respiration of a hexose sugar, and (2) by 
multiplying the milligrams of carbon dioxide produced per hour by each kilo- 
gram of respiring material by the factor 220. 

’ Unpublished work by T. M. Whiteman. 

¢ Unpublished work by Morris Lieberman. 

4 Unpublished work on the respiration of vegetables by R. C. Wright and 
T. M. Whiteman. 

¢ Unpublished work.by W. T. Pentzer. 

f Unpublished work by D. H. Rose and M. H. Haller. 


829965—49——2 


10 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


It will be noted that lettuce, Bartlett pears, peaches, and cherries 
have a much higher respiration rate than potatoes, apples, and onions. 
- This means that the first group require considerably more refrigera- 
tion than the second to keep them at a specified temperature. Less 
pronounced differences occur between other commodities in the list 
and are important to a lesser degree in determining the amount of 
refrigeration necessary to cool them and keep them in sound, usable 
condition. 

It is interesting to note that the storage life of apples (as repre- 
sented by the three varieties with short, medium, and long storage 
périods, table 1) varies inversely as the rate of evolution of heat. The 
same relation holds true if one considers broccoli, lettuce, peas, spinach, 
and sweet corn in comparison with carrots, onions, storage varieties 
of grapes, and potatoes. 

It is difficult to determine the heat to be removed in cooling fruits 
and vegetables to cold-storage temperatures. This depends mainly 
on the following factors: The specific heat of the product, the rate 
at which it produces heat (by respiration), and its initial and final 
temperatures. If the product could be cooled to the storage tempera- 
ture instantaneously, the heat to be removed would be only the num- 
ber of British thermal units (B. t. u.) or calories obtained by multiply- | 
ing the specific heat of the product by the difference between the 
initial and the final temperature, and this result by the weight of the 
product in pounds or kilograms. This is usually called the sensible 
heat. The cooling process, however, requires time, and during this 
interval additional heat is produced by the respiration of the stored 
fruit or vegetable. 

In order to determine the amount of this additional heat it is 
necessary to know the rate of heat production at any temperature 
and the length of time the product is in each temperature range. For 
example, if the respiration rate (or rate of heat production) for a 
given commodity is twice as great at 70° F. as at 50°, the number 
of hours this commodity is at each of these temperatures must be 
known before the total heat produced can be calculated. When fruits 
and vegetables cool, the rate at which they produce heat decreases, 
and the total heat produced depends not only upon the time required 
for cooling but also upon how long the commodity stays in each tem- 
perature range. 

Table 2 shows the approximate amounts of sensible heat and of 
heat produced by respiration which must be removed from eight 
varieties of five kinds of fruit in cooling them from various tempera- 
tures to a temperature of 35° F. These figures are based on experi- 
mental determinations of the rate of respiration at various tempera- 
tures; some of the data are from the tables given by Magness and his 
associates (53, 55), and the remainder from data reported by Haller | 
etal. (34). The figures for Bartlett pears are based on the maximum 
values given by Magness and Ballard (53). The figures given in 
this table have been obtained by assuming that the heat of respiration 
is produced by oxidation of a hexose sugar and can be calculated from 
the rate of production of carbon dioxide, which has been determined 
experimentally; very few calorimetric measurements of heat produc- 
tion by fruits and vegetables have been made, and this assumption 
seems to be the best available basis for calculating heat production at 


11 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 


any given temperature. Recent investigations (30) indicate that cal- 
culations made on this assumption give values that are within 10 per- 
cent of those obtained calorimetrically. 

The assumption has also been made, although it is believed to be 
only approximately correct for fruits and vegetables, that the rate 
of temperature drop at any given time during cooling is proportional 
to the difference between room temperature and fruit temperature 
at that time. With this assumption as a basis, the temperature and 
thus the rate of respiration at any time during the cooling period, as 
well as the total heat produced during the whole cooling period, have 
been calculated. * 


TasLEe 2.—Approximate amounts of heat of respiration and sensible 
heat to be removed from certain fruits in cooling them from 60°, 70°, 
or 80° to 35° F. in a room at 32°, when the cooling takes place in 
3, 4,5, 6,8, or 10 days 


Initial| Heat of respiration per ton of fruit during— 


Sensi- 

Kind of fruit a ble 
ture |3 days|4 days/ 5 days|6 days] 8 days | 10 days heat ! 
CORES ESE FRU SSIEA URIS CALs Dt bs UL Lee be Ue Bit! we: €b- 
Apples: 80] 8, 000]11, 000)14, 000|16, 000] 22, 000] 27, 000/80, 000 
Wanesapet. -= 70| 7,000} 9, 000/12, 000/14, 000} 19, 000} 23, 000/62, 000 
60] 6, 000) 8, 000/10, 000/12, 000) 16, 000} 20, 000/44, 000 
[ 8012, 000/16, 000|20, 000/24, 000] 32, 000] 40, 000/80, 000 
Grimes Golden__|; 70/10, 000/13, 000/17, 000/20, 000] 26, 000] 33, 000/62, 000 
Leap 8, 000/11, 000/13, 000/16, 000) 21, 000) 27, 000/44, 000 
Peaches: 80/13, 000/18, 000/22, 000/26, 000] 35, 000] 44, 000/80, 000 
Hibertate 2 ow 70/10, 000}13, 000/17, 000/20, 000] 26, 000] 33, 000/62, 000 
60| 7, 000/10, 000/12, 000/14, 000] 19, 000] 24, 000/44, 000 
80/16, 000/22, 000/27, 000/32, 000] 43, 000] 54, 000/80, 000 
@armam.20 4. 70/13, 000/17, 000/21, 000/25, 000] 34, 000} 42, 000/62, 000 
pee: 60/10, 000/13, 000/16, 000/19, 000} 25, 000} 32, 000/44, 000 
“Ba, jae i 70/16, 000/22, 000/27, 000/33, 000] 44, 000} 54, 000/61, 000 
Ae ae ara 60/13, 000/17, 000/22, 000/26, 000] 35, 000} 43, 000/43, 000 
Strawberries: { 80/30, 000/39, 000/49, 000/59, 000! 79, 000} 99, 000/83, 000 
Chesapeake _-_ _-_- 70|24, 000/32, 000/40, 000/49, 000] 65, 000} 81, 000/64, 000 
ee eo 19, 000/26, 000/32, 000/39, 000} 51, 000) 64, 000/46, 000 
80/38, 000/51, 000/64, 000/77, 000)102, 000/128, 000/83, 000 
Howard 17_____- 70/31, 000/42, 000/52, 000/63, 000] 84, 000/104, 000/64, 000 
Oranges: 60/25, 000/34, 000/42, 000/51, 000} 68, 000} 84, 000/46, 000 
Florida seed- 80] 9, 000/12, 000/15, 000/18, 000} 24, 000} 30, 000/81, 000 
linge Se. 70| 8, 000/10, 000/13, 000/15, 000] 20, 000] 26, 000/63, 000 
60] 6, 000! 9, 000/11, 000/13, 000] 17, 000} 22, 000!45, 000 


1 For any one kind of fruit at a given temperature these figures are assumed to 
be the same for all cooling periods included in the table. 
2 The rate of respiration is practically the same for both Florida seedling oranges 
and California navel oranges. 
*The authors wish to acknowledge the valuable assistance given by W. V. 


Hukill, Division of Farm Buildings and Rural Housing, Bureau of Plant Industry, 
Soils, and Agricultural Engineering, in making the calculations and in preparing 
this statement on the production of heat by fruits and vegetables. 


12 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


As a result of these calculations it has been found that the heat 
produced by the respiration of fruit while it cools is directly pro- 
portional to the length of the cooling period. The figures for cool- 
ing periods of 3, 4, 5, 6, and 8 days are therefore set at 0.3, 0.4, 0.5, 0.6, 
and 0.8 (to the nearest thousand) of the figure for 10 days. The spe- 
cific heat has been calculated by the formula S=0.008a@ + 0.20, in 
which S signifies the specific heat of a substance containing a percent 
of water; 0.20 is the value that has been assumed to represent the 
specific heat of the solid constituents of the substance in question (88). 

Column 1 of table 2 shows the kind and variety of fruit and col- 
umn 2 the temperature of the fruit at the time cooling started, in a 
room held at 32° F. The next column shows the amount of heat 
evolved by respiration if the fruit reaches 35° at the end of 38 days. 
The next five columns show the amount of heat if cooling to 35° re- 
quires 4, 5, 6, 8, or 10 days, respectively. ‘The last column is the 
sensible heat (obtained by multiplying the specific heat of the fruit 
by the difference between initial and final temperatures and this 
result by the number of pounds in a ton).. For any one kind of 
fruit at a given initial temperature, the specific heat is assumed to be 
the same for all the cooling periods included in the table. 

The values given in table 2 are only approximate. However, in 
view of the results of the investigations mentioned on page 10, it is 
believed that the two assumptions that have been made—namely, (1) 
that the heat of respiration is produced only by the oxidation of a 
hexose sugar and can be computed from observed amounts of carbon 
dioxide produced and (2) that the rate of temperature drop is always 
proportional to the difference between fruit temperature and room 
temperature—probably lead to fairly accurate results. The figures 
are presented to help cold-storage-plant operators estimate the refrig- 
eration required for cooling the specified fruits under the various 
conditions given. As an example of how the figures can be used, 
the following calculation may be of interest: A ton of Bartlett pears 
cooling from 70° to 85° F. in 10 days in a 32° room is shown to be 
capable of producing about 54,000 B. t. u. Its sensible heat at 70° 
(35° above its final temperature) is 61,000 B. t. u. The sum of the 
two is 115,000 B. t. u. If this be multiplied by the capacity of the 
room in tons of fruit, say 600 (the capacity of some of the commercial 
cold-storage rooms in the United States) and divided by 288,000 (the 
number of British thermal units in a ton of refrigeration), the quo- 
tient 239 is obtained; this is approximately the number of tons of 
refrigeration required to cool 600 tons of Bartlett pears to 35° in 
10 days under the conditions specified. The corresponding figure for 
Winesap apples is 177 and that for Grimes Golden apples 200. 


EFFECT OF COLD STORAGE ON SUBSEQUENT 
BEHAVIOR OF FRUITS AND VEGETABLES 


The belief is rather common among those concerned with the mar- 
keting of fresh fruits and vegetables that commodities of this kind 
that have been in cold storage deteriorate more rapidly after removal 
from the low temperature than if they had been held at ordinary 
temperatures. It is difficult, however, to find a basis for judging 
whether or not they actually do so because there is no means known 
other than the use of low temperature for checking the ripening and 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 13 


decay so largely responsible for their deterioration—that is, it is 
impossible in the present state of knowledge to obtain fruits and 
vegetables that have not been refrigerated but still have not changed 
in any way since harvest, for comparison with similar lots that have 
been in cold storage and are eventually removed. However, so far 
as is now known, it is correct to say that for many fruits and vege- 
tables cold storage at or near 32° I’. is beneficial. Important excep- 
tions are bananas, sweetpotatoes, tomatoes, cucumbers, melons, and 
certain other horticultural products mentioned herein, the behavior of 
all of which in storage is discussed later under appropriate headings. 


SWEATING 


When fruits or vegetables are removed from a low temperature to 
a higher one there is frequently a condensation of moisture from the 
air on the cool surface of the commodity. This is known as sweating 
and is more marked the higher the relative humidity of the outside 
air. It should be prevented whenever possible in the case of onions 
and the more tender fruits, because it favors the development of decay. 
This does not mean that when any of these products sweat after 
removal from an iced refrigerator car or a refrigerated room they 
are sure to decay; it does mean that they are more hkely to decay 
than if they were dry after being unloaded and remained dry until 
consumed. In this connection dryness means merely the absence of 
liquid water on the surface. 

Sweating can be prevented to some extent, as in the British prac- 
tice with eggs and certain other commodities, by allowing the fruits 
er vegetables to warm up gradually. Under commercial conditions in 
the United States this is rarely practicable, however, and the best thing 
to do in very damp weather is to realize the risk, handle the product: 
carefully, and get 1t into consumption without undue delay. 


WAXING 


The application of waxing preparations to certain perishable prod- 
ucts has been practiced commercially for several years. It probably 
started with the waxing of citrus fruits and was followed by the wax- 
ing of rutabagas. Its value for both of these products lies in the fact 
that it not only improves their appearance but also prevents shrinkage 
by restricting the loss of water. 

Waxing has been tried on a number of other products. In the case 
of cucumbers and carrots it gives considerable promise and may 
come into general use. It has been tried on cantaloups but did not 
prove to be beneficial. 

(See 76.) 


CHOICE OF STORAGE CONDITIONS 


The storage conditions recommended herein for certain commodi- 
ties represent either a compromise between two undesirable extremes 
of temperature or humidity or a choice of the least harmful of two 
such extremes. Grapefruit, for example, suffers less from decay at 
low humidities than at high; on the other hand, at low humidities. 
it is subject to a pitting which, in fruit from regions where stem- 


14 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


end rot is not prevalent, is usually more damaging to the market value 
of the fruit, because it is more common than decay. Therefore, it is 
recommended that grapefruit from all producing regions be held 
at 85 to 90 percent relative humidity in storage in order to reduce 
pitting. 

The Jonathan variety of apple suffers less from soft scald if stored 
at 34° to 86° F. than if stored at 82°. It is susceptible, however, to the 
more common and more serious condition known as internal break- 
down and also to Jonathan spot and decay if held continuously at 
temperatures above 32°; hence this latter temperature should usually 
be chosen as safer than any higher one. 

Investigations in New York State (97) have shown that one lot of 
apples may have a stimulatory effect on the ripening of another lot 
stored with it. In some tests as much as 50 percent or more of the 
storage life of a lot of apples was lost because of exposure to the emana- 
tions from other apples. The agent causing the stimulation is pre- 
sumed to be ethylene, but it was not identified as such. The most 
effective absorbent for removing the stimulatory agent was activated 
coconut-shell carbon. 

Combinations that should be avoided in storage rooms are apples 
with celery, cabbage, potatoes, or onions; celery with onions; and cit- 
rus fruit with any of the strongly scented vegetables. Under various 
commodities additional undesirable combinations are discussed. 


FRUITS AND NUTS 


By Dean H. Ross, senior physiologist 


The recommended temperature, relative humidity, and approximate 
length of storage period for the commercial storage of fresh, dried, and 
frozen fruits, and nuts are given in table 3. Detailed descriptions of 
these requirements are given in the text. 


APPLES | 
(Temperature, 30° to 32° F.; relative humidity, 85 to 88 percent) 


There is a wide variation in the storage quality of the different 
varieties of apples and of the same variety grown in different regions. 
For example, McIntosh grown in the Middle Atlantic States is prac- 
tically an early-fall apple not suitable for more than 2 or 3 weeks’ 
storage, whereas if grown in northern New York or New England 
it can be held for as long as 4 or 5 months. Such varieties as Northern 
Spy, Baldwin, and Rhode Island Greening grown in the Cumberland- 
Shenandoah Valley district or in the hot, irrigated valleys of the 
Pacific Northwest behave like fall varieties and are short-lived in 
storage, although suitable for winter storage when grown in New 
England, New York, Michigan, and other northern producing districts. 

The keeping quality of apples in storage is also definitely related - 
to the cultural and orchard sanitation practices of the grower, who 
alone is responsible for the production of sound, properly matured 
fruit. To have good keeping quality, apples should be mature and 
well-colored. When they have reached this stage, they are less likely 
to scald in storage and are in better condition generally to be held in 
storage for the maximum period than if they are either immature or 
overmature (55, 56). 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 


15 


Taste 3.—Recommended temperature, relative humidity, and approi- 
mate length of storage period for the commercial storage of fresh, 
dried, and frozen fruits, and nuts, and the average freezing pots 


f Tempera- 
Commodity ies 
OE: 

IND DIGS! as) eee ee na 230 to 32 
Atpnicoitis tents Vangt pret ax 31 to 32 
ANVOCR GOS “eyes ay te pe CI (3) 
Loy ae  . Sea i (4) 
Blackberriesso. (esse 31 to 32 
Cherniese: 2 (hits fakes als 31 to 32 
Coconuts etre) sere se ay 32 to 35 
Cranberries. sneer 36 to 40 
Dates sete ema al eel ee (4) 
Dewhbenriest. 2a ae ees 31 to 32 
Rigs). (fresh) eg! 35" Sex ee a 31 to 32 
Grapefruitee sees 2S. (4) 
Grapes: 

WimMiheha cre: Soyo kaso 30 to 31 

INTACT NCAT se OES 31 to 32 
Hemionget ti GI La ere, ee 55 to 58 
Dimes. *ellypsariy 3 ap eae ? 45 to 48 
Logan blackberries_______- 31 to 32 
Olives? Gresh) os see we 45 to 50 
Oranges 22 2 rea (4) 
Reachest page ac te ieee. 31 to 32 
Pears: 

IS argletgmesae enn 29 to 31 

Fall and winter varieties_| 29 to 31 
Pineapples: 

Mature green______-___ 50 to 60 

1 ats) Osc es See Stee nd a ae 40 to 45 
Plums (including prunes) __| 31 to 32 
Quincese eee eee et ee Sle tOroe 
Raspberries hf st ai Sat 1 31 to 32 
DLEAWOCITICS sap v See 31 to 32 
Dricdtinuits. 28 2. ee ts (4) 
Mrozem fruits= ee ee (4) 
INGGS Mere See hon bees 432 to 45 


Relative 
humidity 


Percent 


85 to 90 


85 to 90 
80 to 85 
85 to 90 
85 to 90 
80 to 85 
85 to 90 
85 to 90 
80 to 85 


85 to 90 
85 to 90 


85 to 90 
'85 to 90 
80 to 85 
80 to-85 
80 to 85 
80 to 85 


(*) 
65 to 75 


Approximate 


length 


of storage period 


2 


7 to 10 days 4 


10 to 14 days_ 


ifto;2 imonthsuu sas 


( 
(tO LOl day cia ae ote ee ee 


Average 
freezing 
point ! 


LO day siete aia selon OS 


6 to 8 weeks_ 


> tO.O Months: =... 5 


3 to 8 weeks 4 


Lito 4mionths lie. 


6 to 8 weeks-_ 
7 to 10 days_ 
4 to 6 weeks_ 


1 These figures are based on previously published work by Wright (116) and 
are subject to revision whenever further investigation makes this necessary. 


2 See text and table 4. 

3 See text and table 5. 

4 See text. 

5 Green: Flesh, 30.2°; peel, 29.8°. 


Ripe: Flesh, 26.0°; peel, 29.4°. 


6 Hastern sour, 28.0°; eastern sweet, 24.7°; California sweet, 24.2°. 


7 Flesh, 28.0°; peel, 27.4°. 
§ Winter Nelis, 27.2°; Anjou, 26.9°. 


® Persian (English) walnuts, 20.0°; pecans, 19.6°; and chestnuts (Italian), 23.8°. 


To insure soundness and good keeping quality, apples must be not 
only properly grown and at the proper stage of maturity, they should 
also be handled in all the operations of picking, grading, packing, 
and hauling with that degree of care necessary to prevent serious 
bruising, skin punctures, or other mechanical injuries; and they should 
be stored as quickly as possible after they are picked. 


16 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


Apples should not be handled while frozen, if such handling can 
possibly be avoided. Water core does not develop or spread in storage, 
and in varieties such as Yellow Newtown and Winesap it may actually 
disappear after a few months’ storage, especially if originally present 
only in mild form. When large portions of the flesh are affected, 
especially in soft-textured varieties like Jonathan, Delicious, Stayman 
Winesap, and Rome Beauty, there is danger of subsequent break-down 
and prompt disposal of the fruit is advisable. The diseases of apples 
in storage are discussed in Farmers’ Bulletin 1160 (8) and Miscel- 
laneous Publication 168 (84). 

For the storage of most varieties of apples the best results are 
obtained by maintaining a temperature of 30° to 32° F. and a relative 
humidity of 85 to 88 percent. However, as the storage temperature 
approaches the freezing point of the fruit the hazard of freezing 
increases unless the temperature is well controlled and there is good 
air circulation. Yellow Newtown apples from the Pajaro Valley, 
Calif., and McIntosh and Rhode Island Greening apples from New 
York should be held at 35° to 38° rather than at 32° to prevent the 
development of internal browning or brown core. Grimes Golden 
apples should be held at 34° to 36° instead of at lower temperatures 
(77) in order to avoid soggy break-down. It should be remembered, 
however, that at these higher temperatures the fruit cannot be held 
as long as when stored at 30° to 32°, because of the possible develop- 
ment of Jonathan spot and internal break-down. 

(See 9, 21, 36, 43, 71, 72, 73, 74.) 

If air-cooled storage is used, the temperature obtainable will usually 
not be much lower than the average of the prevailing outside tempera- 
tures. The nearer this is to 32° F. the better. 

The length of time apples can be held successfully in cold storage 
will vary with the variety and with the district where grown, as well 
as with their condition when harvested. Table 4 shows the normal or 
average storage period and the maximum storage period for the more 
important apple varieties when picked at proper maturity and stored 
immediately at 30° to 32° F. 


Taste 4.—Normal and maximum storage periods for important apple 


varieties 
Storage period Storage period 
Variety Variety 

Nor- Maxi- Nor- | Maxi- 

mal mum mal mum 

Months | Months Months | Months 

Jonathani=, 4 seeee 2 to 3 4 || Arkansas (Black 

Grimes Golden__-—-_-- 2 to 3 4 Areas oS 2 SS 4 to 5 6 
Mcintosh ets se sees 2 to 3 4360-51) SDeliciauste2 ae 1 eae 4 to 5 6 
Golden Delicious____- 3 to 4 5 || Northern Spy__-_-_-_-- 4 to 5 6 
Contlandi=* eset 3 to 4 Salli: Baldwin.) t Fie ht 4 to 5 6 to 7 
Rhode Island Green- || Rome Beauty_------ 4 to 5 6 to 7 
IN GY oe a ee ee 3 to 4 6") Ben’ Dawises ae ore 4 to 5 8 
Stayman Winesap____| 4 to 5 5'to 6) Winesaps 252 Naito’ 8 
York-lmperiales = 4 to 5 5 to 6 || Yellow Newtown_-_-_-_| 5 to 7 8 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 17 


In determining when to remove apples from storage the dealer 
must, of course, consider the market, but he must also allow for 
the more rapid softening that takes place at the higher temperatures 
to which they will usually be removed. Investigations by the United 
States Department of Agriculture (55) have shown that apples soften 
approximately twice as fast at 70° as at 50° F., twice as fast at 50° 
as at. 40°, and about twice as fast at 40° as at 32°, whereas at 30° the 
rate is about three-fourths that at 32°. 

Apples in cold storage should be inspected frequently, in order 
that they may be removed and sold while still in good condition. 
It is highly desirable that apples intended for storage be wrapped 
in oiled paper or packed in shredded oiled paper, in order to reduce 
* damage by scald as much as possible (22). Apples should not be 
stored in the same room with potatoes because of the danger that 
the former will absorb undesirable odors. On the other hand, the 
odors given off by apples are readily absorbed by dairy products; 
consequently the two should not be stored in the same room. (See 
also p 14.) 

(See 8, 81, 90, 19.) : 


APRICOTS 
(Temperature, 31° to 82° F.; relative humidity, 80 to 85 percent) 


Apricots are not stored commercially to any extent, although they 
will keep well for 1 to 2 weeks at 31° to 32° F. When harvested at a 
firmness pernitting storage and shipping, the fruit lacks flavor and 
has poor dessert quality after ripening but is satisfactory for canning 
purposes (7). 


AVOCADOS 
(Temperature, see text; relative humidity, 85 to 90 percent) 


Investigations in California (63) on the storage of avocados have 
shown that the best temperature for all varieties grown there, except 
the Fuerte, is about 40° F. The Fuerte discolors internally at this 
temperature but holds up well at 45°. At temperatures below 40° 
all the varieties investigated are likely to become discolored internally 
and do not soften when removed to a higher temperature. When 
properly stored. the Dickinson, Royal, Taft, and Queen are said to 
hold up well for about 2 months, the Spinks, Sharpless, and Challenge 
for 5 to 6 weeks, and the Rey, Fuerte, and Kist for about 4 weeks. 
Most of these varieties are of the Guatemalan race. 

No general recommendations can be made concerning the storage of 
varieties of avocados grown in Florida, Central America, or the West 
Indies, because of the wide variation among them in susceptibility 
to injury by low temperatures. Many varieties of the West Indian 
race are injured by exposure to temperatures of 50° to 53° F. for 15 
days (99, 100), whereas others (Pollock, Trapp) remain in good con- 
dition for 3 weeks when held at 42° (52). Varieties of the Guate- 
malan race are more resistant to cold, and those of the Mexican race 
are the most resistant of all. Some of the varieties that are least 
affected by cold (Lula, Taylor) can safely be held at 87° to 42° for 4 
weeks (52). Wolfe, Toy, and Stahl (7/5), in their work on the storage 
of Florida-grown avocados, obtained the results given in table 5. 

829965493 


18 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


Taste 5.—E'ffect of storage of various avocado varieties at different 
temperatures on their marketable life * 


< 


5 ea Period 
pti- erio there- : Gain in 
mum | held at | after at aed market- 
Variety 2 tempera-joptimum| room Ble & _| able life 
ture for |jtempera-|tempera- a F SOS ihe tie 
storage ture ture tons: ) eee storage 
softening 
| 
he Days Days Days Days 
Bollocks 8 ae Sn ms 42 21 3 3 17-19 
‘Prapp GWab) eects Ce 42 21 ae) | 2-5 18-20 
Wrallclaar: CVD) iy ie aaa sv as CB) Agee pene lca et as ae Ra re | A 
Collinson <G XG Wai 42-48 14 4-6 3-4 7-8 
Booth) 8 (Glo Wil) See ae see 42 14 4—6 | 1-4 7-9 
Poulan G eB) baie hy ein sein cece ee 37 28 3-6 i=) 20-24 
Paylor (Gye ieee a OE 37 28 4—6 | 2-3 23-26 


1 Adapted from Wolfe, Toy, and Stahl (1/74). ; 

2 WI, West Indian; G, Guatemalan; M, Mexican; G * M, Guatemalan X 
Mexican; G X WI, Guatemalan * West Indian. 

3 Cold injury noted even at 48° storage. 


At the higher temperatures mentioned, in the text, anthracnose, a 
fungus disease of avocados, will probably be an important factor in the 
storage of this fruit. At the lower temperatures decay is not likely to 
be troublesome after long storage. 


BANANAS 


_ (Temperature: ripening, 62° to 70° F.; holding ripe fruit, 56° to 60°. Relative 
humidity: green fruit, 90 to 95 percent; ripe fruit, somewhat reduced but 
not below about 85 percent) 

The banana is one of the fruits that must be shipped to market 
green, because in this condition it can be handled for a longer time 
without becoming overripe and without serious injury from bruising 
during the marketing process. Furthermore, bananas of the Gros 
Michel variety, which make up the great bulk of banana shipments 
into this country, if allowed to ripen on the plant become mealy, lack 
flavor, and are subject to splitting, with subsequent decay. 

The bunches of green bananas as they are received from the Tropics 
are usually ripened at a temperature of about 64° F., with a relative 
humidity of 90 to 95 percent or higher. If it is desired to hasten the 
ripening process, a higher temperature can be used (up to 70°) for the 
first 18 to 24 hours, but temperatures should then be reduced to about 
66°. In any case the relative humidity should be kept at 90 to 95 
percent or higher until the fruit becomes thoroughly colored. After 
this it should be reduced shghtly, but not below about 85 percent. 
Prolonged exposure to high temperatures will cause poor color and 
flavor and weak necks and will hasten decay. 

The lowest temperature at which green bananas can safely be held 
in order to delay ripening is about 56° F.; below this they suffer an 
injury known as chiling—a form of peel injury to which both green 
and ripe bananas are susceptible, caused by low but not freezing tem- 
peratures. Ripe fruit is slightly more susceptible to chilling mjury 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 19 


than green fruit. Fruit chilled in the green stage does not develop 
a bright-yellow color on ripening, but instead a smoky dull color. 
Fruit chilled after ripening will develop a dull-brown color when 
later exposed to higher temperatures and is very susceptible to han- 
dling marks, the slightest bruising causing discoloration. 

The best holding temperature for ripe bananas is generally con- 
sidered to be between 56° and 60° F. At this temperature they will 
retain their good appearance and flavor and remain edible for a week 
or 10 days, although ordinarily they will not keep firm enough for 
shipment more than half that time. 

(See 25, 103.) 

BLACKBERRIES 


Short storage only. See Raspberries. 


CHERRIES 
(Temperature, 31° to 32° F.; relative humidity, 80 to 85 percent) 


The extreme limit for the successful commercial cold storage of 
fresh cherries at shipping point is probably about 10 days to 2 weeks. 
Tt is doubtful whether fresh cherries from California and the Pacific 
Northwest can be held satisfactorily in cold storage for more than 
about a week after arrival at eastern markets. This would mean about 
16 days from harvest. If held longer than the period indicated they 
begin to lose flavor and the bright attractive appearance characteristic 
of the fresh fruit. The stems may also dry out noticeably, especially 
if the relative humidity is low. Color changes and decay can be 
retarded by the use of carbon dioxide gas in transit (28). In fact, 
the treatment has proved to be so beneficial in this respect that it is 
now being used commercially for shipments of cherries from the 
West and Northwest to eastern markets. 

The sweating discussed on page 13 is particularly troublesome on 
cherries, because of the dull appearance that it produces on the fruit 
when displayed for sale soon after removal from iced cars. 


CocoNuts 
(Temperature, 32° to 35° F.; relative humidity, 80 to 85 percent) 


Coconuts are best stored at 32° to 35° F. and can be held satisfac-.. 
torily within that range for 1 to 2 months. ! 


~ CRANBERRIES 
(Temperature, 36° to 40° F.; relative humidity, 85 to 90 percent ) 


A. large part of the cranberry crop is held at the bog every year 
until wanted for Thanksgiving and the Christmas holidays. When 
so held it is usually kept either in common (air-cooled) storage or in 
artificially refrigerated warehouses. The storage period is usually 
not longer than about 3 months and for that length of time the range 
from 36° to 40° F. (whether obtained in air-cooled storage or under 
artificial refrigeration) has been found to be the most desirable. 
Occasional lots of poorly colored fruit may be held at 45° to 50° for a 
few weeks in order to permit more rapid coloring than would occur 
at lower temperatures. Storage for more than about 4 months, in the 


20 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


range from 36° to 40°, is not satisfactory because of the common 
occurrence of end rot, a fungus disease, which can develop at low 
temperatures (J/8). Shrinkage of the berries as a result of water loss 
is also a limiting factor. Further facts that need to be kept in mind 
are that keeping quality depends to some extent on the maturity of 
the fruit, that some varieties keep better than others, and that there 
may be a difference in keeping quality from year to year in crops 
from the same bog. 

Cranberries for long-time storage are best held “in the chaff” in 
the picking crates as they come from the field. Thus handled they 
keep better than if sorted and cleaned previous to storage. 

Cleaned and sorted cranberries can safely be stored at the market 
for 5 to 8 weeks if held at a temperature of 32° F., but fruit held for 
a longer time at that temperature is likely to develop a “low-temper- 
ature break-down.” Berries in this condition are “rubbery” when 
pressed between the fingers, the flesh is permeated with red pigment 
from the skin, and some of the natural luster has disappeared. Such 
berries closely resemble those that have been frozen. Fruit held at 
36° to 40° is less likely to discolor and become rubbery, but if it has 
been cleaned and sorted it is more likely to suffer from decay than if 
held at 32°. 

(See 4, 23.) 


DatTEs 


(Temperature, see text; relative humidity, 65 to 75 percent; or 28° F. and no 
humidity control for cured grades) 


Dates absorb moisture and odors readily from the air. The rate of 
absorption is much less at temperatures below 32° F. than at those 
above 32°. Deterioration caused by humidity above 75 percent is 
slow at storage temperatures below 28°. The dates of commerce are 
of three grades with respect to storage life—dried, cured, and non- 
cured. The cured and noncured grades are perishable. A tempera- 
ture as low as 0° has no deleterious effect upon dates but is actually 
beneficial to them. 

Dates are of two different types, and fruits of each type are likely 
to be either dry, cured, or noncured. The cane-sugar type is usually 
firm, light-colored, and comparatively dry, whereas the invert-sugar 
type is usually softer, darker colored, and inclined to be shghtly sticky 
or sirupy. 

Deglet Noor, the most important variety grown in this country, is 
of the cane-sugar type. Dates of this variety, cured grade, keep well 
until March at 28° to 32° F. and for a year at 24° to 26° or lower, 
whereas the noncured grade requires 18° or lower for storage until 
March and 0° to 10° for a year. In Deglet Noor dates that have be- 
come overripe or have been held under unfavorable storage conditions 
the cane sugar is inverted and the dates become soft, sirupy, and 
darker in color. Such dates are commonly graded as “dark soft.” 
If they can be dried down somewhat, they can be stored at 28° to 32° 
until Christmas without becoming objectionably dark and sirupy, 
although a temperature of 0° to 10° will be needed if they are to be 
stored until March. If such dates are not cured, a temperature of 
0° to 10° is necessary for even short-time storage (39). 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 21 


Halawy, Khadrawy, Zahidi, and Saidy dates are all of the invert- 
sugar type, and the cured grades can be kept until Christmas at 25° 
to 32° F. without forming sugar spots but require a temperature of 
18° or lower if stored until March. Noncured grades of these varieties 
require 0° to 10° for even short storage. After Christmas it is well 
to shift all dates of the invert-sugar type remaining in storage to 
“freezers” at 0° to 10° (4). 


DEWBERRIES 


Short storage only. See Raspberries. 


Fics (FREsH) 
(Temperature, 31° to 32° F.; relative humidity 85 to 90 percent) 


Fresh figs in storage require a temperature of 31° to 32° F. and a 
relative humidity of 85 to 90 percent, but even under these conditions 
they cannot be expected to keep satisfactorily for more than about 10 
days. 


GRAPEFRUIT 
(Temperature, see text; relative humidity, 85 to 90 percent) 


Storage rooms for grapefruit should have a relative humidity of 85 
to 90 percent. Lower humidities are favorable to pitting, and higher 
ones may increase decay. 

For short-time storage, grapefruit can be held satisfactorily at a 
temperature of 32° F. For longer periods the temperature to be used 
will depend on the character of the fruit and the troubles most likely 
to be encountered. For fruit grown in sections where stem-end rot 
is prevalent, this disease is likely to be the determining factor; it will 
generally be advisable to use a comparatively low temperature range 
(32° to 34°). On the other hand if the fruit is grown in sections 
where stem-end rot is not prevalent, the limiting factors are likely to 
be storage pitting and watery break-down, which develop most seri- 
ously at temperatures of 40° or lower. For fruit from these sections a 
temperature of 45° to 55° is satisfactory, and the more rapid develop- 
ment of undesirable high color and the increase in blue mold and green 
mold rots at the higher temperatures have not been found as objec- 
tionable on such fruits as the pitting that results from storage at lower 
temperatures. 

Sound fruit that is not overmature or likely to suffer from stem- 
end rot can usually be held for 6 weeks without serious spoilage at 
the higher temperature mentioned above, and this storage period can 
sometimes be doubled with satisfactory results. Weak or overmature 
fruit requires close watching from the time it is removed from the 
tree, regardless of storage conditions. 

The percentage of stem-end rot in Florida and Texas grapefruit 
will be greatly reduced if the fruit is properly treated with borax or 
sodium metaborate, pulled from the tree instead of being clipped 
(112), and precooled before being shipped. ‘The disbuttoning that 
may occur during handling and packing is also effective in reducing 
loss from stem-end rot. As compared with stem-end rot, blue mold 
and green mold rots are relatively less important on Florida grape- 


22 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


fruit in storage. Stem-end rot is not known to occur on California 
and Arizona grapefruit. _ 
(See 10, 24, 92, 93, 94, 111, 112, 113.) 


GRAPES 
VINIFERA 


(Temperature, 30° to 31° F.; relative humidity, 85 to 90 percent) 


Large quantities of the European, or vinifera, grapes, grown princi- 
pally in California, are stored every year. The most important of the 
varieties stored are Emperor and Ohanez (Almeria). Olivette de 
Vendemian, Malaga, Sultanina (Thompson Seedless), Cornichon, and 
Alphonse Lavallee (Ribier) are also occasionally stored. Al] of these 
have low freezing points, lower in fact than the freezing point of any 
other important fruit, largely because of their high sugar content. 
Although for most varieties there is no danger of freezing injury at 
temperatures as low as 28° F. (73), they are usually held at tempera- 
tures of 30° to 31°. At low air velocities a humidity of 85 percent 
prevents excessive wilting of stems and berries without favoring the 
formation of mold growth. At air velocities of 100 to 150 feet per 
minute a humidity of 90 percent is desirable (2, 67). 

California grapes for cold storage are packed in kegs or drums in 
sawdust or in various types of lidded lugs with or without sawdust. 
Good results are usually obtained, although if the fruit or the sawdust 
is damp at packing time or becomes so in storage there is danger of 
damage by mold. Mold may develop also if the grapes have been 
handled carelessly and if there are numerous cracked or loosened ber- 
ries scattered through the pack. Varieties differ in keeping quality. 
The best storage varieties when properly handled can be held 3 to 6 
months in storage at 30° to 31° F. Emperor, Ohanez, and Alphonse 
Lavallee (Ribier) seem to keep better than any of the other storage 
varieties. Treating grapes with sulfur dioxide has helped to reduce 
spoilage in storage. In recent years the display type of lug has 
grown in favor for storage, because grapes packed in it can be re- 
fumigated. The common practice is to fumigate with concentrations 
of about 1 percent of sulfur dioxide before storage and to refumigate 
with about 0.2 percent of the gas at intervals of 10 days. Under these 
conditions fruit has been held fully as long in this type of package as 
in the more expensive sawdust chests or kegs. When fruit packed in 
display lugs cannot be refumigated in storage, 5 grams of sodium 
bisulfite is often added to the pads before packing, and this is fol- 
lowed by the usual prestorage fumigation. Mixing 5 grams of 
sodium bisulfite (per lug, chest, or keg) with the sawdust used in 
packing grapes has also been effective in checking decay (66). 

Storage quality varies considerably from season to season and 
seems to be adversely affected by rain just before and during harvest. 
Grapes picked before rains usually keep better than those picked after 
rains. : 

(See 73, 65.) 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 23 


AMERICAN 
(Temperature, 31° to 32° F.; relative humidity, 80 to 85 percent) 


The eastern, or American, varieties of grapes, the most important of 
which is Concord, are not adapted to long storage; and most of them 
do not hold up well under storage conditions for more than 3 or 4 
weeks, depending on the variety. After that time they begin to 
deteriorate in flavor and may suffer heavily from decay if the temper- 
ature is not kept close to 32° F. The Catawba keeps better than most 
other eastern varieties and, if in good condition when stored, can be 
held for 3 to 8 weeks even in common storage in the districts where 
this variety is grown on a commercial scale. 

Too low humidity is undesirable for grapes, since it causes shrivel- 
ing, especially of the stems. Stock intended for storage should be 
handled carefully to avoid cracking of the berries or loosening at the 
cap stem, because such injuries allow juice to exude and thus furnish 
favorable conditions for the beginning of decay. 

Muscadine grapes are shipped only short distances if at all and are 
not known to be held in cold storage anywhere in commercial quan- 
tities. | 

(See 2, 48.) 


LEMONS 
(Temperature, 55° to 58° F.; relative humidity, 85 to 90 percent) 


From the standpoint of preventing decay (blue mold rot, green 
mold rot, and alternaria rot) in stored lemons, the lowest temperature 
that can be used without freezing the fruit would seem to be the most 
desirable. The difficulty is that at low temperatures certain non- 
parasitic troubles which are fully as serious as decay develop; among 
these are red blotch, pitting, and membranous stain. With proper 
humidity and at temperatures above 50° F. red blotch and pitting 
practically never occur and membranous stain is greatly reduced. 
The best results are usually obtained by storage at 55° to 58° in a 
relative humidity of 85 to 90 percent. Under such conditions lemons 
can be expected to hold up satisfactorily for periods of 1 month to 
as long as 4 months, depending on their maturity and condition when 
stored (77). ‘Tree-ripened lemons, which are yellow when picked, 
do not keep well in storage. 

It is of the utmost importance that lemons be handled carefully 
during picking and packing in order to avoid clipper cuts, scratches, 
and bruises and consequent damage later by green mold rot and blue 
mold rot. The fungus that causes the latter is able to penetrate the 
uninjured skin of lemons but is likely to cause more loss if the skin 
of the fruit is broken at numerous places. It can also spread from 
one fruit to another in the package and for this reason is frequently 
referred to as “blue contact rot.” Air conditioning as now used in 
some of the lemon storage houses in California furnishes a means of 
preventing condensation of moisture on fruit and so decreases the 
danger of decay. 

Lemons and other citrus fruits should not be stored in the same 
rooms with dairy products because of the readiness with which the 
latter absorb odors. (See also p. 14.) Lemons in storage should be 


24 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


examined frequently to avoid loss from the development of decay or 
other deterioration. 
(See 11, 16, 39, 42.) 


LIMES 
(Temperature, 45° to 48° F.; relative humidity, 85 to 90 percent) _ 


Preliminary investigations with Tahiti (Persian) limes indicate 
that fruit from a well-kept grove may be stored satisfactorily at a 
temperature of 45° to 48° F. for 6 to 8 weeks, provided that the rela- 
tive humidity is kept above 85 percent or the fruits are wrapped 
so as to prevent moisture loss. Prevention of desiccation is very 
important. For best quality, the Tahiti lime should be picked while 
still green but after the fruit has become “full” and smooth, having 
lost the “dimpled” appearance around the blossom end. 

Key (Mexican, or Dominican) limes can be stored satisfactorily at 
the temperatures recommended for Tahiti limes. The preferred color 
for this variety on the markets of the United States is yellow. 

Temperatures above those recommended permit the development 
of stem-end rot, which is often a serious factor in the marketing of 
limes from Florida and the West Indies. 


LocGAN BLACKBERRIES 


Short storage only. See Raspberries. 


Ottves (FREsH) 
(Temperature, 45° to 50° F.; relative humidity, 85 to 90 percent) 


The best storage temperature for fresh olives (67) les between 
45° and 50° F., and the safe storage period is 4 to 6 weeks. At lower 
temperatures the flesh of green fresh olives becomes brown, beginning 
around the seed and at the stem end. Ripe fresh olives develop more 
browning than green ones, showing severe discoloration even at 50° 
if stored for more than about a month. 


ORANGES 
(Temperature, see text; relative humidity, 85 to 90 percent) 


Although oranges are ordinarily stored at about 38° F., experi- 
mental results have shown that for long storage (8 to 10 weeks) a 
range of 34° to 38° gives better results. However, within this range 
some decay, chiefly blue mold rot or green mold rot, may occur during 
storage of 2 months or more, and some fruit may begin to show pit- 
ting and brown stain of the rind. If stored for longer periods, decay 
increases and the spotted fruit may gradually turn brown over all 
or most of the surface. Watery break-down may develop, as in 
grapefruit. Stem-end rot is likely to develop in Florida fruit if the 
storage temperature is higher than about 34°. Among California 
varieties, Washington Navel oranges are more subject to decay (blue 
mold or green mold rots) than Valencia oranges. The Washington 
Navel is also subject to alternaria rot. 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 25 


Careful handling is necessary at all times to avoid injury to the 
fruit. Decay, chiefly blue mold rot or green mold rot, frequently 
follows injuries. 

A free circulation of air around the boxes is desirable for oranges 
as for other citrus fruit. A relative humidity of 85 to 90 percent is 
sufficient to hold the shriveling of packed oranges to a minimum and 
retards decay more than does a higher humidity. 

Oranges should not be stored with eggs or butter, or in places where 
it is possible for the orange odor to penetrate into egg- or butter-storage 
rooms. (See also p. 14.) It is desirable that oranges in storage be 
examined regularly and often to avoid loss from the development of 
pitting or decay. After such examinations, a decision as to how long 
the fruit can safely be left in storage should take account of the fact 
that if pitting and decay are found they may increase rapidly after 
the fruit is removed to higher temperatures. 

(See 72, 92, 93, 94, 111, 113.) 


PEACHES 
(Temperature, 31° to 32° F.; relative humidity, 80 to 85 percent) 


Peaches are not adapted to cold storage. However, if they are 
sound and well-matured but not overripe, they can be held at 31° to 
32° F. for 2 to 4 weeks, depending on the variety, with lttle or no 
bad effect on the flavor, texture, or appearance of the fruit. Storage 
for longer periods is usually harmful to all of these characters. 
The peaches lose their flavor and natural bright color, become dry 
and mealy, or wet and mushy, and show marked browning of the 
flesh, especially around the stone. The loss in flavor is more rapid 
at 36° and 40° than at 32°, and break-down develops sooner at 36° 
and 40° than at either lower or higher temperatures. The best stor- 
age varieties and the periods they can ordinarily be held in storage 
~ are Tuskena (Tuscan), 8 weeks; Early and Late Crawford, and 
Salwey, 4 weeks; Elberta and J. H. Hale, 3 to 4 weeks. Belle, Cham- 
pion, Hiley, and Carman are less desirable as storage varieties and 
cannot be expected to hold up well, even under optimum conditions, 
for more than 2 or 3 weeks. 

(See 7, 29, 33, 37, 38.) 


PEARS 


(Temperature, 29° to 81° F.; relative humidity, 85 to 90 percent) 


BARTLETT PEARS 


The successful storage of Bartlett pears (43, 53, 54) depends not 
only on the temperature and humidity in the storage room but also on 
the condition of the fruit when stored. If the highest quality is to be 
obtained, Bartlett pears for storage should not be removed from the 
tree until the ground color begins to lighten and the lenticels have 
corked over. If picked before reaching that stage, they have a marked 
tendency to wilt, scald, and break down in storage. They also tend 
to break down in storage if picked when too ripe. The most desirable 
temperature for the storage of Bartlett pears is 29° to 31° F. The 
relative humidity should range from 85 to 90 percent. The maximum 

829965494 


26 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


period for storage for canning and local fresh markets is about 90 
days, and for storage at shipping point and at terminal markets, 45 
to 60 days. 


FALL AND WINTER PEARS 


For fall and winter varieties of pears (43, 67, 68), such as Anjou, 
Bosc, Clairgeau, Comice, Easter Beurré, Hardy, Seckel, and Winter 
Nelis, the most desirable storage temperature is 29° to 31° F. A rela- 
tive humidity of 85 to 90 percent is most commonly used. However, a 
relative humidity of 90 to 95 percent is maintained in some pear stor- 
age rooms in order to prevent shriveling. Such humidities are main- 
tained in connection with air velocities of 100 to 200 feet per minute. 
The length of time for which it is safe to store these pears depends 
on the variety and when it is picked and also on whether the fruit 
is Shipped directly to a consuming center and there stored or is stored 
at the shipping point for a time and. later shipped to market. In- 
formation on these points is given in table 6. In using the table it 
should be remembered that wide differences in keeping quality are 
often found in pears from various producing sections of the country. 
If Bose, Flemish Beauty, and Comice pears are held in cold storage 
beyond their season they do not ripen satisfactorily or they may not 
ripen at all (27). For best ripening after storage these and most 
other varieties of fall and winter pears should be held at a temperature 
somewhere in the range from 65° to 70°, preferably about 65°. 


Taste 6.—Length of time at 30° to 31° F. for safe storage of certain 
varieties of pears at shipping point and after shipment to market 
(68) 7 


Length of 
Storage treatment and variety storage End of storage period 
period : 
Stored immediately after harvest: Months 
PRAT Gy ee outs eee St Reeth eee 2 to 3 | September to November. 
@omice et RI eres dy 2to 3 | November to December. 
I BY Ovo neg Medlin. Sucteranahen: Puen vans iis be arric pam el 3 to 3% Do. 
Clairgeauis tise EO ce Ps ec ne 6 | February. 
VARTA OE SU ea Set een oe eae negra 5 to 6 | March. 
Winters Néehs2ae nies Ses eee 6 to 7 | March to May. 
RaASbereBeCUTne 2s.) aeons Seen itor’, Do. 
Stored after 12-day transit period 
(precooled) : 
TD OU hes See wee 4to5 | March. 
LSTEEy ee yg ell I Maen SS agua a ea xtr 2to 3 | September to November. 
Comic eal wir yg he LEST AAR, ae 2to 3 | November to December. 
BOSC VUE Fe hey eb yt LPS a 2 to 3 Do. 
Claing eat Feu wed, Proce Pe OVER Ay 3 to 6 | November to February. 
WanitergNiehics es! tek vivs sph ole 2 6 to-7 | March to May. 


The commonest and most serious decays of fall and winter pears in 
storage are gray mold rot, caused by the fungus Botrytis, and blue 
mold rot, caused by the fungus Penicillium. Gray mold rot is able to 
spread from decaying to sound healthy fruit and for that reason is 
frequently called nest rot. Losses from this rot can be reduced by 
the use of paper wrappers impregnated with copper (74). In the 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 27 


Pacific Northwest blue mold rot, in the form known as pinhole rot, 
is sometimes more important on pears, particularly Winter Nelis, 
than gray mold rot. Losses from blue mold can be greatly reduced 
by careful picking and handling, prompt storage at 29° to 31° F. after 
harvest, and the use of paper wrappers to prevent direct contact. 
between: diseased and sound fruit. 

Kieffer pears, if they are sound, firm, and still green when stored 
and are held under the conditions recommended for other fall and 
winter pears, can be expected to keep satisfactorily for 2 or 3 months. 
If intended for storage, they and other varieties should be handled 
with extreme care during the picking and packing process, because 
even slightly bruised or rubbed places are very likely to turn black 
and seriously damage the sales value of the fruit. Investigations 
(50) by the United States Department of Agriculture have proved 
that a ripening temperature of 60° to 65° F. is essential for the at- 
tainment of maximum quality in Kieffer pears for either dessert ‘or 
canning purposes. 


PINEAPPLES 


(Temperature: mature green, 50° to 60° F.; ripe, 40° to 45°. Relative humidity, 
85 to 90 percent) 


Pineapples are not adapted to long storage. Fully ripe fruits can 
be held satisfactorily at 40° to 45° F. for 2 to 4 weeks. Mature green 
fruits should not be held at temperatures below 50° and even at this 
temperature some of them will retain part of the green color in the 
skin and will fail to develop good flavor in the flesh after removal 
to room temperature. The maximum storage period for such fruit 
at 50° is 3 to 4 weeks. When held at 60°, mature green fruit ripens 
slowly, but after 2 or 3 weeks losses from decay, chiefly black rot, may 
be expected. The relative humidity for pineapples in storage should 
range from about 85 to 90 percent. — 

(See 102, 110.) 


PLums (INcLuDING PRUNES) 
(Temperature, 31° to 32° F.; relative humidity, 80 to 85 percent) 


Plums and prunes (fresh) are not stored extensively and are not 
adapted to long cold storage. Such varieties as Wild Goose and those 
of the damson type store better than the softer fleshed plums, such 
as Santa Rosa, Beauty, Wickson, and Duarte. The storage period, at 
31° to 32° F., ranges from 3 to 8 weeks depending on the variety. 
After that time the soft-fleshed varieties are likely to become too soft 
for commercial handling, may suffer some darkening of the flesh, and 
lose somewhat in flavor. : 

One of the most important commercial shipping and storage varie- 
ties is the Italian Prune. Ata temperature of 32° F., 2 weeks is about 
the maximum cold-storage period for this fruit if a shipping period 
is necessary before it goes on the market. After arrival at market 
prunes shipped immediately after harvest can ordinarily be held in 
cold storage for about 3 weeks. If held longer there is danger that 
shriveling, mealiness, and internal browning, as well as abnormal 
flavor, will develop. Too much confidence should not be placed in 
the appearance and condition of the fruit while it is in storage, as 
more deterioration—decay, shriveling, and internal browning—may 


28 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


take place in 3 days after removal from storage than during the whole 
storage period. Fresh prunes shipped out of storage at shipping 
point cannot safely be stored again after arrival at eastern markets. 
Storage disorders can be prevented by partly ripening the fruit prior 
to storage at 31° to 32° or by holding it at 40° to 45°. 

(See 7, 26, 97.) 


QUINCES 
(Temperature, 31° to 82° F.; relative humidity, 80 to 85 percent) 


The behavior of quinces in storage is about the same as that of early 
varieties of apples such as Jonathan and Grimes Golden. 


RASPBERRIES 
(Temperature, 31° to 82° F.; relative humidity 80 to 85 percent) 


Fresh raspberries, blackberries, Logan blackberries, and dewberries 
are not adapted to storage and are usually not stored commercially. 
For short periods, 7 to 10 days, most of them can be kept in fair condi- 
tion by storage at 31° to 32° F. in a relative humidity of about 80 to 
85 percent. Young and Boysen dewberries cannot be stored satis- 
factorily for more than 8 to 4 days. 


STRAWBERRIES 
(Temperature, 31° to 382° F.; relative humidity, 80 to 85 percent) 


Fresh strawberries are not stored commercially except for very 
short periods; 10 days is probably the maximum. Even for so short 
a time as this the temperature must be kept below 40° F. to prevent 
loss from decay caused by certain low-temperature fungi such as 
gray mold and Phytophthora, the fungus which causes leather rot; 
31° to 82° is still better. After about 10 days, sometimes sooner, the 
fruit loses its fresh bright color, shrivels more or less, and deterio- 
rates in flavor (774). | 


Driep FrutIts 
(Temperature and relative humidity, see text) 


For the preservation of natural color in storage, cut dried fruits 
and dried berries that are not subject to sugaring are held at 26° F. 
with no humidity control or at 32° with a relative humidity of 70 
to 75 percent. 

Figs and prunes are best stored at 40° to 45° F. The relative hu- 
midity should not be over 70 to 75 percent, to prevent excessive absorp- 
tion of moisture. Dried apples, apricots, and peaches keep best at 
26° to 32°. Raisins should be stored at 40° to 45° and require a rela- 
tive humidity of 50 to 60 percent to keep them from absorbing 
moisture. The holding of dried fruit in high humidity at tempera- 
tures above 32° is likely to result in mold. The dried fruits men- 
tioned can be kept in marketable condition for 9 to 12 months at the 
temperatures and humidities specified. 

Dried fruit can be tightly stacked, without stripping, in large sohd 
blocks in storage rooms without injurious effect, and this method of 
handling the packages minimizes the absorption of moisture from the 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 29 


storage-room air. When nonventilated packages, such as those used 
for dried fruit and dates, are removed from cold rooms, the sweating 
that results occurs mostly on the outside of the package and the 
moisture can be prevented from penetrating into the fruit by allowing 
the packages to warm up before they are opened (6). 


FROZEN FRUITS AND VEGETABLES 


(Temperature, see text) 


Frozen fruits should be held at —10° to 0° F. if they are to be stored 
for several months. | 

For the freezing of fruits a temperature of 0° F. or lower is de- 
sirable for both small containers and barrels. If freezing takes place 
too slowly the same undesirable conditions may develop that are en- 
countered if the fruit is stored at too high a temperature after being 
frozen (17, 18, 96). . 

For best results frozen fruits should be held in airtight containers. 

The best temperature for freezing vegetables is from —10° to —5° F, 
For storage after freezing —10° to 0° is satisfactory if ample pro- 
vision is made for rapid cooling until the product reaches the 
freezing point. 


Nuts 
(Temperature, 32° to 45° F.; relative humidity, 65 to 75 percent) 


Most of the commercial nut crop, including walnuts of all kinds, 
filberts, almonds, Brazil nuts, peanuts, and sometimes pecans, is usu- 
ally held in ordinary warehouse storage through the winter follow- 
ing harvest. The portion of the crop (except pecans) that is to be 
kept through the following summer should be placed in cold storage 
early in March. Pecans become stale and rancid much sooner than 
most other kinds of nuts, and it is safer to put them in cold storage at 
32° I. shortly after harvest. Brazil nuts can usually be kept satis- 
factorily in warehouse storage during the winter, but that portion to 
be held over summer should be stored at 32° before warm weather. 
Brazil nuts should be carefully inspected before being accepted for 
storage to see that they are well dried out, or cured. Walnuts, filberts, 
and almonds usually need not be stored below 40° to 45°. Chestnuts 
are rarely held in any other way than in cold storage at 32° to 40°. 

Shelled and unshelled peanuts can be held at common warehouse 
temperature during the winter, but during spring and summer shelled 
peanuts should be kept in cold storage both for protection against 
sects and to prevent development of rancidity. If cold storage is 
not available, common storage can be used, but the peanuts should be 
stored in the shell and shelled out as needed, because if stored shelled 
they are likely to darken and become rancid. In common storage care 
should be taken to prevent infestation by insects. Fumigation is de- 
sirable for both shelled and unshelled peanuts if they are to be held 
in common storage during the summer for any considerable time. 

As all varieties of nuts keep better unshelled than shelled, it is 
usually the best practice to store nuts in the shell and crack them as 
needed unless the kernels can be sealed in vacuum, which will permit 
them to be kept even longer than in the shell. Generally nut kernels 
should be stored at 32° F. 


30 


CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


The relative humidity of the storage room should be 65 to 75 per- 


cent. 


lower humidities there will be undue drying. 


(See 1/7.) 


VEGETABLES 


By R. C. WRIGHT, physiologist 


The recommended temperature, relative humidity, and approximate 
length of storage period for the commercial storage of vegetables 
are given in table 7. Detailed descriptions of these requirements are 


given in the text. 


At higher humidities there is danger of mold growth, and at 


TasiE 7.—Recommended temperature, relative humidity, and approxi- 
mate length of storage period for the commercial storage of various 
vegetables, and the average freezing points 


Commodity 


ASPAT AGUS Oia ems 
Beans: 
Green orisnapi2s 2) 8 
Lima: 
Unshelleds2u2 2. ius 


Broccoli (Italian, or sprout- 
ing). 

Brussels’sproutse) f2 rec) 

Cabbage 


Celeriac 
Celene = A ed ama a 
Corn (ereem) mS em 
Cucumbers@ is) 12 ORS 
Bgeplamtisies ) aye sia ee 
EINGhiiy ee Se er URS ees 
Garlic (dry) 
Horseradish 


TEC TEUCES SF ee nak et tee he 

Muskmelon (cantaloup) - 

Honey Dew and Honey 
Ball. 


CO) ethene lin wi PR tan GE ES aE 


ture 


A oi 
32 


32 to 40 


32 
32 to 35 


32 to 35 
32 


36 to 40 
32 to 34 
36 to 38 
36 to 40 


32 tO oo 
50 


See footnotes at end of table, p. 31. 


Tempera-| Relative 
humidity 


Approximate 


length 


of storage period 


Average 
freezing 
point ! 


Percent 
85 to 90 


85 to 90 


85 to 90 
85 to 90 
85 to 90 
85 to 90 


95 to 98 
85 to 90 
90 to 95 


90 to 95 
90 to 95 


95 to 98 
85 to 90 
85 to 90 
95 to 98 
90 to 95 
85 to 90 
85 to 95 
85 to 90 
90 to 95 
70 to 75 
95 to 98 
90 to 95 
95 to 98 
85 to 90 
90 to 95 


75 to 85 
75 to 78 
75 to 85 
75 to 85 


80 to 85 
85 to 95 


1 to 3 months 


10 to 14 days. _+___- 
CAO iircenge Bk se 


3 LO 4nWwEeeKSE OU WR cpa 


3 to 4 months 


10 to 14 days_______ 
2 sto 3oweeksels Sis 


3 to 4 months 
2 to 4 months 


10 to 14 days______- 
LOndhaiyige! cary 25.05 pee Bye 
DitOye WeCKS Ee Sass 


6 to 8 months 
10 to 12 mont 
2 to 5 months 


Agte) 


2 to 4;rweeksis_ 2s 


1 to 3 months 


2 GO OEWee KS = ae ee 


(aLOnlOKday c= eae 


2*to 4 weeks= S50 


A to Gsweeks4 2) 222 eee ee 


2 to 3 days__ 


Diweekgeis \ oan aaees 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 31 


Taste 7.—Recommended temperature, relative humidity, and approxi- 
mate length of storage period for the commercial storage of various 
vegetables, and the average freezing points—Continued 


; : Average 
5 Tempera-| Relative | Approximate length : 
Commodity. ture humidity of storage period ee 
point 
Celie Percent oe aA 
Onionceeyh. fer en SZ COMCOnLom OO tOre montis.) = a2 5 30. 1 
Onion sets! =e so ese Si neh OMCOnGa| 2 === GOn et ree as 29. 5 
1 EAH STON OFS gy ak Ea S237 90) to, 97 2.to 4: months. 22 = 28. 9 
Peasr(ereem)e — 7.25 23. cs 38214| 85) to 90) a to 2 weeks____- = 30. 0 
Peppers: 
Chili Gres yeaa se sk (2) MOMCOniS al OrtorOumontbhs: 2) aya) a eT 
Syweebe Sefer ane 32 | 85 to 90 | 4 to 6 weeks_______- 30. 1 
Potatoes: 
Byam yes, Ses eS yey ps Aesth (?) Sou Os9O el) sede ey Sees ee Ue En cas Sag 
LE ShaRde Seat ep Seah get fy Les SSELOLO OMe SoOMbOnO OAs (2) iis a ths Melee A. nla s 28. 9 
Pumpkins 2 23 S24 ta ts ro 50 to 55 | 70 to 75 | 2 to 6 months______-_ 30. 1 
Radishes (winter)__.____-- 32 1595 tor 98\ |. Zato;4 months. 28 2) | eee 
ilanulloaing oe oe a Fg oe 32 | 90 to 95 | 2 to 8 weeks________ 28. 4 
ubaloa gases se ae te ey 32 | 95 to 98 | 2 to 4 months- __--__-. 29. 5 
Nalsifys GES EEO SE BLE 9S). TOn9 Sw. 2 GOR Sasha Ae 28. 4 
Dpolmach< wy aye pers we gs 32 | 90 to 95 | 10 to 14 days___-___-_ 30. 3 
Squashes: 
ROLUL OGIO 0) pa sane Senet rem Ay DIRE nm 40 to 50 | 85 to-95 | 2°to 3 weeks_.-.-_--|s_-__=_ 
VWatterie aero, 50 to 55 | 70 to 75 | 4 to 6 months______~_ 29. 3 
DVRCCEDOLAUOCS aa ae es 2'50' to: 50) ,| -SOkto 85 4/2 Uae Goya MU Sas = He 28..5 
Tomatoes: 
Ripe sapere wc tbe AN 10150) 1380 to 850) 71to 10 daysusso222 30. 4 
Matumeroreen s\2 22a 2 255 to 70 | 80 to 85-| 3 to 5 weeks____-__z 30. 4 
AUT NI SP pearsall < beew ty 32 | 95 to 98 | 4. to 5 months... ___= 30. 5 


1 These figures are based on previously published work by Wright (1 16) and 
are subject to revision whenever further investigation makes this necessary. 
2 See text. 3 Flesh. 4 Rind. 


ASPARAGUS 
(Temperature, 32° F.; relative humidity, 85 to 90 percent) 


Fresh asparagus is not usually stored except temporarily when the 
market is overstocked. Experiments have shown, however, that it can 
be kept successfully for 3 to 4 weeks at a temperature of 32° F. At this 
temperature, growth of the stalks, which takes place at higher tem- 
peratures, is practically nil. The original tenderness of fresh aspara- 
gus, which at ordinary room temperatures is lost soon after cutting 
owing to the formation of woody tissue, is preserved at the lower tem- 
perature. Furthermore, the sugar content, to which asparagus owes 
some of its flavor and which after cutting rapidly diminishes at higher 
temperatures, remains practically the same as when the asparagus 
is cut, if it is put in storage at this temperature immediately after 
cutting. Therefore, the sooner asparagus is placed in proper storage 
after harvesting the better will be its condition when used. The loss 
of water while in storage or transit is likely to be great if the stalks 
are not stood on wet moss or other moist absorbent material placed in 
the bottoms of the crates. In storage, asparagus bunches are some- 
times set in water in shallow trays or pans. After a long haul to 
market, asparagus should not be expected to keep in storage for more 
than 3 to 6 days, although the preservation of quality will depend 


32 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


largely on how the product was handled before being received for 

storage. Asparagus that has been precooled immediately after being 

packed will arrive at the market in better condition than if not so 

treated. The principal decays of asparagus in storage are bacterial | 

soft rot and gray mold rot. 
(See 69.) 


BEANS 


(Temperature, 32° to 40° F.; relative humidity, 85 to 90 percent) 
GREEN, OR SNAP 


Green beans are usually stored for only short periods. When 
held at 32° F. they may be expected to keep 2 to 4 weeks provided they 
are in good condition and are placed in storage promptly. At 40° 
the storage period will be about a week shorter. The humidity should 
not be lower than 85 percent, to prevent wilting, and the hampers or 
other containers should be so stacked as to allow abundant air circu- 
lation. If the containers are packed close together the temperature 
may rise somewhat because of the heat given off by the commodity, » 
and more or less rapid decay may be expected. If the beans are stored 
too long, the pods may become moldy or slimy and stick or “nest” 
together. The principal kinds of decay favored by too high storage 
temperature or too long holding period are watery soft rot, slimy 
soft rot, rhizopus rot, and gray mold rot. 


LIMA 


Shelled lima beans are sometimes stored in quart baskets and, if 
fresh and sound when stored, can be expected to keep in good salable 
condition for about 15 days at 32° F. and about 4 days at 40°. If 
stored too long, the beans tend to fade to a light color and become 
sticky. Unshelled lima beans can be held satisfactorily for 2 to 4 
weeks at 32° and 10 days at 40°. 


BEETS 


(Temperature, 32° F. Relative humidity: topped, 95 to 98 percent; bunch, 
85 to 90 percent ) 


Late beets stored at 32° F. may be expected to keep 1 to 3 months 
under suitable storage conditions. Either cold storage or cool cellar 
storage is suitable provided the humidity is kept sufficiently high to 
prevent wilting. Cellar storages often have a higher average tem- 
perature range than is recommended, and under these conditions the 
period of successful storage will be comparatively shorter. ‘The tem- 
perature in such storage should not go above 45°. Beets are subject 
to wilting because of the rapid loss of water and should be kept where 
the humidity is sufficiently high to prevent excessive evaporation. 

Before going into storage, beets should be topped and well sorted 
to remove all diseased specimens and those showing mechanical injury, 
in order to prevent undue shrinkage because of storage decay. Beets 
may be stored in ventilated barrels or better in slat crates. Storage 
in large bulk should be avoided. 

Bunch beets may be stored at 32° F. for 10 days to 2 weeks. See dis- 
cussion of bunch carrots (p. 34), as the same conditions apply as for 
bunch beets. 


STORAGE ‘OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 33 


Broccoit (ITALIAN, OR SPROUTING) 
(Temperature, 32° to 35° F.; relative humidity, 90 to 95 percent) 


Italian, or sprouting, broccoli does not keep well in storage and is 
usually held for only very short periods. The best storage tempera- 
ture is 32° F.. If in good condition and stored with sufficient ventila- 
tion between the packages, broccoli should keep satisfactorily for a 
week or 10 days. Longer:storage is undesirable because the leaves 
are likely to discolor and the buds may drop off (76). 


BrussELs SPROUTS 
(Temperature, 32° to 35° F.; relative humidity, 90 to 95 percent) 


Brussels sprouts are stored only occasionally but when stored they 
require the same conditions as broccoli. They should be held in 
small containers to prevent yellowing and the development of mold. 
The maximum storage period is probably not longer than 3 to 4 
weeks, 

CABBAGE 


(Temperature, 32° F.; relative humidity, 90 to 95 percent) 


A large percentage of the late crop of cabbage is stored and sold 
during the winter and early spring, or until the new crop from the 
Southern States appears on the market. If stored under proper con- 
ditions, cabbage should keep for 3 to 4 months. The longest keeping 
varieties belong to the Danish Ballhead class. Cabbage is most suc- 
cessfully held in common storage in the Northern States, where a fairly 
uniform inside temperature from 32° to 35° F. can be maintained. 
Many such storage houses are to be found, principally in New York, 
Pennsylvania, Michigan, and Wisconsin. Cabbage in quantity 
usually is not held in cold storage because its value does not justify 
the expense of handling. 

Storehouses should be insulated sufficiently to prevent freezing, for 
although shght freezing does no harm, hard freezing is likely to 
cause considerable loss. More ventilating capacity than is required 
for most other vegetables should be provided to carry away the ex- 
cessive moisture given off by the active respiration of this product 
and to obtain the maximum advantage of the cold night air during 
mild weather. Cabbage wilts quickly if held under too dry storage 
conditions; hence the humidity should be high enough to keep the 
leaves fresh and turgid. Bin storage is common, the bins usually be- 
ing 4 to 5 feet wide and 10 to even 20 feet long and about 5 feet deep. 
They are best separated by tight board partitions and ventilating 
slat floors. Tiers of bins may be built.as high as it is convenient to 
elevate the cabbage, and while the bins are being filled ample air space 
between the tiers of bins should be allowed for ventilation. The 
use of slat shelves with the heads piled one or two layers deep is con- 
sidered the best method, but it is too expensive when large quantities 
are to be stored. 

Cabbage should be handled carefully from the field to the storage. 
Before it is stored, the roots and all loose leaves should be trimmed 
away and the damaged and misshapen heads should be culled. On 

82996549 5 


34 CIRCULAR 278, U. S.. DEPARTMENT OF AGRICULTURE 


being removed from storage, the heads should be trimmed again to 
remove loose and damaged leaves. 

Early cabbage, especially southern-grown, should not be expected to 
keep over 3 to 6 weeks even at 32° F. — 

The most common decay found in stored cabbage is slimy soft rot. 


CARROTS 


(Temperature, 32° F. Relative humidity: topped, 95 to 98 percent; bunch, 
85 to 90 percent) 


Carrots are stored in fairly large quantities during the winter. 
The marketing period for stored carrots extends to late winter or 
early spring. They are usually held in common storage in those sec- 
tions where the storage temperature can ordinarily be held sufficiently 
low. In the larger markets stored carrots must be sold in competition 
with fresh stock, which is being shipped practically the year around 
from either southern or western producing sections. Carrots are 
sometimes held in cold storage, although the prices obtained for them 
do not usually justify this kind of treatment. 

It is generally considered that very light freezing causes practically 

no injury, but carrots should be protected from severe freezing and 
are best stored at a temperature of 32° F. They are subject to wilting 
or drying out if the humidity is not fairly high; for this reason they 
are more easily kept in a well-ventilated cellar or bank storage than in 
an above-ground storage. The relative humidity should be main- 
tained at 95 percent or slightly higher. 

Before being placed in storage, carrots should be topped and all 
misshapen orinjured specimens sorted out. The latter are especially 
objectionable, because their presence in a storage lot favors the devel- 
opment of two serious diseases of stored carrots, namely, watery soft 
rot and bacterial soft rot.. Carrots are best kept in slat crates or ven- 
tilated barrels, and provision should be made for air circulation be- 
tween the containers. Under good conditions they should keep 4 
to 5 months. | 

Bunch carrots may be stored at 32° F. for 10 days to 2 weeks, and 
the tops will still retain a fresh appearance if they are not crowded 
in storage. If cold storage is not available bunch carrots from dis- 
tant production sections may be packed in crushed ice and should keep 
at least a week. If it is desired to carry over such carrots for only 
a day or two, icing may not be necessary. Under these conditions, 
however, the crates or containers should be opened and the contents 
loosened so as to allow air to circulate through; otherwise, heating 
will take place, and the foliage will soon become yellow or discolored ; 
this, of course, 1s undesirable. 

Foliage will discolor sooner if kept warm and wet than if kept 
cool and dry. 

CAULIFLOWER 


(Temperature, 32° F.; relative humidity, 85 to 90 percent) 


Cauliflower is not usually kept in cold storage; however, an over- 
supply on the market can be stored for a short time to await more 
favorable conditions. If in good condition cauliflower can frequently 
be held satisfactorily for 2 to 3 weeks at 82° F. Successful storage 
depends not only on preventing decay but also on retarding the ma- 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 30 
turing of the head, or curd. Overmaturity is marked by a browning 
of the otherwise white curd and the development of a ricey appear- 
ance. ‘The leaves also become yellowish and may drop off. During 
storage or transportation the crates should be stacked with the flower 
heads down to protect the curds from discoloration by dirt and mois- 
ture. When it is desirable to hold cauliflower temporarily out of cold 
storage, packing it in crushed ice will aid in keeping it fresh. Freez- 
ing causes a grayish-brown discoloration and softening of the curd, 
accompanied by a water-soaked condition. 


CELERIAC 


(Temperature, 32° F.; relative humidity, 95 to 98 percent) 


Celeriac should be stored under the same conditions as those for 
topped carrots and should keep 3 to 4 months. 


CELERY 


(Temperature, 31° to 32° F.; relative humidity, 90 to 95 percent) 


Much of the late celery grown in the Northern States, notably New 
York and Michigan, is put in cold storage to supply the market up 
to the period in late winter when the competition of new celery from 
California and the South renders further holding unprofitable. Con- 
siderable celery from the South and West is also put in cold storage 
toward the end of the shipping season and field to supply the market 
during the summer, or until supplies of early, northern-grown stock 
appear on the market. 

Celery is a rather perishable commodity and under unsuitable 
storage conditions may suffer severely from watery soft rot. This dis- 
ease originates in the field and is caused by a fungus that is able to 
develop to some extent even at temperatures of 34° to 36° F. For 
this reason celery intended for storage should be as free as possible 
from infection. If held in rooms where a uniformly low temperature 
can be maintained, it should keep for 2 to 4 months. It is best stored 
at a temperature of 31° or 82° with a relative humidity high enough 
to prevent wilting (90 to 95 percent), and with sufficient air circulation 
to keep the temperatures at the top and bottom of the room as nearly 
equal as possible. Considerable heat is given off by celery because of 
active respiration, and the air at the top of a storage room is likely 
to be 8° to 4° warmer than at the bottom unless special precautions are 
taken to avoid such a condition. Air circulation can be maintained 
around the crates by using 1- by 2- or 2- by 2-inch dunnage strips 
between the crates, which should be stacked so as not to touch at the 
sides. If wall or ceiling refrigerating coils are used, fans should be 
located at such positions as will insure adequate air circulation. If 
the storage period is long, celery will keep better in small crates than 
inlarge ones. | 

Celery should not be piled more than four crates high in storage; 
otherwise there is danger of overheating even with stock that is in 
prime condition. If it is piled five to eight crates high, as is sometimes 
done, the room should be watched carefully to see that overheating 
does not occur. 


36 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 

Some growth takes place in celery while in storage. The central 
stalks lengthen considerably, obtaining their food at the expense of 
the outer stalks and the roots. Blanching of the stalks also takes 
place in most varieties that are put into storage. Some celery is 
trimmed and washed as it comes from storage, but probably the 
larger part is moved out in the original crates in which it was 


received. 
(See 95.) * 


Corn (GREEN) 


(Temperature, 31° to 32° F.; relative humidity, 85 to 90 percent) 


Green corn is seldom stored, although there are occasions during 
the southern shipping season when it may be desirable to put an excess 
supply of this commodity temporarily in cold storage; however, 
storage for more than a few days will result in serious deterioration. 
The sugar content, which so largely determines quality in this product 
and which rapidly decreases at ordinary temperatures, is reduced 
very little if the corn is quickly cooled and kept at a relatively low 
temperature. In order to keep this loss of sugar to a minimum and 
preserve the flavor, corn in the husks as it comes from the field for 
consumption in the fresh state should be cooled as quickly as possible. 
This is sometimes done by submerging it in tanks of ice water immedi- 
ately after removal from the field to reduce the temperature to as near 
32° EF. as possible. 

Corn should not be handled in bulk because of its tendency to heat 
but should be put in baskets or crates, which allow air circulation and 
the more rapid removal of field heat and heat produced by respira- 
tion. This commodity as it usually arrives on the market should not 
be expected to keep in marketable condition in cold storage for more 
than 4 to 8 days. 


CUCUMBERS 


(Temperature, 45° to 50° F.; relative humidity, 85 to 95 percent) 


Cucumbers are usually held in storage for only short periods and 
cannot be expected to keep satisfactorily for much over 10 to 14 days. 
The most favorable storage temperature range seems to be between 
45° and 50° F., with a relative humidity of 85 to 95 percent. When 
cucumbers are held at 45° or below for longer periods than recom- 
mended dark-colored watery ateas, which are an indication of low- 
temperature injury, appear. ‘These areas soon become infected, and 
mold growth develops. If the cucumbers are held at 50° little or no 
break-down develops, but they tend to ripen, the color changing from 
green to yellow, and there may be some shriveling and surface pitting. * 

Waxing is practiced commercially, usually with a paraffin-carnauba 
emulsion ‘containing approximately 7 percent solids. This treatment 
reduces weight loss ‘and improves appearance. 


* THOMPSON, H. C. CELERY STORAGE. | Refrig. Res. Found.] TRRF Fact File 
Sheet. 2 pp. October 1947. [Processed.] 

* Morris, L. L., and MANN, L. K. STORAGE OF CUCUMBERS. Refrig. Res. Found. 
Fact File Sheet. 1p. December 1948. [Processed.] 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 37 


EGGPLANTS 


(Temperature, 45° to 50° F.; relative humidity, 85 to 90 percent) 


Eggplants cannot be expected to keep satisfactorily in storage for 
more than about 10 days. 


ENDIVE, OR ESCAROLE 


(Temperature, 32° F.; relative humidity, 90 to 95 perecent) 


Endive, or escarole, is a leafy vegetable and under commercial con- 
ditions is not adapted to long storage. Even at 32° F., which is con- 
sidered to be the best storage temperature, it cannot be expected to 
keep satisfactorily for more than 2 or 3 weeks. The storage require- 
ments for endive are practically the same as for lettuce. Like lettuce 
it should keep somewhat longer than the period just mentioned if it 
is stored with cracked ice in or around the packages. ‘The relative 
humidity in rooms where endive is held should be kept at 90 to 95 
percent in order to prevent wilting. 

A certain amount of desirable blanching usually occurs in endive 
that is held in storage. 


Garuic (Dry) 


(Temperature, 32° F.; relative humidity, 70 to 75 percent) 


Garlic is best stored under the temperature and humidity condi- 
tions required for onions. If in good condition and well-cured when 
stored, this product should keep at 32° F. for 6 to 8 months. In 
Cahfornia, where considerable garlic is grown, it is frequently put 
in common storage, where it may be held for 3 to 4 months or some- 
times longer if the building can be kept cool, dry, and well-ventilated. 
Garlic is stored in loose mesh bags, which are piled two layers deep in 
stacks separated by air spaces. It is essential that garlic be well 
cured in the field before going into storage. 


HorsERADISH 
(Temperature, 32° F.; relative humidity, 95 to 98 percent) 


Horseradish should keep satisfactorily for 10 to 12 months if stored 
under the conditions recommended for topped carrots. 


JERUSALEM-ARTICHOKES 
(Temperature, 31° to 32° F.; relative humidity, 90 to 95 percent) 


Jerusalem-artichokes, if held in storage at a temperature of 31° to 
32° F. in a relative humidity of 90 to 95 percent, may be expected 
to remain in good condition 2 to 5 months. At low humidities they 
shrivel badly and are more likely to decay than if kept in a moist 
atmosphere. They are sometimes stored in barrels or in paper-lined 
bags, in which they keep longer, with less wilting and decay, than if 
left in open containérs. 


38 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


KoHLRABI 
(Temperature, 32° F.; relative humidity. 95 to 98 percent) 


Kohlrabi should keep 2 to 4 weeks if stored under the conditions 
recommended for topped carrots. 


LEEKs (GREEN) 
(Temperature, 32° F.; relative humidity, 85 to 90 percent) 


Green leeks are crated and stored under conditions similar to those 
suitable for celery. If properly handled, they should keep satis- 
factorily for 1 to 8 months in storage. 


LETTUCE 


(Temperature, 32° F.; relative humidity. 90 to 95 percent) 


Lettuce is sometimes put in cold storage when there is a surplus 
on the market, or in certain sections when the fall crop is threatened 
by approaching cold weather. If in good condition when stored, it 
can be expected to keep for 2 or 3 weeks. When lettuce is held tem- 
porarily out of cold storage crushed ice will greatly aid in keeping it — 
fresh. Even when it is held in cold storage, ice tends to keep this prod- 
uct fresher by preventing drying or wilting. 

One of the most troublesome diseases of lettuce in transit and stor- 
age is tipburn of the type that develops in the interior of the head. 
This injury appears in the field, but in the later stages of the market- 
ing process it is frequently followed by a slimy bacterial decay, which 
may result in serious damage. Frequent inspection of stored lots is 
desirable. 


MELONS 


Cold storage is used very little for most kinds of melons. When it is 
used, the storing is generally done at the terminal markets to avoid 
temporary adverse market conditions. 


WATERMELONS 
(Temperature, 36° to 40° F.; relative humidity, 75 to 85 percent) 


The ordinary commercial varieties of watermelons cannot usually be 
expected to keep in storage for more than 2 or 3 weeks. Experimental 
lots have been held at temperatures of 32° F. and did not develop 
decay as rapidly as at the recommended temperatures, but there was a 
tendency for the melons to become pitted or dented and to take on an 
objectionable flavor after 1 week. 


MUSKMELONS (CANTALOUPS) 
(Temperature, 32° to 34° F.; relative humidity, 75 to 78 percent) 


The common commercial varieties of cantaloups can be expected to 
keep about 1 week in cold storage at the recommended temperatures 
after arrival on the market and, under favorable conditions, for a 
few days longer but not as long as 2 weeks. The riper the melons are 
on arrival, the shorter the storage period should be. When they are 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 39 


held too long in storage, decay develops so rapidly on removal that 
they soon become practically worthless. 


(See 70.) 
HONEY DEW AND HONEY BALL MELONS 


(Temperature, 86° to 38° F.; relative humidity, 75 to 85 percent) 


Honey Dew and Honey Ball melons can usually be kept a little 
longer in storage than cantaloups. At 36° to 38° F. or slightly lower, 
such melons can be expected to keep for 2 to 4 weeks and still reach 
the consumer in good condition. If held for a longer time at tem- 
peratures below 36°, low-temperature break-down marked by a watery 
discoloration of the rind, followed by fungus decay, will probably 
result ; at 36° to 88°, decay, which will seriously discolor the rind, may 
occur. All blemishes show more plainly on these melons because of 
their light-colored, smooth surfaces. 


CASABA AND PERSIAN MELONS 


(Temperature, 36° to 40° F.; relative humidity, 75 to 85 percent) 


Casaba and Persian melons are relatively good keepers. They will 
remain in good condition in storage for at least 4 weeks and have been 
reported to keep as long as 6 weeks. 


MusHrooms (CULTIVATED) 


(Temperature, 32° to 385° F.; relative humidity, 80 to 85 percent) 


Mushrooms do not keep well in storage and are therefore stored 
only temporarily for periods of 2 to 8 days or slightly longer. ‘They 
are easily injured by freezing. 


OKRA 
(Temperature, 50° F.; relative humidity, 85 to 95 percent) 


Okra, if in good condition, can be kept satisfactorily in storage for 
a maximum of 2 weeks at a temperature of 50° F. A relative humidity 
of 85 to 95 percent is desirable to prevent wilting. At temperatures 
below 50° okra is subject to chilling injury, which is manifested by 
surface discoloration, pitting, and decay. ° 


ONIONS AND ONION SETS 


(Temperature, 32° F.; relative humidity, 70 to 75 percent) 


Onions are held in either common or cold storage. In the northern 
onion-growing States, strongly flavored varieties, mostly of the globe 
type, are generally held in common or dry storage. ‘The principal 
northern onion-producing States have a sufficiently low average winter 
temperature so that onions can be successfully held in common storage 
there during the winter months. About one-fourth of the onion crop 
of these States, however, is put in cold storage for consumption late 
in the spring. About the first of March is considered as late as onions 
should be held in common storage, because after this time there is 
danger of sprouting. The mild, or Bermuda, types, such as those pro- 


*Morris, L. L., and Mann, L. K. STORAGE OF OKRA. Refrig. Res. Found. 
Fact File Sheet. 1p. December 1948. [Processed.] 


40 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


duced in Washington, southern California, Texas, and other States 
where the climate is not suitable for common stor age, are usually con- 
sumed shortly after being harvested. These onions can be, and limited 
quantities are, held in cold storage, but usually for much shorter 
periods than the globe varieties because of their peorer keeping 
qualities. The Spanish, or Valencia, type of onions grown in this 
country are often stored and, if well- matur ed, are considered capable 
of storage for practically as long as the globe type. 

A comparatively low relative. humidity (70 to 75 percent) is very 
desirable for the successful storage of onions. At higher humidities, 
in which many other vegetables keep best in storage, onions are dis- 
posed to root growth and decay. The commonest form of the latter 
is gray mold rot occurring at the top of the bulb, whence its name 
“neck rot” (98). The fungus causing it can develop to some extent 
even at 32° F.; hence onions intended for storage should be carefully 
sorted over to remove all diseased bulbs. A uniform temperature of 
32° is found to be sufficiently low to keep onions dormant and reason- 
ably free from decay provided they are in good sound condition and 
well-cured when stored. 

Onions are not perceptibly injured by shght freezing if allowed to 
thaw out slowly and without rough handling. In cold storage they 
are usually held in bags of 50 or 100 pounds each, which are best piled 
in pairs laid crosswise in stacks five or six sacks high. The stacks 
should be set a few inches off the floor on 2- by 4- inch strips and the 
individual stacks separated by a few inches of space to allow for air 
circulation. When kept in common storage, onions are best stored 
in slat field crates holding about 1 bushel, rather than in bags. Be- 
fore being placed in storage onions should be well dried or cured in 
the field for a period of 4 to 6 weeks, and all decayed specimens or 
those showing thick, or “bottle,” necks should be sorted out. 

Onion sets are usually held in common storage. They require 
nearly the same conditions as large onions and are best stored in 
shallow slat-bottom crates or trays not over 4 inches deep and about 
5 by 5 feet in some districts or 2 by 3 feet in others. The corner posts 
of the crates should project about an inch above the side pieces in 
order to prevent the crates from resting tightly on each other when 
stacked and to allow air circulation between them. Because of their 
size, onion sets tend to pack closely in the crates; hence it is essential 
to allow as much air circulation as possible and to maintain a com- 
paratively low humidity. If good stock is provided and is held under 
proper storage conditions, it should keep 6 to 8 months. 

(See 57.) 

PAaRSNIPS 


(Temperature, 32° F.; relative humidity, 90 to 95 percent) 


Parsnips have nearly the same storage requirements as topped car- 
rots and should keep for 2 to 4months. They are not injured by shght 
freezing while in storage but should be protected from hard freezing 
aud should be very carefully handled while in a frozen condition. 
Parsnips dry out readily in storage; hence it is essential that the 
humidity of the storage place be kept relatively high. Parsnips are 
sometimes stored in sand or clean soil to prevent wilting, but they 
will keep in good condition when held in barrels or crates if the 
proper humidity is maintained. 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 41 


Pras (GREEN) 


(Temperature, 32° F.; relative humidity, 85 to 90 percent) 


Storage conditions required for green peas are somewhat different 
from those for beans. Green peas tend to lose part of their sugar con- 
tent, on which much of their flavor depends, unless they are promptly 
cooled to near 32° F. shortly after being picked. They cannot be 
expected to keep in salable condition for more than 1 to 2 weeks unless 
packed in crushed ice, in which condition the storage period may be 
extended perhaps a week. Peas keep better unshelled than shelled. 


. PEPPERS 


CHILI PEPPERS (DRY) 
(Temperature, see text; relative humidity, 70 to 75 percent) 


Chili peppers are usually picked when ripe and then dried and 
allowed to equalize in moisture content in covered piles. Water is 
usually added to the peppers after drying, and as a result they become 
less brittle. They are then packed tightly by tamping into sacks 
holding 200 to 800 pounds and stored in nonrefrigerated warehouses 
for 6 to 9 months. | 

The temperature of the warehouses depends to some extent on their 
censtruction and the way in which they are managed but chiefly on 
the outside temperature. In southern California, where a large part 
of the commercial crop of Chili peppers is produced, the outside 
temperature ranges from 50° to 80° F. during the usual storage 
period. 

The moisture content of Chil peppers when stored is generally low 
enough (10 to 15 percent) to prevent mold growth; the chief storage 
trouble is insect infestation. Sometimes manufacturers of Chili pep- 
per products hold part of their supply of the raw material in cold 
storage, but they prefer to grind the peppers as soon as possible and 
store them in the manufactured form in airtight containers. 


SWEET PEPPERS 


(Temperature, 32° F.; relative humidity, 85 to 90 percent) 


Sweet, bell, or bullnose peppers, if in good condition may readily 
be kept fresh in storage for a month to 6 weeks at a temperature of 
32° F. A relative humidity of 85 to 90 percent is desirable to prevent 
shrinkage (46). 


POTATOES 


EARLY CROP 


(Temperature, see text; relative humidity, 85 to 90 percent) 


Early and intermediate potatoes are not often stored. However, 
if need arises in the production area (usually southern) this crop 
can be kept satisfactorily in common unrefrigerated storage through 
the summer in the locality where it is grown (44). The potatoes 
should, of course, be sorted over carefully before storage to remove all 


42 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


those decayed and seriously bruised and cracked. Investigations in 
Oklahoma (75) indicated that refrigerated storage at 50° F. after a 
curing period is best: for the early crop. Some evidence has been 
obtained, however, that, if early potatoes are to be used for chipping 
purposes, a storage temperature of 60° to 70° is better than one of 50°. 
The relative humidity should be the same as that for the late crop. 
If refrigeration is not available, an underground storage cellar or 
cave is next best if the temperature can be held below 70° and a fairly 
moist atmosphere maintained to avoid excessive shrinkage. (See also 
49.) 

It almost goes without saying that under the conditions suggested 
the storage period would never be so long as is possible with late-crop 
potatoes. 


LATE CROP 
(Temperature, 38° to 50° F.; relative humidity, 85 to 90 percent) 


Potatoes are stored either in cold or common storage, but the greater 
part of the crop that is stored is held in common storage (20). Like 
most other vegetables that can be held for relatively long periods in 
common storage, potatoes can be successfully kept through the fall 
and winter months only in those sections where a sufficiently cold 
winter climate prevails. In either cold or common storage a temper- 
ature of 40° F’. is as low as table or seed stock need be kept during the 
first few months after harvest. At temperatures below this, there is 
a tendency for potatoes to become undesirably sweet. However, if 
sweetening occurs, a few days’ exposure to ordinary living-room tem- 
perature will partly restore the natural flavor. Potatoes should always 
be kept in the dark to prevent greening. At 40° potatoes will remain 
dormant 5 to 8 months after harvest, depending on the variety. If it 
is desired to keep them longer than this, as is often the case with seed 
stock, the temperature may be lowered to 38°, at which they should 
remain dormant indefinitely. A storage temperature as low as 32° is 
not only unnecessary but detrimental. | 

Investigations have indicated that potatoes stored at 50° to 60° F, 
have better texture, color, and flavor when cooked or made into chips 
than the same stock stored at lower temperatures (64, 120), although 
the higher temperatures are not suitable for long-time storage. When 
potatoes are stored at these higher temperatures, sprouting will occur 
more quickly. A limited amount of sprouting does not injure potatoes 
tor food purposes, but it makes the stock difficult to market because 
usually only dormant potatoes are wanted. If sprouting has started it 
can be checked by lowering the storage temperature. 

Potatoes that have been kept at 40° F. for a long time are seldom 
suitable for processing, such as chip making, french frying, or dehy- 
drating, without first being conditioned to reduce the quantity of sugar 
that has accumulated. This is accomplished by holding the potatoes at 
70° to 80° until trial cooking tests show that they have recovered sufh- 
ciently for use. The length of the conditioning period will depend on 
the variety and the amount of sugar that has accumulated; usually, 
however, this will be 1 to 3 weeks (123). 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 43 


The relative humidity of a potato-storage house should be 85 to 90 
percent, to prevent undue shrinkage through loss of water. In cold 
storage, potatoes are generally kept in sacks holding 50 or 100 
pounds net ; in common storage they are usually placed in bins holding 
from 150 to as much as 1,000 bushels or more. In Maine and northern 
New York, where the average temperature is sufficiently low, the 
large-bin storage is used with success, but in the milder climate of 
States in the latitude of Pennsylvania potatoes should not be stored 
in such large units. Potatoes are readily injured by even slight freez- 
ing, which takes place at about 29° F. or slightly below ; hence common- 
storage buildings should be sufficiently insulated to prevent freezing. 
Insulation will also prevent the condensation of moisture on the walls 
and ceilings and the consequent undesirable wetting of stored stock, 
which favors the development of decay. Common storages should be 
provided with sufficient ventilation to take advantage of the cool night 
air in mild weather; this will aid in removing excess moisture, 
accumulating especially soon after potatoes are stored in the fall, 
and will maintain a lower average temperature. Ventilators should 
never be opened, however, when the outside temperature is higher 
than that inside the storage house. In addition to damage to potatoes, 
condensed moisture caused by improper ventilation or inadequate 
insulation may also cause serious impairment to the building struc- 
ture (19). 

Potatoes intended for storage should be handled carefully to avoid 
bruises and cuts; otherwise they are lhkely to be damaged by various 
forms of decay before the end of the storage period. 


PUMPKINS AND SQUASHES 


(Pumpkins and winter squashes: temperature, 50° to 55° F.; relative humidity, 
70 to 75 percent. Summer squashes: temperature, 40° to 50°; relative 
humidity, 85 to 95 percent ) 


In general most varieties of pumpkins will not keep in storage as 
long as the usual storage varieties of squash. Such varieties as Con- 
necticut Field and Cushaw are relatively poor keepers and cannot be 
expected to hold in good condition more than 2 to 3 months. Varieties 
like Large Cheese and Table Queen will keep 3 to 6 months. 

Hard-shell squashes, such as the Hubbards, can be successfully stored 
for 6 months or longer. 

All stock should be well-matured, carefully handled, and free from 
Injury or decay when put in storage. The best storage temperature 
appears to be 50° to 55° F. with a relatively low humidity of about 
70 to 75 percent. Investigations have shown that a preliminary - 
curing at 80° to 85° for about 2 weeks is of benefit in ripening immature 
specimens and in healing mechanical injuries incident to harvesting. 

Summer (yellow crookneck) squashes keep best at a temperature of 
about 40° to 50° F. and a relative humidity of 85 to 95 percent. Under 
these conditions, they may show some increase in the yellow color, 
pitting, and some wilting at the necks after approximately 3 weeks; 
but they are not likely to be seriously damaged by decay.’ 


“Morris, L. L., and MANN, L. K. STORAGE OF SUMMER SQUASH. Refrig. Res. 
Found. Fact File Sheet. 1p. December 1948. [Processed. ] 


44 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


RapisHES (WINTER) 
(Temperature, 32° F.; relative humidity, 95 to 98 percent) 


Winter radishes require the same storage conditions as topped 
carrots and should keep in good condition for 2 to 4 months. 


RHUBARB 
(Temperature, 32° F.; relative humidity, 90 to 95 percent) 


Rhubarb stalks, if fresh and in good condition, may be stored for 
2 to 3 weeks. The bunches should be packed in crates, which are 
stacked to allow ample air circulation on all sides; otherwise there 
is danger of heating and mold growth. 


RUTABAGAS 
(Temperature, 32° F.; relative humidity, 95 to 98 percent) 


Rutabagas require the same storage conditions as topped carrots 
and should keep satisfactorily under such conditions for 2 to 4 months. 
Probably most of the rutabagas that now appear on the market have 
been waxed. This treatment improves the appearance and is reported 
to prevent wilting and loss of weight. 


SALSIFY 
(Temperature, 32° F.; relative humidity, 95 to 98 percent) 


Salsify has the same storage requirements as topped carrots. The 
roots are not injured by slight freezing but should be carefully 
handled while frozen. Under the conditions specified, they should 
keep for 2 to 4 months. 


SPINACH 
(Temperature, 32° F.; relative humidity, 90 to 95 percent) 


Spinach is usually stored for only short periods. It should keep 
fairly well for a week or two after being cut. If crushed ice is used 
in the packages, this period can be extended somewhat. 


SQUASHES 
See Pumpkins and Squashes. 


SWEETPOTATOES 
(Temperature, 50° to 55° F.; relative humidity, 80 to 85 percent) 


The requirements for the successful storage of sweetpotatoes differ 
from those recommended for most other vegetable crops. When 
freshly dug sweetpotatoes are to be stored for any length of time they 
should be given a preliminary curing treatment to permit the healing 
of all wounds or abrasions incident to harvesting and handling, in 
order to prevent the entrance of decay organisms. 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 45 


The curing and storing are done in the same house so that the 
potatoes do not have to be moved after the curing treatment. When 
commercial lots are handled, the storage house is generally of special 
construction with sufficient insulation to maintain a uniform tem- 
perature and some means of ventilation that will insure the desired 
humidity. Provision should be made for heating the building during 
the curing process and for holding the proper storage temperature 
afterward. The curing process ordinarily takes about 10 days, during 
which the house is kept at a temperature of 85° F’., with a relative 
humidity of 85 percent. After the curing period the storage tem- 
perature is allowed to drop to about 55°, with a humidity of 80 to 
85 percent. Short periods of a few hours at temperatures somewhat 
lower than 50° need not cause alarm, but prolonged periods of low 
temperature should be avoided because of the danger from certain 
types of decay, which are more likely to develop at temperatures below 
the range given (45,61). Under the recommended conditions, prop- 
erly cured stock should keep satisfactorily for 4 to 6 months. 

Only well-matured stock that is practically free from mechanical 
injury or decay should be used for storage. Sweetpotatoes are usu- 
ally stored in slat crates of about a bushel capacity or in bushel baskets. 
Shallow bins are sometimes used. The roots should be handled as 
little as possible during storage. : 


(See 40.) 
TOMATOES 


(Temperature: ripe, 40° to 50° F.; mature green, 55° to 70°. Relative humidity, 
80 to 85 percent) 


Ripe tomatoes are held in storage only temporarily and, except for 
short storage, should not be stored at temperatures lower than 40° F. 
At 40° to 50°, if not already soft ripe, they will keep in good condition 
for a week to 10 days; at temperatures lower than this they sometimes 
show a tendency to break down. 

Green tomatoes are best kept at a temperature not lower than 55° F. 
At this temperature ripening progresses slowly but satisfactorily, and 
mature green tomatoes can be kept for 3 to 5 weeks before becoming 
overripe. At temperatures much below 55° green tomatoes do not 
ripen well and if kept there more than about 8 days and then moved 
to a warmer place usually do not ripen satisfactorily; on the other 
hand, if they are to be kept for less than 8 days they can be held at 
40° or even somewhat lower. If fairly rapid ripening is desired, 
temperatures from 60° to 70° should be used. At 70° or slightly above, 
ripening is accelerated, but so also is the development of decay, which 
will be found difficult to control. At about 80° coloring will be un- 
even since the development of the red pigment is inhibited at this 
temperature or above. The relative humidity of tomato-storage or 
tomato-ripening rooms should be from 80 to 85 percent. 

(See 101, 119, 121.) 


TURNIPS 


(Temperature, 32° F.; relative humidity, 95 to 98 percent) 


Turnips require the same storage conditions as topped carrots. 
They can be expected to keep for 4 to 5 months. 


46 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


FROZEN VEGETABLES 


For treatment of frozen vegetables see page 29. 


CUT FLOWERS, FLORISTS’ GREENS, RHIZOMES, TUBERS, » 
CORMS, AND BULBS 


By T. M. WHITEMAN, associate horticulturist 


The recommended temperature, relative humidity, and approximate 
length of storage period for the commercial storage of cut flowers, 
florists’ greens, rhizomes, tubers, corms, and bulbs are given in ta- 
ble 8. Detailed descriptions of these requirements are given in the 
text. 


TaBiE 8.—Recommended temperature, relative humidity, and approxi- 
mate length of storage period for cut flowers, florists’ greens, 
rhizomes, tubers, corms, and bulbs 


Commodity nae! Approximate length 
ine of storage period 
Cut flowers: ! an Oe 

Babysbréathisee tlt eee er a ee ee 40 | 3 to 4 days 
Bouvardiaesweete22: fas 5 222. er eee 40 | 7 to 10 days 
Butterflybush, orange-eye______-___-------- 40 | 3 to 4 days 
Calendulat(potsmanicold) Sa. sean 40 | 3 to 6 days 
Callavcommon-and golden - oo _ 49348 Serer 40 | 10 days. 
Candytuits 22226 oe ae 2. ee an eee 40 | 3 to 6 days 
Carnation: 

Rulitbloomsar 24 eee tea ici eer 33-40 | 7 to 10 days. 

BGS 5 Ee sia ss Ded Ea ep ea a yg 33-40 | 10 to 15 days 
Ching-asten 5 e266. oe eee ape ee ge 40 | 7 to 10 days 
Chrysanthemums a eee eee 35 | 2 weeks. 
Clarkia 4 itiect te Pais TOV ee Se 40 | 3 to 6 days 
Columbine =e * eee ee ee 40 | 3 to 4 days 
Wornilowermties. 20) 6. = ae ae ee 40 | 3 to 6 days 
CrOCUSH ars Soe eee atte ge ee Be ee 33-36 | 1 to 2 weeks 
1 E50 FE vest ih as pr ps A eh Si lagi ts Soya Bc Ne A 40 | 7 to 10 days 
Daisy; Pnglishi0 SBAse_ Gyo TOT aaaR: © 40 | 3 to 6 days 
Delphinium: 

Hardyalsrikcs piece te oe Gea eee ee aie 40 Do. 

Annuallarks pure eo feces pee 40 |-3 to 4 days 
WE VETIC Wire See Loe eRe ene ae Se eee ae 40 | 3 to 6 days 
Horget=me-not; trues2_2 235. & fi ant eee 40 | 3 to 4 days 
Foxglove, common and common white- - - --- 40 Do. 
TeGSIAUE: 2 O82). ee eke eee ee 33-36 | 1 to 2 weeks 
Gaillardia, common perennial -_--_---------- 40 | 3 to 6 days. 
Gardeniaes 20 4 ee ee ae 45-50 | 1 week. 
Gladiolus erat Un nae fe ee 35 | 2 weeks. 
He@athe “bal elie 3) teen eee ne ee eee 40 | 7 to 10 days 
Fy acing i ges ey SS oe Poe oe ee oe 33-36 | 1 to 2 weeks 
Tris Cte hes = Soe Se ee eee ee 33 | 1 week. 
Laceflower, blue! 222220 sO 22) Ue Beane eee 40 | 3 to 6 days. 
Lily: 

HaStere oF ee Sees A coe ea ine ere 35 | 1 month 

Gold ban deen 2 es See oo ea 35 Do. 

Realy Hatin Ok ee eT RAC 35 Do. 

Speciostim:. 0p hs ea ire hh eee ee See 35 Do. 
Lily-of-the-valleys_22 5122 See See 40 | 1 week. 
Tiypine 26! oS ink ee es Ee ee ee 40 | 3 to 6 days. 


See footnotes at end of table, p. 47. 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 47 


TasiE 8.—Recommended temperature, relative humidity, and approwi- 
mate length of storage period for cut flowers, florists’ greens, 
rhizomes, tubers, corms, and bulbs—Continued 


Tem- : 
3 Approximate length 
Commodity Fie of storage period 
ure 
Cut flowers: !—Continued 
Narcissus: Stk 
1D) aio Wears oe eS ree pees aioe Serre 33-36 | 1 to 2 weeks. 
Ra perwiiter aes eee ts woe the ets 33-36 Do. 
Ohi eats Se eee ce ae 45-50 | 1 week. 
Peony, Chinese and common: 
Mroatm oS! oe ae a cere a Re oo 35 | 3 to 4 weeks. 
MooscsOMd se sears, see e) bie ep 35 | 10 to 14 days. 
iBhloxarardenes 2A" e sere e see ee cme 40 | 3 to 4 days. 
TPL TOSIS mb se on Se ae fi pe ae ep ae 50 Do. 
GI FOSE Gas ee eee ee ee eS ee 40 Do. 
EROS Cee eles Peo Re ek oye re 35-40 | 1 week. 
Snapdraconscommoneen s'est ee eee = 40 | 3 to 6 days. 
Snowaropiie cise ae see ee hee nee a Eee 33-36 | 1 to 2 weeks. 
STU Ls pa eae alt ee lel ng Sie 33-36 Do. 
SeaticenGeaslavenGen) at a. spe oe oer 35-40 | 3 to 6 weeks 
SUE hao) ors ag Ses SS teen gi Dali Ne ieD ies R EAR ae 40 | 3 to 4 days 
SLOCescOmMm one atta ee bet Ee 40 | 3 to 6 days 
trawl wie steer ei Teeter ween CL EE ier 35-40 | 3 to 6 weeks. 
DSN CX EUBI]OL OLE IRE ot SR yo Slag ae a a eee 40 | 3 to 4 days 
Rt eee ee See ee ee ye ee 33-36 | 1 to 2 weeks 
WiGlet sweeter: seme oe eee Skee 33-36 | 3 to 6 days 
Florists’ greens:! . 
ASparacus ern anGysnoilax © osc ee vite oo! 40-45 | 7 to 10 days. 
Ferns: 
Commonrwoodfernes 2) as Se a ee 32-45 | 2 weeks to 4 months. 
Dale cereal ash) Te Cee AA ee ee oe 32-45 O. 
Crea xe hs ae rin a banger ener ete a) 32-45 Do. 
Croundpine (uycopodium)ia 4 = 32-45 Do. 
FLOW WACHL EL ODGCG) a oes ees 32 | 1 month 
PUGS oer ye ees eee es ER Ne SR ee 32-40 | 3 to 6 weeks. 
Eevuecthoesdroopingwim A wtl iene. pani 32-40 Do. 
Mountain-laurel tat 88 26 salve oper expec 32-40 Do. 
Rhizomes, tubers, corms, and bulbs: ? 
Bevonidecuoena s.r er em ea sak ate 45 | 4 to 6 months. 
Caladiuinr spotted es wks Pies AN eed 45 Do. 
Calla, \common and golden=_/___=- 22-22 35 | 6 to 8 months. 
Canna srs. sxe sha A area ae eel a 40-45 | 6 to 7 months. 
Cr OCU Se ene erat ao ye Sa AE ee epee 55-60 | 4 to 6 months. 
ID FEN UL chy las eet eat att opera Marsares hat 40-45 | 6 to 7 months. 
ee siat yas Ce? ke IPE SER ko) SPUD REEL 55-60 | 4 to 6 months. 
GladioluswA2e _ ashy. lew robes sy ek | Fy os 40-50 | 7 to 8 months. 
1B Lyf: 176)'b 29 0 a ea ene BORD Ie oe Avnet SOR ee nen 55-60 | 4 to 6 months. 
ily: 
ASU CTY Cancer tare eee ee gre aie ers eee 32-35 | 6 to 8 months. 
Regal sc: SOR Gaal tie). “RE SE a OE 32-35 Do. 
inily-of-the=valleyee) ae ty fe hey sete 25-28 | 12 to 17 months. 
Narcissus: 
IDE 06 (6 bl hey pl enero nh one orga Sneath a a aE a 55-60 | 4 to 6 months. 
IR ApeLWteka seem en ere eT nee S42 75-80 | 5 to 7 months. 
Orn ER CD ST LE TA TRC aS UNE 40-45 | 6 to 7 months. 
SNOWOTOp sats rere oe 8 A jel i pe Lest ye) 55-60 | 4 to 6 months. 
SK (SUD gee Sea aa ae Sa eae Meee ar oe ea ee 55-60 Do. 
UST aa es a nae em 45 Do. 
UCT OSC me eee ARISTA TRIAL ANI BAD PD. 40-45 | 6 to 8 months. 
Mp ase Fo ts were be ita 2 wd 110d 5 50-55 | 4 to 6 months. 


1 Approximately 80 percent relative humidity. 
2 Approximately 75 percent relative humidity; for individual species see text. 


4S CIRCULAR 278, U. 8. DEPARTMENT OF AGRICULTURE 


The average freezing points of some flower petals, foliage, bulbs, 
corms, etc., are given in the following tabulation: 


RBPIOWO NOOCDNOCMORUUMNTIONNWO ww 


Foliage: Temperature (°F.)! | Petals—Continued Temperature (° F.) 
Amarylliss 2.2 See ee 31.0 Daisy;.Shastas+-2-s = eee 29. 
Asparagusslernis ta nem e 24. 2 Delphinium (hardy lark- 
ANSDICISCrAa” 2 os 22 Seen 24. 9 SIU) es Sly ye a ae 26. 
Caladium, spotted__...=-__ 30. 6 Gardenmianes.- "0-5 yee aes 28. 
Carnation: 24 2a 27. 4 Gladiohistes. <n eee 28 
Chrysanthemum = ees 29. 6 Heather ees. ee 28 
Columbines a= Sa een 29. 1 Hemerocalliste:: ~ = = sueuee 30 
Daisy; Shasta-s 3s eee = 30. 0 Fiyvacintheeeeeres .> Tree 28. 
Delphinium (hardy lark- irise Japaneseses.. 22 a0 ee 30. 

ON 5) ngs ee Pa i te CE age 9. 2 ihilyacbastert.. soe as 2 
Dracaena: ==) ears © ee 28. 0 Narcissus (daffodil) __-______ 30. 
Ferm’) dagger 2. Bi. yaa 23. 6 Orchid (Catileya) = a2 30. 
Gag oOhUses sac aetence ates 26. 8 Peony, commons. sass. == 29. 
Hiemerocallishs2 22) = = 30. 0 Poimsettiaece ss). ae 29. 
Holly Gilexiopaca) ta 26. 3 Ranunculuss2222 2) Soe 28. 
Tris: Rose; hy bridttea > asses ee 30. 

Datel ares sates 29. 0 qCultp = = 26 =. ae 28. 

Germann. as tee ee 27. 6 Violét, Sweet. 2. ae ae 28. 

Japanese. 22 Pie ce 28. 7) Bulbs, corms, ete.: 
lhily..astere!-=— 3 S24 eee 29. 2 Calltea se ae a ee 27 
PanG@anus= se hae tee ee 30. 4 Dahliaa. = Sees 28 
Rubber, variegated________ 30. 3 Gladiolus2222 34 = 2 Uren 26 
Violet sweets2= 4-2 pe. 22 = 27. 4 Eliy S.C lint nee ee ee 28 
VAN CQ anajOne = eee Ree 28. 6 Dnily, Revalss2sSsss = 2am 20. 

Petals: Narcissus: 

IATICIMIONG se eee oe 28. 1 Daliodils toss 26. 1 
Carnation. Sse t seen 28. 4 Paperwhites.——2 =e 28.9 
Chrysamthentume = — 28. 4 Dulas Fe. Pea ee eer ee 25. 4 
Columbinese= 2 == =5 eae Slew 


1 Many of these figures are based on previously published work by Wright (116) and are subject to revision 
whenever further investigation makes this necessary. 


Cut FLoweErs 


The best prestorage handling of cut flowers not only includes the 
selection of species and varieties known to be good keepers but also 
provides for the use of well-grown blooms only. Well-grown flowers 
are those that have developed under near-optimum water, nutrient, 
temperature, and hght conditions and that have received no mechani- 
cal or other damage while on the plant. Flowers of such quality are 
“firm” and are said to have “substance,” terms which probably could 
be justified on the basis of total solid matter, as contrasted with soft 
blooms that usually develop under conditions of forced growth. Well- 
grown flowers do not have hard, or excessively woody, stems for the 
species; hard stems make water uptake difficult. 

In preparing cut flowers for storage the proper cutting stage, dis- 
cussed under individual species, is of prime importance. Most cut 
flowers are benefited by a freshening period of a few hours (in water) 
at about 50° F. immediately after being cut, unless intended for dry 
storage. Bunching is usually done before storage. Flowers should 
be bunched and tied firmly but not too tightly or in too large bunches. 
Wrapping the bunches with waxed paper with an opening at the top 
prevents tangling of the blooms during the handling of such species 
as delphinium, columbine, and lilies-of-the-valley; of course, in the 
case of roses wrapping of the bunches is customary. If intended for 
storage in water, cut flowers should be stored at the proper tempera- 
ture soon after they have been freshened at 50°. 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 49 


One of the chief factors influencing the storage life of cut flowers 
istemperature. There is no known treatment that is nearly so satisfac- 
tory in extending the life of cut flowers as is storage at comparatively 
low temperatures. If the proper temperature is used, inherent quality 
should not deteriorate appreciably during a short storage period. The 
temperatures given here are recommended to insure a reasonably long 
life for flowers after their removal from storage. Cut flowers for 
which 45° to 50° F. is recommended, when stored at a much lower 
temperature, may not keep well after removal from storage. ‘Those 
for which a storage temperature of 33° is recommended will naturally 
mature more rapidly if stored at any higher temperature. If held at 
33° they develop slowly. Often the changes in storage are not ap- 
parent, but the longer the blooms are stored at 33° the shorter will be 
their life when later they are used for decorations. On the other 
hand, if short-lived blooms that keep best when held at 40° are stored 
at 33° they show the effect of the previous adverse conditions after 
they are placed at ordinary room temperature (70°-75°). 

In case a variety of flowers needs to be stored and only one storage 
room is available it is suggested that a temperature of 40° be used: 
The length of time that the blooms are to be stored should always be 
considered. For a number of items table 8 shows the recommended 
storage temperature as well as the range in the approximate length 
of the storage period. 

The effect of humidity on the storage of cut flowers deserves more 
consideration than it has had in the past. A relative humidity of 
90 to 95 percent is probably too high in that 1t may promote mold 
growth, especially if the blooms are somewhat crowded. At low 
relative humidity, 70 to 75 percent, the petals of certain types of 
flowers may tend to dry to an undesirable degree. A relative humidity 
of 80 percent is therefore recommended. It has been noted with cer- 
tain types of blooms, such as gladiolus, that floret development was 
faster in a room with a temperature of 36° F. and a high relative 
humidity than in a room at the same temperature and a lower 
humidity. 

In constructing a storage room for cut flowers one of the most im- 
portant considerations is air circulation. To get good air circulation 
the vases or buckets of flowers should be set on racks arranged so 
that air can pass through and back of them. Forced but gentle air 
movement should be provided, but the blooms should not be in a direct 
draft. The containers should be spaced so that the blooms are not 
crowded. 

Cut flowers in storage should usually be kept with the stems in 
water. They should not be crowded in the container because of the 
danger of mechanical injury and the decay that may result because 
of insufficient ventilation. In most cases care should be taken not 
to spill water on the blooms as these usually discolor rather readily. 
Experiments in which French marigolds and carnations were stored 
out of water for comparatively long periods have been reported 
(60,61). Peonies are known to keep fairly satisfactorily when stored 
out of water. 

Damage to several kinds of cut flowers when stored in the same 
room or even in the same building with apples, oranges, pears, and 
other fruits should be emphasized. Such damage is evidenced by. 
premature withering or the rapid aging of the blooms and is thought 


50 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


to be caused by the ethylene gas given off by the ripening fruit. Roses, 
carnations, snapdragons, stocks, and daffodils have been shown to 
be affected in this way by the emanations from ripening fruit (47, 
68). Since ethylene gas is used at times to defoliate rose bushes and 
is known to cause premature dropping of foliage from cut sprays ot 
holly, it may also affect such greens as mountain-laurel and huckle- 
berry. Greens therefore should not be stored near the fruits 
mentioned. 

Many kinds of pigmented flowers fade, discolor, or develop an off- 
color appearance during storage, especially under dark storage con- 
ditions. The effectiveness of artificial light in retarding this color 
change has been demonstrated experimentally (59). 

Chrysanthemums (Chrysanthemum hortorwm) and giladioluses 
(Gladiolus) may be stored at 35° F. for 2 weeks and usually remain 
very salable. Longer storage is possible, though frequently not prac- 
ticable. These flowers are usually tied in bunches of 12 and 25, respec- 
tively, but are not wrapped when placed in storage. Pompons, 
however, are sold in bunches of various sizes. 

As a rule the proper cutting stage for the large-flowering types 
of chrysanthemums is just after the green color in the center of the 
flower has disappeared. No general rule can be given for pompons, 
since several varieties, especially some of those in the anemone group, 
need more development on the plant than the single varieties. 

Gladioluses should be cut when three or four of the lower buds are 
partly opened. However, for distant shipments before storage one 
or two of the lowest florets should show color but should be in the 
tight-bud stage. Gladiolus spikes should always be shipped and 
stored in a vertical position (105). | 

Easter lily (Lilium longiflorum), Speciosum lily (L. speciosum), 
Regal lily (LZ. regale), goldband lily (ZL. awratum), common peony 
-(Paeonia officinalis), and Chinese peony (P albiflora) can be held 
at 35° F. for comparatively long periods, although 30 days is usually 
the maximum storage period. ‘The lies in this group should be cut 
for storage when the corolla is about one-half opened or just before 
the tips begin to reflex. Liles forced at relatively high temperatures 
should be kept at a temperature of about 50° for a preliminary period 
of about 24 hours before being put at 35°. Peonies showing color in 
the tight-bud stage may be stored at 35° for 3 to 4 weeks, but they 
seldom give satisfaction for decorative purposes without special han- 
dling; in the loose-bud stage they may be held satisfactorily for 10 
to 14 days. 

The cut blooms of common and golden callas (Zantedeschia aethi- 
opica and Z. elliottiana) may be stored when fresh for as long as 10 
days at 40° F. and will be in good condition upon removal. Calla 
blooms should be gathered just before the spathe shows signs of 
curling downward. Callas that have been subjected to hard forcing 
should be held for about 24 hours at a temperature of 50° previous 
to storage at 40°. When the blooms are gathered they should be 
pulled, not cut, otherwise the stems will split at the cut ends and curl 
after a few days in storage. The pulling method separates the stem 
from the rhizome, leaving no useless appendage. Although the 
blooms are sometimes removed from the plant by cutting, the stub 
left on the plant will probably rot and thus may serve as a place of 
entrance for disease organisms. Pulling, however, is much easier 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 51 


with potted callas than with those planted in beds or benches. Callas 
intended for storage should be tied near the end of the stem and also 
loosely tied below the blooms. They are usually put up in lots of 
1 dozen. 

Lupine (Lupinus), clarkia (Clarkia), common stocks (Matthiola 
incana), candytuft (lberis), delphinium, or hardy or perennial 
larkspur (Delphiniwm formosum and hybrids), cornflower (Cen- 
taurea cyanus), feverfew (Chrysanthemum parthenium), common 
snapdragon (Antirrhinwm majus), blue laceflower (TZ rachymene 
caerulea), English daisy (Bellis perennis), calendula, or pot marigold, 
(Calendula officinalis), and common perennial gaillardia (Gaillardia 
aristata) should not be held at temperatures lower than 40° F., and 
cannot be stored with good results for more than a 3- to 6-day period. 
Sweet violets are usually made up in bunches of 100, supported under- 
neath by a few galax leaves and wrapped with a light waxed paper, 
and should be stored at 33° to 36°. 

Columbine (Aguilegia), stevia (Piqueria trinervia), babysbreath 
(Gypsophila paniculata), annual larkspur, baby primrose (Primula 
jorbesiz), sweet pea (Lathyrus odoratus), true forget-me-not (Myo- 
sotis scorpioides), and orange-eye butterflybush (Buddleia davidii) 
all have flowers whose petals shed quickly, almost regardless of tem- 
perature. They cannot be held much lower than 40° F. or for longer 
than 3 to 4 days without impairing the keeping quality after removal. 
Forced Buddleia is sold by the dozen. ‘The others mentioned are 
bunched in lots of 25. It is not customary to wrap any of these for 
storage, but, as previously mentioned, certain kinds may be wrapped 
to prevent tangling with adjacent bunches. 

Orchids and gardenias (Gardenia augusta, G veitchi, and G. for- 
tunez) are not customarily stored for long periods. However, they 
may be kept in storage in good condition for about a week at a tem- 
perature of 45° to 50° F. They keep best when cut just after they have 
reached a salable condition. 

Carnations (Dianthus caryophyllus) cut in full bloom may be held 
at 33° to 40° F. for 7 to 10 days. If cut in the bud stage they may 
be stored at 33° to 40° for 10 to 15 days. Several varieties of carna- 
tions, although it is not known how many, will develop satisfactorily 
when cut in the bud stage, viz, when one or two petals have unfolded 
from the bud, and will last considerably longer than if cut at a more 
advanced stage of maturity (47). 

‘Roses for nearby markets should be cut in the loose-bud stage; if 
they are to be shipped to distant markets they should be cut in as 
tight a condition as is permissible with the variety in question. In 
the loose-bud stage they may be held at 35° to 40° F. for 1 week; if 
stored at lower temperatures the subsequent keeping quality is often 
impaired (59). These are often tied loosely in bunches of 25 buds 
and wrapped with parchment paper. 

Dahlias (Dahlia), China-asters (Callistephus chinensis), sweet 
bouvardia (Bowvardia humboltiz), and heath (Frica) may be held 7 to 
10 days at 40° F. They usually are handled in lots of 1 dozen, tied, but 
not wrapped. 

Statice, including bigleaf and notchleaf sea-lavender (Limoniwm 
latifolium and L. sinwatum, respectively), and strawflower (Helich- 
rysum bracteatum) may be kept at 35° to 40° F. for 3 to 6 weeks. 
They may be dried and will retain their original color and shape; 


52 CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


strawflowers are usually dried instead of being stored to retain their 
freshness. 

Common foxglove (Digitalis purpurea), common white foxglove 
(D. purpurea var. alba), and garden phlox (Phlox paniculata) are 
not ae satisfactory for storage but may be held for 8 or 4 days at 
40° F. 

The various forced irises, especially the so-called Dutch irises, may 
be held for 1 week at 33° F. (106). 

Cut poinsettias (Z’uphorbia pulcherrima) sold during the Christmas 
season usually need not be stored for the few days between their 
arrival at the wholesale house and the day of sale. If holding is neces- 
sary, storage at about 50° F. is recommended. Any change of en- 
vironment, such as improper storage, will increase the apparently 
inherent tendency of poinsettias to shed their foliage. They should 
be cut when showing sufficient color to be salable. Searing the cut 
ends with boiling water is a practice usually followed to prevent undue 
loss of sap before or during storage. 

Cut lilies-of-the-valley (Convallaria majalis) are kept satisfac- 
torily at 40° F. and may be held for 1 week at this temperature; if 
they are kept longer the lower bells often become watery in appearance 
(104). The proper cutting stage is just after the terminal bell has 
lost its deep-green color. It should be of a yellow-green appearance, 
the lower 3 or 4 bells at this time being well opened. They are usually 
tied with foliage in bunches of 25 and are better wrapped loosely in 
heavy waxed paper, leaving the tops and bottoms of the bunches open. 

Hyacinth (Hyacinthus orientalis), tulip (107), narcissus (daffodil 
and paperwhite), freesias (Freesia refracta var. alba), squills, snow- 
drops (Galanthus nivalis), and crocuses can usually be held satisfac- 
torily for 1 to 2 weeks at 33° to 36° F. 

Spikes, such as snapdragons, should be cut just after the lower five or 
six flowers have fully opened; umbels, such as blue lacefiower, should 
be cut just after they develop to a salable condition; flowers formed in 
heads usually should be cut after the outermost petals are fully de- 
veloped and just before stamens appear in the center of typically 
single heads or after the center has become closed with petals in 
double sorts; corymbs, such as candytuft, are usually best when cut 
after three-fourths of the lower flowers are opened; flowers described 
as thyrses, or corymbose cymes, such as the hlac, should be cut when 
about two-thirds of the determinate branches are developed; cymose 
clusters, such as babysbreath, should be cut after a few of the terminal 
flowers have developed. 

Experiments have been reported on the use of a plastic coating on 
gardenia blooms (87). With further tests on different types of 
blooms, the usé of such materials may become more extensive as a 
supplement for cold storage or during shipment. 


FLorists’ GREENS 


Fern asparagus (Asparagus plumosus) and smilax asparagus (A. 
asparagoides) are usually shipped in crates. The turn-over of these 
greens is rapid and shipments are arranged so that storage for longer 
than 3 or 4 days is usually unnecessary, but they may be kept in the 
case for 7 to 10 days at 40° to 45° F. The sprays of A. plumosus are 
tied in bunches of various sizes. Smilax is packed in the crates in 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 53 


various lengths or “strings.” The commercial popularity of A. 
sprengeri has declined so much that it is produced and used chiefly by 
small retail growers. 

Drooping leucothoe (Leucothoe catesbaez), huckleberry, and moun- 
tain-laurel (alma latifolia) sprays may be held in good condition at 
32° to 40° F. for 3 to 6 weeks. ‘They are usually tied in bunches and 
kept standing in water. 

Galax (Galax aphylla), groundpine (Lycopodium), dagger ferns, 
common woodferns, and various other species including Vryopteris 
intermedia, packed in crates, may be held at 82° to 45° F’., depending 
on the length of time it is desired to keep them. At 32° they may be 
expected to keep 1 to 4 months, whereas at 45° they may be kept for 
but 2 to 3 weeks in good condition. They are not put in water, but if 
stored loose they are kept moist by occasional sprinklings. If they are 
held in wooden cases or packed in such a way as to prevent excessive 
drying out by direct air currents, no sprinkling is necessary. Galax 
is usually tied in bunches of 25; the ferns are tied in lots of 50; but 
both are customarily sold by the hundred. ‘Those who make a busi- 
ness of gathering the greens in this group sometimes store them in 
coldframes, covering them with about 6 inches of some material such 
as sphagnum moss or leaves, and shading or using sash as the weather 
demands. Handled in this way, shipments can be made at any time. 
Others who might be termed “brokers” have many employees gather- 
ing galax and ferns and store their entire holdings in cold-storage 
warehouses; this is the method commonly employed for storage, and 
it usually gives satisfaction. 

Holly (lex opaca) sprays or wreaths can usually be held satisfac- 
torily for approximately 1 month at a temperature of 32° F. and a 
relative humidity of about 80 percent. ‘These should be left in the 
packing cases to prevent the circulation of air immediately around 
them (122). 

A plastic coating has been used on such greens as fern asparagus, 
dagger fern, and cut branches of several evergreen trees (35, 87). . 


RHIZOMES, TUBERS, AND CorMsS 


Lily-of-the-valley pips or crowns may be held in cold-storage ware- 
houses at a temperature range of 25° to 28° F. (104) up to 17 months, 
although after about 12 months increasing deterioration in quality 
becomes noticeable. The pips are tied in bundles of 25 with string or 
willow ties and are packed with sphagnum moss in wooden cases of 
various sizes containing 250 to 2,500 pips. 

Taro (Colocasia esculenta), spotted caladium (C. neoguineensis), 
and tuber begonias (Begonia tuberhybrida) may be held at 45° F. 
and a relative humidity of 75 to 80 percent for 4 to 6 months. They 
should be packed in dry sawdust, sand, or peat. 

Canna (Canna indica), dahlia (Dahlia) (3), and peony (Paeonia) 
may be stored at 40° to 45° F. over a period of 6 to 7 months with a 
relative humidity of 70 to 80 percent. They are usually packed in 
dry soil, sand, sawdust, or peat moss. 

Gladiolus (mostly hybrids) may be stored at 40° to 50° F. with a 
relative humidity of 70 to 75 percent for 7 to 8 months. A tempera- 
ture of 40° will hold these corms dormant during the normal storage 
season, whereas at 50° sprouting will occur after 4 to 6 months’ stor- 


54 CIKCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


age. ‘They should be stored dry in shallow trays with ample ventila- 
tion but only after a curing period of 3 to 6 weeks in an open or well- 
ventilated shed. 

Common and golden callas (Zantedeschia aethiopica and Z. elli- 
ottiana) stored at 35° F. and a relative humidity of 70 to 75 percent in 
dry soil, sand, or peat will keep for many months if it is necessary to 
store them. The normal rest period in the forcing cycle is during 
May, June, and July. Unless water is withheld during this period 
complete dormancy will not exist. Therefore, storage in a dry con- 
dition is essential, with sufficient humidity to prevent undue shriveling. 


BuLss 


Most, varieties of daffodils (Narcissus), hyacinths, freesias, squills 
(Scilla), snowdrops (Galanthus nivalis), Crocus, and similar bulbs 
may be held at 55° to 60° F. for 4to 6 months. However, these as well 
as tulips (Tudipa) are usually not stored, because better results are 
obtained by planting them as soon as they are received. Paperwhite 
narcissus (108) may be stored at 75° to 80° for 5 to 7 months from 
digging time. | 

Tulips are usually not stored in any packing material, although 
the use of such material may be advantageous for long storage. Tem- 
peratures of 36° to 38° F., except for short storage, are considered 
rather low for tulips shipped from the West in which the buds are 
already formed. The best storage temperature for shipped tulips is 
between 50° and 55°, at which they may be kept for 4 to 6 months. 
Tuberoses (Polianthes tuberosa) stored at a temperature of 40° to 45° 
may be expected to keep satisfactorily for 6 to 8 months. 

Bulbs of the genus Lilium, of which there are approximately 50 
species, are like most other bulbs in giving best results if planted when 
received. However, as Easter lihes are forced during the entire year, 
it becomes necessary to provide cold-storage conditions suitable for 
satisfactory holding until they are wanted for planting. The bulbs 
should be packed in boxes in thoroughly dry soil if rooting and desic- 
cation are to be prevented. A temperature of 32° to 35° F., with a 
relative humidity of 75 to 80 percent, is recommended; under these 
conditions they may be held for 6 to 8 months. 

The procedure of storing certain bulbs for a time previous to plant- 
ing at a temperature that will subsequently hasten flowering is in con- 
sidetable use commercially. Bulbs so handled are said to be “pre- 
cooled” (7, 31, 32). 


LIPERATORE-€lieD 


(1) ALLEN, FE. W. 
1932. POSSIBILITIES AND LIMITATIONS OF COLD STORAGE FOR STONE FRUITS. 
Ice and Refrig. 82: 424. 
and PENTzER, W. T. 
1936. STUDIES ON THE EFFECT OF HUMIDITY IN THE COLD STORAGE OF FRUITS. 
Amer. Soc. Hort. Sci. Proc. (1935) 33: 215-228, illus. 
(3) ALLEN, R. C. 
1938. TEMPERATURE AND HUMIDITY REQUIREMENTS FOR THE STORAGE OF 
DAHLIA RooTS. Amer. Soc. Hort. Sci. Proce. (1987) 35: 770-773. 
(4) Bain, H. F., BERGMAN, H. F., and Witcox, R. B. 
1942. HARVESTING AND HANDLING CULTIVATED CRANBERRIES. U. S. Dept. 
Agr. Farmers’ Bul. 1882, 24 pp., illus. 


(2) 


(6) 


(18) 


(19) 


(20) 
(21) 


(22) 


(23) 


(24) 


(25) 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 955 


BARGER, W. R. 
1933. EXPERIMENTS WITH CALIFORNIA DATES IN STORAGE. Date Growers’ 
Inst. Rpt. 10: 3-5. ; 
———— PENTzER, W. T., and FISHER, C. K. 
1948. LOW TEMPERATURE STORAGE RETAINS QUALITY OF DRIED FRUIT. Food 
Indus. 20: F1-F4 (20: [887]—[340] ), illus. 
Biaauw, A. H., VERSLUYS, M. C., LUYTEN, I., and others. 
1925-26. THE RESULTS OF THE TEMPERATURE TREATMENT IN SUMMER FOR 
DARWIN TULIPS. K. Akad. van Wetensch. te Amsterdam, Afd. 
Natuurk. Proe., Sect. Sci. 28: 717-731, illus., 1925; 29: [113]- 
126, [199]-220, illus., 1926. 
Brooks, C., Coontry, J. S., and FisHemr, D. F. 
1930. DISEASES OF APPLES IN STORAGE. U.S. Dept. Agr. Farmers’ Bul. 
1160, 20 pp., illus. (Revised.) 
and HARLEY, C. P. 
1934. SOFT SCALD AND SOGGY BREAK-DOWN OF APPLES. Jour. Agr. Res. 49: 
55-69, illus. 
and McCotiocg, L. P. 
1936. SOME STORAGE DISEASES OF GRAPEFRUIT. Jour. Agr. Res. 52: 319— 
351, illus. 
and McCottocHu, lL. P. 
1937. SOME EFFECTS OF STORAGE CONDITIONS UPON LEMONS. Jour. Agr. 
Res. 55: 795-809, illus. 
Camp, A. F., GappuM, L. W., and STAHL, A. L. 
1933. COLD STORAGE STUDIES ON CITRUS FRUITS. Fla. Agr. Expt. Sta. 
Ann. Rpt. 1932-33 : 104-109. 
CARRICK, D. B. 
1930. SOME COLD STORAGE AND FREEZING STUDIES ON THE FRUIT OF THE 
VINIFERA GRAPE, N. Y. (Cornell) Agr. Expt. Sta. Mem. 181, 
37 pp., illus. ; 
CooLey, J. S., and CRENSHAW, J. H. 
1931. CONTROL OF BOTRYTIS ROT OF PEARS WITH CHEMICALLY TREATED WRAP- 
PERS. U.S. Dept. Agr. Cir. 177, 10 pp., illus. 
CoRDNER, H. B. 
1943. EXPERIMENTS WITH IRISH POTATOES: TIME OF PLANTING; SEED 
SOURCES ; VARIETIES } IRRIGATION ; FERTILIZERS ; TIME OF HARVEST; 
AND STORAGE OF THE SPRING CROP. Okla. Agr. Expt. Sta. Tech. Bul. 
T-18, 27 pp., illus. 
DENNY, F. E. 
1924, EFFECT OF ETHYLENE UPON THE RESPIRATION OF LEMONS. Bot. Gaz. 
77: 322-329, illus. 
DIEHL, H. C. 
1940. QUICK FREEZING FRUITS [AND] QUICK FREEZING VEGETABLES, Amer. 
_ Soe. Refrig. Engin., Refrig. Data Book 2: 19-30, illus. 
MAGNESS, J. R., Gross, C. R., and Bonney, V. B. 
1930. THE FROZEN-PACK METHOD OF PRESERVING BERRIES IN THE PACIFIC 
NORTHWEST. U.S. Dept. Agr. Tech. Bul. 148, 38 pp., illus. 
Epear, A. D. 
1938. STUDIES OF POTATO STORAGE HOUSES IN MAINE. U.S. Dept. Agr. Tech. 
Bul. 615, 47 pp., illus.: 


1947. POTATO STORAGE. U.S. Dept. Agr. Farmers’ Bul. 1986, 38 pp., illus. 
FISHER, D. F. : 
1942. HANDLING APPLES FROM TREE TO TABLE. U.S. Dept. Agr. Cir. 659, 
39 pp., illus. 
and Coo.Ley, J. S. 
1947. APPLE SCALD AND ITS conTROL. U. S. Dept. Agr. Farmers’ Bul. 
1380, 9 pp., illus. (Revised.) 
FRANKLIN, H. J. 
1940. CRANBERRY GROWING IN MASSACHUSETTS. Mass. Agr. Expt. Sta. Bul. 
371, 44 pp., illus. 
FRIEND, W. H., and Bacu, W. J. 
1932, STORAGE EXPERIMENTS WITH TEXAS CITRUS FRUIT. Tex. Agr. Expt. 
Sta. Bul. 446, 40 pp., illus. 
FRuIT DISPATCH COMPANY, EQUIPMENT DEPARTMENT. 
1942, BANANA RIPENING MANUAL. Fruit Dispatch Co., Equipment Dept. 
Cir. 14, 20 pp., illus. 


56 


(26) 
(27) 
(28) 


(29) 


(41) 
(42) 
(43) 
(44) 
(49) 


(46) 


CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


GERHARDT, F., and ENGLISH, H. 
1945. RIPENING OF THE ITALIAN PRUNE AS RELATED TO MATURITY AND STOR- 
AGE. Amer. Soc. Hort. Sci. Proc. 46: 205-209, illus. 
and HZELL, B. D. 
1941. PHYSIOLOGICAL INVESTIGATIONS ON FALL AND WINTER PEARS IN THE 
PACIFIC NORTHWEST. U.S. Dept. Agr. Tech. Bul. 759, 67 pp., illus. 
and RYALtL, A. L. 
1939. THE STORAGE OF SWEET CHERRIES AS INFLUENCED BY CARBON DIOXIDE 
AND VOLATILE FUNGICIDES. U.S. Dept. Agr. Tech. Bul. 631, 20 pp. 
Gore, H. C. 
1911. STUDIES ON FRUIT RESPIRATION. I. THE EFFECT OF TEMPERATURE ON 
THE RESPIRATION OF FRUITS. II. THE EFFECT OF PICKING ON THE 
RATE OF EVOLUTION OF CARBON DIOXIDE BY PEACHES. III. THE RATE 
OF ACCUMULATION OF HEAT IN THB RESPIRATION OF FRUIT UNDER 
ADIABATIC CONDITIONS. U.S. Bur. Chem. Bul. 142, 40 pp., illus. 
GREEN, W. P., HUKILL, W. V., and Ross, D. H. 
1941. CALORIMETRIC MEASUREMENTS OF THE HEAT OF RESPIRATION OF FRUITS 
AND VEGETABLES. U.S. Dept. Agr. Tech. Bul. 771, 22 pp., illus. 
GRIFFITHS, D. 
1936. SPEEDING UP FLOWERING IN THE DAFFODIL AND THE BULBOUS IRIS. U.S. 
Dept. Agr. Cir. 367, 18 pp., illus. 


1936. TULIPS. U.S. Dept. Agr. Cir. 372, 64 pp., illus. 

HALLER, M. H., and Harpine, P. L. 

1939. EFFECT OF STORAGE TEMPERATURES ON PEACHES. U. S. Dept. Agr. 
Tech. Bul. 680, 32 pp., illus. 

HARDING, P. L., Lutz, J. M., and Roskg, D. H. 

1932. THE RESPIRATION OF SOME FRUITS IN RELATION TO TEMPERATURE. 
Amer. Soe. Hort. Sei. Proc. (1981) 28: 583-589. 

HAMNER, C. L., GARTNER, J. B., and O’RouRKE, F. L. 

1948. A NON-TOXIC PLASTIC COATING TO IMPROVE THE KEEPING QUALITY OF 
CUT FOLIAGE. Mich. Agr. Expt. Sta. Quart. Bul. 30: 268-271, illus. 

HARDING, P. L. 

1935. PHYSIOLOGICAL BEHAVIOR OF GRIMES GOLDEN APPLES IN STORAGE. Iowa 
Agr. Expt. Sta. Res. Bul. 182: 318-852, illus. 

and HALLER, M. H. 

1933. THE INFLUENCE OF STORAGE TEMPERATURES ON THE DESSERT AND KEEP- 
ING QUALITY OF PEACHES. Amer. Soc. Hort. Sci. Proc. (1932) 
29 : 277-281, illus. 

and HAtuer, M. H. 

1935. PEACH STORAGE WITH SPECIAL REFERENCE TO BREAKDOWN. Amer. Soc. 
Hort. Sci. Proce. (1934) 32: 160-163. 

Harvey, HE. M. 

1946. CHANGES IN LEMONS DURING STORAGE AS AFFECTED BY AIR CIRCULATION 
AND VENTILATION. U.S. Dept. Agr. Tech. Bul. 908, 32 pp., illus. 

HASSELBRING, H., and Hawkins, L. A. 

1915. RESPIRATION EXPERIMENTS WITH SWEETPOTATOES. Jour. Agr. Res. 
5: 509-517. 

HAWKINS, L. A. 

1929. GOVERNING FACTORS IN TRANSPORTATION OF PERISHABLE COMMODITIES. 
Refrig. Engin. 18: 130-131, 135. 

HENDRICKSON, H. M. 

1940, REFRIGERATION OF LEMONS. Amer. Soc. Refrig. Engin., Refrig. Data 
Book 2: 174-178, illus. 

HUuKILL, W. V., and SmirH, BH. 

1946. COLD STORAGE FOR APPLES AND PEARS. U. S. Dept. Agr. Cir. 740, 
61 pp., illus. 

KImMBprouGcH, W. D. 

1944. STORAGE OF IRISH POTATOES IN THE LOWER SOUTH. La. Agr. Expt. 
Sta. La. Bul. 386, 17 pp. 
LAURITZEN, J. I. 
1931. SOME EFFECTS OF CHILLING TEMPERATURES ON SWEETPOTATOES. Jour. 
Agr. Res. 42 : 617-627, illus. 
and Wricut, R. C. 
1930. SOME CONDITIONS AFFECTING THE STORAGE OF PEPPERS, Jour, Agr, 
Res. 41 : 295-305, illus. 


STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 57 


LUMSDEN, D. V., Wricut, R. C., WHITEMAN, T. M., and ByrngEs, J. W. 
1940. FRUIT AND FLOWERS INCOMPATIBLE, Florists’ Exch, 95 (2):10, 1], 
illus. 
Lutz, J. M. 
1938. FACTORS INFLUENCING THE QUALITY OF AMERICAN GRAPES IN STORAGE. 
U. 8. Dept. Agr. Tech. Bul. 606, 27 pp., illus. 


1947. STORAGE OF SOUTHERN-GROWN POTATCES DURING THE SUMMER. Amer. 
Potato Jour. 24: 209-220, illus. 

CULPEPPER, C. W., Moon, H. H., and MEYERS, A. T. 

1933. OBTAINING OPTIMUM DESSERT AND CANNING QUALITIES FROM THE 
KIEFFER PEAR; SOME INFLUENTIAL FACTORS. Canning Age 14 (10): 
404—406, 414, 428, illus. 

and Simons, J. W. 

1948. STORAGE OF SWEETPOTATOES. U.S. Dept. Agr. Farmers’ Bul. 1442, 
50 pp., illus. (Revised. ) 

LYNCH, S. J., and STAHL, A. H. 

1939. STUDIES IN THE COLD STORAGE OF AVOCADOS. Fla. State Hort. Soc. 
Proc. 52: 79-81. 

Maeness, J. R., and BALLARD, W. S. 

1926. THE RESPIRATION OF RARTLETT PEARS. Jour. Agr. Res. 32: 801-832, 
illus. 

—— DIerH1, H.C., and ALiEn, F. W. 

1929. INVESTIGATIONS ON THE HANDLING OF BARTLETT PEARS FROM PACIFIC 
coASsT DisTRIcTS. U.S. Dept. Agr. Tech. Bul. 140, 28 pp., illus. 

DIEHL, M. C., HALLER, M. H., and others. 

1926. THE RIPENING, STORAGE, AND HANDLING OF APPLES. U.S. Dept. Agr. 

Dept. Bul. 1406, 64 pp., illus. 
and 'Taytor, G. F. 
1925. AN IMPROVED TYPE OF PRESSURE TESTER FOR THE DETERMINATION OF 
FRUIT MATURITY. U.S. Dept. Agr. Dept. Cir. 350, 8 pp., illus. 
MAGRUDER, R., WESTER, R. E., Jonres, H. A., and others. 
1941. STORAGE QUALITY OF THE PRINCIPAL AMERICAN VARIETIES OF ONIONS. 
U.S. Dept. Agr. Cir. 618, 48 pp. 

MILBRATH, J. A., HANSEN, E., and HarTMaAN, H. 

1940. DEFOLIATION OF ROSE PLANTS WITH ETHYLENE GAS. Science 91: 100. 

NEFF, M. S. 

1939. Coren AND KEEPING QUALITIES OF CUT FLOWERS. Bot. Gaz. 101: 501- 
U4, 


1939. PROBLEMS IN THE STORAGE OF CUT CARNATIONS. Plant Physiol. 
14 : 271-284, illus. 
and Loomis, W. E. 
1936. STORAGE OF FRENCH MARIGOLDS, Amer. Soc. Hort. Sci. Proce. (1935) 
33 : 683-685. 
OLNEY, A. J. 
1926. TEMPERATURE AND RESPiRATION OF RIPENING BANANAS. Bot. Gaz. 
82: 415-416, illus. 
OVERHOLSER, HL. L. 
1925. COLD STORAGE BEHAVIOR OF AvocaDos. Calif. Avocado Assoc. Ann. 
Rpt. 1924-25 : 32-40. 
PEACOCK, W. M., WRIGHT, R. C., WHITEMAN, T. M., and FULLER, BE. 
1931. DIFFERENCES IN THE COOKING QUALITY OF POTATOES DUE TO STORAGE 
TEMPERATURES. Potato Assoc. Amer. Proce. (1930) 17: 109-116. 
PENTZER, W. T. 
1931. THE COLD STORAGE OF GRAPES. Ice and Refrig. 81: 84. 
——— and AsBupry, C. E. 
1935. THE SODIUM BISULPHITE TREATMENT OF GRAPES TO RETARD MOLD 
GROWTH. Blue Anchor 12 (5) :6, 26-27. 
and AssBury, C. E. 
1937. OBSERVATIONS ON THE COLD STORAGE OF GRAPES AND OTHER FRUITS 
DURING THE 1936 SEASON. Ice and Refrig. 93: 193-194. 
Magness, J. R., Dient, H. C., and Hater, M. H. 
1932. INVESTIGATIONS ON HARVESTING AND HANDLING FALL AND WINTER PEARS, 
U.S. Dept. Agr. Tech. Bul. 290, 30 pp., illus. 


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CIRCULAR 278, U. S. DEPARTMENT OF AGRICULTURE 


PENTZER, W. T., PERRY, R. L., HANNA, G. C., and others. 

1936. PRECOOLING AND SHIPPING CALIFORNIA ASPARAGUS. Calif. Agr. Expt. 
Sta. Bul. 600, 45 pp.. illus. 

WIANT, J. S., and MacGiitivray, J. H. 

1940. MARKET QUALITY AND CONDITION OF CALIFORNIA CANTALOUPS AS IN- 
FLUENCED BY MATURITY, HANDLING, AND PRECOOLING. U. S. Dept. 
Agr. Tech. Bul. 730, 74 pp., illus. 

PLAGGE, H. H. 

1940. COLD STORAGE OF APPLES AND PEARS. Amer. Soc. Refrig. Engin., 
Refrig. Data Book 2: 163-170, illus. 

and Maney, T. J. 

1928. SOGGY BREAKDOWN OF APPLES AND ITS CONTROL BY STORAGE TEMPERATURE. 
Iowa Agr. Expt. Sta. Res. Bul. 115, pp. 63-118, illus. 

and Maney, T. J. 

1987. FACTORS INFLUENCING THE DEVELOPMENT OF SOGGY BREAK-DOWN IN 
APPLES. Jour. Agr. Res. 55: 739-768, illus. 

MANEY, T. J., and PICKETT, B. S. 

1935. FUNCTIONAL DISEASES OF THE APPLE IN STORAGE. Iowa Agr. Expt. 
ta. Bul. 329, 79 pp., illus. 

PLATENIUS, H. 

1939. WAX EMULSIONS FOR VEGETABLES. N. Y. (Cornell) Agr. Expt. Sta. 
Bul. 723, 43 pp., illus. 

JAMISON, F. S., and THoMpPSoON, H. C. 

1934. STUDIES ON COLD STORAGE OF VEGETABLES. N. Y. (Cornell) Agr. Expt. 
Sta. Bul. 602, 24 pp., illus. 

RAMSEY, G. B., and Link, G. K. K. 

1982. MARKET DISEASES OF FRUITS AND VEGETABLES : TOMATOES, PEPPERS, EGG- 
PLANTS. U.S. Dept. Agr. Mise. Pub. 121, 44 pp., illus. 

and WIANT, J. S. 

1944. MARKET DISEASES OF FRUITS AND VEGETABLES : BEETS, ENDIVE, ESCAROLE, 
GLOBE ARTICHOKES, LETTUCE, RHUBARB, SPINACH, SWISS CH ARD, and 
SWEETPOTATOES. U.S. Dept. Agr. Misc. Pub. 541, 40 pp., illus. 

WIANT, J. S., and LINK, G. K. K. 

1938. MARKET DISEASES OF FRUITS AND VEGETABLES: CRUCIFERS AND CUCUR- 
BITS. U.S. Dept. Agr. Mise. Pub. 292, 74 pp., illus. 

WIANT, J. S., and SmitH, M. A. 

1949. MARKET DIEASES OF FRUITS AND VEGETABLES: POTATOES. U. S. Dept. 
Agr. Mise. Pub. 98, 60 pp., illus. (Revised.) 

RASMUSSEN, E. J. 

1937. EFFECT OF DELAY IN STORAGE TEMPERATURE ON THE KEEPING QUALITIES 
OF APPLES. N. H. Agr. Expt. Sta. Tech. Bul. 67, 55 pp., illus. 

Rose, D. H., BRATLEY, C. O., and PENTZER, W. T. 

1939. MARKET DISEASES OF FRUITS AND VEGETABLES : GRAPES AND OTHER SMALL 
FRUITS. U.S. Dept. Agr. Misc. Pub. 340, 27 pp., illus. 

Brooks, C., BRATLEY, C. O., and WINSTON, J: R. 

1944. MARKET DISEASES OF FRUITS AND VEGETABLES: CITRUS AND OTHER SUB- 
TROPICAL FRUITS. U.S. Dept. Agr. Misc. Pub. 498, 57 pp., illus. 

Brooks, C., FISHER, D. F., and BRATLEY, C. O. 

1933. MARKET DISEASES OF FRUITS AND VEGETABLES : APPLES, PEARS, QUINCES. 
U. S. Dept. Agr. Mise. Pub. 168, 71 pp., illus. 

FISHER, D. F:, Brooxs, C., and BRATLEY, C. O. 

1937. MARKET DISEASES OF FRUITS AND VEGETABLES : PEACHES, PLUMS, CHER- 
RIES, AND OTHER STONE FRUITS. U.S. Dept. Agr. Misc. Pub. 228, 
27 pp., illus. 

SHAPOVALOY, M., and LINE, G. K. K. 

1926. CONTROL OF POTATO-TUBER DISEASES. U. S. Dept. Agr. Farmers’ 
Bul. 1367, 38 pp., illus. (Revised.) 

SHERWOOD, C. H., and HAMNER, C. L. 

1948. LENGTHENING THE LIFE OF CUT FLOWERS AND FLORAL GREENS BY THE 
USE OF PLASTIC COATINGS. Mich. Agr. Expt. Stat Quart. Bul. 30: 
272-276, illus. 

SIEBEL, J. E. 

1918. COMPEND OF MECHANICAL REFRIGERATION AND ENGINEERING ... Ed.9, 
571 pp., illus. Chicago. 

Srevers, A. F., and Barcer, W. R. 

1930. E¥YPERIMENTS ON THE PROCESSING AND STORING OF DEGLET NOOR DATES 
IN CALIFORNIA. U. S. Dept. Agr. Tech. Bul. 193, 24 pp., illus. 


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STORAGE OF FRUITS, VEGETABLES, AND FLORISTS’ STOCKS 59 


(90) Smock, R. M. 
1940. THE STORAGE OF APPLES. N. Y. Agr. Col. (Cornell) Ext. Bul. 440, 
38 pp., illus. 
and SouTHWICcK, F. W. 
1948. AIR PURIFICATION IN THE APPLE STORAGE. N. Y. (Cornell) Agr. Expt. 
Sta. Bul. 848, 52 pp., illus. 
(92) StTaut, A. L., and CAIN, J. C. 
1937. COLD STORAGE STUDIES OF FLORIDA CITRUS FRUITS. III, THE RELATION OF 
STORAGE ATMOSPHERE TO THE KEEPING QUALITY OF CITRUS FRUIT IN 
COLD STORAGE. Fla. Agr. Expt. Sta. Bul. 316, pp. 3-41, illus. 


(91) 


(93) and Camp, A. F. 
1936. COLD STORAGE STUDIES OF FLORIDA CITRUS FRUITS. I. EFFECT OF TEM- 
PERATURE AND MATURITY ON THE CHANGES IN COMPOSITION AND 
KEEPING QUALITY OF ORANGES AND GRAPEFRUIT IN COLD STORAGE. 
Fla. Agr. Expt. Sta. Bul. 303, 67 pp., illus. 
(94) and Firirip, W. M. 


1936. COLD STORAGE STUDIES OF FLORIDA CITRUS FRUITS. II, EFFECT OF VARIOUS 
WRAPPERS AND TEMPERATURES ON THE PRESERVATION OF CITRUS FRUITS 
IN STORAGE. Fla. Agr. Expt. Sta. Bul. 304, 78 pp., illus. 
(95) THOMPSON, H. C. 
1917. CELERY STORAGE EXPERIMENTS. U.S. Dept. Agr. Bul. 579, 26 pp., illus. 
(96) TRESSLER, D. K., and Evers, C. F. 
1936. THE FREEZING PRESERVATION OF FRUITS, FRUIT JUICES, AND VEGETABLES. 
369 pp., illus. New York. 
(97) VINCENT, C. C., VERNER, L., and BLODGETT, EF. C. 
1929. PROGRESS REPORT OF PRUNE STORAGE AND MATURITY STUDIES. Idaho 
Agr. Expt. Sta. Bul. 167, 19 pp., illus. 
(98) WALKER, J. C. 
1947. ONION DISEASES AND THEIR CONTROL. U.S. Dept. Agr. Farmers’ Bul. 
1060, 26 pp., illus. (Revised.) 
(99) WarpLaw, C. W. 
1934. PRELIMINARY OBSERVATIONS ON THE STORAGE OF AVOCADO PEARS. ‘Trop. 
Agr. [Trinidad] 11: 27-835, illus. 


(100) and Lronarp, EH. R. 
1935. THE STORAGE OF AVOCADO PEARS. Trop. Agr. [Trinidad] 12: 132-1383. 
(101) LEONARD, E. R., and BAKER, R. E. D. 
19384. OBSERVATIONS ON THE STORAGE OF VARIOUS FRUITS AND VEGETABLES. 
I. TOMATOES, CAULIFLOWERS, STRING BEANS, EGGPLANT, CUCUMBERS, 
AND MELONS. ‘Trop. Agr. [Trinidad] 11: 196-200, illus. 
(102) LEONARD, HE. R., and BAKER, R. E. D. 


1934. OBSERVATIONS ON THE STORAGE OF VARIOUS FRUITS AND VEGETABLES. 
II. PAPAWS, PINEAPPLES, GRANADILLAS, GRAPEFRUIT, AND ORANGES. 
Trop. Agr. [Trinidad] 11: 230-235. 
(103) = LEONARD, EH. R., and BARNELL, H. R. 
1939. METABOLIC AND STORAGE INVESTIGATIONS ON THE BANANA. ‘Trinidad 
Imp. Col. Trop. Agr. Low Temp. Res. Sta. Mem. 11, 61 pp., illus. 
(104) WHiItTEMaN, T. M. 
1932. COMMERCIAL FORCING OF LILIES-OF-THE-VALLEY. U.S. Dept. Agr. Cir. 
215, 20 pp., illus. 


(105) and McCLELLAN, W. D. 

1946. TIP CURVATURE OF CUT GLADIOLI. Amer. Soc. Hort. Sci. Proc. 47 : 515— 

521, illus. : i 

(106) and WRIGHT, R. C. 

1939. STORAGE OF WEDGWOOD IRIS BLOOMS. Florists’ Exch. 92 (14) : 16. 
(107) WRIGHT, R. C., and GRIFFITHS, D. 

1934. THE STORAGE OF TULIP BLOOMS. Florists’ Exch. 82 (3): 11, 14. 
(108) WariGut, R. C., and GRIFFITHS, D. 


1935. THE FORCING OF PAPER WHITE NARCISSUS BULBS AFTER STORAGE AT VARI- 
OUS TEMPERATURES. Amer. Soc. Hort. Sci. Proce. (1934) 32: 
645-650. 
(109) Wrant, J. S. 
1937. INVESTIGATIONS OF THE MARKET DISEASES OF CANTALOUPS AND HONEY 
DEW AND HONEY BALL MELONS. U. S. Dept. Agr. Tech. Bul. 573, 
48 pp., illus. 
(110) WititamMs, L. J. 
1933. COLD STORAGE OF PINEAIPLES AND PINEAPPLE JUICE. Ice and Refrig. 
85 : 25-26. 


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WINSTON, J. R. 
1935. REDUCING DECAY IN CITRUS FRUITS WITH BORAX. U. S. Dept. Agr. 
Tech. Bul. 488, 32 pp., illus. 


1936. A METHOD OF HARVESTING GRAPEFRUIT TO RETARD STEM-END ROT. U.S. 
Dept. Agr. Cir. 396, 8 pp., illus. 


1937. HARVESTING AND HANDLING CITRUS FRUITS IN THE GULF STATES. U. S. 
Dept. Agr. Farmers’ Bul. 1768, 38 pp., illus. 

WINTER, J. D., and ALDERMAN, W. H. 

1935. PICKING, HANDLING, AND REFRIGERATION OF RASPBERRIES AND STRAW- 
BERRIES. Minn. Agr. Expt. Sta. Bul. 318, 39 pp., illus. 

WOLFE, H. S., Toy, L. R., and STAHL, A. L. 

1946. AVOCADO PRODUCTION IN FLORIDA. Fla. Agr. Col. Ext. Bul. 129, 
107 pp., illus. 

WRIGHT, R. C. - 

1937. THE FREEZING TEMPERATURES OF SOME FRUITS, VEGETABLES, AND 
FLORISTS’ sTocKsS. U.S. Dept. Agr. Cir. 447, 11 pp. 


1941. INVESTIGATIONS ON THE STORAGE OF NUTS. U. S. Dept. Agr. Tech. 
Bul. 770, 35 pp., illus. 

DEMAREE, J. B., and Witcox, M. §S. 

1937. SOME EFFECTS OF DIFFERENT STORAGE TEMPERATURES ON THE KEEPING 
OF CRANBERRIES. Amer. Soc. Hort. Sci. Proc. (1936) 34: 897-401, 
illus. ; 

and GORMAN, E. A. 

1940. THE RIPENING AND REPACKING OF MATURE-GREEN TOMATOES. U. S. 
Dept. Agr. Cir. 566, 8 pp., illus. 

PEACOCK, W. M., WHITEMAN, T. M., and WHITMAN, Hb. F. 

1936. THE COOKING QUALITY, PALATABILITY, AND CARBOHYDRATE COMPOSITION 
OF POTATOES AS INFLUENCED BY STORAGH TEMPERATURE. U. S. Dept. 
Agr. Tech. Bul. 507, 20 pp., illus. 

PENTZER, W. T., WHITEMAN, T. M., and Ross, D. H. 

1931. EFFECT OF VARIOUS TEMPERATURES ON THE STORAGE AND RIPENING OF 
TOMATOES. U. S. Dept. Agr. Tech. Bul. 268, 35 pp., illus. 

and WHITEMAN, T. M. 

1931. DETERIORATION OF CHRISTMAS HOLLY IN TRANSIT AND STORAGE. U. S. 
Dept. Agr. Cir. 207, 12 pp., illus. 

and WHITEMAN, T. M. 

1949. A PROGRESS REPORT ON THE CHIPPING QUALITY OF 33 POTATO VARIETIES. 

Amer. Potato Jour. 26: [117]-120. 


U.S. GOVERNMENT PRINTING OFFICE: 1949 


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