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Das Tierreich.
Kine Zusammenstellung und Kennzeichnung der
rezenten Tierformen.
—_— Begrindet von der Deutschen Zoologischen Gesellschaft. sm _
Im Auftrage der
Kénigl. PreuB. Akademie der Wissenschaften zu Berlin
‘ “
»avra get.
herausgegeben von
Franz Kilhard Schulze.
»Sine systemate chaos.“
39. Lieferung.
Crustacea.
Cumacea
(Sympoda)
by
the Rev. T. R. R. Stebbing
Tunbridge Wells. HS Van
With 137 figures.
PES
Berlin.
Verlag von R. Friedlinder und Sohn.
Ausgegeben im September 1913, |
R. Friedlander & Sohn, Berlin NW 6, Karlstr. 11.
Das Tierreich.
Kine Zusammenstellung und Kennzeichnung der
rezenten Tierformen.
_— Begriindet von der Deutschen Zoologischen Gesellschaft. 2.
Im Auftrage der
Konigl. PreuB. Akademie der Wissenschaften zu Berlin
herausgegeben von
Franz Eilhard Schulze.
So
Seit Linnés Systema naturae ist die Zahl der bekannten Tierformen so angewachsen,
da eine neue, umfassende Ubersicht des Systems, die als Abschlu8 der bisherigen und
als Grundlage kiinftiger systematischer Forschung dienen kann, ein dringendes Bediirfnis
geworden ist. Um diese Aufgabe zu erfiillen, hat die Deutsche Zoologische Ge-
sellschaft das vorliegende Werk begriindet und dessen wissenschaftliche Leitung Herrn
Geh. Reg.-Rat Prof. F. E. Schulze in Berlin anvertraut. Das gewaltige Unternehmen
fand die Unterstiitzung der Koniglich PreuBischen Akademie der Wissen-
schaften, die in Wiirdigung der Bedeutung des Werkes im Jahre 1902 die Heraus-
gabe tibernommen hat.
Die einheitliche Durchfiihrung des Werkes ist durch eine Reihe wohldurchdachter
Bestimmungen gesichert. Fiir die Benennung der Tierformen und ihrer systematischen
Kategorien gelten die von dem 5. internationalen Zoologen-KongreB zu Berlin (1901)
angenommenen Regeln.
Die Herausgabe findet in Lieferungen statt, die je eine oder mehrere nahe-
stehende Gruppen behandeln, jedoch unabhangig von einer systematischen Folge er-
scheinen. Nach AbschluB einer jeden in mehreren Lieferungen behandelten Haupt-
abteilung erscheint ein Gesamtregister.
Jede Lieferung ist einzeln kauflich. Dem Umfang entsprechend ist der Preis
der Lieferungen verschieden; jedoch wird fiir die Subskribenten, die sich auf 5 Jahre
hinaus fiir die Abnahme aller in diesem Zeitraum erscheinenden Lieferungen verpflichten,
der Berechnung der durchschnittliche Preis von Mark 0,80 fiir den Druckbogen zugrunde
gelegt. Der Hinzel-Ladenpreis fiir jede vollstandige Lieferung erhdht sich gegen den
Subskriptionspreis um ein Drittel.
Bearbeiter: Dr. Graf K. Attems-Petzenstein in Wien, R. 8. Bagnall, Oxford, England,
Stadtbaurat T. Becker in Liegnitz, Graf H. v. Berlepsch auf SchloB Berlepsch, Prof. Berlese
in Florenz, Prof. R. Blanchard in Paris, Prof. F. Blochmann in Tiibingen, A. de Bormans
in Turin, Dr. K. Borner in Berlin, Prof. E.-L. Bouvier in Paris, Prof. J. Ch. Bradley in Ithaca (N. Y.),
Geh.-Rat Prof. K. Brandt in Kiel, Staatsrat Prof. M. Braun in Kénigsberg i. Pr., Dr. L. Breitfuss
in Katharinenhafen, F. Bryk in Helyla (Finnland), Prof. 0. Birger in Santiago (Chile), Prof. G.
Canestrini in Padua(+), Geh.-Rat Prof. ©. Chun in Leipzig, P. L. Czerny in Pfarrkirchen, Prof.
C. W. v. Dalla Torre in Innsbruck, Prof. F. Doflein in Freiburg B., Dr. G. Enderlein in Stettin,
Lehrer H. Fahrenholz in Hannover, Dr. 0. Finsch in Braunschweig, Dr. H. Friese in Schwerin,
Prof. A. Giard in Paris, Prof. W. Giesbrecht in Neapel (+), Hofrat Prof. L. v. Graff in Graz, Dr.
K. Grinberg in Berlin, Prof. V. Hacker in Halle, Dir. E. Hartert in Tring, Dr. R. Hartmeyer in
Berlin, Dr. C. E. Hellmayr in Miinchen, Dir. Dr. W. E. Hoyle in Cardiff, Dr. Ihle in Utrecht,
Dir. Prof. A. Jacobi in Dresden, Dr. K. Jordan in Tring, Prof. J. J. Kieffer in Bitsch, Dr. W. Kobelt
in Schwanheim, Prof. C. A. Kofoid in Berkeley (California), Prof. H. J. Kolbe in Berlin, Dir. Prof.
K. Kraepelin in Hamburg, Prof. P. Kramer in Magdeburg (+), Dr. H. A. Krauss in Tiibingen, Prof.
W. Kikenthal in Breslau, Dir. Prof. L. Kulezynski in Krakau, Dr. A. Labbé in Paris, Prof. A.
Lameere in Briissel, Prof. R. Lauterborn in Ludwigshafen a. Rh., Prof. R. v. Lendenfeld in Prag (+),
Prof. H. Lohmann in Hamburg, Geh.-Rat Prof. H. Ludwig in Bonn, Prof. M. Lithe in Kénigsbergi. Pr.,
A. D. Michael in London, Prof. W. Michaelsen in Hamburg, Dr. T. Mortensen in Kopenhagen,
Geh.-Rat Prof. G. W. Miller in Greifswald, Prof. J. P. Mc Murrich in Toronto, Prof. A. Nalepa in
Wien, Dr. G. Neumann in Dresden, Prof. L.-G. Neumann in Toulouse, Dr. Nieden in Berlin, Prof.
A. Nosek in Caslau, W. R. Ogilvie-Grant in Edinburgh, Dr. Ohaus in Berlin, Prof. A. E. Ortmann
in Princeton, Geh. Sanititsrat Dr. A. Pagenstecher in Wiesbaden (+), Prof. K. A. Penecke in Graz,
Prof. G. Pfeffer in Hamburg, Dir. Dr. R. Piersig in Annaberg (+), Prof. L. Plate in Jena, Geh.-Rat
Prof. A. Reichenow in Berlin, Prof. L. Rhumbler in Hannov.-Miinden, H. Riffarth in Berlin, Dir.
Dr. F. Ris in Rheinau (Schweiz), Dr. R, von Ritter-Zahony in Gérz, Dr. C. Fr. Roewer in Bremen,
The Hon. W. Rothschild in Tring, E. H, Riibsaamen in Berlin, Dr. F. Schaudinn in Hamburg(‘),
(Fortsetzung auf Seite 3 des Umschlages.)
Mnf be A. Ratt bar,
Wir KIND REGARDS,
): P ype PTEBBING,
] unsRivcs W ELLS.,
LIBRARY
Division of Crustacea
Das Tierreich.
Eine Zusammenstellung und Kennzeichnung der
rezenten Tierformen.
_.@ Begrindet von der Deutschen Zoologischen Gesellschaft. s_
Im Auftrage der
Konigl. Preuf. Akademie der Wissenschaften zu Berlin
herausgegeben von
Franz Kilhard Schulze.
» Udevra pet.“ »Sine systemate chaos.”
39. Lieferung.
Crustacea.
Cumacea
(Sympoda)
by
the Rev. T. R. R. Stebbing.
Tunbridge Wells.
With 187 figures.
Berlin.
Verlag von R. Friedlander und Sohn.
Ausgegeben im September 1913.
fg Was Pierreich
i Im Auftrage der
Konigl. Preu&f&{S. Akademie der Wissenschaften zu Berlin
herausgegeben von
Franz EHilhard Schulze.
39. Lieferung.
Crustacea.
Cumacea
(Sympoda)
the Rev. T. R. R. Stebbing M. A., F. B.S.
Tunbridge Wells.
With 137 figures.
h
at
Berlin.
Verlag von R. Friedlander und Sobn.
Ausgegeben im September 1913.
Alle Rechte vorbehalten.
A, Hopfer, Burg b. M.
Contents.
Pag.
IPTElAGO iis 8s Nie iat ogee ta eee ae V
Abbreviations of ihe ies of fenane PoC M LAS |. Jee re ee rmeres LVL
Systematic Index . . . ng LS Seer ee Re Neda ene apy. Ul
Ordo Cumacea (Symoda). Pei eae + cee kOe ae eee 1
Alphabetical Index .... Seth arene erty eal ct! “aileare ct le ae ieereaeae ON)
Nomenclator generum et ciSonam Pe eit tee tend ke js Le SOT
(The bibliography is carried down to February 1. 1913.)
Preface.
In regard to the small but very distinct group of malacostracan Crustacea
with which this monograph is concerned, special honour may be ascribed to
the Russian writer Lepechin and the American Say as clear sighted pioneers.
For elaborating our knowledge of the group Henrik Kroyer of Copenhagen is
foremost in the first half of the nineteenth century, while G. O. Sars of
Christiania in the second half outshines all other workers. During the century
now current Dr. Calman and Dr. Zimmer have taken the lead in progress.
That many other authorities have done excellent service will be clear from
the present summary of their labours, and their various merits will be best
estimated by those who have occasion and opportunity to consult the original
memoirs. Among zealous and sagacious collectors David Robertson of Cumbrae,
helpful as he was to Spence Bate, Anton Dohrn, and other well known students,
ought not himself to be forgotten.
For the last twenty years, and recently with more emphasis than before
by precept and practice, I have urged the expedience of assigning the mas-
culine gender to all generic names of animals. As, however, the older custom
is here followed, the Editor of ,,Das Tierreich* obligingly allows me to explain
that this is due to the established rules of his vast undertaking, and does not
imply any change in my personal opinion. Similarly, the ordinal name Cumacea
is retained for old acquaintance sake, although, with the cancelling of the
generic name Cuma, the alternative title of Sympoda for the Order invites,
and has from some authors of repute received, recognition and approval.
It is a pleasure to add, though it scarcely needs adding, that in passing
through the press these pages have been greatly indebted to the constant
care of the editorial staff.
July 26, 1913.
Thomas R. R. Stebbing,
Ephraim Lodge, The Common, Tunbridge Wells.
Abbreviations of the. titles of literature
Acta Ac. Petrop. — Acta Academiae Scientiarum Imperialis Petropolitanae. Petropoli. 4.
Act. Soc. Linn. Bordeaux. — Actes de la Société Linnéenne de Bordeaux. Bordeaux,
Paris, Londres. 8.
H. & A. Adams, Gen. Moll. — The Genera of recent Mollusca; arranged according to their
Organization. By Henry Adams and Arthur Adams. v.1, 2; atl. London.
1858 (1853—58). 8.
L. Agassiz, Nomencl. zool. — Nomenclator zoologicus, continens Nomina systematica
Generum Animalium tam viventium quam fossilium. Auctore L. Agassiz.
1 Vol. & Index universalis. Soloduri. 1842—46,46. 4.
Ann. nat. Hist. — The Annals and Magazine of natural History, including Zoology,
Botany, and Geology. London. 8.
Ann. S. Afr. Mus. — Annals of the South African Museum. Cape Town. 8.
Ann. Sci. nat. — Annales des Sciences naturelles. — [Sér.2—4:] Zoologie ([Sér. 5 & sequ.:]
Zoologie et Paléontologie). Paris, 8.
Ann. Scott. nat. Hist. — The Annals of Scottish natural History. A quarterly Magazine
with which is incorporated the Scottish Naturalist. Edinburgh & London. 8.
Annuaire Mus. St.- Pétersb. — Annuaire du Musée zoologique de |’Académie Impériale
des Sciences de St.-Pétersbourg. St.-Pétersbourg. 8,
Ann. Univ. Lyon — Annales de Université de Lyon. Paris. 8.
Arch. Naturg. — Archiv fiir Naturgeschichte. Berlin. 8.
Arch. Naturv. Kristian. — Archiv for Mathematik og Naturvidenskab. Kristiania. 8.
Arch. Zool. expér. — Archives de Zoologie expérimentale et générale. Paris. 8.
Belcher, Last arct. Voy. — The last of the arctic Voyages; being a Narrative of the
Expedition in H, M.S. Assistance, under the Command of Captain Sir Edward
Belcher, in Search of Sir John Franklin, during the Years 1852—54.
With Notes on the natural History, by Sir John Richardson, [Richard]
Owen, Thomas Bell, J. W. Salter, and Lovell Reeve. Vol. 1,2. London.
ESbo ao!
T. Bell, Brit. stalk-eyed Crust. — A History of the British stalk-eyed Crustacea by
Thomas Bell. London. 1853. 8.
Ber. Komm. D. Meere — Bericht der Kommission zur wissenschaftlichen Untersuchung
der Deutschen Meere in Kiel. Berlin. 2. ;
Bibl. hautes Etudes — Bibliothéque de l’Ecole des hautes Etudes. Section des Sciences
naturelles. Paris. 8.
Bih. Svenska Ak. — Bihang till Kongl. Svenska Vetenskaps-Akademiens Handlingar.
—- Afdelning 4, Zoologi. Stockholm. 8.
Borlase, Nat. Hist. Cornwall — The natural History of Cornwall... by Will. Borlase.
London 1758. 2.
Brit. Ass. Handb. — British Association Handbook. Fauna, Flora and Geology of the
Clyde Area, edited by G. F. Scott Elliot, Malcolm Laurie and J. Barclay
Murdach. Glasgow. 1901. 8.
Bull. Ac. St.-Pétersb, — Bulletin de l’Académie Impériale des Sciences de St.-Péters-
bourg. St.-Pétersbourg. 4 (8).
Bull. Mus. Paris — Bulletin du Muséum d'Histoire naturelle. Paris. 8.
Literatur-Kiirzungen Vil
Bull. Soc. Moscow — Bulletin de la Société Impériale des Naturalistes de Moscou.
Mosecou. 8.
Burmester, Beitr. Cuma — Beitrige zur Anatomie und Histologie von Cuma Rathkii
Kr. Inaugural- Dissertation . . . Kiel vorgelegt von Johannes Burmester,
Kellinghusen. 1883. 8.
Camp. arct. Orléans — Duc d'Orléans Campagne aretique de 1907. Bruxelles, 4.
Cellule — La Cellule. Recueil de Cytologie et d’Histologie générale. Publ. par J. B.
Carnoy, G. Gilson et J. Denys. Lierre & Gand, Lierre & Louvain, 4.
Claparéde & Lachmann, Etud. Infus. — Etudes sur les Infusoires et les Rhizopodes par
Edouard Claparéde et Johannes Lachmann. Partie I—11I. Geneve et
Bale. 1868. 4. [1858, 59, 61 in: Mém. Inst. Genev., v5, 6, 7.]
Claus, Freileb. Cop. — Die frei lebenden Copepoden mit besonderer Beriicksichtigung
der Fauna Deutschlands, der Nordsee und des Mittelmeeres von C. Claus.
Leipzig. 1863. 4.
G. Cuvier, Régne an., ed. 2 — Le Régne animal distribué d’aprés son Organisation, pour
servir de Base 4 I’Histoire naturelle des Animaux et d’Introduction a |’Ana-
tomie comparée. Par [Georges] Cuvier. Nouvelle Edition, revue et augmentee.
[Tom. 4 & 5:| Par [Pierre André] Latreille. Tom. 1—5. Paris. 1829,
29°°30;29,, 29; 8.
Dalyell, Powers Creator — The Powers of the Creator displayed in the Creation; or,
Observations on Life amidst the various Forms of the humbler Tribes of
animated Nature: with practical Comments and Illustrations. By John Graham
Dalyell. Vol. 1—3. London. 1851, 53, 58. 4.
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Systematic Index.
Pag.
Cumacea (Sympoda
T. Stebbing)
1.Fam. Vaunthompsoniidae
1. Gen. Vaunthompsonia Bate
1. V. inermis Zimmer.
2. ,, meridionalis O. Sars 9
3. ,, cristata Bate. 10
4. ,, arabica Calm. : 11
2. Gen. Gaussicuma Zimmer . iil
1. G. vanhoeffeni Zimmer ial
3. Gen. Bathyecuma H. J. Hansen 12
1. B. longicaudatum Calm. . 12
2. ,, brevirostre (Norm.) 138
3. ,, longirostre Calm. 13
4. ,, elongatum H. J. Hansen . 14
5. ,, natalense Stebb. 14
2. Fam. Sympodommatidae 15
1. Gen. Sympodomma TT. Stebbing 15
1. S. weberi (Calm.) 16
2. ,, anomalum (0. Sars) 16
3. ,, diomedeae (Calm.) . 17
4, ,, africanum Stebb. 17
3. Fam. Bodotriidae . 18
1. Gen. Cyclaspoides Bonnier 18
1. C. sarsi Bonnier : 18
2. Gen. Eocuma Mareusen . . 19
1. E. dollfusi Calm. . 20
2. ,, taprobanicum Calm. . 20
3. ,, longicorne Calm. 21
4. ,, hilgendorfi Mareusen 21
5. ,, stelliferum Calm. a ait
6. ,, latum Calm. . ae)
7. ,, Sarsil (Kossm.) Pee
8. ,, ferox (P. Fisch.). sek
9. ., affine Calm. . 5 OR
3. Gen. Bodotria Goodsir 23
1. B. pulchella (0. Sars). 24
a
Far neers oo ree eee eo ee crue
jee)
“
~~
7 2
yy
. Sublevis Calm.
» Scorpioides (Montagu) .
» gibba (O. Sars) .
. montagui Stebb.
australis Stebb. .
pulex (Zimmer) :
arenosa Goodsir
similis Calm.
., Slamensis Calm..
» parva Calm. .
audouinil (M.-E.)
concava D. Roberts.
, Sp., Calm. .
. Cyclaspis G. O. Sane :
; elacialis ile Elansent
sibogae Calm.
, longicaudata O. Sars
spectabilis Zimmer .
» picta Calm.
hornelli Calm.
.. herdmani Calm..
. levis G. M. Thoms. .
varians Calm.
10. ,, pusilla O. Sars
11. ,, longipes Calm.
12. ., persculpta Calm.
13. ,, exsculpta O. Sars
14. ,, elegans Calm.
15. ,, similis Calm.
16. ,, uniplicata Calm.
17. ,, unicornis Calm...
18. ,, cingulata Calm. .
19. ,, thomsoni Calm. .
20. ,, australis O. Sars —
2a quadrituberculata Zimine
22. ,, gigas Zimmer
23. ,, costata Calm.
24. ,, bistriata Zimmer
25. ,, triplicata Calm. .
. 28
“)
. brevipes
Systematic Index
. Stephanomma 0. Sars
. goésil O. Sars
. Zygosiphon Calman
. mortenseni Calm.
. Iphinoé Bate
. crassipes H. J. Hansen
zimmeri Stebb. .
H. J. Hansen.
serrata (Norm.) :
.. trispinosa (Goodsir)
.. tenella O. Sars .
. robusta H. J. Hansen
inermis O. Sars .
. Heterocuma Miers
. Sarsi Miers
. granulatum Miers
en.
C.
Cumopsis 0. Sars .
goodsir (Bened.)
longipes (A. Dohrn)
Pag. |
5 CO)
. 40 |
41
ell
. 42
Bodotriidarum genera dubia et
species dubiae . 51
Gammarus esca F. 51
Cuma fasciata E. Hesse . 51
» parva E. Hesse . 51
» punctata E. Hesse 51
rufa E. Hesse . el
, terginigra E. Hesse 51
» folini P. Fisch. 51
4. Fam. Ceratocumatidae . 51
1. Gen. Ceratocuma Calman 51
1. C. horridum Calm. . 52
5. Fam. Leptocumatidae. . 52
1. Gen. Leptocuma 0. Sars 53
1. L. kinbergii O. Sars 53
2. 5. minus Calm. . 54
6. Fam. Chalarostylidae 54
1. Gen. Chalarostylis A. M. Norman 54
1. C. elegans Norm. 54
2. Gen. Hemilamprops 0. San 55
1. H. rosea (Norm.) 55
2. ,, assimilis O. Sars 56
3. ,, normani Bonnier 56
4 pellucida Zimmer oi
5. ,, cristata (O. Sars) 57
6. ,, uniplicata (O. Sars) 58
7. Fam. Paralampropidae . 58
1. Gen. Paralamprops 0. Sars 59
1. P. serratocostata (O. Sars) 59
2. ,, aspera Zimmer , 60
Xl
Pag
8. Fam. Platysympodidae . 60
1. Gen. Platysympus T. Stebbing 61
1. P. typicus (O. Sars) 61
2. ,, brachyurus (Zimmer) 562
9. Fam. Leuconidae . ; 62
1. Gen. Leucon Kroyer. a63
1. L. siphonatus Calm. 64.
2. ,, tenuirostris O. Sars . 65
3. ,, nasicoides Lilj. . 65
4. ,, australis Calm. . - 66
5. ,, fulvus O. Sars 66
6. ,, nasica (Kroyer) . - 66
7. ,, antarcticus Zimmer . 68
8. ,, nathorsti Ohlin . 68
Y. ,, septemdentatus Zimmer . 69
10. ,, mediterraneus O. Sars 69
11. ,, heterostylis Calm. 70
12. ,, longirostris O. Sars 70
13. ,, kalluropus Stebb. 71
14. ,, pallidus 0. Sars. 71
15. ,, serratus Norm. i
16. ,, kerguelensis Zimmer 72
17. ,, assimilis O. Sars 72
18. ,, acutirostris O. Sars 73
19. ,, sagitta Zimmer . 73
20. ,, vanhoeffeni Zimmer 74.
,, leuckartii Mareusen . 74
2. Gen. Eudorella A. M. Norman . 74
1. E. emarginata (Kroyer) 75
2. ,, gracilior Zimmer 77
3. ,, fallax Zimmer 77
4. ,, nana O. Sars 78
5. ,, monodon Calm. . 78
6. ,, hirsuta (O. Sars) 78
7. ,, hispida O. Sars . 79
8. ,, gracilis O. Sars . 19
9. ,, splendida Zimmer 80
10. ,, truncatula (Bate) . 80
11. ,, pusilla O. Sars . 81
12. ,, abyssi O. Sars whi)!
lisse sordida Zimmer . . 8l
3. Gen. Eudorellopsis 0. Sas 82
1. EK. deformis (Kroyer) 82
2. ,, integra (S. I. Sm.) . . 83
3. ,, resima Calm. . 84
4. ,, biplicata Calm. . 84
4, Gen. Pseudoleucon Zimmer . 85
1. P. sorex Zimmer = 80.
10. Fam. Diastyloididae 86
1. Gen. Diastyloides 0. Sars . 86
XIV
Pag.
1. D. serratus (O. Sars) . 86
., biplicatus (O. Sars) 87
11. Fam. Diastylidae 88
1. Gen. Diastylis Say . 89
1. D. bidentatus Calm. . 91
2. Galliy Calman Oil
3. ,, planifrons Calm. 91
4. ,, alaskensis Calm. 92
5. ., anderssoni Zimmer 92
6. .. hammoniae Zimmer 93
7. ., antillensis O. Sars 93
8. ., koreanus Calm. 93
9, ., algoae Zimmer? 94
10. ., spinulosus Heller . 94
ll. ., armatus Norm. 95
12. ,, bispinosus (Stimps.) 95
13. .. cornutus (Boeck) . 96
14. ,, laevis Norm. 97
15. ,, sulcatus Calm. . 97
16. ., rathkii (Kroyer) 98
17. ,, bradyi Norm. 99
18. ., goodsiri (Bell) . 99
19. ,, stygius O. Sars 100
20. ,, polaris O. Sars. 100
21. ,, lucifer (Kroyer) 101
22. ,, rugosoides A. Walker 101
23. ,, scorpioides (Lepech.) . 102
24. ., rugosus O. Sars 102
25. ,, tumidus (Lilj.). 103
26. ., neapolitanus O. Sars . 104
27. ,, echinatus Bate. 104
28. ., tricinctus (Zimmer) 105
29. ., nucella Calm. 105
30. ,, anomalus Bonnier . 106
31. ,, asper Calm. 106
. ambigua Lilj. 107
» quadriplicata Lilj. 107
2. Gen. Brachydiastylis I. Steb-
bing SOV
1. B. resimus (coon 107
3. Gen. Diastylopsis S. I. Smith 109
1. D. neozealanica(G.M.Thoms.) 110
2. ,, dawsoni S. I. Sm.. el 1G
3. ,, elongata Calm.. > ltd
4. ,, crassior Calm. . pert
5. ,, annulata (Zimmer) = alii
6. ,, robusta (Zimmer) . Se
7. ., thileniusi (Zimmer) eplelis
8. ., dentifrons (Zimmer) a ea!
» Insularum (Calm.). 14...
Systematic Index
Pag.
D. sp., (Calm.) . . 114
4. Gen. Adiastylis T. Stobbine + 114
1. A. acanthodes Stebb.. aes
v. ,, longicaudatus (Bonnier) . 115
3. ,, longipes (O. Sars) . E16
4. ,, costatus (Bonnier). . 116
5 Com Makrokylindrus T. Steb-
bing ety)
1, M. serricauda (r Scott). ewer
2. ,, fragilis Stebb. en
3. ,, cingulatus (Calm.) . 119
4. ,, dubius (Bonnier) org
5. ,, erinaceus (O. Sars) . 120
6. ,, josephinae (O.'Sars) . . 120
6. Gen. Paradiastylis Calman . 121
1. P. brevicaudatus (Zimmer) . 121
2. ,, brachyurus Calm. . e121
3. ,, longipes Calm. . » 129
7. Gen. Leptostylis 0. Sars . . 123
1. L. mancus O. Sars 123
2. ., antipus Zimmer . 124
3. ,, ampullaceus (Lilj.) eE
4. ,, villosus O. Sars 7 125
5. ., crassicauda Zimmer . 125
6. ., borealis Stappers . 125
7. 5. macrurus.O. Sars . . 126
8. ., macruroides Stebb. . 126
9. ,, productus Norm. OF
10. ,, longimanus (0. Sars). . 127
11. ,, gracilis Stappers . 128
8. Gen. Ekleptostylis T. Stebbing 129
walker (Calm. ) . 129
12. Fam. Colurostylidae. . 129
1. Gen. Colurostylis Calman. . 130
1. C. pseudocuma Calm. . 130
2. Gen. Anchicolurus T. Stebbing
130
1. A. occidentalis (Calm.) . . 130
Genus incertae sedis, near
Colurostylidae. eersik
Gen. Pachystylis H. J. Tignten
P. rotundatus H. J. Hansen
13. Fam. Oxyurostylidae
}. Gen. Oxyurostylis Calman
1. O. smithi Calm.
14. Fam. Pseudodiastylidae
1. Gen. Pseudodiastylis Calman
1. ‘P. ferox Calm.
15. Fam. Ekdiastylidae
1. Gen. Ekdiastylis T. Stebbing
131
131
. 182
. 132
. 132
133
133
. 133
. 135
135
SONS oe poe
9
. sculptus (O. Sars) .
politus (S. L. Sm.).
insignis (O. Sars) c
, mystacinus (0. Sars) .
hexaceros (Zimmer)
argentatus (Calm.)
horridus (O. Sars).
abbreviatus (O. Sars).
fimbriatus (O. Sars) .
16. aves Holostylidae.
1. Gen. Holostylis T. Stebbing .
1. H. helleri (Zimmer)
ens.
17. Fam. Pseudocumatidae
1. Gen. Stebbing
ie:
2. Gen.
SS:
Gen.
S.
. Pseudocuma 0.
. cercaroides O. Sars
, Simile O. Sars .
gayi (Nic.) .
Petalosarsia TT.
declivis (O. Sars) .
Schizorhynechus O. Sars
scabriusculus (O. Sars)
eudorelloides (O. Sars)
bilamellatus (O. Sars)
abbreviatus (O. Sars).
. Caspiocuma 0. Sars.
'. campylaspoides (O. Sars)
. Pterocuma 0. Sars
.rostratum (0.
Sars)
pectinatum (Sowinsky)
sowinskyi (O. Sars)
. Stenocuma 0. Sars
. diastyloides (O. Sars).
, tenuicauda (0.
Sars) .
gracile (O. Sars)
graciloides (O. Sars) .
Sars
longicorne (Bate) .
ciliatum 0. Sars
lagunae Baker .
Strauchia Czerniavski
taurica Czern.
18. Fam. Paraleuconidae
1. Gen.
ie
Paraleucon Calman .
suterl Calm.
19. Fam. Hemileuconidae
1. Gen.
Hemileucon Calman.
1. H. uniplicatus Calm. .
2. 5,
comes Calm.
20. Fam. Lampropidae
1. Gen.
Lamprops O. Sars
Systematic
Pag.
5 ey)
. 136
. 136 |
5) LH
5 By
ole
. 138
. 138
5 IBY
. 139
139
. 140
. 140
140
141
. 142
143 |
. 148
. 148
. 144
. 144
. 145
145
. 145
. 145
. 146
. 146
. 147
. 147
. 147
. 148
. 148
. 148
. 149
. 149
5 alee
. 150
oy kay)
lol
5 lap
« 15
ay libil
yelloill
. 152
. 152
. 152
. 152
. 153
. 153
Index
y. Gent StenotyphlopsT. Senne
3. Gen. Platytyphlops be Sepiing
1. P. peringueyi Stebb. .
4. Gen. Bathylamprops Zimmer
1. L. fuseata O. Sars
2. ,, fasciata O. Sars
3. ,, quadriplicata S. I. Sm.
. comata Zimmer
, bering Calm.
1. S. spinulosa Stebb.
2.
ils
» orbicularis (Calm.)
2
.
calmani Zimmer
21. Fam. Dicidae ..
1. Gen. Die T. Stebbing
1. D. calmani Stebb..
ee
22. Fam. Gynodiastylidae .
1. Gen. Gynodiastylis Calman
23. Fam. Heteroleuconidae
1. Gen. Heteroleucon Calman
2, Gen. Paranannastacus T. Steb-
bing
ce
S212) BOS Mie em lep
tubulicauda (Calm.)
1. G. laevis Calm.
2
oO:
- 498
carinatus Calm.
.. costatus Calm. .
. bicristatus Calm.
1. H. akaroénsis Calm. .
24. Fam. Nannastacidae .
1. Gen. Schizotrema Calman
te S. calmani Stebb..
2. , depressum Calm.
a
3
. bifrons Calm.
sordidum Calm.
TE reptans ( (Galea)
Pr
oo)
ae
tardus (Calm.) .
. Nannastacus Bate
zimmer Calm. .
suhmil O. Sars.
gibbosus Calm.
minor Calm.
agnatus Calm. .
» lepturus Calm. .
» longirostris O. Sars
brachydactylus Calm..
ossianl Stebb.
10. ., hanseni Calm. .
11. ,, georgi Stebb.
12. ,, pardus Calm.
13. ,, hirsutus H. J. Hansen
XVI Systematic Index
Pag. Pag.
14. N. brevicaudatus Calm... 174 3. P. tridentata Stebb. . . . 185
15. ,, unguiculatus (Bate) . . 174 4. ,, armata Bonnier . . . 186
IM sae stebbingi Cala ie 176 | o6 Pam Gamprlaspididaemiss
EhvarSuigmna paeecotias 176 " j Ge ; PY j P i
Gen. Diops Paulson. . . . 176 oe ee G. O. Sars 187
D. parvulus Paulson.-. . . 176 5 ak YOPOpMS Calas) 2) gheoe
Nannastacus spinosus (eaaieeay 176 Oct ee ee sie sale
4. Gen. Platyeuma Calman . . 176 > RSS ae ea eRe
1: Pchol Cala 176 4. , pulchella 0. Sars . . ~ 190
ia) ‘Piss ue, ene 5. ;, pacili¢a, OoSars) - 7 190
_N oJ / 7
e. et iia sae ta Ce is 6. ,, orientalis Calm. . . . 190
Sok aoe Ga ee oh . i. ey Cuily.) = 4) ae
BY tate et a + 5) Qtaord> OSSars eno
6. Gen. Cumella 0. Sars... 178 9. ,, spinosa Calm. . . . . 192
1. C. australis Calm.. . . . 179 10. ., costata O. Sars. . . . 193
a 99 eracilliima (Gali, 6 6 4 UC 11. .. vitrea Calm. 2 alas
3. ,, molossa Zimmer . . . 180 12. ,,. ovalissStebbs a2. ee eos
: ” tee vio a 13. ., macrophthalma O. Sars . 195
- » Clavicauda Calm... . 14. ,, undata 0: Sars? iy). 71195
6. ,, leptopus Calm. . - + + 181 1b. f sulcata O. Sars. . . . 195
# ” pee Cat ad 6c aie ie 16. ,, horrida’ O2Sars “= )2 9196
.» Serrata Calm. . . . . 182 eee
9, nevis Calm... 5. en |! fe ieee 2
10. ,, pygmaea O. Sars . . . 182 19: ,,. affinis 70) Sars 1a. n-etom
ner limicola ©) Sars: 2s ss 20.-,,. frigida H. J. Hansen. . 197
» abyssicola (Norm). . . 184 21..,, verrucosa O. Sars. . . 198
» carinata (H. J. Hansen) . 184 | 99 nodulosa 0. Sars . . . 198
95. Fam. Sonar cae ras 23. ,,.antarctica Calms . . . 199
dacra as . 184 | Genera Cumaceorum incertae
1. Gen. Procampylaspis eens 184 sedis). 2: . 199
iP. bonmertsCalm.. 9 2e185 Gen. Galyptecteos Toe . 199
2. ,, compressa Zimmer . . 185 Gen. Olbia Marcusen . . . 199
Cumacea
(Sympoda, '. Stebbing)
1779 Gen. Gammarus (part.), J. C. Fabricius, Reise Norweg., p.249 | 1780 Gen.
Oniscus (part.), Lepechin in: Acta Ac. Petrop., 1778 v.1 p.248 | 1788 Gen. and Subgen.
Cancer (Gammarus) (part.), J. F. Gmelin, Syst. Nat., v5 p. 2993 | 1793 Gen. and Subgen.
Cancer (Oniscus) (part.), J. F. W. Herbst, Naturg. Krabben Krebse, v.21v p.146 | 1804
Gen. Cancer (part.), Montagu in: Tr. Linn. Sqe. London, v.7 p.70 | 1812 Gen. Astacus
(part.), Pennant, Brit. Zool., ed.5 v.4 p.35 | 1818 Gen. Diastylis, Say in: J. Ac. Philad.,
v. lit p.313 | 1828 Gen. Cuma (non [Calonne] Mus. Calonn. 1797 Moll.), H. Milne-Edwards
in: Ann. Sci. nat., v.13 p. 294 | 1829 Gen. Condylura (non Illiger 1811 Mam.),
Latreille in: G. Cuvier, Régne an., ed.2 v4 p.153 | 1851 Ord. Dicladopa (part.),
Latreille, Cours Ent., v.1 p. 159, 414 | 1840 Ord. ,, Diclapodes“, group ,,Cumites“ (part.),
H. Lucas, Hist. An. artic., p.266, 267 | 1846 Ord. Cumacea, Kroyer in: Naturh. Tidsskr.,
ser.2 v.211 p. 202, 203 | 1849 Larval Decapod, Gen. Cuma, H. Nicolet in: Gay, Hist.
Chile, v.38 p.220 | 1852 Subtrib. Cumidea, J. D. Dana in: U.S. expl. Exp., v. 131
p.502 | 1853 Fam. Cumadae, T.Bell, Brit. stalk-eyed Crust., p.321 | 1856 Fam. Diastylidae,
Bate in: Ann. nat. Hist., ser.2 v.17 p.449 | 1863 Fam. D., G.O.Sars in: Nyt Mag.
Naturv., v.12 p.236 | 1863 Division ,Cumaceen”, C. Claus, Freileb. Cop., p.18 | 1865
Ord. Cumacea, G. O. Sars in: Forh. Selsk. Christian., 1864 p.153 | 1883 Group Cuma,
Burmester, Beitr. Cuma (Anatomy and Histology) | 1883 Cuwmacea, Boas in: Morphol.
Jahrb., v.8 p.571 | 1887 C., G. O. Sars in: Rep. Voy. Challenger, v.19 pars 55 p.1 |
1893 Subord. C., T. Stebbing in: Int. sei. Ser., v. 74 p. 291 | 1893 ,,Cumaceen“,
Butschinsky in: Zool. Anz., v.16 p. 386 (Embryology) | 1899 Cumacea, G. O. Sars,
Crust. Norway, v.3 | 1900 Ord. Sympoda, T. Stebbing in: Willey, Zool. Results, pars 5
p. 606, 609 | 1901 Subord. S., T. Scott in: Brit. Ass. Handb., p. 332 | 1905 Ord. Cumacea,
Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p.3 | 1906 Ord. Sympoda, A. M.
Norman & T. Scott, Crust. Devon Cornwall, p.29 | 1909 S., A. M. Norman & G.S.
Brady in: Nat. Hist. Tr. Northumb., ser.2 v.31 p.25 | 1907 S., A. M. Norman in:
Ann. nat. Hist., ser.7 v.20 p.361 | 1908 ,,Cumaceen“, Zimmer in: Ergeb. Tiefsee-Exp.,
v.8 p.155 | 1908 Sympoda, Stappers in: Arch. Zool. expér., ser.4 v.8 Notes p.98 |
1909 §., Stappers in: Cellule, v.25 p.352 (alimentary system) | 1910 S., T. Stebbing
in: Ann. 8S. Afr. Mus., v.61v p.409 | 1911 S., Stappers in: Camp. arct. Orléans, Crust.
Malac. p.99 | 1912 Cumacea, Calman in: P. U.S. Mus., v.41 p.603 | 1912 Sympoda,
T. Stebbing in: Ann. S. Afr. Mus., v.10 p.129 | 1913 S., Giesbrecht in: Lang, Handb.
Morphol., v. 4. p. 13.
Externally the body is divisible into three compartments:
1) a large carapace covering the sessile eye or pair of eyes when
present, the bases of two pairs of antennae, the mandibles, two
pairs of maxillae, and. three pairs of maxillipeds; 2) five leg-
bearing segments all normally free but subject to fusion of the
Das Tierreich. 39. Lief.: T. R. R. Stebbing, Cumacea. 1
2 Cumacea
first, second, and third, with the carapace and of one with an-
other among themselves, each limb of the first pair having its
Antenna 1
Eye-- —- - -7%
L/
---- Antenna 2, peduncle
Carapace *- --—-—-—
Peraeon
segment 2
Peraeon | _
segment 5
Pleon
segment 1 - Antenna 2, flagellum
SAAS 3) eet Pleopod 1
Telsonic segment - - — —— LS eee ee Exopod
of uropod
--
‘
Uropod peduncle
Endopod of uropod
Fig. 1.
Bodotria arenosa (after G. O. Sars).
exopod or swimming-branch, an appendage which may or may
not recur on any or all of the three following pairs but is never
found on the fifth pair; 3) a flexible pleon of six or seven mo-
Cumacea 3
vably articulated segments, of which the first five may carry
each a pair of pleopods, and the sixth is always furnished with
a pair of uropods, while the seventh, the telson, never has appen-
Antenna 1
Peraeopod 1
Eyelobe
Branchial apparatus
Cephalic ganglion
exopodal part
Se am ti }
Peraeopod 2 Stomach
- - - Carapace
Branchial apparatus eS
epipodal part | ;
Tea) oe | 3 liversacs
Nerve chain
Peraeopod 3
Peraeopod 4 -~-
Incubatory plate
(in formation)
\) Intestinal canal
Peraeopod 5 ----"SS
Pleon segment 1
Fig. 2.
Diastylis lucifer (after G. O. Sars).
dages. Internally, the heart (so far as known) within a narrow
pericardial sinus has three pairs of lateral openings, is fusiform,
and does not extend beyond the peraeon segments; a short dilated
one-chambered stomach is followed by along, generally straight,
1*
4 Cumacea
intestinal canal with the anal opening in the telson, or in the
equivalent portion of the telsonic segment, when the telson itself
is in fusion with the sixth segment of the pleon; hepatopan-
creatic glands situated below the stomach form from one to four
pairs of lobes; the nerve chain has a dorsal cephalic ganglion
followed ventrally by ten well developed ganglia and six caudal
ganglia of which all but the lastare very slight, the whole series
connected by double commissures. Ovaries and testes paired.
Development of young carried far forward within the body of
the mother.
Integument calcified, more or less brittle. The carapace usually has
a bell-shaped frontal lobe, with small, often eyeless, eyelobe at its apex, both
enclosed by a pair of pseudorostral lobes drawn round from the sides to
meet in front; the inferolateral margins fold underneath, leaving a space
which is occupied by the third pair of maxillipeds except for a narrow opening
on either side to admit water into the branchial cavities; when the epipods
of the first maxillipeds, commonly furnished more or less with branchial leaflets,
have been sufficiently irrigated, the pair of exopods in combination, in
coalescence, or separately and apart shoot forward to leave room for the
water to pass out in front, and are then withdrawn to close the openings
till a fresh supply has been made use of, this respiratory arrangement being
apparently found only in this Order of Crustacea. The fifth pedigerous
segment is narrower than the rest, its appendages more or less late in
development, and sometimes, it appears, permanently wanting. The narrow
pleon is always without pleopods in the female. The telson, to which the
anal aperture belongs, is often fused with the preceding segment, the term
telsonic segment being applied to the resulting compartment.
Kyes may be entirely wanting, or only represented by pigment, or
efficient with corneal lenses varying in number from 2 or 3 to 11; the
lenses of the single eye are usually but not always in close proximity; eyes
distinctly paired are rare. First antennae never elongate, the peduncle
three-jointed, often stouter in male than in female, principal flagellum scarce-
ly ever as long as the peduncle, tipped with one or two sensory filaments,
and often in the male surrounded at the base by a fascicle of them,
accessory or medial flagellum smaller than the principal, sometimes almost
imperceptible. Second antennae in female from one- to five-jointed, never
elongate; in the adult male sometimes exceeding the length of the body by
the gradually elongated, many-jointed flagellum, the last two jomts of the
peduncle often fringed with brush of sensory filaments, the whole apparatus
resembling similar structures among the Amphipoda and Phyllopoda, sheltered
when out of use at the sides of the body; the flagella in Lamprops are of
moderate length, prehensile, used for clasping the female. Upper lip simple,
often faintly emarginate. Lower lip bilobed. Mandible without palp,
with dentate cutting edge, accessory plate at least on one member of the
pair, a spine-row of spines varying from two to a score, molar stout or
rarely slender, base of the trunk usually narrowing to the rear, sometimes
broad. First maxillae as a rule with medial plate narrower than the
lateral, surrounded by few spines, lateral plate with about ten, the so-called
palp turned straight back or rarely absent, when present carrying distally
two filaments or a single filament, probably employed in cleansing the epipod
of the first maxillipeds and its branchial leaflets. Second maxillae generally
divided into two or three spiniferous plates, occasionally undivided. First
or
Cumacea
maxillipeds with second joint usually produced into a plate furnished with
coupling spines, the third joint obscure, rarely the last joint degraded with
the three preceding joints fused into one, the branchial leaflets varying in
number from two or three, or in female even none, to several dozens.
Second maxillipeds having at the base a pair of plates, perhaps epipodal,
Fig. 3.
Fig. 4.
Later larval stage of Diastylis lucifer (Kroyer), after Sars.
Fig. 3 and 4. a antenna 1, a? antenna 2, br.exp. exopodal part of the branchial apparatus,
h heart, i intestinum, LZ anterior lip, 1 posterior lip, / liver sacs, M mandibles, m* anterior
maxilla, m? posterior maxilla, mp'—mp* maxilliped 1—2, O eye, p'~ peraeopod 1—4, R rostral
plates of carapace, ¢ telson, urp uropod, v remnant of the yolk-mass.
which in the female send a fan of filaments into the incubatory cavity.
Third maxillipeds more or less leglike, almost always provided with exopods.
The five pairs of peraeopods not chelate or subchelate, the 24 joint normally
the longest, stout’ or otherwise in accord with the exopodal furniture; the
third joint in maxillipeds and peraeopods scarcely ever elongate, sometimes
6 Cumacea
evanescent. Pairs of pleopods in the male attached to the first five, the
first three, the first two, the first, or to none of the pleon segments, their
structure consisting of a peduncle generally carrying two rami, which are
never more than two-jointed. The much more conspicuous uropods are
always present, consisting of a peduncle carrying two rami, the exopod always
two-jointed, the endopod having one, two, or three joints.
Size ranging from under 2 mm to 35 mm.
The group has been variously classified, among the Amphipoda by
J.C. Fabricius, the Isopoda by Lepechin, the Macrura by Pennant and
Goodsir, as a larval form of Macrura by H. Milne-Edwards, Nicolet,
Agassiz, and Dana, among the Schizopoda by Say, Desmarest, Latreille,
Kossmann, among the (unrestricted) Stomapoda by De Kay, Bate,
Danielssen, placed near to Mysis by Huxley, as a suborder of Mysidacea
by Boas, among Branchiopoda by Latreille and Stimpson, recognised as
an independent group by Kréyer, van Beneden (sessile-eyed Decapoda),
Claus (between Malacostraca and Copepoda), Fritz Miller (nearer to stalk-
eyed than sessile-eyed Malacostraca), Dohrn (between Edriophthalma and
Mysis), Paulson, G. 0. Sars, T. Stebbing, T. Scott, Zimmer, Calman
and others. Dohrn finds affinity of the group with the Isopoda by the
embryonal development, Delage derives it from the Tanaidae, G. O. Sars
and Stappers agree in placing it between the Podophthalma and Edrioph-
thalma, but G. O. Sars considers its approach nearer to the stalk-eyed
Crustacea, and Stappers thinks that the proximity is incontestably closer
to the sessile-eyed.
Distribution in all Oceans, from shore between tide-marks to great depths, Caspian
Sea to mouth of the Volga.
26 Families, 62 accepted and 4 doubtful genera, 309 accepted and 23 doubtful
species.
Synopsis of Families chiefly based on male specimens:
Five pairs of pleopods in the male — 2.
Fewer than five pairs of pleopods in the
male — 5.
Four pairs of peraeopods in the male with
CXOPOdS 20 sa wel wie eines oo alan ae 1. Fam, Vaunthompsoniidae
Fewer than four pairs of peraeopods in the
male with exopods — 3.
Three pairs of peraeopods in the male with
3 well developed exopods. ....... . 2. Fam. Sympodommatidae
Fewer than three pairs of peraeopods in the
male with well developed exopods — 4.
No separate telson ..........- ++. 3, Fam. Bodotriidae
Telson separate. ..-+......+.+. +. . 4 Fam. Ceratocumatidae
Three pairs of pleopods in the male — 6.
Fewer than three pairs of pleopods in the
male — 9.
No separate telson .......+.+ ++... 5. Fam. Leptocumatidae
Telson separate — 7.
First maxillae with palp. ....... .. 6, Fam. Chalarostylidae
First maxillae without palp — 8.
Cumacea
More than one pair of peraeopods in female
with exopods .
Only the first pair of peracopods i in anne
with exopods .. . ir ee
8
Pleopods in male limited to two pairs — 10.
Pleopods in male fewer than two pairs — 18.
10 { No separate telson
| Telson separate — 11.
eyues er er .6 Te, 6 2
Uropods with three-jointed endopod — 12.
Uropods with fewer than three joints to the
endopod — 16.
Mandibles broad at the base... .
11
12) Mandibles narrow at the base — 18.
Telson with only two apicai spines. .
-13 2 Telson with fewer or more than two Apia
spines — 14.
14 Melsontsmall’ s° 2 3 6°:
Telson large — 15.
15 Telson with no apical spines =e
Telson with three apical spines. .
16 Uropods with two-jointed endopod ... .
Uropods with endopod simple — 17.
17 ) Telson small with no apical spines... -
18 Pleopods in male a single pair. ..... -
Pleopods in male wanting — 19.
19 Mandibles broad at base... . BG)
Mandibles not broad at base — 20.
Telson separate — 21.
20 1) No separate telson — 23.
9 Apical spines of telson three or more .
—
Uropods with three-jointed endopod
Uropods with endopod two-jointed or simple,
telson medium .
22
Only first two pairs of ES eue in male
with exopods :
First four pairs of meraeapods in rale th
exopods — 24.
23
a Telson large with two apical spines .
J
\
{ Mandibles with stout molar
) Mandibles with slender molar — 25.
2
rs
Second maxiila and first maxilliped normal,
| second maxilliped with gies joint peculiarly
dentate ....
| Second maxilla Bhidivided: ack maniliped with
oy 10), 10" (oa -@
25
reduced number of joints, second maxilli-
ped normal
Apical spines of telson fewer than three — 22,
)
12.
13. F
14,
15. F
16.
Nei
18.
19. F
23.
24,
26.
. Fam.
. Fam.
Fam.
. Fam.
. Fam.
. Fam.
. Fam.
. Fam.
Fam.
Fam.
. Fam.
Fam.
Paralampropidae
Platysympodidae
Leuconidae
Diastyloididae e
Diastylidae
. Colurostylidae
. Oxyurostylidae
. Pseudodiastylidae
. Ekdiastylidae
. Holostylidae
. Pseudocumatidae
. Paraleuconidae
. Hemileuconidae
Lampropidae
Dicidae
Gynodiastylidae
Heteroleuconidae
Nannastacidae
Procampylaspididae
Campylaspididae
8 Cumacea: 1. Vaunthompsoniidae
Synoptic table for female specimens.
ss 3 gd |aov Ss 2
S Be NS 2 2 = * 6 | Auxiliary marks of
a as |Se3| 925 distinction !)
“ | 28 ps8] sae
=) am
]
large |[Zormore|) 3 |2-+2r?) 20. Fam. Lampropidae
ample 3ormore; 3 2+ 2r 6. , Chalarostylidae *)
large 3 ? 2+2r a.s.,ace.fl.verysmall| 14. ,, Pseudodiastylidae
large 3 3 212r | mx.1lwithoutpalp | 7. ,, Paralampropidae
large 3 3 1 mx. 1 without palp | 8. ,, Platysympodidae
large | 2 3 |2or2-+2r |11. ,, Diastylidae
large 2 3 2 m. broad at base 10. ,, Diastyloididae
large 2 2 2 15. ,, Ekdiastylidae
large 2 i 2 16. , Holostylidae
large 0 3 ? 21. 5 wWDicidse*)
large 0 3 2+ 21 telson acute 13. , Oxyurostylidae
medium! Oor2r | 2orl 2 (22. , Gynodiastylidae
small 0 2or3| 2+2r | 12. ,, Colurostylidae
small 0 1 2+-2r | branchial sacs few 17. ,, Pseudocumatidae
small 0 1 ? prp. 5 undeveloped | 4. ,, Ceratocumatidae *)
0 yo 2 3+ r 5. , Leptocumatidae
0 : 2 3 1. ,, Vaunthompsoniidae
0 2 3 _ eyelobe linguiform 2. ,, Sympodommatidae
0 2 3 m. broad at base 9. ,, Leuconidae
0 2 3 | m. broad at base 18. , Paraleuconidae
0 2 38 m. broad at base 19. ,, Hemileuconidae
0 1 or2|lorl+2r 3. , Bodotriidae
0 1 2 mxp. 2, dentate apex | 25. , Procampylaspididae
0 1 2 | m.molarstiliform | 26. ,, Campylaspididae
0 1 2 m. broad at base 23. , Heteroleuconidae
0 1 2 or 0 24. , Nannastacidae
1. Fam. Vaunthompsoniidae
1878 Vaunthompsondiiae (err. typ.), G.O. Sars in: Arch. Naturv. Kristian., 0.3
p. 465 | 1879 Vaunthompsoniidae, G.O.Sars in: Arch. Naturv. Kristian., v4 p.10 | 1887
V., G.O.Sars in: Rep. Voy. Challenger, v.19 pars 55 p.5, 22 | 1893 V. (part.), T. Stebbing
in: Int. sci. Ser., v.74 p.303 | 1895 V., H. J. Hansen in: Ergeb. Plankton-Exp., v.2
G.c. p.55 | 1905 Vauntompsoniidae, Calman in: Sci. Invest. Fish. Ireland, 1904 app. 1
p. 16, and Siboga-Exp., pars 36 p.6, 8 | 1906 V., Calman in: Mt. Stat. Neapel, v.17 p. 414 |
1907 V., Zimmer in: Zool. Anz., v.31 p.368 | 1907 V., Calman in: Tr. zool. Soe.
London, v.181 p.28 | 1908 V., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 164 | 1912 Vaun-
thompsoniidae, T. Stebbing in: Ann. S, Afr. Mus., v.10 p. 133, 134.
Telson wanting. Accessory flagellum of antenna 1 very small. Antenna
2 of Q 2- or 23-jointed. Palp of maxilla 1 with 2 filaments. First 4 pairs
of peraeopods in G, first 3 in Q, with well developed exopods. GC with
5 pairs of pleopods. Uropods with 2-jointed endopod.
3 genera.
1) Some of these occur spasmodically in other families.
2) r., rudiment; a.s., acc. fl, accessory flagellum of antenna 1; mx., maxilla;
m., mandible; prp., peraeopod; mxp., maxilliped.
8) Characters of Chalarostylis 2 assumed from the companion genus Hemi-
lamprops.
4) In these families the 9 is unknown, the characters given are assumed to agree
with those ot the 6.
Cumacea: 1. Vaunthompsoniidae, 1. Vaunthompsonia 9
Synopsis of genera:
Kye present; outer apex of second joint of maxilliped
8 little or not produced. ... . SS Pec. 16
Eye wanting; outer apex of second joint of maxilli-
ped 3 much produced — 2.
{ Pseudorostral lobes not meeting in front of the
2 eyelobe ..-+-.+s52«:++-+-+-.. 2, Gen. Gaussicuma
\ Pseudorostral lobes meeting in front of the eyelobe 3. Gen. Bathyeuma
1. Gen. Vaunthompsonia
1. Gen. Vaunthompsonia Bate
1858 Vaunthompsonia (Sp. un.: V. cristata), Bate MS. in: Kinahan in: Nat.
Hist. Rev., v.5 P. p. 203 | 1858 Vauntompsonia, Bate in: J. R. Dublin Soe., v2 p. 102 |
1859 Vaunthomsonia, Bate in: Ann. nat. Hist., ser. 3 v3 p.273 | 1863 Vaunthompsonia
(part.), A.M. Norman in: Tr. Tyneside Club, v. 51v p.271 | 1879 V.; GaOsSarssin:
Arch. Naturv, Kristian., v.4 p.11 | 1887 V., G. O. Sars in: Rep. Voy. Challenger, v.19
pars 55 p.22| 1905 Vauwntompsonia, Calman in: Sci. Invest. Fish. Ireland, 1904 app. 1
p.16 | 1907 V., Calman in: Tr. zool. Soc. London, v. 181 p. 29 | 1908 V.. Zimmer in:
Ergeb. Tiefsee-Exp., v.8 p. 164 | 1909 V., Zimmer in: Ergeb. Schwed. Siidp.-Exp., v.6
or. 3 p.4 | 1911 Vaunthompsonia, T. Stebbing in: P. Linn. Soc. London, p. 64, 69 | 1912
V., T. Stebbing in: P. Linn. Soc. London, p. 16, 78.
General form not very slender. Pseudorostral lobes nearly meeting in
front of the eyelobe. Eye developed. Third maxilliped with the lateral apex
of the second joint not much or not at all produced.
4 species.
Synopsis of species Q:
1 { Carapace of (young) female not carinate 1. V. inermis
\ Carapace of female carinate — 2.
9 { Carina single, median, straight . 3. V. cristata
Carinae paired, submedian, curved 2. V. meridionalis
Synopsis of species ¢:
First legs reaching more than two joints beyond carapace 3, V. cristata
First legs reaching little more than one joint beyond carapace 4. V. arabica
1. V.inermis Zimmer 1909 V.i., Zimmer in: Ergeb. Schwed. Siidp.-Exp., v. 6
nr.3 p.4 t.1 £.4, 5.
Near to V. meridionalis Sars but inferior in size, and carapace
without submedian carinae and group of denticles. Eyelobe without denticles.
L. © with incipient marsupium about 6°5 mm.
South Georgia, outside Kochtopf Bay. Depth 24—52 m.
2. V. meridionalis O. Sars 1887 V.m., G. O. Sars in: Rep. Voy. Challenger,
v.19 pars 55 p.9, 23 t.2 f.1—5 | 1905 Vawntompsonia m., Calman in: Sci. Invest. Fish.
Ireland, 1904 app.1 p.17 | 1908 V.m. and Vauwntrompsonia (laps.) m., Zimmer in:
Ergeb. Tiefsee-Exp., v.8 p.166 t. 38 f. 34, 45; t. 39 £.46—52 | 1909 Vawntompsonia m.,
Zimmer in: Ergeb. Schwed. Siidp.-Exp., v.6 nr.3 p.4.
©. Pseudorostral lobes reaching little beyond the eyelobe, the sinus
rather deep, antero-lateral corner acute. Carapace in medio-dorsal line as
long as the pedigerous segments, but the serrate lateral margin shorter than
that of those segments; behind the large hidenticulate eyelobe the carapace
is smooth, denticulate to the rear, between these spaces having a pair of
curved denticulate carinae converging backwards. Second pedigerous seg-
ment the longest, the last three produced backward in rounded postero-
10 Cumacea: 1. Vaunthompsoniidae, 1. Vaunthompsonia
lateral lobes. Telsonic segment short, with setules on the little produced
apical margin. Eye brownish. Three -jointed flagellum of first antenna as
long as second joint of peduncle. Second antenna 3-jointed (Zimmer), 2-jointed
(G. O. Sars). Apical points of lower lip minute. Fore part of mandible
short, spine-row of 9 spines. First maxilliped with 4 branchial leaflets in
a row. ‘Third maxilliped with lateral apex of long second joint somewhat
produced. First peraeopod reaching apex of pseudorostrum with end of its
5th joint. Second peraeopod with 34 joint very short but distinct, 4 shorter
than 5% or 7%, which are subequal. Uropods stout, peduncle as long as
telsonic segment, about 4/,%8 as long as the subequal rami, with about
13 spines on the medial margin; 1%* joint of endopod two and a half times
as long as 24, its medial margin fringed with unequal spines; 24 joint of
exopod fringed with short spines on lateral margin and feathered setae on
the medial. Colour white with brown markings, stellate on the carapace.
L. © reaching 12mm. Adult d not known, young 3 65 mm.
Kerguelen Island, off Cumberland Bay, depth 232 m, Gazelle Harbour, depth
5—10 m; South Georgia, Cumberland Bay, depth 250—310 m.
3. V. cristata Bate 1856 Scorpionura vulgaris, J. V. Thompson MS. in: W.
Thompson, Nat, Hist. Ireland, v.4 p. 394 | 1858 Vaunthompsonia cristata, Bate in:
Kinahan in: Nat. Hist. Rev., v5 P. p.203 | 1858 Vauntompsonia c., Bate in: J. R.
Fig. 5. Fig. 6. INES 6
V. cristata (2) Bate, V. cristata (2) Bate, an- V. cristata (2) Bate, front
antenna 1 (after Sars). tenna 1 (after Sars), Flagella (after Sars).
(more magnified).
Dublin Soe., v.2 p. 102 tf. | 1859 Vauwnthompsonia c., Kinahan in: Rep. Brit. Ass., 1858
v.28 p.266 | 1863 V.c., A.M. Norman in: Tr. Tyneside Club, v.51v p.271 | 1879 V.c., with
syn. Bodotria longipes, Norman MS., G. O. Sars in: Arch. Naturv. Kristian., v.4 p.12
t. 23—26 | 1893 V.c., T.Stebbing in: Int. sci. Ser., v.74 p.304 | 1905 Vauntompsonia
c., Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p.17 t.1 f.1| 1907 V.c., Calman
in: Tr. zool. Soe. London, v. 181 p. 29 | 1909 Vaunthompsonia c., A. M. Norman & G. S.
Brady in: Nat. Hist. Tr. Northumb., ser. 2 v.31 p. 27.
Pseudorostral lobes meeting underneath the eyelobe, not extended
beyond it; antero-lateral corners not produced in the G, in the Q acute, the
Cumacea: 1. Vaunthompsoniidae, 1. Vaunthompsonia, 2. Gaussicuma 11
preceding emargination denticulate, the lower border of carapace for some
distance backward dentate. Carapace with median carina, armed in Q,
not in G, with close set double row of forward pointing teeth, interrupted
for a short space before its continuance to the hind margin. Five pedigerous
segments conspicuous, but the 1st in dO laterally covered by the gud seoment.
The strongly produced telsonic segment has its broadly triangular apical
margin denticulate. Eyelobe somewhat linguiform, with a pair of little apical
teeth; eye dark reddish, with several corneal lenses. Antenna 1, 1st joint
longest, 34 in © equal to 24, but shorter in G, flagellum rather long, with
short 3¢ joint, accessory minute, one-jointed. Antenna 2 of Q 2-jointed,
with 2 plumose setae on the 1%* joint, the long-jointed flagellum of J not reaching
far along the pleon. Apical points of lower lip conspicuous. Fore part
of mandible not short, spine-row of 17 spines. First maxilliped with
5 branchial leaflets. Third maxilliped with medial margin of the long
gnd joint serrate, the lateral apex not at all produced. Peraeopod 1 as in
V. meridionalis. Peraeopod 2 with 34 joint distinct (Calman), wanting (Sars),
7th joint longer than the 4 or 5, which are subequal. Pleopods of 3
with broad peduncle, armed on medial margin with 5 coupling spines and
4 plumose setae, branches subequal, a little shorter than peduncle, 1st joint
of exopod without setae, which are numerous on_ its 2nd joint and on the
1-jointed endopod. Uropods rather slender, peduncle as long as telsonic segment,
with a dozen spines on medial margin, endopod with 2"4 joint two thirds of
length of the 1°*, spines on medial margin of 1st 9, of 24, 3 or 4, exopod as long
as peduncle, four-fifths of endopod, with slender spines on both margins
and a very long apical spine. L. O 6 mm, OC 5 mm.
British Isles, W. coast of France, Mediterranean. W. Indies? Surface (¢) to 36m,
4. V. arabica Calm. 1907 Vauntompsonia a., Calman in: Tr. zool. Soc. London,
v. 181 p.2, 29 t.7 f. 20—24.
3. Very near to V. cristata, but with medio-dorsal line of carapace
arched instead of straight, and the pleon stouter, 1‘* antenna with shorter
and stouter peduncle, the third joint longer than the second, third maxilliped
with medial margin of 2"4 joint not serrate, peraeopod 1 reaching end of
carapace only with end of the sixth joint, peraeopod 2 with 7% joint less
slender, and the following peraeopods relatively shorter. Colour variable,
patches usually on carapace, dendritic spots sometimes on sides of pleon.
L. 3:1 mm.
Suez Canal; Aden.
2. Gen. Gaussicuma Zimmer
1907 Gaussicuma (Sp. un.: G. vanhoeffeni), Zimmer in: Zool. Anz., v.31 p. 368 |
1912 G., T. Stebbing in: Ann. S. Afr. Mus., v, 10 p. 134.
General form slender. Pseudorostral lobes not meeting in front of
the apically downbent eyeless eyelobe. Pedigerous segments each with a
ventral tooth. Telsonie segment strongly produced between bases of uropods.
Maxilliped 3 with 2™¢ joint long and distally very much produced.
1 species.
1. G. vanhoeffeni Zimmer 1907 G. v., Zimmer in: Zool. Anz., v.31 p. 368.
Pseudorostral lobes with apices separated from the obtuse antero-lateral
corners by a deep incision. Carapacé with creases on hinder part of back
12 Cumacea: 1. Vaunthompsoniidae, 2. Gaussicuma, 3. Bathycuma
and sides, a median carina doubly denticulate along anterior two-thirds of
its length, with denticulate lateral branches starting at about two-thirds of
the length of the frontal lobe, and two rows of denticles on each of the
pseudorostral lobes, which are also denticulate on front and lower margins.
First pedigerous segment short, the hinder ones narrowing gradually to the
pleon. Each pedigerous segment with a median ventral tooth directed forward
and 2 pairs of denticulate carinae, submedian and lateral; the former con-
tinued to end of pleon, the latter to its 5" segment; the pleon has an
additional pair of lateral carinae, and the long telsonic segment has a
median carina. Eyelobe with front part sharply downbent is devoid of an
eye. Maxilliped 3 with lateral front corner of second joint extraordinarily
produced forward. Pedunele of uropods about as long as telsonic segment;
endopod over half as long as the peduncle and considerably longer than
the exopod, with about 7 spines on medial margin of its 1** joint and some
apical spines on the second. L. © with commencing marsupium about 12 mm.
Antarctic: lat. 65° 15‘ S., long. 80° 34' E., depth 3423 m.
3. Gen. Bathycuma H. J. Hansen
1895 Bathycuma (Sp. un.: B. elongata), H. J. Hansen in: Ergeb. Plankton-Exp.,
v.2G.c. p.55 | 1905 B., Calman in: Sci. Invest. Fish. Ireland, 1904 app. Ppa
1905 B., Calman in: Siboga-Exp., pars 36 p.9 | 1908 B., Zimmer in: Ergeb. Tiefsee-
Exp., v8 p.164—166 | 1912 B., Calman in: P. U.S. Mus., v1 41 p.614 | 1912 ee al
Stebbing in: Ann. S. Afr. Mus., v.10 p. 135.
General form elongate. Pseudorostral lobes meeting in the front.
First pedigerous segment short but well exposed. Telsonie segment produced
between the bases of the uropods. Eye wanting. Mandible with long spine-
row. Palp of maxilla 1 with two filaments. Maxilliped 3 with lateral apex
of 294 joint distally produced, 4 joint little expanded.
5 species.
Synopsis of species:
1 { Pleon segments multicarinate. - . - +++ e+ sss: 1. B. longicaudatum
\ Pleon segments not multicarinate — 2.
9 Peduncle of uropod not longer than the exopod . . . . 2. B. brevirostre
Peduncle of uropod longer than the exopod — 8.
3 J Pseudorostrum long Uy etm Tha Se ear Lea ELS 3. B. longirostre
\ Pseudorostrum not long — 4.
Uropods with 2nd joint of endopod nearly as long as the 1st 4. B, elongatum
4 Uropods with 2n4 joint of endopod not nearly as long as
thre 2St 77. teigen lentes Wenien a wie) Ma Become) ie: eles len Ome 5. B. natalense
1. B. longicaudatum Calm. 1912 B. (2) longicaudata, Calman in: P.U.S. Mus.,
v.41 p.604, 614 tf. 10—13 | 1912 B. longicaudatus, T. Stebbing in: Ann. S. Afr. Mus.,
v.10 p.135. ;
Pseudorostrum short, pointed, slightly upturned, angular sinus defined
by blunt antero-lateral tooth. Carapace medio-dorsally arched, front half
carinate with 2 rows of fine serrations more or less alternating, the carina
beginning on the narrow eyeless eyelobe. Pedigerous segments 3 and 4
each with transverse ridge. The long slender pleon having on the first
5 segments dorsal, dorso-lateral and ventro-lateral pairs of ridges. The
telsonic segment produced as a prominent rounded lobe between the bases
of the uropods. Antenna 1 with 24 and 34 joints subequal, slighter than
the Lt, Maxilliped 3 with lateral apex of slender 24 joint produced nearly
Cumacea: 1, Vaunthompsoniidae, 3. Bathycuma 18
to end of 4 joint. Peraeopod 1 with 3° joint longer than broad, 5‘ and
7h subequal, each shorter than 6". Peraeopod 2 with 5 joint longer than
4th, 7th than 5%, Peduncle of uropods subequal to telsonic segment, pro-
bably longer than endopod. L. 9 immature 18°5 mm.
Off San Diego (California). Depth 1174—1218 m.
2. B. brevirostre (Norm.) 1879 Leucon brevirostris, A. M. Norman in: Ann.
nat. Hist., ser.5 v.3 p.71 | 1896 Vawnthompsonia caeca, J. Bonnier in: Ann. Univ. Lyon,
v.26 p.536 t. 28 f.3a—s | 1905 Bathycuma brevirostris, Calman in: Sci. Invest. Fish.
Ireland, 1904 app.1 p.18 | 1908 B. (Vauntompsonia?) b., Zimmer in: Ergeb. ‘Tiefsee-
Exp., v.8 p. 165, 166 | 1912 B.b., T. Stebbing in: Ann. 8. Afr. Mus., 10 p. 135.
©. Pseudorostrum very short, broad, rounded, edge minutely serrate
(Norman), upper margin horizontal, without serrations (Calman), antero-lateral
angle greatly produced, acute. Carapace once and two thirds as long as
deep, widest behind the middle, medio-dorsal line gently arched, its numerous
denticles in an irregularly double alternating line. Maxilliped 1 in GO with
5 branchial leaflets on the epipod. Maxilliped 3 with row of strong teeth
on lower surface of 2"¢ joint of which the apex reaches end of 4% joint
and carries setae reaching end of 6 joint. Peraeopod 1 long, slender,
gud joint strongly spined, 5 and 7 joints each ?/,48 as long as the 6.
Peraeopod 2 with 4% joint longer than 5%, 7% than 4". Pedunele of uropods
subequal to endopod with 5 spines on medial margin in Q,7 in G, 10 spines
on medial margin of endopod’s 1* joint, 5 on the 2™¢ which is nearly */,“
as long as the 1st; exopod a little longer than the endopod. IL. about 10 mm.
S. of Rockall, lat. 56° 26‘ N., long. 14° 28‘ W., depth 205 m; 77 miles W. N.W.
of Achill Head, county Mayo, Ireland, depth 699 m; Bay of Biscay, depth 350—1710 m;
Mediterranean, depth 950—1100 m,
3. B. longirostre Calm. 1905 B. longirostris, Calman in: Siboga-Exp., pars
36 p.2, 9 t.2 f.13, 14 | 1908 B. (Vauntompsonia ?) l., Zimmer in: Ergeb. Tiefsee-Exp.,
v.8 p.166 | 1912 B.1, T. Stebbing in: Ann. 8. Afr. Mus., v.10 p. 135.
Pseudorostrum long, about '/,'* of length of carapace, horizontal, acute,
the sinus nearly semicircular, antero-lateral angle acute. Carapace nearly
thrice as long as wide, and more than twice as long as deep, its medio-
dorsal line rather convex, sharply carinate, with double row of alternating
teeth, the eyeless eyelobe narrowly wedge-shaped. Fourth pedigerous segment
with sideplates produced forwards as narrowly rounded lobes. Pleon rather
long and stout, smooth, telsonic segment produced as a prominent rounded
lobe between the bases of the uropods. Antenna 1 with last joint of peduncle
slightly swollen, flagellum 2-jointed, 1s* joint twice as long as 294, accessory
2-jointed, very short. Maxilliped 3 with a few teeth distally on the 2"¢ joint,
of which the serrate setose apex reaches end of the 4‘ joint. Peraeopod
1 with 2"¢ joint strongly denticulate, 7™ joint longer than 5, a little shorter
than the 6, Peraeopod 2 with 2™¢ joint very short, 7" long and slender.
Peraeopods 3 and 4 subequal in length, but exopod of 34% equal to that of
gnd and nearly double that of the 4%. Peduncle of uropods subequal in length
to telsonic segment, with 13 unequal spines on the medial margin; exopod
about /,48 as long as the peduncle, with spinules on lateral margin and
long setae on the medial; endopod a very little shorter, its first joint not
much longer but much stouter than the second, each with 9 spines of varying
length on the medial edge. L. O (immature) 12 mm.
Lat. 5° 40'7 S., long. 120° 45'5 KE. (Malayan Archipelago), depth 1158 m.
14 Camacea: 1. Vaunthompsoniidae, 3. Bathyeuma
4. B.elongatum H.J.Hansen 1895 B. elongata, H.J. Hansen in: Ergeb. Plankton-
Exp., v.2G.c p.57 t.6 f.6,6a, b; t.7 f.1, la—e | 1905 B.e., Calman in: Sci. Invest.
Fish. Ireland, 1904 app.1 p.18 | 1908 B.e, Zimmer in: Ergeb.
Tiefsee-Exp., v.8 p. 165, 166 | 1912 B. elongatus, T. Stebbing in:
Ann. 8. Afr. Mus., v.10 p. 135.
Body very elongate. Pseudorostrum acute, a little
depressed, antero-lateral corners acute, lower margin for
some distance serrate. Carapace nearly twice as long as
deep, strongly compressed, with a serrate carina only on
the front third portion. Pedigerous segments and pleon
not carinate, second segment of pleon notably shorter than
1st or 34; telsonic segment produced in semi-oval shape
between bases of uropods. Kye wanting. Antenna 1 with
1st joint dentate, 3¢ longer than 2°4, flagellum 2-jointed,
1st joint rather longer than 2™4, the accessory minute,
iets! but also 2-jointed. Antenna 2 with 4 plumose setae.
B. elongatum Hansen, Mandibles elongate, the basal part shorter than that beyond
mandible. the molar, spine-row of 17 spines. Maxilla 1 with dilated
hook-like end to the longer filament on the palp. Maxilliped 1
broad, a longitudinal series of 8 branchial leaflets on the epipod. Maxilliped 2
extremely slender. Maxilliped 3 with 2"¢ joint twice as long as rest of limb, but
produced apex not quite reaching apex of 4%" joint, which distally carries a
submarginal row of teeth. Peraeopod 2 as described differing from other species
of the genus by absence of the third joint. Peduncle of uropods longer than
Fig. 9.
B. elongatum Hansen. a antenna 1 and 2, & telson and uropods.
the equal stout rami, exopod with 13 spines on lateral margin, 3 at apex, and
7 or 8 plumose setae on medial margin, endopod with 7 unequal spines on
medial margin and apex of 1‘ joint, and on the slightly shorter but much
narrower 24 joint 5 small spines to the medial margin and 2 long on the
apex. Colour clear greyish. L. © (not fully adult) 12°5 mm.
Lat. 18° 6‘ N., long. 26° W. (North Atlantic), depth 4980 m.
5. B. natalense Stebb. 1912 B. natalensis, T. Stebbing in: Ann, S. Afr. Mus.,
v.10 p. 135 t. 49.
Cumacea: 1. Vaunthompsoniidae, 3. Bathycuma, 2.Sympodommatidae, 1.Sympodomma 15
Pseudorostrum acute, slightly upturned, the lobes meeting for a short
distance in front of the little triangular eyeless eyelobe. Carapace about a
quarter of the body’s total length, rough with minute denticles, the medio-
dorsal line carinate, the first 3¢ showing spinules in double line, commencing
on the eyelobe and fading away into a single line towards or to the hind
margin. 1%* pedigerous segment seemingly united to the
carapace. Pleon segments 1—5 with lower borders flat-
tened; telsonic segment produced between bases of uropods
to an almost semicircular partially serrate border. Antenna 1
with 18 joint geniculate, 34 joint longer than 2™4, 1st joint
of the 2-jointed flagellum dilated near the base and fringed
with long filaments, the minute 2-jointed accessory having
a close-set fascicle of very long setae. Mandibles with spine-
row of 21 spines, but basal part of trunk longer than the
distal section. Maxilliped 1 with 8 branchial leaflets in
longitudinal series on the epipod. Maxilliped 3 with apex
of 2"4 joint well produced and serrate on the medial margin. riewie
Peraeopod 2 with short 34 joint, the spinose 7" as long as B natalense Steb-
5t and 6 joints combined. Pleopods with peduncle little bing, mandible.
longer than the subequal setose rami. LExopod of uropod
about */,%s of the length of the spinose peduncle, with 8 spines on the medial
margin, endopod scarcely shorter, with 21 spines on medial margin of the
1st joint, 24 joint much thinner, more than half as long, with a dozen little
spines on the medial margin. L. dO 11 mm.
S. Africa, Cape Natal N. by E, 24 miles, depth 805 m.
2. Fan. Sympodommatidae
1912 Sympodommatidae, T. Stebbing in: Ann.S. Afr. Mus., v.10 p. 131, 133, 138.
Telson wanting. Accessory flagellum of antenna 1 very small. Antenna
2 of © 2-jointed. Palp of maxilla 1 with 2 filaments. First 3 pairs of
peraeopods in G and © with well developed exopods. OG with 5 pairs of
pleopods. Uropods with 2-jomted endopod.
1 genus.
1. Gen. Sympodomma 1. Stebbing
1912 Sympodomma, 'T. Stebbing in: Ann. S. Afr. Mus., v.10 p. 138.
General form slender, elongate, width diminishing gradually from cara-
pace to pleon. Hyelobe narrowly linguiform, separating the pseudorostral
lobes, in which the sinus is well defined by the produced antero-lateral angle.
All five pedigerous segments dorsally exposed, the first short. Pleon elongate;
telsonic segment produced between the bases of the uropods. First antenna
with both flagella slight. Third maxillipeds with second joint distally much
produced, fifth not distally widened. First three pairs of peraeopods in both
sexes with, fourth and fifth pairs without, exopods.
4 species.
Synopsis of species:
1 J Crest of carapace without prominent teeth ........ 1. S. weberi
\ Crest of carapace with prominent teeth — 2.
J Crest of carapace with four prominent teeth ....... 2.8. anomalum
\ Crest of carapace with only three prominent teeth — 3.
16 : Cumacea: 2. Sympodommatidae, 1. Sympodomma
Endopod of uropod longer than exopod, second joint much
Endopod of uropod shorter than exopod, its joints subequal. . 3. 8. diomedeae
3
the shorter’ ‘i. 41st et tteulermes Peukes Vs | si! to) ae/ hem eee nena 4. 8. africanum
1. S. weberi (Calm.) 1905 Heterocuma(?) w., Calman in: Siboga-Exp., pars 36
p.6 t. 1 f.7—12 | 1912 H. w., Calman in: P. U.S. Mus., v.41 p.613 | 1912 Sym-
podomma w., T. Stebbing in: Ann. 8. Afr. Mus., v.10 p. 188.
Pseudorostral lobes slightly upturned, apices bluntly rounded, not reaching
beyond eyelobe. Antero-lateral corners in G obtuse. Carapace short, surface
anteriorly beset with forward directed spiniform points, anterior half with
medio-dorsal carina. Pedigerous segment 1 strap-like, 37 and 4" laterally
articulated, 34 to 5‘ faintly showing the three pairs of ridges, dorsal, dorso-
lateral, and lateral, better developed on the first 5 pleon segments, on the
5th the lateral joining the dorso-lateral at about the middle of the segment.
The telsonic segment with some ridges and a dorsal depression is squarely
produced between the uropods. Eyelobe narrowly linguiform, slightly expanded
at apex, with 2 large but indistinct corneal areas. Antenna 1, 1st and 34
joints longer than 2"¢, flagellum of 3 joints, 1‘t in O dilated at base with brush
of sensory filaments; accessory of 2 joints, 1S* dilated, also with brush of
shorter filaments. Maxilliped 3, 2"¢ joint very much longer than rest of limb,
apically produced to denticulate apex of little produced 4th joint. Peraeopod
1 elongate, 24 joint long, but shorter than rest of limb, 7 joint long, but
shorter than 6%, Peraeopod 2, with 34 joint distinct but very short. Exopod
of peraeopod 3 slightly smaller than that of two preceding pairs. Pleopods large,
rami of 1st pair about two-thirds of length of peduncle. Peduncle of uropods
much longer than exopod, in which the 1%* joint is subequal to 2nd on
endopod, the 2™4 longer than the 1%* of endopod, which is about once
and a half as long as its own 2™¢ joint. L. d 286 mm. Q unknown.
Lat. 5° 40°7 S., long. 120° 45°5 E., (Malayan Archipelago) depth 1158 m.
2. S. anomalum (O. Sars) 1871 Leucon anomalus, G. O. Sars in: Ofv. Ak. Forh.,
v.28 nr.6 p.805 | 1873 L.a., G. O. Sars in: Svenska Ak. Handl, n. ser. v.11 nr. 5 pelt
t.3 f.10—13 | 1879 Vaunthompsonia anomala, G. O. Sars in: Arch. Naturv. Kristian.,
v.4 p.12 | 1887 ?V.a., G. O. Sars in: Rep. Voy. Challenger, v.19 pars 55 p.22 | 1893
2V.a., 'T. Stebbing in: Int. sci. Ser., v. 74 p. 304 | 1912 Sympodomma anomalus, Ts
Stebbing in: Ann. S. Afr. Mus., v.10 p. 138.
Pseudorostral lobes extended along the eyelobe to its extremity; antero-
lateral corners acutely produced; behind these the lower margin. of the cara-
pace for some distance minutely denticulate. Carapace much narrowed in
front; medio-dorsal line carinate, strongly in first half, there carrying 4
strong teeth, 18* behind the eyelobe, 4 a little in advance of the centre.
Pedigerous segment 1 strap-shaped, not overlapped by the larger 2nd seoment.
Pleon segments 1—5 cylindrical. Telsonic segment pentagonal with rounded
apex, the sides between the uropods having each a series of 6 or 7 minute
spinules. Eyelobe narrowly linguiform, slightly expanded at apex and
ending in 2 little divergent teeth much as in Vaunthompsonia cristata (p. 10),
character of eye uncertain. Antenna 1,2°¢ and 34 joints subequal, accessory
flagellum obsolete. Maxilliped 3 as in Leucon. Peraeopod 2 with 3% joint
said to be missing. Peduncle of uropods a little longer than the rami, flat-
tened, medial margin carrying 5 large and 3 little spines; endopod shorter than
exopod, its 18* joint with 10 spines, 2 longer than the rest, gnd joint not
shorter but much narrower with 10 spines. L. 9 juv. about 8mm. G unknown.
West Indies, St. Martin; depth 377—565 m.
Cumacea: 2. Sympodommatidae, 1. Sympodomma 17
3. S. diomedeae (Calm.) 1912 Heterocuma d., Calman in: P. U.S. Mus., v.41
p. 612 tf.6—9 | 1912 Sympodomma d., T. Stebbing in: Ann. 8. Afr. Mus., v.10 p. 138.
Pseudorostral lobes separated by the linguiform eyelobe; an angular an-
tennal sinus formed by blunt moderately projecting antero-lateral angle, followed
by serration of lower margin anteriorly. Carina on frontal lobe surmounted by 3
large forward pointing teeth, to which succeed a pair of adjacent tuberculated
ridges, sides of carapace dotted with rounded tubercles. First pedigerous segment
overlapped by sides of second, this and the 3 following segments having each a
median pair of adjacent ridges, and to these are added on the last three a dorso-
lateral pair. Both pairs are continued on the pleon segments, but become less
distinct towards the last of them. The third and fourth pedigerous segments in
the male have each a ventral spine, found also in the female on the fourth. A
pigmented eye occupies the.apex of the eyelobe. The first peraeopods have the
sixth joint not twice as long as the fifth, and nearly a third longer than the
seventh. In both sexes the first three pairs of peraeopods have well developed
exopods. Peduncle of uropods, fringed with a mixture of long and short
spines, is nearly twice as long as the endopod, of which the 2 subequal spinose
and setose joints are together rather shorter
than the exopod. L. © immature, 14:3 mm.
Off Ose Zaki, Honshu Island, Japan; depth
110—128 m.
4. §. africanum Stebb. 1912 S. africanus,
T. Stebbing in: Ann.S. Afr. Mus., v.10 p. 138 t. 50 (2).
Pseudorostral lobes, eyelobe, and frontal
lobe as in the preceding species, but the 3 large
teeth are succeeded by a single carina, not tuber-
culate, nor have the sides of the carapace f/:) ‘jig 4
tubercles, except a single pair anteriorly outside F. ee Gee
occupied by numerous small lenses. The first
antenna has a geniculate first joint as long as
the subequal second and third joints combined;
the minute 2-jointed accessory flagellum is
shorter than the first jomt of the primary.
First maxilla with elongate palp; second nor-
mally armed, unusually long. Second maxilli-
ped with long and slender second joint. The
long first peraeopods have the sixth joint less
than twice the fifth and not a fourth longer
than the seventh. The short second pair have
the 6% joint not longer than the third, the
seventh as long as fifth and sixth combined.
Exopods on the third peraeopods not clearly
made out. Peduncle of uropods carinate, fringed
the frontal lobe. The pedigerous segments and =f... 0!
pleon are in near agreement also with those of ea og
S. diomedeae, but no ventral spine was found (eo; ee ae
on the third or fourth pedigerous segment. ei | eens
The apex of the eyelobe is pellucid, apparently f° He. 020g
H
i
pedigerous
segments
Fig. 11.
: §. africanum Stebbing, with eye
with unequal spines; the endopod about two- (Fig. 12).
thirds as long as peduncle, a little longer than the
exopod, its first joint about twice as long as the second. L., dO subadult, 18 mm.
South Africa, Cape Point EH. by N. 29 miles.
Das Tierreich. 39. Lief.: T, R. R. Stebbing, Cumacea.
ro
18 Cumacea: 3. Bodotriidae, 1. Cyclaspoides
3. Fam. Bodotriidae
1853 Cumadae (part), T. Bell, Brit. stalk-eyed Crust., p. 321 | 1856 Diastylidae
(part.), Bate in: Ann. nat. Hist., ser.2 v.17 p. 449 | 1863 D. (part), A. M. Norman in:
Tr. Tyneside Club, v.51v p.271 | 1879 Cumidae, G. O. Sars in: Arch. Naturv. Kristian.,
v.3 p. 465, 467 | 1887 C., G. O. Sars in: Rep. Voy. Challenger, v.19 pars 55 p.5, 11 |
1893 C., T. Stebbing in: Int. sci. Ser., v.74 p.301 | 1896 C., J. Bonnier in: Ann. Univ.
Lyon, v. 26 p. 530 | 1900 C., Zimmer in: Fauna arctica, v. 1 p. 414, 431 | 1901 Bodotriidae,
T. Scott in: Rep. Fish. Board Scotl., v.19 p.273 | 1907 B., Calman in: Tr. zool. Soc.
London, v. 181 p.3 | 1908 B., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p.159 | 1910 B., T.
Stebbing in: Ann. S. Afr. Mus, »61v p.409 | 1912 B., T. Stebbing in: Ann. S. Afr.
Mus., v.10 p. 131, 133, 140.
Telson wanting. Accessory flagellum of antenna 1 very small or ob-
solete. Antenna 2 of female 1- or 2-jointed. Palp of maxilla 1 ending in
2 filaments or a single filament. Peraeopods with exopods only on the 1**
pair or also with rudiments on the 2™¢ and 3% pairs in both sexes. Five
pairs of pleopods in the male. Uropods with the endopod 2-joited or simple.
9 genera.
Synopsis of genera:
With fewer than 5 free pedigerous segments — 2.
| With 5 free pedigerous segments — 5.
J With only 2 free pedigerous segments ..... . 1. Gen. Cyclaspoides
\ With only 3 or 4 free pedigerous segments — 3.
J Carapace with lateral cornua. ......... . 2. Gen. Hocuma
2
\ Carapace without lateral cornua — 4.
4 Peraeopod 2 with 34 joint obsolete . . .... .. 3. Gen. Bodotria
\ Peraeopod 2 with 34 joint not obsolete... .. . 4. Gen. Cyclaspis
Without distinct pseudorostral lobes; endopod of
uropod simple, svi. qee +. ook. tees et een eae . 6. Gen. Stephanomma
With pseudorostral lobes; We ndenod of uropod 2-jointed — 6.
With 2 separated branchial siphons. ..... . . 6, Gen. Zygosiphon
With branchial siphou simple — 7.
Second and third peraeopods without exopods. . . 7. Gen. Iphinoé
J
6
\
J : hikes
\ Second and third peraeopods with 1-jointed exopods — 8.
~!
Maxilliped 3, 2nd joint distally produced, 5th Sea
Widenedy isch. Reuss ee ince ner . . . 8 Gen. Heterocuma
Maxilliped 3, 2nd fein ee distally peodueed “pth not
distalliyawid erie dite) venenatis . . . 9. Gen, Cumopsis
1. Gen. Cyclaspoides Bonnier
1896 Cyclaspoides (part.), J. Bonnier in: Ann. Univ. Lyon, v.26 p.530 | 1904 C.,
Calman in: Herdman, Rep. Ceylon Pearl Fish., v.2 p.161 | 1905 C., Calman in: Sci.
Invest. Fish. Ireland, 1904 app.1 p.15 | 1905 C., Calman in: Siboga-Exp., pars 36 p. 6.
Carapace without lateral cornua. First three pedigerous segments fused
with the carapace. Second peraeopod 6-jointed. Other characters as in
Cyelaspis (p. 28).
1 species.
1. C. sarsi Bonnier 1896 C. s., J. Bonnier in: Ann. Univ. Lyon, v.26 p. 530 t. 28
f.la—u | 1904 C. sarsii, Calman in: Herdman, Rep. Ceylon Pearl Fish., v.2 p.161 |
1905 C. sarsi, Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p.15 | 1905 C. s.,
Calman in: Siboga-Exp., pars 36 p. 6.
Cumacea: 3. Bodotriidae, 1. Cyclaspoides, 2. Hocuma 19
Pseudorostrum acute, with antero-lateral corner rounded, denticulate.
Carapace a little compressed laterally in front, to the rear embracing the
first three pedigerous segments, with one dorsal suture or none. Intestine
anteriorly coiled, as in Platycuma holti. Body
without spinose armature. Telsonic segment subequal
in length to the preceding, well produced between
the uropods. Eye wanting. Antenna 1 very short,
its single flagellum consisting of 2 very small joints.
Spine-row of mandible of 15 components. Palp of
maxilla 1 ending in a single filament. Exopod of
maxilliped 1 very elongate. Maxilliped 3 with 2™¢ and ;
4th joints produced along lateral margin of 34 and Fig. 13.
5th respectively. Peraeopod 1 with joints 5—7 sub- C.sarsi Bonnier (¢ juv.).
equal. Peraeopod 2 6-jointed, small. Pleopods 1—5
3 with rami as long as the peduncle. Uropods with peduncle slightly shorter
than exopod, equal to endopod, which has 5 spines on medial margin and
2 apical. L. 5 mm.
Bay of Biscay, depth 950 m; 77 miles W.N.W. of Achill Head, Ireland, depth
698 m; ?lat. 5° 46-7 S., long. 134° 0' E. (Malayan Archipelago), depth 1788 m in
atrial cavity of Ascidian.
2. Gen. Hocuma Marcusen
1894 EHocwma (Sp. un.: E. hilgendorfi), Marcusen in: Hilgendorf in: SB. Ges. Fr.
Berlin, p. 170 | 1894 E., Hilgendorf in: SB. Ges. Fr. Berlin, p.171 | 1903 #., Zimmer
in: Zool. Jahrb., Syst. v.18 p.669 | 1904 #., Calman in: Herdman, Rep. Ceylon Pearl
Fish., v.2 p.160 | 1907 E., Calman in: Bull. Mus. Paris, p.121 | 1907 E., Calman in:
Tr. zool. Soe. London, v. 181 p. 20.
Carapace broad in ©, usually narrow in GO, with projecting lateral
cornua in both sexes. First pedigerous segment and at least sometimes the
second firmly united with the carapace. Antenna 1, accessory flagellum
very small or obsolete. Antenna 2 of © with or without a minute 24 joint.
Peraeopod 1 with 2™¢ joint distally produced acutely forward or inward.
Peraeopod 2 6-jointed. Uropods with peduncle much shorter than the sub-
equal rami, endopod 1-jointed.
For mouth-organs see E. taprobanicum (nr. 2) and E. ferox (nr. 8).
9 species.
Synopsis of species:
Lateral borders of carapace carinate — 2.
1 \ Lateral borders of carapace not carinate — 7.
Carapace of 9 not narrowed to the rear; pedigerous seg-
ment 2 fused, dorsally unsutured. ......... . J. EB. dollfusi
Carapace of 2 narrowed to the rear; pedigerous segment
2 free or dorsally sutured — 3.
Carapace with paired dorsal ridges —}5.
Peduncle of antenna 1 not elongate; eon Zt 5 6
distally narrowed. . . .. . . 2. BH. taprobanicum
Peduncle of antenna lL Blonaate: pleon segment 5 6 not
distally narrowed. .... . ; F ots eee Os sn Lon Sicorne
o*
J
\
3 \ Carapace without paired dorsal ridges — 4.
20 Cumacea: 3. Bodotriidae, 2. Kocuma
Antero-lateral teeth separated from the cornua by a deep
5 COVIby 0) 2-2.) te) pot MO RPE EEA ts Weer dota, oo 4. EK. hilgendorfi
Antero-lateral teeth separated from the cornua by nearly
straight margin — 6.
Peraeopod 1, joint 2 distally produced inwards, 5th more
6 than twice: as Jong as /Uhiees 5) s ) \oteeuton teat eens 5. E. stelliferum
Peraeopod 1, joint 2 distally produced forwards, 5th much
less’ than twice (7k i) ce emrcmeen 0) etre Uo. em Meee 6. E. latum
seudorostral lobes not projecting beyond the cornua. . . 7. E. sarsii
P
‘i Pseudorostral lobes projecting beyond the cornua — 8.
C
g Jarapace of G not nearly twice as long as broad... . 8. EB. ferox
Carapace of G nearly twice as long as broad. .... . 9. E. affine
1. E. dollfusi Calm. 1907 H.d., Calman in: Bull. Mus. Paris, p.119 tf. 2—4.
Pseudorostrum apically emarginate in dorsal view, sides gently convex.
Carapace not much longer than broad, leaving free only 3 pedigerous segments,
the lateral cornua not level with apex of pseudorostrum, lateral margin behind
them carinate, and breadth slightly encreased. Pedigerous segment 2 slightly
detached from carapace at sides only. Pleon slender, smooth like the rest
of the body, telsonic segment shorter than the preceding. LKyelobe small,
without pigment. Peraeopod 1 short, 2"¢ joint longer than the remaining 5
combined. Uropods with rami almost 3 times as long as the peduncle. L. Q
645 mm (specimens not adult).
Coasts of France, Bay of Biscay, Turbelle, in the sand; the Channel, Villers-
sur-Mer, in the sand.
2. E. taprobanicum Calm. 1904 E. taprobanica, Calman in: Herdman, Rep.
Ceylon Pearl Fish., v.2 p.161 t.1 f.1—20; t.2 f.21—28 | 1907 EH. t., Calman in: Tr.
zool. Soc. London, v. 181 p. 20.
Pseudorostrum apically notched in dorsal view, sides sinuous, extending
much beyond the small incurved lateral cornua. From these the greatest
breadth of the carapace diminishes to the base greatly in ©, moderately in C,
the sides sharply inflected, with oblique ridge running from the cornua to
the free margin of the flattened ventral surface; a slight medio-dorsal ridge
is more prominent anteriorly in ©, posteriorly in G. First pedigerous
segment not distinguished from carapace unless by faint dorsal marking,
2nd fused but with complete suture. Pleon much more slender in Q than
in CG, 5 segment longer than 6*, its width in CG abruptly decreasing at
middle. Eye with 3 corneal lenses, larger in G. Antenna 1 with 1%* joint
angularly bent, third longer than 2™4, shorter than 1%‘, flagellum 2-jointed
in Q, 3-jointed in G. Antenna 2 © a rounded nodule bearing 2 plumose
setae. Lower lip with broadly rounded lobes. Maxilla 1 with 2 apical filaments
on palp. Maxilliped 1, exopod normal, epipod with branchial sacs in straight
line, 22 in Q, 33 more elongate in GO. Maxilliped 2 with 3¢ joint distinct
though small. Maxilliped 3, 2™¢ joint greatly produced outside the 34 and
4th, 34 unusually large, in figure longer than the 4%, 4 produced along
lateral margin of the narrow 5%, Peraeopod 1 long, 2™¢ joint abruptly narrowed
distally and apically produced, acutely on the medial edge beyond the third
joint. Peraeopod 2 the shortest, its 3¢ joint suppressed, the 4‘ apically
produced and tipped with a long spine. Peduncle of uropod less than half
as long as the subequal rami, both minutely knobbed at apex, endopod in ©
with a few plumose setae and spinules on medial margin, in O densely set
Cumacea: 3. Bodotriidae, 2. Hocuma 21
with plumose setae on medial margin, as on the peduncle. L. Q subadult
11:1 mm, ¢@ adult 9:3 mm.
Gulf of Manaar. Depth 11—16 m.
3. E. longicorne Calm. 1907 E. longicornis + E. producta, Calman in: Tr. zool.
Soe. London, v.181 p.20 t.6 f.1—6; p.20, 24 t.6 f. 18—20.
Pseudorostrum with rounded apices, sides slightly convex, extending much
beyond the small lateral cornua, which are directed outwards in G, forwards
in immature Q. From these the breadth of carapace diminishes very gradu-
ally backward, the sides clearly carinate, transverse ridge from the cornua
not reaching the ventral margin, medio-dorsal ridge partial and faint. A well
marked suture distinguishes 2™¢ pedigerous segment from carapace. Pleon
segment 5 longer than the following segment, its sides parallel. Kye with
3 corneal lenses. Antenna 1, peduncle long, 3¢ joint much longer than 2"4,
a little longer than 1*, flagellum 3-jointed in G. Peraeopods and uropods
nearly as in &. taprobanicum. L. 7°65 mm adult 3, 46 mm immature Q.
Suez (¢), Penang (9).
4. EB. hilgendorfi Marcusen 1894 H.h., Mareusen in: SB. Ges. Fr. Berlin,
p-170 | 1894 E.h., Hilgendorf in: SB. Ges. Fr. Berlin, p.171 | 1903 H.h., Zimmer
in: Zool. Jahrb., Syst. v.18 p.669 tf. D—H | 1904 H#.h., Calman in: Herdman, Rep.
Ceylon Pear! Fish., v.2 p.161 | 1907 H.h., Calman in: Tr. zool. Soe. London, v.181 p.20.
Pseudorostrum having outwardly directed antero-lateral processes level
with the apices but separated from them by shallow emarginations, much in
advance of the incurved lateral cornua which are separated from them by
deep cavities. Greatest breadth of the much depressed carapace subequal to
the length, with sinuous sides gradually diminishing backwards. A slight
median carina runs from the eye-lobe to past the middle of the telsonic
segment. The frontal lobe ends on each side in a pit-like depression from
which runs a longitudinal carina to the end of the third pedigerous segment.
The first pedigerous segment is dorsally indicated by a line on the carapace,
the 224, though ventrally separate in the Q, is dorsally distinguished only
by a faint suture. Eye with 3 corneal lenses. The 3? maxilliped and
peraeopods 1 and 2 are apparently in near agreement with those of E. tapro-
banicum (nr. 2). Peduncle of uropods only a little over one-third the length
of the subequal rami. L. about 10 mm, specimens perhaps not full-grown.
Enosima (Japan), depth 6—23 m.
5. E. stelliferum Calm. 1907 E. stellifera, Calman in: Tr. zool. Soc. London,
v. 181 p. 20, 23 t.6 f. 13—17.
Pseudorostrum with acute antero-lateral processes on a level with the
obtuse apices but separated from them by 2 small unequal emarginations.
The divergent sides lead straight back to the incurved cornua, and thence
the carinate sides of the carapace slope rapidly backward to the narrow hind
margin. The medio-dorsal carina is only apparent to the rear, and the flanking
dorso-lateral carinae are not strongly marked. Under moderate magnification
the carapace appears covered with stellate clear spots, to which the specific
name refers. Eye showing no lenses or pigment. Antenna 1 with 1%* joint
longer than 2™4 and 34 combined, 3¢ longer than 2"¢, Peraeopod 1 very
long and slender, 2™4 joint with long acute distal process directed obliquely
inwards, this joint not longer than the 5 which is longer than the 6‘ and
22 Cumacea: 3. Bodotriidae, 2. Eocuma
more than twice as long as the 7. Peduncle of uropods little more than
a third of the length of the subequal rami. L. 56 mm. Q immature.
Gulf of Siam, depth 15—28 m.
6. E. latum Calm. 1907 E. lata, Calman in: Tr. zool. Soc. London, v. 181
p. 20, 22 t.6 f.7—12. :
Pseudorostrum with acute antero-lateral processes not quite on a level
with the rounded apices and separated from them by an emargination, the
sides diverging straight from the processes to the acute
forward-directed lateral cornua. From these the carinate
sides of the depressed carapace slope rapidly backward
to the narrrow hind margin. No distinct median carina,
but dorso-lateral carinae from ends of frontal lobe to the
hind margin well marked. Transverse ridge from a little
behind the cornua reaches the free ventral margin of the
carapace. Eye showing no lenses or pigment. Antenna 1
with ist joint longer than 2"¢ and 3¢ combined, 3¢ longer
than 2", Peraeopod 1 not very long, 24 joint the longest,
its distal process directed forwards, 5% and 6% joints
equal, each less than one-third longer than the 7‘ joint.
Uropods as in E. stelliferum. L. 3°9 mm. Q immature.
Gulf of Siam, depth 9—19 m.
7. E. sarsii (Kossm.) 1880 Cyclaspis s., Kossmann, Reise
Roth. Meer., Malacost. pars2 p.88 t.4 f.3 | 1901 Cyclaspoides
cornigera (err., non G. O. Sars 1879), A. O. Walker in: J. Linn.
Soe. London, v. 28 p. 294 | 1904 Hocuma sarsii, Calman in: Herd-
man, Rep. Ceylon Pearl Fish., v.2 p. 162, 166 t.2 f.35—88 | 1907
E. s., Calman in: Bull. Mus. Paris, p.120 tf.5 | 1910 #.s., T.
Stebbing in: Ann. S. Afr. Mus., v.61v p.414; in: Mar. Invest. 8.
Afr., Crustacea, pars 5.
Pseudorostrum with broadly truncate apices level
with or a little in advance of the stout curved lateral
cornua, an intervening rectangular process being separated
from the apices by a shallow emargination and from the
cornua by a semicircular cavity. From the cornua the
rounded sides of the pitted and tuberculate carapace
converge only slightly to the rather broad hind margin,
near to which are a pair of bosses; the median carina
on the front half is succeeded by a submedian pair of
ridges on the hind part. The 2™¢ pedigerous segment is
much lower than the carapace, the 4% and 5% have
reehe each 2 dorsal tubercles. All the segments of the stout
E. sarsii (3) (Koss. Pleon have a prominent medio-dorsal carina. Hye of
mann). three corneal lenses. Peraeopod 1 has the 2™¢ joint about
two-thirds as long as the remaining joints combined and
is apically produced forwards; the 5 joint is a little longer than the
6th and nearly twice as long as the 7%, Peraeopod 2 has no spine on
the apex of the 4t» joint. Peduncle of uropods has two oblique ridges on
the dorsal surface and is about half as long as the rami. L. 9 mm @G.
Red Sea; Gulf of Manaar, depth 13 m; Mediterranean, Cannes, depth 27—86 m.
8. E. ferox (P. Fisch.) 1872 Bodotria f., P. Fischer in: Folin & Périer, Fonds Mer,
v.2 p.47 t.1 f.4 | 1873 B. 7, P. Fischer in: Act. Soc. Linn. Bordeaux, ser. 3 v. 8v p. 424 |
Cumacea: 3, Bodotriidae, 2. Hocuma, 3. Bodotria 23
1878 Cyclaspis cornigera, G. O. Sars in: Arch. Naturv. Kristian., v.3 p. 488 t. 7—9 | 1887
C.c., G. O. Sars in: Rep. Voy. Challenger, v.19 pars 55 p.12 | 1896 Cyclaspoides ferox,
J. Bonnier in: Ann. Univ. Lyon, v.26 p. 533 | 1904 Hocuma f., Calman in: Herdman,
Rep. Ceylon Pearl Fish., v.2 p. 161, 162.
_Pseudorostrum with obtuse apices and antero-lateral angle slightly
expressed in advance of the strong laterally divergent and apically incurved
cornua. Behind these the stiff very spinulose carapace is a little constricted,
and then continued with non-carinate lateral borders to the broadly rounded
hind margin which appears truncate in lateral view. The 2"¢ pedigerous
segment is much lower than the carapace. The whole body is spinulose
in the ©, but only faintly so in the G. The pleon is long and especially
in the © very slender. In the CO the 5 segment is narrowed rather abruptly
in the distal half. Kye with distinct pigment and the corneal lenses (as
figured by G. O. Sars) more than 3. Antenna 1, 3¢ joint as long as 15*,
much longer than 2"4, both flagella 1-jointed, the accessory very small but
distinct. Antenna 2 in © with 18* joint angularly bent, carrying laterally
2 small plumose setae, and succeeded by a minute setuliferous 24 joint;
in G the flagellum reaching beyond the uropods. Mouth-organs in near
agreement with those of 1. taprobanicum (nr. 2), but in maxilliped 3 the 34
joint is much shorter than the 4%. Peraeopod 1, 24 joint large, strong,
spinulose, nearly as long as the 5 remaining joints combined, its forward
extended triangular apex overlapping the 34 joint, 5t* and 6% joints longest
of the 5, subequal, not specially long. The small 6-jointed 2"¢ peraeopod
has a long spine at apex of the 4 joint, in this species the 34 joint being
present and the 7‘ suppressed. Peduncle of uropods a little less than half
as long as the subequal rami, with 3 plumose setae on its medial margin,
7 on that of the endopod in the ©, these margins thickly set with setae
and spines in the G. Colour dusky in Q, lighter in d. L. © 5 mm,
GS 68 mm.
Bay of Biscay, Cap Breton, depth 6°5 m; harbour of Bayonne; Mediterranean,
Goletta, depth 4—9 m.
9. EK. affine Calm. 1904 E. affinis, Calman in: Herdman, Rep. Ceylon Pear! Fish.,
v.2 p.162, 165 t.2 f.29—34.
Only 3 known. The affinity is with 1. taprobanicum (nr. 2), but differing
in the narrower and less depressed form of the carapace, which has the lateral
borders nearly parallel, not carmate. There is a transverse dorsal depression
behind the frontal lobe. Peraeopod 1 has the 2™¢ joint narrower and the
spines on its lower surface more numerous, the 7 joint only half as long
as the sixth. Peraeopod 2 has the spine on the produced 4th joint less
extended. Peduncle of uropods nearly one-third of the length of the rami.
L. adult O 6°7 mm.
Gulf of Manaar, depth 13 m.
3. Gen. Bodotria Goodsir
1828 Cuma (non [Calonne] 1797, Mollusea), H. Milne-Edwards in: Ann. Sci.
nat., v.13 p. 287 | 1831 C., Latreille, Cours Ent., v.1 p.416 | 1840 C., H. Milne-Edwards,
Hist. nat. Crust., v.3 p.553 | 1841 C. (part.), Kréyer in: Naturh. Tidsskr., v. 3v1 p. 508 |
1843 C. (part.) + Bodotria, Goodsir in: Edinb. new phil. J., v.34 p. 120, 127 | 1856 C.-++
B., Huxley in: Quart. J. geol. Soc., v.12 p.35 | 1861 Bodothria, P.J.Beneden in: Mém.
Ac. Belgique, v.33 [nr. 3] p. 78, 76, 174 (Bodotria, p. 166) | 1864 Bodotria, Fr. Miiller,
Fiir Darwin, p.12, 54 f.52 | 1875 Bodothria, R.T. Maitland in: Tijdschr. Nederl. dierk.
24 Cumacea: 8. Bodotriidae, 3. Bodotria
Ver., v.1 p.240 | 1900 Bodotria, T. Stebbing in: Willey, Zool. Results, pars5 p. 610 |
1901 B., T. Scott in: Brit. Ass. Handb. Glasgow, p. 332 | 1907 B., Calman in: Tr. zool.
Soe. London, v.181 p.3 | 1909 B., Stappers in: Cellule, v.25 p. 368 (alimentary system) |
1912 B., T. Stebbing in: Ann. S. Afr. Mus., v 10v p.129, 140.
Carapace without lateral cornua; first pedigerous segment inconspicuous,
the second long. Only the first pair of the peraeopods carrying exopods in
either sex; second peraeopods with the third joint indistinct. Endopod
of uropods simple or two-jomted with second joint always the shorter.
Hepatopancreatic glands with one pair of distally bifurcated lobes.
11 species accepted, 3 doubtful.
Synopsis of species:
{ Uropod with two-jointed endopod — 2.
\ Uropod with endopod simple — 7.
J Hye present — 3.
\ Eye wanting — 6.
Carapace with two pairs cf lateral ridges toe as ke . . I. B. pulchella
Carapace with not more than one pair of lateral Sabees — 4.
Lateral ridges faint on carapace, evanescent beyond ... 2. B. sublevis
Lateral ridges distinct on carapace, continued on free pedi-
gerous segments — 5.
{ Free pedigerous segments with dorsal carina simple
Free pedigerous segments with dorsal carina crested .
\ Uropod with endopod about half the length of the peduncle
J
\
—
bo
Oo
. Scorpioides
. gibba
- montagui
. australis
OL
lor)
ee aes
Hho Ww
Uropod with endopod two-thirds of the length of the peduncle
Free pedigerous segments some or all with lateral ridges — 8.
Free pedigerous segments without lateral ridges — 10.
Carapace with lateral ridges partial and famt ...... 7. B. pulex
Carapace with lateral ridges well developed — 9.
Peraeopod 1 with 2nd joint much longer than rest of limb 8. B. arenosa
9 , Peraeopod 1 with 2nd joint little or not at all longer than
(ct) 0) del bet | Ce eM PN Uy Ao Bars \ oho 18} fibranllls
Pseudorostrum with antero- el tooth acute... .... 10. B. siamensis
10 Pseudorostrum with antero-lateral tooth blunt... .. . 11. B. parva
1. B. pulchella (O. Sars) 1878 & 1879 Cuma p., G. O. Sars in: Arch. Naturv.
Kristian., v3 p.485 t.6; v4 p.124 t.60 | 1890 C.p., T. Scott in: Rep. Fish. Board
Scotl., v8 p.329 | 1899 C. p., T. Scott in: Rep. Fish. Board Scotl., v.17 p. 267 | 1901
C.p., A. O. Walker in: J. Linn. Soe. London, v.28 p. 294 | 1907 Bodotria p., Calman
in: Bull. Mus. Paris, p. 116 tf.1 | 1907 B.p., Calman in: Tr. zool. Soe. London, v.18:
p.-3 | 1909 B. p., A. M. Norman & G.S. Brady in: Nat. Hist. Tr. Northumb., ser.2 v.3
p. 26 | 1912 B. pulchellus, T. Stebbing in: Ann. S. Afr. Mus., v 10v p. 141.
Pseudorostrum very short, obtuse, with a little antero-lateral tooth. The
medio-dorsal line of the carapace depressed near the middle in the G, in
both sexes flanked by 2 pairs of longitudinal carinae, the upper nearly straight
and at least sometimes reaching the hind margin, the lower curved and
distally joining the upper one. Only a median carina on the pedigerous
segments, of which the 2"¢ (1S* free) is much the largest and in the dG raised
much above the rest. Pleon segments with median carina in the Gd, in
which the 5% segment is distally constricted. Eye prominent. Antenna
1 with much curved 1st joint. Maxilliped 3 with distal process of 24 joint
apically truncate. Peraeopod 1 with 5 joint not quite so long as 6" and
7‘. combined. Peraeopod 2 having the slender curved 24 joint armed on the
concave margin with some 6 retroverted teeth. Uropods with rami ind less,
Cumacea: 3. Bodotriidae, 3. Bodotria 25
in Q more, than half as long as the peduncle; endopod 2-jointed, the large
1s* joint armed on serrate, medial margin in CO with 8, in © with 3, strong
spines, the much shorter and narrower 2"¢ only with apical spines.. Colour
dusky in front. L. o 3:2 mm, © 2°5 mm.
Mediterranean, Bay of Naples, depth 9—19m, Cannes, 19m; Bay of Biscay,
Bayonne: Scotland, Forth and Clyde; England, Liverpool Bay.
2. B. sublevis Calm. 1907 B.s., Calman in: Tr. zool. Soc. London, v.181 p.3
t.1 f. L—a.
Pseudorostrum nearly horizontal, antero-lateral angle well developed.
Medio-dorsal line of carapace with a depression some distance from the hind .
margin, the carina continued on the pedigerous segments, but nowhere
prominent, the flanking lateral carinae a single pair, distinct only in its
middle portion, not continued on the 2™¢ pedigerous segment and _ barely
indicated on the 3 following segments. Peraeopod 1, 224 joint a little shorter
than all the following combined, and 5‘ than 6 and 7 combined. Rami
of uropods two-thirds as long as peduncle, endopod with 18* joint nearly
4 times as long as 2™4, 1St with 5 spines on medial margin, 2"¢ with 2 at
apex. Immature CG with smoother carapace, the lateral carinae alone
distinct. L. 2°6 mm.
Gulf of Siam, depth 9 m.
3. B. scorpioides (Montagu) 1804 Cancer s., Montagu in: Tr. Linn. Soe.
London, v.7 p.70 t.6 f£.5 | 1812 Astacus s., Pennant, Brit. Zool., ed. 5 v.4 p.35 | 1818
Diastylis 2s., Say in: J. Ac. Philad., v.11 p.314 | 1825 D. ? scorpionides, A. G. Des-
marest, Consid. gén. Crust., p. 245 | 1843 Cuma edwardsii, Goodsir in: Edinb. new
phil. J., v.84 p.123 t.2 f. 1—13, 18; t.4 f.11| 1846 C. scorpoides, Kroyer in: Naturh.
Tidsskr., ser. 2 v.211 p. 123 (nota) | 1853 C. edwardsii, T. Bell, Brit. stalk-eyed Crust., p. 326
tf. on p. 828 (not those on p.826) | 1856 C. scorpioides, Bate in: Ann. nat. Hist., ser. 2
v.17 p.456 t.14 f.2 | 18638 C.s., A. M. Norman in: Tr. Tyneside Club, v.5iv p.271 |
1878 C. edwardsii, G. O. Sars in: Arch. Naturv. Kristian., v.3 p.470 t. 1—3 | 1883 C.
scorpiotdes, Burmester, Beitr. Cuma, p.3 | 1889 C. edwardsii, Hoek in: Tijdschr. Nederl.
dierk, Ver., ser.2 v.21 p.170, ?C. scorpioides var. | 1893 C. scorpioides, T. Stebbing
in: Int. sci. Ser., v.74 p.302| 1899 & 1900 C. edwardsii, G. O. Sars, Crust. Norway,
v.83 p.12 t.4; C. scorpioides, p.106 | 1901 Bodotria scorpioides, T. Scott in: Rep. Fish.
Board Scotl., v.19 p.273 | 1905 B. s., Calman in: Sei. Invest. Fish. Ireland, 1904 app. 1
p.13 | 1906 B.s. A. M. Norman & T. Scott, Crust. Devon Cornwall, p.29 | 1907 B.s.,
Calman in: Tr. zool. Soc. London, v.181 p.3 | 1909 B.s., Stappers in: Cellule, v, 25
p. 368 t.2 f.19 | 1909 B.s., A. M. Norman & G. S. Brady in: Nat. Hist. Tr. Northumb.,
ser.2 v.3 p. 26.
Pseudorostrum short, apically truncate, antero-lateral tooth short, acute.
Carapace hard, with longitudinal median carina and pair of lateral carinae
continued to the end of the 5 pleon segment, more strongly developed
in pleon of GO than in that of Qg. 2"4(18* free) pedigerous segment shorter
but more strongly carinate in G than in @. Pleon segment 5 in CG
narrowed in distal half. Eye more prominent in GO, pigment dark, corneal
lenses 9. Antenna 1, 1%¢ joint straight, in © not so long as 2"¢ and 3°
combined, 3¢ longer than 2"4. Peraeopod 1, 224 joint longer than all the
following joints Combined, 5 much longer than 6', but not so long as the
subequal 52 and 6% combined. Peraeopod 2, 2" joint longer than the
following 5 joints combined. Rami of uropods subequal, rather longer in ©,
a little shorter in G, than half the peduncle, which in CG has the medial
margin bordered with plumose setae and spines, the endopod having on
26 Cumacea: 3. Bodotriidae, 3. Bodotria
that margin 7 spines in GO, 4 in ©, the much shorter 2"¢ joint being without
spines except 2 at the apex. Colour pale with dark patches. L. 5—6 mm.
British Isles, France, Kattegat, Norway, Mediterranean. Depth, between tide-
marks (Torquay), to 46 m (off Tynemouth).
4. B. gibba (O. Sars) 1878 Cuma g., G.O.Sars in: Arch. Naturv. Kristian.,
v.3 p.483 t.4.5 | 1912 Bodotria gibbus, 'l. Stebbing in: Ann.S. Afr. Mus., v. 10v p. 141.
Pseudorostrum short, obtuse, slightly upturned, antero-lateral tooth well
marked. Integument coarsely squamose. Carapace with depression median
in ©, nearer to the front in GC, longitudinal median carina and flanking
pair of carinae, the latter continued some way along the body, while the median
carina is conspicuously upraised on the 4 free pedigerous segments and
first two of the pleon, the crest on the 2™¢ (first free) pedigerous segment
being very high and acutely tipped in the Q but lower and flat-topped
in the O. Kye distinct. Antennal, 18 joint in © longer than 274 and 34
combined. Peraeopod 1, 24 joint much longer than the rest combined, 5%
a little longer than 6*, 6 than 7, Peraeopod 2, 2™4 joint much longer
than the following 5 combined. Armature and structure of uropods differing
but little from B. scorptoides, but rami in both sexes more than half the
length of the peduncle. Colour dusky with a deeper shade anteriorly. L. Q
3, © 47 mm,
Mediterranean, Goletta, depth 4—9 m.
5. B. montagui Stebb. 1912 B.m., T. Stebbing in: Ann. 8S. Afr. Mus., v. 10v
p. 141 t.51 A @A).
Agreeing with B. scorpioides, except in integument little calcified,
carapace with pair of oblique grooves diverging near the middle backwards
from the central carina, 4% pedigerous segment not separated in dorsal
outline from its neighbours by deep incisions, eye wanting, 3? joint of an-
tenna 1 not longer than 24, fifth joint of peraeopod 1 only moderately longer
than 6, 34 joint of peraeopod 2 faintly indicated. L. Q 4°5 mm.
S. Africa, lat. 32° 53’ 30" S., long. 28° 11‘ 00" EK., depth 75 m.
6. B. australis Stebb. 1912 B. a., T. Stebbing in: Ann.S, Afr. Mus., v. 10v p. 142
t.51B (3B).
Pseudorostral lobes with antero-lateral angle well marked. Carapace
with a pair of infero-lateral carinae presenting a scalloped edge, while
the lower margin is fringed with little bosses. Last 3 pedigerous segments and
first 2 of pleon made distinct by deep depressions. Integument little calcified.
Eye wanting. Antenna 1 with 3¢ joint shorter than 2"¢. Maxilliped 2 having
gnd joint widest distally instead of in proximal half as in B. montagua.
Peraeopod 1 with 6 joint not longer than 7% and both together not so
long as the 5%, Peraeopod 2 with 3¢ joint obsolete. Rami of uropods
*/,48 of the length of the peduncle, exopod scarcely longer than endopod, of
which the 24 joint a little exceeds 7/,4 of the 18. L. 9 3°25 mm.
S. Africa, lat. 329 53' 30" S., long. 28° 11‘ 00" E., depth 75 m.
7. B. pulex (Zimmer) 1903 Cuma p., Zimmer in: Zool. Jahrb., Syst: v. 18vi
p. 666 tf. A—C | 1912 Bodotria p., Calman in: P. U.S. Mus., v.41 p. 609.
Pseudorostrum very short, distally truncate, slightly upturned, antero-
lateral tooth well developed. Integument closely pitted. Carapace with
median carina, continued along the body, the pair of lateral carinae only
Cumacea: 3. Bodotriidae, 3. Bodotria 27
developed on the front part of the carapace and on the free pedigerous
segments. 1¢ pedigerous segment indicated by a suture. Eye well developed.
Antenna 1 and peraeopod 1 short. Rami of uropods subequal, about three-
fourths length of peduncle, endopod 1-jointed, its medial margin serrate,
carrying 5 spines, with a much larger spime at the apex. L. © about
35 mm. G not known.
Enosima (Japan), depth 6—22 m.
8. B. arenosa Goodsir 1843 B. a., Goodsir in: Edinb. new phil. J., v.34 p.120,
138 t.2 £.17; t.3 f.8—13 | 1846 B.a., Kroyer in: Naturh. Tidsskr., ser. 2 v.2m p.210 |
1853 B.a., T. Bell, Brit. stalk-eyed Crust., p. 332 tf. (¢) | 1856 B. a., Bate in: Ann. nat.
Hist., ser.2 v.17 p.465 t.15 f.6 | 1861 Bodothria a., P. J. Beneden in: Mém. Ac.
Belgique, v.33 p.78 | 1866 Cuma pusilla, G. O. Sars in: Nyt Mag. Naturv., v.15 p. 108 |
1867 Bodotria arenosa, Marcusen in: Arch. Naturg., v.331 p.357 | 1873 B.a., P. Fischer
in: Act. Soe. Linn. Bordeaux, ser. 3 v.8v p.425 | 1899 & 1900 Cuma scorpioides + C.
arenosa, G. O. Sars, Crust. Norway, v.3 p. 10 t. 1—8; p. 106 | 1901 Bodotria a., T. Scott
in: Rep. Fish. Board Scotl., v.19 p.273 | 1909 B.a., A.M. Norman & G.S. Brady in:
Nat. Hist. Tr. Northumb., ser. 2 v.3u p.25 | 1912 B. arenosus, T. Stebbing in: Ann. S.
Afr. Mus., v. 10v p. 141.
Pseudorostrum very short, truncate, antero-lateral tooth acute. Carapace
with median carina from the eyelobe to the hind margin and continued
along the body to the end of the 5 pleon segment; a pair of lateral
carinae traverse the carapace and free pedigerous segments, but are only
faintly indicated on the pleon. 2™¢ (15t free) pedigerous segment the longest.
Eye composed of 8 corneal lenses. Antenna 1, 18° joint longer than 24
and 34 combined, 3¢ a little longer than 2™4. Antenna 2 Gt, flagellum
reaching end of the body. Peraeopod 1, 24 joint much longer than the rest
combined. Peraeopod 2, 24 joint longer than the rest (4 in number)
combined. Rami of uropods a little under half the length of the peduncle,
which has its medial margin in the G fringed with plumose setae and
spines; the 1-jointed endopod in both sexes has 3 spines on the medial
margin and 2 at the apex. L. 7 mm. — Fig.1 p. 2.
Scotland; S. and S. W. Norway; 30 miles off Sunderland, depth 84 m.
9. B. similis Calm. 1907 B.s., Calman in: Tr. zool. Soc. London, v. 181 p.4
t.1 £.4—9.
Similar to B. avenosa, pseudorostrum and median carina as in that
species. Pair of lateral carinae on carapace and free pedigerous segments
except the last in the ©, with a second lower pair of lateral carinae on the
carapace of the G but not of the @. Peraeopod 1, 2"¢ joint a little shorter
in ©, a little longer in G, than the rest of the joints combined. Rami
of uropods subequal in G, two-thirds as long as peduncle, exopod about
three-fourths of peduncle in ©, rather longer than the 1-jointed endopod,
which has six spines on medial margin in Q, 12 in CO. L. @ 2:15,
SO 2:8 mm.
Gulf of Siam, depth 9—18 m.
10. B. siamensis Calm. 1907 B.s., Calman in: Tr. zool. Soc. London, v. 181
p-5 t.1 f. 10—15.
Near to B. similis. Pseudorostrum very short and blunt, antero-lateral
tooth almost spiniform. Median carina of carapace and pedigerous segments
not very prominent, pair of lateral carinae faint on front part of carapace
28 Cumacea: 3. Bodotriidae, 3. Bodotria, 4. Cyclaspis
and not reaching the end of it. Peraeopod 1, 24 joint about four-fifths as
long as the rest combined. Rami of uropods subequal, five-sixths in 9, about
three-fourths in G, as long as the peduncle, the 1-jointed endopod with
9 or 10 spines on medial margin. A transverse dark stripe precedes the gud
(1st free) pedigerous segment. L. Q 2:1 mm, 0 2:75 mm.
Gulf of Siam, depth 9—18 m.
11. B. parva Calm. 1907 B. p., Calman in: Tr. zool. Soc. London, v.181 p.
t.1 f.16—18 | 1912 B. parvus, T. Stebbing in: Ann. S. Afr. Mus., v 10v p. 141.
Pseudorostrum very short, blunt, antero-lateral tooth rounded. Carapace
having near the hinder end a depression on the dorsal edge bounded by a slight
crescentic transverse ridge, and also a pair of lateral carinae faintly marked,
not reaching the hind margin nor joining the transverse ridge. Free pedi-
eerous segments without carinae. Peraeopod 1, 24 joint a little shorter than
the other joints combined. Rami of uropods subequal, rather less than two-
thirds as long as the peduncle; endopod 1-jointed with small spine near
middle of the medial margin, which then becomes coarsely serrate, the apex
carrying 2 very unequal spines. L. 9 15 mm, S not known.
Gulf of Siam, depth 9 m.
B. ? audouinii (M.-E.) 1828 Cuma a., H. Milne-Edwards in: Ann. Sci. nat.,
v.13 p.294 t.13B f.1—7 | 1840 C. a., H. Milne-Edwards, Hist. nat. Crust., v.3 p. 553 |
1840 ©. a., H. Lueas in: Hist. An. artic., p. 268 | 1843 C. a., Goodsir in: Edinb. new phil.
J., v.34 p.124 t.2 f.14—16; t.4 f.12 | 1899 C. scorpioides (pro: B. arenosa), G10!
Sars, Crust. Norway, v.3 p.10, 11.
Described as having 2 eyes, short 2nd antennae, 5 pairs of natatory feet, colour
yellowish white, length about 7 mm; figures show carapace without lateral ridges,
pleon without pleopods, length 12°5 mm.
B. concava D. Roberts. 1868 B.c., D. Robertson in: P. Nat. Hist. Soc. Glasgow,
v.11 p.69 (deser. nulla).
B.? sp. Calm. 1912 B.? sp., Calman in: P. U.S. Mus., v.41 p. 609.
Q. Carapace and pedigerous segments with lateral ridge. First peraeopods
with fifth joint expanded, about one and one half times as broad as the fourth.
Honshu Island (Japan), surface.
4. Gen. Cyclaspis G. 0. Sars
1865 Cyclapis (err. pro: Cyclaspis) (Sp. un.: Cyclaspis longicaudata), G. O. Sars
in: Forh. Selsk. Christian., 1864 p.206, 207 | 1878 Cyclaspis (part.), G.O. Sars in:
Arch. Naturv. Kristian., v. 3 p. 487 | 1887 C., G. O. Sars in: Rep. Voy. Challenger,
v.19 pars 55 p.11 | 1893 C. (part.), T. Stebbing in: Int. sci. Ser., v.74 p. 302, 303 | 1895
Cyclapsis (err. pro: Cyclaspis), H. J. Hansen in: Ergeb. Plankton-Exp., 2 G.c. p. 51”
1899 Cyclaspis, G. O. Sars, Crust. Norway, v.38 p.15 | 1904 C, Calman in: Herdman,
Rep. Ceylon Pearl Fish., v.2 p. 167 | 1905 C., Calman in: Siboga-Exp., pars 36 p.5 |
1907 C., Calman in: Tr. zool. Soc. London, v.181 p.6 | 1908 C., Zimmer in: Ergeb.
Tiefsee-Exp., v.8 p.160 | 1910 C., T. Stebbing in: Ann. S. Afr. Mus., 0. 61v p. 413.
Carapace without lateral cornua; first pedigerous segment indistinct
except sometimes in adult Q, the second not long. Maxilla 1 with palp ending
in 1 or 2 filaments. Only the first pair of the peraeopods carrying exopods
in either sex; second peraeopods with the third joint distinct. Endopod of
uropods simple.
25 species.
1
2
i
Or
~]
10
ll
12
13
14
15
16
|
|
18 |
|
{rai
Jumacea: 3. Bodotriidae, 4. Cyclaspis
Synopsis of species:
Kye wanting — 2.
Eye present — 5.
Pedigerous segment 2 forming raised border to the
carapace; telsonic segment much shorter than
preceding segment .. . 5 ie
Pedigerous segment 2 not orsiiys meal border te
the carapace; telsonic segment not much shorter
than preceding segment — 8.
Carapace of Q almost globose; apex of eyelobe
acute — 4.
Pseudorostral lobes meeting well in front of eyelobe 3.
Pseudorostral lobes barely meeting in front of eyelobe 4.
C
C
arapace smooth!) — 6.
arapace sculptured or dentate — 12.
Carapace of 9 subcylindrical; eyelobe parallel-sided 2.
J
\
Peduncle of uropods much longer than rami... 0.
Pedunele of uropods not much longer than rami — 7.
J Peraeopod 1 with apical process to 2d joint — 8.
| Peraeopod 1 without apical process to 2nd joint — 9.
Peraeopod 1 long, with 6th joint much longer than 5th 6.
Peraeopod 1 not long, with 6th joint not much ee
than 5th . :
Peraeopod 1 with ond Port cada to Feel of
limb — 10.
Peraeopod 1 with 2nd joint not subequal to rest of
limb — 11.
Peraeopod 1 with setae on apex of 2nd joint .. 8.
\ Peraeopod 1 without setae on apex of 2nd joint . 9.
Peraeopod 1 with 2n4 joint longer than rest of limb 10.
Peraeopod 1 with 2nd joint shorter than rest of limb 11.
J
\
Ridges enclosing quadri-lateral area on each side
of carapace — 183.
Ridges not enclosing quadri-lateral areas on cara-
pace — 16.
Two strong transverse ridges on back of carapace — 14.
No transverse ridges on back of carapace — 15.
Hinder transverse ridge with 2 procurved teeth . 12.
Hinder transverse ridge without teeth. .... . 13.
Lateral quadrangles with 4 prominent tubercles . 14.
Lateral quadrangles with only 1 prominent tubercle 15.
Dorsal carina of carapace with forward pointing
tooth near front — 17.
Dorsal carina of carapace unarmed — 18.
Eyelobe not enclosed by pseudo-lateral lobes — 19.
Eyelobe enclosed by pseudo-lateral lobes — 20.
ae C. varians sometimes denticulate.
1
Carapace with oblique ridge on either side. . . . 16.
Carapace without lateral ridges... . : Mee
Carapace encircled by collar-like ridge ... . 18.
Carapace without collar-like ridge — 21.
Pedigerous segment 5 without median carina. . . 19.
Pedigerous segment 5 with median carina — 22.
(oMohne He)
aaae
. glacialis
. sibogae
. longicaudata
spectabilis
. picta
. hornelli
. herdmani
levis
. varians
. pusilla
. longipes
. perseulpta
. exsculpta
. elegans
. similis
uniplicata
. unicornis
. cingulata
. thomsoni
30- Jumacea: 3. Bodotriidae, 4. Cyclaspis
Three carinae on last 2 pedigerous segments and
91 firste@s of pleon 24% 2) qemiemie oo ty eee 20. C. australis
A single carina from last pedigerous segment to end
Of pleon.2). J Leese ened 3a 21. C. quadrituberculata
Pseudorostral lobes barely meeting in front of
99 eyelobes =). e. 2) mle eleanor te ole aun 22. C. gigas
| Pseudorostral lobes meeting well in front of eye-
lobe — 23.
93 J Carapace with several longitudinal ribs .... . 23. C. costata
“° | Carapace without such ribs — 24.
OA J Carapace with 2 pairs of obliquely transverse ridges 24. C. bistriata
\ Carapace with 3 pairs of transverse ridges... . 25. C. triplicata
1. C. glacialis H. J. Hansen 1908 C.g., H. J. Hansen in: Résult. Voy. Belgica,
Schiz. Cum. p.15 t.3 f. la—g.
Pseudorostral lobes only just touching in front of the eyelobe; antero-
lateral corners each forming a subacute tooth. Carapace with a rounded longi-
tudinal median carina, broad for the first three-fifths of the length, this part
surmounted by a narrow secondary carina and flanked by a pair of hollows,
each occupied centrally by an oblong arched area. Pedigerous segment
2 constituting raised hind margin of the carapace, the abruptly lower 3¢ and
Ath not carinate, the 5 with carina. Pleon with narrowly rounded carina,
gradually vanishing on the} telsonic segment, which is decidedly shorter than
the 5t, Kyelobe rounded in front, then constricted, without corneal lenses.
Maxilliped 3, 2™4 joint broader at base than distally, 4™ distally serrated on
medial side, 7% as long as 6%, Peraeopod 1, 2%¢ joint rather longer than
rest of limb, markedly tapering in distal third of its length, 5 joint as
long as 6%, twice as long as 7%. Peduncle of uropods equal in length to
the rami. Colour yellow with rounded white spots. L. Q without marsupium
10°3 mm, ¢ unknown.
Lat. 719 14’ S., long. 80° 14‘ W. (Antarctic).
2. C. sibogae Calm. 1905 C. s., Calman in: Siboga-Exp., pars 36 p.4 t. 1 f. 4—6 |
1907 C. s., Calman in: Tr. zool. Soc. London, v. 181 p.7.
Pseudorostral lobes short, truncate, slightly upturned, not meeting in
front of the eyelobe; antero-lateral tooth projecting a little in front of the
apices. Carapace subcylindrical, with medio-dorsal keel, evanescent to the
rear, flanked by a low pair of rounded ridges between two strong transverse
ridges, the anterior running nearly vertically down on each side, where it forms
an obtuse tooth with the lateral curve of the second ridge. A third ridge
curves towards the second without reaching it. Pedigerous segment 2 is
distinct, with rounded dorsal crest; 5% segment the longest. Pleon segments
subcylindrical, with 2 ridges on each side converging distally, the lateral
articular processes very slight, telsonic segment little shorter than the. fifth.
Eyelobe long and narrow, extending to the apices of the pseudorostrum,
devoid of corneal lenses. Peraeopod 1, 2"4 joint considerably longer than
rest of limb, 34 and 4 combined little longer than 5‘, all three little longer
than 6t, which is little longer than the 7*. Peduncle of uropods slender,
rami broken. L © subadult 16°5 mm.
Lat. 0° 511 S., long. 129° 48"8 K., depth 411 m.
3. C. longicaudata O. Sars 1865 C.1., G.O. Sars in: Forh. Selsk. Christian.,
1864 p. 207 | 1876 C.1., A. M. Norman in: Jeffreys in: P. R. Soc. London, v.25 p. 214 |
1879 Cyclapis l, A.M. Norman in: Ann. nat. Hist., ser.5 0.3 p.73 | 1896 Cyclaspis l.,
Cumacea: 3. Bodotriidae, 4. Cyclaspis 31
J. Bonnier in: Ann. Univ. Lyon, v.26 p. 534 t. 28 f. 2a—v | 1899 C.1., G. O. Sars, Crust.
Norway, v.3 p. 16 t. 7,8 | 1900 C.1., Zimmer in: Fauna arctica, v1 p.414 (additional
references) | 1905 C.J., Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p.14 | 1912
C.1., Calman in: P. U.S. Mus., v.41 p. 609.
Pseudorostral lobes meeting for a short space in front of the minute
acute eyelobe, obliquely truncate; antero-lateral tooth well marked in Q only,
in both sexes about level with pseudorostral apices. Carapace quite smooth,
almost globose in ©, less wide and deep in G. Pedigerous segment 2 well
developed, segments 3—5 narrowing abruptly. Pleon long and especially
in © very slender, telsonic segment distally widened, little shorter than the
5th segment, which in GC is distally narrowed. Eye wanting. Antenna 1 with
very long sensory filaments on the flagellum and another group in the ¢ at
apex of the peduncle. Antenna 2 of © 2-jointed, with 2 plumose setae, the
flagellum of GO not reaching end of the body. Mandible with numerous
spines in spine-row. Maxilla | with single filament at apex of palp. Mavxilliped
1 with several branchial sacs in straight series. Mavxilliped 2, 2"¢ joint large,
distally denticulate. Maxilliped 3, 2%¢ joint very large, strongly bent, produced
beyond the 4, Peraeopod 1, 2™¢ joint not longer than rest of limb, very
broad except at narrowly tapering distal part, of the others the 6™ joint is the
longest. Peraeopod 2 longer than any succeeding pair. Pleopods of 0 with
interlocking process on lateral margin of endopod. Peduncle of uropods
about half in Q and two-thirds in d the length of the subequal rami, endopod
in GC fringed with dense row of ciliated spinules. Colour pure white. L. ©
8mm, ¢ 7 mm.
N. W. and N. KE. Atlantic; N. Norway; W. Ireland; Bay of Biscay; Mediterranean.
Depth 120—8285 m.
4. C. spectabilis Zimmer 1908 C. s., Zimmer in: Ergeb. Tiefsee-Exp., v8
p. 161 t. 36 f. 1—16.
Pseudorostral lobes meeting only in a point in front of the acute eyelobe,
apically blunt; antero-lateral tooth not quite level with the apices. Carapace
almost globose, but with a slightly angled transverse carina behind the frontal
area; pedigerous segments 2 and 3 almost continuing the curves of the carapace
in dorsal view, segments 4 and 5 abruptly narrowed. Telsonic segment scarcely
shorter than the preceding segment. Eyelobe longer than broad with narrow
apex. Kye wanting. Appendages in near agreement with C. donyicaudata,
but maxilliped 2 has the 2"4 joint less robust, not distally denticulate, mavxilli-
ped 3 has the 2°¢ joint not produced to end of the 4%. Peduncle of uropods
a little over half the length of the exopod which is slightly longer than the
endopod. Colour white, pellucid. L. © about 12 mm.
Lat. 35° 9‘ S., long. 18° 32‘ EK. (South Atlantic), depth 565 m.
5. C. picta Calm. 1904 C. p., Calman in: Herdman, Rep. Ceylon Pearl Fish.,
v.2 p.170 t.3 f.54, 55 | 1907 C. p., Calman in: Tr. zool. Soc. London, v. 181 p. 6.
Pseudorostrum prominent, acute, lobes contiguous well in advance of the
eyelobe; antero-lateral corners sharply produced. Carapace smooth, some-
what compressed with median carina becoming fainter to the rear. Pedi-
gerous segment 1 marked by faint suture. Pleon segments subcylindrical,
with lateral articular processes well marked; telsonic segment not much
longer than broad, shorter than the 5" segment. Eye pigmented, corneal
lenses indistinct. Peduncle of uropods more than twice as long as the sub-
equal rami, endopod rather broad, its apex not acute. L. © subadult 3 mm.
Gulf of Manaar, depth 13 m.
32 Cumacea: 3. Bodotriidae, 4. Cyclaspis
6. C. hornelli Calm. 1904 C. h., Calman in: Herdman, Rep. Ceylon Pearl
Fish., v.2 p. 172 t.4.f. 67—71 | 1907 C.h., Calman in: Tr. zool. Soc. London, v. 181 p.6.
Closely resembling C. herdmani (nr. 7), but carapace still smoother,
dorsal keel simple, feebler, peraeopod 1 much longer, the apically produced
2nd joint about three-fourths as long as rest of limb, 5 half and 7" three-
fourths as long as 6", in peraeopod 5 the 4% joint only two-thirds as long
as the 5‘; peduncle of uropods a little longer than the exopod, which is
slightly longer than the endopod; the latter with 14 spines on medial margin
of SG, 6 on upper part of that margin in Oo. L. Q 53 mm, co 42 mm.
Gulf of Manaar, depth 13—17 m.
7. C. herdmani Calm. 1904 C.h., Calman in: Herdman, Rep. Ceylon Pearl
Fish., v.2 p.171 t.3 f.56—59; t.4 f£.60—66 | 1907 C.h., Calman in: Tr. zool. Soc.
London, v. 181 p. 6.
Pseudorostrum truncate, eyelobe reaching quite to the tip; antero-lateral
tooth subacute. Carapace almost smooth in dG, in Q with median keel
anteriorly between 2 shallow depressions, followed by feebler double keel
starting from a shallow pit. Pedigerous segment 1 exposed in adult Q.
Pleon segments subcylindrical with lateral articular processes. Eyelobe rather
prominent, not longer than broad, corneal lenses indistinctly defined in ©,
large and conspicuous in G. Antenna 1, 1%* joint longer than 2"4 and third
combined, 2™4 a little shorter than 34, flagellum 2-jointed. Antenna 2 of
Q 2-jointed. Peraeopod 1 not elongate, 274 jot as long as rest of limb,
apically produced into a short pointed process as in C. hornelli, 5%, 6%,
and 7th joints subequal, each in © little longer than the 4", but more ex-
tended in G. Peraeopod 2 with 5‘ joint little more than half as long as
the 4th, to which it is subequal in the following limbs. Pedunele of uropod
as long as the subequal rami, which have acute apices without terminal
spines, endopod in © with some 6 spinules along middle of medial margin,
in C rather shorter than exopod, fringed with about 23 spinules decreasing
in size successively and leaving bare the last third of the margin. L. 9
44 mm, 3 4:3 mm.
Gulf of Manaar, depth 13 m.
8. C. levis G. M. Thoms. 1892 C.1., G. M. Thomson in: J. Linn. Soe. London,
v.24 p.264 t.16 f.1—6; t.17 f.7—26 | 1893 C.1, T.Stebbing in: Int. sci.Ser., v.74
p. 303 | 1899 C. laevis, G. O. Sars, Crust. Norway, v.3 p. 16 | 1904 C. levis, Calman in:
Herdman, Rep. Ceylon Pearl Fish., v.2 p.170, 172 | 1907 C. 1. (Thomson?), Calman in:
Tr. zool. Soe. London, v.181 p.6,8,17 t.5 f.6—8 | 1908 C. 1, Zimmer in: Ergeb.
Tiefsee-Exp., v.8 p. 161.
Pseudorostral lobes with apices subobtuse, meeting for a short distance in
front of the hardly visible eyelobe (eyelobe running quite to tip of pseudo-
rostrum, Calman). Carapace described as smooth, destitute of ridges, yet
keeled above, and with a short feeble keel running back from the antero-
lateral tooth (Thomson fig., Calman text). Only the last 4 pedigerous seg-
ments exposed behind the carapace (1** exposed at the sides in adult ©,
Jalman). A dorsal keel and slight lateral keels traversing the pleon, the
segments of which have lateral notches behind and interlocking knobs in
front. Eye not made out (with corneal facets, Calman). Antenna 1, 1% joint
much larger than 24 and 34 combined, 24 notably larger than 3¢ (274 and
34 equal, Calman). Antenna 2 of © 2-jointed. Maxilla 1, the palp ending
in 2 long filaments. Maxilliped 3 with apical process of 2™¢ joint not quite
Cumacea: 3. Bodotriidae, 4. Cyclaspis 33
reaching produced apex of 4‘ joint (no doubt by error regarded as the 3°).
Peraeopod 1, 2"¢ joint (without the long slender apical process) as long as
rest of limb (the supposed apical process, unique in length, really an acci-
dental meeting of 2 long apical setae, Calman), 5‘ and 7 joints subequal,
each shorter than the 6, Pleopods of SG with apically serrate coupling
spines on medial margin of peduncle, 5‘ pair somewhat smaller than the
others. Peduncle of uropods subequal to the exopod which is very slightly
longer than the apically acute endopod; the latter in the male has the
medial margin fringed above with setae, below with spines. L. 7—8 mm.
New Zealand, surface to 15 m.
9. C. varians Calm. 1912 C.v., Calman in: P.U.S.Mus., v.41 p.610 tf. 1—5.
Pseudorostral lobes meeting in front of slightly prominent eyelobe,
truncate more obliquely in © than in G, forming a sinus with antero-
lateral tooth sharper in the Q than in theo. Carapace smooth except for
longitudinal median keel in Q which is sometimes anteriorly denticulate be-
hind the eyelobe. First pedigerous segment exposed. First five pleon seg-
ments with lateral articular processes in the ©. Eye pigmented. First
antennae with 34 joint longer than 2™4, Peraeopod 1 short, 2™! joint sub-
equal to rest of limb, without apical process or apical setae, 7" joint rather
shorter than 5t* or 6t®, Peduncle of uropods in © once and a half, in d
nearly twice as long as the telsonic segment. Rami in Q subequal, more
than two-thirds as long as the peduncle, endopod with medial margin serra-
ted, carrying 4 spines, and an apical spine; endopod in GC less than two-
thirds as long as the peduncle, a little shorter than the exopod, with about
10 spines on medial margin and an apical spine. L. Q 3-4 mm, G 43 mm.
U.S. A., Vineyard Sound, Wood’s Hole; surface.
10. C. pusilla O. Sars 1887 C. p., G.O. Sars in: Rep. Voy. Challenger, v.19
pars 55 p.19 t.1 £.21—23 | 1902 C. argus, Zimmer in: Zool. Jahrb., Syst. v.17 p. 444
tf. A--C | 1903 C.a., Zimmer in: Zool. Jahrb., Syst. v.18 p.668 | 1907 C. pusilla,
Calman in: Tr. zool. Soc. London, v.181 p. 6.
© (C. p.) Pseudorostral lobes very short and obtuse; antero-lateral
tooth well marked. Carapace large and almost globular, laterally smooth,
dorsal line evenly arched. Pedigerous segment 1 not exposed, side plates of
2nd produced far forwards, 4 and 5 not higher or thicker than the slender
cylindrical segments of the pleon, which are smooth except for the slight median
keel continued from the anterior body. Eyelobe constricted at base, the
rounded tumid terminal part provided with 7 corneal lenses, frontal pair very
small. Pedunele of uropods rather longer than the subequal rami. Colour
whitish, with brownish pink patches chiefly antero-lateral. L. Q adult
35 mm. CO not known, unless it be C. argus.
Flinders Passage, between Australia and New Guinea, depth 13 m.
3 (C. argus). Differs from female by greater size, carapace not glo-
bose, antero-lateral tooth more obtuse, eyelobe not constricted at base and
carrying 9 lenses, of which the lateral pairs are very small as well as the
frontal. Peraeopod 1 (imperfect in ©), 2™4 joint longer than rest of limb,
in which all the joints are rather short. Colour yellowish white. L. about
6 mm.
Australia, Plenty Bay.
Das Tierreich. 39. Lief.: T. R. R. Stebbing, Cumacea. 3
34 Cumacea: 3. Bodotriidae, 4. Cyclaspis
11. C. longipes Calm. 1907 C.1., Calman in: Tr. zoo]. Soe. London, v. 18%
p.6,7 +5 f.1—5.
Pseudorostral lobes very short, eyelobe reaching quite to the tip (over-
lying without separating them?); antero-lateral tooth almost spiniform in ©,
less so in G. Carapace of © slightly compressed, not globose, with dorsal
edge almost level, feebly carinate. First pedigerous segment well exposed
in ©, very slightly in dG, second slightly carinate, sideplates well produced
forwards. Pleon segments subcylindrical, with lateral articular processes.
Prominent eyelobe provided with numerous corneal lenses, larger in CG than
in ©, frontal pair seemingly very small. Antenna 1 rather long, 32? joint
longer than 2™4, Peraeopod 1 very long and slender, 2"¢ joint little more
than half as long as rest of limb, not apically produced, 7‘ joint nearly half as
long as 2™4, longer than 5*, shorter than 6%, Peduncle of uropods longer than
the rami, medial margin serrate in Q, setose in; endopod in © three-
fourths as long as exopod, with spines on distal half of medial margin, in
S only a little shorter than the exopod and fringed with 10 spines. L. ©
331 anim, God mm:
Danish West Indies, Cruz Bay, St. Jan.
12. C. persculpta Calm. 1905 C. p., Calman in: Siboga-Exp., pars 36 p.3 t.L
f.1—8 | 1907 C. p., Calman in: Tr. zool. Soc. London, v. 181 p. 6, 16.
Very near to C. exsculpta (nr. 13). Pseudorostral lobes apically ob-
liquely truncate with shallow emargination, the elongate narrow eyelobe over
lying them to the apex; antero-lateral tooth well marked. Carapace strongly
sculptured; a somewhat sinuous carina crossing the frontal lobe not far from
the base of the eyelobe runs out into a prominent tooth on either side,
marking the greatest breadth, its downward continuation meeting a straight
horizontal ridge continued far backwards; a little behind the middle of the
length of the carapace a second transverse carina on either side of a
median notch rises into a laminar tooth with sharp apex bending forward
and downward; laterally to this pair of hooks the carina is continued obliquely
downwards and forwards on each side to join the longitudinal ridge
which separates a dorsal depressed area from a lateral one, forming a tooth
at the junction, from which a vertical ridge runs downwards as hind margin
of the lateral area; to the rear two successive ridges run down to join the
horizontal ridge above mentioned; there is a median dorsal keel, which be-
hind the second transverse carina is elevated and undulating, forming three
lobes; subparallel with this part there are a pair of longitudinal carinae.
Pedigerous segments 3—5 have dorso-lateral carinae, segments 4 and 5
having also each a median carina. Pleon with median and lateral ridges,
the median faint on segments 2—4, the lateral peculiarly arranged on segment
5; a transverse crescentic groove on the telsonic segment. Eyelobe having
at the apex very small corneal lenses, the number undetermined. Peraeopod
1 long, slender, 2" joint about two-thirds as long as rest of limb, 7%
joint subequal to 5%, about two-thirds as long as 6%. Other limbs about
as in C, australis (nr. 20). Peduncle of uropods little longer than the
equal rami, which end acutely without terminal spines, endopod having
more than the proximal half of its medial margin serrate, this part ending
in a spine, the rest bare. L. © with developing marsupium 11°25 mm, 2
unknown.
Lat. 19 425 S., long. 130° 475 E., W. of New Guinea; depth 32 m.
Cumacea: 3. Bodotriidae, 4. Cyclaspis 35
13. C. exsculpta O. Sars 1887 C.e., G.O.Sars in: Rep. Voy. Challenger, v.19
pars 55 p.20 t.1 f.24—26 | 1905 C.e., Calman in: Siboga-Exp., pars 36 p. 3, 4 | 1907
C. e., Calman in: Tr. zool. Soc. London, v. 181 p. 6.
Integument highly indurated. Pseudorostrum and eyelobe as in C,
persculpta. Carapace strongly sculptured; a carina crossing it immediately
behind the frontal lobe and a second at about one-third of its length from
the end, the latter notched in the middle, both connected by a median longi-
tudinal carina and 2 lateral ridges, dividing middle part of carapace into
4 depressed areas, 2 dorsal, 2 lateral of irregular shape. From anterior
transverse carina 2 slight ridges run forward on each side, one ending in
front of the antero-lateral tooth, the other behind it at some distance. The
median carina behind the second transverse one is flanked by a sub-
parallel pair. Pedigerous segments 2—4 with 3 longitudinal crests. Pleon
missing. Eyelobe having at the apex a few small corneal lenses. Colour
uniformly whitish. L. (carapace and pedigerous segments) 2 mm, estimate
for total length under 5 mm.
Flinders Passage, between Australia and New Guinea, depth 13 m.
14. C. elegans Calm. 1907 C. e., Calman in: Tr.zool. Soe. London, v.181 p.9 t.2.
Pseudorostral lobes produced a little in front of the eyelobe, squarely
truncate in Gd, a little obliquely in Q; antero-lateral corners nearly on a
level with the apices. Carapace strongly sculptured, but without transverse
carinae; a median dorsal carina ends in a bluntly raised tooth; on each
side a quadrilateral depressed area is bounded by ridges with a tubercle at
each corner, the front margin sinuous, the hinder subparallel to that of the
carapace, the lower horizontal ridge produced in front and behind to the
edge of the carapace, the upper continued forward to the side of the pseudo-
rostrum. Pedigerous segment 1 in © showing a little laterally, 2™¢ large
with carina ending in a sharp tooth, and side-plates expanded; tooth wanting
and plates not expanded in G, median carina continued to end of pleon
segment 5, accompanied by lateral carinae on pedigerous segments 4, 5, and
first 2 pleon segments, these carinae stronger in CO than in ©. Eyelobe
constricted at base, little longer than broad, with about 9 corneal lenses
but no pigment. Antenna 1 without trace of accessory flagellum, 3¢ joint
of peduncle shorter than 2"¢. Antenna 2 of © 1-jointed, with 3 plumose setae.
About 18 spines in spine-row of mandible. Lower lip with apices very
narrow, incurved, carrying spatulate spines. Maxilla 1 with 2 apical filaments
on palp. Maxilliped with about 13 branchial leaves in O, 17 in G. Manxilli-
ped 3, 2™¢ joint sharply angled, produced to middle of 4 joint which is
also broadly produced. Peraeopod 1, 2™¢ joint longer than rest of limb,
6 joint not long but longer than the 5 or 7%. Pleopods of d differing
from those of C. longicaudata (nr. 3) by absence of process on lateral margin
of endopod. Peduncle of uropods scarcely longer than the rami which are
subequal in ©, endopod serrate on distal half of medial margin and armed
with a small pectinate spine, the apex slightly curved upwards and outwards;
endopod in GC the shorter, strongly fringed like the peduncle with plumose
setae. L. 0 63 mm, CO 6:2 mm.
New Zealand, Lyttleton Harbour: depth 2—9 m.
15. C. similis Calm.. 1907 C. s., Calman in: Tr. zool. Soc. London, v. 181
p.12 t.3 £. 1—8.
Similar to C. elegans (nr. 14) in shape and proportions, but lateral
depressed area on carapace less distinctly excavated, with one tubercle instead
2
3*
36 Cumacea: 3. Bodotriidae, 4. Cyclaspis
of four, the lower horizontal ridge continued behind with even curve into
the hinder vertical ridge, not extending behind to the edge of the carapace;
the hind vertical ridge forks into two obtusely diverging branches; the
front one does not reach the ridge running backwards from the anterior
upper tubercle. Dorsal crest of 24 pedigerous segment. produced backwards
into a large rounded lobe. Dorso-lateral crests of following segments stronger
than in C. elegans. Peduncle of uropods more decidedly longer than the
rami, endopod not curved at the tip. L. 9 5°75 mm, GC (immature, with no
cristiform lobe on 24 pedigerous segment) 5°25 mm.
New Zealand, Lyttleton Harbour; depth 2—9 m.
16. C. uniplicata Calm. 1904 C. sp., Calman in: Herdman, Rep. Ceylon
Pearl Fish., 7.2 p.160 | 1907 C. uniplicata, Calman in: Tr. zool. Soc. London, v. 181
p.7, 13 t.4 f.11—20.
Pseudorostral lobes short, truncate, not reaching beyond the eyelobe;
antero-lateral corner showing 2 teeth, the lower the sharper and more produced.
Carapace sharply carinate, with tooth overhanging base of eyelobe, and sharp
ridge from near the middle curving downwards and forwards on each side
towards but not reaching the antero-lateral corner. First pedigerous segment
narrowly exposed dorsally, 2"¢ feebly carinate. Pleon long, rather slender.
Eyelobe not longer than broad, occupied by about 9 corneal lenses, with
pigment. Antenna 1, 3% joint longer than 2™4, accessory flagellum minute
but distinct. Maxilliped 3, 2™4 joint angled, longer than rest of limb, distally
produced nearly to end of the much produced 4. Peraeopod 1 very long,
ynd joint apart from distal tooth of medial margin little longer than 6™ and
7*h combined, 78 longer than 5” but considerably shorter than 6%. Peraeopod
2 stout, 2™4 joint longer than rest of limb, in which the 7 joint is the longest.
Peduncle of uropods slightly shorter than the exopod, as long as the endopod,
which has about 9 spines on the medial margin, and is distally bare, api-
cally acute. L. © (immature) 4°99 mm, O unknown.
Gulf of Siam, Koh Kam; depth 6—18 m.
17. C. unicornis Calm. 1907 C. u., Calman in: Tr. zool. Soe. London, v. 181
p.7, 14 t.5 f. 9—11.
Pseudorostral lobes slightly prominent, eyelobe reaching quite to the tip;
antero-lateral corner acutely produced, not quite level with pseudorostral apices.
Carapace granular, with median carina produced into a forward pointing
tooth behind the eyelobe, without transverse ridges. First pedigerous segment
not exposed, 2"4 feebly carinate. Pleon rather long. Peraeopod 1, 24 joint about
three-fourths as long as rest of limb, 32 and 4» joints stout, the slender 5%
and 6t equal, 7 much shorter than either. Peduncle of uropods considerably
longer than the rami, exopod slightly longer than endopod, which has 4 small
spines on the serrate medial margin and a long one on the subobtuse apex.
L. O (immature) 3°2 mm, CG unknown.
Danish West Indies, Cruz Bay, St. Jan.
18. C. cingulata Calm. 1907 C.c., Calman in: Tr. zool. Soc. London, v.181
poi t40e 12-10:
©. Pseudorostral lobes short, horizontal, with the long narrow eyelobe
projecting in front; antero-lateral tooth not level with the apices. Carapace
strongly arched, a little in front of middle encircled by a very prominent
collar, notched at the centre, its lateral apices turned a little forwards and
Cumacea: 3. Bodotriidae, 4. Cyclaspis 37
marking the greatest breadth of the carapace; on the last third of the length
a medio-dorsal pair of carinae extend to the hind margin. Pedigerous segment
1 is dorsally exposed, though overhung by the carapace, the 4% and 5%
have well marked dorso-lateral carinae, which are also found on the first 5
Fig. 15.
C. cingulata (2) Calman.
pleon segments; a median keel extends from the 5 pedigerous to the! 5th
pleon segment, but is feeble at beginning and end of the series. Hyelobe
with 11 corneal lenses on the distal end. Antenna 1
with 34 joint longer than 2™¢, accessory flagellum
minute but distinct. Maxilliped 3, 2"¢ joint with
distal process reaching nearly as far as that of the
4th joint, its length without this long process little
more than the rest of limb. Peraeopod 1, 2"4 joint
feebly produced at apex, length less than rest of limb,
5th and 7* joints equal, each a little shorter than 6%.
Pedunele of uropods longer than the exopod but slightly
shorter than the endopod, which has 3 small spines
about the middle of the medial margin, the apex bare,
acute. L. 4:2 mm.
GS. Carapace with only a faint trace of the
encircling collar, and without the hinder pair of dorsal
carinae; infero-laterally a faint horizontal ridge meets
the collar anteriorly. Dorso-lateral keels on last 2
pedigerous segments, but the stout pleon segments
have no distinct keels. Terminal joints of peraeopod 1 Fig. 16.
longer and more slender than in ©. C. cingulata Calman,
maxilliped 3.
Gulf of Siam; depth 9—18 mm.
19. C.thomsoni Calm. 1907 C.¢., Calman in: Tr. zool. Soe. London, v. 181
D714 25 £1216,
Near to C. levis (nr. 8). Pseudorostral lobes short, with eyelobe
reaching to or beyond their apices; antero-lateral tooth minute. Carapace
pitted, rugose, with dorsal carina flattened or faintly doubled to the rear;
on either side behind the middle a short groove is limited by a faint ridge
running obliquely downwards; a slight short ridge runs backward from the
antero-lateral corners. Pedigerous segment 1 exposed at the sides in ©,
not in dt, 2"¢ with dorsal carina, 34 to 5 with lateral carinae. Pleon
segments with dorsal and lateral carinae and articular processes in Q, the
lateral carinae wanting in G. Eyelobe apparently short, occupied by several
corneal lenses. Peraeopod 1, 2"4 joint not distally produced but carrying 2
38 Cumacea: 3. Bodotriidae, 4. Cyclaspis
plumose setae at the apex, length a little less than rest of limb, 7 joint
about two-thirds of 5, three-fourths of 6 in length. Peduncle of uropods
very little longer than exopod, which is a little longer than the endopod;
the latter having 4 spines on its medial margin in Q, plumose setae and
9 spines in G, the apex bare, acute. L. Q 6mm, CO 675 mm.
New Zealand, Bay of Islands; depth 15 m.
20. C. australis O. Sars 1887 C.a., G. O. Sars in: Rep. Voy. Challenger, v.19
pars 55 p.12 t.1 f.1—20 | 1907 C.a., Calman in: Tr. zool. Soe. London, v.181 p.7.
Integument highly indurated, granular or squamose or reticulate. Pseudo-
rostral lobes short, obtusely truncate, not produced beyond the long narrow
eyelobe; antero-lateral corners reaching level of apices, a short carina running
back from the short tooth. Carapace with dorsal cara down the middle,
an obliquely longitudinal ridge on either side limiting the gastric region
and connected with this a slight curved prominence passing down the sides
to the rear. Pedigerous segment 2 firmly connected with the carapace,
following segments abruptly lower, 4" and 5 with 3 carinae. Pleon with
median carina on all segments, 3 carinae on the first two, articular processes
as usual. Eyelobe with 7 or more corneal lenses on the slightly tumid apex,
pigment dark. Antenna 1, 2™4 and 32 joints subequal, flagellum 2-jointed,
accessory minute, l-jointed. Antenna 2 © 2-jointed (text, undivided in fig.)
Upper lip slightly emarginate. Lower lip with apices slightly incurved, armed
with spatulate spines. Maxilla 1, palp ending in 2 long but unequal filaments.
Maxilliped 1, epipod with 14 branchial leaves, exopod ending in an indurated
lamella armed with six curved setae and encircled by a very thin and
pellucid border. Maxilliped 3, 2™¢ and 4 joints apically much produced.
Peraeopod 1, 2™4 joint longer than rest of limb, its distal half abruptly
narrowed, not or scarcely produced at apex, 6‘ joint equal to 5%, longer
than 7th, Peraeopod 2 short, with large 1s joint. Peduncle of uropods
shorter than the exopod, equal to the endopod, which has the medial margin
finely serrulate, each branch with apex acute, bare. Colour (as preserved)
light brown, with darker shadings, especially in front. L. © (not fully adult)
8mm, C (not fully adult) 10 mm.
Australia, Port Philip; depth 69 m.
21. C.quadrituberculata Zimmer 1907 C. q., Zimmer in: Zool. Anz., v.31 p.367.
Integument with larger granules in reticulate arrangement among
smaller ones. Pseudorostral lobes not reaching in advance of the eyelobe;
antero-lateral corners acute. Carapace somewhat compressed, with median
rounded keel, flanked by tubercles, one rather elongate pair on the pseudo-
rostral lobes outside the frontal lobe, the other behind the first about two-
thirds of the way down the length of the carapace. Pedigerous segment 1
completely fused with carapace. A median carina runs from 5‘ pedigerous
segment to end of pleon. Peduncle of uropods scarcely longer than exopod;
exopod longer than endopod, which has spines on the medial margin. Colour
white. L. d (not adult) 12 mm. © unknown.
South Georgia; depth 75 m.
22. C. gigas Zimmer 1907 C.g., Zimmer in: Zool. Anz., v.31 p. 367.
Integument finely reticulate. Pseudorostral lobes meeting in a point
in front of the long eyelobe; antero-lateral corners acute. Carapace very
tumid, in front half medio-dorsally elevated between 2 deep depressions ;
Cumacea: 3. Bodotriidae, 4. Cyclaspis 39
frontal lobe flanked by a pair of blunt tubercles; behind it sublaterally stands
a longer pair, sloping down as it were from the raised hinder division
of the carapace; below these tubercles the pseudorostral lobes have a
longitudinal rounded ridge. The hind margin of the carapace is obtusely
excavate, the fusion with 1st pedigerous segment dorsally well marked.
A median carina extends from the 4 pedigerous segment to end of pleon.
The eyelobe shows two corneal lenses in front. Antenna 1 extremely small.
Peduncle of uropods slightly longer than the endopod, which is rather longer
than the exopod and has on the medial margin a close-set series of spines
encreasing in size towards the apex. Colour white. L. © (with marsupium)
about 15 mm.
German South Pole Exp., Winter Station (lat. 66° 2’ S., long. 89° 38’ E.).
23. C. costata Calm. 1904 C.c., Calman in: Herdman, Rep. Ceylon Pearl Fish.,
v.2 p. 168 t. 3 f. 39—53 | 1907 C. c., Calman in: Tr. zool. Soc. London, v. 181 p. 7.
Pseudorostral lobes meeting well in front of the large subcircular
eyelobe; antero-lateral corners obtuse, especially in the G. Carapace
compressed, anterior half with carina flanked by a depression on either side;
behind and below these the carapace in © is beset with longitudinal
ridges, 12 or more in number, formed by rows of minute granules, in the ¢
only hinder dorsal third of carapace occupied by the ridges, a vertical ridge
separating them from the irregularly granulated anterior portion. Pedigerous
segment 1 exposed, less in GS than in Q, segment 2 with median crest in Q,
feeble in G, segments 4 and 5 each with submedian pair of tubercles in
©, these segments in G depressed, with prominent dorso-lateral corners.
Pleon rather stout, with feeble carina in Q. Eye pigmented, with corneal
lenses, which at least in GO are large and distinct. Antenna 1 very short,
32 segment rather longer than 2"4, flagellum 2-jointed, accessory minute.
Antenna 2 © unjointed, with 2 plumose setae. Maxilliped 3, 27¢ joint
much bent, without the narrow distal process rather longer than rest of limb,
process of 4*® joint large reaching much beyond that of the 24 joint, 5% and
6th joints broad, 7** almost rudimentary. Peraeopod 1 very short, 2"¢ joint
not apically produced, about as long as rest of limb. Peraeopod 2 in G
having front margin of 2™4 joint serrate with recurved teeth, apparently
wanting in Q. Peduncle of uropods a little longer than the subequal
rami in ©, but equal to those of the CG; all the rami have an apical
spine, the endopod of © has also a subapical spine; the much broader
endopod of 3 is fringed on the medial margin with serrate spines, with a
short interval between the series and three on the oblique apex. L. ©
adult 3°75 mm, CG adult 3°9 mm.
Gulf of Manaar, depth 15—16 m.
24. C. bistriata Zimmer 1902 C. b., Zimmer in: Zool. Jahrb., Syst. v.17 p. 447
tf. D—F, | 1907 C. biplicata, Calman in: Tr.zool.Soc. London, v.181 p.7,17 t.3 f.4—15.
Pseudorostral lobes truncate, meeting just in front of eyelobe, antero-
lateral tooth acute, reaching level of the apices. Carapace compressed,
more or less arched in ©, with median carina, rising especially in © in-
to a blunt tooth at hind margin (Calman), flanked by 2 pairs of parallel
striae or very low keels in obliquely transverse curves, the anterior close to
the frontal lobe, the second about half-way down the carapace (Zimmer), or
both further back and converging behind middle of carapace (Calman), feeble
and not meeting in GC. Pedigerous segment 1 only laterally exposed and
40 Cumacea: 3. Bodotriidae, 4. Cyclaspis, 5. Stephanomma
only that in Q, 24 with carina forming a more or less slender tooth, curving
a little forward (Calman), the other segments with a carina, which is continued
along front part of the pleon, which has the usual articulations. Eyelobe
a little longer than broad, with about 9 corneal lenses (Calman), 2 rather
long but small and 2 large but indistinct (Zimmer). Antenna 1, 34
joint longer than 2", Antenna 2 Q 2-jointed. Peraeopod 1, 24 joint
longer than rest of limb. Peduncle of uropods shorter than endopod which is
shorter than exopod (Zimmer), subequal to the subequal rami (Calman in
text, but endopod shorter than exopod in fig.) L. Q almost 5 mm, (Zimmer),
Qo 42 mm, d 4:16 mm. (Calman).
New Zealand, Plenty Bay, Lyttleton Harbour, depth 2—9 m.
25. C. triplicata Calm. 1907 C.t., Calman in: Tr. zool. Soe. London, v. 181
p.7, 19 t.8 £.16—24.
Very like C. bistriata but with the 2 pairs of oblique ridges on the
carapace here much more prominent, and a third pair starting from a pair
of prominent tubercles outside the frontal lobe and running nearly vertically
downwards to join a prolongation of the anterior oblique ridge which runs
horizontally forwards to the anterior lower edge of the carapace; tooth at
end of median carina serrated behind, carina of 224 pedigerous segment
sinuous, with front angle blunt and hinder one overhanging the third segment;
continuance of median carina here faintly marked, but 4" and 5‘ pedigerous
segments and first three of pleon laterally carinate. Peraeopod 1 with 2™¢
joint longer in relation to rest of limb, uropods a little stouter. L. @ 4 mm,
SG 4 mm.
New Zealand, Lyttleton Harbour, depth 2—9 m.
5. Gen. Stephanomma 0. Sars
1871 Stephanomma (Sp. un.: S. goésii), G. O. Sars in: Ofy. Ak. Forh., v.28 nr. 6
p. 808 | 1873 S., G. O. Sars in: Svenska Ak. Handl., n. ser. v.11 nr.5 p.16 | 1907 S.,
Calman in: Tr. zool. Soc. London, v.181 p. 14. .
No pseudorostral lobes meeting in front of the eye, or separated
from the carapace even by a suture, antero-lateral tooth subacute, well advanced.
Carapace not inflated. Five pedigerous segments distinct. Eye composed
of numerous lenses. Maxilliped 3 with 2™4 joint unusually short, 3¢ longer
than broad, presence of exopod in female doubtful. Uropods with one-
jointed endopod.
1 species.
1. S. goésii O. Sars 1871 S.g., G. O. Sars in: Ofv. Ak. Forh., v.28 nr. 6
p.808 | 1873 S. g., G.O. Sars in: Svenska Ak. Handl., n.ser. v.11 nr.5 p.16 t.4
f. 17—23.
Integument much indurated, finely facetted, pitted and reticulate. Cara-
pace with short rounded keels behind each antero-lateral tooth, a central carina
from end to end, raised behind the eye and there presenting a forward-
pointing tooth, this flanked by a pair of depressions with a pair of tubercular ele-
vations. Pedigerous segment | short strap-shaped, 224 with outstanding side-plates
extended forwards over hind margin of the carapace, 3¢ and 4‘ with similar
side-plates extending backwards, 2"¢ to 5 with median carina, the 5 and the
first 5 segments of the pleon having also on each side 3 strongly marked
earinae. The telsonic segment is produced triangularly between the bases of
Cumacea: 3. Bodotriidae, 5. Stephanomma, 6. Zygosiphon 41
the peduneles of the uropods. Eye situated on the broad blunt-ended equi-
valent of an eyelobe, and composed of 10 corneal lenses encircling a central
one. Below this region a semicircular
orifice leading to the branchial cham-
bers can be closed by the triangular
exopods of maxilliped 1. Antenna |
small, the three joints of the pe-
dunecle subequal in length, flagellum
very short, accessory undeveloped.
Maxilliped 3, 2™¢ joint scarcely
longer than the 3¢ and 4 com-
bined, not apically produced, 4%
joint broadly produced to a pointed
apex. Peraeopod 1 slender, attached
unusually far down on the sides
under the carapace. Peraeopod 2
little more than half as long as
the 1st. Peduncle of uropods quite
smooth, considerably longer than
the subequal rami; the lanceolate ane
endopod with medial margin finely S. pout o) Gare.
serrate and carrying a row of 5 short
spies along the middle. L. © adult about 9°5 mm, or including uropods
about 11 mm, @ unknown.
West Indies, St. Martin.
6. Gen. Zygosiphon Calman
1907 Zygosiphon, Calman in: Tr. zool. Soc. London, v. 181 p. 25.
Pseudorostrum with frontal projections wide apart. Branchial siphons
very long and widely separated. All five pedigerous segments free. Uropods
with endopod of two joints, the second joint much the shorter.
1 species.
1. Z. mortenseni Calm. 1907 Z. m., Calman in: Tr. zool. Soe. London,
v.181 p.25 t.7 f. 1—19.
Fig. 18.
Z. mortenseni (2) Calman.
_ Pseudorostrum with plates meeting in front of eyelobe, then broadly
divergent with antero-lateral tooth on each side in advance of their short
42 Cumacea: 3. Bodotriidae, 6. Zygosiphon, 7. Iphinoé
prominences. Carapace with sides more or less straight, which bulge to the
rear in ovigerous specimens; on each side a lozenge-shaped depression bounded
by ridges faint in © and still more so in dG; from the front corner of the
depression a ridge runs forward on to the side of the pseudorostrum; the
hind margin is obtusely raised. The last 3 pedigerous segments are much
lower than the 18¢and 224, and diminish much in width successively. The
Fig. 19.
Z. mortenseni (2) Calman, front.
telsonic segment is well produced between the uropods and is not shorter
than the segment next preceding it. The eyelobe is broad, notched or emar-
ginate in front, with 2 separate masses of pigment but no distinct lenses
in ©. but deeply pigmented and with 9 corneal lenses in G. Antenna 1,
1st joint as long as 2"4 and 34 combined, 34 twice
as long as 224, Mandible with 12 spines in spine-
row. Maxilla 1, palp with 2 apical filaments.
Maxilliped 1, epipod with very few branchial
leaflets in O, about 9 inG’, exopod ending in
a cone of spirally coiled membrane capable of
being extended to a great length forward and out-
ward. Maxilliped 3, 2™4 joint bent, externally
Fig. 20. produced into along pointed lobe, 4% joint also
Z. mortenseni (3) Calman, front. expanded externally into a very large curved
lobe. Peraeopod 1, 2"4 joint in dD about as long
as the rest combined, in © about two-thirds of their length. Peraeopod 2, ou
joint a little less than the rest combined, 5 and 6 joints subequal, very short.
Uropods slender, length of exopod in Q nearly, in Go a little over, three-fifths
that of peduncle, endopod three-fourths as long as exopod, 2"¢ joint rather
over a third as long as 18‘, medial margin with 3 spines in Q, 6 in O.
Li. 0.267, CO 2:7 am.
Gulf of Siam, depth 9—19 m; Gulf of Manaar.
7. Gen. Iphinoé Bate
1856 Halia (Sp. un.: H. trispinosa) (non Risso 1826, Mollusca), Bate in: Ann.
nat. Hist., ser.2 v.17 p.458 | 1856 Venilia (Sp. un.: V. gracilis) (non Duponchel 1829,
Lepidoptera), Bate in: Ann. nat. Hist., ser.2 v.17 p.460 | 1856 Iphinoé (Sp. un.: 72
trispinosa), Bate in: Ann. nat. Hist., ser. 2 v.18 p. 187 | 1856 Cyrianassa (Sp. un.: Gs
:
Cumacea: 3. Bodotriidae, 7. Iphinoé 43
gracilis), Bate in: Ann. nat. Hist., ser. 2 v.18 p. 187 | 1867 Iphithoé (err. pro: Iphinoé),
A. M. Norman in: Rep. Brit. Nee v.36 p.201 | 1869 Iphinoé, A. M. Norman in: Rep.
Brit. Ass., v.88 p.¥72 | 1878 I., G. O. Sars in: Arch. Naturv. Kristian., v.3 p.495 |
1886 Sphinoe (err. pro: Iphinoé), R. Koehler in: Bibl. hautes Etudes, v.32 nr. 4
p.23 | 1893 Iphinoe, T. Stebbing in: Int. sci. Ser., v.74 p.302 | 1895 Iphinoé, H.
J. Hansen in: Ergeb. Plankton-Exp., v.2 G.c. p. 52 | 1899 I, G. O. Sars, Crust. Norway,
v.38 p.12 | 1907 7, Calman in: Bull. Mus. Paris, p.118 | 1908 Iphinoe, Zimmer in:
Ergeb. Tigter divers v.8 p. 162 | 1910 Iphinoé, T. Stebbing in: Ann. S. Afr. Mus., v. 61v
p. 410.
Pseudorostral lobes meeting in front of the eyelobe. Carapace narrow,
without lateral cornua. First pedigerous segment short but distinct. Only
the first pair of peraeopods carrying exopods in either sex; second peraeopods
with the third joint obsolete. Endopod of uropods two-jointed, the second
joint usually the longer.
8 species.
Synopsis of species:
{ Uropod with 1st joint of endopod longer than 2nd. , , . 1. I. crassipes
) Uropod with 1st joint of endopod shorter than 2nd — 2,
{ Maxilliped 3 with 5th joint distally widened . “
) Maxilliped 3 with 5th joint not distally widened — 3.
Carapace slender — 4.
Carapace comparatively short and stout — 7.
Maxilliped 3 with 2ndjoint much more than twice as we
| as the rest of the appendage ... . ise 3, I. brevipes
- | Maxilliped 3 with 2ndjoint not much more Chem twice as
J
\
bo
e
. zimmeri
long as the rest of the appendage — 5.
Carina of carapace not serrate in a8 with not more than
Grtecthe ini Oso. eco le trEispinoss:
Carina of carapace sachs in pot sexes, mich more than
6 teeth in 9 — 6,
Serration of carina not limited to front half in oe limited
6 to hinder half in 3?) : 4. I. serrata
Serration of carina limited to front tale in Both sexes . 6. I. tenella
_ { Peraeopod 2 uncommonly long, rather longer than the 34. 7. I. robusta
: Peraeopod 2 uncommonly short, shorter than the 34 8. I. inermis
1. I. ecrassipes H.J. Hansen 1895 J.c., H.J. Hansen in: Ergeb. Plankton-
Exp., v2 G.c. p.53 t.6 f.4,4a—f | 1904 I. macrobrachium, Calman in: Herdman,
Rep. Ceylon Pearl Fish., v.2 p.173 t.4 f.72—75 | 1910 I. crassipes, T. Stebbing in:
Ann. 8S. Afr. Mus., v. 61v: p. 412 t. 19.
Pseudorostral lobes upturned, distally forming a semicircular cavity,
the produced lateral corners of which carry convergent tufts of setae; antero-
lateral corners acute in Q, not produced in G. Carapace with median keel
-and 2 weak pairs of lateral keels anteriorly. Pedigerous segments 3 and 4
of Gas in J. zimmert (nr. 2) and J. brevipes (nr. 3). Telsonic segment roundly
produced between the uropods at their base and showing a dorsal depression.
Eyelobe rounded, with 10 corneal lenses, 2 of them central, closely united.
Antenna 1, 3¢ joint longer than 2™4, as long as 1S‘, flagellum with 8 filaments
fringing 1S* joint, 274 tipped with long curved spine and 2 filaments. Maxilla 1
with 2 filaments at end of palp, and a distally plumose seta on lateral margin
of lateral lobe. Maxilliped 3, 2™4 joint produced to middle of the broad
1) Or sometimes wanting.
44 Cumacea: 3. Bodotriidae, 7. Iphinoé
prolongation of the 4** joint, this process itself reaching much beyond the
5th joint, its lateral margin fringed with 12 plumose setae, the apical 2 much
the longest. Peraeopod 1, 2™4 joint much shorter than rest of limb, 6 shorter
than 5th, little longer than 7, Peraeopod 2, terminal joint nearly as long
as 3 preceding joints combined. Peduncle of uropods longer than the subequal
rami, its medial margin in adult G closely fringed with spines, as is also
the 18t joint of the endopod, which is more than twice as long as the narrow
Fig. 21. Fig. 22.
I. crassipes Hansen, maxilliped 3. I, crassipes Hansen, uropod and telsonic
(after Stebbing). segment (after Stebbing).
ond joint, that carrying distally 4 spines and 2 long setae; in younger speci-
mens the relations are different, the 18* joint not nearly twice as long as
the 2™4, and in J. macrobrachium the exopod much shorter than the endopod.
L. O adult 8mm, GC juv. 3:2 mm, 9 juv. 1 mm.
Gulf of Guinea, Anecho (Togo); Gulf of Manaar, depth 8—13 m (I. macrobrachium);
South Africa, depth 75 m.
2. I, zimmeri Stebb. 1910 J. z., T. Stebbing in: Ann. S. Afr. Mus., v. 61Vv
p.411 t. 18.
SG. Pseudorostral lobes obliquely subtruncate, meeting a little in front
of the semicircular eyelobe. Carapace smooth; 3¢ and 4% pedigerous seg-
ments with articulating arrangement strongly developed; 5% segment of pleon
distally narrowed, telsonic segment with dorsal depression. Eye obscure.
Antenna 1, 34 joint a little longer than 2™4, Maxilliped 1 with the branchial
leaflets numerous, closely contiguous. Maxilliped 2, 2"¢ joint gradually widening
and then rather abruptly narrowing to the apex. Maxilliped 3, distal process
of 24 joint not nearly reaching apex of distal process of 4% joint, 5" joint
unusually widened distally on the medial side, as in Heterocuma sarsi (p. 48).
Peraeopod 1, 2"4 joint narrowed distally, nearly as long as rest of limb and
subequal to peraeopod 2, in which the 2™¢ joint is distally widened with
Cumacea: 3. Bodotriidae, 7. Iphinoé 45
rounded distal process simulating and perhaps representing a short 34 joint.
Peduncle of uropods subequal to the rami; endopod with 1S joint much
stouter than 2"4, about half as long,
fringed on medial margin with 10
pectinate spines, the apical much the
longest, 2"4 joint fringed with a dozen
spines, the lowest the largest, follow-
ed by an oblique series of 10 spinules
extending to the acute apex which
carries an inward pointing spine. L.
9mm. A specimen without pleopods,
L. 5 mm, agrees as to the uropods
with J. robusta, H. J. Hansen (ny. 7).
Fig. 23. Fig. 24.
I. zimmeri Stebbing, South Africa, lat. 32° 53’ 30” S., I. zimmeri Stebbing,
maxilliped 3. long. 289 11’ 00’ K.; depth 75 m. uropod.
3. I. brevipes H.J. Hansen 1895 J. b., H. J. Hansen in: Ergeb. Plankton-
Exp., «2 G.c. p.54 t.6 £5, 5a—h | 1908 I. africana, Zimmer in: Ergeb. Tiefsee-Exp.,
v.8 p.163 t. 37 £.17—833 | 1910 TF. brevipes, T. Stebbing in: Ann.S. Afr. Mus., v.61v p. 410.
Pseudorostral lobes rather short, horizontal, acute at apex, then truncate,
then emarginate; antero-lateral corners acutely produced in Q. Carapace
very slender, nearly three times as long as high, lower margin for a short
space anteriorly serrate, otherwise quite smooth, with no distinct median
carina, which is confined to the pleon segments, but sometimes having a
longitudinal medio-dorsal row of denticles beginning behind the eyelobe and
extending half the length of the carapace (Zimmer); the pedigerous segments
quite smooth. Eyelobe rounded, eye darkly pigmented, corneal lenses indistinct.
Antenna 1, 34 joint longer than 2"¢, Antenna 2 in © 1-jointed, with 3 or 4
plumose setae, in G reaching to end of pleon. Maxilliped 3, 2"¢ joint unusually
long, but distal process moderately produced, rest of limb little more than
one-third as long as 2™4 joint, 4% joint moderately produced, 5‘ not specially
widened. Peraeopod 1, 2"4 joint considerably longer than rest of limb, 7 joint
nearly equal to 6, Peraeopod 2 very short, shorter than peraeopod 3,
gnd joint abnormal, little longer than broad, the next joint with strong spine
on produced truncate apex, 3% joint wanting. Pedunele of uropods in imma-
ture © equal to, in O adult longer than the equal rami, exopod robust, with
numerous setae on margins and blunt apex, endopod in G with 1* joint little
longer than broad, about half as long as the narrow second joint, carrying
short spines on medial margin and a long apical spine, the 2"¢ joint fringed
with spines successively larger to the long terminal spine. Colour a clear,
almost pellucid, grey. L. d adult 86 mm, 9 immature 6°6 mm, Q ovigerous
about 12—18 mm, ¢ adult 15 mm.
Gulf of Guinea, Cotonu; Great Fish Bay; South Africa, depth 44—62 m.
4. I, serrata (Norm.) 1867 Iphithoé s., A. M. Norman in: Rep. Brit. Ass.,
v.36 p.201 | 1869 Iphinoés., A.M. Norman in: Rep. Brit. Ass., v.38 p.272 | 1878
?T. (misprint Ihpinoé) gracilis (part.) var. s., G.O.Sars in: Arch. Naturv. Kristian.,
v.3 p.496 t.10—14 | 1886 Sphinoes., R. Koehler in: Bibl. hautes Etudes, v.32 nr. 4
p.23 | 1899 ?Iphinoé s., G. O. Sars, Crust. Norway, v.3 p.14 | 1905 I. s., Calman in:
Sci. Invest. Fish. Ireland, 1904 app.1 p.16 | 1907 J. s., Calman in: Bull. Mus. Paris,
nr. 2 p. 118.
Animal very long and slender. Pseudorostral lobes well produced,
slightly bent upwards, obliquely truncate, crenate, ciliated; antero-lateral corners
a
46 Cumacea: 3. Bodotriidae, 7. Iphinoé
acute. Carapace twice and a half (Norman), about twice (Sars), as long as
high, median carina in © with 13 to 20 forward pointing teeth, successively
larger to the front, where 2 or 3 are sometimes separated from the main series,
which may or may not reach the hind margin; in G: (Sars) pseudorostral
lobes shorter, more level, antero-lateral corners obtuse, carapace more com-
pressed, sometimes entirely without serrate carina, sometimes with one but its
serrations not reaching the frontal lobe. First pedigerous segment chiefly
visible in dorsal view, 24 very large, all five and the pleon segments with
median carina. Eyelobe of Q ending in 2 little points, eye red-brown, with
9 corneal lenses in G, those in © smaller and less distinct. Antenna 1,
3° joint longer than 18*, which is longer than 2"¢, flagellum 1-jointed, with
minute 2-jointed accessory. Antenna 2 of © 2-jointed with 4 setae on the
large 18* joint. Lower lip, with apices not rounded, armed with 5 peculiar
spines. Maxilliped 1, branchial leaflets numerous. Maxilliped 3, 224 joint
with distal process reaching middle of 4, and fringed on medial margin with
long plumose setae, the 4% fringed in like manner on lateral margin of its
prolongation. Peraeopod 1 slender, 224 joint shorter than the rest of limb,
having both margins denticulated (A. M. Norman), denticulation not shown
on lateral margin nor mentioned in text (G. O. Sars), 5% and 7 subequal
and about one-third shorter than 6% (A. M. Norman), 7‘ figured about one-
third shorter than the subequal 5*® and 6 (G. O. Sars). Peraeopod 2 very
short, with a joint missing, the 2 joints succeeding the 2™¢ having each a
long apical spine, terminal joint not longer than 2 preceding joints combined.
Peduncle of uropods longer than the slightly unequal rami, its medial mar-
gin in © armed with 12—14 long spines, in the G with many more in
2 or 3 rows; exopod broad with many plumose setae on medial margin of
2nd joint and round the apex, the slightly shorter endopod with 1° joint
about twice as broad but little over half as long as 2"4, carrying 3 or 4
small and a long apical spine, 2"¢ joint fringed in Q with 13, in CO with
16 spines, encreasing in size to the apex. Colour pale straw yellow, flecked
with brown. L. Q 12mm, dO 105 mm.
Shetland Isles, depth 55—147 m (A. M. Norman); Mediterranean, Bay of Naples ete.
(G. O. Sars); W. of Ireland, depth 31—837 m (Calman).
5. I. trispinosa (Goodsir) 1843 Cuma t., Goodsir in: Edinb. new phil. J.,
v.34 p.126 t.3 f.1—7 | 1853 C. ¢., T. Bell, Brit. stalk-eyed Crust., p. 329 tf. | 1856 C. ¢.,
W. Thompson, Nat. Hist. Ireland, v. 4 p. 392 | 1856 Halia t. + Venilia gracilis, Bate in:
Ann. nat. Hist., ser.2 v.17 p.458 t.14 f.5; t.15 f.5; p.460 t. 15 f.7 (4) | 1856 [phinoé
t. + Cyrianassa g., Bate in: Ann. nat. Hist., ser. 2 v.18 p.187 | 1863 Iphithoe t. +
C. g., A.M. Norman in: Tr. Tyneside Club, v. 51v p. 271, 274 | 1869 Iphinoé g. (S
of I. trispinosa), A. M. Norman in: Rep. Brit. Ass., v.38 p. 272 | 1873 Iphithoe
trispinosa, P. Fischer in: Act. Soc. Bordeaux, ser. 3 v.8v p.424 | 1878 Iphinoé gracilis
(part.), G. O. Sars in: Arch. Naturv. Kristian., v3 p.496 | 1886 Sphinoe serrata, R.
Koehler in: Bibl. hautes Etudes, v.32 nr.4 p.23, 60 | 1893 Iphinoé gracilis var. maeotica.
Sowinski in: Mém. Soe. Kiew, v.13 p.357 t.3 f.4 | 1893 J. m., Butschinsky in: Zool.
Anz., v.16 p. 886 (embryology) | 1898 J. gracilis var. m., ‘Sowinski in: Bull. Ae.
St.-Pétersb., ser.5 v.8 p.374 t.3 f.4 | 1899 I. trispinosa, G. O. Sars, Crust. Norway,
v.3 p.14 t.5, 6 | 1901 TF. gracilis, S. Zernow in: Annuaire Mus. St.-Pétersb., v.6 p. 566 |
1905 J. trispinosa, Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p.15 | 1907 J. f.,
Calman in: Bull. Mus. Paris, p.118 | 1909 7. t., A. M. Norman & G.S. Brady in: Nat.
Hist. Tr. Northumb., ser.2 v.3u p.26 | 1912 LF. ¢., Calman in: P.U.S.Mus., v.41 p. 604.
In both sexes very long and slender. Pseudorostral lobes horizontal,
acute; antero-lateral corners triangular in ©, obtuse in G. Carapace of
Cumacea: 3. Bodotriidae, 7. Iphinoé 47
quite smooth above, in © with the hinder half of median carina, rising from
back to front and there followed by a row of 3 or 4 little denticles. Pedi-
gerous segment 1 distinct dorsally, covered below by advanced sides of the
large 224 segment. Pleon longer (G. O. Sars), generally not longer (Calman and
figures by Sars) than the front division of the body. Eye with corneal lenses
prominent and much more distinct in GC than in Q. Maxilliped 3 as in
I. serrata (nr. 5) but perhaps with relatively longer 2¢ joint and less developed
prolongation of the 4%, Peraeopod 1, 24 joint about as long as or longer
than the rest of limb, with both margins showing some denticulation.
Peraeopod 2 much as in J. serrata, with which the uropods also agree.
L. g 10 mm, GC somewhat less.
Norway, very rare; Kattegat; Shetland Isles, depth 55--147 m, and @ in towing-net;
Bay of Biscay; Great Britain and Ireland, shallow water to 26m; Cap Breton, depth
48—65 m.
6. I. tenella O. Sars 1878 J.t., G.O. Sars in: Arch. Naturv. Kristian., v3
p.505 t.15, 16 | 1905 J. t., (with ?Z. trispinosa), Calman in: Sci. Invest. Fish. Ireland,
1904 app.1 p.15 | 1907 J, ¢., Calman in: Bull. Mus. Paris, p. 118.
Body very long and slender. Pseudorostral lobes comparatively short,
horizontal; antero-lateral corners acute in Q, obtuse in GO. Carapace with front
half of carina forming several teeth, successively larger to the front, more
numerous in © than in G. Eye well developed. Antenna 1, 3¢ joint longer
than 2™4, shorter than 15*, both flagella 2-jointed, the accessory minutely.
Maxilliped 3 with process of 4" joint weak and carrying few setae. Peraeopod 1,
the slender 2"¢ joint shorter than rest of limb, 5‘ longer than 6", and 6"
than 7%, Peraeopod 2 with terminal joint considerably longer than 2 preceding
joints combined. Peduncle of uropods considerably longer than the equal
rami, which are similar in armature to those of J. trispinosa, but with more
spines (8 instead of 5) on the first joint of the endopod in the male, the
joint itself much more than half as long as the second. L. © 7 mm,
CS about 8 mm.
Mediterranean, Goletta, Syracuse, Naples, Spezia.
7. I. robusta H. J. Hansen 1895 J. r., H. J. Hansen in: Ergeb. Plankton-Exp.,
v.2 G.c. p.52 t.6 f.3, 3a—d.
Pseudorostral lobes short, with minute apical point, otherwise truncate
with rounded corners; antero-lateral corners not produced. Carapace short
and stout, with feeble medio-dorsal furrow flanked by pair of weak carinae,
which are met at about the middle by a pair of lateral furrows running back
from the front and then curving upwards. Median furrow and carinae con-
tinued on the pedigerous segments and with diminishing vigour on the first
4 pleon segments. Last 3 pedigerous segments with 3 pairs of lateral carinae,
2 above and 1 weaker below. Pleon segments 1—4 have also a feeble
subdorsal pair of carinae; pleon segment 5 has only a single median keel.
Eyelobe broader than long, apically pointed, number of lenses uncertain.
Antenna 1, 34 joint not longer than 24, or than the flagellum, which has
a group of sensory filaments on the 1%¢ joint and 2 long: olfactory setae on
the 24, accessory minute, 1-jointed. Maxilliped 3, 2"¢ joint bluntly produced,
Ath with only 2 plumose setae on its broadly rounded, not elongate process,
5th fusiform. Peraeopod 1, 24 joint little longer than rest of limb, 7" nearly
half as long as 6%, Peraeopod 2 long and slender, a little longer than
the 34, Peduncle of uropods little longer than the subequal rami, with
48 Cumacea: 3. Bodotriidae, 7. Iphinoé, 8. Heterocuma
numerous spines on medial margin; exopod stouter but scarcely longer than
endopod, in which the 15* joint is about three-fourths as long as the more
slender 2™4, with 9 and 11 spines on them respectively, each apical spine
the longest. Colour greyish, with irregularly distributed red-brown spots.
L. 3 adult 6°2 mm, 9 unknown.
Gulf of Guinea, Anecho (Togo).
8. I, inermis O. Sars 1878 [.%., G. O. Sars in: Arch. Naturv. Kristian., v3
p. 508 t. 17, 18.
Pseudorostral lobes rather short, apically truncate; antero-lateral corners
acute in Q, rounded off in dG. Carapace not very elongate, with low carina,
but no teeth in either sex. Pedigerous segment 2 large, laterally acutely
produced over lower part of segment 1 on to the carapace. Eyelobe
broader than long, prominent in dG with 9 corneal lenses of which the middle
one is very large. Antenna 1, 34 joint nearly as long as 1S*, a little less
than twice the 2™4, flagellum 1-jointed, accessory minute, 2-jointed. Maxilli-
ped 3, 24 joint apically produced nearly to the end of the 4, which only
reaches half way along the 5 but is fringed with many plumose setae.
Peraeopod 1 of moderate length, 2"4 joint as long as rest of limb, 4th and
5th joints equal, 6 and 7 successively a little shorter. Peraeopod 2 scarce-
ly as long as the 34 pair. Peduncle of uropods decidedly longer than
the rami, the exopod a little longer than the endopod, of which the 1st joint is
about two-thirds as long as the second and carries 4 graduated spines, among
which in the O 4 spinules are distributed; the narrow 24 joint in the ©
has 11 graduated spines and in the CO a considerably larger number. Colour
red-yellow with brown and chalk-white shadings. L. O 9mm, GC 8 mm.
Goletta, depth a few fathoms.
8. Gen. Heterocuma Miers
1879 Heterocuma, Miersin: P. zool. Soc. London, p.22, 57 | 1893 H., T. Stebbing in:
Int. sci. Ser., v.74 p.304 | 1895 H., H. J. Hansen in: Ergeb. Plankton-Exp., v.2 G.c.
p- 56 | 1905 H., (part.), Calman in: Siboga-Exp., pars 36 p.6,8 | 1908 H., Zimmer in:
Ergeb. Tiefsee-Exp., v.8 p. 165 | 1910 H., Calman in: Ann. nat. Hist., ser.8 v.6 p. 612.
Pseudorostral lobes not strongly produced; first pedigerous segment
only dorsally distinct; telsonic segment not produced between bases of the
uropods. Eye present. Antenna 1 with third joint longer than second, flagella
two-jointed. Branchial leaflets numerous, in a straight series. Mavxilliped 3
with second and fourth joints distally produced laterally, the fifth distally
widened in the medial direction. Second and third peraeopods in both
sexes carrying imperfectly developed exopods. Pleopods of male with lateral
margin of endopod not produced into a narrow process. Endopod of uropods
two-jointed, first joint the longer.
2 species.
Synopsis of species:
J Carapace not granuliferous. .......+..-+-+-.. 1. H. sarsi
\ (Carapace: granulifezouse 2 sa 71 ok a> a eee 2. H. granulatum
1. H. sarsi Miers 1879 H.s., Miers in: P. zool, Soc. London, p.22, 58 t.3 f.3 |
1895 H. sarsii, H. J. Hansen in: Ergeb. Plankton-Exp., v.2 G.c. p.56 | 1910 H. sarsi,
Calman in: Ann. nat. Hist., ser.8 v.6 p.612 t.10 f. 1—13.
Cumacea: 3. Bodotriidae, 8. Heterocuma, 9. Cumopsis 49
Pseudorostral lobes blunt, reaching little beyond the eyelobe; antero-
lateral corners obtuse, especially in the O, and not reaching level with apices
of pseudorostrum. Carapace not elongate, smooth except for median carina
which is flanked by shallow depressions in front half and becomes doubled
in the hinder half. First pedigerous segment only exposed dorsally, this
and the rest devoid of carinae, 3¢ and 4*® segments in the dO with strong
lateral articulation. Pleon segments with feebly developed medio-dorsal and
lateral carinae, telsonic segment with dorsal depression. Eyelobe little longer
than broad, with large dark eye. Antenna 1 stout, 1s joint as long as 2nd and
32 combined, 34 a little longer than 2"4, Mandibles with many spines in
spine-row. Maxilla 1 with 2 apical filaments to the palp. Maxilliped 3, 224
joint with medial margin produced into a short tooth. Peraeopod 1, 2™4 joint
a little shorter than rest of limb, with a pair of stout spines on medial edge
near the base, and medial margin distally produced into a small tooth.
Peraeopod 2, 2"4 joint with setose stiliform exopod extending just beyond it,
the next joint absent or coalesced. Peraeopod 3, 2nd joint with similar exopod
not extending beyond its middle. Peduncle of uropods slightly shorter than
exopod, subequal to the endopod, which is setose on the lateral margin, the
longer 18° joint, like the rather shorter and much narrower 2nd, carrying about
a dozen spines successively larger towards the abruptly larger apical spine.
L. O adult 18 mm, © immature 17 mm.
Corean and Japanese Seas; lat. 329 41‘ — 33°19’ N., long. 128° 56’ — 12997'-5 He
depth 73—91 m.
2. H. granulatum Miers 1879 H. sarsi var. granulata, Miers in: P. zool. Soc.
London, p. 22,58 | 1910 H.s. var. g., ?H. g., Calman in: Ann. nat. Hist., ser.8 v.6 p.614 |
1912 H.s. var. g., Calman in: P. U.S. Mus., v.41 p. 612.
Like H. sarsi, but carapace dorsally carrying small tuberculiform
granules, largest along the median carina, gradually disappearing towards
the lateral margins; first 5 pleon segments with strongly marked dorsal,
dorso-lateral and lateral pairs of carinae, and a feeble ventro-lateral pair;
the appendages, especially the 18* peraeopods and the uropods a little more
elongated.
Corean Channel, lat. 32° 41’ N., long. 128° 57‘ E., depth 73 m, off Goto Island,
Korea Strait, depth 171 m.
9. Gen. Cumopsis 0. Sars
1878 Cumopsis, G. O. Sars in: Arch. Naturv. Kristian., v3 p. 465, 511 | 1893 C.,
1’. Stebbing in: Int. sci. Ser., v.74 p.302 | 1908 C., Zimmer in: Ergeb. Tiefsee-Exp.,
.8 p.159 | 1909 C., Stappers in: Cellule, v.25 p. 369 (alimentary system).
Pseudorostral lobes little produced. All five pedigerous segments distinct.
Telsonic segment not produced between the uropods. Eye well developed.
First antenna of @ with dense whorl of filaments round swollen base of
the flagellum. Third maxilliped with second joint not apically produced.
Second and third peraeopods with very small setiferous exopods. Pleopods
with process on lateral margin of the endopod. Uropods elongate, endopod
9-jointed, first joint longer than second, first joint of exopod rather long.
Hepato-pancreatic glands with two pairs of lobes.
2 species.
Das Tierreich. 39. Lief.: T. R. R. Stebbing, Cumacea. 4
50 Cumacea: Bodotriidae, 9. Cumopsis
Synopsis of species:
Carapace with 2 pairs of latero-dorsal arcuate folds . .. 1. C. goodsir
Carapace without lateral folds ........-... . . 2. C. longipes
1. C. goodsir (Bened.) ?1856 Cuma edwardsii (err., non Kroyer 1846), Bate in:
Ann, nat. Hist., ser.2 v.17 p.457 t.14 f.4 | ?1859 Vauwnthomsonia e., Bate in: Ann.
nat. Hist., ser. 3 v.3 p.273 | 1861 Bodothria goodsir, P. J. Beneden in: Mém. Ac. Belgique,
v.83 [nr. 3] p.76, 77, Bodotria goodsirii, p. 142, 166 t.13 f.1—16 | 1865 Bodotria goodsirii,
Fr. Miiller in: Arch. Naturg., v.3811 p.811—323 | 1869 Cuma goodsiri, A. Dohrn in: Jena. Z.,
v.5 p.56, 61, 63, 71, 75, 76 t. 2; t. 3 f.16,17 | 1878 & 1879 Cumopsis g., G.O.Sars in: Arch.
Naturv. Kristian., v.3 p.512; v4 p.1 t.19—21 | 1906 C. g., A. M. Norman & T. Scott,
Crust. Devon Cornwall, p.30 | 1909 C. g., Stappers in: Cellule, v.25 p.355.
Integument thin, flexible, finely squamose. Pseudorostral lobes meeting
a short space in front of eyelobe; antero-lateral corners shortly produced
in © with adjoining excavation, not produced in GO. Carapace with level
dorsal line, not carinate, but from the antero-lateral corners 2 pronounced
folds sweep round towards the centre, there joining a half elliptical fold
which extends from behind the frontal lobe nearly to the hind margin. Pedi-
gerous segment 1 exposed dorsally, laterally covered by the 2™¢ segment
(carapace leaving the five posterior segments exposed as complete rings in
themselves, Bate), 2"¢ segment having near the first a pair of semicircular
dorsal folds. Telsonic segment short, truncate. Eyelobe nearly heart shaped,
with dark eye and several corneal lenses. Antenna 1 with joints of peduncle
successively shorter, in @ distally widened, both flagella 2-jointed, the accessory
minutely, the principal distinguished in the dO by its swollen base and verti-
cillate equipment of radiating pellucid filaments. Antenna 2 in Q 2-jointed,
without setae on 18* joint, in GO the short-jointed flagellum reaching beyond
the body. Lower lip with slender spines on the incurved apices. Maxilli-
ped 3, 2"4 joint large and broad, longer than rest of limb, its lateral apex
carrying 5 plumose setae, 4” joint little produced, with 2 setae on lateral
apex. Peraeopod 1 comparatively short, 24 joint large, narrowing distally,
6% joint shorter than 5, Peraeopod 2 very small, with one joint missing
(the 2 anterior thoracic legs extremely long, the three posterior extremely
short, and all except the last having an exopod, Bate). Peraeopods 2 and
3 each with a very small rudimentary exopod carrying 3 short setae. Peduncle
of uropods very slender, much longer than the equal, linear rami, in which
the 18t joint of the exopod is as long as the 2"¢ of the endopod; armature
in spines and setae very much greater in the G. Colour dark violet, darkest
on hinder and lower part of carapace and 5% pleon segment. L. 5 mm.
British Isles, Belgium, Mediterranean. Sea-shore and surface of sea.
2. C. longipes (A. Dohrn) 1869 Cuma l., C. (Bodotria) l., A. Dohrn in: Jena. Z.,
v.5 p.77; p.8L t.8 f.1 | 1879 Cumopsis laevis, G. O. Sars in: Arch. Naturv. Kristian.,
v.4 p.8 t.22 | 1893 O.1, T. Stebbing in: Int. sci. Ser., v 74 p. 3803 | 1906 C.1., A.M.
Norman & T. Seott, Crust. Devon Cornwall, p. 29.
Very like C. goodsir, but carapace without lateral folds, having only
the semi-elliptical fold behind the frontal lobe; 24 pedigerous segment quite
smooth; antenna 1 of © more elongate, with 24 joint scarcely shorter than ilu
antenna 2 of Q with 1 plumose seta on 1%* joint; peraeopod 1 comparatively
longer and stronger, with 6 joint subequal to 5; uropods comparatively
longer; colour in © more intense. L. 6 mm.
British Isles (Clyde, S. Devon, on shore sands); Mediterranean (Syracuse, Naples,
Spezia); Bayonne Harbour.
Cum.: 3. Bodotr. — Bodotr. gen. dubia et spec. dubiae — 4. Ceratocumat., 1.Ceratocuma 51
Bodotriidarum genera dubia et species dubiae
Gammarus esca F. 1779 G.e., J. C. Fabricius, Reise Norweg., p. 249.
Antenna 2 longer than the body, colour blackish, herring-food in Norway.
Size of Gammarus pulex.
Cuma fasciata E. Hesse 1868 C.f., E. Hesse in: Ann, Sci. nat., ser.5 v.10
p. 358 (9); p.362 (C. fusciata).
Eyes black, indistinct. Colour deep rust brown mixed with bright green and
a white band. L. 6mm. Coast of France, in tidal pool.
Cuma parva EH. Hesse 1868 C.p., E. Hesse in: Ann. Sci. nat., ser.5 v.10
p. 359, 369 (Q or juv.).
Eyes black. Colour rust brown. L. 1mm. Coast of France, tidal pool.
Cuma punctata E. Hesse 1868 C. p., E. Hesse in: Ann. Sci, nat., ser.5 v.10
p. 355 (3), 869 (C. Ponctata).
Hyes red. Colour dull yellow with green and black markings. L. 3mm, Coast
of France, tidal pools.
Cuma rufa E. Hesse 1868 C. r., E. Hesse in: Ann. Sci. nat., ser.5 v.10
p. 356 (Q).
Kyes black. Colour deep rust brown. L. 12mm. Coast of France in tufts of
»Polysiphonia urceolata Lightf*.
Cuma terginigra E. Hesse 1868 C. t., E. Hesse in: Ann. Sci. nat., ser. 5
v.10 p. 849.
Eyes prominent but generally small and indistinct. Colour of G pale green,
passing through pink to a pleon of golden yellow traversed by a red ray, the whole
diversified by bands and points of black. Front of 2 apple-green framed in deep
black, hinder portion yellow with ray of red and black points. L. reaching 2°5 mm,
Br. 1 mm.
Coast of France, in sandy pools, sometimes abundant.
Cuma folini P. Fisch. 1872 (1874) C. Folini, P. Fischer in: Folin & Périer,
Fonds Mer, v.2 p.110 t.5 f.7,8 | 1907 C. F., Calman in: Tr. zool. Soc. London,
o 181 p: 3.
Probably the same as Bodotria arenosa, Goodsir (Calman).
4, Fam. Ceratocumatidae
1905 Ceratocwmidae, Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p.37 |
1912 Ceratocumatidae, T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 133, 142.
Female unknown. Male with small distinct telson, unarmed; only the
first two pairs of the peraeopods furnished with exopods; the seventh joint
in the two following pairs ending in a curved spine; pleon carrying five pairs
of pleopods; uropods with one-jointed endopod.
1 genus.
1. Gen. Ceratocuma Calman
1905 Ceratocuma (Sp. un.: C. horridwm), Calman in: Sci. Invest. Fish. Ireland,
1904 app. 1 p.37.
Fifth pedigerous segment without appendages; uropods very elongate.
1 species.
4*
52 Cumacea: 4. Ceratocumatidae, 1. Ceratocuma, 5. Leptocumatidae
1. C. horridum Calm. 1905 C. horrida, Calman in: Sei. Invest. Fish. Ireland,
1904 app.1 p.11, 37 t.4 f.57—75 | 1912 C. horridus, T. Stebbing in: Ann. S. Afr.
Mus., v.10v p. 143.
Pseudorostral lobes horizontal, bluntly truncate, meeting for a moderate
distance in front of the small eyeless eyelobe; a rather deep sinus is formed
by the bifid antero-lateral corner produced as far forward
as the pseudorostrum, the lower tooth initiating a long
lateral ridge on the dorsally flattened carapace. This is
diversified by 11 pairs of notable procurved teeth, 3 pairs
submedian, the rest lateral. All five pedigerous segments
are free, each except the first carrying a dorsal and a
lateral pair of spine-like processes, the lateral pairs of
the second and third segments being expanded at the
base and hollowed beneath to receive the end of the
flagellum of the second antenna. Fifth pedigerous segment
without trace of limbs, having ventrally behind a median
spine 2 contiguous eminences with the crescentic slit-like
apertures of the vasa deferentia. The first 4 pleon seg-
ments have each a pair of blunt recurved teeth at the
rear; the 5t® segment is nearly three times as long as
the sixth; the little round-ended telson becomes inconspicuous
when closed down over the anal opening. Antenna 1 has
the 18t joint large, distally widened, the flagellum 4-jointed,
accessory 1-jointed, both provided with sensory hairs.
Antenna 2 with last joint of peduncle nearly twice as
long as the preceding joint. The fully adult character
of both antennae makes it improbable that the absence
of the 5 peraeopods is due to immaturity. Mandibles
with strong molar and numerous spines. Palp of maxilla 1
with 2 filaments. Maxilliped 1 with the epipod scantily
provided with branchial leaflets. Maxilliped 2 with full
number of joints. Maxilliped 3, the long 2”? joint nar-
rowed distally, not apically produced. Peraeopod 1 has 34
joint longer than 4%, 5% equal to both combined, the
short 6 furnished with 2 peculiar processes each armed
with a dense tuft of radiating setae. Peraeopod 2 very
slender, the long 24 joint narrowing rapidly about the
middle, the 5 joint much more than twice the 6 and
Fig. 26. 7¢» cgombined. Fifth and sixth joints much longer in the
C.horridum(3)Calman. 4 than in the 34 peraeopod. Pleopods feebly developed,
in the South African specimen only the first pair clearly dis-
cernible, although the antennae evince maturity. Uropods two-fifths of body's
length, the subequal rami more than thrice as long as the peduncle, each serrate
on medial margin, tapering to a fine point, with minute terminal spine. L. 4°05 mm.
Irish Sea, 77 miles W. N. W. of Achill Head, county Mayo, 699 m; South Africa,
Cape Natal N. by E. about 24 miles, depth 805 m.
én
5. Fam. Leptocumatidae
1912 Leptocumatidae, T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 133.
Telson wanting. Accessory flagellum of antenna 1 very small. Antenna 2
of Q 8-jointed. Palp of maxilla 1 ending in 2 filaments. Peraeopods 1—4
Cumacea: 5. Leptocumatidae, 1. Leptocuma 53
with exopods in both sexes, all well developed, except those on the 4" pair
in the female. Only 3 pairs of pleopods in the male. Endopod of uropods
2-jointed.
1 genus.
1. Gen. Leptocuma 0. Sars
1873 Leptocuma (Sp. un.: L. kinbergit), G.O. Sars in: Svenska Ak. Handl., n. ser.
wll nr.5 p.24 | 1878 & 1879 L., G.O.Sars in: Arch, Naturv. Kristian., v3 p. 465;
v.4 p.12 | 1887 L., G. O. Sars in: Rep. Voy. Challenger, v.19 pars 55 p.6 | 1908 L.,
Zimmer in: Ergeb. Tiefsee-Exp., v.8 p.165 | 1912 Z., Calman in: P. U.S. Mus., v.41 p. 615.
General form slender, elongate, diminishing very gradually from cara-
pace to pleon. Pseudorostral lobes produced shortly or not at all in front
of the semicircular eyelobe. First pedigerous segment exposed only dorsally.
Exopod on peraeopod 4 of © rudimentary.
2 species.
Synopsis of species:
{ Uropods, 2nd joint of endopod more than half as long as Ist 1. L. kinbergii
) Uropods, 2nd joint of endopod less than half as long as 1st. 2. L. minus
1. L. kinbergii O. Sars 1873 L.k., G.O.Sars in: Svenska Ak. Handl., n. ser.
v.11 nr.5 p.24 t.6 f.29—33 | 1893 L.k., T. Stebbing in: Int. sci. Ser., v.74 p. 304 |
1907 L. k., Calman in: Tr. zool. Soe. London, v. 181 p.30 | 1908 L. kinbergi, Zimmer
in: Ergeb. Tiefsee-Exp., v.8 p.165 | 1912 L. kinbergii, Calman in: P.U.S. Mus., v4l p.616.
Body so much compressed in front that the whole in dorsal view appears
almost parallel sided. Pseudorostral lobes little or in maturity not at all
advanced in front of eyelobe, apices
broadly rounded and separated by
a notch from the bluntly rounded
antero-lateral advanced corners. .
Carapace very small. quite smooth.
Pedigerous segment 2 infero-
laterally stretching over 18* seg-
ment on to the carapace; 34
segment below overlapping the
gud and 4, ‘Telsonic segment
not much shorter than the pre-
ceding segment, both narrower
than those which precede. The
flattened semicircular — eyelobe
showing no corneal lenses. An-
tenna 1, flagellum 3-jointed, acces-
sory minutely 1-jointed. Peraeo- Fig. 26.
pod 1, 24 joint about equal to L. kinbergii (2) Sars.
rest of limb, 7 scarcely half as
long as 6%, Peraeopod 2, 3° joint quite short. Exopod of peraeopod 4
small, one-jointed, with small setae. Peduncle of uropods nearly but not
quite so long as the laminar rami (or may equal the exopod, Calman)
endopod a little longer than exopod, of which the 18* joint is not very long,
while the 2"4 begirt with spines and setae is somewhat longer than the first
joint of the endopod; the latter joint on medial margin above has a set of
5 spines and below a graduated series of 12, the second joint about two-
54 Cumacea: 5. Leptocumatidae, 1. Leptocuma, 6. Chalarostylidae, 1. Chalarostylis
thirds as long, carrying 15 spines, of which the apical are large. The 18 spines
on lateral margin of exopod’s 2"4 joint are blunt-ended. L. o 14—22°5 mm.
3 unknown.
South Atlantic, off mouth of Rio Plata, depth 94 m; Straits of Magellan.
2.L.minus Calm. 1912 LZ. minor, Calman in: P.U.S. Mus., v.41 p.616 tf. 14—20.
Pseudorostral lobes truncate, meeting a short space in front of the small
eyelobe; with a small sinus and slightly advanced antero-lateral angle. Carapace
short. Pedigerous segments 2, 3, with side-plates greatly expanded. Telsonic
segment considerably shorter than the one preceding it, about 3/4s of peduncle
of uropod. Eye pigmented. Antenna 1, flagellum 2-jointed, accessory 1-jointed.
Antenna 2 of © 3-jointed, last joint bluntly conical, twice as long as width
at base. Mandible with premolar part long, about 6 spines in spine-row.
Maxilliped 1 with about 8 branchial leaflets. Maxilliped 3 with 2"4¢ joint
slightly produced. Peraeopod 1 with 24 joint slightly longer than rest of
limb, 7 little shorter than 5 or 6, Peraeopod 2, 5 joint little longer
than 4), a little shorter than 6 and 7 combined, 7** not quite as long as
6, Peraeopod 4 of © with rudimentary exopod of 2 joints, the 2"? minute.
Exopod of uropods a little shorter than the pedunele, a little longer than
the endopod; 18* joint of the latter more than twice as long as the 2".
L. Oo 75 mm, co 64 mm.
U.S.A., Wood’s Hole; Gloucester Harbor, depth 15 m.
6. Fam. Chalarostylidae nom. nov. T. Stebbing
1912 Hemilampropidae, T. Stebbing in: Ann. S. Afr, Mus., v. 10v p. 133, 143.
Telson ample, with more than 2 apical spines. Antenna 1 with both
flagella well developed. Palp of maxilla 1 with 2 filaments. Peraeopods 1—4
with exopods in both sexes well developed except those on the 34 and 4* pairs
of the female which are rudimentary. Peraeopod 1 with 24 joint much shorter
than the rest of the limb. Male with only 3 pairs of pleopods. Uropods
with the endopod 3-jointed.
2 genera.
Synopsis of genera:
Telson small in relation to the peduncle of the uropods 1. Gen. Chalarostylis
Telson large in relation to the peduncle of the uropods 2. Gen. Hemilamprops
1. Gen. Chalarostylis A. M. Norman
1879 Chalarostylis, A. M. Norman in: Ann. nat. Hist., ser.5 v.3 p.65 | 1900 C.,
G. O. Sars, Crust: Norway, v.3 p.26 | 1912 C., T. Stebbing in: Ann. S. Afr. Mus.,
v.10v p. 144.
Telson short, with three apical spines. Uropods remarkably long and
slender. Female unknown.
1 species.
1. C. elegans Norm. 1879 C.e., A. M. Norman in: Ann. nat. Hist., ser.5 v3
p. 66 | 1912 C.e., T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 144.
Pseudorostral lobes forming no sinus. Carapace not deep but wide,
spinulose, anteriorly with medio-dorsal crest of about 10 small spines. Anterior
margin of pedigerous segments smooth, side-plates of fifth not produced
Cumacea: 6. Chalarostylidae, 1. Chalarostylis, 2. Hemilamprops 55
backwards. Pleon quite smooth; telson only as long as the sixth and little
over half the fifth pleon segment, with 2 pairs of lateral and 3 apical spines.
Antenna 1, 1%‘ joint spinose, especially below. Antenna 2 as long as cepha-
lothorax, strongly ciliated. Peraeopod 2 (in original description called 1s* by
error) with slender 2"¢ joint, not spined, 3¢ joint very short with 1 long spine,
4th longer with similar spine, 5 nearly as long as 3 preceding joints com-
bined and much longer than the 2 following together, armed with 10 spines,
2 of them much longer than the rest, 6 joint very short, one-third of 7.
Pleopods with long peduncle, and setose rami. Peduncle of uropods thrice
as long as telson, with 24 spines on the medial margin, endopod with i
joint as long as the telson, 8 spines on medial margin, ond with 2, 34 with
no spines: exopod reaching middle of 34 joint of endopod. L. adult G rather
more than 8 mm.
Off Rockall, depth 199 m.
2. Gen. Hemilamprops G. 0. Sars
1883 Hemilamprops, G. O. Sars in: Forh, Selsk. Christian., nr. 18 p. 11, 55 | 1896
H., J. Bonnier in: Ann. Univ. Lyon, v.26 p. 546, 549 | 1908 H., Zimmer in: Ergeb. Tiefsee-
Exp., v.8 p. 169, 171 | 1912 H., T. Stebbing in: Ann. S. Afr. Mus., v.10v p. 144.
Distinguished from Chalarostylis by the greater development of the telson.
6 species.
Synopsis of species:
1 Telson with more than 5 apical spines — 2.
Telson with not more than 5 apical spines — 8.
5 Hye large, red, with 8 lenses; telson with 8 apical spines 1. H. rosea
=) Eye whitish, without lenses, telson with only 6 apical spines 2. H. assimilis
3 Eye wanting — 4.
Eye present — 5.
A Telson with 5 apical spines . ... +... © a cet el fens hat OPEL. TLOPM ATT
Telson with only 3 apical spines... - 2 + + + e+ ee > 4. H. pellucida
Carapace without pair of carinae curving round the frontal
5 IGN. -Gice S Sreld) o) ole Bo olole aha seo Bech sec 5. H. cristata
Carapace with pair of carinae curving round the frontal
WOO Be ARG, Hae SePOOIES cen O Pha o Ge. he RRR er aieo eee mac 6. H. uniplicata
1. H. rosea (Norm.) 1863 Vawnthompsonia r. + Cyrianassa elegans, A. M. Norman
in: Tr. Tyneside Club, v.51v p.271 t.18 £.1—8 9; p.275 t.13 £4—9¢ | 1865 Lamprops
x, G.O. Sars in: Forh. Selsk. Christian., 1864 p.189 | 1883 Hemilamprops r., G20;
Sars in: Forh. Selsk. Christian., 1882 p.11, 55 | 1897 H.r., T. Scott in: Rep. Fish.
Board Scotl., v.15 p.134 | 1899 H.r., G. O. Sars, Crust. Norway, v.3 p. 22 t. 12—14 |
1900 H. r., T. Scott in: Ann. Scott. nat. Hist., v9 p.219 | 1905 H. r., Calman in: Sci.
Invest. Fish. Ireland, 1904 app. 1 p. 41.
Integument thin, semipellucid. Carapace, especially in Q, shorter than
the group of pedigerous segments, dorsal line straight, horizontal, sides
smooth, pseudorostral projection almost obsolete. Second and third pedigerous
segments the largest. Telson rather broad, lamellar, equal in length to peduncle
of uropods, not rapidly narrowing to the rounded apex, which carries 4 pairs
of spines, the outermost the longest, lateral margins generally with 2 pairs
of setiform spines. Eye large, round, red, with 8 lenses. Antenna 1 with
principal flagellum 3-jointed, its 1* joint in the d having a fascicle of 6 setae,
56 Cumacea: 6. Chalarostylidae, 2. Hemilamprops
accessory 2-jointed. Antenna 2 in © with 1S* joint carrying 3 plumose setae,
its length subequal to 3 following joints combined. Peraeopod 1 fully twice
as long as the carapace, 2"4¢ joint much shorter than the rest combined, among
which the 6t joint is much the longest. In peraeopod 2 the 6" joint is
shorter than any but the 34. In the 34 and 4* pairs the 2™¢ joint is much
longer than the rest of the limb, slender in the © with small 2-jointed exopod
attached near the base, but stout in the G with exopod fully developed,
the 3¢ pair in the GO carrying on the 34 joint 2 smooth falciform spines.
Peraeopod 5 small, normal. Uropods slender, peduncle not longer than the
exopod, in which the 1s* joint is about two-thirds as long as the second,
the two together not quite equal to the first 2 of the endopod, which has
about 15 spines on the medial margin of its long 1%t joint, 4 on the second,
and the still shorter 3¢ furnished with 5 little spines round the apex. Colour
crimson, in ramified stellate patches. L. Q 6mm, CG 7 mm.
Norway, Finmark, Shetland, N. E. England, W. of Scotland and Ireland. Depth
38 to 364 m.
2 H.assimilis O.Sars 1883 H. a., G.O.Sars in: Forh. Selsk. Christian., 1882
nr.18 p.11,55 t.1 f.98, 24 | 1899 H.a., G. O. Sars, Crust. Norway, v.3 p.23 t.15|
1905 H.a., Calman in: Sci. Invest. Fish. Ireland, 1904 app. 1 p. 41.
Near to /7. rosea, but eye imperfect, without lenses, pigment whitish;
peraeopod 1 rather less slender, 6* joint scarcely as long as, instead of longer
than the 3 preceding joints combined; peraeopod 3, with the 2 pecuhar
spines on 34 joint in GO more slender, not falciform, distally annulate and
on one side denticulate; telson more narrowed distally, with single pair of
slender lateral spines, only 6 apical, the outermost pair much the longest;
hody semipellucid, unpigmented. L.o 5mm, G 6mm (Sars, 1899); © 6°5 mm
(Sars, 1883).
East Finmark, at Vardé, 56—94 m (Sars, 1883), Vard6, Hammerfest and Hasvig,
113—877 m (Sars, 1899).
3. H. normani Bonnier 1896 H. n.,.J- Bonnier in: Ann. Uniy. Lyon, v.26 p.546
t.29 f.3a—r | 1900 H.n.,-G. O. Sars, Crust. Norway, v3 p.25 | 1905 ? H. cristata,
Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p.41 | 1908 H. normant, Zimmer in:
Ergeb. Tiefsee-Exp., v.8 p.171 |. 1912 H.n,, T. Scott in: Sci. Invest. Fish. Scotl., 1911
urs l p..3: ;
Pseudorostral lobes little prominent, smooth, apically rounded. Carapace
smooth, except for medio-dorsal serrate carina reaching nearly half way along
with 26 teeth, of which 6 are on the prominent but sightless eye-lobe;
2 slight submedian furrows on the hinder half of the carapace. The telson
is about four-fifths of the length of the peduncle of the uropod, its sides
serrate, the hinder half carrying 7 pairs of strong spines in the CG, 5 in
immature Q, the apex 5 spines of which the central one is much the longest.
Antenna 1 with flagellum of 5 short joints, accessory 3-jointed, two-thirds
of length of the principal. Antenna 2 of © with a single plumose seta on
each of first 2 joints. Maxilliped 2 with 34 joint fused or lost. . Peraeo-
pods 3, 4, with rudimentary 2-jointed exopods. Peduncle of uropods shorter
than the rami, of which the endopod is only a little the longer, carrying
on the medial margin of its 3 joints 8, 4, and 2 spines respectively; the
exopod, as figured, has the first joint somewhat longer than the second,
L. d 10 mm, © a little less. None of the specimens fully adult.
Bay of Biscay, depths 650 and 950 m.
or
~]
Cumacea: 6. Chalarostylidae, 2. Hemilamprops
4. H. pellucida Zimmer 1908 A. p., Zimmer in: Ergeb. Tiefsee-Exp., v8
p. 171, 172 t. 39 f.53, 54; t.40 f.55—59 | 1910 H. p., T. Stebbing in: Ann.S. Afr. Mus.,
v.6 p.415 | 1912 H. pellucidus, T. Stebbing in: Ann.S. Afr. Mus., v. lOv p. 144 t. 52 (4).
Integument pellucid, delicate, microscopically scabrous. Pseudorostral
lobes short, subacute, with serrate edges. Carapace in both sexes rather
deep, the small triangular eyelobe without lenses, its medio-dorsal line convex,
carrying 4 or 5 forward-pointing denticles, immediately followed by a nearly
level line of 20 denticles reaching beyond the middle
of the carapace and succeeded by a groove between
the inflated branchial regions. Pedigerous segments
combined shorter than the carapace. Pleon longer than
the preceding portion of the body. Telson with 3 long
apical spines, its lateral margins conspicuously denti-
culate, on the nearly parallel-sided hinder half carrying
6 to 8 pairs of spines, subject to some irregularity;
the telson in the adult male apparently longer than
in the O, with a narrower base. Antenna 1 with
serrate edge to the large 1’ joint, 3¢ joint small, in
3 not longer than broad, flagellum in © 5- or 6-jointed,
accessory of 3 joints, the 3¢ microscopic, flagellum in Jd
4-jointed, accessory of 3 well developed joints. An-
tenna 2 O-small, 4-jointed, with 1 seta on the rather
large 18¢ joint and 1 on the small 2°47. Antenna 2
in subadult G showing a flat process on the side of
the basal joint; the annulated flagellum long. Mouth
organs not affording distinctive characters. Peraeopod 1,
2nd joint much curved, its convex border distally
serrate. Peraeopod 2, 2"4 joint shorter than suc-
ceeding joints combined, of which the strongly spined
5 is longer than the short 6 and long narrow 7
combined. Peraeopods 3 and 4 showing the characteristic
sexual differences, but the 3¢ joint in peraeopod 3 of oh
the subadult male not showing any flattened appendages. oe itd Zanes
Pleopods as in Paralamprops serratocostata (p. 59). telson and uropod (after
Peduncle of uropods in © longer than the telson with Stebbing)-
its terminal spines and longer than both rami, but in the
subadult G not longer than the telson including its median apical spine and
not quite so long as the endopod, which in this sex has the 34 joint a little
longer than the 2™4, and the first much longer than these 2 combined; the
exopod, with its first joint considerably longer than the 2™4, is equal to the
peduncle in the G, but shorter than it in the go. L. 85 mm.
S. Africa, outside the Agulhas Bank, depth 564 m, and off Cape Point N. 81° E. 32 miles.
5. H. cristata (O. Sars) 1870 Lamprops c., G. O. Sars in: Forh, Selsk. Christian.,
1869 p.157 | 1872 ZL. c., G.O.Sars in: Forh. Selsk. Christian., 1871 p.270 | 1879
L.c., A. M. Norman in: Ann. nat, Hist., ser.5 v3 p.68 | 1883 Hemilamprops c., G. O.
Sars in: Forh. Selsk. Christian., nr. 18 p.11,56 | 1886 H. ¢., G.O.Sars in: Norske
Nordhavs-Exp., v.6 Crust.2 p.19 | 1896 H.c., J. Bonnier in: Ann. Univ. Lyon, v.26 p.549 |
1900 H.c., G.O. Sars, Crust. Norway, v.3 p.25 t.18 | 1905 ?A.c., Calman in: Sci.
Invest. Fish. Ireland, 1904 app. 1 p.41 |.1908 H.c., Zimmer in: Ergeb. Tiefsee-Exp.,
v.8 p.171 | 1912 H.c., T. Scott in: Sci. Invest. Fish. Scotl., 1911 nr. 1 p.3.
In near agreement with H. normani (nr. 3) and H. pellucida (nr. 4), but
distinguished from both in the ¢ by absence of the medio-dorsal longitudinal
58 Cumacea: 6. Chalarostylidae, 2. Hemilamprops, 7. Paralampropidae
denticulation of the carapace, agreeing in both sexes with H. pellucida but
differing from H. norman: in the apical spines of the telson; having in peraeopod 1
the 7th joint twice as long as the 5‘, and notably longer than the 6, while
in a © specimen of H. pellucida, 8 mm long, this 7 joint is notably shorter
than the 6t and less than once and a half as long as the 5%. The fragi-
lity of this limb in the southern species commonly makes it unavailable for
comparison. Peraeopod 3 in the G appears to agree with H. pellucida in the
want of peculiar appendages on the 3¢ joint. Telson comparatively short,
scarcely longer than the last segment, not decidedly longer as in the other
two species, with 2 or 3 pairs of lateral spinules, and 3 subequal apical
spines, much larger than the spinules (Sars), or with 3 in G, 4 or 5 in O
pairs of lateral spines, and 3 apical, not greatly longer than the others, the
median sometimes a little the longest (Calman, but perhaps referring to
HH. normani). Colour light orange or golden yellow, eye distinct, bright red
(G. O. Sars), without pigment (Calman). L. 7 mm.
Norway, N. to Lofoten Islands, in depths 150 to over 188 m; South of Rockall, 1152 m.
6. H. uniplicata (O. Sars) 1872 Lamprops u., G.O. Sars in: Forh. Selsk.
Christian., 1871 p.270 | 1883 Hemilamprops u., G. O. Sars in: Forh. Selsk. Christian.,
1882 nr. 18 p.11, 56 | 1886 H.wu., G.O. Sars in: Norske Nordhavs-Exp., v.6 Crust. 2
p.18 | 1896 H. u., J. Bonnier in: Ann, Univ. Lyon, v.26 p.549 | 1899 H. u., G. O. Sars
in: Crust. Norway, v.83 p.24 t. 16,17 | 1905 H. u., Calman in: Sci. Invest. Fish. Ireland,
1904 app.1 p.41 | 1908 H.w., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 171.
Body long and slender. Pseudorostral lobes shortly produced, quadrate.
Frontal lobe of carapace slightly keeled medio-dorsally, and almost surrounded
by a pair of conspicuous curved carinae, which approach without meeting
one another behind the middle of the carapace. Hyelobe very small, trian-
gular. Pleon considerably longer than preceding portion of the body, slightly
tapering. Telson much longer than the sixth segment, subequal to peduncle of
uropods, evenly tapering, with 6 or 7 pairs of lateral spines, and 3 subequal rather
longer spines on the apex. Eye very small, but distinct, red. Antenna 1 of ©
with principal flagellum and accessory both 3-jointed, 34 joint of accessory minute,
flagellum of GO 5-jointed, 18* joint with row of 7 long filaments, accessory
of 3 joints, with perhaps a microscopic 4". Sars figures on 2"¢ joint of antenna 2
in G a laminar process similar to that in . pellucida. Peraeopod 1, 2"4 joint
strongly curved, 7% joint longer than 6, more than twice as long as 5%.
Peraeopod 3 in G without peculiar appendages on the 34 joint but with
special sculpture of the widened short 6% joint. Peduncle of the uropods
subequal to the exopod, in which the 18* joint is rather more than */,% of
the second, both combined being much shorter than the endopod, in which
the 1st joint is far longer than the combined 2"¢ and 34 Colour bright
orange. L. 7 mm.
Norway, N. to Lofoten Islands and Norwegian N. Atlantic, 113—785 m, W. coast
of Ireland, depth 364 m.
7. Fam. Paralampropidae
1912 Paralampropidae, T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 183, 143.
Telson large, with more than 2 apical spines. Antenna 1 with both
flagella well developed. Antenna 2 of female 5-jointed. Maxilla 1 without
palp. Peraeopods 1—4 with exopods in both sexes well developed except
those on the 34 and 4th pairs of the female which are rudimentary. Only
3 pairs of pleopods in the male. Uropods with the endopod 3-jointed.
1 genus.
Cumacea: 7. Paralampropidae, 1. Paralamprops 59
1. Gen. Paralamprops 0. Sars
1887 Paralamprops, G. O. Sars in: Rep. Voy. Challenger, v.19 pars 55 p. 6, 26 |
1908 P., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 169, 170.
Carapace broad, depressed, without antero-lateral angles. Pleon long and
slender. Lelson with three apical spines. Eye wanting. Fifth peraeopod normal.
2 species.
Synopsis of species:
{ Telson not nearly reaching end of peduncle of uropod 1. P. serratocostata
\ Telson reaching end of peduncle of uropod .... - 2. P. aspera
1. P. serratocostata (O. Sars) 1885 Hemilamprops serrato-costata, G.O.Sars
in: Rep. Voy. Challenger, v.1 pars2 p.742 (nom. nud.) | 1887 Paralamprops s., G. O.
Sars in: Rep. Voy. Challenger, v.19 pars 55 p.9, 26 t.2 f.6—13; t.3 | 1908 P.s., Zimmer
in: Ergeb. Tiefsee-Exp., v. 8 p. 169.
as , 2
Fig. 28. Fig. 29.
P. serratocostata Sars (di juv.). P. serratocostata (2) Sars, antenna 1.
Pseudorostral lobes short, somewhat upturned, denticulate. Carapace
nearly parallel-sided, sculptured by 7 longitudinal serrate carinae, one median
on anterior half, an arcuate submedian pair on the hinder half, flanked by
a longer subdorsal pair, and a lateral pair encircling the whole carapace.
60 Cumacea: 7. Paralampropidae, 1. Paralamprops, 8. Platysympodidae
Between the 2 latter pairs is a small interrupted serrate carina, and beneath
the encircling keel there is a small smooth one. Pedigerous segments well
developed, first three with transverse rows of denticles, and their infero-
lateral angles acute. Pleon with telson much longer than preceding part of
the body. Telson very slender, much longer than 6 pleon segment, but
much shorter than the peduncle of the uropods, armed with 10 pairs of
spines laterally, the large last pair similar to the
3 apical spines. Ocular lobe very small, conical.
Antenna 1 elongate, apices of 1S and 24 joints
denticulate, flagellum of 5 long joimts and a ter-
minal very short one, each of the last 2 having
an annulated sensory appendage; accessory 3-jointed
nearly as long as principal. Antenna 2 of O with
basal joint perhaps subdivided, the next joint
short with 2 plumose setae, the terminal joint long
and slender, the penultimate denticulate. Maxilli-
Fig. 30. ped 1 with 6 branchial leaflets on the epipod.
P. serratocostata (2) Sars, | Peraeopod 1 long and slender, 7 joint very slender,
MO ee as long as the 5%, each shorter than the 6*, all three
with slender setae on the- medial margin. Peraeo-
pod 2 with long 5t* joint equal to 6t and 7 combined, 6% very short.
Peraeopod 5 very small. Pleopods with short rami, the endopod 1-jointed with
projection crossing the 2-jointed exopod just below its 18* joint, each ending
in 4 setae. Pedunele of uropods long, but scarcely longer than the endopod,
both carrying numerous unequal spines on the medial margin, 1% joint of
endopod nearly twice as long as the other 2 joints combined; exopod as long
as first 2 joints of endopod, its 18* joint very short. L. @ 12 mm.
Kerguelen Islands, depth 232 m.
2. P. aspera Zimmer 1907 P.a., Zimmer in: Zool. Anz., v.31 p.373 | 1912 P.
asper, T. Stebbing in: Ann. 8. Afr. Mus., v. 10v p. 143.
Very like P. serratocostata, but with 9 carinae on the carapace, the
encircling lateral pair, a 2"¢ within meeting this forward on the pseudorostral
lobes, a third further in on the hinder half of the carapace, a 4% within
this, parallel to it but a little further forward, and a median carina on the
front half; the third carina continued on the pedigerous segments; of these
the antero-lateral angles are not acutely produced; pleon dorsally rough
with denticles; telson reaching end of peduncle of uropods; antenna 1 long,
but not so long as in the preceding species. Colour yellowish white. L.,
young GO, about 9 mm.
German South Pole Expedition, Winter Station (lat. 66° 2’S., long. 89° 38’ E.),
depth 385 m.
8. Fam. Platysympodidae
1899 & 1900 Platyaspidae, G. O. Sars, Crust. Norway, v.3 p.3; p.26 | 1905 P.
(part.), Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p.42 | 1912 Platysympodidae,
T. Stebbing in: Ann. S. Afr. Mus., v. LOV p. 133, 143.
Telson large with more than 2 apical spines. Antenna 1 with both
flagella well developed. Antenna 2 of female 4-jointed. Maxilla 1 without palp.
Exopods on peraeopods 1—4 in the male, only on the 1st pair in the female.
Only 3 pairs of pleopods in the male. Uropods with the endopod 3-jointed.
1 genus.
Cumacea: 8. Platysympodidae, 1. Platysympus 61
1. Gen. Platysympus T. Stebbing
1870 Platyaspis (non Platyaspis Agassiz 1846 pro: Platynaspis Redtenbacher Col.
1843) (Sp. un.: P. typica), G.O. Sars in: Forh. Selsk. Christian., 1869 p.158 | 1900
P., G. O. Sars, Crust. Norway, v.3 p.26 | 1905 P. (part.), Calman in: Sci. Invest. Fish.
Ireland, 1904 app. 1 p.42 [1908 P., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 159, 170 |
1912 P., Calman in: P. U.S. Mus., v.41 p.634 | 1912 Platysympus, T. Stebbing in:
Ann. S. Afr. Mus., v. 10v p. 143.
Carapace broad, depressed, first pedi-
gerous segment conspicuous, flagellum of
antenna 2 in the male not very elongate.
2 species.
Synopsis of species:
as the peduncle of the
ULOPOUS! Sie. 4) ss wel aby pleus
\ Telson little more than half
| as long as the peduncle
of the uropods. . . . 2. P.brachyurus
1. P. typicus (O. Sars) 1870 Platyaspis
typica, G.O. Sars in: Forh. Selsk. Christian., 1869
p. 158 | 1900 P. t., G. O. Sars, Crust. Norway, v.3
p. 27 t.19, 20 | 1905 P. ¢., Calman in: Sci. Invest.
Kish. Ireland, 1904 app. 1 p.12, 42 | 1907 P. t¢.,
Zimmer in: Zool. Anz., v.81 p.374 | 1912 P.t.,
Calman in: P.U.S. Mus., v.41 p. 634 | 1912 Platy-
sympus typicus, T. Stebbing in: Ann. S. Afr. Mus.,
v. 10v p. 144.
Integument squamose. Carapace twice
as long as the pedigerous segments combined,
or in © even more than twice, the shape
quadrangular oval, widest in front in GC, to
the rear in Q, pseudorostrum about hori-
zontal, eyelobe small, the medio-dorsal line
| Telson two-thirds as long
Fig. 31. Higs325
P. typicus (2) (Sars). P. typicus (Sars), maxilla 1.
from this to the rear gently convex, obtusely carinate, the sides lamellarly
expanded, carinate, probably fringed as in Platytyphlops peringueyi. Pedigerous
segment 1 in adults nearly as long as the 3 following segments combined, with
saddle-like dorsal depression; 2°¢ segment with double dorsal carina (Calman).
Pleon very slender, subequal in length to preceding part of the body. Telson
62 Cumacea: 8. Platysympodidae, 1. Platysympus, 9. Leuconidae
two-thirds as long as peduncle of uropods in G, rather longer than that pro-
portion in ©, the side margins very faintly serrate, the apex carrying three
subequal spines. Antenna 1 Q, 1% joint equalling the 2™4 and shorter 34
combined, flagellum 3-jointed, accessory 2-jointed; in GC, 34 joint of peduncle
broader than long, flagellum 5-jointed, 18* joint with brush of long setae,
accessory 3-jointed. Antenna 2 Q, 1 plumose seta on each of first 3 joints,
4th joint tapering, as long as 2™¢ and 34 combined; in C only just reaching
the pleon. Peraeopod 1, 224 joint nearly equal to rest of joints combined,
34 very small, 4th, 5%, 6t subequal, 7 shorter. Peraeopod 2 narrow,
rather long, 2"4, 5% and 6% joints long, the rest short, strong spines on 5%,
Peraeopod 5 small, but not rudimentary. Peduncle of uropods rather longer
than the endopod which carries 2, 1, 1 spines on the serrate medial margin
of its 3 joints respectively; exopod as long as first 2 joints of the endopod.
Colour light yellow, with whitish band across carapace. L. O 6 mm, CO 5°5 mm.
Norway, to Lofoten Islands, depth 226—753 m; W. of Ireland, depths 364 and
728 m; Mediterranean, near Capri 750—1100 m.
2. P. brachyurus (Zimmer) 1907 Platyaspis brachyura, Zimmer in: Zool.
Anz., v.31 p.374 | 1912 Platysympus brachyurus, T. Stebbing in: Ann. 8S. Afr. Mus.,
v.10v p. 144.
General habit as in P. typicus, but carapace not as there twice as long
as group of pedigerous segments, and its lateral margins obtuse, the pseudo-
rostrum rather longer, the first pedigerous segment shorter, the telson little
more than half as long as the peduncle of the uropods, with its side margins,
like the medial margin of the uropods, coarsely serrate. Uropods with 3, 1,
1 spines on the 3 joints of the endopod respectively; exopod missing. Colour
white, very pellucid. L. © about 5 mm, dO unknown.
German South-Pole Expedition, Winter Station (lat. 66° 2’S., long. 89° 38’ E.),
depth 385 m.
9. Fam. Leuconidae
1878 & 1879 Leuconidae, G. O. Sars in: Arch. Naturv. Kristian., v3 p.466; v4
p- 22 | 1883 Z., G. O. Sars in: Forh. Selsk. Christian., nr. 18 p.12 | 1887 Z., G. O. Sars
in: Rep. Voy. Challenger, v.19 pars55 p.8, 32 | 1893 Z., T. Stebbing in: Int. sci.
Ser., v.74 p. 304, 312 | 1900 Z., G. O. Sars, Crust. Norway, v.3 p. 28 | 1907 L., Calman
in: Tr. zool. Soc. London, v.181 p.31 | 1908 Z., Zimmer in: Ergeb. Tiefsee-Exp., v. 8:
p-176 | 1909 Z., Zimmer in: Ergeb. Schwed. Siidp.-Exp., v.6 nr.3 p.5 | 1912 L., T. Stebbing
in; Ann. 8. Afr. Mus., v. 10v .p. 133, 155.
All the pedigerous segments distinct. Telson wanting. Eye wanting.
First antenna with accessory flagellum small. Mandibles broad at the base,
spines few. Palp of first maxilla with a single filament. Branchial leaflets
few. Exopods on first four pairs of peraeopods in the male and the first
three in the female. Two pairs of pleopods in the male. Uropods with the
endopod two-jointed.
4 genera.
First antenna not geniculate ....-+.-+.. . . - 1. Gen. Leucon
\ First antenna geniculate — 2.
J First antenna geniculate between 2nd and 34 joints. . 2. Gen. Eudorella
\ First antenna geniculate between 1st and 2nd joints — 3.
Pseudorostrum obsolete or short, vertical, truncate . . 8. Gen. Eudorellopsis.
3 | Pseudorostrum obliquely upturned above the carapace 4. Gen. Pseudoleucon
Cumacea: 9. Leuconidae, 1. Leucon 63
1. Gen. Leucon Kroyer
1846 Leucon (part.), Kroyer in: Naturh. Tidsskr., ser.2 v. 2m p.208 | 1855 L.,
W. Liljeborg in: Ofv. Ak. Férh., 1855 v.12 p.121 | 1865 L., G. O. Sars in: Forh.
Selsk. Christian., 1864 p.176 | 1871 Z., G.O.Sars in: Svenska Ak. Handl., n. ser.
v.9 nr. 13 p.38 | 1879 Z., G. O. Sars in: Arch. Naturv. Kristian., v.4 p.23 | 1887 L.,
G. O. Sars in: Rep. Voy. Challenger, v.19 pars 55 p.33 | 1896 Leuconopsis, A.O.Walker
in: Rep. Brit. Ass., 1896 v.66 p.417 | 1897 Z., A.O. Walker in: J. Linn. Soc. London,
v.26 p.227 | 1900 Leucon, G. O. Sars, Crust. Norway, v.3 p.29 | 1907 Z., Calman
in: Tr. zool. Soe. London, v.181 p.31 |-1908 Z., Zimmer in: Ergeb. Tiefsee-Exp., v. 8
p. 176 | 1909 Z., Stappers in: Cellule, v.25 p.369 (alimentary system) | 1911 L., Stappers
in: Camp. arct. Ontac. Crust. Malac. p. 99 | 1912 Z., T. Stebbing in: Ann, S. ian Mus.,
v.l0v p. 156.
Pseudorostral projection prominent, carapace with longitudinal, medio-
dorsal, serrate crest in female, often wanting in male. Peduncle of first
antenna without conspicuous geniculation, accessory flagellum usually very
small. Terminal joint of second antenna in female well defined. Hepato-
pancreatic glands with four pairs of lobes.
20 species.
Synopsis of species:
Uropods with exopod shorter than ataeséa — ll.
Uropods with exopod longer than endopod — 14.
Antenna 1 with accessory flagellum shorter than 1st
joint of principal. «3. 3: . 10. L. mediterraneus
sper 1 with accessory fiaceliunn met ironies. then
1st joint of principal — 12.
10
1 J Branchial siphon monstrously extensile ..... - 1, L. siphonatus
\ Branchial siphon not monstrously extensile — 2.
Pseudorostrum 2 more than half length of rest of
Carapacenm. |i eue-n ° 2. L. tenuirostris
Pseudorostrum not more Hace half length of ree ai
carapace — 3.
3 ‘ Uropods with rami subequal — 4.
Uropods with rami not subequal — 10.
Dorsal crest of 9 carapace serrated throughout its
A | length — 5.
| Dorsal crest of 9 carapace with partial or interrupted
serration — 6.
P { Pseudorostrum truncate ..5....+-+.:+e-s 3. L. nasicoides
Pseudorostrum acute. .....+-.-+.. 4, L. australis
Uropods with 2nd joint of endopod more an half as
6 | HONG pastethen SOF perme sts os a eve 5 . 5. L. fulvus
Uropode with 2nd joint of endopod not more than
half as long as the 1st — 7.
Medio-dorsal serrations reaching end of carapace after
7 ; an interval — 8.
Medio-dorsal serrations not reaching end of carapace — 9.
Antenna 1, accessory flagellum much shorter than 1 st
joint of EINCcipaliye ces) oli rs 6. L. nasica
Antenna 1, accessory flagellum as Giccer as 1st dint
of arate Bee ig ame eam Pie ah 7. L. antarecticus
Uropods with 2nd joint of endopod half as long as Lst 8. L. nathorsti
9 : Uropods with 2nd joint of ee be much less than
LAPIN E Bs, Oy ayes) aa UTR obo eee 9. L. septemdentatus
64 Cumacea: 9. Leuconidae, 1. Leucon
Antenna 1 with accessory flagellum longer than 1st
| joint of principal. - <=. . Il. L. heterostylis
| Antenna 1 with accessory angela Pot once than
lst joint of principal — 183.
12
eit. segment greatly produced between the
18 WLOpOdsts Sesh ke ne ones mea eae oes ode 12. L. longirostris
; | Telsonie segment ails eaieenale produced femeee
THERUTO DOGS Ae-e: oie Sameer enue Se eee 13. L. kalluropus
than its apical spines.) can eaien > a 14. L. pallidus
Uropods with 2nd joint of eadoned feoeder! A and
than its apical spine — 165.
| Uropod with 2nd joint of endopod not broader at end
14 \
Carapace with median serrations continuous throughout 15. L. serratus
Carapace with median serrations not continuous
throughout — 16.
Eyelobe small but distinct. .......... . 16. L. kerguelensis
\ Eyelobe rudimentary — 17.
—
or
ee
J Pseudorostrum horizontal — 18.
17 ) Pseudorostrum bent upwards — 19.
serrations not limited to first half of carapace . 17. L. assimilis
Frontal lobe without lateral denticles; median ser-
rations only on first half of carapace. . . .. . 18. L. acutirostris
18
\
, Frontal lobe with 3 denticles on each side; median
nal Uropods with exopod only a little longer than the
peduncle... = = 19. L. sagitta
ee) Uropods with exopod Bean ioaver than ‘ie paderele 20. L. vanhoeffeni
1. L. siphonatus Calm. 1905 Z. s., Calman in: Sci. Invest. Fish. Ireland,
1904 app. 1 p.19 t.1 f.2—4 | 1906 Z.s., Calman in: Mt. Stat. Neapel, v.17 p.416 t. 27
f.9 | 1908 L. s., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 177.
Pseudorostral lobes well advanced, slightly upturned, apically rounded,
with teeth on the dorsal and apical margins in the Q only, the border between
Fig. 33.
L. siphonatus Calman (2 juv.).
the apex and the antero-lateral tooth having 2 emarginations separated by
a serrate prominence and introduced by a strong subapical tooth; the lower
border behind the antero-lateral tooth serrate. In the © the frontal lobe has
one or two teeth at its apex. Anteriorly the sides of the carapace are
sprinkled with denticles. Antenna 1, 34 joint longer than 2"4, flagellum 3-jointed,
the 1-jointed accessory not wart- like but nearly equal in length to 1S* joint
of principal flagellum. 1S* maxillipeds, with the branchial siphon formed by
the exopods capable of protrusion to a distance greater than length of carapace,
Cumacea: 9. Leuconidae, 1. Leucon 65
surrounded by stiff setae springing from the pseudorostral margin. Perae-
opod 1 rather robust, 5 and 6% joints equal, 7 about one-third as long
as either. Peduncle of uropods shorter than the subequal rami, endopod with
1st joint nearly three times as long as 2"4, with 5 spines on medial margin,
2nd joint with 3 spines and at apex a spine and a stout plumose seta.
L. © juv. (Ireland) 3°85 mm, © with marsupium (Mediterranean) 2°7 mm.
W. coast of Ireland; depth 699 m; Mediterranean, near Capri; depth 100—1100 m.
2. L. tenuirostris O.Sars 1887 L.¢., G.O.Sars in: Rep. Voy. Challenger,
v.19 pars 55 p.33, 38 t.5 f.1—4 ! 1905 Z.t., Calman in: Sci. Invest. Fish. Ireland,
1904 app. 1 p.20| 1908 L.¢., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p.177, 178.
General form very slender. Pseudorostral lobes slightly upturned,
tapering to subacute apices for a length exceeding half that of the rest of
the carapace; antero-lateral corners little produced but acute, preceded by a
small sinus between denticles. Carapace compressed, with median carina
having 7 teeth along the front half, and front half of lower margin den-
tate. Fifth pedigerous segment not wider than the pleon, in which the
5th segment is very long, the telsonic segment short, almost pentagonal,
forming an obtuse triangle between the bases of the uropods. Antenna 1,
accessory flagellum not wart-like, its one joint as long as first of principal
flagellum. Peduncle of uropod as long as the endopod, in which the somewhat
tapering first joint has 5 spines on the medial margin, the slender 2™4 joint
a little more than half as long has 4 spines, the apical the largest; the
exopod only reaches half way along the 24 joint of the endopod. L. Q juv.
about 10 mm.
N. Pacific, lat. 836° 10’ N., long. 178° E.; depth 3750 m.
3. L. nasicoides Lilj. 1855 L. n., W. Liljeborg in: Ofv. Ak. Forh., v. 12 p. 122 |
1869 L. n., G.O.Sars in: Nyt Mag. Naturv., v.16 p. 345 | 1872 ZL. n., G.O.Sars in: Forh.
Selsk. Christian., 1871 p.270 | 1879 Z.n., G.O.Sars in: Arch. Naturv. Kristian., v.4 p.15, 22 |
1879 L.n., A.M. Norman in: Ann. nat. Hist., ser. 5 v.3 p.72 | 1879 L.nasicoidis, S.1. Smith
in: Tr. Connect, Ac., v.5 p.115, L. nasicoides, p.125, 129 | 1886 L. nasicoides, G.O. Sars in:
Norske Nordhavs-Exp., v.6 nr.15 p.19 | 1887 L.n., H. J. Hansen in: Vid. Meddel., v. 39
p. 199 | 1900 L. n., G. O. Sars, Crust. Norway, v.3 p.31 t.23 | 1900 ZL. n. (part.), Zimmer in:
Fauna arctica, v.1 p.411, 416 | 1901 L..n., Ohlin in: Bih. Svenska Ak., v.26 nr. 12 p. 41 |
1908 L.n., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p.177, 178 | 1911 ZL. n., Stappers in:
Camp. arct. Orléans, Crust. Malac. p.102 t.4 f.5,7 | 1912 L.n., Calman in: P. U.S.
Mus., v.41 p. 619.
©. Pseudorostral lobes moderately produced horizontally to narrowly
truncate apex; lateral truncation denticulate ; antero-lateral corners triangular,
the lower margin behind them serrate for half its length. Median carina of
carapace serrate from front to hind margin, frontal lobe having also 3 den-
ticles on each side. Telsonic segment short, pentagonal, triangularly pro-
duced between bases of uropods. Antenna 1, 3¢ joint shorter than 24
Peraeopod 1, 6 joint much shorter than 5t*. Peraeopod 2, 3¢ joint wanting.
Peduncle of uropods a little shorter than the subequal rami; first joint of endo-
pod more than twice as long as 2™4, carrying 9 spines on the medial margin,
where the 2"4 has 5 followed by along one at the apex. Colour yellowish
white. L. 6 mm.
SG. More slender than ©. Pseudorostral lobes transversely truncate,
with 2 denticles; antero-lateral corners obtuse-angled. Carapace smooth
Das Tierreich. 39. Lief.: T. R. R. Stebbing, Cumacea. 5
66 Cumacea: 9. Leuconidae, 1. Leucon
except for 2 denticles in line at apex of frontal lobe. Peraeopod 3 with 2
subequal narrow lanceolate appendages on the 3¢ joint. Peduncle of uropods
scarcely shorter than the rami, in which the endopod has a few more spines:
than in the © and its 2"¢ joint rather more than half as long as the first.
L. 6 mm.
Coast of Norway, depth 19—94 m; Kattegat; Bay of Fundy; Gulf of St. Lawrence;
Greenland, depth 75 m; Spitzbergen, depth 1000 m; lat. 45° 29’ N., long. 55° 24’ W.,
depth 103 m; Nova Zembla, 90 m.
4. L. australis Calm. 1907 L.a., Calman in: Nat. antarct. Exp., v.2 Crust.
Cum. p.1 f. 1—3.
Pseudorostral lobes very slightly upturned, acute, not denticulate:
antero-lateral corners triangular, lower margin behind them serrate for a short
distance. Median carina of carapace closely serrate from front to hind
margin. Telsonic segment short, pentagonal, obtusely triangular between
bases of uropods. Antenna 1, accessory flagellum longer than first joint of
principal. Peduncle of uropods slightly shorter than the subequal rami,
endopod with 18* joint twice as long as 2™4, carrying 8 spines on medial
margin, 2™4 joint having 3 on medial margin and a larger one on apex.
L. Q 3°5 mm.
Antarctic.
5. L.fulvus O. Sars 1865 L.f., G.O.Sars in: Forh. Selsk. Christian., 1864
p. 180 | 1886 L. f., G.O. Sars in: Norske Nordhavs-Exp., v.6 nr. 15 p.19 | 1900 L. 7.
G, O. Sars in: Crust. Norway, v.3 p.32 t.24 | 1900 Z.f, Zimmer in: Fauna arctica,
v.1 p.416 | 1908 ZL. 7f., Zimmer in: Ergeb. Tiefsee-Exp., v8 p.177,178 | 1911 L. f,
Stappers in: Camp. arct. Orléans, Crust. Malac. p.105 t.4 f. 8.
©. Pseudorostral lobes slightly upturned to apex in dorsal view
narrowly truncate, laterally denticulately truncate; antero-lateral corners acutely
produced with slight serration adjoining. Median carina of carapace denti-
culate from front to behind the middle and after an interval reappearing
to the rear. Peraeopod 1, 6 joint nearly as long as 5, Peduncle of
uropods subequal to the subequal rami, 2"¢ joint of endopod two-thirds as.
long as 1**, 1st with 6 spines on medial margin, 2"¢ with 4 and a very long
apical spine. Colour bright orange. L. scarcely 5 mm.
GS. Pseudorostral lobes obtusely rounded at apex with its lateral
border smooth; antero-lateral corners smoothly rounded, preceded and followed
by serration. Carapace quite smooth, with minute denticle at apex of frontal
lobe. Peraeopod 3, 34 joint with 2 comparatively short, falciformly curved
appendages (Sars), 4 lanceolate appendages, not very long, slightly incurved,
3 of them subequal (Stappers). Uropods more elongate than in ©, spines 10
and 6 on joints of endopod, 24 joint nearly four-fifths as long as 1%*.
Colour paler than in O. IL. 5°5 mm.
Lofoten Islands; depth 11—23 m; Finmark; Iceland; Spitzbergen; Nova.
Zembla, 90 m.
6. L. nasiea (Kroyer) 1841 Cuma n., Kroyer in: Naturh. Tidsskr., v.3 p. 524,
532 t.6 f.31—33 | 1846 Leucon n., Kroyer in: Naturh. Tidsskr., ser. 2 v.21 p. 189, 209
t.2 f.5a,b | 1849 DL. n., Kréyerj(Petersen del.) in: Voy. Nord, Crust. t.3 f.2a—o |
1855 L.n., W. Liljeborg in: Ofy. Ak. Forh., v.12 p.121 | 1865 ZL. nasicus, G. O. Sars
in: Forh. Selsk, Christian., 1864 p.178 | 1875 L.nasica, Aug. Metzger in: Jahresber.
Comm. D. Meere, v, 2/3 p. 286, 308 | 1877 L.n., Meinert in: Naturh. Tidsskr., ser. 3 v. 1k
Cumacea: 9. Leuconidae, 1. Leucon 67
p. 180 | 1879 L. nasicus, A. M. Norman in: Ann. nat. Hist., ser.5 0.3 p. 70 | 1879 L. n.,
S. I. Smith in: Tr. Connect. Ac., v.5 p.114 | 1884 L. n., J.S. Schneider in: Tromse Mus.
Aarsh., v.7 p.54 | 1887 L.n., H. J. Hansen in: Dijmphna Udb., p. 240; and in: Vid.
Meddel., v.39 p.199 | 1893 L. nasica, T. Stebbing in: Int. sci. Ser., v.74 p. 304 | 1893
Cuma n., Apstein in: Jahresber. Comm. D. Meere, v.6 p. 196 | 1897 Leuconopsis ensifer,
A.O. Walker in: J. Linn. Soe. London, v.26 p.227 t. 17 f. 1—1Lh(d) | 1900 Leucon nasicus,
G. O. Sars, Crust. Norway, v.3 p.30 t. 21,22; p.106 L. nacica (misprint for nasica) |
1900 L. nasicus, Zimmer in: Fauna arctica, v. 1 p.417 | 1901 L.x., Ohlin in: Bih. Svenska
Ak., 0.26 nr.12 p.40 | 1902 L. n., L. nasicoides (err. pro: L. nasica var.), Gas:
Brady in: Nat. Hist. Tr. Northumb., v.14 p.94 | 1908 ZL. nasica, Zimmer in: Ergeb.
Tiefsee-Exp., v.8 p.177 | 1909 L. nasicus, A. M. Norman & G. S.Brady in: Nat. Hist. Tr.
Northumb., ser.2 v.311 p.27 | 1911 ZL. nasica, Stappers in: Camp. arct. Orléans, Crust.
Malac. p.100, L. nasicus t.4 f.1,6| 1912 LZ. n., Calman in: P. U.S. Mus., v.41 p.619.
Fig. 34. Fig. 35.
L. nasica (Q) (Kroyer), peraeopod 3 L. nasica (Kroyer), maxilla 1 (after
(after Sars). Sars).
Body slender and elongate. Pseudorostral lobes in Q upturned, pro-
minent, obliquely truncate with the truncate edges minutely denticulate and
setiferous; antero-lateral corners sharply triangular preceded by a deep sinus,
followed for some distance by fine serration of the lower margin. In the )
the lobes are shorter, little upturned, almost transversely truncate, and the
antero-lateral corners are obtuse with the sinus obsolete. Carapace in © with
median carina serrate beyond the middle and after an interval again serrate
to the hind margin, but this crest is quite wanting in the adult GO. An-
tenna 1, 34 joint as long as 24, accessory flagellum minute, though 2-jointed
in the G. Antenna 2 of Q with 3 plumose setae on 1st joint. Peraeopod 1
rather long, 6" joint not much shorter than 5. Peraeopod 3 in SO with
2 very conspicuous unequal ensiform appendages on the 34 joint. Pedunele
of uropods equal to the equal rami, 2"¢ joint of endopod less than half as
long as the 1s‘, which has on the medial margin numerous spines long and
short interspersed, the 2"¢ joint having a graduated series of spinules followed
by a long apical spine and still longer plumose seta; the exopod carries numerous
5*
68 Cumacea: 9. Leuconidae, 1. Leucon
plumose setae. Colour whitish, L. 9 12 mm, d 10mm. The young d
is much like the ©.
Norway, depth 56—188 m; Kattegatt ; Heligoland; British Isles, depth 82—108 m;
Gulf of St. Lawrence, depths 91 and 128m; Spitzbergen, 175—210 m; Kara Sea;
Greenland, 4—565 m; Afognak Bay, Alaska, 22—31 m; Coal Harbor, Unga, 11—l4m;
Mare Murmanicum, 90 m.
7. L. antareticus Zimmer 1907 ZL. antarctica, Zimmer in: Zool. Anz., v.31 p.227.
© juv. Slender. Pseudorostral lobes much extended to apex in dorsal
view very acute; antero-lateral corners preceded by a broad deep sinus.
Carapace with denticulate carina from front to hind margin, but last 2 denticles
separated by a smooth space from the rest. An eyelobe obscurely present,
the frontal lobe having a pair of lateral denticles in front and a sublateral
pair to the rear, the pseudorostral lobes also having a pair close to the front
pair of the frontal lobe. Antenna 1, the 1-jointed accessory flagellum as long as
1st joint of principal flagellum. Peduncle of uropods shorter than the rami,
exopod slightly shorter than endopod, which has on
medial margin of 18* joint about 9 spines, 3 on 2"4
joint and an apical spine. Colour yellowish white.
L. about 13°3 mm. G juv. differing little from © juv.
German South-Pole Expedition, Winter Station (lat.
66° 2‘ S., long. 89° 38‘ E.)
Fig. 36. Fig. 37.
L. nasica (Kroyer), maxilliped1 (after Sars). L. nathorsti Ohlin.
8. L. nathorsti Ohlin 1900 Z. nasicoides (part.), Zimmer in: Fauna arctica,
v.1 p.417 | 1901 ZL. nathorsti, Ohlin in: Bih. Svenska Ak., v.26 nr. 12 p.41 .6 f.9a—e |
1908 L. n., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p.178 | 1911 Z. n., Stappers in: Camp.
arct. Orléans, Crust Malac. p. 104.
Near to LZ. nasicoides (nr. 3), but pseudorostral lobes longer and more
acuminate, with six teeth to the lateral truncation; the antero-lateral corners
moderately produced, lower margin behind them serrate for a third of its
length. Median carina of carapace with 13—15 teeth reaching a little
Cumacea: 9, Leuconidae, 1. Leucon 69
behind the middle, and usually 2 or 3 more within the remaining space;
frontal lobe usually with only 1 denticle or none (instead of 3) on each
side. Peduncle of uropods about equal to endopod, a little longer than
exopod; 2™4 joint of endopod half as long as 1**. Adult 3 without median
carina. L. O 7 mm.
King Charles Island, lat. 78° 50‘ N., depths 20—110 m. Leucon breviremis,
H. J. Hansen MS., not adopted by Ohlin, as being appropriate only to young specimens.
9. L. septemdentatus Zimmer 1902 L. s., Zimmer in: Ergeb. Hamburg.
Magalh.-Sammelr., v. 2 Cumac. p.3 f.1—5 | 1908 LZ. s., Zimmer in: Ergeb. Tiefsee-Exp.,
v.8 p. 177,178 | 1909 L. s., Zimmer in: Ergeb. Schwed. Siidp.-Exp., v.6 nr, BOO tod
f. 6—16.
©. Pseudorostral lobes slightly upturned; antero-lateral corners shortly
produced, acute. Carapace rather short, median carina variable in length
but not reaching beyond the middle, its teeth 5—9 in variable groups.
Pleon segment 5 not much longer than 4*, telsonic segment short, little
produced between uropods. Antenna 1, single joint of accessory flagellum
equal to 18t of primary. Peraeopod 1, 5‘ joint longer than 6%, 6' than
7th, Peraeopod 2 with full number of joints. Peraeopod 4 with 1-jointed
exopod on right or left limb’s 24 joint, whether sometimes on both uncertain.
Peduncle of uropods longer than exopod, as long as endopod, of which the
1st joint is more than thrice as long as the 2™¢ and carries 12 spines on
the medial margin, the 2™4 having 4, with 2 apical, the latter very long.
Colour yellowish white. L. about 5 mm.
3. More slender than @. Pseudorostral lobes shorter and more blunt;
antero-lateral corners obtuse, not produced. Carapace without denticulate
carina. Flagellum of antenna 2 reaching beyond the pleon. Peraeopod 3
without the peculiar appendages on the 3¢ joint sometimes found in S of this
genus. Peduncle of uropods rather shorter than the subequal rami, its medial
margin and proximally that of endopod having plumose setae intermixed
with the spines. L. about 4 mm.
Fuegian archipelago, depth 6 m; Falkland Isles, depths 1—8 m.
10. L. mediterraneus O. Sars 1879 L. m., G. O. Sars in: Arch. Natury.
Kristian., v.4 p.25 t.27—29 | 1908 L. m., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 177.
Integument thin and flexible, squamose. Pseudorostral lobes strongly
upturned, triangular in 9, shorter, sub-horizontal in dG; antero-lateral
corner in © forming a strong tooth preceded by a smaller one, a sinus,
and 2 denticles, and followed by about 10 denticles on lower margin’s first
half; in the G the corners are rounded with 2 denticles at some distance
from the feebler serration of the lower margin. Median carina of carapace
wanting in G, in © low, furnished with 6—8 spaced teeth, the series ending
just behind the middle. Pedigerous segments 2—4 with somewhat outstanding
side-plates. Pleon segment 5 dorsally setose behind. Telsonic segment short,
bluntly conical between bases of uropods. Antenna 1, 34 joint longer than
1st or 2™4, flagellum 3-jointed in Q, 4-jointed in GO, accessory almost imper-
ceptible. Antenna 2, © 3-jointed, with 2 plumose setae on 1S joint, 34 joint
long and narrow: in GC flagellum reaching nearly length of the animal, the
joints successively longer. Maxilliped 1 with 5 small branchial sacs. Perae-
opod 1, 284 joint unusually narrow and nearly straight. Peraeopod 2, with
34 joint lost or fused. Peduncle of uropods in Q subequal to endopod,
which has 9 spines on medial margin of 1%* joint and 5 on that of the
70 Cumacea: 9. Leuconidae, 1. Leucon
second, the latter two-fifths as long as the former and carrying an apical
spine; the exopod is a little longer than the 18* joint of the endopod. In
the G the endopod is rather longer than the peduncle, its 18* joint equal to
the exopod and carrying more numerous spines. Colour whitish, semi-
pellucida. L. © 6mm, G a little less.
Mediterranean, Spezia, depth 38—56 m.
11. L. heterostylis Calm. 1907 L, (?)h., Calman in: Tr. zool. Soc. London,
v.181 p.32 t.8 f. 1—5.
Pseudorostral lobes straight, horizontal, acute, with coarsely quadridentate
projection preceding the sinus which leads to the antero-lateral tooth; serration
of lower margin behind the tooth begins strongly but soon dies out. Median
line of carapace coarsely and irregularly dentate in anterior half, with one
little tooth near hind margin following a depression. Antenna 1, 15* joint
longer than 2"4, 2™¢ than 34, principal flagellum of 3 joints, the last minute,
accessory l-jointed, nearly as long as first 2 joints of principal combined.
Antenna 2 with 3 joints, 3 plumose setae on the 18%. Maxilliped 1
with 2 papilliform branchial sacs. Peraeopod 2, 34 joint represented by
2 portions of a narrow ring. Peduncle of uropods a little longer than exopod
but a little shorter than 18* joint of endopod, which is two and a half times
as long as the 24 joint, these joints carrying 8 and 4 spines respectively,
the apical spine very long. L. © 3°42 mm, ¢ unknown.
New Zealand, Akaroa Harbour, depth 11 m.
12. L. longirostris O. Sars 1871 LZ. 1., G. O. Sars in: Ofv. Ak. Forh., v.28
nr. 1 p.78 | 1871 ZL. U., G. O. Sars in: Svenska Ak. Handl., n. ser. v.9 nr. 13 p.42 t.15
f.75 | 1879 L.1., A. M. Norman in: Aon. nat. Hist., ser. 5 v3 p.69 | 1906 Z.1., Calman
in: Mt. Stat. Neapel, v.17 p.414 t.27 f.1—8 | 1908 Z.J., Zimmer in: Ergeb, Tiefsee-
Exp., v.8 p.177 | 1912 ZL. 1, Calman in: P. U.S. Mus., v.41 p.619.
©. Pseudorostral lobes more than one-third of total length of carapace,
straight, somewhat upturned, acute, with 2 or 3 teeth below leading to a
frontal row of 3 or 4, which are separated by a narrow sinus from the
strong tooth of the antero-lateral corner; adjoining lower margin serrate for
some distance. Median carina of carapace carrying about 7 forward-pointing
teeth, diminishing in size towards the hind margin, which they do not nearly
reach. Side-plate of 4% pedigerous segment with a little curved tooth at
the hind angle, the 5% segment having a pair of forward-curving ventral
teeth. Pleon segment 5 long, telsonic segment short, with dorsal depression.
Antenna 1, flagellum 3-jointed, its 18 joint as long as 34 joint of peduncle
and as the 1-jointed accessory. Peraeopod 1 long, with row of teeth
of 294 joint on lower surface. Pedunele of uropods a little shorter than the
exopod, which is a little longer than the 18* joint of the endopod; the latter
more than thrice as long as the 2"4 joint, with 15 spines on the medial
margin, 2" joint with 5 on that margin and a much larger apical one.
L. O subadult 5:7 mm.
3. Pseudorostral lobes one-fifth of total length of carapace, level,
obliquely truncate with 2 teeth at end of truncation, the margin then retreating
to a vertical piece with 2 teeth and a small antero-lateral tooth below.
Median carina of carapace in adult without teeth, with 5 in young d (Sars).
Side-plate of 4t pedigerous segment without tooth, 5% segment with the
ventral pair. Telsonic segment more narrowly produced than in the 9
Antenna 1, flagellum 4-jointed, its 18* joint in the adult equalled by the
Cumacea: 9. Leuconidae, 1. Leucon alt
1-jointed accessory, which in the young male is rudimentary, wart-like (Sars),
but not so (Calman). Peraeopod 3 without peculiar appendages to the 30
joint, such as are found in some species. Peduncle of uropods rather shorter
than exopod which is a little shorter than 1st joint of endopod, this being
4 times as long as the 2™4 joint, both joints having more spines than in the
©, most of them small and simple, at least as figured. L. 6 mm.
Off Portugal, lat. 38¢ 7‘ N., depth 1036 m; Davis Strait, lat. 59° 10° N., depth
3200 m; off Capri, depths 950—1200 m; lat. 40° 16‘ 50 N., long Gro Daelb Ws
depth 2322 m.
13. L. kalluropus Stebb. 1912 L.k., T. Stebbing in: Ann. S. Afr. Mus., v. 10v
p.156 t.57 (9).
Seemingly near to L. Jlongirostris
(nr. 12), but with the telsonic segment less
produced, the produced portion much instead
of little shorter than the base, the proportions
and armature of the uropods also differing.
Pedigerous segment 5 with procurved ventral
spines as in the other species, and the 1-
jointed accessory flagellum of antenna 1
as long as 18* joint of the 4 in the principal.
Peraeopods 1—4 with 2"4¢ joint distally
narrowed. In peraeopods 2—5 the 3¢ joint
is obscure. In the 24 pair the much spined
7th joint is as long as the 5 and fully
twice the 6, Peduncle of the uropods a
little longer than the 18* joint of the endo-
pod, which is more than thrice as long as
the 24 joint, its medial margin fringed with
more than 20 serrate spines following 2 or
3 that are setiform, the lateral margin
carrying 8 feathered spines; the 2™¢ joint
has 8 serrate spinules on the medial margin;
the exopod is a little shorter than 1S* joint
of endopod, and has various elongate spines.
L. do about 5 mm, © unknown.
I 526 = ; Fig. 38.
S. Africa, Cape Natal N. by E. 24 miles, L. kalluropus (3) Stebbing, telsonic seg-
depth 805 m.
ment and uropod.
/
14. L. pallidus 0. Sars 1865 ZL. p., G.O.Sars in: Forh. Selsk. Christian.,
1864 p. 182 | 1873 L. p., G. O. Sars in: Svenska Ak. Handl., n. ser. v.11 nr.6 p.8 t.3
f.10 | 1879 L. p., A.M. Norman in: Ann. nat. Hist., ser.5 v3 p.72 | 1886 L. p., G. O.
Sars in: Norske Nordhavs-Exp., v.6 nr.15 p.19 | 1894 LD. p., A. M. Norman in: Ann.
nat. Hist., ser.6 v.13 p.275 | 1900 L. p., G. O. Sars, Crust. Norway, v.3 p.33 t. 25 |
1900 L. p., Zimmer in: Fauna arctica, v.1 p.417, 432, 437 | 1905 L. p.(?), Calman in:
Sci. Invest. Fish. Ireland. 1904 app. 1 p.19 | 1908 L.p., Zimmer in: Ergeb. Tiefsee-Exp.,
v.8 p.178 | 1911 L.p., Stappers in: Camp. arct. Orléans, Crust. Malac. p.106 t.4 1
Slender, much compressed. Pseudorostral lobes in © long, horizontal,
tapering to obtusely acuminate apex, antero-lateral corners acute, preceded
by a deep sinus, followed by serration of lower margin for some distance;
in G lobes much shorter, with lower margin serrate and antero-lateral corners
rounded off. Carapace with dorsal carina denticulate in both sexes, with
72 Cumacea: 9. Leuconidae, 1. Leucon
scarcely an interruption from end to end, and in immature specimens (Calman)
having 1 or 2 small denticles on each side of the frontal lobe. Telsonic
segment very short, produced and rounded triangular. Antenna 1, accessory
flagellum rudimentary. Peraeopod 2 with 34 joint wanting or fused. Peraeopod
3 with three large lanceolate distally denticulate appendages on the 34 joint
in G. Peduncle of uropods shorter than the rami or about equal to the
endopod exclusive of its spine-like terminal, which in immature specimens
is distinct from the trunk (Calman), 2™4 joint apart from terminal less than
half as long as 1S*; exopod longer than endopod with the terminal included.
Colour pure white. L. oO 4mm, ¢ 4:5 mm.
Coast of Norway, depth 75—753 m; Spitzbergen, depth 2636 m; Kara Sea; Nova
Zembla, 90 m.
15. L. serratus Norm. 1876 ZL. s. (nom. nud.), A. M. Norman in: Jeffreys in:
P. R. Soe. London, v.25 p.212 | 1879 ZL. s., A. M. Norman in: Ann. nat. Hist., ser. 5
v.3 p.70 | 1900 ZL. s., Zimmer in: Fauna arctica, v.1 p.418, 432 | 1908 L. s., Zimmer
in: Ergeb. Tiefsee-Exp., v.8 p. 178.
Pseudorostral lobes short, horizontal, triangular, with serrate edge; antero-
lateral corners produced forward and outward in wing-like form, the serrations
of lower margin forming about 10 teeth, the first 3 or 4 large. Median carina
of carapace boldly arched, serrate from end to end, teeth decreasing to the
rear. Peraeopod 1, 24 joint with 7 spines on hind margin. Peduncle of
uropods not longer than 5 pleon segment, endopod much shorter than
exopod, each of its joints with 5 spines on the medial margin, apical spine
two-thirds as long as the 2™¢ joint. L. © 6 mm, Go unknown.
Davis Strait, lat. 50° 10’ N., long. 50° 25‘ W., depth 3200 m.
16. L. kerguelensis Zimmer 1908 L.k., Zimmer in: Ergeb. Tiefsee-Exp., v.8
p. 178 t.41 (6) £.71, 72, t.42 (7) £.73—75.
Body slender. Pseudorostral lobes short, subacute, slightly upturned.
Antero-lateral corners rounded, smooth. Carapace with a small but actual
eyelobe on which a smooth median carina takes its origin, ceasing a little
way to the rear of the frontal lobe. Antenna 1, flagellum 4-jointed, the
1-jointed accessory about half as long as 1S* joint of principal flagellum.
Antenna 2 about reaching end of the body. Peraeopod 1 elongate, 2"¢ joint
in this and the 3 following pairs strongly developed. Peraeopod 3 having
the 34 joint equipped with 2 lanceolate appendages. Peduncle of uropods
closely fringed on medial margin with spines and plumose setae; the endopod
about equal to it in length, with 2™4 joint half as long as the first, is little
more than three-fourths as long as the exopod. Colour white. L. ¢ a little
over 5mm. © unknown.
Kerguelen, Gazelle Harbour.
17. L. assimilis O. Sars 1887 ZL. a., G. O. Sars in: Rep. Voy. Challenger,
v.19 pars 55 p. 33, 34 t.4 | 1908 ZL. a., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 177, 178.
Near to L. nasica (nr. 6). Integument thin, pellucid, with no distinct
sculpture. Pseudorostral lobes not elongate; horizontal, subacute, oblique
margins denticulate and setulose, sinus denticulate; antero-lateral corners
triangular, lower margin denticulate for most of anterior half. Median carina
of carapace arcuate in front, densely denticulate and setulose, the 3 sub-
terminal denticles a little separated from the rest; eyelobe wanting, frontal
Cumacea: 9. Leuconidae, 1. Leucon
=~!
Oo
lobe carrying a series of 3 denticles close to each lateral fissure. Pedigerous
segments 1—5 with laminar sideplates slightly projecting laterally. Telsonic
segment pentagonal, produced end apically rounded. Antenna 1, 2"? joint a
little shorter than 15*, and 34 than 2"¢, flagellum 3-jointed, 18* joint twice
as long as 2™¢ and 34 combined, accessory wart-like. Maxilliped 1, with 5
branchial sacs, the membranous exopod elongate. Peraeopod 1, 5 and 6%
joints subequal, 7** much shorter than either, apically carrying some half
dozen strong, claw-like spines. Peraeopod 2 with 3¢ joint wanting or fused.
Peraeopod 3 in young @ apparently without any peculiar appendages on 34
joint, but seemingly in both sexes the 5 joint carries 3 strong setae, the
distal part of which is densely annulated and apically uncinate. Peraeopod 5
alike in both sexes, and very small. Peduncle of uropods as long as the
endopod, in which the 1%* joint is a little more than twice the second, fringed
on medial margin with a dozen spines, 3 conspicuously larger than the rest;
the 2"¢ joint has 5 medial marginal and 2 larger apical spines; the exopod
is about a fifth of its length longer than the endopod. Colour light yel-
lowish. L. © and young CG, 9 mm.
Kerguelen, Cumberland Bay, depth 232 m.
18. L. acutirostris O. Sars 1865 L.a., G.O. Sars in: Forh. Selsk. Christian.,
1864 p.181 | 1894 Z.a., A.M. Norman in: Ann. nat. Hist., ser.6 v.13 p.275 | 1900
L. a., G. O. Sars, Crust. Norway, v.3 p.34 t.26 | 1900 ZL. a., Zimmer in: Fauna arctica,
v.1 p. 432, 438, 439, 444 | 1908 L.a., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p.178 | 1911
L. a., Stappers in: Camp. arct. Orléans, Crust. Malac. p.107 t.4 f. 10.
Body of © rather short and thick, of Od more slender. Pseudorostral
lobes rather short, especially in G, horizontal, acuminate, margins adjacent
to apex denticulate, setiferous; antero-lateral corners acutely produced in ©
with serration following a narrow sinus, blunt and smooth in G; lower margin
serrate for some distance in both sexes. Median carina on anterior half of
carapace in ©, strongly denticulate, but entirely wanting in C. Telsonic
segment pentagonal, produced part apically rounded. Peraeopod 1, 5‘ and
6th joints equal, 72 much shorter, apically setose. Peraeopod 2 without 3¢
joint. Peraeopod 3 of dD having 3 lanceolate not distally denticulate append-
ages on the 34 joint. Peduncle of uropods equal to endopod, in which the
gnd joint is two-thirds of the 18* in Q, a little over half in the ¢, having
4 spines on medial margin of each joint in O, but in GO 12 on 15 and
6 on 224, with a long apical spine in both sexes; exopod somewhat longer
than endopod. Colour whitish with olive tinge. L. OQ 3mm, C 35 mm.
Coast of Norway, depths 56—548 m; Nova Zembla, 90 m.
19. L. sagitta Zimmer 1907 LZ. s., Zimmer in: Zool. Anz., v.31 p.225 | 1909
L. s., Zimmer in: Ergeb. Schwed. Siidpol.-Exp., v6 nr.3 p.7 t.2 f.17—81.
Body slenderly lanceolate. Pseudorostral lobes moderately long, slightly
upturned, apically acute, the oblique margins feebly denticulate; antero-lateral
comers triangular, under margin of carapace denticulate for a space. Median
carina of carapace reaching to middle or further, with teeth varying from 5
to 12, with sometimes a single tooth near the hind margin. Pleon segment
5 rather short. Telsonic segment with produced part almost truncate but
not sharply. Eyelobe reduced to a little point. Antenna 1, accessory flagellum
about three-fourths as long as 1%t joint of the principal. Peraeopod 1, 7%
joint nearly as long as 6%, 6% much shorter than 5. Peraeopod 2 with
37 joint lost or fused, and this joint not very clearly separated from the 24
74 Cumacea: 9. Leuconidae, 1. Leucon, 2. Kudorella
in the next 2 pairs. Peduncle of uropods intermediate in length between
the shorter endopod and longer exopod; endopod about two-thirds of exopod,
with 18t joint nearly twice as long as 2™4, carrying 6 or 7 spines on the
medial margin, the last much the largest; 24 joint with 4 spines on medial
margin and a large terminal. Colour yellowish white. L. @ about 5 mm.
Adult G not known.
South Georgia, depths 12—310 m.
20. L. vanhoeffeni Zimmer 1907 ZL. v., Zimmer in: Zool. Anz., v.31 p.226.
Body rather narrow. Pseudorostral lobes short, bent very obliquely
upwards, only subacute; antero-lateral corners preceded by a broad sinus.
Median carina of carapace equipped with 5—7 teeth and extending about to
the middle. Eyelobe rudimentary. Antenna 1, 1-jointed accessory about
three-fourths as long as 1s¢ joint of principal flagellum. Peraeopod 1, 6%
and 7* joints equal, 5% longer than either. Peduncle of uropods about
three-fourths as long as the exopod and about as long as the endopod, which
has about 8 spines on medial margin of 18* joint and 6 on that of the a™4
and a strong apical spine. L., O with commencing marsupium, about 5 mm.
Adult G not known.
Kerguelen, on Macrocystis.
L. leuckartii Marcusen 1867 ZL. 1., Marcusen in: Arch. Naturg., v. 331 p. 360
(descr. nulla),
Black Sea.
2. Gen. Eudorella A. M. Norman
1846 Leucon (part.), Kroyer in: Naturh. Tidsskr., ser.2 7.211 p. 208 | 1856 Eudora
(non Hudora, Péron & Lesueur, 1809 Hydrozoa, nee Haan 1833, Crust.), Bate in: Ann.
nat. Hist., ser.2 v.17 p.457 | 1863 Cyrianassa (part.), A. M. Norman in: Tr. Tyneside
Club, v.51v p.273 | 1865 Eudora, G. O. Sars in: Forh. Selsk. Christian., 1864 p. 184 |
1867 Eudorella, A. M. Norman in: Rep. Brit. Ass., v.36 p.194, 197 (nota) | 1869 Eudora
G. O. Sars in: Nyt Mag. Naturv., v.16 p. 346 | 1871 Eudorella (part.), G. O. Sars in:
Svenska Ak. Handl., n. ser. v.9 nr. 13 p. 43 | 1877 E. (part.), Meinert in: Naturh. Tidsskr.,
ser.3 v.11 p.181 | 1879 E., G. O. Sars in: Arch. Naturv. Kristian., 7.4 p.33 | 1879 E.
(part.), S. I. Smith in: Tr. Connect. Ac., v.51 p.115 | 1883 E., G. O. Sars in: Forh.
Selsk. Christian., 1882 nr. 18 p.12 | 1887 E., G. O. Sars in: Rep. Voy. Challenger, v. 19
pars 55 p.7,40 | 1900 H., G. O. Sars, Crust. Norway, v.3 p.35 | 1900 E., Zimmer in:
Fauna arctica, v.1 p. 418, 433 | 1907 H., Calman in: Tr. zool. Soe. London, v.181 p.31 |
1908 E., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 176, 179 | 1909 E., Stappers in: Cellule,
v.25 p.369 (alimentary system).
Body slender. Carapace short, subtruncate. Eyelobe obsolete. Efferent
opening for water circulation dorsal. First antennae geniculate between gnd
and 34 joints. Uropods with exopod shorter than the endopod. Hepato-
pancreatic glands with four pairs of lobes.
13 species.
Synopsis of species:
(ee above antero-lateral corner of carapace simply
concave — 2.
| Sinus above antero-lateral corner of carapace not simply
concave — 4.
| Uropods having apical spine of endopod fused with the 2nd
oy
JONG oye Mo “Gesietsbers che ey coer as <6 3 B82 6 OL. omarginata
| Uropods having apical spine of endopod distinct — 3.
Cumacea: 9, Leuconidae, 2. Eudorella iD
Uropods with 1st joint of endopod more than thrice as
| long as 2nd... . : 2. BE. gracilior
z | Uropods with 1st joint bi enionod ‘little over ice as
[ova eRe is pL ine ie ot (ohare ciec oa eve ee os due taller
Fifth pleon segment with postero-dorsal group of long
| setae — 5.
. | Fifth pleon segment without postero-dorsal group of long
setae — 6.
Uropods with 2n4 joint of Syeee one-third of 1st, apical
| spine fused .. . 4. Bi. nana
2 | Uropod with 2nd joint a ndenod one- Fourth: 68 Ist, apical
spine distinct -...-..-- SP target en eee cas MOnOdor
Anterior margin of pseudorostral lobes beset with forward
pointing setae — 7.
Anterior margin of pseudorostral lobes not so beset — 9.
Anterior margin of pseudorostral lobes in 2 denticulate
throughout ... . 6. E. hirsuta
Anterior margin of arendorcerr™l Tene ona dennienler:
throughout — 8.
Uropods with 2nd joint of a aed not one-fourth of 1st,
~I
————. —_——
P | apical spine fused . . . 7. E. hispida
| Uropods with 2nd joint of endopod ¢ one- eeeah af Ist, aie
spine distinct ..... Ne BeOS ae he . 8. BE. gracilis
| Pseudorostral lobes in 2 with tuft of sons setae on upper
9 ; PRAT OEE trons gure ce ean Wchire ereicaihs, Volt.oasicl wet Tsih : 9. E. splendida
\ Pseudorostral lobes han tuft of pees on upper margin —10.
10 J Uropods with apical spine of endopod fused to 2nd joint — 11.
Uropods with apical spine of endopod distinct — 12.
Pp Pp Pp Pp
u j Pleon clothed with short hairs... . +--+ -+++.-s 10. E. truncatula
lwelconmhairlesss e088 2) cue ois ces ee es > LI). pusilla
12 { Projection within frontal sinus dentate .......+.. 12. E. abyssi
\ Projection within frontal sinus smooth ....... . .13. BE. sordida
J
1. E. emarginata (Kroyer) 1846 Lewcon emarginatus, Kroyer in: Naturh.
Tidsskr., ser.2 v.2 p.181, 209, 211 t.2 f.3a—h | 1849 L. e., Kroyer (Petersen del.) in:
Voy. Nord, Crust. t.5a f.2a—s | 1852 L. e., W. Liljeborg in: Ofv. Ak. Férh., v.9
p.6 | 1863 Cyrianassa ciliata, A. M. Norman in: Tr. Tyneside
Club, v. 5iv p.273 t.13 f.4—9(6) | 1865 EHudora emarginata, ih, oe ve
‘G. O. Sars in: Forh. Selsk. Christian., 1864 p.185 | 1867 Hudo- / oR
rella e., A. M. Norman in: Rep. Brit. Ass., v.36 p. 194, 197 |
1871 E.e., G. O. Sars in: Svenska Ak. Handl., n. ser. v.9 nr. 13
p. 50 t.18 f.98 | 1877 H.e., Meinert in: Naturh. Tidsskr., ser. 3 :
v.11 p.181 | 1879 H.e., S. I. Smith in: Tr. Connect. Ac., v5 ;
p. 115 | 1887 EH. e., H. J. Hansen in: Vid. Meddel., v.39 p. 200 |
1889 E. marginata, D. Robertson in: P. nat. Hist. Soc. Glasgow,
' ser. 2 v.31 Tr. p.47 | 1894 E. emarginata, A.M. Norman in:
Ann. nat. Hist., ser. 6 v.13 p. 275 | 1900 H.e., G.O.Sars, ig. a9,
Crust. Norway, v.3 p.36 +t.27,28 | 1900 H.e., Zimmer in: 4, emaretnata (Kroyer),
Fauna arctica, v.1 p.418, 433 ff. in text | 1901 H. e., Ohlin in: carapace (after Sars).
' Bih. Svenska Ak., v.26 nr. 12 p.43 | 1905 EH. e., Calman in:
Sci. Invest. Fish. Ireland, 1904 app.1 p.21 | 1909 H.e., A. M. Norman & G.S8. Brady
in: Nat. Hist. Tr. Northumb., ser.2 v.31 p.28 | 1911 EH. e., Stappers in: Camp. arct.
‘Orléans, Crust, Malac. p.108 | 1912 H.e., Calman in: P. U.S. Mus., v.41 p. 620.
Body slender, elongate, finely hairy all over. Truncate front fringed
with short hairs, smooth except for a short serrate space reaching the semilunar
76 Cumacea: 9. Leuconidae, 2. Eudorella
sinus, which is bounded below by a somewhat upturned sharp tooth. The sinus
in the G is much smaller than in the Q. The lower margin is serrate for
some distance back from the antero-lateral corner. The long 5™ pleon segment
dorsally projects a pair of long setae over the short telsonic segment. An-
tenna 1 © carrying many plumose setae, 18t joint broader but shorter than
either of the subequal 284 and 34 joints, 3-jointed flagellum as long as 34
joint of peduncle, 1-jointed accessory nearly three-quarters as long as the
long 1st joint of the principal flagellum; in the O the flagellum is 4-jointed,
Fig. 41.
E. emarginata (Kroyer), antennal?
(after Sars).
Fig. 40. Fig. 42.
E. emarginata (d) (Kroyer) E. emarginata (2) (Kroyer), an-
(after Sars). tenna 2 (after Sars).
its 1st joint shorter than the accessory, carrying at the base a dense fascicle
of sensory filaments. Antenna 2 of © with 5 plumose setae on the basal
joint. Peraeopod 1 with 6 joint much longer than 5. Peraeopod 2, ante-
penultimate joint subequal to next preceding joint, terminal joint at least as
long as antepenultimate, broad, spatulate, with many radiating spines. Pe-
duncle of uropods a little longer than the exopod, about equal to the first
joint of the endopod, which is fringed on the medial margin with numerous
unequal spines, and in the CO has also several slender setae on the upper
Cumacea: 9. Leuconidae, 2. Kudorella 77
half; the 2™¢ joint by fusion with the apical spine becomes mucronate, but
even with the fusion is only two-fifths of the length of the 18* joint. Colour
whitish tinged with grey or yellow. LL. about 12 mm.
Norwegian Coast, depths 56—549 m; Spitzbergen, depths 20—435 m; W. Green-
land, 772 m; Atlantic coast N. America, 13—126 m; Heligoland; British Isles, 37—108 m;
Nova Zembla, 61—90 m.
2. B. gracilior Zimmer 1907 E.g., Zimmer in: Zool. Anz., v.37 p. 228 tf. 2, 3 |
1909 E. g., Zimmer in: Ergeb, Schwed. Siidpol.-Exp., v6 nor.3 p.2, 12 t.3 f. 53—55,
t.4 f. 56—68.
Body and limbs as compared with /. sordida (nr. 13) somewhat more
slender. Truncate front fringed with short hairs and smooth except for a
very small denticulate prominence above the smooth sinus, which is bounded
below by a slightly upturned tooth followed by slight denticulation of the
lower margin; the sinus, of moderate depth in the ©, is almost evanescent
in the G. The 5t pleon segment dorsally projects a group of (4) setae
over and beyond the short telsonic segment, which is more produced between
the bases of the uropods in the d than in the Q, ending conically in the
former, almost squarely in the latter. Antenna 1 ©, with many plumose
setae, 222 joint of peduncle the longest, equal to the 4-jointed flagellum,
accessory as long as 1s? joint of the principal. Antenna 2 of © 1-jointed,
with 4 plumose setae. Peraeopod 1, 6 joint more than twice as long as
7th, 5th intermediate in length. Peraeopod 2 having the widened terminal
joint at least as long as antepenultimate, begirt with spines. Peduncle of uropods
a little shorter than the endopod but longer than its 18* joint, having on its
upper side near the end a seta which reaches beyond the rami; exopod
broad, two-thirds as long as endopod, 1st joint of the latter between 3 and
4 times as long as the 2"4, its medial margin fringed with numerous unequal
spines, apical spine of the 2™¢ joint distinct. Colour white. L. about 7 mm.
S. Georgia, depths 75—310 m.
3. E. fallax Zimmer 1909 E. 7., Zimmer in: Ergeb. Schwed. Siidpol.-Exp.,
v6 nr.3 p.2,9 t.2 £.32—389, t.3 f. 40—47.
Pseudorostral lobes with very small hairs on the rounded top, scarcely
any on the slightly convex front which descends without serration to the
shallowly quadrate sinus bounded by a small antero-lateral tooth, with very
feeble serration of the lower margin. In the d even not adult the sinus
disappears. First pedigerous segment firmly united with the carapace. The
5th pleon segment dorsally projects a succession of 6 setae over the short
telsonic segment. Antennal Q, 2™4 joint considerably longer than 1s‘ or 34,
flagellum 4-jointed, accessory almost as long as 1* and 2"¢ joints of principal
combined. Antenna 2 © with 3 setae on 15* joint, small 2"¢ joint faintly indi-
cated (in figure). Peraeopod 1, 6 joint not much longer than 5‘. Perae-
opod 2, spinose 7* joint as long as 5 and 6% combined, firmly fixed in
an excavation of the 6%, Peraeopod 5 comparatively very small. Pedunele
of uropods rather shorter than endopod, of which the 1St joint is subequal
to the exopod and more than twice as long as the 2"? joint, with setae on
the upper part and unequal spines on the lower part of its medial margin,
the series being continued on the 24 joint, which ends in a strong distinct
spine. Colour white. L. © varying round 6°5 mm.
S. Georgia, depths 64—310 m.
78 Cumacea: 9. Leuconidae, 2. Eudorella
4. E. nana O. Sars. 1879 EH. n., G. O. Sars in: Arch. Naturv. Kristian., v4
p. 40 t. 33.
Very like E. truncatula (nr. 10), but much smaller. Pseudorostral lobes
with very few hairs on rounded upper corner, thence descending smoothly
to a single upturned tooth in Q, followed by 2 downward pointed teeth and
then a sinus bounded by a sharp forward directed tooth with some serration
of the lower margin; in the G a somewhat sinuous front is bounded by a
forward directed tooth without other teeth or serration. The 5‘ pleon segment
projects from its dorsal end a group of setae far over the conically produced
telsonic segment. Antenna 1, flagellum in Q 3-jointed, accessory longer than
1st joint of principal, but not as in O equal to 18* and 2"¢ joints combined,
flagellum in G 4-jointed. Peraeopod 1 slender, 6' joint’ somewhat longer
than 5th, twice as long as 7%, Peraeopod 2 having terminal joint as long
as antepenultimate, beset with spines but not densely. Peduncle of uropods
slightly shorter than the exopod, which is slightly longer than 1st joint of
the endopod; lower part of peduncle and upper part of endopod having on
medial margin in G serrate setae in addition to the fringing spines; 2"¢ joint
of endopod one-third the length of the 1', apart from the apical spine
which appears however to be completely fused with the joint, medial margin
having only a subapical spinule. Colour whitish, semipellucid. L. 3 mm.
Mediterranean, Bay of Spezia, depth 38—56 m; Bay of Naples.
5. E. monodon Calm. 1912 E. m., Calman in: P. U.S. Mus., v 41 p. 622
tf. 21—24.
Near to E. hirsuta (nr. 6), but serration at upper part of pseudorostral
lobes pointing downwards, setae on surface of body sparse and inconspicuous,
except those projecting backwards from 5t* pleon segmeut. A minute pro-
curved tooth is situate on the mid-dorsal line of the carapace about one-
third of its length from the front. Telsonic segment short, about four-fifths
as long as the peduncles of the uropods, between the bases of which it is
well produced. Antenna 1, 2"¢ joint dilated, about two-thirds as broad as
long, accessory flagellum not nearly as long as 1** joint of the principal.
Peraeopod 1 with 6% joint longer than 5 and more than twice the 7%,
Peraeopod 2 with 5' jomt about equal to 4‘ and little longer than 6% and
7th combined. Peduncle of uropods as long as the endopod, 1%* joint of
which is 4 times as long as the 2"4, carrying numerous spines on the medial
margin and setae on the lateral; exopod shorter than the endopod, slightly
longer than its 18¢ joint. L. © 4°7 mm, ¢ unknown.
Beach near Calcasien Pass, Louisiana.
6. E. hirsuta (O. Sars) 1869 Eudora h., G. O. Sars in: Nyt Mag. Naturv., v.16
p-317, 847 | 1871 Eudorellah., G. O. Sars in: Svenska Ak. Handl., n.ser. v.9 nr. 13 p.44 t.18
f. 100 | 1879 HE. h. (var.?), A. M. Norman in: Ann. nat. Hist., ser. 5 v.3 p.72 | 1894 H. h.,
A. M. Norman in: Ann. nat. Hist., ser.6 v.13 p.275 | 1900 HE. h., G. O. Sars, Crust.
Norway, v.3 p.38 t.30 | 1900 E.h., Zimmer in: Fauna arctica, v.1 p. 419, 433 tf., 440.
3ody very slender and elongate. Pseudorostral lobes clothed in upper
part with forward curving setae, the rather sinuously truncate front
margin denticulate in Q with upward pointing teeth as far as the sinus,
which is nearly filled by a prominence serrate downwards and bounded by
a very small forward pointing tooth at the end of the serrate part of the
lower margin (antero-lateral angle well rounded off, var. Norman); marginal
Cumacea: 9. Leuconidae, 2. Eudorella 79
denticles and sinus wanting in GO even when young, adult not known. An-
tenna 1, 2™4 joint considerably longer than 18* or 34, accessory not so long
as 18¢ joint of principal flagellum in Q, but longer than 1%* in the 4-jointed
flagellum of SO. Peraeopod 1, 6% joint scarcely longer than 5". Peraeopod 2,
terminal joint less than half the antepenultimate. Peduncle of uropods rather
shorter than exopod, which is longer than 1s* joint of endopod; endopod’s
gnd joint more than a third as long as the 1%*, with 3 spines on its medial
margin and a quite distinct apical spine. Colour whitish, L. 5 mm.
Coast of Norway, depths 282—377 m; between Ireland and Rockall, 2524 m
(Norman).
7. BE. hispida O. Sars 1871 E. h., G. O. Sars in: Ofv. Ak. Forh., v.28 nr. 1
p.80 | 1871 EF. h., G. O. Sars in: Svenska Ak. Handl., n. ser. v.9 nr.13 p.49 t.18
f, 95—97 | 1878 E. h., S. I. Smith in: A. KE. Verrill in: Rep. U. S. Fish. Comm., v1
p. 555 | 1879 E.h., S. I. Smith in: Tr. Connect. Ac., v.51 p. 115 | 1900 EH. h., Zimmer
in: Fauna arctica, v.1 p. 419, 433 tf., 440 | 1905 H. h., Calman in: Sci. Invest. Fish.
Ireland, 1904 app. 1 p. 21.
Near to 2. emarginata (nr. 1). Body very narrow and elongate. Pseudo-
rostral lobes clothed in upper part with forward curving setae, the smoothly
truncate front ending in a tridentate prominence, upper boundary of the
sinus, within which are 2 downward pointing teeth (but this outline variable
Calman), the lower boundary being formed by a sharp tooth strongly pro-
duced forwards and followed by considerable serration of the lower margin.
Fifth pleon segment much the longest, not carrying long backward directed
setae, but like all the rest of the integument closely beset with short hairs.
Antenna 1, 34 joint subequal in length to the 2"¢, flagellum of © figured as
3-jointed; with 18* joint longer than the 1-jointed accessory. Peraeopod 1,
6th joint very little longer than 5%, Peraeopod 2, antepenultimate joint
(true 5%) nearly double as long as the next preceding joint, the 3% joint
being lost or fused; in yeung male antepenultimate joint once and a half
as long as preceding joint (Calman). Peduncle of uropod stout, as long as
the exopod which is very little shorter than the endopod; of the latter the
second joint is less than a quarter the length of the 15* joint, or with inclusion
of the coalesced terminal spine it is half the length of the 15*; spines on
medial margin of peduncle 8, 1%¢ joint of endopod 12, 2™¢ joint 3; there are
numerous setae on both rami. L. © juv. 55 mm, C juv. 5 mm.
Atlantic coast of N. America, off Nova Scotia, depth 126m; Bay of Fundy,
ete., 2—99 m; W. of Ireland, 366 and 585 m.
8. E. gracilis 0. Sars 1871 E. g., G. O. Sars in: Ofv. Ak. Férh., v.28 nr. 6
p. 800 | 1873 E. g., G. O. Sars in: Svenska Ak, Handl.,n. ser. v.11 nr.6 p.8 t.3 f.11—13
1900 E. g., Zimmer in: Fauna arctica, v.1 p. 419, 433 tf., 440.
Body very slender and elongate. Pseudorostral lobes on rounded upper
front margin carrying hairs of moderate length, otherwise smoothly truncate
down to quadridentate upper boundary of the sinus, within which is a bidentate
prominence, the lower boundary being a small horizontal antero-lateral tooth
followed by slight serration of the lower margin. Pleon hairy, but the long
5th seoment not projecting long setae over the telsonic segment, which is
well produced conieally, with rounded apex, between bases of uropods. Antenna 1,
34 joint little over half length of 24, flagellum 3-jointed, accessory as long
as 1st joint of principal. Peraeopod 2 rather large, 3° joint wanting or fused,
5th considerably longer than the next preceding, terminal oval or broadly
80 Cumacea: 9. Leuconidae, 2. Eudorella
lanceolate, surrounded with radiating spines. Peduncle of uropods scarcely
shorter than endopod, of which the 1t joint is thrice as long as the second,
the medial margins carrying respectively 7, 8, and 3 spines, the terminal
spine on 2"4 joint distinct, not so long as the joint; exopod a seventh shorter
than the endopod. Integument pellucid. L. ovigerous Q about 6°5 mm,
unknown.
Spitzbergen, depth 1017 m.
9. E. splendida Zimmer 1902 EH. s., Zimmer in: Ergeb, Hamburg. Magalh.-
Sammelr., v.2 Cumac. p.5, 6 tf.6,7 | 1907 EH. similis, Calman in: Nat. antarct. Exp.,
v.2 Crust. Cum. p.3 tf. 1—6 | 1909 H. splendida, Zimmer in: Ergeb. Schwed. Siidpol.-
Exp, 216 “nrgo: po lt:
Pseudorostral lobes in © subadult (Calman) with tuft of setae on the
top. in Q and young CG the front margin smoothly truncate down to the sinus
which has an almost smooth median prominence and is bounded below by a
horizontal tooth followed by a smooth lower margin; in adult G (Calman)
the front margin is strongly convex, projecting well in front of the single
small tooth which defines it below. Fifth pleon segment projecting a pair
of long setae over telsonic segment (Calman, not Zimmer, whose specimens
were imperfect). Antenna 1 Q, flagellum 3-jointed, 4-jointed with accessory
as long as first 2 joints of principal combined. Peduncle of uropod subequal
to the exopod; endopod longer, with terminal spine fused to 2™4 joint (Zimmer)
or distinet (in figure by Calman, who likens the uropods to those of E. trun-
catula, in which Sars figures the terminal spine as fused). Colour iridescent.
L. 6 6 mm, © subadult 5°8 mm.
S. Georgia, edge of low-tide; Coulman Island, depth 183 m.
10. EB. truneatula (Bate) 1856 Hudora t., Bate in: Ann. nat. Hist., ser.2 v.17
p.457 t. 14 f.3 | 1857 EH. ¢., A. White, Hist. Brit. Crust., p. 151 | 1859 E. ¢., Danielssen in:
Norske Selsk. Skr. 19. Aarh., v.41 p.108 | 1865 E. t., G. O. Sars in: Forh. Selsk.
Christian., 1864 p.186 | 1869 Eudorella t., A. M. Norman in: Rep. Brit. Ass., v.¢8
p.272 | 1871 E.t, G.O.Sars in: Svenska Ak. Handl., n. ser. v9 nr. 13 p.44 t.18
f.99 | 1877 E. inermis + E. t., Meinert in: Naturh. Tidsskr., ser.3 v.11 p. 183 (d) |
1879 E.t., A. M. Norman in: Ann. nat. Hist., ser.5 v.3 p.72 | 1879 E.t., G. O. Sars
in: Arch. Naturv. Kristian., v.4 p.34 t.30—32 | 1894 H#. ¢t., A. M. Norman in: Ann.
nat. Hist., ser.6 v.13 p.275 | 1897 E.t., Ehrenbaum in: Wiss. Meeresunters., v.21 p.412 |
1900 FE. ¢t., G. O. Sars, Crust. Norway, v.3 p.37 t.29| 1900 EH. ¢, Zimmer in: Fauna
arctica, v.1 p. 419, 433 tf., 440 | 1905 EH. t., Calman in: Sci. Invest. Fish. Ireland, 1904
app. 1 p.¥1 | 1907 E. ¢., Calman in: Tr. zool. Soe. London, v.181 p.33 | 1909 F. Tt,
A.M. Norman & G. S. Brady in: Nat. Hist. Tr. Northumb., ser.2 v.311 p.28 | 1912 E. t.,
Calman in: P. U.S. Mus., v.41 p. 620.
Integument thin and flexible, beset with short hairs, clearly squamose.
Body slender, especially the pleon. Pseudorostral lobes in the © with a few
stiff setae on the rounded upper front corner, smoothly truncate in front down
to the denticulate, slightly prominent upper boundary of the sinus, the middle
of which is occupied by a prominence with 3 downward pointing denticles,
while its lower boundary is formed by an upward pointing tooth followed
by some serration of the lower margin. In the male the sinus is replaced
by a smooth convexity. The 2™4 and 3¢ pedigerous segments have the side-
plates rather widely outstanding. The 5% pleon segment does not project
long setae over the short telsonic segment. Antenna 1, 3¢ joint shorter
than 24, flagellum in © 3-jointed, its long 18* joint scarcely longer than the
1-jointed accessory; flagellum in CG 4-jointed, with a tuft of long filaments
Cumacea: 9. Leuconidae, 2. Eudorella 81
on the 2"4 joint. Antenna 2 Q with 3 plumose setae and no division into
9 joints. Maxilliped 3 with a hooked spine on medial margin of the 2"¢ and
another on that of the 4 joint. Peraeopod 1 long and slender, its 7 joint
half as long as 6‘, and about three-quarters as long as 5%", Peraeopod
2 rather stout, not long, its terminal joint spade-like surrounded by a moderate
number of radiating spines. Peduncle of uropods a little longer than the
exopod, a little shorter than the endopod, in which the terminal spine is
fused with the 24 joint; 2" joint apart from the spine about two-fifths of
1st in ©, about one-third in the G, carrying 3 spines on medial margin
in ©, in CO 5 (Mediterranean), 1 (Norwegian). Colour whitish, semipellucid.
L. about 5 mm.
S. and W. Norway, to Lofoten Is., 11—56 m; Mediterranean, 38—47 m; various
seas between these latitudes, at various depths from shallow water, W. Ireland, through
depths reaching 549 m, at Trondhjemfjord and at lat. 55° 11’ N., long. 11° 31’ W.
2639 m; U. S. A., Vineyard Sound, etc., depths 29—97 m.
11. BE. pusilla O.Sars 1871 EH. p., G. O. Sars in: Ofy. Ak. Forh., v.28 vr. 1
p.79 and Svenska Ak. Handl., n. ser. v.9 nr.13 p.46 t.16 f.76—78; t.17 £.79—94 |
1879 E. p., S.1. Smith in: ‘fr. Connect. Ac., v.51 p.116 | 1900 EF. p., Zimmer in:
Fauna arctica, v.1 p.419, 433tf., 440 | 1912 EH. truncatula, Calman in: P. U.S. Mus.,
v.41 p. 620.
In shape and size very near to E. truncatula (nr. 10), but distinguished
by the upper margin of the pseudorostral lobes which is vaulted in profile,
not flattened, and the frontal part almost entirely without setae; the pleon
also is without trace of hairs. Calman mentions a procurved dorsal tooth
on the carapace close to hind margin of respiratory opening as occasionally
present. Antenna 1 comparatively stronger, with the 3¢ joint relatively short
and more widened apically; maxilliped 3 without hooked spine on 2™4 or 48
joint (as figured); peduncle of uropod as long as 1%* joint of endopod which
is longer than the exopod and 4 times as long as the 2"4 joint not including
the fused terminal spine. L. ovigerous 9 4°5 mm, CO undescribed.
Atlantic coast of N. America, depths 2—46 m.
12. KE. abyssi O. Sars 1887 E.a., G. O. Sars in: Rep. Voy. Challenger, v.19
pars 55 p.9,41 t.5 f.5—12.
Body moderately slender, integument thin, beset with scattered short hairs.
Pseudorostral lobes smoothly truncate in front down to tridentate prominence
bounding the sinus, which is nearly filled by quinquedentate prominence and
bounded below by small antero-lateral tooth with succeeding serration of
lower margin. Pleon segment 5 without backward projecting setae. Antenna
1 rather stout, flagellum 3-jointed, 1-jointed accessory as long as 1%* joint
of principal. Peraeopod 1, 7 joint half as long as 6", which is a little
longer than the 5%, Peraeopod 2, terminal joint almost linear, but as usual
fringed with divergent spines. Peduncle of uropods about equal to endopod,
in which the 2™4 joint is less than a quarter as long as the 1S*, with 4
spines on medial margin and a distinct terminal spine, the 1%* joint as
long as the exopod and fringed with numerous spines. L. Q about 9 mm,
GO unknown.
N. Atlantic, lat. 38° 34’ N., long. 72° 10’ W., depth 2268 m.
13. E. sordida Zimmer 1907 E. s., Zimmer in: Zool. Anz., v.31 p. 227 tf. 1 |
1909 E. s., Zimmer in: Ergeb. Schwed. Siidpol.-Exp., v.6 nr. 3 p.10 t.3 f. 48—52.
Das Tierreich. 39. Lief.: T. R. R. Stebbing, Cumacea. 6
82 Cumacea: 9. Leuconidae, 2. Eudorella, 3. Eudorellopsis
Extremely like L. fallax (nr. 3), but the carapace slightly more slender,
the pseudorostral lobes meeting very far to the rear, the respiratory opening
at the top surrounded by long setae, the sinus in front not quadrate but
occupied by a prominence giving a sinuous outline bounded below by an
antero-lateral tooth, variable yet decidedly larger than in the other species,
the same superiority attaching to this tooth also in the male; setae on the 5th
pleon segment either wanting or not nearly as long as the telsonic segment;
antenna 1, with accessory flagellum only about as long as the 1%* joint of
the principal. L. about 7 mm.
S. Georgia, depths from 12—250 m.
3. Gen. Kudorellopsis 0. Sars
1883 Eudorellopsis, G. O. Sars in: Forh. Selsk. Christian., 1882 nr. 18 p. 12, 56 |
1900 E., G. O. Sars in: Crust. Norway, v.3 p.89 | 1900 E., Zimmer in: Fauna arctica,
v.1 p.431.
Near to Eudorella, but body short and compact, pleon relatively short.
First antenna geniculate between the first and second joints. Uropods with
the exopod longer than the endopod.
4 species.
Synopsis of species:
J Pseudorostrum with upturned process behind branchial orifice 1. EH. deformis.
| Pseudorostrum with upturned process frontal — 2.
Sides of carapace: smooth’. = «>= «= =)si*) -. =e see ee 2. EB. integra
Sides of carapace not smooth — 3.
3 J Carapace with longitudinal ridge on each side ..... . . 3. H.resima
) Carapace with 2 oblique curved ridges on each side. . .. . 4. HE, biplicata
1. FE. deformis (Kréyer) 1846 Leucon d., Kroyer in: Naturh. Tidsskr., ser. 2
v.2u p.194,209 t.2 f.4 | 1849 L. d., (Petersen del.) Kroyer in: Voy. Nord, Crust.
t.5a f.3a—h | 1865 Eudora d., G. O. Sars in: Forh. Selsk. Christian., 1864 p. 60 | 1871
Eudorella (2) d., G. O. Sars in: Svenska Ak. Handl., n. ser. v9 nr.13 p. 44,50 t. 1g
f. 101—103, t.20 f.104—118 | 1877 E. d., Meinert in: Naturh. Tidsskr., ser.3 v.11 p.184 |
1883 Eudorellopsis d., G. O. Sars in: Forh. Selsk. Christian., 1882 nr. 18 p. 12, 56 | 1886
E. d., G. O. Sars in: Norske Nordhavs-Exp., v.6 nr. 15 p.20 | 1887 HE. d., H. J. Hansen
in: Vid. Meddel., v.39 p.200 | 1897 E. d., Ehrenbaum in: Wiss. Meeresunters., v.21
p.413 | 1900 E. d., G. O. Sars in: Crust. Norway, 0.3 p.40 t.31, 32 | 1900 E. d., Zimmer
in: Fauna arctica, v.1 p. 420, 433, 440 | 1905 H.d., Calman in: Sci. Invest. Fish. Ireland,
1904 app.1 p.22 | 1909 HE. d., A. M. Norman & G. 8. Brady in: Nat. Hist. Tr. Northumb.,
ser.2 v.3u p.29| 1912 E.d., Calman in: P. U.S. Mus., v.41 p. 623.
Pseudorostral lobes with sharp process erected to the rear of the dorsal
branchial orifice, front steeply truncate and fringed throughout in Q but less:
completely in G with broad lamellar teeth, antero-lateral corners triangularly
produced in ©, smooth above, with serrate lower margin. Pleon segments 1—5
with ventral setae. Telsonic segment with 2 setae and 2 setules (figured)
on hind margin. Antenna 1, 3¢ joint shorter than 15*, longer than 24, flagellum
3-jointed, accessory 1-jointed, about half as long as 1** joint of principal.
Peraeopod 1, 5% and 6 joints subequal, 7** shorter and very narrow. Peraeo-
pod 2, terminal joint at least as long as antepenultimate, with 5 apical
Cumacea: 9. Leuconidae, 3. Eudorellopsis 83
and 2 lateral spines. Uropods more robust in Q than in GC, the peduncle
in the former longer than the endopod, but shorter in the latter, in both
sexes shorter than the exopod, which is much longer than the endopod and
in the © is very broad, carrying on the obliquely truncate apex several setae
and rows of strong spines on its dorsal sur-
face; this ramus in the dis more slender, less
strongly spined but carrying longer setae;
2nd joint of endopod small, a third as long
as the 1S* joimt or less, with small apical
spine. L. OQ 5mm, GO 4°5 mm.
Greenland, Iceland, Norway, Denmark,
German Ocean, Irish Sea, N. of England, U. S. of
America; surface to 56m; off Nova Scotia, depth
103 m.
Fig. 43.
E. deformis (¢) (Kroyer) (after
Sars).
Fig. 44.
E. deformis (3) (Kroyer),
uropod (after Sars).
2. E. integra (S. 1. Smith) 1879 Eudorella (?)7., S. 1. Smith in: Tr. Connect.
Ae., v.51 p.116 | 1887 Hudorellopsis i., H. J. Hansen in: Vid. Meddel., v.39 p.201 t.7
f. 3,3a—d | 1893 H.2., T. Stebbing in: Int. sci. Ser., v.74 p.305 | 1900 E.i., G. O.
Sars, Crust. Norway, v.3 p.39 | 1900 EH. i., Zimmer in: Fauna arctica, v.1 p. 420, 433,
440 | 1912 H.7., Calman in: P. U.S. Mus., v.41 p. 624.
Pseudorostral lobes with dorsal process acute, erected close behind the
frontal margin, which is smooth, bulging above, then descending with slight
obliquity to the little produced antero-lateral rounded corner, followed in
the © by faint denticulation of the lower margin; in the CO no denticulation
6*
\
84 Cumacea: 9, Leuconidae, 3. Eudorellopsis
and the corner not produced. Carapace sometimes with minute denticle dor-
sally one-third of length from front. Pleon segments without ventral setae.
Hind margin of telsonic segment, in: only, carrying 4—6 conspicuous setae.
Antenna 1 © nearly as in E. deformis (nr. 1), but 1-jointed accessory equal
in length to 1St joint of principal flagellum (H. J. Hansen), considerably
shorter than that joint (Smith) though relatively larger than in E. deformis;
in G flagellum 4-jointed, accessory 2-jointed. Peraeopod 1, 7 joint little
shorter than 6. Uropods, like the limbs, less robust than in the preceding
Species; peduncle in both sexes shorter than the endopod, in which the 2"¢ joint
is half as long as the 18* in QO, a little less in GO, where both joints are
strongly setose; exopod considerably longer, tapering, slightly curved outward
at the apex, carrying several setae on distal part of medial margin and at
the apex, very long and closely set in the d. L. © about 5 mm, GC a very
little longer.
W. Greenland, Atlantic coast of N. America, depths 28—506 m; Bering-Sea,
depths 52—65 m.
3. E. resima Calm. 1907 H. resimus, Calman in: Tr. zool. Soe. London,
v. 181 p.33 t.8 f. 6—10.
Pseudorostral lobes directed upwards with slight inclination forwards,
distal end truncated, in lateral view a broad triangle rising above the dorsal
line of the carapace, frontal margin coarsely serrate, receding to the small
antero-lateral tooth which defines a very small sinus. Carapace with forward
pointing median tooth near the end, the sides traversed from end to end
by a curved ridge. Antenna 1, 1-jointed accessory as long as 1%* joint of
3-jointed principal flagellum. Peraeopods 1, 2 short. Uropods short and
stout, exopod nearly twice as long as peduncle, roughened on surface
with irregular tubercles or blunt teeth; endopod only a little shorter, 1%*
joint twice as long as 24, apical spine long and stout. L. Oo 1:75 mm, do
unknown.
New Zealand, Lyttleton Harbour.
4. KE. biplicata Calm. 1912 EH. b., Calman in: P. U.S. Mus., v. 41 p. 625
tf, 25, 26.
Pseudorostral lobes nearly vertical, acute, front margin concave. Cara-
pace little longer than deep, front margin sinuous with
upper part faintly serrate, sides having each 2 oblique
curved ridges. Pedigerous segment 1 firmly united to
the carapace. Telsonic segment about as long as the
short broad peduncle of the uropods, the produced
portion broadly rounded and marked off by a distinct
transverse groove. Maxilliped 3 with 2™¢ joint nearly
twice as long as the rest combined. Peduncle of
uropods scarcely two-thirds as long as the exopod,
which has the apex narrowed and curved outward;
the rather shorter endopod has a 1S* joint thrice as
Fig. 45. 5 : = :
E, biplicata Calman, long as the 2™¢, which is tipped with a short stout
telson and uropod. spine. L. O 55 mm, GO unknown.
Atlantic lat. 45° 29’ N., long. 55° 24‘ W., depth 121 m; lat. 45° 04‘ 00” N,,
long. 59° 36’ 45“ W., depth 103 m.
Cumacea: 9. Leuconidae, 4. Pseudoleucon 85
4. Gen. Pseudoleucon Zimmer
1903 Pseudoleucon (Sp. un.: P. sorex), Zimmer in: Zool. Jahrb., Syst. v.18 v1
p. 676 | 1907 P., Calman in: Tr. zool. Soc. London, v.181 p. 31, 34 | 1908 P., Zimmer
in: Ergeb. Tiefsee-Exp., v.8 p. 176.
Body slender, laterally compressed. Pseudorostral lobes forming an
acute obliquely upward directed pseudorostrum. Branchial orifice not dorsal.
Pleon short. First antenna strong, geniculate between first and second joints.
Second antenna of female with 2 plumose setae. Maxilla 1 with small process
on lateral margin of palp. Peraeopod 2 with 3% joint short but distinct.
Uropods with endopod much shorter than exopod.
1 species.
1. P. sorex Zimmer 1903 P.s., Zimmer in: Zool. Jahrb., Syst. v.18vi p.677
tf. s—v.
Fig. 46.
Fig. 47. Fig. 48. Fig. 49.
P. sorex (2) Zimmer, F.sorex (2) Zimmer, P. sorex (2) Zimmer,
antenna i. maxilliped 3. uropod.
The pseudorostral lobes meet immediately behind the minute eyelobe,
with edges in contact to the obliquely raised acute apex, to which a small
seta is attached, frontal margin except near narrowed apex serrate with in-
tervening setules, sinus small with bulge adjoining the small antero-lateral
86 Cumacea: 9. Leuconidae, 4. Pseudoleucon, 10. Diastyloididae, 1. Diastyloides
tooth. Carapace with a transverse pair of denticles behind the eyelobe, and
a longitudinal couple a little further back. Pedigerous segments together
longer than carapace, 1s* short, 2™4 long, last four well separated, with short
setae on antero-lateral margins. Pleon ventrally with scattered plumose setae,
telsonic segment with setae dorsally on the hind margin. Antenna 1, 3¢ joint
longer than 2"4, Peraeopod 1 with small seta at apex of 7 joint. Peraeopod 2,
7th joint tipped with several setae, one very long, this joint longer than
the 6, but shorter than either the 4% or the 5%. Peraeopods 3—5 with
7* joint minute. Peduncle of uropods as long as endopod, two-thirds of
the length of the exopod, with oblique series of setae on surface and several
setae on apical margin, some very long; second joint of endopod half as
long as 1t, tipped with a strong spine; on the long outward curving exopod
there are many spines, passing distally into plumose setae. Colour brown.
L. about 4°5 mm.
Enosima, Japan, depth 6—23 m.
10. Fam. Diastyloididae
1812 Diastyloididae, T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 133, 146.
Telson narrowly elongate, with only 2 apical spines. Antenna 1 with
slight flagella. Antenna 2 of female not strongly developed, flagellum in
male elongate. Mandibles with base of trunk broad. Exopods on peraeopods
1—4 in male, only on the first 2 pairs in female. Only 2 pairs of pleopods
in the male, the small rami of the 18 both single-jointed, in the 24 confluent
to a single piece. Uropods with slender rami, the 3-jointed endopod not
differing much in length from the exopod.
1 genus.
1. Gen. Diastyloides 0. Sars
1900 Diastyloides, G. O. Sars in: Crust. Norway, v.3 p.60 | 1900 D., Zimmer
in: Fauna arctica, v.1 p. 426,431 | 1908 D., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 181,
182 | 1912 D., T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 146.
Eye wanting. Antenna 2 of female without apical plumose seta on
small terminal joint. Peraeopod 1 with 6% joint elongate. Peraeopod 2
with 24 joint apically produced.
2 species.
Synopsis of species:
{ Carapace without pairs of oblique carinae. . .. +. - - 1. D. serratus
\ Carapace with 2 pairs of oblique carinae ......-. - 2. D. biplicatus
1. D. serratus (O. Sars) 1865 Diastylis serrata, G. O. Sars in: Forh. Selsk.
Christian., 1864 p.169 | 1894 D. serratus, A.M. Norman in: Ann. nat. Hist., ser.6 v.13 p.276 |
1900 Diastyloides serrata, G. O. Sars, Crust. Norway, v.3 p. 61 t. 45 | 1900 D. s., Zimmer
in: Fauna arctica, v.1 p.426 (with synonymy) | 1905 D.s., Calman in: Sci. Invest. Fish.
Ireland, 1904 app. 1 p. 47.
Pseudorostral lobes much produced, especially in © in which the acu-
minate tip is slightly depressed; antero-lateral corners obsolete. Carapace
smooth in G, large in © with serrate lower margin and branchial regions
roughened with minute forward pointing denticles; eyelobe minute. Pedi-
Cumacea: 10, Diastyloididae, 1. Diastyloides 87
gerous segment 5 with postero-lateral angles acute, in the d produced. Telson
much shorter than peduncle of the uropods, with 5 pairs of lateral spines
on distal half in the Q, longer and much narrower with angular projection
in the G. Antenna 1 with 1St joint shorter than 2™4 and 3¢ combined, the
principal flagellum small, 5-jointed in G, 3-jointed in Q, accessory 3-jointed.
Antenna 2 of © with 3 terminal joints very short. Mandibles with conical
process over the molar, spines of spine-row not numerous. Peraeopod 1,
Fig. 52.
D. serratus Sars,
pleopod 1.
Fig. 53.
Fig. 50.
D. serratus (Sars), Fig. 51. D. serratus (Sars),
mandible. D. serratus (Sars), peraeopod 1. pleopod 2.
Qnd joint spinose, much shorter than rest of limb, 68 joint shorter than 24
more than twice as long as 7*, Peraeopod 2 with spine on apex of the
34 joint. Peduncle of uropods longer than the rami, the endopod rather the
longer. especially in the G, the first joint much longer than the 2™¢ and 3¢
combined, spines on medial margin of the 3 respectively in the 9 9, 2, 1,
in the oO 13, 3, 2. Colour yellowish white. L. Q 6mm, G7 mm.
Norway, Christianiafjord, 56—188m, West Finmark, 565m; off Skagen lighthouse;
off Yorkshire coast, 7—11m; Mediterranean, near Capri, depth 950 m.
2. D. biplicatus O. Sars 1865 Diastylis biplicata, G. O. Sars in: Forh. Selsk.
Christian., 1864 p.171 | 1867 D. lamellata, A. M. Norman in: Rep. Brit. Ass., v.36 p.197,
200 | 1879 D. calveri, $ A. M. Norman in: Ann. nat. Hist., ser.5 v3 p.63 | 1894
88 Cumacea: 10. Diastyloididae, 1. Diastyloides, 11. Diastylidae
D. biplicatus, A. M. Norman in: Ann. nat. Hist., ser.6 v.13 p.276 | 1897 D. biplicata,
T. Scott in: Rep. Fish. Board Scotl., v.15 p.134 | 1900 Diastyloides b., G. O. Sars,
Crust. Norway, v.3 p.62 t.46 | 1900 D. b., Zimmer in: Fauna arctica, v.1 p.426 (with
synonymy) | 1905 D. 6., Calman in: Sci. Invest. Fish. Ireland, 1904 app. 1 p.5,
48 | 1909 D.b., A. M. Norman & G.S.
{ | z Brady in: Nat. Hist. Tr. Northumb.,
5 a ITE. ser.2 v.31 p.33.
EL Pseudorostral lobes well pro-
Va duced, slightly upturned, antero-
lateral corners obsolete. Carapace
with 2 pairs of lateral oblique
carinae, and hind margin also cari-
nate, with an infero-lateral carina
added in the G, only anterior half
of lower margin serrate. Telson
shorter than peduncle of uropods,
with 6 pairs of lateral spines,
geniculate in 3 at about one-third
of its length. Peraeopod 1 elongate,
especially in regard to the 6" joint,
which is as long as the 2™¢, Peraeo-
pod 2 with both 2™4 and 3° joints
produced to a spiniform apex.
Peduncle of uropods in the female
little longer than the endopod, which
exceeds the exopod by its 3¢ joint
or more, spines on medial margin
of its 3 joints respectively in Q 9,
2,1, in d 20, 4, 2. Colour tinged
Fig. 54. b F x
D. biplicatus (Sars), peraeopod 1. with reddish brown. L. QO 7 mm,
S about 8 mm.
Norway, with D. serratus; Skagerrak and Kattegat; round Scotland; N. EK. England,
71—82 m; off W. of Ireland, depth 220—2980 m.
1l. Fam. Diastylidae
1856 Diastylidae, Bate in: Ann. nat. Hist., ser.2 v.17 p.449 | 1859 D., Danielssen
in: Norske Selsk. Skr. 19. Aarh., v.41 p. 108 | 1863 D., A. M. Norman in: Tr. Tyneside
Club, v. 51v p. 271 | 1864 D., Fr. Miller, Fiir Darwin, p.55 | 1878 & 1879 D. (restricted),
G. O. Sars in: Arch. Naturv. Kristian., v3 p. 466; v.4 p.44 | 1887 D., G. O. Sars in:
Rep. Voy. Challenger, v.19 pars55 p.7, 43 | 1893 D., T. Stebbing in: Int. sci. Ser.,
v.74 p. 311, 312 | 1900 D., G. O. Sars, Crust. Norway, v.3 p.41 | 1990 D., Zimmer in:
Fauna arctica, v.1 p. 420,433 | 1908 D., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 181 |
1911 D., Calman in: Tr. zool. Soc. London, v.181v p.342, 366 | 1912 D. (further restricted),
T. Stebbing in: Ann. S. Afr. Mus., v. 10v_ p. 133, 146,
Of the pedigerous segments the first and usually the rest distinct. Telson
well developed, with only two apical spines. Accessory flagellum of first
antenna distinct, mandibles normal, not broad at the base. Palp of first
maxilla with two filaments. Branchial leaflets numerous. Exopods on the
first two pairs of peraeopods, also on the third and fourth pairs in the male
but not in the female, except occasionally as rudiments. Two pairs of pleo-
pods in the male. Uropods with the endopod three-jointed. ;
8 genera accepted.
Cumacea: 11. Diastylidae, 1. Diastylis 89
Synopsis of genera:
1 Telson comparatively long — 2.
Telson comparatively short — 6.
J Post-anal portion of telson long — 3.
~ \ Post-anal portion of telson not long — 5.
3 Peraeopods 2, 3, of adult Q not widely separated . 1. Gen. Diastylis ')
Peraeopods 2, 3, of adult 9 widely separated — 4.
Pseudorostrum in Q strongly upturned and peraeo-
pods 8, 4, without exopods. ... . 2. Gen. Brachydiastylis
Pseudorostrum not strongly upturned, and Be aeopeds
3, 4, in Q with rudimentary exopods ... . . 3. Gen. Diastylopsis
P J Telson with lateral spines on the post-anal portion 4. Gen. Adiastylis
\ Telson without lateral spines on the post-anal portion 5. Gen. Makrokylindrus
{ Maxilliped 3 in 9 without exopod ... . . . 6. Gen. Paradiastylis
Maxilliped 3 with exopod in both sexes — A
Pp P
7 | Lateral spines of telson few or none
\ Lateral spines of telson numerous. ....+...-
=
QR
©
i=)
. Leptostylis
. Ekleptostylis
0
Q
i)
6B
1. Gen. Diastylis Say
1780 Oniscus (part.), Lepechin in: Acta Ac. Petrop., 1778 v.1 p.248 | 1818 Diastylis,
Say in: J. Ac. Philad., v 1m p.313 | 1825 D., A. G. Desmarest, Consid. gén. Crust.,
p- 245 | 1829 Condylura (non Mlliger 1811), Latreille in: G, Cuvier, Régne an., ed.2 v4
p- 153 | 1831 C., Latreille, Cours Ent., v1 p.416 | 1840 Condylurus, H. Milne-Edwards,
Hist. nat. Crust., v.3. p.553 | 1841 Cuma (part.), Kroyer in: Naturh. Tidsskr., v3 v1
p. 508 | 1843 Alauwna, Goodsir in: Edinb. new phil. J., v.34 p. 120, 129 | 1844 Diastylis,
De Kay, Zool. N. York, pars6 p. 32 | 1851 Astaricilla (?), Dalyell, Powers Creator, v. 1
p. 224, Asteracilla, v.1 p.226 t.62 | 1856 Diastylis, Bate in: Ann. nat. Hist., ser.2 v.27
p.45i | 1856 Scorpionura (part.), J. V. Thompson MS. in: W. Thompson, Nat. Hist.
Treland, v.4 p.394 | 1865 Diastylis, G. O. Sars in: Forh. Selsk. Christian., 1864 p. 156,
Goniacis, nom. nud. for D. 3 G. O. Sars, p. 159 | 1900 D., G. O. Sars, Crust. Norway,
v.38 p.42 | 1900 D., Zimmer in: Fauna arctica, v.1 p. 420, 434 | 1909 D., Stappers in:
Cellule, v.25 p.368 (alimentary system) | 1912 D., T. Stebbing in: Ann. S. Afr. Mus.,
v.10v p. 147.
Pseudorostral lobes with antero-lateral corners usually little produced.
Telson long, post-anal portion narrowly produced, elongate, with several pairs
of lateral spines. Second antenna of adult O usually very long. Third maxilli-
ped with exopod in both sexes. Third peraeopod as a rule not widely separated
from the second in the adult female. Both pairs of pleopods in the male
well developed, the endopod two-jointed. Hepatopancreatic glands with three
pairs of lobes.
31 species accepted, 2 doubtful.
Synopsis of accepted species:
Peraeopods 3, 4, of 9 with rudimentary exopods — 2.
Peraeopods 3, 4,.of Q without rudimentary exopods — 6.
9 J Maxilliped 3 with 2nd joint not expanded distally — 3.
\ Maxilliped 3 with 2nd joint expanded distally — 4.
Carapace with 2nd pair of carinae een below to a strong
3 | TOOTH Sma 6 eee cats 8 . 1. D. bidentatus
Carapace with 2nd pair SE carinae ee feningd a tooth « 222) D. dali
1) D. sulcatus is an exception.
90
._ | Uropods with endopod longer hee the exo od — +
6 p I % Pp
~
[0 9)
10
11
13
25
26
27
Cumacea: 11. Diastylidae, 1. Diastylis
Carapace
J with ‘few, ‘Tid BES: <9.) cueso ys aa veyee ee ee eee eS
\ Carapace with many ridges — 5.
J Carapace 6—ridged . 4
| Carapace 10—ridged - 5
\ Uropods with endopod not longer than the exopod — 9.
{ Pedunele of uropods about twice as long as the telson. . 6
| Peduncle of uropods not nearly twice as long as the telson — 8.
{ Telson shorter than fifth segment of the pleon
| Telson longer than fifth segment of the pleen .
j Pedigerous segments 3, +, with dorsal separation faint . . 9
| Pedigerous segments 3, 4, with dorsal separation distinct — 10.
{ Telson longer than peduncle of uropods — 11.
| Telson not:longer than peduncle of uropods — 12.
{ Pedigerous segments and pleon spinulose
| Pedigerous segments and pleon smooth ........
J Post-anal part of telson distinctly the longer — 18.
| Post-anal part of telson not distinctly the longer — 25.
~
Pedigerous segment 5 with hind corners acutely pro-
duced — 14.
]
\
\
j
\
J
Pedigerous segment 5 with hind corners not acutely pro-
duced — 19.
J Carapace of female with very conspicuous spines — 15.
Carapace without very conspicuous spines — 16.
Hind margin of carapace conspicuously upturned
= 10:
- 1h
. D. planifrons
. D. alaskensis
. D. anderssoni
. D. hammoniae
. D. antillensis
. D. koreanus
. D. algoae
D. spinulosus
D. armatus
D. bispinosus
Hind margin of carapace not conspicuously upturned . . 13. D. cornutus
Carapace smooth Fee) CO Oooo Ge oho 6 14. D. laevis
Carapace more or less denticalaie — 17.
J No distinct eye . : 15. D. suleatus
| Eye distinct — 18.
| Peraeopod 2 with 7th joint longer than 6th. .... 16. D. rathkii
| Peraeopod 2 with 7th joint not longer than 6th , . 17. D. bradyi
{ Eye wanting or inconspicuous — 20.
\ Eye distinct — 22.
Pleon segments 1-5 with hind angles acutely produced
| outwards Mae We eth fous Seen aes ie MeoMigs Mien Det eM 18. D. goodsiri
| Pleon segments 1—5 Ai hind qaeles ne prodeced out-
wards — 21.
{ Pleon spinulose . hire PEE eC: BG so) Gi Gro 19. D. stygius
\ Bleon-notrspinulese: Ecce os hol eget eneel eee 20. D. polaris
{ Carapace without transverse pleats — 238.
\ Carapace with transverse pleats — 24.
{ Pleon unarmed . Ie sia oth Se . 21. D. lucifer
| Pleon segments 1—3 with Gear ‘dese: fatcal teeth . . . 22. D. rugosoides
{ Carapace with 5 pairs of obliquely transverse pleats
\ Carapace with fewer than 5 pairs of pleats
{ Eye distinct — 6.
\ Eye wanting or obscure — 27.
{ Pseudorostral lobes not much produced. .... - - 25
\ Pseudorostral lobes well produced . 26
l Pedigerous segment 5 with hind corners acutely prodvosdae 28.
Pedigerous segment 5 with hind corners not acutely pro-
| duced — 29.
. D. seorpioides
. D. rugosus
. D. tumidus
. D.neapolitanus
Cumacea: 11. Diastylidae, 1. Diastylis oT
98 Carapace mapped into polygonal areas by spines and spinules 27. D. echinatus
Carapace girdled by 3 nearly parallel ridges. . . . . . . 28. D. tricinctus
99 j Telson scarcely half as long as peduncle of uropods . . . 29. D. nucella
“ \ Telson nearly as long as peduncle of uropods — 30.
30 { Telson with few lateral spinules ......... =. . . 30. D. anomalus
| Telson with many lateral spinules ........ «9 Slo, Dasper
1. D. bidentatus Calm. 1880 D. sp., Stuxberg in: Bih. Svenska Ak., v5 nr. 22
p- 29 | 1882 D. sp., Stuxberg in: Vega-Exp., v.1 p.715 | 1882 D. picta, Stuxberg in:
A.M. Norman, Mus. Norm., p.10 | 1912 D. bidentata, Calman in: P. U.S. Mus., v.41
p. 605, 637 tf.45—50 | 1912 D. picta, Calman in: P. U.S. Mus., v.41 p.636 (corrected in litt.)
In general like DJ. scorpiordes (nr. 23), but surface of carapace smoother,
the ridges very finely crenulated, only 4 oblique ridges on each side, the
2nd a little way from its lower end produced into a strong tooth pointing
sharply outward and forward, eyelobe carrying a few small denticles, and
frontal lobe 2 short transverse denticulated ridges; telson longer than peduncle
of uropods, its post-anal section much the longer with about 10 pairs of lateral
spines. Antenna 1, 34 joint longer than 2™4, 1st shorter than the 2 combined.
Antenna 2 of ©, penultimate joint elongate. Maxilliped 3, 2"4 joint not
expanded distally, 34 wider than 4%. Peraeopod 1, 2™¢ joint longer than rest
of limb, 5, 6t, and 7 subequal. Peraeopod 2, 5* joint little longer than
6t and 7 combined. Exopod on 3¢ and 4 limbs of Q rudimentary,
2-jointed. Endopod of uropods very slightly longer than exopod, about */,ts
as long as the peduncle, 18 joint nearly twice 2°" and 3° combined, spines
of medial margin respectively 10, 2, 2. Colour red and yellow (Stuxberg).
L. about 12 mm, in O varying from less to much more.
At various stations of the ,Alaskan area, including Aleutian Islands, and the
Pacific coast of N. America north of Puget Sound“, from depths between 11 and 166 m.
2. D.dalliCalm. 1912 D.d., Calman in: P. U.S. Mus,, v.41 p.605, 635 tf.40—44.
In general like L). scorpioides (nr. 23), but surface of carapace much
smoother, ridges not crenulated, only 4 oblique ridges on each side, the
2nd not forming a prominent tooth, the pseudorostral lobes less produced than
in J). bidentatus; telson a little shorter than the peduncle of the uropods,
post-anal section much the longer, lateral spines about 11 pairs. Antenna 1,
32 joint longer than 2™4, 1st than the 2 combined. Antenna 2 of Q, penul-
timate joint elongate. Maxilliped 3, 2"¢ joint not distally expanded, 3° not
wider than 4%, Peraeopod 1, 2" joint much longer than rest of limb, 5™
a little shorter than 6 or 7 which are subequal. Peraeopod 2, 5 joint
a little longer than 6 and 7 combined. Exopod on 34 and 4 limbs of
© rudimentary, 2-jointed. Endopod of uropods very slightly longer than
the exopod, about half as long as the peduncle, 1S* joint much longer than
2nd and 3° combined, spines of medial margin respectively 9, 3,2. L. O 24 mm.
Alaskan area, depth 24—146 m; Sea of Okhotsk, 137 m.
3. D. planifrons Calm. 1912 D.p., Calman in: P. U.S. Mus., v.41 p. 605, 643
if. 58 —61.
Pseudorostral lobes well produced, horizontal, acute, antero-lateral corner
obtuse. Carapace with prominent oblique ridge on each side starting from
the pseudorostral lobes and forming a prominent blunt tooth at its junction
with a curved ridge running up from the lower edge; to the rear the oblique
ridges meet a pair of convergent longitudinal ridges, while in front a blunt
92 Cumacea: 11. Diastylidae, 1. Diastylis
median ridge runs between them. Pedigerous segment 5 with hind corners |
acute, little produced; telson about 7/,4 as long as peduncle of uropods, its
post-anal section the shorter, with 5 pairs of lateral spines. Eye with 3 corneal
lenses. Maxilliped 3, 24 joint much widened distally, produced into a small
lobe laterally, 4t® wider than 34, produced into a curved tooth laterally.
Peraeopod 1, 5% joint longer than 6%, Peraeopod 2, 5 joint nearly twice
as long as 6t and 7 combined. Exopod on 3¢ and 4 limbs of © rudi-
mentary, 2-jointed. Exopod of uropods half as long as the peduncle, a little
longer than the endopod, in which the 1%* joint is longer than the 34, the
3 than the 2™*, the 2 combined rather longer than the 1%‘, spines of the
medial margin respectively 6, 4, 4. L. Q 14 mm.
S. America, lat, 51° 34’ S., long. 68° W., 91 m; Straits of Magellan, 112 m.
4. D. alaskensis Calm. 1912 D.a., Calman in: P. U.S. Mus., v.41 p. 605, 641
tf. 51—57.
Pseudorostral lobes much and acutely produced. Carapace narrowing
forwards, more or less completely encircled by 6 transverse ridges in the Q,
the 6t obscure in the CG, the hind margin upturned. Eyelobe rather longer
than broad; hinder angles of 5‘* pedigerous segment blunt; pleon segments 1,
2, of S each with a medio-ventral spine; telson rather less than 7/,°5 as long
as peduncle of the uropods, post-anal section the longer, with about 6 pairs
of lateral spines. Antenna 1, 2"¢ joint shorter than 3° Antenna 2 of ©,
penultimate joint elongate. Maxilliped 3, 2" joint expanded distally, 3¢ narrower
than 4th, Peraeopod 1, 24 joint about as long as rest of limb, 6' longer
than 5‘ or 7%, Peraeopod 2, 5‘ joint much longer than 6% and 7™ combined.
Exopod on 3¢ and 4* limbs of Q minute, 2-jointed. Exopod of uropod a
little less than half as long as the peduncle, and slightly longer than the
endopod, which has the 1%* joint equal to 2™¢ and 3° combined, or less.
spines on medial margin respectively 4, 2, 2. L. © 135 mm, SG 10°6 mm,
Alaskan area, depth 9—110 m.
5. D. anderssoni Zimmer 1907 D. a, Zimmer in: Zool. Anz., v.31 p. 220 |
1909 D. a., Zimmer in: Ergeb. Schwed. Siidpol.-Exp., v.6 nr. 3 p.18 t.6 f. 97—105,
t.7 f. 106—118.
Pseudorostral lobes well produced in front of the eyelobe, which in ©
put not in G carries a pair of denticles, frontal lobe crossed by 2 ridges
in Q, 1 in G, these being followed by 10 more or less encircling ridges,
of which the first 1 and the last 2 are less developed than the rest; they
are finely denticulate, as is also the upturned hind margin. The upper hind
margin of the first 2 pleon segments is finely dentate; the telson almost as
long as the 5 pleon segment in Q, longer in the 3S, with post-anal section
much the longer, carrying 8 or 9 pairs of lateral spines in Q, about 17 im Ge
Eyelobe showing lenses but no pigment. Antenna 1, 1st joint longer than ~
gnd and 34 combined. Maxilliped 3, 2"¢ joint more than twice as long as
rest of limb, 34 much wider than base of 4. Peraeopod 1, 2"¢ joint not as
long as rest of limb, 5‘ rather shorter than 6‘, 6 much shorter than 7%,
Peraeopod 2, 6 joint about 2/,48 of 7%, both together about 2/08) Of noe
Exopods of 3¢ and 4 pairs minute 2-jointed in Q, strongly developed Ingo:
Setae on medial margin of peduncle in the pleopods clearly modified as
retinacula. Peduncle of uropods as long as 5 and 6 pleon segments
combined, with about 26 spines on medial margin in Q, many more in elie.
the rami rather more than a third as long, the exopod a little the shorter
Cumacea: 11. Diastylidae, 1. Diastylis 93
1st joint of endopod the longest, but not equal to 2"? and 3° combined,
spines on its medial margin 13, 8 on each of the others in the 3 5 DA, Ds
in the @. Colour white, with tinge of yellow. L. about 17 mm.
S. Georgia, depth 64—310 m.
6. D. hammoniae Zimmer. 1902 D.h., Zimmer in: Ergeb. Hamburg. Magalh.-
Sammelr., v.2 Cumac. p.6 tf. 8—10.
Pseudorostral lobes slightly upturned, short. Midway between their
apex and the small eyeless eyelobe start denticulate ridges diverging at a
right angle, then running for a space longitudinally and lastly bending
sharply forward; between them are a short feebly denticulate pair of median
carinae and outside them a submedian pair of curved folds; near the lower
margin runs on each side of the carapace a straight, nearly horizontal,
granular line. Pedigerous segments 3 and 4, wider than the 1** and ue
and each with a rather deep lateral furrow. Pleon segment 5 the longest,
6th the widest, a little shorter than the telson, the distal section of which
has about 12 pairs of lateral setiform spines. Peraeopod 3, 4, without rudi-
mentary exopods. Peduncle of uropods about twice as long as the telson,
with some 28 spines on the medial margin; rami rather shorter than the
telson, exopod slightly shorter than the endopod, of which the 1S* joint is
not greatly the longest, spines on the medial margin of the 3 joints respect-
ively 7, 5, 4. Colour clear yellowish white. L. 9 20 mm.
S. Atlantic, off Cape Blanco, E. Patagonia, 151m; lat. 45° 6’S., long. 600 W., 105m.
7. D. antillensis 0. Sars 1871 D. a.. G. O. Sars in: Ofv. Ak. Férh., v.28 nr.6
p. 803 | 1873 D.a., G. O. Sars in: Svenska Ak. Handl., n. ser. v.11 nr.5 p.4 t.1 £.1—3.
Form of © unusually slender. Pseudorostral lobes not far produced
in front of the elongate oval prominent eyelobe; surface of carapace antero-
dorsally densely beset with forward pointing spicules, and in the middle
obsoletely carinate, with a pair of finely crenulate pleats curving round from
the bases of the pseudorostral lobes to the middle of the back. Pedigerous
segments 3, 4, rather wider than the rest; pleon segment 5 much the longest,
6t2 the widest. Telson less than twice as long as the 6 segment, about
5/.ths of the peduncle of the uropods, with 5 pairs of lateral spines. Antenna J,
3° joint elongate. Peraeopod 1 very long, 6 and 7* joints very long,
subequal. Peraeopod 2, 5 joint subequal to 6" and 7 combined. 3° and
4th pairs without rudimentary exopods. Peduncle of uropods subequal to
the endopod, which is slightly longer than the exopod, its 18* joint equal to
2nd and 3° combined, 3° much longer than 2™¢, spines on the medial margin
respectively 6, 2, 4. Integument pellucid. L. 6°5 mm.
W. Indies, Anguilla, depth 377—565 m.
8. D. koreanus Calm. 1911 D. koreana, Calman in: Tr. zool. Soc. London,
v. 181v p. 342, 3882 t.37 f.19—25.
Pseudorostral lobes far and acutely produced in front of the minute eyeless
eyelobe. Surface of carapace setose with spicules interspersed, lower margin
minutely dentate anteriorly, the back with a median depression between the
branchial regions. Pedigerous segments moderately setose, hind corners of
5th rounded with a minute terminal spinule. First 5 pleon segments with
lateral corners produced and spiniform, and each with pair of small dorso-
lateral teeth; telson as long as 5% and 6% segments combined, the shorter
post-anal section having about 10 pairs of almost setiform lateral spines.
94 Cumacea: 11. Diastylidae, 1. Diastylis
Antenna 1, 18* and 3¢ joints subequal, 2"¢ shorter than either. Peraeopod 1,
2n¢ joint shorter than rest of limb, 6 joint longer than 5%, and nearly
twice the 7. Peraeopod 2, 6% and 7 joints together 7/,4 of the 5%.
3¢ and 4t pairs without rudimentary exopods. The 3¢ to 5* pairs with
tubercles produced at lateral end of 4" joint. Peduncle of uropods subequal
to telson, with a dozen spinules on the medial margin; exopod curved,
shorter than the endopod, which is about half as long as the peduncle, the
1st joint longer than 2°¢ and 3 combined, spines on the medial margin
respectively 6, 3, 2. L. 9 12 mm.
Korea, depth 68 m.
9. D. algoae Zimmer 1908 D.a., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 158,
188 t.44 f.96—98; t.45 f. 99—108 | 1910 D. a., T. Stebbing in: Ann. S. Afr. Mus.,
v. 61v p. 418.
General surface finely granular, integument hard, brittle. Pseudorostral
lobes produced much in advance of the eyeless eyelobe, horizontal, each
acute, apices separated by small emargination; carapace covered with micro-
scopic forward pointing denticles, frontal lobe with pair of longitudinal pit-
like depressions. Pedigerous segments 1—4 about as wide as the carapace,
3¢ and 4% dorsally distinguished only by suture line, 5 with sides acutely
extended along 1S* pleon segment; telson much longer than 5 pleon segment
with lateral setules on the proximal section, the narrow distal section nearly
twice as long with some 16 pairs of lateral spines. Antenna 1, 2"¢ joint of
peduncle the shortest, 2"¢ joint of principal flagellum the longest. Maxilliped 3,
3¢ joimt rather wider than 4%. Peraeopod 1 not very long, 2™¢ joint about
as long as rest of limb, 6" joint longer than 5%, 5% than 7%. Peraeopod 2,
5 joint rather longer than 6 and 7 combined. No exopods on the 3¢ and
4th pairs of the ©. Peduncle of uropods longer than the telson, with
21 spines on the medial margins exopod.rather more, endopod a little less,
than half the peduncle, 18* joint of endopod equal to 2"¢ and 3° combined,
spines on the medial margin respectively 6, 4, 3. Colour white. L. 9 9 mm.
S. Africa, Algoa-Bay, depth 40—55 m.
10. D. spinulosus Heller 1875 D. spinulosa, Cam. Heller in: Denk. Ak. Wien,
v.35 p.28 t.1 £.5 | 1879 D. nodosus, G. O. Sars in: Arch. Naturv. Kristian., v.4 p.472 |
1883 D. spinulosus, G. O. Sars in: Forh. Selsk. Christian., nr. 18 p.12, 57 | 1885 D. nodosa,
G. O. Sars in: Norske Nordhavs-Exp., v.6 nr. 14 p.61 t.7 f.1—4 | 1886 D.n. + D.
spinulosa, G. O. Sars in: Norske Nordhavs-Exp., v.6 nr. 15 p.22; 21 | 1887 D.s., H. J.
Hansen in: LDijmphna Udb., p.241 | 1887 D. s., H. J. Hansen in: Vid. Meddei., v.39
p. 205 | 1900 D. s., G. O. Sars, Crust. Norway, v.3 p.55,56 t.42 | 1900 D. s. + D. nodosa,
Pea in: Rennes arctica, v.1 p.425, 434 (with synonymy); p.423, 433 | 1901 D.s., Oblin
: Bih. Svenska Ak., v. 26 nr. 12 p.47 t.6 f. 10a—d.
Pseudorostral lobes well produced in front of eyeless eyelobe, horizontal,
acute, sides minutely denticulate; carapace large, with a pair of spines on front
of frontal lobe, behind which are two outward curving rows of 4 conspicuous
spines, while dorso-laterally large groups of forward pointing spines are very
conspicuous, these being nodose instead of sharp and somewhat less numerous
in young specimens. Pedigerous segments 1—4 are spiniferous, with a
specially conspicuous pair of spines dorsally, the 5% segment having an
unpaired median spine. Pleon segments coarsely denticulate, with postero-
lateral corners produced outwards; telson longer than the 2 preceding segments
combined and than the peduncle of the uropods, with about 8 pairs of lateral
spines. Antenna 1, 1S joint longer than 24 and 3¢ combined. Peraeopod 1
Cumacea: 11. Diastylidae, 1. Diastylis 95
of moderate length, 2™¢ joint spinulose, 5‘ shorter than 6%, 7 than 5t),
Peraeopod 2, 22¢ to 4% joints denticulate, 5 subequal to 6™ and 7 com-
bined. Exopod of uropod longer than endopod, which is half as long as
the peduncle, its 18* joint scarcely equal to 2.4 and 3¢ combined, spines on
the medial margin respectively 4, 2, 3. Colour pale carneous. L. Q 22°8 mm.
N. Norway, depth 188 m; Franz Joseph Land; Barents Sea, 226—414 m; Kara
Sea, 9—168 m; W. Greenland, 90—377 m; lat. 79° 58’ N., 435 m; lat. 81° 14’ N., 150m.
11. D.armatus Norm. 1876 D. armata (nom. nud.), A. M. Norman in: Jeffreys
in: P. R. Soe. London, v. 25 p. 212 | 1879 D.a., A. M. Norman in: Ann. nat. Hist., ser. 5
v.3 p.64 | 1887 D.a., H. J. Hansen in: Vid. Meddel., v.39 p.205 | 1900 D. a., Zimmer
in: Fauna arctica, v.1 p. 420, 434.
Pseudorostral lobes short, but ending acutely, with a pair of conspicuous
erect spines, behind each of these a longitudinal curved row of 4 spines, all
but the 18* very small, and a single spine between each row and the serrate
antero-inferior margin, the surface of the carapace otherwise glabrous, with
sparse shallow pittings. Pedigerous segments together scarcely half as long
as the carapace, smooth, except that the side-plates of the 5 are armed
with a spine, but not produced backward. Pleon segments smooth, telson
longer than the peduncle of the uropods, its proximal section longer than
the preceding segment, lateral spines 3 small pairs, apical spines long. An-
tenna 1, with the 3 joints of the peduncle subequal. Peraeopod 1, 24 joint
strongly spinose. Exopod of uropods rather longer than the endopod, the
medial margin of which has on its 3 joints respectively 2, 2, 1 spines.
i. © 10 mm.
Entrance of Davis Strait, lat. 59° 10’ N., long. 50° 25’ W., depth 3295 m.
12. D. bispinosus (Stimps.) 1853 Cuma bispinosa, Stimpson in: Smithson. Contr.,
v.6 nr.5 p.39 | 1871 Diastylis qvadrispinosa, G. O. Sars in: Ofv. Ak. Forh., v.28 nr. 1
p.72 | 1871 D. q., G. O. Sars in: Svenska Ak. Handl., n. ser. v9 nr.13 p.28 t.10
f, 50—52, t.11 f.53—61 | 1873 D. quadrispinosa, S. I. Smith in: A. E. Verrill in: Rep.
U.S. Fish. Comm., v.1 p.507, 554 t.3 £.13 | 1874 D. q., S. I. Smith in: Tr. Connect. Ac., v.31
p. 28 | 1879 D. quadrispinosus, S. I. Smith in: Tr. Connect. Ac., v.51 p.112 | 1883 D.
bispinosus, G. O. Sars in: Forh. Selsk. Christian., 1882 nr. 18 p. 57 | 1900 D. bispinosa
G. O. Sars, Crust. Norway, v.38 p.47 | 1900 D. b., Zimmer in: Fauna arctica, v.i p. 421,
434 | 1912 D. quadrispinosa, Calman in: P. U.S. Mus., v 41 p. 605, 658.
Pseudorostral lobes well produced, horizontal, acute. Carapace large,
with finely serrate anterior half of lower margin, a pair of conspicuous for-
ward pointing teeth, nodose in the do, followed by a much smaller pair, flanking
the frontal lobes, and the back especially on the branchial regions beset with
spicules and short hairs, the hind margin raised. Pedigerous segments smooth,
5th with front margin crenulate, and small median tooth behind, postero-
lateral corners more produced in dO than in g. Pleon segments 1 and 5
each with a pair of postero-dorsal teeth, a median serrate ridge on 24 to 5th
in the & and a pair of dorso-lateral ridges; telson shorter than peduncle
of uropods (Sars) or a little longer than it (in CG, Calman), with 12—14
pairs of lateral spines. Eye distinct. Antenna 1, 34 joint longer than 24,
1s¢ equal to both combined. Maxilliped 1, with branchial leaflets in spiral
arrangement. Peraeopod 1, 2"¢ joint shorter than rest of limb, 7" shorter
than 5%, 5 than 6%, Peraeopod 2, 2"¢ joint with strong spine at apex,
3¢ with 2 such spines, 5% shorter than 62 and 7 joints combined. The
many-spined peduncle of uropods rather less than twice the exopod, rather
more than twice the endopod, in which the 1* joimt is rather less than the
96 Cumacea: 11. Diastylidae, 1. Diastylis
2nd and 34 combined, spines on the 3 joints respectively 6, 5, 3 (Sars), for
2nd joint 3 or rarely 4 (Smith). Colour pale carneous, with reddish purple
patch on carapace, and 2 small spots of pink. L. Oo nearly 11 mm.
N. America, from Gulf of St. Lawrence to lat. 35° 42‘ N., near Cape Hatteras,
depth 4—373 m.
13. D. cornutus (Boeck) 1859 D. bispinosa, Danielssen (non Stimpson, 1853)
in: Norske Selsk. Skr. 19. Aarh., v.41 p.108 | 1864 Cuma cornuta, A. Boeck in:
Forh. Selsk. Christian., 1863 p.190 | 1865 Diastylis bicornis, Bate in: Ann. nat. Hist.,
ser.3 v.15 p.84 t.1 f.2 | 1865 D. bispinosa, G. O. Sars in: Forh. Selsk. Christian., 1864
p. 164 | 1865 D. cornuta, Bate in: Zool. Rec., 1864 v1 p.287 | 1866 D. bispinosus,
Bate in: Zool. Rec., 1865 v.2 p. 326 | 1879 D. cornuta, A. M. Norman in: Ann. nat.
Hist., ser.5 v.3 p.55 | 1883 D. cornutus, G. O. Sars in: Forh. Selsk. Christian., 1882 nr. 18
p. 12,58 | 1894 D.c., A.M. Norman in: Ann. nat. Hist., ser.6 v.13 p.276 | 1900 D. cornuta,
G. O. Sars, Crust. Norway, v.3 p.45 t.35, 36 | 1900 D. c., Zimmer in: Fauna arctica,
v. 1 p.421, 434 | 1909 D.c., A. M. Norman & G.S. Brady in: Nat. Hist. Tr. Northumb.,
ser. 2 vd p.3l.
Nearly allied to LY. bispinosus, but pseudorostral lobes less produced,
the antero-lateral corners distinct, though obtusely angled; the two pairs of
Fig. 55. Fig. 56. Fig. 57.
D. cornutus (>) (Boeck), D. cornutus (Boeck), D. cornutus (Boeck),
branchiae (after Sars). pleopod 1 (after Sars). pleopod 2 (after Sars).
teeth which flank the frontal lobe of the carapace are in the © more
conspicuous, and are followed by 2 meandering rows of small unequal
denticles; in the G the pseudorostral lobes are minutely serrate at each
side, the antero-lateral corners produced as rounded dentate lobes close to
the eye, an obliquely transverse fold in the front part of the carapace on
each side joins the serrate lateral carina which runs to the hind margin, the
carapace being otherwise almost smooth. Pedigerous segments with spinules
dorsally in double row in Q, not in d, 5% segment in © with a median
spinule on the hind margin, the postero-lateral angles acute, less produced
in © than in Gd. Pleon clothed with spinules. Telson shorter than peduncle
of uropods, with 8—10 pairs of lateral spines. Eye small, imperfectly deve-
loped. Antenna 1, 34 joint longer than 24, 1st not equal to 2"4 and 34
combined, margin denticulate. Peraeopod 1, with 5 joint nearly, 6" more
Cumacea: 11. Diastylidae, 1. Diastylis | 97
than, twice as long as 7%". Peraeopod 2, 5 joint longer than 6% and 7th
combined. Endopod of uropods in Q with spines on the 3 joints respectively
6, 2, x, in GD 16, 7, 7; terminal spine in © continuous with the apical
joint. Colour greyish white, in Q tinged with yellow. L. @ 12 mm, d
14 mm.
Coast of Norway, Kattegat, British Isles, coast of France, at various depths
between 27 and 549m, but in lat. 55° 40’ N., long. 12° 46’ W., at 2700 m.
14. D. laevis Norm. 1869 D.1., A. M. Norman in: Rep. Brit. Ass., v.38 p. 270 |
1871 D. tumida, G. O. Sars in: Svenska Ak. Handl.,, n. ser. v9 nr.13 p.5 | 1877 D. laevis,
Meinert in: Naturh. Tidsskr., ser. 3 v.11 p.176 | 1879 D.1., A.M.Norman in: Ann. nat.
Hist., ser.5 v.3 p.60 | 1883 D.1., D. rostrata?, G. O. Sars in: Forh. Selsk. Christian.,
1882 nr. 18 p.57,58 | 1897 D. 1, Ehrenbaum in: Wiss. Meeresunters., v.21 p.416 | 1900
D. rostrata, G. O. Sars (non Goodsir 1843), Crust. Norway, v.3 p.51 t.39 | 1906 D. laevis,
A. M. Norman & T. Scott, Crust. Devon Cornwall, p.31 | 1909 D.1., A. M. Norman & G.S.
Brady in: Nat. Hist. ‘I’'r. Northumb., ser.2 v.31 p. 32.
Near to YD. twmidus (nr. 25) and D. rathkii (nr. 16). Pseudorostral
lobes well produced, horizontal, ending acutely, antero-lateral corners rounded,
serrate. Carapace of QO smooth, with scattered hairs, in GO with smooth longi-
tudinal carina on each side, crossed in front by an obliquely transverse fold.
Pedigerous segments also smooth, except for serrate front margin of 5%
segment, of which the postero-lateral corners are acute, more produced in
G than in ©. Pleon segments spinulose only in G; telson nearly equal to
the 2 preceding segments combined, its distal portion much the longer, with
11—17 pairs of lateral spines. Eye well developed. Antenna 1, 3¢ joint
nearly as long as first, longer than the 2™4. Peraeopod 1, 24 joint densely
fringed with plumose setae on both margins with spines on surface be-
tween them; 5* joint about as much longer than 7 as it is shorter than
6%. Peraeopod 2, 5% joint subequal to 6% and 7t* combined; 34 and
4th pairs with 2°¢ joint not denticulate. Peduncle of uropods longer than
the telson, with as many as 23 spines on the medial margin; exopod
longer than endopod, which is about half as long as the peduncle, with
7 or 8. 3 or 4, 2 or 3 spines respectively on the medial margin of its 3
joints, the 1st longer than 2"¢ and 34 combined. Colour greyish white.
L. 10—11 mm.
S. coast of Norway, Danish, French, British waters, depth 9—37 m, and between
lat. 52° 25‘ and 54° 15’ N. in depths between 165 and 2980 m.
15. D. sulcatus Calm. 1912 D. suleata, Calman in: P. U.S. Mus,, v.41 p. 605,
654 tf. 76—78.
Near to D. rathki (nr. 16), but distinguished by sculpture and more
elongate form of the carapace. Pseudorostral lobes horizontal, acute; lower margin
of carapace anteriorly serrate; just behind the small eyeless eyelobe a carina,
there dentate, crosses the carapace and diverges downwards and backwards,
becoming denticulate below but not reaching the hind margin; sideplates of
2nd yedigerous segment narrowly rounded in front, lateral expansions of 34
and 4t2 more strongly produced backwards than in J). rathkii, postero-lateral
angles of 5% acutely produced. Pleon segment 5 slightly longer than the
6th, telson about 1/,¢ longer, its distal section long, narrow, with about 7
pairs of lateral spinules. Antenna 1, 1S¢ joint nearly thrice as long as 2"4,
and twice the 3°. Joints of antenna 2 short. Peraeopod 1, 2"¢ joint much
Das Tierreich. 39. Lief.: T. R. R. Stebbing, Cumacea. 7
98 Cumacea: 11. Diastylidae, 1. Diastylis
longer than rest of limb, joints 5—7 subequal. Peraeopod 2, 3¢ joint with
conspicuous spine on medial margin, 5 longer than 6%" and 7‘ combined.
34 and 4th pairs without exopods; but as in Diastylopsis (p. 109) 3¢ pair
distant from 2™4, its 5th joint as broad as the 4, only a little longer than
broad. Pedunecle of uropods scarcely longer than the telson; exopod less
than 2/,48, endopod about half as long, with 1st joint longer than 2°? and
34 combined, spines of medial margin respectively 5,3, 3. L. Q 13 mm, adult
3 unknown.
Alaska, lat. 63° 37‘ N., long. 165° 10’ W., 22 m.
16. D. rathkii (Kroyer) 1841 Cuma r., Kroyer in: Naturh. Tidsskr., v. 3v1 p. 513,
531 t.5 f.19—¥2; t.6 f.17, 18, 23—30 | 1843 Alauna rostrata, Goodsir in: Edinb. new
phil. J., v 34 p.121, 127 t.4 f.1—10 | 1846 Cuma rathkui + C. angulata 3, Kroyer in:
Naturh. Tidsskr., ser, 2 v.211 p. 144, 207 t.1 f.4,6; p.156, 206 t. 2 f. la—i | 1849 C.r. +
C.a., Kroyer in: Voy. Nord Crust., t.5 f.1; t.5 £2 | 1852 C. rathkei, W. Liljeborg in:
Ofv. Ak. Férh., v9 p.6 | 1856 Diastylis rathkii, Bate in: Ann. nat. Hist., ser.2 v.17
p. 451 (8) t.13 | 1867 Alawna goodsiri (non Bell), A.S. Packard in: Mem. Boston Soe.,
v. lit p.301 | 1869 Diastylis spinosa 3, A. M. Norman in: Rep. Brit. Ass.. v.38 p. 271
1873 D. rathkei, G. O. Sars in: Svenska Ak. Handl., n. ser. v.11 nr. 6 p.7, D. rathkia,
t.3 £.8,9 | 1873 D. orbignyi (2), Latreille MS. in: P. Fischer in: Act. Soc. Linn. Bordeaux,
ser. 3 v.8v p.425 | 1878 D. bimarginatusd, Bate in: Ann. nat. Hist., ser.5 v.1 p.409
1878 D. b., Sim in: Ann. nat. Hist.. ser.5 v2 p.453 | 1879 D. b., Bate in: Ann. nat-
Hist., ser.5 v3 p.93 | 1879 D. rathkii, A.M. Norman in: Ann. nat. Hist., ser.5 v3 p.54
1880 D. Rathkei, Stuxberg in: Bih. Svenska Ak,, v.5 nr. 22 p. 35 (distribution, numbers)
1883 Cuma rathkii, Burmester. Beitr. Cuma, p.1 t.1, 2 | 1885 C. r., H. Blanc in: Reeu.
zool. Suisse, v.2 p. 253 t.10 (development of egg) | 1900 Diastylis rathkei, G. O. Sars.
in: Crust. Norway, v.3 p.44 t.33, 34, D. r. var. dg, p.107 t. 70—72 | 1900 D. rathkii, Zimmer
in: Fauna arctica, v.1 p.423 (with extensive synonymy), 434, D. r. var. glabra, Zimmer,
p. 425 | 1901 D.7r., Ohlin in: Bih. Svenska Ak., v.26 nr. 12 p.46 | 1902 D. r. var. sarsi, A.M.
Norman in: Ann. nat. Hist., ser.7 v.10 p.478 | 1906 D. rathkeit, A. M. Norman & T. Scott,.
Crust. Devon Cornwall, p.30 | 1909 D. r., Stappers in: Cellule, v. 2511 p. 341, 358 t. 1
f.9; t.2 f. 1—18 (le tube digestif) | 1909 D.r.. A. M. Norman & G. S. Brady in: Nat.
Hist. Tr. Northumb., n. ser. 2 v.31 p. 29 | 1911 D. rathkii, Stappers in: Camp. arct.
Orléans, Crust. Malac. p.110 (distribution) | 1912 D.7., Calman in: P. U.S. Mus., v.41
p. 605, 652.
Variable with the variations ill defined. ©. Body slender. Pseudorostral
lobes horizontal, conical, antero-lateral corners distinct or almost obsolete,
serrate; frontal lobe of carapace with double row of forward pointing denticles
down the middle, sometimes crossed by 1, 2 or 3 imperfect transverse rows,
and flanked by scattered denticles, sometimes few, sometimes very numerous.
Pedigerous segment 5 serrate in front, postero-lateral angles acute, strongly pro-
duced straight backwards or slightly divergent. Pleon segments simple, smooth,
telson shorter than pedunele of the uropods but reaching its end or equal to it, the
narrow section elongate with about 15 or fewer pairs of lateral spinules, the apical
pair short. Eye small, distinct. Antenna 1, 18* joint with or without small
apical tooth, longer than 34, 34 than 2"¢, Antenna 2, the joints short,
successively shorter. Peraeopod 1, 2™4 joint scarcely longer than rest of limb,
with denticles on side of distal part, joints 5—7 subequal. Peraeopod 2,
7% joint much or not much longer than 6, 5% much longer than both
combined. 32 and 4*® pairs stout, setose, without exopods. Exopod of
uropods about half as long as the peduncle, longer than the endopod, of
which the 1st joint is longer than the other 2 combined, spines of the
medial margin respectively 6, 3, 2 or 3. Colour whitish with carneous tinge.
LL, mm.
Cumacea: 11. Diastylidae, 1. Diastylis 99
SG. Frontal lobe and adjacent parts of carapace without denticles,
except at commencement of the denticulate carina which on each side curves
to run round subparallel to the lower margin. First 5 pleon segments
with longitudinal rows of denticles, telson geniculate, with 12—16 pairs of
slender lateral spines. Endopod of uropods with 11, 5, 3 spines on the 3
joints respectively. L. 12°5—22 mm.
Coasts of N. Europe, West Greenland and Labrador, Atlantie coast S. of Labrador,
Alaskan area and Pacific coast north of Puget Sound; between lat. 81° and 50° 10‘ N.,
and in depths between 9 and 1222 m.
17. D. bradyi Norm. ?1844 Alauna rostrata, W. Thompson in: Ann. nat. Hist.,
v.13 p. 435 | 21856 A, +., W. Thompson, Nat. Hist. Ireland, v.4 p. 392 | 1879 Diastylis
bradyi, A. M. Norman in: Ann. nat. Hist., ser.5 v.3 p.59 | 1888 D.b., A. O. Walker
in: P. Liverp. biol. Soc., v.2 p.178 t.13 f.10, 11 { 1906 D. bradii, A. M. Norman & T. Scott,
Crust. Devon Cornwall, p.32 | 1909 D. b., A. M. Norman & G. 8S. Brady in: Nat. Hist.
Tr. Northumb., ser.2 v. 311 p. 30 t. 8, 9.
Near to D. rathki, but having the carapace of the O mapped into areas
by lines of small denticles, chiefly transverse; the front margin of the anterior
pedigerous segments minutely crenulate; the pleon segments, including the
6th, having a series of small spines on the lower lateral margin; peraeopods
with the 24 joint, especially in the 1%* pair, strongly spinose, 1S* joint of
exopods also spinose; 7 joint of 2"¢ pair (as figured) even shorter than the
6; endopod of uropods with ¥—11, 3 or 4, 3—6 spines on the 3 joints
respectively. L. 11—12 mm.
Lough Swilley, county Donegal, Ireland, 27 m; round England, in shallow water
to 5m,
18. D. goodsiri (Bell) 1855 Alauna g., T, Bell in: Belcher, Last arct, Voy., v2
p. 403 t. 34 f. 2,2a—q (not f.3) | 1859 A. plumosa, M. Sars in: Forh. Selsk. Christian.,
1858 p. 127 | 1861 Cuma gigantea, Danielssen in: Nyt Mag. Naturv., v.11 p.7 | 1871
Diastylis goodsiri, G. O. Sars in: Svenska Ak. Handl., n. ser. v.9 nr. 13 p.5 | 1880 D. g.,
Stuxberg in: Bih. Svenska Ak., v5 nr.22 p.21, 22, 24,51, 52, 53,59 | 1887 D. g.,
H. J. Hansen in: Dijmphna Udb., p.241 t. 22 f.5—5n; t. 23 f.1—1le (detailed description) |
1900 D. g., G. O. Sars, Crust. Norway, v.3 p.54 t.41 | 1900 D. g., Zimmer in: Fauna
arctica, v.1 p. 422 (with synonymy), 434 | 1911 D. g., Calman in: Tr. zool. Soc. London,
v. 181v p.383 | 1912 D.g., Calman in: P. U.S. Mus., v.41 p. 605, 662.
Body robust, hirsute. Pseudorostral lobes produced, conical, horizontal,
antero-lateral corners obsolete. Carapace inflated, surface without denticles
(Sars), with plenty (H. J. Hansen, Calman), with several furrows varying in
the 2 sexes. Pedigerous segments furrowed down the middle in the d, 34
and 4t in © dorsally coalesced, postero-lateral angles of the 5 acute, more
produced in G than in g. Pleon segments 1—5 with postero-lateral corners
acute, produced backwards and strongly outwards, and in the G these segments
have medio-dorsal and lateral rows of denticles; telson in © as long as the
2 preceding segments combined, in GO a little longer, with 20 pairs of lateral
setiform spinules. Hyelobe small, eye not perceived. Antenna 1, 18 joint
longer than 224 and 3¢ combined, 2°¢ longer than 3°, principal flagellum
4-jointed, accessory small, 3-jointed. Antenna 2 of G with very long flagellum.
Peraeopod 1 not very long, the plumose 2?¢ joint nearly as long as rest of
limb, the last 3 joints not very unequal. Peraeopod 2, 2"4 joint denticulate,
5% jonger than 6 and 7 combined, 7 as figured not longer than 6t, 34
q*
100 Cumacea: 11. Diastylidae, 1. Diastylis
and 4th pairs stoutly built, without exopods in Q. Peduncle of uropods
longer than the telson, exopod scarcely half as long as the peduncle, very
little longer than the endopod, of which the 1S* joint is nearly double the
ond and 34 combined, with 9 spines on the medial margin, the 2™¢ having 2,
the 34 none and its apical spine not separate from the joint. Colour whitish.
L. 25—35 mm.
Coasts of N. Europe, Atlantic coast S. of Labrador, Polar Islands of N. America,
Greenland, Jan Mayen, Spitzbergen, Barents Sea, Kara Sea, Siberian Polar Sea, southward
reaching lat. 449 U5‘ 30’ N., long. 63° 31’ 30 W., depths 2—377 m.
19. D. stygius O. Sars 1871 D. stygia, G. O. Sars in: Ofy. Ak. Forh., v.28 or. 6
p. 798 | 1873 D.s., G.O. Sars in: Svenska Ak. Handl., n. ser. v.11 nr.6 p.6 t.2 f.4—7 |
1879 D. s., A. M. Norman in: Ann. nat. Hist., ser.5 v.3 p.57, 58 | 1886 IDs. GeO:
Sars in: Norske Nordhavs-Exp., v.6 nr. 15 p.22 | 1887 D.s., G. O. Sars in: Rep. Voy.
Challenger, v.19 pars 55 p. 44 t.6—8 | 1900 D. s., Zimmer in: Fauna arctica, v. 1 p. 425,
434 | 1901 D. polaris, Oblin in: Bih. Svenska Ak., v.26 nr. 12 p.50 | 1912 4). stygia,
Calman in: P. U. S. Mus., v.41 p. 605, 661.
Very near to D. polaris (nr. 20). Pseudorostral lobes conically pro-
duced, horizontal, antero-lateral corners obsolete; carapace with lower margin
serrate anteriorly, surface in Q beset with small spinules, densest in front,
but only scantily represented in the G, which on the other hand has a
pair of anterior lobes with serrate edges and a lateral keel low down on each
branchial region. Pedigerous segments 1—3 and 5 have the front margin
finely denticulate, 2—4 in the ¢ are firmly connected, 1—3 have a medio-
ventral ridge, postero-lateral angles of the 5' acutely produced and carrying
4 plumose setae in the G, but not acute in the ©, which has 2 small dorsal
denticles, while in the GO it has a ventral serrate crest, which is continued
on the first 2 pleon segments, pleon segments 1—6 otherwise ‘smooth in
both sexes, or 18t in © with a ventral spine (A. M. Norman); telson about
twice as long as 6% pleon segment with 9 pairs of lateral spinules in Q,
thrice as long, with 15 pairs of spinules in the G. Eyelobe small, eye wanting.
Antenna 1 ©, 1St joint much longer than 2"¢ and 3¢ combined, 2"¢ a little
shorter than 34. Antenna 2 d, flagellum very long. Peraeopod 1, 2"¢ joint
spinose and setose, not so long as rest of limb, 5% to 7% joints subequal.
Peraeopod 2, 34 joint with a strong denticle, 4% with 2 such denticles, fie
joint a little longer than 6%, 5% much longer than both combined. 3° and
4th pairs without exopods in the Q. Peduncle of uropods rather longer than
the telson, not quite twice as long as the subequal rami; 1%* joint of endopod
longer than the 24 and 34 combined, spines on the 3 joints respectively 8,
4,23 in (@, 13,19, Dime G. VL. 0) 16 mm) 24 Form:
Atlantic coast of N. America south of Labrador, depth at a single station 267 m,
otherwise only at depths varying between 1134 and 4896 m (or at most 3200, Ohlin).
20. D. polaris 0. Sars 1871 D.p., G. O. Sars in: Ofv. Ak. Férh., v.28 nr. 6
p- 797 | 1873 D. p., G. O. Sars in: Svenska Ak. Handl., n. ser. v.11 or.6 p.4 t.1 f.1—3 |
1879 D. p., A. M. Norman in: Ann. nat. Hist., ser.5 v3 p.56 | 1886 D. p., G. O. Sars
in: Norske Nordhavs-Exp., v.6 nr.15 p.22 | 1900 D. p., Zimmer in: Fauna arctica, v. 1
p. 423, 434 | 1901 D. p., Oblin in: Bih. Svenska Ak., v.26 nr.12 p. 50.
Pseudorostral lobes shortly conical, horizontal, antero-lateral corners obso-
lete; dorsal line of carapace nearly straight, surface beset with very small
forward pointing denticles, sides having 1 or 3 (G. O. Sars) or 2 (A. M.
Norman) very indistinct obliquely transverse pleats or wrinkles, lateral margin
Cumacea: 11. Diastylidae, 1. Diastvlis 101
serrate anteriorly, hind margin raised. Pedigerous segments all free, I, 2,
and 5 with crenulate front margin, postero-lateral corners of the 5%" bluntly
rounded. Pleon segments 1—6 without denticles, with short hairs, telson
narrow, twice as long as the 6 segment, with 10 or 11 pairs of lateral
spinules. Eyelobe very small, eye wanting. Antenna | with elongate peduncle.
Peraeopod 1, 24 joint not as long as rest of limb, last 3 joints subequal.
Peraeopod 2, 7% joint longer than 6, 5% subequal to both combined.
Pedunele of uropod very little longer than the telson, less than twice the
exopod, rather more than twice the endopod, of which the 1S’ joint equals
the 2"4 and 34 combined, with 5 or 6, 2 or 3, 3 spines on the medial margin of
the joints respectively. L. @ 8—12 mm, dO 12 mm (Ohlin, who unites //.
stygius With this species).
Lat. 60° 31/ N., long. 9° 18 W., depth 419 m; NW. of Spitzbergen, depth 1789 m;
eold area between Iceland and N. Norway, depth. 2510 m.
21. D. lucifer (Kroyer) 1841 Cuma lucifera, Kroyer in: Naturh. Tidsskr., v. 3v1
p. 527, 581 t.6 f.34, 35 | 1846 C.l., Kroyer in: Naturh. Tidsskr., ser. 2 v. 2m p.171, 207 |
1849 OC. 1, Kroyer in: Voy. Nord Crust., t. 3a—k | 1859 Diastylis l., Danielssen in: Norske
Selsk. Skr. 19. Aarh., v. 411 p.108 | 1865 D. borealis, Bate in: Ann. nat. Hist., ser. 3
v. 15 p. 85 t. 1 £.3 | 1871 D. lucifera, G. O. Sars in: Svenska Ak. Handl., n. ser. v.9 nr. 13
p.5 | 1879 D. luciferus, S. 1. Smith in: Tr. Connect. Ac., v.51 p.112 | 1894 D. lucifer,
A. M. Norman in: Ann. nat. Hist., ser. 6 v.13 p.276 | 1900 D. lucifera, G. O. Sars, Crust.
Norway, v.3 p.49 t.38 | 1900 D. 1, Zimmer in: Fauna arctica, v.1 p.422 (with synonymy),
434 | 1906 D.1L, A. M. Norman & T. Scott, Crust. Devon Cornwall, p.32 | 1909 D. L.,
A. M. Norman & G. S. Brady in: Nat. Hist. Tr. Northumb., ser.2 v.31 p.32 | 1912
D.1., Calman in: P. U.S. Mus., v.41 p. 605, 662.
Pseudorostral lobes conical, horizontal, antero-lateral corners almost
obsolete; carapace of Q with denticles in transverse rows on the frontal lobe
and less densely to the rear, in the G only 3 denticles, 2 flanking the frontal
lobe, 1 as in the Q on the eyelobe. Pedigerous segment 5 with postero-
lateral angles acutely produced only in the dG. Pleon segments 1—6 unarmed,
telson in © twice as long as the 6% segment with 3 or 4 pairs of lateral
spines, longer and narrower in the G. Hye small but distinct. Antenna 1,
34 joint a little shorter than 1st, longer than 2°4. Peraeopod 1, 2" joint
not as long as rest of limb, last 3 joints subequal. Peraeopod 2, 5 joint
little longer than 7‘, which is decidedly longer than 6%. 34 and 4" pairs
with 2"4 joint not denticulate, without exopods in ©. Peduncle of uropods
longer than the rami, exopod shorter than telson in Q, subequal to it in C,
in both sexes longer than the endopod, of which the 1st joint is not much
longer than the 2™4, spines on medial margin of its joints respectively 3, 2,
2 in O, ¥, 7, 6 in SG. Colour greyish white with olive tinge in ¢, whitish
semipellucid in dG. L. scarcely 6 mm 9, 8 mm OC.
Coasts of N. Europe, Atlantic coast of N. America S. of Labrador; depths from
15—388 m.
22. D. rugosoides A. Walker 1896 D,r., A.O. Walker in: Rep. Brit. Ass.,
v.66 p.459, and in: Herdman in: P. Liverp. biol. Soe., v.40 Tr. p. 45.
Very near D. rugosus (nr. 24), but distinguished by not having the
vertical pleat on the carapace, and by strong dorso-lateral teeth on pleon
segments 1—3.
Ireland, 6 miles off Galley Head, county Cork.
102 Cumacea: 11. Diastylidae, 1, Diastylis
23. D. scorpioides (Lepech.) 1780 Oniscus s., Lepechin in: Acta Ac. Petrop.,
1778 v.1 p. 248 t.8 f.2 | 1841 Cuma edwardsii, Kroyer in: Naturh. Tidsskr., v.3v1 p.504,
531 t.5 f. 1—16 | 1846 C.e. + C. brevirostris 3, Kroyer in: Naturh. Tidsskr., ser. 2 v. 211
p. 128, 207 t.1 f.1,3,5,9—14; p.174, 208 t.2 f.6 | 1849 C.e.+ C.b., Kroyer in: Voy.
Nord Crust., t.4; t.5 A f.a—t | 1871 Diastylis e., G.O.Sars in: Svenska Ak. Handl., n. ser.
v.9 nr. 13 p.5 | 1879 D.e., A. M. Norman in: Ann. nat. Hist., ser.5 0.3 p.61 | 1880
D. scorpioides, Stuxberg in: Bih. Svenska Ak., v5 nr. 22 p. 21, 22,52 | 1900 D.s., G. O.
Sars, Crust. Norway, v.3 p.58 t.44| 1900 D. s., Zimmer in: Fauna aretica, v.1 p. 424
(with synonymy), 433 | 1901 D. edwardsii, Ohblin in: Bih. Svenska Ak., v.26 nr. 12 p.52 |
1911 D. scorpioides, Stappers in: Camp. arct. Orléans, Crust. Malac. p.112 t.4 f.2—6,
11,12 | 1912 D.s., Calman in: P. U.S. Mus., v.41 p. 605, 684.
Pseudorostral lobes not greatly produced, conical,
horizontal, shorter and more obtuse in GC than in ©,
antero-lateral corners faintly marked; carapace sculp-
tured with 5 pairs of crenulate oblique transverse pleats
or ridges, and the hind margin also raised and cre-
nulate, pedigerous segments with front margin more
or less distinctly raised and crenulate, postero-lateral
corners of 5t® obtuse in Q, somewhat acutely pro-
duced in dG. The non-ovigerous Q having medio-
ventral processes (hyposphenes of Gilson) on segments
2—4, and a dorsal tooth on the 5 (Stappers), pleon
segments 1—6 unarmed, telson shorter than peduncle
of uropods, with about 8 pairs of lateral spinules.
Eye distinct. Antenna 1, 18* joint subequal to 24
shorter than usual in this genus. Peraeopod 1, 2"¢
joint shorter than rest of limb, plumose, distally
spinulose, 7* joint shorter than 5% or 6%, Peraeo-
pod 2, 5t joint decidedly longer than 6% and 7‘
combined, 7 longer than 6%, 34 and 4 pairs
without exopods in the ©. LExopod of uropods
about half as long as the peduncle, endopod con-
siderably shorter, its 1st joint a little longer than
2nd and 34 combined, spines on medial margin of
the joints respectively, 4—7, 2 or 3, 2in Q, 7 or8,
4, 30or4inc¢. Colour reddish brown. L. Q 10—20,
3S 11—13 mm.
West Greenland and Labrador, Atlantic coast of
N. America South of Labrador, off Newfoundland, lat. 47°
40’ N., long. 47° 35’ 30” W., depth 377 m, but in Franz
Joseph Fjord, lat. 78° 6‘ N., long. 27° 17 W., depths between
3—9 m.
Fig. 58.
D. scorpioides () (Lepechin)
(after Sars).
24. D. rugosus O. Sars 1865 D. rugosa, G. O. Sars in: Forh. Selsk. Christian.,
1864 p.166 | 1866 D. r. G. O. Sars in: Nyt Mag. Naturv., v.15 p.104 | 1869
D.r., G. O. Sars in: Nyt Mag. Naturv., v.16 p. 342 | 1872 D. r., G. O. Sars in: Forh.
Selsk. Christian., 1871 p.269 | 1879 D. strigata g, A. M. Norman in: Ann. nat. Hist.,
ser.5 v.3 p.62 | 1879 D. rugosa, G. O. Sars in: Arch. Naturv. Kristian., 0.4 p.46 t.34—38 |
1883 D. rugosus, G. O. Sars in: Forh. Selsk. Christian., 1882 nr, 18 p.12 | 1894 D. r.,
A. M. Norman in: Ann. nat. Hist., ser.6 v.13 p.277 | 1897 D. rugosa, Ehrenbaum in:
Wiss. Meeresunters., v.21 p.416 | 1900 D. r., G. O. Sars, Crust. Norway, v.3 p. 48 t. 37 |
1905 D.r., Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p.45 | 1906 D.r., A. M.
and 32 combined. Antenna 2 of d, flagellum much,
Cumacea: 11. Diastylidae, 1. Diastylis 103
Norman & T. Seott, Crust. Devon Cornwall, p.32 | 1909 D.r., A. M. Norman & G.S.
Brady in: Nat. Hist. Tr. Northumb., ser. 2 v.31 p. 32.
Pseudorostral lobes conically produced, horizontal, antero-lateral corners
slightly marked in Q, rounded and serrate in J; carapace of © with 5 pairs
of transverse pleats, some rather indistinct and discontinuous, the frontal lobe
flanked by a conspicuous pair of forward pointing teeth and having to the
rear a smaller more approximate pair; carapace of O with 2 pairs of distinct
transverse pleats flanking the frontal lobe, the longitudinal lateral carinae
quite smooth. Pedigerous segment 5 with front margin crenulate, postero-
lateral angles in Q blunt, acutely produced in G. Pleon segments 1—6 smooth
in ©. in the & 18* with ventral crest of 3 spines, 1s and 2”¢ each with pair
of postero-dorsal denticles, 3¢ to 6 with postero-dorsal denticulate keel and a
pair of similar latero-ventral keels; telson considerably shorter than peduncle
of uropods, with 6—9 pairs of lateral spinules. Eye purple under white,
with 3 lenses, larger in G than in ©. Antenna 1 Q long, 3¢ joint longer
than 1s, 1st longer than 2"4, flagella well developed. Antenna 2 Q, all the
joints small, flagellum in G very elongate. Peraeopod 1, 2nd joint shorter
than rest of limb, 6t longer than 5", more than twice as long as 7%.
Peraeopod ¥, 5th joint longer than 6‘ and 7% combined, 7 longer than 6.
3¢ and 4th pairs without exopods in ©. Exopod of uropods shorter than the
telson, more than half as long as the peduncle, endopod in © less than half
the pedunele, nearly half in d, with 6, 3, 3 spines on medial margin of
the joints respectively in Q, 16, 8, 5, or more in G, 1** joint in both
sexes subequal to 24 and 3¢ combined. Colour grey-white. L. Q 8 mm,
3 9mm.
From Norway, Trondhjem Fjord, round to Messina in the Mediterranean, depths
surface to 73 m.
25. D. tumidus (Lilj.) 1855 Cuma tumida, W. Liljeborg in: Ofv. Ak. Forh.,
v.12 p.119 | 1859 Diastylis t., Danielssen in: Norske Selsk. Skr. 19. Aarh., v. 411
p. 108 | 1872 D.t., G. O. Sars in: Forh. Selsk. Christian., 1871 p. 269 | 1883 D. tumidus,
G. O. Sars in: Forh. Selsk. Christian., 1882 nr.18 p.12 | 1894 D.¢., A.M. Norman in:
Ann. nat. Hist., ser.6 v.13 p.276 | 1900 D. tumida, G. O. Sars, Crust. Norway, v.3 p.52
t.40 | 1900 D. t., Zimmer in: Fauna arctica, v1 p. 425 (with synonymy), 434 | 1909
D.t., A. M. Norman & G. S. Brady in: Nat. Hist. Tr. Northumb., ser.2 v.31 p. 32 |
1912 D.t., Calman in: P. U.S. Mus., v.41 p. 605.
Pseudorostral lobes not strongly produced, slightly upturned, antero-
lateral corners fairly marked, denticulate. Carapace tumid, in © nearly as
broad as long, smooth, in o& with slight longitudinal keel low down on
each side; pedigerous segment 5 with postero-lateral corners obtuse in ©,
acutely produced in G; pleon segments 1—6 unarmed in Q, with scattered
denticles in G; telson shorter than the peduncle of the uropods, the distal
half armed with about 9 pairs of lateral spinules rather smaller than the
apical pair. Eye prominent, showing in G5 lenses. Antenna 1 Q, 1st joint
shorter than 24 and 34 combined, 34 rather longer than 2°47. Antenna 2 C,
flagellum very long. Peraeopod 1, 2"4 joint shorter than rest of limb, 5th shorter
than 6%, 7t scarcely half as long as 6%. Peraeopod 2, with 2 short teeth
on 34 joint, 5t as long as 6 and 7‘ combined, 7% considerably longer
than 6%, 34 and 4th pairs in Q without exopods. Peduncle of uropods
rather less than twice the exopod in Q, rather more in G; exopod rather
longer than endopod exclusive of the coalesced apical spine, spines on medial
104 Cumacea: 11. Diastylidae, 1. Diastylis
margin of endopod’s 3 joints respectively 7, 3, 2, 18¢ joint subequal to 2>¢
and 3¢ combined. Colour whitish with reddish brown tinge. L. O 9 mm,
G 10 mm.
Coasts of N. Europe, from Norway, Lofoten Isles to N. E. England; depths 37
to 942 m.
26. D. neapolitanus O. Sars 1879 D. neapolitana, G. O. Sars in: Arch. Natury.
Kristian., v.4 p.57 t. 39.
Near to J). rugosus (nv. 24), but much smaller and without prominent
processes. Pseudorostral lobes rather long, horizontal, conically pointed, antero-
lateral corners obsolete. Carapace without prominent ridges or dentate processes.
but rough especially to the rear with crowded spinules. Pedigerous segment
5 with front margin not crenulate, postero-lateral angles obtuse; telson rather
shorter than peduncle of uropods, with seven pairs of lateral spinules, and
a rather larger apical pair. Eye distinct, not very prominent. Hxopod of
uropods about 3/4%s as long as the peduncle, longer than the endopod, which has.
5, 2, 2, spines on medial margin of its joints respectively, the 1S* joint
equal to 2.4 and 34 combined without the coalesced apical spine. L. ovigerous
oO 5 mm.
Bay of Naples, depth about 94 m.
27. D. echinatus Bate 1865 D.e., Bate in: Ann. nat. Hist., ser.3 v.15 p. 81
t.1 f.1 | 1866 D. echinata, G. O. Sars in: Nyt Mag. Naturv., v.15 p.105 | 1869 D.e,
G. O. Sars in: Nyt Mag. Naturv., v.16 p.343 | 1872 D.e., G. O. Sars.in: Forh. Selsk.
Christian., 1871 p.269 | 1879 D.e., A. M. Norman in: Ann. nat. Hist., ser.5 v.3 p. 56 |
1883 D. echinatus, G. O. Sars in: Forh. Selsk. Christian., 1882 nr. 18 p. 12 | 1894 D. e.,
A.M. Norman in: Ann. nat. Hist., ser.6 v.13 p.276 | 1900 D. echinata, G. O. Sars,
Crust. Norway, v.3 p.57 t.43 | 1900 D. e., Zimmer in: Fauna arctica, v.1 p.421
(synonymy, part.), 434 | 1905 D.e. juv., Calman in: Sci. Invest. Fish. Ireland, 1904
app. 1 p. 44.
Pseudorostral lobes of moderate size, conically produced, horizontal,
laterally spinulose, antero-lateral corners obsolete in Q, rounded in CG, denti-
culate. Carapace in © tumid, nearly as broad as long, with 3 or 4 con-
spicuous spines extending forwards on each side to base of pseudorostrum,
the sides mapped into polygonal areas by rows of denticles, usually with
the angles accentuated by a larger spinule; in the more slender CG the armature
less pronounced, but with addition as usual of lateral carinae. Pedigerous.
segments each with dorsal pair of teeth, conspicuous in ©, postero-lateral
angles in segments 3—5 of © dentiform. Pleon segments 1—6 dorsally
and laterally and in G also to some extent ventrally denticulate; telson as
long as the peduncle of the uropods, with 3 pairs of lateral spinules, and
a somewhat larger apical pair. Eye inconspicuous. Antenna 1 of ©. 1%t joint
as long as 2"4 and 34 combined, 3° slightly longer than 24. Antenna 2 of d,
flagellum very elongate. Peraeopod 1, 2"¢ joint as long as rest of limb,
the margins in this and more or less in all the pairs spinulose, the 6" joint
a little shorter than the 5, scarcely longer than the 7. Peraeopod 2,
34 joint produced into a conspicuous tooth, 5t much longer than 6 and
7% combined, 7* longer than 6, 34 and 4 pairs of © without exopods.
Exopod of uropods longer than endopod, which is half as long as the peduncle,
with 3, 2, 1 spines on its joints respectively, the 2™¢ joint longer than the 34,
Cumacea: 11. Diastylidae, 1. Diastylis 105
the 18t than both together; in non-ovigerous females the endopod 2-jointed
(Calman). Colour whitish. L. © 10 mm, ¢ 11 mm.
'N. Norway, Faré Channel, Shetland Isles, off Galway, Ireland; depths between
183 and 1006 m.
28. D. tricinctus (Zimmer) 1903 Leptostylis tricincta, Zimmer in: Zool,
Jabrb., Syst. 118 p.691 tf. CC—EE | 1908 Diastylis ¢., Zimmer in: Ergeb. Tiefsee-
Exp., v.8 p. 183.
Pseudorostral lobes not strongly produced, horizontal, antero-lateral
corners obsolete. Carapace narrow, encircled by 3 finely denticulate pleats,
the foremost curving round the frontal lobe to the rear, the second nearer
to the 34 than the 18t. Pedigerous segment 5 with postero-lateral angles
very acutely produced and each surmounted by a row of plumose setae.
Pleon segments 1—4 with circle of postero-ventral spines; telson rather longer
than 5 pleon segment, about 2/348 of peduncle of uropods, with 8 pairs of
lateral spines and a longer apical pair. Antenna 2, flagellum moderately
long. Exopod of uropods slightly shorter than the telson, endopod about half
as long as the peduncle, with 7, 6, 13 spines on medial margin of its joints
respectively, those on the 34 joint small, 1s‘ joint rather longer than 3°
34 than 2™4, Colour clear brown. L. Gc about 6 mm, Q unknown.
Shinagawa (Japan).
Fig. 60.
D. nucella (2) Calman,
carapace and pedigerous
D. tricinctus (¢) (Zimmer). segments.
Fig. 59.
29. D. nucella Calm. 1912 D.n., Calman in: P. U. S. Mus., v.41 p. 605, 645
tf. 62—65.
Pseudorostral lobes short. horizontal, antero-lateral corners almost obso-
lete. Carapace with height, width, and length subequal, eyelobe wanting,
surface granular, dorsally diversified with 3 longitudinal pairs of series of
minute square-cut teeth, lower margin in front similarly serrate. Pedigerous
segment 5 with postero-lateral corners obtuse. Telson scarcely longer than
the preceding segment, half as long as the peduncle of the uropods, with.
lateral spinules variable, not exceeding 3 on either side, the apical pair longer.
Antenna J, 34 joint longer than 1st or 2.2. Antenna 2, penultimate joint
106 Cumacea: 11. Diastylidae, 1. Diastylis
not elongate. Maxilliped 3, 24 joint not expanded or produced distally.
Peraeopod 1, 2"4 joint much bent, not as long as rest of limb, 5th joint
slightly longer than 7%, 6 than 5%, Peraeopod 2, 5% joint subequal to
6 and 7 combined, 7 longer than 6%, 34 and 4% pairs without exo-
pods. Exopod of uropods a little shorter than the telson, a little longer
than the endopod, of which the 1** joint is about twice the 2™¢ or 34, with
3, 1. 1 spines on the medial margin of the joints respectively. L. O 9°5 mm,
3 unknown.
Cape Smyth, Alaska, depth 5 m.
30. D. anomalus Bonnier 1896 D. anomala, J. Bonnier in: Ann. Univ. Lyon,
v.26 p.555 t. 30 f.4a—p.
Pseudorostral lobes well produced, each with subapical spinule, antero-
lateral corners obsolete. Carapace with height nearly equal to breadth, eyeless
eyelobe well developed, surface diversified with numerous denticles small
and large, 3 strong ones on the medio-dorsal line, a pair set back on the
Fig. 61.
D. anomalus (2) Bonnier.
pseudorostral lobes and 3 other pairs on the sides of the carapace, lower
margin with anterior half serrate. Pedigerous segments covered, especially
on the medio-dorsal line, with prominent strong spines, the 5 with 3 dorsal,
2 lateral, and 1 medio-ventral. Pleon segments 1—4 also armed with strong
spines. 62 with 3 pairs; telson with 3 pairs lateral and a stronger apical pair.
Antenna 1 short, both flagella 3-jointed. Antenna 2 of 4 short joints. Mauxilli-
ped 3, 24 joint distally expanded. Peraeopod 1, 2"¢ joint spinose, much shorter
than rest of limb, 5” a little shorter than 6%, a little longer than 7‘.
Peraeopod 2, 284 and 34 joints each armed with 2 strong spines, 7th joint
only a little longer than the 6%, 34 and 4*® pairs without exopods. Peraeo-
pod 5 entirely wanting, but this may be a juvenile character (see Leptostylis
mancus p. 123). Peduncle of uropods rather longer than the telson, rather
less than twice the exopod, rather more than twice the endopod, of which
the 1st joint is a little longer than the 34, the 34 than the 24, with spines
on the medial margin respectively 2, 1, 1. L. Q juv. scarcely 9 mm.
Bay of Biscay, depth 950 m.
31. D. asper Calm. - 1912 D. aspera, Calman in: P. U.S. Mus., v.41 p. 605, 6417
tf. 66—69.
Cumacea: 11. Diastylidae, 1. Diastylis, 2. Brachydiastylis 107
Pseudorostral lobes moderately long, horizontal, acute, antero-lateral
angles barely indicated. Carapace beset with spinules, larger in front, and
lining the more or less prominent ridges, of which one pair, nearly hori-
zontal, adjoins the pseudorostral lobes and is met by a curved pair from the
lower margins; to these are added an oblique pair and a medio-dorsal longi-
tudinal pair. The eyelobe is broader than long, with obscure corneal areas.
Pedigerous segments 1, 2, 5 have the anterior margin finely serrate; postero-
lateral corners of the 5t* obtuse. Pleon segments 1 and 5 each with a pair
of small dorso-lateral teeth. Telson about twice as long as the 6" segment,
with about 9 pairs of lateral spinules. Antenna 1, 3¢ joint rather longer
than 224, Maxilliped 3, 2"¢ joint distally wider than 3° but not produced.
Peraeopod 1, 2"4 joint spinose, not quite so long as rest of limb, 5* joint
a little shorter than 6 and longer than 7%. Peraeopod 2, pen joint subequal
to 6 and 7% combined. 34 and 4 pairs in © without exopods. Peduncle of
uropods about as long as the telson in Q, shorter in young 3; rami unknown.
L. ovigerous 9 12°2 mm, GC juv. 9mm; Q with rudimentary oostegites, flatter
front part of carapace, body plunosely hirsute carapace, 15°6 mm.
Shahafka Cove, Kodiak, Alaska; hirsute variety, Okhotsk Sea, E. of Sakhalin
Island, depth 137 m.
D. ambigua Lillj. 1886 D. a. (deser. nulla), W. Lilljeborg in: C.W.S. Aurivillius
in: Bih. Svenska Ak., v.11 nr. 4 p.4l.
L, 12 mm.
From N. E. of Troms6 or West Finmark.
D. quadri-plicata Lillj. 1886 D. q.-p. (deser. nulla), W. Lilljeborg in: C. W.
S. Aurivillius in: Bih. Svenska Ak., v.11 nr.4 p. 41.
L. 12 mm.
From N. £. Tromso or West Finmark; at Baddern, between depths 75 and 94m,
2. Gen. Brachydiastylis Stebbing
1912 Brachydiastylis, T. Stebbing in: Ann. S, Afr, Mus., v. 10v p. 176.
Pseudorostral lobes upturned in the female and the carapace short,
the third and fourth pedigerous segments tending to coalesce, and the third
peraeopod widely separated from the second in the ovigerous female.
Antenna 1 with third joint of peduncle the longest. Second joint of third
maxilliped distally expanded, third not wider than fourth. Third and fourth
peraeopods of female without rudimentary exopods. Exopod of uropods much
longer than endopod.
1 species.
1. B. resimus (Kroyer) 1846 Cuma resima, Kroyer in: Naturhist. Tidsskr., ser. 2
v. 211 p. 165, 206, 211.t. 2 f.2a,b | 1849 C. r., Kroyer in: Voy. Nord Crust., t.3 f. la—p |
1871 Diastylis r., G. O. Sars in: Svenska Ak. Handl., n. ser. v.9 nr. 13 p.5 | 1880
D.r., Stuxberg in: Bih. Svenska Ak., v.5 nr. 22 p. 22 | 1883 D. resimus, G. O. Sars in:
Forh. Selsk. Christian., 1882 nr. 18 p. 12,58 | 1893 Diastylopsis resima, T. Stebbing in:
Int. sci. Ser., v.74 p. 811 | 1894 Diastylis resimus, A. M. Norman in: Ann. nat. Hist., ser. 6
v.13 p.277 | 1900 Diastylopsis resima, G. O. Sars, Crust. Norway, v.3 p.65 t. 47 | 1900
D.r., Zimmer in: Fauna arctica, v.1 p.426 (with synonymy) | 1901 D.r., T. Scott. in: Rep.
\
108 Cumacea: 11. Diastylidae, 2. Brachydiastylis
Fish. Board Scotl., v.19 p.274 | 1908 D.+., Zimmer in: Ergeb. Tiefsee-Exp., v8 p.1%4 |
1912 D. (2) r., Calman in: P. U.S. Mus., v.41 p. 666.
©. Pseudorostral lobes abruptly_“upturned, with plumose setae radiating
from the apex, antero-lateral cor-
ners forming a sharp tooth, fol-
lowed by a smaller one; eyelobe
obsolete, frontal lobe with 5 median
pairs of denticles, lower margin
of carapace anteriorly denticulate.
Pedigerous segments 3—5)_ with
postero-lateral corners broadly roun-
ded. Pleon segments unarmed, 5t®
Fig. 63.
Fig. 62. B. resimus (Kroyer), peraeopod 1
B. resimus (2) (Kroyer) (after Sars). (after Sars)
not longer than 6%, telson scarcely 2/348 as long as peduncle of uropods,
its slender distal half with about 4 pairs of lateral spines and an apical pair.
Antenna 1, 2"¢ joint shorter than 15*, 34 nearly as long as both combined,
_——
Cumacea: 11. Diastylidae, 2. Brachydiastylis, 3. Diastylopsis 109
setose. principal flagellum 4-jointed, small accessory 2-jointed. Antenna 2
of 4 joints, none elongate. Peraeopod J, 24 joint much shorter than rest
of limb, with 3 strong teeth on distal part of medial margin, 5‘ and 6"
joints equal, each much longer than
the 7; peraeopod 2 short, 5 joint
not equal to 6 and 7 combined:
34 and 4* pairs stout, with 5 joint
broad but shorter than the 4%. Endo-
pod of uropod about as long as the
peduncle, rather over 2/348 as long as
the setose exopod, with only a single
spinule on each of its 3 joints, the
apical spine long. Colour brownish grey.
L. about 5 mm.
GS. Pseudorostral lobes nearly
horizontal with setules at apex, antero-
lateral corners rectangular, unarmed,
the carapace smooth. Pedigerous seg-
ment 5 with postero-lateral corners
acutely produced. Pleon segments
1—3 each with medio-ventral denticle,
5th segment longer than 6", telson
scarcely shorter than peduncle of uro-
pods. Antenna 1, principal flagellum
6-jointed, accessory not very small,
3-jointed; peraeopod 2 with 2™¢ joint /
large, the slender 5 much longer
than 6% and 7t* combined. L. 6 mm. Fig. 65.
Fig. 64.
B. resimus (3) (Kroyer), :
peraeopod 2 (after Sars). depth 6—128 m. (after Sars).
B. resimus (<3)
Arctic, N. Atlantic, Norway, Denmark, (Kréyer), uropod
3. Gen. Diastylopsis S$. I. Smith
1880 Diastylopsis, S. I. Smith in: Rep. geol. Surv. Canada, app. D. p.215B |
1893 D., T. Stebbing in: Int. sci. Ser., v.74 p. 311 | 1900 D., G. O. Sars, Crust. Norway,
v.38 p.64 | 1900 D., Zimmer in: Fauna arctica, v.1 p. 426 | 1908 D., Zimmer in: Ergeb.
Tiefsee-Exp., v.8 p. 181,189 | 1909 D., Zimmer in: Ergeb. Schwed. Siidpol.-Exp., v.6
nr. 8 p.25 | 191l D., Calman in: Tr. zool. Soc. London, v. 181v p. 366, 378 | 1912 D.
Calman in: P. U. S. Mus., v.41 p.662 | 1912 D., T. Stebbing in: Ann. S. Afr. Mus.,
v.10v p. 146, 147.
Pseudorostral lobes of female not strongly upturned with antero-lateral
corners usually produced. Carapace slender. First three pedigerous segments
dorsally short, fourth dorsally long, third and fourth with a tendency to
coalesce, their sides strongly outdrawn backwards, so that in the female the
second and third peraeopods are wide apart. Eye wanting. Second joint of
third maxilliped distally widened. Third and fourth peraeopods in female
with rudimentary exopods; last three pairs of peraeopods stout.
8 species accepted, 2 doubtful.
Synopsis of accepted species:
1 { Telson much shorter than peduncle of uropods — 2.
\ Telson not much shorter than peduncle of uropods — 5.
110 Cumacea: 11. Diastylidae, 3. Diastylopsis .
9 J Fifth pleon segment decidedly longer than sixth .. . 1. D. neozealanica.
Fifth and sixth pleon segments differing little in length — 3.
g 8 g
J
\
Carapace encircled by transverse converging lines . . . 2. D. dawsoni
Carapace without encircling lines — 4.
First pleon segment with prominent medio-ventral tooth,
without latero-ventral spines . Pe ern tt
First pleon segment without prominent ventral tooth,
with latero-ventral spines . = . =. 9. + - »% s « « 4
3. D. elongata
. erassior
s)
J Telson longer than peduncle of uropods ...... . 5. D. annulata
\ Telson not longer than peduncle of uropods — 6.
Carapace shorter than the pedigerous segments combined 6. D. robusta
Carapace not shorter than the pedigerous segments
combined — 7.
Third maxilliped with laminar distal expansion of 2nd
(cl ee Pn annem e me Bowne, ce Cog ce (a IDE loalicpalien
Third maxilliped without laminar expansion of 2nd joint 8. D. dentifrons
1. D. neozealanica (G. M. Thoms.) 1892 Diastylis n., G. M. Thomson in: J. Linn.
Soe. London, v. 24 p. 268 t.18 f. 1—11 | 1908 Diastylopsis (2) neoseelandiae, Zimmer in:
Ergeb. Tiefsee-Exp., v.8 p. 190.
Pseudorostral lobes not much produced, conical, somewhat arched in
dorsal outline, antero-lateral angle obsolete; carapace with eyelobe small or
wanting, dorsally having 4 transverse bars on the anterior half, the last
bilobed, and a fifth parallel to and near the hind margin. Pedigerous seg-
ment 4 the longest, its side-plates directed backwards; postero-lateral corners
of the 5% produced obtusely backwards; 6% pleon segment considerably
shorter than the 5‘ and subequal to the telson, which is about 3/5""S as long
as the peduncle of the uropods and has 8 pairs of lateral spines on the
tapering distal half. Maxilliped 3 with long setae on the widened distal
end of the 2"4 joint, 34 much wider than the small 4% joint. Peraeopod 1
very long, 24 joint shorter than rest of limb, without strong teeth on medial
margin, 5% joint rather longer than 6‘, 6t than 7; peraeopod 2, 5% joint
longer than 6 and 7‘ combined; peraeopods 3, 4 without exopod (text)
but apparently with rudiment (Fig. 8, G. M. Thomson), and their presence
ascertained (Calman). Endopod of uropods about half as long as the
peduncle, 18 joint longer than 24, 24 than 34, with 5, 4, 4, or 6, 3, 4 spines
on the joints respectively, the 2-jointed exopod a little longer. L. @ 8 mm,
3 unknown.
New Zealand, Bay of Islands, depth 15 m.
2. D. dawsoni S.I.Sm. 1880 D.d., S.I. Smith in: Rep. geol. Sury. Canada,
app. D. p.215B | 1900 D.d., G. O. Sars, Crust. Norway, v.3 p. 64 | 1908 D. d., Zimmer
in: Ergeb. Tiefsee-Exp., v.8 p. 183,190 | 1912 D. d., Calman in: P. U.S. Mus., 0 41
p. 605, 662 tf. 81—90.
Pseudorostral lobes well produced, horizontal, acute, antero-lateral
corners acutely produced in Q only; carapace slender, partly encireled anteriorly
by 4 pairs of fine lines, converging downwards, 1st pair nearly meeting
some way in front of the small eyeless eyelobe, 2"¢ and 3° meeting the
sutures of the frontal lobe, which is crossed by lines to the rear of the
2 lateral pairs, the 4” pair not meeting dorsally; lower margin serrate for
a small distance anteriorly. Pedigerous segment 2 with side-plates defined
above by a notch, rounded in front, covering those of segment 1, in the 3
Cumacea: 11. Diastylidae, 3. Diastylopsis 1
themselves concealed by those of segment 3, which in both sexes are strongly
produced backwards: 4 segment very long, firmly united to the 34, 5% with
a pair of ventral teeth. 5 pleon segment not longer than 6%"; telson in ©
D. dawsoni (2 juv.) S. I. Smith (after Calman).
as long as preceding segment, with 9 pairs of lateral spines, in © 3/48 of
preceding segment, with about 4 pairs of lateral spines. Antenna 1, 224 joint
shorter than 34, accessory flagellum 2-jointed. Antenna 2
© of 4 joints, the 34 long. Maxilliped 3, the long 2"¢ joint
much widened distally, medial and lateral apices both pro-
duced, 3¢ joint 3 times as wide as the 4%, Peraeopod 1
uncommonly short, 2™¢ joint much longer than rest of limb.
Peraeopod 2, 5% joint nearly twice as long as 6% and 7t
combined; 34 and 4 pairs in Q with minute rudimentary
seemingly 1-jointed exopod. Peduncle of uropods twice as
long as the telson in © but less than that in the d, more
than twice the endopod but less than twice the exopod; 1S*
joint of endopod equal to the 2°¢ and longer third combined.
L. 9 145 mm, d 9 mm.
Queen Charlotte Islands, 203 m, and other parts off W. coast
of N. America from Alaska to California, at depths 13— 57 m.
3. D. elongata Calm. 1911 D.e., Calman in: Tr. zool. Soc.
London, v. 181v p.312, 379 t.37 f. 1—12.
Fig. 67.
Pseudorostral lobes straight, acute, slightly raised in ©, oe
F ye ele se Ch D. dawsoni (2
not in G, antero-lateral corners rounded, serrate, with the sinus juv.) 8.1. Smith,
above them slightly marked; carapace compressed, smooth; eye- alas eae
; ans 4 riieckee eee 5 : . 7 pedigerous seg-
lobe with no distinct eye, ending in a minute tooth in © only. misntalGatter
Pleon segment 1 with backward pointing medio-ventral tooth, Calman).
5 not much longer than 6, its postero-lateral angles spini-
form, telson differing little in length from the 6% segment, with about 8 pairs
of lateral spines and a short apical pair in the Q, a little longer and narrower
in the d. Antenna 1, 3¢ joint longer than 2™4, flagellum 3- or 4-jointed,
accessory 2- or 3-jointed. Antenna 2 of © 3-jointed. Peraeopod 1, 2"¢ joint
considerably longer than rest of limb, with tooth at distal medial corner,
5th and 6 subequal, each longer than 7‘; peraeopod 2 9, 24 joint con-
siderably longer than rest of limb, 2"4, 34, and 4% each with small tooth
at distal medial corner; rudimentary exopods of 3¢ and 4" peraeopods in ©
2-jointed; peraeopod 2 d, 2"¢ joint scarcely longer than rest of limb, 5% joint
much instead of little longer than 4%, 6 longer than 7, which ends in
2 little curved claws; peraeopods 3, 4, 2%¢ joint toothed at lateral apex, in
4, 5 the 3¢ joint toothed at medial apex. Peduncle of uropods once and
a half as long as the telson, slightly less than that in proportion to the
112 Cumacea: 11. Diastylidae, 3. Diastylopsis
nearly equal rami, exopod slightly the shorter, 15* joint of endopod longer
than 2"4 and 3¢ combined, marginal spines numerous. L. about 9 mm.
New Zealand, Lyttleton Harbour, depth 2—9 m; Akaroa Harbour.
4. D. erassior Calm. 1911 D. c., Calman in: Tr. zool. Soe. London, v. 181v
p. 312, 381 t.37 £.13—18.
Near to D. elongata, but pseudorostral lobes not raised, less acutely
produced; eyelobe not prominent dorsally, without apical tooth; pleon seg-
ments stouter, 18¢ with no strong medio-ventral tooth but with 2 ventro-
lateral rows of denticles, postero-lateral angles of 3¢ and 4% as well as 5!
spiniform, telson less narrowed distally, peraeopod 2 with 244 joint little longer
than rest of limb, produced to a strong tooth at distal medial corner, the a
joint having 2, and the 4% 3 strong teeth, the following legs stouter and
longer than in the companion species; peduncle of the uropods twice as long
as the telson, about once and a half as long as the endopod, which is shorter
than the exopod. L. Q 9°25 mm, GO unknown.
New Zealand, Lyttleton Harbour, depth 2—9 m.
5. D. annulata (Zimmer) 1902 Leptostylis a., Zimmer in: Ergeb. Hamburg.
Magalh.-Sammelr., v.2 Cumac. p.10 tf.15—17 | 1908 Diastylopsis a., Zimmer in:
Ergeb. Tiefsee-Exp., v.8 p.183, 190 | 1909 D. a., Zimmer in: Ergeb. Schwed. Siidp.-Exp.,
v.6 or.8 p.1,2,23, v7 nor.3 t.8 £. 131—183.
Pseudorostral lobes not upraised, well produced in front of the eyeless
eyelobe, antero-lateral corners acute, producing a well marked sinus; carapace
with a longitudinal series of 3 forward pointing teeth on the frontal lobe,
which is crossed by 2 lines and bounded by a bilobed third line; to these
succeed a pair not meeting in the middle, 3 others which nearly meet and
2 more which show no median interruption, the last of them forming the
upraised hind margin. The postero-lateral corners of the 5% pedigerous seg-
ment are not outdrawn. The fifth pleon segment is little longer than the
6th or than the telson, of which the basal section carries 3 pairs of setae
while the narrowed end has 4 pairs of lateral spines and a somewhat larger apical
pair. Peraeopod 2, as figured, has the 4 joint longer than the 5th; the 34
and 4t pairs have rudimentary 2-jointed exopods; the 4% and 5th pairs
have the 24 joint remarkably short. The peduncle of the uropods is a little
shorter than the telson, but longer than the endopod, in which the 1%* joint
is longer than 224 and 3¢ combined, the spines on the medial margin being
respectively 5, 2, 2. Colour yellowish white. L. Q not fully adult, 6 mm.
South Georgia, low tide to 15 m.
6. D. robusta (Zimmer) 1902 Leptostylis r., Zimmer in: Ergeb. Hamburg.
Magalh.-Sammelr., v.2 Cumac. p.13 tf. 22—31 | 1908 Diastylopsis r., Zimmer in:
Ergeb. Tiefsee-Exp., v.8 p. 183, 190.
Pseudorostral lobes shortly produced in front of the very small eyelobe,
antero-lateral corners bluntly produced and forming a distinct sinus. Cara-
pace shorter than the combined pedigerous segments, crossed by a pair of
lines or ridges which end without meeting near the back of the frontal lobe;
the pedigerous segments have the front margin finely denticulate, and the 34
and 4th have each 2 ridges on the expanded sides parallel to the hind margin.
The pleon segments carry dorsally 3—6 plumose setae and have postero-
ventral spines; the 5t segment is rather longer than the 6th, and the 6%
Cumacea: 11. Diastylidae, 3. Diastylopsis 113
than the telson, to which are attached 2 pairs of lateral spines, the upper
very small, and a large apical pair. Antenna 1 O with each flagellum 3-jointed,
in the O the principal is 5-jointed. Antenna 2 © showing 1 large joint
followed by a minute one. The peduncle of the uropods in the Q is as long
as the 6" pleon segment and as the exopod, rather longer than the endopod,
which carries 6, 2, 2 spines respectively on the 3 joints; in the G the
peduncle is rather longer, and the endopod has 12, 4, 3 spines on the medial
margin, the apical spine being continuous with its 34 joint. Colour yellowish
white. L. about 8 mm.
Magellan Strait, Punta Arenas, depth 4 m.
7. D. thileniusi (Zimmer) 1902 Leptostylis t., Zimmer in: Zool. Jahrb., Syst.
v.17 p.449 tf. G—W | 1908 Z.¢., Calman in: Ann. nat. Hist., ser.8 v.1 p. 239 | 1908
Diastylopsis t., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 183, 190.
Pseudorostral lobes bluntly produced in front of the eyelobe, sinus
defined by an acute antero-lateral tooth. Carapace little longer than pedigerous
Fig. 68. Fig. 69.
D. thileniusi (2) (Zimmer). D. thileniusi (Zimmer), maxilliped 3.
segments combined, frontal lobe with 2 transverse lines, a pair of lateral
lines from the middle of its sutures not quite reaching the lower margin of
the carapace, and a shorter pair crossing the ends of the sutures. Pedigerous
segments 1—3 dorsally short, with denticulate front margin, sides of 34
almost encircling the very long 4*, with which it is nearly fused, 4 nearly
surrounding the small 5%, of which the hind corners are little and obtusely
extended. Pleon segments with postero-ventral spines and some dorsal setae;
telson rather longer than the preceding segment, with 2 pairs of setae on
the wide basal section and 6 of lateral spines on the narrowed end. Antenna 1,
accessory flagellum very small. Antenna 2, 3- (text) or 4-jointed (fig.). Maxilli-
ped 3, 2™4 joint greatly widened distally and on lateral apex strongly pro-
duced to an acute point, while the whole distal margin is produced to
form a rounded plate covering (from below) nearly all the remaining joints.
Peraeopod 2 with 4 joint longer than 5%, 34 and 4 pairs with rudi-
mentary exopods, 4 pair with 24 joint not especially short. Peduncle of
uropods nearly as long as the telson, longer than the endopod, which is
Das Tierreich. 39. Lief.: T. R. R. Stebbing, Cumacea. 8
114 Cumacea: 11, Diastylidae, 3. Diastylopsis, 4. Adiastylis
longer than the exopod and has 4, 3, 3 spines on its 3 joints respectively =
colour yellowish white. L. © about 5 mm.
New Zealand, Bay of Plenty.
8. D. dentifrons (Zimmer) 1908 Leptostylis d., Zimmer in: Zool. Jahrb., Syst.
v.18 v1 p.688 tf. Z. AA. BB. | 1908 Diastylopsis d., Zimmer in: Ergeb. Tiefsee-Exp., v. 8
p. 158, 190 t.46 f. 109—120 | 1909 D. d., Zimmer in: Ergeb. Schwed. Siidp.-Exp., v.6
or. 3 p.26. ;
Very near to D. annulata (ur. 5) but here the distal section of the
telson carries 9 pairs of lateral spines and a strong apical pair, and the
telson itself is scarcely longer than the peduncle of the uropods, of which
the rami are only 2/348 as long as the peduncle but subequal one to the other,
the 18* joint of the endopod not so long as the 2nd and 3¢ combined, the
spines of the 3 joints being respectively 7, 3, 2. Colour yellowish white.
L. reaching 11 mm.
Rhodes Bay, Kerguelen, 19 m.
D. insularum (Calm.) 1908 Leptostylis (?) i., Calman in: Ann. nat. Hist., ser. 8
v1 p. 234 tf. 1—5a | 1911 Diastylis i., Calman in: Tr. zool. Soe. London, v. 181v
p. 342, 385.
Agrees closely with D. neozealanica (nr.1) in form and armature of telson and
uropods, in having the 34 joint of the 84 maxilliped wider than the 4th, and the
34 and 4th peraeopods of the Q provided with minute 2-jointed exopods, but is
distinguished by the carapace, which has the pseudorostral Igbes horizontal, acute,
much more produced in @ than in G, eyelobe a little broader than long, inflated, but
without pigment, frontal lobe crossed by 2 low, rounded, crescentic ridges, with a shallow
depression at the end of each of its bounding sutures; a longitudinal depression between
the branchial regions, a slight elevation of the dorsal hind margin, the whole surface
spinulate and an oblique forward running line of spinules on each antero-lateral surface.
Antenna 1 of ¢ with 84 joint much shorter than 1st or 2nd, instead of subequal, as.
in 9, to each. Peraeopod 1 has the 6th joint a little longer than either of the subequal
5th and 7th joints. L. between 9 and 10mm. Seems to be a variety of D. neozealanica.
New Zealand, Bay of Islands; Lyttleton Harbour, 2—9 m.
D. sp. (Calm.) 1908 Leptostylis sp., Calman in: Ann. nat. Hist., ser.8 v.1 p.239.
Agrees closely with D. thileniusi (nr. 7), but has carapace and pedigerous segments.
together not longer than the pleon, 34 pedigerous segment not strongly produced backwards.
at the sides, so that the interval is slight between peraeopod 2 and 3, and the telson
is armed with only 8 pairs of lateral spines. L. largest G 3:3 mm. Probably young
stage of D. thileniusi.
New Zealand, Bay of Islands.
4. Gen. Adiastylis T. Stebbing
1912 Adiastylis, T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 146, 148.
Telson cylindric, with lateral spines on the short, narrowed distal
section. Eye wanting. Maxilliped 1 with branchial leaflets not very numerous.
Maxilliped 3, 24 joint distally widened, the following joints not widened.
First peraeopod elongate.
4 species.
Synopsis of species:
J Peduncle of uropods longer than the telson — 2.
: \ Peduncle of uropods shorter than the telson — 3.
Cumacea: 11. Diastylidae, 4. Adiastylis 115
. A. acanthodes
9 Pedigerous segments dorsally dentate. .... '
A. longicaudatus
A.l
Pedigerous segments not dorsally dentate . :
‘ ongipes
1
Sire
3 J Exopod of uropods much shorter than the endopod . . 3
+. A. costatus
\ Exopod of uropods not shorter than the endopod . .
1. A. acanthodes Stebb. 1912 A. a, IT. Stebbing in: Ann. S. Afr. Mus.,
v, 10v p. 148 t. 53.
With points of resemblance to Diastylis anomalus (p. 106).
Pseudorostral lobes well produced acutely in advance
of the eyeless eyelobe, forming an obtuse angle with the
curved medio-dorsal line of the carapace, their upper surface
like the rest of the carapace diversified with many unequal
denticles. Pedigerous segments much denticulate, the last 4
each with a pair of dorsal teeth specially conspicuous, the
first 2 each with front margin serrate. Pleon longer than
anterior part of the body, with dorsal and some subventral
denticles, 5 segment much longer than 6%, but shorter than
the telson, which is about ?/,¢S as long as peduncle of uropods,
its last third narrow, with 3 or 4 pairs of lateral spines and
a larger apical pair. Antenna 1, principal flagellum 5- (or 6-)
jointed, accessory 4-jointed. Antenna 2, 2"4 and 5t* joints
of peduncle long, flagellum comparatively short. Maxilliped 3,
2nd joint with long setae on the dilated end, none of the other
joints dilated. Peraeopod 1, 2" joint denticulate. Peraeo-
pod 2, with tooth at apex of the short stout 2"¢ joint, 5%
and 7 joints long and slender, 7* more than twice as
long as the 6. Peraeopods 3 and 4 with 2™4 joint striking-
ly narrowed distally. Pleopods 1 considerably longer than
the 2" pair. Peduncle of uropods much longer than exopod,
which is little longer than the first 2 joints of the endopod,
2nd joint of endopod ?/,4 the length of the 18%, 34 joint
missing from the specimen. L. d about 9mm. © unknown.
S. Africa, Cape Natal N. by E. 24 miles, depth 805 m.
2. A.longicaudatus (Bonnier) 1896 Leptostylis longicaudata,
J. Bonnier in: Ann. Univ. Lyon, v.26 p.557 +.30 f.2,2a—o | 1908
Diastylis 1., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p.183 | 1912 Adi-
astylis longicaudatus, 'T. Stebbing in: Ann. S. Afr. Mus., v.10v_p. 148,
Pseudorostral lobes acutely produced, antero-lateral cor-
ners faintly marked, not produced; carapace besprinkled with eae
_ spinules, medio-dorsal line of frontal lobe slightly undulating; Stee ee
pedigerous segments and pleon not dentate, 5t segment of — and uropoa.
pleon not much longer than others, telson shorter than peduncle
of the uropods but much longer than the rami, the short distal section carrying
3 pairs of lateral spines and a larger apical pair. Eye quite wanting. An-
tenna 1, 3° joint of peduncle not longer than 2", principal flagellum 4-jointed,
accessory 3-joited, very short. Antenna 2 of 4 joints, the 1* the longest.
Peraeopod 1 very long, 7 joint a little longer than the 6; peraeopod 2,
5th and 7% joints not very long, 7" only twice as long as the 6th. Perae-
opods 3 and 4 with 2-jointed rudimentary exopods. Exopod of uropods
notably longer than the endopod, of which the 1% joint is not much longer
8*
116 Cumacea: 11. Diastylidae, 4. Adiastylis
than the 2™¢ or 34, with 3, 2, 2 spines respectively on the medial margin.
L. Q juv. nearly 10 mm. GC unknown.
Bay of Biscay, depth 650 m.
3. A. longipes (O. Sars) 1871 Diastylis |., G. O. Sars in: Ofv. Ak. Forh., v.281
p. 74 | 1871 D.1, G. O. Sars in: Svenska Ak. Handl., n. ser. v.9 nr. 13 p.82 t.13 £.65— 67 |
1879 D.1., A. M. Norman in: Ann. nat. Hist., ser.5 v.3 p.58 | 1896 D. l., J. Bonnier in:
Ann. Univ. Lyon, v.26 p.550 t.29 f.4a—p | 1912 Adiastylis L., T. Stebbing in: Ann.
S. Afr. Mus., v. 10v p. 148.
Pseudorostral lobes moderately produced in advance of the small eyeless
eyelobe, horizontal, acute, with a subapical pair of spinules, antero-lateral
angles obsolete; anterior part of carapace with several variously disposed
spinules; 18* pleon segment with ventral longitudinal crest of 4 or 5 con-
spicuous teeth, 5% and 6 segments subequal, telson rather longer than
both combined, and longer than peduncle of uropods, the narrowed distal
part short, with 4 pairs of lateral spines and a small apical pair. Antenna 1
elongate, 24 joint of peduncle much longer than 34, principal flagellum long, »
6-jointed, accessory stout, 3-jointed. Mandibles with 16 or 17 spines in the
spine-row. Maxilliped 3, with a strong spine on the 2nd and also on the
4th joint. Peraeopod 1 extremely long, especially in regard to the last
3 slender joints, of which joint 6 is actually longer than the 2¢ joint. Peraeo-
pod 2, 5t joint more than twice as long as the 6 and 7‘ combined in
the 3 (G. 0. Sars), 7 very short (G. 0. Sars), not very short in the ¢ (Bonnier).
Peraeopods 3, 4, 4% joint more or a little less than twice as long as the
3 following joints combined. Rudimentary exopods on these limbs not recorded
for © juv. Peduncle of uropods much longer than the rami, endopod with
1st joint longer than 2"4 and 34 combined, spines on medial margin respectively
12 or 13, 5, 4; exopod little longer than 1%* joint of endopod. L. C juv.
about 8 mm, © juv. 12 mm.
Atlantic: Lat. 60° 45’ N., long. 4° 49’ W., 933 m; lat. 59° 41' N., long. 79 34’ W.,
838 m; lat. 38° 7/ N., long. 99 18’ W., 1036 m; Bay of Biscay, depth 950 m.
4. A. costatus (Bonnier) 1896 Diastylis costata, J. Bonnier in: Ann. Univ. Lyon,
v.26 p.553 t.30 f.1,la—m,o | 1912 Adiastylis costatus, T. Stebbing in: Ann. 8S, Afr.
Mus., v. 10v p. 148.
Pseudorostral lobes acute, slightly raised. Eyeless eyelobe very small,
behind it runs a median serrate crest, bifurcating about the middle of the
carapace, the 2 arms reaching the hind margin; more or less oblique to the
median are 4 pairs of curved crests, and spinules are irregularly dispersed
in the intervals. Pedigerous segments 3—5 with transverse rows of spinules;
pleon segments 1—5 with longitudinal rows; telson distally with 5 lateral
pairs and an apical pair. Antenna | with 4 spines on 1st joint, 34 joint
longer than 2"4, principal flagellum 4-, accessory 2-jointed. Antenna 2 4-jointed,
34 joint the longest. Mandible with 12 spines in the spine-row. Peraeo-
pod 1, 24 joint very long. Peraeopod 2, the long slender 5th joint more
than twice as long as 6% and 7 together, 7% twice the 6%. Peduncle of
uropods shorter than the telson, but much longer than the exopod; exopod
a little longer than the endopod, the 1&* joint of which nearly equals the gud
and 34 together, with 2, 2, 1 spines on the medial margin of the 3 joints
respectively. L. @ juv. 6 mm.
Bay of Biscay, depth 1410 m.
Cumacea: 11. Diastylidae, 5. Makrokylindrus 117
5. Gen. Makrokylindrus T. Stebbing
1912 Makrokylindrus, T. Stebbing in: Ann. S. Afr. Mus., v. 10v_ p. 129, 146, 150.
Carapace denticulate; no distinct eye; telson elongate, basal portion
cylindric, much longer than the post-anal portion, which carries only the two
apical spines. Peraeopods of the female, so far as known, without rudimentary
exopods on the third and fourth pairs.
6 species.
Synopsis of species:
i! Pélson distally serrate ..-....-+.-+.-.+-.-.-. J. M.serricauda
\ Telson not distally serrate — 2.
» J Carapace with transverse raised girdle — 3.
“ \ Carapace without transverse raised girdle — 4.
3 { Eyelobe distinct ........-. Wis Sey ne om ae seneires te Mi rags
\ Eyelobe wanting . : Ae icant ak 3. M. cingulatus
4 { Pedigerous segments 3 Ne 1 nora faced Seu ey 4. 20 dubins
| Pedigerous segments 3 and 4 not dorsally fused — 5.
- | Pedigerous segments smooth ..... ++ +--+. + 5. M. erinaceus
\ Pedigerous segments spinose ..°. ......- =. + 6. M.josephinae
1, M. serricauda (T. Scott) 1912 Diastylis s., T. Seott in: Sci. Invest. Fish.
Seotl., 1911 nr.1 p.3 t. 1 f.1—4 | 1912 Makrokylindrus s., T. Stebbing in: Ann. S. Afr.
Mus., v. 10v p.150.
Near to JL, josephinae (nr. 6)
but distinguished by the telson.
Pseudorostral lobes well produced to
blunt apex. Carapace dorsally
strongly spinose and spinulose, lower
margin pectinate for more than half
its extent. from the front. Pedigerous
segment 5 the longest, its postero-
lateral angles rounded. Pedigerous
segments and pleon all dorsally
rough with spicules, the pleon seg-
ments also ventrally. Telson rather
longer than peduncle of uropods,
only slightly narrowing in the distal
third, the sides of which are coarsely
serrate, with intervening setules;
the apical spines stout, somewhat
divergent. Endopod of uropods
shghtly longer than the exopod,
< Rigs 71s
half as long as the peduncle, its M. serricauda (2) (T. Scott), telson and uropod.
1st joint twice as long as the 2
subequal following joints combined; spines on the medial margin of the 3
Joints respectively 7, 1, 1. L. © about 12 mm.
- Lat.-59° 386° N., long. 7° 00’ W., depth 140 m.
2. M. fragilis Stebb. 1912 M. 7, IT. Stebbing in: Ann. S. Afr. Mus., 7.10v
p. 150 t. 54, 55.
Pseudorostral lobes well produced subacutely in front of the prominent
rounded eyeless eyelobe, with dorsal denticles near to the meeting line.
118 Cumacea: 11. Diastylidae, 5. Makrokylindrus
Behind the eyelobe a central ridge, elevated at the middle, ascends to a
bilobed girdle which crosses the carapace a little behind the middle, each
lobe in its forward curve meeting a dentate carina which starts from the
pseudorostral processes, while from the meeting point a ridge descends, giving
off two divergent branches which reach the lower margin. Pedigerous seg-
ments 3 and 4 dorsally coalesced, 5t rather long with postero-lateral angles
rounded. Pleon segments 1—3 in the d each with a dorsal pair of den-
ticles, the rest and all in the © smooth; telson cylindric till near the end,
sides smooth at the base, then serrate, the hinder half smooth, the truncate
apex carrying a pair of large spines. Antenna 1, first 2 joints elongate,
f \
; \
Bok
Dg ORS ent RS Saas
Fig. 72. Fig. 73. Fig. 73a.
M. fragilis Stebbing, telson and M. fragilis (2) Stebbing, M. fragilis Stebbing,
uropod. maxilliped 2. maxilliped 2.
principal flagellum 4-jointed, accessory small, 2-jointed. Antenna 2 of 9
4-jointed. Mandibles with about 20 spines in the spine-row. Maxilla 2 with
distal plate seemingly undivided. Maxilliped 1, branchial leaflets numerous.
Maxilliped 2 with long lamina in place of vibratory fan. Maxilliped 3,
2nd joint strongly produced, 3% short, 5%, 6%, and 7 equal in length.
Peraeopod 2 with 6 joint very short. Pleopods of male with short rami.
Uropods slender, peduncle shorter than telson, endopod 2/;°S as long as
the peduncle, with 7, 5, 4 spines on medial margin of its 3 joints
respectively in the Q, 15* joint a little longer than the equal 2™¢ and
34 combined; exopod with short 18 joint, 2"¢ equal to 18* of the endopod.
L. 10 mm.
S. Africa, Cape Natal N. by E. 24 miles, depth 805 m.
Cumacea: 11. Diastylidae, 5. Makrokylindrus 119
3. M. cingulatus (Calm.) 1905 Diastylis c., Calman in: Siboga-Exp., nr. 36
p. 2,19 t.2 f.26—29 | 1908 D. cingulata, Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 183. |
1912 Makrokylindrus cingulatus, T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 150.
Pseudorostral lobes well produced, horizontal,
subacute, with 2 apical spines. Carapace subglobose, —
encircled about at the middle by a stout sinuous 4
ridge closely set with stout spines, in front of
ridge many spines, mostly uncinate and distally
tuberculate, and on each side a single smooth straight
spine longer than the rest; behind the ridge spines
larger, less crowded, short, stout, conical; lower
margin fringed with spines at the rear short and
thick, towards the front long and slender. Eyelobe
and eye wanting. Pedigerous segment 2 at the
sides covering the 18*, 34 and 4 fused, their side-
plates not expanded, those of the 5 not extended
backwards. Pleon segments 1—6 beset with spi-
nules. Telson longer than peduncle of uropods, 3/4'"s
of length cylindric, with sides denticulate near the
base, the tapering post-anal fourth slightly decurved,
with subapical pair of setules and apical pair of
spinules. Antenna 1, first 2 joints of peduncle rather
long, accessory flagellum small. Peraeopod 5 arti-
culated near the dorsal surface of the segment.
Peduncle of uropod about 5/,'*S as long as telson,
with 8 spinules on distal half of the medial margin;
endopod not quite half as long as the peduncle,
with long slender spines on the medial margin, 1**
joint longer than 2™4 and 3¢ combined, 2"? longer
than 34, exopod only reaching end of the 2". L.
G juv. 15 mm.
Malayan Archipelago: Lat. 5° 40‘ 7'S., long. 120°
45‘ 5 B., depth 1158 m; lat. 6° 24'S., long. 124° 39’ E.,
depth 2798 m.
\
4. M. dubius (Bonnier) 1896 Diastylopsis (2) dubia, Fig. 74.
J. Bonnier in: Ann. Univ. Lyon, v.26 p.539 t.30 £3, M. cingulatus (di jnv.) (Cal-
3a—m | 1900 D. (?) tumida (laps. for dubia), G. O. Sars, ace
Crust. Norway, v.83 p. 64 | 1905 D (?) dubia, Calman in:
Siboga-Exp., nr.36 p.23 | 1908 D. (?) d., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 182 |
1912 D. (2) d., T. Scott in: Sci. Invest. Fish. Scotl., 1911 nr.1 p.4 t.1 £.5 | 1912
Makrokylindrus dubius, T. Stebbing in: Ann. S. Afr, Mus., v. 10v p. 150.
Pseudorostral lobes horizontal, acute, with apical pair of spines. Cara-
pace and other parts of body beset with spinules and spines. Pedigerous
segments 3 and 4 dorsally coalesced, the postero-lateral corners not extended
backwards. Telson longer than peduncle of uropods, full 2/35 of length
cylindric, with lateral series of spinules (or serration Scott), last third tapering,
armed only with apical pair of spinules. No eye. Antenna 1 with long
spinose 1st joint, 2"¢ intermediate in length between 1** and 34, flagellum
4-jointed, the small accessory 3-jointed. Mandible with 17 spines in spine-row.
Maxilliped 1 with about 10 small branchial leaflets. Maxilliped 3, 2nd joint
distally widened, moderately produced. Rudimentary exopods on peraeopods
120 Cumacea: 11. Diastylidae, 5. Makrokylindrus
3 and 4 of Q juv. supposed to be inconstant, but possibly the young specimen
showing these appendages may have been a G juv. Endopod of uropods
rather more than half as long as the peduncle, slightly longer than exopod,
1st joint longer than 2"¢ and 3¢ combined, spines on medial margin respect-
ively 4, 2, 2, with a longer apical spine. L. © juv. 9mm, CG juv.
slightly larger.
Bay of Biscay, depth 650—950 m.
5. M. erinaceus (O.Sars) 1887 Diastylis e., G.O. Sars in: Rep. Voy. Challenger,
v.19 pars 55 p.9, 44,58 t.9 f.9—14 | 1905 D. e., Calman in: Sci. Invest. Fish. Ireland,
1904 app.1 p.47 | 1912 Makrokylindrus e., T.Stebbing in: Ann. 8. Afr. Mus., v. LOv p. 150.
Pseudorostral lobes horizontal, acute, well produced in front of the
small eyeless eyelobe. Carapace large, tumid, without carinae, beset with
numerous forward pointing spines of generally uniform size, continued along
sides of pseudorostrum; anterior part of lower margin finely denticulate.
Pedigerous segments smooth with side-plates evenly rounded. Pleon seg-
ment | with serrate ventral keel, this and the next 2 segments each with
paired dorsal spines and postero-dorsal setae. Antenna 1 with spinose 1*t
joint, 2™¢ nearly as long, 34 much shorter, accessory flagellum very small.
Antenna 2 of © 5-jointed, twisted. Peraeopod 1 very long, 18* joint very
setose, 5 and 6% subequal, very long and slender, each much longer than
the 7, Peraeopod 2 with 5 joint nearly twice the combined length of 6%
and 7%, Telson about as long as the 2 preceding segments together, two-
thirds smoothly cylindric, the post-anal third tapering to truncate apex which
carries a pair of small spines. Peduncle of uropods decidedly shorter than
telson, probably longer than the rami (which are broken). L. O 13 mm.
Atlantic: Lat. 369 21‘ N., long. 23° 31’ W., depth 3700 m.
6. M. josephinae (0. Sars) 1871 Diastylis j., G. O. Sars in: Ofv. Ak. Forh.,
v.28 nr. 1 p.77 | 1871 D.j7., G. O. Sars in: Svenska Ak. Handl., n. ser. v.9 nr. 13 p. 36
t.15 f.72—74 | 1879 D.j., A.M. Norman in: Ann. nat. Hist:, ser.5 v.3 p.56 | 1900 D.7.,
Zimmer in: Fauna arctica, v.1 p.422 | 1905 D. J. (?part.), Calman in: Sci. Invest. Fish.
Ireland, 1904 app.1] p.10,44 | 1912 Makrokylindrus j., T. Stebbing in: Ann. S. Afr. Mus.,
v.10v p. 150.
Pseudorostral lobes well produced in front of the small eyeless eyelobe,
horizontal, acute, with apical pair of spines. Carapace large and deep, armed
on surface with crowded spinules interspersed with spines; especially con-
spicuous are series formed by 5 pairs of spines commencing on the pseudo-
rostral lobes, and between these rows 4 submedian pairs. Pedigerous seg-
ments with transverse dorsal rows of spinules, sides of last segment bluntly
extended; pleon segments spinulose all over; telson longer than peduncle of
uropods, basal two-thirds cylindric, having in the type specimen about 14 pairs
of lateral and 5 ventral spines but none on the tapering end except the
apical pair, whereas in an ovigerous Q Calman found basal part with no
lateral serrations but on each side of post-anal part 10 or 11 small teeth
(? M. serricauda Scott, p. 117). Antenna | slender, 2"4 joint the longest. Peraeo-
pod 1 very long. Peraeopod 2, 5t* joint considerably longer than 6' and
7% combined. Endopod of uropods appears to be only 2-jointed in young
specimens and subequal to the exopod, which with advancing age it exceeds
in length, at the same time becoming 3-jointed. L. over 11 mm.
Off Portugal, lat. 389 10‘ 30” N., long. 9° 25’ W., depth 1412 m; N. of Faro
Islands, lat. 48° 50‘—67° 7’ N., long. 3° 6’—11° 9’ W., 629—1326 m; off W. of
Ireland, depth 364—699 m.
Cumacea: 11, Diastylidae, 6. Paradiastylis 121
6. Gen. Paradiastylis Calman
1904 Paradiastylis, Calman in: Herdman, Rep. Ceylon Pearl Fish., v.2 p.175
1905 P., Calman in: Siboga-Exp., nr. 36 p.20 | 1908 P., Zimmer in: Ergeb. Tiefsee-
Exp., v.8 p.181, 182 | 1911 P., Calman in: Tr. zool. Soe. London, v. 181v p. 366 | 1912
P., T. Stebbing in: Ann. 8. Afr. Mus., v. 10v p. 146, 147.
Telson rather short, with few lateral spines or none. Third maxilliped
with exopod only in the male. Third and fourth peraeopods in the female
quite devoid of exopods.
3 species.
Synopsis of species:
Endopod of uropods with Ist joint twice as long as 2nd
| and 34 combined. .... .~ Cant Oe et
| Endopod of uropods with 1st joint not longer than 2nd
and 34 combined — 2.
Peduncle of uropods twice as long as pleon segment 6 2. P. brachyurus
Peduncie of uropods thrice as long as pleon segment 6 3. P. longipes
1. P. brevicaudatus
1. P. brevicaudatus (Zimmer) 1903 Leptostylis brevicaudata, Zimmer in: Zool.
Jahrb., Syst. v.18v1 p. 685 tf. W—Y | 1904 Paradiastylis (2) b., Calman in: Herdman,
Rep. Ceylon Pearl Fish., 7.2 p.175 | 1908 P. b., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p.181.
Pseudorostral lobes horizontal, acute, antero-lateral corners subacute,
forming a well marked sinus. Eyelobe small, eyeless, with a pair of denticles,
carapace much sculptured, 2 strong teeth in line on the frontal lobe, followed
by a deep median depression of the hinder half of the carapace flanked by
a pair of keels from which on either side 4 lateral forward curving keels
take their origin; hind margin dorsally raised. Pedigerous segment 4 the
longest, with a dorsal pair of quadrate impressions, 5‘ with a postero-dorsal
upturned conical process. Pleon segment 5 the longest, telson slightly shorter
than the 6 segment, its narrowed distal portion much shorter than the
broad proximal, with 1 pair of lateral spines and a small apical pair with
pair of setules adjacent. Antenna 1 rather long and slender. Peraeopod
1 long. Exopods of 34 and 4) pairs slightly developed, the specimen being
probably G juv. Peduncle of uropods more than twice as long as the telson,
which is rather longer than the endopod; 1S* joint of endopod about twice
the 284 and 34 joints combined, 2°¢ nearly twice the 34, spines of medial
margin respectively 5, 2, 1; exopod a little shorter than endopod. Colour
brown. L. d juv. about 9 mm.
Japan: Enosima, depth 6—8 m.
2. P. brachyurus Calm. 1904 P. brachyura, Calman in: Herdman, Rep.
Ceylon Pearl Fish., v2 p. 159,174 t.5 f. 76—90 | 1905 P. b., Calman in: Siboga-Exp.,
nr. 36 p.20[ 1908 P. b., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 181.
Pseudorostral lobes horizontal, acute, with a pair of divergent spines
at apex, antero-lateral corners not produced, rounded, serrate. Hyelobe twice
as broad as long, with a pair of denticles, no distinct eye. A median keel
bears 2 procurved teeth on front part of carapace, on each side 4 curved
ridges run obliquely down and forwards, the front one serrate, the lower
edges crossed by a curved row of spines running down and back from
pseudorostral lobes towards the lower margin; hind margin dorsally raised.
Pedigerous segment 5 without dorsal process. Pleon segments with slight
serrate dorso-lateral crests, 5*® not longer than 6, telson a little shorter
122 Cumacea: 11. Diastylidae, 6. Paradiastylis
than either, slightly narrowed at the base, with a pair of tubercles at about
one-third of its length, distal half sharply convergent to exceedingly narrow
apex carrying a pair of spinules, with pair of setules adjacent and a second
pair higher up. Antenna 1, 2™¢ joint of peduncle the shortest, principal
flagellum 3-jointed, accessory 2-jointed. Antenna 2 of Q 3-jointed. Mandibles
with about 9 spines in spine-row. Maxilliped 3, 2°¢ joint much expanded
distally, following joints not expanded. Peraeopod 1 not very long, 24 joint
little longer than 6, which is more than twice as long as the 7. Peraeo-
pod 2, 284 joint broad, 34 a narrow ring, 6% 3/4%s as long as 7. Following
limbs rather stout, 4 joint much longer than 5%. Peduncle of uropods
twice as long as 6t® pleon segment, a little less than twice the exopod apart
from its long apical spine; exopod a little shorter than endopod, of which
the 1st joint is a little shorter than 2"? and 3? combined, spines of medial
margin respectively 2, 1, 1. L. Q 32mm.
Gulf of Manaar, Cheval Paar, depth 13 m.
3. P. longipes Calm. 1905 P. 1., Calman in: Siboga-Exp., nr. 36 p.2, 21
tf.4a—i | 1908 P. l., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p.181 | 1911 P. 1., Calman in:
Tr. zool. Soe. London, v. 181v_ p. 376.
Near to P. brachyurus, but the lateral surface of the carapace has
only 3 oblique ridges, no strong dorsal teeth, only a pair of denticles on
Fig. 75. Fig. 76.
P. longipes (2) Calman, maxilliped 3. P. longipes (dQ) Calman, maxilliped 3.
the eyelobe and a few in front of the anterior oblique ridge, the paired
longitudinal crests on hinder part of dorsal surface finely serrated and more
prominent, the space between them deeply hollowed; pedigerous segments 3,
4 indistinctly separated; pleon more slender, lateral serrations more extensive;
antenna 1 with 32 segment longer and more slender; maxilliped 3 with
narrower 24 joint; peraeopod 1 much longer, following limbs more slender,
peduncle of uropods thrice as long as 6%" segment of pleon. In the adult 3
the sculpture of the carapace is less strongly marked than in the Q, the
broad eyelobe has 3 large but ill-defined corneal lenses; pleon segments
1—6 quite smooth, pre-anal part of telson subcylindrical, relatively longer
than in Q, a pair of spinules adjacent to the apical pair. Antenna 1 with
each flagellum 4-jointed. Maxilliped 3 with 2°¢ joint much broader than
Cumacea: 11. Diastylidae, 6. Paradiastylis, 7. Leptostylis 123
in Q in accord with well developed exopod. Peraeopods 1—4 with 2nd joint greatly
expanded and distally produced. Endopod of uropods with 6, 3, 3 spines on
the 3 joints respectively, the 34 joint longer than the 2"¢. L. 9 3mm, d 3:5 mm.
Sulu Archipelago, surface to 32m; Gulf of Siam, depth 9—18 m.
7. Gen. Leptostylis 0. Sars
1869 Leptostylis, G. O. Sars in: Nyt Mag. Naturv., v.16 p.343 | 1893 L., T. Stebbing
in: Int. sci. Ser., v.74 p.311 | 1900 Z., G. O. Sars, Crust. Norway, v.3 p.67 | 1900
L., Zimmer in: Fauna arctica, v.1 p.427,431 | 1908 L., Zimmer in: Ergeb. Tiefsee-
Exp., v.8 p. 181,182 | 1911 Z., Calman in: Tr. zool. Soc. London, v. 181Vv p. 366, 367 |
1912 L., T. Stebbing in: Ann. S. Afr. Mus., v.10v p.129, 152.
In general agreement with Liastylis, but having a shorter telson, with
lateral spines few or none; postero-lateral angles of fifth pedigerous segment
not acute; eye wanting; first antenna of female with second joint of peduncle
the shortest; first peraeopod usually long; third and fourth pairs in female
with rudimentary exopods; pleopods of male with short rami; exopod of
uropods generally shorter than the endopod.
11 species.
Synopsis of species:
1 | Uropods with exopod not shorter than the endopod. . 1. L., mancus
\ Uropods with exopod shorter than the endopod — 2.
{ Pedigerous segments 1, 2, emarginate in front — 3.
* | Pedigerous segments 1, 2, not emarginate in front — 5.
3 { Lateral ridges enclosing cordiform area on frontofcarapace 2. L. antipus
| Carapace without such ridges — 4.
4 J Carapace sparsely hirsute, lower margin acutely serrate 3. L. ampullaceus
\ Carapace closely hirsute, lower margin squarely serrate 4. L. villosus
5 | Uropods with 1st joint of endopod equal to the 34 — 6.
\ Uropods with Ist and 34 joints of endopod unequal — 7.
Carapace about thrice as long as pedigerous segments
‘a Commoincda teste eee airs teu ee) (One Crassicauds
: | Carapace not nearly thrice as long as pedigerous segments
combined Sa eee ee ee es loos dani Dorealis
7 J Uropods with 1st joint of endopod shorter than 34 — 8.
) Uropods with 1st joint of endopod longer than 34 — 9.
g J Carapace with lower margin acutely serrate .... . 7. L.macrurus
) Carapace with lower margin squarely serrate... . . 8. L. macruroides
on Uropods with exopod not longer than 1st joint ofendcopod 9. L. productus
“| Uropods with exopod much longer than 1st joint of endopod — 10.
Uropods with 1st joint of endopod at least as long as
Ondvandead combineds s.. «5. ser em = . 10. L. longimanus
| Uropods with 1st joint of endopod little longer than
Ur ONIEN tos co. ee fl ke os eens del nae eee ow! oh od Dy eraeiis
10
1. L. mancus O. Sars 1873 LZ. manca, G. O. Sars in: Svenska Ak. Handl., n. ser.
v.ll vr. 5 p.21 t.5 f.24-28 | 1902 L. m., Zimmer in: Ergeb. Hamburg. Magalh.-
Sammelr., v2 Cumac. p.8 tf. 11—14 | 1905 Z. m. Calman in: Sci. Invest. Fish.
Ireland; 1904 app.k p.40 | 1908 Diastylis‘m., Zimmer in: Ergeb. ‘Tiefsee Exp., v.8
p. 183 | 1912 Leptostylis mancus, T. Stebbing in: Ann. S, Afr. Mus., v. 10v p. 153.
Pseudorostral lobes shortly and bluntly produced, forming no distinct
antero-lateral corner. Carapace roughly pentagonal in dorsal aspect, antero-
lateral margin denticulate; with denticles crossing the frontal lobe and the
pseudorostral lobes, from which on either side a subdorsal denticulate crest
124 Cumacea: 11. Diastylidae, 7. Leptostylis
ascends surrounding a slightly impressed space with arched centre. Pedigerous
segments 1, 2 short, conspicuously narrower than the carapace or the next
2 segments, 5% with postero-lateral corners little extended, blunt. Telson
as long as 6% pleon segment, with 1 pair of lateral spines in juv., 2 pairs
in adult. Antenna 1 short, 2"4 joint of peduncle shorter than 3% Peraeo-
pod 1 long and strong, last 3 joints of uniform length. Peraeopods 3, 4
in © juv. with exopods 1-jointed, narrowly conical. Peraeopod 5 well deve-
loped in adult, the specific name alluding to the absence of these limbs
being no doubt due to observation of a young specimen. Pedunele of uro-
pods considerably longer than the telson, with 3 spines on medial margin
of young, 9 on that of adult, rami equal, rather shorter than the telson,
endopod with a single spine on each joint at end of the medial margin
which in the adult has a continuous brush of fine hairs. L. © adult
10°5 mm, © juv. (uropods included) 7°5 mm.
S. Atlantic, off mouth of R, La Plata, depth 98 m.
2. L. antipus Zimmer 1907 ZL. a., Zimmer in: Zool. Anz., v.31 p. 223 | 1909
L.a., Zimmer in: Ergeb. Schwed. Siidp.-Exp., v.6 nr. 3 p.22 t.7 f. 119—127.
Hirsute. Pseudorostral lobes shortly and bluntly produced; commen-
cing on these and running backwards a pair of carinae enclose a cordiform
space on the front of the carapace, of which the hind margin has a sharp
carina, continued some way forwards on the sides. Eyelobe minute. Pedi-
gerous segments 1, 2, produced forwards in a submedian pair of tooth-like
processes. Telson rather shorter than 6 pleon segment, with 2 pairs of
lateral setae on the abruptly narrowed section and a pair of spines on the
anal valves. Antenna 1, 2™¢ joint shortest, 3¢ longest, both flagella 3-jointed.
Maxilliped 3, 24 joint nearly twice as long as rest of limb, 3¢ slightly wider
than 4th, Peraeopod 1 very long, 6% joint longer than ot, 5% than 7%.
Peraeopod 2, 7 joint nearly as long as 5%. Peraeopods 3, 4, with exopods
rudimentary. Peduncle of uropods more than thrice as long as the telson,
and twice as long as the exopod, with 9 spines on the medial margin,
endopod 2/,48 as long as the peduncle, 18* joint much longer than 2™4, 2Qn¢
a little longer than 34, each with 2 spines on the medial margin. Colour
faintly yellowish white. L. @ 4—6 mm. Adult 3 unknown.
S. Georgia, ¢ juv. at 75m; 9 at 12—15m and 252—310 m.
3. L. ampullaceus (Lilj.) 1855 Cuma ampullacea, W. Liljeborg in: Ofv. Ak.
Forh., v.12 p.120 | 1865 Diastylis a., G.O. Sars in: Forh. Selsk. Christian., 1864 p.175 |
1866 D.a., G. O. Sars in: Nyt Mag. Naturv., v.15 p.105 | 1869 Leptostylis a., G. O.
Sars in: Nyt Mag. Naturv., v.16 p.344 | 1877 Z. a., Meinert in: Naturh. Tidsskr., ser. 3
v.11 p.178 | 1879 ZL. ampullaceus, S. 1. Smith in: Tr. Connect. Ac.. v.51 p.114 | 1881
L. ampulacea, Meinert in: Naturh. 'Tidsskr., ser.3 v.12 p.497 | 1894 L. ampullaceus,
A. M. Norman in: Ann. nat. Hist., ser. 6 0.13 p. 277 | 1896 Z. ampullacea, G. O. Sars in:
Norske Nordhavs-Exp., v.6 nr.15 p:24 |.1900 ZL. a.,-G. O. Sars, Crust. Norway, v.3 p.70
t.50 f.1 | 1900 ZL. a., Zimmer in: Fauna arctica, v1 p.427, 434 (with synonymy) | 1908
L.a., Zimmer in: Ergeb. 'Tiefsee-Exp., v.8 p. 182.
Pseudorostral lobes shortly and not sharply produced, antero-lateral
corners rounded off and coarsely serrate. Carapace broad, with scattered
hairs, eyelobe minute. Pedigerous segments 1, 2, produced forwards in a
submedian pair of tooth-like processes. Telson shorter than 6%" pleon seg-
ment, with 1 pair of lateral spinules. Antenna 1, 3¢ joint rather shorter
than 1st, much longer than 2™¢, principal flagellum 3-jointed, accessory
2-jointed. Peraeopod 1 not very long, 2"4 joint not so long as rest of limb,
Cumacea: 11. Diastylidae, 7, Leptostylis 125
6t2 longer than 5‘ which is more than twice the 7. Peraeopod 2, 7" joint
nearly as long as 5, not twice the 6". Rudimentary exopods of 3¢ and
4th pairs very small, 2-jointed. Peduncle of uropods about twice as long
as the telson, not much longer than the endopod, which has 1s joint nearly
as long as the subequal 2"¢ and 34 combined, spines on the medial margin
respectively 3, 1, 1; exopod about 2/378 as long as endopod. Colour dark
yellowish grey or fuscous. L. adult @ 6 mm.
Whole coast of Norway, depth 38—188m; Rédberg, 457—549m; Sweden and
Denmark, 15—30 m; Gulf of Maine, 95—165 m.
4. L. villosus 0. Sars 1869 L. villosa, G. O. Sars in: Nyt Mag. Naturv., v.16
p. 344 | 1872 L. v., G. O. Sars in: Forh. Selsk. Christian., 1871 p.270 | 1900 L. v., G.O
Sars, Crust. Norway, v.3 p.71, 108 t.50 f.2 | 1900 ZL. v., Zimmer in: Fauna arctica, v. 1
p. 427.435 | 1901 Z.v., T. Scott in: Rep. Fish. Board Scotl., v. 19 p.275 t.18 £41 |
1908 L. v., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p.182 | 1912 L. villosus, T. Stebbing
in: Ann. S. Afr. Mus., v. 10v p. 153.
Densely hirsute. Pseudorostral lobes shortly and bluntly produced, the steep
sides and rounded lower border not acutely serrate but cut into square-
topped divisions, machicolated. Carapace large and deep, eyelobe minute.
Pedigerous segments produced forwards in a submedian pair of tooth-like
processes. Pleon very slender. Telson shorter than the 6% pleon segment,
with 1 pair of lateral spinules. Antenna 1, 3¢ joint as long as 1st, longer
than 2™4, Peraeopod 1 elongate, 2™4 joint about 2/;"s of the total length of
the limb, 6% joint much longer than 5% which is fully twice the 7".
Peraeopod 2, 7 joint longer than 5, Rudimentary exopod of 3% and Ao
pairs extremely small, 2-jointed. Peduncle of uropods more than twice as
long as the telson, but little longer than the endopod, of which the 1*' joint is
not nearly equal to the 2"4 and 34 combined, 3¢ rather longer than 2nd, spines
of medial margin respectively 2, 1, 1, exopod a little longer than first 2 joints
of endopod. Colour light fuscous. L. O 4mm. GC unknown.
Norwegian coast, depth 113—188m; Clyde, 73 m; S. of Shetland Isles, 110—120 m.
5. L. crassicauda Zimmer 1907 L.c., Zimmer in: Zool. Anz., v.31 p. 223.
Sparsely hairy. Carapace inflated, about thrice as long as the steeply
sloping group of pedigerous segments, antero-lateral margin denticulate.
Telson about as long as 6 pleon segment, with 1 pair of lateral spines.
Peraeopod 1 long, 5 and 6* joints subequal, 7% shorter than either.
Pedunele of uropods nearly twice as long as the telson, with 6 spines on
the medial margin; endopod longer than the exopod, a little shorter than
the peduncle, 15¢ joint longer than 24, subequal to 34, spines on medial
0]
margin respectively 2, 1, 1. Colour white. L. d juv. about 5°5 mm.
Winter Station, German South Pole Exp. (lat. 66° 2’ S. long 89° 38° E.)
6. L. borealis Stappers 1908 ZL. b., Stappers in: Arch. Zool. expér., ser. 4 v.8
notes p. 102 tf.4,5 | 1911 Z. b., Stappers in: Camp. arct. Orléans, Crust. Malac. p. 120
t.7 | 1912 L. b., T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 153.
Somewhat hirsute. Pseudorostral lobes shortly and obtusely produced,
slightly upturned. Carapace much less than thrice as long as the group of
pedigerous segments. Pleon slender, telson shorter than 6t segment, with
1 pair of lateral spines. Antenna 1, 3¢ joint shorter than 1°‘, longer than aoe
flagella very unequal, both 3-jointed. Peraeopod | elongate, 224 joint much
shorter than rest of limb, 5t* about 3/,S of 6 and nearly twice 7. Peraeo-
pod 2, 7t joint nearly equal to 5" and 6% combined. Exopods of 34 and
126 Cumacea: 11. Diastylidae, .7. Leptostylis
4th pairs minute, 2-jointed. Peduncle of uropods rather more than twice
as long as the telson and a little longer than the endopod,: with 7 or
8 spines on the medial margin; endopod with 1S* joint longer than 24, equal
to 34, spines on medial margin respectively 2, 1, 1; exopod shorter than
endopod, but reaching beyond its 2™¢ joint. L.Q 45 mm.
Off S. coast of Nova Zembla, depth 90 m.
7. L. macrurus O. Sars 1869 L. macrura, G.O. Sars in: Nyt Mag. Naturv.,
v.16 p.344 | 1870 L. m., G. O. Sars in: Forh. Selsk. Christian., 1869 p. 156 | 1886 L. m.,
G. O. Sars in: Norske Nordhavs-Exp., v.6 nr. 15 p.24 | 1900 L. m., G. O. Sars, Crust.
Norway, v.3 p.69 t.49 | 1900 Z. m., Zimmer in: Fauna arctica, v.1 p.427, 4385 | 1908
L.m., Gimmer in: Ergeb. Tiefsee-Exp., v.8 p. 182 | 1912 L. macrurus, T. Stebbing in:
Ann. 8. Afr. Mus., v. 10v p. 153, 154.
Pseudorostral lobes short, a little upturned. Carapace large, hirsute,
more than twice as long as the group of pedigerous segments, in the &
having a pair of very conspicuous, coarsely serrated, infero-lateral carinae.
Pleon long and slender, 5** segment very long, twice as long as the 6%,
telson shorter than the 6, with 1 pair of lateral spinules, the apical spines
long. Antenna 1 Q, 1%* joint longer than 34, 3¢ than 24, flagella 5-jointed,
principal flagellum in GO 4-jointed, surrounded by brush of setae. Antenna
2 3, 5 joint scarcely twice as long as 2"4, Peraeopod 1, 24 joint scarcely
as long as 6% and 7 joints combined, 5t* rather more than twice the 7%,
6th thrice. Peraeopod 2, 5 joint a little longer than the 7, Exopods of
34 and 4 pairs in Q minute, 2-jointed. Pleopod 1 with peduncle distally
narrowed. Peduncle of uropods as long as 5% pleon segment, with 9 spines
on medial margin; endopod about 2/,4° of peduncle,
1st joint longer than 24, shorter than 34, spines
respectively in © 2, 1, 2,in G 4, 2, 3; exopod reaching
middle of 34 joint of endopod. Colour greyish white.
L. about 5 mm.
Coast} of Norway, depth 75—470 m; cold area of N.
Atlantic 989 m.
8. L. macruroides Stebb. 1912 L. m., T. Stebbing
in: Ann. S. Afr. Mus., v. 10v p. 153 t. 56.
Near to /. macrurus, but with the lower margin
and the infero-lateral carinae of the carapace in the
3 machicolated in the anterior half after the fashion
of L. villosus (nr. 4); hind margin a little upturned,
some of the pedigerous segments and first 2 of the
pleon with latero-ventral spines, 5‘ pleon segment not
quite twice as long as the 6%, to which the telson
Fig. 77. is equal, with its distal section more narrowed, but
1. macruroides (3) Steb- similarly armed as in L. maecrurus. Antenna 1, principal
ing, antenna 1. : v6 ae 5
flagellum 5-jointed, accessory 4-jointed, in each the ter-
minal joint minute. Antenna 2, 5'" joint nearly thrice as
long as 2"4, Peraeopod 1, 2Ӣ joint long, strongly bent. Peraeopod 2, 5% joint
decidedly longer than 7%, Pleopods with peduncle parallel-sided. Peduncle of
the uropods about twice and a half as long as the telson, but much less, than
twice the endopod, with 11 spines on the medial margin; endopod’s 1*
joint longer than 2™4, shorter than 34, spines on medial margin respectively 4,
3, 2. L. adult O about 5 mm, © unknown.
S. Africa, Cape Natal N. by E. 24 miles, depth 805 m.
I
Cumacea: 11. Diastylidae, 7. Leptostylis 12
9. L. productus Norm. 1879 L. producta, A. M. Norman in: Ann. nat. Hist.,
ser.5 v.3 p.65 | 1912 L. productus, T. Stebbing in: Ann. S, Afr. Mus., v. 10v p. 153.
Pseudorostral lobes short, blunt, slightly upturned. Carapace short, as
broad as long, nearly smooth, antero-lateral margin strongly serrate. Telson
not longer than 6 pleon segment, nor more than half the 5, without
lateral spines, apical pair rather large. Peduncle of uropods nearly thrice
as long as the telson, with 4 spaced spines on the medial margin; endopod
nearly as long as peduncle, with spines on its 3 joints respectively 2, 1, 0,
exopod as long as 1S* joint of endopod, scarcely longer than the lanceo-
fate 35.) Sex? ©.
Off Butt of Lewis, lat. 59° 41‘ N., long. 7° 34‘ W., depth 838 m.
10. L. longimanus (O. Sars) 1865 Diastylis 1., G. O. Sars in: Forh. Selsk.
Christian., 1864 p.173 | 1869 Leptostylis l., G. O. Sars in: Nyt Mag. Naturv., v.16
p. 343 | 1879 Z.1, S. I. Smith in: Tr. Connect. Ac., v.51 p.114 | 1900 Z, longimana,
G. O. Sars, Crust. Norway, v.3 p.68 t.48 | 1900 Z. 1., Zimmer in: Fauna arctica, v. 1
p. 427, 435 | 1905 D. 1., Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p. 10,48 |
1908 L.1., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 182 | 1912 L. longimanus, T. Stebbing
in: Ann. S. Afr. Mus., vu. 10v p. 153.
Fig. 79.
L. longimanus
(Sars), pleopod 1.
Fig. 80.
Fig. 78. L. longimanus
L. longimanus (Sars), peraeopod 1. (Sars), pleopod 2.
Pseudorostral lobes acute, a little upturned. Carapace large and deep,
nearly smooth, antero-lateral margin minutely denticulate. Pleon slender,
128 Cumacea: 11. Diastylidae, 7. Leptostylis
telson shorter than the 6 segment, with 1 pair of lateral spinules. Antenna
1 of © long, 1%¢ joint longer than 24, subequal to 34, accessory flagellum
much smaller in Q than in G. Antenna 2 in d, 2"4 joint subequal to 5%.
Peraeopod 1 very long, but 2™4 joint scarcely longer than 5‘, much shorter
than 6%, 6t more than thrice the 7. Peraeopod 2, 7 joint longer than 5%,
Exopods of 34 and 4* pairs in Q 2-jointed. Pleopods with the short 1-jointed
rami nearly equal. Peduncle of uropods about equal the 5% and 6 pleon
segments combined, longer than the endopod, in which the 1S* joint is longer
than 2"¢ and 34 combined, spines on the medial margin respectively 3, 1, 1
in Q, 5, 1, 1 in dG, exopod equal to first 2 joints combined. Colour whitish.
L. 9 5°55 mm, ¢C about 5 mm.
Coast of Norway, depth 56—188 m; N. America, Casco’ Bay.
11. L. gracilis Stappers 1908 ZL. g., Stappers in: Arch. Zool. expér., ser. 4 v8
notes p.100 tf.1—3 | 1911 Z. g., Stappers in: Camp. arct. Orléans, Crust. Malae. p.116
t.5,6 | 1912 LZ. g., T. Stebbing in: Ann. S. Afr. Mus., v 10v p. 153
Very near to L. longimanus, but peraeopod 1 with 2™¢ joint longer
than 5th, 6th, and 7% joints combined, 6 twice and a half as long as the 7%;
Fig. 82.
L-gracilis Stappers, part of left
mandible.
Fig. 83.
Fig. 81. L. gracilis Stappers, half
L. gracilis (¢) Stappers. of lower lip.
lay 4
peraeopod 2 with 7 joint as long as 5; pleopods with the small rami
very unequal, the shorter exopod being figured as 2-jointed instead of
l-jointed; in the uropods the 1* joint of the endopod is much shorter than
the 224 and 34 joints combined, but longer than the 34, the 284 being described
Cumacea: 11. Diastylidae, 7. Leptostylis, 8. Ekleptostylis, 12. Colurostylidae 129
as the shortest, though not so figured, spines on the medial margin respect-
ively 2. 1. 1, the exopod a little longer than first 2 joints of endopod combined.
L. d 6mm, 9 unknown.
Off S. of Nova Zembla, depth 90 m.
8. Gen. Ekleptostylis Stebbing
1912 Hkleptostylis, T. Stebbing in: Ann. 8. Afr. Mus., v. 10v p. 146, 153.
Near to Leptostylis (p. 123), but the telson with many lateral spines and
in the male having a lobe broadly laminar produced over the base of the
terminal section and the anal valves each ending in a little tooth, and
peduncle of first antennae with its three joints of equal length.
1 species.
1. E. walkeri (Calm.) 1907 Leptostylis w., Calman in: Bull. Mus. Paris, p. 2,
7 tf.6—9 | 1911 ZL. w., Calman in: Tr. zool. Soc. London, v.181v p.366 | 1912 Ekleptostylis
w., I. Stebbing in: Ann. 8S. Afr. Mus., v. 10v p. 153.
beside generic differences pseudorostral lobes a little more salient, teeth of
lower margin of carapace less distinctly
Laer / We
longitudinal denticulate carina on each side
as in Leptostylis macrurus (p. 126). The telson
lateral spines, with an unusuaily conspicuous
setule issuing from the side of each spine; in the C the spines are larger but only
uropods nearly twice and a half as long as the telson, with some 23 spines
on the medial margin; endopod a little longer than exopod, 2/345 as long as
General form and proportions as in Leptostylis villosus (p. 125), but
squared, and its surface carrying only some
long scattered setae, that of the d having a
is about 4/,"s of the length of the 6% pleon Fig. 84.
seomment, in the ©. having about! 14. pairs of . ™ walkert (¢) (Calman), telson:
5 pairs with smaller setules. Peraeopod 1, 24 joint less than half the total Jength
of the limb, 5 joint 2/,4% of 6*®, 6 thrice as long as the 7%. Peduncle of
the peduncle, 1 joint nearly twice 2™4, a little longer than 34, spines on
medial margin respectively 7, 2, 1. L. ovigerous QO 6°5 mm.
Bay of Biseay, depth probably 100 m.
12. Fam. Colurostylidae
1912 Colurostylidae, T. Stebbing in: Ann. S. Afr. Mus., v. L0v p. 143, 146.
Side-plates of pedigerous segment 3 produced backwards. Telson small,
blunt, without apical spines. Antenna 1 with accessory flagellum small in
male, very small in female. Antenna 2 of female 3- or 4-jointed. Palp
of maxilla 1 ending in 2 filaments. Peraeopods 1-4 with exopods in both
sexes. but those of the 3¢ and 4t® pairs in the female rudimentary. Only
2 pairs of pleopods in the male. Endopod of uropods 2- or 3-jointed.
2 genera.
Synopsis of genera:
Endopod of uropods 2-jointed ......-+-.- . 1. Gen. Colurostylis
Endopod of uropods 3-jointed ....... +... & Gen. Anchicolurus
Das Tierreich. 39. Lief.: T. R. R. Stebbing, Cumacea. 9
130 Cumacea: 12. Colurostylidae, 1. Colurostylis, 2. Anchicolurus
1. Gen. Colurostylis Calman
1911 Colurostylis, Calman in: Tr. zool. Soc. London, v. 181v p. 342,376 | 1912
C., T. Stebbing in: Ann. 8. Afr. Mus., v. 10v p. 146.
Side-plates of pedigerous segments 3 and 4 produced backwards. An-
tenna 2 of female 3-jointed. Maxilliped 3 with apex of 2"¢ joint a little
produced. Peraeopod 2 with 7 joint longer than 6%. Endopod of uro-
pods 2-jointed.
1 species.
1. C. pseudocuma Calm. 1911 C.p., Calman in: Tr. zool. Soe. London, v. 181v
p. 342, 376 t. 36 f.23—36 | 1912 C.p., Calman in: P. U.S. Mus,, v.41 p.674 | 1912
C. p., T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 146.
Pseudorostral lobes straight, acute, scarcely raised, sinus obsolete, antero-
lateral corner rounded. Carapace pitted. Backward curvature of the side-
plates of 34 pedigerous segment in © separates 3¢ peraeopods widely from
the 2™4. Length of telson about 2/34° of preceding segment in Oo, 3/48 in OC.
Kye without pigment. Antenna 1 in © with 3-jointed flagella, the accessory
very small, in the G the 34 joint of peduncle as broad as long, with brush
of sensory filaments, the flagella 4-jointed, the accessory not quite so small
as in the @. Antenna 2 of © 3-jointed, flagellum in CO elongate. Mouth-
parts resembling those in Pseudocuma (p. 148). Mandible with about 9 spines
in spine-row. Mavxilliped 3 with about 4 plumose setae on the expanded
and slightly produced apex of the 2"4 joint. Peraeopod 2 with 7 joint
longer than 6%, but not twice as long. Exopods on peraeopods 3 and 4
of © 2-jointed, attached near the base of the 24 joint and nearly half its
length. Rami of pleopods very small. Peduncle of uropods in Q more
than twice, in CO nearly thrice, as long as 6% pleon segment, in the O a
little, in the G much, longer than the subequal rami, of which the endopod
has a first joint not quite so long as the 2°, both fringed with setae or —
spines. L. O 233mm, CG 2:7 mm.
New Zealand, Lyttleton Harbour, depth 2—9 m; Akaroa Harbour, depth 11 m
2. Gen. Anchicolurus T. Stebbing
1912 Colurostylis (2), Calman in: P. U.S. Mus., v.41 p.605, 670 | 1912 Anchicolurus,
T. Stebbing in: Ann. S. Afr. Mus., v. 10v p.176.
Side-plates of pedigerous segment 3 produced backwards. Antenna 2
of female 4-jointed. Maxilliped 3 with apex of 2™¢ joint expanded but not
produced. Peraeopod 2 with 6 joint longer than seventh. Endopod of
uropods 3-jointed.
1 species.
1. A. occidentalis (Calm.) 1912 Colurostylis (2) occidentalis, Calman in: P. U.
S. Mus., v.41 p.605, 670 tf. 100—112 | 1912 Anchicolurus o., T. Stebbing in: Ann. S. Afr.
Mus., v.10v p.176.
Pseudorostral lobes meeting in front of the small eyelobe, antero-
lateral corners acute in both sexes. Carapace with 4 pairs of oblique ridges.
1st pedigerous segment only dorsally exposed, 22¢ with side-plates greatly
expanded and marked by an incision of the front margin, those of the 34
segment overlapping the 2™4 side-plates above and produced backward below,
32 and 4% segments with medio-dorsal carina, side-plates of 4% marked by
Cumacea: 12. Colurostylidae, 2. Anchicolurus, Genus incertae sedis 131
a suture, 5 with serrate front margin and dorso-lateral carinae extended
into acute postero-lateral teeth. Pleon segments 1 and 2 in dO with mid-
ventral backward pointing tooth and segments 3—5 each with a pair of
ventral articulated spines. Telson carrying distal
setae and slightly longer than broad in 3, but
not longer and unarmed in ©. Eye without
pigment in either sex, larger in the O, with
3 lenses in the o. Antenna 1 with 274 joint
of peduncle the shortest, flagellum 4-jointed in
©, 5-jointed in G, accessory 3-jointed, much
smaller in Q than in G. Antenna 2 of > 4-jointed,
32 joint large, ovate, serrated, 4 minute, flagel-
lum in GO elongate. Maxilliped 3 with apex of 2¢
joint expanded, not produced, carrying plumose
setae, 3¢ joint with comb-like
row of spinules. Peraeopod 1
with long 2"4 joint, distally
narrow; peraeopod 2 with 6%
joint longer than seventh, very
unusual in this pair; 34 and
4th pairs with 2-jointed exo-
: ; Fig. 86.
Fig. 85. pod In 2, conspicuous though A. occidentalis (Calman), telson
A. occidentalis(?)(Cal- Small. Peduncle of uropods ana uropod, » = tip of endopod of
man), antenna 2. not quite twice as long as the uropod (more magnified).
exopod; endopod about 3/4*s
as long as the exopod, 3-jointed, 1%* joint nearly 4 times as long as the
other 2 combined, exclusive of the long spiniform process produced far
backward from the lower surface of the 34. L. © 12mm or more, d
11:8 mm.
Off Oregon, lat. 43° 01‘ 00” N., long. 124° 30’ 30” W., depth 64 m; Pacific
Grove, California; Monterey-Bay, California, depth 16 m.
Genus incertae sedis, near Colurostylidae
Gen. Pachystylis H. J. Hansen
1895 Pachystylis, H. J. Hansen in: Ergeb. Plankton-Exp., v.2 G.c. p.58 | 1908
P., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 169, 181.
General form as in Diastylis (p. 89) but telson short, cylindrical with-
out apical spines. Eye wanting. Antenna 1 with accessory flagellum short.
Mandible with strong molar, few spines in the spine-row on the short distal
section. Maxilliped 3 with second joint much dilated distally, third joint
short but broader than the fourth. First four peraeopods in the male, only
the first two pairs in the female, carrying exopods. Endopod of uropods three-
jointed. (Pleopods of male unkown).
1 species.
P. rotundatus H. J. Hansen 1895 P. rotundata, H. J. Hansen in: Ergeb. Plank-
ton-Exp., v.2 G.c. p.58 t.7 f. 2—2e.
Pseudorostral lobes moderately produced in front of the eyeless eye-
lobe; antero-lateral corners acute in Q, rounded in dG. Antenna 1, principal
9*
132 Cumacea: Genus incertae sedis, 13. Oxyurostylidae, 1. Oxyurostylis
flagellum 3-jointed in Q, 2-jointed in CG juv., accessory 3-jointed, 18* and
32 joints very small. Mavxilliped 3, 2°¢ joint with spine-like process at medial
apex of 24 joint, stronger in © than in G.
Telson in d a little longer than the preceding
segment, length scarcely twice the breadth, evenly
cylindric, end truncate; in © shorter than pre-
ceding segment, narrowing before the middle to
a rounded apex carrying about 4 setules. Peduncle
of the uropods much longer than the telson, and
in the © than the rami, but in the d only a
little longer than the exopod, which is only a
little longer than the endopod; 1st jomt of
endopod not so long as the equal 274 and 34
joints combined, each of the three having a
single lateral spine, the 3¢ a long apical one.
Colour greyish white. L. 9 23 mm, dc juv.
Fig. 87. 18 mm.
P. rotundatus (Jd juv.) Hansen, 3 ;
telson ena Mouth of R. Tocantins, Brazil.
13. Fam. Oxyurostylidae
1912 Oxyurostylidae, T. Stebbing in: Ann. S, Afr. Mus., v. 10v_ p. 133, 146.
Telson large, acute, without apical spines. Antenna 1 with accessory
flagellum small. Antenna 2 of © 4-jointed. Palp of maxilla 1 ending in
2 filaments. Peraeopods 1—4 with exopods in both sexes, but those of the
34 and 4‘) pairs in the female very rudimentary. Only 2 pairs of pleopods
in the male. Endopod of uropods 3-jointed.
1 genus.
1. Gen. Oxyurostylis Calman
1912 Oxyurostylis, Calman in: P. U.S. Mus., v.41 p.605, 666 | 1912 O., T. Stebbing
in: Ann. S. Afr. Mus., v.10v p. 146.
Telson with lateral spines along the narrow distal section. Maxilliped
3 with 24 joint distally widened and produced, 3¢ narrowly produced, 4%
neither expanded nor produced; with exopod in both sexes.
Endopod of the uropods longer than the exopod in the male.
1 species.
1. O. smithi Calm. 1912 0. s., Calman in: P. U.S. Mus.,
v.41 p.605, 667 tf.91—99 | 1912 O.s., T. Stebbing in: Ann. 8.
Afr. Mus., v. 10v p. 146.
Pseudorostral lobes meeting horizontally in front
of the short broad eyelobe, sinus obsolete, antero-lateral
corner not produced. Carapace having the frontal lobe
crossed by 2 prominent transverse ridges, followed in the
© by 2 others much fainter; these are flanked by a pair
of short sinuous ridges, behind which are 2 parallel pairs
of large oblique ridges. All 5 pedigerous segments distinct, Oana
side-plates of 34 extended backwards in the ©, hind maxilliped ‘3.
corners of 5 extended slightly and bluntly in Q, strongly
and acutely in 3. Telson about once and a half the length of the preceding seg-
Cum.: 18. Oxyurostylidae, 1. Oxyurostylis, 14. Pseudodiastylidae, 1. Pseudodiastylis 133
ment in ©, twice its length in G, lateral spines in 6 or 7 pairs, unspined apex
acutely upturned. Eye without pigment, visual elements indistinct. Antenna | ©
with 2"4 joint of peduncle the shortest, principal flagellum 5-jointed in Q with
3-jointed accessory, in G the principal 5- or 6-jointed, with accessory 4-jointed.
Peraeopod 2 with the last 2 joints together not quite equal to the preceding
joint. Exopods on peraeopods 3 and 4 of Q 2-jointed but minute. Peduncle
of uropods scarcely longer than the telson, but longer than the rami, which
are subequal in O, in the G the endopod the longer, armed with many
more spines than in the Q,its 1st joint longer than either of the 2 following
joints. L. Oo 66mm, CO 7:3 mm.
U.S. America, Casco Bay; Vineyard Sound, Woods Hole, surface; Punta Rassa,
Florida, depth 2m; Calcasieu Pass, Louisiana.
14. Fam. Pseudodiastylidae
1912 Pseudodiastylidae, T. Stebbing in: Ann. S, Afr, Mus., v. 10v_ p. 138, 146.
Telson long and narrow, with more than 2 apical spies. Antenna |
large, with very small accessory flagellum. Antenna 2 of female with slender
terminal joint. Maxilliped 3 resembling peraeopod 1. Peraeopods 1—4 of
female with exopods, those of the 3¢ and 4 pairs rudimentary. Uropods
elongate. Male unknown.
1 genus.
1. Gen. Pseudodiastylis Calman
1905 Pseudodiastylis (Sp. un.: P. ferox), Calman in: Siboga-Exp., nr. 36 p. 1,16 |
1908 P., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 169,170,173 | 1912 P., T. Stebbing
in: Ann. S. Afr. Mus., v. 10v_ p. 146.
Pseudorostral lobes greatly produced. Carapace inflated. Telson narrowly
elongate, with three apical spines. Eye wanting. First antenna very large,
flagellum elongate, accessory three-jointed but minute. Second antenna of
female with terminal joint long, stiliform. Third maxilliped leg-like. Third
and fourth peraeopods of female with vestigial exopods.
1 species.
1. P. ferox Calm. 1905 P. f., Calman in: Siboga-Exp.. nr. 36 p.16 t.2 f.15—25 |
1908 P. f., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p.175 | 1912 P. f., T. Stebbing in: Ann.
S. Afr. Mus., vu. 10v p. 146.
Integument armed with numerous spicules and spines, many of those
on the carapace being especially conspicuous and of complicated structure.
Pseudorostral lobes directed slightly upwards obliquely, curved, apically acute,
the production about one-third of the total length of the carapace; antero-
lateral corners spinose, slightly produced. Carapace oval, dorsal outline
strongly arched. The eyeless ocular lobe minute. First pedigerous segment
short but completely exposed; second and third segments seemingly immovably
united. Telson including the terminal spine nearly as long as the 3 preceding
segments combined; except at the base over the anal orifice its width little
over a twentieth of its length; 3 lateral pairs of spines are followed by
3 apical spines, of which the central is fully twice as long as the other 2.
Antenna 1 about three-fourths as long as the carapace, almost pediform,
1st joint equal to 2.¢and 34 combined, a little shorter than the flagellum,
of which the 18* joint is long, the 5t* minute; the 3-jointed accessory scarcely
one-sixth as long as the 1S* joint of the principal. Maxilliped 3 a little
134 Cumacea: 14. Pseudodiastylidae, 1. Pseudodiastylis
shorter than peraeopod 1 and with a more strongly armed 5* joint, but —
otherwise in close resemblance. Peraeopod 2 with shorter 274, longer 5%
joint than in the preceding pair and 6 joint only half as long as the
Eig. 89. Fig. 90.
P. ferox (Q juv.) Calman. P. ferox Calman, telson.
straight needle-like 7. In the 3¢ and 4* pairs the exopod is 2-jointed,
but extremely minute. Peduncle of uropods a fourth longer than the telson;
the rami broken, but the remains indicating a long and slender character.
L. Q with very small oostegites 14 mm.
Malayan Archipelago: Lat. 5° 40'.7S., long. 120° 45'.5K., depth 1158 m.
Cumacea: 15. Ekdiastylidae, 1. Ekdiastylis 135
15. Fam. Ekdiastylidae
1912 Ekdiastylidae, T. Stebbing in: Ann. 8. Afr. Mus., v. 10v_p. 183, 146, 155,
Telson large, with only 2 apical spines. Antenna 1 with accessory
flagellum small. Antenna 2 of female with no elongate joint; flagellum of
male very long. Peraeopods 1—4 in male provided with exopods, but only
the first two pairs in the female. Only two pairs of pleopods in the male.
Endopod of uropods 2-jointed.
1 genus.
1. Gen. Ekdiastylis T. Stedbing
1912 Ekdiastylis, T. Stebbing in: Ann. S, Afr. Mus., v. 10v_ p. 146, 155,
With the characters of the Family. The species E. insignis, E. argen-
talus, E. abbreviatus, E. fimbriatus, appear te be in close alliance, with con-
siderable sexual differences.
9 species.
Synopsis of species:
Carapace strongly sculptured — 2.
Carapace not strongly sculptured — 3.
Peraeopod 2 with 5th joint scarcely as long as 6th and 7th
Combinedtyecuc cite gis 6) 6, «iene eye tis te «+, Ly fe SCUlptuaty
Peraeopod 2 with 5th joint much longer than 6th and 7th
combined .... . 5 Sao oO GOua eee ees 2 epOlitus
—
eee
Distal half of telson with fewer than 3 pairs of lateral
. 3 spines — 4.
Distal half of telson with pairs of lateral spines not fewer
than 3 — 5.
Carapace with pair of infero-lateral processes near the front 3. KE. insignis
A C : ;
“arapace without such processes .....-.+...-... 4. H.mystacinus
Carapace with 3 pairs of lateral processes... .- . 5. BE. hexaceros
Carapace with fewer than 3 pairs of lateral processes — 6.
Carapace with single pair of antero-lateral processes . . . 6. H. argentatus
Carapace without lateral processes — 7.
Carapace coarsely spinose . .....-+..++-+- ee %. EB horridus
Carapace not coarsely spinose — 8.
Telson of 9 with 5 pairs of lateral spines. . ...... 8 EH. abbreviatus
Telson of Q with 3 pairs of lateral spines. . .. +... 9, E. fimbriatus
or
poe eee ee se
1. E. sculptus (0. Sars) 1871 Diastylis sculpta, G. O. Sars in: Ofv. Ak. Férh.,
v.28 nr.1 p.71 | 1871 D.s., G. O. Sars in: Svenska Ak. Handl.,n. ser. v.9 nr. 13 p.24
t.1—9 f.1—49 | 1873 D.s., S.J. Smith in: A. E. Verrill in: Rep. U. 8. Fish. Comm.,
-y, 1 p. 554] 1874 D. s., S. I. Smith & Harger in: Tr. Connect. Ac., v.31 p.29 | 1879
D. sculptus, S. 1. Smith in: Tr. Connect. Ac., v.51 p.111 | 1900 D. sculpta, Zimmer in:
Fauna arctica, v1 p.425 | 1912 D.s., Calman in: P. U.S. Mus., v.41 p. 605, 657 | 1912
Ekdiastylis sculptus, T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 146, 155.
Pseudorostral lobes shortly produced in front of the prominent eyelobe,
horizontal, antero-lateral corners obtuse. Lower margin of carapace sinuous,
finely denticulate in front, hind margin carinate, frontal lobe crossed by 2
straight carinae, from the hinder a short median longitudinal carina passes to
join the foremost of 3 pairs of oblique curved lateral carinae by which the much
sculptured carapace is divided up into larger and smaller areas. The last
136 Cumacea: 15. Ekdiastylidae, 1. Ekdiastylis
3 pedigerous segments are extended backwards laterally, causing in the
ovigerous © a wide separation between the 2"¢ and 3¢ peraeopods; hinder
angles of the 5* segment obtuse, more extended in G than in ©. Telson
shorter than peduncle of the uropod, much in 9, a little in d, the slender
post-anal part having about 7 pairs of lateral spines in the 9, a dozen in
the S. Eye distinct. Antenna 1, 2" joint of peduncle the shortest. principal
flagellum 4-jointed in ©, 5- or 6-jointed in G, accessory 2- or 3-jointed.
Antenna 2 of © of five small joints, the last minute. Maxilliped 1 with
about a score of branchial elements in twisted series on the epipod. Peraeo-
pod 2 with 5 joint not quite equal to length of 6 and 7 joints com-
bined. Rami of pleopods very short. Peduncle of uropods longer than the
rami, its medial margin with about 20 spines in Q, 30 to 40 in CG, exopod
longer than endopod, of which the 1* joint is shorter than the 2"4, with
respectively 7 and 10 spines on medial margin of Q, 11 and 18 in G, but
the numbers variable. L. © about 9 mm, GC about 10 mm.
N. America, from Gulf of St. Lawrence to Long Island, surface to 347 m.
2. E. politus (S.1I.Sm.) 1879 Diastylis p., S.1I. Smith in: Tr. Connect. Ac.,
v.51 p. 108 | 1900 D. p., Zimmer in: Fauna arctica, v.1 p.423 | 1912 Ekdiastylis p.,
T. Stebbing in: Aun. 8S. Afr. Mus., v. 10v p. 155.
Agreeing with /. scu/ptus except that the areolations of the cara-
pace are fewer, less deep, and not extending to the postero-lateral portions.
Pedigerous segments 1, 2, in near agreement with /. sculptus, 3—5 more
nearly with those of Diastylis rathkii (p. 98). Telson with slender distal
section scarcely longer than the basal, with 6—9 pairs of lateral spines
ing, 8—10 in the G. Peraeopod 2 with 5% joint nearly twice as long
as 6% and 7 joints combined. Peduncle of uropods longer than the telson
and also than the rami, of which the exopod is the longer; endopod with
4 spines on medial margin of 1S* joint in OQ, 4—6 on the lanceolate 274,
in the GO with 24—30 spines, of which 10—12 are on the 1%" joint. Colour
of G whitish, pellucid, with pink eyelobe. L. o 12 mm, ¢ 14 mm.
N. America, Gulf of St. Lawrence to Vineyard Sound, surface to 847 m.
3. E. insignis (O. Sars) 1871 Diastylis i., G. O. Sars in: Ofv. Ak. Firh., v.28
nr. 1 p.76 | 1871 D.i., G. O. Sars in: Svenska Ak. Handl., n. ser. v9 nr. 13 p.34 t. 14
f, 68—71 | 1879 D.i., A.M. Norman in: Ann. nat. Hist., ser.5 v3 p.55 | 1905 D.2.,
Calman in: Sei. Invest. Fish. Ireland, 1904 app.1 p.10,12,44 | 1912 Ekdiastylis 2.,
T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 155.
Pseudorostral lobes in front of small eyeless eyelobe horizontal, rather
long, acute, with 2 apical spinules, antero-lateral corners rounded, serrulate.
Carapace large, tumid, laterally reticulate with short rows of tubercular spinules,
in front having on each side a strong subventral forward pointing spiniform
process. Pedigerous segments narrow, first concealed except dorsally, side-
plates of the rest small, irregularly emarginate or divided into spiniform
processes, the last 3 spinose at hind margin and having a large pair of
dorsal teeth, the 5 postero-laterally bifid. Pleon segments densely aculeate.
Telson longer than peduncle of uropods, basal two-thirds broad, subeylindric,
with 6 or 7 denticles on each side, the tapering end with single pair of
lateral spines close to the apex and a much larger apical pair. Antenna
1 small, last 2 joints of peduncle subequal. Peduncle of uropods much
longer than the rami, with few spines on the medial margin; exopod con-
siderably longer than the endopod, in which the 1S* joint is longer than
Cumacea: 15. Ekdiastylidae, 1. Ekdiastylis 137
the 2"4, but with spines on medial margin respectively 2 and 3. L. © juy.
scarcely 8 mm.
Off Portugal; lat 38° 7’ N., long. 9°18’ W., depth 1036 m; W. of Shetland, 4—6 m.
4. E. mystacinus (0. Sars) 1887 Diastylis mystacina, G. O. Sars in: Rep. Voy.
Challenger, v.19 pars 55 p.9,60 t. 10 f. 1—3 | 1912 Ekdiastylis mystacinus, 'T. Stebbing
in: Ann. S. Afr. Mus., v. LOv p. 155.
Pseudorostral lobes shortly and horizontally produced in front of the
small rounded eyeless eyelobe, each with apical denticle and strongly but
irregularly dentate crest continued as pair of slightly flexuous carmae on
front of carapace; no antero-lateral corners: surface of large tumid carapace
microscopically spicate, lower margin anteriorly serrate. Pedigerous seg-
ments 1 and 2 with front margin denticulate, 3¢ with transverse dorsal
carina, 42 with pair of medio-dorsal longitudinal serrate carinae, 5% with
pair of dorsal denticles; first 3 pleon segments similarly armed; telson for
little more than proximal half cylindric, rest slightly tapering to apical pair
of stout spines; only 1 lateral pair, with a spinule above on left side of the
specimen. Peraeopods | and 2 with 2™¢ joint densely spinose. Pedunele of
uropods rather shorter than telson, without spines on medial margin, the
endopod about half as long as peduncle, considerably shorter than the exopod,
its 2 subequal joints armed respectively with 1 and 2 lateral spines. L. ©
with marsupium, about 10 mm.
N. Atlantic, lat. 36° 21‘ N., long. 23° 31’ W., depth 3700 m.
5. E. hexaceros (Zimmer) 1908 Diastylis h., Zimmer in: Ergeb. Tiefsee-Exp.,
v.8 p. 158, 187 t. 44 (9) f.983—95 | 1910 D.h., T. Stebbing in: Ann. 8. Afr. Mus., v. 61v
p-418 | 1912 Ekdiastylis h., T. Stebbing in: Ann. S. Afr. Mus., v, 10v p. 155,
Pseudorostral lobes well produced horizontally in front of the small
rounded eyeless eyelobe, with 3 submarginal denticles but no apical denticle.
Carapace large, with a pair of lateral processes in advance of the eyelobe,
a larger pair to the rear of it, and a pair still larger and much wider apart
to the rear of the frontal lobe; much of the lower margin serrate, surface
scabrous with microscopic denticles. Pedigerous segment 3 with submedian
pair of conical bosses, 4t* with a pair of dentate longitudinal carinae, 5°" with
a pair of denticles. Pleon segment 1 begirt with several denticles; the next
2 segments having only 2 pairs, and the 2 following a single pair a-piece.
Telson for rather more than half its length cylindric, then tapering with
5 spines on one side and 6 on the other. Colour white, pellucid. L. O with
well developed marsupium about 10 mm.
S. Africa, outside the Agulhas Bank, lat, 39° 9’S., long. 18° 32°8’ E., depth 565 m,
6. E. argentatus (Calm.) 1912 Diastylis argentata, Calman in: P. U.S. Mus.,
v.41 p.605, 649 tf. 70—75 | 1912 Ekdiastylis argentatus, T. Stebbing in: Ann. S. Afr.
Mis-ailOwe pslod:
Pseudorostral lobes horizontally produced in front of the minute tri-
angular eyeless eyelobe, slightly denticulate above and notched at the base;
carapace with pitted surface, oblique line of denticles on each side of the
pseudorostrum, with small procurved tooth submarginal, about one-third of
the length from the front, followed in the © by a serrate carina reaching
the hind margin; lower margin anteriorly serrate. Pedigerous segments 2, 3,
with side-plates more or less serrate, last 3 segments with some dorsal
denticles, 5tt segment with postero-lateral lobes rounded in ©, bifid in GC.
138 Cumacea: 15. Ekdiastylidae, 1. Ekdiastylis
Some of the pleon segments carrying a few denticles. Telson in © shorter
than peduncle of uropods, but equal to it in O, with about 6 pairs of lateral
spines on the narrowed section in Q, about 10 in d. Antenna 1 with 2n¢
joint the shortest. Antenna 2 of © with no joints elongate. Maxilliped 3,
24 joint not distally expanded or elongate. Peraeopod 2 with 2™4 joint pro-
duced into a curved spiniform tooth. Exopod of uropods about 2/,48 as long
as the peduncle, rather longer than the endopod, of which the first joint is
subequal to the 2°4 in the © (fig.), but much longer than the 224 in the d
(text). Surface of body with a silvery nacreous lustre. L. about 9:5 mm.
Off Chile, lat. 46° 47‘ 30 S., long. 75° 15‘ 00’ W., depth 112 m.
7. E. horridus (O. Sars) 1887 Diastylis horrida, G. O. Sars in: Rep. Voy.
Challenger, v.19 pars 55 p.9,55 t.9 f.1—8 | 1908 D. h., Zimmer in: Ergeb. Tiefsee-
Exp., v.8 p. 158, 184 t.43 f.79—89; t. 44 £. 90—92 | 1912
Ekdiastylis horridus, T. Stebbing in: Ann. 8. Afr. Mus.,
v. 10v_ p. 155.
Pseudorostral lobes acute, well produced
horizontally in front of the rounded eyeless eye-
lobe, antero-lateral corners wanting. Carapace irre-
gularly grooved (or regularly, Zimmer), armed
with many unequal spinules, the whole lower
margin denticulate. Pedigerous segments with
transverse rows of spinules, side-plates with denti-
culate margins. Pleon spinulose, sometimes with
medio-dorsal row of spines. Little groups of
pellucid spots on sides of pedigerous segments
and pleon (Zimmer). Proximal half of telson
cylindric, the rest tapering with several pairs of
lateral spines and a much larger apical pair.
.Antenna 1 with 2 strong spines at apex of 1*¢
joint and long simple seta near end of lateral
margin, 3% joint shorter than 24. Peraeopod 1,
7th joint very slender, much longer than the 6%
(G. O. Sars), sometimes shorter or subequal (Zim-
mer, Calman). Peduncle of uropods a little longer
than the telson, rather more than twice as long
as the exopod, to which the endopod is nearly
Fig. 91.
E. horridus (Sars), telson and
uropod. equal with 3 and 4 (G. O. Sars) or 4 and 7
spines respectively on medial margin of its sub-
equal 18* and 2"4 joints. Colour yellowish white. L. Q 11—14 mm.
Kerguelen, off Cumberland Bay, depth 239 m, and Gazelle Harbour, among red Algae.
8. BE. abbreviatus (0. Sars) 1871 Diastylis abbreviata, G. O. Sars in: Ofv. Ak.
Forh., v.28 nr.1 p.74 | 1871 D.a., G. O. Sars in: Svenska Ak. Handl., n. ser. v9 nr. 13
p.80 t.12 £.62-—64 | 1873 D.a., S.1. Smith in: A. E. Verrill in: Rep. U.S. Fish. Comm.,
v1 p54; | 1879 2D: ahbreviatus, S. I. Smith in: Tr. Connect. Ac., v.51 p.113 | 1900
D. abbreviata, Zimmer in: Fauna arctica, v1 p.420 | 1912 Hkdiastylis abbreviatus,
T. Stebbing in: Ann. S, Afr. Mus., v 10v p. 155.
Pseudorostral lobes much produced horizontally in front of the small
eyeless eyelobe, with serrulate edges; carapace large with finely roughened
surface, the whole lower margin rather strongly serrate, a pair of small
denticles on hinder part of the frontal lobe and another widely separated
Cumacea: 15. Ekdiastylidae, 1. Ekdiastylis, 16. Holostylidae, 1. Holostylis 139
pair much to the rear. Pedigerous segments with few denticles in trans-
verse rows, side-plates of 24 and 34 coarsely crenulate, 5% segment with
4 denticles of some size, the hind angles shortly but acutely extended. Pleon
segments somewhat nodose and denticulate. Telson longer than peduncle
of uropods, its narrowed hinder half carrying 5 pairs of lateral spines. An-
tenna 1 with 2™4and 3% joints of peduncle subequal. Peraeopod 2 with 7%
joint little longer than 6. Peduncle of uropods nearly twice as long as
the exopod, which is considerably longer than the endopod; 1%* joint. of the
latter rather longer than the 2™4, each with 2 spines on the medial margin,
the apical spine as long as the 2"4 joint. L. © juv. scarcely 6 mm.
N. America, lat. 39° 54‘ N., long. 73° 15’ W., depth 56—66 m, Cape Ann, 66 m,
Caseo Bay, 31 m.
9. E. fimbriatus (0, Sars) 1873 Diastylis fimbriata, G. O. Sars in: Svenska
Ak. Handl., n. ser. v.11 nr.5 p.7 t.2 f.4—9 | 1912 EHkdiastylis fimbriatus, T. Stebbing
in: Ann. S. Afr. Mus., v.10v p. 155.
Pseudorostral lobes apically “bent downwards, carapace with only the
anterior part of the lower margin serrate, while somewhat above the lower
margin each side (in G only) is carinate with a remarkable fringe of teeth
encreasing in size to the rear. Pedigerous segments 2—4 have the side-
plates pectinately fringed and expanded beyond the 1S* and 5 segments.
The last has its sides sharply extended backwards and equipped with spinules
and plumose setae; the 34 and 4 pedigerous and (in the @) the first pleon
segments have 2 longitudinal rows of medio-dorsal backward-directed spinules,
and the first 4 pleon segments extend the postero-lateral corners into a
sharp tooth, which in the 2"¢ to the 4 follows a denticulate ridge. Pleon
segments in © not dentate. Telson subequal to peduncle of uropods, the
narrowed distal half carrying in the G7 pairs, in the © 3 pairs, of unusually
long spines and a rather shorter apical pair. Eyelobe flat, without eye.
Antenna 1 in G short, stout, with brush of sensory filaments at base of
4- (or 5+) jointed principal flagellum, accessory 2-jointed; in Q, 3% joint of
peduncle linear, longer than 2"4, the flagella 3- and 2-jointed. Antenna 2
of G extending beyond the uropods. Peduncle of uropods with 12 spines
on medial margin in G, 4 in Q; exopod about 3/,*S as long as the peduncle,
endopod shorter, with 8 and 7 spines on medial margin respectively of 1°
and 24 joints in Gd, 2 and 3 in g.. L. dO scarcely 6 mm.
S. Atlantic, lat. 23° 41/S., long. 419 47/ W.
16. Fam. Holostylidae
1912 Holostylidae, T. Stebbing in: Ann. S. Afr. Mus., v. 10v_ p. 133, 146.
Telson large with only 2 apical spines. Antenna 1 with accessory
flagellum small. Maxilliped 1 with branchial leaflets numerous. Maxilli-
ped 3 with 24 joint very elongate and distally produced. Peraeopods 1—4
with exopods in the male, only the first two pairs having them in the female.
Only two pairs of pleopods in the male. Endopod of uropods 1-jointed.
1 genus.
1. Gen. Holostylis T. Stebbing
1912 Holostylis, T. Stebbing in: Ann. S. Afr. Mus., v. 10v_ p. 133, 146.
With the characters of the family.
2 species.
140 Cumacea: 16. Holostylidae, 1. Holostylis, 17. Pseudocumatidae
Synopsis of species:
{ Third maxilliped with 34 joint much wider than 5th. os 1 as helleri:
\ Third maxilliped with 5th joint much wider than 34... 2. H. gayi
1. H. helleri (Zimmer) 1907 Diastylis h., Zimmer in: Zool. Anz., v.31 p. 221 |
1909 D. h., Zimmer in: Ergeb. Schwed. Siidp.-Exp., v.6 nr.3 p.2, 15 t.6 f. 84—96 |
1912 Holostylis h., T. Stebbing in: Ann. 8S, Afr. Mus., v. 10v p. 146.
Pseudorostral lobes much and sharply produced in front of the small
eyeless eyelobe; antero-lateral corners subquadrate with denticulate margin
stronger above than below. Carapace strongly beset with
powerful teeth, of which 2 are especially so on each side
of the frontal lobe and one below it; besides there is a
close covering of microscopic denticles and the lower margin
is denticulate. Pedigerous segments 1 and 2 short, with
side-plates beyond width of carapace. All the pedigerous
segments and first 3 of pleon have medio-dorsal pairs of
warts or teeth, the latter form on the last 3 pedigerous and
first 2 pleon segments. On first 5 segments of pleon a
median dentate carina encreasing on successive segments,
Telson begins laterally dentate, narrows before the middle,
with about 9 pairs of lateral spines and a stronger apical
pair. Antenna 1 short and weak. Branchial leaflets about
20. Maxilliped 3, 2"¢ joint nearly thrice the length of rest
combined, 5t® narrower than 3¢ or 4, Peraeopod 1 with
strongly dentate 2™¢ joint, and a medial distal tooth on
34 and 4th joints. Peraeopod 2, 5t joint not so long as
6th and 7t combined, 6t” little shorter than 7%. Peduncle
NUS EE o of uropods about as long as the telson with some 16 spines
Banat: on the medial margin. Rami about half as long as the
“uropod. peduncle (text), exopod with 1st joint much shorter than 2”,
slightly longer than the 1-jointed endopod which has about
11 spines on its medial margin. Colour yellowish white. L. @ about 13 mm.
South Georgia, depth 20—75 m.
2. H. gayi (Nic.) 1849 Cuma g. H. Nicolet in: Gay, Hist. Chile, v.3 p.220,
Crust. t.3 f. 1,la—d | 1912 Holostylis g., T. Stebbing in: Ann. S. Afr. Mus., 7. 10v p. 146.
Pseudorostral lobes somewhat upturned; carapace inflated at the sides,
covered with transverse carinae to the rear; telson less than two-thirds the
length of peduncle of uropods, with 6 or 7 pairs of lateral spines and a larger
apical pair. Maxilliped 3 with 2™¢ joint twice as long as rest combined,
carrying many setae on the dilated and produced apex, 5‘ joint wider than
34 or 4th, Peduncle of uropods with 12 spines on medial margin; exopod
with 2 equal joints, rather more than half as long as the peduncle; endopod
more robust, as long as the telson, 1-jointed, with 7—9 spines on the medial
margin. L. © 65 mm.
San Carlos, Chiloe, among Fucus.
17. Fam. PSeudocumatidae
1878 & 1879 Pseudocumidae, G. O. Sars in: Arch. Naturv. Kristian., 7.3 p. 466;
v4 p.59 | 1887 P., G. O. Sars in: Rep. Voy. Challenger, v.19 pars 55 p. 7 |-1893° P.
T. Stebbing in: Int. sci. Ser., v.74 p.307,312 | 1900 P., G. O. Sars, Crust. Norway,
v.38 p.72|1900 P., T. Scott in: Ann. Scott. nat. Hist., v.9 p. 222 | 1903 P., Zimmer in:
Cumacea: 17. Pseudocumatidae, 1. Petalosarsia 141
Zool. Jahrb., Syst. v. 18v1 p. 694 | 1912 P., Calman in: P. U.S. Mus., v.41 p. 605 | 1912
Pseudocumatidae, T. Stebbing in: Ann. S. Afr. Mus., v. 18v_ p. 133.
Lobes of pseudorostrum contiguous from the eyelobe to the apex
(except in Schizorhynchus p. 143). Protuberances on the pedigerous segments
when present not spiniform. Pleon slender, smooth; telson small, lamellar,
not apically acute or subacute. Eyelobe varying in shape, eye composed otf
3 or 2 corneal lenses or quite indistinct. Antenna 1 with accessory flagellum
rudimentary. Antenna 2 in female very small, in male well developed.
Spines in spine-row of mandible not numerous, molar stout. Epipod of maxilli-
ped 1 scantily furnished with branchial leaflets, at least in the female. Maxilli-
ped 3 with long 24 joint not apically produced but furnished at the lateral
angle with a very long seta. First four pairs of peraeopods in the male
provided with exopods well developed, hut only first two pairs in the female,
the third and fourth pairs in that sex having the exopods represented by
more or less distinctly 2-jointed rudiments attached to the 2"? joint. The
first two segments of the pleon in the male have small pleopods, the first
distinctly 2-jointed, setiferous, the second almost rudimentary, with the 2"¢
joint fused or diminutive. Uropods with 1-jointed endopod.
6 genera accepted, 1 doubtful.
Synopsis of the accepted genera:
Peraeopod 1 with third and fourth joints firmly united
| and the fifth abnormally dilated ........, 1, Gen. Petalosarsia
: | Peraeopod 1 with third, fourth, and fifth joints normal
in size and arrangement — 2.
| Pseudorostral lobes not distally contiguous... . . 2. Gen, Schizorhynechus
~ \ Pseudorostral lobes distally contiguous — 3.
Fifth pleon segment not longer than broad; peduncle
3 | of uropods very short. .....:s...: he 3. Gen. Caspiocuma
| Fifth pleon segment longer than broad; peduncle of
uropods not very short — 4.
| Pedigerous segments, some or all, with dorsal pro-
4 - tuberances. ... ; et wee ic hee. 9 6. 4aGen, Pterocuma.
| Pedigerous segments without dorsal protuberances — 5.
> {Body very slender... .... howe Sea) ee 5. Gen. Stenocuma
all Body not specially slender ........-. . . . 6.Gen, Pseudocuma
1. Gen. Petalosarsia Stebbing
1865 Petalopus (Sp. un.: P. declivis) (non Motschulski 1845), G. O. Sars in: Forh.
Selsk. Christian., 1864 p.196 | 1883 Petalomera (non Stimpson 1858), G. OU. Sars in:
Forh. Selsk. Christian., 1882 nr. 18 p. 18, 58 | 1893 Petalosarsia, T. Stebbing in: Int.
sci. Ser., v.74 p. 308 | 1900 P., G. O. Sars, Crust. Norway, v.3 p.77.
In general character agreeing with Psewdocuma (p. 148), but the eye
feebly developed, first antenna with second joint broadly lamellar and third
very small, second antenna of female distinctly two-jointed, and first peraeo-
pods abnormal in structure, the third and fourth joints being firmly united
to form a small cup for the very large foliaceous fifth joint, over the broad
distal end of which the slender sixth and seventh joints can be folded to
produce a subchelate arrangement; the small second pleopod simple.
1 species.
142 Cumacea: 17. Pseudocumatidae, 1. Petalosarsia
1. P. declivis (O. Sars) 1865 Petalopus d., G. O. Sars in: Forh. Selsk. Christian.,
1864 p.197 | 1883 Petalomera d., G. O. Sars in: Forh. Selsk. Christian., 1882 nr. 18
p. 13, 58 | 1886 P. d., G. O. Sars in: Norske Nordhavs-Exp., v.6 nr.15 p.24 | 1892 P. d.,
T. & A. Scott in: Ann. nat. Hist., ser.6 v.10 p.206 | 1893 Petalosarsia d., T. Stebbing
in: Int. sci. Ser., v. 74 p. 308 | 1894 P. d., A. O. Walker in: Rep. Brit. Ass., v. 64
p. 326 | 1897 Petalomera d., Ehrenbaum in: Wiss, Meeresunters., v.21 p. 419 | 1900
Petalosarsia d., G. O. Sars, Crust. Norway, v3 p.77,108 t.54 | 1900 P. d., Zimmer in:
Fauna arctica, v.1 p.428,437, 441 | 1902 P. d.. T. Scott'in: Rep. Fish. Board Scotl., v.20
p. 509 | 1909 P. d., A. M. Norman & G. S. Brady in: Nat. Hist. Tr. Northumb., ser. 2
v.3 p.34|1911 P.d., Stappers in: Camp. arct. Orléans, Crust. Malac. p.121 t.6 f.11,
12 | 1912 P.d., Calman in: P. U.S. Mus., v.41 p. 675.
Fig. 95.
P. declivis (Sars),
pleopod 2.
Fig. 93.
pod 1.
Fig. 96.
Fig. 94. P. declivis (Sars), telson and
P. declivis (Sars), pleopod 1. uropod.
Body rather short and compact, integument strongly indurated. Pseudo-
rostrum rather prominent, acute, more so in © than in G, antero-lateral corners
minutely projecting with a little apical denticle. From near this point to
the hind margin a pair of strong dorso-lateral carinae traverse the carapace,
to which in the G are added a second pair subparallel to the upper pair.
The pedigerous segments smooth. Pleon rather shorter than preceding part
of the body, telson smooth, broadly semioval. Antenna 1, 2"¢ joint shorter but
broader than the 1S‘, with medial margin partially serrate, 34 joint scarcely
longer than broad in Q and broader than long in G. Antenna 2 in © with
24 joint much narrower than 15*, but nearly as long, flagellum in G reaching
Cumacea: 17. Pseudocumatidae, 1. Petalosarsia, 2. Schizorhynchus 143
to the end of the body. Peraeopod 1, 2"4 joint much curved, not so long
as remainder of the limb fully extended, 5' joint twice as long as broad,
but its breadth exceeding length of the sixth joint. 2™¢ peraeopod scarcely
more than half as long as the 1**, its 5 joint a little longer than the 4'"
but shorter than the 7". Peduncle of uropods little more than half the length
of the endopod, which is rather longer than the exopod. Colour uniformly
yellow. L. oO 4mm, G nearly 5 mm.
Norway, Lofoten Islands, depth 94—113 m, Coast of Finmark, depth 38—75 m,
Spitzbergen; Franz-Josefs Land; Heligoland; N. E. England, depth 68-104 m; Firth
of Forth; Irish Sea; off Newfoundland, depth 160m; U.S. A., off Martha’s Vineyard,
depth 71m; Nova Zembla, 90 m.
2. Gen. Schizorhynchus 0. Sars
1900 Schizorhynchus, G. O. Sars, Crust. Norway, v.3 p. 73.
Pseudorostrum with lobes meeting for a very short space in front of
the eyelobe, then being separated by a deep interval; antero-lateral corners
more or less quadrate, with serrulate front margin. Eyelobe short and broad,
occupied generally by a transverse row of three corneal lenses, of which the
central one may be wanting in the female. Peduncle of uropods not longer
than the rami.
4 species.
Synopsis of species:
Integument everywhere spiculate .......... 1. S, seabriusculus
4 Integument not spiculate — 2.
bo
Uropod with equal rami; telson not denticulate — 3.
Telson nearly quadrate; eye of female with only 2 lenses 3. S. bilamellatus
2 Telson semicircular; eye of female with 3 lenses ... 4. S. abbreviatus
J Uropod with endopod shorter than exopod; telson denticulate 2. S. eudorelloides
\
1. S. seabriusculus (O. Sars) 1894 Pseudocuma scabriuscula, G. O. Sars in:
Bull. Ac. St.-Pétersb., ser. 4 v.4 (v.36) p.332 t.12 f.11—14 | 1900 Schizorhynchus scabri-
usculus, G. O. Sars, Crust. Norway, v.3 p.73 | 1907 Pseudocuma scabriuscula, Lebediew
in: Trudui St.-Peterb. Obshch., v. 381 p.95.
Body short and stout, integument hard, everywhere scabrous with small
projecting spikes. Pseudorostrum produced to widely and deeply separated
subacute points, the antero-lateral corners subquadrate, serrate in front. Cara-
pace broadest behind, a little constricted in the middle, with a pair of carinae
from tips of pseudorostrum embracing the frontal region, branchial regions
tumid. Pedigerous segments not carinate. Pleon short, telson small, semi-
circular. Eyelobe truncate, with all three corneal lenses distinct. Peduncle
of uropods considerably shorter than the subequal rami, endopod with serrulate
medial margin. Colour greyish white. L.o 3mm. GC unknown.
Caspian Sea.
2. S. eudorelloides (O. Sars) 1894 Pseudocuma e., G. O. Sars in: Bull. Ac.
St.-Pétersb., ser. 4 v.4 (v.36) p. 330 t.12 f.1—10 | 1900 Schizorhynchus e., G. O. Sars,
Crust. Norway, v.3 p. 73 | 1901 Pseudocuma e., S. Zernow in: Annuaire Mus. St.-Pétersb.,
v.6 p.568 | 1907 P. e., Lebediew in: Trudui St.-Peterb. Obshch., v. 381 p. 95.
Body rather stout, integument not very hard, squamose. Pseudorostrum
projecting in 2 rounded lobes, narrowly but deeply separated, the antero-
144 Cumacea: 17. Pseudocumatidae, 2. Schizorhynchus
lateral corners nearly rectangular, front margin serrulate. Carapace with a
pair of serrulate submedian carinae between the eyelobe and the middle,
their hinder part flanked by crests which rising from
the sides curve round so as to define medio-dorsally
the branchial regions. Pedigerous segments evenly
arched above. Pleon not very long. ‘Telson sub-
semicircular, the serrulate margin affording 2 decided
teeth on each side of the narrowly truncate apex.
Kyelobe short and broad, concavely truncate, with 2
small corneal lenses, one at each corner. Antenna 1,
gud joint longer than 18¢ or 3¢, which are subequal
in length. Peraeopod 2 particularly short and stout.
Fig. 97.
S. eudorelloides (Sars) :
carapace. Pedunele of uropods equal in length to exopod, endo-
pod a little shorter, with 5 spinules along the medial
margin. Colour whitish, L. © under 5mm. GC unknown.
Caspian Sea.
3. S. bilamellatus (O. Sars) 1894 Pseudocuma bilamellata, G.O.Sars in: Bull.
Ac. St.-Pétersb., ser. 4 v.4 (v.36) p.326 t.10,11 | 1900 Schizorhynchus bilamellatus, G. O.
Sars, Crust. Norway, v.3 p. 73.
Body rather long and slender, integument moderately hard, squamose.
Pseudorostrum projecting in 2 thin oval lamellae some distance apart, point-
ing obliquely upwards, antero-lateral corners nearly rectangular in Q, more
obtuse in G, front edge serrulate. Carapace with a pair of submedian carinae,
serrulate in Q, smooth in G, running from front of eyelobe to the middle,
flanked to the rear by crests which rising from the sides abruptly turn so
as medio-dorsally to define the branchial regions. Pedigerous segments not
carinate, 2"4 and 34 dorsally short, much expanded at the sides, 4% and 5%
dorsally large, 5t* with hind corners a’ little extended backwards. Pleon
rather elongate. Telson square, length and breadth being equal, but hind
angles rounded off. Eyelobe broad, with front corners rounded, especially in
CS which has 3 corneal lenses, while the © has only 2, one at each corner
of the lobe. Antenna 1, 2"¢ joint longer than 1%‘, 3¢ than 294. Upper lip
distally narrowed, distal border emarginate. Peraeopod 1 rather slender.
Peraeopod 2 with 5* joint in 9 a little longer than the 2 preceding joints
combined, but in G more than twice as long and more setiferous, the 7‘
joint in G a little curved and distally armed with 5 retroverted teeth. Pe-
duncle of uropods in Q as long as the equal rami, rather shorter than the
rami in the CG, the endopod in both sexes having the medial margin closely
fringed with spinules. Colour whitish, L. @ 9mm, d 10 mm.
North Caspian Sea,
4. S. abbreviatus (0. Sars) 1897 Pseudocuma abbreviata, G.O. Sars in: Annuaire
Mus. St.-Pétersb., v. 2 p. 283 t.14 f. 9—12 | 1900 Schizorhynchus abbreviatus, G. O. Sars,
Crust. Norway, v.3 p. 73.
Body short and compact. Pseudorostrum rather short, a deep exca-
vation separating its rounded processes, which laterally lead by a
shallow emargination to the serrulate rounded antero-lateral corners. Cara-
pace deeper than wide, slightly narrowed to the rear, without longitudinal
carinae and the branchial regions not sharply defined. Pedigerous segments
smooth. Pleon short and narrow, telson semicircular. Eye composed of 3
corneal lenses in a transverse row on the short broad eyelobe. Antenna 1,
Cumacea: 17, Pseudocumatidae, 2. Schizorhynchus, 3. Caspiocuma, 4. Pterocuma 145
2nd joint distally expanded, carrying above a dense row of upturned setae.
Uropods with peduncle considerably shorter than the subequal rami, endopod
with serrate medial margin. L. © 3 mm. GC unknown.
North Caspian Sea.
3. Gen. Caspiocuma 0. Sars
1900 Caspiocuma, G. O, Sars, Crust, Norway, v.3 p. 73.
Integument indurated. Carapace without carinae, its branchial regions
greatly inflated. First pedigerous segment narrow and inconspicuous, the
rest sloping steeply to the pleon, which is short, depressed, the segments
on each side expanding into a shelf-like carina. Eye inconspicuous or ab-
sent. Peduncle of uropods very short. Appendages in general not known
to differ from those of Pseudocuma.
1 species.
1. C. campylaspoides (O. Sars) 1897 Pseudocuwa c., G.O.Sars in: Annuaire
Mus. St.-Pétersb., v.2 p.286 t.15 f.1—3 | 1900 Caspiocuma c., G. O. Sars, Crust. Norway,
v.3 p. 73 | 1907 Pseudocuma c., Lebediew in: Trudui St.-Peterb. Obshch., v. 381 p. 95.
Body short and compact, integument hard and thickly incrusted, with
general aspect as in Campylaspis. Pseudorostrum short, apically blunt, antero-
lateral corners inconspicuous, obtusely rounded off. Front part of carapace in
dorsal view much narrower than the broad branchial region. Telson semioval,
narrowly rounded at apex. The minute eyelobe showing no trace of pigment
or corneal lenses. Uropods short and stout, peduncle not much longer than
broad, shorter than exopod, which is shorter than the endopod, about in
proportions of 2:3:4. L. o 3mm, GC unknown.
South Caspian Sea, probable depth 75 m.
4, Gen. Pterocuma 0. Sars
1900 Pterocuma, G. O. Sars, Crust. Norway, v.3 p. 73.
Pedigerous segments some or all with dorsal protuberances. Telson
quadrate, broader than long. Eyeiobe triangular or rounded; of the 3 corneal
lenses the central one or all three may be evanescent in the female. Rami
of uropods subequal.
3 species.
Synopsis of species:
1 J Pseudorostral lobes very prominent, acuminate. .... .- 1. P. rostratum
\ Pseudorostral lobes rather short, not acuminate — 2.
9 J Second pedigerous segment with a pair of submedian processes 2, P. pectinatum
\ Second pedigerous segment with a single median process. 3. P. sowinskyi
1. P. rostratum (O.Sars) 1894 Pseudocuma rostrata, G. O. Sars in: Bull. Ac. St.-
Pétersb., ser. 4 v.4 (v.36) p.313 t.4 | 1900 Pterocuma r., G. O. Sars, Crust. Norway, v.3
p.73 | 1907 Pseudocuma r., Lebediew in: Trudui St.-Peterb. Obshch., v. 381 p. 95.
Body rather stout. Pseudorostrum prominent, acuminate, antero-lateral
corners nearly quadrate, serrulate in front. Branchial regions of carapace
bounded by an upper and a lower pair of oblique carinae and traversed
midway between these by a third pair, an intervening wedge-shaped dorsal
Das Tierreich. 39. Lief.: T. R. R. Stebbing, Cumacea. 10
146 Cumacea: 17. Pseudocumatidae, 4. Pterocuma
space separating the carinae of one side from those on the other. Pedige-
rous segments longitudinally carinate, the keel successively encreasing, that
of the last segment the most prominent, triangular. Pleon smooth, telson
small, broader than long, quadrangular, hind angles rounded off. Eye, 2 patches
of pigment in triangular eyelobe. Uropods short and stout, peduncle sub-
equal to the subequal rami, endopod with 1 spinule below the middle of the
serrulate medial margin. Many reddish brown stellate flecks. L. oO 8 mm,
GS unknown. A
North and South Caspian Sea.
2. P. pectinatum (Sowinsky) 1892 Pseudocuma pectinata, Sowinsky in: Mém. Soe.
Kiew, v.11 p.113 | 1894 P. p., G.O.Sars in: Bull. Ac. St.-Pétersb., ser. 4 v. 4 (v. 36) p. 299
t.1,2 | 1897 P.p., G. O. Sars in: Annuaire Mus. St.-Pétersb., v.2 p. 288 | 1899 P. p.,
Sowinsky in: Bull. Ac. St.-Pétersb., ser.5 v.8 p.376 | 1900 Pterocuma p., G. O. Sars,
Crust. Norway, v.3 p.73 | 1901 Pseudocuma p., 8. Zernow in: Annuaire Mus. St.-Pétersb.,
v.6 p. 666,573 t.22 f.19,20 | 1907 BP. p., Lebediew in: Trudui St.-Peterb. Obshch.,
v. 381 p. 95.
Body rather stout, integument squamose. Pseudorostrum short. slightly
upturned, subacute, the antero-lateral corners broadly triangular. Carapace
smooth, the branchial regions tumid, gibbous behind in ©. Of the pedige-
rous segments the 2"¢ has a pair of submedian dorsal processes and the
side-plates much extended forwards in a rounded lobe, the 3¢has a single
medio-dorsal process not large, the long 4th and 5t* having each a large,
laminar, linguiform, erect, median process. The 5 segment has the postero-
lateral corners a little obtusely extended, and in the ¢O fringed with plumose
setae. The first 5 pleon segments have a pair of longitudinal carinae, dis-
tally raised and rounded; each segment also having the hind border raised,
and emarginate on both dorsal and ventral faces, while the sides project at
an obtuse angle. Telson very small, rather broader than long, quadrangular.
Eye with 3 corneal lenses in a rounded eye-lobe. Antenna 1, Ist and 274
joints subequal in length, 3¢ shorter, flagellum 3-jointed, accessory minutely
1-jointed. Uropods long and slender, rami considerably shorter than peduncle.
endopod rather longer than exopod, with serrulate medial margin, proximal
half of which in dis densely fringed with spinules. Colour varied with
reddish brown arborescent patches. L. O 8, d 85 mm.
Black Sea (Sea of Azov); N. Caspian Sea.
3. P. sowinskyi (O. Sars) 1894 Pseudocuma s., G. O. Sars in: Bull. Ae. St.-
Pétersb., ser.4 v.4 (v.36) p.312 t.3 | 1900 Pterocuma s., G. O. Sars, Crust. Norway, v.3
p. 73 | 1901 Pseudocuma s., 8. Zernow in: Annuaire Mus. St.-Pétersb., v.6 p.566 | 1907
P. s., Lebediew in: Trudui St.-Peterb. Obshch., v. 881 p. 95.
Pseudorostrum short, slightly upturned, subtruncate, antero-lateral corners
much produced, with concave front margin coarsely serrate. Carapace with
a pair of sinuous carinae from tips of pseudorostrum embracing the whole
frontal region and then with strong curve ferming serrate submedian boundaries.
to the tumid branchial regions. A median carina is figured along the proxi-
mal half of the carapace. All the pedigerous segments with medio-dorsal
carina, rather low and rounded on the first 3, higher and laminar on 4
and 5%, on the 5 triangular, the carinal edges coarsely serrated. Pleon
segments smooth. Telson transversely quadrangular, much broader than long.
iyelobe rounded triangular, the middle corneal lens quite inconspicuous.
Antenna 1 as in P. peetinatum. Uropods rather short and stout, rami subequal
Cumacea: 17. Pseudocumatidae, 4, Pterocuma, 5. Stenoecuma 147
one to the other and not much shorter than the peduncle. Colour diversified
with reddish brown stellate flecks. L. OQ 11 mm, ¢ unknown.
N. und N. W. Caspian-Sea, Sea of Azov.
5. Gen. Stenocuma 0. Sars
1900 Stenocuma, G. O. Sars, Crust. Norway, v.3 p. 73.
Body slender, integument little indurated. Pedigerous segments without
dorsal carinae or protuberances. Telson quadrate or semielliptic. Eyelobe
semicircular, with or without the three corneal lenses. Rami of uropods
subequal, the exopod not at all shorter than the endopod.
4 species.
Synopsis of species:
Carapace with 2 pairs of longitudinal serrated crests . 1. S. diastyloides
Carapace without serrated crests — 2.
| Peraeopods 3—5 exceptionally stout, Deer exceptionally
MGR. Gad aided, ONG ue oto « -.-.. YS. tenuicauda
Dimensions of peraeopods aad pleon not exceptional — 3.
Carapace very narrow, Phe oa ria rather short and
obtuse: . 3. = : : 2 2 od. 8. eracile
Carapace not very narrow, Baendoresiran ll pounced!
GE ORKCHUG 6-6 Ag. Bh GMA Aeon Gullo. 6 Fonte eS ak . . 4. 8. graciloides
1. S. diastyloides (O. Sars) 1897 Pseudocwma d., G. O. Sars in: Annuaire
Mus. St.-Pétersb., v.2 p.279 t.14 f.1—8 | 1900 Stenocuma i G. O. Sars, Crust, Norway,
v.3 p. 73.
Pseudorostrum very prominent, horizontal, subacute, antero-lateral
corners quadrate with serrate margin. Carapace with a pair of serrate carinae
from ends of eyelobe to about the middle, followed by a pair wider apart,
which turn abruptly downwards as they approach the anterior pair. Pedi-
gerous segments 2—4 broadly expanded laterally, 5‘ with hind corners slightly
extended backwards. Telson quadrate, rather broader than long, distal margin
slightly convex between two acute points. Eyelobe well developed, but showing
no pigment or lenses. Antenna 1, 1%* joint slightly curved, stout, as long
as 34, shorter than 2™4, flagellum very small, 3-jointed. Peraeopod 1, 24
joint as long as the other joints combined, which is not the case with 2n¢
joint of peraeopod 2. Rami of uropods rather shorter than the peduncle,
the 1-jointed endopod having on the medial margin 9 spines and at the
apex 2 that are larger but very unequal. L. Q 10 mm, oO unknown.
South Caspian Sea, depth 53—170 m.
2. S. tenuicauda (O. Sars) 1894 Pseudocuma t., G. O. Sars in: Bull. Ac. St.-
Pétersb., ser.4 v.4 (v.36) p. 323 t.9 | 1900 Stenocuma t., G. O. Sars, Crust. Norway,
v.3 p. 73.
Integument very thin, pellucid. Pseudorostrum short, level, truncate,
antero-lateral corners blunt, little prominent. Carapace narrow, without
carinae. Free pedigerous segments smooth, last 3 laterally expanded and
last 2 laterally tumid. Pleon very narrow, not very long, telson small, semi-
elliptic. Kye with 3 corneal lenses. Antenna 1, 1% joint shorter than either
of the subequal 2"4 and 3% joints. Peraeopod 1, 2"4 joint shorter than the
rest combined, 5 much longer than either of the subequal 6*" and 7, Peraeo-
pod 2, 2"¢ joint much shorter than the rest combined, 6 shorter than 4h,
10*
148 Cumacea: 17. Pseudocumatidae, 5. Stenocuima, 6. Pseudocuma
5th, or 7%, which are subequal in length. Peraeopods 3—5 abnormally: stout,
4th joint not much shorter than 2™¢ in the last 2 pairs, 5‘ joint short, oval,
fringed with strong simple setae on medial margin. Rami of uropods sub-
equal, a little shorter than the peduncle, endopod having on medial margin
3 spines in ©, 10 (7 in fig.) in Dd. Colour whitish. L. 3°5 mm.
N. W. Caspian ‘Sea.
3. 8. gracile (O. Sars) 1894 Pseudocuma gracilis, G. O. Sars in: Bull. Ac. St.-
Pétersb., ser.4 v.4 (v.36) p.318 t.7 | 1900 Stenocuma g., G. O. Sars in: Crust. Norway,
v.83 p.73 | 1907 Pseudocuma g., Lebediew in: Trudui St.-Peterb. Obshch., v. 381 p. 95.
Integument thin, semi-pellucid. Pseudorostrum short, level, apically
somewhat obtuse, antero-lateral corners little prominent. Whole body without
carinae. 24 and 3% pedigerous segments the largest, laterally
very broad and laminar, 5% with hind corners evenly rounded.
Pleon very long and narrow. Telson small, semi-elliptic. Hye
with 3 corneal lenses. Antenna 1, joints 1—3 subequal in
length. Peraeopod 1, 2"¢ joint as long as the rest combined,
5th much longer than either 6 or 7. Peraeo-
pod 2, 2™¢ joint rather shorter than the rest
combined, 6 shorter than 5 or 7*, 7% distally
armed in CO with 3 retroverted hooks.. Peraeo-
pods 3—5 slender, 4 joint shorter than 5!
Fig. 8. = in the 34 and 4* pairs. Rami of uropods sub- ee
S. gracile (Sars) S. gracile (Sars)
pleopod 1. equal one to the other and to the peduncle, pleopod 2.
endopod having medial margin with 8 spines
in Q, ciliated and with 9 spines in 3. L.4:5--6mm. Colour uniformly whitish.
N. W. Caspian Sea.
4. S. graciloides (O. Sars) 1894 Pseudocuma g., G. O. Sars in: Bull. Ac. St.-
Pétersb., ser. 4 v.4 (v.36) p. 321 t.8 | 1900 Stenocuma g., G. O. Sars, Crust. Norway,
v.8 p.73 | 1902 Pseudocuma latiaculeata + P. gracilloides, S. Zernow in: Annuaire Mus.
St.-Pétersb., v.6 p. 566,570 t.22 f.21—24; p.570,577 | 1907 P. graciloides, Lebediew
in: Trudui St.-Peterb. Obsheh., v. 381 p. 95.
More strongly built than S. gracile. Pseudorostrum longer and apically
subacute. Branchial regions of carapace more distinctly defined. Peraeopod 2
with 7% joint decidedly longer than 54, instead of scarcely equal to it.
Uropods with 6 spines on medial margin of endopod in Q, 10 in oO.
L. 6mm. Colour variegated with reddish brown, stellate flecks. Character
of spines on which P. latiaculeata was chiefly based is common to many species.
North Caspian Sea; Sea of Azoy.
6. Gen. Pseudocuma, 0. Sars
1858 Cyrianassa (Sp. un.: C. longicornis) (non Bate 1856), Bate in: Kinahan in:
Nat. Hist. Rev., v.5 P. p.203 | 1861 Leucon (non Kroyer 1846), P. J. Beneden in: Mém.
Ac. Belgique, v. 33 [nr.3] p. 85 | 1865 Pseudocuma, G. O. Sars in: Forh. Selsk. Christian.,
1864 p.68 | 1879 P., G. O. Sarsin: Arch. Naturv. Kristian., v.4 p.60 | 1893 P., T.Stebbing
in: Int. sci. Ser., v.74 p.307 | 1894 P. (part.), G.O.Sars in: Bull, Ac. St.-Pétersb.,
ser. 4 v.4 (v.36) p. 298 | 1897 P. (part.), G. O. Sars in: Annuaire Mus. St.-Pétersb., v.2
p. 289 | 1900 P., G. O. Sars, Crust. Norway, v.3 p.73 | 1909 P., Stappers in: Cellule,
v.25 p.368.
Pedigerous segments without carinae or protuberances. Telson semi-
circular or bluntly semiovate. Eyelobe semicircular or heart shaped with 3
Cumacea: 17. Pseudocumatidae, 6. Pseudocuma 149
corneal lenses. Second antenna of female very small, one-jointed, with a
single apical seta. Peduncle of uropods not very short. Hepato-pancreatic
glands with two pairs of lobes. ;
4 species accepted, 1 obscure.
Synopsis of the accepted species:
Carapace without lateral folds... .... - «'.. « 1. P. cercaroides
Carapace with lateral folds — 2.
iJ
\
J Pseudorostrum with antero-lateral corners denticulate . 2. P. simile
|
(
bo
Pseudorostrum with antero-lateral corners not denticulate — 3.
. {| Antenna 1 with third joint longer than the second . . 3. P. longicorne
3 | Antenna 1 with third joint shorter than the second. . 4. P. ciliatum
1. P. cerecaroides O, Sars 1894 P.c., G. O. Sars in: Bull. Ac. St.-Pétersb.,
ser. 4 v.4 (v.36) p.315 t.5,6/ 1900 P. c., G. O. Sars, Crust. Norway, v.3 p. 73 | 1907
P. c., Lebediew in: Trudui St.-Peterb. Obsheh., v.381 p. 95.
Body short and stout, integument strongly incrusted, squamose. Pseudo-
rostrum rather prominent, level, subacute; antero-lateral corners small, ob-
tuse in ©, nearly quadrate in GO. Carapace without median or lateral longi-
tudinal carinae but with sinuous ridges defining the branchial regions. Pedi-
gerous segments smooth, laterally well expanded, 18 very short, the much
narrowed 5t* with rounded postero-lateral corners. Pleon narrow, telson very
small, nearly semicircular. Eyelobe semicircular, with 3 distinct corneal
lenses, pigment rather dark. Antenna 1, 3% joint shorter than either of the
subequally long 18* and 24 joints. Maxillipeds 2 and 3 and the peraeopods
robust. Peraeopod 1, 2"4 joint shorter than rest of limb; tip of peraeopod
2 not hooked; in the last three pairs the 4% joint is twice as long as the
5th; the rudimentary exopod of the 3¢ and 4 pairs in the © is extremely
small but distinctly 2-jointed. Uropods rather short and in 9 rather stout,
peduncle having on medial margin 4 spines in ©, 5 in GC, the subequally
long endopod with 4 spines in 9, 9 in CO together with fringe of fine cilia,
exopod considerably shorter. Colour yellowish. L. 5 mm.
Caspian Sea.
2. P. simile O.Sars 1900 P. similis, G.O.Sars, Crust. Norway, v.3 p.76 t. 53 |
1900 P.s., T. Scottin: Ann. Scott nat. Hist., v9 p.222 | 1902 P.s., T. Scott in: Rep.
Fish. Board Scotl., v.20 p. 480 | 1903 P. s., Zimmer in: Zool. Jahrb., Syst. v. 18vi
p- 694 | 1905 P.s., Calman in: Sci. Invest. Fish. Ireland, 1904 app. 1 p. 36 | 1906 P. s.,
Gilson in: Mém. Soc. ent. Belgique, v.12 p.78 tf.4,5, 7, 12,15 (detailed comparison
with P. longicorne) | 1909 P. s., Stappers in: Cellule, v.25 p.368 t.1 f. 9a; t.2 f. 20 |
1909 P.s., A. M. Norman & G.S. Brady in: Nat. Hist. Tr. Northumb., ser. 2 v.31 p. 34.
Similar to P. longicorne (nr. 3), but larger and less slender, lobes of
pseudorostrum less prominent, antero-lateral corners armed in front with 3
—denticles. Sculpture of carapace, eye, antennae, mouth-organs and limbs
about as in the species mentioned, but terminal joint of peraeopod 2 in Jd
with 2 denticles and a hook at the apex (Sars), 2 apical hooks and 1 sub-
apical (Gilson). Pleopod 1 of G with only 2 setae on medial margin of 1**
joint, and pleopod 2 with only 1 spine on lateral margin. Peduncle of
uropod longer than the subequal rami, endopod in © having about 6 spines
on medial margin, where the G has 12. Telson rather broader than long,
truncate in ©. Colour light. L. @ 5mm, ¢ 5:5 mm.
W. coast of Norway, depth 11—22 m; North’Sea; Scotland, E., W., N. E.; W.
coast of Ireland, shallow water; off N. E. England, depth 53 m.
150 Cumacea: 17. Pseudocumatidae, 6. Pseudocuma
3. P. longicorne (Bate) 1858 Cyrianassa longicornis (J.V. Thompson MS. sp.),
Bate in: Kinahan in: Nat. Hist. Rev., v5 p. 204 | 1858 C. 1, Bate in: J. R. Dublin Soc..
v.2 p. 101 tf. s | 1861 Leucon cercaria, P. J. Beneden in: Mém. Ac. Belgique, v. 33 [nr. 3]
p-85 t.14 | 1865 Pseudocuma bistriata, G. O. Sars in: Forh. Selsk. Christian., 1864
p-70 | 1877 Cuma bella, Meinert in: Naturh. Tidsskr., ser.3 v.11 p.179 | 1879 Psewdocuma
cercaria, G. O. Sars in: Arch. Naturv. Kristian., v.4 p.62 t,40—42 | 1880 Cuma c.,
Meinert in: Naturh. Tidsskr., ser. 3 v.12 p.497 | 1893 Pseudocuma longicornis, T. Stebbing
in: Int. sci. Ser., v.74 p.307.| 1895 P. 1, H. J. Hansen in: Ergeb. Plankton-Exp., v. 2
G.c. p.59 | 1900 P. cercaria, G. O. Sars, Crust. Norway, v.3.p. 74, 108 t. 51, 52 | 1900
P. longicornis, Zimmer in: Fauna arctica, v.1 p.428 (with additional syn.) | 1905. P. 1.,
Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p.36 | 1906 P.1., Gilson in: Mém,
Soc. ent. Belgique, v.12 p.78 tf. 1—3, 6,8—11, 18, 14 | 1906 Pl, A. M. Norman &
T. Seott, Crust. Devon Cornwall, p.32 | 1909 P.J., Stappers in: Cellule, v.25 p. 368 |
1909 P.1., A. M. Norman & G. S. Brady in: Nat. Hist. Tr. Northumb., ser. 2 v. 311 p. 33.
Body rather slender, especially in G. Pseudorostrum prominent, acute,
in Q more than in CG. Antero-lateral corners unarmed. Carapace sculptured
with 2 pairs of oblique folds from the antero-lateral corners, one curving
close to the hind margin, the other in advance of it; free edges of cara-
pace quite smooth. Pedigerous segments smooth. Pleon slender, telson nearly
semicircular and slightly serrate in Q, fully as long as broad at the base
and smooth-edged in d. Eyelobe somewhat heart shaped with the 3 corneal
lenses larger and more distinct in GO than in ©. Antenna 1, 32 joint shorter
than 1st but longer than 2™4, at least in the QO, flagellum 3-jointed in ©,
4-jointed in Oo. Antenna 2, the long flagellum in the G variable in its
length. Peraeopod 1, 2"¢ joint about equal in length to rest of the limb,
slender 5* joint much longer than 6%, 6 than 7, Peraeopod 2, 7* joint
as long as 5% in ©, but shorter in G and tipped with a single hook (Sars),
2 apical and 1 subapical (Gilson). The rudimentary exopods of the 3¢ and
Ath peraeopods in the © are small, but distinctly 2-jointed, 2"¢ joint longer
than 1st. Pleopod 1 of GO with 4 strong setae distally on medial margin
of basal joint; pleopod 2 with 2 spines distally on lateral margin of that
joint. Peduncle of uropods equal to exopod in ©, shorter in d, endopod
much longer, with 10 spines on medial margin in ©, 14 in G; apical spine
of exopod very elongate; relative dimensions of peduncle and rami variable
(Gilson). Colour pale, variegated with dark brown stellate flecks. L. about
4mm. Gilson describes a form of the @, in which the 24 antenna is short,
and the 2"4 peraeopod has only 2 hooks at the distal end.
From N. coast of Norway round coasts of Europe to Sicily; depth, surface to 68 m.
4. P, ciliatum O. Sars 1879 P. ciliata, G. O. Sars in: Arch. Naturv. Kristian.,
v.4 p.71 t.43 | 1893 P. ciliatus, T. Stebbing in: Int. sci. Ser., #74 p.307 | 1900 P. ciliata,
G. O. Sars, Crust. Norway, v.3 p. 74.
Near to P. longicorne, but more robust, pseudorostrum shorter, apically
blunt; antero-lateral corners more produced, almost quadrate; sculpture of
carapace the same, but somewhat sharper; 2"¢ pedigerous segment with
broader side-plate; integument of pleon with a peculiar irregular reticulate
structure; telson almost rectangular, broader than long, distal margin tend-
ing to truncate. Eyelobe and eye and mouth-organs as in P. longicorne;
antenna 1 short and stout, 34 joint shorter than 2°¢, 2™4 than 1st. Peraeopod
1, 5t and 6t joints not slender. Rudimentary exopod of 3¢ and 4 pe-
raeopods indistinctly 2-jointed. Peduncle of uropod shorter than the exopod,
which is not very much shorter than the endopod; the latter carrying only 2
spinules near the apex but the whole medial margin closely ciliated; apical
Cum.: 17. Pseudocumatid., 6. Pseudoc., Strauchia, 18. Paraleuconidae, |. Paraleucon 151
spine of exopod not very elongate. Integument somewhat pellucid, with
light shade of fawn colour in front, and hinder part of carapace crossed by
a deep brown band of stellate spots. L. @ 3°5 mm, 3 unknown.
Mediterranean, Goletta; taken with P. longicorne.
P. lagunae Baker 1912 P. 1, C. F. Baker in: Rep. Laguna Mar. Lab., v1
p.106 f.57A—F.
Juy.? Carapace with strong transverse fold behind the large eyes, its sides with
83 curved ridges; telson as long as peduncle of uropods, ending subacutely. L. 1:5 mm.
Tide-pool, Laguna beach, Orange County, California.
Gen. Strauchia Czerniavski
1867 S., Czerniavski in: Syezda Russ. Kst., v.1 p. 64.
Telson very short, unarmed.
S. taurica Czern. 1868 S. ¢., Czerniavski in: Syezda Russ. Est., v.1 p.65 t.5 f.1.
It is stated, but probably in error, that the 1st peraeopod is shorter than the
2nd, and that only the 2nd and 84 peraeopods in the 2 have exopods. The statements
would be more likely to apply to the 34 maxillipeds and the 1st and 2nd peraeapods.
L. about 1 mm.
Black-Sea, off Cape St. John, depth 1-4 m.
18. Fam. Paraleuconidae
1912 Paraleuconidae, T. Stebbing in: Ann. 8S. Afr. Mus., v. 10v p.133, 156.
Pseudorostrum obliquely upturned in female, horizontal, truncate in.
male. Telson wanting. Second antenna in male with short flagellum. Man-
dible broad at the base. Mavxilla 1 with single filament to the palp.
Branchial leaflets few. Peraeopods 1—3 in female, 1—4 in male with
exopods. Only one pair of pleopods in male. Uropods with endopod in-
distinctly two-jointed.
1 genus.
1. Gen. Paraleucon Calman
1907 Paraleucon (Sp. un.: P. suteri), Calman in: Tr. zool. Soe. London, v. 181
p. 32, 36.
Character of genus included in that of the family for the present.
1 species.
1. P. suteri Calm. 1907 P.s., Calman in: Tr. zool. Soc. London, v. 181 p. 32,
36 t.9 f.1—20.
Pseudorostral lobes in Q directed obliquely upwards, upper margin
straight, apically acute, in GO horizontal, vertically truncate; antero-lateral
corners in Q forming a triangular tooth with faint serration of lower margin
behind it, in O rounded off. Median carina of carapace in © with a few
variable teeth and a large tooth near the hind margin overhanging a pre-
ceding depression, in the Ga similar arrangement feebler or obsolete. In
the © 3 subparallel ridges curve obliquely across the sides of the carapace,
the first on the frontal and pseudorostral lobes, the second passing from the
median depression towards the antero-lateral corner, the third from the hind
margin to the lower margin a little way behind the antero-lateral corners.
The first of these is wanting in the 3. Antenna 1, 3¢ joint not shorter
152 Cumacea: 18, Paraleuconidae, 1. Paraleucon, 19. Hemileuconidae, 1. Hemileucon
than 2"4, in d decidedly longer, flagellum 3-jointed, accessory 1-jointed. An-
tenna 2 of © 3-jointed, with 2 plumose setae on 1*¢ joint. Maxilliped 1
with 2 small papilliform branchial sacs. Peraeopod 1, 2°¢ joint nearly as
long as rest of limb. Peraeopod 2 with 34 joint distinct. The single pair
of pleopods very small, the rami minute, 1-jointed, each carrying a long
plumose seta. Peduncle of uropods shorter than the endopod, which is shorter
than the exopod; endopod indistinctly divided into 2 subequal joints, with
about 12 spines on the medial margin and 2 apical spines. L. 9 2:9 mm,
S 2:4 mm.
New Zealand, Lyttleton Harbour, depth 2—9 m; Akaroa Harbour, 11 m.
19. Fam. Hemileuconidae
1912 Hemileuconidae, T. Stebbing in: Ann. S. Afr. Mus., v.10v_ p. 133, 156.
Pseudorostrum obliquely upturned in female, horizontal, truncate in
male. ‘Telson wanting. Second antenna in male with short flagellum.
Peraeopods 1—3 in female, 1—4 in male with exopods. Pleopods wanting
in male. Uropods with endopod distinctly two-jointed.
1. Gen. Hemileucon Calman
1907 Hemileucon, Calman in: Tr. zool. Soe. London, v. 181 p. 32, 37.
Character of the genus included in that. of the family for the present.
2 species.
Synopsis of species:
Carapace with a single horizontal ridge on each side . 1. H. uniplicatus
Carapace with two oblique ridges on each side. . . . 2. H. comes
1. H. uniplicatus Calm. 1907 H. u., Calman in: Tr. zool. Soe. London, v. 181
p.37 t.9 f. 21—25.
Pseudorostral lobes straight, in © slightly upturned, obliquely truncate,
in GO horizontal, vertically truncate; antero-lateral angles produced to a tooth
in female, with slight serration in front and behind, in dO without produced
tooth. Carapace with nearly half of the median carina serrate, a lateral
ridge on each side running from the end of the frontal lobe longitudinally
and curving upwards to join the hind margin. Antenna | with 5-jointed
flagellum and one-jointed accessory. Peduncle of uropods rather more than
two-thirds the length of the subequal rami, the second joint of the endopod
bearing the same proportion to the first joint. L. O 26 mm, ¢ 2°35 mm.
New Zealand, Lyttleton Harbour, depth 2—9 m; Akaroa Harbour, 11 m.
2. H. comes Calm. 1907 H.c., Calman in: Tr. zool, Soc. London, v. 181 p. 38
t.9 £. 26—832.
Pseudorostral lobes straight, in © directed obliquely upwards, acute;
antero-lateral corners produced into a tooth, with slight serration of the
lower margin behind it. Carapace with irregular serration of the median
carina anteriorly and a depression towards the rear, an oblique curved ridge
crossing the side of the frontal lobe and the pseudorostral lobe to the middle
of its lower margin and subparallel to this pair another pair starting from
near the median line and approaching but not reaching the antero-lateral
tooth. First antenna as in A. wiplicatus, second antenna of & with flagellum
rather longer than in that species. Peraeopod 2 with 34 joint distinct.
Cumacea: 19. Hemileuconidae, 1. Hemileucon, 20. Lampropidae, 1. Lamprops 153
Pedunele of uropods a little shorter than the endopod, which is a little
shorter than the exopod; second joint of endopod rather more than two-thirds
as long as the first joint. L. O 2°38 mm, ¢ 2°7 mm.
New Zealand, Lyttleton Harbour; depth 2—9 m.
20. Fam. Lampropidae
1878 Lampropidae, G. O. Sars in: Arch. Naturv. Kristian., v.3 p. 462, 465 | 1883
L., G. O. Sars in: Forh. Selsk. Christian., 1882 nr. 18 p.11 | 1887 Z., G. O. Sars in:
Rep. Voy. Challenger, v.19 pars55 p.6,25 | 1899 L., (restricted), G. O. Sars, Crust.
Norway, v.3 p.17 | 1900 Z., Zimmer in: Fauna arctica, v.1 p. 414, 432 | 1908 Z., Zimmer
in: Ergeb. Tiefsee-Exp., v.8 p. 158,159,169 | 1912 ZL. (further restricted), T. Stebbing
in: Ann. S. Afr. Mus., v. 10v p. 133, 158.
Pseudorostral lobes not strongly produced. All the pedigerous segments
distinct. Telson well developed, with more than two apical spines. First
antennae with both flagella well developed. Second antennae of female more
conspicuous than usual. First maxillae with terminal filaments of palp two
or only one. First four pairs of peraeopods with exopods, which in the
female are rudimentary on the third and fourth pairs. No pleopods in either
sex. Uropods with endopod three-jointed.
4 genera.
Synopsis of genera:
{ Pseudorostral lobes with antero-lateral sinus. . . . 1. Gen. Lamprops
- \ Pseudorostral lobes without antero-lateral sinus — 2.
> J Maxilla 1 with single filament to the palp .. . . 2. Gen. Stenotyphlops ')
“ \ Maxilla 1 with two filaments to the palp — 8.
3 Maxilliped 3 with 5th joint not expanded ..... 3. Gen. Platytyphlops
; Maxilliped 3 with 5th joint expanded. ...... 4. Gen. Bathylamprops')
1. Gen. Lamprops 0. Sars
1863 Lamprops, G. O. Sars in: Nyt Mag. Naturv., v.12 p. 239 | 1865 Z., G. O.
Sars in: Forh. Selsk. Christian., 1864 p.188 | 1877 ZLampros, Meinert in: Naturh.
Tidsskr., ser.3 v.11 p.187 | 1879 Lamprops, A. M. Norman in: Ann. nat. Hist., ser. 5
v.3 p.68 | 1880 Lampros, Meinert in: Naturh. Tidsskr.. ser. 3 v.12 p.501 | 1899 Lamprops,
G. O. Sars, Crust. Norway, v.3 p.18 | 1900 Z., Zimmer in: Fauna arctica, v.1 p.414. 432 |
1905 Z., Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p.41 | 1908 Z., Zimmer
in: Ergeb. Tiefsee-Exp., v.8 p. 169,170 | 1912 Z., Calman in: P. U.S. Mus., v.41 p.605,
629 | 1912 Z., T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 159.
Pseudorostral lobes with antero-lateral sinus. Second and third pedi-
gerous segments rather large, with broad side-plates. Telson tapering, truncate
at the apex. Hye well developed. Second antenna of male with flagellum
of moderate length, more or less prehensile. Second peraeopods not much
shorter than first. In the uropods the first joint of the exopod is rather
unusually long.
3 accepted species and 2 incerti generis.
Synopsis of accepted species:
Eye present
I Of the 5 apical spines of the telson median the shortest 1. L. fuscata
Of the 5 apical spines of the telson median the longest — 2.
1) Of these genera, the male being unknown, the systematic position is provisional.
154 Cumacea: 20. Lampropidae, 1. Lamprops
{ Uropods with 1stjoint of exopod decidedly shorter than 2nd 2. L. fasciata
\ Uropods with the 2 joints of exopod subequal . . .. 3. L. quadriplicata
Species incerti generis:
Eye wanting
| Telson shorter than peduncle of uropods, with 5 apicalspines L.? comata!)
2 Telson as long as peduncle of uropods, with only 3 apical
Rj) ome pea oo 5 Gc chk sea Sous -.- iP beringi)
1. L. fuseata O. Sars 1865 L.7., G. O. Sars in: Forh. Selsk. Christian., 1864
p.192 | 1884 L.f, J. S. Schneider in: Tromso Mus. Aarsh., v.7 p.54 | 1887 L. 7,
H. J. Hansen in: Vid. Meddel., v.39 p.198 | 1899 L. 7, G. O. Sars, Crust. Norway, v.3
p.20 t.11 | 1900 Z. f., Zimmer in: Fauna arctica, v1 p. 414,432 | 1912 L. 7, Calman
in: P. U.S. Mus., v.41 p. 629.
Fig. 100.
L. fuscata (Q) Sars, an-
tenna 2.
Fig. 103.
L. fuscata (2)
Sars, telson.
Fig. 101. Fig. 102. Fig. 104.
L. fuscata (Jj) Sars, L. fuscata Sars, uropod. L. fuscata (d¢)
antenna 2. Sars, telson.
Pseudorostral lobes rather produced, acute, antero-lateral corners almost
obsolete. Carapace conically tapering and slightly carinate forwards, sides
quite smooth. Pedigerous segments combined about as long as the carapace.
1) Of these species, the male being unknown, the generic position remains in-
determinate.
Cumacea: 20. Lampropidae, 1. Lamprops 155
Pleon with telson scarcely as long as preceding part of the body; telson
in © tapering considerably, with 2 pairs of small lateral spines and 5 spines
at apex. the middle one the shortest, outer pair longest, divergent; telson
in G little tapering, the small lateral spines far apart, of the apical spines
the long outer pairs subequal, little divergent. Hye small, distinct. An-
tenna 2 of G unusually short, plainly prehensile, proximal section of flagellum
thick with small hooklike projections of the medial margin. 24 peraeopod
nearly as long as the 1S*, its 7 joint considerably longer than the 6. In
peraeopod 3 of Q the rudimentary exopod is rather conspicuous. Peduncle
of uropods as long as the telson, longer than the exopod, which is much
shorter than the endopod. Colour brownish, mottled in front with opaque
white. L. O 6 mm, GC scarcely 5 mm; Alaskan specimens?, ovigerous ©
8°7 mm.
Arctic; Lofoten Islands, Greenland, Franz Joseph Land; depths 6—75 m; off
Newfoundland, depth 121 m; Alaska?
2. L. fasciata 0. Sars 1863 L. f., G. O. Sars in: Nyt Mag. Naturv., v.12 p. 236 |
1865 L. f., G. O. Sars in: Forh. Selsk. Christian., 1864 p.191 | 1877 Lampros f., Meinert
in: Naturh. Tidsskr., ser. 3 v.11 p.187 | 1884 Lamprops f., J. S. Schneider in: 'Tromse
Mus. Aarsh., v.7 p.54 | 1886 L. f., A. M. Norman in: Rep. Fish. Board Scotl., v4
p. 163 | 1892 L. f., A. O. Walker in: P. Liverp. biol. Soe., v6 Tr. p. 104 | 1897 L. 7,
Ehrenbaum in: Wiss. Meeresunters., v.21 p.410 | 1899 L. 7, G. O. Sars, Crust. Norway,
v.3 p.19 t.9,10 | 1900 Z. 7, Zimmer in: Fauna arctica, v1 p. 414,432 | 1901 L. 7,
T. Scott in: Rep. Fish. Board Scotl., v.19 p. 274 | 1902 L. f., T. Scott in: Rep. Fish.
Board Scotl., v.20 p.510 | 1905 Z. 7, Calman in: Sei. Invest. Fish. Ireland, 1904
app. 1 p.4l.
Pseudorostral lobes obtuse, scarcely projecting beyond the blunt antero-
lateral corners. Carapace with 3 pairs of obliquely curved ridges crossing
the branchial regions. First 4 pedigerous segments combined longer than
the carapace. Pleon with telson longer than preceding part of the body.
Telson nearly twice as long as 6 pleon segment, rapidly tapering, with a
single pair of lateral spines, 5 apical spines, of which the divergent outer
pair and the central spine are much larger than the intermediate pair. Eye
conspicuous, dark red. Antenna 2 of d about half the length of the body,
not so conspicuously prehensile as in “. fuscata. Peraeopod 1, 7 joint
shorter than 6, Peraeopod 2, 7 joint subequal to 6%, 5% equal to both
combined. Peraeopods 3, 4. 5 densely setiferous. Peduncle of uropods scarcely
longer than the telson or the exopod, in which the first joint is rather
shorter than the second: the endopod is rather longer, its 1S* joint longer
than the 2™4 and 34 combined. A broad band of dark brownish violet crosses
the branchial regions and is sueceeded by other transverse bands. L. © 9 mm,
© considerably less.
Norway, N. to Vadsé, Kattegat, Heligoland, E. and W. Scotland, Irish Sea; at
no great depths.
3. L. quadriplicata S.I.Sm. 1879 L. q., 8.1. Smith in: Tr. Connect. Ac., v.51
p.118 | 1893 L.q., T. Stebbing in: Int. sci. Ser., v.74 p.309 | 1900 L.q., Zimmer
in: Fauna aretica, v.1 p. 415,432 | 1912 DL. qg., Calman in: P. U.S. Mus., v.41 p. 628.
Pyeudorostral lobes truncate, not produced beyond the slightly projecting,
obtuse, sometimes denticulate, antero-lateral corners. Carapace with 4 pairs
of very distinct, arcuate, nearly parallel ridges, the first from antero-lateral
corners to the sides of the frontal lobe, the second extending behind that
lobe from the lower margins, the third and fourth similarly, the fourth
156 Cumacea: 20. Lampropidae, 1. Lamprops, 2. Stenotyphlops
nearly touching the hind margin above. First 4 pedigerous segments com-
bined subequal in length to the carapace. Pleon with telson longer than
preceding portion of the body. Telson as long as 5th and about twice as
long as 6t pleon segment, with 2 or 3 pairs of lateral spines, and 5 apical
of similar size, only the median spine a little the largest. Antenna 1, 1%*
joint of principal flagellum only a little shorter than the accessory. Peraeopod 1
not very elongate, 5%, 6, and 72 joints subequal in length. Peraeopod 2,
6t and 7t joints subequal, 5th joint subequal to both combined. Peraeo-
pods 3, 4, with 3 or 4 plumose setae on apex of the rudimentary exopod
in 5 and others on the 1st joint of it, the exopods in the C fully deve-
loped. The G without pleopods, in accord with the generic character. Peduncle
of uropods as long as the telson and also subequal to the endopod, the
exopod a little shorter, its 2 joimts subequal; medial margin carrying 10 spines
on peduncle, 16, 5, 2 on the 3 joints respectively of the endopod. LL. ©
9mm, CG not fully mature 8-5 mm.
Gloucester Harbor, Massachusetts, 13-18 m, and Casco Bay, surface; off New-
foundland, depth 67 m.
L.? comata Zimmer 1907 Z.?c¢., Zimmer in: Zool. Anz., v.31 p. 373.
Pseudorostral lobes very short, obtuse. Carapace dorso-ventrally somewhat
depressed; a depression between the eyelobe and the frontal lobe. Pedigerous
segments combined shorter than the carapace. Pleon decidedly longer than
preceding part of the body. Telson about three-fourths as long as peduncle
of uropods, with 5 pairs of small spines on the distal third, and 5 very
strong apical spines. Eye entirely wanting. Antenna 1 almost three-fourths as
long as the carapace, 1S* joint of principal flagellum only a little shorter than the
accessory. Peraeopod 1 elongate, 6% joint rather longer than 5%. Peraeo-
pod 2, 7% joint twice as long as 6 and half as long as 5%. The 2-jointed
exopod of 3¢ peraeopod more strongly developed than is usual in the © of
this family. Peduncle of uropods nearly as long as 5% pleon segment;
endopod with close row of unequal spines on the medial margin. Colour
white. L. about 7mm. GO unknown.
Antarctic: Lat. 65°15 S., long 80° 12‘ E., depth 3423 m.
L. (?) beringi Calm. 1912 Z. (?)6., Calman in: P. U.S. Mus., v.41 p. 605, 630
tf. 27, 28.
Like Hemilamprops uniplicata (p. 58) in having a single pair of lateral
ridges on the carapace, but with well marked sinus to the pseudorostral
lobes defined by acutely produced antero-lateral angle; telson, as long as
pedunele of uropods with 5 or 6 spines on each side not symmetrically
arranged, and 3 apical spines, the median the longest: 1%* peraeopod not
very elongate, with 5%, 6%, and 7 joints subequal; exopod of uropods
only slightly shorter than endoped with its 1st joint scarcely half as long
as the ond, L., ovigerous ©, 12°7 mm. CG unknown.
Bering Island.
2. Gen. Stenotyphlops T. Stebbing
1912 Stenotyphlops, T. Stebbing in: Ann. 8. Afr. Mus., v.10v p. 159, 162.
Carapace narrow, eyelobe without visual elements, pleon slender, telson
carrying 3 apical spines. First antenna with both flagella elongate. Second
antenna of female four-jointed. Palp of first maxilla with a single filament.
Cumacea: 20. Lampropidae, 2. Stenotyphlops, 8. Platytyphlops i Mey7k
First maxilliped with terminal joint peculiarly widened at the base. First and
second peraeopods of the female with exopods, third and fourth pairs with
4
4
i
Fig. 105.
S. spinulosa (2)
Stebbing.
exopods very rudimentary. Fifth peraeopods apparently
wanting.
1 species.
1. 8, spinulosa Stebb. 1912 S. spinulosus, T. Stebbing
in: Ann. S. Afr. Mus., v.10v p. 162 t. 60.
Surface spinulose. Pseudorostral lobes slightly up-
turned, without sinus of antero-lateral margin. Carapace
narrowly oval, with median keel running for some distance
backwards from the small eyeless eyelobe, and behind the
middle of the earapace’s length flanked by a pair of raised
ridges. Fifth pedigerous segment not wider than the pleon.
Telson about 4/7tts of the peduncle of the uropods, the
margins serrate and behind the middle having 3 pairs
of lateral spines. First antenna with 1S* joint much
longer than 2™4 and 3¢ combined, principal flagellum
4-jointed, accessory 3-jointed, about 4/;%S of principal.
Second antenna 4-jointed, the slender
Ath joint as long as the stout 1%*.
Third maxilliped with 2™¢ joint not
distally expanded and 5 also nar-
row. Peraeopod 2 slender, 2" joint
slightly shorter than rest of limb,
needle-like 7*® about thrice as long
as 6, both combined scarcely as
long as the well spined 5%, Third
and fourth peraeopods much shorter
than the second, their 2%4¢ joint car-
rying a minute 2-jointed exopod. Of
fifth peraeopods no trace. Margins
of uropods serrulate, endopod 4/,'s
as long as the peduncle, its 18* joint
about twice the 24 and 34 combined,
spines on the medial margin respec-
tively 10, 3, 1; the exopod, with 1%*
joint little more than 1/;'s of the
2nd. reaches nearly to the middle of
the endopods 34 joint. L. 12 mm. Di me ve a
S. Africa, Cape Point EK. by N.29 miles. . Saeeaie i a
3. Gen. Platytyphlops T. Stebbing
1912 Platytyphlops (Sp. typ.: P. peringueyi), T. Stebbing in: Ann. S. Afr. Mus.,
v. LOv p. 158, 159.
Carapace broad, depressed, without antero-lateral angles. Pleon long
and slender.
Telson with three apical spines. Eye wanting. Antenna 1
with both flagella well developed. Antenna 2 of female 4-jointed. Palp of
first maxilla with two filaments. First and second peraeopods in female with
exopods, third and fourth pairs with microscopic two-jointed rudiments. Fifth
peraeopods rudimentary. Pleopods wanting in both sexes.
2 species.
158 Cumacea: 20. Lampropidae, 3. Platytyphlops
Synopsis of species:
{ Carapace with pair of conspicuous dorsal processes .. . 1. P. peringueyi
| Carapace without conspicuous dorsal processes. . . . - . ¥%. P. orbicularis
1. P. peringueyi Stebb. 1912 P.p., T. Stebbing in: Ann. 8. Afr. Mus., v. 10v
p. 159, 162 t. 58, 59.
Pseudorostral lobes short, upturned. Carapace rounded oval. rather
longer than broad, the carinate margin fringed with pellucid scales, ocular
lobe small, triangular, frontal lobe broad, medio-dorsal line arched, finely
serrate to beyond the middle of the carapace, which in its hinder part has
a submedian pair of carinae rising into thin round-topped processes. Pedi-
gerous segments combined equal about half the length of the carapace, the
first shorter and much narrower than the second, the second and third
depressed and spread out at the sides, though still much narrower than the
carapace, the fifth not wider than the pleon, which is much longer than the
preceding part of the body. Telson much longer than the preceding segment,
shorter than the peduncle of the uropods. with
anal opening near the base, thence narrowing to
the truncate 3-spined apex; 3 pairs of spines are
spaced on the margins of the hinder half, the
largest pair being subapical. Antenna 1, peduncle
rather longer than the flagella, its 18¢ joint more
than twice as long as the 2™4, 34 a little shorter
than 2™4, principal flagellum 5-jointed (or in ©
perhaps 6-jointed), accessory 4-jointed. Antenna 2
of Q with 3 plumose setae on 1%* joint, which is
rather large, but much shorter than the very slender
4th, 34 with tooth near the base. Upper lip emar-
Fig. 107.
P. peringueyi (2) Stebbing, an- ~, : ; ; ;
tenna 2. ginate. Numerous spines in spine-row of mandibles.
Filaments of palp of maxilla | very unequal. Epi-
lord
pod of maxilliped 1 with 7 unequal branchial leaflets. First peraeopods
reaching much beyond carapace, second joint in first and second pairs shorter
than following joints combined, 6* joint longest of these in the first pair but
very short in the second; third and 4 pairs alike in d and Q, except for
minuteness of exopods in the 9. Fifth peraeopods pellucid, vestigial, but
the 2.4 joint, as in the 2 preceding pairs, longer than the following joints
combined. Peduncle of uropods a little longer than the endopod, the 45
joints of which have on the medial margin 8 or 9, 3 or 4, and 2 spines
respectively; the exopod is a little longer than the telson, which equals the
first 2 joints of the endopod. L. about 10 mm.
South Africa, Cape Point E. by N. 29 miles, and 81° KE. 32 miles.
2. P. orbicularis (Calm.) 1905 Platyaspis 0., Calman in: Sci. Invest. Fish.
Ireland, 1904 app. 1 p.12,42,43 t.5 f.77—81L | 1912 Paralamprops o.?, Calman in: P.
U.S. Mus., v.41 p.631 tf. 29—39 | 1912 Platytyphlops o., T. Stebbing in: Ann. S. Afr.
Mus., v. 10v p. 144, 161.
Like P. peringueyi, but carapace rather broader than long, 1:75 long
by 18mm broad; a pair of low submedian carinae on the hinder half;
1st pedigerous segment larger than those which follow it; antenna 1 with
both flagella 4-jointed; antenna 2 © with 2 plumose setae on the small
1st joint, 4*" joint narrowly fusiform; first peraeopods (as figured) not longer
than the carapace. The specimen defective beyond the 1%* pedigerous seg-
ment. Complete specimens, of which the specific identity is rather uncertain,
Cumacea: 20. Lampropidae, 3. Platytyphlops, 4. Bathylamprops, 21. Dicidae = 159
have breadth of carapace a little less than the length, with minute teeth
along from eyelobe to beyond the middle, with no paired ridges to the rear;
telson only a little longer than the 6 pleon segment; antenna 1 with
principal flagellum 6- or 7-jointed, accessory 4-jointed; antenna 2 © as in
P. peringueyt, but 3% joint without tooth, 4! more than twice as long as 3°;
peraeopods as in P. peringueyi; endopod of uropods about as long as the
peduncle. L. © with developing oostegites 16 mm.
Atlantic coast of N. America, S. of Labrador, depths between 600 and 1000 m;
W. coast of Ireland, 77 miles W. N. W. Achill Head, county Mayo, depth 698 m.
4. Gen. Bathylamprops Zimmer
1908 Bathylamprops (Sp. un.: B. calmant), Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 173.
Pseudorostral lobes acutely and much produced. Telson well developed.
Eye wanting. Antenna 1 probably long. Antenna 2 of female with terminal
joint elongate. Palp of maxilla 1 with two filaments. Epipod of maxilliped |
with few (5) branchial sacs. Maxilliped 3 with 5‘ joint much widened.
1 species.
1. B.calmani Zimmer 1908 B.c., Zimmer in: Ergeb. Tiefsee-Exp., v.8 p. 173
t.40 f. 60—62; t.41 f.63—70.
Integument thin, brittle, squamose.
Pseudorostral lobes produced to about
one-fifth of the length of the carapace:
antero-lateral corners obsolete. Ocular
lobe forming a narrow equi-lateral triangle,
without visual elements. Carapace about
as long as the pleon without its telson,
wrinkled by numerous denticulate swel-
lings, chiefly transverse but in part con-
nected by others at angles with them.
First pedigerous segment not elongate.
Pleon segments 1—5 with 3 longitudinal
denticulate swellings. Antenna 2, 1S* joint
nearly spherical, with plumose seta on a
conical process, 4% nearly as long as
preceding 3 joints combined. Peraeopod
1 slender, rather long, 5 joint slightly
longer than 6%. Peduncle of uropods
very slender, subequal to pleon segments
4—6 combined. Colour white. L. 9 at least 13 mm. GC unknown.
Lat. 6° 12‘ S., long. 41° 17’ E., depth 2959 m off Dar-es-Salaam.
Fig. 108.
B. calmani (2 juv.) Zimmer, maxilliped 3.
21. Fam. Dicidae
1910 Dicidae, T. Stebbing in: Ann. S. Afr. Mus., v.61v p.415 | 1912 D., Calman
in: P. U.S. Mus., v.41 p.608 | 1912 D., T. Scott in: Sci. Invest. Fish. Scotl., 1911 nr. 1
p-5 | 1912 D., T. Stebbing in: Ann. S. Afr. Mus., v.10v p. 133, 147.
Telson long, cylindric to the triangular apex, without apical spines.
Exopods on first four pairs of peraeopods at least of the male. Uropods
with three-jointed endopod, not longer than the exopod.
1 genus.
160 Cumacea: 21. Dicidae, 1. Dic
1. Gen. Dic T. Stebbing
1910 Dic, T. Stebbing in: Ann. S. Afr. Mus., v. 61v p. 415 | 1912 Diastylis (part.),
Calman in: P. U. S. Mus., v.41 p.608 | 1912 Dic, T. Scott in: Sci. Invest. Fish. Scotl.,
1911 nr.1 p.5| 1912 D., T. Stebbing in: Ann. 8. Afr, Mus., v. 10v p. 147.
Sixth segment of pleon very short; telson very much longer than that
segment and longer than the peduncle of the uropods.
2 species.
Synopsis of species:
{ Uropods with exopod and endopod equal in length . . 1. D. calmani
\ Uropods with exopod much longer than the endopod . 2. D. tubulicauda
1. D. calmani Stebb. 1910 D.c., T. Stebbing in: Ann. S. Afr. Mus., v. 61V
p.416 t.46 B, 47.
Pseudorostral lobes down bent. Carapace in front with several obliquely
transverse markings. Third and fourth pedigerous segments dorsally coalesced.
goRjans
UO SSULYLeUL
SOATVA [BURA
Fig. 109. Fig. 110.
D. calmani Stebbing, maxilliped 3, left and right. D. calmani Stebbing.
Telson as long as the 3 preceding segments combined, apex cut into
10 denticles. Antenna 1, principal flagellum small, 4-jointed, accessory
minute, 2-jointed. Antenna 2, flagellum 5-jointed, the specimen no doubt
not fully adult. Mandible with numerous spines. Maxilla 1 with 2 filaments.
Maxilliped 3, 2"4 joint distally dilated, 3¢ of unusual size and shape, the
broadly rounded end nearly reaching distal end of the 5. Peraeopod 1, rather
long, 284 pair not more than a third as long, but with stout 27¢ jomt. Exo-
pods of 3¢ and 4* pairs 2-jointed, shorter than the 24 joint, probably not
fully developed. Peraeopod 5 short. Peduncle of uropods more than 2/98 ag
long as the telson, longer than the exopod, which is equal to the endopod; of this
the spines on medial margin of the 3 joints are respectively 4, 3, 3. L. d
not fully adult, about 5mm. A specimen 2°75 mm long, is without 5 peraeopod.
S. Africa lat. 32° 53/ 30” S., long. 28° 11/00” E., depth 75 m.
Cumacea: 21. Dicidae, 1. Dic, 22. Gynodiastylidae, 1. Gynodiastylis 161
2. D. tubulicauda (Calm.) 1905 Diastylis t., Calman in: Sci. Invest. Fish,
Ireland, 1904 app.1 p.10,46 t.5 f.82—86 | 1910 Dic ? t., T. Stebbing in: Ann. S. Afr.
Mus., v.61v p.416 | 1912 D.t., T. Scott in: Sci. Invest. Fish. Scotl., 1911 nr.1 p.4 t.1
f.6 | 1912 D.t., T. Stebbing in: Ann. S. Afr. Mus., v.10v p. 147.
Pseudorostral lobes acute, prominent, horizontal. Carapace spinulose.
Pedigerous segments 3, 4, quite distinct one from the other. Telson as
long as the 4 preceding segments combined, apex sharply triangular, with
2 apical spinules. Peraeopods 3, 4, very long. Peraeopod 5 quite wanting,
probably a juvenile character. Peduncle of uropods 7/48 as long as telson, much
less than twice the exopod, but considerably more than twice the endopod,
which has 2, 1, 1 spines respectively on medial margin of the 3 joints, and
2 apical spines. L. juv., 5-4 mm, sex doubtful.
Ireland, County Mayo, 77 miles W. N. W. of Achill Head, depth 699 m; lat. 59°
25' N., long. 70° 33’ W., depth 1100 m.
22. Fam. Gynodiastylidae
1912 Gynodiastylidae, T. Stebbing in: Ann. 8, Afr. Mus., v. 10v p. 138, 146.
Telson of moderate size, with no apical spines or 2 that are rudimen-
tary. Accessory flagellum of 1st antenna very small. Second antenna small
in both sexes. Palp of maxilla 1 with 2 filaments. Branchial leaflets few or
wanting. Maxilliped 3 without exopods in the female. Peraeopods 1—4 or only
1st and 24 in the male with exopods, and only 18t and 24 in the female.
No pleopods in the male. Uropods with the endopod two-jointed or simple.
1 genus.
1. Gen. Gynodiastylis Calman
1911 Gynodiastylis, Calman in: Tr, zool. Soc. London, v.181v p. 312, 366 | 1912
G., T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 146.
At present with the characters of the family.
4 species.
Synopsis of species:
1 { Uropods with the endopod simple ..+..-+-++-+- > 1. G. laevis
~ \ Uropods with the endopod two-jointed — 2.
9 { Uropods with first joint of endopod longer than second . 2. G. carinatus
\ Uropods with first joint of endopod not longer than second — 3.
3 J Carapace with numerous longitudinal ridges... -.- - 3. G. costatus
\ Carapace with a pair of dorso-lateral ridges... .-+.- - 4, G. bicristatus
1. G. laevis Calm. 1911 G.1., Calman in: Tr. zool. Soc. London, v. 181v p. 342,
367, 371 t. 35 f. 832—39 | 1912 G.1, T. Stebbing in: Ann. S. Afr. Mus, v. 10v p. 147.
Pseudorostral lobes short, slightly deflexed, apically truncate, sinus
slightly defined by tooth in Q, by rounded prominence in G. Surface of
carapace quite smooth, eyelobe slightly prominent. Pedigerous segment 1
very short, 24 very long, with side-plate produced anteriorly as a narrow
lobe, 34 and 4 with side plates extended backwards. Pleon with faint
lateral keels, 6 segment little shorter than 5‘; telson little more than half the
length of the preceding segment. Eyelobe shows neither pigment nor distinct facets.
Antenna 1 short, 2" joint of peduncle about °/,4% of 3¢, the small flagella
each 3-jointed in Q, the principal 4-jointed in the G. Antenna 2 of Go
Das Tierreich. 39. Lief.: T. R. R. Stebbing, Cumacea. 11
162 Cumacea: 22. Gynodiastylidae, 1. Gynodiastylis
apparently much as in G. carinatus. Peraeopod 1, 2™4¢ joint nearly twice as
long as rest of limb, distally produced as a strong tooth, 6th half as long
as 5th, with only 3 long setae on the sub-distal margin, 7th °/,4 of 6%.
Peraeopod 2, 6t and 7 joints subequal, together shorter than 5‘, Peraeo-
pods 3, 4 without exopods in either sex. Peduncle of uropods °*/,45 of 6"
pleon segment, exopod about as long as peduncle (text), shorter in fig., endo-
pod simple, a little longer than the exopod. L. 9 4:1 mm, ¢ (?adult) 2-4 mm.
N. Zealand, Lyttleton Harbour, depth 2—9 m.
Fig. 111. Fig. 112.
G. laevis (2) Calman, G. carinatus (dQ) Cal-
telson and uropod. man, telson and uropod.
2.G. earinatus Calm. 1911 G. carinata, Calman in: Tr. zool. Soe, London, v.181v
p. 342, 367, 388 t.35 f.6—31 | 1912 G. carinatus, T. Stebbing in: Ann. S. Afr. Mus.,
v. 10v p. 146.
Pseudorostral lobes with edges acute, but apex truncate, sinus defined
by slightly produced antero-lateral tooth in Q, by rounded slightly serrate
projection in G. Carapace with 5 or 6 pairs of longitudinal ridges, eyelobe
less than twice as broad as long. Of the pedigerous segments all 5 in GC,
last 2 in ©, have a pair of longitudinal dorso-lateral ridges, segment 3 has
the side-plates expanded forwards and backwards, the attachment of the
limbs in the Q widely separating the 2%¢ and 34 pairs. Pleon with pair of
lateral ridges. Pleon segment 5 in Q not much longer than the 6"; telson
2/48 ag long as the preceding segment, ovate, the bluntly pointed apex having
2 extremely minute setiform points. Eye without pigment, but with 3 large
corneal lenses. Antenna 1 of Q with 2"4 joint of peduncle much the shortest,
both flagella very small. Antenna 2 of © short, 3- or 4-jointed, in the 3
very short, the large setose terminal joint of the peduncle being nearly as
long as the 8-jointed flagellum. Maxilliped 1 almost or quite without
branchial leaflets. Peraeopod 1, 2™4 joint about 3/4ts of rest of limb, 5‘ more
than twice the 6, which is less than twice the 7* and carries on the
oblique subdistal border a series of 10 remarkably long smooth setae. Peraeo-
pod 2 with last 3 joints subequal in length. Peraeopods 3, 4, without
exopods in either sex. Peduncle of the uropods a little shorter than pleon
segment 6, exopod about 4/gts of peduncle, endopod about 4/5 of exopod in
the Q, its 24 joint not half as long as the 1, but in dO the peduncle is
not longer than the exopod, this being little longer than the endopod, in
which the 2™4 joint is nearly as long as the 1%. L. 9 4mm, ¢ 2°85 mm.
N. Zealand, Lyttleton Harbour, depth 2—9 m.
Cumacea: 22. Gynodiastylidae, 1. Gynodiastylis, 23. Heteroleuconidae 163
3. G. costatus Calm. 1911 G. costata, Calman in: Tr. zool. Soc. London,
v.181v p.342, 368, 372 t.36 f.1—10 | 1912 G. costatus, T. Stebbing in: Ann. S. Afr. Mus.,
v.10v p. 147.
Pseudorostral lobes and carapace much as in G. carinatus, but the
ridges more numerous, some ill defined and interrupted, and just behind the
sharp antero-lateral corners a short vertical pair, forming a tubercle behind
the sinus, are continued upwards, carrying a series of thin, transparent, curved
spines. Pedigerous segments of the Q much as in G. carinatus, but only the
third with a pair of slightly marked dorso-lateral ridges. Pleon segment
6 little shorter than 5; telson about 3/4%s of 6%, with 2 extremely minute
setae or spinules at the bluntly pointed apex. Eye without pigment. An-
tenna 1, 2™4 joint only half as long as 34. Antenna 2 in Q short, 4-jointed;
in the G also very short. Peraeopod 1, 2™4 joint scarcely half rest of limb,
5th joint twice the 6, which has about 11 long setae on the oblique distal
part, 7* joint more than half the 6. Exopod on all the first 4 pairs of
peraeopods in the G, on only the first 2 in the Q. Pedunecle of uropods
longer than 6' pleon segment, exopod in © nearly as long as peduncle, a
little shorter than the endopod, of which the 1%* joint is slightly shorter
than the 2™4, but in G the peduncle is twice as long as the subequal rami.
io, 2:35 mm. o 1-9 mm.
Gulf of Siam, Koh Kam, 87m; N. of Koh Chuen, 27m; between Koh Mesan
and Cape Liont, depth 9—15 m.
4. G. biecristatus Calm. 1911 G. bicristata, Calman in: Tr. zool. Soc. London,
v. 181v_ p. 342, 368, 374 t. 36 f. 11—22 | 1912 G. bicristatus, T. Stebbing in: Ann. 8. Afr.
Mus., v.10v p. 147.
Pseudorostral lobes short, truncate, in © horizontal, a little depressed
in G, sinus well defined. Carapace with pair of strong cristiform ridges,
diverging from hind margin forwards and then converging on to the sides
of the pseudorostral lobes; eyelobe broad. Pedigerous segments each with
pair of dorso-lateral crests, indistinct on the 5%; side-plates of 3¢ not greatly
expanded, with no great interval between 2™4 and 3¢ peraeopods. Pleon segment
5 little longer than the 4, Telson a little shorter than the preceding segment.
Eye without pigment. Antenna 1, 24 joint more than half the 34. An-
tenna 2 of © 3-jointed, of GO very short. Maxilliped 1 apparently without
branchial leaflets. Peraeopod 1, 24 joint about half rest of limb, 5 more
than twice 6%, which has distally about 9 very long setae, 7 little over
half the 6". Exopods on all the first 4 pairs of peraeopods in the Go, only
on first 2 in @. Peduncle of.uropods about twice as long as the telson,
exopod shorter than endopod, which is considerably shorter than the peduncle,
with 1s* joint shorter than 2"4. L. Oo 1:9 mm, o& 17 mm.
Japan, 73m; Gulf of Siam Koh Kam, 9—18 m; between Koh Mesan and Cape
Liont, depth 9—15 m.
23. Fam. Heteroleuconidae
1912 Heteroleuconidae, 'T. Stebbing in: Ann. 8. Afr. Mus., v.10v_ p. 133, 156.
Telson wanting. Second antenna in female three-jointed, the flagellum
in male very short. Mandible with base broad, molar obtuse. First maxilla
with single filament to the palp. Only the first two pairs of peraeopods
with exopods in either sex. No pleopods in male or female. Uropods with
endopod simple.
1 genus.
nite
164. Cumacea: 23. Heteroleuconidae, 1. Heteroleucon, 24. Nannastacidae
1. Gen. Heteroleucon Calman
1907 Heteroleucon (Sp. un.: H. akaroénsis), Calman in: Tr. zool. Soc. London,
v. 181 p. 32, 34.
For the single genus the characters of the family for the pre-
sent suffice.
1 species.
1. H. akaroénsis Calm. 1907 H.a., Calman in: Tr. zool. Soe. London, v.18t
p. 34 t.8 f. 11—23.
Q. Pseudorostral lobes directed obliquely upwards with upper margin
straight, apically acute; antero-lateral corners preceded by a double denticle
and small rounded sinus, then beginning a serrate division of the lower
margin. Median line of carapace having the carina serrate with 8—10 teeth
in front half and one larger tooth close to hind margin. Telsonic segment
little produced between bases of uropods. Antenna 1 with first 2 joints
stout, 3¢ shorter and narrower, flagellum 2-jointed, accessory 1-joimted. An-
tenna 2 three-jointed, with 2 setae on the 1%* joint. Maxilliped 1 with
only 2 small papilliform branchial sacs. Peraeopod 1 with 24 joint shorter
Fig. 113. Fig. 114.
H. akaroénsis (2) Calman, antenna 1. H. akaroénsis (2) Calman, antenna 2.
than rest of limb. Peraeopod 2 with very short 3¢ joint. Peduncle of uro-
pods rather shorter than the subequal rami, endopod with about 10 spines on —
medial margin and a strong, dorsally curved, apical spine. L. 2°75 mm.
3S. Pseudorostral lobes very short, horizontal, apically rounded; antero-
lateral corners broadly rounded and serrated. Median line of carapace smooth,
except for one or two small serrations anteriorly. Antenna 1 with 3-jointed
flagellum. Antenna 2 with flagellum very short, not longer than peduncle,
9-jointed. Peduncle of uropods little more than half as long as the rami.
L. 165 mm.
New Zealand, Akaroa Harbour, depth 11m (both sexes), Lyttleton Harbour,
depth 2—9 m (females).
24. Fam. Mannastacidae
1866 Nannastacidae, Bate in: Zool. Ree., v.2 p.329 | 1878 & 1879 Cumellidae,
G. O. Sars in: Arch. Naturv. Kristian., v0.3 p.466; v4 p.92 | 1880 C@., Kossmann,
Reise Roth. Meer., Malac. pars 2 p.90 | 1887 C., G. O. Sars in: Rep. Voy. Challenger,
v.19 pars55 p.62 | 1893 Nannastacidae, T. Stebbing in: Int. sci. Ser., v.74 p. 305,
312 | 1895 N., H. J. Hansen in: Ergeb. Plankton-Exp., v.2 G.c. p.59 | 1897 Cumellidae,
Ehrenbaum in: Wiss. Meeresunters., v.21 p.419 | 1900 Nannastacidae, G. O. Sars, Crust.
Norway, v.3 p.79 | 1900 N., Zimmer in: Fauna arctica, v.1 p. 428, 431 | 1900 V., T. Stebbing
in: Willey, Zool. Results, pars5 p.611 | 1904 N., Calman in: Herdman, Rep. Ceylon
Cumacea: 24, Nannastacidae, 1. Schizotrema 165
Pearl Fish., v.2 p.176 | 1905 N. (part.), Calman in: Sci. Invest. Fish. Ireland, 1904
app. 1 p.22 | 1905 N. (part.), Calman in: Siboga-Exp., nr. 36 p.10 | 1906 N. (part.),
Calman in: Mt. Stat. Neapel, v.17 p.417 | 1907 N. (part.), Zimmer in: Zool. Anz., v.31
p. 369 | 1911 N. (part.), Calman in: Tr. zool. Soc. London, v.181v p.341 | 1912 N.,
T. Stebbing in: Ann. 8S. Afr. Mus., v. 10v p. 133, 164.
Pseudorostral lobes with antero-lateral corners well expressed. Telson
wanting. Eyes sometimes developed as a distinctly separated pair. Accessory
flagellum of first antenna very small. Second antenna of female one-,
two-, or sometimes three-jointed. Mandibles normal. First maxilla with
two filaments to the palp or only a single filament. First four pairs of
peraeopods in male with exopods, in the female only the first two pairs or
none. No pleopods in either sex. Uropods with the endopod simple.
6 genera accepted, 1 doubtful.
Synopsis of accepted genera:
{ Eyes, when present, paired — 2.
| Eyes, when present, not paired — 4.
o J Exhalant respiratory orifices paired - ... . . . 1. Gen. Schizotrema
Exhalant respiratory orifice single — 3.
First and second peraeopods of the female without
EXOPOUS#epienr stem, aitea rs! siren Teer 5 . . . 2. Gen. Paranannastacus
First and second peraeopods of the female with
EXOMOUSH AUS sh ate a ees or et-s ~ .«... 8. Gen. Nannastacus
Sino leit amOnbrrceuegpen ce rel Mie eter enters) so) ae 4. Gen. Platycuma
Alimentary canal not coiled, “1st iiawilla with 2
filaments — 5.
Carapace more or less overhanging anterior pedi-
PETOUSHSCOMENtSS ee bot eye) t= eie fel . . 5. Gen. Cumellopsis
Carapace not overhanging the anterior peieernns
al
Alimentary canal spirally coiled; 18t maxilla with
4 pieyea ca OURS RE GID Geen a ech Net epee 6. Gen. Cumella
1. Gen. Schizotrema Calman
1911 Schizotrema, Calman in: Tr. zool. Soc. London, v.181v p. 341,360 | 1912
S., T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 164, 165.
Respiratory orifices paired, wide apart. Pseudorostral lobes not meeting
in front. Carapace somewhat depressed, not or little overhanging pedigerous
segments. Third maxillipeds and first and second peraeopods with exopods
in female. Other characters as in Nannastacus (p. 168).
4 species.
Synopsis of species:
{ Uropods with peduncle longer than the endopod. . . . . 1. S, calmani
\ Uropods with peduncle not longer than the endopod:— 2.
9 { Uropods with endopod little longer than the peduncle . . 2. S. depressum
\ Uropods with endopod very much longer than the peduncle — 8.
Telsonic segment longer than preceding segment, apically
3 | CULCRee ce Sess os Gn or ihel coiretnon cats te re) oloie = . 3.8. bifrons
| Telsonic segment Sorter aa Heeceding peoaens apically
mONEG! GI OG 6 Gea 6 co 5d 05 6 pio of Bock -..~. 4. §. sordidum
166 Cumacea: 24. Nannastacidae, 1. Schizotrema
1. S. calmani Stebb. 1912 S.c., T. Stebbing in: Ann.S. Afr. Mus., v.10 v p.165 t, 61.
Pseudorostral lobes upturned, widely divergent. Carapace with small
median eyeless eyelobe, median line finely denticulate and setulose, lateral
margins fringed with denticles conspicuous anteriorly, soon
dwindling away. Pedigerous segments 2—5) with laterally
flattened edges cut into denticles. Pleon segments 1—5
with dorsal and ventral lines of denticulation and lateral
ridges, 5 segment long, distally narrowed, telsonic seg-
ment short. Antenna 1, 3¢ joint shorter than 2™4,
Maxilliped 1 with last joint slender; branchial leaflets
about seven. Maxilliped 3 with 6 joint curved, apically
a little widened. Peraeopods 3, 4, with 2"4 joint piri-
form, the rest very slender. Peduncle of uropods denti-
culate along 3 edges, twice as long as the endopod; exopod
about 4/58 of endopod. L. d about 2°5 mm. © unknown.
S. Africa, Cape Natal distant N. by E. 24 miles, 805 m.
Be
2. 8. depressum Calm. 1911 S.d., Calman in: Tr. zool.
aes Soe. London, v. 181v p. 341, 361 t. 34 f. 14—17.
reeds ala lia,
8. calmani (3) Steb- Pseudorostral lobes short, truncate, obliquely up-
pace: pedigerous seg, turned, each surrounding the branchial orifice from which
ments. a long branchial siphon is protrusible; front margin nearly
vertical, antero-lateral corner rounded, ending in curved
spine, from which a dwindling series runs back, with 2 or 3 large ones at the
end. Carapace very broad and depressed, minutely granular, with long setae on
various parts of this section and the body in general. Pedigerous segments
broad and depressed, with large spines on the side-plates. Pleon short, all
segments but the 1S* with stout lateral spines, the telsonic segment a little
shorter than the 5, rotundo-quadrate. Eyes dark, set just behind the
branchial orifices. Antenna 1, 3¢ joint longer than 2°¢, flagellum 2-jointed.
Antenna 2 of © with 2 teeth on anterior margin of the basal part, the apex
conically produced. Maxilliped 3 with 5 and 6 joints somewhat widened.
Peraeopods 3—5 short and stout, in the 34 pair the 5 joint hardly longer
than the 4t and shorter than the 6%, in the 5% pair the 5% joint about
one half longer than the 4 and subequal to the 6. Peduncle of uropods
a little longer than the telsonic segment, with 3 conspicuous teeth on the
medial margin; endopod a little longer than the peduncle, a little shorter
than the spine at its apex; the exopod more than three-fourths as long as
the endopod, with a long apical spine. L. Q 1:5 mm.
Gulf of Siam, depth 2 m.
3. S. bifrons Calm, 1911 S.b.. Calman in: Tr. zool. Soe. London, v.181v p.341,
362 t. 34 f. 18—21.
Pseudorostral lobes short, obtuse, with irregular denticles on the lower
edge, the front margin thence receding to the base of a stout cylindrical
process which supplies the antero-lateral angle, being itself tipped with a
curved spine and fringed with denticles on its surface. The respiratory
orifices and branchial siphons as in the preceding species. Carapace broad,
with scattered spinules or tubercles and short hairs. First pedigerous seg-
ment almost concealed, the rest broad, depressed, tuberculate. Pleon short,
the telsonic segment longer than the 5, apically acute. yes dark, each
Cumacea: 24. Nannastacidae, 1. Schizotrema, 2. Paranannastacus 167
with 3 lenses, placed just behind the branchial orifices. Antenna 1, with
2 spines at the end of 1%* joint. Hinder peraeopods rather long and slender;
in the 5 pair the 5” joint nearly thrice as long as the 4 and a little
longer than the 6%, Peduncle of uropods about half as long as the telsonic
segment, irregularly denticulate on both edges; endopod about two and a
half times as long as the peduncle, with stout apical spine about half its
length; exopod a little less than half as long as endopod, but with apical
spine reaching beyond it. L. Q 165 mm.
Paumben, India, depth 2—9 m.
4. S. sordidum Calm. 1911 S.s., Calman in: Tr. zool. Soc. London, v. 181v
p. 341, 363 t.34 f. 22—24.
Pseudorostral lobes short, truncate, obliquely upturned, the front margin
with concave recession to the sharply produced antero-lateral angle which
bears a transparent apical spine. Orifices and siphons as in the other species.
Carapace rather broad and depressed, with scattered long hairs. Pedigerous
segments broad, depressed, with scattered and a few pellucid spines. Telsonic
segment a little shorter than the 5%, round-ended. Eyes dark, in the usual
position. Antenna 1, 34 joint slightly longer than 2"4, flagellum 3-jointed,
the last joint minute. Peraeopods 3—5 very long and slender; in the 5%
pair the 5‘ joint nearly 4 times as long as the 4‘ and one-third longer
than the 6th, Pedunele of uropods not more than half as long as the telsonic
segment; the endopod more than thrice as long as the peduncle; exopod
about half as long as the endopod. L. o 1 mm.
Gulf of Siam, depth 2 m.
2. Gen. Paranannastacus T. Stebbing
1912 Paranannastacus, T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 164.
Like Nannastacus (p. 168), but with no exopods on either the third
maxillipeds or the first and second peraeopods of the female.
2 species.
Synopsis of species:
Antero-lateral angles of carapace not produced beyond the
| MSCUGOTOSHFUM re hogs Phe) 6 S12, ese yes, sup 8 a 1. P. reptans
| Antero-lateral angles of carapace produced beyond the
PSCudOrosirmmMy ce seh crete pe os) woes ss sw! 2. P. tardus
1. P. reptans (Calm.) 1911 Nannastacus r., Calman in: Tr. zool. Soc. London,
v. 181v p.341, 356 t. 33 f.22—28 | 1912 Paranannastacus r., T. Stebbing in: Ann, S. Afr.
Mus., v. 10v p. 164.
Pseudorostral lobes shortly produced, somewhat upturned, truncate,
meeting in the middle line below the branchial orifice, antero-lateral margin
deeply concave, antero-lateral corner acutely produced. Carapace little broader
than deep, depressed in middle line between the inflated branchial regions
and with a median convexity anteriorly. Pedigerous segment 1 reduced to
the side-plates, 5 pedigerous and 18* pleon segment each with a pair of
small spiniform dorsal teeth. The pair of eyes prominent and darkly
pigmented. Antenna 1, 2™4 and 34 joints subequal, and together subequal
to 15* joint, principal flagellum with minute terminal joint. Maxilliped 3 and
peraeopods 1, 2, without exopods. In maxilliped 3 and peraeopod 1 the 2™¢
joint is very short; not short but not inflated in peraeopod 2. Peduncle of
168 Cumacea: 24. Nannastacidae, 2. Paranannastacus, 3. Nannastacus
uropods about half, exopod less than a fifth, as long as the endopod: terminal
spine of exopod much longer than that of endopod. L. © 15mm. GO unknown.
Gulf of Siam, depth 2 m.
2. P. tardus (Calm.) 1911 Nannastacus t., Calman in: Tr. zool. Soc. London,
v. 181v p.341, 359 t. 34 f.4—11 | 1912 Paranannastacus t., T. Stebbing in: Ann. S. Afr. Mus.;
v.10v p. 164.
Pseudorostral lobes shortly produced, slightly upturned, truncate, meeting
in the middle line below, and apparently also above, the branchial orifice,
antero-lateral margin deeply concave, antero-lateral corner strongly produced,
serrate below. Carapace considerably broader than deep, with branchial regions
little inflated. Pedigerous segment 1 reduced to laterally expanded side-
plates with marginal laminar spines; the next segment with small median
dorsal tooth. Pleon segment 1 with pair of small dorso-lateral teeth, often
(perhaps accidentally) absent. The pair of eyes as figured rather small.
Antenna 1, 2™4 and 34 joints subequal, and combined in length subequal to
the stouter 18¢ joint. Maxilliped % and peraeopods 1, 2, without exopods,
with 24 joint not elongate, but in the maxillipeds having a long narrow distal
process, and in peraeopod 1 having a prominent rounded lobe on the medial
margin. Uropods differing from those of P. reptans (p. 167) chiefly by the
terminal spine of the exopod, which is shorter instead of much longer than that
of the endopod. L. 9 16mm. G not known.
Gulf of Siam; depths between 9 and 28 m.
3. Gen. Nannastacus Bate
1865 Nannastacus (Sp. typ.: Cuma unguiculata), Bate in: Ann. nat. Hist., ser. 3
v.15 p.86 | 1878 & 1879 N., G.O. Sars in: Arch. Naturv. Kristian., v.3 p.566; v.4 p.108 |
1880 N., Kossmann, Reise Roth. Meer., Malac. pars2 p.90 | 1887 N., G. O. Sars in:
Rep. Voy. Challenger, v.19 pars55 p.62 | 1893 N., T. Stebbing in: Int. sci. Ser.,
v.74 p.305 | 1900 N., G. O. Sars, Crust. Norway, v.3 p.79 | 1900 N., T. Stebbing in:
Willey, Zool. Results, pars 5 p.611 | 1904 N., Calman in:, Herdman, Rep. Ceylon Pearh
Fish., v.2 p.176 | 1905 N., Calman in: Sci. Invest. Fish. Ireland, 1904 app. 1 p. 23 |
1905 V., Calman in: Siboga-Exp., nr. 36 p.10 | 1911 N., Calman in: Tr. zool. Soe. London,
v. 181v p.350 | 1912 N., T. Stebbing in: Ann. 8. Afr. Mus., v. 10v p. 164.
Antero-lateral angles of carapace well defined. Eyes, when present,
divided into a distinct pair. Maxilla 1 with 2 filaments to the palp. Third
maxilliped with exopod in the male, but not always in the female. First
two peraeopods with exopods in the female. Uropods with endopod longer,
generally much longer, than the exopod.
16 species.
Synopsis of species:
Third maxilliped without exopod in the female — 2.
! Third maxilliped with exopod in both sexes — 6
9 J Antero-lateral angle of carapace obtuse — 3.
\ Antero-lateral angle of carapace acute — 4.
3 { Antero-lateral angle of carapace armed with a spine . 1. N. zimmeri
\ Antero-lateral angle of carapace without spine . . . . 2. N. suhmii
Uropods with endopod nearly thrice as long as pedunele 3. N. gibbosus
4 Uropods with endopod not nearly thrice as long as
peduncle — 5.
. { Carapace little broader than deep. ......... 4. IN. minor
“ | Carapace considerably broader than deep. ...... 5. N. agnatus
Cumacea: 24. Nannastacidae, 3. Nannastacus 169
Uropods with peduncle at least twice as long as telsonic
: segment — 7.
6 Uropods with peduncle not nearly twice as long as telsonic
segment — 9.
f Pseudorostrum short .... . BBW) No. Polini tenon Oa Nes LOD CULUS
\ Pseudorostrum long — 8.
Pseudorostral lobes strongly produced with divergent
PY DIIGL YS ig Guretee mc cnh a) IomiCa mre aac oe as et WON, Longirostris
Pseudorostral lobes less strongly produced, the truncate
apices not divergent. ..... Lh eos se 6 a es! SAIN, brachydactylus
First five segments of pleon with paired dorsal carinae — 10.
\ Pleon segments without paired dorsal carinae — 1l,
10 \ Pseudorostrum upturned, ridged. ......... . 9 N, ossiani
Pseudorostrum horizontal, smooth ........-.-. 10. N. hanseni
Carapace smooth — 12.
11 \ Carapace rough with hairs, granules, or spines — 18.
Fifth peraeopod with fifth joint little longer than sixth 11. N. georgi
P P J g
12) Fifth peraeopod with fifth joint much longer than sixth 12. N. pardus
P J g
13 Eyes not very wide apart .........-.. -.13. N. hirsutus
Kyes wide apart — 14.
Carapace roughened with granules, without spines. . . 14. N. brevicaudatus
1a Pp g g ’ Pp
\ Carapace with spines — 15.
Antero-lateral angles of carapace serrate; uropods with :
15 | exopod about one-third as long as endopod. . . . 15. N. unguiculatus
| Antero-lateral angles of carapace not serrate; uropod
with exopod nearly half as long as endopod . . - 16. N. stebbingi
1. N. zimmeri Calm. 1911 NW. z., Calman in: Tr. zool. Soc. London, v. 181v
p. 841, 352 t.33 £.4—15.
Pseudorostral lobes short, obliquely upturned, narrowly truncate,
meeting below but not above the branchial channel; antero-lateral margin
deeply concave, antero-lateral angle produced into a blunt, apically setose
spine. Carapace inflated behind, much narrowed in front, dorsal line convex,
a slight elevation at the rear bearing a small tubercle. Pedigerous segment
1 a narrow strip with expanded side-plates. Pledn without teeth or spines.
Maxilliped 3 without exopod, but with blunt distal process on 2Ӣ joint.
Peraeopods 1, 2, with exopods. Peduncle of uropods nearly one-third as
long as the endopod, which is over eight times the length of the exopod,
but has a shorter apical spine. A dark patch of pigment on each side of
the carapace anteriorly, and a band encircling 5 segment of pleon. L. 15 mm.
3. Pseudorostral lobes short, blunt, meeting a short distance above
the branchial channel; antero-lateral margin deeply concave, angle rounded
and followed by a curved spine and some serration. Carapace with surface
minutely granular, not much narrowed in front, dorsal line nearly straight,
with depression between the inflated branchial regions. Pedigerous segment
1. scarcely visible, and its side-plates very small. Pleon with lateral groove
overhung by a serrated crest in each segment. Hyes dark, larger than in ©,
and less wide apart. Antenna 1 has 2™4 joint produced distally into a narrow
process tipped with plumose setae. Maxilliped 3 with well developed exopod,
2nd joint relatively longer than in ©. Peraeopod 5 with 5 and 7‘ joints
long. Uropods with endopod about ten times as long as exopod. Colour
as in Q and sometimes also bands on 3¢ and 4" pleon segments. L. 16 mm.
Trincomali, Ceylon, surface.
170 Cumacea: 24. Nannastacidae, 3. Nannastacus
2. N. suhmii O. Sars 1887 N. s., G.O. Sars in: Rep. Voy. Challenger, v.19
pars 55 p. 9,63 t.10 £.4,5 (M. swhmi in: explanation of plate) | 1900 N. s., T. Stebbing
in: Willey, Zool. Results, pars 5 p.64 | 1904 N. s., Calman in: Herdman, Rep. Ceylon
Pearl Fish., v.2 p.177 | 1905 NV. s., Calman in: Siboga-Exp., nr. 36 p.12 | 1911 N.s.,
Calman in: Tr. zool. Soc. London, v.181v p.354 | 1912 N. s., T. Stebbing in: Ann.
S. Afr. Mus., v. lOV p. 164.
©. Form of body rather stout. Pseudorostral lobes obliquely upturned,
forming a narrow blunt-ended tube tipped with setules, antero-lateral margins
broadly concave, the lower corners strongly produced, ending bluntly, serrate
below. Carapace setulose, without keels or spines (Sars), but perhaps with
medio-dorsal tooth or spine (Calman), boldly arched till near the end which
carries a few small denticles. First 4 pedigerous segments very short, side-
plates evenly rounded, quite smooth. Pleon with a few very small denticles
on dorsal surface of each segment (G. O. Sars), anterior pleon segments with
strong paired teeth on dorsal surface (Calman). Eyes wide apart, each with
8 corneal lenses. Antenna | slender, accessory flagellum knob-shaped. Maxilli-
ped 3 and peraeopods 1, 2, with remarkably small exopods (G. O. Sars),
maxilliped 3 with no trace of an exopod (Calman). Peraeopods 3—5 subequal,
considerably longer than peraeopod 2, their 5 and 6 joints elongate.
Peduncle of uropods unusually short, exopod exceedingly small, though con-
sisting of two well defined joints, endopod rather elongate. Colour rather
pellucid, with a few dark brown patches, one low down on each side of
carapace, and a band on 5 pleon segment. L. about 2—2°6 mm.
GS. Less robust than Q. Pseudorostral lobes little prominent, obtusely
rounded, antero-lateral corners less prominent, lower margin irregularly flexuous,
smooth. Carapace with dorsal line almost horizontal. Pleon segments quite
smooth. Eyes much larger than in ©. Antenna 2 scarcely reaching beyond
34 pleon segment, joints of flagellum very long and thin. Peraeopod 1 with
5th and 6 joints slender, subequal, the 7 shorter. Endopod of uropods
even more elongate than in the Q. Colour much darker and more extended.
In both sexes a dark band on 4* joint of maxilliped 3 and peraeopod 1.
Off Samboanga, Philippine Islands; surface.
3. N. gibbosus Calm. 1911 N.g., Calman in: Tr. zool. Soe. London, v.181v
p. 341, 355 t. 33 f. 16—21.
Pseudorostral lobes short, truncate, meeting in middle line below branchial
orifice. antero-lateral margin deeply concave, lower corner produced, triangular,
sometimes with one or two denticles at apex. Carapace variable, uneven,
setose. with depression between branchial bosses, a small boss behind each
eye. a denticle sometimes behind the eyes and one or two on rearward dorsal
elevation. Pedigerous segment 1 represented only by side-plates, those of
the following segments expanded and rounded, segments 2 and 5 sometimes
with medio-dorsal denticle, such as also occurs sometimes on pleon segment 1.
Eyes prominent, dark. Antenna 1, 34 joint longer than 2™4, flagellum with
minute terminal joint. Maxilliped 3 without exopod, 2™¢ joint rather short,
apical process not long. Peraeopods 1, 2, with well developed exopods,
2nd joint of peraeopod 2 dilated, 5‘ joint longer than 4*, 7% twice as long
as 6: in 5 pair 5t joint fully twice 4t* and subequal to 7 joint. Endo-
pod of uropods nearly thrice length of peduncle and fully 8 times as long
as exopod, but apical spine of exopod considerably longer than that of
endopod. L. Q 18mm. G unknown.
sulf of Siam, 4—9 m.
Cumacea: 24, Nannastacidae, 3. Nannastacus 171
4. N. minor Calm. 1911 N. m., Calman in: Tr. zool, Soc. London, v. 181v
p. 341, 357 t.34 f.1—3 | 1912 N.m., T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 164.
Like Paranannastacus reptans (p. 167), but dorsal surface of carapace
less arched, and branchial regions less inflated, antenna 1 with 3° segment
shorter than 2™4, peraeopods 1, 2, with well developed exopods, 204 pair with
2nd joint inflated. L. Q@ 13mm. G unknown.
Gulf of Siam, depth 2 m.
5. N. agnatus Calm. 1911 N.a., Calman in: Tr. zool. Soe. London, v. 181v
p. 341, 360 t. 34 f.12,13 | 1912 N. a., T. Stebbing in: Ann. S. Afr. Mus., v. 10v_ p. 164.
Like Paranannastacus tardus (p. 168), but telsonic segment shorter,
peraeopods 1, 2, with well developed exopods, 2nd joint of 1S* pair a little
shorter, 24 joint of 2"4 pair a little longer, hinder pairs slightly stouter and
shorter. L. Q 13mm. GC unknown.
Gulf of Siam, depths between 9 and 28 m.
6. N. lepturus Calm. 1911 N.1., Calman in: Tr.
zool. Soc. London, v. 18iv p. 341, 352 t. 33 f. 1—3.
Pseudorostral lobes short, truncate, antero-
lateral margin shallowly concave, lower corner
rounded. Carapace slightly compressed, dorsal line
a little convex. Pedigerous segment 1 distinct,
side-plates of the following segments little expanded
laterally. Pleon segments each with well marked
lateral groove, 5t twice as long as the truncate
telsonic segment. Antenna 1, 2" joint with sub-
distal process not reaching beyond the apex, each
flagellum 2-jointed, accessory a little shorter than
1st joint of principal. Peduncle of uropods about
twice as long as the endopod, which has 7 spines
on the medial margin; exopod about four-fifths
as long as endopod, but with a much longer apical
spine. L. Od 2mm. oO unknown.
Suez.
7. N. longirostris O.Sars 1879 N.1, G. O. Sars
in: Arch. Naturv. Kristian., v.4 p.119 t. 58, 59 | 1905 N.
1, Calman in: Siboga-Exp., nr. 36 p.15 | 1911 N. 1,
Calman in: Tr. zool. Soe, London, v. 181v p. 352.
A slender form, integument thin, setulose
especially in ©. Pseudorostral lobes strongly pro-
duced, horizontal, with divergent apices which are
serrate and setose, antero-lateral margin concave,
lower corner little produced, acute and bidentate
in Q, rounded in G. Carapace depressed along
the middle, in profile nearly horizontal, till it bends
down to the long pseudorostral lobes. Pedigerous
segment 1 short but distinct. Eyes wide apart,
; , Fig. 116.
each with 3 corneal lenses, larger in CO than N. longirostris (3) Sars.
in Q. Antenna 1, 34 joint longer than 274,
nearly as long as 1%, accessory flagellum 2-jointed. Maxilliped 1, exopods
forming two clearly separated very narrow tubes, apically truncate, far-
172 Cumacea: 24. Nannastacidae, 3. Nannastacus
projected, the ends usually a little divergent. Peraeopod 1 with 4—6 laminar
spines on lateral margin of 24 joint, 7 joint more than half as long as
6th, with long thin spine-like claw. Peduncle of uropods long and narrow,
especially in the d, more than twice as long as the exopod which is from
four-fifths to three-fourths as long as the endopod and has a longer apical
spine; endopod with 4 spines on medial margin in 9, 7 in CG. Colour greyish
white, with some darker patches in front. L. 3 mm.
Mediterranean, at Spezia; depths 11—19 m.
8. N. brachydactylus Calm. 1905 WN. b., Calman in: Siboga-Exp., nr. 36. p. 2.
14 tf. 3a—e.
Allied to .V. dongirostris, but integument not setulose, pseudorostral
lobes less elongate, not apically divergent, antero-lateral margin very short,
lower corner quadrate; telsonic segment broader than long; eyes very large,
dark, near together; peraeopod 1 very stout, 7 joint less than one-third
of the length of the 6%, with very short stout curved claw; peduncle of
_uropods a little more than one and a half times the length of the endopod,
which has a short apical spine and 7 spines on the medial margin: exopod
incomplete, apparently agreeing with that of NM. dongirostris. L. CG 1-6 mm.
© unknown.
Sailus Ketjil, Paternoster Islands (Sunda-Sea?); surface.
9. N. ossiani Stebb. 1900 N.o0., T. Stebbing in: Willey, Zool. Results,
pars 5 p. 606, 612 t.64A | 1904 N. o., Calman in: Herdman, Rep, Ceylon Pearl Fish.,
v.2 p.177 | 1905 N. 0., Calman in: Siboga-Exp., nr. 36 p. 12.
Pseudorostral lobes obliquely upturned, apically rounded, the distal part
concentrically ridged, bases of 3 spines below the upturned lobes (shown in
figure); antero-lateral corners blunt, little produced. Carapace tumid. median
line in profile with long anterior and short hinder convexity, branchial regions
inflated. Fourth and fifth pedigerous segments having the centre dorsally
much elevated. Pleon segments with pellucid lateral carinae and pair of
serrate dorsal carinae, distally extended into a strong tooth in segments 1—5,
segment 5 not greatly longer than any other, telsonic segment apically pointed.
Eyes wide apart, dark, apparently composed each of 3 corneal lenses. Flagellum
of antenna 2 comparatively short, 10-jointed. In peraeopod 1, 5% joint sub-
equal to 6, but considerably longer than 6 in 34, 4%, and 5% pairs.
Pedunele of uropods a little shorter than the slender exopod, which is no
doubt shorter than the much stouter (broken) endopod. L.d 2mm. 90
unknown. .
Blanche Bay, New Britain; surface.
10. N. hanseni Calm. 1905 N. h., Calman in: Siboga-Exp., nr. 36 p. 11 tf. la—e.
Pseudorostral lobes horizontal, apically truncate, not ridged, not meeting
either above or below the branchial orifice; antero-lateral corners blunt,
obscurely serrate, slightly produced. Carapace depressed in median line
between the inflated branchial regions, elevated behind into bilobed prominence
overhanging some of the pedigerous segments, of which the last 3 have each
a dorsal pair of spinose prominences. Pleon segments 1—4 have lofty spine-
tipped pairs of prominences, the 5 segment a much lower pair, telsonic
segment apically pointed. Eyes large, dark, each with 3 corneal lenses. Limbs
as in JN, ossiani (nr. 9), and probably also the uropods, the unbroken endopod
co
Cumacea: 24. Nannastacidae, 3. Nannastacus 1
twice as long as the exopod, with a shorter apical spine. Integument beset
with tubercles and granules. L. dG 15 mm. Q unknown.
Paternoster Islands, and between these and Postillion Islands; surface.
11. N. georgi Stebb. 1900 WN. g., T. Stebbing in: Willey, Zool. Results,
pars 5 p. 606,615 t.64B | 1905 N. g., Calman in: Siboga-Exp., nr. 36 p. 12.
Near to NV. sudmit 3 (nr. 2) and N. zimmeri GD (nr. 1). Pseudorostral
lobes subhorizontal, apically rounded, not carinate, antero-lateral margin sinuous,
lower corner acute. Carapace with branchial regions inflated flanking a depression
between long and short convexities of the median line. 1%* pedigerous seg-
ment almost entirely concealed; fifth pedigerous and 1%* pleon segment with
a marked longitudinal medio-dorsal depression. Pleon segments with pellucid
lateral carinae, 5 segment considerably the longest, telsonic segment distally
triangular. Eyes well separated, large, dark, each with 3 corneal lenses. Antenna
2. flagellum of 18 joints or more, distally extremely slender. Maxilliped 1,
branchial leaflets about 16. Peraeopod 5, 5* joint not greatly longer than 6".
Pedunele of uropods more than twice as long as the very short exopod, but
scarcely a third as long as the endopod; apical spine of exopod considerably
longer than that of endopod. L. 3d 25 mm. © unknown.
Blanche Bay, New britain; surface.
12. N. pardus Calm. 1905 N. p., Calman in: Siboga-Exp., nr. 36 p.2, 13 tf.2a—e.
Pseudorostral lobes very short, upturned, apically rounded, not meeting
above, antero-lateral margin forming two convexities, lower corner rounded
with a single curved tooth on its surface. Carapace smooth, median
line forming long and short low convexities, separated by slight depression
between the branchial regions. The 5 pedigerous segments unarmed, dorsally
rounded, with expanded side-plates. Pleon stout, setulose, first 5 segments
with well marked lateral groove, telsonic segment little shorter than 5%,
extended to a blunt apex. Eyes large, dark, each with 3 corneal lenses. Peraeo-
pod 5 long and slender, its 6 joint only two-thirds as long as 5. Peduncle
of uropods about one-third, exopod about one-fourth, as long as the endopod;
apical spine of exopod much longer than that of endopod. Colour characteristic,
large black spots in great numbers, especially dark on the carapace. L. d
17mm. Q unknown.
Paternoster Islands; surface.
13. N. hirsutus H. J. Hansen 1895 N.h., H. J. Hansen in: Ergeb. Plankton-
Exp., v2 G.c. p.59 t. 7 £. 38, 3a, b.
Whole body free from spines or processes but beset with scattered
tolerably long hairs. Pseudorostral lobes somewhat upturned, meeting in
front, obliqely truncate, antero-lateral margin shallowly concave, lower corner
not at all produced, obtusely quadrate and armed with a stout plumose seta.
Carapace plump, medio-dorsal line evenly convex. All five pedigerous seg-
ments distinct. Telsonic segment not much shorter than 5, blunt at apex.
Eyes not far apart. Antenna 1, 2¢ joint rather longer than 34, flagellum
of 2 equal joints, accessory rudimentary. Peduncle of uropods longer than
the exopod, endopod defective. Colour brownish grey. L. ovigerous © 1°7 mm,
S unknown.
Bermudas.
174 Cumacea: 24. Nannastacidae, 3. Nannastacus
\
14. N. brevicaudatus Calm. 1905 N. b., Calman in: Sci. Invest. Fish. Ireland,
1904 app. 1 p.11, 26 t.1 f. 15—19.
o. Near to N. unguiculatus’ (nr. 15), but pseudorostral lobes very
short, not rising above eyes in lateral view, antero-lateral corner less pro-
duced, rounded at apex, finely serrate below. Surface of carapace rough
with prominent granules but with no flattened spines. Pedigerous segments
with a few small spines near lateral margins. Pleon shorter and stouter,
telsonic segment with no median spine on hind margin. Eyes colourless
or with colour easily disappearing. Antenna 1, process on 2™4 joint shorter.
In peraeopod 3 the 2°42 joint equals in length the remaining joints combined,
and in the 4t and 5‘ pairs is nearly two-thirds of their length. Peduncle
of uropods little longer than its distal breadth, endopod once and a half as
long as the peduncle and exopod about two-fifths as long as the endopod.
L. 18 mm.
SG. Near to N. unguiculatus, but scattered setae on surface more
numerous, pleon much shorter, serrations on its dorsal surface less distinct;
eye pigment much less dark, reddish brown; hinder peraeopods with 2¢ joint
relatively longer, that of 4t pair nearly equal to remaining joints combined;
peduncle of uropods about two-thirds as broad as long, a little over half as
long as the endopod, which is about thrice as long as the exopod. Body
as preserved colourless. L. 1°95 mm.
_ Ballynakill Harbour.
15. N. unguiculatus (Bate) 1859 Cuma wnguiculata, Bate in: Ann. nat. Hist.,
ser.3 v.83 p.273 tf.Q, i, m, u | 1865 Nannastacus binoculoides, Bate in: Ann. nat. Hist.,
ser.3 v.15 p.87 t.1 f.4(4) | 1869 Nannasticus b., A. M. Norman in: Rep. Brit.
Ass., v.38 p.259 | 1879 Nannastacus wnguiculatus, G. O. Sars in: Arck. Naturv.
Kristian., v4 p. 109 t.55—57 | 1889 Nannasticus wu. (part.), D. Robertson in: P. nat.
Hist. Soc. Glasgow, ser.2 v.31 Tr. p.49 | 1893 Nannastacus u., T. Stebbing in: Int.
sci. Ser., v.74 p.305 | 1901 N. unguiculata, T. Scott in: Rep. Fish. Board Scotl., v.19
p. 276 | 1902 N. unguiculatus, T. Scott in: Rep. Fish. Board Scotl., v.20 p. 480 | 1904
N. u., Calman in: Herdman, Rep. Ceylon Pearl Fish., v.2 p.176 | 1905 N. w., Calman
in: Siboga-Exp., nr.36 p.12,13 | 1905 N.u., Calman in: Sci. Invest. Fish. Treland, 1904
app. 1 p.25 | 1906 N.u., A. M. Norman & T. Scott, Crust. Devon Cornwall, p.33.
Q. Body short and plump. Integument hard, rough with little tubercles
and scattered hairs. Pseudorostral lobes not very long, upturned nearly
perpendicularly, with serrate edges, meeting below the branchial orifice:
antero-lateral margin deeply concave, the lower corner being strongly and
sharply outdrawn, the margin below coarsely serrate for a long distance
backward. The large carapace obliquely truncate behind, its medio-dorsal
line convex till near the rear, where a short depression is followed by another
rise, which is spinose; the former convexity carries 2 longitudinal rows of
spines, which are small compared with the curved rows of flattened spines
which flank the branchial regions to the rear. One or two similar spines
are found on the margins of the outstanding side-plates of each pedigerous
segment. These segments have also dorsally 2 rows of smaller spines. Such
spines are continued on the pleon, each row having 3 on the 5‘ segment,
2 on the 4, 1 spine only on each of the other segments. The telsonic
segment has a single spine on its obtuse apex. Eyes wide apart, very dark,
each with 3 corneal lenses. Antenna 1, 2"¢ joint apically produced, flagellum
3-jointed, accessory 1-jointed, rudimentary. Antenna 2 1-jointed, with 3 setae.
Exopods of maxillipeds 3 and peraeopod 1, 2, slender. Peraeopods 3—9
Cumacea: 24, Nannastacidae, 3. Nannastacus FAs
all longer than peraeopod 2, and successively longer; 2nd joint in peraeo-
pod 3 nearly as long as the other joints combined, but in 4 and 5 much
shorter, in those pairs the 5 joint longer than the 6". Peduncle of uro-
pods less than half as long as endopod, which is about thrice as long as
the exopod, but with much shorter apical spine. L. about 2 mm.
Fig. 117.
N. unguiculatus (2) Bate, front (after
Sars).
Fig. 119.
N. unguiculatus (2) Bate, maxilli-
ped 2, vibratory lamina (after Sars).
Fig. 118. Fig. 120.
N. unguiculatus (2) Bate, maxilliped 1 N. unguiculatus Bate, maxilliped 3
(after Sars). (after Sars).
GS. Body slightly more slender, without scattered hairs. Pseudorostral
lobes shorter, thicker, less strongly upturned, antero-lateral corners quadrate.
Carapace almost vertically truncate behind, devoid of the flattened spines
flanking the branchial regions. Eyes larger. Antenna 2, flagellum filiform,
of about 18 elongate joints. Maxilliped 1, branchial leaflets (wanting in Q )
about 16 in number. Exopods of maxilliped 3 and peraeopods more strongly
176 Cumacea: 24. Nannastacidae, 3. Nannastacus, 4. Platycuma
developed. In peraeopod 4 the development of the exopod-bearing 2™¢ joint
is strongly contrasted with the slenderness of the following joints. Uropods
rather longer than in the Q. .
Seas of Great Britain and Ireland, from north of Shetland to Plymouth; W. coast
of France: Mediterranean, Messina and Spezia, depths 11—19 m.
16. N. stebbingi Calm. 1904 N. s., Calman in: Herdman, Rep. Ceylon Pearl
Fish., v.2 p. 159,176 t.5 f. 91—93.
Near to N. osstani (nr. 9), but with the last three pedigerous segments
carrying paired serrated crests on the dorsal surface. Here also immediately
below the pseudorostrum the concave antero-lateral margin bears a group of
three curved spines, of which the upper and largest is very conspicuous. These,
however, are perhaps also present in XN. ossiani. L. G, 1°38 mm, Q unknown.
Gulf of Manaar; depth 4 m.
Genus dubium et species dubiae
Diops Paulson 1875 D., Paulson, Unters. Krebse rot. Meer., v.1 p. 129 | 1879
Nannastacus, G. O. Sars in: Arch. Natury. Kristian., v.4 p.108 | 1880 N., Kossmann,
Reise Roth. Meer., Malac. pars 2 p. 90.
Probably a synonym of Nannastacus (p. 168), but (as described) distinguished by
a slender 3-jointed second antenna in the female, and by having only a single apical
filament on the palp of the first maxilla.
Diops parvulus Paulson 1875 D.p., Paulson, Unters. Krebse rot. Meer., v. 1
p. 129 t.21 f,1—I1r | 1880 D. (Nannastacus) p., Kossmann, Reise Roth. Meer., Malac.
pars2 p.90 | 1893 D. p., T. Stebbing in: Int. sci. Ser., v.74 p. 306.
Q. L. 35 mm, larger than any accepted species of Nannastacus, and impossible
to reconcile with N. wnguiculatus, the apical lobes of the pseudorostrum not being
serrate, the antero-lateral corners little instead of much outdrawn and the margin below
little if at all serrate. The telsonic segment (as figured) differs from the accustomed form.
Nannastacus spinosus (Paulson) 1875 Diops s., Paulson, Unters. Krebse rot.
Meer., v.1 p.181 t.21 f.2 | 1880 ?.Nannastacus sarsii, Kossmann, Reise Roth. Meer.,
Malac. pars2 p.90 t.4 f.4—9, t.5 f. 1.
Kossmann’s species, 3, 2°75 mm long, has 2 filaments on the palp of the first
maxilla, Paulson’s specimen, only 1 mm long, was evidently not dissected. Both authors
represent the 5th segment of the spinose pleon as uncommonly short.
4. Gen. Platyeuma Calman
1905 Platycuma (Sp. un.: P. holti), Calman in: Sci. Invest. Fish. Ireland, 1904
app. 1 p.3,30| 1912 P., T. Stebbing in: Ann. 8. Afr. Mus., v.10v p. 164,171.
Carapace depressed and expanded, overhanging the anterior pedigerous
segments. Pedigerous segments except the first laterally expanded. Telsonic
segment short. Eye wanting. Alimentary canal spirally coiled. Mouth-organs
generally as in Cumellopsis (p.177), but first maxilla with single filament to
the palp, epipod of first maxilliped with a single branchial leaflet.
Female unknown.
1 species.
1. P. holti Calm. 1905 P. h., Calman in: Sci. Invest. Fish. Ireland, 1904 app.1
p. 10,30 | 1912 P.h., T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 131.
Integument pellucid. Pseudorostral lobes slightly upturned meeting
before the frontal lobe for one-eighth the length of the carapace, not as far
Cumacea: 24, Nannastacidae, 4. Platyeuma, 5. Cumellopsis 177
forward as the antero-lateral corners, truncate in dorsal view, antero-lateral
margin concave between 2 convexities. Carapace large, depressed, quadrate
in front, slightly widened to the convex hinder part of the carinate sides,
hind margin trilobed, the middle lobe concealing the first 2 and partially
the 34 of the pedigerous segments. Of these the
4th and 5 and of the pleon the first 5 segments
have each a pair of thin serrate dorsal crests. Pleon
less than half the length of the body, the segments
without lateral grooves; telsonic segment distally
wider than its length, scarcely produced between
the uropods. Eyelobe obsolete. Antenna 1, 1°"
joint equal to 2™¢ and 34¢ combined, flagellum
3-jointed, shorter than 3¢ joint of peduncle, 2-
jointed accessory minute. Antenna 2, with sensory
hairs very numerous, flagellum not reaching the
pleon. Mandibles with about 9 spines in the spine-
row, molar stout. Maxilliped 1, the 3¢ joint sup-
pressed. Maxillipeds 2, 3, 34 joint distinct, but
small. Peraeopod 1, 6 joint subequal to 5%,
nearly thrice as long as 7. Peraeopod 2, 7
Fig. 121. Fig. 122.
P. holti (gd) Calman. P, holti Calman, coiled intestine.
joint longer than 5‘ and over thrice as long as 6‘". In the 34 to 5% pairs
the 5th joint more than twice as long as the 6, the 7" very small but
carrying a long curved spine or claw. Uropods three-fourths as long as the
pleon, endopod about two-thirds as long as the peduncle, and exopod in the
same relation to the endopod. L. 4:1 mm.
W. coast of Ireland, 77 miles W. N. W. of Achill Head, county Mayo, depth 699 m,
5. Gen. Cumellopsis Calman
1905 Cumellopsis (Sp. un.: C. helgae), Calman in: Sci. Invest. Fish. Ireland, 1904
app. 1 p.3, 28 | 1906 C., Calman in: Mt. Stat. Neapel, v.171v p. 416.
Carapace subovoid, more or less overhanging anterior pedigerous
segments, antero-lateral corners well pronounced. Eyelobe small, without
eye. First maxilla with two filaments to the palp. Branchial leaflets fairly
developed.
2 species.
Synopsis of species:
J Peduncle of uropods nearly twice as long as the endopod 1. C. helgae
\ Peduncle of uropods not nearly twice as long as the endopod 2. C, puritani
Das Tierreich. 39. Lief.: T. R. R. Stebbing, Cumacea. 12
178 Cumacea: 24, Nannastacidae, 5. Cumellopsis, 6. Cumella
1. CG. helgae Calm. 1905 C. h., Calman in: Sci. Invest. Fish. Ireland, 1904
app. 1 p.28 t.2 f. 20—34 | 1906 @. h., Calman in: Mt. Stat. Neapel, v.171v p. 418.
Pseudorostral lobes slightly upturned, dorsally truncate, shortly produced in
front of the small sightless eyelobe, front margin concave below with prominent
antero-lateral angle leading to 2 teeth on the lower margin. Carapace
elevated above the pedigerous segments, dorsal line slightly carinate to the
rear, each side with broad shallow longitudinal depression, evanescent to the
rear, bounded above and below by low rounded ridges, the lower sharpened
at the antero-lateral corner. Pleon a little longer than the preceding part of
the body, 5% segment about twice as long as the telsonic. Antenna 1,
1st joint shorter than 2"¢ and 34 combined, flagellum 3-jointed, the 2-jointed
accessory minute. Maxilliped 1 with 10 branchial leaflets on epipod of
young SG. Peraeopod 1, 6 joint a little shorter than 5, nearly twice as
long as 7%, Peraeopod 2, 7% joint little more than twice as long as 6".
In 34 to 5th pairs the 5% joint rather long. Uropods considerably longer
than 5 and telsonic segments of pleon combined, peduncle slender, nearly
twice as long as endopod, exopod three-fourths as long as endopod. L. ©
with developing oostegites, 5°8 mm, young d 47 mm.
77 miles W. N. W. of Achill Head, county Mayo; depth 699 m.
2. C. puritani Calm. 1906 C.p., Calman in: Mt. Stat. Neapel, v.17iv p. 418
t.27 £.10—12.
Very near to Cumellopsis helgae, but the size much less, antero-lateral
corner less acute, lateral longitudinal depression of the carapace bounded
below by a sharply marked ridge, the uropods with the peduncle shorter
and rami longer, the endopod in immature male eight-ninths, and in adult
male nearly three-fourths as long as the peduncle. L., adult d, 3°6 mm,
immature CO, 2°7 mm.
Mediterranean, near Capri, depth 950—1100 m.
6. Gen. Cumella O. Sars
1865 Cumella (Sp. un.: C. pygmaea), G. O. Sars in: Forh. Selsk. Christian., 1864
p.198 | 1869 C., A.M. Norman in: Rep. Brit. Ass., v. 38 p. 272 | 1879 Spencebatea.
(Sp. un.: S. abyssicola), A. M. Norman in: Ann. nat. Hist., ser. 5 v.3 p. 67 | 1879 Cumella,
G. O. Sars in: Arch. Naturv. Kristian., v.4 p.93, 126 ; 1900 C., G, O. Sars, Crust. Norway,
v.3 p.80 | 1905 C., Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p.23 | 1911 C.,
Calman in: Tr. zool. Soc. London, v.181v p.344 | 1912 C., T. Stebbing in: Ann. S.
Afr. Mus., v. 10v_ p. 129, 164, 166.
Eyes, when present, confluent, median. Peduncle of uropods usually
longer than the rami, the endopod usually longer than the exopod.
11 species accepted, 2 inadequately known.
Synopsis of accepted species:
| Pedigerous segment 3 with submedian pair of upraised
1 beethy! i; ' oi. ie: fe ne Moe fe 8 Us Tease ogre) Mey Vokes ee oe 1. C. australis
| Pedigerous segment 3 without such teeth — 2.
{ Eyes wanting; second antennae of female 3-jointed — 3.
| Eyes present; second antennae of female 2-jointed — 4.
3 Hyelobewiarrowig <= Ge fers tae ne oe as a as) eR 2. C. gracillima
\ Biyelobe: broad:.. "25 -sejeacr vee eines ae ee 3. C. molossa
Telsonie segment longer than preceding segment — 5.
Telsonic segment not longer than preceding segment — 6.
Cumacea: 24. Nannastacidae, 6. Cumella 179
Pseudorostrum long, the lobes meeting well in front of
the eyelobe. . “Meu ap pes) SOnaee teeny omc. ee
Pseudorostrum short, the lobes scarcely meeting in front
Ofstheseyelobe: sets Weta cen eel) se ay ee . . §. C. clavicauda
4. C. forfieula
Peduncle of uropods more than twice as long as the
telsonic segment ...... RIE Oe dr anes cir
| Peduncle of uropods less than twice as long as the
telsonic segment — 7.
6 6. C. leptopus
Fifth peraeopod with Sth joint very much longer
than 6th — 8,
Fifth peraeopod with 5th joint not very much longer
than 6th — 9.
U
. hispida
. serrata
{ Telsonic segment longer than peduncle of uropods 7
\ Telsonie segment shorter than peduncle of uropods . . 8.
QO
Uropods with endopod not longer than exopod. .. . 9. C. laevis
p P 8 p
Uropods with endopod longer than exopod — 10.
I g
10 J Pseudorostram short, obtuse, horizontal - ... . . .10. C. pygmaea
) Pseudorostrum rather long, acute, somewhat upturned . 11. C. limicola
1. C. australis Calm. 1907 C. a., Calman in: Nat. antarct. Exp., v.2 Crust. Cum,
p.4 tf. 7—13 | 1907 C.a., Zimmer in: Zool. Anz., v.31 p.369 | 1911 C. a., Calman in:
Tr. zool. Soe. London, v. 181v p. 344.
Pseudorostral lobes short, vertically truncate, meeting in front of the
subglobular eyelobe, the front margin forming a semicircular concavity, the
antero-lateral corners rounded, with strong serration continued a short
space along the lower margin. Carapace large, compressed, dorsal line arched,
serrate with about 20 large teeth and others, the foremost tooth overhanging
the eyelobe. Pedigerous segment 3 produced dorsally into a pair of conti-
euous teeth overhanging the following segment but with their concave side
forward. Pleon short, stout, cylindrical, with truncate end, 5" segment the
longest. Maxilliped 3, 24 joint with about 4 teeth on the medial margin, the
distal one very strong. Peraeopod 1 rather short and stout, 2"¢ joint with
5 strong teeth on lateral margin distally. Peraeopod 3 with 2¢ joint slender,
curved, longer than remaining joints combined, 5 nearly twice as long as
4th and once and a half the 6%. Peduncle of uropods not quite twice as
long as the telsonic segment, last two-thirds of medial margin with dwindling
serration; endopod about two-thirds of peduncle, and exopod two-thirds of
endopod. L. © 2°9 mm, CO unknown.
Antarctie Circle.
2. C. gracillima Calm. 1905 C. g., Calman in: Sci. Invest. Fish, Ireland,
1904 app.1 p.23 t.1 f.5—14 | 1911 C.g., Calman in: Tr. zool. Soc. London, v.181v p.344.
©. Pseudorostral lobes separated almost completely or completely by
the narrow sightless eyelobe, bent almost straight upwards with spiniform
process at the apex, obliquely truncate with front margin descending almost
straight to the antero-lateral corner from which projects the largest of a
series of some 10 pellucid spines. Branchial siphons separate, very long,
spirally coiled. Carapace rather short, compressed, deep, with medio-dorsal
carina and about 4 long spiniform teeth. Pedigerous segments 1, 2 very
short, 4t2 with a forward curving dorsal spine (perhaps paired); side-plates
not extensive. Pleon long and slender, much longer than preceding part
of the body, 5 segment more than twice as long as the telsonic. Antenna
1 very long and slender, 1st joint longer than 2™¢ and 3¢ combined, flagellum
12*
180 Cumacea: 24. Nannastacidae, 6. Cumella
of 3 joints, the 3¢ minute, accessory minute, 2-jointed. Antenna 2, 1%" joint
large, 2"4 and 34 small, each with apical seta, that on 24 joint very long.
Maxilliped 3, distal lateral angle of 2"¢ joint not produced. Peraeopod 1,
2nd joint slender, unarmed, 5% and 6* joints subequal; 34 to 5t™ pairs rather
long and slender, 5 joint about thrice as long as 6. Peduncle of uro-
pods long and slender, thrice the length of the telsonic segment, the rami
subequal, a little over half the length of the peduncle. L. subadult, with
developing oostegites, 2°75 mm.
S. Seemingly near to Cumella abyssicola (p. 184). Pseudorostral lobes
more oblique than in Q, less acute, only 4 small teeth at antero-lateral
corner. Carapace with dorsal edge smooth, without teeth. Flagellum of
antenna 2 short. Peduncle of uropods about two and a half times as long
as the telsonic segment, less than twice the length of the rami. L. 24 mm.
Off W. of Ireland, 77 miles W. N. W. of Achill Head, county Mayo, depth 699 m,
64 miles N. W.'/2 W. of Cleggan Head, depth 364 m.
3. C. molossa Zimmer 1907 C. m., Zimmer in: Zool. Anz., v.31 p.370 | 1911
C.m., Calman in: Tr. zool. Soe. London, v. 181v p. 344.
Near C. gracillima (nr. 2), but less slender; the eye-lobe, though without
pigment or lenses, large, broad and well-developed, carrying a pair of postero-
lateral spines, the pseudorostral lobes not meeting in front of it, but keeping
apart, along with the cornet-shaped siphons. The carapace not quite twice
as long as the pedigerous segments combined, the antero-lateral margins
receding, the lower margin serrulate to the end behind the larger denticulation;
traces of medio-dorsal teeth are present. Pedigerous segments 4 and 5 each
with a forward standing dorsal spine, the 5 with a spine on each side,
being in this respect resembled by the first 5 pleon segments. Pleon little
longer than preceding part of the body. Uropods broken. Colour white.
L. ¢ about 4 mm, GO unknown.
German south Pole Exp., Winter station lat. 66° 2‘ S., long. 89° 38‘ E., depth 385m.
4. C. forficula Calm. 1911 C. f., Calman in: Tr. zool. Soc. London, v. 181v
p. 341, 344, 345 t. 32 f. 1—6.
©. Pseudorostral lobes well produced, acute, meeting in front of the
prominent eyelobe; front margin receding, antero-lateral corner obtuse with
a minute tooth. Carapace compressed, with scattered hairs, dorsal line convex,
concave, and then briefly convex, front convexity with a forward curving
spine, so also the hinder but not always. Pleon a little shorter than pre-
ceding part of the body, telsonic segment the longest, inflated, arched above
and below, encreased in width backwards, obtusely extended between the
uropods. Eye well pigmented. Antenna 1 slender, 1-jointed accessory flagellum
minute. Peraeopods 3—5 slender, with elongate 5'* joint, which in the 5%
pair is four-fifths as long as the 2"¢ and more than twice the 4% joint. Uro-
pods very short, less than telsonic segment even including their indistinctly
defined apical spines; peduncle less than half as long as the telsonic seg-
ment, equal to the exopod, a little shorter than the endopod, spine in each
case included, these spines curving inwards like the tail-forceps of an earwig.
Le 1am.
GS. Rather more slender and elongate than the ©; pseudorostral lobes
shorter, with the upper edge concave; carapace with dorsal line more level
and without spines, pleon segments 1—5 with lateral grooves; eye very
- Cumacea: 24. Nannastacidae, 6. Cumella 181
large, corneal lenses large and prominent; antenna 2, flagellum reaching end
of body; uropods a little longer than the telsonic segment, the peduncle more
than half the length of that segment. L. 1°85 mm.
Gulf of Siam, depths from 2 to 19 m.
5. C. eclavicauda Calm. 1911 C.c¢., Calman in: Tr. zool. Soe. London, v. 181v
p- 341, 344, 346 t.32 f.7—10.
Pseudorostral lobes very short, seemingly not meeting in front of the
eyelobe, truncate except for a small tooth at the medial angle, front margin
short, nearly straight, antero-lateral corner rounded off. Carapace with dorsal
line faintly sinuous, without spines. Pleon segments 1—4 with lateral groove;
telsonic segment the longest, inflated, encreasing in width backwards, hind
margin broadly convex, scarcely extended between the uropods. Eye very
large with 9 corneal lenses, one antero-median, pigment forming 2 small
lateral spots, suggestive of paired eyes as in Nannastacus (p. 168). Antenna
1 nearly as in C. forficula (nr. 4) but shorter and stouter. Antenna 2, length
of flagellum agreeing with lateral grooving of the pleon. Peraeopod 5, 5 joint
little more than half the second and about once and a half as long as the
4th joint. Uropods similar to those of ©. forjicu/a, but rami less curved,
peduncle about two-thirds as long as the telsonic segment. L. d 1:9 mm.
CG unknown.
Cruz Bay. St. Jan, Danish West Indies,
6. C. leptopus Calm. 1911 C. 1, Calman in: Tr. zool. Soc. London, o. 181v
p- 841, 344, 347 t.32 f.11—14.
Pseudorostral lobes not produced to meet in front of the eyelobe, front
margin concave, antero-lateral corner slightly prominent, rounded. Carapace
with dorsal line very slightly sinuous, without spines. Pleon segments 1—5
with lateral grooves, telsonic segment less than two-thirds as long as the ote
acutely extended between the uropods. Eye very large, 7 corneal lenses
large and prominent, one antero-median, no conspicuous pigment. Antenna
| rather slender, 1-jointed accessory flagellum minute. Peraeopod 2, 5 joint
more than twice as long as 4t*, Peraeopods 3—5 very slender; in 5‘ pair
the 5th joint about three-fourths as long as the 2™4 and 4 times as long as
the 4th. Uropods long and slender, peduncle more than twice as long as
the telsonic segment, endopod less than half the peduncle apart from its
long apical spine; exopod about five-sixths of endopod, but with a still longer
apical spine. L. Gd 2:2 mm, © unknown.
Cruz Bay, St. Jan, Danish West Indies.
7. C. hispida Calm. 1911 C.h., Calman in: Tr. zool. Soe. London, v. 181v
p. 341, 344, 347 t. 32 f. 15—18.
Scattered hairs on surface rather numerous. Pseudorostral lobes long,
obliquely upturned, acute, meeting in front of the prominent eyelobe; front
margin concave, antero-lateral corner quadrate. Carapace with slightly undu-
lating dorsal line, a tooth behind the eye, and another before the middle.
Pleon a little shorter than preceding part of the body, telsonic segment nearly
as long as the 5", not inflated, obtusely extended between the uropods. Eye
pigmented. Antenna | rather slender, 1-jointed accessory flagellum minute.
Peraeopods 3—5 slender, 5 pair with 5t* joint four-fifths as long as 2™¢,
thrice the 4 and twice the 6 joint. Peduncle of uropods about three-
quarters as long as the telsonic segment, a little shorter than the straight
182 Cumacea: 24. Nannastacidae, 6. Cumella
endopod including its indistinctly separated apical spine; exopod three-quarters
as long as endopod. L. 9 2°55 mm. dG unknown.
Gulf of Siam, depth 9 m
8. C. serrata Calm. 1911 C.s., Calman in: Tr. zool. Soc. London, v.181v p.341,
344, 348 t. 32 f. 19—24.
©. Surface without conspicuous hairs. Pseudorostral lobes, long,
obliquely upturned, acute, meeting in front of the prominent eyelobe; front
margin concave, antero-lateral corner with an acute tooth. Carapace com-
pressed, dorsal line slightly convex, with about 6 procurved teeth, the last
on a prominence at the back, separated from the rest by a shallow depression.
Pleon equal to preceding part of the body, telsonic segment shorter than
the 5', not inflated, obtusely extended between the uropods. Eye with distinct
lenses, without pigment. Antenna 1, 34 joint a little shorter than 2"4, Peraeo-
pod 1, 2™4 joint little more than half as long as the following joints com-
bined. Peraeopods 3—5 rather slender, 5% pair with 5% joint nearly as
long as 27, more than twice the 4 and nearly twice the 6" joint. Peduncle
of uropods longer by a fourth than telsonic segment, and by a third than
the endopod without Ne apical spine; exopod less than two-thirds as long
as the endopod. L. 2:2 mm.
G. Pseudorostral lobes less produced than in Q, obliquely truncate,
upper edge slightly concave, front margin receding, antero-lateral corner
rounded. Carapace with dorsal line nearly straight, without teeth. Pleon
shorter than preceding part of the body, segments 1—5 with lateral grooves.
Kye with a number of large and prominent lenses, but without pigment.
Antenna 2, flagellum as long as the body. Peduncle of uropods longer by
a half than the telsonic segment, the endopod less than two-thirds as long
as the peduncle. L. 2°3 mm.
Cruz Bay, St. Jan, Danish West Indies.
9. C. laevis Calm. 1911 C.1., Calman in: Tr. zool. Soe. London, v.181v p.341,
344, 350 t. 32 f. 25—27.
Surface without conspicuous hairs. Pseudorostral lobes short, truncate,
meeting in front of the prominent eyelobe, front margin receding, antero-
lateral corner rounded. Carapace with dorsal line horizontal, slightly sinuous,
without teeth. Pleon shorter than preceding part of body, telsonic segment
less than two-thirds as long as the 5%, the broadly rounded end little ex-
tended. Eye pigmented. Antenna 1 rather stout, 3¢ joint shorter than 24,
the 1-jointed accessory flagellum minute. Peraeopod 1, 2™4 joint about half
as long as remaining joints combined. Peraeopods 3—5 short, stout, 5 pair
with 5% joint little more than half as long as the 2" and about one half
longer than the 4 and little longer than the 6 joint. Peduncle of uro-
pods nearly twice as long as telsonic segment; endopod more than two-thirds
as long as the peduncle, and subequal to exopod, each with the terminal
spine distinct. L. @ 115 mm. G unknown.
Gulf of Siam, depth 2—15 m.
10. C. pygmaea O. Sars 1865 C. p., G. O. Sars in: Forh, Selsk. Christian.,
1864 p. 199 | 1869 C. agilis (S$), A. M. Norman in: Rep. Brit. Ass., v. 68 p.272 |
1879 C. pygmaea, G. O. Sars in: Arch. Naturv. Kristian., v.4 p.94 t.50—52 | 1897 C.p.,
T. Scott in: Rep. Fish. Board Scotl., v.15 p.135 | 1897 C. p., Ehrenbaum in: Wiss.
Meeresunters., v.21 p.419 | 1900 C. p., G. O. Sars, Crust. Norway, v.3 p.81 t. 55 | 1900
C. p., Zimmer in: Fauna arctica, v.1 p. 428, 438 | 1901 C.p., A. O. Walker in: J. Linn.
Cumacea: 24. Nannastacidae, 6. Cumella 185
Soc. London, v.28 p. 294 | 1902 C. p., G. S. Brady in: Nat. Hist. Tr. Northumb., v. 14
p-95 | 1905 C. p., Calman in: Sci. Invest. Fish. Ireland, 1904 app. 1 p. 23 |, 1909 C. p.,
A. M. Norman & G. S. Brady in: Nat. Hist. Tr. Northumb., ser.2 v. 311 p.34 | 1911
C. p., Calman in: Tr. zool. Soe. London, v. 181v p. 344.
©. Pseudorostral lobes short, meeting in front of the eye, front margin
straight and then deeply emarginate with acute antero-lateral angle. Cara-
pace large and deep, rather compressed in front, dorsal line arched, carinate
throughout, with 8 to 12 procurved denticles. Pleon narrow, a little shorter
than preceding part of the body, telsonic segment shorter
than the 5, obtusely extended between the uropods.
Eye rather conspicuous, circular. Antenna 1, accessory
flagellum minute, 2-jointed. Peraeopod 1, 24 joint distally
strongly serrate on both margins, 5‘ joint as long as
6th and 7t combined; in 2"¢ pair 5 and 7 joints
subequal. 62 half as long as 7; in the 3 following Fig. 123.
pairs 5¢ joint only a little longer than 6. Peduncle ¢. pygmaea (3) Sars,
of uropods serrate on medial margin, considerably longer fron,
than the endopod, which is a little longer than the
much narrower exopod. Colour, dark brown markings. L. 2°5 mm.
3. Pseudorostral lobes shorter, antero-lateral corner less prominent.
Carapace shallower, scarcely compressed in front, dorsal line nearly straight,
unarmed. Eye very large, subquadrate, with 7 lenses, the front median
one very prominent. Antenna 2, flagellum reaching the 3¢ pleon segment.
Peraeopods 1—4 with 2™¢ joint greatly dilated. L. 3 mm.
Norway, British Isles, Heligoland, Mediterranean; shallow water.
11. C. limicola O. Sars 1879 C. l., G. O. Sars in: Arch. Naturv. Kristian.,
y.4 p. 103 t. 53,54 | 1911 C. 1, Calman in: Tr. zool. Soc. London, v. 181v p. 344.
©. Integument pellucid, finely granular, hairy. Pseudorostral lobes
subacute, obliquely upturned, meeting in front of the eye, the front margin
receding. finely serrate above, the antero-lateral corner produced into a little
tooth. The carapace in dorsal view triangular, dorsal line little arched, cari-
nate, on the anterior half having 3 or 4 rather large, suberect teeth. Pleon
. shorter than preceding part of the body, telsonic segment shorter than 5",
longer than any of the others. Eye distinct. Antenna 1, peduncle of 3
subequal joints, 1-jointed accessory flagellum minute. Peraeopods 1 and 2
with the exopods unusually slight; 2"¢ joint of peraeopod 1 without strong
serration, 7 joint little over half the length of the 6%. Uropods short,
peduncle not longer than the telsonic segment, but considerably longer than
the endopod, which in turn is longer than the narrow exopod. L. 3 mm.
GS. Larger but more slender and less hairy than the Q, pseudorostral
lobes shorter, more strongly serrate, the antero-lateral tooth much shorter;
dorsal line of carapace horizontal, without carina or teeth. Pleon segments
laterally grooved, with finely serrate carina for upper boundary of the groove.
Eye of great size, of the 9 lenses 3 larger than the rest are conspicuous
over the back and sides of the pseudorostral projections. Antenna 2, fla-
gellum reaching end of pleon. Epipod of 18* maxilliped furnished with
branchial leaflets which are wanting in the ©. Peraeopods 1—4 with 24 joint
strongly developed. Peraeopod 5 more slender than in the ©. Uropods
rather more elongate than in the Q, peduncle longer than the telsonic seg-
ment. L. 3°5 mm.
Mediterranean, depth 11—19 m.
184 Cum,: 24. Nannastacidae, 6. Cumella, 25. Procampylaspididae, 1. Procampylaspis
C. abyssicola (Norm.) 1879 Spencebatea a., A. M. Norman in: Ann. nat. Hist.,
ser.5 v.83 p.67 | 1893 S.a., T. Stebbing in: Int. sci. Ser., v.74 p.306.
Pseudorostral lobes very short, broadly truncate. Carapace smooth, dorsal line
a bold gradual curve, lateral margin inarched, exposing bases of the legs. Pedigerous
segment 5 with ventral pair of spines and dorsal pair of spine-like points. Eye present.
Uropods very elongate, peduncle much longer than the rami, endopod than the exopod.
L.d4mm. 9 unknown.
W. of Donegal Bay, lat. 54° 53’ N., long. 10° 56‘ W., depth 2487 m.
C. carinata (H. J. Hansen) 1887 Campylaspis c., H. J. Hansen in: Vid. Meddel.,
v.39 p.207 t.7 f.4,4a | 1900 ?C. c., G. O. Sars, Crust. Norway, v.3 p.84 | 1905 ?Cumellopsis
c., Calman in: Sci. Invest. Fish. Ireland, 1904 app.1 p.28 | 1912 ?Cumella c., Calman
in? P. U.S. Mus., v.41 ‘p. 626;
Rather like C. australis (nr.1). Pseudorostral lobes shortly produced, meeting
in front of the prominent eye slightly upturned, apically subacute vertically truncate,
front margin concave below, with serration of the corner continued a short space along
the lower margin. Carapace somewhat compressed, dorsal line carinate throughout and
denticulate with procurved teeth in series dwindling to the rear, numerous irregular
tubercles on back and sides. Pedigerous segments 1—3 similarly armed with tubercles
or spines, 4th and 5th with back smooth but sides armed. Pleon rather shorter than
preceding part of body, segments 1—5 with upper front angle produced into a large
rough tubercle. Telsonic segment of equal length and breadth. Eye having 4 rather
widely separated prominent lenses. Uropods a little shorter than last 2 pleon segments
combined, the peduncle as long as the 5th pleon segment, with dwindling serration,
starting a little below base of medial margin; endopod two-thirds length of peduncle,
exopod a little shorter. L., Q with oostegites rather large, 4-5 mm.
W. Greenland, Disco, Nordfjord, depth 47 m.
25. Fam. Procampylaspididae
1912 Procampylaspidae, T. Stebbing in: Ann. S. Afr. Mus., v.10v p. 153, 167.
Telson wanting. No distinct eye. First antenna with accessory fla-
gellum very small. Mandible with narrow molar. Second maxilla normal.
First maxilliped with seventh joint small, unexpanded. Second maxilliped
with strong teeth projecting from concave margin of the terminal joint. Hxo- _
pods on first four pairs of peraeopods of male, only on first two of female.
No pleopods in either sex. Endopod of uropods simple. |
1 genus,
1. Gen. Procampylaspis Bonnier
1896 Procampylaspis (Sp. typ.: P. armata), J. Bonnier in: Ann. Univ. Lyon, v.26
p. 541 | 1900 P., G.O.Sars, Crust. Norway, v.3 p.83 | 1900 P., T. Stebbing in: Willey,
Zool. Results, pars 5 p.611 | 1905 P., Calman in: Sci. Invest. Fish. Ireland, 1904 app. 1
p. 22,33 | 1906 P., Calman in: Mt. Stat. Neapel, v.171v p.419 | 1907 P., Zimmer in:
Zool. Anz., v.31 p.372.
This being at present the only genus will have the characters of the
family.
4 species.
Synopsis of species:
Carapace with dorsal denticles numerous in male, wanting
1 in female:~ “catenins 6. eeeerrcuner Ber tos cal mn eae 1. P. bonnieri
\ Carapace with dorsal denticles scanty in either sex — 2.
Cumacea: 25. Procampylaspididae, 1. Procampylaspis 185
9 Carapace: bituberculate 95. 6.6.2 es) eM ee we Boe 2. P. compressa
Carapace not bituberculate — 3.
. J Second maxilliped with terminal joint tridentate... . . 3. P. tridentata
Second maxilliped with terminal joint quadridentate . .. 4, P. armata
1. P. bonnieri Calm. 1906 P. b., Calman in: Mt. Stat. Neapel, v.171v p. 422
t.27 £.21—27.
@. Pseudorostral lobes subacutely produced for some distance in front
of the evanescent eyelobe, the antero-lateral margin vertical to a small sinus
and well marked but not far produced acute corner, which is microscopi-
cally serrate. Carapace a little more than two-fifths of the body’s length,
smooth, inflated, narrowed in front as seen from above, dorsal line arched,
with slight depression to the rear. Pedigerous segments with side-plates
expanded, smoothly rounded, 1st and 2"¢ dorsally raised into bifid laminar
teeth procurved. Pleon segments smooth, the 5% widest at first third of
its length. Peduncle of uropods as long as 5‘ pleon and telsonic segment
combined; endopod as long as 5 pleon segment, two-thirds of the length
of the peduncle, exopod little over three-fourths of the endopod. L. 2°3 mm.
CG. In general form as in P, armata (nr. 4), but the eyelobe is evanes-
cent. There are a pair of small spines at tip of pseudorostral lobes above.
Dorsal surface of carapace with group of about 10 curved spines, submedian,
to the rear; spines in close order just over the lower margin from antero-
lateral corner to the hind margin. Side-plates of last 4 pedigerous segments
armed with marginal teeth, a pair of dorsal spines on each of the last 3
of these segments. First 4 pleon segments each with pair of forward slo-
ping straight dorsal spines near the front and a sharp lateral spine. Antenna 2,
last 2 joints of peduncle subequal. Exopod of uropods about four-fifths of
the length of the endopod. IL. 2°3 mm.
Mediterranean, near Capri; depth 950—1200 m.
2. P. compressa Zimmer 1907 P. c., Zimmer in: Zool. Anz., v.31 p. 372 | 1912
P. compressus, 'T. Stebbing in. Ann. S. Afr. Mus., v. 10v_p. 167.
©. Pseudorostral lobes little produced, and without lateral sinus.
Carapace not half the length of the body, a little compressed; eyelobe tuber-
culiform, carrying a pair of denticles; a submedian pair of processes behind
the middle of the length of the carapace carry each on its summit a couple
of procurved denticles. The pedigerous segments combined are about half
as long as the carapace. The peduncle of the uropods is rather longer than
the 5t pleon segment, the endopod about two-thirds as long as the peduncle
and rather longer than the exopod. Colour yellowish white. L. Q with
commencing marsupium, about 4 mm.
German South Pole Expedition, Winter station lat. 660 2‘ S., long. 89° 38’ E,,
depth 385 m,
3. P. tridentata Stebb. 1912 P. t., T. Stebbing in: Ann. S. Afr. Mus., v. 10v
p. 167 t. 62 (14), tf. A, B.
Pseudorostral lobes a little upturned, meeting for a very short space
in front of the narrow bidenticulate eyelobe, laterally forming a shallow
sinus defined by a small tooth, which is followed a little way to the rear
by a similar tooth on the lower margin. Carapace well arched, not actually
carinate, with scattered hairs on the surface and behind the centre a little
denticle. Pleon segments laterally carinate, 5 the longest, distally nar-
186 Cumacea: 25. Procampylaspididae, 1. Procampylaspis
rowed, telsonic segment not longer than broad. Antenna 1, 15° joint longer
than 24, 24 than 34, principal flagellum 3-jointed, accessory 1-jointed.
Antenna 2, 18* joint of the long flagellum knobbed at the base. Branchial
leaflets numerous. Maxilliped 2, 2™4 joint not twice as long as_ broad,
the much curved 7 produced into a sharp point, the concave margin forming
3 strong teeth, of which the middle one is the longest. Maxilliped 3, 2n4
joint stout, much longer than rest of limb, 34 very small, 4th widened dis-
tally. Peraeopod 1, 2"4 joint stout, not longer than rest of limb, 3° unu-
Fig. 124. Fig. 125. Fig. 126. Fig. 127.
P. tridentata P. tridentata (dQ) P. tridentata P. tridentata Stebbing,
Stebbing, mandible. Stebbing, maxilli- Stebbing, maxilli- maxilliped 3.
ped 1. ped 2.
sually long, more than twice as long as broad, 6% longer than any except
the 224. Peraeopod 2, 34 joint short but outdrawn to a conspicuous apical
spine, 7% as long as 4* and 5% combined, 4 times as long as the 6".
In the 34 and 4* pairs the stout 2™4 joint is much narrowed distally. Pe-
duncle of uropods with finely serrate margins; endopod about two-thirds as
long with 10 spines on the medial margin, and a much larger apical spine;
the exopod much more slender and rather over two-thirds as long. L. 3
45mm. © unknown.
S. Africa, Cape Natal distant N. by E. 24 miles; depth 805 m.
4. P. armata Bonnier 1896 P.a.-+ P. echinata, J. Bonnier in: Ann. Univ. Lyon,
7.26 p.541 t.29 f.la—r; p.544 t.29 f.2a—t | 1905 P.a,, Calman in: Sci. Invest. Fish.
Ireland, 1904 app.1 p.33 | 1906 P. a., Calman in: Mt. Stat. Neapel, o.171v p. 419
t.27 f.13—20 | 1912 P. armatus, T. Stebbing in: Ann. S. Afr. Mus., v. L0v p. 167.
Q. Pseudorostral lobes upturned, meeting with blunt apices for a short
distance in front of the long narrow eyeless eyelobe, descending nearly verti-
cally to a small sinus defined by a small blunt antero-lateral tooth. Cara-
pace oval, dorsal line moderately arched, not carinate, but just behind the
centre bearing a strong median tooth, the eyelobe on its slightly raised apex
having a pair of small tubercles. First pedigerous segment raised to a
transverse, procurved lamina with median bifid tooth overlapping hind margin
of carapace; a similar less prominent tooth on second segment is not pro-
Cum.: 25. Procampylaspididae, 1. Procampylaspis, 26. Campylaspididae, 1. Campylaspis 187
curved; fourth and fifth segments dorsally granular. Pleon segments except
the telsonic granular all round, the 5 segment scarcely inflated anteriorly.
Uropods (in Calman, fig. 14) of similar dimensions to those of P. bonnieri,
but peduncle twice the length of the endopod (Calman, text). © immature,
35 mm. Ovigerous Q, with pleon segments nearly smooth.
¢. Antero-lateral corner scarcely produced; median dorsal tooth of
carapace less prominent, sometimes bifid; a pair of small dorsal teeth are
adjacent to the hind margin and on each side about 6 denticles occupy the
last quarter of the length of the carapace a little way above the lower mar-
gin: the eyelobe is like that of Q (Calman); in Bonnier’s P. echinata there
is no eyelobe figured, and the carapace is quite smooth. Pedigerous seg-
ments 1, 2, not produced into dorsal teeth as in Q, segments 2—5 each
with a pair of dorsal tubercles, and having the side-plates expanded and
margined with rows of spiniform teeth. Pleon segments 1—5 have the
granules spiniform above and below the antennal groove. Last segment of
peduncle in second antennae nearly twice as long as the preceding one.
Peduncle of uropods considerably longer than the last two segments of
pleon combined, fully twice as long as the endopod. L. adult 3S nearly
6 mm. (Bonnier), 4 mm (Calman), immature Gd (P. armata, Bonnier)
scarcely 5 mm.
Bay of Biscay, depth 950 m; Irish Sea, 50 miles W. N. W. of Cleggan Head,
212m; and 77 miles W. N. W. of Achill Head, county Mayo 699 m: Mediterranean,
near Capri, 200—1100 m.
26. Fam. Campylaspididae
1878 & 1879 Campylaspidae, G. O. Sars in: Arch, Naturv. Kristian., 0.3 p. 466;
v.4 p.74 | 1887 C., G. O. Sars in: Rep. Voy. Challenger, v.19 pars 55 p. 8,65 | 1900 C.,
G. O. Sars, Crust. Norway, v.3 p.82 | 1900 C., Zimmer in: Fauna arctica, v1 p.451,
435 | 1912 C., T. Stebbing in: Ann. S. Afr. Mus., v. 10v p. 133, 170.
Telson wanting. First antenna with accessory flagellum very small.
Second antenna of female imperfectly developed. Mandible with molar slender,
acute. First maxilla with two filaments to the palp or only one. Second
maxilla an undivided plate. First maxilliped of four joints, the last minute.
Second maxilliped with the terminal joint straight, spiniferous, or spine-like.
Exopods on first four pairs of peraeopods of male, only on first two of female.
No pleopods in either sex. Uropods with endopod simple.
1 genus.
1. Gen. Campylaspis G. 0. Sars
1865 Campylaspis (Sp. typ.: C. rubicunda), G. O. Sars in: Forh. Selsk. Christian.,
1864 p.200 | 1866 C., G. O. Sars in: Nyt Mag. Naturv., v.15 p.105 | 1878 & 1879 C.,
G. O. Sars in: Arch. Naturv. Kristian., v. 3 p.466; v4 p.75 | 1887 C., G, O. Sars in:
Rep. Voy. Challenger, v.19 pars 55 p. 65 | 1896 C., J. Bonnier in: Ann. Univ. Lyon, v. 26
p. 538 | 1900 C., G. O. Sars, Crust. Norway, v.3 p.83 | 1900 C., Zimmer in: Fauna arctica,
v.1 p.429 | 1905 C., Calman in: Sci. Invest. Fish. Ireland, 1904 app. 1 p.34 | 1911 C,,
Calman in: Tr. zool. Soe. London, v.181v p. 364 | 1912 C., T. Stebbing in: Ann, S. Afr.
Mus., v.10v p.171.
This being at present the only genus, the characters of the family
suffice for its definition.
23 species accepted.
188
~l
ea)
10 ;
11
12
13 «
14
15
16
17
18
19
20
|
|
|
|
|
\
|
|
|
|
|
|
\
|
|
|
Cumacea: 26. Campylaspididae, 1. Campylaspis
Synopsis of species:
Uropods broad .
Uropods not broad — 2.
Carapace smooth — 3.
Carapace not smooth — 9.
Eyelobe obsolete ... . Ai Ade Gag
Eyelobe not obsolete — rk:
Carapace less than half the total length of the body
Carapace not less than half the total length — 5.
Uropods with exopod not shorter than the endopod
Uropods with exopod shorter than the endopod — 6.
Eyelobe without corneal lenses — 7.
Eyelobe with corneal lenses — 8.
Eyelobe broad, with whitish pigment .
EKyelobe longer than broad, eye indistinct .
Peraeopod 2 with 7th joint longer than 5th and 6th
combined .. .
Peraeopod 2 with 7th join eiarier nth ‘5th and gth
combined ..... : a oc ic ent dude p
Carapace spinose .. . AG kc
Carapace not spinose — lo.
Carapace with conspicuous carinae, not conspicuously
verrucose — 11.
Carapace verrucose, not conspicuously carinate — 16.
Carapace with 3 aay of lateral keels, the hindmost
bifurcate
Carapace with only 2 pairs af tetera keels — 12.
Kye wanting — 13.
Eye present — 14.
Carapace with transverse keel
Carapace without transverse keel .
eo) <0) fers etre
Eyelobe long, distally widened . :
Eyelobe short, not distally widened — 15.
Peraeopod 2 with 7th joint shorter than 5th and 6th
combined .
Peraeopod 2 with 7th sein longer fan 5th anal 6th
combined Fo oo 6446 0066
Carapace with tubercles are combined — 17.
Carapace without tubercles carinately combined — 19.
HKyelobe long, linguiform
Eyelobe very small — 18.
Peraeopod 2 with 7th joint about equal to 4th, 5th
and 6th combined .. . :
Peraeopod 2 with 7th joint Rarely doneer then “5th .
Carapace with few tubercles; peduncle of uropods
SIMO OLN wenn rentlte
Carapace with numerous taberolens redunele a aropeds
serrate — 20.
Maxilliped 3 with 4th i i ucts
panded . 5
ee ex-
Maxilliped 3 with 4th joint ‘Gala — ‘21.
~l
10.
ite
12.
13.
14.
15.
16.
Whe
18.
19.
20.
. platyuropus
. nitens
. paeneglabra
. pulchella,
. pacifica
. orientalis
. rubieunda
. glabra
. Spinosa
. costata
. vitrea
. ovalis
- macrophthalma
. undata
. suleata
. horrida
. rostrata
. maculata
. affinis
. frigida
Cumacea: 26. Campylaspididae, 1. Campylaspis 189
Pleon segment 5 not transversely sulcate .... . 21. C. verrucosa
f
an \ Pleon segment 5 transversely sulecate — 22.
J
\
99 Pleon segments without dorsal teeth ......-. 22. C. nodulosa
“| Pleon segments with dorsal teeth. .......- 23. C. antarctica
1. C. platyuropus Calm. 1911 C. p., Calman in: Tr. zool. Soc. London, v. 181v
p. 341, 364 t. 34 f. 25—29.
Pseudorostral lobes short, truncate. Carapace about half the whole
length of the body, dorsal line behind the very small eyelobe convex,
flanked by pair of dorso-lateral ridges curving towards it near the end. Pleon
segments stout, slightly depressed, telsonic segment hexagonal, broader than
long. Maxilliped 3 with well developed exopod, 2"4 joint of endopod as long
as next 3 joints combined, 4 greatly expanded, less than twice as long
as broad, longer than 3 following joints combined. Peraeopod 1, 3° joint
with medial margin produced into a large thin triangular tooth. Peraeopod 2,
7th joint rather stout, shorter than 5‘ and 6 joints combined. Uropods
remarkably broad and flattened, peduncle less than twice as long as the
telsonic segment, the broad endopod three-fourths the length of the peduncle,
the narrower exopod a little shorter. L. © with developing oostegites, 1°8 mm.
GS unknown.
Gulf of Siam, depth 9—19 m.
2. C. nitens Bonnier 1896 C. n., J. Bonnier in: Ann. Univ. Lyon, v.26 p. 538
t.28 f.4a—r! 1905 ? C.n., Calman in: Sci. Invest. Fish. Ireland, 1904 app. 1 p.10, 35.
Pseudorostral lobes short, obtuse. Carapace smooth, oval, dorsal line
much arched, eyelobe very small. Telsonic segment very obtusely extended
between the uropods. Eye wanting. Antenna 1, the 2™¢ joint of peduncle
the shortest, flagellum of 3 or 4 joints, accessory a minute tubercle. Maxilla 1,
palp with 2 filaments. Maxilliped 1 with some 20 branchial leaflets on the
epipod. Mavxilliped 2, terminal joint as it were embedded in the preceding and
carrying 3 apical spines. Maxilliped 3, with denticulate medial margin to
joints 3 to 6. Peraeopod 2, 7% joint rather stout or in Q slender (Calman),
shorter than 2 preceding joints combined. Uropods as long as the pleon,
peduncle twice as long as the endopod or more (Calman), exopod a little
shorter than endopod. L. young G, a little less than 5 mm, immature ©,
4-5 mm.
3ay of Biscay, depth 950 m; W. coast of Ireland, 50 miles W. N. W. of Tearaght
Lighthouse, county Kerry, depth 570 m.
3. C. paeneglabra Stebb. 1912 C. paeneglaber, T. Stebbing in: Ann. S. Afr,
Mus., v. l1Ov p.172 t. 64.
Near to C. glabra (nr. 8), size, shape, mouth organs, pellucid spots on
carapace, being in close agreement, but the S. African species is rather
larger, the pseudorostral lobes are slightly upturned, the carapace is not
quite smooth, the eyelobe is more elongate, showing no lenses, the 1%* and
9d pedigerous segments are not transversely ridged, the 2™¢ peraeopod has
the 7 joint longer instead of shorter than the 5% and 6 joints combined,
and the 5” peraeopods are more slenderly built, with the 2"¢ and 5‘ joints
longer relatively to the other joints. L. d about 43 mm. Q unknown.
S. Africa, Cape Natal N. by E. 24 miles, depth 805 m.
190 Cumacea: 26. Campylaspididae, 1. Campylaspis
4. C. pulchella O. Sars 1871 C.p., G. O. Sars in: Ofv. Ak. Férh., v.28 nr. 6
p.806 | 1873 C. p., G. O. Sars in: Svenska Ak. Handl., n. ser. v.11 nr.d p.14 t.1
f. 14—16.
Pseudorostral lobes shortly and rather sharply produced in front of
the prominent oblong eyelobe, the front margin receding without any sinus
or antero-lateral corner. Carapace almost globular, smooth. 1% pedigerous
segment only visible dorsally, 2°¢ raised into a prominent transverse crest,
the last 3 successively much lower and narrower. Pleon thin and_ short,
telsonic segment much shorter than the 5%, very slightly extended between
the uropods, almost truncate. Eye indistinct. Antenna 1 long and slender,
3 joints of peduncle subequal. Peraeopods 3 to 5 very slender, with the
4th joint elongate. Uropods elongate, peduncle with medial margin serru-
late, more than twice the length of the rami, of which the exopod is
described and figured as slightly longer than the endopod. L. © about 3 mm.
G unknown.
West Indies, Anguilla, depth 877—565 m.
5. C. pacifica O. Sars 1887 C. p., G. O. Sars in: Rep. Voy. Challenger, v.19
pars 55 p. 9,66 t. 10 f. 6.
Pseudorostral lobes shortly produced in front of rather broad eyelobe,
antero-laterally receding obliquely to a small sinus with the lower angle
rounded off. Carapace dilated, more than half total length of the body,
overarching the pedigerous segments, boldly arched with faint undulation
of the dorsal line, otherwise smooth. Pleon segments 1—5 slightly cari-
nate laterally, 5% the longest; telsonic segment obtusely extended be-
tween the uropods. Eye showing whitish pigment, without lenses. Uro-
pods about equal in length to pleon segments 3 to 6 combined, the
smooth peduncle more than twice as long as the endopod, which is
longer than the more slender exopod. Colour brownish red. L. OQ 3 mm.
G unknown.
Samboangan (Philippine Islands).
6. GC. orientalis Calm. 1911 C. o., Calman in: Tr. zool. Soc. London, v. 181v
p. 341, 365 t.35 f. 1—6.
Near to C. rubicunda (nr. 7). Pseudorostral lobes short, truncate
in dorsal view, with faint lateral sinus. Ocular lobe longer than broad, eye
indistinct. Pleon segments not depressed, not laterally carimate. Maxilli-
ped 3, 24 joint much longer than rest of joints combined, 3¢ joint slightly
produced on medial margin, 4** not wider than 34, not nearly as long as
5th and 6 combined, 6 much longer than 5, none of the joints conspi-
cuously serrate. Peraeopod 1, 4%" joint not expanded, hardly longer than
5th. Peraeopod 2, 7 joint shorter than 5 and 6% combined, not very
slender. Uropods with peduncle about twice as long as telsonic segment or
as the endopod, with medial margin neither serrate nor spiniferous; endopod
with numerous spines, decidedly longer than the exopod. Colour, traces of
reddish purple. L. 9 55 mm. GC unknown.
Korea, depth 66 m.
7. C. rubicunda (Lilj.) 1855 Cumar., W. Liljeborg in: Ofv. Ak. Forh., v.12
p. 121 | 1865 Campylaspis r., G. O. Sars in: Forh. Selsk. Christian., 1864 p. 202 | 1870
C. r., G. O. Sars in: Forh. Selsk. Christian., 1869 p. 160,161 | 1873 C. r., G. O. Sars
in: Svenska Ak. Handl., n. ser. v.11 nr.6 p.10 t.4 | 1879 C. r., A. M. Norman in: Ann.
Cumacea: 26. Campylaspididae, 1. Campylaspis 191
nat. Hist., ser.5 v.3 p.73 | 1887 C.r., H. J. Hansen in: Vid. Meddel., v.39 p. 207, 209
1897 C. r., T. Scott in: Rep. Fish. Board Scotl., v.15 p. 133 | 1900 C. r., G. O. Sars,
Crust. Norway, v.3 p.84, 108 t. 56,57 | 1900 C. r., Zimmer in: Fauna arctica, v.1 p. 429,
435 | 1909 C. r., A. M. Norman & G. S. Brady in: Nat. Hist. Tr, Northumb., ser. 2
v.31 p.35 | 1911 C. r., Stappers in: Camp. arct. Orléans, Crust. Malac. p.122 | 1912
C. r., Calman in: P. U.S. Mus., v.41 p. 627.
Q Pseudorostral lobes very short, but distally acute, antero-lateral
corners obsolete. Carapace very large, inflated, half the length of the body,
quite smooth. Pedigerous segments 1 and 2 raised to transverse procurved
ridges, each with median produced point. Pleon segments short, depressed,
telsonic segment slightly broader than long, obtusely extended. Hye distinct,
semicircular, rather prominent, with 2 corneal lenses (Hansen). Maxilla 1
with single apical filament on the slender palp. Maxilliped 1, epipod with 10
branchial leaflets, terminal joint rather small, conical, with 1 apical setule.
Maxilliped. 2, the short terminal joint carrying 4 strong spines. Maxilliped 3
with the joints partly serrate, 4'* joint considerably expanded, as long as
5th and 6t combined. Peraeopod 2, 7 joint slender, longer than 2 pre-
ceding joints combined. Uropods not very long, peduncle with serrate medial
margin, twice as long as the endopod, which is a little longer than the exo-
pod. Colour bright red except frontal lobe and adjacent part which are opaque
white. L. 5mm. G Like QO, except that it is more slender, pseudorostral
lobes shorter, more obtuse; carapace shallower, epipod of maxilliped 1 with
18 branchial leaflets, uropods more elongate, peduncle fringed with plumose
setae, not twice as long as the endopod, which carries more spines than in
the ©, and is more decidedly longer than the exopod. L. about 6 mm.
Lat. 75° 45‘ N., depth 1977 m; Greenland, Norway, Kullaberg, Kattegat, British
Isles, Atlantic coast of N. America; depths from 22 to 188m; Nova Zembla, 90 m.
8. C. glabra O. Sars 1879 C. g., G. O. Sars in: Arch. Naturv. Kristian., v. 4
p.77 t.44—47 | 1900 C. g., G. O. Sars, Crust. Norway, v.3 p.86 t. 58 | 1902 C. g., G.S.
Brady in: Nat. Hist. Tr. Northumb., v. 141 p. 95 | 1905 C. g. (?C. rubicunda var.), Calman
Fig. 128.
C. glabra (2) Sars.
in: Sci. Invest. Fish. Ireland, 1904 app. 1 p. 34 | 1906 C. g., Calman in: Mt. Stat. Neapel,
v. 17iv p.423 | 1909 C.g., A. M. Norman & G. S. Brady in: Nat. Hist. Tr. Northumb.,
ser.2 v. 311 p. 35.
Very near to C. rubicunda (nr. 7), but considerably smaller and differ-
ing in colour, pedigerous segments 1 and 2 less strongly produced dorsally ;
eyelobe semioval, with 3 corneal lenses, pigment purple under opaque white
192 Cumacea: 26. Campylaspididae, 1. Campylaspis
covering; terminal joint of maxilliped 1 more rudimentary; peraeopod 2
with terminal joint shorter instead of longer than the 2 preceding joints
combined; peduncle of uropods little serrate on medial margin or distinctly
(Calman), not twice as long as the endopod or a little more than twice
(Calman) and little more than twice the exopod in the 3, a little less than
Fig. 132.
C. glabra Sars,
maxilliped 1, apex.
Fig. 129.
C. glabra Sars, mandibles.
Fig. 130. Fig. 131. Fig. 133. Fig. 134.
C. glabra Sars, maxilla 2. C. glabra Sars, C. glabra (2) Sars, —_Q, glabra Sars, maxilli-
maxilliped 1. maxilliped 2.} ped 3.
twice in the ©. Colour whitish, sometimes with tinge of pink (Calman).
L. © scarcely exceeding 3 mm, CG nearly 4 mm.
Mediterranean, N. of England, W. coast of Ireland, Norway; depths 33—1100 m.
9. C. spinosa Calm. 1906 C. s., Calman in: Mt. Stat. Neapel, v 171Vv p. 412,
426 t.28 f. 35—43.
Q. Pseudorostral lobes sharply upturned, meeting only just in front
of the slitlike eyeless eyelobe, thence receding convexly to a scarcely indica-
ted antero-lateral corner. Surface of carapace beset with spinules and armed
on each side with about 8 strong spines reaching from the pseudorostral
lobes to the rear; there are half a dozen long setae dorsally, and the lower
margin anteriorly carries several procurved spines. First pedigerous segment
almost completely concealed, each of the others having a pair of dorsal spines.
Of the pleon segments the first 4 are similarly armed, the 5 has 3 dorsal
and 2 postero-lateral spines; the telsonic segment is pentagonal, as broad
as long, with 3 medio-dorsal spines and a subapical pair. The mouth-parts
Cumacea: 26. Campylaspididae, 1. Campylaspis 193
distinguished from those of C. rubicunda (nr. 7) by molar of mandible rather
" longer and more slender, palp of maxilla 1 with 2 filaments, epipod of maxilli-
ped 1 with only 3 branchial sacs. Maxilliped 2 with 3“ joint distinct, terminal
joint carrying 3 spines. Maxilliped 3 with dentate margins, 4th joint little
broader than 5, equal in length to 5 and 6 joints combined; its apical
setae very long. Peraeopod 2, 2™4 joint less than a third of the total length,
5th with a serrate margin, 7* as long as 5* and 6" combined. Peduncle
of uropods more than twice as long as the telsonic segment, indistinctly
serrate; exopod a little longer than the endopod, about half as long as the
peduncle, its long terminal spine serrate. L., subadult, 3°1 mm.
3. Imperfectly known. Pseudorostrum more horizontal than in Q,
carapace less deep, the spines for the most part reduced to tubercles, but
those on pleon segments even longer than in Q. L. 3°77 mm.
Mediterranean, near Capri; depth 950—1100 m.
10. C. ecostata O. Sars 1865 C.c., G. O. Sars in: Forh. Selsk. Christian., 1864
p. 204 | 1872 C.c., G. O. Sars in: Forh, Selsk. Christian., 1871 p.272 | 1894 C.c., A. M.
Norman in: Ann. nat. Hist., ser.6 v.13 p.277 t.12 f.9 | 1897 C.c¢., T. Scott in: Rep.
Fish. Board Scotl., v.15 p.135 | 1897 C. c., Ehrenbaum in: Wiss. Meeresunters., v.21
p.420 | 1900 C. c., G. O. Sars, Crust. Norway, v.3 p.87 t.60 | 1900 C. c., Zimmer in:
Fauna arctica, v.1 p. 425,435 | 1909 C.c., A. M. Norman & G. S. Brady in: Nat. Hist.
Tr. Northumb., ser. 2 v.31 p. 35,
Pseudorostral lobes meeting for a short space subacutely in front of
the narrowly oval eyelobe; antero-lateral corner obtuse, slightly produced
below a shallow sinus. Carapace much dilated, but not nearly half the
length of the body, dorsal line evenly vaulted behind, sides sculptured with
3 pairs of distinct oblique folds or subcrescentic carinae, the hindmost at the
middle bifurcating, so that a 4 fold nearly reaches the hind margin and
then turns forward to rejoin the 3¢ in the submedian line. Pedigerous
segments 1, 2, slightly produced dorsally, each with median procurved or
erect point; 5 pedigerous and first 4 pleon segments carry each a pair of
adjacent dorsal tubercles. Telsonic segment pentagonal, length and breadth
equal. Eye distinct, showing 4 lenses (in fig.). Mavxilla 1, palp with 2 filaments.
Maxilliped 1, terminal joint less rudimentary than usual in the genus, Maxilli-
ped 2, terminal joint carrying 3 spines, the smallest intermediate. Maxilliped 3,
4th joint not greatly expanded, nor nearly as long as the next 2 joints combined.
Peraeopod 2, 7 joint, scarcely longer (G. O. Sars in text) or rather shorter
(in fig.) than 2 preceding joints combined. Peduncle of uropods unarmed
in ©, with the usual setae in O, not twice as long as the endopod, which
is a little longer than the exopod; the latter in the © is figured a little more,
in C a little less, than half the length of the peduncle. Colour pale reddish.
L. Oo 5 mm, CO 6°5 mm.
Norway, Heligoland, British Islands; depths 388—274 m.
11. C. vitrea Calm. 1906 C. v., Calman in: Mt. Stat. Neapel, v.171v p.412,
425 t.28 f. 28—34 | 1912 C. v., Calman in: P. U.S. Mus., v 41 p. 628.
Integument thin and semitransparent (much less so in supposed @).
Pseudorostral lobes well produced in front of the short narrow eyeless eye-
lobe, apically acute, convex in dorsal outline, antero-laterally receding to a
shallow sinus and obtuse corner. Carapace more than half the length of
the body, with 2 pairs of finely serrate lateral oblique keels, the upper pair
joined by a straight transverse keel about one-fourth of the length of the
Das Tierreich. 39, Lief.: T. R. R. Stebbing, Cumacea. 13
194 Cumacea: 26. Campylaspididae, 1. Campylaspis
carapace from its end; from the points of junction a pair of short keels
run backwards to the lower lateral keels and are united in the rear by a_
broad bilobed ridge, thus enclosing a concave subquadrate area; the lateral
areas between the lateral keels are also concave, and in front those keels
are connected by a low vertical ridge. Pedigerous segments 1—3 have trans-
verse dorsal crests, and the 4% and 5t" have each a pair of cristiform dorsal
teeth. Pleon segments 1—4 have paired dorsal, lateral, and ventro-lateral
ridges (less prominent in supposed CG’), the 5 has the three pairs on its front
half, but behind has only a medio-dorsal ridge and a lateral pair; the telsonic
segment has a medio-dorsal ridge in front and a lateral pair. Appendages
in general agreement with C. macrophthalma (nr. 13), but antenna 1, 274
joint distinctly longer than 34, distal joints of peraeopod 1 a little longer,
7th joint of peraeopod 2 one-third of the total length of the limb, following
peraeopods rather longer and more slender than im ©. macrophthalma, and
the uropods longer, the peduncle more than two and a half times the length
of the telsonic segment and over twice the length of the subequal rami.
L. Q subadult, 4°77 mm; immature J?, 7:2 mm.
Q Mediterranean, near Capri, depth 950—1100 m; G? lat. 40° 02‘ 00" N., long.
68° 50’ 30” W., depth 985 m.
12. C. ovalis Stebb. 1912 C. 0. T. Stebbing in: Ann. 8S. Afr. Mus., 7 10v
p.171 t. 63.
Integument pellucid. Pseudorostral lobes shortly and subobtusely produ-
ced in front of a very small eyeless eyelobe. Carapace in dorsal aspect
Fig. 135. Fig. 136. Fig. 137.
C. ovalis (Q) Stebbing. C. ovalis (dj) Steb- C. ovalis Stebbing,
bing, maxilliped 1. maxilliped 2.
forming an oval broadest in front, the median part shallowly convex, the
boundary constituted by a somewhat raised keel, below which the sides are
strongly inflexed, with a second longitudinal keel near the lower border.
Pedigerous segments 2—5 successively narrowed, with lateral angles more or
less rounded. Fifth pleon segment the longest, telsonic pentagonal, the two
combined rather shorter than peduncle of uropods. First antenna small,
Cumacea: 26. Campylaspididae, 1. Campylaspis 195
flagellum 3-jointed, accessory minute. Upper lip with obtuse-angled margin.
Maxilla 1, palp with 2 filaments. Maxilliped 1, terminal joint minute, at medial
apex of preceding joint. Maxilliped 2 with long spear-ended spine attached
to the short 3¢ joint, 7° joint spine-like. Maxilliped 3 with very small 7th
joint. Peraeopod 1 very like maxilliped 3, but with the joints longer.
Peraeopod 2, 7% joint rather longer than the 5, Peraeopod 3 with second
joint long, much narrowed distally. Peduncle of uropods serrate, twice and
two-thirds as long as the endopod, which is a little longer than the exo-
pod. L. carapace of somewhat immature male, 3°3 mm long, 2°5 mm broad.
South Africa, Cape Natal N. by E. 24 miles, from depth of 805 m.
13. C. macrophthalma O. Sars 1879 C. m., G. O. Sars in: Arch. Naturv. Kristian.,
v.4 p.87 t. 48,49 | 1906 C. m., Calman in: Mt. Stat. Neapel, v.171v p. 424.
Pseudorostral lobes very shortly produced in front of the eyelobe, ob-
tuse at apex, antero-lateral corner below the short sinus distinct. The cara-
pace scarcely half the length of the body, sculptured with 2 pairs of lateral
longitudinal keels, united in front by a transverse short keel. Eyelobe of
unusual shape, linguiform, widest in front where it is prominent and oceupied
by the 3 corneal lenses of the eye. Pedigerous segment 1 partly concealed,
the 3 following segments with dorsal outline almost forming a right angle
with the carapace. Pleon segments laterally keeled. Antenna 1, with the
3 joints of the peduncle nearly subequal, flagellum much longer than any
one of them. Maxilla 1, palp with 2 filaments. Maxilliped 1, the terminal
joint carrying 3 short spines. Maxilliped 2, 2°¢ joint with peculiarly widened
seta at apex of medial margin, terminal joint with 3 spines of which the
intermediate is very small. Maxilliped 3, 4‘ joint only moderately widened,
not so long as next 2 joints combined. Peraeopod 1, 4 joint much longer
than the 5%, Peraeopod 2, 7% joint shorter than the 2 preceding joints
combined. Peduncle of uropods serrate on both margins, twice as long as
the serrate endopod, which is a little longer than the much narrower exo-
pod. Colour bright red. L. 9 5mm. G not certainly known.
Mediterranean, at Messina, depth 38, near Capri, depth 130—200 m.
14. C. undata O. Sars 1865 C.u., G. O. Sars in: Forh. Selsk. Christian., 1864
p.205 | 1900 C. w., G. O. Sars, Crust. Norway, v.3 p.88 t.61 | 1900 C. u., Zimmer in:
Fauna arctica, v.1 p. 430, 435.
Near to C. costata (nr. 10), but larger, carapace less strongly arched
above, dorsal surface undulated with irregular protuberances and ridges, laterally
sculptured with 2 very prominent rather flexuous carinae, the upper contin-
ued. on to the side of the pseudorostral projection, the lower ending at the
antero-lateral corner; peraeopod 2 with 7 joint not longer than the 5;
peduncle of uropods more than twice as long as the endopod. Colour pale
carneous. L. 9 6 mm, C 7 mm.
Norway, off the Lofoten Islands; depths 188—377 m.
15. C. suleata O. Sars 1870 C.s., G. O. Sars in: Forh. Selsk. Christian., 1869
p. 161 | 1872 C. s., G. O. Sars in: Forh. Selsk. Christian., 1871 p.272 | 1886 C..s.,
G. O. Sars in: Norske Nordhavs-Exp., v.6 nr. 15 p.24 | 1900 C.s., G. O. Sars, Crust.
Norway, v.3 p.86 t.59 | 1900 C. s., Zimmer in: Fauna arctica, v.1 p. 430,435 | 1905
C. s., Calman in: Sci. Invest. Fish. Ireland, 1904 app. 1 p.35 | 1906 C. s., Calman in:
Mt. Stat. Neapel. v. 171v p. 424.
13*
196 Cumacea: 26. Campylaspididae, 1. Campylaspis
Pseudorostral lobes shortly and obtusely produced in front of the semi-
circular eyelobe, antero-lateral corners obsolete below a faintly indicated sinus.
Carapace very large, boldly arched, dorsal surface anteriorly undulated, lateral
faces deeply sulcate between 2 projecting carinae which extend obliquely from
the most prominent part of the branchial region to the base of the pseudo-
rostral projection. Pedigerous segments 1, 2, elevated to procurved lamellae.
Eye apparently having 4 lenses. Maxilla 1, palp with 2 filaments. Maxilliped 1,
terminal joint extremely small. Maxilliped 2, terminal joint with 4 diver-
ging spines. Maxilliped 3, 4% joint not nearly as long as 2 following joints
combined. Peraeopod 2, 7‘ joint longer than 2 preceding joints combined.
The 3 following pairs with medial margin of 5 joint serrate. Peduncle of
uropods with medial margin serrate, nearly twice as long as endopod, and
fully twice the exopod. Colour whitish, middle tinged with red. L. Q 45 mm,
3 5 mm.
Norway, Lofoten Islands, Hardanger and Trondhjem-Fjords, Hus, depths
150-470 m; W. coast of Ireland, depths 238—364 m; Mediterranean, depths 130—200 m.
16. C. horrida O. Sars 1870 C.h., G. O. Sars in: Forh. Selsk. Christian., 1869
p. 162 | 1872 C. h., G. O. Sars in: Forh. Selsk. Christian., 1871 p.272 | 1894 C. h.,
A. M. Norman in: Ann. nat. Hist., ser.6 v.13 p.278 t.12 f.6,7 | 1900 C. h., G. O. Sars,
Crust. Norway, v.3 p.89 t.62 | 1900 C.h., Zimmer in: Fauna arctica, v1 p. 429,435 |
1905 GC. h., Calman in: Sci. Invest. Fish. Ireland, 1904 app. 1 p.35 | 1912 C. h.?, Calman
in? P21. S. Mus.;\0, 41) p.'627.
Pseudorostral lobes shortly produced in front of the linguiform eye-
lobe, truncate, sinus and antero-lateral corners small but definite. Cara-
pace large and tumid, surface uneven with conical protuberances dorsally
crowded, laterally in 2 longitudinal rows, the lower forming an almost contin-
uous carina, both connected in front by a transverse prominence continued
on the side of the pseudorostral projection. Pedigerous segments 1, 2, raised
to thin lamellae, 34 to 5™ of these and first 4 pleon segments each with
dorsal pair of adjacent tubercles and sides acutely produced. Kye imper-
fectly developed. Maxilla 1, palp with 2 filaments. Peraeopod 2, 2"¢ joint much
more than half as long as the rest of the joints combined, 7‘ joint subequal
to the 5t, Pedunele of uropods coarsely serrate on both margins, about
two and a half times as long as the endopod. Colour dark-red. L. © about
6 mm, CO nearly 7 mm.
Norway, north to Lofoten Islands, depths 188—565 m; ?Japan, depth 1107 m.
17. C. rostrata Calm. 1905 C.r., Calman in: Sci. Invest. Fish. Ireland, 1904
app. 1 p.10,85 t.2 f.35—38 | 1910 C. r., Calman in: Bull. Mus. Paris, p. 180.
Closely resembling C. horrida (nr. 16) but pseudorostral lobes strongly
curved downwards to the acute apex, meeting in front of the very small
eyeless eyelobe for a distance thrice its length; the dorsal surface of the
carapace with only few tubercles; the 5 pleon segment carrying a pair of
dorsal tubercles; peraeopod 2 with the 2™¢ joint only half as long as the
rest of the joints combined, the 5" joint little longer than the 4%, the
7th ag long as 4t, 5%, and 6 combined. L., Q with rudimentary oostegites,
5°25 mm.
W. coast of Ireland, depth 570 m; lat. 35° 24‘ 45’ N., long. 10° 19‘ 7” W.,
1205 m; coast of Sudan, 1056 m.
Cumacea: 26. Campylaspididae, 1. Campylaspis 197
18. C. maculata Zimmer 1907 C. m., Zimmer in: Zool. Anz., v.31 p.371 | 1909
C.m., Zimmer. in; Ergeb. Schwed. Siidp.-Exp., v6 nr.3 p.13 t.5 £.69—83.
Not unlike C. horrida (nr. 16) and nearly allied to C. nodulosa (nr. 22).
Pseudorostral lobes produced to a distance not longer than the tuberculiform,
apparently eyeless eyelobe; a sinus well developed. The carapace more than
half the length of the body, beset with numerous tubercles, combined chain-
wise in three lateral lines, 2 close together and parallel to the lower margin,
the 34 at a somewhat greater distance. Pedigerous segments 1, 2, are not
raised dorsally. The pleon segments are without tubercles. Upper lip,
mandibles and other mouth-organs as in C. nodulosa, Maxilliped 1, terminal
joint comparatively large, piriform. Maxilliped 2, terminal joint with 3
apical spines. Maxilliped 3, 4th joint not very broad or very long, lateral
margin produced to a tooth. Peraeopod 1, 2"4 joint longer than the rest com-
bined, 7'* about two-thirds as long as the 5 or 6, Peraeopod 2 as in
C. Nonna. Peraeopod 5 small, fully developed. Peduncle of uropods with
margins weakly serrate, rather more than twice as long as the endopod,
which is a little longer than the exopod. Colour yellowish, spotted freely
with red-brown. L. about 5 mm.
South Georgia, depth 75 m.
19. C. affinis O. Sars 1870 C. a., G.O. Sars in: Forh. Selsk. Christian., 1869
p. 160 | 1900 C.a., G. O. Sars, Crust. Norway, v.3 p.91 t.64 | 1900 C. a., Zimmer in:
Fauna arctica, v.1 p. 429, 435 | 1912 C.a.?, Calman in: P. U.S. Mus., v.41 p. 628.
Named for affinity to C. rubicunda (nr. 7), but with some striking
differences. Pseudorostral lobes meeting for a short space in front of the
oval eyelobe, obtuse at apex, descending to a small sinus and feebly marked
antero-lateral corner. Carapace not half the length of the body, moderately
inflated, with a few low dorsal tubercles anteriorly. Pedigerous segments
1, 2, slightly elevated dorsally, following segments smooth, except that in the
G pleon segments 1—5 have the usual groove for the long 2"¢ antenna.
Maxilla 1 with 2 filaments to the palp as in C. verrucosa (nr. 21), notin C. rubieunda.
Maxilliped 1, terminal joint less rudimentary than usual, oval fusiform, mar-
gin ciliated, apex with 2 unequal spines. Maxilliped 2 with distally widened
seta at apex of medial margin of 2™¢ joint, 6% of quite moderate dimensions,
7% carrying 3 spines, the intermediate the smallest. Peraeopod 2, 7‘ joint
longer than 2 preceding joints combined. Uropods rather slender, peduncle
with unserrated margins, about twice the length of the endopod in the o,
but less than that in the 3; endopod in the © only just longer than the
exopod, but in the GO considerably longer, with spines on medial margin
numerous. Colour pale red, more or less fleeting in spirit. L. © 55 mm,
Oo 6 mm.
Norway, Lofoten Islands, depths 377—471 m; ?U.S. A., off Martha’s Vineyard,
depths 65—70 m.
20. C. frigida H. J. Hansen 1908 C.f., H. J. Hansen in: Résult. Voy. Belgica,
Schiz. Cum. p.16 t.3 f.2a—2n.
Near to C. verrucosa (nr. 21) and C. antaretica (nr. 23). Integument
very thin and flexible. Pseudorostral lobes meeting for a short space in
front of the prominent, parallel-sided, apparently sightless eyelobe, obtuse at
apex, descending to a small sinus and feeble antero-lateral corner. Carapace
covered with unequal tubercles seemingly in symmetrical arrangement.
198 Cumacea: 26. Campylaspididae, 1. Campylaspis
Pleon segment 5 with conspicuous transverse impression behind the middle.
Antenna 1, 24 joint shorter than 18%, 34 than 24, flagellum of 4 joints,
4th twice as long as the very short 1st, 34 longer than the long 2™4, accessory
a small oblong joint. Maxilliped 3 with 4% joint abnormal, triangular, so
strongly expanded on medial side that it is as broad as long, most of the
very long distal margin concave and free, the 5 joint attached to its lateral
apex. Peraeopod 1, 4t» joint little longer than 5%. Peraeopod 2, 7 joint
longer than 5%, Peduncle of uropods twice as long as the endopod, which
is slightly longer than the exopod. L., © with brood, 6 mm. ¢ unknown.
Lat. 70° 48‘ S., long. 91° 54’ W., a single specimen.
21. C. verrucosa O. Sars 1866 C.v., G. O. Sars in: Nyt Mag. Naturv., v.15
p- 105 | 1894 C. v., A. M. Norman in: Ann. nat. Hist., ser.6 v.13 p. 278 t. 12 f.8 | 1900
C. v., G. O. Sars, Crust. Norway, v.3 p.90 t. 63 | 1900 C.v., Zimmer in: Fauna arctica,
v.1 p. 430,435 | 1905 C. v., Calman in: Sci. Invest. Fish. Ireland, 1904 app. 1 p. 35 |
1906 C. v., Calman in: Mt. Stat. Neapel, v.171v p. 424.
Pseudorostral lobes shortly and obtusely produced in front of the ob-
long oval eyelobe, sinus and antero-lateral corner well marked. Carapace tumid,
covered with tubercles more or less symmetrically arranged. Pedigerous
segments and pleon segments as in C. horrida (nr. 16). The imperfectly |
developed eye, antenna and mouth-organs also agree with the species
just named, but the 7** joint of peraeopod 2 is comparatively narrower. Peduncle
of uropods less coarsely serrate, not nearly twice as long as the endopod.
Colour whitish with greenish tinge (G. O. Sars), all speckled with deep brown
or black (A. M. Norman). L. Q 5°5 mm, dO 6°5 mm.
Norway, depths 113—565 m; W. coast of Ireland, depth 219—570 m; Mediter-
ranean, near Capri, depth 200—1100 m.
22. C. nodulosa O. Sars 1887 C. n., G. O. Sars in: Rep. Voy. Challenger,
DAS eparsioo apo, GSmtenlle
Integument strongly indurated. Pseudorostral lobes produced obtusely
in front of the rather large oval but apparently eyeless eyelobe, sinus and
antero-lateral corner well marked. Carapace very large, not overhanging
the pedigerous segments, scarcely half the length of the body, without furrows
or carinae but more or less symmetrically covered with spaced tubercles.
Pedigerous segments somewhat raised dorsally. Pleon segments separated
by deep constrictions, lightly carinate laterally without dorsal teeth, the 5%
with transverse furrow behind the middle. Antenna 1, 2™4 joint shorter
than 15*, 34 than 24, flagellum of 3 joints, the 3¢ minute, the minute ac-
cessory 2-jointed. Upper lip broadly quadrate, emarginate behind, projecting
in front between the 2-jointed 2"¢ antennae as a rounded protuberance.
Mandibles with long narrow molar, left member with accessory cutting plate
and 4 spines, right with 6 spines and no accessory plate. Maxilla 1, with
palp long, carrying 2 filaments. Maxilliped 1, with 9 branchial leaflets on the
epipod. Maxilliped 2, with 3 spines on the terminal joint, the intermediate the
smallest. Maxilliped 3, 4 joint with lateral, margin produced to a tooth,
medial serrate, perhaps in this respect differing from C. maculata (nr. 18).
Peraeopod 1, 2"4 joint not longer than the rest combined, 7‘ about two-
thirds as long as 5 or 6t, Peraeopod 2, 4% and 5% joints each with
bidentate apex (wanting in C. horrida (nr. 16) and C. maculata (nr. 18), 7
joint equal in length to the 5, Peraeopod 5 wanting, probably only as a .
juvenile character. Peduncle of uropods slightly serrate on both margins;
Cumacea: 26. Campylaspididae, 1. Campylaspis 199
endopod three-fourths as long as the peduncle, exopod a little shorter. L.
nearly 5 mm, adult unknown.
Kerguelen Island, depth 232 m.
23. C. antarctica Calm. 1907 C. verrucosa var. a., Calman in: Nat. antarct.
Exp.. v.2 Crust. Cum. p.5 t. f. 14—16, tf.4 | 1907 C. v. var. a., Zimmer in: Zool. Anz.,
v.31 p.370.
As compared with C. verrucosa (nr. 21), hairs on surface much longer,
sides of carapace more flattened or sulcate over an oblique area as in
C. sulcata (nr. 15), dorsal tubercles of pleon segments 1—5 replaced by
sharp teeth pointing backwards, with a decided instead of faintly indicated
pair of dorsal teeth on each segment, 5% segment with post-median con-
striction more strongly marked, hind margin dorsally extended into a curved
median tooth serrate on the upper margin, peduncle of uropods more strongly
serrate, the teeth on medial margin spiniform, curved, irregular in size; medial
margin of endopod more distinctly serrated. L. adult 9 5°35 mm.
Winter quarters of the British Expedition, and of the German South-pole Exp.,
lat. 66° 2‘S., long. 89° 38’ E., depth 385 m.
Genera Cumaceorum incertae sedis
Gen. Calyptoceros Huxley
1856 Calyptoceros, Huxley in: Quart. J. geol. Soe., v.121 p. 34.
Evidently having a distinct telson, and, judging by the name, probably an adult
male with long antennae.
British Channel.
Gen. Olbia Marcusen
1867 Olbia (non Stal, Hemiptera, 1862), Marcusen in: Arch, Naturg., v. 331 p. 359
(deser. nulla).
Black Sea.
Alphabetical Index
abbreviata (Diastylis)
abbreviata (Pseudocuma)
abbreviatus (Diastylis) . -
abbreviatus (Ekdiastylis) . -
abbreviatus (Schizorhynchus) . -
abyssicola (Cumella)
abyssicola (Spencebatea)
abyssi (KHudorella)
acanthodes (Adiastylis) .
acutirostris (eucon) . .
Adiastylis
affine (Hocuma) .-
affinis (Campylaspis)
affinis (Hocuma)
africana (Iphinoé)
africanum (Sympodomma)
atricanus (Sympodomma)
agilis (Cumella) ,
agnatus (Nannastacus) -
akaroénsis (Heteroleucon)
alaskensis (Diastylis) .
Alauna ..
algoae @iaaivtis) Sa asks
ambigua (Diastylis) .
ampulacea (Leptostylis) .
ampullacea (Cuma)
ampullacea (Diastylis). ... -
ampullacea (Leptostylis) .
ampullaceus pee ean
Anchicolurus .
anderssoni (Diastylis) .
angulata (Cuma)
annulata Diastylopsis)
annulata (Leptostylis) .
anomala (Diastylis) .
anomala (Vaunthompsonia)
anomalum (Sympodomma)
anomalus (Diastylis)
anomalus (Leucon) .
anomalus (Sympodomma)
antarctica (Campylaspis)
antaretica (Campylaspis verrucosa var.)
antarctica (Leucon) . .
antarcticus (Leucon)
antillensis (Diastylis) .
antipus (Leptostylis)
Pag.
138
144
138
arabica (Vaunthompsonia)
arabica (Vauntompsonia)
arenosa (Bodothria) . .
arenosa (Bodotria) ...
arenosa (Cuma)
arenosus (Bodotria) . .
argentata (Diastylis)
argentatus (Ekdiastylis). .
argus (Cyclaspis) .
armata (Diastylis)
armata (Procampylaspis) .
armatus (Diastylis) .
armatus (Procampylaspis)
aspera (Diastylis) .
| aspera (Paralamprops)
asper (Diastylis)
asper (Paralamprops)
assimilis (Hemilamprops) .
assimilis (Leucon)
Astacus
Astaricilla . .
Asteracilla .
audouinii (Bodotria) ....
audouinil (Cuma) .«
australis (Bodotria) .
australis (Cumella)
australis (Cyclaspis)
australis (Leucon)
Bathyecuma .
Bathylamprops .
bella (Cuma) . :
beringi (aroronee
bicornis (Diastylis)
bicristata (Gynodiastylis)
bicristatus (Gynodiastylis)
bidentata (Diastylis)
bidentatus (Diastylis) .
bifrons (Schizotrema) -
bilamellata (Pseudocuma)
bilamellatus (Schivothynchusy
bimarginatus (Diastylis) .
binoculoides (Nannastacus)
binoculoides (Nannasticus) .
biplicata (Cyclaspis)
biplicata (Diastylis) .
biplicata (Diastyloides)
biplicata (EHudorellopsis)
biplicatus (Diastylis)
biplicatus (Diastyloides) ©
bispinosa (Cuma) .
bispinosa (Diastylis)
bispinosus (Diastylis) .
bispinosus (Diastylis)
bistriata (Cyclaspis)
bistriata (Pseudocuma)
Bodothria .
Bodotria ‘
Bodotriidae :
bonnieri (eeosarine napisy
borealis (Diastylis)
borealis (Leptostylis) .
brachydactylus (Nannastacus)
Brachydiastylis :
brachyura (Paradiastylis)
brachyura (Platyaspis) .
brachyurus (Bavadiaatylia)
brachyurus (Platysympus)
bradi (Diastylis) .- 2... -.
bradyi (Diastylis). ...
brevicaudata (Leptostylis) .
brevicaudata (Paradiastylis)
brevicaudatus (Nannastacus)
brevicaudatus (Paradiastylis)
brevipes (Iphinoé)
brevirostre (Bathycuma)
brevirostris (Bathyeuma) .
brevirostris (Bathycuma uaun ore:
SCALE) ee a ieee hove)? fois tw cen «>
brevirostris (Cuma) .
brevirostris (Leucon)
caeca (Vaunthompsonia) .
ealmani Bathylamprops)
ealmani (Dic) .
ealmani (Schizotrema)
ealveri (Diastylis) .
Calyptoceros
Campylaspidae
Campylaspididae
Campylaspis
campylaspoides (Caspiobuniays.
campylaspoides (Pseudocuma)
Cancer :
Cancer (Gaenanaye
Cancer (Oniscus) .
carinata (China)
earinata (Cumella)
earinata (Cumellopsis). . .
earinata (Gynodiastylis) .
earinatus (Gynodiastylis)
Caspiocuma
Ceratocuniay ts) a) eh cee ve os
Ceratocumatidae
Ceratocumidae ....
Bor ole
Alphabetical Register
Pag.
84
88
87
95
95, 96
95
96
39
150
23
23
18
185
101
125
172
107
62
121
62
Heyer s/s)
99
121
121
174
121
cerearia (Cuma)
cercaria (Leucon). .-- = -
cercaria (Pseudocuma) ‘
cerecaroides (Pseudocuma) .
Chalarostylidae .
Chalarostylis ..-..-: -
ciliata (Cyrianassa)
ciliata (Psendocuma)
ciliatum (Pseudocuma)
ciliatus (Pseudocuma) .
cingulata (Cyclaspis)
cingulata (Diastylis) .. .
cingulatus (Diastylis)
eingulatus (Makrokylindrus) .
clavicauda (Cumella)
Colurostylidae.
Colurostylis
Colurostylis ;
ecomata (Lamprops) .
comes (Hemileucon) 5
compressa (Procampylaspis)
compressus (Procampylaspis) .
coneava (Bodotria)
Condylura . .
Condylurus . :
cornigera (Cyclaspis)
cornigera (Cyclaspoides) .
cornuta (Cuma)
cornuta (Diastylis)
cornutus (Diastylis) .
| eostata (Campylaspis) .
costata (Cyclaspis)
costata (Diastylis)
costata (Gynodiastylis)
costatus (Adiastylis)
costatus (Gynodiastylis)
crassicauda (Leptostylis)
| erassior (Diastylopsis)
erassipes (Iphinoé) .....
| eristata (Hemilamprops) .
| eristata (Hemilamprops)
7 | cristata (Lamprops) .
eristata (Vaunthompsonia)
cristata (Vauntompsonia)
Cuma .
Cumacea
Cumadae.
Cumnrelllans .-4 eal oon
Cumellidae
Cumellopsis
Cumidae..
Cumidea,.
Cumopsis . Secale eee UE:
ONG Ss GG Gn 6 oo 8%
Cyclapsis
| Cyclaspis .
Cyclaspoides
me Or OL Ot
aN om
ie
202
Pag.
Cyma . 208
Cymatopsis Ao 208
Cyrianassa . : 42,74, 148
dalli (Diastylis) - Saber Ror heey Monee eel:
dawsoni (Diastylopsis) 110
declivis (Petalomera) 142
declivis (Petalopus) . 142
declivis (Petalosarsia) 142
deformis(Hudora)) is 3. sare 0. deny 1
deformis (Hudorella) ....... 82
deformis (EHudorellopsis) .... 82
deformis (Leucon) . . aie; SA mint)
dentifrons @bisstylopeis)) 114
dentifrons (Leptostylis) ... . 114
depressum (Schizotrema) 166
Diastylidae ere 1,18
Dastiy idiom hem mon ae oS
Diastylis . a ces 1, 160
Diastylis G2) avy co 8 ee Aken aoe
Diastyloides .... J £3508 286
diastyloides (rcnloceneye § 147
diastyloides (Stenocuma) ... . 147
Diastylotdida@e <i. 20 tele 180
Diastylopsish (ii-8 -.« e- 109
Dic. A Pou ha ats 160
Dicidae . . ess ce a ee LOO
DrcHadiopiamen. o- INES Btn te 1
diomedeae (Eeteraenma) Aros Le intl) wuTh
diomedeae (Sympodomma) ... 17
Diops : Sa eed ai iberml vl ehy
dollfusi CRacuniayhe idch bebrolitet tedind 40)
dubia (Diastylopsis) . rst ese WLS
dubius (Malcvolkeylinarus) hs Poaces ALLS
echinata (Diastylis)’\.\/2-i\<4 «on LOS
echinata (Procampylaspis) . . .« 186
echinatus (Diastylis) Sige ce, 1lOy
edwardsii (Cuma). . ... . 25. 50,102
edwardsii (Diastylis) 102
edwardsii (Vaunthomsonia} .... 50
Ekdiastylidae 135
Ekdiastylis . 135
Ekleptostylis . ii ot wee eo
elegans (Ghallimoatylia) Pay Hae ene Bo
elezans) (CyClaspis)ie. en -inruee) ee OO
elegans (Cyrianassa) “2 26% 216 =) hOO
elongata (Bathycuma). ...... 14
elongata (Diastylopsis) eiomron ale):
elongatum (Bathycuma)..... 14
elongatus (Bathycuma) ...... 14
emarginata (Hudora) << ...9..2 . 9/75
emarginata (Budorella) ..... 7d
emarginatus (Leucon)......-+ 75
ensifer See ai ok leecietealte CML OIG
EKocuma . . Se ete ee eG
erinaceus (aera : 120
erinaceus (Makrokylindrus A 120
esca .(Gammaris)) 20 wemsiieni as moll:
Alphabetical Register
Pag.
Kudora 5: [TS oe aCmn sag 74
Hudorella. ... °: A eters te.
eudorelloides Childers) 143
eudorelloides (Schizorhynchus) 143
Eudorellopsis . oi, Pah pee 82
exsculpta (erclasnis) z 35
fallax (BHudorella) . 77
fasciata (Cuma) . 51
fasciata (Lamprops) - 155
fasciata (Lampros) on - . 155
ferox. (Bodotria) = 39.3 = 2 arp
ferox (Cyclaspoides) . 23
ferox (Hocuma) ... Jo si Walcot ome
ferox (Paoudodinstylis) sito l eh oeeeliso:
fimbriata (Diastylis) : 139
fimbriatus (Ekdiastylis) . 139
folini (Cuma) . 5 ail
forficula (Cumella) ; 180
fragilis (Makrokylindrus) . eli
frigida (Campylaspis) ..... 197
fulvus (Leucon). 2 66
fuscata (Lamprops) ....... 154
fusciata (Cuma) 51
Gammariss ps scene ces 1
Gaussicuma 11
gayi (Cuma) . tiie De hopkes tems 140
fay (ELOLOStyIS) pie.) Cente 140
georgi (Nannastacus) . 173
gibba (Bodotria) .- 26
gibba (Cuma) woh fone 26
gibbosus CNcnnaatacney a Ne oahe 170
gibbus (Bodotria) . . 26
gigantea (Cuma) 99
gigas (Cyclaspis) Byo. to 38
glabra (Campylaspis) ... .. . 191
glabra (Diastylis rathkii var.) ... 98
glacialis (Cyclaspis). ..... - 380
goésii Scene rae y ; 40
Goniacis. . . i ish Ea pale OO
goodsir (Bodothea) = see. «mete
goodsir (Cumopsis) on aD)
goodsiri (Alauna). ...... - 98,99
goodsir: (Cuma); =. 9 = \-) 1 euneureeee
goodsiri (Cumopsis) - ‘ 50
goodsiri (Diastylis) ....... 99
goodsirii (Bodotria) . -. 50
gracile (Stenocuma) . 148
gracilior (Hudorella) ... . Ue
gracilis (Cyrianassa) .... - 46
gracilis (Hudorella) .... . 79
gracilis (Iphinoé) . 46
gracilis (Leptostylis) 128
gracilis (Pseudocuma) . 148
gracilis (Stenocuma) 148
gracilis var. maeotica (iaiinoayt 46
gracilis var. serrata (Iphinoé) 45
SGITE (Venilia) . - 46
Alphabetical Register 203
Pag. Pag.
gracillima (Cumella) .... . . 179 | josephinae (Diastylis) ... +. - 120
gracilloides (Pseudocuma) . . . . 148 | josephinae (Makrokylindrus) . . 120
graciloides (Pseudocuma) ... . - 148 | kalluropus (Leucon).....-: - 71
graciloides (Stenocuma). ... . 148 | kerguelensis (Leucon)....-.- - 72
granulata (Heterocuma).... .- - 49 | kinbergii (Leptocuma). ..... 53
granulata (Heterocuma sarsi var.) . 49 | kinbergi (Leptocuma)..+-..+-+-s 53
granulatum (Heterocuma). ... 49 | koreana (Diastylis)ueet te bie tee) noe
Gynodiastylidae ..... . 161 | koreanus (Diastylis)......-- 93
Gynodiastylis. . =): .... . .s. 161 | laevis (Cumella); = = -).'. > 182
lalate. Glo eee 1400\ > laevis; (Cumiopsis)): | ae: 4) io ee OO
hammoniae fpinseaiicy Po e111 930) lacvise (Oyclaspis)\i.geamets tite as cjieeros
hanseni (Nannastacus) .... .-. 172 | laevis (Diastylis) ..-...---:- 97
helgae (Cumellopsis) . ... ..- 178 | laevis (Gynodiastylis). ...-.- 161
helleri (Diastylis). . . ....- ..- 140 | lagunae (Pseudocuma) ...-. .- 151
helleri (Holostylis) . . .... . 140 | lamellata (Diastylis). ...-+-.. 87
Hemilampropidae....... 54 | Tampropidae...-.-...-.- 153
Hemilamprops ......... 55 | Lamprops . SIN ec Satan eS:
eriileucon ea eee eh taae) ee, COUP Wa lamprose. sells avis c=) fer seem eta pls
Hemileuconidae...... .- 152 | lata (Kocuma) .. . ect rie 22"
herdmani See a eee es oe) | latiaculeata (Pecudocara) OM en eel
Heterocuma ... ea ees ool latin (MOCUINE er top is cota nromere
ieterocyma, - |. - . -%. = « . - - 208 | Leptocuma . . - Ee Katsoreh ge MOS
eteroleucom.) +... |. - - 164 Teptooimatiane AMER EGOS cri OLS
Heteroleuconidae ...... . - 163 | Leptocyma.. . 5s ene a ED
heterostylis (aeucon) ...... 70 | leptopus (Gamelan) sie oaiae c 18]
hexaceros (Diastylis) ..... . . 137 | Leptostylis... .- oe eet 2d
hexaceros (Ekdiastylis) . ... . 137 | lepturus (Nem ac ore) mea an WE!
hilgendorfi (focuma) ......- 21 | leuckartii (Leucon).....--. 74
irentam(Budona)ie. & 2. et ieiea a eee G0 | WueuGome.. ..6 06 ene mt on 08
intesuta: (Gudorella) sai 6 ew) CON ueucon. =.) 2 22s certs be A SS
hirsutus @Nannastacus) .. . <=. 173 | Ibeuconidae......-:. «- «> 62
Inspida, (Cumella). =. 2... ..)- 181 "| Leuconopsis - . -... 2. + sss 63
hispida (Eudorella) . ...... 79 | levis (Cyclaspis) .......-.- 32
HMollostyludae . . ss 6. 2's 139) limicola,(Cumella).. >. = «)-) 183
Holostylis ......... .. 189 | longicaudata (Bathycuma)..... 12
holti (Platyeuma) . .... .. . 176 | longicaudata (Cyclapis) ..-.-..- 30
hornelli (Cyclaspis)...... . 382 | longicaudata (Cyclaspis) .... 30
horrida (Campylaspis) ... .. 196 | longicaudata (Diastylisy . .... . 115
horrida (Ceratocuma) . . ... .. 52 | longicaudata (Leptostylis) . - . . . 115
horrida (Diastylis) . .... .. . 1838 | longicaudatum (Bathyeuma)... 12
horridum Cas ea. .... . 52 | longicaudatus (Adiastylis)... .- 115
horridus (Ceratocuma). . . . .. . 52 | longicaudatus (Bathyeuma). .... 12
horridus MBkaiastylis) .... . 138 | longicorne (Hocuma) ...... 2l
Thpinoé PELs. ...... 45 | longicorne (Pseudocuma) .... 150
inermis (Hadorella)) See se) 2) 80) | longicornis (Cyrianassa). = - - - «1 150
inermis (Iphinoé)...... .-. . 48 | longicurnis (Hocuma) .....-.. 21
inermis (Vaunthompsonia) .. . 9 | longicornis (Pseudocuma) . ... .- 150
insignis (Diastylis) . ... . . . . 186 | longimana (Leptostylis) ..... . 127
insignis (Ekdiastylis) .... . . 136 | longimanus (Diastylis). . . ... + 127
insularum (Diastylis) . .... . . 114 | longimanus (Leptostylis) . . . . 127
insularum (Diastylopsis) ... . 114 | longipes (Adiastylis) ...... 116
insularum (Leptostylis) . . .. . . 114 | longipes (Bodotria)........- 10
integra (Kudorella) . ee ea Son || loneipes (Cuma). cua 1s!)
integra udoreliopsis) .... . 83 | longipes (Cuma (Bodotria)), ree Fed OO
iiphinoesy. .) 5.1 Mee stele. 48°.|loneipes, (Cumopsis) 3 2227) 5). e050
Epiinocwe sete eenig rs.) «ts .. 42; || longipes! (Cyclaspis) = .:15°5 >. in ei ndt
plithochwewemees «else. 28 2a, 43°)| longipes,(Diastylis)) = ie) 2°!) <...9 ab
204 Alphabetical Register
Pag.
longipes (Paradiastylis)..... 122
longirostre (Bathycuma) .... 13
longirostris (Bathyeuma) ..... 13
longirostris (Bathycuma (Vauntomp-
SONIA) esha ts, ote et ely eee eS
longirostris (Leucon) ...... 70
longirostris (Nannastacus) ... 171
lacifera. (Cama)! ) ten 2) < i) eae meee Ol
lacifera:(Diastylis)i: c.. ta) -tye eee y LOL
fucifer<C(Diastyilis) semaine LO)!
luciferus (Diastylis) . a haghemesst nar LO
macrobrachium deine) ‘eee es 4s
macrophthalma (Gampylaspiayy’ 5 JIGS
macrura (Leptostylis) . ..--.. 126
macruroides (Leptostylis) . . . . 126
macrurus (Leptostylis) ... .. 126
maculata (Campylaspis). .... 197
maeotica (Iphinoé) ......-+.-. 46
maeotica (Iphinoé gracilis var.) . . 46
Makrokylingrius) fap si. nen emote a kes
manca (Diastylis). . =. . 22+. 7. 123
manca (Leptostylis). ....-.. . 128
mancus (Leptostylis) . .... . 123
marginata (Hudorella). .....- 75
mediterraneus (Leucon)..... 69
meridionalis (Vaunthompsonia) . w)
meridionalis (Vauntompsonia) .. . )
meridionalis (Vauntrompsonia) . . . 9
minor (Leptocuma)... .-...% . 954
minor (Nannastacus) ...... I71
minus @s.eptocuma) .. =. .-- » « 04
molossa (Cumella) ....... 180
monodon (Mudorella) ..... . 78
montacui, (Bodotria) ) 0.) le ie 20
mortenseni (Zygosiphon) .... 41
mystacina (Diastylis) ....... 137
mystacinus (Ekdiastylis) .... 137
macica <CeucOM)) par katve Heh kersed)s tccte Ld
nana (@iudorella) 7s 2 5). 2. ve
Nanaia sia @udiaienn ctrl re eee lO
Infaabikeyerauey 6 co5 6 5 4g 6 6 0 4 Js)
INannastacus} es cp sts) laid re feleiemme ns udt
Nannasticus ...... ¢ aL 174
MES, (OM) 6 56 ob oo ooo Oe
kale (AiO) 4 5 66 65 06 0 We
nasicoides (Leucon)....... 65
nasicoides (Leucon)... « - 67, 68
nasicoidis (Leucon) . ...... 65
nasicus)|(eucan)) =f. 2). a a-0- 2 OO
natalense (Bathyeuma) ..... 14
natalensis (Bathycuma) ...... .14
nathorsti Gseucon) . =... . 68
neapolitana (Diastylis) ...... 104
neapolitanus (Diastylis). ... . 104
neoseelandiae (Diastylopsis) . . - . LILO
neozealanica (Diastylis) ..... . 110
neozealanica (Diastylopsis) . . . 110
nitens (Campylaspis)
nodosa (Diastylis)
nodosus (Diastylis)
nodulosa (Campylaspis). . .
normani (Hemilamprops) .
nucella (Diastylis)
| occidentalis (Anchicolurus) .°.
occidentalis (Colurostylis)
Olbia .
Oniscus
orbicularis (Par member ent
orbicularis (Platyaspis)
orbicularis (Platytyphlops)
orbignyi (Diastylis) . .
orientalis (Campylaspis)
ossiani (Nannastacus)
ovalis (Campylaspis)
Oxyurostylidae
Oxyurostylis
Pachystylis....
pacifica (Cammy lannisyl
paeneglaber (Campylaspis) .
paeneglabra (Campylaspis)
pallidus (Leucon) .
Paradiastylis . 5
Paralampropidae
Paralamprops
Paraleucon .
Paraleuconidae
Paranannastacus
pardus (Nannastacus) .
parva (Bodotria)
parva (Cuma).
parvulus (Diops)
parvulus (Diops (Nannastncas))
parvus (Bodotria) ;
pectinata (Pseudocuma) .
pectinata (Pterocuma). .
pectinatum (Pterocuma)
pellucida (Hemilamprops)
pellucidus (Hemilamprops)
peringueyi (Platytyphlops)
persculpta pkey?
Petalomera eee
Petalopus
Petalosarsia
picta (Cyclaspis)
picta (Diastylis) :
planifrons (Diastylis) .
Platyaspidae
Platyaspis .
Platycuma :
Pia tyayanpodaiter
Platysympus
Platytyphlops
platyuropus (Campylaspis)
plumosa (Alauna) . 2 ite
polaris (Diastylis)
polaris (Diastylis) .
politus (Diastylis)
politus (Bkdiastylis)
ponctata (Cuma)
Procampylaspidae.
Procampylaspididae
Procampylaspis . ;
producta (Hocuma) .
producta (Leptostylis) .
productus (Leptostylis) .-
Pseudocuma
pseudocuma (Colteontyiisy
Pseudocumatidae
Pseudocumidae
Pseudodiastylidae
Pseudodiastylis .
Pseudoleucon .
Pterocuma
pulchella (Bedottiay:
pulehella (Campylaspis) .
pulchella (Cuma) . ae
pulchellus (Bodotria) .
pulex (Bodotria)
pulex (Cuma)
punetata (Cuma)
puritani (Cumellopsis)
pusilla (Cuma) .
pusilla (Cyclaspis)
pusilla (EHudorella)
pygmaea (Cumella) .
quadriplicata (Diastylis)
quadriplicata (Lamprops) . .
quadrispinosa (Diastylis) .
quadrispinosus (Diastylis)
quadrituberculata (Cyclaspis) .
qvadrispinosa (Diastylis) .
rathkei (Cuma) .
rathkei (Diastylis)
rathkii (Cuma) .
rathkii (Diastylis)
rathkii var. glabra Gaiety)
rathkii var. sarsi (Diastylis) .
reptans (Nannastacus). .. .
reptans (Paranannastacus)
resima (Cuma) .
resima (Diastylis) .
resima (Diastylopsis)
resima (Budorellopsis)
resimus (Brachydiastylis) .
resimus (Diastylis)
resimus (Kudorellopsis)
robusta (Diastylopsis)
robusta (Iphinoé) .
robusta (Leptostylis)
rosea (Hemilamprops)
rosea (Lamprops). .
Alphabetical Register
Pag.
100
100
136
136
51
184
184
184
21
127
127
148
130
140
140
133
133
85
145
24
190
24
24
26
26
51
178
27
33
81
182
107
155
95
95
38
95
98
98
98
205
Pag.
rosea (Vaunthompsonia) . 5 tee)
rostrata (Alauna) . 98, 39
rostrata (Came leans) 196
rostrata (Diastylis) 97
rostrata (Pseudocuma). . 145
rostrata (Pterocuma) 145
rostratum (Pterocuma) 145
rotundata (Pachystylis) 131
rotundatus (Pachystylis) 131
rubicunda (Campylaspis) 190
rubicunda (Cuma) 190
rufa (Cuma) : 51
rugosa (Diastylis) . 102
rugosoides (Diastylis) . 101
rugosus (Diastylis) 102
sagitta (Leucon) 73
sarsi (Cyclaspoides) : 18
sarsi (Diastylis rathkii var.) . 98
sarsi (Heterocuma) . 48
sarsii (Cyclaspis) 22
sarsii (Cyclaspoides) 18
sarsii (Hocuma) . 22
sarsii (Heterocuma) . 48
sarsii (Nannastacus) . 176
sarsi var. granulata (Heterhenmay 49
scabriuscula (Pseudocuma) . . 143
seabriusculus (Schizorhynehus) 143
Schizorhynehus... . A rG: J 143
Schizotrema : 165
scorpioides (stacusy 25
scorpioides (Bodotria) . AHS
scorpioides (Cancer) 25
scorpioides (Cuma) . F 25
scorpioides (Cuma) 25, 27, 28
scorpioides (Diastylis) . 25
scorpioides (Diastylis) 102
scorpioides (Oniscus) 102
scorpionides (Diastylis) 25
Scorpionura 89
scorpoides (Cuma) 25
sculpta (Diastylis) 135
seculptus (Diastylis) . 135
sculptus (Ekdiastylis). . 135
septemdentatus (Leucon) 69
serrata (Cumella) . 182
serrata (Diastylis) 86
serrata (Diastyloides) . 86
serrata (Iphinoé) ; 45
serrata (Jphinvoé gracilis var.) 45
serrata (Iphithoe) . 45
serrata (Sphinoe) . 45, 46
serratocostata (Hemilamprops) . 59
serratocostata (Paralamprops) . 59
serratus (Diastylis) ......- +. 86
serratus (Diastyloides) 86
serratus (Leucon) . 72
serricauda (Diastylis) 17
206
serricauda (Makrokylindrus)
siamensis (Bodotria)
sibogae (Cyclaspis) .
simile (Pseudocuma)
similis (Bodotria) .-
similis (Cyclaspis)
similis (Kudorella)
similis (Pseudocuma)
siphonatus (Leucon)
smithi (Oxyurostylis) .
sordida (Budorella) .
sordidum (Schizotrema) .
sorex (Pseudoleucon) .
sowinskyi (Pseudocuma) .«
sowinskyi (Pterocuma)
spectabilis Ore epee
Spencebatea
Sphinoe . :
spinosa (Gamipeiacpis)
spinosa (Diastylis)
spinosus (Diops) i ise
spinosus (Nannastacus) .
spinulosa (Diastylis) :
spinulosa (Stenotyphlops) .
spinulosus (Diastylis) -
spinulosus (Stenotyphlops) .
splendida (Eudorella) .
stebbingi (Nannastacus)
stellifera (Hocuma) .«
stelliferum (Kocuma)
Stenocuma
Stenotyphlops .
Stephanomma.
Strauchia .
strigata (Dias)
stygia (Diastylis) .
stygius (Diastylis)
sublevis (Bodotria)
suhmii (Nannastacus) .
suhmi (Nannastacus)
suleata (Campylaspis) .
suleata (Diastylis). .
suleatus (Diastylis) .
suteri (Paraleucon) .
Sympoda
Sympodomma. . ;
Sympodommatidae
taprobanica (Kocuma) .
taprobanicum (Bocuma) .
tardus (Nannastacus) é
tardus (Paranannastacus) .
taurica (Strauchia)
tenella (Iphinoé)
tenuicauda (Pseudocuma)
tenuicauda (Stenocuma) .
tenuirostris (Leucon)
terginigra (Cuma) .
Pag.
117
27
30
149
27
35
80
149
64
132
81
167
85
146 |
146
31
178
43
192
98
176
176
94
157
94
157
80
176
21
21
147
156
40
151
102
100
100
25
170
170
168
168
151
147
147
Alphabetical Register
thileniusi (Diastylopsis) .
thileniusi (Leptostylis). -
thomsoni (Cyclaspis)
tricincta (Diastylis) .
tricincta (Leptostylis) .
tricinctus (Diastylis)
tridentata (Procampylaspis) .
triplicata (Cyclaspis)
trispinosa (Cuma) .
trispinosa (Halia) .
trispinosa (Iphinoé) .
trispinosa (Iphithoe)
truneatula (Hudora) .
truneatula (Kudorella) .
| truneatula (Budorella)
tubulicauda (Diastylis)
tubulicauda (Dic) .
tumida (Cuma) .
tumida (Diastylis)
tumida (Diastylopsis) .
tumidus (Diastylis) .
typica (Platyaspis)
typicus (Platysympus)
undata (Campylaspis) .
unguiculata (Cuma) .
unguiculata (Nannastacus) .
unguiculatus (Nannastacus) .
unguiculatus (Nannasticus) .
unicornis (Cyclaspis)
uniplicata (Cyclaspis) .
uniplicata (Hemilamprops)
uniplicata (Lamprops) . :
uniplicatus (Hemileucon) .
vanhoeffeni (Gaussicuma). .
vanhoeffeni (Leucon) .
varians (Cyclaspis) .
Vaunthompsonia
Vaunthompsondiiae
Vaunthompsoniidae
Vaunthomsonia .
Vauntompsonia . 2
Vauntompsoniidae...
Vauntrompsonia
Venilia
verrucosa (Caripulascian
Ol
verrucosa var. antarctica (Canpeieeor|
villosa (Leptostylis) .
villosus (ientacelin}
vitrea (Campylaspis)
vulgaris (Scorpionura). . .-
walkeri (Ekleptostylis) .
walkeri (Leptostylis)
weberi (Heterocuma)
weberi (Sympodomma)
zimmeri (Iphinoé)
zimmeri (Nannastacus)
Zygosiphon .
Nomenclator generum et subgenerum
Adiastylis T. Stebbing in: Ann. 5S. Afr. |
Sp.: A. |
Mus., v.10v -p.148. 1912.
acanthodes.
Alauna Goodsir in: Edinb. new phil. J.,
v. 34 p. 120, 127. 1843. Sp.: A. rostrata. |
Anchicolurus T. Stebbing in: Ann.S. |
LOT ep. |
Afr. Mus., v.10v p. 176.
A. occidentalis.
Astacus Borlase, Nat. Hist. Cornwall, |
p. 274. 1758. Sp.: A. verus.
Astaricilla Dalyell, Powers Creator, v. 1
p.224. 1851. Sp.: A. caudatae.
Asteracilla [pro: Astaricilla Dalyell 1851]
Dalyell, Powers Creator, v. 1 p.226. 1851.
Bathyecuma H. J. Hansen in: Ergeb.
Plankton-Exp., v.2G.c p. 455. 1895.
Sp.: B. elongata.
Bathylamprops Zimmer in: Ergeb.
Tiefsee-Exp., v. 8 p. 158,173. 1908.
~Sp.: B. calmani.
Bodothria [pro: Bodotria Goodsir 1843]
P. J. Beneden in: Mém. Ac. Belgique,
v. 33 [no. 3] p.73. 1861.
Bodotria Goodsir in: Edinb. new phil.
J., v. 34 p. 120, 128. 1843. Sp.: B.
arenosa.
Brachydiastylis T. Stebbing in: Ann.
S. Afr. Mus., » 10 v p.176. 1912. Sp.:
B. resimus.
Calyptoceros Huxley in: Quart. J. geol.
Soe., v.121 p.36. 1856.
Campylaspis G. O. Sars in: Forh. Selsk. |
Christian., p. 200. 1865. Sp.: C.
rubicunda, C. costata, C. undata.
Cancer Linné, Syst. Nat., ed. 10 p. 344,
625. 1758. Sp.: C. cursor, C. raninus,
C. mutus. C. minutus, C. ruricola, C.
vocans, C. craniolaris, C. philargius, C.
rhomboides, C. maculatus, C. pelagicus,
C. nucleus, C. lactatus, C. maenas, CV.
depurator, C. feriatus, C. granulatus,
C. pagurus, C. chabrus, C. araneus,
C. cuphaeus, C. muscosus, C. per-
sonatus, C. maja, C. longimanus, C.
horridus, C. cristatus, C. superciliosus,
C. cornutus, C. longipes, C. spinifer, C.
cruentatus, C. hepaticus, C. calappa, C.
grapsus, C. aeneus, C. punctatus, C.
dorsipes, C. symmysta. C. bernhardus,
C. diogenes, C. gammarus, C. astacus,
C. curcinus, C. pennaceus, C. squilla, C.
crangon, C. carabus, C. cancharus, C.
pilosus, C. norvegicus, C. homarus, C.
arctus, C. mantis, C. scyllarus, C. pulex,
C. locusta, C. salinus, C. stagnalis.
Caspiocuma G. O. Sars, Crust. Norway,
v.3 p. 73. 1900. Sp.: C. campylaspoides.
Ceratocuma Calman in: Sci. Invest.
Fish. Ireland, 1904 app. 1 p. 11, 37.
1905. Sp.: C. horrida.
Chalarostylis A. M. Norman in: Ann.
nat. Hist., ser. 5 v.3 p.65. 1879. Sp.:
C. elegans.
Colurostylis Calman in: Tr. zool. Soe.
London, v. 18 1v_ p. 342, 376. 1911.
Sp.: C. pseudocuma.
Condylura Illiger, Prodr. Mam. Av.,
p: 125. 18lle Spe Cas cristata, C:
longicaudata. Mamm.
Condylura Latreille in: G. Cuvier,
Régne an., ed. 2 vu 4 p. 153. 1829
Sp.: C. Dorbignyr.
Condylurus [pro: Condylura Latreille
1829] H. Milne- Edwards, Hist. nat.
Crust., v.38 p.553. 1840.
Cuma |Calonne}], Mus. Calonn., p.35. 1797.
Sp.: C. Aulica, C. Moro, C. Prismatica.
Moll.
208 Nomenclator generum et subgenerum
Milne-Edwards in: Ann.
1828. Sp.:
Cuma H.
Sci. Nat., v. 13 p. 294.
C. Audouwini.
O. Sars in:
p. 198. 1865.
Forh. Selsk.
Spe CG:
Cumella G.
Christian.,
pygmaea.
Cumellopsis Calman in: Sci. Invest.
Fish. [reland, 1904 app. 1 p. 11, 28.
1905. Sp.: C. Helgae.
Cumopsis G. O. Sars in: Arch. Naturv.
Kristian., v.3 p.465,511. 1878. Sp.:
C. Goodsiri.
Cyclapis [pro: Cyclaspis G. O. Sars 1865]
G. O. Sars in: Forh. Selsk. Christian.,
p. 206. 1865. Sp.: C. longicaudata.
Cyclapsis [pro: Cyclaspis G. O. Sars
1865] H. J. Hansen in: Ergeb.
Plankton-Exp., v.2G.c p.51. 1895.
Cyclaspis G. O. Sars in: Forh. Selsk.
Christian., p. 207. 1865. Sp.: C. longi-
caudata.
Cyclaspoides Bonnier in: Ann. Univ.
Lyon, v.26 p.530. 1896. Sp.: C. Sarsi.
Cyma pro: Cuma H. Milne Edwards 1828
L. Agassiz, Nomenel. zool., Index
p. 108,112. 1846.
Cymatopsis pro: Cumopsis G.O.Sars 1878
Rye in: Zool. Ree., v.17 Index p.4 1881.
Cyrianassa pro: Venilia Bate 1856 Bate
in: Ann. nat. Hist., ser. 2 v.18 p. 187.
1856.
Diastylis Say in: J. Ac. Philad., v lu
p. 313. 1818. Sp.: D. arenarius.
Diastyloides G. O. Sars, Crust. Norway,
v.3 p.60. 1900. Sp.: D. serrata, D.
biplicata.
Diastylopsis 8. I. Smith in: Rep. geol.
Surv. Canada, 1878—79 p.215 B. 1880.
Sp.: D. Dawsoni.
Die T.Stebbing in: Ann. S. Afr. Mus., |
v.61v p.415. 1910. Sp.: D. calmani.
Diops Paulson, Unters. Krebse rot.
Meer:, v1 pr 128.) 187522. Spyz D)
parvulus, D. spinosus.
Ekdiastylis T. Stebbing in: Ann. S.
Afr. Mus., v.10v p.155. 1912. Sp.: EH.
hexaceros.
Ekleptostylis T. Stebbing in: Ann.
S. Afr, Mus., v.10 v p. 146,153. 1912.
Sp.: HE. walkeri.
EKocuma Marcusen in: Hilgendorf in:
SB. Ges. Fr. Berlin, p.170. 1894. Sp.:
E. hilgendorfi.
Eudora Péron & Lesueur in: Ann.
Mus. Paris, v. 14 p.326. 1809. Sp.:
E. undulosa. Acalepb.
Eudora Subgen. Haan in: Th. Siebold,
Fauna Japon., p.22. 1833. Sp.: Cancer
(E.) tenax, C. (E.) impressus, C. (E.)
incisus. , Deeap.
Eudora Bate in: Ann. nat. Hist., ser. 2
v.17 p.457. 1856. Sp.: E. truncatula.
Eudorella pro: Eudora Bate 1856 A. M.
Norman in: Rep. Brit. Ass., v. 36
p. 194,197 nota. 1867.
Eudorellopsis G. O. Sars in: Forh.
Selsk. Christian., nr. 18 p.56. 1883.
Sp.: H. deformis.
Gammarus J. C. Fabricius, Syst. Ent.,
p. 418. 1875, Sp.: G. Jlocusta,G-
pulex, G. linearis, G. salinus, G. stag-
nals.
Gaussicuma Zimmer in: Zool. Anz.,
v.31 p.368. 1907. Sp.: G. vanhoeffent.
Goniacis G. 0. Sars in: Forh. Selsk.
Christian., 1864 p.159. 1865.
Gynodiastylis Calman in: Tr. zool. Soe.
London, v. 18 1v p. 342, 367. 1911. Sp.:
G. carinata, G. laevis, G. costata, G.
bicristata.
Halia Risso, Hist. Kur. mérid., v. 4
p. 52. 1826. Sp.: H. hélicoides. Moll.
Halia Bate in: Ann. nat. Hist., ser.2 v.17
p. 458. 1856. Sp.: H. trispinosa.
Hemilamprops G. O. Sars in: Forh.
Selsk. Christian., nr. 18 p. 11, 55, 56.
1883. Sp.: H. assimilis.
Hemileucon Calman in: Tr. zool. Soe.
London, v. 181 p. 2, 37. 1907. Sp.:
H. uniplicatus, H. comes.
Heterocuma Miers in: P.
London, p. 22, 57. 1879.
sarsi, H. s. var. granulata.
zool. Soe.
Sp: 2H.
Heterocyma pro: Heterocuma Miers 1879
Rye in: Zool. Rec., v.16 Index p. 5.
1881.
Nomencelator generum et subgenerum
Heteroleucon Calman in: Tr. zool. Soe.
London, v. 181 p..2, 34. 1907.
H. akaroénsis.
Holostylis T. Stebbing in: Ann. 8.
Afr. Mus.. v. 10 vp. 138, 146. 1912.
Sp.: H. helleri, H. gayi.
Thpinoé [pro: Iphinoé Bate 1856] G. O.
Sars in: Arch. Naturyv. Kristian., 0.3
p- 496. 1878.
Ann. nat. Hist., ser. 2 v.18 p.187. 1856.
Iphinoé Subgen. H. & A. Adams, Gen.
4. Moll., v.1 p.280. 1858. Sp.: Tricho-
tropis (I.) ciliata, T. (L) unicarinata,
/ Set T ‘ awit. Aappea th
Pas Com! Ointng = ae Moll. _»
\s IQ54 pare SA nsie Jan, ia! 24.
Iphithoé [pro: [phinoé Bate 1856] A. M.
Norman in: Rep. Brit. Ass., v. 36
p- 197,201. 1867.
Lamprops G. O. Sars in: Nyt Mag.
Naturv., v.12 p. 236,289. 1863. Sp.:
L. fasciata.
Lampros [pro: Lamprops G. O. Sars |
1863] Meinert in: Naturh. Tidsskr., |
Ser. oa walepelet. 1877.
Leptocuma G. O. Sars in: Svenska Ak.
Handl., v. 11 nr. 5 p.24. 1873. Sp.:
L. Kinbergii.
Leptocyma pro: Leptocwma Sars 1874
Rye in: Zool. Ree., v.11 p.552. 1876.
Leptostylis G. O. Sars in: Nyt Mag.
Naturv., v. 16 p. 343. 1869. Sp.: ZL.
longimana, L. villosa.
Leucon Kroyer in: Naturh. Tidsskr.,
ser. 2 v2 p. 181, 208. 1846. Sp.:
L. emarginatus, L. Nasica, L. deformis.
Leuconopsis A. O. Walker in: Rep.
Brit. Ass., v.66 p.419. 1896. Sp.: L.
ensifer. :
Makrokylindrus T. Stebbing in: Ann.
S. Afr. Mus., v.10 v p.150. 1912. Sp.:
M. josephinae, M. erinaceus, M. dubius,
M. cingulatus, M. serricauda, M. fragilis.
WNannastacus Bate in: Ann. nat. Hist.,
ser. 3 v. 15 p. 86. 1865. Sp.: WN.
binoculoides.
Nannasticus [pro: Nannastacus Bate
1865] A. M. Norman in: Rep. Brit.
Ass., v.38. 1868. p.259. 1869.
Das Tierreich.
Spas
209
Olbia Stal in: Ent. Zeit. Stettin, p. 118.
1862. Sp.: O. caprina. Hem.
Olbia Marcusen in: Arch. Naturg., v.331
p. 359. 1867. nom. nud,
| Oniscus Linné, Syst. Nat., ed. 10 p. 636.
|
39. Lief.: T. R. R. Stebbing,
1758. Sp.: O. asilus, O. oestrum, O.
psora, O. physodes, O. entomon, O. ceti,
O. marinus, O. scopulorum, O. aqua-
ticus, O. asellus, O. armadillo.
Iphinoé pro: Halia Bate 1856 Bate in: | Oxyurostylis Calman in: P. U.S. Mus,
v.41 p. 605, 666. 1912. Sp.: O. smithi.
Pachystylis H. J. Hansen in: Ergeb.
Plankton-Exp., v.2 G.c¢ p.58. 1895.
Sp.: P. rotundata.
Paradiastylis Calman in: MHerdman,
Rep. Ceylon Pearl Fish., v. 2 nr. 12
p.173. 1904. Sp.: P. brachyura.
Paralamprops G. O. Sars in: Rep. Voy.
Challenger, v. 19 pars 55 p. 6, 9, 26.
1887. Sp.: P. serrato-costata.
Tr. zool. Soc.
1907. "Spc:
Paraleucon Calman in:
London, v. 181 p. 2, 36.
suteri.
Paranannastacus |’. Stebbing in: Ann.
S. Afr. Mus., v.10 v p.164. 1912. Sp.:
P. reptans, P. tardus.
Petalomera Stimpson in: P. Ac. Philad.,
1858 p. 226. 1859. Sp.: P. granulata.
Decap.
Petalomera pro: Petalopus G. O. Sars
1865) .G.O) Sars: ins Forks -Selsk:
Christian., no. 18 p.13,58. 1883. Sp.:
P. declivis.
| Petalopus Motschulski in: Bull. Soe.
Moscou, v.18 p.107. 1845.
metallicus.
HOHE ae 6
Col.
Petalopus Claparéde & Lachmann,
Etud. Infus., pars 2 p. 438,442. 1859.
Sp.: P. diffluens. Prot.
Petalopus G. O. Sars in: Forh. Selsk.
Christian., p. 196. 1865. Sp.: P. declivis.
Petalosarsia T. Stebbing in: Intern.
sci. Ser., v.74 p.308. 1893. Sp.: P.
declivis.
Platyaspis [pro: Platynaspis L. Redten-
bacher 1843] L. Agassiz, Nomencl.
zool., v.2 Col. p.129. 1846. Col.
Platyaspis G. O. Sars in: Forh. Selsk.
Christian., p. 158. 1870. Sp.: P. typica.
Cumacea. 14
210
Platycuma Calman in: Sci. Invest. Fish. |
Ireland, 1904 p. 11,30, 1905. Sp.: P.
Holti.
Platysympus pro: Platyaspis G. O. Sars
1870 T. Stebbing in: Ann. S. Afr.
Mus., » 10 vy p 143. 1912: Sp.2-P:
typicus, P. brachyurus.
Platytyphlops T. Stebbing in: Ann.
S. Afr. Mus., v.10 v: p. 159. 1912) Sp.:
P. peringueyt.
Procampylaspis Bonnierin: Ann. Uniy.
lyon, 0.26 ps S215 1896s esp. ss cee
armata, P. echinata.
GanO:
Arch.
1878.
Pseudocuma G. O. Sars in: Forh. Selsk.
Christian... p268, 193: W865 iSps:2P:
bistriata.
Pseudocoma [pro: Pseudocuma
Sars 1865] G. O. Sars in:
Naturv. Kristian., v3 p. 466.
Pseudodiastylis Calman in: Siboga-
Exp., pars 36 p.16. 1905. Sp.: P. ferox.
Pseudoleucon Zimmer in: Zool. Jabrb.,
Syst. v.18 p.676. 1908. Sp.: P. sores.
Pterocuma G. O. Sars, Crust. Norway,
v.3 p.73. 1900. Sp.: P. pectinata, P.
Sowimskyi, P. rostrata.
Schizorhynchus G. O. Sars, Crust. Nor-
way, v.3 ».73. 1900. Sp.: S. bilamel-
latus, S. eudorelloides, S. scabriusculus,
S. abbreviatus.
Schizotrema Calman in: Tr. zool. Soe.
London, v. 181v p. 341, 360. 1911. Sp.:
S. depressum, S. bifrons, S. sordidum.
Secorpionura J. V. Thompson MS. in:
W. Thompson, Nat. Hist. Ireland, v. 4
p. 394. 1856. Sp.: S. vulgaris, S.
maxima, S. longicornis.
Spencebatea A. M. Norman in: Ann.
nat, Hist., ser. 5 v.3 p.67. 1879. Sp.:
S. abyssicola.
Nomenclator generum et subgenerum
Sphinoe [pro: Jphinoe Bate, 1856]
Koehler. in: Bibl. hautes Etudes,
v.82 nr.4 p.23. 1886.
Stenocuma G. O. Sars, Crust. Norway,
v.3 p.73. 1900. Sp.: S. gracilis, S.
graciloides, S.tenuicauda, S. diastyloides.
Stenotyphlops T. Stebbing in: Ann,
S. Afr, Mus., » 10v p.162. .1912. Sp.:
S. spinulosus.
Stephanomma G. O. Sars in: Svenska
Ak, Handl., v. Jl nr.5 p.16. 1873; Sp::
S. Goésii.
Strauchia Czerniavsky in: Syezda Russ.
Kst., v.1 p.64. 1868. Sp.: S. taurica.
Sympodomma T’. Stebbing in: Ann.
S. Afr. Mus., v.10 v p.138. 1912. Sp.:
S. africanus.
Vaunthompsonia Bate MS. in: Kinahan
in: Nat. Hist. Rev., 0.5 P: .p. 203:
1858. Sp.: V. cristata.
Vaunthomsonia [pro: Vaunthompsonia
Bate MS. 1858] Bate in: Ann. nat.
Hist., ser. 3 v.38 p.273. 1859.
Vauntompsonia [pro: Vaunthompsonia
Bate MS. 1858] Bate in: J. R. Dublin
Soc., v.2 p.102. 1858. Sp.: V. cristata.
Vauntrompsonia |[pro: Vaunthompsonia
Bate MS. 1858] Zimmer in: Ergeb.
Tiefsee-Exp., v.8 p. 166. 1908.
Venilia Duponchel! in: J. B. Godart
& Duponchel, Hist. Lép., v 7m p. 110.
1829. Lep.
Venilia Alder & Hancock, Ann. nat.
Hist., «0, 13 sp: lel: 1844) sspears
mucroniferda. Moll.
Venilia Bate in: Ann. nat. Hist., ser. 2
v. 17 p. 460. 1856. Sp.: V. gracilis.
Zygosiphon Calman in: Tr. zool. Soe.
London, v.181 p. 2, 25.. 1907. Sp.: Z
mortensent.
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Amathusiidae. Bearbcitet von H.S tichel (Berlin). XV und 248 Seiten
mit 42 Abbildungen. 1912 X.
Subskriptionspreis Mark 13,60. Einzelpreis Mark 18,—.
Turbellaria, II. Rhabdocoelida. Bearbeitet von L. von Graff(Graz).
XX und 484 Seiten mit 394 Abbildungen. 1913 VI.
Subskriptionspreis Mark 29,—. Einzelpreis Mark 38,—.
Pteropoda. Bearbeitet von J.J.Tesch (Helder). X VI und 154 Seiten
mit 108 Abbildungen. 1913 VI.
Subskriptionspreis Mark 10,—. Einzelpreis Mark 13,—.
Gymnophiona (Amphibia apoda). Bearbeitet von Fr. Nieden
(Berlin). X und 31 Seiten mit 20 Abbildungen. 1913 V.
Subskriptionspreis Mark 2,60. Einzelpreis Mark 3,50.
Solenogastres. Bearbeitet von J. Thiele (Berlin). X und 57 Seiten
mit 28 Abbildungen. 1913 VI.
Subskriptionspreis Mark 4,—. Einzelpreis Mark 5,20.
Cumacea (Sympoda). Bearbeitet von T. R. R. Stebbing (Tun-
bridge Wells). XVI und 210 Seiten mit 137 Abbildungen. 1913 IX.
Subskriptionspreis Mark 12,—. Einzelpreis Mark 16,—.
In Vorbereitung befindet sich:
40. Lieferung.
Pyrosomida et Cyclomyaria. Bearbeitet von G. Neumann
(Dresden).
Es wird ersucht, Subskriptions-Anmeldungen baldigst an die unter-
zeichnete Verlags-Buchhandlung direkt, oder durch Vermii rer
ae cee aad Tete saudi
Buchhandlungen, zu richten.
Berlin, September 1913.
NW 6, Karlstr. 11.
| MLR, OST 2B 1913
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