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IH ^%
DESCRIPTIVE CATALOGUE
DF THE
INDIAN DEEP-SEA FISHES
IX THE
INDIAN MUSEUM.
BEING A. BEVISED ACCOUNT OF THE DEEP-SEA PISHES
COLLECTED BT Til E
ROYAL INDIAN MARINE SURVEY SHIP
INVESTIGATOR
Bt
A. ALCOCK, M.B., C.M.Z.S., F.G.S.,
IN'I'IAN MEDI&AX SERVICE, SITKlilN TUN' UK NT OF THE INDIAN MDSEUM AND PROFESSOR OP ZOOl.OOl IX IHP
MEDICAE COLLEGE; CALCUTTA ; FORMERLY NATURALIST 10 THE MARINE SURVEY OF INDIA
6 i': -■
- w»a
(*Ul
CALCUTTA:
PRINTED BY ORDKR OF THE TRUSTEES OF THE INDIAN MUSEUM*
1899.
Price Five Rupees.
'■■'""' *"y
&^5*<236W<
\8
•
.A.
DESCRIPTIVE CATALOGUE
OF THK
INDIAN DEEP-SEA FISHES
IN Till'
INDIAN MUSEUM.
BETNG A REVISED ACCOUNT OF TITF DEEP-SEA FISHES
COLLECTED BY THK
ROYAL INDIAN MARINE SURVEY SHIP
INVESTIGATOR
l!V
A. ALCOCK, M.B., C.M.Z.S., F.G.S.,
INDIAN MEDICAL SERVICE, SUPERINTENDENT OP THE INDIAN MUSEUM AND PROFESSOR OF ZOOLOGY IN THE
MEDICAL COLLEGE, CALCUTTA; FORMERLY NATURALIST TO THE MARINE SURVEY OF INDIA.
CALCUTTA:
PRINTED PA' ORDER OF THE TRUSTEES OF THE INDIAN MUSEUM.
1899.
Price Five Rupees.
CALCUTTA : — BAPTIST MISSION PRESS.
4
K
Preface.
The following extract from the Prefatory Note to the Account of the Deep
Sea Madreporaria collected by the B. I. M. 8. Investigator will serve to explain
how the collection of Deep- Sea Fishes described in the present volume came to
be made and to be presented to the Indian Museum.
" In the year 1871 the Council of the Asiatic Society of Bengal appointed Dr. T. Oldham,
Dr. F. Stoliczka and Mr. J. Wood-Mason to form a suh-committee to report upon the desir-
ability of moving the Government of India to undertake deep-sea dredging in Indian waters.
" The sub-committee drew up an elaborate Memoir on the subject, in which definite pro-
posals for deep-sea dredging were embodied : this Memoir was submitted to Government, and
a copy of it along with a copy of the letter with which it was forwarded, is published iu the
Proceedings of the Asiatic Society of Bengal for 1871.
" The Government received the proposals of the Council of the Asiatic Society with cordial
approval : it gave a small grant-in-aid of carrying them into immediate effect, and when, in
1874, the present Marine Survey Department was established, it sanctioned the appointment,
upon the staff of the Survey, of a Surgeon-Naturalist — an appointment that had also been
strongly advocated by the organizer and first head of the Department, Commander Dundas
Taylor, I. N.
"But in the early days of the Survey (1874-1881) neither machinery nor vessels capable
of deep-sea research were available, so that Surgeon (now Lieutenant-Colonel) J. Armstrong,
I. M.S., the first Surgeon-Naturalist of the Department, had to report that it was " quite impos-
sible to carry into execution the scheme of deep-sea dredging originally proposed by the Council
of the Asiatic Society of Bengal," and had to confine himself to the Zoology of the shallow-
water and littoral, although he did occasionally manage to dredge in water as deep as 100
fathoms.
" However, in 1876, when it had been decided to construct a special vessel for the accom-
modation of the Marine Survey, the Council of the Asiatic Society again addressed the Gov-
ernment of India, and asked that provision for deep-sea dredging might not be forgotten iu
the plans for the new vessel. In reply the Government authorized the Council of the Society
to confer with the Dockyard authorities on the subject of such equipment.
" The Council thereupon appointed a sub-committee, consisting of Dr. John Anderson, then
Superintendent of the Indian Museum, and Messrs. J. Wood-Mason (then Deputy Superin-
tendent of the Indian Museum), W. T. Blanford, H. P. Blanford, and H. B. Medlicott, for the
purpose of advising the Dockyard authorities in this direction.
" The result of this and other measures was that when, in 1881, the new vessel Investigator
was ready for sea, she was properly provided with the means of undertaking deep-sea research
as opportunity should occur.
" Before this, however, Dr. Armstrong had left the Survey, and it was not until the end
of the year 1884, when Commander A. Carpenter, B. N., was appointed to the command of
the ' Investigator,' and Surgeon (now Major) G. M. J. Giles, I.M.S., to the post of Surgeon-
Naturalist, that deep-sea dredging became a recognized, if subordinate, branch of the ship's
routine.
11
" Since 1885 the Zoological collections made by the 'Investigator' have been accumulat-
ing year by year in the Indian Museum, where, in accordance with the recommendations of the
Council of the Asiatic Society of Bengal, they have beeu deposited.
"It must not, however, be supposed that deep-sea dredging occupies a very large part of
the attention of the officers of the Survey ; since, as a rule, it is only possible when the ship
is proceeding to and returning from her systematic surveys of the shores and shallows. It is
rarely indeed that as many as twenty deep-sea hauls are made in one year.
" From October 1888, when regular records began to be kept, up to the present time, 118
more or less successful hauls have been made in depths of over a hundred fathoms (100-1997
fms.).
" As regards the ' Investigator ' herself, she is a paddle-steamer of 580 tons, and for a few
facts as to her history and equipment I may refer to a paper in the Scientific Memoirs of the
Medical Officers of the Army of India for 1898."
"With regard to the contents of the present volume on the Deep-Sea Fishes,
I may state that the species not here described for the first time have already-
been noticed in the following papers : —
Journ. As. Soc. Bengal : Vol. LVIII. pt. 2, 1889, pp. 279-295, pi. xvi-xviii,
and pp. 296-305, pi. xxii ; Vol. LXIII. pt. 2, 1894, pp. 115-137, pi. vi-vii ; and
Vol. LXV. pt. 2, 1896, pp. 301-338.
Ann. Mag. Nat. Hist., (6) IV. 1889, pp. 376-399 and 450-461 ; (6) VI.
1890, pp. 197-222, pi. viii-ix, and pp. 295-311 ; (6) VIII. 1891, pp. 16-34
and 119-138, pi. vii-viii ; (6) X. 1892, pp. 345-365, pi. xviii, and pp. 207-214;
(6) XVI. 1895, pp. 144-146 ; and (7) II. 1898, pp. 136-156.
Proc. Zool. Soc. 1891, pp. 226-227.
All but four of the new species described in the above-cited papers have
been figured in the Illustrations of the Zoology of the R. I. M. S. Investigator,
Fishes, pis. I-XXVI (and XXVII-XXXV in the press).
As it is intended that those plates should be bound up with the present
Catalogue, together with a small appendix on the Shore Fishes discovered by the
Investigator, no special illustrations have been prepared for this volume.
I have however added (facing page 12) a chart, compiled from plate ii of
Dr. Ernst Koken's Die Vorwelt mid ihre Entmckelungsgeschichte, which is meant
to elucidate the theories upheld in the Investigator Reports as to the origin and
past geographical relations of the fauna of the Indian Oligobenthus.
For the opportunity of making this chart and for help in compiling it, I am
indebted to Mr. T. H. Holland of the Indian Geological Survey.
To explain an apparent want of uniformity in the plan of this Catalogue,
I must mention that I have intentionally omitted diagnoses of the families and
genera, and of the one or two species, heretofore well known to inhabit Indian
Ill
seas. The Catalogue is designed to meet the wants of students of the Indian
fauna, and it would have been superfluous to reprint information that can be
found recorded in Day's standard volumes on the Fishes of India.
I cannot let this volume go out without acknowledging the essential and
fundamental obligations that it owes to the Catalogue of the Fishes in the British
Museum, to the Introduction to the Study of Fishes, and to the Report on the Beep
Sea Fishes collected by H. M. S. Challenger.
In any scheme of classification the compass and standing of the various
units will vary with individual opinion ; but the value and influence of a well-
organized system of broad, clear, consistent generalizations must always
last, and I am happy to add my tribute of regard to the author of the works
in which these qualities are so conspicuously manifest.
A. Alcock, Major, I. M. S.,
Superintendent of the Indian Museum.
INTRODUCTION.
46
55
57
2
The Fishes included in this Catalogue were all of them dredged by the
R. I. M. S. " Investigator," in deep water, between the meridians of 65° and 99°
E. and the parallels of 53 and 24° N., during the years 1885-1899.
They number, exclusive of a few mangled remains that cannot be identified
with certainty, 169 species, namely : —
Chondropterygii ... ... ... ... 9 species.
Acanthopterygii
Anacanthini
Physostomi
Plectognathi
Of these — 126 species have, so far as is known, been taken only by the
" Investigator ; " while 43 species are believed to be identical with species found
in other parts of the world, the identities having been assured by actual com-
parison of specimens in 13 instances.
It will be interesting, to begin with, to inquire into the geographical distri-
bution of these 43 species and to see if they throw any light on the outside
relations of the Fish-fauna of the Indian seas.
It appears that the following 23 of them, or over thirteen per cent, of the
whole, are common to these seas and the Atlantic, and many of them to certain
very definite areas of that Ocean : —
Setarches Giintheri Johns.
* Hoplostethus mediterraneum C. V. (Also in the Mediterranean).
Trachichthys Darwinii Johns.
Polymixia nobilis Lowe.
* Antigonia capros Lowe.
Bembrops caudimacula Stdr.
( = *Hypsicometes golioides G. & B.).
Chiasmodus niger Johns.
Chaunax pietus Lowe.
* Dicrolene intronigra G. & B.
Dlplacanthopoma brachysoma Gthr.
\ * Macrurus cavernosus G. & B.
[ (probably identical with the Mediterranean Hymenocephalus italicus Giglioli)
Hacrurus Isevis Lowe. (Also in the Mediterranean).
Bathygadus longifilis G. & B.
* Argyropelecus hemigymnus Cocco. (Also in the Mediterranean).
{
* Species marked with an asterisk have been compared with actual specimens from other localities.
* Stemoptyx diaphana Herni.
Cyclothone elongata (Gfchr. ).
Cy clot hone microdon (Gthr.).
Chauliodus Sloanii Bl. Sclm. (Also in the Mediterranean).
* Nenscopelus macrolepidotus Johns.
Platytroctes apus Gthr.
* Synaplwbranchus pinnatus Gronov.
Uroconger vicinus Vaillant.
? Leptoderma macrops Vaillant.
The remaining 20 are Indo-Pacific species.
Very significant, to my mind, is the occurrence in these seas — it also occurs
in Japan, where it was originally found — of Bembrops caudimacula (=Hypsicometes
gobioides G. & B.). Hardly less significant is the distribution, having regard to
its mode of life, of Ghaunax pictus.
Bembrops caudimacula, which is undoubtedly the young of Bembrops gobioides,
appears to be common off the West Indies and neighbouring coasts of North
America at depths of 68 to 324 fathoms, and a good number of specimens have
been taken in the Andaman Sea at 107 to 194 fathoms. It is a Trachinoid fish
with a large flat head and a big shovel mouth, very much the form of Platy-
cephalic, and is undoubtedly — like most of the members of its family — a dweller
on or near the bottom. A fish that most commonly lives near the 100-fathom
limit cannot be truly called bathybial, nor would anyone who has handled
Bembrops be likely to decide that it belonged to the nectic fauna; so that some
other explanation must be found for its peculiar geographical distribution. And
if this explanation will also serve to throw some light on the distribution of, e.g.,
Lobotes surinamensis, which is so far from being pelagic or nectic that it enters
brackish water ; and if it will also enable us to better understand the curious
distribution, e.g., of Symbranchus, of the Chromides, and of the Gyprinodontidx,
its probability will be enhanced,
The hypothesis that appears to offer the most satisfactory explanation is,
that a very considerable part of the fish-fauna of the Oriental region originated
from, and to a certain extent is a remnant of, the fauna of the Tertiary Mediter-
ranean of Professor Suess — of a Mediterranean that extended from the present
Gulf of Mexico, through the present Mediterranean basin, far into the Eastern
Hemisphere,
In the Account of the Deep Sea Madreporaria collected by the Investigator,
pp. 5-^10, I have discussed some evidence in favour of this hypothesis that is
afforded by certain other elements of the marine fauna; and in the Account of
the Veep Sea Brady ur u collected by the Investigator, pp. 2, 3, 82, 85, I have added
m,i,]o further confirmatory evidence, derived from the present distribution of
* Species marked with an asterisk have been compared with actual »pociincns.
Certain genera and species of Crabs ; so that the evidence to be adduced from the
fish-fauna will already have been corroborated from several other marine groups.
It is no part of my plan, in this series of Reports, to refer to the evidence
that lies outside the boundaries of the " Investigator" collections ; but no one
who remembers the singular distribution of certain Mammals (e.g. the Tapirs),
Birds (e.g. the Trogons, Barbets, and Whistling Teal), and Amphibia (e.g.
the Cmciliidse), and no one, more especially, who has considered the geogra-
phical range of a number of Indian Snake and Lizard families and genera
too numerous to mention, can have failed to have suspicions of the former ex-
istence of some land connexion (which this sea-connexion implies) between the
tropical and subtropical regions of America and the Old World — suspicions
which the comfortable formula " similarity of conditions" alone will not satisfy.
Moreover, as regards marine fishes, the researches of Dr. Giinther have
familiarized us with the idea of a former direct and open connexion between the
Mediterranean and Japanese Seas (Introcl. Study of Fishes, p>- 270).
The testimony supplied by the Indian fish-fauna favourable to the hypo-
theses in question may be briefly summarized as follows : —
A. The Marine Fishes. The total number of genera of Indian marine
fishes is about 350, of which
(1) about 28 per cent, are common to the Atlantic, the Mediterranean, and
the Indo-Pacifie :
(2) nearly 6 per cent, are common to the Atlantic, the Bed Sea, and the
Indo-Pacifie, to the exclusion of the Mediterranean :
(3) nearly 20 per cent, are common to the Atlantic and Indo-Pacifie, to the
exclusion of the Mediterranean and Red Seas :
(4) nearly 3 per cent, are common to the Mediterranean and Indo-Pacifie
to the exclusion of the Atlantic.
That is to say, taking the Atlantic and Mediterranean as one integral region,
over 56 per cent, of the genera of Indian marine fishes are also found in that
region.
Not only so, but a considerable number of species are common to the two
regions, and although it may be argued that most of these — such as the Sword-
fishes, the Pilot fish, the Tunnies, the Sucker fish, the Stemoptychidse, and even
certain Glupeidss and Beryeidx — are oceanic forms of unlimited range, yet this
objection cannot, I think, apply to such species as Lobotes surinamensis, Bembrops
caudimacula, Chaunax pictus, or Macrurus cavernosus, or to Sargus noct and
Crenidens Forslcalii.
B. The Freshwater Fishes. Exclusive of certain immigrants from marine
families, the freshwater fishes of India are Carps (which are nearer two-thirds
than half the whole freshwater fish-fauna), Siluridse, Cyprinodontidse, Ghromides
and Nandinae, Symbranchidae, Notopteridx, Labyrinthici, Ophiocephalidae, Bhyncho-
bdellidae. Excluding the Carps and the Nandinae, we have our attention at once
attracted bj certain curiosities of distribution.
For instance, in the small family of Symbranchidae, we find Symbranchus
with three species, one common in the jheels of the Oriental region, another
common in tropical America. The third is from Indo-Australian waters.
Again, the Gyprinodontidas, of which 2 genera and 5 species occur in India,
have a most suggestive range, being found in tropical and temperate America,
in tropical Africa and the regions of the Mediterranean basin, and in south-
western Asia — one genus, Cyprinodon, being represented in India and all round
the Mediterranean.
Again, the Siluroid genus Arms has a tropical distribution that fits in most
remarkably with the theory of a tropical Mediterranean of wide extent east and
west.
The same is the case with the Chromides, which are freshwater fishes of
tropical America and Africa, and of which three species are found in India.
The other families — Notopteridae, Bhynchohdellidas, Labyrinthici and Ophioce-
phalidse — do not extend further to the west than Western Africa.
I may conclude this Introduction with some Tables of genera and species
that are common, on the one hand to the Atlantic and Mediterranean and, on
the other hand, to the seas of India. Of course many of these are widely ranging
forms, and may be discounted ; but, on the other hand, no mention is made of
several characteristic Indo-Pacific genera that have been discovered, fossil, in
the Tertiary deposits of Northern Italy, and a respectable number of forms that
are common to the Atlantic, the Mediterranean and Japan are not included
here.
I. List of Genera and Species common to the Atlantic, the Mediterranean, and the Indian Fauna.
1. Carcliarias. 10. Rhinobatis.
2. ZygEena. vii. „ halavi.
i. „ blocliii. viii. „ columnae.
ii. „ tudes. 11. Raja.
3. Larana. 12. Trygon.
jii. „ spallanzauii. ix. „ pastinaca.
4. Odontaspis. x. „ bennettii (Atl. not Med.),
5. Alopeoias. 13. Pteroplataaa.
iv. „ vulpes. 14 Myliobatis.
6. Notidanus. 15. RLrinoptera.
7. Scyllium. 10. Dieerobatis.
8. Centrophorus. 17. Serranus.
9. Pristis. 18. Chelidoperca.
v. ,, pcctinata. 19. Piistipoma.
vi. „ perrotteti (Atlantic not Med.). xi. „ stridens (Med. not Atl.).
20.
Lobotes.
60.
Regalecus.
xii.
„ surinamensis.
61.
Heliastes.
21.
Apogon.
62.
Novacula.
22.
Dentex.
63.
Julis.
23.
Smaris.
64.
Coris.
24.
Chastodon.
65.
Physiculus.
25.
Sargus.
66
Fierasfer.
xiii.
„ noct (Med. not Atl.).
67.
Ammodytes.
26.
Pagrus.
68.
Macrurus.
27.
Chrysoplirys.
69.
Mystaconurus.
28.
Pimelepterus.
70.
Malacocepbalus.
29.
Sebastes.
xxii.
,, lams.
30.
Scorpsena.
71.
Arnoglossus.
31.
Hoplostethus.
72.
Platophrys.
xiv.
,, mediterraneum.
73.
Solea.
32.
Utobrina.
74.
Syngnatbus.
33.
Sciaena.
75.
Hippocampus.
34.
Scia?noides.
xxiii.
„ guttulatus.
35.
Histiopborus.
76.
Nerophis.
36.
Tricbiurus.
77.
Balistes.
37.
Tbyrsites.
xxiv.
„ maculatus (Atl. not Med.)
38.
Caranx.
78.
Ostracion.
XV.
,, carangus.
XXV.
„ nasus (Med. not Atl.).
39.
Naucrates.
79.
Tetrodon.
xvi.
„ dnctor.
80.
Diodon.
40.
Ecbeneis.
xxvi.
„ hystrix.
xvii.
„ remora.
81.
Orthagoriscus.
sviii.
„ naucrates.
82.
Argyropelecus.
41.
Seriola.
xxvii.
„ hemigymnus.
42.
Stromateus.
83.
Cbauliodus.
43.
Coryphsena.
xxviii
,, sloanii.
xix.
„ hippurus.
84.
Saurus.
44.
Scomber.
85.
Scopelus.
45.
Thynnus.
86.
Cbloropbthalmus.
XX.
„ tbnnnina.
87.
Belone.
xxi.
„ pelamys.
88.
Hemiramphus.
46.
Cybium.
89.
Exocoatus.
47.
Batrachus.
xxix.
„ evolans.
48.
Lopbius.
XXX.
„ furcatus.
49.
Trigla.
90.
Stomias.
50.
Peristetbium.
91.
Cbtpea.
51.
Dactylopterus.
92.
Engraulis.
52.
Gobius.
93.
Alepocephalus.
53.
Callionymus.
94.
Anguilla.
54.
Salarias.
95.
CoDgromursena.
55.
Tripterygium.
96.
Murasna.
56.
Cepola.
xxxi.
„ afra (Atl. not Med.).
57.
Spbyraena.
97.
Muraenesox.
58.
Atherina.
98.
Nettastoma.
59.
Mugil.
99.
Ophichtbys.
II. List of Additional Genera and Species common to the Atlantic the Red Sea and the Indian
Fauna, excluding the Mediterranean.
100. Tmniiira.
101. Mesoprion.
102. Priacanthus.
103. Holacanthus.
104. Mulloides.
105. Upeneus.
106. Holocentrum.
107. Myripristis.
10S. Acanthurus.
109. Opistbognatbus.
110. Antennarius.
III. List of Additional Genera
120.
Galeocerdo.
121.
Mustelus.
122.
Centroscyllium.
123.
G inglymostoma.
124.
Narcine.
125.
Aetobatis.
xxxiii.
„ narinari.
126.
Dules.
127.
Gerres.
128.
Epbippus.
129.
Letlirinus.
130.
Setarches.
xxxiv.
,, giintheri.
131.
Tracbicbthys.
XXXV.
„ darwinii.
132.
Polymixia.
xxxvi,
„ nobilis.
133.
Melamphaes.
134.
Bathyclnpea.
135.
Polynemus.
136.
Otolitbus.
137.
Antigonia.
xxxvi;
i. „ capros.
138.
Micropteryx.
xxxviii. ,, chrysurus,
139.
Chorincmus.
140.
Tracbynotus.
xxxix.
„ ovatus.
141.
Pseues.
xl.
„ regulus.
142.
Elacate.
xli.
„ nigra.
143.
Malacautkus.
111. Periophthalmus.
112. Salarias.
113. Eleotris.
114. Glypbidodon.
115. Platyglossus.
116. Pseudoscarus.
117. Callyodou.
118. Monacantbus.
119. Albula.
xxxii. „ conorbyiicbus.
and Species common to the Atlantic and the Indian Fauna
excluding the Mediterranean.
144. Chiasmodus.
xlii. „ niger.
145.
Bembrops.
xliii.
,, caudimacula.
146.
Chaunax.
xliv.
„ pictus.
147.
Ceratias.
148.
Onirodes.
149.
Dibrancbus.
150.
Sicydium.
151.
Fistularia.
152.
Pomacentrus.
153.
Cossyphus.
154.
Bregmaceios.
155.
Brotula.
156.
Neobytbites.
157.
Dicrolene.
xlv.
„ intronigra.
158.
Diplacanthopoma.
xlvi.
„ bracbysoma.
159.
Bassozetus.
160.
Batbygadus.
xlvii.
,, longifilis.
xlviii.
Mystaconurus cavernosue.
161.
Citharicbtbys.
162.
Pseudorbombus.
163.
Apboristia.
1G4.
Cynoglossus.
165.
Dovyicbtbys.
166.
Neoscopelus.
xlix.
,, macrolepidotus.
167.
Batbypterois.
168.
Arius.
169.
Opistltopterus.
170.
Raconda.
171.
Cliatoessus.
172.
Spiatelloides.
173.
Elops.
1.
,, saurus.
174.
Pellona.
175.
Megalops.
176.
Sternoptyx.
li.
„ diaphana,
177.
Cyclothone.
lii.
,, microdon,
liii.
„ elongata.
178.
Malacosteus.
179.
Photostomias.
180.
Bathytroetes.
181.
Platytroctes.
liv.
„ apus.
182.
Xenodermichthys.
183.
Leptoderma.
184.
Halosaurus.
185.
Nemiclitbys.
186.
Synaphobranclius.
lv.
..
187.
Uroconger.
Ivi.
,, vicinus
188.
GymnomurEena.
lvii.
Diodon maculatus.
189.
Chilomycterus.
pinnatus.
IV. List of Genera and Species common to the Mediterranean and Indian Fauna, excluding the
' Atlantic.
190.
Lates.
191.
Diagramma.
192.
Crenidens.
lviii.
„ forskalii.
193.
Uranoscopus.
194.
Lepidotrigla.
195.
Cristiceps.
196.
Synaptura.
197.
Odontostomus.
19S.
Saurenchelys.
To sum up : if we estimate the number of Indian genera of marine fishes at
350, and of species at 1200, then over 56 per cent, of the genera and close on 5
per cent, of the species are also found in the Atlantic-Mediterranean region.
This does not include the Cephalochordate forms AmpJdoxus and Asijm-
metron.
Amphioxus occurs in the tropical and temperate parts of the Atlantic, in
the Mediterranean, off the Indian . and Ceylon coasts, and in the tropical and
temperate parts of the Pacific.
Asymmetron has only been found (1) in the West Indies and (2) off the
south-eastern coast of Papua. In the latter locality a species has recently
been discovered by Dr. Arthur Willey, who was at once attracted by the
singular fact of geographical distribution thus revealed.
List of the Indian Deep-Sea Fishes contained in the Collection of the Indian Museum.
[The references for the plates and figures are to the Illustrations of the Zoology of the
Investigator for 1892, 1894, 1895, 1897, 1898, 1899,
1900.]
Page.
Plate
Pig.
CHONDROPTERYGIT :—
*
i.
Family Spinacidse : —
1. Centrophorus Rossi. ... ...
13
XXVI
3
2. Centroscyllium ornatum.
14
... VIII & XXXV .
.. 2&1
ii.
Family Scyllidre : —
3. SCYLLIUM CANESCENS, Gthr. ...
16
4. Scyllium hispidum.
15
VIII
.. 3
5. Scyllium quagga.
17
XXVII
.. 1
iii.
Family Torpedinidse : —
6. Benthobatis Moresbyi.
18
XXVI
1
iv.
Family Rajida? : —
7. Raja mamillidens. ... ...
19
VIII
1
8. Raja Poivelli.
20
XXVI
.. 4
9. Raja Johannis-Davisi.
21
XXVII
.. 2
TELEOSTEI:—
Acanthoptertgii : —
i.
Family Serranidse : —
1. Chelidoperca investigatoris. ...
23
X
.. 1
2. Synagrops philippinensis, (Gthr.).
24
XXVIII
.. 1
3. Brephostoma Carpenteri.
26
XVIII
.. 2
ii.
Family Scorpaanida? : —
4. Sebastes hexanema, Gthr. ...
27
5. Setarches Guntheri, Johns.
28
X
o
6. Pterois macrurus.
30
XVIII
.. 4
7. Minous inermis.
30
XVIII
,.. 3
iii.
Family Berycidae : —
8. Monocentris japonicus, C. V.
32
9. Hoplostethus Mediterraneum, C. V. ...
34
XIV
.. 3
10. Trachichthys Darwinii, Johns.
35
11. Trachichthys intermedics, Hector.
36
12. Melamphaes mizolepis, Gthr.
37
13. Polymixia nobilis, Lowe. ...
38
iv.
Family Kurtidse : —
14. Bathyclupea Hoskynii.
40
XXVIII .
.. 2
v.
Family Trichiuridas : —
15. Thyrsites bengalensis.
42
XV
,.. 10
vi.
Family Carangidre : —
16. Bathyseriola cyanea. ...
43
XVIII
1
vii.
Family Cyttidse: —
17. Antigonia capros, Lowe. ...
44
Page.
Plate.
Fig.
Family Trachinidse : —
18. Uranoscopus crassiceps. ...
Champsodon vorax, Gthr. ...
Bembrops caddjmacdla, Stdr.
Chiasmodds niger, Johns. ...
Family Pediculafci : —
22. Lophius indicus.
Lophius gracilimanus.
Lophius mutilus.
Lophius lugubris.
Ceratias bispinosus, Gthr. ...
Onirodes glomerosus.
Chaonax pictds, Lowe.
Halieutjia stellata, Wahl....
Halieutsea nigra.
Halieutsea coccinea.
Halieutsea fumosa.
Dibranchus nasutus.
Dibranchns micropus.
Malthopsis lutea-
Halicmetus ruber.
Family Cottidas : —
37. Teigla hemisticta, Schleg.
Lepidotrigla spiloptera, Gthr.
Lepidotrigla longipinnis.
Peristethium Rivers- Andersoni.
P eristethium serrulatum.
Peristethium investigatoris.
Family Gobiidse : —
43. Gobius' cometes. ...
44. Amblyopus ar otocephalus.
45. Callionymus carebares.
46. Callionymus kaiancs, Gthr. ...
19.
20.
21.
23.
24.
25.
26.
27.
28.
29.
30.
31.
32.
33.
34.
35.
36.
38.
39.
40.
41.
42.
45 ...
X
.. 4
47 ...
XXVIII
.. 5
48 ...
XX & XXXI ..
. 6 &2
50 ...
XXVIII .,
. 3
53 ...
XIX
,. 3
54 ...
XXIX
. 3
54 ...
X
. 2
55 ...
XIV
I
56 ...
XXXV
. 2
57 ...
XXVIII
. 4
58
60
60 ...
XIX
. 2
61 ...
XIX
. 1
62 ...
XIV
. 2
63 ...
XX
2
63 ...
XX
1
64 ...
XIX
. 4
66 ...
XIX
, 5
67
67
68 ...
XXXI
1
68
69 ...
XXV
2
70 ...
XXV
1
71 ...
XX
3
72 ...
XX
7
73 ...
XX
4
74
Anacanthini :
Family Gadidee : —
1. Bregmaceros Macclellandii, Thomps.
2. Physiculus roseus.
3. Physiculus argyropastus.
Family Ophidiidae : —
4. Neobtthites macrops, Gthr. ...
5. Neobythites conjugator.
6. Neobythites nigripinnis.
7. Neobythites steatiticus.
8. Neobythites pterotus.
9. Neobythites squamipinnis.
10. Dicrolene intronigea, G. & B.
11. Dicrolene multijilis. ...
2
75
76
XI
.. 2
77
XXII
.. 1
80
81
XVII
.. 4
82
XI
.. 3
82
XXI
.. 2
83
... XI and XXIX .
.. 4 and 1
84
XXI
.. 1
85
86
XI
.. 1
10
Page.
Plate.
Fig.
12.
Bicrolene nigricandis.
87 ...
II
... 4
13.
Bassozetus glutinosus.
88 ...
I
... 3
14.
Dermatorus trichiurus.
90 ...
I
... 1
15.
Dermatorus melanocephalus. ...
91 ...
XXI
... 4
16.
Dermatorus melampeplus.
92 ...
XVII
... 3
17.
Glyptophidium argenteum.
93 ...
II
... 3
18.
Glyptophidium macropus.
94 ...
XV
... 6
19.
Lamprogrammus niger.
95 ...
I
... 2
20.
Lamprogrtimmus fragilis.
96
21.
Tauredophidium Hextii.
97 ...
XXI
... 3
22.
Diplacanthopoma Ilivers- Anderson i.
99 ...
XVII
1
23.
DlPLACANTHOPOMA BEACHTSOMA, Gthr.
... 100 ...
XVII
... 2
24.
Diplacanthopoma raniceps.
... 101 ...
XXVI
2
25.
Saccogaster maculata.
... 102 ...
XXIX
2
26.
Hephthocara simum.
... 103 ...
XXII
... 3
Family
Macruridse : —
27.
Macrurus parallelus, Gthr. ...
... 106
28.
Macrurus quadricristatus.
... 106 ...
III
... 1
29.
Macricrus flabellispinis.
... 107 ...
XVI
... 2
30.
Macrurus investigatoris.
... 109 ...
III
... 4
31.
Macrurus Petersonii.
... 110 ...
III
... 5
32.
Macrurus nasutos, Gthr.
... Ill ...
XIII
... 3
33.
Macrurus semiquincuitciatus. ...
... Ill ...
XII
... 2
34.
Macrurus polylepis.
... 112 ...
XXIX
... 4
35.
Macrurus pumiliceps.
... 113 ...
XVI
• • • O
36.
Macrurus Hextii.
... 113 ...
XII
... 3
37.
Macrurus Wood-Masoni.
... 114 ...
XIII
1
38.
Macrurus macrolophus.
115 ...
XII
... 1
39.
Macrurus lophotes.
... 116 ...
III
2
40.
Macricrus Huskynii.
116 ...
IX
... 4
41.
Macruros cavernosds, G. & B.
... 117 ...
III
... 3
42.
Macrurus LjEvis, Lowe.
... 119
43.
Bathtgadds longifilis, G. & B.
120
44.
Bathygadus furvescens.
... 121 ...
XVI
... 1
Family Ateleopodidre : —
45. Ateleopus indicus.
Family Pleuronectidaa : —
46. Chascanopsetta lugubris.
47. Boopsetta prselonga.
48. Boopsetta maculosa.
49. Lmops inacrophlhalmus.
50. Solea umbratilis.
51. Aphoristia Gilesii.
52. Aphoristia Wood-Masoni.
53. Aphoristia septemstriata .
54. Aphoristia trifasciala.
55. Cynoglossus Carpeuteri.
123
II
125 .
XV
.. 3
126 .
.. XV & XVII .
.. 2 & 5
127 ..
XV
1
128 .
XXIII
1
129 .
XV
.. 4
131 .
XIV
.. 4
131 .
XVI
.. 4
132 ,
11
1
133 .
XV
.. 5
133 .
XXII
,. 5
11
Phtsostomi :—
i. Family Sternoptyuhidae : —
1. Argyropelecus hemigymnos, Cocco.
2. Sternoptyx diaphana, Herm.
3. Polyipnus spinosds, Gtlir. ...
4 Cyclothone elongata, (Gthr.).
5. Cyclothone microdon, (Gthr.).
6. Photichthys corythasolus.
7. Chauliodus Sloanii, Bl. Schn.
8. Chauliodus pammelas.
ii. Family Stomiatidse : —
9. Stomias nebulosus.
JO. Stomias elongatus.
11. Mulacosteus, sp — ...
12. Photostomias atrox.
12a. Astronesthes, sp.
iii. Family Scopelidae : —
13. Scopelarchus Giintheri.
14. Harpodon squamosus.
15. Chlorophthalmus comiger. ...
16. Bathypterois Giintheri.
17. Bathypterois insularum.
18. Bathypterois atricolor.
19. SCOPELUS ENGRAULIS, Gthr....
20. Scopelus Domerilii, Blkr....
21. Scopelus pterotus.
22. Scopelus pyrsobolus.
23. Neoscopelus macrolepidotus, Johns.
24. Scnpelengys tristis.
25. Odontostomus atratus.
iv. Family Alepocephalida? : —
26. Alepocephalus bicolor.
27. Alepocephalus Blanfordi.
28. Alepocephalus edentulns.
29. Bathytroctes squamosus.
30. Narcetes erimelas.
31. Platytrootes apds, Gthr. ...
32. Aulastomomorpha phosphorops.
33. Xenodermichthys Giintheri. ...
34. Xenodermichthys squamilaterus .
35. Leptoderma ajjinis.
v. Family Halosaurida? : —
36. Halosaurus anguilliformis. ...
37. Halosaurus mediorostris, Gthr.
38. Halosaurus parvipennis.
39. Halosaurus carinicauda.
40. Halosaurtis nigerrimus.
Page.
135
136
138
139
141
142
144
145
146
147
149
150
211
153
154
155
157
158
159
161
162
162
163
164
166
167
169
171
172
173
175
177
178
180
181
182
184
185
186
187
188
Plate.
Fig.
XXV
XXX
VII
XXXIII
.. 4
XXX
2
XXXV
.. 3
XVII
.. 7
XXX
.. 1
XV
.. 8
VII
.. 6
XXXII
.. 1
XVII
.. 6
IX
.. 3
XXX
.. 3
VII
.. 7
XXXIII
.. 3
IV
2
IX
1
XXXII
.. 4
V
.. 1
IV
.. 1
V
2
XXXII
.. 2
XXV
.. 4
XXXII
.. 3
VII
XXXIII
VII
XXXIII
1
2
2
12
vi. Family
41.
42.
43.
44.
45.
46.
47.
48.
49.
50.
51.
52.
53.
54.
55.
56.
57.
PlECTOGNATHI
i. Family
1.
2.
Page.
Plate.
Fig.
Mura?nidse : —
Nemichthys acanlhonotus.
.. 190 ..
XIV
... 5
Gavialiceps microps.
... 191
Dysomma Bucephalus.
.. 192 ..
VI
... I
Dysommopsis mucipara.
.. 193 ..
XXXIV
... 1
Synaphobranchcts pinnatus, GroDov.
195
Coloconger raniceps.
196 ..
VII
... 4
Gongromursena squaliceps. ...
... 197
Congromursena macrocercus
198 ..
VII
... 5
Congromursena nasica.
198 ..
IX
... 2
Gongromursena musteliceps. ...
199 ..
XV
7
CONGROMUR/ENA GUTTULATA, Gthr.
199
Uroconger vicinus, Vaillant.
.. 200
Promyllantor purpureus.
202 ..
VI
... 2
Sauromursenesox vorax.
.. 203 ..
VI
... 3
Xenomystax trucidans.
.. 205 ..
XVI
... 5
Saurenchelys tseniola.
.. 206 ..
XXXIV
... 2
Nettenchelys Taylori.
.. 207 ..
XXV
... 5
Sclerodermi : —
Triacanthodes ethiops.
Halimochincrgus centriscoides.
208
209
XV
XXXI
N. B. — The species of Astronesth.es was mislaid among some specimens of Shore Fishes, and
was only discovered after the text relating to the Stomiatidse had been printed off.
Explanation of the Chart.
The Chart has been compiled from Plate II of Dr. Ernst Koken's Die
Vorivelt und Hire EntmckeltmgsgescMchte, and shows the supposed coast-lines of
the Tertiary Continents.
The present coast-lines are indicated by dotted lines.
The supposed Tertiary coast-lines of Koken are indicated by thick black
lines.
The Great Inland Sea, stretching from the present Gulf of Mexico to the
present Arabian Sea, is coloured dark blue.
The Chart is meant to elucidate the theory advanced in the text as to the
origin of a considerable part of the Fish Fauna of India, especially that of the
Oligobenthus.
12
Family Muraenidse : —
41. Nemichthys acanthonotus.
42. Gavialiceps microps.
43. Dysomma bucephalus.
44. Dysommopsis mucipara.
45. Synaphobeanchus pinnatds
46. Coloconger raniceps.
Gronov.
Page.
Plate.
Fig.
... 190 ...
XIV
... 5
... 191
... 192 ...
VI
... I
... 193 ...
XXXIV
... 1
195
... 196 ...
VII
... 4
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13
Sub-class CHONDROPTERYGII.
Order Plagiostomata.
Sub-order Selachoidei.
In the Fauna of British India, Fishes, Vol. I. p. 6, five families of Sharks
are recorded for the seas of India, namely, Carchariidas, Lamnidx, Bhinodontidse,
Notidanidgs, Scylliidse.
A sixth family, Spinacidas, with two genera, has been discovered by the
" Investigator " to be also represented.
Family SpindCldCB.
" Two dorsal fins : no anal. Mouth but slightly arched ; a long deep
" straight oblique groove on each side of the mouth. Spiracles present :
" gill-openings narrow. Pectoral fins not notched at their origin. No nie-
" titating membrane." (Giinther) .
Key to the Indian genera of the family Spinacidse.
Each dorsal fin with a spine : no lateral folds of skin along the belly : —
1 . Upper teeth erect, with a single cusp : lower teeth
oblique ... ... ... ••• ••• Centeophoeus.
2. Teeth equal in both jaws, very small, tricuspid ... Centeoscyllium.
Centeophoeus, Midler and Henle.
Centrophorvs, Miiller and Henle, Plagiostomen, p. 88 : Giinther, Catalogue of Fishes, VIII. 419 (et synon.).
" Two dorsal fins, each with a spine which is sometimes hidden below the
" skin : no anal fin. Trunk elongate without lateral folds. Mouth wide, but
" slightly arched : a long deep straight oblique groove on each side of the mouth.
" Teeth of the lower jaw with the point more or less inclined backwards or out-
" wards. Upper teeth erect, triangular or narrow-lanceolate, with a single cusp.
" No membrana nictitans. Spiracles wide, behind the eye. Gill-openings narrow."
{Giinther).
l. Centrophorus Rossi, Alcock.
Centrophoms Rossi, Alcock, Ann. Mag. Nat. Hist. August, 1898, p. 143 : Illustrations of the Zoology op
the Investigator, Fishes, plate XXVI. fig 3.
Snout spathulate, much produced, its length measured from the most convex
part of the upper jaw is one-eighth of the total, or more than 2^ times the inter-
narial space, or about three times the major diameter of the eye-ball. The
nostrils, though completely ventral, cause a distinct notching of the margin of
the snout seen from above.
14
Mouth crescent-shaped, considerably protractile, the distance between its
angles is about two-thirds the length of the snout. The labial fold extends
rather more than half-way between the angle of the mouth and the middle of the
lower jaw.
Teeth of the upper jaw acute, triangular, in two series : those of the lower
jaw very oblique, in a single series.
The posterior borders of all the fins are frayed or fringed, this not being
due to abrasion. The angle of the pectoral is rounded ; the extremity of the
ventral is almost in the perpendicular with the after limit of the base of the
second dorsal. The dorsal spines (measured obliquely as they stand) are not
much more than half the greatest height of their fins, but their points project
well beyond the skin ; the second dorsal is a little larger than the first.
Scales minute, acutely and very elegantly tridentate, or anchor-shaped.
Colour uniform jet-black.
A single specimen, 10 inches long, from off the Travancore coast, 430 fath.
Regd. No. — .
Named in memory of Captain Daniel Ross, who, according to Sir Clements
Markham, was the first Indian " Marine Surveyor-General " (1823-1833) to
introduce a really scientific method into marine surveying in India.
This species is closely related to the Centrophorus calceus Lowe and the
Centrophorus obscurus Vaillant of the Atlantic-Mediterranean gate and to the
Centrophorus foliaceus of the Japanese Sea.
Centroscyllium, Midler and Henle.
Centroscyllium, Miiller and Henle, Plagiostomen, p. 191 : Giinther Catalogue of Fishes, VIII. 425 : Jordan and
Evermann, Fishes of North and Middle America, I. p. 56: Goode and Bean, Oceanic Ichthyology, p. 11.
Paracentroscyllium, Alcock, Ann. Mag. Nat. Hist. Nov. 1889, p. 379.
" Two dorsal fins, each with a strong spine : no anal fin. Mouth crescent-
" shaped : a straight oblique groove at each angle of the mouth. Teeth equal
" in both jaws, very small, tricuspid. No membrana nictitans. Spiracles of
" moderate width. Gill-openings rather narrow." (Giinther).
Centroscyllium contains three species, Centroscyllium fabricii from Green-
land, North America and the coast of " Sahara," Centroscyllium granulatum from
the Falkland Is., and Centroscyllium ornatum from the Arabian Sea and Bay of
Bengal, 690-620 and 405-285 fathoms.
2. Centroscyllium ornatum, Alcock.
PaTacentroscyllium ornatum, Alcock, Ann. Mag. Nat. Hist. Nov. 1889, p. 379 .- Illustrations of the Zoology
or the Invkstioatoic, Fishes, PL. VIII. FIG. 2 (yonng), pi. XXXV. §g. 1.
Centroscyllium ornatum, Alcock, Jouin. As. Soc. Bengal, Vol. LXV. pt 2, IH'M, pp. 30S, 310,
15
All the tissues extremely fragile.
Head very large, very flat and depressed, branchial regions laterally
expanded.
Snout much depressed, somewhat polygonal in outline : nostrils very large,
situated on ventral surface of edge of snout. Under surface of snout with
numerous rather large pores, two rows of which form an elegant Y- or V-shaped
figure that extends between the nostrils.
Eyes very large, their major diameter four-fifths as long as the snout
measured from the mouth, and a fifth the length of the head (branchial region
included).
Spiracles rather small, about a third the major diameter of the eye, situated
on the upper surface of the head, behind the eye.
Mouth crescentic, large, the distance between its angles being as long as,
or slightly longer than the snout : minute tricuspid teeth in both jaws.
Body covered with minute extremely deciduous placoid scales, the spine of
each scale with a stelliform base.
Dorsal spines very strong and acute, the 2nd nearly twice the size of the
1st. The 1st dorsal fin arises in advance of a point midway between the pec-
torals and ventrals, the 2nd arises immediately behind the level of the base of
the ventrals.
Colours uniform jet-black, but the integument is very deciduous. In youn^
specimens the hue is " deep violet black, lighter between the eyes ; head with
minute white spots arranged in the shape of a lute ; ventrals with pale tips."
Bay of Bengal 405-285 fathoms : Arabian Sea 690-620 fathoms.
Regd. Nos. 11664, 11666 : ~ to p
Family Scylliidce.
SCTLLIUM, CUV., M. & H.
3. Scyllium hispidum, Alcock.
Scullium hispidum, Alcock, Ann. Mag. Nat. Hiat., July, 1891, p. 21 : Illustrations of the Zoology of the
Investigator, Pishes, pl. VIII. figs. 3, 3a.
Head broad and depressed. Snout flat, semicircular in outline, its length,
measured from the convexity of the upper jaw, is half its greatest breadth and
twice the width of the inter-narial space. Nasal valves separated by an interval
almost equal to the maximum diameter of the nostril : each has a very short
cirrus.
Eyeballs large, their major diameter being between two-thirds and three-
quarters the length of the snout (measured as above).
16
Spiracles very small, their diameter less than half that of the pupil, situated
immediately behind and below the eye.
Mouth large crescentic, a short labial fold at the angles only. Teeth in
broadish bands in both jaws, small, mostly tricuspid (sometimes 5-cuspid) the
middle cusp much the largest. Surface of palate and of tongue papillated.
The entire surface of the skin closely covered with minute stony tricuspid
spines.
The first dorsal fin, which arises just ahead of the vertical through the
after end of the base of the ventrals, is larger than the second, which is about
opposite the anal : the base of the anal is about twice as long as that of either
dorsal. The pectorals are at least twice as big as the ventrals, and the latter
have a very oblique posterior margin.
Colours : dull stone-grey, rather lighter ventrally.
Andaman Sea, 188-220 fathoms, 185 fathoms, 370-419 fathoms, and 405
fathoms.
The largest specimen is a female 11 inches long.
Regd. Nos. 13120 type of male: 1J* type of female: 2ffi: a38_JLai.
382 385.
1 - 1 •
4. Scyllium canescens, Giinther.
Scyllium canescens, Giinther, Ann. Mag. Nat. Hist. (5) II. 187S, p. 18 : Challenger Deep Sea Fishes, p. 1, pi. i.,
fig. A : Alcock, J. A. S. B., Vol. LX V., pt. 2, 1896, p. 310.
Differs from 8. hispidum, if my identification be correct, in the following
particulars : —
(1) the snout is slightly longer, relatively :
(2) the eye is slightly smaller :
(3) the labial folds at the angles of the mouth are slightly larger :
(4) teeth tricuspid, the cusps subequal :
(5) the body is covered with simple stiff prickles :
(6) the fins have much the same position, but the second dorsal is slightly
larger than the first, and the base of the anal is not so extensive.
Colours in spirit blackish with a hoary gray surface. Some of the fins, as
in some specimens of Scyllium hispidum, may be tipped with white behind.
A single small specimen from the Arabian Sea, 690-620 fathoms.
Regd. No. ^8.
[ The type of this species was dredged by the Challenger, off the south-
western coast of S. America at a depth of 400 fathoms. ]
Distribution. Patagonian fjords : Arabian Sea : at considerable depths.
17
5. Scyllium quagga, n. sp,
Illcstbations op the Zoology op the Investigator, Fishes, pl. XXVII. fig. 1.
Head broad, depressed. Snout flat, elliptical in outline with a bluntly acu-
minate tip ; its length, measured from the convexity of the upper jaw is half its
greatest breadth.
Nasal valves separated by an interval the width of which is rather more
than two- fifths the length of the snout : each with a very short and inconspic-
uous cirrus.
Eyes large, their major diameter rather over two-thirds the length of the
snout.
Spiracles small, their diameter about equal to that of the pupil ; situated
immediately behind and below the eye.
Mouth large, crescentic ; a short labial fold at the angles only. Teeth in
both jaws tricuspid, the middle cusp much the longest and most acute.
Body everywhere covered with minute scales, which are tricuspid or anchor-
shaped, except on the throat and belly, where they are granular.
The dorsal fins are not very unequal : the first arises just in front of the
vertical through the after end of the base of the ventrals : the second arises
just in advance of the vertical through the after end of the anal.
Base of anal a little longer than that of either of the dorsals, between a
half and two-thirds the length of the interval between itself and the caudal.
Posterior margin of ventrals very oblique.
Colours in spirit : very numerous well-defined alternate cross-bands of light
and very dark brown, from snout to tip of tail, not passing on to ventral
surface : the cross-bands are rather irregular in breadth, but the dark ones are
usually the narrowest. The dark cross-bands are not broken up into spots, but
are continuous stripes, and the light cross-bands are unspotted.
An apparently adult male is 11 inches long.
Off Malabar coast, 102 fathoms.
Regd. No. ~.
This species comes nearest to 8. Burgeri M. & H.
Suborder Batoidei.
Family Torpedinidce.
Benthobatis, Alcock.
Benthobatis, Alcock, Ann. Mag. Nat. Hist. August, 1898, p. 144.
The whole animal invested in a loose, soft, naked, glandular skin.
18
Tail very distinct from the oval disk and vrithout distinct lateral folds. Two
dorsal fins on the tail ; caudal fin well developed.
Nasal valves confluent into a quadrangular curtain.
Teeth flat, rhomboidal, with the posterior angle acutely produced.
Spiracles large, situated immediately behind the inconspicuous and quite
rudimentary eyes.
A large electric organ between the head and either pectoral fin.
This curious blind torpedo differs from all other members of the family in
having the eyes quite rudimentary and in the absence of lateral folds of skin on
the tail. In general form it resembles Narcine, though the disk is not quite so
broad. The teeth, like those of Discopyge, have the posterior angle produced,
but more acutely than in Discopyge, judging from Tschudi's figure.
6. Benthobatis Moresbyi, Alcock.
Benthobatis Moresbyi, Alcock, Ann. Mag. Nat. Hiet. August, 1898, p. 145 : Illustrations of the Zoology of
the Investigator, Fishes, pl. XXVI. fig. 1.
The disk is oval, its long axis, which is fore and aft, is a little shorter than
the tail : more than a third of its extent is pre-oral.
The eyes are represented by two small unpigmented spots, each not much
bigger than the head of a pin, situated one in front of either spiracle ; a slender
optic nerve passes to each, and expands into a vesicle so small as to need a lens
for its recognition.
The mouth is small and protractile ; the teeth are small rhomboidal plates,
with the posterior angle strongly and acutely produced, and are arranged in
mosaic in about ten very oblique series in either jaw.
The gill-openings are large and well-spaced : the posterior one is nearer to
the cloacal opening than to the mouth.
The dorsal fins are placed rather close together, the first being a little in
front of the posterior limit of the ventrals, and, like the caudal, are thick and
fleshy. The ventrals are of the usual shape ; though they are separate, the skin
between them is loose and copious. The whole animal is covered with a per-
fectly smooth, soft, glandular, purplish-black skin ; scattered on the disk and
round the edge of it are some small white pores, not much smaller than the
eyes ; in one specimen part of the tips of the second dorsal and caudal fins are
white.
Two males (each about 14- inches long) and a young one, from off the
Travancore coast, 430 fath.
-r, -■ AT 232 234
Regd. JNos. -j- : — .
19
Named in memory of Capt. Moresby, of the Indian Navy, whose surveys
(1834-38) in the seas where this curious fish is found are known to all readers
of Darwin's ' Coral-Reefs.'
Family Rajidce.
Raja, (Artedi) Cuvier.
Raja, Cuvier, Regne Animal, Poissons, p. 375.
Raja and Vraptera, Miiller and Henle, Plagiostomen, pp. 132, 155.
Raja, Giinther Catalogue of Fishes, VIII. 455.
Raja, Jordan and Evermann, Fishes of North and Middle America (Bull. U. S. Nat. Mus. No. 47, 1896)
I. p. 66 : Goode and Bean, Oceanic Ichthyology, p. 24.
" Tail very distinct from the disk, which is of rhombic shape, with a fold on
" each side. Body generally rough or with spines, rarely entirely smooth. Two
" dorsal fins, on the tail, without spine. Tail with a rudimentary caudal fin, or
" without caudal. Each ventral fin divided into two by a deep notch. Nasal
" valves separated in the middle, where they are without a free margin. Teeth
" obtuse or pointed. Pectoral fins not extending forward to the extremity of the
" snout. Sexes differing in the form of the teeth and in the dorsal spines."
(Giinther).
Synopsis of the Indian species of Raja.
I. Snout short, slightly exsert : upper surface of disk covered with
prickles, in addition to definitely placed spines : a single row of
large spines on middle line of back and tail : dorsal fins very close
together ... ... ... ... ... R. maniillidens.
II. Snout much exsert : upper surface of disk with definitely placed
spines, but with few or no prickles : dorsal fins well separated : —
1. Snout, measured from mouth, about two-sevenths the
greatest breadth of the disk in length : a short row of spines
in the middle line of the nape, and several rows on the tail ... R. Powelli.
2. Snout, measured from mouth, about a third the greatest
breadth of the disk in length : a single spine in the middle line
of the nape: a single row of spines on the tail ... ... R. Johannis-Davisi.
7. Maja maniillidens, Alcock.
Raja mamillidens, Alcock, Ann. Mag. Nat. Hist. November 1889, p. 380: Illustrations op the Zoology of
the Investigator, Fishes, plate VIII, fig. 1.
The disk, including the ventral fins, is about half the total length and its
breadth is about the same : it is rhomboidal in shape, with the angles rounded :
the anterior margin of the pectoral fin is slightly and broadly sinuous.
The snout is short, broad, and only slightly exsert, its length, measured
from the mouth or from the eye, is not quite a fourth the greatest breadth
of the disk, and is a little more than the distance between the outer margins of
the nostrils.
20
Major diameter of orbit equal to the width of the inter-orbital space and
much more than that of the spiracle.
Mouth crescentic. Teeth, in the female, having a globular base and a
mamillary point : in twenty-four oblique rows in the upper and eighteen in the
lower jaw.
The whole of the upper surface of the disk (including the ventral fins) and
tail, and all the surfaces of the posterior half of the tail (including the dorsal
and rudimentary caudal fins) are covered with small sharp close-set prickles :
under surface of disk smooth and glandular.
A large spine at either angle of either orbit, and a pair of spines between
the spiracles : one or two on each shoulder girdle, and a single row down the
middle of the back from the occiput nearly to the first dorsal fin.
Dorsal fins adjacent but separate, the posterior the larger.
Colour in life — uniform jet black : in spirit, dark chocolate.
A single female specimen, 11^ inches long, from the Gulf of Manar, 597
fathoms.
Regd. No. 11769.
Apparently allied to the Mediterranean Raja atrata, M. & H.
8. Raja JPowelli, Alcock.
Raja Powelli, Alcock, Ann. Mag. Nat. Hist., August, 1898, p. 145 : Illustrations of the Zoology of the
Investigator, Fishes, pl. XXVI. fig. 4.
The disk without the ventral fins is nearly half the total length, and its
breadth is equal to its length with the ventrals : it is rhomboidal with the angles,
except the rostral, rounded : the anterior margin of the pectoral fin is broadly
sinuous.
The snout is rather slender and much exsert : its length, measured from the
mouth, is two-sevenths the greatest breadth of the disk and is nearly half again
as much as the distance between the outer margins of the nostrils.
Major diameter of orbit hardly equal to the width of the inter-orbital space,
but considerably more than that of the spiracle.
Teeth in the female obtusely pointed or obscurely tricuspid ; in about
55 transverse rows in either jaw. Mouth straight.
Both surfaces of the disk are smooth, except for some prickles near the edge
of the snout and the edge of the anterior half of the pectoral fins.
Two or three spines on the anterior edge of the orbit and one near either
postorbital angle : three in the middle line of the nape. Two or three series
of spines extend from the hinder fourth of the disk to the first dorsal fin, there
21
is a single short series between the dorsal fins, and the sides of the tail from end
to end are thorny.
The distance between the two dorsal fins is greater than the length of
either : the first dorsal is the higher, the second, which is confluent with the
rudimentary caudal, is the longer.
Colours : upper surface warm brown with a pair of large ocelli behind the
shoulder girdle : under surface dirty white.
A single female 12^ inches long from the Gulf of Martaban 67 fathoms.
Regd. No. 2-f.
Apparently allied to the Kerguelen Raja murrayi, Gthr.
Named after Lieutenant Powell of the Indian Navy : a colleague, in the old
Marine Survey branch of the service, of Captain Moresby.
9. Maja Johannis-Davisi, n. sp.
Illustrations of the Zoology of the Investigator, pl. XXVII. fig. 2.
The disk without the ventral fins is nearly half the total length, and its
breadth is much more than its length with the ventrals : it is rhomboidal with
the angles, except the rostral, rounded, and with the two anterior sides much
longer than the other two : the anterior margin of the pectoral fin is broadly
sinuous.
Snout slender and much exsert : its length, measured from the mouth, is
close on a third the greatest breadth of the singularly broad disk and is If times
the distance between the outer margins of the nostrils.
Major diameter of orbit equal to the width of the interorbital space and
nearly twice that of the spiracle.
Mouth straight : teeth obtusely pointed (male) in about 32 very oblique
rows in the upper, and about 30 in the lower jaw.
Both surfaces of disk smooth, except for some star-shaped prickles on the
ventral surface of the rostral cartilage and of the edges of the snout and adja-
cent part of the pectoral fins.
Two strong spines on the anterior margin and one at the posterior angle of
either orbit : a very strong spine in the middle of the nape. An eminence but
not a distinct spine on either side of shoulder and pelvic girdles. Tail smooth
except for a mid-dorsal row of large spines which extend from its base to the
second dorsal fin.
The distance between the two dorsal fins is rather more than half the length
of the base of either : the two fins are of about equal size and the second is
confluent with the caudal.
Colours : smoky black above, black mottled with white below.
22
A single male 8|- inches long from off the Travancore coast 224-284
fathoms.
Regd. No. y'
Near the Mediterranean Raja oxyrhynchus L. Named after the celebrated
Elizabethan navigator and explorer John Davis, who — though best known for
his Arctic voyages — piloted three expeditions to the East Indies and lost his life
in Indian seas.
Besides the Sharks and Rays just described, there is evidence of the
existence of other species in the depths of these seas : —
(1) Off the Travancore coast, in 824 fathoms, an egg, with a very early
embryo, of some gigantic species of, probably, Raja was dredged.
(2) Off the western coast of the Andamans, in 561 fathoms, an empty
egg-capsule of, probably, Gallorhynchus was taken.
(3) Off the Godavari Delta, in 410 fathoms, an empty egg-capsule of,
probably, Chimsera was obtained.
Sub-class TELEOSTOMI.
Order Teleostei.
Sub-order Acanthopterygii.
Family Serranidce.
Chelidopeeca, Boulenger.
Chelidoperca, Boulenger, Catalogue of the Pereiform Fishes in the British Museum, I. p. 304.
" Body compressed ; scales rather large, rough and ciliated. Lateral line
" complete, the tube with ascending tubule. Mouth large, protractile ; maxil-
" lary exposed ; jaws with bands of villiform teeth, a few of the inner ones in
" the middle of the upper jaw enlarged, depressible, hinged at the base ; teeth
" on vomer and palatines ; tongue smooth. Head partly scaled ; pre-opercle
" serrated, without antrorse teeth on the lower border ; opercle with two spines.
" Gill-membranes separate ; seven branchiostegals ; pseudobranchiae present ;
" gill-rakers rather long. Dorsal fins confluent, with X. 10 rays, the spinous
" and soft portions subequal in length. Anal short with III. 6 rays. Pectorals
" subsymmetrical, obtusely pointed, (with 17 rays).* Ventrals anterior to base
" of pectorals, close together, with a strong spine. Posterior process of
" premaxillaries not extending to the frontals ; supra-occipital and parietal crests
" short, smooth area of cranium extending posteriorly to a line connecting the
" preopercular portions" (Boulenger).
* lu the single Indian species the pectoral flu rays are 1-1 or 15 in number.
23
Centropristis investigatoris, Alcock, Ann. Mag. Nat. Hist., September, 1890, p. 199 : Illustrations op thi
SOOLOGT OF THE INVESTIGATOR, FlSHES, PL. X. FIG. 1.
10. Chelidoperca investigatoris, (Alcock).
investigatoris, Alcock, Ann. Mag. Nat. Hist., Septembei
Investigator, Fishes, pl. X. fig. 1.
Chelidoperca investigatoris, Bonlenger, Cat. Perciform Fishes, I. p. 305.
B. 7. D. X. 10. A. III. 6. L. lat. 42. L. tr f .
10
Dorsal and ventral profiles quite symmetrical.
Height of the body between 3^ and 3f , length of the head, from the tip of
the lower jaw to the tip of the operculum about 2jt, in the total, without
caudal.
Head inclined to depression in its anterior half, deep, broad, and inflated in
its branchial region, with the operculum prolonged ; scaly, except on the snout
and upper jaw.
Snout depressed, rounded ; its tip formed by a prominent median knob on
the projecting lower jaw ; its extreme length (including the mandibular element)
is equal to the major diameter of the eye and is less than its breadth.
Eyes in their long diameter 4§ in the head-length ; the upper border of the
orbit enters the dorsal profile ; the breadth of the interorbital space is one-third
the length of the eye. Nostrils superior.
Mouth wide, oblique ; jaws strong, the maxilla reaches the vertical through
the posterior border of the orbit, the mandible closes outside the maxilla ; teeth
in villiform bands in the premaxilla and palatines and in a small patch on the
vomer ; small canines in the mandible and at the maxillary symphysis ; tongue
long and spathulate.
Gill-opening very wide ; operculum with two flat spines ; preopercular
border rounded and serrated throughout ; sub- and interoperculum large ; pseudo-
branchias coarse ; gill-rakers tuberculate.
Scales, except on the lateral line and in the row flanking the dorsal fin,
large ; finely ctenoid, except on the operculum ; eight series on the cheek.
Lateral line salient, with very small scales.
One dorsal, with its spinous and soft portions of equal extent, the fourth
and fifth spines the greatest and one-fourth longer than the eye ; the rays
slightly increasing in length to the ninth, which is less than two-thirds of the
maximum body-height and shorter than the corresponding anal ray. Caudal
emarginate, with the upper lobe the longer, its basal half scaly ; its length is
about equal to that of the pectoral, which is rather longer than the postorbital
portion of the head. Ventrals subjugular, the second ray almost as long as the
pectoral fin.
Pyloric caeca few. Air-bladder small.
24
Colours in life : — Head and body bright pink, belly and throat white; a
broad bright yellow band passes from the tip of the snout through the eye to the
caudal fin ; indefinite bright yellow markings on the cheeks, opercles, and fins.
In spirit, faded yellow, with four incomplete cross bands of grey.
Total length 5| inches.
Two specimens from off the Ganjam coast, 98-102 fathoms.
Regd. No. 12820, 12821.
Synageops, Giinther.
Synagrops, Giinther, Challenger Deep-Sea Fishes, p. 16.
ilelanostoma, Doderlein, Denk. Akad. Wien, XLVIII. 1884, p. 5 (name pre-occupied).
Parascombrops, Alcock, Jonrn. As. Soc. Bengal, Vol. LVIII. pt. 2, 1889, p. 296.
Eypoclydonia, Goode & Bean, Oceanic Ichthyology, p. 236.
Body rather elongate, covered with large, thin, deciduous, cycloid scales.
Muciferous cavities of the vertex of the head well developed. Seven branchios-
tegals : pseudobranchias present : edge of preoperculum more or less serrated :
operculum with two weak points. Villiform teeth in (usually) narrow bands on
the jaws vomer and palatines, with the addition of a pair of large canines in
the upper jaw and an irregular row of canines in the lower jaw. Two
separate dorsal fins, the first with nine slender spines : anal fin with two spines.
An air-bladder. Pyloric caeca in small number.
This genus may probably prove to be identical with Acropoma Temm. and
Schleg.
11. Synagrops philippinensis, (Giinther).
Acropoma philippinense, Giinther, Challenger Shore Fishes, p. 51 : Alcock, Jonrn. As. Soc. Bengal, Vol. LXII
pt. 2, 1894, p. 116.
Parascombrops pellucida, Alcock, Jonrn. As. Soc. Bengal, Vol. LVIII. pt. 2, 1889, p. 296, pi. xxii. fig. 1.
? Melanostoma argyreum, Gilbert and Cramer, Proc. U. S. Nat. Mns. Vol. XIX. 1896, p. 416, pi. xxxix. fig. 3.
Synagrops philippinensis, Illustrations of the Zoology of the Investigator, Fishes, pl. XXVIII. fig. 1.
B. 7. D. IX. 1-9. A. II. 7. L. lat. circ. 28.
Body compressed, its greatest height between 3| and 3^ in the total without
the caudal.
Head, measured from the tip of the underhung lower jaw to the tip of the
semimembranous prolongation of the opercle, 2^ in the total without the caudal,
compressed, the muciferous cavities of its vertex well developed and bounded by
numerous low sharp crests, most of which are oblique : in the skin covering
the vertex of the head numerous tiny scales are embedded and almost concealed.
Snout short, a large part of it is formed by the prominent lower jaw : its
length, including the mandibular element, is less than that of the eye.
25
Eye of good size, its major diameter is equal to the depth of the caudal
peduncle or to two-sevenths of the greatest body height, and is rather more than
the greatest width of the inter-orbital space.
Mouth-cleft wide, very oblique ; the maxilla reaching a little behind the
middle of the pupil ; the mandibles peculiarly emarginate on either side of the
symphysis. Villiform teeth on premaxilla?, vomer, palatines and anterior part
of lower jaw ; in addition there is a pair of large canines at the symphysis of the
upper jaw and a row of irregular (large and small) canines in the lower jaw,
one of them being at the symphysis.
Gill-cleft wide : numerous gill-rakers, the longest of which are nearly two-
thirds the major diameter of the eye, on the first branchial arch. Pseudo-
branchiae large.
Preoperculum with a double border, the angle and the lower limb of the
outer border strongly serrated, the angle of the inner border with three small
spines. Operculum naked, with two weak points. Sub-operculum much pro-
longed behind the angle of the operculum.
Scales cycloid, large, very thin, extremely deciduous.
The dorsal fins are separated by an interspace equal to three-fourths the
length of the eye : the spinous is considerably the higher : the 1st spine is small,
the 3rd is the longest and is equal to two-thirds of the body height, the 4th is
nearly as long. The 1st anal spine is short, the 2nd as long as the eye. Caudal
forked almost to its base, forming two distinct lobes. Pectoral delicate; its
length equals the distance from the tip of the snout to the anterior pre-opercular
edge. Ventral long, reaching two-thirds of the distance to the anal : its spine
is long, and has its outer edge closely sharply and evenly serrated.
Colours in life, transparent light brown suffused with pink from the blood-
vessels : opercular and visceral regions like burnished silver.
The intestine is long and much coiled ; and there are five very large pyloric
caeca.
A small fish : adults of both sexes measure from 3 to 4 inches.
Common in the Bay of Bengal between 60 and 102 fathoms, also found
between 145 and 250 fathoms.
Regd. Nos. 11829: 12437: 12720: 12723 a-j : 12856 a-b : 12857 a-q :
13503-13508, 13510.
I cannot see any difference between this and the species described and
figured by Gilbert and Cramer under the name of Melanostoma argyreum.
Distribution. Bay of Bengal : East Indian Archipelago : (Hawaii).
4
26
Brephostoma, Alcock.
Brephostoma, Alcock, Ann. Mag. Nat. Hist., Nov., 1889, p. 383, and September, 1890, p. 201.
Head-bones and opercles unarmed; preoperculum with a double edge.
Mouth edentulous. Eyes large. Two separate dorsal fins, the first with five
spines. Anal fin with one spine and similar to second dorsal. Scales large,
adherent, ctenoid. Seven branchiostegals. Pseudobranchia? present. Pyloric
caeca in moderate number. No air-bladder.
Its nearest relative is Pomatomus.
12. Brephostoma Carjpentcvi, Alcock.
Brephostoma carpenteri, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 383, and Sept. 1890, p. 201, pi. ix. fig. 4:
Illustrations op the Zoology op the R. I. M. S. Investigatoe, Fishes, pl. XVIII. fig. 2.
2
B. 7. D. V. I. 10. A. I. 9. L. lat. 32-36. L. tr. T.
9
Body elongate, compressed, its greatest height slightly over one-fourth the
total length without the caudal.
Head one-third the total length without the caudal, for the most part
covered with somewhat deciduous scales.
Snout broadish and somewhat depressed, its tip formed by the symphysis
of the lower jaw : its length, including the mandibular element, is about two-
thirds the major diameter of the eye.
Eyes large, their major diameter more than a third the length of the head
and about twice the breadth of the inter-orbital space : supra-orbital margin in
the dorsal profile.
Mouth oblique ; the upper jaw, which is much concealed by the extensive
pre-orbitals, hardly reaching to the pupil ; the lower jaw, the rami of which are
very broad, closes inside the upper jaw except at the symphysis. No teeth.
Tongue free, broad.
Gill-opening very wide, the bones of the gill-cover, like those of the face
and jaws, thin and weak — almost membranous, quite unarmed ; the preopercu-
lum with a double edge. Gill-rakers of the outer side of the first arch long,
close-set, — the longest being nearly half the length of the eye. Pseudobranchiae
large.
Scales strong, thick, very adherent, imbricate, the exposed surface studded
with spines as in most species of Macrurus. The scales of the head, though
otherwise similar to those on the body, are somewhat deciduous. Lateral line
in the form of simple tubes with large round patent orifice : it extends half-way
along the caudal fin.
27
Dorsal fins separated by a snout-length, the second much the higher ; the
first has five stout sharp spines, the three anterior a little longer than the snout ;
the second has one short spine and ten branched rays, and is invested at its base
with scales. Anal with one spine and nine branched rays, situated opposite the
second dorsal, and similar to it in every respect. Caudal short, forked ; its
proximal half scaly. Pectorals well developed, as long as the head without the
snout. Ventrals thoracic, with one spine and five rays with scaly bases.
No air-bladder. Long pyloric cseca in moderate number. Colours in life,
uniform jet-black.
A single specimen, 4 inches long, from Carpenter's Ridge (Lat. 6° 18' to
16' N., long. 90° 40' to 44' B.) 1370 to 1520 fathoms.
Regd. No. 12472.
Named after Captain Alfred Carpenter, R.N., D.S.O., Superintendent of
the Indian Marine Survey from 1884 to 1889, who practically initiated, and who
greatly developed, the deep-sea dredging operations of the " Investigator."
Family Seorpcenidce.
Sebastes, Cuv. & Val.
13. Sebastes hexanema, Giinther.
Sebastes hexanema, Giinther, Challenger Shore Fishes, pi. xvii. fig. B, and Challenger Deep-Sea Fishes, p. 18 :
Alcock, Ann. Mag. Nat. Hist., July 1891, p. 23.
D. XI. 1-9. A. III-5. Sc. circ. 53.
Height of the body about one-third, length of the head nearly half, the
total length without the caudal.
Snout produced, its length is not quite equal to the major diameter of the
eye, which is about two-sevenths the length of the head, and more than twice
the width of the inter-orbital space (in young specimens).
Vertex of head with prominent spines and small scales and with three pairs
of simple tentacles of good length— one at the anterior angles of the orbits,
one above the middle of the eyes, and a third on the nape. (There are other
small tentacles along the preopercular stay and along the lateral line, but they
are liable to be lost by abrasion).
Mouth-cleft wide, the maxilla reaching nearly to the middle of the eye.
Villiform teeth in the jaws, in a narrow band on the palatines, and in a V-shaped
patch on the vomer. Tongue free, pointed.
The 3rd and 4th dorsal spines are the longest, about a third the length of
the head, and shorter than the 2nd of the anal. Pectoral fin reaching to the
anal.
28
Rose-coloured in life with indistinct dusky patches on the back and one on
the first dorsal fin : these fade away in spirit.
Two specimens, the largest just over 3 inches long, from the Andaman
Sea, 188-220 fathoms. They have been compared with one of the Challenger
duplicates.
Regd. Nos. 13031, 13032.
Distribution : Andaman Sea : East Indian Archipelago.
Setaeches, Johnson.
Setarches, Johnson, Proo. Zool. Soo. 1862, p. 177 : Gunther, Challenger Deep-Sea Fishes, p. 19 : Goode and
Bean, Oceanic Ichthyology, p. 262 : Jordan and Evermann, Fishes of North and Middle America, II. p. 1860.
Bathysehastes, Steindachner and DSderlein, Denk. Ak. Wien, XLIX. 1885, p. 207.
? Lioscorpius, Gunther, Challenger Shore Fishes, p. 40, and Challenger Deep-Sea Fishes, p. 20.
Head and body compressed : the vertex of the head with the muciferous
cavities well developed but with few ridges, and those low and indistinct, and
with no erect spines : occiput naked, without a groove. Preorbital and preoper-
culum armed with spines, operculum with two spines. Seven branchiostegals.
Pseudobranchias present. Bands of villiform teeth on the jaws vomer and
palatines.
Body covered with very small cycloid scales. Lateral line very wide, naked.
Vertical fins not elongate. Dorsal fins separate but in contact, the first usually
with eleven spines. Pectorals long and large, without separate appendages.
Pyloric appendages few.
14. Setarches Gunthei'i, Johnson.
Setarches Giintheri, Johnson, Proc. Zool. Soc, 1862. p. 177, pi. xxiii : Vaillant, Exp. Sci. Travailleur et Talisman
Poissons, p. 373 : Goode and Bean, Oceanic Ichthyology, p. 263.
Lioscorpius longiceps, Alcock, Ann. Mag. Nat. Hist., July 1891, p. 23 : Illustbations op the Zoology of the
Investigator, Fishes, PL. X. pig. 3. [It seems to me that Lioscorpius longiceps Gunther, Challenger Shore Fishes,
p. 40, pi. xvii. fig. C is little different from this species, for though the short description does not correspond the
figure does.]
Scorpxna remigera, Gilbert and Cramer, Proc. TJ. S. Nat. Mas. Vol. XIX. 1896, p. 418, pi. xl.
B. 7. D. IX. 1-10. A. III-5.
Height of the body from - to ^, length of the head about - the total length
without the caudal.
m
Head singularly large and clumsy looking, with cavernous bones and well-
developed muciferous cavities, scaly on temples cheeks and opercles, naked
elsewhere.
Three free divergent spines on edge of preorbital ; four or five, of which the
three upper are usually large, on the edge of the preoperculum ; two on the
29
operculum. Preopercular stay prominent, with or without a single small spine
near its posterior end. No true spines on the top of the head, but two low
ridges on the occiput and one behind either orbit end somewhat acutely, as also
does a short ridge at the anterior supra-orbital angle and one above the middle
of the orbit.
The snout (including the sharp knob at the mandibular symphysis which,
fitting into a notch in the pre-maxillary, forms its tip) is about 3^ in the length
of the head. Nostrils of good size, a fleshy papilla above the anterior one (this
is liable to loss by abrasion).
Major diameter of eye from - to - the length of the head : interorbital space
a little wider than the eye, with three very characteristic mucous fossa?, one
large and elliptical in the middle line posteriorly, two smaller and oval, side by
side, anteriorly.
Mouth large, oblique, with prominent lower jaw ; the maxilla reaches as far
as the posterior border of the orbit. Villiform teeth on jaws vomer and palatines.
Tongue ending in a small free spathulate tip.
Gill-opening wide. Pseudobranchia? rather large. Gill-rakers on outer
side of first arch rather short and distant.
Body covered with tiny cycloid scales. Lateral line broad, naked, with
double tubule.
3rd, 4th and 5th dorsal spines the longest, the 3rd about as long as the
snout, and not quite as long as the 3rd anal spine, which is the longest of that
fin. Pectorals very large, reaching beyond the beginning of the anal, consisting
of 21 or 22 rays. Ventrals reaching rather more than halfway to anal.
Colours red with minute black specks, and with some dusky markings that
usually fade.
Most of our specimens come up with everted stomach and dislocated viscera,
so that dissection is difficult. In two specimens I have counted 4 large pyloric
ca3ca, and in one there was a small thick- walled air-bladder.
Andaman Sea 185 and 188-220 fathoms : Bay of Bengal, off Trincomali,
320-296 fathoms.
Largest specimen nearly 5$ inches long.
Distribution: Madeira and C. Verde Is., Bay of Bengal, Andaman Sea,
(East Indian Archipelago?), Hawaii: in moderate depths (up to about 300
fathoms).
Regd. Nos. 13036, 14131-14133, ™ ~^-
30
Pterois, Cuv.
15. Pterois macrurus, Alcock.
Pterois macrurus, Alcook, Journ. As. Soc. Bengal, Vol. LXV. pt. 2, 1896, p. 303 r Illustrations of the
Zoology of the Investigator, Fishes, pl. XVIII. fig. 4.
D. XIII. 9. A. II. 7.
Length of the head about two-fifths, height of the body about one-third of
the total without the caudal.
Snout deep, about as long as the eye, which is about one-fourth the length
of the head. Supra-orbital ridges high, overhanging the deeply concave inter-
orbital space, the width of which is about half the length of the eye. Pre-
orbital with frill-like crests arranged in a star, and with a large tentacle over-
hanging the angle of the mouth. Crests of the head and cheek fairly well
developed, serrated.
Scales finely ctenoid: in from 40 to 45 rows between the temple and the
base of the caudal. The scales cover all parts of the head except the jaws,
throat, and tip of the snout, and the middle line of the narrow interorbital
space.
The pectorals reach to the base of the caudal : the ventrals reach just
beyond the beginning of the anal. The caudal is pointed, one at least of its
upper rays is produced as a slender filament which is as long as the body behind
the eye : the rays below this gradually decrease in length.
Pseudobranchias singularly large. Three large pyloric caeca.
Colours in spirit : sepia with black cross-bars : vertical fins with dark spots
which are distinct only along the upper edge of the caudal: pectorals and
ventrals nearly black, with white spots.
Seven inches from tip of snout to tip of caudal filament.
Two specimens from off the Malabar coast, 68-148 fathoms ; (six specimens
from 45 fathoms).
Regd. Nos. 13823 to 13828 : ~-
Minous, Cuv. & Val.
16. Minous inermis, Alcock.
Minous inermis, Alcock, Journ. As. Soc. Bengal, Vol. LVIII. 1889, pt. 2, p. 299, pl. xxii. fig. 4: Ann. Mao-. Nat.
Hist., Sept. 1892, pp. 207-214: Journ. As. Soo. Bengal, Vol. LXIII. 1894, pt. 2, p. 116 : Illustrations of the
Zoology of the Investigator, Fishes, pl. XVIII. fig. 3.
B. 7. D. XL 12. A. II. 10. P. 11/1. V. I. 5. C. 13.
Head and body compressed : height of body about a third, length of head
about three-sevenths the total without the caudal.
31
Preorbital with two divergent spines on its edge : preoperculum with a
laro-e and a short spine at its angle and some dentations along its lower edge :
operculum with two weak stays hardly projecting as spines. Preopercular stay
crenate. Top of head eroded : two eroded and crenate ridges on occiput, one or
either postorbital region : upper margin of orbit eroded and crenate.
Snout about as long as the eye, or rather more than a fourth the length of
the head : interorbital space hardly narrower than the eye, traversed by ridges.
Mouth-cleft oblique, the maxilla reaches almost to the anterior margin of
the pupil. A barbel about the middle of the limb of the lower jaw. Villiform
teeth in the jaws and palatines.
Gill-opening wide : the posterior gill-cleft is a small foramen. Gill-rakers
short, somewhat club-shaped.
Integument comparatively thin, without scales : it invests all the fins. The
lateral line shows as 17 to 18 tubular papillae.
All the fin-rays are simple. Dorsal fins continuous, the soft portion being
the higher : all the spines are weak and flexible, the 1st being very small and
the 2nd and 3rd somewhat isolated. Anal spines hidden, the 1st being visible
only on reflecting the skin. Pectoral as long as the head : its free filament
reaches to the 3rd anal ray. Ventral two-thirds or more as long as pectoral.
Colours in fife : — rosy red with white and gray mottlings and minute black
dots ; throat and barbels white ; pectoral, ventral, aud anal fins edged with
black, the posterior surface of the pectoral with small light spots.
Two large pyloric caaca : a small air-bladder.
Largest specimen about 4^ inches long.
Off Coromandel coast, 133, 70, and 60 fathoms. Off Malabar coast, 68-148
and 45 fathoms.
Regd. Nos. 12444, 12445 : 13219 : 13220-13223 : 13511-13514 : ~Jp.
This fish, -wherever found — and it has been taken at five widely-distant stations on both coasts of the penin-
sula is always more or less encrusted with the gymnoblastic Hydroid Stylactis minoi.
In the Annals and Magazine of Natural History for September 1S92 I have given reasons for concluding that
the relation between the Fish and the Hydroid is a definite commensalism, and not accidental or parasitic. Since
that paper was published Minous inermis has twice been taken by the " Investigator," — once off the Madras coast
and once off the Konkan coast — and on both occasions Stylactis minoi was found on all the specimens captured.
For the description and figure of Stylactis minoi see Ann. Mag. Nat. Hist., Sept. 1892, pp. 212, 213.
Family Beryciclce.
When Day's latest volumes on the Fishes of India were published, in the
Fauna of British India series, only two Indian genera of this family were
known ; I therefore give a synopsis of the genera now known to inhabit these
seas.
32
Synopsis of the Indian genera of the family Berycidse.
I. No barbels : eight braneliiostegals : —
1. Scales bony, forming a bony mail : ventrals consisting of a huge
spine and 2 or 3 almost rndimentary rays ... ... Monocentris.
2. Scales ctenoid : anal spines 3 or 4 : —
i. One dorsal : abdomen serrated : 6 soft rays in the ven-
trals :
a. No teeth in the vomer ... ... ... Hoplostethus.
b. Vomerine teeth ... ... ... Trachichthts.
ii. Two dorsals : abdomen not serrated : 7 soft rays in the ven-
trals : —
a. No preopercular spine ... ... ... Myripristis.
b. A preopercular spine ... ... ... Holocentrum.
3. Scales cycloid : anal spines feeble and few (1 or 2) ... Melamphaes.
II. Two barbels: four branchiostegals ... ... ... Poltmixia.
Monocentris, Bl. Schn., Cuv. & Val.
Monocentris, Cuv. & Val., Hist. Nat. Poissona, IV. p. 461 : Giinther, Cat. Fishes. I. p. 8 (ubi synon.).
Body deep, compressed, covered with a mail of large irregular bony scales :
those in the middle line of the belly are large and strongly keeled.
Head large, the superficial bones, with the exception of the operculum, are
sculptured to form wide deep muciferous cavities which are covered by spongy
skin. Eye large. Snout blunt, rounded, overhanging the mouth. Mouth
inferior, large, its cleft little oblique. Villiform teeth on jaws and palatines
only.
Eight branchiostegals : large pseudobranchite : no large projecting spine at
the angle of the preoperculum.
Two dorsal fins, the first consisting of a few large coarse isolated spines.
Caudal emarginate but not deeply forked. Ventrals consisting of a huge spine
and two or three small rays.
An air-bladder : a few large pyloric caeca.
17. Monocentris japonicus (Houtt,), Cuv. & Val.
Monocentris japonicus, Cuv. & Val., Hist. Nat. Poiss. IV. 461, pi. xcvii. : Temra. & Schleg. Faun. Japon., Poiss.
p. 50, pi. xxii. fig. 1 : Giiuther Catalogue of Fishes, vol. I. p. 9 : Bleeker, Atlas Ichthyol. Vol. VIII. pi. ccclvi.
fig. 4.
See also Castelnau, P. L. B. New South Wales, vol. III. 1878, p. 365: Hilgendorf, SB. Ges. nat. Freunde,
Berlin. 1879, p. 22: Macleay, P. L. S. N. S. Wales, vol. V. 1880-81, p. 510: Steindachner and DOderlein, Denk.
Ak. Wien, LXVII. 1883, p. 217 : Nystrom, Bihang Svensk. Vet.-Akad. Handl. XIII. iv. no. 4, 1887, p. 4.
B. 8. D. VI. 11. A. 10. P. 13 or 14. V. I. 2 or 3. L. lat. 16.
Height of the body nearly half the total length with the caudal, length of
the head nearly half the total without the caudal.
33
Snout overhanging the mouth, blunt, rounded, about as long as the eye,
which is about a fourth the length of the head : it, like the broad convex inter-
orbital space and the vertex of the head, is occupied by large deep polygonal
mucous cavities which are covered with thick spongy skin. The same sort of
skin covers the cheek and opercles, with the exception of the operculum which
is naked and coarsely striated. Nostrils large, situated near the lower angle of
the orbit.
Mouth inferior, its cleft very slightly oblique : the maxilla is half as long as
the head and completely overlaps the mandible. Broad bands of hard black
villiform teeth in the jaws and palatines.
Gill-cleft wide. Gill -rakers coarse, those on the outer side of the 1st arch
close set and more than half as long as the eye. Pseudobranchias very large.
The operculum completed behind by a flap of thick skin.
Body covered with great irregular bony plates, many of which are acutely
keeled : there are 15 rows of them round the body at the level of the 2nd dorsal
spine, — the mid-ventral row, which are very strongly keeled, being unpaired.
First dorsal fin in the form of 6 coarse isolated spines, of which the first
four are acute and fluted, and the last two are small truncated and eroded at tip.
The 2nd dorsal and the anal are placed far back on the body. The caudal
peduncle is formed very abruptly, its depth being less than a fifth the greatest
body-height. Ventral spine a great fluted spike, about five-sixths as long as the
head : it can be securely fixed like a bayonet, at right angles to the body.
Colours in spirit greenish-yellow, the bony dermal plates neatly outlined in
black.
A single specimen, an adult female with ova, a little over six inches long,
from the Gulf of Martaban, 67 fathoms.
Kegd. No. ^.
Distribution : Japanese Seas, Port Jackson, Andaman Sea, Mauritius.
Hoplostethus, Cuv. & Val„ Gthr.
Hoplostethus, Cnv. and Val., Hist. Nat. Poiss. IV. p. 469 : Giinther Cat. Fishes I. p. 9 (ubi synon.) and Chal-
lenger Deep Sea Pishes, p. 20 : Goode and Bean, Oceanic Ichthyology, p. 189 : Jordan and Evermann, Fishes of
North America I. p. 837.
" Body deep, compressed, covered with scales of moderate size and more or
" less distinct ctenoid structure, rather irregularly arranged, those of the lateral
' ' line being the largest.
" Head very large, the superficial bones being deeply sculptured to receive
" wide muciferous cavities which are covered by thin skin only. Mouth very
"wide, oblique; the jaws and palatine bones armed with villiform teeth, the
" vomer being toothless. Eye very large. Eight branchiostegals ; gill-openings
5
34
" very wide, gill-laminae very short. Preoperculum armed with a flat spine.
"Abdomen protected by dermal scutes which form a serrated edge. One
" dorsal fin the anterior rays of which are spinous. Ventrals with six soft rays :
"pectoral symmetrical. Caudal deeply forked. Air-bladder simple. Pyloric
"appendages numerous." Giinther.
18. Hoplostethus mediterraneum, Cuv. & Val.
Hvplostethus mediterraneum, Cuv. & Val. Giinther, Challenger Deep-Sea Fishes, p. 21 (ubi synon.).
See also Vaillant, Exp. Sci. Travailleur et Talisman, Poiss., p. 378 : Giinther, Ann. Mag. Nat. Hist. IV. 1889,
p. 417: CaruB, Prodr. Faun. Medit II. p. 616: Alcock, Jonrn. As. Soc. Bengal, Vol. LXIII. 1894, pt. 2, p. 116 :
Goode and Bean, Oceanic Ichthyology, p. 189, fig. 208 : Jordan and Evermann, Fishes of North America, I. p.
837 : K. Koehler, Campagne du Caudan, Poissons, p. 484.
Illustrations of the Zoology of the Investigator, Fishes, pl. XIV. fig. 3.
B. 8. D. VI. 13. A. III. 9. P. 18. V. I. 6. L. lat. 28-29.
Height of body rather over 2\, length of head 21- in total without caudal.
As in Monocentris the head cheeks and preopercles are occupied by large
deep square and rhomboidal muciferous cavities, which, however, are covered by
a thin skin. A strong flat supra-clavicular spine, and a much stronger one at
the angle of the preopercle.
Snout not overhanging the mouth, its length (including the prominent tip
of the mandibular symphysis) is somewhat less than the diameter of the large
round eye : the latter is about 3^ to 3^ in the length of the head and a little
less than the width of the convex inter-orbital space. Nostrils large, situated
near the upper angle of the orbit.
Mouth-cleft wide, very oblique (nearer the vertical than the horizontal) ;
the lower jaw, though closing within the upper, distinctly prominent ; the upper
jaw nearly two-thirds the length of the head. Villiform teeth in the jaws, and
in a short and narrow band in the palatines.
Gill-cleft wide : some gill-rakers on the outer side of the first arch are more
than three-fourths the length of the eye : large pseudobranchia?. Opercle striated.
Body covered with scales of uneven size, those of the lateral line, to the
number of 28 or 29, being enlarged. On the back and tail, and on the throat
and near the middle line of the belly, the scales are sharply granular, but behind
the gill-opening and on the sides of the belly they are smooth. In the middle
line of the belly the scales, to the number of 11 to 13, are enlarged and strongly
keeled — the keels ending very acutely.
Pectorals large, reaching more than half-way along the anal. Ventrals
reaching two-thirds of the way to the vent.
Colours in spirit, warm brown or plum-colour with a silvery sheen.
35
Six specimens, the largest nearly a foot long, from off Madras, 145-250
fathoms, off Trincomali, 320-296 fathoms, and off Travancore coast 224 to 430
fathoms.
Regd. Nos. 13711 : 14126-14129: ^-6: ™.
Distribution : Atlantic coast of North America, between 11° and 40° N. lat. :
off the European and African shores of the Atlantic from Ireland to Cape Verde :
Mediterranean : Arabian Sea : Bay of Bengal : Japanese Seas. At moderate
depths (about 150 to about 400 fathoms, where the depth has been recorded).
In the Indian Museum there are also specimens from the Caribbean Sea and from
the Gulf of Gascony.
Teachichthys Shaw, Cuv. & Val., Giinther.
Trachichthys, Cuv. & Val., Hist. Nat. Poiss. III. p. 229 : Giinther, Cat. Fishes, I. p. 10 (ubt synon.), and
Challenger Deep-Sea Pishes, p. 21 : Goode and Bean, Oceanic Ichthyology, p. 187.
Trachichthys differs from Hoplostethus only in having villiform teeth on the
vomer. The pyloric caeca (in some species) are less numerous, and there is a
distinct spine on the operculum.
19. Trachichthys Darwinii, Johnson.
Trachichthys Darwinii, Johnson, Proc. Zool. Soc, I860, p. 311, pi. zzxii.
Trachichthys japonicus, Steindaohner & Doderlein, Denk. Ak. Wien, XLVII. 1883, p. 218, pi. ii.
Trachichthys Darwinii, Giinther, Challenger Deep-Sea Fishes, p. 24 (ubi synon.): Alcock J.A.S.B. LXV. 1896,
pt. 2, p. 314 : Goode and Bean, Oceanic Ichthyology, p. 188, fig. 207.
B. 8. D. VIII. 13. A. III. 11. P. 15. V. I. 6. L. lat. 27-30.
Height of body about 2-, length of head about 2- in the total length without
the caudal.
Bones of vertex of head and of sub-orbital ring, but not of any part of the
opercle, sculptured for muciferous cavities that are covered with a harsh skin in
which tiny scales are embedded : the cheeks, the membranous edge of the
operculum, and the middle line of the chin are also scaly. A flat supra-clavicular
spine of no great size : a very strong flat spine at the angle of the preopercle :
a stay, ending in a prominent spine, across the upper part of the operculum,
which bone has a striated surface.
Snout (measured to the tip of the symphysis of the lower jaw) a little longer
than the eye and about equal to the width of the interorbital space. Eye large,
round, its diameter about one-fourth the length of the head. Nostrils very large,
situated near the upper angle of the eye.
Mouth very wide and oblique, approaching the vertical : the lower jaw,
though closing inside the upper, projecting : the upper jaw two-thirds as long as
the head. Villiform teeth in jaws and vomer and in a long narrow band in the
36
palatines : the inner row of teeth in the lower jaw and in the anterior half of
the upper jaw considerably enlarged.
Gill-opening very wide : some of the gill-rakers on the outer side of the
1st arch are about two-thirds as long as the eye : large pseudobranchiae.
Body closely covered with harsh ctenoid scales of irregular size and dispo-
sition. Those of the lateral line, to the number of 27 to 30, are enlarged.
There are from 10 to 12 enlarged, strongly and acutely keeled, abdominal
scutes.
3rd and 4th dorsal spines the longest, a little longer than the eye. The
pectorals do not reach to the vent : the ventrals reach about two-thirds the way
to the vent.
Colours in spirit : frosted, the cheeks and backs cinnamon, the operculum
and gill-membranes dusky.
One specimen, a foot long, off Trincomali, 320-296 fathoms.
Regd. No. 14130.
Distribution : Madeira, Bay of Bengal, Japan.
20. Trachichthys intermedins, Hector.
Trachichthys intermedins, Hector, Trans, arid Proo. New Zealand Ingt. VII. 1874, p. 245, pi. xi : Giinther,
Challenger Deep-Sea Fishes, p. 24, pi. v. fig. D: Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 380.
B. 8. D. VI. 13. A. III. 10-11. P. 18 or 19. V. I. 6. L. lat. 28.
Height of the body nearly half, length of the head two-fifths the total
length without the caudal.
The skin covering the muciferous cavities of the head and cheeks is not
scaly, nor is the middle line of the chin and the membranous edge of the oper-
culum scaly. Supra-clavicular and preopercular spines sharp and prominent :
spine at upper end of operculum not very prominent.
Snout much shorter than the eye : eye from a third to two-sevenths the
length of the head and equal to the width of the interorbital space.
Cleft of the mouth oblique, but not approaching the vertical : none of the
teeth are appreciably enlarged.
Only 9 keeled scutes in the middle line of the belly.
5th and 6th dorsal spines the longest, — a good deal longer than the eye.
The pectorals reach well beyond the middle of the anal : the first two ventral
rays almost reach the vent, the ventral spine is sharply serrated in the basal
half.
Colours in spirit : warm brown, pectorals and ventrals much darker.
37
The characters noticed above are those that distinguish this species from
T. Darivinii.
One specimen, 2f inches long, from the Bay of Bengal, 272 fathoms.
Regd. No. 11723.
Distribution : New Zealand Seas : Bay of Bengal.
Melamphaes, Giinther.
Melamphaes, Giinther, Cat. Fishes V. p. 433, and Challenger Deep-Sea Fishes, p. 26 (ubi synon.): Goode and
Bean, Oceanio Ichthyology, p. 177.
Plectromus, Gill, Proc. U. S. Nat. Mas. VI. 1883, p. 257 : Goode and Bean, Oceanio Ichthyology, p. 178: Jordan
and Evermann, Fishes of North America, I. p. 840.
Scopelogadus, Vaillaut, Exp. Sci. Travailleur et Talisman, Poiss. p. 140 : Goode & Bean, Oceanic Ichthyology,
p. 181.
" Head large and thick, with nearly all the superficial bones modified into
" wide muciferous channels. Cleft of the mouth of moderate width, obliquely
descending backwards, with the jaws nearly equal in front. A narrow band of
" villiform teeth in both the jaws, palate toothless. Eight branchiostegals :
" pseudobranchia? present. No barbels. Opercles not armed. Scales large,
" cycloid, rather irregularly arranged. One dorsal : caudal forked : anal spines
" very feeble : ventrals with seven rays." {Giinther).
Distribution : Atlantic, Indo-Pacific.
21. Melamphaes mizolejris, Giinther.
Scopelus mizolepis, Giinther Ann. Mag. Nat. Hist. 1878, Vol. II. p. 185.
Melamphaes mizolepis, Giinther, Challenger Deep-Sea Fishes, p. 28 : Alcock, Ann. Mag. Nat. Hist. Sept. 1890
p. 201.
B. 8. D. 11.11. A. 1.8. P.14. V. 1.7.
Height of the body about two-sevenths, length of the head about two-fifths
of the total length without the caudal.
Head very thick, with deep muciferous cavities : the bones of the cranium
stout, the other bones thin and weak : snout short.
Eye very small, about one-seventh the length of the head, half that of the
snout, and half the width of the interorbital space.
Mouth cleft wide, somewhat oblique, the maxilla reaching to the middle of
the pupil, the lower jaw slightly prominent. A very narrow band of villiform
teeth in both the jaws.
Gill-opening very wide : the gill-rakers on the outer side of the first arch
are close-set and nearly as long as the eye. Posterior margin of the preoper-
culum subvertical.
88
Scales deciduous very large : [one left on the thorax of the Bingle " Investi-
gator " specimen measured quarter of an inch in its major diameter, the specimen
itself being just over 3 inches long.]
The dorsal fin arises somewhat nearer to the tip of the snout than to the
base of the caudal, and behind the base of the ventral, which is below that of
the pectoral : the last dorsal ray is above the middle of the anal. Pectoral fin
long, reaching nearly to the end of the anal.
Stomach very large : a few very large pyloric caeca.
Colour black.
A single specimen from the Bay of Bengal, off the Ganjam coast, 1310
fathoms.
Regd. No. 12834.
Distribution : East Indian Archipelago : Bay of Bengal : in deep water.
Polymixia, Lowe.
Polymixia, Lowe, Trans. Camb. Phil. Soc, VI. 1838, p. 198 : Gnnther, Cat. Fishes I. p. 16 (et synon.) : Goode
& Bean, Oceanic Ichthyology, p. 243 {ubi synon.) : Jordan & Evermann, Fishes of North America, I. p. 854.
Body compressed, rather elongate. Eye large. Snout short. Mouth-cleft
very slightly oblique : the upper jaw overhanging the lower. Villiform teeth
in jaws, vomer, palatines, and pterygoids.
Two barbels on the throat.
Gill openings wide : only four branchiostegals : gill-cover unarmed, except
that the edge of the preoperculum is finely serrated : pseudobranchiae present.
Scales moderate : ctenoid. One long dorsal fin, beginning with about five
spines. Ventrals with 6 or 7 rays. Caudal forked. Anal with 4 spines.
Delicate pyloric appendages in moderate number. A thin-walled air-bladder
with more or less distinct vestiges of a pneumatic duct.
22. Polymixia nobilis, Lowe.
Polymixia nobilis, Lowe, Gunther Cat. Fishes, I. p. 17, and Challenger Deep Sea Fishes, p. 34, pi. i. fig. B :
ubi synon. (Nemobrama Webbii Val., Polymixia lowei and japonica Gthr., Dinemus venustus Poey) : Alcock, Ann.
Mag. Nat. Hist., Nov., 1889, p. 381, and July, 1891, p. 23 : Goode and Bean, Oceanic Ichthyology, p. 243, fig. 241 :
Jordan and Evermann, Fishes of North America, I. pp. 854, 855 (foot-note).
B. 4. D. V. 30-38. A. III-IV. 14-18. V. I. 6-7. Sc. circ. 50. L. lat. circ. 36.
Height of the body about equal to the length of the head, which is a little
over one-third the total without the caudal.
All parts of the head, except the snout, the suborbital space, the upper
jaw, the middle line of the chin, and the border of the angle of the pre-oper-
culum, are scaly.
39
Snout not much more than half the length of the eye, which is about one-
third that of the head : interorbital space about a fourth of a head-length in
width. Nostrils of good size, placed near the level of the middle of the eye.
Mouth-cleft large : the upper jaw reaches behind the posterior border of
the orbit and so overhangs the lower that its teeth are quite outside the mouth
when closed. Barbels about as long as the head without the operculum.
Gill-opening very wide : gill-rakers on the outer side of the 1st arch nearly
half the length of the eye : pseudobranchige large.
Scales of moderate size, strongly ctenoid : about 36 of those of the lateral
line are perforated.
The dorsal spines gradually increase in length to the last, which is about
half the length of the anterior dorsal rays : the anterior third, or more, of the
soft part of the dorsal is high, the posterior two thirds or less is very low. The
anal spines also gradually increase in length to the last, which is not nearly so
long as the anterior anal rays : also the anterior half of the soft part of the
anal is high, and the posterior half low.
Pectorals about three-fifths, ventrals about two-fifths the length of the
head : the ventral spine and first ventral ray are very intimately adherent.
Colours in spirit silvery, the cheeks nape and back a warmish light-brown :
tip of the elevated part of the dorsal fin black.
Five specimens from the Andaman Sea 185 to 405 fathoms : the largest is
about 7^ inches long.
Regd. Nob. 11725 : 13034: ~: 2f~
Distribution : "West Indies : Madeira, Canaries and S. Atlantic : Mauritius :
Andaman Sea : Japan.
I accept Dr. Giinther's synonomy of the species. There is no doubt that
our specimens are absolutely identical with P. lowei and P. japonica.
Family Kuvtidm.
Bathyclupea, Alcock.
Balhyclupea, Alcock, Ann. Mag. Nat. Hist., Angust, 1891, p. 130: Goode and Bean, Oceanic Ichthyology,
p. 190 : Jordan and Evermann, Fishes of North America, I. p. 834.
Head and body compressed : the head with the mucous cavities well deve-
loped. Mouth cleft oblique, or very oblique, with the lower jaw prominent.
Minute villiform teeth in the jaws, palatines, and vomer. Gill-openings wide :
seven branchiostegals : pseudobranchiae present. Scales cycloid, deciduous :
lateral line nearly straight.
One short dorsal fin, with the spines weak or absent, situated in the
posterior half of the body. Anal long, with one spine. Pectorals large, the
40
upper rays longest. Ventrals small, subjugular, with one spine and five rays.
Caudal forked.
Pyloric appendages in moderate number.
Air-bladder with a persistent pneumatic duct.
Distribution : Andaman Sea, Bay of Bengal : Caribbean Sea.
I was led by a combination of external characters and by the presence
of a persistent pneumatic duct, to place this genus among the Clupeidse ; but
Messrs. Gill and Goode and Bean have quite properly removed it to the
neighbourhood of the Berycidse. They have made it the type of a distinct family
(Bathyclupeidae) ; but I prefer to emphasize its very obvious relations with
Kurtus and Pempheris and to include it in the same family with them.
23. Bathyclupea Hoskynii, Alcock.
Bathyclupea Hoskynii, Alcock, Ann. Mag. Nat. Hist., August, 1891, pp. 131, 132, fig. 4: Illustrations of the
Zoology of the Investigator, Fishes, pl. XXVIII. fig. 2.
B. 7. D. 10. A. 33. P. 29. V. 6. L. lat. circ. 38.
Soft tissues fragile, bones thin.
Head and body compressed ; the height of the latter almost exactly equals
the length of the former, which is one-third the total without the caudal.
The median abdominal line is neither keeled nor serrated. The mucous cavities
of the skull are large.
Snout rectangular, formed in front by the lower jaw, which in repose is
almost vertical ; its length, including the mandibular element, is not quite equal
to the diameter of the large lateral circular eye, which is one-third the length of
the head ; the width of the flat interorbital space is half the diameter of the eye.
Nostrils small, almost superior.
Mouth wide, its cleft nearly vertical : length of the upper jaw two-fifths that
of the head. Villiform teeth in narrow bands in the premaxillse, mandible, and
palatine, and in an inconspicuous V-shaped patch on the vomer. Tongue large,
bilobed.
Gill-cleft very wide, the membranes entirely ununited; all the opercular
bones well-developed, and the horizontal border of the preoperculum sharply
serrated ; four gills ; the middle gill-rakers on the outer side of the first arch
considerably elongated ; pseudobranchige large.
Head naked. Body and nape covered with large cycloid scales, deciduous
everywhere except on the lateral line. In the largest specimen a scale from the
flank measures 10 millim. in the vertical and 7"5 millim. in the antero-posterior
diameter. Each scale of the lateral line has a deep pocket on its inner side
which opens externally by numerous fine pores.
41
The dorsal fin commences almost exactly midway between the tip of the
snout and the tip of the upper lobe of the caudal fin ; the length of its base is
equal to that of the snout ; it is roughly triangular and its height is a fifth
greater than the diameter of the eye. The anal commences about an eye-diameter
in advance of the dorsal and extends to within a very short distance (equal to
three-fourths of an eye-diameter) of the base of the caudal. Caudal forked, its
length is about one-sixth of the total. Pectorals extending a good deal beyond
the origin of the anal. Ventrals very small, in close contact.
Stomach large, with a csecal sac and a bunch of large pyloric appendages.
A large air-bladder with a persistent pneumatic duct much like that of Glupea.
Vertebrae 9 + 22.
Colours silvery grey becoming black along the back.
14 specimens, from the Andaman Sea, 185 fathoms, 188-220 fathoms, and
370-419 fathoms, and from off Madras 145-250 fathoms. The largest specimen,
an adult female, is 8 inches long : the largest adult male is 6^ inches long.
Registered Nos. 13111, 13112, 13114: 13641, 13642: ^, 2-f, 2™ll™}
254 to 256 381
Named after the late Commander R. F. Hoskyn, R.N., who was Superin-
tendent of the Marine Survey of India from October 1889 until his death on
January 27th, 1892. Most of the success of the " Investigator " during those
years was due to his good-natured sympathy, joined to a singular boldness and
originality in handling the trawl.
Family TrlcMuHdCB.
Thyrsites, Cuv. & Val.
Thyrsites, Cut. & Val , Hist. Nat. Poiss. VIII. p. 196 : Giinther, Cat. Fishes, II. p. 350.
" Body rather elongate : cleft of the mouth wide. The first dorsal continu-
" ous, with the spines of moderate length and extending on to the second. Two
" to six spurious fins behind the dorsal and anal. The greater portion of the
" body naked. Several strong teeth in the jaws : teeth on the palatine bones.
" No keel on the tail. Seven branchiostegals. Air-bladder present. Pyloric
" appendages in moderate number." (Giinther).
Subgenus Peomethichthys, Gill.
Promethichthys, Gill, Mem. Ac. Nat. Soi. Vol. VI. 1893, 115, 123: Goode and Bean, Oceanic Ichthyology, p.
20O ; Jordan and Erermann, Fishes of North America, Vol. I. p. 882.
Two finlets behind the dorsal and two behind the anal fin. Ventrals repre-
sented by a pair of small spines. No dagger-shaped spine behind the vent.
Scales very minute or absent. Lateral line undulating or bifurcating below the
front part of the spinous dorsal.
6
42
24. Thy r sites (PromethicJithys) bengalensis, Alcock.
Thyrsites bengalensis, Alcock, Journ. Aa. Soo. Bengal, Vol. LXIII. pt. 2, 1894, p. 117, pi. vi. fig. 1 : Ilt.ustka-
tions of the Zoology of the Investigator, Fishes, pl. XV. fig. 10.
This species may possibly be identical with, the T. prometheoides of Bleeker,
which I know only from the short description in Giinther's Catalogue.
B.7. D.XVIII. ~-4 LI. A.II.11-12, LI. P.14. V.I.
Length of head two-sevenths of the total (caudal included), and twice the
greatest height of the body.
The snout, which has the usual Trichiurid form, is two-fifths of the head in
length, and twice the diameter of the eye. The nostrils are small pores situated
well in front of the eye.
The mouth is large, and the upper jaw-bones are massive : the maxilla
reaches to a point midway between the anterior border of the orbit and the pupil.
There is a single row of distant fang-like teeth in the premaxillary, which in
front, to the number of three or four, are of great size : the mandibular teeth are
similar in size form and arrangement, but only two — the front one on each side —
are enlarged, and these but slightly. There is a single row of small sharp distant
teeth on each palatine. Grill-opening extremely wide. Pseudobranchi'33 large.
The head and body are invested in a thick silvery scaleless skin. The
lateral line bifurcates at the level of the 5th or 6th dorsal spine, the upper
branch running along the base of the dorsal fin, the lower descending with a
curve to the middle line, or a little ventrad of it, and then taking a somewhat
sinuous course to the caudal.
The longest (middle) spines of the long first dorsal fin are two-thirds the
greatest body-height in length : the second dorsal, like the anal, is low and
short : the two spurious finlets are incompletely isolated in both fins.
The caudal is large and deeply forked.
The delicate pectorals are not quite half as long as the head. The ventrals,
which arise close together on the abdominal profile, a little in advance of the
pectorals, are each reduced to a single fluted spine.
In correlation with the strong jaws and large fangs the stomach is huge, its
length being one-third of the total (caudal included). In the specimen dissected
there is a small air-bladder and seven large but delicate pyloric casta.
Colours in spirit : burnished silver, with the mid-dorsal line, from snout to
caudal, blue-black : fins hyaline, the spinous dorsal with a black edge which is
broadest in front, the tips of the lobes of the caudal fin dusky.
Five specimens are in the Indian Museum, from off Madras 145-250
fathoms, The largest specimen is 5^ inches long.
Regd. Nos. 13518, 13520-13522, 13524.
43
Family Carangidce.
Bathyseriola, Alcock.
Bathyseriola, Alcock, Ann. Mag. Nat. Hist., Sept. 1890, p. 202.
Body fusiform but much compressed; edge of the belly sharp, grooved
along the middle line. Scales small, deciduous, cycloid : lateral line unarmed.
One dorsal fin with the spinous portion rather feeble : the soft portion, and the
anal, long. No finlets. Anal spines forming an integral part of the anal fin.
Ventral with a continuous membranous attachment to the belly.
Snout conical, cleft of mouth narrow : villiform teeth in the jaws only.
Preopercular border entire : seven branchiostegals : pseudobranchiae present.
No air-bladder. Pyloric appendages numerous. Vertebra? 10+14.
25. Bathyseriola cyanea, Alcock.
Bathyseriola cyanea, Alcock, Ann. Mag. Nat. Hist. Sept. 1890, p. 202 : Illustrations of the Zoology of thi
Investigator, Fishes, pl. XVIII. fig. I.
B. 7. D. VIII-IX. 24-25. A. III. 22. P. 22. V. I. 5.
Body oblong and compressed ; its height about 3- in the total and one-
ninth less than the length of the head.
Head compressed and thin in its lower, broad and heavy in its upper half ;
its muciferous cavities well developed. Snout rounded, a little inflated at the
tip, the jaws equal in front ; its length, which is hardly equal to its greatest
breadth, is equal to the diameter of the eye. Eyes circular, their diameter not
quite one-fourth of the length of the head ; they are encircled by a sharp-edged
adipose fold, widest fore and aft ; interorbital space wider than the eye, convex
from side to side. Nostrils large, situated almost superiorly at the tip of the
snout.
Cleft of mouth narrow, the maxillary hardly reaching the vertical through
the middle of the eye ; jaw-bones weak, with a trenchant edge, which bears a
narrow band of villiform teeth; tongue large and fleshy; buccal folds very
broad. Gill-cleft wide ; gill-membranes united only quite anteriorly ; gill-covers
with thin, almost membranous, bones, the operculum with two diverging weak
stays above, the preoperculum bulging backwards as a large, striated, entire
lobe; gill-laminae broad, gill-rakers on the first arch long, close-set, acute;
pseudobranchiae fleshy. The mucosa of the whole pharynx black.
Scales extremely deciduous ; the few that still adhere are small and mem-
branous, and those of the lateral line, which are — inch in their major diameter,
have each a salient membranous tube.
The dorsal and anal fins have thick gelatinous bases ; the dorsal spines are
short and rather weak, and their interconnecting membrane is delicate ; the anal
44
spines are in close contact with each other and with the rest of the fin. Caudal
symmetrically forked. Pectorals pointed, their length rather more than four-
fifths the height of the body. Ventrals much shorter than the pectorals ; they
are adherent to the abdomen throughout their inner border, and can be retracted
within a shallow furrow in the middle abdominal line.
Peritoneal cavity large, the membrane black ; numerous pyloric caeca in an
arborescent mass ; no air-bladder. Vertebra? 10/14.
Colours in life, uniform bluish black, with an uneven silvery sheen.
In the Indian Museum are four adult specimens, the largest being nearly
7 inches long, from off the Ganjam coast 98-102 fathoms and off the Godavari
coast, 240-276 fathoms.
Regd.Nos. 12816, 12817, 12819: 13035.
This species has affinities on the one hand with Seriola and on the other
hand with the Nomeidse.
Family Cyttidce.
Antigonia, Lowe.
Antigonia, Lowe, Proo. Zool. Soc. 1843, p. 45.
Gaprophonos, Miiller and Troschel Horse Ichthyol, iii. p. xxviii.
Bypsinotos, Temminck and Schlegel, Faun. Japon. Poiss. p. 84.
Antigonia, Gunther, Cat. Fishes, ii. p. 497 and Challenger Deep-Sea Fishes, p. 44, (ubi synon) ■ Steindachner
and Doderlein, Denk. Ak. Wien, XLIX. 1885, p. 187 : Goode and Bean, Oceanic Ichthyology, p. 229.
Body compressed and elevated (rhomboidal) covered with rather small spiny
scales. One dorsal fin with eight spines : the soft portion, like the soft portion
of the anal, long. Anal with three spines, which though continuous with, are
somewhat isolated from, the soft portion. Ventrals composed of a spine and five
rays.
Mouth small, little protractile ; small teeth in the jaws only. Lower limb
of preoperculum serrated. Six branchiostegals. Pseudobranchire present.
Pyloric appendages few. Air-bladder present, large.
26. Antigonia capros, Lowe.
Antigonia capros, Lowe, Proc. Zool. Soc. 1843, p. 85: Gunther, Cat. Fishes, II. 497 and Challenger Deep-Sea
Fishes, p. 44 : Steindachner and Doderlein, Denk. Ak. Wien. XLIX. 1885, p. 187, pi. v : Goode and Bean, Oceanic
Ichthyology, p. 229, fig. 235.
Caprophonus aurora. Mull. & Trosch. Hor. Ichth. III. p. xxviii., pi. v. fig. 1.
Hypsinotus rubescens, Temm. & Schleg. Faun. Japon. Poiss. p. 84, pi. xlii. fig. 2. Gunther' Challenger Shore
Fishes, p. 44.
B. 6. D. VIII-IX. 27-34. A. III. 27-33.
The height of the body varies with age : in adults it is greater than the
length without the caudal, in specimens about a third grown it equals the length
of the body to the beginning of the caudal peduncle.
45
Head everywhere covered with harsh ctenoid scales; its length is from
two-fifths to a third the total without the caudal. The superficial bones of the
cranium are sculptured and striated : the edge of the preorbital is denticulated
and the horizontal limb of the opercle serrated.
Snout about equal to the width of the inter-orbital space in length and
rather shorter than the eye. Bye round, a third or more the length of the head.
Nostrils almost superior.
Mouth-cleft very small, the maxilla not nearly reaching to the anterior
border of the orbit. Gill-rakers short. Pseudobranchige very large.
Scales small, harsh, ctenoid; in from 60 to 65 rows between the gill-
opening and the caudal fin. Small scales extend some way along the spines
and rays of the dorsal and anal fins. Lateral line strongly curved.
2nd or 3rd dorsal spine the longest and by far the stoutest : its length is two-
thirds that of the head and twice that of the longest soft rays. 1st anal spine
the longest, about as long as the eye and about one-fourth longer than the
longest soft rays. 12 or 13 rays in the pectoral : the upper, which are the
longest, are more than three-quarters the length of the head. Ventral spine
very strong, its length is equal to that of the snout and eye combined : its edge
like the inner edge of the ventral rays is scabrous.
Colour in life red : in spirit light brown or yellow.
5 pyloric caeca, of which 2 are much enlarged.
Two specimens from off Trincomali 320 to 296 fathoms, and one from off
the Malabar coast 68 to 148 fathoms.
Eegd. Nos. 14134, 14135 : y8-
Distribution : West Indies, Madeira, East Indian Seas, Japan.
Family Trachifiidce.
Ueanoscopus, Cuv.
27. Uranoscopus crassiceps, Alcock.
TJranoscopus trassiceps, Alcock, Ann. Mag. Nat. Hist. Sept. 1890, p. 205 : Illustrations of the Zoology of
t-he Investigator, Fishes, pl. X. fig. 4.
B. 6. D. IV. ^ A. 13. P. 18. V. 1.5.
Length of the head 2- to two-fifths of the total without the caudal ; its
maximum breadth in repose (that is, when the opercles are not expanded for
defence) is two-thirds its length ; its greatest height (and that of the body) is
about one-fourth the total length of the body without the caudal.
Bones of the head rugose : the antero-inferior angle of the preorbital pro-
duced and subacute : a spine on the lower border of the suboperculum and 4
46
or 5 along the lower border of the preoperculum. Two small coarse spines or
tubercles on the supra-clavicular region : the clavicular spine about as long as the
major diameter of the orbit. The points of the pubic bones project as a pair
of spines between the clavicular symphysis.
Eye from a sixth to a seventh the length of the head, according to age, and
rather less than the width of the inter-orbital space. A small filament in front
of the nostril.
Lips rather fleshy, papillated, especially the lower lip. A large prelingual
filament, more than two-fifths the length of the head. No barbel.
No scales on the throat and belly.
Nine pyloric appendages.
Colours in life : back dirty greenish, below marbled with lighter shades,
belly silvery white, first dorsal black.
The type specimen is eight inchea long.
Numerous specimens from off the Ganjam coast 98 to 102 fathoms, off the
Coromandel coast 128 fathoms, and off the Malabar coast 45, 100 and 68 to 148
fathoms.
Regd. Nos. 12784-12786, 12788, 12791, 12797-12798, 12800, 12803,
13214, 13451, 13495, 13496, 13498-13500, 14093-14103: ^-f*:6-?? -If.
In the original description, I stated that there is no prelingual filament ;
but, as a matter of fact, the prelingual filament is very large.
The nearest relative of this species seems to be U. Jcaianus, Giinther
(Challenger Shore Fishes, p. 43, pi. xix. fig. A).
It is a voracious species : in the stomach of the specimen dissected seven
entire individuals of Scopelus pterotus were found.
Champsodon, Giinther.
Champsodon, Giinther, Proc. Zool. Soc. 1867, p. 102 ; Challenger Shore Fishes, p. 52 : Goode and Bean
Oceanic Ichthyology, p. 291.
Body elongate, compressed, covered with minute granular or spiny scales.
Two indistinct lateral lines, with transverse branches that lodge minute gland-
like papillae.
Mouth wide, oblique : irregular cardiform and setiform teeth of unequal size
in the jaws — largest in the lower jaw : cardiform teeth in two patches on the
vomer. Eye lateral, directed somewhat upwards. Two naked spines on edge of
preorbital bone.
Seven branchiostegals. Gill-openings very wide. Posterior border of pre-
operculum finely denticulated : a strong dagger-shaped spine at its angle.
47
Two dorsal fins, the first short, the second long and similar to the anal.
C audal forked. Pectorals small, median. Ventrals jugular, of good size.
An elongate air-bladder. A few large pyloric casca.
28. ChampSOdOfl VOrax, Giinther.
Champsodon vorax, Giinther, P. Z. S. 1867, p. 102, and Challenger Shore Fishes, pp. 43, 52, 56, pi. xxiii. 6g. A,
and Challenger Deep-Sea Fishes, p. 49: Alcock, Journ. As. Soc. Bengal, Vol. LV1II. pt. 2, 18S9, p. 302 : Illustra-
tions of the Zoology of the Investigator, Fishes, pl. XXVIII. fig. 5.
B. 7. D. V. 21. A. 19.
Length of head 3-, height of body 5 to 5^ in the total without the caudal.
All parts of the head, except the edge of the operculum, the throat and
branchiostegal membranes, and the sides of the chin, are covered with very
small granular scales, among which are some rows of tiny gland-like papillae .
The crown of the head is flat, and is traversed, from the snout backwards, by a
pair of ridges, which are fairly parallel as far as the occiput and then diverge to
end each in a supra-clavicular spinule. There are two diverging spines on the
edge of the preorbital, two spinules on the lower edge of the interoperculum,
and a long dagger-like spine at the angle of the preoperculum.
The snout, measured to the tip of the prominent lower jaw, is nearly twice
the length of the eye, which, in the adult, is about one-fifth the length of the
head. Inter-orbital space flat ; its width equals the diameter of the eye. Pupil
small. Nostrils almost superior, near the tip of the snout.
Mouth-cleft wide, very oblique, the lower jaw prominent, the maxilla
reaches well behind the posterior border of the orbit. The larger teeth in the
jaws are depressible.
Gill-opening extremely wide ; gill-membranes quite free. Operculum thin,
striated. Pseudobranchige small.
Body covered with minute ctenoid scales among which are some rows of
small gland-like papilla?. Two indistinct lateral fines, with numerous trans-
verse branches in which the gland-like papilla? are generally found.
First dorsal small, lower than the anterior part of the second dorsal, from
which it is separated by about an eye-length. The pectorals, which are not
much longer than the snout, arise high up on the side and consist of very
slender branched rays. Ventrals rather longer than the head without the
snout.
Six or seven large pyloric caeca.
Colours warm brown, gradually becoming whitish on the belly : the whole
surface with a frosted appearance : first dorsal black.
The largest specimen in the Indian Museum — an adult female — is 5 inches
long.
48
Numerous specimens from the Bay of Bengal 100 to 40 fathoms, and off
the Malabar coast 124 to 28 fathoms.
Regd. Nos. 11663, 11831, 11832, 11835, 11923, 11924, 12438,12506,
12948, 13935-13939, 14334-14350, —^ ~> 4-f •
Distribution : East Indian Archipelago, Bay of Bengal, Arabian Sea.
In the Indian Museum is also one of the Challenger Duplicates from the
East Indian Archipelago.
Bembrops, Steindachner.
Bembrops, Steindachner, SB. Ak. Wien, LXXIV. 1877, i. p. 211 ; Alcock, Jonrn. As. Soc. Bengal, Vol. LXIII.
1894, pt. 2, p. 118, and Vol. LXV. 1896, pt. 2, p. 316.
Hypsicometes, Goode, Proc. U.S. Nat. Mas. III. 1880, p. 347 ; Giinther, ' Challenger ' Deep-sea Fishes, p. 85;
Jordan and Gilbert, Bull. U.S. Nat. Mus. XVI. p. 808 ; Goode and Bean, Oceanic Ichthyology, p. 290 : Jordan &
Evermann, Fishes of N. America, III., p. 2293.
Bathypercis, Alcock, Journ. As. Soc. Bengal, Vol. LXII. 1893, pt. 2, p. 177.
Body elongate, subcylindrical : head large, depressed. Snout broad spathu-
late : cleft of the mouth wide, oblique, with the lower jaw projecting : villiform
teeth in the jaws vomer and palatines. Eyes large, close together, almost super-
ior.
Gill-opening very wide, the membranes free. Seven branchiostegals.
Opercle with 3 spines : angle of preoperculum feebly armed. Pseudobranchias
large.
Scales large, cycloid or very finely ctenoid, rather deciduous. Lateral fine
continuous from occiput to caudal, the scales of its anterior portion keeled or
feebly spinate. A small supra-clavicular spine.
Two separate dorsal fins, the first short, the second long. Anal similar to
the 2nd dorsal. Ventrals jugular.
Pyloric appendages few. No air-bladder.
29. Bembrops caudimacilla, Steindachner.
Bembrops caudimacula, Steindachner, Sitzungsb. Ak. Wien, LXXIV. 1877, i. p. 212 : Alcock, Jonrn. As. Soc.
Bengal, Vol. LXIII. pt. 2, 1894, p. 118.
Hypsicometes gobioides, Goode, Proc. U.S. Nat. Mus. III. 1880, p. 347, and Oceanic Ichthyology, p. 290, fig.
263 : Giinther, Challenger Deep-Sea Fishes, p. 85 ; Jordan and Evermann, Fishes of N. America III. p. 2294.
Bathypercis platyrhynchus, Alcock, J. A. S. B. Vol. LXII. pt. 2, 1893, p. 177, pi. ix. fig. 1.
Bembrops platyrhynchus, Alcock, J. A. S. B. Vol. LXIII. pt. 2, 1894, p. 118 : Illustrations of the Zoology
of the Investigator, Fishes, pl. XX. fig. 6.
B. 7. D. VI. 14. A. 16-17. P. circ. 25. V. I. 5. L. lat., from
origin on occiput, 50.
Head large, broad, depressed, its extreme length, measured from the tip of
the projecting mandible to the apex of the prolonged opercular flap is from two-
fifths of to 2^ in the total, caudal excluded. Body elongate, cylindrical, low,
49
and tapering to the large caudal : its greatest height is about one-third the
length of the head.
The snout is broad, much depressed, and spathulate, resembling the bill of
Bathypterois ; its extreme length is a little more than the major diameter of the
orbit which is about one-fourth the extreme length of the head. Mouth-cleft
wide, slightly oblique, the maxilla reaching nearly to the vertical through the
middle of the eye, and ending in a fleshy horizontally-disposed barbel. Teeth
in villiform bands on the jaws, vomer, and palatines. Tongue large, spathulate.
The large eyes are placed close together on the summit of the head, separ-
ated from each other by a narrow groove ; but the visual axis is lateral. The
gill-cleft is very wide, the gill-membranes being free of the isthmus throughout :
the preopercular angle is spinate, and the operculum, which is prolonged in
membrane nearly to the level of the 4th dorsal spine, has two spines above, and
one belonging to the suboperculum below. Four gills with setiform gill-rakers
and broad lamina? : pseudobranchias large.
The body, and the head and the snout above, are covered with rather large
cycloid or finely ctenoid scales. The lateral line beginning on the occiput as a
close-set row of weak re-curved spines, or strongly carinated scales, curves in-
wards towards the first dorsal fin and then downwards along the lower half of
the tail, being salient but unarmed in this part of its course.
The first dorsal fin is short, and is separated from the second by four or
five rows of scales : the second, which is much more elevated than the first,
extends from the level of the vent to within an eye-length of the base of the
caudal. The anal fin is similar to the second dorsal. The pectorals are large
and long and reach to, or even beyond, the origin of the anal. The ventrals
arise an eye-length in front of the pectorals and reach half-way to the anal.
Three rather large pyloric casca.
Colours in spirit, yellowish brown with thirteen incomplete and indefinite
darker cross-bands : a golden-green ocellus on the crown of the head and one in
the apex of each opercle, but all these ocelli tend to fade away : spinous dorsal
white at the base, black in the upper half.
In the young there is a large oval black and white ocellus in the upper part
of the basal half of the caudal fin ; but this gets broken up in older individuals.
An adult female in the Indian Museum is 81 inches lona:.
2 &
In the Indian Museum are 14 specimens from off the Coromandel coast 107
and 128 fathoms, from the Gulf of Manar 143 fathoms, and from the Andaman
Sea 185 and 194 fathoms : [also a specimen labelled Htjpsicometes gobioides from
the Gulf of Mexico 280 fathoms] .
Eegd. Nos. 13437, 13493, 13494: 2-^9> ™> ^
50
Distribution : Caribbean Sea and Atlantic coasts of N. America to 40° N.,
between 68 and 324 fathoms : Bay of Bengal and Andaman Sea 107 to 194
fathoms : Sea of Japan.
A series dredged in the Andaman Sea shows that B. gobioides is the adult
of B. caudimacula. The latter name has the priority.
Chiasmodus, Johnson.
Chiasmodus, Johnson, Proc. Zool. Soc. 1863, p. 408 and Ann. Mag. Nat. Hist. (3) XIV. 1864, p. 76 : Giinther,
Cat. Fishes V. p. 435, and Challenger Deep-Sea Fishes, p. 99: Goode and Bean, Oceanic Ichthyology, p. 291
Jordan and Evermann, Fishes of N. America, III. p. 2291.
Ponerodon, Alcock, Ann. Mag. Nat. Hist. (6) VI. 1890, p. 203, and (6) VII. 1891, pp. 9, 10.
Pseudoscopelas, Liitken, Vid. Selsk. Skr. 1892, 6 Rask. nat. math. Afd. VII. 6. pp. 285, 297.
Body elongate, naked. Eyes lateral. Two separate dorsal fins, of which the
second is much the longer and is equal opposite and similar to the anal ; ven-
trals thoracic ; pectoral rays branched. Caudal forked, cleft of mouth extreme-
ly wide ; jaws distensible and armed with canine teeth, as are also the palatines.
Gill-openings very wide, the gill-membranes united only quite anteriorly ;
preoperculum with a (small) spine at its angle ; seven branchiostegals ; pseudo-
branchia?. Lateral line single, uninterrupted. Abdominal cavity enormously
distensible. An air-bladder. No pyloric cseca. No anal papilla. Vertebra?
14/24. Mucous system of the head well developed.
30. Chiasmodus niger, Johnson.
Chiasmodus niger, Johnson, P. Z. S. 1863, p. 408 and Ann. Mag. Nat. Hist. (3) XIV. 1864, p. 76: Giinther,
Cat. Fishes, V. p. 435 and Challenger Deep-Sea Fishes, p. 99 : Carte, P. Z. S. 1866, p. 35, pi. ii : Jordan and
Gilbert, Cat. Fish. N. America, p. 1)9 : A. Agassiz, Ball. Mns. Comp. Zool. XV. 1888, p. 29, fig. 208 : Goode and
Bean, Oceanic Ichthyology, p. 292, fig. 264.
Ponerodon vastator, Alcock, Ann. Mag. Nat. Hist. (6) VI. 1890, p. 203, pl.ix. fig. 5.
Cliiasmodus niger, Alcock, Illustrations of the Zoology ok the Investigator, Fishes, pl. XXVIII. fig. 3.
B.7. D.X.29. A.29. V.I.5.
Body somewhat elongate and compressed, its height being 4- in the total
without the caudal.
Head low, long, and compressed, its length being 3- in the same standard ;
its surface is studded with pores, those on the crown being elliptical and
arranged in numerous longitudinal rows. A very large jwre, almost as big as
the anterior nostril, in front of the upper angle of the orbit.
Snout depressed, tapering, and rounded, its length being twice the diameter
of the eye and about one-fourth the length of the head ; the lower jaw projects
slightly. Eyes lateral, small, circular, deep-set; interorbital space nearly twice
the diameter of the eye and nearly flat from side to side ; it is traversed by two
anteriorly-converging ridges which enclose a V-shaped space. Nostrils large,
superior, situated near the tip of the snout.
51
Cleft of mouth oblique, extremely wide, the maxilla which is a very slender
bone, reaches almost to the angle of the preoperculum. Depressible hinged
fangs in two rows — those of the inner row being much the larger — in both jaws :
a row of distant, fixed, recurved teeth in each palatine. The front tooth on each
side of both jaws is also fixed. Tongue free, thin, almost spathulate.
Gill-openings wide ; gill-covers thin and flexible, the preoperculum with a
very oblique edge, a small, stout, obliquely decurrent spine at its angle, and a
thick muscular covering ; gill- membranes attached only quite anteriorly ; four
gills, the last gill-cleft a small foramen, branchial arches extremely weak and
flexible ; no gill-rakers ; pseudobranckias well developed.
Skin entirely scaleless, thin, covered with a uniformly thick adherent layer
of mucus ; a single lateral line of pores, which follows the dorsal profile from
occiput to base of caudal.
Two dorsal fins, separated by an interval equal to two-thirds the length of
the snout : the first, which begins slightly in advance of the vertical through
the base of the pectoral, consists of ten slender but well-ossified spines, of which
the longest (third) is barely as long as the snout and eye combined ; the second
contains twenty-nine slender articulated rays, branched at the tip and decreasing
regularly in length from before backwards, the longest (second) being about
half the length of the head. Anal equal, opposite and similar to the second
dorsal. Caudal symmetrically forked. Pectorals slender, as long as the post-
orbital portion of the head, all the rays branched. Ventrals thoracic, equal in
length to the eye and snout combined.
The abdomen is a great elastic sac, which extends behind the normally
situated vent into the tail ; it contains a vast collapsed stomach, but no pyloric
caaca. There is an air-bladder similar to that of Ghampsodon.
There are 14 abdominal and 24 caudal vertebras.
Colours in life : blotchy violet-black to black.
In the Indian Museum is a single specimen, just over six inches long, from
the Bay of Bengal, off the Godavari coast, 920 to 690 fathoms.
Regd. No. 12836.
Distribution : West Indies, North and Mid Atlantic, Madeira, Bay of Bengal.
Dr. Giinther places Chiasmodus among the Gadidse, but I feel pretty sure
that its place in the system is close to Ghampsodon.
Family JPecliculati.
When the second volume of the Fishes, in the Fauna of British India, was
published in 1889, only two genera of Pediculates were known to occur in
Indian Seas, namely, Antennarius and Ealieutsea.
52
We now know of nine Indian genera, which are shown in the following
table.
Synopsis of the Indian genera of the family Pediculati.
I. Gills three and a half or two and two halves: no pseudobranchiae :
an air-bladder : no pyloric caeca. Body compressed and elevated :
the spinous dorsal consists of 3 isolated spines, of which the first
is a rostral tentacle ... ... ... ... Antennarius.
II. Gills three : small pseudobranchiae present : no air-blader : two
pyloric caeca. Head and anterior part of body depressed, disk-
like : the spinous dorsal consists of #from 3 to 6 spines, of which
the first 3 are long isolated tentacles ... ... ... Lophids.
III. Gills two and a half : no air-bladder : no pyloric caeca : —
1. Body elevated: mouth-cleft oblique or nearly vertical: the
spinous dorsal consists of 1 or 2 unprotected tentacles.
No pseudobranchia? : —
i. Ventral fins present ... ... ... Chaunax.
ii. Ventral fins absent : —
a. Skin covered with prickles ... ... Ceratias.
b. Skin smooth ... ... ... ... Onikodes.
2. Head and anterior part of body depressed, disk-like : mouth
horizontal : the spinous dorsal consists of a single tentacle
lodged in a bony cavity above the mouth. Pseudobran-
chiae rudimentary, but distinct ... ... ... Halieutma.
IV. Gills two : no pseudobranchiae : no air-bladder : no pyloric caeca.
Head, etc., depressed, disk-like : mouth-cleft horizontal, not wide :
the spinous dorsal consists of a single tentacle lodged in a bony
cavity above the mouth : —
1. Soft dorsal fin present : —
i. Teeth in the jaws only ... ... ... Dibranchcs.
ii. Teeth in the jaws, vomer, and palatines ... ... Malthopsis.
2. No soft dorsal fin ... ... ... ... Halichetus.
Lophius, Artedi.
Lophius, Artedi, Genera Piscium p. 62 : C a v. and Val. Hist. Nat. Poiss. XII. p. 339: Giinther, Cat. Fishes,
II. p. 178 : Goode and Bean, Ocpanic Ichthyology, p. 485 :
Lophius and Lophiomus, Jordan and Evermann, Fishes N. America III. pp. 2713, 2714.
Head exceedingly large, forming a broad disk with the eyes on its upper
surface. Mouth exceedingly wide, the lower jaw projecting: jaws and palate
with cardiform teeth. Skin naked : bones of the head with numerous spines.
The tbree anterior dorsal spines, which are situated on the head, are isolated
and are modified to form tentacles : the posterior dorsal spines vary in number
from one to three, and when numbering three form a continuous fin. Soft
dorsal and anal short. Gills three. Pseudobranchias present, small. Pyloric
appendages two.
53
Distribution. American, European, and African coasts of N. Atlantic : Medi-
terranean : Seas of India and Archipelago : Seas of China and Japan.
Key to the Indian species of the genus Lophius.
I. Six dorsal spines, the last 3 of which form a continuous fin : —
1. Pectorals broad : 3rd dorsal spine a simple filament ... L. indicus.
2. Pectorals narrow : 3rd dorsal spine fringed with tags of
skin ... ... ... ••• ... L, gracilimanut.
II. Five dorsal spines, the last 2 rudimentary and hidden beneath the
skin ... ... ... ... ••• L. mutilus.
III. Four dorsal spines, the 4th not hidden ... ... ... L. lugubris.
31. Lophius indicus, Alcock.
lophius indicus, Alcock, Journ. As. Soc. Bengal, Vol. LVIII. pt. 2, 1889, p. 302 : Illustrations of the Zoology
of the Investigator, Fishes, pl. XIX. fig. 3.
B. 6. D. I. I. I. III. 8-9. A. 6-7. P. circ. 23. V. I. 5.
Disk subcircular, half the total length, caudal included, fringed all round
with tassels of skin which also extend along the sides of the tail and caudal fin
and on to the dorsal surface of the pectoral fins.
Two spines on the preorbital, and two on the elevated upper border of the
orbit. Humeral spine coarse, multifid.
Byes very small, their diameter being about one-tenth the length of the
cephalic disk and about two-thirds the width of the interorbital space.
Tongue hyoid and neighbouring parts of branchial arches coloured much
like the body.
Depressible teeth of unequal size in 2 or 3 irregular series in both jaws,
the inner series the largest (especially in the lower jaw) and most regular. Two
teeth on either side of the vomer, and three or four along each palatine.
First and third dorsal spines the longest, — rather over two-fifths the length
of the disk : the first spine a bristle ending in a large tuft, the second fringed
throughout, the third a simple bristle. Of the next three connected spines the
first is much the longest, the 2nd and 3rd being short. Caudal about a fifth the
total length (itself included). Pectorals broad and fan-like, all but the first
one or two and the last five or six rays being of approximately equal length.
Colour in life, dorsal surface dark grey or brown with either a network
of fine black lines or numerous small black rings : ventral surface colourless.
In the Indian Museum are eight specimens, the largest being just over five
inches long.
Malabar coast, 28 fathoms, Bay of Bengal 25 to 60 fathoms, Gulf of
Martaban 67 fathoms, Andaman Sea 90 fathoms.
54
Regd. Nos. 12450, 12451, 12504, 13216, 2-^^, ™.
This species is very closely related to, and may possibly be only a dwarf
variety of, Lophius setigerus Wahl, the common Fishing-frog of Japan.
32. Lophius gracilimaniis, n. sp.
Illustrations of the Zoology of thb Investigator, Fishes, pl. XXIX. fig. 3.
B. 6. D. I. I. I. III. 8. A. 6. P. circ. 18. V. I. 5.
Disk elliptical, nearly half the total length, caudal included, fringed as in
the preceding species, but more scantily.
Two spines on the preorbital, supra-orbital margin elevated and dentated.
Humeral spine truncated and bifid at tip.
Eyes of moderate size, their major diameter about one-sixth the length of
the disk and nearly equal to the width of the interorbital space.
Hyoid and neighbouring parts of branchial arches not or only slightly
pigmented.
Teeth as in the preceding species, except that the premaxillary teeth beyond
the vicinity of the symphysis become a single series.
Second dorsal spine the longest, its length being at least three-quarters that
of the disk : the first dorsal spine is a bristle ending in a small tassel, the second
is a simple bristle, and the third which is little shorter than the second is fringed
throughout its length. Of the next three connected spines the first is the
longest, but the 2nd and 3rd are also of good length. Caudal a fourth the total
length (itself included). Pectorals narrow, pointed.
Colours, mottled dark sepia to blue-black.
Only one of the pyloric caeca is large.
Three specimens, the longest a little over four and a quarter inches long,
from off the Malabar coast 68 to 148, and 100 fathoms.
This species is distinguished from L. indicus by the larger eye, the narrow
pointed pectoral fin, the long caudal fin, and the different form and length of the
isolated dorsal spines.
^ j -f.T 488-490 672
Regd. Nos. — - — , — .
33. Lophius mutilus, Alcock.
Lophius mutilus, Alcock, Journ. As. Soc. Bengal, Vol. LXII. pt. 2, 1893, p. 179 : Illustrations of the
Zoology of the Investigator, Fishes, pl. X. fig. 2.
B. 0. D. I. I. I. II rudimentary. 8-9. A. 5. P. circ. 15. V. 1.5.
Cephalic disk subcircular, not quite half the total length caudal included,
scantily fringed — like the sides of the tail and the dorsal surface of the pectoral
fins — with slender tassels.
55
A single distinct spine on the preorbital : three teeth on the sharp over-
hanging upper border of the orbit. Humeral spine trifid.
Byes large, their diameter being a fifth the length of the disk and equal to
the width of the interorbital space.
Hyoid and neighbouring parts of branchial arches dusky, but not distinctly
pigmented.
Small depressible fangs of unequal size in three irregular series in the
mandible, in two series near the symphysis of the upper jaw, but in a single
series along the greater part of the upper jaw. A pair of rigid fangs on each
side of the vomer, and five or six along each palatine.
The third dorsal spine is much the longest, being as long as the cephalic
disk : all three are plain bristles. The second portion of the spinous dorsal
consists of two small spines only visible after dissection. Caudal about a fourth
the total length (itself included). Pectorals narrow.
Colours in spirit, mottled brown.
A single specimen, 5; inches long, from the Bay of Bengal, off the Madras
coast, 128 fathoms.
Regd. No. 13438.
This species is at once distinguished by the rudimentary second part of the
spinous dorsal.
34. Lophius lagubris, Alcock.
Lophius lugubris, Alcock, Jonrn. As. Soc. Bengal, Vol. LXIII. pt. 2, 1894, p. 118: Illustrations of thk
Zoology of the Investigator, Fishes, pl. XIV. fig. 1.
B. 6. D. I. I. I. I. 7-8. A. 5-6. P. arc. 13. V. 1.5.
Cephalic disk subcircular, about half the total length, caudal included,
scantily fringed with slender tassels as in Lophius mutilus.
A single distinct spine on the preorbital ; 3 teeth on the sharp overhanging
upper border of the orbit. Humeral spine trifid.
Byes moderate, their diameter about a seventh the length of the disk, and
rather less than the width of the interorbital space except at its fore end.
Hyoid and neighbouring parts of mouth slightly dusky, not distinctly
pigmented.
Teeth as in Lophius mutilus.
All the dorsal spines are simple filaments : the third is much the longest,
being as long as or even longer than the cephalic disk. The second part of the
dorsal fin consists of a single short slender filament, about twice as long as the
eye. Caudal rather more than a fourth the length of the total (itself included).
Pectorals narrow.
56
Colours in spirit, dark sepia mottled with black.
Three specimens, tlie largest 5^ inches long, from off Colombo, 143 and 142
to 400 fathoms.
Regd. Nos. 13467, ™, ™.
This species is distinguished from L. nmtilus by the smaller eye, and by the
second portion of the spinous dorsal fin, which consists of a single filament. In
one of the specimens this filament is not present, or any trace of it.
Ceratias, Kroyer.
Ceratias, Kroyer, Natnrhist. Tidsskr. 1844-45, p. 639 : Giinther, Cat. Fishes, III. p. 205 and Challenger Deep
Sea Fishes, p. 52.
Ceratias, Diceratias, Mancalias, Typhlopsaras, Cryptopsaras, Gill : Goode and Bean, Oceanic Ichthyology, pp.
488^19 L (subgenera).
Ceratias and Mancalias (p. 2729), Cryptopsaras (p. 2731), Jordan and Evermann, Fishes of N. America,
Vol. III.
Head enormous, body and tail short ; both often elevated and compressed.
Skin with minute scutes or prickles. Mouth very large, approaching the
vertical, the mandible projecting. Depressible teeth of unequal size in the
jaws and sometimes on the vomer. Eyes small.
Gills two and a half. No pseudobranchia?.
Spinous dorsal reduced to one or two spines, which are generally modified
into tentacles. Soft dorsal and anal short.
Yentrals absent.
Distribution : Arctic and North Atlantic : Seas of India and Archipelago
Japanese Seas.
Subgenus Diceratias.
Two dorsal spines situated on the anterior part of the head. Vomerine
teeth present. No pyloric appendages.
35. Ceratias (Diceratias) Mspinosus, Giinther.
Ceratias hispinosus, Giinther, Challenger Deep Sea Fishes, p. 53. Alcock, Illustrations of the Zoology
OF THE INVESTIGATOB, FlSHES, PL. XXXV. FIG. 2.
D. I.I. 6. A. 4. C. 9. P. 14.
The height of the body, when the stomach is empty, is about half the total,
caudal included.
Head much larger than the body and tail combined. Each frontal bone with
a strong outstanding spine, situated above and behind the eye.
57
On top of the snout are the two isolated dorsal spines, the anterior of which
bears a long stout tentacle (about a third as long as the body) ending in a fleshy
knob, while the second is almost a rudiment.
Eye small, subcutaneous, about a third as long as the snout : in front of it
is a tubular nostril.
Mouth-cleft enormous, the length of the maxilla being nearly one-third of
the total, caudal included. A series of large and small depressible teeth in each
jaw : a few large teeth, decreasing in size from without inwards, on each side
of the vomer.
Skin of head and body covered with minute prickles.
Dorsal and anal fins placed close to the caudal, which is very large and is
pointed.
Colour black.
A single specimen, 5^ inches long, from off the Malabar coast, 636 fathoms.
Regd. No. 14008.
Distribution : Banda Sea, Arabian Sea.
Onieodes, Liitken.
Oneirodes, Liitken, Oversigt Dansk. Vid. Selsk. Forhandl. 1871, p. 56 : Gill, Proc. U. S. Nat. Una. I. 1878,
p. 227: Giinther, Challenger Deep Sea Pishes, p. 56: Jordan and Gilbert, Bull. U. S. Nat. Mas. XVI. p. 848:
Goode and Bean, Oceanic Ichthyology, p. 492 : Jordan and Evermann, Fishes of N. America III. p. 2732.
Paroneirodes, Alcock, Ann. Mag. Nat. Hist. Sept. 1890, p. 206.
Head enormous, body and tail short, both compressed and elevated. Skin
naked. Mouth large, oblique, the lower jaw a little prominent. Depressible
teeth of unequal size in the jaws and on the vomer. Byes small.
Gills two and a half. No pseudobranchiaa.
Spinous dorsal reduced to two spines, which are modified into tentacles.
Soft dorsal and anal short.
Ventrals absent. No pyloric cgeca.
Distribution : off coasts of Greenland : Bay of Bengal in deep water.
36. Ofiirodes glomerosus, Alcock.
Paron irodes glomerosus, Alcock, Ann. Mag. Nat. Hist. Sept. 1890, p 206, pi. ix. fig 6 : Illustrations of the
Zoology of the Investigator, pl. XXVIII. fig. 4.
D. LI. 6. A. 4. C. 8.
When captured the form of the body was ovoid, though unstable ; hardened
in spirit it becomes compressed and oval. The length of the head is five
eighths, its greatest height nine-sixteenths of the total, without the caudal. The
eye is small, being deeply buried beneath a circular patch of transparent (unpig-
58
merited) skin; above each eye is a prominent, coarse, procumbent spine.
Mouth moderately large, its cleft obliquely ascending ; the length of the maxilla
is one-third that of the head ; a narrow band (?) of small teeth in each jaw and
on the vomer ; tongue large ; only the floor of the mouth pigmented.
Gills 2:2 ; gill-opening a small circular aperture just beneath the root of the
pectoral fin.
Skin thin and perfectly smooth and scaleless ; it is protected by a thick
coat of mucus.
Two clavate cephalic tentacles, the first being rather more than twice the
length of the second, situated close together in the after part of the interorbital
space, with luminous organs imbedded in their enlarged tips. Second dorsal
and anal placed far back on the tail, almost in contact with the caudal, which is
pointed and in length a little more than one-fourth of the total ; all the rays
of the vertical fins simple ; pectorals very short, pointed ; ventrals absent.
Colours : — Body and fins jet-black ; in spirit the tip of the cephalic ten-
tacles become white. Pharyngo-branchial and peritoneal membranes unpig-
mented.
One specimen, 1- inch long from the Bay of Bengal, 1260 fathoms.
Re^d. No. 12840.
"t3
Chaunax, Lowe.
Chaunax, Lowe, Trans. Zool. Soc. III. 1849, p. 339 : Giinther, Cat. Fishes, III. p. 200 and Challenger Deep
Sea Fishes, p. 58: Goode and Bean, Oceanic Ichthyology, p. 487 : Jordan and Evermann, Fishes N. America, III.
p. 2726.
Head enormous, cuboidal. Skin covered with minute prickles. Mouth-
cleft wide, approaching the vertical, the lower jaw heavy and prominent.
Bands of small teeth in the jaws and palate.
Spinous dorsal reduced to a short tentacle situated on the snout. Soft
dorsal of moderate length. Anal short. Ventrals present.
Gills two and a half : no pseudobranchias. No pyloric ca?ca.
37. Chaunax irictus, Lowe.
Chaunax pictus, Lowe, Trans. Zool. Soc. III. 1849, p. 339, pi. li : Giinther, Cat. Fishes, III. p. 200 and
Challenger Deep Sea Fishes, p. 58, pi. x. fig. A : Goode, Proc. U. S. Nat. Mus. III. 1881, p. 470: Jordan and Gilbert,
Bull. D. S. Nat. Mas. XVI. p. 846: Vaillant, Exp. Sci. Travaillenr et Talisman, Poiss. p. 343, pi. xxviii. figs.
1-11 : Alcock, Ann. Mag. Nat. Hist. Nov. 1889, p. 381 : Goode and Bean, Oceanic Ichthyology, p. 487, fig. 398.
Chaunax fimbriatm, Ililgendorf, Sitzungsb. Ges. Naturf. Freunde, 1879, p. 80 : Steindachner and Doderlein,
Denk. Ak. VVien XLIX. 1885, p. 194.
B.7. D.I. 11. A. 6-7. P. 11. V. 4.. C. 8.
Shape like that of Diodon or Tetrodon.
59
The great cuboidal head is at least half the total length without the caudal,
and its greatest height, behind the eyes, is from a third to two-fifths the same
measure.
Byes large, subcutaneous, lateral although placed high up near the dorsal
profile.
Snout short, squarish, underhung by the massive square-cut lower jaw.
Nostrils two tiny pores situated near the edge of the snout. On the top of the
snout, folding backwards on to a shallow smooth depression of the skin, is a
tentacle, about half as long as the eye, ending in a leaf-like tassel : this repre-
sents the first dorsal fin.
Mouth-cleft wide, the length of the maxilla being two-ninths the total
without the caudal.
Skin extremely loose, covered with minute granules or prickles which are
finest on the throat and belly, traversed by chain-like rows of mucous pores as
follows : —
(1) one extending, on each side, from the snout over the eye, and then
curving downwards to run along the ventral surface of the tail and on to the base
of the caudal :
(2) one extending from the tip of the chin, on each side, along the lower
border of the head, to near the gill-opening :
(3) one across the head (" like a headstall ") behind the level of the eyes
from (2) to (2) :
(4) one on each side from the snout, in continuation of (1), round and across
the cheek, to (3) :
(5) one across the chin from (2) to (2).
Pectorals fairly broad, ventrals small and narrow.
Colours in spirit, either uniform light yellow, or light yellow with large
faint-brown spots ; throat white.
In the Indian Museum are numerous specimens from the Bay of Bengal
193, 272 and 145 to 250 fathoms, and two from off Colombo 14*2 to 400 fathoms
and 480 fathoms.
Regd. Nos. 11687, 11690, 11693-11695, 11719, 11720, 13468, 13183-13492,
528
T •
Distribution : West Indies and neighbouring Atlantic coasts of America,
Madeira, C. Verde and neighbouring coasts of Africa : Arabian Sea, Bay of
Bengal : Fiji, Japan : at moderate depths, 130 to about 400 fathoms.
E.
fumosa.
E. stellata.
H.
nigra.
E.
coccinea.
60
Halieuma, Cuv. and Val.
Diagnosis, etc., recorded in tbe Fishes of India.
Key to the Indian species of the geyius Halieutaea.
I. Under surface of disk covered with a thick, perfectly smootb, glandular
skin
II. Under surface of disk finely granular ; interorbital space decidedly
concave : —
1. Pour rays in the dorsal fin
2. Five rays in the dorsal fin...
III. Under surface of disk with stellate spines ; interorbital space hardly
concave ; five rays in the dorsal fin
38. JIalieutcea stellata, Wahl.
Srnonomy recorded in the Fishes of India.
B. 6. D. 4. A. 4. P. 13. V. I. 5. C. 9.
Tail, including the caudal fin, about three-fourths the length of the disk.
Disk broader than long, very little elevated anteriorly. Dorsal surface
covered with stout sharp spines having a broad star-shaped base. The spines
on the edge of the disk and along the side of the tail are bifid or multifid, and
usually have numerous short filaments between them.
Skin of the under surface of the disk finely granular.
Eyes between g and - the length of the disk in diameter, and about a dia-
meter apart. Interorbital space decidedly concave.
Rostral tentacle three-lobed.
Caudal about - the total length (itself included), not quite as long as the
pectoral. Ventrals moderately broad, between - and ^ the length of the pecto-
rals.
Parietal peritoneum moderately pigmented.
Colours in life, pink : in spirit the dorsum is light brown with some black
streaks and patches.
Has been taken in the Bay of Bengal at 98 to 102 fathoms and off the
Malabar coast at 68 to 148 fathoms.
Distribution: Seas of India, Malay Archipelago, China, and Japan.
39. Halieutcea nigra, Alcock.
Halieutxa nigra, Alcock, Ann. Mag. Nat. Hist. July, 1891, p. 24: Illustrations of thk Zoology of thb
INVE3TIGATOB, FlSUES, PL. XIX. FIG. 2.
B. 6. D. 5. A. 4. C. 9. P. 13. V. I. 5.
61
Differs from H. stellata in the following particulars : —
(1) the cephalic disk is circular and is decidedly elevated anteriorly, and
there are no cutaneous filaments along its edge, except at the chin :
(2) the eyes are about i the length of the disk, and are rather more than a
diameter apart :
(3) the caudal is - the total length (itself included) :
(4) the ventrals, which are only about half the length of the pectorals, are
slender.
Colours in life, blue-black with jet black vermicular lines on the dorsal
surface : in spirit bluish with the black lines more distinct.
A single specimen, just over 2- inches long, from the Andaman Sea, 188 to
220 fathoms.
Regd. No. 13027.
It is quite possible that this is only a variety of H. stellata.
40. Halieutcea coccinea, Alcock.
Halieutsea coccinea, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 382 : Illustrations of the Zoology of thi
Investigator, Pishes, pl. XIX. fig. 1.
B. 6. D. 5. A. 4. C. 9. P. 13-14. V. I. 5.
Differs from H. stellata in the following particulars : —
(1) the cephalic disk is circular and is distinctly elevated anteriorly, and the
cutaneous filaments on its edge are few and inconspicuous :
(2) the spines of the dorsal surface have needle-like points, and the under
surface of the disk is well covered with stellate spines : [in the young the spines
of the under surface are scattered and very small] :
(3) the interorbital space is but slightly concave and its width is much more
than a diameter of the eye :
(4) the caudal is about a fifth the total length :
(5) the ventrals are slender :
(6) the parietal peritoneum and branchial mucosa are jet black and parti-
cularly thick.
Colours in life : " dorsum bright pink, with fine black vermicular lines ;
under surface crimsou " : in spirit white, with the black lines very distinct and
the black peritoneum and branchial mucosa showing through on both sides of
the disk.
Numerous specimens from the Andaman Sea, 265 and 185 fathoms : the
largest is just over 7 inches long.
Regd. Nos. 11741: 2££J.
62
This species may perhaps be only a variety of H. stellata. An accident to
the unique specimen known in 1889 led me to describe the rostral tentacle as
bilobed : in the specimens received since then the tentacle has the usual three-
lobed form.
41. Halieiitcea fumosa, Alcock.
Halieutma futtwsa, Alcock, Jonrn. As. Sue. Bengal, Vol. LXIII. pt. 2, 1894, p. 119 : Illustrations of the
Zoology of the Investigator, Fishes, pl. XIV. fig. 2.
B. 6. D. 4. A. 4. C. 9. P. 13. V. I. 5.
Differs from H. stellata in the following particulars : —
(1) the spines of the dorsal surface are mere spinules, though their bases
are pyramidal or star-shaped : the filaments along the edge of the disk and sides
of tail are excessively delicate :
(2) the skin of the under surface of the disk is thick, gelatinous and abso-
lutely smooth :
(3) the length of the eye is between ^ and - that of the disk, and the
interorbital space is slightly concave :
(4) caudal one-fourth the total length, and equal to the pectorals :
(5) ventrals slender, more than - the length of the pectorals.
Colours in spirit : upper surface smoky blue, becoming hyaline near the
edge of the disk ; under surface grey, finely and closely speckled with silver ;
dorsal fin blackish ; pectorals and caudal with a broad black cross-band and
commonly a milk-white tip.
Numerous specimens, from the Bay of Bengal, 145 to 250 fathoms, and off
the Malabar coast, G8 to 148 fathoms. The largest is about 4 inches long.
Regd. Nos. 13716-13720, 13722-13725, 13727, 12823, ^^.
It is quite possible that this species also is merely a variety of II. stellata.
Dibranchus, Peters.
Dibranchus, Peters, Monatsb. Akad. Berlin, 1875, p. 736 : Gill, Proo. U. S. Nat. Mus. I. 1878, p. 231 : Giinther,
Challenger Deep Sea Fishes, p. 59 : Goode and Bean, Oceanic Ichthyology, p. 500 Jord*nn and Evermanu, Fishes
of North America, III. p. 27-13.
Head and anterior part of body forming a large subtriangular or ovate
disk the edge of which is armed with horizontal spines. Skin beset with spines
having stelliform bases, or with tubercles and granules.
Cleft of mouth horizontal, of moderate width. Minute teeth in the jaws
only.
Forehead with a transverse bony bridge forming a cavern above the mouth,
in which a retractile tentacle, that represents the 1st dorsal fin, is lodged.
63
Soft dorsal and anal short.
Two gills (on the 2nd and 3rd branchial arches). No pseudobranchiae.
No air-bladder : no pyloric appendages.
Distribution : "West Indies and Atlantic coasts of the United States : Cape
Verde and neighbouring coasts of Africa : Arabian Sea, Bay of Bengal, Anda-
man Sea. At moderate depths.
42. Dibrancluis nasutus, Alcock.
Dibranchus nasutus, Alcock, Ann. Mag. Nat. Hist, July 1891, p. 24, pi. vii. fig. 1 : Illustrations of thk
Zoology of the Investigator, Fishes, pl. XX. fig. 2.
B. 6. D. 6. A. 4. C. 9. P. 12-13. V. 1. 5.
Disk, measured to the gill-opening, shorter than the tail (including caudal),
its cranial portion very slightly elevated.
The frontal bridge projects considerably beyond the mouth, forming a
snout.
The rostral tentacle ends in a pair of fleshy balls, with a pair of filaments
above and between them.
Eyes about one-sixth the length of the disk and not much more than one
diameter apart anteriorly.
Dorsal surface closely covered with rigid spines having a stelliform base :
ventral surface much more sparsely beset with similar but smaller spines or
acute tubercles.
Dorsal fin in the anterior half of the tail, but some distance behind the gill-
opening : anal fin entirely behind the dorsal.
Caudal fin 4^ in the total length, equal to the pectorals. Ventrals narrow,
nearly as long as the pectorals.
Colour in life, blue black to jet black.
The largest specimen is not quite 3| inches long.
Andaman Sea, 188 to 220 and 405 fathoms : off Travancore coast, 406
fathoms.
Regd. Nos. 13028, 14116-14118, if3.
43. Dibranchus mici*opus, Alcock.
Dibranchus micropus, Alcock, Ann. Mag. Nat. Hist., July 1891, p. 25, pl. vii. figs. 2,2a, 2b,: Illustrations
of the Zoology of the Investigator, Fishes, pl. XX. fig. 1.
B. 6. D. 5. A. 4. C. 9. P. 15. V. 5.
Disk as long as, or longer than, the tail, its cranial portion very decidedly
elevated.
64
Edge of the frontal bridge flush -with the chin, not projecting.
The rostral tentacle ends in a pair of fleshy lobes, surmounted by a third,
median, foliaceous fimbriated lobe.
Eyes between a sixth and a seventh the length of the disk, somewhat more
than two diameters apart.
Dorsal surface closely covered with spines which have a stelliform base and
a flexible, almost setaceous, shaft ; on the tail they are almost rigid : ventral
surface with similar but smaller bristle-like spines.
Dorsal fin close behind the gill-opening, anal fin not entirely behind the
dorsal.
Caudal fin 4- to 4- in the total length, equal to the pectorals. Ventrals
very small, not a third the length of the pectorals.
Colour in life, uniform blue black.
The largest specimen is not quite 3 inches long.
Bay of Bengal, off Vizagapatam coast, 240 fathoms ; Andaman Sea, 370 to
419 and 405 fathoms ; off Travancore coast, 406 fathoms.
Regd. Nos. 13029, 13030, 14120, y' y •
Malthopsis, Alcock.
Malthopsis, Aleock, Ann. Mag. Nat. Hist., July 1891, p. 26.
Head and anterior part of body forming a large depressed sub-triangular
disk. Bones of the snout produced to form a sharp projecting spine, overhanging
a cavity above the mouth, in which a retractile tentacle is lodged.
Skin more or less beset with large conical striated tubercles.
Mouth-cleft rather narrow, horizontal. Villiform teeth on the jaws vomer
and palatines.
Soft dorsal and anal short.
Two gills (on the 2nd and 3rd branchial arches). No pseudobranchia?.
No air-bladder : no pyloric appendages.
Distribution : Indian Seas, Mid Pacific. At moderate depths.
44. Malthopsis lutea, Alcock.
Malthnpsis lutea, Alcock, Ann. Mag. Nat. Hist., July 1891, p. 26, pi. viii. figg. 2, 2a : Illustrations of tub
Zoology of the Investigator, Fishes, pl. XIX. fig. 4.
B. 5. D. 5. A. 4. C. 9. P. 11. V. 1.5.
Disk not quite as long as the tail (caudal included), its cranial part moder-
ately elevated. Snout projecting horizontally or obliquely upwards as a stout
striated spine.
65
Beneath this nasal prolongation is a deep narrow vault, flanked on each side
by a pair of large, almost confluent nostrils, and containing a short, fleshy,
clavate tentacle.
Eyes large, lateral, nearly circular ; their diameter is about one-seventh of
the total length, caudal not included ; they are strongly convergent and ante-
riorly are barely half a diameter apart ; the anterior limit of the orbit is in the
same vertical line with the anterior limit of the mouth.
The mouth-cleft, which is horizontal, is about two-thirds of an eye-diameter
in width. Teeth villiform, in bands in the jaws and in broad patches on the
vomer and anterior ends of the palatines.
Gill-cleft a small foramen, in width about one-fifth of an eye-diameter,
situated superiorly in the axilla ; two gills; no pseudobranchias. Sub-operculum
prolonged and ending in a stout trifid or multifid spine.
Body more or less covered with hard granular adherent plates, each with
a large radially-striated conical tubercle in its centre. On the dorsal surface of
the cephalic disk they are of moderate size, in contact along the middle line, but
distant and slightly sunken laterally ; on the ventral surface of the cephalic disk
they are very few and distant (except on the belly, where they may be numerous)
and sunken ; on the tad they are large and in close contact throughout.
The dorsal fin is in the anterior half of the tail, the anal is completely
behind the dorsal : the ventrals are very long, nearly equal to the pectorals,
which are equal to the caudal, which is two-ninths of the total.
Colours in life : pinkish yellow ; some specimens with a few irregular dark
rings on the dorsum of the cephalic disk.
Five abdominal and thirteen caudal vertebras, the neural spines of the
former fused into a trenchant ridge as in Malthe and Halieutsea.
The largest specimen is 3- inches long.
Andaman Sea, 185, 188 to 220, and 405 fathoms.
Regd. Nos. 13014-13016, 13018-13020 : 2|6, ^, 'f : l^p.
Haiicmetus, Alcock.
Halicmetus, Alcock, Ann. Mag Nat. Hist., July 1891, p. 319.
Head and anterior part of body forming a large depressed sub-triangular
disk. Front with a transverse bony bridge roofing in a cavity that lies above
the mouth and lodges a fleshy retractile tentacle representing the spinous dorsal
fin.
Skin covered with granules and tubercles.
9
66
Mouth-cleft rather narrow, horizontal. Villiform teeth in jaws vomer and
palatines.
No dorsal fin whatever (except the rostral tentacle) : anal fin very short.
Two gills (on the 2nd and 3rd branchial arches). No pseudobranchias.
No air-bladder : no pyloric appendages.
45. Halicmetus ruber, Alcock.
Htilicmetns ruber, Alcock, Ann. Slag. Nat. Hist., Jul^ 1891, p. 27, pi. viii. figs. 1, la-b. Illustrations of the
Zoology of the Investigator, Fishes, pl. XIX. fig. 5. (" Halieuteea coccinea," Goode and Bean, Oceanic Ichthyo-
logy, fig. 410.)
B. 6. D. 0. A. 3-4. C. 9. P. 11. V. 1.5.
Disk not quite as long as the tail (caudal included), its cranial part little
elevated.
The truncated snout is occupied by a bony rugose orbital bridge, beneath
which is a cavity lodging a fleshy tentacle which ends in three lobes, the middle
(superior) lobe being crested by a pair of papillae or small bifid filament. The
eyes are about one-seventh the length of the disk and are about half a diameter
apart anteriorly.
The nostrils are minute papilla? situated on each side of the rostral tentacle,
almost within the subrostral cavity.
Mouth horizontal, with the lower jaw slightly projecting ; its cleft is a little
wider than the eye. Villiform teeth in bands in the jaws vomer and on the
anterior ends of the palatines.
Gill-cleft a small foramen, less than half an eye-diameter in width, situated
superiorly in the axilla. The sub-operculum ends in a stout multifid spine.
Surface of the body uniformly invested with minute close-set graniform
spines, which also cover the eyes up to the corneal margin. The edge of the
cephalic disk bears in addition large finely granular multifid spines in three
longitudinal series, and the tail is clad with large granular conical tubercles —
of which there are five longitudinal series on each side — in close contact.
There are also some smaller tubercles scattered on the dorsal surface of the
disk.
Fins in form and position as in Halieufsea, Malthopsis, &c, but the soft dorsal,
as well as the spinous, is entirely wanting, and the anal is almost rudimentary.
The pectorals, which are abont a third longer than the ventrals and a little
longer than the caudal, are nearly one- fifth the total length.
Colours in life, uniform light pink.
Five abdominal and thirteen caudal vertebra?.
The largest specimen, a gravid female, is nearly 3- inches long.
Andaman Sea, 188 to 220, and 405 fathoms : Arabian Sea, off Travancore
coast, 406 fathoms.
Regd. Nos. 13025, 13026: 14122-14125 : ™.
Family Cottidce.
Teigla, Artedi.
46. Trig la hemisticta, Temm. & Schleg.
Synonomy and diagnosis recorded in the Fishes of India, p. 791, and Fanna of British India, Fishes II. p.
241.
Bay of Bengal, off Ganjam coast, 98 to 102 fathoms.
Regd. Nos. 12748, 12751, 12752, 12757, 12761, 12762, 12766, 12767, 12773,
12774, 12776, 12777.
Lepidotrigla, Giinther.
Lepidotrigla, Giinther, Cat. Fishes, II. p. 196.
" Head parallelopiped, with the upper surface and the sides entirely bony :
" the enlarged infra-orbital covering the cheek. Body with scales of moderate
" size, regularly arranged. Two dorsals, the first much shorter than the second.
" Three pectoral filaments. Villiform teeth in both the jaws and on the vomer,
" none on the palatine bones. Air-bladder generally with lateral muscles, often
" divided into two lateral halves. Pyloric appendages in moderate number."
(Giinther).
47. Lepidotrigla spiloptera, Giinther.
Lepidotrigla spiloptera, Giinther, Challenger Shore Fishes, p. 42, pi. xviii. fig. C ; and Challenger Deep Sea
Fishes, p. 64.
B. 7. D. IX. 15. A. 15. L. lat. 60.
Scales feebly serrated, those of the lateral line unarmed, those that imme-
diately flank the dorsal fins with well developed spines.
Profile of snout concave : pre-orbital projecting as a broad spine, about half
as long as the eye. Interorbital space very concave, its width is nearly equal
to the vertical diameter of the eye. A deepish transverse groove behind the
orbits, not well marked in the young.
The 1st dorsal spine, which is the highest, is not very much more than half
the length of the head. Pectoral fin reaching to the 4th or 5th anal ray.
Colours in life : reddish ; pectoral dark blue on its inner surface, with
numerous white spots and a white margin.
68
One specimen, nearly four inches long, from the Gulf of Martaban 67
fathoms.
Regd. No. -j-.
This species was taken by the Challenger in the seas of the Bast Indian
Archipelago at 140 fathoms.
48. Lepidotrigla spiloptera var. longipinnis, Alcock
Lepidotrigla spiloptera, var. longipinnis, Alcock, Ann. Mag. Nat. Hist., Dec. 1890, p. 429.
Only differs from the type in the great length of the pectoral fins, which
reach to, or beyond, the 9th anal ray.
Largest specimen five inches long.
Off Ganjain coast 18 fathoms, Gulf of Martaban 67 fathoms, Andaman Sea
55 fathoms, off Malabar coast 68 to 148 fathoms and 100 fathoms.
-r. i tvt innntr 295 416-417 499-527 680-688
Regd. Nos. 12925, -p, — j — , — j — , — — .
Family Cfltaphracti.
Peristethium, Lacepede.
Diagnosis, etc., recorded in the Fauna of British India, Fishes, II. p. 241.
Key to the Indian species of the genus Peristethium.
I. Pre-opercubx ridge not prolonged to form a spine ... ... P. Rivers- Andersnni.
II. Pre-opercular ridge prolonged to form a spine : —
1. Preorbital processes long, narrow, spatliulate ... P. serridatnm.
2. Preorbital processes short, broad, triangular : —
i. Twenty rays in the soft dorsal ... ... P. investigatoris.
ii. Fifteen rays in. the soft dorsal ... ... P. Halyi.
49. Peristethium Rivers- Ander so ni, Alcock.
Peristethus Rivers- Andersoni, Alcock, Journ. As. Soc. Bengal, Vol. LXIII. pt. 2, 18SN, p. 121, pi. vi. figs. 2,
2a, 26.
B. 7. D. VI. 22. A. 21. L. lat. 32.
The length of the narrow sub-spathulate pre-orbital processes is nearly
equal to the distance between their base and the anterior border of the orbit :
each has, on the upper surface near its base, a recurved upstanding spine.
The pre-opercular ridge is remarkably salient but is sharply truncated, not
forming a spine. The opercular ridge forms a short blunt spine.
The lower jaw is thickly fringed with small tentacles. The long labial
tentacles when laid back hardly surpass the angle of the mouth.
69
The interorbital space, the breadth of which is equal to the major diameter
of the orbit, is deeply concave, and is traversed fore and aft by a deep median
groove. Each supra-orbital margin is surmounted posteriorly by a strong
recurved spine, and there is a similar spine on each side of the occiput.
The body-shields are in four rows on each side : each shield is strongly
carinated, the carina being produced behind into a strong spine ; and in the case
of the shields of the posterior third of the lateral line the carinse are slightly
produced and pointed in front also.
The length of the anterior ventral shields is more than twice their greatest
breadth.
Colours in spirit : body flesh-coloured; the pectorals with a broad jet-black
band in their posterior half and with a milk-white tip ; the spinous dorsal black
in its upper half, and the soft dorsal with a black edge.
Length 3^ inches.
Off Colombo, 142 to 400 fathoms. One specimen.
Regd. No. 13469.
Named after Captain A. R. S. Anderson, I.M.S., Surgeon-Naturalist to the
Marine Survey of India from 1893 to 1899.
50. Pevistethhim serrulatum, Alcock.
Peristethus serrulatum, Alcock, Ann. Mag. Nat. Hist., August 1898, p. 153: Illustrations op the Zoology of
the Investigator, Fishes, pl. XXV. figs. 2, 2a.
B. 7. D. VII. 22. A. 21. L. lat. 33.
The length of the narrow spathulate preorbital processes is equal to more
than two-thirds the distance between their base and the anterior border of the
orbit.
Pre-opercular ridge sharply serrulate, ending in a curved rather narrow
spine, which is nearly as long as the eye. All the bony ridges of the head are
finely serrulate or serrate ; in addition there are, on either side, a preorbital, a
post-orbital, an occipital, a post-temporal and an opercular spine, and on the
forehead there are at least five small spines.
Interorbital space concave, less than the major diameter of the eye.
The large labial tentacles, when laid back, reach to the after limit of the
orbit.
All the shields of the body carry a stout recurved spine — eight rows in all ;
those of the posterior third of the lateral line are not simple spines, but are
acutely produced both forwards and backwards. The anterior ventral plates
70
are nearly twice as long as broad and nearly twice as long as the posterior
ventral plates.
Colours in spirit : flesh-colour, rather dusky dorsally ; distal half of pec-
torals and edges of vertical fins blackish.
Length nearly six inches.
Andaman Sea, 185 fathoms.
-r, , >T 296 297
Kegd. JNos. — > — .
51. Peristethiiim investigatoris, Alcock.
Peristethus investigatoris, Alcock, Ann. Mag. Nat. Hist., August 1898, p. 152: Illustrations of tub Zoologt
of the Invrstigator, Fishes, PL. XXV. F[GS. 1. In.
B. 7. D. VII. 20. A. 21. L. kit. 35-36.
The length of the broad triangular preorbital processes is equal to consider-
ably less than half the distance between their base and the anterior border of the
orbit.
Preopercular riclge trenchant, ending in a sharp spine, which is about two-
thirds as long as the eye.
A spine at the posterior angle of the orbit, one on either side of the occiput,
one on either post-temporal region, one at the upper angle of the operculum ;
in young specimens only there are three small inconspicuous tubercles, disposed
in a triangle, on the forehead.
Interorbital space concave, less than the major diameter of the eye.
The large labial tentacles, when laid back, reach far behind the posterior
border of the orbit.
All the shields of the body carry a stout recurved spine — eight rows in all.
The anterior ventral plates are irregular in shape, their greatest length,
measured diagonally, is nearly twice their breadth, and is half again as much as
the greatest length of the posterior ventral plates.
Colours in life : adults red, young dusky violet ; pectorals, first dorsal, and
distal half of labial tentacles black, second dorsal with a black edge.
The largest specimen is a little over 6 inches long.
Andaman Sea, 188 to 220 and 405 fathoms : off Travancore coast, 224 to
284 fathoms.
r» i tvt -.«.-.<-.,-, i i/^oo 121 140 529 530
Kegd. Nos. 13037, 13038, T, — , — , —.
This species appears to be near P. platycephakm Goode and Bean, from
Barbados.
71
Family GoMidce.
Gobius, Artedi.
52. Gobius cometes, Alcock.
Gobius cometes, Alcock, Ann. Mag. Nat. Hist., Sept. 1890, p. 208, pi. viii. fig. 2 : Illustrations of the
Zoology of the Investigator, Pishes, pl. XX. fig. 3.
B. 5. D. VI. 10-11. A. 10-11. L. lat. 23-24. L. tr. 5-6. C. 18-20.
P. 23. V. 1.5.
Length of the head about 2^, height of the body about 4- in the total with-
out the caudal.
Snout broad, its length about two-thirds that of the eye. Eyes entering
the dorsal profile, separated by a very narrow shallow groove, their major dia-
meter about 3- in the length of the head.
Mouth cleft oblique, the lower jaw a little prominent, the maxilla reaching
the vertical through the middle of the eye.
In each jaw an inner band of villiform teeth, and an outer regular row of
slightly enlarged, acute, slightly curved teeth ; tongue large and fleshy.
Gill-covers large, the suboperculum much larger than the operculum ; gill-
lamina? broad ; gill-rakers small and weak.
Scales large (0-23 inch in the vertical, 0"18 inch in the antero-posterior
diameter) microscopically ctenoid ; they cover the crown of the head as far as
the eyes, leaving the cheeks and opercles scaleless ; there are five or six rows of
scales between the second dorsal and the anal fins.
All the fins are elongated ; the second and third dorsal spines are about
half as long as the head; the rays of the feathery second dorsal and anal
increase in length from before backwards as far as the antepenultimate ray,
which is almost as long as the head. The caudal is long and pointed, its longest
rays, which are on the dorsal aspect, are nearly one-third the total length. The
ventrals are united, but are not adherent to the abdomen ; their length is about
equal to the height of the body. Pectorals with a long fleshy base, their longest
(middle) rays are equal to the length of the head without the snout.
Intestine short; anal papilla long and slender. A large thin- walled air-
bladder is present. Vertebras 11/13.
Colours in life : — Transparent grey, with seven broad bright-yellow cross
bands not quite reaching the middle line of the abdomen, and the gills showing
through the gill-cover as a bright pink blotch. Second dorsal and caudal fins
beautifully pencilled black and white like a feather, anal with a broad dark
border, ventrals blue-black. In spirit the yellow cross-bands almost entirely
fade.
72
Length between 4 and 5 inches.
Very numerous specimens from off the Ganjam coast, 98 to 102 fathoms,
and 107 fathoms.
Regd. Nos. 12729 et seg.
Ambltopus, Cuv. & Val.
53. Amblyopus arctocephalus, Alcock.
Amblyopus arctocephalus, Alcock, Ann. Mag. Nat. Hist., Dec. 1890, p. 432 : Illustrations of the Zoology
of the Investigator, Fishes, pl. XX. fig. 7 (eye far too distinct).
D. VI. 43. A. 41. Scales 50-60 rows.
Head angular, its opercular region somewhat inflated, its vertex compressed
into a sharp carina, its length one-sixth of the total, caudal included.
Body compressed, its height, which is 7 to 7-; in the total, caudal included,
diminishes very slightly from nape to base of caudal. Eyes completely hidden
and aborted, though the optic nerve is distinct.
Snout broad, with the lower jaw prominent. Mouth-cleft oblique, rather
wide, the length of the maxilla being 2^ in that of the head ; the upper lip with
a short broad barbel on each side; the mandibular symphysis prominent. In
each jaw a row of small, close, even, acute teeth, and external to these in the
front of the premaxilla, on each side, two large canines, and in the mandible
five, of which two are lateral and one (the largest) median.
Head naked ; body covered with thin, smooth, hardly imbricate scales,
which increase in size from before backwards.
Dorsal and anal fins low, enveloped in skin, confluent with the pointed
caudal. Pectorals with the four or five upper rays as long as the maxilla, the
lower rays extremely short. Ventrals jugular, small, cohering ; their length
varies from nearly two-thirds to not quite one-third the body-height.
Stomach large, saccular ; no pyloric casca. A large, globular, thick-walled
air-bladder. Anal papilla large, bilobed. Eleven abdominal, seventeen caudal
vertebra?.
Colours in life mottled pink, fins hyaline.
Length 5 inches.
Off Orissa coast, 50 fathoms, off Vizagapatam, coast 20 to 25 fathoms, off
Indus Delta 137 to 131 fathoms.
Regd. Nos. 12926-12931, 1 3 1-57-13459, ™, 3-^^.
Tn the drawing the artist has mistaken the dissected orbit for the eye. As
a matter of fact the eye-ball is indistinguishably fused with the connective-tissue
of the orbit, though the optic nerve is of normal size.
73
Callionymus, Linn.
54. Callionymus carebares, Alcock.
Callionymus carebares, Alcock, Ann. Mag. Nat. Hist., Sept. 1890, p. 209 : Illustrations of the Zoologt of
the Investigator, Fishes, pl. XX. fig. 4.
B. 7. D. IV. 9. A. 9. C. 9 + r. P. 21. V. I. 5.
The upcurved branchiostegal rays are prolonged considerably beyond the
suboperculum, so that the extreme length of the head is about two-fifths of the
total without the caudal. The height of the body is about one-sixth of the same
measure, and is less than the height of the head.
Eyes large, their major diameter being rather over one-fourth of the extreme
head-length and one-fourth longer than the snout; they are separated by a
narrow shallow groove.
Floor of the mouth dusky.
Preopercular spine upcurved, very fine and acute ; its length is two-thirds
the long diameter of the eye ; its base is advanced forwards as a sharp spine
of considerable length ; and on its upper border, close behind the angle of the
preoperculum, are one or sometimes two spinelets.
The gill-opening is not much smaller than the orbit and is rather more on
the side than on the top of the head ; the branchial arches are slender and flexi-
ble, the gill-rakers almost rudimentary.
The skin is loose and very thin. Lateral line single. The first dorsal fin
is lower than the second, its spines decreasing in length from before backwards ;
the height of the second dorsal and of the anal is not quite twice the greatest
body-height ; the length of the caudal is rather more than one-fourth of the
total in the female and about one-fourth the total in the male ; the pectorals are
rather shorter than the ventrals, which are as long as the postorbital portion
of the head and reach to or just beyond the origin of the anal, when laid back.
The intestine is convoluted ; the anal papilla is very slender, and in the
male it is very much longer than it is in the female. Vertebra? 8/13.
Colours in life : — the upper half of the head and body and all the fins range
from sepia-grey to blotchy black, and the ventral surface of the body is trans-
parent and colourless ; the first dorsal fin has in the male a central black patch,
and in the female a central, black, white-edged ocellus.
Total length 5 inches.
Numerous specimens, from off the Ganjam coast, 98 to 102 fathoms, and
from off the Malabar coast 100 fathoms.
Regd. Nos. 12740 et seq., 2±2»
In this species the secondary sexual characters are developed in the female,
not the male.
10
74
55. Ccillionymus kaianus, Gthr.
Callionymus kaianus, Giinther, Challenger Shore Fishes, p. 44, pi. xix. fig. B.
D. IV. 9. A. 9. C. 10. P. 21. V. 1.5.
Length of the head nearly a third, height of the body about a ninth of the
total length without the caudal. Eyes as long as or a little longer than the
snout, one-fourth the length of the head ; separated by a very narrow bridge.
Preopercular spine shorter than the eye ; its base is advanced forwards as
a sharp spine ; on its upper edge are two spinelets, the anterior of which is
very small.
Gill-opening a small aperture, not half the diameter of the eye, on the
upper side of the neck.
Lateral line single.
The anterior dorsal spine is prolonged, especially in the male, in which sex
it is not much shorter than the head. The rays of the second dorsal fin, in
both sexes, are as long as the postorbital portion of the head. The middle
caudal rays are prolonged in both sexes, being between a third and a fourth the
total length (caudal included).
The ventrals are a little longer than the pectorals ; in the female, but not
in the male, they reach beyond the origin of the anal.
Anal papilla large in the male.
Colours : reddish, with irregular large rounded violet spots along the
middle of the body : a lunate black spot, in both sexes, between the 3rd and 4th
dorsal spines; second dorsal with a series of large subocellated bands, which
are very conspicuous in the male.
Two specimens, adult male and female, from off the Malabar coast 102
fathoms. The male is 7 inches long.
Regd. Nos. — ' -r.
Distribution : Sea of New Guinea : Arabian Sea.
A large number of young, which may perhaps belong to this species, were
taken off the Malabar coast in 56 to 58 fathoms.
Suborder Anacanthini.
Anacanthini Gadoidei.
The Gadoidei of the Fauna of British India, include two families (Gadidse
and Ophidiidse), three genera exclusive of Ammodytes, and five species only.
To these we have now to add two families (Macrv/ridse and Ateleopodidse),
seventeen genera, and forty-four species, all of which have been brought to light
by the dredge of the " Investigator."
75
The following synopsis shows the inter-relations of the Indian families of
the Gadoidei : —
I. At least two dorsal fins : scales present : the ventrals in all Indian
species contain six rays or more : air-bladder and pyloric append-
ages generally present: —
1. Second dorsal fin well developed : a normal caudal fin in all
the known Indian species
2. Second dorsal more or less rudimentary : the tail tapers to a
filament
II. One dorsal fin : the ventrals consist of one or two filaments, or may be
wanting : —
1. Dorsal fin short, corresponding with the first dorsal fin of
Macruridee : each ventral consists of one filament. No
scales: no air-bladder : no pyloric creca
2. Dorsal fin long, occupying the greater part of the back :
ventrals, when present, consisting of one or two filaments.
Scales generally present, and air-bladder also. Pyloric
appendages present or not ... ... ... OphidiidjE.
GADIDiE.
Macrdrimi.
Ateleopodids.
Family Gaclidce.
Two genera are now known to inhabit the seas of India.
I. First dorsal fin above the pectorals, and consisting of several rays
an air-bladder
II. First dorsal consisting of a single ray placed on the occiput
Phtsiodlus.
Bregmaceros.
Bregmaceros, Thompson.
56. Bregmaceros 3Iacclellandii, Thompson.
Diagnosis and synonomy in Fauna of British India, Fishes, II. p. 433.
Numerous specimens from the Bay of Bengal, 10, 65, 95 and 145 to 250
fathoms ; from off the Andamans ; and from off the Malabar coast 56 to 58
fathoms.
Regd.Nos. 11830, 12387, 12475, 13442-13447, 13563-13587,
580
Physiculus, Kaup.
Physiculus, Kaup, Wiegmann's Archiv. f. Natnrges. 1858, p. 88 : Giinther, Challenger Deep Sea Pishes, p. 87 :
Goode and Bean, Oceanic Ichthyology, p. 365 : Jordan and Evermann, Fishes of N. America, III. p. 2547.
Physiculus and Pseudophycis, Giinther, Cat. Pishes, IV. pp. 348, 350 : vide Challenger Deep Sea Fishes,
p. 87.
Body elongate, covered with small scales. A separate caudal fin. Two
dorsals and one anal fin. Ventrals with a narrow flat base ; composed of
several rays. A band of villiform teeth in the jaws : no teeth on the vomer or
palatines. Chin with a barbel. Seven branchiostegals. Small glandular
pseudobranchiae in some of the species.
76
Distribution : West Indies, Madeira and neighbouring parts of Atlantic,
Mediterranean : India, Australia, Japan.
Key to the Indian species of the genus Physiculus.
I. First ray of the dorsal fin prolonged : the longest ventral ray
reaches only just beyond the origin of the anal ... ... P. roseus.
II. Dorsal fin not prolonged : the longest ventral ray reaches far beyond
the origin of the anal ... ... ... ... P. argyropastus.
57. Physiculus roseus, Alcock.
Physiculus roseus, Alcock, Ann. Mag. Nat. Hist., July, 1891. p. 28 : Illustrations of the Zoology op the
Investigator, Fishes, pl. XI. fig. 2 : Joarn. As. Soc. Bengal, Vol. LXIII. pt. 2, 1894, p. 122.
B. 7. D. 6-7/57. A. 55. V. 7.
Head and trunk broad ; tail compressed, higher than the trunk anteriorly.
Length of the head very nearly one-fourth of the total, including the caudal ;
greatest height of the body, just behind the origin of the dorsal fin, about one-
sixth of the total.
Snout depressed, broader than long, obtusely rounded ; its length, which is
equal to the major diameter of the eye and slightly exceeds the width of the flat
interorbital space, is one-fourth that of the head. Nostrils superior, situated
immediately in front of the orbit.
Mouth wide, oblique, with the upper jaw overlapping the lower ; the maxilla
reaches beyond the vertical through the middle of the orbit. Teeth villiform,
in broadish bands in the jaws only.
Barbel stout, about as long as the eye.
Gill-openings very wide. Small glandular pseudobranchia3.
Body and head covered with a thick mucilaginous skin, which is invested
everywhere with small deciduous scales, of which there appear to be six rows
between the first dorsal fin and the lateral line. The dorsal and anal fins, which
are invested with a fold of thick scaleless skin, extend to within an eye-length
of the caudal. The first dorsal, which is separated from the second only by a
notch, begins in the vertical through the base of the pectoral ; its first ray is
prolonged and nearly equals the postrostral portion of the head in length. The
ventrals arise on flattened bases ; their outer ray is prolonged only j ust beyond
the origin of the anal. The pointed pectorals arise on oblique bases ; their
length is about equal to that of the head behind the eye.
The vent is situated well in advance of the origin of the anal fin, but behind
the base of the pectorals, and there is a small postanal papilla. A large simple
air-bladder.
Colours in life uniform rose-red.
77
Length 7 inches.
From the Andaman Sea, 185 and 188 to 220 fathoms.
Regd. Nos. 13047 : 3-^-4.
58. Physiculus argyropastus, Alcock.
Physiculus argyropastus, Alcock, Joura. As. Soc. Bengal, Vol. LXII. pt. 2, 1893, p. 180, pi. ix. fig. 2, and
Vol. LXIII. pt. 2, 1894, p. 122 : Illustrations of the Zoology op the Investigator, Fishes, pl. XXII. fig. 1.
B. 7. D. 9/55. A. 57. V. 6.
Differs from P. roseus in the following particulars : —
(1) the length of the head is more than a fourth of the total, caudal in-
cluded :
(2) the length of the snout is barely equal to the width of the inter-orbital
space and exceeds the major diameter of the eye, which is about a fifth the
length of the head :
(3) the maxilla reaches nearly to the posterior border of the orbit :
(4) the first dorsal is not prolonged, its length being less than a third that
of the head :
(5) the vent is situated between the bases of the pectoral fins :
(6) the upper rays of the pectoral fin are as long as the head behind the
middle of the eye :
(7) the prolonged ventral ray reaches to the 6th or 7th anal ray, or even
beyond :
(8) the margin of the air-bladder is fimbriated.
Colours in spirit light pinkish brown with a silvery sheen : belly throat and
gill-membranes black.
Length 9 inches.
Bay of Bengal 107 and 128 fathoms : Gulf of Manar, 180 to 217 fathoms.
Regd. Nos. 13439-13441, 13541-13545, 13549, y.
Family OphidlidW-
Excluding Ammodytes, the genera included in the Fishes of India are two,
namely Brotula with 3 species, and Fierasfer with a single species.
The ' Investigator ' has since brought to light thirteen more genera and 23
more species, a few, indeed, of which belong to the fauna of the abysses, but
the majority of which are inhabitants of moderate depths.
78
Synopsis of the Indian genera of the family Ophidiicte.
* Vent situated at the throat
* Vent situated at least about a head-length behind the gill-opening : —
** Barbels present, on the chin
** No barbels : —
I, Caudal completely free
II. Caudal more or less united with the dorsal and anal fins : —
A. Head more or less scaly. Oviparous : —
1. Eyes well developed: —
i. Bones of the head firm, without spines (except per-
haps on the gill-covers) : greatest height of the body
from a fourth to a seventh the total length : pre-
operculum of moderate size : —
o. Pectorals entire : —
a. Lateral line distinct, extending well on to
the tail : —
k. Pyloric casca very small : pseudobran-
chia? consisting of 2 or 3 filaments ...
y. Pyloric cseca large : pseudobrancbire
absent
/3. Lateral line indistinct or absent
b. Lower pectoral rays prolonged and more or less
detached from each other and from the rest of
the fin
ii. Bones of the head thin and soft : greatest body-
height an eighth to an eleventh of the total length,
the tail ending in a long lash : preoperculum very
large and expanded : —
a. No spines on the head (except one on the oper-
culum")
b. Bones of the head with spiny crests
iii. Bones of the head thin and soft, with frill-like
crests : greatest body-height about a sixth the total
length : preoperculum of moderate size : —
a. No lateral line : ventral fins consisting of one
or two filaments
b. No ventral fins : lateral line peculiarly large
and conspicuous
2. No eyes : gill-covers armed with enormous spines
B. Head covered with a peculiar loose glandular scale-less
skin. Viviparous : —
1. Ventral fins consisting each of a single filament, which
however may be made up of more than one ray : —
i. Scales of the body imbricate : none of the teeth
enlarged
ii. Scales not or hardly imbricate : some of the teeth
enlarged
2. No ventral fins
[Fierasfer],
[Brotula].
[DlNENATICHTHYS].
Neobtthites.
Pycnocraspedum.
Bassogigas.
DlCROLENE.
Bassozetus.
Dermatorus.
Glyptophipiuji.
Lamprogrammlis.
TAOREPOPHimiM.
Dll'LACANTHOPOMA.
Saccogaster.
Hephthocara.
79
Neobythites, Goode & Bean.
Neobythites, Goode and Bean, Proc. U. S. Nat. Mus. VIII. 1886, p. 600: Giinther, Challenger Deep Sea Fiehe3
p. 100.
Pycnocraspedum, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 386.
Monomitopus, Alcock, Ann. Mag. Nat. Hist., Oct. 1890, p. 297.
Neobythites, (p. 325), Dicromita (p. 319), Benthocometes (p. 327), Bassogigas, (p. 328), Goode and Bean,
Oceanic Ichthyology : Jordan and Evermann, Pishes of N. America, III. pp. 2512, 2506, 2514, 2515.
Body elongate, compressed ; head not compressed, its bones firm : both head
and body covered with small cycloid scales : tail not filamentous.
Lateral line never continued to the end of the tail, sometimes very indis-
tinct.
Snout slightly overhanging the lower jaw ; without barbels. Mouth wide.
Villiform teeth in bands on the jaws and palatines, and in a A-shaped band or a
patch on the vomer.
Eye of moderate size.
Gill-openings wide : operculum with a spine, which is usually long sharp
and styliform, but may sometimes be weak and flat. Eight branchiostegals.
Pseudobranchias rudimentary (usually consisting of 2 or 3 filaments) or absent.
Dorsal and anal fins more or less confluent with the caudal. Each ventral
fin consists of two rays which may either be intimately fused to form a single
filament, or (more commouly) be separate in all or part of their extent : the
ventral fins are inserted, either close together or some little distance apart, just
behind the clavicular symphysis.
Air-bladder present. Pyloric caeca usually present.
Another character by which spirit specimens of Neobythites may be recog-
nized is that the dorsal profile of the cranium and snout form a single common
curve of no great convexity.
Distribution : Atlantic : Indo-Pacific. At moderate depths usually.
Key to the Indian species of the genus Neobythites.
I. The lateral line runs halfway along the tail, or further : pectoral fins
broad and short : pyloric casca present : —
1. A strong sharp styliform spine at the upper angle of the opercu-
lum : numerous long gill-rakers along the outer side of the 1st
branchial arch : each pseudobranch consists of two small fila-
ments : pyloric caeca very short : —
i. Very short pyloric cseca in a ring round the pylorus and in
two short rows along the mesenteric attachment of the
neighbouring part of the intestine : —
a. Two spines or spinules at the angle of the preoper-
culum: —
a. Each ventral fin consist of 2 rays coherent
only in their basal moiety . ... ... N. macrops.
80
j8. Each ventral fin consists of 2 rays coherent
throughout so as to form a single filament ... N. conjugator.
b. No spines or spinules at the angle of the preoper-
culum ... ... ... ... N. steatiticus.
ii. Short pyloric caeca in a ring round the pylorus only : angle
of preoperculum merely excised : each ventral fin consists
of a single filament ... ... ... ... N. nigripinnis.
2. A flat weak point at the upper angle of the operculum : only 4 or
5 long gill-rakers on the outerside of the 1st branchial arch : no
pseudobranchiaa whatever: pyloric cseea large [Pycnocraspedum] N. sqttamipinnis.
II. The lateral line is indistinguishable and appears to be present only close
to the head : pectoral fins long and feathery : no pyloric creca [Basso-
GIGAs] ... ... ... ••• ••• ••• N- fterotus.
The Indian species of Neobythites are so much alike that it will be sufficient
to give a diagnosis of one, and then to note merely the specific differences of the
others.
59. Neobythites maerops, Gthr.
Neohythites maerops, Giinther, Challenger Deep Sea Fishes, p. 102, pi. xx. fig. A : Alcock, AnD. Mag. Nat.
Hist. (6) IV. 1889, p. 385 and VIII. 1891, p. 30.
D. circ. 100. A. circ. 80. P. circ. 26. V. 2 coherent at base.
(I) Length of head 4l- to 4<2- in the total. (2) Greatest body height about
equal to the length of the head without the snout.
(3) Snout broad, rounded, hardly overhanging the upper jaw, as long as
the eye and about equal to the width of the flat interorbital space.
(4-) Major diameter of eye about two-ninths the length of the head.
(5) Nostrils rather far apart, — one in front of the eye, the other, subtu-
bular, near the edge of the snout.
(6) Upper jaw half as long as the head, overhanging the lower jaw. Teeth
in broad bands in the jaws, in a A-shaped patch on the vomer, and in an ellip-
tical patch on each palatine. None of the teeth in any way enlarged.
(7) Opercular spine long and sharp. (8) A spinule at the angle of the
preoperculum and another a short distance above it. (9) Gill-rakers on the outer
side of the first branchial arch numerous, of good length. (10) Each pseudo-
branch consists of 2 filaments.
(II) Body, head, and bases of fins covered with small scales, of which
there are 8 or 9 series between the 1st dorsal ray and the lateral line. (12) The
lateral line extends more than halfway along the tail.
(13) The dorsal and anal fins are confluent with the caudal : the longest
dorsal rays are equal to between a third and a fourth the greatest body
height.
81
(14) The distance between the first anal ray and the base of the pectoral
fin is equal to the length of the head without the snout.
(15) Pectorals pointed, their length is equal to the post-orbital portion of
the head.
(16) The ventrals are bifid but at some considerable distance from their
base, the inner branch being considerably the longer and both branches being
slender. The total leugth of the ventrals is equal to the length of the head
behind the middle of the eye.
(17) Pyloric casca extremely short, in a ring round the pylorus and in two
short series along the mesenteric attachment of the neighbouring part of the
gut.
Colours in spirit, yellowish grey mottled with brown ; some large black
blotches on the dorsal fin.
The largest specimen in the Indian Museum — an adult — is 8^ inches long.
Andaman Sea, 188 to 220, 271, and 405 fathoms : Arabian Sea, off Travan-
core coast, 224 to 284 fathoms.
Regd. Nos. 11646, 11647, 11649: 13053—13056, 13060, 13062—13064,
-ictrinf 151 156 585
130o6 : — ' — ' -j-.
Distribution : Fiji Is. : Andaman Sea.
60. Neobythites conjugator, Alcock.
Neobythites conjugator, Alcock, Journ. As. Soc. Bengal, Vol. LXV. 1896, pt. 2, p. 30-1: Illustrations of thk
Zoology of the Investigator, fishes, pl. XVII. fig. 4.
D. circ. 90. A. circ. 72. P. circ. 28. V. 2 (fused to form a single fila-
ment). Scales 100-110 rows.
This species differs from N. macrops only in the following particulars, which
for easy reference are numbered to correspond with the numbers relating to the
diagnosis of that species.
(1) Length of head about 4^ in total.
(6) The outer row of teeth in the upper jaw is distinctly enlarged.
(15) The pectorals are hardly half as long as the head.
(1 6) The ventrals are half the length of the head : each consists of two
rays intimately fused to form a single filament.
Colours in spirit, sepia ; caudal, distal two-thirds of pectorals, and outer
part of dorsal and anal fins black.
Length 9 inches.
Off Ceylon, 296-320 fathoms : off Travancore coast 406 fathoms.
Resfd. Nos. -; r.
° 11
11
82
61. Neobythites nigripinnis, Alcock.
Siremho nigripinnis, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 384
Monomitopus nigripinnis, Alcock, Ann. Mag. Nat. Hist., Oct. 1890, p. 295 : Illustrations of the Zoology of
the Investigator, fishes, pl. XI. fig. 3 (lateral line incorrect).
D. 95-100. A. 85-88. P. 28. V. I.
Differs from N. macrops in the following particulars : —
(3.) The snout, though as long as the eye, is only about - to - the width of
the interorbital space in length.
(6) The upper jaw is rather more than half the length of the head.
(8) The preopercular angle is excised, and the angles bounding the excision
are pronounced but are not distinct spines.
(12) The lateral line extends only about halfway along the tail.
(15) The pectorals are not half the length of the head.
(16) Each ventral is a single slender filament not half as long as the head.
(17) There is a ring of short pyloric casca round the pylorus only.
Colours in spirit, sepia ; fins black.
Length 9 inches.
Arabian Sea, 719, 740 and 824 fathoms ; Bay of Bengal, 561, 599, 753 and
696 fathoms ; Andaman Sea, 490 fathoms.
Regd. Nos. 11764, 12864, 13448, 13449, 13464, 13536-13540, p "» T'
329 331-333
T* "" 1 "
62. Neobythites steatiticus, Alcock.
Neobythites steatiticus, Alcook, Journ. As. Soc. Bengal, Vol. LXII. pt. 2, 1893, p. 181, pl. ix. fig. 3 : Illustrations
of the Zoology of the Investigator, Fishes, pl. XXI. fig. 2 (lateral line drawn too long).
D. circ. 85. A. circ. 65. P. circ. 22. V. 2 (coherent at base).
Differs from N. macrops only in the following particulars : —
(1) Length of head about 3'- in the total.
(4) Major diameter of the eye about one-sixth the length of the head.
(6) The bands of teeth in the jaws are not very broad.
(8) There are no spinules at the angle of the pre-operculum, which is
rounded.
(13) The longest dorsal rays are about two-fifths of the greatest body height.
(14) The distance between the first anal ray and the base of the pectoral
fin is equal to the length of the post-orbital portion of the head.
(15) Pectorals rounded, their length not much more than half that of the
head.
83
(16) The ventrals are equal in length to the post-orbital portion of the
head: each consists of two stout filaments — the inner of which is slightly the
longer — bound together in their basal half.
Colours in spirit : creamy yellow, clouded marbled and mottled like soap-
stone with shades of light brown ; a large oval ocellus, consisting of a black
centre and a creamy white ring, on the dorsal fin between the 20th and 30th
rays or beyond : anal jet black with a milk-white border.
Length 7 inches.
Bay of Bengal, 107, 128 and 145 to 250 fathoms.
Regd. Nos. 13435, 13474, 13476, 13478—13482.
63. Neobythites (Bassogigas) pterotus, Alcock.
Neohythites pterotus, Alcock, Ann. Mag. Nat. Hist., Sept. 1890, p. 210 ; Oct. 1890, p. 297 ; July 1891, p. 30 :
Illustrations of the Zoology of the Investigator, fishes, pl. XI. fig. 4 (female), ani> pl. XXIX. fig. 1 (male).
D. circ, 120. A. circ. 95. P. 18. V. 2.
Differs from N. 'inacrops in the following particulars : —
(1) Length of head about a sixth of the total : (2) greatest body height
equal to the length of the head.
(3) Snout nearly twice as long as the eye and about three-fourths the width
of the interorbital space ; somewhat overlapping the upper jaw.
(4) Major diameter of the eye about a seventh the length of the head.
(5) Anterior nostril large, not subtubular though pierced in a circumscribed
patch of naked skin.
(6) Upper jaw more than half the length of the head.
(8) No spinulea at the angle of the preoperculum.
(12) Lateral line either absent, or present only quite near the head : (thirty
rows of scales between the base of the dorsal and the vent).
(13) The longest dorsal rays are half the greatest body height.
(14) The distance between the base of the pectoral and the first ray of the
anal fin is 1- times the length of the head.
(15) Pectorals feathery : in the male they are 1- times the length of the
head, in the female they are as long as the head behind the anterior nostril.
(16) Bach ventral consists of two rays which are separate from their base ;
the inner ray, which is the longer, is about two-fifths the length of the head.
In the male both rays have spathulate tips.
(17) Pyloric caeca absent.
Colours : body brown ; head, abdomen, and all the fins black.
84
The largest specimen is nearly a foot long.
Bay of Bengal, 1310 and 1748 fathoms : Arabian Sea, 1000 fathoms.
Regd. Nos. 12832, 12863, 13046.
As in N. squamijriunis, the basal half or more of the dorsal and anal fins is
particularly fleshy and scaly.
64. Neobythites (Pycnocraspedwm) squamip hints, Alcock.
Pycnocraspedum squamipinne, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 38R.
Neobythites squamipinnis, Alcock, Journ. As. Soc. Bengal, Vol. LXI1I. pt. 2, 1894, p. 123 : Illustrations of
the Zoology of the Investigator, Fishes, pl. XXI. fig. 1.
Differs from N. macrops in the following particulars : —
(1) The length of the head is about 3- in the total.
(2) The greatest body height is about equal to the length of the head
behind the eye.
(3) The length of the snout though about equal to that of the eye, is only
between two-thirds and three-fourths that of the interorbital space.
(4) Major diameter of eye between a fifth and a sixth the length of the
head.
(5) Anterior nostril not subtubular.
(6) The teeth in the vomer and palate bones, though disposed in the same
way are in narrow bands.
(7) Opercular spine flat short and weak : (8) two or three rather indis-
tinct points at the angle of the preoperculum : (9) only four or five enlarged
gill-rakers on the outer side of the 1st branchial arch. (10) Pseudobranchia?
entirely absent.
(lfi) The ventral fins though otherwise similar are not half the length of
the head.
(17) Thirteen large long pyloric caeca.
Colours in spirit ; yellowish or greenish grey, fins darker.
The largest specimen is 11- inches long.
Bay of Bengal 193 and 145 to 250 fathoms.
Regd. Nos. 11700, 11701, 13525, 13526.
The vertical fins are more thickly scaled than usual, in this species, and the
caudal, though confluent with the dorsal and anal fins at its base, is free in a
considerable part of its extent.
85
Subgenus Dicrolene, Goode & Bean.
Dicrolene, Goode and Bean, Bull. Mas. Comp. Zool. X. 1883, p. 202, and Oceanic Ichthyology, p. 337 : Giinther
Challenger Deep-Sea Fishes, p. 107 : Jordan and Evermann, Pishes N. Amer. III. 2522.
Pteroidonus, Giinther, Challenger Deep-Sea Fishes, p. 106 : Goode and Bean, Oceanic Ichthyology, p. 337.
Paradicrolene, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 387.
Differs from Neobythites only in having the lower pectoral rays, to the num-
ber of 6 to 10, prolonged, and more or less isolated from each other and from
the rest of the pectoral fin, as free filaments. This one character is, in any case,
insufficient to justify the separation of Dicrolene from Neobytldtes ; but the
character itself is variable, for not only in the young of one species, but also in
the adults of another species, the lower pectoral rays are, for a considerable
distance, united to one another and to their pectoral fin by membrane.
Key to the species of Dicrolene.
I. Pectoral filaments free and independent in the adult : no circumscribed
cross-band on tlie tail : —
. 1. Twenty-seven rows of scales between the dorsal fin and the vent ... D. intronigra.
2. Tliirty-four rows of scales between the dorsal fin and the vent ... D. multifilis.
II. Pectoral filaments inter-connected by membrane in their basal moiety : a
broad black cross-band, involving also the dorsal and anal fins, on the
posterior third of the tail ... ... ... ... ... D.nigricaudis.
65. Dicrolene intronigra, Goode & Bean.
Dicrolene intronigra, Goode and Bean, Bull. Mus. Comp. Zool. X. 1883, p. 202, and Oceanic Ichthyology, p. 338,
fig. 297 A, B : Giinther, Challenger Deep Sea Fishes, p. 107 : Vaillant, Exp. Sci. Travaillear et Talisman, Poissons,
p. 258, pi. xxiii. fig. 2.
Paradicrolene Vaillanti, Alcock, Ann. Mag. Nat. Hist. (6) VI. 1890, p. 297.
D. circ. 100. A. circ. 85. P. 18-19/8-9. V. 2.
(1) Length of head one- fifth the total or less: (2) greatest height of the
body equal to the length of the head without the snout.
( 3) Snout broad, rounded, hardly overhanging the upper jaw, as long as the
eye, but hardly equal to the width of the inter-orbital space. Nostrils rather
large and far apart, one being in front of the eye, the other near the edge of the
snout.
(4) Major diameter of eye two-ninths to one-fourth the length of the head.
(5) Upper jaw decidedly more than half the length of the head, overhanging
the lower jaw. Villiform teeth in broadish bands on the jaws and palatines
and in a narrow A-shaped patch on the vomer : none of the teeth enlarged.
(6) Opercular spine a long sharp style : three spinules at the angle of the
preoperculum.
(7) Gill-rakers on the outer side of the first branchial arch numerous and
of good length. Each pseudobranch consists of two small filaments.
86
(8) Body, head, and bases of fins covered with small scales, of which there
are about twenty-seven rows between the dorsal fin and the vent. Lateral line
very indistinct, apparently not continued halfway along the tail.
(9) Dorsal and anal fins confluent with the caudal : the longest dorsal rays
are about two-fifths the greatest body height.
(10) The distance of the 1st anal ray from the base of the pectoral is
slightly more than the length of the head.
(11) Each pectoral consists of two portions, — a normal, pointed, upper
portion which is a little longer than the head, and a lower portion consisting of
eight or nine free filamentous rays the longest of which (2nd and 3rd) are
from 1^ times to twice the length of the head.
(12) Each ventral consists of two entirely separate rays, the inner and
Ion o-er of which is equal to the length of the head behind the middle of the eye.
(13) There are a few extremely short almost rudimentary pyloric casca.
Colours in spirit, yellowish brown : gill-membranes, pectoral fins and their
tree filaments, ventrals, and outer part of dorsal and anal fins, black.
The largest specimen is 10 inches long.
In the Indian Museum are 4 specimens from the Andaman Sea, 669 fathoms,
and the Arabian Sea, -406 and 740 fathoms, (besides a specimen from the Atlantic,
presented by the Smithsonian Institution).
Distribution : West Indies and Atlantic coasts of the United States : Atlantic
coasts of Morocco and north-west Africa: Arabian Sea. At considerable depths.
Eegd. Nos. 12862, ~, y, ™.
66. Dicrolene mwltifilis, Alcock.
Paradicrolene multifdis, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 387 and Nov. 1892, p. 3-48 : Illustra-
tions OF THE ZOOLOGT OF THE INVESTIGATOR, PL. XI. FIG 1.
D. arc. 100. A. arc. 85. P. 18/8-10. V. 2.
Differs from Dicrolene intronigra only in the following particulars, which
are numbered to correspond with the paragraphs relating to the diagnosis of
that species : —
(1) The length of the head is about two-ninths of the total.
(4) The major diameter of the eye is not quite two-ninths the length of
the head.
(8) Scales in 34 or 35 rows between the dorsal fin and the vent : the
lateral line ends in the last third of the tail.
(10) The distance of the 1st anal ray from the base of the pectoral is
i qual to the length of the head without the snout.
87
(11) The pectoral filaments are never as much as twice the length of the head.
(12) The inner ray of the ventral fin is not half the length of the head.
Colours in spirit yellowish or sepia, fins dark grey.
A mature female is only 6- inches long.
Bay of Bengal, 193 and 281 to 258 fathoms : Andaman Sea, 271 fathoms.
Regd. Nos. 11648, 11704, 11707, 11715-11717, 13166-13169.
67. Dicrolene nigrlcaudls, Alcock.
Paradicrolene nigricaudis, Alcock, Ann. Mag. Nat. Hist., July 1S91, p. 30 . Illustrations of the Zoology of
the Investigator, Fishes, pl. II, Fia 4.
D. circ. 90. A. circ. 75. P. 19-20/6-7. V. 2.
Differs from Dicrolene intronigra only in the following particulars : —
(I) The length of the head is two-ninths of the total.
(3) The length of the snout is quite equal to the width of the interorbital
space.
(4) The major diameter of the eye is not quite two-ninths the length of the
head.
(8) There are 30-31 rows of scales between the dorsal fin and the vent.
The lateral line is very distinct : it runs four scales below the first dorsal ray
and ends in the posterior fourth of the tail.
(10) The distance between the first anal ray and the base of the pectoral
fin is equal to the length of the head.
(II) The pectorals are as long as the postrostral portion of the head : their
free rays are connected with one another for a considerable distance, and with
the rest of the fin, by membrane, and are never more than 1^ times as long as
the head.
(12) The inner ray of the ventral fin is equal in length to the postorbital
portion of the head.
(13) The intestinal wall round the pylorus is puckered, but there are no
distinct cascal pouches.
Colours in spirit, brown : the posterior third of the tail, including the cor-
responding parts of the dorsal and anal fins, is black : the caudal fin and pectoral
filaments are white.
The largest specimen, which is not far off maturity, is a little over 8 inches
long.
Andaman Sea, 188 to 220 fathoms : Arabian Sea, off Travancore coast, 224
to 284 fathoms.
Regd. Nos. 13040, 13044, 5-^~.
88
Bassozetus, Gill.
Bassozetus, Gill, Proo. U. S. Nat. Mug. VI. 1883, p. 259: Goode and Bean, Oceanic Ichthyology, p. 321 :
Jordan and Evermann, Fishes of North America, III. p. 2507.
Bathyonus, Goode and Bean, Proo. U. S. Nat. Mus. Till. 1886, p. 603 -. Giinther, Challenger Deep Sea
Fishes, p. 321.
? Moebia, Goode and Bean, Oceanic Ichthyology, p. 331 : Jordan and Evermann, Fishes of North America, III.
p. 2510.
I follow Goode and Bean in substituting the name Bassozetus for Bathyonus, but the fact remains that
Bassozetus was quite insufficiently characterized by its author.
Head and body compressed, the body low and very elongate, the end of
the tail lash-like : the bones of the head soft, almost membranaceous. Both
head and body covered with small, deciduous, cycloid scales.
Lateral line indistinguishable.
Snout hardly overhanging the lower jaw, somewhat depressed ; without
barbels. Mouth wide. Villiform teeth in bands on the jaws and palatines and
in a A-shaped band on the vomer.
Bye small.
Gill-openings wide : operculum with a feeble spine above. Eight branchi-
ostegals. Pseudobranchiaa rudimentary or absent. Preoperculum large, usually
extending far back over the other opercular bones, its edge entire.
Dorsal and anal fins confluent with the caudal The ventrals arise close
together at the clavicular symphysis : each consists of either one or two fila-
ments.
Air-bladder present. No pyloric caeca.
This genus is very closely related to Neobythites, from which it only differs
in the following particulars : —
(1) the tail is long and lash-like and the body altogether lower and more
elongate :
(2) the bones of the head are very thin and soft :
(3) the angle of the preoperculum forms a sort of semicircular lobe extend-
ing some way over the other opercular bones.
Distribution : Atlantic and Indo-Pacific, at great depths. Nematmus and
Mixonus are probably not distinct from Bassozrias.
08. Bassozetus glutinosus (Alcock).
Bathyonus glutiiiosus, Alcock, Ann. Mag. Nat. Hist., Sept. 1890, p. 211 : Illostratio.ns of the Zoology of thk
Ikvestigatob, Fishes, pl. I. fig. 3.
D. cirr. 125. A. circ. 105. P. 29-30. V. 1.
Length of the head about one-fifth the total : greatest Leight of the body
three-fifths the length of the head.
89
Snout depressed, rounded, somewhat inflated at the tip ; its length, which
is less than its breadth, is nearly one-fifth the length of the head. Byes situated
in the uppermost part of the anterior third of the head, their major diameter
being one-tenth to one-eleventh of the head-length and one-third the width of
the convex interocular space. Nostrils large, one at the antero-superior limit
of the orbit, the other midway between the first and the tip of the snout. Mouth
wide, oblique ; the maxilla, which is half as long as the head, completely encloses
the mandible in repose ; villiform teeth in narrowish bands in the jaws, palatines,
and vomer, the last arranged in a V with incurved limbs.
Gill-covers large ; the preoperculum overlaps large portions of all the other
opercular bones, extending almost to the hinder edge of the operculum ; the
operculum with a feeble flat spur at the postero-superior angle, and another
below concealed by the overlying preoperculum ; seventeen long scabrous gill-
rakers on the first branchial arch, besides some rudimentary ones above; no
pseudobrancniae .
Scales small, thin, deciduous ; there are twenty-five rows between the dorsal
fin and the vent.
All the fin-rays delicate. The dorsal and anal fins are thick and fleshy ;
the highest rays of the dorsal — near the middle of the fin — are higher than the
corresponding anal rays, and measure nearly half the maximum body-height ;
the dorsal begins well in advance of the gill-opening. Caudal very narrow, its
length nearly one-twelfth of the total ; it is confluent with the other vertical fins
only at its base. Pectorals entire, pointed, half as long as the head. Ventrals
consisting each of a single filament, which is as long as the postorbital portion
of the head.
Colours in spirit : head belly and pectorals black, body brown, vertical fins
light grey.
The largest specimen is about 7- inches long.
Bay of Bengal, off the Ganjam coast, 1310 fathoms : Arabian Sea, off Malabar
coast, 636 and 891 fathoms.
Regd. Nos. 12824, 12825, 12827 : 14006, 14007, f .
Deematoetts, Alcock.
Dermatorw, Alcock, Ann. Mag. Nat. Hist., Oct. 1890, p. 298.
Celema, Goods and Bean, Oceanic Ichthyology, p. 329.
Head and body compressed, the body low and very elongate, the end of the
tail lash-like.
The bones of the head, though thin, are fairly firm, and are armed with
numerous upstanding spines.
12
90
Both head and body are covered with small deciduous cycloid scales — so
deciduous on the head that except in very well preserved specimens that part
may be thought to be naked.
Lateral line indistinguishable.
Snout hardly overhanging the lower jaw, depressed ; without barbels.
Mouth very wide. Villiform teeth in bands on the jaws and palatines ; few and
scattered, deciduous or absent, on the vomer.
Eye of moderate size.
Gill-openings wide : gill covers armed with spines. Eight branchiostegals.
Pseudobranchiaa rudimentary or absent. Preoperculum expanded much as in
Bassozetus. The gill-rakers on the outer side of the first branchial arch are long
and most curiously compound.
Dorsal and anal fins confluent with the caudal. The ventrals arise close
together at the clavicular symphysis : each consists of a single filament, which
may be bifid at its extremity.
Air-bladder small. Pyloric caeca absent or quite rudimentary.
This genus is very closely related to Bassozetus, from which it only differs
in the following particulars : —
(1) the bones of the head are firmer and are armed with numerous spines :
(2) the teeth on the vomer are few and scattered, or are entirely wanting.
Alcochia G-oode and Bean, Oceanic Ichthyology, p. 329, probably should be
united with Dermatorus.
Key to the species of Dermatorus.
I. Spines of the head rigid : diameter of eye about two-ninths the length
of the head : some vomerine teeth present ... ... ... D. trichiurus.
II. Spines of the head weak and flexible : diameter of eye a sixth or a
seventh the length of the head ; —
1. Some vomerine teeth ... ... ... ... B. melanocephalus.
2. No vomerine teeth ... ... ... ... B. melampeplus.
69. Dermatorus trichiurus, Alcock.
Dermatorus trichiurus, Alcock, Ann. Mag. Nat. Hiac, Oct. 1890, p. 298 : Illustrations of the Zoology op
THE INVESTIGATOB, FlBHES, PL. I- FIG. 1.
D. 160 + x. A. 140 + 0. P. circ. 16. V. 1. (split at the end).
Head between a sixth and a seventh the total length : greatest height of the
body equal to the length of the head behind the posterior border of the pupil.
Two small preorbital spines. A s'rong recurved spine at the anterior angle
of the orbit from which two irregular series of spines pass backwards to the
91
supraclavicular angle. In addition both edges of a broad mucous channel
excavated in the edge of the preoperculum are spinate, and there is an acute
spine at the upper angle of the operculum.
Snout depressed ; its length is a little more than that of the eye and equal
to the width of the interorbital space. A large nostril in front of the eye, and
a smaller one midway between it and the edge of the snout.
Diameter of the eye about two-ninths the length of the head.
The length of the upper jaw is nearly two-thirds that of the head. Teeth
in narrow bands in the jaws and palatines ; few and scattered on the vomer.
About 20 long gill-rakers on the outer side of the first branchial arch.
Each pseudobranch consists of two inconspicuous papillae.
Scales in about 20 rows between the dorsal fin and the vent. No lateral
line.
The rays of the vertical fins are weak : the first anal ray is a little less than
a head-length behind the axilla. Pectorals pointed, as long as the post-orbital
portion of the head. Each ventral consists of a filament, as long as the post-
rostral portion of the head, split into two near its end.
The intestine is puckered round the junction with the pylorus, but there are
no distinct cgeca.
Colours in spirit : head and belly black, body light brown, fins light grey.
An adult female is a little over 7 inches long.
Arabian Sea, 1000 and 931 fathoms.
Regd. Nos. 12865 : ^.
This species may probably prove to be identical with the previously de-
scribed Porogadus nudus Vaillant (Exp. Sci. Travailleur et Talisman, Poissons,
p. 262, pi. xxiv. fig. 2).
70. Dermatorus melanocephalus, Alcock.
Dermatorus melanocephalus Alcock, Ann. MagyNat. Hist., July 1891, p. 32: Illustrations of the Zoology
of the Investigator, Fishes, pl. XXI. fig. 4.
Differs from D. trichiurus only in the following particulars : —
(1) the bones of the head are thin and soft and the spines are all weak and
flexible, bending, instead of pricking the hand as they do in D. trichiurus ; there
are no spines on the edge of the preoperculum :
(2) the diameter of the eye is only one-sixth the length of the head and half
that of the snout :
(3^ the long gill-rakers on the outer side of the first branchial arch are less
numerous : and there are no pseudobranchias :
92
(4) the pyloric cgeca are somewhat less rudimentary.
An adult female is nearly 8 inches long.
Bay of Bengal, 1644 and 1748 fathoms.
Regd. No. 13073.
This species may very probably prove identical with Porogadus subarmatus
Vaillant (op. cit. p. 265, pi. xxiv. fig. 3).
71. Dermatorus melmnpeplns, Alcock.
Dermatorus melampeplus, Alcock, Journ. Aa. Soc. Bengal, Vol. LXV. pt. 2, 1896, p. 305 : Illustrations of
the Zoology of the Investigator, Fishes, pl. XVII. fig. 3.
Differs from D. trichiurus in the following particulars : —
(1) As in D. melanocephalus the spines of the head are weak and flexible ;
but there are some on the edge of the preoperculum :
(2) the eye is half the length of the snout and less than a sixth the length
of the head :
(3) there are no teeth on the vomer.
Colour uniform black.
Length a little over 9 inches.
One specimen from off the Malabar coast, 931 fathoms.
Regd. No. y-
Gltptophidium, Alcock.
Glyptophidium, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 390, and Jonrn. As. Soc. Bengal, Vol. LXV. pt. 2,
1896, p. 309.
Body elongate, compressed, of good height ; tail long, tapering to a lash-like
filament : head compressed, with soft, almost membranaceous, frilled and crested
bones. Both head and body covered with thin caducous scales.
Lateral line absent.
Snout not overhanging the lower jaw, without barbels. Mouth wide :
villiform teeth in bands on the jaws and palatines and in a A-shaped band on
the vomer.
Eye of good size.
Gill-openings wide: operculum with a feeble flat spine. Eight branchiosteg-
als. Each pseudobranch consists of from 6 to 12 rather long lax filaments.
Dorsal and anal fins confluent with the caudal.
The ventral fins arise close together just behind the clavicular symphysis :
each consists of either 1 or 2 filamentous rays.
93
Air-bladder present. Pyloric appendages small, almost rudimentary.
The tail is so extremely slender and filiform that the end of it is often lost,
and when the caudal fin grows again it appears to be free from the other vertical
fins.
The lash-like tail, the frilled crests of the head bones, and the compara-
tively large pseudobranchia? distinguish Glyptophidium. In no other Indian
Ophidioid do the pseudobranchise consist of more than 2 small filaments.
Key to the species of Glyptophidium.
I. Each ventral consists of a single filament ... ... ... 0. argenteum.
II. Each ventral consists of two filaments ... ... ... G. macropus.
72. Glyptophidium argenteum, Alcock.
Glyptophidium argenteum, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 390 : Illustrations of the Zoology
of the Investigator, Fishes, pl. II. fig. 3.
Length of the head about a fifth the total, greatest body height (at the
shoulder) equal to the length of the head without the snout. The body rapidly
tapers to a long filamentous tail.
Length of the snout equal to the major diameter of the eye (which is nearly
a fourth the length of the head), barely equal to the width of the interorbital
space.
The upper jaw overlaps the lower, except at the tip, where a sharp knob at
the mandibular symphysis projects slightly : its length is half that of the head.
Villiform teeth in a broadisb band on the upper jaw, in a narrow band on the
lower jaw, in a slightly curved narrowly-elliptical band on the palatines, and
in a A-shaped band on the vomer.
Three weak points at the angle of the preoperculum. Numerous long
gill-rakers on the outer side of the 1st branchial arch. Each pseudobranch
consists of about a dozen filaments.
Scales rather large, excessively thin and deciduous, — more especially on the
head, so that in specimens that are not exceptionally well preserved the head
appears naked.
Dorsal and anal fins confluent with the caudal, the rays of the anal being
much shorter than those of the dorsal and the rays of all three being very
slender : the longest dorsal rays are nearly half the body height.
Pectorals pointed, a little longer than the head without the snout, the rays
very slender and about 22 in number.
Each ventral consists of a single slender ray which is about as long as the
post-orbital portion of the head.
94
A few rudimentary caeca round the pylorus only.
Colours in spirit : blackish, silvered over ; fins blackish grey.
Numerous specimens, the largest 9- inches long.
Andaman Sea, 271 and 405 fathoms : off Travancore coast, 360 and 406
fathoms.
t» , at ,,»», H 17 125 144 160 161 164 334 336 337
Regd. JNos. 11661, y - y, — , — , -j-, ~> T' T' T' T'
73. Glyptophidium, macropus, Alcock.
Qlyptophidium macropus, Alcock, Journ. Aa. Soc. Bengal, Vol. LXIII. pt. 2, 1894, p. 122, pi. vi. fig. 3 :
Illustrations of the Zoology of the Investigator, Fishes, pl. XV. fig. 6.
Differs from 0. argenteum only in the following particulars : —
The length of the head is only about a fourth the total.
The angle of the preoperculum is notched, the angles of the notch being
rounded off.
Each pseudobranch consists of from 5 to 8 filaments.
The length of the pectorals is equal to that of the head behind the middle
of the eye.
Each ventral consists of two rays of which the inner one is an eye-length
longer than the head.
The pyloric caeca are longer.
The largest specimen is 5^ inches long.
Colours in spirit : head, iris and body silvery, the body finely speckled with
black : vertical fins greyish with blackish tips, pectorals blackish, ventrals milk
white.
Bay of Bengal, 145 to 250 fathoms.
Regd. Nos. 13529-13535.
Lamprogrammus, Alcock.
Lamprogrammus, Alcock, Ann. Mag. Nat. Hiat., July 1891, and Jonrn. As. Soc. Bengal, Vol. LX V. pt. 2, 1896,
p. 309.
Body elongate, compressed, of good height ; tail long, tapering to a lash-
like filament : head compressed, with soft, almost membranaceous, frilled and
crested bones : both head and body covered with thin deciduous scales.
Lateral line broad, very conspicuous, continued at least halfway along the
tail ; its scales much enlarged, each being furnished with a (luminous) gland.
Snout not overhanging the jaws, without barbels. Mouth wide. Villiform
teeth in bands on the jaws and palatines and in a A-shaped band on the vomer.
Eye of moderate size.
95
Gill-openings wide: operculum with a feeble flat spine. Eight branchi-
ostegals. No pseudobranchia?.
Dorsal and anal fins confluent with the caudal.
No ventral fins.
A small air-bladder present. Pyloric appendages few and small.
The curious Halosanrus-like lateral line and the absence of ventral fins and
pseudobranchiae distinguish this genus from Glyptophidium, which it otherwise
closely resembles.
Key to the species of the genus Lamprogrammus.
I. The angle of the preoperculum is simply notched ... L. niger.
II. Three weak flat teeth at the angle of the preoperculum ... ... L.fragilis.
74. Lamprogrammus niger, Alcock.
Lamprogrammus niger, Alcock, Ann. Mag. Nat. Hist., Jnly 1891, p. 33, fig. 2 : Illustrations of the Zoology
of the Investigator, Fishes, pl. I. fig. 2.
D. circ. 110. A. circ. 90. P. 17. V. 0.
Length of the head about one-fifth of the total, greatest body height (at
the shoulder) equal to the length of the head without the snout.
Length of the snout about twice that of the eye, less than the width of the
convex interorbital space.
The major diameter of the eye is an eighth or a ninth the length of the
head.
The upper jaw, the length of which is half that of the head, overlaps the
lower. Villiform teeth in a broad band in the upper jaw, in a narrow band on
the lower jaw and on each palatine, and in a narrow broken A-shaped band on
the vomer.
Angle of the preoperculum notched, the angles of the notch rounded off.
About ten long gill-rakers on the outer side of the first branchial arch.
Body and head covered with deciduous, almost membranaceous, scales of
moderate size.
The scales of the very conspicuous lateral line are adherent and greatly
enlarged; they lie beneath a continuous sheath of black skin, which is loopholed
over a long narrow groove with raised margins situated along the vertical
diameter of each scale. These grooves are filled with an opaque white substance,
which probably has a luminous function. The lateral line, in fact, is exactly
similar to that of several species of Halosaurus.
96
The rays of the fins are weak and are damaged beyond description by cap-
ture : those of the dorsal fin are much more strongly developed than those of the
anal. The pectorals appear to be short.
Six small pyloric casca.
Colour : jet black.
The largest specimen is 15^ inches long.
Bay of Bengal, near the An damans, 561 fathoms : Andaman Sea, 405
fathoms.
Regd. Nos. 13048, 13049.
75. Lamprogr animus fragilis, Alcock.
Lamprogrammus fragilis, Alcock, Ann. Mag. Nat. Hist., Nov. 1892, p. 348.
D. circ. 90. A. circ. 75. P. 17. V. 0.
Differs from L. niger only in the following particulars : —
The greatest height of the body is equal to the length of the head.
The major diameter of the eye is between a sixth and a seventh the length
of the head.
The upper jaw is not quite half as long as the head.
There are three weak teeth at the angle of the preoperculum.
The scales of the lateral line lie beneath a continuous tube of black skin
which is traversed along the middle by a continuous bright stripe.
The pyloric caeca are a little longer.
Colour : jet black.
The largest specimen is 2 feet long.
Bay of Bengal, off Godavari poast, 678 fathoms : Arabian Sea, off Travancore
coast, 406 fathoms.
Regd. Nos. 13171 : \ - j .
Taubedophidium, Alcock.
Tauredophidium, Alcock, Ann. Mag. Nat. Hist., Sept. 1890, p. 213.
Body elongate, compressed : head large and broad, not compressed ; its
bones firm : both head and body covered with small cycloid scales.
Eyes atrophied and completely hidden beneath the skin.
Operculum and preoperculum armed with enormous spines.
Lateral line indistinguishable.
97
Snout broad, not overhanging the jaws; without barbels. Mouth wide.
Villiform teeth in bands on the jaws and palatines, and in a A-shaped band on
the vomer.
Gill-openings wide : eight branchiostegals. Pseudobranchia? rudimentary
or absent.
Dorsal and anal fins confluent with the caudal. The ventrals arising far
apart, on distinct bony bases, about midway between the pectorals and the
clavicular symphysis : each consists of two filaments.
An air-bladder and pyloric caeca present.
76. Tauredophidium Hextli, Alcock.
Tauredophidium Hextii, Alcock, Ann. Mag. Nat. Hist., Sept. 1890, p. 213, pi. viii. fig. 1 : Illustrations of the
Zoology of the Investigator, Fishes, pl. XXI. fig. 3.
D. 64. A. 58. C. 10. P. 18. V. 2.
Head broad, pyramidal, its length about a fourth of the total, its bones
hard and firm. Greatest height of the body nearly equal to the length of the
head.
The operculum, which is a short narrow bone, is armed with a great thick
spine half as long as the head : at the angle of the preoperculum are three
similar spines, the middle and longest one of which is three-fourths the length
of the opercular spine. The occipital crest projects as a coarse subcutaneous
eminence, and behind it the first (?) neural spine projects similarly but more
strongly.
The eyes are completely atrophied ; the small orbital cavities are hidden
beneath thick scaly skin, and are filled with connective tissue, deeply imbedded
in which is a small pigmented eyeball about the size of an ordinary pin-head.
Nostrils large. Muciferous cavities of snout and mandible well developed and
opening to the exterior by pores. Mouth large, its cleft nearly horizontal ;
maxilla more than half the length of the head, much expanded behind, com-
pletely including the lower jaw in repose. Teeth in narrowish villiform bands
in jaws, vomer, and palatines. Ten long pointed scabrous gill-rakers on the
first brancial arch, besides some rudimentary ones above and below.
Scales in 22 rows between the dorsal fin and the vent.
Vertical fins united; the dorsal begins just behind the vertical through
the base of the pectoral, its longest rays — about the middle of the fin — are
rather over one-third the maximum body-height and exceed the corresponding
anal rays in length. Caudal long and pointed. Pectorals entire, pointed, as
lone as the head without the operculum. Ventrals separated from each other
by an interspace equal to one-third the length of the head ; each consists of two
13
98
filaments, of which the inner is much the longer, reaching beyond the origin of
the anal fin.
A bimch of about six slender ca?ca situated above the pylorus.
Colours in the fresh state : — uniform chocolate ; fins blackish ; throat and
belly black, owing to the pigmentation of the peritoneum.
The largest specimen is just over four inches long.
Bay of Bengal, off Ganjam coast, 1310 fathoms.
Regd. No. 12829, 12830.
Named after Rear- Admiral Sir John Hext, R. N., formerly Director of the
Royal Indian Marine, who was always a good friend to the Zoological depart-
ment of the " Investigator."
Diplacanthopoma, Giinther.
Diplacanthopoma, Giinther, Challenger Deep-Sea Fishea, p. 115 : Goode and Bean, Oceanic Ichthyology, p. 318.
Body elongate, compressed, covered with small thin imbricate scales. Head
broad, somewhat depressed, covered with a thick, glandular, scaleless skin,
sharply defined from the scaly skin of the trunk : its bones firm.
Lateral hue indistinct and incomplete (or absent?).
Snout broad, depressed, slightly overhanging the lower jaw ; without barbels.
Mouth wide : villiform teeth in bands on the jaws and palatines and in a A-shaped
band on the vomer.
Eye of moderate size.
Gill-openings wide : operculum with two radiating ridges each ending in a
spine : free edge of preoperculum rounded, unarmed. Eight branchiostegals (in
two species), but the branchiostegal membranes are so thick that the rays cannot
be counted without reflecting the skin. No pseudobranchiaa.
Dorsal and anal fins confluent with the caudal. The ventrals arise close
together a short distance behind the clavicular symphysis : each consists of a
single filament, which sometimes is made up of two (or more ?) rays intimately
fused together throughout their extent.
Air-bladder present. No pyloric cosca.
In one of the three known species of this genus the female is viviparous, and
in another species the male has a peculiarly elaborate penis.
Key to the species of Diplacanthopoma.
I. The two spines of the operculum are hidden in loose skin: length
of the adult female about six inches ... ... ... D. raniceps.
II. The two opercular spines project freely : —
1. Adult female about four inches long ... ... D. brachysoma.
2. Adult female about fifteen inches long ... ... D. Rivers-Andersoni.
99
Except in point of size, the species of Diplacanthopoma hardly differ from
one another.
77. Diplacanthopoma Rivers- Antler soni, Alcock.
Diplacanthopoma Rivers- Andersoni, Alcock, Ann. Mag. Nat. Hist. Ang. 1895, p. 144: Illustrations of the
Zoology of the Investigator, Fishes, pl. XVII. fig. 1.
Bight branchiostegals.
Head about a fourth the total length : greatest height of the body, at the
shoulder, equal to the length of the head without the snout.
Snout depressed, on a very much lower plane than the occiput, its length as
much exceeds that of the eye as it falls short of the width of the interorbital
space. Nostrils large : one immediately in front of the eye, the other near the
edge of the snout.
Major diameter of the eye between a sixth and a seventh the length of the
head.
Upper jaw half as long as the head, overlapping the lower jaw. Jaw-teeth
in broadish bands, palatine and vomerine teeth in narrowish bands.
The spines of the operculum project freely. Only three enlarged gill-
rakers on the outer side of the first arch, and those entirely in the upper half of
the arch.
The bases of the dorsal and anal fins are invested in a thick scaleless
glandular skin similar to that of the head. The anal begins a head length
behind the axilla. Pectorals and ventrals of no great length.
Each ventral fin consists of a single stout fluted filament, looking like two
or more rays intimately fused together.
Colours in spirit : body purplish brown ; head and fins much darker.
A single specimen from off the Indus Delta: length a little over 15 inches.
The specimen is a pregnant female : the ovaries open on a fleshy cushion,
behind the vent.
Regd. No. 14136.
The ovaries consist of a densely packed mass of embryos and ova enclosed
in a thin but extremely tough capsule. The capsule is abundantly supplied with
blood by the ramifications of a large branch of the mesenteric artery.
There is no attachment or adhesion of any kind between the ovarian capsule
and its contents.
The embryos form a thick surface layer immediately beneath the capsule,
enclosing a central mass of largish (a little over 1 millim. diameter) ova, which
consist entirely of yolk-spherules, without any trace of an embryo or even of a
germinal area.
100
Whether these unchanged ova would have developed subsequently to the
birth of the present superficial layers of embryos, or whether they were destined
for the ultra-ovarian nourishment of the present embryos, are questions which
it is impossible in an isolated case to discuss ; but from their large size, which
precludes any suggestion of immaturity, it would seem probable that they were
intended for present use rather than for a future brood.
The embryos, which are long and eel-like — 6 to 8 millim. long — lie matted
together, firmly adhering to one another by their tails, by means of a coagulated
secretion.
The vertical fins only are represented by a median fold of integument, which
runs from the occiput, round the tip of the tail, to the vent. This fold of integu-
ment consists of very numerous layers of large-nucleated cells. The remains
of the yolk-sac are enclosed in the abdomen, causing a bulging of the abdominal
wall along its whole length, from the throat to the vent ; but there is no vitelline
constriction or pedicle.
I am inclined to think that the vertical fold of the integument, which is
really only an extended sheet of embryonic cells, is an absorbent (nutritive)
surface, somewhat as in the embryos of certain fishes of the family Embiotocidse,
in which the interradial membranes of the vertical fins have been shown to act
as a foetal placenta.
In the present case, however, there is no vascular connexion, at any rate on
the foetal side ; and I am inclined to think that the nutrient material is absorbed
not so much from the thin tough ovarian capsule as from the ovary itself, perhaps
from those ova in which no trace of a germinal vesicle can be found.
An embryo taken at random measures 8 millim., namely 2 millim from the
snout to the vent and 6 millim. from the vent to the tip of the tail.
78. Diplacaiithopoma brachysoma, Giinther.
Diplacanthopoma brachysoma, Giinther, Challenger Deep Sea Fishes, p. 115, pi. xxiii. fig. C: Goode & Bean,
Oceanic Ichthyology p. 319: Alcock, Illustrations of the Zoology of the Investigator, Fishes, pl. XVII.
fig. 2, [specimen with an injured tail).
Differs from D. Rivers-Andersoni only in the following particulars : —
The branchiostegal rays appear to be only six in number.
The snout, though otherwise similar, is only as long as the eye.
The diameter of the eye is nearly a fourth that of the head.
The distance between the 1st anal ray and the axilla is less than the length
of the head.
The ventral fins arise nearer to the clavicular symphysis : though stout, they
are not fluted.
101
The colour is light brown fading on the head and belly : the fins are
grey.
The size is much smaller : a gravid female in the Museum collection is only
4- inches long.
Andaman Sea, 490 fathoms.
Regd. No. 11768.
Distribution : off Atlantic coast of Brazil, 350 fathoms : Andaman Sea, 490
fathoms.
79. Diplacanthopoma raniceps, Alcock.
Dipiacanthopoma raniceps, Alcock, Ann. Mag. Nat. Hist., Aug. 1898, p. 154: Illustrations of the Zoologt
of the Investigator, Fishes, pl. XXVI. fig. 2, 2a.
Differs from D. Rivers- Andersoni only in the following particulars : —
The head is conical, the slope from the occiput to the tip of the snout being
gentle and not abrupt, the whole head being frog-like.
The length of the snout is equal to the diameter of the eye, which is nearly
a fourth the length of the head.
The spines of the operculum hardly project through the skin.
The ventrals arise nearer the clavicular symphysis : each consists of a stout
filament, more than half as long as the head, and flattened and broadened
in the middle of its course : each is made up of at least two intimately-fused
rays.
The size is much smaller, a gravid female in the collection is not quite six
inches Ions;.
Colours in spirit as in D. Bivers-Andersoni.
Andaman Sea, 405 fathoms.
ry i AT 117 118 139
Regd. JNos. — , — , — .
The male is furnished with a most elaborate copulatory organ, which is
almost as long as the snout. It is hollow and is lined and strengthened by the
peritoneum, which is as thick and tough as leather. The intestine opens at its
base, and the testes are prolonged into its cavity. Its free end has almost the
consistence of cartilage and is thrown into several broad rigid lip-like folds
which inclose two deep cavities. From the smaller (anterior) cavity a pair of
papilla? project, and into the larger (posterior) cavity the testes open.
The corresponding organ of the female, though smaller than that of the
male, is large. It is a hollow cone, lined by peritoneum, and lodging the ends
of the ovaries, which open by a large common orifice at its spongy tip.
102
Subgenus Saccooastek.
Saccogaster, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 389.
Differs from Diplacanthopoma, which it very closely resembles, only in the
following characters : —
The scales are not, or hardly, imbricate, so that the peculiar scale-less skin
of the head is not so abruptly demarcated from the integument of the body.
A row of teeth in the mandible, and a few teeth near the symphysis of the
upper jaw are enlarged.
The operculum has two radiating ridges but they do not end in spines.
The only known species is viviparous.
80. Diplacanthopoma {Saccogaster) maculatum, Alcock.
Saccogaster maculata, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 389 ; and July 1891, p. 30, pi. vii. fig. 3 :
Proc. Zool Soc, 1891, pp. 226, 227, fig: Illustrations of the Zoology of the Investigator, Fishes, pl. XXIX.
fig. 2.
Eight branchiostegals.
Length of head 3- to 4 in the total : greatest body-height (except in the
pregnant female) equal to the postorbital portion of the head.
Snout depressed, not overhanging the lower jaw, its length is half again
that of the eye and equal to the width of the interorbital space. Nostrils incon-
spicuous ; one in front of the eye, the other near the edge of the snout.
Diameter of the eye about a tenth the length of the head.
Upper jaw half as long as the head.
Two or three gill-rakers on the outer side of the first branchial arch are
somewhat enlarged, as in Diplacanthopoma.
The bases of the dorsal and anal fins are invested in a thick scaleless srlandu-
lar skin. The first anal ray is a head-length behind the axilla.
The fleshy bases of the pectorals are particularly long : the fin is about as
long as the postorbital portion of the head. The ventrals arise a short distance
behind the clavicular symphysis ; each consists of a single filament not half as
long as the head.
Colours : light brown, the scales showing as white dots, head and fins
darker.
The largest specimen is 4 inches long.
Bay of Bengal, 193, 240, 145 to 250, and 195 to 210 fathoms.
Regd. Nos. 11073, 11674, 13045, 13527, 13528.
103
The male is furnished with a penis that consists of a large bilobed papilla
with a pore, or genital opening, between the lobes. Sometimes there is also a
long filament between the lobes.
The female is viviparous as in D. Bivers-Andersoni.
Hephthocara, Alcock.
Hephthocara, Alcock, Ann. Mag. Nat. Hist., Nov. 1892, p. 349.
Body elongate, compressed, tail tapering to a filament, covered with small
thin deciduous slightly imbricate scales. Head large and broad, covered with
a thick gelatinous scaleless skin ; its bones thin.
Lateral line indistinguishable.
Snout broad, depressed, not overhanging the jaws ; without barbels.
Mouth wide, with oblique cleft ; the lower jaw slightly prominent. Villi-
form teeth in the jaws and palatines and in a crescent on the vomer.
Bye of moderate size.
Gill-openings wide : operculum with two feeble radiating ridges, the upper
of which ends in a spine. Eight branchiostegals. No pseudobranchise.
Dorsal and anal fins confluent with the caudal.
No ventral fins.
Air-bladder present. No pyloric casca.
The only known species is viviparous.
Hephthocara differs from Diplacanthopoma, to which it is very closely
related, in the absence of ventral fins and in having a long lash-like tail.
81. Hephthocara simum, Alcock.
Hephthocara simum, Alcock, Ann. Mag. Nat. Hist., Nov. 1892, p. 349, PL. xviii. fig. 1 : Illustrations of the
Zoology of the Investigator, Fishes, pl. XXII. fig. 3.
Head large, deep, broad, much inflated posteriorly ; its length is from 4 to
5 in the total ; its greatest height, which is more than the greatest height of the
body, is nearly equal to its length ; its bones are wafer-like and smooth ; its
integument is smooth and scaleless and, in life, forms a thick mucous cap of
gelatinous consistence.
The small snub snout, the end of which is formed by the projecting man-
dible, is equal in length to the width of the interocular space, this being rather
more than twice the major diameter of the deep-set eye, which again is
about one-seventh the length of the head. The nostrils are inconspicuous and
are situated one in front of the angle of the eye, the other at the tip of the snout.
Mouth large, with its cleft oblique, and with the mandible projecting beyond
the thin broad maxilla, which last is a little more than half as long as the head.
104
Villiform teeth in broaclisk bands in the premaxillaa and mandible, and in very
narrow bands on the palatines and expanded head of the vomer.
The edge of the preopercnlum is smooth. As in Diplacanthopoma and
Saccogaster there are only three enlarged gill-rakers on the outer side of the
first branchial arch, and these are in the upper half of the arch.
The fin-rays are all extremely delicate ; the dorsal fin, which begins about
a snout-length behind the level of the gill-opening, and the anal, which begins
nearly a head-length behind the same level, are confluent with the caudal at its
base. The narrow pointed pectorals are a little longer than the combined
eye and snout. There are no traces of ventrals.
Colour : uniform sepia, fins black.
The largest specimen, which is a gravid female, is a little over 11 inches
long.
Off Travancore coast, 824 and 902 fathoms : Gulf of Manar, T 58' N„ 937
fathoms : Bay of Bengal, near the Andaman Islands, 606 fathoms.
T> l AT ioi»7<i 339 340 372 590
Regd. Nos. 13172, — , — , —, T.
This species is viviparous in almost exactly the same way as Diplacantho-
poma Rivers-Andersoni.
Family Bfacruvidm.
Head large, with the muciferous cavities well developed ; trunk short :
tail long, compressed, gradually tapering to a filament. Scales present on
the body and generally on the head. Two dorsal fins, the first, which
arises just behind the head, being short ; the second, which arises either
immediately or a short distance behind the first, being continued to the
tip of the tail. Anal nearly similar in extent to the second dorsal. No
caudal. Ventral fins thoracic or jugular, composed of several (6-12) rays.
No pseudobranchiae. Six or seven branchiostegals. Air-bladder present.
Pyloric appendages numerous.
This family has been added to our knowledge of the Indian Fauna by the
researches of the " Investigator." Eighteen Indian species, belonging to two
genera, are now known. Of these, thirteen seem to be peculiar to Indian waters,
and five occur in other seas. The five that are found elsewhere are Macrwms
(Coelorhynchus) parallehts and Macrurus nasutus from Japan; Macrurus (Mysta-
conurus) cavernosas (probably identical with Macrurus italicus of the Mediterra-
nean) and IJnllii/ijiiihis Itmi/iflis which are also known from the West Indian and
Madeiran regions ; and Macrurus (Malacocephahis) laevis from the North Atlantic
and Brazil coast.
105
Key to the Indian genera of the family Macruridae.
I. Snout projecting beyond the mouth : rays of the anal fin better
developed than those of the second dorsal : gill-rakers on the outer
side of the first branchial arch tuberculiforru ... ... Macrurus.
II. Snout not projecting beyond the mouth : rays of the second dorsal
fin better developed than those of the anal : gill-rakers on the outer
side of the first branchial arch long and setiform ... ... Bathtgadus.
Macrurus, Bloch, Giinther.
Macrurus, Giinther, Challenger Deep-Sea Fishes, pp. 122-124.
Macrurus, Coelorhynchus, Coryphaenoid.es, Hymenocephalus, Lionurus, Trachonurus, Cetonurus, Chalinura,
Optonurus, Malacocephalus, Nematonurus, Moseleya, Abyssicola, Goode and Bean, Oceanic Ichthyology, pp. 390-417.
Malacocephalus, Moseleya, Nematonurus, Albatrossia, Bogoslovius, Chalinura, Coryphaenoides, Hymenocephalus,
Macrourus, Coelorhynchus, Trachonurus, Lionurus, Jordan and Evermann, Fishes N. America, pp. 2569-2592.
Head short and thick with the nmciferous cavities well developed ; tail long
and tapering.
The snout usually projects well beyond the mouth, which may be either
quite inferior or lateral and subterminal. Teeth in the jaws only. A barbel at
the symphysis of the lower jaw.
Gill-openings wide, the gill-membranes slightly united in front. The first
branchial arch is broadly connected by membrane with the outer wall of the gill-
chamber, so that the slit between that arch and the gill-cover is very much
narrower than the other branchial clefts. The gill-rakers on the outer side of the
first branchial arch are mere scabrous tubercles.
Scales either strongly and typically ctenoid or, more rarely, cycloid. No
scaleless fossa on the side of the nape.
Second dorsal fin separated from the first by an interval ; its rays, or at least
its anterior rays, are more or less rudimentary. Anal rays much better develop-
ed than those of the second dorsal. Ventrals arising below, or slightly in
advance of, the pectorals.
Key to the Indian subgenera of Macrurus (adapted from Dr. Giinther).
I. Villiform teeth iu bands in both jaws : scales imbricate and spini-
gerous : —
1. Dorsal spine serrated : mouth inferior ... ... Macrorus.
2. Dorsal spine smooth : —
i. Mouth inferior ; the infraorbital region divided
into a vertical and a subhorizontal portion by a
longitudinal ridge ... ... ... Coelorhynchus.
ii. Mouth wide, in the ordinary lateral position ... Mystaconurus.
II. Teeth in two series in the upper jaw and in a single series in the
lower jaw. [Dorsal spine smooth : mouth lateral : scales imbricate
and spiuigerous] ... ... ... ... .,. Malacocephalus.
14
106
Subgenus Ccelorhynchus, Gunther.
Mouth comparatively small, entirely inferior, on the lower side of the head.
a longitudinal ridge dividing the infraorbital region into a vertical and a sub-
horizontal portion. Teeth in bands in both jaws . Scales imbricate, with strong
serrated keels ; no series of enlarged dorsal scales. Dorsal spine smooth.
Key to the Indian species of the subgenus Ccelorhynchus.
I. Scales of the body with subparallel spiny ridges: pyloric cneca 12
in number : seven branchiostegals ... ... •■• M. parallelus.
II. Scales of the body with radiating spiny ridges : pyloric easca about.
40 in number : six branchiostegals : —
1. Body-scales with 5 spiny ridges: 6 to 6| series of scnles
between the first dorsal fin and the lateral line ... M. quadrici-istntns.
2. Body-scales with 8 or 9 spiny ridges : 4 to 5 series of scales
between the first dorsal fin and the lateral line ... M. flabellispinis.
82. ? Macrurus (Ccelorhynchus) parallelus, Gimther.
Macrurus (Ccelorhynchus) parallelus, Gunther, ChalleDger Deep-Sea Fishes, p. 125, pi. XXIX. fig. A : Alcock,
Ann. Mag. Nat. Hist., (6) IV. 1889, p. 391, and Jonrn. As. Soc, Bengal, Vol. LX1II. pt. 2. 1894, p. 126.
B. 7. D. 10. A. 90. P. 16. V. 7.
Scales with usually 5 spiny ridges which are nearly parallel to one another,
and of which the middle one is the strongest.
Twelve pyloric caaca.
Two young specimens in extremely bad preservation possess the above cha-
racters, and are therefore referred, though with hesitation, to M. parallelus Gthr.
Gulf of Manar, 597 fathoms.
Regd. Nos. 11760, 11761.
Distribution : New Zealand waters ; Japan ; Gulf of Manar.
83. Macrurus (Cailorhyriclius) quadricristatus, Alcock.
Macrurus (Calorhynchus) quadricristatus, Alcock, Ann. Mag. Nat. Hist., Angnst, 1891, p. 119: Jonrn. As. Soc.
Bengal, Vol. LXIII. pt, 2, 1894, p. 126: Illustrations of the Zoology of the Investigator, Fishes, pl. HI.
fig. 1.
B. 6. D. 11. A. circ. 90. P. 16. V. 7.
Head much exceeding the trunk in all three dimensions: tail low, compress-
ed, tapering.
Length of the head nearly a third the total : greatest height of the body
barely equal to the length of the snout.
Length of the snout nearly half that of the head, twice the major diameter
of the eye and twice the width of the interorbital space.
107
The suborbital crest is strongly salient and serrated and terminates acutely
at the preopercular angle. The posterior half of the head is longitudinally
traversed on each side by two strongly serrated ridges, one extends from the
interorbital space to the occiput, the other from the supra-orbital ridge to the
shoulder.
Nostrils situated immediately in front of the eye ; the posterior is very
large.
The mouth is a small, completely inferior, crescentic orifice ; its front limit
is in the vertical through the anterior nostril, and the maxilla reaches a little
behind the vertical through the middle of the eye. Villiform teeth in bands in
the jaws, the outer row in the upper jaw slightly enlarged. Barbel slender, less
than half the eye in length.
Gill-rakers in the form of tubercles.
Scales of the head with about 3, scales of the body with about 5, thin salient
slightly-divergent serrated crests. 6 or 6- series of scales between the last ray
of the first dorsal fin and the lateral line.
The first spine of the first dorsal fin is very small, the second is smooth
throughout. The interval between the first and the very inconspicuous second
dorsal is hardly half the extent of the base of the first. Pectorals narrow and
pointed, their length slightly exceeds that of the postorbital portion of the head.
Ventrals with the outer ray prolonged well beyond the origin of the anal.
Stomach large, siphonal ; very many long slender caeca in a thick cluster
round the pylorus ; a thin- walled air-bladder.
Colours in life : — Chocolate ; body and tail with numerous broad black cross
bands, which do not reach the mid-abdominal line. In spirit the colouration is
unchanged, but much lighter.
The largest specimen is 7 inches long.
Andaman Sea, 185, 188 to 220, and 405 fathoms.
Registered Nos. 13070-13072, 5£
84. Macrurus (CcelorhynchusJ fiabellispinis, Alcock.
Macrurus (Cailorhynchus) fiabellispinis, Alcock, Journ. As. Soc. Bengal, Vol. LXIII. pt. 2, 1894, pp. 123, 126.
Illustkations of the Zoology of the Investigator, Fishes, pl. XVI. figs. 2,2a.
B. 6. D. 1/8. A. 95. P. 16. V. 7.
Differs from M. quadricristatus in the following particulars : —
(1) the length of the snout is about two-fifths that of the head :
(2) the suborbital crest ends bluntly, and the four crests on the nape are
low and inconspicuous :
108
(3) the mouth is much larger, the maxilla reaching almost to the posterior
border of the orbit :
(4) the scales of the head have from 3 to 8, those of the body have 8 or 9,
strongly -radiating serrated crests; and there are 4 or 4- rows of scales between
the last ray of the first dorsal fin and the lateral line :
(5) the interval between the two dorsal fins is longer than the base of the
first :
(G) the pectorals are decidedly shorter than the postorbital portion of the
head, and the prolonged first ventral ray does not reach to the anal fin.
There are about 40 long pyloric caeca, and there is a thin- walled air-bladder.
Colours : dark stone-grey ; fins and gill-membranes black.
Length 19 inches.
Arabian Sea, 719 fathoms.
Reffd. No. 13472.
■^£>v
Subgenus Maceueus, Gthr.
Mouth small or moderate, entirely inferior, the infraorbital ridge distinct.
Teeth in bands in both jaws. Scales imbricate with spines or serrated keels :
no series of enlarged dorsal scales. Dorsal spine serrated.
The following key is substituted for that given by me in J. A. S. B. Vol.
LXIII. pt. 2, 1894, pp. 126-127, which was reprinted by Goode and Bean in
Oceanic Ichthyology, p. 532.
Key to the Indian species of the subgenus Macrurus.
I. Seven branchiostegals : vent between the ventral fins: —
1. Eight rays in the ventral fin : —
i. Scales with strong serrated (parallel) ridges: barbel hardly
half as long as the eye ... ... ... ... M. investigatoris.
ii. Scales with spinelets : barbel at least as long as eye ... M. J'etersoni.
2. Ten rays in the ventral fin : scales with densely packed spinelets M. nasntus.
:>. Eleven or twelve rays in the ventral fin : —
i. Length of the head about a fifth of the total : —
a Scales witli about fifteen series of capillary spinelets M.semiquincunciatus.
h. Scales with about seven series of capillary spinelets M. polylepis.
ii. Length of head about an eighth the total: scales with from
three to eight (usually six) series of short spinelets ... M. pumiliceps.
II. Six brancliiostegals : vent immediately in front of the first ray of the
nnal fin : —
1. Seven lays in the ventral fin ... ... ... M. Hextii.
109
2 Eight (rarely 9) rays in the ventral fin: —
i. Second spine of the first dorsal fin remarkahly prolong-
ed :—
a. Scales with twelve to seventeen rows of spinelets ... M. macrolophus.
b. Scales with five or six very short series of spinelets M. lirphotes.
ii. Second spine of the first dorsal fin nut abnormally long ... M. Wood-Masoni.
3 Nine rays in the ventral fin : one or more of the spinelets of the
middle series in each scale much enlarged ... ... M. Hoskynii.
The species of Macrurus so much resemble each other, that it will be suffi-
cient to describe one species and then to give only the differential points of the
others, leaving the points of agreement to be understood. The commonest of
the Indian species — M. investigatoris — may be first described.
85. Meicrurus (Meter urus) investigatoris, Alcock.
Macrurus investigatoris, Alcock, Ann. Mag. Nat. Hist., Not. 1889, p. 391 : Journ. As. Soc. Bengal, Vol. LXIII.
pt. 2, 1891, p. 126. Illdstrations of the Zoology of the Investigator, Fishes, pl. III. fig. 4.
B. 7. D. r+ 11. A. circ. 100. P. 20-21. V. 8 (rarely 9).
Length of the head rather over a fifth of the total. Greatest height of the
body (at the eminence of the first dorsal fin) equal to the length of the head
behind the anterior nostril.
Snout almost as long as the eye, overlapping the mouth ; with 3 well
developed tubercles (median and lateral) : nostrils in a scaleless fossa, the
posterior much the larger.
Major diameter of the eye 3^ in the length of the head: width of inter-
orbital space about two-thirds the length of the eye.
Mouth rather small, completely inferior : the upper jaw, which overlaps the
lower and is about as long as the snout, about reaches to the middle of the eye.
Teeth in the jaws in broadish bands, none of the teeth enlarged.
Barbel slender, hardly half as long as the eye.
Scales on all parts except the mouth, throat, and gill-membranes. Scales
of the body of moderate size; each with from 9 to 13 quite similar, nearly
parallel, longitudinal ridges which project slightly — sometimes not at all —
beyond the edge of the scale. To the naked eye the ridges appear entire, but
they are really finely serrated.
Five rows of scales between the last ray of the first dorsal fin and the
lateral line.
First spine of the first dorsal fin rudimentary ; second spine about as long
as the head without the snout.
Pectorals pointed ; as long as the head behind the middle of the eye.
Outer ray of the ventral fins filamentous, slightly longer than the pectorals.
no
Vent situated between the ventral fins. About twelve long filiform pyloric
caeca.
Distinctive colours : first dorsal fin black with white root and tip.
Bay of Bengal 193, 240, 270, and 320 to 296 fathoms : Andaman Sea, 265,
271, 405, 490 fathoms.
Regd. Nos. 11654, 11655, 11658, 11676, 11679, 11766, 11772, 12447, 13177.
47 4S 120 123 145 148 152 158
T' 7' Is l ' X' l ' 1 ' T-
86. Macrurus (Macrurus) Petersonii, Alcock.
Macrurus petersonii, Alcock, Ann. Mag. Nat. Hist., Aug. 1891, p. 121 : Journ. As. Soc. Bengal, Vol. LXIII. pt.
2, 189-t, p. 127: Illustrations of the Zoology of the Investigator, Fishes, pl. III. fig. 5.
? Macrurus (Chalinurus) hispidus, Aluock, Ann. Mag. Nat. Hist., Nov. 1880, p. 397 : Illustrations of the
Zoology of the Investigator, Fishes pl. xiii. fig. 2 (is probably the young of M. Petersonii).
B. 7. D. r + 10. A. circ. 100. P. 18-20. V. 8.
Differs from M. investigatoris only in the following particulars : —
The snout is quite as long as the eye, and its tubercles are not very much
pronounced.
The eye is about a fourth the length of the head, and very little more than
the width of the interorbital space.
The mouth is large, and the upper jaw, which is much longer than the
snout, reaches behind the middle of the eye. The outer row of teeth in the
upper jaw are enlarged.
The barbel is at least as long as the eye.
The scales are covered, but not very densely, with sharp conical spinelets,
not in rows, some of which project beyond the edge of the scale. Some scales
along the edge of the gill-opening, and in a patch immediately behind the first
dorsal fin, are quite smooth.
Six rows of scales between the last ray of the first dorsal fin and the lateral
line.
Second dorsal spine as long as the head behind the middle of the eye.
Length of pectorals about equal to the postorbital portion of the head.
Outer ventral ray hardly prolonged, its length not much more than two-
fifths that of the head.
Over twenty longish vermiform pyloric caeca of great delicacy.
Distinctive colouration : head and iris silvery ; first dorsal black with white
base and tip.
Length of adult, 9- inches.
Andaman Sea, 185 and 188 to 220 fathoms.
Ill
Named after Mr. B. Peterson, for many years gunner of the Investigator,
who in his zeal for zoology, was once nearly wound round the surging-drum of
the dredsinsr-winch, and once fell overboard anions' sharks.
Regd. Nos. 13117, 13118, 5±±* 3-f8.
87. Macrurus (Macrurus) uasutus, Gtinther.
Macrurus nasutus, Giinther, Ann Mag. Nat. Hist., 1877, Vol. XX. p. 440: Challenger, Deep-Sea Fishes, p. 132,
PI. XXX. fig. B.
Macrurus brevirostris, Alcock, Ann. Mag. Nat. Hist. (6) IV. 1889, p. 393 ; Journ. As. Soo. Bengal, Vol. LXIII.
pt. 2, 1894, p. 127. Illustrations of the Zoology of the Investigator, Fishes, pl. XIII. fig. 3.
B. 7. D. r+11. A. arc. 110. P. 20-21. V. 10.
Differs from M. investigatoris only in the following particulars : —
The head is between a fifth and a sixth of the total.
The median tubercle of the snout is prominent, but the lateral tubercles are
indistinct.
Major diameter of eye about 3- in the length of the head.
Outer row of teeth of the upper jaw considerably enlarged.
Barbel decidedly more than half as long as the eye.
Scales very closely covered with sharp conical spinelets of equal size, not
arranged in rows, some of them projecting beyond the edge of the scale.
Five and a half or six rows of scales between the last ray of the first dorsal
fin and the lateral line.
2nd dorsal spine a little longer than the head.
About 35 large long pyloric caeca.
Largest specimen, 11 inches long.
Andaman Sea, 490 fathoms : Arabian Sea, off Travancore coast, 738
fathoms.
Regd. Nos. 11762, 13115.
(In the Indian Museum is also one of the " Challenger " duplicates from
Japan).
88. Macrurus (Macrurus) semiquincunciatus, Alcock.
Macrurus semiquincunciatus, Alcoek, AnD. Mag. Nat. Hist., Nov. 1889, p. 392 ; Jonrn. As. Soc. Bengal, Vol.
LXIII. pt. 2, 1894, p. 127. Illustrations of the Zoology of the Investigator, Fishes, pl. XII. fig. 2.
B. 7. D. r+11. A. arc. 100. P. 21. V. 11.
Differs from M. investigatoris only in the following particulars : —
The median tubercle of the snout is not very prominent and the lateral ones
are very indistinct.
112
The major diameter of the eye is 3g in the length of the head.
Outer row of teeth in the upper jaw considerably enlarged.
Barbel stout, as long as the eye.
Scales of the body small, densely covered with sharp capillary spinelets
some of which project far beyond the edge of the scale.
Eight rows of scales between the last ray of the first dorsal fin and the
lateral line.
2nd spine of the dorsal fin as long as the head.
Twenty-two long vermiform pyloric caeca.
Largest specimen 10 inches long.
Andaman Sea 130 to 250 fathoms : Bay of Bengal, 240 and 410 fathoms.
Kegd. Nos. 11660, 13133, 13173.
Palatksy, quoted by Goode and Bean in Oceanic Ichthyology p. 531, has
changed the name of this species to M. sesqiricnnciatus. The word sesqui-
cunciatus, if it has any meaning, might mean 'an ounce and a half.' The name
semiquincunciatus was meant to express the fact that the rows of spinelets on the
scales are arranged like the principes and hastati of the Roman legion, — that is
to say in quincuncem, but without the triarii, or in a semi-quincunx. The idea
may perhaps be a little far-fetched, but the etymology appears to me to be
perfectly correct.
89. Macrurus (Macrurus) polylepis, Alcock.
Macrurus polylepis, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 395; Jonrn. As. Soc. Bengal, Vol. LXIII.
pt. 2, 1894, p. 127. Illustrations of the Zoology of the Investigator, Fishks, pl. XXIX. fig. 4.
B. 7. D. r+11. P. 19. V. 11-12.
Differs from M. investigatoris only in the following particulars : —
The greatest body-height (at the eminence of the first dorsal fin) is only
equal to the length of the head behind the middle of the eye.
Snout decidedly shorter than the eye, the median tubercle far more distinct
than the lateral ones.
Major diameter of the eye nearly a third the length of the head, and nearly
twice the width of the interorbital space.
Though the mouth is not large, the upper jaw is much longer than the
snout and reaches behind the middle of the eye.
Outer row of teeth in the upper jaw much enlarged.
Barbel about as long as the eye.
Scales small, with about seven series of sharp capillary spinelets, some of
which project beyond the edge of the scale.
113
Bight series of scales between the last ray of the first dorsal fin and the
lateral line.
[2nd dorsal spine and pectoral fins broken at tip].
Outer ray of the ventral fin as long as the head behind the middle of the eye.
[Pyloric ca3ca present, of moderate size].
Largest specimen (immature) just over 6 inches long.
Bay of Bengal, 193 and 272 fathoms.
Reffd. Nos. 11678, 11724.
"S^
90. Macrurus (Macrurus) pwmilicepsf Alcock.
Hacrurus pumiliceps, Alcock, Journ. As. Soc. Bengal, Vol. LXIII. pt. 2, 1891, pp. 125, 127: Illustrations of
thf. Zoology of the Investigator, pl. XVI. fig. 3.
B. 7. D. r+12. P. 18. V. 12.
Differs from M. investigatoris only in the following particulars : —
The head is just over an eighth the total length.
The greatest height of the body is equal to the length of the head.
The major diameter of the eye is almost a third the length of the head.
The upper jaw is a little longer than the snout.
The barbel is about three-quarters as long as the eye.
Scales with from 3 to 8 (usually 6) short rows of spinelets, which seldom
project beyond the edge of the scale.
Lateral line indistinguishable.
2nd dorsal spine as long as the head.
The pectorals and the outer ray of the ventral fin are as long as the head
without the snout.
Nine or ten very small pyloric ca?ca.
Largest specimen 11 inches long.
Arabian Sea, between Maldives and Travancore coast, 719 fathoms.
Regd. Nos. 13561, 13562.
[The number of rays in the anal fin is far over 100].
91. Hacrurus (3Iacrurus) Itextii, Alcock.
Macrurus Seitii, Alcock, Ann. Mag. Nat. Hist., Oct. 1890, p. 299 ; Jonrn. As. Soc. Bengal, Vol. LXIII. pt. 2,
1894, p. 126. Illustrations of the Zoology of the Investigator, Fishes, pl. XII. fig. 3.
B. 6. D. r+11. A. arc. 110. P. 21. V. 7.
Differs from M. investigatoris only in the following particulars : —
The snout is as long as the eye, and its tubercles are inconspicuous.
15
114
The major diameter of the eye is 4j to •£> in the length of the head and
equal to the width of the interorbital space.
Month rather large, the upper jaw, which is as long as the snout plus half
the eye, extending behind the middle of the eye.
Outer row of teeth in the upper jaw considerably enlarged, in the lower jaw
slightly enlarged.
Barbel stout, three-fourths as long as the eye.
Scales closely covered with tiny capillary spinelets which are deciduous and
do not project beyond the edge of the scale.
[Five rows of scales between the last ray of the first dorsal fin and the
lateral line, as in M. investigatoris.]
Outer ventral ray as long as the head behind the anterior nostril.
Vent far behind the tip of the mass of the ventral fins.
Fourteen or fifteen very large pyloric casca.
Length of largest specimen nearly 23 inches.
Arabian Sea, between the Laccadives and the Malabar coast, 360 and 1000
fathoms.
Regd. No. 12866. 251.
The structure of the ovary and of the maturing ova in this species, have been
described by me in Ann. Mag. Nat. Hist., Nov. 1892, pp. 351, 353.
92. Macrurus (Macrurus) Wood-Masoni, Alcock.
Macrurus Wood-Masoni, Alcoek, Ann. Ma?. Nat. Hist., Oct. 1830, p. 301 ; Nov. 1892, p. 353: Jonrn. As. Soc.
Bengal, Vol. LXI11. pt. 2, 1894, p. 126. Illustrations of the Zoology of the Investigator, Fishes, pl. XIII.
i ii. 1.
B. 6. D. r + 10. A. arc. 105. P. 21. V. 8.
Differs from M. investigatoris only in the following particulars : —
The snout is about quarter again as long as the eye ; all its tubercles are
indistinct.
Major diameter of the eye 4- to 47, in the length of the head, and very
slightly more than the width of the interorbital space.
Mouth rather large, the upper jaw slightly longer than the snout. Outer
row of teeth in the upper jaw slightly enlarged.
Barbel a mere papilla hardly a quarter the length of the eye.
Scales covered with numerous close-set series of tiny spinelets.
Four or live rows of scales between the last ray of the first dorsal fin and
the lateral Line.
115
Outer ventral ray considerably shorter than the postorbital portion of the
head.
Vent situated far behind the tip of the ventral fins.
11 or 12 long large pyloric caeca.
Largest specimen just over 19 inches long.
[The end of the 2nd dorsal spine is broken in all the specimens].
Arabian Sea, between the Malabar coast and the Laccadives, 360, 559, 902
and 1000 fathoms.
r> i XT -ino*?^ 356 358-360
Regd. Nos. 12867, — , — j — .
93. Macrurus (Macrurus) macrolophus, Alcock.
Macrurus macrolophus, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 394; August, 1891, p. 121; Nov. 1892'
p. 351, 352, fig. 1 : Journ. As. Soc. Bengal, Vol. LXIII. pt. 2, 1894, p. 126. Illustbations of the Zoology of
the Investigator, Fishes, pl. XII. FIG. 1 {the tail is a healed " stump" in this figure).
B. 6. D. r + 9. A. cite. 85. P. 20-21. V. 8.
Differs from M. investigatoris only in the following particulars : —
The head is about two-ninths of the total length.
The greatest height of the body is equal to the postrostral portion of the
head.
The snout is decidedly longer than the eye, and the lateral tubercles are
indistinct.
Major diameter of the eye about 4- in the length of the head.
Outer row of teeth in upper jaw slightly enlarged.
Barbel very inconspicuous, not a third as long as the eye.
Scales with 12 to 17 rows of spinelets, the rows converging towards the
middle line of the scale, the last in each row projecting far beyond the edge of
the scale.
[Five rows of scales between the last dorsal ray and the lateral line, as in
M. investigatoris].
Second spine of the first dorsal fin much prolonged, — 1\ to 1? times as long
as the head.
Pectorals as long as the postorbital portion of the head.
Outer ventral ray about as long as the head behind the middle of the eye.
Vent behind the mass of the ventral fins.
About 10 or 11 slender pyloric casca.
The largest specimen (adult) is 11 inches long.
116
Bay of Bengal, 240 and 410 fathoms : Andaman Sea, 265 and 370 to 419
fathoms : Arabian Sea, off Travancore and Malabar coast, 360 and 406 fathoms.
Regd.Nos. 11776,13130, 13175, ?-f, 3±9 - 3|i, »
94. Macrurus (Macrurus) lophotes, Alcock.
Macrurus lophotes, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 385; Jonrn. As. Soc. Bengal, Vol. LX1II.
pt. 2, 1894, p. 126. Illustrations of the Zoology of the Investigator, Fishes, pl. III. fig. 2.
B. 6. D. r + 11-12. V. 8-9.
Differs from M. investigatoris only in the following particulars : —
The greatest height of the body is equal to the length of the head without
the snout.
The snout is nearly half again as long as the eye.
The barbel is not a quarter the length of the eye.
Outer row of teeth in the upper jaw slightly enlarged.
Scales small, with 5 or 6 very short series of spinelets, the last in each series
projecting beyond the edge of the scale.
Five or six rows of scales between the last ray of the first dorsal fin and the
lateral line.
2nd dorsal spine nearly twice as long as the head.
Vent situated behind the mass of the ventral fins.
The number of pyloric ca?ca is unknown, the specimens being far too much
damaged for dissection.
Length of the immature specimens, five inches.
Bay of Bengal " Swatch of No-ground " 405 to 285 fathoms.
Regd. Nos. 11670, 11071.
95. Macrurus {Macrurus) Hoskynii, Alcock.
Macrurus Hoskynii, Alcock, Ann. Mag. Nat. Hist., Sept. 1890, p. 214: Jonrn. As. Soc. Bengal, Vol. LXIII.
pt. 2, 1894, p. 126. Illustrations of the Zoology of the Investigator, Pishes, pl. IX. fig. 4.
B. 6. D. r+10. P. 19-20. V. 9.
Differs from M. investigatoris only in the following particulars : —
Snout slightly but distinctly longer than the eye.
Major diameter of the eye 4- in the length of the head : interorbital space
one-fourth again as broad as the major diameter of the eye.
Outer row of teeth in the upper jaw slightly enlarged.
Barbel not a fourth as long as the eye.
117
Scales with 13 to 18 short series of acute conical spinelets, the last spinelet
of the middle series greatly enlarged above the others.
Four rows of scales between the last ray of the first dorsal fin and the
lateral line.
2nd dorsal spine almost as long as the head.
Outer ventral ray as long as the head behind the middle of the snout.
Vent situated behind the tips of the mass of the ventral fins. Nine pyloric
ca?ca.
One specimen just over 14 inches long. In the fresh state it emitted a
powerful and disagreeable musky odour.
Bay of Bengal, 1310 fathoms.
Regd. No. 12833.
Subgenus Mystaconurus, Giinther.
Mouth wide, in the ordinary lateral position, the infra-orbital ridge quite
indistinct and not marking off two distinct planes. Teeth in bands in both jaws.
Scales imbricate, spinigerous ; no series of enlarged dorsal scales. Dorsal spine
smooth.
96. Macrurus (Mystaconurus) cavernosus, Goode and Bean.
[an Macrurus (Mystacomirus) italicus, Giglioli.]
Bathygadus cavernosus, Goode and Bean, Proo. U. S. Nat. Mas. VIII. 1885, p. 598 : Giinther, Challenger Deep
Sea Fishes, p. 156.
Symenocephalus cavernosus, Goode and Bean, Oceanic Ichthyology, 1896, p. 408, fig. 341.
Macrurus (Mystacomirus) heterolepis, Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 396; Ang. 1891, p. 122:
Jonrn. As. Soc. Bengal, Vol. LXV. pt. 2, 1896, p. 309. Illustrations of the Zoology of the Investigator,
Fishes, pl. III. fig. 3.
B. 7. D.r+ 10-11. A. circ. 100. P. 13-14. V. 10-11.
Length of the head not quite a fifth of the total : greatest height of the body
not quite equal to the length of the head without the snout : tail lash-like.
Snout about three-quarters as long as the eye, distinctly but not very
greatly overhanging the mouth, with median and lateral angles but not tubercles.
Nostrils as in all the previous species.
Major diameter of the eye about 3- in the length of the head and about
four-fifths the width of the interorbital space.
Mouth wide, the upper jaw, which is about half as long as the head, reach-
ing the level of the posterior border of the orbit. Villiform teeth in rather
narrow tapering bands in both the jaws : none of the teeth enlarged.
118
Barbel very slender and inconspicuous, not half as long as the eye.
Scales apparently absent from the head ; large thin and very deciduous on
the body. The scales on the anterior half of the trunk and posterior half of the
tail are smooth and cycloid ; but between these parts — at any rate in adults — the
scales bear 6 or 7 series of rather distant granules or semi-erect spinules.
Seven rows of scales between the last ray of the first dorsal fin and the
lateral line.
1 st dorsal spine rudimentary ; the second is as long as the head without the
snout.
Pectorals narrow, pointed ; about as long as the postorbital portion of the
head.
Outer ray of the ventral fins filamentous, nearly as long as the head without
the snout.
Vent situated behind the mass of the ventral fins, immediately in front of
the 1st anal ray. 12 or 13 not very long pyloric caaca.
Colours in life : head and iris silvery, body pinkish brown with a silvery
sheen, throat and belly black : first dorsal white, with a black patch about the
middle.
Length between 6 and 7 inches.
In some specimens there is a curious circular spot or pimple in the middle
line of the belly in front of the ventral fins.
Andaman Sea, 188 to 220, 240, 265, 271 and 405 fathoms : Gulf of Manar,
180 to 217 fathoms.
Regd. Nos. 11644, 11774, 12449, 13551-13560, y - £ lf\ if6, if7.
Distribution: G. of Mexico, 227 fathoms : G. of Manar and Andaman Sea,
180 to 405 fathoms.
I am satisfied by actual comparison of our specimens with one received from
the Smithsonian Institution (and now in the Indian Museum collection), that the
species described by me as Mystaconurus het&rol&pis is the same as Goode and
Bean's Hyuinnocephalus cavernosas : and I believe that both these names will prove
to be synonyms of Giglioli's Hymenocephalus italicus from the Mediterranean.
Subgenus Mvlacocephalus, Gthr.
Mouth wide, in the ordinary lateral position, the infra-orbital ridge quite
indistinct. Teeth in the upper jaw in two rows ; in a single row in the lower
jaw. Scales imbricate, spinigerous. Dorsal spine smooth.
119
97. Macrurus (Malacocephalus) Icevis, Lowe.
Macrurus Isevis, Lowe, Proc. Zool. Soc. 1843, p. 92.
Malacocephalus Uevii, Gunther, Cat. Pishes, Vol. IV. p. 397: Liitken, Vid. Meddel. Nat. Foren., Kjobenhavn
1872, p. 1.
Macrurus Isevis, Gunther, Challenger Deep-Sea Fishes, p. 148, pi. xxxix. fig. B : Smitt, Hist. Scandinavian
Fishes by Fries, Ekstrom & Snndevall, II. p. 593, fig. 141 : Alcock, Ann. Mag. Nat. Hist., Nov. 1889, p. 398 ; and
Aug 1891, p. 123: Goode and Beau, Oceanic Ichthyology, p. 415 : Koehler, Result. Sci. " Caudan," Poissons, Fasc.
III. p. 492.
Malacocephalus Izevis, Gilbert & Cramer, Proc. V. S. Nat. Mns. XIX. 1896, p. 432.
B. 7. D.r + 11-12. A. ciro. 200. P. 17. V. 9.
Head nearer a sixth than a fifth the total length : greatest body-height
equal to the length of the head behind the anterior nostril : tail peculiarly low,
long and tapering.
Snout blunt-pointed ; the median tubercle hardly, and the lateral tubercles
not at all, distinguishable ; slightly projecting beyond the mouth ; nearly as long
as the eye. Nostrils, as usual, in a scaleless fossa high up in front of the eye.
Major diameter of the eye a third to three-sevenths the length of the head,
barely equal to the width of the flat interorbital space.
Mouth wide ; the upper jaw, which is half as long as the head, reaches
behind the middle of the eye. Teeth of the upper jaw in two rows — an inner
row smaller and closer-set, and an outer row more distant and much larger. A
single row of large rather irregular teeth in the lower jaw.
Barbel about two-thirds as long as the eye.
Scales on all parts, except the mouth throat and branchiostegal membranes
and certain definite patches to be presently noticed near the ventral fins. Those
of the head are rough and very small : those on the body are small and are
covered with very short bristle-like spinelets.
Eleven or 12 rows of scales between the last ray of the first dorsal fin and
the lateral line.
First dorsal spine rudimentary ; the second a little longer than the post-
orbital portion of the head.
Pectorals rather narrow, pointed, about the same length as the second dorsal
spine.
V entrals very short ; the outer ray very slightly prolonged — about as long
as the eye.
The vent lies at the end of an oval naked depression between the ventral
fins : there is a second naked fossa, but with its long diameter transverse, just
in front of, or between, the bases of the ventral fins.
There are 60 or more slender pyloric casca.
120
Andaman Sea, 188 to 220, 265, and 370 to 419 fathoms : Arabian Sea, be-
tween the Maldives and Cape Comorin, 719 fathoms.
Eegd. Nos. 13116, 13517, 3-f9, ™.
Distribution. : off the coast of Brazil ; North Sea, Bay of Biscay, Madeira
and North Atlantic ; Mediterranean ( ? ) ; Arabian Sea, Andaman Sea, Hawaii
(Sandwich Is.).
I have compared our specimens with the figures and descriptions given by
Gunther and by the authors of the History of Scandinavian Fishes, and I am
perfectly sure that they are one and the same species.
Bathygadus, Gunther.
Bathygadus, Gunther, Ann. Mag. Nat. Hist., II. 1878, p. 23 : Challenger Deep-Sea Fishes, p. 154 : Goode and
Bean, Oceanic Ichthyology, p. 420 : Jordan and Evermann, Fishes N. America, III. p. 2563.
Head short and thick with the muciferous cavities well developed : tail long
and tapering.
Snout not projecting beyond the mouth, which is large and lateral, with the
jaws even in front. Vilhform teeth in narrow bands in the jaws only. Barbel
present or absent.
Gill-openings wide, the gill-membranes slightly united in front. No mem-
branous connexion or diaphragm between the first branchial arch and the wall of
the gill-chamber. Numerous long setiform gill-rakers on the outer side of the
first branchial arch.
Scales small, deciduous, cycloid.
Second dorsal fin almost continuous with the first, its rays are well
developed. Anal rays feeble. Ventrals arising below the pectorals.
Key to the Indian species of Bathygadus.
I. Barbel present : anterior dorsal, upper pectoral and outer ventral rays
very greatly prolonged ... ... ... ... B. longifilis.
II. No barbel : anterior dorsal, upper pectoral and outer ventral rays
sliglitly prolonged ... ... ... ... ... B. furvescens.
98. Bathygadus longifilis, Goode & Bean.
Bathygadus longifilis, Goode and Bean, Proc. U. S. Nat. Mus. VIII. 1885, p. 599, and Oceanic Ichthyology,
1896, p. 422: Gunther, Challenger Deep-Sea Fishes, p. 157: Alcock, Ann. Mag. Nat. Hist. (6) VI. 1890, p 302, and
VIII. 1891, p. 123.
Bathygadus multifilis, Gunther, Challenger Deep-Sea Fishes, p. 155.
Hijmenoccphalus longifilis, Vaillant, Exp. Sci. Travaillenr et Talisman, Poiss. p. 218, pi. xxiii. fig 1.
B. 7. D. r+ 10 + are. 140. P. 13-15. V. 8.
Head and body compressed. Head about a sixth the total length : greatest
height of the body about equal to the length of the head behind the anterior nostril.
121
Snout barely longer than the eye, which is nearly a fourth the length of the
head and slightly more than the -width of the interorbital space. Nostrils in a
scaleless fossa in front of the eye ; the posterior the larger.
Mouth wide, the maxilla, which is decidedly more than half as long as the
head, reaching the level of the posterior border of the orbit : villiform teeth in
bands of moderate breadth in the jaws.
Barbel slender, from half to nearly two-thirds the length of the head.
Operculum with two feeble points, which are often subcutaneous and in-
distinguishable : angle of preoperculum full and rounded. Very numerous long
close-set gill-rakers on the outer side of the first branchial arch.
Deciduous cycloid scales on the head and body : six rows of them between
the last ray of the first dorsal fin and the lateral line.
First dorsal ray rudimentary, the second is produced to a filament and is
about twice as long as the head, or even longer. The longest rays of the 2nd
dorsal fin are about as long as the snout and eye combined.
Upper pectoral ray nearly three times, 'outer ventral ray nearly twice as
long as the head.
About 20 long rather slender pyloric cseca.
The largest specimen is a little over a foot long.
Bay of Bengal, off west coast of Andamans, 683 fathoms : Arabian Sea, in
the neighbourhood of the Laccadives and Maldives, 459, 636, and 740 fathoms.
Eegd. Nos. 12860, 12861, 13132, 14004, ^ ^.
Distribution : Atlantic coasts of N. America ; Atlantic coasts of Morocco ;
Arabian Sea and Bay of Bengal ; off Philippine Islands.
99. Bathygadus furvescens, Alcock.
Bathygadus furvescens, Alcock, Journ. As. Soc. Bengal, Vol. LXIII. pt. 2, 1894, p. 128 : Illustrations of the
Zoology of the Investigator, Fishes, pl. XVI. fig. 1 (first dorsal, upper pectoral and outer ventral rays broken).
B. 7. D. r + 9-10. P. 15. V. 8.
Differs from B. longifilis, Goode and Bean, only in the following particulars : —
Length of head from a fifth to a sixth (young) the total.
Snout distinctly though not very greatly longer than the eye.
Major diameter of the eye a fourth (young) to a fifth (adult) the length of
the head.
No barbel.
Seven rows of scales between the last ray of the first dorsal fin and the
lateral hue.
16
122
Second dorsal spine from a half to two-thirds the length of the head. The
upper pectoral rays vary from about two-thirds (adults) the length of the head
to the entire length of the head (young). The outer ventral ray is about equal
to the post-orbital portion of the head in the adult, but is equal to the post-
rostral portion of the head in the young.
There are 20 very large pyloric casca.
The longest specimen is 20- inches in length.
Arabian Sea, 406, 480, 719 fathoms ; Gulf of Manar, 142 to 400 fathoms ;
Bay of Bengal, 410 fathoms ; Andaman Sea 405 fathoms.
Regd.Nos. 13213, 13470, 13550, ^_3-3, !£ ^5 ^-9.
This species may possibly be identical with B. melanobranchus Vaillant.
Family Ateleopodid.ee.
Head rather large, with the muciferous cavities fairly well developed ;
trunk short ; tail long, compressed, gradually tapering to a filament.
A single, short, dorsal fin, situated above the pectorals. A long anal
fin, continuous with the caudal.
Ventrals jugular.
No pseudobranchiae : no air-bladder : no pyloric appendages.
This is another family which has been found to range into Indian waters
since the publication of the Fishes and Fauna of India.
Ateleopus, Schleg.
Ateleopus, Temminck and Schlegel, Faun. Japon., Poiss, p. 255 : Giinther, Cat. Fishes IV., p. 398.
Skeleton semi-cartilaginous.
The snout projects well beyond the mouth, which is small, quite inferior
and strongly protractile downwards. Teeth villiform, minute ; in a band in the
upper jaw only, or in both jaws : palate smooth. No barbel.
Gill-openings fairly wide, the gill-membranes slightly united anteriorly.
The first branchial arch is rather broadly connected, at its upper part, by mem-
brane, with the wall of the gill-chamber, so that the slit between that arch and
the gill-cover is narrower than the other branchial clefts. Gill-rakers on the
outer side of the first branchial arch cartilaginous, tuberculiform. Eight bran-
chiostegals.
Head and body covered with a thick, gelatinous, scaleless skin.
The ventrals arise immediately behind the clavicular symphysis : each con-
sists of a single stout filament made up of two intimately coherent rays.
Distribution : Japan, Andaman Sea, Laccadive Sea.
123
100. Ateleopus indicus, Alcock.
Ateleopus indicus, Alcock, Ann. Mag. Nat. Hist., Ang. 1891, p. 123: Illustrations of the Zoology of the
Investigator, Fishes, pl. II. fig. 2.
B. 8. D. 8. A. + C. 76-80. P. 12. V. 2 (fused to form a single ray).
Length of the head about equal to that of the rest of the trunk, and con-
tained about 5j times m the total : greatest height of the body, at the shoulder,
equal to the length of the head behind the middle of the snout.
The broadly-pointed, depressed, projecting, marginally-inflated snout is one-
third of the head in length and twice the major diameter of the oval eye ; about
half its extent is preoral. The nostrils, which are very large, are situated
superiorly immediately in front of the eye.
The mouth is a small, quite inferior, crescentic orifice, in width hardly more
than equal to the diameter of the eye-ball, its angle barely reaching the vertical
through the anterior border of the orbit, though the maxilla reaches nearly to the
vertical through the middle of the orbit ; it is strongly protractile downwards,
and looks as if adapted for suction. There is a short band of very minute teeth
in the inner aspect of the upper jaw; but the lower jaw is quite toothless.
Gill-rakers short, coarse, cartilaginous.
Head, body, and fins uniformly invested with a soft, thick, gelatinous, scale-
less skin. The lateral line follows the dorsal curve of the trunk, and then runs
along the middle of the tail.
The base of the dorsal fin is about three-quarters as long as the snout : the
height of the fin is about equal to the length of the head. The longest rays of
the anal fin are about equal to the postorbital portion of the head.
Pectorals pointed ; their length is almost equal to that of the dorsal fin.
Each ventral consists of a stiffish cartilaginous rod, about half as long as the
head, and made up of two intimately coherent rays.
Stomach long, simple ; intestine short and wide, opening in front of the
first anal ray. The ovaries in the adult female consist of a pair of thin-walled
sacks loosely filled with largish eggs (over 1 millim. in diameter in spirit) and
opening by a common orifice behind the vent.
Colours : mottled dark brown to purple-black ; all the fins except the
ventrals black.
The largest specimen, an adult female, is 15 inches long.
Andaman Sea, 188 to 220 and 405 fathoms : Arabian Sea, off Travancore
coast, 224 to 284 fathoms.
T) J TV.T 1 onaa lil 456-458 4fi0
Regd. jNos. 13069, — , — j — , — •
124
Anacanthini Pleuronectoidei.
To the 8 genera and 39 species recorded in the Fauna of British India the
" Investigator " has added 8 more genera and 2-i more species. Not all of these,
however, belong to the fauna of the deep-sea.
The following is a synopsis of the genera of Pleuronectidte at present known to inhabit
Indian seas. Those that are represented in the depths are marked with an asterisk : those
that have recently been added to the Indian fauna by the " Investigator," are marked with a
dagger.
I. Jaws and dentition nearly equally developed on both sides : —
1. The dorsal fin begins on the neck ... ... ... Psettodes.
2. The dorsal fin begins in front of the eyes, on the snout : —
i. Eyes on the left side : lateral line with a curve above
the pectoral fin : —
a. Eyes not widely separated : teeth minute, equal,
in a single series : maxilla about a third the length
of the head
I. Eyes not widely separated : teeth unequal, in a
single series : maxilla nearly half as long as the
head
c. Eyes not widely separated : teeth large, unequal,
in a single series : maxilla about three-quarters
the length of the head
d. Eyes separated by a concave space, which is usu-
ally of considerable width
e. Eyes as in Rhomboidichthys : no scales
Eyes on the left side : lateral line nearly straight
Eyes on the right side: lateral line nearly straight,
slightly ascending anteriorly : mouth small
Eyes on the right side : lateral line with a wide curve
anteriorly: mouth large
i
II. Jaws and dentition much more developed on the blind side : —
n.
iii.
IV.
ARNOGLOSSUS.f
PSEUDORHOMBUS.
*CHASCAKOPSETIA.t
Rhomboidichthys.
PsETTYLLIS-f
ClTHARICHTHTS.
SAMARIS.f
BKACHTPLEORA.t
Caudal fin independent : —
i. Eyes on the left side : lateral line with a curve above
the pectoral: pectorals well developed
ii. Eyes on the right side : lateral line with a curve above
the pectoral : pectorals well developed
iii. Eyes on the right side : lateral line straight : pectorals
absent or present
Vertical fins confluent with the caudal : —
Eyes on the right side : pectorals present or absent
Eyes on the left side : pectorals absent : —
a. No lateral line
b. Two or three lateral lines on the left side
with tentacles ...
c. Two or three lateral lines on the left side
without tentacles
i.
ii.
*L.EOPS.t
*BoOPSETTA.f
Solea and *Achirus.
Stnaptura.
*Al>HORISTIA.t
ips
ips
Pl.AGUSIA.
*C\'NOGLOSSUS.
125
Family Pleuronectidm.
Chascanopsetta, Alcock.
Chascanopsetta, Alcock, Journ. As. Soc. Bengal, Vol. LXIII. pt. 2, 1891, p. 128.
Pelecanichthps, Gilbert and Cramer, Proc. U. S. Nat. Mns. XIX. 1896, p. 432.
Mouth very wide, the maxillary being very much more than half the length
of the head. Jaws and teeth equally developed on both sides, each jaw being
armed with a single row of long slender depressible teeth. Byes on the left side.
The dorsal fin commences near the tip of the snout, its rays, and those of the
anal, being simple, slender, and scaleless. Caudal free. Scales minute, mem-
branous, hardly imbricate. Lateral line with a strong curve above the pectoral.
Gill-openings wide, the gill-membranes united in front. Gill-rakers none.
Seven branchiostegals. Pseudobranchise large.
101. Chascanopsetta lugubris, Alcock.
Chascanopsetta lugubris, Alcock, Journ. As. Soc. Bengal, Vol. LXIII. pt. 2, 1894, p. 129, pi. vi. fig. 4 : Illus-
trations of the Zoology of the Investigator, Fishes, pl. XV. fig. 3.
B. 7. D. 115. A. 80. C. 16. V. 6.
Body long, low, tapering, the dorsal profile considerably more convex than
the ventral. The greatest height of the body is about one-fourth, and the length
of the head about one-fifth of the total, caudal included.
Mouth-cleft very wide, oblique, with the lower jaw strongly projecting : the
maxilla, which is hardly expanded posteriorly, is about three-quarters the length
of the head, — reaching nearly to the angle of the j:>roperculum. Each jaw is
armed with a single row of sharjD curved teeth of two sizes, the larger fairly
regularly alternating with the smaller : those of the lower jaw are very close-set,
and are strongly depressible inwards across the floor of the mouth : those of the
upper jaw are more distant, not so strongly depressible, and rather smaller.
Tongue large, free, with a long styliform point. The rami of the lower jaw are
capable of very wide divarication.
The eyes, which are on the left side, are large (their major diameter being
about two-sevenths of the length of the head', close-set (less than a- third of a
diameter apart), and nearly equal in front. The snout proper is short — about
two-thirds the length of the eye. The nostrils are small pores situated in front
of the interorbital space.
The gill-openings are wide, the gill-membranes being free posteriorly : the
gill-arches are extremely weak and slender, the gill-lamina? are delicate, and
there are no gill-rakers.
The body and the post-orbital portion of the head are covered with minute
membranous hardly imbricating scales, which are somewhat enlarged along the
126
lateral line. The lateral line on both sides has a strong sinuous curve above the
pectoral fin.
The fin-rays are weak and filiform : the dorsal begins in front of the eye,
on the snout. The caudal peduncle is strongly constricted, and expands again
at the insertion of the fin, which is long and pointed, — 6r> in the total length.
The pectorals are slender : that on the coloured side is much larger than its
fellow, its upper rays being nearly as long as the caudal. Both ventrals are well
developed.
Colours : dull dusky brown, the peritoneum showing through as a black
patch ; iris and fins black ; tongue dusky brown.
The largest specimen is 8- inches long.
Bay of Bengal, off Madras coast, 145 to 250 fathoms : Gulf of Manar, 143
fathoms.
Regd. Nos. 13728, 13729 : 5-^3.
Boopsetta, Alcock.
Boopsetta, Alcock, Jonrn. As. Soc. Bengal, Vol. LXV. pt. 2, 1896, p. 305.
Cleft of the mouth very narrow : teeth in the jaws only, in broadish villi-
form bands on the blind side, gradually becoming obsolescent on the coloured
side. Eyes on the right side, very large, almost in contact, the upper bulging
beyond the dorsal profile, the lower in advance of the upper. The dorsal fin
begins above the middle of the upper eye. Both pectorals and both ventrals
well developed. Scales of moderate size, stout ctenoid* and adherent on the
coloured side, thin cycloid and deciduous on the blind side. Lateral line with a
strong curve above the pectoral.
Gill-openings somewhat contracted, the membranes very broadly united
below the isthmus : gill-rakers numerous, of good length, pointed. Six branchi-
ostegals.
Closely allied to Pleuronectes.
102. Boopsetta pr&longa, Alcock.
Pacilopsetta prcelonga, Alcock, Journ. As. Soc. Bengal, Vol. LXIII. pt. 2, 1S94, p. 130, pi. vii. fig. 2: Illustra-
tions of thk. Zoology of the Investigator, Kishes, pl. XV. fig. 2 (youny).
Boopsett'i umbrarum, Alcock, Jonrn. As. Hue. Bengal, Vol. LXV. pt. 2, 1896, p. 305: Illustrations of thb
Zoology of the Investigator, Fishes, pl. XVII. fig. 5.
B. G. D. 60. A. 51. C. 18. P. d. 8, s. 6. V. 6. L. lat. circ. 110.
Height of the body one-third, length of the head one-fourth the total, with-
out the caudal.
* In the young the scales on both sides are cycloid.
127
The mouth is short and broad, the cleft approaching the vertical, the maxilla
being a little over three-fourths the length of the eye, which is slightly more
than one-third the length of the head. The teeth are in broad villiform bands
in the jaws on the blind side, the bands gradually becoming narrow and dis-
appearing on the coloured side. The upper eye bulges very strongly beyond the
general dorsal profile. The length of the snout — i.e., of the space between the
front wall of the lower orbit and the tip of the knob of the mandibular
symphysis — is less than half the length of the eye.
The body, and the head excepting only the snout throat and gill-membranes,
are covered with scales.
The rays of the vertical fins are stout, the longest are more than two-fifths
the greatest body-height. The caudal is large, with a distinct though broad
peduncle. The dorsal begins just behind the middle of the eye.
The coloured pectoral is rather longer than its fellow, the latter being more
than half the length of the head. The coloured ventral is rather longer than its
fellow, the latter being as long as the eye.
Colours in spirit : right side blackish-brown, with traces of six opalescent
cross-bands : irides and coloured pectoral fin blue-black, the pectoral with a
narrow white cross-stripe. Vertical fins (on coloured side), and right ventral,
almost black, tipped with milk white. Left side rather dusky.
In the young, both sides are coloured, being grey with numerous large black
blotches, disposed in series.
The largest specimen is 67 inches long.
Gulf of Manar, 142 to 403 and 180 to 217 fathoms : Andaman Sea, 185
fathoms.
Regd. Nos. 13733, 13734, 2, 3™^1, 3-^, *±±?.
103. Boopsetta maculosa, Alcock.
Pcecilopsetta maculosa, Alcock, Jonrn. As. Soc. Bengal, Vol. LXII1. pt. 2, 1894, p. 130, pi. vii. fig. 1. Illus-
trations op the Zoology of the Investigator, Fishes, pl. XV. pig. 1.
B. 6. D. 56. A. 46. C. 18. V. 5.
Differs from B. prsdonga only in the following particulars, specimens of
about the same size being compared : —
The height of the body is half the total length without the caudal.
The maxilla is almost as long as the eye, which is a third as long as the
head.
The dorsal fin begins above the middle of the eye : and the longest rays
are barely a fourth the greatest height of the body.
128
The pectorals are only about as long as the eye.
The largest specimen is nearly 4- inches long, and is not adult.
Colours of the young : both sides of the body grey with numerous large dark
grey and black blotches disposed in series : caudal with two black blotches :
right pectoral with a black blotch.
Judging from the analogy of B. prselonga the coloration of the adult would
be different.
Bay of Bengal, off Madras coast, 145-250 fathoms : Andaman Sea, 185
fathoms.
Regd. Nos. 13732, 555.
L^ops, Giinther.
L&ops, Giinther, Challenger Shore Fishes, p. 29.
Scianectes, Alcock, Ann. Mag. Nat. Hist., Sept. 1890, p. 216.
Body elongate-oval or piriform : head small. Cleft of the mouth very
narrow, with the dentition much more developed on the blind side than on the
coloured : palatine and vomerine teeth none. Eyes on the left side. The dorsal
fin begins in front of the eye. Scales small, thin, deciduous. Lateral line with a
curve above the pectoral.
Gill-openings moderately broad, the membranes united anteriorly and rather
broadly. Gill-rakers sufficiently numerous, short, pointed. Six branchiostegals.
This genus is represented in India by 3 species, only one of which is an
inhabitant of the deep-sea. Of the other two, one is hardly distinguishable from
L&ops pa/rviceps, Giinther.
104. Lceops macrophthalmus, Alcock.
Scianectes macrophthalmus, Alcock, Journ. As. Soc. Bengal, Vol. LVIII. pt. 2, 1889, p. 292, pi. xvi. fig. 4 : Ann.
Mag. Nat. Hist., Sept. 1890, p. 216 : Illustrations of the Zoology of the Investigator, Fishes, pl. XXIII.
FIG, 1.
B. 6. D. 85-88. A. 68. C. 17. P. s. 14. d. 12. V. 6. L. lat. circ, 95.
Body piriform, very delicate, its greatest height about 2l- in the total with-
out caudal. The length of the head is one-third, or rather less, of the same
standard. Snout obtuse, about half as long as the eye. Eyes close together,
separated by a salient decliving ridge, the lower slightly in advance ; their major
diameter rather more than one-fourth the length of the head. Cleft of the
mouth nearly vertical ; length of the maxilla a little more than one-fourth that
of the head ; the lower jaw projecting in repose. Villiform teeth in a band on
the blind side of each jaw.
129
Opercles thin ; the branchiostegal rays prolonged.
Scales minute, thin, smooth, deciduous. Lateral line salient, curved above
the pectoral, continued right along the caudal fin.
The longest dorsal rays are about half the length of the head and are
slightly shorter than the corresponding anal rays.
The coloured (left) pectoral, which is more developed than its fellow, is
nearly two-thirds the length of the head.
The coloured ventral is in the same line as the anal ; its longest rays are
equal to the major diameter of the eye.
Colours : left side dark sepia ; vertical fins and left ventral black ; left
pectoral grey and black ; edge of branchiostegal membrane, on left side, black.
The largest specimen is nearly 5 inches long.
Bay of Bengal, 100, 98 to 102, and 107 fathoms : Arabian Sea, off Malabar
coast, 100 fathoms.
Regd. Nos. 11721, 12805-12809, 13632, 13633, ^—.
After renewed examination I am convinced that the vomerine teeth, the
supposed presence of which led me to keep this species distinct from Lasops, are
not true teeth, but are only irregularities of the surface of the bone.
Solea, Cuv., Gthr.
105. Solea umbratilis, Alcock.
Solea (Achirus) umbratilis, Alcock, Journ. As. Soc. Bengal, Vol. LXIII. pt. 2, 1894, p. 131, pi. vii. fig. 3 :
Illustrations of the Zoology of the Investigator, Fishes, pl. XV. fig. 4.
D. 70. A. 50. C. 18. P. 0. V. 5. L. lat. circ. 80.*
The height of the body is 2g in the total without the caudal. The length of
the head is sometimes a little more, sometimes a little less than a third of the
total without the caudal (in adults). The snout is but slightly hooked : its length
is twice that of the eye and from two-sevenths to a quarter that of the head.
The eyes are nearly or quite a diameter apart. The nostril of both sides is a
slender tube. The mouth-cleft reaches to the posterior limit of the lower eye.
No pectoral fins whatever.
Ventral fins symmetrical, separated from the anal by more than an eye-length.
Colours in spirit : warm olive brown, with numerous large black blotches
which form four or five irregular transverse series and three irregular longitu-
dinal series : dorsal and anal fins bluish-black or black : underside smoky in
rather more than the posterior half. The black blotches are sometimes circum-
scribed by a light areola.
* From its commencement behind tbe npper eye.
17
130
Length (of a nearly adult female) 4- inches.
Bay of Bengal, off Madras coast 107 and 91 fathoms : Arabian Sea, off
Kattiwar coast, 82 fathoms, and off Malabar coast, 100 and 68 to 148 fathoms.
Regd. Nos. 13615-13618, 13621-13628, %**, ^~.
In form and colour this species at first sight resembles Solea cyanea (Ann.
Mag. Nat. Hist., Dec. 1890, p. 439), from which it is distinguished — specimens
of equal size being compared — by the much larger head and mouth, by the
larger and more widely separated eyes, and by the fewer rays in the dorsal and
anal fins. It is however closely allied to Solea cyanea, and also to Solea
melanosticta, Peters (MB. Ak. Berl., 1876, p. 845), and Solea kaiana, Gthr.
(' Challenger ' Shore fishes, p. 49, pi. XXI., fig. 0.) — all being comparatively
deep-water forms of the East Indian Seas.
Aphoristia, Kaup.
AphoriUia, Kanp, ArclnV. fur Naturges. XXIV. 1858, i. p. 106 : Giinther, Cat. Fishes, IV. p. 490 : Goode and
Bean, Oceanic Ichthyology, p. 458.
Symphurus, Eafinesque, Jordan and Evermann, Fishes N. Amer. III., p. 2704.
Body elongate, lanceolate. Mouth narrow, more developed on the blind
(right) side, where alone there are minute teeth in the jaws. Snout hardly
hooked. Eyes on the left side. The dorsal fin begins above the middle of the
eye : it and the anal are confluent with the caudal. No pectoral fins. Only one
ventral — the left — present. Scales of moderate size, ctenoid. No lateral line.
Gill-openings narrow, the gill-membranes rather broadly united. Gill-
rakers minute.
Distribution : "West Indies and American coasts of North Atlantic ; Bay of
Bengal, Andaman Sea, and Gulf of Mandr.
Key to the Indian species of Aphoristia.
I. Height of the body rather more than a fourth of the total length :
no cross-stripes : some coloration on the right side : —
1. Eyes so far forward that the mouth-cleft reaches to at least
the middle of the lower one : the origin of the anal is
hardly an eye-length behind the base of the ventral ... A. Gilesii.
2. The mouth-cleft does not reach to the middle of the lower
eye : the origin of the anal is at least two eye-lengths be-
hind the base of the ventral ... ... ... A. Wood-Masoni.
II. Height of the body a fourth, or less than a fourth of the total
length : dark cross-bands on the left side, no coloration on the right
side : —
1. Length of the head a fourth of the total : 3 (sometimes 4)
cross-bands... ... ... ... ... A. trifasciata.
2. Length of the head a fifth of the total : 7 (sometimes 6)
cross-bands... ... ... ... ... A. septemstriata.
131
106. Aphoristia Gilesii, Alcock.
Aphoristia Oilesii, Alcock, Jonrn. As. Soc. Bengal, Vol. LVIII. pt. 2, 1889, p. 293, pi. xvii. fig. 2 : Illustra-
tions of the Zoology of the Investigator, Fishes, pl. XIV. fig. 4.
D. 97-98. A. 83-85. C. 14. V. 4. Scales about 90 transverse rows
behind head : about 38 longitudinal series behind gill-opening.
1 2
Length of the head a fifth, greatest body height 3- to 3^ in the total length
without the caudal.
Snout between a fifth and a sixth the length of the head : the posterior
nostril is a pore immediately in front of the inter-orbital space, the anterior is
a slender tube situated above the lip midway between the eye and the tip of the
snout. No nostrils on the blind side.
Eyes well within the anterior third of the head ; their major diameter is
about two-thirds the length of the snout and hardly an eighth the length of the
head : they lie not very far apart, between the same verticals.
Mouth-cleft nearly horizontal, about two-ninths the length of the head,
reaching to or slightly beyond the middle of the eye.
Body and every part of the head except the edges of the mouth, covered
with scales, which are sharply ctenoid on the coloured side and less strongly
ctenoid on the blind side.
The longest rays of the dorsal fin are a little more than a third the greatest
body-height in length.
The anal fin begins about an eye-length behind the base of the ventral,
which arises between the after border of the gill-covers.
Colours : left side dark brown with numerous close fine parallel longitudinal
lines : right side grey with irregular dark brown patches.
The largest specimen is over 5- inches long.
Bay of Bengal, 193 and 210 fathoms.
Regd. Nos. 11684, 13630.
Named after Major G. M. J. Giles, I. M. S., who was Surgeon-Naturalist to
the Marine Survey from 1884 to 1888.
107. Aphoristia Wood-Masoni, Alcock.
Aphoristia Wood-Masoni, Alcock, Journ. As. Soc. Bengal, Vol. LVIII. pt. 2, 1889, p. 294 : Illustrations of
the Zoology of the Investigator, Fishes, pl. XVI. fig. 4.
D. 90-98. A. 78-84. C. 14. V. 4. Scales about 90 transverse rows
behind head : about 34 longitudinal series behind gill-opening.
Except in the following particulars this species agrees with A. Gilesii : —
The length of the snout is between a fourth and a fifth that of the head.
132
The eyes are only just within the anterior third of the head, and their
diameter is not much more than half the length of the snout.
The mouth-cleft is a fourth the length of the head and does not reach to the
middle of the eye.
The longest dorsal rays are not a third the greatest body height.
The anal fin begins a snout length behind the base of the ventral.
Colours : left side as in the preceding species, but the ground-colour is
lighter : on the right side the fine parallel lines may be as distinct as they are on
the left.
The largest specimen is nearly 55 inches long.
Andaman Sea, 490 fathoms : Bay of Bengal, 475 fathoms.
Eegd. Nos. 11765, 13180.
Named after my predecessor James Wood-Mason. He was deputed as
Naturalist with the " Investigator " in April and May 1888.
108. Aphoristia septem-striata, Alcock.
Jphoristia septem-striata, Alcock, Ann. Mag. Nat. Hist., Aug. 1891, p. 125: Illustrations of the Zoology op
the Investigator, Fishes, pl. II. fig. 1.
D. 97. A. 80. C. 12. V. 4. Scales from 92 to 100 transverse rows
behind the head : about 40 longitudinal series behind the gill-opening.
Except in the following particulars this species agrees with A. Gilesii: —
The greatest height of the body is a fourth of the total length.
The length of the snout is between a fourth and a fifth that of the head.
The eyes are only just within the anterior third of the head and are almost
in contact.
The mouth-cleft is about a fourth the length of the head and does not reach
to the middle of the eye.
The longest dorsal rays are about a third the greatest body-height.
The anal fin begins rather more than a snout-length behind the base of the
ventral.
Colours : left side warm brown, with six or seven broad dark cross-bands :
right side colourless.
The largest specimen is 4- inches long.
Andaman Sea, 188 to 220 fathoms : Gulf of Manar, 142 to 400 fathoms.
Regd. Nos. 13109, 13110, 13613, 13614.
133
109. Aphoristia trifasclata, Alcock.
Aphoristia trifasciata, Alcock, Jonrn. As. Soc. Bengal, Vol. LXIII. pt. 2, 189-4, p. 132, pi. vii. fig. 4. Illus-
TRATIONS OF THE ZOOLOGY OF THE INVESTIGATOR, FlSHES, PL. XV. FIG. 5.
D. 87-89. A. 75-77. C. 12. V. 4. Scales 80-82 transverse rows be-
hind the head : 38-40 longitudinal series behind the gill-opening.
Except in the following particulars this species agrees with A. Gilesii : —
The length of the head is a fourth of the total, without the caudal, as also
is the greatest height of the body. The eyes are in contact.
The mouth-cleft is about a fourth the length of the head and reaches con-
siderably behind the middle of the eye.
The longest dorsal rays are about two-fifths the greatest body-height.
The anal fin begins a snout-length behind the base of the ventral.
Colours : left side warm brown with 3 broad blackish cross-bands : right
side colourless.
The largest specimen is 4- inches long.
Bay of Bengal, off Madras coast, 145 to 250 and 195 to 210 fathoms.
Regd. Nos. 13595-13606, 13608-13612.
This species and the last are both akin to Symphurus leei Jordan and
Bollman, Symphwrus fasciolaris Gilbert, and Symphurus atramentatus Gilbert.
Ctnoglossus, Ham. Buch., Gthr.
110. Cynoglossiis Carpenteri, Alcock.
Cynoglossus' Carpenteri, Alcock, Journ. As. Soc. Bengal, Vol. LVIII. pt 2, 18S9, p. 287, pi. xviii. fig. 1 : Illus-
trations of the Zoology of the Investigator, FisnES, PL. XXII. fig. 5.
D. circ. 105. A. circ. 85. C. 12. V. 4. Scales about 95 rows from top
of gill-opening to caudal.
Body lanceolate : tail tapering. The height of the body is about the same
as the length of the head, or from 3^ to 3g in the total without the caudal.
The snout is obtusely triangular and about a third the length of the head :
the rostral hook ends in the vertical through the anterior border of the eye.
Two nostrils : the posterior, which is a pore lying behind a papilla, is situated in
front of the interorbital space ; the anterior, which is tubular, is situated a little
in advance of the lower eye.
Eyes prominent, about half a diameter apart : the lower one, which is
slightly the larger and slightly posterior, is about one-eleventh the length of the
head in its major diameter.
Mouth large ; its angle is nearly an eye-length behind the posterior border
of the lower orbit, and about equidistant between the tip of the snout and the
edge of the gill-cover. Minute teeth in the posterior two-thirds of the jaws on
the blind side only. Lips not fringed.
134
The opercle is conspicuously expanded below and behind, and the branchio-
steo-al rays and membrane extend a considerable distance behind its edge, giving
the appearance of a broad fringe.
The integument is invested with small strong scales, which on the blind side
and anterior half of the coloured side are cycloid, and on the posterior half of
the coloured side sharply ctenoid. Three lateral lines on the left side, the middle
separated from the upper by 16 or 17, and from the lower by 22 to 24 rows of
scales at the respective points of greatest divergence : none on the right. Those
on the left side are connected by an irregular cross line on the head.
The longest rays of the dorsal and anal fins are about equal to the snout.
The ventral fin is almost indistinguishably confluent with the anal.
Colours in spirit : warm brown, the opercles and caudal blackish : right side
colourless.
Length 6 to 8 inches.
Bay of Bengal, 68, 98 to 102, and 107 fathoms : Arabian Sea, off Malabar
coast, 100 fathoms.
Regd. Nos. 12433, 12434, 12726 a to g, 12727 a to h, 12728 a to h, 13589-
-i o'ni 647-659
13o94, j- .
Sub-order Physostomi.
Family Ster iiopty chidce.
Eight species of this family, belonging to six genera, are now known to
inhabit the seas of India. They probably all belong to the " Necton," though
they probably live below the depths to which daylight penetrates.
Body naked, or with very thin and deciduous scales. Barbels none.
Margin of the upper jaw formed by the maxilla and premaxilla, both of
which are toothed. Gill-opening very wide, the opercular apparatus not
always completely developed. Pseudobranchiae present or absent. Air-
bladder simple, if present. Adipose fin present, but generally small.
Series of phosphorescent bodies along the ventral aspect of the body.
Synopsis of the Indian Genera of the family Sternoptychidae.
' I. Body short and elevated, its height more than half its length without
the caudal. Pseudobranehioe present : —
1. Dorsal fin preceded by a large triangular transparent osseous
plate : no teeth on the vomer : —
i. Body hatchet shaped ... ... ... Augyuopelecis.
ii. The ventral constriction between the body and the tail is
filled by a curious transparent fold of skiu supported by
long interhsenial rays ... ... ••• Sfernoptyx.
2. Dorsal fin preceded by a forked spine : teeth on the vomer ... Polyipnos.
CO
135
'II. Body very low and elongate. No pseudobranchiee : —
1. Dorsal fin in the after half of the body, arising opposite the
origin of the anal : anal long ... ... ... Cyclothone.
2. Dorsal fin in the middle of the body, well in advance of the anal :
anal moderately long ... ... ... ... Photichthys.
3. Dorsal fin placed far forward, hardly a head-length behind the
occiput: anal short : no gill-rakers ... ... ... Chauuodus.
Argyropelecus, Cocco.
Argyropelecus, Cocco, Giorn. Sci. Sicil. 1829: Cuvier and Valenciennes, Hist. Nat. Poiss. XXII. p. 392 :
Giinther, Cat. Fishes, V. p. 389 : Goode and Bean, Oceanic Ichthyology, p. 125 : Jordan and Evermann, Fishes N.
Amer., p. 603.
Vleurothyris, Lowe, Fishes of Madeira, p. 64.
Head and body elevated and compressed : tail narrow, abruptly delimited
from the trunk, like the handle of a hatchet. Body covered with a thin silvery
skin, which is superficially wrinkled, as if thin scales had been loosely attached
to it. Series of " bull's-eye "-like luminous organs along the ventral border of
head, body, and tail. A series of imbricate scutes forms a serrature along the
middle line of the belly, from the throat to the ventral fins.
Cleft of the mouth wide, vertical, with the lower jaw prominent, and the
margin of the upper jaw formed by the premaxilla and maxilla : minute teeth in
a single row in the jaws and palatines. The jaw-bones, like all the bones of
the head, are very thin, but well-ossified.
Eyes rather large, lateral but directed upwards and very close together.
Gill-opening very wide, very long gill-rakers on the outer side of the first
branchial arch. Nine branchiostegals. Pseudobranchias present.
Dorsal fin short, situated about the middle of the back, preceded by a thin
transparent triangular osseous plate in which the ends of several neural spines
are visible. Anal short. Caudal forked. Adipose dorsal rudimentary.
Pectorals well developed, situated near the ventral profile : ventrals small :
both the humeral arch and the pubic bones project strongly, as flat spines, in the
middle line of the belly.
An air-bladder. Four pyloric appendages.
Distribution: Atlantic, Mediterranean, Bay of Bengal.
ill. Argyropelecus hemigymnus, Cocco.
Argyropelecus hemigymnus, Cocco, Giorn. Sci. Sicil. 1829 : Bonaparte, Fann. Ital. Pesc, text : Cut. & Val., Hist.
Nat. Poiss. XXII., p. 398: Giinther, Cat. Fishes, V., p. 385, and Challenger Deep-Sea Fishes, p. 167 : Vaillant, Exp.
Sci. Travaillenr et Talisman, Poiss., p. 103 : Goode and Bean, Oceanic Ichthyology, p. 126, fig. 147 : Jordan and
Evermann, Fishes N. Amer. p. 604.
Sternoptyx mediterranea, Cocco, Giorn. 11 Faro, 1838, IV., p. 7, fig. 2 : Bonaparte, Faun. Ital. Pesc, fig.
Sternoptysc hemigymnus, Cuvier, Uegne Animal, Poiss., pi. 103, fig. 3.
B. 9. D. 7-8. A. 11. P. 9. V. 5.
Greatest height of the body about 2- times the greatest height of the tail,
and about half the total length without the caudal fin. Major (vertical) diameter
136
of eye from half to two-fifths the length of the head. A small spine at the
posterior corner of the mandible, and another at the angle of the preoperculum.
Tail without spines.
The pectoral fin extends nearly to the origin of the anal.
Bay of Bengal, 1803 fathoms.
Distribution : Atlantic, Mediterranean, Bay of Bengal. Regd. No. 13119.
Besides the " Investigator " specimen there are several specimens from the
Mediterranean in the Indian Museum.
Sternoptyx, Hermann.
Sternoptyx, Hermann, Naturforscher, 1781, XVI : Cnvier & Valenciennes, Hist. Nat. Poiss. XXII., p. 412 :
Giinther, Cat. Fishes, V., p. 386, and Challenger Deep-Sea^Fishes, p. 168: Gootle and.Bean, Oceanic Ichthyology,
p. 123 : Jordan and Evermann, Fishes N. Amer., p. 603.
Form of the body as in Argyropelecus, but with a peculiar drag downwards
and forwards, and with the wide corner between the ventral angle of the trunk
and the tail filled up by a curious transparent fold of skin supported by long
interhaemal rays. Body covered with a skin like that of Argyropelecus, but not
quite so brilliant. Series of luminous organs as in Argyropelecus, but less
conspicuous and less " bull's-eye "-like. The middle line of the belly between
the humeral and pelvic spines is trenchant, not serrated.
Mouth as in Argyropelecus, but not quite so wide : palatines toothless.
Eyes large, lateral, without any upward cast.
Gill-opening wide : a few rather distant gill-rakers on the outer side of the
first branchial arch are elongate. Branchiostegals five. PseudobranchiaB present.
Dorsal fin short, situated farther back than in Argyropelecus, preceded by a
thin triangular osseous plate which has a serrated edge but no series of indepen-
dent spines. Anal long, extending from near the vent to near the caudal ; its
rays are feeble. Caudal forked. The anal is preceded by a pair of little spines
similar to those at the pectoral and pubic symphyses.
Pectorals well developed, not far from the ventral profile. Ventrals small.
Distribution : Atlantic, Arabian Sea, Western Pacific.
112. Sternoptyx diaphana, Hermann.
Sternoptyx diaphana, Hermann, see Giinther, Cat. Fishes, V., p. 387 aud Challenger Deep-Sea Fishes, p. 169,
pi. xlv , figs. D, D' : Vaillant, Exp. Sci. Travaillenr et Talisman, p. 102 : A. Agassiz, Hull. Mus. Comp. Zool. XV.,
p. 22, fig. 195 : Goode and Bean, Oceanic Ichthyology, p. 124, fig. 146 : Jordan and Evermann, Fishes, N. Amer.,
p. 603: Collett, Hirondelle Poissons (Monaco, 1896), p. 125.
B. 5. D. 9-12. A. 12. P. 10. V. 3.
Greatest height of the body twice that of the tail (not including the trans-
parent fold that fills up the corner between the tail and the belly), and more
137
than two-thirds the total length without the caudal. Diameter of the circular
eye nearly half the length of the head.
Besides the spines connected with the fins and the pectoral and pelvic
arches, there are a pair of spines on the occiput, a pair on the nape, one at the
angle of the preoperculum and one at the posterior end of the mandible.
The pectorals are short, being about half again as long as the eye. The
ventrals are very short and narrow.
Arabian Sea : off Malabar coast, 912 to 931 fathoms.
Distribution : Atlantic, Arabian Sea, "Western Pacific.
Regd. No. y.
Besides the " Investigator " specimen there is one of the " Challenger "
duplicates in the Indian Museum.
Polyipnus, Giinther.
Polyipnus, Giinther, Challenger Deep-Sea Fishes, p. 170 (1887) : Goods and Bean, Oceanic Ichthyology, p. 128.
Form of the body as in Argyropelecus but without the abrupt ventral con-
striction between the body and the tail. Body covered with a thin silvery
burnished skin, like that of Argyropelecus, and with large extremely thin exces-
sively deciduous scales. Series of conspicuous luminous organs along the ventral
border of the head, body, and tail, and also on the sides of the head and belly.
A series of scutes form a serrature along the mid-ventral line from the pectoral
symphysis to the base of the caudal, the series being broken by the ventral and
anal fins.
Mouth as in Sternoptyx, but the teeth in the jaws appear to be in more than
one series, at any rate anteriorly, and there are teeth on the vomer.
Eyes large, lateral.
Gill-opening wide, numerous very long gill-rakers on the outer side of the
first branchial arch. Pseudobranchia? present. Branchiostegals six (?)
Dorsal fin beginning about the middle of the body, rather short, preceded
by a small bifurcate spine but not by any large triangular osseous plate. Anal
rather short. Caudal forked. Adipose dorsal present.
Pectorals well developed, situated near the ventral profile : ventrals small :
the humeral and pelvic symphyses project in the middle line of the belly, but
not so strongly as in Argyropelecus.
An air-bladder.
Distribution. Tropical Pacific, Andaman Sea.
18
138
113. Polyipnus SpiflOSUS, Griinther.
Polyipnus spinosuss, Giinther, Challenger Deep-Sea Fishes, p. 170, pi. li. fig. B : Alcook, Ann. Mng. Nat. Hist.,
Dee. 1889, p. 398, and Aug. 1891, p. 126 : Gilbert and Cramer, Proc. U. S. Nat. Mus. XIX., 1896, p. 416.
D. 12-13. A. 15-16. P. 12. Y. 5.
Body oval, its greatest height between two-thirds and three-fourths its
length without the caudal. Major diameter of eye vertical, more than half the
length of the head. Snout very short. Nostrils large, situated close together
on top of the snout, the posterior the larger.
The upper part of the head is narrow compressed and concave, and is
bounded on each side by a serrated ridge that ends in a large sharp semi-recumb-
ent spine. The edge of the preoperculum is serrated near the angle, which is
occupied by a claw-like spine pointing vertically downwards. The lower edge
of the mandible is finely serrated and ends in a spinule.
The scutes along the mid-ventral line are spiny.
The dorsal fin is preceded by a small forked spine: the adipose dorsal
occupies the middle of the space between the dorsal and caudal. The pectoral
reaches almost as far as the base of the ventrals.
The luminous organs are as follows, on each side : —
(1) a series of six small ones in the intervals between the bases of the
branchiostegal rays :
(2) a series of six larger ones along the isthmus :
(3 1 a series of ten still larger ones along the abdomen between the humeral
symphysis and the base of the ventral fin :
(4) a series of five between the ventral and the anal :
(5) a series of twelve above the anal fin :
(6) a series of four or five along the ventral border of the caudal peduncle :
(7) a small one at the anterior angle and a small one at the posterior angle
of the eye :
(8) a very large one below the middle of the orbit, on the preoperculum :
((.t) a small one on the suboperculum :
(10) two above the base and three behind the base of the pectoral fin, these
form a second tier on the abdomen :
(11) one behind the gill-opening and one much further back, forming a
third tier on the abdomen.
The scales are extremely thin and deciduous : one from the side of the trunk
is 7'"> millim. in vertical and about 25 inillim. in horizontal diameter; one from
the middle of the tail is 6"5 millim. and not quite 2 inillim. in its diameters.
The largest specimen is between 2 and 2- inches long.
139
Cyclothone, Goode & Bean.
Cyclothone, Goode and Bean, Bull. Mns. Coinp. Zool. X., 1883, p. 221, and Oceanic Ichthyology, p. 99 : Jordan
and Evermann, Fishes N. Amer., p. 581.
SigmoTs, Gill, Proc U. S. Nat. Mas. VI., 1883. p. 256.
Neostomii, Vaillaut, Exp. Sci. Travaillenr et Talisman, Poiss., p. 96.
Body elongate, compressed, without scales. Series of inconspicuous lumin-
ous spots along the ventral aspect of the body from the chin to the caudal fin.
Head compressed, the bones thin but ossified. Cleft of mouth oblique, extremely
wide, the maxilla, which extends nearly to the angle of the preoperculum,
forming much the greater part of the margin of the upper jaw ; the lower jaw
projecting. Both jaws with a single series of needle-like teeth, large ones
usually alternating with small ones. Teeth on the palatine and pterygoid bones
usually present. Bye moderate, or small.
Gill-openings very wide : numerous long bristle-like gill-rakers. No
pseudobranchiaa. Branchiostegals numerous.
Dorsal fin short or of moderate length, situated in the after half of the body,
arising opposite the origin of the anal. Anal long. Caudal forked. Adipose
fin, when present, small. Pectorals and ventrals well developed.
A long slender air-bladder is present in one of the Indian species.
Distribution : Atlantic, Antarctic, Indo-Pacific.
114. Cyclothone elongata (Gthr.).
Gonostoma elongat am, Giinther, Ann. Mag. Nat. Hist. II. 1878, p. 187 ; and Challenger Deep-Sea Fishes, p. 173,
pi. slv., tig. B : Alcock, Ann. Mag. Nat. Hist. (6) VIII., 1891, p. 127, and X., 1892, p. 354.
Sigmops stigmaticus, Gill, Proc. 0. S. Nat. JIua. VI., 1883, p. 256.
Cyclothone elongata, Goode and Benu, Oceanic Ichthyology, p. 101, fig. 119: Jordan nnd Evermann, Fishes
N. Amer , p. 583.
B. 11. D. 13. A. 29-30. P. 11-12. V. 7-8.
Length of the head two-ninths, height of the body one-seventh of the total
length without the caudal.
Length of the eye about two-thirds that of the snout, between a seventh
and an eighth that of the head, less than the width of the interorbital space.
Nostrils small, situated on top of the snout ; the posterior the larger.
Mouth-cleft extremely wide, the maxilla reaching to the angle of the pre-
operculum. About 14 or 15 large needle-like teeth (two or three of which
belong to the premaxilla) with minute teeth between them, on each side of the
upper jaw ; about ten large teeth, with small teeth between them, on each side
of the lower jaw. Small teeth on the palatines and pterygoids, the anterior
palatine tooth somewhat enlarged.
Gill-laminae very short ; gill-rakers numerous, long and bristle-like. Bran-
chiostegals very short.
140
The dorsal fin commences nearly opposite the commencement of the anal,
and is higher than the body below it. The anal ends a short distance in front
of the caudal; its anterior portion (about 13 rays) is much the highest. Caudal
forked.
Pectorals narrow, arising near the ventral profile, two-thirds as long as the
head, and reaching two-thirds the way to the base of the ventrals. Ventrals
narrow, nearly two-thirds as long as the pectorals ; almost, but not quite, reach-
ing to the vent.
The body is covered with a black skin which is scaleless, and is coated in
life with thick tenacious mucus.
Two rows of luminous organs run along each side of the ventral border of
the body.
In the lower series are 4 in front of the pectoral, 10 between the pectoral
and ventral, 4 between the ventral and anal, and 22 above the anal fin.
The upper series, which are much the most conspicuous, consist of twelve
organs and extend from the pectoral to the vent.
In addition, there are small organs in each of the spaces between the bases
of the branchiostegal rays ; a large one below each eye, one near the upper end
of the preoperculum, one on each side of the mandibular symphysis and a short
series on both the dorsal and ventral edges of the caudal peduncle.
In life, the luminous organs are bright rose-pink, with silvery margins.
The vent is midway between the root of the caudal fin and the eye. An air-
bladder is present, and six large pyloric casca.
A fine male and a female with ripe ovaries from the Arabian Sea, in the
neighbourhood of the Laccadive Islands, 738 and 1200 fathoms. In both the
length is 7'- inches.
Distribution : New Guinea and Banda Sea : Arabian Sea : off Atlantic coasts
of North America.
The female, taken off Minnikoy, when the dredge was shot in 1200 fathoms,
was alive and active when taken from the dredge, and lived on board for about
quarter of an hour afterwards ; so that it is most improbable that it came from
any great depth. No display of luminosity was observed, though it was watched
for.
I have described the ovaries, and the microscopic structure of the stomach,
in Ann. Mag. Nat. Hist, for November 1892. The ovaries are long narrow
tubes in which the developing ova form a closely-pleated band with very little
interstitial stroma.
In the stomach the submucous coat is of remarkable thickness, and consists
of a network of connective tissue of great regularity, the meshes of which are
141
crammed with leucocytes : the microscopic appearance is very much like that of
the cortex of mammalian lymphatic gland.
Regd. Nos. 13122, 13181.
115. Cyclothoiie microdon (Gunther).
Gonostoma microdon, Gunther, Ann. Mag. Nat. Hist. II. 1878, p. 188, and Challenger Deep-Sea Fishes, p. 175 ■'
Alcock, Ann. Mag. Nat. Hist. Dec. 1889, p. 399 : Lutken, Vid. Selsk. Skr. (6) Nat. Math. Afd. VII. 1892, 6, p. 280,
pi. ii. figs. 4, 5.
Cyclothone lusca, Goode and Bean, Ball. Mus. Comp. Zool. X. 1883, p. 221 : A. AgasBiz, Ball. Mus. Comp. Zool.
XV. 1888, p. 22, fig. 196.
Cyclothone microdon, Goode and Bean, Oceanic Ichthyology, p. 99, fig. 114: Jordan and Evermann, Fishes N.
Amer. p. 582 : Collett, "Hirondelle" Poissons (Monaco, 1896) p. 130.
B. 9. D. 12-13. A. 17-21. P. 9-10 V. 5?
The chief differences, besides the much smaller size and much shorter anal
fin, that separate this species from C. elongatum, are the following : —
The eye is minute.
The teeth in the upper jaw are numerous, and gradually increase in size
from before backwards, one or two in the premaxilla and a few in the posterior
half of the maxilla being slightly enlarged : the teeth in the lower jaw, which are
also extremely numerous and close set, also gradually increase in size from before
backwards.
The narrow ventrals reach beyond the vent.
There are no glandular (luminous) masses on the edges of the caudal
peduncle.
Bay of Bengal, off Andaman Is. 485 fathoms : Andaman Sea, 265 fathoms.
Regd. Nos. 12455, 12468.
Besides the " Investigator" specimens there are several of the " Challenger "
duplicates in the Indian Museum.
Photichthys, Hutton.
Phosichthys, Hntton, Cat. Fishes New Zealand, p 55.
Photichthys, Gunther, Challenger Deep-Sea Fishes, p. 177 : Goode and Bean, Oceanic Ichthyology, p. 104.
Body elongate compressed, with thin extremely deciduous scales. Series
of conspicuous luminous organs extending along the ventral aspect of the body
from the chin to the caudal fin. Head compressed, the bones thin but ossified.
Cleft of mouth oblique, extremely wide, the maxilla, which extends nearly to the
angle of the preoperculum, forming the greater part of the margin of the upper
jaw ; the lower jaw projecting. Both jaws with a single series of teeth. A
fang on either side of the head of the vomer. Teeth on the palatines and
sometimes on the pterygoids. Eye moderate.
142
Gill-openings very wide : numerous long bristle-like gill-rakers. No pseudo-
branchiae. Branchiostegals numerous.
Dorsal fin short, in the middle of the body, standing above the interval
between the ventrals and the anal. Anal rather long. Caudal forked. Adipose
dorsal small. Pectorals and anals well developed.
A long slender air-bladder present.
Distribution. Seas of India and New Zealand.
Phofichthys differs from Cyclothone (1) in having scales, though they are
thin and deciduous, (2) in the very conspicuous luminous organs, (3) in the
position of the dorsal fin, which stands over the space between the ventrals and
the anal instead of commencing opposite the origin of the anal.
116. Phofichthys corythceolus, Alcock.
Diplophos corythxolum, Alcock, Ann. Mag. Nat. Hist. August, 1898, p. 147: Illustrations of the Zoologt of
the Investigator, Fishes, pl. XXV. fig. 3.
B. 12. D. circ. 11. A. circ. 24. P. 10. V. 7.
Length of head about one-fourth, height of body between one-fifth and one-
Bixth of the total without the caudal.
The snout, which has the lower jaw prominent, is hardly longer than the
eye, which is not quite a fourth the length of the head. The eyes are not quite
a diameter apart.
The maxillary almost reaches to the preopercular angle. There is a single
row of small, rather distant, acicular fangs of unequal size in either jaw, and a
row of close-set acicular teeth on part of the palatines ; the whole surface of the
mesopterygoids is studded with sharp little denticles ; and there is a fang on
either side of the head of the vomer.
Gill-openings extremely wide ; four gills with short laminae ; gill-rakers,
especially those on the first arch, long and bristle-like.
The body has evidently been covered with large thin and deciduous scales.
The dorsal fin arises about an eye-length behind the base of the ventrals,
and its last few rays are just above the first few anal rays ; its first ray is slightly
nearer to the snout than to the base of the caudal. The long anal fin approaches
within less than half a head-length of the base of the caudal. The pectorals
are on almost the same plane as the ventrals, and these arise about midwav
between the base of the former and the origin of the anal.
The luminous organs, which are of the " bull's eye " type, are disposed as
follows on either side : —
(1) one at the mandibular symphysis :
(2) one between the bases of all the branchiostegal rays :
143
(3) 12 or 13 between the tip of the isthmus and the base of the pectoral :
(4) eight between the pectoral and the ventral:
(5) eight between the ventral and the anal :
(6) 15 along the bases of the anal rays :
(7) eight between the anal and the caudal.
The above form the lowermost tier.
The upper tier consists of 17 between the gill-opening and the level of the
anal fin, where it stops.
In addition, there are some luminous organs on the head, notably one at the
anterior angle of the orbit.
Length 5 inches.
Andaman Sea, 185, 188 to 220, and 405 fathoms.
Regd. Nos. 13076, 13077, ^ x-f°, ^, ^.
Chauliodus, Bl. Schn.
Chauliodus, Schneider, Bioch, Syst. Ichth. p. 430: Gilnther, Cat. Fishes, V., p. 392: Goode and Bean, Oceanic
Ichthyology, p. 96: Jordan and Evermann, Fishes N. Amer., p. 585.
Body elongate, compressed, covered with exceedingly thin and deciduous
scales. Head elevated, compressed, with the bones thin but ossified, and the
gill-cover very narrow. Series of luminous spots along the ventral aspect of the
body from the chin to the caudal fin. Cleft of the mouth extremely wide, the
upper jaw, about half of the edge of which is formed by the premaxilla, reaching
almost to the angle of the preoperculum ; the mandible projecting. About four
enormous fangs in each premaxilla, and about five fangs, the anterior one of
which is more than half the length of the head, in the mandible : none of these
fano-s are received within the mouth. Edge of maxilla finely denticulated :
palatine with a single series of small teeth : no teeth on the tongue. Eye of
moderate size.
Gill-opening very wide : no gill-rakers : no pseudobranchiaj. Branchioste-
gals numerous.
Dorsal fin short, placed far forwards on the body, hardly a head-length
behind the head. Anal fin also short, placed far back near the caudal. Caudal
forked. Adipose dorsal small, opposite the anal. Pectorals and ventrals well,
developed, the ventrals long.
A long slender thin-walled air-bladder.
Distribution : Atlantic, Mediterranean, Arabian Sea and Bay of Bengal,
Western Pacific.
144
117. Chauliodiis Sloanii, Bl. Schn.
Chauliodus sloanii, Schneider, Bloch, Syst. Ichth. p. 430 : Cnvier and Valenciennes, Hiat. Nat. Poiss. XXII.
p. 383 : Giinther, Cat. Pishes, V. p. 392, and Challenger Deep-Sea Pishes, p 179: Goode and Bean, Bull. EsBex
Inst. XI. 1879, p. 22: Goode, Proc. U. S. Nat. Mus. III. 1880, p. 483 : Facciola, Nat. Sicil. II. p. 206 : Vinciguerra,
Ann. Mns. Genov. (2) It. p. 4f>9: A. Agassis, Bull. Mus. Comp. Zool. XV. 1888, p. 32, fig. 214: Cams, Prodr.
Faun. Medit. II. p. 570: Alcock, Ann. Mag. Nat. Hist. (6) IV. 1889, p. 399; VIII. 1891, p. 127: X. 1892, p. 355 :
Goode and Bean, Oceanic Ichthyology, p. 96, fig. 115 : Jordan and Evermann, Fishes N. Amer. p. 5S5.
Chauliodus setinosus, Schneider op. cit pi. lxxxv : Bonaparte, Fann. Ital. Pesc. fig.
Chauliodus Schneideri, Risso, Faun. Ear. Merid. III. p. 4t2 fig. 37.
Stomias Schneideri {Stomias boa) Cnvier, Regne An., Poiss., pi. 97, fig. 3 (too many teeth and lnminous spots).
? Chauliodus Macouni, Bean, Proc. D. S. Nat. Mus. 1890, p. 44.
B. 16-18. D. 6. A. 12. P. 12-13. V. 7. L. lat. 56.
Head much compressed squarish, its length, which is not much more than
its height, is from a sixth to a seventh of the total, without the caudal. Eye
circular, about as long as the snout proper, between a third and a fourth the
length of the head. Lower jaw prominent beyond the snout.
The dorsal fin begins less than a head-length behind the occiput ; the
filamentous first ray is from twice the length of the head to more than half the
body in length.
The longest ventral rays are nearly twice as long as the pectorals and about
a third as long again as the head.
Scales hexagonal. The luminous organs are disposed, on each , side, as
follows, and are conspicuous : —
(1) a aeries between the bases of the branchiostegals :
(2) a series of 61 in a lower tier from the chin to the caudal, of which 30
are between the chin and the ventrals :
(3) a series of 38 between the base of the pectoral and the anus, of which
17 are between the base of the pectoral and the base of the ventral.
(4) one below the eye, and (5) one in the upper part of the suboperculum.
Besides these there are countless tiny spots, in rows and clusters, all along
the abdominal line from the chin to the caudal.
Colours in spirit, silvery more or less ; fins white.
A ripe female in the Indian Museum is six inches long, but there is a muti-
lated specimen much longer.
Gulf of Manar, 597 fathoms ; Bay of Bengal, 922 and 1590 fathoms; Lacca-
dive Sea.
Dist7-ibuHon : as for the genus.
Regd. Nos. 11731-11733, 12473, 12837, 12838.
In one specimen dissected there were 3 large pyloric caeca, in another 3
large ones and a small one.
145
The ova are not arranged in a pleated band in the ovaries as they are in
Gonostoma.
The stomach is much like that of Gonostoma, having a remarkably thick
submucous coat, which, under the microscope, has much the structure of the
cortex of mammalian lymph-gland.
118. Chauliodus pammelas, Alcock.
Chauliodus pammelas, Alcock, Ann. Mag. Nat. Hiat., Nov. 1892, p. 355 : Illustrations of the Zoology op thi
Investigator, Fishes, pl. XXX. fig. 4.
B. 16. D. 6. A. 12. P. 11-12. V. 7.
Differs from C. Sloanii in the following points : —
The length of the eye is considerably more than that of the snout proper,
and is a third that of the head.
The skin is naked, although it is mapped out into rhomboidal and hexagonal
depressions each of which has a luminous spot in the centre, and in life was
covered with a thick coat of transparent mucus tissue full of capillary blood-
vessels.
The luminous organs are much less conspicuous.
The body fins and iris are uniform jet black, which has not altered after
nearly seven years' immersion in spirit.
Length 10 inches.
Arabian Sea, in the neighbourhood of Minnikoy, 1370 fathoms.
Regd. No. 13183.
This is undoubtedly a good species.
Family Stomiatidce.
Skin generally naked, sometimes with thin deciduous scales. A hyoid
barbel, which may be either free or attached to the mandibular sym-
physis. Margin of the upper jaw formed by the premaxilla and maxilla,
which are both toothed. Opercular apparatus little developed or rudi-
mentary. Gill-opening, like the mouth-cleft, very wide. Pseudobranchiae
none. Adipose fin present or absent. Air-bladder present or absent.
The presence of four representatives of this family has been brought to
light by the " Investigator."
I have followed Dr. Gunther in keeping this family distinct. My own
opinion is that it might be united with the Sternoptychidae.
19'
146
Synopsis of the Indian genera of Stomiatidse.
I. Pectorals present : hyoid barbel free : skin mapped out into subhexagonal
areolffi : teeth on tbe palatines and vomer ... ... ... Stomias.
II. Pectorals rudimentary : skin perfectly smooth and scaleless : hyoid barbel
attached to tbe mandibular symphysis : no teeth on the palate ... Malacosteps.
III. Pectorals absent : skin perfectly smooth and scaleless : hyoid barbel
attached to the mandibular symphysis : teeth on the palatines ... Photostomias.
Stomias, Cuv., Gthr.
Stomias, Cuvier, Regne Animal, Poiss. p. 232: Giinther, Cat. Fishes, V. p. 426: Goode and Bean, Oceanic
Ichthyology, p. 107 : Jordan and Evermann, Fishes of N. Araer. p. 588.
Body low, elongate, compressed, vent situated at no great distance from the
caudal fin. Skin with subhexagonal impressions in which deciduous scarcely-
imbricate scales may be present. Head compressed, with the snout very short
and the mouth-cleft oblique and very wide. Teeth pointed, unequal in size,
those of the premaxilla and mandible the longest ; maxilla finely denticulated :
vomer with a pair of fangs : palatines and tongue with smaller pointed teeth.
Eye of moderate size. A fleshy barbel in the centre of the hyoid region.
Opercle narrow, incomplete.
Dorsal opposite the anal, close to the caudal. Pectorals and ventrals
narrow, the ventrals situated very far back in the posterior third of the body.
No adipose dorsal.
Series of phosphorescent dots along the lower side of the head body and tail.
An air-bladder. No pyloric appendages.
Distribution : North Atlantic and West Indies ; Atlantic gate of Medi-
terranean ; Mediterranean ; Arabian Sea.
Key to the Indian species of the genus Stomias.
I. Height of the body about a twelfth of the total length (with caudal) ... S. nebulosus.
II. Height of the body about a sixteenth of the total length (with caudal) ... S. elongatus.
119. Stomias nebulosus, Alcock.
Stomias nebulosus, Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 451. Illustrations of the Zoology of the
Investioator, Fishes, pl. VII. fig. 1.
This species may prove to be identical with the West Indian S. afflnis.
D. 17. A. 21. P. 6. V. 5.
Length of the head one-ninth, height of the body one-twelfth of the total.
Snout shorter than the large eye.
147
Cleft of mouth oblique, enormous ; the limbs of the mandible widely dis-
tensible. Teeth fixed ; upwards of twenty-five small, unequal, and curved in
each premaxilla, and about the same number, in the form of minute, close-set,
even serrations, in each maxilla ; a fang on each side of the vomer ; one or two
moderate-sized teeth in the palatines. The teeth of the lower jaw are very
large, curved and acute, and stand out laterally, eight or nine on each side,
almost at right angles to the jaw.
Barbel about as long as the head and ending in three longish filaments.
The bony part of the opercle is reduced to a small preoperculum.
The surface of the body is covered with a tenacious slime. There are no
scales, but the body is mapped with regular rows of hexagonal depressions, each
with a minute central white point.
A salient white line, composed of a multitude of (luminous ?) specks, runs
along the mid- ventral line, from the throat to the anal fin. Two rows of lumin-
ous organs on each side of the abdomen, the lower one extending from the
isthmus to the base of the caudal, the upper extending from the base of the
pectoral to the origin of the anal.
Those of the lower row are disposed as follows : — 6 from the isthmus to the
base of the pectoral, 34 from the pectoral to the base of the ventral, 9 from the
ventral to the anal, 15 above the course of the anal, — in all 64.
Those of the upper row are 85 or 36 in number.
The dorsal fin begins in the last fifth of the body, a little in rear of the
commencement of the anal, which is also the deeper. Caudal not forked. The
pectorals arise on very narrow bases near the ventral profile ; their length is
equal to the height of the body. The ventrals are also narrow and are exceed-
ingly prolonged, reaching beyond the origin of the anal.
Colours in spirit : — Uniform black ; fins and barbel white, with black tips.
Two specimens, rather mutilated, the longer 3- inches.
Gulf of Manaar, 597 fathoms.
Regd. Nos. 11734, 11735.
120. Stomias elongatiis, Alcock.
Stomias elongatus, Alcock, Ann. Mug. Nat. Hist., August, 1891, p. 129.
D. 19. A. 21. P. 6. V. 6.
Length of the head one-tenth, height of the body one-fifteenth of the total
without the caudal. Eye longer than the snout, not quite a fourth the length of
the head.
148
The widely-distensible mandible projects much beyond the upper jaw.
Five large, distant, fixed fangs in each premaxilla, as well as a freely movable
one near the symphysis ; a few minute, inconspicuous, distant denticulations in
the maxilla? ; eight or nine moderate-sized laterally-projecting fangs on each
limb of the mandible, decreasing in size from before backwards ; a fang on each
side of the vomer, and two small, distant, incurved teeth on each palatine.
The barbel, which is as long as the caudal fin, is trifid at its extremity.
Opercular bones membranaceous.
No scales ; the body, which is coated with tenacious mucus, is mapped out
into silvery hexagonal areolae. There are on each side along the ventral surface
of the body two rows of small luminous organs ; the internal extends from the
mandibular symphysis to the base of the caudal, but, owing to the denudation of
the integuments of the tail, the number of its constituents cannot be determined
beyond the origin of the anal fin, up to which point there are 57, namely, to the
base of the pectorals 9, to the base of the ventrals 51, to the origin of the anal
57 ; the external extends from the base of the pectoral to the origin of the anal,
and numbers 45. There is a single luminous organ on the barbel and a row
along the base of the branchiostegal rays. The dorsal fin arises at the level of
the third anal ray. Caudal pointed, its length is about one-twelfth of the total.
The pectorals, which arise near the ventral profile, are equal in length to the
caudal. The ventrals are very long, reaching to the sixth anal ray.
Colours in the fresh state : — jet black, with silvery hexagonal markings. A
long slender air-bladder is present.
One specimen, a little over 5 inches long.
Arabian Sea, off the Laccadives, 738 fathoms.
Re^d. No. 13075.
o
Malacosteus, Ayres.
Malncosteun, Ayrea, Journal Bostou Soc. Nat. Hist. 1849, p. 53 : Giinther, Cat. Fishes, V. p. 427. and Chal-
lenger Deep Sea Fishes, p. 212: Goode and Bean, Oceanic Ichthyology, p. 113: Jordan and Evermann, Fishes N.
America, p. 592.
Differs from Stomias in the following respects : —
The body is not so low and elongate, and the skin is perfectly smooth and
sea It 'less. The head is not so much compressed, and the gape is even wider, the
ends of the jaws extending beyond the root of the pectoral. There are no teeth
on the palate. The opercle is membranaceous. The pectorals are rudimentary,
and the ventrals are not placed so far back, being well in front of the posterior
third of the body.
Distribution: Atlantic; Andaman Sea; off Philippines.
149
121. Malacosteus sp.
? Malacosteus niyer, Ayres, I. c. pl. v. Goode and Bean, Oceanic Ichthyology, p. 114, fig. 138.
? M<dacosteus indicus, Giinther, Alcock, Ann. Mfig. Nat. Hist., (6) IV. 1889, p. 452 : Illustrations of thi
Zoology of the Investigator, Fishes, pl. XXXIII. fig. 4.
A species of Malacosteus was dredged in the Andaman Sea in 650 fathoms.
It is represented by a single damaged specimen, and appears to be identical with
Malacosteus indicus Giinther, of the Challenger Report, p. 214, pl. liv. fig. B.
At the same time, judging from the descriptions and figures of the two
species, it seems to me that Malacosteus indicus and Malacosteus wiger are
the same.
The following is a brief description of the " Investigator " specimen, so far
as description is possible : —
D. 18. A. 20. P. 3. V. 6.
The gill-cleft is so wide, and the neck so narrow, that the head can be
turned completely upside down over the back without hurting the specimen.
Length of jaws and mouth-cleft more than a third of the distance between
the tip of the snout and the origin of the anal fin.
There is almost no snout distinct from the rim of the orbit : eye between a
third and a fourth the length of the lower jaw.
The largest teeth are a pair of curved outstanding fangs at the fore end of
the mandible. Of the lateral mandibular teeth two exceed the others in size,
the anterior one being not very far from the mandibular symphysis, the posterior
one being a good way back.
Skin smooth, soft, black, everywhere covered with tiny white dots. A large
petal-shaped luminous organ, as long as the orbit, beneath the eye, and a smaller
oval one on the cheek near the middle of the upper jaw.
Dorsal and anal fins nearly equal and opposite, close to the caudal. Caudal
small, its peduncle narrow, — the height midway between the anal and the caudal
fin being about half an eye-length. The ventrals arise about an eye-length
behind the middle of the body.
Length 3- inches.
Regd. No. 11642.
Photostomias, Collett.
Photostomias, Collett, Ball. Soc. Zool Frnnce, 1889, p. 291, and Hirondelle Poissons (Monaco, 1896), p. 131 :
Goude and Bean, Oceanic Ichthyology, p. 114.
Thaumastomias, Alcock, Ann. Mag. Nat. Hist., Sept. 1890, p. 220.
Body elongate, compressed, scaleless, with the vent not far distant from the
caudal fin. Head compressed, with the cranium small, the snout short, and the
150
cleft of the mouth exceedingly wide. A long elastic muscular band passing from
the hyoid bone to the inner aspect of the mandibular symphysis. Teeth acute,
unequal, in single series in premaxilla?, maxilla?, mandibles, and palatines ; none
on the tongue. Bye moderate. Gill-covers rudimentary. One dorsal fin
opposite to the anal, situated in the posterior fourth of the body, near the caudal.
No adipose dorsal. No pectoral fins. Ventral fins situated in the anterior half
of the body. Gill-openings very wide. No air-bladder.
A large luminous organ behind the orbit : two long rows of small luminous
organs on either side of the body, on the ventral aspect, from the gill-opening to
or nearly to the caudal fin.
Distribution : North Atlantic and Azores ; both sides of the Bay of Bengal.
122. Pliotostomias atrox (Alcock).
Thaumastomias atroz, Alcock, Ann. Mag. Nat. Hist., Sept. 1890, p. 220, pi. viii fig. 7, and Aug. 1898, p. 147:
Illustrations of the Zoology of the Investigator, Fishes, pl. XXX. fig. 2. [See also Goode and Bean, Oceanic
Ichthyology, fig. 141, which, however, has been incorrectly copied from my figure in the Annals and Magazine of
Natural History, a second large post-orbital luminous organ, of which there is no trace either in the specimen or
in my drawing and description, having been added.
D. 23. A. 25. C. eirc. 25. P. 0. V. 6.
Length of the head one-fifth, height of the body one-tenth, of the total
without the caudal.
Snout truncated, broad, with a slightly concave vertical profile, its length
one-third the diameter of the eye. Eye large, circular, its diameter about one-
fourth the length of the head ; interorbital space wider than the eye, convex.
On each side there are two luminous organs, one about the size and shape of a
caraway-seed, below and partly in front of the eye, the other large, salient,
slipper-shaped, and more than one-third the length of the head, lying parallel
with the upper jaw behind the eye. Mouth enormous, its cleft as long as the
head ; its floor is completely wanting except quite anteriorly, its place being
taken by a long elastic muscular band which extends from the tip of the hyoid to
the inner surface of the mandibular symphysis ; the mouth-cleft and the gill-cleft
being thus continuous beneath, almost divide the head from the rest of the body ;
the lower jaw projects beyond the upper. Teeth, everywhere except in the
maxilla, in the form of slender acute rigid fangs ; in each premaxilla eight or
nine, and three remote stouter ones at the symphysis; in each half of the
mandible an uneven row of over twenty, and five (one median flanked on each
side by a pair) of superior size at the symphysis ; in each palatine a row of seven
or eight, increasing in size from before backwards, and a patch on the upper
pharyngeal bones; maxillary teeth in the form of even, close-set, recurved
serrations, of which there are over thirty in each bone.
151
Gill-cleft extremely wide and oblique, its superior limit being above the
middle of the eye ; gill-cover reduced apparently to a narrow straight preoper-
culum, very obliquely articulated, furnished with a narrow membranous fringe ;
four branchial arches, extremely weak and flexible, bearing very narrow lamina? ;
gill-rakers rudimentary.
Body scaleless. Skin thick, soft, velvety, and uniformly covered with
adherent tenacious mucus ; apparently no lateral line. Besides the large lumin-
ous glands already described, there are two regular rows of minute luminous
organs along the ventral half of the body on each side : the upper, numbering
about fifty, extending from the gill-opening to the base of the caudal ; the lower,
numbering about forty, skirting the ventral profile from the isthmus to the fifth
anal ray ; a few similar luminous organs on the crown of the head.
The dorsal fin begins slightly in advance of the posterior fifth of the body,
and is equal and opposite to the anal. The longest (central) anal rays are a little
longer than the corresponding dorsal rays, and are equal to the depth of the tail
at their point of origin. The caudal is deeply forked, with the lower lobe the
broader and longer and about 57, of the total length.
Pectorals absent. The ventrals arise in the anterior half of the body, their
point of origin being 1- times as far from the vent as from the margin of the
gill-cleft ; the two outer rays are thickened, coherent throughout, and prolonged,
their length being two-fifths of the total length including the caudal ; the inner
rays are short, weak, and inconspicuous.
Stomach siphonal, its cul-de-sac extending halfway along the abdominal
cavity ; intestine straight, opening at the origin of the anal fin. No pyloric caeca.
Colours intense black : the large postocular luminous organs very conspi-
cuous, one being naked and rose pink in colour, the other being silvery and
almost entirely covered by a fold of black skin. The small luminous organs
were not visible until after immersion in spirit.
The largest specimen is just under 5 inches long.
Bay of Bengal, off Ganjam coast, 1310 fathoms, and off the Andamans, 606
fathoms.
Regd. Nos. 12835, ^.
Family Scope\ida>.
In the Fauna of British India four genera and six species are recorded.
We now know of six more genera and thirteen more species, all belonging to the
deep sea.
152
In the following synopsis the genera added to the Indian list by the
Investigator are marked with an asterisk, and those that are represented in the
deep sea by a dagger.
Synopsis of the Indian genera of Scopelidge.
I. Pseudobranchise well developed : —
1. Body scaly, without phosphorescent spots : —
i. Maxilla not dilated posteriorly : —
a. Dorsal in front of the ventrals : some of the teeth
enlarged and barbed at point : eyes with a strong
upward cast ... ...
6. Dorsal fin above or behind the ventrals : none of
the teeth barbed : eyes quite lateral : —
a. Teeth on each palatine in a single band ...
/3. Teeth on each palatine in a double band ...
ii. Maxilla rudimentary or absent : some of the teeth barbed
at point : caudal three-lobed ... ... ...
iii. Maxilla dilated behind : —
a. Eyes lateral, very large : no prolonged rays
b. Eyes very small : some of the rays of the pectoral
(and also sometimes of the caudal and anal fins)
enormously prolonged
2. Body scaly, with phosphorescent spots : —
i. Snout high, short, bluntly rounded : mouth-cleft nearly
reaching the angle of the preoperculum : anal fin close
behind the dorsal
ii. Snout depressed, rather long : mouth-cleft wide, but not
nearly reaching the angle of the preoperculum : anal a
considerable distance behind the dorsal
3. Body naked : some enormous fangs in the mandible, vomer and
palatines
II. Pseudobrauchioe rudimentary : otherwise closely related to Neoscopelus
•ScOPELARCHOS.f
Saurus.
Saurida.
Earpodon.f
*CHLOROPHTEALMUS.f
*BATHVPTEKOIS.t
Scopelus.f
*NEOSCOPELUS.f
*ODONTOSTOMUS.t
*SC0PELENTGYS.f
SCOPELARCHUS, Alcock.
Scopelarchus, Alcock, Journ. As. Soc. Bengal, Vol. LXV. pt. 2, 1896, p. 306.
Body elongate, compressed, covered with scales which, except on the lateral
line, where they are adherent, are deciduous.
Cleft of mouth very wide : premaxilla very long, tapering, firmly attached
to the long slender maxillas. A single row of small teeth in the premaxilla : a
double row of teeth in the mandible, the inner row being large depressible and
barbed at tip ; an incompletely double series of similarly enlarged teeth on either
palatine, and a long narrow row of almost similar teeth on the tongue and
hyoid. Eye large. Gill-openings very wide, gill-membranes not attached to the
isthmus, branchiostegals not very numerous (about G ? ) ; pseudobranchire
large.
153
The dorsal fin is short, it arises well in the anterior third of the body
(measured with the caudal) and all its extent lies between the pectorals and
ventrals : the anal is long, occupying the greater part of the tail. Pectorals
large. Ventrals with 8 rays. An adipose dorsal fin. Caudal forked. No
luminous spots.
This is a remarkable generalized form of Scopeloid, showing affinities with
Saurus, CMorophthalmus, Scopelus, Odontostomus, and Paralepis. To casual view
it looks just like a Scopelus devoid of luminous organs.
123. Scopelarchus Guntheri, Alcock.
Scopelarchus Ountheri, Alcock, Journ. As. Soc. Bengal, Vol. LXV. pt. 2, 1896, p. 307: Illustrations op the
ZOOLOGT OF THE INVESTIGATOR, FlSHES, PL. XVII. FIG. 7.
D. 9. A. 26. P. 19. V. 8. L. lat. (of enlarged adherent scales) circ. 50.
Head and body compressed : shape as of Scopelus.
Length of head (with gill-cover) not quite one-fourth, height of body about
two-elevenths of the total (without caudal). Snout about three-fourths the
length of the eye : the lower jaw in repose fitting within the upper. The eyes
are large — about one-third the length of the head — they are separated from one
another by a mere linear space, and their visual axis is rather more superior
than lateral.
The mouth-cleft forms a slightly oblique sweep, and the maxilla extends a
considerable distance behind the posterior border of the orbit.
The scales of the lateral line are much enlarged, and their vertical diameter
is much greater than their antero-posterior diameter ; each is chambered, the
chamber opening dorsally and ventrally.
The first dorsal ray arises about an eye-length behind the base of the
pectorals, the last stands a little in advance of the base of the ventrals. The
first anal ray arises near the middle of the body (measured with the caudal), the
last is less than an eye-length distant from the rudimentary rays at the base of
the caudal. The adipose fin stands in the posterior third of the distance between
the dorsal and caudal.
Pectorals broad and falciform, reaching to the base of the ventrals, which
are small and do not nearly reach the anal.
Colours in spirit white, occiput and caudal peduncle black.
A single specimen, an adult female, about five inches long, from off the
Indus Delta, 947 fathoms.
Regd. No. y-
20
154
Harpodon, Lesueur, Giinther.
124. Harpodon squamosus, Alcock.
Earpodon squamosus, Alcock, Ann. Mag. Nat. Hist. August, 1891, p. 127 : Illustrations op the Zoology or
thb Investigator, Fishes, pl. XXX. fig. 1.
B. 17. D. 12-14. A. 13-15. P. 10. V. 9.
The length of the head, measured to the edge of the operculum and not to
the end of the produced branchiostegal rays and membrane, is about one- fifth,
the height of the body about one-sixth of the total, without the caudal. The
vertex of the head with numerous minute mucous pores.
Snout broad, depressed ; its tip is formed by the projecting lower jaw, and
its length, including the mandibular element, slightly exceeds the major diameter
of the eye, which is about one-eighth the length of the head as above limited.
The width of the flat interorbital space is twice the vertical diameter of the
eye.
Mouth-cleft oblique, wide : the maxilla is nearly two-thirds the length of
the head as above limited. Introrsely-depressible cardiform teeth in bands in
both jaws ; one series in the lower jaw enlarged, with barbed hastate tips, and
one series in the upper jaw less enlarged ; in each palatine an outer irregularly-
double row of teeth, of which the anterior and external are enlarged, and a very
short inner irregularly- double row ; hyoid bone and all the branchial arches
toothed.
Gill-openings extremely wide ; the branchiostegal rays and membrane much
produced beyond the operculum.
Body, posterior part of head, and cheeks covered with deciduous cycloid
scales, which are less deciduous on the posterior half of the tail.
The dorsal fin arises within the anterior half of the body (measured with
the caudal) almost opposite to the origin of the ventrals. The anal arises
about an eye-length behind the vent, which is nearly twice as far from the gill-
opening as from the base of caudal. The fimbriated adipose dorsal is situated
far back, above the posterior half of the anal. Caudal deeply forked, with an
inconspicuous median lobe. Ventrals long, delicate, and feathery, the longest
(middle) rays reach to within two eye-lengths of the vent in the adult. Pectorals
very narrow and fragile ; they arise almost on the same plane with the eyes, and
their longest (middle) rays do not nearly reach to the dorsal fin, being about as
long as the postorbital portion of the head.
Stomach with a very long ca)cal sac ; eighteen large pyloric cteca in a
pectinate arrangement.
155
Colours in life: — hyaline grey; paired fins and caudal black, visceral
peritoneum black, buccal and branchial cavities partially and slightly pigmented.
Bay of Bengal, 240 to 276 and 281 to 258 fathoms.
Mature females are from 9 to 10^ inches long ; the males are from 7g to 8-
inches long.
Regd. Nos. 13084 to 13095, 13209, 13210.
Chlorophthalmus, Bonaparte.
Chlorophthalmus, Bonaparte, Fann. Ital. Pesci: Giinther, Cat. Fishes, V. p. 403, and Challenger Deep Sea
Fishes, p. 192 : Goode and Bean, Oceanic Ichthyology, p. 60 : Jordan and Evermann, Fishes N. Amer., p. 541.
Hyphalonedrus, Goode, Proc. U. S. Nat. Mus. IV. 1881, p. 483.
Form of the body elongate, subcyhndrical or compressed, covered with
scales. Head rather long, with the lower jaw usually projecting. Mouth-cleft
wide, the maxilla dilated bebind. Teeth minute, in narrow bands on the jaws
vomer and palatines. Eye large.
Gill-openings very wide : pseudobranchia? well developed. Branchiostegals
10 to 7.
Dorsal short, situated in the anterior half of the body : anal short, situated
in the posterior half of the body. Caudal forked. Adipose fin small.
Pectorals and ventrals well developed : the ventrals inserted at no great
distance behind the pectorals, under or somewhat behind the dorsal.
Distribution : "West Indies and Atlantic coasts of North America : Mediter-
ranean : Bay of Bengal : "Western Pacific.
125. Chlorophthalmus comiger, Alcock.
Chlorophthalmus comiger, Alcock, Jonrn. As. Soc. Bengal, Vol. LXIII. pt. 2, 1894, p. 133, pi. vi. fig. 5 : Illus-
trations of the Zoology of the Investigator, Fishes, pl. XV. fig. 8.
B. 8. D. 11. A. 9. P. 14. V. 9. L. lat. circ. 55.
Length of the head about two-fifths of the total without the caudal, greatest
height of the body about half the length of the head.
Eye not quite so long as the snout (including the mandibular element) not
quite a third the length of the head, about 3 times the width of the interorbital
space.
The mandibular symphysis forms a strongly projecting, transverse, hori-
zontal plate, of which the angles are dentiform. The maxilla reaches to the
anterior edge of the pupil. Teeth minute, in narrow bands in the jaws : very
inconspicuous on the vomer and palatines.
Gill-rakers of the first arch numerous, close-set, bristle-like.
156
The first few rays of the dorsal fin are in front of the ventrals. The
adipose fin is as far behind the dorsal as the dorsal is behind the anterior edge
of the pupil.
The pectorals are as long as the head behind the snout, and reach nearly
to the tips of the ventrals : the latter are rather more than half the total length
of the head.
Colours in spirit : silvery grey with numerous broad ill-defined dusky cross-
bands ; fins hyaline, the tip of the caudal and the base and tip of the dorsal black :
numerous parallel oblique rows — very conspicuous on the thorax and belly —
of tiny black specks with a silvery centre, resembling incipient luminous
spots.
Seven large pyloric casca.
The largest specimen is a little over 3 inches long.
Bay of Bengal, off Madras coast, 145 to 250 fathoms.
Regd. Nos. 13712 to 13715.
"ov
Bathypteeois, Giinther.
Bathypterois, Giinther, Ann. Mag. Nat. Hist., 1878, Vol. II., p. 183 ; Challenger Deep Sea Fishes, p. 185 : Goode
and Bean, Oceanic Ichthyology, p. 61 : Jordan and Evermann, Pishes N. Amer., p. 544.
Form of the body elongate, slightly compressed. Scales cycloid, of moderate
size. Head low ; with a long broad depressed bill-like snout, the end of which
is formed by the prominent mandible. Cleft of the mouth wide ; maxillary
much developed, very movable, dilated behind. Villiform teeth in narrow
bands on the jaws : vomerine teeth present or absent : no teeth on the palatines
or tongue. Eye very small.
Gill-openings very wide : gill-laminaa well developed : gill-rakers long and
numerous. Branchiostegals numerous. No pseudobranchias.
Dorsal fin in the middle of the back, above or just behind the root of the
ventral, rather short. Anal rather short, below or just behind the dorsal.
Caudal forked, its lowermost rays sometimes prolonged. Adipose dorsal present
or absent.
Pectorals remarkably developed, at least their uppermost rays are isolated
and enormously prolonged. Ventrals abdominal, well developed, their outer-
most rays usually prolonged.
No luminous spots.
Distribution : Atlantic ; Arabian Sea, Bay of Bengal and Andaman Sea ;
Western and Southern Pacific.
157
Key to the Indian species of the genus Bathypterois.
I. Ventral edge of the caudal peduncle not notched : —
1 . Lowermost caudal and ventral rays enormously prolonged, nearly 3
times as long as the head in the adult, longer in the young ... B. Guntheri.
2. Lowermost caudal and ventral rays moderately prolonged, the former
about once aud a half, the latter not quite twice the length of the head B. insular um.
II. Ventral edge of the caudal peduncle curiously notched : the outer ventral rays
slightly, the lowermost caudal rays very slightly prolonged ... ... B. atricolor.
126. Bathypterois Guntheri, Alcock.
Bathypterois Oiintheri, Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 450, and August 1891, p. 129: Illustra-
tions of the Zoology of the Investigator, Fishes, pl. VII. fig. 6.
B. 12. D. 13. A. 11. P. 2/6/5. V. 7-8 L. lat. circ. 55.
Body elongate and compressed, its height about one-sixth of the total,
without caudal. Head contained nearly three and a half times in the same
measure ; depressed, flat-crowned, nearly as broad as deep. Snout broad, de-
pressed, rounded, duck-bill shaped, with a median intermaxillary notch, into
which a strong recurved projection of the very prominent mandible fits ; its length
one-third that of the head; its surface with numerous large pores. A wide
mucous channel with a line of large pores along the under surface of the broad
mandible. Eyes minute, situated near the middle of the maxilla, close to its
edge, a snout-length apart ; the orbital margins rounded and inflated. Interorb-
ital space nearly flat from side to side. Nostrils small, superior, far in advance
of the eye.
Cleft of mouth extremely wide, slightly oblique ; the maxilla, which has a
dilated, abruptly-truncated, hinder end, is nearly two-thirds the head-length.
Villiform teeth in broad bands on the outer edges of the strong jaw-bones, and
in a minute patch on each side of the expanded vomer.
Gill-cleft reaching to the fore end of the isthmus ; gill-lamina? broadish ;
gill-rakers numerous, close-set, long, bristle-like, except on the fourth arch.
Body and head, except the jaws and front part of the vertex of the snout,
covered with large, thin, smooth scales, those on the sides of the head rather
deciduous, those on its crown enlarged.
The interradial membrane of all the fins except the caudal is covered with a
thick, black, velvety, deciduous integument. The dorsal begins a little in
advance of the middle line, and is just entirely in advance of the anal, the two
fins being of nearly equal extent and height. A very small adipose dorsal in the
posterior half of the tail. Caudal large and deeply forked ; its lowermost ray
rigid, prolonged, curved, with a spatulate tip, the total length of the ray from
base to tip being nearly two-thirds that of the rest of the body.
158
The pectoral consists of three distinct portions : — (1) an upper portion of
two detached, rigid filaments, the first of which is the longest and, though broken,
reaches to the tip of the upper lobe of the caudal ; (2) a middle portion of six
comparatively short branched rays, diminishing in length from above downwards
and connected together by a stout interradial membrane ; and (3) a lower portion
of five free, simple, elongated rays, which reach at least halfway along the
tail.
The ventrals arise just in front of the dorsal ; the outermost ray of each
fin forms a long, curved, rigid, spatulate appendage, which is nearly as long as
the body in the adult and longer than the body in the young.
Colours in spirit : — Head nearly black ; body dark brown, with two broad,
transverse, white bands, one just in front of the dorsal fin, the other near the
middle of the tail ; caudal white ; the other fins black, except their prolonged
rays, which are translucent white, with black tips. A large, opaque-white,
three-lobed body shows through the bones of the crown of the head and snout,
and there is a white streak along the mucous canal of the mandible.
Length of adult female 10 inches (not including the prolonged caudal ray).
Andaman Sea, 490 fathoms ; Bay of Bengal, off Andaman Is., 561 fathoms ;
Arabian Sea, off the Laccadives and Maldives, 636 and 719 fathoms.
Regd. Nos. 11770, 13706, 14001, 14002.
In the young the prolonged caudal and ventral rays are relatively much
longer than they are in the adult.
127. Bathypterois insularum, Alcock.
Bathypterois insularum, Alcock, Ann. Mag. Nat. Hist., Nov. 1892, p. 356.
B. 13-14. D. 12-13. A. 10. P. 2/12-13. V. 9. L. lat. 48-51. L. tr. 13.
Body elongate, its height a little more than half the length of the head,
which is about one-fourth of the total without the caudal. The snout, which has
the typical duck-bill shape, is in length a little more than one-third the length of
the head. The very small eyes are not quite a snout-length apart. There is
nothing peculiar about the mouth, but there are no teeth on the vomer. The
branchial structures are identical with those of other species of the genus. The
body and the head, except the jaws and snout, are covered with thin deciduous
cycloid scales.
The dorsal fin begins half a snout-length behind the base of the ventrals,
and the anal immediately behind the vertical through the last dorsal ray ; there
is a small adipose fin nearly midway between the dorsal and the base of the
caudal. The two uppermost pectoral rays are intimately coherent in their basal
159
half and reach at least as far as the adipose dorsal ; the other pectoral rays,
which are slender and rigid, reach at least as far as the vent. The ventral fins
are very large, their two outermost rays, which are very stout and stiff, are
prolonged, being about 1- times to twice as long as the head. The two or three
lowermost rays of the forked caudal are prolonged, their length being at least
one-third that of the rest of the body.
Colour black ; fins hyaline grey.
Length of the adult female, h- inches.
Arabian Sea, off the Laccadive Islands, 1140 fathoms.
Eegd. Nos. 13187, 13188.
128. Bathypterois atricolor, Alcock.
Bathypterois atricolor, Alcock, Journ. As. Soc. Bengal, Vol. LXV. pt. 2, 1896, p. 306 : Ann. Mag. Nat. Hist.,
Aug. 1898, p. 146: Illustrations of the Zoology op the Investigator, Pishes, pl. XVII. fig. 6.
B. 12. D. 15. A. 10. P. 2/12. V. 9. L. lat. 52. L. tr. 15.
Length of head a little more than one-fifth, height of body about one-eighth,
of total (without caudal).
Length of snout a little more than one-third that of head, and a little more
than 5 times that of eye, equal to width of interorbital space.
Few or no teeth on the vomer.
The dorsal fin arises just behind the vertical through the base of the
ventrals, and nearly half its extent is in the anterior half of the body (measured
without caudal) : the anal arises just behind the level of the last dorsal ray : the
adipose fin is halfway between the end of the dorsal and the base of the caudal :
the lower caudal lobe is hardly prolonged. Upper two pectoral rays prolonged
at least as far as the end of the caudal fin, coherent in basal part but not fused :
outer two ventral rays thickened, unbranched, prolonged as far as 7th or 8th
anal ray, not quite half again as long as the head.
Colours, uniform black, except the pectoral filaments.
The ventral edge of the caudal peduncle is most curiously notched as in
B. dubius Vaillant, to which this species is closely related.
Length, a little over 8 inches.
Arabian Sea, near the Laccadives, Maldives, and C. Coinorin, 891, 459 and
824 fathoms.
-n •, AT 62 166
Eegd. JNos. yj — .
160
Scopelus, Cuv., Giinther.
Scopelus:, Giinther, Cut. Fishes, V. pp. 404, 405 ubi synon.
Myctophidie (exo. Nannobrachium and Scopelosaurus) Goode and Bean, Oceanic Ichthyology, p 70, and Jordan
and Evermann, Fishes N. Amer., pp. 551, 5.52.
Body moderately elongate and compressed, covered with large scales of
which those of the lateral line are generally the largest. Series of luminous
spots run along the lower side of the head body and tail, and a luminous body
often occupies the snout and the back of the tail. Head generally compressed,
with the bones thin but ossified. Cleft of the mouth very wide : premaxilla long,
styliform ; maxilla well developed. Villiform teeth in bands in the jaws, on the
palatines pterygoids and tongue, and sometimes on the vomer. Eye large.
Dorsal fin short ; in, or nearly in, the middle of the back ; anal short or of
moderate length. Caudal forked. Adipose fin small. Pectorals and ventrals
well developed, the latter inserted in front of, or below, the anterior part of the
dorsal, and eight-rayed.
Gill-opening very wide, the outer branchial arch with numerous long gill-
rakers. Branchiostegals from 8 to 10. Pseudobranchia? present.
Air-bladder small. Pyloric caaca few.
Distribution : Pelagic and Nectic.
The Indian species of this genus are now known to be nine in number.
Two of them, however, are represented in the Investigator collections only by
specimens recovered from the stomach of other fishes, and are too much damaged
for recognition. The remaining seven fall into the following sub-genera.
Indian sub-genera of the genus Scopelus.
I. Snout deep, short, rounded : dorsal fin just in advance of the anal : pseudo-
branchiffi well developed : the anal fin with as many rays as, or more rays
than, the dorsal: eye large : scales smooth. ... ... ... Myctophum.
II. Snout rather long, somewhat depressed, not rounded : dorsal fin a consider-
able distance in advance of the anal : the anal fin with as many rays as,
or more than, the dorsal : —
1. Pseudobrauckioe well developed : eye moderate : scales with minute
spines ... ... ... ... ... ... Neoscopelus.
2. Pseudobranchia? rudimentary : eye small. (Scales unknown) ... Scopelengys.
Key to the Indian species of the sub-genus Myctophum.
I. A large luminous mass on the snout immediately in front of the orbit :
pectorals short : —
1. The pectorala do not reach to the base of the ventrals : all the
phosphorescent spots are divided by a median transverse black
septum ... ... ... ... ... ... S. engraxdis.
2. The pectorals reach just beyond the base of the ventrals : phosphores-
cent organs without a black septum ... ... ... S- dumerilii.
161
II. No large luminous mass on the snout : pectorals of moderate length or
very long : —
1. The pectorals reach the middle of the ventrals : snout about half as
long as the eye ... ... ... ... ... S. indicus.
2. The pectorals reach the 1st anal ray : snout about half as long as the
eye ... ... ... ... ... ... S. pterotus.
3. The pectorals reach at least to the sixth anal ray : snout about a
quarter as long as the eye ... ... ... ... S. pyrsobolus.
129. Scopelns engraulis, Gthr.
Scopelus engraulis, Giinther, Challenger Deep Sea Fishes, p. 197, pi. li. fig. C : Alcock, Ann. Mag. Nat. Hist.
(6) VIII. 1891, p. 129.
D. 14. A. 14-15. P. 12. V. 9. L. lat. 38. L. tr. |.
Height of the body about a fifth, length of the head about two-sevenths of
the total, without the caudal. The eye is near the extremity of the extremely
short snout, its diameter is about a fifth the length of the head and rather less
than the width of the interorbital space.
Operculum thin and narrow, scarcely covering the gill-opening : posterior
margin of preoperculum oblique.
Mouth oblique, very wide, the upper jaw overlapping the lower. The
maxilla extends back to the mandibular joint and is not dilated posteriorly.
The first dorsal ray is midway between the end of the snout and the adipose
fin and a little in advance of the root of the ventral : the last dorsal ray is just
in advance of the vent. Pectoral short, not reaching to the ventral. Ventral
reaching to the vent.
Scales perfectly smooth, those of the lateral fine not enlarged.
Colour : black ; mandibles whitish, with a broad black cross-band below the
eyes.
The luminous organs are arranged as follows on each side : —
one on the preoperculum, near its lower angle :
three between the isthmus and the root of the pectorals, and one above the
pectoral :
three, in an oblique series, between the pectorals and the root of the ventral,
and an isolated one higher up on the side :
three between the ventrals and the vent, and two or three higher up on the
side:
three in an oblique series running from the vent towards the lateral line :
eleven along the base of the anal and a little behind, and one higher up,
opposite a break in this series :
four along the root of the lower caudal rays.
21
162
All these luminous organs are divided into two by a black septum.
A large whitish, gland on the snout immediately in front of the eye.
A mature female is nearly 5- inches long.
Andaman Sea, 188 to 220 fathoms.
Regd. Nos. 13127, 13128.
Distribution : off Philippine Is. ; Andaman Sea.
130. ScopelUS DumerilU, Bleeker.
Scopelus Dumerilii, Bleeker, Act. Soo. Sc. Indo-Neerl. i. Manado and Macassar, p. 66 : Giinther, Cat. Fishes,
V. p. 410, and Challenger Deep Sea Fishes, p. 198.
D. 14-15. A. 14-15. P. 10. V. 9. L. lat. 37.
Very closely related to Scopelus engraulis, from which it only differs in the
following characters : —
The eye is larger, its diameter being from a third to two-sevenths the length
of the head and equal to the width of the interorbital space.
The first dorsal ray is rather nearer to the adipose fin than to the end of the
snout.
The pectoral fin reaches somewhat beyond the root of the ventral, and the
latter reaches beyond the vent.
The luminous organs correspond with those of S. engraulis, except that the
uppermost tier of four distant organs are placed much higher up, the last two
(which correspond respectively with the first and last anal rays) being on the
lateral line.
None of the luminous organs have a median black septum ; and the large
glandular body in front of the eye is much brighter.
A specimen nearly 3 inches long, from off the Malabar coast, 172 fathoms.
Regd. No. 13730.
Distribution : East Indian Archipelago ; Fiji ; Arabian Sea.
131. Scopelus pterotus, Alcock.
Scopelus {Myctophum) pterotus, Alcock, Ann. Mag. Nat. Hist., Sept. 1890, p. 217 : Illustrations of the
Zoology of the Investigator, Fishes, pl. IX. fig. 3.
D. 11-12. A. 17. P. 15. V. 8. L. lat. circ. 30.
Length of the head about a third, greatest height of the body about a fourth
the total without the caudal. Snout obtuse, its length hardly half the diameter
of the eye, which is a third the length of the head and rather more than the
mean width of the interorbital space.
163
Mouth large, moderately oblique ; the jaws perfectly equal in repose ; the
maxilla nearly reaches the preopercular angle and is dilated at its hinder end ;
no vomerine teeth. Opercles large ; the operculum produced into a membranous
spur behind ; the vertical border of the preoperculum very oblique.
Scales extremely deciduous, smooth, cycloid, their average diameter one-
twelfth of an inch.
The dorsal fin begins nearer to the tip of the snout than to the base of the
caudal, but behind the bases of the ventrals, which are much advanced, its last
ray falls in the vertical through the first or second anal ray ; adipose dorsal
entire. Pectorals long, extending beyond the tip of the ventral to the first or
second anal ray.
The luminous organs are arranged on each side as follows : — A series ex-
tending close to the mid-ventral line from the isthmus to the base of the caudal,
and numbering four to base of ventral, three more to origin of anal, ten or
eleven more to hinder end of anal, and one more at base of caudal ; above this series
are the following, rather more scattered — one at the angle of the preoperculum,
two along the ramus of the mandible, two along the edge of the gill-opening,
one on the base of the pectoral, two on the base of the ventral, three in a
straight line along the middle of the flank, two above the anal, and one at the
base of the caudal. No luminous organ on the back of the tail.
Nine pyloric ca?ca. An air-badder.
Colours in the fresh state : — Uniform silvery, with thickly scattered black
specks ; opercles and iris burnished silver.
Length of mature females from not quite two inches to a little over two
inches.
Bay of Bengal, 98 to 102 fathoms; Andaman Sea, 370 to 419 fathoms.
132. Scopelus pyrsobolus, Alcock.
Scopelus pyrsobolus, Alcock, Ann. Mag. Nat. Hist. Sept. 1890, p. 21S, pi. VIII. fig. 3; Illustrations of thb
Zoology of the Investigator, Fishes, pl. XXX. fig. 3.
D. 12. A. 13. P. 12. V. 8.
Length of the head (not including a membranous prolongation of the sub-
operculum) about a third, greatest body-height about a fourth of the total length
without the caudal. Snout about a quarter as long as eye. Eyes large, round,
strongly convergent, bulging beyond the dorsal profile ; their diameter is a third
of the length of the head proper, and more than the mean width of the inter-
orbital space. Mouth wide, oblique ; the maxillary, which does not quite reach the
preopercular angle, is slightly dilated behind. Jaws equal in front. Villiform
teeth developed on the vomer. Opercles large, extremely thin, the vertical
border of the preoperculum oblique.
164
The scales and almost all the luminous organs have been denuded : there
are opaque white glandular organs still left on the caudal peduncle, both dorsally
and ventrally.
The dorsal fin begins nearer to the tip of the snout than to the base of the
caudal, its first ray is almost in the vertical through the base of the ventrals ;
the entire fin is nearly half an eye-length in advance of the anal. Adipose fin
well developed.
The pectorals reach at least to the sixth anal ray.
About five large pyloric caeca. A well developed air-bladder.
Colours, apparently black. Iris and lower part of opercles like burnished
silver, the opercles in the fresh state brilliantly coruscating.
A mature female is over 3 inches long.
Bay of Bengal, off Madras coast, 920 to 690 fathoms.
Re^d. No. 12839.
"Bv
Neoscopelus, Johnson.
Neoscopelus, Johnson, Proc. Zool. Soc, 1863, p. -11 : Giinther, Cat. Fishes, V. p. 405 : Goode and Bean, Oceanic
Ichthyology, p. 92.
Neoscopelus differs from Scopelus in having a rather long and distinctly
depressed snout; a narrower mouth-cleft, which reaches only as far as the
posterior border of the orbit ; a smaller eye ; scales with minute spines, and an
anal fin much more remote from the dorsal. The maxilla also is much more
dilated posteriorly.
The argument, therefore, for recognizing it as a distinct genus is very
strong.
'&■
133. Neoscopelus macrolepidotUS, Johnson.
Neoscopelus macrolepHotus, Johnson, P. Z. S. 1863, p. 44, pi. 7 : Vaillant, Exp. Sci. Travaillenr et Talisman,
Poiss. p. 119 : Alcock, Ann. Hag. Nat. Hist., (6) VIII. 1891, p. 129 : Goode and Bean, Oceanic Ichthyology, p. 93,
figs. 108, 109.
Scopeliis macrolepidotus, Giinther, Cat. Fishes, V. p. 414, and Challenger Deep-Sea Fishes, p. 196.
B. 9. D. 13. A. 13. P. 15-16. Y. 8. L. lat. 30.
Length of the head about a third, height of the body about two-ninths the
total length without the caudal. Snout broad, somewhat depressed, decidedly
longer than the eye, its tip formed by the prominent mandible. Nostrils almost
superior. Eye situated almost midway between the tip of the snout and the
vertical limb of the preoperculum, its length about a fifth that of the head and
decidedly less than the width of the interorbital space.
165
Mouth-cleft very oblique ; the maxilla, which is dilated behind, only reaches
to the posterior border of the orbit. Villiform teeth are present on the vomer.
The first dorsal ray is almost midway between the tip of the snout and the
adipose fin : the anal begins half a head-length behind the last dorsal ray. The
pectorals are as long as the head without the snout, and when unbroken, reach
almost to the vent and as far as, or beyond, the tips of the ventrals.
The posterior margin of each scale is covered with minute spines.
Under surface with numerous very regular longitudinal series of large
luminous organs : beginning with two series at the tip of the isthmus, becoming
7 or 8 series on the throat, 5 or 6 series on the belly, 2 series on either side of
the anal fin, and 3 series — of which that in the middle line is very small — between
the anal and the caudal.
A large air-bladder is present.
Mature females are nearly 7^ inches long, mature males are somewhat
smaller.
Andaman Sea, 188 to 220 and 405 fathoms : Arabian Sea, off Travancore
coast, 360 fathoms.
Eegd. Nos. 13124, 13125, if7, 1-f, 3J2.
Distribution: West Indies; Madeira and Morocco coast; Arabian and
Andaman Seas ; off Kermadec Is.
In the Indian Museum is also one of the " Challenger " duplicates.
SCOPELENGTS, Alcock.
Scopelengys, Aloock, Ann. Mag. Nat. Hist., October, 1890, p. 303.
Head and body compressed. Eye small. Mouth very wide ; the maxilla
dilated behind. Acute villiform teeth, in bands uncovered by the lips in the jaws,
and in the palatines and vomer. Gill-openings very wide ; gill-covers complete.
Pseudobranchige rudimentary. Dorsal fin near the middle of the body, short;
an adipose dorsal. Anal fin short. Caudal forked. Pectorals well developed.
Ventrals with eight rays. [Scales, if present, very deciduous.] No air-bladder.
Pyloric caeca present in moderate number.
"When I described Scopelengys I did not know Neoscopelus by autopsy. I
now feel sure, though the specimen is in a very bad state of preservation, that it
is very closely related to Neoscopelus.
Apart from any differences that may exist in the scales, it differs from
Neoscopelus in having a smaller eye, rudimentary pseudobranchiae, and no air-
bladder.
166
134. Scopelengys tristis, Alcock.
Scopelengys tristis, Alcock, Ann. Mag. Nat. Hist., October, 1890, p. 303 : Illustrations of the Zoology op
the Investigator, Fishes, pl. VII. fig. 7.
B. 8. D. 12. A. 13. P. 15. V. 8.
Head and body rather elongate, compressed. Eye situated high up, very
small ; its major diameter is a little more than - the length of the snout, which
is about j the length of the head, which is not quite 3 the total without the
caudal. Mouth wide, its cleft very oblique, approaching the vertical, with the
lower jaw projecting in repose ; the maxilla, which is widely dilated behind,
measures more than half the length of the head ; the premaxilla is a stout bone,
firmly attached to the maxilla, which it equals in length. Acute villiform teeth,
in rather broad bands uncovered by the lips in the premaxilla? and mandible,
in narrow bands in the palatines, and in a small patch on each side of the head
of the vomer ; no teeth on the tongue.
Gill-openings very wide ; gill-covers complete ; long close-set gill-rakers on
the first arch. Pseudobranchige rudimentary, consisting of three or four small
lamellae on each side.
The dorsal fin begins above the origin of the ventrals ; the whole fin is
included in the anterior half of the body measured with the caudal. Adipose
dorsal rather large, fimbriated. The anal fin begins a little more than a snout-
length behind the .posterior limit of the dorsal. Caudal forked. Pectorals
entire, about as long as the maxilla, and reaching just beyond the origin of the
ventrals ; they arise close to the ventral profile.
Eight large pyloric caeca. No air-bladder.
Colour in the fresh state apparently uniform black throughout.
One specimen, 6- inches in length, nearly mature.
Arabian Sea, off the Laccadive Is., 1000 fathoms.
Odontostomus, Cocco.
Odontostomiis, Cooco, Lett su Alcun. Salmon , p. 32 : Giinther, Cat. Fishes, V. p. 417, and Challenger Deep-
Sea Fishes, p. 200: Ooode and bean, Oceanic Ichthyology, p. 121.
Head compressed; body compressed, moderately elongate, naked. Snout
short. Cleft of the mouth very wide : premaxilla and maxilla slender, the
former with a series of small teeth of equal size : the lower jaw, the vomer and
the palatines with a few deprcssible fangs of enormous size. Eye large; the
orbit of great vertical depth and with a broad transparent membranous lateral
fold or wall.
Gill-openings wide: no gill-rakers : eight branchiostegals. PseudobrancLme
avoII developed.
167
Dorsal fin short, in the middle or anterior half of the body : anal fin long,
in the posterior half of the body. Caudal forked. Adipose fin small, placed far
back.
Pectorals and ventrals well developed, the pectorals inserted near the
ventral profile, the ventrals inserted below the dorsal some way behind the
pectorals.
Distribution : Mediterranean, Bay of Bengal, Andaman Sea.
135. Odontostomus atrattis, Alcock.
Odontostomus atratus, Alcock, Journ. As. Soc. Bengal, Vol. LXII. pt. 2, 1893, p. 182, pi. ix. fig. 4 : Illustrations
of the Zoology of the Investigator, Fishes, pl. XXXIII. fig. 3.
B. 8. D. 11. A. 26. P. 12. V. 8.
The extreme length of the square, high, compressed head is a little more,
and the greatest height of the compressed tapering body is a little less, than one-
fourth of the total, caudal included.
The snout has the form of a pointed wart beyond which the upper jaw
slightly projects, the lower jaw again projecting a little beyond the upper.
The eyes, which are situated about a diameter apart, near the top of the
head, have their major diameter obliquely vertical, and are capable of such
strong rotation inwards as to bring the visual axis obliquely upwards, the orbit
being walled in laterally by a stout but transparent fold of skin in its lower half.
The cleft of the mouth extends almost to the posterior edge of the preoper-
culum : the premaxillge are armed with a series of close uniform serrations for
the most part pointing forwards, the vomer bears on each side a sabre-shaped
depressible fang nearly half as long as the head, the palatines have each an
exactly similar fang succeeded by a row of close serrations, and the mandible
has on each side a distant series of similar fangs of unequal size, the largest of
them however being hardly half the length of those on the vomer and palatines.
Gill-cleft extremely wide and high : four gills with wide lamina3 and gill-
rakers inconspicuous or absent : pseudobranckias large.
Body covered with a glandular scaleless skin in which the lateral line
appears in spirit as a white streak. Rows of white dots (luminous organs ?)
exist along the free border of the preoperculum and the inner border of the
broad boat-shaped mandible.
The dorsal fin lies altogether within the anterior half of the body : the anal
begins about half a head length behind the vertical through the last dorsal ray,
and extends to the rudimentary basal rays of the forked caudal. The large
pectorals arise close to the ventral profile, almost in the same plane with the
168
ventrals, the bases of which they touch when laid back. The ventrals arise
under the middle of the dorsal.
Length of the adult 3^ inches.
Bay of Bengal, 573 fathoms ; Andaman Sea, 370 to 419 fathoms.
Regd. Nos. 13434, 5Z5.
Family Alepocephalidce.
Head almost always naked. Body usually covered with thin cycloid
scales, but sometimes naked. Barbels none. Margin of the upper jaw
formed by the premaxillse and maxillae, the former being placed along the
upper anterior edge of the latter: the maxilla commonly broad and
resembling that of the Clupeidce. Gill-openings very wide: opercles
complete. Pseudobranchiae present.
No adipose fin : the dorsal fin belongs to the caudal portion of the
back-bone.
Pyloric caeca few. No air-bladder.
This family is represented in the depths of the Indian Seas by 12 species
belonging to 7 genera.
Synopsis of the Indian genera of the family Alepocephalidae.
I. Body scaly : —
1. Suout normal : body rather elongate : ventrals present : —
i. Six branchiostegals : a single series of teeth on the pre-
maxilla?, none on the maxilla? ... ... ... Alepocephalus.
ii. Seven branchiostegals : a single series of teeth on the pre-
maxilla? and maxilla? ... ... ... ... Bathytroctes.
iii. Seven branchiostegals : several series of teeth on the pre-
maxillse, a single series on the maxilla? ... ... Narcetes.
2. Snout normal : body short and elevated : no ventrals ... Platttroctes.
3. Anterior bones of the head produced to form a tubular snout at
the end of which the mouth is situated : body rather elongate ... Aolastomomorpha.
II. Body naked: —
1. Body moderately elongate: dorsal and anal fins of moderate
length ... ... ... ... ... Xenodermichthts.
2. Body exceedingly elongate : dorsal and anal fins very long ... Leptoderma.
Alepocephalus, Risso.
Alepocephalus, Uisso, Mem. Ac. Nat. Sci. Turin, XXV. 1820, p. 270: Muller, Abh. Ak. Wise. Berlin, 184R,
p. 171 : Cuvier and Valenciennes, Hist. Nat. Poiss. XIX., p. 169 : Gunther, Cat. Fishes, VII., p. 477 : Goode and
Bean, Oceanic Ichthyology, p. 86.
Alepocephalus and Mitchillina, Jordan and Evermann, Fishes N. Araer., pp. 452, 453.
Body moderately elongate, compressed, covered with thin cycloid scales.
Head nuked. Cleft of the mouth of moderate width, Avith the jaws nearly even
169
in front. A series of small teeth in the premaxilla? mandibles and palatines, and
sometimes on the vomer. Eye large.
Gill-openings wide, the opercles large and thin. Six branchiostegals. Gill-
rakers numerous and rather long.
Dorsal and anal fins nearly equal and opposite, placed far back, in the
posterior half of the body. Caudal forked. Pectorals and ventrals well deve-
loped, but rather small.
Distribution : Atlantic ; Mediterranean ; Arabian Sea and Bay of Bengal ;
Western South Pacific.
Key to the Indian species of Alepocephalus.
I. The maxilla reaches a little beyond the anterior border of the orbit, and the
snout is more than a fourth the length of the head : tlie anal fin begins well
behind the middle of the body (measured with the caudal) : —
1. Eyes of adult more than a diameter apart, and between a fifth and a
sixth the length of the head : the gill-membranes overlap each other
broadly: 9 pyloric cseca ... ... ... ... A.bicolor.
2. Eyes of adult less than a diameter apart, and between a third and a
fourth the length of the head : the gill-membranes overlap slightly: 14
pyloric casca ... ... ... ... ... A. Blanfordi.
II. The maxilla reaches nearly to the posterior border of the orbit, and the snout
is less than a fourth the length of the head : the anal fin begins exactly in
the middle of the body : 4 pyloric cseca ... ... ... ... A, edentulus.
136. Alepocephalus bicolor, Alcock.
Alepocephalus bicolor, Alcock, Ann. Mag. Nat. Hist., August, 1891, p. 133 : Illustrations of the Zoology of
the Investigator, Fishes, pl. IV. fig. 2.
B. 6. D. 21. A. 28. P. 10. V. 8. L. lat. 62. L. tr. 18.
Length of the head slightly more than a fourth, height of the body nearly
a fifth the total without the caudal.
The length of the obtusely- pointed depressed snout is contained about
3.- times in that of the head. The eyes, which converge anteriorly, are between
one-fifth and one-sixth of the head-length in diameter, and are more than their
own diameter apart. The large nostrils are situated close together immediately
in front of the eye.
Mouth-cleft slightly oblique; the maxilla reaches just behind the vertical
through the anterior border of the orbit. A row of small teeth in each jaw and
on the palatines.
Gill-openings very wide, the membranes entirely separate and overlapping
broadly ; a great part of the gill-cover is formed by the broad flat branchiostegal
rays, which are uncovered by the opercle from their very bases ; the opercular
22
170
bones, which are extremely thin, are invested by the same tough black skin that
covers the head ; the gill-lamina? are coarse and the gill-rakers on all the arches
long and lamellar ; pseudobranchias small.
Head naked, body covered with large cycloid scales, which are deciduous
everywhere but on the lateral line ; small scales also invest the bases of all the
fins. A scale from the flank measures about 7*5 millim. in the horizontal and
about 5'5 millim. in the vertical diameter.
The dorsal and anal fins arise just in advance of the posterior third of the
body (measured without the caudal), and the base of the former, which begins a
little in advance of the latter, is two-thirds that of the latter in extent. Caudal
deeply forked, with very numerous rudimentary rays at its base. Pectorals
broad, in length a little more than the postorbital portion of the head. The
ventrals arise just abaft of midway between the pectorals and anal ; they are
broad and reach more than halfway to the anal.
Stomach small, siphonal. The intestine, which, when unravelled, is about 2-
times the entire length of the fish, consists of two portions, which both in
structure and arrangement are quite different from one another : the anterior
five-sixths is thin-walled and of small calibre, and is intricately coiled in a globu-
lar mass situated in the anterior fourth of the abdomen, the coils being held by
a long mesentery ; the posterior sixth is wide, but with walls so thick as to
almost block the lumen (in the contracted state), the mucosa in this condition
being thrown into numerous longitudinal folds ; it passes straight down the
middle of the abdominal cavity unsupported by mesentery. There are nine
large long pyloric casca in a pectinate arrangement.
In a female with much-enlarged ovaries containing ova nearly -i millim. in
diameter the ovaries extend back to the wide genital pore, through which they
open to the exterior.
Colours in life: — Head, including sclerotic and iris, black; body uniform
dull slate-blue ; pharyngo-brancbial mucous membrane and parietal peritoneum
black.
Adult females are just under a foot in length : adult males are a good deal
smaller.
Bay of Bengal, off Ganjam coast, 240 to 276 fathoms; Arabian Sea, off
Malabar coast, 360 fathoms.
Regd. Nos. 13079, 13080, 13081, 13083, ™.
The microscopic structure of the hind-gut of Alepocephalus bicolor.
In transverse section the appearance somewhat resembles that of the human
vas deferens. Externally there is a thin fibrous coat containing blood-vessels,
and internal to this and intimately adherent to it is a thin layer of longitudinally-
171
arranged muscular fibres. Inside this is a layer, averaging about balf a milli-
metre in thickness, of dense, circularly-arranged, muscular fibres. Internal to
this is a submucous layer thrown into numerous wide longitudinal folds, and
invested by a single row of long columnar epithelium, with numerous large
goblet-cells. The submucous coat in all the sections made is everywhere infil-
trated with round or oval, deeply-pigmented, highly granular corpuscles, which
measure from -r^- to -£- of an inch in diameter ; in shape they resemble large
leucocytes, but they are so granular that no nucleus can in any instance be
detected.
The thick muscular coat, the dense infiltration of the submucosa with these
pigmented granular corpuscles, and the large and numerous goblet-cells of the
mucosa characterize this part of the intestine.
137. Alepocephalus Blanfordi, Alcock.
Altpocephalu* BJanfordi, Alcock, Ann. Mag. Nat. Hist., Nov. 1892, p. 357: Illustrations of the Zoology or
the Investigator, Fishes, pl. IX. tig. 1. (reduced.)
B. 6. D. 16. A. 17. P. 11. V. 6-7. L. lat. circ. 70.
Length of head one-third, height of body two-elevenths, of the total
without the caudal.
The length of the obtusely-pointed depressed snout is barely greater than
the diameter of the huge orbit, or two-sevenths of the length of the head.
The eyes are hardly half a diameter apart, with the large nostrils placed
close together in front of their angle.
The mouth-cleft is almost horizontal, and the upper jaw, which reaches just
beyond and rests upon the anterior border of the orbit, completely encloses the
mandible on all sides ; a row of fine teeth in each jaw and on each prominent
palatine.
Gill-openings very wide, the gill-membranes entirely separate and only
slightly overlapping; the branchiostegal rays are but little concealed by the
opercular bones, and the whole gill-cover is clothed by a continuation of the thick
scaleless skin that covers the head; gill-rakers numerous, close-set, broadly
lanceolate, acute ; pseudobranchia? large and coarse.
Body covered with thick deciduous cycloid scales ; a scale from the abdomen
is nearly 5 5 millim. in the horizontal and 5 millim. in the vertical diameter.
The dorsal and anal fins, which are similar in form, equal in extent, and opposite,
lie well within the posterior third of the body (measured without the caudal) ;
the caudal is deeply forked, with many rudimentary rays at its base. The ventrals
arise almost in the middle of the body, nearer to the anal than to the pectorals.
172
Stomach siphonal ; a row of fourteen very large and long pyloric cseca em-
braces its pyloric moiety ; the intestine, which when unravelled is about twice
the entire length of the fish, is arranged as in Alepocephalus bicolor, but the wall
of the coiled up small intestine is much thicker, and the straight hinder gut is
held by a stout mesentery.
Colour : head and fins black ; body lavender-grey.
Length 14" inches.
Arabian Sea, off Cape Comorin, 902 fathoms.
Regd. No. 13191.
The microscopic structure of the hind-gut, etc., of Alepocephalus Blanfordi.
The straight large gut in this species, as in Alepocephalus bicolor, is remark-
able for the great thickness of its wall and for its contracted lumen ; only in the
present case, although the circular muscular coat is conspicuously thick, it is not
this but the highly glandular mucous coat that contributes most to the thickness
of the wall. The great development of the glands of the mucosa, which are
compact little branching follicles, is in marked contrast to A. bicolor, where the
mucous membrane consists of simple columnar epithelium. The loose submucous
coat is honeycombed with (lymphatic ?) channels and crowded with leucocytes ;
but the large pigmented granular corpuscles which were so numerous in
.1. bicolor are here few in number.
The small intestine at its duodenal end, and the pyloric ca?ca, appear, in
transverse sections, to be identical in structure. In both the mucous membrane
is thrown into apparently permanent longitudinal folds, and contains in its depth
a regular series of racemose glands opening to the surface by a longish duct.
Microscopic cylinders of glandular substance, which in stained sections has
exactly the appearance of mammalian pancreas, run in the mesentery, parallel
with the pyloric caeca and in contact with them.
138. Alepocephalus edentulus, Alcock.
Alepocephalus edentulus, Alcock, Ann. Mag. Nat. Hist., Nov. 1892, p. 358, pi. xviii. fig. 2: Illustrations of
THE Zoology of the Investigator, Fishes, pl. XXXII. fig. 4.
B. 6. D. 29. A. 35. V. G. P. 9. L. lat. circa 50. L. tr. 15.
The length of the head is a little more than one-fourth, and the height of
the much compressed body nearly one-fifth, of the total with the caudal included.
The blunt snout is barely equal in length either to the width of the interorbital
space or to the diameter of the eye, which is very nearly two-ninths the length
of the head. The mouth-cleft is almost horizontal, the jaws are even anteriorly,
and the maxilla reaches considerably behind the vertical through the centre of
the eye. Minute teeth occur in a row in the premaxillse and mandibles, and
173
there are a few inconspicuous and deciduous teeth on the prominent edges of the
palatines only.
Gill-openings very wide, the gill-membranes being attached to the isthmus
only quite anteriorly ; gill-rakers conspicuous on all the branchial arches, and, to
the number of about twelve in the middle of the first arch, long and setaceous ;
pseudobranchise small. Head covered with a velvety scaleless skin ; body with
scales that are so deciduous as to have entirely disappeared, leaving only
imprints.
The long anal fin begins an eye-length behind the middle of the body,
measured without the caudal, and the shorter dorsal arises in the vertical through
the sixth or seventh anal ray ; the caudal is completely divided down to its base
into two long feathery lobes. The small ventrals, which arise midway between
the base of the pectoral and the origin of the anal, reach rather more than half-
way to the latter point.
Stomach siphonal ; a row of four stout pyloric caeca ; intestine slightly
coiled, with its terminal end enlarged and thick-walled.
Colours : head and eyes jet-black ; body and fins greyish black.
A mature male is nearly 7 inches long.
Bay of Bengal, off Madras coast, 475 fathoms.
Re^d. No. 13192.
■'6'-
Bathytroctes, Giinther.
Bathytroctes, Giinther, Ann. Mag. Nat. Hist., 1878, Vol. II.. p. 249 ; and Challenger Deep Sea Fishes, p. 225 :
Goode and Bean, Oceanic Ichthyology, p. 40 : Jordan and Evermann, Fishes of N. Amer., p. 454.
Talismania, Goode and Bean, Oceanic Ichthyology, p. 41 : Jordan and Evermann, Fishes of N. Amer., p. 455.
Differs from Alepocephalus only in the following particulars : —
The maxilla, as well as the premaxilla, has a series of small teeth. The
opercles and part of the cheeks are sometimes, but not commonly, scaly. There
are seven branchiostegals. The anal fin commonly begins below the posterior
part of the dorsal.
Distribution : Atlantic ; Arabian and Andaman Seas ; "Western Pacific.
139. Bathytroctes squamosus, Alcock.
Bathytroctes squamosus, Alcock, Ann. Mag. Nat. Hist , Oct. 1890, p. 304 : Illustrations of the Zoology of
the Investigator, Fishes, pl. V. fig. 1.
B. 7. D. 17. A. 17. C. arc. 35. P. 10. V. 9. L. lat. circ. 50. L. tr. 15.
Length of the head a little more than the greatest height of the body and a
little more than a fourth of the total.
174
Snout much shorter than the eye, which is a third the length of the head
and almost enters the dorsal profile. Nostrils large, situated immediately in
front of the eye.
Mouth-cleft wide, approaching the transverse ; premaxilla short and slender ;
the broad maxilla, composed of three longitudinal plates, of which the innermost
(uppermost) is movable, reaches just behind the level of the mid-orbit, and in-
cludes the mandible in repose, except anteriorly, where the latter strongly pro-
jects. Small, even, acute, uniserial teeth, recurved in the premaxillaa, mandible,
palatines, and vomer, procurrent or procurved in the maxillse. Tongue large.
A row of pores along the limb of the mandible.
Gill-openings very wide, the membranes entirely separate ; fourth gill-cleft
occluded ; gill-rakers long and close-set on the first three arches, longest on the
first. Pseudobranchiae large and coarse.
Scales large, deciduous, except on the lateral line where they are adherent
and perforated. There are scales on the cheeks and opercles.
The dorsal fin begins just behind the origin of the ventrals, which are situat-
ed in the vertical through the middle of the body measured without the caudal.
The anal begins in the vertical through the third dorsal ray. Both these fins
have fleshy succulent bases, and the rays increasing in length regularly and
steeply to the fourth, and then decreasing as regularly but more gradually to the
last. Caudal symmetrically forked. Pectorals long and narrow ; their longest
rays equal the length of the head behind the anterior nostril, and in repose almost
touch the bases of the ventrals. Ventrals broad, reaching slightly beyond the
vent.
Stomach large ; intestine coiled in a spiral ; five or six large pyloric caeca.
Colours in the fresh state : — Head uniform deep black, body pinkish brown,
fins transparent grey ; buccal membrane and entire peritoneum black.
A female specimen, 10j inches long, with gravid ovaries, the mature ova
measuring - of an inch in diameter.
Arabian Sea, off the Lacadives, 740 fathoms.
Regd. No. 12869.
Besides this species the remains of two others from the Andaman Sea are in
the collection. One of them may be B. macrolepis, Gthr., the other may be
IS. microlepis, Gthr.
Narcetes, Alcock.
Nareeiee, Alcock, Ana. Mag. Nat. Hist., Oct. 1890, p. 305.
Differs from Alepocephalus only in the following particulars : —
There are teeth on the maxilla?, as well as on the premaxilla?. There are
seven branchiostegals. The anal fin is entirely behind the dorsal.
175
Differs from Bathytrodes in that the teeth of the premaxillae and mandibles
are in several series.
Head naked. Body rather elongate, compressed, covered with scales of
moderate size. Eye rather small. Mouth wide ; the maxilla extending beyond
the vertical through the middle of the orbit. Fine teeth in premaxillae, maxillae,
mandible, palatines, and vomer, those in the premaxillas and mandible pluriserial ;
no teeth on the tongue.
Gill-openings wide ; gill-covers complete ; seven branchiostegals ; four gills,
with narrow lamina?; gill-rakers long. Pseudobranchia? present. The dorsal
arises in the posterior half of the body and the anal is entirely behind it. No
adipose dorsal fin. Caudal forked. Pyloric cseca in moderate number. Ovaries
with an oviduct.
140. Narcetes erimelas, Alcock.
Narcetes erimelas, Aloock, Ann. Mug. Nat. Hist., Oct. 1890, p. 305: Illustrations of the Zoologt of the
Investigator, Fishes, pl. IV. fig. 1.
B. 7. D. 15-16. A. 12. C. circ. 35. P. 10-11. V. 9. L. lat. 68.
Head broad, pyramidal, its length 3- to 3^ in the total without the caudal ;
body elongate, its greatest height, just behind the gill-oj)ening, about 5^ in the
same standard.
Head-bones sculptured, specially the operculum and preoperculum, both of
which have their border augmented by a semimembranous striated fringe.
Snout nearly as broad as long, depressed, rounded from side to side, its
dorsal and ventral profiles meeting at an acute angle; its length is a little
over g that of the head, and more than half as long again as the eye. Nostrils
very large.
Eye rather small, its major diameter 5- in the head-length, and not quite
equal to the width of the deeply concave interorbital space.
Mouth wide, oblique ; the maxilla reaches well behind the vertical through
the posterior border of the orbit. The premaxilla is a short strong bone ; the
maxilla is composed of three longitudinal plates, of which the innermost (upper-
most) is movable ; the mandible is very strong and broad, and its under surface
is excavated for a wide mucous channel which opens by six large circular pores
on each side.
Teeth small, even, uniform, acute ; those in the jaws standing, uncovered
by the lips, outside the mouth ; those in the premaxilla? and mandible recurved,
quadriserial anteriorly and triserial laterally in the premaxillaa, biserial in the
mandible ; those in the maxillaa uniserial, procurrent or procurved ; those in the
176
palatines uniserial, incurved ; those in the vomer recurved, in a group of two or
three on each side. Tongue large, toothless.
Gill-openings very wide ; gill-membranes entirely separate ; gill-covers large,
complete ; gill-rakers on the first arch close-set, finely pointed, and as long as
the eye ; fourth gill-cleft rather wide ; gill-laminas very narrow, the individual
lamella3 extremely delicate. Pseudobranchiae large.
Head naked ; body covered with deciduous scales of moderate size. The
lateral line runs straight along the middle of the body.
The dorsal fin begins almost in the vertical through the origin of the vent-
rals, which are situated a snout-length behind the middle of the body measured
without the caudal. The anal fin begins two rows of scales behind the last
dorsal ray. Pectorals and ventrals well developed, broad, fragile.
Stomach very large, with thick walls thrown into deep longitudinal folds.
Intestine coiled in a spiral ; ten very large pyloric casca in a bunch.
Colours in the fresh state : — head, iris, body, fins, inside of mouth and gill-
chamber, and entire peritoneum, deep black.
Two female specimens, measuring respectively 13- and 9^ inches.
Both, when brought on board, were in a curious state of cataleptic rigor.
Arabian Sea, near Laccadive Is., 740 fathoms.
Regd. Nos. 12870, 12871.
Platytroctes, Giinther.
Platytroctes, Giinther, Ann. Mag. Nat. Hist. 1S78, II. p. 249 : and Challenger Deep-Sea Fishes, p. 229 : Goode
and Bean, Oceanic Ichthyology, p. 45 : Jordan and Evermanu, Fishes N. Amer., p. 458.
Body rather abbreviated and elevated, much compressed, covered with small
scales many of which are keeled. Head naked. Mouth of moderate width : a
single series of small teeth in premaxilla, maxilla and mandible : a few (two in
the only known species) teeth on the vomer. Eye large.
Gill-opening wide : six branchiostegals. Gills very narrow : pseudobranchio?
present : gill-rakers long, lanceolate.
The dorsal and anal fins, which are in the posterior half of the body, are
equal, opposite, and of moderate length. Caudal forked. No adipose dorsal.
Pectorals small. Each clavicle ends below in a long freely projecting spine,
which is fused with its fellow except perhaps at tip.
No ventrals.
Pyloric appendages rudimentary.
Distribution : Atlantic ; Arabian Sea.
177
141. Platytroctes apus, Gimther.
Platytroctes apus, Giinther, Ann. Mag. Nat. Hiet. 1878, II. p. 249, and Challenger Deep-Sea Fishes, p. 229, pi.
lviii. fig. A: Alcock, Ann. Mag. Nat. Hist. (6) VI. 1890, p. 307: Goode and Bean, Oceanic Ichthyology, p. 46,
fig. 53 : Jordan and Evermann, Fishes of N. Amer., p. 458.
D. 18. A. 17. P. 20. L. lat. circ. 100.
Length of the head two-sevenths, greatest height of the body, at its middle,
rather more than a third the total length without the caudal. More than half
of this height, however, is contributed by simple dorsal and ventral folds of skin
into which neither muscles nor viscera enter.
The snout is shorter than the eye, which is a third or more the length of
the head and almost enters the dorsal profile. Nostrils large, superior, nearer
to the edge of the snout than to the eye.
The narrow triangular interorbital space and the occiput are sharply con-
cave, the concavity being bordered on each side by a mucous canal with large
pores. A similar mucous canal with pores runs along the preorbital, and another
one along the free edge of the preopercle.
Mouth rather short but broad, the lower jaw projecting when the mouth is
open. The maxilla, which is a broad petal-shaped bone, reaches to or a little
beyond the anterior edge of the eye. The limbs of the mandible make a curious
boat-shaped bone. A single series of small even teeth in the premaxilla and
maxilla and in the front half of the mandible : a small tooth on either side of the
head of the vomer.
Gill-laminse very short : gill-rakers on the first branchial arch long, ex-
tremely numerous and close-set.
The clavicles project freely at their symphysis as a pair of spikes separated
only at tip.
The vent is much nearer to the root of the caudal than to the gill-opening :
the dorsal fin begins immediately above it and the anal immediately behind
it.
Pectoral fin short, about half as long as the eye, its base nearly horizontal.
Scales small, cycloid : those near the dorsal and ventral profiles, and many
of the others, have a keel, like the scales of many snakes.
Colours in spirit : brown ; head, pectoral region, vent, and edges of caudal
peduncle black. Length six inches.
Arabian Sea, in the neighbourhood of the Laccadive banks, 740 fathoms.
Regd. No. 12868.
Distribution : Mid- Atlantic ; Arabian Sea.
23
178
AULASTOHOMOEPHA, AlcOck.
Aulastomatomorpha, Alcock, Ann. Mag. Nat. Hist., Oct. 1890, p. 307.
Head naked. Body elongate, covered with minute hardly imbricate scales.
Anterior bones of the head produced into a long tube terminating in a narrow
mouth. Margin of the upper jaw formed equally by the premaxilla? and maxilla?.
Uniserial teeth, in the jaws only. Eye large. Gill-cover apparently complete.
Gill-opening wide below, contracted above, where it does not surpass the level
of the pectoral fin ; four gills with narrow lamina?. Pseudobranchia? almost
rudimentary. Dorsal fin short, quite in the posterior part of the body. Pecto-
rals and ventrals well developed. Anal fin very long. Caudal forked. Pyloric
caeca few, small.
142. Aulastomomorpha phosphorops, Alcock.
Aulastomatomorpha phosphorops, Alcock, Ann. Mag. Nat. Hist., Oct. 1890, p. 307: Illustrations of the
Zoology of the Investigator, Fishes, Plate V. Fig. 2.
B. 5? D. 21. A. 41. P. 7. V. 6.
Body elongate and compressed, surrounded from the mid-dorsal line behind
the nape to the mid-ventral line behind the vent by a continuous thick succulent
fold of the integuments, like, but not so wide as, that of Platytroctes ; its greatest
height, including this fold, is a little more than ^ of the total without the caudal.
Head completely covered with a thick, spongy, dazzling white, probably lumin-
ous, skin.
Head low and rather depressed, its length 3g in the total without the
caudal ; produced anteriorly into a long tubular snout, at the end of which is
the small mouth.
The snout is a little less than half the length of the head, or 6"- in the total
without the caudal.
The eyes are very large and extremely prominent ; the major diameter of
the eye-ball is slightly over - the head-length, but owing to the encroachment
up to the margin of the cornea of the broad posterior orbital fold, the diameter
of the exposed " eye " is only a little more than i of the same standard ; the true
(bony) interorbital space is less than half the diameter of the eye in width.
Nostrils situated high up, above the anterior orbital angle. Mouth at the
extreme end of the tubular snout, small, the jaws apparently with limited
motion. The upper jaw, which projects slightly beyond the lower, is formed in
its anterior half by the premaxilla, in its posterior half by the maxilla. Minute,
acute, recurved teeth in a single row in the premaxilla? and mandible ; no teeth
in the maxilla?.
30
179
Gill-openings very wide below, contracted above, where they do not surpass
the level of the pectorals. Gill-covers apparently complete ; their constituent
bones, including the branchiostegal rays, though well calcified, are extremely
thin and fragile, and are completely concealed within a continuous fold of skin
and mucous membrane. Four gills, with narrow laminae and coarse lamella? ;
the fourth gill-cleft wide ; gill-rakers well developed on all the arches, moder-
ately long on the first, short on the fourth and fifth. Pseudobranckia? rudi-
mentary, consisting of four or five delicate short lamella? on each side.
Body covered with minute, hardly imbricate, cycloid scales, about — by ^
of an inch respectively in the shortest and longest diameters. The lateral line
traverses the middle of the body uninterruptedly.
The dorsal fin begins slightly in advance of the posterior fourth of the body
measured without the caudal ; the length of its base is shorter than the snout ;
its rays, like those of the anal, increase gradually in length from before back-
wards, the longest being not quite equal to the major diameter of the eye-ball.
The anal begins an eye-length behind the middle of the body as above limited,
and ends a short distance behind the last dorsal ray ; its longest rays slightly
exceed the longest dorsal rays. Caudal symmetrically forked, its rudimentary
rays very numerous, both dorsally and ventrally. Pectorals narrow, rather
more than ^ of the head in length. Ventrals short, arising immediately behind
the middle of the body, as above limited, and reaching just behind the vent.
Stomach subsiphonal ; intestine long, coiled in a spiral ; four small pyloric
caeca, arranged in a ring. Reproductive glands very large, apparently discharg-
ing in the male through a well-developed post-anal papilla.
Colours in the fresh state : — head snow-white, iris black, body chocolate,
fins blackish grey ; mouth, gill-chamber and entire peritoneum intense black.
One specimen, apparently a male near maturity, measuring 11 inches in
length.
Arabian Sea, near the Laccadives, 1000 fathoms.
Regd. No. 12872.
Xenodermichthts, Giinther.
Xenodermichthys, Gunther, Ann. Mag. Nat. Hist., 1878, II., p. 250, and Challenger Deep-Sea Fishes, p. 230:
Goode and Bean, Oceanic Ichthyology, p. 46.
Body low, rather elongate, compressed, without true scales, but with
numerous tiny more or less regularly arranged nodules which are probably
luminous in function, and often also with scattered rudimentary scales. Mouth
small, or moderate, with feeble jaws and small teeth in the premaxilla, maxilla,
and mandible. Palate toothless.
Gill-opening wide. Pseudobranchiae present. Gill-rakers long and numerous.
180
Dorsal and anal fins equal and opposite, of moderate length, placed far back
in the posterior half of the body. Caudal forked. Pectorals and ventrals well
developed, but rather small.
Pyloric casca present.
Distribution : European and African side of the Atlantic ; Arabian Sea, Bay
of Bengal, Andaman Sea ; Japanese Seas.
Key to the Indian species of Xenodermichthys.
I. The lateral line is indistinct ... ... ... ... X, Giintheri.
II. The lateral line is a salient tube supported by regularly arranged
subcutaneous scales ... ... ... ... ... X. squamilaterus.
143. Xenodermichthys Guntheri, Alcock.
Xenodermichthys Giintheri, Alcock, Ann. Mag. Nat. Hist., Nov. 1892, p. 359, pi. xviii. fig. 3 : Illustrations of
the Zoology op the Investigator, Fishes, pl. XXXII. fig. 2.
B. 6. D. 15. A. 14. V. 6. P. 5?
Body elongate, compressed, covered with a thick, scaleless, longitudinally -
wrinkled, black skin, in which scattered granular yellowish-coloured nodules are
imbedded. The dorsal and anal profiles are symmetrically similar in life. The
length of the head is slightly over two-sevenths, and the height of the body
immediately behind the gill-opening about one-sixth of the total without the
caudal.
The obtuse snout, surmounted by an acutely-pointed tubercle which projects
from the prominent symphysis of the lower jaw, is not quite equal in length to
the diameter of the circular eye. The eyes, which in life encroach upon the
dorsal profile, are about two-sevenths of the length of the head, and are about
two-thirds of a diameter apart.
The mouth-cleft is oblique, and the jaws are even in front, except for the
symphysial tubercle on the mandible. The premaxillse, which form on each side
nearly one half the extent of the margin of the upper jaw, are armed with a row
of minute close-set teeth, as are also the maxilla? (which have the typical
Alepocephaloid structure and reach to the vertical through the posterior border
of the orbit) and the broad boat-shaped mandible ; no teeth on the palatines or
vomer.
The gill-cleft is extremely wide, extending forwards almost to the mandibu-
lar symphysis and upwards almost to the post-temporal region; the opercle
appears to be perfect, and, together with the branchiostegal rays, is enveloped
in a thick membranous skin, as in Alepocephalus ; four gills, with numerous long
close-set gill-rakers on the first arch ; pseudobranchise present.
No lateral line can be distinguished in life.
181
The dorsal and anal fins, which are equal, opposite, and similar, lie in the
posterior third of the body, and approach within an eye-length of the long series
of rudimentary rays that form the base of the deep-forked caudal. The ventrals
lie well within the posterior half of the body, and the pectorals arise on the
ventral profile, almost in the same horizontal line with the ventrals.
The stomach is siphonal and its pyloric end is embraced by a row of seven
or eight csecal appendages, the posterior six of which are relatively enormous ;
the intestine has an anterior much-coiled portion and a hinder portion which
passes perfectly straight backwards, much as in Alepocephalus bicolor and
A. Blanfordii, to its orifice just in advance of the posterior third of the body.
Colour uniform jet-black.
A mature female is nearly 6 inches long.
Bay of Bengal, off Madras coast, 678 fathoms ; Arabian Sea, off Travancore
coast, 430 fathoms.
Regd. Nos. 13193, ^, H2.
This species chiefly differs from X. socialis Vaillant, with specimens of which
I have compared it, in having a longer and sharper snout, and much fewer dorsal
and anal fin-rays.
I have described the ova of this fish in the Annals and Magazine of Natural
History for November 1892. The largest ova are between 2 and 3 millim. in
diameter.
144. Xenodermichthys squamilaterus, Alcock.
Xenodermichthys squamilaterus, Alcock, Ann. Mas;. Nat. Hist., Ang. 1898, p. 148 : Illustrations of the
ZOOLOGT OF THE INVESTIGATOR, FlSHES, PL. XXV. FIG. 4.
B. 6. D. 20. A. 18. P. 6. V. 6.
Distinguished from X. Giintheri, which it very closely resembles, by the
following characters : —
1. The lateral line is a salient tube which runs straight down the middle
of the body and is stiffened by thin subcutaneous equidistant scales : between
every two scales there is a pore.
2. The snout is shorter and blunter, and the eye is rather smaller, its dia-
meter being one-fourth the length of the head.
Length 6 inches.
Colour : uniform jet black.
Andaman Sea, 370 to 419 fathoms.
Regd. No. --.
182
This species is distinguished from X. socialis by the prominent scaly lateral
line and by the much fewer rays in the dorsal and anal fins.
Leptoderma, Vaillant.
Leptoderma, Vaillant, Exp. Sci. Travaillenr et Talisman, Poiss. p. 165 : Goode and Bean, Oceanic Ichthyology,
p. 18.
Body low, very elongate, tail tapering almost to a filament ; skin naked.
Head moderate, with enormous eyes. Cleft of the mouth small, the edge of the
upper jaw formed nearly equally by the premaxilla and maxilla. A series of
small teeth in both jaws, none on the palate.
Gill-opening wide but not reaching much above the level of the pectorals,
the upper arc of the gill-arches also truncated. Numerous close-set lanceolate
gill-rakers. Pseudobranchia? present, small.
Dorsal and anal very long, ending near the caudal, the anal the longer.
Caudal very small, forked. Pectorals and ventrals well developed.
Distribution : Atlantic coast of Morocco ; Bay of Bengal.
145. Leptoderma affinis, n. sp.
Leptoderma macrops, Alcock, Ann. Mag. Nat. Hist. (6) X. 1892, p. 361, (an Vaillant ?) : Illustrations of the
Zoology of the Investigator, Fishes, pl. XXXII. fig. 3.
D. circ. 66. A. circ. 85. V. 5.
Greatest height of the body, at the shoulder, about half the length of the
head, which is about two-ninths of the total.
Eye-ball considerably more than a third the length of the head and nearly
twice as long as the snout.
The snout is squarish ; the mouth is terminal, much as in Aulastomomorpha,
the maxilla being vertical when the mouth is opened in a perfectly natural
manner. The maxilla is very broad and consists of three pieces. The rami of
the mandible are also of great breadth, except anteriorly where there is a series
of small teeth. A series of small teeth on the premaxilla ; none on the maxilla
or palate.
The anal begins an eye-length nearer to the snout than to the base of the
caudal fin ; the dorsal begins about half a head-length behind the first anal ray :
both fins extend nearly to the caudal. The distance of the ventrals from the
<rill-opcning is equal to the length of the head without the snout.
The skin is naked and intensely black. In life it is uniformly covered with
a thick velvety opalescent epidermis which is probably luminous in function.
The lateral line, which consists of a row of pores, extends from the occiput to
the caudal.
183
In spirit the colour is purple, the contracted opaline epidermis forming a
sort of "bloom."
Length 8- inches.
Bay of Bengal, off Kistna coast, 753 fathoms.
Regd. No. 13197.
This species seems to differ from Leptoderma macrops, Vaillant, in having
the body less elongate, the lateral line very distinct, and the rays of the dorsal
and anal fins more numerous. It agrees fairly well with the figure but not with
the description of that species, and is probably identical with it.
Family Halosauridce.
Body elongate, tapering, covered with cycloid scales : head either
scaly or almost naked. Margin of the upper jaw formed by the premax-
illse mesially and the maxillae laterally. Opercles incomplete. The short
dorsal belongs to the abdominal portion of the backbone. No adipose
dorsal. Anal exceedingly long. Stomach with a caecal sack. Pyloric
appendages in moderate number. Gill-openings wide. No pseudobran-
chiae. Air-bladder large, simple.
Halosaueus, Johnson.
Halosaurus, Johnson, Proc. Zool. Soc. 1863, p. 406 : Giinther, Cat. Fishes, VII. p. 482, and Challenger Deep-
Sea Fishes, p. 232.
Halosaurichthys, Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 454.
Halosaurus, Aldrovandia and Halosaurichthys, Goode and Bean, Oceanic Ichthyology, pp. 129, 132, 136.
Halosaurus and Aldrovandia, Jordan and Bvermann, Fishes of N. Amer., pp. 607, 608.
Body elongate, compressed ; terminating in an exceedingly long, tapering,
lash-like tail ; abdomen rounded. Snout much projecting beyond the mouth,
which is inferior and of moderate width. Facial bones with large muciferous
channels ; a series of luminous organs generally developed in connexion with
the mucous canal system.
Suboperculum large, preoperculum rudimentary, interoperculum mem-
branous. Gill-openings wide, gill-membranes entirely separate. Branchiostegals
numerous. No pseudobranchia?.
Eye large or moderate. Villiform teeth in compact bands in the jaws,
rudimentary palatines, and pterygoids : none on the vomer. A band of hyoid
teeth. Dorsal fin short, above the space between the ventrals and the vent.
Anal fin exceedingly long, occupying the entire length of the tail. No caudal fin.
Lateral line running near the ventral profile, composed of scales which are
usually enlarged and bear the luminous organs.
Distribution : N. & S. Atlantic, Indian, and Pacific Oceans.
184
Key to the Indian species of Halosaurus.
I. Ventral fins not united together : —
1. Head naked, except for a few scales on the temples and upper part
of the cheeks : —
i. Scales on the temples only : nearly half the snout is pre-oral H. anguilliformis.
ii. Scales on the temples and cheeks : not quite a third of the
snout is pre-oral ... ... ... ... E. mediorostris.
2. Head and snout scaly : half the snout is pie-oral ... ... H. parvipinnis.
II. Ventral fins united together by membrane [Halosaurichthys] : —
1. Head naked, except for a few scales on the temples and cheeks ;
about a third of the snout is pre-oral ... ... ... H. nigerrimus.
2. Head scaly : not quite two-fifths of the suout is pre-oral ... H. carinicauda.
146. Halosaurus anguilliformis, Alcock.
Halosaurus anguilliformis, Alcock, Ann. Mag. Nat. Hist. Dec, 1889, p. 453.
Halosaurus Hoskynii, Alcock, Ann. Mag. Nat. Hist., Oct. 1890, p. 309: Illustrations op the Zoology of the
Investigator, Fishes, pl. VII. fig. 3.
B. 10. D. 11-12. A. circ. 175. P. 13. V. 1/8. L. tr. 13-14, between D. and V.
Head naked except for a few scales on the temples, its length about - of the
total, and exceeding the distance between the gill-opening and the base of the
ventral fin by about an eye-length.
Length of the snout 2- to 2- in that of the head, the pre-oral portion being
not quite a half of the whole.
The major diameter of the eye nearly equals the width of the interorbital
space, and is contained six and a half times in the head-length and just over 3
times in the length of the postorbital portion of the head.
The maxilla does not quite reach the vertical through the anterior margin
of the orbit.
The pterygoid band of teeth is very broad and is separated from the pala-
tine band by a considerable interval.
Eight moderately long gill-rakers on the middle of the first arch, besides
some small ones above and below.
Scales extremely deciduous, those on the lateral line larger and more adher-
ent than the rest, measuring - of an inch in diameter and having a small central
perforation. The scales of the lateral line are thirty in number between the gill-
opening and the vent, beyond which they are in contact with the rays of the
anal fin.
The pectorals are slightly larger than the post-rostral portion of the head.
The dorsal fin begins about an eye-length behind the insertion of the first
(outermost) ventral ray.
185
Seven or eight large pyloric casca in a longitudinal row embracing the ascend-
ing limb of the stomach.
Colours in the fresh state : — body and fins uniformly dark sepia-brown,
opercles silvery, throat black.
Four specimens, the largest 21 inches long.
Arabian Sea, off the Laccadive Islands, 1000 fathoms ; Gulf of Manar, 675
fathoms.
Regd. Nos. 11541, 12874, 12875, 12876, 12877.
This species may possibly prove to be identical with H. affinis, Giinther,
from Japanese Seas.
147. Halosaurus mediorostris, Gthr.
Halosaurus mediorostris, Giinther, Challenger Deep-Sea Fishea, p. 239, pi. lis. fig. C : Alcock, Jonrn. As. Soc.
Bengal, Vol. LXI1I pt. 2, 1894, p. 136.
B. 11. D. 11. P. 8. V. 1/8.
Head with some scales on the temples and upper part of cheeks ; otherwise
naked.
Length of the snout about 2- in that of the head ; not quite a third its
extent is pre-oral.
Eye small, its diameter is less than a fourth the length of the postorbital
portion of the head and about two-thirds the width of the interorbital space.
The maxilla reaches to, or slightly beyond, the anterior border of the
eye.
Pterygoid teeth separated from the palatine teeth by a considerable interval.
Numerous, close-set, longish gill-rakers on the outer side of the first bran-
chial arch.
Scales deciduous, those of the lateral line enlarged.
The dorsal fin begins about an eye-length behind the outermost ventral
ray.
The pectorals, which are extremely slender, are not quite equal in length to
the postrostral portion of the head.
Colour light brown, greater part of head blackish.
Arabian Sea, between the Maldives and Cape Comorin, 719 fathoms.
Regd. No. 13710.
Distribution : off Philippines ; Arabian Sea.
24
186
This species is distinguished from H. anguilliformis by the short pre-oral
portion of the snout and the numerous close-set gill-rakers on the outer side of
the first branchial arch.
148. Halosaurus parvipennis, Alcock.
Halosaurus parvipennis, Alcock, Ann. Mag. Nat. Hist., Nov. 1892, p. 362 : Illustrations of the Zoology of
THE INVBSTIGATOR, FlSHES, PL. XXXIII. FIG. 1.
B. 12-13. D. 9-10. P. 12-13. V. 1/9. L. tr. 14-15.
Head and snout scaly.
Length of head about an eighth of the total, and less than the distance be-
tween the gill-opening and the base of the ventral fin.
Length of the snout a little over a third that of the head, half its extent is
preoral.
The major diameter of the eye is much more than twice the width of the
interorbital space, is from a fifth to two-elevenths the length of the head, and
nearly half the length of the postorbital portion of the head.
The maxilla does not quite reach the vertical through the anterior margin
of the orbit.
The pterygoid band of teeth is short and narrow and is hardly separated
from the palatine band.
The gill-rakers on the outer side of the first branchial arch are short and
distant.
Scales adherent, especially on the lateral line. Those of the lateral line are
but slightly enlarged, especially at the anterior end of the line, and number
between 50 and 60 from the gill-opening to the vent, beyond which they are
separated from the anal fin, at first by two, and then by one series of scales.
An irregular series of scales in the middle line behind the dorsal fin are consi-
derably enlarged and elongated.
The dorsal fin begins about an eye-length behind the insertion of the outer-
most ventral ray. The very narrow pectorals are about as long as the post-
orbital portion of the head.
About eight extremely short and inconspicuous pyloric caaca.
Colours : sepia-brown, opercles silvery : throat, etc., black.
The largest specimen (adult) is about 15r inches.
Arabian Sea, off Malabar coast, 459, 636, and 865 to 880 fathoms.
Eegd. Nos. 13198, 14005, ^9, ^.
187
149. Halosaurus carinicauda, Alcock.
Ealosaurichthys carinicauda, Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 454: Illustrations of the Zoology
of the Investigator, Fishes, pl. VII. figs. 2, 2a.
B. 12-13. D. 11. P. 14-15. V. 1/9. L. tr. 15-16 between D. and V.
Head and snout scaly.
Length of the head about a seventh of the total and considerably less than
the distance between the gill-opening and the base of the ventral fin.
Length of snout a little over a third that of the head, not quite two-fifths of
its extent is preoral.
Major diameter of the eye not twice the width of the interorbital space,
about a seventh the length of the head and a little more than a fourth the length
of the postorbital portion.
Maxilla not reaching the anterior margin of the orbit.
Pterygoid band of teeth narrow, scarcely separated from the palatine
band.
Seven or eight distant, short gill-rakers — in addition to some smaller ones —
on the outer side of the first branchial arch.
Scales deciduous, except on the lateral line where they are but little en-
larged. Those of the lateral line number about 60 between the gill-opening and
the vent, beyond which they are separated from the anal fin, at first by two and
then by one series of scales. Some scales in the middle line behind the dorsal
fin are enlarged and elongated, and in the posterior part of the tail these are set
in a low median fold of skin.
The dorsal fin begins about an eye-length behind the outermost ventral ray.
The pectorals are not quite equal in length to the post-rostral portion of the
head.
The ventrals are coherent into a single plate, the union being membranous.
Five or six extremely small and inconspicuous pyloric ca?ca.
Colours : sepia-brown, the greater part of the head blackish.
Length 15- inches.
Andaman Sea, 490 fathoms.
Regd. No. 11763.
This species is very closely related to Halosaurus parvipennis, from which
it differs chiefly in having the ventral fins united with one another, the preoral
part of the snout much shorter, and the eyes smaller and farther apart.
188
150. Halosaurus nigerrimus, Alcock.
Halosaurus nigemmns, Alcock, Ann. Mag. Nat. Hist., Aug. 1898, p. 149 : Illustrations of the Zoologt of
the Investigator, pl. XXXIII. fig. 2.
B. 12. D. circ. 12. P. circ. 9. V. 8.
Temples and cheeks scaly; head otherwise naked. Length of the head
about a seventh of the total and nearly equal to the distance between the
gill-opening and the base of the ventral fin. Length of the snout about 2-
in that of the head ; about a third of its extent is preoral. Eyes about two-
thirds of a diameter apart.
The maxilla reaches to the anterior edge of the pupil.
Teeth as in H. carinicauda.
Numerous long close-set gill-rakers on the outer side of the first branchial
arch.
Scales deciduous : those of the lateral line adherent and much enlarged and
about 30(?) in number between the gill-opening and the vent. There is a low
median fold of skin, with some enlarged scales, behind the dorsal fin, as in
H. carinicauda.
Fins as in H. carinicauda, but the ventrals are united with one another only
in their basal portion.
Colour : everywhere uniform jet black.
One young specimen about 75 inches long, from off the Maldives, 459
fathoms.
Regd. No. if.
Family Muraenidce.
To the ten genera and thirty-nine species recorded in the Fauna of British
India, the " Investigator " has added other ten genera and seventeen species.
Key to the Indian Deep Sea genera of Muraenidie.
The branchial openings into the pharynx are wide slits : the heart is
situated between or close behind the gills : —
1. Vent close to the gill-opening : nostrils close together in a hollow in
front of the eye : jaws produced to form a long slender beak :
NEMICHTHYTNA :—
i. Pectorals present, the vent being between their roots ... Nemichthys.
ii. Pectorals absent : vent about a snout length behind the gill-
opening ... ... ... ... ... Gavialiceps.
2. Vent at no great distance from the gill-opening : nostrils distant from
one another, lateral : snout not beak-like : eyes small : DYSOM-
MINA:—
i. Pectorals present, the vent being between their roots ... Dysomma.
189
ii. Pectorals absent : vent nearly tliree-quarters of a bead-length
behind the gill-openings
3. The distance of the vent from the gill-opening is either equal to or
very much more than the distance between the gill-opening and the
tip of the snout : the nostrils are distant from one another : —
i. Gill-openings united in a longitudinal slit: STNAPHOBRAN-
CHINA
ii. Gill-openings separate : —
a. Nostrils superior or lateral : tongue free : vertical fins con-
fluent round the tip of the tail : pectoral fins present :
ANGUILL1NA :—
a. Scaleless : the cleft of the mouth extends slightly beyond
the middle of the eye : —
Teeth in a single series in the jaws ; none on the vomer
Teeth in bands in the jaws ; vomerine teeth
Teeth in the jaws biserial ; vomerine teeth
p. Scaleless : the cleft of the mouth reaches the eye : teeth
in the jaws in broad bands, and in a broad confluent
patch on the palate ...
b. Nostrils superior or lateral : tongue not free, or if free curi-
ously truncated : vertical fins confluent round the tip of the
tail: no scales: MUR^ENESOCINA :—
a. Pectorals present : snout of good length but not bill-like :
teeth in the mandible and maxilla practically uniserial ;
a single row of teeth in the vomer
j3. Pectorals present : snout very long almost bill-like :
teeth in the jaws in broad bands and very sharp : a
single row of teeth in the vomer. Maxilla with a broad
longitudinal groove, running the whole length of the
bone and dividing the band of teeth into two portions ...
y. Pectorals absent : snout very long aud bill-like : teeth
in the jaws in two or three series, small and sharp : a
lone row of enlarged teeth on the vomer : no air-bladder
8. Pectorals absent : snout very long and bill-like : small
stout conical teeth in very broad bands in the jaws and
vomer: air-bladder present : lateral line indistinct
Dysommopsis.
Synaphorrancbus.
coloconger.
congromcr&na.
Uroconger.
PrOMYLLAN TOR.
Sauromurj:nesox.
Xenomystax.
Sadrenchelys.
Nettenchelys.
Nemichthyina, Gunther.
Nemichthys, Richardson.
Nemichthys, Richardson, Samarang Pishes, p. 25 : Gunther, Cat. Fishes, VIII. p. 21 : Goode and Bean, Oceanic
Ichthyology, p. 151 : Jordan and Evermann, Pishes of N. Amer., p. 369.
Leptorhynchus, Lowe, Ann. Mag. Nat. Hist. X. 1852, p. 54.
Belonopsis, Brandt. Mem. Ac. Petergb. Sav. Strang. VII. 1854, p. 171.
Investigator, Goode and Bean, Oceanic Ichthyology, p. 518.
Body exceeding elongate, tail lash-like. No scales. Lateral line distinct.
Vent near the gill-opening and pectorals, at the anterior end of the long
abdominal cavity.
190
Jaws produced to form a long slender bill, of which the upper part is
formed by the vomer and premaxillas. The inner surface of the bill covered
with small close-set teeth. Similar teeth on the mandible. Eye large. The
nostrils of each side are close together, in a hollow before the eye. Gill-openings
wide, nearly confluent. Pectoral and vertical fins well-developed. The dorsal
fin begins close behind the head.
Distribution : Atlantic ; Indian Ocean ; Pacific.
151. NemichtJiys acanthonotus, Alcock.
Nemichthys acanthonotus, Alcock, Jouro. As. Soc. Bengal, Vol. LXIII. pt. 2, 1894, p. 136: Illustrations of
the Zoology of the Investigator, Fishes, pl. XIV. fig. 5.
Investigator acanthonotus, Goode and Bean, Oceanic Ichthyology, p. 518.
The posterior third or more of the long slender body is very abruptly con-
stricted to form a lash-like tail.
The head, rather more than four-sevenths of which is formed by the long
tapering snout, is between one-seventh and one-ninth of the total. The diameter
of the subcutaneous eye is between one-third and one-fourth the length of the
post-orbital portion of the head, and between one-sixth and one-seventh the
length of the snout. The nostrils have the usual position, and the jaws are
curved at tip as in N. infans. Small recurved asperities in crowded bands form
the dentition of the jaws and vomer.
The vent is situated immediately behind the gill-opening and the root of the
pectoral fin.
The gill-openings, which are wide, are separated from one another only by
a thin fold of skin.
No scales. The lateral line is marked by a series of small glistening pores
which are arranged with beautiful regularity in " fives" (quincunces). The head
is studded with similar pores.
The dorsal fin commences on the occiput, and is continued to the tip of the
tail : in a part of its extent, somewhat less than the middle third, the long
slender rays are replaced by strong short spines — like those of Notacanthus —
interconnected by a low membrane. The anal fin, which commences immediately
behind the vent, has its rays well-developed throughout, — the longest rays being
considerably more than half the length of the post-rostral portion of the head.
The pectorals are large, and are half as long as the post-orbital portion of
the head.
Colours : uniform dark sepia becoming black ventrally : gill-covers and fins
black.
The largest specimen is over 30 inches long.
Bay of Bengal, 475 fathoms ; Arabian Sea, 636 fathoms.
Regd. Nos. 13643, 14003.
191
This species is distinguished from all its congeners by the long series of
strong sharp spines in the middle of the dorsal fin. On this account Messrs.
Goode and Bean (Oceanic Ichthyology, p. 518) separated it from the genus
Nemichthys and gave it the generic name Investigator, a course in which I am
unable to follow them.
G-avialiceps, Wood-Mason MS., Alcock.
Qavialiceps, Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 460, and Nov. 1892, p. 364.
Body exceedingly elongate, tail tapering. No scales. Lateral line indis-
tinct. Vent about a snout-length behind the gill-opening, near the anterior end
of the long abdominal cavity.
Jaws produced to form a long slender bill, of which the upper part is
formed by the premaxilla? and vomer. Sharp teeth on the vomer, premaxilla?,
and mandibles.
Eye small. Nostrils as in Nemichthys. Gill-openings wide, nearly conflu-
ent. Pectorals absent. Anal well developed. Dorsal ill-developed and begin-
ning a considerable distance behind the head.
Gavialiceps differs from Nemichthys in having (1) no pectoral fins, (2) the
dorsal fin ill developed and beginning some way behind the head, (3) the eye
small, (4) the vent not quite so near the throat, and (5) the lateral line indistinct.
152. Gavialiceps microps, Alcock.
Qavialiceps microps, Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 461 and Nov. 1892, p. 364.
The head, of which rather more than two-fifths is formed by the long taper-
ino- snout is between a fifth and a sixth of the total length. The diameter of the
small subcutaneous eye is about one-eleventh the length of the post-orbital por-
tion of the head (measured to the gill-opening) and about a ninth the length of
the snout.
The premaxillary teeth are arranged in a single row, and diminish in size
but increase in number from behind forwards, where they end in a narrow band
of minute asperities : the vomerine teeth posteriorly are long and sharp and are
disposed in a long, close-set, comb-like series ; anteriorly they form a fine rasp-
like band of minute asperities : in the mandible a row of large distant needle-like
teeth stands up from an uneven band of small denticles. Gill-openings close
together, wide. The scaleless integument is thin and deciduous and thickly en-
veloped in mucus ; no lateral line is apparent. The dorsal fin is feebly develop-
ed, and, indeed, hardly distinguishable, except in the posterior part of the body :
it begins about a head-length behind the head. The pectoral fin is represented
by an inconspicuous knob, without any rays.
192
The abdominal cavity extends at least halfway along the tail. The siphonal
stomach, which has its pyloric end long, tapering, and much constricted, leads
into a widely expanded duodenum, which, in the single specimen dissected, is
furnished with a small diverticulum near the pylorus. The vent is about a snout
length behind the gill-opening.
Colour uniform black, with a silvery sheen on the head.
The largest specimen is about 14- inches long.
Bay of Bengal, off the Andamans and Nicobars, 1045 and 869 to 913
fathoms ; Arabian Sea, off the Laccadives, 1045 fathoms ; Gulf of Manar 902
fathoms.
Regd. Nos. 11662, 13201, 25,
This species is perhaps identical with Nemicldhys infans Vaillant (nee
Giinther), described and figured in Exped. Sci. du ' Travailleur ' et du ' Talis-
man,' Poiss. pp. 93 and 94, pi. vii. fig. 1, and there only doubtfully referred to
Dr. Giinther' s type.
Dysommina.
Dysomma, Alcock.
Vysomma, Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 459.
The branchial openings into the pharynx are wide slits. The heart is situa-
ted between the gills. The tail is exceedingly long, the vent being situated
immediately behind the gill-opening and between the bases of the pectoral
fins.
Tail tapering. Snout short but projecting beyond the mouth and lower
jaw. Eyes very small, subcutaneous. Nostrils large, lateral, those on the same
side distant from each other. Cleft of the mouth wide. Small sharp teeth in a
single row or extremely narrow band in each jaw : a short row of large teeth on
the vomer. Tongue not free.
Gill-openings rather small, well separated. No scales. Lateral line of
minute pores. Pectorals and vertical fins well developed, the dorsal beginning
immediately behind the gill-opening, the anal immediately behind the vent.
153. Dysomma bucephalus, Alcock.
Dysomma bucephalus, Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 459 : Illustrations of the Zoology of the
Investigator, Fishes, pl. VI. fig. 1 : Ann. Mag. Nat. Hist., Aug. 1891, p. 137, fig. 5.
Length of the head, measured to the gill-opening, between a fourth and a
fifth of the total. Vent situated with the abdominal pore on a large round fleshy
" clitellum," immediately behind the gill-opening.
193
Snout short, between a fifth and a sixth of the length of the head measured
to the gill-opening, round-pointed, somewhat depressed, studded, like the head
and cheeks, with minute pores.
Byes very small, their diameter about a fifth the length of the snout, deeply
sunk beneath the skin. Nostrils large, the anterior tubular, the posterior simple.
Mouth-cleft wide, rather more than a third the length of the head ; lips
inflated, with many small pores ; jaws weak. A row or very narrow band of
minute teeth in the upper jaw ; a row of small teeth in the lower jaw ; a short
row of large teeth at the fore end of the vomer.
Gill-covers formed of tough skin, in which neither bony opercles nor bran-
chiostegal rays can be detected. Branchial arches very weak : gill-laminae broad.
No scales : the lateral line, which consists of a row of small pores, follows
the dorsal curve.
Vertical fins fairly well developed. Pectorals rather longer than the snout,
rounded or truncated.
Colour brown, paler beneath ; vertical fins darker, edged with white.
The body-cavity extends more than halfway along the tail, and the intestinal
loop follows it. Air-bladder thick-walled, nacreous, consisting of three lobes — a
large middle one with a small one on each side.
An adult female is 11 inches long.
Bay of Bengal, 112, 128, 193, 145 to 250, and 240 fathoms.
Regd. Nos. 11675, 13108, 13436, 13644-13646.
Dysommopsis, Alcock.
Dysommopsis, Alcock, Ann. Mag. Nat. Hist., Ang. 1891, p. 137.
Closely related to Dysomma, from which it differs, much as Gwvialkeps differs
from Nemichthys, in the following particulars : —
The vent is situated nearly three-quarters of a head-length behind the gill-
opening, and there are no pectoral fins : the dorsal fin begins some way behind
the gill-opening.
154. Dysommopsis mucipara, Alcock.
Dysommopis muciparus, Alcock, Ann. Mag. Nat. Hist., Aug. 1891, p. 137.
Head a little inflated in the branchial region, tapering anteriorly ; its length
a little more than one-eighth of the total. The vent lies with the genital pore
in an unpigmented circular depression, which is situated at a distance from the
gill-opening equal to the length of the post-rostral portion of the head ; the tail,
25
194
which tapers very slightly, is therefore more than four times the combined head
and trunk in length.
Snout acutely pointed, overhanging the upper jaw; its length is one-fifth
that of the head and 2- times that of the small deeply subcutaneous eye ; its
surface is densely crowded, like the lips, with minute pores. Nostrils large ; the
anterior, which is tubular, is situated near the tip of the snout, the posterior is a
valved foramen lying immediately before the angle of the eye.
Mouth wide, its cleft being nearly half the head in length ; small sharp
teeth in a single row in the mandible and a double row in the maxilla ; vomer
with a row of three large teeth. The mandibular teeth are small and close-set
posteriorly, larger and more distinct anteriorly.
Gill-openings small, close together near the mid-abdominal line ; the gill-
covers are formed of tough skin, in which branchiostegal rays are faintly ap-
parent ; branchial arches weak, gill-lamina? broad.
Skin scaleless, enveloped in thick, very tenacious mucus. Lateral line a
row of indistinct pores. Vertical fins confluent, the dorsal beginning halfway
between the gill-opening and the vent, the anal immediately behind the vent.
No pectoral fins.
The abdominal cavity extends almost to the tip of the tail, its posterior part
being occupied solely by the genital glands and air-bladder.
Stomach with a long tapering csecal sac reaching some distance behind the
vent, intestine forming a single loop, the convexity of which embraces the gastric
caecum. Air-bladder a long nacreous tube extending from the occiput almost to
the tip of the tail ; much inflated anteriorly, and tapering posteriorly to a fine
thread.
Colours in life deep purple-black.
Bay of Bengal, 240 to 270 fathoms.
Regd. Nos. 13106, 13107.
Synaphoekanchina, Gthr.
Synaphobranchus, Johnson.
Synaphohranchuf, Johnson, Proc. Zool. Soc, 1862: Giinther, Cat. Fishea, VIII., p. 22, and Challenger Deep-Sea
Fishes, p. 253 : Goods and -Bean, Oceanic Ichthyology, p. 142 : Jordan and Evermann, Fishes N. Amer., p. 351.
Gill-openings ventral, united into a longitudinal slit between the pectoral
fins, separate internally. Pectoral and vertical fins well developed. Nostrils
lateral, the anterior subtubular. Cleft of the mouth very wide ; teeth small.
Body scaly.
Distribution : Atlantic, Arabian Sea, Western Pacific.
195
155. Synapliobranchus pinnatus, (Gronov.) Gthr.
Synaphobranchws pinnatus, Giinther, Cat. Pishes, VIII., p. 23, and Challenger Deep-Sea Fishes, p. 253, pi. lxii.,
fig. A. (ubi synon.) : Vaillant, Exp. Sci. Travailleur et Talisman, Poiss. p. 88, pi. vi., fig. 2 : Goode and Bean,
Oceanic Ichthyology, p. 143, fig. 164 : Jordan and Evermann, Fishes N. Amer., p. 351.
Length of the head not quite a seventh of the total, and equal to the distance
between the gill-opening and the vent ; length of the tail, therefore, considerably
over twice that of the rest of the body.
Length of the snout about a third that of the head and rather over twice
that of the eye.
Mouth-cleft rather over half the length of the head. Teeth in the jaws in
a narrow band with some enlarged ones anteriorly : a single short row of largish
teeth on the preniaxillary, and a single row of small teeth on the vomer.
Scales rudimentary, forming a diagonal pattern. Lateral line distinct.
Cheeks and occiput scaly.
The dorsal fin arises about a head-length and a third behind the gill-opening,
and therefore a short distance behind the anal : both fins are low. Pectorals half
as long as the head.
Colour purple-black or brown.
171 31R
Arabian Sea, 459 and 824 fathoms. Registered No. — : — .
In the Indian Museum are one of the " Challenger " duplicates, and a speci-
men from America.
Anguillina, Gthr.
CoLOCONGER, Alcock.
Coloconger, Alcock, Ann. Mag. Nat. Hist., Dec. 18S9, p. 456.
The branchial openings into the pharynx are wide slits. The heart is
situated between the gills. The tail is a good deal shorter than the combined head
and trunk. The gill-openings are separate. The muscular and osseous systems
are well developed. The posterior nostril is superior. The tongue is free. The
end of the tail is surrounded by the fin. Pectoral fins are present. No
scales.
Snout short. Eyes large. Cleft of the mouth wide, extending beyond the
middle of the eye. The teeth form a sharply prominent serrated ridge in each
jaw : there are none on the vomer. The vertical fins are well developed, the
dorsal beginning above the base of the pectoral.
196
156. Coloconger raniceps, Alcock.
Coloconger raniceps, Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 456: Illustrations of the Zoology of the
Investigator, Fishks, pl. VII. fig. 4.
Head frog-like, its length (measured to the gill-opening) is half, or some-
times a little more than half, the distance between the gill-opening and the vent,
and a fifth, or sometimes a little more than a fifth, the total.
Snout blunt, shorter than or as long as the eye. Bye about a fourth the
length of the head and equal to the -width of the interocular space. Nostrils
large, the anterior sub-tubular, the posterior above the angle of the eye.
Mouth cavernous, its cleft extending to the hinder edge of the pupil. Jaws
slender, equal. Tongue short, broad, fleshy, free in its anterior third. In each
jaw a row of small uniform teeth in continuous contact, except at their extreme
tips, which show as minute recurved asperities on a sharp-edged ridge. No
vomerine teeth. A large, oval, horny, granular plate in the fauces behind the
superior pharyngeal bones. A mucous channel with numerous pores along the
lower jaw beneath.
Gill-lamina3 narrow ; gill-openings of moderate size, a broad fold extends
from their outer edge to the base of the pectoral fin.
No scales. Head with numerous black tubular papilla?. Lateral line a
salient tube, with upwards of a hundred similar papilla?.
Vertical fins confluent; the dorsal, which begins above the base of the
pectoral, is considerably higher than the anal. Pectorals two-fifths of the length
of the head, or a little longer.
Colours in spirit : brown ; abdomen speckled with black, due to the peri-
toneal pigment showing through.
Visceral peritoneum black. Stomach with a cascum half as long as the
body-cavity. Intestine sinuous. Only the left lobe of the liver developed. Air-
bladder large, globular.
Length 6^ to 10- inches.
Andaman Sea, 265, 271, and 405 fathoms; Bay of Bengal, 200 to 400
fathoms ; Arabian Sea, off Malabar coast, 224 to 284 fathoms.
Regd. Nos. 11637, 11639, 11778, 11779, ^ ™, 'f, '-^j^-
CONGROHURiENA. Kaup.
One species of this genus is included in the Fauna of British India ; the
"Investigator" has brought five more to light, all being inhabitants of deep-,
water.
197
ii.
Key to the Indian species of Congromurasna.
The length of the head measured to the gill-opening, is less than the
distance between the gill-opening and the vent : —
1. The head is at least three eye-lengths less than the distance bet-
ween the gill-opening and the vent : —
i. The band of vomerine teeth is of good length... ... C. anago.
ii. The band of vomerine teeth is extremely short : series of
small black dots above and below the lateral line ... C. guttulata.
2. The head is an eye-length shorter than the distance between the
gill-openiug and the vent ... ... ... ... C. macrocercus.
The length of the head, measured to the gill-opening, is equal to or
greater than the distance between the gill-opening and the vent : —
1. The length of the snout is a fifth that of the head : the dorsal fin
begins in advance of the gill-opening ... ... ... C. squaliceps.
2. The length of the snout is about a fourth that of the head : the
dorsal fin begins above the gill-opening: —
i. Outer teeth in the jaws decidedly enlarged : vomerine teeth
in a short very narrow band ... ... ... 0. nasica.
ii. None of the teeth eularged : vomerine teeth in a very
broad band of some length ... ... ... C. musteliceps.
157. Congromurcena squaliceps, Alcock.
Oongromursena squaliceps, Alcock, Joarn. As. Soc. Bengal, Vol. LXII., pt. 2, 1893, p. 183 .
Batkycongrus squaliceps, Ogilby, Proc. Lino. Soc. N. S. Wales, XXIII., 1898, p. 293.
Head, measured to the gill-opening, about an eye-length, longer than the
distance between the gill-opening and the vent, which is not quite one-fourth the
length of the tail.
The snout, which projects far beyond the mouth, is one-fifth the head in
length : it is broadish and blunt pointed. The major diameter of the very elliptic-
al eye is not quite two-thirds of the length of the snout. The anterior nostril
is a short wide tube situated on the lip near the end of the snout, the posterior
is a wide foramen situated in advance of and above the angle of the eye.
The mouth-cleft is wide, extending almost to the vertical through the
posterior border of the orbit, and the lips are greatly developed : the minute
teeth are in bands in the jaws, and in a broad rasp-like patch outside the mouth
in the premaxillary ; there are a few teeth on the vomer quite anteriorly.
Gill-openings comparatively wide, separate. No scales : lateral line with
small pores. Pectorals narrow, half an eye-length longer than the snout.
Vertical fins confluent, the dorsal beginning a little in advance of the gill-
opening.
Colour in spirit, grey, the vertical fins in their after half to two-thirds with
a black edge, which in the anal tends to involve the whole fin. Pectorals
198
hyaline. A very large air-bladder extending half a head-length beyond the vent.
Visceral peritoneum silvery.
The largest specimen is 20 inches long.
Bay of Bengal, 128 and 195 to 210 fathoms.
Regd. Nos. 13693-13695, 13697, 13450.
158. Congromurcena macrocercus, Alcock.
Congromursena longicauda, Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 455 (name preocc.) : Illustrations of
the Zoology of the Investigator, Fishes, pl. VII. fig. 5.
Congromursena macrocercus, Alcock, Journ. As. Soc. Bengal, Vol. LXIII., pt. 2, 1894, p. 134.
Bathycongrus macrocercus, Ogilby, loc. cit. supra.
Differs from G. squaliceps only in the following particulars : —
The head, measured to the gill-opening, is an eye-length shorter than the
distance between the gill-opening and the vent, which is not quite a fourth the
length of the tail.
The snout, which has the same form and the same relations to the mouth, is
a little more than a fourth the length of the head.
The eye is very little more than half the length of the snout.
The mouth-cleft reaches to the posterior border of the pupil.
The dorsal fin begins above the gill-opening.
The narrow pectorals are as long as the snout.
Colour brown, the fins grey.
Length 16 inches.
Andaman Sea, 265 fathoms ; Bay of Bengal, 240 to 276 fathoms.
Regd. Nos. 11781, 11782, 13452.
159. Corigromurceim nasica, Alcock.
Congromureena nasica, Alcock, Journ. As. Soc. Bengal, Vol. LXII., pt. 2, 1893, p. 183 : Illustrations of the
Zoology of the Investigator, Fishes, pl. IX. fig. 2.
Bathycongrus nasicus, Ogilby, loc. cit. supra.
Differs from G. squaliceps only in the following particulars : —
The head, measured to the gill-opening, is very slightly longer than the
distance between the gill-opening and the vent, which is considerably more than
a fourth (1 : about 3'4) the length of the tail.
The snout is a little over a quarter the length of the head and is rather
sharp-pointed.
The eye is not much more than half the length of the snout.
199
The teeth are in two bands in each jaw, the outer band being enlarged : the
vomerine teeth also are enlarged.
The dorsal fin begins over the gill-opening. The pectorals are as long as
the snout.
The colour is grey : the vertical fins in their posterior third or fourth have
a very narrow black edge.
Length of adult females from 10 to 12 inches.
Bay of Bengal, 128 and 195 to 210 fathoms.
Regd. Nos. 13453-13456, 13649-13692.
160. Cotigromnrcena musteliceps, Alcock.
Congromursena musteliceps, Alcock, Journ. As. Soc. Bengal, Vol. LXIII., pt. 2, 1894, p. 133, pi. vii., fig. 5 :
Illustrations of the Zoology of the Investigator, Fishes, pl. XV. fig. 7.
Bathycongrus musteliceps, Ogilby, loc. cit. supra.
Differs from C. squaliceps only in the following particulars : —
The snout is sharp-pointed ; its length is a quarter that of the head and fully
twice that of the eye.
The mouth-cleft reaches to the posterior border of the pupil.
The vomerine teeth are in a broad band of some length.
The dorsal fin begins above the gill-opening.
Colours dark brown, pectorals and vertical fins nearly black.
Length 15 inches.
Bay of Bengal, 145 to 250 fathoms.
Kegd. Nos. 13698-13702.
161. Congromurcena guttulata, Gthr.
Congromurxna guttulata, Giinther, Challenger Deep-Sea Fishes, p. 252.
Congrellus guttulatus, Ogilby, loc. cit. supra.
This species, if my identification be correct, differs from G. squaliceps, only
in the following particulars : —
The head measured to the gill-opening is more than three eye-lengths shorter
than the distance between the gill-opening and the vent, which is not quite a
third the length of the tail.
The snout, which is blunt-pointed, is a fourth the length of the head.
The diameter of the eye is between half and two-thirds the length of the
snout.
The mouth-cleft reaches to the posterior border of the pupil.
The dorsal fin begins behind the root of the pectoral fin.
200
Colours, yellowish brown, with series of small black dots and splashes above
and below the lateral line.
Length nearly two feet.
Off Malabar coast, 636 fathoms.
Regd. No. 14009.
Distribution : Off Fiji Is. 315 fathoms; Arabian Sea.
Ueoconger, Kaup.
162. Uroconger vicinus, Vaillant.
Vroconger vicinus, Vaillant, Exp. Sci. Travailleur et Talisman, Poiss. p. 86, pi. vi., fig. 1 : Alcook, Ann. Mag.
Nat. Hist., X., 1892, p. 363 : Goode and Bean, Oceanic Ichthyology, p. 138.
Length of the head, measured to the gill-opening, not much more than half
the distance between the gill-opening and the vent, and between a seventh and
an eighth of the total.
Body high for an eel ; its greatest height being rather more than half the
length of the head.
Muciferous cavities of the head well developed, the pores on the snout and
in a line running from the mandibular symphysis towards the gill-opening being
very conspicuous.
Snout about twice as long as the eye and between a third and a fourth the
length of the head. Anterior nostril subtubular, situated not far from the end
of the snout, difficult to distinguish from the mucous pores : posterior nostril
situated just above the eye.
The mouth-cleft reaches at least to the middle of the eye. Teeth large,
especially on the premaxillae and at the anterior end of the maxillary and mandi-
ble. Vomerine teeth reduced to two large ones — one behind the other — at the
anterior end.
Gill-openings of moderate size, separated by an interval greater than their
own greatest diameter.
Lateral line very distinct, its pores white.
The dorsal fin begins about a snout-length behind the gill-opening. Pec-
torals narrow, pointed, nearly as long as the combind eye and snout.
Colours purplish brown.
The ripe female is 25 inches long.
Bay of Bengal, 475 fathoms ; Arabian Sea, off Malabar coast, 430 and 636
fathoms.
Distribution : Cape Verde and Atlantic coasts of Morocco ; Indian seas.
201
This species is distinguished from U. lepturus Richardson, (1) by the reduced
number and larger size of the vomerine teeth, (2) by the widely -separated gill-
openings, (3) by the increased distance from the gill-opening of the dorsal fin.
The microscopic structure of the wall of the stomach of Uroconger vicinus.
The stomach has a lining membrane of two different kinds : in the anterior
half the mucous membrane is of an almost horny hardness ; in the posterior half
it is soft and glandular ; and the transition between the two is abrupt.
In vertical longitudinal sections of the stomach-wall, carried through the
abrupt line of demarcation between the two different regions of mucous mem-
brane, examined under the microscope, the following structure is seen : —
(1) Common to both regions of the stomach : (a) an external thin fibrous coat,
one-fortieth to one-sixth of a millimetre thick, with many longitudinal bundles
of muscular fibres and large blood-vessels ; (b) a very compact thin coat of trans-
verse nmscular fibres, about one-eighth of a millimetre thick ; (c) another very
compact layer of longitudinal muscular fibres, about one-seventh of a millimetre
thick ; (d) a very thick (j-lg millimetre) submucous coat made up of a loose
mesh-work of branching and anastomosing small-nucleated cells, the meshes be-
ing filled with lymphoid cells ; this coat also contains many blood-vessels, which
frequently traverse in their course large, compact, sharply-circumscribed nodules
of lymphoid tissue, and a great many branching pigment-cells.
(2) The mucous membrane of the anterior part, which is about one-eighth of a
millimetre thick, appears at first like a superficial layer of pure fibrous tissue ;
but good sections show that it consists of a stratified epithelium with its consti-
tuent cells compressed somewhat as in the horny layer of the human epidermis.
These compressed (horny) cells, however, are not flattened into plates to form a
smooth surface, but are angularly concreted to form a broken rough surface.
Beneath the superficial horny layer are several rows of cells of which the granu-
lar protoplasm seems to be fused into a solid mass, leaving only the nuclei dis-
tinct ; and beneath this again comes fibrous tissue gradually passing into the loose
submucosa.
(3) The boundary -line between the anterior horny mucosa and the posterior
soft mucosa is very abrupt, and in every section there is seen a conspicuous
thickening of the submucous coat at the expense of both the mucous and the
muscular coats. The mucous coat is made up of the compact ramifications of
an acino-tubular gland lined with granular, large-nucleated, cubical epithelium.
(4) The mucous membrane of the posterior part, which is rather over one-
fourth of a millimetre thick, is formed entirely of long tubular glands packed
close together, side by side, at right angles to the surface. These glands, which
much resemble mammalian gastric glands, are lined with a granular cubical
26
202
epithelium having large prominent vesicular nuclei ; they have broadish mouths,
and in their deepest third they end by subdividing into two or three long sinuous
branches, which lie in a plane parallel to that of the rest of the gland.
Promyllantor, Alcock.
Promyllantor, Alcock, Ann. Mag. Nat. Hist., Oct. 1890, p. 310.
Closely related to Congromurxna.
Body stout, with the muscular and osseous systems well developed. Tail
about as long as the trunk. Muciferous cavities of the head well developed.
Eye rather small. Cleft of the mouth narrow, not extending behind the middle
of the eye. Villiform teeth in broad bands in the jaws and in a broad confluent
patch on the palate. Tongue free. Nostrils lateral. Gill-openings widely
separate ; four gills with wide clefts. No scales. Pectoral and vertical fins well
developed, the latter confluent. The dorsal begins some distance behind the occiput.
163. Promyllantor purpureas, Alcock.
Promyllantor purpureas, Alcock, Ann. Mag. Nat. Hist., Oct. 1890, p. 310: Illustrations of the Zoologt
of the Investigator, Pishes, pl. VI., fig. 2.
The head is about a sixth, the tail is about half-a-head-length more than
half the total length ; the body is massive, its greatest height equals the length
of the post-orbital portion of the head.
Head with its muciferous cavities highly developed, low, broad, inflated,
ending in a broad, pointed, swollen snout, which is twice the length of the
eye or - the total length of the head, and conspicuously prominent beyond the
mouth. Eyes circular, set high up on the side of the head, deep beneath a small
transparent area of skin, a diameter and a half apart.
Anterior nostril a short wide tube situated inferiorly at the tip of the snout.
Posterior nostril a large circular foramen just above the anterior orbital angle.
Mouth-cleft reaching slightly behind the vertical through the anterior
border of the orbit ; the jaws completely hidden by the very thick inflated lips.
Villiform teeth in broad bands in the jaws, and in a broad, confluent, triangular
patch covering the palate. Tongue free.
Gill-openings small widely separated foramina, hardly larger than the eye ;
four gills with narrow lamina? and coarse lamellae and wide clefts ; no gill-rakers.
Integument thick, coriaceous, scaleless, investing the vertical fins and
completely concealing their rays. The lateral line traverses the middle of the
body.
Vertical fins confluent ; the dorsal begins a distance behind the occiput
equal to the length of the post-rostral portion of the head, or just behind the
203
level of the tips of the pectorals when laid full back. The anal begins immedi-
ately behind the vent. Pectorals small, pointed, equal in length to the rostr-
orbital portion of the head.
Stomach with a cul-de-sac of moderate size ; intestine wide, little convolu-
ted; liver large, indistinctly lobated, embracing the oesophagus. Air-bladder
very large, with very thick spongy walls and a small central cavity.
Colours in the fresh state : — body and fins uniform purple-black.
A mature female is 17 inches long.
Arabian Sea, off the Laccadive islands, 1000 fathoms.
Kegd. No. 12878.
Morjinesocina, Giinther.
Sauromur&nesox, Alcock.
Sauromur&nesox, Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 457.
The branchial openings into the pharynx are wide slits. The heart is
situated between the gills. The tail is nearly as long as the head and trunk
combined, and is much lower than the trunk. The snout is pointed, is of good
length and overhangs the mouth and lower jaw, but is not specially prolonged.
Byes large. Nostrils lateral. Cleft of the mouth wide. Sharp teeth in a single
row in the maxilla and mandible : some enlarged teeth in the premaxilla and at
the anterior end of the mandible : a row of fangs on the vomer. Tongue free.
Gill-openings separate. No scales. Lateral line distinct but not conspicu-
ous : each pore at the end of a small branch.
Vertical fins ill-developed confluent, the dorsal beginning a little in advance
of the gill-opening. Pectorals well developed.
164. Sauromurcenesox vorctx, Alcock.
Sauromurmnesox vorax, Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 458 : Illustrations of the Zoology op
the Investigator, Fishes, pl. VI., fig. 3.
Form of the body lizard-like, especially in the adult female, the body being
high with an arched back, and the tail being low, even at its junction with the
trunk, and tapering.
Length of the head, measured to the gill-opening rather over two-thirds
the distance between the gill-opening and the vent, or about two-ninths of the
total.
The length of the snout is twice the width of the interorbital space and
more than twice the diameter of the large circular eye ; it tapers to a fine point,
which is slightly hooked. Nostrils large, the anterior subtubular, at some
204
distance from the tip of the snout ; the posterior in front of the middle of the
eye.
Cleft of mouth wide, extending an eye-length behind the posterior border
of the orbit ; the upper jaw overlapping the lower. Tongue free, bicylindrical,
truncated. In maxillge and mandibles a single row of close-set, equal, acute
teeth of moderate size ; also in the maxilla? an inner very incomplete series of
similar teeth, and in the mandibles near their symphysis three pairs of canine
teeth, the middle of which are very large and fit when the mouth is closed into
a notch between the maxillaries and premaxillaries ; four large equal canines
in a row in the vomer ; premaxillae with three smaller canines, which project
when the mouth is closed.
Gill-openings wide, extending obliquely from the upper border of the base
of the pectoral fins to near the middle line of the abdomen ; a broad flap of skin
connects their anterior margin with the base of the pectoral fin ; gill-laminae
broad.
Integument thin, without scales. The lateral line follows the dorsal curve
and ends in the posterior half of the tail.
Vertical fins, especially the anal, feebly developed, confluent ; the dorsal
begins considerably in advance of the gill-opening, the anal behind a very large
abdominal pore. Pectorals longer than the snout.
Colours : chocolate above, whitish or silvery below ; vertical fins whitish,
pectorals dark brown edged with grey.
The ripe female is 14 inches long.
Bay of Bengal, 193 and 145 to 250 fathoms.
Eegd. Nos. 11672, 13648, 13703.
Xenomystax, Gilbert.
Xenomystax, Gilbert, Proo. Unitod States Nat. Mus. XIV., 1891, p. 348 : Goode and Bean, Oceanic Ichthyology,
p. 146 : Jordan and Evermann, Fishes of N. Amer., p. 360.
The branchial openings into the pharynx are wide slits. The heart is
situated between the gills. The tail is very much longer than the combined
head and trunk. The gill-openings are wide and are separated by a narrow
space. The posterior nostril is superior. The tongue is not free. The end of
the tail is surrounded by the fin. Pectorals present. No scales.
Snout long. Eyes large. Mouth-cleft very wide, extending behind the
eye, the upper jaw projecting much beyond the lower.
Teeth conical slender and sharp, mostly depressible, those in the jaws in
broad bands. Maxillary with a broad longitudinal groove, running the whole
205
length of the bone and dividing the band of teeth into two. A clump of teeth
on the premaxillary. A row of enlarged teeth on the vomer.
The vertical fins are well developed and confluent ; the dorsal begins a little
in advance of the gill-openings.
Lateral line conspicuous. Air-bladder present.
165. Xenomystax trucidans, Alcock.
Xenomystax trucidans, Alcock, Journ. As. Soc. Bengal, Vol. LXIII., pt. 2, 1894, p. 134: Illustrations of the
Zoology of the Investigator, Fishes, pl. XVI., fig. 5.
Length of the head, measured to the gill-opening, equal to the distance
between the gill-opening and the vent, or slightly more than a sixth of the total.
The depressed and sharply pointed snout is a little more than one-third of
the head in length and nearly four times the major diameter of the eye : its
mucous pores, like those of the mandible and of the rest of the head, are large
slits : the anterior nostril is a large sub-tubular slit situated on the lip close to
the tip of the snout, the posterior is a wide elliptical foramen situated, almost
superiorly, partly in the posterior and partly in the middle third of the snout.
The mouth-cleft is wide, extending an eye-length behind the posterior border of
the orbit, or more than half way along the head, and the maxillae are most
remarkably massive. The teeth are in broad crowded bands, acicular or canini-
form, and for the most part depressible : those in the upper jaw are in two bands
— an outer very broad band of large depressible teeth in four series which
increase in size from without inwards, and an inner narrow band or very close-
set row of small rigid teeth — the two bands being separated throughout their
whole extent by a broad groove : the pre-maxillary teeth, which are much
enlarged, are in a broad patch standing outside the closed mouth : the mandi-
bular teeth are in at least five series increasing in size from without inwards,
and at the symphysis, where they are greatly enlarged, they form a patch which
fits into a wide notch in the upper jaw : the vomerine teeth form a short row of
fangs. Tongue small and intimately adherent throughout to the floor of the
mouth. Skin scaleless, glandular. Lateral line formed by a row of large brilli-
ant close-set pores. Gill-openings wide, crescentic, separated by a very narrow
interspace.
Vertical fins well developed, the dorsal beginning just in advance of the
gill-opening. Pectorals narrow, pointed, more than half the snout in length.
The stomach is large, extending the whole length of the abdominal cavity,
and is very distensible : the intestine in its posterior portion is coiled in a series
of close pleats : only the left lobe of the liver is developed : pancreas large : a
large air-bladder extending behind the vent.
206
Colour : body and fins blue-black ; pectorals with narrow whitish edge and
tip : margin of gill-opening and of all the mucous pores of the head and lateral
line brilliant white.
A mature female is between 25 and 26 inches long.
Arabian Sea, between the Laccadives and the Malabar coast, 360, 406 and
719 fathoms.
Regd. Nos. 13704, ^, 5*1.
This species appears to differ from X. atrarius, dredged by the TJ. S. Fish
Commission in 401 fathoms off the coast of Ecuador, only in the greater relative
length of the tail, the nearer approximation of the gill-openings, and the greater
length of the pectoral fins.
Saurnchelys, Peters.
166. Sauretichelys taeniola.
Gavialiceps tseniola, Wood-Mason MS., Alcock, Ann. Mag. Nat. Hist., Dec. 1889, p. 460.
Nettastoma tseniola, Alcock, Ann. Mag. Nat. Hist., Ang. 1891, p. 135, and Oct. 1892, p. 364.
Length of the head, measured to the gill-opening, about five-eighths of the
distance from the gill-opening to the vent, and about a seventh of the total.
Length of the snout five times that of the eye, and contained about 2^
times in that of the head. The nostrils are difficult to distinguish from the large
elliptical symmetrically disposed mucous pores of the snout.
The mouth-cleft extends to, or beyond, the after limit of the eye, and the
upper jaw projects well beyond the lower.
Broadish bands of small sharp teeth in both jaws, the band in the upper
jaw subdivided by a median longitudinal toothless space. A patch of somewhat
enlarged teeth on the premaxillary, separated from the maxillary teeth by a
notch into which a patch of similarly enlarged teeth on the mandibular symphysis
fits. Three long rows of teeth on the vomer, the outer rows very small and
sometimes absent, the middle row very large. Tongue short, the edge of the tip
just free.
Gill-openings of moderate size, close together.
No scales : the lateral line is very distinct and consists of a row of large
pores which is continued right across the gill-cover to the occiput.
Vertical fins confluent ; the dorsal begins above the gill-opening.
No pectoral fins.
No air-bladder.
Colour : black in adult life, the young silvery.
Adult females are about 24 inches long.
207
Andaman Sea, 265 fathoms ; Bay of Bengal, 240, 270, 281 to 258 and 260
fathoms.
Regd. Nos. 13111-13113, 12467, 13098-13105, 13202-13206.
Nettenchelys, Alcock.
Nettenchelys, Alcock, Ann. Mag. Nat. Hist., Aug. 1898, p. 149.
Body stout; tail longer than the combined head and trunk. Muciferous
cavities of head well developed ; snout much produced, broad, depressed ; mouth-
cleft very wide, broad bands of small conical teeth in jaws and vomer ; tongue
not free ; a tubular nostril situated dorsally near the tip of the snout on either
side. Gill-openings of moderate size, well separated ; four gills with wide clefts.
No scales ; the lateral line, which consists of a single row of pores, is very
indistinct in the greater part of its extent. Dorsal and anal fins well developed,
confluent with the broad caudal only in the basal half of the latter. No pectoral
fins. An air-bladder ; no pyloric appendages.
167. Nettenchelys Taylori, Alcock.
Nettenchelys Taylori, Alcock, Ann. Mag. Nat. Hist., Aug. 1898, p. 150: Illustrations of the Zoology of the
Investigator, Fishes, pl. XXV. fig. 5.
Head, measured to gill-opening, one-seventh the total, and half as long as
the distance between the gill-openings and the vent ; the tail is thus a good deal
more than half the total. Snout a third the length of the head (measured to the
gill-opening), elongate, broad, depressed, bill-like, the upper jaw overlapping the
lower. There is a series of large pores along the upper lip, as also along each
side of the lower jaw and along the top of the snout on either side, but the only
undoubted nostrils are a largish tubular pair at the tip of the snout. Byes sub-
cutaneous, not much more than a fourth the length of the snout and not much
more than half a diameter apart.
The mouth-cleft reaches behind the eye ; the dental surface of both jaws is
broad and is crowded with row upon row of close-set conical teeth, which are
little more than villiform, though the innermost row in either jaw is slightly
enlarged. On the vomer is a long broad convex band of similar teeth — about
six longitudinal rows of them. The tongue is large and thick and tapers to a
point ; it is firmly adherent to the floor of the mouth.
Gill-openings of moderate size, lateral, well separated.
No scales. Although the mucous system and pores of the head are so well
developed, those of the lateral line, which are in a single row, very soon become
distant, small, and inconspicuous, though they are continued to the end of the tail.
Though the tail tapers it does not end in a point, but in a broad caudal fin,
the outer rays of which are confluent only in their basal half with the dorsal and
208
anal fins; these latter are well developed, the dorsal beginning immediately
behind the gill-opening. No pectorals.
The stomach forms a cul de sac of great length ; the air-bladder extends a
long way behind the vent.
Colours : dark lavender-grey ; dorsal and anal fins with basal half whitish
and free half blackish.
A ripe female, 22 inches long, from off the Travancore coast, 430 fathoms.
Regd. No. ~.
Named after Commander A. Dundas Taylor, formerly of the Indian Navy,
the founder and first chief of the present Marine Survey of India.
Sub-order Plectognathi.
Family Sclerodermi.
Teiacanthodes, Bleeker.
Triacanthodes, Bleeker, Act. Soc. Sc. Indo-Neerl. iii. Japan, iv. p. 37: Giinther, Cat. Fishes, VIII. p. 208.
Body elevated and compressed, with a short tail ; covered with small spiny
scales. No lateral line. Teeth very small, conical, close-set, from 14 to 20 in
the upper, and about twenty-two in the lower jaw, in a single series, often with
two or three in a second series. Anterior dorsal fin formed by from 4 to 6
strong spines. Ventrals formed by a pair of strong spines joined to the pelvic
bone, with one or two rudimentary rays.
Distribution : Japan ; Indian Seas.
168. Triacanthodes ethiops, Alcock.
Triacanthodes ethiops, Alcock, Journ. As. Soc. Bengal, Vol. LXIII. pt. 2, 1894, p. 137, pi. vii., fig. 6: Illustki-
TIONS OF THE ZOOLOGY OF THE INVESTIGATOR, FlSHES, PL. XV. FIG 9.
D. VI. 14-16. A. 14. P. 12-13. V. I. 1. C. 12.
Height of the body a little more than half the total without the caudal.
Head and body covered with small spiny scales, which are continued — much
reduced in size — on to the basal half of the fin-rays. In the young the spinelets
of the scales are embedded each in a fleshy papilla.
Eye very large.
Spinous dorsal well developed ; the first spine, which is the longest, is
rather shorter than the spine of the ventral fin, which is a third the length of the
body without the caudal. All the spines are rough with small barbs.
In the axil of eacli ventral spine is a small filamentous soft ray.
209
Colour in spirit : in the young blue-black, the papillae white : in large speci-
mens mottled black and white.
Largest specimen rather over 3- inches long.
Bay of Bengal, off Madras coast, 145 to 250 fathoms ; Andaman Sea, 185
fathoms.
Regd. Nos. 13709, 3™, 2Z2.
The peculiar delicacy of the tissues and the large goggle eyes proclaim this
fish to be an inhabitant of the depths.
Halimochirurgus nov. gen.
Body low and moderately compressed but not elongate, except the snout,
which is produced to form a long curved perfectly tubular organ, with the mouth
near the end, remarkably like the surgical instrument known as a catheter.
Tail short. Skin everywhere covered with small villiform and capillary spinelets.
No lateral line.
Mouth small, placed superiorly and transversely, near the end of the tubular
snout. A row of minute conical teeth in each jaw.
Gills pectinate. Gill-opening a narrow slit in front of the pectorals.
The spinous dorsal consists of two large spines. The soft dorsal is short
and almost opposite the anal.
Bach ventral consists of a huge spine articulating with the pelvic bone.
The mandibular symphysis is sharp and very prominent beyond the mouth,
but there is no barbel.
This curious fish is undoubtedly a very close relative of Triacanthus, but
has, at the same time, well-marked affinities with Monaeanthus and Anacanthus.
169. Halimochirurgus centriscoides, Alcock.
Halimochirurgus centriscoides, Alcock, Proc. Asiatic Soc, Bengal, Jnly, 1899, p. 78.
Illustrations of the Zoology of the Investigator, Fishes, pl. XXXI. fig. 3.
D. II. 13. A. 12. P. 11-12. V. I. C. 12.
Greatest body-height not quite half the length of the tubular snout, and
a fifth the total length without the caudal.
Snout tubular with an upward curve, ending in a point formed by the
mandibular symphysis. On its upper surface, a short distance from its end, is
the transverse mouth, well inside of which are two broad curtains of mucous
membrane — one connected with each jaw — forming an inner valvular orifice.
A row of minute distant conical teeth in each jaw.
Eye large. Nostrils small, in a hollow immediately in front of the eye.
27
210
Skin covered with granular, villiform, and capillary spinelets, which also
extend some way on to the fin-rays.
First dorsal spine situated above the ventrals, and equal in length to them
but hardly so stout : its length is about nine-tenths the greatest body-height.
Second dorsal spine about seven-tenths the greatest body-height.
Each ventral fin consists of a long stout movable spine having a movable
catch-joint with the pelvic bone. All the spines are rough.
Caudal not quite two-fifths the length of the snout.
Colour white, with a silvery sheen.
Length a little over 4^ inches.
Off Cape Comorin, 143 fathoms.
Regd. No. '— .
The delicacy of the tissues, the large eye, and the contents of the stomach
show that this fish came from a considerable depth and from the bottom.
211
Addenda [to Family STOMUTrDin].
Astronesthes, Richardson.
Astronesthes, Richardson, Ichth. Voy. Sulph., p. 97 : Giinther, Cat. Fishes v".,4>. 42-1 : Goode and Bean, Oceanic
Ichthyology, p. 105 : Jordan and Everraann, Fishes N. Amer., p. 586.
Phcenodon, Lowe, Proc. Zool. Soc, 1850, p. 250.
" Body rather elongate and compressed, scaleless, with the vent situated at no great distance
from the caudal fin. Head rather compressed, with the snout short and with the cleft of the
mouth wide. Teeth pointed, unequal in size, two pairs in the upper jaw and one in the lower
being long curved canine teeth : maxillary finely and subequally denticulated : vomer with a pair
of small fangs: palatines with a single series of small pointed teeth, similar to those of the
tongue. Eye of moderate size. A fleshy barbel is suspended from the centre of the hyoid region.
Dorsal fin rather long, opposite the interspace between the ventrals and the anal : caudal forked :
pectoral and ventral fins well developed. Series of luminous dots run along the lower side of the
head body and tail. Gill-opening very wide, the outer branchial arch with minute gill-rakers :
pseudobranchise none : air-bladder none. Stomach caecal " (Giinther).
Distribution : Atlantic : Indo-Pacific.
122 a. Astronesthes sp.
Illustrations of the Zoology of the Investigator, pl. XXXV. fig. 3.
A very small but perfect specimen of Astronesthes was taken, off the Travancore coast, in
224-284 fathoms.
It does not agree with any of the described species, but is nearest related to A. niger, from
which indeed it seems to differ only in having (1) the ventrals placed further back, (2) the anal
fin shorter — consisting of only 12 rays, and (3) the barbel very much thicker, and of nearly equal
thickness throughout.
As it is obviously immature I do not describe it : possibly it is a young Astronesthes niger.
Synopsis of the Indian genera of Stomiatidae.
[To be substituted for the Synopsis on p. 146.]
I. Hyoid barbel free : pectorals well developed : teeth on the palatines and
vomer : —
1. Body elongate: skin mapped into hexagonal areolae: dorsal and
anal fins opposite to one another and situated close to the caudal ... Stomias.
2. Body of moderate length : skin perfectly smooth and scaleless :
dorsal fin above the space between the ventrals and the anal ... Astronesthes.
II. Hyoid barbel attached to the mandibular symphysis: skin smooth and
scaleless : —
1. Pectoral rays reduced in number : no teeth on the palate ... Malacosteus.
2. Pectorals absent : teeth on the palatines ... ... ... Photostomias.
INDEX.
Page.
I
'age.
acanthonotus (Investigator)
„ (Nemichtliys)
AcANTHOPTERYGII
... 190
... 190
... 22
Arnoglossus
Astronesthes
Ateleopodidj; ...
74
,75,
124
211
122
Achirus
... 124
Ateleopus
122
Acropoma philippinense
affinis (Halosaurus)
... 24
... 185
,, indicus
atramentatus (Symphurus) ...
123
133
„ (Leptoderma)
„ (Stomias)
... 182
... 146
atrarius (Xenomystax)
atratus (Odontostomus)
206
167
Alcockia
... 90
atricolor (Bathypterois)
157,
159
Alepocephalid^e
... 168
atrox (Photostomias)
...
150
Alepocephalus ... 168,
„ bicolor
169,
169,
174, 180
170, 181
„ (Thaumastomias)
Aulastomomorpba
168,
178,
150
182
Blanfordi 169,
171,
172, 181
,, phospborops
178
,, edentulus
169, 172
aurora (Caprophonus)
...
44
Amblyopus
,, arctoceplialus
... 72
... 72
Bassogigas
Bassozetus
7^
1,83,
7S
90
Ammodytes
74, 77
,, glutinosus
88
Anacanthini
... 74
Bathyclupea
39
„ Gadoidei
... 74
„ Hoskynii
...
40
„ Pleuronectoidei
... 124
Bathycongrus macrocercus
198
Anacanthus
... 209
„ musteliceps
199
anago (Congromuriena)
... 197
„ nasicus
198
anguilliformis (Halosaurus)
184, 186
,, squaliceps
...
197
Anguillina
189, 195
Bathygadus
105,
120
Antennarius
51, 52
„ cavemosus
117
Antigonia
... 44
„ furvescens
120,
121
,, capros
... 44
„ longifilis
120,
121
Aplioristia
124, 130
„ melanobranchus ...
...
122
Gilesii 130,
131,
132, 133
„ multifilis
...
120
,, septem-striata
130, 132
Batht/onus glutinosus
...
88
„ trifasciata
130, 133
Bathypercis platyrhynchus
...
4S
,, Wood-Masoni
130, 131
Bathypterois 49,
152,
156,
157
apus (Platytroctes)
... 177
„ atricolor
157,
159
arctoceplialus (Amblyopus)...
... 72
,, dubitis
159
argenteum (Glyptophidium)
93, 94
,, Guntheri
...
157
argyreum (Melanostoma)
24, 25
,, insularum
157,
158
argyropastus (Pliysiculus) ...
76, 77
Bathyseriola
...
43
Argyropelecus ... 134,
135,
, 136, 137
„ cyanea
43
„ hemigymnus ...
... 135
Batkytroctes
168,
173,
17.S
Page.
Page.
Batnytroctes macrolepis
... 174
cavemosus (Hymenocephalus)
117, 118
„ microlepis
... 174
„ (Mystaconurus) ...
... 117
„ squamosus
... 173
centriscoides (Halimocbirurgus)
... 209
Batoidei
... 17
Centropborus
... 13
Bembrops
... 48
,, calceus
... 14
„ caudimacula
48, 50
,, foliaceus
... 14
„ gobioides
... 50
„ obscurus
... 14
,, platyrhynchus
... 48
,, Rossi
... 13
bengalensis (Proruethicbtliys)
... 42
Centropristis investigatoris
... 23
,, (Thyrsites
... 42
Centroseylliuni ...
13, 14
Benthobatis
... 17
,, ornatum
... 14
,, Moresbyi
... 18
Ceratias
52, 56
Bertcidj;
... 31
„ (Diceratias) bispinosus
... 56
bicolor (Alepocepbalns)
169, 170, 181
Champsodon
46, 51
bispinosus (Ceratias)
... 56
„ voras
... 47
,, (Diceratias)
... 56
Cbascanopsetta ...
124, 125
Blanfordi (Alepocepbalus) 169,
171, 172, 181
,, lugubris
... 125
boa (Stomias)
... 144
Cbanliodus
135, 143
Boopsetta
124, 126
„ Macouni
... 144
„ maculosa
... 127
„ pammelas
... 145
,, prselonga
126, 127, 128
„ Schneideri
... 141
,, umbrarum
... 126
„ setinosus
... 144
Brachyplenra
... 124
,, Sloanii
144, 145
bracbysoma (Diplacantbopoma]
I 98, 100
Chaunax
52, 5S
Bregmaceros
... 75
„ pictus ...
... 58
,, Macclellandii ...
... 75
Cbelidoperca
... 22
Brepbostoma
... 26
„ investigatoris ...
... 23
„ Carpenteri
... 26
Cbiasmodus
50, 51
brevirostris (Macrurus)
... Ill
„ niger
... 50
Brotula
77, 78
Cbloroplitbalmus
152, 153, 155
bucepbalus (Dysomma)
... 192
,, corniger ...
... 155
calceics (Centrophorus)
... 14
Chondroptertgii
... 13
Callionymus
... 73
Cilharichthys
... 124
„ carebares
... 73
coccinea (Halieutsea)
60, 61
„ kaiauus
... 74
Ccelorhyncbus ...
105, 106
Callnrhyncbus, egg-capsule of
... 22
Coloconger
189, 195
canescens (Scyllium)
... 16
,, raniceps
... 196
Caprophonus aurora
... 44
cometes (Gobius)
... 71
capros (Antigonia)
... 44
Gongrelhts guttulatus
... 199
Carangid.e
... 43
Cougromursena
189, 196, 202
Carchariidj:
... 13
anago
... 197
carebares (Callionymus)
... 73
„ guttulata
197, 199
carinicanda (Halosaurichthys)
... 187
„ longicauda
... 198
„ (Halosauius) ...
184, 187, 188
,, macrocercus ...
197, 198
Carpenteri (Brepbostoma) ...
... 26
,, musteliceps ...
197, 199
„ (Cynoglossus) ...
... 133
„ nasica
197, 198
Catai-hracti
... 68
,, squaliceps
197, 198, 199
caudimacula (Bembrops)
48, 50
conjugator (Neobytbites) ...
80, 81
cavtmosus (Bathygadus)
... 117
corniger (Chloropbtbaluius) ...
... 155
Page.
corythasolum (Diplophos)
142
corytheeolus (Pboticbtbys) ...
142
CorriDM
67
crassiceps (Uranoscopus)
...
45
cyanea (Batbyseiiola)
...
43
,, (Solea)
...
130
Cyclothone
135,
139
„ elongata
...
139
,, lusra
...
141
„ microdon
141
Cynoglossus
124,
133
„ Carpenteri
133
Ctttidj;
...
44
Darwinii (Traebicbtbys)
35,
37
Dermatoius
78
,89,
90
„ melampeplus
90,
92
„ melanocepbalus...
90
,91,
92
,, trichiurus
00;
,91,
92
diapbana (Sternoptyx)
...
136
Dibranchus
52,
62
„ micropus
...
63
„ nasutus
...
63
Diceratias
...
56
Dierolene
78,
85
„ intronigra
85,
,86,
87
,, multifilis
85,
86
„ nigricaudis
85,
87
DinematiehtJiys ...
...
78
Dinemus venustus
...
38
Diodon
.. .
58
Diplacantbopoma 78, 98, 102, 103, 104
,, bracbysoma 98, 100
„ raniceps ... 98, 101
„ Rivers-Andersoni98,99, 100, 101, 103
„ (Saccogaster) maculatum ... 102
Diplophos corythseolum ... ... 142
dubius ( Bathypterois) ... ... 159
Dumerilii (Myctopbum) ... 160, 162
Dysomma ... ... 188, 192, 193
Dysomma bucephalus ... ... 192
Dysomhina ... ... 188, 192
Dysominopsis ... . ... 189, 193
„ niucipara ... ... 193
edentulus (Alepocephalus) ... 169, 172
egg of Raja ... ... ... 22
egg-capsule of Callorbyncbus ... 22
elongata (Cyclotbone) ... ... 139
elongatum (Gonostoma) ... ... 139
elongatus (Stomias) ... 146, 147
Page.
Embiotocidse ... ... ... 100
engraulis (Myctopbum) ... 160, 161, 162
erimelas (Narcetes) ... ... 175
etbiops (Triacanthodes) ... ... 208
fasciolaris (Symphurus) ... ... 133
Fierasfer ... ... 77, 78
flabellispinis (Coeloi'hyncbus) ... 107
„ (Macrurus) ... ... 106
foliaceus (Centrophorus) ... ... 14
fragilis (Lamprogrammus) ... 95, 96
fumosa (Halieuteea) ... 60, 62
furvescens (Batbygadus) ... 120, 121
Gadim: ... ... 51, 74, 75
Gavialiceps ... ... 188, 191, 193
„ microps ... ... 191
,, tseniola ... ... 206
Gilesii (Aphoristia) 130, 131, 132, 133
glomerosus (Onirodes) ... ... 57
„ (Paronirodes) ... ... 57
glutinosus (Bassozetus) ... ... 88
„ (Bathyonus) ... ... 88
Glyptophidium ... ... 78, 92, 93, 95
„ argenteum ... 93, 94
„ macropus ... 93, 94
G0BHD.E ... ... ... 71
gobioides {Bembrops) ... ... 50
„ (Hypsicometes) ... 48, 49
Gobius ... ... ... 71
„ cometes ... ... ... 71
Gonostoma ... ... ... 145
„ elongatum ... ... 139
„ microdon ... ... 141
gracilimaims (Lopbius) ... 53, 54
Guntberi (Batbypterois) ... ... 157
„ (Scopelarcbus) ... ... 153
,, (Setarcbes) ... ... 28
„ (Xenodermicbtbys) 180, 181
guttulata (Congrellus) ... ... 199
„ (Congroniura?na) ... 197, 199
Halicmetus ... 52, 65
„ ruber ... ... 66
Halieutsea 51, 52, 60, 65, 66
„ coccinea ... 60, 61
,, fumosa ... 60, 62
,, nigra ... ... 60
,, stellata ... 60, 61, 62
Halimocbiiurgus ... ... 209
„ centriscoides... ... 209
Halosaurichthys carinicauda ... ... 187
Page.
Page.
Halosadrid*
... 183
italicus (Mystaconurus)
, 117
Halosaurus
95, 183
japonica (Polymixia)
38
, 39
„ affinis
... 185
japonicus (Monocentris)
32
„ anguilliformis
184, 186
Johannis-Davisi (Raja)
19
, 21
,, carinicauda
184, 187, 188
kaiana (Solea) ...
130
„ Hoskynii
... 184
kaianus ( Callionymus)
74
,, mediorostris
184, 185
„ (Uranoscopus)
...
46
„ nigerrimus
184, 188
Kurtidj;
39
„ parvipennis
184, 186, 187
LaBops
124
, 128
Halyi (Peristethium)
... 68
„ macropbthalmus
128
Harpodon
152, 154
Ik vis (Macrurus)
. * .
119
„ squamosus
... 154
„ (Malacocephalus)
...
119
hemigymnus (Argyropelecus)
... 135
Lamnidse
13
„ (Sternoptyx)
... 135
Lamprogram m us
78,
94
hemisticta ( Trigla)
... 67
„ fragilis
95,
96
Hephthocara
78, 103
» niger
95,
96
„ simum
... 103
Leei (Symphurus)
. . .
133
heterolepis (Mystaconurus) ...
117, 118
Lepidotrigla
...
67
hexanerna (Sebastes)
... 27
,, spiloptera
67
Hextii ( Macrurus )
108, 113
„ „ var. Ion
gipinnis ...
68
„ (Tauredophidium) ...
... 97
Leptoderma
168,
182
hispidum (Scylliurn)
15, 16
,, affinis
182
hispidus ( Clialinurus)
... 110
,, viacrops
182,
183
Holocentrum
... 32
lepturus (Uroconger)
■ ■ •
201
Hoplostethus
32, 33, 35
Lioscorpius longiceps
. ..
28
„ mediterraneum...
... 34
longicauda (Congromursena) ...
198
Hoakynii (Bathyclupea)
... 40
longiceps (Lioscorpius)
• ••
28
„ (Halosaurus)
... 184
longifilis (Bathygadus)
120,
121
„ (Macrurus)
109, 116
„ (Ilymenocephalus) ...
120
Hymenocephalus eavemosus ...
117, 118
longipinnis, var. (Lepidotrigla) spiloptera
68
,, italicus
... 118
Lopbius
...
52
,, longifilis
... 120
,, gracilimanus
53,
54
Hypsicometes gobioides
48, 49
„ indicus ...
53,
54
indicua (Ateleopus)
... 123
„ lugubris ...
53,
55
„ (Lophiua)
53, 54
„ mutilus ...
53, 54, 55,
56
„ (Malacosteug)
... 149
„ setigerus ...
54
„ (Myctophum)
... 161
lopbotes (Macrurus)
109,
116
inennis (Minous)
30, 31
Lowei (Polymixia)
38,
39
infans (Nemichthys)
190, 192
lugubris (Chascanopsetta) ...
125
instilarum ( Batliypterois) ...
157, 158
„ (Lopbius)
53,
55
intermedius (Tracliichtliya)...
... 36
lusca (Cyclothone)
141
intronigra (Dicrolene)
85, 86, 87
lutea (M;iltliopsis)
...
64
Investigator acanthonotus
... 190
Macclellandii (Bregmaceros)
• • •
75
investigatoria (Centropristis)
... 23
Macouui (Cltanliodus)
...
144
„ (Chelidoperca)
... 23
macrocercus (Bathycongrus) ...
...
198
„ (Macrurus) 108,
109, 110, 111,
„ (Congromursena)...
J 97,
198
112,11s
;, 114, 115, 116
macrolepidotua (Neoscopelus)
...
164
,, (Peristethium)
68, 70
„ (Scopelus) ...
...
164
italicus (LTymenocephalus)
... 118
macrolepis (Bathytroctes) ...
*•«
174
rnacrolophus (Maerurus)
macrophthalmus (Laeops) ..
„ (Scianectes)
macrops (Leptoderma)
Page.
109, 115
... 128
... 128
182, 183
(Neobythites) 79, 80, 81, 82, 83, 84
macropus (Glyptophidium) ... 93, 94
maculatum (Diplacanthopoma) ... 102
,, ( Saccogaster) ... ... 102
maculosa (Boopsetta) ... ... 127
(Pcecilopsetta) ... ... 127
Macrurid.£ ... ... 74, 75, 104
Maerurus .., ... 26, 105
Maerurus [sub-genus] ... ... 108
„ brevirostiis ... ... Ill
„ (Chalinurus) hispidus ... 110
„ (Ccelorhynchus) flabellispinis 107
,, ( ,, ) parallelus ... 106
,, ( » ) quadricristatus 106
Maerurus flabellispinis ... ... 106
Hextii ... 108, 113
Hoskynii ... 109, 116
„ investigatoris 108, 109, 110, 111,
112, 113, 114, 115, 116
„ laevis ... ... ... 119
lophotes ... 109, 116
„ (Malacocephalus) laevis ... 119
,, macrolophus ... 109, 115
„ (Mystaconurus) cavernosus ... 117
( „ ) heterolepis 117, 118
„ ( „ ) italicus ... in
„ nasutus ... 108, 111
„ parallelus ... ... 106
„ petersoni ... 108, 110
„ polylepis ... 108, 112
maerurus (Pterois) ... ... 30
Maerurus pumiliceps ... 108, 113
„ quadricristatus ... 106, 107
„ semiquincunciatus 108, 111, 112
„ sesquicunciatus ... ... U2
„ Wood-Masoni ... 109, 114
Malacocephalus ... 105, 118
» ]®vis ... ... H9
Malacosteus ... ... 146, 148
Malacosteus indicus ... ... 149
» "iger ... ... 149
sp. ... ... 149
Maltke ... ... ... 65
Malthopsis ... ... 52, 64, 66
lutea... ... ... 64
Page.
mamillidens (Raja) ... ... 19
mediorostris (Halosaurus) ... 184, 185
mediterranea (Sternoptyx) ... ... 135
mediterraneum (Hoplostethus) ... 34
melampeplus (Dermatorus) ... 90, 92
Melamphaes ... ... 32, 37
,, mizolepis ... ... 37
melanobranchus (Bathygadus) ... 122
nielanocephalus (Dermatorus) 90, 91, 92
melanosticta (Solea) ... ... 130
Melanostoma argyreurn ... 24, 25
microdon (Cyclothone) ... ... 141
„ (Gonostoma) ... ... 141
microlepis (Bathytroctes) ... ... 174
microps (Gavialiceps) ... ... 191
micropus (Dibranchus) ... ... 63
Minous ... ... 30
„ inermis ... 30, 31
Mixonus ... ... ... 88
mizolepis (Melamphaes) ... ... 37
,, (Scopelus) ... ... 37
Monacanthus ... ... 209
Monoceutris ... 32, 34
,, japonicus ... ... 32
Moresbyi (Benthobatis) ... ... 18
mucipara (Dysommopsis) ... ... 193
multifilis (Bathygadus) ... ... 120
„ (Dicrolene) ... 85, 86
,, (Paradicrolene) ... ... 86
Mdr,enesocina ... 189, 203
Mur.snid.£: ... ... 188
Murrayi (Raja) ... ... 21
mustelicep's ( Bathycongrus) ... ... 199
„ (Congromursna) 197, 199
mutilus (Lophius) ... 53, 54, 55, 56
Myctophum (sub-genus) ... ... 160
Myripristis ... ... 32
Mystaconurus (sub-genus) ... 105, 117
Narcetes ... ... 168, 174
„ erimelas ... ... 175
nasica (Congromursena) ... 197, 198
nasicus (Bathycongrus) ... ... 198
nasutus (Dibranchus) ... ... 63
„ (Maerurus) ... 108, 111
nebulosus (Stomias) ... ... 146
Nematonus ... ... ••• 88
Nemichthyina ... ... 188, 189
Nemichthys ... 188, 189, 191, 193
acanthonotus ... ... 190
VI
Page.
Page.
NemichtJiys infant
190,
192
Peristethium
...
68
Nemobrama Wtbbii
38
„ Ealyi
...
, 68
Neobythites
78, 79, 85,
88
„ investigatoris ...
68,
70
,, (Bassogigas) pterotus
...
83
„ Rivers- A ndersoni
68
„ conjugator
80,
81
„ serrulatum
68:
, 69
,, maciops 79, 80
, 81, 82, 83,
84
Petersoni (Macrurus)
108,
110
„ uigripinnis
80,
82
philippinense (Acropoma)
...
24
,, pterotus
80,
83
philippinensis (Synagrops) ...
24
„ squaraipinnis ...
80,
84
phosphorops (Aulastomoniorpha)
178
,, steatiticus
80,
82
Pboticbthys
135,
141
Neoscopelus
152,
164
„ corytheeolus
...
142
„ macrolepidotus
...
164
Pbotostomias
146,
149
Nettastoma txniola
...
206
,, atrox
150
Nettenchelys
189,
207
Physiculus
75:
, 7fi
Taylori
...
207
„ argyropastus
76,
77
niger (Chiasmodus)
...
60
„ roseus
7€
1 77
„ (Lamprogranimus)
95,
96
Phtsostomi
...
134
„ (Malacosteus) ...
149
pictus (Cbaunax)
58
m'gerrimus (Halosaurns)
184,
188
pinnatus (Synaphobranchus)
...
195
nigra (Halieuteea)
...
60
Plagiostomata
...
13
nigricaudis (Dicrolene)
85,
87
Plagusia
...
124
,, {Paradicrolene) ...
...
87
platyrhynchus (Bathypercis) ...
...
48
nigripinnis (Neobythites) ...
80,
82
„ (Bembrops)
...
48
nobilis (Polymixia)
...
38
Platytroctes ... ... 168
,176,
178
Notacanthus
...
190
,, apus
177
Notidanidm ... ,.t
...
13
Plectognathi
208
•nudus (Porogadus)
91
Pleuronectidj; ...
124,
125
obscurtcs (Centrophorus)
...
14
Poecilopsetta maculosa
...
127
Odontostomus
152,
153,
166
,, preelonga
...
126
,, atratus
...
167
Polyipnus
134,
137
Oniiodes
52,
57
,, spinosus
138
,, glomerosus
...
57
polylepis (AJacrurus)
108,
112
Ophidiice
74, 7£
',77,
78
Polymixia
32,
38
ornatum (Centroseyllium) ...
...
14
„ japonica
38,
39
,, (Paracentroscyllium)
...
14
„ Lowei
38,
39
(xryrhynchus (Raja)
22
,, nobilis
...
38
pammelas (Chauliodus)
145
Pomatomus
...
26
Parucentroscylliuin ornatum ...
...
14
Ponerodon vastator
...
50
Paradicrolene nmltifilis
86
Porogadus nudus
...
91
„ nigricaudis
...
87
„ subarmatus
...
92
„ Vaillanti
85
Powelli (Raja) ...
19,
20
parallelus (Ccelorhyuchus) ...
106
prajlouga (Boopsetta) ... 126,
127,
128
„ (Macrurus)
106
„ (Pcecilopsetta)
...
126
Paralepis
153
prometheoides (Thyrsites)
...
42
Parascombrops pellucida
24
Prometbicbtliys ...
...
41
Paronirodes glomerosus
...
57
Promyllautor
189,
202
parvipennis (Halosaurus)
184,
186,
187
„ purpurcus
...
202
Pediculati
51
Psettodes
124
pellucida (Parascombrops)
...
24
Psettylis
...
124
Page.
Page.
Pseudorhombus ...
...
124
Scopelengys ... ... 152
160,
165
Pterois
...
30
„ tristis
...
166
„ macrurus
30
Scotelidj:
151,
152
pterotus (Bassogigas)
83
Scopelus ... 152, 153
160,
164
„ (Myctophum)
161,
162
,, macrolepidotus
164
„ (Neobythites)
80,
83
„ mizolepis
37
,, (Scopelus)
...
46
,, Dumerilii
160,
162
pumiliceps (Macrurus)
108,
113
,, engraulis ... 160
161,
162
purpureas (Promyllantor) ...
...
202
„ indicus
161
Pycnocraspedum ...
...
78
„ pterotus
161,
162
„ squamipinne...
...
84
„ pyisobolus
161,
163
pyrsobolus (Myctophum)
161,
163
Scorpmna remigera
28
quadricristatus (Ccelorhynchus)
106
ScouFMrnvM
...
27
„ (Macrurus) ...
106,
107
SCTLLIID^!
13,
15
quagga (Scyllium)
...
17
Scyllium
...
15
Raja
...
19
,, canescens
16
„ egg of
...
22
„ hispidum
15,
16
„ Johannis-Davisi
19,
21
quagga
17
,, mamillidens
...
19
Sebastes
...
27
„ Murray i
...
21
„ hexanema
...
27
„ oxyrliynclms...
22
Selachoidei
13
„ Powelli
19,
20
semiquincunciatus (Macrurus) 108
Ill,
112
Rajids
19
septem-striata (Aphoristia) ...
130,
132
rauiceps (Coloconger)
...
196
SERRAN1D3!
22
,, (Diplacanthopoma) ...
98,
101
serrulatum (Peristethium ) ...
68,
69
remigera (Scorpsena)
...
28
sesquicunciahis (Macrurus) ...
112
~RMnodontid.se
...
13
Setavcbes
...
28
Rhomboidichthys ...
...
124
„ Guntheri
...
28
Rivers-Andersoni (Diplacanthop
nma) 98
99,
setigerus (Lophius)
.. .
54
100, 101
103
setinosus (Chauliodus)
.**
144
„ (Peristethiurn)
...
68
Sigmops stigmaticus
139
roseus (Physiculus)
76,
77
simum (Hephthocara)
103
Rossi (Centrophorus)
...
13
sloauii (Chauliodus)
144,
145
ruber (Halicmetus)
...
66
socialis (Xenodermichthys) ...
181,
182
rubescens (Hypsinotus)
...
44
Solea
124,
129
Saccogaster
78, 102,
104
,, (Achirus) umbratilis ...
...
129
Samaris
...
124
,, cyanea
...
130
Saurencbelys
189,
206
„ Tiaiana
...
130
„ tasniola
206
„ melanosticta
...
130
Saurida
...
152
spiloptera (Lepidotrigla)
...
67
Sauromursenesox
189,
203
Spinacids
...
13
„ vorax
203
spinosus (Polyipnus)
...
138
Saurtis
152,
153
squaliceps (Bathycongrus)
197
Schneideri (Chauliodus)
...
144
,, (Congromursena) ...
197,
198
„ (Stomias)
...
144
squamilaterus (Xenodermichthys)
180,
181
Scianectes macrophthalmus ...
...
128
squamipinne ( Pycnocraspedum )
...
84
SCLERODERMI
...
208
squamipinnis (Neobythites)...
80,
84
Scopelarchus
152
squamosus (Bathytroctes) ...
...
173
,, Guntheri
...
153
„ (Harpodon)
...
154
Till
Page.
Page.
steatiticus (Neobythites)
80,
82
Trachichtbys Darwinii
35,
37
stellata (Halieutsea)
60, 61,
62
,, intermedius
...
36
STERNOPTYCHID.E
134,
145
Trachinidj;
...
45
Sternoptyx
... 134, 136,
137
Triacanthodes
■ ••
208
„ diaphana
...
136
„ etbiops
208
„ hemigymnns
...
135
Triacanthus
...
209
„ mediterranea
...
135
TRICHIUKIDiE
...
41
stigmaticus (Sigmops)
...
139
tricbiurus (Dermatorus)
90, 91,
92
Stomias
146,
148
trifasciata ( Aphoristia)
130,
133
,, aflinis ...
...
146
Trigla
67
„ boa
...
14i
,, bemisticta
67
„ elongates
146,
147
tristis (Scopelengys)
166
,, nebulosus
..
146
trucidans (Xenomystax)
...
205
Stomias Sehneideri
...
144
umbrarum (Boapsetta)
12*
Stomiatibj:
... 145, 146,
211
umbratilis (Achirus)
129
subarma his ( Poroga dus )
...
92
„ (Solea)
129
Symphurus atramentatns
...
133
Uranoscopus
45
„ fasciatus
133
„ crassiceps
45
„ Leei
...
133
„ kaianus
46
Synagrops
...
24
Uroconger
189,
200
„ philippinensis
...
24
„ lepturus
201
Stxaphobkanchina
189,
194
,, vicinus
200,
201
Synapbobranchu s
189,
194
Vaillanti (Paradicrolene)
85
;, pinrtatus
...
194
vastator (Ponerodon)
50
Synaptura
124
venustus (Dinemus)
38
taauiola (Qavialiceps)
...
206
vicinus (Uroconger)
200,
201
,, (Saurencbelys
...
206
vorax (Cbarapsodon)
...
47
Taureclopbidium
78,
96
„ (Sauromimenesox)
203
„ Hextii
97
Webbii (Nemobrama)
38
Taylori (Nettencbelys)
•••
207
Wood-Masoui (Apboristia) ...
130,
131
Teleostei
...
22
(Macrurus) ...
109,
114
Tetrodon
. .. . ..
58
Xenoderniichthys
168,
179
Thaumastomias atrox
...
150
„ Guntheri
180,
181
Thyrsites
41
„ socialis
181,
182
,, prometheoides
...
42
;, squamilaterus
180,
181
„ (Promebchtbys)
bengalensis ...
42
Xenomystax
189,
204
TORPEDINIDjE
17
,, atrarius
206
Tracliicbthys
32,
35
„ trucidans
...
205
LIST OF PUBLICATIONS ISSUED BY TL CJSTEES OF THE INDIAN MUSEUM.
Bs. As. P.
Account of the Deep-Sea Brachvura collected by the R.I.M.S. Investigator. By
A. Alcock, M.B., C.M.Z.S., F.G.S. .'.. ... ... ' ... 6 0 0
Account of the Deep-Sea Madreporaria collected by the R.I.M.S. Investigator. By
A. Alcock, M.B., C.M.Z.S., F.G.S. ... ... ...c. ..'. 4 0 0
Catalogue of Archaeological Collections in the Indian Museum. Parts I and II. By
L Anderson, M.D., F.R.S., &c. ... ... ... ... ■ ... 4 12 0
Catalogue of Coins of the Indian Museum, Parts I to IV. By C. J. Rodgers, M.R.A.S.,
M.N.S. ... ... ... ... ... ... ... 24 0 0
Catalogue of the Indian Deep-Sea Fishes in the Indian Museum. By A. Alcock, M.B.,
C.M.Z.S., F.G.S. ... ... ... ... ... ... 5 0 D
Catalogue of Mammalia in the Indian Museum, Part I. By 3. Anderson, M.D., F.R.S.,
Part II. By AY. L. Sclater, M.A., F.Z.S. ... ... ... ... 6 0 0
Catalogue of Mantodea in the Indian Museum, Parts I and II. By J. Wood-Mason,
F.Z.S...&C. ... ... ... ... ... *... 2 0 ('
Catalogue of Moths of India, Parts I to VII. By E. C. Cotes, and C. Swinhoe, F.L.S.,
F.Z.S., &c. ... ... ... ... ... ... 5 12 0
Echinoderma of the Indian Museum : Ophiuroidea collected by the R.I.M.S. Investigator.
By K. Koehler ... ... ... ■ ... ... ... 10 0 0
Figures and Descriptions of Nine Species of Squillida? from the Collection in the Indian
Museum. By J. Wood-Mason, F.Z.S., &c. Edited by A. Alcock ... ... 2 0 0
Guide to the Zoological Collections Exhibited in the Invertebrate Gallery of the Indian
Museum. By A. Alcock, M.B., C.M.Z.S., F.G.S. ... ' ... ... 0 10 0
Guide to the Zoological Collections Exhibited in the Reptile and Amphibia Gallery of the
Indian Museum. By A. Alcock, M.B., C.M.Z.S., F.G.S. ... ... ... 0 3 0
(laud List of Mollusca in the Indian Museum, Parts I, II, and Fasciculus E. By G.
Nevill, C.M.Z.S., &c. Index, Parts I and II. By TO. Theobald ... ... ... 7 4 0
List of Batrachia in the Indian Museum. By W. L. Sclater, M.A., F.Z.S. ... ... 1 (> o
List of Snakes in the Indian Museum. By W. L. Sclater, M. A., F.Z.S. ... ... 1 0 0
Monograph of the Asiatic Chiroptera and Catalogue of the Species of Bats in the Indian
Museum. By G. E. Dobson, M.A., M.B., F.R.S., &c. ... ... ... 3 0 b
Monograph of the Oriental Cicadidab, Parts I to VII. By W. L. Distant, F.E.S. ... £.2 2 6
Tin above can be obtained from the Superintendent of Jhe Indian Museum, Calcutta, and from
Messrs. Friedlander Sf Sun, 11, Garlsirasse, Berlin.
Other Publications sold by tub Superintendent of the Indian Museum (aZ.w obtainable from Messrs. Friedlandei &
DlRECIOK OP THE BOYAL INDIAN MARINE.
Illustratious of the Zoology of the R.I.M.S. Investigator, 1892. Fishes, Plates i-vii. Crustacea. Plates' i-v
,, 1894. Fishes, Plates viii-xiii. Crustacea, Plates vi-viii. Echinoderma, Plates i-iii
1895. Bchinoderma, Plates iv-v
L895. Pishes, Plates xiv-xvi
., Is.).*, Crustacea, Plates ix-xv
,, 1896. Crustacea, Plates xvi-xxvii ... ... ... ... ... ■
189' Pishes, Plate xvii. Crustacea, Plates xxviii-xxxii. Mollusca, Plates i-vi
Pishes, Plates xviii-xxiv. Crustacea, Plates xxxiii-xxxv. Mollusca, Plates vii-viii
., 1899. Pishes, Plates xxv-xxvi. Crustacea, Plates xxxvi-xlv ... ...
ft.
A.
P.
... 1-'
0
0
... J2
0
0
'1
0
0
... 3
0
0
7
0
0
... VI
0
0
... 12
0
0
... 12
0
0
... 12
0
0
QL622 A4 1899
catalog hull
Harvard MCZ Library \rv> I