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DYNAMIC  THEORY  OF  FISHERIES  ECONOMICS  —  II; 
DIFFERENTIAL  GAME  THEORETIC  APPROACH 


T.  Takayama 


#441 


College  of  Commerce  and  Business  Administration 

University  of  Illinois  at  U  r  ba  n  a  -  C  ha  m  p  a  ig  n 


FACULTY  WORKING  PAPERS 
College  of  Commerce  and  Business  Admlnistratix>n 
University  of  Illinois  at  Urbana-Champaign 

October  12,  1977 


DYNAMIC  THEORY  OF  FISHERIES  ECONOMICS  —  II; 
DIFFERENTIAL  GAME  THEORETIC  APPROACH 

T.  Takayama 

#441 


■i  l-i  ;•.,:,»...  •:  '•'■•■:',■ 


■i.'.iK 


October  2nd,  1977 


DYNAMIC  THEORY  OF  FISHERIES  ECONOMICS— II; 
DIFFERENTIAL  GAME  THEORETIC  APPROACH 

T.  Takayama 
University  of  Illinois 


Abstract 


A  two  countries-single  speriss  dynamic  optimizacion  problem  in  fisheries 
is  formulated  in  differential  game  theoretic  framework.  The  Cournot-Nash  solution 
concept  is  introduced  and  two  strategies  in  arriving  at  the  Cournot-Nash  solutions; 
are  discussed.   These  strategies  are  (1)  closed-loop  catch  strategy  and  (2)  open- 
loop  catch  strategy. 

A  quadratic  benefit  function  is  then  used  as  the  objective  function  of  each 
country  that  is  to  be  maximized  subject  to  a  linear  population-catch  dynamics.   Th'^ 
solutions  are  then  derived  for  each  catch  strategy  and  their  implications  are  d^''- 
cussed. 

The  major  conclusions  are  as  follows: 

(i)   The  stable  optimal  catches  show  that  each  country  must  regulate 
the  tonal  catch  of  her  fleet, 
(ii)   The  mash-size  snould  also  be  regulated. 
(XXI)      The  opeu-locp  optimal  catch  strategies  exist  only  when  two 
countries'  future  discount  rates  are  the  same, 
(iv)   The  higher  the  future  discount  rates,  the  more  fish  vri.ll  be 

caught  now  and  in  the  near  future,  and  the  slower  the  conver- 
gence of  this  fish  population  to  the  desired  level. 


October  2nd,  1977 


DYNAMIC  THEORY  OF  FISHERIES  ECONOMICS— II: 
DIFFERENTIAL  GAME  THEORETIC  APPROACH* 

T.  Takayama** 
Introduction 

Even  after  the  imposition  of  the  two  hundred  mile  territorial  waters 
limit  by  many  countries,  there  are  many  fish  species  being  caught  in  the  open 
international  waters.  The  two  hundred  mile  territorial  waters  reg;ulation  is  a 
revelation  of  inherent  conflicts  among  a  number  of  countries  whose:  fishing  fleets 
are  after  one  or  more  fish  species  of  their  common  interest  and  need.   Of  course, 
the  regulation  does  not  solve  all  the  problems  of  the  host  country  in  relation  to 
the  desired  catch  o?  fish  species,  but  alleviates  the  intermediate  long-run  burden 
of  driving  the  fich  population  to  an  undesirably  low  level  close  to  extinction  [2]. 

The  cod  war  between  the  United  Kingdom  and  Iceland  is  another  example  of 
international  conflicts  in  the  field  of  fisheries  economies.   Can  they  resolve 
this  type  of  conflict  in  any  manner  whatsoever?  This  is  an  important  question  in 
both  theory  and  politics.   In  this  paper  we  plan  to  partially  answer  this  question 
from  a  theoretical  point  of  view. 

As  a  natural  axtension  of  the  dynamic  optimization  approach  related  to 
fisheries  economics  [1,  10,  11],  we  apply  the  differential  game  theory  to  a 
two  country-one  fish  species  situation. 

In  section  1,  the  fish  population  dynamics  and  general  properties  of  the 
differential  game  model  applicable  to  fisheries  economics  are  discussed.   Qua- 
dratic objective  fuiictionals  and  linear  fish  population  dynamics  cases  are  then 
solved  for  the  Cournotr-Nash  equilibrium  and  various  implications  of  the  solution 

are  discussed  in  section  2.   In  the  last  section  we  discussed  future  research 

*This  project  is  partially  supported  by  the  Ford  Foundation  Grant,  lEO 
#750-0111. 

**Professor  of  Economics  at  University  of  Illinois,  Champaign-Urbana,  Illi- 
nois 61801.   He  is  grateful  to  his  colleagues,  professors.  Royal  Brandis,  H.  Brems, 
T.  Sawa,  M.  Simaan  for  their  encouragement  in  this  work.   He  alone  is  responsible 
for  any  error  in  this  paper. 


■'-;..!  rJfe 


•J    li! 


directions  in  this  field  and  conclude  with  some  positive  notes  for  future  research 
in  the  near  future. 

1.   Differential  Game  Formulation  of 
Fisheries  Economic  (Conflict)  Problem 

The  problem  the  fisheries  economicsts  face  is  that  of  determining  and 
recommending  the  fishing  intensity  that  will  maximize  the  economic  value  to  the 
consuming  societies  and  also  maximize  the  producers'  surplus  at  a  level  of  pro- 
duction in  perpetuity.   In  the  previous  paper,  we  took  the  stand  that  the  con- 
suming societies  can  be  looked  upon  as  one  society  with  one  market  for  a  fish 
species  and  also  the  producers'  supply  response  can  be  represented  by  a  single 
market  supply  function.   In  this  paper,  we  deal  with  two  countries  or  consuming 
societies  pursuing  their  individual  economic  value  or  goal  such  as  a  maximum  wel- 
fare or  profit  or  whatnot.   The  two  producer  societies  are  also  separated  in  the 
sense  that  the  cost  function  of  each  country  is  expressed  by  its  own  currency 
unit.   We  also  assume  in  this  paper  that  there  is  no  trade  of  fresh  fish  or  pro- 
cessed fish  products  between  the  two  countries. 

Thus,  country  1  tries  to  maximize  its  welfare  from  the  consumption  and 
production  (catch)  of  a  fish  species.   The  same  or  different  objective  may  hold 
true  for  country  2.   For  simplicity's  sake,  let  us  write  this  objective  function 
as  a  function  of  the  (recruitable)  fish  population,  p(t),  at  time  t  (^  0)  and 
the  intensity  of  catch  by  the  ith  country  fleet  x.(t)  at  time  t  (^0);  that  is, 


1 


/-S 


(1.1)       W^  =   /  e'''i^W^(p(t),x^(t))dt 


where 

r  denotes  the  rate  of  the  future  discount  by  country  i,  i  =  1,2.  W^(p(t), 
x(t))  denotes  the  welfare  or  benefit  or  profit  or  any  other  objective  that 


the  ith  country  tries  to  maximize,  which  depends  on  the  fish  popula- 
tion (to  a  lesser  extent)  and  the  catch  of  fish  by  the  country's  fleet 
for  its  own  consuming  society. 

Thus  far,  the  objective  function  of  each  country  is  completely  indivi- 
dualized.  This  is  a  clear-cut  departure  that  the  differential  game  theoretical 
approach  makes  from  the  ordinary  optimization  approach.   There  is,  however,  one 
common  object  between  them  that  is  to  be  carefully  observed;  that  is,  the  ocean  in 
which  the  population  of  a  fish  species  of  their  interest  grows  or  declines  in 
relation  to  the  fishing  intensities  of  fishermen  of  the  two  countries.   The  fish 
population  in  this  paper  is  considered  as  a  directly  observable  state  variable, 
and  the  catch  by  the  fishermen  in  each  country  is  considered  as  a  control  vari- 
able;  there  are  naturally  two  control  variables  in  a  two  countries-one  fish 
species  case. 

The  fish  population  (and  catch  interaction)  dynamics  is  written  as 

(1.2)  p(t)  =  f(p(t),  Xj^(t),t),  te[0,  T) 

where  p(t)  is  defined  as  before,  (recruitable)  fish  and  population  at 
time  t,  state  variable;  x(t)  denotes  total  intensity  of  catch  by  two 
countries;  t  denotes  actual  time;  and  T  is  the  terminal  time  greater 
than  zero  and  can  be  finite  or  infinite.   In  our  two  countries  model, 
the  total  catch  intensity  x(t)  can  be  divided  into  two  parts  as 
follows : 

(1.3)  x(t)  =  Xj^(t)  +  X2(t) 

where  x  (t)  is  defined  in  relation  to  (1.1)  and  denotes  the  Intensity 

X 

of  catch  by  the  ith  country  fishermen,  at  time  t,  i  =  1,2. 


The  simplest  dynamics  we  can  consider  is  the  linear  first-order  dynamics 


such  as 


(1.4)         p  =  a  +  bp  -  X 

where  we  assume  that  a  and  b  are  constants  and  most  likely 


(1.5) 


a  ^  0  and 
b  >  0  . 


The  Figure  1  below  shows  this  catch  fish  population  dynamics. 


-a/b  p 

Figure  1:   Catch-fish  population  djmamics;  linear  case. 


A  nonlinear  dynamic  case  is  also  of  great  theoretical  interest.   For 
instance,  the  Lotka  type  or  Voltera  type  quadratic  dynamics  [  5 ]  may  be  important 
in  application.   However,  we  confine  ourselves,  in  deriving  some  practical  infer- 
ences, to  the  linear  dynamics  case,  (1.4),  in  this  paper. 

In  Figure  1,  the  steady-state  locus  of  (p,  x)  combinations  is  shown  by 
p=0=a+bp-x  equation.  When  catch  is  zero,  the  minimum  sustainable  popula- 
tion (below  which  the  species  become  extinct)  is  -(a/b).   To  the  right  of  the 


"^The  dynamic  identification,  t  of  p(t),  x(t),  etc.,  will  be  omitted,  unless 
otherwise  stated  in  the  following  development. 


ii.1    ;e">lj3r 


d  •>:{/■.) 


.)    I'x    .  q)    ^o 


line,  the  population  (stock)  to  catch  relationship  is  favorable  to  increase  in 
fish  population.   To  the  left  of  the  line,  due  to  relatively  heavy  catches,  the 
fish  population  tends  to  be  zero. 

In  formulating  the  differential  game  problem  within  the  framework  of 
two  countries  and  one  fish  species,  let  us  employ  as  general  a  formulation  as 
possible.   The  problem  is  clearly  divided  into  the  following  two  subproblems: 

Fisheries  Differential  Game  Problem  (F  D  G  P) ; 

(i)   Find  x  (t)  that  maximizes       (ii)   Find  x  (t)  that  maximizes 

W  in  tl.l)  subject  to  (1.2)         W,  in  II. 1)  subject  to  (1.2) 
for  t£  [0,  T).  for  t£[0,  T) . 

This  problem  looks  very  much  like  a  djmamic  duopoly  problem  formulated 
in  differential  game  theoretic  framework  [7,  8].   The  substantial  differences 
between  these  formulations  are:   (a)  the  duopolists  are  assumed  to  pursue  the 
same  kind  of  objective,  profit  in  case  of  [7,  8],  while  in  this  case  the  objec- 
tives can  be  (i)  profit  in  different  monetary  terms,  or  (ii)  satisfaction,  or 
(iii)  any  combination  thereof,  or  (iv)  any  other  objective  one  country  claims 
to  attain,  and  (b)  the  duopolist  control  their  own  supply  quantity  looking  at 
the  market  dynamics  represented  by  the  price  change,  while  in  this  case  the 
intensity  of  catch  is  controlled  by  the  fishing  industry  in  each  country  looking 
at  the  fish  population  d5mamics. 

By  carefully  checking  the  objective  function,  W  ,  (1.1),  the  fish 
population  dynamics,  (1,2),  and  the  total  and  individual  country  catch  rela- 
tionship, (1.3),  we  can  conclude  that  the  objective  function  depends  only  on 
Xj^(t)  and  X2(t); 

(1.6)       W^  =  W^(x^(t),  X2(t)),  i  =  1,  2. 


(n 


As  in  static  duopoly  situations  there  are  many  different  strategies  the 
two  parties  can  employ;  there  may  be  equally  many  or  more  strategies  our  dynamic 
controllers  (nations)  can  assume.   In  this  paper,  we  confine  ourselves  to  the 
following  two  cases:   (i)  closed-loop  strategy,  and  (ii)  open-loop  strategy. 
The  close-loop  strategy  is  the  one  that  takes  the  fish  population  information  p(t) 
as  time  goes  on  and  adjusts  its  catch  to  it.   On  the  other  hand,  the  open-loop 
strategy  is  the  one  that  looks  at  the  initial  fish  population  and  then  decides 
the  whole  course  of  catch  intensity  decisions  as  a  function  of  time  t  only. 

Of  course,  in  optimal  control  theory,  where  there  is  only  one  controller 
or  decision  maker,  there  is  no  difference  between  the  outcomes  of  these  two 
optimally  (in  some  sense)  chosen  strategies.   However,  these  two  strategies  are 
known  to  bring  about  two  quite  different  catch  intensity  histories  for  these  two 
countries  [  9],  as  will  be  revealed  later  in  the  next  section. 

From  the  game  point  of  view,  we  deal  only  with  the  so-called  "Coumot- 
Nash"  equilirim  game  [3] ,  that  can  be  defined  as  a  pair  of  catch  histories 
(x*^(t,  p(t)),  x*(t,  p(t))  for  the  closed-loop  strategy,   or  (x*Q(t),  x^^Ct)) 
for  the  open-loop  strategy  that  satisfies  the  following  conditions: 
W^(x*^(t,  p(t)),  x*^(t,  p(t))  ^ 


(1.7.1) 


(1.7.2) 


W^(x^^(t,  p(t)),  x*^(t,  p(t))  and 
W2(x*^(t,  p(t)),  x*^(t,  p(t))  ^ 
W2(x*^(t,  p(t)),  X2^(t,  p(t)). 


or 


W^(x*Q(t),  x*Q(t))  ^ 


\^^10^^>'  ^20(^>) 
W2(x*Q(t),  x*Q(t))  ^ 
W2(x*Q(t),  X2Q(t))  , 


and 


(1.7.1)  states  that  the  pair  of  catch  histories  using  the  closed-loop 
strategy  by  both  countries,  (x*  (t,  p(t)),  X2c^*^'  P(t)))  is  better  or  equally 
as  good  as  any  other  closed-loop  catch  strategies  that  this  country  can  assume 
when  another  country  sticks  to  the  star -marked  closed-loop  strategy. 

(1.7.2)  similarly  states  that  the  pair  of  catch  strategies  using  the  open- 
loop  strategy  by  both  countries,  (x*  (t),  x*  (t))  is  better  or  equally  as  good  as 
any  other  open-loop  catch  strategies  that  this  country  can  employ  when  another 
country  sticks  to  the  star-marked  open-loop  strategy. 

In  the  sense  stated  above  (1.7.1)  and  (1.7.2)  are  similar  to  static 
Cournot  or  Nash  equilibrium. 

There  are  other  dynamic  games  such  as  the  dynamic  Stackelberg  game  [3  ] , 
but  due  to  its  operational  difficulty  we  confine  ourselves  to  this  Cournot-Nash 
differential  game  in  this  paper. 

The  necessary  conditions  for  optimality  of  dynamic  Cournot-Nash  (hereafter 
"Cournot"  only)  catch  strategy  in  closed-loop  form,  can  be  derived  as  follows. 

The  Hamil  onian  of  the  Fisheries  Differential  Game  Problem  (FDGP) 
can  be  written  as 


(1.8)        H^  =  W^(x^(t,  p(t)),  X2(t,  p(t)))  +  A^(i:)f(p(t),  x^(t,  p(t))  + 

X2,(t,  p(t)). 

for  i  =  1,2. 

Following  Ho  [3],  Starr  and  Ho  [8],  and  Simaan  and  Takayama  [7],  the 

following  coupled  necessary  conditions  are  derived: 

Country  1 

■   p  =  f(p(t),  x^(t,p(t))  +  X2(t,  p(t))),  p(o)  =  p^ 


(1.9) 


(i) 


^1  =  ^iH(^>  -  h^^^^i  ^  M,  vi) 

^    +  X, (t)  |i  =0 
3x.      1    dx^ 

X  (T)  =  0  or  lim  e"^^X(t)  =  0. 

•^  ^-4<x>  1 


(1.9) -continued 


(1.9) 


(ii) 


Country  2 

p  =  f(p(t),  x^(t,  p(t))  +  x^it,   p(t)),  p(o) 


=  P, 


/  3f  _^  9f  3xA 


^2   "  ^2'*'2^^^  "  '''2^^^ 


f2  =  A2(t)||=0 
9x„    2    dx- 

X      (T)  =  0  or  lim  e"^2^  X(t)  =  0. 

t^co         2 


9f  ^^1 

Here  In  the  second  condition  for  each  country  we  find  a  term —  for  the 

dX,  dp 

first  country  and  ^ —  -r^l  for  the  second  country  (these  are  like  "conjectural 

9x^  9p 

variation  term"  in  static  Cournat  case)  representing  "interdependence"  or  "inter- 
connectedness"  of  their  actions. 

Of  course,  each  country  can  follow  their  strategies  that  ignores  this 
interdependence;  that  is,  each  assumes  that  the  other  country's  strategy  is  deter- 
mined by  the  initial  fish  population,  p  ,  and  time,  t,  only.   Then,  by  constructing 
the  Hamiltonian,  the  following  necessary  conditions  result 


(1.10)    H°=  W.(x-(t),  x_(t))  +  Y.(t)f(p(t),  x/t)  +  X   (t)) 

1     1   1        .id  X  J-         ^ 

for  i  =  1,2. 
Country  1 


(1.11) 


(1) 


p  =  f(p(t),  x^(t)  +  X2(t)),  p(0),  p(o)  =  p^ 


9f 

V  '^i  \^'^  "■  \^'^  9F 

|^l  +  Y,(t)f  =0 

dx^     1     ox^ 

Y(T)  =  0  or  lim  e"^l^Y'^(t)  =  0 


(1.11) — continued 
(1.11) 


(ii) 


p  =  f(p(t),  x^(t)  +  X2(t)),  p(o) 

3f 


=  P, 


Y2  =  r^  Y2(t)  +  Y2(t)  3p 


?2  +  Y2  (t)  If  =0 

ax„     I  OX- 


Y^(T)  =  0  or  lim  e  ^l^Xit)   =  0 


9f   3x 
Here  in  this  open-loop  catch  strategy  case,  the  interaction  term  -r—  —^r^   is 

ox,    dp 

missing  from  the  second  condition  for  each  country.   In  the  next  section,  we  will 
trace  out  the  consequence  of  this  difference  in  detail. 


2.   Quadratic  Benefit — Linear  Population 
Dynamics  Case 


The  main  purpose  of  explicibly  formulating  FDGP  in  the  quadratic  benefit- 
linear  fish  population  dynamics  framework  is  to  firmly  quantify  optimal  strategies 
and  resultant  fish  population  dynamics,  and  consequently  to  derive  policy  impli- 
cations from  the  results. 

In  this  section  we  deal  with  the  following  two  cases: 
Case  1:  (Competitive  Markets — Close-Loop  Game) 

The  social  benefit  function  is  defined  as  the  over-time  integral  of 
the  instantaneous  (we  omit  this  term  hereafter)  consumers'  and  producers'  surplus 
accruing  from  the  fish  market ,  and  the  two  countries  use  the  closed-loop  catch 
strategy. 
Case  2 :  (Competitive  Markets — Open-Loop  Game) 

The  social  benefit  is  the  same  as  defined  in  Case  1,  but  the  two  countries 
use  the  open-loop  catch  strategy. 

Another  pair  of  cases  dealing  with  a  monopolist  controlling  the  domestic 
Tiarket  in  each  country  can  be  solved  easily,  but  we  will  not  go  into  them  in  this 
paper. 


10 


There  are  other  combinatorial  possibilities  for  our  case  studies  of 
the  Coumot  game,  and  we  plan  to  do  some  exhaustive  work  later  in  this  area. 

2.1.   Competititve  Markets — Closed-Loop 
Catch  Strategy  Differential  Game 

Following  Sarauelson  [6]  and  Takayaraa  [10],  let  us  express  the  consumers' 
benefit  functional  as  the  time  integral  over  [0,  T) ,  where  T  could  be  infinity, 
of  the  integral  of  the  instantaneous  demand  (felicity)  function: 

(2.1)         Consumers'  Benefit  in  the  ith  country  = 

T 

-r  t        12 
CB.  S  /  e  i  [a.x,  -  tt  3.  x.  ]dt 
1   /         1  i   2  1  1 


./ex  [a.x^  -  2 
*'o 


for  i  =  1,2. 
where  the  instantaneous  demand  function  of  the  ith  country  for 
the  fish  species  is  given  as 


(2.2)         P^,  =  a.  =  6.  X.,  i  =  1,2 
di    X    XX 


We  assume  in  the  two  cases  we  handle  in  this  section,  that  the  catch  x.  is  consumed 
without  any  wastage  in  the  process  or  the  consumption  is  measured  in  the  fresh 
fish  unit. 

The  total  cost  of  catching  x,  units  of  fish  is  expressed  as  the  over- 
time integral  of  the  integral  instantaneous  producers'  supply  function. 

T 
(2.3)        TC.  E  f     e'^-i^Lyx.,  +  ^  e,x/]dt 


/-r  .t  r      ,  1  rN     2i 
e  X  [^x..  +  -  e.x.  ] 


where  the  instantenous  producers'  supply  function  is  expressed  as 


11 


(2.4)  P  .  =  y.  +  0.  x.(t) 

SX      1      11 


for  i  =  1,2, 


The  social  benefit  function  of  the  individual  countries  can  be  expressed 
as  the  difference  between  the  consumers'  benefit,  CB  ,  and  the  total  cost  of 
catch,  TC. ;  that  is 


i 
SB.(x   (p(t))  =  CB.  -  TC   E  /*  e"''i''[(a.  -  y.)x.  -  y  (6.  +  0,)x? 


(2.5)       SB^.(x,  (p(t))  =  CB,  -  TC,   E  /  e  ^i  [  (a,  -  y,)x,  -  ^  (6,  +  0,)x,  ]dt 

o 

for  1  =  1,2. 


Now,  let  us  define  our  competitive  markets — closed-loop  catch  strategy 
differential  game  as  follows: 

CMCLDG: 

Find  the  d5mamic  paths  of  the  pair  (x*  (p(t)),  x*  (p(t)))  that  satisfy 
the  following  Cournot  conditions 

SB^(x*^(p(t))  ^  SB^(x^^(p(t)))   and 

SB2(x*^(p(t))  ^  SB2(x2^(p(t))) 

subject  to: 

(1.4.C)  p  =  a  +  b  p  -[x^^(p(t))  +  X2^(p(t))]. 

The  Hamiltonianfor  this  problem  is  given  as 

(2.6)   Hf  =  (a.  -  ii.)x.   -^(3.  +0.)x^ 
1     1    1  ic   2   1    1  ic 

+  A.^  (a+bp  -  K^^  -x^^). 
The  necessary  conditions  for  optimality  of  the  pair  of  catches  are: 


12 


(2.7)-, 


(i) 


Cii) 


(a)  p  =   2   +  bp  -  x^^   -  x^^,    p(o)    =  p^ 

(b)  \,  — (b   -  r^   -  |f2c)X^^ 


(c)        (Ci^    -   ti^) 


(3,     +0i)xie-\c   =   ° 


(d)      A      (T)    =  C  or  lim  e  ^1*^   ^0^,^^)   =  0 


(a) 


3   +  bp    -  X 


Ic 


^'2c'   P    ^°^    =  Po 


^(b)    A^^  =   (-b  -r^   -|fl)A2^ 

(c)  (a„   -  ^2)    -    (^2   +  Q2)^'2c   -   ^2c  =  ° 

(d)  A^    (T)    =  0  or   lira  e"^2^A^    (t)    =  0. 


lor   the  siipjlicity  of   Cv^n^pucation,    in  the  following 
we  assume   ti  at 


(2.8) 


a^  =  a. 

- 

a 

Bt^   =   02 

s 

6 

'i  =  h 

= 

y 

e,  =  G. 

5 

0 

whicli  is  ^i^aivaien^  to  siaCriig  that  i:be  t^wo  contries  possess  the  same  demand  and 

cost  structures.   In  i;a::.s  case,  aftar  scxe  calculations  involving  the  solving  of 

2 
Ricatti  equations,  we  get  the  following  solutions.    The  first  pair  are 


(2.9) 


x^*(pu))  =  x|^(p(t))  =  1^ 


whicn  are  not  df,sirable  or  stable  solutions  for  these  two  countries  unless  the 
fish  population  at  the  initial  period  was  already 


(2.10) 


-,*   = 


L  2  + 


e 


-  a 


The  detailed  d'?rivation  of  the  solutions  is  given  in  the  Appendix 
at  the  end  of  this  paper. 


13 


(2.11) 


-  a 


A  pair  of  optimal  stable  catch  strategies  exist  and  are 

(a)  X*  (D(t)^=  g  -  y  _  (2b-2r^+r?)    2 (a  -  y) 

,  2b-2ri+r9  .. 
+   2  —  "P^^J 

(b)  X*  (p(t)  =  "  -  ^  -  (2b-2r^+rj)  l(a  -  P) 

^2b:ir2±rip(t)  . 


-  a 


The  fish  population  moves  along  the  following  converging  path. 


(2.12) 


(2.13) 


=  (b-ri -r2) 


3b 


-{^^} 


2(a  -  u) 

3  +  e 
P(t)  , 


-  a 


where  we  assume 


(i)  b  -  r^  -  r^  >  0 


(U,  H|^.,,„ 


The  fish  population  converges  to 


(2.14) 


1  r2(a  -  I 

b  L  B  +  G 


-  P) 

0 


-  a 


>  0 


Due  to  (2.12),  we  find  the  ultimate  fish  population  as  t  tencfg 
infinity  as 


to 


(2 


•^»    PJ  -  I  ["^^  -]  >  0- 


Thus,  the  fish  population  history  accompanied  by  the  optimal  catch 
(2.11)  is  given  as 


14 


(2.16) 


P^(t)  =  (p^ 


p*)e  +  p* 

^c  '  c 


as 


From  (2.11)  and  (2.1A)  we  get  the  ultimate  optimal  catch  intensity 


(2.17)        -!e  =  ^fc=fTi  • 

The  following  comparative  d3mamics  conclusions  can  be  derived 
from  the  above  results.   The  discount  rates  do  not  affect  the  optimal  target 
(t  -*<»)  catches  or  the  target  population;  that  is 


(2.18)     .^_^^o   ^,^=0,    1  =  1,  2. 


9r. 

1 


3r, 


dr, 


Given  the  social  discount  rate  of  country  1,  r  ,  an  increase  of  the  discount 
rate  of  country  2,  r  ,  will  decrease  the  catch  intensity  of  country  1  at  any  level 
of  the  fish  population  less  than  p*  ,  and  increase  that  of  country  2.   Similarly, 
an  increase  of  the  discount  rate  of  r^ ,  given  that  of  country  2,  r„ ,  will 
increase  the  catch  intensity  of  country  1,  and  decrease  that  of  country  2  at  any 


p(t)  <  p*. 


(2.19) 


In  other  words,  we  have 


9x   (p(t)) 

9r.    =  ¥^^'^   -   P^> 


3x^^(p(t))   2 


3r. 

1 


3^ 


p(t)) 


for  i,  j  =  1,2. 


As  a  result,  if  covmtry  1  becomes  more  conscientious  about  the  future 
generations  than  country  2  in  relation  to  fish  consumption  and  production,  r^ '  > 
r^ ,  country  1  becomes  more  conservation-oriented  than  before  in  order  to  attain 


J    .0 


J.  rfji<!<j-t  '*')  %■' ^ar/'^^:Ji    ,4.i3,-'.'   atfj 


,io')    );■*  i^.'iz   •=>s6ft'j';i.'li   t:nti    .  ' 


15 

the  maximum  pcosible  social  baneflt  in  the  long  future  and  in  perpetuity. 

Thi.5  conclusion  establishes  the  principle  of  conservation  for  renewable 
resources  including  fish,  deer,  forestry,  and  others. 

Also  this  sfTiie  prirclple  has  a  potential  element  that  makes  one  country, 
say  country  1,  legislate  the  two  hundred  mile  territorial  waters  limit  if  she 
sees  foreign  vessels  exploit  her  territorial  fishing  beds  to  an  undesirable  extent . 

Traditional  contro/ersies  over  whether  or  not  some  catch  control  regula- 
tion is  necessary  can  be  answered  in  this  two  country-one  fish  species  framework. 

As  in  the  single  country-single  species  case  [10],  we  can  conclude  that 
the  total  catch  tor  each  country  stipulated  by  (2.il)  is  necessary  for  both 
countries  to  en^^oy  ultimate  maximum  social  benefit.   The  reason  is  rather  straight- 
forward.  Since,  hy   (2.7)  (i,c)  and  (ii.c),  the  marginal  social  value  of  a  unit 
of  fisti  sold,  in  ta--  narket  always  excaads  the  marginal  individual  cost  of  catching 

the  same  by  A.  'D(t/)>  0  cr  X.    (t)  >  0  for  p  <  p*  or  p  <  p^ ,  individual  fish- 
j      -,  f.  ..  V  .  /        ro  "^o    c      o    o 

ericin  wou;  d  like  ro  fish  moxe,  if  left  to  follow  their  own  individual  profit 
maximisation  p-rinclples,  thar.  ths  socially  opti&al  (2.11).   This  overcatch  rep- 
resented by  v.2.9)  at  each  mom.ir  t  of  tiKe.  when  p(t)<  p*  or  p*  ,  must  be  stopped. 
An  internatiCiiax  regulatici-s  controlling  the  catches  by  both  countries  must  be 

worked  out  in  thl^  ca^e.   Kow  to  acc-omplish  this  lies  outside  the  scope  of  this 

3 

paper . 

Mesh-cizt  regulation  arguments  have  been  carried  out  in  a  static  frame- 
work in  [1,  11],  and  in  dynamic  one-ccuntry  on3-fish  species  framework  in  [10]. 
Following  [10],  one  can  develop  similar  arguments  for  some  kind  of  mesh  regulations. 
The  reason  is  simple  enough:   the  iniormatd  on  on  the  optimum  ri.esh-size  for  the 
industry  and  the  consuroing  societies,  if  it  exists,  is  external  to  the  individual 

For  seme  arguments  in  this  direction,  the  reader  is  referred  to  [1,  11]. 


16 


fishermen  and  the  industry  concerned  in  each  country. 

2.2.   Competitive  Markets-Open-Loop  Catch  Strategy  Differential  Game 

If  the  two  countries  decide  to  take  the  open-loop  strategy  in  the  compe- 
titive market  within  their  national  boundary,  the  objective  fimctions  remain  to 
be  the  same  as  (2.5)  for  i  =  1,2.   The  population  dynamis  is  the  same  as  before, 
(1,4),  with 


(2.20) 


X  =  x^Q(t)  +X20  (t) 


The  problem  can  be  defined  as  follows : 

CMOLDG    (Competitive-Markets-Open-Loop  Catch  Strategy  Differential  Game) 

Find  the  dynamic  paths  of  the  pair  (x*  (t),  x*^(t))  that  satisfy  the 


20 


following  conditions 


SB^(x*Q(t))  ^  SB^(x^Q(t))    and 
SB2(x*Q(t))  ^  SB2(x2Q(t)) 
subject  to 


(1.4.0)    p  =  a  +  bp  -C^^qM   +  X2Q(t)) 


The  Hamiltonian  for  this  problem  is  given  as 


(2.21) 


H.  =  (a.  -  y.)x. 
1     X    1  xo 

+  A.  (a  +  bp 
xo 

for  i  =  1,2. 


v(B.  +  G.)x^ 


X   xo 


""lO  ~  ""20^ 


(2.22) 


The  necessary  conditions  for  optimality  of  the  pair  of  catches  are: 


(i) 


(a)   p  =  a  +  bp  -  x^Q  -  x^Q,  p(o)  =  p^ 


(b)  X 


10 


•(b  -  r)A 


10 


(c)  (a^  -  u-,^)  =  (3-1^  +  G^)  x^Q  -  \o  ^  ^ 

(d)  A^q(T)  =  0,  lim  e  -"  X^QCt)  =  0 


17 

(2 . 22) — continued 

(a)   p  =  a  +  bp  -  x^^  -   x^Q,  p(o)  =  p^ 


(ii) 


(b)   A^g  =  -(b-r)X2o 


(c)  (a^  -  ■.^■)    -   (B2  +  0,)x,o  -  X^o  =  0 

-r  t 

(d)  X^q(T)  =  0,  lim  c  ^   X^^(t)  =  0 


I 


As  in  the  previous  case.  Case  1,  wt  assume  for  simplicity's  sak^,  that 
(2.8)  holds;  the  two  countries  are  the  same  in  their  demand  and  supply  structure. 

In  this  case,  hov/ever,  there  is  no  optimal  stable  (convergent)  open- 
loop  catch  strategies  unless  zhe  following  condition  holds: 

(2.23)     r  =  r„  (-■  v   herearter) . 

The  implicaticn  oj;  this  condition  is  quite  Interesting  and  suggestive. 
If  the  two  countries  with  the  same  demand  and  supply  (cost)  structure  engaging 
fisiiing  the  -jana  fish  ^p-acies  in  the  common  ocean  (fishing  banks),  and  if  one 
country's  future  disccunt  rate  is  different  from  the  other,  these  countries  can- 
not find  any  reasonable  catch  strategy  that  will  eventually  bring  them  to  their 
target  catch  and  fish  population  defined  and  discussed  in  the  previous  subsection. 

Wrat   luakes  the  opef-loo^.^  catch  stratrgy  case  so  diagonally  different 
from  the  closec'-locp  ca^ch  strategy  game?   In  the  case  of  the  closed-loop  case, 
the  parties  involved  observe  carefully  the  history  of  fish  population  over  time, 
p(t).   This  makes  ons  party's  catch  responsive  to  the  other  party's  catch  through 
the  observed  fish  population.   Thus,  each  party  responds  to  each  other  sensitively, 
and  this  interaction,  brings  about  an  optimum  catch  strategy  to  each  party  (as 
long  as  (2.13)  holds).   However,  in  the  case  of  the  open-loop  strategy  game, 
each  party  decides  that  she  can  determine  her  o',jn  optimal  strategy  on  the  basis 
of  the  initial  fish  population  and  time.   Thus,  unless  complete  symmetry  exists 


18 


in  their  environment,  they  cannot  come  up  with  any  optimal  catch  strategy. 

After  superimposing  (2.8)  and  (2.23),  we  get  the  following  results. 
A  pair  of  singular  solutions  exist  and  are: 


(2.24)    x^*(t)  =  X2*(t)  =  f^     . 

They  are  not  desirable  or  stable  solutions  unless  the  fish  population 
is  already  at  the  level  represented  by  (2.10).  Optimal  and  stable,  open-loop 
catch  strategies  exist  and  are 


(2.25)  x*Q(t)  =  x*Q, 


f^\  ct  -  y   /2b-r  \  g  -  y    |  ,  /2b  -  r  ^  /.  >, 


The  fish  population  dynamics  using  the  open-loop  catch  strategies  is 
then 
(2.26)    P  =\-^  I'a   a.  q'  -  a  -  (b-r)p 


?,  ,/b-rY2(a  -  v) 
P   \  b  ^  6  +  0 


(2.27)       (i)   b-r  >  0 

v...      2(a  -  y)   ,  .  n 


This  population  converges  to 

(2.28)     p;   =  i  plVe^  -  a] 

which  is  exactly  the  same  as  the  closed-loop  desired  or  target  population, 
(2.14). 


(2.29) 


The  ultimate  optimal  catch  in  perpetuity  is  given  by 

10   ^20   S  +  0 
as  expected . 


The  optimal  time  profile  of  the  fish  population  in  this  case  is  given 


as 


•(b-r)t 


(2.30)    p^(t)  =  (p^  -  pj  )e 


+  P*, 


19 


As  a  consequence, the  optimal  open-loop  catch  strategy  brings  a  much 
faster  convergence  to  the  common  target  fish  population 


ifzca  -y)  _  J. 
b  |_  B  +  0    J 


At  the  same  time,  for  a  given  initial  fish  population,  the  optimal  open- 
loop  catch  is  always  smaller  than  the  optimal  closed-loop  counterpart  as  the 
following  computation  shows: 


(2.31) 


x*^(p(0))-  x*^(0) 


(2b  -  r)(D 
■ -o- 


^H  0, 


for  p(0)  <  p*  =  p*  =  p* 


which  is  shown  in  Figure  2  below. 


X 


* 

X      = 

c 

X* 

o 

(        X* 

^        c 

= 

X*        + 

ic 

■X* 

■^ic 

2  ""  ~ 
^  B  + 

y 

0    =  ■ 

X*       + 
lO 

X*        = 

xo 

X* 
O 

) 

p  =  0 


Zx*^'tp(t))// 

Ac  Zx?_(t) 


xo 


'Ao 


p« 


Figure  2.   Optimal  catch  strategies^x*  (p(t))  andJxf-Ct), 


As  a  consequence,  the  open-loop  catch  strategy,  assuming  of  course  that 
the  two  countries  use  the  same  future  discount  rate,  brings  a  much  faster  con- 
vergence to  the  target  fish  population. 


<«1i   Ob   .3-iKt,-i;  I  .»ic 


0   =   (< 


A,;    ,    -^.q 


X  51 


('  \i'. 


\ 


20 


i  U(a   -p)  _   1 
b  L  B  +  ©     J" 


Thus,  declaring  that  both  countries  employ  the  same  future  discount  rate  and 

stipulate  and  monitor  the  optimal  history  of  catch  each  country  is  to  follow, 

may  be  a  faster  way  to  reach  the  target  population  and  catch. 

It  is  also  easy  to  conclude  that  along  the  stable  convergent  linear  paths, 

A  E  and  A  E,  the  catches  and  the  fish  population  increase  till  they  reach  the 
c       o 

long-run  equilibrium  point,  the  Cournot-Nash  equilibrium  point,  if  these  two 
countries  employ  the  closed-loop  and  open-loop  catch  strategy,  respectively 
(provided  p  <  p*  =  p*  ) . 

The  future  discount  rate  has  no  effect  on  the  target  catch  or  fish  popu- 
lation.  However,  as  r  increases,  the  optimal  intial  catch  will  increase  due  to 
the  fact  that 

(2.32)    ^'^io'^"''     1   r  *     r  ^^s^   •    i    -y 
T^ =  2      (P*  -  P(o))>0,  1  =  1,2, 

as  long  as  p(o)  <  p*. 
This  is  consistent  with  the  larger  future  discount  rate  or  the  assertion  of  the 
"more  now,  less  later"  attitude  of  a  society. 

The  same  conclusions  as  those  for  the  closed-loop  case  apply  to  this 
case  as  to  the  catch  control  and  mesh-size  issues. 

Conclusion 

In  this  paper,  we  have  formulated  the  fisheries  economies  problem 
involving  two  countries  and  one  fish  species  in  a  differential  game  framework. 
A  quadratic  social  benefit  and  linear  fish-catch  dynamics  problem  is  then  solved 
for  the  Cournot-Nash  differential  game  solutions.   Obviously,  there  are  at  least 
two  strategies  to  play  within  this  game  framework,  and  the  solutions  are  derived 


21 


for  the  closed-loop  catch  and  open-loop  catch  strategies. 

We  find  that,  if  the  demand  and  (cost)  supply  structures  of  the  two 
countries  are  exactly  the  same,  the  closed-loop  catch  strategy  generates  a 
pair  of  optimal  catch  paths  to  follow  for  any  future  discount  rate  of  indivi- 
dual countries  as  long  as  b  -  r^  ~  ^9  '**-'>  while  the  open-loop  counterpart  pro- 
duces a  pair  of  twin  catch  paths  to  follow  only  if  the  future  discount  rates  of 
individual  countries  are  exactly  the  Game.   This  is  an  outstanding  feature  of 
this  renewable  resource  differential  game. 

In  both  cases,  the  increasing  appreciation  of  the  present  over  the 
future  increases  the  optimal  catch  at  present,  slows  down  the  process  of  con- 
vergence of  the  fish  population  to  the  target. 

In  the  case  of  the  closed-loop  catch  strategy,  an  increase  of  country  I's 
future  discount  rate,  r  ,  increases  the  initial  optimal  catch  of  country  1, 
while  country  2  accepts  the  fact  that  r  has  increased  and  curtails  the  optimal 
catch  accordingly  (see  (2.19)).  VThether  country  2  is  willing  to  follow  this 
course  of  action  may  not  be  an  issue  at  all.   Rather,  that  r.  and  r„  are  histori- 
cally given  and  accepted  as  such  must  be  the  basic  framework  of  our  model. 

Of  course,  if  the  sum  of  the  two  countries'  discount  rates,  r  ■*"  r„ , 
exceed  the  rate  of  increase  of  the  fish  population,  b,  there  will  not  be  any 
economically  meaningful  solution  for  our  closed-loop  strategy  game.  While  the 
open-loop  strategy  game  is  much  less  restrictive  in  this  respect  since  that 
the  common  rate  of  future  discount,  r,  must  be  less  than  the  growth  rate  of 
the  fish  population,  b,  is  the  only  requirement.  However,  since  the  condition 
for  the  existence  of  optimal  catches  depends  on  the  equality  of  the  two  rates, 
the  open-loop  catch  strategy  may  be  considered  too  restrictive. 

A  dynamic  cheory  of  fisheries  economies  we  have  developed  so  far  has  estab- 
lished a  conservation  principle  that  a  scarce  renewable  resource  such  as  a  fish 


22 

species  for  human  consumption  requires  a  total  catch  control  or  a  control  over 
the  catch  by  the  fishing  fleets  of  every  individual  country. 

Also,  we  argue  that  some  workable  mesh-size  regulation  must  be  enforced 
since  individual  fishermen  have  no  way  of  knowing  what  mesh-size  is  optimal  from 
the  consuming  societies'  point  of  view  (as  well  as  their  own  in  the  long-run), 
that  is,  this  information  is  external  to  both  consumers  and  fishermen. 

There  are  many  directions  to  go  and  many  topics  to  cover  in  the  future 
research  in  the  area  of  renewable  resources  economics.   A  natural  extension  of 
our  two  countries-single  species  formulation  is  a  two  countries-single  species 
formulation  with  different  demand  and  supply  structures.   Also,  finite  time 
horizon  problems  should  be  solved  for  more  practical  problems.   A  system  of 
matrix  Ricatti  equations  must  be  solved  to  obtain  any  meaningful  quantitative 
results  from  a  multiple  species-two  countries  formulation.   In  this  direction, 
computer-based  algorithms  are  absolutely  necessary,  and  many  diverse  and  practi- 
cal problems  may  be  solved  efficiently. 

Another  extension  is  to  solve  more  than  two  countries-one  species  model 
by  effectively  utilizing  our  model  developed  here  along  with  the  concept  of 
coalitions. 

An  ultimate  extension  is  to  formulate  and  solve  multiple  countries- 
multiple  species  problems  in  generality,  but  quantitatively. 

Fish  population-catch  djmamics  can  be  made  nonlinear  to  attain  generality. 
One  observation  dealing  with  a  quadratic  population  dynamics  case  is  already  dis- 
cussed elsewhere  [10],  and  may  shed  some  new  light  on  the  predator-prey  dynamics 
[5]  widely  accepted  by  the  economics  profession  for  some  time. 

Econoraetrically,  estimation  of  parameters  in  population-catch  (or  forestry 
growth  and  harvesting)  dynamics  is  a  challenging  field.   This  is  a  field  in  which 
various  interdisciplinary  activities  will  prove  most  productive.   Cooperative 


23 

>f forts  in  these  directions  and  topics  stated  above  by  fisheries  specialists, 
)ptimal  control  and  differential  game  specialists,  economists,  and  environ- 
lentalists  will  make  regulations  and  control  economically  and  politically  viable 
md  sound . 


24 


References 


[1]   J.  H.  Boyd;  "Optimization  and  Suboptimization  in  Fishery  Regulation: 

Comment,"  American  Economic  Review,  June  1966,  vol.  56,  pp.  511-517. 

[2]  H.  E.  Crowther;  "Our  Fishing  Industry  in  the  World  Race:  A  Strategy  or  an 
Awakening  Challenge?"  in  Recent  Developments  and  Research  in  Fisheries 
Economics  (Eds,  F.  W.  Bell  and  J.  E.  Hazleton) ,  1967,  pp.  19-27. 

[3]   Y.  C.  Ho;  "Differential  Games,  Dynamic  Optimization  and  Generalized 

Control  Theory."  Journal  Optimization  Theory  and  Application,  September 
1970,  Vol.  6,  pp.  179-209. 

[4]   P.  A.  Samuelson;  "A  Universal  Cycle?"  The  Collected  Scientific  Papers  of 
Paul  A.  Samuelson,  Vol.  3,  1972,  pp.  473-486. 

[5]  ;  "Generalized  Predator-Prey  Oscillations  in  Ecological  and 

Economic  Equilibrium"  ibid.,  pp.  487-490. 

[6]   ;  "Intertemporal  Price  Equilibrium:   A  Prologue  to  the  Theory  of 

Speculation,"  ibid.,  Vol.  2,  1966,  pp. 946-984. 

[7]   M.  Simaan  and  T.  Takayama;   Dynamic  Duopoly  Game:   Differential  Game 
Theoretic  Approach,  Faculty  Working  Paper  #155,  February  1974. 

[8]  and  ;  "An  Application  of  Differential  Game  Theory  to  a 

Dynamic  Duopoly  Problem  with  Maximum  Production  Constraints,"  to  appear 
in"  Automat ica,  March  ,  1978. 

[9]   A.  W.  Starr  and  Y.  C.  Ho;"Nonzero-Sum  Differential  Games."  Journal  of 

Optimization  THeory  and  Application,  March  1969,  Vol.  3,  pp.  184-206. 

[10]   T.  Takayama;  Dynamic  Theory  of  Fisheries  Economics  -  1:  Optimal  Control 
Theoretical  Approach,  Faculty  Working  Paper  #437  ,  College  of  Commerce 
and  Business  Administration,  September,  1977. 

[11]   R.  Turvey;  "Optimization  and  Suboptimization  in  Fishery  Regulation," 
American  Economic  Review,  March  1964,  Vol.  54,  pp.  64-76. 


25 


Appendix 

In  this  appendix  only  Case  1  problem  is  solved  in  detail.   The  reader 
irf.ll  find  it  easy  to  solve  Case  2  problem  by  following  the  development  below. 

From  (2.7)  and  assuming  (2.8)  we  have  the  following  necessary  condi- 
tions 


(A.l) 


(i) 


(ii) 


(a)  p  =  a  +  bp  -X,   -  X  ,  p(o)  =  p 
ic    Zc  o 


(b)  \,  —i^-r,-  -^  )\, 


(c)  (a  -u)  -  (B  +  0)x^  -  X  ^  =  0 

Ic    i^c 

-r  t 

(d)  lim^  "■     X^^(t)  =  0 


(a)  p  =  a  +  bp  -  x^^  -  x^^,  p(o)  =  p^ 

(b)  L  =  -(b  -  r.  -  ^^Ic 

(c)  (a  -  y)  -  (3  +  0)X2^  "  ^2c  "  ° 


(d)  lim  e"'^2'^X„  (t)  =  0 
zc 


By  setting 

(A. 2)    X^   =  K.p  +  E.,  i=  1,2, 
ic    X     X 


we  have,  from  (A.l)  (i,c)  and  (ii,c). 


(A.  3)   X 


q  -  y 


K,- 


ic    e  +  Q    B  +  e*^   B+0 


^P  - 


Ei 


r,  i  =  1,2. 


By  differentiating  (A. 2)  with  respect  to  t,  and  equating  the  results  to 
(A.l)  (i,b)  or  (ii,b)  respectively,  one  gets 


26 


K    K  K 

(A.  A)        {K.  +  K^(b  +  g^  g-fg)  +  (b  -r.  +  ^)\}   P 


+  {K,(a--f^ 


2(a  -  y) 

e 


F        F  K 


(A.  5) 


for  i  =  1,2. 

Equality  in  (A. 4)  should  hold  for  all  variations  of  p.   Thus, 

we  get 

K       K  K 

^i  -^  ^i  (b  +  sTT  -^  3^^  +  (b  -  r,  +  3^)  K^  =  0 

^i<-  -'r^  ^  FT^-^3^>  +  (b  -  r.  +  3^)E^  +  E^  =  0 


Assuming  that  K  and  E  converge  to  zero  as  t  tends  to  infinity,  we  get 

K.      K^ 


(A.  6) 


(A.  7) 


for  i  =  1,2 
that  K  and 

(2b  -  r,  +  -34-^;  + 
K^(a  - 


K. 

1 


iB+OB+ee+o'^i 

2(a  -  y) 


;)  K,  =  0 


E  E  K 

+  ^r^-^  +   (b  -  r.   +  ^ 


B+0B+0       B+0 


for  i  =  1,2. 
One  set  of   signular  solutions  is  obtained  as; 


^1  "  ^2  "  ° 


i   6  +  0'  i 


•)E.  =  0 


\=  E2=0. 


The  other  set  of  stable  solutions  is  obtained  after  some  algebraic 
manipulations 


(A.  8) 


=  (2b  -  2r  +  rg)(B  +  0) 
1  3 


_  _  (2b  -  2r^+  r,)(B  +  0) 

E  =  (2b  -  2r;^  +  r,)  [2(a  -  p)  -  a(B  +  0)1 
■"•  3b 


E  =  (2b  -  2r^  +  r.^)[2(a  -  u)  -a(e  +  0)  ] 
^  3b 

By  substituting  (A. 8)  into  (A. 2),  (A. 3)  and  (A.l)  (ii,a)  or  (ii,a)  in  that  order, 
we  get  the  desired  results. 


•  «tJ-'- i 


2  ■ 

J  Of 


-  i^^riir'  t^--^  ^  e^T- 


T      c;  vy 


•,6P    l":";iBj' 


^l&:id  jglifl    i.-n.  a   195"' G   faD«?±r>t' 


0 
.0 


(B.i±) 


27 


The  two  other  unstable  pairs  of  strategies  as  other  solutions  of  the 
Ricatti  equations  are 


(A.9) 


(a)   X* 


g  -  U 


ic   B  +  0 


(b)  x*^(p(t))  =  f-ri  {^{ffi   -a)  ^  ^= 


and 


r   \         *  -  ot  -  U    t  2b-r  Vg  -  y    ) 


(A. 10) 


J.  2b-r  1 


/,  >,   jt  _  g  -  y 


Corresponding  to  these  strategies,  the  fish  population  moves  along  the  paths 
stipulated  by  the  following  dynamics 


(A.  11) 


•  _    r.  (2(a  -  y)    \         r 

p  -  -  iS  Vb  +  0  -  7  "^  2-" 


p   for  i  =  1,2, 


which  are  unstable  due  to  the  fact  that  r.  >  0  for  i  =  1,2. 

It  is  obvious  that  these  two  solutions  do  not  really  satisfy  (2.7) 
(i,  d)  or  (ii,  d) .   Therefore,  even  though  these  are  solutions  of  the  Ricatti 
equations,  they  are  not  those  of  (2.7). 

similar  results  as  above  can  be  obtained  for  the  open-loop  strategy  case. 


3-9*