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375' 

^AUNA HAWAIIENSIS 

BEING THE LAND-FAUNA OF THE 
HAWAHAN ISLANDS 




''■ks 



VOLUME II ^^^.^ 



BY VARIOUS AUTHORS 



1 899—1 91 



PUBLISHED IN SIX PARTS 
EACH WITH A SEPARATE TITLE-PAGE 



CAMBRIDGE 
AT THE UNIVERSITY PRESS 

1913 



©ambriUge : 

PRINTED BY JOHN CLAY, M.A. 
AT THE UNIVERSITY PRESS. 



JUN15 1S55 
i/BRARl 



352072 



FAUNA HAWAIIENSIS 




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CAROLO R. BISHOP, 

VIRO HONORATISSIMO, 

QUI ANIMO BENEVOLENTE ET MANU PROFUSA 

INSULIS HAWAIIENSIBUS 

SCIENTIAE CAUSA 

INSTITUTIONEM MAGNIFICAM 

STABILIVIT, 

OPUS HOC 

INSCRIBITUR. 



CONTENTS OF VOLUME II. 



Orthoptera by R. C. L. Perkins (Plates I and II) 

Neuroptera by R. C. L. Perkins (Plates III — V) 

COLEOPTERA PhYTOPHAGA BY D. SHARP (Plate VI) 

Coleoptera Rhynchophora (p. 117), Proterhinidae (p. 183), Heteromera (p 
CioiDAE (p. 253) BY R. C. L. Perkins (Plates VII— X) .... 



MOLLUSCA BY E. R. Sykes (Plates XI and XII) . 

Earthworms by F. E. Beddard 

Entozoa by a. E. Shipley (Plates XIII and XIV) 
Arachnida (except Acari) BY E. Simon (Plates XV— XIX) 

CRU.STACEA LSOPODA BY A. DOLLFUS (Plate XX) . 

Cru-stacea Amphipoda BY T. R. R. Stebbing (Plate XXI) 
Hemiptera (supplement) BY G. W. Kirkaldy 
Hymenoptera (supplement) by R. C. L. Perkins 
Orthoptera (supplement) by R. C. L. Perkins 
Neuroptera (supplement) by R. C. L. Perkins 
Diptera (supplement) by R. C. L. Perkins 
Twenty- one plates as mentioned above 



246), 



PAGE 
I 

31 
91 

17 ETC. 
271 

427 

443 

S2I 

527 

531 
600 
687 
691 
697 



NOV 5., 

^""eton, 0, Q 



FAUNA HAWAIIENSIS 



VOL. II. PART I. 



ORTHOPTERA 



R. C. L. Perkins. 



Price Eight Shillings. 
To Subscribers Four Shillings. 






The Fauna Hawaiiensis is being published in parts at irregular inter- 
vals, and will it is hoped be completed in about two years. 



Contributions have been made or promised by the following, viz. 



F. E. Beddard, F.R.S. 

Prof. AUGUSTE FOREL 

Dr L. O. Howard and W. H. Ashmead 

G. W. KiRKALDY, F.E.S. 
Sir John Lubbock, F.R.S. 
E. Meyrick, F.Z.S. 

R. C. L. Perkins, B.A. 

R. I. POCOCK 

D. Sharp, F.R.S. ... 
A. E. Shipley, M.A. 
M. EuG. Simon 

E. R. Sykes, F.Z.S. 
The Lord Walsingham, F.R.S. 
Mynheer F. M. van der Wulp 



Earthworms. 

Formicidae. 

Hymenoptera Parasitica. 

Hemiptera. 

Thysanura and Collembola. 

Macrolepidoptera. 

{Hymenoptera Aculeata, Orthoptera, 
\ Neiiroptera, and part of Coleoptera. 

Myj'iapoda. 

Part of Coleoptera. 

Parasitic Worms. 

Arachnida. 

Molliisca. 

Microlepidoptera. 

Dipt era. 



It is also intended to give a list of the Vertebrates, with their distribution, 
in the Islands. 



N.B. The parts of Volumes I. and II. are being published concurrently in order 
to expedite the completion of the work. 

The price of each part will vary according to its extent and the number of Plates. 
Subscribers to the whole work will be charged half-price for each part. The parts will 
be sent, as published, to each subscriber who has paid for the preceding part. 

Those who wish to be subscribers, should send name and address either directly to 
C. J. Clay & Sons, Cambridge Warehouse, Ave Maria Lane, London, E.C., or through 
any bookseller. Payments for each part should also be made to Messrs C. J. Clay & Sons. 



FAUNA HAWAIIENSIS 

OR THE 

ZOOLOGY OF THE SANDWICH (HAWAHAN) ISLES 

Being Results of the Explorations instituted by the Joint Committee 

appointed by 

THE ROYAL SOCIETY OF LONDON FOR PROMOTING NATURAL KNOWLEDGE 

AND THE BRITISH ASSOCIATION FOR THE ADVANCEMENT OF SCIENCE 

And carried on with the assistance of those Bodies and of the Trustees of 

THE BERNICE FAUAHI BISHOP MUSEUM AT HONOLULU. 



EDITED BY 

DAVID SHARP, M.B., M.A., F.R.S. 

SECRETARY OF THE COMMITTEE. 



VOLUME II. PART I. 

ORTHOPTERA 
By R. C. L. PERKINS, B.A. 

Pages I — 30; Plate I coloured and Plate II uncoloured. 



CAMBRIDGE: 

AT THE UNIVERSITY PRESS. 

1899 

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ORTHOPTERA 

By R. C. L. PERKINS, B.A. 



ORTHOPTERA. ''■^■t^- 

I, ''0. 



'^'.r-' 



By R. C. L. Perkins. 



c. 



§ 1. General considerations on the Orthoptera. 

The Hawaiian Orthoptera, although not very numerous in species, are neverthe- 
less an interesting and important group. In all seventy-three species are at present 
known, but of these a large proportion have no place in the indigenous fauna. Of 
the seven great divisions of the Order, the Phasmodea alone are altogether unrepre- 
sented. The Mantodea and Acridiodea have each a single species, both of which are 
known from elsewhere, and have certainly been imported by man. 

The Dermaptera and Blattodea are more numerously represented, and together 
form a considerable portion of the whole fauna, but nearly all the species are well-known 
insects outside the islands, and the importation of many of them has been rendered easy 
by their domestic habits. In the Dermaptera there are 4 genera represented, and 
7 species, none of which are endemic ; but they are nevertheless of some interest from 
the fact that some of them are in all probability natural immigrants. Thus Anisolabis 
pacifica has been found only on one of the islands, and there only at a considerable 
elevation above the sea in the mountain forests, where it is well established. Some 
of the species too exhibit very considerable variation, when a series of examples is 
examined, but whether these species are similarly variable in other countries I do not 
know, as the material necessary for such a comjaarison has not been available. 

The Blattodea are a heterogeneous collection of 1 5 species, four of which belong 
to the genus Phyllodromia, the remaining eleven representing, each one, a different genus. 
Three species are not at present identified from any locality outside the islands, but in 
spite of this, two of them {Phyllodromia hospes and Loboptera extraned) will certainl)' 
prove to be recent importations, both being found in company with the foreign forms 
in the neighbourhood of houses, and in localities, where no native fauna now exists. 
The third species {Phyllodromia obtusatd), on the other hand, is no doubt truly in- 
digenous, frequenting the mountain forests, where it is found amongst the leaves of 
native plants, and beneath the bark of the larger trees. This species is notably variable, 
and on some of the islands the variation is in a definite direction, so that it would 

F. H. II. I 



2 FAUNA HAWAIIENSIS 

appear to be in the process of becoming differentiated into other distinct species. All 
the other species are well-known foreign forms, and in the islands they are generally 
found in the neighbourhood of settlements, and often within the houses themselves, 
although some have now spread widely over the lowlands and the lower slopes of the 
mountains. 

Thus of the 24 species representing the four primary divisions of the Order thus far 
considered, one only has any claim to be considered indigenous, the rest, with the 
exception of two or three species, which may possibly be natural immigrants, having 
certainly been imported by man. 

In striking contrast with these are the remaining two divisions of the Order, viz. 
the Locustodea and Gryllodea. 

The Locustodea are represented by 13 species distributed in 4 genera. Two of 
these genera, each with a single species, are foreign, and both are certainly recent 
importations. The Elimaea has now spread throughout the group, over the lowlands, 
but the XipJiidiwn so far has been found only in Honolulu and the immediate 
neighbourhood, and its introduction is probably of very recent date. The other two 
genera, one with a single species, the other with ten, are peculiar to the islands, 
and they are evidently allied to one another, and the more highly peculiar Brachy- 
metopa with its 10 species may well have developed from some such form as the genus 
Conocephaloides. 

The Gryllodea are of great interest, and are the most important and extensive 
section of the Hawaiian Orthoptera. Ten genera are known, and these include 36 
species. Four of the latter may be at once set aside, as being of foreign origin, and 
each of these four represents a different genus. The three species Gryllus innotabilis, 
Gryllodes poeyi, and Gryllotalpa africana are all well-known elsewhere, and a Myrme- 
cophila, although now described as new, is certainly a recent importation, since it lives 
only in the nests of foreign ants, and has been found only in the city of Honolulu. The 
remaining 32 species are all peculiar to the islands, and are equally distributed between 
two divisions of the Gryllodea. The 16 species of the Trigonidiides are all referable to 
the genus Paratrigonidiuni, and no doubt others of this genus yet remain to be dis- 
covered on some of the islands. Elsewhere the genus is known from Asia. The 
other 16 indigenous species of Gryllodea are distributed in 5 allied genera, which are 
themselves also peculiar to the islands, and are so highly remarkable as to warrant the 
formation of a separate group for their reception. Three of the genera are represented 
by only a single species, but there is hope that others may yet be discovered, since 
neither genus has yet been found on more than one of the several islands. Of the 
other two genera, PrognatJiogryllus contains five, and Leptogryllus 8 species ; the former 
has occurred on Kauai and Oahu only, while the latter is widely distributed, Kauai 
and Hawaii, at either end of the chain of forest-bearing islands, each having more than 
one species. 



ORTHOPTERA 3 

The contrast between the indigenous portion of the Hawaiian Orthoptera and the 
foreign (whether introduced or naturally immigrant) is most strongly marked. 

Thus the 29 foreign species are scattered through all the six great divisions of the 
Order represented in the islands, while the endemic, to the number of 44, are limited to 
three of these divisions. Moreover, the foreign species represent no less than 24 genera, 
with an average of little more than one species to a genus, whereas the indigenous 
represent but 9 genera, with an average of nearly five species to the genus. 

Confining our attention to the indigenous portion of the fauna, the number of 
species which fail to extend their range beyond a single island is quite remarkable, more 
so, I believe, than is the case with any of the other Orders of insects. 

The Phyllodroinia is found throughout the whole group, but, as has been remarked, 
on certain of the islands the examples vary in a definite direction, as if tending to form 
distinct species. 

In the Locustodea the unique Conocephaloides has been found only on 
Hawaii, and of the 10 species of Brachymetopa not one is common to any two 
of the islands. 

Of the Trigonidiides two species of Paratrigonidhun [P. varians and P. pacificum) 
are widely, the latter indeed universally, distributed over the 6 larger islands, but it 
should be noted that both these species exhibit local variation, and owing to the great 
difficulty of differentiating the species of this genus, it is possible that more than one is 
included under each of these names. So far as is known each of the remaining 14 species 
limits its range to one or other of the islands. 

In the Prognathogryllides the genus Leptogi-yllus alone has species with a range 
extending over more than one of the islands, L. nigrolineatus having occurred on Oahu 
and Maui, and L. forficularis on Maui and Hawaii. 

Thus of the 44 endemic species five only have been found to inhabit more than 
one of the islands, but the foreign species, excepting a few, which have certainly been 
only very recently imported, are mostly of general distribution over the group. 

There are now added 44 species (35 being truly indigenous) to the list of Ortho- 
ptera, published by Herr Brunner in his paper in the Proceedings of the Zoological 
Society for 1895, wherein 29 species (9 only being indigenous) were enumerated. We 
are very much indebted to Herr Brunner for help in the identification of some foreign 
species discovered since 1895, as well as for the great assistance we have derived from 
the paper alluded to. 

It may be noticed that a large Phasmid Anchiale confusa Sharp {CypJiocrania 
manilata West.), brought home by the Beechey expedition, is said to have been taken 
in the Sandwich Islands, as also are several other large and conspicuous Orthoptera of 
other families. Certainly none of these have any place in the fauna of these islands, 
although they may have been taken in those other islands in the Pacific known under 
the name of Sandwich. 



FAUNA HAWAIIENSIS 

§ 2. Systematic account of the Orthoptera. 

DERMAPTERA. 

Anisolabis Fieb. 

(i) Anisolabis littorea White. 

Anisolabis littoj^ea, Brunner P. Zool. Soc. London, Dec. 1895, ?• ^92- 
Hab. Oahu and Maui (Blackburn). I have never met with this species. 

(2) Anisolabis maritinia Bon. 

Anisolabis maritinia, Brunner I.e. 

Hab. Common all over the group, in the mountains. 

(3) Anisolabis pacifica Erichs. 

Anisolabis pacifica, Brunner I.e. 

Hab. In the mountains of Kauai ; not found elsewhere. 

(4) Anisolabis annulipcs Luc. 

Anisolabis anmilipes, Brunner I.e. 

Hab. Generally abundant, both on the plains and in the mountains. 

Labia Leach. 

(i) Labia pygidiata Dubr. 

Labia pygidiata, Brunner I.e. 

Hab. Widely distributed ; found under bark of trees in the mountains. 

Chelisoches Scudder. 

( I ) Chelisoches mo'rio Fab. 
Chelisoches morio, Brunner I.e. 
Hab. Taken on nearly all the islands, usually between the leaves of Freycinetia, 



Dn 



or uracaena. 



ORTHOPTERA 5 

Sphingolabis Borm. 

(i) Sphingolabis hawaiiensis Borm. 
Forficula kazvaiiensis, Brunner I.e. 
Hab. Common generally throughout the islands. 

BLATTODEA. 

Phyllodromia Serv. 
(i) Phyllodromia germanica L. 

Phyllodromia germanica, Brunner Nouv. Syst. Blatt. p. 90. 
Hab. In houses. 

(2) Phyllodromia hieroglyphica Brunn. 
Phyllodromia hieroglyphica, Brunner P.Z.S. Dec. 1895, p. 892. 
Hab. Found on several of the islands. 

(3) Phyllodromia hospes, sp. no v. 

Tota testacea. Frons inter oculos nonnunquam plus minusve obscurata vel rufes- 
cens. Pronotum paullo latius quam longius, antice angustatum, margine postico rotun- 
dato. Tegmina testacea, venis testaceis. Alae hyalinae (antice testaceae), venis 
pallidis. Vena ulnaris 4-ramosa. Abdomen unicolor, testaceum. Lamina supra-analis 
$ apice obtuso, late levissimeque emarginato ; lamina subgenitalis ampla. $. 

Long, corporis, 1 3 mm. ; Long, pronoti, 3 mm. ; Lat. pronoti, 4 mm. ; Long, 
tegminum, 1 1 mm. 

The normal number of branches of the vena ulnaris appears to be 4, but some 
examples have only 3, and others 5. The left and right wings sometimes differ in the 
number of branches. The lamina supra-analis is sometimes hardly more than truncate 
at the apex. 

Hab. Kauai, and Honolulu. Found in houses, and under stones on the plains, 
and is no doubt an imported species. Herr Brunner has informed us that it is allied to 
P. conspersa Br. 

(4) Phyllodromia obticsata Brunn. 

Phyllodromia obhisala, Brunner P.Z.S. Dec. 1895, p. 892. 

Hab. Found throughout the group in the mountain forests. The species varies 
greatly ; examples from Molokai are remarkable for their small size, those from Kauai 
are the largest, examples from Hawaii being on the average intermediate. 



6 FAUNA HAWAIIENSIS 

LoBOPTERA Brunn. 

(i) Loboptera extranea, sp. nov. 

Nigrescens vel castanea. Frons inter oculos pallide signata. Antennae pallidae, 
plus minusve fuscescentes. Pronotum subtransversum, margine antico lateribusque 
flavo-niarginatis. Tegmina lobiformia, marginibus internis contiguis, lateribus flavo- 
marginatis. Abdomen utrinque flavo-marginatum. Coxae pallidae, nigro-notatae ; 
femora omnia (cum tibiis) pallida, nigro- vel fusco-notata, postica spinis compluribus 
bilineatim subtus armata. Cerci ad basim nigricantes, apicibus pallidis. Lamina supra- 
analis [$ $) transversa, triangularis, parum producta ; lamina subgenitalis $ ampla, 
medio margine apicali saepe levissime exciso. $^. Long. 8"5 — 9"5 mm. Tegmen 
2 '5 mm. 

Hab. Maui, on the coast. Hilo, Hawaii (Bro. Matthias Newell). Probably on 
all the islands, generally living in company with the young of Periplaneta australasiae. 
In the development of the tegmina the sexes are quite alike. The wings are repre- 
sented by pale lobes at the sides of the metanotum, the inner margins being marked by 
a deep suture, but not free. 

Stylopyga Fisch. 
(i) Stylopyga decorata Brunn. 
Stylopyga decorata, Brunner P.Z.S. 1895, p. 893. 
Hab. Honolulu (Blackburn). 

Methana StSl. 

(i) Methana ligata Brunn. 

Methana ligata, Brunner I.e. 
Hab. Honolulu (Blackburn). 

Polyzosteria Burm. 

(i) Polyzosteria soror Brunn. 

Polyzosteria soror, Brunner Nouv. Syst. Blatt. p. 219. 
Hab. Common, in company with P. aiistralasiae. 

Periplaneta Burm. 

( I ) Periplaneta americana L. 

Periplaneta americana, Brunner P.Z.S. 1895, P- 893- 
Hab. Abundant throughout the islands. 



ORTHOPTERA 7 

(2) Periplaneta australasiae Fab. 
Periplaneta australasiae, Brunner Nouv. Syst. Blatt. p. 233. 
Hab. Abundant throughout the islands. 

Eleutheroda Brunn. 
(i) Eleutheroda dytiscoides Serv. 
Eleutheroda dytiscoides, Brunner P.Z.S. 1895, p. 893. 

Hab. a common and injurious species ; very abundant in Honolulu, and else- 
where. 

Leucophaea Brunn. 
(i) Lejtcophaea surinamensis Fab. 
Leucophaea surinamensis, Brunner I.e. 
Hab. Oahu, Kauai, Maui ; and probably on all the islands. 

Nauphoeta Burm. 

(i) Nauphoeta bivittata Burm. 

Nauphoeta bivittata, Brunner Nouv. Syst. Blatt. p. 287. 
Hab. Two examples taken under bark of trees in Honolulu. 

Oniscosoma Brunn. 

(i) Oniscosoma pallida Brunn. 

Oniscosoma pallida, Brunner P.Z.S. 1895, p. 893. 
Hab. Haleakala, Maui (650 m.). Blackburn. 

EuTHYRRHAPHA Burm. 

( I ) Euthyrrhapha pacifica Coqueb. 

Euthyrrhapha pacijica, Brunner I.e. 

Hab. Taken on several islands, and probably to be found on all. 

MANTODEA. 

Orthodera Burm. 

(i) Orthodera prasina Burm. 
Hab. Kauai ; introduced with fruit trees. 



8 FAUNA HAWAIIENSIS 

ACRIDIODEA. 
OxYA Serv. 

(i) Oxya velox Fab. 

Oxya velox, Brunner P.Z.S. Dec. 1895, p. 893. 

Hab. Abundant on Kauai and Oahu, but had not spread to the other islands 
in 1897. 

LOCUSTODEA. 

Elimaea Stal. 

(i) Elimaea appendiculata Brunn. 

Elimaea appendiculata, Brunner I.e. 

Hab. Very abundant throughout the islands, on the plains and lower slopes of 
the mountains. 

Brachymetopa Redt. 

The ten species, which represent this genus, are closely allied to one another, 
and for the most part very similar in general appearance. The genus, though peculiar 
to these islands, is allied to the widely distributed Coiioccp/ialus, but still more closely 
to the Hawaiian genus Conocephaloides, described hereafter. The latter indeed, with 
the general appearance of a Conocephalus, combines the characters of that genus with 
some of those which distinguish Brachymetopa from it. All the species of Brachy- 
metopa have the tegmina and wings in a more or less rudimentary condition, and they 
are useless for purposes of flight in either sex, but serve as stridulating organs in 
the <J. In B. deplanata Z the tegmina are extremely short, being only as long as 
the pronotum, but in some others they extend to the apex of the abdomen, while 
in others again they are intermediate in length between these extremes. 

Several, and perhaps most, of the species have two distinct forms, a green and 
a darker (or at least not green) one, which are so strikingly different as to suggest 
at first sight that they are distinct species. There appear to be no really intermediate 
forms, although the examples that are not green are themselves variable, the colour 
varying from testaceous or ochreous to dark fuscous in some species. It is quite 
possible that the two forms are really tending to become distinct species, at least 
in certain cases, and the two Oahuan species B. discolor and B. blackbjcjnti may not 
improbably have originated from a single simply dimorphic species. Certain it is that, 



ORTHOPTERA g 

although numerous examples of both species have been examined, no green form of 
the former, or not-green one of the latter has yet been discovered. In this respect 
they stand alone, for of all the other species which have been examined in such 
numbers as these, the two distinct forms have been obtained. All the species are 
nocturnal in their habits, and rest by day on the leaves of various forest trees or 
plants. The males stridulate freely at dusk and through the night, and can be heard 
for a long distance, but it is extremely difficult to locate them by sound, and we 
have only occasionally been able to do so. 

The modified cerci of the males, which form organs for copulation, in most cases 
furnish excellent specific characters. 

(i) Brachymetopa discolor Redtb. 

Brachymetopa discolor', Redtenb. Verb. z. b. Ges. Wien, 1891, p. 431 ; Brunner 
P. Z. S. Dec. 1895, p. 894. 

(Plate I. figs. I & ii? ; and Plate II. figs, i & \a.) 

This is a very distinct species, and may be known at once from dark varieties 
of several of the green species by the shining coal-black front of the head, and the 
black-spotted legs. 

Hab. Both mountain ranges of Oahu. Mts. near Honolulu (1500 — 2000 ft.). 
Waianae Mts. (2000 ft.). 

(2) Brachymetopa blackburni Bormans. 

Conocephalm blackbtirni de Bormans Ann. Mus. Genova, xviii. 1882, p. 346. 

Brachymetopa blackburni Redtenbacher Verb. Ges. Wien, 1891, p. 431 ; Brunner 
P. Z. S. 1895, P- 894- 

(Plate I. fig. 2.) 

Hab. Mountains near Honolulu, 1500 ft. and upwards. 

De Bormans says that this species is found on several of the islands, but the 
examples taken on islands other than Oahu, no doubt belong to different species. 
Although I have frequently taken the adult ?, and young of both sexes, I have never 
seen an adult Z. 

(3) Brachymetopa nitida Brunn. 

Brachymetopa nitida, Brunner Proc. Zool. Soc. Lond. 1895, P- 894- 
(Plate I. figs. 3, 3«, & lb; and Plate II. figs. 2 & 2a.) 

Hab. The original examples were taken in Kona, Hawaii. I have since collected 
it on the lower slopes of Mauna Kea, above Hilo, and freely at Olaa in the Puna 
F. H. II. 2 



lo FAUNA HAWAIIENSIS 

district. In neither of these localities do the specimens agree with the typical ones, 
or with each other, but owing to the variation exhibited, they cannot be regarded 
as distinct species. 

Brachymetopa nitida, van hildensis, van nov. ^ with the tegmina shorter than 
in typical specimens, the greatest length shown by these examples being only lo mm. 

Brachymetopa nitida, var. ptmae, var. nov. Generally larger than the other forms, 
attaining; the followinsf dimensions. 

Long, corporis (haud contracti) $ 25, $ 27 mm. ; pronoti $ 6'5, ? 7 mm. ; tegminum 
$ II, ? II — 13 mm.; femorum post. $ 16, $ 17mm.; ovipositoris 12 mm. 

One or two of the $$ however have the femora of only 13 mm. and therein 
agree with the var. hiloensis, but still remain distinct from the typical specimens by 
the short tegmina, the length of which is only 8 mm. 

Brachymetopa nitida, var. crassipes, var. nov. The tibiae are evidently stouter 
than in the other forms. $. 

Long, corporis 23 mm. ; pronoti 6'5 mm. ; tegminum io'50 mm. ; femorum 
post. 13 mm. 

A single ^ was taken at Kilauea (4000 ft.). This is the highest elevation at 
which the genus has occurred. 

The ^ characters seem to be identical in the various forms ; the upper hooks of 
the cerci are bent upwards, the lower are very long and rise up behind the former 
to a considerably higher level. 



(4) Brachymetopa kauaiensis, sp. nov. 

Viridis. Mandibulae in parte apicali nigrae. Labrum cum fronte tota pallidum. 
Tegmina abbreviata, tympano $ minus infuscato, haud nitido. Tibiae posticae utrinque 
spinulis armatae. Unci cercorum $ inferiores decumbentes. $'^. 

Long, corporis $ 19, $ 21 mm.; fastigii vert. $ r2, ? 1*5 mm.; pronoti $ 5'5, 
$ 6 mm.; tegminum ^ 775, ? 8 mm. ; femorum post. $ 12, $ 13 mm. ; ovipositoris 
1 1 mm. 

Hab. Makaweli, Kauai (2000 ft.). Rare, i $ taken in 1895, ^^^ i <? in 1897. 



(5) Brachymetopa unica, sp. nov. 

Viridis. Mandibulae pallidae, margine interno nigro. Frons media plaga lata 
triangulari rufo-brunnea ornata. Fastigium verticis latum, apice fere truncato. Teg- 
mina ad apicem abdominis se extendentia, subacuminata, densissime venosa. Tibiae 
posticae interne spinis raris, externe fere nullis (una tantum), armatae. ?. 



ORTHOPTERA ii 

Long, corporis 20 mm. ; fastigii vert, i -5 mm. ; pronoti 6 mm. ; tegminum 1 1 mm. ; 
femorum post. 12 mm. ; ovipositoris io'5 mm. (Plate I. fig. 4.) 

Hab. Mountains near Honolulu, Oahu. A single ? taken in 1S96, amongst 
B. blackburni and B. discolor. It is a very distinct species, and the tegmina with 
their somewhat sharp apices with the margins narrowly reddish, are unlike any other 
species. 

(6) Brachymetopa affinis, sp. nov. 

Viridis. Mandibulae ad apices nigrae. Frons tota pallida. Tegmina sublonga, 
capite cum pronoto evidenter longiora. Tibiae posticae interne spinulis perpaucis 
armatae. $. 

Long, corporis 22 mm. ; fastigii vert. 2 mm. ; pronoti 6 mm. ; tegminum i r5 mm. ; 
femorum post. 1 3 mm. ; ovipositoris 1 1 mm. 

Hab. Mountains of Kauai, 3000 ft. One ? taken in June 1894. This individual 
was referred by Brunner to his B. deplanata, a species peculiar to Lanai. From an 
examination of the much larger material obtained during my second visit, I feel sure 
that no species of the genus is found on more than one island of the group. The 
present species is most closely allied to B. blackburni, but the absence of the distinct 
black transverse marks on the face at the base of the mandibles, and the less spinose 
posterior tibiae will easily distinguish it. The latter on their outer margin have only 
four spines. 

(7) Brachymetopa parviila, sp. nov. 

Laete viridis, subnitida. Mandibularum pars apicalis plus minusve nigra. Frons 
tota pallida. Tegmina capite cum pronoto baud longiora. Tibiae posticae externe 
spinulis nullis armatae. Unci cercorum $ superiores validi, fortiter erecti. Ovipositor 
$ perlongus, femore postico longior. $^. 

Long, corporis $ 18, ? 19 mm.; fastigii vert. ^ i, ? i mm.; pronoti ,? 5, ? 5"5 mm.; 
tegminum $ 7'5, ? 8 mm.; femorum post. $ 9*5, $ 11 mm.; ovipositoris 12-5 mm. 
(Plate L fig. 6.) 

Brachymetopa parvula, var. brunnea, var. nov. Corpus totum pallide brunneum. 
(Plate L fig. 6a.) 

Hab. Mountains of West Maui above Lahaina (3000 ft.). One $ and several ? 
taken ; var. brunnea a single pair taken off the same branch of a tree in the same 
locality. This is on the whole the smallest species of the genus. The male characters 
are quite distinct from those of any other species, the upper of the two processes of 
the modified cercus rises erect, and is both long and stout ; the lower one is much 
bent and directed backwards. In the % the ovipositor is of unusual length for the 
size of the species. 

2 — 2 



12 FAUNA HAWAIIENSIS 



(8) Brachymetopa mauiensis, sp. nov. 

Nitida, viridis. Mandibulae apices versus nigrae. Frons pallida, juxta basim 
mandibularum utrinque nigropunctata. Tegmina fortiter reticulatim venosa, capite 
cum pronoto breviora. Tibiae posticae externe inermes. Unci cercorum $ superiores 
breves, inferiores suberecti et his multo longiores. ^?. 

Long, corporis $ i8, $ 21 mm.; fastigii vert. ^ i, ? r25mm.; pronoti $ 5-5, 
$ 5"5 mm.; tegminum $ 8 — 8'5, $ 8-5 mm.; femorum post. $ 10, ? 11 '5 mm.; ovi- 
positoris 10 mm. (Plate I. fig. 5; and Plate II. figs. 4 & ^a.) 

Brachymetopa mauiensis, var. ochracea, var. nov. % major (long. 24 mm.), colore 
ochraceo. 

Hab. Maui. West Maui Mts. in the lao Valley. Z%- Haleakala (4000 ft.) t- 
The ti from the latter locality have the tegmina slightly longer, but otherwise agree 
exactly with that from the distant western mountains. Var. ochracea, lao Valley i %. 



(9) Brachymetopa molokaiensis, sp. nov. 

Viridis, praecedenti cognatissima, sed minus nitida. $ unco superiore cercorum 
multo longiore et fortiore : ? tegminibus apices versus magis longitudinaliter venosis, 
minus reticulatis, bene distinguenda. <??. 

Long, corporis $ 20, ? 25 mm.; fastigii vert. ^ i"5, ? 175 mm.; pronoti $ 5*5 — 6, 
^ 6 mm.; tegminum $ 8*5 — 10, ? 10 mm.; femorum post. $ 1 1 '5, ? i2"5 mm.; ovi- 
positoris 1 1"5 mm. 

Hab. Mountains of Molokai (3000 — 4000 ft.). 

The species here described was assigned partly {$) to B. nitida, and partly (?) to 
B. deplanata by Herr Brunner. The Z is quite distinct from the former by the 
unarmed outer margin of the posterior tibiae, and by the cerci, and the % taken in 
the same locality agrees so well in most respects with the Z that I have little doubt 
of its identity. 

(10) Brachymetopa deplanata Brunn. 

Brachymetopa deplanata, Brunner Proc. Zool. Soc. Lond. 1895, p. 894. 

(Plate I. fig. 7; and Plate II. figs. 3 & 3«.) 

The Z of this species (I have not seen an adult $) is very distinct from any other, 
by the very short tegmina, which are subequal to the pronotum in length, and the 
cerci are also peculiar, the upper hook being simply transverse in direction and slightly 



«l 



I 



ORTHOPTERA 13 

curved, the lower is also transverse and decumbent with its apex inclined downwards. 
The females assigned to this species in the original description, I have otherwise 
disposed of (vide B. affinis and B. molokaiensis). 
Hab. Lanai (2000 ft.). 

DiSPOSITIO SPECIERUM GENERIS BRACHYMETOPAE. 

1. (<?) Tibiae posticae utrinque spinulis multis armatae. 

2. (j) Frons capitis tola pernigra discolor. 

3. (2) Frons pallida. 

4. (j) Mandibulae subtotae nigrae ; frons capitis juxta basim mandibularum 

utrinque linea nigra transversa signata blackburni. 

5. {4) Mandibulae ad basim pallidae ; frons capitis haud nigrolineata. 

6. (7) $ uncus cercorum inferior decumbens ; ? tegmina capite cum pronoto 

evidenter breviora kaiiaiensis. 

7. ((5) $ cercorum uncus inferior erectus ; 9 tegmina capite cum pronoto 

haud evidenter breviora nitida. 

8. (7) Tibiae posticae in margine interno inermes vel perpaucis spinulis 

(4 ad max.) armatae. 

9. {10) Tegmina i brevissima, pronoto longitudine subaequalia deplanata. 

10. (p) Tegmina $ pronoto evidenter longiora. 

11. {12) Frons capitis plaga magna triangulari rufo-brunnea ornata. Tegminum 

apices subacuminati laiica. 

12. (11) Frons unicolor ; tegminum apices rotundati. 

13. {14) Tegmina capite cum pronoto evidenter longiora affinis. 

14. (/j) Tegmina capite cum pronoto haud longiora. 

15. {16) Uncus superior cercorum $ erectus, supra inferiorem elevatus. $ ovi- 

positor femore postico longior .parviila. 

16. (/J) (J uncus superior cercorum haud erectus. ? ovipositor femore 

postico brevior. 

1 7. {18) Magis nitida. $ uncus superior cercorum brevis. $ tegmina apices 

versus fortiter reticulatim-venosa tnauiensis. 

18. (77) Minus nitida. S uncus superior cercorum minus brevis. 9 tegmina 

apices versus ex majore parte longitudinaliter venosae molokaiersis. 



CoNOCEPHALOiDES, gen. nov. 

A genere Conocephalo differt lobis genicularibus inermibus, a Bi^achymetopo lobis 
meso- et metasterni angustis, femoribus posticis perlongis, intermediis ter longioribus, 
alis tegminibusque perfecte explicatis, capite et corpore toto (excepto ovipositore) 
multo longioribus. 

(i) C. hawaiiensis, sp. nov. 

Viridis. Mandibulae flavae. Labrum pallidum, roseotinctum. Fastigium verticis 
inferum haud in spinam productum, vix longius quam latius, apice rotundato. Anten- 
narum articulus basalis viridis, caeteri subtestacei, apicem versus fuscescentes. Pronotum 



14 FAUNA HAWAIIENSIS 

supra utrinque fusco-lineatum, lateribus flavomarginatis. Alae tegminibus vix bre- 
viores. Femora omnia viridescentia, posticorum marginibus inferioribus spinis armatis. 
Tibiae pallide fuscescentes, posticae biseriatim spinosae. Tarsi graciles. Ovipositor 
rectus. %. 

Long, corporis cum ovipositore 46 mm. ; pronoti 8 mm.; tegminum 38 mm.; fern, 
post. 23 mm.; ovipositoris 17 mm. (Plate I. fig. 8.) 

Hab. Olaa, Hawaii (2000 ft.); i % taken, Dec. 1896. 

XiPHiDiUM Serv. 

( I ) Xiphidium fusctini Fab. 

Xiphidiuni fusctun, Brunner P. Z. S. 1895, p. 894. 
Hab. Only in and around Honolulu. 

GRYLLODEA. 

Gryllotalpa Latr. 

( I ) Gryllotalpa africaua Fab. 

Hab. This introduced species abounds on the windward side of Oahu, and is 
injurious on the sugar plantations. 

Myrmecophila Latr. 

(i) Myrmecophila quadrispina, sp. nov. 

Atrofusca, ore, antennis, pedibus, cercisque, vel nonnullis ex his, pallidis. Frons 
capitis breviter pilosa. Corpus subtilissime sericeo-pubescens. Cerci conspicue pallido- 
pilosi. Femora postica obscurata, fortissime dilatata ; tibiarum posticarum marginibus 
interioribus spinis quattuor armatis, prima (sive basali) brevi, secunda quartaque multo 
longioribus, tertia, quam spina basalis quoque, evidenter multo breviore. Z ?• Long. 
2'2 mm. (Plate H. fig. 7.) 

Hab. Honolulu, in gardens in the city. An imported species, living in the nests 
of foreign species of ants. 

Gryllus L. 

(i) Gryllus innotabilis Walk. 

Gryllus innotabilis, Brunner P. Z. S. Lond. 1895, p. 895. 
Hab. Abundant on all the islands. 



ORTHOPTERA 15 

Gryllodes Sauss. 

(i) Gryllodes poeyi Sauss. 
Gryllodes poeyi, Brunner loc. cit. 

Hab. Common generally on the plains, and lower slopes of the mountains. 
Obs. An immature example of a second species of Gryllodes was obtained at 
Lahaina, Maui, but no adult. 

Paratrigonidium Brunn. (1893). 

The sixteen species of this genus are closely allied to one another, and in some 
cases the specific characters are difficult to appreciate. This is partly due to the varia- 
bility of some of the species, but also to the changes which take place after the death 
of the insect in the process of drying. Thus P. viridescens in life is quite different in 
appearance (Plate I. fig. 11) to any other species of the genus, and its peculiar habits 
leave no doubt that it really is distinct, yet in dried examples its resemblance to several 
other species is extreme. P. pacijicum, the first-described Hawaiian species, is in some 
respects the most aberrant of the series, since it differs from all the others in the structure 
of the ^ genitalia, and of the % ovipositor, as well as in the neuration of the lateral field 
of the tegmina (Plate II. fig. to). 

As to the absence of wings in this genus, it is noteworthy that although in 
the adult insect the metathoracic wings are entirely wanting, yet in the penultimate 
state they are as well developed as the mesothoracic (or tegmina). Thus both the 
tegmina and the wings develope equally during the earlier stages up to the last 
ecdysis, when the tegmina undergo a further and very great developement, while the 
wings, on the other hand, not only cease to develope, but entirely abort. 

The habits of the various species are interesting, and a large number are found to 
inhabit only one particular species of tree or plant. Thus P. viridescens and P. filicum 
live amongst ferns, but each is found only on one particular species of fern. P. saltator 
hides at the bases of the leaves of Freycinetia, P. atroferrugineiini amongst those of 
Metrosideros. Several species are found only beneath the bark of large trees, but 
P. pacifiaim is to a large degree terrestrial. Most of the species are nocturnal in their 
activities, others are active by day, but generally only in damp shady gulches and dense 
forests. P. pacijicum in such situations sings the whole day through, with a plaintive 
chirp, but at night all the species are most active and noisy. There is a considerable 
difference between the songs of some of the species, and in some cases the sound can 
be heard at a great distance. The power of leaping of most of the species is prodigious, 
and the time that must be spent to collect a series of the most active ones is very 



1 6 FAUNA HAVVAIIENSIS 

considerable. The ground-frequenting P. pacificum is on the whole the most easily- 
captured, the arboreal species in spite of their small size will clear several feet in one 
jump, and several leaps will be made with astonishing rapidity. After this however 
the distance covered at a leap becomes shorter and shorter till eventually the power of 
leaping is nearly entirely lost, and the insect attempts to escape by running. 



( I ) Paratrigonidium freycinetiae, sp. nov. 

Frons pallida. Antennarum articulus primus et secundus nigricantes. Pronotum 
atrum, postice nonnunquam pallidius. Femora antica et intermedia nigricantia. Femora 
postica supra nigra ; ad apices etiam et externe et interne nigra, ad basini pallida. 
Tibiae pallidae, anticae et intermediae distincte nigronotatae. Tegmina, cerci, et 
antennae (basi excepta) pallide flavescentia. $. 

Long, corporis 6 mm., pronoti i'^ mm., tegminum 4'2 mm., femorum post. 
5 "3 mm., ovipositoris 2 mm. 

Hab. Olaa, Hawaii (2000 ft.). Very rare, three females taken from Freycinetia. 
I saw one .?, but was unable to capture it. It is excessively wild and active. 

(2) Paratrigonidmm saltaior, sp. nov. 

Frons cum pronoto pallida. Femora antica et intermedia nigra ; femora postica 
supra nigricantia, externe interneque ex majore parte pallida. Tibiae intermediae anti- 
caeque, plus minusve distincte, maculatae. Tegmina tota pallide flavescentia. Anten- 
narum articulus primus sat pallidus. $. (Long. sp. praecedentis.) 

This species is closely allied to the preceding, and only the male sex of the one and 
the female of the other was taken, but the different colour of the pronotum, and of the 
hind femora, will readily separate them. The superior tegmen of P. saltator is uni- 
colorous, pale yellowish, with no dark mark adjoining the vena stridulans, and the front 
and intermediate femora are entirely of a dark pitchy or black colour. 

Hab. Mountains of Oahu, above 2000 ft. Rare. Apparently attached to the 

Freycinetia. 

(3) Paratj'igonidi^im roseuni, sp. nov. 

Caput cum articulo primo antennarum, et femora omnia laete rosea. Tibiae tarsi- 
que pallidi. Tibiae posticae roseo-tinctae. Pronotum ex majore parte nigrum. An- 
tennae, tegmina, et cerci pallide flavescentia. %. 

Long, corporis 7 mm., pronoti I's mm., tegminum 4'5 mm., femorum post. 5 mm., 
ovipositoris 2*5 mm. (Plate L fig. 9.) 



ORTHOPTERA 



17 



Hab. Mountains of West Maui (3000 ft.). A single ? taken from Metrosideros. 
Like the preceding it is a very quick and strong jumper and difficult to seize. I saw- 
several other specimens, when the individual described above was taken, but was un- 
able to revisit the locality. 

(4) Pai'atrigonidium atroferrugineuni Brunn. 

Paratrigonidium atroferrugineuni Brunner P. Z. S. 1895, P- 895. 

(Plate I. fig. 10.) 

This is a very distinct species, and cannot be confused with any other. The black 
tegmen of the Z with its orange-coloured border, and the black femora with red apices 
in both sexes, give it a very distinctive appearance. The tegmina of the % are generally 
entirely pale, but sometimes black in the centre with a ferruginous border, very much 
as in the ^. 

Hab. Island of Molokai only. Lives in the leaves of Metrosideros polymorpJia, 
but chiefly if not solely in those of one special form of this very variable tree. The 
young of the cricket is entirely green in all its stages. 

(5) Paratrigonidium subroseum, sp. nov. 

Totum pallidum, plus minusve roseo-tinctum, femoribus tibiisque omnibus imma- 
culatis. Z %. 

Long, corporis t 7'5. ? r'5 mm. ; pronoti t i'5, ? i'5 ; tegminum t 5'6, ? 4"8 mm.; 
femorum post. ? 5, ? 5'2 mm. ; ovipositoris 2-5 mm. 

This is a very distinct species. There is no dark spot on the tegmen behind the 
vena stridulans, such as is present in most of the Hawaiian species. In the immature 
stages the insect is entirely green. After death the rosy pigment has a tendency to 
become massed in certain parts, as is also the case with the green pigment in other 
species, but the legs are in reality quite unspotted. 

Hab. Mountains of Oahu (2000 ft.). Attached to bushy trees o{ Metrosideros. 

(6) Paratrigonidium Jilicum, sp. nov. 

Brunneum, testaceum, vel olivaceum. Frons pallida, nonnunquam notis fuscis, 
plerumque minus distinctis, ornata. Antennae subinfuscatae, basim versus articulis 
paucis, rare dispositis, obscurioribus. Tibiae anticae minus distincte fusco-notatae. 
Femora postica interna fere concoloria, vel tantum duabus notis minus distinctis fuscis 
signata. Tibiae posticae supra pallidae. t Tegmen de.xtrum post venam stridulantem 
plaga triangulari nigra ornatum : hac, majore ex parte, laevi, minus rugosa. Cerci 
minus elongati. (Plate II. fig. 6.) <? $. 

F. H. II. 3 



1 8 FAUNA HAWAIIENSIS 

Long, corporis (cum tegminibus) t 7'2, ? 67 mm. ; tegminum ^ 5-5, % 4*5 mm. — 
Lat. tegminum ^ 3 mm.— Long, femorum post, t 5'4, % 5'8 mm.; ovipositoris 27 mm.; 
cerci % 3 mm. 

Hab. Olaa, Hawaii (2000 ft.). In dense forest frequenting a tall soft fern, which 
covers the ground beneath the trees. Distinguished from the two following species by 
the much less distinct markings of the legs generally, and the almost entirely pale pos- 
terior tibiae. Immature examples are entirely green. 



(7) Paratrigonidium viridescens, sp. nov. 

Pallidum, subviridescens (vivum eleganter viride), nigro-ornatum. Frons nigro- 
notata. Pronotum lateribus nigris. Femora omnia cum tibiis distincte nigro-notata. 
Tegmen [$) dexterum maculis distinctis circa venam stridulantem, et saepe ad latera, et 
ad apicem plus minusve ornatum. $ $. 

Long, corporis J 7*5, $ 7 mm. ; pronoti $ i"5, ? i'5 mm.; tegminum $ y2, % 4"5 
mm. — Lat. tegminum $ 3 mm. — Long, femorum post. $ 5"2, $ 5'8 mm. ; ovipositoris 
2-5 mm. (Plate L fig. 11.) 

This species is very distinct in life, the general colour being then of a delicate 
light green. This unfortunately soon fades in dried specimens although a slight tint 
still remains. The most noticeable characters are the unusually sharply defined black 
markings, which vary in extent, but are scattered over the tegmen, and not entirely 
confined to the neighbourhood of the vena stridulans. The black marks on the legs are 
very distinct, and the tegmen is rather long in proportion to its width. 

Hab. Olaa, Hawaii, 2000 ft. Lives amongst a beautiful creeping fern, which 
clothes the tree trunks in wet forests. 



(8) Paratrigonidium varians, sp. nov. 

Colore brunneo vel testaceo. Frons pallida, lineis maculisque nigris vel fuscis 
variabilibus ornata. Antennae basim versus articulis nigricantibus, rare dispositis, 
variegatae. Pronotum maculis fuscis vel nigris supra ornatum, lateribus fusco- vel nigro- 
marginatis. Tegmen $ dextrum macula nigra post venam stridulantem signatum, ibique 
ex majore parte fere planum, vix rugulosum ; pars caetera (nonnunquam fere tota) plus 
minusve infuscata. Tibiae anticae et intermediae nigro-notatae ; femora postica intus 
satis distincte nigro- vel fusco-notata. Tibiae posticae juxta basim pallidae ; hac parte 
excepta, fere totae nigricantes, tantum ad basim spinarum pallidae. 

Long, corporis J6'5, ? 6"5 mm. ; pronoti ^ i'2, $ 1-2 mm.; tegminum ^ 5, $ 4*4 mm. — 
Lat. tegminum $ 2-5 mm. — Long. fem. post. ^ 5, $ 5*5 mm. ; ovipositoris 27 mm. 



ORTHOPTERA 



19 



This species greatly resembles P. viricicscens, in dried examples, although when 
fresh, and not discoloured, it has a totally different appearance. It appears to be a 
variable species, and the examples taken in different localities are rarely altogether 
similar. 

Hab. Puna (2000 ft), and Kau (4000 ft.), Hawaii ; mountains of West Maui ; 
Honolulu (2000 ft.); Makaweli, Kauai (2000 ft.). Apparently always rare. It is an 
arboreal species. 

(9) Paratrigonidiuni grande, sp. nov. 

Statura majore, colore testaceo vel brunneo. Frons cum pronoto plus minusve 
atro- vel fusco-notata. Antennae articulis multis nigricantibus variegatae. Tibiae 
anticae et intermediae distincte fusco- vel nigro-maculatae. Femora postica interna 
notis compluribus nigris ornata. Tibiae posticae supra ex majore parte fuscae vel 
subnigrae, ad basim spinarum pallidae. $ Tegmen dextrum post venam stridulantem 
plaga triangulari nigra, laevi, haud rugosa. $ $. 

Long, corporis (cum tegminibus) $ 9*5, $ 87 mm.; pronoti $ 17, $ 2 mm.; 
tegminum ^ 7, ? 575 mm. — Lat. tegminum $ 3'8 mm. — Long, femorum post. $ 6'2, 
$ 6'2 mm.; ovipositoris 3 mm. (Plate I. fig. 12.) 

Hab. Hawaii, in the Puna, Kau, and Kona districts. Lives beneath the bark 
of trees, coming outside only at night. The young are of the same colour as the 
adult, not green like those of the three preceding species. The large size, combined 
with the smooth area behind the vena stridulans of the $, will distinguish this species 
from any other. 

(10) P aratrigonidium crepitans, sp. nov. 

Testaceum, capite plus minusve fusco-variegato, antennarum articulis basalibus 
pallidis. Pronotum fusco-variegatum. Femora omnia distincte nigro-notata ; tibiae 
posticae variegatae. Cerci pallidi, breviores. Tegmen ^ dextrum post venam stridu- 
lantem macula nigra ornatum, hac plaga tota rugulosa. i%. 

Long, corporis (cum tegminibus) J" 7, ? 6 mm. ; pronoti % i "5 mm. ; tegminum 
t 5'5> ? \'- mm-; femorum post. ^ 5'2, % 5'5 mm.; cerci ^ 3'25 mm.; ovipositoris 
2'5 mm. 

Very like P. filicimi and the allied species in general appearance, but the $ is 
quite distinct by the evidently less smooth (more rugulose) dark area behind the 
vena stridulans. 

Hab. Kauai (4000 ft.), rare, or at least very difficult to obtain, living amongst 
heaps of dead wood, or in low vegetation. 

3—2 



20 FAUNA HAWAIIENSIS 

( 1 1 ) Paratrigonidium robustuni, sp. nov. 

Forma et colore fere praecedentis, sed major et cercis evidenter longioribus 
distinguendus. ^. 

Long, corporis 9 mm. ; pronoti i '6 mm. ; tegminum 6 mm. — Lat. tegminum 
3'5 mm. — Long, femorum post. 6 mm.; cercorum 4'6 mm. 

Very like the preceding species but decidedly larger and with longer cerci. The 
species is altogether more robust than most others of the genus. The black area 
behind the vena stridulans well-marked, and rugulose as in the preceding. 

Hab. Kauai (4000 ft.); i ,^ taken from beneath the bark of a tree. 

(12) Paratrigonidiiiin molokaiense, sp. nov. 

Testaceum, fronte inter antennas tota nigra. Clypeus totus vel ex majore parte 
niger. Articulus antennarum primus et secundus nigri vel picei. Pronotum cum 
capite inter setas distincte pallide pubescens. Femora omnia cum tibiis distinctissime 
nigro-notata ; femora postica notis minimis compluribus nigris intus ornata. Tegmen J 
dextrum plaga post venam stridulantem nigra, tota rugosa, maculisque compluribus 
postice, necnon in campo laterali, signatum. Tegmina % supra pallida, campo laterali 
toto nigricante. Cerci pallidi, minus longi. 1; %. 

Long, corporis J' 8, ? 7 mm.; pronoti $ 1-4, % 1-4 mm. ; tegminum ^ 5'5, 
% 4"5 mm — Lat. tegminum ^ 3"2 mm. — Long, femorum post. ^ 5*2, % 5"2 mm.; 
ovipositoris 27 mm.; cercorum % 3'2 mm. 

Certainly a distinct species, the hind femora (at least in the ^) rather more 
strongly clavate, than in most of the genus. 

Hab. Mountains of Molokai (3000 ft.); rare, i ^ i ? taken amongst low vege- 
tation. 

(13) Paratrigonidium attenuattmi, sp. nov. 

Pallide testaceum, angustum, elongatum, femoribus omnibus nigro-notatis. Teg- 
men T; dextrum post venam stridulantem plus minusve evidenter nigro-notatum, plaga 
nigra rugulosa, speculo angustissimo, fortiter elongato. ^ $. 









Var. 


majoi 








Var. minor, 


Long. 


corporis $ 


8, 




? 


8-25 


mm. 


$ 


7-25 mm. 




pronoti 


2 






2 


fi 




17 .. 




tegminum 


5-5^ 






5 


?) 




5 


Lat. 


?j 


3. 












275 .. 


Long. 


fem. post, 
ovipositoris 


6-5, 






6'S 
3-5 


J) 


- 


5-5 .. 



(Plate I. fig. 13; and Plate II. fig. 5.) 



ORTHOPTERA 21 

Apparently there are two forms, a larger and a smaller, of this species, the former 
being found at a greater elevation in the mountains than the latter. Except for 
the difference in size I see no other noticeable distinction between the adults, yet, 
if I am not mistaken, the immature forms of the larger examples are green, while 
those of the smaller are not. 

This species is easily distinguished by the very narrow speculum of the ^ tegmen. 
The large examples are not much less long than P. graiide, but the ^J is a much 
narrower insect. 

Hab. Kauai (4000 ft), the larger form. The smaller at an elevation of 2000 — 
3000 ft. Both live beneath the bark of large trees. 



(14) P aratrigonidiuni dcbi/c, sp. nov. 

Angustum, testaceum, fronte cum pronoto pallida, fusco-notata. Tegmen $ dextrum 
circa venam stridulantem nigro-maculatum. Tibiae anteriores et intermediae distincte 
fusco- vel nigro-notatae. Femora postica notis parvis nigricantibus nonnullis interne 
signata. Cerci pallidi. $. 

Long, corporis 6"8mm. ; pronoti V2 mm.; tegminum 4*5 mm. — Lat. tegminum 
2 '5 mm. — Long, femorum post. 5 mm. 

Hab. Mountains of Oahu (2000 ft. and upwards). 3 $ taken. 



(15) Paratrigonidiiun exiguum, sp. nov. 

Minimum, brunneum, fronte et pronoto variegatis, baud concoloribus. Pro 
notum ex majore parte nigro-fuscum, pallido-notatum. Antennae articulis basalibus 
exceptis infuscatae. Femora omnia cum tibiis fortiter nigro-notata. Cerci fuscescentes. 

Long, corporis ^ 6, $ 6 mm.; pronoti % I'S mm.; tegminum ^ 4, ? 4 mm. — Lat. 
tegminum t 2"5 mm. — -Long, femorum post. ^ 4-2, $ 5 mm. 

A very small species, bearing a great superficial resemblance to P. pacificum Scud., 
but not really allied at all to that (so far as the Hawaiian species are concerned) 
isolated species. The variegated pronotum, short cerci &c., will distinguish it at a 
glance, while the different neuration of the lateral field of the tegmina, and the 
different form of ovipositor, show that it is really more nearly allied to any of the 
preceding species, than to P. pacificum, which it superficially so greatly resembles. 

Hab. a single pair taken in the Waianae mountains, Oahu, at an elevation of 
3000 ft. It is an arboreal species. 



2 2 FAUNA HAWAIIENSIS 



(i6) P aratrigonidium pacijicum Scudd. 

Trigonidimn pacificum Scudder P. Boston Soc. xii. 1868, p. 139. 

Parat7'igonidumi pacific7im Brunner P. Z. S. Lond. 1895, p. 895. 

This species may be known from any of the preceding by the different neuration 
of the lateral field of the tegmina. (Plate II. fig. 6ff.) The pronotum is never 
variegate, nor have the posterior femora towards the base outwardly a large number 
of closely-set small, but separate, spots, as most of the species of the genus. The 
cerci are unusually long and in the % extend far back behind the ovipositor. The 
^ genital armature (often withdrawn into the body) is quite different to that of any 
of the preceding, in all of which it terminates in two elongate free processes, which 
bear minute denticles. In the present species there are no such processes. The 
ovipositor is also of different form, being interrupted at about the middle of the 
length of its superior margin. In all the other species, the interruption is nearer 
the apex than the middle. The ^ tegmina (which vary greatly in colour) are never 
maculate. In its habits this species is largely terrestrial, although it often ascends 
certain plants, especially frequenting the tree ferns, and the stems and dead leaves of 
the banana. In colour it varies from very dark brown (nearly black) to pale testaceous, 
and it also varies considerably in size. In the wet woods of Hawaii there would 
appear to be two races, a larger and generally paler one, and a smaller and very 
dark one with brightly shining thorax, the latter being the more strictly terrestrial 
in habits. When, however, examples from all the islands are compared, it seems 
impossible to keep the two as distinct, without making a number of other and 
ill-defined species. 

Hab. Found on all the islands in the mountain forests, preferring damp and 
shady places, and moving freely by day. 

Obs. There are several examples of this genus, which probably belong to other 
and distinct species, but they are too closely allied to some of those described above 
to be separated without the inspection of more material. 



Group PROGNATHOGRYLLIDES, new group. 

In the Proceedings of the Zoological Society (Dec. 1895) Herr Brunner v. 
Wattenwyl described a genus ProgiiatJiogryllus to include two species of Hawaiian 
crickets. The discovery of 14 additional species allied to these, has made it necessary 
to form several new genera, and to separate the two original species. The characters 
of the genus Prognathogryllus have in consequence required some modification. The 



ORTHOPTERA 23 

five genera characterized below are well marked, and their peculiarities seem to 
warrant the establishment of a special group, Prognathogryllides. They are dis- 
tinguished from the Podoscirtes group by the form of the head, which is strongly 
porrect ; and by the armature of the apex of the posterior tibiae, there being only 
two calcaria on either side of each. The number given by Herr Brunner is five, 
three outer and two inner, but the examination of many more species and examples 
than were submitted to him, has led to the conclusion that the upper of what he 
considered to be the three outer calcaria should rather be counted as the apical spine 
of its series. In the first place its position and appearance are rather those of a 
spine than of a true calcar, and secondly in some of the species, in certain examples 
it may be altogether absent, whilst in others of the same species it is present. If 
this spine be admitted as a calcar, the apical spine of the inner series must in many 
of the species be also considered as such, since the appearance of both, and their 
position as regards the calcaria, are precisely similar. This apical spine of the inner 
series is also sometimes wanting in some examples of a given species, in which case 
there are no spines adjoining the calcaria, and these stand out distinctly as two on 
each side. (Cf. Plate II. figs. \ob, xor, \od\ 15, 15^, \^b; etc.) \\i Prognathogryllns 
as now restricted, the apical spine of the inner series is very distinct from the calcaria, 
and this fact renders it probable that in the allied genera the true inner calcaria are 
also two, that which might at first sight be considered a third being merely the apical 
spine of the inner series. 

All the species are of elongate and slender form, except Prognathogryllus robustns, 
which is comparatively short and robust. In Thaitmatogryllus and Leptogryllus the 
tegmina are very small and scale-like, and sometimes only visible at the sides of 
the body at the hind margin of the pronotum, and the wings are wanting. ApJiono- 
gryllus has no free tegmina. In the ? of Prognathogryllus (the I being unknown) 
they are about equal in length to the head and pronotum together, but in P. robustus 
they extend nearly to the apex of the abdomen. The wings in this genus are ill- 
developed, being about as long as the short tegmina in P. alatus, and much shorter 
than the tegmina in P. robustus, but they are always present. 

In Nesogryllus $ (the ? being unknown), the tegmina are well developed for 
stridulation, but they do not extend back as far as the apex of the long and slender 
abdomen, and the wings are rudimentary. 

Prognathogryllus and Nesogryllus have a distinct tympanum on the inner face 
of the front tibiae. The other genera have none. 

The males of those species of Leptogryllus, which have the metanotum exposed, 
bear on this part two punctures, one on either side of the middle line, which are 
represented by two smooth points in the females. When the scale-like tegmina are 
larger, and conceal (at least for the most part) the metanotum, its basal part is 
depressed, and the punctures or orifices, which lie within, this cavity, are furnished 



24 FAUNA HAWAIIENSIS 

outwardly with a small curved process. In Thawnatogrylliis this cavity is very deep 
and abrupt and apparently extends beneath the pronotum, which with the tegmina 
entirely conceals it. Instead of the two minute orifices there appears to be a large 
transverse slit, with a small hard spine at either extremity. As in Leptogrylhis these 
structures are peculiar to the ^. 

All the species of this group are entirely nocturnal in their activities. In the 
day-time they conceal themselves beneath the bark of trees, in hollow stems, or 
amongst dead leaves attached to trees, but we have never met with any species that 
is not arboreal in its habits. All the species of Prognathogryllus, Nesogryllus and 
Aphonogryllus are extremely rare insects, while the Thaiimatogryllus and several of 
the species of Leptogrylhis have been taken in some numbers, but even these are 
rarely or never to be found unless a special search be made for them. 

The genera of Prognathogryllides may be tabulated as follows : 

1. {2) Cerci tribus articulis tarsorum posticorum conjunctis multo longiores Prognathogryllus. 

2. (/) Cerci tribus articulis tarsorum posticorum haud longiores. 

3. {&) Spinae tibiarum posticarum internae externis evidenter longiores. (Tegmina 

aut longa aut nulla.) 

4. (j) $ tegmina nulla Aphonogryllus. 

5. {4) i tegmina longa .., Nesogryllus. 

6. (j) Spinae tibiarum posticarum internae externis haud evidenter longiores. 

(Tegmina squamiformia, saepe minutissima.) 

7. (t?) Metatarsus anterior longissimus : calcar tibiarum posticarum externum et 

superius longum Thautnatogryllus. 

8. (7) Metatarsus anterior minus longus, tantum bis terve longior quam latior : 

calcar tibiarum posticarum externum et superius breve Leptogrylhis. 



Prognathogryllus (sensu strictiori). 

Prognathogryllus Brunner P. Z. S. 1895, p. 896. 

Tegmina abbreviata, vel fere ad apicem abdominis extensa. Alae sat distinctae, 
ad apices tegminum extensae, vel, si haec longa, his multo breviores. Tibiae anticae 
tympano instructae ; tibiae posticae spinis utrinque armatae, internis, quam externae, 
multo longioribus. Cerci ovipositori subaequales. %. 

(i) Prognathogryllus alattts Brunn. 

Prognathogryllus alatus Brunner P. Z.S. (1895) p. 896. 
(Plate II. figs. 8, 8«, & 8(5.) 



ORTHOPTERA 



25 



(2) Prognathogryllus elotigattis, sp. no\'. 

Gracilis, elongatus, colore testaceo. Tegrnina pallida, fere ad apicem segmenti 
abdominis secundi se extendentia, venis flavis. Femora postica gracillima, spinis internis 
externis evidenter longioribus. $. 

Long, corporis 22 mm., pronoti 4, tegminum 5-5, femorum post. 11, ovipositoris 
7 mm. (Plate I. fig. 15.) 

A very distinct species, which, although of nearly the same length as the preceding, 
appears much longer, owing to its much narrower form. The posterior femora are very 
slender and much less wide at the base, and the inner series of spines on the posterior 
tibiae are much less strong. 

Hab. The high plateau of Kauai. July, 1896. 



(3) Prognathogryllus robustus, sp. nov. 

Brunneo-niger, robustus. Tegmina fere ad apicem abdominis se extendentia. 
Tibiae posticae cum femoribus minus elongatae, spinis internis quam externis multo 
longioribus. %. 

Long, corporis 16 mm., pronoti 4, tegminum 9, femorum post. 9, ovipositoris 
10 mm. (Plate L fig. 14.) 

Quite unlike the other species in general appearance, of a dark brownish colour, 
and with well developed tegmina. The wings are much shorter than these. The 
posterior femora are formed much like those of P. alatus. The inner series of spines 
on the posterior tibiae is much more strongly developed than the outer, but they are 
shorter than those of P. alatus. 

Hab. High central plateau of the Kauai mountains. Very rare, a single % only 
taken, high up in a dead tree amongst the dry leaves that still adhered. 



(4) Prognathogryllus oahuensis, sp. nov. 

Haud robustus, flavo-testaceus, pronoto fusco-variegato. P. elongato simillimus, 
femoribus tibiisque posticis multo brevioribus facile distinguendus. %. 

Long, corporis 16 mm., pronoti 4 mm., tegminum 5 mm., femorum post. 9 mm., 
ovipositoris 6 mm. 

Easily known from P. elongattis, which (of the preceding species) it most resembles, 
by the shape of the posterior femora, which are much wider near the base in proportion 
to their length. 

F. H. II. 4 



26 FAUNA HAWAIIENSIS 

The inner series of spines on the posterior tibiae is more strongly developed, and 
the whole insect is less slender and elongate. 

Hab. Waianae Mountains, Oahu. A single % taken under bark in Feb. 1896. 
It was being attacked by the introduced ant, Pheidole viegacepJiala, which had already 
bitten off the greater part of the antennae and tarsi. 

(5) Prognathogryllus inexspectattis, sp. nov. 

Gracilis, elongatus, testaceo colore. Pronotum totum pallidum, haud fusco-varie- 
gatum. Tegmina abbreviata, venis flavis. Femora postica minus incrassata. %. 

Long, corporis 19 mm., pronoti 375 mm., Lat. pronoti 3 mm., Long, tegminum 
5 mm., femorum post, circa 9 mm., ovipositoris 6'50 mm. 

Readily distinguished from P. oalmensis by the much -narrower posterior femora, 
the unicolorous pronotum, which is also narrower, and the shorter spines of the pos- 
terior tibiae inwardly. From P. elongatus the evidently shorter posterior legs at once 
separate it. 

Hab. Kauai (4000 ft.). Under ha.rk of A cada koa. 

Aphonogryllus, gen. nov. 

Gracilis, elongatus, tegminibus alisque carens. Tibiae anticae tympano nullo 
instructae ; posticae utrinque spinis armatae, internis, quam externae, multo longioribus; 
calcar internum et superius multo longius quam dimidium metatarsi. Cerci brevissimi, 
tarsorum articulis tribus posticorum, una conjunctis, haud aequales. $. 

(i) Aphonogryllus apteryx, sp. nov. 

Testaceus, dorso plus minusve fusco-variegato. Tegmina libera nulla, sed latera 
mesonoti lobate-producta. Femora postica minus fortiter elongata. Tibiae posticae 
supra infuscatae, spinis utrinque armatae, internis multo longioribus, basi nigris, media 
parte pallidis. ^. 

Long, corporis 16 mm., pronoti 3'25 mm., femorum post. 7*5 mm., cercorum 2 mm. 
(Plate IL figs. 9, 9^;, 9^, gr.) 

Hab. Mountains of Oahu. Two males taken in 1896, one at an elevation of 
2500, the other of 3000 feet. 

Nesogryllus, gen. nov. 

Gracilis, elongatus. Tegmina capite una cum pronoto bis longiora. Alae teg- 
minibus multo breviores. Tibiae anticae tympano instructae ; posticae utrinque spinis 
armatae, internis, quam externae, multo longioribus ; calcar internum et superius haud 



ORTHOPTERA 27 

minus longum quam dimidium metatarsi. Cerci breves, metatarsis posticis subaequales. 
Venae tegminis superioris obliquae 2, necnon venae 3 breves ab angulo venae stridu- 
lantis externo emissae. Speculum vena transversa una in partes subaequales divisum. 
Campus apicalis brevissimus. ^. 



(i) Nesogrylhis stridulans, sp. nov. 

Gracilis, flavo-testaceus, tegminibus fere hyalinis, parte basali infuscata et opaca, 
venis pallidis. Antennae longissimae, testaceae, concolores. Pronotum fusconotatum. 
Femora omnia cum tibiis tarsisque pallide testacea et translucida. Tibiae posticae spinis 
pallidis utrinque armatae. ^. 

Long, corporis 22 mm., pronoti 4 mm., tegminum 13 mm., femorum post. 
io"5 mm., cercorum 375 mm. (Plate II. figs. 11 & \\a.) 

Hab. Mountains of Oahu, 3000 ft. A single ^ taken (in June 1895) from the 
interior of a hollow twig. 



Thaumatogryllus, gen. nov. 

Elongatus cylindricus. Tegmina minima, maris metanotum fere totum, feminae 
dimidiam fere partem metanoti celantia. Tibiae anticae tympano nullo instructae. 
Metatarsi longissimi ; tibiae posticae spinis brevibus utrinque subaequalibus armatae ; 
calcaria superiora, et e.xterna et interna, longissima et subaequalia. Cerci breves, parte 
tertia ovipositoris vix longiores. t ?• 



(i) Thauinatogrylhis variegatus, sp. nov. 

Flavo-testaceus, colore fusco nigroque variegatus. Femora omnia cum tibiis inter- 
mediis et anticis fortiter nigronotata. Tarsi pallidi. Tegmina brevia, metanotum totum 
haud tegentia. Z %■ 

Long, corporis 15 mm., pronoti 4 mm., femorum post. 10 mm., ovipositoris 
875 mm. (Plate I. fig. 16; and Plate II. figs. 10, \oa, \ob, \oc, \od.) 

Some individuals are considerably darker than others and there is some variation 
in size. The anterior legs are remarkably long and slender, their metatarsus being 
many times as long as wide. The posterior femora are somewhat more strongly and 
suddenly dilated on their basal part than is usual in the allied forms. The posterior 
tibiae are very long, and the spines on each side very short. 

Hab. Mountains of Kauai (4000 ft.). 

4—2 



28 FAUNA HAWAIIENSIS 



T,Fi'Tot;KVi ii's. orn. nov. 

Gracilis, sulxlrprossus. roj^iniiia niiniina siiuamilonnia. iioniumquain im'taiiiHiiin 
totuin, vol tViT totuiii, ti-m'iuia, saepc taiUuni ad latera extroina mcsonoti antice vix 
disccrnciula. Alao nullac. Tibiae anticac tympano nullo instnictae ; metatarsus tantum 
bis tcrve longior qiiani latior. ribiae posticae spinis brex'ibus utrinque subaequalibus 
armatae ; calcar internum et superius, i^uani caetera. longius, parti dimidiae metatarsi 
liaud aequale. Cerci brevissimi. jT if. 



^i) Lcptog^ryllus uigroliucatiis, sp. nov. 

Pallide tlavo-testaceus. Femora cum libiis immaculata. Abdominis latera late nioru. 
Femora postica perlonga, minus dilatata ; tibiar utrinque spinis brevibus armatae. 
Tegmina metanotum totum hand tegentia. / ^. 

Long, corporis iS mm., pronoti 4"5 mm., tcmorum post. ^ \i mm., ^ ii mm., 
ovipositoris 7 mm., cercorum 2 "5 mm. 

H.A.B. Mountains of Oahu and Maui. 



{2) Lcpfoy;ry//us niiiiviiiacu/att(s, sp. now 

Forma et colore fere praecedentis ; femoribus posticis et tibiis evidenter brevioribus 
facile distinguendus. ^f ^f . 

Long, corporis (baud contracti) 15"5 mm., pronoti 35 mm., femorum post. 9 mm., 
ovipositoris mm. (Plate L tig. 17 : and Plate IL tigs. 12 Cs: 12a.) 

This species is very like the preceding, but the shorter posterior legs will distinguish 
it at once. Moreover the abdomen, instead of the continuous broad lateral stripes, has 
a series of diamond-shaped spots on each side. The tegmina as in the preceding. 

H.\B. Kauai. In the Mountains 3000 — 4000 ft. 



(^3) Lcpii\^ry/liis sif/ii/ts, sp. no\-. 

Praecedentibus colore simillimus : pallide testaceus ; abdomen utrinque late nigro- 
lineatum; femora antica et intermedia cum tibiis immaculata: metatarsi postici apex 
niger. Tegmina minutissima, tantum ad latera corporis videnda. r. 

Long, corporis 12*5 mm., pronoti 3 mm., femorum post. 725 mm. (Plate II. 

fig- 13-) 

Hab. Mountain above Hilo, Hawaii, 2000 ft. 



ORTHOPTERA 29 

(4) Leptogryllus elongatus, sp. nov. 

Praecedentibus forma et colore simillimus ; femora antica et intermedia cum tibiis 
plus minusve distincte nigro- vel fusco-notata ; pronotum utrinque saepius indistincte 
fuscum ; abdomen utrinque fusco- vel nigro-lineatum, aut bilineatum. Tegmina (^ %) 
minutissima. Z ?. 

Long, corporis (haud contracti) 14-5 mm., pronoti 3'5 mm., femorum post. 9 mm., 
ovipositoris 6 mm. (Plate I. fig. 18.) 

Hab. Hilo, Puna, and Kau, Hawaii (2cxx) — 4000 ft.). 

(5J Leptogryllus kauaiensis, sp. nov. 

Forma fere praecedentium, colore variabili, testaceus, plerumque parte corporis 
majore nigro- et fusco-variegata ; femora antica et intermedia cum tibiis nigro-annulata. 
Tegmina metanotum totum tegentia (^), vel multo minora, unum ab altero longe dis- 
tantia, et brevissima ($). ^ $. 

Long, corporis 16 mm., pronoti 4 mm., femorum post. 8*5 — 9 mm., ovipositoris 
6 mm. 

Hab. Mountains of Kauai (2000 — 4000 ft.). 



(6j Leptogryllus fusconotatus, sp. nov. 

Praecedentis colore et facie ; J, tegminibus minutissimis bene distinctus. t- 
Long, corporis 14 mm., pronoti 375 mm., femorum post. 9 mm. 

Hab. Mountains of Oahu (2000 ft.). One J, taken in 1896. 

(7) Leptogryllus forficularis Brunn. 

Prognatlwgryllus forficularis Brunner P. Z. S. 1895, P- 897. 

(Plate H. figs. 14, 14a, & 141^.) 

In the % of this species the tegmina are extremely small and visible only at the 
extreme sides beyond the margin of the pronotum. It closely resembles the preceding, 
but its shorter and less wide posterior femora easily distinguish it. The length of these 
is only about 6^ mm. I have taken a single I which appears to belong to this species, 
although taken on a different island. Apart from the sexual characters it does not seem 
to differ at all from the %. 

Had. Kona, Hawaii, 3000 ft. lao Valley, Maui, i Z- 



FAUNA HAWAIIENSIS 



(8) Leptogryllns simillinms, sp. nov. 

L. forfindari simillimus, tibiis posticis minus spinosis distinguendus. ^ %. 

Very closely allied to L. forficularis, but generally if not always with the body 
darker in colour. The spines on the tibiae are evidently less close and do not extend 
nearly so far back towards their base. I have examined a series of specimens. The ^ 
is often slightly shorter and less robust than the $. 

Long, corporis (haud contract!) % 14 mm.; pronoti 3'25 mm., femorum post. 
6-5 mm., ovipositoris 5 mm. (Plate I. fig. 19 ; Plate II. figs. 15, 15a, & \'^b) 

Hab. Haleakala, Maui (4000 — ^5000 ft.). 



The following works are all that contain matter relating to the indigenous Hawaiian Orthoptera. 

1. ScuDDER, S. H. A century of Orthoptera. Decade I. Gryllides. P. Boston Soc. xn. 1868, 

pp. 139—143. 
Trigonidium pacifiaiin, n. sp., p. 139. 

2. BoRMANS, A. DE. Fauna orthopterologique des iles Hawai ou Sandwich. Ann. Mus. Genova, 

xvin. 1882 and 1883, pp. 438 — 448. 
17 species, 2 new. 

3. Redtenbacher, J. Monographic der Conocephaliden. Verh. Ges. Wien, xli. 1891, pp. 

315—562. 
Description of Brachymetopa, p. 430, and of B. discolor, blackbiirni, p. 431. 

4. Brunner von Wattenwyl, C. On the Orthoptera of the Sandwich Islands. P. Zool. Soc. 

London, 1895, pp. 891 — 897. 

29 species, 6 new. This paper deals with the collections received by the Committee 
previous to 1895. 



ZOOLOGrlCAL RESULTS based on material from New Britain, New Guinea, Loyalty 

Islands and elsewhere, collected during the years 1895, 1896, and 1897, by 

Arthur Willey, D.Sc. Lond., Hon. M.A. Cantab., Late Balfour Student of the 

University of Cambridge. Demy 4to. Parts 1, II, and III. Price i2i-. 6d. each. 

PART I. 

1. The anatomy and development of Peripatus novae-britanniae. By ARTHUR WiLLEY, M.A., D.Sc. 

With Plates I. — IV. and 7 figures in the text. 

2. Metaprotella sandalensis, n. sp. [Caprellidae]. By Dr PAUL MAYER. 

With 6 figures in the text. 

3. On a little-known sea-snake from the South Pacific. By G. A. BOULENGER, F.R.S. 

With Plate V. 

4. Report on the Centipedes and Millipedes. By R. I. POCOCK. 

With Plate VI. 

5. Account of the Phasmidae with notes on the eggs. By D. SHARP, M.A., F.R.S. 

With Plates VII.— IX. 

6. Scorpions, Pedipalpi and Spiders. By R. I. PocoCK. 

With Plates X.— XI. 

PART II. 

7. Report on the specimens of the genus Millepora. By Sydney J. HiCKSON, M.A., D.Sc, F.R.S. 

With Plates XII.— XVI. 

8. Report on the Echinoderms (other than Holothurians). By F. JEFFREY Bell, M.A. 

With figures on Plate XVII. and one figure in the text. 

9. Holothurians. By F. P. BEDFORD, B.A. 

With figures on Plate XVII. 

10. Report on the Sipunculoidea. By ARTHUR E. Shipley, M.A. 

With Plate XVIII. 

11. On the Solitary Corals. By J. Stanley Gardiner, M.A. 

With figures on Plates XIX. and XX. 

12. On the postembryonic development of Cycloseris. By J. STANLEY GARDINER, M.A. 

With figures on Plates XIX. and XX. 

13. On a collection of Earthworms. By FRANK E. Beddard, M.A., F.R.S. 

With Plate XXI. 

14. The Gorgonacea. By ISA L. HiLES, B.Sc. 

With Plates XXII. and XXIII. 

PART IIL 

15. Orthogenetic variation in the shells of Chelonia. By Hans Gadow, M.A., Ph.D., F.R.S. 

With Plates XXIV. — XXV. and one text-figure. 

16. Enteropneusta from the South Pacific, with notes on the West Indian Species. By ARTHUR 

WiLLEY, D.Sc, Hon. M.A. Cantab. 

With Plates XXVI.— XXXII. and 7 text-figures. 

17. On a collection of Echiurids from the Loyalty Islands, New Britain and China Straits, with an attempt 

to revise the group and to determine its geographical range. By ARTHUR E. SHIPLEY, M.A. 
With Plate XXXIII. 

PART IV. {In the Press:\ 

18. On the anatomy of a supposed new species of Coenopsammia from Lifu. By J. STANLEY 

Gardinep;, M.A., Fellow of Gonville and Caius College, Cambridge. With one plate. 

19. On the Insects from New Britain. By D. Sharp, M.A., M.B., F.R.S. With one plate. 

20. Report on the Stomatopoda and Macrura. By L. A. BORRADAILE, M.A., Lecturer of Selwyn 

College, Cai|hbridge. With four plates. 

21. Report on the /Slugs. By Walter E. Collinge, F.Z.S., Mason College, Birmingham. 

With two plates. 

22. Report on thefPolyzoa. By E. G. Philipps, Newnham College, Cambridge. With two plates. 

23. The Hydroid Zoophytes. By LAURA ROSCOE Thornely, University College, Liverpool. 

With one/ plate. 

24. Asti'osdera ivUleyana, the Type of a new Family of Calcareous Sponges. By J. J. LISTER, M.A., 

St John's College, Cambridge. With two plates. 

25. A contribution towards our knowledge of the Pterylography of the Megapodii. By W. P. 

Pycraft, A.L.S., British Museum. With one plate. 

26. The Stolonifera and Alcyonacea. By SYDNEY J. HiCKSON, M.A., D.Sc., F.R.S., Beyer Professor 

of Zoology in the Owens College, and ISA L. HiLES, B.Sc, Owens College, Manchester 
With wo plates. 

27. Report on /the Xeniidae. By J. H. AsHWORTH, B.Sc., Demonstrator in Zoology, Owens College, 
Manchjbster. With two plates. 

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FAUNA HAWAIIENSIS 



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NEUROPTERA 



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Earthworms. 

Formicidae. 

Hymenoptera Parasitica. 

Hemiptera. 

Thysanura and Collembola. 

Macrolepidoptera. 

[Hymenoptera Aculeata, Orthoptera, 
\ Neuroptera, and part of Coleoptera. 

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Part of Coleoptera. 

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FAUNA HAWAIIENSIS 



OR THE 



ZOOLOGY OF THE SANDWICH (HAWAUAN) ISLES 

Being Results of the Explorations instituted by the Joint Committee 

appointed by 

THE ROYAL SOCIETY OF LONDON FOR PROMOTING NATURAL KNOWLEDGE 

AND THE BRITISH ASSOCIATION FOR THE ADVANCEMENT OF SCIENCE 

And carried on with the assistance of those Bodies and of the Trustees of 

THE BERNICE PAUAHI BISHOP MUSEUM AT HONOLULU. 



EDITED BY 

DAVID SHARP, M.B., M.A., F.R.S. 

SECRETARY OF THE COMMITTEE. 



VOLUME II. PART II. 

NEUROPTERA 
By R. C. L. PERKINS, B.A. 

Pages 31 — 89; Plates III, IV, V, nncolonrcd. 



September 25, 1899. 



CAMBRIDGE : 

AT THE UNIVERSITY PRESS. 

1899 

\_All Rights reserved.] 



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ILsiDjis: F. A. BROCKHAUS. 
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JSombao: E. SEYMOUR HALE. 



NEUROPTERA 

By R. C L. PERKINS, B.A. 



NEUROPTERA. 
By R. C. L. Perkins. 






§ 1. General considerations on the Neuroptera. 

The Neuroptera are rather numerously represented in the islands, but the fauna 
as represented by this group is of the same fragmentary character, as is seen in the 
other Orders of insects, some of the extensive divisions of Neuroptera being totally 
unrepresented. 

At present the total number of species known to us amounts to iii, of which 54 
or nearly one-half belong to the Hemerobiidae (including herein Chrysopides and 
Myrmeleonides), 29 to the Odonata or dragon-flies, and 25 to the Psocidae. The 
remaining three species belong to two other divisions and consist of two Termitidae, 
and a solitary species of Embiidae. 

The divisions Ephemeridae and Trichoptera (or Phryganeidae), and the smaller 
groups Perlidae, Sialidae and Panorpidae are absolutely unrepresented. The entire 
absence of the first four of these divisions is a very remarkable fact, as they are aquatic 
in habits, and the numerous mountain streams and rivers would appear to be admirably 
adapted for many species ; so that one must conclude that these water-frequenting 
insects are ill-adapted for passing over wide expanses of ocean. 

The Hemerobiidae, excluding the Myrmeleonides, have 52 species which belong to 
6 genera. Not one of these species is known to occur in other countries, but two of 
the genera, which are represented by single species, have no alliance with the other 
forms, and when this family has been extensively collected in other countries, these two 
species will probably be found elsewhere. This is almost certain to be the case with 
the small Chrysopa tnicrophya, a species which is abundant in gardens in towns and 
settlements, although it has extended its range to a considerable elevation up the 
mountain sides. A single species, referred temporarily to Megalomus (although not a 
true member of that genus), is more remarkable ; it may even prove to be peculiar to 
the islands, but will more probably ultimately be found elsewhere. It is noteworthy 
that both these insects are of general distribution over the islands, in contrast with 
most other species of this family, which are so often confined to a single island. 

Nineteen species are included in the genus Nesomicromus, which has been founded 
F. H. II. 5 



32 FAUNA HAWAIIENSIS 

for their reception. The simpler forms of this genus are very similar to the well-known 
genus Micromus, but the more remarkable species have the wing-contour of Drepa- 
nopteryx, and bear to Micromus a relation very similar to that which Drepanopteryx 
bears to Megalonius or Hemerobius. Between the extreme forms of Nesomicromus 
species are found in quite intermediate conditions, so that one is compelled to treat the 
whole series of species as forming but a single genus, and their relationship is strongly 
evidenced by the apical abdominal appendices of the tt, which though differing in 
detail in many species, nevertheless possess notable peculiarities common to all. 

Allied to Nesomicromus, and no doubt evolved within the islands from some such 
form, are the other two genera, Pseudopsectra and Nesothauma, each with but one 
species at present known. The former is in most respects intermediate between the 
latter and Nesomicromus. Both these insects, but especially the Nesothauma, have 
claim to be considered amongst the most remarkable of all known species oi Neuroptera. 
Nesothauma has no trace of posterior wings and the front pair are almost of the 
consistency of the elytra of a Coleopterous insect. Pseudopsectra is likewise incapable 
of flight, but the front pair of wings are less abnormal, and the posterior pair are 
represented by small lobes. Although reminding one of the anomalous and rare genus 
Psectra, Psetidopsectra is, without doubt, not related in any way to that genus. The $ 
characters of both Pseudopsectra and Nesothatima are formed entirely on a similar plan 
to those of Nesomicromus. The two insects included in these abnormal genera are 
very local and rare, and both frequent the same locality, Haleakala on Maui, where 
they are found at an elevation of five or six thousand feet above sea-level. 

As above mentioned the single species of Chrysopa is almost certainly foreign, but 
another genus, Anomalochrysa, not known from elsewhere, includes no less than 29 
species. These species form a most interesting series, but are excessively difficult to 
differentiate, owing to the great variability in colour of many of them, and the instability 
of the characters afforded by the nervuration. Here again the terminal segments of the 
^ afford great help in many instances, and on these characters there would appear to be 
two good genera, but the females of the two sections appear to present no points for 
generic division, at least in dried examples, the distortion of the body after death being 
much greater in this sex than in the ^. 

The species of Hemerobiidae have in general a much more restricted range than 
have the indigenous Agrionidae of the Odonata. Excluding the single Megalomus and 
Chrysopa as probably foreign, of the genus Nesomicromus 1 5 out of the 1 9 are restricted 
each one to a single island, the remaining four being widely distributed insects, Hawaii 
and Maui each having four species peculiar, while the latter likewise has also its 
peculiar genera Psetidopsectra and Nesothau7iia. The species of Anomalochrysa are 
even more localized, two only of the 29 occurring on more than one of the islands. In 
this genus the island of Hawaii is extraordinarily rich, since it has 12 species peculiar to 
itself, and both of the two more widely distributed species are also found there. The 



NEUROPTERA 33 

following table of the indigenous Hemerobiidae gives the percentage of species peculiar 
to each of the islands. 





Species peculiar to. 


Kauai 


7 


Oahu 


6' 


Molokai 


6 


Lanai 


— 


Maui 


9 


Hawaii 


16 



ber of species. 


Percentage of peculiar 




species. 


8 


87-5 


9 


60 


8 


75 


2 


— 


12 


75 


20 


80 



Of the two Myrmeleonides I have not met with the Eormicaleo perjurus Walk., 
which is almost certainly becoming extinct ; the other, Eormicaleo wilsoni, is locally 
common in open places on the lava-flows of Hawaii, where there is a stunted vegetation, 
and it also is found on other of the islands. Very probably it will prove to be a 
natural immigrant, and will ultimately be found elsewhere. 

If we compare this table with that of the Agrionidae given on p. 34 it will at once 
be seen that the percentage of species peculiar to the several islands is much more 
nearly equal in the Hemerobiidae than is the case with the dragon-flies. In both tables 
Kauai stands first in the proportion of its peculiar species to the total number found 
upon it, and this is no doubt due, at least to a considerable extent, to the greater 
distance between it and the neighbouring island of Oahu, and probably in a lesser 
degree to its position at the extremity of the forest-bearing islands. The extraordinary 
richness of Hawaii in species of Anomalochrysa may perhaps point to that island as the 
spot where these insects first became established. The small percentage of species of 
Agrion peculiar to most of the islands is probably partly due to their greater powers of 
flight as compared with the Hemerobiidae, the high percentage of species peculiar to 
the more remote island of Kauai rather pointing to this conclusion, but it may be that 
they have been established for a shorter time in the islands than the Hemerobiids, 
or, even if antecedent to these, they may be less susceptible to the effects of isolation 
and the changes in environment thereby produced. However that may be, there is no 
doubt that several of the species of Agrion are at the present time in the process 
of forming other distinct species, owing to the isolation and change of conditions 
brought about by individuals having spread to more than one of the islands. 

The Odonata or dragon-flies. The 29 species of Odonata are distributed in 5 genera, 
but no less than 24 are assigned to the genus Agrion. No doubt they will ultimately 
be separated from that genus, and will themselves form not less than three distinct 
genera, but at present it seems better to leave them under that name, as many of the 
species are difficult, being very variable even in important characters, and some have 
already formed more or less distinct local races, or subspecies, so that the question as 

' Anomalochrysa riifcscens M'^L., the locality of which is not recorded, is included here as being 
probably Oahuan. 

S— 2 



34 FAUNA HAWAIIENSIS 

to which are true species is as yet by no means absolutely settled. In the females the 
climax of difficulty is reached, for these have no structures suitable for distinguishing the 
species that can compare with the terminal appendages of the abdomen of the ^, while 
the superficial distinctions, such as colour, details of nervuration, etc. are notably variable. 
All the species of Agrioit are peculiar to the islands and are probably all related to one 
another, as more or less transitional forms are found between the different groups, and 
it is probable that all originated from some single species which reached the islands in 
remote times. Of the other four dragon-flies the Tramea, Pantala, and Anax jimius, 
are no doubt natural immigrants, being all species of powerful flight and all are distri- 
buted throughout the group, whereas a large proportion of the species of Agrion are 
either restricted in range to one island, or when occurring on more than one are tending 
to form other distinct species, or local forms. The larger Anax, which is a very fine 
insect, is so far as is known peculiar to the islands, and has probably been established there 
sufficiently long to acquire characters which separate it from any other of the genus. 
It is very powerful on the wing, and is widely distributed over the islands, and extends 
its range far up the mountain sides. Another moderate-sized species, assigned to 
Sympetrum by Karsch, but for which Kirby has established a genus, Nesogonia, is also, 
so far as is known, peculiar to the islands. It is generally distributed over the group, 
in the mountains, and is notably variable in size, colour and nervuration. 

If we limit our investigation to the species of Agrion it will be seen that in the 
number of species peculiar to it, Kauai ranks easily first of the six larger islands. 
Excluding A. xanthome las and A. pacificum, species ubiquitous over the islands, seven 
species are restricted to Kauai, nor has it any species that even extends to the neigh- 
bouring island of Oahu. The latter island has four species peculiar to itself, and five 
others which are found also on one or more of the others. Maui, Molokai and Hawaii 
have each but a single species peculiar to them, while the little island of Lanai with 
seven or eight species has none. It should be added however with regard to these 
islands that some of the species upon them form varieties very distinct from the typical 
examples, and this is especially the case with several of the species found on Hawaii, 
where diminution in size and corresponding changes in nervuration are often evident. 
Nevertheless the occurrence of examples in a condition intermediate between the typical 
and extreme forms of such species renders it inadvisable to consider them as belonging 
to more than one species. The following table shows the percentage of species of 
Agrion peculiar to each of the islands. 

Species peculiar to. Total number of species. Percentage. 

9 777 

lO 40 

8 12-5 

7 — 

10 10 

9 ii'i 



Kauai 


7 


Oahu 


4 


Molokai 


I 


Lanai 


— 


Maui 


I 


Hawaii 


I 



J 



NEUROPTERA 



35 



Psocidae are richly represented in the islands and twenty-five species are dealt with 
in the present paper. No doubt many others yet remain to be discovered, indeed other 
species are certainly included in those collected by me, but owing to their small size 
and poor state of preservation it was not advisable to attempt the description of the 
species. Nearly all are subject to much distortion and contraction of the body-segments 
after drying, as well as discoloration, and owing to the exudation of a sticky substance 
they are with difficulty relaxed, so as to be suitable for examination. Evidently there 
are good characters in the terminal abdominal segments of the ^, but these are not 
available in dried examples. Many of the species exhibit great variability (in colour, 
nervuration, etc.), which is so remarkable a feature of so many Hawaiian insects. The 
difficulties of study in this group are so great from the causes above enumerated, that 
the present paper on these insects can only be regarded as a preliminary sketch, especially 
as regards the species referred to the genus Elipsocus, in which the instability of 
nervuration is so great, as to render the discrimination of species almost hopeless, without 
special attention to the insects in the field. In this preliminary study I have not con- 
sidered it advisable to enter minutely into the generic question, but have referred all 
the species to three well-known genera. The solitary species referred to Stenopsocus 
temporarily, is clearly generically distinct therefrom, but only one example, in mutilated 
condition, was secured. It is very different to any other Hawaiian Psocid, and is inter- 
esting as having been taken at a high elevation in the mountains, where the nights, 
even in August, were cold, with hard frosts. Of the other 24 species, 14 are assigned 
to Psocus, most species of which appear to be confined, each one, to a single island, 
while 10 are placed in the genus Elipsocus. Some species of this latter genus are 
so variable in nervuration, that not only generic, but even characters of superior value, 
are affected. 

The other components of the Neuropterous fauna are comparatively of little interest. 
The two Termites belong to the genus Calotermes. The smaller of these, C. margini- 
pennis Latr.,i is certainly an introduced species and has done great damage to wooden 
buildings in the city of Honolulu. The larger one, referred to C. castaneus Burm., is 
very possibly distinct from that species, which was described from winged forms. The 
soldiers of the Hawaiian species possess well-developed eyes, and it is noteworthy that 
they are found (and probably only found) in the native forests, and series of them 
from three of the islands vary distinctly in the length of the gular area on the under-side 
of the head, as though they were already forming distinct races on the various islands. 
It is possible however that this variation, although affecting an important character, is 
merely such as occurs in different communities, and is not due to isolation on different 
islands, the material at hand not being sufficient for deciding this point. 

The solitary Embiid {Oligotoma insularis M'Lach.) is an interesting insect on 
account of the dorsal sclerites of the thorax in the winged Z not being of the simple 
structure usual in these insects, and very different to those of the apterous ?. An 



o 



6 FAUNA HAWAIIENSIS 



account of its habits and development is published in the Entomological Monthly 
Magazine, xxxiii. (1897), p. 56. 



§ 2. Systematic account of the Neuroptera. 

HEMEROBIIDAE. 

HEMEROBIIDES. 

The Hemerobiidae represented by the genera Megalomus, Nesomicromus, Psetido- 
psectra, and Nesothauma, as well as the numerous species of Anojnalochrysa and the 
solitary' one oiChrysopa of the subdivision Chrysopides and including also Myrmeleonides 
form the most extensive section of the Hawaiian Neuroptera. Nearly all the species 
are confined to the mountain forests. The larvae of the Hemerobiids proper appear to 
feed on the species of Psocidae, those of Anomalochrysa, I have little doubt, prey on 
Lepidopterous larvae. Their mandibles are sharp and strong, so that they can give 
quite a sharp bite, when they fall upon the face or neck, as is often the case when one 
is beating the branches of trees. All the species appear to be nocturnal, although some 
are very readily disturbed, as one walks through the brush in the daytime. 



Megalomus Ramb. 

(i) Megalomus hospes, sp. nov. 

Head, thorax, and abdomen varying in colour from nearly uniform dark brown or 
blackish, with obscure pale markings, to a nearly uniform yellowish colour ; generally 
brown with pale markings, or yellow with fuscous markings. 

Anterior wings grey, more or less fulvescent, with a more or less distinct dark spot 
on the cubitus posticus, towards the base. Nervuration set with hairs, and with alternate 
light and darker spaces. The gradate nervules form two more or less distinct, trans- 
verse, oblique darker lines, and there is a third near the base. In one example the 
whole middle portion of the wing between the inner and the basal series of gradate 
nervules is deeply infuscate, forming a wide blackish transverse fascia. Radius with 
three sectors. The posterior wings are almost hyaline, and beautifully iridescent, 
their nervuration pale. 

' We have excluded the Chrysopa oceanica Walk, from the list as probably not belonging to the Hawaiian 
fauna. Several species brought home by the Beechey expedition with the locality " Sandwich Is.'' appear to 
have come from the other islands of that name. 



NEUROPTERA 2,7 

$ appendices very long and narrow, the sides subparallel, with an apical depression 
outwardly, rounded at their extremities ; inwardly on their apical portion furnished with 
short somewhat spinose hairs, which are directed inwardly. Between the appendices 
there may be seen a short chitinous process, with the apex bifid, forming two spines. 
Ventral valve very narrow, sublinear, its sides parallel in dorsal or ventral aspect, curved 
upwards and clothed with long hairs. (Plate IV. figs, i, 2, 16 & i6«.) 

Expanse 13 — 17 mm. 

Hab. Found all over the group, in the mountains, but not very abundant. This 
species has no relation with the rest of the Hawaiian Hemerobiidae, and I suspect it 
has been introduced. 



Nesomicromus, gen. nov. 

Allied to Micromtis, some of the species having the superficial appearance, as well 
as the nervuration of that genus. The wings are either rounded at the apex, or falcate, 
somewhat resembling Drepanopteryx, the one form passing gradually into the other, 
so that the species cannot be subdivided on this character, although the extreme forms 
are vastly different. The species with simply rounded wings can (so far as I can see) 
only be separated from Micromus by the length of the joints of the maxillary palpi, in 
which they also agree with the species which have falcate wings. The penultimate joint 
of these palpi is very short, being only about half as long as the terminal. 

The c/ characters are very similar throughout the genus, the appendices being rarely 
very conspicuous, usually of triangular form and pointed at the apex. In all the species 
they give off each from their lower margin towards the base a fine spine, and slight 
differences in the length, form, etc. of these spines furnish useful specific characters. 
The general similarity in the form of the appendices, which extends also to the two 
following genera, is very remarkable. As in Micromus, there is no free cellule formed 
by a recurrent nervule at the base of the wing. 



(i) NesomicroTnus vagus, sp. nov. 

Brown or black ; face, legs and palpi paler. Antennae varying in colour from 
testaceous to nearly black. Thorax with short pale pubescence. 

Anterior wings brownish or grey-brown (grey in immature examples), generally 
with a small pale spot towards the base, situated on the cubitus posticus. Gradate 
nervules more or less infuscate, forming two transverse darker lines on the wings, often 
irregular and broken, sometimes wanting. Apical margin evidently, but slightly excised, 
giving the wings a hooked appearance at the tip. Nervuration dark, sometimes (with the 
pterostigma) more or less pink, the nervures with short and very inconspicuous hairs. 



38 FAUNA HAWAIIENSIS 

Radius normally with 5, rarely 4 or 6 sectors ; in the latter cases the number is often 
different on the two sides of the same insect. 

Posterior wings subinfuscate, but paler than the front wings and more transparent, 
nervuration generally dark, the apical margin very slightly excised below the tip of the 
wing. 

^ appendices viewed laterally, sub-triangular, narrowly rounded at the apex, on 
their inferior margin inwardly they are furnished each with a fine spine, curved upwards, 
and under a very strong lens finely serrate or dentate, the spines extending backwards 
rather beyond the apices of the processes. (Plate IV. fig. 3.) 

Expanse 12 — 18 mm. 

Hab. Common all over the group in mountain forests, and sometimes found on 
the coast. 

(2) Nesomicromus latipennis, sp. nov. 

Allied to the preceding, and of similar colour. 

Anterior wings brown, posterior pair infumate. Radius with 5 sectors. Readily 
known by the very broad and short wings, the apical margin not perceptibly emargi- 
nate ; the posterior pair very obtuse, their apical margin almost straight, not slightly 
concave. (Plate IV. fig. 4.) 

Expanse 15 mm. 

Hab. Kona, Hawaii (3500 ft.); i % June, 1892. 

(3) Nesomicrojnus angustipennis , sp. nov. 

Closely allied to N. hawaiiensis, but with the wings narrower and the apex 
somewhat more produced. The radius gives off 6 or 7 sectors. The cellules formed 
between the outer and inner series of the gradate nervules are extremely narrow in 
proportion to their length, and this fact gives a character to the wing by which the 
species may be easily recognized. 

The body and anterior wings are brown, but only gradually arrive at this colour, 
being pale for some time after the emergence of the insect. Normally there is a small 
pale spot towards the base of the wing on the cubitus posticus, and the radius bears 
alternate pale and dark spots as in N. hawaiiensis. The variation of the present species 
also appears analogous to that ot the other, the nervuration and pterostigmata being pink 
in some examples, and the wings in some are evidently less narrow than usual, etc. 

The Z characters differ, in that the spines of the apical processes are shorter, and 
do not reach to the apices of the processes themselves. (Plate IV. fig. 5.) 

Expanse 12 — 16 mm. 

Hab. Mountains of Kauai (4000 ft.). 



NEUROPTERA 39 

(4) N esomicrotnus drepanoides, sp. nov. 

Dark brown or nearly black, legs and antennae pale. Anterior wings dark brown 
more or less mottled with lighter yellowish-brown, or nearly entirely pale yellowish- 
brown with the gradate nervules delineated by transverse dark lines, the radius with 
the usual alternate dark spots, and some dark markings near the base of the wing. 
Posterior wings more or less infuscate. Pterostigma sometimes pink. 

Allied to the preceding species, but with the apical margin of the anterior wings 
more deeply excised, and the dorsal margin from the apex to about the middle, 
evidently, but very slightly, concave. The wings are also less narrow. The sectors of 
the radius are 6 or 7 in number, whereof the two which are nearest the base sometimes 
unite close to their point of origin. The general appearance of the nervuration is that 
of the preceding. In the $ the spines of the apical processes appear to extend about 
to the apex of each process, and they cross each other near the base. (Plate IV. 
fig. 6.) 

Expanse 14 — 16 -5 mm. 

Hab. Kauai (4000 ft), i ,?, 3 ? taken. Of the latter one example is much paler 
than the others (as described above), and it also has broader wings, but I doubt whether 
it is specifically distinct ; in fact no two individuals out of the four agree. 

(5) Nesomicronius paradoxus, sp. nov. 

Nearly black, the legs and antennae and some marks on the dorsum of the thorax 
pale. Anterior wings dark brown, the costal area hyaline for the most part, and some 
pale spots around the margins, especially along the apical. Posterior wings nearly 
entirely infuscate. 

The apical margins in both pairs of wings are excised, in the inferior, very lightly. 
The dorsal margins of the superior pair are also deeply excised to about the middle, 
forming there a conspicuous rounded lobe, after which they are slightly concave to the 
base. The nervuration is black and distinct, and hardly perceptibly furnished with 
hairs. There are 6 sectors to the radius. (Plate IV. fig. 7.) 

Expanse 12 mm. 

Hab. Kilauea, Hawaii (4000 ft.) ; i %. 

(6) Nesomicromus fulvescens, sp. nov. 

Dark brown with pale markings, antennae and legs pale. Anterior wings pale 
yellowish-brown, largely but not deeply infuscate from the region of the inner gradate 
series to the apical margin. The outer gradate nervules are blackish and infuscate, 

F. H. II. 6 



40 FAUNA HAWAIIENSIS 

forming a distinct transverse dark line, the dorsal margin near the base is also dark, as 
also spots on the costa, radius, and the cubital nervures. Posterior wings subhyaline, 
with rather pale nervuration, the outer series of gradate nervules darker. 

Anterior and posterior wings very broad in proportion to their length, their apical 
margins hardly perceptibly emarginate. In the former the radius gives off 5 sectors, 
the 5th twice furcate before reaching the outer series of gradate nervules. 

No doubt some examples of this species will have 6 sectors to the radius. 

Expanse i6'5 mm. 

Differs from N. d^^epanoides and angustipennis , much as N. latipennis does from 
N. hawaiiensis. 

Hab. Waianae Mts., Oahu (3000 ft.) ; i ?. 



(7) Nesomicromus belhilus, sp. nov. 

Black, or more or less brown, head and sometimes the prothorax with pale spots, 
legs pale, front and middle tibiae with two distinct black rings. Antennae with the two 
first joints black, the rest pale at their base, and dark at the apex. Anterior wings 
$ whitish, subtransparent, iridescent, with a large dark irregular and broken blotch at 
the base, and another more or less evident at the apex, bounded inwardly by the 
suffused nervules of the outer gradate series. All the nervuration with alternating dark 
and pale spaces, the dark ones on the radius very distinct. In the $ the anterior wings 
are much suffused with ochreous-brown, a round spot between the upper parts of the 
two gradate series being less suffused, but not very distinct. The posterior wings are 
transparent and iridescent, infumate at the extreme base along the dorsal margin, and 
their nervuration is to a large extent pale, becoming dark towards the apex. In the 
? these wings are more clouded, especially along the dorsal margins, and the nervuration 
is on the whole darker. 

The anterior wings are simply rounded at their apices, the apical margin not at all 
emarginate, but forming a continuous curve with the hind margin. There are 5 sectors 
to the radius, sometimes 6, at least on one side. 

Abdomen $ with the terminal appendices yellow ; their apices very narrow and 
somewhat produced, slightly turned upwards and inwards and subtuberculate. The 
spines are long and very fine, their extremities curved upwards, but they do not extend 
backwards so far as the apices of the appendices themselves. (Plate IV. fig. 8.) 

Expanse 16 — 17 mm. 

Hab. Haleakala, Maui (5000 ft.); very rare, 3 ^, i ? taken. 



NEUROPTERA 41 



(8) Nesomicromus molokaiensis, sp. nov. 

Female, very closely allied to the preceding, the anterior wings much more 
infuscate, dark brown in colour, with paler markings, the chief of which is a roundish 
spot near the apex, between the two gradate series. The posterior wings are entirely 
infumate, except for a pale apical spot on each, corresponding to those on the front 
wings, their nervuration is chiefly dark, and the nervures do not bear alternate light and 
dark spots, although some are entirely pale. 

In the anterior wings the radius has 6 sectors. 

Expanse i6'5 mm. 

Hab. Molokai, above 4000 ft.; i ? taken in June, 1893. It is possible that this 
form may prove to be a variety of the preceding species. 



{9) Nesomicromus 7ninor, sp. nov. 

Female closely allied to N. bellulus, but smaller, the wings much shorter, the 
anterior pair for the most part infumate, costal area and the upper part of the wing 
below and along the radius clearer. 

Five sectors to the radius, the sectors themselves, at least towards the base, with 
alternate dark and light spaces, as also the radius. Posterior wings entirely, but lightly 
infumate, except for an ill-defined pale spot near the apex. Nervuration for the most 
part dark, without distinct alternating light and dark spaces. 

Antennae pale, testaceous, the joints infuscate at their apices, the two basal joints 
brownish. 

Expanse 14 mm. 

Hab. Waianae mountains, Oahu (3000 ft.); i % taken in February, 1896. 



(10) Nesomicromus infumattis, sp. nov. 

Female closely allied to N. belluhis, anterior wings darkly infumate, with only 
sparse small pale spots, especially along the dorsal margin ; costal area paler than the 
rest of the wing ; at the base there are some dark markings, and the gradate nervules 
are distinct, as two zigzag transverse lines, darker than the ground colour. 

The wings are very narrow in proportion to their length. There are 5 or 6 sectors 
to the radius (5 one side and 6 the other). 

Posterior wings infumate, but transparent, no trace of a paler apical spot, the 
cubitus posticus very strong, black, except at the extreme base. 

6—2 



42 FAUNA HAWAIIENSIS 

Two basal joints of antennae dark, the other joints infuscate, sHghtly paler at their 
bases. 

Expanse i6 mm. 

Hab. Haleakala, Maui (5000 ft.); i % taken in Oct. 1896. 



(11) Nesomicromus longispinosus, sp. nov. 

Dark brown or blackish, antennae, including the two basal joints, testaceous, the 
apices of the joints lightly infuscate. Legs testaceous, front and intermediate tibiae 
largely fuscous. 

Anterior wings brownish-grey, nervuration for the most part dark, but interrupted 
by pale spots, radius very distinctly alternately light and dark. At the extreme base of 
the dorsal margin and about the cubitus towards its base, there is more or less dark 
infuscation, and one or both of the series of gradate nervules are more or less infuscate, 
and form dark lines. The wings are rounded at the apex, and the radius gives off 
6 sectors. 

Posterior wings hyaline, nervuration pale, but darker along the outer series of 
gradate nervules, so as to form an evident dark line. Radius connected with the sector 
by several transverse nervules. 

Z appendices pale, narrow, the spines unusually strongly developed, and very finely 
spinulose along one edge, very long, extending far behind the apices of the appendices, 
and crossing one another. (Plate IV. fig. 15.) 

Expanse of ^ 13 mm.; $ 15 mm. 

Hab. Kilauea, Hawaii (4000 ft.) ; i t and i % taken. Remarkable for the long 
spines of the Z appendices, and the additional transverse nervules in the posterior wings 
of both sexes. 



(12) Nesomicromus haleakalae, sp. nov. 

Female closely allied to the preceding, rather larger and with the anterior wings 
more grey, less tinged with brown. 

The black markings of the wings are more distinct, the dark markings along the 
cubitus forming with the blackish suffusion along the gradate nervules of the inner 
series a distinct curved blackish line. The radius gives off 5 sectors, and the elongate 
cellules formed between the two series of gradate nervules, are evidently less narrow and 
numerous than those of the preceding species. 

Posterior wings very much as in N. longispinosus, nearly hyaline, the radius and 
sector connected by several transverse nervules. 



I 



NEUROPTERA 43 

The antennae agree with those of the preceding species in having the two basal 
joints testaceous, but the rest are much darker, pale narrowly at the base. 
Expanse 16 mm. 
Hab. Haleakala, Maui (4000 ft.) ; i $ taken in April, 1894. 

(13) Nesomicromus brunnescens, sp. nov. 

Brown or blackish brown, legs and antennae testaceous, the joints of the latter 
infuscate on their apical portion. 

Anterior wings brown, or brownish grey (the latter colour probably only in 
examples not fully mature), rounded at their apices, radius giving off 7 — 9 sectors, the 
gradate nervules forming two obliquely transverse, fine dark lines, sometimes very in- 
distinct, the cellules between the two series very narrow in proportion to their length. 

Posterior wings subhyaline, somewhat iridescent, nervuration light brown or 
yellowish, the outer series of gradate nervules darker. 

$ appendices narrow towards the apex, not strongly produced, their spines some- 
what strong, curved upwards, serrulate, and extending back to the apices of the appen- 
dices or even slightly beyond them. (Plate IV. fig. 9.) 

Expanse 14 — 16 mm. 

Hab. Molokai, Lanai, and Haleakala, Maui. Rare (2000 — 5000 ft.). 

( 1 4) Nesomicromus rubrinervis, sp. nov. 

Head, thorax, legs and antennae testaceous, abdomen darker. 

Anterior wings narrow, their apices rounded, pale brown, tinged with pink, the 
nervuration pink. Radius with four sectors. Nervules of inner gradate series sub- 
infuscate, forming a faint dark line. In this series four of the transverse nervules are 
nearly continuous and form a slightly oblique line, but the two upper ones of the series 
are greatly separated from the four lower, and from one another. Posterior wings 
nearly hyaline, the nervuration and pterostigma pink. 

Spines of the appendices of ^ reaching about to their apex, and curved upwards. 

Expanse 13 — 14"5 mm. 

Var. a. Dark brown in colour, wings dark, shorter and wider than in the type, 
pterostigmata and nervuration pink. Radius with 5 sectors. 

The dark colour may really be normal, the two examples described above being 
possibly immature. The pinkish tinge to the wings and the condition of the inner series 
of gradate nervules is so similar, that I have little doubt that the two forms are one 
species, in spite of the additional sector to the radius. 

Expanse 14 mm. 

Hab. Kilauea, Hawaii (4000 ft.) ; i ^ 2 $. 



44 FAUNA HAWAIIENSIS 



(15) N esomicromiis forcipaius, sp. nov. 

Head, thorax and abdomen all pale, subtestaceous. Basal joint of the antennae 
suffused with pink. 

Anterior wings rounded at the apex, dull yellowish, with obscure fuscous spots. 
Nervuration pale, interrupted by spaces of a pink colour. Radius with 4 sectors. 
Posterior wings with pale nervuration, more or less pink in parts. 

Appendices of $ long and strong, of about equal width (in lateral view) from near 
the base to the apex, their extremities slightly turned inwards, the spines on their inferior 
margin near the base very short, not nearly extending back to their apices. (Plate IV. 
figs. 14 & i4«.) 

Expanse of ^ 1 1 mm. ($ unknown). 

Hab. Makaweli, Kauai (above 2000 ft.) ; i $ taken. The form of the appen- 
dices will distinguish it at a glance from any other species. 

(16) Nesoniicronius distinctus, sp. nov. 

Female, with the head and thorax yellowish-brown, metathorax and abdomen 
darker. Legs and antennae testaceous. 

Anterior wings rounded at their apices, pale yellowish-brown ; the outer series of 
gradate nervules black, and with blackish infuscation around them, forming a very 
distinct, and but slightly oblique, transverse dark line. Inner series hardly infuscate. 
Radius with distinct black spots at the points of origin of the sectors, which are five in 
number. Posterior wings pale, subhyaline, pterostigma and nervuration pink, nervures 
in the region of the outer gradate series deep black, forming a conspicuous curved 
marking at the apices of the wings. 

Expanse 14 mm. 

Hab. a single % taken in the mountains on Molokai, August, 1893. 

(17) Nesomicrontus subochraceus, sp. nov. 

Head and thorax yellowish or testaceous, abdomen generally darker, legs and 
antennae testaceous, the latter with darker annulation. 

Anterior wings rounded at the apex, pale greyish-fulvous, more or less mottled 
with fuscous. Nervures alternately dark and light; radius with 4 sectors, the number 
being constant in the series examined. Posterior wings hyaline and iridescent with 
very pale nervuration, the nervures near the apical margin becoming black and forming 
a distinct marking. 



NEUROPTERA 45 

^ appendices of the usual form, the spines hardly reaching to their apex. Apical 
ventral segment narrow, tongue-like. (Plate IV. fig. 10.) 

Expanse 13 — 16 mm. 

Hab. Molokai, Maui, and Hawaii (from 3000 — 5000 ft.). 



(18) Nesomicronms sienopteryx, sp. nov. 

Small, blackish, antennae and the posterior legs testaceous, anterior and middle 
tibiae and femora for the most part dark, their tarsi testaceous. 

Wings very narrow ; anterior pair rounded at the apex, greyish-fuscous, slightly 
fulvescent, with a black streak from the base, uniting with the dark inner series of 
gradate nervules, to form a curved dark line. Radius with 5 sectors, nervuration dark, 
interrupted by pale spaces. Posterior wings subhyaline, very lightly infumate, nervu- 
ration somewhat dark for the most part, the nervures not conspicuously blackened 
towards their apices ; apical margin very faintly, but just perceptibly excised. 

^ with the spines of the appendices, somewhat strong, extending to their apices, 
crossing each other towards the apex. 

Expanse 1 1 mm. 

Hab. Haleakala, Maui (5000 ft.) ; i $. 



(19) Nesomicromus minimtis, sp. nov. 

Usually of a dark brown or black colour, sometimes paler, prothorax generally with 
brown or yellowish markings in the darker specimens. 

Anterior wings rounded at the apex, narrow, greyish, or yellowish-grey, near the 
base with two small black spots, one above the other and close together, situated one 
on the cubitus anticus the other on the cubitus posticus. Other black or fuscous spots 
are often present, but these are the most conspicuous, and apparently are never absent. 
Radius normally with 4, rarely with 3 sectors. 

Posterior wings hyaline, nervuration pale and inconspicuous, but towards the apical 
margin the nervures become black, and form a delicate marking. Pterostigma often 
quite pallid and inconspicuous, but passing from yellow in some to testaceous in others, 
and then very conspicuous. 

$ with the appendices of the usual form, their spines very finely serrulate, curved 
upwards, extending slightly beyond their apices. (Plate IV. fig. 11.) 

Expanse 10 — 14 mm. 

Hab. Mountains of Molokai and Hawaii (3000 — 4000 ft.). 



46 FAUNA HAWAIIENSIS 



PsEUDOPSECTRA, gen. nov. 

Allied to Nesomicroinus. Antennae longer than the anterior wings. Maxillary 
palpi with the terminal joint long, acuminate, twice as long as the preceding. Prothorax 
short. 

Anterior wings very short, rounded at the apex, strongly convex above, and 
concave beneath, coriaceous and opaque, costal and apical margins fringed, the dorsal 
one bare. Nervuration without hairs. Costal area with no recurrent nervule at the 
base, and not strongly dilated. Six or seven sectors to the radius. 

Posterior wings very minute in both sexes, forming small subtriangular lobes. 

Z characters as in Nesomicronms, the appendices each furnished towards the base 
on their inferior margin with an upturned spine, which is finely serrulate. 

(i) Psendopsect7'a lobipennis, sp. nov. 

Brown or nearly black, the legs and antennae testaceous, the latter with darker 
annulations, the former with the front and intermediate tibiae with more or less distinct 
fuscous markings. 

Anterior wings brown or yellowish-brown, gradate nervules infuscate forming dark 
lines, nervuration with alternate darker and paler spaces, and there are distinct dark 
spots placed all round the margins of the wings. Posterior wings subtriangular narrowly 
rounded at the apex, with one very thick longitudinal nervure, and one or two others 
much less distinct. 

Appendices of t short, clothed with long hairs, their spines strongly developed, 
crossing one another, and extending to the apices of the appendices. (Plate IV. 

fig- 12.) 

Expanse 9 mm. 

Hab. Haleakala, Maui (5000 ft.). One ^ and one ? taken. 



Nesothauma, gen. nov. 

Antennae short, apical joint of maxillary palpi long, about twice as long as the 
preceding. Head and thorax strongly, densely and roughly punctured. 

Prothorax bilobate in front. Anterior wings very small, their texture almost that 
of the elytra of a Coleopterous insect, the dorsal margin very strongly rounded, the 
costal margin much less strongly. Their surface is strongly convex, but somewhat 
depressed along the margins, which are reflexed. At the base, for about one-third its 
length, the wing is strongly compressed into a strong longitudinal carina, which in the 



NEUROPTERA 47 

natural position of the wings marks off a dorsal from a lateral field. The nervuration 
is not to be definitely made out, but the transverse nervules are excessively numerous 
and divide the wing up into great numbers of small square or subcircular cellules ; the 
nervures bear no hairs, nor is there any trace of a marginal fringe. Posterior wings, 
none. 

The (T characters are similar to those of Nesoniicroimis and Pseudopsectra. 



(i) Nesothauma haleakalae, sp. nov. 

Black, head with pale markings and sometimes the thorax. Antennae variable in 
colour, the basal joint sometimes black, sometimes testaceous. 

Wings black with yellow markings, or yellow with black and fuscous markings, 
very variable in colour. (Plate IV. figs. 13, 13a & i3<5.) 

Appendices of Z narrow towards the apex, and pale in colour, their spines slender, 
long, crossing each other, and extending considerably behind the extremity of the 
appendices. 

Length about 4 mm. Expanse 6 — 7 mm. 

Hab. Haleakala, Maui (5000 ft.). Rare. 



CHRYSOPIDES. 
Anomalochrysa M'Lachl. 

(i) Anomalochrysa princeps, sp. nov. 

Head, thorax, abdomen, legs and antennae flavous. A large species of slender 
form, but variable in size. 

Prothorax with pale and rather long pubescence, gradually attenuate from the base 
forwards, and with a transverse impressed line near the base. 

Wings with pale nervuration, the anterior much broader than the posterior, the latter 
subfalcate. The former are ornamented with sparse black, or blackish, spots. Their 
nervuration is not conspicuous, many of the nervules being in part, or wholly, almost 
colourless, and for the rest of a pale yellow colour, the whole set with pale hairs. The 
gradate nervules form four longitudinal rows of cellules, the two middle ones sometimes 
more or less confused, and one or other of them incomplete. Dividing nervule of third 
cubital cellule received in the apical side of the cellule, instead of in the upper (i.e. the 
cubital nervure), as is usual in the genus. Posterior wings subfalcate, three distinct 
rows of cellules formed by the gradate nervules, a fourth sometimes more or less 
indicated. All the pterostigmata pallid, the wings subhyaline, tinged with yellow. 
F. H. II. 7 



48 FAUNA HAWAIIENSIS 

Abdomen in t with pale pubescence, its apical dorsal plate not strongly dilated, 
erect, clothed along the margins with pale hairs, and armed, on either side, on the 
margin at the base beneath with a long fine spine, the apex of which is strongly 
incurved. Apical ventral valve narrow, tongue-like, not closed against the dorsal plate, 
its surface clothed with long pale hairs. (Plate III. fig. i and Plate IV. figs. 20 
& 20a;.) 

Length 11 — 14 mm. Expanse 33 — 43 mm. 

Hab. Hawaii ; various localities in dense and damp forests (2000 — 3000 ft.). 



(2) Anomalochrysa molokaiensis, sp. nov. 

Closely allied to the preceding, flavous with a bright sulphur-yellow median 
longitudinal stripe extending the whole length of the insect. May be distinguished 
at once from the preceding by the condition of the third cubital cellule, which is of the 
usual form, the dividing nervule received by the cubitus. 

The wings have no black dots. The nervuration is rather more complex ; there are 
four rows of cellules and indications of a fifth formed by the gradate nervules in the 
anterior wings ; four more or less complete rows in the posterior. 

Expanse of $ about 43 mm. 

Hab. Mountains of Molokai (4000 ft.). A single % found drowning in a pool of 
water. 

(3) Anomalochrysa sylvicola, sp. nov. 

Flavous, with a median longitudinal stripe of a sulphur-yellow colour more or less 
distinct. 

Allied to the preceding species but smaller, the wings of very similar shape, the 
posterior pair very distinctly pointed at their apex. Nervuration and hairs pale, but the 
gradate nervules in the anterior wings are more or less black or blackish, and form 
evidently four longitudinal rows of cellules with a tendency to a fifth, the intermediate 
series being more or less irregular ; posterior wings with four, more or less complete, 
rows, one of these sometimes consisting of but few cellules. In the anterior wings the 
gradate nervules (except the lowest series slightly), although dark themselves, have not 
an evident infuscation along their margins. Abdomen clothed with pale hairs, the 
apical dorsal plate not greatly dilated, formed much as in the preceding. (Plate HI. 
fig. 2.) 

Expanse 35 — 37 mm. 

Hab. High plateau of Kauai (4000 ft.). 



NEUROPTERA 49 

(4) Anomalochrysa debilis, sp. nov. 

Form and colour as in A. sylvicola (the yellow longitudinal stripe not alv/ays 
visible in dried examples), but considerably smaller. Nervuration pale ; gradate nervules 
in the anterior wings black or dark forming four rows of cellules, the two middle ones 
not generally completely separated ; posterior wings with three rows. All the cellules 
in the gradate series distinctly but lightly infuscate on each side of the gradate nervules. 
Abdomen clothed with pale hairs, apical dorsal plate of ^ not strongly dilated, erect, the 
spine on each side at the base beneath, exceedingly fine and hair-like, both dorsal and 
ventral plates clothed with pale hairs. 

Length 8 mm. Expanse 24 — 27 mm. 

Hab. Kona, Hawaii (about 3000 ft.). 

(5) Anomalochrysa peles, sp. nov. 

Form, colour and general appearance much as in the two preceding species. From 
A. sylvicola it may be at once distinguished by the infuscation of the front wings, which 
is more conspicuous than in A. debilis, nearly all the cellules of the anterior wings being 
narrowly but distinctly clouded along the nervules ; and from either of those species it 
may be known by the form of the third cubital cellule, the apical portion of which has 
its apical and inferior angle strongly produced outwardly. Of the cellules formed by 
the gradate nervules the upper and lower rows are distinct and very similar in size ; 
between these there are at least three other rows more or less confused. In the 
posterior wings four rows are more or less evident. Abdomen of ^ much as in the 
preceding species. In the only example I have seen, the basal joint of the antennae, 
the head, and front of the prothorax, are darker in colour, with a reddish tinge, but this 
is hardly likely to be a constant character. The hairs on the nervures of the wings are 
rather long and conspicuous. Ventral valve of apical abdominal segment with long 
pale hairs. 

Expanse 32 mm. 

Hab. Kilauea, Hawaii ; a single Z taken. 

(6) Anomalochrysa montana Blackb. 

Anomalochrysa montana Blackburn, Ann. Nat. Hist. (5) xiv. (1884), p. 419. 
Hab. Mauna Loa, Hawaii, at an elevation of nearly 7000 ft. (Blackburn). 
Kilauea, Hawaii (4000 ft.). 



50 FAUNA HAWAIIENSIS 



(7) Anomalochrysa angulicosta, sp. nov. 

Thorax, abdomen, legs and antennae yellowish, probably more or less green in 
life, as traces of that colour can still be detected on the metathorax of the ?. Prothorax 
rather short. 

Wings somewhat broad, costal margin in the $ angulated about the middle, the 
costal area being suddenly narrowed at that point. In the ? the costal margin is simple 
except that it is perceptibly but slightly emarginate before the pterostigma. Nervura- 
tion pale yellowish or green, pterostigma pale. 

In the anterior wings the gradate nervules form three distinct rows of cellules, 
whereof the upper are extremely high, the lower towards the apex show a tendency to 
further division. In the posterior wings there are also three rows. All the cellules of 
the anterior wings containing fuscous spots or lines. Apical portion of the third cubital 
cellule pentagonal. Abdomen with pale pubescence, the apical ventral valve with long 
hairs. (Plate III. fig. 3.) 

Expanse 32 — 34 mm. 

Hab. Mountains of Molokai ; i ? taken at an elevation of 4500 ft. in June, 1893, 
and I $ at 4000 ft. June, 1896. 



(8) Anomalochrysa cognata, sp. nov. 

Female very closely allied to the preceding, but smaller and with narrower wings. 
Colour very similar, probably with a bright sulphur-yellow mediodorsal stripe in life. 
Anterior wings with the cellules infuscate along their nervules ; gradate nervules 
forming four rows of cellules, of which those of the upper row are not very high, and 
have their sides straight (not bent as in the preceding) ; the third row consists of small 
and somewhat irregular cellules. In the posterior wings there are three rows. Dividing 
nervule of the third cubital cellule meeting its apical side, somewhat as in ^. p7'inceps, 
but nearer its upper extremity. The lower portion of this cellule is therefore quadran- 
gular, instead of pentagonal, as is usual in the genus. 

Expanse 31 mm. 

Hab. Mountains near Honolulu (3000 ft.); i % taken in 1896. 

(9) AnomalocJirysa rufescens M'^Lachl. 

Anomalochrysa rufescens, M'Lachlan, Ann. Nat. Hist. (5) xii. (1883), p. 300. 
Hab. Hawaiian Islands (loc. ?). Blackburn. 



NEUROPTERA 51 



(10) Anomalochrysa viridis, sp. no v. 

Green in life, generally fading after drying to yellow, testaceous, or brown. 
Prothorax usually with more or less distinct brown spots. 

Posterior wings considerably narrower than the anterior pair, pointed at the apex. 
Pterostigmata olivaceous in mature examples, at least in the %, paler apparently in 
the t. 

The nervuration although fine is clear and conspicuous in mature examples, but is 
much darker in some than in others, and is set with very fine hairs, which are evidently 
easily abraded, being much more numerous in pallid examples which have recently 
emerged. In the anterior wings, which are moderately broad, but somewhat variable 
in this respect, four rows of longitudinal cellules are formed by the gradate nervules, of 
which the two intermediate rows consist of cellules not much higher than wide, and are 
often more or less confused and incomplete, the nervules, which divide them, failing 
towards the apex of the wing. One example has three rows only. Posterior wings 
with three rows. 

The abdomen is clothed with subdecumbent pubescence, the apical dorsal plate in 
the t, is erect, not very wide, and fringed with dark hairs, which on its ventral (or inner) 
surface at the extreme base extend across it for some distance on each side. I can 
detect no sign of the two fine lateral spines which are present in all the preceding 
species examined, at about the spot where in this species the fringe of hairs is directed 
transversely, as just mentioned. These internally-placed hairs are however themselves 
of a spinose nature, and at their apices are beautifully curved inwards. The apical 
ventral valve is clothed with somewhat long hairs. (Plate III. fig. 4.) 

Expanse 28 — 32 mm. 

Hab. Mountains of Kauai (4000 ft.). 



(11) Anomalochrysa soror, sp. nov. 

Closely allied to the preceding, but probably of smaller average size, with narrower 
wings, and the nervuration, which is green, paler. It may be known at once by the 
shape of the wings which are almost perfectly rounded at their apices, instead of 
forming a distinct angle thereat. 

The form and pubescence of the abdomen is much like that of the preceding, the 
apical dorsal plate is furnished with similar spinose hairs, which are situated along the 



52 FAUNA HAWAIIENSIS 

lateral margins of the plate beneath, forming a longitudinal row ; their colour is dark 
and their apices are strongly curved. 

The characters of the nervuration are those of the preceding species. 

Expanse 25 — 28 mm. 

Hab. Haleakala, Maui, 5000 ft. ; 3 ^, i ?. Probably common but overlooked. 



(12) Anomalochrysa f rater, sp. nov. 

Closely allied to and with the general appearance of A . viridis, which it resembles 
in the form of the wings, and in the nervuration, and varies in the same way. The 
pterostigmata are sometimes pale, sometimes olivaceous and very distinct. 

The ^ is easily known by the pubescence of the abdominal segments, the apical 
portions of which are covered especially towards the sides with long and generally black 
pubescence, which is directed towards the base of the abdomen, while the basal half of 
each segment has only short and inconspicuous hairs. The apical dorsal plate has curved 
hairs, similar to those of the two preceding species, but they are differently disposed, 
occupying a considerable portion of the lower surface of the plate. 

I see no constant differences between the % of this species and that of A. viridis, in 
spite of the conspicuous distinctions between the ^^, but the abdomen of the former sex 
is invariably so greatly distorted in dried examples, that it is useless for the investigation 
of specific characters. (Plate IV. fig. 18.) 

Expanse 26 — -32 mm. 

Hab. Island of Hawaii (2000 — 4000 ft.) ; common and generally distributed. 



(13) Anomalochrysa nana, sp. nov. 

A very small species, bright apple-green in life, with a sulphur-yellow mediodorsal 
stripe from the front of the vertex to the apex of the abdomen. In the dried example 
the body has for the most part become testaceous, or yellowish, and the yellow line has 
to a considerable extent become indistinct, or disappeared. 

Anterior wings narrow, obtuse at the apex, hyaline and somewhat iridescent, the 
nervuration green partly faded to yellowish, and rather conspicuously clothed with hairs, 
which are not very closely set. Gradate nervules very regular, forming an upper and a 
lower entire row of cellules, and between these other two rows for the length of a 
few cellules, after which one of the series of gradate nervules fails and there is only a 
single row of cellules between the upper and lower. There are only about 19 ante- 
pterostigmatic cellules. 



NEUROPTERA 53 

In the posterior wings the nervuration is less distinct. The gradate nervules form 
three rows of cellules. 

Abdomen clothed with fine pale hairs. 

Expanse 23 mm. 

Hab. Molokai mountains (3000 ft), June, 1892 ; i % taken. 



(14) Anomalochrysa pmirosticta, sp. nov. 

Yellow, pronotum with some fuscous markings, meso- and metanota towards the 
sides greenish. Head, legs and antennae concolorous with the body. 

Wings hyaline, iridescent, with yellow nervuration and pterostigmata, and a few 
small black spots on the basal portion of the anterior pair, the spots being situated on 
the nervuration, the gradate nervules are also mostly black, and form four rows of 
cellules, of which the upper and lower are complete, the part between them not being 
divided for its whole length. The third cubital cellule has the apical inferior angle 
greatly produced, so that the length of the upper side of the whole cellule is about 
equal to the lower. The posterior wings are distinctly pointed at the apex, and have 
three rows of cellules in place of the four of the anterior. These rows are quite 
distinct, although the two lower may not be completely divided. The nervuration of 
the wings is set somewhat sparsely with rather long hairs, which are not at all 
conspicuous. 

Male unknown. 

Expanse 37 mm. 

Hab. Olaa, Hawaii (2000 ft.); i $, December, 1896. 



(15) Anomalochrysa longipennis, sp. nov. 

Dark brown, base of antennae and front legs rosy red, thorax especially the 
prothorax also with red markings, meso- and metathorax also partly pale. (Plate HI. 

fig- 5.) 

Wings hyaline, but not perfectly transparent, the anterior pair with a few intra- 
cellular dark spots along the dorsal margin. Nervuration green in life, more or less 
yellow after drying, very similar to that of the preceding species, to which it is very 
closely allied. It differs from A. paurosticta as follows : the dark spots on the wings 
are placed within the cellules, instead of on the nervuration, the wings themselves 
are longer, the hairs on the nervules are evidently closer and more conspicuous, the 
third cubital cellule Is more widely produced apically, and the thickening of the dorsal 



54 FAUNA HAWAIIENSIS 

margin near the base of the wing is more conspicuous, and closes, or nearly closes, the 
apical portion of the cellule above it. 
Male unknown. 

Expanse 42 mm. 

The entirely different colour of this insect and the preceding cannot be taken into 
account in separating the two, as they belong to a group in which different individuals 
of the same species often show differences in coloration precisely similar to that exhibited 
by these two insects, but I believe they are really distinct. 

Hab. Kilauea, Hawaii; i ? taken in August, 1896. 



(16) Anomalochrysa maclachlani Blackb. 

Anomalochrysa maclachlani Blackburn, Ann. Nat. Hist. (5) xiv. (1884), p. 418. 
Hab. Mauna Loa, Hawaii (6000 ft.) in May, 1882. (Blackburn.) 



(17) Anomalochrysa deceptor, sp. nov. 

This is a very variable species, and the extreme forms are totally unlike one 
another in general appearance. 

The following three forms of coloration no doubt constantly occur. 

(i) Head, thorax and abdomen entirely flavous, or partly greenish (in life 
probably sometimes entirely green). (2) Abdomen and sides of the thorax yellowish 
or green, face yellow or pink, and a crimson stripe extending from the front of the 
vertex of the head to the mesothorax, antennae at the base in these examples often 
pink. (3) Whole body dark brown, or with the meso- and metathorax, or one of these 
parts more or less pale, green or yellow. 

Intermediate forms occur, e.g. a large part of the thorax may be yellow or greenish, 
the abdomen dark, and the face pink, &c. 

The wings also vary, and may be hyaline and colourless, or themselves slightly 
greenish, or they may be whitish and opaque ; nor is this condition of the wings 
confined to examples with a particular coloration of the body. These white-winged 
examples generally have dark spots on the anterior pair, but this is not invariably 
the case, and the spots are generally few, and confined to the base of the wing along 
the dorsal margin, but sometimes are more extensive. 

The nervuration is always pale, green or yellow, except that the gradate nervules 
are usually, if not always, more or less dark, and the dorsal margin of the anterior wings 
is sometimes pink. 



NEUROPTERA 55 

Anterior wings in the ? long and narrow, generally somewhat broader in the t ; 
the gradate nervules form four rows of cellules, in the upper of which the cellules are 
very high and narrow ; the intermediate rows are subject to further division, so that in 
some examples as many as 5 or 6 cellules may sometimes be seen in a transverse line at 
some portion of the wing. In one example there are 5 complete rows. 

In the posterior wings there are also 4 rows, but one is often incomplete or may be 
entirely obsolete. The superior row, as in the anterior wings, consists of extremely high 
and narrow cellules, with the sides more or less curved. 

The third cubital cellule has its inferior apical angle considerably produced, and 
the dividing nervure is evidently shorter than its apical margin (i.e. the nervure between 
it and the fourth). 

Apical dorsal plate of <? strongly dilated, with very short hairs on its margin above, 
on the margin beneath they are also short, and form a fringe directed inwardly. 
Apical ventral valve tongue-like, its surface nearly glabrous. (Plate III. fig. 6 and 
Plate IV. figs. 19 & 19 a.) 

Expanse 33 — 38 mm. 

Hab. Hawaii, various localities (2000 — 4000 ft.). Haleakala, Maui (5000 ft.). 
Koolau range, Oahu (above 2000 ft.). 



(18) Anomalochrysa simillima, sp. nov. 

Extremely closely allied to the preceding, flavous, fading to testaceous, or brown, 
the latter variety with the wings whitish, and subopaque. Probably varies in colour 
like the preceding. 

On the anterior wings 5 rows of cellules result from the series of gradate nervules, 
and there is a tendency to further division. 

The species may be distinguished by the following characters ; the cellules of the 
upper row formed by the gradate nervules, especially in the hind wings, are less high, 
and not so narrow in proportion to their height. The third cubital cellule is less 
produced (as a rule hardly at all) at its inferior apical angle, and the dividing nervure is 
about equal to the apical side of the cellule. 

In this, and the other species of the genus, the dorsal margin of the anterior wing 
is greatly thickened at the base just beyond the petiole, and with the nervure above an 
elongate cellule is formed. In the preceding species this cellule, although narrow, is 
distinct and open, but in the present one it is nearly obliterated, owing to the fact that 
the greatly dilated margin in parts touches, or almost touches, the nervure above. 

Hab. High plateau of Kauai (4000 ft.). 

F. H. II. 8 



56 FAUNA HAWAIIENSIS 

(19) AnomalocJirysa gayi, sp. nov. 

Closely allied to A. deceptor, and exhibiting analogous variation, but I have not 
seen any dark-bodied forms such as are found in that species. 

Entirely flavous, or the thorax more or less green, sometimes with a bright 
crimson dorsal stripe on the thorax, and the head and base of the antennae of that 
colour. 

Wings with pale nervuration, green or yellow ; anterior pair with 3 very distinct 
rows of cellules formed by the gradate nervules. Third cubital cellule with its inferior 
apical angle extremely strongly produced, dividing nervule much shorter than the apical 
side, and only about one-third the length of the lower side of the cellule. 

Posterior wings with 3 more or less complete rows of cellules in the gradate series. 

Abdominal characters of t as in A. deceptor. (Plate III. fig. 7.) 

Expanse 32 — 39 mm. 

Hab. High plateau of Kauai. I have much pleasure in naming this distinct 
species after my friend Mr Francis Gay of Kauai, on whose estate the specimens were 
captured, and to whom I am indebted for much assistance when working on that 
island, and for much botanical information. 



(20) Anontalochrysa zoe, sp. nov. 

This species is also closely allied to A. deceptor and probably varies in much the 
same way. 

Yellow with a dorsal longitudinal stripe of a brownish colour, extending over the 
vertex, pro- and mesothorax, or entirely dark, except that the metathorax is more or 
less greenish, and the apical segments of the abdomen pale. Face, antennae and legs 
sometimes crimson, as also the dorsal margin of the anterior wings and sometimes 
of both pairs. 

Wings with pale nervuration, green or yellowish, the wings themselves also some- 
times with a greenish tinge. Gradate nervules in the anterior pair forming 4 distinct 
rows of cellules ; third cubital cellule with its inferior apical angle strongly produced, 
but evidently less so than in A. gayi, its apical side much longer than the dividing 
nervule, which is about half as long as the lower side of the cellule. Sometimes there 
are intracellular spots on the wings, in which case these are less transparent than in 
unspotted examples, just as was the case in A. deceptor. 

Posterior wings with three distinct rows of cellules formed by the gradate series. 

Expanse 33 — 39 mm. 

Hab. Mountains of Molokai (4000 ft. and upwards). Very rare. 



NEUROPTERA 57 

(21) Anomalochrysa raphidioides, sp. nov. 

Head, legs and antennae usually red in the %, the antennae and head sometimes 
nearly black ; in the ^ the antennae and legs are paler, yellowish, and sometimes also 
the head. Thorax and abdomen nearly black in the ?, and sometimes nearly as dark 
in the ^, but probably usually paler, the apical dorsal plate yellow. 

Wings with very distinct and conspicuous nervuration, in the % nearly entirely dark 
brown in colour, in the t paler, although many of the nervules are quite dark. The 
anterior wings are suffused with brown, very conspicuously in the $, in the t less deeply, 
the colour paler (more yellow). The posterior wings are at least for the greater part 
hyaline and finely iridescent. All the pterostigmata in the % are dark brown or 
reddish brown, and extremely conspicuous, but much less so in the t, and pale in 
colour. 

Prothorax attenuate in front, and conspicuously elongate. 

Anterior wings very narrow in the $, much wider in the ^, the gradate nervules 
form three longitudinal rows of cellules, but the second is often partially divided to 
form a few cellules of an additional row, and rarely this row is complete. In the 
posterior wings, which in the % are not much narrower than the anterior, there are three 
distinct rows, which are usually complete or very nearly so. 

The apical dorsal plate of the $ is conspicuously dilated, its pubescence pale, some- 
what short and inconspicuous. The ventral plate, which is applied to the dorsal, has 
only very short and inconspicuous hairs. Apex of the abdomen of % reddish. (Plate 
III. fig. 8.) 

Expanse 31 — 2,"] mm. 

Hab. Kona district of Hawaii, on Mauna Loa and Hualalai, at elevations of 
4000 ft. and upwards. I have taken about a dozen examples of this very distinct 
species, three only being males. 

(22) Anomalochrysa retictdata, sp. nov. 

Dark brown, head more or less red, thorax paler in parts which are probably 
greenish or yellowish in life ; when immature, entirely pale, testaceous, no doubt green 
or yellow in life. Having seen but two mature, and one immature example I cannot 
judge of the variation. 

Wings hyaline and iridescent, scarcely infuscate, pterostigmata olivaceous, nervu- 
ration for the most part dark in both pairs of wings, and very clear and distinct 
throughout. In the anterior wings the gradate nervules form three complete rows of 
cellules, and in each of the three examples examined another row between the second 
and third is represented by 3 cellules. The third cubital cellule is scarcely produced at 
its apical and inferior angle. 

8—2 



58 FAUNA HAWAIIENSIS 

In the posterior wings, there are three distinct rows of cellules formed by the 
gradate nervules, the rows being almost or quite complete. I have not seen a ^ of this 
species. It is probably allied to A. proteus, but the darker and more distinct nervura- 
tion, which is of a simpler character, easily distinguishes it. In size, shape of wings, 
&c. it resembles that species. 

Expanse 35 — 36 mm. ($). 

Hab. Kona, Hawaii (4000 ft.). 

(23) Anomalochrysa biseriata, sp. no v. 

Dark brown, olivaceous, or more or less testaceous. A rather small species with 
the wings clear, hyaline, or almost so, and brightly iridescent. In general appearance, 
but for its smaller size, it greatly resembles the preceding species. 

Wings with the nervuration for the most part dark and very distinct. In the ante- 
rior pair the gradate nervules form three complete rows of cellules, in the posterior pair 
only two. In this respect this species agrees with A. kaematui'a, but that is readily 
separated by the crimson terminal segments of the abdomen, and the dense clothing of 
somewhat long black hairs with which the ventral surface of the abdomen is covered. 
In A. biseriata the hairs on this part are very short, and comparatively inconspicuous. 
The hairs with which the nervuration is set are very fine and but little evident. (Plate 
III. fig. 9.) 

I have not seen a ^ of this species. 

Expanse 24 — 27 mm. 

Hab. Waianae mountains, Oahu (2000 ft.) ; 5 ? taken. 

(24) Ano)nalochrysa hae})iafura, sp. nov. 

Female dark brown, or nearly black, head and scape of the antennae red, legs 
yellow or reddish yellow. Apical segments of the abdomen bright crimson, sometimes 
yellowish. Pterostigmata pale olivaceous, or brightly pink. 

Wings very clear, hyaline, and iridescent, nervuration very distinct, for the most 
part very dark brown, or black. In the anterior pair the gradate nervules form three 
rows of cellules, the lower of which consists of 4 or 5 cellules only, the upper of 
about twice as many. In the posterior wings there are only two rows. 

The abdomen is clothed with obscurelv-coloured hairs on the dark segments, with 
pale ones on the apical. 

Expanse 27 — 32 mm. 

I have not seen the $, but the $ is very distinct from that of any other species. 

Hab. a single specimen taken in October, 1892, and three more in 1896 in the 
mountains near Honolulu (3000 ft.). The three latter were all at rest on a low plant. 



NEUROPTERA 59 

(25) Anonialochrysa ornatipennis Blackb. 

Anomalochrysa ornatipennis Blackburn, Ann. Nat. Hist. xiv. (1884), p. 419. 
Hab. Mauna Loa, Hawaii (4000 ft.) ; \ % (Blackburn). 

(26) AnoinalocJirysa hepatica M'^Lachl. 

Anomalochrysa hepatica M'^Lachlan, Ann. Nat. Hist. xii. (1883), p. 299. 

(Plate HI. fig. 10 and Plate IV. fig. 17.) 

Hab. Haleakala, Maui (4000 — 5000 ft.). Not rare. 

(27) Anonialoclirysa proteus, sp. nov. 

An extremely variable species, entirely yellow, green, or liver-coloured, or reddish, 
often particoloured, the thorax at least being usually more or less yellow or green in 
some parts. 

Wings normally hyaline, colourless or nearly so, sometimes however suffused with 
pale brown, and sometimes with few or many intracellular blotches of a brown colour. 
In these spotted examples the wings tend (as has been noticed in regard to other 
species) to become white and opaque. 

In the anterior wings of the ^ the gradate nervules divide off 5 rows of cellules, 
sometimes a sixth is partly formed, rarely there are only 4 complete, and a fifth indicated 
by a few cells. The cellules of the middle rows are generally more or less irregular 
and confused. In the % there are 4 rows and indications of a fifth, but the wing in this 
sex sometimes has the nervuration as complicated as in the t, although in general 
there seems to be a tendency to a diminution of the number of cellules. 

Posterior wings with 4 rows in the same parts, sometimes more or less of a fifth 
in the ^ ; in the % 4, with one more or less incomplete, sometimes only 3. 

The nervuration in general is pale, yellow or green, but some of the nervules are 
more or less dark, at least those of the gradate series. The third cubital cellule has its 
inferior apical angle not at all, or only a little produced, and the dividing nervule is 
equal, or nearly equal, to the apical side of the cellule. 

Abdomen in the ^ somewhat densely clothed with pubescence, which is generally 
black or dark, and is always directed towards the base of the abdomen, the hairs being 
suberect. The apical dorsal plate is strongly dilated and its ventral valve glabrous or 
nearly so. The % in life probably somewhat resembles the t in the character of the 
abdominal pubescence, as appears from the better preserved examples, but as a rule the 



6o FAUNA HAWAIIENSIS 

abdomen in this sex is greatly distorted in drying, and the characters are lost. (Plate 
III. figs. II, 12 & 13.) 

Expanse from about 29 to about i^ mm. 

Hab. Oahu and Hawaii. Found at elevations of from 2000 — 8000 ft. I have 
examined about 100 examples of this species. The Oahuan specimens seem to be of 
smaller average size. 

(28) Anomalochrysa fulvescens, sp. nov. 

Face and basal joint of the antennae red, brown, or yellowish, pro- and mesothorax 
generally red or reddish brown, the sides of the latter and the metathorax often pale, 
testaceous or yellowish. Abdomen varying in colour from yellow to dark brown. 
Legs varying in colour from pale yellow to red. 

Anterior wings strongly fulvescent, often more or less spotted with dark fuscous, 
or banded, or even entirely suffused with that colour, but always with a fulvous tinge. 

The nervuration in the field of the gradate nervules is extremely complicated, and 
consists for the greater part of large numbers of small cellules, most of which are but 
little higher than wide, forming a dense reticulum, and not regularly disposed in rows. 
The third cubital cellule has its basal side exceedingly strongly thickened in the $, and 
in both sexes the dividing nervule almost meets the upper extremity of the apical side, 
both being received in an evident thickening on the cubital nervure. The colour of the 
nervuration varies from yellow to brown through bright pink. The posterior wings are 
hyaline and iridescent distinctly tinged with fulvous, but much less deeply coloured 
than the anterior pair. Their nervuration is also irregular and confused, but the gradate 
nervules are generally less numerous than in the anterior wings. The pterostigmata 
vary in colour from very pale yellow to reddish. 

In the $ the abdomen has only an excessively minute and short pubescence, the 
apical dorsal plate is not strongly dilated, and has an apical fringe of pale hairs ; on its 
lower margin inwardly it is furnished with a row of spinose hairs, directed inwardly, and 
with curved apices. The ventral valve with only some short pubescence. (Plate III. 
figs. 14. 15 & 16.) 

Expanse 28 mm. (very small $) to t,"/ mm. 

Hab. Haleakala, Maui (4000 — 5000 ft.). 1 have examined about 50 examples of 
this remarkable insect. Of the examples with dark-spotted or banded wings there are 
no two alike, nor do the markings ever agree on the two sides of the same insect. 

(29) Anomalochrysa rhododora, sp. nov. 

Brown, more or less tinged with red, head, basal joint of the antennae, and front 
and intermediate legs red. 



NEUROPTERA 61 

Wings obtuse, the nervuration pale, somewhat densely set with a conspicuous 
pubescence. Anterior pair greyish, the nervures, especially those at the base of the 
wings, and the gradate nervules, with a somewhat faint brownish infuscation. The 
form of the nervuration is very similar to that of A. fulvescens, and the third cubital 
cellule is as in that species, pterostigmata pale. Posterior wings hyaline and iri- 
descent. 

$ characters much as in the preceding. 

Expanse t^t^ — 35 mm. 

Var. $ xerophylla, van nov. 

Anterior wings brown and white, opaque ; posterior pair, white, dull and opaque, 
with a few obscurely coloured spots. The colour of the antennae, legs and body 
is less bright than that of typical examples. The insect in a resting position bears an 
extraordinary resemblance to a small dead leaf that has been attacked by insects. 
Although its general appearance is utterly unlike the type, there is no doubt it is 
an extreme form of that species, the variation being analogous with that of other 
species of the genus. 

Expanse 34 mm. 

Hab. Kilauea, Hawaii. Very rare, i $ \ % taken ; var. xerophylla i %. The 
species is evidently allied to the preceding, but its general appearance is altogether 
different. 

Chrysopa Leach, 
(i) Chrysopa micropliya M^Lachl. 
Chrysopa microphya M'^Lachlan, Ann. Nat. Hist. (5) xii. 1883, p. 299. 
Hab. Common all over the Islands, in the mountains, and in gardens in 
Honolulu, and elsewhere. Probably introduced. 



MYRMELEONIDES. 

(i) Formicaleo perjtirus Walk. 

Myrincieon perjurus Walker, Cat. Neuropt. Brit. Mus. 1852, p. 340. 
Forniicaleo perjurus M^L., Ann. Nat. Hist. (5) xii. 1883, p. 301. 
Myrmeleon violentus Walker, op. cit. p. 348 ; Formicaleo pei'jurus var., M'^L., 
Ann. Nat. Hist. (5) xii. 1883, I.e. 

Hab. Honolulu, Oahu (Beechey expedition) ; Maui (Blackburn). 



62 FAUNA HAWAIIENSIS 

(2) Formicaleo wilsoni M^Lachl. 

F. wilsoni M-^Lachlan, Ann. Nat. Hist. (6) x. 1892, p. 178. 
Hab. Hawaii, locally common, from sea-level to 5000 ft. Lanai (Scott B. 
Wilson). 

ODONATA, 

LIBELLULINA. 
Pantala Hag. 

( I ) Pantala flavescens Fab. 

Hab. Abundant all over the Islands in open country, and in the streets and 
gardens of Honolulu. 

Tramea Hag. 
(i) Tramea lacerata Hagen. 
Hab. All the Islands, in open country on the lowlands. 

Sympetrum^ Newm. 
(i) Sympetrum blackbjirni M'^Lach. 
Lepthemis blackburni, M'Lachlan, Ann. N. H. (5) xii. (1883), p. 229. 
Hab. On all the Islands, in mountain forests, or deep valleys, but not very 
abundant. The species is variable in colour, size, &c. 

AESCHNINA. 
Anax Leach. 
(i) Anax Junius Drury. 
Hab. Very abundant all over the Islands. 

(2) Anax stremms Hagen. 

Anax strenuus, Hagen, Verh. Ges. Wien, 1S67, p. 34 (?) ; Blackburn, Ann. 

N. H. (5) XIV. 1S84, p. 413, (^). 

Hab. Kauai, Maui and Hawaii, and probably all the Islands, in the mountains. 

Not rare but often difficult to catch. 

' A new genus, Nesogonia, Kirby, Ann. Nat. Hist. (7) u. 1898, p. 347, has now been established 
for this insect. 



NEUROPTERA 63 

AGRIONINA. 
Agrion Fabr. 

The dragon-flies of the genus Agrion are amongst the most important repre- 
sentatives of the Order of Neuroptera in the Hawaiian Islands. Several species are 
found on all the more important islands of the group, and the range of many of them 
extends over several islands ; wherein they differ from the greater part of the endemic 
insects, which are for the most part confined to a single island ; or to one or two 
of those which lie most closely together. At the same time, when a series of examples 
of a species from different islands is compared, certain more or less constant differences 
are often observable, especially as regards size. 

Perhaps the most interesting facts relate to the earlier stages, or nymphs, which 
are aquatic and carnivorous. Excluding these dragon-flies and a few water-beetles, 
the insect fauna of the streams and pools is almost non-existent. The Ephemeridae, 
Perlidae, and Trichoptera, usually so numerous, are entirely unrepresented in the 
Islands, although the mountain streams, rising at high altitudes, with their superb 
waterfalls, and various temperatures, appear admirably adapted for many of these. 
It is therefore not a little surprising to find the group of dragon-flies so well represented, 
and that the individuals are so numerous, being on the whole the most conspicuous 
of all the endemic insects. In the absence of the groups above mentioned, I believe 
that their main food-supply comes from without, consisting of such creatures as 
accidentally fall into the water. Under ordinary circumstances this is not great, but 
after rain, when the streams rise very quickly, food becomes abundant. When the 
streams, as is often the case, become nearly dry, large numbers of creatures resort to 
the pools that are left, for the sake of the moisture, and the numbers that come to grief is 
often astonishing, the whole surface being covered with the drowned and drowning. 

There are, however, other species, the nymphs of which live under very different 
circumstances. These have given up their aquatic life, and live hidden at the bases 
of the leaves of a liliaceous plant — Asteiia veratroides. Sometimes a little water 
is held by the plant around the stem, but more often there is merely a collection 
of damp earth and dead leaves. These nymphs would even appear to dislike the 
collections of water, for in wet weather they often crawl half-way up the leaves, instead 
of remaining at the base, where the water accumulates. They differ in some points 
from those which frequent the water ; they are shorter and stouter, and much more 
sluggish, and the caudal appendages are very short and thick, differing therein 
greatly from some of the aquatic species, the appendages of which form beautiful 
tracheal gills. 

On the whole they are without doubt better off as regards a food supply than 
the aquatic species, for there is generally abundance of animal life around them. 
F. H. II. 9 



64 FAUNA HAWAIIENSIS 

A number of interesting beetles breed only in this plant, and minute young of molluscs 
and earthworms are generally abundant in the same, as well as the larvae of small 
moths. Moreover nymphs of various sizes often frequent a single plant, and if hard 
pressed for food the larger, no doubt, devour the smaller individuals. 

In consequence of these habits, some of these species of dragon-flies, although 
their powers of flight are feeble, may often be seen in numbers in localities remote 
from water, and where they would not naturally be looked for. 

These terrestrial nymphs are able to endure extreme drought. On one occasion 
when out shooting, having no more convenient receptacle, I carried a number for 
the greater part of the day in an envelope. In the evening, although very dry, they 
were still quite lively. They were then placed in a tumbler of water, where they 
remained on the bottom, not being able to crawl up the sides. Here they remained 
for a day, apparently as happily as on dry land, when they were taken out and 
preserved. 

M^Lachlan (Ann. Nat. Hist. (5) xii. (1883), p. 238) established a new genus 
Megalagrion for the two species A. blackburni and A. oceanicuin, on the character 
that the post-costal area was complicated by the division of its cellules. We have not 
adopted that genus on account of the instability of the nervuration in this respect. 
(Cf description of A. kauaiense, infra?) 

(i) Ag7'ion xanthomelas Selys. 

Agrion (.?) xanthomelas Selys, Synop. Agrionines, legion Agrion, p. 174. 
M'=Lachlan, Ann. Nat. Hist. (5) xii. 1883, p. 232. 

(Plate V. figs. I, 4 & \a.) 

Hab. Probably occurs all over the Islands. Very common on Oahu, Maui 
and Hawaii. Found both on the coast and in the mountains as high as 3000 ft. 

(2) Agrion pacificum M^Lachl. 

Agrion {?) pacificum M'^Lachl., Ann. Nat. Hist. (5) .xii. 1883, p. 234. 

(Plate V. figs. 6 & 6fl.) 

Hab. Not so abundant generally as A. xanthojfielas , but found on all the Islands. 

(3) Agrion hawaiiense M'^Lachl. 

Agrion (?) hawaiiense M^LachL, Ann. Nat. Hist. (5) xii. 1883, p, 232. 

Hab. " Oahu, at no great elevation above the sea." (Blackburn.) I have not 
met with this species. 



NEUROPTERA 65 



(4) Agrion nigro-hamatum Blackb. 

Agrion (?) nigro-haniatiim Blackburn, Ann. Nat. Hist. (5) xiv. 1884, p. 414. 
(Plate V. figs. 5 & 5«.) 

Race nigro-lineattwi, van nov. 

Of considerably smaller average size than typical specimens and with a distinct 
black line on the upper side of the femora. This point of difference had already 
been noticed by Mr Blackburn in his note on the species (/. c), but the other 
distinctions between the type and his Oahuan examples, as there given, will hardly 
hold in a series of the two forms. 

Hab. Typical examples common on Maui and Molokai. Race nigro-lineatum 
common on Oahu and also found on Hawaii. 

Obs. The bright yellow face and the colour of the eyes, which are bright green 
or turquoise blue on the lower half, and red on the upper, give this species a most 
remarkable appearance when flying around the streams. The colour of the eyes 
fades after death. 



(5) Agrion koelense Blackb. 

Agrion (?) koelense, Blackburn Ann. Nat. Hist. (5) xiv. 1884, p. 417. 

This species varies considerably in several of the points that are used in the ori- 
ginal description. The number of cellules surmounted by the pterostigma is variable in 
all, or nearly all, the Hawaiian species and of no specific value here or in other species. 
The number of cellules between the quadrilateral and nodus is three or four. The 
post-cubitals are usually 16 or 17 in number. The upper margin of quadrilateral 
usually about one-third the length of the lower, in the front wings, but more than 
one-third in the posterior pair. Sometimes, as is also the case in allied species, the 
t assumes more or less the colour of the ?, the femora being pale beneath, and the 
sides of the thorax bearing pale longitudinal stripes as in that sex ; in fact these 
are the only characters of colour by which the % is usually distinguished. The 
valvules are sometimes pale, sometimes black, their appendages, as also the superior 
pair, are black. 

In the t the superior appendages, viewed laterally, have their upper and lower 
margins subparallel, the apical angles produced into two processes of which the upper 
is somewhat more strongly developed than the lower. (Plate V. figs. 7 & 7«.) 

Hab. Mountains of Lanai (2000 ft. and upwards) and in the lao Valley of 
the West Maui Mountains. 

9—2 



66 FAUNA HAWAIIENSIS 



(6) Agrion asteliae, sp. nov. 

Extremely like the preceding in general appearance, sometimes larger, sometimes 
of equal size. Post-cubital nervules usually 19 to 21 in the anterior wings, but in one $ 
there are only 14. The $ characters are extremely like those of A. koelense, but are 
quite sufficiently distinct by the greater development of the inferior apical process 
of the superior appendages which is rather larger than the upper process, and both 
of these processes are somewhat more strongly bent inwards, and also towards each 
other. 

Female with the femora pale beneath, some pale markings on the sides of the 
thorax and sometimes two lateral longitudinal stripes on the dorsum. Superior 
appendages black, valvules pale, their appendages black. In one example the abdomen 
is almost entirely black, in the other the bases of the segments have a very narrow 
distinct pale band, hardly indicated in the former. 

Hab. Mountains of Oahu, 3000 ft. Nymphs taken freely in the leaves of 
growing plants of Astelia veratroides. A single pair taken in copula at Kilauea 
Hawaii (4000 ft.). The individuals from the two Islands do not altogether agree, but 
I doubt whether they could be separated, even with a long series of examples. 



(7) Agrion amaurodytum, sp. nov. 

Dull black, or in the $ (and rarely in the ^) greenish black with metallic lustre. 
t with the head, thorax and base of the abdomen usually more or less covered with 
a slate-coloured pruinose efflorescence, rarely altogether absent in this sex, though 
invariably wanting in the %. Labrum pale along the apical margin. Sides of face 
along the inner margin of the eyes below the antennae pale, these markings connected, 
or nearly so, by a pale line in the %. Vertex black, the region of the post-ocular spots 
occupied usually by two patches of bluish pruinosity. In the $ the post-ocular spots 
are sometimes absent, often very small, but sometimes well-developed, and yellow in 
colour. Prothorax with a transverse pale line in front in the %, and sometimes some 
spots posteriorly, usually unspotted in the ^, but occasionally with markings like those 
of the $. Thorax with a yellow stripe on each side of the dorsum in front in the $, 
which is rarely, more or less distinctly, present in the t ; the sides with pale markings 
(variable) in the $, which are generally less bright in the ^, and sometimes entirely 
absent. Legs black in the ^, the femora more or less pale beneath in the %. Wino-s 
with 19 — 21 post-cubital nervules. Four or five cellules between the quadrilateral and 
nodus. Pterostigma dark, not very elongate. Abdomen in the ? with a more or less 



NEUROPTERA 67 

distinct very narrow pale band at the base of most of the segments ; sometimes these 
bands appear also in the t\ ist and 2nd segments often with a pale spot or line at 
the sides. 

t superior appendages as long as the loth segment, entirely black, or sometimes 
more or less pale inwardly, the apices bent inwards and armed with an excessively 
short and minute spine. On the inferior margin about half-way between the apex 
and base of the appendage there is a stout blunt spine directed inwards, so that the 
appendages may be looked upon as being produced into a long superior and a short 
inferior process. The inferior margin, as viewed inwardly, is not at all strongly 
rounded, so that the upper and lower margins are subparallel. Inferior appendages 
short, pale or black, their apices directed inwards. 

$ superior appendages black, subacute ; valvules pale, their appendages dark. 

Var. A. Two or three males taken high up on the ridges of the West Maui Mts. 
are larger than the typical specimens, and show none of the pale-bluish colour on 
the dorsum of the thorax and abdomen, which would appear to be the case with 
specimens from Molokai only when the % colouration is assumed. 

Race Waianaeanum, var. nov. Oahuan specimens from the Waianae Mts. differ 
from typical ones as follows : 

t With no bluish pruinose efflorescence. Post-ocular spots sometimes present. 
Prothorax with pale spots. Dorsum of thorax with longitudinal lateral stripes. Femora 
pale beneath. Abdomen with distinct narrow pale basal bands to the segments. 

% Rhinarium and part of the post-clypeus pale, legs almost entirely pale, as also 
the appendages of the valvules. Abdomen with a yellowish transverse medio-dorsal 
band on the second segment. 

Race peles, var. nov. 

Much smaller than the preceding forms. Post-cubital nervules usually 13 — 16. 
Typically this race exactly resembles the Molokai examples in general appearance, and 
exhibits similar variation, the pruinose efflorescence being absent, when the markings of 
the % are assumed. There are only three cellules between the quadrilateral and the 
nodus. 

Var. fallax, var. nov. 

Where the typical form abounds an extraordinary variety is sometimes found. 
This has the abdomen more or less red in both sexes, the third segment generally 
almost entirely so ; the legs are almost wholly pale. The post-ocular spots are red 
or yellow, large and connected (or almost so) by a red or yellow line. The pro- 
thorax is much spotted and the longitudinal lateral lines of the dorsum of the 



68 FAUNA HAWAIIENSIS 

thorax are broad and distinct, while the pale marks which border the eyes inwardly 
are connected by a transverse band just behind the posterior margin of the clypeus. 

The measurements of this species are about as follows : 

Typical form and race waianaeaniim. Length of abdomen 36 — 38 mm.; of post, 
wings 25 — 27 mm.; expanse 52 — 54 mm. 

Var. from W. Maui. Length of abdomen 42 mm. 

Race peles and var. fallax Length of abdomen 30 mm. ; of post, wings 19 mm. ; 
expanse 43 mm. 

Hab. Typical specimens from Molokai (3000 — 4000 ft.), and from the lao Valley, 
Maui. Larger and darker var. from high ridges of W. Maui Mts. (4000 ft.). Race 
waianaeanum from the Waianae Mts., Oahu {2000 — 3000 ft.). Race peles widely 
distributed on Hawaii (2000 — 4000 ft.), the var. fallax found with it. 

(8) Agrion eudytum, sp. nov. 

Closely allied to the preceding and very similar in most respects. Black ; ^ with 
the post-ocular spots, those on the prothorax, the lateral longitudinal lines of the dorsum 
of the thorax, the greater part of its sides, the first two segments of the abdomen, 
and the femora beneath pruinose, of a bluish, almost white, colour. Face below the 
antennae obscurely pallid. 

Female with the face yellow below the antennae, as also the post-ocular spots 
which are distinct and a line on the posterior margin of the vertex. Prothorax with 
yellow spots. Thorax with the latero-dorsal lines reaching half-way to the insertion 
of the wings, its sides for the most part pale. Femora almost entirely pale. Abdomen 
with the first two segments and base of the third pale at the sides and also partly 
above. 

Anterior wings with 3 or 4 cellules between the quadrilateral and the nodus, and 
about 20 post-cubitals. Upper side of quadrilateral very short ; one-fourth as long as 
the lower in the anterior, one-third the length of the lower, in the posterior wings. 

^ superior appendages as long as the loth segment, pale inwardly on the basal 
portion, formed very like those of A. amaurodyhim, but more dilated, the inferior 
apical spine evidently smaller, between which and the base, on the inferior margin of 
the appendage there is a minute black tubercle. Inferior appendages black, pale at 
the base, curved upwards, the narrow apical portion longer than in the preceding 
species and much less strongly bent inwards. 

% superior appendages black, valvules and their appendages pale. 

Hab. Kauai (about 1000 ft.), i ^, i $ taken. Probably not rare as I spent 
only an hour or so in the locality and did not revisit it. 



NEUROPTERA 69 



(9) Agrion adytum, sp. nov. 

Closely allied to the preceding, and like it of very slender form in the t- 
Neither sex pruinose. Abdomen dark, black or more or less piceous. Apex of labrum 
pale, otherwise the face below the antennae nearly black in the Z, in the % it is nearly 
entirely pale. Post-ocular spots wanting in the three t examples, but one has a pale 
line on the posterior margin of the vertex in the middle. Dorsal thoracic markings 
present or absent in this sex, in the single $ the latero-dorsal stripes are abbreviated 
into a spot on either side in front. Legs black or piceous, femora more or less pale 
beneath. Abdomen of ? with a yellow spot on the sides of the first and second 
segments. Wings with 18 — 22 post-cubitals in the front pair, and 3 or 4 cellules 
between the quadrilateral and the nodus. 

^ characters very like those of the preceding, but the apical process of the superior 
appendages is shorter and the minute tubercle on the inferior margin between the 
apical inferior tooth and the base is absent. They are distinct at once from those of 
A. amaurodytnm by the small size of that tooth. 

$ superior appendages black, valvules and their appendages pale. 

Hab. Mountains of Kauai (4000 ft.). 



(10) Agrion oresitrophum, sp. nov. 

Male of very slender form with red abdomen, the 6th and 7th segments at least 
black. Face below the antennae red or pale, post-ocular spots of the same colour and 
connected. Prothorax spotted, the front and hind margins also red or yellow. Latero- 
dorsal lines of thorax and median crest similarly coloured. Legs red or pale, with 
black spines. Wings with 14 — 17 post-cubitals in the front pair, and 3 cellules between 
the quadrilateral and the nodus. 

Superior appendages very short, much shorter than the loth segment in lateral 
view, the apex black and directed downwards, viewed inwardly the inferior margin is 
very strongly rounded, and near the base gives rise to an acute spine directed upwards, 
its point attaining to about the level of the superior margin of the appendage. Lower 
appendages long extending considerably behind the superior, curved inwards and 
upwards, their apices black, very slightly emarginate (hardly perceptibly in some 
examples), to form two exceedingly minute spines. Tenth segment angulately excised. 

Two female examples from the same locality may belong to this species ; the 
abdomen is entirely blackish above, with a narrow pale ring at the base of the 
segments. Clypeus and labrum black or blackish at the base. Wings as in the ^. 
(Plate V. figs. 8 & 8fl.) 



^Q FAUNA HAWAIJENSIS 

Length of abdomen 31 — 36 mm. Posterior wing 20*5 — 23 mm. Expanse 
44 — 49 mm. 

Hab. Mountains of Kauai (4000 ft.). Not noticed at lower altitudes. 

A. leptodevias is closely allied to this species and has very similar ^ characters but 
the tubercle of the superior appendages of that species inwardly is differently situated, 
not reaching higher than half the height of the appendage, and the lower appendages 
extend only about as far back as the superior pair. 



(11) Agrio7t orobates, sp. nov. 

Male of the form, size and colour of the preceding. Superior appendages short, 
in lateral view shorter than the loth segment, resting on the lower pair, and terminating 
in a black tubercle. Viewed inwardly, they are much less strongly rounded on the 
lower margin than is the case with that species, and on this margin towards the base 
they are armed with an extremely minute black spine, which is not directed upwards 
to the level of the dorsal margin. Lower appendages long, directed upwards and 
inwards, their apices meeting in the middle line, each armed with two minute but 
distinct spines, the inferior rather the stronger. Tenth segment extremely deeply 
excised, almost to its basal margin. 

A single ? taken in the same locality may belong to this species. It resembles 
those rnentioned under the preceding in most respects but has the abdomen nearly 
entirely red, a dorsal longitudinal line on the second segment and the apex of this 
and the five following segments being black. 

Hab. Mountains of Kauai above Waimea (4000 ft.). A single $ taken in 
May, 1894. 



(12) Agrion lepiodemas, sp. nov. 

A small and very slender species. Face below the antennae entirely red, vertex 
of head posteriorly with a red transverse line. Prothorax with the front and hind 
margin and four spots on the dorsum yellow ; thorax above and at the sides with 
yellow or red stripes, the median ridge red. Legs red, the spines black. Four basal 
and three apical segments of the abdomen red, the intermediate ones dark. Thirteen 
post-cubital cellules in the front, eleven in the hind wings. Three cellules between the 
quadrilateral and the nodus. Upper side of quadrilateral ^ in the upper wings, in 
the lower \ the length of the lower side. 

Superior appendages very short, in lateral view hardly half as long as the loth 
segment, very strongly dilated, almost from the extreme apex, which forms a small 



NEUROPTERA 71 

black tooth directed transversely and downwards as is easily seen in an apical view. 
The strongly curved margin of the dilatation is dark, and inwardly, at about the middle 
of the height of the appendage, gives rise to a distinct black spine or tubercle directed 
upwards. Inferior appendages with acute black apices, and directed inwards and 
upwards. 

Length of abdomen, 29 mm. ; post, wing, 18 "5 mm. ; expanse, 40 mm. 

Hab. Halemano, Oahu, above 2000 ft. A single <J taken in February, 1893. 



(13) Agrion calliphya M^Lachl. 

Agrioui?) calliphya M-^Lachl., Ann. Nat. Hist. (5) xii. 1883, p. 236 (<?). 

I have examined many examples of this species. The abdomen (^) of those from 
Haleakala attains a length of 40 mm. and that of the largest examples from Lanai is 
only a little shorter. In those from Molokai the length is about 35 mm., but some are 
considerably shorter, as is also the case in some from Lanai. The number of cellules 
between the quadrilateral and the nodus is variable, 3, 4, or 5, the number to some 
extent apparently depending on the size of the individual. 

The % typically is very unlike the t in general appearance, the abdomen being 
for the most part black, the base of segments 1 — 7 very narrowly pale or red and 
(except on the first and second) with a red spot near the apex of each of these 
segments. The three terminal segments are red in the middle, black at the sides. 
Clypeus black on the basal portion, thoracic markings as in the ^. Posterior margin 
of prothorax strongly raised and sinuate, its middle portion being strongly produced 
posteriorly. Superior appendages dark, valvules and their appendages pale. (Plate V. 
figs. 9 & 9 a.) 

Race niicrodenias, var. nov. 

Like the typical individuals in colour and form but of uniformly smaller size ; 
length of abdomen about 30 mm. or less, posterior wing 19 mm. Usually 12 — 14 
post-cubitals. Three cellules between the quadrilateral and the nodus. 

This dwarf form of the species is analogous to the race peles of A. amattrodytum, 
which indeed is nearly always found with it. 

The colour of the ^ is somewhat variable, the darkest example has only the first 
three and the last two segments of the abdomen red, while in others it is nearly entirely 
this colour. I have several times taken the sexes coupled. 

Hab. Molokai, Maui and Lanai. Race inicrodemas, abundant on Hawaii. 

F. H. II. 10 



72 FAUNA HAWAIIENSIS ' 

(14) Agrion nesiotes, sp. nov. 

Very slender in form, with the abdomen black, the two basal segments entirely or 
for the most part red above, as also the two apical. Face pale below the antennae, 
clypeus black above in the %. Posterior margin of head pale, post-ocular spots present 
or absent. Thorax with the median carina and lateral dorsal lines yellow or red. Legs 
red or pale. Abdomen with five or six segments more or less pale in tesseral examples, 
otherwise as above. Wings with 16 — 20 post-cubitals. Three cellules between the 
quadrilateral and the nodus. Post-pterostigmatic cellules of the posterior wings more or 
less duplicated. Posterior margin of the prothorax in the % produced backwards in the 
middle, the lobe so formed subtruncate. 

^ superior appendages very long, twice as long as the loth segment, gently curved 
on their outer margins, the extreme apex bent inwards, dilated only for a short distance 
at the base, the inferior margin of the dilated portion armed at its apex inwardly with a 
black spine. Inferior appendages bent inwards, acute at the apex, short, extending 
only about as far back as the apex of the dilated portion of the superior pair. Tenth 
segment strongly raised at the apex about the middle of its margin, and narrowly 
excised, with a regular fringe of hair along the excision. 

$ superior appendages short, black ; valvules and their appendages pale. (Plate V. 
figs. 10 & 10 a.) 

Length of abdomen 36 mm. ; posterior wing 23 mm. ; expanse 49 mm. 

Hab. Hawaii (2000 — 4000 ft); widely distributed, but rarer than the other 
species found on this Island. Taken in Kona, Kau, and Puna districts. 

(15) Agrion Jugoj'um, sp. nov. 

Very slender and elongate in form ; abdomen with several of the basal segments 
red. 

Face below the antennae entirely or almost entirely red or yellowish in the $, base 
of clypeus and labrum more or less black in the $. Post-ocular spots red or yellow and 
always united by a line of the same colour. Pronotum with the anterior and posterior 
margins and some spots on the dorsum red or yellow. Median crest of the thorax, 
a line on either side of the dorsum and the sides more or less of one or other of these 
colours. Legs red or yellow with black spines. Wings with 22 — 24 post-cubital 
cellules, pterostigma bright wine-red in the Lanai examples, generally much less 
brightly coloured in those from Maui. Post-pterostigmatic cellules often forming two 
complete rows, and nearly always forming more or less of a double series at least in one 
wing, but very variable, sometimes slightly infuscate. Three, four, or five cellules 
between the quadrilateral and the nodus. Abdomen of $ with the first four segments 
nearly entirely red, the next three usually black, or dark, the three apical for the most 
part red. In the ? three to five segments from the base are red with their apical 



NEUROPTERA JZ 

margins black, as well as two or three of the terminal segments, but the colour is very 
variable. 

$ superior appendages red on the basal portion, curved inwards but not very 
strongly, about equal in length to the loth segment, strongly and suddenly dilated on 
more than their basal half. The black apex is armed inwardly with an extremely 
minute tubercle ; the dilated portion bears on its apical margin a well-developed black 
tooth, directed inwardly, and situated just above its inferior apical angle (when the 
appendages are viewed inwardly). The lower appendages are short and bent inwards, 
their apices forming a black spine. The apex of the loth segment is emarginate. 

Female superior appendages short, red or piceous, rounded at the apex, not very 
acute. Valvules more or less pale, their appendages of the same colour or piceous. 
(Plate V. figs. 2, II & II a.) 

Length of abdomen 40 — 47 mm. Wing post. 27 mm. Expanse 56 mm. 

Hab. Mountains of Lanai and high ridges of the W. Maui Mts. (4000 ft.). 

(16) Agrion molokaiense, sp. nov. 

Almost exactly like the preceding in general form and appearance and of the same 
dimensions. Three males (one much mutilated) differ as follows. The basal part of 
the clypeus is black, the post-ocular spots are somewhat smaller. The pterostigma is of 
a dark brownish colour. The post-pterostigmatic cellules, especially in the posterior 
wings, are very distinctly clouded. The third segment of the abdomen and the three 
apical ones are nearly entirely red, the two basal are much suffused with black, the rest 
black or blackish. It is probable that none of these distinctions would hold good in a 
long series, but the species is easily known by the genitalia. 

Superior appendages in strict dorsal view, very little produced beyond the extremity 
of the dilated portion : in lateral view the apex is obliquely truncate and each of these 
angles is produced into a minute tooth or spine. The dilatation of the appendages is 
very gradual and the spine at its inferior apical angle is evidently smaller than that of 
the preceding species. 

A single $ which belongs to this species differs from the preceding, so far as I can 
see, only in the smaller size of the post-ocular spots and the almost black basal two 
segments of the abdomen. 

A single $, which is evidently somewhat immature, almost exactly resembles this ? 
in colour, and is so different in colour to that sex as described above, as to make it 
doubtful whether the species could be separated from the preceding without the exami- 
nation of the appendages. 

Length of abdomen, &c. as in A. jtigorum. 

Hab. Mountains of Molokai, above 4000 ft., but one taken at about 1000 ft. less 
elevation. 

10 — 2 



74 FAUNA HAWAIIENSIS 



(17) Agrion oahuense Blackb. 

Agrion (?) oahuense Blackburn, Ann. Nat. Hist. (5) xiv. 1884, p. 414 (^). 

Female with the abdomen black or more or less piceous, most of the segments 
with a narrow pale basal band. Apex of labrum widely, and a transverse band behind 
the clypeus pale. Post-ocular spots present, connected, sometimes obscure. Posterior 
margin of prothorax slightly produced in the middle. Latero-dorsal lines of thorax and 
median carina reddish. Legs pale. Three cellules between the quadrilateral and the 
nodus. Post-pterostigmatic cellules at least in the posterior wings more or less duplicated. 
This series consists of 12 — 14 cellules at least, the divided cellules, which vary from one 
only to many, being considered as single cellules in the count. Superior appendages 
dark, valvules and their appendages generally of an obscure colour, sometimes quite 
pale. Size about that of the ^, and of similar slender form, the abdomen very slightly 
stouter. (Plate V. figs. 12 & 12 a.) 

Hab. High ridges of mountains on Oahu (3000 ft.). Nymphs living between the 
leaves of Astelia veratroides. 



(18) Agrion satelles Blackb. 

Agrion (?) satelles Blackburn, Ann. Nat. Hist. (5) xiv. 1884, p. 414. 

Hab. Haleakala, Maui (4000 ft.). (Blackburn.) I cannot identify this species 
with any known to me. A large form of Agrion deceptor and a similar one of 
A. calliphya are common in the locality. 



(19) Agrion deceptor M"^Lachl. 

Agrion (?) deceptor M'^Lachlan, Ann. Nat. Hist. xii. 1883, p. 235. 

A common species in most parts of the Islands. Examples from Maui, Lanai and 
Molokai are generally larger than the Oahuan specimens, the largest attaining a length of 
46 mm., the front wings often having as many as 22 post-cubitals. The thoracic markings 
and the amount of red on the abdomen are variable. In Maui specimens the two basal 
abdominal segments are often nearly entirely black above in the $. In dark-coloured 
%% the two basal segments are black above, with or without red spots, the third black 
at the base and apex and with a median black line in the central red portion, the 
following segments except for a narrow pale basal band nearly entirely dark, the apex 
of the 8th and 9th and the whole of the loth more or less red. In the ^ the small lower 



NEUROPTERA 75 

tooth of the upper appendages is rather less developed in some specimens than others, 
but there is no constancy in this respect even with examples from the same locality. 
Some specimens show an evident tendency to the form of nervuration on which the 
genus Megalagrioii was founded. 

Hab. All the Islands from Oahu to Hawaii inclusive. 



(20) Agrion vagabundum, sp. nov. 

Allied to A. deceptor M'^Lachl., larger specimens of which it sometimes exactly 
resembles in general appearance. Abdomen usually with the first 6 segments red. 
Face below the antennae pale, clypeus more or less dark in the %. Post-ocular spots 
large and connected. Prothorax spotted, thorax with longitudinal latero-dorsal stripes. 
Legs reddish. Wings usually with 4, sometimes 5, cellules between the quadrilateral 
and the nodus. Four to six post-pterostigmatic cellules in the posterior wings. Ptero- 
stigma not brightly coloured. 

Z superior appendages as long or longer than the loth segment, curved inwards 
but not very strongly, widely dilated on the basal half the inferior margin within 
bearing a very minute tubercle, sometimes hardly visible, towards the apical angle, 
which is rounded off No distinct tooth or spine at the inferior apical angle, such as is 
seen in A. deceptor. Inferior appendages acute, extending rather beyond the dilated 
portion of the upper ones, curved inwards and upwards. 

% superior appendages red or blackish, valvules pale. (Plate V. figs. 13 & 13a.) 

Length of abdomen 31 — 36 mm. ; hind-wing 23*5 mm. ; expanse 50 mm. 

Hab. Widely distributed on Kauai from 1000 ft. to 4000 ft. or more, in the 
mountains. 

Some females which certainly belong to this species show hardly any red colour on 
the abdomen, the variation being much as in A. deceptor, from which the % can hardly 
be distinguished. 

(21) Agrion kauaiense, sp. nov. 

Face below the antennae red or pale, as also the post-ocular spots which are 
connected. Prothorax with red or pale spots ; thorax above and at the sides with red 
or pale lines. Legs red with long black spines. Pterostigma bright red. Abdomen 
with the basal five segments and the two apical ones (more or less) red ; 6th, 7th and 
8th black usually, but the 6th sometimes red. First five segments often with a narrow 
apical black band. 



76 FAUNA HAWAIIENSIS 

Wings with i6 — 22 post-cubital cellules. Upper side of the quadrilateral about \ 
the length of lower in the front wings, somewhat longer in the posterior pair. Post- 
costal area with either a single row of cellules, or with a double row for a considerable 
part of its length, extremely variable in the number of cells that are divided. 

t superior appendages terminating above in a stout hook very strongly curved 
inwards, below this is a distinct but much smaller black tooth, and on the lower margin 
inwardly nearly half-way between the latter tooth and the base of the appendage is 
a third very minute tooth or tubercle. The appendages themselves more or less 
red, the teeth black. Inferior appendages red with black tips curved upwards and 
inwards. 

$ superior appendages generally black, sometimes red, pointed ; valvules more or 
less pale, their appendages generally dark. 

Length of abdomen 33 — 40 mm. ; posterior wing 25 — 28*5 mm. ; expanse 57 mm. 

Allied to^. deceptor but generally larger and with the nervuration of the postcostal 
area more complicated. The t is very distinct by the form of the superior appendages 
which are much more strongly curved inwards at the apex, and also by the position of 
the second tooth which, if the appendage be viewed inwardly, is seen to be situated 
about in the middle of its greatest height, whereas in A. deceptor the tooth is situated 
at the lower angle. The bright colour of the pterostigma is also a useful character. 

Hab. Kauai, widely distributed (2000 — 4000 ft.). 



(22) Agrion oceanicum M'^Lachl. 

Megalagrion oceanicum M'^Lachl., Ann. Nat. Hist. (5) xii. 1883, p. 239 (<^). 

Female with the wings, &c. as in the Z, which it resembles in most respects, but 
the abdomen is stouter and the amount of black colour is more extensive than is usual 
in that sex. Posterior margin of the prothorax strongly raised, angulate in the middle, 
and fringed with long hairs. (Plate V. figs. 14 & \\a.) 

Hab. Island of Oahu, common and widely distributed, frequenting the streams of 
both the mountain ranges. 



(23) Agrion blackburni M'^Lachl. 

Megalagrion blackburni M'^Lachl., Ann. Nat. Hist. (5) xii. 1883, p. 238. 

(Plate V. figs. 15 & 15 «.) 

Hab. Maui, Lanai (Scott B. Wilson), Molokai and Hawaii (windward side). 
Common generally. 



NEUROPTERA 



71 



(24) Ag7'ion heterogamias, sp. nov. 

Male with the abdomen red, that of the % greenish-black. Labium, apex of labrum, 
a space along the inner margin of each eye, connected by a transverse line, pale. 
Vertex of head black with pale post-ocular spots, which are nearly connected by a pale 
line. Prothorax more or less spotted. Dorsum of the mesothorax with the median 
crest and two lateral lines pale, rarely reddish ; sides of thorax more or less pale. Legs 
pale or reddish, with long black spines. Wings with elongate pterostigma of an obscure 
colour. Post-costal area of both pair with a double row of cellules for a great part of 
its length. Abdomen in the ^ generally with the apical half of the 6th, the 7th, and 
more or less of the 8th black, the rest nearly entirely red, and not very variable ; in the 
% the abdomen is nearly entirely greenish-black above, with the basal margins of the 
3rd to the 7th segments very narrowly pale and the apices of the following obscurely 
and narrowly reddish, or testaceous. 

Z superior appendages as long as the loth segment somewhat incurved, the apex 
black and armed at its extremity with a minute tubercle directed inwards ; inwardly, a 
short way behind the apex, is a second minute spine or tubercle, from which the 
appendage is greatly produced downwards and dilated : at the apex of the lower margin 
of the dilated portion inwardly there is a small black spine directed inwardly, and the 
margin itself now becomes thickened and strongly raised, and near the base of the 
appendage gives rise to another black spine directed upwards. Lower appendages 
elongate-triangular, extending beyond the dilated portion of the superior pair, hardly 
curved inwards, red, terminating in a minute tubercle or spine. Tenth segment angu- 
lately excised. 

Female superior appendages acute, triangular, black or obscurely reddish ; valvules 
pale, their appendages darker. (Plate V. fig. 3.) 

Length of abdomen 36 — 40 mm. ; of hind-wing 29 mm. ; expanse 64 mm. 

Hab. Kauai, widely distributed from near sea-level to 4000 ft. Allied to A. 
oceanicum and A. blackburni, the females of which are quite distinct from the present 
species by their general appearance, and the males by the form of the appendages. 

PSOCIDAE. 
In this group we have followed Renter in the description of wing-nervuration. 

Psocus Latr. 
(i) Psocics haleakalae, sp. nov. 

Head with the labrum dark, the post-clypeus pale with longitudinal dark lines, the 
front more or less pale, with dark markings. Eyes in the t not very large, the space 
between them in a front view of the head being (where least) fully equal to, or rather 



78 FAUNA HAWAIIENSIS 

greater than, the width of the two together. In the % they are somewhat more remote. 
The antennae of the ^ near the base are clothed with long and conspicuous hairs, which 
stand out on all sides nearly erect from the surface, but towards the apex they become 
less so. In the $ the antennal hairs are much shorter, and inconspicuous. 

The surface of the mesothorax is dull, and its margins more or less pale. 

Wings hyaline, the anterior slightly infuscate along the margin about the apex. 
At the extreme base there is a dark band, and a very distinct unbroken transverse fascia 
about the middle. The apical portion of the pterostigma, a spot beneath and adjoining 
this, and one on the transverse portion of the inner branch of the cubitus, are also black 
or dark. Posterior wings with an infuscation on the dorsal margin towards the base. 

Legs pale, the tarsi dark. 

Abdomen more or less pale, at least in some examples. The apical ventral 
segment of the ^ is flat at the base, but the apical portion is bent upwards at right 
angles to this and forms a sort of long process. 

Expanse 8"5 — lo mm. 

Hab. Haleakala, Maui (5000 ft.). 

(2) Psociis simulator, sp. nov. 

Resembles the preceding in general appearance. It is rather larger, the median 
transverse fascia of the anterior wings is slightly interrupted about the middle. There 
is a small spot on the cubitus about half-way between the base of the wing and the 
median fascia. In the ^ the eyes are much larger, the width of one of them (in a front 
view of the head) being considerably greater than the distance between its inner 
margin and the nearest ocellus. 

Expanse 10 — W2 mm. 

Hab. Haleakala, Maui (5000 ft.). The ^ is quite distinct by the size of the eyes 
should the slight colour distinctions prove inconstant. 

(3) Psocus imictcs, sp. nov. 

One of the largest Hawaiian species, wings hyaline, faintly clouded. The ptero- 
stigma is for the most part brownish and there is no spot adjoining it. There is a faint 
fuscous stain, forming a transverse band, which passes through the cubitus at its 
furcation. Otherwise the wings are without markings. The median lobe of the 
mesothorax is faintly shining in front. The unique example appears to be a $, but the 
eyes are more prominent than is usual in that sex, although they are widely separated. 
The pubescence of the antennae is short and inconspicuous. 

Expanse i3'5 mm. 

Hab. Haleakala, Maui (5000 ft.) ; unique. 



NEUROPTERA 79 



(4) Psocus hualalai, sp. nov. 

Allied to P. haleakalae, but readily distinguished by the less conspicuous wing- 
markings. The apical portion at least of the pterostigma is black, and there is a spot 
below adjoining it, and one on the transverse portion of the inner branch of the 
cubitus, as in that species. The median fascia is usually represented by two or three 
disconnected spots, which, however, are sometimes sufficiently developed to form a 
transverse band, but this is always more or less interrupted. The head above the 
antennae is for the most part dark. The antennae of the Z have a pubescence similar 
to that of P. haleakalae, and the eyes are widely separated, the width of one of these 
being only about equal to the distance between its inner margin and the nearest ocellus. 
In the % the eyes are still smaller. 

Expanse circa 9 mm. 

Hab. Mauna, Hualalai (about 8000 ft.). 

(5) PsocHS konae, sp. nov. 

Allied to P. simttlator, but distinguished by the lack of the distinct median fascia 
of the anterior wings, which is represented by two or three detached spots. The small 
spot on the cubitus between the base of the wing and the median fascia is also wanting. 
In the wing-markings this species exactly resembles some examples of P. hualalai, but 
from that species the large eyes of the t, which are similar in size to those of P. 
simulator, will easily separate it. 

I have not seen the % of this species. 

Expanse 10 mm. 

Hab. Kona, Hawaii (4000 ft). 

(6) Psocus kauaiensis, sp. nov. 

Of smaller average size than any of the preceding species, and varying greatly in 
the wing-markings. Sometimes the anterior pair have an entire submedian transverse 
dark fascia, as in P. haleakalae, &c., sometimes this is broken up into two or three 
separate spots, as in P. hualalai, &c., or the wings may be without any markings, 
except the dark pterostigma. The most usual form is that which resembles typical P. 
hualalai in markings. The transverse portion of the inner branch of the cubitus is 
dark on the part towards the costal margin, pale towards the other extremity, but it is 
not traversed by a distinct black spot as in all the preceding species, and at the most 
shows a hardly perceptible infuscation along its margins. 

F. H. II. II 



8o FAUNA HAWAIIENSIS 

The Z is readily distinguished by the antennae, the pubescence of the third joint 
being less long and conspicuous ; nor does it stand out suberect from the surface, but 
on both upper and lower surfaces is strongly inclined in the direction of the apex. The 
eyes are large and in a front view of the head the width of one of them is much greater 
than the distance from its inner margin to the nearest ocellus. 

Expanse to 9 mm., generally less. 

Hab. Mountains of Kauai (4000 ft.). Examples from Haleakala (5000 ft.) 
appear identical with those from Kauai, but of slightly greater average size. 



(7) Psocus molokaiensis, sp. nov. 

A small species, the dark markings of the anterior wings as follows : the apex 
of the pterostigma, and a spot adjoining it, the costal margin to the pterostigma 
(generally), more or less infuscation along the cubitus, a spot close to the base of 
its inner branch, and another on the transversely-directed portion of the same, a band 
or line along the dorsal margin from the base to the middle of the wing. These 
markings vary, but are sufficient to distinguish the species, from the fact that their 
general tendency is longitudinal and not transverse as in the other species. In the $ 
the eyes are moderately large, but the space between them is rather wide, being 
about equal to the width of the two taken together. The pubescence of the antennae 
is short and inconspicuous and not erect. Thorax in both sexes with the surface dull. 

Expanse to about 8 mm. 

Hab. Mountains of Molokai (3000 ft.). On Cyathodes. 



(8) Psocus distinguendus, sp. nov. 

Wing-markings and general appearance as in P. haleakalae. The transverse 
fascia is narrow, and sometimes interrupted, and, as in the other species, the markings 
are generally rather more developed in the % than in the ^. 

P. disiinguendus may be known from any of the preceding species by the polished 
surface of the mesothorax, at least in front both of the median and lateral lobes. 

The hairs on the third joint of the antennae of the ^ are shorter than those of 
P. haleakalae. Its eyes are very large, the space (where least) between their inner 
margins being hardly wider than the width of one of these organs. In the $ the eyes 
are comparatively small, and widely separated. 

Expanse min. 8, max. 1 1 mm. 

Hab. Molokai (3000 ft.) ; Lanai, Maui, Hawaii. 



NEUROPTERA 8i 

(9) Psocus oahuensis, sp. nov. 

I have not seen the Z of this species ; the female is very Hke that of P. dis- 
tinguendus, to which it is closely allied. The mesothorax is highly polished, the 
abdomen has bright yellow transverse stripes above. It may be distinguished by the 
evidently wider and more distinct submedian dark fascia of the anterior wings ; the 
absence of the black spot on the cubitus at about the middle of its length, and that on 
the transverse portion of the inner branch of the same nervure is almost obsolete. 

Expanse 10 mm. 

Hab. Waianae Mountains, Oahu. 



(10) Psocus lanaiensis, sp. nov. 

Somewhat similar in general appearance to the other Hawaiian species. The 
anterior wings have a more or less evident transverse median band, the pterostigma 
contains a large dark spot, and there is another adjoining this, without it. The radius 
is pallid, a character which readily distinguishes the species from any of the preceding. 
The cubitus is also often pale (yellow), but sometimes infuscate. Elsewhere also the 
nervuration is widely interrupted by pale spaces, but the terminal furcations of the 
nervures are always dark. 

The face in the ^ in front of the ocelli bears a yellow mark enclosing a dark spot : 
around the ocelli it is dark ; in the $ the face is pale with dark markings ; antennae 
very long, and towards the base in the ^ clothed with rather long hairs ; the eyes 
exceedingly large, the width of one of them even greater than the distance (where 
least) between their inner margins ; in the ? the eyes are small and distant. 

Mesothorax dark with yellow margins, and sometimes a median yellow stripe. 
Abdomen more or less pale above. 

Expanse 8 — 8-5 mm. 

Hab. Mountains of Lanai. 

(11) Psocus sylvestris, sp. nov. 

Very closely allied to the preceding, but the ? has the mesothorax pale. The $ 
appears to vary somewhat in this respect ; it may be distinguished, however, by the 
less conspicuous pubescence of the third joint of the antennae, and the hairs on the 
lower side of this joint are less erect. 

Expanse as in the preceding species. 

Hab. Kona, Hawaii (4000 ft.). 

II — 2 



82 FAUNA HAWAIIENSIS 



(12) Psoctis heterogairdas, sp. nov. 

General appearance very like that of the two preceding species. Thorax almost 
entirely pale. Nervuration almost entirely of a pale yellow colour, the apical furcations 
of the nervures being all pale. There is an evident, but not deep, infuscation along 
the apical margin of the anterior wings, which extends also on to the dorsal margin and 
thence inwardly to the pterostigma, thereby enclosing a clearer space. The eyes of 
the t are evidently smaller, and more widely separated, than those of the preceding 
species. The pubescence of the antennae is subdecumbent and inconspicuous. The ? 
is micropterous, and robust in life, but distorts so in drying that its characters cannot 
be satisfactorily determined from pinned specimens. 

Expanse 8 — 8"5 mm. 

Hab. Mountains of Oahu. 



(13) Psoctis vtonticola, sp. nov. 

Closely allied to P. heterogamias. The apical margin of the anterior wings has an 
infuscate band as in that species, but it is rather more definite, narrower, and less 
diffuse. Of the furcations of the inner branches of the radius and cubitus, which 
terminate in the apical margin of the wing, those of the latter are all dark ; of those 
of the radius the outer is yellow at the extreme base, the inner for fully half its length, 
the apical half dark. The eyes in the $ are not very large, the width of both together 
being subequal to the distance between their inner margins. The pubescence of the 
antennae is inconspicuous and subdecumbent. In the $ the eyes are smaller and more 
remote, and the pubescence of the antennae extremely short and inconspicuous. 

Expanse 8*5 — 9 mm, 

Hab. Mountains of Kauai (4000 ft.). 



(14) Psociis vittipennis, sp. nov. 

Extremely like the preceding in general appearance. The single ^ that I have 
seen has the radius in the anterior wings dark, but it is pale in the females, as in 
all the allied species, and probably would often be so in the ^. Superficially the 
present species may be distinguished as follows : the apical margin of the wing has 
a darker border, which is very distinct and definite, and the inner apical branch of the 
radius is pale for a shorter distance, the yellow part not extending so far as the apical 



NEUROPTERA 83 

furcation of the inner branch of the cubitus. The mesothorax in both sexes is more or 
less smooth and shining. The third joint of the antennae in the t is conspicuously 
clothed with long hairs, which stand out strongly from the surface of the joint. The 
eyes are of about the size of those of the preceding. 

Expanse 9 — 10 mm. 

Hab. Mountains of Kauai (4000 ft.). 

STENOP.SOCU.S Hag. 
( I ) Stenopsocus pulchripennis, sp. nov. 

Head black, the face shining, behind the post-clypeus with indefinite brown 
markings. Antennae black, with very long and slender joints, the third with the 
apex dark, the rest pale, the pubescence is short in general, but at intervals there 
are placed rather long single hairs. 

Anterior wings hyaline, with three transverse dark fasciae, the basal one broad, 
extending from near the radius to the dorsal margin, the middle one very narrow 
along the inner branch of the cubitus. The ajaical fascia extends transversely from the 
costa, at the apex of the pterostigma, to the inner branch of the cubitus, when it 
becomes deflected along that nervure and its terminal furcations, forming an irregular 
dark apical mark, containing two clear spaces. All the nervuration pale as far as the 
basal fascia from the base of the wing, beyond this, dark. Discoidal area quadrangular. 
Pterostigma at the apex obliquely truncate, its inner angle connected by a pale nervule 
to the point of furcation of the inner branch of the radius. Area postica very small, 
triangular, appendiculated at its vertex, the area adjoining it on the apical side very 
large, longer than high, as large or larger than the other two areas formed by the 
apical furcations of the inner branch of the cubitus, taken together. Posterior wings 
clear, nervuration dark. 

Length of anterior wing 2 '5 mm. 

Hab. IVIount Hualalai, Hawaii (8000 ft.). This is certainly no true Stenopsocus, 
but as 1 have seen only one much mutilated example, I have only been able to examine 
it very imperfectly. The wings show no trace of a marginal fringe, nor of hairs on the 
nervuration, but it is possible that this may be due to abrasion. 

Elipsocus Hag. 

(i) Elipsocus montanus, sp. nov. 

Body black or dark brown, the front of the head, the margins of the thorax, 
and generally some lines on the abdomen more or less pale. 



84 FAUNA HAWAIIENSIS 

Post-clypeus and the front of the head along the inner margins of the eyes, 
somewhat shining. Eyes in the Z widely separated, the space between them greater 
than their combined width. Antennae with short pubescence. Wings subhyaline, 
anterior pair usually evidently clouded, but not deeply. Their markings are very 
faint and consist of five fuscous spots along, but not touching, the apical margin, and 
two others placed in a line with the second of the series, at equal distances apart 
towards the base of the wing. There is often also a spot adjoining the pterostigma, 
one on the cubitus about the middle, and another in the angle at the meeting of the 
anal and dorsal nervures, the latter sometimes forming, with infuscations placed inwardly 
to it, a very faint transverse band. The pterostigma varies in colour from being 
entirely pale to a dark brown colour, its apical margin is long. There is a distinct 
but narrow space between the inner branch of the cubitus and the vertex of the 
area postica. The nervuration is nearly entirely dark in both pairs of wings. 

Expanse circa 9 mm. 

Hab. Haleakala, Maui (5000 ft.). 



(2) Elipsocus inconstans, sp. nov. 

This species varies greatly in size and in the wing-markings. In strongly marked 
examples nearly one-half of the anterior wings is fuscous from the base ; there is then 
a transverse hyaline or subhyaline fascia, and the apical portion of the wing is strongly 
variegate with fuscous, enclosing hyaline spots. In other examples the wings are but 
little infuscate on the basal portion, and the other markings may be disconnected and 
form only detached spots, in which case the species bears a considerable resemblance 
to E. montanus, but the markings are never so faint as in that species, and the area 
postica is always, at least partly, infuscate. It may further be at once distinguished 
from the preceding species by the entirely dull surface of the post-clypeus, and of the 
front of the head, which is more depressed transversely. 

The area postica is sometimes free, with a distinct space between it and the inner 
branch of the cubitus, but sometimes its vertex touches (or is connected by an 
excessively short transverse nervule with) that branch. In the latter case a closed 
discoidal area is formed and the examples have no longer the nervuration of the genus. 
This variation is exhibited by examples of the most different superficial appearance, 
as well as in those of similar general aspect, and the nervuration on the two sides of the 
same insect may be markedly different, so that it is evidently in a very unstable 
condition. Examples from the most widely separated islands of the group show 
analogous variation. 



NEUROPTERA 85 

In any case the more highly variegated wings easily distinguish the species from 
any other of the genus found in the Islands. 

Expanse 6'5 — 8'5 mm. 

Hab. Found throughout the group in mountain forests (2000 — 5000 ft.). 

(3) Elipsocus psylloides, sp. nov. 

Head pale, yellowish or testaceous ; thorax and abdomen also more or less pale. 
Wings hyaline, nervuration on the basal part mostly pale, becoming dark towards the 
apex. Space between the inner branches of radius and cubitus evidently, but faintly, 
infuscate. Pterostigma pale. The extremities of the nervures in the dorsal and apical 
margins are slightly infuscate, giving them the appearance of being slightly thickened 
at their apices. The inner branches of the radius and cubitus do not directly meet 
at the points where they are angulated near their base, but the angles are connected 
by a very short transverse nervule. This character, however, appears to vary, as the 
angles approach each other more nearly in some examples than in others. Antennae 
with short pubescence in both sexes. 

Expanse circa 8 mm. 

Hab. Haleakala, Maui (5000 ft.) ; Kona, Hawaii (2000 — 4000 ft.). 

(4) Elipsoc2is criniger, sp. nov. 

Very like the preceding, the nervuration pale, darker towards the apex in the 
anterior wings, but the extremities of the nervures have not the appearance of being 
thickened. There is a distinct (but somewhat faint) fuscous blotch within the 
pterostigma, and another in the area postica, and the basal portion of the inner 
branches of the cubitus and radius, to their point of contact, is infuscate. The species 
may further be recognized by the longer hairs on the front of the head, as well as 
those of the antennae, which are somewhat long and irregular. 

Expanse 8 mm. 

Hab. Kona, Hawaii (2000 ft.); one example. 

(5) Elipsocus deb His, sp. nov. 

Allied to E. psylloides, but smaller, the wings less clear, with a slight yellowish 
tinge. Head, thorax and abdomen for the most part pale. As in that species the 
terminations of nervures are slightly infuscate, and have an appearance of being 
thickened at the margins of the wings. Near the base there is a somewhat extensive, 



86 FAUNA HAWAIIENSIS 

but vague, infuscation, and the clouding of the apical part of the wings is more 
extensive, not being confined to the area between the inner branches of the cubitus 
and radius, although that part is generally slightly more deeply clouded than the rest. 
The apex of the pterostigma is more obtuse, and the inner branches of the radius 
and cubitus are directly in contact at the point where they are angulated. 

Expanse 6 mm. 

Hab. Waianae Mountains, Oahu (above 2000 ft.). 



(6) Elipsocus erythrostictus, sp. nov. 

Allied to the preceding, the apical extremities of the nervures with a similar 
appearance. Anterior wings with a yellowish tinge, the nervuration pale at the base, 
dark towards the apex. This species is readily distinguished by the two crimson or 
pink marks in the pterostigma, one of which is small, and situated at the basal 
extremity, the other at the apical is paler and more diffuse. 

Expanse 5 '2 mm. 

Hab. Kona, Hawaii (2000 ft.). 



(7) Elipsoais vinostis M^Lachl. 

Elipsocus vinostis M^Lachlan, Ann. Nat. Hist. (5) xii. 1883, p. 228. 

The colour of the pterostigma which is given as the main character of this species 
is probably not constant. The species appears to vary greatly in size and colour and 
also in the shape of the pterostigma itself. Some examples have the wings entirely 
infuscate. 

Hab. Many and probably all of the Islands, in the forests. 



(8) Elipsocus inaequi/uscus, sp. nov. 

Allied to E. vinostis, but easily distinguished by the more variegate appearance 
of the anterior wings. These are rather deeply infuscate for the most part but clearer 
in places. The greater part of the area postica, and more or less of the wing adjoining 
it, are always pale and subhyaline, and the region between this area and the nearest 
branch of the inner division of the cubitus is always very perceptibly more deeply 
infuscate than the wing in general. The nervuration is dark and very similar to that 
of E. vinostis, but in that species the infuscation of the wings, whether deep or light, 



NEUROPTERA 87 

is much more uniform over the whole surface. The pterostigma is uniformly infuscate 
like the wings, but not otherwise darkened. 

Expanse 5 '5 — 6 mm. 

Hab. Haleakala, Maui (5000 ft.). 

(9) Elipsocus ii/icrajjiaunis, sp. nov. 

Face yellowish, the post-clypeus with dark lines, generally with a radiate arrange- 
ment, front dark about the ocelli, and a number of very conspicuous dark spots, placed 
closely together near the inner margins of the eyes, on each side. Thora.x and 
abdomen dark. The anterior wings are strongly infuscate, the infuscation extending 
from near the base to the apex, and occupying a large part of the middle of the wing 
and about the apical branches of the inner division of the cubitus. The general appear- 
ance of the wings is dark with paler areas. The latter are chiefly the region around 
the pterostigma, some small spots on the apical margin, part of the area postica and the 
region adjoining this on the basal side, and sometimes more or less of the wing at the 
base. The nervure forming the apical side of the area postica is colourless to the 
vertex, which touches and is confluent for some distance with the inner branch of the 
cubitus. The pterostigma is abruptly truncate at the apex, its apical margin, and more 
or less of the inner margin towards the apex conspicuously blackened, as also the spot 
at its base. The inner branches of radius and cubitus at the point where they are 
angulated do not meet, but the angles are rather distant, and connected by a transverse 
nervule of a white colour. 

Expanse 4"5 mm. 

Hab. Kona, Hawaii (4000 ft.). This minute species is easily distinguished by 
the form of the area postica, and the nervuration is not that of the genus Elipsocus at 
all. Nevertheless I have not cared to separate it, because of the fact that other species 
have a tendency towards a similar nervuration, e.g. E. inconstans (supra, q.v.). 



(10) Elipsocus frigidus, sp. nov. 

Size and general appearance of the preceding, from which it differs in the less 
strongly blackened apical margin of the pterostigma, and the fact that the nervure 
defining the vertex of the area postica, though contiguous to, does not become confluent 
with the inner branch of the cubitus at that point. I have seen but two examples 
which are in poor condition, and it is not improbable that a good series of examples 
would show this to be a variation of the preceding species. (Cf variation of E. 
inconstans)^ 

Hab. Hualalai, Hawaii (8000 ft.). 
F. H. II. 12 



88 FAUNA HAWAIIENSIS 

TERMITIDAE. 
Calotermes Hag. 

(i) Calotermes castanetts Burm. 

C. castanens Burm., M^Lachlan, Ann. Nat. Hist. (5) xii. 1883, p. 227. 

Soldier with distinct eyes and variable in the length of the head beneath. Several 
royal pairs often found in a small colony. The abdomen of the $ appears to undergo 
little or no increase in size after the perfect state is attained. 

Hab. All the Islands, in the mountain forests. Colonies often occupy the whole 
of a large forest tree. 

(2) Calotermes marginipennis Latr. 

C. marginipennis Latr., M'^Lachlan, loc. cit. 

Hab. Towns and settlements. Abundant and destructive to the wooden buildings 
in Honolulu. 

EMBHDAE. 

Oligotoma Westw. 

( I ) Oligotoma insularis M'^Lach. 

Oligotoma insularis M"'Lachlan, Ann. N. H. (5) xii. jo. 227 ; Perkins, Ent. Mag. 
1897, p. 56 (development and habits). 

Hab. Most and probably all of the Islands. Kauai, Oahu, Molokai, Maui and 
Hawaii, from sea-level to 3000 ft. 



NEUROPTERA ' 89 



BIBLIOGRAPHIC LIST (arranged chronologically). 

1. Walker, F. Catalogue of the specimens of Neuropterous insects in the collection of the 

British Museum. London, 1852. 

' Myrmeleon perjunis, n. sp., p. 340 ; 71/. violentus^ n. sp., p. 348. 

2. Hagen, H. Notizen beim Studium von Brauer's Novara-Neuropteren. Verh. Ges. Wien, xvii. 

1867, PP' 31—62. 
2 spp. — Anax sirenuus, n. sp., Oaliu, p. 34; A. severus and ocellatiis, synonyms oi A. Junius. 

3. Selys-Longchamps, E. de. Synopsis des Agrionines. 5"" Legion. Bruxelles, 1877. 

A.? xanthomclas, n. sp., p. 174. 

4. M'^Lachlan, R. Neuroptera of the Hawaiian Islands. Part L Pseudo-Neuropiera. Ann. 

Nat. Hist. (5) XII. 1883, pp. 226 — 240. 

17 spp., 9 of them new. Megalagrion, n. g. 

5. . Neuroptera of the Hawaiian Islands. Pait II. Planipennia. T. c. pp. 298 — 303. 

6 spp., 4 of them new. Anoma/ochi'ysa, n. g. 

6. Blackburn, T. Notes on Hawaiian Neuroptera, with descriptions of new species. Ann. Nat. 

Hist. (5) XIV. 1884, pp. 412 — 421. 

7 n. spp. and numerous notes on habits. 

7. M'^Lachlan, R. Supplementary note on the Neuroptera of the Hawaiian Islands. Ann. Nat. 

Hist. (6) X. 1892, pp. 176 — 178. 

Formicaleo ivilso?ii, n. sp. 

8. Perkins, R. C. L. A collecting trip on Haleakala, Maui, Sandwich Is. Ent. Mag. xxxii. 

(1896), p. 194. 

Brief note on habits of Agrion. 

9. . Notes on Oligotoma insularis ArLach. (Embiidae), and its immature conditions. Ent. 

Mag. 1897, pp. 56—58. 

10. . Notes on some Hawaiian Insects. P. Cambridge Soc. ix. 1897, pp. 373—380. 

Note on habits of Agriott, pp. 373, 374- [Repeated here. Cf p. 63 ante.] 

11. KiRBY, W. F. Description of a new genus of Odonata. Ann. Nat. Hist. (7) 11. 1898, 

PP- 346—348. 
Nesogonia, n. g. for Lepthemis blackburni M'^Lachl. 



ZOOLOGICAL RESULTS based on material from New Britain, New Guinea, Loyalty 
Islands and elsewhere, collected during the years 1895, 1896, and 1897, by 
Arthur Willey, D.Sc. Lond., Hon. M.A. Cantab., Late Balfour Student of the 
University of Cambridge. Demy 4to. Parts I, II, and III. Price 12s. 6d. each. 

PART I. 

The anatomy and development of Peripahis novae-britanniae. By ARTHUR WiLLEY, M.A., D.Sc. 

With Plates I. — IV. and 7 figures in tlie text. 
Metaprotella sandalensis, n. sp. [Caprellidae]. By Dr PAUL MAYER. 

Witii 6 figures in the text. 
On a little-known sea-snake from the South Pacific. By G. A. BOULENGER, F.R.S. 

With Plate V. 
Report on the Centipedes and Millipedes. By R. I. POCOCK. 

With Plate VI. 

5. Account of the Phasmidae with notes on the eggs. By D. Sharp, M.A., F.R.S. 

With Plates VII.— IX. 

6. Scorpions, Pedipalpi and Spiders. By R. I. PocoCK. 

With Plates X.— XI. 

PART II. 

Report on the specimens of the genus Millepora. By SYDNEY J. HiCKSON, M.A., D.Sc, F.R.S. 

With Plates XII.— XVI. 
Report on the Echinoderms (other than Holothurians). By F. JEFFREY BELL, M.A. 

With figures on Plate XVII. and one figure in the text. 
Holothurians. By F. P. BEDFORD, B.A. 

With figures on Plate XVII. 

10. Report on the Sipunculoidea. By ARTHUR E. SHIPLEY, M.A. 

With Plate XVIII. 

11. On the SoHtary Corals. By J. STANLEY GARDINER, M.A. 

With figures on Plates XIX. and XX. 

12. On the postembryonic development of Cycloseris. By J. STANLEY GARDINER, M.A. 

With figures on Plates XIX. and XX. 

13. On a collection of Earthworms. By FRANK E. Beddard, M.A., F.R.S. 

With Plate XXI. 

14. The Gorgonacea. By ISA L. HiLES, B.Sc. 

With Plates XXII. and XXIII. 

PART III. 

15. Orthogenetic variation in the shells of Chelonia. By HANS Gadow, M.A., Ph.D., F.R.S. 

With Plates XXIV. — XXV. and one text-figure. 

16. Enteropneusta from the South Pacific, with notes on the West Indian Species. By ARTHUR 

WlLLEY, D.Sc, Hon. M.A. Cantab. 

With Plates XXVI.— XXXII. and 7 text-figures. 

17. On a collection of Echiurids from the Loyalty Islands, New Britain and China Straits, with an attempt 

to revise the group and to determine its geographical range. By ARTHUR E. SHIPLEY, M.A. 
With Plate XXXIII. 

PART IV. {In the Press.'] 

18. On the anatomy of a supposed new species of Coenopsammia from Lifu. By J. STANLEY 

Gardiner, M.A., Fellow of Gonville and Caius College, Cambridge. With one plate. 

19. On the Insects from New Britain. By D. SHARP, M.A., M.B., F.R.S. With one plate. 

20. Report on the Stomatopoda and Macrura. By L. A. BORRADAILE, M.A., Lecturer of Selwyn 

College, Cambridge. With four plates. 

21. Report on the Slugs. By Walter E. Collinge, F.Z.S., Mason College, Birmingham. 

With two plates. 

22. Report on the Polyzoa. By E. G. Philipps, Newnham College, Cambridge. With two plates. 

23. The Hydroid Zoophytes. By LAURA ROSCOE Thornely, University College, Liverpool. 

With one plate. 

24. Astrosclera willeyaiia, the Type of a new Family of Calcareous Sponges. By J. J. Lister, M.A., 

St John's College, Cambridge. With five plates. 

25. A contribution towards our knowledge of the Pterylography of the Megapodii. By W. P. 

Pycraft, A.L.S., British Museum. With one plate. 

26. The Stolonifera and Alcyonacea. By Sydney J. HiCKSON, M.A., D.Sc, F.R.S., Beyer Professor 

of Zoology in the Owens College, and ISA L. HiLES, B.Sc., Owens College, Manchester. 
With two plates. 

27. Report on the Xeniidae. By J. H. ASHWORTH, B.Sc, Demonstrator in Zoology, Owens College, 

Manchester. With two plates. 

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D. Sharp. 

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Earthworms. 

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FAUNA HAWAIIENSIS 

OR THE 

ZOOLOGY OF THE SANDWICH (HAWAHAN) ISLES 

Being Results of the Explorations instituted by the Joint Committee 

appointed by 

THE ROYAL SOCIETY OF LONDON FOR PROMOTING NATURAL KNOWLEDGE 

AND THE BRITISH ASSOCIATION FOR THE ADVANCEMENT OF SCIENCE 

And carried on with the assistance of those Bodies and of the Trustees of 

THE BERNICE PAUAHI BISHOP MUSEUM AT HONOLULU. 



EDITED BY 

DAVID SHARP, M.B., M.A., F.R.S. 

SECRETARY OF THE COMMITTEE. 



VOLUME II. PART III. 

COLEOPTERA. I. 
By D. SHARP and R. C. L. PERKINS. 

Pages 91—270; Plates VI, VII, VIII, IX, X, uncoloiired. 



II 



Feb. ^ih, 1900. 



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AT THE UNIVERSITY PRESS. 
1900 

[All Rights reserved.'] 



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15 


iftl 



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j^eto Horft: THE MACMILLAN COMPANY. 

Sombaa: E. SEYMOUR HALE. 



COLEOPTERA PHYTOPHAGA 

By D. sharp. 

COLEOPTERA RHYNCHOPHORA, PROTE- 

RHINIDAE, 

COLEOPTERA HETEROMERA, CIOIDAE 

By R. C. L. PERKINS. 



91 



o^s 






■•»o. 






COLEOPTERA\ 

I. COLEOPTERA PHYTOPHAGA. 
By D. Sharp. 

Contents. § i, General remarks ; § 2, Systematic account; § 3, Bibliographic list. 

§ 1. General Remarks. 

The remarkably successful zoological work of Mr Perkins has thrown fresh 
light on many points connected with the entomology of the Hawaiian Islands. 
It would be out of place to discuss most of these points at present, but there is one 
to which I may here allude. It was formerly supposed that very few specimens of any 
of the precinctive^ species could be obtained. As the geographical area of the islands 
is very small, it of course followed as a corollary that the number of individuals existing 
of a species was extremely small. Although there is still much truth in this idea, it has 
nevertheless become clear that in many cases the limitation is far from being so great as 
we supposed it to be, Mr Perkins having procured a large number of individuals of 
numerous precinctive species. He has however succeeded in doing this in most cases 
only by tracing the exact conditions under which they exist. It thus remains true that, 
compared with the great majority of Insects in other parts of the world, the individuals 

^ Owing to the great extent of the Hawaiian Coleoptera this Order will be dealt with in divisions. 
Editor. 

- I use the word precinctive in preference to endemic or peculiar — both of which are in common use 

in the sense of "confined to the area under discussion." The word endemic has been objected to on the 
grounds that its derivation does not indicate geographical restriction, and that it is actually used in medicine 
to signify constant, but not necessarily exclusive, presence in a locality. The word peculiar has no special 
connection with geography and when used by itself is often inadequate to convey the meaning intended ; so 
that when one speaks of " peculiar " forms it is necessary to add some expression to make it clear that 
geographical peculiarity is meant. Hence we require another word that can be used when there is no context 
(as in tables), or when the context does not make it clear that geographical restriction is the point in view. 
Precinctive appears to me suitable for this purpose and I see no objection to the use of the word restricted. 
" Precinctive forms " means therefore forms that are confined to the area specified. 

F. H. II. 12 



92 FAUNA HAWAIIENSIS 

of a species actually existing are very few ; the species having not only a very small 
geographical distribution, but also being found in but few stations in the area. 

The fact that species are sometimes found in considerable numbers at some of 
these stations renders it doubtful whether there is diminished fertility of the individuals, 
as I formerly supposed. At present there is but little evidence on this point. 

Whether the Insects are unusually specialised in their modes of life is also doubtful. 
Most of them are either forest species, or are of alpine or subalpine habits. 

Many forest Insects in other parts of the world are notorious as being but rarely 
met with. In the New Forest, in the south of England, this is certainly the case, and 
it is probably due to the fact that trees are only attacked when they are in certain 
stages of decay, or offer some special kind of shelter, or are accompanied by some 
particular kinds of fungoid growths. 

Variation. In the case of two or three of the species dealt with in this paper 
I have had at my disposal some hundreds of specimens for examination. It therefore 
seems necessary that I should make some remarks on their variation, though on 
considering this I find that the data are very inadequate for a complete discussion. 
Nevertheless it will be well to present certain facts with regard to the genus Plagithmy- 
sus which appears to be one of the most suitable genera for the study of variation in 
Hawaiian Insects. 

Twenty-nine species of this genus are now known. But about nineteen of these 
have each been found only in a single station, and of the remaining ten several are so 
rare that no opinion can be pronounced as to their variation. We have little or no 
information as to the distribution of the species beyond the fact that no species has yet 
been found on more than one island. It would scarcely be possible to find species more 
closely allied than are some of these Plagithniysus and yet, so far as the specimens yet 
procured entitle us to judge, though found in the same locality and at the same season 
they appear to be really distinct. P. varians, P. darwinianus and P. lamarckianus are 
examples of this, all of them having been procured at Kilauea in August ; they are as 
closely allied as any three species can be. 

Varieties of the same species may be found together, and copulate indiscriminately. 
The variation is not indiscriminate. It appears probable that each species has its own 
set of variations and in some cases closely allied species could be defined by their 
variation. As an example I may refer to the colour of the hind legs. In Z'. aeqiialis 
these are either black or red, there are no intermediates although the two kinds copulate 
indiscriminately. The species is as regards this character dimorphic. In the three 
closely allied species — P. varians, P. darwinianus, P. lamarckianus — the facts are 
different. Here also there is variation in the redness or blackness of the legs, but the 
varieties of one species are not completely discontinuous. Moreover the species 
differ from one another in their variation in this respect. P. va^dans is very variable as 
regards the character in question. A highly remarkable variety of it has each leg half 



COLEOPTERA 



93 



red, half black. This variety is very common in this species but does not occur at all 
in either of the two allies though each of them is variable to a greater {P. lamarckianus) 
or less {^P. darwitiianus) extent as to the redness and blackness of the legs. P. varians 
is almost trimorphic as regards the colour of the legs, but the three forms are not 
completely distinct, and one of them is very much rarer than the other two. 

Of P. daj'2ui7tiamis I have not a large series before me, but Mr Perkins examined 
a large number of specimens of this species captured by Mr A. Koebele and tells me 
that the legs are always red or nearly so in it'. 

Thus as regards the character here considered we find that P. aequalis is 
dimorphic, that P. varians is imperfectly trimorphic, and that P. darwinianus is only 
slightly variable. 

Very little information is available as to local variation. We have not received 
any one species of Plagithmysus from more than one locality in any considerable 
number ; the specimens we have received lead me however to anticipate that consider- 
able local difference in the variation exists. As an example I may mention that the 
five specimens of P. aequalis — a species confined to the island of Kauai — found at 
Waimea in April cannot be quite matched by any specimens of the large series found at 
Makaweli in January and February. Whether greater information as to local variation 
would lead to the union of some of the species at present treated as distinct, is a point 
on which I am not at all positive. 

I may however point out that the three closely allied species I have already 
mentioned as being found in one locality have there each a different food-tree. In 
other words they are segregated by food though not by locality. 

The two closely allied species, P. blackburni and P. dai'wiitiamis, exhibit the 
converse of this. Though both are found in the island of Hawaii and have the same 
food-tree they have not yet been found in the same locality. Each of these species has 
however as yet only been once or twice met with. It would be very interesting 
to know whether they ever occur together, and if so whether they still remain 
distinct. 

In connection with this I must not omit to call attention to the peculiar case of 
P. vitticollis with its var. longulus, and of P. bishopi with its V2S . gracilis. In the case 
of these species the variety and typical form inhabit either different localities or different 
trees. We know however but little about these two species, and it is possible that 
these "varieties" may be incipient species, or even closely allied, but actually distinct 
forms. At present it holds good that all the forms I have treated as distinct species 
are segregated either geographically or by food : and this also applies to the two 
varieties just mentioned. 

' A small series only of P. lamarckianus having been procured I do not mention its variation here, but 
may refer the reader to p. no. It appears to be different from either of the other species as regards 
the character under discussion. 

13—2 



94 FAUNA HAWAIIENSIS 

Distribution. Turning our attention to distribution, we note that the Coleoptera 
Phytophaga of the world consist of three great famihes : Chrysomehdae, Bruchidae, and 
Cerambycidae. 

(i) Chrysomehdae. There are about 18,000 species of Chrysomehdae known, 
they are found nearly everywhere except in the Hawaiian Islands, where there is only a 
single species ; and even this has apparently been introduced quite recently. Chry- 
somehdae live on foliage, and the Hawaiian Islands are doubtless well adapted to sustain 
them, so that the absence of this great family has an important significance. 

(2) Bruchidae. A family of about 700 species, of very little importance in this 
connection. The species live in seeds and are carried about by commerce. A few 
specimens have been found at Honolulu. 

(3) Cerambycidae. One of the important families of Coleoptera, consisting of 
upwards of 12,000 species. Fifty-four have been discovered in the Hawaiian 
Archipelago, of which number forty-five are precinctive, while nine have been found 
elsewhere. These nine species belong to as many different genera ; the individuals do 
not differ in the islands to any appreciable extent from those found elsewhere. Owing 
to their comparative lack of interest they have however been but little collected. There 
is nothing in their distribution that throws any light on the origin of the Hawaiian 
fauna. The forty-five precinctive species of Cerambycidae belong to two very distinct 
categories. First there are single species of two genera of Prionini. Each of the 
genera to which they belong has a wide distribution ; the Hawaiian forms are not 
closely allied to any of the external forms ; they are in fact rather more distinct than is 
usual in their genera, so that there is no indication of special affinity with species of any 
other locality. The forty-three precinctive species of Cerambycini are totally 
different from the two Prionini we have discussed : they belong to three genera, very 
closely allied inter se, and form a special group or tribe — Plagithmysides — peculiar 
to the islands, distinguished by the wings having no transverse fold. They seem to be 
nearest allied to the genera Neoclytus and Euryscelis, which are found in North and 
Central America and the Greater Antilles. They are in appearance the most remarkable 
of the Hawaiian Coleoptera and must be considered as a very highly specialised 
group of Cerambycidae. They are remarkable in their habits'. 

Thus there is a very great distinction between these Plagithmysides and the 
non-precinctive Cerambycidae found in the islands ; but the two endemic Prionini 
occupy as it were an intermediate place in their characters. These latter may be 
compared with the Corvus hawaiiensis amongst the birds, while the Plagithmysides 
appear to be analogous with the Drepanidae. 

' See on this point Perkins, Ent. Mag. xxxii. p. 190, and P. Cambridge Phil. See. ix. p. 373. 



COLEOPTERA 95 

§ 2. Systematic account of the Coleoptera Phytophaga. 

Fam. CHRYSOMELIDAE. 

Epitrix Foudr. 

(i) Epitrix parvula Fabr. 
Crioceris parvula Fabr., Syst. El. i. p. 468. 

Hab. Oahu, Waianae mts. ; Molokai. On "poha" or "cape gooseberry" — a 
plant which has no precinctive insect attached to it. 

Mr Jacoby has examined some of our specimens of this insect, and informs me 
that they are not to be distinguished from E. parvula, a species widely distributed 
in Central America and the Antilles. The species was not found by Mr Blackburn 
and has probably been recently introduced. 

Fam. BRUCHIDAE. 

Bruchus L. 

Bruchus, sp. 

Species of this family are doubtless occasionally carried to islands with peas and 
beans, or other seeds. Mr Perkins has brought back two or three specimens found 
at Honolulu. 

Fam. CERAMBYCIDAE. 

Subfam. PRIONINI. 

Parandra Latr. 
(i) Parandra puncticeps Sharp. 

Parandra puncticeps Sharp, Tr. Ent. Soc. London, 1878, p. 202. 

Rare, of nocturnal habits, chiefly found in the month of September, in several 
islands. 

Hab. Kauai, 4000 ft. May, July, September, and October, Perkins. — Oahu, in 
'' Acacia falcata" in the mountains, Blackburn. — Molokai, September 1893, Perkins. 
— Hawaii ; Kona, July and September, Perkins. 



96 FAUNA HAWAIIENSIS 

This species exhibits a great deal of variation in the form and proportions of the 
prothorax, epistome, mandibles, etc. and there may possibly be more than one form in 
the islands. The material before me is not sufficient to decide as to this, owing to the 
development of the individual being subject to much variation ; but the varieties appear 
to be to some extent located in different islands. 

Aegosoma Serv. 
(i) Aegosoma reflextim Karsch. 
Aegosoma refiexum Karsch, Berlin, ent. Zeitschr. xxv. 1881, p. 7, PL I. fig. 11. 

Hab. Kauai, in July, Perkins. — Maui ; Grove Ranche, Karsch ; Haleakala, 
Perkins. — Hawaii ; Kona and Kilauea, Perkins. 

This species is common in the larval state, in wet or dry, decayed wood, and is 
also found in trees that are not entirely dead. The imago is rare, being nocturnal, 
and only occasionally attracted by light. 

Subfam. CERAMBYCINI. 
Group OEMIDES. 

Xystrocera Serv. 
(i) Xystrocera globosa Olivier. 
Cerambix globosus Oliv., Ent. iv. 67, p. 27, p. xii, fig. 81. 

Hab. Oahu ; Honolulu (Perkins). No doubt recently introduced. The species 
is widely distributed outside the islands, and its occurrence in Honolulu has no special 
interest. 

Group HESPEROPHANIDES. 
AsTRiMus Sharp. 

(i) Asirimus hirtus Fairm. 

Stromatium hirtum Fairm., Rev. Zool. i860, p. 50. 

(n. syn.) Astrimus obsciirtis Sharp, Tr. Ent. Soc. 1878, p. 204. 

Hab. Oahu ; Honolulu, Blackburn. ■ 

This was no doubt introduced, and it is doubtful whether it has become naturalized 
or has died out. It was not met with by Mr Perkins. The species is widely dis- 
tributed, having been found in Siam, Fiji, and the Philippine Islands as well as in 
Tahiti. Mr Gahan considers Gelonaetha Thorns, to be the same genus as Astrhmis 
Sharp. I was formerly of the opinion that Stromatium hirtum was a different species, 
but I now accept Mr Gahan's identification of the two. 



COLEOPTERA 97 

Group CALLIDIOPSIDES. 

Ceresium Newman. 

(i) Ceresium simplex Gyll. 

Stenocoriis simplex Gyll. in Schonherr Syn. Ins. App. i, 3, p. 178. 
Ceresium simplex Sharp, Tr. Ent. Soc. London, 1878, p. 203. 

Hab. Oahu ; Honolulu, Blackburn, Perkins; Waialua, Perkins. — Molokai, 
Perkins. — Maui ; Wailuku, Perkins. 

The species is widely distributed in the Pacific islands, but the record of it as 
occurring in New Zealand [Callidium zelandicum Blanchard, Voyage Pole sud, 
IV. p. 272, PL XVII. fig. 4) was probably erroneous; Blanchard's figure does not 
appear to represent this species. It probably occurs on all the islands where the 
introduced Acacia — Prosopis, sp. } — is found. It comes freely to light and has been 
noticed by Mr Perkins on all the islands except Lanai, though I have mentioned above 
only localities that have been specially noted. 



CuRTOMERUS Steph. 
( I ) Curtomerus pilicornis Fab. 

Callidium pilicorne Fab., Ent. Syst. i. 2, p. 327. 
Curtomerus hiteus Steph., 111. Brit. Ent. iv. p. 249. 
Sotenus setiger Sharp, Tr. Ent. Soc. London, 1878, p. 205. 

Hab. Oahu ; Honolulu, Blackburn, Perkins. — Also seen on Maui at Lahaina. 
The species is no doubt liable to be carried about, having been found on one occasion 
in England. Other localities are South America, the W. India islands, and Florida. 



Group CLYTIDES. 

Clytus Laich. 
(i) Clytus cidnicornis Chevr. 

Clytus crinicornis Chevr., Ann. Soc. Ent. France, i860, p. 460. 

Hab. Oahu; Blackburn, Perkins. — Kauai; Perkins*.— Molokai, Perkins. — Abun- 
dant all over the islands, on the introduced Acacia, Prosopis, sp. This is a Central 
American species. 



98 FAUNA HAWAIIENSIS 

Group PLAGITHMYSIDES. 

The difficulty of distinguishing the three genera of Plagithmysides has considerably 
increased in consequence of the discovery of additional forms. The characters 
distinguishing Clytarlus from Plagithmysiis are, that in the former genus the hind 
legs are clubbed and possess a long slender basal stalk. A sexual difference in the 
form of the legs is constant in Clytarlus inasmuch as the femora of the female are less 
clubbed than they are in the male (Plate VI. figg. 2 and 3). On the other hand 
in the large majority of the species of Plagithniysus there is very little difference in 
the legs of the two sexes, and the femora are never abruptly clubbed, and with a 
long thin basal stalk. The females of Plagithniysus funebris, arachnipes and cristatus 
possess however much more slender legs than their males, and therefore considerably 
approach Clytarhis. The female of Clytarlus pennatus, has the legs but little clubbed, 
and therefore there is but little difference between it and the females of the three 
species of Plagithmysiis mentioned. 

The male of Plagithniysus adstatus approximates in the form of the legs to 
Callithmysus. Thus this species is as it were the central point of the group, its male 
approaching Callithmysus, while its female comes near to Clytarlus. 

It is also desirable to mention, in connection with the extraordinary reduction in 
size of the abdomen in this group, that this character is subject in the females of the 
same species to great variation. This is probably in connection with the growth of 
the ovaries. Some of the females have the abdomen very small, while in others, 
of the same sex and species, it is of the size normal in Coleoptera. 

Mr Perkins has observed that the extraordinarily complex stridulating organs of 
these insects are brought into play during breeding, and that very frequent coupling 
occurs, and this to an extent that is almost unknown otherwise in insects ; the 
Plagithmysides are extremely active and are apparently both polyandrous and poly- 
gamous. He has also suggested that the size of the abdomen influences the stridulating. 
So that all the observations that have been made tend to show that the unequalled 
complexity of the stridulating organs is correlative with the unusual system of copula- 
tion and probably plays an important part in it. 

Clytarlus Sharp. 

Clytarlus Sharp, Ent. Mo. Mag. Oct. 1896, p. 238. 
Clytarlus (pars) Sharp, Tr. Ent. Soc. London, 1879, p. 102. 

All the species of Clytarlus, except C. abnormis, are attached to one of the two 
Acacias, Sophora chrysophylla and Acacia koa. They are rare and difficult to obtain. 
The material before me is not adequate for the satisfactory elucidation of these very 
difficult insects. 



COLEOPTERA 99 



(i) Clytarlus filipes Sharp. 

Clytarlus filipes Sharp, Tr. Dublin Soc. (2) iii. 1885, p. 196. 

This species has the punctuation of the upper surface denser and finer than in the 
allies. It. is readily distinguished, so far as the male is concerned by the peculiar struc- 
ture of the dorsal plate of the genital segment, which is unusually large and very deeply 
notched, so that the two angles project as a sort of forceps at the tip of the body. 
(Plate VI. fig. I a) 

Hab. Hawaii ; Mauna Loa, " on Acacia sp." Blackburn : Kona and Kilauea in 
August and September ; Kau, the half-way house, Sepr. Perkins. Apparently rare, we 
having received only 1 5 examples. Attached to Sophora chrysophylla. 

(2) Clytarlus mediocris, sp. no v. 

Minor, fusco-niger, depressus, opacus, albido-vestitus, elytris plus minusve conspicue 
transversim fasciatis, antennis pedibusque fusco-testaceis, femoribus basibus pallidis. 
Long. 4-^ — 6^millim. 

(Plate VI. figs. I and i b.) 

Closely allied to C. filipes, but with a shorter stalk to the hind femora, and with 
the terminal dorsal plate of the male much less deeply excised. 

Hab. Maui ; a series of 38 specimens was found on Haleakala, 3000 — 5000 ft. 
May 1896. Attached to Sophora chrysophylla. 

(3) Clytarlus fragilis Sharp. 

Clytarlus fragilis Sharp, Tr. ent. Soc. London 1881, p. 534; Tr. Dublin Soc. (2) 
III. 1885, PL V. fig. 48. 

Hab. Oahu ; Palolo Valley, November, Blackburn : Waianae mountains, Feb- 
ruary 1896, and April 1892 ; beaten from dead branches of Koa, Perkins. Apparently 
very rare, only four specimens having been obtained. On Acacia koa. 

(4) Clytarlus dedilis, sp, nov. 

Minor, subdepressus, testaceo-brunneus, in elytris subfasciatus, dense punctatus, 
pedibus ex parte majore testaceis, femoribus posterioribus clava abrupta. Long. ? 
9 millim. 

Closely allied to C fragilis, but with the club of the hind femora rather longer, 
F. H. II. 14 



lOO FAUNA HAWAIIENSIS 

and the punctuation of the elytra denser ; the punctuation on the brown band behind 
the scutellum denser and finer. The antennae are rather long, yellow, but little thicker 
at the tip. The pronotum is transversely evenly convex without median or lateral 
crests, there being a single ante-median, and two smaller post-median transverse eleva- 
tions. The punctuation on the apical parts of the elytra is obsolete ; the colour of the 
hairs forming the obscure elytral fasciae is yellow rather than white ; the pubescence on 
the femora is long, and that on the under surface of the body is long and scanty. The 
stalk of the hind femur is very pallid and long, the club is abrupt, brown. Only five 
abdominal segments can be seen in the male, and the hind-margins of the dorsal and 
ventral plates of the fifth are broad and nearly straight. 
Only two specimens of the male sex have been found. 

A third specimen, from Mauna Loa 17. vii, 92, is I have no doubt the female ; it is 
much larger, and has much less abruptly clubbed femora. 

Hab. Hawaii ; Mauna Loa (W.) Sepr. 1892, 4000 ft. ; Hualalai, 5, viii, 92, 5000 ft. 
On Acacia koa. 



(5) Clytarhis obscurus, sp. nov. 

Minor, rufo-obscurus, vel fuscus, albido-pubescens, in elytris subfasciatus, antennis 
pedibusque testaceis, femoribus clava fusca. Long. 4^ — 6-J millim. 

Van Antennis pedibusque ex parte majore nigris. 

This is an excessively variable species in the colour of the elytra, legs and antennae, 
in the length of the white hairs, and in the fasciae of the elytra ; it also varies consider- 
ably in the elongation of the legs ; the variation in size seems to be less than it is in 
some other species. The thorax usually appears somewhat depressed longitudinally on 
each side of the middle, so that the middle appears slightly crested, but the transverse 
elevations on the middle are very small. The legs bear a very distinct, fine long 
pubescence ; the hind femora have a long basal stalk yellow in colour, and a darker, 
moderately long and abrupt club, which is much thicker and more abrupt in the male 
than it is in the female. The genital segment of the male is simple. The punctuation 
in this species is less dense and fine than in C. Jilipes and mediocris, and the simple 
genital .segment of the male clearly distinguishes C. obscuncs from both those species. 
The paler varieties closely resemble C. fragilis and C. dedilis, but may be separated 
therefrom by the less abrupt club of the femora, and by the darker colour. 

Hab. Kauai ; in several localities at a height of about 3000 feet, Perkins ; nearly 
fifty specimens. On Acacia koa. 



COLEOPTERA 



(6) Clytarlus modestus Sharp. 



101 



Clytarlus tnodestus Sharp, Tr. ent. Soc. 1879, p. 104. 

Hab. Maui; Haleakala, 4 — 5000 ft. in April and May, "on Acacia falcata" 
Blackburn ; Haleakala, 5000 ft. May, Perkins. On Acacia koa. 



(7) Clytarlus claviger, sp. nov. 

Secundum sexum diversus. Mas, rufus, albido-pubescens, elytris fere nudis, in 
medio longitudinaliter breviter albido-signatis, femoribus abrupte clavatis, clava nigra. 
Fern., fusca, albido-pubescens, antennis pedibusque testaceis, his gracilibus, clava fusca, 
gracili. Long. % 7^ millim. 

This is closely allied to C. modestus. The thorax is a little cristate along the 
middle. The punctuation of the elytra is dense and fine, and leaves the surface a little 
shining. 

Hab. Hawaii ; Kilauea, June 1895. O^i Acacia koa. 

Although only two males and one female have been obtained, and the two indi- 
viduals of the male sex are very different in appearance, yet I have little doubt that 
they belong to one species, and are distinct from C. niodesttis. The male is readily 
distinguished by the black femoral club, and the female by its larger size and darker 
colour. The individual of the female sex is very different from the males. 



(8) Clytarlus laticollis, sp. nov. 

Rufo-testaceus, parum pubescens ; pedibus minus elongatis, femoribus clava rufa, 
elongata ; thorace densisissime punctato, medio late obtuseque elevato ; elytris fortiter 
punctatis, medio plagis obsoletis albido-pubescentibus. Long. 775 millim. 

Only one specimen — a male — has been obtained. The species has the clubbed 
legs of Clytarlus, but the slender stalk is less elongate than in the normal Clytarlus, 
being in fact only as long as the club. Thorax broad and short, extremely densely 
punctured, almost without pubescence, with a broad elevation — rather than a crest- 
along the middle, and with five or six transverse elevations on this. Elytra much 
narrowed behind, strongly punctured at the base, obsoletely at the tip, with very litde 
pubescence but with a small divided mark of scanty white hairs on the middle. 

Hab. Maui; Haleakala, 5000ft. May 1896, Perkins. On Acacia koa. 

14—2 



I02 FAUNA HAWAIIENSIS 

(9) Clytarlus abnorinis, sp. nov. 

Rufus, pectore fusco ; pedibus minus elongatis ; thorace densisissime punctate, 
supra inaequali, haud cristate ; elytris dense, subtiliter punctatis, singulo posterius ad 
suturam linea pubescentiae albidescente. Long. 9*5 millim. 

Only one specimen — a male — -has been received of this distinct species ; the legs 
are formed much as in C. laticollis, but in several other respects the two species are 
very distinct. The thorax is not like that of any other species ; it is very densely 
punctured above, and has no transverse elevations or scabrosities along the middle, but 
has an obtuse conical tubercle in front, and a broad obscure elevation behind ; the 
surface is also more prominent on each side, so that the dorsum is very uneven. The 
elytra are rather shining ; the line of pubescence along the suture of each extends from 
the tip to a little in front of the middle, and there are a few additional pallid hairs on 
the outside of each line where it ends. The club of the hind femora is elongate and 
very gradual. 

Hab. Hawaii; Olaa, September 1896. Probably attached to Metrosideros or 
Straussia ; not to Acacia, as there are none growing in the locality of its occurrence. 

(10) Clytarlus pennatus Sharp. 

Clytarlus pennatus Sharp, Tr. ent. Soc. London 1881, p. 532. 

(Plate VL fig. 2 $, fig. 3 ?.) 

The specimen originally described was a male. Both sexes vary much in size ; 
the female is more slender, and the legs are much thinner and bear but little pubescence. 
A fine series has now been obtained ; the variation is considerable as regards colour, 
size (8 — 1 2^ millim.) and pubescence, but is nothing like so great as it is in several 
species of Plagithmystis. 

Hab. Maui ; Haleakala, Blackburn ; Haleakala, 5000 ft. February and May, 
Perkins. On Acacia koa. Mr Blackburn originally met with this species "on the 
same tree as Proterhinus lecontei." C. pennatus and P. lecontei, according to Mr 
Perkins' observations, live on different species of trees, so that the earlier record was 
probably based on a mere accidental occurrence. 

(11) Clytarlus nodifer, sp. nov. 

Fuscus, vel testaceus, dense albido-pubescens, elytris vel fasciis interruptis duabus, 
vel maculis quatuor denudatis ; pedibus testaceis minus elongatis ; maris femoribus 
abrupte clavatis, clava fusca dense albido-vestita ; feminae clava gracili fusca, parum 
dense albido-setosa ; tibiis apice fusco. Long. 6 — 12 millim. 



COLEOPTERA 103 

Allied to C. pennatus, but with shorter legs, and more abrupt club to the posterior 
femora. Thorax strongly cristate, with an abrupt anterior vertical elevation, and a well- 
marked posterior elevation bearing two carinae. The colour of the elytra varies, it is 
usually blackish, but is sometimes yellow along the outer margins ; the white pubescence 
is more dense in the male than in the female ; the denuded spaces are densely and 
finely punctured. The femoral club is very dark, in abrupt contrast to the pallid stalk, 
and it is variegated with white pubescence which is much more extensive in the male 
than it is in the female ; in the latter sex the club is only slender, and is not abrupt. 

Hab. Hawaii, Kona, 3000 ft. March 1896. A small series of eleven specimens, 
unfortunately much broken. Attached to Acacia koa. 



( 1 2) Clytarlus longipes, sp. nov. 

Gracilis, dense subtiliter punctatus, opacus ; mas rufus, elytris parce albido-vestitis, 
ante medium fascia angulata denudata, pedibus testaceis, femoribus clava elongata rufa ; 
femina supra fusca, thorace plus minusve evidenter rufo-vittato, elytris tenuiter albido- 
vestitis, haud signatis ; pedibus fuscis, femoribus basi testacea. Long. 6 — 8^ millim. 
(Plate VI. fig. 4 t.) 

This 'species exhibits considerable difference between the sexes. The male to a 
certain extent reminds one of Plagithniys2is both in form and colour, while the female is 
extremely similar to certain females of C. obscuriis. The thorax is scarcely at all 
cristate, but has a well-marked elevation in front, and two behind the middle, it is very 
densely punctate, and has very little pubescence ; it is red in the male, black, obscurely 
striped with red in the female. The elytra are finely, very densely punctured, quite 
dull. In the male they are red, with an angular space covered with white pubescence 
about the scutellum, and also with the apical half bearing a good deal of white pubes- 
cence. In the female they are nearly black, red only at the basal margin, and there is 
white pubescence scantily distributed all over them and not forming a pattern. The 
legs are very different in the two sexes ; in the male the middle femora are shaped as 
in Plagithmystis, but the hind legs are those of Clytarlus, with a rather long club : in 
the female the middle and hind legs are long and slender, with long slender clubs to the 
femora. 

This species was found in sufficient numbers to make it probable that the sexual 
distinctions are fairly constant. It appears to have been found in company with 
C. obsciirus, and some of the females of the two are so similar that comparison of the 
form and length of the legs is necessary to distinguish the most similar examples ; while 
between the males of the two species there is but little resemblance. 

Hab. Kauai ; thirty specimens. On Acacia koa. 



I04 FAUNA HAWAIIENSIS 



(13) Clytarlits annectens, sp. nov. 

Gracilis, rufus, dense punctatus, elytris in dimidio apicale guttis minutissimis 
albidis ; pedibus basi antennisque flavis : femoribus posterioribus clava elongata, 
suboblonga. Long. 7*5 millim. 

Only one individual, a male, has been found of this species which might be placed 
with as much propriety in Plagithmysus as in Clytarliis. The thorax is not crested 
along the middle, but there is a slight anterior elevation, and numerous minute irregular 
scabrous elevations behind it : the sculpture is excessively dense and fine and there is 
almost no pubescence. The elytra are much narrowed behind, and are densely and 
rather closely punctured : there are some excessively minute spots of white hairs on the 
apical part near the suture, and also a few below the shoulder. There is a dense white 
pubescence on the sides of the breast. The middle legs have a long club and a rather 
short but slender stalk ; on the hind legs the stalk is longer and slender, so that I refer 
the species to Clytarlus rather than to Plagithmysus. 

Hab. Kauai, 1894. On Acacia koa. 



Plagithmysus Motsch. 

Stenopterus? Motschoulsky, Bull. Soc. Moscou, 1845, p. 85. 

Plagithmystis, id. op. cit. Pt. II. 1845, p. 41 ; Sharp, C. R. Soc. ent. Belgique, 
1885, p. Ixxiv ; id. Ent. Mo. Mag. 1896, p. 237. 

CEnemona (erroneously), Motschoulsky, Etudes ent. 1852, p. 76. 

Neoclyius (pars), Horn, Tr. Amer. ent. Soc. v. p. 150. 

Clytarlus Sharp, Tr. ent. Soc. London, 1878, pp. 206 and 208. 

Clytarlus (pars), id. op. cit. 1879, p. 102. 

Mr Perkins has obtained fine series of this genus, enabling me to feel pretty 
certain that the species are fairly distinct, and can be readily recognised notwithstanding 
their being in many cases very closely allied. No species has been found on two 
islands. Apparently each species is confined to one kind of tree. Nine of the species are 
attached to one or other of the precinctive Acacias. Closely allied species are attached 
in certain cases to the same species of tree, but it would seem that they are in this case 
geographically separated. P. blackburni and P. darwinianus are both attached to 
Sophora chrysophylla, and both occur on the island of Hawaii, but they have not been 
found in the same locality there. Closely allied species {^P. darwinianus, varians and 
lainarckianus) live in the same locality but are attached to different trees. 



COLEOPTERA 



105 



(i) Plagithmysus vitticollis Sharp. 

Plagithmysus vitticollis Sharp, Ent. Mo. Mag. xxxii. 1896, p. 240. 

(Plate VI. fig. 5 ; ?.) 

Hab. Hawaii, Kilauea, August 1895. On Akala {Rubus sp.) Perkins. 

Var. longuhis Shp. 1. c. 

Hab. Hawaii, Olaa. 

Mr Perkins has now procured a fair series of the variety longulus. None of the 
individuals have any signs of being red on the elytra ; the thoracic vittae are less 
definite, the lateral yellow patch being considerably less extensive and there are few or 
no white hairs on the hind feet. One or two of the individuals are however intermediate 
in these respects, so that it is probable that both var. longuhis and the typical form 
belong to one species which has a slightly different variation in the two localities. As 
the two forms of the species live on different trees, it would be possible to infer that we 
have in this case a species in process of dividing into two. 

(2) Plagithmysus pernmiidus, sp. nov. 

Gracilis, niger, thorace nigerrimo, vittis albidis angustis perdiscretis ; elytris dense 
punctatis, signaturis irregularibus albido-ochraceis ubique ornatis ; femoribus basi flava, 
tibiis tarsisque rufo-obscuris, tarsis posterioribus, dense albido-hirsutis. Long. 1 2 millim. 
(Plate VI. fig. 6.) 

A very distinct species, remarkable on account of the very definite white stripes on 
the jet-black thorax, and by the peculiar irregular but very definite pallid marks on the 
elytra, which along the suture are more confluent than elsewhere so as to form a pair of 
much broken irregular lines ; the white tarsi, in contrast with the dark tibiae, are also 
remarkable. The antennse are thin and long. The hind and middle legs are very long, 
but not very thick. The under surface is black, and very definitely marked with 
patches of pale sulphureous scales ; the posterior aspect of the hind coxae has a very 
definite white patch on it. The femora are sparingly clothed with very short white 
hairs. The sexes apparently differ very little. 

Hab. Kauai, 2000 ft. Feb. 1897: nine examples, Perkins. This feeds on the 
same tree — Bobea sp. ? as P. vitticollis var. longulus, and Callithinysus liirtipes. The 
tree is called "ahakea" by the natives. 

(3) Plagithnysus newelli Sharp. 

Plagithmysus newelli Sharp, Ent. Mo. Mag. xxxii. 1896, p. 240. 

(Plate VI. fig. 7.) 

Hab. Maui ; Brother Matthias Newell, a single example. 



io6 FAUNA HAWAIIENSIS 

(4) Plagithmysjis concolor Sharp. 

Plagithmysus concolor Sharp, t. c. p. 241. 
(Plate VI. fig. 8 ; ?.) 

Hab. Kauai; on " Ohia-ha." = £2i£-eitm, sp. Perkins; Kaholuamano 4,000 ft. 
G. C. Munro. This species varies but little. The series of 27 specimens sent by 
Mr Munro differ but little in colour and markings, and range in length between 
8f 



!J — 18 millim. 



(5) Plagithmys2is solitarins Sharp. 

Plagithnysiis solitarius Sharp, t. c. p. 241. 

Hab. Oahu, Nuanu Valley, 2000 ft., October 1892; Perkins. Unique. 



(6) Plagithmysus cuneahis Sharp. 

Plagithmysus cuneatus Sharp, t. c. p. 241. 

(Plate VI. fig. 9 ; ?.) 

Hab. Oahu, Kaala 1000 ft., March 1893; Perkins. Mr Perkins is unfortunately 
not acquainted with the name of the tree on which this species was found. 

(7) Plagithmysus finschi Har. 

Clytarhis finschi Harold, Mitt. Munchen. ent. Ver. iv. 1880, p. 166; Karsch, 

Berlin, ent. Zeitschr. xxv. 1881, p. 8, PI. I. fig. 13. 
Plagithmysus finschi Sharp, Ent. Mo. Mag. 1896, p. 242. 

Hab. Maui, Olinda, Dr Finsch ; 4000 ft., Blackburn ; Haleakala 4000 ft.. May ; 
Perkins. On Acacia koa (Perkins). According to Blackburn on " A. falcata" but this 
gentleman appears to have erroneously called A. koa, A. falcata. 

(8) Plagithmysus fuuebi'is Sharp. 

Plagithmystis ftmebris Sharp, 1. c. p. 273. 

(Plate VI. fig. 10; t.) 

Hab. Maui, Haleakala in May, and Sept. — Oct. Perkins. On Sophora 
chrysophylla. 



COLEOPTERA 



107 



(9) Plagithmysus pulverulenhis Motsch. 

Stenopterus pidverulentus Motsch., Bull. Mosc. 1895, i. p. 85, PI. I. f. 12. 
Plagithmysiis pulveridentus Motsch., Bull. Mosc. 1845, 11. p. 41, PI. VI. f. 7 ; 

Sharp, Ent. Mo. Mag. xxxii. 1896, p. 242. 
Clytarlus rohistus Sharp, Tr. ent. Soc. 1878, p. 206. 

Hab. Oahu, both ranges ; on Acacia falcata. Blackburn (erroneously) ; Perkins, 
on A. koa. 

(lo) Plagithviysus diana, sp. nov. 

Nigerrimus, dense regulariter griseo-ornatus : thorace subgloboso, griseo, nigro 
trivittato : elytris griseis, areis denudatis nigris, ad suturam lineis latis discretis ante 
medium leniter divergentibus pallide pubescentibus, ante lineas has plaga angulariter 
nigro-pubescente. Long, i 3 millim. 

(Plate VI. fig. II.) 

An extremely distinct, beautifully, though soberly, ornamented species, with the 
sides of the thorax more strongl)- rounded than they are in any other species of the 
genus. There is a slight longitudinal carination of the middle of the thorax. The legs 
are entirely black even at the base, the posterior are elongate, but less incrassate than 
in most other species. The sexes appear to be very similar. 

Hab. Kauai, 4000ft., July, 1896. Eight examples; on Pelea sp. 

(11) Plagithmysus collaris, sp. nov. 

Niger, elytris, antennis pedibusque rufo-obscuris, his femoribus extrorsum tibiisque 
plus minus nigricantibus ; pube pallide sulphurea ornatus ; elytris densissime punctatis, 
singulo linea pubescente pallida anterius versus latus divergente, ante lineas plaga 
nigro-velutina, basi rugosa, parcissime sulphureo-pubescente. Long. 8^ — 16 millim. 
Mas, prothorace ad latera densissime punctato. Fem., prothorace ad latera plus minus 
late laevigato. 

(Plate VI. fig. 13; Z.) 

This is a very distinct species, somewhat similar to P. bishop i and P. vicinus. 
There are no definite thoracic stripes, but the thorax is broadly cristate and scabrous 
along the middle, and there are distinct lateral elevations ; it is a good deal constricted 
at the base. The elytra are dark red, and remarkable on account of their dense 
punctuation. 

Hah. Maui; Haleakala. The larva of this species was found in October, 1S96, 
feeding- in the wood of Pelea sp. ; by carrying this wood to Honolulu Mr Perkins 
obtained a series of 20 specimens in the following December. 

F. H. II. IS 



io8 FAUNA HAWAIIENSIS 

(12) Plagithmysiis bishopi Sharp. 

Plagithmystis bishopi Sharp, Ent. Mo. Mag. xxxii. 1896, p. 242. 

(Plate VI. fig. 12.) 

Hab. Hawaii; Kilauea, 4000 ft., August 1S95, and 1896. On Pelea sp. 
(Perkins). 

Plagithmysus bishopi var. gracilis, Sharp, /. c. The unique individual of this variety 
was found on another tree resembling " pua " but with lighter, smoother bark. 

(13) Plagithmysus vicinus Sharp. 

Plagithmysus vicinus Sharp, t. c. p. 243. 

Hab. Hawaii; Mauna Loa, 3000ft., October 1892; Perkins. 

Undoubtedly a distinct species. On Pelea sp., but not the same species as that to 
which P. bishopi is attached. 

(14) Plagithmysiis bilineatus Sharp. 

Plagithmysus bilineatus Sharp, /. c. p. 243. 

(Plate VI. fig. 14; ?.) 

Hab. Hawaii ; Kilauea, also in the Kona and Puna districts. On the " Ohia 
lehua " tree, Metrosideros polymorpha, Perkins. 

Besides being found in more than one locality this species was obtained in plenty. 
The variation in size (from 9 — 20 millim. long) is considerable, but in respect of colour 
it is less than in some other species. Some specimens are considerably blacker than 
others ; but the blackness shews itself chiefly on the elytra. The hind legs are 
sometimes much more broadly tinted with black towards the tips than they are in 
others, and this is in nearly all cases correlative with a greater extension of the black 
colour on the wing-cases. The head and thorax remain rfed in the whole of the series 
before me. 

(15) Plagithmysus lanaiensis Sharp. 

Plagithmysus lanaiensis Sharp, /. c. p. 244. 

Hab. Lanai ; Halepaakai, 3000 ft, July 1894, Perkins. Probably (but only 
inferentially) attached to Metrosideros polymorpha, this being the predominant tree 
in the locality of capture. 



COLEOPTERA 



109 



(16) Plagiihniysus pcTkinsi Sharp. 

Plagithinysits perkinsi Sharp, t. c. p. 244. 
(Plate VI. fig. 15 ; %.) 

Hab. Hawaii; Mauna Loa, in two localities, June and July. On Myopontm, 
or bastard sandal tree. 

(17) Plagitlunysus v avians Sharp. 

Plagithmysus varians Sharp, /. c. p. 245 (excl. var. y). 

Hab. Hawaii ; Mauna Loa, Kilauea ; Perkins. On Acacia koa. 

This has been found in greater plenty than any other species and I have been 
able to examine about 200 individuals. The variation in the colour of the legs is of 
considerable interest. The extreme base of the femora being always yellow, the other 
parts vary. It may be said that the normal colour of the rest of the femur is black, 
but there are many specimens in which the apical third is bright red : all the inter- 
mediates between this state and the black form occur. The black and the red colours 
are not sharply delimited in these cases. There are two or three specimens in 
which the black colour is nearly entirely absent. And there is another, apparently very 
rare, variety, of an opposite character, in which the black colour is absent from the 
middle but present at the tips. There are only two or three examples of this variety 
known ; and they resemble in this character P. laniarckianus rather than P. darwiiii- 
anus. Though the black colour may be absent from either the tip or the middle it 
is never completely absent from both, there being no individual with quite red legs, 
though the black pigmentation is in a few cases but slight. In consequence of this 
deficiency these individuals bear a resemblance to P. daTwinianus (in which species 
the legs are red). But these specimens do not approach P. darwiniamis in other 
respects. 

The former var. y of P. varians is now P. laniarckianus. 

(18) Plagithmysus darwinianus Sharp. 

Plagitlunysus darivinianjis Sharp, f. c. p. 271. 

(Plate VI. fig. 16 ; Z.) 

Hab. Hawaii; Kilauea, in August. On the "IMamane" tree Sophora chryso- 
pliylla, Perkins. In this species the legs never become black, but in some individuals — 
especially when the size is large — the red is somewhat darker, showing a very imperfect 
condition of blackness, analogous with that exhibited on the apical part of the femur 

15—2 



no FAUNA HAWAIIENSIS 

in some of the Intermediate varieties connecting the forms of P. varians. .In P. 
dm'winianas the colour is however uniformly distributed. 

It should be noticed that the red-leggedness of P. dartoiniamis is not due to mere 
deficiency of black pigmentation as compared with P. varians ; for the black pigmenta- 
tion of the wing-cases is much more extensive and decided than it is in even the 
darkest varieties of P. varians. 



(19) Plagithmysus laniarckianus, sp. nov. 

Rufo-flavus, antennis pedibusque rufis, femoribus basi flava, apice plus minus 
late nigricante ; thorace vel nigro vel rufo, sat conspicue albido-vittato ; elytris 
flavescentibus, dense punctatis, dorso late longeque nigro, singulo ad suturam linea 
albido-pubescente anterius versus latus divergente. Long. 9 — 15 millim. 

Plagithmysus varians, var. y, Sharp, t. c. p. 245. 

Van a, femoribus rufis. 

This species is somewhat doubtfully distinct from P. darivinianiis. The pubescence 
of the under surface and of the legs is less elongate, the antennae are slightly thinner 
and usually darker in colour towards the tip, and the legs are usually more or less 
broadly black. As minor distinctions we may mention that the elytra are usually 
broader at the base, and are there never distinctly marked with white hairs ; their 
black portions and the white lines on them are usually a little more extensive ; the 
thoracic crest is a little more elevated on its posterior part. 

Hab. Hawaii; Kilauea, August 1896, on the " mamake " or paper-mulberry. 
Although this lives in the same locality as P. darwinianiis it frequents different trees, 
being attached to the Urticaceous trees, Pipturus albidus, and Urera sandwicensis. 
I have seen -^t^ specimens ; the varieties with entirely red femora are very similar to 
P. dai-winianus, and it is possible that it may prove that the two forms are not distinct 
species when more specimens are obtained. There were several specimens of P. 
laniarckianus in the series I originally referred to P. darwinianus, and I am indebted 
to Mr Perkins for directing my attention to the fact that there are probably two 
distinct forms. 



(20) Plagithmysus albert isi Sharp. 

Plagithmysus albert isi Sharp, Ent. Mo. Mag. xxxiii. 1897, p. 12. 

(Plate VI. fig. 17; 2.) 

Hab. Oahu ; West Honolulu, Feb. 25th, 1874; Signor d'Albertis. There is 
reason to fear that this species may now be extinct. 



COLEOPTERA 1 1 1 

(21) Plagithmysiis pidvillatus Karsch. 

Clytarlus pnhillatus Karsch, Berlin, ent. Zeit. xxv. 1881, p. 9, PI. I. f. 14. 

Hab. Maui ; " Grove Ranche," Karsch ; Haleakala, 5000 ft., Perkins. We have 
received only three individuals, and the species was described by Karsch on one female. 
It is one of the least attractive species, being remarkable for its uniform colour and 
freedom from definite ornamentation as well as for the very dense sculpture of the 
elytra. The species is probably verging on extinction owing to the destruction in this 
locality of the forest tree it inhabited. Insects are no longer to be found at Grove 
Ranche ; but Mr Perkins procured his examples about 2000 ft. directly above its 
situation, on Metrosideros polymorpha. 

{22) Plagithmysus blackburni Sharp. 

Clytarlus blackbui'iii ShsiV^, Tr. Dublin Soc. (2) in. 18S5, p. 195, PI. V. f 47. 

Plagithmysus blackburni 'i\\'AX^, Ent. Mo. Mag. 1896, p. 271. 

Hab. Hawaii; found by Mr Blackburn [on " Acacia falcata'''] on Mauna Loa 
in May ; by Mr Perkins at Kona in the autumn, on Sophora chrysophylla. The only 
variation that occurs in the colour of the legs is the one that I have mentioned as being 
so rare in P. varians. 

(23) Plagithmys7is sulphurcscens Sharp. 

Plagithmysus sulphurescens Sharp, Ent. Mo. Mag. xxxii. 1896, p. 271. 

(Plate VI. fig. 17; $.) 

Hab. Hawaii ; Kilauea, July 1895. The name of the tree to which this species 
is attached is unfortunately not known. 

Since P. lamarckianus has been distinguished from P. varians, it should be 
mentioned that P. sulphurescens is allied to the former more than to the second of 
these species. 

(24) Plagithmysus speculifer Sharp. 

Plagithmysus speculifer Sharp, t. c. p. 272. 

Hab. Maui; head of the Jao Valley, June I S94. Unique. Food-tree unknown. 

(25) Plagithmysus aestivus Sharp. 

Plagithmysus aestivus Sharp, t. c. p. 272. 

Hab. Molokai; in June and August, Perkins. On Metrosideros polymorpha. 



112 FAUNA HAWAIIENSIS 

(26) Plagithmysus aequalis Sharp. 

Plagithmysus aequalis Sharp, Ent. Mo. Mag. 1896, p. 273. 

Hab. Kauai ; Mr Perkins procured a very large series of this species at MakaweU, 
January and February 1897, on Acacia koa. And I have also seen a few specimens 
from Waimea and Koholuamano. 

This species is dimorphic as regards the colour of the legs, which are either red or 
black without intermediate states. The individuals with red legs are very numerous. 
Specimens with red thorax are very rare. The chief variations in these respects are as 
follows. 

Van /3, pedibus capiteque rufis. 

Var. y, pedibus, capite thoraceque rufis. 

I have not before me any specimens in which the legs are black without the head 
being so, but it is probable that such specimens occur, and, if so, would form var. a. 

Four individuals found in April 1895 form a variety with shorter thorax, having 
more rounded sides, with more white pubescence on the upper surface, and the posterior 
femora a little differently shaped. One of the two originally described females is a very 
peculiar individual and may perhaps belong to another species. 

(27) Plagithmysus mimroi, sp. nov. 

Rufo nigroque variegatus ; elytris rufis, singulo posterius plaga nitida nigricante, 
fere aequaliter ubique albido-guttulatis ; thorace albido vittato, femoribus bast flava. 
Variat thorace femoribusque rufis, vel nigris. Long. 11 — 12 millim. 

(Plate VT. fig. 19; ?.) 

Closely allied to P. aequalis, but with the elytra nearly evenly spotted all over with 
white flecks of pubescence, there being however a small area in front of the middle 
densely and finely punctured. There is no definite spot of white on the mesothoracic 
episterna. The hind legs are very long, the femora moderately thick, and there is 
extremely little difference between the sexes. 

Hab. Kauai. We are indebted to G. C. Munro, Esq., for the discovery of this 
species : he obtained si.x specimens of it in July 1897, 2000 ft. above Waimea. 

(28) PlagitJiinys7t,s arachnipes Sharp. 

Plagithmysus arachnipes Sharp, /. c. p. 274. 

(Plate VI. fig. 20; ?.) 

Hab. Kauai. May 1895. Perkins. On Acacia koa. 



COLEOPTERA 



113 



Both sexes of this species may be readily distinguished from P. aequalis by the 
longer, slender basal part of the femora : as this part is pallid yellow in colour, the 
discrimination of the two species is easy, though apparently the two are closely allied. 

(29) Plagithniysiis cristatits Sharp. 

Clytarlus cristatus Sharp, Tr. ent. Soc. London, 187S, p. 207. 

Plagithmysus cristatus Sharp, Ent. Mo. Mag. 1896, p. 274. 

(Plate VI. fig. 21 ; t-) 

Hab. Oahu ; Honolulu, Blackburn ; on Acacia koa, Perkins. 

In this species the hind femora are peculiar in shape, being much like what they 
are in Callithinystis. The legs are much more slender in the female than they are in 
the male. The specimen supposed to be Clytus atteniiatus Boisd. in the collection of 
the British Museum is, as Mr Gahan surmised, a female of this species. Boisduval's 
name has however no sufficient claim to validity, as will be seen from his description, 
repeated here on p. 115. 

Callithwvsus Sharp. 
Callithmysiis Sharp, Ent. Mo. Mag. x.xxii. 1896, p. 238. 

(i) Callithmysits nticrogaster Sharp. 

Clytarlus viicrogaster Sharp, Tr. ent. Soc. 1879, p. 103; Tr. Dublin Soc. (2), 

III. 1885, PI. V. fig. 46. 
Hab. Oahu ; near Honolulu, 2000 ft., in June, very rare ; Blackburn. 

Callitkmysjis microgaster var. ? Jiirtipes, var. nov. Tibiis posterioribus minus 
elongatis, usque basin densissime pubescentibus. 

Hab. Oahu : one individual on the ridge leading from the head of Pauoa Y^allev 
to the peak called Konahuanui, Oct. 31st, 1892. On Bobea sp. 

Subfam. LAMIINI. 

Group ACANTHOCINIDES. 

Lagocheirus Thorns. 

(i) Lagoclieints obsolctiis Thoms. 

Lagocheirus obsoletus Thoms, Class. Longic. i860, p. 10. 

Lagocheirus araneiformis var., Sharp, Tr. ent. Soc. London, 1878, p. 210. 

Hab. Oahu. Apparently confined, so far as this archipelago is concerned, to the 



114 FAUNA HAWAIIENSIS 

island of Oahu. This form is now considered to be a distinct species, though it is 
extremely close to the S. American insect with which it was previously identified — as a 
small form — by the late Mr Bates and myself. Gahan considers that L. longipennis 
Bates is a mere synonym of L. obsoletiis. The form is widely distributed, having been 
found in Loo Choo islands, Tahiti, the W. Indies, and Mexico. 



Group NIPHONIDES. 

Prosoplus Blanch. 

(i) Prosophts bankii Fabr. 

Lamia bankii Fabr., Syst. Ent. p. 176. 

Micracantha iiisii/aris Pascoe, Tr. ent. Soc. London (2), v. 1859, p. 40. 

Micracantha nutans Sharp, Tr. ent. Soc. 1878, p. 209. 

Hab. On the introduced Acacia, Prosopis sp. Probably on all the islands. 
Mr Perkins, knowing it not to be native, procured but few examples. The species is 
very widely diffused, having been found in Madagascar, Port Essington, Tondano, 
Amboyna. L. bankii was described as found at the Cape of Good Hope. The 
identification is due to an examination of the Banksian type in the collection of the 
British Museum made by Mr Gahan. The genera Prosoplus and Micracantha are 
considered by him to be one and the same. 

Group APOMECYNIDES. 

Apomecvna Serv. 

(i) Apomecyna pertigera Thorns. 

Mecynapus pertigera Thorns. Physis, i. 6, j^. 160. 

Hab. Oahu. In a garden, Honolulu, November 1896, Perkins. Kauai, July 
1897, Munro. The species comes from E. India and China. 

Group PTERICOPTIDES. 
Oopsrs Fairm. 

(i) Oopsis nit tat or Fabr. 

Lamia nutator Fabr., Mant. i. p. 142. 

Stasilca curvicornis Karsch, Berlin, ent. Zeitschr. xxv. p. 8, Plate I. fig. 12. 

Hab. Probably on all the islands. Abundant at low elevations. Widely 
distributed in Polynesia, and found in Australia. 



COLEOPTERA 115 



§ 3. Bibliographic List (arranged chronologically). 

1. BoiSDUVAL, — . Faune entomologique de I'Oceanie. Paris (no date, but 1835 '" preface), 705 pp. 

[P. 485. " C/yfus atfeiu/atus, Dejean. Fuscus ; thorace cristate, albido lineato ; elytris apice 
valde attenuatis, albido subvariegatis ; subtus ferrugineus. Nouvelle Hollande. Collec- 
tion de M. Dejean. Cet insecte n'est point un vrai Clyttis, il devra probablement former 
un nouveau genre." The specimen in Dejean's collection shows this to be Plagithmysus 
cristatus ? . D. S.] 

2. MoTSCHouLSKY, V. DE. Remarques sur la collection de Coleopteres russes de Victor de 

Motschoulsky. Bull. Soc. Moscou, xviii. 1845, P'- i- PP- 3 — 127, Plates I. — III. 

P. 85, description of Stenopierus piilverulenius, PI. I. fig. 2. Said to be from California. 

3. • Observations sur le Musee entomologique de I'Universite imperiale de Moscou. Op. cit. 

1845, Pt- II- PP- 332—388, Plates I., VII. 

Plagithmysus, n. g., p. 369, for the Stenopterus puh'eniknlits, p. 370, PI. VII. figg. 7 — 7, iv. 
Again said to be from California. 

"4. . Etudes entomologiques, 1852, p. 76. 

Erroneous synonymy of Plagithmysus. 

5. Sharp, D. On some Longicorn Coleoptera from the Hawaiian Islands. Tr. ent. Soc. London, 

1878, pp. 201 — 210. 

Nine species, 6 new (but 3 not really new). Three genera supposed new (i not really so). 

6. . On some Coleoptera from the Hawaiian Islands. Tr. ent. Soc. London, 1879, pp. 77 — 105. 

Two n. spp. of Clytarlus. Indicates, without naming, three genera. 

7. Harold, E. von. Einige neue Coleopteren. Mitth. Miinchen. ent. Ver. iv. 1880, pp. 148 — 171. 

Clytarlus finschi, n. sp., p. 166. 

8. Karsch, F. Zur Kaferfauna der Sandwich-, Marshall- und Gilberts-Inseln. Berlin, ent. Zeitschr. 

XXV. 1881, pp. I — 14, PI. I. 

Aegosoma reflexum, Stasilea curvicornis = Oopsis nutator, Clytarlus pulvillatus, n. spp., with 
figg. ; C. finschi, figure. 

9. Sharp, D. On some new Coleoptera from tlie Hawaiian Islands. Tr. ent. Soc. London, 1881, 

PP- 507—534- 

Clytarlus pennatus, fragilis, n. spp. 

10. Blackburn, T. and Sharp, D. Memoirs on the Coleoptera of the Hawaiian Islands. Tr. Dublin 
Soc. 2nd ser. in. 1885, pp. 119 — 290, Plates IV. and V. 

General work. Clytarlus blackburni and filipes, spp. n. 

F. H. II. 16 



ii6 FAUNA HAWAIIENSIS 

11. Sharp, D. Note on the genus Plagithmysus Motsch. C.R. Soc. ent. Belgique, 1885, pp. 

Ixxiv — Ixxvi. 

Synonymy and habitat of PIag;ithmysus. 

12. . On Plagithtnysus : a Hawaiian genus of Longicorn Coleoptera. Ent. Mag. xxxii. 1896, 

pp. 237—245, 271—274. 

Twenty-two spp., 18 new. Rearrangement of genera. Callithmysns, n. g. 

13. . Supplement. Op. cit. xxxiii. 1897, p. 12. 

P. albertisi, n. sp. 

14. Perkins, R. C. L. A collecting trip on Haleakala, Maui, Sandwich Islands. Ent. Mag. xxxii. 

1896, pp. 190—195. 

Stridulation and habits of Plagithmysus (as Clytarhis). 

15. . Notes on some Hawaiian Insects. P. Cambridge Phil. Soc. ix. 1897, pp. 373 — 380. 

Habits and variation of Plagithmysides. 



I 



117 



o, f- 5^'^ 






II. COLEOPTERA RHYNCHOPHORA, PROTERHINIDAE, 
HETEROMERA AND CIOIDAE. 

By R. C. L. Perkins. 

COLEOPTERA RHYNCHOPHORA. 

Earn. CURCULIONIDAE. 

§ 1. General Remarks on the Curculionidae \ 

The Curculionidae are represented by 137 species, and of these no less than 132 
are at present only known from the islands. It is probable however that of the others 
some will be found to be not truly indigenous, so that we will first briefly review the 
various tribes with respect to these doubtfully indigenous forms. Of the Otiorhynchini 
the species oi Rhyncogomis are all peculiar to the islands, and the only one to which any 
suspicion can attach is R. vcstitiis, which in many respects is very different to, and indeed 
less remarkable than the others. It is the only species which frequents the low lands, 
and is found upon plants which are known elsewhere, whereas the others are all attached 
to the forest trees or plants which are themselves loeculiar to the islands. Still on the 
whole we are disposed to consider it indigenous. The solitary species of Pantomorus 
on the other hand is certainly an introduced species, as we have found it in gardens in 
Honolulu, and on Maui it has increased to such an extent as at times to be injurious. 
The Cyladini are represented only by one species of Cylas known to occur elsewhere, 
and which is attached to a common coast species oi Ipomoea. No doubt this beetle is a 
natural immigrant. 

The Cryptorhynchini have 20 species of Acalles, certainly all indigenous, and two 
other species representing each a different genus. Both appear to be allied to Acalles, 
the one indeed closely allied, and there is no reason to doubt that both are indigenous. 

The Calandrini have three species of Calandra, two being widely distributed 
outside the islands. The third is a remarkable species and is attached to cactus and 
banana, and although not known elsewhere it may well have been imported, and cannot 
safely be considered indigenous, nor even a natural immigrant. Besides these a single 
species of Sphenophoriis, the well-known cane-borer of the sugar-planters, is known 
elsewhere, and is either an importation by man or a natural immigrant. 

' For remarks on the Proterhinidae and other groups see subsequent pages. 

16 — 2 



ii8 FAUNA HAWAIIENSIS 

The Cossonini are the most extensive and important representatives of the Curcu- 
Honidae. The 46 species of Oodemas and the 10 of Heteramplnis may be at once 
dismissed as being all indigenous, and the same may be said of the three allied species 
of Anotheoriis, and the same number of Nesotociis. The 17 species of Dryophthor^is 
are a varied assemblage and might be distributed in two or three genera, but there is 
no reason to suppose that any of them will be found elsewhere. The unique example 
which represents the genus Orothreptes and the hardly less rare species for which we 
have made the genus Deinocossoniis are almost certainly indigenous, for they are moun- 
tain forms, attached to indigenous trees. The latter indeed appears to have already 
formed to some extent distinct races on several of the islands. The species for which 
the genus Dysomjna is formed must also be considered indigenous, the unique example 
having been found by sifting dead leaves on the boggy summit of the Kauai mountains. 
Other two genera now described, each for a single species — Thallatodora and Ha- 
loxenus — are both found on the coast in logs of driftwood. It is probable that both 
will be found elsewhere, being natural immigrants. The two species of Pseudolus must 
both be excluded from the indigenous weevils ; one is found freely in banana and cactus 
in company with Calandra remota, the other has been found in and near Honolulu only, 
and on one of these occasions in boards of foreign wood, which formed the floor of a 
room. The Phloeophagoso7na is abundant in decaying wood oi Aleurites at low eleva- 
tions, in company with Pentarthrum obsairum, and both these species I have little 
doubt occur outside the islands. It may be noted that the Aleurites or candle-nut-tree 
produces but very few indigenous insects, and such as frequent it are mostly poly- 
phagous species, and although the tree now forms in many localities a large part of the 
lower forest, I suspect that it has been comparatively but a short time in the islands, 
and was very probably brought there by the natives when they settled in the country. 
Another species of Pentarthrum. found on the lowlands near Honolulu must also be 
considered as foreign. The third species of this genus is very different in appearance 
and structure to the others, and is truly indigenous, being confined to the mountain 
forests, where it is attached solely to the tree-ferns, which are so conspicuous a feature 
of these forests. Thus the species which in my opinion are foreign amount to 13, and 
are distributed in no less than nine genera, and it may be remarked that those 
Cossonini, which we consider as foreign, although they have not yet been obtained 
elsewhere, are all small and obscure insects such as have been little collected in most 
countries. 

The indigenous Curculionidae on the other hand with their 123 species are distri- 
buted in 13 genera, which represent three tribes only — Otiorhynchini, Cryptorhynchini, 
and Cossonini. The genus Rhyncogonus, which was established by Sharp for the 
members of the first tribe, is not peculiar to the islands, and has since been found in 
New Zealand and other of the Pacific islands {vide Ent. Mo. Mag. 1899, p. 56). 

The Cryptorhynchini have two genera peculiar, each with only one species 



COLEOPTERA 



119 



known, but the great majority of the species belong to the widely distributed genus 
AcalL's. 

or the Cossonini, Oodemas, Heteramphus, Anotheoriis, Nesotocus, Dysomnia, 
Deinocasoims, and Orothreptes are all genera peculiar to the islands, and it is probable 
that some members of the genus Dryoplithoriis will at some time be separated as 
distinct, while the rest will remain in that or be placed in very closely allied genera. 
The indigenous Pentarthriim is a remarkable insect, very different to the two foreign 
exponent > of the genus, but said to be allied to New Zealand forms (Sharp, Tr. ent. 
Soc. Lond. 1878, p. 25). Thus nine of the 13 genera of indigenous Curculionidae are 
peculiar to the islands, while Rhyncogonus with its numerous species probably originated 
in the Pacific region and will be found to be widely distributed therein. With regard 
to Acalles Sharp has remarked (Tr. Dublin Soc. 1885, p. 269) that it "is one of the 
most frequent components of insular faunae, being very richly represented in New 
Zealand, Polynesia, and the Atlantic islands." 

Distribution of genera and species in the islands. Of the six genera which 
contain each but a single species, the two Cryptorhynchids, H ypei'omorpha and Chae- 
nostermmi, have both been found on Oahu alone, and the Cossonideous genera Dysomma 
and Orothreptes are peculiar, the first-named to Kauai, the latter to Hawaii, whereas 
Deinocossonus is probably to be found on all, since it has been taken on Kauai and 
Hawaii at either end of the group as well as on the intermediate island of Oahu, and 
the indigenous Pentarthruvi is likewise ubiquitous. It must however be remembered 
that all the above-named insects, excluding the Pentarthriun, are excessively rare and 
difficult to procure, so that more extended observations might show that those apparently 
peculiar to one island also occur on some of the others. Nesotocus has been found 
alive on Kauai, Maui and Hawaii, and fragments also on Molokai, so that it is no 
doubt of general distribution, but Anotheorus is wanting on Hawaii, its range being 
from Kauai to Maui. The distribution of Heteraniphus is similar to that of 
Anotheorus, except that whereas we failed to collect the latter on Molokai, so Heter- 
aniphus was not found on Lanai, but both are found on the neighbouring island of 
Maui. The extensive genera Oodemas and DryophtJiorus are ubiquitous over the 
group, as also is Acalles, but Rhyncogonus apparently becomes scarcer on the windward 
islands, and is altogether absent from Hawaii. 

The species of the genus Rhyncogonus, e.xcepting only R. sordidus, which was 
iound alive on Lanai and in fragmentary condition on Molokai, are confined each one 
to a single island, and of the 19 species Kauai has no less than nine, Oahu four, 
Molokai three, Lanai and Maui two each, whilst Hawaii, as has been remarked, has 
none. 

Of the 20 species of Acalles the three islands, Kauai, Oahu and Hawaii have each 
four species, and Maui and Lanai three each, while one species is found throughout the 



I20 FAUNA HAWAIIENSIS 

group, and another has occurred on three of the intermediate islands. These fif,ures 
are however subject to correction, for the Hawaiian species oi Acalles are excessively 
difficult to collect, and some of them exhibit considerable variability, so that it is neither 
certain that all the species described are certainly distinct, nor that the forrrs from 
different islands assigned to a single species are really identical. The species of 
Dryophtho7'us appear to be less restricted in their distribution ; Kauai has s'x, Oahu 
two, and Hawaii one peculiar species, but the remaining eight are found o i two or 
more of the islands ; three indeed are found throughout the group. 

The extensive genus Oodemas has no less than 46 species, and is most richly 
represented on Kauai, which has 18 species peculiar to it. Oahu and Maui each have 
just half as many peculiar, whilst Hawaii, Lanai and Molokai have respectively three, 
two and one. The remaining four species occur on two or more of the islands, but 
neither of them is found on Kauai. 

Heteramplms would appear to have its head-quarters on Oahu, where are five of 
the ten known species, and these five are all peculiar to it ; Maui has three species 
peculiar, and Molokai and Kauai each have one ; that on the latter island would have to 
be separated generically were it not connected by the Oahuan H. hirtellns with the 
more usual forms. Three of the Oahuan species are usually found in company on 
Astelia veratroides, but would appear to be of extremely restricted range within that 
island, since they are not to be found in many places where the plant is extremely 
abundant, nor have we found any species attached to the Astelia on the other islands, 
the Molokai form having been obtained from wet moss, and the others from wet 
decaying logs. No species of this genus has been found on the large island of Hawaii. 

Anotkeorus has one species quite peculiar to Kauai, Oahu likewise has one, but it 
is very closely allied to the Maui form, and as both vary, I have some doubt as to their 
being distinct. A single example from Lanai agrees best with the Maui species. 

The extraordinary genus Nesotocus has one species peculiar to Kauai and one to 
the Western range of mountains on Maui, the third has been taken both on Haleakala, 
the Eastern division of this island, and also on Hawaii. 

The remaining indigenous species have been sufficiently referred to above under 
the genera to which they belong. 

The percentage of the indigenous species of Curculionidae peculiar to the several 
islands is given in the table. 





Species 


peculiar 


to. 


Total 


number of ; 
found upon. 


species 


Percentage of species 
peculiar to. 


Kauai 




41 






48 




85 


Oahu 




27 






39 




69 


Molokai 




4 






16 




25 


Maui 




18 






31 




58 


Lanai 




6 






19 




31 


Hawaii 




9 






20 




45 



COLEOPTERA 



12 I 



These figures fully show the remarkable character of the Kauai fauna in this family 
of Coleoptera. Not only is it considerably the richest in species, but the percentage of 
peculiar species is much higher than is the case with any other island. Moreover the 
species themselves are in many cases highly remarkable, as compared with those of the 
same genus on the other islands. Thus in Oodewas, O. longirostre, O. costatnm, 
O. striahan, O. pulchrum, are species of the most highly evoluted form, each in its own 
particular direction. So also the Kauai species of Rhyncogonus form a remarkable 
series. These facts suggest the probability that the ancestors of the now existing fauna 
of Curculionidae may, at least to a large extent, have first gained a footing on this 
island, possibly at a time when some of the more recent islands were not adapted for the 
mode of life of these creatures, or possibly even before these islands existed at all. This 
suggestion is strengthened by a consideration of the condition of the weevils on the 
other islands ; thus Oahu comes second both in number of species and in the per- 
centage of those peculiar to it, and it is noteworthy that a number of the most 
remarkable forms are found in what yet remains of the once fine forest of the Waianae 
range, itself one of the oldest portions of the whole group of islands. Very different is 
the case with Hawaii, which from its large size, lofty mountains, and very varied climate, 
as well as its position at the end of the series of islands, might be expected to have a 
large number of remarkable forms. This however is not the case, for it is not only 
altogether wanting in Rhyncogonus, Hetcramphus and Anotheot'us, but it possesses but 
three or four species of Oodenias, and these of the most commonplace forms. In the 
group we are now considering the condition of the fauna of the big island contrasts 
curiously with its Aculeate Hymenoptera, which are both richly represented and 
remarkable ; indeed it is not necessary to go beyond the order of Coleoptera to see a 
similar contrast, for the indigenous Cerambycidae are well represented there and some 
of the forms are amongst the most remarkable of those found in the group. 

Variation. We have already when dealing with the Aculeate Hymenoptera, 
Neuroptera and Orthoptera remarked on the variability of so many of the island species. 
Not less remarkable is this in the case of the Curculionidae. Mr Blackburn in describing 
the species of Oodenias notes that their variability in certain points is embarrassing to 
the describer, and with a large collection this becomes so much so, that in many cases 
it is almost impossible to write a description which will apply in all respects to every 
individual specimen. This variability too is not confined to superficial and trivial 
characters, but affects those which are considered of decided importance for the separa- 
tion of species. Even structures which are almost always constant vary in some species, 
as for example parts of the prosternum in some species of Oodenias. We may observe 
that there is no doubt as to the examples which exhibit this variability belonging to the 
same species, and we are excluding from consideration such a species as O. niultifornie 
of Hawaii, which may possibly include several allied and variable species; but is more 



122 FAUNA HAWAIIENSIS 

probably one which is now in the process of becoming divided into several, the division 
not yet being complete. Some of the species of Rhyncogonus too are variable, as also 
are those belonging to the genera Dryophthorus and Heteramplms, and there is no 
doubt that many of the species of Acalles will prove to be equally so, when they have 
been collected in numbers sufficient for an investigation on this point. 

The Curculionidae of the islands are as a whole remarkably isolated. The RJiynco- 
goni of the Otiorhynchini and the solitary indigenous species of Peiitarthruiu and 
Orothreptes all probably originated in the Pacific islands, for the latter appears to be 
most allied to the New Zealand Sericoirogus, although this alliance is by no means 
close. The very important Cossonideous genera Oodemas, Anotheorus, Heteramplius 
and Nesoiocus, on the other hand, are of quite uncertain origin, no near ally to any of 
these forms having been found elsewhere. Indeed great as are the differences between 
the first three of these genera it would appear that they are at least as closely connected 
inter se as they are with anything outside the islands. Thus some of the species of 
Oodemas, entirely different as they are in facies, do make some approach to Heteramplius, 
while Anotheorus and Oodemas appear to be more nearly allied. 

Wollaston remarks that metallic colouring is of rare occurrence in the Cossonini, 
so that it is worthy of note that no less than 49 species, in fact all the members of the 
genus Oodemas and Anotheorus, are so coloured. The extreme similarity in colour and 
clothing between the very diverse genera Orothreptes and Deinocossomcs, is quite extra- 
ordinary, both being clothed with a golden pubescence, in which they greatly resemble 
the New Zealand Sericotrogiis, although Deinocossomis does not even belong to the 
same section of the Cossonini as the other two. 



§ 2. Systematic account of the Curculionidae. 

Tribe OTIORHYNCHINI. 

Rhyncogonus Sharp. 

(i) Rhyncogonus nitidus, sp. nov. 

Niger, nitidus, supra fere glaber. Caput cum pronoto laeve, et distincte punctatum. 
Elytra nitida, seriatim punctata. Abdominis segmentum ventrale apicale d'-is latum, 
opacum, pubescens, densissime punctatum, apice truncate, $-ae nitidum, parcius pubes- 
cens, apice angusto, subacuto. Long. 12 — 17 mm. (Plate VII. fig. i.) 

Black, smooth, and shining, often with a small spot of pale pubescence on each 
side of the thorax at the base, the % generally larger and more robust than the ^. 



COLEOPTERA 123 

Head and thorax distinctly and rather finely punctured, the punctures of unequal size ; 
second joint of the antennae rather longer than the third. Elytra shining, each with 
about 1 2 rows of punctures dorsally, more strongly attenuate towards the apex in the % 
than in the ^, and in the former sex with the edges flattened for some distance from the 
shoulders. Abdomen beneath coarsely punctured at the base, and widely depressed in 
the ^, finely punctured and but little depressed in the ?. Apical ventral segment of the 
t dull, finely and densely punctured, pubescent but not very densely so, the apex very 
broad and subtruncate. In the % this segment is shining, clothed with inconspicuous, 
and very short pubescence, subtriangular and narrowly rounded at the apex. 

Hab. Kauai Mountains (3000 ft.). 



(2) Rhyncogonus funereus, sp. nov. 

Niger, subnitidus, supra haud pubescens. Caput strigosum et punctatum. Pro- 
thorax dense rugoso-punctatus. Elytra grosse punctata. Antennarum articulus secundus 
tertio multo brevior. $. Long. 12 "5 mm. 

Black, with the surface shining, and coarse sculpture. Head strongly strigose, and 
punctured. Second joint of the antennae much shorter than the third. Thorax strongly 
and densely rugosely punctured, its sides hardly rounded, subparallel. Elytra shining, 
subdepressed, without pubescence, coarsely punctured, the interstices narrow, irregular. 
Posterior femora with a transverse spot of pale pubescence outwardly about the middle ; 
tibiae, tarsi and apex of the rostrum with fine pubescence. 

Hab. Oahu, Waianae mountains. A single $, found dead and somewhat muti- 
lated, beneath bark. 



(3) Rhyncogonus sqtiamiger, sp. nov. 

Niger, nitidus, antennis, tibiis, tarsisque rufescentibus. Caput cum prothorace 
tenuiter pubescens. Elytra squamulis pallidis, plus minusve seriatim dispositis, vestita. 
Antennarum articulus secundus tertio multo longior. $. Long. 14 mm. (Plate VH. 
fig. 2.) 

Black and shining, the antennae and all the tibiae and tarsi of an obscure reddish 
colour. Head with a thin clothing of short pale hairs which become scale-like, and form 
a pale mark around the inner and posterior margins of the eyes, rugosely punctured. 
Antennae with the first joint of the funiculus very long, considerably longer than the 
second, the club unusually long and slender, its terminal evidently longer than its basal 
joint. Thorax with sparse pubescence above, but forming a rather denser line along 
each side, as wide as long, its sides rather strongly rounded, the surface shining and 

F. H. II. 17 



124 FAUNA HAWAIIENSIS 

closely punctured. Elytra with regular rows of very closely-set punctures, the interstices 
narrow, and very smooth and shining ; along the striae there are rows of minute pale 
scales for the most part placed singly, but in places, especially at the shoulders, and 
towards the apex of the elytra, forming more or less dense patches. 

Hab. Kauai Mountains near Lihue (3000 ft.) ; i $ taken. 



(4) Rkyncogoniis stygms, sp. nov. 

Niger, capite strigoso et punctato. Antennarum articulus secundus et tertius 
subaequilongi. Elytra opaca, pube inconspicua et brevissima vestita, seriatim punctata, 
interstitiis tuberculis minutissimis, rare dispositis, mvmitis. Long. 12 "5 — 15 mm. 
(Plate VII. fig. 3.) 

Black, the antennae, tibiae, and tarsi sometimes with a reddish tinge. The head is 
strigose and punctured, the antennae have the second and third joints nearly equal in 
length, the apical joint of the club is evidently shorter than its basal one. The thorax 
is densely punctured, and has a small pale pubescent spot on each side at the base. 
The elytra are quite dull, each with about a dozen regular rows of punctures dorsally, 
and clothed with an excessively short pubescence, usually hardly visible without close 
examination, but in some examples sufficient to give the surface a slightly greyish 
appearance. The interstices are furnished with minute scattered tubercles, which are 
slightly less dull than the general surface. The intermediate and posterior femora have 
a distinct patch of pale pubescence outwardly. 

Hab. Kauai, Halemanu (4000 ft.). 



(5) Rhyncogonus viinor, sp. nov. 

Niger, parum nitidus, capite pubescentia pallida parcius vestito. Antennarum 
articulus secundus tertio vix longior. Prothorax utrinque linea pubescentiae pallidae 
continua ornatus. Elytra, humeris et parte apicali exceptis, baud pubescentia. ?. 
Long. io'5 mm. 

In general sculpture this species bears a strong resemblance to the preceding, but it 
is a much smaller insect and it is comparatively less elongate. The pubescence of the 
elytra is confined to some scanty patches at the extreme base, and a very sparse clothing 
on the apical portion. Their surface is evidently less dull, and except towards the 
sides and apex there are no evident minute tubercles on the interstices. Their form 
is different, for they are much more suddenly narrowed to the apex, the narrowing 
beginning only about one-third of their length from the apex. The front of the head 
is evidently more pubescent, and there is a wide and distinct stripe of pubescence on 



COLEOPTERA 125 

each side of the thorax from base to apex. In the $ (the only sex obtained), the dorsal 
edges of the elytra are very evidently flattened for a considerable distance from the 
base. The puncturation of the whole insect, allowing for the difference in size, does 
not materially differ from that of R. stygins. 

Hab. Kauai Mountains (4000 ft.); i % taken. 



(6) Rhyncogonus molokaiensis, sp. nov. 

Niger, subnitidus, antennis pedibusque saepe rufescentibus. Caput circa basim 
dense subtilius punctatum. Antennarum articulus tertius secundo vix brevior. Pro- 
thorax dense (nee grosse) punctatus, macula utrinque ad basim pubescente ornatus. 
Elytra subnitida, fere glabra. Long. 9*5 — 12 mm. 

Black, and somewhat shining, the legs and antennae more or less of an obscure 
red colour. The head is closely and finely punctured at the base, the second joint of 
the antennae subequal to, or rather longer than, the third. Thorax densely, but not 
coarsely, punctured ; the punctures, however, are much larger and stronger than those 
at the base of the head. There is a very distinct smooth median dorsal line extending 
from base to apex, and a small spot of pale pubescence on each side at the base. The 
elytra are not very dull, and are almost without pubescence, and the surface is slightly 
roughened by delicate corrugations proceeding from the punctures. In both sexes the 
two apical segments of the abdomen beneath are clothed with pubescence, which is 
very dense on the apical one. The other segments have only very sparse and minute 
hairs. In the ^ the apical segment is broad and truncate, in the $ narrow and some- 
what pointed at the apex, as is usual throughout the genus. 

Hab. Molokai, mountains (3000 ft.) ; rare. 



(7) Rhyncogon7is ditbins, sp. nov. 

Niger, vix nitidus, elytris vix evidenter pubescentibus, Z praecedenti cognatissimus, 
capite circa basim baud dense punctato, elytris magis corrugatis, distinguendus. ^. 
Long, fere 10 mm. 

I have seen only a single Z of this species, which is closely allied to that of 
R, violokaiensis, but apparently distinct. The front of the head (including the rostrum) 
is by no means densely punctured, and at the base the punctures are larger and much 
more sparing. The smooth dorsal line on the thorax is much more definite, and the 
surface of the elytra less smooth, owing to the stronger corrugations. 

Hab. Molokai, Kalae (1700 ft.); i t taken. 



126 FAUNA HAWAIIENSIS 



(8) Rhyncogonus freycinetiae, sp. nov. 

Niger, subnitidus, elytris vix pubescentibus, capite circa basim minus dense 
punctato. Prothorax latus, trans medium evidenter latior quam longior, lateribus linea 
pubescentiae continua ornatis. Elytra subnitida, interstitiis laevioribus. Abdominis 
segmentum tertium ventrale $ aeque ac quartum pubescens. $. Long. 1 1 mm. 
(Plate VII. fig. 4.) 

Female black, and somewhat shining, in general appearance very like that of R. 
molokaiensis. From that species it may easily be distinguished by the less close 
puncturation of the base of the head ; the much wider and more depressed thorax, with 
continuous lateral lines of pubescence. The ventral surface of the hind body is also 
more pubescent, the third segment hardly less clothed than the fourth. 

Hab. Oahu, Halemano (2000 ft.) ; i % taken at the bases of the leaves of 
Freycmetia. 

(9) Rhyncogonus blackburni Sharp. 

Rhyncogonus blackburni Sharp, Tr. Dublin Soc. in. (Ser. 11.) p. 177. 

Hab. Oahu, mountains near Honolulu (Blackburn) ; i ^ in the same locality 
(2000 ft.). 



(10) Rhyncogonus koebe/ei, sp. nov. 

Niger, baud nitidus, antennis pedibusque rufescentibus. Caput sublaeve, sparsim 
punctatum. Prothorax subtilissime pubescens, densissime rugoso-punctatus, lateribus 
fortiter rotundatis. Elytra peropaca, parcissime pubescentia, apicibus liberis. $, Long. 
II mm. (Plate VII. fig. 5.) 

Dull blackish, the antennae and all the legs dull reddish. Head smooth and 
shining, sparsely punctured, the eyes very little prominent. Second joint of the 
antennae rather longer than the third, the basal joint of the club longer than the apical. 
Thorax very finely pubescent, densely rugosely punctured, its sides strongly rounded. 
Elytra dull, with very little pubescence, the puncturation somewhat irregular, their 
apices free, subdivergent. All the femora clothed with very short pubescence, that on 
the tibiae longer. Apical ventral segment of the hind-body widely truncate at the 
apex, densely clothed with pale pubescence. The species is very remarkable for the 
condition of the eyes, which are much less prominent than in any other. 

Hab. Oahu, mountains near Honolulu (2000 ft.) ; i $ taken by Mr A. Koebele, 
whilst collecting with me. 



COLEOPTERA 127 

(11) Rhyncogomis kaiiaiensis, sp. nov. 

Niger, vel plus minusve rufescens, capita et corpore toto subtiliter griseo- vel rufo- 
pubescente. Caput strigosum et punctatum. Prothorax dense punctatus, utrinque vitta 
pubescente ornatus. Elytra vix opaca, subtiliter pubescentia. Long. 10 — 13 mm. 

Colour varying from black to reddish brown. The male generally much depressed, 
the % more convex, but neither sex is constant in this respect. Head more or less 
strigose and punctured, clothed all over with fine pale hairs, which form a denser line 
around the inner margins of the eyes. Thorax densely punctured, clothed with fine 
pubescence like that of the head, and on each side with a line of dense pale pubescence. 
Elytra usually more or less shining, but much duller in some examples than others, 
clothed with fine grey or reddish pubescence, the series of punctures for the most part 
distinct, but towards the sides becoming more or less confused, and the surface more or 
less rough, often tuberculate. The pseudepipleural region of the elytra always with 
conspicuous patches or spots of dense pubescence. 

Hab. Kauai (4000 ft.) ; 15 examples taken. Apparently a very variable species. 

(12) Rhyncogomis sordidus, sp. nov. 

Niger, vel subrufescens, opacus, subtiliter pubescens, capite punctate. Prothorax 
dense nee grosse punctatus. Elytra opaca, pubescentia, seriatim punctata, lateribus 
maculis pubescentibus haud ornatis. $%■ Long. 10 — 12 mm. 

The colour of this species varies from black to obscure red. The front of the head 
and rostrum is almost without pubescence, the puncturation of the latter is usually not 
very close and it shows little or no trace of longitudinal strigosity, its surface is more or 
less flattened or depressed. The thorax is densely, but not coarsely, punctured, more 
or less impressed near the base, clothed with very short and fine pubescence, and with 
a line of denser and rather longer hairs on each side, but even in fresh examples these 
lines are obscure. The elytra have a very fine pubescence, which is more conspicuous 
in the ?, and of a reddish colour. Their surface is dull, but not rough, and the 
punctures towards the sides remain quite distinct, although the series may be more or 
less confused. On their deflexed portion there is no trace of spots or lines of dense 
pubescence, such as are found in the preceding species. In the ^ the apical ventral 
segment only of the hind-body is densely hairy, the others but sparsely so, the two 
basal more or less strongly impressed. 

Hab. Lanai, mountains (3000 ft.). A short and rather variable series taken. 
The relative length of the second and third joints of the antennae appears to vary, as 
also the puncturation of the insect. Fragments of a Rhyncogomis found on Molokai 
are probably referable to this species. In one spot these were very abundant, but the 
species had no doubt been extinct (so far as that particular locality is concerned) for 
years, the vegetation consisting only of small stunted bushes. 



128 FAUNA HAWAIIENSIS 



(13) Rhyncogonus lanaiensis, sp. no v. 

Forma, facieque praecedentis ; $ oculis fortiter prominentibus, abdominis segmentis 
ventralibus tribus apicalibus dense pubescentibus distinguendus. $. Long. 1 1 '5 mm. 
($ ? 14 mm.) 

Extremely like the preceding, but distinct by the strongly prominent eyes. The $ 
also differs in having the three apical ventral segments of the hind-body densely 
pubescent, and the head somewhat strigose, as well as punctured. The first joint of 
the funiculus of the antennae is considerably longer than the second. 

A single $ probably belongs to this species. It differs from the preceding in its 
larger size, more prominent eyes, and the rather more pubescent ventral segments of 
the hind-body. 

Hab. Lanai (2000 — 3000 ft.) ; i $ and i ?. 



(14) Rhyncogonus laliainae, sp. nov. 

Nigricans, pubescentia pallida vestitus, pedibus antennisque rufescentibus ; ^ 
thorace densissime punctato, interstitiis nitidis. Elytra pubescentia, sensim ad apices 
attenuata. ^. Long. io"5 mm.. 

Blackish, the antennae, tibiae, tarsi, and more or less of the femora, red. The 
surface of the rostrum iyZ) is somewhat smooth, punctured, but not strigose, and with a 
scanty short pubescence. The antennae have the first joint of the funiculus evidently 
longer than the second. The sides of the prothorax are rounded, its surface smooth 
and shining between the dense punctures, and clothed with short fine pubescence, which 
at the sides becomes longer and coarser, and forms a somewhat indefinite lateral band 
on each side. The elytra are dull, and on each there are five or six lines of pale 
pubescence, which are not very distinct, and due to the fact that the interstices are 
alternately more densely or more sparsely clothed. The puncturation is distinct 
throughout, but the second and third series originate at some distance from the base of 
the elytra. 

Hab. Maui, mountains behind Lahaina (3000 ft.) ; \ ^ taken. 



(15) Rhyncogonus depressus, sp. nov. 

Niger, densius pallido-pubescens ; Z fortiter depressus. Rostrum pubescentia 
tectum, strigoso-punctatum. Pronotum cum elytris pubescens, nitidum ; elytra seriatim 
punctata, tuberculisque minutis asperula. Long. io"5^ — 13"5 mm. (Plate VII. fig. 6.) 



COLEOPTERA 129 

Black, the whole insect clothed with somewhat dense pale pubescence, which is 
rather in the form of very fine elongate scales, and to a large extent conceals the 
surface, which is shining. The head in front is strigose and punctured ; the prothorax 
densely punctured, the sides not at all strongly rounded, the surface shining, and with a 
smooth narrow dorsal line, the sides more densely pubescent than the general surface. 
The elytra in the Z are much depressed, with closely-set series of punctures, the surface 
where not hidden by the pubescence, shining, and roughened with small tubercles. All 
the legs clothed with pubescence, that on the front tibiae, as well as that of the scape of 
the antennae, longer and more conspicuous than that of most of the species. The single 
% which probably belongs to this species is much less depressed, and has a shorter club 
to the antennae, its middle joint being not longer than wide, and the joints themselves 
less distinct. 

Hab. Kauai, Halemanu (4000 ft.) ; 3 t and i %. 

{16) Rhyncogoims vittatiis, sp. nov. 

Niger, antennis pedibusque saepe plus minusve rufescentibus. Caput cum pronoto 
squamulis pallidis vestitum. Elytra tuberculis nitidis asperula, fasciis longitudinalibus 
pallidis ornata. ^ %. Long. 10 — 11 mm. (Plate VI. fig. 7.) 

Black, the antennae and legs often more or less red or piceous. The J is very 
strongly flattened or depressed, the % sometimes of more convex form. The head 
is strigose and punctured, but the surface is for the most part concealed by the covering 
of scales and hairs. The sides of the prothorax are but little rounded, and slightly 
convergent in front, the surface densely punctured and concealed by the pale squamous 
pubescence. The elytra have each about six distinct longitudinal bands, formed of 
minute pale scales, the spaces between these bands being sparsely clothed with similar 
scales. The surface is roughened by shining tubercles, and the puncturation much 
obscured. All the legs and the antennae are clothed with pale hairs. In both sexes 
the entire ventral surface of the hind-body is conspicuously pubescent, but the basal 
segments are less densely clothed than the apical. 

Hab. Kauai, mountains (3000 ft.) ; not common. 



(17) Rliyncogomis tuber ailatus, sp. nov. 

Minor, haud nitidus, totus pubescens, nigro-piceus, antennis pedibusque rufescenti- 
bus. Caput strigosum. Elytra tota fusco-pubescentia, seriatim punctata, tuberculis 
rare dispositis munita, apicibus haud liberis. ^ %. Long. 8*5 — 9 mm. 

A small species, entirely clothed with fuscous pubescence, the ^ strongly depressed. 
Head strigose ; antennae with the first joint of the funiculus subequal to the second ; in 



I30 FAUNA HAWAIIENSIS 

the Z these joints are unusually short, and either of them is only about the length of the 
basal joint of the club. The sculpture of the prothorax is largely concealed by the 
pubescence, its surface is roughened by minute tubercles, and the puncturation is 
indistinct. The elytra are seriately punctured, and entirely covered with pubescence, 
which shows no tendency to form lines, and are united to the extreme apex. There are 
very distinct tubercles sparsely scattered over the surface, many of which terminate in a 
single short bristle. The apical ventral segment of the hind-body in the ^ is of a 
reddish colour, densely pubescent, and truncate at the apex. The rest of the ventral 
surface is conspicuously, but less densely, pubescent. 

Hab. Kauai, Halemanu (4000 ft.) ; x ^ i $ taken. 



(18) Rhyncogonns sylvicola, sp. nov. 

Minor, niger, vix nitidus, pubescens, prothorace subelongato, rugoso-punctato. 
Elytra ad basim prothorace latiora, tuberculis rare dispositis munita, apicibus breviter 
liberis. $. Long. 9 — 9-5 mm. 

Allied to the preceding, but blacker in colour, and the surface less dull. The 
prothorax is evidently longer, and its puncturation, which is dense and rugose, is 
distinct. The elytra have scattered tubercles much as in that species, but they are 
evidently wider at the base, so that the humeral angles stand out somewhat from 
the hind angles of the prothorax, and their extreme apices are free. 

Hab. Kauai, Halemanu (4000 ft.) ; 2 % taken. 

(19) Rhyncogonus vestitus Sharp. 

Rhyncogomis vestitus Sharp, Tr. Dublin See. iii. 1885, p. 177. 

(Plate VH. fig. 8.) 

Hab. Maui. Very abundant on the sandhills between the mountains of East and 
West Maui, but not found elsewhere. 



Pantomorus Schonherr. 

(1) Pantomorus olindae, sp. nov. 

Nigricans, squamis pallidis vestitus. Capitis pars praeantennalis longitudinaliter 
impressum, post hanc linea impressa mediali sat distincta. Antennarum articulus 
secundus tertio multo brevior. Oculi fortiter convexi. Pronotum pallidis squamis 
celatum, lateribus parum rotundatis. Elytra substriata, striis confertim punctatis. 



COLEOPTERA 



J31 



squamis pallidis necnon postice setulis pallidis brevissimis vestita, lateribus fusco- 
squamosis. Abdominis segmenta ventralia subtilissime rugulosa, vix evidenter punctata. 
Long, (cum rostro) 8- — 8*5 mm. 

This species is found in Honolulu, and is sometimes very abundant at Makawao 
and Olinda, Maui, and is found as high up as 5000 ft. on Haleakala. It differs from 
Pantomorus proper in having the antennal scrobes less deep and less definitely marked 
posteriorly. 

Hab. Oahu and IVIaui ; no doubt an imported species. 



Tribe CYLADINl. 

Cylas Latr. 

(i) Cylas turcipennis Boh. 



Cylas turcipetmis Boh. Sch. Gen. Cure. i. p. 369. 
Hab. Oahu and Maui ; coast. 



Tribe OR YP TORN YNCHINI. 

AcALLES Schonherr. 
(i) Acalles lateralis Sharp. 

Acalles lateralis Sharp, Tr. Dublin Soc. iii. 1885, p. 178. 

(Plate VH. fig. 9, t.) 

Hab. Oahu etc. The typical specimens are from Oahu ; I have taken others 
which must be referred to this species, on Kauai, Molokai, Maui, and Hawaii. Very 
scarce. The insects of this genus are with one or two exceptions extremely difficult to 
procure, being generally taken singly, and in very different conditions as regards the 
covering of scales ; this, added to the fact that they are certainly variable in colour, 
size etc., makes a comparison of allied forms very difficult. It is quite possible that 
there are several species very closely allied to A. lateralis, the number and condition 
of the examples taken making it impossible to decide on this point. The single pair 
taken on Kauai are very large (5 mm. excl. rostr.), the male has the prothorax wider 
and more strongly constricted in front, and the prominences caused by the unevenly 
raised interstices of the elytra are more strongly developed. The % has the rostrum 
black, the thorax wider, and the prominences on the elytra stronger than those of 
typical A. lateralis. 

F. H. II. 18 



FAUNA HAWAIIENSIS 



(2) Acalles Inuneralis, sp. nov. 

Minor, haud latus, squamis pallidis (griseis vel subferrugineis) vestitus, antennis 
rostroque plus minusve rufescentibus. Pronotum totum squamis pallidis vestitum, 
medium sulcatum, antice constrictum, <J-is sat latum, trans medium vix minus latum 
quam longum, ?-ae fortiter elongatum, lateribus minus fortiter rotundatis. Elytra 
squamis pallidis vestita, ad humeros utrinque macula nigra distincta ornata, striis 
distinctis, obscurius punctatis, interstitiis omnibus convexis, secundo (praecipue ^) forti- 
ter inaequaliter elevato. 

t% Long, (rostr. excl.) 3 — 3'5 mm. 

Allied to A. lateralis but quite distinct, 

Hab. Maui, Haleakala (4500 ft.) ; one pair taken together. 



(3) Acalles callichronia, sp. nov. 

Haud latus, niger, rostro, antennis, tarsis, tibiarumque anticarum parte apicali, 
rufis. Rostri basis squamis albescentibus, caput ferrugineis ornatum. Pronotum 
elongatum, antice constrictum, medium sulcatum, squamis ferrugineis nigrisque varie- 
gatum. Elytra ad basim plaga magna nigra ornata, post banc fascia transversa pallida, 
ad latera dilatata, turn fascia transversa nigricante, parte apicali pallide squamosa, 
maculis nigris ornata ; interstitiis primo et tertio haud convexis, secundo fortiter 
interrupte elevato, partibus elevatis squamis nigris erectis densissime vestitis, quarto, 
quinto et sexto elevatis, striis haud evidenter punctatis. 

$ Long, rostr. excl. 3*5 mm. (Plate VII. fig. 10.) 

Hab. Hawaii, Kilauea, July 1895 ; one example taken. 



(4) Acalles iiielanolepis, sp. nov. 

Haud brevis, niger, antennis, rostro, tarsisque rufescentibus. Caput cum pronoto 
nigro-squamosum, hoc vitta laterali albida utrinque ornato. Pronotum angustum, 
elongatum, antice minus fortiter constrictum, obscurius longitudinaliter sulcatum, sulco 
medio plagam laeviorem elongatam rubricolorem ferente. Elytra nigro-squamosa, post 
media macula albida, trans suturam posita, ornata, interstitio 2° et 4° sat evidenter nee 
fortiter elevatis, squamis erectis vestitis, 1° et 3° parum convexis, striis obscure punctatis. 
Femora omnia nigro-squamosa, squamisque suberectis albidis conspersa, maculis vel 
cingulis pallidis carentia. Long, (rostr. excl.) circa 3 mm. (Plate VII. fig. 11.) 



COLEOPTERA 133 

In this species tlie elytra except for the well-marked transverse white spot, beyond 
the middle, are nearly entirely clothed with black scales, but there is an indication 
of an obscure paler area on each side extending from behind the shoulders towards the 
suture. The sides as seen from above are simply rounded from the shoulders, and 
not at all irregular. The second and fourth interstice are evidently more strongly 
raised than the first and third, the first being hardly, and the third but little convex. 
The second is slightly unevenly raised, and is densely clothed with erect black scales 
at the base and again just before the region of the transverse white spot, which 
accentuate the appearance of inequality. The most conspicuous feature of the species 
is the smooth red lanceolate space lying between the inner ridges of the prothorax, and 
the entirely black appressed scales of the femora. 

Hab. Hawaii. A single example taken at Kilauea. 



(5) A calks fraier, sp. nov. 

Praecedenti simillimus, nigro-squamosus, rostro rufo, antennis, tarsisque testaceis. 
Pronotum squamis ferrugineis supra maculatum, sulco medio plagam rubram haud 
ferente. Elytra plaga ferrugineo-squamosa fasciam latam formante ornata, apice 
squamis ferrugineis variegato. Femora media distincte pallide cingulata. Long, 
(rostr. excl.) circa 3 mm. 

Very closely allied to A. melanolepis, but the thorax is without the red smooth 
lanceolate area between the inner ridges, and is ornamented above with ferruginous 
spots, and the elytra have a large lateral band of similarly coloured scales, extending 
from behind the shoulders, and forming a wide transverse fascia which is less distinct 
towards the suture ; their apex is also variegated with paler scales. All the femora 
have a pale band of appressed scales at the base and another about the middle. 

A single example from Molokai is very similar to the above but the pale scales 
of the thorax and elytra are greyish, the transverse fascia of the latter less defined, 
the antennae, rostrum and tarsi much less clearly coloured, and the thorax evidently 
shorter. The difference in the latter respect is probably sexual, but I suspect that the 
two forms are distinct, though very closely allied. 

Hab. Maui, Haleakala (5000 ft.); one example. Van on Molokai (4000 ft.); 
one example. 

(6) Acalles koae, sp. nov. 

Haud brevis, rostro piceo, antennis tarsisque rufescentibus. Rostri basis cum 
capite antico pallide squamosa. Pronotum angustum, elongatum, antice constrictum, 
postice angustatum, longitudinaliter sulcatum, sulco medio obsolete carinato, squamis 

H S. Bsrbor, iS 2 

U. s. Notion^ Muwum, 
Washington, D- *" 



134 FAUNA HAWAIIENSIS 

fulvescentibus depressis, erectisque nigris vestitum. Elytra in parte basali squamis 
albescentibus vestita, nigro variegata, post banc fascia transversa nigra, maculaque alba 
trans suturam posita, apice fulvo-squamoso ; interstitiis 2° et 4° minus fortiter elevatis, 
1° et 3° baud convexis, striis (squamis baud remotis) impunctatis. Femora nigro- 
squamosa, media pallide notata. Long, (rostr. excl.) 3 mm. 

This prettily marked species appears to be quite distinct. The prothorax is 
clothed with appressed scales of a fulvous colour, and black erect ones along the two 
inner ridges. The basal half of the elytra is covered with pale squamosity of a whitish 
and pale brown colour, variegated with black markings, which are most extensive about 
the shoulders, and are partly formed by the basal elevation of the second and fourth 
interstices, which are there covered with dense erect black scales. A little behind the 
middle of the elytra there is a complete transverse black fascia, which posteriorly 
encloses a pure white trans-sutural spot. The first and third interstices are fiat, not 
perceptibly convex, the second and fourth are elevated but by no means strongly so, 
and apparently unevenly, though this appearance is partly at least due to the erect setae 
which are placed on them at the base and about the middle of the elytra, and give them 
an appearance of being more elevated than is really the case. The sides of the elytra 
are simply and regularly rounded from the shoulders, the striae show no visible 
puncturation without the removal of the scales. 

Hab. Hawaii. A single example beaten from Acacia koa at an elevation of 
about 5000 ft. in Kona. 



(7) Acalles innotabilis, sp. nov. 

Minus brevis, piceo-niger, rostro, antennis, tarsisque rufescentibus. Caput plus 
minusve pallide squamosum. Pronotum angustum, elongatum, obscurius longitudinaliter 
sulcatum, antice minus fortiter constrictum, squamis pallidis fuscescentibus vestitum. 
Elytra ex majore parte cum pronoto concoloria, squamisque nonnullis erectis nigris, 
maculaque albida postice trans suturam posita, ornata, humeris rotundatis, striis omnibus 
grosse punctatis, interstitio 2° subinaequaliter elevato, 1° et 3° subconvexis, 4°, 5°, 6° 
sat distincte subaequaliter elevatis. Tibiarum apices pallide squamosi. Long, (rostr. 
excl.) 3*2 mm. 

Hab. Kauai. A single example taken in the lower forest (about 2500 ft.). 



(8) Acalles leptothorax, sp. nov. 

Niger, rostro, antennis, tarsisque rufis, baud latus, sat elongatus, squamis pallidis 
nigrisque intermixtis variegatus. Rostrum post antennas squamis pallidis vestitum. 
Pronotum elongatum, lateribus leviter, fere aequaliter, rotundatis, antice baud subito 



COLEOPTERA 135 

angTJstatum vel constrictum, antice setis nonnullis erectis nigris, et ubique squamis 
nigris albidisque variegatim vestitum, supra vix evidenter sulcatum vel inaequale. 
Elytra squamis albidis nigrisque variegatim vestita, interstitiis convexis, 2" parum 
fortiter inaequali, striis parum distincte punctatis, lateribus sat aequaliter rotundatis. 
Femora nigra plus minus distincte albido-cingulata, tibiis nigro-squamosis, apices versus 
(circa \ — \ longitud.) squamis pallidis vestitis, ibique rufescentibus. Long, circa 3 mm. 
(Plate VII. fig. 12.) 

Remarkable for the very narrow long prothorax, which is quite evenly rounded at 
the sides. The extreme apex of the elytra is covered with pale scales, in front of which 
is a wide band of black squamosity enclosing a quadrate pale trans-sutural spot, which 
itself is dark in the centre. In front of this the elytra are variegated with black and 
pale scales intermixed. 

Hab. Kauai. A single example was taken at Halemanu (4000 ft.). 

(9) Acalles decoratus Blackburn. 

Acalles decoratus Blackb., Tr. Dublin Soc. iii. 1885, p. 180. 
H.AB. " Lanai, 2000 ft." " A single specimen " (Blackburn). 

(10) Acalles mauiensis Blackburn. 

Acalles Jtmuiensis Blk., t. c. pp. 179, 181, 254. 

Hab. Maui. " A single specimen beaten from Aleurites triloba, at an elevation 
of about 4CXX) ft. on Haleakala." (Blackburn.) 

The specimen referred to on p. 181, /. c. is said, p. 254, /. c, to be probably 
a distinct species. 

(11) Acalles angusticollis Sharp. 

Acalles angusticollis Sharp, Tr. Dublin Soc. in. 1885, p. 179. 

Hab. Maui, Haleakala (Blackburn), where I have myself taken a single example. 
Minute examples taken on Oahu (var. 7?ii?ior, Shp.) by Mr Blackburn are probably 
distinct, and almost agree with abraded specimens taken by myself on Lanai. 

(12) Acalles duplex Sharp. 

Acalles diiplex Sharp, /. c. p. 1 78. 
(Plate VII. fig. 13. ?; fig. 14, ?■) 

Hab. Oahu. Widely distributed on the island. Varies greatly in size, the 
females apparently not being always of diminutive size as compared with the males. 



I -,6 FAUNA HAWAIIENSIS 



(13) Acalles ignotus Blackburn, 

Acalles ignotus Blackb. t. c. p. 180. 

Hab. Oahu. A single example (Blackburn). 



(14) Acalles lanaiensis, sp. nov. 

Valde convexus, pallide squamosus, rostro piceo, antennis tarsisque rufescentibus. 
Caput cum pronoto squamis palUdis vestitum. Pronotum sat latum, antice fortius con- 
strictum, medium postice distincte sulcatum. Elytra pallide squamosa, plaga humerali 
utrinque saepe nigra, angulis humeralibus sat distinctis, fere rectis, interstitio 2° plus 
minusve inaequaliter elevato, tertio quartoque elevatis, illo angulos humerales formante, 
striis obscure punctatis. Long, (rostr. excl.) vix 3 mm. 

Apparently allied to Acalles dttplex, Shp., but much smaller and the sculpture of 
the elytra much less strongly pronounced. 

Hab. Lanai Mountains ; several examples taken. 



(15) Acalles chlorolepis, sp. nov. 

Brevior, robustus, griseo-squamosus, rostro, antennis, tarsis, tibiarumque apicibus 
plus minusve rufescentibus. Caput griseo-squamosum ; antennarum articulo tertio brevi. 
Pronotum baud latum, squamis griseis depressis, paucisque erectis nigricantibus vestitum, 
longitudinaliter 5-sulcatum, lateribus minus fortiter rotundatis. Elytra griseo-squamosa, 
apicem versus utrinque plaga nigricante, interstitiis 2° et 4° sat fortiter inaequaliter 
elevatis, 1° et 3° vix vel baud convexis, striis internis obscure punctatis, angulis humera- 
libus sat distinctis. Femora nigro-squamosa, squamis erectis griseis conspersa. Long, 
(rostr. excl.) vix 3 mm. 

Hab. Kauai (4000 ft.) ; two or three examples taken. 



(16) Acalles nigripennis, sp. nov. 

Sat latus, antennis tarsisque rufescentibus, rostro rufopiceo. Caput pallide squamo- 
sum. Pronotum basi latum, antice fortiter constrictum, squamis pallidis vestitum, 
postice fortius sulcatum. Elytra tota nigro-squamosa, lateribus baud aequaliter rotun- 
datis, interstitiis 2" et 4° subfortiter nee aequaliter elevatis, striis obscure punctatis. 
Femora omnia ubique pallide squamosa. Long, (rostr. excl.) 3 mm. (Plate VI L 
fig- 1 5-) 



COLEOPTERA 137 

The elytra have the second interstice rather strongly raised, rather more strongly 
at the base and at about their middle than between these points ; the third is hardly 
convex ; the fourth strongly and unevenly raised, forming the humeral angles ; the fifth 
and sixth are also more or less unevenly raised, the latter in dorsal aspect forming the 
lateral outline of the elytra behind the shoulders, which owing to the form of this 
interstice has an angulate appearance, instead of being regularly rounded from the 
base. 

Hab. Lanai, a single example taken in the mountains. 



( I 7) Acalles tuberculatus, sp. nov. 

Piceus, rufopiceus, vel rufescens, minus brevis, rostro, antennis tarsisque rufescen- 
tibus, squamis pallidis nigrisque vestitus, squamis erectis elongatis et gracilibus. 
Pronotum haud latum, plus minusve evidenter longitudinaliter 5-sulcatum, sulco medio 
tuberculo elongate laevi munito, antice fortius constrictum, squamis ex majore parte 
nigris vestitum. Elytra squamis nigris pallidisque variegata, striis obscure punctatis, 
interstitiis 2° et 4° aequaliter, 6° inaequaliter elevato, humeris prominulis. Long. 
275—3 mm. 

This species is very distinct by the smooth tubercle situated about the centre of 
the prothorax, and the form of the erect scales, which are longer and more slender than 
those of other species. The elytra have their humeral angles somewhat acute and 
prominent, the 2nd and 4th interstices moderately strongly, and evenly raised, the 6th 
is somewhat strongly raised a short distance behind the shoulders, and forming the 
outline of the elytra in dorsal aspect, gives it at that point an appearance of being 
angulated. One example has the elytra nearly entirely clothed with pale scales. Most 
of them are for the most part or entirely denuded. 

Hab. Hawaii ; Kona (4000 ft.). About 10 examples found under small logs. 



(18) Acalles pallidicollis, sp. nov. 

Minus brevis, niger, rostro piceo, tarsis antennisque rufescentibus. Caput rostrique 
basis squamis pallidis vestita. Pronotum totum pallide squamosum, elongatum, antice 
constrictum, obscurius sulcatum. Elytra squamis pallidis nigrisque variegata, basi 
apiceque ex majore parte nigris, media parte pallide squamosa, obscure nigro fasciata, 
striis internis obscure grosse punctatis, interstitiis 2° et 4° fortius, primo et tertio his 
levins elevatis. Femora omnia pallide vestita, nigro-cingulata, squamis erectis pallidis. 
Long, (rostr. excl.) 3-25 mm. 

Hab. Kauai, above 2000 ft. One example taken. 



138 FAUNA HAWAIIENSIS 



(19) Acalles monticola, sp. nov. 

Minus brevis, pallide squamosus, rostro piceo, antennis basi rufescentibus. Caput 
cum basi rostri squamis depressis erectisque pallidis vestitum. Pronotum elongatum, 
antice constrictum, obscurius 5-sulcatum, squamis pallidis depressis, nonnullisque nigris 
erectis vestitum. Elytrorum pars basalis squamis pallidis vestita, nigro-punctata, post 
media plaga transversali nigricante, quae maculam albidam, trans suturam positam, 
continet, apice pallido; interstitiis 2° — 6° fortius aequaliter convexis, striis obscure grosse 
punctatis. Femora omnia pallide squamosa, media parte late nigrocincta. Tibiae ad 
basim nigro-squamosae. Long, (rostr. excl.) 3'5 mm. 

In this species all the interstices of the elytra are raised, except the first, which is 
only slightly convex. There is very little difference between the convexity of the other 
interstices, that of the 3rd being slightly less than the 2nd and 4th. 

Hab. Oahu. A single example taken in the Waianae mountains. 



(20) Acalles oahuensis, sp. nov. 

Minus brevis, pallide squamosus, rostro antennisque rufis. Caput squamis pallidis 
depressis, nigrisque erectis, vestitum. Pronotum elongatum, antice subfortiter constric- 
tum, 5-sulcatum, lateribus rotundatis, squamis pallidis nigrisque variegatum, aliisque 
erectis nigris vestitum. Elytra squamis pallidis vestita, plaga magna basali nigra, 
fasciisque post media duabus irregularibus transversis ornata, leviter striata, striis parum 
distincte punctatis, interstitiis 2°, 4°, et 6° aequaliter elevatis, 1°, 3° et 5° baud convexis. 
Femora cum tibiis ex majore parte squamis nigris vestita. Long, (rostr. excl.) 3 mm. 

A very distinct species, each of the elytra with three very well-marked longitudinal 
costae, which continue the three ridges on either side of the thorax. The alternate 
interstices are not at all raised or convex. 

Hab. Oahu. A single example taken in the Waianae mountains (2000 ft.). 



Chaenosternum Blackburn. 

( I ) Chaenosternum konamun Blackburn. 

Chaenosternum konanum Blackb., Tr. Dublin Soc. iii. 1885, p. 181. 
Hab. Oahu. One example taken near Honolulu (Blackburn). 



COLEOPTERA 



139 



Hyperomorpha Blackburn, 
(i) Hyperomorpha squamosa Blackburn. 
Hyperomorpha squamosa Blackb., /. r. p. 182. 

Hab. Oahu. A single example taken in wet moss in the Pausa valley (Black- 
burn). I have not met with this, or the preceding genus. 

Tribe CALANDRINI. 

Sphenophorus Schonherr. 

(i) Sphenophorus obsairiis Boisd. 

Calandra obsciira Boisd. Voy. Astr. 11. p. 448; Fairm. Rev. Zool. 1S49, p. 474. 
The various islands. 

Hab. a destructive species in the fields of sugar-cane, well-known as the ' cane- 
borer.' Also on banana in the mountains and sometimes found under rocks. The 
species has probably a wide distribution outside the Archipelago, though very little has 
been written about it. 

Calandra Clairville. 
(i) Calandra reniota Sharp. 

Cala7idra reviota Sharp, Tr. Dublin Soc. iii. 1885, p. 183. 
Hab. Oahu (Blackburn). Maui ; in banana stems ; common. 

(2) Calandra oryzae, L. 
Calandra oryzae Linn., Amaen. Ace. vi. p. 395. 
Hab. All the islands ; in food. 

(3) Calandra linearis van striata Thunb. 
Calandra linearis var. striata Thunb. Nov. Act. Ups. vii. p. 112. 
Hab. Oahu ; in decaying tamarinds (Blackburn). 

Tribe COSSONINI. 

Dryophthorus Schonherr. 

(i) Dryophthorus squalidus Sharp. 

Dryophthorus squalidus Sharp, Tr. Ent. Soc. London, 1S78, p. 22. 
Hab. Common throughout the islands in the mountains. 

This abundant species is extremely variable in size, the smallest examples are less 
than four, the largest more than six, mm. in length. All the interstices of the elytra 
F. H. II. 19 



I40 FAUNA HAWAIIENSIS 

are covered with a cinereous bloom. The apical portion of the rostrum in the ? (from 
the insertion of the antennae to the apex) is impunctate, or at the most has a few 
excessively feeble and fine punctures. 

(2) Diyophthortis distinguendus, sp. nov. 

Niger, piceus, vel rufescens, haud robustus, grosse punctatus. Rostri (?-is pars 
anterior opaca, rugoso-punctata, $-ae nitida, subtilius sat distincte fere ad apicem 
punctata. Oculi sat magni. Pronotum densissime punctatum, antice fortiter constrictum. 
Elytra satis profunde sulcata, sulcis confertim grosse punctatis, interstitiis haud cinereo- 
sericeis, subtilissime punctulatis, et brevissime pallide setulosis. Long, (rostr. incl.) 
4—5-5 mm. 

This species greatly resembles D. squalidzis Shp., and is often found in company 
with that species, but is certainly distinct. The interstices of the elytra are never 
covered with the whitish bloom as in that species, and the minute punctures which they 
bear give rise to short pale-coloured setae. The % is readily distinct from that of 
D. sqtialidiis by the fact that the rostrum has an evident, though not coarse, punctura- 
tion on its anterior shining portion. Like most of the Hawaiian species this varies 
considerably, but it is by no means improbable that I have included more than one 
species under the above name. The raised apical margin of the elytra is continuous 
with the sixth interstice as in D. squalidus. 

Hab. Found on all or nearly all the islands of the group. 

(3) Dryophthorus peles, sp. nov. 

Niger, robustus, grosse punctatus. Rostri % pars praeantennalis nitida, subtiliter 
sat evidenter punctata. Pronotum latum, grosse punctatum, antice fortiter constrictum. 
Elytra fortiter seriatim punctata, interstitiis distinctissime punctulatis, brevissime pallide 
setulosis, haud cinereo-sericeis, margine apicali elevato cum interstitio sexto continuo. 
Long, (rostr. incl.) 6 — 6 '5 mm. 

This species somewhat resembles D. distingttendus, but it is altogether larger and 
more robust. The interstitial puncturation of the elytra is larger, and more distinct and 
definite than in most of the species. The punctures in each row are closely set, and the 
spaces dividing one from another are more than usually convex, grooved on each side 
where they meet the interstices. This convexity gives them a strongly marked knob- 
like appearance in certain aspects. The whitish bloom seen in D. squalidus is entirely 
absent, but the interstitial punctures bear excessively minute pale setae. The interstices 
themselves are all of nearly similar height, and the sixth is confluent with the elevated 
apical side-margin. 

Hab. Hawaii. Kilauea (June 1895). Half-a-dozen examples, no doubt obtained 
in a batch, as they stand under the same number. The species appears to me to be 
most closely allied to D. nesiotes. 



COLEOPTERA 141 

(4) Dryophthorus nesiotes, sp. nov. 

Niger, robustus, statura magna. Rostri pars praeantennalis ? nitida, baud, vel vix 
evidenter, punctata. Pronotum latissimum, antice fortissime constrictum, fortiter dense 
punctatum. Elytra seriatim fortiter punctata, interstitiis cinereo-sericeis, tuberculis 
minutis, sat distinctis, munitis. Long, (rostr. incl.) 6 — 775 mm. (Plate VII. fig. 16.) 

This species also belongs to the group in which the elevated apical margin is 
continuous with the sixth interstice of the elytra. The interstices themselves (except in 
very much rubbed examples), have a distinct ashy covering. The formation of the 
spaces between the punctures in each row on the elytra is nearly similar to that in 
D. peles. The present species is distinct by its large size, the distinct minute asperities 
with which the interstices are set, and their ashy covering. 

Hab. Kauai mountains (4000 ft.). 

(5) Dryophtho7'ns g^^avidiis Sharp. 

Dryophthor7ts gravidus Sharp, Tr. Ent. Soc. London, 1878, p. 22. 

To this species I refer a great number of e.xamples taken on Molokai, Lanai, Maui, 
and Hawaii. The difierences between the extremes are so great that they do not 
appear, when examined together, to be even very closely allied species, but after 
examining the whole series from the various localities, I am forced to consider all 
to belong to one variable form. Minute individuals appear to be always distinguishable 
from D. squalidus etc. by the shorter and wider elytra, differences in puncturation etc. 
Some examples from Maui are very remarkable, the punctures on the elytra becoming 
distorted and tending to become obsolete, the elytra themselves being laterally com- 
pressed. A similar tendency is also seen in some specimens from other islands. The 
elytra are much more narrowed posteriorly in some examples than in others. The 
puncturation of the prothorax varies in the size and density of the punctures ; those on 
the elytra also are variable. There is a decided tendency towards the sculpture of the 
elytra exhibited by D. crassus Shp. in some Maui individuals. 

Hab. Oahu (Blackburn) ; all the islands to windward of Oahu. 

(6) Dryophthorus crassus Sharp. 

Dryophthorus crassus Sharp, Tr. Ent. Soc. London, 1878, p. 23. 

Hab. Oahu and Maui (Blackburn). I have taken it singly near Honolulu, but 
not elsewhere, unless some of the examples from Maui, included under D. gravidus, 
should be referred to this species. 

19 — 2 



142 FAUNA HAWAIIENSIS 

(j) Di'yophthorus homoeorhynchus, sp. nov. 

Niger, vel piceus, haud robustus, rugoso-punctatus. Rostrum utriusque sexus 
(apice extremo excepto), opacum, rugoso-punctatum. Pronotum densissime rugoso- 
punctatum, antice fortissime constrictum. Elytra sat profunde sulcata, sulcis grosse 
punctatis, interstitiis punctulatis, punctis setas brevissimas pallidas, satis conspicuas, 
ferentibus. Long, (rostr. incl.) 4"5 — 6 mm. (Plate VII. fig. 17; fig. 17a, head and 
rostrum of ?; fig. 17^ of ^.) 

This species, although the elevated apical margin appears rather continuous with 
the sixth than the fourth interstice of the elytra, evidently belongs to the group of 
D. declivis, from which it is also separated at once by the fact that there is little 
difference in the elevation of the interstices, whereas in that species the second, fourth 
and sixth are very noticeably more elevated than the others. From those species 
of the group of D. squalidus which it somewhat resembles, the female is distinguished 
at once by the dull apical half of the rostrum, with its rugose puncturation, and the ^ by 
the more outstanding pterygia, so that the rostrum is more widened about the point of 
insertion of the antennae. The sexes are easily known by the more slender rostrum of 
the female. 

Hab. Kauai mountains (2000 — 4000 ft.). 

(8) Dryophthorus declivis Sharp. 

Dryoplithot'iis dcclii>is Sharp, Tr. Ent. Soc. London, 1878, p. 23. 
(Plate VII. fig. 18.) 

This is a very distinct species easily known by the costate 2nd, 4th and 6th 
interstices of the elytra, the two latter uniting to form the strongly raised apical side- 
margin. These interstices vary somewhat in the amount of elevation, but it is always 
very apparent. The female has the rostrum more slender than the ^, less widened at 
the insertion of the antennae, but the dull surface and rugose puncturation to near the 
apex is common to both sexes. 

Hab. Oahu, Molokai, Hawaii. Not so common as many of the species. 

(9) DryophtJiorus modestiis Sharp. 

DryophtJioi-us nwdestus Sharp, Tr. Ent. Soc. London, 1878, p. 23. 

The sexual distinctions in this species are similar to those exhibited by the sexes 
in D. homoeorliynchtLs and D. declivis. It varies considerably in size and the elevation 
of the apical margin, the connection between which and the fourth interstice is rather 
less manifest in some examples than others. 

Hab. Oahu, Maui, and Hawaii. 



COLEOFTERA 143 



(10) Dryophthoi'us pnsillus Sharp. 

Dryophihorns pusillus Sharp, t. c. p. 24. 

Hab. Oahu (Blackburn) ; in stems of the tree-fern. I have never met with this 
species. 



(11) Diyopht horns oahuensis, sp. nov. 

Species minima, minus lata, rufescens, capite nigricante. Rostrum (^) latum, 
rugosum, et opacum. Pronotum antice fortiter constrictum, grosse rugoso-punctatum. 
Elytra seriatim grosse punctata, interstitiis omnibus distinctis, 2° et 4° quam 3° et 5° 
paullo fortius elevatis, 6° ad apicem declivem fortissime carinato-elevato. Long. Z 
(rostr. incl.) 27 mm. 

The rostrum is rather short and broad, the pterygia strongly outstanding. The 
eyes are feebly facetted. The puncturation of the prothorax is coarse and rugose but 
apparently shallow. All the interstices of the elytra are distinct, but not wide, the 
second and fourth a little more raised than the third and fifth, and not interrupted, the 
sixth alone forms the extremely strongly elevated apical side-margin. The species is 
very distinct. 

Hab. Oahu. A single example (.?) taken in the Waianae mountains (3000 ft). 



(12) Dryophthorus kauaiensis, sp. nov. 

Species minima, rufescens. Caput (^) cum rostro rugoso-punctatum, squamosum. 
Pronotum dense grossissime rugoso-punctatum, antice constrictum. Elytra sat grosse 
seriatim punctata, interstitio 2°, 4° et 6° haud interrupte carinato-elevatis, tertio sub- 
obsoleto. Long. ^ vix 3 mm. 

This species is allied to D. oaJmensis, but it is a little larger, and abundantly 
distinct by the sculpture of the elytra, the sharply carinate form of the second and 
fourth interstices readily separating the two. The apical margin is formed by the sixth 
interstice, which is strongly raised towards the apex. In D. ptisilbis Shp., which is a 
small species, the fourth interstice is continuous with that margin, as also is the case 
with D. niodestus Shp. Both in D. kauaiensis and the preceding species the eighth 
interstice forms a part of the lateral outline towards the base of the elytra, in dorsal 
aspect. In the present species indeed it is rather strongly raised. 

Hab. Kauai. A single J taken in the mountains (4000 ft.). 



144 FAUNA HAWAIIENSIS 

(13) Dryophthorus insignis Sharp. 

Dryophthorus insignis Sharp, /. c. 1878, p. 24. 

(Plate VII. fig. 19.) 

I have examined numerous specimens which I refer to this species from the islands 
of Oahu, Maui, and Hawaii. It also is found on the other islands. The examples 
from the islands specified do not altogether agree, but as the series from each locality 
exhibits some variation I see no means of separating them as distinct forms. Certainly 
in the majority of cases the Oahuan examples can be distinguished by the more 
imperfect eyes, but there is evident individual variation in this respect. The Maui 
form has the raised interstices of the elytra always very strongly broken, and the eyes 
are more developed. In specimens from Hawaii the eyes are well-developed, convex 
and a little prominent. From the other islands I have seen only odd specimens. 

Hab. Found under bark of trees in the mountains of all the islands. 

(14) Dryophthoriis insignoides, sp. nov. 

Piceus, vel plus minusve rufescens, sat grosse punctatus. Oculi mediocres. Rostri 
($) pars praeantennalis nitida, rugoso-punctata. Pronotum elongatum, antice fortiter 
constrictum, obscurius grosse punctatum. Elytra seriatim grosse punctata, interstitio 
2° et 4° inaequaliter carinato-elevatis, setis vestitis, primo sat distincto. Long, (rostr. 
incl.) 4 — 475 mm. 

The general sculpture of this species is as in D. insignis Shp., from which it is 
readily distinguished by the distinct first interstice of the elytra, the carinae of which 
are less strongly elevated. All the raised interstices (i.e. 2nd, 4th and 6th) bear erect 
setae, and the second and fourth are evidently interrupted or uneven. The third and 
fifth interstices are generally more or less distinct. The eyes are fairly well developed. 
The rostrum of the female on the part in front of the antennae has the surface shining, 
and is strongly and rugosely punctured, in the $ this part is dull and covered with 
squamosity. 

Hab. Kauai mountains. Single examples from other islands, Molokai, Lanai, 
and Oahu, do not altogether agree with the typical specimens, but are too close to 
separate without the examination of a fair series. 

(15) Dryophthorus brevipennis, sp. nov. 

Praecedenti simillimus, statura minore, elytris brevioribus distinguendus. Long. 
3—375 mm. 

Very similar to the preceding but a decidedly smaller insect. The second, fourth 



COLEOPTERA 145 

and sixth interstices of tlie elytra are more or less raised and bear fine setae, but the 
elevations are feeble. The other interstices are quite distinct. The elytra themselves 
are decidedly shorter than in D. insignoides, the rostrum in the $ is more finely and less 
rugosely punctured than in that species. In a strict dorsal aspect of the elytra the 
acute 8th interstice always forms a portion of their lateral outline near the base. 

Hab. Kauai mountains (4000 ft.). 



(16) Dryophthorus fuscescens, sp. nov. 

Haud latus, statura minore, fuscescens. Oculi minimi. Pronotum antice fortiter 
constrictum, grossius punctatum. Elytra seriatim grossius punctata, interstitiis cunctis 
sat distinctis, secundo quartoque inaequaliter distincte carinato-elevatulis, cumque 
sexto, setis brevissimis vestitis. Long. 3 '5 mm. 

A small and very distinct species, with unusually small eyes, which consist of 
comparatively few coarse facets. The rostrum of the % is closely, coarsely and rugosely 
punctured between the smooth ape.x and the line of insertion of the antennae, and is 
hardly shining. In the ^ the sculpture of the rostrum is entirely concealed, and this 
may possibly be the case with the % when alive, but the covering is removed in the 
single example of this sex. 

Hab. Kauai (4000 ft.) ; 2 ^ and i % taken. 



(17) Dj-yophthorus verticalis, sp. nov. 

Piceus, subrobustus. Rostri (?) pars praeantennalis nitida, fortiter distincte punc- 
tata. Oculi sat magni. Pronotum grossius punctatum, antice fortiter constrictum. 
Elytra seriatim grosse punctata, interstitiis sat distinctis, secundo quartoque inaequaliter 
elevatis et setulosis, postice fortissime declivia, fere verticalia. Long, (rostr. excl.) 
3 — vix 4 mm. (Plate VIL fig. 20.) 

The male of this species is at once distinguished from any other of the section by 
its short robust form as seen in dorsal view, the elytra becoming wider at the base of 
the posterior declivity than they are across the shoulders. Posteriorly they are strongly 
vertical and of slightly recurved form. The female is less remarkable but it is more 
robust and larger than the allied species. The rostrum is shining and strongly punc- 
tured. In both sexes the sinuation of the 6th interstice of the elytra causes the 8th 
to be distinctly visible in dorsal aspect, so that it evidently forms part of the lateral 
outline. The second and fourth interstices are unevenly raised, or carinate, and 
setulose. 

Hab. Kauai mountains (4000 ft.). 



146 FAUNA HAWAIIENSIS 



Thallatodora, gen. nov. 

Corpus subgracile, fusiformi- ovale, rugoso-sculpturatum. Rostrum elongatum, ad 
insertionem antennarum fortissime curvatum, sive geniculatum, parte praeantennali 
fortius dilatata. Antennae longe ante medium rostrum insertae, scapo elongato, funiculo 
capituloque una conjunctis baud minus longo, funiculi 5-articulati articulo secundo 
sequentibus evidenter longiore, capitulo brevi, robusto. Oculi subrotundi, fortiter 
prominentes, in rostro, a capite distantes, siti. Pronotum elongatum, antice constric- 
tum, elytris angustius. Scutellum vix discernendum. Tarsi breves, articulo tertio 
bilobato, lobis parvis. Corpus totum subtus dense ac grosse punctatum ; coxae poste- 
riores late, intermediae his minus late, anteriores intermediis minus late separatae, 
haudquaquam tamen contiguae. Metasternum sat longum. Abdominis pars basalis 
(sc. segmentum i et 2, quae parum distincte sunt divisa), mesosterno una cum meta- 
sterno baud minus longa. 



( I ) Thalattodora insignis, sp. nov. 

Piceo-nigra, tarsis antennisque rufescentibus, opaca. Rostrum rugosum. Pronotum 
elongatum, antice constrictum, lateribus parum fortiter rotundatis, grosse denseque 
rugoso-punctatum. Elytra profunde striata, striis grosse confertim punctatis, interstitiis 
asperis, punctulatis, setis brevissimis pallidis, uniseriatim dispositis. Long, (rostr. incl.) 
3'5 mm. (Plate VIII. fig. i; i«, antennae.) 

Hab. Lanai. A single example taken under logs on the coast. 



Pentarthrum Wollaston. 

(i) Pentarthrum prolixum Sharp. 

Pentartlirtiin prolixum Sharp, Tr. Ent. Soc. London, 1878, p. 25. 

Hab. Found on all the islands, living on tree-ferns. Varies greatly in size. 

(2) Pentarthrum obscuriim Sharp. 

Pentarthrum obscurum Sharp, /. c. 

Hab. Oahu. Abundant in dead wood up to an elevation of 2000 ft. Often in 
company with Phloeophagosoma and Pseudolus. 



COLEOPTERA 147 

(3) Pentarthrum blackbiirni Sharp. 

Pentarthrum blackbtirni Sharp, /. c. 

Hab. Oahu. Near Honolulu (Blackburn). I did not meet with this species. 

Orothreptes, gen. nov. 

Subfusiformis, haud gracilis, totus pube aurea vestitus. Rostrum elongatum, levius 
curvatum, lateribus parallelis. Oculi magni convexi. Antennae breves, robustae, fere 
ad medium rostrum insertae, scapo minus longo, funiculo 5-articulato, articulo primo 
brevi, caeteris transversis, capitulo elongate. Pronotum latum, antice subconstrictum. 
Tarsi breves, sat robusti, articulo tertio late bilobato. Coxae omnes sat late separatae, 
sed posteriores et intermediae multo latius quam anteriores. Metasternum longum. 
Elytra pronoto evidenter latiora. Scutellum patens. 

Perhaps most nearly allied to Sericotrogus Woll., but differing greatly in its much 
less narrow and elongate form, the less strongly curved rostrum, the pterygia being not 
in the least outstanding, the larger but less prominent eyes, short wide prothorax, 
shorter metasternum, and in many other respects. The lobes of the tarsi are very 
largely developed, the apical joint is somewhat stout. 

(i) Orothreptes callithrix, sp. nov. 

Rufotestaceus, totus aureo-pubescens. Rostrum opacum, subtilissime punctatum, 
lateribus parallelis. Pronotum sat latum, antice subconstrictum, densissime grossius 
punctatum, lateribus rotundatis. Elytra profunde striata, striis haud distincte punctatis, 
interstitiis confuse crebreque rugoso-punctatis. Long, (rostr. incl.) circ. 4 '2 5 mm. 
(Plate VIII. fig. 2 ; 2a, antenna). 

Hab. Hawaii. A single example taken in the Kona district (3000 ft.) 

Deinocossonus, gen. nov. 

Forma subcylindrico-fusiformi, totus pube demissa vestitus. Rostrum latum, breve, 
a capite parum distincte divisum, medium postice longitudinaliter sulcatum, ad inser- 
tionem antennarum subangustatum, ibique utrinque tuberculo munitum. Oculi per- 
magni, convexiusculi, a pronoto distantes. Antennae ad apices oculorum insertae, ab 
apice rostri distantes, funiculo 7-articulato, articulo primo caeteris, qui omnes sunt bre- 
vissimi, longiore, capitulo sat robusto. Pronotum subelongatum, antice angustatum. 

F. H. II. 20 



148 FAUNA HAWAIIENSIS 

Elytra ac pronotum aeque lata. Scutellum conspicuum. Tarsorum articulus tertius sat 
latus, bilobatus. Uncus tibiarum anteriorum apicalis validus. Femora robusta. Coxae 
omnes late separatae. Metasternum satis longum. Abdomen subtus pubescens, fortiter 
punctatum. 

A genus, the species of which is of small size, and apparently without any allied 
form known from elsewhere. It is very remarkable for the short wide rostrum, the very 
large, but not prominent eyes, and the dense covering of golden pubescence. In the 
testaceous colour, and the clothing, these insects have an extraordinary resemblance to 
the insect, for which I have made the genus OrotJireptes, but that has a totally different 
form of rostrum, and a 5-jointed funiculus, and differs in many other respects. Probably 
in habits the two are identical. 

Deinocossonus is a form of excessive rarity, or at least very difficult to procure, and 
the pubescence is so easily removed, that I have been unwilling to subject the few 
specimens taken to much manipulation. 

(i) Deinocossomts nesiotes, sp. nov. 

Angustus, testaceus, aureo-pubescens, capite rostroque nigris. Pronotum dense, 
nee profunde punctatum, inter puncta subtilissime rugulosa. Elytra striata, striis obscu- 
rius, plus minusve evidenter punctatis, interstitiis obscure punctulatis, et pubescentibus. 
Femora anteriora supra testacea, parte inferiore nigricante. Long, (rostr. incl.), 2 '5 — 
vix 3 mm. (Plate VIII. fig. 3 ; 3a, antenna.) 

Var. a. elytrorum striis haud evidenter punctatis, rostri parte apicali testacea. 

Var. y8. rostro et capite nigris, femora omnia cum tibiis nigricantia. 

Hab. Oahu. Two examples taken together at an elevation of 3000 ft. on Kaala 
mountain. Var. a. Kauai, 4000 ft. ; Oahu, Waianae mountains. Var. ^. Hawaii. 
One specimen at Kilauea. 

Haloxenus, gen. nov. 

Corpus elongatum, gracile, fusiforme, haud pubescens. Rostrum latum, breve 
subconicum (sc. lateribus ad apicem evidenter convergentibus), a capite minus distincte 
divisum. Oculi rotundi, latissime separati, a pronoto sat distantes. Antennae paullo 
ante medium rostrum insertae, funiculi 7-articulati articulo primo robusto, caeteris 
longiore, secundo caeterisque brevissimis, capitulo sat magno, apice acumlnato. Pro- 
notum elongatum antice angustatum, vix constrictum. Pedes sat robusti ; tarsi breves, 
articulo tertio parvo, haud evidenter bilobato, articulo ultimo elongato, caeteris conjunctis 
haud minus longo. Metasternum perlongum. Coxae omnes late separatae. Elytra 
elongata, angusta, lateribus parallelis. Scutellum patens. 

This genus comes nearest to Eutornns WolL, but has the rostrum narrowing to 
the apex, and the point of insertion of the antennae is different. 



COLEOPTERA 149 



(i) Haloxenus immigrans, sp. nov. 

Piceus, vel rufopiceus, subnitidus, elongatus, gracillimus. Rostrum basim versus 
dense sat fortiter, ad apicem subtiliter punctatum. Pronotum fortiter punctatum, antice 
sensim nee fortiter angustatum. Elytra striata, striis sat grosse punctatis, circum 
suturam subtilissime punctulata. Long, (rostr. incl.) circ. 3 mm. (Plate VIII. 
% 4-) 

Hab. Molokai and Lanai coasts. Not rare under logs of drift-wood. 



PsEUDOLUS Sharp. 

(i) Pseudolus longulus Boheman. 

Rhyncolus longiihts Boh., Eug. Res. p. 149. 

Hab. Oahu and Maui. Abundant, and no doubt inhabits other of the islands, 
at low elevations. On Cactus, Aletiriies, Musa, &c. 

(2) Pseudolus Jiospes, sp. nov. 

Niger, gracilis, elongatus, subnitidus. Rostrum confertim punctatum, parte prae- 
antennali fortius dilatata. Pronotum elongatum, fortiter crebre punctatum, antice 
fortiter constrictum. Elytra striata, striis grosse confertim punctatis, interstitiis 
punctulatis, setas minimas pallidas ferentibus, et transversim subtilissime rugulosis. 
Tarsorum articulus tertius, secundo evidenter latior, bilobatus, lobis parvis. Long. 
6"5 mm. 

Hab. Oahu. Four examples taken, two in Honolulu and two a short distance 
outside. 

This species has only been met with near Honolulu, and I suspect it has been 
introduced. Generically it appears to be somewhat intermediate between Psetidolus 
and Phloeophagosovia. 

Phloeophagosoma Wollaston. 
( I ) Phloeophagosoma tenuis Gemm. 

Rhyncolus tennis Gemm. Mun. Cat. viii. p. 2667. 
Rhyncolus gracilis Boh. Eug. Res. p. 149. 

Hab. Oahu, up to 2000 ft. ; common. Hawaii, in Kona (2000 — 3000 ft.). 

20 — 2 



I50 FAUNA HAWAIIENSIS 

Nesotocus, gen. nov. 

Forma subfusiformi, statura magna. Rostrum cylindricum, perlongum, maris 
rugose sculpturatum, saepissime spinulosum, feminae parte praeantennali laevi, nitida, 
impunctata. Oculi magni, convexiusculi, a pronoto sat distantes. Antennae longissimae, 
graciles, maris paullo ante vel fere ad medium rostrum, feminae longe post medium 
insertae, funiculo (^) perspicue piloso, 7-articulato, articulis basalibus fortissime elongatis, 
capitulo elongato, subfusiformi. Pronotum antice fortiter angustatum, truncato-conicum. 
Scutellum patens. Pedes longissimi, femoribus anterioribus ^ gracilioribus, % robusti- 
oribus et magis incrassatis, tarsorum articulo primo et secundo elongatis, tertio late 
bilobato, articulo ultimo gracili elongato, apicem versus crassiore, unguiculis longis et 
curvatis. Coxae anteriores (necnon etiam intermediae et posteriores) late separatae. 
Metasternum sat longum, postice subangulariter emarginatum, medium longitudinaliter 
impressum vel sulcatum. Abdominis segmentum primum ventrale secundo paullo 
longius, tertium una cum quarto hoc paullo brevius, segmentum apicale ^ latius $ minus 
late rotundatum. Elytra et pronotum pube pallida decorata, hoc et illis subaequilatis. 

There is apparently no known ally to the three remarkable insects for which the 
above genus is formed. They all live in the wood of the tree Cheirodendron, and, I 
believe, of another tree which has a considerable resemblance to that genus. In 
superficial appearance the three species almost e.xactly resemble one another, and this 
is rather that of Erirhinini than Cossonini. 

( I ) Nesotocits imtnroi, sp. nov. 

Niger, pubescentiae pallidae maculis ornatus, antennis tarsisque nonnunquam 
rufescentibus. Rostrum $ (saltem in exemplis majoribus), rugoso-sculpturatum, utrinque 
serrato-spinosum, in exemplis parvis vi.x plus quam simpliciter punctatum, % parte prae- 
antennali laevi impunctato. Antennae t evidenter propius ad apicem rostri quam ad 
oculos, % longe post medium rostrum insertae. Pronotum sat dense punctatum, antice 
fortiter angustatum, conico-truncatum. Elytra striata, striis parum fortiter punctatis, 
apicem versus compressa. Long, (rostr. incL), ^ majoris 21 mm., ^ minoris i2'5 mm. ; 
% 15 mm. 

Ill-developed males have the front femora more incrassate than large examples, 
tending to resemble the % in this respect, as also to some extent in the smoother 
rostrum. 

Hab. Hawaii, Maui. Various localities on Hawaii (2000 — 4000 ft.) ; Kohala, 
Kau, Puna; Haleakala on Maui. This species was given to me, soon after my arrival 
in the islands, by Mr G. C. Munro, who obtained specimens in the Kohala mountains, 
and I have named it after this excellent ornithologist. A fragment (elytra and sternum) 
found in the mountains of Molokai may belong to this species, but the pubescence is 
more deeply yellow. 



COLEOPTERA 151 

(2) Nesotocus nczuelli, sp. nov. 

Praecedenti cognatissimus. Differt statura majore, antennaruni scapi apice magis 
incrassato, funiculo densius longiusque piloso, articulo secundo pro primo magis elongate. 
Z- Long. 26 mm. 

The single t of this species I believe to be distinct from N. muiu-oi, and not 
merely a very large and finely developed example of that species. Besides the 
characters given above the apical joint of the fnnicnlns of the antennae is nearly equal 
to the basal portion of the club, which is clothed with sub-erect hairs (the apical portion 
being densely covered with appressed pubescence). In N. munroi the apical joint of 
the funiculus is very greatly shorter than the basal portion of the club. 

Hab. Maui. A single example has been taken by Bro. Matthias Newell in the 
lao valley. 

(3) Nesotoctis kauaiensis, sp. nov. 

Praecedentibus simillimus, niger, pubescentia flava ornatus. Differt ^ antennis 
haud propius ad apicem rostri quam ad oculos insertis, % rostri parte postantennali 
minus fortiter denseque punctata. Long. 13 — 16"5 mm. (Plate VIII. fig. 5.) 

The ^ of this species is very distinct from either of the preceding, by the different 
point of insertion of the antennae, this being nearly equidistant from the eyes and apex 
of the rostrum. The % is much more difficult to separate, but the basal portion of the 
rostrum appears to be smoother, and more finely and less closely punctured. In fresh 
examples the pubescence is decidedly more deeply yellow, and that on the elytra is 
rather more extensive. 

Hab. Kauai (4000 ft.). 



DvsoMMA, gen. nov. 

Corpus sat gracile, cylindrico-fusiforme, haud pubescens. Rostrum elongatum, 
leviter curvatum, pronoto longitudine subaequale, parte praeantennali basali parte sat 
latiore. Oculi parvi, subobsoleti. Antennae longe ante medium rostrum insertae, 
funiculi 7-articulati articulo basali elongato, secundo bis longiore, hoc tertio longiore, 
capitulo elongato-ovali. Pronotum magnum, elongatum, elytris latius, antice fortiter 
angustatum, vix constrictum. Uncus tibiarum anticarum apicalis validus. Tarsi brevi- 
ores, articulo tertio parvo, lobis brevissimis. Unguiculi graciles, breves. Coxae omnes 
late separatae, metasterno longo. Abdominis segmentum basale inter coxas posteriores 
antice rotundatim productum. Elytra angustiora, lateribus subparallelis. Scutellum 
patens. 



152 FAUNA HAWAIIENSIS 



(i) Dysomma sylvicola, sp. nov. 

Elongatus, sat gracilis, colore castaneo, subnitidus. Rostrum subrugoso-punctatum, 
setis brevissimis paucis vestitum. Pronotum dense aequaliter punctatum. Elytra 
pronoto angustiora, striata, striis confertim punctatis, interstitiis uniseriatim punctulatis. 
Long, (rostr. incl.) 5 mm. (Plate VIII. fig. 6.) 

Hab. Kauai. A single example taken in the mountains (4000 ft.). 



Heteramphus Sharp. 

(i) Heteramphus filiatni, sp. nov. 

Latus, niger, opacus, tarsis antennisque rufescentibus. Pronotum latum, antice 
angustatum, vix constrictum, creberrime punctatum, brevissime pubescens, dorso juxta 
basim haud, vel vix evidenter, impresso. Elytra seriatim punctata, interstitiis usque 
ad basim parcius sed distincte pallido-pubescentibus. Long, (rostr. incl.) 8 — 9 mm. 
(Plate VIII. fig. 7.) 

Closely allied to H. wollastoni Shp., but at once distinguished by the much more 
densely punctured thorax, which is clothed with a short minute pubescence, and lacks 
the basal impression of the allied species. The pale hairs on the interstices of the 
elytra are not confined to the apical portion, but extend to the extreme base. 

Hab. Oahu, in the mountains. Attached to the tree-fern ; only a few examples 
taken, the species being much more difficult to procure than H. wollastoni. 



(2) Heteramplms wollastoni Sharp. 

HeterampJms wollastoni Sharp, Tr. Dublin Soc. iii. 1885, p. 188. 

Hab. Oahu. Common in the mountains, but local. Lives at the bases of the 
leaves of Astelia veratroides. 



(3) Heteramphus foveatus Sharp. 

HeterampJms foveattis Sharp, /. c. 

Hab. Oahu. Common in the same localities as, and in company with, H. 
wollastoni. 



COLEOPTERA 153 



(4) Hctcranipluis lialcakalae, sp. nov. 

Piceus vel rufo-piceus, rostro, antennis pedibusque rufescentibus. Rostrum rugoso- 
punctatum, parte postantennali carinatim compressa. Pronotum subgrosse (antice 
quam postice minus dense) punctatum, sat elongatum, minus latum, dorso juxta basim 
leviter vel vix evidenter impressum. Elytra sat elongata, striata, striis parum fortiter 
nee confertim punctatis, parce usque ad basim pubescentia. Long, (rostr. incl.) 6 — 7 mm. 

The general appearance and form of this species is that of H. foveatiis Shp., but 
it is easily distinguished by the fact that it is more elongate, both the prothorax and 
elytra being narrower, and the former has only a shallow, sometimes hardly perceptible, 
impression near the base, and the punctures of the elytral striae are finer and more 
feeble. It is also very closely allied to the following species. 

Hab. Maui, Haleakala (5000 ft.) ; three examples ; taken from dead logs. 



(5) HcterainpJms /rater, sp. nov. 

Piceus vel rufo-piceus, rostro, antennis pedibusque plus minusve rufescentibus. 
Rostrum rugoso-punctatum, parte postantennali carinatim compressa. Pronotum grosse 
punctatum, postice juxta basim profunde foveatum. Elytra minus lata, usque ad basim 
parce pubescentia, striis minus fortiter punctatis. Long, (rostr. incl.) 6 — 7 mm. 

Somewhat intermediate between H. haleakalae and H. foveatns, the base of the 
prothorax having a strong round impression. It is a narrower and more elongate 
insect than H. foveatus, and has the striae of the elytra less strongly punctured. From 
H. haleakalae it differs in the strong impression on the prothorax, and in the fact that 
the shoulders of the elytra are capable of being more closely applied to the base of the 
prothorax. The two examples taken do not agree well together, the one being more 
bulky than the other, and having the thoracic puncturation coarser, while the punctures 
of the striae of the elytra are finer and less closely placed. The smaller example has no 
impression on the apical ventral segment of the hind-body, whereas in the larger this 
segment is distinctly impressed, as is also the case with all the individuals of the 
preceding species. 

Hab. Maui, Haleakala (5000 ft.) ; very rare. 



(6) HeterampJms molokaiensis, sp. nov. 

Piceus, antennis, rostro pedibusque plus minusve rufescentibus. Rostrum distincte 
punctatum, parte postantennali carinatim compressa. Pronotum angustum, elongatum, 
antice angustatum, vix grosse punctatum, parum evidenter pubescens. Elytra pronoto 



154 FAUNA HAWAIIENSIS 

paullo latiora, angustula, vix evidenter pubescentia, apicem versus rarissime setosa, striis 
obscurissime obsolete punctatis. Long, (rostr. incl.) 5 mm. (Plate VIII. fig. 8.) 

Most nearly allied to H. halcakalae, but at once distinguished by its much more 
slender and elongate form. 

Hab. Molokai mountains (4000 ft.). A single example taken out of wet moss in 
June 1893. 

(7) Hete7'ainplms cylindriais Sharp. 

Heteramphus cylindriats Sharp, /. c. p. 189. 

(Plate VIII. fig. 9 large form; fig. 10 small form.) 

Hab. Oahu ; common, at the bases of the leaves and in the stems of Astelia. 
Varies greatly in size, some examples being much larger and more robust than others. 



(8) Hcte7-amphus hirtellns Sharp. 

HeterainpJms hirtellus Sharp, /. c. 

Hab. Oahu. A single e.xample was taken by Mr Blackburn in the mountains 
near Honolulu. I have never met with this species. 

(9) HeterainpJms kauaiensis, sp. nov. 

Piceus, sat latus, nitidus, rostri parte basali, pronoto, elytrisque, dense pallide 
setosis. Rostrum apicem versus dilatatum, nitidum, et subtiliter punctatum. Pronotum 
sat latum, nitidum, dense grosseque punctatum, juxta basim obsolete impressum, linea 
dorsali laevi. Elytra brevia, nitida, striis grosse confertim punctatis, interstitiis setis 
longis flavescentibus vestitis. Long, (rostr. incl.) 5*5 mm. (Plate VIII. fig. 11.) 

This remarkable species is allied to H. Jiirtcllus Shp., the rostrum, viewed from 
in front, being widened at some distance before the insertion of the antennae on the 
basal side, owing to the long outstanding pterygia. Its form is much the same as in 
H. foveatus, but it might reasonably be considered as the type of another genus. 

Hab. Kauai. A single example taken on the high plateau. 

(10) Heteranipltus nivicola, sp. nov. 

Nigro-piceus, opacus (capite cum pronoto absente), elytris apices versus setas 
nonnullas ferentibus. Elytra distinctissime striata, striis remote nee grosse punctatis, 
interstitiis baud convexis subseriatim rugoso-punctatis. Metasternum grosse punctatum. 



COLEOPTERA 



155 



Abdominis segmentum basale subtus nitidum, grosse et remote punctatum, segmentum 
2"", 3"" et 4" rugoso-punctata, segmentum apicale densius fortiter distincte punctatum. 
Long. ?. 

The genus of the insect above described is not certain, since the head, prothorax 
and most of the appendages are wanting, but it is probably a true Hetcramphus. It 
is easily distinguished from any of the other species, by the sculpture of the elytra and 
the under-parts. The former are less wide at the base and comparatively more elongate 
than those of H. foveatits (than which the species is of larger size), the striae are very 
distinct and are wide apart, their punctures are somewhat fine, and especially on the 
striae towards the sides, remote from one another, the interstices are not the least 
convex, and are rather largely and rugosely and somewhat seriately punctured, so that 
when looked along from the apex they even appear somewhat grooved or concave. 
The humeral angles of the elytra are rounded off and indistinct. The abdomen beneath 
is shining, the basal segment remotely and very strongly punctured, the three following 
ones are rugosely punctate, the apical one strongly and closely, but less coarsely than 
the basal one. 

Hab. Maui. A single example in fragmentary condition was found near the 
summit of Haleakala. 



OoDEMAS Bohem. 

(i) Oodeinas olindae Blackburn. 

Oodemas olindae Blackburn, Ent. Mo. Mag. xvii. p. 199. 

(Plate VIII. fig. 12, ?.) 

Hab. Maui, Haleakala (5000 ft.) ; i ^ and i % taken. Two examples, now in the 
British Museum, taken by Mr Blackburn, are both $, and apart from sexual differences 
agree closely with my <y. The % taken by me is less shining, the prothorax entirely 
dull, and strongly punctured, but there is little doubt that it belongs to the same species. 
The ^ differs from the $ in the shorter rostrum, and the very large second joint of the 
anterior and intermediate tarsi, which are hardly less wide than the third joint. 



(2) Oodeinas longirostre, sp. nov. 

Nigrum, aenescens, elongato-ovale, nitidum. Rostrum longissimum, valde curva- 
tum, apice fortiter dilatato, basali parte gracillima, subtiliter dense punctatum. Oculi 
fortiter prominentes. Prothorax nitidus, sat elongatus, subtiliter punctatus. Elytra 

F. H. II. 21 



156 FAUNA HAWAIIENSIS 

nitida, basi obscure marginata, antice obscure striata, striis grossius punctatis, interstitiis 
crebre punctulatis, postice fortiter convexis. $. Long, (cum rostro) 7 mm. (Plate VIII. 
fig. 13, I? ; I3«, anterior tarsus of ^.) 

Very distinct from any other species ; most nearly allied to O. olindae. The 
excessively long slender rostrum, which is much more strongly curved, will at once 
distinguish it from that species. The two basal joints of the funiculus are much 
the same in both species. 

Hab. Kauai. A single % taken in the mountains (4000 ft.). 



(3) O ode mas dilatatipes, sp. nov. 

Nigroaeneum, elongatum, nitidum. Rostrum nitidum, subtiliter sat dense puncta- 
tum, apice dilatato. Oculi fortiter prominentes. Antennarum articulus secundus et 
tertius fortiter elongati, subaequales. Pronotum latum, antice fortiter angustatum, 
subtiliter punctatum. Elytra seriatim (subgrosse) punctata, interstitiis convexis, sub- 
tiliter punctatis. Tarsorum ^ intermed. et antic, articulus secundus aeque latus ac 
tertius. Z- Long. 5 mm. 

Black, slightly aeneous, shining. The Z (the only sex obtained), has the rostrum 
long, strongly dilated on its apical portion, the surface shining, finely and densely 
punctured. The eyes are very strongly convex, and prominent. Antennae with the 
first joint of the funiculus long, the following hardly longer. The prothorax is wide, 
strongly narrowed in front, and finely punctured. Elytra shining, with rows of 
moderately large punctures, the interstices slightly and narrowly raised, or convex, and 
finely punctured. The hind portion of the elytra is of the abrupt form usual in the 
section, and the apical segments of the hind-body beneath are inclined to the basal part. 
The femora are extremely stout, and the second joint of the front and intermediate 
tarsi very wide. 

Hab. Oahu, Waianae mountains. A single t only taken. Differs from the ^ of 
O. olindae Blk., in the shorter rostrum, the coarser puncturation of the striae of the 
elytra, which are not distinctly margined at the base, etc. 



(4) Oodemas nivicola Blackburn. 

Oodemas nivicola Blackb., Ann. Soc. Ent. Belg. xxi. p. 75. 

Hab. Maui, Haleakala (4000 ft. to the summit), outside the forest. Common 
and very variable in size, sculpture &c. Sometimes brightly shining, often quite dull. 



COLEOPTERA 157 

(5) Oodcmas costatiim, sp. nov. 

Ovatum, robustum, nitide aeneum. Rostrum nitidum, subtiliter punctatum, apice 
subdilatato. Antennarum articulus secundus et tertius fortissime elongati, fere aequi- 
longi. Elytra seriatim punctata, interstitiis punctulatis, a basi usque ad apicem elytrorum 
distinctissime costatis. Tarsorum antic, et intermed. (j) articulus secundus maximus, 
articulo tertio baud minus latus. ^. Long. 5 '5 mm. 

Ovate, shining, bronzy black. Rostrum shining, finely punctured, dilated at the 
apex. Eyes convex, subprominent. Antennae with the first two joints of the funiculus 
very long, subequal. Prothorax shining finely and feebly punctured. Elytra striate, 
the striae strongly punctured, the interstices sharply raised from base to apex, giving 
the elytra a ribbed appearance, and finely punctured. Second joint of the anterior and 
intermediate tarsi of the ^ very large, about as wide as the apical width of the lobate 
third joint, and rather longer than wide. 

Hab. Kauai Mountains (4000 ft.). A very distinct species, of which only a 
single $ was taken. 

(6) Oodemas longicorne, sp. nov. 

Ovatum, robustum, aeneum, nitidissimum. Rostrum sat longum, apice dilatato, 
nitidum, subtiliter dense punctatum. Antennae longae, funiculi articulo primo fortiter 
elongato, secundo hoc vel longiore. Prothorax latus, nitidus, subtiliter punctatus. 
Tarsorum articulus secundus anticorum et intermediorum aeque latus ac tertius, et 
hoc multo longior. Elytra grossius seriatim antice punctata, interstitiis obscure costato- 
elevatis. Z- Long. 5 mm. 

This species is closely allied to O. costahtm, but is very distinct by the sculpture 
of the elytra, the interstices only showing a faint tendency to the costate form of that 
species. 

Hab. Kauai. Four examples (all $), taken on the high plateau. 

(7) Oodemas ptinctulatissimtim , sp. nov. 

Nigroaeneum, nitidum, ovale, robustum. Antennae testaceae, graciles, sat elon- 
gatae, articulo secundo brevi, tertio minus fortiter elongato, sed .secundo longiore. 
Rostrum nitidum, apice dilatato, fortiter deflexo, subtilissime punctatum. Oculi parum 
prominentes. Prothorax latus, nitidus, subtilissime (vix evidenter) punctatus. Elytra 
nitida, antice vix evidenter, postice paullo distinctius, striata, dense punctulata, apice 
fere recurvo. Pedes rufotestacei ; tarsorum {$) anticorum et intermediorum articulus 
secundus dilatatus, tertio vix minus latus. $. Long. 3 '5 mm. 

21—2 



158 FAUNA HAWAIIENSIS 

Remarkable in its group for the unusually short second joint of the funiculus which 
though evidently longer than the short first joint is less elongate than in the allied 
species, and for the dense puncturation of the elytra, the punctures on the striae (which 
are hardly perceptible in front) being very little larger than those on the interstices. 

Hab. Oahu, Waianae mountains; i ^ taken in April 1892. 



(8) Oodenias molokaiense, sp. nov. 

Ovatum, nigroaeneum, nitidum. Rostrum dense subtilius punctatum, apice dilatato, 
Oculi magni, prominuli. Antennarum articulus tertius fortissime elongatus, secundo 
fere bis longior. Elytra minus fortiter seriatim punctata. Tarsorum (^) anticorum 
articulus secundus permagnus, aeque latus ac tertius. Long. 3'5 — 4*5 mm. 

(Plate VIII. fig. 14, ,? ; fig. \\a, anterior tarsus of ^.) 

Ovate, aeneous, more or less shining, generally brightly so, of more or less robust 
form. The rostrum is finely and closely punctured, its apical portion evidently dilated. 
The eyes are large and somewhat prominent. Antennae long, slender, the second 
joint of the funiculus very elongate, twice, or nearly twice, as long as the first. 
Prothorax broad, much narrowed in front, finely punctured, obsoletely so in some 
examples. Elytra with rows of fine punctures, sometimes with scarcely perceptible 
striation in front, in others with evident striae, the interstices very finely, subobsoletely 
punctured. Posteriorly the elytra are vertical, almost recurved, especially in the ^. 

Second joint of the front and intermediate tarsi in the <? very large, as wide as 
the lobate third joint, and rather longer. Apical ventral segments of the hind-body in 
a plane inclined to that of the basal segments. Rostrum shorter than that of the $. 

Two examples (<^, %) from Lanai have the elytra more strongly striated, the 
punctures on the striae rather larger, and the interstitial puncturation rather more 
developed. 

Hab. Molokai (3000 — 4500 ft.); Lanai (2000 ft.). Rare. M}^\&di to O . pzmctula- 
hsshnum, but readily distinguished by the longer second joint of the funiculus, and 
the different puncturation of the elytra. The examples from Lanai might almost pass 
as a distinct species. 

(9) Oodenias chrysodortim, sp. nov. 

Castaneum, nitidum, conspicue aureo-micans, ovale, elytrorum lateribus bisinuatis. 
Rostrum longum, apice dilatato, rugoso-punctatum, saepe longitudinaliter carinatum. 
Oculi perdepressi. Pronotum baud transversum, antice fortiter angustatum. Elytra 
(antice saltern) obscure striata, striis saepe impunctatis, vel parce subtilissime punctatis. 
^ tarsorum anticorum articulus secundus fere aeque longus ac latus, articulus tertius 



COLEOPTERA 159 

lobis magnis. Antennae crassiusculae, articulo 2° et 3° aut subaequalibus aut hoc 

longiore. t %. Long. 375 — 5-2 mm. 

(Plate VIII. fig. 15, $ ; fig. 15a, anterior tarsus of ^; fig. I'^b, of ?.) 

This species is readily distinguished by its bright golden-brown colour, the long 

rostrum (especially in the $), which has the apex slightly but evidently widened, and 

the feeble sculpture of the elytra. 

Hab. Maui, Haleakala (5000 ft.) ; lives in the stems of Rubtis viacraei. 

(10) Oodcnias briinneum, sp. nov. 

Anguste ovale, nitidum, brunneum, aureo-micans. Rostrum nequaquam dilatatum, 
rugoso-punctatum. Oculi convexi, haud fortiter depressi. Antennae crassiusculae, 
funiculi articulo secundo et tertio brevissimis, hoc illo longiore. Pronotum nitidum, 
subelongatum, subtiliter nee dense punctatum. Tarsorum ^ anticorum articulus 
secundus latior quam longior, tertio evidenter angustior. Elytra nitida antice vix 
evidenter striata, seriatim subtiliter punctata, interstitiis etiam punctatis. ^. Long. 
3-25 mm. 

This species somewhat resembles the preceding in colour, but the rostrum, which 
is not widened on the apical portion, the more convex eyes, the elytra much less 
strongly rounded behind the shoulders, and their more distinct puncturation (the 
punctures of the series are slightly larger than those of the interstices), will easily 
distinguish it. 

Hab. Molokai, mountains (3000 ft.); i <?. 

(11) Oodcmas aipreum, sp. nov. 

Ovale, plus minusve nitidum, cupreum. Rostrum sat longum, apice haud dilatato, 
subtilissime punctatum. Antennarum funiculi articuli 2 basales parum longi, sub- 
aequales. Pronotum subelongatum, antice angustatum. Elytra subtiliter striata, 
seriatim, nee fortiter punctata, interstitiis sat crebre punctulatis. Long. 4 — 5 mm. 

Elongate-oval, more or less shining, the elytra reddish or piceous, and of a distinct 
copper colour. Legs and often the antennae reddish. Rostrum rather long, not 
widened towards the apex, finely and more or less sparsely punctured. The two basal 
joints of the funiculus of the antennae are short, stout and of about equal length. 
Prothorax finely punctured, rather long and much narrowed in front. Elytra more or 
less feebly striate, the punctures of the striae distinct, but not coarse, the interstices 
with a much finer but distinct and more or less dense puncturation. The lobes of the 
front tarsi are small in all the specimens examined, but it is uncertain whether the t sex 
is represented. 

Hab. Maui, Haleakala (5000 — 6000 ft.) ; rare. 



i6o FAUNA HAWAIIENSIS 

(12) Oodemas crassicorne Blackburn, 

Oode^nas crassicorne Blackb. Tr. Dublin Soc. iii. 1885, p. 184. 
The sexual distinctions are important in this species, the second joint of the front 
tarsi of the ^ being much wider than that of the %. 

Hab. Lanai Mountains (2000 — 3000 ft.). Not rare. 

{^i^ Oodemas haleakalae, sp. nov. 

Elongato-ovale, nitidum, aenescens. Rostrum rugoso-punctatum ; oculi parum 
convexi. Antennarum articulo secundo brevi, tertio elongate. Pronotum baud trans- 
versum, antice angustatum. Elytra antice levissime vel vix striata, lateribus bisinuatis, 
striis subtiliter remote punctatis, interstitiis saepe crebre punctulatis, postice (prae- 
cipue ?) sat convexis. Tarsorum ant. ^ articulus secundus transversus, tertius hoc 
multo latior, lobis magnis. Long. 3 — 4*5 mm. 

This species is closely allied to O. crassicorne, Blk. It varies greatly in size, small 
males being only about half the bulk of the female, and in sculpture, the puncturation 
of the elytra being much more developed in some examples than in others. It may be 
distinguished from the allied species by the longer rostrum and less convex eyes, and 
the former is generally more rugosely punctured, especially in the female. The rostrum 
is generally more or less evidently widened towards the apex, and is often distinctly 
carinate down the middle. The interstitial puncturation of the elytra is as a rule hardly 
less developed than that of the striae. Towards the apex (especially in the $) the 
interstices become rather strongly convex, the fourth, fifth and sixth terminating 
somewhat abruptly at the same point, a character which will distinguish this species 
from some others of very similar appearance (e.g. O. iarduni, Blk.). 

Hab. Maui, Haleakala (5000 — 6000 ft.) ; common. 



(14) Oodemas montamim, sp. nov. 

Nigroaeneum, nitidum, ovale. Antennae crassae, scapo brevi, funiculi articulo 
primo elongato, crasso, secundo subaequali. Rostrum nitidum, subfortiter punctatum, 
apice leviter dilatato, fortius decurvato. Oculi convexi et prominuli. Pronotum sat 
longum, subtilissime punctatum, antice fortiter angustatum. Elytra nitida, seriatim nee 
grosse punctata, vix evidenter striata, interstitiis subtiliter minus dense punctatis. 
(Sexus ?) Long, vix 4 mm. 

This species very closely resembles O. tardum Blackb., but the very stout 



COLEOPTERA i6i 

antennae, the first and second joints of the funiculus being elongate, subequal, and 
unusually stout, and the less punctured interstices of the elytra easily distinguish it. 
The single example taken is probably a ^. The second joint of the front tarsi is short, 
subrotundate, and much less wide than the bilobate third joint. 

Hak. Kauai. A single example taken on the high plateau in August, 1S96. 

(15) Oodevias tardum Blackburn. 

Oodemas tardum Blackb. Tr. Dublin Soc. in. 1885, p. 184. 

Several specimens taken on Haleakala I refer to this species, which in most 
respects very closely resembles O. haleakalae (q. v.). It would appear to be somewhat 
variable, but the material taken is quite insufficient for a proper understanding of the 
species. 

Hab. Maui, Haleakala (4000 — 5000 ft.). Rare. 

(16) Oodemas obsairum Blackburn. 

Oodemas obsairum Blackb. Ann. Soc. Ent. Belg. xxi. p. 75. 
van substrictum Blackb. Ent. Mo. Mag. xvii. p. 200. 

The characters afforded by the front tarsi of the Z are important for distinguishing 
this species. The lobate third joint is unusually small and the lobes short, the second 
joint is rather wide. 

Hab. Maui, Haleakala (4000 — 5000 ft.) ; not rare. 

(17) Oodemas aequale Blackburn. 

Oodemas aequale Blackb. Tr. Dublin Soc. in. 1885, p. 184. 
Hab. Lanai (2000 — 3000 ft.), where it is not rare. 

(18) Oodemas apionoides, sp. nov. 

Angustum, elongato-ovale, nigroaeneum, plus minusve nitidum. Rostrum puncta- 
tum, parte apicali levissime dilatata, prae insertione antennarum sat evidenter decurvata. 
Antennarum articulus secundus brevis, robustus, tertius huic subaequalis, vel paullo 
longior. Pronotum elongatum antice sensim angustatum. Elytra subtilissime striata, 
striis grossius remote punctatis, interstitiis parce (saepe subobsolete) punctatis. Tarsorum 
anticorum (^) articulus secundus brevis, rotundus, tertius hoc multo latior, lobis parum 
magnis. Z%- Long. 3—3-5 mm. 



1 62 FAUNA HAWAIIENSIS 

A rather variable species, the punctures on the striae of the elytra being decidedly 
coarser in some examples than others, and sometimes also more numerous, and there- 
fore less remote. The elytra are often of a bright bronzy colour. The t is shorter 
than the ?, and readily distinguished by the less elongated rostrum. The striation of 
the elytra is always very fine and sometimes hardly discernible in front. The widening 
of the rostrum towards the apex is sometimes hardly perceptible, and its puncturation 
is variable, being decidedly rugose in some examples. 

Hab. Kauai Mountains (3000 — 4000 ft.). Not common. 



(19) Oodemas affinc, sp. nov. 

Praecedenti cognatissimum, rostro breviore, elytris fortius et confertius seriatim 
punctatis distinguendum. $. Long. 3"25 mm. 

Closely allied to O. apionoides. The rostrum is closely punctured, the apex some- 
what strongly curved downwards, and it is evidently wider than that of the preceding 
species. The first joint of the funiculus of the antennae is very short and stout, the 
second rather longer than this. The prothorax is dull and remotely punctured, its 
sides slightly convergent from the base to front. The elytra are shining, brassy, 
distinctly striate ; the striae coarsely punctured, and not very remotely. The interstices 
are distinctly but sparsely punctured. Second joint of the front tarsi (^) very small, 
transverse, lobes ot the third small. 

Hab. Kauai, Makaweli (2000 ft.) ; i t taken. 



(20) Oodc)}ias parallchim, sp. nov. 

Laete aeneum, nitidissimum, angustissimum. Antennarum articulus secundus 
brevissimus, tertius fortiter elongatus secundo bis longior. Pronotum nitidum, antice 
minus fortiter angustatum, subfortiter punctatum. Elytra nitidissima, subtilissime 
striata, striis sat fortiter nee confertim punctatis, interstitiis parce distincte punctulatis. 
Tarsorum anticorum ^ articulus secundus brevis transversus, tertius hoc multo latior, 
lobis magnis. ^. Long, circa 3 mm. 

This is a very distinct species. Its bright bronzy colour, very narrow form, 
comparatively strongly punctured thorax, long second joint of the funiculus of the 
antennae, &c., &c., readily distinguish it. 

Hab. Oahu. A single ^ taken on Kaala (Waianae Mountains) in December, 
1892. 



COLEOPTERA 163 



(21) Oodemas graciliforme, sp. no v. 

Angustum, elongato-ovale, nitide aeneum. Rostrum subnitidum, subtiliter punc- 
tatum, apice levissime dilatato, evidenter decurvato. Antennarum articulus secundus et 
tertius elongati, subaequales. Pronotum hand transversum, antice fortius angustatum. 
Elytra nitida, levissime striata, striis remote nee fortiter punctatis, interstitiis baud 
crebre punctulatis, apicem versus elytrorum plus minusve convexis. Tarsorum ^ 
anticorum articulus secundus parvus, subrotundus, quam dimidia pars tertii vix latior, 
hujus lobis baud magnis. Z%- Long. 375 — 4-5 mm. (Plate VIII. fig. 16.) 

An elongate very narrow species of moderate size and bright bronze colour. The 
rostrum is slightly widened towards the apex, and bent downwards from about the line 
of insertion of the antennae. The first and second joints of the funiculus are elongate, 
and subequal. The prothorax is generally shining, and varies considerably in its 
puncturation, which is closer and stronger in some examples than others ; its form is 
rather long and narrow, and it is much narrowed in front. The elytra are narrow, 
shining, bright brass-coloured or golden, with very fine but evident striae, which are a 
little deeper posteriorly, where the interstices are more or less convex. The punctures 
on the striae are not at all coarse, and are remote ; the interstitial puncturation is much 
finer than these, and not dense. 

In the t the second joint of the front tarsi is small, about as long as wide, half the 
width of the lobate third joint, which is transverse, and not largely developed. 

This species is distinct by the very narrow elongate form, bright bronzy colour, the 
subequal first and second joints of the funiculus of the antennae, fine striation of the 
elytra, and the fine puncturation of the striae. 

Hab. Kauai, Halemanu ; rare. 



(22) Oodemas ranmloruin, sp. nov. 

Nigroaeneum, nitidum, ovale. Rostrum baud dense subtiliter punctatum, apice 
levissime dilatato, decurvato. Antennarum articulus secundus brevis, robustus, tertius 
gracilis, elongatus. Pronotum opacum, subtilissime, nee dense, punctatum, antice an- 
gustatum. Elytra nitidissima, plus minusve cupreo-nitentia, levissime striata, striis 
subtiliter punctatis, interstitiis minus dense subtilissime punctulatis. Long. 3 — 375 mm. 

The two examples of this small elongate-oval species were obtained from the pith- 
cavity of a dry twig, resting side by side. It is probable that they may be the sexes of 
one species, although they differ somewhat in shape &c. That which is probably the $ 
has the prothorax slightly shorter and wider, and of a dull brassy-green colour. The 

F. H. II. 22 



1 64 FAUNA HAWAIIENSIS 

elytra are black, with a copper-coloured metallic tint. The second joint of the anterior 
tarsi is small, the lobes of the following joint rather large. In the other example, both 
thorax and elytra are of a very marked copper colour, the former is rather longer, and 
the sides of the latter less strongly rounded. 

Hab. Oahu. Mountains near Honolulu (3000 ft.) ; two examples taken. 



(23) Oodemas leiothorax, sp. nov. 

Nigroaeneum, nitidum, ovale. Rostrum c/ breve, ? sat longum, apice baud dilatato, 
distincte punctatum. Antennarum articulus secundus elongatus, tertio paullo {i) vel 
multo (?) longior. Pronotum nitidum, subtilissime (vix evidenter) punctatum. Elytra 
nitida, levissime (vix evidenter) striata, striis subtilissime remote punctatis. Tarsorum 
anticorum c/* articulus secundus brevis, subrotundus, tertius lobis parvis. Z%- Long. 
3-5 mm. (Plate VIII. fig. 17). 

Apparently a very distinct species, by its antennae, the very feeble puncturation of 
the thorax, and the delicacy of the series of punctures on the elytra. The metallic tint 
of the latter is bronze-coloured and very distinct. 

Hab. Kauai. High plateau ; 2 ^, i ? taken. 



(24) Oodanas fiexirostre, sp. nov. 

Ovale, nigroaeneum, nitidum. Rostrum fortiter punctatum, ante medium fortius 
decurvatum, apice baud dilatato. Antennarum articulus secundus et tertius subaequi- 
longi. Pronotum nitidum, sat distincte punctatum. Elytra nitida, levissime striata, 
striis remote parum fortiter punctatis, interstitiis subtiliter, nee dense, punctatis. Tarso- 
rum anticorum lobi parvi. ^ (?). Long. 3-5 mm. 

Very similar to O. leiothorax in most respects, but distinguished by the more 
strongly curved rostrum, the much more distinct puncturation of the prothorax, and the 
less fine punctures of the elytra. 

Hab. Kauai (4000 ft.). One example. 



(25) Oodemas dubiosuni, sp, nov. 

Nigroaeneum, ovatum, elytris (j) subnitidis. Rostrum (saltern in media parte) 
parce punctatum, apice haud dilatato. Antennae graciliores, articulo funiculi primo et 
secundo subaequilongis [$), vel hoc longiore ($). Pronotum latum antice fortius angus- 
tatum, subtiliter punctatum. Elytra striata, striis subfortiter punctatis, interstitiis dis- 
tincte punctulatis. Tarsorum anticorum $ lobi parvi. Long, vix 4 — 4^5 mm. 



COLEOPTERA 165 

Three examples taken in the same locality and at the same time are referred to 
this species, although they do not agree very closely in some respects. I have regarded 
the two smaller, and partly shining, examples as t. the larger, which is entirely dull, as 
the ?. 

The rostrum is narrow, the apex not at all dilated, the puncturation sparse 
throughout, or at least down the centre. The antennae are slender, the first and 
second joints of the funiculus in the ^ are not very elongate, subequal in length, or with 
the second very slightly the longer. In the % the difference between them is more 
pronounced, the first being evidently less long than the second. The prothorax is 
wide, dull, or at least not brightly shining, and finely punctured. Elytra somewhat 
shining in the J^, quite dull in the %, lightly striate in the former, much more distinctly 
so in the latter, in which the punctures are decidedly coarse, more so than in the t- 
The lobate third joint of the anterior tarsi is very small for the size of the species. The 
metallic colour is more pronounced on the elytra than on the rest of the insect, and is 
bronzy. The general form is rather robust, especially the %. 

Hab. Kauai mountains (Makaweli, 3000ft.). 



(26) Oodemas sh^iatuin, sp. nov. 

Aeneum, elongato-ovale, nitidum. Rostrum subnitidum, crasse punctatum, apice 
haud dilatato. Antennae crassiusculae, funiculi articulo primo secundo evidenter 
breviore. Pronotum sat latum, dense punctatum. Elytra nitida, basi submarginata, 
fortissime striata, striis confertissime grosse punctatis, interstitiis convexis, conspicue 
punctulatis. Tarsorum anticorum lobi sat magni. (Ansexus?) Long. 4-5 mm. (Plate 
VIII. fig. 26.) 

This is a most distinct species of a bright bronzy colour and of elegant form, the 
elytra narrowing very gradually to the apex at a distance not far from their base. 
Their strong striation, with coarse close punctures, and conspicuous interstitial punctura- 
tion, together with the shape of the insect, readily distinguishes it from all others. 

Hab. Kauai. A single example taken in the mountains behind Lihue (3000 ft.). 

(27) Oodemas piincticolle, sp. nov. 

Nigroaeneum, nitidum, oblongo-ovatum. Rostrum strigoso-punctatum, apice haud 
dilatato. Oculi convexi. Antennarum articulus secundus brevis, tertius elongatus, illo 
multo longior. Pronotum sat longum, crebre distincteque punctatum. Elytra nitida, 
seriatim nee fortiter punctata, postice fere verticalia, apicibus late rotundatis, striis 
obsoletis, interstitiis crebre conspicueque punctulatis. Tarsorum anticorum articulus 
bilobatus parvus. (An sexus ?) Long. 4 mm. 

22 — 2 



1 66 FAUNA HAWAIIENSIS 

Apparently a very distinct species. The elytra are much more parallel-sided than 
is usual in the genus (as in O. pulchrum, &c.), and are comparatively widely rounded at 
the apex. The interstitial puncturation is very distinct, and the traces of striation are 
very feeble even towards the apex. The metallic colouring is not very pronounced. 

Hab. Kauai, Halemanu. One example taken. 



(28) Oodemas purpurascens, sp. nov. 

Robustum, oblongo-ovatum, nitidum. Rostrum rugoso-punctatum, apice baud 
dilatato. Antennarum articulus secundus brevis, tertius hoc multo longior. Pronotum 
sat longum, subtiliter densius punctatum. Elytra nitida, metallescentia, parum distincte 
striata, seriatim grossius foveolatim-punctata, interstitiis plus minusve, (nonnunquam 
crebre), punctulatis. Tarsorum anticorum Z articulus tertius parvus. J $. Long. 

4— 5"25 mm. 

The head and thorax are black with little or no metallic colouring, the elytra are 
very distinctly metallic, of a dark colour inclining to purple. The interstices are very 
faintly, sometimes hardly perceptibly, convex, their puncturation in the single ^ is faintly 
impressed and not conspicuous as in the two females examined, but of the latter one has 
the interstitial puncturation considerably more strongly developed than the other. The 
^ is much smaller than either of the females. The species appears to be most nearly 
allied to O. puncticolle, from which it differs in the colour, in its more robust form, 
coarser punctures of the elytra, which are less vertical behind, and more pointed at the 
apex, &c. 

Hab. Kauai (4000 ft.), 2 % ; near Makaweli (3000 ft.), i t- 



(29) Oodemas pulchrum, sp. nov. 

Elongatum, oblongo-ovatum, nitidum, laete aenescens. Rostrum Z breve, % sat 
elongatum, rugoso-punctatum. Antennarum articulus secundus (praecipue $) fortiter 
elongatus, tertius hoc longior. Elytra grosse seriatim foveolato-punctata, interstitiis 
apicem versus subacute elevatis. Tarsorum anticorum Z articulus secundus brevis, 
baud latior quam pars dimidia art. tertii. ^ %. Long. 475 — 6 mm. (Plate VI I L 
fig. 18.) 

Shining, elongate, oblong ovate, prothorax bronze-coloured, elytra bright metallic 
green. Rostrum very short in the $, considerably longer in the female, rugosely punc- 
tured. First joint of the funiculus of the antennae elongate, very strongly so in the ?, 
the second joint generally still longer, but sometimes the two are nearly equal in length. 
Prothorax shining, much narrowed in front, not very wide at the base. Elytra very 
coarsely punctured, their surface brightly shining, the sides but little rounded, giving 



COLEOPTERA 167 

the insect a more parallel-sided appearance than is usual in the genus ; the striae on the 
apical portion of the elytra are very deeply impressed, so that the interstices are narrow, 
strongly convex, and almost sharp. In the ^ the second joint of the front tarsi is small, 
short and transverse, about half as wide as the bilobate third joint. 

Hab. Kauai (4000 ft.); rare. This is an extremely distinct species by its bright 
metallic colour, its extremely coarse elytral punctures, and its general form. In the 
latter respect the following species closely resembles it. 

(30) Oodcvias oblongum, sp. nov. 

Praecedenti affine, nigrum, nitidum, parum aenescens, oblongo-ovatum. Rostrum 
opacum, rugoso-punctatum. Antennarum articulus tertius secundo longior. Pronotum 
nitidum, subtiliter punctatum antice angustatum. Elytra seriatim fortiter punctata, 
postice striis quam praecedentis minus fortiter impressis. J %. Long. 575 — 6'5 mm. 
(Plate VIII. fig. 19.) 

Similar in form to the preceding, black, and only slightly aeneous. The great 
difference in colour separates the two at once, as well as the finer puncturation of the 
elytra of the present species, the punctures though less coarse being more definite in 
form, and the striae towards the apex are less deeply impressed, the interstices conse- 
quently standing out less sharply. The interstitial puncturation is fine, more or less 
dense, and very distinct. 

Hab. Kauai (4000 ft.); rare. 

(31) Oodeiiias grande, sp. nov. 

Nigrum, robustum, ovatum, parum distincte aenescens. Antennarum articulus 
secundus et tertius sat elongati, subaequales, vel hoc longiore. Rostrum rugoso- 
punctatum. Pronotum latum, nitidum, subtiliter punctatum. Elytra nitida, seriatim et 
confertim fortiter punctata, interstitiis dense et distinctissime punctulatis, postice haud 
convexim elevatis, striis vix videndis. %. Long. 7 mm. (Plate VIII. fig. 20.) 

A large robust elongate-oval species, black, with a slight aeneous tint. The 
rostrum is rugosely punctured, rather long in the ?, the sides nearly parallel, or slightly 
converging from the base. The two basal joints of the funiculus are both long, sub- 
equal, or the second the longer. The prothorax is very wide at the base, strongly 
narrowed in front, the surface shining, finely but distinctly punctured. Elytra shining, 
the striation excessively feeble even to the apex, the punctures of the series moderately 
large and close, the interstices densely and conspicuously punctured. 

Hab. Kauai (4000ft.); 2 % taken. A very distinct species distinguished by its 
large size, elongate basal joints of the funiculus, the rows of large punctures on the 
elytra and dense and distinct interstitial puncturation. 



1 68 FAUNA HAWAIIENSIS 

(32) Oodemas corticis, sp. nov. 

Nigroaeneum, plus minusve nitidum, elongato-ovatum, sat robustum. Rostrum 
crebre punctatum, apice baud dilatato. Oculi convexi. Antennarum funiculi articulus 
secundus primo longior. Pronotum latum, antice fortiter angustatum. Elytra crebre 
punctulata, lateribus bisinuatis, punctis, quae majora sunt, seriatim dispositis, striis 
parum distinctis, vel obsoletis. Lobi tarsorum anticorum parum magni. ^ %. Long. 
5 — 7 mm. (Plate VIII. fig. 2[.) 

This species of which I have examined a great number of examples, is very 
variable in almost every character. The rows of larger punctures on the elytra are 
sometimes very distinct, with the punctures themselves closely set, sometimes much less 
so, and the punctures remote from one another, or even for the most part obsolete. 
These punctures are shallow and by no means coarse, but larger than those of the 
interstices, which are nearly always well developed and distinct. Most examples have 
the surface shining, some brilliantly so, others are nearly dull. In spite of the variation 
exhibited, the species by its large size, and general form, &c., is one of those most easily 
recognized on a casual inspection. 

Hab. Lanai, Molokai mountains, and Maui on Haleakala. Very abundant under 
the bark of trees. 



^'^'^ Oodemas pachysoma^ sp. nov. 

Ovatum, robustum, nigroaeneum, plerumque plus minusve nitidum. Rostrum 
distincte punctatum, apice baud dilatato. Oculi convexi. Antennarum articulus secun- 
dus et tertius subaequilongi. Pronotum latum, antice fortiter angustatum, subtiliter 
punctatum. Elytra levissime vel vix striata, seriatim vix fortiter punctata, interstitiis 
distincte subtiliter punctatis. Tarsorum anticorum utriusque sexus lobi parum magni. 
t %. Long. 4-5 — 6 mm. (Plate VIII. fig. 22.) 

A rather distinct looking species, of robust form, and not brightly aeneous. The 
rostrum in the ^ is rather short, and wide at the base, becoming narrower towards the 
apex. The elytra are (at least in the <J) wide near the base, the sides being strongly 
rounded from the shoulders. The sculpture is variable, consisting of rows of larger 
punctures which are sometimes closely placed (i.e. the punctures in each row), but in 
other examples are decidedly irregular. The interstitial puncturation is much more 
dense in some examples than in others, but it is nearly always conspicuous. The lobate 
third joint of the anterior tarsi is unusually small for the size of the species in both 
sexes. The general surface is more or less shining, but there is considerable variation 
in this respect. 

Hab. Kauai mountains (4000 ft.). 



COLEOPTERA 169 

(34) Oodcnias acohsoiiia, sp. nov. 

Ovatum, nitidum, elytris $ aenescentibus, $ subcupreo-nitentibus. Rostrum $ 
brevius, nitidum, subtiliter punctatum, apice baud dilatato. Antennae graciles, articulis 
2 basalibus funiculi elongatis, subaequilongis. Oculi convexi. Pronotum nitidum, 
subtiliter punctatum, antice fortiter angustatum. Elytra nitida, crebre punctulata, 
punctis quae majora sunt, seriatim dispositis, striis levissimis, vel obsoletis. Tarsorum 
anticorum lobi mediocres. $ %. Long. 4 — 4'5 mm. 

This species is closely allied to the two preceding, but is a smaller and more 
shining insect. The puncturation of the elytra shows some variation, the punctures 
which are disposed in series being considerably coarser in some examples than in 
others, the density of the interstitial puncturation also varies. 

Hab. Kauai, a few examples taken in the mountains at an elevation of 4000 ft. 

(35) Oodevias aenescens Bohem. 

Oodemas aenescens Boh., Eug. Res. p. 138, t. 2, f. 6. 

(Plate VIII. fig. 23.) 

Hab. Oahu and Lanai mountains. Remarkable for the extremely coarse punctu- 
ration of the base of the abdomen beneath. The length of the first and second joints of 
the funiculus of the antennae is a little variable, sometimes one and sometimes the other 
being slightly the longer. 

(36) Oodemas angustiim Blackburn. 

Oodemas angustum Blackb., Ann. Soc. Ent. Belg. xxi. p. 75. 
Hab. Oahu, Waianae mountains (Blackburn). 

{^l"]) Oodemas halticoides Blackburn. 

Oodemas halticoides Blackb., Ent. Mo. Mag. xiv. p. 5. 
Hab. Oahu mountains, 2000 — 3000 ft. (Blackburn). 

(38) Oodemas robustiim Blackburn. 

Oodevias robustum Blackb., Ann. Soc. Ent. Belg. xxi. p. 75. 
Hab. Oahu, Waianae mountains (Blackburn). 



170 FAUNA HAWAIIENSIS 

(39) Oodemas insulare Blackburn. 

Oodemas insulare Blackb., Ann. Soc. Ent. Belg. xxi. p. 74. 
Hab. Oahu (Blackburn). 

(40) Oodemas sculpturatum Blackburn. 

Oodemas sculpfurahim Blackb., Ann. Soc. Ent. Belg. xxi. p. 74. 

(Plate VIII. fig. 24.) 

Hab. Maui, Haleakala (4000 — 5000 ft.). Not rare. 

(41) Oodemas nitidissinuim, sp. nov. 

Nigroaeneum, nitidissimum, ovale. Rostrum subtiliter crebre punctatum, apice 
haud dilatato. Antennarum articulus secundus fortiter elongatus, tertio bis longior. 
Pronotum nitidissimum, subtiliter subobsolete punctatum, latum, antice fortiter angus- 
tatum. Elytra levissime (parum distincte) striata, striis remote subtiliter punctatis, 
interstitiis conspicue punctulatis. Long. 3 "5 mm. 

A very distinct species, easily known by the very long first joint of the funiculus of 
the antennae, the highly polished surface, and puncturation of the elytra. 

Hab. Oahu, a single example taken in the Waianae mountains in April 1892. 

(42) Oodemas jjiauiense Blackburn. 

Oodemas niauiense Blackb., Ann. Soc. Ent. Belg. xxi. p. 75. 

(Plate VIII. fig. 25.) 

Hab. Maui, Haleakala (Blackburn). I have taken this species on Molokai and 
Hawaii in some numbers. It varies greatly in size, and otherwise. Examples from 
Hawaii are generally much more shining than those from Molokai. 

(43) Oodemas borrei Blackburn. 

Oodemas borrei Blackb., Ann. Soc. Ent. Belg. xxi. p. 75. 

This species is sometimes quite brightly shining, and varies greatly in size. 

Hab. Maui, Haleakala, above the forest. Commonly found in company with 
O. nivicola (from 6000 — loooo ft.), attached to roots of grasses and low plants. 



COLEOPTERA 171 

(44) Oodenias konamim, sp. nov. 

Ovatum, robustum, nigroaeneum, opacum vel minus nitidum. Rostrum strigoso- 
punctatum. Antennae graciles, minus breves, articulo funiculi primo brevi, quam 
secundus evidenter breviore. Pronotum basi latum, opacum (rare subnitidum) subtiliter 
punctatum. Elytra sat lata, subopaca, rarius nitida, grosse seriatim punctata, plerumque 
obscure vel vix evidenter striata, interstitiis nonnunquam convexiusculis, haud dense 
punctulatis. Long. 5 — 6 mm. 

A large, wide, and robust species, generally dull, with the elytra a little shining, 
sometimes quite brightly so. The rostrum is so punctate as to have an appearance of 
longitudinal strigosity, but in some examples this is less evident. Prothorax generally 
quite dull from the microscopic rugulosity of the surface, finely, obscurely, and remotely 
punctured. In a few examples the puncturation is more evident. Elytra with series 
of coarse punctures, sometimes placed in evident striae, owing to the slight but evident 
convexity of the interstices, but the striation is vague and shallow. In many examples, 
however, there is little or no trace of interstitial convexity. The metasternum and base 
of the abdomen beneath are coarsely punctured. In spite of the variation in detail of 
the sculpture, this species is not difficult to recognize, by its large size and robust form, 
and the more or less dull surface. It is perhaps most nearly allied to O. scidpturahiin, 
but that species is of shorter and more subquadrate form, and generally has the 
interstices of the elytra more distinctly convex. 

Hab. Hawaii, Kona district (4000 ft.). About 40 examples were taken. 

(45) Oodenias rnultifornte, sp. nov. 

Nigroaeneum, nitidum, ovatum. Antennae sat graciles et elongatae, articulo 
funiculi primo minus elongate, quam secundus breviore. Elytra nitida, seriatim sat 
grosse punctata, interstitiis punctulatis. Long. 3 — 5 mm. 

To the above species I refer all the specimens of Oodenias taken by me on Hawaii, 
except the series of O. konanuni and a few specimens, which I consider a variety of that 
distinct species O. mauiense. If I am right in my conclusions, the variation exhibited 
by O. multiforme is much greater than that of any other species, so much so that the 
extreme forms would not only appear to be totally distinct, but not even very closely 
allied. When, however, a great many examples are compared, these extreme forms are 
so connected by intermediates, as to make it impossible to decide where any line 
of separation can be drawn. For this reason we have not attempted to draw up any 
detailed description of the species, as such could apply to but a fraction of the examples 
we have examined. Indeed apart from generic characters, there are probably none 
which do not exhibit more or less variation. The brief description given above applies 
to a common form of the insect, and a number of this form were taken at Kilauea, 

F. H. II. 23 



172 FAUNA HAWAIIENSIS 

Hawaii. The chief points of variation which we have observed are as follows. Size 
very variable, the largest examples being three or four times the bulk of the smallest. 
Form sometimes robust, sometimes narrow and elongate, to some extent the variation 
being probably due to sex, as it is usual in other species for the ^ to be shorter and 
wider than the %. In some the elytra become a good deal wider either a little behind 
the shoulders or still further back, while in some the curve of their sides is very even and 
forms an almost regular continuation of that of the sides of the prothorax. The surface 
of the insect is normally shining, others are less so, especially anteriorly, some are rarely 
quite dull. The rostrum varies in sculpture, sometimes it is finely and by no means 
closely punctured, sometimes the puncturation is dense and rugose. The antennae, 
which are slender, are not extremely variable, but they are longer in some examples 
than in others, and there is no doubt that some have the first joint of the funicle more 
slender and less short than others. The prothorax varies in width and in sculpture, 
generally it is finely punctured, sometimes more strongly and closely, sometimes very 
shining and almost or quite impunctate. The elytra are rarely dull, but frequently 
instead of being merely seriately punctured, they have the punctures placed in evident 
striae, and the individual punctures of the rows are often remote, but sometimes close 
and regular, even in those near to the suture. Small examples from the Kona district 
are often excessively shining and have the puncturation of the elytra much more sparse 
and irregular. The eyes which normally are a little convex in a few specimens are 
hardly so at all. 

We have attempted the division of the species on nearly all these characters and 
on minute differences in the rostrum, with the result that we have found that either a 
number of species must be made, with very indefinite distinctions between them, or that 
the whole series must be considered as one very variable species, which is possibly at 
the present time in the process of dividing into several, the division as yet being quite 
incomplete. It is of course possible that extended research into the habits etc. of some 
of these forms might prove that the views we hold are erroneous, and that there are 
really several variable species. 

Hab. Hawaii, taken in various localities on both sides of the island from 
2000 — 5000 ft. 

(46) Oodemas infermim Blackburn. 

Oodemas infernum Blackburn, Ent. Mo. Mag. xvii. p. 199. 

Hab. Hawaii, Kilauea (Blackburn). I think it doubtful whether this is not a 
form of the preceding species, but as Mr Blackburn in his final consideration of the 
species (Tr. Dublin Soc. 18S5, p. 187) includes it in those, which have a long first joint 
to the funiculus, whereas O. multiforme appears to belong to those which have this joint 
short, I have thought it safer to consider the two distinct. 



COLEOPTERA 1 73 



Anotheorus Blackburn. 

(i) Anotheorus montamcs Blackburn. 

Anof/ieortis montanus Blackb., Ent. Mo. Mag. xiv. p. 5. 
Hab. Oahu mountains, on Acacia koa. 

(2) Anotheorzis ignavus Blackburn. 

Anotheorus ignaims Blackb , op. cit. xvii. p. 201. 

Hab. Maui, Haleakala (4000 — 5000 ft.) ; var. on Lanai ; rare. This species 
approaches very closely to the preceding, and both vary in the shape of the thorax, 
and it is doubtful whether they are distinct. The Maui examples are most variable, and 
a single specimen from Lanai agrees better with these than with the Oahuan form. 

(3) Anotheorus robustus, sp. no v. 

Nigroaeneus, robustus, antennis tarsisque plus minusve testaceis. Rostrum sub- 
tilissime punctatum. Pronotum antice angustatum, plerumque nitidum, elytris multo 
angustius. Elytra lata, striata, striis confertim punctatis, interstitiis creberrime punctu- 
latis. Long, (rostr. incl.) 6 — 8'5 mm. Lat. (exempli minim.) 3 mm. (Plate VIIL 
fig. 27.) 

Although this species, of which I have examined 80, or more, examples, varies 
greatly in size, and sometimes in sculpture, its great size distinguishes it at once, the 
smallest examples being of about twice the bulk of either of the preceding. 

Hab. Kauai mountains ; common. 



Fam. SCOLYTIDAE. 

The Scolytidae are represented by twenty-six species, three only being at present 
known to inhabit other countries, although some of the others will no doubt ultimately 
be found to be likewise of foreign origin. Only three genera are represented, Xyleborus 
with 19 species being the most extensive and important. There are six species of 
Hypothenemus known, and one of Crossotarsus. The latter, C. externedentatus, is 
already known to occur elsewhere, as also is the Hypothenemus eruditus, and one of 

23—2 



174 FAUNA HAWAIIENSIS 

Xyleborus {X. confiisus). The greater number of the species of Xyleboriis and Hypo- 
thenemus are true forest insects and are no doubt peculiar to the islands. Owing to the 
extreme sexual differences exhibited by the species of Xylebortis it is impossible in most 
cases to unite the sexes from a mere inspection of the specimens, so that it is probable 
that some of the males described will prove to belong really to females described under 
another name, and therefore the species will be less numerous than they appear to be. 
So far as is at present known most of the species of this genus are restricted each one 
to a single island, and of those with a wider distribution some I suspect are not truly 
indigenous — X. iviniaturtis for example — and will prove to have been imported by man. 
The species assigned to Hypothenenius are at present very imperfectly known, and 
would appear to be rare insects, and it is doubtful whether, excluding H. erudiius, they 
are true members of that genus. It may be remarked that in addition to the species 
here enumerated a small Scolytid has been recendy imported into the islands, and is said 
to be very injurious to certain fruit-trees in some localities, but I neglected to collect or 
examine this insect. 

Tribe TOMICINI. 
Xyleborus Eichh. 

( I ) Xyleboriis molokaieiisis, sp. nov. 

Elongatus, cylindricus, piceo-niger, setis aureis elongatis postice sparsim vestitus. 
Elytra subopaca, sat distincte seriatim punctata, parte declivi juxta suturam utrinque 
tuberculis minutissimis 2 vel 3, lineariter dispositis, munita. $. Long. 4' 25 mm. 

Distinguished by its large size and dull elytra, which bear on the apical declivous 
portion two or three very minute tubercles on each side of and near to the suture, and 
placed nearly in a line with one another. From the base of each of these tubercles 
springs a long fine golden seta. Exterior to each of these series of tubercles may often 
be detected one or two other excessively minute ones on either wing-case, and the 
general surface of this posterior portion of the elytra under a very strong lens is 
minutely asperulous. 

Hab. Molokai, above 4000 ft. In the wet decaying wood of Cheirodendron. 

(2) Xyleborus kmiaiensis, sp. nov, 

X. molokaiensi magnitudine et forma simillimus, elytrorum parte postica declivi 
similariter tuberculis munita, sed colore testaceo vel rufo-testaceo, elytris sat evidenter 
nitentibus distinguendus. %. Long. 4 mm. 

Hab. Kauai, Halemanu and above Waimea (4000 ft.). Four examples taken. 



COLEOPTERA 175 

(3) Xylcborus maiiiensis, sp. nov. 

AngListus, elongatus, cylindricus, niger, antennis pedibusque testaceis. Elytra vix 
nitida, seriatim punctata, puncturatione minus distincta, subobsoleta, postice setis pallidis 
pluribus vestita, parte declivi suturam juxta tuberculis 4 vel 5 minutissimis, lineariter 
dispositis, utrinque munita, et ubique minutissime sat distincte asperula. $. Long. 

3"2— 3"5 mm. 

Closely allied to X. molokaiensis but much smaller and narrower, and with the 
posterior declivous portion of the elytra with more numerous setae. This area has also 
the appearance of being covered with rows of longitudinal and very minute asperities, 
apparently due to the mode in which the punctures are impressed. The tubercles are 
very minute, and form a row on each side near the suture, in each of which rows 3 to 5 
tubercles can be distinguished on careful examination. The better developed ones are 
usually those in the middle of the series. On either side exterior to these rows other 
very obscure and minute tubercles may be sometimes distinguished. 

Hab. Maui, Haleakala (5000 ft.). On Cheirodendron. 

(4) Xyleborus hawaiiensis, sp. nov. 

Niger, antennis pedibus testaceis, haud nitidus, X. ;;/a;//r«.f?' cognatissimus. Elytra 
opaca, subobsolete seriatim punctata, parte declivi suturam juxta tuberculis minutissimis 
2 vel 3 lineatim utrinque armata, pilis brevioribus parum conspicue vestita, vix asperula. 
%. Long. 3 mm. 

Apparently slightly less elongate than X. mauiensis, but extremely similar to it in 
nearly all respects. It may be distinguished from that species by the decidedly less 
hairy apical surface of the elytra, the hairs being less evident than those on the dorsal, 
and the general surface of the declivous portion is moreover not distinctly covered with 
very minute asperities. 

Hab. Hawaii, Hilo (2000 ft.) ; four examples taken. 

(5) Xyleborus trjmcatns Sharp. 

Xyleborus triincatus Sharp, Tr. Dublin .Soc. iii. 1885, p. 192. 

The elytra of this species are evidently shining, and it is smaller than any of the 
preceding. I have seen only two examples besides the types, and they are both darker 
than the latter, but otherwise nearly identical. 

Hab. Oahu (Blackburn). Lanai, Hawaii, in the Kona district; one from 
each locality. 

Obs. The five species enumerated above are extremely closely allied, and almost 
agree together in the character of the tubercles on the apical portion of the elytra, but 
so far as I can decide on the scanty material e.xamined, the various forms are readily 



176 FAUNA HAWAIIENSIS 

distinguished by the characters given. It may be noted that four of the species 
apparently are Hmited in range to one or other of the islands, while the fifth has been 
taken on three different islands. Of this group of species I have further seen three 
examples, which appear to belong to none of the forms described above, nor yet do they 
agree together themselves, but the material is quite insufficient for deciding as to the 
specific value of the characters they exhibit. 

(6) Xyleborus obliquus Sharp. 

Xyleborus obliquus Sharp, Tr. Dublin Soc. iii. 18S5, p. 192. 

Hab. Oahu and Hawaii (Blackburn) ; I have not met with this species. 

(7) Xyleborus lanaiensis, sp. nov. 

Nigricans, parum nitidus, elongatus, cylindricus, antennis pedibusque testaceis. 
Elytra subtiliter minus distincte punctata, postice sat abrupte declivia, parte declivi 
utrinque suturam juxta tuberculis 2 fortioribus lineatim munita, et subdepressa. $. 
Long. 3 "6 mm. 

Readily known from any of the preceding by the stronger development of the four 
tubercles of the posterior declivous portion of the elytra, these tubercles being placed 
two in a line on each side of and near to the suture, the upper pair near the com- 
mencement of the declivity, the others near the apex. From the base of each pos- 
teriorly springs a long fine seta, directed downwards and backwards. Outwardly to 
these larger tubercles there are on each wing-case one or two considerably smaller 
ones. 

Hab. Lanai 2000 ft., January 1894. Three or four examples taken. 

(8) Xyleborus similliimis, sp. nov. 

Praecedenti (X. lanaiensi) simillimus et cognatissimus, parte declivi elytrorum 
eodem modo armata. Elytra subnitida, distinctius et minus subtiliter punctata, tuber- 
culis 2 superioribus setas breviores ferentibus. ?. Long. 3 '4 mm. 

Extremely like X. Ianaie7isis, but the elytra are somewhat striate, and the punctures 
are decidedly larger and more distinct, the surface somewhat shining. Apical armature 
much as in X. lanaiensls, the four larger tubercles being well-developed and there are a 
minute pair anterior to the upper pair of these, but these may be more or less evident 
in the preceding species also. The setae which spring from the base of the anterior 
pair of the larger tubercles are evidently less developed than those in the same position 
on X. lanaiensis. 

Hab. Hawaii, above Hilo (1800 ft.); two examples taken. 



COLEOPTERA 



(9) Xyleborus oahuensis, sp. nov. 



^n 



Nigricans, antennis pedibusque testaceis, spp. duabus praecedentibus simillimus, 
sed paullo major. Elytra subtilissime sed subdistincte punctata, subnitida, parte declivi 
depressiuscula, tuberculis 6 sat distinctis, setas longas ferentibus, armata, horum 4 
anterioribus curvatim dispositis. %. Long, vix 4 mm. 

Very like the preceding two species, having four well-developed tubercles similarly 
disposed on the posterior portion of the elytra, two in a line on one side and two so 
placed on the other side of and near to the suture. Behind the anterior tubercles on 
either wing-case towards the side there is another distinct tubercle, these two and the 
anterior pair being placed in a curve. These sublateral tubercles are well-developed 
but smaller than the anterior and about equal to the posterior pair in size. From the 
base of each of the tubercles posteriorly a long fine seta rises. There are no evident 
smaller tubercles in this species. 

Hab. Oahu. A single e.xample was taken in the northern part of the Koolau 
rang-e, at an elevation of about 1000 ft. 



&^ 



(10) Xyleborus ditbiosus, sp. nov. 

Castaneus, subnitidus, pedibus antennisque testaceis. Elytra subtilissime seriatim 
punctata, parte declivi minus abrupta, tuberculis 2 minus fortibus lineariter dispositis 
utrinque juxta suturam armata, necnon latera versus utrinque tuberculis aliis minutioribus 
2 vel 3 lineariter dispositis munita, setis brevioribus. %. Long. 3 "5 mm. 

The single example above characterized is a very obscure species. The four larger 
tubercles of the posterior declivous portion of the elytra are less strong than those of 
the several preceding species, yet more so than those of the series ending with 
X. truncaius. In front of the anterior pair of these there is a third pair smaller and ill- 
developed, and in a line with the others. They are situated close to the suture just at 
the beginning of the posterior declivity, and in the unique example are placed quite 
obliquely and not opposite the one to the other. Besides this series there is a second 
one of about three minute tubercles on each wing-case towards the side. 

Hab. Maui. A single example was taken in the lao valley in 1894. 

(11) Xyieboriis confustis Eichhoff. 

Xyleborus confusus Eichhoff, Ratio, Descr. emend. Tomicinorum (1S79). 

Xyleborus insularis Sharp, Tr. Dublin Soc. iii. 1885, p. 193. 

Hab. Oahu and Kauai (Blackburn) ; Oahu (both ranges), Maui, Kauai, Hawaii. 
At elevations from 1500 — 4000 ft. above sea-level. We are indebted to Mr W. F. H. 
Blandford for the identification of this species. 



178 FAUNA HAWAIIENSIS 



(12) Xyleborus rugatus Blackb. 

Xyleborus 7'ugatus Blackburn, Tr. Dublin Soc. iii. (1885), p. 192. 
Hab. Oahu. A single specimen taken. (Blackburn.) 



(13) Xyleborus frigidus Blackb. 

Xyleborus frigidus Blackburn, Tr. Dublin Soc. iii. 1885, p. 193. 

Hab. Maui. A single specimen taken on Haleakala, 4000 ft. (Blackburn.) 



(14) Xyleborus ininiaturus Blackb. 

Xyleborus immaturus Blackburn, Tr. Dublin Soc. in. 1885, p. 193. 

I have taken what is probably the t of this insect. It is testaceous or rufotesta- 
ceous in colour, but otherwise bears no resemblance to the $. Form short and robust, 
the whole insect hardly twice as long as wide, longitudinally convex. Thorax very 
narrowly rounded, or somewhat pointed, in front, of ovate form, truncate at the base. 
The sculpture is very feeble, and there is none of the coarse rugosity in front such as is 
seen in the $, and the clothing there consists of shortish and inconspicuous hairs. 
Elytra short, rather longer than wide in dorsal aspect, rather distinctly substriate, the 
striae finely punctured. There are no evident tubercles on their posterior portion. 
Length hardly 2 mm. 

Hab. Oahu and Hawaii (Blackburn) ; Hawaii in various localities. 



(15) Xyleborus agaiiius, sp. nov. 

$ castaneus, parum elongatus, tibiis omnibus plus minus obscuratis, tarsis pallidis. 
Pronotum magnum, antice acuminatum, ruguloso-punctatum, anterius et latera versus 
pilis longis vestitum, lateribus sat fortiter rotundatis. Elytra parce pilosa, desuper 
visa longiora quam latiora, haudquaquam striata, obscure confuse punctata, lateribus 
subcompressis. $. Long. 2 mm. 

The form of the thorax is much like that of the $ assigned to X. immattirus, but 
its sculpture and clothing are very different, as also is the colour of the entire insect. 
The elytra are different in form and sculpture, having no trace of striation, and the 
punctures are very feeble and indistinct. 

Hab. Lanai. Two examples were taken at an elevation of 3000 ft. in Jan. 1894. 



COLEOPTERA 179 



(16) Xyleborus exsectus, sp. nov. 

Nigricans, elytrorum basi nonnunquam rufescente, pedibus antennisque rufes- 
centibus vel testaceis. Pronotum suboblongum, antice sat profunde excisum, et in 
processum latum, cujus apex truncatus, productum, lateribus pilis vestitis. Elytra 
leviter striata, striis subtiliter punctatis, setis parce vestita, parte declivi baud tuber- 
culata. 1^. Long. 3 '2 mm. 

Hab. Maui. Three examples taken on Haleakala (5000 ft.). One of these is of 
a testaceous colour and is no doubt immature. 



( I 7) Xyleborus viilcanus, sp. nov. 

$ nigricans, elytris thoracisque basi subpiceis, pedibus testaceis, oblongus. Pro- 
notum parum distincte sculpturatum, elytris vix latius, lateribus parallelis, setis elongatis 
aureis parce vestitis, anterius excisum, et in processum triangularem productum. Elytra 
sat elongata, rugulosa, vix evidenter punctata, parte declivi tuberculo minutissimo 
suturam versus utrinque munita. $. Long. 2'5 mm. 

Much smaller than the preceding species, and very distinct by the pointed process 
of the pronotum. The posterior declivous portion of the elytra is somewhat abrupt, 
and on its upper portion on either side of the suture an excessively minute tubercle can 
be seen, each of which is furnished with a longish seta at its base, while some even 
more minute and hardly visible tubercles are placed in a transverse line with these. 

Hab. Hawaii. A single example taken at Kilauea. 



(iS) Xyleborus littoralis, sp. nov. 

$ castaneus, pronoto plus minus nigricante, pedibus testaceis, oblongus. Species 
prima facie X. vtilcano simillima, sed major, et minus angusta, pronoto antice fere 
similariter producto. Elytra substriata, grossius distincte seriatim punctata, parte 
declivi juxta suturam utrinque tuberculis minutissimis 2 vel 3 lineariter dispositis, 
aliisque paucis vel minutioribus et obscuris, munita. Z- Long. 3 mm. 

Easily distinguished from A', viilcaims by the substriate and somewhat coarsely 
punctured elytra, and the more numerous and rather more developed tubercles of the 
posterior truncation. These tubercles form two series on each side, those nearest the 
suture, though very minute, being fairly distinct, those external and subparallel to these 
are very minute and obscure. 

Hab. a single example taken at sea level on Molokai, July 1893. 
F. H. II. 24 



i8o FAUNA HAWAIIENSIS 



(19) Xyleboriis ignobilis, sp. nov. 

Nigricans, thorace nonnunquam obscure testaceo vel piceo, antennis pedibusque 
testaceis. Pronotum fere ad basim rugulosum, antice setis pallidis vestitum, lateribus 
rotundatis haudquaquam parallelis. Elytra plus minus obsolete seriatim punctata, 
setis pallidis brevibus, lineariter dispositis, vestita, postice baud abrupte declivia, parte 
declivi tuberculis nullis munita, nee impressa. Long. 27 mm. 

This insect bears little resemblance to any other Hawaiian species of the genus, 
the elytra being simply rounded off behind and not at all abrupt, nor tuberculated, but 
sculptured as on the dorsal surface. The prothorax becomes gradually less rough 
from the front margin backwards, but it is more or less asperulous or rugulose to 
very near the basal margin, near which the rugulosities have a somewhat concentric 
arrangement. 

Hab. Hawaii. Three examples taken, each in a different locality, two on the 
windward and one on the lee side of the island (2000 ft.). 



Hypothenemus Westw. 

(i) Hypothenemtis eriiditus Westw. 

Hypothenemus erudihis Westwood, Tr. Ent. Soc. London, i. p. 34 ; Sharp, op. cit. 
1879, p. 102. 

Hab. Oahu ; Honolulu (Blackburn). 

(2) Hypothenemus viaadicolhs Sharp. 

Hypothenemus maculicollis Sharp, Tr. Ent. Soc. London, 1879, Pt. I. p. loi. 

Hab. Oahu ; mountains near Honolulu (Blackburn). I did not collect this 
species. 

(3) Hypothenemus griseus Blackb. 

Hypothenemus griseus Blackburn, Tr. Dublin Soc. iii. 1885, p. 194. 

Hab. Oahu ; a single specimen taken on the plains near Honolulu (Blackburn). 



COLEOPTERA i8i 

(4) Hypothcncimis sylvicola, sp. nov. 

Minus elongatus, cylindricus, pallide testaceus, capite et pronoto plus minusve 
infuscatis. Pronotum subtilissime granulato-sculpturatum, parte anteriore asperata, 
opacum, parce pubescens. Elytra pallida, setis pallidis vestita, subtilissime vix evi- 
denter punctata, baud evidenter striata. Long, i '5 mm. 

In colour this species must greatly resemble H. griseus Blk., but that species has 
the elytra deeply striate. In some examples the asperities of the prothorax are but few, 
in others they are much closer and more numerous, and I suspect that the difference is 
sexual. The sculpture otherwise is excessively feeble and consists of a very dense 
and excessively minute granulation or puncturation, not definitely distinguishable even 
with a very strong lens. The clothing of the elytra consists of short pale setae, which 
in certain aspects appear to have a linear arrangement, and also of some longer fine 
hairs, which are particularly noticeable at the sides about the apex. 

Hab. Lanai, 2000 ft. several examples. — Kauai, 4000 ft., one example taken. 
The latter has the thorax entirely pale and concolorous with the elytra. 

(5) Hypothenemus insularis, sp. nov. 

Cylindricus, angustulus, sat elongatus, niger, antennarum basi pedibusque rufo- 
testaceis. Pronotum anterius fortiter asperulum et parce setosum, posterius opacum, 
subtilissime granulatum (an punctulatum ?). Elytra setis brevibus griseis conspicue 
vestita, sat elongata (circa bis longiora quam basi latiora), evidenter punctata. Long. 
I — I "4 mm. 

Allied to H. ruficeps, but very different in superficial appearance, owing to its 
entirely black colour. The punctures on the elytra are not so very fine, but they are 
ill-defined. Apparently they are more or less serially arranged, but not in the definite 
manner seen in the last species, in fact the puncturation appears somewhat rugose. 

Hab. Kauai. Two examples were taken at Makaweli, above 2000 ft. 

(6) Hypothenemus ritjiceps, sp. nov. 

Cylindricus, sat elongatus, capite et pronoto rufescentibus, elytris nigris. Pronotum 
anterius sat fortiter asperulum, postice opacum et parum distincte sculpturatum, parce 
pubescens. Elytra nitida, setis brevibus griseis sat conspicue (postice saltem) vestita, 
subobsolete striata, striis subgrosse punctatis. Long, i "5 mm. 

Distinct by the red head and prothorax, and black elytra. The punctures on the 
latter are somewhat coarse but not very definite, and are placed in subobsolete striae. 
The short whitish setae are dense and conspicuous on the posterior portion of the 
elytra, less so in front, but they have been partly abraded on this part. 

Hab. Oahu ; Kaala, Waianae range (2000 ft.) ; one example taken. 

24 — 2 



1 82 FAUNA HAWAIIENSIS 

Tribe PLATYPINI. 

Crossotarsus Chapuis. 

(i) Crossotarsus externedentatus Fairm. 

Crossotarsus externedentattis Fairmaire, Rev. Zool. 1S50, p. 51. 

Hab. Oahu ; Honolulu (Blackburn). — I cut a pair of this species out of a 
hard trunk of Acacia koa, at an elevation of 1000 ft. in the northern part of the 
Koolau range. 

Fam. ANTHRIBIDAE. 

The Anthribidae are very poorly represented in the islands, only three species 
having occurred. Two of these species are at present not known from elsewhere, 
and one of them constitutes a peculiar genus, but there is little doubt that all are foreign 
insects, and will ultimately be found in other countries. 

Araeocerus Schonherr. 
(i) Araeocerus fasciculatus De Geer. 

Ctirculio fasciculatics De Geer, Ins. v. 276, /. 16, fig. 2. 

Hab. Abundant all over the islands in the mountains. The large number of 
constant varieties in markings, as well as the great variation in size of this species is 
remarkable. 

(2) Araeocerzis const an s, sp. no v. 

A. fasciculato simillimus sed statura majore, totus dense griseo-pubescens, haud 
variegatus, tarsis anterioribus t robustioribus, et tarsis % longioribus distinguendus. 
Long, circa 5 mm. 

Very closely allied to A. fasciculatus, but larger than the larger examples of that 
species, not varying much in size, and uniformly covered with dense grey pubescence. 
In colour the integument is blackish and reddish fuscous, the antennae towards the 
base and the tibiae are more or less red. When the pubescence is removed, the 
pronotum is seen to be dull and very densely punctured, and the elytra bear each about 
ten rows of very distinct and regular punctures, the interstices having a dense shallow 
and very minute rugulose puncturation. Both sexes may be distinguished structurally 
from A. fasciculatus by the thicker anterior tarsi, so that the t of the latter rather 
resembles the % of the former, and these parts are longer in the % oi A. const an s. 

Hab. Hawaii, Kona ; on the coast ; found in the flowers of the white poppy. 



COLEOPTERA 183 

Mauia Blackb. 
(i) Mania satellcs Blackb. 

Majiia satelles Blackburn, Tr. Dublin Soc. in. 1885, p. 195. 

Hab. Maui ; Wailuku valley, where one specimen was taken by Blackburn. 
I have not met with this species. 



Fam. PROTERHINIDAE. 



This remarkable family is peculiar to the Hawaiian islands, and all the species, of 
which 122 are at present known, are assigned to the genus Protcrhinus. These species 
exhibit so much diversity in structure that the extreme forms would appear to be 
certainly generically distinct, but they are so connected by intermediate forms, that it is 
inadvisable to attempt a further division of the genus until the species are more 
thoroughly known. At present, owing to the great number of the species, the close 
alliance between many of them, their variability, and the great sexual differences, the 
mere separation of the species is attended with the greatest difficulty. The figures on 
Plates I X. and X. will show to some extent the diversity in structure and appearance 
exhibited by these insects, and the specimens figured have been chosen for this purpose, 
rather than with a view to exhibiting the minute distinctions between closely allied 
species — an almost hopeless task in dealing with insects so variable both in structure 
and in superficial appearance. 

In the Cambridge Natural History in the classification of the Coleoptera Dr Sharp 
leaves the Proterhinidae as well as the somewhat similar insects comprised in the 
family Aglycyderidae unclassified. The former have much more the appearance of 
Rhynchophora than the latter, since the beak is always distinct and definite in 
Proterhinus, at least in the female sex. In the t however it is hardly more than a 
simple prolongation of the head, and only very rarely does the rostral portion attain a 
length greater than its width. The female beak on the other hand is in most cases 
elongate, and on account of its shining surface and absence of squamosity extremely 
definite. It is noteworthy that in a considerable number of species, which have the 
rostrum in the ? below the average in development, the rostrum of the ? is above the 
average, so that the sexual differentiation is less than usual, and I believe that these 
species are the more primitive, and their resemblance to the members of the Aglycyde- 
ridae is greater than that of the other forms. The Proterhinidae are however always 
distinct from Aglycyderes by the definite rostrum of the ?, and by the structure of the 
tarsi which are really four-jointed, the small division at the base of the claw-joint, being 
a true joint and not a mere constriction as in Aglycyderes. With regard to the 



1 84 FAUNA HAWAIIENSIS 

structure of the prothorax in these Insects to which considerable importance was 
attached by Dr Leconte, who regarded Proterkinns as belonging to the Rhynchophora 
and Aglycyderes as related to the Colydiidae (see quotation from a letter, Trans. Ent. 
Soc. Lond. 1879, Pt. I. p. 78), it may be noticed that the differentiation between the 
dorsal surface and flanks is much more marked in a few species than in the majority of 
P7'oter]mms, which, as Sharp (/. c. p. 80) has remarked, is also the case with the two 
species of Aglycyderes, the New Zealand species having the differentiation between the 
parts of the prothorax less definite than the Canarian species. 

The characters which I have found most useful in the separation of the species are 
the nature of the clothing, the size and form of the eyes and of the lobes of the front 
tarsi, the condition of the humeral angles of the elytra, the shape of the prothorax, the 
length of the antennal joints, and the development of the club (or 3 apical joints), and 
the puncturation of the abdomen beneath. In few species are any of the characters 
constant, when a long series of examples are examined, the size of the individuals being 
extremely variable, and these depauperated examples often have the structures charac- 
teristic of the species much modified. For these reasons I have not found it easy to 
identify the numerous species hitherto described, although through the kindness of 
Dr Sharp I have been able to carefully examine the types of all the species described 
by him. The description of most of these species was drawn up from only one or two 
examples, and except in the case of a few of the most distinct species, such material is 
quite inadequate for an accurate knowledge of the species. In several cases when 
I have referred a species taken by myself to one already described by Dr Sharp, I have 
not been able exactly to match the types with any individual even in a series of 
specimens, and it is probable that this is due to the specimens having been captured in 
a slightly different locality to that whence the typical examples came, but it is of course 
possible that some of the species have themselves undergone slight changes during the 
last quarter of a century. 

Of the 122 species known 8 only appear to extend their range beyond a single 
island, and in few, if any, of these do the examples from different islands altogether 
agree, while several of these species inhabit only two of the closely adjoining inter- 
mediate islands of the group. Of the species that restrict their range to a single island 
Kauai has 29, Oahu 28, Maui 27, Lanai 9, Molokai 9, and Hawaii 12. To facilitate 
the discrimination of the species I have arranged the species in seven divisions, one for 
each island and comprising the species peculiar to it, the seventh containing the few 
species which are found on more than one island. 

Nearly all the species are attached to the dead or dying wood of the forest trees, 
and the individuals frequently congregate in small batches beneath the bark, in which 
the larvae feed, and they are very sluggish in their habits. Several are found in the 
stems of tree-ferns, and one or two are attached to smaller ferns {Pteris etc.), while a 
few live on small shrubs and woody creepers. 



COLEOPTERA ' 185 

A. Species insulae Kauai propriae. 

(i) Proterliiniis gigas, sp. nov. 

Robustus, pallida squamosus, nigricans, antennis, pedibus elytrisque saepe plus 
minusve rufescentibus. Oculi magni, fortiter prominentes. Antennae graciliores, 
articulis elongatis, tertio quam secundus bis longiore. Pronotum inaequale, 3-impressum, 
rugoso-punctatum, antice constrictum, pallide squamosum, margine antico lateribusque 
setis curvatis vestitis. Elytra lata, squamis pallidis necnon setis erectis, sat conspicuis, 
vestita, basi juxta scutellum utrinque tuberculata, humeris fortiter productis. Pedes 
setis erectis conspicue vestiti. Abdominis segmentum primum ventrale circa medium 
obsoletissime, latera versus et ad basim fortiter punctatum. $ %. Long. 4 — 5 '5 mm. 
(Plate IX. fig. I, $.) 

This large species can hardly be confused with any other, although it varies 
considerably in size and in other respects. The 8th joint of the antennae is sometimes 
a little longer, sometimes a little shorter than the 9th, and generally they are subequal. 
In a few examples out of a long series the 8th is considerably the shorter of the two, as 
is usual in most species of the genus. The surface of the elytra, except for the 
depression formed between the shoulders and the tubercle on either side of the 
scutellum, is nearly evenly convex. 

Hab. Kauai (4000 ft.). Under bark of Cheirodendron only. 

(2) Proterhinus crassicoTiiis, sp. nov. 

Forma, facie, et magnitudine fere praecedentis, sed antennis brevioribus, articulis 
multo crassioribus, abdominis segmento 1° ventrali circa medium fortiter et distincte 
punctato, bene distinctus. $. 

Hab. Kauai mountains (2000—3000 ft.). Only two examples were taken, owing 
to the erroneous supposition that they were identical with those of the preceding 
species, which is found on the high plateau of the same island. 

(3) Proterhinus antJu-acias, sp. nov. 

Niger, vel piceo-niger, latus, antennis pedibusque saepe rufescentibus. Oculi 
magni, prominentes. Antennae graciliores, scapo brevi, articulo secundo tertioque una 
conjunctis multo breviore, clava gracili. Pronotum antice fortiter et abrupte constrictum 
(vel nonnunquam tantum angustatum), anteriusque profunda foveatum, angulis posterio- 
ribus macula pallida squamosa sat conspicue signatis. Elytra lata, maculatim squamosa, 



1 86 FAUNA HAWAIIENSIS 

setisque erectis et elongatis vestita, fortiter punctata, humeris productis. Abdominis 
segmentum primum ventrale, vel ad medium, grosse fortiterque punctatum. Z ?• 

Long. 3—475 mm. 

A species of wide and somewhat depressed form, rather scantily clothed with 
squamosity, forming in fresh examples a number of roundish spots on the elytra, which 
also bear pale erect setae. The scape of the antennae is not long, being evidently 
shorter than the two following joints united. Second joint evidently longer than wide, 
but very much shorter than the third. All the other joints elongate, the 8th shorter 
than the gth, the latter with the two following forming the long and slender club. Eyes 
large and prominent. Thorax usually strongly and abruptly constricted in front, but 
variable in this respect. In front there is a deep and conspicuous impression, and 
sometimes two lateral, and one posterior (near the hind-margin about the middle) 
as well, but all except the anterior one may be obsolete. The posterior angles of the 
thorax are very clearly marked, being covered with a spot of dense pale squamosity. 
Elytra wide, roughly punctured, generally more or less flattened, the sides somewhat 
strongly rounded, the shoulders prominent. Lobes of the front tarsi only moderate in 
size, or even small for the size of the insect. Whole body beneath coarsely and closely 
punctured. 

Hab. Kauai, widely distributed and not rare (2500 — 4000 ft.). Remarkable for 
its dark colour, wide form, the distinct spots of squamosity on the posterior angles of 
the thorax, and the clothing of the elytra, which consists of long fine erect setae, and 
numerous, often rather ill-defined, spots of appressed scales. 



(4) Proterhinus eugonias, sp. nov. 

Sat latus, nigricans, piceus, vel rufescens, pallide squamosus. Antennae graciles, 
clava sat evidenter 3-articulata. Oculi parum fortiter prominentes. Pronotum profunde 
3-impressum. Elytra sat lata, convexa, parte basali depressa, minus fortiter nee dense 
punctata, pallide squamosa, setisque erectis parce vestita, humeris late ac fortissime 
productis. Segmentum abdominis primum (saltern ad medium) vix evidenter punctatum. 
^ %. Long. 2 — 3-5 mm. (Plate IX. fig. 2, ?.) 

This species is allied to P. basalis Sh. It varies greatly in size but is a smaller 
insect than the other. The form is wide and somewhat short, the colour varies from 
black to red. The squamous covering is of a golden or silvery colour. 

Antennae moderately long, slender, generally clear red with the apical joints dark, 
the club distinctly formed of three joints but rather slender and elongate. Eyes 
moderately large, but not at all strongly prominent ; no ridge between the vertex and 
front of head. Thorax with three very deep impressions, the anterior the largest; 
immediately in front of each of the posterior ones the squamosity forms a dense patch, 



COLEOPTERA 187 

different to the general covering. Elytra short, seen from the side strongly convex 
longitudinally, with golden or greyish appressed squamosity, and sparse erect white 
setae, which are chiefly found towards their apex, the basal portion is transversely 
flattened or depressed. Humeral angles widely and extremely strongly produced. 
Tibiae, tarsi, and sometimes the femora, red ; lobes of the front tarsi of moderate size. 
Basal segment of the abdomen beneath with the puncturation obsolete, at least over its 
middle portion. Hind coxae widely separated, the distance between them considerably 
greater than the length of the metasternum. 

Hab. Kauai mountains (4000 ft.). Differs from P. basalis Sh. in the smaller 
average size, more strongly produced humeral angles of the elytra, their less numerous 
erect setae, and the patches of dense squamosity in front of the posterior impressions of 
the thorax. Both species vary much, but no doubt they are distinct. 



(5) Proterhinus basalis Sharp. 

Proterhimis basalis Sharp, Tr. Ent. Soc. London, 1879, p. 98. 

I have examined a series of examples of this species. The humeral angles of the 
elytra are a good deal more strongly produced in some individuals than others. The 
squamous covering is sometimes of a ferruginous colour, but more often greyish. The 
shape of the thorax is variable, but usually is more or less abruptly constricted in front. 
The length of the insect varies from 2 '5 — 4 mm. 

Hab. Kauai mountains (2000 — 4000 ft.) ; not common, but widely distributed. 



(6) Proterhinus dubiostis, sp. nov. 

Sat latus, rufescens vel piceus, aureo-squamosus. Oculi parum fortiter prominentes. 
Antennarum articuli 3 apicales clavam formantes. Pronotum sat evidenter 3-impressum. 
Elytra lata, aureo-squamosa, minus fortiter nee dense punctata, basali parte plus minusve 
impressa, scutellumque juxta utrinque tuberculata, setis erectis paucis, brevibus et 
inconspicuis, humeris fortiter acute productis. Abdominis segmentum primum ventrale 
ad medium vix evidenter punctatum. $ $. Long. 2 — 3 '2 mm. 

The above characters are drawn up from typical examples, but as a matter of fact 
almost all the structures vary in this species, and series of individuals from different 
localities nearly always present more or less considerable differences. Thus in the 
typical examples the antennae are somewhat stout, the club distinctly of three rather 
wide joints. In some localities the antennae are decidedly more elongate and slender. 
The squamosity appears to be always of a golden colour. The thorax has three impres- 
sions and is not very densely squamose. The elytra have the humeral angles strongly 

F. H. II. 25 



1 88 FAUNA HAWAIIENSIS 

and acutely produced, their basal portion is more or less flattened or impressed (not 
very definitely so as in P. basalts), they have only a few short and inconspicuous erect 
setae, are rather finely and not very densely punctured, and are evenly convex trans- 
versely, except at the base. The lobes of the tarsi are of moderate size. The basal 
abdominal segment is hardly visibly punctured, at least on its middle portion. Examples 
of this species taken at low elevations (2000ft.), differ from typical ones in all or some 
of the following characters. The antennae are shorter and thicker, the insect is of 
narrower form, the elytra have the humeral angles less produced, and are less impressed 
at the base, while the front tibiae are often extremely short and wide. Some of the 
abnormal individuals taken with typical examples however possess some of these 
characters. 

Hab. Kauai. Common above Waimea at an elevation of 4000 ft. The 
examples most aberrant in one direction come from Halemanu (4000 ft.), in the other 
from Makaweli (2000 ft.). 

(7) P voter Iiinus difficilis, sp. nov. 

Rufescens vel piceus, sat latus, minus dense pallide squamosus. Antennae graci- 
liores. Oculi haud fortiter prominentes. Pronotum minus dense squamosum, 3-im- 
pressum, antice saepe abruptius constrictum. Elytra pallide squamosa, setisque erectis 
brevibus sed conspicuis vestita, longitudinaliter obscure carinata, humeris productis. 
Abdominis segmentum primum ventrale circa medium sparsim subobsolete punctatum. 
$ ?. Long. 2-25— 3-5 mm. 

In form and general appearance very similar to P. diibiosus but a rougher 
insect. The squamosity is less appressed, the erect setae of the elytra are more 
numerous, their puncturation is more pronounced, and the surface less even, there being 
traces of longitudinal ridges extending back from the shoulders. These ridges in fresh 
examples are rendered more distinct by the squamosity which is somewhat condensed 
along them. The humeral angles are less strongly produced forwards than in well- 
developed examples of P. dubiosus. The examination of a series of individuals of both 
species is necessary for an appreciation of the specific characters, as both vary greatly 
in size, and in the development of the structural characters which distinguish them. 

Hab. Kauai mountains. Less common than P. dubiosus. 



(8) Pi'oterhinus eulepis, sp. nov. 

Sat latus, nigricans, elytris rufescentibus, nigro-maculatis. Antennae graciles sat 
elongatae. Pronotum squamis appressis vestitum, angulis posterioribus distinctis, 
maculaque pallida squamosa signatis. Elytra squamosa, setisque erectis brevioribus 



COLEOPTERA 1S9 

pallidis vestita, minus dense punctata, humeris acutis et productis. Abdominis segmen- 
tum primum ventrale vel ad medium fortiter et distincte punctatum. Z ?• Long. 
2'5 — 3"25 mm. ; van minor, 175 — 2 mm. (Plate IX. fig. 3. $.) 

This is another variable species, very closely resembling typical P. dubiosus in form. 
The antennae are slender and rather long, the three-jointed club being also slender. 
The thorax is covered with squamosity, and has sometimes three impressions, but the 
two posterior ones may be obsolete. Its hind angles are a little prominent, and 
rendered more distinct by bearing a spot of pale squamosity, which is more dense than 
that of the general surface. The elytra appear to be always red, more or less spotted 
with black, the black spots being free from squamosity. They are nearly evenly 
convex in shape, the basal tubercles near the scutellum being absent or very ill- 
developed, their basal margin is rather deeply and regularly emarginate, so that the 
shoulders are produced and acute, and are free from the base of the thorax. The erect 
setae are short and white, but rather conspicuous in fresh examples, especially at the 
sides of the elytra. The puncturation is not very dense. All the femora are strongly 
clavate. Beneath, the basal abdominal segment is strongly and distinctly punctured, 
although towards the middle the punctures are more remote than at the sides. Very 
small examples (var. minor) apparently referable to this species are not infrequent, but 
there appears to be no good character to separate them, unless the more obsolete 
puncturation of the base of the abdomen beneath should prove constant. 

Hab. Kauai mountains (4000 ft.). 



(9) Proterhinus neglecHcs, sp. nov. 

Rufo-piceus, sat latus, elytris rufescentibus, nigro-maculatis. Antennae totae rufae, 
sat elongatae, sed crassiusculae, clava distincte 3-articulata. Pronotum aureo-squamosum, 
antice fortiter impressum, dense rugoso-punctatum, postice minus fortiter angustatum, 
angulis posterioribus sat distinctis. Elytra minus dense punctata, submaculatim squa- 
mosa, setisque brevibus pallidis vestita, sat lata, humeris acutis, evidenter productis. 
Femora omnia fortiter incrassata. i ?. Long. 2-8 — 3-2 mm. 

This species has very much the form and general appearance of large typical 
examples oi P. eidepis. In the two examples taken the anterior thoracic impression is 
large and distinct, the others are obsolete. The hind angles are distinct, and appear to 
be marked by spots of denser squamosity than that of the rest of the thorax. The 
elytra are nearly evenly convex, there being little or no trace of the tubercles at the 
base near the scutellum, but they are a little flattened transversely behind their basal 
margin. The humeral angles are acute and formed like those of P. culcpis. From 
that species P. neglectus is at once distinct by the thicker antennae, the intermediate 

25 — 2 



I90 FAUNA HAWAIIENSIS 

joints being quite stout, whereas in the other they are very slender. This seems to be 
the only good character to distinguish the two, and it may be observed that the specific 
•differences between P. gigas and P. crassicornis are quite similar. 

Hab. Kauai. A single pair (^ ?) were taken together on the high plateau in 
April, 1895. 



(10) Proterhinus collaris Sharp. 

Proterhinus collaris Sharp, Tr. Ent. Soc. London, 1879, p. 96. 

This species is very similar in form and general appearance to P. eiilepis, but may 
be distinguished by the unusual length of the scape of the antennae. The second joint 
is very short, the third elongate and slender. In the $ the eyes are unusually remote 
from the anterior margin of the thorax. The squamosity of the elytra appears to be 
much broken up to form roundish spots. The thorax is decidedly elongate. In their 
systematic catalogue Blackburn and Sharp have sunk this species as a variety of 
P. nigricans, but I cannot think the two are identical, P. collaris being a larger insect, 
and very different in general appearance, the ^ having relatively smaller eyes, and much 
more remote from the thorax, while the basal joint of the antenna in the % is much 
more elongate and more slender, than in P. nigtncans %. 

Hab. Kauai mountains (Blackburn). I have not met with this species, and have 
seen only the two types in Dr Sharp's possession, and two or three individuals in the 
British Museum from Blackburn's collection. 



(11) Proterhinus amaurodes, sp. nov. 

Statura parva, vel saepe minima, nigricans vel rufo-piceus. Antennae sat graciles. 
Pronotum dense rugoso-punctatum, parum squamosum, antice impressum, baud con- 
strictum, lateribus rotundatis. Elytra obscure sed dense punctata, ad humeros macula 
pallida squamosa, et postice maculis rotundatis compluribus ornata, humeris plus minusve 
productis, semper acutis et distinctis, setis erectis inconspicuis et brevissimis. Tarsorum 
lobi parvi. Abdominis segmentum primum ventrale subfortiter punctatum. ^ %. Long. 
1-5 — 2-5 mm. 

In well-developed examples this is rather a distinct little species. The squamosity 
of the thorax is sparse, and that of the elytra is much broken up, forming distinct spots. 
There is always a distinct patch extending back from the humeral angles, and a number 
of roundish spots on the posterior parts of the elytra. The antennae are moderately 
long and slender, with 3-jointed club. There is usually a distinct impression on the 
thorax in front, the posterior impressions being absent or nearly obsolete. The humeral 



COLEOPTERA 



191 



angles of the elytra are always sharp and distinct, and in well-developed examples 
evidently, though not strongly, produced. The lobes of the front tarsi are small. The 
colour of the insect is typically black, the squamosity of the elytra being placed on paler 
(piceous or reddish) spots, but in many examples (especially from certain localities), the 
whole elytra are rufo-piceous or reddish. The species appears to be allied to P. 
vestitus. 

Hab. Kauai. Common generally (from 2000 — 4000 ft.). Often so covered with 
a wax-like excretion as to be only recognizable after the most careful cleaning. 



(12) Proterhinus nigricans Sharp. 

Proterhinus nigricans Sharp, Tr. Ent. Soc. London, 1879, p. 95. 

In its typical condition this species is remarkable for the size of the basal joint of 
the antennae in both sexes, that of the Z being also considerably longer than is usual in 
the genus. I have not met with any examples that altogether agree with the types, but 
one Z has the antennae similarly formed, while its eyes are somewhat smaller. Others 
which have the eyes as in the type have the basal joint of the antennae less developed. 
The colour of the elytra varies from black to reddish-pitchy with black markings. The 
squamosity of the thorax is not dense, but the posterior angles are distinctly marked by 
a small dense spot of pale scales. The species is no doubt distinct, although closely 
allied to P. amaurodes, collaris, &c., with which it agrees in the strong puncturation of 
the basal ventral segment of the abdomen. 

Hab. Kauai. On Waialeale, 3000 ft. (Blackburn). Makaweli (2000 — 3000 ft.), 
and behind Waimea (4000 ft.). Apparently not common. 



(13) Proterhinus binotatus, sp. nov. 

Rufescens, capite thoraceque saepe obscuratis, elytris plaga utrinque magna nigri- 
cante maculatis. Antennae nigrae articulis basalibus rufescentibus, clava baud abrupte 
3-articulata. Pronotum ex majore parte parce squamosum, latera versus densius 
squamoso-maculatum, rugoso-punctatum, antice subimpressum. Elytra, partibus nigri- 
cantibus exceptis, pallide squamosa, setisque brevissimis, albidis, et erectis vestita, 
angulis humeralibus parum productis, sed distinctis, et subacutis. Pedes omnes setis 
brevibus erectis vestiti, lobis tarsorum anticorum magnitudine mediocribus. Abdominis 
segmentum primum ventrale minus distincte punctatum. t ?• Long. 2-2 — 2"6 mm. 

This obscure species appears to be closely allied to P. simplex Sh. of Oahu, but it 
is readily distinguished from the well-developed examples of that species. The elytra 



192 FAUNA HAWAIIENSIS 

are widely red at the base, and covered with pale squamosity, generally over a sub- 
triangular area ; behind this are two large black, or dark, lateral spots, extending nearly 
to the suture, and free from squamous covering ; the apical portion is often more or less 
infuscate, and the squamosity broken up into spots. 

Hab. Kauai mountains (4000 ft.) ; found very sparingly on several occasions. 



(14) Proterhinus scutatus Blackb. 

Proterhinus scutatus Blackburn, Tr. Dublin Soc. iii. 1885, p. 169. 

I have taken only three or four examples of this species, which mostly quite agree 
with those collected by Mr Blackburn. This insect is much less clothed with squamosity 
than most of the species, the thorax being nearly bare, and the elytra in fresh examples 
very little squamose, except for two longitudinal lines extending backwards from their 
shoulders ; but the latter bear numerous short and rather thick erect setae, which are 
pale in colour and quite conspicuous. In structure this species closely resembles some 
examples of P. simplex, but it is no doubt distinct. 

Hab. Kauai, 2500 ft. (Blackburn) ; (2000 — 4000 ft.), in the Waimea district. 



(15) Proterhimis sctiilostts, sp. nov. 

Ferrugineus, elongatus, angustulus, setulis erectis brevibus vestitus, elytris non- 
nunquam nigro-maculatis. Antennae plerumque breviores, concolores, rufae. Rostrum 
$ breve, fortiter longitudinaliter rugosum, paullo vel baud longius quam latius. Prono- 
tum minus latum, plerumque plus minusve evidenter 3-impressum, setis appressis parce, 
aliisque erectis, vestitum. Elytra fortiter elongata, lateribus subparallelis, subseriatim 
punctata, setis brevibus erectis vestita, vix aequaliter convexa, angulis humeralibus 
distinctis, paullo productis, vel fere rectis. Femora postica parum fortiter clavata, 
tarsorum anticorum lobis parvis. $ $. Long. 2*5 — 3 mm. 

In its rufescent colour, elongate elytra, and the absence of squamosity, this species 
somewhat resembles the P. loiigulus Sh. of Oahu, and the other species allied thereto. 
It is however more nearly allied to P. blackburni Sh. &c., the second joint of the 
antennae in certain aspects being not much shorter (though more robust) than the 
third, and the short rostrum of the ? is coarsely sculptured. The seven or eight 
examples taken exhibit some variation, the antennae being decidedly less short in some 
than in others. The dorsal surface of the elytra is somewhat flattened, and, at least in 
some examples, there are traces of longitudinal raised lines. 

Hab. Kauai (2500 — 4000 ft.); not common. 



COLEOPTERA 193 

(16) Proterliimts an/iqinis, sp. nov. 

Rufescens vel rufo-fuscus, angustus, elongatulus, elytris nonnunquam nigro- 
maculatis, setis appressis albidis vel argenteis, aliisque erectis sat conspicuis, sed 
brevioribus, vestitis. Antennae plerumque breviores. Oculi minuti. Pronotum setis 
appressis, aliisque erectis, vestitum, baud latum. Elytra angusta, elongata, lateribus 
saepe fere parallelis, plus minusve irregulariter argenteo- vel albido-squamosa, setisque 
erectis brevioribus conspicue vestita, parum conspicue inaequalia. Rostrum $ brevissi- 
mum, sulcis longitudinalibus sat fortibus. $ $. Long. 2 — 275 mm. 

Allied to P. blackburni Sh , but distinguished by the white or silvery appressed 
setae which cover parts of the elytra, the narrower form, shorter erect setae, etc. 

Hab. Kauai mountains (4000 ft.); a few examples were taken on several 
occasions. 

(17) Proterhinus setiger, sp. nov. 

Rufescens vel rufo-piceus, setis erectis sat crebre vestitus, statura majore. Antennae 
graciles, articulo secundo subelongato. Oculi prominuli. Pronotum antice abrupte 
angustatum, setis erectis vestitum, baud squamosum, grosse denseque punctatum, 
lateribus baud rotundatis. Elytra baud squamosa, setis erectis perconspicue vestita, 
inaequalia, lineis longitudinalibus elevatis setis appressis pallidis vestitis, grosse 
punctata, basi arcuatim sat fortiter emarginata, humeris acute productis, lateribus parum 
rotundatis. Rostrum % evidenter longius quam latius, longitudinaliter fortiter rugosum, 
lateribus basim versus levissime convergentibus. ^ ?. Long, vix 4 mm. 

A much larger species than P. blackburni, to which it is evidently allied. I have 
seen only one $ and one female, and the former has the prominent lateral angles 
of the prothorax, which are formed by its sudden constriction in front, much more 
strongly developed than those of the latter, and the puncturation more distinct. Each 
wing-case has three raised dorsal lines, one extending back from the humeral angles, 
another from the basal tubercles on either side of the scutellum, while between these on 
each side is a third elevation which does not however reach to the basal margin of 
the elytra. In the % the basal abdominal segment beneath is less coarsely punctate 
than the very coarsely and closely punctured metasternum ; the apical segments are 
impunctate or nearly so. 

Hab. Kauai (4000 ft.) ; i t and i % taken. 

(18) Proterhinus solitariiis, sp. nov. 

Piceus, setis erectis ex majore parte vestitus, baud squamosus, statura majore, baud 
robustus. Oculi minuti. Antennae graciles, articulo secundo elongato, subclavato. 
Pronotum antice abrupte angustatum et impressum, dense rugoso-punctatum. Elytra 



194 FAUNA HAWAIIENSIS 

lineis elevatulis longitudinalibus inaequalia, setis erectis vestita, basi leviter emarginata, 
humeris acutis. Femora postlca parum fortiter clavata, tarsorum lobis anticorum 
minutis. Rostrum $ brevissimum, fere aeque latum ac longum, fortiter longitudinaliter 
rugosum. $. Long. 3 •5 mm. 

Female in size and superficial appearance very similar to P. setigei'-, but differing in 
the much shorter rostrum, less deeply emarginate base of the elytra, smaller lobes to 
the front tarsi, etc. A single $ example, although taken at a higher elevation than the 
?, is probably referable to this species, since it agrees generally with the individual 
above described, but is entirely of a reddish colour. The lobes of the front tarsi are 
very minute, as in the ?. 

Hab. Kauai, a single $ taken at an elevation of from two to three thousand feet 
on the mountains. The $ mentioned above was taken on the high plateau of the same 
island. 

Obs. a single $ with the rostrum still slightly longer than that of P. setiger, the 
elytral puncturation less definite, and the basal abdominal segment beneath very closely 
and coarsely punctured all over, is very probably another species, while a $ from 
Molokai will not agree with any of these. It has the rostrum very short, but less so 
than that of P. solitarius. 



(19) Proterhimis squalidus, sp. nov. 

Atro-piceus vel sordide rufescens, antennis rufescentibus, setis brevibus erectis 
nonnullisque appressis vestitus, parum convexus. Antennae sat graciles, articulo 
secundo sat robusto, sed elongato, quam tertius vix breviore. Oculi minuti. Pronotum 
plus minusve impressum, sat latum, setis erectis brevibus, aliisque erectis, vestitum, 
confuse riigoso-punctatum. Elytra setis brevibus erectis, aliisque pallidis et depressis 
irregulariter (sive submaculatim), vestita, puncturatione parura distincta, humeris parum 
productis, fere rectis. Rostrum % brevissimum, tarsorum lobis anticorum minutis. 
Z ?. Long. 175 — vix 3 mm. 

Differs from P. antiqints in being of less narrow and elongate form, and generally 
in having the antennae longer and more slender. It appears to me to be really more 
closely allied to P. archaeus of Oahu, but that species has the elytra decidedly more 
elongate, and differs in various small points. The description given above has been 
made from examples from which the muddy substance, with which they are nearly 
always covered when captured, has been carefully removed. 

Hab. Kauai mountains (2000 — 3000 ft.). 



COLEOPTERA 195 



(20) Pi'oterhinus wikstroemiae, sp. nov. 

Subrobustus, nigricans vel rufo-niger, minus dense pallido-squamosus. Antennae 
elongatae, graciles, articulis omnibus (secundo plerumque excepto) sat elongatis, clava 
gracili, minus distincta. Oculi baud magni, sed fortius prominentes. Pronotum dense 
rugoso-punctatum, latera versus sat dense squamosum, parte reliqua nudiuscula, antice 
angustatum et impressum, lateribus subfortiter rotundatis. Elytra sparsim squamosa, 
punctata, fere aequaliter convexa, setisque albidis erectis brevibus parum conspicue 
vestita, angulis humeralibus fere rectis, minus distinctis. Femora sat fortiter incrassata. 
^ ?. Long. 3—3-5 mm. (Plate IX. fig. 4, ^.) 

This species is not at all closely allied to any other found on Kauai, nor indeed 
is it very similar superficially to any of the other species. I have no doubt it is 
nearest to P. dispar, although it differs from that species in numerous small points. 
Both appear to be solely attached to the same species of poisonous shrub, and a 
minute examination of the structural characters leaves no doubt that the two are more 
nearly related than would appear to be the case on a superficial examination. Nearly 
all the more remarkable characters of P. dispar are less developed in P. wikstroemiae. 

Hab. Kauai (4000 ft.), in various localities. Attached to Wikstroemia foetida. 



(21) Protcrliinus serricornis, sp. nov. 

Species parva, angustula, rufo-testacea, capite obscuriore, elytris utrinque plus minus 
nigro-signatis. Antennae breves, robustae, articulis brevibus et latis, clava distincta nulla. 
Pronotum squamulis pallidis minus dense vestitum, antice impressum. Elytra angusta, 
pallide squamosa, setisque nonnullis brevissimis erectis et parum conspicuis vestita, 
humeris baud productis. ^. Long, circa 2 mm. (Plate IX. fig. 6, J antenna.) 

This little species is quite distinct from any other (at least in the J' sex), by the 
form of the antennae, which are unusually stout and short. The intermediate joints are 
all of subrotundate form and so closely resemble the ninth and tenth that practically 
there is no club. The eyes and lobes of the front tarsi are both very small. In what 
I believe to be the ? of this species, the antennae are quite differently formed and by 
no means remarkable in structure, since they terminate in an evident three-jointed club. 
The intermediate joints, however, are rather less slender than in most species. The 
posterior femora are not very strongly clavate. 

Hab. Kauai, Halemanu, May 1895; three or four examples taken. 

F. H. II. 26 



196 FAUNA HAWAIIENSIS 



(22) ProterJiinus laticornis, sp. nov. 

Minor, angustulus, rufescens, capite femoribusque, et nonnunquam etiam pronoto, 
obscurioribus, elytris circa medium utrinque nigro-maculatis. Antennae ^ nigrae, basi 
rufa, ab articulo tertio vel quarto sensim usque ad apicem incrassatae, clavam quasi 
perelongatam formantes. Species pallida squamosa, faciem P. serricornis omnino 
habens, sed antennis longe aliter formatis, femoribus posterioribus valde incrassatis 
distinctissima. Z- Long. 2 — 2-25 mm. (Plate IX. fig. 5, <J.) 

This minute but very remarkable species is clearly allied to P. serricornis, with 
which it agrees in colour, clothing, and shape. Probably on the average it is of rather 
larger size. The joints of the antennae from the fourth or fifth become gradually wider 
to the apex, the whole forming as it were a very gradual and elongate club. The lobes 
of the front tarsi are minute, and the hind femora very strongly incrassate. The female 
which no doubt belongs to this species is quite unlike the <J in its antennal characters, 
since the intermediate joints are comparatively slender, and there is a distinct and 
strongly developed 3-jointed terminal club. This sex is therefore very similar to 
P. serricornis $, from which it may be known by the more strongly incrassate 
femora. 

Hab. Kauai, Halemanu (4000 ft.). 



(23) ProterJiinus eurhopahcs, sp. nov. 

Piceus, elytris plus minusve rufescentibus nigro- vel fusco-notatis, forma fere spp. 
2 praecedentium, sed plerumque colore obscuriore. Antennae ^ graciliores, basi rufa, 
clavae (distinctissime 3-articulatae) articulis latis. Femora postica minus fortiter 
incrassata. Z- Long, vix 2 — 2-2 mm. (Plate IX. fig. 7, Z antenna.) 

Closely allied to the two preceding species, the general form and appearance being 
the same, the colour perhaps more obscure. The Z has the antennae comparatively 
slender, indeed more nearly resembling the % of the preceding two species, and the club, 
consisting of three very distinct joints, is unusually developed. The eyes and tarsal 
lobes are small, the posterior femora not very strongly incrassate. Two females 
taken in company with these males are no doubt of the same species, resembling 
them very closely in all respects, except for the usual sexual differences in the 
rostrum. 

Hab. Kauai, Makaweli, at an elevation of 2000 — 3000 ft. 



COLEOPTERA 197 



(24) Proterhinns homocochronnts, sp. nov. 

P. eiii'hopalo forma et colore simillimus, sed paullo major, et antennis longioribus, 
clava paullo minus abrupta, capite inter oculos latiore distinguendus. ^ %. Long. 
2 — 2'6 mm. 

Very closely allied to P. cur/iopahis, but a rather larger species with decidedly 
longer antennae, and the club less marked, owing to its basal joint being less short and 
wide. The head is also decidedly wider between the eyes in the present species. 

Hab. Kauai, above Waimea (4000 ft.). 



(25) ProterJiinus cogJiatus, sp. nov. 

Elongatus, angustus, rufescens, capite obscurato, elytris utrinque nigro- vel fusco- 
notatis. Antennae graciles elongatae, articulis omnibus sat elongatis, clava gracili, haud 
abrupta. Pronotum subelongatum, antice angustatum. Elytra angusta, pallide squamosa, 
setisque nonnullis erectis perinconspicue vestita, basi leviter emarginata. Species forma, 
colore, et caeteris, P. homoeocJironw simillimus, sed paullo major, et antennis evidenter 
gracilioribus et magis elongatis, clava minus distincta, distinguenda. ^ $. Long. 
2"6 mm. 

Very closely allied to P. homoeochronius but of larger average size, the antennae 
elongate and very slender, the club decidedly more slender and less definite. 

The five species described above form a very remarkable series ; and although all 
were obtained from only a very small area of the island of Kauai, excepting perhaps 
the two Halemanu species, no two were found in the same locality. It is probable 
however that P. serricorms and P. laticornis were not actually obtained together, since 
two very distinct districts were collected over at Halemanu, and it is more than likely 
that each of these regions had its peculiar species. It may also be observed that the 
two species taken in the mountains behind Waimea occurred at different altitudes. 

Hab. Kauai, mountains above Waimea (3000 ft.). 



(26) Proterhinus angusti/ormis, sp. nov. 

Angustus, fortiter elongatus, plus minusve rufescens, pallide squamosus, elytris 
nigro-maculatis. Antennae graciles, articulis basalibus rufis, clava vi.x 3-articulata, 
articulis duobus apicalibus praecedenti multo majoribus. Oculi minores parum fortiter 
prominentes. Pronotum minus latum, pallide squamosum, lateribus minus fortiter 

26 — 2 



198 FAUNA HAWAIIENSIS 

rotundatis. Elytra rufa, nigro-maculata, pallide squamosa, setisque brevissimis erectis 
vestita, perelongata, humeri's vix productis, sed distinctis. Femora postica minus 
fortiter clavata. Abdominis segmentum primum ventrale ex majore parte vix evidenter 
punctatum. t ?• Long. 2 — 2-2 mm. 

A small and very narrow species remarkable for the great length of the elytra. 

Hab. Kauai, Halemanu. Four or five examples taken in May 1895. 



(27) Proterhinus leptophyas, sp. nov. 

Forma elongata et angustissima, nigricans, plus minusve rufo-variegatus, antennis 
rufis. Oculi minuti. Antennae breves, clava vix plus quam 2-articulata. Pronotum 
inaequale, antice angustatum, medium longitudinaliter subsulcatum, impressionibus 
duabus sublateralibus distinctis aut obsoletis. Elytra perelongata, angustissima, nigra, 
plus minus rufo-variegata, maculis rufis tantum squamis vestitis, parte nigra nuda, dense 
punctata, humeris hand productis. Femora gracilia. <? $. Long. 2 — 2*4 mm. (Plate IX. 
fig. 8, ?.) 

This small species is very distinct, and remarkable for its very narrow form and 
extremely elongate elytra. The antennae are unusually short, and entirely of a clear 
red colour, as also are the tarsi, and sometimes also the tibiae. The elytra usually bear 
a number of pale spots, which alone are covered with squamosity, the dark parts being 
bare. In the darkest individuals the elytral markings are reduced to a single spot at 
the extreme base, and such examples have nearly the whole surface of the elytra 
without squamous clothing. The hind femora are not at all strongly clavate. 

Hab. Kauai (4000 ft). Probably a local species, since all the specimens obtained 
were taken on the same day (April 15th, 1895). 



(28) Proterhinus linearis Blackburn. 

Proterhinus linearis Blackburn, Tr. Dublin Soc. iii. 1885, p. 169. 

The unique example of this species in the British Museum is in poor condition and 
I cannot identify it with any of those taken by me with certainty. 

Hab. Kauai (Rev. T. Blackburn). 

(29) Proterhinus maculifer, sp. nov, 

Angustulus, depressiusculus, piceus (nonnunquam nigricans vel rufescens), elytris 
maculis rotundatis squamiferis pallidioribus plus minusve variegatis. Oculi parvi. 
Rostrum Z haud brevissimum. Antennae graciles, breviores, clava baud abrupta. 



COLEOPTERA 



199 



Pronotum minus latum, antice impressum, minus dense squamoso-vestitum, lateri- 
bus parum fortiter rotundatis. Elytra angustula, maculatim squamosa, asperula et 
punctata (puncturatione et sculptura saepe exudatione lutosa celatis), setis erectis 
brevissimis nee conspicuis, angulis humeralibus plerumque fere rectis. Femora postica 
minus fortiter clavata, lobis tarsorum anticorum parvulis. ? %. Long, vix 2 — 3 mm. 
(Plate IX. fig. 9, t.) 

Chiefly remarkable for its flattened, or even sub-depressed form, and generally for 
the narrow elytra, which bear a number of roundish squamose spots, these spots 
being paler in colour than the rest of the surface, whether this be red or blackish. The 
sculpture is generally obscure, the punctures at least being generally filled up with a 
muddy excretion, and not infrequently the insect has nearly the whole surface thickly 
covered and concealed. The antennae, which are rather short, are generally entirely 
of a clear red colour ; the rostrum of the $ is also red or reddish, its surface smooth and 
shining, with the longitudinal grooves fine. 

Hab. Kauai, common generally. Many examples taken exhibiting variation in 
colour, size and other respects. 



B. Species insulae Oahu propriae. 

(30) Proterhinus kamptarthriis, sp. nov. 

Niger vel piceus, antennis tibiis tarsis elytrisque rufescentibus, his plerumque plus 
minusve nigro- vel fusco-maculatis. Antennarum articulo primo tertioque perelongatis, 
illo incrassato. Pronotum minus latum, 3-impressum. Elytra pallide squamosa, humeris 
productis. Antennarum $ articulus tertius subcurvatus. ^ ?. Long, circa 3 mm. 
(Plate IX. fig. 10, t.) 

Black or piceous, with red elytra, which are usually sparsely spotted with black. 
Femora dark, tibiae, tarsi and antennae wholly or partly red. Eyes prominent, in the ^ 
very remote from the thorax. Antennae long, extremely so in the J, the scape 
incrassate, and very elongate, second joint short, third extremely long, slightly but 
distinctly curved in the <y, and perceptibly so in the ?, club distinctly of three joints, the 
9th being considerably wider at the apex than the 8th. Prothorax rather narrow, with 
three more or less evident impressions, rather scantily covered with pale squamosity. 
Elytra covered with grey appressed scales, and with sparse erect white setae posteriorly, 
the sides sub-parallel. The humeral angles are strongly produced, and the basal 
tubercles near the scutellum are present, but there is no trace of longitudinal ridges. 
Lobes of the tarsi by no means large. 

Hab. Oahu, Waianae mountains (3000 ft.) ; three % taken in April 1892, and 
two males February 1896. 



200 FAUNA HAWAII ENSIS 



(31) Proterhinus leiorhynchus, sp. nov. 

Niger, vel piceus, antennis elytrisque rufescentibus, his nigro-maculatis, squamis 
pallidis vestitus. Antennae fortiter elongatae, conspicue pubescentes, articulo primo 
perlongo et incrassato, secundo sat elongate sed tertio breviore, 9° quam 10"' vix minus 
lato. Pronotum distincte 3-impressum, lateribus aequaliter rotundatis. Elytra pallide 
squamosa, setisque erectis elongatisque sat conspicue vestita, juxta scutellum utrinque 
tuberculata, humeris acutis. %. Long. 3 — 375 mm. 

This is a very distinct and somewhat large species, having the antennae unusually 
long. The basal joint is greatly incrassate and very long, the second is also elongate, 
although shorter than the third, the three apical joints form a very distinct club, the 
ninth at the apex being hardly less wide than the tenth, but very much more so than 
the long and slender joint preceding it. The rostrum is smooth and shining, the 
longitudinal impressed lines very fine. There is no transverse ridge between the 
vertex and front of the head. The eyes are prominent but not very large for the size 
of the insect. Thorax with the sides evenly rounded, densely punctured, and having 
three more or less distinct impressions. Elytra with pale squamosity, except over the 
black markings, and bearing elongate pale erect setae, which are quite conspicuous. 
The basal tubercles on either side of the scutellum are distinct, the humeral angles 
sharp and somewhat produced. Lobes of the tarsi somewhat small. 

Hab. Oahu, Waianae mountains (3000 ft.). Four females taken in February 1896. 
One of these has the second joint of the antennae abnormally short. There is also 
slight variation in the width of the thorax, and the elytra vary greatly in length. 

(32) Proterhinus ruficornis, sp. nov. 

Totus rufescens, aut capite, pronoto, femoribusque obscuratis, elytris nigro- 
maculatis. Antennae rufae, sat elongatae, clava distinctissime 3-articulata, articulo 9°, 
quam 8'", multo latiore, et aeque lato ac art. 10. Oculi magni fortiter prominentes. 
Pronotum antice abrupte constrictum, et impressum, impressionibus posterioribus ob- 
soletis. Elytra griseo-squamosa, humeris subproductis. Species haec P. laorhynclio 
cognatissima, antennarum articulis 2 basalibus minus elongatis, oculis et tarsorum lobis 
paullo majoribus, pronoti impressionibus posterioribus obsoletis, distinguenda. ^%. 
Long. 3—3-5 mm. 

Extremely closely allied to P. leiorhynchtis, from which it may be distinguished by 
the characters given above. Also very close to P. separandus, of Maui and Molokai, 
but that species has the antennae less elongate, and differs slightly in other respects. 

Hab. Oahu. A single pair were taken in cop. at an elevation of nearly 3000 ft., 
in the mountains near Honolulu. 



COLEOPTERA 



20I 



(33) Proterhinus deinops, sp. nov. 

Piceus, elytris rufescentibus, nigro-maculatis. Caput post oculos fortisslme con- 
strictum. Oculi fortissime prominentes, postice acuti. Antennarum articuli 3 apicales 
clavam distinctam formantes. Pronotum 3-impressum, sat elongatum. Elytra grosse 
et profunde punctata, basi utrinque juxta scutellum tuberculata, humeris acutis, fortiter 
productis. ^ $. Long, circiter 3 mm. (Plate IX. fig. 11, $.) 

One of the most remarkable species of the genus, having the head very strongly 
constricted behind the eyes, so that the front and vertex are separated by a strong 
ridge. The eyes themselves are excessively prominent, and are produced behind into 
a distinct point. The antennae are not long, the 9th joint is much wider than the 8th, 
so that the club is very definitely formed of three joints. The thorax is unusually long 
and narrow. The elytra have a deep coarse puncturation, and their humeral angles are 
very strongly produced, and acute. In fresh examples their red portions are covered 
with pale squamosity, which is absent on the black spots ; the erect setae appear to be 
few, and not conspicuous. 

Hab. Oahu, Waianae mountains, 3000 ft. Rare ; 6 examples were taken in 
February 1896. 

(34) Proterhinus squamicollis, sp. nov. 

Haud latus, niger, piceus, vel rufescens, pallide squamosus. Antennae graciles, 
clava distincte 3-articulata. Oculi magni, prominentes, coniformes. Capitis vertex a 
fronte cannula transversa distincte separatus. Pronotum haud latum, antice fortiter 
angustatum, et depressum, squamis pallidis ubique dense vestitum, impressionibus 
duabus posterioribus sat distinctis, lateribus fortiter rotundatis. Elytra distincte 
punctata, subinaequalia, squamis pallidis maculatim, necnon setis elongatis pallidis 
conspicue vestita, basi scutellum juxta utrinque tuberculata, tuberculis albido-squamosis, 
humeris productis et acutis. $%. Long. 2'25 — 3 mm. 

A distinct species, and readily distinguished from the other Oahuan Proterhini by 
the following combination of characters. Eyes large, prominent, and of conical form ; 
a distinct transverse ridge between the front and vertex of the head ; prothorax densely 
squamose all over, much narrowed and deeply impressed in front, the sides very 
strongly rounded ; elytra generally with more or less evident longitudinal ridges, with a 
tubercle on each side of the scutellar region at the extreme base, the shoulders produced, 
acute, and much deflexed ; their squamous covering is more or less broken up to form 
a number of spots, and the erect setae are long, fine, and quite conspicuous. 

Hab. Oahu. Widely distributed in the Koolau range, but rare. Ten or eleven 
examples taken at altitudes of 2000 — 3000 ft. 



202 FAUNA HAWAIIENSIS 



(35) Proterhimis adehis, sp. nov. 

Elongatus, parum convexus, nigricans, vel subobscure rufescens, elytris rufis, nigro- 
variegatis. Antennae modice elongatae, graciles, clava distincte 3-articulata, basim 
versus rufae. Vertex capitis a fronte baud divisus. Oculi vix magni, sed fortiter 
prominentes. Pronotum minus dense squamosum, antice angustatum, et profunde 
impressum. Elytra basim versus subangustata, maculatim pallide squamosa, parte 
basali rufa, postice nigro-maculata, setis erectis parce et parum conspicue vestita, dorso 
parum convexo, basi leviter arcuatim emarginata, humeris distinctis. Abdominis seg- 
mentum primum ventrale fortiter ubique punctatum, suturis ventralibus sat profundis. 
cT?. Long, circiter 3 mm. 

Of this apparently distinct species only a single pair, $ and ?, were taken. Apart 
from the usual sexual differences they are very similar, except that the </ being some- 
what abraded, has lost nearly all the curved setae from the sides of the thorax (which 
appears less abruptly narrowed in front), as well as the erect ones from the elytra. 
Otherwise it differs only in having the thorax and legs redder in colour, and the eyes 
slightly, but quite evidently, larger. 

Hab. Oahu. A pair were taken in the Koolau range at an elevation of 2500 ft. 
in the winter of 1892. 



(36) ProterJiinus obscuricolor, sp. nov. 

Nigricans, maculatim squamoso-vestitus, antennis pedibusque obscuricoloribus, 
P. vestito forma et magnitudine simillimus. Antennae ? graciles, clava minus abrupte 
3-articulata. Pronotum minus dense squamosum, rugoso-punctatum, 3-impressum, 
impressione anteriore magis profundo et majore, antice angustatum, haud abrupte 
fortiterque constrictum, lateribus sat rotundatis, setisque curvatis ciliatis. Elytra 
maculatim squamosa, setisque pallidis erectis gracilibus conspicue vestita, basi emar- 
ginata, angulis humeralibus distinctis. %. Long. 2-25 — 3 mm. 

Allied to P. vestitiis, but differing in colour, the shape of the prothorax, &c. 
The elytra are black, with a somewhat vague humeral or subhumeral spot of pale 
squamosity, and some other roundish spots behind these ; at the base they are formed 
much as in typical vcstitiis. A single female only of this species was taken, but there 
are two males in Dr Sharp's collection, which no doubt belong to the same. They 
differ from the ? in their antennae being shorter, and the club more distinct (i.e. the 
9th joint is more stout in proportion to the preceding), and they are considerably 



COLEOPTERA 203 

smaller, but are themselves of different size. The smaller example is a very fresh one 
and not quite mature, it being of a generally more pallid colour, and the legs and 
antennae are entirely red. These examples are marked as P. vesiitus, var. b and van c, 
and bear the number 352. 

Hab. Oahu. Mountains near Honolulu, 2000 — 3000 ft. ; i ? taken in the winter 
of 1892 ; 'mountains Oahu' (Blackburn). 



(37) Protcrhiiius denudatus, sp. nov. 

Elongatus, rufescens, elytris plus minusve nigro-notatis, parce aureo-squamosus. 
Antennae nigrae, articulis basalibus rufis, clava vix distincte 3-articulata. Oculi minus 
fortiter prominentes. Pronotum aequaliter, nee dense, aureo-squamosum, crebre 
punctatum, evidenter 3-impressum, antice haud abrupte constrictum. Elytra parce 
aureo-squamosa, setisque albidis brevioribus vestita, % fortissime elongata, fere aequaliter 
convexa, humeris acute productis. ^ %. Vix 3 mm. 

In the elongate form (especially of the $), in the colour and clothing this species 
is remarkably similar to Proterhinit.s detritus from Lanai. It lacks, however, the 
extremely abrupt constriction of the prothorax in front, the second joint of its antennae 
is less short, and they themselves are only red at the base. The rostrum of the ? 
detritus, however, is so different, being very coarsely sulcate, that it is doubtful whether 
the two are in reality at all closely allied. The t of the two examples examined is 
larger and wider, and much less elongate, than the ?, but I suspect that this is not 
usually the case, but rather an individual variation. A second male, taken in the other 
mountain range on Oahu, I refer at present to this species as a variety, but it is very 
likely a distinct species. It differs in being of a more obscure red colour, with the 
antennae entirely red, and is as narrow and elongate as the $, there are traces of whitish 
squamosity on the elytra, their erect white setae are longer, and they are faintly 
impressed at the base. 

Hab. Oahu. One of each sex taken in the mountains near Honolulu (2000 — 
3000 ft.) ; a variety of the t in the Waianae mountains. 



(38) Proterhinus compadus, sp. nov. 

Nigricans, elytris plus minusve rufescentibus. Caput cum pronoto aureo-squam- 
osum. Antennae mediocriter graciles et elongati. Pronotum obscure 3-impressum. 
Elytra albido-squamosa, setisque albidis erectis brevioribus sat conspicue vestita, fortiter 
denseque punctata, longitudinaliter subcarinata, basi scutellum juxta utrinque tuber- 
culata, humeris productis, minus acutis, basi thoracis adaptatis. Rostrum % fortiter 

F. H. II. ^7 



204 FAUNA HAWAIIENSIS 

longitudinaliter rugosum, ad basim distinctissime angustatum. ^ ?. Long. 2"6 — 
275 mm. 

Species, ut opinor, P. difficili (sp. Kauaiensi) cognatlssima, sed forma angustiore. 

Very similar to some examples of P. difficilis of Kauai, but distinguished by the 
evidently narrower elytra, which in lateral view are less convex longitudinally. The 
humeral angles are a good deal produced, but are not sharp, and are closely applied to 
the base of the prothorax, the puncturation is strong and dense. This species also 
bears a strong resemblance to the following i^P. platygonias), but the deep thoracic 
impressions, the extremely coarsely punctured elytra, and the strong constriction of 
the head behind the eyes, in the latter species, will easily distinguish it. 

Hab. Oahu, Waianae mountains (3000 ft.) ; two examples (^ and $) taken. 



(39) Proterhimis platygonias, sp. nov. 

Nigropiceus, pallide squamosus, elytris plus minusve rufescentibus et nigro- 
maculatis. Caput post oculos fortiter constrictum. Pronotum profunde 3-impressum, 
minus dense squamosum, grossius rugoso punctatum. Elytra inaequalia, grosse punc- 
tata, utrinque juxta scutellum tuberculata, humeris fortiter, nee acute, productis. 
%. Long. 2-5 — vix 3 mm. (Plate IX. fig. 12, $.) 

Black, the elytra reddish with black markings posteriorly, antennae red at the base 
or nearly altogether black. Head constricted behind the eyes, so that a ridge is formed 
between the vertex and front. Antennae not stout, nor long, the basal of the three 
terminal joints not very different to the 8th, the two terminal ones much wider, and 
short. Thorax very uneven owing to the three extremely deep depressions, its 
puncturation dense and rugose. Elytra with pale squamosity, and some rather short 
white erect setae, the surface uneven, the ridges which extend back from the humeral 
angles being evident at the base, and sometimes distinct to nearly the apex. On either 
side near the scutellum is a rather strong tubercle, covered with white squamosity, and 
between these and the shoulders the elytra are deeply impressed, as is readily seen in a 
lateral view. The humeral angles are strongly produced, but not very sharp, and 
closely applied to the base of the thorax. The puncturation is extremely coarse. 
Lobes of the front tarsi unusually small. 

Hab. Oahu, Waianae mountains (3000 ft.). Two female e.xamples taken in 
February, 1896. In one the antennae and tarsal lobes are black or nearly so, in the 
other (excepting the apical joints of the former) both are red. The latter example 
has the longitudinal ridges of the elytra less distinct, and the eyes decidedly less 
prominent. 



COLEOPTERA 205 

(40) Proterhinus vestittis Sharp. 

Protej-hinus vestitus Sharp, Tr. Ent. Soc. London, 1878, p. 16. Tr. Dubhn Soc. 
III. 1885, PI. V. fig. n^-] and 38. 

Typical examples of this species have a very distinctive appearance, ov^^ing to the 
extremely sudden constriction of the thorax in front. I have only seen one rather 
small example v^^hich quite resembles the types in structure, but specimens from the 
Waianae range have the thorax in some cases hardly less abrupdy constricted, and must 
be referred to the same species, although they form a well-marked local variety 
{^P. vcstitns, var. heterostictiis). These examples also have the setae of the elytra and 
legs generally less elongate, and the black or dark markings of the elytra are more 
lateral in position. The species is closely allied to P. eulcpis but the elytra are in 
general less wide and short, the shoulders less produced, and the erect setae longer. 

Hab. Oahu, mountains near Honolulu (Blackburn) ; Waianae mountains, both 
sides. 

(41) Proterhinus robiistus Blackb. 

Proterhimis robusttis Blackburn, Tr. Dublin Soc. iii. 1885, p. 171. 

Extremely close to P. vestitus but probably distinct. Like that species it is 
variable, but in its typical form it differs from typical examples of P. vestitus, in having 
the thorax but little constricted in front and less narrowed behind, the hind angles 
distinctly marked by a spot of pale squamosity. The elytra have shorter and less 
conspicuous erect setae, those on the legs being also much shorter. The basal joint of 
the antennae is unusually long, being fully as long as the two following joints together. 
Apart from the pale spots at the hind angles the thorax has very little squamosity, so 
that the puncturation is quite conspicuous, being very dense, and somewhat coarse. 
The curved setae at the sides are short and very feebly developed. I have not seen 
a ^ of this species. Length 2 — 3 mm. 

Hab. Oahu, Waianae mountains, found in the lowest forest. Only a few female 
examples were collected. 

(42) Proterhinus sicbplanatus, sp. no v. 

Nigricans, parum dense squamosus, P. anguiai'i cognatissimus, antennis plerumque 
brevioribus, minus gracilibus, forma magis depressa (sive deplanata), distinguendus. 
^ $. Long. 2 — 2 '5 mm. 

A somewhat obscure species, evidently closely allied to P. angularis. From 

27 — 2 



2o6 FAUNA HAWAII ENSIS 

typical individuals of that species it is readily distinguished by the much less con- 
spicuous setae on the elytra, and the hardly visible curved setae at the sides of the 
prothorax. From the less typical examples which I have referred to P. angtdaris, it may 
be known by the shorter and less slender antennal joints, and its evidently flattened, or 
subdepressed form. 

The shoulders of the elytra are formed as in P. angtdaris, and are likewise marked 
with pale squamosity, but as a rule less conspicuously, and the basal ventral segment of 
the abdomen is strongly punctate. 

Hab. Oahu, Halemano (Koolau range) (2000 ft.), in January, 1893. 



(43) Proterhimis simplex Sharp. 

Proterhimis simplex Sharp, Tr. Ent. Soc. London, 1878, p. 17. 

The examples from which this species was described, appear to me to be not only 
immature, but also particularly small and ill-developed specimens. Large individuals 
are twice the size of the specimens sent by Mr Blackburn, and altogether wider and 
more bulky, the colour pitchy or reddish pitchy, the elytra red, with dark markings 
laterally, about the middle, and extending sometimes to the suture. The squamosity 
is not continuous, there being an oblique spot near the shoulders of the elytra and 
others behind these. The setae are shorter, and less conspicuous, than those of fresh 
examples of P. obscuricolor, vestitus and others. The prothorax is much wider, and has 
the sides much more strongly rounded than in the type, and is often distinctly 3-im- 
pressed. The eyes, the tarsal lobes, and humeral angles of the elytra are often all 
much more developed, but all of them vary, even in the larger examples. Length 
2 — nearly 3 mm. 

Hab. Oahu. Both mountain ranges, at elevations of 2000 — 3000 ft. 



(44) Proterhimis oxygonias, sp. nov. 

P. scutato forma facieque simillimus, colore plerumque minus obscuro, elytrorum 
angulis humeralibus fortius et acutius productis, distinguendus. ^%. Long. 2*25 — 
275 mm. 

This species appears to be very closely allied to P. scutatus, from which it differs 
generally by its less obscure {i.e. more red) colour, the less evenly rounded sides of the 
thorax, and the more strongly and acutely produced humeral angles of the elytra. 

Hab. Oahu, mountains near Honolulu (2000 ft.). A single example taken in the 
Waianae range of the same island appears to be a variety of this species. 



COLEOPTERA 207 



{45) Proterhimis leptothrix, sp. nov. 

Haud latus, rufescens, elytris plus minus nigro- vel fusco-signatis. Oculi minores. 
Antennae mediocres, clava 2-articulata, articulo 9° vix quam 8"' evidenter latiore. Pro- 
notum parum squamosum, dense rugoso-punctatum, setisque erectis gracillimis versus 
latera sat conspicue vestitum, antice impressum, angulis posticis macula pallida squamosa 
signatis. Elytra plus minusve albido-squamosa, setis erectis gracillimis et elongatis 
conspicue vestita, humeris acutis, plus minusve productis. Pedes setis conspicue 
elongatis vestiti, tarsorum anticorum lobis parvis. $. Long. 2 — 2 "5 mm. (Plate IX. 

% 13. ?■) 

A very distinct species, owing to the form of the antennae, and the e.xtremely fine 
and more than usually elongate setae, which clothe the elytra and legs. The club of the 
antennae can hardly be considered as consisting of more than two joints, as the 9th 
in whatever aspect it is viewed is hardly different in form to those that precede it. 
A single example taken in the same locality as the $ above described is no doubt the $, 
as it differs only in being smaller and in the usual se.xual characters, but it is a good 
deal abraded. 

Hab. Oahu ; Kaala (3000 ft.), Waianae range ; 2 ? taken together in December, 
1892, and a single $ in the same month but on another occasion. 



(46) Proterhinus seticollis, sp. nov. 

Praecedenti cognatissimus, sed colore obscuriore, setisque erectis elytrorum minus 
longis gracilibusque distinguendus. ^ $. Long. 2 — 2*75 mm. 

This species is very closely allied to P. leptothrix, and may possibly on the 
examination of a long series prove to be a form of that species. It is very easily 
distinguished, so far as the few examples taken are concerned, by the different clothing, 
the erect setae being much less elongate and slender. It is also generally of a darker 
colour, and superficially greatly resembles P. scutatiis, from which it is abundantly 
distinct by the 2-jointed club of the antennae, the stronger longitudinal furrows of the 
rostrum in the ?, the patch of white squamosity at the hind angles of the thorax, which 
bears conspicuous erect setae, &c. 

Hab. Oahu, Waianae mountains (3000 ft.) ; a few examples taken on the opposite 
side of the range to the locality for P. leptothrix. 



2o8 FAUNA HAWAIIENSIS 



(47) Proterhinus longulus Sharp. 

Proterhimis longulus Sharp, Tr. Ent. Soc. London, 1S79, p. 97. 

I have examined only the types, one t and one %, of this species, which are in 
Dr Sharp's collection. It is very closely allied to P. epitretus of Lanai, and P. ferru- 
gineus of Hawaii, under which species I have pointed out the characters whereby these 
allied forms may be distinguished. 

Hab. Oahu mountains, in stems of fern (Blackburn). 



(48) Proterhinus paradoxus Sharp. 

Proterhimis paradoxus Sharp, Tr. Ent. Soc. London, 1879, p. 100. Tr. Dublin 

Soc. III. 1885, PI. V. fig. 36. 
I have never met with this species, which is evidently allied to the following 
(T-". oahuensis). 

Hab. Mountains near Honolulu (Blackburn). 



(49) Proterhinus oahuensis, sp. nov. 

Haud latus, nitidus, ferrugineus, setis erectis flavescentibus vestitus, hand squa- 
mosus. Antennae graciliores, articulis 2 basalibus sat robustis. Oculi parvi, a pronoto 
longe remoti. Pronotum subnitidum, antice fortiter impressum, impressionibus posteri- 
oribus minus distinctis. Elytra nitida, sat elongata, grosse subseriatim punctata, suturam 
versus utrinque oblique longitudinaliter carinata, carina quoque longitudinali ab angulis 
humeralibus extensa, setis erectis flavescentibus elongatis, aliisque brevioribus, conspicue 
vestita. Femora omnia gracilia, tibiis anticis gracillimis et elongatis, lobis tarsorum 
minutissimis. ^. Long. 2 '5 mm. (Plate X. fig. 4.) 

This species is very distinct, and obviously allied to P. paradoxus, but has 
the elytra of different form, these not being rectangularly deflexed laterally, nor con- 
spicuously flattened above. The shoulders are nearly rectangular, and from each a 
distinct carina extends backwards, and internal to these towards the suture other two of 
the interstices are distinctly raised and form obliquely longitudinal carinae. The basal 
abdominal segment beneath is very coarsely punctured and the segmental sutures are 
extremely deep. 

Hab. Oahu, mountains near Honolulu; i t taken in May, 1896. 



COLEOPTERA 209 

(50) Pi'oterhinus kaalae, sp. no v. 

Piceo-niger, antennis tarsisque rufotestaceis, minutus, sat robustus. Antennae 
graciles, elongatae, articulo primo fortissime incrassato, clava distincte 2-articulata. 
Oculi parvi. Pronotum grosse punctatum, setis erectis pallidis vestitum, haud squa- 
mosum. Elytra brevia, lata, pronoto fere bis latiora, grossissime punctata, haud 
squamosa, setis per elongatis albidis supra sex-seriatim vestita. Femora omnia cum 
tibiis setis longis albidis vestita, tarsorum lobis anticorum sat magnis. Z- Lono-. vix 
2 mm. (Plate X. fig. 3.) 

One of the smallest, but quite one of the most remarkable of all the known species. 
There is no squamous clothing, but the erect setae are whitish, and very long and 
conspicuous, and form three rows on each wing-case, those of the series next to the 
suture being shorter than the others. The setae in each row are distant and very 
regular, and of even length, not consisting of some short and some long ones intermixed, 
as in most of the conspicuously setae species. Similar setae clothe the legs and thorax, 
and the sculpture of the latter and of the elytra is very coarse. The lobes of the front 
tarsi are rather large for the size of the insect, the claw-joint short, not projecting very 
far beyond the lobes. The basal joint of the antennae is extremely wide, the inter- 
mediate joints slender, the club consisting of 2 joints only. The elytra are short and 
wide. 

Hab. Oahu. A single t was taken on Kaala in the Waianae mountains. 

(51) Proterhimis archaetts, sp. nov. 

Depressus, nigricans vel piceus, parum squamulosus, setis conspersis erectis sat 
conspicue vestitus. Antennae rufae, articulo secundo sat elongate sed, quam tertius, 
robustiore. Oculi prominuli, setigeri. Pronotum dense rugoso-punctatum, setis erectis 
elongatis vestitum. Elytra parum vel haud squamosa, setis erectis sparsim sed con- 
spicue vestita, crebre confuseque punctata, dorso deplanato vel depresso, basi utrinque 
subtuberculato, angulis humeralibus distinctis, fere rectis. Femora minus fortiter 
clavata, tarsorum anticorum lobis parvis. Abdominis segmentum primum ventrale 
grosse et sat crebre punctatum. Rostrum % breve, vix longius quam latius, fortiter 
longitudinaliter rugosum. ^ %. Long. 2'2 — 3"2 mm. 

Allied to P. blackburni but quite distinct by its depressed or flattened form, 
less numerous erect setae, &c. 

Hab. Oahu. Koolau range behind Waialua. A single example (var. diversus), 
taken in the same range near Honolulu, has the elytra less flattened, the thorax strongly 
transverse, and the erect setae of the elytra decidedly shorter. It will probably prove 
to be a distinct species. The typical form lives beneath the bark of Straussia. 



2IO FAUNA HAWAIIENSIS 

(52) ProterJiimis obsciirus Sharp. 

Proterhimis obsairus Sharp, Tr. Ent. Soc. London, 1878, p. 18. 

This species was described on a single example of the female, but subsequently in 
Trans. Dublin Soc. Vol. iii, 1885, was referred in the systematic Catalogue (p. 250) to 
P. simplex as a variety. I believe it to be distinct from that species, but both varying 
greatly, it becomes impossible to form an adequate description that will apply to all the 
individuals of each. Certainly in what I consider to be well-developed examples of the 
two the species are readily distinguished. 

In its well-developed and typical form the ^ of P. obscitrus is shorter and wider 
than the female, and generally rather darker in colour. The antennae are moderately 
long, but the intermediate joints are not very slender, and the first of the three club 
joints is not very different to the following. The basal joint is short and very stout. 
Eyes but little prominent. Thorax wide and very strongly rounded at the sides, with 
the squamosity forming dense lateral patches, but sparser on the dorsum. There is an 
evident anterior impression, but the curved erect setae at the sides are altogether 
absent, or very indistinct. The elytra vary in colour from nearly entirely black, to a 
dull red with dark markings, and are rather scantily squamose, and bear, especially 
posteriorly, some short, pale, erect, but inconspicuous setae. The humeral angles are 
not acute, nor very distinct, and are sometimes quite effaced. All the femora are 
strongly incrassate and clavate, and the lobes of the front tarsi are moderately large. 

The female is generally more slender, the thorax less wide, the elytra longer, and 
their colour less dark, and the squamous covering more regular. 

In the Waianae mountains there is found what appears to be a dwarf form of the 
above species, the smallest examples of which are very minute, and most of the structural 
characters are accordingly modified. 

Hab. Oahu. Koolau range (1500 — 3000 ft.); small van in the Waianae moun- 
tains at similar elevations. 

(53) Proterhiims oscillans Sharp. 

Proterhimis oscillans Sharp, Tr. Ent. Soc. London, 1878, p. 18. 

As I understand it, this is not a very variable species, and I have examined 
numerous examples that agree well enough with the types. As a rule the species is 
constant in colour and may be recognised easily by this. Small examples have the 
tarsal lobes diminished in size, and in some the humeral angles of the elytra are less 
indistinct than in others, and the elytra do not always become distinctly narrowed 
towards the base. The rostrum of the $ is normally very decidedly narrowed from the 
apex to the base. Length from r5 to 275 mm. 

Hab. Oahu. Both mountain ranges. Not rare from 1500 — 3000 ft. 



COLEOPTERA 211 



(54) Proterhinus pachycnemis, sp. nov. 

Rufescens, elytris nigrosignatis. Antennae mediocres, articulo secundo, quam 
tertius, haud minus longo, sed multo crassiore, clava sat evidenter 3-articulata. Prono- 
tum latera versus sat dense squamosum, lateribus minus fortiter rotundatis. Elytra 
pallide squamosa, setisque pallidis erectis conspersim vestita, humeris subobscuris, haud 
productis. Femora antica et postica miro modo incrassata, intermediis multo crassiora. 
t. Long. 2 mm. (Plate IX. fig. 14. ^.) 

This species, which somewhat resembles P. oscillans or P. obscnrus, is quite distinct 
by the elongate second joint of the antennae, and the extraordinarily incrassated femora, 
the hind femur being about twice as wide as the intermediate, which itself is more 
robust than in most of the species. A second and much damaged ^, which I refer to 
this species, is totally different in superficial aspect, being black, with only small parts 
of the thorax and elytra obscurely red, and the latter less nearly parallel-sided. 

Hab. Oahu. Kaala (2500 ft.), Waianae range. Two ^ examples taken, one on 
the windward and the other the opposite side of the range. 



(55) Proterhinus laticollis, Blackb. 

Proterhimcs laticollis Blackburn, Tr. Dublin Soc. in. 1885, p. 170. 

(Plate IX. fig. 15, and 15a. Z-) 

The male of this species is a very remarkable insect, not only on account of its 
extremely wide thorax, as indicated by Blackburn in his description, and by the name 
which he applied to it, but still more so by the structure of the terminal joint of the 
front tarsi, which is unlike that of any other of the genus. This joint is so extremely 
shortened that it only just extends as far as the apex of the lobes of the preceding one. 
The antennae are more than usually elongate and slender, and the front femora very 
strongly incrassate, as compared with the hinder ones. In some examples the elytra 
are quite evidently impressed across the base, but in others no trace of this is to be 
detected. The $ of P. laticollis, on the other hand, is comparatively commonplace, 
since neither thorax nor tarsi present the peculiar features exhibited in the $. It is 
only to be recognised by its general similarity in colour, clothing, and the elongate and 
slender antennae. As in the Z, the base of the elytra is sometimes sub-impressed. 
Length 2 — 2'5 mm. 

Hab. Waianae mountains, Oahu. About 17 examples taken. 
F. H. ir. 28 



212 FAUNA HAWAIIENSIS 

(56) Proierhhms vicimis, sp. nov. 

Species parva, angustula, rufescens, capita, necnon saepe pronoto plus minusve 
infuscatis, elytrorum marginibus lateralibus, et nonnunquam maculis quibusdam dorsali- 
bus, niericantibus. Antennarum $ articulus basalis robustus, sed minus brevis, hoc 
cum sequentibus rufo, articulis apicalibus nigris, clava distincte 3-articulata. Pronotum 
baud latum, antice impressum, aequaliter (vix dense) squamis vestitum. Elytra angusta, 
lateribus saepe fere parallelis, pallide, baud maculatim, squamosa, setisque brevibus 
albidis erectis parce inconspicueque vestita, angulis humeralibus baud productis. Oculi 
et tarsorum anticorum lobi parvi. $ ?. Long, vix 2 — 2-25 mm. 

The affinities of this insect are with five of the Kauai species, P. serricor7iis and 
the four following, rather than with any other of the Oahuan forms. With those species 
it agrees in general form, colour, and clothing, as also in the very short rostral portion 
of the head in the t, and, considering its minute size, in the rather strongly developed 
basal joint of the antennae. It cannot however be referred to any of those species, for 
the antennae, which are neither long nor yet particularly slender, and of which neither 
the 3-jointed club, nor the intermediate joints present any special features, will agree 
with none of them. The sexes apparently are almost similar, except for the usual 
sexual differences. 

Hab. Oahu. Taken only in the Waianae mountains, and not numerously. 

(57) Proterhinus pusillus Sharp. 

Proterhinus pjisillus Sharp, Tr. Ent. Soc. London, 1879, p. 97. 

This is a very minute and obscure insect, and may not improbably prove to be a 
depauperated form of some other species. It belongs to the most difficult and obscure 
section of the genus, and has no striking characters of any sort. It appears to approach 
closely to some varieties of P. deceptor, and also to minute examples of P. obscurus. 

Hab. Oahu mountains near Honolulu, 1500 — 2000 ft. (Blackburn.) 



C. Species insulae Maui propriae. 

(58) Proterhinus validus Sharp. 

Proterhinus validus Sharp, Tr. Ent. Soc. London, 1881, p. 531. 
(Plate IX. fig. 16. i.) 

The antennae in this species are decidedly more elongate and less thick in some 
examples than in others. The length varies from 4 — 6 mm. 

Hab. Maui, Haleakala (4000 — 5000 ft.) ; many examples taken on Acacia koa. 



COLEOPTERA 2 1 3 

(59) Proterhinus comes, sp. nov. 

Major, piceus, vel obscure rufescens, subbrunneo-squamosus, pronoto elytrisque 
setis erectis crebre conspicueque vestitis. Oculi sat magni, fortiter prominentes. 
Antennae graciles, articulo secundo brevissimo, clava gracili, elongata, 3-articulata. 
Pronotum antice fortiter angustatum et transversim impressum, impressionibus pos- 
terioribus rotundatis et distinctis. Elytra dense punctata, longitudinaliter subcari- 
nata, margine basali scutellum juxta utrinque tuberculato, humeris fortiter acute 
productis. Pedes omnes conspicue setosi, lobis tarsorum anticorum minoribus. ^. 
Long. 3-5—4 mm. 

Apparently closely allied to P. validus, but a considerably smaller insect 
and of less elongate and oblong form. The antennae are quite differently formed, 
the intermediate joints being slender and elongate. The eyes also, and the lobes 
of the front tarsi, are less developed, and the longitudinal ridges of the elytra 
generally more obscure. 

Hab. Maui. Three examples of the $ taken on Haleakala (4500 — 5000 ft.) ; 
it is probably a variable species, as these examples do not altogether agree in 
small points. 



(60) Proterhinus lecontei Sharp. 

Proterhinus lecontei Sharp, Tr. Ent. Soc. London, 1879, p. 99 ; Tr. Dublin 

Soc. III. 1885, PI. V. fig. 42 ?, fig. 43 t. 
Varies greatly in size, some examples being only about half the bulk of a normal 
specimen. Length (excluding rostrum) 3*2 — nearly 5 mm. 

Hab. Maui, Haleakala (5000 ft.) ; not rare, but only on one species of tree, of 
which I neglected to obtain a specimen for identification. 



(61) Proterhinus sharpi, sp. nov. 

Ferrugineus, elytris saepe plus minusve nigricantibus, ferrugineo-squamosus. 
Antennae rufescentes, sat graciles. Oculi fortiter prominentes. Pronotum minus 
latum, dense squamoso-vestitum, 3-impressum, antice angustatum. Elytra suboblonga, 
lateribus subparallelis, utrinque longitudinaliter carinata, carinis postice abrupte trun- 
catis, humeris sat fortiter productis. $ %. 2"6 — 375 mm. (Plate IX. fig. 17. $.) 

In the form of the elytra this species can only be compared with P. lecontei, 
the longitudinal ridges of the elytra being well-marked and abruptly vertical behind, 
so that in a posterior view each appears as a strongly elevated tubercle. In 

28—2 



214 FAUNA HAWAIIENSIS 

general appearance it is very different to lecoKtei, but in reality the two are really 
related. 

Colour ferruginous, as also the squamosity, the elytra sometimes partly or even 
entirely black. Eyes large, strongly prominent ; antennae slender, with evidently 
3-jointed club, generally entirely of a clear red colour. Prothorax subelongate, densely 
squamose, with three well-marked impressions, at the sides with some curved suberect 
setae. Elytra elongate, subparallel-sided, tuberculate at the base on either side of 
the scutellar region, depressed between these tubercles and the produced humeral 
angles, flattened or even somewhat depressed dorsally between the well-marked longi- 
tudinal ridges, which terminate abruptly behind. When the elytra are reddish in colour 
they have a ferruginous squamosity, but when their colour is black the squamosity dis- 
appears ; erect, thin setae are also scattered over the surface, especially at the sides 
and posteriorly. Lobes of the front tarsi somewhat large. 

Hab. Maui, Haleakala (5000 ft.), in the same localities as P. lecontei, but with 
very different habits, this species being found only on a species of fern, while P. lecontei 
frequents one of the forest trees. 



(62) Proterhinus tuberculiceps, sp. nov. 

Rufescens, elytris plus minusve nigro-maculatis, antennis, pedibusque omnibus 
rufis. Caput inter oculos fortiter bituberculatum. Oculi fortiter prominentes. 
Antennae graciles, elongatae, clava distincte 3-articulata, sat elongata. Pronotum 
perinaequale, fortissime 3-impressum, impressione anteriore transversa, antice abrupte 
constrictum. Elytra longitudinaliter subcarinata, setis erectis gracillimis, sat conspicuis, 
vestita, humeris fortiter productis. Z%- Long. 3*5 mm. (Plate IX. fig. 18. Z-) 

A very distinct species, entirely reddish in colour, except for some small, and more 
or less inconspicuous, dark spots on the elytra. The eyes are strongly prominent, and 
there is a strong tubercle close to the inner margin of each. The prothorax is very 
uneven, the anterior impression extending right across it, while the two posterior ones 
are deep, but smaller, and roundish. In front the constriction is strong and abrupt, 
and there and at the sides the suberect, curved setae are very evident. The lobes of 
the front tarsi are well developed. The elytra have their shoulders strongly produced, 
the suture a little raised, and a sufficiently evident longitudinal ridge running backwards 
from the humeral angles of each. The erect setae are long, fine, and conspicuous. All 
the femora are strongly clavate, a little more so in the ^ than in the ?. The rostrum 
of the % is only moderately long, and is smooth and shining, with the longitudinal 
grooves fine. 

Hab. Maui, Haleakala (4000 ft.) ; 2 ^, i ? taken. 



COLEOPTERA 215 

(63) Proterlmms sternalis Sharp. 

Proterhintis sternalis Sharp, Tr. Ent. Soc. London, 1879, p. 98. 

(Plate IX. figs. 19 and 19 a. t) 

This species is remarkable for the great developement of the eyes, and tarsal lobes, 
the concavity of the prosternum from the front to the base, and the unusual clothing of 
the intermediate tibiae, which bear long and soft hairs, instead of the usual short and 
stiff setae. The ridges of the elytra are quite well-marked, especially when these are 
observed in lateral view. Length from 2 '2 — 2 '3 mm. 

Hab. Maui, Haleakala (4000 — 5000 ft.) ; not very common, only about 20 
examples were taken. 

{64) Proterhintis sternalioides, sp. nov. 

Sat latus, nigricans vel rufo-niger, antennis, pedibus, elytrisque plus minusve 
rufescentibus, his riigro-notatis, praecedenti sp. cognatissimus. Oculi magni. Prono- 
tum antice constrictum, fortiter 3-impressum, minus latum. Elytra subinaequalia, 
pallide squamosa, setisque erectis elongatis vestita, basi scutellum ju.xta utrinque 
tuberculata, humeris fortiter productis. $%. Long. 2'2 — 375 mm. 

This species is extremely like P. sterna/is, which it resembles in colour, form, 
&c. The longitudinal ridges of the elytra are rather less pronounced, and the face 
of the $ (including the rostrum) is evidently more elongate. The rostrum of the 
$ is slightly longer, its surface slightly rugulose all over. The intermediate tibiae 
are clothed with shorter setae, and the lobes of the front tarsi of the ^ are of rather 
different form, being rather more narrow and elongate, the whole joint having a less 
quadrate form. The hind coxae of the d' are distinctly more widely separated, and the 
prosternum is impressed only along its anterior margin, not longitudinally sulcate down 
the middle from front to base as in Z'. sternalis. 

Hab. Maui, Haleakala (5000 ft.) ; several examples taken, probably in company 
with P. sternalis. 

(65) Proterhinus microtarsus, sp. nov. 

Atro-piceus, elytris rufescentibus, nigro-notatis, antennis pedibusque totis rufis. 
Antennae elongatae, graciles, articulo i elongato sed robusto, quam tertius longiore, 
caeteris elongatis, clava gracili, distinctissime 3-articulata. Oculi magni. Vertex a 
fronte capitis haud distinctus. Pronotum perinaequale, antice subito fortiterque con- 
strictum. Elytra squamulosa, longitudinaliter subcarinulata (lateraliter compressa), setis 



2i6 FAUNA HAWAIIENSIS 

pallidis elongatis sat conspicue vestita, humeris fortiter productis. Tibiae anticae 
graciles, elongatae ; femora omnia fortiter incrassata, lobis tarsorum anticorum minimis. 
cT. Long. 3 mm. (Plate IX. fig. 20 and 20a. rf".) 

This remarkable species is very distinct from any other. The antennae are long 
and slender, concolorous with the legs, of a clear rufotestaceous colour. The basal 
joint is elongate but rather stout, all the rest are elongate and slender, as also is the dis- 
tinctly 3-jointed club. Prothorax very abruptly constricted, and transversely impressed 
in front, the impression deeper about the middle. The two posterior sublateral impres- 
sions are also distinct, and behind these the middle part of the thorax is seen (especially 
in lateral view) to be somewhat strongly elevated, the elevation subsulcate. All the 
femora are strongly incrassate, the front tibiae unusually long and slender, the lobes of 
the front tarsi excessively small for the size of the insect. Elytra subparallel-sided for 
their basal half with pale squamosity, and conspicuous, very fine, erect, pale setae. 
Seen from the side they are somewhat compressed laterally, to form the rather obscure 
longitudinal ridges. Their humeral angles are strongly produced. 

A female taken in the same locality as the c/ no doubt belongs to the same 
species, although at first sight of very different appearance. The antennae are formed 
like those of the <?, the eyes are smaller but more prominent. Rostrum short, not 
nearly twice as long as wide, the surface entirely rugulose. The whole insect is 
blackish, except the reddish tibiae and tarsi and the greater part of the antennae, the 
club-joints being obscure in colour. The prothorax is less elevated posteriorly than 
that of the ^, and the elevated portion is not evidently sulcate. The elytra are a good 
deal longer, entirely black, the sides nearly parallel except at the ape.x. Femora 
strongly clavate as in the Z, the front tibiae also long, and the lobes of the tarsi small. 

Hab. Maui, Haleakala (4000 ft.) ; rare, only one <S and one % having been taken. 



(66) Protei'hinus megalotarsiis, sp. nov. 

Nigricans, elytris rufescentibus vel piceis, pedibus antennisque saepe plus minusve 
rufescentibus. Oculi sat magni, subconici, fortiter prominentes. Antennae mediocres, 
clava distincte 3-articulata. Pronotum pallide squamosum, profunde 3-impressum, vix 
latum (feminae elongatum), lateribus rotundatis. Femora ^ fortiter incrassata, lobis 
tarsorum anticorum magnis. Elytra pallide squamosa, setisque gracilibus erectis sat 
conspicue vestita, fortius (nee inaequaliter) convexa, sutura subelevata, humeris fortiter 
acute productis. ^ $. Long. 2-8 — 3-25 mm. 

A rather distinct species, at least in the ^, but apparently variable, the antennae 
being a good deal shorter, in some examples than in others. There is no distinct ridge 
between the vertex and front of the head in either sex. The eyes are large, very 



COLEOPTERA 



217 



prominent and subconical. In the Z the lobes of the front tarsi are largely developed 
and wide, but evidently less so in the female, in which sex the rostrum is smooth and 
shining, with two fine longitudinal grooves. The elytra have only faint, sometimes 
indeed no traces of longitudinal ridges, but the suture itself is quite evidently raised. 

Hab. Maui, Haleakala ; a few examples taken at and below an elevation of 
4000 ft. 

(67) Protcrhinus heniichlorus, sp. nov. 

Nigricans, pallide squamosus, antennis pedibus elytrorumque basi plus minusve 
rufescentibus, P. /^/«;/t77r// cognatissimus. Antennae, oculi, tarsorumque anticorum lobi, 
mediocres, haudquaquam insignes. Pronotum minus dense squamosum, setis curvatis 
parum conspicuis, 3-impressum, impressione antica transversa, duabus posticis distinc- 
tissime rotundatis. Elytra plus minusve nigricantia, plaga basali conspicue pallido- 
squamosa parceque nee fortiter punctata, sutura subelevata, humeris productis. Femora 
postica haud fortiter incrassata. t%. Long. 17 — 2-8 mm. 

A small species with rather a distinctive appearance, owing to a patch of dense 
pale, often nearly white, squamosity, which covers the basal third or even half of the 
elytra, over which part the puncturation appears sparse and feeble. The erect setae are 
chiefly placed on the hinder part of the elytra, but they are sparse and not very con- 
spicuous. The femora are only moderately robust. Closely allied to P. hmieralis 
but distinguished by the less developed tarsal lobes, eyes, and antennae, the less 
incrassate femora, and other less important characters. As in that species the elytra 
are without evident longitudinal ridges, and there is none between the front and vertex 
of the head. 

Hab. Maui, Haleakala (5000 ft.) ; a few examples of each sex taken. 



(68) Proterhinus humeralis Sharp. 

Proterhimis humeralis Sharp, Tr. Ent. Soc. London, 1879, p. 96. 

A rather distinct species, at least in the ^. Apparently allied to P. sternaiioides, 
but a decidedly narrower and more elongate insect. The antennae of the ^ are rather 
longer and at the same time stouter than in most of the species, and the lobes of the 
front tarsi are strongly developed, although not to the extent of those of P. sierna/is. 
In both sexes, but especially so in the $, the femora are more strongly incrassate than 
is usual, but all the specific characters of the ? are much less pronounced than in the $. 
There is no ridge formed between the front and vertex of the head. 

Hab. Maui, Haleakala (4000 — 5000 ft.); I have seen only the two type specimens, 
and two or three others taken by myself. 



2i8 FAUNA HAWAIIENSIS 



(69) P roterhinus bi'evipennis, sp. nov. 

Rufescens vel rufo-piceus, antennis pedibusque totis, vel ex parte, rufis, pallide 
squamosus, forma breviore. Antennae graciles. Oculi prominentes. Pronotum sat 
dense aureo-squamosum, antice fortiter angustatum, aut nonnumquam abrupte con- 
strictum, profunde 3-impressum, medio longitudinaliter subsulcato, antice et ad latera 
setis curvatis suberectis, sat conspicuis, vestitum. Elytra rufescentia, pallide squamosa, 
parte apicali nigra vel nigro-maculata, dorso plus minusve inaequali, irregulariter 
punctato, setis erectis brevibus longioribusque intermixtis sat conspicuis, humeris fortiter 
productis. Femora minus fortiter incrassata. tt- Long. 2 '5 — y2 mm. 

A variable species, typical examples of which are easily distinguished. The basal 
two-thirds of the elytra are covered with pale squamosity (except generally on some 
black spots), and this pale portion is often flattened or subdepressed, between the more 
or less evident longitudinal ridges. In well-developed examples there are also traces of 
transverse ridges. The puncturation has a peculiarly irregular appearance. The 
antennae are quite slender, clear red except at the apex, but not very long. The eyes 
are of moderate size and prominent, and there is more or less trace of a dividing ridge 
between the vertex and front of the head. The rostrum of the % is finely longitudinally 
rugulose and dull. The femora are not very strongly clavate. The coxae are all 
equally widely separated, and the basal segment of the abdomen beneath has a dull and 
densely and minutely granulate appearance. 

Hab. Maui, Haleakala (5000 ft.); about two dozen examples taken. 

(70) Proterhinus epichrystis, sp. nov. 

Rufo-niger, elytris rufescentibus, plus minusve nigro-maculatis. Antennae 
graciles, sat elongatae. Oculi fortiter prominentes. Frons capitis a vertice carina 
distincta transversa separata. Pronotum squamis pallidis aureis densius vestitum, 
fortiter 3-impressum, setis suberectis curvatis ad latera conspicuis. Elytra nigro- 
maculata, pallide squamosa, setisque albidis elongatis conspicue vestita, fortiter punctata, 
fere aequaliter convexa, sutura elevatula, basi scutellum juxta utrinque tuberculata, 
humeris fortiter acute productis. ^ $. Long. 27 — 4 mm. 

A rather large species, but very variable in size, ill-developed examples being only 
about one-third or one-fourth the bulk of the larger ones. The head and thorax are 
densely clothed with a very pale golden squamosity, the eyes are strongly prominent 
and usually of conical shape, the antennae elongate and slender with well-marked 
3-jointed club. Thoracic impressions deep and distinct. Elytra red with black 
markings, coarsely punctured, the longitudinal ridges little or not at all evident, tubercu- 
late at the base on either side near the scutellum, the humeral angles strongly produced 



COLEOPTERA 



219 



and acute. Their clothing consists of pale squamosity, which is not very dense, and is 
absent from the dark markings, and numerous elongate white, erect setae, which are 
conspicuous. The femora are not very stout, the lobes of the tarsi are somewhat 
variable in size, but never very largely developed. The rostrum in the % is short, 
generally slightly narrowed to the apex, and the head in both sexes has a very distinct 
transverse ridge (rarely interrupted) between the vertex and the front. All the coxae 
are widely, and about equally, distant, the basal ventral abdominal segment is minutely 
granular, rather than punctured. 

Hab. Maui, Haleakala (5000 ft.) ; about 40 examples were taken. 

(71) Proierhiiuis halcakalac, sp. nov. 

Niger, pedibus antennisque rufescentibus. Caput post oculos constrictum, his 
fortiter prominentibus. Antennae graciliores. Rostrum % fere opacum, crebre longi- 
tudinaliter rugulosum, lateribus parallelis. Pronotum elongatum, lateribus subaequaliter 
rotundatis, distincte 3-impressum, impressione antica lata, duabus posticis rotundatis. 
Elytra elongata, parce pallide squamosa, setisque erectis albidis parcius vestita, fortiter 
punctata, ad basim scutellum juxta utrinque tuberculata, fere aequaliter convexa, humeris 
peracute fortiter productis. Femora minus fortiter clavata, subgracilia, lobis tarsorum 
anticorum minoribus. $. Long. 3'5 mm. 

Only a single $ of this species was collected, and it appears to be most nearly 
related to P. epichrysiis of the Maui species, but more closely still to P. lanaiensis of 
Lanai. In colour it is entirely black, except for the dark reddish legs and the rather 
more brightly coloured basal joints of the antennae. The transverse ridge between the 
vertex and front of the head is very distinct, the eyes are strongly prominent. The 
basal of the three joints of the club of the antennae is not very much more developed 
than the preceding, so that the club is not very distinct. The anterior impression of 
the elongate thorax is large and vague, the two posterior ones small, round, and definite. 
The elytra are long, nearly parallel-sided for more than half their length, with traces of 
the two longitudinal ridges, at least in certain aspects. All the femora are somewhat 
slender, and the lobes of the front tarsi are decidedly small for the size of the insect. 

Hab. Maui, Haleakala. A single ? was taken on April ist, 1894, at an elevation 
of about 5000 ft. 



(72) ProterJiiuHS arhopalus, sp. nov. 

Nigricans vel piceo-niger, tibiis tarsisque obscurius rufescentibus. Oculi magni, 
prominentes. Antennae incrassatae, articulo primo elongato, robuste clavato, secundo 
elongato, subtriangulari, tertio hoc paullo longiore, 4°, 5°, 6° et 7° haud longioribus quam 
F. H. II. 29 



220 FAUNA HAWAIIENSIS 

latioribus, angulis apicalibus posterioribus productis, 9°, 10° et 11° his evidenter gracili- 
oribus. Pronotum 3-impressum, lateribus subfortiter rotundatis. Elytra subsquamosa, 
setisque erectis pallidis vestita, longitudinaliter subcarinulata, sutura subelevata, basi 
scutellum juxta utrinque tuberculata, humeris acute fortiter productis. Femora 
antica fortiter incrassata, lobis tarsorum maximis. ^. Long, vix 4 mm. (Plate IX. 
fig. 21. Z.) 

This very remarkable species cannot be confused with any other, on account of the 
structure of the antennae, the middle joints of which are produced at their hinder apical 
angles, and are actually wider than the terminal ones, which usually form the club. 
The front and vertex of the head are not divided by any evident ridge, the eyes are 
strongly prominent, and the tarsal lobes of the front feet very large. The elytra when 
closely examined are seen to have each a fine ridge about midway between the suture 
and lateral margin. These ridges are a good deal more evident in one of the examples 
than in the other, and the development of the antennae also is decidedly greater in one 
of the two specimens. 

Hab. Maui, Haleakala. Two male examples taken at an elevation of 5000 ft. in 
October, 1896. 

{j-^ Proterhiims epitracJiys, sp. nov. 

Niger, statura majore, antennis pedibusque nigricantibus. Antennarum articulus 
primus brevis et robustus, secundus vix longior quam latior, caeteris elongatis, clava 
elongata distinctissime 3-articulata. Oculi permagni, fortiter prominentes. Pronotum 
squamosum, lateribus setis conspicuis erectis curvatis vestitis, fortissime 3-impressum, 
medium longitudinaliter subsulcatum. Elytra variegato-squamosa, grossissime rugoso- 
punctata, basi scutellum juxta utrinque tuberculata, setis gracilibus erectis conspicue 
vestita, sutura subdepressa, humeris fortiter productis. Femora antica sat incrassata, 
lobis tarsorum parvis. ^. Long. 4 mm. 

A large species, with very large eyes, antennae thick at the base, but becoming 
slender before the club-joints, and unusually small lobes to the second joint of the front 
tarsi. Thorax deeply impressed in the middle in front, and this impression connected by 
a longitudinal groove with a postero-median impression, the two sublateral impressions 
very deep and distinct, and rather large, the hind angles marked by a distinct spot of 
white squamosity. Elytra uneven, submaculately squamose, impressed between the 
basal tubercle and the strongly produced humeral angle on either side, the puncturation 
extremely coarse and rugose, the sutural portion subdepressed. The fine erect setae 
of the elytra and the curved ones on the prothorax unusually conspicuous. Tibiae 
with the setae short and somewhat dense. 

Hab. Maui. A single ^ taken on Haleakala in April, 1894, at an elevation of 
less than 4000 ft. 



COLEOPTERA 221 



(74) Protcrliinus separandus, sp. nov. 

Rufescens, capite, thorace (plus minusve), antennis, femoribusque obscurioribus. 
Capitis vertex a fronte baud divisus. Oculi magni, fortiter prominentes. Antennae 
subgraciles, articulo primo crasso et brevi, clava distincte 3-articulata, baud notabili. 
Pronotum pallide squamosum, antice abrupte constrictum et transversim impressum, 
impressionibus posterioribus minus distinctis, dense rugoso-punctatum. Elytra griseo- 
squamosa, sparsim nigro-maculata, obscurius longitudinaliter carinata, basi extrema 
bituberculata, humeris productis. Femora sat fortiter incrassata, lobis tarsorum anti- 
corum sat magnis. $. Long. 3'2 mm. 

This species is quite distinct from any other species known on Maui, but is very 
closely allied to the Oahuan P. ruficornis. The single J example described above is 
probably somewhat abraded, but there are evident erect pale setae still remaining on the 
elytra. There is no trace of a ridge between the front and vertex of the head, and the 
eyes are very large and strongly prominent. The antennae are not very slender, their 
basal joint is very short and stout, the second slightly longer than wide, the third about 
twice as long as this, the club is distinctly formed by three joints, but is not unusually 
elongate or in any way remarkable. This will easily distinguish the species from the 
Oahuan P. ruficornis, which has the club very strongly developed, its basal joint being 
more than usually elongate, very narrow at the base and much widened at the apex. 
Basal segment of the abdomen beneath rather strongly punctured, evenly at the sides, 
the puncturation becoming obsolete about the middle. Coxae rather less widely 
separated than in most of the species. 

Hab. Maui. A single example taken in the lao valley in March, 1S94 ; and one 
from the mountains of Molokai in 1893 may perhaps belong to the same species. 



(75) Protcrhimis mactilatns, sp. nov. 

Rufescens, femoribus nigricantibus, elytris nigro-maculatis. Antennae graciles, 
rufae, clava vix plus quam 2-articulata. Oculi minores. Pronotum squamosum, 
medium sat latum, antice fortiter abrupte constrictum, fortiter 3-impressum, lateribus 
setis curvatis conspicue marginatis. Elytra maculatim squamosa, setisque erectis 
pallidis parcius vestita, linea obliqua pallide squamosa posthumerali maculisque 
rotundis compluribus postice ornata, basi emarginata, humeris distinctis, vel productis. 
Femora omnia fortiter incrassata. Abdominis segmentum primum ventrale fortiter 
sed minus dense punctatum. $%. Long. 2*6 — 3 mm. 

I have seen only two examples of this species, and the ? is so different to the 
$ in form, owing to the extreme elongation of its elytra, as at first sight to appear 

29 — 2 



222 FAUNA HAWAIIENSIS 

to be a quite distinct species. No doubt this will prove to be a variable character, 
and probably common to either sex, as in many other species of the genus. The 
species is not closely allied to any other of those taken on Maui, but rather re- 
sembles P. vestitns of Oahu and its allies on Hawaii. It is remarkable for its red 
colour, entirely red antennae, the club of which is hardly more than 2-jointed, since 
(especially in the $) the ninth joint is but little different to the preceding, and the 
strongly clavate black femora in either sex. The elytra have a distinct oblique post- 
humeral elongate spot of pale squamosity, and several small roundish ones behind 
these, they are subcompressed at the sides but can hardly be said to bear longi- 
tudinal ridges. Their puncturation is close and coarse, and their form in the 
$ is very long and narrow. 

Hab. Maui, lao valley, March 1S94. 

(76) Proterhinus osculans, sp. nov. 

Parum latus, piceus, rufo-piceus vel rufescens, pedibus plerumque rufis, antennis, 
articulo basali saepe excepto, nigris. Oculi minimi sed prominentes. Antennae 
graciles, clava vix plus quam 2-articulata, articulo 8° et 9° inter se simillimis (hujus 
apice paullo latiore). Pronotum parce squamosum, antice transversim impressum, 
depressionibus posticis vel absentibus vel fere obsoletis. Elytra parum squamosa, 
post media saepe squamositate pallida binotata, setis erectis brevioribus vestita, 
densissime grosse rugoso-punctata, humeris acutis et productis. Femora omnia con- 
spicue gracilia. ^%. Long. 2 — 2 '5 mm. (Plate X. fig. 6.) 

This is very unlike any of the other Maui species, and it does not indeed appear 
to be very closely allied to any other species of the genus. It is chiefly remarkable for 
the unusually slender femora, the hardly more than two-jointed antennal club (the tenth 
joint being much wider than the ninth, and this not very different to the preceding), and 
the somewhat long rostrum of the %, which is narrowed towards the base, the surface 
being entirely longitudinally rugose. The eyes are quite small, but prominent, the 
squamous covering of the insect is sparse, revealing the dense puncturation, the surface 
itself being sometimes even shining, and set with shortish erect setae. The elytra are 
without longitudinal ridges, their humeral angles acute and somewhat produced. 
Usually the entire insect is covered when captured with an exudation, which conceals 
all the structure, and is difficult to remove. 

Hab. Maui, Haleakala (5000 ft.). About 20 examples taken. 

{"]"]) Proterhimis lepiorhynchns, sp. nov. 

Parum latus, rufescens, pronoto cum capite obscuriore, irregulariter squamosus. 
Oculi parvi, prominuli. Antennae basi excepta nigrae, graciles, clava vix plus quam 



COLEOPTERA 223 

2-articulata. Pronotum latum, antice abrupte constrictum, densissime punctatum, im- 
pressionibus posterioribus absentibus, linea longitudinal! squamositatis densae utrinque 
notatum, caetera parte parce squamosa, setis curvatis sat distinctis lateraliter vestitum. 
Femora parum fortiter clavata. Elytra irregulariter albido-squamosa, baud inaequalia, 
setis perpaucis erectis albidis inconspicue vestita, basim versus sat fortiter angustata, 
humeris acutis. Rostrum % gracile, fortius elongatum, basim versus evidenter angus- 
tatum. %. Long. 2 '5 mm. 

This species is perhaps the most closely allied to P. oscidans, having the 
rostrum of the % of very similar form, elongate and widening towards the ape.x, and also 
very similar antennae. The general aspect of the two is very different, the abundant 
short erect setae, which form the chief portion of the clothing in P. oscidans, being 
almost absent in the present species, and there are numerous other distinctions. The 
basal abdominal segment beneath closely and strongly punctured at the sides. 

Hab. Maui, lao valley. A single % example taken. 



(78) ProterJdiius vdrabilis, sp. nov. 

Nigricans, tibiis, lobisque tarsorum, rufescentibus. Caput post oculos subcon- 
strictum, oculis fortiter prominentibus. Antennae elongatae, vix graciles, articulo 
primo fortiter clavato et elongato, secundo robusto, longiore quam latiore, tertio 
quartoque elongatis, lateribus fere parallelis, clavae 3-articulatae articulo basali 
longiore et evidenter latiore quam articulus octavus. Pronotum profunde 3-impressum, 
antice fortiter angustatum, pallide (latera versus densissime) squamosum. Femora 
antice fortissime incrassata, lobis tarsorum latis et permagnis. Elytra maculatim 
squamosa, setisque erectis albidis parcius vestita, rugoso-punctata, subaequaliter con- 
vexa, basi scutellum juxta utrinque tuberculata, humeris productis. ^. Long, vix 
4 mm. (Plate IX. fig. 22.) 

A somewhat large species, with the lobes of the front tarsi very strongly developed. 
The antennae are also remarkable, the basal joints being elongate but stouter than usual, 
the first long and strongly clavate, the next two or three nearly parallel sided, whereas 
the two or three preceding the club are short and of triangular shape. The three 
terminal joints form a distinct club, the 9th joint being considerably wider at the apex 
than the preceding. The head is constricted behind the eyes, an obscure ridge being 
formed between the front and the vertex. The thorax is wide across the middle, but 
very strongly and abruptly narrowed in front, the two posterior impressions are very 
deep, roundish, and small, the front one wide and vague, but deepened at the middle. 
The pale squamosity towards the sides is very dense. The elytra have no really 
definite longitudinal ridges, but are nearly evenly convex, tuberculate on either side 



224 FAUNA HAWAIIENSIS 

of the scutellar region at the extreme base, the humeral angles being produced, but 
hardly sharp. The front femora are extremely strongly incrassata, and the white 
erect setae on all the tibiae are dense and conspicuous. 

Hab. Maui. A single ^ of this fine insect was taken on Haleakala at an 
elevation of 5000 ft. in May 1896. 



(79) Proterhimis calliphyas, sp. nov. 

Nigrescens, tibiis, tarsisque, necnon saepe parte elytrorum et fronte, et rarius 
antennis, rufescentibus. Oculi parvi sed prominuli. Antennae graciles, articulis 
duobus basalibus sat crassis, clava 3-articulata. Pronotum perinaequale, a latere 
visum postice elevatum, medium longitudinaliter plus minusve distincte sulcatum, 
latera versus densissime pallide squamosum, lateribus fortissime rotundatis. Elytra 
maculatim squamosa, setisque erectis albis parce vestita, paullo post basim vage, sed 
evidenter, depressiusculum, humeris productis. Femora omnia sat fortiter incrassata. 
Abdominis segmentum primum ventrale dense minute granulato-asperulum, vix 
punctatum. ^ $. 2'2 — 3 mm. 

Although by no means a large insect this species is rather robust in form, especially 
in the Z sex. It is very distinct from most of the other species found on Maui which 
have the humeral angles produced, by the form of the prothorax, which, seen from the 
side, has the posterior half much elevated above the anterior, by the slender antennae 
with the two basal joints stout, and by the rather obscure depression of the elytra a 
little behind the basal margin. When in fine condition the elytra are prettily maculate, 
and the prothorax towards the sides is very densely squamose, the middle portion much 
less so. The dorsal and lateral portions of the thorax are very clearly marked off, the 
two forming a sharp edge where they meet, and this edge is set with curved setae. 

Hab. Maui, Haleakala (4000 — 5000 ft.) ; generally attached to Alyxia. Allied to 
the larger P. mirabilis but very distinct. 



(80) Proterhimis persimilis, sp. nov. 

Major, fusco-niger, elytris pedibusque fusco-squamosis, P. dispart cognatissi- 
mus. Antennae elongatae et gracillimae, articulo secundo brevissimo, tertio fortiter 
elongato et gracillimo, clava 3-articulata, perelongata. Oculi fortissime prominentes. 
Pronotum antice angustatum et impressum, impressionibus duabus posterioribus sat 
distinctis, rotundatis. Elytra squamosa, setisque albidis erectis parce vestita, basi 
scutellum juxta utrinque tuberculata, dorso sutura excepta longitudinaliter impresso, 
humeris leviter productis. Femora (praesertim $) insignissime incrassata, lobis tarsorum 
anticorum latis. J $. Long. 3-5 — 5 mm. (Plate X. fig. 7.) 



COLEOPTERA 225 

Closely allied to P. dispar. Sharp, of which species it might be regarded as a 
highly-developed local race. The t is remarkable for the development of the rostrum, 
which is less short than usual, and the extreme thickness of the femora. The eyes are 
large and very prominent, the antennae long and very slender, the lobes of the front 
tarsi strongly developed, being unusually wide, and the joint strongly transverse. The 
tibiae have a clothing of somewhat fine hairs. These are also mainly the characters of 
P. dispar, but that species may be distinguished by the less elongate and slender third 
joint of the antennae, and indeed generally by the following joints also being shorter, 
and it never attains the size of the larger e.xamples of P. persiniilis. 

Hab. Maui, lao valley ; attached to JVikstroemia. 



(81) Proterhimis nivicola, sp. nov. 

Totus niger, angustus, parce squamosus. Antennae nigrae, graciles, breviores, 
clava vix 3-articulata. Oculi minores, minus prominentes. Pronotum subelongatum, 
minus distincte 3-impressum, parum squamosum, sed ad angulos posteriores macula 
albido-squamosa signatum. Elytra angusta, fortiter punctata, parcius obscure sub- 
maculatim squamosa, setis erectis brevissimis et perinconspicuis, basi utrinque scutellum 
juxta distincte tuberculata, angulis humeralibus fere rectis. Femora omnia minus fortiter 
clavata, lobis tarsorum anticorum minutis. %. Long, vix 3 mm. 

Only a single example of the % of this species having been taken nothing is known 
as to its variability. It is chiefly distinguished by its general black colour and narrow 
elongate form, but possesses no very remarkable characters. The ninth joint of the 
antennae is but little different to the preceding, so that the club is apparently 2-jointed. 
To the characters given above it may be added that the rostrum is rather short, shining, 
and with the longitudinal grooves somewhat strong, and that the basal ventral segment 
of the hind-body is strongly and closely punctate over its whole surface. The species 
is of interest as being the only one found far above the limit of the true forest. 

Hab. Maui, Haleakala (9000 ft.) ; taken not many weeks after the disappearance 
of a heavy fall of snow. 

(82) Proterhinus punctipennis Sharp. 

Proterhimis pimctipcnuis Sharp, Tr. Ent. Soc. London, 1881, p. 530. 

The examples which served for the original description of this species were large 
and well-developed specimens. It is, however, very variable, the development of the 
antennae being much less strong in some examples than in others. The puncturation 
of the elytra, which is in reality very coarse and close, appears much less striking in 
individuals in which the squamous covering is more abundant, and frequently the insect 



2 26 FAUNA HAWAIIENSIS 

is entirely or to a large extent covered with an exudation, which altogether conceals or 
very much changes the appearance of the sculpture. Generally the insect is red, with 
dark lateral markings on the elytra, but some examples are nearly entirely blackish or 
piceous, with black antennae. Small individuals with slender antennae certainly closely 
approach the Oahuan P. oscillans, but I cannot consider the two species identical. 
Length 2 — 3 mm. 

Hab. Maui, Haleakala (4000 — 5000 ft.) ; upwards of seventy examples of this 
species have been examined. 



(83) Proterhimis epiinelas, sp. nov. 

Nigricans, tibils tarsisque et rarius femoribus rufescentibus, maculatim pallide 
squamosus. Oculi parvi. Antennae graciles, articulo secundo elongato, haud minus 
longo, vel etiam (praesertim in %) nonnunquam longiore quam tertius. Pronotum 
antice depressum, lateribus fortiter rotundatis, setisque curvatis marginatis, utrinque 
plaga sublaterali densissime albido-squamosa. Elytra maculatim squamosa, parcissime 
albido-setulosa, punctata, humeris parum distinctis. ^ $. Long, vix 2 — 2"5 mm. 

A small species, remarkable for its dark colour, the tibiae and tarsi alone being as a 
rule of a red or pitchy colour, and for the elongation of the second joint of the antennae, 
which is subequal to the third in length, or not infrequently decidedly the longer of the 
two. The prothorax is very densely squamose towards the sides, less so towards the 
middle. The elytra have no longitudinal ridges, but are more or less perceptibly, 
though vaguely, impressed transversely a little behind the base, and therefore have a 
slight appearance of elevation at the extreme base in the region of the scutellum. The 
humeral angles are obscure. 

Hab. Maui, Haleakala (5000 ft.) ; an extensive series taken. This insect varies 
a good deal in size, but is never a large and often a minute species. 



(84) Proterhinus 7'ujicollis, sp. nov. 

Rufescens, elytris nonnunquam plus minusve nigro-marginatis, supra rarius nigro- 
infuscatis, parum dense squamosus. Antennae graciles, sat elongatae, ruficolores, 
articulis apicalibus nonnunquam nigricantibus. Pronotum plerumque (^ saltem) sat 
latum, medium parce, latera versus densius, aureo-squamosum, vix evidenter punctatum, 
antice transversim impressum. Elytra irregulariter nee dense squamosa, squamis 
saepe maculas minus distinctas rotundatas formantibus, dense et sat distincte punctata, 
setis erectis evidenter vestita, angulis humeralibus haud acutis. Femora postica minus 
fortiter clavata, lobis tarsorum anticorum parvis. ^ %. Long. r8 — 2*2 mm. 



COLEOPTERA 227 

One of the smallest species of the genus generally remarkable for being entirely, 
or for the most part, of a rufescent colour. The series of over a score of individuals 
exhibit not a little variation, and the tendency in some of them to become very narrow 
and elongate in form is remarkable, although a similar tendency is observed in several 
other species. Certain individuals approach very closely to some varieties of P. deceptor, 
but I cannot consider the two species identical. 

Hab. Maui, lao valley. A series of examples taken in March 1894. All were 
obtained at the same time and place. A single example taken on Molokai perhaps 
belongs to this species. 



D. Species insulae Lanai propriae. 

(85) Proterhimis insignis Sharp. 

Proterhimis insignis Sharp. Tr. Dublin Soc. iii. 1SS5, p. 173. PI. V. fig. 44 
and 45. 

This species is distinct at a glance by its large size (although it is of rather narrow 
form), and the very strongly elongate slender antennae of the ^, with their exceeding 
long but stout basal joint. In the % this joint also is extremely elongated, but less so 
than that of the ^. Length 3 '5 — 4*5 mm. 

Hab. Lanai, mountains (2000 — 3000 ft.) ; rare, only about a dozen examples 
were taken. 

(86) Proterhimis loiigicornis Sharp. 

Proierhimis longicornis Sharp, Tr. Dublin Soc. iii. 1885, p. 173. 

Z readily distinguished by the unusually long antennae, and especially of the 8th 
joint, which is longer than the following or basal joint of the club. The antennae of 
the % are much shorter, but the unusual length of the 8th joint is noticeable also in this 
sex. The species appears to be most closely allied to P. kamptarthrus of Oahu, but 
the two are very distinct. Length 2-5 — 375 mm. 

Hab. Lanai, mountains ; rare, less than a score of specimens taken. 



(87) Proterhinus lanaiensis, sp, nov. 

Statura majore, suboblongus, rufo-niger, elytris plus minusve variegatis. Antennae 

gracillimae, rufae, apicibus nonnunquam nigris, clava distincte 3-articulata. Oculi 

magni, coniformes, fortiter prominentes. Caput post oculos fortiter constrictum. Pro- 

notum dense aureo-squamosum, distincte 3-impressum, setis curvatis conspicuis. Elytra 

F. H. II. 30 



2 28 FAUNA HAWAIIENSIS 

saepe nitida, grosse punctata, plus minus irregulariter vel maculatim squamosa, setis 
erectis gracillimis perconspicuis crebre vestita, basi scutellum juxta utrinque tuberculata, 
humeris fortiter acute productis, lateribus saepe fere parallelis. ^ %. Long. 3 — 4 mm. 

When in fresh condition this species is readily distinguished by the distribution of 
its squamous covering, and the very conspicuous and extremely fine erect setae, with 
vi^hich the elytra are clothed, combined with the characters afforded by its shape, 
the elytra being generally of more than usually parallel form, the distinct ridge formed 
between the vertex and front of the head, the conical prominent eyes, the coarse 
puncturation, and sometimes the shining surface of the elytra. I'here are on these, 
in some examples, traces of longitudinal ridges, but they are generally indistinct. 
Most closely allied, I think, to P. haleakalae, but certainly distinct. The rostrum 
in the % of that species is quite dull, while in P. lanaiensis it is, at least to a con- 
siderable extent, shining. 

Hab. Lanai, 2000 — 3000 ft. Not common. 



(88) Proterhimis ineptus Sharp. 

Proterhinus ineptus Sharp, Tr. Dublin Soc. iii. 1885, p. 171. 
Pi'oterhinus integer Sharp, /. c. p. 172, van 

An extraordinarily variable species, of which P. integer Sharp is no doubt a narrow 
and highly-developed variety. The prothorax is very variable in length and roundness, 
and the size of eyes, tarsal lobes, length of the antennae are all variable characters. 
Many of the varieties of this species occur quite constantly, and it is quite probable 
that some of these are in the process of becoming distinct species, but between such 
forms as P. ineptus and P. integer intermediate varieties certainly occur. The larger 
individuals attain a length of about 4 mm., the smallest I have seen is less than 2 mm. 
in length, but the latter is quite exceptional. 

Hab. Lanai, throughout the forest, from 2000 — 3000 ft. Many examples (from 
one to two hundred) examined. 



(89) Proterhinus detritus Sharp. 

Proterhinus detritus Sharp, Tr. Dublin Soc. in. 1885, p. 172. 

(Plate X. fig. I.) 

Of this species I have seen very few examples, the type (j") in Dr Sharp's 
collection, and four or five examples taken by myself The former is in very abraded 
condition. The examples in the British Museum, which are placed under this name, as 



COLEOPTERA 



229 



determined by Mr Blackburn, are of a quite different species, and not allied to the true 
P. detrihis^. The species is remarkable for its red or pitchy red colour, very elongate 
elytra, the abrupt narrowing of the prothorax in front, whereby a more or less distinct 
angle is formed on each side, the very short second joint to the antennae, and the 
deep sulci of the rostrum of the $. The squamosity has not a maculate arrangement, 
and is very scanty on the elytra, less so on the thorax, and dense on the face, and is 
of a golden colour in fresh examples, in which the erect flavescent setae on the elytra 
are quite conspicuous. 

Hab. Lanai, 2000 ft. and upwards ; a very scarce species. 

(90) Proterhiims epifretus, sp. nov. 

Elongatus, ferrugineus, setis erectis sat conspicue vestitus. Antennae ferrugineae, 
articulo secundo minus brevi, plus minusve elongato. Pronotum minus latum, saepe 
subelongatum, setis appressis, aliisque curvatis et suberectis vestitum, 3-impressum. 
Elytra elongata, grossissime punctata, setis elongatis, erectis, et gracillimis vestita, 
dorso parum vel baud convexo, angulis humeralibus productis et acutis. $ $. Long. 
275 — vix 4 mm. (Plate X. fig. 2.) 

On account of the character of its clothing, form, and colour, this species could 
only be confused with P. detritus Shp. of those species found in Lanai, but it is 
distinct from that species by the form of the prothorax, less short second antennal joint, 
and finer and longer erect setae of the elytra, as well as by the smoother and less 
strongly grooved rostrum of the $, &c. P. epitretus on Lanai represents the P. longiilus 
of Oahu, from which it differs in the more extremely coarse puncturation of the 
elytra, and the longer and finer erect setae, and the former character will also readily 
separate it from P. ferrugineus of Hawaii. The species varies greatly in size, and 
considerably in the length of the antennae. Mirmte examples are only one-half or 
one-third the size of the well-developed individuals. 

Hab. Lanai, mountains (2000 — ^3000 ft.) ; attached to the stems of the fronds 
of the tree-ferns. 

(91) Proterhimis hrez'i/orniis, sp. nov. 

Sat latus, parum convexus, niger, tibiis tarsisque et nonnunquam antennis etiam 
et femoribus rufis, elytris maculatim squamosis. $ antennae fortius elongatae et 
graciles, clava elongata 3-articulata. Pronotum latissimum, antice sat abrupte angus- 

' Since writing the above I find that three insects were sent by Mr Blackburn to Dr Sharp under the 
number 449, and are marked A, B, and C. One of these is the true F. detritus, the other two are examples 
of what I consider to be the commonest form of P. angularis, the type specimens of that species being an 
unusual variety. These two examples are very different to one another, one being very narrow in form, a 
variety which I have taken on several islands in company with the more usual one of P. angularis. The 
British Museum examples under P. detritus are also vars. of P. angularis. 

30—2 



2 30 FAUNA HAWAIIENSIS 

tatum, ibique impressum, ad medium minus dense, latera versus densissime squamosum. 
Elytra post basim depressiuscula, maculatim aureo-squamosa, setis perpaucis erectis 
parum conspicue vestita, asperula, minus distincte punctata, angulis humeralibus ob- 
scurioribus. ^ %. Long, circa 2-5 mm. 

This little species is quite distinct, and rather reminds one of P. dispar. It 
appears to be very constant in its general appearance, and may be at once recognized 
from its nearest allies by its short and wide form, very wide thorax, long, slender 
antennae, dark colour, and the maculate arrangement of the squamosity of its elytra. 
The % appears to be generally more slender, and of less distinctive appearance than is 
the <?, but superficially the sexes much resemble each other. 

Hab. Lanai, 2000 ft. Not a common species, about a dozen examples were 
taken in the winter of 1894. 



(92) Protc7-hi7ms analcis, sp. nov. 

Angustulus, sordide niger, vel piceus, squamis aureis irregulariter minus dense 
vestitus. Antennae breviores, graciles, nigrae, basi rufa, clava parum abrupte 3-articu- 
lata. Pronotum minus latum, saepe plus minusve evidenter 3-impressum vel antice 
tantum impressum, lateribus minus fortiter rotundatis. Elytra subirregulariter squamosa, 
setisque brevissimis erectis vestita, basi scutellum juxta utrinque plus minusve tuber- 
culata, angulis humeralibus plerumque fere rectis. ^ ?. Long. r5 — 275 mm. 

Quite one of the most obscure species of the genus and generally of minute size, 
and chiefly distinguished by the fact that it possesses, at least so far as I can see, 
no notable peculiarity of structure. The antennae are shorter than in most species, 
generally black, with one or more of the basal joints red, sometimes entirely black. 
The prothorax frequently shows three impressions, and is not at all wide. The elytra 
are usually subtuberculate at the extreme base on each side near the scutellum, and the 
squamous clothing is irregular and not dense, the erect setae short. These points and 
the obscure colour furnish the best characters, but the insect in reality is quite unlike 
any other species found on Lanai, nor will it agree with any of those taken on other of 
the islands. 

Hab. Lanai, mountains ; some dozens of this species have been examined. 



(93) Protcrhimis epichlorus, sp. nov. 

^ rufescens, rarius rufo-niger, pallide, vix vel baud maculatim, squamosus. Anten- 
nae fortiter elongatae, articulo secundo elongato, quam tertius vix vel haud minus longo. 
Pronotum latum, subaequaliter sat dense aureo-squamosum, lateribus fortiter rotundatis. 



COLEOPTERA 231 

Elytra pallida squamosa, necnon setis brevibus erectis vestita, basi scutellum juxta 
subelevata, angulis humeralibus obscuris. Femora sat incrassata, lobis tarsorum anti- 
corum haud minutls. Z ¥• Long, vix 2 — 2 '5 mm. 

Amongst the species with indistinct humeral angles, this is chiefly remarkable 
for the following characters, at least in the Z sex. The antennae are strongly developed, 
the basal joint robust and somewhat long, the second elongate, and when seen in some 
aspects as long as, or hardly less long than, the following. The club is of three joints, 
but not very distinct, as the intermediate joints are generally less slender than in many 
species. Thorax very strongly rounded at the sides. Lobes of front tarsi fairly well 
developed. Elytra with the erect setae short, and at the base in the region of the 
scutellum a little elevated. The colour of the insect is generally rufescent, sometimes 
with some darker markings on the elytra, but occasional specimens are much obscured 
with blackish colouring. 

The $ which I assign to these males resembles it in general appearance but 
appears to vary in the length of the antennal joints. 

The elongation of the second antennal joint in this species calls to mind P. cpimelas 
of Maui, but the superficial appearance of the two is entirely different. 

Hab. Lanai (2000 — 3000 ft.) ; not rare. 



E. Species insulae Molokai propriae. 

(94) Protcrhiims molokaiensis, sp. nov. 

Major, elongatulus, sordide niger, elytris magna ex parte rufescentibus. Antennae 
subvalidae, articulo primo fortius elongate, sed robustissimo, secundo brevissimo, clava 
evidenter 3-articulata, sed parum conspicua. Oculi magni fortiter prominentes. Pro- 
notum subelongatum, dense punctatum, antice subabrupte et fortiter constrictum, 
3-impressum, pallide squamosum, setis curvatis suberectis anterius marginatum. Elytra 
et longitudinaliter et transversim subinaequalia, dorso rufescente et pallide squamoso, 
subdeplanato, setis erectis pallidis conspersim sed conspicue vestita, remotius punctata, 
basi utrinque scutellum juxta fortiter tuberculata, humeris acutis, fortiter productis. 
Femora fortiter incrassata, lobis tarsorum anticorum parum elongatis, sed robustis. 
t. Long. 3-5 mm. 

Allied to P. validiis of Maui, this species is quite distinct. 

Hab. Molokai. A single t taken in the mountains at an elevation of 3000 ft., in 
June 1896. 



232 FAUNA HAWAIIENSIS 

(95) Proterhimis convexiusculus, sp. nov. 

Nigricans (nonnunquam rufescens), elytris plerumque plus minus rufescentibus, sat 
convexis, pallide aureo-squamosus. Oculi magni, prominentes. Pronotum pallide 
squamosum, forma variabili, impressionibus sat profundis (nonnunquam lutoso-repletis), 
medium saepe longitudinaliter subsulcatum. Elytra sat convexa, pallide aureo- vel 
griseo-squamosa, squamis baud remotis minus distincte punctata, angulis ipsis humerali- 
bus fortiter acute productis, sutura elevatula, setis erectis parce conspersis. Abdominis 
segmentum primum ventrale subtiliter granulatim asperulum. ^ %. Long. 2 '6 — 3 "5 
mm. 

A variable species, individuals differing a good deal in size, colour, and distribution 
of the squamosity, as well as in the length and thickness of the antennae, &c. I have 
had some doubt as to whether it can be absolutely separated from that excessively 
variable species P. integer of Lanai, but have thought it better to keep the two apart. 
One almost entirely black example of this species has been taken, even the antennae 
being entirely of that colour, and only the tarsal lobes are piceous, and this individual 
has the elytra covered with grey squamosity. Usually the squamous covering is 
co-extensive with the red colour of the wing-cases, which typically are red at the base 
and there squamose, the apical dark parts being nearly free from squamosity, the extent 
of the red colour and squamous covering varying together. In P. convexiusculns it 
may be observed there is little or no trace of a transverse ridge between the front and 
vertex of the head. 

Hab. Molokai, mountains, generally at an elevation of about 4000 ft. Numerous 
examples were taken, but many of them are in indifferent condition. 

(96) Proterhimis echinoides, sp. nov. 

Nigricans, elytris rufis, nigro-signatis, tibiis tarsisque rufescentibus, antennis fere 
nigris, pallide squamosus. Antennae breviores, subcrassae, clava 2-articulata. Oculi 
magni. Pronotum minus profunde 3-impressum, medium dense, latera versus minus 
dense, squamis pallidis vestitum. Elytra sat fortiter et fere aequaliter convexa, squamis 
pallidis necnon setis multis elongatis erectisque perconspicue vestita, basi scutellum 
juxta utrinque tuberculata, angulis humeralibus fortiter productis. Femora omnia cum 
tibiis setis elongatis pallidis conspicue vestita, tarsorum anticorum lobis sat minutis. ^. 
Long. 2 '5 — 27 mm. 

A small species, but very easily recognized by the unusual development of the 
erect setae on the elytra and legs, the short stout antennae, with the club hardly more 
than 2-jointed, large eyes, small tarsal lobes, and nearly evenly and somewhat strongly 
convex elytra. One example has the antennae rather more elongate than the others. 

Hab. Molokai, mountains (3000 ft.) ; three males taken in June 1896. 



COLEOPTERA 233 

(97) Proterhimis le^icothorax, sp. nov. 

Nigro- vel rufo-piceus, pedibus antennarumque basi plus minus rufescentibus. 
Oculi sat magni, prominuH. Pronotum fortiter 3-impressum, setis appressis squamisque 
albidis dense vestitum (impressionibus 2 lateralibus densissime squamis albicantibus 
vestitis), medium postice subsulcatum. Elytra grossissime dense rugoso-punctata, 
maculis duabus albidis squamosis postice signata, ex majore parte squamis carentia, sed 
setulis brevibus erectis albidis sat evidenter vestita, sutura distincte elevatula, cariniformi, 
angulis humeralibus fortiter productis. Femora postica parum fortiter incrassata. ^ $. 
Long. 2 — 2'25 mm. 

One of the smallest species of those which have the humeral angles of the elytra 
strongly produced forwards. It is chieHy remarkable for the gross sculpture of the 
elytra which are nearly free from squamous covering, and especially for the peculiarly 
dense whitish clothing of the thorax. I see no trace of a transverse ridge between the 
front and vertex of the head in the t, and only the faintest trace thereof in the %. The 
species is allied to P. innotabilis. 

Hab. Molokai. Three examples (one of which has lost most of the squamous 
covering) were taken at an elevation of about 4000 ft. 

(98) Proterliiniis angiistior, sp. nov. 

Subgracilis, nigricans, elytris plus minusve (praecipue basim versus) rufescentibus. 
Antennae graciliores, minus fortiter elongatae, articulis basalibus plerumque rufescenti- 
bus, clava 3-articulata. Pronotum baud latissimum, % saepe angustulum, baud dense 
squamosum, subfortiter 3-impressum. Elytra angustiora, nigricantia, partibus pallidiori- 
bus (sc. rufescentibus) squamas pallidas ferentibus, setis erectis, elongatis, parce 
conspersis, sat evidenter vestita, fere aequaliter convexa, basi scutellum juxta utrinque 
plus minus tuberculata, sutura levissime elevata, humeris fortiter productis. Oculi 
parum fortiter prominentes. Femora omnia fortiter incrassata, lobis tarsorum anticorum 
minoribus. t ?• Long. 2-4 — vix 3 mm. 

The affinities of this species are evidently with P. htimeralis of Maui, which 
in many respects it greatly resembles. All the femora are quite strongly incrassate, 
very much as in that species, but the antennae, especially those of the t, are much less 
developed, as also are the eyes and the lobes of the front tarsi. The chief variation 
exhibited by the seven examples taken lies in the shape of the prothorax, which is a 
eood deal more dobose in some than in others. No doubt in a long series the tendency 
would be to a greater elongation generally in the females, as I have observed to be the 
case with other species. The sides of the rostrum of the ? are slightly convergent 
from base to apex. 

Hab. Molokai, mountains (3000 ft.). 



234 FAUNA HAWAIIENSIS 

(99) Proterhinus heterotarsus, sp. nov. 

Nigricans, elytris rufis plus minusve nigro- vel fusco-notatis, tibiis, tarsis, antennis- 
que plerumque rufis, his apicem versus nigricantibus. Antennae graciles, clava distincte 
3-articulata. Pronotum aequaliter minus dense squamosum, minus fortiter 3-impressum, 
globosum, lateribus fortiter rotundatis. Elytra griseo-squamosa, setisque erectis pallidis 
conspersim vestita, basi emarginata, humeris productis et acutis. Femora sat incrassata ; 
tarsorum anticorum articulus secundus brevissimus, fortiter transversus, lobis brevibus. 
Species praecedenti simillima. ^ %. Long. 2 — 27 mm. 

Apparently closely allied to the preceding species (/". angnsiior), but generally of 
less narrow form, and with the thorax more globose, the antennae rather longer and 
more slender, and the elytra more nearly covered with squamosity, the dark markings 
being reduced. The variation of both species makes it difficult to distinguish the two 
in description, but the tarsi appear to be constantly different in form, the lobes of the 
second joint of the front pair being in this species extremely short, and the whole joint 
very small and strongly transverse, while the rostrum of the $ is nearly parallel-sided, 
not gradually but slightly narrowed to the apex. The extent to which the humeral 
angles of the elytra are produced is decidedly variable. 

Hab. Molokai, mountains (3000 ft.) ; only six or seven examples taken. 

(100) Proterhinus erythrodes, sp. nov. 

Minor, rufescens, totus fere aequaliter pallido-squamosus, elytris nonnunquam 
fusco- vel nigro-notatis. Antennae graciles, elongatae, clava distincte 3-articulata, 
rubricolores, nonnunquam articulis apicalibus nigricantibus. Oculi minores, parum 
prominentes. Pronotum globosum, parum distincte impressum, aequaliter pallide 
squamosum, lateribus fortiter rotundatis. Elytra pallide squamosa, setisque nonnullis 
pallidis erectis parum conspicue vestita, aequaliter convexa, basi leviter emarginata, 
humeris distinctis, sed vix acutis vel productis. Femora omnia cum tibiis tarsisque 
rufescentia (rarius obscurata), lobis tarsorum anticorum minutis. ^ %. Long, i '6 — 
2'5 mm. 

A small and obscure species, without any striking characters. The antennae are 
rather long and quite slender, with a distinctly 3-jointed apical club ; they are often 
entirely of a clear red colour, but the apical joints are sometimes more or less dark. 
The colour of the insect is red, that of the thorax being sometimes more or less 
obscured, as also is the head, and the elytra often have some dark markings. The 
squamous covering is evenly distributed over the greater part of the surface. The 
elytra at the base are lightly emarginate, but the humeral angles are hardly produced, or 
distinctly acute. The eyes are small and the lobes of the front tarsi minute. 

Hab. Molokai, mountains (3000 ft.) ; only nine examples of this species were 
taken. 



COLEOPTERA 235 



(loi) Proter-himis ombrophiliis, sp. nov. 

Totus niger, haud latus, squamis pallidis inaequaliter vestitus. Rostrum $ totum 
opacum, subtiliter longitudinaliter rugulosum, basim versus sensim angustatum. An- 
tennae nigrae, breviores, clava parum distincte 3-articulata (sc. articulo 8° et 9° inter se 
haud multo diversis). Oculi minores, sed subprominuli. Pronotum haud dense 
squamosum, rugoso-punctatum, obscurius 3-impressum, lateribus minus fortiter rotun- 
datis, et macula albida squamosa densiore signatis. Elytra haud inaequalia, basim 
versus haud distincte angustata, obscure punctata et ubique asperula, irregulariter 
(submaculatim) albido-squamosa, setis erectis brevibus et subconspicuis, basi leviter 
emarginata, humeris distinctis, sed minus fortiter prominulis. Femora haud fortiter 
incrassata, tarsis omnibus nigris. $. Long. 2 mm. vix superans. 

This species is closely allied to the unique P. nivicola of Maui, with which it closely 
agrees in colour, clothing, form of antennae, and rostrum, as well as in the somewhat 
peculiar sculpture. It differs from that species at once in its much less elongate elytra, 
which do not exhibit the same tendency to inequality of the surface. P. 07nbropliilus has 
the elytra of only moderate length. 

Hab. Molokai. A single example taken in the highest forest of the island, in 
May 1893. 



(102) Protcrliinns pte7'idis, sp. nov. 

Elongatus, angustissimus, rufescens, elytris saepe plus minus fusco- vel nigro- 
maculatis. Antennae brevissimae, incrassatae, articulo primo robusto, triangulari (? 
etiam fortiter elongato), caeteris brevissimis, clava brevi tantum 2-articulata. Pronotum 
squamosum, saepe plus minus evidenter 3-impressum, impressione antica majore, lateri- 
bus minus fortiter rotundatis. Elytra perelongata pulchre maculatim squamosa, setisque 
albidis vestita, humeris haud productis. Pedes sat robusti, lobis tarsorum minutioribus. 
Z ?. Long. 175 — 2*5 mm. (Plate IX. fig. 24. Z-) 

Although varying a good deal in the development of some of the characters, this 
small species by its elongate and very narrow form, its maculate elytra, and extra- 
ordinarily short and thick antennae, is one of the most remarkable and distinct species 
of the genus. It is I think allied to P. navita, which makes some approach to it in the 
structure of the two basal joints of the antennae. 

Hab. Molokai (3000 ft.) ; in the leaf-stalk of a species o{ Ptej-is. 

F. H. II. 31 



2 36 FAUNA HAWAIIENSIS 



F. Species insulae Hawaii propriae. 
(103) Proterhimis kawaiiensis, sp. nov. 

Nigro- vel rufo-piceus, statura majore. Antennae graciles, elongatae, articulo 
tertio cum sequentibus sat fortiter elongate, clava distinctissime 3-articulata, articulo 
9°, quam 8"^ multo longiore. Oculi permagni, fortiter prominentes. Pronotum profunde 
3-impressum (impressione quarta postice plus minus distincta), setis depressis aliisque 
curvatis et erectis vestitum, baud latum. Elytra pallide squamosa, setisque erectis sat 
crebre conspicueque vestita, grosse punctata, basi scutellum juxta utrinque tuberculata, 
humeris productis. Femora minus fortiter incrassata, tibiis anterioribus elongatis et 
gracilibus, tarsorum lobis parum magnis. $ $. Long, vix 4 — 475 mm. (Plate X. 
fig. 8.) 

Only a pair of this large species have been e.xamined. The elytra are much more 
elongate in the $ than in the $, but otherwise, apart from the usual sexual differences, 
the two are very similar structurally. The puncturation of the elytra is apparently very 
coarse and rough, but is not clearly seen owing to the squamosity. The species is 
easily recognised by its large size, very large and prominent eyes, long slender antennae, 
uneven setose thorax, the elytra somewhat impressed along the suture, the more than 
usually elongate and slender clavate hind femora, the long slender front tibiae, and for 
the size of the insect the small lobes of the front tarsi. It is allied to P. epitrachys of 
Maui. 

Hab. Hawaii, Kona, 3000 ft. September 1892, i % ; Olaa, 2000 ft. November 
1896, I ^. No doubt a rare species. 

(104) Proterhin2is vulcanus, sp. nov. 

.Niger, tarsis antennisque plus minusve rufescentibus. Antennae sat elongatae sed 
crassiusculae, clava distinctissime 3-articulata (sive articulo 8", quam 9"^ multo minore). 
Oculi magni, fortiter prominentes. Pronotum profunde 3-impressum, pallide (sat dense) 
vestitum, antice (saepe abrupte) angustatum. Elytra inaequalia, dorso subdeplanato, 
irregulariter aut maculatim cinereo-squamosa, setis erectis pallidis sat conspicuis, basi 
scutellum juxta utrinque tuberculata, humeris productis. Tarsorum anticorum articulus 
secundus major, lobis haudquaquam parvis. Rostrum % longitudinaliter rugulosum. 
t ?• Long, vix 3—3-25 mm. 

This species appears to be allied somewhat to the P. lecontei of Maui, and 
also perhaps to the preceding species (/-". kawaiiensis). There are distinct traces of 
longitudinal and transverse unevenness of the surface of the elytra, but it is much less 
marked than that of P. lecontei. The thickness of the antennae is no doubt variable, 



COLEOPTERA 



237 



and one t has these organs evidently more elongate and more slender than those of the 
other examples of either sex. The general resemblance in colour and clothing between 
the few individuals secured is very great. It is one of the darkest species, and of rather 
large size. 

Hab. Hawaii (3000 — 4000 ft.) ; rare, but widely distributed. Five examples only 
were secured in the Kona and Kau districts. 

(105) Proterhinus peles, sp. nov. 

Niger, lobis tarsorum antennisque nonnunquam rufescentibus, parce ferrugineo- 
squamosus. Antennae graciles, elongatae, clava distinctissime 3-articulata. Oculi 
magni, subconici, fortiter prominentes. Pronotum minus latum, minus profunde 
3-impressum, antice subabrupte angustatum, aequaliter nee dense squamosum. Elytra 
subinaequalia, sparsim ferrugineo-squamosa, setisque gracillimis albis sat conspicue, nee 
dense, vestita, basi utrinque scutellum juxta tuberculata, humeris acute productis. 
Femora postica minus fortiter incrassata, lobis tarsorum anticorum magnis. ^ %. Long. 

275—3 mm. 

A somewhat distinct-looking species, of dark colour and moderate size. The 
antennae are fairly long and quite slender, the club very well-marked, and consisting of 
three joints. The eyes and tarsal lobes are both large. The elytra are not much 
covered with squamosity, this being chiefly noticeable along the two by no means 
strongly marked longitudinal ridges. The suture is also a little raised. The humeral 
angles are produced and very acute, and the basal tubercles well marked, as also is the 
puncturation. The colour of the insect is black or obscure pitchy-black, the lobes of 
the tarsi and antennae sometimes paler. Rostrum of the female rugulose, dull, or 
hardly shining. 

Hab. Hawaii, Kilauea (4000 ft.) ; i t and i % taken in 1895. 

(106) Proterhiniis rufesce^is, sp. nov. 

Rufescens, capite, thorace, femoribusque, saepe obscurioribus, elytris plus minusve 
nigro-maculatis. Antennae minus fortiter elongatae, articulis saepe crassiusculis, non- 
nunquam sat gracilibus. Oculi magnitudine mediocres, prominuli. Pronotum sat 
fortiter 3-impressum, aequaliter subferrugineo-squamosum. Elytra rufa, nigro-maculata, 
subseriatim fortiter punctata, minus dense squamosa, albidis duabus postice maculis 
ornata, setisque albis elongatis rarius conspersa, fere aequaliter convexa, basi scutellum 
juxta utrinque tuberculata, humeris fortiter acute productis. Femora postica minus 
fortiter incrassata, lobis tarsorum anticorum magnitudine mediocribus. ^ %. Long. 

175—3 mm. 

31—2 



2 38 FAUNA HAWAIIENSIS 

This species in its typical condition is distinct enough from any other of those 
taken on Hawaii. The antennae (as above indicated) vary in stoutness, and the insect 
varies greatly in size, while the colour is sometimes much obscured, dark varieties being 
occasionally found. Minute examples naturally have the eyes and other parts less 
developed. In general, the red colour, and strongly produced humeral angles of the 
nearly evenly convex elytra, which are deeply and distinctly punctured (at least in clean 
examples), and which have two distinct spots of whitish squamosity, one on either side 
of the suture towards the apex, will distinguish it from the several following species, to 
which it is most nearly allied. It may be observed that there is no distinct transverse 
ridge formed between the vertex and front of the head as in some species on the other 
islands, to which this is otherwise very similar in appearance. 

Hab. Hawaii, Kau district (4000 ft.) ; 40 or more examples taken. 

(107) Proterhintis affinis, sp. nov. 

Elongatulus, minus latus, nigricans vel obscure rufescens, parte elytrorum dorsali 
tota vel ex majore parte rufa. Oculi magni. Antennae forma variabiles, saepius 
crassiusculae, nonnunquam graciles, articulo secundo brevissimo, clava distincte 3-articu- 
lata. Pronotum minus latum, vel subelongatum, minus dense pallide squamosum, plus 
minusve evidenter 3-impressum. Elytra irregulariter, plerumque maculatim, squamosa, 
dorso rufo, saepe subdepresso vel deplanato, basi utrinque scutellum juxta tuberculata, 
humeris distinctis, subacutis, parum productis. t ?• Long. 2-25 — 3-5 mm. 

This variable species in the smallest examples bears a great resemblance to large 
ones of P. gracilis, but the two are certainly distinct, the eyes and tarsal lobes 
being normally more developed in P. affinis, and the antennae thicker. These latter 
organs, however, exhibit some variability both in stoutness and length. Examples of 
the species taken in Kona generally have the elytra more evenly convex, less flattened 
or depressed above. In those from Kau, they are, in some cases, not only distinctly 
flattened, but show traces of longitudinal and transverse ridges. The humeral angles 
are distinct, although but little produced, and the tubercles in the region of the scutellum 
are distinct. The hind femora are not very strongly incrassate. There is no distinct 
ridee between the vertex and front of the head. 

Hab. Hawaii (3000—4000 ft.) ; about 50 examples were taken from various 
localities. 

(108) Proterhimis gracilis Sharp. 

Proterhinus gracilis Sharp, Tr. Ent. Soc. London, 1881, p. 529. 
This species in its typical condition is distinct enough, by its narrow and elongate 
form (especially in the ?), the slender antennae, and dark elytra, marked with red at the 



COLEOPTERA 



239 



base. I have assigned to it specimens of mucli more obscure appearance, and with the 
humeral angles of the elytra less produced, but it is quite possible some of these belong 
to another distinct species. In any case P. gj'acilis varies greatly in size, colour and 
length of the elytra. It is allied to the preceding species, P. affinis, as I have remarked 
under that insect. 

Hab. Hawaii. Various districts and localities (3000 and 4000 ft.); a great 
number of examples were taken. 



(109) Protcrhinus air/iynchus, sp. nov. 

Nigricans, elongatulus, elytris irregulariter pallide squamosus. Oculi magni, pro- 
minentes. Antennae subcrassae, rufescentes, clava 3-articulata. Rostrum ^ elongatum, 
aeque longum ac latum, vel paullo longius, lateribus pubescentia pallida densiore margi- 
natis. Pronotum minus latum, fortiter 3-impressum, setis appressis aliisque curvatis et 
suberectis vestitum. Elytra basim versus obscure rufescentia, ibique pallide squamosa, 
subinaequalia, basi scutellum juxta utrinque distincte tuberculata, angulis humeralibus 
subproductis et distinctis. $. Long, circiter 3 mm. (Plate X. fig. 9.) 

This species is very like P. affinis in form and general appearance, but is of a 
darker colour. The antennae are quite stout, the intermediate joints being a good deal 
widened from the base to apex, being of elongate triangular form. The $ is easily 
known by the long rostrum, which is even a little longer than its width at the base, and 
is clothed at the sides with a dense outstanding pale pubescence. I cannot assign any 
? to the two males with the above characters. 

Hab. Hawaii, Kilauea ; 2 $ taken in July 1895. 



(iio) Proterhinus tarsalis Blackb. 

Proterhinus tarsalis Blackburn, Tr. Dublin Soc. iii. 1S85, p. 171. 

Only three or four examples of this species were taken by me, the ? having the 
antennae a good deal more elongate, and thinner, than those obtained by Mr Blackburn. 
The species is a distinct one in superficial appearance, and is allied to P. vestitus Shp. 
ind P. angularis Shp., having the basal abdominal segment closely and strongly 
punctured beneath, as in those species. 

Hab. Hawaii, Mauna Loa, at an elevation of 6000 ft. (Blackburn) ; Kona 
(3000 ft.); a few examples taken. 



240 FAUNA HAWAIIENSIS 



(ill) Froterhinus hypotretus, sp. nov. 

Sordide niger, vel rufescens, elytris nigro-maculatis. Antennae minus fortiter 
elongatae, crassiusculae. Oculi minus fortiter prominentes. Pronotum minus dense 
squamosum, plerumque plus minusve evidenter 3- vel 4-impressum, impressione anteiiore 
sat distincta. Elytra maculatim squamosa, macula albida humerali aliisque posteria'ibus 
subrotundatis, sat distinctis, setisque erectis gracilibus albidis conspicue vestita, basi 
leviter emarginata, humeris distinctis, sed parum productis. Abdominis segmentum 
primum ventrale dense ac fortiter punctatum. Long. 2 '2 — 3 mm. 

Allied to the preceding species, but of wider form, and differing in colour, the 
insect being redder. Also closely allied to F. vestitus of Oahu, some varieties of 
F. angularis, &c., but certainly distinct from any of these. In fresh examples the 
maculate arrangement of the squamosity of the elytra, and the numerous erect fine 
setae, are very conspicuous. The elytra themselves are usually flattened or slightly 
impressed along the suture, the condition of the humeral angles is very similar to that 
seen in the same parts of F. vestitus. The lobes of the front tarsi are by no means 
large. 

Hab. Hawaii. 



(112) Froterhinus desqiianiatns, sp. nov. 

Ferrugineus (elytris cum capite nonnunquam obscuratis sive nigro-rufis), elongatus, 
parum squamosus. Antennae concolores, rufae, articulo secundo brevissimo. Prono- 
tum parum latum vel subelongatum, parum profunda 3-impressum, subaequaliter nee 
dense squamulosum, angulis posticis macula squamosa densiore signatis, lateribus 
rotundatis. Elytra elongata, vix squamosa, satis erectis pallidis, minus elongatis, 
sparsim sed conspicue vestita, dense fortiterque punctata, basi emarginata, humeris 
acutis. Abdominis segmentum primum ventrale fortiter et distincte punctatum. ^ %. 
Long, circiter 3 mm. 

On account of its elongate narrow form, ferruginous colour, and lack (at least for 
the most part) of squamous covering, this species could only be compared with F. fer- 
ruginetis, so far as the species of Hawaii are concerned. It is distinct at once from that 
insect by the distinctly and strongly punctured basal segment of the hind-body beneath. 
Extremely closely allied to F. detritus of Lanai, it may be distinguished from that 
species by the more evenly rounded sides of the prothorax, which are not so strongly 
and abruptly narrowed in front, and the puncturation of the elytra is apparently rather 
less coarse. 

Hab. Hawaii, Kona (2000 — 3000 ft.) ; rare, 2 ^ and i $ only having been taken. 



COLEOPTERA 241 

(113) Proterhiims ferrtigineus, sp. nov. 

Elongatus, ferrugineus, elytris saepe nigro-notatis, setis appressis erectisque ves- 
titus, hand squamosus. Antennae concolores, ferrugineae. Pronotum haud latum, setis 
appressis, aliisque curvatis et suberectis latera versus vestitum, plus minusve evidenter 
3-impressum, antice saepe abruptius constrictum. Elytra fortiter elongata, haud squa- 
mosa, setis erectis brevissimis, aliisque longioribus, conspicue et densius vestita, fortiter 
dense punctata, basi scutellum juxta utrinque tuberculata, angulis humeralibus productis 
et subacutis. Femora postica gracilia, parum fortiter clavata. Abdominis segmentum 
primum ventrale circa medium parum distincte punctatum, granulatim asperulum. Z ?■ 
Long. 3 — 4'2 mm. 

Very closely allied to the Oahuan P. longuliis, of which species it is the repre- 
sentative on Hawaii. Of P. longulits I have only seen the types in Dr Sharp's 
colkction, but P. feri'-ugineus is no doubt on the average a larger species, and yet has 
the hind femora still less clavate, and also appears to differ slightly in the sculpture and 
clothing of the elytra, and in other small points. 

Hab. Hawaii (4000 ft.). In the leaf-stems of the tree-ferns, to which P. longuliis 
is also attached. 

(114) Proterhinus similis Blackb. 

Proterhimis similis Blackburn, Tr. Dublin Soc. iii. 1885, p. 170. 

Species maxime variabilis, antennis nigris vel nigrescentibus, articulis 2 basalibus 
haud distincte rufis, elytrorum basi saepe emarginata, sed angulis humeralibus ipsis 
haud acute productis, saepe minus distinctis. ^ %. Long. 2 — 3 "5 mm. 

This species is so extraordinarily variable, that it would be useless to draw up any 
lengthy and minute description, as it would apply to but few examples. It belongs to 
the section in which the humeral angles themselves are not distinctly produced forwards 
and acute, although the base of the elytra is often emarginate. The insect is black, red, 
or parti-coloured, the elytra sometimes entirely covered, sometimes nearly free from 
squamosity, which varies in colour from golden to grey ; sometimes they are maculately 
squamose. Often they are depressed or flattened, often evenly convex, but nearly 
always they appear slightly elevated at the extreme base near the scutellum, and at that 
point have a small area along the suture free from squamosity, even when elsewhere 
they are entirely covered. The antennae are exceedingly variable, short or of medium 
length, with the joints robust and minutely asperated, or decidedly elongate, even the 
second joint being sometimes somewhat long and slender, and there are various inter- 
mediate conditions. In colour they are black, even the basal joints being black or at 
least dark. This character readily distinguishes the species from the other species of 
this group (the Hawaii form oi P. decepior), which has the basal two joints (at least) of 



242 FAUNA HAWAIIENSIS 

the antennae clear red, and the second joint more rounded. In those examples with 
the longest and most slender antennae, the front tibiae are often much lengthened, and 
the tarsal lobes appear to be very minute, and decidedly smaller than are those of 
certain other forms. After the examination of a vast amount of material (consisting of 
hundreds of examples) I have come to the conclusion that this species is at the present 
time in the process of becoming differentiated into a number of distinct forms. Some 
of these forms indeed may already be really distinct species, but to decide whether this 
is the case would probably require very careful observations of the living insect in 
various localities. 

Hab. Hawaii. Common on the mountains, especially at an elevation of about 
4000 ft. The whole insect is often covered with a muddy substance. 

G. Species quae duas vel complures insulas incolunt. 

(115) Proterhinus innotabilis, sp. nov. 

Nigricans vel piceus, rare rufescens, elytrorum basi plerumque rufescente et pallide 
squamosa. Oculi prominuli. Antennae graciliores, minus fortiter elongatae. Capitis 
vertex a fronte linea transversa elevata (nonnunquam fracta) divisus. Pronotum minus 
latum, subferrugineo-squamosum, fortiter et distincte 3-impressum. Elytra minus elon- 
gata, fortiter crebre punctata, sutura saepe elevatula, basi plerumque rufescente, ibique 
pallide squamosa, postice maculis duabus albidis squamosis utrinque suturam juxta 
saepe notata, setis erectis albidis minus fortiter elongatis sparsim vestita, ad basim 
utrinque scutellum juxta tuberculata {tuberculis saepe albido-squamosis), angulis ipsis 
humeralibus fortiter acute productis. Z ?■ Long, (exemplorum in insula Maui capt.) 
2 — 3 mm. 

So far as examples from Maui are concerned this is a small species, or at least only 
of moderate size in very largely developed individuals. It is one of the most obscure 
of the numerous species found on Haleakala, and is allied to P. epickrystts and P, brevi- 
pennis, but cannot be referred to either. On Molokai I have taken specimens of a 
Proterhinus exceeding like P. innotabilis but of larger average size, and which appa- 
rently pass from this form into one, which becomes largely different to the typical 
Haleakala form, and indeed in many respects approaches the Molokai species P. con- 
vexiusciilits. These examples from Molokai are themselves a most puzzling series, and 
may represent more than one species, but I have not cared to consider them at present 
as more than varieties of P. innotabilis, especially as this species is itself not a little 
variable. 

Hab. Maui, Haleakala (4000 — 5000 ft.), Molokai. Not rare, many examples 
having been taken. The examples taken on Molokai are apparently referable to the 
same species. 



COLEOPTERA 24: 



(116) Proterhinus angularis Sharp. 

Proterhinus angularis Sharp, Tr. Ent. Soc. London, 188 r, p. 530. 

An extremely variable species, no examples of the very long series examined 
agreeing altogether with the original types. In the most common form the antennae 
are long and slender, and clothed with stiff and conspicuous setae. Thorax very 
variable in shape, generally rather narrow, rarely constricted in front as in the type. 
The anterior depression is usually deep and distinct, the two posterior ones often 
obsolete. The posterior angles are not distinctly marked by spots of dense squamosity. 
Elytra long and narrow, shoulders always produced, but not very sharp, marked with a 
patch of pale squamosity, and behind there are often a number of more or less distinct 
and roundish pale squamous spots. 

Small examples are often of extremely narrow and elongate form. The dark 
colour, form of the humeral angles of the elytra, and patches of pale squamosity thereon, 
and the strongly and closely punctured basal ventral segment of the abdomen are the 
most constant characters. The type specimens are remarkable for the more obscure 
colour of the very conspicuous erect setae on the elytra, and I have only seen a few- 
examples resembling them in this respect. Usually the longer setae are quite white, 
and I am by no means satisfied as to the identity of the two forms, which may be 
closely allied and variable but distinct species. The length varies from 2 — 2>'5 ^n^n- 

Hab. Oahu, Molokai, Maui, Lanai, Hawaii. Generally attached to Straussia. 
Variable in each locality. 



(117) Protcrhiims dispar Sharp. 

Proterhinus dispar Sharp, Tr. Ent. Soc. London, 1881, p. 52S; Tr. Dublin Soc. 
III. 1885, PI. V. fig. 41, ?. 

I have seen but few Oahuan examples of this species, but have taken it very freely 
on Molokai, and have also met with it on Lanai. It varies very greatly in size and 
otherwise, and the distinctive characters become much obscured in diminutive .specimens. 
Some examples from Oahu have the head and thorax as well as part of the elytra of a 
distinct red colour, and the antennae vary from black to red. The great differences in 
size are not a sexual character, for of the Oahuan examples taken by me one of the 
females is twice as large as either of the males, and e.xactly the reverse is the case with 
the types in Dr Sharp's collection. The species is an interesting one on account of the 
F. H. II. 32 



244 FAUNA HAWAIIENSIS 

rather large development of the rostrum in the ^, and in the same sex the posterior 
coxae are much less widely separated than in many species, much less widely than the 
front pair. In the % the distance between the hind coxae is greater, so that it does not 
greatly differ from the majority of species in this respect. 

Hab. Oahu, Molokai and Lanai ; attached to Wikstroemia. 



(ii8) Proterhinus alyxiae, sp. nov. 

Latus, brevis, robustus, ferrugineus, pulcherrime squamosus. Antennae elongatae, 
articulis basalibus notabiliter robustis (secundo subquadrato), clavam versus sensim 
gracilescentibus, clava gracili, vix evidenter 3-articulata. Pronotum trans medium latis- 
simum, perinaequale, antice transversim fortiter impressum, parte posteriore fortiter 
elevata, antice posticeque ferrugineum, caetera parte densissime pallido-squamosa. 
Elytra brevia, lata, submaculatim squamosa, setisque albidis erectis conspersim vestita, 
dorso juxta basim impresso, ibique nigricante, basi emarginata, humeris productis. 
Femora omnia fortiter incrassata. Z ?• Long, vix 3 — 3 '2 mm. (Plate IX. fig. 23, ^.) 

Var. pauper. $ $ antennis gracilioribus, ? statura multo minore et forma graci- 
liore distinguenda. $ $. Long. $ 275, $ 2 mm. 

This beautiful species, on account of its short, wide form, reminds one at first sight 
of some Kauai species. It is not really closely allied to these, however, but it is so to 
P. calliphyas of Maui. It is distinct from that species by the accentuation of most of 
those characters, which render the latter remarkable. The distinction between the 
dorsum and flanks of the prothorax is clearly marked by the sharp edge formed at their 
meeting. 

Hab. Molokai mountains (3000 ft.); rare, taken from stems oi Alyxia. Four 
females (var. pauper) taken on Lanai are only half or less than half the size of the 
smaller Molokai examples, and have much more slender antennae. The single <J taken 
with these is about as large as the type, but its antennae are decidedly thinner. 



(119) Proterhinus navita, sp. nov. 

Haud latus, rufescens, sat dense pallide squamosus, elytris saepe plus minusve 
nigro-notatis. Antennarum ^ articuli 2 basales perconspicue robusti, nigricolores ; 
articuli sequentes graciliores, plerumque evidenter, sed minute, tuberculati, clava parum 
distincte 3-articulata (sive fere 2-articuIata). Pronotum densius aureo-squamosum, antice 
plus minus impressum. Elytra aureo- vel griseo-squamosa, setisque brevissimis nonnullis 
erectis inconspicue vestita, angulis humeralibus obscuris. Tarsorum anticorum articulus 
secundus minor, lobis minoribus. ^ $. Long. i'5 — 3 mm. 



COLEOPTERA 245 

The male of this species is distinguished chiefly by the stoutness of the two basal 
joints of the antennae, the first, in well-developed examples at least, being very stout 
and subtriangular in form, the second very short and quadrate, and their colour is black 
or nearly so. Normally it is a red insect, with only some fuscous or black spots on the 
elytra, and even these are sometimes absent, but in some examples the black colour is 
a good deal extended over the insect, the basal half of the elytra remaining pale. In 
general the $ resembles the ^, but the peculiar character of the basal antennal joints is 
not always so pronounced. In faded examples the squamosity is grey, instead of 
golden. 

Hab. Lanai (2000 — 3000 ft.) ; Molokai (3000 ft.). 



( 1 20) Proterhinus debilis Sharp. 

Proterkinus debilis Sharp, Tr. Ent. Soc. London, 1878, p. 19. 

I have not met with any examples on Oahu that agree with this species, and have 
only examined three or four specimens, so I do not know whether the antennae vary in 
structure. In its typical form it is quite distinct from any other Oahuan species. Mr 
Blackburn also referred some specimens taken on Hawaii to this species. 

Hab. Oahu, Waianae mountains ; and on Hawaii (Blackburn). 

(121) Proterhimts deceptor, sp. nov. 

Rufescens, elytris latera versus nigris vel nigro-notatis, horum angulis humera- 
libus haud distinctis. P. oscillanti simillimus sed setis elytrorum erectis magis distinctis, 
et rostro % evidenter minus elongate et latiore. ^ ?. Long. 175 — 2*25 mm. 

This species is very similar in most respects to P. oscillans, the humeral angles of 
the elytra being usually very indistinct or effaced. The squamosity, which likewise has 
a tendency to form a number of roundish spots on the elytra, is thereon greyish or 
silvery rather than golden. Their erect setae in fresh examples are longer, more 
numerous and much more evident, and so also are those upon the legs. Usually the 
elytra are red, and black or infuscate only along the lateral margins, but sometimes 
about the middle the black colour extends inwards to form lateral spots as in oscillans. 
These distinctions alone would perhaps appear hardly sufficient for the formation of 
another species, were it not for the fact that the rostrum of the % is decidedly shorter 
and wider than that of oscillans, and is not distinctly narrowed towards the base. 

Hab. Oahu. Waianae range. Examples from slightly different localities do not 
altogether agree, and it is uncertain whether the series examined is really all of one 
species. 



246 FAUNA HAWAIIENSIS 

Obs. On Kauai a variable insect is found, which I refer to the above species. 
These Kauai examples are generally darker than those from Oahu, and have the elytra 
often entirely blackish, or only obscurely red. On Molokai or Lanai the species is 
again found, and the individuals again are subject to much variation. On Hawaii the 
individuals attain a considerably larger size than the type, and on this island form two 
distinct races, one found on the windward and southern parts of the island, the other in 
the Kona district on the west. These Kona examples are readily distinguished from 
the others by the more elongate and conspicuous erect setae of the elytra, and they are 
of smaller average size. For these two races I propose the names P. deceptor van 
major and van konafuis. 



(122) Proterhinus blackburni Sharp. 

Proterhinus blackhirni Sharp, Tn Ent. Soc. London, 1878, p. 17. 

P. hystrix Sharp, op. cit. 1881, p. 527, =var. of this species. 

(Plate X. fig. 5, ^, van disignatus.) 

This species varies greatly in size, colour, form of thorax, length of elytra, &c., and 
I do not think the P. hystrix Sh. is a distinct species. On Kauai there are two well- 
marked varieties, the van eiigeniae, van n., of somewhat robust form, the thorax with 
strongly prominent lateral angles in front, the erect setae of the elytra dense, and with a 
good deal of pale appressed clothing, which tends to form four spots, the antennae 
longish and slender ; and the van bisignatus, van n., which a good deal resembles the 
preceding form, but is darker, and has the basal tubercles of the elytra conspicuously 
marked with white or silvery appressed setae. The sculpture and clothing of this 
species is often entirely concealed by a mud-like covering, and it is not improbable that 
I have included several distinct species under one name, for it is one of the most 
difficult species to study, and many of the examples taken are in indifferent condition, 
but supposing there are several allied species, they are certainly all variable. 

Hab. Found on all the islands of the group, in the forests from 1500 — 4000 ft. ; 
the var. hystrix on Hawaii, van bisignatus peculiar to Kauai, and var. engeniae on the 
same island, but with very similar examples on Molokai. 



COLEOPTERA HETEROMERA. 

All the Heteromera, with the exception of the Cistelidae, are probably foreign. 
Those that are not known at present to occur outside the islands will probably be 
discovered elsewhere as the study of Coleoptera progresses. 



COLEOPTERA 247 

Fam. OEDEMERIDAE. 

OxAcis Leconte. 

(i) Oxacis collaris Shp. 

Ananca collaris Sharp, Tr. Dublin Soc. 1885, p. 169. 

Hab. Oahu, Molokai, and probably all the islands on the plains. I observed this 
species in great numbers on the quarantine island at Honolulu, where it was crawling 
over shrubs covered with the scales of Ceroplastes. 

Fam. ANTHICIDAE. 

Anthicus Payk. 

(i) Anthicus oceanicus Laf. 

Anthicus oceanicus Laferte, Mon. Anth. p. 1 70. 

Hab. Kauai, Oahu and Maui and probably all the islands ; on the coast. 

(2) Anthicjis mjindtclus Shp. 

Anthicus mundulus Sharp, Tr. Dublin Soc. 1885, p. 168. 

Hab. Oahu and Kauai (Blackburn). Salt marshes near sea-level. 

Fam. CISTELIDAE. 

The Cistelidae are represented by two genera, Cistela with seven, and Labetis with 
three ^ species. The latter genus is peculiar to the islands, as are all 10 of the species 
of this family. All are true forest insects, and are certainly indigenous. Species of 
both genera have been found on Kauai and Hawaii, but not on all of the intermediate 
islands, so that no doubt several species are yet to be discovered. So far as is known 
no species of either genus extends its range to two of the islands. 

' The two examples taken on Oahu by Mr Blackburn and which probably represent other two distinct 
species are not included in these remarks. 



248 FAUNA HAWAIIENSIS 

CiSTELA Fabr. 

(i) Cistela crassicornis Sharp. 

Cistela crassicornis Sharp, Tr. Dublin Soc. 1885, p. 168, PI. IV. fig. 25. 

Hab. Oahu. Mountains near Honolulu, Palolo valley (Blackburn) ; my single 
example was taken at the head of Pauoa valley, which is not far distant. It is, I have 
no doubt, of the % sex. It has the elytra densely punctured, and I fancy that Sharp's 
description of the ^ ' elytris parcius punctatis ' is rather apparent than real and is due to 
the condition of the specimen, which is not thoroughly clean. 

(2) Cistela kauaiensis, sp. nov. 

Rufo-brunnea, ad colorem testaceum varians, capita thoraceque opacis, densissime 
punctatis, elytris subnitidis, suturam versus plus minusve evidenter striatis, ibique saepe 
impressis, sat crebre punctatis, punctis nonnullis seriatim dispositis. Long. 7*5 — 9 mm. 
Antennarum 4 — 4' 2 5 mm. (Plate X. fig. 22.) 

Extremely like C. crassicornis but larger and with very evidently longer antennae. 
The species agrees generally in sculpture with that species. It varies a little in the 
length and structure of the antennae, irrespective of sex, but even in the most highly 
developed ^ the joints are elongate even towards the apex. In this sex the three which 
precede the apical joint are distinctly grooved beneath. The apical ventral segment 
in the t has the apical margin somewhat raised, and this is to some extent the case in 
some of the $$, but usually this segment in the Z has a more flattened or impressed 
form than in the other. The apical joints of the antennae in the % are narrower than 
the ^, and but little serrate inwardly. 

Hab. Kauai 4000 ft. Not common. 

Obs. Two or three examples from Lihue on the same island taken at an elevation 
of 3000 ft. have shorter antennae and are only of the size of the smaller individuals of 
C. kaziaiensis. They are therefore somewhat intermediate between that species and C. 
crassicornis but probably are distinct from either. 

(3) Cistela nigricollis, sp. nov. 

Nigra, elytris brunneis, antennis pedibusque testaceis, capite nitido. Long. 7 mm., 
antenn. 3'5 mm. $. 

Very closely allied to C. kauaiensis, but smaller and readily distinguished by the 
black head and thorax, the former in front being distinctly punctured, not very densely 



COLEOPTERA 249 

and rugosely, and with the surface between the punctures distinctly shining. The joints 
of the antennae (^) towards the apex are also rather shorter and wider. 

A single example taken at Halemanu, Kauai, is larger with longer elytra and has 
the front of the head more closely punctured, but otherwise resembles the type. I 
should think it is probably a distinct species. 

Hab. Kauai. High plateau (4000 ft.). 

(4) Cistela subaenescens, sp. nov. 

Nigra ($ piceo-nigra), pedibus antennisque testaceis, his apices versus obscurioribus, 
elytris subaeneo-micantibus. Long, circa 7 mm., antenn. circa 3 mm. 

Black, or in the ? piceous, the elytra with distinct aeneous glitter. Head and 
thorax densely punctured. Antennae of the ^ with the terminal joints shortish, and 
serrate inwardly, deeply grooved beneath, less wide in the ?, testaceous at the base, 
becoming infuscate towards the apex. Elytra shining, pubescent, aeneous, somewhat 
densely punctured. 

Hab. Oahu, Waianae mountains (3000 ft.). 

(5) Cistela apicalis, sp. nov. 

Elongata, angustula, nigricans, antennis, pedibus et parte elytrorum apicali, testa- 
ceis, oculis permagnis. Long. 7 "6 mm. 

Extremely like the preceding, a little more elongate, the elytra with a very faint 
indication of brassy reflection in certain lights, testaceous on their apical portion, less 
shining, and less strongly and more obscurely punctured than in C. subaenescens, the 
antennae entirely clear testaceous, and the eyes decidedly larger. 

Hab. Oahu. Two examples were taken in the mountains at Honolulu in the 
summer of 1896. 

(6) Cistela montana, sp. nov. 

Nigricans vel piceus, elytris aeneo-micantibus, antennis testaceis, apices versus 
saepe obscurioribus, antennis brevioribus, articulis apicalibus parum fortiter dilatatis. 
Long. 6 — 6'5 mm., antenn. vix 2*5 mm. 

A rather smaller species than C. subaenescens, which it greatly resembles, and easily 
distinguished by the shorter antennae, which have the apical joints much less strongly 
dilated in the $. In colour and sculpture the two species are identical, and one e.xample 
of a rufo-piceous colour I suspect is the $ of C. montana. 

Hab. Kauai. Four examples were taken on the high plateau. 



2 50 FAUNA HAWAIIENSIS 



(7) Cistela konae, sp. nov. 

Fusco-brunneus, pedibus antennarumque basi testaceis, elytris nitidis, submetallicis, 
sat evidenter sed parum profunde striatis, crebre punctatis, punctis nonnullis seriatim 
dispositis, interstitiis convexiusculis. Long, circa 6 mm. 

Of an obscure brownish colour, the head and prothorax darker, the face in front of 
the antennae and the legs testaceous. Head very densely punctured, the eyes by no 
means large. Prothorax nearly dull, very densely punctured, strongly transverse, 
truncate or slightly emarginate in front. Elytra shining, distinctly and closely punc- 
tured, with the interstices even towards the sides quite distinctly convex, so that the 
striation is more evident than in any other species. In the single example taken only 
the basal joints of the antennae remain, and these present no noteworthy characters. 
Perhaps most nearly allied to C. montana, but quite distinct from this and other species 
by the condition of the elytral interstices, and also apparently remarkable for the very 
wide prothorax, the sides of which are nearly straight and parallel for about two-thirds 
of their length from the base. The eyes are smaller than in most species of the genus. 
The single mutilated example is probably a %. 

Hab. Hawaii, Kona, 5000 ft. A single example (mutilated) was taken from a 
spider's web. 



Labetis Waterhouse. 

(i) Labetis hawaiiensis, sp. nov. 

Nigra, subnitida, elytris testaceis vel brunneis, antennis (articulis basalibus exceptis) 
nigricantibus, pedibus testaceis. Long. 10 — 12 mm. Antenn. 5 — 7 mm. 

Head black, closely punctured in front, but the surface more or less shining 
between the punctures. Prothorax black, the surface shining, closely punctured, with a 
narrow smooth median longitudinal impression, more distinct in some examples than in 
others, and in some there are also indefinite impressions, rendering the surface more or 
less uneven. Elytra varying in colour from testaceous to brownish, striate, the striae 
nearest the suture often deeper than the outer ones, closely punctured, the interstices 
conspicuously punctate, the punctures closer in some examples than in others, the 
surface shining. At the apex the first and second interstices are about equally convex. 
Anterior tibiae with the superior apical angle greatly produced (varying in development), 
and rounded at the apex. Abdomen beneath black, finely punctured, the apical 
segment very deeply impressed in the ^, and faintly but evidently so in the %. 

Hab. Hawaii, Kilauea ; a series of over a dozen examples taken in August. 



COLEOPTERA 251 

(2) Label is tibialis Waterhouse. 

Labelis tibialis Waterh., Ent. Mag. xv. p. 267. 

Haud nitida, testacea, pronoto saepe fusco-testaceo, antennis plerumque testaceis, 
elytrorum interstitio primo et secundo apicem versus subaequaliter convexis, haud 
inaequaliter elevatis. Long. io"5 — 12'5 mm. 

Closely allied to the preceding, but easily distinguished by the dull surface, and the 
pale colour of the prothorax, the antennae also are pale. One example has the prothorax 
largely blackish, but it is pallid about the middle. Abdomen black (or nearly so) 
beneath. The depth of the impression of the apical ventral segment of the ^ evidently 
varies. 

Hab. Kauai, 4000 ft., on the high plateau. That the examples taken by 
Blackburn on Oahu and referred to this species are, as I suspected, distinct, has been 
confirmed for me by Mr Waterhouse, who kindly examined the Oahuan specimen in 
the British Museum. Probably the Oahuan species is very near to if not identical 
with my L. /lazuaiieusis. The Blackburnian collection should contain two species from 
Oahu, probably one from either mountain range (vide Tr. Dublin Soc. 1885, p. 248), 
for there is no such variation in size in the species of this genus, as that given by 
Blackburn for his two examples (1. c. p. 167), one of which is much smaller than the 
other known species of the genus, the other probably the largest. 

(3) Labelis coniilans, sp. nov. 

Praecedenti cognatissima, opaca, testacea, capite pronotoque nonnunquam plus 
minus infuscatis, hoc saepe piceo, elytrorum interstitio secundo apicem versus fortissime 
convexo-elevato, primo plus minus obsoleto, abdominis segmentis ventralibus (saltem ex 
magna parte) pallidis. Long. 10 — 12-5 mm. (Plate X. fig. 21.) 

Quite like the preceding in general appearance, and differing constantly, so far as I 
can see, only in the paler underparts of the body, and the sculpture of the elytra near 
their apex. In this species the second interstice is extremely convex, as also is the 
sutural margin, while the first interstice is little or not at all raised, and a deep groove 
is thereby formed between the suture and the second interstice. The elytral interstices 
are generally more strongly and densely punctured than in either of the preceding 
species, but all vary in this respect. 

Hab. Kauai. Taken in company with L. tibialis. Of the 18 examples of 
Labetis taken in company, 8 belong to the present species, and 10 to Z. tibialis. I can 
detect no variety in the least intermediate between these, and therefore must consider 
them distinct. Nearly all the examples of both are of the ^ sex. 

F. H. II. 33 



252 FAUNA HAWAIIENSIS 



(4) Labetis, sp. ? 

Hab. Oahu. Mountains near Honolulu (Blackburn). See remarks under 
L. tibialis (ante). 

(5) Labetis, sp. ? 

Hab. Oahu (PWaianae mountains), Blackburn. See remarks unAe.r L. tibialis 
(ante). This and the preceding species are stated (Tr. Dublin Soc. 1885, p. 248) to 
have been obtained in localities 20 miles apart. 



Fam. TENEBRIONIDAE. 

Epitragus Latr. 
(i) Epitragus diremptzis Karsch. 

Epitragus diremptus Karsch, Berlin. Ent. Zeitschr. xxv. p. 6. 
Hab. Generally distributed throughout the islands. 

Opatrum Fabr 

(i) Opatrum seriatum Boisd. 

Opatru7n seriatum Boisd., Voy. Astr. Col. p. 252. 
Hab. All the islands of the group, on the lowlands. 

Platydema Cast. 

(i) Platydema obscurum Shp. 

Platydema obscurum Sharp, Tr. Dublin Soc. 1885, p. 166. 
Hab. Oahu. Probably characteristic of the lowlands. 

Gnathocerus Thunb. 

(i) Gnathocerus cormctus Fabr. 

Trogosita cornuta Fabr., Ent. Syst. Supp. p. 51. 
Hab. Oahu, Honolulu. 



COLEOPTERA 253 

Tribolium Macl. 

( I ) Tribolium ferrugiiieum Fabr. 

Tencbrio ferruginejis Fabr., Sp. Ins. i. p. 324. 
Hab. Oahu, Honolulu &c., in food. 

Alphitobius Steph. 
(i) Alphitobius diaperinns Panz. 
Tenebrio diaperinus Panz., Ins. Germ. 37, 16. 
Hab. Lowlands of most or all the islands. 

(2) Alphitobius piccus 01. 
Helops piceus 01., Ent. in. 58, p. 17. 
Hab. Generally distributed in the islands. 

(3) Alphitobius lateralis Boh. 
Heterophaga lateralis Boh., Eug. Res. 1858, p. 94. 

Hab. Oahu, Honolulu. — Maui. Introduced, probably from China. This species 
was kindly determined for me by Mr Champion. 

ScioPHAGUS Sharp. 

(i) Sciophagus pandanicola Fairm. 

Heterophaga pandaiiicola Fairm., Rev. Zool. 1849, p. 446. 
Hab. Oahu and Kauai (Blackburn). 



Fam. CIOIDAE. 



The Cioidae is a family of doubtful position. They are represented by 42 species, 
29 of which are referred to the genus Cis, and 13 to a new genus, Apterocis. The 
members of the latter genus are the most remarkable island representatives of the 
family, although most of those assigned to Cis are themselves of peculiar facies, and 
very unlike the ordinary representatives of the genus. There is however no doubt 
that both these genera, as represented in the islands, will hereafter be further divided. 

33—2 



254 FAUNA HAWAIIENSIS 

Thus in Apterocis some of the species e.g. A. ephistenioides are quite regularly 
convex, the prothorax being very closely adapted to and continuing the curve of the 
elytra. In others, e.g. A. rufo-notatus, A. variegatiis, &c., the outline is not regularly 
convex, but is interrupted at the point of contact of the elytra and prothora.x. Some of 
the species of this latter group closely resemble A. ephistemoidcs in general appearance, 
but others e.g. A. variegatus have a very coarse sculpture, and one {^A. hystrix) is 
unique in the genus as being conspicuously clothed with erect hairs. We also notice 
differences in the structure of the mesosternum in front of the coxae, and in A. varie- 
gatus and probably in the allied A. omatipennis, the metasternum is less short than is 
normal in the genus. 

Of the species assigned to Cis, C. pacificiis is the most commonplace and 
greatly resembles European species, and will probably be found to be not indigenous. 
It has some resemblance to C. breviformis but to no other species, and the latter also 
has a very different appearance to the truly indigenous species, and I suspect may 
prove to be also foreign. Another species C. alienus may also prove to have 
been introduced. The remaining 26 species are all of peculiar facies, and exhibit 
considerable variety of form and appearance, and must be considered as peculiar to the 
islands. Cis violokaioisis (and probably C. kaleakalae, the unique example of which 
could not be examined from this point) is wingless or nearly so, but we have not 
separated it generically on that account, because the wings of another species, C. fallax, 
are in a more or less rudimentary condition, and we are not even certain whether this 
species is more than a variety of the normally full-winged C. signatus. Besides 
these species, C. miiints and C. diviinuiivus appear to me to belong to the same 
group, and will probably prove to have the wings either absent or rudimentary, 
the abortion of the wings being accompanied, in most of the species at least, by a 
shortening of the metasternum. In these characters it will be seen that the insects 
above mentioned approach to the genus Apterocis, but they lack the oval and highly 
convex form, which is so striking a feature of the latter. Nevertheless the complex 
affinities between the Hawaiian species of Cis (excluding the doubtfully indigenous 
forms) and Apterocis render it highly probable that the latter originated in the islands, 
through forms somewhat similar to some of the species of Cis, which now inhabit 
them. 

Some few of the species of Cis are found in large fungi growing e.xternally on 
trees, but the greater number are attached to dead branches of trees, or found beneath 
dead bark, where they no doubt feed on the smaller fungi which grow in such places. 
Individuals of many of the species are numerous, and they are no doubt parasitised by 
the minute Hymenoptera of the genera Sierola and Scleroderma, which are taken 
in their company. We have lately bred forms (winged and wingless) allied to these 
Hymenoptera from the burrows of Eimearthron, another genus of Cioidae, in this 
country. 



COLEOPTERA 255 



Cis Latr. 



(i) Cis alienus Sharp. 

Cis alienus Sharp, Tr. Ent. Soc. London, 1879, p. 91. 

This is very different to any other Hawaiian species, and is probably either a 
natural immigrant, or has been introduced by man. Its general appearance, the 
clothing of the under parts, the great length of the prosternum in front of the coxae &c., 
are all foreign to the true native type. 

Hab. Oahu, Honolulu, mountains ; two examples taken (Blackburn) ; Waianae 
mountains, several examples, Perkins. It is probably a scarce insect. 



(2) Cis pacijictts Sharp. 

Cis pacificiis Sharp, Tr. Ent. Soc. London, 1879, p. 91. 

The most common-place species of the genus as represented in the islands, and 
probably not truly indigenous. The processes on the head of the ^ vary greatly 
in development. 

Hab. Throughout the islands in the mountain forests. 



(3) Cis breviformis, sp. nov. 

Parum elongatus, ferrugineus, pronoto plus minusve infuscato, pedibus antennis- 
que rufo-testaceis, harum articulis apicalibus nigricantibus. Pronotum opacum vel 
minus nitidum, brevissime griseo-pubescens, sat latum, densius subtiliter punctatum. 
Elytra breviuscula, griseo-pubescentia, haud maculata, parum profunde ruguloso- 
punctata. Long. 2 mm. 

Somewhat like Cis pacificus in form, and quite unlike any other Hawaiian species. 
The thoracic puncturation is rather close and very fine, that of the elytra is larger, but 
shallow, rugose and irregular. Both the thorax and elytra are clothed with very short 
pale setae, which hardly stand out from the surface, but are quite conspicuous, and with 
the short form and ferruginous colour give the species a very distinctive appearance. 

Hab. Molokai. Two examples taken at an elevation of above 4000 ft. 



2 56 FAUNA HAWAIIENSIS 

{4) Cis nesiotes, sp. nov. 

Subcylindricus, haud fortiter convexus, pronoto aeneo, antice posticeque saepe 
pallescente, elytris testaceis, marginibus lateralibus fasciaque transversa circa media 
nigris, setulis pallidis brevissimis inconspicue vestitis. Pronotum nitidum distincte sat 
fortiter punctatum. Elytra grosse denseque rugoso-punctata. Pedes articulique anten- 
narum basales testacei, femoribus saepe antennarumque clava nigricantibus. Long, vix 
2 — 2'5 mm. 

A distinct species with the thorax always brassy or coppery, elytra pale, black 
along the margins and about the middle of their length with a transverse fascia, which 
forms two sharp angles. This fascia is sometimes broken up into detached spots. 
The species is chiefly remarkable for the coarse (but shallow) and very rugose 
puncturation of the elytra, which even on the apical portions is hardly different in 
character. The very short pallid setae are quite evident but not conspicuous. There 
are no long hairs on any part of the elytra. There is considerable variation in the 
puncturation of the prothorax, the punctures being very much coarser and closer in 
some examples than in others, and in many the surface of this part is evidently uneven. 

(5) Cis cognatissiinus, sp. nov. 

Praecedenti forma et colore simillimus, pronoto aeneo, nitido distincte punctato, 
elytris testaceis nigro-signatis, basi grossius rugoso-punctata, setulis pallidis brevissimis 
vestitis, apicem versus minus dense rugoso-punctatis et laevioribus. Long. 17 — 
2"25 mm. 

Extremely like the preceding, and very variable in size ; in some examples the 
median transverse fascia of the elytra is represented by two spots, but usually it forms 
a sharp angular mark on either wing-case just as in the preceding. The thorax is 
distinctly and clearly punctured, more closely in some examples than others, the surface 
shining. The elytra are less rugosely punctured, towards the apex a good deal 
smoother and the punctures less close and rugose than in C. nesiotes, and for this 
reason the two forms cannot be considered identical. The clothing is of the same 
character in both the species. 

Hab. Kauai, Oahu, Lanai, Molokai, Hawaii, in the mountains from 2000 — 
4000 ft. 

(6) Cis bicolor Sharp. 

Cis bicolor Sharp, Tr. Ent. Soc. London, 1879, p. 93; Blackburn, Tr. Dublin 

Soc. 1885, p. 163, PI. IV. fig. 22. 
This is a distinct species and is I believe quite distinct from C. tabidiis Shp., 
which in Tr. Dublin Soc. (1. c. supra) was sunk as a variety of it, as well as from the 



COLEOPTERA 257 

forms with aeneous thorax mentioned in Blackburn's note (1. c). The latter at least in 
part are no doubt referable to Cis cognatissinius or C. nesiotes, or to both of these 
species. Usually C. bicolor may be distinguished at a glance from these species by its 
general appearance. The prothorax is black, not aeneous, and is widely pale in front 
and more or less so posteriorly. The elytra are testaceous with a black spot on each 
about the middle, and in front of these is another spot placed transversely across 
the suture. This transverse spot is absent in only a few e.xamples in a long series 
and it is doubtful whether these are really specifically identical with the others. The 
elytra in very fresh specimens bear some excessively short setae rather like those of 
C. cognatissimus, but there is no trace of any long fine hairs at the sides or apex. The 
species is of rather elegant form owing to its convexity, and the rather rounded sides of 
the elytra. The chief variation is seen in the puncturation especially of the prothorax, 
which in some examples except at the extreme base is excessively smooth and shining 
and nearly impunctate, but we have taken examples distinctly punctured in company 
with the very finely punctured individuals. The length varies from r5 — 2*5 mm. 

Hab. In the mountains of all the islands. The van in which the trans-sutural 
black spot on the elytra is wanting has occurred on Kauai and Hawaii, and these 
individuals are above the average in size. 

(7) Cis bimaailatus Sharp. 

Cis bimaculatus Sharp, Tr. Dublin Soc. 1885, p. 161. 

The largest Hawaiian species of the genus, but very variable in size, some examples, 
both from Hawaii and Maui, being only about one-third the bulk of the larger indi- 
viduals. These dwarf specimens are taken in company with those of normal size. 
Otherwise this species exhibits little or no variation. There are no long hairs on the 
elytra, but excessively short pale setae are present, at least in fresh examples. Length 
2"2 — 3"2 mm. 

Hab. Maui, Haleakala.— Hawaii, at elevations of 3000—5000 ft. Not rare, 
about 30 examples have been examined. 

(8) Cis nigrofasciatus Blackb. 

Cis nigrofasciatus Blackburn, Tr. Dublin Soc. 1885, p. 162. 

(Plate X. fig. 18.) 

Seven examples of this species were taken and they exhibit no noteworthy varia- 
tion, except that some are considerably larger than others. The setae of the elytra 
are excessively minute, and hardly visible even under a very strong lens. Length 
2 — 2'7 mm. 

Hab. Lanai, 2000—3000 ft. Blackburn's unique example was taken on the 
same island. 



258 FAUNA HAWAIIENSIS 

(9) Cis unicus, sp. nov. 

Testaceus, nitidus, haud setosus, capite infuscato, elytris in parte basali lateraliter 
nigro-fuscis. Pronotum nitidum, subtilius vix dense punctatum. Elytra nitida, grossius 
irregulariter ruguloso-punctata. Long. 2 mm. 

Closely allied to C. nigrofasciatus, but smaller than the smallest example of that 
species, and decidedly more shining. The puncturation of the elytra is coarse but 
shallow, somewhat rugose and irregular, the punctures appearing to differ in size. The 
colour of the elytra is testaceous, but the basal half is to a large extent blackish or 
infuscate, except along the suture, and the dark colour at a point rather beyond the 
middle of the length of the wing-cases extends inwards nearly to the suture, to form a 
vague interrupted band. I can detect no trace of setae either on the thorax or the 
elytra. 

Hab. Hawaii. A single example was taken at Kilauea. 

(10) Cis porcatus Sharp. 

Cis porcatus Sharp, Tr. Ent. Soc. London, 1S79, p. 92. 

This minute species is easily recognized by its dark colour, very dense sculpture 
and short grey clothing, which however is easily abraded. The head of the cT is pro- 
duced into strong angular processes, which vary greatly in development. 

Hab. One of the commonest of the island species and of general distribution in 
the forests from 1 500 to 4000 ft. 

(11) Cis instilaris Sharp. 

Cis insiilaris Sharp, Tr. Dublin Soc. 1885, p. 164. 

(Plate X. fig. 19.) 

A unicolorous, or nearly unlcolorous, testaceous species, with strong and very 
dense puncturation over the whole surface, the elytra and also the prothorax in 
fresh examples bearing excessively short setae. The thorax and base of elytra are 
sometimes infuscate, and in some the elytra are quite pallid, but there are no distinct 
markings. The c/ has the front of the head produced into two processes, which, as 
usual, vary greatly in development. Length 2 — 27 mm. 

Hab. Kauai, Oahu, Lanai, and Hawaii. Taken on most of the islands rather 
sparingly, and it no doubt occurs on all. It is found in the forests from 2000 — 4000 ft. 
About a score of examples were captured. 



COLEOPTERA 259 



(12) Cis ckioj'oiiciis Sharp. 

Cis chloroticus Sharp, Tr. Dublin Soc. 1S85, p. 164. 

This is another normally immaculate species of a testaceous colour, the thorax and 
base of elytra sometimes more or less inclined to fuscous. The surface in this species 
is not quite glabrous, as in a profile view of the prothora.x some short, fine, erect hairs 
can easily be detected, although they are sparsely distributed. Length V] — 2"25 mm. 

Hab. Maui, Haleakala (4000 — 5000 ft.) ; not common. 



(13) Cis simulator sp. nov. 

Elongatus, subconvexus, nitidus, testaceus, capite nigricante, pronoto saepe plus 
minusve infuscato, elytris utrinque nigro-maculatis, apices versus lateraliter setis graci- 
libus elongatis sparsim vestitis. Pronotum subremote distincte punctatum. Elytra 
basim versus obscure punctata, puncturatione subobsoleta. Long. 175 — 22 mm. 

In the condition of its clothing this species resembles the following {Cis iabidus), the 
head and prothorax bearing fine, erect setae, very short on the latter, and easily abraded, 
the elytra at the sides towards the apex having a sparse clothing of long, fine hairs. It 
differs from that species in its larger average size, and more convex form, and the elytra 
are more pointed behind, being decidedly less parallel-sided. The insect would appear 
to be very constant in its general appearance, being testaceous in colour, with dark 
head, and two black spots at about the middle of the length of the elytra. In some 
examples the thorax is more or less obscured with fuscous or blackish clouding. At 
the base the elytra have an obscure, indefinite, and subobsolete, but rather large 
puncturation. In general appearance and form this species is extremely similar to 
C. chloroticus, but that insect has immaculate elytra, and is devoid of the long hairs at 
the sides of the elytra towards the apex. 

Hab. Maui, Haleakala (5000 ft.) ; about a dozen examples taken. 



(14) Cis tabidns Sharp. 

Cis tabidns Sharp, Tr. Ent. Soc. London, 1879, p. 93. 

Distinct from Cis bicolor, under which species it was sunk by Blackburn. It is 
very variable in size, colour &c. and also in the puncturation of the prothorax, which is 
much closer in some than others. In fresh examples the elytra bear some long fine 
hairs at the sides towards the apex, and even in some specimens all along the sides, as 

F. H. II. 34 



26o FAUNA HAWAIIENSIS 

well as some shorter erect ones on the thorax, often abraded. The prothorax varies in 
colour from black to testaceous, and the elytra are sometimes nearly wholly infuscate, 
rarely entirely testaceous and immaculate. Cis tabidus appears to be most closely allied 
to the even more variable C. setariiis, and although the colour, prothoracic puncturation 
&c., are normally different, still it is by no means easy to know to which of the two species 
certain examples should be assigned. The shape of the prothorax, length and convexity 
of the elytra &c. appear to vary analogously in either species. 

Hab. Oahu and Kauai. Common. Most and probably all of the islands, 

(15) Cis set arms Sharp. 

Cis setarius Sharp, Tr. Dublin Soc. 1885, p. 162. 

Cis apicalis Shp. loc. cit. ~ var. 

Cis concolor Shp. loc. cit. p. 163 = var. 

A most variable and perplexing species allied to the preceding but darker in 
colour, often entirely black or castaneous, generally with the anterior or both this and 
the posterior margin of the prothorax pale, the base of the elytra dark but more or less 
of the apex often pale, sometimes more than the apical half of a testaceous colour. 
Fine long hairs can generally be detected at the apex of the wing cases, sometimes also 
they form a regular but not dense clothing all along the sides, and hairs of a similar 
character can be detected standing erect on the prothorax. This part is usually 
shining, strongly but not densely punctured, and the elytra have a large and more 
or less indefinite puncturation at the base. In some examples the puncturation of 
the prothorax is extremely fine, and it appears that the species is more or less different 
according to the locality, but varies much in each. The length of the elytra and their 
convexity as well as the width of the prothorax are also variable. Length \-2 — 2 mm. 

Hab. Abundant on Hawaii in the mountains, and also found on the other islands 
including Kauai, so that it is of general distribution. 

(16) Cis caiidtis Sharp. 

Cis calidus Sharp, Tr. Dublin Soc. 1885, p. 164. 

Apparently a rather variable species, the puncturation being denser and more 
regular both on the prothorax and elytra in some examples than in others. It also 
varies in size and colour, the latter being sometimes quite black and sometimes 
castaneous. The dense regular clothing of fine erect setae is easily abraded and then 
the species is not very easily distinguished from some well clothed individuals of the 
very variable C. setarius. Length i'^ — -2 mm. 

Hab. Kauai and Oahu. A few examples taken in the mountains. 



COLEOPTERA 261 

(17) Cis signatus Sharp. 

Cis signattis Sharp, Tr. Ent. Soc. Lond. 1879, p. 92. 

Cis attenuahis Shp., Tr. Dublin Soc. 1885, p. i65 = var. of C. signatus. 

(Plate X. figs. 20, 20 «, 20 b and 20 r.) 

Extremely variable in colour. The palest examples are nearly entirely testaceous, 
having only two or four dark spots on the elytra, one example indeed apparently 
referable to this species is entirely immaculate. The darkest individuals are black with 
only the anterior and posterior margins of the prothorax, and two spots near the apex 
and one on the suture of the elytra in front of these, testaceous, so that probably entirely 
black examples of the species may occur. The prothorax is not infrequendy of an 
entirely testaceous colour, and is much wider in some examples than in others. Cis 
attenuatus Shp. is no doubt a variety of this species. The puncturation of the elytra is 
a little stronger in some examples than is usually the case. Length vix \'^ — 2 '25 mm. 

Hab. Taken on all the islands, commonly on Maui, Lanai and Hawaii, from 
1500 — 5000 ft. in the mountains. 

(18) Cis 7-oridns Sharp. 

Cis roi-idus Sharp, Tr. Dublin Soc. 1885, p. 165. 

Apparently most nearly allied to C. signatus but differing in colour &c. and 
with a stronger sculpture. It varies considerably in size, but not very greatly in other 
respects. Length i "6 — 2*2 mm. 

Hab. Most, and probably all, of the islands. A series was obtained in Kona, 
Hawaii (4000 — -5000 ft.), and we have taken it on Kauai, Molokai and Lanai. 

(19) Cis kaiiaiensis, sp. nov. 

Testaceus, capite nigro, elytris maculis compluribus plus minusve conjunctis 
variegatis, setis nullis vestitus, minus fortiter convexus. Pronotum rufo-testaceum, 
densissime punctatum, postice evidenter utrinque subimpressum. Elytra dense fortiter- 
que rugoso-punctata, macula nigra humerali, aliisque compluribus ad media plus minusve 
conjunctis, ornata. subnitida et nuda. Long. 2 "6 mm. 

A very distinct species somewhat allied to C. signatus, but more convex, with 
different markings, much stronger and coarser puncturation, and apparently with no 
trace of setae on the elytra or prothorax. Curiously enough the peculiar elytral 
markings are almost, if not quite, identical with those of some examples of Apterocis 
variegatus and A. ornatipennis. The scutellum is very distinct, and the wings are fully 
developed. The prothorax is very finely, but distinctly, margined at the sides, but not 
at all posteriorly. The surface of the elytra is distinctly shining between the dense and 
somewhat coarse punctures. 

Hab. Kauai. A single example was taken on the high plateau (4000 ft.). 

34—2 



262 FAUNA HAWAIIENSIS 



(20) Cis molokaiensis, sp. nov. 

Niger, brevissime, sed conspicue, griseo-setulosus, pronoto antice posticeque plus 
minusve testaceo-signato. Pronotiim latum, lateribus fortiter rotundatis, postice baud 
evidenter marginatum, densissime ruguloso-punctatum. Elytra densissime ruguloso- 
punctata, punctis ipsis vix discernendis. Metasternum breve. Abdominis segmenta 
ventralia punctata. Alae nullae (an rudimentariae ?). Long, i "4 — 2 mm. 

This and the following four or five species will probably hereafter be separated 
generically from the other Hawaiian Cis. As it would appear that Cis signaitcs in 
certain varieties makes an approach in form to these species, we have not thought it 
advisable to form a new genus at present, until the question of variation in that species 
has been thoroughly investigated. Cis molokaiensis on account of its dark colour and 
grey setulosity somewhat resembles C. po7'cat7is Shp., but it is abundantly distinct 
therefrom by its stronger sculpture, wider prothorax (which, however, varies in width in 
different specimens), and more elegant form, as well as by the shorter metasternum and 
the fact that the wings, if present at all, are rudimentary. There is a faint development 
of tubercles on the head of the ^, but they are very small compared with those of well- 
developed C. porcatus. 

Hab. Molokai. High wet forests, Sept. 1893. 

(21) Cis haleakalae, sp. nov. 

Niger, pronoti margine antico, pedibus, antennarumque basi, testaceis, praecedenti 
affinis sed major et latior, baud evidenter setulosus. Pronotum subtiliter densissime 
ruguloso-punctatum. Elytra dense ubique rugulosa, punctis ipsis baud vel vix dis- 
cernendis, lateribus sat rotundatis, apicibus subacuminatis. Long. 2 mm. (Plate X. 

fig- 1 7-) 

Somewhat of the form of the preceding, but with the elytra more rounded at the 
sides, and more pointed behind. The sculpture of the elytra is very remarkable, being 
of a finely strigose character. There is little doubt the species is wingless, although we 
have not been able to investigate this and other points of structure. 

Hab. Maui. A single example on Haleakala (5000 ft). 



(22) Cis mirabilis, sp. nov. 

Niger, ore, antennarum basi, pronoti margine antico maculaque postica, elytrorum- 
que fascia subapicali testaceis. Pronotum fortiter densissime punctatum. Elytra 
convexiuscula, fortiter rugosa, et punctata, baud setosa. Long. 2 mm. 



COLEOPTERA 263 

A very remarkable species of the same curious form as C. haleakalae, and no doubt 
either wingless or with only rudimentary wings, though it has not been examined in 
this respect. The coarse rugose sculpture of the elytra, amongst which distinct 
puncturation is evident, distinguish it at once. 

Hab. Kauai, 4000 ft. One example captured. 

(23) Cis fallax, sp. no v. 

Testaceus, pronoto saepe medio nigricante vel fusco, elytris nigro-maculatis, 
C. signato colore similis, et eodem vestitu et puncturatione, sed forma magis convexa, 
elytrorum lateribus magis rotundatis, alls brevioribus, ad apices elytrorum baud extensis, 
distinguendus. Long. i'5 — vix 2 mm. 

I have found it necessary to separate a few examples taken in company with 
C. signatus from that species, although they do not differ from certain of its varieties 
either in colour, clothing or sculpture. They are decidedly more convex than C. signatus 
and the sides of the elytra are more rounded, so that they often appear rather shorter 
and more pointed at the apex. On dissecting one of these specimens we find the wings 
to be less developed than those of the preceding species, and in repose they do not 
nearly reach to the apex of the abdomen, as is the case in the numerous examples 
of C. signatus that we have examined. For these reasons it is advisable for the present 
to consider the two forms as distinct species. 

Hab. Oahu. Three or four examples have been taken in the Waianae mountains 
in company with C. signatus. 

(24) Cis miiJius, sp. nov. 

Nigricans, pedibus, antennarum basi, pronoti margine antico et postico (vel horum 
altero), elytrisque testaceis, his nigro-signatis ; colore variabilis, nonnunquam totus 
testaceus, elytris nigro-notatis ; C. signato vestitu et colore simulans, setulis brevis- 
simis crebre ubique vestitus. Pronotum densissime punctulatum. Elytra subinae- 
qualia, basim versus grossius rugoso-punctata. Long, i '5 — 2 mm. 

Like many specimens of C. signatus in colour and setulosity, but easily distinguished 
by the coarse but shallow punctures on the elytra, the surface of which is somewhat 
rugose and uneven, and the insect is more convex. The wings are apparently not 
fully developed as compared with the ordinary condition of these organs in the genus, 
but we have not been able to fully examine the species with regard to this point. There 
are normally four black spots on the dorsum of the elytra, which may be connected 
more or less laterally. 

Hab. Maui, Haleakala (5000 ft.) ; three examples taken. 



264 FAUNA HAWAIIENSIS 



(25) Cis diminutivus Sharp. 

Cis dimimitivtts Sharp, Tr. Ent. Soc. London, 1879, p. 94. 

Hab. Oahu. Two examples were taken on Konahuanui by Mr Blackburn ; we 
have not met with this species, which appears to be quite distinct. 



(26) Cis evanescens Sharp. 

Cis evanescens Sharp, Tr. Ent. Soc. London, 1S79, p. 95. 

This minute species is easily distinguished from any but the following species by 
the very feeble and fine, often hardly perceptible, puncturation of the prothorax. In 
some examples the prothoracic punctures though sparse and feeble are distinguishable, 
in others the surface is practically impunctate, and the surface usually dull. The elytral 
sculpture is also variable, but always feeble, and consists of a rugulosity of the surface 
rather than puncturation, and is sometimes hardly perceptible. In some examples the 
thorax is of a testaceous colour as is usual in the following species. The entire insect 
is without clothing. Length i — 1*5 mm. 

Hab. Found on all the islands in the mountain forests from 2000 — 4000 ft. 



(27) Cis laeticulus Sharp. 

Cis laeticiihts Sharp, Tr. Ent. Soc. London, 1879, p. 94. 

Resembles C. evanescens for the most part in sculpture, but the elytra are, at least 
in most examples, decidedly more elongate, and the hinder angles of the prothorax are 
very slightly prominent and acute. This point of distinction between the two species 
is generally difficult to appreciate without separating the elytra from the thorax, but 
appears to be quite constant. In typical examples the prothorax is of a pallid colour 
and contrasts greatly with the elytra, which also are sometimes pale on their apical 
portion. Examples from Lanai probably referable to the same species have the elytra 
and thorax concolorous, or at least the latter is not much paler than the former. One 
of these Lanai examples, however, hardly differs from the typical examples, although 
the individuals were all taken in company. Length 12 — v^ mm. 

Hab. Oahu and Lanai. Probably on all the islands but not collected on the 
others. If I remember rightly the species is found on withered leaves of Freycinetia, 
where it no doubt feeds on some species of fungus. 



COLEOPTERA 265 

(28) Cis longipennis Blackb. 

Cis longipennis Blackburn, Tr. Dublin Soc. 1885, p. 162. 

Hab. Kauai. ' A single specimen was found in dry wood on the mountains.' 
(Blackburn.) We have not met with this species, which must be very closely allied to 
the following. 

(29) Cis angustiformis, sp. nov. 

Angustus, elongatus, parum convexus, fusco-niger ad colorem testaceum varians, 
pronoto nonnunquam testaceo, C. laeticulo cognatissimus, sed elytris sat evidenter 
longioribus distinguendus. Long. r2— i"5mm. 

Very closely allied to C. laetiadus, with which it agrees in the obsolete, or nearly 
obsolete, sculpture, and in having the hinder prothoracic angles a little acute and 
prominent. Darker examples are concolorous, except that the apices of the elytra are 
generally a little paler ; the paler individuals have the elytra of a fusco-testaceous 
colour, and the prothorax rather clear testaceous, so that there is some contrast between 
the colour of these parts, but it is not so striking as in typical C. laetiadus, while the 
darker specimens resemble the Lanai form of this species. It cannot however be 
treated as a form of C. laeticuhis, as the elytra are decidedly longer in proportion to 
their width, indeed the great elongation of the insect is quite remarkable. 

Hab. Oahu. Seven examples were taken in company in theWaianae mountains, 
at an elevation of 3000 ft. 

Apterocis, gen. nov. 

Forma ovali, fortissime transversim et longitudinaliter convexa, metasterno parum 
elongate, alls nullis. 

Allied to Cis with the antennae &c. similarly formed, but of oval shape, and 
extremely convex longitudinally and transversely. The species are wingless and the 
metasternum is shorter than in Cis. There are no processes on the front of the head 
of the ^ in any of the species known, but this sex may be known, in many of the 
species at least, by the presence of a tubercle on the basal abdominal segment 
beneath. 

(i) Apterocis ephistemoides Sharp. 

Cis ephistemoides Sharp, Tr. Dublin Soc. 1S85, p. 165, Plate IV. fig. 21. 
This species varies greatly in size and in some examples a few long hairs can be 
detected at the sides of the elytra. Length i — 2 mm. 
Hab. Common generally in the mountains. 



266 FAUNA HAWAIIENSIS 

(2) Apterocis vagepunctatus Blackb. 

Cis vagepundatiis Blackburn, loc. cit. p. 166. 

Hab. Oahu. A single specimen in the mountains near Honolulu (Blackburn) ; I 
have not met with this species. 

* 

(3) Aptej'ocis lanaiensis, sp. nov. 

Nigricans, pronoto nonnunquam plus minusve piceo vel rufescente, elytrorum apice 
humerisque rufo-maculatis, antennarum basi pedibusque rufescentibus. Nitidus, valde 
aequaliter convexus, pronoto ruguloso-punctato, postice baud evidenter marginato. 
Elytra circa basim sparsim vage punctata. Abdominis segmenta ventralia evidenter 
densius punctata. Long, circa 2 mm. (Plate X. fig. 14.) 

A very distinct species by the rugulosely punctured prothorax, vague scattered 
punctures at the base of the elytra and the distinct humeral and apical red spots on the 
latter. The general form of the insect is like that of Cis cphistemoides. 

Hab. Lanai. Three examples were taken on the summit of the mountains. 

(4) Apterocis montamts, sp. nov. 

Brunneus, valde aequaliter convexus, capite pronotoque nigricantibus, hoc antice 
posticeque pallido-marginato, antennis testaceis. Pronotum remote subtiliter sub- 
obsolete punctatum, postice levissime marginatum. Elytra basim versus obsoletissime 
punctata, vage subrugulosa, setulis brevissimis pallidis, vi.x videndis, ferentia. Ab- 
dominis segmenta ventralia necnon metasternum nitida et impunctata. Long, circa 
2"2 mm. 

Apterocis nioiitanus var. minor, var. nov. 

Minor, pronoto fere impunctato, elytris plaga vaga transversa obscuriore. Long, 
circa 1*5 mm. 

Much smaller than the typical specimens, but obtained in company with these. 
They differ in the nearly or quite impunctate prothorax, and the elytra have the base 
and apex paler than the middle portion, giving them a vague banded appearance. I 
can detect both sexes in these small examples and it is possible that they are really a 
distinct species. 

Two or three examples from Molokai appear to belong to this species and in some 
respects unite the typical examples with the var. minor by the following characters : 



COLEOPTERA 267 

Brown, the prothorax with some very fine remote subobsolete punctures, the base of 
the elytra with some very feebly impressed nearly obsolete but rather large ones. The 
apical portion of the elytra is of a pale testaceous colour and much paler than the brown 
basal portion much as in the var. minor. 

Hab. Maui, Haleakala, 5000ft., typical form and the var. minor; also on Molokai 
above 4000 ft. Rare. 

(5) Apterocis variabilis, sp. nov. 

Nitidus, valde aequaliter convexus, colore variabili, sed elytrorum apice extremo 
semper pallido. Pronoto distincte subtiliter nee dense punctato, postice haud distincte 
marginato. Elytra basim versus remote punctata, punctis saepe obsoletis, vix videndis, 
nonnunquam sat distinctis sed parum profundis. Abdominis segmenta ventralia dense 
subtiliter punctata. Long. r5 — 2 mm. 

Variable in colour, like A. ephistemoides in form, but of larger average size. Black 
with the thorax more or less piceous, the apex of the elytra pale and with a reddish spot 
at either shoulder sometimes united to form a transverse fascia, or entirely reddish-brown, 
with only the apex of the elytra paler than the general colour, the elytra being a little 
darker in colour in front of the pale apex, thereby rendering the paleness more distinct. 
Intermediates occur between the extreme forms. The dark varieties (.^ males) have 
the thorax more closely punctured, and the puncturation of the basal part of the elytra 
generally rather large and distinct, although shallow. In the pale forms (? females) the 
elytral puncturation is often hardly observable, but there is evident variation in this 
respect. The mesosternum in front of the middle coxae is I believe formed as in 
A. ephistemoides, so that the front pair are almost contiguous with these. 

Hab. Oahu, Waianae mountains, several examples taken. In some varieties this 
species greatly resembles A. lanaiensis but the puncturation of the prothorax is not 
rugulose as in that species, others resemble A. montanus, which has the thorax much less 
distinctly punctate. 



(6) Apterocis liawaiiensis, sp. nov. 

Niger, nitidus, valde fere aequaliter convexus, elytrorum apice rufescente. Pro- 
notum nitidum, subtiliter remote punctatum, postice haud evidenter marginatum. 
Elytra nitida, basali parte praecipue versus latera grossius remote punctata. Meta- 
sternum et abdominis segmenta ventralia dense punctata ; segmentum basale t tuberculo 
forti munitum. Mesosternum breve, parte sculpturata praecoxali fortiter obliqua. 
Long, circa 2 mm. 

F. H. II. 35 



268 FAUNA HAWAIIENSIS 

The half-dozen examples of this species show no noteworthy variation in colour, 
but some have the puncturation of the thorax less fine, and the punctures at the base of 
the elytra more numerous and coarser. I suspect these differences are for the most 
part sexual. Very short pale and inconspicuous setae can be detected in fresh 
examples. 

Hab. Hawaii (4000 ft.). 

(7) Apterocis ru/o-noiaiiis, sp. nov. 

Niger, nitidus, prothorace nonnunquam rufo-piceo, elytrorum apice rufescente, 
humerisque maculis rufis, saepe in fasciam transversam conjunctis, signatis. Pronotum 
fortiter minus sparsim punctatum. Elytra basim versus grosse remote punctata. 
Mesosternum dense sculpturatum, parte antica vix obliqua. Abdominis segmenta 
ventralia dense subtiliter punctata. Van elytris nigris, baud rufo-notatis. Long. 
I "5 — 2 mm. 

Found in company with A. hazvaiiensis, but distinguished by the general form, 
the prothorax and elytra not forming together such an even and continuous curve. 
The mesosternum in front of the coxae is so little oblique that it nearly continues 
the plane of the metasternum. Some examples have the thorax decidedly more 
finely and remotely punctured than others, the variation being analogous with that 
exhibited in the same parts of A. hawaiiensis. 

Hab. Hawaii (4000 ft.). Six examples taken. — Molokai (4500 ft.). One 
mutilated example taken, apparently identical with those from Hawaii. 

(8) Apterocis impunctattis, sp. nov. 

Nigricans vel nigro-piceus, antennis pedibusque rufo-testaceis. Pronotum im- 
punctatum, nitidum, postice distincte marginatum. Elytra nitida, impunctata, corpore 
subtus impunctato. Long, i — 1"5 mm. 

In many respects very like A. ephistemoides but readily distinguished by its form, 
the outline of the thorax and elytra not forming the very even curve that is seen 
in that species. Posteriorly the prothorax is finely but very distinctly margined. 

Hab. Oahu, Waianae mountains. — Kauai, Makaweli (2500 ft.) one very minute 
individual. 

(9) Apterocis variegates, sp. nov. 

Niger, sat elongatus, ovalis, glaber, elytris testaceis, nigro-variegatis, pronoti 
margine antico maculaque basali, antennarumque articulis basalibus, testaceis. Pronotum 
densissime grosseque rugoso-punctatum, subimpressum et subinaequale, postice vix 



COLEOPTERA 269 

marginatum. Elytra grosse rugoso-punctata, basim versus sulcis longitudinalibus com- 
pluribus, male definitis, inaequalia, testacea, maculis nigris plus minusve conjunctis 
eleganter variegata. Metasternum fortiter punctatum. Abdominis segmenta 2 — 5 
dense subtilius punctata. Long. 2 '5 mm. 

Remarkable for its fusiform-ovate shape, and the excessively dense coarse sculpture. 
The metasternum is not so much shortened as is usual in the genus, and the mesosternum 
in front of the coxae is rather long, horizontal, and densely sculptured except along 
the narrow articulating margin. No doubt this insect and others allied to it will here- 
after be separated from typical Apterocis, which are much more strongly convex and 
differ in the form of the mesosternum, but if so it will be necessary to form several new 
genera for this remarkable group of insects. 

Hab. Maui, Haleakala (4000 — 5000 ft.). Rare, four examples only having been 
met with. 



(10) Apterocis ornatipennis, sp. nov. 

Forma, magnitudine, et colore praecedentis, sed pronoto minus dense rugoso- 
punctato, punctis ipsis facile distinguendis, distinctus. Long. 2 — 2 '5 mm. (Plate X. 

% 1 5-) 

Very closely allied to A. variegatits, and only differing in the sculpture of the 
prothorax, which is less extremely dense, and though the puncturation is very rugose 
the individual punctures are moderately distinct, and do not run into one another to the 
extent that is seen in the other species. The pale colour of the margins of the 
prothorax is also of greater extent, and in one example the prothorax is entirely piceous 
in colour. 

Hab. Lanai (2000 — 3000 ft.) ; very rare, two examples only having been met 
with. 



(11) Apterocis sfrigosiis, sp. nov. 

Castaneus, rufo-brunneus, vel testaceus (an immaturus ?), elytris prothorace palli- 
dioribus, latissime nigro-fasciatis, maculisque duabus nigris ad basim plerumque ornatis. 
Pronotum fortiter punctatum et longitudinaliter strigosum, nitidum, postice haud evi- 
denter marginatum, margine antico pallido, et ad angulos anteriores flavo-maculatum. 
Elytra fortiter convexa, ad basim latiuscula, apices versus attenuata, basi grosse rugoso- 
punctata, setas pallidas brevissimas inconspicuas sparsim ferentia. Metasternum sat 
breve, fortiter punctatum. Abdominis segmenta 4 ultima distincte subtiliter punctata. 
Long. 17 — 2 '2 mm. (Plate X. fig. 16.) 

In this species the markings on the elytra are variable, the two spots at the basal 



2 70 FAUNA HAWAIIENSIS 

margin being sometimes connected with the wide transverse fascia, but sometimes are 
altogether wanting. The extreme apex of the elytra is usually dark. The meso- 
sternum is prolonged subhorizontally in front of the coxae, almost continuing the plane 
of the metasternum. It is coarsely sculptured except for the very narrow front margin 
which is overlapped by the prosternum. 

Hab. Molokai. Highest forests above 4000 ft. 



(12) Apterocis subaeneus, sp. nov. 

Praecedenti cognatissimus, sed forma paullo angustiore, prothorace nigro, subaeneo- 
micante, ad angulos anteriores haud flavo-maculato, distinguendus. Long, vix 2 mm. 

Extremely like A. strigosus but a narrower insect with black slightly aeneous 
prothorax, which is not maculate at the front angles. The pale setae are observable on 
the thorax, and on the elytra they are decidedly less inconspicuous than is the case in 
the preceding species. A series of examples may, however, prove this to be only a 
variety of A. strigosus. 

Hab. Maui. A single example taken on Haleakala at an elevation of 5000 ft. 



(13) Apterocis Iiystrix, sp. nov. 

Unicolor, testaceus, aut capite nigricante, ovalis, sat convexus, setis erectis elon- 
gatis dense ubique vestitus. Pronotum latum, postice haud marginatum, densissime 
subtiliter punctatum. Elytra dense irregulariter rugoso-punctata. Long, circa 2 mm. 

Unlike any other species, in form something like A. strigosus and its allies, the 
convexity of the elytra and prothorax not forming a very strong and even curve, when 
viewed laterally. The elytral puncturation is dense, rugulose and indefinite, the punc- 
tures apparently not being of even size, and but shallowly impressed. 

Hab. Lanai. Two examples (one mutilated) were taken on the top of the 
mountains. 



ZOOLOG-ICAL RESULTS based on material from New Britain, New Guinea, Loyalty 
Islands and elsewhere, collected during the years 1895, 1896, and 1897, by 
Arthur Willey, D.Sc. Lond., Hon. M.A. Cantab., Late Balfour Student of the 
University of Cambridge. Demy 4to. Parts I, II, and III. Price 12s. 6al. each. 

PART I. 

1. The anatomy and development of Peripatus novae-britanniae. By ARTHUR WiLLEY, M.A., D.Sc. 

With Plates I. — IV. and 7 figures in the text. 

2. Metaprotella sandalensis, n. sp. [Caprellidae]. By Dr PAUL MAYER. 

With 6 figures in the text. 

3. On a little-known Sea-snake from the South Pacific. By G. A. BoULENGER, F.R.S. 

With Plate V. 

4. Report on the Centipedes and Millipedes. By R. I. POCOCK. 

With Plate VI. 

5. Account of the Phasmidae with notes on the eggs. By D. SHARP, M.A., F.R.S. 

With Plates VII.— IX. 

6. Scorpions, Pedipalpi and Spiders. By R. L PocoCK. 

With Plates X.— XI. 

PART 11. 

7. Report on the specimens of the genus Millepora. By Sydney J. HiCKSON, M.A., D.Sc, F.R.S. 

With Plates XII.— XVI. 

8. Report on the Echinoderms (other than Holothurians). By F. JEFFREY Bell, M.A. 

With figures on Plate XVII. and one figure in the text. 

9. Holothurians. By F. P. BEDFORD, B.A. 

With figures on Plate XVII. 

10. Report on the Sipunculoidea. By ARTHUR E. SHIPLEY, M.A. 

With Plate XVIII. 

11. On the Solitary Corals. By J. STANLEY GARDINER, M.A. 

With figures on Plates XIX. and XX. 

12. On the postembryonic development of Cycloseris. By J. STANLEY GARDINER, M.A. 

With figures on Plates XIX. and XX. 

13. On a collection of Earthworms. By Frank E. Beddard, M.A., F.R.S. 

With Plate XXI. 

14. The Gorgonacea. By ISA L. HiLES, B.Sc. 

With Plates XXII. and XXIII. 

PART III. 

15. Orthogenetic variation in the shells of Chelonia. By HANS Gadow, M.A., Ph.D., F.R.S. 

With Plates XXIV.— XXV. and one text-figure. 

16. Enteropneusta from the South Pacific, with notes on the West Indian Species. By ARTHUR 

WiLLEY, D.Sc, Hon. M.A. Cantab. 

With Plates XXVI.— XXXII. and 7 text-figures. 

17. On a collection of Echiurids from the Loyalty Islands, New Britain and China Straits, with an attempt 

to revise the group and to determine its geographical range. By ARTHUR E. SHIPLEY, M.A. 
With Plate XXXIII. 

PART IV. \In the Press:] 

18. On the anatomy of a supposed new species of Coeitopsammia from Lifu. By J. STANLEY 

Gardiner, M.A., Fellow of Gonville and Caius College, Cambridge. With one plate. 

19. On the Insects from New Britain. By D. Sharp, M.A., M.B., F.R.S. With one plate. 

20. Report on the Stomatopoda and Macrura. By L. A. BORRADAILE, M.A., Lecturer of Selwyn 

College, Cambridge. With four plates. 

21. Report on the Slugs. By Walter E. Collinge, F.Z.S., Mason College, Birmingham. 

With two plates. 

22. Report on the Polyzoa. By E. G. Philipps, Newnham College, Cambridge. With two plates. 

23. The Hydroid Zoophytes. By LAURA ROSCOE Thornely, University College, Liverpool. 

With one plate. 

24. Astrosclera willeyana, the Type of a new Family of Calcareous Sponges. By J. J. LiSTER, M.A., 

St John's College, Cambridge. With five plates. 

25. A contribution towards our knowledge of the Pterylography of the Megapodii. By W. P. 

Pycraft, A.L.S., British Museum. With one plate. 

26. The Stolonifera and Alcyonacea. By Sydney J. HiCKSON, M.A., D.Sc, F.R.S., Beyer Professor 

of Zoology in the Owens College, and ISA L. HiLES, B.Sc, Owens College, Manchester. 
With two plates. 

27. Report on the Xeniidae. By J. H. ASHWORTH, B.Sc, Demonstrator in Zoology, Owens College, 

Manchester. With two plates. 

iContron: C. J. CLAY and SONS, 

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The Lord Avebury, F.R.S 
F. E. Beddard, F.R.S. ... 

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Prof. AUGUSTE FOREL 

p. H. Grimshaw, F.E.S. 

G. W. KiRKALDY, F.E.S. 

E. Meyrick, F.Z.S. 
R. C. L. Perkins, B.A. 

R. I. POCOCK 

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R.S. 



Hymetioptera Parasitica. 

Thysanuva and Collembola. 

Earthworms. 

Isopod Cntstacea. 

Formicidae. 

Diptera. 

Hemiptera. 

Macrolepidoptera. 

[Hymenoptera Aculeata, Orthoptera, 
\ Neuroptera, and part of Coleoptera. 

Myriapoda. 

Part of Coleoptera. 

Parasitic Worms. 

Arachnida. 

Molhisca. 

Microlepidoptera . 



It is also intended to give a list of the Vertebrates, with their distribution, 
in the Islands. 



N.B. The parts of Volumes I. and II. are being published concurrently in order 
to expedite the completion of the work. 

The price of each part will vary according to its extent and the number of Plates. 
Subscribers to the whole work will be charged half-price for each part. The parts will 
be sent, as published, to each subscriber who has paid for the preceding part. 

Those who wish to be subscribers, should send name and address either directly to 
C. J. Clay & Sons, Cambridge Warehouse, Ave Maria Lane, London, E.C., or through 
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FAUNA HAWAIIENSIS 



OR THE 



ZOOLOGY OF THE SANDWICH (HAWAHAN) ISLES 

Being Results of the Explorations instituted by the Joint Committee 

appointed by 

THE ROYAL SOCIETY OF LONDON FOR PROMOTING NATURAL KNOWLEDGE 

AND THE BRITISH ASSOCIATION FOR THE ADVANCEMENT OF SCIENCE 

And carried on with the assistance of those Bodies and of the Trustees of 

THE BERNICE PAUAHI BISHOP MUSEUM AT HONOLULU. 



EDITED BY 

DAVID SHARP, M.B., MA., F.R.S. 

SECRETARY OF THE COMMITTEE. 



VOLUME II. PART iV. 

MOLLUSCA BY E. R. SYKES : EARTHWORMS by F. E. BEDDARD 

ENTOZOA BY A. E. SHIPLEY. 

Pages 271—441; Plate XI, coloured; Plates XII, XIII, XIV, imcoloured. 



CAMBRIDGE: 

AT THE UNIVERSITY PRESS. 
1900 

[^// liiglits resereed.'] 
May 19, 1900. 



aonDon: C. J. CLAY and SONS, 
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TLeippf. F. A. BROCKHAUS. 

0cto Jgorft: THE MACMILLAN COMPANY. 

Bombao: E. SEYMOUR HALE. 



MOLLUSCA 

By E. R. SYKES, F.Z.S. 

EARTHWORMS 

By F. E. BEDDARD, F.R.S. 

ENTOZOA 

By a. E. SHIPLEY, M.A. 



271 



MOLLUSCA. 



""■ '■ s-b., 



ByE.E.Sykes, "■=;£-.;.., 

WITH INTERCALATIONS ON ANATOMY 
By Lt.-Col. Godwin- Austen. 

Contents. § i, General remarks, p. 271 ; § 2, Systematic account, p. 275 ; § 3, Biblio- 
graphic list, p. 400 ; § 4, Alphabetical list of names placed as synonyms, p. 407 ; 
§ 5, Alphabetical list of nnideiitifed or erroneously recorded names, p. 412. 

The material upon which this study of the land and fresh-water Mollusca is based, 
in addition to the collection formed by Mr Perkins, consists in a great measure of the 
collection in the British Museum (Natural History), where the bulk of Newcomb's and 
Pfeiffer's type-specimens are to be found. Thanks to the kindness of Prof. A. Hyatt, 
an examination has been made of the type-specimens of the species described by 
Gulick, of the genus Leptachatina, and now preserved at Boston. Mr D. D. Baldwin, 
of Maui, has also very kindly sent over a number of specimens and Mons. Ancey has 
lent the types of some species described by him. A collection formed in the Islands by 
Mr Hutchison has also been placed in my hands for examination by Mr Fulton. 

Lt.-Col. H. H. Godwin- Austen, F. R.S., has enabled me to add to the interest of 
this work in a great degree, by very kindly dissecting some of the species and permitting 
me to incorporate here the results ; it is hoped that he may be able to give a further 
account of the anatomy in the Ackatinellidae. 

To Mr Edgar A. Smith, I desire, in conclusion, to express my most grateful thanks 
for his unfailing help and courtesy. 

§ 1, General Remarks on the Mollusca. 

To the student of the Mollusca, the Hawaiian Islands fauna is probably more 
familiar by name than that of many better known places, owing to the occurrence there 
of the well-known Achatinelloid group of forms. 

Tables of the distribution of the fauna are given below, but a few general remarks 
here may be of interest. 

F. H. II. 36 



272 FAUNA HAWAIIENSIS 

The Limacidae yield nothing very peculiar or very striking and the few forms 
peculiar to the Islands may well have been developed from introduced European 
ancestors. 

The Zonitidae are scattered over the Islands ; all are peculiar, but they are 
nearly related to forms found in other islands of the Pacific : similar remarks apply 
to the Endodontidae, one group of which [Pterodiscits), however, appears to be peculiar. 

The presence — and that strongly contested — of only a single indigenous species 
of the Helicidae again indicates affinities with Polynesia. 

The Pupidae as a family, have a very wide geographical range, and hence no 
deductions can be drawn from their presence ; it should be noted that here — if the 
identification be correct — the fauna includes a species not peculiar to the Islands. 

With reference to the Achatinellidae it may at once be noted that several divisions 
of the family may be made. First, the brightly coloured forms which fall into the 
genus Achatinella proper and which are replaced in the Southern Pacific Islands by 
the genus Partula. The metropolis of distribution of all these forms seems to be 
Oahu, save in the case of the subgenus Partulina when Maui and Molokai appear to 
divide the honour. No species has been found on Kauai and only two on Hawaii at 
the other end of the group. Species have been described by authors upon coloration 
and band-formations ; in my opinion numbers even of the ' species ' here admitted will 
prove, when their anatomy is carefully investigated, to be varying forms of one common 
species. Consider, for example, such a shell as Tacliea nemoralis dealt with in the 
same manner as the Hawaiian forms have been ! Still, even when reductions are made, 
the fauna will remain remarkable for its numerical strength in species. 

Secondly, passing through Perdicella and Newcombia, confined to the islands of 
Molokai and Maui, we come to the second great division, typified by Leptachatina and 
Amasti'a. Here, while the metropolis again seems to be Oahu, Kauai, the oldest 
island geologically considered, ranks well with the rest. 

Thirdly, passing through the interesting and recently described Thaammiia of 
Oahu, we come to Carelia, which is confined to Kauai save for one subfossil species on 
the Island of Niihau (the only mollusc on that island). 

Fourthly, we have the little group of Aufiadella and Frickella, which leave the 
impression that they are linking forms between Achatinella and Tornatellina, and, 
again, belong in the main to Oahu. It should be borne in mind, as illustrating the 
peculiarity of the fauna, that only about half a dozen out of, approximately, 330 
species of Achatinellidae are found on more than one island, and indeed some of 
these may be due to errors of identification. In our present state of knowledge a 
faunal list is largely influenced by the ' personal equation ' of the writer. 

From the residue of the fauna but little is to be learnt ; the development of 
Succinea appears abnormal and further research will probably reduce the so-called 
' species ' of this group. 



MOLLUSC A 273 

The following general conclusions may, however, be drawn : 

1. The Molluscan fauna is nearly related to that of the Polynesian islands, and 
shows hardly any trace of continental influence, Asiatic or American. 

2. The species are nearly always confined to one island ; but it is very doubtful 
if, as has been stated, "each valley has its peculiar species." 

3. When the genera found are confined to the islands, the majority of living 
species usually occur on Oahu. 

I give below tables of distribution ; but, owing to the fauna being so restricted in 
distribution, have not added percentages of peculiar species. 

(i) Families Limacidae, Zonitidae, Endodontidae, Helicidae, Pupidae. 

Species peculiar to one Island. 

Pupidae. Total. 

1 1 

4 14 

2 

3 

8 

2 8 

Species occurring in more than one Island. 

Limacidae. Two species (Z. gagates and Agriolmiax laevis) are found elsewhere, 
and may have been introduced. One species is common to Kauai and Maui, one to 
Maui and Hawaii, and one to Kauai, Oahu, and Maui. 

Zonitidae. One species common to Kauai, Oahu, and Maui ; one to Oahu, 
Molokai, and Lanai ; and one, respectively, to Kauai and Maui, Maui and Oahu. 

Philomycidae. One found in Kauai, Oahu, and Hawaii, and one in Oahu only; 
these species, however, are not peculiar to the Hawaiian fauna. 

Endodontidae. Two species are common to Kauai and Oahu ; one, respectively, 
to Kauai and Lanai, Maui and Oahu, Lanai and Oahu, Lanai and Molokai ; while 
three are of uncertain habitat. 

Helicidae. One species — introduced — in Kauai and Oahu. 

Pupidae. One in Kauai, Oahu, and Hawaii ; one, respectively, in Kauai and 
Oahu, Oahu and Hawaii ; further, a single species is found outside the Islands. 

\6 — 2 





Limacidae. 


Zonitidae. 


Endodontidae. 


Helicidae. 


Kauai 




3 


8 




Oahu 




7 


3 




Molokai 




I 


I 




Lanai 


I 


2 






Maui 


I 


5 


2 




Hawaii 


I 


3 


I 


I 



J'- 



274 



FAUNA HAWAIIENSIS 



(2) Achatinellidae. In view of their interesting characters I have here dealt 
with the distribution by genera. 

Species occurring in only one Island. 





ACHATINELLA group. 












^ 














-a 


























c 








1 


g 


'-I 


1 


.g 


!3 


.51 


•2 

S 


.«■ 








53 




1 


1 




5 

^ 

1 






« 

S 


<5 




e2 


Kauai 














7 


15 




8 


I 


31 


Oahu 


17 


19 


I 


29 






40 


33 


I 




10 


15° 


Molokai 






9 


2 


3 


8 


IS 


3 






2 


42 


Lanai 






3 








12 


5 








20 


Maui 






16 


3 


5 


I 


17 


4 






I 


47 


Hawaii 






2 








3 


7 






I 


13 



The only forms of Achatinellidae found on more than one island are in the 
genera Leptachatina and Aiu-icidella ; in the former one species is said to be found 
on Kauai and Oahu, and two on Maui and Oahu ; in the latter similar notes occur 
with regard to Oahu and Maui, Molokai and Lanai, Maui and Molokai, and (doubt- 
fully) Oahu and Hawaii. 

Further a single subfossil species of Carelia is recorded from Niihau ; and the 
following are of uncertain habitat: Bulimella i, Partidina 3, Achatinellastriim i, 
Aniastra 7, Leptachatina 6, and Aitriatletla 2. 

(3) The residue of the fauna. 

Species occurring in only one Island. 







a 




















< 




u 




c 










i5 


Q 

5 


S 


< 


« 


'C 


I 


5 


<A 










< 
Z 


1 


1 




.■V. 







K 






J 




^ 








Kauai 


3 




3 


2 










8 


Oahu 


I 




2 


3 


I 


I 


3 




II 


Molokai 





















Lanai 


I 




I 












2 


Maui 


3 




4 


I 


I 








9 


Hawaii 


2 


I 


12 


I 










16 



MOLLUSC A 275 

Species occurring in more than one Island. 

Tornatellina. One species, said to be found in Oahu, occurs in the Tonga 
Islands. Two species are common to Kauai and Oahu, one to Hawaii and Oahu, 
and one to Kauai, Oahu, and Hawaii. 

Stenogyridae. Opeas j^mceus is said to be found in all the Islands, and both this 
and O. prestoni (Hawaii) occur elsewhere. One species is of uncertain habitat. 

Succinea. Two species in Oahu, Molokai, and Hawaii ; one in Lanai, Oahu, and 
Maui ; one in Kauai and Hawaii, one in Maui and Molokai. 

Limnaeidae. One species in 'all the Islands'; one in Kauai and Oahu, one 
in Oahu and Maui ; three of uncertain habitat. 

Melania. One common to Kauai, Oahu, Maui, and Molokai ; one to Kauai 
and Oahu ; one to Kauai and Molokai ; one of uncertain habitat. 

Helicina. One common to Kauai, Oahu, Lanai, and Molokai ; one to Maui 
and Lanai. 

Neritina. One common to Maui, Oahu, and Hawaii ; two of uncertain habitat. 
Two said to be found in 'all the Islands.' 

These three tables show that Kauai has 50 species peculiar to it, Oahu 175, 
Molokai 44, Lanai 25, Maui 64, Hawaii 37. 

§ 2. Systematic account of the fauna. 
Fam. LIMACIDAE. 

Amalia Moquin-Tandon. 

Avtalia M.-T. Hist. Moll. France, 1855, 11. p. 19 [first species Lhnax gagates, 
Drap.]. 

Milax Gray, Cat. Pulm. Brit. Mus. 1855, p. 174 [has the same type; there are, 
also, older names supposed to be identical, but founded on erroneous characters or 
improperly described]. 

While dealing with slugs it may be convenient to note that Semper has recorded a 
species stated to be very near Limax tene/lus Nilsson ; further Dr Cooper is said to 
have seen a species of Jmiella from these islands, but I have been unable to trace his 
note from the reference given (see CoUinge, P. Malac. Soc. London, 11. p. 50). 

(i) Amalia babo7'i Collinge. 
Amalia babori ZoWmgt, P. Malac. Soc. London, 11. (1897), p. 294. 
Hab. Maui, at 5000 ft., Haleakala.— Hawaii, 2000 to 4000 ft., Olaa to Kilauea 
(Perkins). 



276 FAUNA HAWAIIENSIS 

(2) Amalia gagates Draparnaud. 

Limax gagates Draparnaud, Tabl. Moll. France, 1801, p. 100 ; Hist. Moll. France, 

1805, p. 122, pi. IX. figs. I, 2. 
Amalia gagates Drap., Collinge, P. Malac. Soc. London, 11. p. 49. 
Hab. Maui (Perkins). 

Agriolimax Morch. 

Agriolimax Morch, J. Conchyl. xiii. (1865), p. 378. 

As to the correct name for this genus, see Cockerell and Collinge, Conchologist, 
II. pp. 199, 200. 

(i) Agriolhnax bevenoti Collinge. 

Agriolimax bevenoti Co\\\ng&, P. Malac. Soc. London, 11. (Nov. 1897), p. 295. 
Hab. Kauai, at 4000 ft. — Oahu, 2000 ft., Honolulu. — Maui, 5000 ft., Haleakala 
(Perkins). 

(2) Agriolimax globosus Collinge. 

Agriolimax globosus Collinge, P. Malac. Soc. London, 11. (April 1896), p. 47. 
Hab. Hawaii, Mauna Loa (Perkins). 

(3) Agriolimax laevis Muller. 

Limax laevis Muller, Hist. Vermium, 11. (1774), p. i. 

Agriolimax laevis Muller, Collinge, P. Malac. Soc. London, 11. p. 295. 

Hab. Kauai, at 2000 ft., Lihue. — Maui, 5000 ft., Haleakala (Perkins). 

(4) Agriolimax perkinsi Collinge. 

Agriolimax perkinsi Collinge, P. Malac. Soc. London, 11. (April 1896), p. 47. 
Hab. Lanai, at 2000 ft. (Perkins). 

(5) Agriolimax i^') sandivichiensis Souleyet. 

Lim/ix sandwichiensis Souleyet, Voy. Bonite, Zool. 11. (1852), p. 497, pi. xxviii. 

figs. 8 — 1 1 [animal and shell]. 
Hab. Hawaiian Islands (?). 

It seems uncertain whether this be really Hawaiian, or even accurately repre- 
sented ; see Collinge, P. Malac. Soc. London, 11. p. 46. 



MOLLUSCA 277 

Fam. ZONITIDAE. 

GoDwiNiA, n. gen. 

This new genus is proposed for the Vitrina capcrata of Gould, which has, of 
recent years, usually been placed in Helicarion ; it will be seen from the valuable 
anatomical notes of Lt.-Col. Godwin-Austen that there are differences which separate 
the species from that genus. Probably the Vitrina tenella of Gould also belongs here. 
The types oi Helicarion Ferussac (Tabl. Moll. 182 1, pp. xxxi, 24) appear to have been 
the Australian iorms freycineii and cuvieri. 

(i) Godivinia caperata Gould. 

Vitrina caperata Gould, P. Boston Soc. 11. (1847), p. 181 ; U. S. Explor. Exped. 

Moll. 1856, pi. I. fig. 9. 
Helix newcombi Pfeiffer, P. Zool. Soc. London, 1S54 (Jan. 1855), p. 51 ; Reeve, 

Conch. Icon. Helix, pi. clxxxix. fig. 132 1. 

Plate XII. figs. 6— 12. 

Hab. Kauai (Gould, Perkins). — Oahu (Pfeiffer). 

Very possibly the habitat of ' Oahu ' is a mistake. 

" The animal is dark, with a rather broad pale pallial margin ; foot with a well 
defined central area beneath ; the specimen was so much contracted that the mucous 
gland could not be decisively made out ; from analogy, however, one should be present. 
There are no shell-lobes, the mantle-edge is curved and well defined. The right dorsal 
lobe is small, and the left lobe is long, narrow, and continuous. 

" The visceral sac has three coils. The buccal mass has a strong, broad, muscle on 
the lower posterior side ; the oesophagus is short, leading into a very capacious stomach ; 
the salivary gland is in one compact, rounded mass. Jaw solid, dark sienna in colour, 
with a very straight cutting edge ; odontophore long and narrow, with a few large 
median teeth ; at first sight these centrals appear to be simple and straight-sided in form, 
and they are very nearly so, but closer examination shows that the centre and adjoining 
teeth have very small notches on the outer side ; these are not cusps. The laterals are 
all curved and aculeate. The dental formula is : 

18— 5— I— 5-18 

23—1—23 

Unfortunately the generative organs were not seen by me, all this portion being lost 
during dissection, as will sometimes occur in these small species. 



2 78 FAUNA HAWAIIENSIS 

"It will be seen from the above characters that this species cannot be placed in the 
Helicarionidae — the absence of shell-lobes forbids this. Aculeate laterals are hardly 
ever met with even in the genera of Zonitidae possessing shell-lobes ; I can only recall 
one species, Macrochlamys castaneolabiata. The solid jaw, divided foot, and, in all 
probability, the presence of a mucous gland place it in the Zonitidae. In so many 
points is it distinct from any of the Indian and Malayan forms that I am acquainted 
with that I the more regret that the generative organs have still to be made out " 
(H. H. Godwin-Austen). 



(2) Godwinia (.'') tenella Gould. 

Vitrina tenella Gould, P. Boston Soc. 11. 1847, p. 18 1 ; U. S. Explor. Exped. 
Moll. 1856, pi. I. fig. 10. 

Hab. Kauai (Gould). — Maui, Haleakala, 5000 — 9000 ft. (Perkins). 

The specimens found by Mr Perkins appear to be identical with Gould's species, 
which, so far as I can trace, has not been rediscovered on Kauai. In fresh specimens 
the lip is margined with black. 



ViTREA Fitzinger. 

Vitrea Fitz., Beitr. Landeskund. Oesterr. iii. p. 99. 

Fitzinger's type, as I understand him, was diaphana Studer. 

Until the anatomy of these Hawaiian species is known, I can suggest no better 
reference than to the present genus. 

( I ) Vitrea lanaiensis Sykes. 

Vitrea (?) lanaiensis Sykes, P. Malac. Soc. London, 11. (1897), p. 298. 

Plate XI. figs. 43, 44. 

Hab. Lanai, mountains behind Koele (Perkins). 

(2) Vitrea molokaiensis Sykes. 

Vitrea (?) molokaiensis Sykes, P. Malac. Soc. London, 11. (1897), p. 298. 

Plate XI. figs. 45, 46. 

Hab. Molokai, forest above Pelekunu (Perkins). 



MOLLUSC A 279 



(3) Vitrea pauxilliis Gould. 

Helix pnsillus Gould, P. Boston Soc. 11. 1846, p. 171 \jion H. pusilla, Lowe, 

1831]. 
Helix paiixillus Gould, U. S. Explor. Exped. Moll. p. 40, pi. in. fig. 46. 
Hyalinia baldicini Ancey, Bull. Soc. Malac. France, vi. 1889, p. 192 ; Sykes, P. 

Malac. Soc. London, in. pi. xiii. figs, i — 3. 

Hab. Maui (Gould) ; West part of Maui (Ancey) ; Haleakala, 5000 feet (Perkins). 
See, for a note on the synonymy, P. Malac. Soc. London, 11. p. 298. 



PsEUDOHVALiNA Morse. 
The original type was, I gather, Helix exigua Stimpson. 

(i) Psettdohyalina kauaiensis Pfeiffer. 

Helix kaivaiensis Pfeiffer, P. Zool. Soc. London, 1854 [1855], p. 52; Reeve, 

Conch. Icon. Helix, sp. 1256. 
Hab. Kauai (Pfeiffer). — Maui and Oahu (Baldwin). 

I follow M. Ancey in the generic reference, as I do not know how, at present, the 
nomenclature may be bettered. 

MiCROCYSTiNA Morch. 
Type Nanina rinkii, Morch. 

(i) Microcystina (?) ciyptoportica Gould. 

Helix cryptoportica Gould, P. Boston Soc. 11. (1846), p. 20; U. S. Explor. Exped. 

Moll., pi. V. fig. 72. 
Hab. Oahu (Pease, Baldwin). 
I place this here as the description states "columella valde intorta." 

Microcystis Beck. 
For a discussion as to the type, see P. Malac. Soc. London, 11. pp. 130—2. 

F. H. II. 37 



28o FAUNA HAWAIIENSIS 

(i) Microcystis cJiainissoi Pfeiffer. 

Helix chamissoi Pfeiffer, P. Zool. Soc. London, 1855, p. 91 ; Bland and Binney, 
Ann. Lye. New York, x. p. 338, pi. xv. fig. 3 {jaw and radula ; copied 
in Ann. New York Ac. in. pi. xvii. fig. O). 
Hab. Kauai, Waioli and Haena (Baldwin) ; Makawele and Mountains above 
Waimea (Perkins). 

Bland and Binney give " W. Maui " on the authority of Newcomb, but this 
seems very dubious ; Mons. Ancey gives Oahu, but I think this must be an error. The 
figures in the Maimal of Conchology (Vol. 11. pi. xxxviii. figs. 74 — 6) are not good. 



Philonesia, gen. nov. 

Recently', I discussed the genus Microcystis Beck, and expressed the opinion that 
these small Zonitoid forms so characteristic of the Hawaiian Islands, and scattered over 
the Islands of the Central Pacific, could not be placed in that genus. I, further, 
referred them to Macrochlamys, stating that " whether our small forms are in accord 
with the typical group of this genus anatomically, remains to be proved ; but, concho- 
logically, they only appear to differ in size." 

Specimens of a form which I refer to the unfigured Microcystis baldivini Ancey, 
and which were collected by Mr Perkins, contained the animal, and Lt.-Col. Godwin- 
Austen has most kindly made an examination of it. His full report will be found on 
p. 281, but I may here summarize it by saying that this species does not belong to 
Macrochlamys at all, and the query I suggested has been answered. He points out its 
affinity to Sitala and Kaliclla and here it is interesting to note that Mr Perkins found 
a species in the Hawaiian Islands that I have referred to the latter genus. 

Under these circumstances, and as the shells are distinct by the conchological 
characters of the columella from both Microcystina and Lajuprocystis — anatomically, 
also, from the former — I have ventured to create a new genus and propose to take 
Microcystis baldivini Ancey, as the type. Probably the bulk of the Hawaiian Zonitoid 
forms belong to this group. 

(i) Philonesia abeillei Ancey. 

Microcystis abeillei Kncoy, Bull. Soc. Malac. France, vi. (1889), p. 199. 
Hab. Molokai (Ancey) ; Mapulehu (Baldwin) ; wet forest above Pelekunu 
(Perkins). — Oahu, Waianae Mts. (Perkins). — Lanai (Perkins). 

All the specimens are young, but I cannot sever them from this species. 

' P. Malac. Soc. London, 11. p. 130. 



MOLLUSC A 281 



(2) Philonesia baldivini Ancey. 

Microcystis baldwini Ancey, Bull. Soc. Malac. France, vi. (1889), p. 204. 
Plate XII. figs. 1—5. 

Hab. Oahu and west part of Maui (Ancey); Head of Panoa Valley, Nuuanu, and 
Honolulu Mts. (Perkins). 

"The animal is brown ; spotted and splashed with pure white (Plate XII. fig. i a) 
on the inteofument which covers the branchial chamber and visceral sac, these markines 
shew clearly through the transparent shell and give it a very pretty, mottled ap- 
pearance. The extremity of the foot is truncated ; with a mucous gland. In the 
specimen examined the foot (Plate XII. fig. 2) is very much contracted, but there is 
every indication that a small lobe overhangs the mucous gland. The foot, which is 
regularly segmented, has a central area (Plate XII. fig. 2«); the pallial margin appears 
unusually broad, but this is deceptive and due to the extreme lateral contraction under- 
gone ; the two grooves above are similarly widened. The mantle edge has a well- 
developed, tongue-like, right shell lobe near the respiratory orifice, with an indistinct, 
narrow, left shell lobe. The right dorsal lobe is black and well developed, the left 
paler and moderately broad. Tentacles black. 

" Plainly seen through the shell were four embryonic shells, lying one behind the 
other in the uterus, in various stages of development. The enveloping integument is 
transparent and so thin that the small shells, being comparatively heavy bodies, very 
readily break away, and the spermatophore adjacent was not made out. 

"The odontophore has a formula of 

30 : 9 : I : 9 : 30 
39 : I : 39- 

" The basal plates of the central teeth are quadrate in outline. The central tooth is 
tricuspid, the side cusps basal, blunt ; the central point with convex sides. The 
median teeth have a blunt cusp only on the outer basal side, the ninth tooth is a 
narrower basal plate and is intermediate in form, the next eighteen being curved and 
bicuspid ; the most interesting character is seen at this part of the row, for all the 
succeeding and outermost teeth are tricuspid, occasionally with even four points. The 
radula is remarkable for the similarity of the outermost teeth to those of Kaiiella 
barrakpurensis" \ those of Sitala attegia and 5. infula" should also be compared, in 
which latter the pectiniform teeth are seen on the whole length of the row. The 
present shell shows an approach to Kaiiella in a few of the outermost laterals, but it 

1 Land and F. W. Moll. India, i. pp. 19, 20, pi. v. fig. ii. 
- Tom. cit. pi. vni. figs, i ^ & 2 e, after Stoliczka. 

37—2 



2 82 FAUNA HAWAIIENSIS 

must be noted that the median teeth have a single outer cusp, while Kaliella has both 
outer and inner cusps ; this latter characteristic is, however, not present in Sitala. 
Kaliella has few teeth in the row, Sitala many; 33 : i : 33- i53 : i ^ 153- respectively. 
A more important link with the genus Sitala is displayed by the presence of right and 
left shell lobes, which Kaliella does not possess ; the close parallel lines of contraction 
across the right shell lobe shew that it has considerable extension in life. Stoliczka 
also mentions in Sitala infula the swollen uterus and the advanced state of development 
of the ova ; pointing to similar embryonic stages in these molluscs. Yet another 
character is in common, namely, the absence of any amatorial organ. The male organ 
of the present species is also slightly different ; I am unable, having only one specimen 
to dissect, to examine this in section. 

"The jaw is very thin and delicate, and so colourless that its detection and ex- 
traction are very difficult. It has a well defined central projection on the cutting edge. 

" The generative organs (Plate XII. figs. 3, 3«) cannot be described so fully as one 
would wish, owing to the expanded state of the uterus. The hermaphrodite duct and 
albumen gland were perfect : and the male portion thence complete. The prostate — as 
it is called by Semper, shewn in his figure of Microcystis myops as a loose fringe-like set 
of convolutions — appears in this species as a closely packed and thickened mass of 
oblong form, flattened on one side, where the oviduct would be lying attached if perfect. 
The vas deferens is given off at the anterior end. The penis is a thickened muscular 
tube, broad and bulbous below, tapering upwards to where the very short thickened 
retractor muscle is given off: the vas deferens at this point has three sharp con- 
volutions ; seen with transmitted light a short, sharp, ' kink ' occurs in the bulbous 
portion near the generative aperture. 

" The sculpture of the shell, magnified about thirty times, presents a very fine, 
regular, slighdy wavy, longitudinally striated surface ; this striation is strongest near 
the suture, becoming finer outwards. There are about 1 1 striae to -003 inch. The 
most advanced embryonic shell consists of 2\ whorls, the sculpture is well shewn on it. 

" The point now to be solved is whether we are to retain this species in Microcystis. 
Mr Sykes regards' M. ornatella as the type of the genus ; this was also the opinion of 
H. Nevill. Further Mr Sykes goes on to say ' Now these small Zonitoids \i.e. those 
of the Hawaiian Islands] hardly fit into the same genus as this species and therefore 
some other generic title is required for them.' The anatomy now described, shews, for 
many reasons, that the shell cannot be placed in Macrochlamys as Mr Sykes, guided by 
the shell characters, proposed. In my opinion it is undoubtedly close to Kaliella, 
still closer to Sitala, and yet there are sufficient differences in the generative organs to 
separate this Hawaiian form from both. If we take the shell alone into account, the 
sculpture presents one character, viz. fine, close longitudinal striation, not found in the 
Indian species of Sitala, in which the general surface is smooth, with spiral liration. 

' P. Malac. Soc. London, 11. i8g6, p. 131. 



MOLLUSC A 283 

The sculpture of Kaliella is finer and transverse to the whorl, so differs still more. 
It therefore may become necessary, if this shell be generally distinct from M. ornatella, 
to create a new genus. 

"When we consider the immense area on the Equatorial belt over which Kaliella, 
Sitala, and this allied form are distributed, it appears that they fall naturally into a 
subfamily of their own which may be called the Sitalinae, Godwin-Austen, noni. nov. ; 
one that is sufficiently distinct from the Durgellinae on the one hand, with which they 
are associated over a large portion of their range, and from the Macrochlaminae on the 
other, where the area of association is more restricted and the differences in the animal 
much greater." (H. H. Godwin-Austen.) 



(3) Philonesia cicercida Gould. 

Helix cicernila Gould, P. Boston Soc. 11. (1846), p. 171 ; U.S. Explor. Exped. 
Moll. pi. V. fig. -]--,. 

Hab. Hawaii (Gould) ; Kohala (Perkins). 



var. boettgeriana Ancey. 

Microcystis cicercula var. boettgeriana Ancey, Bull. Soc. Malac. France, vi. p. 206. 
Hab. Hawaii, Kona (Ancey). 



(4) Philonesia exaequata Gould. 

Helix exaequata Gould, P. Boston Soc. 11. (1S46), p. 171 ; U.S. Explor. Exped. 

Moll. pi. V. fig. 61. 
Helix discnins Pfeiff'er, Zeitschr. flir Malak. vii. 1851, p. 68 \_non Deshayes]. 
Helix obtusangula Pfeiffer, t. c. p. 153. 
Nanina discus Pfeiffer, Tryon, Man. Conch. Ser. 11. Vol. 11. p. i 14. 

Hab. Kauai (Gould, Perkins). 



(5) Philonesia hartmanni Ancey. 

Microcystis hartmanni Anc&y, Bull. Soc. Malac. France, vi. (1889), p. 198. 
Hab. Oahu (Ancey) ; Kalaikoa (Baldwin). 



284 FAUNA HAWAIIENSIS 

(6) Philonesia indefinita Ancey. 

Microcystis indefinita Ancey, Bull. Soc. Malac. France, vi. (1889), p. 203. 
Hab. Maui, east part (Ancey) ; Makawao (Baldwin). 

(7) Philonesia lynnanniana Ancey. 

Microcystis lymanniana Ancey, Mem. Soc. Zool. France, vi. (1893), p. 329. 
Hab. Oahu, Waialae (Ancey). 

(8) Philonesia oahuensis Ancey. 

Microcystis oahuensis Kncey, Bull. Soc. Malac. France, vi. (1889), p. 202. 
Hab. Oahu (Ancey) ; Halemano (Perkins). 

I refer, with some doubt, a single specimen found by Mr Perkins, to this unfigured 
species. 

var. depressinsciila Ancey. 

M. oahuensis var. depressiusciila Ancey, /. c. p. 203. 
Hab. Oahu (Ancey). 

(9) Philonesia perhtcens Ancey. 

Microcystis perlucens Ancey, Bull. Soc. Malac. France, vi. (1889), p. 207. 
Hab. Maui, east part (Ancey). 

(10) Philonesia perkinsi Sykes. 

Macrochlamys pe7'kinsi Sykes, P. Malac. Soc. London, 11. (1896), p. 126. 

Plate XI. figs. 41, 42. 

Hab. Lanai. — (?) Oahu, a single specimen (Perkins). 

(11) Philonesia platy la Ancey. 

Mic7'ocystis platyla Ancey, Bull. Soc. Malac. France, vi. (1889), p. 196; Sykes, 
P. Malac. Soc. London, iii. pi. xiii. figs. 13 — 15. 

Hab. Oahu (Ancey) ; Waianae Mts. (Baldwin, Perkins). 



MOLLUSC A 285 

(12) Philonesia plicosa Ancey. 

Microcystis plicosa Ancey, Bull. Soc. Malac. France, vi. (1889), p. 200. 
Hab. Oahu (Ancey) ; Palolo (Baldwin). 

(13) Philonesia sericans Ancey. 

Microcystis sericans Ancey, P. Malac. Soc. London, iii. (1899), p. 268. 
Hab. Hawaii, Olaa (Ancey). 

(14) Philonesia subrutila Mighels. 

Helix subrutila Mighels, P. Boston Soc. 11. (1845), P- i9- 

Hab. Oahu (Mighels, &c.). — Mr Baldwin gives Kauai, but I doubt this ; the 
species is unknown to me. 

(15) Philonesia S7ibtilissima Gould. 

Helix subtilissima Gould, P. Boston Soc. 11. (1846), p. 177 ; U.S. Explor. Exped. 

Moll. pi. V. fig. 62. 
Unknown to me ; from the figure I am not certain of its generic position. 
Hab. Maui (Gould). 

(16) Philonesia turgida Ancey. 

Microcystis turgida Ancey, Bull. Soc. Malac. France, vii. (1890), p. 339; Sykes, 
P. Malac. Soc. London, iii. pi. xiii. figs. 5 — 7. 

Hab. Maui (Ancey) ; Makawao (Baldwin) ; Mts. at 4000 ft. (Perkins). — A speci- 
men found on Lanai by Mr Perkins, may belong to a variety. 

Obs. The Helix niisella of Fdrussac has been recorded with a query from the 
islands, but does not really belong to their fauna. 

Kaliella Blanford. 
Type the group of Helix barrakporensis Pfr. 



286 FAUNA HAWAIIENSIS 

(i) Kaliella konaensis Sykes. 

Kaliella konaensis Sykes, P. Malac. Soc. London, ii. p. 299. 

Plate Xl/ fig. ■x,i. 

A remarkable little shell which seems to fall between Kaliella and Trochoconulus. 

Hab. Hawaii, Mt. Kona, at 3000 ft. (Perkins). 

Fam. PHILOMYCIDAE. 

Tebennophorus Binney. 

Tcbcnnophorns Binn., J. Boston Soc. iv. 1844, p. 171 (Type Lijuax carolinensis, 
Bosc). 

(i) Tebennophorus bilineatus Benson. 

Incilaria bilineata Benson, Ann. Nat. Hist. ix. 1842, p. 486. 

Philomycus bilineattis Benson, Martens, Preuss. Exped. Ost-Asien, Mollusca, p. 16, 

pi. V. fig. I. 
Tebennophorus australis Bergh ?, Collinge, P. Malac. Soc. London, 11. p. 50. 
Tebennophorus striahis Hasselt, Collinge, /. c. p. 295. 

Hab. Oahu, Mount Tantalus, Honolulu at 2000 ft. — Kauai, Lihue at 2000 ft. — 
Hawaii, Olaa at 2000 ft. (Perkins). 

See Collinge, J. Malac. vii. 1900, p. 80. 

(2) Tebennophorus stjHatus Hasselt. 

Meghimatium striatum Hasselt, Bull. Sci. Nat. Geol. in. 1824, p. 82. 
Tebennophorus striatus Hasselt, Collinge, P. Malac. Soc. London, 11. p. 50. 

Hab. Oahu, Mount Tantalus (Perkins). 

Fam. ENDODONTIDAE. 

Endodonta Albers. 

Ejidodonta Alb., Die Heliceen, 1850, p. 89 (first species Helix lainellosa, Fer.) ; 
Op. cit. Ed. 2, 1S60, p. 90 (" typus Helix lamellosa, Fer."). 



MOLLUSC A 287 



(i) Endodonta apiculata Ancey, 



Eiidodoiita apiculata Ancey, Bull. Soc. Malac. France, vi. (1889), p. 189. 
Hab. Kauai, Dr Newcomb (Ancey). 



(2) Eiidodoiihi lainellosa Ferussac. 

Helix laviellosa Ferussac, Hist. Moll. i. p. 369, pi. li. a, fig. 3 ; Quoy and 
Gaimard, Voy. Freycinet, Zool. p. 469, Pfeifter, Conchylien-Cabinet, Helix, 
p. 197, pi. c. figs. 6 — 8. 

Helix fricki Pfeiffer, P. Zool. Soc. London, 1858, p. 21, pi. xl. fig. 3. 

According to Mdrch (J. Conchyl. xiii. p. 395) this species "depose ses oeufs 
dans rombilic." 

The teeth or lamellae seem to be variable ; some specimens shew traces of a second 
tooth in the upper portion of the outer lip, thus having nine teeth in all. Considerable 
variation is also shewn in the relative proportions of height and breadth, and in the 
width of the umbilicus. 

Hab. Oahu (Pease, Ancey) ; Waianae Mts. and Konahuanui (Baldwin) ; Mt. 
Kaala. — Lanai Mts. behind Koele (Perkins). 



(3) Endodonta laiiiinata Pease. 

Helix laminata Pease, Amer. J. Conch. 11. (1806), p. 292. 

According to the diagnosis this differs from the last by being spirally sculptured 
as well as transversely ribbed, thereby becoming decussated. The teeth appear to 
be identical in number and position and I believe it will, eventually, only prove to be a 
local race. 

Tryon (Man. Conch. Ser. 2, in. p. 70) considered it to be a form of E. cavemula, 
Hombr. and Jacq., stating " I have before me two trays of shells named Helix 
laminata Pease, from the 'Sandwich Is.,' one of them from the describer, which 
undoubtedly represent the same species." Since the two forms differ, from the 
diagnoses, so widely in the armature, there must, 1 think, be some error. 

Hab. Kauai (Pease) ; Kahiliwi to Haena (Baldwin). 

F. H. II. ' 38 



288 FAUNA HAWAIIENSIS 

Sub-genus Thaumatodon Pilsbry. 

(4) Endodonta ( Thaumatodoii) coniorta Ferussac. 

Helix contorta Ferussac, Hist. Moll. i. p. 10, pi. li. a, fig. 2. 
Helix intercarinata Mighels, P. Boston Soc. 11. (1845), P- '8. 

Specimens found by Mr Perkins, and which I refer to this species, appear to shew 
considerable variation and may be divided as follows : 

A. Six specimens, fairly typical in shape and size, but only one is furnished with 
five palatal teeth, the others having four. 

B. One specimen, darker in colouration, the colour markings being very distinct, 
palatal teeth five, these being remarkably incrassated. 

C. A long series (from Makaweli), larger, slightly more strongly sculptured and 
very variable in colouration, sometimes the dark brown colour predominating, at others 
a greenish yellow. All appear to have four palatal teeth only. 

Hab. Oahu (various authors). — Kauai (Perkins). 

(5) Endodonta [Thaumatodon) hystricella Pfeiffer. 

Helix hystricella Pfeiffer, P. Zool. Soc. London, 1S59, p. 25. 

The original examples of this unfigured species, referred to as in Mus. Cuming, 
do not appear to be now in the British Museum. Two specimens found by Mr Perkins 
agree well with Pfeiffer's diagnosis and dimensions ; they also accord in the number of 
teeth. 

Hab. Kauai (Pease). — Oahu, Kaala (Perkins). 

(6) Endodonta [Thaumatodon) nuda Ancey. 
Endodonta [Thatimatodoti) nuda Ancey, P. Malac. Soc. London, iii. (1S99), p. 268, 

pi. XII. fig. I. 

Hab. Hawaii, Olaa (Ancey). 

(7) Endodonta [Thaitmatodon) 7-ingcns Sykes. 

Endodonta [Thazimatodon) ringens Sykes, P. Malac. Soc. 11. (1896), p. 126. 

Plate XL figs. 39, 40. 

In describing this species, I referred to it as having four teeth within the outer 
lip; perhaps it would be more correct to say "one basal tooth and three within the 



MOLLUSC A 289 

outer lip." The ribs appear to be at varying distances apart. The Molokai specimens 
appear to belong to a large variety. 

Hab. Lanai Mountains, behind Koele. — Molokai in wet forest above Pelekunu 
(Perkins). 

(8) Endodonta ( Thaumatodon) rugata Pease. 

Helix riigata Pease, Amer. J. Conch. 11. (1866), p. 291. 
Hab. Maui (Pease). 

Sub-genus Nesophila Pilsbry. 

The following table may assist in separating the species of Nesophila. 

A. Parietal lamella; absent. — E. capillata Pease. 

B. Parietal lamella single. — E. decussatula Pease ; E. elisae Ancey ; E. jugosa 
Mighels ; E. lanaiensis Sykes ; E. stellula Gould. . 

C. Parietal lamellae two. — E. binai'ia Pfeiffer ; E. liyshdx (Mighels MS.) Pfeiffer ; 
E. patuicostata Pease. 

D. Parietal lamellae several. — E. baldwini Ancey ; E. distans Pease ; E. tiara 
Mighels. 

(9) Endodonta {^Nesophila) baldwini Ancey. 

Charopa baldwini Ancey, Bull. Soc. Malac. France, vi. (1889), p. 176. 
Mons. Ancey also records a white variety. 
Hab. Hawaiian Islands (Ancey). 

(10) Endodonta {Nesophila) binaria Pfeiffer. 

Helix binaria Pfeiffer, P. Zool. Soc. London, 1856, p. n- 

I am unable to trace the type of this species, which should have passed with 
Cuming's collection into the British Museum. 
Hab. Kauai (Pease). 

(11) Endodonta {Nesophila) capillata Pease. 

Helix capillata Pease, Amer. J. Conch. 11. (1866), p. 292. 

Hab. Kauai (Pease). 

38—2 



290 FAUNA HAWAIIENSIS 

(12) Endodoiita {^Nesophila) decussatit/a Pease. 

Helix deciissatjila Pease, Amer. J. Conch. 11. (t866), p. 291. 
Hab. Molokai (Pease) ; Mountains at 4000 ft. (Perkins). 

Mr Baldwin gives " Kauai" as the habitat, but, since he marks it as a species he 
has not seen, I think there is probably some error. 

(13) Endodonta [Nesophila) distans Pease. 

Helix distans Pease, Amer. J. Conch. 11. (1866), p. 290. 
Hab. Kauai (Pease). 

(14) Endodonta [Nesophila) elisae Ancey. 

Pitys elisae Ancey, Bull. Soc. Malac. France, vi. (1889), p. 180. 

Unknown to me. 

Hab. } Hawaiian Islands (Ancey). 

(15) Endodonta {Nesophila) hystrix (Mighels MS.) Pfeiffer. 

Helix hystrix Pfeiffer, Symb. Hist. Hel. in. p. 67 ; Gould, U. S. Explor. Exped. 

Moll. pi. IV. fig. 52*. 
Helix setigera Gould, P. Boston Soc. i. p. 174 [nee Sowerby]. 

It is, of course, not the Helix hystrix of Cox, an Australian species. 

Hab. Oahu (authors) ; Mount Kaala, Oahu (Perkins). 

(16) Endodonta [Nesophila) j'ugosa Mighels. 

Helix jugosa Mighels, P. Boston Soc. 11. (1845), p. 19. 

Helix rubiginosa Gould, P. Boston Soc. 11. (1846), p. 173; U. S. Explor. Exped. 
Moll. pi. IV. p. 49. 

The two forms were first united by Pease ^ who stated that the synonymy was 
accepted by Gould ; recently Mons. Ancey^ has revived E. rubiginosa as a species, 
referring to it some shells from Oahu. In this state of conflict 1 have followed Pease, 
considering that he and Gould were in the best position to form an opinion. 

Hab. Kauai, Waioli to Kapaa (Baldwin) ; Kauai (Perkins, etc.). 

' J. Conchyl. xix. (1870), p. 95. 

■ Bull. Soc. Malac. France, vi. (1889), p. 179. 



MOLLUSC A 291 

(17) Endodonta {^Nesophild) lanaiensis Sykes. 

Endodonta [Nesopki/a) lanaiensis Sykes, P. Malac. Soc. London, 11. (1S96), p. 127. 

Plate XI. figs. Z7^ 38. 

Specimens from Kauai which I refer to this species are strongly hispid in the 
young state, but with age the hairs appear to be rubbed off ; none of those from Lanai 
are very young, and only traces of hairs can be seen. The species appears to be near 
E. decussahila, but it almost lacks decussation and is darker in colour ; the interstices ot 
the ribs are closely, finely, striate. Save for the presence of a parietal lamella, the 
Kauai specimens approach Pease's diagnosis of E. capillata. 

Hab. Lanai Mountains, behind Koele. — Kauai, Makaweli, on Dracaena and 
Clieirodendron (Perkins). 

(iS) Endodonta (Nesop/ii/a) paucicostata Pease. 

Helix paiicicostata Pease, J. Conchyl. xviii. (1870), p. 395. 
Helix filocostata Pease, P. Zool. Soc. London, 1871, p. 454. 

Hab. Kauai (Pease). 

(19) Endodonta {Nesophila) stcllula Gould. 

Helix stellnla Gould, P. Boston Soc. i. (1844), p. 174; U. S. Explor. Exped. 

Moll pi. IV. fig. 52t. 
Hab. Maui (Gould). 

(20) Endodonta {Nesophila) tiara Mighels. 

Helix tiara Mighels, P. Boston Soc. 11. (1845), p. 19; Kuster, Conch.-Cab. Helix, 

pi. cxxv. figs. 9 — II. 
According to Mons. Ancey^ this species possesses several parietal lamella ; if so, 
the character has been omitted from the various diagnoses. 
Hab. Kauai (various authors). 

(21) Endodonta {Nesophila), sp. 

Two interesting little specimens with a depressed spire were found on Molokai by 
Mr Perkins, the exact habitat being " Forest above Pelekunu "; they have 4— 4i whorls, 
with two parietal lamellae and no teeth within the other lip, but appear not to be adult. 

Hab. Molokai. 

' Bull. Soc. Malac. France, vi. p. 175. 



292 FAUNA HAWAIIENSIS 

Pterodiscus Pilsbry. 
Type, P. ivesleyi Sykes. 

(i) Pterodiscus digonophorus Ancey. 

Patida digonophora Ancey, Bull. Soc. Make. France, vi. (1889), p. 171 ; Sykes, 

P. Malac. Soc. London, iii. pi. xiii. figs. 9 — 11. 
Hab. Oahu (Ancey); Waianae Mts. (Baldwin). 

(2) Pterodiscus petasus Ancey. 

Pterodiscus petasus Ancey, P. Malac. Soc. London, in. (1899), p. 268, pi. xii. 

fig- 4- 
Hab. Oahu, Waianae Mts. (Ancey). 

(3) Pterodiscus wesleyi Sykes. 

Endodonta [Pterodiscus) weskyi Syk^s,, P. Malac. Soc. London, iii. (1896), p. 127. 
Endodonta [Pterodiscus) alata Pfeiffer, Pilsbry, Man. Conch. (2) ix. p. 36, pi. iv. 
fig. 44 [nee Helix alata, Pfeiffer]. 

Hab. Hawaiian Islands. 

The following two species, placed in this group by Mr Pilsbry', with the habitat of 
Hawaiian Islands, are unknown to me. They were originally described from " Islands 
of the Central Pacific " by Pease : the first has been recorded from Tahiti, but never 
again found there, and Mons. Ancey has suggested a Hawaiian origin ; the second has 
been localized as from (.-*) Lanai. They are Helix prostrata and H. depressifortnis 
(P. Zool. Soc. London, 1864, p. 670). 

Fam. HELICIDAE. 

Papuina von Martens. 

Papuina Mart., Die Heliceen, Ed. 2, i860, p. 166 (type Helix littms. Lesson). 

(i) Pap2tina barnaclei Smith. 

Helix [Merope?) barnaclei Smith, Ann. Nat. Hist. (4) xx. p. 242. 

I am informed that careful search has been made, in the neighbourhood indicated, 
but that no trace of the species can be found. At present, bearing in mind on the one 

' Man. Conch. (2) ix. p. 36. 



MOLLUSCA 293 

hand the improbability of a species of Papuina occurring in Hawaii, and on the other 
the positive statement of the original collector, I can but include it, with this note of 
warning. 

Hab, Hawaii, eight miles from Kailua (Smith). 



EuLOTA Hartmann. 

Eulota Hart., Erd- und Susswasser Gasteropoden, p. 179 (type Hciix Jriitiaun, 
Mull.). The date usually given is 1842, but the title-page of the copy in the British 
Museum bears that of 1840. 



(i) Eulota similaris Fdrussac. 

Helix similaris Ferussac, Prodrome, 1822, p. 47 {iiom. sol); Hist. Moll. i. p. 171, 
pi. XXV. B, figs. I 4, XXVII. A, figs. I — 5. 

A widely scattered species ; presumably not indigenous. 

Hab. Kauai (Pilsbry). — Oahu, Tantalus (Perkins). 

The following have been described under the term Helix and recorded from the 
Islands. 

Helix fornicata Gould, P. Boston Soc. 11. (1846), p. 172. 

Supposed to come from Kauai. Tryon notes': "In the corrigenda to the Mollusca 
of the Wilkes Exploring Expedition, Dr Gould states that the only specimen was lost, 
and H. tongana Quoy, figured by the artist for this species." 

Helix sandwichensis Pfeiffer, P. Zool. Soc. London, 1849, p. 128. 
Appears to be the young of a South American Systrophia. 
Helix exserta Pfeiffer, P. Zool. Soc. London, 1856, p. 32. 
Only a fragment of the type remains ; it has never been figured. 

Fam. PUPIDAE. 

Pupa Draparnaud (1801). 

Pupa Drap., Tabl. Moll. France, pp. 32, 56 (first species Turbo luuscoruin L.) ; 

1805, Hist. Moll. France, p. 59. 
There appears to be a Pupa of Lamarck of even date (Syst. anim. sans Vert, 
p. 88) with Turbo uva as type ; also, through the kindness of Mr Sherborn, I have 

' Man. Conch. (2), 111. p. 27. 



294 FAUNA HAUAIIENSIS 

examined the Museum Bolteniamnn, Ed. i. 1798, and Bolten proposed Pupa (p. no) 
for Vohita flaiuinea and V. solidula. The first species belongs to Actacon (iSio) and 
the second is the type of Solidula (1807). If, therefore, Bolten's names are to be 
used, Pupa Drap. cannot stand unless it prove to be the same as Pupa Humphrey, 
1797 (Mus. Calonnianum). 

An excellent study of the Pupidae of Oceania has been written by Dr Boettger ; 
the attention of subsequent authors does not appear to have been sufficiently directed 
to this paper. 

( r ) Pupa acantliinula Ancey. 

Pupa acantliinula Ancey, Mem. Soc. Zool. France, v. (1892), p. 709. 
Hab. Oahu. Makiki (Ancey). 

(2) Pupa ad)nodcsta Mighels. 

Pupa admodesta Mighels, P. Boston Soc. 11. (1845), p. 19; Boettger, Conch. 
Mittheil. i. p. 58, pi. xii. fig. 15. 

Hab. Oahu (Mighels). 

(3) Pupa bacca Pease. 

Pupa bacca Pease, P. Zool. Soc. London, 1S71, p. 462. 
Hab. Hawaii, Kalapana (Pease). 

(4) Pupa cost at a Pease. 

Pupa costata Pea.se, P. Zool. Soc. London, 1871, p. 462. 
Hab. Hawaii (Pease). 

(5) Pupa lyonsiana Ancey. 

Pupa lyonsiana Ancey, Mem. Soc. Zool. France, v. (1892), p. 713. 
Hab. Oahu, Punahou (Ancey). 

(6) Pupa lyrata Gould. 

Pupa lyrata Gould, P. Boston Soc. i. (1S43), p. 139; Boettger, Conch. Mittheil. 

I. p. 61, pi. XII. fig. 17. 
Pupa striatula Pease, P. Zool. Soc. London, 1871, p. 461. 
Pupa magdalenae Ancey, Mem. Soc. Zool. France, v. (1892), p. 716. 



MOLLUSCA 295 

I follow Dr Boettger in placing P. striahda with P. lyrata, and have added 
P. magdalenae, since the character of two parietal teeth, on which Mons. Ancey lays 
special stress, is found in some undoubted specimens of P. lyrata. 

Hab. Hawaii (Pease). — Oahu (Baldwin); Palaina (Ancey). 



(7) Pupa luirabilis Ancey. 

Pupa mirabilis Ancey, Bull. Soc. Malac. France, vii. (1890), p. 339. 
Hab. Oahu (Ancey). 

(8) Pupa newcombi Pfeiffer. 

Pzipa newcombi Pfeiffer, P. Zool. Soc. London, 1852 [1854], p. 69 ; Kuster, Conch.- 

Cab. Pupa, pi. xx. figs. 23, 24. 
Pupa costulosa Pease, P. Zool. Soc. London, 1871, p. 462. 

Hab. Hawaii (Pease, Baldwin). — Oahu and Kauai (Baldwin) [sed quaere, E.R.S.]. 

var. seniimilum Boettger. 

Pupa iieiocoinbi var. seniinu/uni Boettger, Conch. Mittheil. i. p. 5S, pi. xii. 
fig. 14. 

Hab. Probably Hawaii (Boettger). 

(9) Pupa pedicuius Shuttl. var. nacca Gould. 

Vertigo nacca Gould, P. Boston Soc. viii. (1862), p. 280. 

Dr Boettger has pointed out that this is only a form of P. pedicuius, Shuttleworth 
(Bern. Mittheil. 1852, p. 296), and barely of varietal rank. The typical form appears 
not to be found in the Hawaiian Islands. 

Hab. Hawaii (Gould). 

(10) Pupa perlonga Pease. 

Pupa perlonga Pease, P. Zool. Soc. London, 1S71, p. 462 ; Boettger, Conch. 
Mittheil. i. p. 60, pi. xii. fig. 16. 

Only two specimens, which I refer here with some hesitation, since the teeth are 
rather obscure. 

Hab. Oahu (Pease). — ^Kauai, Makaweli (Perkins). 
F. H. II. 39 



296 FAUNA HAWAIIENSIS 

Fam. ACHATINELLIDAE. 

The various species of this family form probably one of the most puzzling groups 
of land-shells known : numerous ' species ' have been described, founded almost entirely 
on shell colouration or banding, and this, bearing in mind such protean forms as Tachea 
nemoralis or Polytnita picia, is a course which often leads to endless trouble. At present, 
such genera as Achatinella s. s. and Achatinellastriim are in utter confusion, and any 
attempt at a list of species simply reflects the general view of the writer and cannot be 
regarded as authoritative. 

Of recent years some species have been described, based on shell-colouration com- 
bined with that of the mantle, but it should be remembered that some malacological 
characteristics are frequently as liable to variation as are conchological ones. The 
severance of species by consideration of habitat is, again, not a necessarily certain 
guide, since one species of mollusc may range widely, while at different points of its 
range, local variations may occur. 

The history of the Achatinellidae in literature commences with Dixon's description 
of Turbo apexftdva ; subsequently stray species were described by various authors, but 
no serious attempts were made at their study until about 1850-60, when Newcomb and 
Pfeiffer added greatly to our knowledge. To Mr Gulick, in 1858, we owe large addi- 
tions to the catalogue — so far as names go — but his views of species were somewhat 
too narrow, and he described many trifling variations as species, mainly on the ground 
of geographical range. Later, Pease did useful work and gave a catalogue of the family. 
Dr Hartman, in 1888, listed the known forms in a similar way. Of recent years Mr 
Baldwin has done the chief work on the group, and his very valuable catalogue has been 
of great assistance. 

Numerous genera and sections have been described, almost all on purely concho- 
logical grounds, and so variable are the forms that linking species are easily found. 
Still they prove useful in the arrangement of species and therefore have been admitted 
in this work. 

A list of them arranged in chronological order may be of use. 
1828. Achatinella Swainson, O. J. Sci. Lit. and Arts, p. 81. 



1847 
1854 
1854 
1854 
1854 
1854 
1854 



Leptachatina Gould, P. Boston Soc. 11. p. 201. 
Partulina Pfeiffer, Malak. Blatt. i. p. 114. 
Newcoftibia Pfeiffer, /. c. p. 1 1 7. 
Bulimella Pfeiffer, /. c. p. 11 9. 
Laniinella Pfeiffer, t. c. p. 126. 
Ackalinellastriiin Pfeiffer, t. c. p. 133. 
Labiclla Pfeiffer, /. c. p. 142. 



MOLL use A 



297 



1855 
1855 
1855 
1855 
i860, 
1870, 
1870, 

1875 
1899 
1899 



Carelia H. & A. Adams, Gen. Rec. Moll. 11. p. 132. 

Ainastra H. & A. Adams, /. c. p. 137. 

Auriculella Pfeiffer, Malak. Blatt. 11. p. 3. 

Frickella Pfeiffer, /. c. p. 3. 

Apex Martens, Die Heliceen, p. 248. 

Ebiirnella Pease, P. Zool. Soc. London, 1869, p. 647. 

Perdicella Pease, /. c. p. 649. 

Carijiella Pfeiffer, Novit. Conch, iv. p. 116. 

Thaanumia Ancey, P. Malac. Soc. London, iii. p. 269. 

Baldwinia Ancey, /. c. p. 270. 



Mr Pilsbry's remark, that the use of Ferussac's term Helicteres " would open the 
door to an endless series of vagaries in nomenclature," appears to me to be thoroughly 
justified ; for the converse view see Pease, P. Zool. Soc. London, 1862, p. 3. The 
Achatinella of Schliiter (1838) appears to be a mixture of Ferussacia and allied groups. 

Our knowledge of the anatomy is due primarily to Bland and Binney, who pointed 
out that the Achatinellidae may be divided into two main groups based on the cha- 
racters of the radula and jaw ; also that Carelia, while it agrees in the radula with the 
Leptachatina and Amastra group, differs in the characters of the jaw. Heynemann has 
also written on the subject, and recently notes have been published by Messrs Gwatkin, 
Pilsbry and Suter. Mr Pilsbry has recently^ stated with reference to Pariula and 
Achatinella that " these forms have no relations with the Bulimulidae and Achatinidae, 
with which conchologists associate them, but lie at the base of the terrestrial pulmonate 
tree." 

The classification here adopted is : 

Ganw?, Achatinella. Suhgenn. Achatinella s. s. [Syn. Apex.) Bidimella. Partu- 
lina. (Syn. Baldzvinia.) Acliatinellastrum. (Syn. Eburnella.) 

Genus Perdicella. 

Genus N^ewconibia. 

Genus Amastra. Subgenn. Amastra s. s. Laminclla. Amastrella, n. subgen. 
Kauaia, n. n. \_Carinella, Pfr., nee Sby.J 

Genus Leptachatina. 

Genus Thaanumia. 

Genus Carelia. 

Genus Aiiricnlella. 

Genus Frickella. 



Ann. nat. Hist. iv. (1S99), p. 156. 



39—2 



298 FAUNA HAWAIIENSIS 



ACHATINELLA S. Str. 

Synonym. Apex von Martens. 

Well has Dr Hartman remarked, that the species of this group "are involved in 
almost inextricable confusion." Our ignorance of the animal in most forms, combined 
with the fact that authors have described frequently from single specimens, or from 
series of two or three, entirely at present prevents one ascertaining the true specific 
limits. 

In the one or two instances where I have attempted 'lumping,' the results are due 
to a long series of specimens in which I have been unable to arrive at a definite break 
between the one 'species' and the other. 

The conclusions I have drawn are founded on Newcomb's and Pfeiffer's type 
specimens ; specimens compared with Gulick's types ; further a few of the types de- 
scribed by Mr Smith ; and, finally, the specimens collected by Mr Perkins and a collec- 
tion formed by Mr Hutchison. 

All the species of this group are confined to the Island of Oahu. 

(i) Achatinella albospira Smith. 

Apex albospira Smith, P. Zool. Soc, London, 1873, p. "]"], pi. x. fig. 8. 

Unknown to me; it has been united by Dr Hartman with A. turgicia Newc. 
[ = A. cestus Newc.]. 

Hab. Oahu, Ewa. 

(2) Achatinella apexfoilva Dixon. 

Turbo apex fulva Dixon, Voyage round the world, 1789, p. 344, figd. on an un- 
numbered plate. 

Turbo lugubris Chemnitz, Conch. -Cab. xi. 1795, p. 278, pi. ccix. figs. 2059, 
2060. 

Cocldogena [Helicteres) lugubris Chemnitz, Ferussac, Prodrome, p. 60, no. 431. 

Monodonta seminigra Lamarck, Hist. Nat. Anim. sans Vert. vii. (1822), p. -i,-]. 

Achatinella pica Swainson, Zool. Ill, Moll, sen 11. pi. xcix. fig. i. 

Hab. Oahu, Kawailoa (Baldwin). 

Dixon was thoroughly binomial, as a reference to his work will show : his figures 
are good, and the only point that can be made against the utilization of his name is that 
it appeared in three words. This, probably, was a printer's error, or a slip, as other 
names in the same work are proper, and suitable descriptions are given. 



MOLLUSC A 299 

(3) Achatinella apicata (Newcomb MS.) Pfeiffer. 

Achatiiiella apicata Pfeiffer, P. Zool. Soc. London, 1855 [1856], p. 210. 
Hab. Oahu, Halemano (Perkins). 

van polynnorpJia, Gulick. 

Apex polymorpha Gulick, P. Zool. Soc. London, 1873, p. 81, pi. x. fig. 5. 

Hab. Oahu, Wahiawa, Kalaikoa, Ahonui (Gulick) ; Kawailoa Gulch, and above 
Ewa (Perkins). 

van vespertina Baldwin. 

Achatinella {Apex) vespertina Baldwin, P. Ac. Philad. 1895, P- 219 [shell and 

animal], pi. x. fig. 14. 
Hab. Oahu, Kawailoa (Baldwin) ; from a ridge between Waala and Kawailoa 
Gulches (Perkins). 

var. alba, var. n. 

Shell snow-white, save for the peristome being margined with lilac, similar tinting 
appearing on the columella plait, and inside the outer lip at its junction with the body- 
whorl of the shell. 

Hab. Oahu, near head of Kawailoa Gulch (Perkins). 

A fine series of this species. It appears to be distinct from A szuiftii Newc. 
\^=A. cestus Newc. var.], to which Newcomb subsequently referred it, and the type of 
which I have examined. The shell is larger, different in form, and more polished, the 
ground colouring darker, and the banding not so conspicuous in the typical form : 
the protoconch is in general brown, and but rarely white. The prevailing tint 
is brown, sometimes becoming black with a white area below the suture, rarely 
altogether snow-white. A few, from near Kawailoa, are tinted with lilac on an 
ashy ground, occasionally having a chestnut sutural line ; these form a passage to 
A. vespertina Baldwin, in which the lilac tint has changed to cream-colour. This 
latter is the only form in which the animal appears to have been noticed. 

(4) Achatinella bicolor (Gulick) Pfeiffen 

Achatinella bicolor Pfeiffer, Mon. Hel. Viv. iv. 1859, p. 529. 
Near, apparently, to A. cookei Baldwin. 
Hab. Oahu, Lehui (Pfeiffer). 



300 FAUNA HAWAIIENSIS 

(5) Achatinella cestus Newcomb. 

Achatinella cestus Newcomb, P. Zool. Soc. London, 1853 [1854], p. 132, pi. xxii. 

fig. 8. 
Achaiitiella turgida Newcomb, torn. cit. p. 134, pi. xxii. figs. 10, \oa. 

Hab. Oahu, Palolo and Ewa (Newcomb); Ewa and Halemano (Perkins). 

var. swiftii Newcomb. 

Achatinella swiftii Newcomb, P. Zool. Soc. London, 1853 [1854], p. 133, 
pi. XXII. fig. 9 ; Ann. Lye. New York, vi. p. 325 [animal]. 

Achatinella {Achatinellastrum) valida Pfeiffer, P. Zool. Soc. London, 1855, p. 6, 
pi. XXX. fig. 24. 

Apex flavidtts Gulick, op. cit. 1873, p. 80, pi. x. fig. i. 

Apex tubei^ans Gulick, torn. cit. p. 81, pi. x. fig. 3. 

Apex liliaceus Gulick, toni. cit. p. 79, pi. x. fig. 4. 

Apex tu7'biniforniis Gulick, torn. cit. p. 81, pi. x. fig. 7. 

Hab. Oahu, Ewa (Newcomb) ; as A. Jlavidus, Kalaikoa and Ahonui (Gulick) ; 
as A. tuberans, Kalaikoa, Ahonui, Wahiawa and Halemano (Gulick) ; as A. turbi- 
ni/ormis, Kalaikoa and Lehui (Gulick). 

var. forbesiana Pfeiffer. 

Bulimella forbesiana Pfeiffer, P. Zool. Soc. London, 1855, p. 5, pi. xxx. fig. 16. 

Apex gtdickii Smith, loc. cit. 1873, p. 78, pi. ix. fig. 19 \jion 17]. 

Hab. Oahu as A. gulickii, Kalaikoa, Ahonui and Waialei (Smith). 

The type form has the bands interrupted ; in the variety szuiftii they are continuous 
and almost confluent, recalling a dwarf A. bigubris. From the variety swiftii, given a 
fair number of specimens, we may pass by slight variations up to a nearly white shell 
with hardly a trace of banding. One specimen is light yellow and almost unmarked 
except the latter half of the last whorl, which is strongly banded with dark brown (Plate 
XL figs. 6, 7). The general brown colouring is more persistent in the variety than in 
the typical form. The variety forbesiana is somewhat more elongate and slightly 
differently banded and coloured. 

(6) Achatinella cookei Baldwin. 

Achatinella (Apex) cookei Baldwin, P. Ac. Philad. 1895, P- -20, pi. x. fig. 15 
[shell and animal]; Suter, /. c. p. 239 [radula]. 

Hab. Oahu, Waiau (Baldwin). 



MOLL use A 301 

(7) Achatiiiella decora Ferussac. 

Cochlogena {Helicteres) decora Ferussac, Prodrome, 1822, p. 60, no. 430. 

Turbo lugubris sinistrorsus Chemnitz, Conch. -Cab. xi. 1 795, p. 307, pi. ccxiii. 

figs. 3014, 3015. 
Achatinella perversa Swainson, Quart. Journ. Sci. &c. i. 1828, p. 84; Zool. 111. 

1833, ser. II. pi. xcix. fig. 2 ; Newcomb, Ann. Lye. New York, vi. 1858, 

p. 309 [animal]. 
Achatinella concidcns Gulick, Ann. Lye. New York vi. 1858, p. 234, pi. viii. 

fig- 54- 
Hab. Oahu, Halemano, Waimea. 

var. leucopJuiea Guliek. 

Apex leucophaeus Guliek, P. Zool. Soc. London, 1873, p. 82, pi. ix. fig. 16. 
Hab. Oahu, Waialei. 

var. neglecta Smith. 

Apex neglectus Smith, P. Zool. Soe. London, 1873, p. 78, pi. ix. fig. 22. 

Apex coniforrnis Guliek, toni. cit. p. 81, pi. ix. fig. 17. 

Hab. Oahu, Wahiawa, Kalaikoa, Ahonui, Halemano (Smith. Gulick). 

var. versicolor Guliek. 
Apex versicolor QviXxck, P. Zool. Soc. London, 1873, p. 80, pi. ix. fig. 18. 
Hab. Oahu, Ahonui, Kalaikoa. 

var. innotabilis Smith. 

Apex innotabilis Smith, P. Zool. Soc. London, 1873, p. 78, pi. ix. fig. 23. 
Hab. Oahu. 

subspecies vmstelina Mighels. 

Achatinella mustelina Mighels, P. Boston Soc. 11. 1845, p. 21 ; Reeve. Conch. Icon. 

Achatinella, pi. iii. fig. 21; Newcomb, Ann. Lye. New York. vi. p. 313 

[animal]. 
Hab. Oahu. Waianae, Kaala. 



302 FAUNA HAWAIIENSIS 

var. kitcorraphe Gulick. 

Apex leucorraphe Gulick, P. Zool. Soc. London, 1873, p. 79, pi. x. fig. 2. 
Hab. Oahu, Kalaikoa. 

var. cinei'osa Pfeiffer. 

Achatinella {Bulimella) cinerosa Pfeiffer, P. Zool. Soc. London, 1855, P- 2, 
pi. XXX. fig. 5. 

Hab. Oahu. 

It is with much dififidence that the above 'lumping' has been attempted. While 
not very difficult as a general rule to identify single specimens, I have been unable to 
divide the long series of specimens which, partly collected by Mr Perkins and partly by 
Mr Hutchison, have passed through my hands. Shape, colour, and form, seem to be 
as nothing, and one passes by infinitesimal graduations from one so-called species to 
another. True it is that the animals are said to differ in colour, but this alone is not, in 
my opinion, sufficient for a specific character ; even if it be so, it can, in general, only 
avail the field naturalist, and not the Museum student. 

The variety neglecta Smith is not really so greenish as represented in the original 
figure ; fig. 23 on the same plate gives a better idea of the real colour. 

Ferussac appears to have confounded more than one distinct form under his decora ; 
the name has therefore been here used for the species he first referred to. 

(8) Achatinella doliuni Pfeiffer. 

Achatinella {Bulimella) doliuin Pfeiffer, P. Zool. Soc. London, 1855, p. 5, pi. xxx. 

fig- 15- 
Hab. Molokai (Baldwin). 

I fancy this habitat must be wrong and that the species really belongs to Oahu ; 
the shell is very close to A. hanleyana Pf., and may prove to be only a colour variety. 

(9) Achatinella hanleyana Pfeiffer. 

Achatinella {Bulimella) hanleyana Pfeiffer, P. Zool. Soc. London, 1855, p. 202. 

Related to the form of A. lorata described as A. nobilis, and may prove to be an 
extreme variety. 

Hab. Oahu. 



MOLL use A 303 

(10) Achatinella lorata Ferussac. 

Helix {Cochlogend) lorata Ferussac, Prodrome, 1822, p. 60. 

Achatina lorata Ferussac, Deshayes, Hist. Moll. 11. p. 193, pi. clv. figs. 9 — 11. 

Achatinella lorata Ferussac, Newcomb, Ann. Lye. New York, iv. p. 310 [animal]; 

Semper, Reis. im Philippinen, Landmollusken, pi. xvi. fig. 23 [anatomy]. 
Achatinella alba Nuttall, Jay, Cat. Shells, Ed. in. 1839, p. 58 \jio)nen soliini\. 
Achatinella pallida Nuttall, Jay, loc. cit. ; Reeve, Conch. Icon. Achatinella, sp. 2. 
Achatinella [Btilimella) nobilis Pfeiffer, P. Zool. Soc. London, 1855, p. 202. 
Achatinella ventrosa Pfeiffer, op. cit. 1855, p. 6, pi. xxx. fig. 20. 
Non A. lorata Ferussac, Reeve, Conch. Icon. Achatinella. sp. 6. 

A very variable shell, with or without colour bands, and, occasionally, pure white. 
Hab. Oahu (various authors) ; Manoa to Halawa (Baldwin) ; Nuuanu, Head 
of Panoa Valley, Mount Tantalus (Perkins). 

(11) Achatinella multilineata Newcomb. 

Achatinella 7nultilineata Newcomb, P. Zool. Soc. London, 1853 [1854], p. 138, 
pi. XXII. fig. 23. 
Achatinella {Bulimella) monacha Pfeiffer, op. cit. 1855, p. 3, pi. xxx. fig. 9. 

Hab. Oahu, Waianae Mountains (Baldwin) ; Koolau poko (Newcomb). Dr 
Hartman referred this species, apparently by error, to Maui. 

(12) Achatinella napus Pfeiffer. 

Achatinella {A chatinellastrum) naptts Pfeiffer, P. Zool. Soc. London, 1855, p. 5, 

pi. xxx. fig. 19. 
Achatinella {Bulimella) concavospira Pfeiffer, op. cit. 1859, p. 30. 
Apex leucozonus Gulick, op. cit. 1873, p. 83, pi. x. fig. 6. 

Hab. Oahu. 

I reo-ret to be unable to agree with Newcomb that A. napus is the same as 
A. ptUcherrima Swainson. A. concavospira seems to be only an elongate variety ; the 
types of both species are in the British Museum (Natural History). 

(13) Achatinella ovum Pfeiffer. 

Achatinella {Achatinellastrum) ovum Pfeiffer, P. Zool. Soc. London, 1856, p. 334. 

Hab. Oahu. 
F. H. II. 40 



304 FAUNA HAWAIIENSIS 

(14) Achatinella pulchella Pfeiffer. 

Achatinella [Achaiinellastncm) pulche/la Pfeiffer, P. Zool. Soc. London, 1855, p. 6, 
pi. XXX. fig. 2. 

A small species, very variable in colour, with a blunt apex, and somewhat 
depressed in form. 

Hab. Oahu, mountains behind Ewa (Perkins). 

(15) Achatinella sordida Newcomb. 

Achatinella sordida Newcomb, P. Zool. Soc. London, 1853 [1854], p. 139, pi. 
XXIII. fig. 27. 

Some specimens run very close to A, decora Fer. 
Hab. Oahu, Lihue (Newcomb). 

(16) Achatinella swainsoni Pfeiffer. 

Achatinella i^Bulimella) swainsoni Pfeiffer, P. Zool. Soc. London, 1855, p. 4, pi. 
XXX. fig. 13. 

Newcomb suggested that this might be only a form of A. sordida; it appears, 
however, to be distinct, being broader, brown in general coloration, and having a 
brown, in place of a white lip. It is a little doubtful, from its form, if it be correctly 
placed in this group, but the sections are very artificial. 

Hab. Oahu. 

(17) Achatinella vittata Reeve. 

Achatinella vittata Reeve, Conch. Icon. Achatinella, 1S50, sp. 9. 

Achatinella simiilans Reeve, loc. cit. sp. 15. 

Achatinella {Achatinellastrum) globosa Pfeiffer, P. Zool. Soc. London, 1855, p. 7, 

pi. XXX. fig. 25. 
Apex albofasciatiis Smith, op. cit. 1873, p. 78, pi. ix. fig. 21. 
Apex timiefactus Gulick, torn. cit. p. 82, pi. ix. fig. 20. 
Helix decora F^russac, Quoy and Gaimard, Voy. Uranie et Phys. 1824, Zool. 

p. 478 [nee H. decora Fer. i%2 2 = A. perz'ersa Swainson]. 
Achatina decora Ferussac, Deshayes, Hist. Moll. 11. pt. 2, p. 191, pi. CLV. 

figs. 5, 7. 
Achatinella decora Ferussac, Newcomb, Ann. Lye. New York, vi. ^. 307 

[animal]. 
? ? Achatinella vestita Mighels, P. Boston Soc. 11. 1845, p. 20. 
Hab. Oahu, Waheawa, Halemano, Nuuanu Valley, &c. 



MOLLUSC A 305 

van cinerea, n. var. 

Banding almost black on the last whorl, ash coloured on the whorl above, the 
upper whorls tinted with pale brown banding above the suture, replaced by an almost 
black line at the apex. 

Hab. Oahu, Nuuanu (Perkins). 

Having examined the types of the first five species mentioned in the above 
synonymy, I am unable to separate them specifically ; with a fair series of specimens 
the forms shade one into another. The variety is noteworthy for its banding being 
ashy and almost black, while in the typical form it is red-brown in various patterns. If 
A. vestita be really this species it takes precedence in date : I have never seen a 
specimen. 

subgen. Bulimella Pfeiffer. 

Bulimella Pfeiffer, Malak. Blatt. i (1854), p. 119 (as section oi Achatmella, first 
species A. rosea Swainson). 

(18) Achatinella {^Bulimella) abbreviata Reeve. 

Achatinella abbreviata Reeve, Conch. Icon. Achatinella, sp. 19; Newcomb, Ann. 

Lye. New York, vi. p. 317 [animal]. 
Achatinella bacca Reeve, loc. cit. sp. 45 ; Newcomb, loc. cit. p. 318 [animal]. 
Achatinella nivosa Newcomb, P. Zool. Soc. London, 1853 [1854], p. 132, pi. 

XXII. fig. 6. 
Achatinella [Achatinellastrum) Clementina Pfeiffer, P. Zool. Soc. London, 1855 

[Feb. 1856], p. 205. 

Hab. Oahu, Palolo and Konahuanui (Baldwin) ; Niu (Newcomb) ; Head of 
Kawailoa Gulch (Perkins). 

The specimens found are of a puzzling form, shewing links between Clementina and 

colorata. 

The animal, as described by Newcomb, seems to vary a good deal in colour. 

(19) Achatinella {^Btilimella) anipla Newcomb. 

Achatinella anipla Newcomb, P. Zool. Soc. London, 1853 [1854], p. 137, pi. 

XXII. fig. 19. 
Mr Baldwin considered this a synonym of A. colorata Rve. ; the only specimen I 
have seen is the type, which is somewhat injured, and I incline to place it near 
A. abbreviata Rve. 

Hab. Oahu, Koolau (Newcomb). 

40 — 2 



3o6 FAUNA HAWAIIENSIS 



(20) Achatinella {Bjilimella) bulimoides Swainson. 

AcJiatinella bulimoides Swainson, Brand's Journ. 1828, p. 85; Zool. Illustr. ser. 2, 
II. pi. cviii. fig. I ; Reeve, Conch. Icon. Achatinella, sp. 8 ; Heynemann, 
Malak. Blatt. xiv. (1867), p. 146, pi. i. fig. 2 [anatomy]. 

Achatinella obliqua Gulick, Ann. Lye. New York, vi. p. 245, pi. viii. fig. 63. 

Achatinella ooinorpha Gulick, t. c, p. 246, pi. viii. fig. 64. 

A. obliqua was united with this species by Newcomb ; Mr Baldwin, however, 
gives it as distinct. This latter view may be correct, but the two forms are very 
closely related. 

Hab. Oahu, Kahana (Gulick) ; Kawailoa (Baldwin). 



(21) Achatinella [Bulimella) byronii Wood. 

Helix byronii Wood, Index Test. Suppl. p. 22, pi. vii. fig. 30. 

Achatinella melanostoina Newcomb, P. Zool. Soc. London, 1853 [1854], p. 132, 

pi. xxii. fig. 7. 
Achatinella limbata Gulick, Ann. Lye. New York, vi. p. 252, pi. viii. fig. 70. 
Achatinella ma ho£-ani Gulick, t. c, p. 254, pi. viii. fig. 72. 
Achatinella pidcherrinia Swainson, Zool. 111. pi. cxxiii. fig. 2; Gwatkin, P. Ac. 

Philad. 1S95, P- 238 [radula]. 

Hab. Oahu, Ewa (Newcomb) ; Ahonui, Kalaikoa (Gulick) ; Panoa Valley, 
Halemano, and ridges between Opaeula and Kawailoa Gulches (Perkins). 



var. recta Newcomb. 

Achatinella recta Newcomb, P. Zool. Soc. London, 1853 [1854], p. 143, pi. xxiii. 

fig- 45- 
Bulimella multicolor Pfeiffer, op. cit. 1S55, p. 4, pi. xxx. fig. 11 \^pars, non 
fig. 1 1 a]. 

Hab. Oahu, Waialua (Newcomb) ; Halemano and Nuuanu Valley (Perkins). 



var. nympha Gulick. 

Achatinella nynpha Gulick, Ann. Lye. New York, vi. p. 251, pi. viii. fig. 69. 

Hab. Oahu, Ahonui, Wahiawa, Halemano, Kawailoa, Waimea (Gulick) ; Hale- 
mano (Perkins). 



MOLLUSC A 307 

The variation is, as usual in the group, very great. A long series collected by 
Mr Hutchison, added to those of Mr Perkins, has led me to be unable to form any 
definite break between the various described species which are here placed as varieties. 
A. pulcherriina appears to be a large race in which the colouring has been broken into 
bands. A. imdticolor and A. recta are, I think, only dwarf varieties. The sinistral 
shell figured by Pfeiffer {loc. cit. pi. xxx. fig. 1 1 a) as a variety of A. multicolor 
belongs really to A. ovifonnis. A. nympha seems a small, elongate, almost colourless 
variety, with a white lip. 

(22) Achatinella [Bulimella) decipiens Newcomb. 

Achatinella decipiens Newcomb, P. Zool. Soc. London, 1853 [1854], p. 153, 
pi. XXIV. fig. 68 ; Ann. Lye. New York, vi. p. 332 [animal]. 

Achatinella viridans Pfeiffer, Mai. Blatt. 1854, p. 121 [nee Mighels, fide New 
comb]. 

Achatinella planospira Pfeiffer, P. Zool. Soc. London, 1855, p. 3, pi. xxx. fig. 8. 

Achatinella herbacea Gulick, Ann. Lye. New York, vi. p. 233, pi. viii. fig. 52. 

Achatinella scitula Gulick, /. c. p. 241, pi. viii. fig. 61. 

Hab. Oahu, Kahana (Newcomb, Baldwin); Koolauloa (Hartman); Waimea, 
Kawailoa, Hakipu (Gulick). 

(23) Achatinella {Btiliviella) /aba Pfeiffer. 

Achatinella {Btdiinella) faba Pfeiffer, P. Zool. Soc. London, 1859, p. 30. 
Hab. Hawaiian Islands. 

I cannot trace this species in the Brit. Mus. ; it seems not to have been recognized 
by any recent author. 

(24) Achatinella [Buliniella) glabra Newcomb. 

Achatinella glabra Newcomb, P. Zool. Soc. London, 1853 [1854], p. 139, pi. xxii. 

% 25. 

Achatinella fricki Pfeiffer, op. cit. 1855, p. 3, pi. xxx. fig. 7. 

Achatinella platystyla Gulick, Ann. Lye. New York, vi. p. 196, pi. vr. fig. 25. 

Achatinella wheat leyi Newcomb, MS. 

Hab. Oahu, Kawailoa to Hauula (Baldwin); Koolau poko (Newcomb); Ka- 
waiawa (Hartman) ; Kawailoa (Guhck and Perkins). 

Only two dead specimens. I think A. fricki, which Newcomb placed with 
A. ovata, really belongs here ; fig. 7 a, however, belongs to A. ovata. The determina- 
tion of ^. wheatleyi is from specimens so named in the Brit. Mus. 



3o8 FAUNA HAWAIIENSIS 



(25) Achatinella {Bidinielld) elegans Newcomb. 

Achatinella elegans Newcomb, P. Zool. Soc. London, 1853 [1854], p. 149, 
pi. XXIV. fig. 57. 

Hab. Oahu, Hauula (Newcomb); Hauula and Kaipapau (Baldwin). 



(26) Achatinella [Buliiiiella) luteostonia Baldwin. 

Achatinella [Buliniella) luteostonia Baldwin, P. Ac. Philad. 1895, P- 217, pi. x. 
figs. 7, 8 [with a note on the animal]. 

Hab. Oahu, Palolo to Niu (Baldwin). 

(27) Achatinella [Btilimella) lymaniana Baldwin. 

Achatinella {^Btdimella) lymaniana Baldwin, P. Ac. Philad. 1895, P- 219, pi. x. 
figs. 12, 13. 

Hab. Oahu, Waianae mountains (Baldwin). 

(28) Achatinella {Bulimella) lyonsiana Baldwin. 

Achatinella iyBulimella) lyonsiana Baldwin, P. Ac. Philad. 1895, p. 218, pi. x. 
figs. 9 — 1 1 [with note on animal] ; Suter, t. c. p. 239, pi. xi. fig. 52 
[radula]. 

Hab. Oahu, Konahuanui mountain (Baldwin). 

(29) Achatinella {Bulimella^ ovata Newcomb. 

Achatinella ovata Newcomb, Ann. Lye. New York, vi. p. 22 [May, 1853]; T. c. 

p. 324 [animal]; P. Zool. Soc. London, 1853, p. 130, pi. xxii. fig. 2. 
Biili77iella Candida Pfeiffer, P. Zool. Soc. London, 1855, p. 2, pi. xxx. fig. 4. 
Achatinella phaeozona Gulick, Ann. Lye. New York, vi. p. 215, pi. vii. fig. 40. 
Achatinella spadicea Gulick, t. c. p. 247, pi. viii. fig. 65. 
Achatinella loi'ata Reeve, Conch. Icon. Achatinella, sp. 6 [nee Ferussae]. 

Hab. Oahu, Kahana, Waianae (Newcomb); Kawailoa (Baldwin); 2i% A. phaeo- 
zona, Keawaawa, Kailua, Olomana (Gulick) ; as A, spadicea, Kahana (Gulick) ; Hauula 
to Kahana (Baldwin). 

Gulick's two species are unknown to me. I follow Newcomb in placing them 
here ; Mr Baldwin has, however, given them rank as species. 



MOLL use A 309 



(30) Achatinella {Bulimella) oviformis (Newcomb) Pfeiffer. 

Achatinella oviformis Pfeiffer, P. Zool. Soc. London, 1855, p. 208. 
Achatinella multicolor Pfeiffer, t. c. p. 4, pi. xxx. fig. 11 a [nee fig. 11, which 
equals A. byronii, var.]. 

Hab. Oahu (various authors). 



{31) Achatinella {Btilimella) rosea Swainson. 

Achatinella bidimoides var. rosea Swainson, Brand's Journ. 1828, p. 85. 
Achatinella rosea Swainson, Zool. Illustr. sen 2, pi. cxxiii. fig. i ; Reeve, 

Conch. Icon. Achatinella, sp. 28 ; Newcomb, Ann. Lye. New York, vi. p. 309 

[animal]. 
Bulimella rosea Swainson, Hartman, P. Ac. Philad. 1888, p. 30, pi. i. fig. 4. 

A good series, including some varieties approaching A. ovata. 

Hab. Oahu, Wahiawa to Kawailoa (Baldwin); Waialua (Hartman); Halemano 
(Perkins). 



(32) Achatinella {^Bulimella) rottmda Gulick. 

Achatinella rotunda Gulick, Ann. Lye. New York, vi. p. 249, pi. viii. fig. 67. 
This form has, with much doubt, been allowed specific rank. In this I have 
followed Mr Baldwin ; Newcomb considered it a variety of A. ovata. 

Hab. Oahu, Kaawa and Kahana (Gulick) ; Head of Kawailoa (Perkins). 



(33) Achatinella {Bulimella) riigosa Neweomb. 
Achatinella ru^osa Newcomb, P. Zool. Soc. London, 1853 [1854], p. 138, 

pi. XXII. fig. 2 2. 

Achatinella corrugata Gulick, Ann. Lye. New York, vi. p. 248, pi. viii. fig. 66. 

Achatinella torrida Gulick, t. c. p. 250, pi. viii. fig. 68. 

I strongly suspect that this will prove to be only a roughened form of A. byronii 
Wood. 

Hab. Oahu, Ewa (Newcomb) : as A. corrugata, Hakipu (Gulick) ; Kahana 
(Baldwin) : as A. torrida, Kahana, Kaawa, Waikane, Waiolu (Gulick). 



3IO FAUNA HAWAIIENSIS 



(34) Achatinella {^Bidimella) sowerbyana Pfeiffer. 

BtLlimella sowerbyana Pfeiffer, P. Zool. Soc. London, 1855, p. 4, pi. xxx. 
fig. 14. 



var. ficscobasis Smith. 

Btilimelia fuscobasis Smith, P. Zool. Soc. London, 1873, p. "]•], \A. ix. fig. 15. 

I think Mr Smith's species is only a colour variety. 

Hab. Oahu (type form, authors) ; Mount Kaala (variety, Smith). 

(35) Achatinella [Bulimella) taeniolata Pfeiffer. 

Achatinella taeniolata Pfeiffer, P. Zool. Soc. London, 1846, p. 38; Reeve, Conch. 

Icon. Achatinella, sp. 7, 
Achatinella rubiginosa Newcomb, P. Zool. Soc. London, 1854 [1855], p. 154, 

pi. XXIV. fig. 69. 
Bulimella macrostoma Pfeiffer, P. Zool. Soc. London, 1855, p. 2, pi. xxx. 

fig. 6. 

Newcomb was of opinion that A. macrostoma was identical with A. riitila, but, 
after examining the type, I prefer to place it here. 

Hab. Oahu, Palolo (Newcomb); Niu to Palolo (Baldwin). 

(36) Achatinella [Bulimella) vidua Pfeiffer. 

Btiltmella vidiia Pfeiffer, P. Zool. Soc. London, 1855, p. 3, pi. xxx. fig. 10. 

Newcomb placed this as a synonym of A. ovata ; Mr Baldwin regarded it as a 
distinct species. The columellar plait is very small in the specimens in the Brit. Mus. 

Hab. Oahu (Baldwin, &c.). 

(37) Achatinella [Bulimella) viridans Mighels. 

Achatinella viridans Mighels, P. Boston Soc. 11. (1845), P- -O I Newcomb, P. Zool. 

Soc. London, 1854, p. 310 [animal]. 
Achatinella radiata Pfeiffer, P. Zool. Soc. London, 1845 [1846], p. 89; Reeve, 

Conch. Icon. Achatinella, sp. 35. 



MOLLUSC A 311 

Achatinella subviretis Newcomb, P. Zool. Soc. London, 1853 [1854], p. 136, 

pi. XXII. fig. 18. 
Achatinella rutila Newcomb, t. c. p. 138, pi. xxii. fig. 21 ; Op. cit. 1854, p. 310 

[animal] ; Ann. Lye. New York, vi. p. 326 [animal]. 

Hab. Oahu, Niu (Newcomb) ; Palolo, Niu, Konahuanui (Hartman) ; Nuuanu to 
Waialae (Baldwin) ; Nuuanu, Waialae (Perkins). 



subgen. Partulina Pfeiffer. 

Partulina Pfeiffer, Malak. Blatt. i. 1854, p. 114. 

Pfeiffer had no fixed type for his section, but the species all belong to one group, 
and I would suggest that his first-named, A. virgtilata Migh., be treated as the type. 

Pease, in his review of the genus in 1869, did not alter the grouping, so far as 
regards Partulina. 

Mons. Ancey has recently (P. Malac. Soc. London, in. 1899, p. 270) proposed to 
place P. pliysa Newc, and its allies, in a new subgenus Baldwinia. 

(38) Achatinella (Partulina) anceyana Baldwin. 

Achatinella {Partulina) anceyana Baldwin, P. Ac Philad. 1895, p. 223, pi. x. 

fig. 16 ; Gwatkin, t. c. p. 238 [radula]. 
Hab. Maui, Makawao (Baldwin). 

(39) Achatinella {Partulina) aptycka Pfeiffer. 
Achatinella {Newcombia) aptycha Pfeiffer, P. Zool. Soc. London, 1855 [March], 

p. I, pi. XXX. fig. I. 

Hab. Hawaiian Islands. Probably from Maui. 

(40) Achatinella {Partulina) compta Pease. 

Partulina compta Pease, J. Conchyl. xvii. 1869, p. 175. 

Curiously enough some specimens collected on Maui, and sent to me by Mr 
Baldwin, exactly agree with a specimen from Molokai presented by Pease to the 
British Museum under this name. 

Hab. Molokai (Pease) ; Kawela (Baldwin)— Maui (Baldwin). 

F. H. II. 41 



312 FAUNA HAWAIIENSIS 

(41) Achatinella [Pariulina) confusa nom. nov. 

Achatinella physa Newcomb, P. Boston Soc. v. 1855, p. 218 ; Amer. J. Conch. 11. 

1866, p. 214, pi. XIII. fig. 10. 
Achatinella {Partulind) physa Newcomb, Baldwin, P. Ac. Philad. 1895, p. 225 

[animal]. 
Nee A. physa Newcomb, 1854, q. v. (p. 316). 

An inspection of the figures and descriptions given by Newcomb (P. Zool. Soc. 
London, 1853, p. 152, pi. xxiv. fig. 64, and as given above) will, I think, show that 
he was confusing two species, under the belief that the first description related only 
to a young specimen. It therefore becomes necessary to restrict his name to the 
species he first referred to, which unfortunately appears to be the same as A. hawaii- 
ensis Baldwin, and to rename the other form, which, it is to be regretted, is the species 
universally known as A. physa. It may be noted that the habitat originally given by 
Newcomb agrees with that of Hamakua given by Mr Baldwin for his A. hawaiiensis, 
whilst Kohala is a different, but adjoining, district. 

Hab. Hawaii, Kohala (Newcomb). 

(42) Achatinella [Parttilina) crassa Newcomb. 

Achatinella crassa Newcomb, P. Zool. Soc. London, 1853 [1854], p. 155, pi. xxiv. 
fig. 71. 

H.^B. Lanai (Newcomb) ; near Koele (Perkins). 

(43) Achatinella [Partulina) dolei Baldwin. 

Achatinella i^PartiUind) dolei Baldwin, P. Ac. Philad. 1895, P- 221, pi. x. figs. 17, 
18 ; Suter, t. c. p. 238, pi. xi. fig. 55 [radula]. 

Belongs to the group of A. tappaniana C. B. Ad. ; specimens, precisely similar to 
some kindly sent me by Mr Baldwin, were identified by Mr Gulick as a variety of his 
A.fasciata ( = tappaniana). 

Hab. Maui, Honomanu (Baldwin). 

(44) Achatinella {Partiilina) dubia Newcomb. 

Achatinella dubia Newcomb, Ann. Lye. New York, vi. p. 23 (May, 1853) ; P. Zool. 
Soc. London, 1853 [1854], p. 152, pi. xxiv. fig. 65. 

Hab. Oahu, among stones, and Waianae on bushes (Newcomb) ; Makaha Valley, 
Waianae Mts (Perkins). 



MOLLUSC A 313 

(45) Achatinella {Partulina) dwightii Newcomb. 

Achatiiiella divightii Newcomb, Ann. Lye. New York, vi. p. 145 (Oct. 1855); 
Amen J. Conch, n. p. 213, pi. xiii. fig. 9; Gwatkin, P. Ac. Philad. 1895, 
p. 238 [radula]. 

Closely related, apparently, to some of the varieties o^ A. redfieldi Newc. 

Hab. Molokai, Kamalo (Baldwin) ; Mountains (Perkins). 

(46) Achatinella [Parhilina) fusoidea Newcomb. 

Achatinella fusoidea Newcomb, Ann. Lye. New York, vi. p. 144 (Oct. 1855); Amer. 
J. Conch. 11. 1866, p. 213, pi. XIII. fig. 8. 

Hab. Maui, Haleakala (Newcomb). 

(47) Achatinella [Partuliua) goiildi Newcomb. 

Achatinella gottldi Newcomb, Ann. Lye. New York, vi. p. 21 (May 1853); P. Zool. 

Soc. London, 1853 [1854], p. 129, pi. xxii. fig. i. 
Achatinella talpina Gulick, Ann. Lye. New York, vi. p. 212, pi. vii. fig. 38 (Dee. 

1856). 
Achatinella myrrhea Gulick, Pfeiffer, Mon. Helic. Viv. iv. p. 517. 

Hab. Maui, on Tutui trees, Wailuku Valley (Newcomb) ; Wailuku (Gulick). 

(48) Achatinella (^Partiilina) g7'isea Newcomb. 

Achatinella grisea Newcomb, P. Zool. Soc. London, 1853 [1854], p. 153, pi. xxiv. 

fig. 66. 
Hab. Maui, Makawao (Newcomb, &c.). 

(49) Achatinella {Partulina) hayseldeni Baldwin. 

Partulina hayseldeni Baldwin, Nautilus, x. p. 31, July 1896. 

Plate XL fig. 2. 

Belongs to the group o{ A. variabilis Newc. 

Hab. Lanai (Baldwin) ; Lanaihale, near highest point of Mountains (Perkins). 

(50) Achatinella [Partulina) horneri Baldwin. 

Achatinella {Partulina) horneri Baldwin, P. Ac. Philad. 1895, p. 224, pi. x. figs. 
20, 21, 22 ; Gwatkin, t. c. p. 238 [radula]. 

Hab. Hawaii, Hamakua (Baldwin). 

41—2 



314 FAUNA HAWAIIENSIS 

(51) Achatinella [Pariu/ina) lignaria Gulick. 

Achatinella lignaria Gulick, Ann. Lye. New York, vi. p. 209, pi. vii. fig. 35 (Dec. 

1856). 
Hab. Maui, Wailuku (Gulick). 

van crocea Gulick. 

Achatinella crocea Gulick, t. c. p. 21 i, pi. vii. fig. 36 (Dec. 1856). 

I think A. crocea is only a variety; both were placed by Newcomb as synonyms 
of his A. terebra. 

Hab. Maui, Waihee (Gulick). 

(52) Achatinella [Partulina) jnarmorata Gould. 

Achatinella marniorata Gould, P. Boston Soc. 11. p. 200 (1847); U. S. Explor. 
Exped. Moll. fig. 94; Newcomb, Ann. Lye. New York, vi. p. 311 [animal]; 
Gwatkin, P. Ac. Philad. 1895, P- 238 [radula]. 
Achatinella adamsi Newcomb, Ann. Lye. New York, vi. p. 19 (May, 1853); 

P. Zool. Soc. London, 1853 [1854], p. 137, pi. xxii. fig. 20 (as A. adamsii). 
Achatinella induta Gulick, Ann. Lye. New York, vi. p. 207, pi. vii. fig. 34 (Dec. 

1856). 
The synonymy of this species is difficult ; Newcomb united two other forms de- 
scribed by Gulick from a different district of Maui ; Mr Baldwin on the other hand 
regards them as species. For the present I have left them, with some hesitation, specific 
rank ; they are A. ustulata and A. pltimbea. 

Hab. Maui, Haleakala (Gould) ; Makawao (Newcomb, Baldwin) ; Wailuku 
(Gulick). 

(53) Achatinella {Partulina) mighelsiana Pfeiffer. 

Achatinella mighelsiana Pfeiffer, Mon. Hel. Viv. 11. p. 238 ; Newcomb, Ann. Lye. 
New York, vi. p. 319 [animal]; Gwatkin, P. Ac. Philad. 1895, p. 238 [radula]; 
Reeve, Conch. Icon. Achatinella, sp. 40. 

The typical form is a whitish shell with a single black band at the periphery ; this 
single band is occasionally split into two smaller ones. Some lovely varieties were 
collected by Mr Perkins, which may be tabulated as follows : 

(a) White and bandless. 

(/8) Bandless, of a rich orange hue with strigations of a slightly darker shade, 
tubercle white. 



MOLLUSC A 315 

(y) One-banded, the whitish shell tinted with yellow, ashy, or slaty strigations. 
A few are white above the band, yellowish below, and show traces of a second band in 
the umbilical area. 

(S) Two- and even three-banded, ground-colouring white, tinted faintly with 
ashy strigations, shell not quite so attenuate. 

Hab. Molokai, Kalae (Baldwin) ; the Mountains (Perkins). 

(54) Achatinella l^Parhtlina) 7)iorbida Pfeiffer. 

Achatinella [Bu/imella) morbida Pfeiffer, P. Zool. Soc. London, 1859, p. 30. 

Hab. ?Oahu. 

The only authority I am aware of for the exact habitat is Mr Baldwin, who gives 
Oahu, but he marks it as one of the species he has not seen. 



(55) Achatinella i^Parhilina) niucida Baldwin. 

Achatinella [Pai'tiiliiui) mucida Baldwin, P. Ac. Philad. 1S95, P- 222, pi. x. fig. 23. 

A series of about 60 specimens. It is generally of an ashy colour with a dark zone 
at the periphery ; smaller colour lines are also present in most specimens. The brown 
stain at the base of the columellar plait is also noteworthy. 

Hab. Molokai, Makakupaia (Baldwin) ; Makakupaia, and Mountains of Molokai 
(Perkins). 

(56) Achatinella [Partulina) nivea Baldwin. 

Achatinella [Partulina) nivea Baldwin, P. Ac. Philad. 1895, p. 222, pi. x. fig. 19. 
Hab. Maui, Makawao to Huelo (Baldwin). 

(57) Achatinella [Partiilina) perdix Reeve. 

Achatinella pei'dix Reeve, Conch. Icon. Achatinella, sp. 43 (1850); Newcomb, 
Ann. Lye. New York, vi. p. 317 [animal]; Gwatkin, P. Ac. Philad. 1895, 
p. 238 [radula]. 

Achatinella iindosa Gulick, Ann. Lye. New York, vi. p. 205, pi. vii. fig. n (Dec. 
1856). 

A. undosa was wrongly placed by Clessin (Norn. Helic. Viv. p. 305) as a synonym 
of A. radiata Gould. 

Hab. Maui, Lahaina (Baldwin); Olinda at 4000 ft. (Perkins). 



3i6 FAUNA HAWAIIENSIS 



(58) Achatinella {Partulina) physa Newcomb. 

Achatinella physa Newcomb, P. Zool. Soc. London, 1853 [1854], p. 152, pi. xxiv. 

fig. 64. 
Achatinella [Partulina) liawaiiensis Baldwin, P. Ac. Philad. 1895, p. 225, pi. x. 

figs. 24 — -26 ; Gwatkin, t. c. p. 238 [radula]. 
Nee A. physa Newc. subsequently. 

See for notes on the synonymy under A. confiisa Sykes. 
Hab. Hawaii, Mauna Kea (Newcomb) ; Hamakua (Baldwin). 



(59) Achatinella [Partulina) plumbea Gulick. 

Achatinella plumbea Gulick, Ann. Lye. New York, vi. p. 213, pi. vii. fig. 39. 
Hab. Maui, Kula (Gulick). 

(60) Achatinella (Partulina) porcellana Newcomb. 

Achatinella porcellana Newcomb, P. Zool. Soc. London, 1853 [1854], p. 146, 
pi. xxiii. fig. 47. 

In appearance recalling a dwarf specimen of A. terebra Newc. of W. Maui ; only 
known to me from the type. 

Hab. E, Maui (Newcomb). 

(61) Achatinella [Partulina) proxima Pease. 

Helicter proxivins Pease, P. Zool. Soc. London, 1862, p. 6. 

Partulina proxima Pease, Hartman, P. Ac. Philad. 1888, p. 27, pi. i. figs, i, 2. 

Achatinella proxima Pease, Gwatkin, 1. c. 1895, p. 238 [radula]. 

A fine series. A variety is interesting as showing a link towards A. theodorei 
Baldwin ; it is much more slender and smaller than the typical form, generally lighter 
in colour, and the colour-markings are much finer in pattern. It was found with the 
typical form. 

Hab. Molokai, Waikolu (Baldwin) ; Kahanui, and mountains of Molokai 
(Perkins). 



MOLLUSCA 317 



(62) Achatviella {Pariulvia) pyramidalis Gulick. 

Achatinella pyramidalis Gulick, Ann. Lye. New York, vi. p. 204, pi. vii. fig. 32 
(Dec. 1856). 

Newcomb regarded this as a variety of A. perdix Reeve ; not having seen speci- 
mens which unite them I have left it as a species. Clessin (Norn. Helic. Viv. p. 
306) placed it — erroneously — under A. mannorata Gould. 

Hab. Maui, Lahaina (Gulick) ; Huelo (Baldwin); Waihee (Perkins). 



(63) Achatinella {Partulina) radiata Gould. 

Achatinella radiata Gould, P. Boston Soc. 11. 1845, P- -7- 
Btdinms gouldiV{€\^(tr, Zeitsch. fiir Malak. 1846, p. 116. 
Partula deiisilineata Reeve, Conch. Icon. Partttla, sp. 9. 

Hab. ? Maui (Baldwin). 

The specimens in the British Museum are labelled "Oahu", but probably this is 
erroneous and Maui is the correct habitat. 



(64) Achatinella [Parttilina) redfieldi Newcomb. 

Achatinella redfieldi Newcomb, Ann. Lye. New York, vi. (May 1853), p. 22 ; t. c. 
p. 325 [animal]; P. Zool. Soc. London, 1853 [1854], p. 131, pi. xxii. fig. 5 ; 
Gwatkin, P. Ac. Philad. 1895, P- 238 [radula]. 

The long series collected by Mr Perkins has given me considerable difficulty. 
Newcomb originally gave both Maui arid Molokai, Clessin (Nom. Helic. Viv. p. 306) 
gave Molokai and Kauai (the latter being obviously wrong), and Mr Baldwin gives 
Mapulehu, Molokai. I think Maui was a slip, due to confusion with the very closely 
allied A. splendida, and that A. redfieldi is really a Molokai shell. Next arises the 
question of what the typical form may be ; Newcomb states that the shell is either plain 
or banded on the third whorl only, while he gives six as the number of whorls, the shell 
figured being banded (as A. splendida) on all the whorls. The forms I refer to this 
species are : 

a. Typical (Plate XL fig. 15). Varies from nearly white to chestnut, sometimes 
being particoloured. 

Hab. Makakupaia, Molokai (Perkins). 



3i8 FAUNA HAWAIIENSIS 

yS. Light to dark fawn colour, banded with brown, the Hp being sometimes white. 
This is the form figured by Newcomb. 

Hab. Molokai, towards or above Kamalo (Perkins). 

y. Lip white, shell chestnut, a white band at the periphery and often a smaller 
one above it, upper whorls finely tessellated. (Plate XL fig. i6.) 

Hab. Molokai, Makakupaia and Kamalo (Perkins). 



(65) Achatitiella {^Parhtlina) ru/a Newcomb. 

Achatinella rtifa Newcomb, Ann. Lye. New York, vi. p. 21 (May 1853); t. c. 
p. 324 [animal]; P. Zool. Soc. London, 1853 [1854], p. 130, pi. xxii. fig. 3. 

Hab. Molokai, Kalae (Baldwin) ; mountains (Perkins). 

Dr Hartman gave, erroneously, Maui for this shell. The figure is not good, being 
too elongate and too highly coloured : a pale variety exists. 



(66) Achatinella i^Partulina) splendida Newcomb. 

Achatinella splendida Newcomb, Ann. Lye. New York, vi. p. 20 (May, 1853) ; P. 

Zool. Soc. London, 1853 [1854], p. 131, pi. xxii. fig. 4. 
Achatinella baileyana Gulick, Ann. Lye. New York, vi. p. 202, pi. vii. fig. 31 

(1856). 
Achatinella solida Gulick, Pfeiffer, Mon. Helic. Viv. iv. p. 516. 

Hab. Maui, Wailuku (Newcomb, &c.) ; Lahaina and Wailuku (Baldwin). 



(67) Achatinella [Partulina) tappaniana C. B. Adams. 

Achatinella tappaniana C. B. Adams, Contrib. to Conch, p. 126 (1850) [with var. 

dtibiosa\. 
Achatinella ebnrnea Gulick, Ann. Lye. New York, vi. p. 199, pi. vi. fig 28 ; 

Gwatkin, P. Ac. Philad. 1895, p. 238 [radula]. 
Achatinella ampulla Gulick, t. c. p. 200, pi. vii. fig. 29. 
Achatinella fasciata Gulick, t. c. p. 201, pi. vii. fig. 30. 
Achatinella tuba Gulick, Pfeiffer, Mon. Helic. Viv. iv. p. 523. 

Hab. Maui, (as A. tappaniana^ Lahaina (Baldwin); (as A. eburned) Honuaula 
(Gulick) ; (as A. ainpitlla and A. fasciata) Honukawai (Gulick). 



MOLLUSC A 319 



(68) Achatinella [Partulina) terebra Newcomb. 

Achatinella terebra Newcomb, P. Zool. Soc. London, 1853 [1854], p. 144, pi. 

XXIII. fig. 40. 

Btdimella attenuata Pfeiffer, P. Zool. Soc. London, 1855 [March], p. 4, pi. xxx. 

fig. 12. 
Achatinella corusca Gulick, Pfeiffer, Mon. Helic. Viv. iv. p. 525. 
Achatinella perforata Gulick, Pfeiffer, pag. cit. 

Hab. Maui; W. Maui (Newcomb) ; Wailuku (Hartman) ; Honokowai (Baldwin). 

(69) Achatinella [Partulina) tessellata Newcomb. 

Achatinella tessellata Newcomb, Ann. Lye. New York, vi. (May, 1853), p. 19; 
t. c. p. 327 [animal] ; P. Zool. Soc. London, 1853 [1854], p. 139, pi. xxiii. fig. 
26; Gwatkin, P. Ac. Philad. 1895, p. 238 [radula]. 

A very fine series. The forms found at Pelekunu are generally dextral and of 
large size ; recalling in shape and colouring A. virgiilata, but as they possess the 
mottled colouring of the earlier whorls, so characteristic of the present species, I have 
placed them here. 

Hab. Molokai, Kalae to Waikolu (Baldwin) ; Pelekunu, Makakupaia, Kahanui, 
&c. (Perkins). 

(70) Achatinella [Partulina) ustiilata Gulick. 

Achatinella ustiilata Gulick, Ann. Lye. New York, vi. p. 211, pi. vii. fig. 2,7- 
Nee A. ustulata Newcomb MS.; fide Pfeiffer, Malak. Blatt. i. p. 136 {=A. 

colorata Reeve). 
Hab. Maui, Beautiful Valley (Gulick) ; Lahaina (Baldwin). 

(71) Achatinella {Partulina) variabilis Newc. 

Achatinella variabilis Newcomb, P. Zool. Soc. London, 1853 [1S54], p. 154, pi. 

XXIV. fig. 70. 

Achatinella fulva (Newcomb) Pfeifter, loc. cit. 1855 [1856], p. 208. 
Achatinella lactea Gulick, Ann. Lye. New York, vi. 1858, p. 19S. pi. vi. fig. 27 
[bad]. 

Hab. Lanai (Newcomb, &c.) ; windward side on ridges facing Maui, above 
Waiapaa, behind Koele, and Lanaihale (Perkins). 

F. H. II. 42 



320 FAUNA HAWAIIENSIS 

van semicarinata Newc. 

Achatinella semicarinata Newcomb, P. Zool. Soc. London, 1853 [1854], p. 156, 
pi. XXIV. fig. 76. 

From an examination of the very fine series collected by Mr Perkins, I think 
Newcomb was quite right in placing A. fulva and A. lactea in the synonymy. The 
former is a straw-coloured form without banding and the latter a white form with a 
reddish-brown stain in the interior of the aperture. Mr Baldwin remarks that A. 
variabilis is ' invariably dextral,' but sinistral specimens, typical in every other respect, 
were found by Mr Perkins. In placing A. semicarinata as a variety I have been 
guided by the great difficulty I found in endeavouring to separate this form from A. 
fulva, the type specimens of which shew traces of the carina. 

Mr Perkins remarks that ' the broader form with ridge more raised ' is ' from 
higher elevations ' : it appears to be gradually replaced by the form fulva at lower 
altitudes and this latter shades into A. variabilis (typical). 

Hab. Lanai (Newcomb, &c.) ; mountains (Perkins). 

(72) Achatinella [Parttilina] virgulata Mighels. 

Partiila virgjilata Mighels, P. Boston Soc. 11. 1845, P- 20. 

Achatinella virgulata Mighels, Reeve, Conch. Icon. Achatinella, sp. 3 ; Newcomb, 

Ann. Lye. New York, vi. p. 312 [animal]; Gwatkin, P. Ac. Philad. 1895, 

p. 238 [radulaj. 
Bulimtis rohri Pfeiffer, Zeitsch. f. Malak. 1846, p. 115. 
Bulimies insignis Mighels, Reeve, Conch. Icon. Achatinella, sp. 3. 

Hab. Molokai, Kaluaaha to Halawa (Baldwin); Mapulehu and mountains 
(Perkins). 

It is a very variable species and the following, which I take to be a variety, is 
perhaps worthy of note. 

var. a. Either entirely white or slightly tinted with brown on the last whorl ; 
mouth varying from dusky to white ; the spiral black line on the upper whorls either 
present or absent. 

Hab. Molokai, Pelekunu (Perkins). 

subgen. Achatinellastrum Pfeiffer. 

Achatinellastrum Pfr., Malak. Blatt. i. (1854), p. 133 (first species A. venulata 
Newc). 



MOLLUSC A 321 

{T^ Achatinella {Achatinellastrum) augnsta Smith. 

Achatinella augtista Smith, P. Zool. Soc. London, 1S73, p. 74, pi. ix. fig. 7. 
Dr Hartman referred this shell, as A. angusta, to A. fidgens Newc. 
Hab. Oahu, Waialae, Waialupe, Palolo (Smith). 

(74) Achatinella {Achatincllastrum) bclla Reeve. 

Achatinella bella Reeve, Conch. Icon. Achatinella, sp. 17; Newcomb, Ann. 
Lye. New York, vi. p. 316 [animal]; Gwatkin, P. Ac. Philad. 1895, p. 238 
[radula]. 

Pease (P. Zool. Soc. 1869, p. 652) united the species, I think erroneously, with 
A. polita Newc. 

Hab. Molokai (various authors and Perkins) ; Kalae to Waikolu (Baldwin). 

(75) Achatinella {Achatincllastj-mn) bilineata Reeve. 

Achatinella bilineata Reeve, Conch. Icon. Achatinella, sp. 22. 

Achatinella johnsoni 'H&v;comh, P. Zool. Soc. London, 1854, p. 147, pi. xxiii. fig. 50. 

Achatinella aphistre Newcomb, t. c. p. 147, pi. xxiii. fig. 51. 

Hab. Oahu, Koolau (Newcomb) ; Manoa to Nuuanu (Baldwin). 

(76) Achatinella {Achatinellastruni) buddii Newcomb. 

Achatinella buddii Newcomb, P. Zool. Soc. London, 1853 [1854], p. 155, pi. xxiv. 

fig- 11- 
Achatiyiella pexa Gulick, Ann. Lye. New York, vi. p. 197, pi. vi. fig. 26. 

Achatinella pliiniata Gulick, t. c. p. 217, pi. vii. fig. 41. 

Achatinella casia Gulick, t. c. p. 234, pi. viii. fig. 53. 

Achatinella fusco zona Smith, P. Zool. Soc. London, 1873, p. 76, pi. ix. fig. 9. 

I follow Mr Baldwin in uniting Mr Smith's species, with which I am unacquainted. 
Dr Hartman (P. Ac. Philad. 18S8) places it (on p. 32) amongst the synonyms of ^. 
bnddii ; possibly this may be a slip as further on (p. ^^t^ he leaves it specific rank, 
remarking ' this may be a good species, though it approaches very near to A. fusco- 
lineata. Smith,' a comparison which appears to me inaccurate. 

Hab. Oahu, Palolo (Newcomb); Niu, Wailupe, Waialae, Palolo, Kailua, and 
Waimea (Gulick) ; Makiki, Palolo (Smith). 

42 — 2 



322 FAUNA HAWAIIENSIS 

{'jj) Achatinella [Achatinellastrum) casta Newcomb, 

Achatinella casta Newcomb, P. Zool. Soc. London, 1853, p. 134, pi. xxii. fig. 12. 
Achatinella juncea Gulick, Ann. Lye. New York, vi. p. 230, pi. vii. fig. 49. 
Achatinella cognata Gulick, t. c. p. 240, pi. viii. fig. 60. 

A. cognata is only known to me from the description : I incline to think Newcomb 
was right in suppressing it as a species ; Mr Baldwin, however, considers it distinct. 

Hab. Oahu, Ewa (Newcomb, Baldwin) ; Kalaikoa, Wahiawa, Halemano, Haiki- 
puu, and Waikane (Gulick) ; above Ewa (Perkins). 

(78) Achatinella [Achatinellastrum) cervina Gulick. 

Achatinella cervina Gulick, Ann. Lye. New York, vi. p. 241, pi. viii. fig. 62. 

Newcomb placed it as a variety of A. ovata ; Mr Baldwin, on the other hand, gives 
it rank as a species and places it in Achatinellastruni. If the specimens in the Brit. 
Mus. are correctly identified, it is very close to A. buddii Newc. 

Hab. Oahu, Kahana (Gulick). 

(79) Achatinella [Achatinel/astriiin) colorata Rve. 

Achatinella colorata Reeve, Conch. Icon. Achatinella, sp. 18 ; Newcomb, Ann. 
Lye. New York, vi. p. 316 [animal]. 

Hab. Oahu, Ahuimanu (Hartman) ; Kalihi (Baldwin). 

(80) Achatinella {Achatinellastruvi) concolor Smith. 

Achatinella concolor Smith, P. Zool. Soc. London, 1873, p. 75, pi. ix. fig. i. 
Dr Hartman considered it to be a form of ^. colorata Rve. 
Hab. Oahu, Ewa (Smith). 

(81) Achatinella {^Achatinellastrum) ciicuniis Gulick. 

Achatinella cucumis Gulick, Ann. Lye. New York, vi. p. 225, pi. vii. fig. 45. 

Hab. Oahu, Kalihi (Gulick); Kalihi to Moanalua (Baldwin); Kaliua (sic) 
(Hartman). 



MOLLUSCA 



(82) Achatinella iyAchatinellastrjim) cnneus Pfeiffer. 



323 



Achatinella (^Achalinellastrzmt) cuneus Pfeiffer, P. Zool. Soc. London, 1855, 
p. 205. 

Newcomb considered this a form of A. decipiens; Dr Hartman appears to have 
been in some confusion, as he placed it (P. Ac. Philad. 1888) at p. 29 under A. decipiens, 
and at p. 30 under A. viridans. I have seen a long and characteristic series found on 
the Island of Oahu by Mr Hutchison. 

Hab. Oahu (authors) ; Halawa (Baldwin) ; mountains behind Ewa (Perkins). 

(83) Achatinella [Achatinellastrum) curta Newc. 

Achatinella airta Newcomb, P. Zool. Soc. London, 1853, p. 144, pi. xxiii. fig. 43. 
Achatinella Jindiilata Newcomb, P. Boston Soc. v. (1855), p. 219; Amer. J. 

Conch. II. (1866), p. 216, pi. XIII. fig. 15; Pfeiffer, P. Zool. Soc. London, 

1855, p. 208. 
Achatinella dimorpha Gulick, Ann. Lye. New York, vi. p. 236, pi. viii. fig. 56. 
. Achatinella albescens Gulick, t. c. p. 237, pi. viii. fig. 57. 
Achatinella contracta Gulick, t. c. p. 239, pi. viii. fig. 59. 
Achatinella rhodoraphe Smith, P. Zool. Soc. London, 1873, p. 75, pi. ix. fig. 10; 

Gwatkin, P. Ac. Philad. 1895, p. 238 [radula]. 
Achatinella pyg-ntcea Smith, P. Zool. Soc. London, 1873, p. 75, pi. ix. fig. 11. 

• Hab. Oahu, Waialua (Newcomb); various localities (Gulick); Halemano, 
Waipio, &c. (Smith) ; between Kawailoa and Waala gulches, generally between 
Kawailoa and Halemano, Waimea (Perkins). 

(84) Achatinella {Achatitiellastrtim) delta Gulick. 

Achatinella delta Gulick, Ann. Lye. New York, vi. p. 231, pi. viii. fig, 50. 
Newcomb considered A. delta to be a more banded variety of A. airta Newc. ; 
from the material I have seen I incline, with doubt, to leave them distinct. 
Hab. Oahu, Kalaikoa, Halemano, &c. (Gulick). 

(85) Achatinella {Achatincllastritm) diluta Smith. 

Achatinella dilnta Smith, P. Zool. Soc. London, 1873, p. 75, pi. ix. fig. 14. 
It is near to, but seems distinct from, A. ligata Smith, with which Dr Hartman 
placed it. 

Hab. Oahu, probably (Smith). 



324 FAUNA HAWAIIENSIS 

(86) Achatinella [Achatinellastrum) ernestina Baldwin. 

Achatinella [Achatinellastrtcni) ernestina Baldwin, P. Ac. Philad. 1895, p. 217, 
pi. X. figs. 5, 6 [animal described]. 

Hab. Oahu, Nuuanu Valley (Baldwin). 

(87) Achatinella {A chatinellastrum) formosa Gulick. 

Achatinella for^nosa Gulick, Ann. Lye. New York, vi. p. 235, pi. viii. fig. 55. 
Hab. Oahu, Waimea (Gulick). 

(88) Achatinella [Achatinellastrum) fulgens Newc. 

Achatinella fulgens Newcomb, P. Zool. Soc. London, 1853 [1854], p. 131, pi. xxii. 

fig. 24. 
Hab. Oahu, Niu (Newcomb) ; Waialua, south-east end (Hartman). 

(89) Achatinella [Achatinellastfuin) germana Newcomb. 

Achatinella germana Newcomb, P. Zool. Soc. London, 1853 [1854], p. 151, pi. 
XXIV. fig. 61. 

Hab. Maui, Makawao (Newcomb). 

(90) Achatinella i^Achatinellastruni) jttddii Baldwin. 

Achatinella i^Achatinellastrmn) jiiddii Baldwin, P. Ac. Philad. 1895, P- 216, pi. x. 

figs. 3- 4- 
Hab. Oahu, Halawa (Baldwin). 

(91) Achatinella [Achatiucllastntin) leluiiensis Smith. 

Achatinella lehuiensis Smith, P. Zool. Soc. London, 1873, P- 7^' P'- ^'^- fig- 4- 

I have not seen the species, but it appears from the figure to be near A. zonata 

Gulick ; Dr Hartman has suggested that it is a form of A. multicolor Pfr. 

( = ovifortnis Pfr. ). 

Hab. Oahu, Lehui (Smith). 



MOLLUSC A 325 

(92) Achatinella {Achatinellastrum) ligata Smith. 

Achatinella ligata Smith, P. Zool. Soc. London, 1873, p. 76, pi. ix. fig. 13. 
Achatinella bellula Smith, t. c. p. 77, pi. ix. fig. 8. 

I fancy these two forms are only varieties of one species ; they approach A. 
nympha Gulick. 

Hab. Oahu, Waimolu (Smith) ; Panoa and Nuuanu (Baldwin) ; ridges round 
Nuuanu, Waimea, and beyond head of Panoa Valley (Perkins). 

(93) Achatinella {^Achatinellastrum) livida Swainson. 

Achatinella livida Swainson, Zool. 111. pi. cviii. fig. 2. 

Achatinella emmersonii Newcomb, P. Zool. Soc. London, 1853 [1854], p. 156, pi. 

XXIV. fig. 74. 
Achatinella viridans Reeve, Conch. Icon. Achatinella, sp. 25 [nee Mighels]. 
Achatinella reevei C. B. Adams, Contrib. to Conch. 1850, p. 128. 
Achatinella consanguinea Smith, P. Zool. Soc. London, 1873, p. ']i, pi. ix. fig. 3. 
Nee A. livida Pfeiffer, P. Zool. Soc. London, 1845, p. 89 [ = ^. vnlpina Per.]. 
According to Dr Hartman, Mr Smith's species is probably a variety of A. 
colorata; from the specimens I have seen, I think it rather belongs here. 
Hab. Oahu, Waialua (Newcomb, Baldwin); Ahuimanu (Smith). 

(94) Achatinella {Ac hat inc I last rum) longispira Smith. 

Achatinella longispira Smith, P. Zool. Soc. London, 1873, p. Ti^ pi. ix. fig. 5. 

Placed by Dr Hartman as a synonym of A. viUpina, but the present species is 
much more slender in form; I should be inclined rather to refer it to the group of 
A. olivacea. 

Hab. Oahu, Halawa, Ahuimanu (?) (Smith). 

(95) Achatinella {Achatinellastrum) multizonata Baldwin. 

Achatinella {Achatinellastrum) multizonata Baldwin, P. Ac. Philad. 1S95, p. 215, 
pi. X. figs. I, 2 [animal described]. 

The shells collected by Mr Perkins from ' round Nuuanu' are in no sense typical 
of this species, they appear to be forms shewing links between it and A. bellula Smith 
{ = ligata Smith) ; indeed the two may prove to be forms of one variable species. 

Hab. Oahu, Nuuanu Valley (Baldwin); ridges round Nuuanu and Waimea 
(Perkins). 



326 FAUNA HAWAIIENSIS 



(96) Achatinella {Ackatinellastrum) nattii Baldwin and Hartman. 

Achatinella nattii Baldwin and Hartman in Hartman, P. Ac. Philad. 1888, p. 34, 
pi. I. fig. 3 [as nealii in explanation of plate]; Gwatkin, 1. c. 1895, P- 238. 
[radula]. 

Hab. Maui, Makawao to Honomu (Baldwin). 



(97) Achatinella [Achatinellastruni) olivacea Reeve. 

Achatinella olivacea Reeve, Conch. Icon. Achatinella, sp. 20. 

Achatinella prasina Reeve, 1. c. sp. 27. 

Hab. Oahu, Manoa to Nuuanu (Baldwin); Nuuanu and Mt. Tantalus (Perkins). 



(98) Achatinella [Achatinellastriim) papyracea Gulick. 

Achatinella papyracea Gulick, Ann. Lye. New York, vi. p. 229, pi. vii. fig. 48. 
Hab. Oahu, Kalaikoa, Ahonui, Wahiawa (Gulick). 



(99) Achatinella [Achatinellasti-uin) polita Newcomb. 

Achatinella polita Newcomb, Ann. Lye. New York, vi. p. 24(1853, May); t. c. 

p. 328 [animal]. 
Pease considered this to be identical with A. bella. 
Hab. Molokai (Newcomb) ; Kaluaaha to Halawa (Baldwin). 

(100) Achatinella [Achatinellastrum) producta Reeve. 

Achatinella p7'odncta Reeve, Conch. Icon. Achatinella, sp. 13; Newcomb, Ann. 
Lye. New York, vi. p. 315 [animal]; Bland and Binney, Ann. Lye. New 
York, X. p. 336, pi. XV. figs. 2, 4 [radula and anatomy]. 

Achatinella venulata Newcomb, P. Zool. Soc. London, 1854, p. 146, pi. xxiii. 
fig. 48. 

Achatinella hybrida Newcomb, t. c. p. 147, pi. xxiii. fig. 52. 

Achatinella dunkeri (Cuming MS.) Pfeiff"er, op. cit. 1855, p. 208. 

Hab. Oahu, Koolau (Newcomb, Hartman); Manoa to Nuuanu (Baldwin). 



MOLLUSC A 327 



(loi) Ackatinella [Achatinellasirum) saccata Pfeiffer. 

Achatinella [Ackatinellastrum) saccata Pfeiffer, P. Zool. Soc. London, 1859, p. 30. 

Unknown to me. 

Hab. Hawaiian Isles (Pfeiffer); Oahu (?) (Baldwin). 



(102) Achatinella [Achatinellasirum) solitaria Newcomb. 

Achatinella solitaria Newcomb, P. Zool. Soc. London, 1853 [1854], p. 150, 
pi. XXIV. fig. 60. 

Hab. Oahu, Palolo (Newcomb). 



(103) Achatinella [Achatinellastrum) trilineata GuJick. 

Achatinella trilineata Gulick, Ann. Lye. New York, vi. p. 226, pi. vii. fig. 46. 
Hab. Oahu, Palolo, Waialae, Wailupe, and Niu (Gulick). 



(104) Achatinella {Achatinellastruni) versipellis Gulick. 

Achatinella versipellis Gulick, Ann. Lye. New York, vi. p. 224, pi. vii. fig. 44. 
Hab. Oahu, Kailua (Gulick). 



(105) Achatinella {Achatinellastrmn) vulpina Ferussac. 

Helix vtilpina Ferussac, Voy. Freycinet, Zool. p. 447, pi. lxviii. figs. 13, 14; 

Souleyet, Voy. Bonite, Zool. 11. p. 508, pi. xxix. figs. 3, 4 [animal]. 
Achatina vulpina Ferussac, Deshayes, Hist. Moll. 11. pt. 2, p. 193, pi. clv. fig. 1. 
Achatinella vulpina Reeve, Conch. Icon. Achatinella, sp. 29. 
Achatinella castanea Reeve, 1. c. sp. 24. 
Achatinella adusta Reeve, I. c. sp. 30. 

Achatinella virens Gulick, Ann. Lye. New York, vi. p. 254, pi. viii. fig. -] i (1858). 
Achatinella fuscolineata Smith, P. Zool. Soc. London, 1873, p. 75, pi. ix. fig. 2. 
Achatinella livida Pfeiffer, P. Zool. Soc. London, 1845, P- ^9 [n^c Swainson]. 
Hab. Oahu, Palolo (Baldwin); Kailua, Palolo, Halawa (Smith); Manoa to 
Nuuanu (Baldwin); Nuuanu Yalley and Mt. Tantalus (Perkins). 

F. H. II. 43 



328 FAUNA HAWAIIENSIS 

van stewarti Green. 

Achatina stewarti Ore&n, Contrib. Macl. Lye. Philad. i. (1827, July), p. 47, pi. iv. 

figs. 1—4. 
Achatinella stewarti Green, Reeve, Conch. Icon. Achatinella, sp. 26. 
Achathiella pulcherrima Reeve, 1. c. sp. 23, fig. a [nee Swainson]. 
Achatinella tricolor Smith, P. Zool. Soc. London, 1873, p. 70, pi. ix. fig. 6. 
Hab. Oahu (various authors); Heia (Smith); Nuuanu and Mt. Tantalus 
(Perkins). 

van crassidentata Pfeiffer. 

Achatinella {Achatinellastruni) crassidentata Pfeiffer, P. Zool. Soc. London, 1855, 

p. 6, pi. XXX. fig. 23. 
Achatinella diversa Gulick, Ann. Lye, New York, vi. p. 220, pi. vii. fig. 42 

(1856, Dec). 
Achatinella varia Gulick, t. c. p. 222, pi. vii. fig. 43. 
Achatinella arialoga Gulick, t. c. p. 227, pi. vii. fig. 47. 

Hab. Oahu, Halawa (Baldwin) ; Halawa, Palolo, Waialae and Wailupe (Gulick) ; 
Waialae and Nuuanu (Perkins). 

van liliacea Pfeiffer. 

Achatinella [Achatinellastrum) liliacea Pfeiffer, P. Zool. Soc. London, 1859, 
p. 31. 

Hab. Oahu (Baldwin). 

The difficulty of arriving at a satisfactory dividing line between A. vulpina and 
A. producta is very great. As at present arranged, A. vulpina is the brown shell, 
van stewarti the greenish coloured form, van crassidentata the parti-coloured, and 
van liliacea the bandless variety ; all the above being sinistral. A. producta on the 
other hand is reserved for the larger and, usually, dextral form. 

(106) Achatinella [Achatinellastrum) ivailuaensis, sp. nov. 

Testa dextrorsa, subperforata, nitida, turrita, solidula, levissime striata, alba, lineis 
castaneis picta, apud peripheriam zona alba, in sutura linea nigro-castanea notata ; 
anfr. 5 — 5-^, regulariter crescentes, convexi ; apertura auriformis ; margine columellari 
plica fusca mediocri munita, margine dextro acuto, callo parietali tenuissimo. Long. 
15-5, alt. 8-4 mill. Plate XI. fig. 19. 

A pretty little shell of the group of A. Bella Reeve, of Molokai. A variety also 
occurred (Plate XI. fig. 20) in which the banding is almost obsolete, save in the suture 
of the earliest whorls and in one strong dark band below the periphery. 

Hab. Maui, Wailua (Perkins). 



MOLLUSC A 329 



(107) Achatinella {Achat ine/lasti'uin) zonata Gulick. 

Achatinella zonata Gulick, Ann. Lye. New York, vi. p. 237, pi. viii. fig. 58. 
Achatinella glauca Gulick, t. c. p. 232, pi. viii. fig. 51. 

United by Newcomb with A. trilincata Gulick ; it appears however to have 
much flatter whorls, and I follow Mr Baldwin, with some little doubt, in restoring it to 
specific rank. According to Newcomb, A. glaiica is a synonym of A. livida Swain., 
but specimens in the Brit. Mus. "named from Gulick's type" as a variety, lead me to 
place it here. 

Hab. Oahu, Waimea, Pupukea, Waialei, Kahuku, Hauula, and Kaawa (Gulick); 
above Ewa (Perkins). 

The following appears to be only a manuscript name : 

Achatinellastrum olesonii Baldwin, Cat. Shells Hawaiian Islands, 1893, p. 5. 

Hab. Oahu, Nuuanu. 



Perdicella Pease. 

Perdicella Pease, P. Zool. Soc. London, 1869, p. 649. 

Pease, unfortunately, having named no type, it becomes necessary to select one 
and I propose to take A. helena Newc. The species come from Maui and Molokai. 



( I ) Perdicella fnlgurans, sp. nov. 

Testa subperforata, dextrorsa, ovato-turrita, nitida, sub lente lineis spiralibus 
confertim sculpta, albida, strigis fulgurantibus castaneis elegantissime picta, sutura 
modice impressa, apice obtusulo ; anfr. 5^^, plano-convexi, ultimis | longitudinis testae 
aequans ; apertura ovato-pyriformis, intus lilacina ; peristoma margine dextro simplici, 
columellari subreflexo ; plica columellaris torta, subprominens, mediocris, rapide 
ascendens. Long. 16 ; lat. 8 ; long, apert. 8-i ; lat. apert. 4-9 mill. (Plate XL fig. 5.) 

This very pretty shell is akin to I^. zebrina Pfr., but may be readily separated 
from it by its greater size, by being much broader in proportion to the length, and by 
the colour-pattern being finer in design and more zigzag. The protoconch is brown, 
then becoming paler with a dark shade near the sutural line. It is the Partiilina 
zebrina Pfr. of Mr Baldwin's valuable catalogue. 

Hab. E. Maui, Makawao to Huelo (Baldwin); Maui (Hutchison). 

43—2 



330 FAUNA HAWAIIENSIS 

(2) Perdicella helena Newcomb. 

Achatinella helena Newcomb, Ann. Lye. New York, vi. (May, 1853), p. 27 ; P. Zool. 

Soc. London, 1853 [1854], p. 151, pi. xxiv. fig. 63. 
Hab. Molokai, on Ti-tree (Newcomb) ; Kamalo to Kalae (Baldwin) ; Kalae and 
Makakupaia (Perkins). 

(3) Perdicella mauiensis (Newcomb) Pfeiffer. 

Achatinella maniensis (sic, err. typ.) (Newcomb) Pfeiffer, P. Zool. Soc. London, 
1855 [1856], p. 207 ; Newcomb, Amer. J. Conch. 11. p. 217, pi. xiii. fig. 16. 

Hab. Maui, Makawao to Huelo (Baldwin). 

(4) Perdicella juinuscula Pfeiffer. 

Achatinella {Newcombia) minuscula Pfeiffer, P. Zool. Soc. London, 1858, p. 22. 
Hab. Maui, Lahaina (Baldwin). — Molokai Mts. at 4000 feet (Perkins). 
Both these habitats can hardly be correct ; I suspect the former may be an error 
of identification. 

(5) Perdicella ornata Newcomb. 

Achatinella ornata Newcomb, P. Zool. Soc. London, 1853 [1854], p. 149, pi. xxiv. 

fig- 55- 
Hab. Maui ; E. Maui (Newcomb); Lahaina (Baldwin). 

(6) Perdicella theodorei Baldwin. 

Achatinella {Partulina) theodorei Baldwin, P. Ac. Philad. 1895, p. 226, pi. x. 

fig. 27. 
Hab. Molokai, Kawela (Baldwin); Makakupaia and the mountains (Perkins). 

(7) Perdicella zebra Newcomb. 

Achatinella zebra Newcomb, Ann. Lye. New York, vi. p. 142 [Oct. 1855]. 
Placed by Dr Hartman, in his list, both in Achatinellastrttm and LaminellaX 
Hab. E. Maui (Newcomb). 



MOLLUSC A 331 

(8) Perdicella zebrina Pfeiffer. 

Newconibia zebrma Pfeiffer, P. Zool. Soc. London, 1855 [1856], p. 202. 
Hab. E. Maui (Baldwin as P. zebra Newc). 

Newcombia Pfeiffer. 

Newcombia Pfeiffer, Malak. Blatt. i. 1854, p. 117; Pease, P. Zool. Soc. London, 
1869, p. 649. 

Pfeiffer's list of species was very heterogeneous and included shells of diverse 
groups, his first species being A. hclena Newcomb ; fortunately Pease in 1869 properly 
confined the group to the shells it is now used for. 

Two sections may be formed: L Spirally lirate ; N. lirata, etc. IL Nearly 
smooth, usually more elongate ; N. cumingi, etc. 

(i) Newcombia canaliculata Baldwin. 

Achatinella [Newcombia) canaliculata Baldwin, P. Ac. Philad. 1895, p. 226, pi. x. 

figs. 28, 29 ; Gwatkin, t. c. p. 238 [radula]. 
Hab. Molokai, Halawa (Baldwin). 

(2) Neivcombia ciniiamoniea Pfeiffer. 

Achatinella cinnavwmea Pfeiffer, Malak. Blatt. iv. 1857, p. 230. 

Achatinella {^Newcombia) cinnamomea Pfeiffer, P. Zool. Soc. London, 1858, p. 22. 

Newcombia cinnamomea Pfeiffer, Gwatkin, P. Ac. Philad. 1895, p. 238 [radula]. 

Conchologically this is very close to N . cumingi Newc. 

Hab. Molokai, Mapulehu (Baldwin) ; Makakupaia and the mountains (Perkins). 

(3) Neivcombia cumingi Newcomb. 

Achatinella cumingi Newcomb, Ann. Lye. New York, vi. 1853, p. 25; P. Zool. 

Soc. London, 1853 [1854], pi. xxiv. fig. 59. 
Hab. Maui, Haleakala (Newcomb) ; Lahaina and Makawao (Baldwin). 

(4) Newcombia ge7nma Pfeiffer. 

Achatinella gemma Pfeiffer, Malak. Blatt. iv. 1857, p. 323. 

Achatinella {Newcombia) gemtna Pfeiffer, P. Zool. Soc. London, 1858, p. 22. 

Akin to N. lirata Pfr., but the sculpture is almost obsolete. 

Hab. Molokai Mts. (Perkins). 



332 FAUNA HAWAIIENSIS 



(5) Newcombia plicata (Mighels MS.) Pfeiffer. 

Achatinella plicata Pfeiffer, Mon. Helic. Viv. 11. 1848, p. 235 ; Newcomb, Ann. 

Lye. New York, vi. 1858, p. 312 [animal]; Reeve, Conch. Icon. Achatinella, 

sp. 44. 
Bulimus liratus Pfeiffer, P. Zool. Soc. London, 1851 [1853], p. 261. 

I cannot trace the supposed description by Mighels in P. Boston Soc. as 
Bulhmts plicatus. 

Hab. Molokai, Kalae (Baldwin) ; Mountains (Perkins). 



(6) Newcombia perkinsi Sykes. 

Newcombia perkinsi Sykes, P. Malac. Soc, London, 11. 1896, p. 130. 
(Plate XL fig. 36.) 

Hab. Molokai Mts. (Perkins). A fine series of this handsome shell. 

(7) Newcombia pfeifferi Newcomb. 

Achatinella pfeifferi Newcomb, Ann. Lye. New York, vi. p. 25 (May, 1853) ; 

P. Zool. Soc. London, [853 [1854], p. 150, pi. xxiv. fig. 58. 
Bulimus newcombiamis Pfeiffer, P. Zool. Soc. London, 1851, p. 261 [Dec. 1853]. 

Hab. Molokai, Kaluaaha (Baldwin). 

(8) Newcombia philippiana Pfeiffer. 

Achatinella philippiana Pfeiffer, Malak. Blatt. iv. 1857, p. 89. 
Hab. Molokai, Makakupaia (Baldwin). 

(9) Neivcombia sulcata Pfeiffer. 

Achatinella sulcata Pfeiffer, Malak. Blatt. iv. 1857, p. 231. 

Achatinella {^Newcombia) sulcata Pfeiffer, P. Zool. Soc. London, 1858, p. 22. 

Newcombia sulcata Pfeiffer, Gwatkin, P. Ac. Philad. 1895, p. 238 [radula]. 

Hab. Molokai, Pohakupili (Baldwin). 



MOLLUSC A 333 



Amastra H. and A. Adams. 

Amastra H. and A. Adams, Genera of Recent Mollusca, ii. p. 137. 

Type : the group of A. magna Ad. 

This large genus may for convenience be subdivided into groups somewhat in the 
following manner ; perhaps the large first section might be more broken up, though I 
think no sectional name will prove necessary. 



subgen. Amastra (s. str.). 

(i) Amastra affinis Newcomb. 

Achatinella affinis Newcomb, P. Zool. Soc. London, 1853 [1854], p. 142, pi. xxiii. 

fig- 35- 
Achatinella [Laminella) goniostoma Pfeiffer, P. Zool. Soc. London, 1855, p. 203. 

Amastra rtistica Gulick, P. Zool. Soc. London, 1873, p. 84, pi. x. fig. 17. 

Dr Hartman has suggested that A. riistica may equal A. variegata Pfr., an Oahu 

species. 

Hab. E. Maui, Kula (Newcomb, Gulick). 



(2) Amastra albolabris Newcomb. 

Achatinella albolabris Newcomb, P. Zool. Soc. London, 1853 [1854], p. 149, 

pi. XXIV. fig. 56. 
Achatinella nucleola Reeve, Conch. Icon. Achatinella, sp. 39 \_non Gould]. 

One young specimen I refer to this species. See a note under A. subrostrata 
Pfr. 

Hab. Oahu, Waianae Mts. (Newcomb, Perkins) ; Kapalama and Kalihi (Baldwin). 



(3) Amastra amicta Smith. 

Amasti-a amicta Smith, P. Zool. Soc. London, 1873, p. 86, pi. x. fig. 20. 
Dr Hartman notes that this species "may (t(\\x-e\ petricola" ; it appears to me quite 
distinct. 

Hab. Hawaiian Islands (Smith). 



334 FAUNA HAWAIIENSIS 

(4) Aniastra assiniilis Newcomb. 

Achatinella assimilis Newcomb, P. Zool. Soc. London, 1853 [1854], p. 148, 

pi. XXIII. fig. 53. 
Amastra assimilis Newcomb, Gwatkin, P. Ac. Philad. 1895, p. 238 [radula]. 
Achatinella dcshayesii Morelet, Bull. Soc. Hist. Nat. Moselle, 1857, p. 27 

[pars]. 
It has been suggested that this is a variety oi A. nubilosa Mighels, but the present 
species is a more slender shell with much flatter whorls. 

Hab. W. Maui (Newcomb). 

(5) Amastra azirostoma Baldwin. 

Amastra atcrostoma Baldwin, Nautilus, x. (July, 1896), p. 31. 
Hab. Lanai (Baldwin). 

(6) Amastra badia Baldwin. 

Amastra badia Baldwin, P. Ac. Philad. 1895, p. 230, pi. xi. fig. 40. 
Hab. Oahu, Ewa (Baldwin). 

(7) Amastra biplicata Newcomb. 

Achatinella biplicata Newcomb, P. Zool. Soc. London, 1853 [1854], p. 156, 
pi. XXIV. fig. 75. 

Achatinella deshayesii Morelet, Bull. Soc. Hist. Nat. Moselle, 1857, p. 27 [pars]; 
Pease, P. Zool. Soc. London, 1869, p. 652. 

Morelet's original series, now in the British Museum, consists of three specimens, 
one belonging to this species, and two to A. assimilis Newc. ; his diagnosis however 
refers to a form with only one columellar plait. From Mr Perkins' long series, it 
appears that the upper plait is variable and sometimes becomes obsolete ; in one 
specimen, which has received an injury, both plaits are dwarfed so as to show only as a 
slight thickening of the columella. 

Hab. Lanai (Newcomb); Waiapaa and Koele (Perkins). 

(8) Amastra breviata Baldwin. 

Amastra breviata Baldwin, P. Ac. Philad. 1895, p. 231, pi. xi. figs. 45, 46. 
Hab. Oahu, Palolo and Halawa (Baldwin). 



MOLLUSCA 335 

(9) Amasira citrca Sykes. 

Amastra citrea Sykes, P. Malac. Soc. London, 11. (1896), p. 129. 

Plate XI. fig. 4. 

Hab. Molokai (Hutchison). 

(10) Amastra conicospira Smith. 

Amastra conicospira Smith, P. ZooL Soc. London, 1873, p. 86, pi. x. fig. 10. 

Dr Hartman places this in the synonymy of A. assimilis Newc. ; I have never 
seen the present species, but from the figure it appears distinct. 
Hab. Hawaiian Islands (Smith). 

(11) Amastra conifera Smith. 

Amastra conifera Smith, P. Zool. Soc. London, 1873, P- SS- P^- ^- fig- ii- 
Hab. E. Maui, Kula (Smith). 

(12) Amastra cornea Newcomb. 

Achatinella cornea Newcomb, P. Zool. Soc. London, 1853 [1854], p. 141, pi. xxiii. 

fig- 32. 
Hab. Oahu, below Kaala (Perkins). 

Newcomb appears not to have noted the exact habitat ; the type-tablet in the 
British Museum is, however, labelled ' Oahu.' 

(13) Amastra crassilabrum Newcomb, 

Achatinella crassilabrnm Newcomb, P. Zool. Soc. London, 1853 [1854], p. 141, 
pi. XXIII. fig. 31. 

Hab. Oahu, Waianae (Newcomb, &c.). 

(14) Amastra cylindrica Newcomb. 

Achatinella cylindrica Newcomb, P. Zool. Soc. London, 1853 [1854], jj. 134, 
pi. XXII. fig. 1 1 ; Ann. Lye. New York, vi. p. 325 [animal]. 

Hab. Oahu, Waianae (Newcomb). 

F. H. II. 44 



336 FAUNA HAWAIIENSIS 

(15) Amastra decorticata Gulick. 

Amastra decorticata Gulick, P. Zool. Soc. London, 1873, P- ^4' pl- ^- fig- '^^■ 
Dr Hartman has united this with A. ellipsoidea of Gould, from Maui, but a glance 
at Gould's figures would have shewn him their distinctness. 

Hab. Oahu, Kawailoa, Halemano, and various localities (Gulick) ; ridges of 
Nuuanu (Perkins). 

(16) Amastra dia'andi Ancey. 

Amastra durandi Ancey, Naturaliste, 1897, p. 178. 
Hab. Oahu (Ancey). 

(17) Afnastra ellipsoidea Gould. 

Achatinella ellipsoidea Gould, P. Boston Soc. 11. (1847), p. 200; U. S. Explor. 
Exped. Moll. pi. VII. fig. 96. 

A species unknown to me : Newcomb united it with h\s A . ptipoidea, but it appears 
not to be so produced in form. 
Hab. Maui (Gould). 

(18) Amastra elliptica Gulick. 

Amastra elliptica Gulick, P. Zool. Soc. London, 1873, p. 83, pi. x. fig. 15. 
Hab. Oahu, Waialei, Kahuku, Hauula, Kawailoa (Gulick); Waianae (Hartman). 
Two specimens, collected by Mr Perkins on " Waianae Mts. Oahu," may belong to 
a large, incrassate variety. 

(19) Amastra extincta Pfeiffer. 

Achatinella [Laminelld] extincta Pfeiffer, P. Zool. Soc. London, 1855 [1856], 

p. 204. 
? ? Leptachatina hart )iia7ii {'^ ewe. MS.) Hartman, P. Ac. Philad. 188S, p. 54. 

I fancy that the new name given by Dr Hartman, on the ground that recent 
examples had been found, was due to an error of identification. Specimens submitted 
to me under the name of A. extincta by Mr Baldwin appear to be only a form of 
A. similaris Pease. 

Hab. Oahu, subfossil (Pfeiffer). 



MOLLUSC A 237 

(20) Amas(ra flavescens Newc. 

Achatineila flavescens '\>lQ:\\comh, P. Zool. Soc. London, 1853 [1854], p. 151, pi. 
XXIV. fig. 62. 

Hab. Hawaii (Newcomb). 

Dr Hartman gives also " Wanoa, Oahu (Newcomb)," but this must be an error. 

(21) Amastra frosti Ancey. 

Amastra frosti Ancey, Mem. Soc. Zool. France, v. (1892), p. 719; Sykes, P. 
Malac. Soc. London, iii. pi. xiii. fig. 12. 

Mons. Ancey has also described (P. Malac. Soc. London, in. p. 269, pi. xii. 
fig. 1 1 ) a variety unicolor. 

Hab. Oahu, Waianae (Ancey). 

(22) Amastra gray ana Pfeiffer. 

Achatinella [Laminella) grayana Pfeiffer, P. Zool. Soc. London, 1855 [1856, Feb.], 
p. 204. 

A single specimen. Dr Hartman has suggested that this is a form of A. magna, 
but I have not seen linking specimens. It is marked with spiral bands, due probably 
to the periostracum. 

Hab. Oahu (Clessin, Nomencl. Helic. Viv. 1881); ? Oahu (Baldwin). — Lanai, 
Lanaihale, a ground shell (Perkins). 

(23) Amastra humilis Newcomb. 

Achatinella humilis Newcomb, Ann. Lye. New York, vi. (Oct. 1855), p. 143; 

Amer. J. Conch. 11. (1866), p. 211, pi. xiii. fig. 4. 
Hab. Molokai, Kalae (Newcomb); Makakupaia and the mountains (Perkins). 

(24) Amastra inflata Pfeiffer. 

Achatinella {Laminella) inflata Pfeiffer, P. Zool. Soc. London, 1855 [Feb. 1856] 

p. 203. 
Hab. Oahu (Clessin, Nomencl. Helic. Viv.) ; Koolauloa (Baldwin). 

44—2 



338 FAUNA HAWAIIENSIS 

(25) Aniastra intermedia Newcomb. 

Achatinella intermedia Newcomb, P. Zool. Soc. London, 1853 [1854], p. 135, 
pi. XXII. fig. 13. 

Hab. Oahu, Waianae (Newcomb) ; ridges of Nuuanu, and Waianae Mts. below 
Kaala (Perkins). 

A long series. 

(26) Amastra irregularis Pfeiffer. 

Achatinella irregularis Pfeiffer, P. Zool. Soc. London, 1855 [Feb. 1856], p. 205. 
Hab. Hawaiian Islands (Pfeiffer). 

(27) Amastra lineolata Newcomb. 

Achatinella lineolata Newcomb, Ann. Lye. New York, vi. (1853), p. 29; P. Zool. 
Soc. London, 1853 [1854], p. 140, pi. xxiii. fig. 29. 

Hab. Hawaii (Newcomb). 

The habitat ' Maui,' originally given, seems to have been a slip. 

(28) A^nastra longa Sykes. 

Amastra longa Sykes, P. Malac. Soc. London, 11. (Oct. 1896), p. 129. 

Plate XL fig. 35. 

Hab. Lanai (Newcomb) ; windward side, apparently extinct (Perkins). 

(29) Amastra luctuosa Pfeiffer. 

Achatinella [Laminella) luctttosa Pfeiffer, P. Zool. Soc. London, 1855 [Feb. 1856], 

p. 204. 
Laminella luctuosa Pfeiffer, Bland and Binney, Ann. Lye. New York, x. pp. 

335 — 6 [jaw and radula]. 

Hab. Oahu, Waialee (Baldwin). 



MOLLUSC A 339 

{2>o) Ainastra magna C. B. Adams. 

Achatinella magna C. B. Adams, Contrib. to Conch, p. 125, 1850; Newcomb, 

Ann. Lye. New York, vi. (1858), p. 319 [animal]. 
Achatinella baldwinii Newcomb, P. Zool. Soc. London, 1853 [1854], p. 155, 

pi. XXIV. fig. 72. 

I have seen a good series of this handsome shell. 
Hab. Lanai (Newcomb) ; behind Koele (Perkins). 

(31) Aniastra malleata Smith. 

Amastra malleata Smith, P. Zool. Soc. London, 1873, P- ^5, pi. x. fio-. 18. 

Mr Baldwin has united this with A. affinis Newcomb ; I have not seen a specimen, 
but, from the figure and description, they appear distinct. 

Hab. E. Maui, Kula (Smith). 

(32) Amasti'a mastersi Newcomb. 

Achatinella mastersi Newcomb, P. Zool. Soc. London, 1853 [1854], p. 153, pi. 

XXIV. fig. 67 ; Ann. Lye. New York, vi. p. 332 [animal]. 
Laminella mastersi Newcomb, Bland and Binney, Ann. Lye. New York, x. p. 335, 

pi. XV. figs. 7, 9 — 1 1 [jaw and radula]. 
Amastra mastersi Newcomb, Gwatkin, P. Ac. Philad. 1895, p. 239 [radula]. 
Achatinella rubens Pfeiffer, Malak. Blatt. 1854, p. 129 [nee Gould, fide Newcomb]. 

Hab. Maui (Newcomb) ; ? Haleakala at 5000 feet (Perkins). 

Two, apparently immature, specimens, which I refer here with some hesitation. 
Four specimens from " Molokai Mts." appear to me to be very close to this species. 

{-^■^ Amastra melanosis Newcomb. 

Achatinella melanosis Newcomb, P. Zool. Soc. London, 1853 [1854], p. 144, 
pi. xxiii. fig. 41. 

Hab. Hawaii (Newcomb). 

(34) Amastra modest a C. B. Adams. 

Achatinella modesta C. B. Adams, Contrib. to Conch. 1850, p. 128. 
Achatinella pumila Guliek, Clessin, Nom. Helic. Viv. 1881, p. 313. 

Hab. Hawaiian Islands (Adams). — Molokai (Hartman, Baldwin). 



340 FAUNA HAVVAIIENSIS 



(35) Amastra moesta Newcomb. 

Achatinella moesta Newcomb, P. Zool. Soc. London, 1853 [1854], p. 157, pi. xxiv. 

fig- 77. 
Achatinella obscura Newcomb, t. c. p. 157, pi. xxiv. fig. 78. 

According to Pease (P. Zool. Soc. London, 1869, p. 651), and he is followed by 
Dr Hartman (P. Ac. Philad. 1888, p. 47), these two forms are one species. Probably 
this is correct. 

Hab. Lanai (Newcomb). 



(36) Amastra mucronata Newcomb. 

Achatinella mucronata Newcomb, Ann. Lye. New York, vi. (May 1853), p. 28; 

P. Zool. Soc. London, 1853 [1854], p. 146, pi. xxiii. fig. 49. 
Achatinella {Laminella) fusiformis Pfeiffer, P. Zool. Soc. London, 1855, p. 5, 

pi. XXX. fig. 18. 

Hab. Molokai (Newcomb, Baldwin). 

Newcomb, in his later paper, gave the locality of ' Maui,' but probably this is a 
slip due to the fact that he was, as he subsequently stated, unable to see the proofs. 
Two specimens, collected on Molokai by Mr Perkins, appear to belong to a dwarf race. 



[t,j) Amast7'a nana Baldwin. 

Aitiastra nana Baldwin, P. Ac. Philad. 1895, ?• 232, pi. xi. figs. 48, 49 [with 
animal] ; Gwatkin, t. c. p. 239 [radula]. 

Hab. Maui, Makawao at 4000 feet (Baldwin). 



(38) Amastra nigra Newcomb. 

Achatinella nigra Newcomb, P. Boston Soc. v. (Sept. 1855), p. 219; Amen J. 
Conch. II. (1866), p. 210, pi. XIII. fig. 3. 

According to Clessin (Nomencl. Helic. Viv. 1881, p. 311) A. globosa, Gulick nee 
Pfeiffer, is a synonym. 

Hab. E. Maui (Newcomb). 



MOLLUSC A 341 

(39) Amastra mibilosa Mighels. 

Achatinella nubilosa Mighels, P. Boston Soc. ir. (1845), P- 20; Reeve, Conch. 

Icon. Achatinella, sp. i ; Newcomb, Ann. Lye. New York, vr. p. 312 

[animal]. 
Achatinella nubilosa Gould, U.S. Explor. Exped. Moll. pi. vii. fio-. 95. 
Hab. Molokai (Newcomb) ; Kalae (Baldwin). 

It has been suggested, but I think erroneously, that this species comes, in reality, 
from Oahu. 

(40) Amastra micula Smith. 

Amastra nucula Smith, P. Zool. Soc. London, 1873, p. 85, pi. x. fig. 19. 
Hab. Lanai ? (Smith). 

(41) Amastra peasei Smith. 

Amastra peasei Smith, P. Zool. Soc. London, 1873, p. 86, pi. x. fig. 13. 
Hab. Hawaiian Islands (Smith). — Oahu ? (Baldwin). 

(42) Amastra pellucida Baldwin. 

Amastra pellucida Baldwin, P. Ac. Philad. 1895, p. 231, pi. xi. figs. 41, 42 [with 

animal]. 
Hab. Oahu, Waianae Valley (Baldwin). 

(43) Amastra petricola Newcomb. 

Achatinella petricola Newcomb, Ann. Lye. New York, vi. (Oct. 1855), p. 143; 

Amer. J. Conch. 11. (1866), p. 211, pi. xiii. fig. 6. 
Achatinella {Laminella) umbilicata Pfeifier, P. Zool. Soc. London, 1855 [Feb. 

1856], p. 205. 
Amastra umbilicata Pfr., Hartman, P. Ac. Philad. 1888, p. 50, pi. i. fig. 11. 

Hab. Molokai (Newcomb) ; Mapulehu (Baldwin). 

(44) Amastra porphyria Newcomb. 

Achatinella porphyria Newcomb, P. Zool. Soc. London, 1853 [1854], p. 136, 

pi. XXII. fig. 16. 
Achatinella {Laminella) grossa Pfeiffer, P. Zool. Soc. London, 1855 [Feb. 1856], 

p. 204. 
Hab. Oahu, Waianae (Newcomb). 



542 FAUNA HAWAIIENSIS 



(45) Ajnastra porphyrostonia Pease. 



Amastra porphyrostoma Pease, J. Conchyl. xvir. (1869), p. 172 ; Hartman, P. Ac. 
Philad. 1888, p. 48, pi. I. fig. 6. 

Hab. Oahu (Pease). 



(46) Amastra piillata Baldwin. 

Amastra pullata Baldwin, P. Ac. Philad. 1895, p. 228, pi. xi. figs. 31, 32 ; 

Gwatkin, t. c. p. 239 [radula]. 
Amastra twibrosa Baldwin, t. c. p. 229, pi. xi. figs. 36, 37 ; Gwatkin, t. c. p. 239 

[radula]. 

After an examination of the long series collected by Mr Perkins I have been 
unable to sever these two species. The animals are said to differ in colour, and they 
are said to inhabit different districts ; conchologically they seem to shade into one 
another, and the radula appears to be identical. Probably they will prove to be local 
races. 

Hab. Molokai, Kamalo (Baldwin, as A. rnnbrosa) ; Waikolu (Baldwin, as A. 
pjtUata) ; Kamalo and Makakupaia Mts. (Perkins). 



(47) Amastra pupoidea Newcomb. 

Achatinella pupoidea Newcomb, P. Zool. Soc. London, 1853 [1854], p. 144, 
pi. xxiii. fig. 42. 

The colouring is somewhat exaggerated in the figure. See also a note under A. 
ellipsoidea Gould. 

Hab. E. Maui (Newcomb). 



(48) Amastra pitsilla Newcomb. 

Achatitiella pusilla Newcomb, Ann. Lye. New York, vi. (Oct. 1855), p. 144; 

Amer. J. Conch. 11. (1866), p. 211, pi. xiii. fig. 5. 
Achatinella pidla (Newcomb) Pfeiffer, P. Zool. Soc. London, 1855 [Feb. 1856], 

p. 209. 

Hab. Lanai (Newcomb). 



MOLL use A 343 



(49) Amasti-a reticulata Newcomb. 

Achatinella retiatlata Newcomb, P. Zool. Soc. London, 1853 [1S54], p. 148, 

pi. XXIV. fig. 54. 
Achatinellastruyn conspersa Pfeiffer, P. Zool. Soc. London, 1855 [March], p. 7, 

pi. XXX. fig. 26. 

Hab. Oahu, Waianae (Newcomb). 



(50) Aviastra nibcns Gould. 

Achatinella ritbens Gould, P. Boston Soc. 11. (1845), P- -7 ! Newcomb, Ann. Lye. 
New York, vi. p. 314 [animal]; Reeve, Conch. Icon. Achatinella, sp. 42, 
pars b [fide Newcomb]. 

Hab. Oahu, W. Mts. (Newcomb) ; Kaala (Baldwin). 



(51) Aj7tastra rubicitnda Baldwin. 

Amastra rubicimda Baldwin, P. Ac. Philad. 1895, p. 229, pi. xi. fig. 38 [with 
animal] ; Suter, t. c. p. 240, pi. xi. figs. 54 [jaw], 56 [radula]. 

Hab. Oahu, Konahuanui Mt. (Baldwin). 



(52) Aviastra rtibida Gulick. 

Amastra rubida Gulick, P. Zool. Soc. London, 1873, p. 84, pi. x. fig. 12. 
Hab. Oahu, Kahuku (Gulick). 

(53) Amastra sericea Pfeiffer. 

Achatinella {Lamhiella) sericea Pfeiffer, P. Zool. Soc. London, 1859, p. 31. 
Hab. Hawaiian Isles (Pfeiffer). — ? Oahu, Waialua (Baldwin). Unknown to me. 

(54) Amastra simularis Hartman. 

Amastra simularis Hartman, P. Ac. Philad. 1888, p. 252, pi. xiii. fig. 7. 
Amastra similaris Hartman, Gwatkin, op. cit. 1895, p. 239 [radula]. 
Hab. Molokai (Hartman, Perkins); Mapulehu (Baldwin). 

F. H. II. 45 



344 FAUNA HAWAIIENSIS 

var. roseotincta Sykes. 

A, sinmlaris Hartman, var. roseotincta Sykes, P. Malac. Soc. London, ii. p. 130. 
Plate XI. fig. 3. 

Hab. Molokai mountains (Perkins). 

Varieties under the names of maura and semicarnea have recently been described 
by Mons. Ancey (P. Malac. Soc. London, in. p. 270, pi. xiii. figs, 8, 16). 

• (55) Amastra solida Pease. 

Amasti-a solida Pease, J. Conchyl. xvii. (1869), p. 173. 
Hab. Oahu (Pease). 

(56) Amastra spirizona Ferussac. 

Helix [Cochlogena) spirizona Ferussac, Prodrome, no. 433. 
Achatina spirizona Yh.x., Hist. Moll. 11. pt. 2, p. 196, pi. clv. figs. 14, 15. 
Achatinella spirizona Fen, Newcomb, Ann. Lye. New York, vi. p. 307 [animal]. 
Achatinella actita Swainson, Quart. J. Sci. Lit. Arts, i. (1828), p. 84; Zool. 

Illustr., ser. 2, pi. xcix. fig. 3. 
Achatinella bcstica Mighels, MS. 

Hab. Oahu, Waianae Mts. (Baldwin); Waianae Mts. below Kaala on lee side, 
and Halemano (Perkins). 

var. nigrolabris Smith. 

Amastra nigrolabris Smith, P. Zool. Soc. London, 1873, p. 85, pi. x. fig. 9. 
Hab. Oahu, Wahiawa, Kalaikoa, Waimea (Smith) ; Halemano (Perkins). 

var. rttdis Pfeiffer. 

Achatinella rudis Pfeiffer, P. Zool. Soc. London, 1855, p. 5 (pars). 

Laminella albida Pfeiffer, t. c. p. 203. 

Newcombia chloi'otica Pfeiffer, t. c. p. 203. 

Hab. Oahu. 

A. nigrolabris Smith, is, in my opinion, only a broader variety, in which the light 
band below the suture is wider : in a box of specimens from Halemano forms are found 
showing a graduation from it to the typical form. It is possible that A. rudis may be 
a distinct species, but I fancy it is only a paler colour variety. Mr Perkins notes that 
he found the species "mostly on dead branches of trees, covering itself with mucus to 
which debris of bark and wood stick, and therefore very well concealed." 



MOLL use A 345 

(57) Ajuastra stibrostrata Pfeiffer. 

Achatinella [Laminella) siibrostrata Pfeiffer, P. Zool. Soc. London, 1859, p. 31. 

Hab. Hawaiian Islands (Pfeiffer). — ? Oahu (Baldwin). 

Dr Hartman suggests that this is the same as A. albolabris Newc. ; it may possibly 
be an elongate variety, but I doubt it. 

(58) Amastra tenuilabris Gulick. 

Amastra tenuilabris Gulick, P. Zool. Soc. London, 1873, p. 83, pi. x. fig. 16. 

Hab. Oahu (Gulick, with some doubt). 

Dr Hartman remarks that from "a comparison of types" this is a synonym oi A. 
flavesccns Newc. I have, equally, examined the types, and this species differs in being 
stumpier, with a larger mouth, and more shouldered whorls. 

(59) Amastra temtispira Baldwin. 

Amastra tenuispira Baldwin, P. Ac. Philad. 1895, p. 232, pi. xi. fig. 51. 
Hab. Oahu, Kaala Mt. (Baldwin). 

(60) Amastra tcxtilis Ferussac. 

Helix [Helic teres) textilis Ferussac, Voy. Freycinet, Zool., p. 482. 

Achatinella textilis Ferussac, Binney, Ann. Lye. New York, xi. p. 190, pi. xiv. 

fig. G [radula]. 
Amastra textilis Ferussac, Hartman, P. Ac. Philad. 1888, p. 50, pi. i. fig. 8. 
Achatinella microstoma Gould, P. Boston Soc. 11. (1845), p. 28. 
Achatinella ventulus Ferussac, Reeve, Conch. Icon. Achatinella, sp. 31 ; Pfeiffer 

in Conch. -Cab. Achatinella, p. 287, pi. Lxvii. figs. 12, 13 [nee Ferussac]. 

Hab. Oahu (Baldwin, Hutchison, &c.). 

A single specimen from Nuuanu, Oahu, appears to be a varietal form. 

(61) Amastra transversalis Pfeiffer. 

Achatinella [Laminella) transversalis Pfeiffer, P. Zool. Soc. London, 1855 [Feb. 

1856], p. 204. 
United by Newcomb with A. reticulata, but appears distinct. 

Hab. Oahu, Keawaawa (Baldwin). 

45—2 



346 FAUNA HAWAIIENSIS 



(62) Amastra tristis Ferussac. 

Helix iyHelicteres) tristis Ferussac, Voy. Freycinet, Zool. p. 482, pi. lxviii. 

figs. 6, 7. 
Achatinella tristis Ferussac, Reeve, Conch. Icon. Ac/mtinella, sp. 37 ; Pfeiffer in 

Conch. -Cab. Achatinella, pi. lxvii. figs. 10 — 11 [not good]. 
Achatinella ftiliginosa Gould, P. Boston Soc. 11. (1845), p. 28. 

Hab. Oahu, Palolo (Hartman); Nuuanu to Palolo (Baldwin). 



(63) Amastra tiiry-itella Ferussac. 

Helix [Cochlogena) tttrritella Ferussac, Prodrome, No. 434 ; Voy. Freycinet, 

Zool. p. 481. 
Achatina t2irritella Fer., Hist. Moll. 11. pt. 2, p. 196, pi. CLV. fig. 13; Souleyet, 

Voy. Bonite, Zool. 11. p. 509, pi. xxix. figs. 7 — 8 [animal]. 
Achatinella tur'ritella Fer., Newcomb, Ann. Lye. New York, vi. p. 307 [animal]. 
? Helix {Cochlogena) hiteola Ferussac, Voy. Freycinet, Zool. p. 480. 
? Achatina luteola Fen, Hist. Moll. 11. pt. 2, p. 196, pi. clv. fig. 12. 
? Laniinella luteola Fer. (sic), Hartman, P. Ac. Philad. 1888, p. 42. 
Achatina oahuensis Green, Contrib. Macl. Lye. Phil. i. (1827), p. 49, pi. iv. fig. 5. 
Achatinella inornata Mighels, P. Boston Soc. 11. (1845), p. 21. 

Newcomb states that he was unable to trace the type oi A. luteola, and apparently 
it is lost : Pease (P. Zool. Soc. London, 1869, p. 652) united it with A. turritella, and 
probably this will prove to be correct. 

Hab. Oahu (authors) ; Kalihi to Palolo (Baldwin) ; ridges of Nuuanu Valley 
(Perkins). 



(64) Amastra iindata Baldwin. 

Amastra zmdata Baldwin, P. Ac. Philad. 1895, P- 23°' P^- ^i- ^g- 39- 
Hab. Oahu, Nuuanu (Baldwin). 

(65) Amastra uniplicata Hartman. 

Amastra zmiplicata Hartman, P. Ac. Philad. 1888, p. 50, pi. i. fig. 7. 
Hab. Molokai (Hartman). 



MOLLUSC A 347 

(66) Aniastra variegata Pfeiffer. 

Achatinclla variegata Pfeiffer, Zeitschr. flir Malak. 1849, p. 90; Conch. -Cab. 

Acliatinella, p. 282, pi. lxvii. figs. 14, 15. 
Amastra riibens, Reeve, pars, Conch. Icon. Acliatinella, fig. 42 a [fide Newcomb]. 
Acliatinella decepta C. B. Adams, Contrib. to Conch. 1850, p. 127. 
Hab. Oahu, head of Boothes Valley (Hartman) ; Waianae (Baldwin). 

(67) Aniastra ventulus Ferussac. 

Helix [Helicteres) ventulus Ferussac, Voy. Freycinet, Zool. p. 481. 
Achatinella ventulus Ferussac, Newcomb, Ann. Lye. New York, vi. p. 306 

[animal]. 
Achatinclla niclampoides Pfeiffer, P. Zool. Soc. London, 1851 [Dec. 1853], p. 262 ; 

Pfeiffer in Conch. -Cab. Achatinella, p. 288, pi. lxvii. figs. 8, 9. 
Acliatinella (^Amastra) manoaensis Newc, Clessin, Nom. Helic. Viv. 1881, p. 311. 
May prove to be a Leptachatina. 

Hab. Oahu, Nuuanu to Palolo (Baldwin) ; Panoa Valley and ridges of Nuuanu 
(Perkins). 

(68) Amastra violacea Newcomb. 

Acliatinella violacea Newcomb, Ann. Lye. New York, vi. (1853, May), p. 18; 

P. Zool. Soc. London, 1853 [1854], p. 135, pi. xxii. fig. 14. 
Achatinella gigantea Newcomb, P. Zool. Soc. London, 1853 [1854], p. 136, pi. 

XXII. fig. 17. 
These two species have been united by Pease and Dr Hartman, the latter 
remarking "The only example oi gigantea ever found is in the British Museum. It 
probably equals a large example of ^. violacea, Newc." Probably the locality of Maui, 
given by Newcomb, was an error, as his specimen appears to be only an elongate 
form of the Molokai shell. This varies very much in size and shape, as may be seen 
from the following : 

Alt. 34; diam. 16; alt. ap. 15 ; lat. ap. 9-5 mill. 
,, 31 ; ,, 12 ; ,, ,, I 1-5 ; ,, ,,7 
,, 3 ^ ' " 15'" " 13 ' " "9 " 
Mr Baldwin has left A. criaantea in his list as a Maui shell, but the fact that this 
diligent collector has marked it as a shell unknown to him, lends confirmation to the 
view that it does not really come from that island. 

Hab. Molokai, Haleakala (Newcomb) ; Mapulehu to Halawa (Baldwin) ; Halawa 
and Pelekunu (Perkins).—? Maui -Sl?, A. gigantea, Haleakala (Newcomb). 



348 FAUNA HAWAIIENSIS 

subgenus Laminella Pfeiffer. 

Laminella Pfr., Malak. Blatt. i. 1S54, p. 126. 

Pfeiffer's original group was very heterogeneous, as was that of Pease under 
this name (P. Zool. Soc. London, 1869, p. 648); the latter author also proposing 
Perdicella for a portion of the group. I would propose to select A. gravida Fer., the 
old and well-known species, as the type. 

(69) Amastra i^Larninella) alexandri Newcomb. 

Achatinella alexandid Newcomb, P. Calif. Ac. in. (1865), p. 182 ; Amer. J. Conch., 

II. (1866), p. 216, pi. XIII. fig. 14. 
Hab. West Maui, at 7500 feet (Newcomb) ; top of West Maui (Baldwin). 

(70) Amastra {^Laminella) citrma (Mighels MS.) Pfeiffer. 

Achatinella citrina Mighels, Pfeiffer, Mon. Helic. Viv. 11. (1848), p. 234; Reeve, 
Conch. Icon. Achatinella, sp. 33 ; Newcomb, Ann. Lye. New York, vi. p. 312 
[animal]. 
Pease united (P. Zool. Soc. London, 1869, p. 652) this species with A. vemista. 
Conchologically, they differ in the periostracum, shape of whorls, &c., while, from the 
descriptions given by Newcomb, the animals are distinct in colouration. 
Hab. Molokai, Kalae to Waikolu (Baldwin) ; Molokai (Perkins). 

(71) Amastra [Laminella) concinna Newcomb. 

Achatinella concinna Newcomb, P. Zool. Soc. London, 1853 [1854], p. 157, 
pi. XXIV. fig. 79. 

Newcomb's type was a bandless dextral shell. In the very fine series collected, 
both dextral and sinistral forms occur ; black bands are either absent or present, and, 
in the latter event, vary from one to even four in number. 

Hab. Lanai (Newcomb, &c.) ; Koele side of highest point, side of highest point 
furthest from Koele, near Koele at 3000 feet (Perkins). 

(72) Amastra {Laminella) depicta Baldwin. 

Laminella depicta Baldwin, P. Ac. Philad. 1895, p. 228, pi. xi. figs. 33 — 5 [animal 
described]. 

A very fine series, shewing a range of colour from pale yellow to rich orange, 
tinged with crimson ; it is sometimes dextral, but sinistral forms predominate. 

Hab. Molokai, Kamalo (Baldwin); mountains, and above Pelekunu (Perkins). 



MOLL use A 349 

{^2,) Aniastra {Laminella) elongata Newcomb. 

Achatinella elongata Newcomb, Ann. Lye. New York, vi. (May, 1853), p. 26. 
Achatmella acuta Newcomb, P. Zool. Soc. London, 1853 [1854], p. 142 [nee 
Swainson]. 

The figure given in P. Zool. Soc. London (1. c.) under this name does not (fide 
Newcomb) represent the present species, but A. soror. Dr Hartman gives ' Makawao, 
Maui ' as the habitat, but this must be an error ; further he unites the species, 
erroneously in my opinion, with A. hutchinsonii Pease. 

Hab. Oahu, Lehue (Newcomb) ; Waianae Mts (Baldwin). 

(74) Aniastra {Laiinnella) erecta Pease. 

Laminella erecta Pease, J. Conchyl., xvii. (1869), p. 174. 
Close to A. viicans Pfeiffer. 
Hab. Maui (Pease). 

(75) Aniastra {^Laminella) farcimen Pfeiffer. 

Achatinella {^Laminella) farcimen Pfeiffer, P. Zool. Soc. London, 1856, p. 334. 
nee Amastra farcimen, Pfeiffer, Hartman, P. Ac. Philad. 1888, p. 46, pi. i. fig. 5. 

Dr Hartman states that his figure is "typical"; this is obviously incorrect as the 
type is a sinistral specimen, of considerable size, while the figure represents a smaller, 
dextral, shell of another group. What species his shell may belong to, I am unable to 
determine, but it appears to possess no columellar plait. 

Hab. Maui (Newcomb, fide Pfeiffer). 

(76) Amastra {L^aininclUi) fratcrna Sykes. 

Amastra fratcrna, Sykes, P. Malac. Soc. London, 11. (Oct. 1896), p. 129. 

Plate XL fig. 23. 

Hab. Lanai, mountains behind Koele (Perkins). 

{^'Jl^ Amastra (^Laminella) gravida Ferussac. 

Helix gj^avida Ferussac, Voy. Freycinet, Zool. p. 478, pi. lxviii. figs. 4, 5. 
Achatina gravida Ferussac, Deshayes, Hist. Moll. 11. p. 192, pi. clv. figs. 3, 4. 
Achatinella gravida Fen, Newcomb, Ann. Lye. New York, vi. p. 307 [animal]. 



350 FAUNA HAVVAIIENSIS 

Achatinella sitffusa Reeve, Conch. Icon. Achatinella, sp. ii. 

Achatinella dimondi C. B. Adams, Contrib. to Conch. 1850, p. 126 (with van 

lata). 
The specimen described by Reeve does not now appear to exist in the Brit. Mus. 
collection. 

Hab. Oahu, Kalihi to Niu (Baldwin) ; Nuuanu (Perkins). 

(78) Avtastra [Laiiiiiic/ia) helvina Baldwin. 

Achatinella [Lajuiiiella) helvina Baldwin, P. Ac. Philad. 1895, P- 227, pi. xi. 

fig. 30 [shell, animal, and anatomy] ; Gwatkin, t. c. p. 239 [radula]. 
Some specimens, given to Mr Perkins by Mr O. Meyer, are broader and have 
more periostracum, forming a link towards A. picta. 

Hab. Molokai, Ohia Valley, near Kaluaaha (Baldwin); Molokai (Perkins). 

(79) Amastra [Latnitiella] hutchinsonii Pease. 

Helictcr htUchinsonii Pease, P. Zool. Soc. London, 1862, p. 7. 

Amastra hutchinsonii Pease, Hartman, P. Ac. Philad. 1888, p. 45, pi. i. fig. 9. 

Dr Hartman suggests, I think erroneously, that this is a synonym of A. elongata 
Newc. 

Hab. Maui (Pease). 

(80) Amastra [Laminclla) niicans Pfeiffer. 

Achatinella [Laniinella) niicans Pfeiffer, P. Zool. Soc. London, 1859, p. 31. 
Ainastra niicans Pfeiffer, Hartman, P. Ac. Philad. 1888, pi. i. fig, 10. 
Dr Hartman's figure is not very good. 
Hab. Oahu (Baldwin, Hutchison). 

(81) Aniasti-a {Laniinella) picta Mighels. 

Achatinella picta Mighels, P. Boston Soc. 11. (1845), p. 21 ; Newcomb, Ann. Lye. 

New York, vi. p. 311 [animal]; Reeve, Conch. Icon. Achatinella, sp. 36; 

Pfeiffer, Conch.-Cab. Achatinella, p. 284, pi. lxviii. figs. 28, 29 [not very 

good] ; Bland and Binney, Ann. Lye. New York, x. p. 335, pi. xv. fig. 6 

[jaw J. 
Achatinella picta Pfeiffer, P. Zool. Soc. London, 1845 [1846], p. 90. 

Hab. Maui, Lahaina and Makawao (Baldwin) ; Haleakala, at 4000 feet, and lao 
Valley (young shells) (Perkins). 



MOLLUSC A 35; 



var. bulbosa Gulick. 

Achatinella bitlbosa Gulick, Ann. Lye. New York, vi. (1858), p. 253, pi. viii. 
fig. 71. 

Newcomb placed A. bulbosa as a synonym ; it appears to me to be of varietal 
rank, and to differ in being larger and in the whorls being more flattened. I fancy 
the species will prove to be variable, as Mr Perkins' shells are more slender than the 
series in the Museum. Specimens sent by Mr Hutchison as from 'Maui' are still 
more slender, and may possibly prove to be distinct. Mighels gave, by error probably, 
' Oahu.' 

Hab. E. Maui, Honuaula and Kula (Gulick). 



(82) Ainastra [Laininel/a] remyi Newcomb. 

Achatinella remyi Newcomb, Ann. Lye. New York, vi. (Oct. 1855), P- H^ ^ 
Amer. J. Conch. 11. (1866), p. 215, pi. xiii. fig. 13. 

Hab. Lanai (Newcomb). 

Only known to me from the original series in the Brit. Mas. ; Pfeiffer 
(P. Zool. Soc. London, 1855, p. 207) gave Hawaii as the habitat, but probably this 
was an error. 

(83) Amastra {Laniinclla) sangitinea Newcomb. 

Achatinella sangttinca Newcomb, P. Zool. Soc. London, 1853 [1854], p. 135, 

pi. XXII. fig. 15 ; Ann. Lye. New York, vi. p. 326 [animal]. 
Laminella fernssaci Pfeiffer, P. Zool. Soc. London, 1855 [1856], p. 203. 
Hab. Oahu, Lehui (Newcomb); Waianae and Halemano (Baldwin) ; Halemano, 
Kawailoa, and Makaha Valley (dead) (Perkins). 

(84) Amastra {Laminella) soror Newcomb. 

Achatinella soror Newcomb, P. Zool. Soe. London, 1853 [1854], p. 143, pi. xxiii. 

fig. 38 [also fig. 36, sub nom. A. acuta']. 
Hab. Maui (Newcomb). 

The additional locality of Lanai given, subsequently, by Newcomb, really, I think, 
refers to my A. fratcrna. 

F. H. II. 46 



352 FAUNA HAWAIIENSIS 

(85) Amastra {Laminella) straminea Reeve. 

Achatinella straminea Reeve, Conch. Icon. Achatinelia, sp. 38 ; Newcomb, Ann. 

Lye. New York, vi. p. 318 [animal]. 
Hab. Oahu, Panoa to Palolo (Baldwin) ; Nuuanu (Perkins). 

(86) Amastra [Laminetla] tetrao Newcomb. 

Achatinella tetj'ao Newcomb, P. Boston Soc. v. (1855), p. 219; Ann. Lye. New 
York, VI. p. 334 [animal]; Amer. J. Conch. 11. (1866), p. 214, pi. xiii. 
figs. II, 12. 
From the fine series collected it appears that the ground-colouring, under 
the zigzag periostracum, varies considerably. Shades of crimson or rich orange pre- 
dominate, but occasionally the colour is confined to a band below the suture, the 
rest of the shell being whitish. 

Hab. Lanai (Newcomb); mountains and behind Koele (Perkins). 

(87) Amastra {^Laminella) veimsta Mighels. 

Achatinella vetmsta Mighels, P. Boston Soc. 11. (1845), p. 21 ; Newcomb, Ann. Lye. 
New York, vi. p. 311 [animal]; Reeve, Conch. Icon. Achatinella, sp. 32; 
Binney, Ann. Lye. New York, xi. p. 191, pi. xiv. fig. D. 

Hab. Molokai, Mapulehu (Baldwin); mountains (Perkins). 

Mighels gave, but erroneously, ' Oahu ' as the locality. 

(88) Amastra [Laminella) villosa Sykes. 

Amastra villosa Sykes, P. Malac. Soc. London, 11. (1896), p. 129. 
Plate XI. fig. 24. 

The specimen here figured is not the one whose measurements were given in the 
original diagnosis, but a slightly smaller shell whose periostracum is better preserved. 

Hab. Molokai (Perkins). 

subgenus Amastkella, n. subgen. 

This name is proposed for a group of rotund, generally incrassated, small forms, 
which have been usually placed in Amastra. I take as the type A. rugnlosa Pease. 
They are nearly all natives of Kauai, but a few species are found on other islands. 



MOLLUSC A 353 



(89) Aniastra iyAmast^'ella) antkojiii Newcomb. 
Achatinella anthonii Newcomb, P. Calif. Ac. 11. (1861), p. 93 ; Amer. J. Conch. 11. 

(1866), p. 210, pi. XIII. fig. 2. 

Hab. Kauai (Newcomb). 



(90) Amastra [Amastre/la) antiqiia Baldwin. 

Amastra antiqiia Baldwin, P. Ac. Philad. 1895, p. 233, pi. xi. fig. 47. 
Hab. Oahu, Ewa (Baldwin, as fossil). 



(91) Aviastra [Auiastrel/a) carinata Gulick. 

Amastra carinata Gulick, P. Zool. Soc. London, 1873, p. 83. 

Achatinella obesa var. agglutinans Newcomb, P. Zool. Soc. London, 1853 [1854], 

p. 143, pi. XXIII. fig. 39 «. 
This appears to be specifically distinct from A. obesa Newc. 
Hab. W. Maui, Wailuku (Gulick). 



(92) Amastra [Amastrella) cyclostoma Baldwin. 

Amastra cyclostoma Baldwin, P. Ac. Philad. 1895, p. 234, pi. xi. fig. 53 [animal 

and shell]. 
Hab. Kauai, Makaweli (Baldwin). 



(93) Amastra [Amastrella) nuclcola Gould. 

Achatinella luicleola Gould, P. Boston Soc. 11. (1845), p. 28. 

Achatinella brevis Pfeiffer, P. Zool. Soc. London, 1845 [1846], p. 90. 

Hab. Kauai (Newcomb); Hanalei (Baldwin).— ? Oahu, Manoa Valley (Clessin, 
Nomenc. Helic. Viv.). 

I feel doubtful as to this last locality : the A. njicleola Gould, of Reeve, is 
A. albolabj'is Newc. (cf. p. 333). 

46 — 2 



354 FAUNA HAWAIIENSIS 



(94) Amastra {Amastrella) obesa Newcomb. 

Achatinella obesa Newcomb, Ann. Lye. New York, vi. (May, 1853), p. 24; t. c. 
p. 329 [animal]; P. Zool. Soc. London, 1853 [1854], p. 143, pi. xxiii. fig. 39 ; 
Binney, Ann. Lye. New York, xi. p. 191, pi. xiv. fig. H [radula and jaw]. 

Hab. Maui, Makawao and Kula (Baldwin) ; Haleakala (Newcomb). 



(95) Amastra i^Ainastrelld) riigidosa Pease. 

Amastra riigulosa Pease, P. Zool. Soc. London, 1869, p. 649 (nom. sol.); 
J. Conchyl. xviii. (1870), p. 95 ; Crosse, 1. c. xxiv. (1876), p. 99, pi. i. fig. 4. 



var. similaris Pease. 

Amastra similaris Pease, P. Zool. Soc. London, 1869, p. 649 (nom. sol.). 
Amastra rugulosa var. similaris Pease, J. Conchyl. xviii. (1870), p. 96. 

Mr Perkins' specimens are small but otherwise agree with some presented by Pease 
to the British Museum. I have seen specimens collected by Mr Hutchison as from 
Oahu, but think there must be an error as to the locality. 

Hab. Kauai (Pease, type and var.) ; Kapaa (Baldwin) ; Lihue (Perkins). — E. 
Maui, Kula (Hartman) [? an error]. 

(96) Amastra {Amastrella) sphaerica Pease. 

Amastra sphaerica Pease, P. Zool. Soc. London, 1869, p. 649 (nom. sol.); 
J. Conchyl. xviii. (1870), p. 94; Crosse, I.e. xxiv. (1876), p. 98, pi. i. 
figs. 5, 5rt. 

Hab. Kauai (Pease). 

The habitat is given as "? Lanai " by both Mr Baldwin and Dr Hartman, but 
I know not on what authority. 

(97) Amastra {Amastrella) vctiista Baldwin. 

Amastra vettista Baldwin, Cat. Shells Hawaiian Islands, 1893, p. 10 (nom. sol.); 

P. Ac. Philad. 1895, P- '^11^ ?'• -'^i- %• 5°- 
Hab. Oahu, near the base of Punchbowl Hill, Honolulu, fossil (Baldwin). 



MOLL use A 355 



siibgen. Kauaia, nom. nov. 

Carinella Pfr. (1875) nee Sowerby (1839). 

The type of Pfeiffer's group is A. kauaiensis Newc, : the subgeneric name was 
used first by Sowerby for a different group of Molluscs. Whether A. alata and 
A. helicifortnis really belong here I am not clear. 



(98) Aniastra [Kauaia) alata Pfeiffer. 



1 -> 

JO- 



Helix alata Pfeift'er, P. Zool. Soc. London, 1856, p. 

I have elsewhere (P. Malac. Soc. London, 11. p. 127) pointed out that all authors 
have overlooked the fact that this shell has a columellar plait. It is, in my opinion, 
not a Helicoid at all, but belongs to an aberrant group of Amastra. The columellar 
plait does not ascend rapidly into the shell, but stands almost horizontally, and has no 
final 'knob.' The single specimen found by Mr Perkins measures diam. max. 8; alt. 4; 
alt. apert. 3 mill. 

Hab. Lanai (Pfeiffer) ; Mts. behind Koele (Perkins). 



(99) Amastra [Kajiaia) heliciformis Ancey. 

Amastra helicifonnis Ancey, Bull. Soc. Malac. France, vii. (1890), p. 340. 
Hab. Oahu, Waianae (Ancey). 

(100) Amastra [Kauaia) kauaiensis Newcomb. 

Achatinella kauaiensis Newcomb, Ann. Lye. New York, vri. (April, i860), 
p. 145; Amer. J. Conch. 11. (1866), p. 209, pi. xiii. fig. i. 

Achatinella [Carinella) kauaiensis Newc, Pfeiffer, Novit. Conch, iv. p. 115, 
pi. cxxvi. figs. 8 — 1 1. 

A good series, principally however dead shells, of this almost extinct species. 
Mr Perkins notes that one specimen was found "with embryonic shells in mouth." 

Hab. Kauai (authors) ; Halemanu (Baldwin) ; Makaweli at 2000 ft. and Hale- 
manu at 4000 feet (Perkins). 



356 FAUNA HAWAIIENSIS 

(loi) Amastra {Katiaia) knudseni Baldwin. 

Amastra knudseni Baldwin, P. Ac. Philad. 1895, p. 234, pi. xi. figs. 43, 44. 

Hab. Kauai, Halemanu (Baldwin, Perkins). A single specimen of this very 
fine species. 

The following appear to be only MS. names : Amasti^a ferrit,ginea Baldwin, 
Cat. Shells Hawaiian Islands, 1893, p. 9 (nom. sol.). Hab. Oahu, Ewa and Waianae 
(Baldwin). — Amastra testndinea Baldwin, t. c. p. 10. Hab. Oahu, Ewa (Baldwin). 

Leptachatina Gould. 

Leptachatina Gould, P. Boston Soc. 11. p. 201 ; type Achatinella acuminata 
Gould. 

It is frequently difficult to draw the line between this group and Amastra, and 
perhaps such species as A. melampoides Pfr. { = A. ventubis Per.) may eventually be 
transferred to Leptachatina. 

Pfeiffer proposed Labiella (Malak. Blatt. 1. 1854, p. 142) for the group with an 
incrassated lip, such as A. labiata Newc, and perhaps it may, conchologically, form a 
convenient section. 

The species are principally from Oahu, but an elongate and generally striate group 
characterizes the older Islands, such as Kauai. 



(i) Leptachatina accincta Mighels. 

Achatina accincta (err. typ.) Mighels, P. Boston Soc. 11. (1845), p. 20; Reeve, 

Conch. Icon. Achatina, sp. loi. 
nee Achatinella accincta Gould, U.S. Explor. Exped. Mollusca, pi. vii. fig. 97. 
Achatinella gj-anifera Gulick, Ann. Lye. New York, vi. (1856), p. 185, pi. vi. 

%• '3 ; Sykes, P. Malac. Soc. London, iii. pi. xiv. fig. 5. 
Achatinella {Leptachatina) margarita Pfeiffer, P. Zool. Soc. London, 1855, p. 206. 

Gulick admitted (P. Zool. Soc. London, 1873, p. 91) the identity of his species 
with Pfeiffer's. If Mighels' dimensions and description are accurate, I think the above 
identification will prove correct. The shell figured by Gould does not appear to be 
Mighels' species. See also a note under L. grana Newc. 

Hab. Oahu (Mighels, Pfeiffer) ; Keawaawa (Gulick). 



MOLLUSC A 357 

(2) Leptachatina acwninata Gould. 

Achatinella acuminata Gould, P. Boston Soc. 11. (1847), P- 200; U. S. Explor. 
Exped. Mollusca, pi. vii. fig. 100. 

Plate XII. figs. 13, i3«. 

The type of the genus ; the radula is figured from a dissection by Lt.-Col. H. H. 
Godwin-Austen, F.R.S. 

Hab. Kauai (Gould); Hanalei (Baldwin); Kaholuamano (Perkins). 

(3) Leptachatina antiqua Pease. 

Leptachatina antiqiui Pease, P. Zool. Soc. London, 1869, p. 651 (nom. sol.) ; 
J. Conchyl. xviii. (1870), p. 94 ; Crosse, J. Conchyl. xxiv. p. 98, pi. iii. fig. 6. 
Leptachatina antiquata Pease, J. Conchyl. xviii. (1870), p. 87 [err. typ.]. 
Hab. Kauai (Pease); Mana (Baldwin). 



(4) Leptachatina approxinians Ancey. 

Leptachatina approximans Ancey, Naturaliste, 1897, P- 222. 
Hab. Waianae, Oahu (Ancey). 

(5) Leptachatina arborea, n. sp. 

Testa ovato-oblonga, turrita, tenuis vel tenuiuscula, dextrorsa, cornea, longitudina- 
liter levissime striatula ; anfr. 6 — 7, plano-convexi, ultimus | altitudinis testae 
aequans ; sutura bene impressa ; apertura quadrato-ovata, margine dextro subincras- 
satulo, columellari verticali, incrassato, reflexo, plica parva vix conspicua munito. 
Alt. 8; diam. 3-6 mill. Plate XI. fig. 21. 

The plica is very inconspicuous; the colour becomes lighter in adult specimens, and 
then the polished, transparent gloss disappears and the shell becomes of a straw colour. 
Over thirty specimens were collected by Mr Perkins ; they vary slightly in shape, a 
few being broader in proportion to the length, and having more inflated whorls. 
Mr Baldwin sends me the following note: "It is found on the Candle-nut tree 
[Aleurites mohtccana), among the leaves of the Bird-nest fern {Asplcniuvi nidus), some- 
times at a height of 30 or 40 feet. All the other known species of Leptachatina are 
terrestrial — under rocks or on dead leaves and decaying wood." 

Hab. Hawaii, Kona at 4000 feet (Perkins); Olaa, Hilo (Baldwin). 



358 FAUNA HAWAIIENSIS 

(6) Leptachatina balteata Pease. 

Leptachatina balteata Pease, P. Zool. Soc. London, 1869, p. 651 (nom. sol); 
J. Conchyl. xviii. (1870), p. 91 ; Crosse, 1. c. xxiv. (1S76), p. 96, pi. iv. fig. 4. 

Four, apparently immature, specimens ; they approach this species very closely 
save that they do not possess the colour band, and the last whorl measures just over, 
rather than under, half the length of the shell. As the species is only known to me 
from description and figure, I think it safer to refer them here with a query than to 
describe them. 

Hab. Kauai (Pease); Wahiawa (Baldwin); at 4000 feet (Perkins). 

(7) Leptachatina brcznciila Pease. 

Leptachatina brevicula Pease, J. Conchyl. xvii. (1869), p. 169. 

Only known to me from the description. The specimens, while slightly larger 
than the dimensions stated by Pease, agree well with the proportions given. The 
plait, which he states is '" valida, fere transversa^' seems to vary much in size and 
prominence. 

Hab. Kauai (Pease) ; Kaholuamano, and at 4000 feet (Perkins). 

(8) Leptachatina [Labie/la] ca//osa Pteiffer. 

Achatinella [Labie/Ia] callosa Pfeiffer, P. Zool. Soc. London, 1856 [1857], p. 334. 
Only known to me from the unique type in the British Museum. 
Hab. Oahu (Pfeiffer). 

(9) Leptachatina ccrealis Gould 

Achatinella cerealis Gould, P. Boston Soc. 11. (1S47), p. 201 ; U. S. Explor. 
Exped. Mollusca, pi. \ii. fig. 99; Hartman, P. Ac. Philad. 1888, pi. i. fig. 13. 
Two specimens only, which, if not this species, are probably undescribed. 
Hab. Oahu, Waianae (Gould) ; Waianae Mts. below Kaala (Perkins). 

(10) Leptachatina chrysallis Pfeiffer. 

Achatina chrysallis Pfeiffer, P. Zool. Soc. London, 1855, p. 99. 

This species has been united with L. obtusa Newc, by Mr Baldwin, but appears 
to me to be quite distinct ; the habitat he gives of ' Wahiawa to Kawailoa, Oahu ' 
probably really refers to L. obtusa. 

Hab. Hawaiian Islands (Pfeiffer). 



MOLL use A 359 



(ii) Leptachatina cingiila Mighels. 

Achatinella cingula Mighels, P. Boston Soc. ii. (1845), p. 21. 

Leptachatina cingula Mighels, Hartman, P. Ac. Philad. 1888, pi. i. fig. 14. 

Achatinella {^Leptachatina) diviidiata Pfeiffer, P. Zool. Soc. London, 1855, p. 205. 

The L. cingula Mighels is unknown to me ; I quote the following from Mr 
Hartman, "Achatinella dimidiata Pfeiffer, equals cingula Migh. in coll. Newcomb ex 
Auct. The figure of this shell in Chemnitz [i.e. Conch. -Cab. Bulimacea, pi. lxvii. 
figs. 5 — 7J does not represent the species, but equals an Amastra" 

Hab. Oahu (Mighels, Pfeiffer) ; Halemano, Kawailoa Gulch (Perkins). 



(12) Leptachatina clausina Mighels. 

Bitlinms clausimis Mighels, P. Boston Soc. 11. (1845), p. 20. 

Leptachatina clausiana (sic) Mighels, Hartman, P. Ac. Philad. 1888, p. 52. 

Unknown to me. 

Hab. Hawaii (Mighels). 

(13) Leptachatina columna Ancey. 

Leptachatina columna Ancey, Naturaliste, 1889, p. 266; Sykes, P. Malac. Soc. 
London, in. pi. xiii. fig. 18. 

Near L. cJirysallis Pfr. 

Hab. Oahu (Ancey). 

(14) Leptachatina compact a Pease. 

Labiella conipacta Pease, J. Conchyl. xvii. (1869), p. 172. 

The specimens appear to agree with Pease's description ; the species has not been 
figured. 

Hab. Maui (Pease) ; E. Maui (Baldwin) ; Haleakala, at 5000 feet (Perkins). 

(15) Leptachatina conicoides, sp. nov. 

Testa conico-ovata, imperforata, dextrorsa, tenuiuscula, cornea, apud suturas 
crenulata ; anfr. 6, ultimus f altitudinis testae aequans ; sutura subimpressa ; apertura 
subverticalis, sinuato-oblonga, margine dextro sub-incrassatulo, columellari reflexo, 

F. H. II. 47 



36o FAUNA HAWAIIENSIS 

adnato, plica obliqua, parva, compressa munito, marginibus callo tenui junctis. Alt. 
7-5 ; diam. 3-5 mill. 

Plate XL fig. 26. 

A somewhat conic shell, in which, when adult, the columella plait becomes incon- 
spicuous. One adult and three young specimens. 

Hab. Molokai (Perkins). 

(16) Leptachatina convexiuscula, sp. nov. 

Testa turrita, elongata, gracilis, tenuiuscula, dextrorsa, brunneo-cornea, laevis, polita, 
nitida, apice obtusulo ; anfr. 6|, convexi, turgiduli, ultimus ^^g- altitudinis testae aequans ; 
sutura bene impressa ; apertura pyriformis, margine columellari sinuato, plica minima 
munito, marginibus callo tenuissimo junctis. Alt. 8 ; diam. 2 '8 mill. 

Plate XI. fig. II. 

A shell of the group of L. exilis Gulick, but with more inflated whorls, slightly 
more tapering towards the apex, and the mouth not so drawn out to the right. Three 
specimens. 

Hab. Oahu, Waiolani (Perkins). 

( 1 7) Leptachatina corneola Pfeiffer. 

Achatinella corneola Pfeiffer, P. Zool. Soc. London, 1845 [1846], p- 90. 
Achatinella corneola Pfeiffer, Reeve, Conch. Icon. Achatinella, sp. 4. 

Hab. Oahu ? (Baldwin) ; Oahu, one young specimen (Hutchison). 

(18) Leptachatina coruscans Hartman. 

Leptachatina cornscans Hartman, P. Ac. Philad. 1888, p. 52, pi. i. fig. 16. 

A variable shell in thickness and colouration. 

Hab. Molokai (Hartman) ; Kamalo (Baldwin) ; Kapanui, Kolamaula, and at 
4000 ft. (Perkins). 

(19) Leptachatina costulata Gulick. 

Achatinella costulata Gulick, Ann. Lye. New York, vi. (1856), p. 177, pi. vi. 
fig- 5 ; Sykes, P. Malac. Soc. London, iii. pi. xiv. fig. 4. 

Newcomb united this shell with L. semicostata Pfeiffer, but Gulick's type is much 
more slender than that species, the mouth is of a different shape, and other minor 
differences exist, all leading me to regard it as a good species. 

Hab. Oahu, Pupukea, Waimea, and Kawailoa (Gulick). 



MOLLUSC A 361 



(20) Leptachatina costulosa Pease. 

Leptacliatina costulosa Pease, P. Zool. Soc. London, 1869, p. 651 (nom. sol.); 
J. Conchyl. xviii. (1870), p. 90 ; Crosse, 1. c. xxiv. p. 96, pi. in. fig. 4. 

Hab. Kauai (Pease) ; Waimea and Kealia (Baldwin). 



(21) Leptachatina ciystal/ina Gulick. 

Achatinella crystallina Gulick, Ann. Lye. New York, vi. (1856), p. 186, pi. vi. 

fig. 14. 
Newcomb united this species with his L. nitida. 
Hab. Oahu, Mokuleia, Kamoo, Waialua (Gulick). 

(22) Leptachatina cmerita, sp. nov. 

Testa elongata, subcylindrica, imperforata, dextrorsa, cornea vel hyalina vel 
flava, tenuiuscula, sub lente longitudinaliter tenuiter striata, apice obtusulo ; anfr. 6^, 
plano-convexi, ultimus |- altitudinis testae fere aequans ; sutura impressa, marginata ; 
apertura ovata, margine dextro sub-incrassatulo, columellari sub-reflexo, plica parva 
inconspicua ascendente munito. Alt. 8 ; diam. 3 '5 mill. 

Plate XL fig. 10. 

Variable in colour, shading from brown to a hyaline tint ; adult specimens lose 
their gloss and become of a straw-yellow. The columellar plait is small and incon- 
spicuous. 

Hab. Molokai, Kalamaula, and at 4000 feet (Perkins). 

(23) Leptachatina exilis Gulick. 

Achatinella exilis Gulick, Ann. Lye. New York, vi. (1S56), p. 188, pi. vi. fig. 16 

[bad]; Sykes, P. Malac. Soc. London, in. pi. xiv. fig. 18. 
Leptachatina cylindrata Pease, J. Conchyl. xvii. (1869), p. 168; P. Zool. Soc. 

London, 1869, p. 650 (nom. sol.). 
Remarkable though the distribution may be, I am unable, after a comparison of 
Gulick's type with specimens of L. cylindrata presented by Pease to the British 
Museum, to sever these two species. 

Hab. Oahu, Keawaawa (Gulick).— Kauai (Pease) ; at 4000 feet (Perkins). 

47—2 



o 



62 FAUNA HAWAIIENSIS 



(24) Leptachatina extensa Pease. 

Leptachatina extensa Pease, P. Zool. Soc. London, 1869, p. 651 (nom. sol.); 
J. Conchyl. xviii. (1870), p. 92. 

Four specimens, agreeing fairly well with Pease's diagnosis, are referred to this 
species. 

Hab. Kauai (Pease) ; Kaholuamano (Perkins). 



(25) Leptachatina fiiniida Gulick. 

Achatinclla fnmida Gulick, Ann. Lye. New York, vi. (1856), p. 181, pi. vi. fig. 9 ; 
Sykes, P. Malac. Soc. London, in. pi. xiv. fig. 15. 

Newcomb united this with his L. vitrea, but they appear to me to be quite 
distinct. 

Hab. Oahu, Waialei, Pupukea, Waimea, Kawailoa, Halemano (Gulick). 



(26) Leptachatina fumosa Newcomb. 

Achatinclla fumosa Newcomb, P. Zool. Soc. London, 1853 [1854, Nov.], p. 140, 
pi. x.xiii. fig. 28. 

Hab. Oahu, Manoa (Newcomb) ; Kawailoa Gulch (Perkins). Only a single 
specimen. 



(27) Leptachatina fusca Newcomb. 

Achatinella fusca Newcomb, Ann. Lye. New York, vi. (1853), p. 28; P. Zool. 
Soc. London, 1853 [1854, Nov.], p. 145, pi. xxiii. fig. 44. 

Hab. Oahu, Manoa (Newcomb). 



(28) Leptachatina fuscida Gulick. 

Achatinella fuscula Gulick, Ann. Lye. New York, vi. (1856), p. 180, pi. vi. fig. 8. 
Hab. Oahu, mountain forests of Mokuleia (Gulick). 



MOLL USCA 363 

(29) Leptachatina glutinosa Pfeiffer. 

Achatinella i^Laminelld) glutinosa Pfeiffer, P. Zool. Soc. London, 1855 [1856, 

Feb.], p. 204. 
Achatinella lacrima Gulick, Ann. Lye. New York, vi. (1856, Dec), p. 176, pi. vi. 

fig. 4 ; Sykes, P. Malac. Soc. London, iii. pi. xiv. fig. 10. 
Hab. Oahu, Lihue, Kalaikoa, Wahiawa, Halemano, Peula (Gulick) ; Waianae 
Mts., below Kaala (Perkins). Only a single specimen. 

(30) Leptachatina gracilis Pfeiffer. 

Achatinella [Achatinellastnnn) gracilis Pfeiffer, P. Zool. Soc. London, 1855, p. 6, 

pi. xx.x. fig. 22. 
Achatinella elevata (Newcomb) Pfeiffer, t. c. [1856, Feb.], p. 209. 
Achatinella siibula Gulick, Ann. Lye. New York, vi. [1856, Dec], p. 191, pi. vi. 

fig. 19; Sykes, P. Malac Soc. London, in. pi. xiv. fig. 16. 
Gulick's species appears to be a bandless and slightly more attenuate variety. 
Hab. Oahu (various authors) ; Palolo Valley (Gulick) ; Kaala (Baldwin) ; 
Waianae Mts., below Kaala, lee side (Perkins). 

(31) Leptachatina grana Newcomb. 

Achatinella grana Newcomb, Ann. Lye. New York, vi. (1853), p. 29 ; P. Zool. Soc 

London, 1853 [1854], p. 146, pi. xxiii. fig. 46. 
Leptachatina grana Newcomb, Bland and Binney, Ann. Lye. New York, x. 

p. 336 [radula]. 
The types of this species have met with an accident and are entirely broken. 
Newcomb believed that L. granifera Gulick [= /-• accincta Mighels] was a 
synonym, but I feel doubtful of this. 

Hab. Maui, Makawao (Newcomb) ; Haleakala, at 5000 feet (Perkins). 

(32) Leptachatina guttnla Gould. 

Achatinella guttnla Gould, P. Boston Soc. 11. (1847), p. 201 ; U. S. Explor. 

Exped. Mollusca, pi. vii. fig. 98. 
Achatinella gummea Gulick, Ann. Lye. New York, vi. (1856), p. 182, pi. vi. 

fig. 10; Sykes, P. Malac. Soc London, iii. pi. xiv. fig. i. 
Achatinella fragi lis Gulick, t. c. p. 183, pi. vi. fig. ( i ; Sykes, t. c pi. xiv. fig. 2. 
Newcomb united — I think correctly — the two Gulickian species with Gould's ; 
Mr Baldwin has, however, in his 'Catalogue' allowed them specific rank. 

Hab. Oahu, Mokuleia, Lihue, Punaluu, Hauula, Halemano (Gulick).— Maui (Gould). 



364 FAUNA HAWAIIENSIS 



^}i'-i) Leptachatina imitatrix, sp. nov. 

Testa elongata, turrita, imperforata, dextrorsa, tenuiuscula, flavido-cornea, sub lente 
longitudinaliter minute striata ; anfr. 6^, plano-convexi, ultimus f altitudinis testae 
aequans ; sutura impressa ; apertura elongato-ovalis, margine dextro arcuato, acuto, 
columellari incrassatulo, reflexo, plica minima, inconspicua munito, marginibus callo 
tenui junctis. Alt. 7 ; diam. 2 '6 mill. 

Plate XI. fig. 9. 

Only a single specimen. It recalls L. exilis of Gulick, but is more conic — i.e. the 
upper whorls are narrower in proportion — and is of a light straw-yellow. The 
columellar plait is deeply-seated and inconspicuous. 

Hab. Hawaii, Mauna Loa at 4000 feet (Perkins). 



(34) Leptachatiiia impressa Sykes. 

Leptacliatina hnpi'essa Sykes, P. Malac. Soc. London, 11. (1S96), p. 127. 
Plate XI. ficr. 8. 

o 

Hab. Lanai, Mountains behind Koele (Perkins). 

(35) Leptachatina isthmica Ancey. 

Leptachatina isthinica Anc&y, P. Malac. Soc. London, iii. (1899), p. 270; Sykes, 
t. c. pi. XIII. fig. 20. 

Hab. Maui, Sand Hills between East and West Maui, subfossil (Ancey). 

(36) Leptachatina konaensis, sp. nov. 

Testa elongato-ovata, imperforata, dextrorsa, tenuiuscula, cornea vel pallide cornea, 
longitudinaliter tenuiter striata, apice obtusulo ; anfr. 6, planati, ultimus |- altitudinis 
testae aequans ; sutura impressa, marginata ; apertura sinuato-ovata, columella arcuata, 
margine dextro intus subcalloso, columellari subreflexo, plica mediocri ascendente 
munito. Alt. 8 ; diam. 4 mill. 

Plate XI. fig. 13. 

Akin to L. simplex Pease, but is much more swollen and inflated. Six specimens. 

Hab. Hawaii, Kona at 4000 feet (Perkins). 



MOLLUSC A 365 

{tjI^ Leptachatina i^Labielld) labiata Newcomb. 

Achatinella labiata Newcomb, Ann. Lye. New York, vi. (1853), p. 27; P. Zool. 
Soc. London, 1853 [1854], p. 141, pi. xxiii. fig. ^-^^ ; Gwatkin, P. Ac. Philad. 
1895, p. 239 [radula]. 
Achatinella lagena Gulick, Ann. Lye. New York, vi. (1856), p. 175, pi. vi. fig. 3 ; 

Sykes, P. Malac. Soc. London, iii. pi xiv. fig. 9. 
Achatinella dentata Pfeiffer, P. Zool. Soc. London, 1855, p. 7, pi. xxx. fig. 27. 
I follow Newcomb in including Gulick's species, but the latter's type does not 
fully shew the thickening on the columellar wall, nor the denticle on the outer lip. 

Hab. Oahu, Lehui (Newcomb) ; Halemano, Wahiawa, Kalaikoa (Gulick) ; 
Mount Kaala (Perkins). 

(38) I^eptachatina laevis Pease. 

Leptachatina laevis Pease, P. Zool. Soc. London, 1869, p. 651 (nom. sol.); 
J. Conchyl. xviii. (1870), p. 91 ; Crosse, 1. c. xxiv. (1876), p. 96, pi. iv. fig. 6. 
Hab. Kauai (Pease) ; Waimea (Baldwin). 

(39) Leptachatina Iciicoclnla Gulick. 

Achatinella leucochila Gulick, Ann. Lye. New York, vi. (1856), p. 173, pi. vi. 

fig. I ; Sykes, P. Malac. Soc. London, iii. pi. xiv. fig. 12. 
Newcomb united this with L. pyramis Pfr. ; I think it is quite distinct 
specifically. 

Hab. Kauai (Gulick). 

(40) Leptachatina lineolata Newcomb. 

Achatinella lineolata Newcomb, Ann. Lye. New York, vi. (1853), p. 29; P. Zool. 

Soc. London, 1853 [1854], p. 140, pi. xxiii. fig. 29. 
The real habitat seems somewhat uncertain ; Newcomb originally gave Maui, 
subsequently Hawaii, which is more probably correct. 

Hab. Maui (Newcomb and Hartman).— Hawaii (Newcomb and Baldwin). 

(41) Leptachatina lucida Pease. 

Leptachatina lucida Pease, J. Conchyl. xviii. (1870), p. 93. 
Hab. Kauai (Pease) ; Kealia (Baldwin). 



366 FAUNA HAWAIIENSIS 

(42) Leptachatina marginata Gulick. 

Achatinella marginata Gulick, Ann. Lye. New York, vi. (1856), p. 179, pi. vi. 

fig- 7- 
United by Newcomb with L. succincta Newc, but the present species is smaller 
and more slender. 

Hab. Oahu, Kalaikoa (Gulick). 

(43) Leptachatina nitida Newcomb. 

Achatinella nitida Newcomb, Ann. Lye. New York, vi. (1853, May), p. 29; 

P. Zool. Soc. London, 1853 [1854], p. 140. 
Leptachatiiia nitida Newcomb, Bland and Binney, Ann. Lye. New York, x. p. 336, 

pi. XV. fig. 8 [radula]. 

The figure given by Newcomb (P. Zool. Soc. London, 1853, pi. xxiii. fig. 30) 
apparently has been taken by error from some other shell, and does not represent this 
species. The form found by Mr Perkins appears to be a variety. 

Hab. E. Maui (Newcomb). — Maui and Oahu (Hartman). — Oahu, Mt Kaala 
(Perkins). 

(44) Leptachatina obsoleta Pfeiffer. 

Spii^axis obsoleta Pfeiffer, P. Zool. Soc. London, 1856, p. 335. 

A species of the group of L. sandiuicensis. Mr Perkins found a single young shell 
on ' Haleakala at 5000 feet,' Maui, which may be the young of this species. 
Hab. ? Oahu (Baldwin). 

(45) Leptachatina obtnsa (Newcomb) Pfeiffer. 

Achatinella obtusa Newcomb, Pfeiffer, P. Zool. Soc. London, 1855 [1856], p. 209. 

Mr Baldwin has suggested that this species is identical with L. chrysallis Pfeiffer, 
but I cannot agree with him. 

Hab. Hawaiian Islands. — ? Oahu (Baldwin). 

(46) Leptachatina octogyrata Gulick. 

Achatinella octogyrata Gulick, Ann. Lye. New York, vi. (1856), p. 190, pi. vi. 

fig. 18 ; Sykes, P. Malac. Soc. London, iii. pi. xiv. fig. 7. 
Newcomb placed it as a synonym of L. obclavata, Pfr. [ = Z. sandwicensis Pfr.]. 
Hab. Oahu, Palolo Valley (Gulick). 



MOLLUSC A 367 



(47) Leptachatina oryza Pfeiffer. 

Achatinella {Leptachatina) oryza Pfeiffer, P. Zool. Soc. London, 1855 [1856, Feb.], 

p. 206. 
Achatinella triticea Gulick, Ann. Lye. New York, vi. (1856, Dec), p. 184, pi. vi. 

fig. 12; Sykes, P. Malac. Soc. London, iii. pi. xiv. fig. 8. 
Hab. Oahu, subfossil (Pfeiffer) ; Keawaawa (Gulick). 



(48) Leptachatina {Labiella) pachystoma Pease. 

Labiella pachy stoma Pease, J. Conchyl. xvii. (1869), p. 171. 
I am not sure if this be a true Labiella. 
Hab. Kauai (Pease). 

(49) Leptachatina perkinsi Sykes. 

Leptachatina perkinsi Sykes, P. Malac. Soc. London, 11. (1896), p. 128. 

Plate XI. fig. 30. 

Hab. Lanai, Mts. behind Koele (Perkins). 



(50) Leptachatina petila Gulick. 

Achatinella petila Gulick, Ann. Lye. New York, vi. (1856), p. 189, pi. vi. fig. 17 ; 

Sykes, P. Malac. Soc. London, iii. pi. xiv. fig. 14. 
United by Newcomb with L. fusca Newc, but appears to me to be quite 
distinct. 

Hab. E. Oahu, Koko (Gulick). 

(51) Leptachatina pyramis Pfeiffer. 

Achatinella pyramis Pfeiffer, P. Zool. Soc. London, 1845 [1846], p. 90 ; Reeve, 

Conch. Icon. Achatinella, sp. 41 [good]. 
Appears from its form to be an Oahu species, and I am not sure that Pease's 
localization will prove correct. 
Hab. Kauai (Pease). 
F. H. II. 48 



368 FAUNA HAWAIIENSIS 



(52) Leptachatina resinula Gulick. 

Achatinella resinula Gulick, Ann. Lye. New York, vi. (1S56), p. 174, pi. vi. fig. 2 ; 
Sykes, P. Malac. Soc. London, iii. pi. xiv. fig. 11. 

Hab. Oahu, Kawailoa, Waialei, and other localities (Gulick). 

(53) Leptachatina saccula Hartman. 

Achatinella {Leptachatina) saccula Hartman, P. Ac. Philad. 1888, p. 55, pi. i. 

fig- 15- 
Hab. Hawaiian Islands (Hartman). 



(54) Leptachatina sandwicensis Pfeiffer. 

Achatina sandwicensis Pfeiffer, P. Zool. Soc. London, 1846 [May], p. 32. 
Achatinella {Leptachatina) obclavata Pfeiffer, Op. cit. 1855 [July], p. 98. 
Leptachatina octavula Paetel, Clessin, Nomenc. Helic. Viv. 1881, p. 316. 

Pfeiffer placed his Achatina sandwicensis in the synonymy of L. accincta Mighels ; 
the above identification is from an examination of Pfeiffer's types. 

Hab. Oahu (Pfeiffer) ; Waianae Mts. (Perkins). One young specimen only. 



(55) Leptachatina saxatilis Gulick. 

Achatinella saxatilis Gulick, Ann. Lye. New York, vi. (1856), p. 187, pi. vi. 

fig. 15 ; Sykes, P. Malac. Soc. London, iii. pi. xiv. fig. 17. 
Leptachatina saxitilus Gulick, Hartman, P. Ac. Philad. 1888, p. 55. 

Hab. Oahu, Mokuleia (Gulick). 



(56) Leptachatina sculpta Pfeiffer. 

Achatina sctdpta Pfeiffer, P. Zool. Soc. London, 1855 [1856], p. 211. 
Hab. Oahu (Pfeiffer) ; (Hutchison, one specimen). 

(57) Leptachatina scutilus Mighels. 

Bulimus scutilus Mighels, P. Boston Soc. 11. (1845), P- 20. 
Hab. Oahu (Mighels). 



MOLL use A 369 



(58) Leptackatina semicostata Pfeiffer. 

Achatinella {Leptachatina) semicostata Pfeiffer, P. Zool. Soc. London, 1855 [1856, 
Feb.], p. 206. 

Dr Hartman remarks (P. Ac. Philad. 1888, p. 55) " Dr Newcomb thinks it 
questionable if this species be not a synonym of L. fusca Newc." ; it is quite distinct. 
Hab. Hawaiian Islands (Pfeiffer). 



(59) Leptackatina semipicta Sykes. 

Leptackatina semipicta Sykes, P. Malac. Soc. London, 11. (1896), p. 128. 

Plate XI. fig. 12. 

Hab. Lanai, Mts. behind Koele (Perkins). 



(60) Leptackatina simplex Pease. 

Leptackatina simplex Pease, P. Zool. Soc. London, 1869, p. 651 (nom. sol.) ; 
J. Conchyl. xvii. 1869, p. 170. 

Dr Hartman notes (P. Ac. Philad. 1888, p. 55) that "Examples L. nitida Newc. 
(coll. Newc.) and L. simplex Pse. (coll. Pse.) are similar." There must be some error 
here, as specimens presented by Pease to the British Museum are quite distinct from 
L. nitida Newc. ; further, Newcomb's species does not come from Hawaii. 

Hab. Hawaii (Pease) ; Kona, at 3000 — 4000 feet (Perkins). 



(61) Leptackatina smitki Sykes. 

Leptackatina smitki Sykes, P. Malac. Soc. London, 11. (1896), p. 128. 

Plate XI. fig. 29. 

Hab. Lanai, Mts. behind Koele (Perkins). 



(62) Leptackatina stiria Gulick. 

Leptackatina stiria Gulick, Ann. Lye. New York, vi. (1856), p. 194, pi. vi. fig. 2. 
Hab. Oahu, Halemano, Peula, Kawailoa (Gulick). 

48—2 



370 FAUNA HAWAIIENSIS 

(63) Leptachatina striata Newcomb. 

Tornatellina striata Newcomb, P. Calif. Ac. 11. (1861), p. 93. 
From the description this appears to be close to L. hicida Pease. 
Hab. Kauai (Newcomb). 

(64) Leptachatina striatella Gulick. 

Achatinella striatella Gulick, Ann. Lye. New York, vi. (1856), p. 178, pi. vi. fig. 6 ; 

Sykes, P. Malac. Soc. London, iii. pi. xiv. fig. 19. 
United by Newcomb with L. fiisca Newc, but appears to me to be distinct. 
Hab. Oahu, Keawaawa (Gulick). 

(65) Leptachatina striatula Gould. 

Achatinella striatula Gould, P. Boston Soc. 11. (1845), p. 28. 

Achatinella clara Pfeiffer, P. Zool. Soc. London, 1845 [1846, Jan.], p. 90; Reeve, 
Conch. Icon. Achatinella, sp. 5. 

A nice series, shewing both the form with the sutural band and the unicolorous 
variety. 

Hab. Kauai (various authors); Makaweli, Kaholuamano, Lihue, and at 4000 ft. 
(Perkins). 

(66) Leptachatina succincta Newcomb. 

Achatinella succincta Newcomb, P. Boston Soc. v. (1855), p. 220; Amer. J. Conch. 

II. (1866), p. 213, pi. xiii. fig. 7. 
Leptachatina succinata Newcomb, Hartman, P. Ac. Philad. 1888, p. 55 (err. typ.). 

Hab. Oahu, Ewa (Newcomb) ; Halemano (Perkins). One specimen only. 

(67) Leptachatina supracostata, sp. nov. 

Testa elongata, turrita, imperforata, dextrorsa, tenuis, cornea, polita ; anfr. 8, 
ultimus ^ longitudinis testae fere aequans, primi apud suturas subcostulati, reliqui 
fere laeves ; sutura impressa, marginata, linea spirali notata ; apertura lunata, columella 
sub-arcuata ; margine dextro sub-incrassatulo, columellari sub-reflexo, plica minima 
oblique torta munito. Alt. 6*3 ; diam. 2 mill. 

Plate XI. fig. 22, 



MOLLUSC A 371 

Only two specimens. It belongs to the group of L. exilis Gulick ; is in- 
conspicuously costulate below the suture, the sculpture gradually fading out, until the 
last whorl hardly shews any marking beyond the lines of growth. There is a faint 
spiral line just below the suture. 

Hab. Lanai, Mts. behind Koele (Perkins). 

(68) Leptachatina tenebrosa Pease. 

Labiella tenebrosa Pease, P. Zool. Soc. London, 1869, p. 651 (nom. sol.). 
Leptachatina tenebrosa Pease, J. Conchyl. xviii. (1870), p. 92 ; Crosse, 1. c. xxiv. 

(1876), p. 96, pi. III. fig. 5- 
Hab. Kauai (Pease) ; Wahiawa (Baldwin) ; Kaholuamano, and at 4000 feet 
(Perkins). 

(69) Leptachatina tenuicostata Pease. 

Leptachatina temticostata Pease, J. Conchyl. xvii. (1869), p. 170. 
Hab. Hawaii (Pease). — Oahu (Baldwin). 

I feel doubts as to the accuracy of the last locality, as Mr Baldwin marks it as a 
species he has not seen. 

(70) Leptachatina terebralis Gulick. 

Achatinella terebralis Gulick, Ann. Lye. New York, vi. (1856), p. 193. P^- vi. 

fig. 21 ; Sykes, P. Malac. Soc. London, iii. pi. xiv. fig. 3. 
Hab. Oahu, Kawailoa (Gulick); Waianae Mts., below Kaala (Perkins). 

(71) Leptachatina teres Pfeiffer. 

Achatinella {Leptachatina) teres Pfeiffer, P. Zool. Soc. London, 1855 [1856], 

p. 206. 
Near L. obtusa Newcomb. 
Hab. Hawaiian Islands. 

(72) Leptachatina turgidula Pease. 

Labiella ttirgidula Pease, P. Zool. Soc. London, 1869, p. 651 (nom. sol.). 
Leptachatina turgidula Pease, J. Conchyl. xviii. (1870), p. 89; Crosse, 1. c. xxiii. 

(1876), p. 96, pi- IV. fig. 5- 
Hab. Kauai (Pease); Halemanu (Baldwin); Makaweli (Perkins). Five 

specimens. 



372 FAUNA HAWAIIENSIS 



{j-^ Leptachatina turrita Gulick. 

Achatinella turrita Gulick, Ann. Lye. New York, vi. (1856), p. 192, pi. vi. fig. 20 ; 

Sykes, P. Make. Soc. London, iii. pi. xiv. fig. 6. 
United by Newcomb with L. obclavata Pfr. [=^. sandivicensis Pfr.], but 
L. turrita is a broader and stouter shell, of a darker colour. 

Hab, Oahu, Lihue (Gulick). 



(74) Leptachatina vana sp. nov. 

Testa pyramidato-conica, dextrorsa, imperforata, tenuiuscula, brunneo-cornea, nitida, 
sub lente obsolete longitudinaliter striata, sutura marginata ; anfr. 8, lente accrescentes, 
ultimus I altitudinis testae aequans ; apertura pyriformis, margine dextro acuto, 
columellari sub-reflexo, sinuato, plica mediocri munito, marginibus callo tenuissimo 
junctis. Alt. 7-8; diam. 3-9 mill. Plate XL fig. 27. 

Four specimens of a brownish-horny, pyramidal shell, which has no striking 
characteristics. 

Hab. Oahu, Mt. Kaala (Perkins). 



(75) Leptachatina vitrea Newcomb. 

Achatinella vitrea Newcomb, P. Zool. Soc. London, 1853 [1854], p. 142, pi. xxiii. 

fig- 34- 
Hab. Oahu, Manoa (Newcomb). 



(76) Leptachatina vitreola Gulick. 

Achatinella vitreola Gulick, Ann. Lye. New York, vi. (1856), p. 194, pi. vi. fig. 23. 
Achatinella parvti la Gulick, t. c. p. 195, pi. vi. fig. 24; Sykes, P. Malac. Soc. 

London, in. pi. xiv. fig. 13. 
Both were united by Newcomb with his L. grana ; they appear to me to be quite 
distinct from that species. 

Hab. Hawaiian Islands (Gulick) ; W. Maui (Baldwin for L. parvula). 



MOLLUSC A 373 

Thaanumia Ancey. 

Tliaamiviia oiuphalodes Ancey. 

Thaanumia omphalodes Ancey, P. Malac. Soc. London, iii. (1899), p. 269, pi. xii. 

fig. 8. 
The type, and only, species. 
Hab. Oahu, Waianae Mountains (Ancey). 

Carelia H. and A. Adams. 

Carelia H. and A. Adams, Genera of Recent Mollusca, 11. (Feb. 1855) p. 132. 

This interesting genus, confined to Kauai save for one species on the Island of 
Niihau, was described by Messrs H. and A. Adams, with no named type. 

The anatomy has been described by Binney, P. Ac. Philad. 1876, p. 185, who 
points out that it agrees in general with the Amastra group, but differs in having a 
costate jaw. 

(i) Carelia bicolor Jay. 

Achat ina bicolor Jay, Cat. Shells, Ed. in. (1839), p. 119, pi. vi. fig. 3. 
Carelia bicolor Jay, Binney, P. Ac. Philad. 1876, p. 185, pi. vi. [anatomy]. 
Achatina adusta Gould, P. Boston Soc. 11. (1845), p. 26. 

Carelia adusta Gould, var. angulata Pease, J. Conchyl. xviii. (1870), p. 403. 
Achatina fuliginea Pfeiffer, P. Zool. Soc. London, 1852 [1854], p. 66; Conch.- 

Cab. Achatina, p. 267, pi. xliii. figs. 21, 22. 
Hab. Kauai (various authors). 

(2) Carelia cochlea Reeve. 

Achatina cochlea Reeve, Conch. Icon. Achatina, sp. 5. 

The spiral sculpture is nearly obsolete in some specimens ; I have seen one 
measuring 61 mill, in length. 

Hab. Kauai (various collectors). 

(3) Carelia cuiningiana Pfeiffer. 

Spiraxis cumingiana Pfeiffer, P. Zool. Soc. London, 1855, p. 106, pi. xxxii. fig. i. 
Hab. Kauai (Pfeiffer, &c.). 



374 FAUNA HAWAIIENSIS 

(4) Carelia dolei Ancey. 

Carelia dolei Ancey, Mem. Soc. Zool. France, vi. (1893), p. 328. 
Hab. Kauai, Hanalei (Ancey) ; Haena, subfossil (Baldwin). 

(5) Ca7'elia glutinosa Ancey. 

Carelia glutinosa Ancey, Mem. Soc. Zool. France, vi. (1893), P- 324- 
Hab. Probably Kauai. Unknown to me. 

(6) Carelia olivacea Pease. 

Carelia olivacea Pease, Amer. J. Conch. 11. (1866), p. 293. 

Carelia variabilis Pease, J. Conchyl. xviii. (1870), p. 402 [with var. vitadis']; 
P. Zool. Soc. London, 1871, p. 473. 

I do not quite follow why Pease described C variabilis, when, in the same paper, 
he stated that it and C. olivacea were varieties of one species. 

Hab. E. Kauai (Pease). 

(7) Carelia paradoxa Pfeiffer. 

Spiraxis paradoxa Pfeiffer, P. Zool. Soc. London, 1853, p. 128. 
Differs from all others known to me in its strongly granulated surface. 
Hab. Kauai. 

(8) Carelia sinclairi Ancey. 

Carelia sinclairi Ancey, Mem. Soc. Zool. France, v. (1892), p. 720. 
Hab. Niihau, subfossil (Ancey). 

(9) Carelia tuj-ricula Mighels. 

Achatina turriciila Mighels, P. Boston Soc. 11. (1845), p. 20. 

Carelia turricula Mighels, Kobelt, J. B. Malak. Ges. 11. (1875), p. 225, pi. vii. 

fig- I- 
Achatina obeliscus Reeve, Conch. Icon. Achatina, sp. 129. 
Achatina newcombi Pfeiffer, P. Zool. Soc. London, 185 1 [1853], p, 262. 

Hab. Kauai, Hanalei (Baldwin, Perkins). 



MOLL use A 375 

AuRicuLELLA Pfeiffer. 

Aurictdella Pfeiffer, P. Zool. Soc. London, 1855, p. i ; Mai. Blatt. 11. p. 3. 
The type appears to be the Partula auricula Fer. 

(i) Auriculella amditsta Pease. 

Auriculella ambusta Pease, J. Conchyl. xvi. (1868), p. 345, 
Probably the locality suggested by Mr Baldwin is correct. 
Hab. Oahu 1 (Baldwin). 



(2) Auriculella auricula Fer. 

Partula auricula Ferussac, Prodr. p. 66, no. 6 ; Voy. de Freycinet, Zool. 

p. 486. 
Auricula owaihiensis Chamisso, Nov. Act. Leop. xiv. (1829), p. 639, pi. xxxvi. 

fig. I. 
Auricula sinistrorsa Chamisso, torn. cit. p. 640, pi. xxxvi. fig. 2 [spec. juv.]. 
Partula dumartroyi Souleyet, Rev. Zool. v. (1842), p. 102. 
Bulimus armatns Mighels, P. Boston Soc. 11. (1845), p. 19. 

This species varies greatly in size and shape ; it is generally unicolorous, varying, 
from nearly white, through shades of yellow and green, to brownish green ; a few 
specimens have a single brown band. 

Hab. Oahu, Mount Tantalus, Mount Kaala, Halemano, Head of Kawailoa 
Gulch (Perkins). 



(3) Auriculella brnnnca Smith. 

Auriculella brtmnea Smith, P. Zool. Soc. London, 1873, p. 88, pi. x. fig. 23; 
Gwatkin, P. Ac. Philad. 1895, p. 238 [radula]. 

Two Lanai specimens have a single darker band at the periphery ; others are 
unicolorous. 

Hab. Molokai and Lanai (Smith); Molokai, Kalamaula, also Lanai, behind 
Koele (Perkins). 

F. H. II. 49 



376 FAUNA HAWAIIENSIS 

(4) Auriailella cerea Pfeiffer. 

Achatinella cerea Pfeiffer, P. Zool. Soc. London, 1855, p. 2, pi. xxx. fig. 21. 
Pease has suggested (J. Conchyl. xvi. p. 343) that this is identical with A. petitiana 
Pfeiffer ; he is not improbably correct, but I have only seen the single type specimen. 
Hab. Molokai (Nevill, fide specimens from Newcomb). 

(5) Auriculella chamissoi Pfeiffer. 

Achatinella [Auriculella) chamissoi Pfeiffer, P. Zool. Soc. London, 1855, p. 98. 
Hab. Oahu (Baldwin). — Hawaii (fide tablet in Brit. Mus.). 

(6) Auriculella crassula Smith. 

Auriculella crasstda Smith, P. Zool. Soc. London, 1873, p. 88, pi. x. fig. 22. 
Auriculella ponderosa Ancey, Bull. Soc. Malac. France, vi. (1889), p. 225. 

Hab. Maui, Makawao (Baldwin) ; lao Valley, Olinda, and Haleakala at 4000 feet 
(Perkins). 

(7) Auriculella diaphana Smith. 

Auriculella diaphana Smith, P. Zool. Soc. London, 1873, p. 87, pi. x. fig. 25. 
Auriculella patula Smith, torn. cit. p. 88, pi. x. fig. 24. 

Hab. Oahu, various localities (Smith) ; Mount Tantalus, and head of Panoa 
Valley (Perkins). 

(8) Auriculella expansa Pease. 

Auriculella expansa Pease, J. Conchyl. xvi. (1868), p. 343, pi. xiv. fig. 8. 
Hab. Hawaiian Islands (Pease). — Probably Maui (Ancey). — Kauai (Baldwin). 

(9) Auriculella lurida Pfeiffer. 

Tornatellina castanea Pfeiffer, Mon. Helic. Viv. in. (1853), p. 524. 
Achatinella (Auriculella) lurida Pfeiffer, Mon. Helic. Viv. iv. p. 570. 

Pfeiffer re-named the species, apparently to avoid confusion with Achatinella 
castanea Reeve. 

Hab. Maui ? (Baldwin). — Oahu, Mount Tantalus (Perkins). 



MOLLUSC A 377 



(lo) Auriculella newconibi Pfeiffer. 



Balea neivcovibi Pfeiffer, P. Zool. Soc. 1852- [1S54], p. 67. 
AchatineUa obeliscus Pfeiffer, Malak. Blatt. 11. (1855), p. 166. 
Hab. Molokai, Kalamaula (Perkins). 



(11) Auriculella obliqua Ancey. 

Auriadella obliqua Ancey, Mem. Soc. Zool. France, v. (1S92), p. 721 ; Sykes, 
P. Malac. Soc. London, iii. p. 275, pi. xiii. fig. 17. 

Appears to be very near A. a))ibiista Pease. 

Hab. Oahu, Waianae Mts. (Baldwin). 



(12) Auriculella perkinsi sp. nov. 

Testa subperforata, elongato-conica, brunnea aut corneo-brunnea, linea brunnea 
ad peripheriam saepe notata, nitida ; anfr. 6 — 6^, planiusculi, ultimus f altitudinis 
testae aequans ; apertura auriformis, intus brunnea, margine parietali lamina obliqua 
intrante, columellari lamina volvente munitis; peristoma leviter reflexum, incrassatulum. 
Alt. 8 ; lat. 4 mill. 

Plate XI. figs. 17, 18. 

van a. Magis elongata et tenuior, flavida, peristomate albido. 

I cannot identify this species with any of the numerous varieties of A. atiric7ila, 
and therefore describe it. It is very variable in colour, shading from rich brown to 
light yellow : when brown the band — if present — is yellowish, and conversely. The 
lip varies in colour from dark brown to white. It is a fairly thin shell and appears to 
be common. 

Hab. Oahu, ridges round Nuuanu, and Mount Tantalus (Perkins). 



(13) Auriculella petitiana Pfeiffer. 

Tornatellina petitiana Pfeiffer, Zeitsch. Malak. iv. (1847), p. 149 ; Kuster, Conch. - 
Cab. Tornatellina, p. 153, pi. xviii. figs. 24, 25. 

Hab. Hawaiian Islands. 
See a note under A. cerea. 

49—2 



378 FAUNA HAWAIIENSIS 

(14) Auriculella perpusilla Smith. 

Auriadella perptisilla Smith, P. Zool. Soc. London, 1873, p. 87, pi. x. fig. 26. 
Hab. Oahu, Kahalu (Smith). 

(15) Auriculella pule hr a Pease. 

Auriculella pulchra Pease, J. de Conchyl. xvi. (1868), p. 346, pi. xiv. fig. 6. 

Specimens presented by Pease to the British Museum under this name do not 
quite agree with his diagnosis, and his figure appears to have been drawn from a 
variety which he notes, and not the type form. I have followed the identified specimens ; 
possibly it is a variable species, or an error may have occurred in translating his paper. 

Hab. Oahu (authors); Mount Tantalus and Mount Kaala (Perkins). 

(16) Auriculella tenella Ancey. 

Auriculella tenella Ancey, Bull. Soc. Malac. France, vi. (1889), p. 232. 
Hab. Oahu, Waianae (Ancey). 

(17) Auriculella tenuis Smith. 

Auriculella tenuis Smith, P. Zool. Soc. London, 1873, p. 87, pi. x. fig. 27. 
Mons. Ancey has described (Bull. Soc. Malac. France, vi. p. 230) a var. solida. 
Hab. Oahu, various localities (Smith). 

(18) Auriculella triplicata Pease. 

Auriculella triplicata Pease, J. de Conchyl. xvi. (1868), p. 346. 
Hab. Maui (Hartman). — Oahu, Tantalus and Panoa (Baldwin). 

(19) Auriculella utiiplicata Pease. 

Auriculella uniplicata Pease, J. de Conchyl. xvi. (1868), p. 344, pi. xiv. fig. 7. 
Hab. Maui (Pease) ; West Maui (Baldwin). — ^Molokai, Kalamaula, and above 
Pelekunu (Perkins). 



MOLLUSC A 379 

(20) Auriculella westerlundimia Ancey. 

Auriculella wesier/iindiana hncay, Bull. Soc. Malac. France, vi. (1889), p. 218; 
Sykes, P. Malac. Soc. London, iii. p. 275, pi. xiii. fig. 21. 

Hab. Hawaii, Kona, and Waimea (Ancey) ; Kona at 3000 feet, and Olaa 
(Perkins). 



Insufficiently known or erroneously recorded species. 

The following appear to be only manuscript names : Jucunda Smith ; solida 
Gulick ; solidissima Smith (confer Ann. Lye. New York, x. pp. 331 — 2). 

Bulimus pumicatus MigheLs, P. Boston Soc. 11. p. 19. 

Hab. Oahu. 

Probably this is really an Aiirimlella ; I am totally unacquainted with it. 

Partula pusilla Gould, P. Boston Soc. 11. p. 197; U. S. Explor. Exped. Mollusca, 
pi. VII. fig. 90. 

This species has been referred to Aurictdella, and consequently a Hawaiian 
habitat has been suggested for it ; it is really, however, a Tornatellina and was 
described from Metia [ = Mata]. 



Frickella Pfeiffer. 
Frickella Pfeiffer, P. Zool. Soc. London, 1855, p. 2 ; Mai. Blatt. 11. p. 3. 

Frickella ainoena Pfeiffer. 

Achatinella {^Frickella) amoena Pfeiffer, P. Zool. Soc. London, 1855, p. 2, pi. xxx. 
fig- 3- 

This aberrant species appears to be a link between Achatinella and Tornatellina. 
The single young shell, found by Mr Perkins, does not quite agree with the type, as 
the whorls are flatter, but I am unable to sever it specifically. 

Hab. Oahu, Konahuanui (Baldwin) ; Halemano (Perkins). 



38o FAUNA HAWAIIENSIS 

TORNATELLINIDAE. 

TORNATELLINA Beck. 

Beck (Index Moll. 1837, p. 80) proposed this name as a subgenus oi Achatina, and 
placed in it four species, all of them undiagnosed. Pfeiffer in 1841 (Symb. Hist. Helic. 
pt. 2, p. 5) diagnosed the genus and gave (p. 130) a list of species. Previously to this 
Anton had proposed (1839) Strobilus, but in considering his claims it should be borne 
in mind that Strobila had twice previously been used in Zoology. 

The Hawaiian species appear to be but little understood : the only attempt at 
figuring them was made by Gould, whose six figures, under one name, represent three 
different species. 

I have endeavoured to avoid the creation of synonyms by a careful study of the 
descriptions and measurements given by the various authors. The habitat in the case 
of these very small shells is not always reliable, as they are very liable to be transported 
with plants, &c. 

( 1 ) Tornatellina baldwini Ancey. 

Tornatellina baldwitti Anc&y, Bull. Soc. Malac. France, vi. (1889), p. 238. 
Hab. Oahu, Tantalus (Ancey) ; Waianae Mts. (Perkins). — Kauai (Baldwin). 

(2) Tornatellina compact a sp. nov. 

Testa perforata, ovata, brunneo-cornea, nitidula, tenuis ; spira curta, apice obtusulo ; 
anfr. 5 — 5^, lineis incrementi bene notati, convexiusculi, regulariter et lente crescentes, 
sutura bene impressa ; apertura ovato-pyriformis, lamina unica pygmaea volventi. 
interdum praedita ; peristoma simplex, margine columellari reflexo et expanso. 
Alt. 2'2, diam. I'l mm. 

Plate XL fig. I. 

A compressed, compact little form, the aperture measuring about |- of the length ; 
the whorls are somewhat convex. 

Hab. Hawaii, Mauna Loa at 2000 feet, on hilo grass (Perkins). 

(3) Tornatellina confiisa sp. nov. 

Pupa peponum Gould, P. Boston Soc. 11. (1847), p. 197; U. S. Explor. Exped. 
Mollusca, pi. VII. figs. 104 a — c. 

See for remarks under Tornatellina peponum Gould ; this is the edentulous form 
figured by him. 

Hab. Kauai, Makaweli (Perkins). 



MOLLUSC A 381 

(4) TornatcUitia cylindrica sp. nov. 

Testa elongata, cylindrica, cornea, perforata ; anfr. 5 — 5^, convexiusculi, striatuli, 
ultimus rotundatus, |- altitudinis testae aequans, sutura impressa ; apertura ovata vel 
lunaris, lamellam in pariete gerens ; columella incrassata, albida, contorta, interdum 
denticulo mediocri munita. Alt. 2 '2, lat. vix i mm. 

Plate XI. fig. 28. 

This species may be distinguished from the true T. pepomun, by its smaller size 
and more slender shape. 

Hab. Oahu, Waianae Mts. (Perkins). — Kauai, Makaweli, one specimen (Perkins). 

(5) Tornatcllina dentata Pease. 

Tornatellina dentata Pease, P. Zool. Soc. London, 1871, p. 460. 
I identify Mr Perkins' specimens with some doubt ; if not this species they 
belong to no other recorded Hawaiian form. 

Hab. Hawaii (Pease) ; Puna (Baldwin) ; Kona at 3000 feet (Perkins). 

(6) Tornatellina euryomphala Ancey. 

Tornatellina euryomphala Ancey, Bull. Soc. Malac. France, vi. (1889), p. 239. 
Not found by Mr Perkins ; I have specimens from another source, without indica- 
tion as to which island they come from. 
Hab. W. Maui (Ancey). 

(7) Tornatellina extinct a Ancey. 

Tornatellina extinda Ancey, Bull. Soc. Malac. France, vii. (1890), p. 341. 
Hab. Central Maui, subfossil (Ancey). 

(8) Tornatellina gracilis Pease. 

Tornatellina gracilis Pease, P. Zool. Soc. London, 1871, p. 460. 

A single shell, found by Mr Perkins, agrees well with Pease's description and 
measurements, save that Pease speaks of the shell being sometimes spirally sulcate, 
while Mr Perkins' specimen shews traces of a single spiral thread at the periphery. 

Hab. Kauai (Pease).—? Hawaii, Kona at 3000 feet (Perkins). 



382 FAUNA HAWAIIENSIS 

(9) Tornatellina neiucombi PfeifFer. 

Tornatellina newcombi Pfeiffer, P. Zool. Soc. London, 1856, p. 335. 

I am not quite clear if the localities are to be relied on ; the figure given by 
Gould (as T.peponum, U. S. Explor. Exped. Moll. pi. vii. fig. 104 e) does not, I think, 
represent this species, as has been suggested. 

Hab. Maui and Oahu (Ancey).— Kauai, Oahu, and Maui (Baldwin). 

(10) Tornatellina oblonga Pease. 

Tornatellina oblonga Pease, P. Zool. Soc. London, 1864, p. 673; Binney, Ann. 

Lye. New York, xi. p. 190 [radula]. 
Tornatellina bacillaris Mousson, J. Conchyl. xix. (1871), p. 16, pi. in. fig. 5. 
Tornatellina oblongata Pease, Clessin, Norn. Helic. Viv. 1881, p. 343 (err. typ.). 
Unknown to me as Hawaiian ; it was described from the Tonga Islands. 
Hab. Oahu, Manoa (Ancey). 

(11) Tornatellina peponum Gould. 

Pupa peponum Gould, P. Boston Soc. 11. (1847), p. 197; U. S. Explor. Exped. 
Mollusca, pi. VII. figs. 104, 104 d. 

Gould has undoubtedly confused three species under this name : which it therefore 
becomes necessary to restrict to one of his forms. I propose that it should be used 
for the shells figured by him as fig. 104 and fig. 104^^; namely the slender species with 
a parietal lamina and no columellar tooth : of this I have Hawaiian specimens. 

The next form, that figured as figs. 104 « — c, has no parietal lamina, and equally 
no teeth on the columella ; this has been found by Mr Perkins on Kauai, and is here 
named T. confusa. 

The third form, figured as fig. 104 e (enlargement of mouth only) is a shell nearly 
related to T. euryomphala, Ancey ; it is not, I think, T. newcombi. 

Hab. Hawaii, Hilo, also Oahu (Gould). 

(12) Tornatellina perkinsi, sp. nov. 

Testa elongato-pyramidalis, subperforata, cornea, sub lente striatula ; anfr. 7, 
planiusculi, lente accrescentes, sutura subimpressa ; apertura ovata, lamellam sat minu- 
tam in pariete gerens ; columella incrassata, albida, plicis duabus inconspicuis praedita. 
Alt. 3, lat. 1-2 mm. 

Plate XI. fig. 14. 



MOLLUSC A 383 

Its most noteworthy features are the very elongately pyramidal shape, flattened 
whorls, and the two inconspicuous pHcae on the columella, the upper one being more 
deeply seated than the lower. 

Hab. Kauai, Kaholuamano at 4000 feet (Perkins). 



(13) Tornatellina trochoides sp. nov. 

Testa profunde perforata, cornea, pyramidalis, pellucida, fere laevis ; spira conoidea, 
apice acutiusculo ; anfr. 7 — 7^, regulariter lenteque accrescentes, planiusculi, ultimus 
ad peripheriam subcarinatus, sutura impressa ; apertura quadrato-ovata, superne angu- 
lata, laminam conspicuam validam in pariete gerens ; columella brunnea, incrassata, 
plicis duabus munita. Alt. 4, lat. 2 mm. 

Plate XI. fig. 31. 

Closely related, apparently, to T. tnubilicata Ancey ; but differs from it in the 
relative proportions of height and breadth. 

Hab. Lanai Mountains (Perkins). 



(14) Tornatellina umbilical a Ancey. 

Auriculella ttmbilicata Ancey, Bull. Soc. Malac. France, vi. (1889), p. 232. 
Tornatellina ttmbilicata Ancey, Mem. Soc. Zool. France, v. p. 721. 

Hab. Maui, Lahaina (Ancey). 



Fam. STENOGYRIDAE. 
Opeas Albers. 

Opeas K\h&rs, Die Heliceen, 1850, p. 175; Martens, Die Heliceen, Ed. 2, i860, 
p. 265 (type Buliniiis goodalli Miller). 

( I ) Opeas j mucus Gould. 

B !t I iums junceiis Go\Ad, P. Boston Soc. 11. (1847), p. 191 ; U. S. Explor. Exped. 

Moll. pi. VII. fig. 87. 
Hab. Hawaiian Islands (Gould); all the islands (Baldwin). — Oahu, Waianae 
Mts. (Perkins). 

F. H. 11. so 



384 FAUNA HAWAIIENSIS 

(2) Opeas pj'estoni Sykes. 

Opeas prestoni Sykes, P. Make. Soc. London, iii. (1898), p. "Ji, pi. v. fig. 4. 

A long series, which I am entirely unable to sever from the shell recently 
described by me from Ceylon. The whole genus, however, is notorious for having 
been discovered in widely separated localities, and probably this species has been 
transported with plants. 

Hab. Hawaii, Kawailoa, Mauna Loa at 1500 feet (Perkins). 

(3) Opeas pyrgiscus Pfeiffer. 

Bulimus pyrgiscus Pfeiffer, P. Zool. Soc. London, 1861, p. 24; Malak. Blatt. 

1861, p. 15; Novit. Conch, in. p. 425, pi. xcvi. figs. 10 — 12. 
This form does not seem to have been found in recent years. 

Hab. Hawaiian Islands (Pfeiffer). 

Opeas striolata Pease, is recorded as Hawaiian by Nevill (Handlist Ind. Mus. 
pt. I. 1878, p. 166) ; it appears to be a manuscript name only. 

Caecilioides (Blainville) Herrmannsen. 

(i) Caecilioides baldwini Ancey. 

Caecilianella baldwini Ancey, Mem. Soc. Zool. France, v. (1892), p. 718. 
I have followed Mr Smith (J. Conch, vi. pp. 341—342) as to the generic name. 
Hab. Oahu, Manoa (Ancey). 

Fam. SUCCINEIDAE. 
SucciNEA Draparnaud. 

S^tccinea Draparnaud, Tabl. des Moll. i8oi,p. 55 (first species S. amphibia Drap.). 

The large number of unfigured species described from the Hawaiian Islands 
renders the identification of specimens belonging to this puzzling group by no means 
an easy task. 

Pease proposed (J. Conchyl. xviii. 1870, p. 89) the genus Catinella for Succinea 
rubida; also the genus Tmella (P. Zool. Soc. London, 1871, p. 459) for S. elongata. 
I have thought it best to leave all the forms in Succinea, and have listed the species in 
alphabetical order. 



MOLL use A 385 

Two species collected by Mr Perkins still await identification ; they are not 
improbably new, but so many of the described species are unknown to me that I 
consider it safer to leave them for the present. One is from Honolulu and Waianae 
Mts. in Oahu, with a dwarf variety from the mountains of Molokai at 4000 feet ; the 
other from Kau, Hawaii : both belong to the group of S. canella Gould. 

( I ) Succinea aurulenta Ancey. 

Succinea aurulenta Ancey, Bull. Soc. Malac. France, vi. (1889), p. 242 ; Sykes, 
P. Malac. Soc. London, in. p. 275, pi. xiii. fig. 4. 

Hab. Hawaii, North Kona (Baldwin); Kona at 3000 feet (Perkins). 

(2) Succinea baldwini Ancey. 

Sticcinea baldwini Ancey, Bull. Soc. Malac. France, vi. (1889), p. 250. 

The specimens are identified from the description alone. 

Hab. Maui, Lahaina (Baldwin); Haleakala at 9000 feet (Perkins). 

(3) Sticcinea bicolorata Ancey. 

Succinea bicolorata Ancey, P. Malac. Soc. London, in. (1899), p. 271, pi. xii. 

fis:. 2. 
Hab. Hawaii, Waimea (Ancey). 



(4) Succinea cadiica Mighels. 

Sticcinea caduca Mighels, P. Boston Soc. 11. (1845), p. 21 ; Gould, U. S. Explor. 
Exped. Mollusca, pi. n. fig. 30. 

Hab. Oahu, Waianae Mts. (Baldwin).— Molokai Mts. (Perkins). — Lanai Mts. 
(Perkins). 

(5) Succinea canella Gould. 

Succinea canella Gou\d, P. Boston Soc. n. (1847), p. 184; U. S. Explor. Exped. 
Mollusca, pi. II. fig, 20 ; Bland and Binney, Ann. Lye. New York, x. p. 338 
[jaw and radula]. 

Mons. Ancey has named (Bull. Soc. Malac. France, vi. pp. 245 — 7) varieties 
crassa, obesiila, mamillaris, and liicida. 

Hab. Maui, Lahaina (Baldwin). — Molokai and Maui (Ancey). — Maui, Haleakala 
at 5000 feet; Molokai Mountains (Perkins). 

50 — 2 



386 FAUNA HAWAIIENSIS 

(6) Succinea casta Ancey. 

Succinea casta Ancey, P. Malac. Soc. London, iii. (1899), p. 272, pi. xii. fig. 10. 
Hab. Hawaii, Olaa (Ancey). 

(7) Succinea cepulla Gould. 

Succinea cepulla Gould, P. Boston Soc. 11. (1847), p. 182; U. S. Explor. Exped. 

Mollusca, pi. II. fig. 15. 
Succinea fragilis Souleyet, Voy. Bonite, Zool. 11. (1852), p. 501, pi. xxviii. figs. 

18 — 20 [shell and animal : nee S. fragilis King]. 
Sticcinea souleyeti Ancey, Bull. Soc. Malac. France, vi. (1889), p. 255. 

The synonymy given here and under S. rotundata has been arrived at from a 
careful comparison of the original descriptions and figures, and a study of the identifica- 
tions made by Pfeiffer amongst the specimens in the British Museum. The shell 
figured by Reeve (Conch. Icon. Succinea, fig. 69), purporting to be Hawaiian, under 
the name of S. fragilis King, is not the present species. 

Hab. Hawaii (Gould, &c.). — Oahu, Tantalus, Head of Panoa Valley (Perkins). — 
Molokai Mountains (Perkins). 

(8) Succinea cinnamomea Ancey. 



Succinea cinnamomea Ancey, Bull. Soc. Malac. France, vi. (1889), p. 247. 

Plate XL fig. 32. 

Hab. Oahu, Waianae Mts. (Baldwin) ; Mount Kaala (Perkins). 



(9) Succinea delicata Ancey. 

Succinea delicata Ancey, Bull. Soc. Malac. France, vi. (1889), p. 243. 
Hab. East Maui (Ancey);. Kula (Baldwin). 

(10) Succinea elongata Pease. 

Succinea elongata Pease, J. Conchyl. xviii. (1870), p. 96. 

A single specimen was found by Mr Perkins ; the species was only known to me by 
the description. Authors refer to a 6". elongata Beck, but I have failed to trace the 
reference. 

Hab. Kauai (Pease) ; Waimea (Baldwin) ; Kaholuamano (Perkins). 



MOLL use A 387 

(11) Succiiiea cxplanata Gould. 

Succinea explaiiata Gould, U. S. Explor. Exped. Mollusca, p. 13, pi. ii. fig. 31. 
Hab. Kauai (Gould); North side (Baldwin). 

(12) Succinea garrettiana Ancey. 

Succinea garrettiana Ancey, P. Malac. Soc. London, iii. (1899), p. 272, pi. xii. 

fig- /• 
Hab. Hawaii, Rainbow Falls, Hilo (Ancey). 

(13) Succinea inconspicua Ancey. 

Succinea inconspiciia Ancey, P. Malac. Soc. London, in. (1899), p. 273, pi. xii. 

fig- 9- 
Hab. Hawaii, Waimea (Ancey). 

(14) Succinea konaensis Sykes. 

Succinea konaensis Sykes, P. Malac. Soc. London, 11. (1897), p. 299. 

Plate XI. fig. 34. 

Hab. Hawaii, Mount Kona at 4000 feet (Perkins). 

(15) Succinea lumbal is Gould. 

Succinea lumbalis Gould, P. Boston Soc. 11. (1847), p. 183; U. S. Explor. 
Exped. Moll. pi. II. fig. 18. 

Hab. Hawaii, Mauna Kea (Baldwin). — Kauai (Gould) ; Makaweli, and above 
Waimea at 4000 feet (Perkins). 

The specimens from 'above Waimea' have a more drawn out spire, but only 
belong, I think, to a varietal form. 

(16) Succinea lutulenta Ancey. 

Succinea lutulenta Ancey, Bull. Soc. Malac. F" ranee, vi. (1889), p. 244. 
Hab. Maui (Ancey) ; Ulapalakua (Baldwin). 



3SS FAUNA HAWAIIENSIS 

(17) Slice inca mauicnsis Ancey. 

Succinea ntauiensis Ancey, Bull. Soc. Malac. France, vi. (1SS9), p. 24S. 
Hab. INIaui (Ancey) ; Makawao (Baldwin) ; Haleakala at 5000 feet (Perkins). 



(iS) Siicctiica neiocombianci Garrett. 

Succinea netocombiana Garrett, P. Calif. Ac. i. (1S57), p. 103. 

A species closely related to 5". ccpulla and 6". rohtndata, but. if my identification of 
the latter is correct, this is smaller, the mouth is rounder, and the last whorl not so 
much inflated. 

Hab. Hawaii, district of Waimea (Garrett) ; Kohala Mountains (Perkins). 

( 1 9) Succinea protracta sp. now 

Testa tenuis, aureo-rufa vel pallide-cornea, lineis increment! obliquis bene 
notata : spira producta, apice acutulo, mamillato ; anfr. 3^ con^■exi, sutura bene 
impressa discreti, rapide accrescentes : apertura o\ato-pyriformis, fere recta ; peristoma 
simplex, tenue, margine columellari arcuato, baud plicato. Long. 127, diam. maj. 6; 
alt. ap. 8. laL ap. 5 mm. 

Plate XI. fig. 25. 

Belongs to the group of 5". aurulenta Ancey, but is much more elongate and 
slender, and the suture is more impressed. A single white specimen from ' Mauna Loa 
at 3500' feet I also refer to this species. 

Has. Hawaii, Kau ^^ Perkins). 

(20) Succinea punctata Pfeiffer. 

Succinea punctata Pfeiffer, P. Zool. Soc. London, 1854 [May, 1855], p. 297; 
Reeve, Conch. Icon. Succinea, fig. 29. 

The specimens from Kohala Mts. are young and have the spire slightly more 
depressed, but appear to belong to this form. The white spots shewn in Reeve's 
figure are much exaggerated : most specimens have a few blotches of a paler tint, but I 
have seen none regularly spotted in the way the artist has represented them. 

Hab. Ha^^•aii (Pfeiffer) : Kohala Mountains, Olaa Puna, and Kona at 3000 feet 
(Perkins). 



MOLL use A 389 



(21) Succinea roimtdala Gould. 

Succinea patula Mighels, P. Boston Soc. 11. (1845J, P- 21 [nee S. patula Brug,, 

nee King]. 
Sziccinea rotundata Qo\Ai., P. Bcston Soc. 11. (1847), p. 182 ; U. S. Explor. Exped. 

Moll. pi. II. fig. 14 [shell and animal]. 
Succmea newcomhi Pfeiffer, P. Zool. Soc. London, 1854 [May, 1855], p. 297; 

Novit. Conch, i. pi. ix. figs. 3 — 5 ; Reeve, Conch. Icon. Succinea, fig. 61. 

Hah. Oahu (Gould, &c.).— Molokai (Pfeiffer).— Hawaii, Kohala (Perkins). 

See the remarks under S. cepulla, which species seems to be, like the present one, 
widely scattered over the islands. 



(22) Succinea riibella Pease. 

Succinea rubella Pease, P. Zool. Soc. London, 1871, p. 460. 

The coloration of this species is very variable, and some specimens are found of a 
straw-colour ; it appears to be close to S. canella Gould. 

Hab. Lanai (Pease) ; Mountains (Perkins). 

(23) Stcccinea rubida Pease. 

Siucinea rubida Pease, J. Conchyl. xviii. (1870), p. 97. 
Hab. Kauai (Pease) ; North side (Baldwin). 

(24) Succinea thaanunii Ancey. 

Succinea t haanumi ^nc&y, P. Malac. Soc. London, in. (1899), p. 273, pi. xii. fig. 3. 
Hab. Hawaii, Olaa (Ancey). 

(25) Stuciiiea venusta Gould. 

Succinea venusta Gould, P. Boston Soc. 11. (1847), p. 186; U.S. Explor. Exped. 

Moll. pi. II. fig. 25. 
Hab. Hawaii, Mauna Kea (Baldwin) ; Kona at 3000 feet, and Olaa Puna 
(Perkins). 

The specimens from Kona are whitish and slightly more shouldered. 



390 FAUNA HAWAIIENSIS 

(26) Succinea vesicalis Gould. 

Stucinea vesicalis Gould, P. Boston Soc. 11. (1847), p. 183; U.S. Explor. Exped. 

Moll. pi. II. fig. 1 7. 
The figure given by Reeve (Conch. Icon. Succinea, fig. 85) does not represent this 
species. 

Hab. Hawaii, Mauna Kea (Gould) ; Kau, and Mauna Loa at 2000 ft. (Perkins). 

(27) Sticcinea waianaensis Ancey. 

Succinea waianaensis Ancey, P. Malac. Soc. London, iii. (1899), p. 273, pi. xii. 

fig. 12. 
Hab. Oahu, Waianae Mts. (Ancey). 



Species insufficiently known or erroneously recorded. 

Succinea aperta Lea. Unknown to me ; it has been doubtfully referred to 

S. rotundata. 
Succinea approximata Shuttleworth, Reeve, Conch. Icon. Succinea, fig. 27. 

Apparently really refers to the West Indian 5. approximans . 
Succinea pudorina Gould, Reeve, Conch. Icon. Succinea, figs. 43, 75. 
Two apparently distinct species are figured by Reeve under this name ; he records 
it, I think erroneously, from the 'Sandwich Islands.' 

The following appear to be only manuscript names. 

^. apicalis Ancey, Makawao, Maui. 
S. tenerriina Ancey, Hilo, Hawaii. 

Fam. LIMNAEIDAE. 
Limnaea Lamarck. 

Limnaea Lamarck, Mem. Soc. Hist. Nat. Paris, 1799, p. 75 (X.y^^ Helix stagnalis L.). 

The name Limnaea appears to have been used as early as 1791 by Poli for the 
animals of Unio, Anodonta and Chama ; but his work is so peculiar, that I feel doubts 
if this can be regarded as preoccupying the name for a genus. 



MOLLUSC A 391 

The question whether these forms — when sinistral — belong to Pkysa or Limnaea, 
has for long proved to be a difficult one. Gould, when describing his Pkysa reticulata, 
remarked " its form is so much that of a reversed Litnnaea that I am almost tempted to 
consider it one." Pease, in his review of the Hawaiian species, stated that he was 
"confident," having e.xamined the animals of several hundred specimens, that there was 
no species of Pkysa in the Islands. He also remarked that sinistral and dextral 
specimens of the same species may be found together. Prof. E. von Martens, in 1866, 
expressed the view, here adopted, that these Hawaiian forms were really sinistral 
species of Limnaea. 

As shewing how little the literature has been studied, I may remark that as late 
as 1889 Mr Cooke (P. Zool. Soc. London, 1889, p. 142), when pointing out the 
affinities with Limnaea of the so-called Pkysae of Australia, notes that " Tryon 
mentions, but I have failed to trace on what authority, that sinistral Limnaeas occur 
in the Sandwich Islands." 

Mighels described (P. Boston Soc. 11. p. 21) two species, /;W//f/a and umbilicata, 
as Pkysae : according to Pease the types were destroyed by fire and, as the species 
are unidentifiable from the diagnoses only, I have omitted them. 



(i) Limnaea aiilacospira Ancey. 

Limnaea aiilacospira Ancey, Naturaliste, (2) xi. (Dec. 1889), p. 290; Sykes, 
P. Malac. Soc. London, in. pi. xiii. fig. 19. 

Hab. Maui (Ancey); Haleakala at 5000 feet, and lao Valley (Perkins). 



(2) Limnaea binom,inis, nom. nov. 

Pkysa sandwichensis Clessin, Conchylien-Cabinet, Pkysa, p. 342, pi. xlviii. fig. 7 

(1886). 
nee Limnaea sandwichensis Philippi. 

Hab. Hawaiian Islands (Clessin). — Oahu, Mts. near Honolulu (Perkins). 



(3) Limnaea compact a Pease. 

Limnaea compacta Pease, Amer. J. Conch, vi. (1870), p. 6, pi. in. fig. 4. 

Limnaea ambigiia, Pease, t. c. p. 6, pi. in. fig. 5. 

Pkysa flavida Clessin, Conchylien-Cabinet, Pkysa, p. 364, pi. Li. fig. 9 (1886). 

Hab. Oahu (Pease). — (as L. ambigna) Kauai, Kapaa (Baldwin). — (as L. compacta) 
all the Islands (Baldwin). 

F. H. II. 51 



392 FAUNA HAWAIIENSIS 



(4) Limnaea hartinamii Clessin. 

Physa hartmanni Clessin, Conchylien-Cabinet, Physa, p. 371, pi. liv. fig. 9 (1886). 

Hab. Hawaii (Clessin). 

Some catalogues give a Limnaea hartmanni of Studer and of Charpentier, but I 
cannot trace a described species of that name ; there appears to be a variety of 
L. ovatus Drap bearing the name. 



{5) Limnaea moreletiana Clessin. 

Physa moreletiana Clessin, Conchylien-Cabinet, Physa, p. 341, pi. xlviii. fig. 3 
(1886). 

Unknown to me, but from the figure I think it may be a form of L. tiirgidula 
Pease. It appears not to be the Limnaea -moreletiana Gassies, of Adams (Gen. Rec. 
Moll. II. p. 253). 

Hab. Hawaiian Islands (Clessin). 



(6) Limnaea naticoides Clessin. 

Physa naticoides Clessin, Conchylien-Cabinet, Physa, p. 341, pi. xlviii. fig. 5 

(1886). 
Hab. Hawaiian Islands (Clessin). 



(7) Limnaea oahuensis Souleyet. 

Limnaea oahuensis Souleyet, Voy. Bonite, Zool. 11. (1S52), p. 527, pi. xxix. 

figs. 38 — 41 [with animal] ; Reeve, Conch. Icon. Limnaea, sp. 90. 
Limnaea affinis Souleyet, Voy. Bonite, Zool. 11. p. 528, pi. xxix. figs. 42-^44. 
Limnaeus sandwichensis Philippi, Arch. Naturg. 11. (1845), p. 63 ; Kuster, 

Conchylien-Cabinet, Limnaea, p. 26, pi. iv. figs. 25, 26. 
Limnaea voliitata, Gould, P. Boston Soc. 11. (1847), p. 211; U. S. Explor. Exped. 

Moll. pi. IX. fig. 142. 

I defer to Pease's experience and unite Souleyet's two species ; though, from the 
figures, I should have regarded them as distinct. It is not the Litnnaea affinis 
of Beck. 

Hab. Oahu (Souleyet, Pease, &c.). — Oahu and Maui (Baldwin). 



MOLL use A 393 

(8) Limnaea peasei Clessin. 

Physa peasei Clessin, Conchylien-Cabinet, Physa, p. 339, pi. XLVii. fig. 8 (1886). 
Judging from specimens received by the British Museum from the Morelet 
collection, the figure is by no means good. 
Hab. Hawaiian Islands (Clessin). 

(9) Limnaea reticulata Gould. 

Physa reticulata Gould, P. Boston Soc. 11. (1847), P- 214; U. S. Explor. Exped. 

Moll. pi. IX. fig. 140; Sowerby, Conch. Icon. Physa, fig. 56; Clessin, 

Conch. -Cab. Physa, p. 330, pi. XLVi. fig. 4. 
Limnaea reticulata Gould, Pease, Amer. J. Conch, vi. p. 5. 
Neither Sowerby nor Clessin appears to have been aware of Gould's published 
description of this species. 
Hab. Kauai (Pease). 

(10) Liinnaca rubella Lea. 

Lymnaeus rtcbellus Lea, Tr. Amer. Phil. Soc. ix. (1843), p. 12. 
Limnaea rubella Lea, Pease, Amer. J. Conch, vi. p. 5, pi. iii. figs, i — 3. 
Pease was of opinion that this might prove to be a variety of L. oahuensis Soul. 
Hab. Oahu (Lea). — Kauai (Pease) ; Mts. between Lihue and the sea, also 
Wailua river (Perkins). 

(11) Limnaea turgidula Pease. 

Limnaea turgidula Pease, Amer. J. Conch, vi. (July, 1870), p. 5, pi. iii. fig. 3. 
Hab. Oahu (Pease). 

Erinna a. Adams. 
Erinna newcombi A. Adams. 

Erinna newcombi A. Adams, P. Zool. Soc. London, 1855, p. 120; H. and 

A. Adams, Gen. Rec. Moll. 11. p. 644, pi. cxxxviii. fig. 9 ; Bland and Binney, 

Ann. Lye. New York, x. p. 349 [jaw and radula] ; Binney, P. Ac. Philad. 

1874, p. 54, pi. v. figs. 7 — 10 [jaw and radula]. 

Hab. Kauai, Hanalei River (Baldwin, &c.). H. and A. Adams give as locality 

" Henata River, Kami." 

See also a note on the genus by Dr Jousseaume, Rev. Mag. Zool. (3) 11. (1S74), 

P- 25. 

SI— 2 



394 FAUNA HAWAIIENSIS 



Ancylus, Geoffroy. 

Ancylus Geoffroy, Traite sommaire des coquilles, aux environs de Paris, 

1767, p. 122 [type apparently A. lacusiris]. 

Ancylus sharpi sp. nov. 

Testa pygmaea, convexiuscula, hyalino-flavida ; apertura elongato-elliptica, apice 
obtusulo. Long. 2; lat. ri ; alt. "8 mill. 

Plate XII. figs. 14, 14 a. 

An insignificant little form with no striking characters ; there being no other 
species recorded from the Islands, I venture to give these shells a name ; they are 
probably not adult. 

Hab. Oahu, on pali, head of Nuuanu Valley (Coll. Dr B. Sharp, commisit 
H. A. Pilsbry). 

Fam. MELANIIDAE. 

Melania Lamarck. 

The genus appears to have been first put forward by Lamarck in 1799 (Mem. 
Soc. Hist. Nat. Paris, p. 75) and to have been also characterised by him in 1801 (Syst. 
An. sans Vert. p. 91). In both cases the species named by him was Melania amarula 
Lam., which is therefore the type. 

(i) Melania baldwini Ancey. 



Melania baldiuini Knay, P. Malac. Soc. London, in. (July, 1899), p. 273, pi. xii. 
fig. 6. 

Hab. Maui, Lahaina (Ancey). 

(2) Melania indefinita Lea. 

Melania indefinita Lea, P. Zool. Soc. London, 1850, p. 187; Reeve, Conch. 

Icon. Melania, fig. 56 ; Brot, Conch. -Cab. Melania, pi. xxiii. fig. 7. 
Melania newcombii Lea, Pease, Amer. J. Conch, vi. p. 6 [nee Lea, fide Brot]. 

Hab. Oahu (Pease). 

The Philippine specimens in coll. Cuming seem identical with some from Oahu, 
named M, newcombii by Pease. 



MOLL use A 395 

(3) Alelania katiaiensis Pease. 

Melania kaiiaiensis Pease, Amer. J. Conch, vi. (July 1870), p. 7, pi. m. fig. 6. 

Hab. Kauai (Pease). — Molokai, Pelekunu (Perkins). 

Probably the species of Melania are scattered over the various islands and not 
confined to any single locality ; M. mauiensis, for example, has been found on 
Maui, Molokai, Kauai, and Oahu. 

(4) Melania mauiensis Lea. 

Melania mauiensis Lea, P. Ac. Philad. viii. (1857), p. 145; Brot, Conch. -Cab. 
Melania, p. 322, pi. xxxiii. figs. 7, 8, 8 a. 

Hab. Maui (Lea).— Maui, Oahu, Kauai (Pease).— Maui, Molokai (Brot).— 
Molokai, in taro patches, Pelekunu (Perkins). 

Large specimens were found on Molokai by Mr Ferkins, exact spot not recorded, 
and a small race, kindly identified for me by the late Dr Brot, on Pelekunu. Melania 
tahitensis Pease MS. is stated by Brot to be a synonym. Schepman (Notes Leyden 
Mus. XIV. p. 158) has recorded the present species from the Island of Soemba. 

(5) Melania newconibii Lea. 

Melania newconibii Lea, P. Ac. Philad. viii. (1857), p. 145 ; Brot, Conch. -Cab. 

Melania, p. 213, pi. xxiv. figs. 2, 2 a. 
Melania contigua Pease, Amer. J. Conch, vi. (July 1870), p. 7. 

I follow Brot in uniting M. contigua Pease ; he also places M. oahuensis Pease 
MS. and M. paidla Dunker MS. in the synonymy. 

Hab. Oahu (Lea) ; In stream in mountain gulch near Honolulu (Perkins). — 
Kauai (Pease). 

(6) Melania verreauxiana Lea. 

Melania verrauiana (sic) Lea, P. Ac. Philad. viii. (1857), P- H4- 
Melania verreauxiana Lea, J. Ac. Philad. n. s. vi. pi. xxii. fig. 27; Brot, Conch.- 
Cab. Melattia, p. 32, pi. iv. fig. 2. 

Unknown to me and may not really be Hawaiian. Dr Brot considered it might 
be a form of M. largillierti Phil. 

Hab. Hawaiian Islands (Lea). 



596 FAUNA HAWAIIENSIS 

Fam. PALUDESTRINIDAE. 

Paludestrina D'Orbigny. 
Paludestrina porrecta Mighels. 

Paludina porreda Mighels, P. Boston Soc. ii. (1845), p. 22. 
Hab. Oahu (Mighels). 



Fam. HELICINIDAE. 
Helicina Lamarck. 

In 1799 (Mem. Soc. Hist. Nat. Paris, p. 77) the genus was described but no type 
or species named ; in 1801 (Sypt. \n. sans Vert. p. 94) the only species named was 
Heliciiia nerite//a Lam., which may be taken as the type. Lamarck refers for a figure 
to Lister (Hist. Conch, fig. 59), and this illustration appears to represent a Helicina, 
though it is hard to be certain whether it be H. neritella or not. 



(i) Helicina laciniosa Mighels. 

Helicina laciniosa Mighels, P. Boston Soc. 11. (1845), p. 19 ; Gould, U. S. Explor. 
Exped. Moll. pi. VII. fig. 108. 

A very variable shell in size and coloration ; it appears to be always more 
compact and elevated than H. sandwichiensis. 

Hab. Oahu (Mighels). — Kauai (Baldwin). — Lanai, behind Koele; also Kalamaula, 
Molokai ; Kaala, Oahu ; and between Lihue and the sea, Kauai (Perkins). 



(2) Helicina magdalenae Ancey. 

Helicina magdalenae Ancey, Bull. Soc. Malac. France, vii. (1890), p. 342. 
Helicina constricta Pfeiffer, P. Zool. Soc. London, 1848, p. 120; Conch.-Cab. 
Helicina, p. 22, pi. vii. fig. 1'] — 9 [both relate to his variety only]. 

Pfeiffer's typical form came from ' Otaheite ' and appears to belong to a different 
species to his variety, which seems to be identical with this. Possibly forms may be 
found linking H. magdalenae to H. uberta. 

Hab. Oahu, Tantalus (Ancey). 



MOLL use A 397 

(3) Helicina rotcUoidea Mighels. 

Helicina rotelloidea Mighels, P. Boston Soc. 11. (1845), p. 19 ; Pfeiffer, Conch. - 

Cab. Helicina, p. 23, pi. iii. fig. 40 — 3. 
Helicina bronniana Philippi, Zeitsch. Malak. iv. (1847), p. 124. 
Hab. Oahu (Mighels, &c.). 

(4) Helicina sandivichiensis Souleyet. 

Helicina sandwichiensis Souleyet, Voy. Bonite, Zool. 11. (1852), p. 529, pi. xxx. 

figs. 1—5. 
nee ? H. sandivicJiiensis Sowerby, Thes. Conch, iii. pi. CCLXX. figs. 173 — 4. 

A variety "^" has been recorded by Pfeiffer as from the Loyalty Islands; 
probably this is an error. See Crosse, J. Conchyl. xlii. p. 405. 

Hab. Oahu, Waianae Mts. (Baldwin) ; at and below Kaala (Perkins). 

(5) Helicina tiberta Gould. 

Helicina iiberta Gould, P. Boston Soc. 11. (1847), p. 202 ; U. S. Explor. Exped. 
Moll. pi. VII. fig. 1 14. 

Hab. Maui and Oahu (Gould). — Oahu, below Kaala (Perkins). 

Species doubtful or erroneously recorded. 

Helicina a7itoni'?{&\ii&T. Originally recorded without locality; subsequently Pfeiffer 
gave the Hawaiian Islands and the Gamblers. It really appears to come from 
Honduras, and the Hawaiian habitat is probably erroneous, these supposed Hawaiian 
specimens belonging, as undoubtedly the Gambler Island shells do, to H. pazi Crosse 
(J. Conchyl. xiii. p. 221, pi. vi. fig. 8). 

Helicina crassilabris, Philippi. It has been suggested by Pfeiffer that this is 
Hawaiian, but it really comes from Venezuela or the Caribbean Region. 

Helicina fjilgoj-a Gould, originally described from Manua, Samoa Islands; it has 
also been noted, but, I think, erroneously, from the Hawaiian Islands. 

Helicina pisum Philippi. I think "Sandwich Is." must have been a mistake and 
possibly refers to Vate or Sandwich I.: it may be a slip for Savage I., from which 
specimens, inseparable from this, undoubtedly do come. This appears not to be the 
H. pisum Hombr. and Jacq., which equals H. taliitensis Pease. 



398 FAUNA HAWAIIENSIS 

Fam. NERITIDAE. 

Neritina Lamarck. 

I have not seen the first Edition of the ' Philosophie Zoologique' (1809) in which 
this genus is said to occur, but in the second edition (1830) the name appears in French 
only, with no diagnosis or named species (Vol. i. p. 321). However in his ' Hist. An. 
sans Vert.' it is duly given in Latin with named species (Vol. vi. pt. 2, p. 182). 
The first is N. perversa Gmel., which is the type of Montfort's Velates (1810) 
under the more correct name of V. conoidea, but the others belong to Neritina as 
we understand it to-day. 

(i) Neritina cariosa Gray. 

Nerita cariosa Gray, Wood, Index Test. Suppl. Nerita fig. 9 (1828). 

Neritina sandwichensis Deshayes, An. sans Vert. Ed. 2, viii. (1838), p. 579. 

Neritina convexa Nuttall, Jay, Cat. Shells, Ed. 3, 1839, p. 66 (nom. sol.). 

Neritina nuttalli Recluz, Rev. Zool. 1841, p. 276 ; Souleyet, Voy. Bonite, Zool. 11. 
pi. XXXIV. figs. 43 — 46. 

Neritina solidissima Sowerby, Thes. Conch. 11. p. 541, pi. cxvi. fig. 573. 

I have not sufficient material to determine whether the large synonymy given by 
Tryon (Man. Conch, x.) is fully justified. Prof von Martens (Conch. -Cab. Neritina) 
expressed the opinion (p. 276) that Neritina cariosa Gray does not really belong here, 
but is a form of N. mauritii: this has been dealt with by Mr Smith (P. Zool. Soc. 
London, 1884, p. 275). 

Hab. Hawaiian Islands (various authors). — Maui and Oahu (Baldwin). — Hawaii, 
Hilo (Smith). 

(2) Neritina granosa Sowerby. 

Neritina granosa So-werhy, Tank. Cat. App. p. xi. (1825); Conch. 111. Neritina 

fig. 6. 
Neritina papulosa Jay, Cat. Shells, Ed. 2, 1839, pi. iv. fig. 11. 
Neripteron gigas Lesson, Rev. Zool. 1842, p. 187. 
Hab. All the Islands (Baldwin). — Molokai, Pelekunu (Perkins). 

(3) Neritina liigubris Philippi. 

Neritina lugubris Philippi, Abbild. Conchylien, i. pt. 2, p. 20, pi. i. fig. 9 (1845). 

This has been placed as a synonym of N. cariosa, but from the description and 
figure it seems to be distinct. 

Hab. Hawaiian Islands (Philippi). 



MOLL use A 399 

(4) Neritina neglect a Pease. 

Neritina negleda Pease, P. Zool. Soc. London, i860, p. 435. 
Hab. Hawaiian Islands (Pease). 

(5) Neritina vespertina Nuttall. 

Neritina vespertina Nuttall, Jay, Cat. Shells, Ed. 3, 1839, p. 66 (nom. sol.); 

Reeve, Conch. Icon. Neritina, sp. 61. 
? Neritina sandwichensis Desh., Reeve, Conch. Icon. sp. 82 [nee Deshayes]. 
Hab. All the Islands (Baldwin). 



In conclusion I may call attention to three species, attributed to the Islands, which 
do not really belong to their fauna. 

Partula terrestris Pease. Apparently a manuscript name ; it has appeared in 
Paetel's 'Catalog' and in the Mon. Helic. Viv. (Vol. viii. p. 209) with the 
habitat of ' I. Sandwich.' According to Dr Hartman, it is a synonym of 
P. approximans Pease, from Raiatea. 

Spiraxis sandwicensis was described by Pfeiffer (P. Zool. Soc. London, 1856, 
P- 335) ^s from the Hawaiian Islands. It appears to me to be a form of the 
Btilimus lactifljtus of Pfeiffer, described from Chili, and 1 feel no doubt the 
Hawaiian habitat is erroneous. 

Bulimus kaziaiensis was described by Pfeiffer in the same volume (p. 332). It is 
probably also Chilian and very close to Bulinms albicans Brod. ; but 1 am not 
quite sure of the identity, as the shell is slightly more succineiform. 

Finally, it may be noted that a specimen of Vivipartis ckinensis Gray, doubtless 
imported for food, was collected by Mr Perkins at "Wailuku," Maui. 



F. H. II. 52 



§ 3. Bibliographic List (arranged alphabetically). 

Adams, A. Descriptions of two new genera and several new species of MoUusca, from the collection 

of Hugh Cuming, Esq. P. Zool. Soc. London, 1855 [August to December], pp. 119— 124. 
Adams, C. B. Descriptions of new species of Pariula and Achatinella. Ann. Lye. New York, v. 

(1852), pp. 41—44- 

. Contributions to Conchology. Vol. i. No. 8, 1850, pp. 125 — 128. 

Adams, H. & A. The Genera of recent MoUusca. London, 3 vols. 8vo, 1853—1858. 

Albers, J. C. Die Heliceen. Berlin, 8vo, 1850. 

Ancev, C. F. fetude sur la faune malacologique des iles Sandwich. Bull. Soc. Malac. France, vi. 

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. Mollusques nouveaux de I'Archipel d'Hawai, de Madagascar, et de I'Afrique equatoriale. 

Op. cit. VII. (1890), pp. 339—347- 
. Diagnoses de Mollusques nouveaux. Naturaliste, ser. 2, an. iii (1889), p. 266 S^Leptachatina 

columna, n. sp.]. 
. Descriptions de Mollusques nouveaux. T. c. pp. 290, 291 [Lmmaea aulacospira, n. sp.]. 

. Etudes sur la faune malacologique des iles Sandwich. Mem. Soc. Zool. France, v. (1892), 

pp. 708 — 722. 
. Etudes sur la faune malacologique des iles Sandwich. Op. cit. vi. (1893), pp. 321 — 330. 

. Descriptions de deux nouvelles especes de Mollusques. Naturahste, ser. 2, an. xi (1897), 

p. 178 \Ainast)-a durandi, n. sp.]. 
. Description d'un mollusque nouveau. T. c. p. 222 S^Leptachatina approximaus, n. sp.]. 

. Some notes on the non-marine molluscan fauna of the Hawaiian Islands, with diagnoses 

of new species. P. Malac. Soc. London, iii. (July, 1899), PP- ^68 — 274, pis. xii, xiii pars. 

Baldwin, D. D. Catalogue [of] land and fresh-water shells of the Hawaiian Islands. Honolulu, 

Bvo, 1893, 25 pp. 
. Descriptions of new species of Achatinellidae from the Hawaiian Islands. P. Ac. Philad. 

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. Descriptions of two new species of Achatinellidae from the Hawaiian Islands. Nautilus, 

X. (July, 1896), pp. 31, 32. 
Barnacle, H. G. Musical sounds caused by Achatinellae. J. Conch, iv. (1883), p. 118. 
Beck, H. Index Molluscorum praesentis aevi musei Christiani Frederici. Havniae, 1837, 

fasc. I, 2. 

Benson, W. H. General features of Chusan, with remarks on the Flora and Fauna of that Island. 
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Bergh, R. Anatomische Untersuchung des Triboniophortis schuttei, Kefstr., sowie von Philomycus 
carolimnsis (Bosc) und australis (Bergh). Verh. Ges. Wien, xx. (1870), pp. 843 — 868, 

pis. XI — XIII. 



MOLL use A 401 

BiNNEY, W. G. On some of the species of naked Pneumonobranchous Mollusca of the United 
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. On the anatomy and lingual dentition oi ArioHtnax and other Pulmonata. P. Ac. Philad. 1874, 

PP- 33—62, pis. II— XI. 

. On the genitalia, jaw, and lingual dentition of certain species of Pulmonata [with a note 

on the classification of the Achatinellae, by Thomas Bland]. Ann. Lye. New York, xi. (1875), 

pp. 166 — 196, pis. XII XVIII. 

. On the lingual dentition, jaw, and genitalia of Carelia, Otichidella, and other Pulmonata. 

P. Ac. Philad. 1876, pp. 13 — 92, pi. vi. 

. Notes on the jaw and lingual dentition of Pulmonata Mollusks. Ann. N. York Ac. in. 1884, 

pp. 79 136, pis. II — XVI. 

Bland, T. & Binnev, W. G. On the lingual dentition and anatomy of Achatine/la and other 
Pulmonata. Ann. Lye. New York, x. (Nov. 1873), pp. 331 — 350, pis. xv, xvi. 

Boettger, O. Die /"///a-Arten Oceaniens. In von Martens' Conch. Mittheil. i. (1880), pp. 45 — 72, 

pis. X — XII. 

Chamisso, a. de. Species novas Conchyliorum terrestrium ex insulis Sandwich dictis attulit. Acta 
Ac. German, xiv. (1829), pp. 639, 640, pi. xxxvi. 

Clessin, S. Nomenclator Heliceorum Viventium. Cassel, 8vo, 1881. 

Collinge, W. E. On a collection of Slugs from the Sandwich Islands. P. Malac. Soc. London, 11. 
(April, 1896), pp. 46—51, figs. 

• . On a further collection of Slugs from the Hawaiian (or Sandwich) Islands. Tom. cit. 

(Nov. 1897), pp. 293—297, figs. 

. On the anatomy and systematic position of some recent additions to the British Museum 

collection of Slugs. J. Malac. vii. (1900), pp. 77 — 85, pis. iv, v. 

Cooke, A. H. On the generic position of the so-called Physae of Australia. P. Zool. Soc. London, 
1889, pp. 136—143, figs. 

Crosse, H. Note comple'mentaire sur quelques especes de mollusques habitant File Kauai (iles 
Hawaii). J. Conchyl. xxiv. (1876), pp. 95 — 99, pis. i pars, iii pars, iv pars. 

Deshayes, G. p. [See Ferussac & Deshayes.] 

Dixon, G. A voyage round the world. London, 1789 [cf. p. 354]. 

Draparnaud, J. Tableau des mollusques terrestres et fluviatiles de la France. Paris, 8vo, 1801. 

— . Histoire naturelle des mollusques terrestres et fluviatiles de la France. Paris and Montpellier, 

1805. 

Ferussac, D. de. Tableaux Systdmatiques des animaux mollusques suivis d'un Prodrome 

gene'ral Paris, folio, 1822. 

. [Review of Swainson's description oi AchatinellaJ] Bull. Sci. Nat. xvi. (1829), pp. 138 — 141. 

& Deshayes, G. P. Histoire naturelle des mollusques terrestres et fluviatiles Paris, 

folio, 1820 — 1851. 
FiTZiNGER, L. J. Verzeichniss der im Erzherzogthum Oesterreich vorkommenden Weichthiere, als 

Prodrom einer Fauna derselben. Beitr. Landesk. Oesterr. in. (1833), pp. 88 — 122. 
Gaimard, — . [See Quoy & Gaimard.] 
Garrett, A. On new species of marine shells of the Sandwich Islands. P. Calif. Ac. i. (1854 — 7) 

(second edition, 1873), pp. 114, 115 \_Succinea newcombiatta, n. sp.]. 
Geoffroy, E. L. Traitd sommaire des coquilles, tant fluviatiles que terrestres, qui se trouvent aux 

environs de Paris. Paris, 1767, i2mo. 

52 — 2 



402 FAUNA HAWAIIENSIS 

Gould, A. A. [Descriptions of shells from the Sandwich Islands.] P. Boston Soc. i. (1843 — 4), 

pp. 139, 174. 

. Descriptions of land shells from the Sandwich Islands. Op. cit. 11. (1845), PP- 26 — 28. 

. Descriptions of new shells collected by the United States Exploring Expedition belonging 

to the genus Helix. Tom. cit. (1846), pp. 171 — 173, 177. 
. Descriptions of new shells collected by the United States Exploring Expedition. Tom. cit. 

(1847), PP- 181, 182—7, 191, 197, 200 — 2, 211, 214. 
. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842, under the 

command of Charles Wilkes, U.S.N. MoUusca and Shells. Philad. 1852, 410, with folio 

atlas (1856). 

. Descriptions of new genera and species of shells. P. Boston Soc. viii. (Feb. 1862), pp. 280^284. 

Gray, J. E. Catalogue of the Pulmonata in the British Museum. Part I. London, 8vo, 1855. 
Green, J. New species of Achatina, with remarks on the Ti, or the Dracena terminalis, of the 

Sandwich Islands. Contrib. Maclurian Lye. i. no. 2 (July, 1827), pp. 47 — 50, pi. iv. 

. Remarks on the Achatina sieivartii. T. c. no. 3 (Jan. 1829), pp. 66, 67. 

Gulick, J. T. Descriptions of new species of Achatinella from the Hawaiian Islands. Ann. Lye. 

New York, vi. pp. 173 — 255, pis. vi — viii [pp. 173 — 230 bear date Dec. 1856, while pp. 231 — 255 

are dated Feb. 1858: diagnoses reprinted Malak. Blatt. v. (1858), pp. 198 — 224]. 
. On the variation of species as related to their geographical distribution, illustrated by the 

Achatinellinae. Nature, vi. (July 18, 1872), pp. 222 — 224. 

. On the classification of the Achatinellinae. P. Zool. Soc. London, 1873, pp. 89 — 91. 

. On diversity of evolution under one set of external conditions. J. Linn. Soc. Zool. xi. 

pp. 496—505. 
. Lessons in the theory of divergent evolution, drawn from the distribution of the land shells of 

the Sandwich Islands. P. Boston Soc. xxiv. (1890), pp. 166, 167. 
& Smith, E. A. Description of new species of Achatinellinae. P. Zool. Soc. ^London, 1873, 

PP- 73—89, pis. IX, X. 
GwATKiN, H. M. & SuTER, H. with prefatory note by Pilsbrv, H. A. Observations on the dentition 

of Achatinellidae. P. Ac. Philad. 1895, pp. 237 — 240, pi. xi pars. 
Hartman, W. D. a bibliographic and synonymic catalogue of the genus Auricidella, Pfeiffer. 

P. Ac. Philad. 1888, pp. 14, 15. 
. A bibliographic and synonymic catalogue of the genus Achatinella. Tom. cit. pp. 16 — 56, 

pi. I. 
. New species of shells from the New Hebrides and Sandwich Islands. Tom. cit. pp. 250 — 252, 

pi. XII. [? 1889']. 
Hartmann, J. D. W. Erd- und Siisswasser-Gasteropoden. St Gall, 1840. 
Hasselt, F. C. van. Extrait d'une lettre de F. C. van Hasselt sur les mollusques de I'ile de 

Java, adressee au Prof, van Swinderen a Groningue. Bull. Sci. Nat. Geol. iii. (1824), pp. 81 — 87. 
Heynemann, F. D. Die Zungen von Fartula und Achatitiella. Malak. Blatt. xiv. (1867), 

pp. 146—150, pi. I. 
. Die Kiefer von Philomycics carolinensis, Bosc, und atistralis, Bergh. Nachrbl. Deutsch. 

malak. Ges. iii. (1871), pp. i, 2, pi. i pars. 
Hyatt, A. Evolution and migration of Hawaiian land-shells. P. Amer. Ass. xlvii. (1898), 

PP- 357. 358- 

Jay, J. C. A catalogue of the shells in the collection of John C. Jay. Ed. 3, 1839, 4to. 

Jousseaume, Dr. Des genres Eri?ma et Laiitzia. Rev. Mag. Zool. (3) 11. (1874), p. 25. 

1 The sheet is certified by a note in the volume, dated Feb. 6, 1S88 {sic), signed E. J. Nolan, to have been 
presented on Oct. 23, 1888. 



MOLL use A 403 

Keferstein, W. Ueber die Anatomic der Gattungen Incillaria, Benson, und Mcghimatium, Hasselt, 

im Vergleich mit der von Fhi/ofnyais, Rafinesque. Malak. Blatt. xiii. (1866), pp. 64 — 70, pi. i. 
KoBELT, W. Conchologische Miscellen. J.B. Deutsch. malak. Gas. 11. (1875), pp. 222 — 228, pi. vii. 

. Die geographische Verbreitung der Mollusken. iii. Die Inselfaunen. Op. cit. vi. (1879), 

pp. 195—224. 

Lamarck, J. B. Prodrome d'une nouvelle classification des coquilles. Mem. Soc. Hist. Nat. Paris, 
1799, pp. 63—91. 

. Systeme des animaux sans vertebres. Paris, 1801. 

. Histoire naturelle des animaux sans vertebres. Paris, 7 vols., 1815 — 1822. 

Lea, L On freshwater and land shells. Tr. Amer. Phil. Soc. ix. (1843), pp. i — 31 [^Lymnaea 
rubella, n. sp., p. 12]. 

. Descriptions of fifteen new species of exotic Melaniana. P. Ac. Philad. viii. (1857), 

pp. 144, 145. 

. Observations on the genus Unio, &c. Vol. xi. 1866. 

. New Unionidae, Melaniidae, &c., chiefly of the United States. J. Ac. Philad. n. s. vi. (1867), 

pp. 113 — 187, pis. XXII — XXIV. 

& H. C. Description of a new genus of the family Melaniana and of many new species of the 

genus Melania, chiefly collected by Hugh Cuming, Esq., during his voyage in the east, and now 
described. P. Zool. Soc. London, 1850, pp. 179 — 197 [J/, indefinita, n. sp., p. 187]. 

Lesson, R. P. Description d'une espece nouvelle de Neriptere. Rev. Zool. 1842, pp. 187, 188 

\Neripteron gigas\. 
Lyons, A. B. A few Hawaiian land-shells. Hawaiian Annual, 1892, pp. 103 — 109, pis. i, 11. 
Martens, E. von. Die Heliceen. Ed. 2. Leipzig, 8vo, i860. 

. Conchological Gleanings, iii. The Sandwichian species of Limnaeus. Ann. Nat. Hist. (3) 

XVII. (1866), pp. 207 — 210. 

. Preuss. Exped. nach Ost-Asien. Zool. Theil, Band 11. Berlin, 1867. 

Martini & Chemnitz. Conchylien-Cabinet. Various monographs in Editions i and 2. 

MiGHELS, J. W. Descriptions of shells from the Sandwich Islands and other localities. P. Boston 

Soc. II. (1845), PP- 18—25. 
Moquin-Tandon, a. Histoire naturelle des mollusques de France. Paris, 1855, 2 vols, and atlas. 
MciRCH, O. A. L. Quelques mots sur un arrangement des mollusques pulmone's terrestres (Geophiles, 

Per.) base sur le systeme naturel (suite). J. Conchyl. xiu. (1865), pp. 376 — 396. 
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deshayesii, n. sp., p. 27]. 

. Des genres Erinna, Lithotis, el Lanizia. J. Conchyl. xxiii. (1875), pp. 280, 281. 

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docteur Ed. Graeffe. J. Conchyl. xix. (1871), pp. 5 — 34 \Tornaiellina bacillaris, n. sp., p. 16]. 
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Testaceorum, non Marinorum, succincta historia. Havniae et Lipsiae, 1773 — 1774, 2 vols. 

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New York, vi. (May, 1853), pp. 18 — 30. 
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Sept. 1853), pp. 218 — 220. 
. Descriptions of seventy-nine new species of Achatinella, a genus of pulmoniferous mollusks, in 

the collection of Hugh Cuming, Esq. P. Zool. Soc. London, 1853 [Nov. 1854], pp. 128 — 157, 

pis. xxii — xxiv. 



404 FAUNA HAWAIIENSIS 

Newcomb, W. Abstract of descriptions of some animals of Achatinella, and other remarks. P. Zool. 
Soc. London, 1854 [May, 1855], pp. 310, 311. 

. Descriptions of new species of Achatinella. Ann. Lye. New York, vi. (Oct. 1855), pp. 142 — 147. 

. Synopsis of the genus Achatinella. Tom. cit. (Sept. 1858), pp. 303 — 336. 

. Descriptions of new species of the genera Achatinella and Pupa. Op. cit. vii. (April, i860), 

PP- 145—147- 

. Description of new shells. P. Calif. Ac. 11. (1861), pp. 91 — 94. 

. Description of new species of land-shells. Op. cit. in. (1865), pp. 179 — 182. 

. Descriptions of Achatinellae. Amer. J. Conch. 11. (July, 1866), pp. 209 — 217, pi. xiii. 

Pease, W. H. Descriptions of forty-seven new species of shells from the Sandwich Islands, in the 
collection of Hugh Cuming, Esq. P. Zool. Soc. London, i860, pp. 431 — 438 [Neritina negkcta, 
n- sp., p. 43S]- 

. Descriptions of two new species of Helicter (= Achatitiella, Swainson), from the Sandwich 

Islands, with a history of the genus. Op. cit. 1862, pp. 3 — 7. 

. Description of new land-shells from the Islands of the Central Pacific. Op. cit. 1864, 

pp. 668 — 676 \Tornatellina oblonga, n. sp., p. 673]. 

. Descriptions of new species of land-shells inhabiting Polynesia. Amer. J. Conch. 11. (Oct. 1866), 

pp. 289—293. 

. Descriptions d'especes nouvelles d'Auricuklla provenant des iles Hawaii. J. Conchyl. xvi. 

(1868), pp. 342—347. 

. Descriptions d'especes nouvelles du genre Helicter, habitant des iles Hawaii. Op. cit. xvii. 

(1869), pp. 167—176. 

. On the classification of the Helicterinae. P. Zool. Soc. London, 1869, pp. 644 — 652. 

. Observations sur les especes de coquilles terrestres qui habitent I'ile Kauai (iles Hawaii), 

accompag'nees de descriptions d'especes nouvelles. J. Conchyl. xviii. (1870), pp. 87 — 97. 

. Remarques sur certaines especes de coquilles terrestres habitant la Polynesie, et descriptions 

d'especes nouvelles. T. c. pp. 393 — 403. 

. Synonymie de quelques genres et especes de coquilles terrestres habitant la Polynesie. Op. cit. 

XIX. (1871), pp. 92 — 97. 

. Remarks on the species of Melania and Limtiaea inhabiting the Hawaiian Islands, with 

descriptions of new species. Amer. J. Conch, vi. (1871), pp. 4 — 7, pi. in pars. 

. Catalogue of the land-shells inhabiting Polynesia, with remarks on their synonymy, distribution, 

and variation, with descriptions of new genera and species. P. Zool. Soc. London, 187 1, 
pp. 449—477. 

Pfeffer, G. Anatomische Untersuchung der Achatinella vulpina. J.B. Deutsch. malak. Ges. iv. 

(1877), pp. 330—334, figs. 
Pfeiffer, L. Symbolae ad Historian! Hehceorum. Cassel, 8vo, 1841 — 1846. 

. Monographia Heliceorum Viventium. Leipzig, 8 vols., 1848 — 1877. 

. Novitates Conchologicae. Ser. i. Cassel, 5 vols., 1854 — 1879. 

. Uebersicht der mit innern Lamellen versehenen Ifelix-Arten. Zeitschr. Malak. 11. (1845), 

pp. 81 — 87 [Helix lamellosa, Fer., p. 85]. 

. Ueber neue Landschnecken von Jamaika und den Sandwichinseln. Op. cit. in. (1846), 

pp. 113 — 120. 

. Remarks on the genus Achatinella, Swainson, and description of six new species from 

Mr Cuming's collection. P. Zool. Soc. London, 1845 [Jan. 1846], pp. 89, 90. 



MOLL use A 405 

Pfeiffer, L. Descriptions of thirty new species of Helicea, belonging to the collection of H. Cuming, Esq. 

P. Zool. Soc. London, 1846, pp. 28 — 34. 
. Descriptions of twenty new species of Helicea, in the collection of H. Cuming, Esq. Tom. cit. 

PP- 37—41- 
. Diagnosen neuer Landschnecken. Zeitschr. fiir Malak. iv. (1847), pp. 145 — 151 \Tornatellina 

petitiaiM, n. sp., p. 149]- 
■ . Descriptions of twenty-nine new species of Helicina from the collection of H. Cuming, Esq. 

P. Zool. Soc. London, 1848 [April, 1849], PP- 119 — 125. 
• . Nachtrage zur L. Pfeiffer Monographia Heliceorum : zu Vol. 11. Zeitschr. fiir Malak. vi. (1849), 

PP- 85—95- 

• . Beschreibungen neuer Landschnecken. Zeitschr. Malak. vii. (1850), pp. 65 — 80 [Helix 

discuhis, n. sp., p. 68; cf. p. 153]. 

. Nothwendige Vertauschung einiger Nahmen. Zeitschr. Malak. ix. (1852), pp. 62 — 64. 

. Descriptions of fifty-four new species of Helicea from the collection of Hugh Cuming, Esq. 

P. Zool. Soc. London, 185 1 [July to Dec. 1853], pp. 252 — 263. 
. Descriptions of sixty-six new land-shells from the collection of H. Cuming, Esq. Op. cit. 1852 

[March to May, 1854], pp. 56 — 70. 
. Descriptions of nineteen new species of Helicea, from the collection of Mr Cuming. Op. cit. 

1853 [1854], pp. 124—128. 

. Skizze einer Monographie der Gattung Achatinella. Malak. Blatt. i. (1.854), pp. 112 — 145. 

. Versuch einer Anordnung der Heliceen nach natiirlichen Gruppen. Op. cit. 11. (1855), 

pp. 112— 185. 
. Descriptions of forty-two new species of Helix, from the collection of H. Cuming, Esq. 

P. Zool. Soc. London, 1854, pp. 49 — 57. 
. Descriptions of fifty-seven new species of Hehcea from Mr Cuming's collection. Tom. cit. 

1854 [May, 1855], pp. 286—298. 

. Descriptions of twenty-seven new species oi Achatinella from the collection of Hugh Cuming, Esq., 

collected by Dr Newcomb and by Mons. D. Frick, late Consul-General of PVance at the Sandwich 
Islands. Op. cit. 1855 [March], pp. i — 7, pi. xxx. 

. Descriptions of forty-seven new species of Helicea from the collection of H. Cuming, Esq. 

Tom. cit. [July], pp. 94 — loi. 

. Descriptions of nine new species of Helicea from Mr Cuming's collection. Tom. cit. [August], 

pp. 106 108, pi. XXXII. 

. Descriptions of twenty-three new species of Achatinella, collected by Mons. D. Frick in the 

Sandwich Islands ; from Mr Cuming's collection. Tom. cit. [Feb. 1856], pp. 202 — 206. 
. Descriptions of sixteen new species of Achatinella, from Mr Cuming's collection, collected 

by Dr Newcomb in the Sandwich Islands. Tom. cit. [Feb. 1856], pp. 207 — 210. 
. Descriptions of five new species of Terrestrial Mollusca, chiefly from the collection of 

H. Cuming, Esq. Tom. cit. [Feb. 1856], pp. 210, 211. 
. Weitere Beobachtungen iiber die Gattung Achatinella. Malak. Blatt. 11. (1854 — 1855), 

pp. 1—7, 64—70. 
. Versuch einer Anordnung der Heliceen nach natiirlichen Gruppen. Tom. cit. (1S55 — 1856), 

pp. 112— 185. 
— . Descriptions of twenty-five new species of land-shells, from the collection of H. Cuming, Esq. 

P. Zool. Soc. London, 1856, pp. 32 — 36. 
. Descriptions of fifty-eight new species of Helicea from the collection of H. Cuming, Esq. 

Tom. cit. [March, 185 1], pp. 324—336. 
. Ueber die in Gould's Expedition Shells beschriebenen und abgebildeten Landschnecken. 

Malak. Blatt. iv. (1857), pp. 29 — 37. 



4o6 FAUNA HAWAIIENSIS 

Pfeiffer, L. Neue Landschnecken. Tom. cit. pp. 85 — 89. 

. Diagnosen neuer Heliceen. Tom. cit. pp. 229 — 232. 

. Descriptions of eleven new species of land-shells from the collection of H. Cuming, Esq. 

P. Zool. Soc. London, 1858 [March], pp. 20 — 23, pi. xl. 
. Descriptions of twenty-seven new species of land-shells, from the collection of H. Cuming, Esq. 

Op. cit. 1859, pp. 23 — 29 \Helix hystricella, n. sp., p. 25]. 
. Descriptions of eight new species of Achatinella, from Mr Cuming's collection. Tom. cit. 

pp. 30—32. 
. Descriptions of forty-seven new species of land-shells from the collection of H. Cuming, Esq. 

Op. cit. 1861, pp. 20 — 29 \BuHmus pyrgisais, n. sp., p. 24]. 
Philippi, a. R. Diagnosen einiger neuen Conchylien. Arch. Naturg. 1845, "• PP- 5° — 7i \Limnaea 

volutafa, n. sp., p. 63J. 

. Testaceorum novorum centuria (continuatio). Zeitschr. Malak. iv. (1847), pp. 113 — 127. 

. Abbildungen und Beschreibungen Conchylien. Cassel, 1845 — 1851, 3 vols. 

PiLSBRY, H. A. Relations of the land-molluscan fauna of South America. P. Ac. Philad. 1899, 

p. 226 [reprinted Ann. Nat. Hist. iv. (1899), p. 156]. 

. [See also Gwatkin, Tryon.] 

QuoY, — & Gaimard, — . Voyage autour du monde Uranie et Physicienne Paris, 1824. 

Recluz, C. a. Descriptions de quelques nouvelles especes de Ne'rites vivantes. Rev. Zool. 1841, 

pp. 274 — 276. 
Reeve, L. Conchologia Iconica. Monographs relating to Achatinella, Helix, &c. 
Semper, C. Reisen im Archipel der Philippinen. Band in. Landmollusken. Wiesbaden, 410, 

1870 — 1894. 
Semper, O. Note relative aux genres Balea et Temesa. J. Conchyl. xiv. (1866), pp. 41 — 45. 
Senoner, — . Extrait d'une lettre par M. Senoner. Bull. Soc. Malac. Belgique, vii. pp. cxx, cxxi. 
Smith, E. A. Description of a new species o{ Helix. Ann. Nat. Hist. (4) xx. (1877 Sept.), p. 242. 
. An account of the land and fresh-water moUusca collected during the Voyage of the ' Challenger' 

from December 1872 to May 1876. P. Zool. Soc. London, 1884, pp. 258 — 281, pis. xxi, xxii. 

. [See also Gulick.] 

SoULEYET, — . Voyage autour du monde sur la corvette Bonite. Vol. 11. 1852, Paris. 

. Descriptions de quelques coquilles terrestres appartenant aux genres Cyclostome, Helice, &c. 

Rev. Zool. 1842, pp. loi, 102. 

Sowerby, G. B. a catalogue of the shells of the late Earl of Tankerville London, 8vo, 1825. 

. The Conchological illustrations. London, 8vo, 1841. 

. Thesaurus Conchylioruni. Various Monographs. 

SuTER, H. [See Gwatkin.] 

SwAiNSON, W. The characters of Achatinella, a new group of terrestrial shells, with descriptions 

of six species. Quart. J. Sci. Lit. and Arts, i. (1828), pp. 81 — 86. 

. Zoological Illustrations. Ser. 11. London, 1832 — 1833. 

Sykes, E. R. Preliminary diagnoses of new species of non-marine moUusca from the Hawaiian Islands. 

Parts I, 2. P. Malac. Soc. London, 11. pp. 126 — 132 (Oct. 1896), 298, 299 (Nov. 1897). 
. Contributions towards a list of papers relating to the non-marine mollusca of the Hawaiian 

Islands. Hertford, 8vo, 8 pp., 1896; second edition, 1897. 
. Illustrations of, with notes on, some Hawaiian non-marine mollusca. P. Malac. Soc. London, 

III. (July, 1899), pp. 275, 276, pi. XIII pars, and xiv. 
Tryon, G. W. Manual of Conchology. Series 2. Vols. 11 (1886), ix (1894). Philadelphia. 
Wood, W. Index Testaceologicus. Supplement. London, 1828. 



MOLLUSC A 



407 



4. List of named forms which are placed in this work as 

varieties or synonyms. 



aciit'i Newcomb (Ac/i.) = elorigata Newcomb {Ainastra) 

acuta Swainson {Ach.) = spirizona Fer. {Amastra) 

adamsi Newcomb (Air/i.) = marmorata Gould {Ach.) . 

adiista Reeve {Ach.) = vulpina Fer. {Ach.) 

adusta Gould {Ach.) = bicolor Jay {Carelia) 

affinis Souleyet {Limnaed) = oahiiensis Souleyet {Limnaea) 

alba Nuttall {Ach.) = /orata Fer. {Ach.) 

albescens Gulick {Ach.) = curia Newcomb {Ach.) 

albida Pfeiffer {Amastra) = spirizona Fer., var. {Amastra) 

albofasciatus Smith {Apex) = vittata Reeve {Ach.) 

ambigua Pease {Limnaea) = compacta Pease {Limnaea) 

ampulla Gulick (^it//.) = tappaniana Adams {Ach.) 

analoga Gulick {Ach.) = vulpina F<§r., var. {Ach.) 

aplustre Newcomb {Ach.) = bilineata Reeve {Ach.) 

armatus Mighels {Bulimus) = auricula Fe'r. {Auric.) . 

attenuata Pfeiffer {Bui.) = terebra Newcomb {Ach.) 

bacca Reeve {Ach.) = abbrei'iata Reeve {Ach.) 

bacillaris Mousson {Tornat.) = oblonga Pease {Tornat.) 

baileyana Gulick {Ach.) = splendida Newcomb {Ach) . 

baldwini Ancey {Hyalinia) = pauxilla Gould {Vitrea) 

baldtvinii Newcomb {Ach) = magna Adams {Amastra) 

bellula Smith {Ach.) =■ ligata Smith {Ach.) . 

brevis Pfeiffer {Ach.) = nucleola Gould {Amastra) 

bron?iia>ta Philippi {Helicina) = rotelloidea Mighels {Helicina) 

bulbosa Gulick {Ach.) = picta Mighels, var. {Amastra) 

caesia Gulick {Ach.) = buddii Newcomb {Ach.) . 

Candida Pfeiffer {Bui) = ovata Newcomb {Ach.) 

castanea Reeve {Ach.) = vulpina Fer. {Ach.) 

chlorotica Pfeiffer {Neivcombia) = spirizona Fer., var. {Amastra) 

cinerosa Pfeiffer {Ach.) = decora Fer., var. {Ach.) 

clara Pfeiffer {Ach.) = striatula Gould (Leptach.) 

Clementina Pfeiffer {Ach.) = abbrei'iata Reeve {Ach.) . 

cognata Gulick {Ach.) = casta Newcomb {Ach.) . 

concavospira Pfeiffer {Ach.) = napus Pfeiffer {Ach.) 

cnncidens Gulick {Ach.) = decora Fer. {Ach.) 

coniformis Gulick {Ach.) = decora Fer., var. {Ach.) 

consanguinea Smith {Ach.) = livida Swainson {Ach) 

conspersa Pfeiffer {Ach) = reticulata Newcomb {Amastra) 

contigua Pease {Mclania) = newcombii Lea {Melania) . 

contracta Gulick {Ach) = curta Newcomb {Ach) 

convexa Nuttall {Neritina) = cariosa Gray {Neritina) . 

corrugata GuUck {Ach) = rugosa Newcomb {Ach) 



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conisca Gulick {Ach) = terebra Newcomb {Ach.) 
costulosa Pease {Pupa) = nnvcombi Pfeiffer {Pupa) 
crassidentaia Pfeiffer {Ach.) = vulpina Fer., var. {Ach.) 
crocea Gulick {Ach.) = lignaria Gulick, var. {Ach>i 
cylindrata Pease {Leptach.) = exilis Gulick {Leptach.) . 
decepta Adams {Ach.) = variegata Pfeiffer {Amastra) . 
detisilineata Reeve {Partula) = radiata Gould {Ach.) . 
dentata Pfeiffer {Ach.) = labiata Newcomb {Leptach.) . 
deshayesii Morelet, pars {Ach.) = assimilis Newcomb {Amastra) 
deihayesii Morelet, pars {Ach.) = biplicata Newcomb {Amastra) 
dimidiata Pfeiffer {Ach.) = cingula Mighels {Leptach.) 
dimondi Adams {Ach.) = gravida Yii. {Afiiastra) 
dimorpha Gulick {Ach.) = curia Newcomb {Ach.) 
disculus Pfeiffer {Helix) = exaequata Gould {Pliilonesia) 
discus Pfeiffer {Helix) = exaequata Gould {Philonesia) 
diver sa Gulick {Ach.) = vulpina Fer., var. {Ach.) 
dumartroyi Souleyet {Partula) = auricula Fer. {Auriculella) 
dunkeri Pfeiffer {Ach.) = producta Reeve {Ach.) . 
eburnea Gulick {Ach.) = tappania}ia Adams {Ach) 
emmcrsoiii Newcomb {Ach.) = livlda Swainson {Ach.) 
fascia ta Gulick {Ach.) = tappaniana Adams {Ach.) 
ferussaci Pfeiffer {Lam.) = sanguinea Newcomb {Amastra) 
filocostata Pease {Helix) = paucicostata Pease {Endodonta) 
flavida Clessin {Physa) = compacta Pease {Limnaea) 
flavidus Gulick {Apex) = cestus Newcomb {Ach.) 
forbesiana Pfeiffer {Bui.) = cesttis Newcomb {Ach.) 
fragilis Gulick {Ach.) = guttula Gould {Leptach.) 
fragilis Souleyet {Succinea) = cepulla Gould {Succinea) 
fricki Pfeiffer {Ach.) = glabra Newcomb {Ach.) . 
fricki Pfeiffer {Helix) = lamellosa Fer. {Endodonta) 
fuliginea Pfeiffer {Ach.) = bicolor Jay ( Carelia) . 
fuliginosa Gould {Ach.) = tristis F^r. {Amastra) . 
fulva Pfeiffer {Ach.) = variabilis Newcomb {Ach.) 
fuscolineata Smith {Ach.) = vulpina Fe'r. {Ach.) . 
fuscozona Smith {Ach.) -■ buddii Newcomb {Ach.) 
fusiformis Pfeiffer {Ach.) = mucrofiata Newcomb {Amastra) 
gigantea Newcomb {Ach.) = violacea Newcomb {Amastra) 
gigas Lesson {Neripteron) = granosa Sby. {Ahritina) . 
glauca Gulick {Ach.) = zonata Gulick {Ach.) 
globosa Pfeiffer {Ach.) = vittata Reeve {Ach.) 
goniostoma Pfeiffer {Ach.) = affinis Newcomb {Amastra) 
gouldi Pfeiffer {Bulimus) = radiata Gould {Ach.) 
granifera Gulick {Ach.) = accincta Mighels {Leptach.) . 
grossa Pfeiffer {Ach.) = porphyria Newcomb {Amastra) 
gulickii Smith {Apex) = cestus Newcomb {Ach.) . 
gummea Gulick {Ach) = guttula Gould {Leptach.) 
hartmani Newcomb {Leptach.) = ? extincta Pfeiffer {Amastra) 
hawaiietisis Baldwin {Ach.) = physa Newcomb {Ach.) 
herbacea Gulick {Ach.) = decipiens Newcomb {Ach.) 



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hybrida Newcomb (Ach.) = producta Reeve {Ach.) 
indtita Gulick {Ach.) = tnarmoraia Gould (Ach.) 
iniiolahilis Smith (Apex) = decora Fer., var. (Ach.) 
inortiatd Mighels (Ach.) = turritclla Fer. (Amastrd) 
insignis Reeve (Ach.) = virgulata Mighels (Ach.) 
intercarinata Mighels (Helix) = cotitorta Fe'r. (Endodonta) 
johnsotii Newcomb (Ach.) = bilincata Reeve (Ach.) 
juncea Gulick (Ach.) = casta Newcomb (Ach.) 
lacrima Gulick (Ach.) = glutinosa Pfeiffer (Leptach.) 
lactea Gulick (Ach.) = variabilis Newcomb (Ach.) 
lagena Gulick (Ach.) = labiata Newcomb (Leptach^ 
leucophaeus Gulick (Apex) — decora Fer., var. (Ach.) . 
leucorraphe Gulick (Apex) = decora Fe'r., var. (Ach.) . 
leucosonus Gulick (Apex) = napus Pfeiffer (Ach.) . 
liliacea Pfeiffer (Ach.) = viilpiiia Fer., var. (Ach.) 
liliaceus Gulick (Apex) = cestus Newcomb (Ach.) 
litnbata Gulick (Ach.) = byronii Wood (Ach.) 
liratits Pfeiffer (Bulimns) = plicata Pfeiffer (Neivcombia) 
livida Pfeiffer (Ach.) = vulpina Fer. (Ach.) . 
lugubris Chemn. (Turbo) - apexfulva Dixon (Ach.) 
lurida Pfeiffer (Ach.) = castanea Pfeiffer (Auriculella) . 
licteola Fer. (Helix) = 1 turritella Fer. (Amastra) 
macrosloma Pfeiffer (Ach.) = taeiiiolata Pfeiffer (Ach.) . 
tnagdalenae Ancey (Pupa) = lyrata Gould (Pupa) 
mahogani Gulick (Ach.) — byronii Wood (Ach.) . 
tnaiioaensis Newcomb (Ach.) = ventulus Fer. (Amastra) 
margarita Pfeiffer (Ach.) = accincta Mighels (Leptach^ 
tnelampoides Pfeiffer (Ach.) = ventulus Fer. (Amastra) 
melanostoma Newcomb (Ach.) = byronii Wood (Ach.) . 
microstoma Gould (Ach.) = textilis Fer. (Amastra) 
monacha Pfeiffer (Ach.) = multilineata Newcomb (Ach.) 
multicolor Pfeiffer, pars (Bui.) = byronii Wood, var. (Ach.) 
multicolor Pfeiffer, pars (£ul.) = oviformis Pfeiffer (Ach.) 
mustelina Mighels (Ach.) = decora Fer., subsp. (Ach.) 
myrrhea Gulick (Ach.) = gouldi Newcomb (Ach.) 
?mcca Gould (Pupa) = pediculus Shuttlw., var. (Pupa) 
neglectus Smith (Apex) = decora Fer., var. (Ach.) 
newcombi Pfeiffer (Helix) = caperata Gould (Godwinia) 
newcombi Pfeiffer (Ach.) = turricula Mighels (Carelia) 
fiewcombi Pfeiffer (Succinea) = patula Mighels (Succinca) 
navcombianus Pfeiffer (Bulimus) = pfeifferi Newcomb (Newcombia) 
nigrolabris Smith (Amastra) = spirizona Fer., var. (Amastra) 
nivosa Newcomb (Ach.) = abbreviata Reeve (Ach.) 
fiobilis Pfeiffer (Ach.) = lorata Fer. (Ach.) . 
nucleola Reeve (Ach.) = albolabris Newcomb (Amastra) 
nuttalli Recluz (Neritina) = cariosa Gray (Neritina) . 
nympha Gulick (Ach.) = byronii Wood, var. (Ach.) 
oahuensis Green (Achatina) = turritella Fer. (Amastra) 
obclavata Pfeiffer (Ach.) = sandivicensis Pfeiffer (Leptach. 



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FAUNA HAWAIIENSIS 



obeliscus Reeve {Achafiiia) = turricula Mighels (Carelia) 
obeliscus Pfeiffer {Ach.) = navcombi Pfeiffer {Auricuklla) 
obliqua Gulick {Ach.) = bulimoides Swainson {Ach.) 
obscura Newcomb {Ach.) = tnoesia Newcomb {Amasira) 
obtusangiila Pfeiffer (Helix) = exaeqiiata Gould {Philonesia ) 
odavula Paetel {Leptach.) = sandwicensis Pfeiffer {Leptach.) 
oomorpha Gulick {Ach.) = bulimoides Swainson {Ach.). 
o-amihiemis Chamisso {Auric.) = auricula Fer. {Auriculella) 
pallida Nuttall {Ach.) = lorata Fer. {Ach) .... 
papulosa Gray {Neritind) = granosa Sby. {Neriiina) 
parvula Gulick {Ach.) = vitreola Gould {Leptach.) 
patula Sniitli {Auric.) = diaphana Smith {Auric.) 
perforata Gulick {Ach.) = terebra Newcomb {Ach.) 
perversa Swainson {Ach.) = decora Fe'r. {Ach.) 
pexa Gulick {Ach.) = buddi Newcomb {Ach.) 
phaeozona Gulick {Ach.) = ovata Newcomb {Ach.) 
pica Swainson {Ach.) = apexfulva ],)ixon {Ach.) . 
planospira Pfeiffer {Ach.) = decipiens Newcomb {Ach.) 
platystyla Gulick {Ach.) = glabra Newcomb {Ach.) 
plumata Gulick {Ach.) = buddii Newcomb {Ach.) 
polyinorpha Gulick (Ach.) = apicata Pfeiffer, var. {Ach.) 
ponderosa Ancey {Auric.) = crassula Smith {Auric.) 
prasina Reeve {Ach.) = olivacea Reeve {Ach.) 
pulcherrima Swainson {Ach.) = byjvnii Wood {Ach.) . 
pulcherrima Reeve {Ach.) = vulpina Fer., var. {Ach.) . 
pulla Pfeiffer {Ach.) = pusilla Newcomb {Amasira) 
pumila Gulick {Ach.) = modcsta Adams {Amastra) 
pusillus Gould {Helix) = pauxillus Gould {Vitrea) 
pygmaea Smith {Ach.) = curia Newcomb {Ach.) . 
radiata Pfeiffer {Ach.) = viridans Mighels {Ach.) 
recta Newcomb {Ach.) = byronii Wood, var. {Ach.) 
reeve i Adams {Ach.) = livida Swainson {Ach.) 
rhodoraphe Smith {Ach.) = curia Newcomb {Ach.) 
rohvi Pfeiffer {Bulimus) = virgulata Mighels {Ach.) 
rotundata Gould {Succinea) = patula Mighels {Succinea) 
rubens Pfeiffer {Ach.) = mastersi Newcomb {Amastra) 
rubiginosa Gould {Helix) =jugosa Mighels {Endodonta) 
rubiginosa Newcomb {Ach) = taeniolata Pfeiffer {.4ch.) 
rudis Pfeiffer {Ach.) - spirizona Fer., var. {Ar?iastra) . 
rustica Gulick {Amastra) = affinis Newcomb {Amastra) 
rutila Newcomb {Ach.) = viridans Mighels {Ach.) 
satid7viclunsis Clessin {Pliysa) = binominis Sykes {Limnaea) 
sandwichcnsis Philippi {Limnaea) - oahuetisis Soule)'et {Limnaea 
sandivichcnsis Deshayes {Neritina) = cariosa Gray {Neritina) 
scitula Gulick {Ach.) = decipiens Newcomb {Ach.) 
semicarinata Newcomb {Ach) = variabilis Newcomb, var. {Ach. 
seminigra Lamarck {Monodo7ita) = apexfulva Dixon {Ach.) . 
setigera Gould {Helix) = hystrix Pfeiffer {Endodonta) . 
similaris Pease {Amastra) = rugulosa Pease, var. {Amastra) 



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simulans Reeve {Ach.) = vitlata Reeve {Ach.) . 
sinistrorsa Chamisso {Auric.) = auricula Y€x. {Auric.) 
solida Gulick {Ach.) = spkndida Newcomb {Ach.) 
solidissima Sby. {Neritina) = cariosa Gray {Neritina) 
soukyeti Ancey {Succinea) = cepulla Gould {Succinea) 
spadicea Gulick {Ach.) = ovata Newcomb {Ach.) . 
stewarti Green {Achat ina) = vidpina Fer., var. {Ach.) 
siriatula Pease {Pupa) = lyrata Gould {Pupa) 
subula Gulick {Ach.) ^ gracilis Pfeiffer {Leptach.) 
suffusa Reeve {Ach.) = gravida Fer. {Amas/ra) . 
swiftii Newcomb {Ach.) = cestus Newcomb, var. {Ach.) 
talpina Gulick {Ach.) = gouldi Newcomb {Ach.). 
torrida Gulick {Ach.) = rugosa Newcomb {Ach.) 
tricolor Smith {Ach.) = vulpina Fer., var. {Ach.) . 
triticea Gulick {Ach.) = oryza Pfeiffer {Leptach.) . 
tuba Gulick {Ach.) = tappaniana Adams {Ach.) . 
tuberans Gulick {Apex) = cestus Newcomb {Ach.) 
timief actus Gulick {Apex) = vittata Reeve {Ach.) 
turbiniformis Gulick {Apex) = cestus Newcomb {Ach.) 
turgida Newcomb {Ach.) = cestus Newcomb {Ach.) 
umbilicata Pfeiffer {Ach.) — petricola Newcomb {Amastra) 
umbrosa Baldwin {Amastra) = pullata Baldwin {Amastra) 
undosa Gulick {Ach.) ~ perdix Reeve {Ach.) 
undulata Newcomb {Ach.) = curta Newcomb {Ach.) . 
valida Pfeiffer {Ach.) = cestus Newcomb, var. {Ach.) . 
va}-ia Gulick {Ach.) = vulpina Fer., var. {Ach.) . 
variabilis Pease {Carelia) = olivacea Pease {Carelia) . 
vetitrosa Pfeiffer {Ach.) = lorata Fer. {Ach.) 
veuulata Newcomb {Ach.) = producfa Reeve {Ach.) 
versicolor Gulick {Apex) = decora Fer., var. {Ach.) 
vespertina Baldwin {Ach.) = apicata Pfeiffer, var. {Ach.) 
vestita Mighels {Ach) = ? vittata Reeve {Ach.) . 
virens Gulick {Ach.) = vulpina Fer. {Ach.) . 
viridans Reeve {Ach.) = livida Swainson {Ach.) . 
volutata Gould {Limnaea) = oahuensis Souleyet {Limnaea) 
wheatleyi Newcomb {Ach.) = glabra Newcomb {Ach.). 



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§ 5. List of unidentified, or erroneously recorded, forms. 



antoni Pfeiffer {Helicind) . 
aperta Lea {Succinea) 
apicalis Ancey {Siccci?iea) 
approximata Shuttlw. {Succinea) 
crassilabris Philippi {Helicina) 
exserta Pfeififer {Helix) . 
ferruginea Baldwin {Amasfra) 
fornicata Gould {Helix) . 
fulgora Gould {Helicina) . 
kauaiefisis Pfeiffer {Bulimus) 
olesonii Baldwin {Achatinella) 
pisum Philippi {Helicina) 
pitdorina Gould {Succinea) 
pumicatus Mighels {Bulimus) 
pusilla Gould {Partula) . 
sandwicensis Pfeiffer {Spiraxis) 
sandiiiicliensis Pfeiffer {Helix) 
striolata Pease {Opeas) 
ienerritna Ancey {Succinea) 
terrestris Pease {Partula) 
testudinea Baldwin {Amastra) 



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413 



THE EARTHWORMS OF THE HAWAHAN ARCHIPELAGO. 

By Frank E. Beddard, M.A., F.R.S., Prosector and Vice-Secretary of the 

Zoological Society of London. 



Some of the specimens which I have received from the Hawaiian archipelago have 
been already described by me'. Since the publication of that paper Mr Perkins has 
sent a second series of bottles containing a large number of fresh individuals. In the 
present memoir upon the earthworm fauna of this part of the world I deal with the 
entire series of specimens and attempt to give a complete account of all the earthworms 
which have been described from the Hawaiian islands, whether they are or are not 
contained in the collections which I have myself examined. The collections made by 
Mr Perkins consist of so many individuals that they probably present a very fair speci- 
men of the Oligochaetous fauna of Hawaii. It is therefore permissible to point out what 
appear to me to be justifiable deductions from the material examined. The fact that the 
second set of specimens contained hardly anything that was not in the first set supports 
my contention that I have been able to study a very representative collection. 

Dr Michaelsen" in criticising my previous paper upon this subject advanced the 
opinion that there are no truly indigenous worms in these oceanic islands. I myself 
pointed out the absence of really peculiar forms, a general feature of oceanic islands and 
which at least argues their comparatively short existence. Dr Michaelsen attributes 
the entire earthworm fauna to transference by man. A further study of the matter 
inclines me to agree with him. 

There are many species of Lumbricidae contained in the collections which I 
have examined ; and the list which I gave originally can be increased. But the subject 
does not demand, I believe, more than a mere list of the species. They are clearly to 
be regarded as importations due to man. 

' On some Earthworms from the Sandwich islands, &c. P. Z. S. 1896, pp. 194-2 11. 
^ Oligochaeten von den Inseln des Pacific. Zool. Jahrb. Syst. xii. 1899, p. 211. 



414 FAUNA HAWAIIENSIS 

Allolobophora Savigny. 

(i) Allolobophora putris Hoffm. (This apparently is the same as Kinberg's 
' ' Hypogaeon kavaicum. ' ' ) 

(2) A. foetida (Savigny). 

(3) A. caliginosa (Savigny). 

(4) A. nordenskioldii Eisen. 

(5) A. limicola Michaelsen. 

(6) A. rosea (Savigny). 

PoNTOscoLEX Schmarda. 

Pontoscolex hawaiiensis Beddard, P. Z. S., 1896, p. 196. 

The ubiquitous genus Pontoscolex occurs in the Hawaiian archipelago. I formed a 
new species for the representatives of this genus which were collected by Mr Perkins 
chiefly on the ground that the dorsal vessel was usually double for a certain extent. 
Dr Michaelsen has criticised my conclusion ; and it may be that he is right. In any 
case the genus and species which are at least hard to distinguish from the South 
American P. corethrurus occur in the most widely separated regions of the globe. 
Dr Eisen however' has lately commenced a detailed study of this genus, so that the 
matter of the specific identity or difference of the specimens of Pontoscolex found 
scattered over the world had better be left alone for the present. 

Amyntas Kinberg. 

The main earthworm inhabitants of this archipelago belong to the genus Amyntas 
as I think (following Michaelsen') it should now be called. The much better name 
Perickaeta was used for a Dipteran genus before it was applied to an earthworm ; and it 
appears to me, in spite of the ingenious protest of Horst^ that there is no way of escaping 
from the conclusion that a name once used cannot be resuscitated. One unrecognisable 
species "Perickaeta corticis" has been described by Kinberg''. The remaining species 
are the following : — 

(i) Amyntas peregrinus Fletcher. 

Perickaeta peregrhia Fletcher, Proc. Linn. Soc. N. S. W. (2), i. p. 969. 
Perickaeta 7ttolokaiensis Beddard, P. Z. S. 1896, p. 201. 
Perickaeta floiveri Benham, Journ. Linn. Soc. xxvi. p. 217. 

' Researches in American Oligochaela, &c. P. Calif. Ac. Sci. (3), 11. p. 87. 

' Terricolen von verschiedeiien Gebieten der Erde. JB. Hamb. wiss. Anst. xvi. Beiheft 2. 

^ Zool. Anzeig. 1890, p. 6. 

^ Annulata nova. Ofv. k. Svensk. Ak. Forh. 1866. 



EARTHWORMS 415 

Michaelsen has suggested that the species which I described in my prehminary 
paper as Perichaeta molokaieusis, is really identical with Fletcher's Perichaeta peregrina, 
or is at least to be regarded as a " fragliches synonym." At the time that I described 
that species I was not so convinced as I am now of the unimportance of size as a 
distinguishing characteristic of species of this genus. Fletcher described his species as 
being 19 cm. in length, i.e. nearly double the length of the individuals of ''Perichaeta 
molokaieusis" examined by myself Moreover Fletcher has not given any details about 
the clitellar setae, beyond stating that they are present. This again is a matter which 
is apparently not of such importance as I thought ; that is to say, the same species may 
have setae upon one, two or three or perhaps even none of the clitellar segments. 

There can I think be no doubt as to the identity of Benham's ''Perichaeta 
flowej'i" with the present species. Benham states that there are 12 setae between the 
male pores, which is the number given by Fletcher. I counted 15 in my specimens. 
But the difference is clearly negligible. Benham particularly mentions the clitellar 
setae as being present on segment 16 only, a state of affairs which I found also. None 
of the three forms comprised in the present species possess copulatory papillae ; hence 
it is now probably to be taken as a character of this species. In this as in all other 
points 1 can detect no differences between the descriptions of Benham and of myself. 
We are clearly dealing with the same species, which being with very great probability — 
almost amounting to certainty — identical with that described as " Perichaeta peregrina" 
by Fletcher, must bear that name. I now give for the sake of others who may doubt 
this identification a description of my specimens. 

The two individuals of this species which I have examined were 103 and 81 mm. 
respectively in length. The former specimen possessed 88 segments, the latter 93. 

The prostomium is small and continued by grooves on to the first half of the first 
segment. 

The dorsal pores commence upon segments 10, 11, and are visible upon the 
clitellum. 

The clitellum occupies the usual segments, 14 — 16, and has few setae upon its last 
segment. 

The male pores are not jarominent and are separated by i 5 setae. 

I observed no genital papillae. 

The first septum separates segments 4, 5 ; none are thickened specially. 

The gizzard occupies the usual segments which are not divided by septa. 

The intestine begins in 1 5 ; the caeca are present and not large. 

The sperm sacs are large and occupy segments 11, 12. 

The spermiducal glands extend from segments 17 — 21 or 22; they are broken 
up into lobes which have to some extent a relation to the segmentation of the gland. 
The curved duct communicates directly with the exterior and not through the inter- 
mediary of a terminal dilated sac. 

F. H. II. 54 



41 6 FAUNA HAWAIIENSIS 

The spermathecae are four pairs in 6 — 9. The pouch is sharply marked off from 
the long duct. The diverticulum ending in an oval dilatation is about as long as the 
duct part of the main pouch. 

Hab. Molokai and Mauna Loa. 



(2) Ainyntas heterochaeta Mich. 

Perichaeta heterochaeta Michaelsen, Abhandl. nat. Vereins Hamb. xi. p. 6. 
P. indica Michaelsen, Arch. f. Naturg. 1892, jj. 33; nee/*, indica Horst, Vermes 
in Midden Sumatra, iv. p. 4. 

It is rather a curious fact that the non-identity of the worm which has been called 
by many persons, including myself, Perichaeta indica with the species described by 
Dr Horst under that name in the memoir quoted above has not been noticed. In that 
memoir Dr Horst distinctly figures a terminal sac (" Kopulationstasche ") to the duct of 
the "prostate" gland. His figures of " Eine Perichaeta von Java"' on the other hand 
do not show this duct with such a terminal swelling and refer to the species which has 
since been called Perichaeta indica. It is clear that the proper name to refer to this 
specimen of Dr Horst must be Dr Michaelsen's name of Perichaeta heterochaeta, in 
which no such copulatory pouch is mentioned and which in other respects agrees with 
the worm which has everywhere received the name of Perichaeta indica. If it were 
certain, which it is not, that M. Vaillant described only one species under the name of 
Perichaeta cingulata, then that would have to be the name for the species described by 
Horst, for it agrees in the presence of the terminal sac where the male gland opens on 
to the exterior, and in some other points. 

Dr Michaelsen would include as synonymous with this species my Perichaeta 
nippojiica ; I think that that species may be synonymous. But that is a matter which I 
shall enter into on a future occasion. 

I now think that I was wrong in differentiating the species P. perkinsi. Dr 
Michaelsen, chiefly for the reason that he received an example from Ceylon with papillae 
near to the male pores, identified my species with the one called here Amyntas hetero- 
chaeta. I should mention however that the fact that the union of the vas deferens 
with the male duct is not until near to the external orifice appears to characterise at least 
the individual which I examined. I found in glycerine preparation of two examples 
of undoubted "indica" that there was the more general union shortly after the duct 
emero;ed from the gland. 

Dr Michaelsen and I myself have called attention to the variability which this 
species exhibits in the presence and number of the anterior papillae and in the presence 
or absence of the glandular part of the male terminal apparatus. Among the very 

' Niederl. Arch. f. Zool. iv. 



EAR THWORMS 4 1 7 

numerous examples which I have examined from the Sandwich Islands I find the 
following state of affairs with regard to these variable structures. In 22 examples there 
were no glands at all ; 13 had glands ; in 26 specimens the glands were either small and 
on both sides or only present and small or well developed on one side. The proportions 
seem to show that the gland is disappearing. As to the head papillae — there were none 
at all in 24 ; in 14 there were three pairs on 7, 8 ; in 3 there were pairs on 7, 8, 
9 ; in one there were pairs on 8, 9 ; in 10 there was a pair on 8 ; in one there 
were pairs on 6, 7, 8 ; in 52 there were various degrees of asymmetry, sometimes 
none being present on one side. 

Examples of the species were obtained on Maui, Mauna Loa Hawaii, Halemanu 
Kauai, Kilauea Hawaii, Olaa Hawaii, Haleakala Maui, lao valley Maui, Honolulu 
in imported earth from China. 



(3) Amyntas hesperidum Beddard. 

Perichaeta hesperidum F. E. Beddard, Proc. Zool. Soc. 1892, p. 169. 
Perichaeta sandvicensis Id., ibid. 1896, p. 203. 

In my earlier paper upon this species founded upon the first gathering of Hawaiian 
worms I instituted a new species for some smallish worms from several islands of the 
archipelago. I have since re-examined the two original specimens of Amyntas hespei'i- 
diim which I have still by me, and have compared them with some fresh individuals 
undoubtedly belonging to the same species but coming from Hong Kong. The result 
is that I have to make one or two slight corrections in my earliest account of 
A. hespeHdum. I thought that I had noted a small terminal muscular bursa in that 
species ; but on again studying the specimens and comparing them carefully with others 
I find that what I took to be this distinctive structure was only the commencement of 
the thick investing layers of the spermiducal gland duct as it traverses the body wall. 
There is in fact no terminal bursa. In all the specimens the spermathecae, though 
lying in segments 7 and 8, as I correctly stated, open backwards, i.e. in the inter- 
segmental furrows 7, 8 ; 8, 9, as I also stated. I now find that this is also the case 
with Perichaeta sandvicensis. The spermathecae as a rule lie in the 7th and 8th 
segments but open at the posterior margins of those segments. In both worms the 
diverticulum is coiled and the spermiducal gland has a rather sinuous duct which passes 
rather forwards on its way from the gland to the exterior. In short I can detect no 
differences at all between the individuals which I have referred to two species. The 
older name must clearly have the priority and thus I must term these Sandwich Island 
worms Amyntas hesperidum, inapt though the name undoubtedly is. 

This is a prevalent species in the gatherings from the islands. In my preliminary 
account of the Sandwich Island worms I recorded it from Mauna Loa, Lanai, Hawaii 

54—2 



4i8 FAUNA HAWAIIENSIS 

and Molokai. I have seen in the second collection forwarded to me additional speci- 
mens from Mauna Loa. I can thus improve somewhat upon my original description of 
this species. It is a smallish slender worm measuring up to loo mm. in length with a 
diameter of about 3 mm. The number of segments is curiously constant in the indivi- 
duals which I selected for counting. In two the number was 105, in a third 104. The 
lengths of these specimens varied somewhat — from 82 through 98 to 100 mm. It is 
interesting to note the constancy of the number of segments. The differences in length 
are of course not sufficient to be of importance and are to be accounted for by the 
different degrees of contraction of the individuals. 

The dorsal pores commence between segments 11, 12, and are visible upon the 
clitellum. 

The setae of a given series of segments number as follows : i. 2 i ; 5. 33 ; 12. 52 ; 
16. 53. The setae of the first two segments are small; those of the next four are 
stronger, after which segments they again diminish in size. 

The clitellum is sharply marked off from the segments adjacent to it and both 
commences and ends with its own proper segments. I could not discover any setae 
upon it. 

Neither could I find anyvi^here upon the body of the worms genital papillae. The 
male pores are upon the usual segment ; they are slightly expanded transversely and 
have therefore an eye-like outline. They are fairly conspicuous ; in one case the end 
of the spermiducal gland duct was protruded for a little way. The two pores are 
separated by 18 setae. 

There are three fairly stout intersegmental septa in front of the gizzard which are 
bound to each other by numerous muscular threads in the usual way ; after the gizzard 
come two strong septa to the anterior of which the gizzard itself is bound at its posterior 
end by at least five muscular straps. In a specimen from Lanai septum 8/9 was 
present but thin. 

The gizzard has the usual position that it has in this genus. The intestinal caeca 
are present, but are small and simple ; they occupy not more than two segments. 

The last of the "hearts" is in segment 13. 

The two pairs of sperm sacs are in segments 11, 12 ; the sperm reservoirs as also 
usual in segments 10, 11; of the latter the anterior pair are sometimes larger than the 
posterior 

The spermiducal glands are much incised and occupy not more than three 
segments. Their muscular duct is longish and curved and is unprovided with a 
terminal copulatory dilatation. As to the form of the spermiducal glands it is often 
possible to use their characteristics as apparently valid specific distinctions. But it is 
necessary to be accurate in their delineation and cautious as to laying too much stress 
upon certain features in distinguishing species, as is shown by the present species. In 
most of those which I examined the gland had a somewhat ear-like form, the lower 



EARTHWORMS 419 

margin curving upwards and forwards like the lobe of the ear. But in one example the 
gland was quadrangular though only occupying four segments and deeply incised in 
correspondence therewith. 

I'he spermathecae are two pairs and lie in segments 7 and 8, but open on 7, 8 ; 
8, 9. The oval pouch has a moderately long duct to which is appended a small twisted 
diverticulum which is sometimes longer and sometimes shorter. 

We can extract from the foregoing the following definition of the species : 
Size small, 100 mm. ; number of segments 105. Dorsal pores from 11, 12. 
Number of setae per segment up to 53. Clitellum 14 — 16, without setae. 
No papillae. Septum 8/9 missing. Caeca present. Last heart in 13. 
Sperm sacs 11, 12. Spermiducal glands not very large; duct without end sac. 
Spermathecae 7, 8, with twisted tubular diverticulum. 

Remarks. Dr Michaelsen has put forward grounds for believing that this 
species is really Dr Horst's Amyntas annulata. I myself suspected a possible identity. 
Dr Horst's original description of annitlafa, written some years ago when there was 
no difficulty in distinguishing from each other the very few species of the genus at 
that time known, was hardly complete enough for present requirements. He gives me 
moreover by letter good reasons for denying the identity. 

Nor is there much change required to derive this form from the prevalent Amyntas 
hawayanus. I desire again to refer in connection with this possibility to a species 
described by myself some years since as Perichaeta kcsperidu»i. That species, two 
individuals, arrived together with a form which I shall refer to here, barbadcitsis. 
The worm differs however from barbadensis (I have satisfied myself by a renewed 
examination) in a number of points. The clitellum begins and ends sharply at the 
boundary lines of segments 13, 14, and 16, 17, and has no setae. The spermathecae 
are in segments 8, 9, or at any rate open on to the boundary lines 7, 8 ; S, 9. The 
pouches differ from those of hazvayaniis in having much coiled diverticula ; the duct of 
the spermiducal gland thins towards its end, is longish and rather curved and has not 
really a terminal bulbus as I .said in my original description. I have since met with 
other examples of the same worm from Hong Kong in a bottle containing also examples 
of barbadensis. I have examined four of these, all that I had. They have no setae on 
the clitellum, which commences and ends "sharply." The length is from 80 — 100 mm, 
There are no genital papillae nor are there setae upon any segment of the clitellum. 
The last heart is in segment 13 as is usual ; the caeca are quite normal in position and 
present no special features of interest. The sperm sacs have a constricted-off free end 
as in so many forms. The spermathecae are two pairs and lie in 8 and 9 or at least 
open in the intersegmental grooves 7, 8 ; 8, 9. The diverticulum of the pouches is only 
of moderate length — not so long or not longer than the pjouch — and is more or less 



420 FAUNA HAWAIIENSIS 

closely coiled. The duct of the spermiducal gland is directed rather forward, as is so 
often the case in barbadensis, and is rather curved, especially at the end, where it is 
distinctly thinner. This thin termination was not observable in one individual in which 
the male pores had the appearance of being somewhat everted. These worms are 
undoubtedly my hesperidum. 

It is interesting to find from three distant parts of the world specimens of a worm 
associated with a form from which they can be easily derived, by a reduction of the 
number of spermathecae, and by an emphasising of the slightly coiled diverticulum of 
the parent (.'') form, by the loss of genital papillae and setae on the clitellum. The 
coincidences are at least noteworthy. 

If we are to assume that the migration of the genus Aviyiitas from the Oriental 
region is due always to the interference of man, it is most peculiar that they should have 
been exported in lots of corresponding species. I do not however at present do more 
than emphasise the facts which are as has been stated above. 



(4) Amyntas hawayanus Rosa. 

Perichaeta hawayana Rosa, Ann. k. k. Hofmus. Wien, 1891, p. 396. 

PericJiaeta berviudensis Beddard, P. Z. S. 1892, p. 160 

Perichaeta barbadensis Beddard, ibid. p. 167. 

Perichaeta morrisi Beddard, ibid. p. 166. 

Perichaeta mauritiana Beddard, ibid. p. 1 70. 

Perichaeta mandhorensis Michaelsen, Arch. f. Naturg. 1892, p. 241. 

Perichaeta pallida Michaelsen, ibid. p. 227. 

Perichaeta amazonica Rosa, Atti R. Ac. Torino, 1894, p. 4. 

Perichaeta ciipnlifcra Fedarb, Proc. Zool. Soc. i8g8, p. 445. 

The collection contains a considerable number of examples of A. hawayanus. 
These show so many variations that I believe myself to be able to justify the above 
rather formidable list of synonyms, which are a little more extensive than the list given 
by Dr Michaelsen' in a recent paper. My original description of Perichaeta bermu- 
densis was published when I was unaware of Dr Rosa's Perichaeta hawayana, though 
his publication' seems to antedate mine. I was led in my " Monograph of the Oligo- 
chaeta " to adhere to my species bcrmitdensis on account of the fact that Dr Rosa did 
not mention in his description the larger size of the setae upon the anterior segments, 
nor the presence of setae upon the last segment of the clitellum. The number of 
papillae in the neighbourhood of the male pores seemed too to be different in the two 
series of worms from Hawaii and from the Bermudas. In the series of specimens in the 

^ Die Terricolen des Madagassischen Inselgebietes. Abh. senck. naturf. Ges. 1897, p. 234. 
^ Die exotischen Terricolen, &c. Ann. k. k. Hofmus. Wien 1891, p. 396. 



EARTHWORMS 42 1 

collection made by Mr Perkins I find the following variations in structure, from what 
may perhaps be regarded as the typical organisation of this species. The number of 
the papillae in the neighbourhood of the male pores varies ; I have found only one or 
two, three or four; Rosa says two or three. In one example however I found six of 
these papillae on each side. 

The number of the papillae therefore does not allow of a separation of /iazmya?nts 
and bermudcnsis. 

The setae upon the anterior segments are larger than those which follow ; in one 
example segments 4 — 7 were furnished with these larger setae. This was originally 
one of the reasons for separating bei^7nudensis from Jiawayanus. 

The clitellum was described by Rosa to stop short at the middle or thereabouts of 
the 1 6th segment. I have observed both this arrangement and that generally found 
in bermudensis, i.e. that the clitellum does not commence accurately at the beginning of 
the 14th segment while it stops short as in the typical hawayanus. In one example the 
clitellum was exactly coincident with segments 14 — 16. 

The clitellum has usually setae upon its last segment, i.e. the i6th of the body. 
There are ten to fifteen of these setae. In two examples I could see no setae anywhere 
upon the clitellum. This seems to have been the case with the individuals examined 
by Rosa. A. bermudensis appears to always have setae upon this segment. 

A feature not yet recognised in the worms which I referred to the species hernui- 
densis was found in two examples ol hawayanus. In one of them there was a pair of 
papillae anterior in position lying on the 7th segment near to its posterior end and the 
orifices of the spermathecae. In a ripe individual there was but one of these papillae, 
that of the right side. This fact will be seen presently to bear upon the identity of the 
present species with others hitherto supposed to differ specifically from it. As to 
internal characters the caecum of the intestine has not always the series of short out- 
growths on the under surface that has been described for this species and for bermuden- 
sis. The spermiducal gland is generally long, occupying segments i 7 — 22 about. Some- 
times the duct is given off at the top when the gland commences in segment 17. In 
one specimen the gland was much abbreviated and lay only in 18, 19 on one side and 
17, 18 on the other. This looks like a commencing loss of the gland which is known 
to occur in some other species, e.g. Aniyntas heterochaeta. The spermathecae seem 
always to lie in segments 6, 7, 8. In one example they were particularly large ; but, as 
this individual had no other features which seemed to remove it from the species, the 
difference in size (the pouches were as large as the gizzard) does not seem to be more 
than a variation to be neglected for systematic purposes. 

The size of Amyntas hawayanus varies to some extent. The greatest and least 
lengths which I observed were 150 and 69 mm. The number of segments varied 
between 97 and 73. 

Next as to the identity of Amyntas haivayanus with A. barbadensis — the original 



42 2 FAUNA HAWAIIENSIS 

specimens of the latter were described by me from Kew Gardens, where they had been 
received from Barbados. In the collection of Sandwich- Islands worms before me there 
are a number of specimens of this species found at Honolulu in earth imported from 
China. Of these I have examined seven individuals. 

Their size presents no difficulty for identification. They vary from 99 to 140 mm. 
The clitellum occupies segment 14 to about the middle of segment 16. In two 
specimens I found setae to be limited to the i6th segment; in the others there were 
setae on all the clitellar segments, but very few on 14 and 15. In one specimen the 
numbers on the three segments beginning with 14 are 8, 3, 15 ; in another 2, 2, 10. 
These figures agree broadly with my previous observations upon this species. 

In several cases I found that the setae upon the anterior segments of the body are 
as in Aniyntas hawayanus larger than those posteriorly. Segments 3 — 8 appeared to 
be thus distinguished. 

The arrangement of the genital papillae is as follows : 

There are either two or three in the neighbourhood of the male pores, sometimes 
only one. They lie either in an oblique row or in the case where there are two, one 
behind the other. There is in fact no difference here from the conditions which obtain 
in haivayanus. 

In addition to these posteriorly placed genital papillae there are anterior papillae. 
One individual had a pair on segment 7 ; another a median papilla on the same seg- 
ment. There is here again no practical difference from A. haiuayanus. 

As to internal characters the prevalent number of spermathecae is three pairs 
situated as are those of A. hawayanus. In one specimen only were there but two pairs 
of these organs placed in segments 6, 7. The sperm sacs often, but not always, show a 
constriction near to the free end, by which a small " knob " is divided off from the rest 
of the sac. As in A. hawayanus there are at least often two pairs of egg sacs in 
segments 13, 14. The spermiducal glands are long, occupying segments 17 — 21, as in 
A. haivayanus, and as in that species there is no terminal " Kopulationstasche " into 
which the duct of the gland opens. A character which seems to be peculiar to these 
worms is the occasional duplication of the dorsal vessel. I found this in four out of 
seven examples ; the doubling commenced at the 20th segment or thereabouts, and the 
tube became single again about the 25th. The doubling was complete, the two halves 
not uniting at the septa where they traversed those plates. Of these variable characters 
there are only three which do not seem to occur in examples which have been referred 
to A. hawayanus and A. bermudensis. These are : setae upon segments 14, 15 ; occa- 
sional doubling of dorsal vessel ; knob-like processes of sperm sacs ; the presence of 
only two pairs of spermathecae. Were these or some of these characters united 
invariably together we might indeed separate the specimens as a different species ; but 
they do not. The one example with spermathecae in 6 and 7 only had, it is true, no 



EARTHWORMS 423 

marked difference in size between the anterior and the posterior setae found elsewhere 
among the examples ; and it had a median papilla upon the 7th segment, this segment 
being occupied in others by a pair of similar papillae. These characters however do 
not always coincide, for in my original paper describing the species Perichaeta barba- 
densis, I recorded the fact that in an individual with two pairs of spermathecae there 
was a single median papilla upon the 7th segment, as well indeed as another occupying 
a similar place in the i8th. To make a species of this worm we must characterise it by 
the two pairs of spermathecae and the median anterior instead of paired anterior 
papillae, as well as by the greater uniformity in the size of the body setae generally. 
In view of the variations which occur in individuals which no one would thus separate 
it seems to be unreasonable at least in the meantime to do this. 

I may perhaps be allowed to point out that I was justified on the facts as originally 
known in making a new species for these worms. They then differed as far as was 
known from Perichaeta haivayatia in having setae upon all segments of the clitellum, in 
possessing anterior as well as posterior genital papillae, and finally by generally having 
but two pairs of spermathecae. 

I shall now consider the probable identity of these forms with Perichaeta morrisi. 
This species was originally distinguished from its allies by the following assemblage of 
anatomical features: small size, 52mm. with however 93 segments; two pairs of 
spermathecae in 6, 7 ; median papillae in 7, 8 ; glandular bodies in the neighbourhood 
of the male pores were not seen to open by papillae ; but such glands are usually 
associated with papillae. Rosa' described later examples of what appears to be the 
same species. His examples were larger (up to 80 mm.); setae present on all of the 
segments instead of only 16; clitellum occupying the whole of segments 14 — 16 
instead of stopping towards the middle of 16 ; glands near male pores ; in one example 
a median papilla upon tS was noted ; others showed variation in the anterior papillae, 
in one a median papilla on 6 and a pair on 7 closely approximated in the middle line 
with a more lateral pair on the same segment and a median papilla on 8. Another had 
median papillae on 6 — 8 ; a third one only on 7. Dr Rosa also found, though it was in 
a rudimentary condition, the septum separating segments 8, 9. 

I have been able to compare with these descriptions some worms from Hong 
Kong among my stores of Oligochaeta. I examined many of these which I refer to 
the same species. 

One was 93 mm. long ; two papillae lay by each male pore, and on 7 there were 
three papillae, one median and two lateral. Setae were present on all segments of the 
clitellum. In other characters I found no differences from Perichaeta morrisi as 
described. 

In a second individual of 80 mm. length there were also two pores in the neigh- 

' Lombrichi raccolti a Sumatra &:c. Ann. Mus. civ. Geneva ser. 2a xvi. p. 516. 
F. H. II. SS 



424 FAUNA HAWAIIENSIS 

bourhood of the male pores, but side by side, instead of one in front of the other. 
Setae of clitellum only on i6. 

In a third there was but one papilla to each male pore, and the setae on the 
clitellum were limited to i6. None of these latter had any anterior genital papillae. 
A fourth example was 94 mm. in length, with setae only on the last segment of 
clitellum; the sperm sacs as in the species generally in segments 11, 12, but provided 
with the small terminal knobs such as I have just referred to in Perichaeta barbadensis. 
The spermathecae appeared to be in 7, 8, instead of 6, 7. In all the prostates were 
long and had no terminal bulb. I need not enumerate in detail the various arrange- 
ments of the genital papillae in these examples from Hong Kong ; but I may state 
generally that they varied excessively in this particular. There were often two to four 
papillae on the i8th segment between the male pores; it was very general to find a 
pair of papillae on segment 19 corresponding in position to the male pores on the fore- 
going segment. I observed a median papilla on each of segments 6 — 8 in one 
individual ; one was anomalous by reason of the fact that the 7th segment had no less 
than six papillae arranged in an irregular line along the middle of that segment. The 
spermathecae were as a rule two pairs in 6, 7. But this character was not absolutely 
fixed. One specimen had an additional spermatheca in the 8th segment, but on the 
right side only. In this individual moreover the generally missing septum 8 was present, 
a circumstance which Rosa has stated for Perichaeta niorrisi. Among the same worms 
there were three specimens of rather larger size. One of these was 135 mm. long and 
was the largest. It has 90 segments. In it the papillae were as much reduced as they 
ever are in this species. The larger worms with the fewer papillae and three pairs of 
spermathecae I consider to be the more typical hawayanzis. In this individual (to 
resume) the sperm sacs had constricted apices ; the spermathecae three pairs in 6 — 8. 
One papilla only to inside of male pore; setae on 16, those of segments 3 — 7 about 
enlarged. Obviously the same as this, but a little smaller, was a worm with two 
papillae by male pore and a single median one on 7. These larger specimens have the 
duct of the spermiducal gland bent into an U-shape ; in the smaller and more papillated 
worms the duct is usually slightly curved more in the direction of a large semicircle. 
If we are to accept this as a species we can find no character not found in examples of 
the forms already treated of, except that the two pairs of spermathecae may be a 
segment further behind. This seems to be hardly enough as a character whereby to 
separate the species. 

With regard to the identity of Perichaeta majiritiana I must chiefly refer to 
Dr Michaelsen. I may observe however that in the position of the spermathecae and 
the presence of setae upon one segment only of the clitellum, this supposed species 
agrees with an individual which I found myself unable to definitely distinguish from the 
form which I have called Perichaeta niorrisi. 

In two individuals which I refer to this species, and which are not the same that 



EARTHWORMS 425 

formed the basis of my original description of the species, I found the following 
characters. The length of one was 76 mm. There are a row of four papillae to the 
inside of each male pore. The setae on segments 3 — 7 are particularly strong. I 
found setae on the last segment of the clitellum, i.e. 16. The sperm sacs have a 
constricted extremity. The spermiducal glands are long, extending through segments 
17 — 22. The spermathecae are in segments 6 — 8; on one side of the body was an 
additional pouch in segment 9. A second individual was much the same, but had only 
three papillae by each male pore and no traces of an additional spermatheca. In my 
original description of Perichaeta mauritiana I described only two pairs of spermathecae 
in segments 7, 8. I cannot now lay my hands on that specimen. I may however 
observe that a renewed examination of one of the worms which I originally referred to 
barbadensis, seems to have its two pairs of spermathecae in 7, 8, and not as I stated in 
6, 7. In any case the difference does not seem to me to be important. With the 
present species will have to be merged I think Perichaeta cupulifera. There are at 
least no differences of great importance to distinguish that form from Dehra Dun. 
There is to be seen the same kind of range in the variability of the papillae which are 
from as small a number as only one in front of and behind each male pore to twelve or 
so in the neighbourhood of those pores. 

Dr Michaelsen thinks that his Perichaeta pallida is not to be confused with 
Perichaeta hazvayana. He bases this distinction upon the fact that in pallida the 
anterior setae are not much enlarged, as they are in hawayana, and that the male pores 
are more closely approximated. As to the former it would be necessary to separate 
from barbadensis one of the individuals which I have described above as belonging to 
that " species " if this opinion is correct. There is at least quite as much reason for 
uniting this species with the series concerning which the present remarks are offered, 
as for including Rosa's P. amazonica. Rosa says nothing about the increased size of 
the anterior setae. The fact that the clitellum has none will not I hope, after the 
remarks contained in the present paper, be considered as sufficient to discriminate the 
species. 

In Dr Michaelsen's description of Perichaeta mandhorcnsis there are no salient 
points which serve to discriminate it from the present species. It has larger setae on 
segments 2 — 9 : the caeca have the crenated appearance below that is at least often 
found in hawayana. There is one papilla near each male pore ; the three spermathecae 
occupy the same segments ; the spermiducal glands are without the terminal sac. 
The sperm sacs are divided (as in some individuals of the present species) by a 
constriction. There is in short nothing of importance in the description which warrants 
a separation. 



55- 



426 FAUNA HAWAIIENSIS 



(5) Amyntas schmardae Horst. 

Megascolex schmardae Horst, Notes Leyd. Mus. 1883, p. 194. 
Perichaeta trityphla Beddard, P. Zool. Soc. 1896, p. 205. 
Perichaeta vesiculata Goto and Hatai, Annot. Zool. Japon. in. p. 21. 

It is rather curious that in an appendix to my account of the earthworms of the 
Sandwich Islands I should have described from Barbados a species which I regarded 
as new and described as Perichaeta trityphla. Curious since I have subsequently found 
many specimens of this worm in gatherings from Honolulu at 2000 feet of altitude and 
also in earth imported from Hong Kong. I think that Michaelsen' is probably right in 
identifying my trityplila with the long known species schmardae. But at the same time 
it must be borne in mind that the condition of the caeca, which I thought to be distinc- 
tive of trityphla, has not been described in the original specimens of schmardae. 
These caeca, I may say, vary in number from three to six on each side, and the two 
sides are not always symmetrical in this respect. The occurrence of such caeca seems 
to mark the species as a native of Japan — one of the most prominent characteristics of 
the species of that island being the frequent complication of the caeca. I think that 
there can be no doubt that Perichaeta vesiculata of Goto and Hatai is the same species. 
There is positively no feature in their rather short description of the so-called vesiculata, 
which does not fit in with the facts observed in the anatomy of schmardae and trityphla. 

' Oligochaeten von den Inseln des Pacific. Zool. Jahrb. Syst. xu. p. 224. 



427 



ENTOZOA'. "^^^'--.-o- 



By Arthur E. Shipley, Fellow and Tutor of Christ's College, Cambridge, and 
University Lecturer in the Advanced Morphology of the Invertebrata. 



The Hawaiian Archipelago or the Sandwich Islands are separated by some 2350 
miles from the mainland and by about the same distance from any other group of 
inhabited islands. Hence, as might be expected, their fauna is highly specialized, and 
although we know very little about the Entozoa of this island group, two at least of the 
forms described, living within the bodies of birds characteristic of the country, are new 
and up to the present time have not been recorded from elsewhere. 

But although the Hawaiian Archipelago is so far from other lands it is a much 
frequented spot. Since the Spaniards first found it, before the visits in 1778 of Captain 
Cook, it has by degrees become one of the meeting places of the world. Its position 
"at the cross-roads of the North Pacific" on the line of the great marine trade-routes 
between Northern America, Japan, China and Australia has attracted to its harbours 
men of all nations, so that, like Singapore, it has a most mi.xed population. And man 
has brought not only his own parasites with him but has imported his domesticated 
cattle with their entozoa. 

What little I have been able to find out about the human parasites in the Sandwich 
Islands I owe to a paper by Dr A. Lutz^, and as the parasites he mentions are well 
known and almost cosmopolitan I have said little about them, still they should be 
mentioned in a Fauna Hawaiiensis, for though probably the majority of them have been 
introduced and may not be endemic, this is not certainly the case and is susceptible of 
no proof 

' I have much pleasure in thanking Prof. E. Ray Lankester for permission to reprint matter and figures 
from the Quarterly Journal of Microscopical Science ; Mr Shipley has availed himself of this permission in 
his communication. Editor. 

' Centrbl. Bakter. xni. 1893, p. 126. 



428 FAUNA HAWAIIENSIS 

I. NEMATODA. 
The following Nematodes are recorded by Dr Lutz : — 

(i) Ascaris himbricoides L. 

This was found with Ancylostoma duodenale but also — and this was especially the 
case in children — alone. It is said to be very common among the poorer inhabitants. 

(2) Trichocephalus dispar Rud. 

This was determined only from eggs which passed from the intestine. It appeared 
to be widely distributed but not in very great numbers. 

(3) Oxyurus verniicularis L. 

Observed in one white family who had become infected through an adopted child. 
It is a very common parasite among the poor. 

(4) Strongyloides intestmalis Bavery. 

This parasite was observed in company with Ancylostoma duodenale, but in the 
larval state. It apparently produced no special symptoms, and Dr Lutz repeats his 
doubts as to its beins: the cause of the so-called Cochinchina-diarrhoea. 



'& 



(5) Ancylostoma duodenale Dubini. 

The anaemia and accompanying troubles due to the presence of this parasite are 
very common in the Hawaiian Archipelago. Dr Lutz found it only amongst the 
Portuguese, employed for the most part in the sugar plantations and careless about 
their supply of drinking water. These Portuguese came not from Europe but from 
Madeira and the Azores and it seems probable that they brought their parasites with 
them. Dr L. F. Alvarez of the "Hospital for the Treatment of Leprosy" tells me 
that this entozoon is very common amongst the Portuguese labourers of the city of 
Hilo and its neighbourhood in the Island of Hawaii. It produces a very severe form 
of anaemia. 

(6) Sclerostomum armatum Dies. 

This parasite was found in the Horse and at least in one district was the cause of 
the death of the host. 

(7) Filaria iminitis Leidy. 

Found in the heart of dogs. It is widely spread through the Pacific Islands'. 

' Shipley, P. Cambridge Soc. vni. 1892 — 5, p. 211. 



ENTOZOA 429 

II. PLATYHELMINTHES. 

A. Trematoda. 
(i) Distoma clavatuni Rud. 

Found in the stomach of the Coryphaena hippziris. Several other unidentified 
species of Trematode were met with in other fish. 

(2) Distoma hcpaticitni'^ L. 

The presence of this parasite caused a veritable epizootic amongst the cattle on 
many of the islands of the Hawaiian Archipelago. Horses, though to a less extent, 
and wild swine are also said to have suffered. Dr Lutz was successful in cultivatino- 
the embryos of the Fluke, and he also succeeded in finding and observing the develop- 
ment of the Redia in the fresh-water snail Limnaeus pereger. In his second paper' 
Dr Lutz gives the opinions of certain distinguished conchologists on the species of water 
snails which he found infected with the larvae of Distoma. These Molluscs seem to 
afford matter for a considerable amount of disagreement and the reader is referred to 
this paper for the details of the subject. Dr Alvarez, to whose kindness I am indebted 
for several details in this paper, tells me that this Fluke sometimes attacks man. 

B. Cestoda. 
(i) Drtpanidotacjiia hemignathi'- Shipley. 

Plate XIII. 

The specimens of this tapeworm, of which I received but ten, are all small ; they 
vary in length from 10 mm. to 22 mm. The head is very small ; immediately behind 
it, there being practically no neck, the body begins to broaden out, and in some 
specimens the proglottides attain a width of 2 mm. The segmentation of the body 
commences immediately behind the head, and is very well marked a little further back. 
The posterior border of each segment overlaps the succeeding one with a prominent 
edge or rim ; this is well shown in longitudinal section (fig. 6). The number of 
segments varies from some fifty to sixty to over a hundred. The measurements given 
above are about the average, but, as is well known, tapeworms are extremely extensible 
animals, and this to a great extent diminishes the value of figures quoted in reference to 
their size. In some of my specimens the body is stretched, and the length of the seg- 
ments equals one-half or even two-thirds of their breadth, but in the commoner forms 
the segments are very short and broad, sometimes eight or ten times as broad as long. 
They are flattened, as is seen in transverse section, and sometimes, especially towards 

' Centrbl. Bakter. ix. 1892, p. 783, and xiii. 1893, p. 320. 

'- The description of this species is reprinted (with certain aherations) from the Quart. J. Micr. Sci. XL. 
1898, p. 613. 



430 FAUNA HAWAIIENSIS 

the posterior end, the whole body is hollowed so that each segment is curved. The 
most posterior segments, which are crowded with embryos well advanced in their 
development, are rounder, less flattened, longer, and they readily broke off. 

I was not able to detect any genital pore on the exterior even with the aid of 
powerful lenses, but sections (figs. 4 and 6) and stained mounted specimens show that 
it is on the same side of the body in all the segments. 

The head of the tapeworm bears four suckers, and in the midst of them is the 
rostellum (figs. 3, 8 and 9). The shape of the head is very various : in some cases the 
suckers are, as it were, hunched up and lie at each corner of a square, the lateral 
diameter of which does not exceed the dorso-ventral (fig. 8) ; in other specimens the 
head is not separated from the body by a deep constriction, but is flattened and spread 
out (fig. 7), so that the lateral suckers are separated from one another by a space con- 
siderably wider than that which lies between the dorsal and the ventral suckers. 

The rostellum is minute and sunk in a pit (fig. 3) ; it bears a wreath of ten hooks. 
In all the specimens which I cut into sections, and I think in the others as well, the 
rostellum was retracted, the points of the hooks folded in against the a.xis of the 
rostellum, and not reaching so far forward as the mouth of the pit. When the animal is 
fixed to the mucous membrane of its host this rostellum is doubtless protruded from its 
sheath, and the hooks are divaricated. Certain muscle-fibres which run from the base 
of the rostellum, and lose themselves in the parenchyma, probably serve to retract it. 

The hooks are slightly curved, and the projection which corresponds with the inner 
fork of the more triradiate hooks of other genera is hardly, if at all, marked (fig. 2). 
Measuring in a straight line from the base to the tip the hooks are 18 — 23/^ in 
length, thus corresponding pretty closely with those of Drepanidotaenia tenuirostris 
which, according to Railliet', measure 20 to 23 ^a, and to those of D. lanceolata, which 
measure 25 to 31 /a. 

The four suckers present no peculiarities ; they are deeply cupped, with a small 
orifice to their lumen, but probably they are capable of considerable change of form 
(fig. 9). They are probably retracted by some muscle-fibres which cross one another 
and run into the parenchyma. 

The segmentation of the body begins immediately behind the suckers ; at first the 
segments are very short, but they gradually increase in size throughout the first three- 
quarters of the length of the body. For the last quarter the segments are crowded 
with embryos ; they become in this region much narrower, more cylindrical in shape, 
and longer, and are very easily broken off. The posterior free edge of the segments of 
the anterior two-thirds of the body is sharp, and may overlap the segment behind, or 
may stand out clearly from it. 

The water-vascular system is well developed; on each side of the body are two 
longitudinal canals, — one, the ventral, much bigger than the other, or dorsal. The 
' Traite de Zoologie medicale et agricole, Paris, 1895. 



ENTOZOA 431 

lining of the former seems to be a structureless cuticle with no cells especially related to 
it, but the wall of the dorsal vessel is surrounded by a number of small deeply stained 
cells (fig. 4). I did not see any communication between the vessels of one side, but 
the larger vessels communicate as usual, one with another, by a transverse vessel 
running from side to side along the posterior border of each segment. In the head the 
vessels all communicate. In some of the better preserved sections such structures as 
are depicted in fig. 10 were seen : these may or may not be flame-cells ; they look 
rather like them. No valves were seen in the course of the vessels. 

The lateral nerve-cords are well marked, lying externally to the ventral excretory 
canals ; they fuse together in the head, forming a ganglion which is indicated in fio-. 3. 
No traces of the nerve-ring described by Tower' as running round the posterior end of 
each segment of Moniezia, or of the secondary nerves described by the same observer, 
were to be seen. But these, if present, probably require fresh material and special 
methods of preservation to make them manifest. Special nerve-cells, described below, 
are scattered through the parenchyma of the body. 

The histology — at least in some specimens — could be fairly well made out, and 
agrees roughly with what Blochmann has described in Ligtila monogramma' . The 
whole body is covered by a cuticle, the outer fifth of which stains more deeply than the 
remainder. Within this, with a high power, a number of dots or knobs become visible 
(fig. 10). These are the swollen terminations of certain strands or processes of the 
ectoderm cells. The cells themselves, as Blochmann has shown, lie removed to some 
distance from the cuticle they secrete, but are in contact with it by means of the above- 
mentioned processes ending in the knobs. 

The ectoderm cells are not all at one level, but on the whole form a fairly well- 
marked layer. Each cell is fusiform in shape, and produced into two or three processes, 
which project both peripherally and centrally. They contain large and well-marked 
nuclei. Neither the cells nor their processes are laterally in contact ; they are separated 
one from another to varying extents by the intrusion of some of the parenchymatous 
network which makes up so much of the body of a Cestode. 

This parenchyma consists of a meshwork which permeates everywhere the body of 
the tapeworm, surrounding all the organs, and often, as is the case with the ectoderm 
and the muscles, passing in between their constituent cells. In the spaces of the mesh- 
work there is believed to be a fluid. The meshwork itself is secreted and nourished by 
certain large star-shaped cells which are irregularly scattered through the parenchyma, 
and which give off processes in all directions (fig. 10). 

Round the generative glands this parenchymatous network becomes condensed, 
the spaces disappear, and it forms a close sheath to the ovary, testis, &c. At the 
posterior end of each segment it is also somewhat condensed, and in section presents 

' Zool. Anz. vol. XIX. 1896, p. 323. 

' Die Epithelfrage bei Cestoden und Trematoden, Hamburg, 1896. 

F. H. II. 56 



432 FAUNA HAWAIIENSIS 

the appearance of a well-marked double line, which is very characteristic, and is well 
shown in fig. 6. 

Scattered amongst the parenchyma are certain faintly stained cells which seem to 
be bipolar, and which differ from the cells of the parenchyma both in shape and in their 
powers of absorbing the staining reagents. These I take to be nerve-cells which are 
in communication with the nerve-fibres of the lateral cords. The latter are entirely 
devoid of any nerve-cells on their course. 

Muscle-fibres are scattered through the substance of the body, and one set of 
longitudinal muscles are most definitely arranged. This layer is situated just below the 
epidermis in the anterior part of the segment, but as the latter increases in size 
posteriorly, the cylinder of muscle-fibres, which retains the same diameter throughout, 
comes to lie more deeply in the tissues. These muscles, like the nervous system and 
excretory canals, run from segment to segment ; some of them, if not all, end in the 
cuticle, where it is most bent in at the posterior end of each segment. Laterally the 
fibres are not in contact, being separated by considerable intervals. Their regular 
arrangement is shown in fig. 5. 

In the posterior segments, which are so ripe that the slightest touch breaks them 
off, the parenchyma has undergone considerable degeneration, the cells are less clear, and 
the spaces of the meshwork are larger and more irregular. 

The generative organs begin to arise very early in the series of segments. Already 
in the eighth or tenth segment clusters of cells are segregating, and their deep staining 
shows that they belong to the gonads. In the se.xually ripe segments the ovary is 
centrally placed, and is supported on each side by a lobe of the testis. From the latter 
a fine vas deferens leads into an extensive vesicula seminalis, which is as a rule crowded 
with spermatozoa ; from this a muscular duct leads to the unilateral genital pore. I 
was unable to make out the details of the penis, and similarly I failed to detect any 
yolk-gland amongst the female genitalia. 

The vagina leads at once into a large receptaculum seminis, whose walls were 
strengthened by a series of cuticular-looking rings, whose cut ends are shown in 
figs. 4 and 6. This communicates both with the oviduct and with the uterus. The 
latter presents no special points of interest ; in the posterior segments it contains the 
typical three-hooked larvae, each segment containing at least one hundred and probably 
more. 

Classification. 

In his paper on taenias in birds, Dr Fuhrmann' remarks that of the 240 odd 
species of tapeworm described from avian hosts, only twenty-one have been studied 
anatomically ; the remainder are but little more than names, and probably many of the 
names are of doubtful validity. 

' Rev. Suisse Zool. tome in. 1895 — 6, p. 433. 



ENTOZOA 433 

A certain amount of order has been introduced into this mass of material by the 
establishment of certain sub-groups, and by the giving of a new generic name to the 
members of these subdivisions; thus in 1891 Blanchard and Railliet' established the 
genus Davainea ; in 1892 Railliet" suggested two new generic names, Drepanidotaenia 
and Dicranotaenia, for certain tapeworms inhabiting, for the most part, domestic birds. 
These are characterised chiefly by the nature of the hooks. In the following year 
Diamare' founded the genus Cotiignia, in which the generative organs are double and 
have two pores, but which is distinct from the genus Dipylidium of Leuckart. All 
these genera are characteristic avian tapeworms, and are, with but very few excep- 
tions, confined to birds. 

There is little doubt that the tapeworm which I have described above from the 
intestine of Heviignatlms proceriis corresponds with a Drepanidotaenia of Railliet^ who 
defines his genus as follows : 

" Tapeworms provided with a simple crown of uniform hooks, which are usually 
few in number ; the outer limb (manche) of the forked base of the hooks is much longer 
than the inner (garde), which is always slight ; the point is directed backwards when 
the rostrum is withdrawn. The majority live in the intestines of aquatic birds. Their 
larva is a Cysticercoid, and is found encysted in the bodies of small fresh-water 
Crustacea." 

Railliet describes eight species of Drepanidotaenia ; in one of these the genital 
pores are on alternate sides of the body in successive segments ; the remaining seven 
species are unilateral in this respect, but they fall into two groups, — one, with three 
species, in which the number of hooks is eight ; and the other, with four species, in 
which the number of hooks is ten. 

It is to this latter group that we must add the tapeworm from H. procerus. The 
four species D. anatina, D. sinaosa, D. setigera, and D. tennirostris differ inter se in 
several respects, but perhaps the simplest way of determining the species is by measuring 
their hooks. Of these four species, D. hemignathi most nearly resembles D. tenni- 
rostris, which occurs in certain of the ducks ; it differs, however, markedly in size, being 
hen mature about 4 to -Ar the length of the last named. It resembles D. tennirostris 



w 



5 '-'-' T2- "-'ic '^"g 



in the length of its hooks in the head, which in the latter are 20 — 23 /a, in the former 

are 18 to 23 /x ; but whereas the hooks of the embryo are about the same length in the 

new species, i.e. about 20 /^, in D. tennirostris they are but 7 /a. The neck is short, not 

long as in the last-named species, and the eggs are small, about 40 — 50 ^u, in diameter, 

and spherical in shape, not cylindrical as Krabbe' figures them, with a length of 85 /x. 

The hooks also differ in shape ; those of D. tennirostris have a much more strongly 

' Mem. Soc. Zool. France, tome iv. 1891, p. 420. 

" Ibid, tome xvn. 1892, p. 115. 

' Boll. Soc. Napoli, ser. i, vol. vii. 1893, p. 9. 

* Traite de Zoologie medicale et agricole, Paris, 1895, p. 298. 

^ Danske Selslc. Skr. vni. 1870, p. 249. 

56—2 



434 FAUNA HAWAIIENSIS 

developed process corresponding with the inner limb of the forked base than occurs in 
D. hemignathi. 

The species, which I named after its host, may be characterised as follows : 

(i) Drepanidotaenia he7iiignathi Shipley. 

D. hemignatJii Shipley, Quart. J. Micr. Sci. xl. p. 620. 

Length i — 2'2 centimetres; breadth, in the middle of the body, 2 millimetres. 
Head flattened and compressed, rostrum with a crown of ten hooks ; each hook 
18 — 23 /LA in length, and with but a slight trace of the inner limb of the forked base. 
Neck short. The first segments are short, but they very soon (eighth or tenth) show 
traces of reproductive organs. Genital pore unilateral. The posterior limit of each 
segment is sharply defined, and forms an angle of about 45 degrees with the sides. 
Egg spherical, diameter about 40 — 50 ju,. The three pairs of embryonic hooks measure 
about 20 /u, each in length. 

Hab. Hemignathus procerus. Sandwich Islands : in the intestine. 

(2) Mr Perkins has also given me two or three specimens of a tapeworm from 
a Loxops, sp. This bird, like the Hoiiignatlms, is a member of the family Dre- 
panididae, which is confined to the Sandwich Islands. Unfortunately the specimens 
are without their head, and I am unable to identify them. They differ markedly 
from the Drepanidotaenia described above. 

(3) Echinococcits ?. 

Echinococcus is mentioned by Dr Lutz as occurring occasionally amongst cattle 
killed for the market. 

(4) Taenia crassicollis Rud. 
This cysticercus larva of this species was found by Dr Lutz in Mus decumajius. 

(5) Taenia solium L. 
The entozoon is said to be very uncommon, but is occasionally met with. 

III. ACANTHOCEPHALA. 
(i) Apororhynchus heinignathi^ Shipley. 

In the summer of 1894 I received from Mr Perkins seven small parasites which he 
had noticed adhering lightly to the skin around the anus, but beneath the skin, of a 
species of bird, Hemignathus procerus, which he collected in the island of Kauai. Each 

Quart. J. Micr. Sci. xxxix. p. 207 and xlii. p. 361. 



ENTOZOA 435 

of these parasites was divided into three regions,— a head, a collar, and a trunk ; and, in 
fact, they have an almost ludicrous resemblance to a young Balanoglossus with one or 
two gill- slits (figs. II, 12, and 13). On investigating their anatomy it at once became 
evident that the animals belonged to the group Acanthocephala, and, further, that they 
differed from the other members of the group in the absence of what is perhaps their 
most characteristic organ,— from which, indeed, they take their name — the hooked 
proboscis or introvert. Careful inspection failed to reveal any trace of a scar or mark 
where the introvert might have been broken off; and although in the absence of hooks 
and introvert sheath, &c., the anterior part of the body which I have called the head is 
as unlike the typical introvert as possible, still in its relation to the lemnisci and to the 
ligament it occupies the position of that organ, and until we can get further information 
I think the best plan is to regard this part of the body as equivalent to the eversible 
part of more normal forms. 

The second of the three regions into which the body is externally divided is shorter 
than the head and smaller in diameter ; it may be termed the collar. The third or 
posterior region, which may be called the trunk, is the longest and the most slender of 
the three ; behind it tapers to a point where the orifice of the genital duct is situated, 
and this end of the animal is always a little turned up (figs. 11, 12, 13, 17 and 23). The 
exterior of the collar and trunk are smooth or lightly wrinkled, but the head is covered 
with a number of small depressions or pits which give it a very characteristic appear- 
ance, and which are well seen in sections. The head is attached to the collar by a 
narrow neck, which is surrounded and concealed by the edge of the collar. This is 
obvious in sections (figs. 15 and 23). All the specimens were somewhat shrivelled and 
apparently distorted. The largest measured 3 '5 mm. in length, the smallest 2-5 mm. ; 
had they been fully distended they would probably have been i to i '5 mm. longer. 
The body-cavity of the head is continuous with that of the neck, and the latter opens 
freely into the cavity of the trunk (fig. 23). The first-named space is by far the largest. 
The lumen of the collar region is reduced by the great thickness of the walls of this 
part of the body, and both here and in the trunk much of the internal space is occupied 
by the lemnisci and the reproductive organs. 

The skin is one of the most characteristic features of the Acanthocephala, and as 
far as I know is only paralleled by that of the Nematodes, but it possesses certain 
features not found in the last-named group. The whole body is covered by a thin 
cuticle which does not vary much in thickness in the different regions of the body, and 
which is invaginated a short distance into the genital pore. Beneath this is the true 
epidermis, or subcuticle as it is called; this has in my specimens the usual structure met 
with in the group so well described by Hamann, and consists of a matrix of a fibrillar 
nature, the fibrils being as a rule arranged radially, in which are embedded a certain 
number of amoeboid nuclei (figs. 16 and 20). This tissue is much thicker in the region 
of the collar than elsewhere, and it is thicker in the trunk than in the head. It is pierced 



436 FAUNA HAWAIIENSIS 

in all directions by a series of tubes or lacunae which have no definite lining, but which 
seem to be mere splits in the fibrillar matrix. The lacunae — except in the head — have a 
general circular direction which is very well marked in the trunk region where each runs 
into a lateral longitudinal split (figs. 20 and 24). They contain a small amount of coagu- 
lum, the remnant of the fluid which circulates in them ; during life this fluid, in other species, 
holds in suspension fat and coloured oil globules. If these are present in my species 
they must have been dissolved out in the processes which precede embedding. The 
circular lacunae of the trunk not only communicate with one another by means of the two 
longitudinal lateral lacunae (figs. 20 and 24), but they open into one another by numerous 
small branches which have an oblique or longitudinal direction. In the head the lacunae 
have a general longitudinal course ; they are not, however, straight, but twist in and out 
between the pits on the surface ; they anastomose freely (fig. 14). Thus in a transverse 
section of the head the lacunae appear as round holes more or less uniformly arranged 
in the skin, and the same effect is produced by a longitudinal section of the trunk. 

In the collar region the subcuticular tissue is much thickened, and the lacunar 
system forms a single more or less definite ring which gives off numerous branching 
anastomosing twigs (fig. 15). 

Although the above account attempts to give the general course of the lacunae in 
the skin, it should be mentioned that there is considerable irregularity in the arrange- 
ment, and one is almost inclined to believe that the canals do not remain permanent, 
but that they sometimes close up and new ones appear. As they have no lining of any 
kind, such a closing would leave no trace. 

As Schneider^ Hamann", and Kaiser' have shown in the species investigated by 
them, the lacunar system of the introvert is completely shut off from that of the neck — 
if it be present — and of the trunk, by a fold inwards of the cuticle which cuts the 
subcuticular tissue in two. I have not been able to find any such cuticular ring in the 
species in question, but the state of preservation of my specimens does not allow me 
to say definitely that it does not exist. 

The lemnisci are two elongated sac-like prolongations of the subcuticular tissue 
which are attached anteriorly to the skin at the junction of the head and collar. They 
extend backwards to the extreme posterior end of the body, and are slightly bent so 
that a longitudinal section may cut them in two or three places (fig. 23). Histologically 
they are composed of the same substance as the subcuticle in direct continuity with 
which they arise, and they are traversed by a similar system of canals. Physiologically 
they seem, as Hamann suggests, to act as reservoirs for the fluid of the canal system of 
the introvert ; when the fluid they contain is forced into the spaces of the introvert the 
latter is everted. It is withdrawn again into the body by special muscles. In most 

' Arch. Anat. 1868, p. 584. 

° Die Nemathelminthen, Heft i and 2, Jena, 1891 and 1895. 

' Bibl. Zool. Heft 7, 1892, p. i. 



ENTOZOA 



437 



species the canal system of the lemnisci opens into that of the introvert in front of the 
cuticular ring, and is thus completely independent of that of the trunk. If we assume 
that the head of my species corresponds with the introvert of other forms which have 
lost its introvert sheath, the lemnisci open into the same region of the skin as they do 
in other Acanthocephala. 

The nuclei of the subcuticle and of the lemnisci are very remarkable ; they corre- 
spond in structure with those described by Hamann in Neorhynclms clavaeceps, in which 
species according to this observer both the skin and the lemnisci retain in the adult 
their embryonic condition. As in Neorhynclms the number of nuclei is very small, some 
twelve to twenty seem to suffice for the whole of the subcuticle, and perhaps two to four 
for each lemniscus. The structure of the nucleus shows a most striking- resemblance to 
an amoeba with rather short pseudopodia (figs. i6, 20, and 23). No single nucleolus 
can be detected, but numerous chromatin particles are present, and in some a distinct 
vacuole can be observed. These nuclei are scattered about in a most irregular fashion ; 
not one may be seen in a number of consecutive sections, and then perhaps three or 
four may appear, and from their large size persist through several sections. The nuclei 
lie, as a rule, embedded in the substance of the subcuticle ; more rarely they are found 
in the lacunae. Although there is no proof, one is tempted to believe that the nuclei 
wander through the subcuticle and lemnisci in an amoeboid manner, and that the small 
number of nuclei which are found in these tissues is compensated for partly by the large 
size of each, but more especially by their mobility. Similar amoeboid nuclei undoubtedly 
move about, fuse with one another, and undergo fission in the subcuticle of the larval 
forms of Neorhyuchus clavaeceps. 

Within the subcuticle and completing the skin on the inner side, is a layer of 
circular muscles, and still more internally a layer of longitudinal muscles (figs. 16 and 
25). The muscles of these layers are but a single fibre thick, and they are not very 
uniformly present. The circular layer is most complete in the region of the trunk, and 
I have figured a section to show this (fig. 22). The longitudinal layer is even less 
definite, but scattered fibres can be detected here and there (figs. 16 and 25). Each 
fibre appears to be spindle-shaped, and in the circular muscles has the striated portion 
only on its outer face, forming a thin band ; the inner half of the fibre consists of vacuo- 
lated strands of protoplasm in which is a nucleus. The longitudinal layer of muscles 
alone is continued over the lemnisci (figs. 19 and 24). These muscles are not covered 
on their inner side by any layer of epithelial cells, neither does any such layer cover the 
ligament, but both tissues lie freely exposed to the fluid of the body-cavity. 

In the more typical Acanthocephala the anterior end of the body terminates in a 
hollow eversible portion provided with rows of hooks whose number and shape have a 
certain systematic value. This introvert can be withdrawn, not into the general body- 
cavity, but into the cavity of the introvert sheath, which is shut off from the general 
body-cavity by a double (Echinorhynchidae) or a single (Neorhynchidae) wall. The 



438 FAUNA HAWAIIENSIS 

extrusion of the introvert is believed to be effected by fluid being forced into its lacunae 
by the lemnisci. It is retracted by special muscles attached to the inside of its tip ; 
besides these, other retractor muscles run from the outside of the introvert sheath, and 
these serve to retract the whole sheath and its contents into the trunk. The chief nerve 
ganglion lies as a rule on the posterior end of the introvert sheath, usually in the middle 
line, but in the Gigantorhynchidae it is placed to one side. From the posterior end of 
the introvert sheath, and having its origin between its two walls when they are present, 
the ligament runs backward, traversing the body cavity, and ending in the funnel-shaped 
internal opening of the oviduct in the female and in the vas deferens in the male. 

Owing to the absence of an introvert and its sheath, the relations of the ligament 
in the present species is somewhat altered. It takes its origin from the anterior end of 
the head, and at first seems to consist of a few strands of muscular fibres which arise 
from the muscles of the skin (fig. 21). All my specimens but one proved to be mature 
females, whose ovaries had broken up into the &v^ masses which are characteristic of 
the Acanthocephala. These egg masses consist of packets of a dozen or more cells of 
which the peripheral layer develop into ova at the cost of the central cells which serve 
them as a food supply (figs. 14, 16, and 23). These packets coexisted in my specimens 
with ova in various stages of development, some without any egg shell, whilst others 
were provided with a thick deeply-staining membrane. The whole lumen of the head 
was crowded with these ova. In the region of the collar the ova were confined by a 
thin-walled membrane, and in the trunk there were two such masses of ova, which, 
however, seemed less mature than those lying in the head. Lying amongst the 
various organs in the body-cavity were a number of very finely granular masses, which 
I take to be the masses of spermatozoa (figs. 16 and 20). Of the complex system 
by means of which the ova leave the body, little could be made out beyond the fact 
that a well-marked funnel is present opening into the posterior end of the body- 
cavity of the trunk (fig. 19). I failed, however, to find a second opening near the 
narrow end of the funnel such as occurs in other forms, but this may have been 
due to the poor state of preservation. The funnel leads into a duct which opens on 
the posterior end of the trunk. 

The testes are two in number, and lie one behind the other in the ligament, 
though owing to its looping both may appear in the same transverse section. The 
spermatozoa do not escape into the body of the male as the ova do into that of 
the female, but pass down a duct in the ligament which opens at the end of the 
body. Traces of accessory glands were seen, but the details were not clear. 

The brain lies on or in the ligament just behind its point of attachment to the 
skin of the head (figs. 21 and 23). Owing to the disruption of the ovaries in my female 
specimens the ligament could not be traced very far, but in the only male it reached 
from one end of the body to the other. The brain consists of a few large ganglion 
cells with a clear homogeneous cytoplasm and deeply-stained nuclei ; the divisions 



ENTOZOA 439 

between the cells were very sharp and straight (fig. 21). In the females this mass 
of cells lay on the ligament ; in the male, on the other hand, it occupied the centre 
of the fibrous and muscular strands which compose that body (fig. 25). In the former 
I could trace no nerves leaving the brain, but in the male two nerves surrounded 
by muscles pass backward ; these obviously correspond with the retinacula of other 
forms. 

Classification. 

Until recently the group Acanthocephala included but one genus, Echinorhynchus, 
which comprised several hundred species. Recently, however, Hamann' has pointed 
out that these species present certain differences which enable him to divide the group 
into three families, each with a corresponding genus. To these I venture to add a 
fourth, to include the remarkable form above described. This family may, I think, 
be called the Apororhynchidae, and the new genus Apororhynchus", which name refers 
to the absence of the eversible introvert ; and, inasmuch as it is convenient in naming 
a parasite to have some indication of its host, I think the specific name may be 
Jieniignathi. 

If these terms be adopted, the classification of the Acanthocephala will be as 
follows, the characteristics of each of the first three families being taken from 
Hamann's papers. 

I. Family Echinorhynchidae. The body is elongated and smooth. The in- 
trovert sheath has double walls, and the introvert is invaginated into it. The nerve 
ganglion is in the introvert sheath, mostly embedded in it and central in position. 
The hook papillae are only covered with chitin at their apex, and the hooks have 
a process below. 

Genus EcJiinorJiynclnis, with the characters of the family. 

The vast majority of Acanthocephala belong to this family ; a few may be 
mentioned. E. proteus, found in many fishes and varying in size with its host ; its 
larval forms inhabit the Amphipod Gammariis pulex, and are also found in the body- 
cavity of numerous fresh-water fishes. E. clavula occurs in many fishes and in the 
intestine of a species of Bufo. E. angustahis is found also in fishes, with its larval 
form in the Isopod Asellus aqtiatints. E. moniliformis is said to attain maturity 
in the human intestine ; its usual host is a mouse, and its larval host is the larva of 
a beetle, Blaps mucronata. E. porrigens invests the intestine of the rorqual, and 
E. strumosus that of a seal. There are many others. 

' Loc. cit. and Zool. Anz. Bd. xv. 1892, p. 195. 

' In my original paper I suggested the name Arhynchus, but as Professor C. Wardell Styles and Professor 
A. Hassall have pointed out that this name is preoccupied, having been used by Dejean in 1834 for a beetle, 
I later (Quart. J. Micr. Sci. xlh. p. 361) suggested the name Apororhynchus. 

F. H. II. 57 



440 FAUNA HAWAIIENSIS 

II. Family Gigantorhynchidae. Large forms, whose body is ringed and 
flattened during life like that of a Taenia. The hooks are like those of a Taenia, 
the hook-papilla being entirely covered with chitin. There are two root-like processes 
in each hook. The introvert is muscular, has no lumen, and the introvert cannot be 
retracted into it, but the whole retracts into the body-cavity. The ganglion is excentri- 
cally placed to the side, behind the middle of the so-called sheath. The body-cavity is 
enclosed in a structureless membrane, and is traversed by membranes stretched trans- 
versely. The lemnisci are long, coiled, with a central lacuna. 

Genus Gigantorhynckus, with the characters of the family. 

Hamann includes three species in this family — G. echinodiscus, G. taenioides, and 
G. spira ; and points out that E. gigas agrees with them in all points but that of the 
external annulation. The first of the above-named species occurs in the intestine of 
anteaters, and has been found in Myrmecopliaga jiibata and Cycloturus didactyla. 
G. taenioides has been found in a species of Cariania, DicJiolophus cristatus ; and 
G. spira lives in the king vulture, Sarcorhampus papa. E. gigas in the adult stage 
occurs in the small intestine of swine, and its larval host is believed to be the grubs 
of MelolontJia vulgaris and Cetonia aurata in Europe and of Lachnosterna arcuata 
in the United States \ It is recorded once from the human intestine. 

III. Family Neorhynchidae. Sexual maturity is reached in the larval state. 
The introvert sheath has a single wall. A few giant nuclei only are found in the 
subcuticle and in the lemnisci. The circular muscles are very simply developed, and 
the longitudinal muscles only present in places. 

Genus Neorhynchus, with the characters of the family. 

This genus includes but two species, A^. clavaeceps and A^. agilis. They both 
present interesting cases of paedogenesis, the large embryonic nuclei of the young 
larva do not break up into numerous nuclei as they do in the commoner species. 
A", agilis is found in Mugil auratus and M. cephalus ; A^. clavaeceps in the Carp, 
Cyprinus carpio, its larva form according to Villot" in the fat bodies of the Neuro- 
pterous insect Sialis niger ; it has also been found in the alimentary canal of the 
leech NepJielis octocula, and specimens of the water-snail Linmaea have been artificially 
infected with it. 

IV. Family Apororhynchidae. Short forms, with the body divided into three 
well-marked regions, — head, collar, and trunk. The head is pitted, the collar smooth, 
and the trunk wrinkled, not annulated — in spirit specimens. There is no eversible 
introvert, and no introvert sheath, and no hooks. The sub-cuticle and the lemnisci 
have a few giant nuclei, and the lemnisci are long and coiled. 

' C. W. Styles, Zool. Anz. xv. 1892, p. 52. 
' Zool. Anz. VIII. 1885, p. ig. 



ENTOZOA 



441 



Genus Apororhynchus , with the characters of the family. 

This family in the length and curvature of its lemnisci resembles the Giganto- 
rhynchidae, and in the persistence of the embryonic condition of the nuclei in the 
sub-cuticle and the lemnisci, the Neorhynchidae ; but in the shape of the body, its 
division into three well-marked regions, the absence of eversible introvert, introvert 
sheath, and hooks, it stands alone, though to some extent nearer to the Neorhyn- 
chidae, in which the introvert is relatively small, the introvert sheath simple, and 
the number of hooks reduced, than to either of the other families. 

The single species Apororhynchtis hemignathi was found attached to the inner side 
of the skin in the neighbourhood of the anus of a Sandwich Island bird, Hemignathus 
procerus. This bird is a member of a family Drepanididae, which is entirely confined 
to the Sandwich Island group. Professor Newton tells me that it is probable that 
the "food of Hemignatlms consists entirely of insects which it finds in or under the 
bark of trees " ; hence it is probable that the second host of this parasite, if such 
exists, must be looked for amongst the Insecta. 

(2) Echiiior/iyjichus caiupajiulaius Dies. 

Found by Dr Lutz in Water-rats. This species is said to be a facultative 
parasite of man. 

The Zoological Laboratory, Cambridge. 
March, 1 900. 



ZOOLOGICAL RESULTS based on material from New Britain, New Guinea, Loyalty 

Islands and elsewhere, collected during the years 1895, 1896, and 1897, by 

Arthur Willey, D.Sc. Lond., Hon. M.A. Cantab., Late Balfour Student of the 

University of Cambridge. Demy 4to. Parts I, II, and III. Price 125. 6d. each. 

PART I. 1898. 

1. The anatomy and development of Peripahts novae-britanniae. By ARTHUR WiLLEY, M.A., D.Sc. 

With Plates I.— IV. and 7 figures in the text. 

2. Metaprotella sandaknsis, n. sp. [Caprellidae]. By Dr PAUL MAYER. 

With 6 figures in the text. 

3. On a little-known Sea-snake from the South Pacific. By G. A. BoULENGER, F.R.S. 

With Plate V. 

4. Report on the Centipedes and Millipedes. By R. I. POCOCK. 

With Plate VI. 

5. Account of the Phasmidae with notes on the eggs. By D. SHARP, M.A., F.R.S. 

With Plates VII.— IX. 

6. Scorpions, Pedipalpi and Spiders. By R. I. PocoCK. 

With Plates X.— XI. 

PART II. 1899. 

Report on the specimens of the genus Millepora. By SYDNEY J. HiCKSON, M.A., D.Sc, F.R.S. 

With Plates XII.— XVI. 
Report on the Echinoderms (other than Holothurians). By F. JEFFREY BELL, M.A. 

With figures on Plate XVII. and one figure in the text. 
Holothurians. By F. P. BEDFORD, B.A. 

With figures on Plate XVII. 

10. Report on the Sipunculoidea. By ARTHUR E. SHIPLEY, M.A. 

With Plate XVIII. 

11. On the Solitary Corals. By J. STANLEY GARDINER, M.A. ' 

With figures on Plates XIX. and XX. 

12. On the postembryonic development of Cycloseris. By J. STANLEY GARDINER, M.A. 

With figures on Plates XIX. and XX. 

13. On a collection of Earthworms. By Frank E. Beddard, M.A., F.R.S. 

With Plate XXI. 

14. The Gorgonacea. By Isa L. Hiles, B.Sc. 

With Plates X.XII. and XXIIl. 

PART III. 1899. 

15. Orthogenetic variation in the shells of Chelonia. By Hans Gadow, M.A., Ph.D., F.R.S. 

With Plates XXIV. — XXV. and one text-figure. 

16. Enteropneusta from the South Pacific, with notes on the West Indian Species. By ARTHUR 

Willey, D.Sc, Hon. M.A. Cantab. 

With Plates XXVI.— XXXII. and 7 text-figures. 

17. On a collection of Echiurids from the Loyalty Islands, New Britain and China Straits, with an attempt 

to revise the group and to determine its geographical range. By ARTHUR E. SHIPLEY, M.A. 
With Plate XXXIII. 

PART IV. 1900. 

18. On the anatomy of a supposed new species of Cooiopsammia from Lifu. By J. STANLEY 

Gardiner, M.A., Fellow of Gonville and Caius College, Cambridge. With one plate. 

19. On the Insects from New Britain. By D. Sharp, M.A., M.B., F.R.S. With one plate. 

20. Report on the Stomatopoda and Macrura. By L. A. BORRADAILE, M.A., Lecturer of Selwyn 

College, Cambridge. With four plates. 

21. Report on the Slugs. By WALTER E. COLLINGE, F.Z.S., Mason College, Birmingham. 

With two plates. 

22. Report on the Polyzoa. By E. G. Philipps, Newnham College, Cambridge. With two plates. 

23. The Hydroid Zoophytes. By LAURA ROSCOE Thornely, University College, Liverpool. 

With one plate. 

24. Astrosclei'a tvilleyatia, the Type of a new Family of Calcareous Sponges. By J. J. LISTER, M.A., 

St John's College, Cambridge. With five plates. 

25. A contribution towards our knowledge of the Pterylography of the Megapodii. By W. P. 

Pycraft, A.L.S., British Museum. With one plate. 

26. The Stolonifera and Alcyonacea. By Sydney J. Hickson, M.A., D.Sc, F.R.S., Beyer Professor 

of Zoology in the Owens College, and ISA L. HiLES, B.Sc, Owens College, Manchester. 
With two plates. 

27. Report on the Xeniidae. By J. H. AsHWORTH, B.Sc, Demonstrator in Zoology, Owens College, 

Manchester. With two plates. 

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FAUNA HAWAIIENSIS 



OR THE 



ZOOLOGY OF THE SANDWICH (HAWAHAN) ISLES: 

Being Results of the Explorations instituted by the Joint Committee 

appointed by 

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443 



ARACHNIDA. ^I'tt 



^«Cl^•- 



""Pon, 



Par Eugene Simon. 

§ 1. General considerations on the Arachnida. 

Les patientes recherches de Mr R. C. L. Perkins nous ont fait connaitre loi 
especes d'Arachnides superieurs' des iles Sandwich, dont yy speciales, proportion 
qui ne s'observe nulle part ailleurs. 

La faune de cet archipel peut done etre consideree comme fortement specialisee 
et riche relativement au pen d'etendue de son territoire ; elle se rattache d'une maniere 
generale aux faunes Malaise et Australienne, avec I'absence de certaines formes 
tropicales tres repandues telles que Gasteracantha, Nephila etc., et par contre I'ad- 
jonction de certaines formes palearctiques et nearctiques telles que Dysdei-a, Labulla etc. 

Les divers groupes d'Arachnides y sont tres inegalement distribues : sur une 
quarantaine de families que Ton admet generalement aujourd'hui dans I'ordre des 
Araneae, 28 font completement defaut, et trois {Sicariides, Clubionides et Agclenides) 
n'y sont representees chacune que par une seule espece ubiquiste probablement 
introduite. II en est de meme de I'ordre des Scorpiones dont la seule espece, Iso- 
fneirus macttlatus de Geer, n'est certainement pas autochtone ; les ordres des 
Pedipalpi et des Soltfitgae font defaut. 

Les families qui dominant sont celle des Argiopidcs qui y compte 27 especes 
dont 20 speciales, appartenant en grande partie au groupe des Tetragtiat/ia ; celle des 
Thomisidcs qui y compte 25 especes, toutes speciales; celle des Theridiides avec 17 
especes dont 13 speciales; celle des Attides avec 12 especes dont 8 speciales. 

II est a remarquer que les families qui manquent sont, sauf de rares exceptions 
{Ai'iadna), celles dont les especes vivent cachees dans des terriers comme les Avicu- 
larides, sous les pierres et les ecorces comme les Drassides et Chibionides, ou celles 
qui filent des toiles pres de terre et dans les endroits obscurs comme les Agelenides, 
tandis que les families dominantes sont celles dont les especes filent leurs toiles a 
decouvert sur les plantes comme les Argiopides et les Theridiides, ou qui chassent 
leur proie au soleil comme les Thomisides et les Attides. 

' Les Opiliones et les Acari ne sont pas compris dans ce memoire. 

Only one or two species of Acari have been obtained in the islands. Editor. 

F. H. II. 58 



444 FAUNA HAWAIIENSIS 

Dans chacune des families dominantes le nombre des genres est relativement faible, 
mais certains genres, Tkeridion, Teiragnaika, Sandalodes, etc. sont representes par de 
longues series d'especes voisines les unes des autres, les unes egalement repandues 
dans toutes les iles, les autres confinees dans une seule ; sous ce rapport I'ile la plus 
occidentale de Kauai nous a paru surtout interessante. 

Les genres nouveaux sont au nombre de six : le genre Do^yonyclms, voisin des 
Tetragnatlia, sen distingue par la structure toute speciale de ses tarses anterieurs ; le 
genre Syroloma appartient a la famille des Lycosides ; les genres Mecaphesa, Proernus 
{Pterelas), Pagiopalns et Adrastidia a celle des Thomisides ; les trois derniers, com- 
prenant ensemble huit especes, doivent former dans la sous-famille des Pliilodi^ominae 
un groupe nouveau, particulier a la faune des Sandwich. 



§ 2. Systematic account of the Arachnida. 

Ordo ARANEAE. 

Fam. SICARIIDAE. 

ScYTODES Latreille. 

( I ) Scytodes marmorata L. Koch. 

Scytodes marmorata L. Koch, Ar. Austr. 1872, p. 292, tab. xxiv. fig. 4. 

Hab. Oahu (1500 ft.), Perkins, x. 1896; Schauinsland. — Molokai and Hawaii; 
Schauinsland. 

Espece tres repandue dans la Malaisie Orientale, I'lndo-Chine, I'Australie, et la 
Polynesie ; deja indiquee des Iles Sandwich (Karsch, Thorell) ; trouvee par le Dr 
Schauinsland a Hawaii, Molokai et Oahu (E. Simon). 

Fam. DYSDERIDAE. 

Dysdera Latreille. 

(i) Dysdera crocata C. Koch. 

Dysdera crocata C. Koch, Arachn. t. v. 1839, p. 81, fig. 392 — 394. 
Dysdera rubicunda Blackwall, Spid. of Gr. Brit. p. 371. 
Dysdera interrita Hentz, in : Boston Journ. Nat. Hist. iv. p. 223. 
Hab. Hawaii: Kona. — Oahu; Schauinsland. 

Trouve anterieurement par le Dr Schauinsland a Oahu. Espece commune a 
I'Europe, a I'Amerique du Nord et a I'Amerique du Sud extratropicale, sans doute 
introduite aux Sandwich. 



ARACHNIDA 445 

Ariadna Audouin. 
(i) Ariadna perkiiisi, sp. nov. 

^. Long. 9-5 mm. Cephalothorax pallide fusco-rufescens, tenuiter nigro-margi- 
natus, longe ovatus, subtilissime coriaceus et punctis impressis parvis conspersus, 
acute marginatus, et fovea thoracica superficial! lata impressus. Oculi subaequales, 
medii ovati, inter se contigui sed antice attenuati et leviter divaricati. Oculi laterales 
utrinque ovati, contigui, et prominuli. Oculi medii superne visi cum lateralibus lineam 
recurvam designantes. Clypeus verticalis, planus, subtiliter transversim striatus, oculis 
anticis latior. Abdomen oblongum, teretiusculum, albido-testaceum, antice vitta longi- 
tudinal! postice sensim ampliata et truncata, dein maculis transversis seriatis 6 vel 
7, medianis triquetris, posticis angustis et linearibus atro-violaceis, supra notatum. 
Chelae fusco-olivaceae, longae et attenuatae, crebre transversim striatae. Partes oris 
sternumque pallide fusco-rufescentia, laevia et nitida. Pedes fulvi, antici, femoribus 
exceptis, paulo obscuriores. Pedes i' paris femore curvato supra ad apicem trla- 
culeato aculeoque interiore subapicali armato, patella mutica, tibia longa, tereti, subtus 
ad marginem exteriorem aculeis seriatis decern iniquis, apicali reliquis validiore et 
leviter uncato, ad marginem interiorem aculeis binis subapicalibus, extus aculeis 
lateralibus quatuor, intus aculeis lateralibus trinis armata, metatarso mutico sed longe 
et tenuiter piloso, cylindraceo et valde sinuoso-arcuato. Pedes-maxillares fulvi, tibia 
patella saltem duplo longiore crassiore et ovata, tarso parvo obtusissimo et longe 
piloso, bulbo subgloboso sed depressiusculo, spina longa ad basin recta ad apicem 
flexuoso-arcuata. 

Plate XV. fig. I. 

$. Long. 10 — 12 mm. A mari differt cephalothorace antice obscuriore fere nigro, 
chelis robustioribus nigris sublaevibus, abdomine longiore teretiusculo, maculis dorsa- 
libus majoribus ornato, pedibus anticis robustioribus et brevioribus, apice valde infus- 
catis, femore i' paris aculeis dorsalibus parvis trinis aculeisque interioribus validis et 
longissimis trinis armato, tibia aculeis inferioribus biseriatis 10, 11 vel 12 validis, 
longissimis parvisque intermixtis, armata, sed aculeis lateralibus carcnte, metatarso 
leviter curvato, haud flexuoso, subtus aculeis validis, singulariter elevatis, parvis 
longisque mixtis, 10 — 10, armato. Pedes-maxillares breves et robusti, fulvo-rufuli, 
tibia tarsoque nigris. 

Hab. Kauai. — Lanai (2000 ft.).— Oahu ; Mts. Waianae, Lolle. 

De toutes les especes de ce genre qui me sont connues A. Perkiiisi E. Sim. 
se rapproche surtout 6! A. Bosenbergi Keyserl., de la Republique Argentine, qui a le 
meme dessin abdominal ; elle sen distingue tout de suite par les epines de ses tibias 
anterieurs beaucoup plus nombreuses et alternativement plus longues et plus 
petites. 

58-2 



446 FAUNA HAWAIIENSIS 

Fam. PHOLCIDAE. 

Smeringopus E. Simon. 

(i) Smeringoptis elongaius Vinson. 

Pholcus elongatus Vinson, Aran. Reun. etc. 1864, p. 135. 
Pholcus phalangioides Dolesch., in: Act. Soc. Ind.-Neerl. v. 1859, p. 47. 
Pholcus tiptdoides L. Koch, Ar. Austr. 1872, p. 281, tab. xxiii. fig. 5. 
Pholctis distinctus Cambr., in: Linn. Soc. Journ. x. 1869, p. 380, tab. xi. figs. 

28—30. 
Pholcus inargarita Workman, in: Ann. Mag. Nat. Hist. 1878, p. 451, tab. 

XVIII. figs. I, 2. 
Hab. Oahu, Kaala Mts. (3000 ft.), March, 1893 ! Perkins. 

Espece repandue dans presque toutes les regions tropicales du monde. Sans 
doute introduite aux Sandwich. 

Artema Walckenaer. 

( I ) A rteina sisyphoides Doleschall. 

Indique des Sandwich par plusieurs auteurs (Thorell, E. Simon) ; n'y a pas ete 
retrouv^ par Mr Perkins. ■ 

Fam. THERIDIIDAE. 

Argyrodes E. Simon. 

(1) Argyrodes argyrodes Walckenaer. 

Linyphia m-gyrodes Walck., H. N. Ap. t. 11. 1841, p. 282. 
Theridion trigonimn Hentz in : Bost. Journ. N. H. vi. 1850, p. 280, tab. ix. 
figs. 24, 25. 

Argyrodes epeirae E. Simon, Ann. Soc. ent. Fr. 1866, p. 281, tab. i. figs. 4 — 7. 
Argyrodes argyrodes E. Simon, Ar. Fr. t. v. 1881, p, 16. 

Hab. Hawaii, Kona (3000 ft.). 

Cette espece est repandue dans la region mediterraneenne, dans presque toute 
I'Afrique, dans I'Asie tropicale et dans I'Amerique du Nord. 

Dans I'ancien monde elle accompagne presque toujours le Cyrtophora citricola 
Forsk., dont elle est parasite, mais aux lies Sandwich elle vit sur la toile de 
XArgtope avara Thorell, en meme temps que I'espece suivante. 



ARACHNIDA 447 

(2) Argyrodes hawaiiensis, sp. nov. 

(?• Long. 3— 4mm. Cephalothorax subtilissime coriaceus, fusco-olivaceus, utrinque 
et postice fere niger, parte cephalica antice sensim elevata vi.\ attenuata et obtu- 
sissima. Oculi quatuor antici in lineam valde procurvam, medii paulo majores a 
lateralibus quam inter se multo remotiores. Oculi postici parvi, aequi, in lineam 
subrectam, medii inter se quam a lateralibus remotiores. Oculi quatuor medii 
aream multo latiorem quam longiorem et postice quam antice multo latiorem 
occupantes. Clypeus verticalis, altissimus, baud cornutus, sed incrassatus, a regione 
oculari depressione profunda et pilosa, valde arcuato-angulosa, discretus. Abdomen 
postice sensim elevatum, in tuberculum crassum baud attenuatum, apice obtuse 
truncatum, productum, nigricans, supra vitta latissima, linea media exili secta, 
et utrinque vitta vel macula magna confusa et dentata luteo-argenteolis ornatum. 
Chelae longae et partes oris fusco-piceae, laeves. Sternum nigro-nitidum. Pedes 
olivacei, tarsis cunctis, femoribus metatarsisque quatuor posticis dilutioribus et luteis. 
Pedes-maxillares lutei, tibia tarsoque nigricantibus, femore sat longo tereti, patella 
longa versus basin sensim attenuata, tibia patella plus duplo breviore paulo angustiore 
sed extus obtuse ampliata, tarso breviter ovato, apice oblique truncato. 

Hab. Hawaii, Kona. 

Cette espece se rapproche un peu, par la structure de son bandeau, des A. 
minax Cambr. (de Madagascar) et A. affinis Cambr. (du Parana). 



Ariamnes Thorell. 

(i) Ariamnes corniger, sp. nov. 

%. Long. 8 — 10 mm. Cephalothorax longus, humilis, vix pone medium stria trans- 
versa, leviter procurva, impressus, in parte antica valde attenuatus subacuminatus, in 
parte altera fere parallelus, postice vix attenuatus et recte truncatus, fusco-rufescens, 
leviter reticulatus, vitta media angusta et Integra et utrinque vitta marginali, diluti- 
oribus et fulvis, notatus, fronte, inter oculos medios anticos, tuberculo fulvo, obtuse 
conico et antice oblique directo, insigniter munita. Oculi antici in lineam subrectam, 
medii reliquis oculis multo majores, nigri et convexi, inter se distantes sed a latera- 
libus vix separati. Oculi postici, superne visi, in lineam subrectam, vix procurvam, 
mediocres et inter se aequales, medii inter se quam a lateralibus saltem triplo 
remotiores. Clypeus verticalis, leviter convexus, area oculorum angustior. Abdomen 
angustum, antice truncatum, postice in tuberculum, corpore toto longius, cylindraceum 
sed apicem versus attenuatum et subacutum, postice oblique directum et plus minus 
arcuatum, insigniter productum, utrinque argenteolum, supra vitta lata saepe fusco- 



448 FAUNA HAWAII ENSIS 

marginata, obscuriore, aureo vel rufulo tincta, ornatum, tiiberculo utrinque saepe linea 
nigra abbreviata et punctata et subtus, pone mamlllas, macula nigricanti ornato. 
Chelae, partes oris sternumque fusco-rufula vel olivacea, sternum vitta media dilutiore 
notatum. Pedes graciles, inter se valde inaequales, pallide lutei, femoribus patellis 
tibiisque (apice excepto) i' paris fusco-rufulis, femoribus 2' paris antice, saltern ad 
apicem, rufulo-lineatis, femoribus tibiisque 4' paris apice minute rufulo-cinctis ; tibia i' 
paris metatarso saltern aequilonga femore vix breviore. Pedes-maxillares fusco-rufuli, 
femora dilutiora ; femore gracili apice haud incrassato ; tibia tereti ; tarso acuminate, 
tibia cum patella multo longiore. 

Plate XV. fig. 2. 

Hab. Maui, Haleakala. — Lanai, Koele. 

Cette espece remarquable fait presque le passage du genre Ariamnes au genre 
Rhoviphaea ; elle se rattache au premier par son bandeau etroit et vertical et par sa 
patte-machoire a femur et tibia cylindriques ; mais ses pattes anterieures dont le tibia 
est au moins aussi long que le metatarse ressemblent davantage a celles des Rhoni- 
phaea. Sa region oculaire est pourvue, meme chez la femelle (seul sexe connu), d'un 
tubercule median conique, caractere commun a plusieurs Rkomphaea mais jusqu'ici 
etranger aux Ariamnes. 



Theridion Walckenaer. 

Le genre Theridion est represente aux lies Sandwich par deux especes probable- 
ment introduites et au reste presque cosmopolites, les Theridion riifipes Lucas et tepi- 
dariorum C. Koch, et aussi par une nombreuse serie d'especes propres qui appar- 
tiennent toutes au groupe ayant pour types les Theridion frondeum Hentz et rusticum 
E. Sim. (cf Hist. Nat. An t. i. p. 537). 

La distribution de ces especes dans les di verses lies de I'archipel est curieuse : 

Hawaii : Theridion praetextiim E. S., grallator E. S., vielimun E. S., riifipes 
Lucas, tepidariorum C. Koch. 

Manai : Theridion praetextuni concolor E. S., melinum E. S., campestratiim E. S., 
manaiense E. S., haleakalense E. S., Perkinsi £. S., lepidarioj'mn C. K. 

Oahu : Theridion melinum E. S. 

Molokai : Theridion posticatum E. S., tepidariorum C. K. 

Lanai: Theridion posticatum E. S., tepidariorum C. K. 

Kauai : Theridion Perkinsi E. S., kauaicnsc E. S., acutitarse E. S. 



ARACHNIDA 449 

Synopsis specierum propriarum. 

1. Oculi medii antici lateralibus paulo minores. Pedes longissinii fere riwlci gralhxtor. 

Oculi medii antici lateralibus plus minus majores vel saltern haud minores. 

Pedes normales 2. 

2. Oculi antici inter se aequidistantes 3. 

Oculi medii antici a sese quam a lateralibus remotiores 5. 

3. Oculi postici in lineam parum procurvam, medii lateralibus paulo majores, 

spatio interoculari oculo multo latiore. Abdomen subtus impunctatum ( ? ) kauaiense. 

Oculi postici in lineam valde procurvam, medii a lateralibus quam inter se fere 
duplo remotiores, spatio oculo non multo latiore a sese distantes. Abdomen 
subtus, prope mamillas, minute fusco-bipunctatum 4. 

4. Bulbus maris dente interiore lata obtusa et subtriquetra. Plagula genitalis 

feminae convexa, laevis, semicircularis, antice puncto impresso, postice, 

in declivitate, foveola magna superficiali transversa, impressa Perkinsi. 

Bulbus maris dente interiore gracili acutissima et valde uncata. Plaga genitalis 
feminae ovato-transversa, plana, subtiliter coriacea, antice et praesertim 
postice transversim striata .praetexttim. 

5. Cephalothorax luteus, concolor. Pedes quatuor antici haud annulati. Femora 

4' parisapice late nigro-annulata. Areaoculorum mediorum subquadrata ( ? ) campestratum. 

Cephalothorax vitta media lata nigricanti vel olivacea notatus 6. 

6. Cephalothorax vitta media lata ornatus, sed linea marginali carens. Oculi 

medii postici lateralibus paulo majores. Oculi quatuor medii inter se 

subaequales. Femora 4' paris haud vel parum distincte annulata 7. 

Cephalothorax vitta media lata lineaque marginali exili nigricantibus notatus. 
Oculi medii postici lateralibus saltern haud majores et medii antici minores 
(c??) 9- 

7. Abdomen subtus impunctatum {$ 9) melinum. 

Abdomen subtus, prope mamillas, fusco nigrove binotatimi 8. 

8. Abdomen subtus, prope mamillas, punctis nigris parvis binis notatum ( c? 9 ). 

Bulbus maris apophysi interiore lata brevi et subtriquetra munitus haleakalense. 

Abdomen subtus, prope mamillas, maculis binis nigris subquadratis notatum 
((??). Bulbus maris apophysi interiore longa sat gracili recta et acuta 
munitus .posticatitm. 

9. Abdomen subtus, prope mamillas, nigricanti binotatum. Femora 4' paris apice 

late nigricanti-annulata. Peduni-maxillarium maris tarsus anguste ovatus 

et obtusus, bulbum haud superans matiiensf. 

.\bdomen subtus impunctatum. Femora cuncta apice niiimte fusco-notata. 
Pedum-maxillarium maris tarsus angustissimus, longe acuminatus et 
bulbum superans • • .acutiiarse. 



(i) Theridion tepidariorurn C. Koch. 

Thaddion tepidariorinn C. Koch, Arachn. viii. 1841, p. 75, figs. 647, 648. 
Theridion vtilgare Hentz, in : Bost. Journ. N. Hist. vi. p. 271. 
Hab. Hawaii, Kau, Olaa. — Molokai, Koloe. — Lanai. — Maui, Haleakala. 
Espece repandue dans toutes les regions chaudes et temperees du monde et dont 
I'orieine est incertaine. Sans doute introduite aux Sandwich. 

o 



450 FAUNA HAWAIIENSIS 

(2) Theridioii rjifipes Lucas. 

Theridion riifipes Lucas, Expl. Sc. Alg. Ar. p. 263, tab. xvi. fig. 5. 
Theridion borboniaim Vinson, Aran. Reun. etc. 1864, p. 283, tab. xiv. fig. 6. 
Theridion luteipes Cambr., in : Linn. Soc. Journ., x. 1870, p. 382, tab. xii. 

figs. 46—51- 
Theridion bajnlans L. Koch, J^%. u. Abyss. Ar. 1875, p. 21, tab. 11. figs. 4, 5. 
Tlieridion fiavoaurantiacum E. Simon, in: Ann. Soc. ent. Belg. C. R. Nov. 1880. 
Theridion longipes V. Hasselt, Midd. Sumatra, etc. Ar. 1882, p. n. 

Hab. Hawaii, Kona. 

Espece repandue dans la region mediterraneenne, dans I'Asie et I'Afrique tropi- 
cales, a Madagascar, en Malaisie et en Oceania. Probablement introduite aux 
Sandwich. 

(3) Theridion campestratiim, sp. nov. 

$. Long. 3*5 mm. Cephalothorax pallide luteus concolor, laevis et glaber, parte 
cephalica setis tenuibus triseriatis munita. Oculi quatuor antici in Hneam leviter pro- 
curvam, medii nigri et nigro-Hmbati lateralibus vix majores et inter se quam a laterali- 
bus remotiores (spatio inter medios oculo saltern baud angustiore, inter medios et 
laterales fere duplo minore). Oculi quatuor postici in lineam magis procurvam, medii 
lateralibus majores et a lateralibus quam inter se paulo remotiores. Oculi quatuor 
medii inter se aequales, aream parallelam subquadratam occupantes. Abdomen sub- 
globosum, supra albo-opacum, antice in declivitate vittis binis convergentibus, postice, 
supra mamillas, macula transversa laciniosa nigris notatum, subtus omnino pallide 
luteum. Chelae, partes oris sternumque fulvo-rufula, laevia. Pedes sat longi, femori- 
bus tibiisque 4' paris reliquis paulo robustioribus, sat longe setosi, lutei, patellis tibi- 
isque quatuor anticis rufulo-tinctis, pedum 4' paris femore ad apicem patellaque nigris, 
tibia apice anguste nigricanti-annulata. Plaga genitalis parva, fusca, simplex. 

Plate XV. fig. 3. 

Hab. Maui, Haleakala. 

(4) Tlieridion praetexttim, sp. nov. 

t. Long. 5 mm. Cephalothorax laevis et glaber sed parte cephalica setis longis 
triseriatis munita, luteus, vitta media fusco-olivacea, antice latissima sed postice acumi- 
nata, notatus. Oculi antici in lineam rectam, inter se fere aequidistantes, medii nigri 
lateralibus, albis et ovatis, majores. Oculi postici in lineam valde procurvam, medii, 
ovati et obliqui, a lateralibus quam inter se plus duplo remotiores. Area mediorum 
paulo longior quam latior et antice quam postice paulo latior. Abdomen breviter 



ARACHNIDA 451 

ovatum, parce et longissime albido-crinitiim, pallide testaceum, supra antlce macuHs 
nigris binis, dein utrinque nigro-punctatum, vitta media, in medio ampliata et saepe 
angulosa, antice lutea postice albo-opaca, ornatum, margine anteriore abdominis durius- 
cula, fulvo-rufula, in medio late et obtuse emarginata, utrinque lobo semicirculari, minu- 
tissime serrulato, munita, subtus regione epigasteris magna, convexa, fulvo-nitida, 
regione ventrali albido-testacea, utrinque linea obliqua exili fusca, postice punctis 
minutis nigris binis, plerumque notata. Chelae, partes oris sternumque fulvo-rufula. 
Chelae longae, ad basin convexae subgeniculatae, dein leviter depressae, apice atten- 
uatae et paululum divaricatae. Pedes longi, longe et valde setosi, antici reliquis multo 
longiores et saltem femoribus crassiores, lutei, femoribus, patellis tibiisque anticis leviter 
rufescenti-tinctis, metatarsis (3" exceptis) apice minute fuscis. Pedes-maxillares sat 
longi, lutei, tibia, superne visa, patella saltem aequilonga, ad basin multo angustiore sed 
apicem versus sensim ampliata, apice obtusa et setis longissimis seriatis 5 vel 6, ad 
marginem, munita, tarso mediocri, subacuminato, bulbo ad marginem exteriorem pro- 
cessu membranaceo apice ampliato et obtuso, ad marginem interiorem, prope medium, 
spina nigra longa sat angusta, acutissima et valde uncata (fere semicirculari) armato. 

?. Long. 4 — 6 mm. Cephalothorax pallide luteus, linea marginali carens, vitta 
media paulo obscuriore olivacea, antice lata evanescente et saepe obsoleta, postice 
attenuata notatus. Oculi antici in lineam subrectam (vix procurvam), inter se aequi- 
distantes (sed spatiis oculis mediis baud latioribus distantes), medii nigri et rotundi, 
lateralibus albis et ovatis majores. Oculi postici in lineam sat procurvam, mediocres, 
aequi, medii late ovati, a lateralibus quam inter se remotiores. Area mediorum paulo 
longior quam latior et antice quam postice paulo latior, medii antici posticis majores. 
Abdomen breviter ovatum vel subglobosum, parce et longe albido-crinitum, luteo- 
testaceum, vitta media confusa albo-punctata, utrinque punctis fuscis inordinatis paucis 
et postice, in declivitate, maculis nigris binis parallelis, valde sinuosis, notatum, inter- 
dum antice maculis nigris binis convergentibus, rarius maculis dorsalibus munitum, 
subtus, prope mamillas, punctis nigricantibus binis minutissimis, notatum. Partes oris 
fusco-rufulae. Chelae, sternum, pedesque lutea, tibiis i' paris apice leviter rufescenti- 
tinctis, interdum annulo fusco apicali angustissimo cinctis. Pedes robusti, longe et 
valde setosi. Plaga genitalis ovato-transversa, fusca, subtil iter coriacea, antice et 
praesertim postice tenuiter transversim striata. 

Plate XV. fig. 4. 

Hab. Hawaii, Kona, Kau. 

Var. Tkeridion praetexhim concolor. 

Cephalothorax omnino luteus, vittis carens. Abdomen pallide luteo-testaceum, 
vitta media albo-opaca, fere parallela, marginem anticum haud attingente atque antice 
punctis nigris binis supra ornatum. 

Hab. Hawaii, Kau. — Maui, Haleakala. 

F. H. II. 59 



452 FAUNA HAWAIIENSIS 



(5) TJieridion mainertse, sp. nov. 

$. Long. 4"5 mm. Cephalothorax fulvo-rufescens, vitta media lata fere parallela 
lineaque marginal! exili nigricantibus, notatus, laevis et glaber, parte cephalica setis 
tenuibus triseriatis munita. Oculi antici in lineam rectam, medii lateralibus majores et 
inter se quam a lateralibus remotiores (spatio inter medios oculo circiter aequilato, inter 
medios et laterales fere duplo angustiore). Oculi postici inter se subaequales, in lineam 
procurvam, medii a lateralibus quam inter se paulo remotiores. Area mediorum 
circiter aeque longa ac lata et antice quam postice paulo latior. Abdomen breviter 
ovatum, luteo-testaceum, utrinque punctis nigris iniquis et inordinatis, in medio vitta 
lata dilutiore valde flexuosa, et saltem in parte apicali tenuiter albo-marginata, notatum, 
margine antico abdominis rufulo duriusculo, utrinque lobo semicirculari, granulis obtusis 
uniseriatis 5 — 6 armato, munito, subtus regione epigasteris baud producta, fulvo-rufula, 
nigro-marginata, regione ventrali late et confuse nigricanti-plagiata. Chelae fulvo- 
rufulae, laeves, fere parallelae, sed extus ad basin, leviter convexae. Partes oris fusco- 
olivaceae. Sternum fulvo-rufulum, apice utrinque tenuissime nigro-marginatum. Pedes 
sat longi, sat longe setosi, femoribus, praesertim anticis, robustis, fulvo-rufescentes 
femoribus ad basin metatarsis tarsisque dilutioribus et luteis, sed femoribus 4' paris 
annulo apicali lato nigricanti fulvo-lineato, et metatarsis cunctis, praesertim anticis, apice 
minute fusco-annulatis. Pedes-maxillares lutei, mediocres, tibia, superne visa, patella 
vix breviore, paulo angustiore, apicem versus vix incrassata, subtus parum producta, 
obtusa et seriatim setosa, tarso tibia paulo longiore, sat anguste ovato sed obtuso et 
bulbum baud superante, bulbo ad marginem exteriorem processu membranaceo leviter 
lanceolato, ad marginem interiorem, prope medium, spina fusca, perpendiculari, gracili, 
acuta et leviter curvata, instructo. 

?. Long. 5 — 6 mm. Cephalothorax luteus, vitta media lata et fere parallela, 
postice leviter reticulata, lineaque marginali exillima antice evanescente, nigricantibus, 
notatus. Oculi antici magni, in lineam vix procurvam, medii lateralibus vix majores, 
inter se quam a lateralibus remotiores (spatio inter medios oculi latitudinem vix 
aequanti, inter medios et laterales fere 1/3 minore). Oculi postici in lineam vix 
procurvam subrectam, medii lateralibus paulo minores et a lateralibus quam inter se 
plus duplo remotiores (medii a sese spatio oculo angustiore distantes). Area mediorum 
circiter aeque longa ac lata et antice quam postice multo latior, medii antici posticis 
multo majores. Abdomen magnum, subglobosum, parce et longe albido-crinitum, fulvo- 
testaceum, supra crebre et grosse nigro-punctatum (in lateribus punctis minoribus zonas 
obliquas designantibus), vitta media dilutiore confusa, antice acuminata et alba, postice 
nigro-segmentata, ornatum, subtus prope mamillas nigricanti-binotatum. Chelae, sternum 
pedesque fulvo-rufescentia, partes oris, saltem ad basin, valde infuscatis, coxis 4' paris 
fusco-notatis, femoribus sex anticis, tibiis metatarsisque cunctis, annulo medio annuloque 



ARACHNIDA 453 

apicali fusco-rufulis, femoribus 4' paris annulo medio carentibus sed annulo apicali 
latiore nigricanti et fulvo-lineato ornatis. Plaga genitalis simplex, nigra, transversim 
semicircularis. 

Plate XV. fig. II. 



Hab. Maui. 



(6) Tlieridioii lialeakalejisc, sp. nov. 

t. Long. 4-5 mm. Cephalothorax luteus, vitta media nigricanti, antice lata et 
pone oculos lineata sed postice acuminata, notatus, laevis et glaber sed parte cephalica 
setis longis triseriatis munita. Oculi antici in lineam subrectam, medii nigri lateralibus 
vix majores, inter se quam a lateralibus remotiores (spatio inter medios oculi latitudinem 
vix aequanti, inter medios et laterales fere duplo angustiore). Oculi postici in lineam 
procurvam, medii lateralibus majores, plani et obtuse triquetri, a lateralibus quam inter 
se paulo remotiores. Area mediorum fere parallela et paulo longior quam latior, medii 
postici anticis majores. Abdomen breviter ovatum, pallide testaceum, supra grosse et 
inordinate albo-punctatum, utrinque punctis nigris magnis, marginem confusam desig- 
nantibus, et postice in declivitate linea nigra transversa notatum, margine anteriore 
abdominis coriacea, rufula, utrinque lobo minute serrulato munito, subtus regione 
epigasteris convexa rufula, regione ventrali testacea sed postice, prope mamillas, punctis 
nigris binis notata. Chelae fulvo-rufulae, laeves, fere parallelae, sed ad basin leviter 
convexae. Partes oris infuscatae. Sternum luteum. Pedes lon^i sat lonee setosi, 
femoribus, praesertim anticis, robustis, lutei, femoribus quatuor anticis apice leviter 
infuscatis, femoribus 4' paris annulo apicali lato nigricanti fulvo-lineato notatis, tibiis 
metatarsisque annulo medio vix expresso annuloque apicali distinctiore fusco-rufulis 
notatis. Pedes-maxillares lutei, tibia ad basin infuscata, tibia, superne visa, patella 
saltem haud breviore, paulo angustiore, apicem versus leviter ampliata, subtus producta, 
obtusa et seriatim setosa, tarso tibia paulo longiore, sat anguste ovato sed obtuso et 
bulbum haud superante, bulbo ad marginem exteriorem processu membranaceo, ad 
basin lato ad apicem angustiore striato et plicato, ad marginem interiorem, pone medium, 
processu fusco valde compresso, brevi et lato, instructo. 

?. Long. 5 mm. Cephalothorax luteus, linea marginali carens sed vitta media 
lata fere parallela nigricanti, leviter reticulata et saepe antice fulvo-punctata ornatus. 
OcuH antici in lineam leviter procurvam, medii lateralibus majores et a sese quam a 
lateralibus remotiores (spatio inter medios oculi latitudinem saltem aequanti sed spatio 
inter medios et laterales saltem duplo minore). Oculi postici in lineam procurvam, 
medii lateralibus paulo majores, obtuse triquetri, a lateralibus quam inter se paulo 
remotiores, sed spatio oculo paulo latiore a sese distantes. Oculi quatuor medii inter 
se subaequales, aream fere parallelam et subquadratam occupantes. Abdomen magnum. 

59—2 



454 FAUNA HAWAIIENSIS 

subglobosum, parce et longe albido-crinitum, obscure fulvo-testaceum, supra crebre 
nigro-punctatum, antlce macula longa, postice, supra mamillas, vitta abbreviata, albis, in 
medio vitta dilutiore lata confusa sed parce albo-punctata ornatum, subtus testaceum et 
prope mamillas minute fusco-bipunctatum. Chelae, sternum pedesque fulvo-rufula, 
partes oris valde infuscatae. Pedes late et confuse fusco-rufulo-annulati sed femoribus 
4' paris plerumque concoloribus. Plaga genitalis simplex, transversa et semicircularis, 
nigra, antice marginata. 

Plate XV. fig. 5. 

Hab. Maui, Haleakala. 



(7) Theridion perkinsi, sp. nov. 

$. Long. 5 — 6 mm. Cephalothorax laevis et glaber sed in parte cephalica setis 
nigris tenuibus et longis triseriatis munitus, luteus, vitta media obscuriore olivacea, 
antice latissima et plus minus reticulata, postice angustiore notatus. Oculi antici in 
lineam subrectam, inter se fere aequidistantes, medii lateralibus majores. Oculi postici 
mediocres, aequi, in lineam valde procurvam, medii obtuse triquetri, a lateralibus quam 
inter se plus duplo remotiores et spatio oculo vix latiore a sese distantes. Area 
mediorum paulo longior quam latior et antice quam postice latior. Abdomen breviter 
ovatum, parce et longissime fulvo-crinitum, albido-testaceum, supra utrinque nigro- 
punctatum et postice subsegmentatum, vitta media dilutiore, punctos albos includente, 
et interdum macula media magna subquadrata alba, ornatum, subtus utrinque linea alba 
obliqua et postice, ante mamillas, punctis nigris binis notatum, regione epigasteris fulva, 
convexa et laevi. Chelae fulvo-rufulae, laeves, longae et sat angustae, in medio 
depressae, sed extus ad basin convexae, ungue valido et longo. Partes oris fusco- 
rufulae, laminae longissimae, angustae et parallelae, pars labialis latior quam longior, 
tertiam partem basilarem laminarum vix attingens. Pedes longi, antici posticis multo 
longiores, femoribus saltern anticis sat robustis, valde et longe setosi, lutei, tibiis quatuor 
anticis apice sensim rufulo-tinctis, tibiis 4' paris in medio atque ad apicem anguste et vix 
distincte annulatis, metatarsis cunctis apice minute fuscis. Pedes-maxillares longi, lutei, 
tibia, superne visa, patella longiore, ad basin angustiore, apicem versus sensim ampliata, 
subtus producta obtuse truncata atque ad marginem setis longis seriatis munita, tarso 
brevi, sat lato et obtusissimo, bulbo brevi, ad marginem exteriorem processu mem- 
branaceo sat angusto, apice leviter lanceolato et canaliculato, ad marginem interiorem 
dente nigra vel fusca magna lata, depressa, subtriquetra, sed apice breviter subacuta, 
armato. 

$. Long. 6 — 7 mm. Cephalothorax luteus, vitta media lata paulo obscuriore 
confusa sed reticulata ornatus, linea marginali carens. Oculi antici in lineam rectam, 
inter se fere aequidistantes (medii a sese quam a lateralibus vix remotiores), medii 



ARACHNIDA 455 

lateralibus evidenter majores. Oculi postici aequi, in lineam sat procurvam, medii 
obtuse triquetri nigro-limbati, a lateralibus quam inter se multo remotiores. Area 
mediorum paulo longior quam latior et antice quam postice latior. Abdomen convexum, 
ovatum, valde et longe fulvo-setosum, albido-testaceum, supra grosse et inordinate nigro- 
punctatum (in lateribus punctis zonas obliquas confusas designantibus), vitta media lata 
alba, vel saepius in medio parce biseriatim albo-punctata et postice alba ornatum, subtus 
prope mamillas punctis fuscis binis notatum. Pedes lutei, tibiis ad apicem leviter rufulo- 
tinctis et subannulatis, metatarsis (3'' exceptis) apice minute fuscis. Plaga genitalis 
convexa, laevis, nigricans, paulo longior quam latior, antice rotunda et foveola parva 
nigra impressa, postice truncata et fovea majore transversim semicirculari notata. 

Plate XV. fig. 6. 

Hab. Maui, Haleakala. — Kauai, Koholuamana, Waimea Alts. 



(8) TheridioH kauaiense, sp. nov. 

?. Long. 7 — 10 mm. Cephalothorax laevis et glaber, sed parte cephalica setis 
triseriatis munita, luteus, vitta media obscuriore, antice lata et confusa, prope medium 
angustiore, nigricanti et reticulata, notatus. Clypeus convexus. Oculi antici in lineam 
rectam, inter se aequidistantes (vel medii a sese quam a lateralibus paulo remotiores), 
medii leviter prominuli lateralibus majores. Oculi postici in lineam leviter procurvam, 
medii ovati lateralibus vix majores et a lateralibus quam inter se vix remotiores (a sese 
spatio oculo multo latiore distantes). Area quatuor mediorum saltem baud longior 
quam latior et antice quam postice paulo latior. Abdomen ovatum, convexum, parce et 
tenuiter setosum, albido-testaceum, supra vittis duabus nigris, antice posticeque conver- 
gentibus, validissime flexuoso-dentatis et saepe interruptis, et interdum linea media 
nodosa abbreviata, in lateribus zonis obliquis fusco nigrove punctatis, subtus sensim 
evanescentibus, ornatum, subtus concolor. Chelae fusco-castaneae, laeves. Partes oris 
apice fusco-castaneae, basi luteae. Sternum luteum laeve. Pedes longi, longe et valde 
setosi, fulvo-rufuli, femoribus dilutioribus, tibiis late et parum distincte fusco-rufulo- 
triannulatis, metatarsis anticis ad basin atque ad apicem leviter infuscatis. Area geni- 
talis utrinque crasse pilosa, in medio fovea rufula superficiali subrotunda, antice et 
utrinque tenuiter marginata, atque antice, prope marginem, tuberculis minutissimis binis 
geminatis notata, impressa. 

Plate XV. fig. 7. 

Hab. Kauai, Waimea Mts. 



456 FAUNA HAWAIIENSIS 



(9) Theridioii gral/ator, sp. nov. 

$. Long. 4 mm. Cephalothora.x laevis et glaber, sed parte cephalica setis albis 
tenuibus triseriatis munita, albido-testaceus, parte cephalica vittis duabus pallide 
olivaceis, postice convergentibus et V designantibus, notata. Clypeus latus et planus. 
Oculi antici in lineam leviter procurvam, medii lateralibus paulo minores et inter se 
quam a lateralibus paulo remotiores. Oculi postici parvi et subaequales, in lineam 
leviter procurvam, medii a lateralibus quam inter se vix remotiores sed spatio oculo 
saltern duplo latiore a sese distantes. Area mediorum magna, parallela et subquadrata. 
Abdomen longe oblongum, pallide luteum, macula anteriore magna albo-opaca postice 
emarginata, antice et utrinque tenuiter nigro-marginata, dein punctis nigris quatuor, 
aream quadratam designantibus, supra ornatum. Chelae, partes oris pedes sternumque 
albido-testacea, laevia et nitida, tibiis i' paris apice leviter rufulo-tinctis. Pedes longis- 
simi, fere Pholci, sat longe setosi. Pedes-maxillares albidi, tibia patella paulo longiore 
vix graciliore, apicem versus vix incrassata, subtus parum producta, truncata et seriatim 
longe setosa, tarso parvo et ovato, tibia vix longiore, bulbo simplici, apice biaculeato. 

% (pullus). Long. 4"5 mm. Abdomen ovatum, macula maxima albo-opaca, postice 
leviter attenuata et truncata, antice vittam fulvam acute lanceolatam includente, supra 
fere omnino obtectum. Pedes albido-testacei, subpellucentes, femoribus tibiisque i' 
paris et 4' paris apice anguste nigricanti-annulatis. 

Plate XV. fig. 8. 

% varietas. Abdomen in parte basali macula nigra maxima trapezoidali notatum, 
in parte altera maculis parvis albis biseriatis ornatum. 

Hab. Hawaii, Kona. 



(10) Theridio7i acutitarse, sp. nov. 

t. Long. 3 mm. Cephalothorax laevis et glaber, sed parte cephalica setis 
triseriatis munita, fulvo-rufescens, linea marginali exili nigricanti, vittaque media nigri- 
canti et reticulata, antice lata postice attenuata, ornatus. Oculi antici in lineam levissime 
procurvam, medii leviter prominuli, lateralibus majores et inter se quam a lateralibus 
remotiores (spatio inter medios oculi latitudinem saltem aequanti, inter medios et 
laterales circiter 1/3 minore). Oculi postici, superne visi, in lineam leviter procurvam, 
medii lateralibus paulo majores, ovati vel obtusissime triquetri, a lateralibus quam inter 
se remotiores, spatio oculo haud latiore a sese distantes. Area mediorum saltem haud 
longior quam latior et antice quam postice latior. Abdomen breviter oblongum, parce 
et longe fulvo-setosum, fulvo-testaceum, plus minus albo-punctatum et maculis nigris 
iniquis et angulosis biseriatis, posticis saepe confluentibus et vittis binis valde dentatis 



ARACHNIDA 457 

formantibus, supra ornatum, subtus antice utrinque infuscatum, regione epigasteris 
convexa, fulvo-rufula et nitida. Chelae fulvo-rufulae, laeves, longissimae et angustae, 
sed extus ad basin leviter convexae, intus longe leviter emarginatae, angulo interiore 
apicali dente sat gracili sed truncato armato. Partes oris sternumque fusco-olivacea. 
Sternum tenuiter nigro-marginatum et macula parva apicali nigricanti notatum. Pedes 
fulvi, sat longe setosi, femoribus, tibiis metatarsisque annulo submedio vel subbasilari 
(saepe obsoleto) annuloque apicali fuscis vel rufulis ornatis. Pedes-maxillares fulvi, 
femore sat robusto, patella longiore quam latiore parum convexa, tibia patella longiore, 
paulo graciliore, apicem versus leviter ampliata, subtus parum producta sed longe setosa, 
tarso parvo, tibia vix longiore haud latiore, apice acuminate et bulbum simplex 
superante. 

%. Long. 3 '5 — 4 mm. Cephalothorax fulvo-rufescens, tenuissime, vix distincte, 
nigro-marginatus, parte cephalica antice leviter infuscata, postice macula triquetra nigri- 
canti, punctos binos fulvos includente, notata. Oculi antici in lineam levissime procur- 
vam, medii majores et inter se quam a lateralibus remotiores (spatio inter medios 
latitudinem oculi circiter aequanti, inter medios et laterales saltern duplo minore). Oculi 
postici in lineam subrectam, inter se fere aequidistantes. Area mediorum paulo latior 
quam longior et antice quam postice vix latior. Abdomen subglobosum, fulvo-testa- 
ceum, albo-pvmctatum, antice maculis binis magnis convergentibus, saepe confluentibus, 
dein maculis quatuor biseriatis minoribus et angulosis, postice utrinque macula majore, 
valde anguloso-dentata, nigris, ornatum, subtus antice utrinque infuscatum. Chelae, 
partes oris, sternum, pedesque fulvo-rufula, femoribus apice minute fusco-notatis, anticis 
subtus confuse infuscatis, tibiis metatarsisque annulo medio, saepe obsoleto, annuloque 
apicali, fusco-rufulis notatis. Area genitalis parva simplex et rufula. 

Plate XV. fig. 9. 

Hab. Kauai. 



(11) Theridion melinum, sp. nov. 

^. Long. 3*5 mm. Cephalothorax laevis et glaber, sed parte cephalica setis 
longis triseriatis munita, luteus, vitta media lata obscuriore olivacea, saepe obsoleta, 
notatus. Oculi antici in lineam subrectam, medii nigri et rotundi, lateralibus ovatis et 
albis paulo majores et inter se quam a lateralibus remotiores (spatio inter medios oculo 
saltern haud angustiore). Oculi postici in lineam sat procurvam, medii lateralibus paulo 
majores et a lateralibus quam inter se paulo remotiores. Oculi quatuor medii magni, 
aequi, aream circiter aeque longam ac latam et antice quam postice vix latiorem occu- 
pantes. Abdomen breviter oblongum, parce tenuiter et longe setosum, luteo-testaceum, 
supra nigro-punctatum, macula anteriore parva maculaque media maxima et subquadrata 
albis ornatum, interdum vitta Integra flexuosa, in medio valde dilatata et angulosa 



458 FAUNA HAWAIIENSIS 

postice sensim attenuata, ornatum, margine anteriore abdominis leviter coriacea, 
utrinque lobo obtuse serrato munito, subtus regione epigasteris leviter convexa, fulva, 
fusco-marginata, regione ventrali concolore vel in medio confuse infuscata. Chelae 
fulvae, laeves, ad basin leviter convexae, apice attenuatae et leviter divaricatae. Partes 
oris infuscatae, laminae ad basin dilutiores. Sternum luteum. Pedes longi, femoribus, 
saltern anticis, sat robustis, longe et valde setosi, patellis quatuor anticis, tibiis prope 
medium, seta erecta longiore armatis, lutei, interdum concolores, rarius tibiis anticis 
in medio atque ad apicem, tibiis posticis ad apicem metatarsisque plus minus fusco- 
rufulo-annulatis. Pedes-maxillares mediocres, lutei, tibia, superne visa, patella paulo 
breviore et angustiore, apicem versus sensim ampliata, subtus producta, obtuse truncata 
et setis validis et longis seriatis munita, tarso anguste ovato et obtuso, bulbo ad margi- 
nem exteriorem processu membranaceo, ad basin lato dein angusto et obliquo sed apice 
obtuso et tarsum paulo superante, in medio plagula fusca semicirculari, ad marginem 
interiorem spina brevi, antice directa, apice obtusa et minute uncata, munito. 

%. Long. 4 — 4'5 mm. Cephalothorax luteus, linea marginali carens, sed vitta 
media fusco-olivacea, antice lata, postice sensim attenuata notatus. Oculi antici in 
lineam leviter procurvam, medii lateralibus vix majores et a sese quam a lateralibus 
remotiores (spatio inter medios latitudine oculi paulo latiore). Oculi postici in lineam 
procurvam, medii lateralibus paulo majores, subrotundi vel obtusissime triquetri, a 
lateralibus quam inter se evidenter remotiores, spatio oculo saltem baud latiore a sese 
distantes. Oculi medii inter se aequales, aream paulo longiorem quam latiorem et 
antice quam postice paulo latiorem, occupantes. Abdomen subglobosum, luteo-testa- 
ceum, pictura dorsali valde variabili, plerumque punctis maculisque nigris parum 
regulariter biseriatis vittaque media dilutiore, valde dentata, antice albo-marginata, 
postice valde attenuata et alba ornatum, interdum omnino nigro-punctatum et postice 
nigro-segmentatum, sed antice vitta transversa alba, in medio anguloso-arcuata, notatum, 
rarius nigrum antice arcu transverso lato, postice, in declivitate, maculis parvis binis 
geminatis vittaque abbreviata albis, ornatum, subtus concolor vel in medio confuse 
infuscatum. Chelae fulvae, partes oris infuscatae. Sternum pedesque lutea, tibiis 
metatarsisque plus minus fusco-annulatis. Plaga genitalis fusca, transversim semi- 
circularis, ad marginem posticym foveola media minutissima nigra, punctiformi impressa. 

Plate XV. fig. lo. 

Hab. Hawaii, Kona, Kau. — Maui, Haleakala. — Oahu, Kaala Mts. 2000 ft., 
Waianae Mts. 

(12) Theridion posticatuni, sp. nov. 

t. Long. 4 mm. Cephalothorax fulvo-rufescens, vitta media nigricanti, antice lata 
postice sensim attenuata, notatus, laevis et glaber sed parte cephalica setis longis 
triseriatis munita. Oculi antici in lineam rectam, medii, rotundi nigri, lateralibus, albis 



ARACHNIDA 459 

et ovatis, paulo majores et inter se quam a lateralibus multo remotiores (spatio inter 
medios oculo saltern \ latiore, inter medios et laterales fere dupio minore). Oculi 
postici in lineam procurvam, medii sat longe triquetri, lateralibus vix majores sed a 
lateralibus quam inter se remotiores. Area mediorum paulo longior quam latior et 
antice quam postice latior. Abdomen (valde detritum) oblongum, parce et longe 
setosum, nigricans, supra vitta media alba, in medio ampliata et subangulosa, ornatum. 
Chelae fulvae, laeves, longae et angustae, sed ad basin leviter convexae. Partes 
oris infuscatae. Sternum pedesque lutea, tibiis cunctis ad basin atque ad apicem 
confuse infuscatis et subannulatis, femoribus 4' paris apice nigricanti-annulatis, meta- 
tarsis anticis ad basin atque ad apicem minute fuscis. Pedes longi, valde et longe 
setosi, antici posticis multo longiores. Pedes-maxillares mediocres, lutei, apice rufes- 
centi-tincti, tibia, superne visa, patella vix breviore angustiore sed apicem versus 
sensim ampliata, subtus producta, truncata et setis longis seriatis munita, tarso sat 
anguste ovato et obtuso, bulbo ad marginem exteriorem processu membranaceo valde 
arcuato apice bifido, ramulo exteriore obtuso, interiore tenui et acuto, ad marginem 
interiorem dente nigra valida et acuta antice recte directa et apicem tarsi saltern 
attingente. 

%. Long. 5 mm. Cephalothorax fulvo-rufescens, linea marginali carens, vitta 
media lata obscuriore, antice saepe confusa, in parte thoracica nigricanti et reticulata 
ornatus. Oculi antici in lineam leviter procurvam, magni et subaequales, medii latera- 
libus vix majores, inter se quam a lateralibus remotiores, sed spatio oculi latitudinem 
vix aequanti a sese distantes. Oculi postici in lineam sat procurvam, medii lateralibus 
paulo majores, obtuse triquetri, intus recti, a lateralibus quam inter se remotiores, 
spatio oculo evidenter angustiore a sese distantes. Oculi quatuor medii, magni et 
subaequales, aream antice quam postice paulo latiorem occupantes. Abdomen ob- 
longum, parce et longe fulvo-setosum, luteo-testaceum, supra punctis nigris iniquis et 
inordinatis, vittas duas latas et confusas designantibus, ornatum, subtus utrinque 
leviter nigricanti-variatum et postice, prope mamillas, maculis binis nigris sub- 
quadratis notatum. Chelae, sternum, partes oris pedesque fulvo-rufescentia, parte 
labiali laminisque intus infuscatis. Pedes sat robusti, valde et longe setosi, tibiis 
metatarsisque in medio atque ad apicem late rufescenti-tinctis et subannulatis. Plaga 
genitalis fusca, leviter coriacea plana, semicircularis vel subrotunda. 

Plate XV. fig. 12. 

Hab. Lanai, Koloe, 2000 ft. — Molokai. 



F. H. II. 



60 



46o FAUNA HAWAIIENSIS 

Teutana E. Simon. 

(i) Teutana grossa C. Koch. 

Theridion grosstim C. Koch, Arachn. iv. 1838, p. 112, fig. 321. 

Hab. Hawaii, Kona, Kau. — Maui, Haleakala. 

Espece tres repandue dans la region mediterraneenne, les iles de I'ocean atlan- 
tique et TAmerique du Sud extratropicale. Sans doute introduite aux Sandwich. 

Ulesanis L. Koch, 
(i) Ulesanis oahuensis, sp. nov. 

%. Long. I "5 mm. Cephalothorax brevissimus, superne omnino obtectus, in medio 
convexus, antice posticeque valde declivis, laevis et nitidus, fusco-piceus, nigricanti- 
marginatus. Oculi quatuor antici inter se contigui, in hneam rectam, medii nigri 
lateraHbus albis vix minores. OcuH postici in hneam vix procurvam, inter se anguste 
distantes, medii albi et leviter angulosi lateraHbus paulo majores. Area quatuor 
mediorum longior quam latior, in medio tuberculis obtusis parvis binis et geminatis 
ornata. Abdomen maximum ovatum, antice posticeque rotundum, altissimum et 
tuberculo medio grosso et obtusissimo munitum, albido-testaceum, supra valde et fere 
inordinate nigricanti-reticulatum et subniaculatum, subtus infuscatum. Sternum ru- 
fescens, nigro-marginatum. Pedes breves et robusti, tibiis anticis leviter clavatis, fulvo- 
rufescentes, femoribus tibiis metatarsisque subtus nigricanti-vittatis et apice subannulatis. 

Plate XVI. fig. II. 

Hab. Oahu, Waianae. 

Par son abdomen pourvu d'un seul gros tubercule median tres obtus, cette espece 
se rapproche surtout de V Ulesanis capensis E. Sim. de I'Afrique australe. 

Fam. ARGIOPIDAE. 

Subfam. LINYPHIINAE. 

Erigone Audouin. 

(i) Erigone vagans Audouin. 

Erigone vagans Audouin in Savigny, Descr. de I'Egypte, 2° ed. t. xxii. p. 319, 

tab. I. fig. 9. 
Neriene spinosa Cambr., Pr. Zool. Soc. Lond., 1872, p. 292, tab. xiii. fig. 2. 



ARACHNIDA 461 

Erigone litoralis L. Koch, Beitr. kenn. Ar. Fn. Tirols, 1868, p. 274. 
Erigone vagans E. Simon, Ar. France, t. v. 1884, P- 530. 
Hab. Maui, Haleakala. 

Cette espece, probablement introduite, a un habitat des plus vastes ; elle est en 
efifet repandue dans toute la region mediterraneenne et les lies de I'Atlantique, et a 
etd trouvee aussi a Singapore et au Cap de Bonne-Esperance, oil elle est commune. 



MicRONETA (Menge) E. Simon. 

Ce genre est represente dans les lies Hawaii et Maui par une espece de forme 
normale, differant cependant des especes europeennes et americaines par les cheliceres 
du male, pourvues en avant, pres le bord interne, d'une forte dent aigue, dirigee en 
bas ; caractere frequent dans le genre Tmeticus, mais jusqu'ici etranger au genre 
Microneta. 

( I ) Microneta insiilana, sp. nov. 

$. Long. 2 mm. Cephalothorax nigro-castaneus, subtilissime coriaceus et opacus, 
sat longe ovatus, parte cephalica antice leviter elevata, parum attenuata, fronte lata. 
Oculi antici in lineam rectam, medii inter se contigui, a lateralibus, paulo majoribus, 
anguste sejuncti. Oculi postici in lineam subrectam, sat magni, inter se aequi, sat 
anguste et fere aeque distantes. Clypeus latus, verticalis, sub oculis leviter depressus. 
Abdomen longe ovatum, nigrum, supra antice sensim dilutius et rufescenti-tinctum. 
Sternum nigrum. Chelae fusco-castaneae, subtiliter coriaceae, longae, validae et 
convexae, sed apice valde attenuatae et leviter divaricatae, antice, pone medium et 
prope marginem interiorem, dente acuto armatae, marginibus sulci longis et leviter 
depressis, dentibus parvis paucis remotissimis munitis, ungue longo. Pedes sat longi, 
fulvi. Pedes-maxillares mediocres, fulvi apice infuscati, tibia patella vix breviore, 
extus ad apicem fere supra, apophysi brevi bifida, ramulo superiore acuto et erecto, 
altero acuto sed antice secundum tarsum directo, armata, tarso sat anguste ovato. 

Plate XVIII. fig. 15. 

Hab. Hawaii, Kona. — Maui, Haleakala. 



Cnephalocotes E. Simon. 

Je rapporte avec doute a ce genre une espece des lies Sandwich qui differe de 
ses congeneres europeennes par ses yeux posterieurs presque equidistants et le groupe 
de ses yeux medians a peine plus long que large en arriere, mais qui sous tous les 
autres rapports (notamment par la structure de ses organes sexuels) est tres analogue 

60 — 2 



462 FAUNA HAWAIIENSIS 

aux C. pusillus Menge et citrtus E. Sim. Contrairement a ce qui a lieu dans les 
especes europeennes, le front du male, obtus et convexe, ne presente ni lobe ni 
impressions, et son bandeau, tres haut et un peu proclive, n'en pas convexe. 

Le C. simpliciceps E. Sim. a aussi des rapports avec le genre Entclecara. 

Le genre Cnephalocotes ne renfermait jusqu'ici que sept ou huit especes europeennes 
dont I'une (C. airtiis E. Sim.) a ete retrouvee en Egypte ; nous lui avons rapporte 
avec doute une espece du Bresil, dont la femelle est seule connue. 



(i) Cnephalocotes simpliciceps, sp. nov. 

$. Long. I "8 mm. Cephalothorax nigro-nitidus, brevis, parte cephalica con- 
vexa et obtusa, nee gibbosa nee impressa. Oculi antici in lineam sat procurvam, 
medii lateralibus paulo minores et a lateralibus quam inter se paulo remotiores. 
Oculi postici mediocres, in lineam latiorem leviter procurvam, inter se fere aeque 
€t sat late distantes. Area quatuor mediorum multo latior postice quam antice et vix 
longior quam postice latior. Clypeus area oculorum tota multo latior, leviter proclivis 
sed planus (nee convexus nee prominulus). Abdomen oblongum, convexum, supra 
scuto nigro-nitido, postice truncato, fere omnino obtectum. Partes oris chelaeque 
nigrae. Sternum nigro-opacum, magnum, convexum, postice late truncatum et coxis 
posticis subglobosis latius. Pedes debiles et breves, sat breviter et fere aequaliter 
pilosi, fulvo-olivacei, coxis femoribus ad basin patellisque dilutioribus. Pedes-maxillares 
sat breves, lutei apice infuscati, patella parva nodosa, tibia patella paulo longiore 
multo crassiore, extus apophysibus trinis iniquis armata : apophysi superiore longa sat 
gracili sed obtusa apice leviter plicata, mediana brevi dentiformi, inferiore multo 
longiore crassa apice truncata, tarso ovato et obtuso, supra ad basin depressiusculo, 
bulbo ovato nigro modice complicato. 

Plate XVIII. fig. 14. 

Hab. Hawaii, Kona. 



Labulla E. Simon. 

Le genre Labulla est represente aux lies Sandwich par deux grosses especes 
qui s'eloignent des formes typiques (Z. thoracica Wider) par certains caracteres qui 
semblent les rapprocher des Tetragnatha, notamment par leurs lames-maxillaires qui, 
au lieu d'etre arrondies, sont tronquees a I'extremite et anguleuses au cote externe 
■et par leurs cheliceres dont les marges sont armees de dents beaucoup plus nombreuses, 
mais il est a noter que ce dernier caractere est variable dans le genre Labulla, chez 
L. thoracica Wider, en effet la marge inferieure offre quatre petites dents tandis 



ARACHNIDA 463 

qu'elle n'en presente que deux chez L. altioculata Keyserl. et rnpicola E. Sim. ; la 
disposition de ces dents est au reste la meme : celles de la marge superieure sont 
plus fortes et plus longues que celles de la marge inferieure comme dans tous les 
genres de la sous-famille des Linypliimae. Leurs yeux medians posterieurs sont plus 
rapproches I'un de I'autre que des lateraux, ce qui s'observe aussi, mais a un moindre 
degre, chez L. altioculata Keyserling. 



(i) Labulla grapliica, sp. nov. 

^. Long. 8 '5 mill. Cephalothorax laevis, glaber, fusco-nigricans, reticulatus, 
vitta media lata fulvo-rufula, antice utrinque fusco-reticulata, postice attenuata et 
leviter dentata, ornatus, parte cephalica longa et attenuata, striis obliquis, utrinque 
profundis in medio evanescentibus, discreta, parte thoracica ovata, in medio late sed 
parum profunde canaliculata, haud striata. Oculi cuncti magni et subaequales, lineas 
binas subrectas designantes, medii antici et praesertim postici a lateralibus quam 
inter se fere \ remotiores. Oculi quatuor medii aream leviter prominulam, longiorem 
quam latiorem et antice quam postice angustiorem, occupantes. Oculi laterales 
utrinque a sese juxta contigui. Clypeus verticalis, area oculorum mediorum circiter 
aequilatus. Abdomen ovatum, valde convexum, nigricans, antice latissime dilutius et 
albido-testaceum, utrinque vittis latis binis valde flexuoso-dentatis, saepe confluentibus, 
postice lineolis obliquis seriatis albido-testaceis ornatum, subtus fusco-testaceum, regione 
epigasteris leviter coriacea. Chelae longae, parallelae, sed apice leviter divaricatae, 
fusco-rufulae, in parte basali laeves sed extus tenuiter rugosae, in parte apicali crebre 
et minute rugosae, marginibus sulci longis, superiore dentibus quatuor parvis, obtusis 
et inter se distantibus, dein, pone angulum, dentibus majoribus et acutis quinque, 
margine inferiore dentibus parvis subaequis 8 — 10, i — 4 inter se distantibus, reliquis 
subcontiguis, armatis. Laminae fusco-rufulae, apice late et recte truncatae, cum angulo 
exteriore leviter prominulo et subacuto. Pars labialis nigricans, haud longior quam 
latior et dimidium laminarum haud superans, apice obtusa et marginata. Sternum 
nigricans, opacum. Pedes sat robusti et, praesertim antici, longi, longe setosi et 
aculeis setiformibus armati, fulvo-rufuli, femoribus annulo subapicali lato annuloque 
apicali minore pallide fuscis, tibiis annulo medio fusco lato annuloque apicali nigro 
notatis, patellis fuscis, metatarsis rufulo-tinctis in medio atque ad apicem vix distincte 
infuscatis. Pedes-maxillares fulvi apice fusci, femore gracili curvato, patella parva, 
tibia patella saltem duplo longiore, supra ad apicem obtuse prominula, extus longe 
et valde nigro-setosa, tarso longe ovato, extus, prope basin, obtuse emarginato atque 
ad angulum inferiorem apophysi depressa acuta et antice directa armato, bulbo 
magno valde complicato. 

%. Long. 10 — 12 mill. Alari subsimilis, sed oculis quatuor anticis, antice visis, 
in lineam leviter recurvam, pedibus paulo brevioribus, femoribus haud vel vix distincte 



464 FAUNA HAWAIIENSIS 

annulatis, chelis robustioribus, antice setosis baud vel vix distincte rugosis, margine 
superiore sulci dentibus validis 6 vel 7 (1° angulari et ultimis binis minoribus) anticis 
trinis inter se distantibus reliquis subcontiguis, armato, pedibus-maxillaribus fulvis, 
apice sensim infuscatis. Area genitalis leviter convexa. postice subverticalis, in medio 
depressa et carinula testacea tenui divisa, utrinque fovea subrotunda vel leviter 
angulosa, crasse nigro-marginata, impressa. 

Plate XVI. fig. 2. 

Hab. Hawaii, Olaa. 

(2) Labiilla torosa, sp. nov. 

Z. Long. ceph. th. long. 6 mill. Cephalothorax laevis, fulvus, antice sensim 
rufescenti-tinctus, parte thoracica utrinque late infuscata et reticulata, parte cephalica 
striis profundis obliquis sed in medio evanescentibus discreta, thoracica obtuse cana- 
liculata baud striata. Oculi sat magni, inter se subaequales, quatuor postici, superne 
visi, in lineam levissime procurvam, medii a lateralibus quam inter se saltem duplo 
remotiores. Oculi antici in lineam subrectam, medii lateralibus paulo minores et a 
lateralibus quam inter se plus triplo remotiores. Oculi medii leviter prominuli, aream 
longiorem quam latiorem et antice quam postice angustiorem occupantes. Oculi late- 
rales utrinque a sese juxta contigui. Clypeus verticalis, area oculorum mediorum latior. 
Chelae fusco-rufulae, longissimae, apice attenuatae et leviter divaricatae, extus, prae- 
sertim ad apicem crebre et minute rugosae, marginibus sulci longis, superiore in parte 
apicali dentibus minutissimis et obtusis 4 vel 5, prope medium dentibus multo majoribus 
atque acutis 4 (2° paulo longiore), dein dentibus minoribus binis remotis, margine 
inferiore carinato, prope medium dentibus 6, parvis aequis et subcontiguis, lineam 
leviter arcuatam designantibus, instructis, ungue longo. Partes oris fusco-rufulae, 
laminae leviter inaequales, apice late truncatae cum angulo exteriore prominulo et acuto, 
pars labialis circiter aeque longa ac lata, valde inaequalis, apice rotunda et crasse margi- 
nata. Sternum fuscum opacum. Pedes longi, setis longis aculeisque setiformibus 
muniti, fulvi, patellis fusco-rufulis, tibiis in medio vi.x infuscatis sed annulo apicali 
nigricanti notatis, metatarsis apicem versus leviter infuscatis. Pedes-maxillares magni, 
fulvi apice nigri, femore gracili curvato, patella parva convexa, tibia patella longiore, 
graciliore, sed apice incrassata, longe nigro-crinita, tarso longe ovato, extus in medio 
valde dilatato et anguloso, intus ad basin apophysi nigra depressa acuta et antice 
directa, dein tuberculo rufulo lato et obtuso munito, bulbo maximo, apophysibus 
plurimis rufulis nigrisque insigniter instructo. 

%. Long. 12 — 15 mill. Cephalothorax opacus, fusco-castaneus, ad marginem 
obscurior, regione oculari nigra. Oculi sat magni et subaequales, quatuor postici, 
superne visi, in lineam leviter procurvam, medii inter se quam a lateralibus fere \ 
remotiores, quatuor antici, antice visi, in lineam vix recurvam, subrectam, medii 



ARACHNIDA 465 

lateralibus paulo minores et a lateralibus quam inter se saltern duplo remotiores. 
Oculi quatuor medii leviter prominuli, aream longiorem quam latiorem et antice 
quam postice angustiorem occupantes, antici posticis paulo minores. Oculi laterales 
utrinque a sese juxta contigui. Clypeus area oculorum mediorum paulo latior, ver- 
ticalis sed sub oculis leviter depressus. Abdomen oblongum, convexum, sat longe et 
tenuiter pilosum, nigricans, supra antice late dilutius et testaceum, et utrinque vittis 
binis obliquis testaceis, postice obsoletis, notatum, subtus in medio dilutius. Chelae 
longae, fusco-castaneae, opacae, minute et parce rugosae, marginibus sulci longis, 
superiore dentibus acutis validis inter se fere aequis (ultimis binis paulo minoribus) 
et fere aequidistantibus, margine inferiore dentibus multo minoribus aequis et sub- 
contiguis 10 vel 11 armatis. Partes oris castaneae. Sternum fuscum opacum. Pedes 
longi et validi, longe setosi et aculeis setiformibus muniti, obscure fulvo-rufuli vel 
castanei, tibiis in medio late infuscatis et annulo apicali nigricanti notatis, metatarsis 
apicem versus infuscatis. Pedes-maxillares castanei, femore patellaque dilutioribus. 
Vulva fere praecedentis, fovea media triquetra cannula tenui divisa et utrinque 
foveola crasse nigro-marginata notata. 

Plate XVI. fig. I. 

Hab. Hawaii, Olaa. — Maui, Haleakala. — Molokai Mts. — Kauai, Waimea. 

Subfam. TETRAGNATHINAE. 
DoRYONYCHUs, nov. gen. 

A. Tetragnatha cui subsimilis est differt pedibus omnino muticis, quatuor anticis 
reliquis multo longioribus, tarsis onychio sat longo munitis et unguibus binis, inter se 
valde iniquis et dissimilibus, instructis : ungue exteriore brevi, gracili et recto dentibus 
parvis paucis munito, ungue interiore plus decuplo longiore, ad basin valde curvato, 
dein subrecto, gracili et acutissimo, subtus in parte basali dentibus obliquis seriatis, 
minutissimis armato, subtus in parte apicali mutico sed valde compresso et acute 
carinato. 

L'espece pour laquelle nous proposons ce genre ne differe absolument des 
Tetragnatha que par la structure tres remarquable de ses tarses et de ses griffes 
anterieures, qui est unique dans I'ordre entier des Araneae. 

Les tarses des deux premieres paires sont coupes, vers leur quart apical, d'une 
fausse articulation, formant un onychium assez long, cylindrique, garni en dessous 
de crins rudes, et arme a I'extremite de deux griffes tres inegales et tres dissemblables ; 
I'externe est petite, grele, presque droite et armee de quelques petites dents, I'interne 
est au moins dix fois plus longue, fortement courbee a la base en forme d'agrafe, 
ensuite presque droite, tres grele et terminee en pointe effilee, pourvue en dessous, 
dans sa partie basale seulement, d'une serie de tres petites dents contigues, carinul^e 
et mutique en dessous dans toute sa partie apicale. 



466 FAUNA HAWAIIENSIS 



(i) Doryonychus raptor, sp. nov. 

^. Long. lo mill. Cephalothorax angustus, longus et humilis, parte cephalica 
striis integris profundis discreta, parte thoracica utrinque obtuse marginata, striis 
radiantibus profundis depressioneque media longitudinali, antice profunda et triquetra, 
postice evanescente, impressa, fronte angusta et obtusa, in medio leviter prominula, 
pallide luteus, oculis singulariter nigro-cinctis. Oculi postici inter se aequidistantes, in 
lineam valde recurvam. Oculi antici in lineam circiter aequilatam recurvam. Oculi 
quatuor medii inter se subaequales, aream subquadratam occupantes. Oculi laterales 
utrinque spatio oculo (postico) baud vel vix latiore a sese distantes, posticus antico 
fere duplo major. Abdomen angustum et longissimum, teretiusculum sed postice acu- 
minatum, albido-argenteum, supra confuse aurantiaco-tinctum. Chelae luteae nitidae, 
sat longae et proclives, supra et extus convexae, supra in parte apicali apophysi sat 
brevi valida sed subacuta et arcuata munitae, margine superiore sulci dente i° 
mediocri recto, dente 2°, primo paulo longiore, acuto, dein dentibus minoribus seriatis 
quatuor munito, margine inferiore dente 1° subapicali sat parvo, dente 2° paulo 
longiore, dein dentibus seriatis minutissimis (1° reliquis vix majore) 4 — 5 armato, 
ungue simplici baud dentato. Partes oris, sternum pedesque lutea, parte labiali 
leviter infuscata, tibiis, metatarsis tarsisque ad apicem leviter infuscatis. Sternum 
paululum convexum, nitidissimum. Pedes omnino mutici, quatuor antici reliquis 
multo longiores, tarsis subtus ad apicem onychiisque subtus rude-pilosis. Pedes- 
maxillares Tetragnathae, graciles et longi, tibia patella multo longiore, ad basin angus- 
tiore sed apicem versus sensim ampliata, apophysi tarsali tibia circiter aequilonga, 
gracili et subacuta, subtus in medio angulosa, tarso gracili et longo, bulbo depres- 
siusculo sed spina tenui et acuta vix arcuata et leviter contorta, tarso vix breviore, 
munito. 

$. Long. 12 — 15 mill. Mari subsimilis sed chelis apophysi superiore carentibus, 
abdomine longissimo, paulo crassiore, antice posticeque attenuato, albido-argenteo, 
superne aurantiaco-tincto, vulva, longe pone spiracula locata, postice plagula parva 
rufula ovata, fere verticali, munita. 

Plate XVI. fig. 8— Plate XIX. fig. i. 

Hab. Kauai. 

Tetragnatha Latreille. 

Le genre Tetragnatha, tres abondant dans I'archipel, y est represente par neuf 
especes qui lui paraissent speciales, a I'exception cependant du T. mandibulata Walck., 
qui y a peut-etre ete introduit. 

Ces Tetragnatha peuvent se rapporter a trois groupes ; les T. mandibulata W., 



ARACHNIDA 467 

hawaiensis, iieirix, Perkinsi E. Sim. ; rentrent dans le premier groupe ayant pour 
type le T. extcnsa L. (Hist. Nat. Ar. t. i. p. 720), a cela pres cependant, que 
leur premiere ligne oculaire, vue en dessus, est un peu plus large que la seconde ; 
leurs pattes ne sont armees que d'epines assez courtes et peu nombreuses et le tibia 
de la patte-machoire de la femelle est mutique. 

Les T. restricta, kaimiensis et ttncifcra E. Sim., qui pourraient former un groupe 
special et nouveau, ont cependant la meme disposition oculaire, mais leurs pattes 
sont armees d'epines divergentes beaucoup plus longues et plus nombreuses, rappelant 
celles des T. gracilis Stol. et gemmata L. Koch ; le tibia de leur patte-machoire chez 
la femelle est arme, de chaque cote, d'une longue epine divergente, enfin leur abdomen 
est le plus souvent rhomboedrique, plus ou moins elargi, convexe et parfois tubercule 
au milieu. 

Cette forme est encore exageree dans le T. cuncivcntris E, Sim., qui differe de 
ses congeneres par ses tibias et metatarses anterieurs mutiques et qui pourrait aussi 
devenir le type d'un groupe partlculier. 

Le tableau suivant resume les caracteres des Tetragnatha des lies Sandwich. 



Synopsis specierum. 

Mares. 

1. Chelarum margo superior dente apicali magno, reliquis dentibus multo majore 

et antice oblique directo arniatus mandibulata. 

Chelarum margo superior dente apicali parvo vel nullo 2. 

2. Chelarum margo superior dente apicali carens, dente 1° parvo et remote, dente 

2° maximo, prope medium sito 3. 

Chelarum margo superior dente apicali parvo, ad radicem unguis sito, munitus 5. 

3. Chelarum margo superior dente 1° mediocri remote et uncato uncifera. 

Chelarum margo superior dente 1° remoto, parvo, recto et acuto 4. 

4. Chelarum margo inferior dente apicali minutissimo et obtuso, dente 2" sub- 

apicali valido et longo armatus cimeiventris. 

Chelarum margo inferior dente apicali carens, dente subapicali sat valido d,xxndX\x%...restricta. 

5. Chelarum margo superior dente apicali minutissimo obtuso subrotundo 6. 

Chelarum margo superior dente apicali compresso truncato cariniformi et 

minute bifido 7- 

6. Chelarum margo inferior dente apicali parvo et obtuso, dente 2" subapicali 

valido longo et leviter sinuoso-arcuato, reliquis dentibus seriatis a dentibus 

apicalibus longe remotis iietnx. 

Chelarum margo inferior dente apicali parvo et obtuso, dente 2° subapicali 
valido sed sat brevi et recto, reliquis dentibus seriatis a dentibus apicalibus 
parum distantibus perktnsi. 

7. Chelarum margo superior dente 3° submedio longissimo, reliquis dentibus 

seriatis inter se subcontiguis sed a dente 3" remotis kauatensis. 

Chelarum margo superior dente 3° submedio mediocri, reliquis dentibus seriatis 

inter se fere aeque et sat late distantibus sobrina. 

F. H. II. 6' 



468 FAUNA HAWAIIENSIS 

Fetninae. 

1. Tibia pedum-maxillarium utrinque aculeo setiformi longo et divaricato vel seta 

niunita. Tibiae anticae subtus aculeis tenuibus longissimis et divaricatis 

armatae 2. 

Tibia pedum-maxillarium mutica. Tibiae anticae utrinque aculeis brevibus 

pronis 3 vel 4 armatae, rarius muticae 4. 

2. Tibiae anticae aculeis 4 — 4 subtus munitae. Abdomen rhomboidale restricta. 

Tibiae anticae aculeis longissimis 5 — 5 vel 6 — 6 subtus munitae 3. 

3. Chelarum margo inferior dente apicali sat valido (dentibus seriatis baud minore) 

armatus. Abdomen angustum et teretiusculum. Tibiae anticae aculeis 

longis 6 — 6 armatae kauaiensis. 

Chelarum margo inferior dente apicali niinutissimo (dentibus seriatis multo 

minore) armatus. Abdomen in medio ampliatum et convexum imcifera. 

4. Tibiae metatarsique antici omnino mutici. Abdomen breve supra elevatum 

et turbinatum ciineivejitris. 

Tibiae metatarsique antici aculeati. Abdomen longum et teretiusculum 5. 

5. Chelarum unguis subtus ad basin minute dentatus, margo inferior dente apicali 

maximo cultriformi recto et antice directo armatus (tab. xvii. fig. 7) matidibulata. 

Chelarum unguis simplex baud dentatus 6. 

■6. Chelarum margines duos dente apicali simili, longo, acuto et recto sed leviter 

obliquo muniti. Abdomen subtus concolor nctrix. 

Chelarum margo inferior dente apicali parvo munitus. Abdomen subtus 

vittatum 7 . 

7. Chelarum margo inferior dente apicali parvo et obtuso. Abdomen sat breve 

subtus vitta obscuriore confusa notatum ...perkinsi. 

Chelarum margo inferior dente apicali niinutissimo sed acuto et antice directo 

armatus. Abdomen longum, vitta lata nigra tenuiter albido marginata 

subtus notatum haivaietisis. 

( I ) Tctraguaiha viandibiilata Walckenaer. 

Tetragnatha viajtdibttlata Walck. Apt. t. xi. 1841, p. 211. 
Tetragnatlia minatoria E. Simon, in: Ann. Soc. ent. Fr. 1877, p. 83. 
Teti'agnatha leptognatha Thorell, St. Rag. Mai. etc. t. i. 1877, p. 101 (441). 
Tetragnatha minatoria Thorell, in: Ann. Mus. civ. Geneva, 1887, p. 133. 
Hab. Oahu. — Kauai (Schauinsland). — Molokai (Perkins). 

Espece tres repandue dans la Malaisie orientale, I'lndo-Chine et les Philippines; 
decrite de I'lle Guam (Mariannes) par Walckenaer. 

(2) Tetragnatha netrix, sp. no v. 

$. Long. 8 — 10 mm. Cephalothorax angustus et longus, pallide fulvo-rufescens, 
pilis longis albis vestitus, fovea thoracica profunda transversa sulcisque cephalicis 
impressus, fronte angusta. Oculi quatuor postici, superne visi, in lineam valde 
recurvam, quatuor antici, antice visi, in lineam paulo latiorem subrectam. Oculi 
quatuor medii aream subquadratam occupantes, antici posticis saltem \ majores. 
Oculi laterales utrinque aequi, spatio oculo fere duplo" latiore a sese distantes. Ab- 



ARACHNIDA 469 

domen longissimum, teretiusculum, apice oblique sectum et mamillas pauIo superans, 
albido-luteum, apice punctis fuscis minutissimis quatuor notatum. Chelae longissimae, 
proclives, fulvo-rufulae, laeves, supra, in parte apicali, apophysi arcuata, apice minu- 
tissime bifida, armatae, margine superiore sulci dente i", ad radicem unguis remote, 
parvo recto, antice oblique directo, dente 2° maximo, crasso sed apice sat abrupte 
acuto, dein dentibus acutis multo minoribus quinque, versus basin sensim minoribus 
et inter se fere aequidistantibus, munito, margine inferiore, ad apicem, dentibus angu- 
laribus binis, 1°, prope radicem unguis sito, minutissimo et obtuso, 2° multo majore, 
intus directo, leviter flexuoso et arcuato, dein dentibus parvis seriatis 5 vel 6 fere 
aequidistantibus, r et 2° reliquis paulo minoribus, armato, ungue longo, simplici baud 
dentato. Pars labialis fusca. Laminae, sternum pedesque lutea. Pedes longissimi, 
aculeis tenuibus et longis armati, tibiis anticis utrinque aculeis quatuor aculeoque 
dorsali submedio, metatarsis aculeis basilaribus binis aculeoque interiore tantum 
munitis Pedes-maxillares ordinarii, apophysi tarsali angusta subrecta et obtusa intus, 
ante medium, angulosa et tuberculo rufulo obtuse truncato, munita. 

%. Long. 10 — 12 mm. A mari, cui subsimilis est, differt chelis paulo brevioribus 
et validioribus, apophysi superiore carentibus, margine superiore sulci dente apicali sat 
valido recto acuto et leviter retro obliquo, dein dentibus seriatis 5 vel 6, inter se 
anguste et fere aeque distantibus, versus basin sensim minoribus, margine inferiore 
dente apicali, dente marginis superioris simili, dein dentibus seriatis quinque (a dente 
apicali parum remotis), versus basin sensim minoribus, 1° leviter curvato, armatis. 
Pedes lutei, tibiis anticis aculeis sat longis utrinque trinis, metatarsis aculeis basi- 
laribus verticillatis aculeoque submedio interiore, armatis. Pedum-maxillarium patella 
seta spiniformi apicali, tarsus setis similibus plurimis armati sed tibia mutica. 

Plate XIX. fig. 6. 

Hab. Hawaii, Olaa, Kau. 

Cette espece, qui a le facies du T. mandibulata Walck., s'en distingue par ses 
yeux lateraux anterieurs a peine plus petits que les postt^rieurs ; chez le male par 
I'apophyse des cheliceres plus eloignee de I'apex et par I'armature des marges ; a la 
marge superieure en effet la dent angulaire manque tandis qu'elle est tres forte chez T. 
mandibulata, la premiere dent, assez eloignee de la base du crochet, est petite, 
tandis que la seconde est tres developpee, longue et robuste, a la marge inferieure la 
seconde dent angulaire est au contraire beaucoup plus longue que celle de T. man- 
dibulata et d'une forme speciale. 

La femelle se distingue tres facilement de T. mandibulata car ses cheliceres 
manquent de la dent apicale inferieure dirigee en avant et d^passant la base du 
crochet, tandis que chacune de ses marges offre une dent angulaire semblable droite 
aigue et un peu oblique. 

Pour T. mandibulata voy. PI. XIX. fig. 7. 

61—2 



470 FAUNA HAWAIIENSIS 



(3) Tetragnatha hatvaiensis, sp. nov. 

?. Long. 10 — 12 mm. Cephalothorax fulvus, in medio confuse olivaceo-tinctus. 
Oculi antici, antice visi, in lineam recurvam. Oculi quatuor medii inter se sub- 
aequales, aream paulo longiorem quam latiorem et antice quam postice angustiorem, 
■occupantes. Oculi laterales antici posticis multo minores. Abdomen longe oblongum, 
in medio leviter convexum, luteum, vitta foliiformi latissima, leviter flexuosa, albido- 
marginata et, saltern antice, linea albida tenui secta, supra ornatum, subtus vitta 
media lata nigricanti albido-marginata ornatum. Sternum nigricans. Chelae robustae, 
oblique proclives, margine superiore sulci dente angulari minutissimo et acuto serieque 
remotissima dentium 6, versus basin sensim minorum, munito, margine inferiore dente 
angulari mediocri acuto antice directo, serieque, parum remota, dentium 7 — 8, versus 
basin sensim minorum, munito. Pedes fulvo-olivacei, femoribus tibiis metatarsisque 
apice minute fuscis. Pedes antici sat breviter aculeati, tibiis utrinque aculeis brevibus 
3 vel 4, metatarsis aculeis basilaribus binis et intus aculeis minoribus binis (2° 
submedio) munitis. Pedum-maxillarium patella supra ad apicem seta spiniformi, meta- 
tarsus utrinque setis similibus armati sed tibia mutica. 

Plate XIX. fig. 4. 

Hab. Hawaii, Olaa, Hilo. 

Cette espece, dont le male est encore inconnu, est assez voisine du T. man- 
dibulata Walck., dont elle se distingue cependant par I'armature des marges de ses 
cheliceres qui manquent de la dent apicale cultiforme caracteristique. Elle se distingue 
de T. netrix E. Sim. par ses yeux lateraux anterieurs beaucoup plus petits que les 
post^rieurs, par son abdomen marque d'une bande noire ventrale tres nette, par les 
epines de ses pattes beaucoup plus courtes, enfin par ses cheliceres dont la marge 
superieure offre une dent angulaire tres petite, rudimentaire, I'inferieure une dent 
angulaire plus forte, obliquement dirigee en avant. 



(4) Tetragnatha perkinsi, sp. nov. 

<?. Long. 7 — 9 mm. Cephalothorax longus, fulvo-rufescens, fovea thoracica 
magna, subrotunda, sulcisque cephalicis impressus, fronte sat lata. Oculi postici, 
superne visi, in lineam valde recurvam, medii lateralibus paulo majores et a lateralibus 
quam inter se paulo remotiores. Oculi antici, antice visi, in lineam leviter recurvam, 
vix latiorem. Oculi quatuor medii inter se subaequales, aream vix longiorem quam 
latiorem et antice quam postice angustiorem occupantes. Oculi laterales utrinque sub- 
aequales leviter prominuli et a sese parum separati, spatio interoculari oculo non 



ARACHNIDA 471 

multo latiore. Abdomen cylindraceum sat breve, obscure fulvum, parce nitido- 
punctatum, subtus baud punctatum. Chelae fulvo-rufulae, laeves, longissimae et 
proclives, supra, in parte apicali, apophysi arcuata, apice minute et inaequaliter bifida, 
armatae, margine superiore sulci dente 1° apicali minutissimo, dente 2° remoto 
parvo, dente 3° remoto et submedio, multo majore et acuto, dein dentibus minoribus 
seriatis quatuor, versus basin sensim minoribus, armato, margine inferiore, dente 
apicali parvo et obtuso, dente 2° majore et compresso a praecedenti subgeminato, dein 
dentibus quinque mediocribus inter se fere aequidistantibus, \" et 2° reliquis paulo 
minoribus, munito, ungue longo, simplici, baud dentato. Laminae fusco-olivaceae. 
Pars labialis nigra. Sternum obscure fulvum. Pedes longi, fulvi, minute et parum 
distincte rufulo-annulati, aculeis parvis paucissimis (ordinariis) armati. Pedes-maxil- 
lares fulvi, longi ordinarii, tibia patella multo longiore, ad basin angustiore sed 
versus apicem sensim incrassata, apophysi tarsali tibia circiter aequilonga subrecta et 
obtusa, subtus prope medium tuberculo rufulo truncate, cum angulo anteriore minute 
prominulo, munita. 

%. Long. 8 — 12 mm. A mari differt chelis brevioribus et validioribus, convexis, 
apophysi superiore carentibus, margine superiore sulci dente angulari mediocri et 
obtuso, dein dentibus seriatis quinque, ad apicem remotis, sed inter se appropinquatis, 
versus basin sensim minoribus, margine inferiore dente apicali mediocri acuto et 
leviter curvato, dein dentibus inter se aequidistantibus quinque, i, 2, et 3 inter 
se aequis (1° leviter curvato), reliquis sensim minoribus, armatis. Abdomen sat 
breviter ovatum, obscure fulvum, sat crebre flavo-niticlo-punctatum, subtus vitta media 
lata obscuriore, parum expressa, notatum, pedibus fulvis, distinctius fusco-variatis et 
subannulatis, sat longe setosis sed aculeis paucis mediocribus armatis, tibiis utrinque 
aculeis sat brevibus trinis aculeoque dorsali setiformi submedio, metatarsis aculeis 
basilaribus similibus plurimis aculeoque submedio munitis. Pedum-maxillarium patella 
seta spiniformi apicali, tarsus setis similibus plurimis armati sed tibia mutica. Vulva 
pone spiracula sita, postice late et recte truncata. 

Plate XIX. fig. 5. 

Hab. Maui, Haleakala. — Hawaii, Kilauea. 

(5) Tetragnatha sobrina, sp. nov. 

t. Long. 6 mm. T. perkinsi affinis. Cephalothorax similis sed parte thoracica 
in medio atque ad marginem confuse infuscata et olivacea. Oculi postici, .superne 
visi, in lineam valde recurvam, medii lateralibus paulo majores et a lateralibus quam 
inter se paulo remotiores. Oculi antici, antice visi, in lineam subrectam vix latiorem. 
Oculi quatuor medii aream circiter aeque longam ac latam et antice quam postice 
angustiorem occupantes, antici posticis paulo minores. Oculi laterales utrinque sub- 
aequales, leviter prominuli et a sese parum distantes, spatio interoculari oculo non 



472 FAUNA HAWAIIENSIS 

multo latiore. Abdomen cylindraceum, sat breve, obscure fulvum, supra parce albo- 
punctatum sed vitta media impunctata, antice arcubus tenuibus fuscis postice punctis 
nigricantibus biseriatis 5 — 5 ornata, notatum. Chelae fulvo-rufulae, laeves, longissimae 
et proclives, supra in parte apicali apophysi sat brevi, subrecta et subacuta, armatae, 
margine superiore sulci dente 1° apicali parvo compresso et oblique truncato, dente 
2° parvo, dente 3° majore recto et acuto, dein dentibus seriatis 4 vel 5, inter se fere 
aequidistantibus et versus basin sensim minoribus, armato, margine inferiore dente 
apicali minutissimo, obtuso, subrotundo, ad radicem unguis sito, dente 2° subapicali 
majore et acuto, dein dentibus parvis sex, inter se fere aequidistantibus, ultimis binis 
reliquis minoribus, munito, ungue longo simplici, baud dentato. Laminae fusco- 
olivaceae. Pars labialis nigricans. Sternum pedesque fulva, femoribus anticis leviter 
fusco-maculatis, patellis tibiisque apice leviter infuscatis. Pedes-maxillares a praece- 
denti differt processu apicali tarso breviore et crassiore, apophysi ad marginem in- 
feriorem tuberculo parvo subrotundo, baud truncato et longe ante medium sito, 
instructa. 

Plate XIX. fig. 8. 

Hab. Hawaii, Kona. 

T. sobrina E. S. est voisin de T. perkinsi E. Sim. dont il differe surtout par 
ses yeux medians anterieurs un peu plus petits que les posterieurs, par la pointe 
tarsale de sa patte-machoire plus courte et plus epaisse, par son apophyse tarsale pour- 
vue, au bord inferieur, bien avant le milieu, d'un petit tubercule obtus, ni tronque ni 
malleiforme, enfin par I'armature des marges de ses cheliceres ; a la marge superieure 
en effet la dent apicale est obliquement tronquee et un peu bifide au lieu d'etre 
conique, la 3" dent est relativement plus petite et moins reculee ; a la marge inferieure 
les deux dents apicales sont presque semblables dans les deux especes, mais chez 
T. sobrina les dents seriees sont a peu pres semblables entre elles, sauf les deux 
dernieres qui sont un peu plus petites, tandis que chez T. Perkinsi les deux premieres 
dents sont un peu plus petites que les autres. 



(6) Tetragnatha kanaiensis. 

$. Long. 6 mm. Cephalothorax angustus et longus, pallide fulvo-rufescens, 
parte cephalica postice, thoracica ad marginem, leviter infuscatis, fovea thoracica ovata 
postice attenuata sulcisque cephalicis impressus, fronte angusta. Oculi quatuor postici, 
superne visi, in lineam valde recurvam, inter se aequales et fere aequidistantes. Oculi 
quatuor antici in lineam subrectam vix latiorem, Oculi quatuor medii aream paulo 
latiorem postice quam longiorem et antice quam postice angustiorem, occupantes, antici 
posticis paulo majores. Oculi laterales utrinque aequi et inter se spatio oculo non 
multo latiore distantes. Abdomen angustum et longum, obscure fulvum, luteo-nitido 



ARACHNIDA 473 

punctatum et supra maculis fuscis paucis biseriatis notatum. Chelae longissimae et 
proclives, fulvo-rufulae, laeves, supra in parte apicali apophysi subrecta, apice nigra et 
minutissime bifida, armatae, margine superiore sulci dente apicali, ad radicem unguis 
sito, minutissimo compresso truncate cariniformi et minutissime bilobato, dente 2° 
remote parvo, dente 3° maximo recto et acuto, dein dentibus remotis, acutis 4 vel 5, 
versus basin sensim minoribus, armato, margine inferiore dentibus mediocribus 8, inter 
se fere aequidistantibus, sed leviter iniquis, munito, ungue longo, simplici, haud dentato. 
Pars labialis fusca. Laminae, sternum pedesque fulvo-rufescentia. Pedes aculeis te- 
nuibus sed longissimis instructi, metatarsis anticis ad basin aculeis longis verticillatis, 
et utrinque prope medium aculeo parvo armatis. Pedes-maxillares fulvi, ordinarii, 
apophysi tarsali tibia circiter aequilonga, angusta sed obtusa, intus, prope basin, tu- 
berculo rufulo et obtuso munita. 

%. Long. 8 mm. A mari difiert chelis brevioribus et validioribus, apophysi 
superiore carentibus, margine superiore sulci dente apicali mediocri acuto, dein den- 
tibus seriatis subcontiguis sex, 1° valido reliquis sensim minoribus, margine inferiore 
dentibus parvis seriatis (apicali vix majore) 6 — 8 armato. Abdomen longum et 
angustum, supra flavo-aureum, subtus fulvum. Pedes longi, lutei, aculeis tenuibus 
longissimis et numerosis muniti, femoribus anticis utrinque aculeis longissimis fere 
setiformibus 6 — 6, tibiis subtus aculeis similibus 6 — 6, metatarsis ad basin aculeis 
verticillatis, dein (ante medium) aculeo simili munitis. Pedes-maxillares patella supra 
ad apicem, tibia metatarsoque utrinque aculeis divaricatis longissimis munitis. 

Plate XIX. fig. 9. 

Hab. Kaui, Halemanu. 



(7) Tetragnatha rest rid a, sp. nov. 

Z. Long. 5 '5 mm. A T. kauaiensi, cui valde affinis est, differt cephalo- 
thorace breviore fronte latiore, obscure fulvo-rufescenti, parte cephalica postice macula 
dilutiore confusa notata, oculis quatuor posticis, superne visis, in lineam valde re- 
curvam, inter se aequalibus, mediis a lateralibus quam inter se vix remotioribus, 
oculis quatuor anticis, antice visis, in lineam vix latiorem, subrectam, oculis quatuor 
mediis inter se aequis, aream evidenter latiorem quam longiorem et antice quam 
postice angustiorem, occupantibus, oculis lateralibus utrinque inter se spatio oculo non 
multo latiore distantibus, antico postico paulo minore, abdomine paulo breviore, antice 
obtuse truncato et emarginato, prope medium leviter ampliato, apicem versus longe 
attenuate, obscure fulvo, crebre albo-argenteo-punctato, utrinque, prope medium, leviter 
infuscato, subtus vitta media fulva (impunctata) notato. Chelarum margines ut in 
T. kauaiensi armati sed margo superior dente parvo apicali cariniformi carens et 
dente maximo (2°) paulo remotiore. Caetera fere T. kanaiensis. 



474 FAUNA HAWAIIENSIS 

%. Long. 6 — 7 mm. A mari differt chelis brevioribus et validioribus, margine 
superiore sulci dente apicali parvo et acuto, dein dentibus seriatis, ab apice remotis 
sed inter se contiguis, quinque, i° et 2° sat validis et inter se acquis, rellquis versus 
basin sensim minoribus, margine inferiore fere similiter dentato, sed serie dentium a 
dente apicali minus remota. Abdomen rhomboidale, in medio ampliatum et convexum, 
antice et praesertim postice declive et valde attenuatum, obscure fulvum, crebre albo- 
argenteo-punctatum. Pedes obscure fulvi, antici leviter et confuse fusco-variati, patellis 
anticis aculeo apicali gracili, tibia utrinque aculeis quatuor tenuibus longissimis et 
divaricatis, metatarsis ad basin aculeis trinis verticillatis (lateralibus longissimis, dorsali 
parvo) aculeoque exteriore submedio armatis. Pedum-maxillarium patella supra ad 
apicem tibiaque utrinque seta longissima et divaricata armatae. 

Plate XIX. fig. ,o. 

Hab. Hawaii, Kona. 



(8) Tetragnatha uticifera, sp. nov. 

t. Long. 6—7 mm. Cephalothorax modice longus, fronte lata, fovea thoracica 
lata subrotunda, striis cephalicis profundis impressus, obscure fulvo-rufescens vel oli- 
vaceus, tenuiter nigro-marginatus, parte cephalica utrinque infuscata et lineolis mediis 
binis fuscis exillimis et subgeminatis, antice evanescentibus, notata, oculis singulariter 
nigro-cinctis. Oculi postici superne visi, in lineam valde recurvam, inter se aequi et 
aequidistantes. Oculi antici, antice visi, in lineam vix recurvam, vix latiorem. Oculi 
quatuor medii aream circiter aeque longam ac latam et antice quam postice angus- 
tiorem occupantes, antici posticis majores. Oculi laterales utrinque leviter prominuli 
et a sese parum separati, spatio interoculari oculo vix latiore, anticus postico paulo 
minor. Abdomen longe oblongum, in medio plus minus convexum. Chelae fulvo- 
rufulae laeves, longissimae et proclives, supra in parte apicali apophysi arcuata, apice 
minute bifida, armatae, margine superiore sulci, dente 1°, apice longe remoto, valido 
sed acuto et uncato, dente 2° longiore acuto et recto, dein dentibus minoribus 5, 
inter se aequidistantibus et versus basin sensim minoribus, armato, margine inferiore 
dentibus subapicalibus binis mediocribus, 2° 1° paulo majore, dein dentibus minutis- 
simis binis, inter se remotis, dein dentibus seriatis 6 — 7, vix majoribus et versus 
basin sensim minoribus, munito, ungue longo, simplici baud dentato. Partes oris 
sternumque fusca, parte labiali fere nigra. Pedes longi, obscure fulvi, patellis fuscis, 
femoribus tibiisque annulis fuscis mediis apicalibusque notatis, metatarsis tarsisque apice 
sensim infuscatis, aculeis tenuibus et lonms armati, tibiis anticis aculeis inferioribus 
4 — 4 aculeisque dorsalibus binis, metatarsis aculeis basilaribus verticillatis 2 vel 3, 
aculeoque submedio munitis. Pedes-maxillares lutei fusco-variati, longi, ordinarii, tibia 



ARACHNIDA 



475 



patella multo longiore, ad basin angustiore sed versus apicem sensim ampliata, apo- 
physi tarsal! angusta, subrecta et obtusa, intus, ante medium, angulosa et tuberculo 
obtuso munita. 

?. Long. ■]■ — lo mm. A mari differt chelis brevioribus et validioribus convexis, 
apojDhysi superiore carentibus, margine superiore sulci dente angulari mediocri et 
obtuso, dein dentibus seriatis, ad apicem remotis sed inter se appropinquatis, 5 — 6, 
1° et 2° validis et inter se acquis, reliquis multo minoribus, margine inferiore dente 
apicali parvo, dein serie dentium 6 — 7 validorum sed versus basin sensim minorum, 
armato, abdomine in medio sensim ampliato, supra convexo et saepe obtuse tuber- 
culato, cinereo-testaceo et crebre albido-punctato, vitta foliiformi lata fusca, utrinque 
nigro-marginata, in medio abrupte ampliata et angulosa, postice sensim attenuata et 
flexuosa, supra ornato, interdum vitta obsoleta sed macula media triquetra et utrinque 
maculis parvis binis nigris ornata, subtus in medio confuse infuscato, regione epi- 
gasteris rufulo-tincta, pedibus distinctius fusco-annulatis, femoribus anticis utrinque 
aculeis 5 — 5, exterioribus parvis, interioribus longioribus, patellis aculeo apicali longis- 
simo, tibiis aculeis inferioribus tenuibus sed longissimis 5 — 5 vel 4 — 5, aculeoque 
dorsali submedio, metatarsis aculeis basilaribus verticillatis similibus aculeoque interiore 
submedio instructis. Pedes-maxillares lutei fusco-annulati, patella supra ad apicem 
seta spiniformi longa, tibia utrinque seta simili, tarso setis plurimis munitis. Vulva 
pone spiracula sita, postice late et recte truncata. 

Plate XIX. fig. 2. 

Hab. Maui, Haleakala. — Hawaii ; Kau, Kona, Mauna Loa. — Kauai. 



(9) Tetragnatlia ameiventris, sp. nov. 

^. Long. 5 — 6 mm. A praecedenti differt area oculorum mediorum antice quam 
postice paulo angustiore et saltem baud latiore quam longiore, chelarum margine 
superiore dente 1° mediocri, ad apicem minus remoto, acuto et recto, baud uncato, 
dente 2° 1° plus duplo longiore recto et acuto, dentibus minoribus basilaribus trinis 
munito, margine inferiore dente apicali minutissimo et obtuso, dente 2" multo majore 
acuto et recto, dente 3° minore, dente 4° singulariter sito, parvo, dentibusque basi- 
laribus seriatis trinis armato, pedum aculeis paucis, debilibus et brevibus, metatarsis 
anticis, utrinque ad basin, minutissime aculeatis sed aculeo medio carentibus. 

%. Long. 7 mm. A mari differt abdomine breviore et latiore, antice minute 
emarofinato, in medio dilatato, altissimo et obtuse turbinato, obscure cinereo, crebre 
albido-punctato, prope apicem transversim nigro-notato et subvittato, postice in 
declivitate utrinque macula magna obliqua nigerrima et maculis mediis fuscis minoribus 
seriatis ornato, subtus vitta media nigricanti notato, chelis brevioribus et validioribus, 
subverticalibus, nitidis, obscure fulvis, antice et extus fusco-vittatis, dentibus marginum 

F. H. II. 62 



476 FAUNA HAWAIIENSIS 

fere praecedentis, laminis fulvis extus fusco-marginatis, parte labiali sternoque nigri- 
cantibus, pedibus fulvis, valde fusco-annulatis, femoribus anticis aculeis paucis seti- 
formibus munitis sed reliquis articulis muticis, patellis seta apicali, tibiis setis erectis 
binis supra munitis. Pedum-maxillarium patella seta apicali, tarsus setis plurimis 
muniti sed tibia mutica. 

Plate XIX. fig. 3. 

Hab. Maui, Haleakala. 



Subfam. ARGIOPINAE. 

Argiope Audouin. 

(i) Argiope avara Thorell. 

A. avara Thorell, Freg. Eug. Resa, in Vetensk. Akad. Handl. 1868. 

A. avara, McCook, Amer. Spid., t. iii. p. 222, tab. xiv. f i. 

$. Long. 18 — 25 mm. Cephalothorax fulvo-rufescens, longe et crebre albo- 
sericeo-pubescens, clypeo leviter infuscato. Area oculorum mediorum longior quam 
latior et antice quam postice angustior, oculi antici posticis vix majores. Oculi 
laterales utrinque valde prominuli et contigui, anticus ovatus postico rotundo plus 
duplo minor. Abdomen late ovatum, antice rotundum, postice breviter attenuatum 
et obtusum, superne leviter deplanatum, albido-cereolum, antice lineolis transversis 
binis exilibus et abbreviatis, prope medium lineis transversis integris, saepe flexuosis, 
postice vittis transversis 5 — 7, sensim latioribus, densioribus et saepe albido-punctatis, 
interdum confluentibus, nigerrimis decoratum, in lateribus nigrum valde albido- 
punctatum et maculatum, subtus regione epigasteris testacea, regione ventrali nigra, 
vittis binis rectis sed postice leviter divaricatis, mamillas baud attingentibus, et maculis 
medianis parvis binis albis notata. Sternum atrum, vitta latissima, utrinque lobata, 
fulvo-rufula notatum. Coxae fulvo-rufulae fusco-marginatae et variatae. Pedes robusti 
et longi, fusco-rufescentes, femoribus anticis subtus fere nigris, posticis confuse annu- 
latis, sericeo-pubescentes, crebre et sat longe nigro-setosi. Tuberculum genitale 
profunda foveolatum, et carinula alta testacea, antice leviter canaliculata, postice 
sensim ampliata et plana, munitum. 

Z. Long. 8 — 10 mm. Cephalothorax fuscus, longe et crebre albo-pilosus, versus 
marginem dilutior et vitta media albido-lutea, antice latissima, postice sensim angus- 
tiore, notatus. Abdomen angustum et parallelum, antice rotundum, postice breviter 
acuminatum, supra crebre albo-punctatum et vitta media latissima et parallela paulo 
obscuriore et tenuiter nigro-marginata, notatum, subtus nigricans et vittis albis binis 
integris ornatum, mamillis fulvo-rufulis. Sternum albido-luteum, utrinque late nigri- 



ARACHNIDA 477 

canti-marginatum. Pedes obscure fulvi, femoribus, patellis tibiisque crebre nigro- 
punctatis, metatarsis tarsisque apice leviter et sensim infuscatis, aculeis debilibus 
armati, metatarsis anticis aculeis basilaribus et submediis munitis sed apicalibus caren- 
tibus. Pedes-maxillares lutei, apice fusci ; patella parva nodosa supra seta longissima 
et erecta munita ; tibia brevi et transversa, intus prominula et longe setifera ; tarso 
longe lacinioso, supra ad basin processu magno, intus obtuse conico et rufulo, extus 
nigro, transversim cariniformi et bisinuoso ; bulbo maximo, supra plagula rufula, clypei- 
formi semicirculari, tenuissime concentrice striata, ad apicem lamina styliformi nigra, 
valde plicata munita. 

Hab. Forma typica ; Lanai, Koele. — Hawaii ; Kau, Kona. — Oahu, Waianae 
Mts. — Maui, Haleakala. 



Var. a. avara kauaiensis, subsp. nov. 

Abdomen supra nigrum, antice vitta transversa lata saepe flexuoso-dentata, dein 
maculis octo, transversim biseriatis, quatuor anticis majoribus ovatis et saepe dentatis, 
albo-cereolis, supra decoratum. 

Hab. Kauai Mts. 

Espece decrite des Sandwich par Thorell qui I'indique egalement d'Australie a 
Sydney (St. Rag. Mai., t. iii. p. 69) et de Californie, retrouv^e depuis dans I'Arizona 
et le Texas (McCook). 

Nous avons decrit son cocon ovigere qui ressemble a celui de XA. bruennichi 
Scopoli, d'Europe (Zool. Jahrb. 1899, p. 416). 

C'est probablement de cette espece dont le D' Karsch parle sous le nom d'Argiope 
aenmla Walck. (Sitz.-Ber. Ges. Naturf. Freunde, Mai 1880, p. ']^). 



Cyclosa Menge. 

Le genre Cyclosa est represente aux lies Sandwich par deux especes tres re- 
pandues dans I'ancien et le nouveau monde, C. oculata et hcrbinata Walck., par 
une espece C. albisternis E. Sim. que nous connaissions deja des lies Andamans et de 
I'lnde et par cinq especes qui paraissent speciales. C. perkinsi, simplicicaiida et 
xanthomelas E. Sim. sont voisins des precedents et de forme normale, mais le dernier, 
C. xanthomelas, offre un style de coloration tres special. 

Les deux autres C. olorina et cucurbitula E. Simon, rentrent dans le groupe du 
C. micula Thorell, caracterise par un abdomen brievement ovalaire ou globuleux 
(cf. Hist. Nat. Ar. t. i. p. 780). 

62 — 2 



478 FAUNA HAWAIIENSIS 



(i) Cyclosa oculata Walckenaer. 

Aranea oculata Walckenaer, Fn. Par. ii. 1802, p. 428. 

Epeira oculata Walckenaer, Apt. 11. 1841, p. 145. 

Cyclosa oculata E. Simon, Ar. Fr. i. 1874, p. 41. 

(?) Epeira Walckenaeri Keyserling, Sp. Amer. Epeirldae, 1892, p. 98, tab. v. f. 85. 

(?) Cyclosa Walckenaeri McCook, Amer. Spid. etc. in. p. 226, tab. xvii. f. i. 

Hab. Oahu, Honolulu. 

Espece probablement introduite, dont I'habitat est fort etendu, car elle se trouve 
en France surtout dans les Jardins (ou elle a sans doute aussi ete introduite), dans 
I'Afrique australe, au Natal, et au Cap, dans le sud des Etats-Unis (MacCook), au 
Guatemala ; je I'ai recue aussi de la Jamaique et de St Domingue, et je I'ai trouvee 
au Venezuela. 



(2) Cyclosa turbinata Walckenaer. 

Epeira turbinata Walck., Apt. 11. 1841, p. 140. 

Epeira caudata Hentz, J. Bost. S. vi. 1850, p. 23. 

Cyclosa turbinata McCook, Amer. Spiders etc. iii. 1893, P- 224, tab. xvii. f. 

5-6. 
Epeira strangulata L. Koch, Ar. Austr. i. p. 118, tab. ix. f. 6. 

Hab. Hawaii, Kona. — Maui, Haleakala. — Kauai ; Waimea, Koholuamano. 

Cette espece a, comme la precedente, un habitat tres vaste : elle est repandue 
en Malaisie (Sumatra), aux lies Viti (L. Koch), en Nouvelle-Caledonie et dans I'Ameri- 
que du Nord, du Canada a la California et a la Floride. 

Epeira camelodes Thorell (d'Amboine) en est peut-etre aussi synonyme (?). 



(3) Cyclosa albisternis E. Simon. 

C. albisternis E. Simon, Journ. As. Soc. Beng., lvii. 1887, p. i. No. 3 (188S) 
p. 285. 

Hab. Kauai. 

Tout a fait semblable au type des lies Andamans ; I'espece existe aussi dans le 
sud de rinde. 



ARACHNIDA 479 



(4) Cyclosa perkinsi, sp. nov. 

?. Long. 5 mm. Cephalothorax niger et nitidus, parte cephalica parce albo- 
setosa, parte thoracica late ovata convexa, cephalica abrupte angustiore, postice sulco 
profundo leviter procurvo discreta. Oculi ordinarii. Abdomen oblongum, antice altum 
et tuberculis obtusis binis munitum, postice leviter attenuatum et declive sed apice 
truncatum et obtusissime trilobatum, supra plerumque cinereo-testaceum, nigro- 
punctatum et reticulatum, pone tubera punctis nigris densioribus vittas duas confusas 
designantibus, subtus utrinque late albido-plagiatum, in medio nigrum maculis albis 
quatuor, anticis alteris duplo majoribus subquadratis et inter se transversim late 
remotis. Sternum nigrum, in medio dilutius et confuse vittatum. Pedes lutei, fe- 
moribus (praesertim anticis) apice late nigricanti-annulatis et subtus in parte basali 
annulo multo angustiore notatis, tibiis metatarsisque in medio atque ad apicem fusco- 
annulatis, tarsis apice fuscis. Vulva scapo alto transverso olivaceo-nitido, utrinque 
obtuso, in medio leviter depresso, atque unco brevi, obtuso et linguiformi, munita. 

Var. Abdomen supra albido-opacum, utrinque et antice, in declivitate, leviter 
nigro-reticulatum. 

A Cyclosa turbinata W., cui valde affinis est, differt imprimis abdomine postice 
baud turbinato sed obtusissime trilobato, subtus maculis albis quatuor notato sed 
macula alba anteriore carente ; a C. ociilata W. differt coxis luteis, sterno in medio 
subvittato sed ad marginem baud maculato, maculis albis ventralibus majoribus etc. 

Hab. Hawaii, Kau. 



(5) Cyclosa siiiiplicicauda, sp. nov. 

?. Long. 8 — 12 mm. Cephalothorax fulvo-olivaceus, laevis, parce albido- 

setosus, parte cephalica saepe antice leviter fusco-reticulata et postice plerumque lineis 
exilibus fuscis binis convergentibus notata, thoracica saepe utrinque et postice sensim 
infuscata, parte thoracica ovata, leviter convexa, impressione media, antice truncata, 
munita, cephalica abrupte angustiore, sat longa et convexa, sulco profundo semi- 
circulari discreta. Oculi quatuor medii aream paulo longiorem quam latiorem et antice 
quam postice latiorem occupantes, postici anticis minores, spatio oculo plus duplo an- 
gustiore a sese distantes. Oculi laterales utrinque aequi et contigui, a mediis late 
distantes. Abdomen longe oblongum, plus duplo longius quam kitius, antice breviter 
attenuatum et obtusum, postice, pone mamillas, sat longe productum, sensim attenuatum 
sed obtusum fere ut in C conica, interdum postice haud attenuatum et obtusissime 
truncatum, supra albido-testaceum, breviter sericeo-pubescens, vitta media cinerea 
latissima, antice inordinate et minute nigro-punctata, postice utrinque linea nigra, 



48o FAUNA HAWAIIENSIS 

valde flexuoso-dentata marginata, et punctos albos biseriatos saepe includente, no- 
tatum, utrinque, in declivitate, nigricanti-reticulatum, rarius nigrum albido-vittatum et 
maculatum, subtus regione ventrali atra utrinque vitta albida recta, marginata. Chelae 
fulvo-nitidae, antice ad basin confuse infuscatae. Partes oris nigrae vel fuscae, late 
testaceo-marginatae. Sternum inaequale, luteo-olivaceum vel albidum et fulvo- 
reticulatum, saepe in medio infuscatum et confuse plagiatum sed linea media dilutiore 
sectum. Pedes lutei, sat longe et fere aequaliter setosi, femoribus ad apicem (anticis 
late posticis, anguste) infuscatis et sex posticis annulo subbasilari obliquo et angusto 
fusco notatis, tibiis metatarsisque annulo medio parvo saepe obsoleto annuloque 
apicali fuscis notatis, tarsis apice fuscis, femoribis anticis pluriaculeatis, patellis aculeo 
exteriore munitis, tibiis intus, prope medium, aculeis binis prope apicem aculeis 
plurimis minoribus, armatis, metatarsis muticis. Tuberculum vulvae ovato-transversum, 
olivaceum et nitidum, postice plagula magna cordiformi, ad marginem anticum utrinque 
lobo parvo nigro foveolato, atque in medio unco gracili et recto, sat longo, fulvo, 
apice rufulo et nitido munitum. 

Hab. Hawaii. — Molokai. — Oahu. — Maui. — Lanai. 

A Cyclosa albisterni E. Sim., cui praesertim affinis est, differt abdomine antice 
haud tuberculato postice simpliciter turbinato. 
Plate XVI. fig. 4. 



Var. Cyclosa simplicicaiida rufescens. 

%. Long. 6 mm. Cephalothorax fulvo-testaceus parce et longe albido-pilosus. 
Abdomen fere parallelum, antice rotundum, postice haud vel vix attenuatum, parum 
productum et obtuse truncatum, supra fulvum vel rufescens, vitta foliiformi latissima 
cinerea, antice evanescente, postice tenuiter et flexuose nigro-marginata plerumque 
ornatum, interdum supra late nigricans, fulvo-testaceo-marginatum et punctatum, subtus 
vitta atra lata, in medio leviter coarctata et albido-marginata notatum. Sternum 
nigrum, interdum rufescens et antice confuse infuscatum. 

Hab. Hawaii, Kona. — Molokai. 



(6) Cyclosa xanthomelas, sp. nov. 

%. Long. 8 — 10 mm. Cephalothorax nigro-nitidus, parte thoracica late ovata, 
convexa, fovea media obtuse triquetra profunde impressa, parte cephalica abrupte 
angustiore, angusta et convexa, sulco semicirculari profundo discreta. Oculi quatuor 
medii aream antice quam postice multo latiorem occupantes, postici a sese spatio 
oculo plus triplo angustiore distantes, antici posticis paulo majores. Oculi late- 



ARACHNIDA 481 

rales a mediis sat distantes, utrinque contigui et leviter prominuli, anticus postico 
major. Abdomen oblongum, antice rotundum, postice, ultra mamillas, productum, 
leviter attenuatum sed obtusum (fere ut in C. conica Pallas), nigrum, breviter et 
parce pilosum et utrinque macula fulva maxima subtriquetra et leviter dentata 
ornatum. Chelae nitidae, ad basin fuscae vel nigrae, ad apicem fulvae. Partes oris 
nigrae, intus late luteo-marginatae. Sternum nigrum. Pedes sat longi, sat longe 
setosi, nigri, tibiis ad basin anguste luteo-annulatis, metatarsis tarsisque cunctis pallide 
luteis, metatarsis 4' paris ad apicem et saepe in medio minute fuscis, tibiis anticis 
aculeis setiformibus paucis munitis, sed metatarsis muticis. Pedes-maxillares lutei, 
femore ad apicem patellaque nigris vel fuscis. Vulvae scapum ovato-transversum 
fusco-nitidum, uncus rectus sat gracilis, niger sed apice leviter ampliatus et rufulus, 
marginem scapi paulo superans. 

Plate XVI. fig. 3. 

Hab. Molokai Mts. 



(7) Cyclosa olorina, sp. nov. 

$. Long. 3 "5 mm. Cephalothorax atro-olivaceus, nitidus, parte thoracica sat late 
ovata, convexa, fovea triquetra impressa, parte cephalica abrupte angustiore, sat 
lata, convexa, sulco semicirculari profundo postice discreta. Area oculorum quatuor 
mediorum paulo longior quam antice latior et multo latior antice quam postice, 
medii postici spatio oculo paulo angustiore a sese distantes. Oculi laterales utrinque 
contigui, a mediis sat remoti. Abdomen breviter ovatum vel subrotundum (fere 
Aranei cuctirbitini CI.), supra omnino albo-opacum praeterea atrum, in lateribus et 
postice inordinate albido-punctatum, subtus antice, pone rimam genitalem, linea trans- 
versa, dein utrinque punctis 3 vel 4, lineam longitudinalem designantibus, punctisque 
mediis binis albidis, notatum. Chelae fulvae, antice ad basin late fusco-plagiatae. 
Partes oris sternumque fusca vel nigricantia, sternum utrinque, ad marginem, maculis 
testaceis 2 vel 3 notatum. Pedes breves, sat robusti, pallide lutei, coxis leviter 
infuscatis, femoribus subtus, ante medium, annulo nigricanti angusto (in femoribus 
4' paris latiore) annuloque apicali latiore olivaceo (saepe in femoribus 4' paris lineato), 
tibiis metatarsisque annulo medio angusto annuloque apicali latiore fuscis vel olivaceis, 
saepe obsoletis, ornatis. Scapum vulvae ovato-transversum, fuscum et nitidum, utrinque 
fovea sinuosa impressum, in medio unco testaceo depresso et obtuso, marginem scapi 
paulo superante, munitum. 

Plate XVI. fig. 5. 

Hab. Hawaii, Kona. 



482 FAUNA HAWAIIENSIS 



(8) Cyclosa cucitrbihda, sp. nov. 

?. Long. 2 mm. Cephalothorax laevis et glaber, pallide luteo-testaceus, parte 
thoracica in medio minute et confuse infuscata, parte cephalica brevi et convexa, 
sulco profundo semicirculari postice discreta. Area oculorum quatuor mediorum 
circiter aeque longa ac antice lata et multo latior antice quam postice, medii postici 
spatio oculo paulo angustiore a sese distantes. Oculi laterales utrinque contigui, a 
mediis sat remoti. Abdomen breviter ovatum (fere Aranei cuairbitini CI.), supra 
omnino albo vel flavido-testaceum tenuiter et parce albo-pilosum, subtus vitta latissima, 
mamillas includente, atro-testacea notatum. Partes oris sternumque pallide fusco- 
olivacea laevia. Pedes breves, sat robusti, pallide lutei, omnino mutici. Scapum 
vulvae ovato-transversum, fuscum et nitidum, incisura media uncoque albo-nitido, 
recto, gracili, sed apice leviter incrassato obtuso et marginato, marginem scapi vix 
superante, munitum. 

Hab. Oahu. 



Araneus Clerck. 
(i) Araneiis nautictis L. Koch. 

Epeira nautica L. Koch, Aegypt. u. Abyss. Arachn. 1875, p. 17, tab. 11, f. 2. 
Epeira pullata Thorell, Stud. Rag. Mai. etc. i. 1877, p. 385 (45). 

Espece abondamment repandue dans toutes les regions tropicales, en Afrique, en 
Asie, Malaisie, et dans I'Amerique du Sud ; sans doute introduite aux lies Sandwich. 

Hab. Oahu. — Hawaii. 



(2) Araneus evwiae, sp. nov. 

%. Long. 8 — 10 mm. Cephalothorax obscure fulvo-rufescens, regione oculari 
paulum infuscata, crasse et longe albido-sericeo-pubescens. Area oculorum mediorum 
haud vel vix prominula, circiter aeque longa ac antice lata et antice quam postice 
multo latior, sed spatio inter posticos oculo vix angustiore, antici posticis majores. 
Oculi laterales a mediis late distantes et utrinque angulum frontalem occupantes, 
inter se angustissime separati, anticus postico major. Clypeus oculis mediis anticis 
vix latior. Abdomen longius quam latius, antice attenuatum et subacuminatum, 
postice longius attenuatum atque obtusum, supra cinereo-albidum, parce albido- 
crinitum, atrum, antice utrinque linea obliqua, dentata et punctata, postice vitta folii- 



ARACHNIDA 483 

formi angusta et attenuata, maculas 3 vel 4 includente, nigris ornatum, subtus regione 
ventrali atra, utrinque linea longitudinal!, postice leviter ampliata et incurva, et se- 
cundum mamillas maculis parvis binis albis, decoratum. Chelae fuscae, apice rufes- 
centes, albido-crinitae, margine superiore sulci dentibus quatuor, 3° reliquis majore, 
inferiore dentibus trinis subaequis et contiguis armatis. Partes oris fuscae, late 
testaceo-marginatae. Sternum fuscum plerumque antice paululum dilutius. Pedes, 
praesertim antici, longi, aculeis numerosis albidis subpellucentibus sed ad radicem 
minute fuscis, instructi, fulvi, sericeo-pubescentes, femoribus anticis apicem versus 
sensim infuscatis, posticis apice fusco vel nigricanti-annulatis, patellis tibiisque plus 
minus fusco-variatis, metatarsis tarsisque plerumque concoloribus, interdum valde 
nigricanti-annulatis. Vulva scapo parvo fere obtecto, postice (sub unco) plagula 
olivacea trapezoidali munito, unco testaceo transversim rugato, ad basin lato con- 
vexo et subtriquetro, ad apicem abrupte angustiore gracili et recto, munita. 

Plate XVI. fig. 6. 

Pictura abdominalis valde variabilis : 

Varietas. A. Abdomine supra omnino fusco. 

B. Abdomine fusco, linea alba subintegra longitudinaliter secto. 

C. Abdomine fusco-cervino, antice macula media longitudinali ovata, maculis 
binis ovatis et obliquis cruciata, albo-opacis, postice vitta foliiformi parum expressa 
sed punctis nigris biseriatis notata, supra ornato. 

D. Abdomine supra nigro, in parte basali macula maxima transversa albido- 
testacea obtecto. 

E. Abdomine nigro, antice macula apicali, postice maculis seriatis 2 vel 3 
albo-opacis supra decorato. Pedibus valde et late nigro-annulatis (Hawaii, Olaa). 

Hab. Hau^aii ; Kau, Olaa. — Molokai. — Lanai, Koele Mts. — Kauai, Waimea Mts. 



(3) Araneus kapio/aniae, sp. nov. 

$. Lono-. 15 mm. Cephalothorax fulvo-rufescens, parte cephalica fusca vel 
nio-ricanti, crasse sed breviter fulvo-crinitus. Area oculorum mediorum valde pro- 
minula, vix longior quam antice latior et antice quam postice multo latior, sed 
spatio inter posticos oculo baud latiore, antici posticis multo majores. Oculi laterales 
a mediis late distantes et utrinque angulum frontalem occupantes, inter se evidenter 
separati, anticus postico major. Clypeus oculis mediis anticis baud latior. Abdomen 
longius quam latius, antice breviter attenuatum et obtusum, postice longius et sensim 
attenuatum, supra cinereo-virescens, leviter fusco nigrove vermiculatum, antice in 

V. H. II. ^3 



484 FAUNA HAWAIIENSIS 

declivitate oblique nigro-marginatum, dein transversim dilutius, dein maculis trans- 
versis nigerrimis albo-limbatis, series duas (5 — 5), postice convergentes, desig- 
nantibus, anticis obliquis, reliquis recte transversis, ornatum, in lateribus valde 
nigricanti-reticulatum, subtus regione ventrali atra, utrinque vitta lutea, postice 
ampliata et incurva, marginata. Chelae nigrae, apice rufulae, parce sed valde fulvo- 
crinitae. Partes oris nigrae, intus luteo-marginatae. Sternum nigrum. Pedes, 
praesertim antici, longi, aculeis fulvis, intus in tibiis et metatarsis anticis numerosis 
brevibus et validioribus, armati, femoribus quatuor anticis fusco-rufulis, quatuor posticis 
nigricantibus, cunctis ad basin luteis, patellis fulvo-rufulis fusco-variatis, tibiis fusco- 
rufulis, annulo subbasilari annuloque subapicali angustis fulvis, metatarsis tarsisque 
nigricantibus, metatarsis annulis angustis binis, tarsis (3° excepto) annulo parvo 
subbasilari fulvis, notatis. Vulva scapo late ovato subrotundo, fulvo sed utrinque 
fusco-notato, unco testaceo longo, usque ad basin sat gracili, recto et supra trans- 
versim rugato, munita. 

Plate XVI. fig. 7. 

Hab. Hawaii, Hilo. 



Fam. THOMISIDAE. 

Subfam. MISUMENINAE. 

MisuMENA Latreille. 

Ce genre est represente aux lies Sandwich par un certain nombre d'especes qui 
peuvent se rapporter a trois groupes ou sections. 

Sectio i"" i^Mistimena oreadcs E. Sim.). 

Les caracteres du Misumeua oreades sont ambigus ; cette espece se rapproche en 
effet des Pistms par son front etroit, son groupe oculaire median un peu plus long 
que large et plus etroit en avant qu'en arriere ; d'un autre cote cependant elle se 
rattache au genre Misumeua par ses tubercules oculaires tres obtus, non saillants au 
dessus de I'oeil lateral posterieur, par son aire oculaire plane, ses teguments lisses, 
son abdomen ovale, ni anguleux ni tronque. 

Elle est en outre remarquable par son cephalothorax longuement attenue en 
avant et par son bandeau un peu proclive. 



ARACHNIDA 48 = 



(i) iMisinuena oreades, sp. nov. 

?. Long. 8 mm. Cephalothorax paulo longior quam latior, antice attenuatus, 
fronte sat angusta, laevis, glaber, luteo-testaceus, area oculorum semilunari albo-opaca. 
Oculorum lineae binae fere aequaliter recurvae. Ociili quatuor antici inter se fere 
aequidistantes, medii lateralibus saltem \ minores, quatuor postici inter se aequales, 
medii a lateralibus quam inter se paulo remotiores. Area mediorum paulo longior 
quam postice latior et antice quam postice evidenter angustior. Abdomen sat longe 
oblongum, postice leviter ampliatum, omnino albido-testaceum. Chelae, partes oris 
sternum pedesque luteo-testacea. Pedes quatuor antici reliquis multo longiores et 
robustiores, tibiis aculeis pellucentibus sat brevibus sed inter se iniquis, basin 
articulo baud attingentibus, 4 — 5 vel 3 — 4, metatarsis aculeis, paulo longioribus, 5 — 6 
vel 4 — 5, subtus instructis, sed aculeis lateralibus carentibus. 

Hab. Hawaii, Kea Mts. — Oahu, Wainae. — Molokai. 

Sectio 2^ [illistiniena nesiotcs E. Simon). 

Le Misnmena nesiotcs E. Simon, fait presque le passage des Misumeiia aux 
Diaea ; sa premiere ligne oculaire est un peu plus recurvee que la seconde, comme 
celle des Misuniena tyj^iques, mais avec les yeux lateraux visiblement plus gros que 
les medians, de chaque cote son tubercule oculaire oblique est marque, entre les yeux, 
d'une legere depression. 

(2) Misumena nesiotes E. Simon. 

M. nesiotes E. Simon, in Zool. Jahrb. xii. 1899, p. 416. 

%. Long. 7 mm. Cephalothorax subtilissime coriaceus, setis longis paucis seriatis 
conspersus, fulvo-rufescens, utrinque valde nigro-variegatus et reticulatus et postice 
macula nigra oblique notatus (parte fulva vittam latam et parallelam designante), 
tuberibus ocularibus albo-opacis. Oculi antici inter se fere aequidistantes, laterales 
mediis vix majores. Oculi postici in lineam latiorem, fere aequaliter recurvam, 
parvi et aequi, medii a lateralibus quam inter se paulo remotiores. Oculi quatuor 
medii parvi et aequales, aream subquadratam, postice quam antice vix latiorem, 
occupantes. Abdomen vix longius quam latius, postice ampliatum et obtusum, 
fulvo-testaceum, nigro-plagiatum, supra antice macula parva, dein maculis binis den- 
tatis, postice arcubus transversis, utrinque latis in medio tenuibus, subtus postice 
lineis fuscis binis punctatis et subgeminatis, parum distinctis, notatum. Sternum 
fulvum, parce nigro-setosum. Pedes quatuor antici reliquis multo longiores, obscure 

63—2 



486 FAUNA HAWAIIENSIS 

fulvi, (tarsis exceptis) crebre fusco-punctati, femoribus subtus fere nigris, tibiis sub- 
annulatis. Pedes quatuor postici lutei. Tibiae quatuor anticae aculeis mediocribus 
fusco-pellucentibus 6 — 4, metatarsi aculeis similibus 5 — 4 subtus instructi, sed aculeis 
lateralibus carentes. Vulva tuberculo fulvo parvo obtuso et subrotundo, postice 
excavate, munita. 

Var. Cephalothorax utrinque fulvo-rufulo baud nigro variegatus et punctatus. 
Abdomen albido-testaceum, postice punctis nigris grossis, maculas binas transversas 
confusas designantibus, notatum. Pedes fere omnino lutei, antici vix distincte 
fulvo-punctati. 

Hab. Hawaii, Kona. — Molokai. 



Sectio 3^ {Alisiimeiia afiguliventris E. Simon). 

Les especes de ce groupe se font remarquer par leurs yeux anterieurs a-peu-pres 
equidistants avec les lateraux plus gros que les medians, leur aire oculaire mediane 
environ aussi large que longue et plus etroite en avant qu'en arriere, enfin par 
leurs tubercules oculaires tres obtus mais saillants et, de chaque cote, un peu 
deprimes entre les yeux. 

Leur abdomen est, le plus souvent, fortement elargi et tronque en arriere 
comme celui des Thomisus ; les teguments de leur cephalothorax sont finement cha- 
grines comme ceux des Xysticus. 



(3) Misuniena nigrofrcnata, sp. nov. 

%. Long. 5 mm. Cephalothorax sublaevis, setis validis nigris paucis seriatis 
munitus, albo-opacus plus minusve pallide olivaceo-variatus, regione clypei intense 
nigra, regione thoracica postice sinuoso nigricanti-marginata. Oculi quatuor antici 
in lineam valde recurvam, inter se fere aequidistantes, medii lateralibus paulo minores. 
Oculi quatuor postici parvi aequi, inter se fere aequidistantes. Area oculorum quatuor 
mediorum multo latior postice quam antice et circiter aeque longa ac postice lata. 
Abdomen breviter ovatum, postice ampliatum et obtusum, supra albo-opacum, in 
parte apicali maculis binis, inter se appropinquatis, valde biangulosis et litteram Z 
fere designantibus, et utrinque macula marginali majore, acute excisa, nigris deco- 
ratum, subtus nigrum. Chelae antice albae, nigro-maculatae et marginatae, interdum 
nigrae apice testaceae, opacae, setis spiniformibus paucis munitae. Partes oris, 
sternum, coxae, trochanteresque nigra, laevia. Pedes albi, femoribus ad basin atque 
ad apicem minute nigro-notatis (subtus baud lineatis), tibiis quatuor anticis ad basin 
anguste ad apicem latissime nigris, metatarsis tarsisque ad apicem late nigris, tibiis 



ARACHNIDA 487 

metatarsis tarsisque quatuor posticis ad apicem nigro-annulatis ; tibiis quatuor anticis 
subtus in parte apicali aculeo exteriore unico, aculeis interioribus longioribus trinis, 
metatarsis aculeis 4 — 5 instructis. Pedes quatuor postici fere mutici. Pedes-maxil- 
lares nigri, tarso ad basin albo. 

Plate XVII. fig. I. 

Hab. Oahu, Mountains 2000 f., Kohol. 

Var. Chelae antice nigrae. Abdomen macula anteriore parva subquadrata, 
maculis binis posticis, a sese confluentibus et vittam transversam, utrinque profunde 
et acute bifidam formantibus, supra ornatum. 

Hab. Kauai ; Waimea, Halemanu. 

Var. Chelae antice nigrae. Cephalothorax supra albus, vittis duabus nigris, 
valde laciniosis, notatus. Abdomen praecedentis. Femora antica, praesertim subtus, 
valde nigro-variata et guttulata. 

Hab. Hawaii, Kau. 



(4) Misumena crefacea, sp. nov. 

$. Long. 5 mm. Cephalothorax sublaevis, setis validis nigris paucis seriatis 
munitus, albido-opacus, leviter olivaceo-tinctus, tuberibus oculorum lineaque media 
frontali albidioribus, regione clypei fusco-rufulo-variata. Oculi fere praecedentis, sed 
laterales antici mediis saltern duplo majores. Abdomen breviter ovatum, postice 
ampliatum et obtusum, supra albido-opacum, antice utrinque, in declivitate, nigricanti- 
marginatum, subtus utrinque nigro-punctatum, in medio late nigrum. Chelae testaceae, 
fusco-maculatae, setis spiniformibus paucis armatae. Partes oris, sternum, coxae 
trochanteresque nigricantia, laevia. Pedes albidi, femoribus ad basin atque ad apicem 
minute nigro-notatis et subtus linea exili nigra ornatis, tibiis metatarsis tarsisque ad 
apicem late infuscatis et nigro-punctatis et tibiis (in parte fusca) subtus nigro-lineatis ; 
tibiis anticis subtus, in parte apicali, aculeis nigris sat brevibus i — 2, metatarsis 
aculeis inferioribus, paulo longioribus 3 — 3, et utrinque, ad apicem, aculeo laterali 
minore, munitis. Pedes quatuor postici parcissime aculeati. Pedes-maxillares lutei, 
parce nigricanti-variati. 

Var. Cephalothorax vittis binis fuscis punctatis et dentatis notatus. Abdomen 
antice testaceo-cinereum et nigro-punctatum, postice albidum, macula media magna 
nigro-marginata, postice truncata, utrinque valde et acute bidentata, supra notatum. 
Femora antica valde nigricanti-punctata et maculata et subtus nigro-lineata. 

Hab. Molokai, Koele. 



488 FAUNA HAWAIIENSIS 



(5) Misume7ta anguliventris , sp. nov. 

%. Long. 6 — 7 mm. Cephalothora.x subtilissime coriaceus, parce et longe nigro- 
crinitus, obscure fuscus, prope marginem saepe fulvo-variatus, vitta media latissima 
fulva postice attenuata et truncata, lineam mediam saepe antice bifidam et utrinque 
lineolam abbreviatam fuscas includente notatus, regione clypei nigra. Oculi antici 
inter se fere aequidistantes, medii lateralibus saltem duplo minores. Oculi postici 
parvi, inter se aequales et fere aequidistantes. Area oculorum mediorum circiter 
aeque longa ac postice lata et antice quam postice vix angustior. Clypeus latus. 
Abdomen antice rotundum, postice valde ampliatum et utrinque obtuse angulosum, 
fere Thomisi, supra fuscum vel nigricans, vitta fulva laciniosa marginatum. Chelae 
nigrae, apice dilutiores et rufulae. Partes oris sternum coxaeque fusco-castanea seu 
nigra. Pedes quatuor antici fusco-rufuli, femoribus subtus et in lateribus nigricanti 
variatis et guttulatis atque in medio nigro-lineatis, patellis, tibiis metatarsisque prope 
basin fulvo-annulatis. Pedes quatuor postici fulvi, femoribus ad apicem fusco- 
annulatis. Tibiae anticae aculeis 4 — 5 vel 5 — 6, metatarsi aculeis 6 — 7 vel 7 — 7 
subtus muniti sed aculeis lateralibus carentes. Pedes postici parce et debiliter 
aculeati. Fovea genitalis parum profunda, antice posticeque margine tenui et recto 
discreta. 

t. Long. 4 mm. Abdomen minus, ovatum, antice truncatum, fuscum albido- 
guttulatum, interdum utrinque albidum et macula media maxima nigra trapeziformi 
notatum. Pedes quatuor antici multo longiores, femoribus fuscis, nigro-variegatis, 
patellis tibiis metatarsisque nigricantibus vel fuscis annulo medio lato albido ornatis. 
Pedes postici lutei, femoribus ad apicem, tibiis ad basin atque ad apicem fusco- 
annulatis. Tibiae metatarsique antici aculeis inferioribus longioribus, aculeis latera- 
libus atque in tibiis aculeis dorsalibus armati. Pedes-maxillares fusci, fulvo-variati, 
tibia patella breviore et angustiore subtus apophysi apicali parva subacuta, extus 
apophysi multo majore, antice, secundum tarsum, directa, valde compressa fere 
lamellosa, leviter sinuosa (superne excavata, inferne convexa), atque ad angulum 
superiorem abrupte aciculata, armata, tarso mediocri, ovato, bulbo parvo, ovato, 
simplici. 

Plate XVIL fig. 11. 

Hab. Hawaii ; Kau, Kona, Kilauea. — Oahu ; Wainae, Kaala. — Maui, Halea- 
kala. — Kauai. 

Var. Cephalothorax abdomenque fusca, regione oculari dilutiore. Hawaii. 

Var, Cephalothorax fere omnino fulvo-albidus, utrinque leviter fusco-reticulatus. 
Abdomen supra late albidum, plerumque fusco-marginatum et postice transversim 
fusco-reticulatum . 



ARACHNIDA 489 

Var. Abdomen fulvum, postice in declivitate, vittis transversis fuscis confusis 
notatum. Pedes fulvi, femoribus subtus ad apicem infuscatis. Hawaii, Kauai. 

Var. Abdomen rubro-castaneum, valde albo-opaco-variegatum et reticulatum. 
Pedes quatuor antici rubro-castanei, femoribus supra leviter albido-variegatis, tibiis 
annulo medio lato albo notatis. Kauai. 



(6) Mismnena velaia, sp. nov. 

?. Long. 4 — 5 mm. Praecedenti valde affinis et sulDsimilis. Cephalothorax 
fusco-nigricans, vitta media lata dilutiore, antice confusa et infuscata sed linea media 
exillima albida, oculos baud attingente, secta, postice late albida et leviter dentata. 
Abdomen supra fuscum et nigro-punctatum, utrinque nigrum, postice, in declivitate, 
albidum, antice maculis binis subrotundis, utrinque maculis similibus binis albis, 
ornatum, subtus atrum utrinque leviter dilutius. Chelae antice fuscae fulvo-variatae, 
setis validis paucis munitae. Partes oris sternumque nigro-olivacea, sternum in medio 
paulo dilutius. Coxae olivaceae. Pedes quatuor antici obscure fulvo-rufuli, crebre 
et grosse nigro-punctati, femoribus subtus nigro-lineatis, tibiis in medio dilutioribus 
parcius punctatis, metatarsis tarsisque annulo medio albido notatis. Pedes quatuor 
postici lutei, anguste nigro-annulati. Tibiae anticae aculeis inferioribus sat longis 
subpellucentibus i (vel 2) — 3, metatarsi aculeis inferioribus similibus 3 — 4 aculeoque 
laterali apicali minore, armati. Plaga genitalis plus duplo latior quam longior, fusco- 
nitida, utrinque valde attenuata et obtuse triquetra, apice minute foveolata. 

$. Long. 3 — 5 mm. Abdomen utrinque et postice albidum, macula dorsali 
maxima nigra trapeziformi ornatum. Pedes-maxillares obscure fulvi, tibia nigricanti, 
tibia patella breviore et angustiore, subtus apophysi apicali minutissima, extus 
apophysi multo majore antice, secundum tarsum, directa, compressa, apice profunde 
et inaequaliter fissa : ramulo superiore gracillimo acuto et subrecto, inferiore leviter 
divaricato fulvo et obtuso, armata, tarso mediocri, ovato, bulbo parvo, simplici. 

Plate XVH. fig. 12. 

Hab. Molokai, Koele. 

Var. B. Cephalothorax niger, vitta media lata, oculos haud attingente, antice 
fusco-rufula et linea media exili obscuriore secta, postice .sensim dilutiore lutea et 
maculis parvis binis geminatis fuscis notata, ornatus. Abdomen supra luteo-fulvum, 
crebre fusco-punctatum, macula maxima trapeziformi obscuriore postice truncata et 
utrinque nigro-bidentata notatum. Pedes quatuor antici obscure fulvo-rufuli, cre- 
berrime nigro-punctati, patellis supra, tibiis metatarsisque in medio paulo dilutioribus 
et confuse annulatis, tarsis fusco-rufulis ad basin atque ad apicem luteis. 

Hab. Maui, Haleakala. 



490 FAUNA HAWAIIENSIS 

Var. C. Cephalothorax atro-olivaceus, leviter testaceo-variatus, vitta lata dilu- 
tiore, antice fusco-variata et pone oculos lineata, postice maculam albidam V-magnum 
designantem, notatus. Abdomen supra albidum, macula magna nigra triquetra, antice 
acuta, postice valde et sensim ampliata, truncata et plerumque utrinque bidentata, 
ornatum. Pedes pallide lutei, femoribus anticis subtus confuse olivaceo-punctatis et 
parum distincte fusco-lineatis, tibiis ad apicem, metatarsis, ad basin atque ad apicem, 
tarsisque in medio late nigro-annulatis. 

Hab. Maui, Haleakala. 



DiAEA Thorell. 
Synopsis specierum. 

Femora quatuor antica subtus linea exillima nigra notata insulana. 

Femora baud lineata vitellina. 



Diaea insulana Keyserling. 

D. insulana Keyserling, in L. Koch, Ar. Austr. ii. 1889, p. 261, tab. xxiv. figs. 3 — 4. 

$. Long. 4 — 5 mm. Cephalothorax laevis, setis tenuibus paucis seriatis munitus, 
luteo-rufescens, utrinque et postice late fulvo fuscove marginatus, tuberibus ocu- 
laribus albido-cinereis opacis, clypeo saepe tenuiter nigro-marginato. Oculi antici 
in lineam leviter recurvam, medii lateralibus plus duplo minores, a sese quam a late- 
ralibus vix remotiores. Oculi postici in lineam multo latiorem, magis recurvam, 
inter se subaequales, medii a lateralibus quam inter se evidenter remotiores. Area 
quatuor mediorum circiter aeque longa ac postice lata et antice quam postice paulo 
angustior, antici posticis paulo majores. Clypeus oculis mediis anticis haud triplo 
latior. Abdomen oblongum, postice ampliatum et obtusum, pallide fulvo-testaceum, 
praesertim ad marginem albido-reticulatum, parcissime et minute fusco-punctatuin et 
plerumque postice tenuiter transversim albo-striatum, subtus nigricanti-marginatum et 
macula ventrali magna trapeziformi nigricanti notatum. Chelae, sternum pedesque 
pallide fulva, pars labialis leviter infuscata. Pedes quatuor antici reliquis multo 
longiores et robustiores, femoribus subtus linea exili intense nigra semjDer ornatis, 
antice aculeis parvis quatuor, supra aculeis similibus binis, tibiis aculeis nigris 
tenuibus et sat longis 2 — 2 (rarius 3 — 3), metatarsis aculeis 3 — 4 vel 4 — 5 (api- 
calibus minoribus) subtus armatis, sed aculeis lateralibus carentibus. Pedes quatuor 
postici minute et parce aculeati. Area genitalis antice foveola testacea transversa, 
tenuiter marginata et carinam latam convexam et cordiformem includente, impressa, 
postice plagula nigra convexa, obtuse triquetra nigra, munita. 



ARACHNIDA 491 

$. Long. 4 mm. Cephalothorax utrinque plerumque obscurior. Abdomen 
minus, sat anguste oblongum, supra albidum, crebre nigro-punctatum, in medio late 
nigricans et postice tenuiter transversim albo-striatum. Pedes quatuor antici multo 
longiores et apice graciliores, femoribus subtus plus minus distincte nigro-lineatis. 
Pedes-maxillares mediocres, obscure fulvi, tarso, praesertim ad basin, infuscato, tibia 
patella breviore et paulb angustiore, subtus ad apicem apophysi parva subrotunda, 
extus apophysi multo longiore antice, secundum tarsum, directa, apice leviter ampliata, 
rotunda sed supra calcare acuto, leviter curvato instructa, tarso ovato, bulbo simplici, 
rotundo, disciformi, stylo circumdato. 

Plate XVII. fig. 13. 

Hab. Hawaii. — Molokai. — Maui. — Kauai. 

Var. B. Abdomen in medio parce nigro-punctatum, pone medium macula 
transversa nigra, utrinque valde bidentata et plerumque transversim albido-striata, 

notatum. 

Hab. Hawaii ; Kona. 

Var. C. Abdomen fere praecedentis sed macula dorsali majore. Femora i' 
paris antice punctis nigris paucis notata. Tibiae quatuor anticae nigricanti-biannulatae. 

Hab. Hawaii ; Kona, Mauna Kea. 

Var. D. Abdomen fere praecedentis sed obscurius. Cephalothorax regione 
frontali obscuriore et utrinque vitta fusca valde dentato-laciniosa, marginem haud 
attingente, notatus. Pedum quatuor anticorum femora, tibiae metatarsique grosse et 
inordinate nigro-punctata. 

Hab. Hawaii ; Kona, Kau, Olea. 

Var. E. Abdomen obscure fulvum, macula nigra magna, marginem anticum fere 
attingente, postice medium paulo superante, sensim ampliata, truncata et utrinque 
bidentata, supra ornatum. Pedum anticorum femora infuscata, subtus nigricanti- 
vittata, tibiae nigricanti-biannulatae. 

Hab. Hawaii ; Kona, Kau. — Molokai. 

Cette espece, abondamment repandue dans tout I'archipel, est tres variable comme 
coloration, au point qu'il est difficile de rencontrer deux individus exactement sem- 
blables. Keyserling a figure une forme tres pale. 

(2) Diaea vitellina, sp. nov. 

?. Long. 2 "5 — 3 mm. Cephalothorax laevis, .setis tenuibus paucis seriatis 

munitus, fulvo-rufescens, parte cephalica, praesertim postice, dilutiore, area oculorum 

albo-opaca. Oculi antici in lineam vix recurvam, inter se fere aequidistantes, medii 

lateralibus plus triplo minores. Oculi postici in lineam recurvam multo latiorem, 

F. H. II. 64 



492 FAUNA HAWAIIENSIS 

inter se subaequales, medii a lateralibus quam inter se evidenter remotiores. Oculi 
medii aequi, aream vix longiorem quam postice latiorem et antice quam postice vix 
angustiorem occupantes. Clypeus oculis mediis anticis plus quintuple latior. Abdomen 
sat longe oblongum, pallide luteum, in parte prima vitta reticulata, marginem anticum 
baud attingente, in parte altera maculis transversis brevibus quinque uniseriatis nigris, 
supra ornatum. Chelae, sternum pedesque luteo-rufescentia. Pedes quatuor antici 
reliquis multo longiores, tibiis aculeis sat longis pellucentibus 2 — 2, metatarsis i' paris 
aculeis similibus 3 — 3 (apicalibus minoribus), metatarsis 2' paris aculeis 3 — 4 subtus 
armatis. 

^. Long. 2 '5 mm. A femina differt cephalothorace utrinque, prope marginem, 
aculeis parvis nigris paucis consperso, pedibus anticis multo longioribus, apice 
gracilioribus, femoribus patellisque usque ad basin, tibiisque ad apicem rufulo-tinctis. 
Pedes-maxillares parvi, fulvo-rufuli, tibia patella breviore, subtus ad apicem apophysi 
parva obtusa, extus apophysi multo longiore, antice secundum tarsum directa, com- 
pressa, apicem versus leviter attenuata sed obtusissime truncata cum angulo inferiore 
rotundo, superiore calcare parvo et acuto instructo, tarso ovato, bulbo simplici rotundo- 
discoidali, stylo circumdato. 

Plate XVII. fig. 14. 

Hab. Hawaii, Kau. — Oahu, Waimea. 



Synaema E. Simon. 

Le genre Synaema est represente par un groupe d'especes remarquables par 
leur bandeau etroit, ayant rarement plus de deux fois le diametre des yeux medians 
ant^rieurs, par leur abdomen ovale allonge, leurs metatarses anterieurs, generalement 
armes en dessous de plus de deux paires d'epines mais manquant d'epines laterales 
sauf les apicales, enfin par leurs pattes des deux paires posterieures mutiques a 
I'exception d'une ou de deux tres petites epines inferieures aux tibias et aux metatarses. 

Synopsis specierum. 

1. Coxae cunctae nigrae. Pedes quatuor postici lutei valde fusco-annulati naevigerum. 

Coxae quatuor anticae nigrae, quatuor posticae luteae. Pedes quatuor 

postici lutei 2. 

2. Tibiae anticae aculeis debilibus 4 — 4 vel 4 — 5, metatarsi aculeis 4 — 5 vel 

4 — 6 subtus muniti. Pedes postici omnino lutei dimidiatipes. 

Tibiae anticae aculeis 2 — 2 subtus munitae. Metatarsi postici apice anguste 

fusco-annulati 3. 

3. Metatarsi antici aculeis 5 — 5 subtus muniti .fronto. 

Metatarsi antici aculeis 2 — 2 subtus muniti impotens. 



ARACHNIDA 493 



(i) Synaema diniidiatipes, sp. nov. 

$. Long. 8 — 9 mm. Cephalothorax fere laevis, fusco-castaneus, ad marginem 
posticum paulo dilutior. Oculi quatuor antici inter se fere aequldistantes, laterales 
mediis saltem duplo majores. Oculi medii postici lateralibus vix minores, a lateralibus 
quam inter se paulo remotiores. Clypeus fere laevis, oculis mediis anticis vix duplo 
latior. Abdomen longe ovatum, antice obtuse truncatum, nigro-opacum, parce et 
minute albo-punctatum, postice maculis albis parvis, valde laciniosis et biseriatis 3 — 3, 
interdum maculis majoribus, inter se confluentibus et vittam marginalem valde den- 
tatam formantibus supra ornatum, subtus atrum, regione epigasteris dilutiore et rufula. 
Chelae latae, antice fere planae, nigro-nitidae, parce et longe nigro-crinitae. Partes 
oris, sternum coxaeque quatuor anticae, nigra, laevia. Pedes quatuor antici fusco- 
castanei, femoribus patellisque nigricantibus, femoribus anticis, in parte basali, aculeis 
longis quatuor uniseriatis munitis, tibiis aculeis inferioribus sat debilibus 4 — 4, in- 
terioribus exterioribus longioribus, metatarsis aculeis robustioribus 4 — 5 vel 4 — 6 
subtus armatis. Pedes postici omnino lutei, tibiis aculeis debilibus fere setiformibus 
I — 2 vel 2 — 2, metatarsis 3' paris aculeis submediis binis, metatarsis 4' paris aculeo 
unico subtus munitis. Fovea genitalis testacea, parva, transversa, obtuse triquetra, 
nitido-marginata. 

Plate XVII. fig. 3. 

Hawaii, Kona. 



(2) Synaema fronto, sp. nov. 

%. Long. 8 mm. Cephalothorax supra laevis, utrinque in declivitate leviter 
coriaceus, nigro-castaneus. Oculi quatuor antici in lineam parum recurvam, medii 
inter se quam a lateralibus evidenter remotiores. Oculi quatuor postici inter se fere 
aequldistantes. Area oculorum quatuor mediorum fere parallela et multo latior quam 
longior. Abdomen longe oblongum, supra atrum, utrinque, praesertim postice, obscure 
fulvo-variatum, vitta media lata albo-opaca abbreviata, in medio coarctata, ad apicem 
minute dentata, supra ornatum. Chelae, partes oris, sternumque nigra. Pedes 
quatuor antici nigro-castanei, tibiis subtus aculeis debilibus 2 — 2, metatarsis aculeis 
paulo robustioribus 5—5 subtus armatis. Pedes quatuor postici fulvi, metatarsis apice 
minute infuscatis, tibiis metatarsisque aculeis submediis parvis binis subtus munitis. 

A praecedenti, cui subsimilis est, differt imprimis area oculorum mediorum fere 

parallela et multo latiore quam longiore et tibiis quatuor anticis aculeis 2—2 tantum 

armatis. 

Hab. Oahu, Waianae Mts. 

64-2 



494 FAUNA HAWAIIENSIS 



(3) Synaema impotens, sp. nov. 

?. Long. 5 '5 — 6 mm. Cephalothorax nigro-castaneus, fere laevis. Oculi medii 
antici lateralibus plus duplo majores et inter se quam a lateralibus paulo remotiores. 
Oculi postici fere S. dimidiatipedis. Clypeus oculis mediis anticis plus duplo latior. 
Abdomen lonee obloneum, nigfrum, ad marginem anticum maculis binis geminatis, in 
parte apicali maculis parvis laciniosis triseriatis, decoratum. Chelae, sternum et partes 
oris nigro-nitida, chelae antice parce et valde crinitae. Pedes quatuor antici nigri- 
cantes, patellis tibiisque supra ad apicem anguste albo-marginatis, femoribus antice 
aculeis longis trinis uniseriatis, tibils aculeis 2 — 2 longis et tenuibus, apice setiformibus, 
metatarsis aculeis similibus 2 — 2, subtus armatis. Pedes quatuor postici lutei, meta- 
tarsis ad apicem anguste fusco-annulatis, tibiis aculeo medio, metatarsis 3' paris aculeo 
medio aculeoque apicali, metatarsis 4' paris aculeo medio, subtus munitis. 

Hab. Hawaii, Kona. 



(4) Synaema naevigerum, sp. nov. 

?. Long. 5 — 6 mm. Cephalothorax subtilissime coriaceus, niger, tuberibus ocu- 
laribus minute rufulo-notatis. Oculi medii antici lateralibus plus duplo minores, inter 
se quam a lateralibus vix remotiores. Oculi postici inter se fere aequidistantes, medii 
lateralibus vix minores. Clypeus leviter coriaceo-rugatus, oculis mediis anticis plus 
duplo latior. Abdomen longe oblongum, nigro-opacum, antice maculis binis geminatis 
utrinque et postice punctis inordinatis, interdum confluentibus et vittam marginalem 
dentatam formantibus, albis, supra decoratum. Chelae, partes oris, sternum coxaeque 
cunctae nigra et laevia. Pedes quatuor antici nigro-nitidi, patellis tibiisque ad 
apicem albo-marginatis, tarsis ad basin anguste rufulis, femoribus antice aculeis 
3 vel 4 uniseriatis, tibiis aculeis exterioribus binis, interioribus longioribus 3 vel 4, 
metatarsis aculeis paulo robustioribus 3 — 3 subtus armatis. Pedes quatuor postici 
lutei, articulis cunctis ad apicem tibiisque ad basin fusco nigrove annulatis, tibiis 3' 
paris aculeis debilibus binis, tibiis 4' paris aculeo unico, metatarsis aculeo simili 
submedio subtus armatis. Fovea genitalis testacea, parva, ovato transversa, nitido- 
marginata. 

$. Long. 4 mm. A femina differt abdomine minore, pedibus anticis longioribus, 
aculeis tibiarum et metatarsorum tenuibus fere setiformibus. Pedes-maxillares parvi, 
fusci ; patella convexa ; tibia patella paulo breviore et angustiore, subtus ad apicem 
apophysi nigra minutissima et truncata, extus apophysi longa, antice, secundum tarsum 



ARACHNID A ' 495 

directa, extiis convexa. apice abrupte angustiore et acutissima ; tarso breviter ovato, 
supra convexo ; bulbo simplici, rotundo, discoidali. 
Plate XVII. fig. 2. 

Var. Abdomen supra nigrum, minute et parum regulariter albo-punctatum et 
vitta latissima alba, postice dentata, omnino cinctum (Molokai). 

Var. Abdomen nigerrimum, vitta lata albo-opaca, in parte secunda utrinque 
valde bilobata et apice acuminata, supra ornatum. 

Hab. Hawaii ; Kau, Kona. — Molokai. 



Mecaphesa, gen. nov. 

Cephalothorax pauIo longior quam latior, parte cephalica antice leviter acclivi 
fronte mediocri. Oculi postici inter se subaequales, in lineam valde recurvam, medii 
a lateralibus quam inter se evidenter remotiores. Oculi antici in lineam multo angus- 
tiorem, fere aequaliter recurvam, inter se fere aequidistantes, medii lateralibus saltern 
duplo minores. Area quatuor mediorum parallela, paulo longior quam latior. Clypeus 
angustus. Tegumenta coriacea, setis simplicibus acutis paucis instructa. Caetera fere 
Oxyptilae. 

Ab Oxyptila praesertim differt oculorum linea antica postica aequaliter recurva, 
ab Heriaeo differt oculorum linea postica multo magis recurva, clypeo angusto, 
tegumentis parcius et brevius setosis. 



(i) Mecaphesa cincta, sp. nov. 

%. (pullus) long. 3 mm. Cephalothorax paulo longior quam latior, utrinque 
ample rotundus, antice valde attenuatus, fronte haud declivi sat angusta et truncata, 
crebre coriaceus et setis validis paucis seriatis munitus, niger, utrinque parce et vix 
distincte rufulo-variegatus. Oculi antici in lineam angustam valde recurvam, medii 
lateralibus plus duplo minores, inter se quam a lateralibus vix remotiores. Oculi 
postici in lineam latiorem valde (fere aequaliter) recurvam, medii lateralibus vix 
minores, a lateralibus quam inter se fere \ remotiores. Oculi medii inter se aequi, 
aream parallelam, longiorem quam latiorem, occupantes. Clypeus oculis mediis anticis 
vix duplo latior, ad marginem setis spiniformibus fulvis uniseriatis instructus. Ab- 
domen antice rotundum, postice valde ampliatum et obtuse truncatum, supra atrum, 
postice in declivitate transversim fulvo-striatum, vitta marginali lata albida dentata 
et parce nigro-punctata omnino cinctum, subtus fulvo-testaceum. Chelae nigrae, 
apice luteae, subtiliter coriaceae et parce crinitae. Partes oris sternumque nigra. 
Coxae quatuor anticae fusco-olivaceae, quatuor posticae luteae. Pedes lutei, quatuor 



496 FAUNA HAWAIIENSIS 

antici valde nigricanti-punctati et maculati, tibiis in parte apicali nigricantibus, 
metatarsis fuscis sed prope basin fulvo-annulatis. Pedes quatuor postici articulis 
cunctis, tarsis exceptis, apice anguste sed intense nigro-annulatis. Tibiae anticae 
aculeis fulvis longis sed iniquis, suberectis 4 — 4 vel 5 — 5, metatarsi aculeis similibus 
(apicalibus minoribus) 5 — 5 subtus instructi. Pedes postici fere mutici. 

Hab. Maui, Haleakala. 



(2) Mecaphesa seniispinosa, sp. nov. 

%. Long. 4 mm. Cephalothorax subtilissime coriaceus, fulvo-rufescens, iitrinque 
fuscus et leviter fulvo-variatus, linea media exillima alba sectus, tuberibus ocularibus 
obscure cinereis, parte cephalica antice levissime acclivi. Oculi antici in lineam sat 
angustam, valde recurvam, inter se fere aequidistantes, laterales mediis fere triplo 
majores. Oculi postici inter se aequi, in lineam multo latiorem, valde (fere aequaliter) 
recurvam, medii a lateralibus quam inter se fere \ remotiores. Oculi medii inter se 
subaequales (postici anticis vix majores), aream parallelam, longiorem quam latiorem 
occupantes. Clypeus oculis mediis anticis saltern duplo latior, ad marginem setis 
uniseriatis munitus. Abdomen antice rotundum, postice valde ampliatum et obtuse 
truncatum, omnino fusco-rufescens. Chelae coriaceae, partes oris sternumque fusco- 
rufescentia. Pedes fulvo-rufuli, antici posticis paulo obscuriores, tibiis aculeis fulvis 
validis et suberectis 3 — 3, partem apicalem articuli tantum occupantibus, metatarsis 
aculeis similibus 3 — 3 (apicalibus minoribus) subtus instructis. Pedes postici fere 
mutici. Plaga genitalis fulvo-nitida, convexa, obtuse triangularis, fovea subrotunda, 
crasse marginata et plagulam parvam longam includente, impressa. 

Plate XVII. fig. 4. 

Hab. Hawaii, Mauna Kea. 



Subfam. PHILODROMINAE 
Proerneae. 

Les especes assez nombreuses qui representent aux lies Hawaii la sous-famille 
des P hilodrominae se distinguent toutes des Philodronms ordinaires par leurs cheliceres, 
dont la marge sup^rieure au lieu d'etre transverse et arrondie est longuement oblique, 
garnie d'une serie tres dense de longs crins incurves et armee de deux dents coniques 
(la i"^ plus forte que la 2") eloignees de la base du crochet qui est plus long que celui 
des Philodronms. Ces caracteres indiquent que ces especes font le passage des 
Thomisides aux Chibionides, particulierement au groupe des Sparassus. 

Nous avons deja indique (Zool. Jahrb. 1899, p. 418) les rapports des Proermis 



ARACHNIDA 497 

{Plerelas) et des Psellomis et nous avions meme propose de les rapprocher dans un 
meme groupe, mais la decouverte de types nouveaux {Pagiopalus, Adrastidia) faisant 
graduellement le passage des Proernns aux Pkilodromus nous fait penser aujourd'hui 
que ces araignees se rattachent plus intimement a la sous-famille des P hilodrominae 
dans laquelle elles doivent cependant former un groupe special. 

Les Proernus et genres voisins different des Psellomis par leurs pieces buccales 
beaucoup plus courtes, construites comme celles des Thomisides ordinaires, par leur 
cephalothorax plus ou moins epais et leurs cheliceres semblables dans les deux sexes, 
jamais divergentes. 



Proernus, nom. nov. 
Pterelas E. Simon, in Zool. Jahrb,, 1899, P- 4'^ (nom. praeocc.^). 

Synopsis specierum. 

1. Metatarsi quatuor antici usque ad basin crebre scopulati. Tibia pedum- 

maxillarium maris apophysi parva, apice minute et fere aequaliter bidentata, 

armata 2. 

Metatarsi quatuor antici, apice excepto, baud scopulati. Tibia pedum- 
maxillarium maris apophysi parva, apice inaequaliter bidentata, dente 
superiore inferiore multo minora, armata velox. 

2. Metatarsi quatuor antici aculeis basilaribus trinis tantum muniti 3. 

Metatarsi quatuor antici aculeis inferioribus 3 — 3 usque ad apicem armati aculeatus. 

3. Tibiae 2' paris subtus, usque ad basin, aculeis parvis 3 — 3 armatae. Tibiae 

metatarsique postici aculeati Schauinslandi. 

Tibiae 2' paris subtus, in parte apicali, aculeis interioribus binis aculeoque 
exteriore unico tantum munitae. Metatarsi postici aculeati sed tibiae 
posticae muticae castaneus. 

(i) Proernus schauinslandi E. Simon. 

Pterelas schauinslandi E. Simon, in Zool. Jahrb., 1899, p. 418. 

^. Long. 12 mm. Cephalothorax fulvo-rufescens, parte cephalica antice sensim 
infuscata, postice lineolis abbreviatis fuscis trinis, media recta alteris divaricatis, notata, 
breviter fulvo-pubescens et setis nigris parvis spiniformibus conspersus. Abdomen sat 
breve, antice obtuse truncatum, postice leviter ampliatum et obtusum, supra nigricans, 
antice paulo dilutius et confuse testaceo-variatum, crebre et breviter fulvo-pubescens, 
subtus dilutius et cinereo-albido-pilosum. Chelae fusco-rufulae, attenuatae et parum 
convexae, antice opacae, transversim rugatae et setis spiniformibus inordinatis con- 
spersae. Partes oris, sternum pedesque fulvo-rufescentia, patellis tibiis metatarsisque 

' Pterelas, Guerin, in Mag. Zool. vi. 1836, cl. 7, pi. 20. 



498 FAUNA HAWAIIENSIS 

quatuor anticis fusco-castaneis. Pedes i' paris tibiis subtus aculeis brevibus 3 — 3 
atque extus aculeis lateralibus trinis minoribus, metatarsis aculeis binis inferioribus 
aculeoque laterali exteriore subbasilaribus, armatis. Pedes 2' paris subtus similiter 
aculeati, sed aculeis lateralibus carentes. Pedes quatuor postici minute et parce 
aculeati. Pedes-maxillares mediocres, fulvi, patella intus tarsoque obscurioribus ; tibia 
patella paulo longiore et paulo graciliore, tereti, extus apophysi apicali nigra parva et 
obtusa, sed apice minutissime bidentata, armata ; tarso mediocri, ovato, tibia vix 
longiore ; bulbo parvo et ovato, simplici. 

Cephalothorax long. 5-3 mm., lat. 5-5 mm. Abdomen long. 7 mm. Pedes i 
long. I9'8 mm.; Pedes 11 long. 25 mm.; Pedes in et iv long. 14 mm. 

Plate XVII. fig. 6. 

Hab. Oahu : Koolau range (Perkins, x. 1892). 



(2) Proernus aciileatus, sp. nov. 

^. Long. 12 mm. Cephalothorax pallide fulvo-testaceus, parte cephalica antice 
sensim infuscata et rufescenti-tincta. Abdomen sat longum, depressum, antice obtuse 
truncatum et leviter emarginatum, supra punctis impressis longis et obliquis 3 — 3 
munitum, fulvo-testaceum, sericeo-pubescens, postice utrinque puncto nigro minutissimo 
notatum. Chelae fusco-castaneae, transversim leviter rugatae et setis validis iniquis 
conspersae. Partes oris fusco-castaneae, laminae intus albido-marginatae. Sternum 
pedesque fulva. Pedes i' paris tibiis subtus aculeis 3 — 3, e.xterioribus parvis interioribus 
majoribus, extus aculeis lateralibus trinis parvis uniseriatis atque intus aculeo laterali 
subapicali, metatarsis aculeis inferioribus 3 — 3 aculeoque laterali exteriore subbasilari 
armatis. Pedes 2' paris anticis multo longiores, tibiis aculeis inferioribus minutissimis 
3 — 3, metatarsis aculeis paulo majoribus 3 — 3 munitis, sed aculeis lateralibus carentibus. 
Pedes quatuor postici parce aculeati. Pedes-maxillares ut in praecedenti. 

Hab. Kauai, Koholuamano. 

(3) Proernus castajieus, sp. nov. 

t. Long. 12 mm. Cephalothorax obscure fulvo-rufescens, regione frontali in- 
fuscata, parte cephalica postice vittis obliquis fuscis marginata, fulvo-pubescens et setis 
validis nigris conspersus. Abdomen sat breve, antice obtuse truncatum, postice 
obtusum, nigricans sed antice et subtus paulo dilutius, breviter fulvo-pubescens, punctis 
impressis coriaceis obliquis 3 — 3, supra notatum. Chelae fulvo-rufulae ad basin con- 
vexae et leviter geniculatae, vix rugatae, setis nigris validis conspersae. Partes oris 
sternumque fusca. Pedes obscure fulvo-ravidi, patellis tibiis metatarsisque quatuor 
anticis fusco-castaneis. Pedes i' paris tibiis subtus aculeis sat brevibus 3 — 3 atque 



ARACHNIDA 499 

extus aculeis lateralibus minoribus binis, altero submedio altero subapicali, metatarsis 
usque ad basin crebre scopulatis, aculeis binis inferioribus aculeoque laterali exteriore 
subbasilaribus armatis. Pedes 2' paris in parte apical! aculeis parvis exterioribus binis, 
aculeoque interiore unico subtus munitis, metatarsis ad basin biaculeatis. Pedes quatuor 
postici metatarsis aculeatis sed tibiis muticis. Pedes-maxillares fere ut in praecedentibus 
sed tibia paulo breviore. 

Hab. Maui, Haleakala. 



(4) Proe7-mis velox, sp. no v. 

Z. Long. 8 mm. Cephalothorax obscure fulvo-rufescens, linea marginali lineo- 
lisque radiantibus obscurioribus notatus, breviter sed crasse flavido-pubescens et setis 
validis et longis nigris conspersus. Abdomen oblongum, antice obtuse truncatum, 
postice breviter attenuatum et obtusum, punctis impressis 3 — 3, mediis longis et obliquis, 
notatum, supra obscure testaceum, utrinque crebre nigricanti-punctatum et confuse 
marginatum, breviter et crasse fulvo-rufulo-pubescens, subtus dilutius et luteo-pilosum. 
Chelae sat debiles, fusco-rufulae, apice dilutiores, vix rugatae, setis nigris conspersae. 
Partes oris, sternum pedesque fulvo-rufula, parte labiali, sterno antice, patellis tibiis 
anticis metatarsisque ad apicem leviter infuscatis. Pedes i' paris tibia subtus aculeis 
gracilibus et longis 3 — 3, aculeo laterali interiore subapicali, aculeisque lateralibus 
exterioribus multo minoribus trinis, metatarsis gracilibus et longis, apice excepto, baud 
scopulatis, aculeis inferioribus longis 3 — 3, aculeisque lateralibus exterioribus minoribus 
binis, armatis. Pedes 2' paris anticis multo longiores, subtus similiter aculeati, sed tibiis 
utrinque aculeo laterali parvo tantum munitis. Pedes quatuor postici tibiis metatarsisque 
aculeatis. Pedes-maxillares fere praecedentium sed paulo longiores, apophysi tibiali 
subtus ad basin leviter emarginata, ad apicem inaequaliter bidentata, dente superiore 
inferiore multo minore. 

Plate XVII. fig. 5. 

Hab. Maui, Haleakala. 



Pagiopalus, gen. nov. 

A Procrno differt fronte angustiore, oculis quatuor anticis in lineam magis re- 
curvam angustiorem, mediis lateralibus majoribus et inter se quam a lateralibus 
remotioribus, a lateralibus anguste separatis, oculis mediis posticis a sese quam a 
lateralibus distantioribus, sed area quatuor mediorum, antice quam postice angustiore, 
paulo longiore quam latiore vel saltem baud latiore, pedum anticorum metatarsis 
tarsisque tenuibus, aculeis inferioribus lateralibusque longioribus et numerosioribus 
armatis, tarsis breviter et parce scopulatis, metatarsis baud scopulatis. 

F. H. II. ^5 



500 FAUNA HAWAIIENSIS 



Synopsis specierum. 

Cephalothorax fulvus, nigro-punctatus et lineatus 2. 

Cephalothorax fulvus, nigro-variegatus, macula magna vel vitta albo-opaca 

supra ornatus 3. 

Tibiae i' paris aculeis 4 — 4 subtus armatae. Plaga genitalis feminae area 

parva acute triquetra postice impressa. Pedes-maxillares maris longis- 

simi atomarhis. 

Tibiae t' paris aculeis 3 — 3 subtus armatae. Plaga genitalis feminae area 

parva ovata postice impressa. Pedes-maxillares maris modice longi semipiinctatiis. 

Sternum venterque pallide lutea .persona/us. 

Sternum venterque atra nigriventrts. 



(i) Pagiopalus atomarius, sp. nov. 

%. Long. 7 — 8 mm. $. Long. 6 mm. — Cephalothorax obscure fulvus, luteo- 
nitido-pubescens, fusco nigrove punctatus et lineatus, regione oculari et clypei in- 
fuscata, parte cephalica seriatim punctata, Hneis longitudinalibus trinis, media recta, 
alteris exillimis et sinuosis, postice macula angulosa V magnum formante, parte 
thoracica punctis parvis, in medio lineas radiantes designantibus, versus marginem 
majoribus densioribus et subconfluentibus, notatis. Oculi antici lineam sat angustam 
recurvam designantes, medii lateralibus circiter \ majores et a sese quam a lateralibus 
plus duplo remotiores (spatio inter laterales et medios oculo angustiore). Oculi 
postici in lineam multo latiorem, modice recurvam, medii lateralibus paulo minores et 
a sese quain a lateralibus circiter \ remotiores. Abdomen depressiusculum, oblongum, 
antice obtuse truncatum et minute emarginatum, obscure testaceum, luteo-nitido- 
pubescens et crebre nigricanti-punctatum, prope medium maculis geminatis binis, ad 
apicem macula foliiformi confusa nigricantibus, notatum, subtus dilutius, minutissime 
et parcissime nigro-punctatum. Chelae luteae, antice maculis nigricantibus binis 
magnis, basali in medio dilutiore, notatae. Partes oris leviter infuscatae. Sternum 
pedesque lutea. Pedes, coxis saepe exceptis, crebre nigricanti-punctati. Pedes i' 
paris tibia subtus aculeis gracilibus et longis 4 — 4, utrinque aculeis paulo brevioribus 
trinis, aculeisque dorsalibus binis, metatarso aculeis inferioribus longis 3 — 3 et 
utrinque aculeis trinis (apicalibus minoribus) munitis. Pedes 2' paris anticis multo 
longiores, fere similiter aculeati, sed tibiis aculeis 5 — 5 vel 6 — 6 subtus munitis. 
Pedes quatuor postici numerose aculeati. 

%. Pedes-maxillares sat breves, lutei, nigro-punctati et subannulati. Plaga geni- 
talis nigra rugosa, postice laevior et rotunda, striis binis, antice convergentibus, 
aream minutam, longe triquetram limitantibus, notata. 

Z. Pedes-maxillares lutei, longissimi ; femore recto, cephalothorace multo longiore ; 



ARACHNIDA 501 

tibia tereti subrecta, patella multo longiore, ad apicem apophysi exteriore parva et 
truncata apophysique inferiore cariniformi, antice subacuta, armata ; tarso brevi et 
angusto, bulbo simplici. 

Plate XVII. fig. 7. 

Hab. Hawaii, Kona. — Molokai, Koloe. — Kauai. 



(2) Pagiopalus scmipunctatits, sp. nov. 

t ?. Long. 5 mm. Cephalothorax circiter aeque longus ac latus. utrinque ample 
rotundus, fulvo-rufescens, in medio minute et parce nigro-punctatus et macula media 
parva triquetra notatus, ad marginem valde nigro-reticulatus et clathratus, clypeo in 
medio et area oculari utrinque nigris. Oculi fere praecedentis, sed medii antici latera- 
libus vix majores. Abdomen breve, antice obtuse truncatum et minute emarginatum, 
postice valde ampliatum atque obtusum, supra pallide testaceum minute et inordinate 
nigro-punctatum et late nigro-marginatum, antice macula parva, postice macula majore 
foliiformi confusa nigricantibus notatum, subtus minutissime et parcissime nigro- 
punctatum et macula postica nigra notatum. Chelae luteae, antice maculis nigris 
binis dentatis, basali in medio dilutiore, notatae. Partes oris, sternum pedesque lutea, 
pars labialis leviter infuscata, sternum ad marginem minutissime nigro-notatum. Pedes 
parce nigro-punctati et valde nigro-subannulati. Tibiae i' paris aculeis inferioribus 
gracilibus et longis 3 — 3 et utrinque aculeis lateralibus minoribus binis munitae sed 
aculeo dorsali carentes, metatarsi aculeis inferioribus 3 — 3 longis (apicalibus mino- 
ribus) aculeisque lateralibus muniti. Tibiae metatarsique 2' paris multo longiores, 
aculeis inferioribus 4 — 4 muniti. 

$. Pedes-maxillares lutei, femore intus, tibia metatarsoque supra ad basin nigro- 
notatis. Plaga genitalis antice nigra et coriacea, postice dilutior et plagula parva 
fulva ovata praedita. 

t. Pedes-maxillares lutei, femore intus tibiaque ad basin nigro-punctatis, modice 
longi, femore compresso, tibia patella longiore tereti, extus ad apicem apophysi brevi, 
valde compressa haud attenuata, apice recte secta et subtus ad basin minute dentata 
armata, torso sat longe ovato, bulbo simplici. 

Plate XVII. fig. 15. 

Hab. Maui, Haleakala. 

(3) Pagiopalus personatus, sp. nov. 

%. Long. 5 mm. Cephalothorax latior quam longior, utrinque ample rotundus, 
obscure testaceus et breviter albo-pilosus, sed in medio late albo-opacus, regione 
oculari nigra linea alba divisa, clypeo nigro, regione thoracica utrinque nigricanti 

65 — 2 



502 FAUNA HAWAIIENSIS 

sed ad marginem albo-notata, in medio lineis radiantibus nigricantibus notata. Oculi 
quatuor antici in lineam recurvam, medii lateralibus evidenter majores, a sese quam 
a lateralibus plus duplo remotiores (spatio inter laterales et inedios oculo multo 
minore). Oculi postici in lineam multo latiorem, modice recurvam, inter se sub- 
aequales, medii a sese quam a lateralibus vix \ remotiores. Abdomen antice obtuse 
truncatum et emarginatum, postice valde ampliatum, supra nigrum, antice atque in 
medio late et confuse albido-vittatum, postice arcubus tenuibus transversis albis 
seriatis, i° longo, reliquis parvis, ornatum, subtus albidum et albo-pubescens. Chelae 
antice nigrae, prope medium luteo-maculatae. Partes oris sternum pedesque lutea, 
parte labiali infuscata, laminis extus fusco-notatis, sterno anguste nigricanti-marginato, 
pedibus valde punctatis et late nigro-annulatis. Pedes i' paris tibia subtus aculeis 
gracilibus et longis 4 — 4, utrinque aculeis trinis aculeoque dorsali, metatarso aculeis 
inferioribus similibus 2 — 2 et utrinque aculeis trinis, apicalibus minoribus, munitis. 
Pedes 2' paris anticis multo longiores, fere similiter aculeati, sed tibia aculeis in- 
ferioribus 5 — 5 et metatarso aculeis inferioribus 3 — 3 armatis. Plaga genitalis fusca, 
pilosa, subrotunda, postice plagula transversa angulosa, glabra, notata. 

Plate XVI 1 1, fig. 8. 

Hab. Lanai, Koele Mts. 



(4) Pagiopalus nigriveittris, sp. nov. 

$. Long. 5 mm. Cephalothorax latior quam longior, utrinque ample rotundus, 
obscure fulvo-rufescens et breviter albo-pilosus, utrinque valde nigro-reticulatus sed 
ad marginem anguste albo-cinctus, vitta media lata, oculos fere attingente, leviter 
dentata, antice punctos nigros, lineam transversam valde flexuosam designantes et in 
medio maculam nigram subrotundam includente notatus, regione clypei et oculorum 
nigra sed inter oculos posticos paulo dilutiore et nigro-lineata. Oculi fere praece- 
dentis. Abdomen sat longe oblongum, antice rotundum et vix emarginatum, postice 
leviter et sensim ampliatum, supra nigrum, albo fulvoque pubescens, ad marginem 
anticum maculis binis confusis, prope medium utrinque vitta marginali abbreviata 
arcuata et flexuosa albidis et albo-pilosis ornatum, subtus atrum cinereo-pubescens, 
mamillis testaceis. Chelae antice nigrae, in lateribus et subtus testaceae. Partes 
oris luteae, laminae extus nigfro-notatae. Sternum niero-nitidum, ad marginem an- 
ticum testaceo-variatum. Pedes lutei, femoribus parce tibiis metatarsisque crebre 
nigro-punctatis et subannulatis, aculeis gracilibus et longis, ut in praecedenti ordi- 
natis, armati. Plaga genitalis fusca, pilosa, leviter depressa, multo longior quam 
latior, antice attenuata et obtusa, postice truncata et minute emarginata. 

Hab. Hawaii, Kau. 



ARACHNIDA 503 



Adrastidia, gfen. nov. 

A Proerno differt oculis mediis anticis lateralibus minoribus et a lateralibus quam 
inter se remotioribus, sed mediis posticis a sese quam a lateralibus duplo vel triple 
distantioribus, area oculorum mediorum multo latiore postice quam antice et multo 
latiore quam longiore, abdomine sat longo, antice emarginato, postice sensim ampliato, 
pedum anticorum tarsis gracilibus, breviter scopulatis, metatarsis tantum ad apicem 
leviter scopulatis. 

(i) Adrastidia stigniaiica, sp. nov. 

?. Long. 7 — 8 mm. Cephalothora.x obscure fulvus, breviter et crasse albo-pilosus, 
in medio parce in lateribus crebre nigro-punctatus, punctis densioribus lineas radiantes 
designantibus, parte cephalica in medio lineolis trinis abbreviatis et postice macula 
magna triquetra crebre nigro-punctata, ornata, tuberibus ocularibus albidis. Abdomen 
multo longius quam latius, antice sat angustum et obtuse emarginatum, postice sensim 
ampliatum, elevatum et obtuse truncatum, obscure fulvum, albo-pubescens, minute et 
crebre nigro-punctatum, antice linea media fusca lanceolata, postice, ad apicem, maculis 
parvis nigris, plerumque quatuor, transversim uniseriatis, notatum, subtus pallide 
testaceum et albo-pilosum. Chelae luteo-testaceae, antice ad basin nigro-punctatae. 
Partes oris sternumque lutea, parte labiali leviter infuscata, sterno tenuiter nigro-mar- 
ginato et plerumque in medio minute nigro-punctato. Pedes fulvi, antici crebre et fere 
uniformiter nigro-punctati, postici parcius punctati, tibia metatarsoque 3' paris plerumque 
supra nigro-maculatis. Tibia i' paris subtus aculeis robustis 4 — 5 et e.xtus aculeis 
parvis binis, metatarsus aculeis inferioribus robustis 3 — 3, aculeis lateralibus exterioribus 
trinis aculeoque laterali interiore apicali, armati. Tibia metatarsusque 2' paris subtus 
similiter aculeati sed aculeis lateralibus carentes. Pedes quatuor postici parcissime 
aculeati. Pedes-maxillares lutei, femore ad apicem patellaque nigro-punctatis. 

Plate XVII. fig. 10. 

Hab. Hawaii, Kona. — Lanai, Koele Mts. — ^Maui, Haleakala. — Kauai. 

Var. Abdomen laete rufescens, punctis parvis nigris albo-areolatis crebre ornatum, 
macula media ovata et postice vittis transversis binis nigris supra notatum. 

Hab. Oahu ; Kiwailou Riv., Kaala Mts. 

(2) Adrastidia nebulosa, sp. nov. 

%. Long. 5 mm. Cephalothorax obscure fulvus, parte cephalica antice infuscata, 
linea media exili, utrinque linea exili sinuosa vix expressa et postice macula magna 
acute triquetra fuscis notata, parte thoracica utrinque valde infuscata et lineis fuscis 



504 FAUNA HAWAIIENSIS 

abbreviatis radiantibus notata, tuberibus ocularibus fuscis. Abdomen fere praecedentis, 
fulvo-testaceum, supra confuse infuscatum et lividuni, in medio parcissime utrinque 
densius fusco-punctatum, postice in declivitate, punctis lineas obliquas parum regulares 
designantibus. Chelae, partes oris sternumque luteo-testacea, chelae antice ad basin 
fusco-maculatae. Pedes pallide fulvi, femoribus subtus crebre fusco-punctatis, supra 
nigricanti-variegatis et confuse triannulatis, tibiis metatarsisque, praesertim posticis, 
crebre fusco-punctatis, ut in praecedenti aculeati. Pedes-maxillares pallide lutei, femore 
ad apicem patellaque fuscis. 

Hab. Hawaii, Olaa. — Maui, Haleakala. 



{3) Adrastidia longula, sp. nov. 

%. Long. 8 mm. Cephalothorax fulvo-testaceus albo-pubescens, parte cephalica 
linea media lineisque lateralibus valde flexuosis et ramosis, postice convergentibus, 
notata, parte thoracica utrinque ad marginem valde fusco-reticulata et lineis fuscis 
exillimis et abbreviatis radiantibus, ornata, tuberibus ocularibus albis. Abdomen longe 
oblongum, antice profunde emarginatum, postice leviter et sensim ampliatum, nee 
angulosum nee truncatum, supra fulvo-lividum, fuseo nigroque guttulatum, albo- 
marginatum et maculis paucis albis, vittam mediam abbreviatam designantibus, supra 
ornatum, subtus albido-testaceum. Chelae, partes oris, sternum pedesque pallide fulva, 
chelae antice fuseo-maculatae, pedes supra vix distincte pallide fusco-punctati ; tibiis 
metatarsisque i' paris subtus aeuleis 3 — 3 et extus aculeis lateralibus minoribus binis, 
tibiis metatarsisque 2' paris subtus similiter aculeatis sed aculeis lateralibus carentibus. 

Plate XVII. fig. 9. 

A praecedentibus differt area oculorum mediorum minus transversa, vix duplo 
latiore quam longiore, oculis mediis posticis a sese minus distantibus. 

Hab. IMaui, Haleakala. 



Fam. CLUBIONIDAE. 

Heteropoda Latreille. 

(i) Heteropoda regia Fabricius. 
H. venatoria auct. (ex Linn.). 

Hab. Hawaii, Kona. — Oahu, Wainae Alts. — Kauai. 
Espece repandue dans toutes les regions tropicales du monde. 



ARACHNIDA 505 

Fam. AGELENIDAE. 
Tegenaria Latreille. 

(i) Tegenaria do))iestica Clerck. 

Araneus domestiais Clerck, Aran. Siiec, 1757, p. 102 (ad max. part.). 

Tegenaria civilis Walckenaer. 

Tegenaria Dchranii Thorell (ex Scopoli). 

Hab. Hawaii. 

Espece cosmopolite. 

Nota. Les deux families des Clubionides et des Agelenides, qui occupent ordi- 
nairement une si grande place dans la nature, paraissent etrangeres a I'archipel des 
Sandwich ; les deux especes ubiquistes citees plus haut, y ont sans doute 6te introduites. 

Fam. LYCOSIDAE. 
Lycosa Latreille. 

Ce genre est represente par quatre especes appartenant chacune a un groupe 
special : L. oahuensis Keyserl. (Z,. stygialis E. Sim.), au groupe Alopecosa (type 
L. fabrilis CI.), L. Iiawaiiensis E. Sim., au groupe du L. I'adiata Latreille, L. 
Kabikanai E. Sim., au groupe Trocliosa (type L. rnricola de Geer), enfin L. 
Likelikeae E. Sim. a un groupe particulier de caracteres ambigus. 

Deux de ces especes, L. Kalnkanai et Likelikeae E. S. sont particulieres a Kauai, 
les deux autres sont plus largement distributes, le L. oahuensis Keyserl. a meme ^t6 
trouve a Laysan, en dehors de I'Archipel. 

(i) Lycosa oahuensis Keyserl ing. 

L. oahuensis Keyserling, in L. Koch, Arachn. Austr., 11, 1889, p. 267, tab. xxiv, 
fig. 8. 

L. stygialis E. Simon, in Ann. Soc. ent. Belg., t. xin, 1S98, p. 29. 

Hab. Oahu. — Hawaii. — Laysan. 

Decrit de I'ile Oahu par Keyserling ; nous le connaissons aussi de Hawaii (Museum 
de Paris), et de Laysan (Schauinsland). M. Perkins n'en a capture qu'un seul individu 
dans les Montagnes de Waimea (Oahu). 



5o6 FAUNA HAWAIIENSIS 



(2) Lycosa hawaiiensis E. Simon. 

L. hawaiiensis E. Simon, in Zool. Jahrb., 1899, p. 419. 
Hab. Hawaii. — Maui. 

Decouvert a Hawaii (Kilauea) par le Dr Schauinsland, retrouve en nombre par 
M. Perkins a Olaa et Kona (Hawaii) et a Haleakala (Maui). 

Cette espece ressemble beaucoup au L. oahuensis Keyserl. i^stygialis E. Sim.) ; 
elle appartient cependant a un autre groupe, celui ayant pour type L. radiata Latr., ses 
cheliceres offrant a la marge inferieure trois fortes dents subcontigues. 



(3) Lycosa [T^-oc/iosa) kaiukanai, sp. nov. 

?. Long. 1 5 mm. Cephalothorax ovatus, humilis, fusco-rufescens, omnino crebre 
fulvo-cervino-pubescens, facie obtusissima. Oculi quatuor antici inter se aequidistantes, 
medii lateralibus paulo majores, in lineam subrectam vi.x procurvam, latiorem quam 
lineam 2^"". Oculi sen i""' mediocres, spatio oculo paulo minore a sese distantes. Oculi 
quatuor postici, superne visi, aream multo latiorem quam longiorem et postice quam 
antice fere duplo latiorem occupantes. Clypeus oculis parvis anticis duplo latior. 
Abdomen oblongum, omnino crebre fulvo-cervino-pubescens. Chelae validae, convexae, 
nigrae, fulvo-cinereo-crinitae, margine inferiore sulci dentibus trinis subcontiguis, ultimo 
minore, armato. Partes oris nigricantes. Sternum pedesque fulvo-rufescentia, meta- 
tarsis tarsisque anticis leviter infuscatis. Pedes robusti, quatuor antici patellis muticis, 
tibiis subtus aculeis mediocribus 3 — 3 (aculeis apicalibus carentibus) aculeoque parvo 
laterali interiore submedio armatis, metatarsis usque ad basin crebre scopulatis, aculeis 
pronis 3 — 3, apicalibus parvis, subtus munitis. Fovea genitalis multo longior quam 
latior, antice leviter attenuata et obtusa, postice truncata, rufulo-marginata, carina Integra 
canaliculata, antice angusta et attenuata, postice abrupte ampliata et trapezoidali, 
omnino divisa. 

Plate XVHI. fig. 4. 

Hab. Kauai, Halemanu. 



(4) Lycosa likclikeae, sp. nov. 

%. Long. 25 mm. Cephalothorax ovatus, parte cephalica leviter convexa, sat 
angusta, fronte truncata, obscure fulvo-rufescens, supra flavido utrinque, praesertim 
antice, albidius pubescens, parte thoracica vittis radiantibus abbreviatis, extus ampliatis, 
marginem baud attingentibus, fuscis, notata. Oculi quatuor antici in lineam procurvam 



ARACHNIDA 507 

angustiorem quam oculos maximos sen 2^^, medii lateralibus saltern i majores et a 
lateralibus quam inter se paulo remotiores. Oculi ser. 2^= maximi, spatio oculo duplo 
angustiore a sese distantes. Oculi quatuor postici, superne visi, aream latiorem quam 
longiorem et postice quam antice latiorem, occupantes. Clypeus oculis anticis haud 
latior. Abdomen longe oblongum, postice leviter ampliatum, fulvo-testaceum, flavido- 
cervino-pubescens, supra parce fusco-punctatum et in parte basali maculis fuscis parvis, 
parum regulariter quadriseriatis, notatum, subtus lineolis binis subgeminatis pallide 
tuscis, abbreviatis et parum expressis, munitum. Chelae robustae et convexae, nigrae, 
crebre et valde sordide albido-crinitae, subtus ad marginem interiorem crebre pilosae, 
praeterea glabrae, in medio transversim profunde et regulariter striatae, margine 
superiore sulci dentibus binis, 1° altero majore, margine inferiore dentibus trinis validis 
aequis et contiguis, instructis. Partes oris castaneae. Sternum obscure fulvo- 
olivaceum. Pedes sat longi, fulvo-rufuli, versus extremitates sensim infuscati, supra 
leviter fusco-variati et parce flavido-pilosi, subannulati ; tibiis anticis subtus aculeis 
longis pronis 3 — 3 apicalibusque minoribus, intus aculeis lateralibus binis armatis, meta- 
tarsis cum tarsis usque ad basin sat crebre scopulatis, aculeis longis et pronis 2 — 2 
aculeoque medio apicali subtus armatis. Pedes postici numerose et valde aculeati, tarsis 
subtus omnino crebre setosis sed utrinque linea tenui scopulata marginatis. Fovea 
genitalis- multo longior quam latior, parallela, antice rotunda, postice truncata, rufulo 
marginata et carina rufula, antice parallela, postice sat abrupte ampliata et triquetra, 
omnino secta. 

Plate XVIII. fig. 3. 

Hab. Kauai, Koholuamano. 

Cette espece remarquable pourrait former un groupe special dans le genre Lycosa, 
caractdrise par les yeux de la seconde ligne tres gros et rapproches I'un de I'autre, leur 
intervalle ayant a peine leur rayon, par les yeux anterieurs en ligne assez procurvee 
avec les medians plus gros et un peu plus resserres que les lateraux. A part cela 
elle se rapproche surtout des especes du premier groupe (type L. tarciitu/a Rossi). 



Lycosella Thorell. 

(1) Lycosella spiiiipcs, sp. nov. 

%. Long. 6 mm. Cephalothorax laevis, fusco-olivaceus, parce fulvo-rufulo-pilosus, 
regione oculari utrinque late nigro- marginata, vitta media latissima, postice sensim 
attenuata, vittaque submarginali angusta dilutioribus, obscure fulvis et albido-luteo- 
pilosis decoratus. Oculi quatuor antici in lineam valde procurvam semicircularem, 
inter se aequi et fere aequidistantes, laterales ab oculis maximis ser. 2'''= vix separati. 
F. H. II. ^ 



5o8 FAUNA HAWAIIENSIS 

Area oculorum quatuor dorsalium parallela et subquadrata. Abdomen ovatum, 
nigricans, obscure rufulo-pubescens, linea media integra fulva, luteo-pilosa et, pone 
medium, macula maxima, ovato-transversa albo-pilosa, laete decoratum, subtus fulvo- 
rufulo-pubescens. Chelae nigricantes, intus atque ad apicem rufescentes, parce fulvo- 
crinitae, margine inferiore sulci dentibus trinis subcontiguis, \° et 2° inter se acquis, 
altero minutissimo, armato. Pars labialis nigricans. Laminae-maxillares, sternum, 
pedes-maxillares pedesque fulvo-rufula, femoribus, praesertim subtus, annulis nigricanti- 
bus binis, valde sinuoso-dentatis, ornatis, tibiis metatarsisque cunctis latissime fusco- 
biannulatis, tibiis anticis aculeis inferioribus tenuibus longissimis 5 — 5 (apicalibus 
reliquis minoribus), metatarsis aculeis inferioribus similibus 3 — 3 et utrinque aculeis 
lateralibus minoribus binis (exterioribus setiformibus) armatis. Plaga genitalis fusco- 
nitida semicircularis, incisura profunda, processum linguiformem fulvum includente, 
notata. 

Plate XVI. fig. 9 and Plate XVIII. fig. 5. 

Hab. Kauai, Koholuamano. — Maui, Haleakala. 



(2) Lycosella annulata, sp. nov. • 

?. Long. 5 mm. Cephalothorax niger, vitta submarginali angusta et sinuosa 
atque in parte thoracica linea media paulo dilutioribus et rufescentibus, vix expressis, 
notatus, pilis brevibus rubris albidisque mixtis, sat crebre vestitus. Oculi antici In 
lineam validissime procurvam semicircularem, medii lateralibus vix majores. Area 
oculorum quatuor dorsalium parallela (postice quam antice vix latior) et subquadrata. 
Abdomen oblongum, fuscum, fulvo-rufulo-pubescens, parce et longe albido-crinitum, in 
parte basali nigro-marginatum, prope medium vitta transversa lata, postice vitta trans- 
versa angustiore et leviter sinuosa, nigris, supra ornatum. Chelae nigrae, intus atque 
ad apicem dilutiores et rufescentes, parce fulvo-rufulo-crinitae, margine inferiore sulci 
dentibus trinis, 1° et 2° inter se acquis, altero minutissimo, armato. Partes oris 
sternumque fusco-nigricantia. Pedes fulvo-rufuli, parce lutco-pilosi, femoribus annulis 
nigris trinis valde sinuosis et interruptis ornatis, tibiis cunctis metatarsisque posticis 
late fusco-biannulatis, tibiis anticis aculeis inferioribus 4 — 4, intcrioribus (praesertim 
mediis) longissimis, aculeoque laterali interiore parvo, metatarsis aculeis similibus 
(apicalibus minoribus) 3 — 3 subtus armatis, tibiis metatarsisque 2" paris subtus similiter 
aculeatis, sed utrinque aculeo laterali parvo munitis. Vulva tuberculo conico, obtuso 
nitido, nigro sed apice rufulo, munita. 

Plate XVI. fig. 10. 

Hab. Oahu, Honolulu. 



ARACHNIDA 509 



Syroloma, gen. nov. 

A Trabaea, cui affinis est, differt cephalothoracis facie humiliore et obtusiore, 
oculorum linea antica recurva, latiore quam linea secunda, oculis mediis a lateralibus 
quam inter se plus duplo remotioribus, oculis ser. 2"' paulo minoribus, tarsis pedum 
fascicule parvo setarum claviformium sub unguibus munitis. 

Ce nouveau genre, propre aux lies Sandwich, se rapproche du genre Trabaea par 
les yeux medians de la premiere ligne beaucoup plus rapproches I'un de I'autre que des 
lateraux, mais il s'en eloigne par le front tres obtus et arrondi comme celui des Trochosa, 
par la premiere ligne oculaire plus large que la seconde dont les yeux sont relativement 
plus petits, enfin par la structure des tarses qui est exceptionnelle pour la famille des 
Lycosides ; ces tarses offrent en effet, au dessous des griffes superieures, de petits fascicules 
de polls spathules analogues a ceux des Clubionides et masquant la griffe inferieure. 

Le genre Syroloina peut etre considere comme formant un lien entre les Lycosides 
et certains Clubionides, particulierement les Ctenus et les Mitwga. 



( I ) Syroloma major, sp. nov. 

Z. Long. 13 mm. Cephalothorax fusco-castaneus fere niger, parte cephalica 
postice late dilutiore et rufescenti, sed punctis parvis fuscis 4 vel 6 biseriatis notata, 
utrinque fere glaber, sed linea marginali exili fulvo-pilosa cinctus, supra vitta media 
latissima crebre albido-cinereo-pilosa ornatus. Abdomen ovatum, fusco-testaceum, 
fulvo-sericeo-pubescens et nigro-crinitum, in parte basali maculis parvis binis sub- 
geminatis, dein lineolis transversis tenuibus, postice utrinque macula parva nigricantibus, 
notatum. Chelae validae, ad basin convexae, nigrae, parce fulvo-pilosae, longe nigro- 
crinitae, margine superiore sulci dentibus trinis, medio reliquis duplo majore, inferiore 
dentibus trinis, ultimo reliquis minore, armatis. Partes oris nigrae, laminae intus vix 
dilutiores, pars labialis longior quam latior, apice leviter attenuata et truncata. Sternum 
sat anguste ovatum, fusco-testaceum, parce pilosum et crinitum. Pedes fulvo-rufuli, 
fulvo-sericeo-pilosi, longe et tenuiter hirsuti, annulis fuscis angustis et sinuosis cincti, 
tibiis metatarsisque anticis valde infuscatis, longe et numerose aculeati, tibiis anticis 
subtus aculeis 4—4 aculeoque laterali interiore, metatarsis subtus aculeis 3 — 3, apicalibus 
reliquis minoribus et utrinque aculeis lateralibus binis (altero submedio, altero apicali), 
armatis. Pedes-maxillares rufuli, tarso, saltem ad basin, nigricanti, mediocres et 
graciles ; femore leviter curvato supra ad apicem aculeis 1—3 armato ; patella longiore 
quam latiore ; tibia patella circiter aequilonga, vix graciliore, leviter curvata et longe 

66—2 



5IO FAUNA HAWAII ENS IS 

crinita ; tarso tibia cum patella simul sumptis breviore, tibia non multo latiore, apice 
acuminato et subtus longe crinito ; bulbo nigro, subrotundo, parum complicato. 

$. Long. 17 — 18 mm. A mari differt abdomine majore, sat longe ovato, supra 
nigricanti sed vitta media latissima, leviter sinuosa et postice utrinque acute coarctata 
rufescenti et crebre albido-cinereo-pilosa, ornato, subtus obscure fulvo-testaceo, pedibus 
brevioribus. Vulva fulvo-nitida, semicircularis, fovea subrotunda, antice carinulam 
postice plagulam parvam trapezoidalem includente, impressa. 

Plate XVI II. fig. I. 

Hab. Hawaii, Kona. — Kauai, Halemanu. 



(2) Syroloma minor, sp. no v. 

Z. Long. 8 — 9 mm. Cephalothorax nigricans, in medio vix dilutior, parce fulvo- 
pilosus, linea marginali fulvo-pilosa cinctus, parte cephalica maculis mediis binis 
obliquis, parte thoracica linea longitudinali abbreviata albidius pilosis, ornatus. 
Abdomen nigricans postice dilutius, rufulo-variatum et fulvo-cinereo-pubescens, antice 
linea longitudinali, postice punctis biseriatis albo-pilosis, supra ornatum. Chelae 
robustae, nigrae, ad basin et extus fulvo-pilosae, et, praesertim intus, valde nigro- 
crinitae, marginibus sulci ut in praecedenti. Partes oris nigrae, laminae ad apicem 
saepe dilutiores et rufescentes. Sternum nigro-nitidum. Pedes fulvo-rufescentes, fusco 
vel castaneo-annulati, femoribus annulis trinis angustis et sinuosis (altero submedio 
alteris binis subgeminatis subapicalibus), tibiis annulo basali angusto apicalique latiore, 
metatarsis annulo medio annuloque apicali, notatis, tibiis anticis aculeis inferioribus 3 — 4 
vel 4 — 4 longis, sed apicalibus minoribus et utrinque aculeis lateralibus plurimis, meta- 
tarsis aculeis inferioribus 3 — 3 et utrinque lateralibus binis armatis. Pedes-maxillares 
fere praecedentis sed tarso paulo latiore, fulvo-rufescentes, femore biannulato, tarso 
fusco-castaneo vel nigro. 

$. Long. 8 mm. Cephalothorax obscure fuscus vel nigricans in medio dilutior, 
fulvo-pubescens, linea marginali vittaque media lata sed postice attenuata albidius pilosis, 
ornatus. Abdomen oblongum, atrum, antice plus minus fulvo-variatum, postice abrupte 
dilutius et fulvum, albido fulvoque pubescens, antice linea longitudinali alba sed postice 
sensim rufulo-pilosa, prope medium lineolis transversis arcuatis albido-pilosis, supra 
ornatum. Sternum nigricans. Pedes ut in mari sed breviores. 

Plate XVIII. fig. 2. 

Hab. Molokai. — Maui, Haleakala. — Kaui, Halemanu. 



ARACHNJDA 511 

Fam. ATT I DAE. 
Bavia E. Simon. 

(1) Bavia acriccps E. Simon. 

Bavia aericeps E. Simon, in Ann. Soc. ent. Fr,, 1877, p. 61. 
Acompse suavis L. Koch, An Austr., p. 1 146, tab. xcix, fig. 6, 7. 

Hab. Oahu, Kaala Mts. 

Espece repandue dans toute la Malaisie, I'Australie et la Polynesie. 



Hasarius E. Simon. 

(i) Hasariits adansoni Audouin. 

Attus Adansoni et tardigradus Aud. in Savigny, Descr. de I'Egypte, r" ed. .xxii, 

p. 169, 170, tab. vii, f. 8 — f. 13. 
Salticus striatits Lucas, in Rev. Mag. ZooL, 1853, p. 521. 
Attus capita Lucas in Barker Webb et Berthelot, H. N. Canaries, An. Art., 

p. 27, tab. VII, fig. 8. 
Salticus citus O. P. Cambr., in Zoologist, 1863, p. 8561. 
} Salticus ruficapilius Dolesch., in Act. Soc. Ind.-Neerl., v. 1859, p. 13, tab. 11, 

%• 3- 
Attus nigrofusctts Vinson, Aran. Reun. Maur. Madag., 1864, p. 59, tab. x, fig. 8. 

Eris niveipalpis Gerstaecker, in C. Decken's Reisen in Ost-Afrika, iii, 2, 1873, 

?• 477- 
Salticus scadellatus Butler, in Ann. Mag. Nat. Hist., 1876, p. 441. 
? Salticus acutus Blackwall, in Proc. R. Irish Acad., 2. sen, in, p. 3, tab. i, f. 2. 
Plexippus ardelio Thorell, Studi Rag. Mai. etc., i, 1877, p. 602 (262). 
Hasarius Adansoni Thorell, id., iv, 1892, p. 426. 
'^.Hasarius Garctti Keyserling in L. Koch, An Austn, 1882, p. 1289, tab. ex, 

fig- 4- 

Hab. Hawaii, Kona. — Oahu, Honolulu. 

Espece repandue dans toutes les regions chaudes de I'Ancien Monde ; introduite 
aux Sandwich. 



5X2 FAUNA HAWAIIENSIS 

Plexippus C. Koch. 
(i) Plexippus payktdli Audouin. 

Attics Paykulli Aud. in Savigny, Descr. de I'Egypte, 2" ed. xxii, p. 172, tab. vii, 

f. 22. 
Atttis ligo Walck., H. N. Apt. i, 1837, p. 426. 

Plexippus ligo C. Koch, Arachn. xiii, 1846, p. 107, fig. 1168 — 1169. 
Salticus Vaillanti Lucas, Expl. sc. Alg., Zool. i, 1848, p. 136, tab. v, fig. 2. 
Alius africanus Vinson, Aran. Reun. Maur. Madag., 1864, p. 52, tab. x, fig. 3. 
Euophrys delibuta L. Koch, in Verh. z. b. G. Wien, 1865, p. 874. 
Hasarius Paykulli E. Sim., An Fr. in, 1876, p. 79. 
Menemerus Paykulli Thorell, St. Rag. Mai. etc. iii, 1881, p. 501. 
Plexippus Paykulli Thorell, id., iv, 1892, p. 372. 

Hab. Kauai, Makaweli 2000 ft. vi. 1894, Perkins. 

Espece repandue dans toutes les regions chaudes du monde ; introduite aux 
Sandwich. 

MoLLiCA G. et E. Peckham. 

(1) Mollica microphthalmus L. Koch. 

Jotus niicrophthalnius L. Koch, Arachn. Austr., i, p. 1246, tab. cvii, f. 2 — 3. 
Hab. Oahu, Honolulu. 
Decrit de Tahiti ; crouve depuis dans plusieurs iles de la Polynesie. 

Sandalodes Keyserling. 

Mopsus Keyserl. in L. Koch, Ar. Austr. i, 1882, p. 1330 (non Mopsus Karsch). 

Sandalodes Keyserl, loc. cit., p. 1476 (nota). 

Thyene E. Simon, in Bull. Soc. Zool. Fr., x, 1885 [T. seinicuprea). 

Les trois especes precedentes ont sans doute ete introduites aux Sandwich; les 
Attides autochtones rentrent tons dans le genre Sandalodes, propose par Keyser- 
ling pour deux especes australiennes, qu'il avait d'abord rapportees a tort au genre 
Afopsus de Karsch. Mais les Sandalodes des Sandwich sont plus petits que les 
especes typiques, ils manquent chez le male de crete frontale, et les caracteres du 
genre s'y affaiblissent graduellement, depuis les 5". pubens et validus qui ressemblent 
encore au 6". albobarbatits Keyserl. jusqu'au S. cruciatus E. Sim. qui a le facies d'un 
Icius. 



ARACHNIDA 513 



Synopsis specierum. 

1. Chelae antice, praesertim ad basin, creberrime et inordinate fulvo alboque 

barbatae 2. 

Chelae antice pilis longis albis seriatis, lineas designantibus, ornatae 3. 

2. Pili oculorum albi albociliatus. 

Pili oculorum rubri vel fulvcrufuli .piibens, canosus, validus. 

3. Pili oculorum albi seninilus. 

Pili oculorum fulvo-nifuli verecundus, /lavatiis, cruciatus. 



( I ) Sandalodes piibejis, sp. nov. 

t. 7 — 8 mm. Cephalothorax crassus, leviter ovatus, antice utrinque, pone 
oculos laterales, convexus, niger, pilis simplicibus crassis, longis et pronis, fulvo-ravidis 
crebre vestitus, parte thoracica vittis dorsalibus latis binis lineaque marginali exili 
albido-luteo-pilosis ornata. Pili oculorum fulvo-ravidi. Clypeus fere glaber, oculis 
mediis saltem duplo angustior. Oculi quatuor antici in lineam sat recurvam, laterales 
a mediis anguste separati. Area oculorum dorsalium evidenter latior postice quam 
antice. Oculi parvi ser. 2" ab oculis posticis quam ab anticis saltem \ remotiores. 
Abdomen oblongum, supra ravido-pubescens, parce et longissime albo nigroque 
crinitum, antice vitta marginali lineisque medianis binis arcuatis, postice linea media 
exili abbreviata et utrinque vittis obliquis binis latis albido-pilosis ornatum, subtus 
fuscum, breviter et parce luteo-pilosum. Chelae robustae, rugosae et transversim 
rugatae, nigrae, crasse et dense fulvo-ravido-pilosae et intus longe albo-crinitae, sed 
apice subglabrae, marginibus sulci longis, superiore dentibus binis subgeminatis remotis, 
1° 2° duplo majore, inferiore dente mediocri, armatis, ungue longo, ad basin crasso. 
Partes oris nigrae, laminae ad apicem paulo dilutiores et rufescenti-tinctae, extus ad 
angulum leviter dilatatae et convexae sed obtusae. Sternum luscum. Pedes albo- 
pubescentes, longe nigro alboque hirsuti, fulvo-ravidi, femoribus, praesertim anticis, 
valde infuscatis fere nigris, metatarsis tarsisque dilutioribus ; pedes i' paris reliquis 
robustiores, femore compresso, tibia subtus aculeis mediocribus 3 — 3, interioribus 
exterioribus paulo majoribus, metatarso aculeis similibus 2 — -2 munitis, sed aculeis 
lateralibus carentibus ; pedes 2' paris tibia aculeis inferioribus 3 — 3 aculeoque laterali 
interiore subapicali armata, patella intus uniaculeata ; pedes quatuor postici valde 
aculeati, inter se longitudine subaequales (pedes 3' paris, saltem femore et patella, paulo 
longiores). Pedes-maxillares fulvo-rufuli, graciles et longi ; femore curvato ; patella 
tereti, plus duplo longiore quam latiore ; tibia patella longiore, graciliore et tereti, 
utrinque, praesertim intus, longissime nigro-crinita, extus ad apicem apophysi nigra, 
simplici et subrecta, antice, secundum tarsum, directa armata ; tarso tibia breviore, 
anguste ovato, apice attenuate sed obtuso ; bulbo simplici disciformi, intus stylo gracili, 
subrecto, marginato. 



514 FAUNA HAWAIIENSIS 

%. Long. 8 mm. Cephalothorax niger, parte cephalica fulvo-cinereo-pubescenti, 
subsquamulata, utrinque rubro-pilosa, parte thoracica subglabra sed linea marginali exili, 
lineisque dorsalibus binis, postice divaricatis, albo-pilosis, ornata. Pili oculorum rufuli. 
Clypeus angustus, pilosus. Abdomen longe oblongum, nigrum, supra fulvo-cinereo 
leviter micanti pubescens, antice arcu marginali, postice utrinque vitta lata, intus 
valde dentata, albo-pilosis, decoratum, subtus dilutius et parce albo-pilosum, sed vitta 
media lata nigfra subglabra, notatum. Chelae robustae, nio-rae, ad basin crasse albo- 
luteo-pilosae, ad apicem subglabrae. Partes oris nigrae, apice rufulae. Sternum 
fuscum. Pedes albo nigroque hirsuti, obscure castanei vel nigricantes, coxis sex 
posticis, metatarsis tarsisque dilutioribus. Area genitalis leviter depressa, plagula 
media nigra parva, transversim triquetra et bifoveolata munita. 

Plate XVI II. fig. 7. 

H.\B. Hawaii ; Kona, Kau, INIauna Loa. — Molokai. 

(2) Sandalodcs validus, sp. nov. 

^. Long. 8 mm. A 5". pubenti. cui valde affinis est, differt cephalothorace 
antice et utrinque rubro-pubescenti, linea marginali exili et postice vittis latis binis 
divaricatis luteo-albido-pilosis decorato, pilis oculorum obscure rubris (non fulvis), 
chelis antice valde coriaceis, crebre flavo-pilosis et hirsutis, pedum 2' paris tibia subtus 
aculeis exterioribus trinis aculeoque interiore apicali atque intus aculeis lateralibus binis 
munita. Caetera fere S. pubentis. 

Plate XVIII. fig. 6. 

Hab. Hawaii, Olaa. 

(3) Sandalodcs albociliatus, sp. nov. 

^. Long. 6 — 7 mm. A 6". pubenti, cui affinis est, differt cephalothorace nigro 
fulvo-cinereo-pubescenti, parte thoracica in declivitate sensim glabra sed in medio lineis 
binis abbreviatis et divaricatis albo-pilosis ornata, oculis anticis in lineam minus recur- 
vam, pilis albis cinctis, clypeo glabro, abdomine atro, supra crebre ravido-pubescenti 
atque antice arcu albo marginato, subtus fere nudo, chelis nigris, glabris, subtiliter 
coriaceis et opacis, pedibus fulvo-rufulis, femoribus, praesertim anticis, valde infuscatis 
fere nigris, pedibus 1' paris patella intus minute uniaculeata, tibia aculeis inferioribus 
3 — 3 aculeoque laterali interiore subapicall, tibia 2'' subtus tantum aculeis i — 3 sed 
aculeis lateralibus interioribus binis, munitis. Pedes-maxillares fere S. pubentis sed 
femore nigro. 

Plate XVIII. fig. 8. 

Hab. Lanai, Koele Mts. 



ARACHNID A 5,5 

(4) Sandalodes canosus, sp. nov. 

?. Long. 8 mm. Cephalothorax modice altus, ovatus, antice posticeque atten- 
uatus, niger, parte cephalica parteque thoracica in medio, pills albis pronis depressis et 
leviter ovatis crebre vestitis, atque in medio setis longissimis albis erectis paucis et 
mordinatis, munitis, utrinque in declivitate fulvo-rufulo sed versus marginem sensim 
albidius pubescens. Pili oculorum obscure rubri. Clypeus glaber. Oculi antici in 
Imeam sat recurvam, laterales a mediis distincte separati. Abdomen atrum, supra 
cmereo-nitido-pubescens et setis longissimis albis conspersum, antice sensim albidius et 
utrinque vittis obliquis abbreviatis albis notatum, subtus vitta media latissima nip-ra et 

o 

subglabra, notatum. Chelae robustae, convexae, nigrae, rugosae et rugatae, antice 
creberrime albido-luteo-pilosae, sed apice glabrae. Partes oris sternumque nigra. 
Pedes robusti, albo-pilosi et longissime hirsuti, nigri, antici tarsis dilutioribus, postici 
coxis metatarsis ad basin tarsisque castaneis, aculeis, ut in 5. piibenti ordinatis, armati. 
Area genitalis postice marginata, utrinque foveola ovata impressa. 

Plate XVIII. fig. 13. 

Hab. Maui, Haleakala. 



(5) Sandalodes navafus, sp. nov. 

t. Long. 6-5 mm. Cephalothorax niger, sublaevis, parte cephalica haud vel 
vix distincte tuberculata, in medio crebre albido-cinereo vel fulvo-nitido pilosa, antice, 
prope oculos, vitta longitudinali albidiore sed confusa, notata, utrinque fulvo-rufulo vel 
cervino pilosa, parte thoracica vitta media latissima, postice abbreviata et furcata, 
lineaque marginali exili albo-pilosis notata. Pili oculorum fulvo-rufuli, inter oculos 
medios albidi. Clypeus in medio parce luteo-barbatus, utrinque in genis fulvo-rufulo- 
pilosus. Abdomen oblongum, nigricans, vitta longitudinali dilutiore, antice confusa, 
postice ampliata et arcubus transversis tenuibus fuscis secta, notatum, rufulo-pubescens 
sed antice arcu marginali, dein vitta longitudinali lata et postice vitta transversa, saepe 
confusa, albo-pilosis decoratum, subtus parce albo-pilosum. Chelae sat longae et 
parallelae, atrae, coriaceo-opaceae, pilis longis albis, lineas longitudinales designantibus, 
antice ornatae, margine inferiore sulci dente valido armato. Partes oris sternumque 
nigra. Pedes albo nigroque hirsuti, antici nigri, metatarsis ad basin tarsisque fulvis, 
postici nigricantes, coxis trochanteribus femoribus ad basin, metatarsis ad basin, 
tarsisque fulvis, tibiis 4' paris annulo medio fulvo saepe notatis. Pedes i' paris ut in 
6". puhenti aculeati. Pedes 2' paris tibia aculeis exterioribus trinis aculeoque interiore 
apicali (rarius aculeis binis) subtus armata et intus aculeis lateralibus binis munita, 
patella intus minute uniaculeata. Pedes-maxillares ut in praecedentibus sed fusco- 
castanei, parce albo-pilosi. 

F. H. II. 6^ 



5i6 FAUNA HAWAIIENSIS 

%. Long. 6 mm. Cephalothorax niger, longe cinereo-albido-pubescens, parte 
cephalica antice fulvo-pilosa, sed vitta media albidiore, oculos attingente, notata, parte 
thoracica postice area triquetra glabra munita. Pili oculorum fulvi. Pili clypei 
densissimi et longi, albidi. Chelae parce et longe albido-crinitae. Abdomen longe 
oblongum, cinereo-albido-pubescens, utrinque fusco-variatum, postice arcubus parvis 
fuscis seriatis notatum. Pedes fere maris sed breviores et paulo dilutiores. 

Plate XVI II. fig. 9. 

Hab. Hawaii, Kona. — Lanai, Mt. Koele. — Maui, Haleakala. — Kaui, Waimea. 

(6) Sandalodes verectmdus, sp. nov. 

$. Long. 4 — 5 mm. Cephalothorax niger opacus, in medio pallide luteo utrinque 
fulvo-rufulo-pubescens, parte thoracica postice vittis binis divaricatis abbreviatis albidius 
pilosis, ornata. Pili oculorum fulvo-rufuli, inter oculos lutei. Clypeus crebre luteo- 
barbatus. Pars cephalica postice, inter oculos, tuberculo humillimo vix distincto 
munita. Area oculorum dorsalium vix latior postice quam antice et postice cephalo- 
thorace vix angustior. Abdomen oblongum, obscure testaceum, antice vitta longitudi- 
nal! angusta lanceolata, utrinque zonis obliquis latissimis trinis subcontiguis, fusco- 
violaceis, supra notatum, luteo rufuloque pubescens. Chelae angustae et parallelae, 
atrae, valde coriaceo-opaceae, pilis luteis longis subseriatis ornatae, margine inferiore 
sulci dente valido armato. Partes oris nigricantes. Sternum fuscum. Pedes i' paris 
fusco-castanei, femore compresso fere nigro, metatarso tarsoque dilutioribus, tibia 
subtus aculeis mediocribus 3 — 3 aculeoque laterali interiore subapicali, metatarso 
aculeis longioribus 2 — 2 munitis. Reliqui pedes fulvi, temoribus, tibiis metatarsisque 
apice leviter infuscatis, tibia 2' paris subtus aculeis exterioribus trinis aculeoque interiore 
parvo apicali et intus aculeo laterali armata, tibiis metatarsisque posticis numerose 
aculeatis. Pedes-maxillares fulvo-rufuli, graciles, fere ut in S. pubeiiti. 

Plate XVI II. fig. 10. 

Hab. Oahu, Kawailoa River. 

(7) Sandalodes cniciaiits, sp. nov. 

$. Long. 4 mm. Cephalothorax niger, sublaevis, pilis longis depressis et squami- 
formibus albis supra crebre vestitus, utrinque parcius fulvo-pilosus sed linea marginal! 
exili albo-pilosa cinctus, parte thoracica vittis albis binis abbreviatis et divaricatis 
notata, parte cephalica postice tuberculo humillimo vix expresso notata. Pili ocu- 
lorum fulvi. Pili clypei pauci lutei. Abdomen oblongum, nigrum, antice arcu mar- 
ginal!, dein vitta media lata, parallela vel postice attenuata et dentata et plerumque 
postice vitta transversa cruciata, niveo-pilosis supra decoratum, subtus parce albo- 
pilosum. Chelae parallelae, atrae, coriaceo-opaceae, pilis luteis longis, subseriatis. 



ARACHNIDA 517 

ornatae, margine inferiore sulci dente valido armato. Partes oris sternumque nigra. 
Pedes nigro alboque hirsuti. Pedes i' paris nigricantes, metatarso ad basin, tarsoque 
fulvis. Reliqui pedes obscure fulvi, fusco-annulati, femoribus, basi excepto, fere nigris, 
aculeis, ut in praecedenti ordinatis, armati. Pedes-maxillares fere ut in S. vereamdo 
sed nigri, tarso parvo apice dilutiore. 

Plate XVIII. fig. II. 

Hab. Hawaii, Kona. 

(8) Sandalodes seniculus, sp. nov. 

^. Long. 4 mm. Cephalothorax niger, sublaevis, pilis longis depressis (squami- 
formibus) pronis albis supra crebre vestitus, utrinque subglaber sed linea marginali 
exili alba cinctus, parte cephalica utrinque et antice fulvo-pilosa sed antice vitta media 
alba notata, postice tuberculo medio humillimo, vix perspicuo, notata. Pili oculorum 
supra oculos rufuli, inter oculos et subtus albi. Clypeus in medio parce utrinque in 
genis crebre albo-barbatus. Abdomen oblongum, atrum, in medio confuse dilutius et 
postice arcubus transversis fuscis notatum, sed omnino crebre et crasse albo-pubescens. 
Chelae, partes oris, sternum pedesque praecedentis sed metatarsis anticis fulvis. 
Pedes-maxillares paulo breviores, fusco-castanei. 

Plate XVIII. fig. 12. 

Hab. Maui, Haleakala. 

Ordo SCORPIONES. 

IsoMETRUS Hempr. et Ehrbg. 

( 1 ) Isometrus macuialus de Geer. 

Hab. Hawaii. — Maui. — Oahu. 

Espece repandue dans toutes les regions chaudes du monde ; introduite aux 
Sandwich; deja indiquee de Honolulu par Karsch et par nous-meme de Maui. 

Ordo CHERNETES. 

Fam. CHELIFERIDAE. 

Chelifek Geoffroy. 

(i) Chclifer bifissus E. Simon. 

C. bifissus E. Simon, in Ann. Soc. ent. Belg., xliii 1899, p. 121. 

Hab. Hawaii, Olaa. 

Nous avons decrit cette espece de Sumatra ; elle est sans doute repandue dans 
toute la region Malaise. 

Le C. bfissHS de Hawaii ne differe en rien de celui de Sumatra. 

67 — 2 



5i8 FAUNA HAWAIIENSIS 



(2) Che lifer hawauensis, sp. nov. 

$. Long. 5 mm. Cephalothorax anophthalmus, subtilissime coriaceus et opacus, 
baud granulosus, breviter et simpliciter parce pilosus, multo longior quam latior, antice 
parum attenuatus, obtusus et convexus, sulcis transversis binis subrectis impressus, 
sulco 1°, vix ante medium sito, profundissimo, altero leviore. Segmenta abdominalia 
opaca, setis fulvis minutissimis simplicibus sed truncatis, transversim ordinatis, munita. 
Pedes-maxillares longissimi, coxa nitida, reliquis articulis subtiliter coriaceis et opacis, 
brevissime et simpliciter pilosis, sed digitis pilis tenuibus albis longioribus munitis, 
trochantere breviter pediculato dein subgloboso sed paulo longiore quam latiore, postice 
convexo, prominulo sed obtusissimo, femore baud pediculato, a basi ad apicem sensim 
ampliato, intus recto, extus, in parte apicali, leviter convexo, tibia femore vix breviore, 
baud latiore et saltern triplo longiore quam latiore, et intus et extus leviter convexo, 
ad basin attenuato, manu tibia paulo longiore et paulo latiore, teretiuscula, apicem 
versus leviter ampliata, digitis manu paulo brevioribus, atque ad basin abrupte 
angustioribus. 

Cephalotborax et pedes-maxillares pallide fusco-testacei. Segmenta abdominalia 
dorsalia fulva, seg. i — iv maculis fuscis binis subgeminatis, seg. v — vi transversim 
late infuscata, reliqua maculis fuscis parvis quadriseriatis notata. Coxae pedum- 
maxillarium fusco-castaneae. Pedes lutei, subpellucentes, coxis olivaceis. 

?. Long. 4 mm. A mari differt magnitudine paulo minore, sulco thoracico 1° 
recurvo, pedibus-maxillaribus paulo brevioribus, femore ad basin minus attenuato, manu 
latiore, apice, versus digitos, leviter attenuata. 

Hab. Hawaii, Kona. — Kauai. 

Cette espece, relativement de grande taille, ressemble, par la structure de sa patte- 
macboire (dont le femur est graduellement attenue a la base, non pedicule) aux Ch. 
cancrotdes L. et mertciimms L. Koch, mais elle s'en distingue par le cepbalotborax 
anopbthalme et plus parallele, a peine attenue en avant. Elle pourrait former a elle 
seule un groupe special dans le genre Clielifer. 



Fam. GARYPIDAE. 

Garvpus L. Kocb. 

( I ) Garypits personaitis, sp. nov. 

$ — ?. Long. I '5 mm. Brevis, latus et valde depressus, subtilissime rugosus, pilis 
carens. Cephalotborax vix longior quam postice latior, antice, usque ad oculos, leviter 
attenuatus, ante oculos abrupte angustior, longe productus et obtuse canaliculatus, 



ARACHNIDA 519 

ante medium stria transversa superficiali vix expressa notatus. Oculi utrinque bini, 
inter se contigui, anticus postico paulo minor. Pedes-maxillares minute et crebre 
rugosi, digitis setis tenuibus longis paucis munitis, reliquis articulis nudis ; trochan- 
tere brevissime pediculato, dein lato, intus convexo et vix longiore quam latiore ; femore 
parallelo, baud pediculato ; tibia femore evidenter breviore, circiter aequilato sed versus 
basin sensim attenuate ; manu tibia circiter aequilonga sed duplo la^tiore, ovata, intus 
convexa ; digitis gracilibus, manu paulo longioribus. Pedum quatuor posticorum tro- 
chanteres longi et convexi. Truncus pallide luteo-testaceus, cephalothorace antice, 
usque ad oculos, fusco vel nigricanti, segmentis abdominalibus cunctis utrinque 
minute fusco-notatis, segmentis i et 11 maculis parvis medianis binis subgeminatis, 
seg. in immaculato, segmentis reliquis maculis parvis binis inter se remotis et 
lineas duas designantibus notatis. Pedes-maxillares obscure fusci, digitis rufescenti- 
bus. Pedes pallide lutei, subpellucentes. 

Cette tres petite espece est fort voisine du G. unnigatus E. Sim., de Sumatra ; elle 
s'en distingue surtout par ses teguments beaucoup plus finement rugueux et depourvus 
de polls claviformes. 

Hab. Oahu, Kaala Mts. 

Amblyolpium E. Simon, 
(i) Amblyolpium longiventer Keyserling. 
Olpium longiventer Keyserl. in L. Koch, Ar. Austr., 11, 1885, p. 50, tab. vi, 

fig- 9- 

Hab. Hawaii, Kau. 

Decrit d'Australie (Queensland) par Keyserling; indique depuis de Funafuti 
par R. I. Pocock. (Ann. Mag. Nat. Hist. ser. 7, vol. i, p. 321.) 

Le genre Amblyolpium differe du genre Olpium par le front tronque au niveau 
des yeux anterieurs, nullement prolonge en forme de museau. 



521 



CRUSTACEA ISOPODA. 



Par Adrien Dollfus. A , 

Armadillo Latr. ^'"^■fe 






(i) Armadillo kazvaiensis Dana {Spherillo). 

{i = Armadillo bidens, Budde-Lund. $, ^. australis, Budde-Lund.) 

La description et les figures du Spherillo kazvaiensis, Dana (Explor. Expedit., 
Crust, p. 722, pi. 47 (5« — «')), que nous avons pu consulter dans la bibliotheque de 
M. Milne-Edwards, ne nous laissent pas de doute sur I'identite de cette espece avec 
celle dont Mr Budde-Lund a decrit le $ sous le nom ^Armadillo bidens et la ? sous 
celui ^A. anstralis. — Cette espece presente un caractere de dimorphisme sexuel 
tres-particulier : en effet, le clypeus (partie cephalique situee au dessus du labre) est 
muni chez le ^ de deux processus dentiformes qui ne se voient pas chez la ?. 

Hab. Lanai, 2000 pieds, Perkins, 1892. — Kauai, Makaweli, 3000 pieds, 1897. — 
Hawaii, Lihue, 2000 pieds (vn. 1896). 

(2) Armadillo albospinostis, sp. nov. 

Corps tres-convexe, assez etroit, couvert d'epines coniques sur tout le corps, plus 
longues a la partie superieure. — Cephalon : bord frontal un peu incurve vers le centre, 
nettement margine ; epistome muni anterieurement d'un relief en ecusson assez accentue 
mais qui n'atteint pas la ligne frontale dont il est separe par une aire deprimee ; clypeus 
presque rectangulaire, a processus lateraux peu developpes ; yeux assez grands, environ 
16 ocelles ; antennes a premier article du fouet trois fois plus court que le second. — 
Pereion : premier article du pereion a partie coxale bien developpee, depassant sensible- 
ment Tangle du somite lequel presente I'aspect d'un relief triangulaire a la partie 
postero-laterale du premier segment ; deuxieme segment sans duplicature coxale 
distincte.- — Pleon, Telson : Pleotelson un peu plus long que large, un peu arrondi 
posterieurement et muni dorsalement de quatre epines coniques tres-fortes (chez le $) ; 
exopodite du premier pleopode (^) large et court avec une incurvation bien marquee ; 
endopodite tres-developpe, un peu recourbe au sommet ; uropodes peu obliques : 



522 FAUNA HAWAIIENSIS 

endopodites courts atteignant la moitie dii pleotelson ; exopodites petits, situes sur la 
face dorsale de I'article basilaire, a une petite distance de Tangle interne, et par 
consequent non visible sur la face inferieure de I'uropode. 

Couleur : gris brun vers le centre, un peu marbre sur les cotes, a epines blanchatres. 
— Dimensions : 8 x 3 millim. 

Obs. La % differe du ^ par ses granulations beaucoup moins spinescentes et 
etroites surtout vers la region mediane, et par une couleur generale qui parait uniforme- 
ment grfis-brun, I'extremite des ofranulations restant seules blanchatres. — Peut-etre 
s'agit-il la de deux especes differentes? 

Hab. Oahu, Kawailoa, i exemplaire 3". Perkins, iv. 1892. — Kauai, Makavk^eli, 
3000 pieds, [ exemplaire ?, 1897. 



(3) Armadillo perkinsi, sp. nov. 

Corps peu convexe, un peu deprime lateralement, ponctue et couvert de granu- 
lations assez fortes mais obtuses, disposees en rangees transversales assez confuses 
sur les trois premiers segments, au nombre de deux sur les quatre derniers segments 
pereiaux ; les granulations sont presqu'effacees sur le pleon. — Cephalon muni anterieure- 
ment de deux forts tubercules, depassant la ligne frontale, celle-ci est presque droite, 
un peu inflechie au centre ; epistome a relief a peine sensible ; clypeus court et peu 
large a processus lateraux peu developpes ; yeux moyens, environ 16 ocelles ; fouet des 
antennes a premier article trois fois plus court que le second. — Pereion : les deux 
premiers segments ne presentent aucune duplicature coxale distincte. — Pleon, Telson : 
Processus lateraux des segments du pleon peu divergents ; pleotelson un peu plus long 
que large, a bord posterieur egalant la moitie de la base ; uropodes a base allongee, 
etroite, a endopodite atteignant aux f du pleotelson, a exopodite assez developpe, situe 
vers la moitie du cote interne de la base. 

Couleur : ambre avec taches irregulieres brunes et rougeatres. — Dimensions : 7 y- I'k 
millimetres. 

Hab. Maui, Haleakala, 5000 pieds, iv. 1894, un exempl. $. 

(4) Armadillo danae, sp. nov. 

Corps peu convexe, un peu deprime lateralement, ponctue et presentant anterieure- 
ment des granulations espacees tres-obtuses, qui s'effacent posterieurement. — Cephalon 
muni anterieurement d'un relief (non bi-tubercule comme dans I'espece precedente), 
formant processus frontal et qui est borde anterieurement par une ligne marginee 
distincte, dedoublement de la ligne marginee frontale qui est un peu inflechie au centre ; 
epistome a relief peu sensible ; clypeus court et peu large, a processus lateraux peu 



CRUSTACEA 523 

developpes ; yeux assez petits, environ 1 2 ocelles ; fouet des antennes a premier article 
pres de trois fois plus court que le second.— Pereion : les deux premiers segments ne 
presentent aucune duplicature coxale distincte.— Pleon, Telson : Processus lateraux des 
segments peu divergents, ceux du ^' segment a bord posterieur un peu sinueux; 
pleotelson un peu plus long que large, a sommet depassant en largeur la moitie de la 
base; uropodes peu obliques, a cote interne sinueux, endopodite depassant les | du 
pleotelson, exopodite petit, situe vers les f du bord interne de la base. 

Couleur : ambre avec taches irregulieres brunes.— Dimensions : 7 x i\ millimetres. 

Hab. Kauai, Makaweli, 3000 pieds, 1897. 

(5) Armadillo s/iarpi, sp. nov. 

Corps assez convexe, finement ponctue, presentant anterieurement des tubercules 
obtus et sur le bord posterieur de chaque segment une rangee de granulations dont la 
mediane est la plus accentuee, donnant un aspect vaguement carene a la ligne mediane 
du corps. Cephalon : le front presente un processus large, analogue a celui de I'espece 
pr^cedente mais plus accentue encore, limite comme lui par un dedoublement de la 
marge frontale ; epistome a relief peu sensible ; clypeus court et peu large, a processus 
lateraux peu developpes ; yeux moyens, environ 1 2 ocelles ; fouet des antennes a 
premier article 2^ fois plus court que le second. — Pereion : les deux premiers segments 
ne presentent aucune duplicature coxale distincte. — Pleon, Telson : processus lateraux 
des segments du pleon peu divergents ; pleotelson un peu plus long que large, a sommet 
depassant en largeur la moitie de la base ; uropodes peu obliques, endopodite depassant 
les |- du pleotelson, exopodite petit, situe a une petite distance du sommet, sur le bord 
interne de la base. 

Couleur: gris presqu'uniforme, avec quelques petites taches plus claires. — Di- 
mensions : 7 X 3:1: millimetres. 

Hab. Kauai, 4000 pieds, viii. 1896, un exemplaire ?. 

Ces trois dernieres especes sont tres-voisines de \ Armadillo bifrons B. L., de 
Rockhampton (Australie), qui se rapproche d! A. perkinsi par le processus frontal 
bitubercule et ^A. sharpi par les granulations qui donnent a la partie mediane du 
corps un aspect carene. — Ces especes, auxquelles il faut joindre sans doute A. nigrinus 
B. L. (indique avec doute comme provenant du Cap(?)), — et peut-etre A. {Spherillo) 
spinosus, Dana, de la Nouvelle-Zdande, forment une section bien nette du grand genre 
Armadillo, caracterisee par I'absence de parties coxales distinctes sur les deux premiers 
segments pereiaux. Un autre caractere que je mets en evidence sur les trois especes 
Hawaiennes etudiees ci-dessus est presente par la disposition du front qui presente 
anterieurement un relief bien accentue, soit bitubercule comme dans A. perkinsi, soit 
continu comme dans A. danae et A. sharpi ; la marge frontale parait se dedoubler vers 
F. H. II. 68 



524 



FAUNA HAWAIIENSIS 



le tiers de son parcours en une ligne posterieure qui delimite le bord anterieur un peu 
inflechi de I'epistome, et une ligne anterieure qui suit le relief frontal ; toutefois dans 
A. perki7isi, cette ligne anterieure n'est pas nette ; les deux forts tubercules qui forment 
le relief frontal appartiennent certainement a la portion cephalique tergale et ne parais- 
sent etre qu'un acheminement a la disposition qu'offrent les deux autres especes ou 
la jonction entre le relief frontal et I'epistome parait plus complete ; la ligne posterieure, 
qui delimite ce dernier, semble meme s'attenuer encore plus dans A. sharpi que dans 
les deux autres especes. 



A. perkinsi. 



A. danae. 



A. sharpi. 



PoRCELLio Latr. 
(i) Porcellio laevis Latreille. 
Hab. Lanai, 2000 pieds, Perkins, 1892. 

(2) Porcellio scaber Latreille. 

Hab. Hawaii, Kona, Perkins, 1892. 

Cette espece, si commune dans I'Europe et meme I'Amerique septentrionale, avait 
ete trouvee en grande abondance dans certains pays temperes et froids de I'hemisphere 
sud. — On ne I'a signalee que tres-exceptionnellement en quelques localites des regions 
chaudes ; c'est la premiere fois quelle a ete rencontree a Hawaii. 



Alloniscus Dana, 
(i) Alloniscus floresiamis Dollfus. 

Hab. Lanai, Monts Koele, i. 1894. 

Nous avions decrit cette espece d'apres des exemplaires recueillis par le prof. Max 
Weber, a Sikka (Flores) {Isop. terr. des Indes Neerland., dans Zool. Ergebn. einer 
Reise in Niederl. Ost-Indien, Bd. iv. 1898, p. 374). \J Alloniscus d'Hawaii est bien 
identique a celui de Flores ; il est done probable qu'il faut lui assigner une assez grande 
extension. 



CRUSTACEA 525 



Philoscia Latr. 

(i) ? Philoscia angiisticaiida, Budde-Lund. 

Hab. Hawaii, Kona, 2000 a 3000 pieds, ix. 1892. — Oahu ; Mts. Kaala, 2000 
pieds, III. 1893 ; Honolulu, 2000 pieds, ix. 1892 ; Mts. Waianae, iv. 1892. 

C'est avec un certain doute que nous rapportons les exemplaires de Hawaii a 
I'espece decrite par Budde-Lund d'apres des exemplaires de Borneo (Crust. Isop. 
terrestria, p. 216), mais il ne nous est pas possible de Ten differencier nettement. 

Nous avions deja remarque (Zool. Ergebn. p. 377) que certaines especes des Indes 
Neerlandaises, notamment Ph. truncata Dollfus et Ph. variegata Dollfus, provenant 
de Celebes (nee Ph. variegata Dollfus, du Venezuela, Ann. Soc. Ent. France, 1893), 
etaient tres-voisines de Ph. angusticauda. Nous croyons qu'il faudra les identifier 
aussi avec I'espece de Borneo. 



Geoligia Dollfus. 

J'ai etabli en 1893 (Isop. rec. par M. E. Simon au Venezuela, d. Ann. Soc. Ent. 
France), sous le nom de Geoligia, un genre de Ligiens fonde sur un exemplaire tres- 
interessant decouvert par M. Eug. Simon au Venezuela, dans les forets, vers 1200 
metres d'altitude. J'indiquais comme caractere permettant de distinguer ce genre 
terrestre du genre Ligia qui parait eminemment maritime, I'absence complete de separa- 
tion des coxopodites (ou epimeres) qui sont au contraire bien distincts chez Ligia. 



(i) Geoligia perkinsi, sp. nov. 

Corps ovale, lisse. — Cephalon arrondi anterieurement ; parties laterales formant un 
processus triangulaire ; epistome muni d'une ligne transversale en relief assez accentu^ 
mais non flexueux ; clypeus tres-court et large ; labre tres-developpe ; yeux occupant 
environ § de la largeur du cephalon, pareils a ceux des Ligia ; antennes longues, a 
fouet plus long que la tige, forme d'environ 27 articles ; antennules tres-petites, a 
premier article large. — Pereion : les segments 2 a 7, munis lat^ralement d'un leger 
sillon, se terminent par un angle aigu ; I'incurvation posterieure du segment va en 
s'accentuant du 2*= au f segment ; la separation des coxopodites (epimeres), non distincte 
sur le premier segment, est indiquee par une ligne a peine visible sur les segments 
suivants.— Pleon : angles postero-lat<;raux des segments pleonaux tres-aigus ; pleopodes 
de la premiere paire (<?) a endopodite bien developpe, styliforme et exopodite large et 

68—2 



526 FAUNA HAWAIIENSIS 

court ; ceux de la 2^ paire a endopodite extremement long et se terminant en massue. 
■ — Pleotelson a bord posterieur muni de 5 dents obtuses, rappelant la forme de celui de 
Ligia exotica ; uropodes tres-longs, article basilaire beaucoup plus long que le pleotelson, 
appendices filiformes, artiadds, I'exopodite plus long que I'article basilaire, tri-articule, 
I'endopodite tres-grele, deux fois plus long que I'exopodite, sex-articule. 

Couleur : gris verdatre pale, muni sur tout le corps de tres-petits points pigmentes 
noirs, sauf sur les lineoles qui s'observent de part et d'autre de la ligne mediane. — 
Dimensions : longueur (J* et $) 16 a 17 millimetres ; largeur 6, 5 a 7 millimetres. 

Hab. Hawaii, Lihue, un exemplaire $ (avec un uropode complet), 2000 pieds, vii. 
1896; Olaa, 2000 pieds, ix. 1896; Hilo, 2000 pieds, i. 1896. — Kauai, Monts Waimea, 
4000 pieds, VI. 1894; Koholuamano, 4000 pieds, v. 1895. 

Les exemplaires recueillis par M. Perkins a Hawaii, egalement a une assez grande 
altitude, et qui me paraissent devoir etre unis generiquement a Geoligia simoni, m'ont 
permis de constater que dans I'espece d'Hawaii, la ligne de separation entre les 
coxopodites et le somite, nulle sur le premier segment, existe sur les segments suivants, 
mais elle est a peine distincte. Les appendices des uropodes sont articules ; ce dernier 
caractere est tres-remarquable ; il nous a ete malheureusement impossible de constater 
s'il existait aussi dans Geoligia simoni, les uropodes du seul exemplaire qui ait ete 
recueilli jusqu'a present etant incomplets. 

Nous devons done etablir la caracteristique du genre ainsi qu'il suit : 
Coxopodites (epimeres) non ou tres-peu distincts. Appendices des uropodes 
articules. — Le reste, et notamment les parties buccales, comme dans le genre Ligia. — 
Especes terrestres. 



Bibliographie (ouvrages cites). 

Dana, James D. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842. 
Crustacea, Part 11 (Philadelphia 1852). 

BuDDE-LuND, Gust. Crustacea Isopoda terrestria per familias et genera et species descripta. Hauniae 
1885. 

DoLLFUS, Adrien. Voyage de M. E. Simon au Venezuela (1887— 1888). Isopodes terrestres (.-\nn. 

See. ent. France, 1893, PP- 339—346, pis. 9, 10). 
. Isopodes terrestres des Indes Neerlandaises (Zoolog. Ergebn. einer Reise in Niederland. Ost- 

Indien, Band iv. Leiden, 1898, pp. 359 — 382, pis. xin — xv). 



527 



CRUSTACEA AMPHIPODA. ^'^S' 



■ O 



By Rev. T. R. R. Stebbing. 

Fam. TALITRIDAE. 

Gen. Orchestia Leach. 

Orchcstia plat ens is Krdyer. 

Plate XXI. A. 

1845. Orchestia platensis Kroyer, Naturhistorisk Tidsskrift, Sen 2, vol. i. 
PP- 304 — "o'^o, pi. 2, fig. 2, a— i. 

Kroyer's specimens were taken on a rocky margin of the river Plate just 
north-west of Montevideo. The agility with which the animal skips, he says, is so 
great that it is pretty difficult to catch it. Specimens apparently belonging to this 
species have been entrusted to me by the authorities of the Copenhagen Museum. The 
labels show that these examples were taken at the Bermudas, at St Croix (West 
Indies), and at Beloxi (Mississippi, U.S.A.). Specimens of Orchestia agilis S. I. Smith, 
from the east coast of the United States, have been given me by my friends, Canon 
Norman and Mr S. J. Holmes. These also in my opinion should be included in 
Kroyer's species. 

Now, after much hesitation, I feel constrained to increase its range by applying 
the name to the specimens taken by Mr Perkins at an elevation of 3000 feet and 
2000 feet respectively in the Sandwich Islands. At the greater height there was but 
one example, labelled " Kona, Hawaii." The others were labelled " Oahu " and 
" Honolulu," and " Lanai." Others were labelled, without specifying elevation, 
" Waianae Mts., Oahu, shrimps in dead wood." 

The eastern specimens differ to a slight degree from the western in having the 
second antennae more slender, with flagella not always limited to thirteen or fourteen 
joints, one of the female specimens having twenty-three, and one of the males twenty- 
four joints ; also the first gnathopods of the male have the distal part of the sixth joint 



528 FAUNA HAWAIIENSIS 

less conspicuously widened, nor can I guarantee that the armature of spines on the 
uropods is precisely the same in both forms. 

In the second gnathopods of the male the large oval sixth joint has a smoothly convex 
palmar margin, bordered with submarginal spinules. Round this the long finger curves, 
closely fitting it, but laying its apical part within a long groove which succeeds the 
palmar margin. Kroyer, it should be observed, neither figures nor describes this apical 
part of the finger, which is membranaceous in appearance, slender, but not very acute 
at the point. According to Kroyer the telson is at the apex flatly rounded off or nearly 
truncate. In the specimens which 1 have examined in this respect it is minutely 
emarginate. 

The peraeopods are without specially dilated joints. The second joint of the 
fourth peraeopods is much more narrowly oval than that of the fifth pair; the second 
joint of the latter has its hind margin rather strongly serrate. The first pair of uropods 
have the outer ramus free from spines except at the apex, but this character is common 
to various other species in the family. 

Length reaching 12 mm. 



Orchestia pickeringii Dana. 

Plate XXI. B. 

1S53. Orchestia pickeringii Dana, U. S. Exploring Exp., Crustacea, pt. 2, 
p. 882, pi. 59, f. 9 a — f 

Dana, at p. 1595 of his great work, states that 'the locality of Orchestia 
pickeringii, was Kauai or Oahu, Sandwich Islands.' He does not give any further 
detail as to the habitat. Stimpson records it from California, without comment. The 
specimens obtained by Mr Perkins are labelled " Honolulu. Oahu. 2000 ft." They 
were taken on the same date and at the same height as specimens of Orchestia platensis. 
and the question suggested itself whether by chance the two species might be identical. 
There is considerable general resemblance, but that perhaps is only as much as to say 
that both are species of Orchestia. The idea of uniting them is, 1 think, untenable. 

The second gnathopod of the male has a very large sixth joint, the sloping palmar 
margin ot which is in these specimens diversified b\- two spinulose projections, in 
correspondence with which the strongly curved finger has a very sinuous inner margin, 
its apex entering a small pocket or groove of the somewhat flattened hind margin of the 
sixth joint. Dana describes the palm as having two very low prominences near base of 
finger, whereas these are very marked, and one of them is remote from the base of the 
finger. But an Oahu specimen from the Copenhagen Museum and an Australian 
specimen from the Australian Museum, show modifications approximating to Dana's 



AMPHIPODA 529 

form. These have the inner margin of the finger smoothly concave. In Dana's figure 
the finger has a sHght convexity near the base, and also has a produced apex, corres- 
ponding with a long groove in the sixth joint, which in a manner recall the same 
features in Orchestia platensis. 

Dana gives the fjagellum of the first antennae three-jointed. This variable feature 
has sometimes five joints. The ultimate and penultimate joints of the peduncle of the 
second antennae are much stouter than in any of the specimens referred to Orchestia 
platensis, and yet the flagella are not more than fourteen-jointed. It seems unlikely 
that the larger and more strongly armed male should have a shorter flagellum than a 
less highly developed male of the same species. The specimens referred to Orchestia 
pickeringii have the fourth and fifth joints of the fifth peraeopods, though not dilated, 
thickened in a way not exhibited by the other set. This, however, might be a 
character of advanced maturity. 

Length reaching 14 mm. 

One female specimen, 12 '5 mm. long, which I am inclined to refer to this species, 
differs from those of the same sex referred to Orchestia platensis in rather superior bulk, 
and also in having a distal boss on the fourth joint of the second gnathopods, agreeing 
in that respect with Dana's figure of the limb in question in Parorchestia haivaiensis, $. 
From that species it is completely distinguished by the short, few-jointed, upper 
antennae. 

Gen. Parorchestla., Stebbing. 

1899. Parorchestia Stebbing, Trans. Linn. Soc. London, vol. vii, pt. 8, pp. 397, 

402. 
This genus was instituted to receive three of Dana's species of Orchestia, namely, 
temiis, haivaiensis, and sylvicola. 

Parorchestia haivaiensis Dana. 

Plate XXI. C. 

1853. Orchestia haivaiensis Dana, U.S. Expl. Exp., Crustacea, pt. 2, p. 880, 

pi. 59, f. 8 a— g. 
1899. Parorchestia haivaiensis Stebbing, Trans. Linn. Soc. London, vol. vii, 

pt. 8, p. 402. 
Dana's description and figures of this species, obtained at ' Oahu or Kauai, 
Hawaiian Islands,' refer only to the female. Specimens of both sexes obtained by Mr 
Perkins were labelled as coming from " Koholuamano, Kauai," and from " Waianae 
Mts., Oahu." It has also been taken by Dr Arthur Willey at Lifu. 



530 FAUNA HAWAIIENSIS 

From the species of Orchcstia above discussed the present species is separated by 
many particulars in addition to the minute distinction on which the genus Parorchestia 
is founded. 

(/. The upper antennae extend along at least two-thirds of the last joint of the 
peduncle of the lower antennae. The second joint of the peduncle is longer than the 
first, the third as long as the first plus the second ; the flagellum of nine joints is as 
long as the peduncle. 

The lower antennae are slender, the penultimate joint of the peduncle more than 
twice as long as the preceding joint, and the ultimate as long as both of these together; 
the flagellum of twenty-one joints is as long as the peduncle or rather longer. As 
usual, these dimensions and numbers are subject to some variation. 

The upper lip appears to be rather wider and shorter than in Orchcstia, but no 
tangible differences could be discerned in mandibles, lower lip, and the two pairs of 
maxillae. The maxillipeds differ from those of Orc/iesfni platensis in having both the 
inner and outer plates smaller, the first and second joints of the palp less expanded, 
the third joint with less convex margins, as well as in possessing a distinct though 
minute fourth joint ; also the spinules on the various joints are fewer and more 
scattered, none being seen on the outer margin of the inner plate, while on the other 
hand the outermost spine-tooth on its distal margin is exceptionally large. 

The first gnathopods are slight, the fourth joint having a sort of boss or distal 
expansion of the hind margin ; the fifth joint widens gradually to the rounded distal end, 
and the sixth is rather similarly expanded, the small finger not quite reaching the 
extremity of the palmar margin, which is not at all oblique. 

The second gnathopods have the sixth joint massive, broadly oval, the palmar 
margin oblique, not strongly convex, ending in a small groove or pocket, to receive the 
apex of the finger. The inner margin of the finger is slightly irregular, at its base not 
fitting exactly the palmar margin, while its apex has a membranaceous aspect as in 
Orchcstia platensis. 

The uropods scarcely differ in any important particular from those of the above- 
named Orchcstia, but the telson has a truncate apex, with a spinule at each corner, two 
smaller ones in between them, and two on each side above them. 

Length 13 mm. 

Except in regard to the two pairs of gnathopods and the purely sexual characters, 
the male and female have close resemblance. Dana's figures of the female are unfortu- 
nately very poorly reproduced in the Catalogue of the Amphipodous Crustacea in the 
British Museum, 1862, a work which might have been of extreme value, had the plates 
been executed with sufficient care to make them trustworthy. 



ZOOLOGICAL RESULTS based on material from New Britain, New Guinea, Loyalty 

Islands and elsewhere, collected during the years 1895, 1896, and 1897, by 

Arthur Willey, D.Sc. Lond., Hon. M.A. Cantab., Late Balfour Student of the 

University of Cambridge. Demy 4to. Parts I, II, and III. Price 12s. 6d. each. 

PART I. 1898. 

The anatomy and development of Peripatus novae-britanniae. By ARTHUR WiLLEY, M.A., D.Sc. 

With Plates I. — IV. and 7 figures in the text. 
Metaprotella sandalensis, n. sp. [Caprellidae]. By Dr PAUL MAYER. 

With 6 figures in the text. 
On a little-known Sea-snake from the South Pacific. By G. A. BOULENGER, F.R.S. 

With Plate V. 
Report on the Centipedes and Millipedes. By R. I. POCOCK. 

With Plate VI. 

5. Account of the Phasmidae with notes on the eggs. By D. SHARP, M.A., F.R.S. 

With Plates VII.— IX. 

6. Scorpions, Pedipalpi and Spiders. By R. I. PocoCK. 

With Plates X.— XI. 

PART II. 1899. 

Report on the specimens of the genus Millepora. By SYDNEY J. HiCKSON, M.A., D.Sc, F.R.S. 

With Plates XII.— XVI. 
Report on the Echinoderms (other than Holothurians). By F. JEFFREY BELL, M.A. 

With figures on Plate XVII. and one figure in the text. 
Holothurians. By F. P. BEDFORD, B.A. 

With figures on Plate XVII. 

10. Report on the Sipunculoidea. By Arthur E. Shipley, M.A. 

With Plate XVIII. 

11. On the Sohtary Corals. By J. STANLEY GARDINER, M.A. 

With figures on Plates XIX. and XX. 

12. On the postembryonic development of Cycloseris. By J. STANLEY GARDINER, M.A. 

With figures on Plates XIX. and XX. 

13. On a collection of Earthworms. By FRANK E. Beddard, M.A., F.R.S. 

With Plate XXI. 

14. The Gorgonacea. By ISA L. HiLES, B.Sc. 

With Plates XXII. and XXIII. 

PART III. 1899. 

15. Orthogenetic variation in the shells of Chelonia. By Hans Gadow, M.A., Ph.D., F.R.S. 

With Plates XXIV.— XXV. and one text-figure. 

16. Enteropneusta from the South Pacific, with notes on the West Indian Species. By ARTHUR 

Willey, D.Sc, Hon. M.A. Cantab. 

With Plates XXVI.— XXXII. and 7 text-figures. 

17. On a collection of Echiurids from the Loyalty Islands, New Britain and China Straits, with an attempt 

to revise the group and to determine its geographical range. By Arthur E. Shipley, M.A. 
With Plate XXXIII. 

PART IV. 1900. 

18. On the anatomy of a supposed new species of Coenopsammia from Lifu. By J. Stanley 

Gardiner, M.A., Fellow of Gonville and Caius College, Cambridge. With one plate. 

19. On the Insects from New Britain. By D. Sharp, M.A., M.B., F.R.S. With one plate. 

20. Report on the Stomatopoda and Macrura. By L. A. Borradaile, MA., Lecturer of Selwyn 

College, Cambridge. With four plates. 

21. Report on the Slugs. By Walter E. Collinge, F.Z.S., Mason College, Birmingham. 

With two plates. 

22. Report on the Polyzoa. By E. G. Philipps, Newnham College, Cambridge. With two plates. 

23. The Hydroid Zoophytes. By LAURA RoscoE Thornely, University College, Liverpool. 

With one plate. 

24. Astrosclera willeyaiia, the Type of a new Family of Calcareous Sponges. By J. J. Lister, M.A., 

St John's College, Cambridge. With five plates. 

25. A contribution towards our knowledge of the Pterylography of the Megapodii. By W. P. 

Pycraft, A.L.S., British Museum. With one plate. 

26. The Stolonifera and Alcyonacea. By SYDNEY J. HiCKSON, M.A., D.Sc., F.R.S., Beyer Professor 

of Zoology in the Owens College, and ISA L. Hiles, B.Sc, Owens College, Manchester. 
With two plates. 

27. Report on the Xeniidae. By J. H. ASHWORTH, B.Sc, Demonstrator in Zoology, Owens College, 

Manchester. With two plates. 

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FAUNA HAWAIIENSIS 



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1 



Contributions have been made or 
W. H. ASHMEAD ... 

G. H. Carpenter ... 

Prof. FiLIPPO SiLVESTRI 

F. E. Beddard, F.R.S. 

M. A. DOLLFUS ... 
Prof. AUGUSTE FOREL 

p. H. Grimshaw, F.E.S. 

G. W. Kirkaldy, F.E.S. 
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Hugh Scott, B.A. 

R. C. L. Perkins, D.Sc. 

D. Sharp, F.R.S. ... 
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M. EuG. Simon 
The Rev. T. R. R. Stebbing, F.R.S. 

E. R. Sykes, F.Z.S. 
The Lord Walsingham, F.R.S 



promised by the following, viz. 

Hymetwptera Parasitica. 

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FAUNA HAWAIIENSIS 

OR THE 

ZOOLOGY OF THE SANDWICH (HAWAHAN) ISLES 

Being Results of the Explorations instituted by the Joint Committee 

appointed by 

THE ROYAL SOCIETY OF LONDON FOR PROMOTING NATURAL KNOWLEDGE 
AND THE BRITISH ASSOCIATION FOR THE ADVANCEMENT OF SCIENCE 

And carried on with the assistance of those Bodies and of the Trustees of 
THE BERNICE PAUAHI BISHOP MUSEUM AT HONOLULU. 

EDITED BY 

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SECRETARY OF THE COMMITTEE. 



VOLUME II. PART VI. 

SUPPLEMENT 

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By G. W. KIRKALDY. 

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To ORTHOPTERA, NEUROPTERA and DIPTERA. 
By R. C. L. PERKINS. 



Pages 531—700. 



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531 






Supplement by G. W. Kirkaldy 

to 

HEMIPTERA. 

Vide pp. 93 — 174 hujus operis torn. in'. 

This second contribution deals principally with the Auchenorrhynchous Homo- 
ptera ; I have, however, added species of the Heteroptera etc. described since 1902, as 
well as new species and notes on previously known forms. 

The total number of the Hemiptera so far described from the Hawaiian Islands 
is 305, or, if we exclude the 63 determined species of Coccidae, Aleyrodidae and 
Aphidae, which greatly swell the non-endemic total, 242. Of these 226 are not 
known elsewhere, though I have grave doubts as to the endemicity of some of them. 

The Hemipterous fauna is now seen to lack representatives in many of the most 
important families, only the following six containing undoubtedly endemic forms, viz. 
Myodochidae, Nabidae, Reduviidae, Anthocoridae, Miridae and Acanthiidae, among the 
26 recognized Heteropterous families, and by four only of the 14 Homopterous families, 
viz. Tettigoniidae, Fulgoridae, Asiracidae and Chermidae, that is to say, by 10 out of 
40, or exactly one-fourth ! But this scanty representation is still better shown by the 
fact that out of these ten, only seven are represented by more than ten species each, 
and that none contain more than one subfamily, except the Myodochidae, which have 
an endemic subfamily, Metrarginae. 

Although there are several species still to be described, I think that all but a very 
few genera have been accounted for. 

In the Cimicinae, Oechalia and Coleotichus are possibly endemic, as regards their 
species, but, on the other hand, these may very possibly be found in Samoa or Tahiti, 
the endemic Hemipterous fauna of which is unknown. 

' I regret that I have to announce the decease of Mr Kirkaldy, which has occurred while these pages 
have been in the press, so that they have not had the benefit of his revision. The residence of Mr Kirkaldy 
in the islands for some years, induced a change in his views on numerous points, so that the alterations in 
this supplement are of considerable extent and importance. The species with a number prefixed are not 
hitherto enumerated in the Fauna, and are practically additions, though some were described previous to 
Mr Kirkaldy's first treatment of the subject. Editor. 

F. H. II. 69 



532 FAUNA HAWAIIENSIS 

The following is a list of the families represented endemically in these Islands : 
Myodochidae, with Metrarginae and Cyminae ; Nabidae; Reduviidae with 
Ploiariinae only; Anthocoridae ; Miridae ; Acanthiidae ; Tettigoniidae ; with the 
tribe Phrynomorphini only; Fulgoridae, with Cixiinae only; Asiracidae; and 
Chermidae. 

Cimicidae. Excluding an undoubted introduction, we have three species that are 
not known elsewhere, but may occur in Samoa or some other Pacific Island. 

Urolabididae and Aradidae are entirely absent. 

Lygaeidae. Excluding an undoubted introduction, we have Ithaniar, which will 
probably be found elsewhere. 

Myodochidae. In the Metrarginae, we have an apparently endemic subfamily, 
which however may occur in Samoa and other islands. Nysms has between 20 and 30 
species, all the endemic ones being arboreal. I think all the latter could have 
originated from one immigrant. Sephora and Nesocymns are, I think, endemic, but 
possibly not. Reclada I do not know, and, I think, it is not endemic. 

Pyrrhocoridae and Tingidae are entirely absent endemically, as also Gerridae, 
Enicocephalidae, Nepidae and Macrocephalidae. 

Nabidae are well represented by more than 20 endemic species. Milu is doubtless 
a direct offshoot of a Reduviohis like sharpianus. The endemic Reduviolus proper are 
arboreal, but their offshoot Nesotyphlias are found on the ground under dead leaves etc. 
and on tree-ferns. 

Reduviidae are represented endemically only by the slender, long-legged Ploiariinae, 
of which I have recorded seven species ; one of these is however probably not endemic. 

Anthocoridae have six or more endemic forms. 

Clinocoridae, Polyctenidae and Dipsocoridae are absent. 

Miridae require a revision with more material, and there is a considerable number 
of species undescribed. Siilamita, Kalania, Pseudoclerada and Kavieha^neha are 
interesting endemic genera. Tichorhinus and Sarona seem to be the dominant forms. 

Acanthiidae are represented by five or more endemic species of Acanthia, which 
are very variable in colour and pattern and need fresh study with more material. The 
other families of the Notonectoidea are not represented endemically. 

In the Homoptera, Cicadidae, Cercopidae and Membracidae are absent, as also 
(endemically) Aphidae, Aleyrodidae and Coccidae. Tettigoniidae are represented 
by the endemic Nesophrosyne with a large number of forms, extremely variable 
and difficult to limit, by NesopJiryne (one to four species) also apparently endemic, 
and by one or two other forms in the same tribe (Phrynomorphini) also possibly 
endemic. 



HEMIPTERA 



533 



FuLGORiDAE consist only of Cixiini, in two genera, the endemic lolania (several 
species, only one yet described) and the cosmopolitan Oliarus, with several endemic 
forms. 

AsiRACiDAE are well represented and comprise several endemic genera and a large 
number of species. 

PoEKiLLOPTERiDAE, IssiDAE, Tettigometridae and Derbidae are absent, the first 
named having a single immigrant. 

Chermidae are rather numerous, but very variable, and scarcely studied. 

My thanks are due to the following gentlemen : to Dr R. C. L. Perkins for loan 
of specimens and much help in various ways ; to Mr W. M. Giffard for the gift of his 
valuable Hemipterous collections made in various localities, but principally on Mt. 
Tantalus, Oahu ; and to my colleagues Messrs F. W. Terry and O. H. Swezey, for 
specimens. 



Heteroptera 

For a sketch of the classification of the Heteroptera see Kirkaldy, 1907, P. Haw. 
E. S. I. 135 et seqq. ; and 1908, Can. Ent. xl. 



Fam. CIMICIDAE. 

Oechalia. 

As stated elsewhere, P. Haw. E. S. i. pp. 141 — 3, I feel certain that there are 
two valid species of this genus, as there are two kinds of ova ; at the same time I do 
not believe that the synonymy already given is incorrect. 

Oechalia grisea, Burmeister. 

Oe. griseus, huj. op. in. p. 171. 

I have described some of the stages and figured an egg. Dr Perkins writes in 
reference to other remarks in that paper, "You give a wrong impression as Oechalia 
often swarms in Myoporum, Acacia koa and many others. I have seen hundreds in 
a few yards of sugar cane and it is often very common near the coast and even in the 
outskirts of Honolulu on foreign vegetation. The whole remark certainly gives no 
idea of the ubiquity of this very abundant bug. It is on nearly all native trees, both in 
egg and other stages, as well as on low vegetation." Recently, I have taken it at 
Kilauea (Hawaii) on Myoporum, Acacia koa and ferns. 

69 — 2 



534 FAUNA HAWAIIENSIS 

C. blackburniae. 

C. blackburniae, huj. op. iii. p. 172. 

I have described and figured the ova, and described the nymphal stages (1907, 
op. cit. p. 144, figs. 2 — 4). Dr Perkins writes " I have seen it swarming near the 
coast on Dodonaea as well as in the mountains. I never saw it on Koa in anything 
like such numbers as on Dodonaea. The latter covers acres of land where is no Koa. 
The fact is it is partial to ' pods ' and ' capsules ' and the tree itself is of minor 
consideration, if it bears these." Maui should be added to the list of islands. 

[Van Duzee (1905, Bull. Amer. Mus. xxr. 207) has cited Piezodorus rubrofasciatus 
as a Hawaiian Insect, but in error. See Kirkaldy, 1908, P. Haw. E. S. i. 172 and 187. 
Oshanin has given these islands as a habitat of Carpocoris ptidicus van fuscispina, also 
in error.] 

Subfam. CYDNINAE. 

Geotomus pygmaeus Dallas. 

G. pygmaeus, huj. op. iii. p. 172. 

This little species is common under dried cow-dung, and under stones. It is 
nocturnal, coming to "light" in the evenings. Add Kauai and Lanai to the islands. 

Fam. LYGAEIDAE. 

Ithamai' hawaiiensis Kirkaldy. 

/. hawaiiensis, huj. op. in. p. 1 70. 

Dr Perkins writes, " this species is not attached to Sida, it abounds on Molokai at 
3000 ft. on other plants, and at the Volcano (Kilauea), and as high as 8000 — 10,000 ft. 
on Haleakala. You refer merely to the coast-line." It has been common in Kaimuki 
this year (1908) on Sida cordifolia, but is attacked by Zelus refiardii. I have briefly 
described the eggs and first nymphal instar (1907, P. Haw. E. S. i. p. 149). Add 
Hawaii to the islands. 

This species, although unknown elsewhere, must be a recent introduction as it was 
not taken by Blackburn who would certainly have collected it near Honolulu had it 
been here then. 

Rhopalus hyalinus, Fabr. 

R. hyalinus, huj. op. iii. p. 1 70. 

I have described the ova and nymphs (1907, op. cit. pp. 146 — 8). This has been 
common this year on Sida cordifolia in Kaimuki, as well as on Sonchus oleraceus, its 
more natural food plant. It feeds also on Euphorbia cordata and is preyed on by Zelus 
renardii. Add Kauai to the islands. 



HEMIPTERA 535 

Fam. MYODOCHIDAE. 
Metrarga. 

Metrarga, huj. op. in. p. 165. 

I have recently founded two subgenera in this, viz. Nesocryptias and Nesocliniacias 
(1908, P. Haw. E. S. I. p. 189). 

The forms now known are : 

a. typical subgenus : 

M. nuda (F. H. in. p. 165) typically apparently an Oahuan species, with 

var. mauiensis (1908, P. Haw. E. S. i. p. 188). 
M. obscura (F. H. iii. p. 166) ; I have recently taken this at Kilauea, Hawaii, 

on Ohia lehua i^Nani polyniorpha). 

b. subg. Nesocliniacias. 

contractu (F. H. in. p. 166); with vas. picea (1908, P. Haw. E. S. i. p. 188). 

(i) lanaiensis (1908, 1. c), sp. n. 

c. subg. Nesocryptias. 
villosa (F. H. in. p. 167). 

Dr Perkins writes me that the species of Metrarga are " certainly not bark- 
dwellers, unless as a rare and exceptional case." 

Sephora crinigera White. 

SepJwra criniger, huj. op. in. p. 161. See also Kirkaldy, 1907, Can. Ent. 
XXXIX. p. 244. 

Nesomartis. 
Nesomartis Kirkaldy, 1907, op. cit. p. 245. 

(i) Nesoviartis psammophila Kirkaldy. 
Nesomartis psammophila' Kirkaldy, 1907, 1. c. 

Hab. Oahu, "on the coast on the ground amongst Sida" grass etc., and 
"Hawaii, Kona coast in similar situations" (Perkins). Probably introduced. 

Nesocymus. 

Nesocy^nus, Kirkaldy, 1907, 1. c. 

Apparently allied to Cymoninus Breddin, but there is no percurrent main vein on 
the corium, at least not one at all which " teilt das Corium in zwei annahernd gleiche 
Telle," and while the corium is partly punctured, partly smooth, it is the middle oblique 



SS6 FAUNA HAWAIIENSIS 

third which is punctured, almost all the rest being smooth ; moreover, the clavus is 
closely punctured. The first segment of the antennae is distinctly longer than the long 
diameter of an eye and extends beyond the clypeus. The third and fourth segments 
are about equally long. The impressed line on the pronotum is situated about the 
middle (as stated on p. 245 ; a misprint occurs on p. 244 in the table). 

Nesocymus calvus. White. 

Sephora calvus, Kirkaldy, 1902, F. H. in. 162. 

Nesocymus calvus, Kirkaldy, 1907, Can. Ent. xxxix. 245. 

Hab. Oahu, at roots of herbage in the mountains, from 1500 — 2000 ft. (Perkins). 
At the top of Tantalus (Perkins, Gififard). One of Giffard's specimens has the second 
and third segments of the right antenna coalesced and shortened, a comparatively 
common occurrence in the Cymini. 

Nvsius Dallas (huj. op. in. p. 162). 

In the ensuing descriptions, I have, owing partly to the fact that several species 
are represented only by one or two carded specimens, paid no attention to the bucculae, 
structures considered by Stal to be so important, but have used other characters which 
appear to me better for specific differentiation. The form of the metapleura, utilized 
here for subgenera, is considered by Stal and later authors as of generic importance, in 
other Cyminae, but I cannot regard it as such in Nysius. Orsillus, distinguished 
among other things by the armed fore femora, has the labium passing beyond the hind 
coxae, Nysius being separated by the same authors by the unarmed femora and shorter 
labium. I cannot, however, regard the length of the labium here as a generic character. 

I have therefore united all the species now described, in a single genus Nysius, 
but have separated off two subgenera, Oceanides and Neseis. 

It is unfortunate that I have not seen the types of the species proposed by 
Blackburn and White, and that I am unable to identify ten of them. Most of them 
were recorded from Kilauea, Hawaii, a locality rather closely investigated on several 
occasions by Dr Perkins and Messrs Swezey and GifTard, as well as by myself. 

Conspectus quaruTndam speciei^ufn hawaiiensium. 

I. Metapleura postice satis rotundatim emarginata, angulo exterolaterali nonnihil 

prominente, margine postica late reflexa (2). 

\a. Metapleura postice truncata, plerumque anguste, interdum late, reflexa; 

labium post coxas posticas extensum [subj. Cr^a«/«^« HOV., 

nimbato typo]. 

Labium ad metasternum extensum (3). 

Labium ad coxas posticas extensum (6). 



2. 

2(7. 



HEMIPTERA 



537 



2t>. Labium post coxas posticas extensum; callositas pronoti flavescens; pro- 

notum antice ad margines laterales subito rotundatum [subj, Neseis nov.] 

28 monticola nov. 

3. Species pubescentia satis fortiter vestita 7 delectus White'. 

3a. Species glabrae, nitidae iA. 

4. Callositas pronoti nigrescens (5). 

4a. Callositas pronoti pallida i saundersianus Kirkaldy^ 

5. Color ex parte maxima fuscescens vel nigrescens 2 Mauiensis Blackburn. 

5a. Color ex parte maxima pallida 8 hylaeus nov. 

6. Species pubescentia vestitae (7). 

6a. Species glabrae, nitidae ; vel pubescentia minutissime vestitae (9). 

7. Longitudo ultra 7 mm 9 Kamehameha Kirkaldy. 

la. Longitudo 4 — 4! mm (8). 

8. Supra pallidior ; femora postica pallida, fuscomaculata 10 coenosulus StSl". 

8a. Nigrescens ; femora postica nigrescentia 12 Itchenicola nov. 

g. Caput flavescens, ex parte leviter brunneosufifusum 13 ochriasis Kirkaldy. 

9a. Caput nigrum (10). 

10. Femora postica pallida, brunneomaculata 14 insu/ko/a nov. 

10a. Femora postica pallida, immaculata, apice roseosuffuso 15 sitvestris nov. 

11. Pronotum nigrum, maculis pallidis tribus parvis ad marginem posticam 

ornatum 16 oresitrophus nov. 

i\a. Pronotum rufescens vel brunneoflavescens, areis laevigatis nigris vel nigre- 

scentibus vel fuscescentibus (12)- 

1 2. Vertex linea rufescente mediana longitudinali ornatus 17 tiubico/a nov.^ 

1 2a. Vertex niger, vix maculatus (13)- 

13. Tegmina baud maculata vel ad marginem apicalem corii prope medium 

brunnep-notata, interdum fortius maculata. Pedes pallidiores, leviter 

maculatae 19 nimbaius nov. ^ 

13a. Tegmina fortiter fusconotata. Pedes magis fuscatae (14)- 

14. Labii segmentum primum post marginem apicalem prosterni extensum (15). 

14(2. Labii segmentum primum baud ad capitis basim extensum 24 oribasus nov. 

15. Species glabra, vel sparsissime pubescens 26 montivagus nov. 

15a. Species pubescens 25 insulivagus nov. 

Nysius saundersianus Kirkaldy. 

Nysius saundersianus Kirkaldy, huj. op. iii. p. 163. 

I recorded this formerly from Lanai, Molokai and Hawaii ; Mr Giffard has since 
taken it on the former island, Koele Mts., 2000 ft. (Oct. cf. Giffard, 1908, P. Haw. 
E. S. I. 180). 

The pronotal callosity is pale, and extends from one lateral margin to the other ; 
the lateral margins are somewhat sinuate. 

' Verisimiliter prope banc speciem pertinet N. dallasi White, species mihi incognita. 

* Verisimiliter prope banc speciem pertinent N. arboricola White, N. longicol/is White, iV. whitei 
Blackburn, species mihi incognitae. 

' Verisimiliter prope banc speciem pertinet N. hlackbur>n White, species mihi incognita. 

* Prope banc speciem pertinet verisimiliter N. iwmorivagus White. 

* Prope banc speciem pertinent verisimiliter N. rnbescens, nitidiis, pteridicola, vulcan White 



538 FAUNA HAWAIIENSIS 

(i) Nysius mauiensis Blackburn. 

Nysius mauiensis Blackburn, 1888, P. Linn. Soc. N. S. W. (2) iii. 

I have identified a number of specimens as this species, but am hampered by the 
fact that I do not know N. arboricola, with which Blackburn compares it. The length 
of the labium is not stated, but I suppose it to be as in A^. arboricola. 

This species seems somewhat variable, the antennae being feebly ringed, con- 
spicuously so, or almost entirely fuscate ; the tegmina are in some specimens principally 
pale, in others principally dark. 

Hab. Maui, Haleakala, 4500 ft. (Blackburn), 5000 ft. (Mar., Apr., Perkins). — 
Lanai, 2000 ft. (Jan., Sept., Perkins), 3000 ft. (Feb., Perkins). — Hawaii, Kona, 4000 ft. 
(Aug., Perkins); Kilauea, 4000 ft. (Perkins). 

(2) Nysius arboricola White. 

Nysius arboricola F. B. White, 1878, A. M. N. H. (5) i. 368. 
I do not know this species. Blackburn took it "high up in the mountainous 
district," i.e. of Oahu, probably Mt. Tantalus. 

(3) Nysius whitei Blackburn. 

Nysius whitei Blackburn, 1888, op. cit. 346. 

This was obtained from Hawaii, Mauna loa, abt. 4000 ft. (Feb.). I do not 
know it. 

(4) Nysius longicollis Blackburn. 
Nysius longicollis Blackburn, 1888, op. cit. 344. 
This was obtained on Oahu. I do not know it. 

(5) Nysius dallasi White. 
Nysius dallasi F. B. White, 1878, op. cit. 367. 

This was taken on Oahu, near Nuuanu Pali. It is probably an introduced form 
near N. delectus, but I have been unable to determine it, though it may be the species 
I incorrectly, according to Dr Bergroth, determined previously as A^. vinitor Bergroth. 

(6) Nysius delectus White. 

Nysius delectus F. B. White, 1878, op. cit. 367 ; Kirkaldy, 1907, P. Haw. 
E. S. I. 152. 

An introduced form, widely distributed ; near the coast it is common on 
Bidens pilosa. 

Hab. Apparently on all the islands at almost all elevations ; not arboreal. 



HEMIPTERA 



539 



(7) Nysins kylaetis, sp. nov. 

Pale yellowish testaceous, glabrous. Vertex with some dark markings, principally 
longitudinal, which vary in situation and depth of colour, in the three specimens before 
me. Second and third (except extreme apex), a spot on fourth, and the whole of the 
fourth segment of antennae, blackish brown (in one example, second and third paler). 
Eyes red-brown. Ocelli red. Pronotum with a short, oblique, subpyriform, laevigate, 
black spot on each side of the middle, not nearly reaching the lateral margins of the 
pronotum. Pronotum closely punctured, punctures mostly nearly colourless, except on 
and near the apical margin. Postero-lateral angle in some examples dark, also the 
hind margin is soiled in some. Scutellum similarly punctured, a blackish spot on the 
anterior half in the middle. Sterna dark brown, polished ; pleura closely punctured, 
a little darkly suffused in part ; hind margin of metapleura pale, not punctured. 
Tegmina rather feebly mottled with brownish grey, apical margin of corium in some 
examples more darkly so, apical angle in some blackish brown. Membrane hyaline, 
mottled with brown. Femora with a subapical dark brown ring, tinged with rose 
apical of this. Abdominal sternites tinged with rose, a sublateral black blotch on basal 
half (this is somewhat variable). Antennae with the first segment extending just 
beyond apex of vertex, second slightly longer than the third, and slightly shorter than 
the clavate fourth. Labium reaching to metasternum. Pronotum with the lateral 
margins very slightly sinuate, not much wider basally than apically, toothed minutely 
at the antero-lateral angle. $ not remarkable. $ 5th sternite apically obtusangulate, 
6th acutangulate. Length 45 mm. 

Hab. Kauai, high plateau (Aug., Perkins), Waimea Mts., 4000 ft. (May, Perkins). 

This species is easily recognized among the pale glabrous forms by the short, 
pyriform, laevigate areas on the pronotum. 



Nysius kantehameha Kirkaldy. 

Nysiiis kamehameha Kirkaldy, huj. op. iii. p. 164. 

Hab. Hawaii, Hualalai, 5000 ft. (Aug., Perkins). 

I have seen no specimens beyond the carded type of this, for a Nysius, handsome 
species. The labium extends practically as far as the hind coxae. The tegmina are 
parallel for about one-seventh of their length, thence gently rounded. The exterior 
half of the metapleura black, the hind margin and the interior half, together with the 
orifice, pale. 

F. H. II. 70 



540 FAUNA HAWAIIENSIS 



(8) Nysms coenosuhis Stal. 

Nysius coenosulus Stal, 1859, Eugenics Resa, Zool. p. 243; F. B. White, 1878, 
A. M. N. H. (5) I. p. 369. 

Through the kindness of Dr Aurivillius, I was able to see the type — unfortunately 
in poor condition — of this doubtless introduced species. It occurs in mixed sweeping, 
principally, I think, on Erigeron canadensis. 

Hab. Oahu, near Honolulu (Perkins and others) ; Waianae Coast (March, 
Perkins). — Lanai, 3000 ft. (Perkins). — Hawaii, 2000 — 4000 ft. (Perkins). 



(9) Nysms blackburni White. 
Nysius blackburni F. B. White, 1881, A. M. N. H. (5) vii. p. 53. 
Hab. Hawaii, Kilauea, 4000 feet on ferns (White). I do not know this. 

(10) Nysius lichenicola, sp. nov. 

Head and pronotum yellowish testaceous, closely punctured, and more or less 
suffused, with dark brown ; lateral margins of head, anterior third of pronotum, 
posterior margin very narrowly (usually interrupted medially and at the lateral angles), 
scutellum etc., dark. Tegmina typically yellowish testaceous, closely irrorated with 
blackish grey, usually more so towards the middle, apical angle of corium dark. 
Membrane vitreous, variegated varyingly with brownish grey, apical angle usually 
blackish. Antennae, labium, femora etc., blackish or piceous ; coxae and apex of 
femora testaceous or pale ferruginous. Underside black, orifices dark testaceous. 
The fourth segment of the antennae is about twice as long as the third, and a little 
longer than the second. Pronotum much as in N. coenosulus. Length ^ 3^, 
¥ 3i— 4 mm- 

Hab. Hawaii, Kilauea, among the lichen on the trunks of fallen Nani polymorpha 
and Acacia koa (April, Kirkaldy) ; Kaumana, above Hilo, 2000 ft. (Jan., Perkins, 
no. 679). — Maui, Haleakala, 5000 — 9000 ft. (March, Apr., Perkins). Dr Perkins tells 
me he has taken it from various fruits, Mynah-bird's dung etc. This species is possibly 
not endemic. 

It is variable, but I cannot find structural differences. In a Mauian female, the 
tibiae are more testaceous and the apical angle is almost entirely pale. In one male, 
the hind margins of the pleura are paler, but much more material is necessary for 
comparison, before elevating the following varieties to possible specific rank. 



HEMIPTERA 541 

var. (<5) brunnealis nov. 
Tegmina pale ferruginous inwardly. 
Hab. Haleakala, 5000 ft. (April). 

var. {c) atralis nov. 
Upper surface blackish brown ; a small spot at base of head, one at middle and 
one at each lateral angle of hind margin of pronotum, pale ; a few testaceous specks on 
tegmina, mostly near the lateral margin. 

Hab. Haleakala, 5000 ft. (April). 

The type of this species is one of the specimens from Haleakala. 

(11) Nysius ochriasis Kirkaldy. 
Nysius ochriasis Kirkaldy, 1902, huj. op. iii. 162. 
Food plant : Duhautia sp. 

(12) Nysius insulicola, sp. nov. 

Similar in general appearance to N. mauiensis, but the pronotum is maculate only 
at the postero-lateral angles. 

Head black ; clypeus and a ring around the eyes, brownish yellow. Antennae pale 
brownish yellow, indistinctly infuscate medially. Labium pale yellow, apically blackish. 
Pronotum brownish yellow (callosities browner), confusedly and comparatively sparsely 
brownly punctured, postero-lateral angles blackish brown. Scutellum brownish yellow, 
basally and laterally punctured with dark brown, posterior angle whitish. Tegmina 
subhyaline, pale brownish yellow, commissure and a short oblique stripe apically on 
exocorium dark fuscous. Membrane hyaline, a broad fuscous stripe down the middle. 
Beneath pale yellowish brown with a pink tinge ; sterna mostly blackish, abdomen 
medially pale yellow ; ambulacra and orifices whitish ; legs pale yellow, femora spotted 
with dark brown, hind femora with a dark brown subapical ring. Second and third 
segments of the antennae subequal, each longer than the fourth. Pronotal callosity 
percurrent ; lateral margins very slightly sinuate. % 6th and 7th sternites acute- 
angularly emarginate apically. Length 5 mm. 

Hab. Lanai, over 2000 ft. (Jan., Perkins). 

(13) jVysius silvestris, sp. nov. 

Head black, clypeus pale yellow, ocelli red ; eyes black, interiorly margined with 
testaceous. Antennae and labium pale brownish testaceous, the former with segments 
I — 3 feebly ringed with pale fuscous, 4th pale ferrugineotestaceous, apically pale 
fuscous, the apex of labium blackish. 

70—2 



542 FAUNA HAWAIIENSIS 

Pronotum pale ferruginous, with 5 longitudinal, parallel, elongate spots (or short 
stripes) on the basal third, the middle one extending anteriorly as far as the 
transverse swelling, the sublateral ones sometimes rather indistinct. Scutellum piceous, 
basal angles pale ferruginous, posterior angle white. Tegmina piceous, extero-lateral 
margin on basal half brownish yellow. Membrane hyaline, basal third dark fuscous. 
Sterna polished black, pleura brownish yellow tinged with red, and more or less marked 
with blackish. Legs and ambulacra pale, apical half of hind femora rosy. Abdomen 
beneath pale reddish brown, more or less clouded with blackish on sternites 2 — 6. 
Orifices pale, slightly rosy. Three apical segments of antennae subequal, the fourth 
a little the longer, rather elongately incrassate. Pronotal callosities percurrent, 
pronotum rather sparsely punctured. 

$ 6th sternite very obtusely, 7th acutely, emarginate. Length 3^ — 4 mm. 

Hab. Oahu, Waianae Mts., about 3000 ft., Feb. (no. 547). 



(14) N'ysius oresitrophus, sp. nov. 

Black ; apex of clypeus, apex of first, second and third segments of antennae, 
a small spot on middle of hind margin of pronotum, labium, legs, etc. brownish 
testaceous or brownish yellow, femora and tibiae broadly banded with piceous ; tarsi 
partly fuscous. Postero-lateral angles of pronotum and posterior angle of scutellum 
yellowish testaceous. Tegmina yellowish testaceous, a large spot at base, another at 
intero-apical angle of corium, and a larger one on apical margin of corium, black. 
Membrane yellowish brown, apparently basally blackish. Antennae with the second and 
fourth segments subequal, a little longer than the third. Labium reaching well beyond 
the hind coxae. Pronotum rather superficially punctured, almost striate. Length 
4^ mm. 

Hab. Maui, Haleakala, over 5000 ft. (Oct., Perkins, no. 636) i carded example. 



(15) Nysius nubicola, sp. nov. 

Head blackish with a red line down the middle ; antennae reddish or reddish 
testaceous. Pronotum reddish or reddish testaceous punctured with blackish, the 
laevigate areas blackish (sometimes reddishly suffused). Scutellum rufescent, punctured 
with blackish. Tegmina yellowish hyaline, in some examples almost immaculate, in 
others blotched apically. Labium, legs, ambulacra, orifices, etc. rufescent or yellowish 
testaceous, apical segment of labium black, femora spotted with black. Underside 
black, the pleura partly rufescent, abdomen varyingly pale in part. Head and pronotum 
sparsely pubescent. Labium extending beyond hind coxae, first segment reaching to 
apical margin of prosternum. Second .segment of antennae distinctly longer than the 



HEMIPTERA 543 

third, which is longer than the fourth. Pronotum closely punctured ; lateral margins 
scarcely sinuate, not at all carinate. Length 4 — 5^ mm. 

Hab. Hawaii, Kona, 4000 ft. and over, Aug. (no. 231, Perkins); Hualalai, 5000ft. 
(Perkins). One of the Kona specimens is the type. 

(16) Nysitis neniorivagus White. 

Nysius neniorivagus F. B. White, 1881, op. cit. 54. 

Hab. Hawaii, Mauna Kea. — Maui, Haleakala, 5000 — 6000 ft. I do not know 
this. 

( 1 7) Nysius nimbatus, sp. nov. 

Head and most of underside black. Pronotal callosities piceous. Antennae, 
pronotum, scutellum, tegmina etc. pale olive-testaceous, a brown irregular ^ near the 
middle of the apical margin of the corium. Pronotum and scutellum with blackish 
brown punctures, the anterior half of the former sometimes being suffused with black. 

Legs, orifices etc. pale brownish yellow, fore and middle femora sparsely spotted 
with pale brown, hind femora strongly spotted with blackish brown. Abdomen beneath 
sometimes largely pale apico-medially. Antennae in some examples tinged or marked 
with brown. Second and fourth segments of antennae subequal, each shorter than the 
third. Pronotum rather sparsely punctured, laevigate areas reaching the lateral margins, 
which are very slightly sinuate. Length 5^ — 6^ mm. 

Hab. Oahu, Honolulu, 2500 — 3000 ft., Oct. — Nov. (Perkins). 

(18) Nysius rubescens White. 

Nysius rubescens F. B. White, 1881, A. M. N. H. (5) vii. 55. 

This was described from Hawaii, Kilauea, 4000 ft. 1 do not know it. 

(19) Nysius pteridicola White. 

Nysitis pteridicola F. B. White, 1881, 1. c. 

This was described from the same place as the preceding. I do not know it. 

(20) Nysitis nitidus White. 
Nysius nitidus F. B. White, 1881, A. M. N. H. (5) vii. 53. 
Hab. Maui, Haleakala. I do not know it. 

(21) Nysitis vtdcan White. 
Nysius vulcan F. B. White, 1881, op. cit. 56. 

Hab. Hawaii, Mauna loa, elevation not stated. I do not know it. 



544 FAUNA HAWAIIENSIS 

{22) Nysius oribasus, sp. nov. 

Elongate. Head black. Antennae, pronotum and scutellum piceous ; base and 
apex of first three segments of antennae pale, as also angles and keel down scutellum, 
and postero-lateral angles of pronotum. Labium, ambulacra, orifices and legs pale ; the 
first and fourth segments of the labium, and the tibiae partly, fuscate ; femora thickly 
spotted on the basal half, or so, with dark brown, coxae basally piceous. Tegmina 
hyaline, strongly blotched with dark brown. Beneath blackish, abdomen in one 
example a little pale in part. Second segment of antennae slightly longer than the 
third, which is longer than the fourth. Pronotum coarsely punctured ; lateral margins 
very slightly sinuate, not at all carinate ; laevigate areas blackish. 

Hab. Oahu, Waialua, Koolau range, 2000ft., Feb. (Perkins); Kaala, 1500 ft. 
Jan. (Perkins). The Waialua specimen is the type. 

(23) Nysius insiclivagus, sp. nov. 

Sparsely yellowish pubescent. Head, sterna and pleura black, propleura partly 
brownish piceous. Pronotum, scutellum, and tegmina brownish piceous ; laevigate 
areas of pronotum rather darker ; scutellum partly darker, carina down the middle 
yellow ; tegmina very closely mottled pallidly, veins paler, especially the forked median. 
Beneath, the abdomen is piceous, becoming more palely ferruginous towards the apex. 
Antennae, labium and legs pale ferruginous ; apex of first three segments of antennae 
dark ; orifices pale. First segment of labium extends beyond the apical margin of the 
prosternum. Second segment of antennae distinctly longer than the third, which is 
subequal to the fourth. Lateral margins of pronotum scarcely sinuate, obsolescently 
carinate. 

Hab. Hawaii, Hualalai, 8000 ft., Aug. (Perkins) ; Kilauea, 4000 ft. (Perkins) and 
at the same locality on Nani polymorpha (" Ohia lehua," April). — Maui, Haleakala, 
10,000 ft., Aug. (Perkins). One of the Hualalai examples is the type. 

(24) Nysius montivagus, sp. nov. 

Differs from the preceding by the glabrous, more shining appearance, paler 
pronotum (laevigate areas dark), darker antennae, maculate femora etc. 

Hab. Lanai, Molokai, Hawaii. — Lanai, Halepaakai, July (Perkins). — Molokai, 
4000 ft. (May, Perkins). Hawaii, Kilauea (Kirkaldy). 

(25) Nysius monticola, sp. nov. 

Superficially somewhat like N. ochriasis, but larger and more sordid, the fourth 
segment of the antennae dark, and the pronotum laterally sinuate. 

Yellowish brown, a paler clear line on the vertex along the middle. Eyes blackish 



HEMIPTERA 545 

brown, ocelli reddish brown. A ring at base and another near apex of second 
segment, one at base of third, and all the fourth (except extreme base) segment of 
antennae, dark fuscous. Apex of labium black. A clearer line on anterior two-thirds 
of pronotum down the middle, clearer also medio-laterally ; rather sparsely punctured 
with brown, as also on pleura. Scutellum punctured with dark brown on the basal 
margin and between the keels and the lateral margins. Tegmina and membrane 
immaculate brownish yellow, veins concolorous. Ambulacra, orifices and legs im- 
maculate brownish yellow. The whole insect glabrous, shining. Pronotum laterally 
sinuate, roundedly narrowing laterally in front of the callosity, slightly reflexed in 
part ; impressed behind the callosity. Antennae with the second and third segments 
subequal, a trifle longer than the fourth. Labium reaching nearly to the middle of 
the abdomen, first segment beyond the base of the head. % 6th sternite very obtusely, 
7th rectangularly, emarginate. Length 6 mm. 

Hab. West Maui Mts., 4000 ft. (no. 357, April, Perkins, i %)\ 



( I ) Orthoea vincta, Say. 

Pamera vincta (Say) Stal. 

Orthoea periplanios Kirkaldy, 1907, Canad. Ent. xxxix. p. 246. 

O. pacifica Kirkaldy, 1907, P. Haw. E. S. i. 150; and 1908, P. Linn. Soc. N.S. W. 

O. vincta Kirkaldy, 1908, P. Haw. E. S. i. 189 and P. Linn. Soc. N. S. W. 

This little bug "swarmed in 1900, but was not seen prior to 1897 " (Perkins). It 
has been described under 8 names and has spread over about the whole of America, 
and occurs in Ceylon, India, Burma ; Christmas Islands (Indian Ocean) ; South Africa ; 
Australia, Tahiti and Fiji! I have described the last nymphal instar (1907, P- Haw. 
E. S. I. 150), which has also with long- and short-winged adults, been figured (1908, 
P. Linn. Soc. N. S. W.). In Hawaii, O. vincta is found in Cynodon dacty/on, but does 
not, I think, feed on it. 

Hab. I have collected, or seen, specimens from Kauai, Oahu, Maui and Hawaii, 



Clerada apicicornis Sign. 

Clcrada apicicornis Sign., huj. op. iii. p. 160. 

I have described the final nymph (1907, P. Haw. E. S. i. 151). Dr Perkins has 
seen this species " feeding on a dead Blattid (which had perhaps died before it was fed 
upon) " and does not think it catches Lepisma. I have only seen this on Oahu. 

1 The printed label bears the indication "West Maui 4°oo ft.," but no. 357 (on the underside of the card 
to which the specimen is affixed) refers to " Haleakala 4000 ft.," which is in East Maui. 



546 FAUNA HAWAIIENSIS 

Merragata hebroides White. 

Merragata hebroides, huj. op. iii. p. i68. 
Common and widely distributed \ 

Fam. NABIDAE. 

Vide Subfam. Nabinae, huj. op. iii. p. 153. 
Conspectus generum ac siibgene7'um hawaiiensium. 

1. Caput ordinarium i Reduviolus W. Kirby (2). 

\a. Caput utrimque spina obtusa, prominente; segmentum primum antennarum 

incrassatum (subj. 2) Miln Kirkaldy. 

2. Clavus ac corium baud discreta ; membrana perminuta ; ocelli plus minus 

obsolescentes (subj. \b) Nesotyphlias Kirkaldy. 

za. Tegmina ordinaria; ocelli adsunt (3). 

3. Scutellum immaculatum ; antennae pedesque baud annulatae ; margines 

laterales pronoti fere recti, lobo postico pronoti vix elevato... (subg. \a) Nesomachetes Kirkaldy. 
2,a. Scutellum maculatum ; antennae pedesque annulatae ; margines laterales 

pronoti sinuati, lobo postico pronoti elevato (subg. typicum). 

Reduviolus. 
(i) Reduviolus kahavalu Kirkaldy. 

Reduviolus innotatus Kirkaldy, huj. op. iii. p. 154, PI. v. fig. 32. 
R. kahavalu Kirkaldy, 1907, P. Haw. E. S. i. p. 156. 

Delete the Oahuan record. I have founded a subg. Nesomachetes for this ( 1 908, 
P. Haw. E. S. I. p. 190). 

Reduviohis capsiformis Germar. 

Nabis capsifoj'mis Germar, 1837, Silb. rev. Ent. v. p. 132. 

Reduviolus innotatus F. B. White, 1877, A. M. N. H. (4) xx. p. 112; Kirkaldy, 

1907, P. Haw. E. S. I. p. 156. 
R. blackburni (part) Kirkaldy, huj. op. iii. p. 155 ; Perkins, 1903, Bull. Board 

Agr. Hawaii Ent. i. ; Swezey, 1905, Bull. H. S. P. Ent. i. p. 234, PI. 17, 

figs. 1—4. 
R. capsiformis Reuter, 1908, Mem. Soc. ent. Belg. xv. p. 114. 
The adoption of the above name is on Reuters authority ; in his remarks on 
p. 115 anent the figure of R. innotatus, he has overlooked my paper in the P. Haw. 
E. S. I. 156. R. capsiformis has been described, or alluded to, under eleven names, 

' A Tingid of the genus Teleonemia is now firmly established in the Islands, but has nothing to do with 
the original fauna, having been purposely introduced to check the Lataiia camara. 



HEMIPTERA 547 

and is practically cosmopolitan. It has nothing to do with R. blackburni which is 
a much darker, probably endemic, insect, found at Kiiaucc, Hawaii and elsewhere, 
ferns, trasses etc. 



on 



Reduviolus niorai Kirkaldy. 

Reduviohis worae Kirkaldy, huj. op. in. p. 155, PI. v. fig. 39 ; and 1908, P. H. E. S. 

I. 191. 
Delete Lanai from the islands. 

Reduviolus subrufus White. 
Reduviolus subrufus, huj. op. in. p. 156. 

Delete R. koelensis and oscillans from the synonymy; only fig. i"], in the "Fauna," 
applies to this species. 

(2) Reduviolus koelensis Blackburn. 

Nabis koelensis Blackburn, 1888, P. Linn. Soc. N. S. W. (2) in. p. 352. 

Hab. Lanai, Koele. 

1 am not sure that I know this form, but at least it cannot, I think, be included 
under subrufus. 

(3) Reduviolus oscillans Blackburn. 

Nabis oscillans Blackburn, 1. c. 
The same remarks apply to this. 

Hab. Hawaii, Mauna loa, about 4000 ft. 

(4) Reduviolus arrogans Kirkaldy. 
Reduviolus arrogans Kirkaldy, 1908, P. H. E. S. i. p. 191. 
Hab. Molokai. 

(5) Reduviolus trtictdenhis Kirkaldy. 

Reduviolus subrufus Kirkaldy, 1902, huj. op. iii. PL v. fig. 38, but not of the 

text p. 156. 
R. truculentus Kirkaldy. 1908, P. H. E. S. i. p. 191. 

Hab. Oahu, Honolulu Mts. on Pipturus albidus. 

(6) Reduviolus nubigemis Kirkaldy. 
Redtiviolns nubigenus Kirkaldy, 1908, 1. c. 

Hab. Lanai — also, I think, Maui and Molokai. 
F. H. II. 71 



548 FAUNA HAWAIIENSIS 

(7) Reduviolus kaonohiula Kirkaldy. 

Rcduviolus kaonohiula Kirkaldy, 1908, op. cit. p. 192. 
In fresh examples, the abdomen is grass-green. 

The last nymph, which, with the adult, is found on CyatJiodes tameiameiae at 
Kilauea, Hawaii, is grass-green with yellow markings. 

(8) Reduviolus montivagiis Kirkaldy. 

Reduviolus montivagus Kirkaldy, 1908, 1. c. 
Hab. Kauai. 

{c) Reduviolus subgen. Nesotvphlias. 
Kirkaldy, 1907, P. H. E. S. i. 155. 

Reduviolus lusciosus White. 

Nabis ? lusciosus F. B. White, 1877, A. M. N. H. (4), xx. 112. 

Reduviolus lusciosus Kirkaldy, huj. op. in. p. 157 (part), PI. v. fig. 35; Reuter, 

1908, Mem. Soc. ent. Belg. xv. p. 124. 
Nesotyphlias lusciostis Kirkaldy, 1907, P. H. E. S. i. p. 155. 
Hab. Oahu. Not yet found on any other of the islands. 

(9) Reduviolus silvicola Kirkaldy. 

Redtiviolus silvicoia Kirkaldy, 1908, P. H. E. S. i. 192. 
Hab. Molokai. 

(10) Redtiviolus monticola Kirkaldy. 
Reduviolus monticola Kirkaldy, 1908, 1. c. 

Hab. Oahu. 

(11) Reduviolus procellaris Kirkaldy. 
Reduviolus procellaris Kirkaldy, 1908, op. cit. 193. 

Hab. Molokai. 

(12) Reduviolus volcanicola Kirkaldy. 
Reduviolus lusciosus Kirkaldy, huj. op. in. PI. v. figs. 34 and 340;. 
R. volcanicola Kirkaldy, 1908, P. H. E. S. i. p. 193. 

Hab. Hawaii, Kilauea, on tree-ferns and among dead leaves on the ground. 



549 



HEMIPTERA 

{13) Reduviolus curtipennis (Blackburn). 
Nabis ? curtipennis Blackburn, 1888, op. cit. 353. 
Hab. Hawaii, Waimea. 

(14) Reduviolus paludicola Kirkaldy. 
Reduviolus paludicola Kirkaldy, 1908, 1. c. 
Hab. Molokai. 

(15) Reduviolus lolupe Kirkaldy. 
Reduviolus lolupe Kirkaldy, 1908, 1. c. 
Hab. Uncertain. 

(16) Reduviolus silvestris Kirkaldy. 
Reduviohis silvestris Kirkaldy, 1908, op. cit. 194. 
Hab. Kauai. 

{d) Reduviolus subgen. Milu Kirkaldy. 

Milu Kirkaldy, 1907, Can. Ent. xxxix. p. 247. 

Reduviolus subg. Milu, Reuter, 1908, Mem. Soc. ent. Belg. xv. p. 109. 

(17) Reduviolus kerasphoros Kirkaldy. 

Reduviolus 7'ubritinctus Kirkaldy, huj. op. in. p. 157, PI. v. fig. 33, nee Blackburn. 

Milu kerasphoron Kirkaldy, 1907, Can. Ent. xxxix. p. 248. 

M. kerasphoros Kirkaldy, 1908, P. Haw. E. S. i. p. 194; with var. purpurea, 

P- 195- 
Reduviolus kerasphoron Reuter, 1908, 1. c. 

Hab. Oahu. , 

Reduviolus rubritinctus Blackburn. 

Nabis rubrilinctus, Blackburn, 1889, P. Linn. Soc. N. S. W. (2) in. p. 351. 
Milu ? rubritinctus Kirkaldy, 1908, 1. c. 

Hab. Maui. 

I have not seen this species, which is apparently distinct from the preceding. 

71 — 2 



550 FAUNA HAWAIIENSIS 

Fam. REDUVIIDAE. 

Zelus renardii Kolenati. 

Zehis renardii Kolenati, 1856, Bull. Soc. Nat. Moscou, xxix. 460, PI. 3, fig. 2 ; 

Kirkaldy, 1908, P. Haw. E. S. i. 195. 
Z. laevicollis Champion, 1899, B. C. A. Het. 11. 252 and 260, PI. w. fig. 24. 
Z. peregrimis Kirkaldy, huj. op. in. p. 149; Perkins, 1903, Bull. Agr. Hawaii 
Ent. I. 20; Kirkaldy, 1904, Haw. For. Agr. i. 183; Swezey, 1905, Bull. 
H. S. P. Ent. I. 232, PI. XVI. figs. I- — 3; Kirkaldy, 1907, P. Haw. E. S. i. 
156; and Canad. Ent. xxxix. 247. 
This species was first seen in Honolulu in 1897, but had become common and 
generally spread over Oahu by 1900. It is now common in the lowlands of all the 
islands, the brown ^<g^ masses being very conspicuous on Saccharum officinarum. 
Hibiscus rosasinensis, Citrus aurantiiwi etc. In the younger stages, it feeds on 
Aphids, young leafhoppers etc., but the adult preys on several very destructive beetles 
as well, though sometimes becoming injurious by sucking Coccinellids. It is apparently 
distributed over the Western States (California, Arizona etc.) and Mexico. 
Swezey' and I'- have contributed to the knowledge of the life-history. 



Triatoma Lap. 

Triatoma Laporte, 1832, Essai Hem. pp. 6 and 11. 
Conorhinus Laporte (1832 ?) Essai Hem. p. "]"]. 

(i) T. rubrofasciata de Geer. 

Cifuex riibrofasciatus de Geer, 1773, Mem. in. 349, PI. xxxv. fig. 12. 

Triatoma rubrofasciata Kirkaldy, 1904, Haw. For. Agr. i. 185. 

This has been taken by Koebele and Perkins around labourers' huts on certain 
Oahuan plantations. It is a native of Brazil, the Antilles etc., but has become 
distributed over the Philippines, China, Burma, Malay Peninsula, Ceylon, Andamans, 
India, West Africa and Madagascar. 

Luteva insolida White. 

Luteva insolida White, huj. op. in. p. 152. 

This occurs also around Honolulu ; it is probably not endemic. 

' 1905, op. cit. pp. 232-4, PI. XVI. figs. I — 3. 
' 1907, P. Haw. E. S. I. pp. 156-8. 



HEMIPTERA 551 

(i) Luteva insulicola Kirkaldy. 
Luteva insulicola Kirkaldy, 1908, P. Haw. E. S. i. p. 196. 
Hab. Oahu. Waialua. 

(i) N'esidiolesles insularis Kirkaldy. 

Nesidiolestes insularis Kirkaldy, 1908, op. cit. p. 195. 

Hab. Oahu, Tantalus, 1800 ft. 

I have seen only a single example of this, taken up Tantalus by Mr O. H. Swezey, 
and now in my collection. 

Fam. ANTHOCORIDAE. 
Vide Miridae subfam. Anthocorinae, huj. op. in. p. 125. 

Triphleps persequens White. 

Triphleps persequens White, huj. op. in. p. 125. 

This is distributed throughout the islands in the plains, being very common in 
sugar-cane fields and on cultivated Hibiscus etc. It is doubtless introduced. It feeds 
on Aphidae, Psocidae, and other small insects. Swezey' has described the nymph 
as yellowish, and has figured the adult. T. persequetis is now known from Queensland 
and Fiji". 

PhysopleuTella mundula White. 

Physopleurella mundulus White, huj. op. in. p. 1 26. 

This is also widely distributed similarly to the last, and has similar food-habits. 
Swezey has briefly described and figured the ova and nymphs, as well as the adult". 

Lasiochihis dcnigratus Wliite. 

Lasiochilus denigrata White, huj. op. in. p. 126. 

I followed Reuter in placing L. decolor as a synonym, but, as I now think, wrongly. 
So far as the material now before me goes, L. denigratus is restricted to Hawaii. 

Hab. Hawaii. 

* 

(i) Lasiochilus decolor White. 

Dilasia (f) decolor White, 1879, Ent. Mo. Mag. .xvi. p. 147. 

Lasiochilus denigratus (part) Reuter ; and L. denigrata (part) Kirkaldy, huj. op. 

III. p. 126. 
Hab. Oahu. This seems to be restricted to Oahu. 

1 1905, Bull. H. S. P. Ent. I. 23s, PI. XVI. fig. 7. 
- 1908, Kirkaldy, P. Linn. Soc. N. S. W. xxxii. 784. 
" 1905, loc. cit., PI. XVI. figs. 4—6. 



552 FAUNA HAWAIIENSIS 

(2) Lasiochilus silvicola Kirkaldy. 
Lasiochilus silvicola Kirkaldy, 1908, P. Haw. E. S. i. 196. 
Hab. Kauai, Koholuamano. 

(3) Lasiochilus montivagus Kirkaldy. 
Lasiochilus mofitivagtis Kirkaldy, op. cit. 197. 
Hab. Lanai, Koele Mts. (also doubtfully from Hawaii). 

(4) Lasiochihis mibigenus Kirkaldy. 
Lasiochilus nubigenus Kirkaldy, 1. c. 
Hab. Maui, Haleakala. 

Fam. CLINOCORIDAE. 

Subfam. CACODMFNAE, huj. op. III. p. 129. 
For Klinophilus use Clinocoris Fallen 1829. 

Fam. MIRIDAE. 

Subfam. MIRINAE, huj. op. ill. p. 129. 

Sulamita lunalilo Kirkaldy. 

Sulaniita lunalilo, huj. op. iii. p. 130. 

The type was a specimen from Kilauea, Hawaii. PI. IV. fig. \2a refers to this. 

(i) Sulamita dryas Kirkaldy. 

Sulamita lunalilo, var. Kirkaldy, huj. op. in. PI. iv. fig. i 2. 
S. dryas Kirkaldy, 1908, P. Haw. E. S. i. p. 197. 

(2) Sulamita oreias Kirkaldy. 

Sulamita lunalilo, brach. form, Kirkaldy, huj. op. iii. PI. iv. fig. 13. 
S. oreias Kirkaldy, 1908, P. Haw. E. S. i. 197. 

TiCHORHiNUS Fieber. 

Tichorhinus Fieber, 1858, Wien. E. Mon. 11. p. 314. 

Orthotylus Fieber, 1858, op. cit. p. 315 ; Kirkaldy, 1902, huj. op. iii. p. 132. 



HEMIPTERA 553 

(i) Tichorhinus kassandra Kirkaldy. 
Orthotylus daphne var. kassandra Kirkaldy, huj. op. in. p. 135. 

Cyrtopeltis hawaiiensis Kirkaldy. 

C. hawaiiensis, huj. op. iii. p. 138. 

Hab. Add, Oahu, Tantalus (Dec. Perkins, Swezey, Giffard) ; Maunawili (Giffard) 
on Touchardia. 

An error has crept into the description, viz. the base of the pronotum is wider 
than half the length of the second segment of the antennae. 

Dr Perkins informs me this species is rather variable, some specimens being much 
darker. 

Opuna hawaiiensis Kirkaldy. 
Opuna hawaiiensis, huj. op. in. p. 140. 
Hab. Oahu, on Sida cordifolia. 

(i) Pseudoclerada kilaueae Kirkaldy. 

Pseudoclerada morai ^a.rt, Kirkaldy, huj. op. in. p. 141, PI. iv. fig. 19. 
P. kilaueae Kirkaldy, 1908, P. H. E. S. i. p. 198. 

Hab. Hawaii, Kilauea. 

Kalania Kirkaldy. 

Baracus Kirkaldy, huj. op. in. p. 143. 

Kalania Kirkaldy, 1904, Entom. xxxvii. p. 280. 

Kalania hazvaiiensis. 

Baracus hawaiiensis, huj. op. in. p. 143. 

Hab. This seems restricted to Lanai. It was retaken there by Giffard (1908, 
P. H. E. S. I. 180). 

Hyalopeplus pellucidus Stal. 

Hyalopeplns pelhtcidus, huj. op. in. p. 143. 

This has been redescribed from the type by Reuter (1905, Ofr. Finska Vet. Forh. 
XLVii. no. 12, p. 2). It is distributed from sea level up to 5000 ft. and is certainly not 
endemic. Near Honolulu it is common on Hibiscus rosasinensis, but swarms, further 
from the town, on Guava, Sida, Dodonaea etc. I have described the last nymph (1907, 
P. H. E. S. I. 159). 



554 FAUNA HAWAIIENSIS 

Koanoa hawaiiensis Kirkaldy. 

Koanoa hawaiiensis, huj. op. in. p. 1 36. 
This is arboreal and endemic. 

Nesiomiris hawaiiensis Kirkaldy. 

Nesiomiris hawaiiensis, huj. op. in. p. 145. 

This also is arboreal and endemic. On the other hand Oronomiris hawaiiensis is 
graminivorous and an introduction. 

Acanthia exulans White. 
Acanthia exulans, huj. op. iii. p. 146. 

Hab. Oahu. I can only definitely refer this typically now from Oahu, possibly 
also from Kauai. I have described (1908, P. Haw. E. S. i. p. 198) var. molokaiensis 
from Molokai. 

Acanthia oahuensis Blackb. 
Hab. Oahu. I can refer this also only from Oahu now. 

(i) Acanthia humifera Kirkaldy. 
Acanthia humifera Kirkaldy, 1908, op. cit. p. 199. 
Hab. Oahu, N. W. Koolau Range. 

(2) Acanthia nubigena Kirkaldy. 
Acanthia nubigena Kirkaldy, 1908, 1. c. 
Hab. Maui, Haleakala. 

(3) Acanthia procellaris Kirkaldy. 
Acanthia procellaris Kirkaldy, 1908, op. cit. p. 200. 
Hab. Molokai, 4000 ft. 

Arctocorisa blackburni White. 

Corixa blackburni, huj. op. in. p. 148. 

Corixa blackburni is now referred to Arctocorisa, recently raised to generic rank. 

BuENOA Kirkaldy. 

Buenoa Kirkaldy, 1904, Wien. E. Z. xxni. p. 120. 
Included under Anisops, huj. op. in. p. 148. 



HEMIPTERA 555 

(i) Biienoa pallipes Fabricius. 
Notonecta pallipes Fabricius, 1803, Syst. Rh. 103. 
Anisops sp. ? Kirkaldy, huj. op. in. p. 148. 
Buenoa pallipes Kirkaldy, 1904, Wien. E. Z. xxiii. p. 123. 

Homoptera. 

Fam. MEMBRACIDAE. 

A single introduced species (from New Zealand, where doubtless introduced from 
Australia) occurs. I have seen a specimen in Dr Perkins' collection. It is a small 
Centrotine, probably a Centrotypus sp. 

Fam. TETTIGONIIDAE. 

The Hawaiian species — endemic and introduced — belong entirely to the tribe 
Phrynomorphini, though Tettigoniini and Eurynielini have been recorded in error. 
They are comprised in three endemic, closely related, genera, and three more or less 
cosmopolitan. 

Tribe Phrvnomorphjnj'^. 

Conspectus generuni. 

1. Venae alarum prima secundaque apicem versus baud confluentes (2) 

irt. Hae venae apicem versus confluentes 5 Nesosteles Kirkaldy- 

2. Tegmina cellulis subapicalibus duabus instructa (3). 

2a. Tegmina cellula subapicali unica instructa (5). 

3. Cellula subapicalis exterior (minor) basi et apice pedicellata 2 Nesophrosyne Kirkaldy. 

2,a. Cellula subapicalis exterior (minor) baud et basi et apice pedicellata ...(4). 

4. Vertex brevis ac latus i Nesophryne Kirkaldy. 

i,a. Vertex antice ultra oculos extensus 3 Conosaniis Osborn and Ball. 

5. Cellula subapicalis tegminis subovalis, medio paullo constricta 2a (subg.) Nesoreias nov. 

5«. Cellula subapicalis basi acutangularis 4 MacrosUles Fieber. 

Hie conspectus consanguinitatem non monstrat ; Nesosteles e Macrosteli, 
Nesoreias e Nesophrosyni derivatae. 

On the face of many Phrynomorphini, there is a characteristic pattern of two dark 
stripes up the middle, with more or less concentrically curved, subparallel, transverse 
stripes. Sometimes this is very well marked, sometimes very obscurely, sometimes 
absent. I propose, for the sake of brevity, to term this the "craticuliform pattern" or 
"craticulum," i.e. the "grill." 

' For horismology etc. see my recent classification of the .\uchenorrhyncha, Bull. Haw. Plant. Ent. i. 
269—479, Pis. 21—32 (1906) and op. cit. iii. i— 186, Pis. 1—20 (1907). 

F. H. II. 72 



556 FAUNA HAWAIIENSIS 

Nesophryne Kirkaldy. 

Nesophryne Kirkaldy, Proc. Haw. Ent. Soc. i. p. i6o, 1907. 

Allied to Eutettix Van Duzee, but the habitus is quite different, and the vertex 
much shorter and more declivous, with the eyes scarcely so wide as the transversely 
striolate pronotum. Frons very wide, almost obliterating the fasciolae ; lateral margins 
sloping roundly to the clypeus, the sides of the latter very slightly roundly emarginate. 
Antennae inserted a little nearer to the ocelli than to the base of the lora ; they have a 
short flagellum. Clypeus about twice as long as its width, the lora do not nearly reach 
its apical margin. The labium reaches to the apex of the middle coxae. Tegminal 
venation not altogether unlike that of Eutettix, as represented by E. sellata and 
melaleucae, but the two subapical cells extend equally far apically and are truncate 
there, while the long median cell is apically angular. There is a well defined appendix, 
four apical cells and a nodal cell. Wing venation normal. Type filicicola. The 
genus is confined to Kauai. 

There are probably four species, as it is very likely that my Bythoscopiis kukanaroa 
and kamaiainao belong here. The declivous vertex and the position of the ocelli on 
the apical part of the bend of the head, probably led me to place them in Bythoscopns. 
I cannot refer to them now, however. The two species known to me are separable 
thus : 

I. Tegmina brunneo-flavescentia venis fuscis, cellulis sparse medio brunneo- 

irroratis. Valvula maris pallida ; pygophor feminae brunneo-flavescens ...\ filicicola Kirkaldy. 

1(7. Tegmina forte nigrescenti-suffusa, clavo paucis areis maculisque albidis ornato; 
cellula subcostal! partim brunneo-flavescenti, subhyalina. Valvula maris 
nigra; femina mihi incognita 2 microkpiae sp. nov. 

(i) Nesophryne filicicola Kirkaldy. 

Nesophryne filicicola Kirkaldy, 1907, 1. c. 

t yellowish ; pronotum tinged with greenish, sparsely clouded etc. with dark 
fuscous and olive brown. A large transverse band on the bend of the head, dark 
fuscous, enclosing the conspicuous yellowish red ocelli ; a central line down the clypeus, 
most of the lora, and a spot on the genae, dark fuscous. Tegmina yellowish cinereous, 
veins olive brown, the principal cells obscurely irrorate inside in the middle. The long 
median cell has two subhyaline spots on its apical half, the interior subapical has two, 
and there are others, not so distinct, on the other cells. Wings smoky with dark veins. 
Fore and middle femora rather obscurely banded with fuscous, base of spines of hind 
tibiae blackish. Vertex somewhat declivous, distinctly wider than long, wider than an 
eye, flattened, not produced in front of eyes. Valve pale, wider than long, angularly 
rounded apically ; plates a little longer in the middle than the valve, the two together 
truncately rounded apically. 



HEMIPTERA 557 

% a little darker than the male. Tergites dark ; pygophor pale yellowish ferruginous, 
ovipositor sheath extending beyond the apex of the tegmina (and of the pygophor) ; last 
sternite trisinuate apically. Clypeus blackish, genae and lora ferruginous. Length 
t l\, % 6-J- mm. 

Hab. Kauai, Kalihiwai, 400 ft., on fern {Microkpia strigosa, not as first stated 
Gleichenia dichotonui) (Oct., Giffard)'. 

Nymph (last instar) : pale brownish yellow, more or less infuscate (varyingly so) 
on the nota, a pale fuscous band across the bend of the head. Ocellar rudiments 
reddish. Basal half of tergites clouded in the middle with pale fuscous, blackish 
sublaterally ; apical half laterally broadly blackish ; beneath mostly pale yellowish : 
sutures blood red. Like the adults, but the head is a little more produced subangularly. 

(2) Nesopliryne microlepiac, sp. nov. 

t closely allied to the last, but much darker. Black ; basal margin of vertex, 
a few irregular markings on pronotum and scutellum, a large central suffusion on frons, 
rostrum (except the apex) etc., brownish testaceous. Tegmina cinereous yellow heavily 
suffused with black, except in a few places, i.e. a few pale spots on clavus, and the extero- 
lateral fourth of the corium (except the suffused cross-veins). Legs brownish yellow 
and black. 

Pygophor black with very short, sparse yellow hairs. Length ^ 5^ mm. 

Hab. Kauai, Kalihiwai, 400 ft, on fern Microlepia strigosa (Oct., Giffard). This 
may be only a dark var. of the preceding, but I do not think so. 

Nesophryne kukanaroa, Kirkaldy. 
Bythoscopus kukanaroa Kirk. huj. op. in. p. 114. 
Hab. Kauai, Halemanu, 4000 ft. 

I cannot refer to the unique type, but I suspect it is allied to the two preceding 
forms, if not identical. 

Nesophryne kaiaiiiamao, Kirkald)'. 

Bythoscopus kaiamaniao Kirkaldy, huj. op. iii. p. 115. 

Hab. Kauai, high plateau. 

The unique type is, in this case too, not available for inspection by me, but 
I believe it is allied to the three preceding. If this is so, then a little group of four 
stoutly built forms in this genus, occur only on Kauai. 

' One nymph was labelled " Kilauea, Hawaii," certainly by mistake. It should have been "Kilauea, 

Kauai." 

72 — 2 



558 FAUNA HAWAIIENSIS 

Nesophrosyne Kirkaldy. 

N^esophrosyne Kirkaldy, 1907, Proc. Haw. Ent. Soc. i. p. 160. 

Typa perkmsi. This genus is closely allied to Nephotettix, but the venation will 
separate it'. Typically there are two discoidals, the median cell (interior discoidal) 
running undivided to the subapical line ; there is only one normal subapical cell, which 
is a little constricted medianly, but in all the species but two, a second subapical cell is 
formed by the forking and quick reunion of the radial vein, this small cell being one- 
third of the length of the other subapical cell, and of course pedicellate at both ends. 
This small subapical is joined at its base by a straight cross (nodal) vein to the 
subcostal vein, sometimes there are other nodals. There are four apical cells and 
a well-defined appendix. The pronotum is very short at the sides. In two species, 
however, insjtlaris and oceanides, the exterior subapical cell has disappeared^ 
These two last may form a subgenus Nesoreias (type insularis). N. haleakala I 
formerly separated subgenerically, on account of the longer head and shorter wings, 
but N. halemanu seems to form too strong a link with the more typical species. 

This genus — Nesophrosyne — is the most difficult of the Hemipterous genera of these 
islands to deal with specifically. It is impossible in some instances, from the material 
before me, to say whether certain forms are species or only local varieties. A much 
more adequate material, a knowledge of the range of variation, of the foodplants, and of 
the nymphs, is necessary before the synonymy can be settled. The variation in some 
forms known to me is quite bewildering. 

(a) Subgenus typicum. 

(l) Nesophrosyne umbratilis, sp. nov. 

^ black ; a longitudinal stripe down the vertex, the face (base excepted), sterna, 
legs, scutellum (except the lateral angles broadly) white. Ocelli red. Tegmina black, 
three pale yellowish elongate spots basally and two white ones apically, on clavus ; four 
(or so) obscurer ones on the apical cells internally, and a broad white exterior band 
from near the base to near the apex, broken only by the dark fuscous first apical vein. 
Tergites black, laterally pale ; sternites basally and laterally whitish, the rest — including 
the valve — black, except the creamy plates which are fringed with fine, short, pale hairs. 
Vertex produced about two-thirds of the length of an eye beyond the eyes, a little 
longer than wide at base. Length 4 mm. 

Hab. Kauai, Kalihiwai, 400 ft. (Oct., Giffard), on fern (probably Microlei>ia 
strigosa). 

A striking species, of which only a single male is yet known. 

' This is figured 1908, P. Haw. E. S. i. textf. 3, on p. 208. 

^ In one species the exterior cell is open exteriorly in one example, the only exception I have seen. 



HEMIPTERA 



559 



(2) Nesophrosyne halemanu, sp. nov. 

Brownish yellow ; vertex with a mediolongitiidinal fuscous line, suffused a little 
anteriorly, and four specks, two anteriorly and two posteriorly, of the same hue. Face 
with a strong dark radiating pattern, antennae pale. Tegmina hyaline, tinged with 
yellow, veins brownish fuscous, more or less suffused, apex of tegmen suffusedly 
brownish fuscous. There are some rather faint brownish fuscous smudgings in the 
subcostal and interior discoidal cells Front and middle femora blackish annulate, hind 
tibiae strongly spotted with black. Vertex rounded anteriorly, forming a curve with 
the eyes, subcircularly depressed medianly, a trifle longer than wide at base and a 
little longer than the pronotum, which is a little longer than the scutellum. 

$ sternites more or less greyish black medianly, valve blackish, plates yellowish. 
Length 4^1: mm. 

Hab. Kauai, Halemanu, 4000 ft. (May, 1S95, Perkins, 503, one J and one 
without abdomen). 

(3) Nesophrosyne silvigena, sp. nov. 

Pale ferruginous ; face black, this encroaching on to the vertex ; grill only visible 
at sides as h, ferruginous. Sterna (.^ entirely), femora (except apically) etc., blackish ; 
hind tibiae striped longitudinally with black, bristles pale ferruginous. Membrane 
clouded with black inwardly, in one example with a pale spot on the clavus. Length 
4i— 5i mm. 

Hab. Kauai, Koholuamano (April, no. 519, Perkins). 

(4) Nesophrosyne pei'kinsi, Kirkaldy. 

Eutettix perkinsi Kirk., 1904, Entom. xxxvii. p. 178; and Haw. For. Agr. i. 183. 

Nesophrosyne per kinsi Kirk., 1907, Pr. Haw. Ent. Soc. i. 160; and 1908 op. cit. 
fig. 3 on p. 208. 

^ creamy, eyes pale olive green. Venation partly pale brownish fuscous, partly 
suffused on the cells, but very variably. Head with eyes as wide as the pronotum. 
obtusely rounded anteriorly ; vertex about as long as its width. Tegmina with the 
exterior subapical cell remote from the subcostal vein ; one nodal vein. 

% like the $, but usually immaculate, sometimes the venation in part a little 
brownish. Two specks on apical margin of last sternite and the ovipositor sheath 

fuscous. Length $ 3-3— 3|. ? 4— 4i mm- 

Nymphs described in Pr. Haw. Ent. Soc. i., 1907, p. i6o. 

Hab. Oahu, coast range around Leahi. and up into Kaimuki, on Sida cordifolia. 



56o FAUNA HAWAIIENSIS 

(5) Nesophrosyne maritima, sp. nov. 

$. Head and scutellum pale yellowish brown ; pronotum and tegmina pale greenish, 
subcostal and apical cells hyaline, second apical faintly smoky ; veins on apical half of 
tegmina and some on the basal half, more or less dark fuscous, some more or less 
suffused, especially at their apices. Beneath and the legs, whitish. Vertex scarcely 
produced in front of eyes, transversely impressed anteriorly. 

?. Head and scutellum pale creamy, sometimes suffused with yellowish ; ovipositor 
pale ferruginous ; seventh sternite sinuate, very minutely notched medianly. Otherwise 
like the male. Length 3^ — 4 mm. 

Nymph : $ pale yellowish ferruginous, irregularly suffused. Vertex with two 
black specks close to the middle ; four specks on the top of the head in an arched row. 
Pronotum with four specks in two sublateral rows and another on each side under the 
eyes. Mesometanotum with about a dozen rather irregular specks. 

Hab. Oahu : Waianae Coast (Perkins, 781, Jan. 2 exx.) : Makua (Perkins coll.). 

(6) Nesophrosyne gouldiae, sp. nov. 

t- Whitish ; claval veins (mostly) suffusedly, and two blotches on the apical 
third, inwardly, of the tegmen, dark brownish ; wings hyaline. Sometimes the rest of 
the hopper is whitish, sometimes the base of the vertex and the disk of the pronotum 
irregularly, dark brownish, with a faint fuscous grill, scarcely apparent laterally. Some- 
times the tegmina are more darkly suffused, but always only on the interior half. 
Vertex obtuse-angled in front of the eyes, scarcely longer than its basal width, some- 
what flat. 

%. Immaculate (practically) whitish ; ovipositor sheath fuscous. Vertex more 
convex. Length J 4^, % 5 mm. 

This has very much the appearance of N. perkinsi, but beyond the difference in 
habitat and foodplant, the vertex is convex and not impressed transversely anteriorly, 
and is also longer, while the size is greater ; in perkinsi the shorter head is distinctly 
flattened and impressed in front of the ocelli. The type is a Tantalus specimen. 

Hab. Oahu, Tantalus, 1400 ft. (Sept., Terry and Perkins) on Gouldia sp. ; 
Palolo, 1400 ft. (Kirkaldy) on Gouldia. 

(7) Nesophrosyne pipttiri, sp. nov. 

Sexes almost monomorphic. Head and underside whitish, ocelli honey yellow. 
Eyes greyish black ; a brownish black speck on the tempora. Pronotum and scutellum 
pale testaceous, the latter whitish posteriorly. Tegmina subhyaline, somewhat milky, 
pattern obscure, a faint St Andrew's cross on the clavi when closed. Subcostal cell 



HEMIPTERA 561 

with two greyish brown suffused spots. Veins (except the colourless radial and claval 
veins) mostly yellowish fuscous, cells slightly suffused. Posterior tibiae with brown 
piliferous tubercles. Lora remote from apical margin of genae. Anal vein of clavus 
not hooked or strongly curved. Vertex obtuse, forming more or less of a curve with 
the eyes, a little produced. 

t. Genital segments pale. V^alve short. Plates pointed apically, Haccid, lateral 
margins reflexed, about three-fourths of the length of the mostly blackish pygophors 
which are bristly apically. Sternites sometimes more or less dark. 

?. Hypopygium somewhat long, laterally rounded, gradually produced at the 
middle which is slightly notched. Pygophor elongate, not quite so long as the black 
ovipositor, apical half with scattered pale yellow bristles. Length 4^ mm. 

Hab. Oahu, Tantalus, 1300 ft. (Feb. — Apr., Kirkaldy, Terry, Giffard and 
Swezey), and Maunawili, 820 ft. (Feb., Giffard) on Mamake [Piptitriis albidus). The 
type is a Tantalus specimen. 

(8) Nesophrosyne ponapona, sp. nov. 

Allied to the last, but darker and with a different pattern. Head etc. pale 
yellowish testaceous, vertex with a thin brown transverse line and two specks of the 
same tint. Ocelli orange. Basal two-thirds of frons yellowish brown with dark brown 
grill ; apical third testaceous. Clypeus and lora apically blackish brown. Pronotum 
pale orange brown tinged with olivaceous, with obscure longitudinal streaks. Pro- 
sternum and sternites black (except laterally, t orange yellow, % yellowish). Scutellum 
whitish, anterolateral and posterior angles widely pale orange brown. Tegmina milky, 
basal two-thirds suffused with pale ochreous brownish, subcostal cell and a number of 
spots on the rest, milky white ; veins yellowish fuscous, apical ones mostly darkly suffused, 
two dark spots in the subcostal cell. Tegmina with two nodal veins, one from the 
middle, one from the base, of the exterior subapical cell, suffused ; subapical cell remote 
from the subcostal vein. Wings smoky ; veins dark fuscous. 

Z- Sternites black, lateral margins orange yellow. 

?. Sternites blackish, lateral margins yellowish, pygophor orange brown with 
yellowish brown bristles ; ovipositor sheath blackish. 

Van Sometimes very dark and suffused. Length 4:j: mm. 

Hab. Same localities and foodplant as N. pipturi, but easily distinguished by the 
pattern of the tegmina. 

(9) Nesophrosyne opalescens, sp. nov. 

Allied to N. pipturi. Vertex, frons, antennae, pronotum and scutellum pale, some- 
what soiled, luteous. Clypeus, lora and genae mauvish opalescent. Ocelli crimson- 
ringed. Tegmina brownish yellow, with fuscous markings much as in A^. pipturi, the 



562 FAUNA HAWAIIENSIS 

whole strongly mauvish opalescent. Wings strongly opalescent, veins dark. Abdomen 
black. Femora dark mauvish opalescent, tibiae pale. 

Z. Pygophor with crimson and white tufts. Length 4 mm. 

Hab. Oahu, Waianae (no elevation stated, April, Perkins). 



(10) Nesophrosytie, sp. ? 

Adult unknown. 

Nymph : t- Vertex as in adult. Pale creamy or whitish, eyes greenish brown ; 
first segment of antennae whitish, rest greenish fuscous. Pronotum with two black 
submedian specks on posterior margin and one under the eye laterally on each side. 
Mesometanotum with two pairs of submedian black spots, one anteriorly and one 
posteriorly, and a curved line laterally on the tegminal pad. Tergites 2 — 7 with two 
black bristly hairs on each side laterally, eighth with about a dozen. Legs pale with 
black, or pale fuscous, bristly hairs. 

Hab. Oahu. 1 took three H off " Ohia lehua " [Nani polymofpha) on Palolo 
Ridge at about 1300 ft. (Sept.), but have not yet connected them with an adult. 

(11) Nesophrosyne tnonticola, sp. nov. 

Vertex whitish, with a blackish brown inverted short-handled trident ; frons pale 
yellowish, narrowly whitish basally with a slightly curved narrow blackish transverse 
line just apical of the yellowish brown ocelli ; then just apical of this, suffused with 
sanguineous ; clypeus, lora and genae mostly blackish. Pronotum yellow, anterior 
margin blackish ferruginous, continued in a narrow line down the middle. Scutellum 
yellow, more or less suffused with ferruginous. Tegmina ferruginous, a large yellowish 
white black-ringed spot on the middle of the clavi closed together ; corium with a few 
small subopaque whitish spots interiorly, and the subcostal in part (apicalwards) hyaline ; 
first and fifth subapical cells hyaline, veins dark ferruginous, sometimes verging on 
black, suffused. 

Hab. Oahu, Kaala, over 2000 ft. (Dec, Perkins). Unfortunately only one ^ of 
this pretty and distinct little hopper. 

(12) jVesop/ij'osyiie koleae, sp. nov. 

J. Vertex and a large round spot on the middle of the closed tegmina, orange 
yellow. Face and legs pale yellow, second segment of posterior tarsi blackish. Base 
of vertex (narrowly and irregularly), pronotum and scutellum (except the usually 
yellowish posterior angle) blackish. Tegmina variegated : obscure subhyaline, 
irregularly but strongly suffused (especially the veins) with dark fuscous, with a bluish 



HEMIPTERA 563 

bloom basally ; apical half of the subcostal cell white with two suffused brownish dark 
brown veins towards the apex ; a large yellowish cinereous spot occupying most of the 
basal half of the clavus ; apical cells mostly smoky ; there is also the orange yellow spot 
mentioned before. Wings dark smoky, veins blackish. Abdomen and sterna black, 
mostly. 

? duller and less variegated, the orange yellow spot sometimes almost absent. 
Pronotum basally and the scutellum olivaceous. Sternites partly pale. Genital 
segments pale yellowish ferruginous, ovipositor black. Length ^ 3f, % i,\ mm. 

Hab. Oahu, Tantalus, 1300 ft. (Jan. and Apr., Giffard) on Koolea {Myrsine 
lessertiand). 

{13) Nesophrosync giffardi, sp. nov. 

,J. Head and scutellum bright pale yellow. Pronotum and tegmina French grey, 
the latter with a broad black longitudinal stripe along the inner margin of the corium 
as far as the apex of the clavus, the stripes on the two tegmina being connected by 
a short transverse stripe. Subcostal cell and membrane whitish or hyaline ; apical and 
subapical veins suffused with black or smoky. Wings smoky, veins dark. Legs 
whitish, second segment of posterior tarsi dark. Sterna blackish. Abdomen black 
and white annulate, somewhat irregularly. 

% more obscure, the tegminal band absent, or faindy indicated. Sternites mostly 
pale. Length ^ 4, $ 4^ mm. 

Hab. Hawaii, Kilauea (Aug., Perkins). Oahu: Tantalus, 1300 ft. (Jan., Feb., 
Apr., Giffard). The type is a Kilauea specimen. 

(14) Nesophrosync ulaula, sp. nov. 

%. Rosy ferruginous, posterior margin of vertex very narrowly pale yellow. Ocelli 
ringed around with fuscous. Clypeus narrowly lined exteriorly with fuscous. Meso- 
notum, tergites, sternites, femora (at least basally) dark fuscous or black. Labium, 
rest of legs etc., pale brownish testaceous (the latter sometimes a little fuscous). 
Tegmina dark yellowish ferruginous ; apical cells (except the first) and appendix pale. 
Wings pale smoky, veins fuscous. Genital segments dorsally fulvous, ventrally pale 
ferruginous, ovipositor dark fuscous. Hypopygium apically sinuate, minutely notched 
medially. Length 5I mm. 

Hab. Oahu, "Honolulu Mts" (Aug., Perkins). 

Var. nigrolineata nov. 

Di^"- s from the type by the slightly more angulate vertex, by a dark fuscous, 
tran? .rse, percurrent stripe apical of the ocelli, and by the rather yellower tegmina. 

Hab. Oahu, Maunaloa, 2000 ft. (Dec, Giffard). 

This may be a good species, but I have seen only a single female, and only two 
females of the typical form. 

F. H. 11. 73 



564 FAUNA HAWAIIENSIS 

(15) Nesophrosyne bobeae, sp. nov. 

Much the appearance of caelicola, but the mark on the vertex is of different 
form, and the grill is more clearly defined. Head and scutellum ivory white, the vertex 
with a subcrescentic mark (the base of the grill), the lateral margins concave to enclose 
the pale ocelli. Frons with a well-defined brown grill, the middle line widening 
apically ; clypeus with lateral margins blackish brown, genae and lora sparsely marked 
with blackish brown. Pronotum pale olive green. Scutellum with pale olive brown 
anterior angles. Tegmina hyaline, tinged with olivaceous, with a slight bronzy gleam ; 
subcostal, brachial and claval veins yellowish testaceous, the rest dark olive brown, 
suffused at the nodal veins and the first two apicals. Clavus a little soiled along the 
commissure, the median cell and inner discoidal cell a little nebulose in the middle. 
Wings a little smoky, veins dark. Tergites black. Sternites and legs pale, base of 
bristles of hind tibiae blackish. Vertex subangular anteriorly. Pronotum aciculate- 
punctate. Exterior subapical cell remote from subcostal vein, angulate exteriorly, one 
nodal vein arising from the middle, suffused. Length ^ 4^ , % 5 mm. 

Hab. Oahu, Tantalus, 1800 ft., on Bobea elatior (Oct., 1906, Perkins). 

(16) Nesophrosyne paludicola, sp. nov. 

%. Blackish ; a yellowish ferruginous grill on a purplish brown frons ; a whitish spot 
on the pronotum near the lateral margins. Scutellum sordid whitish, more or less 
suffused (especially apically) with brownish. Tegmina whitish hyaline, claval veins 
suffused brown, apical angle brown ; inner half of corium brownish except a narrow 
claval margin and one or two apical spots, whitish ; exterior half whitish ; apical 
cells mostly brownish. Abdomen mostly blackish brown, hind tibiae blackish brown, 
with white spines. Pygophor pale with pale hairs, ovipositor sheath blackish. 
Length \\ mm. 

Hab. Molokai, 4000 ft. (June, Perkins, i %). 

(17) Nesophrosyne palustris, sp. nov. 

Vertex, pronotum, scutellum and clavus rose pink ; a faint grill, face more or less 
fuscous apically. Corium testaceous, closely sprinkled with rose pink. Coxae and 
femora (except apex) piceous, rest of legs and abdomen brownish testaceous. Hind 
tibiae at base of bristles piceous. Other characters those of N. ulaula, from which it is 
readily distinguished by the piceous femora and the rosy hue. Length 5 mm. 

Hab. Molokai, Kahanui (Feb., Perkins coll.). One rather immature specimen 
in bad condition. 



HEMIPTERA 565 

{18) Nesoplirosync procel/aris, sp. nov. 

Pale yellowish brown, or yellowish testaceous, the tegmina hyaline brownish 
testaceous, veins rather yellower (except the subcostal), the two claval veins rather 
widely so apically. Exterior subapical cell oval, remote from the subcostal vein, one 
nodal vein. Length 5-|^ mm. 

Hab. Molokai, Kalae (Aug., 1893, "o. 167). 



(19) Nesophrosyne iii})ihicola, sp. nov. 

Head black, the frons with a faint ferruginous radiating pattern. Antennae pale. 
Pronotum basal of the eyes luteous, above this, also the scutellum, ferruginous. 
Tegmina brownish yellow, a large luteous spot on the clavus, a dark fuscous spot of 
variable size exterior to this, and a dark fu.scous line from apex of clavus to apex 
of tegmina, then curving outwardly a little ; two subhyaline specks near the apex of the 
tegmina outwardly. Wings smoky, veins dark. Underside mostly blackish, pleurites 
pale or reddish ferruginou.s. Tibiae and tarsi brownish yellow, hind tarsi partly black, 
hind femora with brownish yellow bristles. 

? pygophor reddish ferruginous (fuscous exteriorly at the base) with paler hairs, 
ovipositor-sheath blackish. Length \\ mm. 

Hab. Lanai, over 2000 — 3000 ft. (2 %% Perkins, Jan.). 

(20) Nesophrosyne tmibricola, sp. nov. 

Differs from the last as follows : 

Frons with better developed pattern. Pronotum yellow. Scutellum sordid yellow. 
A fuscous smudge at the base of the tegmina in the middle, fourth and fifth (and a small 
part of third) apical cells smoky, first apical and apex of median cell hyaline. Wings pale 
yellowish, veins darker. Sternites pale ferruginous. Length \\ mm. 

Hab. Lanai, 2000 ft. (i ?, Perkins, Dec). 

(21) Nesophrosyne luiiu, .sp. nov. 

^. Blackish brown, with a violet tinge. Ocelli pale. Eyes reddish piceous. Fore 
and middle tibiae and tarsi mostly pale, hind tibiae with white bristly hairs ; apex of 
the first and all the third segment of the tarsi, pale. Tegmina blackish brown, opaque 
violet black on clavus, with a small yellow spot ; two hyaline spots near the apex of the 
tegmina. Wings hyaline smoky, veins dark. Abdomen blackish brown. Length 4 mm. 

H.\B. Lanai, 3000 ft. (Jan., Perkins, i $). 

73—2 



566 FAUNA HAWAIIENSIS 

(22) Nesophrosyne imbricola, sp. nov. 

$. Head, pronotum and scutellum pale yellow, with four yellowish ferruginous lines, 
the two inner short (almost spots), the two outer longer and united transversely 
anteriorly. Face pale yellowish ferruginous with a rosy tint, a fuscous more or less 
faint pattern on the frons, lateral margins of clypeus narrowly fuscous (or altogether so). 
Anterolateral angles of scutellum yellowish ferruginous. Tegmina yellowish ferruginous, 
with many faint whitish spots (mostly basally) and four or five small pale yellowish 
opaque ones nearer the middle ; two subhyaline spots, one about the middle on the 
exterior margin and one apical of that ; apical cells more or less decolored ; there are 
also three or four small, dark fuscous spots. Wings smoky hyaline, veins dark. Legs 
yellowish brown, fore and hind femora with the basal half blackish, middle femora 
entirely black except the extreme apex. Abdomen mostly fuscous, lateral margins 
broadly pale, apical margin of sternites pale. Pygophor ferruginous red, with sparse 
ferruginous yellow hairs, ovipositor-sheath fuscous. Length 5^^ mm. 

Hab. Lanai, over 2000 ft. (Jan., Perkins). 

(23) Nesophrosyne oneanea, sp. nov. 

%. Head brownish testaceous, vertex with two basal dots, a transverse line just 
basal of the ocelli and two thin lines at right angles thence to base of vertex at eyes, 
dark fuscous. A well marked grill, the apico-interior part suffused, rest of face suffused, 
the genae a little less so. Pronotum irregularly dark sublaterally on the hind margin. 
Tegmina pale brownish yellow, veins suffusedly brownish fuscous, apical half of radial 
and part of subapical-cell-veins, partly yellowish. Wing veins dark fuscous. Legs pale. 
Sterna partly fuscous, pygophor orange yellow, sheath black. Verte.x subangular. 
Exterior subapical cell subremote, joined to subcostal vein by a suffused broad dark 
spot. Length 5-|- — 5:j: mm. 

Hab. Lanai, 3000 ft. (Jan., Feb., Perkins). 

In the other specimen the vertical mark is feeble, while the face is more blackly 
suffused, the apical margin of the genae clearly pale, but the tegmina are characteristic. 

(24) Nesophrosyne caelicola, sp. nov. 

This has very much the characters of bobeae, but I do not think it is the same. 
The principal difference is that the vertical marking encircles the ocelli as well, while in 
bobeae it is quite clear. A much better series than I have of this and other dubious 
forms, is necessary to settle them definitely. Length 5^ mm. [$). 

Hab. Lanai, 3000 ft. (Jan., Perkins). 



' 



HEMIPTERA 567 

(25) Nesophrosyne nubigena. sp. nov. 

?. Very close to the last, but (except faint traces of the grill), the whole face, the 
fore and middle femora (except apices), hind femora, tibiae and tarsi (except the pale 
third segment), greater part of sterna and abdomen, the pygophor and sheath etc., black. 
Fore and middle tibiae and bristles of hind tibiae, brownish yellow. Some orange 
yellow bristles on pygophor. Length 5I mm. 

Hab. Lanai, 2000 ft. (Perkins). 

This may be the % of caelicola, but I do not think so. 

(26) N^csophrosyne haleakala, sp. nov. 

Head, pronotum and scutellum pale yellow, the vertex with some small inconstant 
fuscous markings, frons with strong, dark fuscous, radiating pattern, apex of frons and 
the clypeus blackish brown, almost immaculate, genae and lorae marked inconstantly 
with fuscous. Tegmina milky, veins dark fuscous, some of the transverse and apical 
ones suffused ; there are fuscous smudgings in most of the cells and the apex of the 
middle apical cell is fuscous. Wings smoky, veins dark. Vertex acute-angled, longer 
than wide at base and much longer than pronotum, subcircularly impressed in the 
middle. Scutellum very small, shorter and much narrower than pronotum. Tegmina 
a little shorter than abdomen. 

? pygophor pale fulvous, ovipositor blackish brown. Length 3^ mm. 

Hab. Maui, Haleakala, 9000 ft. (April, Perkins, 2 %%). 

(27) Nesoplwosync nimbigcna, sp. nov. 

Vertex pale brownish yellow ; a basal dot on each side of the middle and a some- 
what sinuate subapical transverse line fuscous. Face brownish yellow, clearer basal of 
the ferrugineo-fuscous grill which is fairly well defined but somewhat suffused ; apex of 
frons brownish yellow ; rest of face yellowish testaceous, a dark fuscous spot on the 
middle of each lora ; the base of the genae and the entire clypeus, dark fuscous. 
Pronotum olivaceous, anterior half dark fuscous. Scutellum brownish yellow, antero- 
lateral angles (excluding the lateral margins very narrowly) and a submedian spot on 
each side near the anterior margin, dark fuscous. Tegmina cinereo-hyaline, veins 
brownish or yellowish ; a nebulous band across the middle of the clavus and an apical 
blotch on the same, also an irregular blotch on the corium about the middle, part of the 
first and most of the second apical cell, dark fuscous. The exterior subapical cell is 
remote from the exterolateral margin ; there are two nodal veins, one at each end of 
the cell, connecting it with the lateral margins, also another vein basal of the nodals, 



568 FAUNA HAWAIIENSIS 

the two cells thus formed having a transverse, nearly percurrent, line in the middle 
of each. Underside blackish brown. Antennae, labium, legs, incisures of segments etc., 
pale, a blackish spot at base of middle femora [fore femora missing]. 
^ plates pale ferruginous, extreme apex yellowish. 

Hab. Maui, Haleakala, over 5000 ft. (Oct., Perkins, no. 636). 

(28) Nesophrosyne myrsines, sp. nov. 

^. Blackish brown, more or less suffused with bluish ; base of vertex very narrowly, 
antennae and ocelli, pale yellow. Pronotum a little paler irregular locally, posterior 
angle of scutellum ferruginous. Tegmina hyaline, tinged with yellowish brown, on 
the middle third is a broad dark band which starts at the exterior margin and on 
reaching the clavus sends out a narrow fork at an obtuse angle, the space on the clavus 
between this and the main band being pale yellowish white with a green tinge ; at the 
commissure the main band starts back at right angles (thus forming a broad V), and at 
the inner margin of the subcostal cell forks, sending one broad, short, branch across the 
subcostal and another on to the first apical cell. The exterior area forming the contents 
of the "V" is subhyaline, the veins suffused with pale yellowish brown ; the apical part 
of the subcostal cell, between the last mentioned fork, is hyaline. The apical part 
(obliquely) of the tegmen is smoky, with a paler spot ; wings smoky, veins dark fuscous. 
Abdomen black. Genital segments black, with crimson and white hairs. Legs pale 
yellowish brown, femora suffused with dark fuscous and tibiae laterally striped with the 
same. Length \\ mm. 

Hab. Hawaii, Kilauea, on Myi'sine. 

■ Nymph : ^. Vertex and basal three or four tergites, pale yellow, paler beneath and 
on the sides and legs. Eyes and pronotum crimson, the latter anteriorly yellowish. 
The mesometanotum and tegmina crimson, tegmina pale yellowish laterally, the former 
more or less suffused with blackish brown locally. Apical half of abdomen above 
crimson, darkening locally to blackish, eighth segment with particolored hairs (pale 
yellowish basally, darkening apically). Tergites 2 — 7 with a lateral blackish bristly 
hair close to apical margin. Vertex convex, subangularly produced in front of the eyes, 
longer than wide basally. 

(29) Nesophrosyne pluvialis, sp. nov. 

Superficially not unlike a pale form of Thamnotettix clitellaria. 

Black : anterior margin of vertex very narrowly, frons (except apically a little 
obscurely), a line down middle of clypeus, antennae, etc., brownish yellow; posterior 
half of pronotum, a large semicircular spot on each clavus (united in repose) opaque 
creamy ; apex of tegmina smoky, subiridescent ; two large hyaline spots on the apical 



HEMIPTERA 569 

half exteriorly and a few smaller scattered hyaline specks. Wings smoky hyaline, veins 
dark. Legs pale except the posterior tibiae which are greyish black, with whitish 
bristles. Apical margin of tergites and lateral margin of pleurites pale. Vertex 
scarcely produced beyond eyes, not longer than pronotum. 

? pygophor and ovipositor black, the former with sparse whitish hairs. 

Hab. Hawaii, Olaa (Dec, Perkins, i ?). 

(30) Nesophrosyne ehu, sp. nov. 
%. Allied to A^. ponapona. Pale immaculate ferruginous. Tegmina translucently 
spotted, subcostal area and apical cells etc., translucent. Wing veins dark fuscous. 
Ovipositor-sheath black. Length 4 mm. 

Hab. Hawaii, Kilauea (Aug., Perkins), Hilo (Apr., Swezey). The type is a 
Kilauean example. 

(31) Nesophrosyne oreadis, sp. nov. 

Pale ferruginous ; vertex ivory white anteriorly and basally, base of frons very 
narrowly fuscous. Clypeus, sterna partly, fore and middle femora mostly, blackish 
brown, pleurites yellow. Tegmina brownish yellow, or pale ferruginous ; a large pale 
yellow spot on clavus with a fuscous edging basally ; corium spotted with hyaline and 
the nodals are fuscous. Two nodals, one at the apex and one at the middle of the 
remote exterior subapical. 

Hab. Hawaii, Kilauea (Aug., Perkins, no. 656). 

(32) Nesophrosyne montivaga, sp. nov. 

Somewhat like N. pipturi, but the veins are more suffused etc. 

Yellowish (sometimes more testaceous, sometimes tinged with orange or reddish) ; 
a narrow curved transverse line at base of frons, genae, lora, clypeus (and sometimes 
apical half of frons), sterna, abdomen etc. black. Pronotum varying speckled with 
blackish. Tegmina yellowish cinereous, veins mostly blackish brown, varyingly 
suffused ; inside some of the cells is a median nebulosity ; two nodal veins almost 
united. Length t l\ — \< ? 4 mm- 

Hab. Hawaii, Kilauea (Aug., Perkins), Hamakua (Apr., Swezey). A Kilauea 
specimen is the type. Separated at once from N. pipturi, ponapona and montium by 
the clypeus, lora and genae being always blackish. 

kii) Nesophrosyne montium, sp. nov. 
Ngslv ponapona and montivaga ; from the latter it differs by the genae being always 
pale, except the extreme inward part ; from the former by the clypeus and lora being 
always dark piceous, except a short ferruginous line on the clypeus, and by the dark 
suffused tegminal subapicals. Beneath black. 



570 FAUNA HAWAIIENSIS 

Z- Pygophor black. 

%. Pygophor blackish ferruginous, sheath black. Length, ,? 3^, ? 4 mm. 

Hab. Hawaii, Mountain View (March, Swezey). 

(34) Nesophrosyne sihicola, sp. nov. 

Pale olivaceous, paler beneath ; vertex and scutelIum[vbrownish testaceous, a brown 
line on the anterior margin of the former, joining the testaceous ocelli archingly and 
also encircling them, a thinner line joining them straightly. Frons tinged with 
olivaceous and with a faint grill, the rest of the face yellowish brown, the clypeus 
apically more or less fuscate at the sides. Tegminal veins brownish olivaceous, some- 
times fading in places ; in others, especially apically and at the nodes, slightly suffused. 
The clavus is sometimes a little marked interiorly with brownish olivaceous. Wings 
hyaline. Legs testaceous, base of spines on hind tibiae blackish. Pronotum a little 
longer than the head, aciculate punctate. Tegmina with the exterior subapical cell 
remote from the subcostal vein ; one nodal (median) vein, suffused. 

$. Ovipositor sheath blackish. Length, % 5^ mm. 

Hab. Hawaii, Kilauea (August, Perkins), Kona, 4000 ft. (July) ; Hamakua 
(April, Swezey). Probably also from Lanai, over 2000 ft. (Jan., Perkins). The type is 
a specimen from Kilauea. 

(35) Nesophrosyne ignigena, sp. nov. 

$. Bright yellow, with a greenish tinge ; frons (and sometimes vertex), underside, 
legs, etc., yellowish white. Clavus and interior half — irregularly — of corium, opaque 
yellowish (sometimes partly greenish), except a couple of hyaline spots each on median 
and interior subapical cells ; rest of corium hyaline. Apical cells hyaline, second yellow, 
third and fourth a little smoky. Wings smoky basally, veins blackish. Tergites black, 
lateral margins broadly pale yellow. Vertex basally about as wide as an eye, interior 
lateral margins of eyes slightly divergent, vertex rounded apically, very little produced 
beyond eyes, not quite as wide as pronotum. Ovipositor black. First (exterior) sub- 
apical cell minute. Length, % \\ mm. 

Hab. Hawaii, Kilauea (July, Giffard), in the Koa-forest (not necessarily on Koa). 

(36) Nesophrosyne pele, sp. nov. 

Yellowish, tinged with green, vertex with four fuscous specks ; frons with a warm 
brown, well defined, radiating pattern ; rest of face (except a small part of the apical 
margin of the genae), abdomen (except pleurites partly yellowish) etc., black. Antennae 
yellowish. Clavus opaque greenish yellow, veins yellow, two large black spots exteriorly 
and three basal and three interior specks. Corium hyaline, varyingly (usually mostly, 



HEMIPTERA 571 

except the subcostal cell) suffused with blackish brown, veins yellowish, a whitish spot 
basal of the middle. In the subcostal cell there is a spot close to the radial cell, and 
two nodal veins suffused, blackish brown. Apical cells hyaline, veins blacki.sh brown, 
mostly suffused, apical margin suffused. Wings smoky. Exterior subapical cell 
exteriorly obtusetriangular, with two short nodals at the angle (very close together) 
and another at the base of the cell. 

?• Pygophor and last sternite pale, ovipositor sheath black. 

Hab. Hawaii, Kilauea (Aug., Perkins), Olaa (Sept., Perkins) ; probably also 
Kauai (no. 631). 

kol) Nesophrosyne umbrigena, sp. nov. 

Dark piceous ; ocelli, hind margin of vertex narrowly, and antennal peduncle, 
yellowish ; posterior half of scutellum orange brown. Grill merely a yellow lateral line 
on frons, with very obscure transverse lines. Tegmina pale ferruginous ; apical half of 
clavus dark fuscous with a large whitish spot ; subcostal area medially and apically 
subhyaline, with a dark spot at the base, and one or two smaller spots interiorly, a large 
dark fuscous spot around the nodal vein. Apical two-thirds of membrane dark fuscous. 
Sternites black, pleurites partly brownish yellow. Legs as in N. silvicola. . Length 
4f— 5 mm. 

Hab. Hawaii, Kilauea (July— Aug., Perkins). In a damaged example, the 
pronotum is largely whitish, and the dark area of the clavus reduced. 



(38) Nesophrosyne craterigena, sp. nov. 

Orange red, pronotum a little soiled ; apical margin of scutellum irregularly, 
narrowly, blackish. Clavus and interior half of corium pale olive greenish, opaque, the 
rest hyaline ; two dark fuscous spots in median cell, a spot at the apex of the basal 
third of the subcostal cell, one at the nodal vein, and the apex (elongately) of clavus. 
The claval veins yellowish medially. Apical third of membrane smoky, first and second 
apical veins etc. a little suffused. Length, 4^ mm. 

Hab. Hawaii, Kona (March, i ^, Perkins). 

(39) Nesophrosyne arcadiico/a, sp. nov. 

Ferruginous ; the head largely suffused with blackish, except a transverse line near 
the apex of the vertex, the frontal margins of the genae and the genal margins of the 
frons (on the latter indeed, faint transverse lines, the remains of the radiating pattern). 
The hind margin of the pronotum is white, and on the pronotum and scutellum are 

F. H. II. 74 



5/2 FAUNA HAWAIIENSIS 

sparse blackish marks. Tegmina with a broad band across the middle rather obliquely, 
a second nearer the apex, and the apex, blackish. A large whitish spot on the clavus, 
bordered basally with black. Rostrum and legs pale ferruginous, or brownish testaceous ; 
the basal three-fourths of the fore femora, the middle femora, a ring on the hind femora, 
the bases (minutely) of the spines on the hind tibiae etc., black. Sternites black (and 
probably tergites). 

t. Valve very small. Length 4^^ mm. 

Hab. Hawaii, Hilo (April, Swezey). 



(40) NesopJu'osync niieniie, sp. nov. 

$. Yellowish testaceous ; vertex with a more or less broken cross and four specks, 
brownish ; frons with a broken and feeble brownish black radiating pattern. Pronotum 
(tinged with green) and scutellum somewhat soiled. Tegmina cinereous, tinged with 
green, the closed clavi with a large pale greenish white spot, irregularly and very 
narrowly margined with pale fuscous ; corium exteriorly mostly hyaline, veins dark 
fuscous, cross veins somewhat suffused ; two subcontiguous nodal veins and one near the 
base of the interior' subapical ; some of the cells a little nebulose medianly. Sternites 
black, apical segments and pygophor pale greenish white, hairs pale. Head of the 
pipturi type. 

%. The grill fainter ; pronotum more or less variegated with pale fuscous. 
Pygophor brownish yellow, hairs concolorous, ovipositor-sheath ferruginous, narrowly 
margined with blackish brown. Head not so angulate anteriorly as in the ^. Eyes 
pale castaneous, with a transverse paler bar, narrowly bordered with blackish on each 
side, not so distinct in the t as in the %. Length, ^ 4, $ 4^ mm. 

Nymph : pale yellow brown, with the following blackish marks : two subcontiguous 
specks in the middle and four on the apical margin of the vertex ; two spots anteriorly 
and four on the hind margin of the pronotum ; a submedian longitudinal line on the 
mesometanotum and a short transverse line at the base which turns off upwards at 
right angles for a little space, also one or two spots etc. on the pads ; eight sublateral 
and five lateral (basal) spots on the tergites and two submedian specks on the last 
tergite. Each of the 26 lateral and sublateral spots emits a black bristle and there are 
some of the latter also on the last tergite and on the pygophor. Legs pale, tibiae more 
or less striped with black, bristles of hind tibia pale. Antennae more or less fuscous, 
flagellum black. (Hamakua, April, Swezey.) 

Hab. Hawaii, Kilauea (April, Swezey, Kirkaldy). 
' On one tegmen there is only one subapical, the exterior, both the discoidals being undivided. 



HEMIPTERA 573 

Subgen. 2. Nesoreias nov. 

I. Face dark, with two paler spots i insularis. 

\a. Face yellow 2 oaanides. 

(41) Nesophrosyne {Nesoreias) insularis, sp. nov. 

Vertex and pronotum whitish yellow, the former suffused with fuscous, usually 
leaving simply two large subcontiguous pale spots at the base ; pronotum sometimes a 
little suffused. Ocelli pale reddish orange, probably not functional. Scutellum yellowish. 
Frons blackish brown, basally narrowly obscure, two large contiguous pale fulvous spots 
close to this ; genae, clypeus and lorae blackish brown, antennae pale. Underside 
yellowish white. Tegmina hyaline yellowish ; clavus opaque pale greenish white ; 
subapical line and apical veins suffusedly fuscous ; apex of clavus and apical cells smoky. 
Wings smoky, veins dark. Vertex scarcely extending before the eyes. A slender 
species. 

i. Seventh sternite apically black, two small spots medianly, genital segment red 
and black. Length 4 rnm. 

Hab. Hawaii, Kilauea (Aug., Perkins), Hamakua (April, Swezey). The type is 
a Kilauea specimen. 

(42) Nesophrosyne {Nesoreias) oceanides, sp. nov. 

Bright yellow ; pronotum a little suffused with greenish. Tegmina with the clavus 
bright yellow, a spot at the base and a smaller one at the apex, brownish black, rest of 
tegmina brownish black except the apex of the subapical vein, the apical cells, and the 
subcostal cell (except basally) etc., hyaline ; veins on these hyaline parts rather suffused 
dark fuscous. Subcostal vein pale yellow, except basally. Wings black. Legs pale 
yellowish, hind tibiae with the base of the spines etc., blackish. Vertex two-thirds 
wider between the eyes at ocelli than at base, one-third wider there than the length. 
Pronotum slightly longer than the head. Length 4-^ mm. 

Hab. Hawaii, Olaa (Aug., Perkins), Hilo (April, Swezey). The type is an Olaa 
specimen. 

Nymph : Head pale yellowish with some obscure sanguineous lines on vertex ; 
nota sanguineous (except lateral margins of pads) ; rest of body whitish or yellowish 
testaceous, except apical half of tergites which is mostly sanguineous with a blackish 
median line, suffused basally (on apical half). Head has much the shape of the adult's. 

Nesosteles Kirkaldy. 

Nesosteles Kirkaldy, 1906, Bull. Ent. H. S. P. Ent. i. 343. 

This is very close to Balchitlia Kirkaldy, and its synonymy depends on whether 
N. glauca and N. hebe are congeneric ; the latter seems to differ very little from 

74—2 



574 FAUNA HAWAIIENSIS 

Balclutha, but I have not sufficient material of the latter now for effective comparison, 
so that I prefer, for the present, to leave matters as they are. 

Conspectus specieruni hawaiiensium. 

1. Caput pronotumque viridi-testacea maculis parvis ac lineis inclaris, pallidis 

rufo-fuscis. Tegminum venae colore tegminum pallidiores (2). 

\a. Caput pronotumque viridescentia vel aureo-brunnea. Tegminum venae 
virides (3)- 

2. Vertex ante oculos vix productus. Tegminum venae late pallidae. Longitudo 

3| — 4 mm I hebe Kirkaldy. 

2a. Vertex leviter rotundatim productus. Tegminum venae inclarae, pallidae. 

Longitudo 2 mm 2 volcanicola nov. 

3. Forma elongata, angusta, satis parallela ; caput pronotumque aureo-brunnea; 

tegmina viridescentia ; vertex subtriangulariter productus. Longitudo 

4 mm 3 plutonis sp. nov. 

3<z. Brevior ? latior, tegminibus ad latera magis rotundis. Vertex ante oculos 

rotundatim paullo productus. Longitudo 2\ mm 4 peregrina sp. nov. 

(i) Nesos teles hebe Kirkaldy. 

Nesosteles hebe, Kirkaldy, 1906, 1. c. 

H.\B. Well distributed over the Islands. I have seen it from Kauai, Kealia 
(Oct., Giffard) ; Oahu, all over, frequently to light (Kirkaldy, Perkins, Giffard, Swezey, 
Terry). Hawaii, Kilauea (Perkins). 

This seems to be hebe from Viti, though all the examples seen are much less 
reddish on the head, being more yellowish green. It may be convenient to distinguish 
it varietally as Aospes, 

(2) Nesosteles volcanicola, sp. nov. 

Closely allied to the above, but much smaller, the verte.x more produced before the 
eyes, and the tegminal veins more obscure. The face also has a somewhat incomplete 
grill. Length 2 mm. 

Hab. Hawaii, Kilauea (Perkins, 1906). 

I have only seen a few carded examples from Kilauea, but it is probably not 
endemic. 

(3) Nesosteles plutonis, sp. nov. 

Head, pronotum, legs etc. golden brown ; tegmina yellowish green, membrane 
hyaline, with yellowish green veins. Abdomen mostly black. Elongate and slender, 
head distincdy wider medianly than at the eyes, a trifle more angular in the $ than in 
the $. Length 4 mm. 

Hab. Hawaii, Kilauea. 

• I have seen only a carded pair in Dr Perkins' collection. It is probably endemic. 



HEMIPTERA 575 

(4) Nesosteles peregrina, sp. nov. 

Yellowish green or green. Abdomen below partly faint fuscous. Membrane 
hyaline, veins yellowish green. Wings hyaline, veins faint fuscous. Vertex flatter 
than in the other three species, distinctly wider in the middle than at the eyes. 
Length 2^ mm. 

A Kilauea specimen is the type. 

Hab. Oahu, Honolulu, comes to light (Kirkaldy) ; Hawaii, Kilauea (Perkins and 
Giffard). Probably not endemic. 



Macrosteles Fieber. 

Macrosteles Fieber, 1866, Verh. zool. bot. Ges. Wien xvi. 

I know only a single Hawaiian species, which appears to be endemic. 



(i) Macrosteles kilaueae, sp. nov. 

Bright yellow, legs etc. paler. Eyes greenish grey. Apical cells of tegmina 
hyaline, with yellow veins. Wings hyaline, veins white, sometimes partly fuscous. 
There is no appreciable sculpture, except microscopic. Vertex wider between the eyes 
than its length, and wider than the eyes together, lateral margins slightly divergent ; 
rounded obtusely anteriorly, somewhat convex, roundly obtuse in profile ; frons some- 
what convex. Pronotum wider than the head and a trifle longer, lateral margins short, 
not keeled, hind margin slightly obtuse-angulately emarginate. Tegminal venation not 
very unlike that of M. punctifr-ons, but the first apical vein is sublongitudinal, not sub- 
transverse, and the fourth vein is also longer and more longitudinal. 

%. Sheath blackish brown. Length 4^ mm. 

Hab. Hawaii, Kilauea. I have seen only one carded ?, in Dr Perkins' collection. 



Phrynomorphus Curtis. 

Phrytiovwiphiis Curtis, 1833, Ent. Mag. i. p. 194. 

Athysanus Burmeister, 1838, Gen. Ins. i., Jasstis. 

Athysanus subg. Athysanus and Conosanus, Osborn and Ball, 1902, Ohio Nat. u. 
232 (not subg. Commellus which is part oi Deltocephalus, sens. lat.). 

The synonymy of this and the allied genera is complicated and as the Hawaiian 
representative is an introduced form, it is unnecessary to dilate here on the generic 
nomenclature. 



576 FAUNA HAWAIIENSIS 

(i) P hrynomorphus hospes, Kirkaldy. 

Deltocephalus hospes Kirkaldy, 1904, Entom. xxxvii. 177. 

Phrynomorphus hospes Kirkaldy, 1907, Bull. H. S. P. A. Ent. iii. 60, PI. i. 

figs. 13—17- 
The last nymphal instar is described and figured in the work last cited (p. 13). 
This introduction from Australia and Fiji feeds here on Cynodon dactylon. 

Tribe Eurvmelini. 

There do not appear to be any of this tribe in the Hawaiian Islands, the two 
species collected by Dr Perkins and referred by me to Bythoscopus, belonging really to 
the genus Nesophryne. 

Stal described two species, B. peregrinans (referred to incorrectly on p. 1 1 5 of 
vol. III. as B . peregrimis) and B. vidiius from Oahu, but erroneously\ 

B . peregritians v^a.s described from Tahiti, Oahu, California and Rio de Janeiro. 
It has been since recognized as an Agallia (sens, lat.) by Berg and recorded from the 
Argentine Republic ; recently Osborn and Ball (now the highest North American 
authorities on Homoptera), believe that a species from California answers to Stal's 
description ; they place it close to A. sanguinolenijis, which is an Aceratagallia. It is 
perhaps a little doubtful however whether the same species actually has this distribution 
from Vancouver to the Argentine. It is well known that the localities recorded in the 
" Eugenies Resa" are often quite inaccurate and as B. viduzis is recorded from Tahiti 
as well as Oahu, and as nothing like either peregrinans or viduus has ever been recorded 
from the Hawaiian Islands, both may safely be expunged from the Hawaiian lists. 

Fam. ASIRACIDAE. 

The Hawaiian Asiracidae are, as regards the endemic, arboreal forms, characterized 
by the spur of the hind tibiae being solid, subcultrate, with rather large teeth ; a 
condition known, in outside faunas, only in Proterosydne arboricola Kirkaldy, from 
Queensland. 

In this family, the male genitalia are of the highest specific, and even generic, 
importance ; hence in these islands, as in other countries, females are often very difficult 
to determine, unless taken with the males. The colour and pattern of the nymphs are 
generally of specific value. 

The genera in Hawaii may be separated as follows, but I must remark that 
Nesosydne is probably heterogeneous. A considerable proportion of the species, how- 
ever, is known only in the brachypterous state, if indeed these latter forms are ever 
macropterous, and I therefore prefer waiting a little before founding the probably 
necessary new genera. 

' Cf. Kirkaldy, 1907, A. S. E. Belg. li. 



2. 

2a. 



HEMIPTERA ^yy 

Conspectus generum. 

Calcar mobile tibiarum posticarum laminatum, bifidum, dentibus invalidis 
instructum. Formae graminivorae [Tribus Criomorphini]' (2). 

Calcar solidum, subcultratum, dentibus fortioribus, paucioribus instructum. 
Formae arborivorae vel filicivorae, Nesosydne ipovweicola, forma herbivora, 
excepta. [Tribus Proterosydnini] (4). 

Antennae fere cylindricae (3). 

Antennae depressae ac dilatatae 2 Perkinsiella Kirkaldy. 

3. Carinae frontis fere ad basim frontis in unum conjunctae 3 Kelisia Fieber. 

3a. Carinae frontis fere medio oculorum conjunctae i Peregrinus Kirkaldy. 

4. Caput ordinarium ( 5 ) . 

4a. Caput perelongatum, abdomine thoraceque simul sumptis longius ... 10 Dictyophorodelphax Swezey. 

5. Frons carinis duabus, interdum obsolescentibus, instructa (6). 

Sa. Frons carinis duabus, in unum fere ad basim conjunctis, instructa (9). 

6. Tegmina baud reticulata (7). 

da. Tegmina brevia, venis baud faciliter distinguendis, subreticulatis 6 Nesorestias Kirkaldy- 

7. Frons elongata, carinis duabus perdistinctis, subparallelis, instructa 5 Nesopleias gen. nov. 

la. Frons minus elongata, carinis lateralibus baud semper faciliter distinguendis ; 

si distinctis, tunc rotundatis (8). 

8. Frons immaculata 4 Aloha (typ.) Kirkaldy. 

2>a. Frons pallida maculata 4a (subg.) Leialoha nov. 

9. Formae pergraciles 9 Nesodryas Kirkaldy. 

9(7. Formae robustae (i°)- 

10. Frons baud maculata 7 Nesosydne Kirkaldy. 

loa. Frons pallide maculata 8 Nesothoe Kirkaldy. 



Peregrinus Kirkaldy. 
Peregrinus Kirkaldy, 1904, Entom. xxxvii. p. 175. 

( I ) Peregmnus viaidis, Ashmead. 

Delphax maidis Ashmead, 1890, Psyche v. p. 323, figs. 

D. psylloides Lethierry, 1896, Ind. Mus. Notes in. p. 105, fig. 

Peregrinus maidis Kirkaldy, 1906, Bull. H. S. P. Ent. i. p. 407; and 1907 op. cit. 
III. p. 132, PI. 12, figs. 7 — 8 ; PI. 10, fig. 14. 

Pundaluoya simpiicia Distant, 1906, Faun. Ind. Rh. in. 468, fig. 255. 

Liburnia psylloides Distant, op. cit. 484. 

Hab. All the islands on Zea mays, the adults also on Saccharum officinarum, 
Sorghum vulgare, etc. One specimen was taken by Dr Perkins in 1892. This intro- 
duced species is found also on Bromus unioloides and Cynodon dadylon in Queensland 
(Tryon), and has been collected as well in Fiji, New South Wales, Java. Ceylon, Florida, 
Texas, and the Antilles. 

' Asiraca atque genera proxima tribum ' Asiracini ' constituunt. Habent haec calcar subuliforme. 



578 FAUNA HAWAIIENSIS 

Perkinsiella Kirkaldy. 
Perkinsiella Kirkaldy, 1903, Entom. xxxvi. p. 179. 

(i) Perkinsiella saccharicida Kirkaldy. 

Perkinsiella saccharicida Kirkaldy, 1903, 1. c. ; 1906, Bull. H. S. P. Ent. i. p. 405, 
Pis. 26 & 27, figs. 1—5 ; 1907, op. cit. III. p. 137, PI. VIII. figs. 5—8 ; PI. xii. 
figs. 2—13. 

P. saccharida Kirkaldy, 1906, op. cit. i. p. 406. 

Hab. Wherever Saccharum officinaruni is grown in the islands ; now greatly 
reduced by various parasites. It has been found in Eastern Australia and Java. 

The metamorphoses are partly described and figured in the papers cited above. 

Kelisia Fieber. 

Kelisia Fieber, 1866, Verh. zool. bot. Ges. Wien xvr. p. 519. 

Three species, apparently not endemic, have been found in the islands. I cannot, 
however, connect them with any previously described. 

(1) Kelisia sporobolicola, sp. nov. 

t ?. Yellowish testaceous, keels paler. Face and antennae brownish testaceous, 
the frons speckled with yellowish testaceous. Tegmina tinged with cinereous, veins 
whitish testaceous, rather thickly granulated with blackish brown. Abdomen above 
spotted laterally with blackish brown. The tegmina reach to the middle of the 
abdomen, and there are no clear apical cells. 

J. Pygophor pale, partly more or less infuscate ; with black genital styles, 
oedeagus etc. ; genital styles long and narrow, divergent, apical third acuminate. 

$. Pygophor concolorous with the underside. 

The type is an Oahuan specimen. 

Hab. Oahu, Honolulu coast (Oct. i ^, 2 $? Perkins ; March, Swezey and 
Kirkaldy); Hawaii, Puako (May, Swezey). 

(2) Kelisia szvezeyi, sp. nov. 

$. Superficially not unlike the last described, and with not very dissimilar genital 
organs, but the pronotal keels are less divergent posteriorly, the tegmina are longer and 
less granulate, and the frons is different. 

Head, pronotum and scutellum pale fuscous, frons and clypeus bordered with black 
inside the pale lateral keels ; rest of body yellowish testaceous, the abdomen sparsely 



HEMIPTERA 



579 



marked with black laterally and on the pygophor. Tegmina rather feebly granulate 
with brown, a black speck near the apex of the clavus on the commissure. Head 
dorsally very slightly produced beyond the eyes, frontal keels blunt. Tegmina reaching 
to about three-fourths of the length of the abdomen, with somewhat clear apical cells. 
Pygophor testaceous, a longitudinal line dorsolaterally, the ventral wall of the anal tube, 
the genital styles, etc., black ; the last very similar in form to those of K. sporoboli. 
Length 2\ mm. 

Hab. Oahu, Kalihi (March, Swezey). Only a single specimen, kindly lent to me 
by Mr Swezey ; it is the only one he has seen. 

(3) Kelisia paludunt, sp. nov. 

Z. Brownish yellow, abdomen often largely infuscate ; a dark fuscous spot at the 
apex of the commissural area. Tibial spur with 18 minute teeth. Tegmina very feebly 
granulate. 

The t pygophor is more or less dusky ; genital styles long and narrow, slightly 
curved, shortly bifid apically. 

%. Immaculate yellow. 

Hab. Oahu, Waikiki (Jan. and March, Swezey and Kirkaldy), on Herpestis 
ni07inieria and Junciis. These were closely intermingled, and eggs not having been 
found, the actual foodplant is not certain. 

Aloha Kirkaldy. 

Aloha Kirkaldy, 1904, Entom. xxxvii. p. 177. 

Corresponds to Criomorphus in the Criomorphini. The median keels of the frons 
are roundedly arched outwardly, but are sometimes obsolescent in A. myoporicola, which 
probably does not belong to this genus. Type A. ipomoeae Kirkaldy. 

Conspectus speciemim hawaiiensiuni. 

1. Frons nigrofusca, parte apicali quarta saltern, cum sex saltern maculis 

utrimque inter carinas submedianas atque margines laterales, pallida, 
maculis quibusdam basin versus minoribus. Antennarum segmentum 
primum nigrescens [subgen. Leialoha nov., typo naniicola Kirkaldy (2). 

\a. Frons concolor, interdum maculis subobsoletis pallidis ornata. Antennarum 
segmentum primum nigrescens apud has, pallidum apud illas species... (5). 

\b. Frons nitida, nigra, immaculata 7 myoporicola nov. 

2. Tegmina vitrea, notis fuscis paucis ornata (3)- 

2(7. Tegmina lactea basi ; apice pallida, fusco-notata ; fascia lata, fusco-fuliginea 

trans medium ornata. Tegminum pili albi i pacifica nov. 

3. Tegmina pallida testaceo et inconspicue granulata, pilis albis i naniicola nov. 

3rt. Tegmina rude albo-granulata, pilis fuscis 2 oceanides nov. 

F. H. II. 75 



58o FAUNA HAWAIIENSIS 

5. Tegmina maculata 4 lehuae nov. 

5a. Tegmina baud maculata, commissura apice excepta (6). 

6. Forma plus minus rubra 5 ohiae nov. 

6<7. Forma haud rubra " 6 ipomoeae Kirkaldy. 

(i) Aloha naniicola, sp. nov. 

Vertex and pronotum testaceous, the latter fuscous laterally with whitish specks. 
Frons darkish fuscous, apical third and some specks, yellowish white ; genae yellowish 
white, extreme base fuscous. Antennae dark fuscous. Clypeus fuscous with pale keels, 
Tegmina hyaline milky with an irregular band along the middle of the membrane, apex 
of clavus, etc., fuscous. Femora pallid, with a subapical fuscous ring, the apex sub- 
sanguineous. Tibiae testaceous, anterior and intermediate pair with two fuscous rings 
on each ; posterior tibiae with one near the base. Length 4 — \\ mm. 

Hab. Oahu, Waianae, 3000ft. (Feb., Perkins); Tantalus, 1300ft. (Jan., April, 
Giffard, Kirkaldy, Swezey). Hawaii, Kilauea (Perkins, Kirkaldy). On Ohia lehua 
{Nani polymorpha, also known generically as Metrosideros). The type is a Kilauea 
example. 

(2) Aloha oceanides, sp. nov. 

Vertex, frons, genae and nota dark brown ; all the keels, the basal specks on frons 
and the specks on the pronotum, yellowish brown ; apical fourth of frons, apical third of 
genae, apical specks on frons, and some at base of genae whitish ; clypeus dark fuscous, 
basal half of middle keel pallid. Antennae dark fuscous or blackish (second segment in 
immature examples paler dorsally). Ventral part of pronotum and the sterna dark 
fuscous. Tegmina milky hyaline, veins (more or less suffusedly) dark brown, strongly 
and closely studded with white granules which emit white hairs ; marginal vein yellowish 
brown. Tegmina suffused with dark brown as follows : extreme base, a short irregular 
transverse blotch extending from marginal vein to median just apical of the junction of 
the latter with the brachial (basally it is fairly straight, apically it extends along the 
median and subcostal veins about as far again), a large rhomboid spot on the subcostal 
cell, at about three-quarters of its length, which extends obliquely on to the median at 
its forking. The apical cells have a large irregular T-shaped mark, and the veins are 
all suffused, at least apically. Anal and axillary suffused just basal of their union. 
Wing veins brownish black. Legs yellowish brown, femora and tibiae annulate with 
dark fuscous. Tergites mostly black ; pleurites yellowish brown and dark fuscous 
chequered. Length 5^ — 5^ mm. 

Hab. Kauai, 4000 ft. (July — Aug., Perkins, three examples, also one in bad 
condition). 

In immature examples, the vertex and the disk of the pronotum, etc., may be 
pallid. 



HEMIPTERA 581 

(3) Aloha pacifica, sp. nov. 

%. Similar to A. oceanides, but proportionately slightly more robust. The whole 
of the tegmina basal of the apical cells is dark smoky except the basal fourth of that 
space, the subcostal cell in part, the apex of the clavus, and the tegminal granules, the 
latter supporting white hairs. Apical keels of the vertex and the middle keels of the 
pronotum and scutellum subsanguineous. Length 5I mm. 

This is a quite distinct species, but the male and definite localities are desiderata. 

Hab. "Kauai? Molokai ? " (i ?, Perkins). 

(4) Aloha lehuie, sp. nov. 

Head and pronotum pale ochraceous, the latter a little fuscate partly, scutellum 
dark ferruginous. Tegmina hyaline, veins pale, granules pale brown ; a dark fuscous 
band across the tegmina close to the base, a short fuscous line at apex of claval cell and 
a fuscous smudge along the middle of the membrane. Wings pale smoky, veins 
blackish brown. Length 4^ mm. 

Hab. Oahu, Tantalus ; on Ohia lehua {Naiti polymorpha). 

(5) Aloha ohiae, sp. nov. 

Head pale yellow ; antennae, legs, scutellum, underside, red (sometimes paler). 
Tegmina hyaline, tinged with yellowish red, veins and granules red. Length 5 mm. 

The type is a Waialua specimen. 

Hab. Oahu, Waialua, Palolo, Pacific Heights, Tantalus. Hawaii, Kona. 
(Perkins, Kirkaldy, Giffard, Terry, Swezey) on Nani polymorpha. 

(6) Aloha ipomocae Kirkaldy. 
Aloha ipomocae Kirkaldy, 1904, Entom. xxxvii. p. 177; and 1908, P. H. E. S. 1. 

PI. 4. fig. 9. 
This species is usually micropterous, but macropterous forms occur. 

Hab. Oahu, Tantalus, etc., up to the summit on Ipomoea pes-caprae, batatas, 
bonanox, tuberculata, insularis, etc. (Perkins, Kirkaldy, Terry, Swezey, Giffard) ; 
Makiki, beneath Hilo grass (Nov., Swezey). Hawaii, Paauhau (March, Swezey). 

(7) Aloha f myoporicola, sp. nov. 

t. Vertex, frons, pronotum and scutellum shining polished black or blackish 
brown ; keels of vertex and of extreme base of frons, apical margin (narrowly) of frons. 
the clypeus, genae (except a slight line), posterior angle, very minutely, of scutellum, 
legs (except femora, a little fuscous locally, and tibiae more or less obscurely annulate), 

75— 2 



582 FAUNA HAWAIIENSIS 

pale yellowish. Antennae yellowish. Tegmina subhyaline pale yellow, veins yellowish 
white, sparsely granulate with brown ; middle two-thirds of tegmen from subcosta to 
suture, smoky. Tergites yellowish, sutures blackish, lateral margins variegated with 
black, sternites mostly blackish. Labium reaching to hind coxae ; second segment of 
antennae scarcely twice as long as the first, the second being about as long as the 
median width of the frons. Tegmina micropterous, extending about as far as the apex 
of the abdomen, with 5 — 6 apical cells. Spur with eight spines. 

?. Usually much paler than the male, the vertex, pronotum, and scutellum being 
more or less pallidly suffused, but sometimes these approach the coloration of the male. 
Tergites dark with yellowish markings apically. Length 4 mm. 

Hab. Hawaii, Kilauea, 4000 ft. (July — Sept., nos. 578, 656, 686 & 691, Perkins, 
July, Giffard, Dec, April — May, Kirkaldy) on Myoporum sandwicense ; Kona, 4000 — 
5000 ft. (June — July, Perkins). 

The type is a Kilauean example. 

Nesopleias, gen. nov. 

This genus has very much the appearance of Nesorestias, but the tegmina are not 
reticulate, and the frons has two entire keels. From Aloha, etc. it differs by the form 
of the tegmina, and by the elongate frons, which has two fine, very distinct, subparallel 
keels, which enclose a sublinear space throughout, meeting just basal of the truncate 
apical margin. The genal carina meets the frontal keels acutely. Pronotum with three 
straight keels, the lateral ones meeting archedly in front ; all meeting the hind margin 
straightly. Scutellum tricarinate. 

Conspectus specierum hawaiiensiuni. 

I. Tegmina cellulis apicalibus destituta ; margine postica densata i ntmbata nov. 

I a. Tegmina cellulis apicalibus instructa ; margine postica ordinaria 2 dubautiae nov. 

(i) Nesopleias ttinibata, sp. nov. 

%. Pale brownish yellow, paler beneath, the tegmina with a fuscous band just 
exterior to the clavus and crossing it at the apex, where it darkens and thickens ; basal 
of this band the colour is paler. Tergites blackish brown laterally. Tegmina scarcely 
reaching beyond the middle of the hopper, roundedly arched externally, rounded 
apically. Subcostal cell and apical margin strongly thickened, especially the latter, 
which is not venose. Radial and brachial veins forked obscurely immediately basal of 
the apical margin. Length 3 mm. 

Hab. Oahu, Tantalus, 1500 ft. (Feb., 1907, Perkins). I have seen only a single 
female of this curious form. 



HEMIPTERA 583 

(2) Nesopleias f dubautiae, sp. nov. 

Highly variable in colour, in some examples almost immaculate pale brownish 
yellow ; in others, the tegmina and abdomen above are heavily blotched with blackish 
brown. These colour variations are not sexual. Spur with nine spines. The pygophor 
has a minute spine on the lip. The ^ styles are contiguous to the oedeagus at their 
laterobasal angle and are produced ventralwards in a triangular lobe ; using the inner 
side of this as a base, they are acuminately and obliquely triangular and are very 
different to those of any other Hawaiian species. Length 3 mm. 

Hab. Oahu, Tantalus (1907, Perkins) ; Konahuanui, 2000 ft. (May, Dec, Swezey) 
on Naenae {Dubautia pianiaginea). 

Nesorestias Kirkaldy. 
Nesorestias Kirkaldy, 1908, P. H. E. S., i. p. 205. 

(i) Nesorestias Jilicicola Kirkaldy. 
Nesorestias filicicola Kirkaldy, 1908, 1. c. 
Hab. Oahu, Tantalus, 2000 ft. (Kirkaldy, Swezey, Terry) on ferns. 

Nesosvdne Kirkaldy. 

Nesosydne Kirkaldy, 1907, op. cit. 161. 

This genus corresponds more or less with Delphacodes Fieber in the section with 
non-compressed spur. The type is N. koae, but the genus as now comprehended is 
doubtless heterogeneous. 

(i) Nesosydne koae Kirkaldy. 

Nesosydne koae Kirkaldy, 1907, 1. c. ; and 1908, op. cit., PI. iv. fig. 2. 

^ %. Pale green, usually yellowing after death. Tegmina hyaline, veins pale 
greenish, with fuscous granules. Sutures of tergites narrowly fuscous. 

^. Base of tergites and the lateral margins greenish, rest blackish. Genital styles 
narrow, elongate, slightly truncately bifid at apex. 

%. Ovipositor dark fuscous. Length ^ 3!, ? 4 mm. 

Hab. Oahu, Tantalus, 1 200— 1 500 ft., on the young leaves of Acacia koa (Perkins, 
Kirkaldy, Giffard, Terry, Swezey). 

Nymphs pale green. 



584 FAUNA HAWAIIENSIS 

var. rubescens Kirkaldy, 1907, op. cit. p. 161. 

Similar to the type and found with it, but larger and with a rosy brownish tinge. 
Length ? 4. ¥ 5 ^m. 

Nymphs green tinged with roseate. 

I have examined some numbers of this sometimes common hopper, but have never 
found a brachypterous form. 

(2) Nesosydne pipttiri Kirlcaldy. 
Nesosydne pipturi Kirkaldy, 1908, op. cit. p. 202, PI. iv. fig. 3. 

<?. Head, pronotum and scutellum fuscous or blackish, with pale keels, the dorsal 
one broadly so and rather suffused. Antennae, clypeus, sterna and legs pale ; femora 
partly fuscous. Tegmina whitish hyaline, a fuscous smudge across the middle third, 
accentuated by the radial and brachial veins being blackish medially and the ape.x of the 
clavus also blackish ; sometimes the darkening is continued more or less to the apex of 
the clavus. Abdomen pale sanguineous, with a few scattered blackish marks. Head 
dorsally scarcely reaching in front of the eyes, apical margin smoothly rounded ; frons 
very smooth, keels almost obsolescent, except as pale lines. Second segment of 
antennae |^ths longer than the first. Labium reaching to the base of the hind trochanters. 
Lateral margins of pronotal keels curving outwards, but reaching hind margin. Tegmina 
rounded apically, not extending as far as apex of abdomen ; with two or three apical 
cells. Spur with 7 — ^8 spines, first segment of hind tarsi much longer than the 
other two. 

%. Pale yellowish, intracarinal areas of vertex and frons rather irregularly fuscous, 
not so deeply or entirely as in the Z- Tegmina hyaline, veins pale yellow, a short black 
line on the commissure. Tergites sometimes lightly stained with fuscous. Length ^ i|-, 
% 2\ mm. 

Easily distinguished by the smooth, wide, somewhat polished frons, with scarcely 
raised keels and by the green nymphs. 

t. Genital styles broad, constricted medially, flatly bifid apically. 

Nymphs pale green, paler beneath. Labium reaching to middle of the hind 
trochanters. Spur with seven spines. 

Hab. Oahu, N.W. Koolau range (825, 786, Perkins, 3 ti)> Tantalus T500 — 
2000 ft. (Feb., May, Aug., Oct., Perkins, Giffard, Kirkaldy, Swezey, Terry), Kona- 
huanui ridge (Feb., Swezey, Jan., Kirkaldy), on Mamake {Piptunis albidtis); Molokai, 
Pelekunu (Oct., Perkins, 3 ??). 

(3) Nesosydne halia Kirkaldy. 
Nesosydne halia Kirkaldy, 1908, op. cit. PL 4, fig. 8. 

$. Pale luteous ; frons between the keels, pronotum and scutellum (usually except 
the keels), dark fuscous. Tegmina cinereohyaline, entire margin pale luteous ; with a 



HEMIPTERA 585 

tiny dark fuscous speck near the apex of the commissure, or also with an obscure obHque 
stripe. Abdomen above sometimes dark apicolaterally ; beneath with dark lateral 
wedges on the three middle segments. Frons elongate, broader at the base than at the 
apex, keels broad and rather flat. Antennae extending to about the middle of the 
clypeus, second segment a little longer than the first. Pronotal keels reaching the hind 
margin, then curving around slightly. Hind tibiae longer than tarsi, first segment of 
the latter one-third longer than the second and third together. Spur with eight teeth. 
Tegmina with three apical cells not nearly attaining the apex of the abdomen. Genital 
styles broad at the base, sinuate, apically acuminate, but truncate at the apex, the inner 
angle acute. The oedeagus is horned on each side basally. 

%. Similar to the ^ but larger ; luteous, the keels paler. Abdomen above mottled 
with fuscous laterally on the apical two-fifths, also similarly at the base. Beneath as in 
the I?. Tegmina also with a dark fuscous blotch at the apex of the commissural cell. 
Length t I, % ^^ mm. 

Hab. Oahu, Tantalus, 1300 ft. {Perkins, Kirkaldy, Giffard), Pacific Heights ridge 
(May, Swezey). 

van %. A dark fuscous oblique stripe on the tegmen, varying to pale. Head, 
pronotum and scutellum (except keels) dark fuscous. 

(4) Nesosydne pele, sp. nov. 

Of the general appearance of N. koae, but larger and differently coloured. 

Head and pronotum brownish testaceous, or pale olivaceous, the clypeus between the 
keels (and sometimes the vertex and frons between the keels), genae and pronotum partly 
anteriorly, dark brown or blackish. Antennae testaceous, extreme apex of first segment 
dark. Scutellum blackish, or piceous. Tegmina hyaline ; apex of anal cell of clavus dark 
brown ; veins brown, rather feebly granulate, black-haired. Legs sordid testaceous, 
more or less suffused. Abdomen dorsally and the body beneath mostly blackish. 
Pronotal keels divergent, not reaching the hind margin. Spur with about nine spines. 
Length 5 mm. 

Hab. Hawaii, Kilauea (Perkins, 1906). 

(5) Nesosydne nmbratica, sp. nov. 

Pale, rather sordid, greenish yellow ; head between the keels fuscous. Tegmina 
with apex of clavus and of subcostal cell dark fuscous. Fore tibiae apically fuscous. 
Labium reaching to the apex of the hind coxae. Second segment of the antennae about 
one-fifth longer than the first. Tibial spur with 9—10 teeth. Tegmina with five apical 
cells, clavus not completely sutured off; apex of subcostal very narrow, truncate, 
continuation of the subapical transverse vein. Tegmina not reaching to the apex of 
abdomen. Length 4| mm. 

Hab. Hawaii. 



586 FAUNA HAWAIIENSIS 

(6) Nesosydne gouldiae, sp. nov. 

Brownish white ; frons and ciypeus between the keels, dark fuscous ; pronotum 
and scutellum between the keels, genae, etc., more or less fuscous. Tegmina yellowish 
brown, hyaline, veins more or less fuscous, the apical ones yellowish brown, apex of 
clavus and of subcostal dark fuscous. Legs and sterna pale yellowish. Tergites more 
or less variegated with dark fuscous. Second segment of antennae one-third longer 
than the first. Tegmina with the clavus not completely sutured off; three apical cells, 
subcostal cell apically acute. Spur with lo — 1 1 spines. Tegmina not reaching to the 
apex of the abdomen. Length 4 — 5 mm. 

The nymphs of the last instar have the same general colour as the adult, but the 
tergites are fuscous except a narrow longitudinal row of spots down the middle (making 
a broken line), and two sublateral longitudinal brownish yellow stripes on two or three 
segments. There are also some pale specks, etc. Second antennal segment about 
twice as long as the first, fore and middle tarsi unjointed, hind tarsi trisegmentate, the 
spur with 7 — 8 spines. 

Hab. Oahu, Honolulu Mts. (Sept., Perkins, nos. 793 & 805, Nov. — Dec, Perkins) 
and Konahuanui (no. 867, March, Perkins) ; also Tantalus (Perkins, Terry) ; Palolo, 
1800 ft. (Terry), on Manono {GoJi/dia sp.). 

(7) Nesosydne nephrolepidis Kirkaldy. 
Nesosydne neph'olepidis Kirkaldy, 1908, op. cit. p. 203, PI. 4, fig. i. 
Hab. Oahu, Tantalus. 



(8) Nesosydne iponioeicola Kirkaldy. 

Delphax pulcJira Stal, 1854, O. V. A. F. xi. 246; and 1859, Eugenies Resa, 
Zool. 275. 

Nesosydne ipomoeicola Kirkaldy, 1908, op. cit. 202, PL 4, fig. 4. 

Somewhat like A'", kalia, but the pronotal keels are more divergent and curving, 
and do not reach the hind margin ; the brachial and radial veins are forked near the 
subapical line ; there are five apical cells and the tegmina are a little longer. The frons 
is nearly parallel-sided, very slightly (and gradually) narrower basally than apically. 
Antennae shorter, scarcely reaching beyond the base of the ciypeus. Basal margin of 
tegmina fuscous. 

^. Head, pronotum and scutellum dark fuscous, or blackish, keels of head pale. 
Sterna, abdomen more or less, etc., dark fuscous or blackish. 

$. Similar to the same sex in N. kalia, but the abdomen is more obscure. 



HEMIPTERA 587 

Vars. Both sexes vary more or less in the following directions, head often scarcely 
fuscate. Apex of commissural and of subcostal cells, etc. dark fuscous ; these may 
meet and be extended in a large median black blotch in the $. Abdomen often orange 
red, more or less marked with black. Length ,? 2-J; ? 3^ — 3-| mm. 

Hab. Kauai, 400 ft. (Oct., Giffard) ; Oahu, Honolulu (Stal), Tantalus, 1300 ft. 
(Kirkaldy, Giffard, Perkins, Swezey, Terry), on Ipomoea bonanox, etc. ; Honolulu and 
Kalihi, sea-level to 50ft., on Jussiaea villosa (June, Swezey); Makiki, 150ft., on 
Dolichos lablab (March, Swezey). Hawaii, above Hilo, 1800 ft. (Dec, Perkins, two tt), 
Olaa (Nov., Perkins, one $). 

This is perhaps the commonest endemic Asiracid. I have not seen Stal's type, 
but I have no doubt of the correctness of the identification ; the name given by him, 
however, is preoccupied. 

Macropterous form. Head with keels brownish yellow, the intracarinal spaces more 
or less fuscous. Nota brownish yellow, more or less strongly and entirely suffused /^ 
with dark fuscous, the keels usually excepted. Tergites mostly blackish brown. 
Tegmina hyaline, veins pale testaceous, basal margin and a mark on the commissure 
blackish, apical veins (at least apically) and apical margin, also veins of wings, fuscous. 
Length 4 mm. 

(9) Nesosydne haniadryas, sp. nov. 

%. Shining dark fuscous ; keels of frons and clypeus narrowly margined with 
fuscous. Antennae, labium and legs, pale yellowish white. Scutellum piceous. Teg- 
mina hyaline iridescent, appearing blackish basally, owing to the abdomen showing 
through, veins fuscous. Head dorsally scarcely longer than broad, extending very little 
beyond the eyes ; lateral margins of frons subparallel. Antennae reaching well beyond 
the base of the clypeus, second segment one-half longer than the first. Tegmina reach- 
ing well beyond the abdomen, with six apical veins, second and third with a common 
base, and the third forked near its base. Pronotum with the lateral keels not reaching 
the hind margin but also not curving away. Spur with 10 spines. Length % 4^ mm. 

Hab. Oahu, Tantalus (i ?, Perkins, 1906). 

I am indebted to Dr Perkins for the loan of this, the only specimen he has taken. 



(10) Nesosydne haleakala, sp. nov. 

$. Brownish yellow ; genae, clypeus and apical part of frontal keel blackish brown. 

Teo-mina with a fuscous stripe obliquely from the base (exteriorly) to apex of clavus, 

darkening at the two ends. Tarsi blackish. Tegmina scarcely reaching beyond the 

middle of the abdomen, clavus not completely sutured off ; three apical veins, the sub- 

F. H. II. 76 



588 FAUNA HAWAIIENSIS 

costal cell acute apically. Second segment of antennae about one-fourth longer than 
the first. Length 3|- mm. 

Hab. Maui, Haleakala, 5000 ft. (by "grubbing," April, no. 251, 2 $?, also an 
example in very bad condition, Perkins). 



(11) Nesosydne nephelias, sp. nov. 

Z- Head, pronotum and scutellum blackish, the keels widely pale ; antennae and legs 
yellowish testaceous, femora long, striped with dark fuscous. Abdomen yellowish 
testaceous, partly suffused with brown. Tegmina hyaline cinereous, veins partly 
fuscous, a spot at the apex of the subcostal cell and the commissure at the apex of the 
clavus, dark fuscous. Venation irregular. 

%. Pronotum, scutellum and abdomen less dark. 

Hab. Lanai, 2000 ft. and over (Jan. — Feb., Perkins). 



(12) Nesosydne procellaris, sp. nov. 

^ %. Brownish yellow ; a line on the genae and the lateral margins of the pronotum 
and scutellum (rather feebly), dark fuscous. Tegmina brownish yellow, hyaline, base 
very narrowly and a transverse smudgy stripe just basal of the subapical veins, dark 
fuscous ; veins mostly pale brownish. Apices of tibiae, tarsi etc., dark brown. Teg- 
mina brachypterous, with five apical veins (sometimes one may be forked very close to 
the apical margin) ; the subcostal is undivided and its apical margin is a continuation of 
the subapical transverse vein. 

Z- Frons between the keels, apical part of vertex and posterolateral margins of 
scutellum all faintly fuscous. Sternites blackish brown, broadly bordered with yellowish, 
last segment yellowish, pygophor piceous, styles blackish ferruginous. Tegmina not 
reaching as far as the apex of the abdomen. Last tergite truncate apically ; pygophor 
not spined in profile, subcircular, lateral margins a little thickened and flattened. 

$. Frons between the keels, apical part of vertex, posterolateral margin of scutellum 
very narrowly, dark fuscous. Sternites yellowish, marked with brownish. Pygophor 
yellowish brown suffused basally with brown, ovipositor yellowish. Tegmina reaching 
just beyond the apex of the abdomen. Ovipositor narrow and parallel-sided for more 
than its apical half, basally considerably widened, lateral margins of pygophor sub- 
parallel , nebenplatten narrow and oblique. Length 4^^ mm. 

Hab. Molokai, 3000 ft. (June, 1^,1?, Perkins). 

This species is very probably now extinct. It is the bulkiest of the Hawaiian 
Asiracidae. 



HEMIPTERA 589 

(13) Nesosydne pahistris Kirkaldy. 

Nesosydne palustris Kirkaldy, 1908, op. cit. p. 202, PI. iv. fig. 7. 

^. Head, pronotum and scutellum sordid brownish yellow, scutellar keels darker, the 
median suffused ; anterior margin of head darker ; genae and clypeus with paler keels. 
Tegmina brownish yellow, hyaline, immaculate except a short, dark stripe on the 
commissure at the apex of the clavus ; also .some of the veins dark. Tergites pale 
brownish yellow, variegated with blackish brown. Legs and sterna pale yellowish. 
Second segment of antennae two-sevenths longer than the first. Spur with nine spines. 
There is no continuous subapical transverse line, and the clavus is not completely 
sutured off from the corium. There are four apical veins, the subcostal cell being acute 
apically. (In one example, there is a supplementary transverse vein in the interior 
discoidal cell of one tegmen.) Tegmina reaching not quite as far as the apex of the 
abdomen. Last tergite deeply acute-angularly emarginate ; pygophor in profile with a 
horizontal spine on the dorsal margin. Length 3^^— 3f mm. 

Hab. Molokai, 4950 ft. (Sept., Perkins) and 4500 ft. (Sept., Perkins), two tt on 
trees in the excessively wet bogs in the highest points of Molokai. This species is also 
probably extinct now. 

(14) Nesosydne mibigena, sp. nov. 

$. Closely allied to the last, but smaller. Tegmina reaching to about the middle 
of the pygophor, with two apical cells only. Spur with eight spines. Length 3 mm. 

Hab. Molokai, forest above Pelekunu (July, one $, Perkins). Also probably 
extinct. 



(15) Nesosydne cyathodis, sp. nov. 

Head brownish yellow, the apical margin of frons and genae, clypeus, antennae, 
legs (except spur), etc., dark fuscous or blackish brown. Tegmina whitish. Pronotum 
and scutellum fuscous with pale keels. Abdomen blackish with pale spots and a broken 
pale line down the middle. Tegmina reaching to about three-fourths the length of the 

abdomen. 

i pygophor black, styles, etc., black ; genital styles in shape somewhat like those 
o{ ckainhersi, but more elongate and narrower. Length if mm. 

The nymphs of the last instar are of the same general colour as the adult, but the 
frons is pale only basally, and the abdomen is rather more marked pallidly. 

Hab. Hawaii, Kilauea, 4000 ft. (April, Swezey, Giffard, May, Kirkaldy), on 

Pukeawe {Cyathodes tameiameiae). 

76 — 2 



590 FAUNA HAWAIIENSIS 

(i6) Nesosydne raillardiae Kirkaldy. 
Nesosydne raillardiae Kirkaldy, 1908, op. cit. p. 203, PI. iv. fig. 5. 
Hab. Hawaii, Kilauea, 4000 ft. (Swezey) on Raillardia. 

(17) Nesosydne chanibersi Kirkaldy. 

Nesosydne chanibersi Kirkaldy, 1908, op. cit. PI. iv. figg. 10 — 12. 

Yellowish, or pale ferruginous. Tegmina subhyaline, veins mostly concolorous 
with the prominent brown granules ; apical margin of tegmina narrowly suffused with 
brown, also a very little suffusing round each granule. Each tergite margined apically 
with dark fuscous. Length 3 mm. 

PL IV. figg. II — 12 of the work cited above, show the venational discrepancy in 
the same individual. 

Hab. Hawaii, Kilauea, 4000 ft. (April, Swezey). 

(18) Nesosydne argyroxiphii Kirkaldy. 

Nesosydne argyroxiphii Kirkaldy, 1908, op. cit. p. 201, PI. iv. fig. 6, and text-fig. i. 

Black ; keels of head and nota, a broken, very narrow line down the middle of the 
abdomen etc., testaceous. Antennae dark fuscous, partly stained with dirty testaceous. 
Tegmina subopaque, milky, veins fuscous. There are no apical cells and the tegmina 
do not reach to the middle of the abdomen. Length 3 mm. 

The nymphs of the last instar are brownish testaceous, largely suffused with dark 
fuscous. 

Hab. Maui, crater of Haleakala (Oct., Perkins, two %%, Swezey, Terry), on 
Ahinahina {Argyroxiphiuni sandwicense), at about 10,000 ft. Mr Swezey has kindly 
lent me a male, the only one I have seen, to. study the genitalia. 



(19) Nesosydne imbricola, sp. no v. 

^. Dark fuscous ; keels brownish yellow ; frons and clypeus with broad brownish 
yellow median and lateral keels. Antennae and legs brownish yellow, largely suffused 
with dark fuscous. Exterior half of clavus and apical fourth of tegmina cinereous 
hyaline, also most of subcostal cell, the rest fumate with one or two small hyaline spots ; 
a spot at apex of commissural area and another at apex of subcostal area, the subcostal 
vein, etc., dark fuscous. Tegmina feebly granulate, not reaching beyond about three- 
fourths of the length of the abdomen. Length 3 mm. 

Hab. Maui, Haleakala, 5000 ft. (March, April, one ^, Perkins). 



HEMIPTERA 591 

(20) Nesosydne monticola, sp. nov. 

?. Black ; submedian keels of vertex (and sometimes of the base of the frons) pale. 
Pronotal keels obscurely pale ferruginous. Tegmina cinereo-hyaline, largely suffused 
with fuscous (almost altogether except a hyaline transverse stripe near the apical 
margin); veins mostly dark fuscous. The frons has a few pale specks along the lateral 
margin and three along the middle keel on each side (the latter sometimes obsolescent). 
Antennae more or less dark fuscous. A row of pale specks down the middle of the 
tergites, the apical two or three segments spotted or specked (individually) pallidly. 
Tegmina reaching to about three-fourths of the length of the abdomen ; mostly four 
apical cells. Length 3 mm. 

Hab. Maui, Haleakala, 8000 ft. (April, three %%, Perkins). 



Nesothok Kirkaldy. 

Nesotho'c Kirkaldy, 1908, P. H. E. S. i. p. 202, fig. 2. 

An endemic genus of ten species, males of which 1 have seen in three species. It 
corresponds somewhat to Chloriona in the "laminate-spurred" series. Beyond the 
spur, it differs by the sudden compression of the tegmina at the base of the apical cells, 
the stronger excavation of the vertex and the union of the submedian keels of the frons 

o 

closer to the base of the latter. First segment of the antennae not, or scarcely, as long 
as wide, about one-third of the length of the second segment (typically), which is rather 
flattened ; clypeus tricarinate. Tibial spur subcultrate, compressed, solid, narrow and 
elongate, with 7 — 10 strong teeth. In all the species, the frons is marked transversely, 
more or less clearly, with pale, broken lines and spots. Type N. Jietus. 

The species are separable as follows : 

1. First segment of antennae blackish or dark fuscous (or if pale, then the second 

is blackish) ( 2 ) 

I a. Antennae pale (6) 

2. Frons basally dark with pale markings, apically white (3) 

2a. Frons pale brownish yellow with whitish specks and a whitish suffused blotch 

in the middle ; tegmina brown with a broad white basal band and an 

exterolateral crescent of the same hue i y?t'///j- Kirkaldy. 

3. Vertex and pronotum white ; tegmina immaculate 2 Imla Kirkaldy. 

3a. Vertex and pronotum pale yellowish or brownish; tegmina maculate (4) 

4. Apical third of tegmen not maculate, some of the veins narrowly suffused ■>> frigidula Kirkaldy. 

4a. Apical third of tegmen irregularly suffused, at least one of the cells smoky ...(5) 

5. Tibiae whitish, clearly annulate with brown 4 bobeae Kirkaldy. 

5a. Tibiae brownish yellow, feebly annulate $ perkinsi Kirkaldy. 

6. Apical part of tegmen not blotched (7) 

6a. Apical part of tegmen blotched with brown (9) 

7. Tegmen with an oblique band near the base (8) 

7a. Tegmen only suffusedly smoky 6 /aka Kirkaldy. 



592 FAUNA HAWAIIENSIS 

8. Larger, tegmina milky 7 piilani Kirkaldy. 

8«. Smaller, tegmina not milky, face more clearly spotted apically 8 terryi Kirkaldy. 

9. Second segment of antennae four times as long as the annuliform first q pluvialis Kirkaldy. 

9a. Second segment less than three times as long as the non-annuliform first 10 silvestris Kirkaldy. 

These species are distributed as follows : 

Kauai: liula, phivialis. Oah\i: perkinsi, bobeae, terryi. yioXokdJi: piilani. Lanai : 

silvestris. Maui : fletus, laka. Hawaii : frigidula. 

(i) Nesotho'c fietits Kirkaldy. 

Nesotho'e fletiis Kirkaldy, 1908, op. cit. p. 204. 

t%. Head and pronotum yellowish, with paler keels, the pronotum fuscous laterally 
with pale granules ; frons with a number of white specks all over the middle third, and 
a suffused white blotch about the same place ; clypeus fuscous. First segment of 
antennae black, second fuscous. Scutellum brownish ferruginous, with paler keels. 
Tegmina warm brown, the base broadly white, produced a little on to the clavus, a 
large crescentic white area on the apical half exteriorly and a few spots apically ; veins 
mostly pale with brownish granules. Sterna and sternites pale. Femora and tibiae 
piceous ; the former pale or sanguineous, basally and apically, the latter annulated with 
brownish yellow or sanguineous ; tarsi pale. Frons slightly more than twice as long as 
broad, laterally gently rounded. Second segment of antennae scarcely three times as 
long as the first, somewhat more rounded in profile than the other species, first about as 
broad as long. Spur with eight spines. Length $ \\; ? 4|- mm. 

The name "fletus" is in allusion to the battle of " Ke Paniwai " in 1776, when the 
lao River was " choked " with the slain. 

Hab. Maui, Western Mts, lao Valley (March, Perkins, one ?, and July, one ^, 
Swezey). Easily recognized by the broad warm brown band down the white tegmen. 

(2) Nesotho'e hula Kirkaldy. 

Nesotho'e hula Kirkaldy, 1908, op. cit. 

(?. Vertex and pronotum white, lateral margins of the latter blackish ; base and apex 
of frons rather narrowly, middle keel, three short transverse lines in the middle etc., 
white, the rest dark fuscous ; genae (except the white apex), clypeus, first segment of 
antennae, scutellum (except the posterior angle minutely, and a few specks on the 
posterolateral margins) black, keels of scutellum sometimes ferruginous; second segment 
of antennae fuscous. Tegmina hyaline, veins on clavus and corium pale, with minute 
fuscous granules, subcostal veins and veins on membrane fuscous, two or three blackish 
marks on commissure ; marginal vein fuscous. Legs pale, femora longitudinally striped 
with black. Abdomen apparently mostly black. Frons narrowed basally. Second 
segment of antennae about three times as long as the first, which is longer than broad. 
Spur with eight spines. ' 



HEMIPTERA 593 

%. The black in the male is replaced by yellowish brown, except the first segment 
of the antennae, and two or three spots on the vertex and a few on the genae, also 
the marks on the commissure. Sternites pale, ovipositor dark. Length ^ \\, % ^\ mm 

Easily recognized by the spotless tegmina and dead white head and pronotum. 

Hab. Kauai, high plateau (Aug., Perkins, two H, two ??, nos. 96 and 631). 

(3) Nesotho'e frigidula Kirkaldy. 

Nesotho'e frigidtda Kirkaldy, 1908, op. cit. 

%. Pale (or dark) yellowish brown, paler beneath ; apical third, or two-thirds, of 
frons and specks on the basal part pale ; second segment of antennae blackish. Tegmina 
hyaline tinged with cinereous, feebly granulate, veins almost concolorous, except the 
subcostal and 2nd — ^4th apicals, which are fuscous. Near the base of the tegmen there 
is a broad, suboblique, fuscous band. Wings hyaline, veins fuscous. Length 4^ mm. 

Hab. Hawaii, Kona, over 2000 — 3000 ft. (Sept. — Oct., 2 %%, Perkins). 

(4) Nesotho'e bobeae Kirkaldy. 

Nesotho'e bobeae Kirkaldy, 1908, op. cit. text-fig. 2 (venation). 

t %. Vertex and pronotum cinereous, the latter fuscous laterally with pale granules. 
Frons fuscous, (apically whitish), three or four elongate pale spots medially, the fourth 
nearly on the apical pale part, and very narrowly ringed with brown ; also pale specks 
laterally. Clypeus fuscous ; genae dark fuscous basally, pale apically. Scutellum sordid 
pale brown, fuscous laterally. Tegmina cinereous hyaline, veins pale, with dark fuscous 
granules closely set, corium and clavus suffused discally with fuscous, also sparsely on 
apical cells (especially the middle one), ist— 3rd apical veins suffused with dark fuscous. 
First segment of antennae black, second dark fuscous. Sterna and legs pale, the latter 
annulate with dark fuscous. Abdomen dark fuscous mostly. Frons about 2^ times as 
long as wide, narrow ; lateral margins vertically reflexed. Labium reaching nearly to 
hind coxae. Second segment of antennae nearly three times as long as the first, which 
is as wide apically as long. Spur with seven spines. Length 4 mm. 

In the male, the penultimate sternite is deeply subangulately emarginate, the last 
slightly so. 

Hab. Oahu, Honolulu Mts. (Dec, 2 <?^, Perkins), Tantalus, 1300 ft. (Dec, 
Giffard, Perkins), on Bobea. 

(5) Nesotho'e perkinsi Kirkaldy. 

Nesotho'e perkinsi Kirkaldy, 1908, op. cit. 

$. Vertex, basal two-thirds of frons, base of genae, scutellum, etc. piceous ; 
pronotum dark fuscous; keels, granules, and 15 — 16 specks on frons — some of which 
may fuse, forming three short transverse lines — pale brownish yellow. Apical third of 



594 



FAUNA HAWAIIENSIS 



frons and apical f of genae, whitish. First segment of antennae black, second brownish 
yellow. Tegmina cinereous hyaline, veins pale with fuscous granules, apical veins 
fuscous, first three suffused ; an irregular smoky smudge near the middle of the corium, 
another on the apical cells interiorly, spreading a little on to the corium. Legs pale 
brownish yellow, femora marked and tibiae annulated, with brown. Tergites piceous 
or blackish, sutures pale. Frons scarcely twice as long as broad, lateral margins 
regularly rounded, frons little wider apically than at the base. Second segment of the 
antennae more than three times as long as the first. Spur with nine spines. Length 
? 4|— 5 mm. 

Hab. Oahu, Honolulu Mts. (Sept., Perkins, over 2000 ft., Oct., Perkins, 5 %%) ; 
Tantalus (1906, Perkins); Palolo (March, 1800 ft., Swezey), on Myrsine sp. 



V--^ 



(6) Nesothoe piilani Kirkaldy. 

Nesothoe piilani Kirkaldy, 1908, op. cit. 

Pale yellow, disk of scutellum fuscous. Vertex and base of frons sordid pale 
brownish yellow, rest of frons paler and clearer, with scarcely perceptible paler, short, 
cross lines. Antennae testaceous, extreme apex of first segment partly fuscous. 
Tegmina milky hyaline, an undulating fuscous line near the base of the tegmen and 
parallel with it, faintly smoky a little apical of this. Veins mostly pale, apical ones 
a little darker, granules brown. Legs pale, apex of femora and base of tibiae a little 
fuscate. Tergites more or less fuscate. Second segment of antennae more than three 
times as long as the first, which is scarcely longer than wide, lateral margins regularly 
rounded, frons a little wider apically than at base. Ovipositor dark. Length 5 mm. 

Hab. Molokai, 3000 ft. (June, Perkins, i $ in bad condition). 



(7) Nesothoe terryi Kirkaldy. 

Nesothoe terryi Kirkaldy, 1908, op. cit. 

Very close to the preceding, but smaller, the frons more clearly spotted apically 
and the tegmina not milky. Length i^^ mm. 

Hab. Oahu, Waialua, Koolau range (Perkins, 3 $$, no. 428). 



(8) Nesothoe laka Kirkaldy. 

Nesothoe laka Kirkaldy, 1908, op. cit. 

%. Pale brownish yellow, scutellum fuscous between the keels. Frons slightly 
suffused with fuscous, with pale median keels and four specks on each side in a longi- 
tudinal row. Tegmina hyaline, tinged with yellowish cinereous, a little smoky in part ; 
marginal vein tinged with sanguineous ; other veins pale with closely set, small, fuscous 



HEMIPTERA 595 

granules ; a small short fuscous line on the corium near the apex of the clavus, the 
apices of most of the apical veins a little suffused. Legs yellowish white, fore tarsi 
banded with fuscous. Frons somewhat like that of N. fletus. Second segment of 
antennae about three times as long as the first, which is rather less cylindric than in the 
genotype, and which is much shorter than broad. Ovipositor dark. Length 5 mm. 

Hab. Maui, lao Valley, Western Mts. (March, Perkins, i %). 

(9) Nesothoc pluvialis Kirkaldy. 

Nesotho'e pluvialis Kirkaldy, 190S, op. cit. 

$. Pale yellowish (or tinged with sanguineous) ; keels, three transverse lines — not 
nearly reaching the lateral margins — a few spots on frons etc., paler. Tegmina hyaline, 
basal half (except the base, etc.) faintly smoky, an irregular brown blotch on the apical 
cells, apices of apical veins suffused ; granulations dark, close ; marginal vein sanguineous. 
Tibiae and tarsi pale, base of forme' piceous ; femora piceous, apex sanguineous. 
Tergites partly fuscous. Frons widening towards the apex, about three-fourths longer 
than wide aoically. Antennae like those of the genotype, second segment about four 
times as long as the first, which is annuliform. Spur with seven or eight spines. 
Length 4 — 4-^ mm. 

Hab. Kauai, Halemanu, 4000 ft. (no. 510, May, Perkins, i $), and Koholuamano, 
4000 ft. (no. 526, April, Perkins, 2 %%) ; Head of Koloko Ditch, 800 ft. (Oct., Giffard). 

(10) N^esotlioc silvestris Kirkaldy. 

Nesotho'e silvestris Kirkaldy, 190S, op. cit. 

Pale sordid fuscous ; the frons with three transverse lines and some specks, and the 
apical margins narrowly, whitish yellow ; genae apically pale ; clypeus dark fuscous ; 
antennae brownish white. Tegmina c'^^^reous hyaline ; veins naturally pale, but often 
suffused with dark fuscous ; a curved dark fuscous stripe on the subcostal, extending 
over the middle three-fourths and touching the radial internally about the middle of the 
scutellar margin triangularly from this inwards, the corium and a part of the clavus 
smoky ; apical and subapical veins mostly dark fuscous suffusedly, a blotch on the 
interior part of these cells ; granules rather feeble, with pale hairs ; subcostal cell pale 
brownish yellow, except where the curved stripe touches it basally and apically. Femora 
and tibiae brownish yellow, biannulate with dark fuscous, tarsi dark fuscous. Labium 
reaching to hind co.xae. Spur with eight spines. Frons about twice as long as broad, 
about as wide apically as basally, lateral margins gently rounded ; keels of genae and of 
frons not meeting apically. Second segment 2| times as long as the first, which is 
longer than broad. Length % \\ — 5 mm. 

Hab. Lanai, Koele Mts., 2000 ft. (Jan., Perkins, 2 $$). 

F. H. II. 71 



-\ 



596 FAUNA HAWAIIENSIS 

Nesodryas Kirkaldy. 

Nesodryas Kirkaldy, 1908, op. cit. pp. 201 & 203. 

Somewhat allied to Megamelus Fieber, but the basal segment of the antennae is 
shorter than wide, not more than one-fourth of the length of the thickened second ; the 
hind tibiae are distinctly longer than their tarsi, and the first segment of the latter is 
more than twice as long as the others together. Veins of tegmina feebly granulate ; 
five apicals, the second and third with a common stalk, fourth forked near apex. Tibial 
spur solid, narrow, with 4 — 7 strong spines. Type N. freycinetiae. The species of 
this genus are perhaps the most delicate of all the Asiracidae. 

The four species may be separated as follows : 

1. Vertex and pronotum each with two small, but conspicuous, dark fuscous 

spots I freycinetiae Kirkaldy. 

\a. Vertex immaculate (2) 

2. Vertex and pronotum immaculate, pale (3) 

2a. Vertex black 5 dryope, sp. nov. 

2b. Vertex pale, pronotum dark smoky 2 elaeocarpi \\\x\jd\A-^. 

3. (? $ • Pale greenish testaceous; tegmina milky coloured 3 giffardi Kirkaldy 

3a. (J 9 . Pale brownish yellow. $ tegmina greyish hyaline ; $ tegmina 

yellowish hyaline with the interior half pale orange brown, or yellowish 

fumate 4 ei/geiiiae Kirkaldy. 

(i) Nesodryas freycinetiae Kirkaldy. 

Nesodryas freycinetiae Kirkaldy, 1908, op. cit. p. 203. 

Z %. Greenish testaceous (the green often fading locally), or even changing to 
pale orange brown ; clavus and inner half of coriuni and of membrane, the abdomen, 
etc., suffused with pale reddish brown (sometimes also pronotum and scutellum). Vertex 
and pronotum each with a couple of small conspicuous blackish brown spots. Apex of 
first apical cell and adjoining exterior margin of tegmen, apex of inner apical cell, etc., 
fuscous. Vertical keels faint. Length ^ $ 4 — 4^ mm. 

The t pygophor has a very long, subascendant, lateral spine on each side ; the 
genital styles are narrow. 

The nymphs have been described by Swezey (1908, P. H. E. S. 11.). 

Hab. Oahu, Honolulu Mts., 2000 ft. (Nov., Perkins, i %) ; Pacific Heights ridge, 
1500 ft. (May, Swezey); Tantalus, 1300 ft. (Jan., April, Oct., Dec, Giffard), on leie 
{Freycinetia ai'borea) ; also reported from Acacia koa, but certainly in error. 

(2) Nesodryas elaeocarpi Kirkaldy. 

Nesodryas elaeocarpi Kirkaldy, 1908, op. cit. 

Head, antennae, etc., greenish testaceous. Eyes blackish grey. Pronotum dark 
smoky. Labium and legs (except the black tip of the former, and black hind femora 
and base of hind tibiae) testaceous. Tegmina hyaline with a broad dark fuscous stripe 



HEMIPTERA 597 

along the middle ; veins not on this stripe testaceous. Wings pale smoky with dark 
brown veins. Tergites black ; sternites mostly pale greenish. Pygophor, etc., black. 
Length % 4^ mm. 

Hab. Oahu, Tantalus (April, 1906, Perkins, Swezey), on Elaeocarpus bifidiis. 

(3) Nesodryas giffardi Kirkaldy. 

Nesodryas giffardiYJvcVsXAy, 1908, op. cit. 

Pale greenish, fading after death. Eyes and claws dark fuscous. Tegmina sub- 
hyaline milky, with white hairs ; veins greenish testaceous (fading), a short fuscous line 
at the apex of the commissural cell ; the base of the tegmina varyingly narrowly and 
rather faintly fuscous ; interior apical cell usually faintly fuscous. Labium reaching 
to, or a little beyond, the base of the hind femora. Length Z % 4 — 4i ittti- 

Hab. Oahu, Maunawili, 800 ft. (Febn, March, Giffard, Swezey), on Olona 
{Touchardia latifolid) and Cyrtandra grandiflora \ Tantalus (Perkins). 

(4) Nesodryas eugeniae Kirkaldy. 

Nesodryas eugeniae Kirkaldy, 1908, op. cit. 

J. Head, pronotum, scutellum and underside pale yellow. Tegmina subhyaline, 
appearing silvery grey basally owing to the dark body beneath, apical third pale smoky. 
Wings hyaline, veins dark. 

%. Pale luteous, or pale brownish yellow, testaceous beneath, immaculate ; scutellum 
and tergites pale orange brown. Tegmina with exterior half of entire length more or 
less hyaline, inner half yellowish fumate, or pale orange brown, veins and granulations 
pale ; sometimes the base of the commissural cell, apices of apical cells, etc., a little 
fuscous. Ovipositor blackish. Length Z 4. % Ah "Tirn. 

Hab. Oahu, Tantalus, 1300 ft. (Perkins; Jan., March, April, Giffard); Palolo 
(Sept., Swezey) on Ohiaha {^Eugenia sandwicensis). 

(5) Nesodryas dryope, sp. nov. 

$. Vertex shining black, also the pronotum (except the varyingly pale brownish 
posterior margin). Frons, genae, and antennae pale yellowish white, first segment of 
last partly blackish. Eyes reddish. Scutellum fuscous or blackish. Abdomen above 
blackish brown Tegmina hyaline, basal veins colourless, the apical veins, the subcostal, 
and the veins of the wings, fuscous ; an oblique fuscous smudge on the tegmina near the 
base, a dark fuscous mark near the apex of the clavus. Femora, and tibiae basally, 
dark fuscous, rest pale. Head dorsally longer than wide, produced a little in front of 
the eyes ; keels strong and well cut ; frontal keels uniting close to the base, filiform, not 

77—2 



598 FAUNA HAWAIIENSIS 

parallel. First segment of antennae annuliform, second short, not reaching to the base 
of the clypeus. Tegminal veins feebly granulate, five apicals, the second forked near 
the base. Spur with 5 — 6 spines. $. Ovipositor scarcely as long as the pygophor. 
Tegmina reaching well beyond the apex of the abdomen. Length 3-| mm. 

Hab. Oahu, Tantalus (Feb., 1906, Perkins, ?), ridge near Tantalus (Dec, 
Swezey, $) ; Kauai, Koloko (800 ft., Giffard, Oct.). 

The type is a Tantalus specimen. 

DiCTYOPHORODELPHAX Swezey. 
Dictyophorodelp/iax Svjeze.y, 1907, P. H. E. S. i. p. 104. 

(i) Dictyophorodelphax mirabilis Swezey. 

Dictyophorodelphax mirabilis Swezey, 1907, op. cit. p. 105 ; and 1908, op. cit. 11. 

2, text-figg. I — 4. 
Swezey has described and figured the ultimate nymph (1908, I.e.). 

Hab. Oahu, Konahuanui, 2200 — 2500 ft., in mixed sweeping, but probably on 
sedges (Swezey, Terry, Feb., Dec). 

Fam. CHERMIDAE. 

( = Psyllidae auctt. et huj. op. in. p. 113.) 
Dr Perkins' collections from 1892 — 1902 are in the hands, I understand, of 
Mr E. A. Schwarz. I add only two to my former enumeration, though I took at 
least ten new species recently during a visit to Kilauea, Hawaii. 

Hevaheva Kirkaldy (huj. op. in. p. 113). 

(i) Hevaheva monticola Kirkaldy. 

Hevaheva vionticola Kirkaldy, 1908, P. H. E. S. i. p. 205. 

I originally described this from a carded pair taken recently by Dr Perkins at an 
elevation of about 2000 ft. on Tantalus, Oahu. I have since secured a pair taken about 
the same place by Mr Swezey. 

Hab. Oahu. 

(2) Hevaheva stives tris Kirkaldy. 

Hevaheva silvestris Kirkaldy, 1908, op. cit. p. 206. 

Described from a single example taken by Dr Perkins on Tantalus ; I have since 
secured another specimen taken at the same place by Mr Swezey. 

Hab. Oahu. 



I 

J 



HEMIPTERA 



Fam. APHIDAE. 



599 



H. &. Barbsr, 
U. S. National Mu»oum, 
VVaa.iington, 0. C. 



There are apparently no endemic Aphidae in Hawaiia. In a recent paper (1907, 
P. H. E. S. I. pp. 99—102), I recorded four species, viz.: Myzus ciiricidus Kirkaldy, 
Aphis rosac (Linn.), Loxerates saccJiari (Zehntner) and Myzocallis kahaivaluokalani 
Kirkaldy. Aphids on Sorghum, Sonchiis, Hibiscus and Cabbage, have not been studied 
sufficiently fully yet, and I recently found another species at Kilauea, Hawaii, on 
grasses etc. All these are probably introduced. 

Fam. ALEYRODIDAE. 

Six species of Aleyrodes, none apparently endemic, have been enumerated by 
Kotinsky in my recent Catalogue of this family (1907, Bull. Board Agric. Hawaii 11.), 
viz. : giffardi, hibisci, kirkaldyi and sonchi of Kotinsky, vaporariorum Westwood, and 
an unnamed species on Coffea. 



Fam. COCCIDAE. 

Since the first part of this contribution was published, (huj. op. iii. pp. 102 etc.), a 
revised list has been issued (1904, Entom. xxxvii. pp. 226 — 30, with a bibliography), 
totalling 53 species, a net increase of five. 



6oo 



Supplement 
by R. C. L. Perkins 

to 
HYMENOPTERA. 

Previously treated in Vol. I. 

HYMENOPTERA. 

In describing numerous additional species of Hawaiian Hymenoptera, I have, for 
convenience, arranged the genera and species nearly in accordance with Ashmead's 
Classification, as set forth in Vol. i. pp. 279 — 281 of this work, and have adopted the 
superfamily Divisions, as enumerated by him. At the same time, I think it would be 
more natural to combine the whole of the Aculeata together with the Chrysididae, 
Bethylidae and the Proctotrupoidea in one large group, as in fact Sharp has already 
suggested. Failing this, a greater number of superfamilies, equivalent to those of 
Ashmead, should be adopted, e.g. one for the Chrysididae and another for the Myma- 
ridae. The latter appear to me to be equivalent to all the other families of the 
Chalcidoidea taken together. In any case Ashmead's characters for superfamilies and 
lesser groups need defining anew, for those that he gives are frequently altogether 
inadequate or incorrect. I need only instance the Chrysididae and Dryininae (Bethy- 
lidae) which he places in his Vespoidea, in spite of the fact that hosts of the species 
have not the thoracic structure he assigns to the superfamily. Similarly some charac- 
teristic Sphecoidea have the pronotal angles attaining the tegulae and would be placed 
in the Vespoidea in accordance with his tables. His characters for the Mymaridae are 
quite erroneous. In drawing up his generic characters I should imagine that his 
extraordinarily acute sight led him to almost entirely neglect the precaution of dissecting 
the smaller forms. The Monograph of North American Proctotrypidae, the most 
elaborated of his works, suffers greatly from this neglect. Most of the genera, that I 
have examined, in that group, have quite different mouth-parts from those described 
by him. It is much to be desired that a thorough revision of Ashmead's tabular 
classifications should be made by those entomologists, who have access to his collections. 
It is greatly to be deplored that he should not have lived to accomplish this himself, 
after having laboured so long and so successfully for the advancement of our knowledge 
of his favourite Order of insects. 



yi 



HYMENOPTERA 6o^i» 

APOIDEA. 

Fam. PROSOPIDAE. 

Nesoprosopis Perkins. 

Nesoprosopis Perkins, huj. op. i. p. 75. 

I have made a dichotomous table for distinguishing the numerous species of this 
large genus. The females I am unable to tabulate at present. Unfortunately one 

species N'. nivalis, of which the male is known, was omitted in the table. It would ' 

there be identified as N. anthracina, from which it is at once distinguished by the ' 

unusually long hairs clothing the front of the head and the thorax and the yellow base 
to the hind tibiae. 

Another species, N. ci'abronoides, was not known to me in the male sex, when I 
made the table, but I have since taken it numerously at Kilauea, the only known ) 

locality for this remarkable little insect. It may be included in the followine manner : 

20. Prothorax with yellow marking 20a. ' 

Prothorax without yellow marking 23. 

20a. Hind tibiae widely yellow at the base but the tarsi entirely dark N. crabronoides. 

Hind tibiae or tarsi or both otherwise coloured 21. 

Synopsis of Nesoprosopis $$. 

1. Genital armature with the stipites produced so that in dorsal aspect they 

extend far behind the apex of the sagittae, more or less membranous 
apically and forming an apical (often blunt) angle at the end of their inner 

margin, this apical part being n. ^re or less deflexed 2. 

Genital armature usually with the stipites hardly or not at all extending 
beyond the apex of sagittae in dorsal aspect, their apex well rounded not 
forming an angle inwardly with the inner margin and not membranous ...5. 

2. Posterior tibiae not distinctly whitish or yellow at the base 3. 

Posterior tibiae white or yellow at the base 4. 

3. Stipites of genital armature fringed with hairs right to the apical inner angle ...N'. simplex. 
Stipites with hairs on their outer margin, which become very short and incon- 
spicuous or altogether fail near the apex N. facilis. 

4. Process of eighth ventral segment with long bifurcation at apex; face not very 
wide N. chlorostida. 

Process of eighth segment without long apical bifurcation ; face very wide N. hirsutula. 

5. Second segment of the abdomen without a definite puncturation and never 
with a distinct metallic reflection, species rarely large 6. 

Second segment evidently punctured or with a metallic reflection ; species 
large 37- 

6. Process of eighth ventral segment linear or sublinear, not dilated ; face below 
the antennae never with three widely separated yellow spots 7. 

Process of eighth ventral segment nearly always dilated, at least expanded at 
its flexure; or if not dilated the face below the antennae bears three 
widely separated yellow spots • lo- 




] 

\ 

\ 



602 FAUNA HAWAIIENSIS 

7. Scape of antennae not much dilated, about twice as long as wide on its broad 

surface 8. 

Scape of antennae strongly dilated, subtriangular, not twice as long as wide ...N. anthracina. 

8. Cheek between the eye and mandible distinct and rather long, plate above the 

clypeus generally narrow and elongate, basal joint of antennae less arched 

beneath N. volcanica. 

Cheek short or very short, plate above the clypeus short ; scape more strongly 
arched beneath 9. 

9. Process of eighth ventral segment without a longish apical bifurcation ; pubes- 

cence of mesonotum rather long N. difficilis. 

Process of eighth segment with longish bifurcation : pubescence of meso- 
notum short N. laeta. 

10. Scape of antennae little or not at all dilated, twice as long as wide or more 

than twice, face below the antennae wholly or nearly all yellow, at most 
the post-clypeal plate and the clypeal sutures being black, the yellow 
colour always continued back above the line of insertion of the antennae 
along the inner orbits ; abdomen never conspicuously ferruginous on the 

basal segments 11. 1 

Scape nearly always distinctly, often strongly, dilated, nearly always less than j 

twice as long as wide ; in species, where least dilated, with the hind 
margin at least strongly curved not very gently curved as in the above 
group, frequently subtriangular or even subcordiform ; face in front of the 
antennae rarely largely yellow and with this colour produced behind the 
antennae along the eye-margins and if so then either the hind margin of 
the scape is quite strongly rounded or the basal abdominal segments are 
ferruginous 15. 

11. Face wholly yellow in front of the antennae 12. 

Face not wholly level in front of the antennae, at least part of the post-clypeal 

plate black, the yellow colour produced back along the eye-margins as a 

narrowing vitta N. ohscurata. 

12. Scape of antenna with a yellow mark apically or with a complete yellow line ...N. blackburni. 
Scape not so marked ; face elongate 13. 

13. Posterior tibiae yellow with a fuscous stain behind N.flainpes. 

Posterior tibiae black at most yellow at the base and apex 14. 

14. Face longer, apical margin of post-clypeal plate much shorter than its sides to 

the antennal fossa N. longiceps. 

Face shorter, apical margin of post-clypeal plate about as long as its side N. finitima. 

15. Abdomen not conspicuously ferruginous on the basal segments; at most (as 

a variation) with a piceous, dull reddish or immature appearance 16. 

Abdomen conspicuously ferruginous basally 35- 

16. Scape of antennae not arched beneath, its lower edge straight or almost 

straight 17. 

Scape distinctly arched beneath 20. 

17. Anterior area of propodeum somewhat coarsely rugose all over N. haleakalae. 

Anterior area at most with short wrinkles in front; species very small 18. 

18. Process of eighth ventral segment not distinctly dilated N. tmitata. 

Process dilated dorsoventrally 19. 

1 9. A very conspicuous deep black median area behind the antennae N. dnmetorum. 

Deep black area not or hardly noticeable N. angustula. 

20. Prothorax with yellow marking 21. 

Prothorax without yellow marking 23. 



HYMENOPTERA 603 

21. Face extremely wide; the supra-clypeal plate extremely wide and short 22. 

Face not wide and transverse ; supra-clypeal plate with the apical margin 

about equal in length to its side as far as the antennal fossa N. kona. 

22. Clypeus for the most part or entirely yellow ; wings nearly clear ^V. kauaiensis. 

Clypeus with a yellow spot, wings dusky N. unica. 

23. Yellow markings of face continued up far behind the antennae along the eye- 

margins as a broad vitta N. inquilina. 

Yellow markings rarely continued back behind the antennae and then only for 
a short distance or as a rapidly narrowing vitta 24. 

24. Face extremely wide and transverse across the eyes in front view ; clypeus 

black, but yellow spots on each side of the face between the clypeus and 

the eyes are sometimes present N. laticeps. 

Clypeus generally yellow-marked, if black the face is not transverse 25. 

;5. Scape of antennae very strongly dilated, subcordiform ; face below the an- 
tennae entirely yellow or almost entirely N.flavifrons. 

Scape not so shaped, often elongate triangular, the anterior margin not 

strongly rounded 26. 

6. Clypeus with a large yellow spot or all yellow, sometimes there is a small spot 

outside the clypeus near to or touching its side margin on each side 27. 

Clypeus either with a small yellow spot or entirely black, or if with a large 
yellow spot or nearly all yellow then there is a large yellow generally more 
or less triangular-shaped spot between the clypeus and the eye-margin on 

each side, these spots always distinct from the yellow clypeal spot 28. 

Wings dusky, hind tibiae not yellow at the base, plate above the clypeus 

black N. koae. 

Wings clearer, hind tibiae yellow at the base, supra-clypeal plate more or less 

yellow N. vicina. 

\ Very small narrow species, with excessively short wrinkles at the anterior 

' margin of the propodeum, the rest of the dorsal surface of which is 

smooth, except for microscopic surface sculpture ; clypeus black, meso- 

thorax not clothed with dark hairs N. mauiensis. 

Species not very small and narrow ; propodeum generally rugose or rugulose 
to the brow or nearly ; clypeus generally yellow-marked, if black, the 

mesonotum is clothed with long black hairs 29. 

Clypeus and sides of face either black or yellow-marked, variable, antennal 
scape very deeply arched beneath ; mesonotum with long erect black 

hairs N. inelanothrix. 

\ntennal scape gently or moderately arched, mesonotum not clothed with long 

black hairs 30. 

ate above the clypeus always short and wide, wider than long to the point 

where its sides meet the antennal fossae 31. 

ite above the clypeus as long or longer than wide to the point where its 

sides meet the antennal fossae 3r. 

■peus only bearing a yellow spot N. comes. 

:e below the antennae with three yellow spots N. coniceps. 

peus black N. ombrias. 

peus yellow or marked with yellow 33. 

ad posteriorly strongly emarginate, so that the vertex is much shorter 

medially than at the sides N. sphecodoides. 

:ad lightly emarginate, the length of vertex in the middle and at the sides 
not greatly different 34. 

I. 78 






6o4 FAUNA HAWAIIENSIS 

34. Wings darker and facial markings generally much smaller N. assimulans, var. oahuensis. 

Wings clearer and facial markings larger N. assimulaiis. 

35. Yellow colour of the face continued far behind the antennae as a broad vitta 

along the eye-margins 36. 

Yellow colour if continued behind the antennae forming a narrowing vitta, 

pointed at its termination N. volatilis. 

36. Scape of antenna pale above anteriorly, mandibles with a yellow line N. hostilis. 

Scape entirely pale above, mandibles without a yellow line N. hilaris. 

37. Abdomen ferruginous N. paradoxica. 

Abdomen not ferruginous 38. 

38. Scape excessively dilated, the front margin very strongly rounded (prothorax 

never with a yellow line or spots) i(). 

Scape not excessively dilated, its front margin at most only gently rounded, 
prothorax sometimes with a pale line or spots 40. 

39. Scape of antennae strongly transverse, front margin extremely strongly rounded N. insignis. 
Scape less wide and its front margin less strongly rounded N. saiel/es. 

40. Face dotted with rather short erect black hairs, for the most part evenly, 

between the antennae and apex of clypeus ; two large facial spots below 

the antennae and sometimes the clypeus also with pale marking N. setosifrons. 

Face not so clothed 41. 

41. Prothoracic pale band very broad, hind tibiae almost entirely creamy white, 

wings very dark with steely iridescence, face black (? always) N. perspicua. 

Without some or any of these characters 42. 

42. Wings above (when expanded) fuscous, without conspicuous blue or violet 

iridescence 43. 

Wings with conspicuous blue or violet iridescence 45. 

43. Face below the antennae with pale markings N. anomala. 

Face without pale markings 44. 

44. Pronotum with an entire or almost entire pale band N. fuscipetinis. 

Pronotum rarely with an entire band sometimes all black N. puhesceiis. 

45. Apical abdominal segments with dark hairs, second ventral segment less raised 

from the base, pronotum always with an entire or almost entire band AL caeruleipennis. 

Apical segments pale-haired, second ventral segment more raised, pronotum 

black, or with pale spots or a broken band, rarely with an entire one N. puhesceiis var. 



( I ) Nesoprosopis ombrias, sp. nov. 

Male black, the legs more pitchy, the tarsi paler, the front tibiae yellow in front 
the flagellum of the antennae dull reddish beneath, clypeus black, a yellow spot on the 
face on either side of it, probably sometimes absent. Cheek very short, the base of the 
mandibles nearly touching the eye, the plate above the clypeus not very wide, its apical 
side about as long as its lateral one to the antennal fossa, the front of the head densely 
and shallowly punctured. Scape of antennae subdilated, its upper side strongly rounded, 
about twice as long as its greatest width, seen from in front slightly arched beneath. 
Mesonotum excessively dull with dense minute surface sculpture, finely and very shallowly 
punctured, the scutellum more closely. Propodeum with its anterior area entirely 
rugose. Wings smoky brown, with some blue iridescence. Abdomen less dull than 



HYMEN OP TERA 



605 



the thorax, having a very fine and close surface sculpture, and excessively fine indefinite 
puncturation. 

Female, except for the usual sexual differences, like the male in sculpture, etc., the 
face entirely black, the front tibiae somewhat reddish in front. Mesonotum and 
scutellum excessively dull, finely and very shallowly punctured and the propodeum 
rugose as in the other sex. Wings smoky brown. Puncturation of the abdomen 
excessively fine and feeble or indefinite, apical segment clothed with fuscous hairs. 
Larger than the Z. Length 7 — 10 mm. 

Closely allied to A^. assimulans, but quite easily distinguishable. The female is 
one of the largest of the species having no well-defined abdominal puncturation. 

Hab. Hawaii, Kona district. 

(2) Nesoprosopis flavipes, Sm. 

Nesoprosopis flavipes Smith, huj. op. i. p. 99. 

This is apparently a distinct species, which I have now taken on Hawaii. It is 
extremely close to A'^. blackburni Sm. 

Hab. Hawaii, a coast or low-land species. 



Fam. MEGACHILIDAE. 
(i) Lithiirgus sp. ? 
Hab. Oahu, widely distributed and sometimes abundant. 



First appeared 



about 1 900. 



SPHECOIDEA. 

( = P'ossores huj. op. i. p. 7.) 

Fam. PEMPHREDONIDAE. 

Subfam. PSENINAE. 



Stigmus Jur. 



(0 



Stiomus mordinatus Fox. 



Stiznms mordinatus Fox. 

A single specimen of a Stigmus was taken in Honolulu some years ago, but the 
species has not since been met with. 1 sent a description to Dr Ashmead of Washington, 
who informed me that it agreed well with the above, found in California and Colorado. 

Hab. Oahu, Honolulu. 

78—2 



1 



i 



v"l 



606 FAUNA HAWAIIENSIS 

Fam. TRYPOXYLONIDAE. 

Trypoxylon Latr. 

(i) Trypoxylon bicolor Sm. 

Identified by Dr Ashmead. It is an Oriental species, not known to occur in 1897, 
but common in 1900 at Hilo, Hawaii. It has now spread to Oahu and probably to the 
other islands, as it often nests in furniture, chairs, etc. 

Hab. Hawaii, Oahu. 



PisoN Spinola. 



Vide huj. op. i. p. 14. 



( I ) Pi son sp. ? 

A third species of this genus has been introduced recently and is now very abundant 
in Honolulu. I find that P. iridipcnnis was common here in 1900, but it is now lost 
amongst the great numbers of the new species. 

Hab. Oahu, Honolulu. 

Fam. CRABRONIDAE. 

In the Trans. Ent. Soc. London, 1902, p. 145 et seq. I divided the Hawaiian 
Crabronids into five groups or genera, including N'esocrabro, already characterized in 
this work. 

Xcnocrabro includes all the species other than those previously referred to Nesocrabro, 
excepting Crabro adnormis now Oreocrabro, Crabro tumidoventris now Hylocrabro, and 
Crabro curtipes now Melanocrabro. The species assigned to Xenocrabro are clearly very 
closely allied to the common European species called Crabro or Solenius vagus, and 
I believe the other Hawaiian forms are all related to or derived from Xenocrabro. 

Nesocrabro Perkins. 

Nesocrabro adspectans, Blackb. 
Nesocrabro daemonius Perkins, huj. op. i. p. 28. 

Having seen the types of N. adspectans, I see no specific characters to distinguish 
N. daemonius. 

Nesocrabro rubrocaudaius, Blackb. 
Nesocrabro bidecoratus Perkins, huj. op. i. p. 27. 

As I anticipated, this is nothing more than a very remarkable variety of the above 
species. 



HYMENOPTERA 607 

Fam. EUMENIDAE. 
PsEUDOPTEROCHEiLUS Perkins. 

Pseudopterocheilus Perkins, Ent. Mo. Mag. xxxvii. p. 266. 
To this genus belong the following species. 

Pseudopterocheilus pte7'ocheiloides, P. 
Odynerus pterocheiloides P., huj. op. i. p. 6g. 

Pseudopterocheilus congruus, Sm. 
Odynerus congruus Smith, huj. op. i. p. 69. 

Pseudopterocheilus relicttis P. 

Pseudopterocheilus relictiis P., Proc. Hawaii Ent. Soc. i. p. 63 ; not Odynerus 
relicttis P., huj. op. i. p. 74. 

Obs. There is found on the coast of Kauai a species, the male of which is almost 
inseparable from P. congrmis and P. hatvaiiensis. Until the female is discovered, it, 
however, would be rash to consider it as being identical with either of these species, as 
the males of Pseudopterocheilus exhibit specific differences much less strongly than the 
females. 

Chelodynerus P. 
Chelodynerus Perkins, Tr. Ent. Soc. London, 1902, p. 136. 

Chelodynerus chelifer, P. 

Odynerus chelifer P., huj. op. i. p. 70. 
Chelodynerus P., Proc. Hawaii Ent. Soc. i. p. 68. 

Nesodynerus p. 

Nesodynerus Perkins, Ent. Mo. Mag. xxxvii. p. 277. 

The following species mostly referred originally to Odynerus belong to my genus 
Nesodynerus. 

Nesodynerus I'udolphi, D. T. 
Odynerus rudolphi D. T., huj. op. i. p. 46. 

Nesodynerus conifer, P. 
Odynerus conifer P., huj. op. i. p. 38. 



V 



6o8 FAUNA HAWAIIENSIS 

Nesodynerus dilatatipes, P. 
Odynerus dilatatipes P., huj. op. i. p. 37. 

Nesodynerus vittativentris, P. 
Odynerus vittativentris P., huj. op. i. p. 65. 

Nesodynerus cooki, P. 
Odynerus cooki P., huj. op. i. p. 64. 

( I ) Nesodytierus paractias P. 
Nesodynerus paractias P., Pr. Hawaii Ent. Soc. i. p. -Ji. 

Nesodynerus oblitus, P. 
Odynerus oblitus P., huj. op. i. p. 74. 

Nesodynerus eupferyx, P. 
Odynerus eupteryx P., huj. op. i. p. 47. 

(2) Nesodynerus optabilis P. 

Nesodynerus optabilis P., Proc. Hawaii Ent. Soc. i. p. 61. 

This species has not been described at length, but only distinguished by such 
characters, as are given in my tables of the species of Odynerus of the island of Oahu. 
It is superficially somewhat like N. rudolphi, Odynei'us montanus, O. unicus and others 
from Oahu. 

Black, shining, wings deeply infuscate and with purple iridescence. Clypeus 
impressed at the apex and distinctly emarginate. Head shining, puncturation inclining 
to be rugose. Mesonotum with the larger punctures very sparse and irregular, the finer 
ones between these distinct, irregular and not dense, parapsidal furrows complete and 
distinct ; scutellum finely and sparsely and rather indefinitely punctured, propodeum 
nearly smooth, shining, trenchant at the sides. Basal abdominal segment shallowly and 
remotely punctured, second very convex, subtuberculate in dorsal aspect ; beneath, the 
costae moderate, the depression subtriangular, distinct, but not deep. Wings with the 
two recurrent nervures about as distant from each other, as the first is from the basal 
lower angle of the second cubital cell, distance from second recurrent nervure to the 
apical lower angle of the cell rather greater than the distance between the recurrent 
nervures ; the second cubital cell is wider above, less triangular, than in N. rudolphi. 

Hab. Oahu, Waialua district, outside the forest, flying over ferns. 



HYMENOPTERA 609 

Odynerus iopteryx P. 

O. iopteryx Perkins, huj. op. i. p. 63. 
O. illudens P., Ent. Mo. Mag. (2) xii. p. 267. 

The unique specimen, on whicli I described O. iopteryx, was a somewhat abnormal 
example. 

Hab. Oahu ; widely distributed, coast and mountains. 

Odynerus acoelogaster P. 

O. acoelogaster P., huj. op. i. p. 65. 

O. relictiis P., t. c. p. 74. 

O. lithophilus P., Ent. Mo. Mag. (2) xii. p. 267. 

O. lithophilus, Pr. Hawaii Ent. Soc. i. p. 72. 

The original description was based on an abnormal e.xample ; O. relictus was 
founded on a very old unique specimen in the British Museum from the Beechey 
collection. The proper description of this variable species is in the Proceedings of the 
Hawaiian Entomological Society as cited above. The original name is fortunately 
quite applicable to the species in all its varieties. 

Odynerus egens P. 

O. egens P., huj. op. i. p. 61. 
O. infanstus P., t. c. p. 73. 

The latter described from a very old specimen of the Beechey collection is no doubt 
identical with O. ep-ens. 



i' 



(i) Odyneriis thersites, sp. nov. 

Black, teeth of mandibles more or less red, sometimes a spot behind the eyes and 
a median frontal spot yellow. Wings hyaline and subinfuscate, with a bluish or 
violaceous iridescence in some lights. General appearance that of O. peles, but the 
clypeus is quite deeply emarginate and bidentate ; the head above the antennae with 
excessively dull surface and a very scanty and feeble, in fact hardly discernible punctu- 
ration. Mesonotum excessively dull, with remote and very faint punctures, the 
propodeum rugose. Basal segment of the abdomen with largish but shallow punctures, 
the second simply convex, beneath the costae well-developed, the depression not defined 
but represented by a faint median impressed line. Male ; length 6 — 8 mm. 

An overlooked species, very likely previously confounded by me with the variable 
^)eles, but nevertheless distinct. 
PP- '-f AB. Hawaii in the mountains at 4000 ft. or more. 

F. H. 



6io FAUNA HAWAIIENSIS 

Odynerus peles P . 

O. peles P., huj. op. i. p. 39. 

The puncturation of this species varies a good deal in intensity and I do not feel 
sure that, even after the segregation of O. thersites described above, there are not more 
than one species under the name of O. peles. Specimens taken in wet districts at 
moderate elevations of two or three thousand feet and others found below the 1000 ft. 
line are generally more strongly punctate than those taken at 4000 ft. and upwards. 

(2) Odynerus lipocharis, sp. nov. 

Black, the first abdominal segment inconspicuously or interruptedly margined with 
yellow. Wings very darkly infuscate and with blue iridescence. Clypeus impressed at 
apex and bidentate, but only slightly emarginate, closely punctate. Front very closely 
rugosely punctate. Mesonotum coarsely and closely punctured, the interstices between 
the coarse punctures shining and punctulate ; scutellum strongly punctured. Propodeum 
rugose, the side-margins explanate. Basal abdominal segment strongly and closely 
punctured, second segment very strongly tuberculately raised, highest point rather in 
front of the middle of its length ; beneath with rather shallow, but distinct, depression, 
the middle costae long. Male ; length 11 mm. 

Allied to O. konanus, hiloensis etc., and superficially unlike any other Kauai species, 
as well as being quite distinct structurally. 

Hab. Kauai, near Lihue, at no great elevation above the sea. 

(3) Odynerus subegens, sp. nov. 

Very like Odynerus egens P., but readily distinguished by the puncturation of the 
thorax, which, though very fine and feeble, is distinct. The apex of the clypeus is 
not red. 

Hab. Hawaii, 4000 ft. 

The following species of Odynerus have been described by me since the publication 
of Vol. I. pt. I of this work, in the Proc. Hawaii Ent. Soc, Vol. i. pp. 70 — 74 and in 
the preceding tables, pp. 61 — 70. 

(4) Odynerus homoeophanes P. 
O. homoeophanes P., t. c, p. 70. 

Hab. Oahu. 

(5) Odynerus euc harts P. 
O. eucharis P., t. c. p. 71. ii 

Hab. Oahu. 



HYMEN OP TERA 6 1 1 

(6) Odynerus pseudochronioidcs P. 
O. pseudochromoides P., t. c. p. 71. 
Hab. Oahu. 

(7) Odynerics paludicola P. 
O. paludicola P. , t. c. p. 71. 

Hab. Oahu. 

(8) Odynerus paranaias P. / 
O. paranaias P., t. c. p. 71. / 
Hab. Oahu. 

(9) Odynerus epipseustcs P. 
O. epipseustes P., t. c. p. 72. 

Hab. Oahu. 

(10) Odynerus taupe P. 
O. lempe P., t. c. p. 72. 

Hab. Maui. 

(11) Odynerus monas P. 
O. monas P., t. c. p. T}^. 

Hab. Molokai. 

(12) Odynerus aprepes P. 
O. aprepes P., t. c. p. ']'iy. 
Hab. Maui. 

(13) Odynerus xerophilus P. 

(9. xerophilus P., t. c. p. 72. 

Represents on Oahu the O. scoriaceus P. of Hawaii and the O. niolokaiensis of the 
intermediate islands. 
Hab. Oahu. 

(14) Odynerus acyanus P. 
O. acyanus P., t. c. p. 63. 
Hab. Oahu. 

The following species were described in the Trans. Ent. Soc. London, 1902, 
pp. 131 — 140. 

F. H. II. 79 



I 

1 



f 



6i2 FAUNA HAWAIIENSIS 

(15) Odynerus cypris P. 
O. cypris P., t. c. p. 138. 
Hab. Hawaii. 

(16) Odynerus eutretus P. 
O. ezdretus P., t. c. p. 138. 
Hab. Hawaii. 

(17) Odynerus holomelas P. 
O. holomelas P., t. c. p. 138. 
Hab. Hawaii. 

(18) Odynerus crypterythrus P. 
O. crypterythrus P., t. c. p. 139. 
Hab. Hawaii. 

(19) Odynerus newelli P. 
O. newelli P., t. c. p. 139. 
Hab. Hawaii. 

(20) Odynerus mesospilus P. 
O. mesospilus P., t. c. p. 140. 
Hab. Hawaii. 

Fam. BETHYLIDAE. 

Vide p. 282 huj. op. Tom. i. 

The Hawaiian species of Scleroderma and Sierola are endemic, those of Holepyris 
are introduced and usually found in houses. One or two other genera also occur in 
houses or more rarely outside, being parasitic on insects attacking grain or seeds, while 
one is parasitic on the minute beetle, Hypothcncmiis eritditus Westw. 

(t) Scleroderma euprepes, sp. nov. 

Head black, two basal joints of antennae, the legs, and the pronotum yellow, rest 
of the antennae sordid yellow or infuscate, the rest of the thorax brown, very dark at 
the sides, along the anterior margin of the propodeum, abdomen dark brown. 

Head much wider than the thora.x, with minute microscopic reticulation, which 
prevents it from being very shining, and some sparse feeble punctures, sparingly pilose. 
Length of eye about half the distance from its upper edge to the occiput. Antennae 



HYMEN OP TERA 6 1 



J 



rather long, the pedicel about as long as the two following joints together, the third a 
little longer than the fourth, none of the funicle joints being at all strongly transverse. 
Thorax duller than the head, similarly sculptured, the propodeum and pronotum sub- 
equal, the former widening posteriorly and impressed along its front margin. Abdomen 
much more shining than the thorax and as long or rather longer than the head and 
thorax together. Length, % 4-25 mm. 

Hab. Kauai, 4000 ft. 



(2) Scleroderma poc diodes, sp. no v. 

Head dark brown, pronotum yellow, rest of thorax dull brown, the propodeum 
more or less yellowish. Abdomen brown at base and apex, the intermediate segments 
black or nearly so, the apical margins being pale, the basal segment yellowish at the 
sides basally. Legs with the femora and tibiae brown, more or less yellow-tinged, the 
tarsi paler, yellowish. Antennae yellow, the flagellum with the joints more or less 
sordid or darkly annulate. 

Head shining, much wider than the thorax, with very fine reticulate sculpture of 
the surface, the eyes about half as long as the distance from their upper margin to the 
occiput ; antennae stout, the pedicel about as long as the two first funicle joints, the 
funicle joints short and most of them strongly transverse. Thorax much duller than 
the head, similarly but rather more strongly sculptured, and like it sparsely pilose, the 
proportion of the parts much as in 6'. ejtprepcs ; abdomen shining, with very fine but 
evident sculpture of the surface, as long or rather longer than the head and thorax 
together. Length, % 3 mm. 

Hab. Oahu ; near Honolulu, 1500 ft. 

(3) Scleroderma chlorodes, sp. nov. 

Head yellowish brown or testaceous, the thorax a little paler, yellow, legs entirely 
pale, concolorous with the thorax or almost so. Abdomen with the basal and apical 
segments more or less pale, being brown or rufescent, the intermediate segments black 
or blackish brown with pale apical margins. Scape and pedicel of the antennae yellow, 
the rest also yellowish, but infuscate, or with dark annulations. 

Head somewhat, but not much, shining, owing to the surface sculpture, and bearing 
a few very fine feeble punctures ; antennae shortish and thick, formed much as in 
S. poecilodes, with the funicle joints mostly very short and transverse. Thorax dull, 
propodeum a good deal wider posteriorly than on its anterior margin. Abdomen more 
shining than the thorax, but with the surface sculpture distinct, sparsely pilose, about as 
long as the head and thorax together. Length, $ 3 mm. 

Hab. Oahu ; Waianae mountains. 

79—2 



6 14 FAUNA HAWAIIENSIS 

(4) Scleroderma sophorae, sp. nov. 

Head and thorax brown, rarely altogether concolorous, the prothorax usually being 
distinctly paler (more yellowish) than the sordid parts behind it, the head too (except 
anteriorly) is more often concolorous with the paler pronotum, than with the mesonotum 
and propodeum. Antennae entirely pale, the funicle joints with fuscous rings. All the 
legs are yellowish-tinged, but they are for the most part of a dark colour, brownish or 
sordid, with yellow tarsi. The basal segment and tip of the abdomen are brown or 
yellowish brown, the intermediate segments black or dark brown, with the apices some- 
what widely pallid. When the intermediate segments are paler than usual, the basal 
segment is always paler still (more yellowish). 

Head very long and narrow, parallel-sided, the eyes only about one-third as long 
as the distance from their upper margin to the occiput ; the surface appears smooth, 
the minute reticulation, distinct enough under high powers of the compound microscope, 
being invisible under a very strong lens ; on the thorax, however, the reticulation can 
be distinguished with the latter, appearing as an excessively fine puncturation or 
shagreening. The antennae are short, the funicle joints for the most part strongly 
transverse, the pedicel as long as the two next joints together. Abdomen more shining 
than the thorax, and as long or a little longer than the latter and the head together. 
Length, ? 2*5 mm. 

Hab. Hawaii, 4000 ft. ; bred from the dead wood of Sophora. 

Obs. It is clear to me from the examination of this species and from others, of 
which I have seen a number of examples obtained in company, that slight differences 
of colour are not of specific value in this genus. Further, the abdominal segments are 
subject to retraction and I suspect that Ashmead's S. breviventre is an unnaturally con- 
tracted specimen. 

SiEROLA Cameron, 
(i) Sierola dichroma, sp. nov. 

Female black, the lower part of the face, the head about the posterior angles, the 
pronotum and mesonotum ferruginous, legs, scape of antennae and mandibles paler, 
yellow testaceous or whitish, rest of antennae infuscate or sordid, abdomen brown-black, 
paler in parts. 

Male ferruginous, a dark cloud about the ocelli, a black transverse one along the 
scutellar and propodeal suture, the base and about five transverse indefinite fasciae on 
the abdomen dark, mandibles whitish, legs pale yellow, base of antennae clear yellow, 
the rest more sordid or brownish. 

Head in both sexes very broad, the temples rounded, the surface dull and cori- 
aceous and with distinct, remote, shallow punctures. Antennae moderately long, the 



HYMENOPTERA 6 1 5 

flagellar joints moniliform, not transverse, the pedicel and first funicle joint cylindrical, 
the former slightly the longer, both being decidedly elongate. Thorax not quite so dull 
as the head, the mesonotum with shallow sparse punctures, the rest almost impunctate, 
but with minute surface sculpture. Wings subinfuscate, stigma and parastigma dark 
fuscous, the veins lighter, but still dark. Abdomen smooth and shining, the apical 
margins impressed so as to make large angular emarginations in the surface of the 
segments in the middle. Length 3 — 3-5 mm. 

In some examples the abdomen is nearly uniformly dark brown. 

Hab. Oahu ; 1500 ft. and upwards. 



HoLEPVRis Kieff 

( I ) Holepyris f hospes, sp. nov. 

Black, the antennae and legs ferruginous, the coxae mostly dark brown, the front 
femora posteriorly and the hind ones outwardly more or less infuscate or darkened, 
mandibles, clypeus, apical abdominal segment, and the impressed margin of the pro- 
notum also ferruginous. 

Clypeus with a longitudinal median carina, head dull, finely and not densely, but 
distinctly, punctured, eyes pilose. Antennae 13-jointed, the pedicel subequal to or 
rather longer than the first funicle joint ; joints of the funicle mostly subequal and a 
little longer than wide. Pronotum as long as the mesonotum and scutellum together, 
very dull and very finely punctured, the punctures of unequal size. Mesonotum much 
shorter than the scutellum, the latter with a transverse basal impression, both dull and 
finely punctured ; propodeum longer than the mesonotum and scutellum together, 
abruptly truncate posteriorly, the truncation margined ; a median raised line reaches 
right through to the truncation, on each side of this are two ler lines, which do not 
attain the truncation, and between all these lines the surface is ^ igreened ; external to 
these lines, between the outer ones and margined sides of the pr,podeum the surface is 
obliquely rugulose. There is a well-developed tooth on the underside of the claw of 
the anterior tarsi near its middle. Wings hyaline, neuration yellow, stigma divided in 
the middle by a white space, so as to form a prostigma, at least not much less developed 
than in Sierola. Abdomen smooth and shining, the apices of the segments except the 
first pale. Length, ? 4*5 mm. 

Hab. Oahu ; Honolulu. 

Obs. This species seems to me certainly congeneric with Epyris hawaiiensis 
Ashm., which Kieffer refers to his genus Holepyris. E. haivaiiensis is a domestic 
species, common in houses, and rarely seen outside. It is certainly introduced, as also 
is H. hospes now described. 



6i6 FAUNA HAWAIIENSIS 

Fam. COSILIDAE. 

SiEROLOMORPHA Ashm. 

(i) Sierolojnorpha kospes, sp. nov. 

Female black, smooth and shining, legs, except the coxae, which are more or less 
dark, tip of abdomen, apex of clypeus, second and third antennal joints all pale-coloured, 
yellowish brown or testaceous ; scape of antennae dark brown but paler at base and 
apex, antennae except the above-mentioned joints sordid, but more or less reddish- 
tinged ; wings infuscate, stigma fuscous. 

Head smooth and shining, very remotely and sparsely punctured, the ocelli in a 
triangle widest at base, antennae 12-jointed, scape rather short, the pedicel subovate 
and subequal to the first funicle joint, which is decidedly shorter than the second, apical 
joint much longer than the preceding one. Thorax very sparsely punctate and sparingly 
pilose, smooth and shining except the impressed front of the pronotum, parapsidal 
furrows subconvergent posteriorly, not reaching the front margin of mesonotum ; 
scutellum with a deeply impressed transverse line in front ; propodeum with a transverse 
costate sulcature in front and a median rough triangular area, the rest smooth. Abdomen 
smooth and shining, very sparsely pilose and punctate. Wings with only one cubital 
and two discoidal cells bounded by true nervures, the second recurrent nervure and the 
second of the transverse cubiti indicated only by white lines, the first transverse cubitus 
also incomplete above. Length 4 mm. 

Ashmead figures the wing of the type of this genus. Bull. U.S. Nat. Mus. xlv. 
PI. IV. fig. 2, as Sierola Cam. ? Kieffer assigns Sieroloniorpha to the Bethylidae, as 
Ashmead did originally, removing it later to the Cosilidae. 

Hab. Oahu, Honolulu, 1906. 



PROCTOTRUPOIDEA. 

Fam. CERAPHRONIDAE. 

Subfam. CERAPHRONINAE. 

Ceraphron Jur. 

( I ) Ceraphron plebems, sp. nov. 

Black, the scape of the antennae and the legs except the coxae brownish yellow or 
testaceous, the front femora more or less darker, brown. 

Head and thorax very densely finely punctured or shagreened ; pedicel of the 
antennae elongate, longer than the next joint, which is itself elongate and longer than 
the following one ; second, third and fourth funicle joints small and short, the following 
ones a good deal larger than these and subequal, and not evidently transverse ; scutel- 



HYMENOPTERA 617 

lum distinctly shining and smoother posteriorly; posterior lower angles of the propodeum 
spinose. Wings hyaline and lightly infuscate, radius given off from the marginal vein 
before the apical third of the length of the wing. Second abdominal segment longi- 
tudinally costate on about its basal third, for the rest polished and impunctate. Length, 

% I — 1-5 mm. 

Hab. Oahu ; Honolulu. 



(2) Ceraphron abnormis, sp. nov. 

Ferruginous, the apex of the scape of the antennae black, as well as all the other 
joints, excepting that the pedicel may be somewhat pale. Abdomen with a dark 
transverse basal band, and the apical segments also black or dark, this colour beginning 
usually about the middle of the large second segment. 

Head dull densely sculptured, shagreened, scape of antennae attenuate towards 
the apex, pedicel in the Z very small, the first flagellar joint about three times as long 
as wide, a little longer than the following, which are subequal ; pedicel in the % elongate, 
rather more than half as long as the very elongate first funicle joint, which is double 
the length of the second funicle joint ; subapical joints of the funicle rather longer than 
wide. Thorax densely punctate, or shagreened, the scutellum smooth and polished 
posteriorly, the metanotum prominently angulate in the middle. Abdomen with long 
regular longitudinal costae at the base, for the rest smooth and shining, glabrous, the 
head and thorax bearing a very short pubescence. Length about 2 mm. 

It is doubtful whether this insect is a true Ceraphron, as it has the parapsidal 
furrows distinct, at least in front. According to Ashmead's definition of the group it 
differs from all Ceraphroninae in that the maxillary palpi are three-jointed, and there 
are two distinct spurs on the hind tibiae. The generic characters are as follows : 

Head transverse ; ocelli in a small subequilateral triangle, the lateral ones very 
remote from the eye-margins, the eyes pubescent, not nearly reaching the mandibles. 
Mandibles bidentate, maxillary palpi with three elongate and subequal joints, the first 
broadest basally, the second near the middle and the third subtriangularly dilated. 
Labial palpi very short, with an elongate pilose joint and probably a short basal one 
before this. Antennae oi $ long, filiform, 11 -jointed, subattenuate apically; in the % 
lo-jointed and with subclavate flagellum. Mesonotum with the parapsidal furrows 
marked by punctate lines, which usually fail before reaching the hind margins ; medio- 
dorsal grooved line complete and distinct. Axillae triangular, large, meeting inwardly, 
marked off from the scutellum, which is acutangular in front, by crenate lines. Posterior 
tibiae with two distinct calcaria. Marginal vein linear, less than one-third of the length 
of the long curved radius, separated by a pale spot from the submarginal. Abdomen 
with the second segment forming most of its surface, and costate at the base. 



6i8 FAUNA HAWAIIENSIS 

Hab. Oahu and one or more of the other islands; introduced from North America, 
as I have collected it in California. Swezey discovered it to be a parasite on Dryinids, 
Haplogonatopus and Echthrodelphax. 

Fam. SCELIONIDAE. 

Subfam. TELENOMINAE. 

(i) Teleiiomits despiciendus, sp. nov. 

Black, the scape dark brown, pale beneath, and also above at the base, the legs 
yellowish or testaceous, the tibiae and femora, seen from above, sometimes more or less 
darker or sordid, the pedicel of the antennae more or less brown. Head and meso- 
notum not much shining, closely and finely sculptured or punctate, the face polished, 
with a few microscopic punctures. Antennae with the pedicel obconical and much 
longer than the small third joint, three following joints all small and short and not 
differing much from one another, the seventh joint is a good deal larger than these, but 
still is much smaller than the eighth, which with the three following" may be said to 
form the club. Scutellum smooth and shining, lower lateral angles of the propodeum 
not prominent. Basal abdominal segment and the second at the base regularly longi- 
tudinally rugose, the latter segment much longer than its basal width and except at the 
base polished. Front wings with short apical cilia. Length, $ "6 mm. 

Hab. Oahu ; Honolulu. 

Telenonius rhopali, sp. nov. 

Closely allied to T. despiciendus, the structure of the antennae being in most 
respects similar. 

Black, with black antennae and dark legs, which are black or in part dark brown ; 
knee-joint of front legs, apex of tibiae more or less, and the base of tarsi paler ; extreme 
base of middle and hind tibiae, and the basal joint or two joints of their tarsi yellowish. 
Antennae with the pedicel about as long as the two following short joints together ; the 
seventh joint is transverse, and considerably larger than the very small sixth joint, but 
much smaller than the eighth, which with the three next forms a four-jointed club. 
Sculpture of abdomen as in T. despiciendus. Length, % '6 mm. 

The male closely resembles the female, excepting that the antennae are not clavate, 
the pedicel is not longer than the third joint, the latter and the two following being 
longer than the sixth and following joints, which are short and subquadrate, the apical 
joint pointed and much longer than the penultimate. Legs sometimes paler than in the 
?, dark sordid brown or pitchy, and the tarsi etc. more clearly and extensively yellow. 

Hab. Oahu ; parasitic in eggs of Rhopaltis hyalinus, but in no wise related to the 
other species of Telenonius infesting these eggs. 



HYMENOP TERA 6 1 9 

(3) Telenonms paractias, sp. nov. 

Black, the scape of the antennae and legs clear testaceous, the tip of the scape and 
of the tarsi infuscate, pedicel a little pallid, at least apically. Wings lighdy infuscate, 
neuration rather pallid, the fringe of front wings moderately short. 

Head polished in front, on the vertex dull, densely and minutely punctured ; pedicel 
of the antennae obconical, rather shorter than the slender and elongate third joint, fourth 
rather similar to the third, subequal to it or slightly shorter, fifth much shorter than the 
fourth and wider, sixth and following short, subquadrate, wider than the fifth, so as to 
form a fairly distinct six-jointed club, the apical joint being smaller than the preceding 
and pointed. Mesonotum finely punctured, sparsely pilose, lower posterior angles of 
the propodeum prominent. Basal abdominal segment regularly longitudinally rugose, 
the second with short regular wrinkles at the base, and forming the greater part of the 
dorsal surface, much longer than its basal width, smooth and shinine. Leneth, 2 i mm. 

H.\B. Honolulu ; parasitic in eggs of Rhopalns hyalimis. 

(4) Telenonius adelphus, sp. nov. 

Extremely like T. paractias and only distinguishable so far as I can see by the 
more widely darkened apex of the scape of the antennae, the more confusedly rugose 
basal abdominal segment and chiefly by the antennal structure. Though the length of 
the antennal joints is nearly similar in either species, in T. adelphus the club is only 
5-jointed, the sixth joint of the antennae being not very different from the fifth, while 
the seventh is much wider than the former. 

The male of this species is known to me, and in general resembles the female, but 
the scape is dark pitchy except at the base and the hind and middle tibiae are more or 
less darker, the former at least being somewhat infuscate above. Antennae filiform, 
pubescent, 12 -jointed, the scape a good deal shorter than the elongate third joint, the 
latter slightly shorter than the fourth, the fifth also shorter than the latter, the following 
joints tending to decrease in length to the apical one, which is longer than the 
preceding and pointed. 

Hab. Oahu, 1500 — 2000 ft. 

(5) Telcnornits vulcanus, sp. nov. 

Black, the tarsi brown or obscure, wings hyaline, iridescent, subinfuscate, clearer at 
base. The head is not very much wider than the thorax, and both are sparsely clothed 
with white hairs and are punctate. The sculpture of the mesonotum under high powers 
of the microscope is seen to consist of .scattered punctures, and of very delicate reticulate 
lines, which are not close enough to prevent the surface from being shining ; under 
F. H. II. So 



620 FAUNA HAWAIIENSIS 

similar powers the rugosity of the basal abdominal segment and the transverse costate 
impression at the base of the second can easily be seen, but under a strong lens the 
sculpture is seen with difficulty. 

The antennal club may be said to be four-jointed, since the seventh joint of the 
antennae is notably smaller than the eighth, though likewise notably larger than the 
sixth ; the pedicel is ovate or obconical, and much wider and also longer than the third 
antennal joint, which is elongate and about twice as long as wide, the third does not 
differ much in length from the second, the penultimate joint is about as long as its 
greatest width. The cilia of the front wings are rather long, the longest being about 
half the length of the width of the wings, where widest. Abdomen narrow, elongate 
and subparallel-sided. Length % '8 mm. 

The male has filiform-moniliform antennae, the pedicel and first funicle joint about 
equal, the latter, in one aspect, more than usually wide, with the sides rounded, subequal 
in length to the next, which is narrower, the apical joint pointed, and one and a half 
times as long as the preceding. 

This species is remarkable for its narrow elongate form. 

Hab. Hawaii ; Kilauea. 

Subfam. BAEINAE. 
The two following species do not agree with the subfamily characters, as given by 
Ashmead, since they possess a quite distinct postmarginal vein. They are easily 
separated from each other as follows : 

1 (2). Abdomen petiolate, posterior ocelli close to the eye-margins Pseudobaeus. 

2 (i). Abdomen sessile, posterior ocelli very remote from the eye-margins Dyscritobaeus. 

These insects are certainly introduced ; they are found in gardens in Honolulu, in 
company, usually, with introduced species of bugs. I suspect they are egg-parasites of 
these, but theoretically they should be bred from eggs of spiders. 

In Dyscritobaeus I have considered the segment, which is apparently the first and 
fits close to the propodeum as the second, as I believe a preceding segment exists, but 
is not readily observed. 

PsEUDOBAEUs, gen. nov. 
Head as wide or wider than the thorax, eyes pubescent, lateral ocelli close to the 
eye-margins. Antennae of the Z filiform, moniliform, 12-jointed; of % 7-jointed, with 
large solid club, several lines of hairs no doubt indicating the effaced articulations. 
Mesonotum with the parapsidal furrows indicated only posteriorly. Wings pubescent, 
shortly fringed, basal nervure present, but faint, marginal vein shorter than the stigmal, 
the latter oblique, but not extremely so, knobbed at the tip, rather shorter than the 
postmarginal, which bears three or four bristly long hairs, similar to those regularly 
disposed on the marginal and submarginal veins. Abdomen petiolate, the basal segment 



HYMENOPTERA 621 

much narrower than the second segment at apex or than the metanotum, second 
segment narrow at base and smaller than the third segment, which is the largest. 




Dyscritobaeus, front wing. 





Pseudobaeus, front wing and female antenna. 

( I ) Pseudobaeus peregrinus, sp. nov. 

Male : head and thorax dark brown or blackish, abdomen brown more or less 
suffused with black or very dark brown, the first segment always paler, yellowish brown; 
scape of antennae more or less yellowish brown, legs for the most part sordid. 

Female much lighter brown than the male, the thorax darker than the face, the 
abdomen with dark margins, scape in front concolorous with the face or nearly so, 
the flagellum more sordid, legs testaceous. 

Head punctured, the face smooth and shining, impunctate, mesothorax and 
scutellum closely punctured, clothed with short white hairs, lower posterior angles of 
propodeum prominent or spinose. First and second abdominal segments longitudinally 
striate, the third with excessively fine striae. Antennal scape of i about as long as the 
four following joints together, pedicel evidently smaller than the first funicle joint, 
which is longer than the following, the second to sixth being roundish, small and 
subequal ; seventh, eighth and ninth rather larger, the apical joint elongate, acuminate. 
Scape of female long, cylindrical, longer than the pedicel and funicle together, pedicel 
longer and larger than the first funicle joint, which is evidently longer than wide and 
about equal to the three following joints together, these being very short or transverse ; 
club ovate, equal to the scape in length. Length 75 mm. 

Hab. Oahu, Honolulu. 

Dyscritobaeus, gen. nov. 
Male : head fully as wide as the thorax, the eyes pubescent, the vertex concave 
behind, the ocelli in a triangle, widest at the base, but the outer ones are very remote 

80—2 



622 FAUNA HAWAIIENSIS 

from the eye-margins and often difficult to see. Antennae filiform 12-jointed. 
Mesonotum without furrows. Marginal vein of front wings shortish, shorter than the 
postmarginal, which is well developed, rather longer than the stigmal, the latter knobbed 
at the apex, very oblique or almost longitudinal in direction, originating about the middle 
of the length of the wing ; marginal, submarginal and postmarginal veins with regular 
series of bristles ; front wings shortly fringed, the cilia longest near the apex of the 
dorsal margin, basal nervure not defined. Abdomen very wide at base adapted to the 
propodeum, sessile, the second and third segments not differing greatly, or the latter 
rather longer. 

( I ) Dyscritobaeus comitaits, sp. no v. 

Male black, with whitish pubescence, the scape of the antennae in front, the legs, 
the whole of the second abdominal segment, the third at least on its basal portion 
yellowish brown or testaceous. 

Head dull, appearing very densely punctate or shagreened, antennae with tlie 
scape extended about as long as the whole insect, the pedicel subtriangular, about as 
long as the first funicle joint, the latter and all the following not differing much in 
appearance, appearing slightly longer than wide, apical joint acuminate and longest. 
Thorax dull, closely punctate (under high powers of the microscope the sculpture is 
seen to consist of punctures and a reticulation of fine lines) metathorax a little angulate 
in the middle. Second and third abdominal segments longitudinally striate, the former 
more strongly and distinctly than the latter. Length 75 mm. 

Hab. Oahu, Honolulu ; in company with the preceding, which it very closely 
resembles superficially. 

Baeus Hal. 
( I ) Baeus persordidus, F.p. nov. 

Dark pitchy brown, the head almost black, the abdomen rather paler than the 
thorax, legs infuscate or sordid yellow, paler than the body, hind tarsi yellow above, 
the tips dark. 

Head much wider than the thorax, somewhat shining and smooth, very finely and 
not closely punctured ; thorax and abdomen finely punctured and clothed with short 
white hairs. Hind tibiae long, but not twice as long as the tarsi, basal joint of the 
latter about equal to the three following together. Abdomen very strongly decurved, 
in dorsal aspect the second segment forms nearly the whole visible surface. 

The antennae have an elongate ovate club, rather longer than all the funicle joints 
and the pedicel together ; the latter is nearly as long as all the funicle joints, which are 
very short, the first being the longest. Length (!?) "5 mm. 

Hab. Oahu ; about 2000 ft. 



HYMEN OP TERA 



623 



Subfam. SCELIONINAE. 

Opisthacantha Ashm. 

(i) Opisthacantha dubiosa, sp. no v. 

Black, the mandibles, scape of the antennae wholly or in part and all the legs 
yellowish brown or testaceous, abdomen often distinctly brownish on the basal portion, 
rarely black. Head and thorax with white pubescence, both very densely and roughly 
sculptured, but the head more finely than the thorax ; postscutellum in profile appearing 
as a short spine projecting over the propodeum. First segment of the abdomen short, 
strongly transverse, much shorter than the second, both of these strongly longitudinally 
striate, third as long as the two preceding together, and much more finely longitudinally 
striate, the following segments excessively finely punctured. 

Antennae of the male filiform moniliform, the pedicel extremely small, the first 
funicle joint decidedly longer than those following, which are short, but not transverse, 
and do not differ much from one another, apical joint longer than the preceding and 
pointed. 

Antennae of female with a large six- or seven-jointed club, as long or longer than 
the pedicel and funicle joints together, pedicel about as long as the first funicle joint, 
which is longer than the following. The basal of the seven joints, which appear to me 
to form the club, may really belong to the funicle. Length about 2 mm. 

Differs from the type of Opisthacantha in the total absence of parapsidal furrows and 
the want of a basal cell. Ashmead appears to have seen specimens agreeing in these 
respects with the above (vide his classifications). Although he describes the third 
abdominal segment as "a little longer" than the second, he figures it as being much 
longer. 

Hab. Oahu ; males very common in company with introduced bugs. 

Anteris Forst. 
(i) Anteris montana, sp. nov. 

Female : black, the abdomen for the most part, the funicle of the antennae dark 
fuscous or pitchy, scape of antennae and legs testaceous, the pedicel of the former and 
coxae of the latter being darker, or more brown ; scutellum with the margin testaceous. 

Head pubescent, densely punctate ; first funicle joint of antennae much longer than 
the pedicel, and rather longer than the second, which like the third is elongate ; the 
latter about as long as the next two together, the fourth being rather longer than the 
fifth and not transverse ; basal joint of the club wider than long and much less wide 
than the next joint. Mesonotum and scutellum dull and very finely punctured, the 
postscutellum prominent and angulate in the middle posteriorly, propodeum rugulose- 
punctate, its inferior lateral angles minutely dentate, the superior ones slightly prominent. 



624 FAUNA HAWAIIENSIS 

Wings slightly yellowish tinged, the neuration yellowish, the stigmal vein paler. 
Abdomen with the basal segment regularly strongly striate, the second much more 
finely striate and crenate along the basal margin, the third very finely rugulosely 
punctate or subaciculate, the following very minutely punctured and hardly shining. 
Length 275 mm. 

The male, which no doubt belongs to this female, agrees very closely with it in 
sculpture, but is generally more elongate. The antennae are filiform, becoming 
attenuate apically, the pedicel is very small, the funicle joints are subequal in length, 
mostly about twice as long as wide, the apical joint much longer than the preceding and 
pointed. Hind coxae and hind and middle tibiae dark brown or dark fuscous, much 
darker than in the female. Allied to A. perkinsi Ashm. but with different sculpture of 
the abdomen and slightly different in other points. 

Hab. Oahu ; in the mountains, the male commoner than the female. 

(2) Anteris oahuensis. sp. nov. 

Female : head black, the mesonotum ferruginous, scutellum, except the margin, 
postscutellum and propodeum more or less darkly infuscate, abdomen obscure dark 
brown. Antennae testaceous, the funicle more sordid, the club nearly black. Legs 
yellow, tarsi appearing darker from the clothing of black hairs. 

Head somewhat coarsely sculptured, punctate and transversely rugose, funicle 
joints of antennae slender, the first three not differing much in length, and each several 
times as long as wide, fourth joint much shorter than the third, but considerably longer 
than wide, basal joint of club not at all strongly transverse, and only a little narrower than 
the following. Mesonotum and scutellum very finely punctured, postscutellum longitu- 
dinally rugose, with rounded well-raised posterior margin, not at all angulate ; propodeum 
emarginate posteriorly, its hind margin raised. Basal segment of the abdomen striate 
and with a smoother median elevation on its basal half, second more finely striate, third 
and fourth still more finely so, fifth and sixth very finely punctured. Wings lightly 
infuscate, veins yellow. Length 3 -5 mm. 

Hab. Oahu; near Honolulu, 1800ft. 

Caloteleia Westw. 

(i) Caloteleia elegans, sp. nov. 

Ferruginous, the funicle joints of the antennae, the scape, legs, base of second and 
of the terminal abdominal segment paler in colour, more yellow, but these parts them- 
selves are not uniform in tint. Five apical joints of the antennae, third, fourth and fifth 
abdominal segments, most of the second and the apical part of the sixth, black. Pedicel 
of the antennae and basal club joint more or less infuscate, but not black, apex of 



HYMENOPTERA 625 

posterior femora also dark. Head well rounded in front and emarglnate behind, the 
eyes pubescent, not nearly attaining the mandibles. Scape of antennae long, cylindrical, 
pedicel small, nearly equal to the first funicle joint, which is longer than the second, the 
latter about as long as its greatest width, club with six joints, transverse except the 
apical one, the basal one pale ; mandibles bidentate. 

Head and thorax quite dull, with dense, excessively minute sculpture, propodeum 
excavated and carinated on each side of the excavation. Front wings blackish smoky, 
with two transverse white bands, one considerably before and the other just beyond the 
marginal vein, the area between these pale bands is the darkest part of the wing. 
The wings are very narrow (petiolate) on less than the basal half marginal vein short, 
thickish, the stigmal knobbed at the tip, the postmarginal difficult to see, but longer 
than the stigmal. In some aspects a basal cell can be traced, but the nervures are very 
faint. Parapsides of the mesonotum evident. Basal abdominal segment longer than 
wide, its sides subparallel, the large suberect, shagreened basal process, attaining the 
postscutellum ; second segment elongate narrow at base, longer than the third, longi- 
tudinally striate. Third segment, wider than long, much longer than fourth, smooth, 
polished and impunctate, sixth only half the width of the base of the fifth, about as 
long as this and the fourth together, pointed at apex. Female: length 2-25 mm. 
Remarkable for the petiolate wings. 

Hab. Oahu, Honolulu ; certainly introduced. 



(2) Caloteleia exit I, sp. nov. 

Black, the mandibles and scape of the antennae, except more or less of the tips of 
each of these, testaceous or ferruginous, pedicel also more or less pale, legs testaceous. 

Head and thorax very densely and finely sculptured, dull, the mesonotal parapsides 
distinct. Pedicel of antennae about as long as the first funicle joint which is longer 
than either of the three following short joints, club six-jointed. Wings hyaline faintly 
clouded, neuration dark, basal cell faintly indicated, postmarginal vein longer than 
stigmal. Propodeum concave and carinate on each side. First abdominal segment 
dull, shagreened, the process, fitting the propodeal concavity, smoother, the extreme 
apex of the segment smooth ; second segment crenate at base and with some delicate 
striae reaching to about the middle of the length of the segment on its disc, otherwise 
smooth and shining, like the third, to which it is about equal in length, or rather longer; 
apical segment very narrow subtriangular, longer than the preceding. Female : length 
2"5 mm. 

Hab. Honolulu; no doubt introduced. 



626 FAUNA HAWAIIENSIS 

Fam. PLATYGASTERIDAE. 

Inostemma Hal. 
(i) InostetmJia almormis, sp. nov. 

Black, the tegulae brown, the abdomen dark brown, legs flavotestaceous, coxae 
browner. Wings hyaline, pubescent. Head dull, lateral ocelli extremely close to the 
eye-margins. Mesonotum dull, with dense minute sculpture, under high powers of 
the microscope showing a reticulation of fine lines ; posteriorly between the parapsidal 
furrows, which are distinct, the sculpture is less fine, and there are three short raised 
lines. Scutellum consute at the base, shagreened, propodeum with a spinose angle on 
each side. Basal abdominal segment rugose, its erect process reaching about to the 
dorsal surface of the thorax, but not extending over it, this process on its upper part 
densely covered with small pointed or spinose elevations ; second segment with a basal 
sculptured depression, longer than al! the following segments together and, except at 
the base, smooth and shining. Length i 6 mm. 

The single example I have seen is in poor condition and deprived of its antennae. 
Its generic position is uncertain, but clearly it is very near to Inostemma, even if it does 
not actually belong there. It is the only Platygasterid yet found in the islands, except- 
ing the very abnormal genus Aphanomerus, introduced by myself and Koebele to 
destroy the eggs of Siphanta. Aphanomerus pusillus'' is now thoroughly established. 

Hab. Oahu, near Honolulu. 

Fam. DIAPRIIDAE. 

Phaenopria Ashm. 
( I ) Phaenopria frater, sp. nov. 

Black, smooth and shining, impunctate, the scape and pedicel of the antennae red, 
the latter sometimes only red-tinged, all the legs ferruginous, the front tibiae and all 
the femora usually more or less obscured and darker than the hind tibiae and tarsi. 

Head with sparse pale hairs, the eyes small, very slightly convex, the ocelli in a 
nearly equilateral triangle. Antennae about two-thirds the length of the insect, the 
scape about equal to the following four joints together, the first flagellar joint distinctly 
longer than the second, longer but not so thick as the pedicel, the flagellar joints 
gradually widening to the tip of the antennae, but without a definite club, except that 
the apical joint is as long as the two preceding and wider than these. Pronotum rather 
densely hairy, the scutellum without a fovea in front, but with the axillae marked off, 
the propodeum conspicuously hairy towards the sides, and bearing a strong median 
carina, which seen in profile is bent into a distinct angle near the base. Petiole of the 

' Aphanomerus would probably be more correctly referred to the subfam. Baeinae, p. 620 ante. R. C. L. P. 



HYMENOPTERA 627 

abdomen sometimes reddish, with dense clothing like the propodeum, the abdomen 
itself elongate-ovate, the second segment about twice as long as all the following 
together. Wings with the thickened or punctiform marginal vein blackish fuscous. 
Hab. Oahu ; 1500 — 2000 ft. 

(2) Phaenopria soror, sp. no v. 

Very like P. frater, but differing in the colour of the antennae, the scape being 
almost black above, except at the tip ; in the darker colour of the legs, the clavate 
portion of the hind tibiae being darkly infuscate ; and in the more elevated and sharper 
basal angle of the median carina of the propodeum and the rather larger club joint of 
the antennae. 

Hab. Hawaii, Kilauea ; taken while ovipositing in larvae of a species of Droso- 
phila. 

(3) Phaenopria subtilis, sp. no v. 

Extremely like P. soror and P. frater and variable in colour. 

Black, polished, the legs and more or less of the antennae being red. Scape some- 
times wholly red or partly or all except the base and apex suffused with black or dark 
brown ; pedicel and flagellum sometimes wholly pale, the former and some of the basal 
joints of the latter always red beneath, though nearly the whole flagellum may be dark 
above. Legs sometimes to a considerable extent embrowned. 

Antennae with the pedicel thicker and as long or rather longer than the first 
funicle joint, the latter a little longer than the second, the funicle joints being mosdy 
small and longer than wide to about the sixth joint, the seventh, eighth and ninth are 
notably wider than the preceding, the two latter appearing in some aspects fully as 
wide as long or even somewhat transverse, the apical joint or club about as long as the 
three preceding together or at least not much shorter. Basal abdominal girdle of 
yellow hair very dense ; carina of propodeum in lateral aspect with its anterior angle 
not very prominent. Length I'S — 17 mm. 

Hab. Oahu; Koolau range, 1500 ft. and above. 

(4) Phaenopria ambulator, sp. nov. 

Female reddish brown or castaneous, the head and the abdomen except at the base 
black or nearly so ; the antennae black ; the scape and pedicel more or less red-tinged, 
the legs ferruginous or brownish yellow. 

Head with the sides slightly convergent behind the eyes basally, sparsely clothed 
with pale hairs, the eyes small ; ocelli absent ; antennae not much shorter than the 
whole insect, scape about as long as the four following joints, the pedicel and first 
flagellar joint nearly equal, the latter much longer than the second, the joints widening 
towards the apex of the antennae, there being no definite club ; the penultimate joint 

F. H. II. ^^ 



628 FAUNA HAWAIIENSIS 

subquadrate, about as long as its greatest width, the apical joint ovate, just as long as 
the two preceding together, but not quite twice as long as its greatest width. Head, 
thorax and abdomen smooth and polished, the head sparsely pilose ; the pronotum, 
propodeum and abdominal petiole with dense pale ochreous wool-like hair. Neck when 
exposed coarsely transversely rugose. Scutellum distinct, not impressed at base, the 
axillae not defined. Wings quite rudimentary, mere filaments, clothed like the propo- 
deum and easily overlooked. Abdomen ovate, pointed at apex, the second segment 
twice or more than twice the length of all the following together. 

Male closely resembles the female ; the antennae are longer, being as long as the 
whole insect, the first and second flagellar joints elongate, the second being distinctly 
longer than the pedicel, most of the following joints roundish and moniliform. 

Van a. Smaller, the scape and pedicel concolorous with the legs, the abdomen 
brown, black only apically and at the sides, the head also less black. 

Hab. Oahu ; 1 500 — 2000 ft. 

(5) Phaenopi-ia montana, sp. nov. 

Black, the legs rufotestaceous, the antennae red, becoming darker apically so that 
towards the tips they are black or almost so ; thorax sometimes dark brown or pitchy, 
a little rufescent in front. 

Smooth and shining, the antennae formed much as in P. ambulator, the third joint 
quite strongly elongate, decidedly longer than the pedicel. Posterior part of thorax 
very densely covered with appressed yellowish hair, the abdomen with a rather con- 
spicuous clothing of subdecumbent pubescence. When seen from in front it is notice- 
able that these abdominal hairs are more or less directed towards the middle line from 
each side. Lengfth 2 mm. Readilv disting-uished from P. ambulator bv colour and 
abdominal vestiture. 

Hab. Honolulu mountains, about 2000 ft. 

(6) Phaenop7'ia hylaea, sp. nov. 

Closel)^ allied to P. ambulator. Head, thorax, two basal joints of the antennae 
and the legs ferruginous or testaceous ; abdomen and the flagellum of the antennae 
black. 

Head smooth, flattish above, and thinly clothed like the thorax and abdomen with 
short erect hairs. Antennae much shorter than those of P. ambulator ; the pedicel is 
rather wider and longer than the following joint, which is subelongate, the fourth and 
following joints are small and nearly round, while those preceding the apical one are 
strongly transverse, the latter conic-ovate and shorter than the two preceding together. 
Thorax and abdomen smooth and shining, the hairy covering of the posterior part of 
the former yellowish, and less dense than that oi P. ambulator. Length r5 mm. 

Hab. Honolulu mountains, about 2000 ft. 



HYMENOPTERA 629 

DiAPRiA Latr. 
(i) Diapria drosophilae, sp. nov. 

Black, the thorax, abdominal pedicel and sometimes the second segment more or 
less red or dull red, in the ^ the thorax is sometimes nearly black or only slightly 
reddish tinged ; antennae in the $ red excepting the dark club, in the $ the two basal 
joints only are wholly yellow or red, but the basal stalk of the other joints is usually 
red, legs yellow with the apical joint of tarsi dark. 

Head smooth and shining, with sparse long hairs, and a patch of pale pubescence 
along the sides of the occipital margins. Antennae of the male very long, the pedicel 
not as long as the first funicle joint, which is much shorter than the following and lacks 
the long basal stalk, the second and following joints of the funicle are subequal, each 
with a very thin stalk, as long or longer than the thickened apical part of the joint, 
which bears a whorl of long hairs ; the apical funicle joints are noticeably shorter than 
the basal ones, the terminal joint being the smallest of all. Antennae of $ short, with 
well-marked three-jointed club, the pedicel much larger and longer than the first funicle 
joint, which with all the following joints is elongate or subelongate, the seventh funicle 
joint is noticeably wider than the preceding, but still much less wide than the basal 
joint of the club. Thorax polished and bearing sparse long hairs, the pronotum densely 
clothed with appressed hair ; scutellum foveated at the base, propodeum with a median 
carina forming a projecting angle anteriorly, in lateral aspect, pubescent on each side. 
Abdominal petiole short, about as long as wide in the ?, rather longer in the $, hairy 
and with appressed pubescence ; the rest of the abdomen smooth and polished. Wings 
subinfuscate, the cilia shortish. Length about 2 mm. 

Hab. Oahu, Honolulu; commonly parasitic on introduced species oi Drosophila. 



(2) Diapria xettica, sp. nov. 

Male black, the two basal antennal joints and all the legs yellow. 

Head globose, polished, clothed with sparse pale hairs ; antennae with the funicle 
moniliform and with whorls of long hairs, the pedicel very much shorter than the first 
funicle joint, which is subequal to the second, these two being the longest, but the latter 
is more incrassate, and in one aspect concave on one side, convex on the other, the 
following joints have at the most a very short stalk and are ovate, the several subapical 
joints being shorter than the basal ones and nearly round, the terminal joint somewhat 
shorter than the two preceding together. Thorax smooth and polished, sparsely clothed 
with erect hairs, the prothorax with a pubescent collar, the scutellum with a basal 
impression ; propodeum dull, with a median carina. Wings hyaline, pubescent, with 
rather short cilia, the stigma yellow. Abdominal pedicel hardly longer than wide, 

81—2 



630 FAUNA HAWAIIENSIS 

dull, hairy and pubescent, the rest of the abdomen smooth and polished. Length about 
I '5 mm. ^. 

Possibly this species should be referred to Ashmead's genus Ceratopria. 

Hab. Oahu, Honolulu ; no doubt introduced. 

CHALCIDOIDEA. 

The endemic Chalcids are chiefly, if not solely, represented by the Families 
Miscogasteridae, Encyrtidae, Eulophidae, and possibly a few species of Pteromalidae 
of the subf. Spalangiinae. In these families only a few genera are represented by 
native species, but there are a good many unquestionably introduced ones, represented 
each by only one or a few species. New forms are constantly appearing in Honolulu, 
having been accidentally introduced with plants from other countries, or in some cases 
purposely for economic reasons. 

Two or three species of Chalcididae have been recently introduced and one has 
now become a very common insect and will soon, no doubt, be ubiquitous. 

Fam. EURYTOMIDAE. 

EURYTOMA 111. 

(i) Euryionia sp. ? 

Hab. Oahu, Hawaii and probably all the islands. A recent introduction now 
common, parasitic on various Braconids. 

IsosoMA Walker. 

Isosoma spp. ? 

Two species of this genus occur in foreign grasses and are now widely distributed 
in the islands. 

Fam. ENCYRTIDAE. 
Subfam. EUPELMINAE. 
In addition to the species described below I know two or three other species of 
Eupelmines, all introduced and not yet determined. 

EuPELMUS Dalman. 

The species of this genus are numerous in the islands and I suspect that Ashmead 
included many species under the two names E. vtUgaris Ashm. and E. Jlavipes Cam. 
To these two species he assigned the whole of my numerous specimens, excepting nine 
individuals, of which five were named E. konae, the other four each representing a 



HYMENOP TERA 6 ; i 



J' 



distinct species. E. niger Ashm., xanthopus Ashm., and molokaiaisis seem all to be 
very easily distinguished and have not again been met with. E. hawaiiensis Ashm., 
and E. flavipes Cam. could hardly be recognized without inspection of the type, with 
any certainty, and the same might be said of E. vulgaris Ashm. I have made no 
attempt to include these in my tables, in fact it could not possibly be done, since 
characters of essential importance are not mentioned in the descriptions. As far as 
Ashmead's "table" is concerned, I may say that examples bred from such distinct 
hosts as bees of the genus Prosopis, cocoons of Chrysopidae, eggs of Orthoptera, etc. 
(each of the parasites, bred from these, forming a distinct species) would run down to 
E. flavipes Cam. 

One may doubt whether all the Hawaiian species rightly belong to one genus, but 
at present until the genera of Eupelmine Encyrtids are thoroughly revised I do not 
-^are to detach any from the genus Eupclnuis. The impossibility of the use of the hairs 
on the eyes for generic separation is obvious on examination of the Hawaiian forms, 
since all conditions between conspicuously hairy eyes, and those on which only a few 
hairs can be detected with difficulty, under high powers of the microscope, are to be 
found. 

Unfortunately the males cannot be associated with their females except by breeding, 
and they are likely to prove more difficult to separate than the latter. All the males 
I have bredi rhat belong to females with almost entirely yellow legs, have the legs dark. 
Ashmead, however, refers the males with dark legs to similarly coloured females, males 
with pale legs to females so coloured, and his determinations are certain to prove 
erroneous in many of these cases. 

Of course not too much stress can be laid on the metallic colouration of the head, 
thorax, etc. in determining species, yet in some cases this colouration is very fairly 
constant. The dichotomojs table, which I have made, not without considerable trouble, 
would probably fail anyone aLtempting to name isolated species, but will I believe prove 
to be of considerable assistance to the worker, after he has given a good deal of time to 
the preliminary study of a fairly representative collection. To see the characters 
properly it is necessary to relax and manipulate all the specimens, so that the parts are 
well exposed for examination. 

Synopsis of species (females). 

Wings smoky black, white for a short distance basally, scape of antennae 
distinctly dilated (apical joint of maxillary palpi dark) i. 

Wings uniformly hyaline, at most faintly yellow-tinged or infuscate ; scape 
simple 2. 

1. Hind tarsi with_ the two apical and (excepting the base and apex) the two 

basal joints dark above setiger. 

Hind tarsi with a band on the basal joint, and the apical joint black subseiiger. 

2. Palpi pale 3- 

Palpi dark i/ 



632 FAUNA HAWAIIENSIS 

3. Scape of antennae to a large extent and the mesopleura yellow E. rhyitcogoni. 

Scape darl^, yellow at most at the base above, mesopleura not yellow ...4. 

4. Facial impression above the interantennal elevation black, blue-black, or 

purple-black, more or less shining, at least in some aspects, finely 
shagpeened or quite smooth, never extremely dull and densely and 

evenly shagreened over its whole surface 5. 

Facial impression generally metallic, of various hues, but very rarely as 
above, the surface very dull, conspicuously and densely shagreened, 
generally shallow 11. 

5. Front and hind femora black or metallic green or blue-black, the apices 

only yellow 6. 

Front and hind femora largely or entirely pale or the hind femora only pale... 7. 

6. Head brilliantly shining E. xestias. 

Head not very polished, owing to minute surface sculpture E. axestias. 

7. Hind tarsi blackish or dark fuscous above 8. 

Hind tarsi pale 9. 

8. Front femora black, metallic E. ombrias. 

Front femora yellow E. parombrias. 

9. Ovipositor black and very short ......E. dysombrias. 

Ovipositor pale, except at base 10. 

10. Facial impression smooth and highly polished E. xestops. 

Facial impression only somewhat shining, being feebly shagreened E. paraxestips. 

11. Facial impression dark, blue-black or purple-black, and the sides of the 

face between the impression and the margins of the eyes also with 

purple or purple-red reflections E. axsstops. 

Facial impression of various metallic tints, as also the sides of the face 
adjoining it, rarely the impression itself is of a dark obscure tint and 
only slightly metallic 12. 

12. Scape with a clear yellow or brownish yellow band at the base 13. 

Scape without a yellow band at base 14. 

13. Abdomen (e.xcept basally) largely of a very dull red or rosy colcar and 

excessively densely shagreened, the surface being very dull E. rhodias. 

Abdomen nearly uniformly metallic and in some aspects shining E.xanthodorus. 

14. Very large species 5 mm. or more ; mesonotum extremely dril and densely 

sculptured, its colour for the most part dull metallic coppery, posterior 
femora dark and metallic outwardly in the lower j-art, the posterior 

tibiae infuscate outwardly • E. ei/prepes. 

Smaller species without some or any of the above characters 15. 

15. Front femora dark and metallic posteriorly except at the apex and more or 

less dark in front also at the base; eyes rather conspicuously hairy 

under a strong lens E. oribates. 

Front femora yellowish or brown, eyes very little hairy 16. 

16. Front tibiae yellow, not at all dark or metallic, mesonotum mostly vivid green ...E. oreias. 
Front tibiae metallic and usually distinctly darkened above ; mesonotum 

more golden or brassy green E. chrysopinus. 

17. Antennae short, and thick, basal funicle joints short not or only a little longer 

than wide 41. 

Antennae slender, several of the basal joints of funicle elongate and much 

longer than wide 1 7 (7. 

17a. Ovipositor hardly exserted behind the abdomen 43. 

Ovipositor well exserted ,. iS. 



HYMENOPTERA 633 

18. Facial impression blue-black or with purple leflections, always shining in 

certain aspects, often nearly smooth, more or less deep and narrow... 19. 
Facial impression rarely black or with purple reflections, usually green, 
golden or of other metallic tints, densely and evenly shagreened all over, 

quite dull, more or less wide <ind often shallow 26. 

19- Ovipositor black or dark fuscous above, at most a little pale apically 20. 

Ovipositor largely pale, being at least partly yellow or dull reddish 24. 

20. Front femora for a large pait or altogether yellow or yellowish brown 21. 

Front femora dark, pitchy and metallic, or metallic black, at most pale at the 

tips 23. 

21. Larger, legs except front and hind coxae clear brownish yellow, the hind 

femora a little metallic outwardly E. monas. 

Smaller, legs more or less darkened or sordid in various parts 22. 

22. Middle tibiae brown above, except at base and apex E. astheiies. 

Middle tibiae pallid yellowish throughout E. parastheties. 

23. Front tibiae above uniformly dark or pitchy, metallic E. kptophyas. 

Front tibiae black or metallic black and brightly yellow at the apex E. epilamprops. 

24. Facial impression above with a strong median carina E. dryas. 

Facial impression above with at most a faint trace of a carina 25. 

25. Legs brownish yellow or yellowish brown E. peles. 

Legs pallid yellow E. chloropus. 

26. Ovipositor as long as the abdomen E. euoplias. 

Ovipositor shorter than the abdomen 26a. 

26a. General colour of the front and hind femora and tibiae dark or black metallic 

green, the base and apex usually yellow ■ 27. 

General colour of these parts yellow or brownish yellow, at most somewhat 
darkened and metallic above or outwardly 34. 

27. Middle femora dark brown or pitchy above and beneath (excepting at the 

extreme base and at the apex) and hardly metallic ; ovipositor black, 

at most pale at the extreme tip E. pelopus. 

Middle femora pale beneath, usually more or less dark only on the posterior 
face ; ovipositor in some species pale to a considerable extent medially, 
yellow or obscure reddish 28. 

28. Middle tibiae almost uniformly pale and not, or hardly perceptibly, metallic 

above ; ovipositor black E. ntelanacrias. 

Middle tibiae more or less darkened and metallic above over a considerable 
portion of their surface, or if not very noticeably darkened, the ovipositor 
is more or less pale, at least dull reddish in part 29. 

29. Ovipositor black E. epimelas. 

Ovipositor pale, yellowish, reddish or obscure red in the middle 30. 

30. Abdominal pleura with wide dark bands, that are hardly metallic, separated 

by metallic bands E. cluilcoprepes. 

Abdominal pleura nearly uniformly metallic 31- 

31. Intermediate abdominal segments metallic blue or green with narrow, dark 

and hardly metallic apical margins E. caerulophanes. 

Intermediate abdominal segments above dark blackish fuscous, dull bluish 
black or purple-black, with bright metallic tints, at most, only at the 
extreme base 32. 

32. Head in front aspect between the front ocellus and the facial impression 

hardly shining or dull, densely sculptured ; mesonotal ridges red or rosy 
metallic 33 • 



634 FAUNA HAWAIIENSIS 

Head in front above the facial impression rather shining and smoother ; 

mesonotal ridges not reddish E. heterosomus. 

33. Smaller; marginal vein yellow, nearly uniform in colour and like the stigmal ; 

facial impression sordid metallic E. rhododorus. 

Larger ; marginal vein infuscate, much paler at the extreme base, darker than 

the stigmal ; facial impression green metallic — E. amaurndes. 

34. Scape of antennae yellow or brownish yellow basally %%. 

Scape not yellow at the base 35. 

35. Ovipositor very short about half the length of the hind tarsi E. achreiodes. 

Ovipositor more than half the length of the hind tarsi 36. 

36. Front tibiae at most a little browner and slightly metallic above E. aporostichus. 

Front tibiae with a dark conspicuous metallic line above 37. 

37. Middle tibiae with a dark metallic line above ; facial impression brightly 

metallic green or golden E. eustichus. 

Middle tibiae at most a little deeper brown above and not or hardly metallic ; 

facial impression of sordid metallic colour E. pelodes. 

38. Fully the basal half of the scape of antennae yellow E. kemixanthus. 

Less than half of the scape yellow 39. 

39. Front tibiae above yellow without a conspicuous dark metallic line ; ab- 

dominal pleura brightly metallic basally, dark apically E. pauroxanthits. 

Front tibiae above with a dark metallic line ; abdominal pleura not so 
coloured, very dull and densely sculptured 40. 

40. Mesothoracic pleura with distinct green metallic tint in part E. leucothrix. 

Mesothoracic pleura without distinct green tint E. paraleucothrix. 

41. Mesonotum medially fine metallic green, the ridges purple and strongly 

contrasted with the other E. basileius. 

Mesonotum without these strongly contrasted colours 42. 

42. Middle tibiae above dark and metallic except at the extreme base and at the 

apex; hind tarsi black or dark above, except at the base E. melanotarsus. 

Middle tibiae above only somewhat sordid in the middle ; hind tarsi mostly 

yellow E. xaiithoiarsiis. 

43. Hind femora and tibiae dark, the latter yellow apically ; middle legs entirely 

yellow, excepting the tips of the tarsi E. dysoplias. 



(i) Eupelnms setiger, sp. nov. 

Very dull, the head and thorax very densely shagreened and clothed with short, 
thick, white, subappressed setae. Head posteriorly with a greenish metallic tinge in 
front of this obscure coppery; mesonotum posteriorly dark fuscous, very little 
metallic, in front and externally to the ridges dull reddish, scutellum and axillae still 
more conspicuously red, mesopleura with a dull green tint in the middle. Abdomen 
shining metallic red at the extreme base, for the rest dull bluish or greenish black and 
densely sculptured, its sides with white setae like those of the thorax. Legs for the 
most part purplish black, the knee joints pale, basal joint of the front and middle tarsi 
pale at base and the former also at the apex, second and fourth joints of front tarsi 
brownish above, third yellow, fifth black ; third and fourth joints of middle tarsi yellow, 
hind tarsi like the front ones with third joint entirely yellow, the second and fourth 



HYMENOPTERA 635 

brown. Ovipositor white, black on more than its apical third, and also at the extreme 
base. 

Head with the facial impression shallow, dull, densely shagreened, not defined 
above ; eyes distinctly pubescent, scape of antennae dilated, the basal part of the funicle 
metallic green or golden green and densely clothed with appressed white hair, apical 
two or three joints of funicle and the club not so clothed, black and only slightly 
metallic. Wing smoky blackish, with a white band near the base, neuration sordid 
yellowish. Ovipositor short, about as long as the first two or three joints of the hind 
tarsi. Length % 3 mm. 

A robust species, very unlike any other, except the following, and probably not a 
true Etipeimus. 

Hab. Hawaii, Kilauea. 

(2) Enpelmiis subsetiger, sp. nov. 

Extremely like the preceding in colour, clothing and sculpture, but only about half 
the bulk, length 2 mm. Front tarsi with the basal and apical joints dark, the rest 
yellow, hind tarsi with the basal joint widely pale at base, and narrowly at the apex, the 
three following joints yellow, the apical joint dark, ovipositor black at extreme base, 
lightly infuscate apically. 

Hab. Oahu, on the coast. 

(3) Eupelniiis rhyncogoni P. 
Eupelmus rhyncogoni Perk., Proc. Hawaii Ent. Soc. i. p. 132. 
Hab. Oahu ; 1 200 ft. and upwards. 

(4) Enpelmtis xestias, sp. nov. 

Head above with brassy and red reflections, smooth and shining, with some feeble 
punctures ; facial impression deep purplish black, smooth and shining with a median 
carina above, palpi pale yellow or whitish. Antennae elongate, clavate, the first two 
funicle joints elongate and the longest, the third stouter than the second, but longer 
than wide, the apical joints transverse ; scape with bright metallic green reflection. 
Mesonotum golden, with redder reflections in the front and on the scapulae, very finely 
shao-reened and with scattered punctures. Front and hind coxae, front and hind 
femora, except at the apex, and half or more of the anterior tibiae black or dark, rest 
of the leo-s brownish yellow, or in parts yellow. Neuration yellow. Abdomen with 
brio-ht green metallic reflection at base, the rest dull, densely shagreened or sculptured, 
black or at the most faindy metallic. Ovipositor black, about as long as the first two 
or three joints of the hind tarsi. Length ? 2 mm. 

Hab. Hawaii ; 4000 ft. 

F. H. II. ^- 



636 FAUNA HAWAIIENSIS 

(5) Eupelntus axestias, sp. nov. 

Head above dull or only slightly shining, the sculpture very fine and indefinite, 
metallic green or yellowish green and with some coppery red reflection, facial impres- 
sion dark with purple reflections, smooth and shining, deep, its upper edge well margined, 
and with a median carina ; palpi whitish. Antennae subclavate, the pedicel much 
stouter and as long or longer than the first funicle joint, which with the next two is 
elongate, the last funicle joint being as wide or wider than long ; scape with metallic 
green reflection, eyes distinctly hairy. Front and hind coxae, front and hind femora, 
except apically, dark and metallic, the front tibiae also dark, but widely yellow apically, 
middle tibiae a little brown above near the base, the rest of the legs brownish. yellow or 
in parts still paler. Mesonotum with white pubescence, dull, bronzy, very finely and 
closely shagreened, and with scattered feeble punctures, externally to the ridges more 
coppery; abdominal pleura dull purplish black ; ovipositor black, about as long as the 
three first joints of the hind tarsi. Length % about 2 mm. 

Hab. Oahu, 1800 ft. 

(6) Evipelnius ombrias, sp. nov. 

Head black, with slight metallic green reflection, finely rugulose behind the ocelli, 
but otherwise very feebly sculptured, yet hardly shining. Paljai pale yellow ; facial 
impression very deep, smooth and shining, with a median carina, and its upper margin 
well defined, deep black or with slight purple lustre. Eyes very little pubescent. An- 
tennae of the usual form, subclavate, the pedicel and first flagellar joint subequal, the 
several basal joints of the funicle all being elongate, scape metallic green-black. Meso- 
notum more or less black or dark in front and at the sides, metallic with various hues, 
but not very brightly. Mesopleura blackish, slightly metallic. Abdomen green or 
blue-black above. Ovipositor missing, probably very short. Front and hind coxae 
dark, metallic, and also the front femora, rest of legs for the most part brownish yellow, 
at most a little darker in parts (e.g. the middle coxae basally); the hind tarsi con- 
spicuously dark. Length % 2 mm. 

Hab. Molokai ; 2500 ft. 

(7) Eupelinus parombrias, sp. nov. 

Head above slightly metallic, not shining ; the facial impression deep, black with 
slight purplish tinge, smooth and shining in some aspects, its upper margin well-defined, 
without a definite median carina. Palpi pale, scape of antennae largely brown or 
yellowish brown basally, darker and slightly metallic apically; the basal joints of the 
funicle elongate, the first being the thinnest and longest, the last funicle joint as wide or 
wider than long. Mesonotum greenish black and with some golden or brassy reflection, 
its pubescence dark and inconspicuous ; pleura of the thorax pitchy brown, dull and 



HYMENOPTERA 637 

metallic. Abdomen blackish, with green metallic reflection at extreme base. Legs 
yellow, the front coxae browner, the hind tarsi entirely brown or infuscate. Wings 
hyaline, neuration yellow. Ovipositor dark or pitchy, shorter than the hind tarsi. 
Length % about 2 mm. 

Has. Oahu ; Koolau range, 1500 ft. 

(8) Eupelmus dysombrias, sp, nov. 

Head finely punctured and somewhat shining and smooth, black, with metallic 
reflections behind the ocelli, blue-black in front of these, the facial impression deep, 
black or with slight purple tinge, viewed from the apex appearing nearly smooth, in 
front view very finely shagreened, its upper margin distinctly defined, no distinct 
median carina. Antennae with the basal joints of the funicle elongate ; eyes very 
sparsely hairy. Mesonotum brassy, but not very brightly metallic, densely shagreened, 
clothed with short and not very conspicuous, white hairs ; mesopleura for a large part 
dull coppery. Abdomen at the base and sides bright copper-coloured, in the depression 
blackish with coppery reflections. Front and hind coxae dark and metallic, middle 
coxae also dark beneath, rest of legs yellow, the front femora outwardly with metallic 
green reflection, but not darkened. Neuration yellow. Ovipositor black, short, about 
as long as the two basal joints of the hind tarsi. Length % rS mm. 

Hab. Oahu, Honolulu, not 100 ft. above sea-level. 

(9) Eupelmus xe stops, sp. nov. 

Head smooth, with a few fine punctures, polished ; facial impression deep, smooth, 
blue or purplish black, a trace of a median carina at its upper extremity, the margin of 
which is well defined. Eyes distinctly but rather sparsely hairy, palpi pale ; scape of 
antennae with metallic green reflection, basal funiclar joints elongate, the apical one 
about as long as wide. Mesonotum with white, short hairs, quite dull and densely 
shagreened, dull brassy; mesopleura in front and posteriorly more coppery. Front 
and hind coxae dark and more or less metallic, rest of legs brownish yellow or testaceous 
and not metallic. Abdomen brilliant coppery at the base, the rest of the surface and 
the pleura also conspicuously metallic, but less bright than the base. Ovipositor yellow 
or yellowish brown, black at the base, as long as the hind tarsi, or rather longer. 
Length % 3 mm. 

Hab. Hawaii, Kau, 4C00 ft. 

(10) Eupelmus paraxestops, sp. nov. 

Head dull, or at least only faintly shining in part, metallic with distinct green 
reflection in part at least, the surface finely rugulose and with some faint punctures ; 
facial impression deep, blue or purplish black, smooth, with indication of a median 

82—2 



638 FAUNA HAWAIIENSIS 

longitudinal carina, its upper margin distinctly defined. Palpi pale, antennae obscurely- 
brown basally, of the usual structure, eyes apparently with very few hairs or almost 
glabrous. Mesonotum metallic green, with brassy or golden reflections, mesopleura 
with a bluish space near the middle. Abdomen brilliantly metallic green at the base, 
its depressed portion dark bluish or purplish black, the pleura metallic green and 
coppery. Legs, e.xcept the dark and metallic front and hind coxae, entirely yellow or 
brownish yellow. Neuration yellow. Ovipositor yellow, black at the base, shorter 
than the hind tarsi. Length % about 3 mm. 
Hab. Oahu ; Koolau range, 1500 ft. 

(11) Etipeh7t2is axestops, sp. nov. 

Head brightly metallic above, brassy or golden with more or less red or coppery 
red reflections ; facial impression deep, blue- or purple-black, dull, distinctly and closely 
shagreened, its upper margin distinctly defined. Antennae slender and elongate, the 
four basal funiclar joints all long and slender, scape brightly metallic, usually more 
or less brownish basally. Palpi pale, eyes somewhat pubescent, but not very con- 
spicuously so. Mesonotum bright green, or brassy green, the scapulae metallic red or 
coppery red, the pubescence white, the surface dull with dense distinct shagreening ; 
pleura with red and coppery reflections. Abdomen at the extreme base bright green, 
behind this dull black or purplish black, the apical segments metallic, the pleura nearly 
uniformly coppery red. Front and hind coxae dark and metallic, rest of legs yellow or 
brownish yellow. Ovipositor black at the extreme base, the rest yellow or brownish 
yellow, nearly as long as the hind tarsi. Length $ 3 '5 — 4 mm. 

Hab. Oahu ; widely distributed at elevations of 1 200 ft. and upwards. 

(12) EiLpelmus rliodias, sp. nov. 

Head and thorax of an infuscate or sordid copper colour, extremely dull and very 
densely shagreened, clothed conspicuously with pale hairs. Facial excavation shallow, 
very densely shagreened and dull, not distinctly margined above ; antennae widely pale 
(yellowish brown) at the base, on the apical portion dark brown or fuscous ; palpi pale ; 
eyes conspicuously hairy ; first funicle joint of antennae longer than the pedicel, the 
second also elongate but shorter than the first ; mesopleura somewhat like the 
mesonotum in colour, but with dull green tints in part. Legs with the front and hind 
coxae dark and metallic, for the rest the legs are brown and yellowish, being darker in 
some parts than others, the apices or base and apices of the joints being usually more 
yellow. Neuration yellow. Abdomen very dull and densely shagreened, black and red 
or dull rose-coloured for the most part, at the extreme base brilliantly metallic, but even 
here with a quite distinct surface sculpture. Ovipositor sordid yellowish, black at the 
base, longish, as long or longer than the hind tarsi. Length ? 3"5 mm. 

Hab. Molokai, 2500 ft. 



HYMENOPTERA 639 

(13) Eupelmus xanthodorus, sp. nov. 

Head dull, rough with shallow punctures and surface sculpture, obscure coppery 
metallic. Facial impression not very deep, very dull, densely shagreened, with greenish 
tint in the antennal scrobes, its upper edge not distinctly margined, faint traces of a 
median carina, which is probably not a constant character. Eyes sparsely and incon- 
spicuously hairy ; scape of antennae clear yellowish at the base, three basal funicle 
joints all quite strongly elongate. Palpi pale. Mesonotum dull, roughly shagreened, 
copper-coloured, clothed with white hairs, mesopleura coppery, except the usual dull 
median area. Front and hind coxae dark and metallic, rest of legs yellow or brownish 
yellow, hind femora outwardly rather browner and with metallic reflection. Abdomen 
above metallic, most conspicuously so at base, the pleura nearly uniform coppery and 
shining in some aspects. Ovipositor sordid yellowish brown, black at base, and nearly 
white close to the tip, hardly as long as the hind tarsi. %. Length 4 mm. 

Hab. Hawaii, Kau ?. The locality of this species is a little uncertain and 
possibly it may be from Oahu. 

(14) Eitpebnus euprepes, sp. nov. 

Head of a dull copper colour, its surface extremely dull and densely shagreened, 
facial impression shallow, very densely shagreened, dull, its sculpture finer than on the 
sides of the face adjoining. Eyes with excessively short, inconspicuous hairs. Palpi 
pale. Mandibles except the tips and the pedicel at the base of the scape of the antennae 
yellowish, scape metallic, first funicle joint very long, longer than the pedicel or than 
the second joint, which with the next two is strongly elongate. Mesonotum dull copper- 
coloured, very densely shagreened, not the least shining, thoracic pleura to a large 
extent metallic green. Front and hind coxae dark and metallic, the rest of the legs for 
the most part yellow or brownish yellow, the front femora with a dark metallic line 
behind, the front tibiae largely dark and metallic, posterior femora dark and metallic 
outwardly on their lower part at least, the hind tibiae subinfuscate and with metallic 
reflection ; middle tibiae similarly somewhat sordid and metallic, except at the base and 
apex. Abdomen with brilliant green and golden reflections at the extreme base, behind 
this banded with blackish hardly metallic and dull greenish metallic transverse areas, 
pleura dull metallic-coloured (mostly green or coppery) with narrow dark apical bands. 
Ovipositor yellow, rather widely black at base, about one and a half times the length of 
the hind tarsi. Length 5-5 mm. ?. 

Hab. Hawaii, Kilauea. 

(15) Eupelmus oribates, sp. nov. 
Head above golden and green metallic, more or less sordid anteriorly, facial 
impression very dull, shagreened, moderately deep, its upper limits definite, but without 
a distinct margin, colour dull metallic green to a large extent. Palpi pale, antennae 



640 FAUNA HAWAIIENSrS 

without special characters, the several basal joints of the funicle elongate, the first the 
longest. Eyes distinctly hairy. Mesonotum roughly shagreened, golden metallic, the 
scapulae reddish coppery, very conspicuously clothed with white hairs. Front and hind 
coxae dark and metallic, legs otherwise mostly brownish yellow, the front femora are 
darkened at the base and have a dark metallic line beneath, the front tibiae are some- 
what embrowned and metallic. Posterior femora more or less darkened outwardly and 
with green metallic reflection. Abdomen brightly metallic at base, the rest blackish 
and hardly metallic, except the apex, which, as often, is again brightly metallic ; ab- 
dominal pleura coppery metallic. Ovipositor yellow, black at base, shorter than the 
hind tarsi, about equal to the first two or three joints. Length % about 3 mm. 

Hab. Oahu, Pali, Deer. 1900, no. 803. 

(16) Eupelnuis oreias, sp. nov. 

Head metallic green, the facial impression dark coloured, with slight bluish or 
purplish tint in some lights, densely shagreened, dull, rather deep ; the sides of face 
between the impression and the eyes metallic coppery or golden. Antennae normal, 
palpi pale, eyes hardly noticeably hairy. Mesonotum largely bright green, densely 
shagreened, scapulae more golden or golden green ; clothing of white hairs not very 
conspicuous, chiefly observable at the sides ; thoracic pleura less brightly metallic than 
the mesonotum, having dull green and golden reflections, abdominal pleura coppery. 
Front and hind coxae dark and metallic, legs otherwise entirely pale. Abdomen above 
metallic, dark banded. Ovipositor short, yellow, black at the base, about half the length 
of the hind tarsi. Length $ 275 mm. 

Hab. Oahu, Koolau range, 1500 ft. 

(17) Eiipelmus chrysopinus, sp. nov. 

Head metallic, green, golden green or golden, the facial impression dull metallic of 
various colours, sometimes dull greenish or purplish black, but always with blue or 
purple reflection about the interantennal elevation, the surface densely shagreened and 
dull. Palpi pale ; antennae without special characters, eyes very little hairy. Meso- 
notum dull or nearly so, brassy or golden green, the scapulae coppery or reddish, 
wholly or in part. Front and hind coxae dark and metallic, the rest of the legs pale, 
yellow or brownish yellow, the front tibiae above metallic and usually dark on this 
metallic line. Abdomen at the base brilliantly green, behind this dark purplish black 
and with more or less blue or green reflections, the apical segment bright metallic 
coppery, the pleura uniformly copper-coloured. Ovipositor yellow, black at the base, 
short, about half the length of the hind tarsi. Length % 3 — 3 '8 mrn. 

Hab. Hawaii, Kau at various elevations. 



HYMENOP TERA 64 1 

(18) Etipelnius monas, sp. nov. 

Head somewhat shining, golden with some green and coppery reflections, finely 
rugulose and with some sparse fine punctures, facial impression, purple or reddish purple, 
with the surface rugulose and not shining, with a more or less distinct median longi- 
tudinal carina. Antennae of the usual form, eyes with very short hairs, palpi dark. 
Mesonotum rather dull golden, densely shagreened, quite conspicuously clothed with 
white hairs, scapulae more or less infuscate. Neuration yellow. Front and hind coxae 
dark and metallic, the rest of the legs yellow, hind femora appearing a little dark above. 
Abdomen bright metallic coppery at extreme base and at the apex, the intermediate 
parts dull blackish, the pleura uniformly metallic coppery, ovipositor black or blackish 
fuscous, a little shorter than the hind tarsi, about equal to the three or four basal joints 
together. Length % 3 mm. 

On account of the dull rugulose facial impression, which is less narrow at the 
bottom than is usual in the species, in which the colour of this part is purple, and also 
because the impression is of more metallic tint than in the other species, it is doubtful 
whether this species should really be placed with those having a blue-black or purple- 
black face. 

Hab. Oahu, near Honolulu, 1500 ft. 

(19) Eupeltnus asthenes, sp. nov. 
Head shining, golden, more or less rugulose and with fine punctures, facial im- 
pression very deep, smooth, deep black or purplish black ; eyes scarcely hairy, antennae 
of the usual form. Palpi dark. Mesonotum dull, finely and closely shagreened, deeply 
depressed in the middle, dull brassy, in front and the scapulae coppery. Front and hind 
coxae black and metallic, general colour of rest of legs yellow or yellowish brown, the 
front femora behind darkened and metallic, front tibiae excepting the apex and extreme 
base dark and metallic above ; hind femora, hind and middle tibiae also more or less 
embrowned and with metallic reflection, the apex of the tibiae being quite pale. Base 
of abdomen brightly metallic green, behind this the segments are blackish and hardly 
metallic, the terminal segment bright coppery ; abdominal pleura also copper-coloured. 
Ovipositor black, about equal in length to the hind tarsi. Length $2-25 mm. 

Hab. Hawaii, Kilauea. 

(20) Eupelmus parasthenes, sp. nov. 
Head above somewhat shining, green or golden, the facial impression very deep, 
purple-black in colour, and sometimes with more or less evidence of a median carina, in 
some aspects the surface appearing nearly smooth. Palpi dark, eyes collapsed after 
death, probably very little hairy, antennae with no special characters, Mesonotum with 
conspicuous white pubescence, brassy with the front lobe and the scapulae more reddish 



642 FAUNA HAWAIIENSIS 

or copper-coloured. Front and hind coxae dark and metallic, rest of legs generally 
yellow, the front femora behind are somewhat darkened and metallic, the front tibiae a 
little sordid, the hind femora are embrowned outwardly and have a metallic reflection, 
the hind tibiae are also somewhat darkened above and slightly metallic, their apices and 
the base of the tarsi being white. Abdomen bright coppery at the base and tip, 
between these dark and hardly metallic, the pleura wholly copper-coloured. Ovipositor 
blackish, about as long as the hind tarsi. Length % 2*6 mm. 

Hab. Oahu, on Piptui'tis, no. 857. 



(21) Eiipelmus leptophyas, sp. nov. 

Head scarcely shining, finely and sparsely punctured, golden or golden green ; the 
facial impression deep, purple-black, smooth and shining, face between the impression 
and eye margins also black ; palpi dark, antennae of the usual form, eyes with sparse 
hairs. Mesonotum brassy, green tinged, finely shagreened, the scapulae more reddish 
golden or coppery, clothed with white hairs. Neuration yellow. Front and hind coxae, 
the front and hind femora (except at extreme base and apex) dark and metallic, front 
tibiae dark and metallic above (except at base and apex), hind tibiae at most slightly 
embrowned, rest of legs brownish yellow or yellow, except the tips of the tarsi. 
Abdomen brilliantly green at extreme ba.se, behind this dull blackish, the apex reddish 
coppery ; abdominal pleura fiery copper-coloured, the intermediate segments, when not 
contracted, green metallic basally. Ovipositor blackish or dark fuscous, paler at the 
extreme tip, long and slender, rather longer than the hind tarsi. Length $ 4*5 mm. 

The abdominal segments in this example were fully extended, so that some speci- 
mens might be considerably shorter. 

Hab. Hawaii (Mauna Loa), 4000 ft. 

(22) Eupelvius epilamprops, sp. nov. 

Head finely rugulose, in some aspects shining about the ocelli, and golden metallic, 
facial impression deep, purplish black, the surface nearly smooth seen from the apex, 
and with a more or less distinct median carina ; antennae of the usual form, the palpi 
dark, the eyes inconspicuously hairy. Mesonotum coppery in front and on the scapulae, 
posteriorly more golden, conspicuously clothed with white hairs. Front and hind coxae 
dark and metallic, front femora almost entirely dark brown or pitchy, metallic above and 
behind, front tibiae much the same as the femora, tarsi brown or fuscous above. 
Middle legs yellowish, the tibiae dark above except at the base and apex. Hind femora 
embrowned, more yellow apically, slightly metallic, hind tibiae dark or brown above 
except at the apex, the tarsi dark above, except the basal joint, which is mostly pale. 



HYMENOPTERA 643 

Neuration pale, yellowish. Abdomen brilliantly metallic at the extreme base and 
coppery at the tip, between these mostly dull blackish, the pleura coppery. Ovipositor 
slender, black, about equal in length to the hind tarsi. Length % 3 mm. 

Hab. Hawaii, Kau, 4000 ft. 



(23) Eupelnms dryas, sp. nov. 

Head in some aspects a little shining, finely rugulose and sparsely punctured, green 
or golden green, facial impression purple-black, smooth or nearly so at least in some 
aspects and with a median carina ; palpi dark, eyes bare or nearly so, antennae of the 
usual structure, with elongate basal funicular joints. Mesonotum in front and on the 
scapulae coppery, posteriorly more or less green, dull, densely shagreened, clothed with 
white pubescence. Neuration yellow. Front and hind coxae dark and metallic, all the 
rest of the legs brownish yellow, front femora with metallic tint posteriorly. Abdomen 
brilliant metallic green at base, the apex coppery, most of the intermediate segments 
dull black, or purplish black, pleura copper-coloured. Ovipositor reddish, longer than 
the hind tarsi. Length $375 mm. 

Van dubius, the legs paler yellow, the middle tibiae except at base and apex 
subinfuscate above, the front femora without metallic tint posteriorly, facial impression 
less smooth and with fine surface sculpture. 

Hab. Molokai, 3000 ft. ; var. dubius Oahu, 1500 ft. I have seen only a damaged 
specimen of the latter, which has lost its ovipositor ; it is probably a distinct species. 

(24) Eupelinus chloropits, sp. nov. 

Head above green and usually with some golden reflections, more or less shining 
anteriorly, in some aspects at least ; facial impression deep, somewhat rugulose or 
shagreened, but still appearing smooth and shining when viewed from the apex, usually 
with fine reddish purple reflections ; palpi dark, antennae normal, with several of the 
basal funicular joints elongate. Eyes usually collapsed in dry specimens, probably with 
very few hairs. Mesonotum metallic green, densely shagreened, the front lobe more 
golden and the scapulae with coppery red or fiery reflections, conspicuously clothed with 
white hairs. Neuration pale yellowish. Legs excepting the dark and metallic front 
and hind coxae almost entirely pale yellow. Abdomen brilliant metallic green at the 
extreme base, behind this dull blackish, the apex coppery ; pleura copper-coloured. 
Ovipositor black at base and more or less fuscous at the apex, the rest yellow, rather 
longer than the hind tarsi. Length $ 3— 4'25 "'"'"• 



& 



This species is unusually slender and elongate. 

Hab Oahu ; mountains near Honolulu, 1200 ft. and upwards. 

F. H. II. ^ 



644 FAUNA HAWAIIENSIS 

(25) Eupelmus peles, sp. no v. 

Head metallic, with coppery and green, or golden reflections, rugulose and sparsely 
punctured ; facial impression deep, wide, shagreened, but the antennal grooves are 
smoother and shining in some aspects, purplish black, as also the sides of the face between 
its impression and the eye margins. Palpi black, antennae of the usual form, eyes bare 
or nearly so. Mesonotum coppery or reddish coppery on the front lobe and on the 
scapulae, posteriorly dull golden or greenish, densely shagreened, the surface dull. 
Neuration yellow. Legs brownish yellow or yellowish brown, the front and hind coxae 
dark and metallic, the front femora above and the hind ones outwardly with slight 
metallic reflection, but not dark. Abdomen brilliantly golden green at the base, behind 
this for the most part blackish ; the bases of the intermediate segments, when exposed, 
are metallic, at least on each side, the pleura coppery. Ovipositor black at the base, 
the rest yellowish, decidedly longer than the hind tarsi. Length ? 4"2 5 mm. 

Hab. Hawaii, Kilauea. 

(26) Eitpelmus etioplias, sp. nov. 

Head above coppery or more golden in some lights, dull, rugulose and sparsely and 
finely punctured ; facial impression dull, densely shagreened, wide at the bottom, metallic 
like the sides of the face adjoining it, and more or less green about the antennal 
grooves, but somewhat less bright, antennae slender with elongate basal funicular joints 
as usual, palpi dark, eyes sparsely hairy. Mesonotum coppery or reddish metallic on 
the front lobe and on the ridges of the scapulae, dull and densely shagreened. Front 
and hind coxae, front and hind femora and tibiae dark and metallic, except the knee 
joints of the front legs and the apex of their tibiae, the knee joint of the hind legs which 
are more widely yellowish, the extreme base of the hind femora, and the apices of the 
hind tibiae ; middle tibiae distinctly embrowned and metallic above except at the base 
and apex, middle coxae also partly dark and metallic, tarsi brown or infuscate above, 
those of the middle legs less dark than the others, basal joint of hind tarsi whitish at 
base. Neuration yellowish, the stigmal vein postmarginal and apical part of the 
marginal vein more sordid or infuscate. Abdomen brightly metallic at the extreme 
base, behind this blackish, the ape.x coppery, as also the pleura. Ovipositor long, black, 
but on about its apical half it is sordid yellowish, about as long as the abdomen, when 
this is not contracted, interrupted a little before the basal fourth of its length. Length % 

375 mm- 

This species must be very close to E. konae Ashm., but difi"ers so much in detail, 
that I hesitate to consider it to be that species. 

Hab. Hawaii, Kilauea. 



HYMENOPTERA 645 

(27) Eiipelnius peloptis, sp. nov. 

Head green, in dorsal aspect dull, rugulose, with a few, nearly effaced, punctures, 
the facial impression wide, shallow, dull, densely rugulose, even in the antennal grooves, 
metallic greenish. Antennae of the usual form, eyes glabrous or nearly, palpi dark. 
Mesonotum dull, densely shagreened, green, the front lobe more golden, the ridges 
with coppery reflections. Front coxae, femora and tibiae almost wholly dark and 
brightly metallic, the trochanters black, slightly metallic, tarsi for the most part dark 
brownish above ; middle legs with the femora blackish, pale at base and apex, tibiae 
also darkish brown except at base and apex, tarsi mostly pale ; hind legs with the coxae 
and femora brilliant metallic green, the tip of the latter yellow, the tibiae dark fuscous 
and much less metallic than the femora, their base and apex pale, basal joint of tarsi 
pale, the rest dark above. Neuration yellow, appearing darker from the clothing of 
black hairs. Abdomen metallic green at the base, coppery at the tip. Ovipositor 
black, the tip pale, a little shorter than the hind tarsi. Length % ? 3-5 mm. 

Having seen but one example, which is in a much contracted state, I cannot give 
its length exactly. 

Hab. Hawaii, Kau district, about 4000 ft. 

(28) Eupelmus melanacrias, sp. nov. 

Head above golden in some aspects with green reflections, dull and rugulose, but 
seen obliquely somewhat shining; facial impression dull blackish, as also the sides of 
the head between it and the eye-margins, the surface quite dull, rugulose or shagreened ; 
palpi dark, antennae of the usual form, eyes with excessively short hairs, fairly numerous 
in parts, but very inconspicuous. Mesonotum dull and shagreened, the front lobe and 
the parapsides or scapulae coppery or reddish coppery, the rest with some greenish tint. 
Front coxae, femora and tibiae dark and metallic (at most pale at the apices) the 
trochanters also mostly dark and metallic, tarsi yellow, the apical joints dark above ; 
middle legs brownish yellow, hind legs with the coxae and femora dark and metallic, the 
latter yellow at the tips, tibiae dark except at base and apex and metallic, trochanters 
and tarsi, except the apical joints, yellow. Neuration yellow. Abdomen metallic blue 
and green above, the tip coppery ; ovipositor blackish, inclining to pitchy, about as long 
as the hind tarsi without their apical joint. Length % 3 mm. 

Hab. Molokai, 4000 ft. 

(29) Eupelmus chalcoprepes, sp. nov. 

Head green or golden green, the surface dull and roughened with rugulosity and 
some shagreening ; facial impression somewhat shallow, the surface dull, very densely 
shagreened or somewhat rugulose, the colour metallic, but not of vivid hues ; scape fine 

83—2 



646 FAUNA HAWAIIENSIS 

metallic green in some lights, the palpi dark, the eyes sparsely and inconspicuously 
pilose. Mesonotum with pallid yellowish or whitish hairs, green and more or less 
golden, the ridges sometimes coppery. Front and hind coxae, femora, and tibiae dark 
and metallic green, their apices and bases (excepting the coxae) more or less yellow, 
their trochanters also for the most part dark, all the joints of their tarsi more or less 
darkened above, the basal joint pale at the base ; middle legs yellowish, their coxae 
outwardly dark and metallic, the femora with a dark metallic line, the tibiae brown and 
metallic above except at the base and apex, the tarsi pale. Abdomen brilliant metallic 
green at the base, behind this the segments are dark, blackish, with basal metallic green 
or blue bands, tip of abdomen in apical view brilliant coppery, pleura blackish, with 
basal metallic bands ; ovipositor black at base, and darkened towards the apex, between 
these parts yellowish or reddish, about as long as the first two or three joints of the hind 
tarsi. Marginal and stigmal veins of front wings subinfuscate. Length % 4 — 4'5 mm. 

Hab. Hawaii, Kau district, 4000 ft. 



(30) Eupelmus epimelas, sp. nov. 

Head above dull golden, the surface minutely but roughly sculptured, dull and 
rugulose, facial impression rather shallow, greenish, densely shagreened or rugulose, dull, 
palpi dark, antennae of the usual form, eyes hairy, but not conspicuously so. Mesonotum 
greenish metallic, finely shagreened, dull, with sparse and fine but roughish punctures, 
the ridges with reddish reflections. Legs almost similar to those of E. chalcoprepes, 
but the middle coxae and the hind trochanters are yellow. Abdomen metallic green at 
the base, behind this purplish black, the tip green, seen from above, coppery in apical 
view ; pleura coppery. Ovipositor black, shorter than the hind tarsi, about equal to the 
three basal joints. Length % about 3 mm. 

Hab. Hawaii, Kilauea. 

(31) Eupelmus caeriilopJiantes, sp. nov. 

Head above metallic green or golden, densely finely rugulose and shagreened, so 
as to be rough and dull in dorsal aspect, facial impression shallow, densely rugulosely 
shagreened, the surface dull, with green and coppery metallic colour ; eyes incon- 
spicuously pilose, palpi dark, antennae of the usual form. Mesonotum with rather 
sparse white hairs, dull, densely shagreened, golden, the parapsidal ridges reddish 
coppery. Legs much as in E. chalcoprepes, but the basal joint of the hind tarsi is 
entirely yellow or whitish ; hind trochanters yellow, middle tibiae subinfuscate above, a 
little beyond the base, and metallic, the base and the apex more widely pale. Abdomen 
brilliant blue-green at the base, the intermediate segments dull metallic blue or green 
with dark fuscous apical bands, fifth segment entirely bluish metallic. Ovipositor black 



HYMENOPTERA 647 

at the base and apex, between pale, of an obscure yellowish brown colour, a good deal 
shorter than the hind tarsi, about as long as the two basal joints together. Length 
? 375 mm. 

Hab. Oahu, Waimea, no. 829. 

(32) Eupelmus heterosomus, sp. nov. 

Head above dull in one aspect, finely rugulose, in other positions considerably 
shining, with coppery and brassy reflections, facial impression wide but deep, entirely 
rugulosely shagreened, metallic with some purplish lustre ; antennae of the usual form ; 
eyes with a good many hairs, but they are very short and not conspicuous ; palpi dark. 
Mesonotum conspicuously white-haired, shagreened, dull, of a golden or brassy metallic 
colour, darker on the front lobe and on the parapsides. Legs almost as in E. caerulo- 
phantes, the basal joint of hind tarsi entirely yellow, the hind tibiae widely yellow at 
the apex, the middle femora entirely or almost entirely yellow or brownish yellow, the 
middle tibiae very dark and metallic above, except at the base and. apex. Abdomen 
brightly metallic at extreme base, the rest dull black or bluish black, the tip, viewed 
apically, bright copper colour ; pleura coppery ; ovipositor black at base and more or 
less dark at the apex, the rest reddish, shorter than the hind tarsi. Length $ about 
3 mm. 

Hab. Hawaii, Kau, 4000 ft. 

{■^l) Eiipelmus rhododoriis, sp. nov. 

Head above reddish golden, dull, roughly sculptured, the facial impression dull, 
densely rugulosely shagreened, of much duller metallic colour than the sides of the face 
along the margin of the eyes, the latter with a good many short hairs on their outer 
portion, the antennae of the usual form, the palpi dark. Mesonotum bright metallic 
o-reen, more golden on the anterior lobe, the parapsides metallic red or rosy. Front 
and hind legs much as in the several preceding species, middle coxae with bright 
metallic tint and darkened, middle femora with a metallic line, middle tibiae whitish 
yellow at base and apex, brownish and metallic, but not dark, between ; basal joint of 
hind tarsi yellowish white, like the apex of the tibia. Abdomen brilliant metallic green 
and o-olden at the base, the rest dull blackish, the tip, viewed apically, copper-coloured ; 
ovipositor black at base, beyond this sordid reddish or brown, not differing much from 
the hind tarsi in length. Length ? 3-5 mm. 

Hab. Oahu, near Honolulu, 1500 ft. 

(34) Eupelmus amaurodes, sp. nov. 
Head above green, more golden in front, dull with rough surface sculpture, and 
anteriorly with remote, largish, feebly impressed punctures ; facial impression very 
densely shagreened and dull, shallow on its upper part, green, the sides of the face 



648 FAUNA HAWAIIENSIS 

adjoining it golden or reddish golden ; antennae of the usual form, eyes evidently 
pilose, but the hairs extremely short, palpi dark. Mesonotum brightly metallic, green 
and golden, densely and rather strongly shagreened, the parapsides rosy red. 
Front and hind legs as in the preceding species, the basal joint of hind tarsi entirely 
pale, middle femora with a darkened metallic line, the tibiae dark brown above, except 
at base and apex, and metallic. Front wings with marginal, stigmal and postmarginal 
veins fuscous. Abdomen with bright green metallic colour at the base, behind this for 
the most part dull blackish, the tip in apical view coppery red, the pleura coppery; 
ovipositor black at base, beyond this brown, short, about as long as the two or three 
basal joints of the hind tarsi. Length % 4*5 mm. 

Hab. Hawaii, Kilauea. 



(35) E^ipelimts achreiodes, sp. nov. 

Head green and golden above, rugulose and with largish feeble or subobsolete 
punctures anteriorly, facial impression shallow and very wide, dull, densely shagreened, 
blackish green, the upper extremity of the interantennal elevation purplish, sides of face 
between the impression and the eyes metallic ; antennae of the usual structure, palpi 
dark, eyes very little hairy. Mesonotum densely shagreened, clothed with white hairs, 
green or golden green, the parapsides with reddish coppery reflections. Front and 
hind coxae dark and metallic, the rest of the legs yellow, front tibiae darkened above. 
Neuration yellow. Abdomen bright metallic green at the base, behind this mostly 
metallic green or blue, and duller, the tip and the pleura coppery. Ovipositor yellow, 
black at base, short, about half as long as the hind tarsi. Length $ 3'5 mm. 

Hab. Hawaii, Kau, about 2000 ft. 

(36) Eupelmus aporostickus, sp. nov. 

Head above green, rugulose, facial impression green, dull and rugulosely 
shagreened, wide, the sides of face adjoining it golden ; antennae of the usual form, 
palpi dark, eyes apparently glabrous. Mesonotum fine metallic green, less dark than 
that of the facial impression, the parapsides reddish golden. Front and hind coxae 
dark and metallic, the middle ones dark brown and metallic beneath, rest of le<ys 
yellow, the front tibiae a little darkened and metallic above. Abdomen brilliantly 
green at the base, behind this metallic green-black, the tip green above, coppery in 
apical view, pleura coppery. Neuration yellow. Ovipositor black at base, the rest 
yellowish brown and more or less subinfuscate apically, as long as the three basal joints 
of the hind tarsi. Length % 3 mm. 

Hab. Oahu, Honolulu mountains, on Pipturus, July 1900, no. 786. 



HYMENOPTERA 649 

{^^) Eupchims eustichus, sp. nov. 

Head green or golden and green, the surface rugulose, with shallow punctures in 
front near the eye margins, the facial impression densely rugulosely shagreened, and 
sometimes with a slight median raised line, dull and wide ; the antennae of the usual 
form, palpi dark, eyes glabrous or nearly so. Mesonotum green or golden green, the 
parapsides coppery or reddish coppery; front and hind coxae dark and metallic, front 
tibiae with dark metallic line above, as also the middle tibiae, front tarsi more or less 
brownish or lightly infuscate above, rest of the legs mostly yellow or brownish yellow. 
Neuration yellow. Abdomen brightly metallic green or blue at the extreme base, 
behind this mostly metallic blue-black or green-black, tip of abdomen in apical view 
coppery red, pleura coppery. Ovipositor black at base, the rest yellowish or reddish, 
rather shorter than the hind tarsi, or about equal to the basal four joints. Length % 
3-25—4 mm. 

Hab. Oahu, mountains near Honolulu, 1500 ft. or more. 

(38) Etipelnius pelodes, sp. nov. 

Head above more or less green, in front with reddish reflections, the surface quite 
rough, with fine wrinkles or shagreening and wrinkles together, the facial impression 
sordidly metallic, dull, densely rugulosely shagreened, sides of the face adjoining 
golden or coppery red ; antennae of the usual form, palpi dark, eyes with very few 
hairs. Mesonotum very dull, and densely shagreened, clothed with pale hairs, of green 
and dull golden colour, the parapsides dull reddish golden or coppery. Neuration 
yellow. Front and hind coxae dark and metallic, rest of legs mostly yellow or 
brownish yellow, front tibiae with a metallic dark line above, the middle one slightly 
embrowned above between the pale base and apex, apical two joints of posterior tarsi 
dark brown or blackish. Abdomen brilliant green at the base, behind this dull bluish 
black, not metallic, tip of abdomen in apical view bright coppery, pleura uniformly 
copper-coloured. Ovipositor yellow, black at the base, about equal in length to the 
hind tarsi. Length % 4 mm. 

Hab. Hawaii, Kilauea. 

(39) Enpelnins hemixantluis, sp. nov. 
Head above of a dull metallic colour, its surface densely finely shagreened and 
very dull, clothed with white hairs, the facial impression very shallow, with dull golden 
and green metallic tints, very densely shagreened all over and the surface quite dull ; 
eyes considerably hairy, antennae of the usual form, the palpi dark ; scape of antennae 
yellow on the basal half or more. Mesonotum very densely shagreened and dull, 
metallic dull golden and green, the parapsides redder, clothed conspicuously with white 
hairs. Neuration yellow. Front and hind coxae dark and metallic, the front and 



650 FAUNA HAWAIIENSIS 

middle tibiae with a dark metallic line above, the hind tibiae brownish except at the 
base and apically, rest of the legs yellow, including the tarsi, yellow, except more or 
less of the apices of these. Abdomen brilliantly metallic green at the base and reddish 
coppery at the tip, between mostly blackish ; pleura unusually dull with dense surface 
sculpture, and conspicuously clothed with white hairs, coppery; ovipositor yellow, black 
at base and more or less darkened at the apex, about as long as the three basal jomts 
of the hind tarsi. Length % about 3 mm. 
Hab. Oahu, near Honolulu, 1500 ft. 

(40) Eupelmus pauroxanthus, sp. nov. 

Head above green and golden, the surface rugulose, dull or very little shining, the 
facial impression, mostly dull metallic coppery, wide, very densely finely shagreened, 
dull ; antennae of ordinary form, the scape pale brown or yellowish for a short distance 
at the base, then becoming darker and metallic brown ; palpi dark, eyes sparsely and 
inconspicuously hairy. Mesonotum green with golden tints, dull, densely shagreened, 
the parapsides also for the most part green. Front and hind coxae dark and metallic, 
rest of legs yellow, the posterior femora outwardly slightly metallic, but not darkened, 
the posterior tarsi, except the white basal joint, lightly infuscate above. Abdomen 
brightly metallic green at the base, behind this the segments are brightly metallic 
basally, dull blackish apically, the pleura of the segments fiery metallic basally, dark 
apically ; ovipositor yellow, black at base, about as long as the first four or five tarsal 
joints. Length $ about 2 '5 mm. 

Hab. Oahu, Honolulu, 200 ft., July 1900, no. 769. 

(41) Eupdnuis leiuothrix, sp. nov. 
Head above green, finely and extremely densely shagreened, very dull ; facial 
impression wide, shallow, very dull and densely shagreened, green and golden ; antennae 
of the usual form, the scape yellow at the base, the rest mostly black and metallic ; 
palpi dark ; eyes considerably and noticeably hairy. Mesonotum dull golden, or 
green and golden, very densely shagreened, parapsides viewed from the sides more 
reddish. Front and hind coxae dark and metallic, front tibiae dark and brightly 
metallic above, the tarsi subinfuscate above ; middle tibiae dark brown above except at 
the base and apex ; hind femora with a darkened and brightly metallic area outwardly, 
the tibiae dark above except at base and apex, the basal tarsal joint white, the apical 
ones dark above ; the general colour of the legs is brownish yellow. Neuration yellow. 
Abdomen bright metallic green at base, behind this blackish, the tip in apical view fiery 
copper-coloured, the pleura dull with dense surface sculpture, clothed with white 
appressed hairs, coppery ; ovipositor yellow, black at base, and more or less dark at the 
apex, shorter than the hind tarsi. Length % 2 — 3 mm. 
Hab. Oahu, near Honolulu, 1500 ft. 



HYMENOP TERA 65 1 

(42) Etipclmns paraleticothrix, sp. nov. 

Head above more or less golden, densely minutely shagreened and dull, facial 
impression wide, not deep, very densely shagreened and dull, of rather dark metallic 
colour, antennae of the usual form, the scape yellow for a short distance at the base, the 
rest dark and metallic, eyes considerably hairy, palpi dark. Mesonotum very densely 
shagreened and clothed conspicuously with white hairs, deep or reddish golden behind 
and still more red on the parapsides, in front greenish golden, pleura with hardly any 
green tint. Legs nearly as in P. leucotkrix, all the tibiae with a dark line or darkened 
above, the front and hind ones conspicuously metallic on the dark line. Abdomen 
brilliant metallic at the base, and the tip, in apical view, reddish coppery, between 
for the most part dull bluish black, pleura coppery ; ovipositor yellow, black at base, 
as long as the first two or three joints of the hind tarsi. Length ? 3"5 mm. 

Hab, Hawaii, Kilauea. 



(43) Etipehnus basileius, sp. nov. 

Head above green punctate and more or less rugulose, facial impression deep, 
in some aspects nearly smooth at the bottom, black ; antennae short and stout, the 
several basal joints of the funicle not elongate, but short ; palpi dark, eyes apparently 
glabrous. Mesonotum green or blue-green, e.xtremely finely shagreened, somewhat 
shining, the parapsidal portion purple. Front and hind legs dark and metallic green, 
tip of the front tibiae and the tarsi mostly pale ; knee joints, and apical part of tibiae 
of hind legs yellow. Middle femora black and submetallic, also the tibiae except 
narrowly at the base and widely at the apex, which parts are yellow. Abdomen 
conspicuously metallic blue, brightly green at the base, pleura green ; ovipositor black, 
as long or nearly as long as the hind tarsi. Length % 2 mm. 

Hab. Oahu, Tantalus and elsewhere, 500 — 1500 ft. 



(44) Eupelmus xanthotarsus, sp. nov. 

Like E. basileius with short thick antennae, the head green, punctate or shagreened 
and with black face. 

Mesonotum entirely green, densely and evenly and more strongly shagreened and 
duller. Legs very similarly coloured, but the middle tibiae are less dark above, though 
embrowned, except at the base and apex. The hind tarsi yellow. Ovipositor black 
above, more or less yellowish tinged at the sides. Length % 2'\ mm. 

Hab. Oahu, near Honolulu, 1500 ft. 

F. H. II. 84 



652 FAUNA HAWAIIENSIS 

(45) Eupelmus melaiiotarstts, sp. nov. 

Very like E. xanthotarsus, but with the head above much less dull. 

Mesonotum densely and rather strongly shagreened, legs much as in the two 
preceding species, but the front and hind tarsi are black or dark fuscous above. The 
middle femora and tibiae are black or very dark, except at the extreme base and at the 
tips. Abdomen blue or green with fine purple reflections. Ovipositor black or 
blackish, more or less pale at the extrem.e tip, as long as the hind tarsi or nearly. 
Length $ 2'5 mm. 

Hab. Oahu near Honolulu, about 1500 ft. 

(46) Etipeimus dysoplias, sp. nov. 

Head above punctate or shagreened, blackish with metallic reflections, green or 
brassy, facial impression deep and narrow, finely shagreened, dull, with greenish tint at 
the bottom ; antennae slender basally, the first two or three funicle joints being quite 
elongate ; palpi dark, eyes collapsed, probably glabrous or nearly. Thorax blackish, 
but with brassy or golden reflections quite distinct. Front and hind coxae black, and 
somewhat metallic ; front femora and tibiae and the last four tarsal joints blackish 
or dark fuscous, the knee joint, apex of tibiae and basal tarsal joint yellow or pale. 
Middle legs yellow ; hind legs with the femora and tibiae black, the latter widely yellow 
at the apex, the knee joint brownish, the basal joint of the tarsus yellow, the following 
joints infuscate above. Neuration mostly fuscous. Abdomen bright green at the base, 
the rest blackish and very little metallic, pleura black ; ovipositor hardly exserted behind 
the abdomen, the tip somewhat pale. Length ? 2'25 mm. 

Hab. Oahu, Honolulu ; not much above sea level, under 100 ft. 

Subfam. MIRINAE. 

Anagyrus How. 

(i) Anagyrus major, sp. nov. 

Head reddish or yellowish brown, scape and pedicel of the antennae blackish or 
fuscous, the former being more or less yellow at the sides, five first funicle joints yellow, 
the sixth and the club black or dark fuscous. Thorax sordid fuscous above, more or 
less reddish-tinged, prothorax whitish, sides and ventral surface of thorax yellow. 
Abdomen dark fuscous at the base, the rest sordid yellowish or brown ; legs yellow or 
brownish yellow, the hind tibiae and femora at least more or less sordid, the colour of 
the legs apparently rather variable ; front tarsi dark. Marginal vein dark fuscous, the 
stigmal vein paler, and with an ill defined small infuscation about it. 

Head with a dense shagreening or close fine sculpture, pubescent, a rounded ridge 
between the antennae, the ocelli in a triangle, the outer ones well separated from the 



HYMENOPTERA 653 

margins of the eyes, the latter hairy. Scape of antennae somewhat dilated on the 
apical part beneath, pedicel obconical, elongate, considerably longer than the first joint 
of the funicle, the latter not differing much in length, the apical ones appearing some- 
what shorter than the basal, club with three subequal joints and about as long as the 
three preceding joints together. Thorax densely sculptured, the sculpture, which 
appears to be a close shagreening, more or less concealed by the conspicuous clothing 
of short white pubescence. Marginal vein subpunctiform, postmarginal very little 
developed, stigmal shortish. Hairless, oblique line very distinct, rest of wing closely 
and regularly set with hairs. Length 1-5 mm. Ovipositor very little prominent. 
Female. 

Hab. Hawaii, Kilauea. 

(2) Anagyrus xanthogaster, sp. nov. 

Very litce the preceding, but in its typical form smaller, the antennae shorter and 
very differently coloured, the general appearance being sordid yellowish, the apical 
joints darker, but without the strongly contrasted coloration of A. major. The abdomen 
yellow above. 

On Oahu I have found a larger form, not apparently differing in structure from the 
above, the mesonotum is reddish brown, the face nearly entirely pale yellow. 

Hab. Hawaii, Kilauea ; Oahu, on the coast. 

(3) Atiagyrus nigricans, sp. nov. 

Black, the legs yellow, the hind femora and tibiae slightly clouded. Wings hyaline 
but not quite clear. 

Head dull, with fine feeble and not dense puncturation ; antennae pale-tinted, not 
quite black, with the scape subdilated beneath, the pedicel obconical about as long as 
the first funiclar joint, which is much longer than wide, the following joints not differing 
much in length, all being more or less elongate, the club longer than the two preceding 
together. Mesonotum and scutellum slightly shining, very finely, remotely punctured. 
Abdomen triangular, acute at ape.x, deeply concave in dry specimens, and more or 
less pallid in the concavity. Neuration etc. as in the preceding. Length rather more 
than I mm. Female. 

Hab. Oahu, near Honolulu, 1500 ft. 

(4) Anagyrus fraternus, sp. nov. 

Very similar to A. nigricans, but the scape of the antennae is very little dilated, 
though somewhat thickened towards the apex, and the thoracic puncturation is rather 
more distinct and the punctures more numerous on the scutellum. The hind femora 

84—2 



654 FAUX A HAWAIIEXSIS 

are black or pitchy black, but the tibiae brightly yellow. The pedicel and the elongate 
funidar joints are all subequal and the club is longer than the two preceding joints, 
the antennal joints not differing much in their proportional length from those ot 
A. nigricans. 

\\'hat I take to be the male of this species has the antennae longer, and set with 
longish hairs, the thorax duller and more distinctly punctate, the pedicel shorter than 
the first funicle joint, the club about as long as the two preceding joints together. It is 
of a less black colour, more inclining to pitchy, especially at the sides of the thorax, and 
the legs are for the most part infuscate. or dirt)- yellow. The species appears to be 
rather larger than A. nigi^ans. 

Hab. Hawaii, Kilauea. 

(5) Anagynis laeviceps, sp. nov. 

Ver\" similar to A. fratemus and A. nigricans. The head in front is nearly 
smooth and the remote punctures are extremely faint : the mesonotum is very finely, 
evenly punctured, but not closely, the scutellum faintly and a good deal more sparsely. 
The antennae are slender with elongate funicle joints, the scape widened apically, but 
without a conspicuous dilatation, the proportional length of the funicle joints and 
pedicel much as in the above species, or subequal. Wings dusky h\-aline, legs black or 
dark pitchy, hind tibiae with a paler spot at the base, and darker than the tarsi. 
Length i-5 mm. Female. 

In the specimen described the sheath of the ovipositor is suberect or bent back- 
wards, and in side view appears as a pointed blade-like process. Probably it is usually 
closed down on the h\-popygium. 

Hab, Oahu : near Honolulu, 2000 ft. 

(6) Anag^rns tantaUus, sp. nov. 

Black, the posterior part of the thorax and the abdomen above within the concavit)- 
obscure or pitchy brown. Antennae black, the pedicel reddish at the apex. Legs for 
the most part yellow, the hind coxae dark, the hind femora infuscate especially on their 
outer surface. Wings lightly infuscate or yellowish-tinged. \'er)- similar to the several 
preceding species. 

Front ot the head above the antennae evenly and distinctly punctured, dull : scape 
ot antennae rounded on the lower side so as to be slightly thickened or dilated, pedicel 
obconical, rather longer than the first funicle joint, which is subelongate, as are the 
following and subequal. Mesonotum rather more finely punctured than the head. 
the scutellum still more finely. Abdomen distincdy longer than the head and thorax 
together. Length about 2 mm. Female. 

Hab. Oahu ; mount Tantalus. 



HYMEXOPTERA 655 

Blepvrus Howard. 

(i) Blepyrus marsdeni How. 

Blepynis ynarsdeni Howard, Proc. U.S. Nat. Mus. xxi. (1898), p. 234. 

This is a very abundant species throughout the islands and I do not feel certain 
that it is not synonymous with Eiicyrhis nisnlarts Cam. described on a single $. It 
would appear that if B. insularis is distinct from the above, it must now have become 
extinct, for it was found by Blackburn on several islands and, being no doubt a parasite 
of scale insects, could hardly have failed to have been bred by myself, Koebele or 
others, if it still existed. 

MiCROTERYs Thom. 

(i) Alicroterys flavus Howard. 
Hab. Common throughout the islands, on introduced scale-insects. 

Aphidenxvrtus Ashm, 
( I ) Aphidencyrhis ? sp. ? 
Hab. Oahu ; a common parasite of some imported Aphides. 

Helegonatopus Perk. 
( I ) Helegonatopus, sp. .'* 
Hab. Oahu ; parasitic on Eclithrodelphax and other Dryinidae. 
Obs. Several other minute and obscure Mirine Encyrtids, all no doubt imported, 
have been collected, but not yet determined. 

Fam. PTEROMALIDAE. 

It is almost certain that this vast family is unrepresented in the islands by endemic 
forms, excepting possibly by a few species of the subf Spalangiinae. Several introduced 
species have been found and, as some of them are spreading, they will no doubt become 
ubiquitous. Species of the genus Pteronialus have been imported for economic reasons. 
Most conspicuous of the Pteromalidae, that are foreign, are two or three species 
belonging to the subf Sphegigasterinae. 

Subfam. SPHEGIGASTERINAE. 
Tomocera How. 

(i) Tomocera ceroplastis Perk. 
Tomocera ceroplastis Perk., Proc. Hawaii Ent. Soc. i. p. 76. 
Hab. Oahu ; introduced. 



656 FAUNA HAWAIIENSIS 

Pachyneuron Walk. 
Pachyneuron spp. 
Two or more species of this or allied genera are parasitic on species known to be 
of foreign origin. 
Hab. Oahu. 

Subfam. SPALANGIINAE. 

Spalangia Latr. 
( I ) Spalangia canieroni, sp. nov. 

Female : Aeneous black, the abdomen deep black or blue-black, the tarsi yellow, 
dark at the tips. Head elongate, coarsely punctate, the punctures well separated from 
one another, the eyes with erect white hairs, longer than is usual on these organs. 
There is an elongate smooth median space at the bottom of the depression above the 
insertion of the antennae. Antennae with the second joint or pedicel elongate, very 
much longer than, but hardly twice as long as the first joint of the funicle, four terminal 
joints of the funicle quadrate, club nearly as long as the three preceding joints. 
Pronotum very densely rugose-punctate in front and at the sides between which is 
a smooth, more or less triangular, area, with at most a few punctures, this area bounded 
behind by a transverse costate line, the apical margin behind the line smooth. 
Mesonotum with a smooth median line, and a densely punctate area on each side of 
this ; the parapsides marked out by a costate or punctate furrow, the scapulae bearing 
some fine punctures ; axillae smooth and shining, like the scutellum, and separated 
from this by costate lines and both impunctate or nearly, the scutellum having a trans- 
verse costate line in front of the hind margin and the suture between it and the 
postscutellum is costate. Propodeum very smooth and shining, with a median narrow 
costate channel, divided by a longitudinal raised line, and punctate around its margins ; 
the pleura marked off by a costate suture, very densely and finely punctured below, but 
smoother near the suture. Wings clear hyaline, neuration brownish. Abdominal 
pedicel with about half a dozen raised longitudinal lines, the other segments very smooth, 
the fourth the largest, the third emarginate posteriorly, the apical ones with pale hairs. 
Length about 3 mm. 

Male very like the female, but the head is shorter, the pedicel of the antennae is 
minute, hardly longer than wide, much shorter than the first funicle joint, which is much 
longer than the following ; the other funicle joints longer than in the female, being 
longer than wide. Length 3 mm. but much smaller examples occur. This is probably 
the species referred to 5". hirta by Cameron. 

Distinguished at once from 5. Iiirta Hal. by the long second antennal joint in the 
female. 

Hab. Oahu, Hawaii, Molokai and no doubt all the islands. 



HYMENOPTERA 657 



(2) Spalangia simplex, sp. nov. 

Black, the head usually with slightly greenish tinge, the thorax more brassy and the 
fourth segment of the abdomen also slightly metallic, the post-petiolar one deep black. 
Tarsi yellowish or testaceous, apex of tibiae, the knee joints and the trochanters (more 
or less) also pale, reddish or testaceous. Front of head smooth and shining, with 
at most a few feeble punctures, but with a distinct fine medio-frontal crenate or punctate 
line reaching the anterior ocellus. Antennae of the female with the funicle joints short, 
increasing in width towards the club, and being mostly transverse, pedicel elongate, 
about as long as the two first funicle joints together, club as long or rather longer than 
the three last joints of the funicle. In the male the antennae are more elongate, the 
funicle joints all much longer than wide, the first much the longest, three times as long 
as wide, nearly as long as the next two together, the pedicel very small, hardly half the 
length of the first funicle joint, club about as long as the two preceding joints. 

Pronotum smooth and shining, but closely punctate on the neck and with a trans- 
verse curved crenate line near the hind margin ; mesonotum with a similar submedian 
transverse crenate line and with one or two pits behind it on the mediodorsal line. 
Axillae marked off posteriorly by strong crenate lines, a less distinct one between 
scutellum and postscutellum ; a distinct transverse crenate line on the former behind the 
middle ; propodeum acutely spined on each side, with a median longitudinal double 
crenate line and with crenate margins enclosing two large smooth spaces, the pleura 
roughly sculptured. Mesonotum and scutellum smooth and shining except as above 
noted. Wings clear, neuration brown. Pedicel of the abdomen with numerous fine 
longitudinal raised lines, twice as long as wide in the J, but a good deal shorter in the 
female, rest of abdomen smooth and polished. Length r^ — 2 mm. 

Hab. Oahu, Honolulu. 



Fam. EULOPHIDAE. 

To this enormously large and difficult family belong a considerable number of 
endemic Chalcids. The specimens preserve very badly in the tropics and, a few hours 
after death, distort and shrivel in such a manner as to preclude any proper description 
from being made, unless they are preserved in a special manner. They are best 
described when freshly killed. Though I have collected a considerable amount of 
material, I do not think it worth while to describe any new forms, until still more 
comprehensive and better preserved material is available. Many species of Aphelininae 
and some species of other subfamilies have been introduced for economic purposes, 
many also accidentally. To the species described by Ashmead in Vol. i. of this work 
two species only need be added here. 



658 FAUNA HAWAIIENSIS 

Paraphelinus Perk, 
(i) Paraphelinus xiphidii Perk. 
Paraphelinus xiphidii Perk., Bull. H. S. P. A. Exp. Station i. p. 264. 
Hab. Oahu, Maui, Kauai and probably all the islands. 

Melittobia Westw. 
(i) Melittobia hawaiiensis P. 
Melittobia hawaiiensis Perkins, Proc. Hawaii Ent. Soc. i. p. 124. 
Hab. Hawaii, Oahu ; Oahu, Honolulu ; Hawaii, Kilauea and no doubt throughout 
the islands. 

Fam. TRICHOGRAMMIDAE. 
There are several species of this family known in the islands, but it is doubtful 
whether there are any endemic forms. Those here described are almost certainly all 
foreign. Several other species are known to me, e.g. a species commonly parasitic in 
the eggs of the introduced Locustid Eliniaea appcndiculata, and one parasitic in the 
eggs of the Chrysopine genus Anomalochrysa. All the species invariably shrivel up 
and collapse within a few hours after death, if not placed in fluid, and are best described, 
when freshly killed. The genus Westwoodella belongs to the subfamily Oligositinae, all 
the other known Hawaiian Trichogrammids (including those undescribed) belong to the 
Trichogramminae. I have described two species under the genus Pentarthron, but this 
appears to me to be synonymous with West wood's genus Trichogramma. P.flavJtm is 
perhaps identical with Trichogramma pretiosa. 

Westwoodella Ashm. 

(i) Westwoodella hilaris, sp. nov. 

Head and thorax orange-red or brownish red, varying in tint in dried specimens, 
abdomen black, with a basal yellowish or reddish, wide ring, sides of thorax posteriorly 
dark, antennae sordid, darker in parts ; front legs sordid, hind legs with the femora 
dark, the tibiae and tarsi pale, tips of all the tarsi black or dark fuscous. 



Pedicel, funicle and club of antenna 
of Westwoodella hilaris 9 . 



Head transverse, collapsed in dry specimens, eyes pilose ; scape of antennae 
stoutish, narrowed apically, pedicel obconical, longer than thick, funicle joint rather like 
the pedicel, but considerably less thick and a little shorter, club long pointed, of three 
subequal elongate joints ; with the ring joint the antennae are seven-jointed. 



HYMENOPTERA 659 

Wings pedicellate, narrow, and with a long fringe, hyaline with a fuscous shade 
where the wing widens after the pedicel and another distinct blotch about the stigmal 
vein. Marginal vein as long as the submarginal, stout, bearing about six long bristles, 
the submarginal with one bristle near its middle, stigmal vein short, thick. Mesothorax 
with a few black bristles, scutellum with one on each side about its middle. Abdomen 
in dried specimens concave above, narrow and elongate, as long as head and thorax 
together. Length "5 mm. Female. 

Hab. Oahu, Honolulu ; amongst foreign grasses. 



Pentarthron Ril. 
(i) Pentarthron semifumatum, sp. nov. 

Black, the thorax at the sides and posteriorly more or less pitchy or obscurely 
reddish, head red, collapsing- in dried specimens, antennae sordid yellowish. Legs with 
the femora black or blackish, the tibiae and tarsi yellow, but the hind tibiae at least are 
sometimes black on their basal half. Wings with a smoky cloud on the front pair from 
their base to the region of the stigmal vein, and occupying all the basal part of the wing. 




7 



Antenna Pentarthron semifumatum $ . 

Antennae with five distinct joints and apparently a very minute ring joint, the scape 
long, the pedicel ovate, longer than the two funicle joints together and much wider, first 
funicle joint in some aspects appearing much wider than the second, club rather longer 
than pedicel and funicle together and without perceptible segmentation. Thorax and 
abdomen more or less shining in some aspects, the mesonotum under the microscope 
with a sculpture of very delicate reticulate lines. Abdomen just about as long as the 
head and thorax together, the ovipositor very little exserted. 

The truly marginal portion of the neuration of the front wings, i.e. the true 
marginal vein, is very short and bears a long bristle, there is another a little before this 
on the submarginal and one a little after it at the base of the stigmal, the three veins 
appearing as a continuous curve. There are many hair lines, some of which, about six 
or eight, stand out more distinctly than the others, owing to a considerable space on 
each side of them being hairless. Between these are many less distinctly separated 
hair lines. Length '5 mm. Female. 

Hab. Hawaii, Oahu, and probably all the islands, since it occurs both in the town 
of Honolulu and in the mountains of Hawaii above 4000 ft. 

F. H. II. 8s 



66o FAUNA HAWAIIENSIS 

(2) Pentarthron flavtim, sp. nov. 

Female, entirely flavous, the head in life probably still paler, the eyes and ocelli 
red, lateral ocelli near the eye-margins. 

Antennae six-jointed with a ring joint, the club solid, and equal in length to 
the pedicel ring joint and funicle together, the two funicle joints in some aspects appear 
not very different from one another, the scape with its basal pedicel is a good deal 
longer than the pedicel and funicle joints together. The mesonotum has a bristle 




Pentarthron Jlavum ? . 
Antenna in two rather different aspects. 

in front on each side, and another near the posterior angles, the scutellum one on each 
side near the posterior margin. The axillae encroach greatly on the parapsides of the 
mesonotum, so that the latter are subsecuriform, being dilated in front and sublinear 
posteriorly. Abdomen equal in length to the head and thorax together, the ovipositor 
not, or hardly, exserted. 

Wings hyaline about 7 or 9 more prominent hair lines, between which are 
a number of others closer together, or more confused, or incomplete. Length % -8 mm. 

Hab. Oahu, Honolulu. 

Fam. MYMARIDAE. 

The Mymarids are a very distinct group of Hymenoptera, to be ranked as 
a superfamily more distinct from any one other than, for instance, is the superfamily 
Apoidea from the Sphecoidea. They show affinities with the Proctotrupoidea, 
especially with the Diapriidae, and also with the parasitic Cynipoidea. Of the Chalcid 
series they appear more nearly related to some Eulophidae and Trichogrammidae, but 
the affinity is very remote. 

The endemic Hawaiian species are probably confined to the genus Polynema, but 
some species introduced either casually, or purposely for economic reasons, are also 
present, and now more numerous individually than the native forms. I have not 
thought it advisable at present to describe specimens from the intermediate islands, 
which may represent mere varieties of those occurring on Hawaii and Oahu. Con- 
sidering the difficulty of differentiating species from the different islands in the case of 
larger insects, it may naturally be expected that in minute parasitic Hymenoptera these 
difficulties will reach a climax. Various species, of which only one sex is known, 
I have also left undescribed at present. 



HYMEN OP TERA 66 1 

Subfam. GONATOCERINAE. 
Alaptus Hal. 

(i) Alaptus immattiiiis P. 
Alaptus immattirus Perk., Bull. H. S. P. A. Exp. Station i. p. 197. 
Hab. Oahu and probably all the islands ; bred from eggs of Psocidae. 

Leim.\cis Forst. 

(i) Leitnacis peregrina, sp. nov. 

Dark, blackish fuscous, more or less yellowish or tinged with yellow, especially 
along the sutures, the legs pale yellow, but in parts a little smoky, antennae smoky 
blackish, the scape and pedicel, more fuscous or yellow tinged in lateral view. Front 
wings hyaline except round the margin, and the marginal vein is dark. 

Antennae 8-jointed, the scape and club nearly equal, the pedicel large, obconical, 
about as long as the two following joints together, first funicle joint stouter and shorter 
than the second, which is more slender than the third, these two being nearly equal in 
length, fourth and fifth wider than the third, but not differing much from it in length, 
club almost as long as the whole of the funicle joints together. Thorax and abdomen 
nearly of equal length, the former truncate posteriorly, on the head and thorax a very 
few fine bristles are noticeable. Front wings spatulate, the longest cilia four or five 
times as long as the greatest width of the wing, the disc with a line of only three or 
four hairs, marginal vein long. Length '25 mm. 

Hab. Oahu, Honolulu. 

Subfam. MYMARfNAE. 

POLYNEMA Hal. 

(i) Polynema gigas, sp. nov. 

Black, the front legs yellow or brownish yellow, the middle and hind legs brown or 
partly yellowish brown, their tibiae and tarsi blackish ; abdominal petiole for the most 
part dark brown ; pedicel of the antennae yellowish at the sides, scape laterally pitchy 
or brown. 

Antennae of male long and filiform, the pedicel compressed, ovate, the first funicle 
joint decidedly shorter than the second, which is about six times as long as wide ; first 
funicle joint in the female short, barely one-half the length of the very long second joint, 
the three terminal ones elongate and not very unequal, the fifth joint being about four 
times as long as wide. Mesonotum with delicate rugulose or reticulate surface sculpture, 
propodeum with smooth and shining surface. Abdominal petiole narrowing to the 

85—2 



662 FAUNA HAWAIIENSIS 

apex, its upper side curved, wlien seen in profile ; ovipositor exserted to about the length 
of the claw joint of the hind tarsus. Wings with dark, ovate, punctiform marginal 
vein ; the longest cilia hardly half as long as the greatest width of the front wing. 
Length i"5 — 2 mm. 

Hab. Hawaii, 4000 ft. ; bred from shoots of Myrsine bored by the caterpillar of 
a very large species of Archips (Tortricidae), but I cannot conjecture the true host of 
this parasite. 



(2) Polynema ten-estris, sp. nov. 

Black, two basal antennal joints and all the legs yellow ; abdominal petiole yellow 
or brownish yellow or in the Z partly dark brown, third antennal joint in the % more or 
less pale. 

Antennae of $ very long and filiform, one and a half times as long as the whole 
insect, the joints of the funicle elongate and about equal, the third being five times (or 
more) as long as wide. First funicle joint in % hardly half as long as the second, which 
is considerably longer than the third, the fourth, fifth and sixth subequal, the last about 
three times as long as its greatest width, club rather longer than the two preceding. 
Sculpture of the thorax, which is somewhat shining, of the usual kind, but hardly 
visible under the strongest lenses ; front wings with dark, punctiform marginal vein, the 
longest cilia hardly half as long as the greatest width of the wing. Ovipositor of % 
very slightly exserted. Length r5 — 175 mm. 

Hab. Oahu, 1500 ft. and upwards. 



(3) Polynema pyrophila, sp. nov. 

Black, the second joint of the antennae beneath paler, more or less yellowish 
or brown, in the <? sometimes obscurely pitchy ; coxae and trochanters generally more 
or less brownish, tips of femora, except sometimes the hind ones, distinctly pale. 

Antennae of i with the pedicel roundish in lateral aspect, the first funicle joint 
rather shorter than the second, the following joints subequal and more than three times 
as long as wide. Antennae of ? very slender, the long ovate club rather longer than 
the two preceding joints together, the pedicel dilated, the first funicle joint hardly half 
as long as the second, which is about one and one-fifth longer than the third. Pronotum 
short, mesonotum hardly shining, with excessively fine reticulate surface sculpture, 
visible only under a very strong lens, the parapsidal furrows very distinct, the scutellum 
elongate and large, divided by a fine transverse line about the middle ; the propodeum 
and abdomen smooth and polished. Second abdominal segment much narrowed from 
the apex forwards to its junction with the petiole. Ovipositor prominent, as lono- 



HYMENOPTERA 663 

as the second joint of the hind tarsus. Wings evenly hairy, the marginal vein thick, 
punctiform, the longest cilia of the front wing only about half the greatest width of 
this. Length 1-3 — 1-4 mm. 

Hab. Hawaii ; 4000 ft. 



(4) Polynema scrutator, sp. nov. 

Male black, the abdominal pedicel and all the legs yellowish or brownish yellow, 
the pedicel sometimes somewhat darkened ; all the tibiae, the middle and hind femora 
and sometimes also the front ones brown or dark brownish, claw joint of tarsi dark. 
Antennae with the pedicel yellow laterally, the scape sometimes brown or piceous at the 
sides. 

Antennae long and filiform, the joints of the funicle subequal, the third about three 
times as long as its greatest width, the pedicel compressed, dilated, being very strongly 
rounded beneath, first funicle joint a little shorter than the second. Thorax with 
distinct microscopic rugulosity under strong magnification, the abdomen smooth, shining, 
with the pedicel much narrowed from base to apex. Front wings with the marginal 
vein ovate, subpunctiform, dark, the longest cilia much shorter than the greatest width 
of the wing, or only about half as long. Length i mm. 

Hab. Oahu, Honolulu. Though I have on various occasions found this species 
in the town of Honolulu, I have never at any time taken a female of the males 
described above. Males and females of P. reduvioli occur with it, in some localities. 



(5) Polynema rubriventris, sp. nov. 

Female black, the abdomen ferruginous, sometimes partly dark or infuscate ; 
the legs, except the apical tarsal joints, entirely yellow, as also the two basal and some- 
times the third antennal joint ; the club and three preceding funicle joints black, while 
the more basal funicle joints are mostly more or less sordidly yellowish or brown. 

Antennae slender, the pedicel compressed, obconic, the first funicle joint half as 
long as the very slender and elongate third, the three apical joints of the funicle 
subequal, slender, three or four times as long as wide, the club as long as the two 
preceding joints. Thorax hardly shining, the minute surface sculpture or rugulosity 
distinct, the impresso-punctate transverse line on the scutellum about two-thirds the 
length of the latter from its front margin. Abdomen smooth and shining, the ovipositor 
distinctly subexserted. Marginal view of front wings subovate, the longest cilia not 
half as long as the greatest width of the wing. Length f6 — 175 mm. 

Hab. Oahu ; 1500 ft. and upwards. 



664 FAUNA HAWAIIENSIS 

(6) Polynema tantalea, sp. nov. 

Male ferruginous, the head and the abdomen, except its base and the petiole, and 
the apical joints of the tarsi black ; middle and hind tibiae blackish and their tarsi more 
or less infuscate. Antennae black, the two basal joints clear yellow. Legs, except as 
noted above, yellow. 

Antennae very long and slender filiform, much longer than the whole insect, the 
first funicle joint considerably shorter than the second, the several following this not 
differing much in length, the third being about five times as long as its greatest width, 
whilst several of the apical joints are noticeably shorter than these basal ones. Thorax 
dull, with very fine surface sculpture, the transverse line near the hind margin of the 
scutellum distinct. Abdomen smooth and shining. Front wings evenly pubescent, 
marginal vein punctiform, dark, the submarginal yellow, longest cilia not half as long as 
the greatest width of the wing. Length r25 mm. 

Hab. Oahu ; 1500 — 2000 ft. 

(7) Polynema oahuensis, sp. nov. 

Ferruginous or yellowish brown, the head black, but the occipital region and 
the face below the antennae is concolorous with the rest of the body ; apical joint 
of the tarsi black, as well as the club and the three preceding antennal joints. 

Antennae of the ? with the first funicle joint quite short, not half as long as 
the very long second joint, the pedicel obconic, the club as long as the two preceding 
joints together. Thorax hardly at all shining, the surface sculpture very fine ; transverse 
impressed line near the hind margin of the scutellum distinct ; abdomen smooth and 
shining, the ovipositor distinctly subexserted, black. Wings evenly pubescent, marginal 
vein punctiform, dark ; longest cilia of front wings, not more than half the length 
of the greatest width of the wing. Length $ r5 mm. 

Hab. Oahu ; 1 500 — 2000 ft. 

(8) Polynema Iriscia, sp. nov. 

Ferruginous, a transverse line across the head in front, the apical joints of the 
middle and hind tarsi, and the apical part of the abdomen, black or dark-coloured. The 
hind tibiae infuscate on their apical portion. Basal two joints of antennae clear yellow, 
the third yellowish but infuscate, the following shows more or less of a yellow tinge, 
while most of the flagellum is black. 

Antennae of male very long, decidedly longer than the whole insect, filiform, 
attenuate apically, scape short, wide in lateral aspect, pedicel very short, ovate or 
subrotundate, first flagellar joint elongate, three times as long as its greatest width and 
just about equal to the second, the latter and the following joints being all nearly equal 
in length, and all fluted on their flattened sides. Mesonotum and scutellum dull, with 



HYMENOPTERA 665 

the usual minute sculpture (reticulated under a compound microscope) distinct ; 
propodeum smooth and shining. Wings with the neuration fuscous, the marginal vein 
of the front wings subovate, beneath which is a fuscous fascia, a second fascia near the 
middle of the wing, while a third occupies the apical part of the wing and is broader 
than the clear space between itself and the middle fascia ; cilia short, the longest 
not half the length of the greatest width of the wing. Abdomen shining. Length ^ 
(abdomen somewhat contracted) i"5 mm. 

Hab. Oahu ; Koolau range, 1500 ft. 

(9) Polynema apicalis, sp. nov. 

Ferruginous, the antennae except the two basal joints, which are more or less 
yellow or brown, black ; the middle tibiae are noticeably darkened, the hind ones, 
except basally, nearly black ; the hind femora are also dark on the apical portion ; and 
the apical joint of all the tarsi is black. Wings hyaline, but widely black on the apical 
portion. Vertex of head with a transverse black line in front. 

Antennae with the pedicel subtriangular or obconic, the first funicle joint very short, 
about equal to the pedicel but much more slender, the second very long, more than 
twice as long as the preceding, and also much longer than the third ; fourth, fifth and 
sixth not differing much in length and shorter than the third, club ellipsoidal, somewhat 
shorter than the three preceding joints together. Thorax not shining, appearing 
densely minutely punctate, from the microscopic reticulation of the surface. Marginal 
vein dark, elongate-ovate, the longest cilia of front wings not equal in length to half the 
greatest width of the wing. Abdomen smooth and shining, the ovipositor very slightly 
exserted. Length i"6 mm. 

Hab. Oahu ; Honolulu mountains, about 2000 ft. 

(10) Polynetna perforator, sp. nov. 

Yellow or brownish yellow, the head, abdomen except the pedicel, and the claw 
joints of all the tarsi black. Antennae with the three basal joints in the ^, and two in 
the $, yellow ; the following joints more or less infuscate, the club fuscous or blackish. 

Antennae in the t long and filiform, the pedicel compressed and dilated, strongly 
rounded beneath, the first funicle joint rather shorter than the second ; in the % the first 
funicle joint much shorter than the next, not two-thirds of its length, the club large and 
as long as the three preceding joints together. Thorax sculptured as in the other 
species, the abdomen smooth and shining. Ovipositor as long or rather longer than the 
claw joint of the hind tarsi. Front wings with marginal vein subpunctiform, dark- 
coloured, ovate, the longest cilia hardly half as long as the greatest width of the wing. 
Length 75 — V2 mm. 

Hab. Oahu, in the mountains. 



666 FAUNA HAWAIIENSIS 

(ii) Polynema ciliata, sp. nov. 

Black, the two basal joints of the antennae, the pedicel of the abdomen, and 
all the legs, pale yellow, the pulvillus of all the tarsi dark. 

Head smooth and shining, the pedicel of the antennae strongly compressed, 
obconical, or ovate, as long or a little longer than the slender first joint of the funicle, 
the latter about half the length of the very elongate second joint, the third rather 
shorter than the second, the three following joints on their wide surface much wider 
than the preceding joints, the two first of these being subequal, the last conspicuously 
more elongate, the club just about equal to the three preceding joints together. The 
club and funicle joints are blackish or smoky, the basal funicle joints sometimes showing 
a yellowish tinge ; the scape and pedicel are almost entirely yellow, the upper edge 
of the former and the tip of the latter being darkened. Mesonotum shining, hardly or 
not visibly sculptured under a strong lens, the propodeum quite smooth and shining. 
Abdomen smooth and polished, the ovipositor very slightly exserted. Front wings 
hyaline basally to considerably beyond the marginal vein, the rest appearing darker, 
with even clothing of black hairs, marginal vein short more or less pale, longest cilia as 
long as the greatest width of the wing, or rather longer. Length $ hardly i mm. when 
fully extended. 

Hab. Oahu ; 1500 ft. ; parasitic in the eggs of a small Delphacid. 



(12) Polynema jassidaruni, sp. nov. 

Black, the antennae black or fuscous in the male, the second joint is always 
noticeably yellowish, and the scape and first funicle joint may be more or less obscurely 
or sordidly pale. In the female the scape is dark or black, the pedicel and most of the 
funicle joints yellow, the club and preceding funicle joint always dark. Legs and 
abdominal pedicel yellow or brownish yellow, the femora and tibiae usually more or less 
embrowned, the apical joint of the tarsi black. 

Antennae of the ^ considerably longer than the whole insect, the pedicel roundish 
in lateral view, the funicle joints subequally elongate, the third about two and a half or 
three times as long as wide. Female with the first funicle joint short, only about half 
the length of the second, the latter slender and the longest of all, the club as long as 
the three preceding joints, which are subequal and shorter than the third funicle joint. 
Form and sculpture of the thorax as in the other species ; the abdomen smooth and 
shining, with the ovipositor distinctly exserted. Wings with the marginal vein puncti- 
form, dark brown or brownish yellow, variable in colour ; the longest cilia of the front 
wings rather less than the greatest width of the wing. Length '65 — 'Ss mm. 

Hab. Oahu near the coast ; parasitic on eggs of a Jassid. 



HYMENOPTERA 667 

(13) Polyncma nana, sp. nov. 

Black, the antennae entirely black above, but the scape and pedicel in lateral view 
are usually more or less pale, piceous or sometimes yellowish ; legs blackish or dark 
brown, but the tarsi are paler, generally yellowish, sometimes infuscate above, the apical 
joint being conspicuously black. 

Antennae of t considerably longer than the whole insect, pedicel compressed, 
dilated, strongly rounded beneath, the first funicle joint decidedly shorter than the 
second, the third and following subequal, the latter being rather more than twice as long 
as its greatest width. In the % the first funicle joint is short, much shorter than the 
second, the club as long as the three preceding joints together, the pedicel compressed, 
ovate. Pronotum short and hardly noticeable in dorsal aspect, the mesonotum shining, 
with distinct parapsidal furrows and a very delicate reticulate sculpture under high 
powers of the microscope. Scutellum large and long as in the other Hawaiian species. 
Wings evenly hairy, marginal vein dark, punctiform, the cilia, where longest, as long as 
or longer than the greatest width of the front wing. Abdomen smooth, shining, the 
ovipositor slightly, but distinctly, exserted. Length about '5 mm. 

Hab. Hawaii, Kilauea. One apterous female, taken with the winged ones, 
appears identical with the above. 

(14) Polyncma reduvioli P. 
Polynenm reduvioli Perk., Bull. H. S. P. A. Exp. St. i. p. 196. 
Hab. Oahu, Kauai, Hawaii, and no doubt all the islands. 

CYNIPOIDEA. 

Fam. FIGITIDAE. 

Subfam. EUCOILINAE. 

The species of Eucoiline Cynipoids are numerous in the islands, and unquestionably 
difficult to determine specifically. Ashmead referred the Hawaiian species described by 
him to a number of genera, but to me all the Hawaiian species appear intimately allied 
and likely to prove still more intimately so, as new species are found. Kiefifer in 
Wytsman's Genera Ins. refers Ashmead's Hawaiian species to three genera, and 
I believe the Pilinothrix of Ashmead should have been placed with the forms, which 
Kieffer refers to Cothonaspis, subg. Anectocleis. It is also noteworthy that the 
Hawaiian species with closed marginal cell, referred by Kieffer to Eticoila, subg. 
Psichacra, very closely resemble some of the forms with open marginal cell and that 
this cell in different species varies very much as to how far it is open above. In fact in 
some Hawaiian forms it is not at all easy to decide whether the marginal cell is really 
closed or open. 

F. H. II. 86 



668 FAUNA HAWAIIENSIS 

Kieffer gives as a character of Cothonaspis (s. 1.) "cupula non convexe," but in all 
or nearly all the Hawaiian forms it is more or less convex and sometimes very strongly 
so. Only in the species, for which 1 have formed the subg. Nesodiranchis is the 
scutellar cup depressed. 

Under the circumstances, I think it better to place all the Hawaiian species with 
more or less open marginal cell (excepting the one just referred to) in a subgenus 
Hypodh^anchis Ashm., treating this as a section of Cothonaspis. Ashmead's characters 
for Hypodiranchis will not fit most of the species here described, nor in fact do the 
examples he described agree with the generic characters he laid down. In the sense 
used here, Hypodiranchis will refer to a division of Cothonaspis in which the cup of the 
scutellum is generally more or less convex, sometimes strongly so, and never impressed, 
and the antennae of the female are without a definite club. 

I find it impossible to refer males, caught at large, to their proper females with any 
certainty and therefore have described none of this sex. Had I done so the list 
of species would have been much increased, but no doubt many or most of the names 
given to males would become mere synonyms of previously described females. By 
breeding, or special collecting in the field, the sexes will no doubt be rightly assigned in 
the future. 

I may note that several species of parasitic Cynipids have been introduced into the 
islands for economic purposes, but these are not considered in this account. 

Cothonaspis Hart. 

Nesodiranchis, subgen. no v. 

Like Diranchis, as defined by Ashmead, in most characters, but with the antennae 
not at all clavate, the more basal flagellar joints being larger and longer than those at 
the apex, so that the antennae become distinctly attenuate towards the tips. Third 
antennal joint slightly longer than the fourth, all of these beyond the pedicel being 
elongate, the sides slightly rounded, the surface very finely striate or rugulose rather 
than fluted, fourth and fifth joints about twice as long as wide, apical joint about one 
and one-half times the length of the preceding. Scutellar cup large, ovate, depressed, 
the posterior fovea large, transverse ; marginal cell in front wings entirely open above, 
cubitus effaced, represented by a hair line ; abdomen about as long as the thorax, with 
a slight hairy girdle at base, hypopygium not conspicuously prominent. 

(i) Cothonaspis {Nesodiranchis) ashnieadi, sp. nov. 

Black, the mandibles, the two basal antennal joints (and the third obscurely) red ; 
legs yellow, tarsal joints mostly black. Neuration brownish towards the base, but the 
radius darker, the last abscissa black or almost so. 



HYMENOPTERA 669 

Antennae shortly pilose, pedicel short, hardly longer than wide ; mesonotum 
smooth and shining, scutellar cup distinctly concave, with a few punctures on each side. 




Nesodiranchis ashmeadi ? . 
Antenna and cup of scutellum. 

in front of the posterior fovea, distinct ; sides of cup pilose, with close rugose punctura- 
tion ; propodeum with a carina on each side of the middle, the area between these 
shining ; on each side of this area externally the propodeum is pubescent. Abdomen 
smooth and shining, with a thin basal pubescent girdle. Length 2 mm. Female. 

Hab. Oahu, 1500 — 2000 ft. 

Subgen. CoTHONASPis Forst. 
(2) CotJionaspis {Hypodiranchis) pelc, sp. nov. 

Black, polished, the mandibles and all the legs ferruginous, the tarsi, at least the 
hind and middle ones, more or less infuscate above, the pedicel of the antennae 
obscurely red or pitchy. 

Antennae moderately long, rather longer than the head and thorax together, 
13-jointed, the pedicel hardly longer than wide, the first funiclar joint elongate, more 
than twice as long as wide, distinctly, though not greatly, longer than the second, and 
about equal to the third ; the first and second distinctly more slender than the latter, 
the latter more so than the fourth, this and all the following being subequal and 
moniliform, each longer than wide, and all fluted. Mesonotum very smooth and 
shining, with sparse pale hairs ; scutellar cup narrowly ovate, pointed anteriorly, twice 
as long as its greatest width, with a large fovea posteriorly, and with some distinct 
piligerous punctures on each side in front of the fovea ; sides of scutellum beneath the 
disc distinctly rugose and posteriorly pilose. Meso- and metapleura smooth. Wings 
hyaline, ciliated, neuration dark and strong, the marginal cell open above for about half 
its length, the cubitus much fainter than the rest of the neuration. Propodeum carinate 
on each side of the middle, the surface dull and thinly pubescent. Abdomen about 
equal in length to the head and thorax together, with a rather sparse and inconspicuous 
hairy girdle at the base, the hypopygium strongly exserted. Length 2 mm. $. 

Hab. Hawaii, Kilauea ; ovipositing in Dipterous larvae, belonging to the genus 
Drosophila. 

86—2 



670 FAUNA HAWAIIENSIS 

(3) CotJionaspis {H.) intermedia, sp. nov. 

Female black, polished, the legs brown, the hind ones darker, the front and middle 
pair more yellowish brown or testaceous. Antennae obscure brownish or pitchy, the 
apical joints black, or nearly so, the pedicel being the palest joint. 

The joints of the flagellum are all elongate, the first being distinctly longer than 
the second, the following joints subequal, the more apical slightly wider than the first 
and second ; the third joint is nearly 2^ times as long as its greatest width, the ante- 
penultimate joint only about twice as long as wide. Scutellar disc very convex above, 
very long and narrow, subparallel-sided, the very fine margins generally, in some 
aspects, appearing slightly sinuate inwardly behind the middle, the fovea very large, 
with one or two very fine punctures on each side in front of the fovea ; below the disc 
the scutellum is finely and regularly longitudinally striate and sparsely hairy ; propodeum 
with the median area between the carinae punctate, the sides with yellow pubescence, 
like that of the abdominal girdle, the latter also having an apical ciliation of longer hairs. 
Meso- and metapleura smooth. Wings hyaline, somewhat yellow-tinged, especially 
about the nervures, which are rather dark fuscous ; cubitus pallid, not distinct, marginal 
cell closed above for about half its length from the base. Hypopygium rather strongly 
exserted. Length 2 '5 mm. $. 

Male very like the female, but with much longer, filiform antennae, the fiagellar 
joints subequally elongate, the third being about 2\ times as long as wide, the 
antepenultimate about 4 times, owing to the attenuation of the antennae apically, the 
first distinctly, but slightly, curved. 

Var. Scutellum at the sides, more or less of the sides of the abdomen at the base, 
more or less of the basal joints of the antennae, or some of these parts, reddish, or 
pitchy. 

Hab. Oahu ; widely distributed in the mountains above 1300 ft. 

(4) Cothonaspis {H.) tantali, sp. nov. 

Polished black, the mandibles and the pedicel of the antennae dark red, all the legs 
ferruginous, the apical joint of the front and middle tarsi and most of the posterior tarsi 
above, infuscate. 

Antennae long and filiform, 13-jointed, slightly longer than the whole insect, the 
pedicel short, hardly longer than wide, the first flagellar joint slightly longer than the 
second, the latter three times as long as its greatest width, the joints towards the apex 
of the antennae a little wider in proportion to their length, the penultimate joint being 
rather more than twice as long as its greatest width, the terminal joint long, one and 
two-thirds the length of the preceding. Mesonotum polished, sparsely pilose ; scutellar 
disc narrow, two and a half times (or more) as long as its greatest width, deeply and 



HYMENOPTERA 671 

largely foveated posteriorly, and with a series of distinct marginal piligerous punctures 
on each side, from near the base to the fovea ; sides of scutellum below the disc 
longitudinally wrinkled and pilose ; propodeum between the carinae very dull and 
densely rugulose, at the sides with a patch of dense woolly hair ; mesopleura smooth, 
metapleura longitudinally strigose. Wings hyaline, faintly yellow-tinged, ciliated ; 
neuration for the most part rather pale yellowish brown, marginal cell open above, the 
cubitus much fainter than the other nervures. Abdomen elongate-ovate pointed at apex, 
with a dense basal girdle of yellowish pubescence, the hypopygium not conspicuously 
exserted, but more so in some examples than in others. The basal flagellar joints of 
the antennae are sometimes more or less reddish-tinged. Length 2 '25 mm. $. 

The antennae are so slender that they can hardly be called subclavate, and 
possibly this species could be referred to Forster's genus Anectoclis. 

Hab. Oahu, near Honolulu, above 1500 ft. 

(5) Cothonaspis {H.) strigosa, sp. nov. 

Extremely similar to P. tantali, having the same scutellar structure, similar 
antennae etc. It is readily distinguished, however, by the mesopleura posteriorly being 
strongly longitudinally strigose, like the metapleura, instead of being altogether smooth 
and shining. 

Hab. Oahu; Honolulu, above 1500ft. 

(6) Cothonaspis (//.) declivis, sp. nov. 

Black, the two basal antennal joints red, mandibles and legs testaceous or brownish 
yellow. 

Antennae long, about as long as the whole insect, all the flagellar joints elongate, 
the flagellum appearing subclavate, owing to the more apical joints being proportionately 
rather wider and shorter than the more basal ones, in some aspects ; first and second 
flagellar joints nearly equal in length, the latter slightly stouter, about equal to the 
following, but rather more slender, third about three times as long as wide, the ante- 
penultimate joint about twice as long as wide. Mesonotum polished, sparsely pilose, 
pleural region smooth and polished, including the metapleura. Scutellar cup more than 
twice as long as wide, very convex longitudinally and very strongly declivous posteriorly 
before the large round fovea, also strongly convex transversely, sides of the cup 
rugulose, but partly smooth ; propodeum dull between the carinae, but very indistinctly 
sculptured. Wings ciliated, neuration brownish yellow, the marginal cell open above, 
except for a short distance at base, cubitus very faint. Hypopygium not prominent. 
Length 2 mm. %. 

Hab. Oahu; near Honolulu, above 1500 ft. 



6/2 FAUNA HAWAIIENSIS 

Note. With P. dcclivis and P. tantali I take a male Cynipid, which I cannot 
refer to either with any certainty. It may be distinct from either, but I suspect this is 
not the case. 

(7) Cothonaspis {H.) naias, sp. nov. 

Black, polished, five basal joints of the antennae red, mandibles and legs testaceous 
or ferruginous, hind coxae browner, hind tarsi evidently infuscate. 

Antennae two-thirds the length of the whole insect, the basal joint of the flagellum 
equal in length to the third, but evidently more slender, the second joint being much 
shorter, about two-thirds the length of these, fourth and following joints wider than the 
third, all elongate, submoniliform, subequal, the antepenultimate joint about twice as 
long as wide. Scutellar cup long and narrow, acuminate in front, foveated at its 
posterior end, about three times as long as wide, with several evident punctures on its 
smooth surface, its sides distinctly, finely rugulose. Propodeum with a dense patch of 
pubescence outside each of the longitudinal carinae ; meso- and metapleura smooth and 
polished. Wings clear, hyaline, ciliated, marginal cell apparently closed above on less 
than its basal half, neuration yellowish brown, cubitus faint. Hypopygium somewhat 
prominent. Length 175 mm. $. 

Hab. Oahu ; 1 500 ft. 

(8) Cothonaspis {H.) debilis, sp. nov. 

Black, polished, mandibles and two basal antennal joints yellowish brown, some 
other of the basal flagellar joints also more or less pale, legs testaceous, apices of tarsi 
dark. 

Antennae as long as the whole insect, slender, but gradually subclavate, the two 
basal flagellar joints being notably more slender than those towards the apex. First 
flagellar joint subequal to or a trifle longer than the second, which is subequal to the 
following joints ; fluted sculpture distinct ; antepenultimate joint of antennae twice as 
long as wide. Scutellar cup elongate-ovate, twice as long as its greatest width, foveate 
at its posterior end, and with a few fine punctures before the fovea ; sides very finely 
rugulose ; propodeal carinae distinct, the pubescence not dense ; mesothoracic and 
metathoracic pleura smooth, polished. Neuration brown, yellow-tinged, cubitus much 
paler and fainter than radius, but easily seen, marginal cell closed above for only a short 
distance at the base. Hairy girdle of abdomen not dense, hypopygium not strongly 
prominent. Length hardly 1-5 mm. $. 

C. debilis van siniilis van nov. 

Very like the above in api^earance and structure, but the hind coxae are rather 
darker brown, the scutellar cup is slightly wider, its margins more distinct and at the 
sides it is shining and smooth instead of being rugulose. 



HYMENOPTERA 673 

C. debilis var. subdcbilis van nov. 

Antennae paler than in typical debits (var. shnilis being intermediate), scutellar 
cup decidedly narrower than in either of the above forms, about 2\ times as long 
as wide, shining and nearly smooth on its sides, or at least hardly visibly sculptured 
even under a very strong lens. 

Hab. Hawaii ; Kilauea. Without a long series of examples the three forms 
described can hardly be considered as species. 

(9) Cothonaspis {H.) abnormis, sp. nov. 

Black, polished, two basal antennal joints, mandibles and all the legs ferruginous 
or testaceous. 

Antennae long and slender, as long as the whole insect, the flagellum feebly 
subclavate, the apical joints being evidently wider, when compared with the basal joints, 
but all are elongate, the antepenultimate joint being about three times as long as wide. 
Three basal joints almost of equal length, though the basal one, on actual measurement 
is a trifle shorter. Scutellar cup elongate, not distinctly margined, and best seen when 
looked at obliquely from in front. It is of narrow ellipsoidal form and very little raised, 
the sides being extremely low, rugosely sculptured, and there is no posterior fovea ; 
propodeum bicarinate, dull and hardly perceptibly sculptured between the carinae, 
pubescent at the sides ; pleura smooth, the mesopleura with an impression. Wings 
more or less yellowish-tinged, marginal cell closed above on about the basal half, 
neuration brownish yellow, cubitus rather distinct. Abdomen strongly compressed, 
with complete hairy girdle, the hypopygium prominent. Length 1-5 — 175 mm. $. 

Hab. Oahu ; 1500 — 2000 ft. 

Obs. This species might be placed in a distinct genus, but the possibility of the 
occurrence in the islands of others, which would connect it with more usual forms, 
renders it inadvisable to separate it generically at present. 

(10) Cothonaspis {H.) dubiosa, sp. nov. 

Black, polished, two basal joints of antennae and mandibles distinctly red, legs 
flavo-testaceous, abdomen blackish or very dark brown over most of the basal portion, 
becoming reddish apically, as also widely so at the sides beneath. Front and middle 
tarsi brownish, the posterior tarsi dark fuscous. 

Antennae about two-thirds the length of the insect, the flagellum subclavate, fluted, 
its third joint being evidently wider than the second, the latter and the first almost 
equal in length, the third and fourth very slightly longer, the antepenultimate joint 
twice as wide as long, the third rather longer, about 2^ times its width and evidently 
fluted. Scutellar cup elongate, twice as long as its greatest width, very distinctly 



674 FAUNA HAWAIIENSIS 

marked by its pale margins throughout, a few scattered punctures (in the type specimen) 
in front of the posterior fovea ; sides of the cup somewhat shining, with some very fine 
punctures, and extremely fine longitudinal rugulosity, sparsely pilose ; pleura smooth 
and polished ; propodeum with the two longitudinal carinae as in the other species. 
Wings ciliate, the neuration yellowish at base, becoming darker apically, the apical 
side of the marginal cell being- dark fuscous ; this cell closed on its basal half above. 
Length rather more than 2 mm. $. 

I cannot be sure that the marginal cell in this species is not altogether closed, as 
in some aspects the nervure on the upper side of the cell appears continuous to the 
apex, though fainter beyond the middle. 

Hab. Oahu ; 1500 ft. I have only examined one e.xample and the abdomen is 
partly crushed. 

(11) Cothonaspis {H.) dichroina, sp. nov. 

Black, polished, the two basal joints of the antennae distinctly red, as well as the 
mandibles, legs testaceous, the tarsi more or less infuscate, the hind ones dark fuscous 
or blackish above, abdomen ferruginous. 

Antennae rather more than two-thirds the length of the whole insect, the flagellum 
subclavate, its second joint being considerably less wide than the third, its first joint 
more slender still ; the first of these joints very little longer than the second, and all the 
flagellar joints are elongate, submoniliform, the antepenultimate (with its contracted 
base) being about twice as long as wide. Scutellar cup elongate, rather more than 
twice as long as wide, pointed anteriorly and foveated at the other e.xtremity, one or 
two fine punctures inside either margin ; at the sides it is densely rugosely sculptured 
or rugose-punctate and sparsely pilose. Propodeum between the carinae impressed, 
slightly hairy, hardly sculptured. Pleura smooth, not sculptured. Hypopygium some- 
what exserted. Length 2 mm. %. 

Hab. Oahu, about 1500 ft. 

EucoiLA Westw. 
( I ) Eucoila {Psichacra) oreias, sp. nov. 

Black, smooth and shining, mandibles red on their apical part, coxae black or very 
dark ; front and middle femora mostly dark brown above, the apical part being pale 
brownish yellow, front and middle tibiae yellowish brown, the latter sometimes darker 
than the former ; hind legs darker than the others, the femora black or nearly, excepting 
the yellowish apices, tibiae dark brown except at base, the tarsi entirely blackish or 
dark fuscous. Neuration yellowish, the apical side of the marginal cell paler than most 
of the neuration. 

Antennae as long as the entire insect, or almost so, the first two joints beyond the 
pedicel being somewhat thinner than the next. The flagellar joints are subequal in 



HYMENOPTERA 675 

length, the first being slightly shorter than the second, both elongate and slender, the 
antepenultimate joint of nearly the same length as these, but much stouter, about twice 
as long as wide. The scutellar cup is elongate-ovate, its greatest width being less than 
half its length ; its margins are not very distinct, its dorsal surface bears no punctures 
in front of the posterior fovea, or at most one or two very feeble ones, but its sides are 
somewhat roughened by a number of punctures. Abdominal hairy girdle not dense. 
Length $ r75 mm. 

Hab. Hawaii, Kilauea. 



(2) Eiicoila (P.) orobates, sp. nov. 

Extremely like E. oreias, with similar long antennae, which do not differ much in 
structure from those of that species. The legs, however, are less dark in nearly all 
parts, brown and, where paler, yellowish, the neuration of the wings is darker, fuscous ; 
the scutellar cup is well margined, wider, about twice as long as its greatest width, and 
bears punctures on each side dorsally in front of the posterior fovea, its sides are quite 
smooth, polished and impunctate or nearly. Length % 175 mm. 

Hab. Hawaii, Kilauea. 



(3) Eucoila [P.) hygropJiila, sp. nov. 

Black, polished, legs ferruginous or testaceous, hind coxae and sometimes the 
middle ones darkish brown, the hind femora and tibiae also browner, the hind tarsi 
dark fuscous ; several of the basal antennal joints more or less distinctly reddish. 
Neuration fuscous, more or less yellow-tinged. Antennae considerably shorter than 
the whole insect, without a definite club, but the fourth joint of the flagellum is 
considerably thicker than the third, the first joint being decidedly longer than the 
second or third, which are subequal and shorter than the fourth ; the antepenultimate 
joint is considerably less than twice as long as its width. The scutellar cup is well 
defined, not at all impressed, but with a distinct posterior fovea, very elongate, about 
three times as long as wide ; the sides are finely longitudinally rugulose ; sides of the 
propodeum and the basal abdominal girdle densely hairy. Length % i "5 — 2 mm. 

Eucoila hygrophila \'ar. pliilygra nov. 

Like the above in sculpture etc. but with the neuration yellower, and the basal 
joints of the flagellum rather different, the first joint being slightly but distinctly longer 
than the second, and about equal to the third. Length % i "2 mm. 

Hab. Oahu ; 1500 — 2000 ft. 

F. H. II. %j 



676 FAUNA HAWAIIENSIS 

EucoiLiDEA Ashm. 

(i) Ejuoilidea micromorpha, sp. nov. 

Black, abdomen beneath at least at the base yellowish or brownish yellow. Legs 
mostly testaceous, the front and middle femora distinctly dark brown above except 
apically, tarsi and sometimes the posterior tibiae also more or less dark. Head, 
abdomen, and mesonotum smooth and polished. Antennae with short, globose pedicel, 
funicle with the joints increasing slightly in width towards the apex of the antennae, 
and fluted, the first and second joints nearly equal in length and the slenderest, the 
third being slightly wider than the latter, and also of about the same length, anteapical 
joints longer than their greatest width, but not much longer. Scutellar cup with a 
distinct complete series of marginal punctures just within its border, rugose-punctate 
below this. Mesopleura smooth and shining. Propodeum very finely sculptured in the 
middle between the carinae, and with a pubescent patch external to the carinae on each 
side in front. Abdominal petiole longitudinally costate. Neuration sordid yellowish. 
Length \'2 mm. Female. 

Var. a. Smaller, the antennae more pitchy, or blackish fuscous, abdomen more 
widely yellowish brown from base to apex, posterior femora as dark, or almost as dark, 
as the others. 

Hab. Oahu, Honolulu on the lowlands and in the mountains to 1500 ft. 

ICHNE UMONOIDEA . 

Fam. ICHNEUMONIDAE. 

Subfam. CRYPTINAE. 
Cryptine Ichneumons are represented, in addition to the introduced species 
enumerated in Vol. i., by a single species of the tribe Hemitelini, a common parasite 
in the nests of spiders. I have little doubt the species is introduced. It is widely 
distributed in the islands. 

Subfam. PIMPLINAE. 
PiMPLA Fabr. 

( I ) Pimpla, sp. ? 

A commonplace species of Pimpla is now common on Oahu, where it was first 
noticed in 1901. 

Pimpla hawaiiensis Cam. 

This species, described from the islands, also inhabits Mexico, and I have bred it 
from pupae of Mexican Tortricidae. It is no doubt an importation into the islands. 



HYMENOPTERA 677 

Glvptogastra Ashm. 

(i) Glyptogastra aslimeadi, sp. nov. 

Black, the apex of the clypeus, the labrum, antennae, front legs except a large part 
of the coxae, the middle tibiae, the sides (more or less) and the apical margins of the 
second to sixth abdominal segments, the whole or nearly the whole of the seventh and 
eighth and the genital armature ferruginous or testaceous. Middle and posterior femora 
red, the hind tibiae obscure red, as also the middle and hind trochanters, the coxae and 
tarsi black or blackish. Apical half of the antennae blackish above, beneath paler, but 
sordid. Abdomen dull, metallic greenish black. 

Face below the antennae and nearly the whole thorax conspicuously clothed with 
pale brownish yellow hairs ; antennae with 32 joints and with the first funicle joint much 
longer than the second, the rest decreasing in length towards the apex, each joint sub- 
compressed so as to be wider at its base and apex than in the middle. Thorax dull and 
densely sculptured beneath the pubescence, formed as in G. haivaiiensis Ashm., the 
propodeum with very shallow punctures, which are larger than those of the mesonotum, 
the posterior median impression without hairs. Elevation of first abdominal segment 
concave above, second to sixth with lateral oblique lines, meeting at their apices the 
more or less distinct transverse impressed lines, which are distinct only towards the 
sides of each segment. Apical three or four segments recurved. Wings uniformly 
light shining fuscous, the neuration and stigma dark. Length (abdomen not straightened 
out) 9 mm. 

Obs. G. hawaiiensis is not simply black as described, but is distinctly metallic. 

Hab. Oahu ; Honolulu, 1500 ft. I have not seen the female. 

Subfam. OPHIONINAE. 

Enicospilus Curt. 

The Hawaiian species of Enicospibis present unusual difficulties owing to the 
variation of important structural characters, as well as of colour. In some species the 
entire absence of the blister-like spot on the front wings decharacterizes the genus. It 
is, however, quite impossible to separate such species generically, because these spots 
are subject to variation in other species, so that in some cases, while the blister spot 
is tolerably distinct in some examples, in others it may be almost or entirely effaced. 
The transverse carina of the propodeum is also variable. Thus in E. kaalae Ashm. 
the male (as in the type) may be without the carina, or it may be well developed, 
as it always is in the female. 



87—2 



678 FAUNA HAWAIIENSIS 

Enicospilus kaalae Ashm. 

Enicospilus semiriifus Perkins, Tr. Ent. Soc. London, 1902, p. 142. 

My E. semirufus is a variety of the above. I characterized it as distinct from 
the fact that the male of E. kaalae was said to resemble the female. In the type, 
however, the male of E, kaalae lacks the distinct carina of the female. The colour 
description likewise did not altogether agree. The examination of a large number of 
examples shows that the carina on the propodeum of E. kaalae may be present or absent 
and is not a specific character. 

Hab. Oahu ; very common throughout the island. The type is from Kauai. 

( I ) Enicospilus tyrannus, sp. nov. 

Male, head and face yellow, the ocelli very large, dark brown ; the mandibles and 
mouth-parts brownish, the impressions on each side of the clypeus and a median spot 
beneath the antennae brown. Antennae ferruginous or testaceous, towards the apex 
fuscous. Pronotum mostly yellowish, the mesonotum dark brown, bordered with 
yellowish, and with a yellow longitudinal line on each side of the middle, dividing the 
dark colour into an anterior and two postero-lateral lobes ; scutellum and postscutellum 
brown medially, more yellow laterally, propodeum blackish pitchy ; more brown on the 
smooth anterior area. Legs ferruginous, the tibiae and tarsi paler. Abdomen dark 
brown or dark fuscous, in parts more reddish brown, especially in lateral aspect. 

Female with the inner orbits widely pale yellow below the antennae, narrowly so 
above ; middle of the face brown, the clypeus more yellowish, palpi and mandibles 
testaceous or ferruginous, the latter with dark teeth, occiput mostly pale yellow. 
Pronotum dark brown with a yellow area on each side, mesonotum blackish fuscous, the 
lateral margins in front and a space in the middle posteriorly, yellowish or reddish ; 
scutellum at sides yellow, more testaceous above ; propodeum black. Legs with the 
coxae black or very dark brown, the femora brown, tibiae and tarsi ferruginous. 
Abdomen dark blackish fuscous, a little red-tinged in parts. 

Antennae of male extending well beyond the apex of the spread wings, in the 
female reaching about to the apex, the eyes very large, leaving only a very short cheek 
between the eyes and mandibles. Mesonotum dull, with very fine and dense rugulose 
puncturation, more pubescent in the female ; propodeum nearly smooth and somewhat 
shining in front in the male, and at the most with faint traces of a carina dividing off the 
posterior finely rugose portion ; in the female the carina is distinct, the part in front of it 
finely and densely rugose, but less dull and more finely sculptured than the posterior 
portion. Wings distinctly yellowish-tinged in the male, still more conspicuously so in 
the female ; stigma of male brown, of female dark brown, discocubital cell with two 
distinct yellowish spots, the basal larger and of triangular shape. Length ^ 1 7, 
$ 22 mm. 

Hab. Molokai, 4000 ft. 



^ 



HYMENOPTERA 679 

(2) Enicospilus capnodes, sp. nov. 

Male black, or in parts blackish fuscous, labrum testaceous, mandibles except the 
tip reddish, the inner orbits narrowly, the posterior widely, yellow ; second joint of 
trochanters and apex of first joint ferruginous. Wings evenly fuscous ; though trans- 
lucent, and not very dark, they are darker than in most of the Hawaiian Enicospilus. 

Head with the eyes very large, only separated from the mandibles by a very 
narrow cheek, the face below the antennae densely and finely punctate, the vertex 
smooth and shining between the large ocelli ; antennae very long, reaching far beyond 
the apices of the expanded wings. Mesonotum very minutely punctured, the propodeum 
clothed with short pale pubescence, slightly shining in front in some aspects, the trans- 
verse carina represented only in the middle by a short line or tubercle, behind this closely 
rugose, dull and much less smooth than in front of it. Discocubital cell of front wings 
with a small dark spot, showing signs of total obsolescence, the radius thickened at the 
base, transverse median nervure not interstitial with the basal nervure, first recurrent 
not interstitial with the second transverse median nervure, but the space between them 
is very short, one-fourth the length of the second recurrent nervure. Abdomen very 
finely punctured or appearing granulate and clothed with pale fuscous pubescence. 
Length 14 mm. 

Hab. Hawaii ; Kona, 3000 ft. 

(3) Enicospihts dispilus P. 
Enicospihis dispilus Perkins, Tr. Ent. Soc. London, 1902, p. 143. 
Hab. Oahu ; abundant in the mountains. 

(4) Enicospilus dimidialus P. 
Enicospilus dimidialus Perkins, loc. cit. supra. 
Hab. Oahu ; common in the mountains. 

Athvreodon Ashm. 

Abanchogastra Perkins, Tr. Ent. Soc. London, 1902, p. 141. 

My genus Abanchogastra is synonymous with the above. A serious misprmt or 
error, of which there are several in Ashmead's table of the genera of Ophionini, misled 
me into supposing my genus undescribed. 

Atkyreodon liawaiiensis Ashm. 

Athyreodon kazoaiiensis, Ashmead huj. op. i. p. 343- 

A colour variety, separable by no structural character, so far as I can see, is found 

on Maui. 

Hab. Maui, West Maui, and probably general, if looked for specially. 



68o FAUNA HAWAIIENSIS 

(i) Athyreodon debilis P. 
Abanchogastra debilis Perkins, Tr. Ent. Soc. London, 1902, p. 141. 
Hab. Oahu ; mountains round Honolulu. 

Banchogastra Ashm. 

(i) Banchogastra vitreipennis, sp. nov. 

Female black, the front tarsi somewhat brownish, the wings hyaline, iridescent, 
only slightly infuscate. 

Mesonotum very densely and finely punctured, the propodeum dull and rugose 
behind the transverse carina, in front of this smoother and more shining-. Abdomen 
narrow, the second segment nearly parallel-sided, elongate, one and a half times (or 
more) as long as its basal width. 

Closely allied to B. nigra but distinguished by its comparatively clear wings and 
long second abdominal segment. Length 9 — io'5 mm. 

Hab. Maui ; Haleakala, 5000 ft. and upwards. 

Pycnophion Ashm. 

( 1 ) Pycnophion fuscipennis, sp. nov. 

Rufous, the head, a median stripe on the mesonotum and the abdomen black. 
Legs mostly black, but the coxae for the most part (the hind ones being more or less 
dark) are red, as well as the extreme apex of the trochanters and base of femora of the 
middle and hind legs. Posterior orbits narrowly yellow and the anterior ones to the 
height of the antennal insertions. Wings shining fuscous, and iridescent. 

Eyes very large, there being a short cheek between these and the mandibles, 
the first funicle joint of antennae nearly as long as the next two together. Mesonotum 
dull, finely punctate, the propodeum without a transverse carina, irregularly finely rugose. 
Ovipositor rather short, extruded to about the length of the three preceding dorsal 
segments. Female. Length 9 mm. 

In this species the first recurrent nervure is almost interstitial with the discoidal. 

Hab. Kauai, 3000 ft. and at lower elevations. 

Pristomerus Curtis, 
(i) Pristomerus hawaiiensis, sp. nov. 

Male black, the two basal joints of the antennae, the mandibles, palpi, apex of 
clypeus, all the legs (except the tips of the tarsi, the apex of the hind tibiae and more or 
less of the hind coxae) yellow, brownish yellow, or testaceous. Scutellum and middle 
of mesonotum on its posterior half red, this colour continued forwards along the furrows 



» 



HYMENOPTERA 68 1 

and in front at the sides to the tegulae ; upper edge of anterior and posterior orbits 
narrowly red, with a dilatation of this colour on the vertex. Basal abdominal segment 
red at the base for half its length or nearly, second segment red-marked at extreme base 
and at the apex, second and third for a large part or wholly red, claspers testaceous. 

Scape of antennae short, not or hardly longer than its greatest width, first and 
second flagellar joints subequally elongate. Mesonotum dull, shallowly but distinctly 
punctured on its posterior half, the punctures becoming effaced anteriorly ; scutellum 
shining and punctate, the deep transverse impression in front of it longitudinally rugose; 
propodeum strongly areolated, the areoles of the posterior surface rugose. Wings 
slightly yellowish-tinged, neuration yellowish fuscous, stigma very large, triangular, 
fuscous. Basal abdominal segment finely longitudinally rugose on its apical half, second 
segment very finely so, third excessively finely rugulose at the base laterally. Hind 
coxae with a strong tooth beneath near the middle, between which and the apex are 
several minute teeth. 

Female generally like the male, but the basal antennal joints are dark brown or 
infuscate, the orbits and thorax wholly black, the hind coxae nearly entirely black 
or very dark brown, the hind femora nearly wholly pitchy brown, and much more slender, 
the tooth is more slender and placed nearer to the apex of the joint, the minute teeth 
following it being absent or hardly visible, the third, fourth and fifth abdominal segments 
with narrow red apical margins. Ovipositor nearly as long as the abdomen. Length 
5"5— 7"5 mm. 

Hab. Hawaii, widely distributed at an elevation of looo — 2000 ft. 

(2) Pristomertis hilaris, sp. nov. 

Very closely allied to P. hazuaiiensis, but the J is easily distinguished by having 
the whole thorax red, excepting more or less of the propodeum ; the antennae have the 
basal flagellar joints pale, as well as the scape. In some examples the propodeum is 
nearly entirely black, in others it is half red. The basal abdominal segments are a little 
more finely longitudinally rugose, and the metapleura above the hind coxae less strongly 
punctured. Length $ 7 mm. 

I have not seen a female of this species, which replaces the preceding on Oahu, 
Maui and Molokai, but may prove to be merely a variety and not a good species. 

Hab. Oahu, Molokai and Maui ; 1000 ft. or more above sea-level. 

Atrometrus Forst. 

(i) Atrometrus tephrias, sp. nov. 
Male with the whole head, excepting the occiput and a broad median band 
extending from this to the antennae and enclosing the ocelli, lemon-yellow ; scape, 
pedicel and base of flagellum more or less pale beneath ; the whole of the prothorax. 



682 FAUNA HAWAIIENSIS 

the mesosternum and mesopleura, excepting a spot beneath the tegulae, and the 
scutellum, pale yellow. Thorax above for the most part blackish, the margins of the 
mesonotum, a sjsot on the disc, one on the postscutellum and at the anterior margin of 
the propodeum, as well as a band on the sides of the latter, all pale in colour, red, 
testaceous or yellow, but never bright yellow like the head. Abdomen beyond the 
second segment brownish, with black bands. Middle and front legs brownish yellow, 
hind legs pitchy brown, the coxae yellow, marked with black, the trochanters also more 
or less pale. Wings very iridescent, neuration blackish. 

Female with head markings as in the ^, but the pale colour, is more testaceous and 
the sides of the head behind the eyes largely blackish. Prothorax reddish, mesonotum 
red with a median anterior and two lateral dark areas, mesosternum black, the pleura 
dark, but partly reddish, especially posteriorly, scutellum and propodeum red, the 
former paler ; legs much as in the male, the hind pair generally darker than the others, 
the coxae brownish. Basal joints of antennae more or less pale, especially beneath. 
Basal two segments of abdomen black or nearly so, the rest obscure brown. 

Head above the antennae finely and densely punctate or coriaceous ; thorax also 
very finely and densely so ; propodeum closely irregularly rugose, with short and rather 
dense, pale pubescence. Antennae of female with about 24 joints, of ^ 25 or 26 joints. 
Length 6 mm. 

Hab. Oahu ; not rare at about 1500 ft. in the mountains. 



(2) Atrometrjts sociabilis, sp. nov. 

Black, the whole face below the antennae, the scape of the antennae beneath, the 
orbits for a short distance above the yellow of the face, the tegulae, the anterior and 
intermediate coxae and trochanters, sulphur yellow or pale yellow. The pronotum is 
reddish or yellowish in the middle above, more or less blackish or infuscate at the sides, 
where the posterior and inferior angles are both paler, being either yellow or testaceous. 
Mesonotum and propodeum entirely black. Front legs, except the still paler coxae and 
trochanters, clear testaceous, the hind coxae black, yellow at the tips, the femora pitchy, 
the tibiae and tarsi sordid, brownish yellow. Abdomen black, or in part brownish- 
tinged, third and following segments ventrally yellowish brown. Neuration dark brown 
or blackish. Antennae with the small ring joint 22- or 23-jointed in either sex. Head 
above the antennae very densely and evenly punctate, propodeum densely irregularly 
rugose, and pubescent, mesonotum excessively finely and closely punctured. Length 
5 mm. Apparently allied to A. flavifrons Ashm. 

Hab. Hawaii ; Kilauea. 



HYMENOPTERA 6S3 

(3) Atroinetus satelles, sp. no v. 

Male black, the face beneath the antennae bright yellow and this colour is continued 
upwards for a very short distance along the orbits ; posterior orbits black, scape of 
antennae yellow beneath, the following two short joints and the next long joint also 
yellowish brown ; seen from above all these joints are also pale, but more sordid. 
Pronotum, tegulae, front and intermediate coxae and trochanters pale yellow, scutellum 
in front with a reddish mark, becoming yellow at its anterior angles ; a reddish spot on 
the postscutellum ; front and middle legs, except as above noted otherwise, clear 
testaceous, a minute yellow spot at the apex of the mesopleura; posterior coxae yellow, 
with a dark stripe outwardly and inwardly ; first joint of trochanters blackish, second 
yellow above ; femora pitchy brown, tibiae and tarsi more or less sordid yellowish. 
Abdomen pitchy, third, fourth and fifth segments brown, with a dark apical band. 

Antennae 23-jointed, head above the antennae smooth, the sculpture nearly 
effaced, propodeum finely granulate or shagreened, a little rugulose posteriorly. 
Length 5 mm. 

Hab. Hawaii ; Kilauea. 

(4) Atro7nettis solitaries, sp. nov. 

Male black, the face below the antennae, the whole orbits except for an interruption 
on the vertex, the whole prothorax, the tegulae, the anterior and intermediate coxae and 
trochanters, a mark more or less reddish on the scutellum, a band (reddish anteriorly) 
on the mesopleura, sulphur yellow. First three antennal joints (counting the minute 
ring joint) brown beneath, darker above. Front and intermediate legs (except as above 
noted otherwise) clear yellowish brown, the tarsi of the latter darker ; posterior legs 
with the coxae reddish, with a black stripe on each side, basal joint of trochanters black, 
pale at the tip ; femora pitchy brown, pale at extreme base ; tibiae and tarsi blackish 
pitchy. Abdomen blackish, after the second segment dark fuscous, pubescent and finely 
punctured, the venter yellowish brown. Antennae 24-jointed, with the minute annular 
joint. Head above the antennae dull, densely punctured ; mesonotum extremely finely 
punctured, the propodeum densely rugose. Length 5 mm. 

Hab. Hawaii ; Kilauea. 

Fam. BRACONIDAE. 

All Hawaiian Braconids are no doubt of foreign origin, e.xcepting Ecphylopsis and 
possibly Ischiogonus palliatus Cam. Of recent introduction are a species oi Brae on, one 
or two of Macrodyctium, several species of Apaiiteles, three species herewith described, 
and a few still undetermined. Chelonus blackbui-ni Cam. is also found in Australia, 
while Phanej-otoma hatvaiiaisis x\shm. was probably introduced by Koebele (? from 
Japan). 

F. H. II. 88 



684 FAUNA HAWAIIENSIS 

Subfam. MICROGASTERINAE. 

Apanteles Forst. 

Apanteles, spp. 

Several species have become established, one is known to me from North America 
and one is parasitic on caterpillars of domestic Lepidoptera. 

Subfam. BRACONINAE. 
Bracon Fabr. 
( I ) Bracon, sp. ? 
Hab. Oahu, at and near Honolulu. 

Macrodvctium Ashm. 
(i) Macrodyctinm omiodivoruvi Terry. 

Macrodycthmi omiodivoruvi Terry, Bull. H.S.P.A. Exp. Station. 
A very variable species. 
Hab. All the islands. 

Habrobracon Ashmead. 

(i) Habrobracon, sp. 1 
Hab. Oahu and probably the other islands; often found in houses. It attacks 
caterpillars living in beans etc. 

Subfam. RHOGADINAE. 

IscHioGONUs Wesm. 

(i) Ischiogomis pallidiceps, sp. nov. 

Female black, the head yellowish brown, the tegulae and legs pale testaceous, basal 
joints of antennae also more or less pale, apex of mandibles dark. Antennae with the 
second joint of flagellum shorter than the first, the head smooth polished. Mesonotum 
shining in front and at the sides, in the middle more dull and with indefinite rugulose 
puncturation ; scutellum at the base longitudinally costate ; propodeum with a median 
raised line or carina, the anterior areas separated by this carina, rugulose ; areola and 
petiolar areas confluent, and more strongly rugulose ; pleural area marked off by a very 
fine longitudinal carina. Wings hyaline, iridescent, nervures brown, the extremities of 
the nervures at the base of the stigma yellowish. Basal abdominal segment coarsely 
rugose, second longitudinally strigose, dull and sculptured between the strigae, third 
shortly strigose at base, the rest smooth and shining, ovipositor exserted to about the 
leneth of the abdomen. 



■ HYMEN OP TERA 685 

Male agrees generally in structure with the female, but is pale brown in colour, 
the basal or more than one of the abdominal segments often darker, sometimes the 
whole abdomen of a darker hue than the head and thorax. Length 3 — 5 mm. 

Hab. Oahu ; appeared first rather numerously in 1900, since which time it has 
been very common. It attacks various Longicorn Coleoptera, notably the species of 
Clytarlus and Plagithmysus. I have seen a specimen, apparently identical, from 
New Zealand. 

Subfam. SPATHIINAE. 

HoRMius Nees. 

(i) Hormius{^.) peregi'inus, sp. no v. 

Head and thorax ferruginous or reddish brown varying in depth of colour, the rest 
of the thorax darker, blackish, more or less tinged with red, basal and usually the apical 
abdominal segments dark, the second segment always pale yellowish, legs testaceous, 
the hind femora more or less dark at least on the apical portion outwardly, their tibiae 
also dark except on the whitish basal portion, antennae testaceous basally, becoming 
dark apically, ovipositor black on the apical third or less, the rest pale. 

Head dull, transversely rugose posteriorly ; mesonotum very finely punctured, dull, 
in the middle posteriorly with some more or less longitudinal wrinkles, scutellum with 
a few strong costae anteriorly ; propodeum with a close more or less reticulate rugose 
sculpture like that of the first abdominal segment, the areolation effaced ; stigma dark 
fuscous, pale at base and sometimes at the apex. Hind femora exceedingly robust and 
incrassate, armed beneath with teeth, the basal of these placed before the middle, a 
longer one is placed nearly at the middle and is closely followed by about five or six 
others which decrease in size towards the apex of the femora. Basal abdominal 
segment subparallel-sided, about twice as long as wide in the male, less elongate in the 
female, only about half as wide as the apex of the second segment, which like the 
following is only very minutely sculptured. Ovipositor longer than the abdomen. 
Length 3-5— 4'5 mm- 

Hab. Oahu, Honolulu ; of recent introduction. 

Obs. I am unable to place this insect generically with any certainty. It appears 
to show some affinity with the Helconinae. 

Spathius Nees. 

(i) Spathius perdebilis, sp. nov. 

Male brown, the propodeum, petiole of abdomen and its apical segments, the 

lateral lobes of the mesonotum more or less, dark brown or infuscate, head paler 

yellowish brown, legs yellow, as also the several basal antennal joints, the other joints 

infuscate. Neuration mostly pallid, the stigma fuscous, pale at base. 

88—2 



686 FAUNA HAWAITENSIS 

Head shining, microscopically rugulose, first flagellar joint of antennae longer than 
the second. Mesonotum dull, densely and minutely punctate or shagreened; propodeum 
with a delicate median carina in front, the surface finely shagreened, the petiolar area 
defined, very narrow, but dilated or somewhat diamond-shaped anteriorly. Petiole long 
and very narrow, many times as long as wide, dull, densely and finely shagreened, its 
upper edge in profile well curved ; rest of abdomen polished and smooth. Length 
2 '5 mm. I have not seen the female. 

Hab. Oahu, Honolulu, introduced, without doubt. 



68; 



Supplement by R. C. L. Perkins 

to 

ORTHOPTERA. 

Vide pp. I — 1 6 hujus tomi. 

ACRIDIOIDEA. 
Atractomorpha Sauss. 
( I ) Airactomorpka crenaticeps Bl. 
Hab. Oahu ; now common round Honolulu; first appeared about 1900. 

LOCUSTOIDEA. 

HOLOCHLORA Stal. 

(i) Holochlora veitosa. 

Hab. Oahu ; the eggs inserted in young shoots of trees have been frequently 
found since 1S96, but the mature insect is still rarely met with. 

The determination of the above species and of the Mantid named below, were, 
I believe, kindly made by the authorities at the Dept. of Agriculture in Washington for 
some of the economic entomologists in the islands. 

Braciivmetopa Redt. 
Brachymetopa blackburni de B. 

Brachynietopa discolor Redt., Perkins, Fauna Haw. 11. p. 9. 

In the Part ' Orthoptera ' of the Fauna I have referred to the peculiar case of 
B. discolor and B. blackburni, but at the time of publication of that Part I had not 
sufficient evidence to warrant my uniting these as forms of one species. There is no 
doubt that they merely represent a striking case of dimorphism. Some years ago 
I spent two days investigating the matter and carefully examining every mature insect, 
that I could find. Of twenty-six adult Zi 24 belonged to the van discolor, the brownish- 
coloured black-faced form. Two males only were the green blackburtii form. One of 
the latter was so far intermediate as to have a large triangular brownish or darkened 
area on the face, this marking being characteristic of the distinct Oahuan species 
B. unica P. 



688 FAUNA HAWAIIENSIS 

Of twenty-one females, one only was of the discolor var. but one or two of the 
other twenty (typical green blackbtirni) showed the triangular dark facial area as in the 
t above referred to. In reality this variety can hardly be considered as a true 
intermediate form, but rather as a third constant variation, rare in either sex. 
It is certainly interesting to find this peculiarity of marking occurring as a constant 
specific character in B. tmica of which I have, since its description, examined 
additional specimens. 

It would appear then that about one ^ in 13 of ^. blackburni is green, typical or 
subtypical, the others being var. discolor, while of the females one in twenty-one only 
is var. discolor the others being the green form. Consequently there is exhibited a 
sexual dimorphism in B. blackburni, which is not yet quite complete, while a third form 
is of uncommon occurrence in either sex, in some respects forming a slight intermediate 
condition between the blackburni and discolor forms and resembling in colour another 
species of the genus, very distinct from and much rarer than blackburni but occurring 
with it. 

I ought to add that all the specimens here considered were taken in the same 

locality. 

GRYLLOIDEA. 

(i) Gryllus, sp. ? 

A commonplace species of Gryllus appeared in Honolulu in 1901 and has now 

spread into the country. 

Paranemobius Alfk. 

(i) Paranemobius schauinslandi Alfk. 

What I take to be this species is an evidently' imported species, found in dirty and 
dusty houses, in closets, cellars, etc., in Honolulu and elsewhere. 

Leptogryllus p. 
(i) Leptogryllus cylindricus, sp. nov. 

Testaceous or ferruginous, becoming often more or less discoloured after death, the 
front tibiae and femora with faint fuscous spots or annulations, apex of first, the second 
and apex of third joints of hind tarsi and sometimes of the middle ones infuscate. Tip 
of ovipositor blackish, with numerous small teeth on each side, the apical ones the 
larger. Hind femora very slender, with many small fuscous spots outwardly. Tegmina 
squamiform, about covering the metanotum in the male, smaller in the female, their 
inner margins not nearly touching the middle line. Ovipositor of the female very long, 
considerably longer than the hind femur. Length of body without ovipositor 15 mm., 
ovipositor 13 mm., hind femur 10 mm. 

Hab. Kauai, about 2500 ft. 

' An examination of the figures of P. schauinslandi has since convinced me that it is not the household 
insect above referred to, and it is still unknown to me. R. C. L. P. 



OR THOP TERA 689 



(2) Leptogryllus apicalis, sp. nov. 

Colour, size and form generally as in L. nigrolineatus, the tegmina in the male 
covering nearly all the metanotum. the posterior femora dark or black for a large extent 
on the basal portion. Length ^ 18 mm., hind femora 12 mm. 

Hab. Molokai and Maui (3000 ft). 



(3) Leptogryllus deceptor, sp. nov. 

Extremely like L. nigroliHeaius having the same colouration but distinguished at 
once by the small laterally placed tegmina, leaving the metanotum mostly exposed, and 
the ovipositor is apparently longer. Female. Length of hind femora 12 mm.; 
ovipositor 9 — 10 mm. 

Hab. Oahu, 1500 — 2000 ft. 



Prognathogryllus Brunn. 

The genus Nesogryllus is the male of Prognathogrylltis and was originally separated 
largely on account of the quite different cerci. It is noteworthy that in the allied genera 
there is no such sexual difference in these organs. The species are difficult and 
specimens from the other islands are so close to the Kauai and Oahu forms, as to be 
inseparable without a good series of specimens, especially as those taken in numbers 
appear to show considerable variation. 



MANTOIDEA. 

A second species of this group has been imported to the island of Hawaii, where it 
is now common in some localities and spreading. 



Paratenodera Rehn. 
(i) Paratenodera sinensis Sauss. 
Hab. Hawaii ; now common on the windward side-; first taken in 1900. 



690 FAUNA HAWAIIENSIS 

BLATTOIDEA. 

Panchlora Burm. 

(i) Panchlora inaderae F. 

I omitted this species in my earlier paper on the Orthoptera, although it was 
known to me that it had occurred in Maui. It has since been found on Oahu, Molokai 
and Hawaii, and is no doubt generally distributed. 

DERMAPTERA. 

At least three species of earwigs have been imported and become established since 
my former list was made up. They are of little importance for the purposes of this 
work. The most conspicuous is a species of Labidura. 

Labidura Leach. 

(i) Labidtira icterica Serv. 

Hab. Oahu, Honolulu and in the country. It occurs as high as looo ft. or more 
in the mountains also. 



691 



Supplement by R. C. L. Perkins 

to 

NEUROPTERA 

Vide pp. 31 — 89 hujus tomi. 

Fam. HEMEROBIIDAE. 

Subfam. CHRYSOPINAE. 

Anonialochrysa reticulata P. 

At Kilauea, Hawaii, I ha\e taken specimens of an Anonialochrysa agreeing well 
with A. reticulata, but they are rather small and, as in many other species, there is 
some variation in the neuration. The male characters are remarkable. The apical 
dorsal plate is fringed with hairs and bears a specially developed stout spinose one on 
each side at the base ; the apical ventral valve is hairy beneath, subtriangular, being 
subacute at the apex, which forms a recurved hook. The abdomen has the clothing 
normal, the hairs not directed basally, as in A. f rater P., which it much resembles. 

Hab. Hawaii, Kilauea. 

Anonialochrysa rufescens McL. 

I suspect that my A. biseriata is the female of A. rufescens McL., the type of 
which I have not seen. 

Subfam. HEMEROBIINAE. 

(i) Nesomici'onins angtilaris, sp. nov. 

Head pale, yellowish, clothed with pale hairs, the pronotum also pale, but dark- 
spotted at the sides and thereon with dark hairs, rest of thorax pale but more or less 
dark in parts, abdomen brownish, with pale hairs. Antennae pale yellowish. Anterior 
wings light brown mottled with darker and with a large triangular dark brown or 
blackish fuscous triangular area on the basal half, connected with dark spaces on the 
costa, apical margin whitish hyaline, this pale area interrupted by a dark space a little 
above its middle, neuration pink in the pterostigmatic region. Posterior wings hyaline 
with dark fuscous markings along the outer gradate nervures and beyond these, with 
a good deal ot tuscous strigulation ; pterostigmatic nervures and many of the others 
pink. Front and middle tibiae annulate, the hind ones very long, pale, subfusiform 
in shape. 

F. H. II. 89 



692 FAUNA HAWAIIENSIS 

Anterior wings with emarginate apical margin, the costa bisinuate, the dorsal 
margin rounded, seven to nine radial sectors, hind wings with apical margin distinctly 
concave. Apical dorsal appendages of abdomen long but not extending far beyond the 
ventral plate, which is triangular, between the appendages two chitinous spines crossing 
one another. Expanse about 16 mm. 

Hab. Molokai, 4500 ft. 

(2) Nesomici-07ims phaeostictus, sp. nov. 

Brownish {after drying) the head above yellow, clypeus and middle of face brown, 
antennae pale yellow, the joints fuscous-ringed ; front and middle tibiae conspicuously 
dark-ringed. 

Wings hyaline, the front pair with four small but conspicuous dark spots on the 
basal fourth of the costa, and nearly equidistant one from another, three similar spots 
below these on the radius, three nearly equidistant dark blotches on the costa beyond 
the small spots, and with one or two other small spots between each of these blotches, a 
transversely elongate spot beyond the latter, a spot at the tip of the wing, sometimes 
connected with the last of the costal blotches, two elongate dark spots on the apical 
margin. For the rest the wings are strigulated with brown or fuscous with darker spots 
on the cubitus and the gradate nervules ; the nervures bear conspicuous light and dark 
hairs, the former more numerously. Hind wings with seven or eight dark spots on the 
costa, a dark suffusion in the costal cells beneath the three or four basal spots, and a 
second suffusion beneath two of the other spots ; a large smoky cloud on the lower part 
of the apical margin and a small one, distant from this, on the dorsal. 

Front wings falcate, the apical margin conspicuously concave, the dorsal margin 
nearly straight or faintly concave on the basal half; radius with about seven sectors. 
Hind wings with the apical margin distinctly concave. Female : expanse about 16 mm. 
Allied to the preceding species. 

Hab. Oahu, 1 500 ft. Koolau range. 

(3) Nesomicronnis ombrias, sp. nov. 

Dark brown or blackish, the lower part of the face testaceous, a pale marking on 
the head above, some spots on the pronotum, the mesonotum in the middle in front, 
yellowish. Basal two joints of the antennae brown, the following testaceous. Legs 
pale, yellowish, the front and middle tibiae with two dark wide rings, the hind pair very 
long, dilated after the basal third, with a more or less dark cloud near the base and 
another at the apex ; femora also more or less infuscate at apex, apex of abdomen 
yellowish. 

Front wings darkish brown or fuscous, mottled over most of their surface with pale 
spots or strigulations, interrupting the dark nervures with pale spaces, the gradate 



NEUROPTERA 693 

nervules dark and with darker suffusion at their sides, especially the outer series. The 
dilated costal cells form the clearest part of the wing, beyond the outer gradate nervules 
the mottling is sparse but there is a marginal series of five or six conspicuous pale spots, 
widely separated from one another, extending from the apical on to the dorsal margins. 
There is a large basal area between the radius and dorsal margin of a generally darker 
colour than the rest of the wing, forming a large blotch, angulated below and more or 
less curved above, where it is broken by pale spaces about the cubital nervures. About 
its angle it is uniformly dark. Hind wings smoky hyaline, iridescent, with a con- 
spicuously darker smoky area along the dorsal margin, on the basal third or fourth of 
the wingf length. 

Front wings rounded at the tips, apical margin not at all excised, costa strongly 
curved on its basal part, where the costal cells are unusually high. Six sectors to the 
radius. Apical segment of the abdomen raised or somewhat bent back, angulately 
emarginate in dorsal aspect, the appendices in side view elongate triangular, bluntly 
pointed, conspicuously clothed with curved hairs, extending far beyond the apical 
ventral segment. Male: expanse 15 mm. 

Hab. Oahu ; mountains near Honolulu, 1500 ft. 



ODONATA. 
Fam. AGRIONIDAE. 

The Hawaiian species referred to the genus Agrion, I believe, are none of them 
really referable to that genus, and the species though apparently all allied to one another, 
yet themselves form a number of groups. Probably in the hands of a specialist the 
whole will form a peculiar genus, with several subgenera. I have previously spoken of 
the great variability exhibited by the various species, and were it not for the characters 
shown by the terminal appendages of abdomen in the $, the difficulty of separating the 
species would be intense. Having now examined many more specimens from the 
different islands, I am the more convinced of the extreme importance of these characters. 
In many species, if one only had the extreme forms, one would certainly consider these 
to form distinct species, but when one has large series, especially from different localities 
and different islands, one finds that the apparently strong distinctions between extremes, 
apart from those afforded by the appendages, all vary and intermediate forms occur. 
Characters usually considered of specific, or even of more than specific importance, 
whether of colour, size, neuration, etc. all vary and one is finally driven to the male 
structures to find constant characters. 

The following tabulation is probably a fairly natural grouping of the species that I 
have examined. It is based on the male characters. 

89—2 



694 FAUNA HAWAIIENSIS 

1. Superior appendages of the $ abdomen short, the inferior ones extending 

behind them ; most of the abdominal segments dark and metallic, and 
at most with a pale band at the base ; lower appendages bidentate at 

the apex or obliquely truncate in side view A. xanthomelas, pacificum, nigra hamatum. 

Without some or all of the above characters 2. 

2. Superior appendages generally shorter than the lower and with a black tooth on 

the inner side towards the base, this tooth being usually directed upwards ; 

abdomen largely red A. oresitrophum, orobates, leptodemas, calliphya. 

Without some or any of these characters 3- 

3. Legs mostly dark brown or black ; abdomen usually black or pitchy and 

metallic, rarely with red segments behind the two basal ones ; insect often 
with more or less whitish pruinosity ; upper appendages in lateral view 

emarginate at apex so as to form two apical processes or teeth 

A. koelense, asteliae, amaurodytiim, eudvium, adyUim. 
Without some or all of these characters 4- 

4. Upper appendages extremely long and forcipate, dilated only at the extreme 

base A. nesiotes. 

Appendages not so formed 5- 

5. Upper appendages turned outwards, their broad surface exposed in dorsal 

aspect ; apical abdominal segment strongly and obliquely raised from 

base to apex A. oahuense. 

Without these characters 6. 

6. Postcostal area of front wings with a single row of cellules, rarely with a few 

divided, to form a double row; abdomen slender or very slender 

A. deceptor, vagabundum, kauaiense, molokaietise, Jugonim. 
Postcostal area of front wings always with a double row of cellules for a 
considerable part ; species of large size and the abdomen relatively stout 
compared with the above species A. oceantcum, blackburni, heierogamias. 

The last three species form the genus Megalagrion of McLachlan, but individuals 
of some of the preceding group might be included therein. 

(i) Agrion culver ti, sp. no v. 

Head dull black, apex of clypeus and labrum, orbits to the height of the antennae, 
mouth-parts and a spot on the basal joint of the antennae yellow or cream-coloured. 
Postocular spots orange, not connected. Prothorax yellow in front and at the sides. 
Dorsum of mesothorax black with purplish metallic lustre, a spot on each side in front 
connected with a sutural line not reaching the hind margin, a spot on each side near this 
margin, a band on the sides of the mesothorax extending back from the middle coxae 
for less than half its length, but represented by a detached spot near the hind margin, 
the whole lower part of the mesopleura extending back from the posterior coxae to the 
hind margin, except a spot posteriorly, yellow. Median longitudinal ridge of mesonotum 
reddish. Legs with the femora and tibiae sordid brownish or fuscous above, beneath 
and the trochanters yellow, the coxae infuscate in part. Abdomen black or pitchy, with 
a green metallic lustre, basal segment yellow at the sides and with very narrowly pale 
apical margin, third to the seventh with narrow yellow basal band, terminal segment 
reddish, but sordid in parts. 



NEUROPTERA 695 

Front wings with about 18 postcubitals, the stigma fuscous, its lower side covering 
one cellule ; three cellules between the quadrilateral and the nodus. Postpterostigmatic 
cellules forming a single row. 

Apical abdominal segment broadly emarginate ; the superior appendages about as 
long as this, dilated on more than their basal half, the dilated part reddish and without a 
distinct tooth on its lower margin apically, at most with a very minute notch in the 
margin just before its apical angle. Seen from above the upper appendages are slightly 
curved inwardly at the tips, the lower ones are red, black at the acute tips, and more 
strongly curved inwards. 

Length of abdomen 35 mm., expanse about 45 mm. Female not known. Colour 
variation not known. 

Hab. Oahu, near Honolulu, 1200 ft. Named after Prof. P. Calvert. 



Agt'ion adytum var. tillyai'di nov. 

Very slender and elongate, head and thorax dull black, the latter with slight 
metallic reflection. Face black, apex of clypeus and labrum and the orbits to about 
the height of the insertion of the antennae white or pale-coloured. Occipital margin of 
the vertex red ; postocular spots wanting. Prothorax with the front margin yellow, as 
also the whole underside ; dorsum of mesothorax with a yellow spot on each side at the 
base, sometimes continued back as a fine sutural line to or nearly to the wings ; sides of 
thorax yellowish, sometirries with a black spot in front of the insertion of the hind wings, 
or with a continuous black longitudinal band in place of this spot. Thorax posteriorly 
reddish. Legs for the most part blackish or brown above, yellow beneath, as also are 
the tarsi for the most part above ; spines long slender and black. Basal two abdominal 
segments black, the first yellowish or red at the base, sides, and along the apical margin, 
the second with red apical margin and yellow on the sides, third segment reddish pitchy 
or pitchy black and this like the preceding sometimes submetallic, the apex still darker, 
fourth, fifth and base of sixth segments either quite red or else pitchy, the apices darker, 
seventh with the extreme base red, eighth entirely red, or only its extreme base red, 
two last segments red, sometimes more or less sordid or infuscate. Appendages red, 
the tips black. Beneath the abdomen is yellow or reddish. 

Front wings with 16 — 20 postcubital cellules before the stigma, the latter wine-red, 
overlying one and part of a second cell, three cellules between the quadrilateral and the 
nodus. Superior appendages t as long as the terminal segment, much dilated basally, 
when viewed inwardly ; in external profile view emarginate at apex to form a large 
superior and small inferior process or tooth, both of which are black, the upper ones 
seen from above a little bent inwards at the tip , lower appendages acute at tips, which 
reach back to the small tooth of the upper ones. 



696 FAUNA HAWAII EN SIS 

The postpterostigmatic cellules do not form a double series as is usual in A. 
eudytum P., though sometimes one or more of them is petiolate. Length of abdomen 
35 mm. ; expanse about 50 mm. I do not know the female. 

Hab. Kauai; near Lihue on a mountain stream. Named for Mr R. J. Tillyard, 
who has done so much work on Australian Odonata. 

(2) Agrion tvillianisoni, sp. nov. 

Agrees generally in size and appearance with the darker example of A. tillyardi, 
but the prothorax is black or dark above anteriorly, and the middle and hind tibiae are 
less dark or embrowned; the second abdominal segment is dark at the sides, unicolorous 
with the dorsal surface ; antepenultimate segment with only its extreme apical margin 
red, the terminal segment, with an apical red band, occupying about half its whole 
surface. Pterostigma fuscous, only a little red-tinged. 

Superior appendages of the male strongly curved inwards, minutely tuberculate at 
the tip, strongly dilated beneath on the basal two-thirds or more of their length, the 
dilatation forming a great prominent angle, but bearing no spine or tooth. Lower 
appendages pale and dilated at the base, black on the apical portion and acute at the 
tips, about as long as the superior appendages. Female not known. 

Hab. Kauai ; in the same locality as A. adytum var. tillyardi. Named for 
Mr E. B. Williamson of Indiana. 



697 



Supplement by R. C. L. Perkins 

to 

DIPTERA. 

Vide Tom. iii. hujus operis, pp. i — 92. 

Fam. PIPUNCULIDAE. 
(i) Pipunctihis vulcanus, sp. nov. 

Head with the front and face black, with inconspicuous fuscous tomentum and in 
the female smooth and shining for half the distance between the ocelli and antennae ; 
posteriorly the head is greyish tomentose, at least towards the sides, but not densely so. 
Antennae entirely dark, black or piceous, third joint acuminately produced at the apex. 
Mesonotum somewhat shining, especially in the male, with hardly noticeable fuscous 
tomentum, and further clothed with short and sparse, but very distinct, erect, black 
hairs ; in the female the sides as far as the tegulae and the anterior margin with a 
narrow border of whitish tomentum ; the pleura with sparse whitish tomentum in both 
sexes ; scutellum with a marginal fringe of bristly hairs, which are more developed than 
the mesonotal ones; metanotum with whitish tomentum. All the legs entirely dark 
above, the femora being black or very dark brown, with short bristly black or spinose 
hairs beneath, the middle ones with a very regular series of longer hairs, curved 
downwards, on their posterior face ; tibiae often less dark than the femora, piceous 
or reddish pitchy, and without special bristles on the hind pair ; tarsi black, or at least 
dark, above. Wings strongly infumate or fuscous in the male, nearly clear in the 
female, third costal segment much shorter than the fourth, posterior cross-vein obliquely 
transverse, its upper extremity about opposite the apex of the second longitudinal, 
terminal segment of fourth distinctly bisinuate ; stigma brown. Abdomen shining on 
the apical segments of the male, very little tomentose except rather obscurely on the 
apical part of the basal segment, which has the lateral series of bristles well developed, 
the rest of the abdomen bearing short black hairs. In the female the abdomen is 
greyish tomentose and clothed with short black hairs, dull, except the fifth segment. 
Length 3'5 — 4 mm. 

Allied to P. jiivator, a variable and widely distributed species, which is found with 
it, and also occurs on other of the islands. Superficially P. vidcmms is most readily 
distinguished by the darker legs. 

Hab. Hawaii, Kilauea. 



698 FAUNA HAWAIIENSIS 

(2) Pipunathts acroihrix, sp. nov. 

Female black, the abdomen shining metallic aeneous-black. Face below the 
antennae with dense white tomentum, that on the head posteriorly more fuscous. 
Third antennal joint clear yellow, the beak thin and long, white. Mesonotum with 
thin fuscous tomentum, dense only at the sides, sparsely and shortly pilose ; marginal 
bristles of scutellum well developed ; metanotum and pleura with denser yellowish grey 
tomentum. Wings long and clear, neuration as in P. vtdcanus, stigma brown, about 
half as long as the fourth costal segment. All the coxae dark, the trochanters and 
tibiae all clear yellow, the femora of this colour at base and apex, the middle being 
black, the tarsi, except the basal joint, being more or less dark above. Posterior tibiae 
with three or four long bristles outwardly near the middle. Abdomen with a distinct 
band of pale tomentum on the basal segment, the rest brightly shining and clothed with 
short hairs, the fifth at the apex with many long black bristles. Length 475 mm. 

Very distinct by the colour of the legs, bristles of hind tibiae, the metallic colour 
and clothing of fifth abdominal segment. 

Hab. Hawaii, Kilauea. Male not known to me. 

(3) Piptmctihis pyrophilus, sp. nov. 

Deep black, more or less shining, especially the male, the very sparse tomentum 
of the surface being mostly fuscous and nowhere conspicuous, except perhaps an apical 
band on the first abdominal segment of the female. Legs in both sexes entirely black, 
or at most a trifle reddish or pitchy at some of the articulations. Antennae entirely 
black ; the face of the female above the antennae not at all shining. 

General appearance that of a minute P. vidcamis, but easily distinguished by being 
even more entirely black in both sexes ; by the erect hairs of the mesonotum, which are 
unusually long, and as well, or almost as well, developed as the scutellar bristles. Wings 
smoky hyaline, differing very little in the sexes, though rather darker in the male. 
Stigma very short compared with the long fourth costal segment. Neuration generally 
as in P. vulcanus. Abdominal hairs black, longer in the female than in the male. 
Length 3 mm. 

Hab. Hawaii, Kilauea. Attached to one or both of the small Delphacid leaf- 
hoppers, that are so common on Raillardia around the crater. 

(4) Pipunculus holoinelas, sp. nov. 

Female entirely black and for the most part shining, the antennae entirely dark, 
piceoVs. Mesonotum apparently without tomentum (? abraded) shining and with very 
short erect hairs, metanotum with greyish tomentum. Halteres pale in the middle, 
more or less dark at base and apex. Legs with the coxae and femora black, the 



DIPTERA 699 

trochanters and some of the articulations alone being more or less yellowish or brown, 
tibiae and tarsi piceous or dark brown above. Posterior tibiae with three or four long 
bristles outwardly about the middle. Wings nearly clear, only slightly smoky, neuration 
much as in P. vuicamis. Abdomen with greyish tomentum on the first two segments, 
the rest deep black and shining, the ovipositor yellow, short and strong. Length 4 mm. 

Hab. Molokai ; readily known from the other species, that have the long bristles 
on the hind tibiae, by the almost entirely dark legs. The male will no doubt have 
darker wings as is usual in Hawaiian species. 

(5) Pipunculus juvator P. 
P.JHvator Perkins, Bull. H. S. P. A. Exp. Station i. 152. 
Hab. Hawaii, Oahu and no doubt other of the islands. 

(6) Pipunculus oahuensis P. 
P. oahuensis Perkins, loc. cit. p. 153. 
Hab. Oahu, 100 — 1000 ft. 

(7) Pipunculus terryi P. 
P. terryi Perkins, loc. cit. pp. 153 and 154. 
Hab. Kauai, Lihue in cane-fields. 

(8) Pipunculus siveezeyi P. 
P. sweezeyi Perkins, loc. cit. p. 1 54. 
Hab. Oahu, 1500 — 1800 ft. not usually common. 

(9) Pipunculus hawaiiensis P. 
P. hawaiiensis Perkins, loc. cit. p. 155. 
Hab. Hawaii, Hamakua, 1200 ft. 

Fam. DROSOPHILIDAE. 

( 1 ) Idiomyia heteroneura, sp. nov. 
Front yellow or golden with a dark transverse mark anteriorly, orbits posteriorly 
and the vertical triangle dark, forming part of a transverse dark band, face yellow, 
antennae reddish yellow, the third joint blackish, the proboscis and palpi yellow. 
Thorax yellowish, two narrow black stripes, well separated, extend on each side of 
the middle from the anterior margin for about two-thirds of the length of the 
mesonotum ; just external to these anteriorly is a largish black spot on each side 
F. H. II. 90 



700 . FAUNA HAWAIIENSIS 

and behind their middle externally on each side there starts another black stripe 
reaching to the scutellum, while between them on the posterior half of the mesonotum 
is a median dark stripe (or perhaps two very closely apposed stripes). Scutellum dark 
on the disc, but broadly bordered with yellow, metanotum mostly dark ; pleura with 
three or four dark spots. Halteres yellow. Legs yellowish, tips of tibiae noticeably 
infuscate. Wings hyaline, their base as far as the apex of the first vein for the most 
part infuscate, the infuscation irregular in depth and distribution, marginal cell infuscate, 
as well as the tip of the wing on either side of the apex of the second, third and fourth 
veins ; transverse veins and a small portion of the fourth longitudinal vein just before 
the apical one of the two transverse veins that are between it and the third, distinctly 
infuscate on each side. This second or apical transverse vein oblique, placed much 
before the posterior transverse, the sector of the fourth vein between these being not 
much shorter than the first named. Abdominal segments hairy, more or less shining, 
black medially and apically, so as to leave a well-marked series of yellow spots on each 
side. Ovipositor reddish yellow. 

Hab. Hawaii. 

(2) Idioviyia silvestris, sp. nov. 

Front golden yellow, the orbits as far as the apical fronto-orbital bristle, the vertical 
triangle and a band connecting these posteriorly, blackish and with grey tomentum ; 
face, palpi and proboscis yellow. Antennae dark, the second joint pitchy brown, the 
arista with about four bristles beneath, the basal jDart being bare, and about nine above. 
Mesonotum yellowish, most of its surface suffused with brown or blackish infuscation, 
forming vague lines and spots, but not a definite pattern ; scutellum dark brownish 
black, metanotum black, tomentose ; mesopleura mostly blackish. Halteres pale 
yellow. Legs yellowish or brownish yellow with the middle and hind femora dark 
brown or blackish, the tibiae also more or less dark. Wings hyaline, basally as far as 
the termination of the first vein largely but irregularly infuscate, marginal cell faintly 
brownish, tip of wing darkly infuscate on either side of veins two, three and four. 
Transverse veins infuscate, as also the part of vein four just before the origin of the 
second or apical of the two transverse veins between it and the third. This second 
transverse vein is oblique and placed much before the posterior cross-vein, so that the 
sector of the fourth that lies between them is not much shorter than this second 
transverse vein itself. Abdomen black, at the base a little pale, pilose and shining, the 
ovipositor paler. 

Hab. Hawaii. 



I 



r 



ZOOLOGICAL RESULTS based on material from New Britain, New Guinea, Loyalty 
Islands and elsewhere, collected during the years 1895, 1896, and 1897, by 
Arthur Willey, D.Sc. Lond., Hon. M.A. Cantab., Late Balfour Student of the 
University of Cambridge. Demy 4to. Parts I, II, and III. Price 12s. 6d. each. 
Parts IV and V. Price 21s. each. Part VI (completing the work). Price 12s. 6d. 

PART I. 1898. 

1. The anatomy and development of Pcripatus novae-britauniae. By Arthur Willey, M.A., D.Sc. 

With Plates I. — IV. and 7 figures in the text. 

2. Mctaprotella sandalcnsis, n. sp. [Caprellidae]. By Dr PAUL MAYER. With 6 figures in the text. 

3. On a little-known Sea-snake from the South Pacific. By G. A. BoULENGER, F.R.S. 

With Plate V. 

4. Report on the Centipedes and Millipedes. By R. I. POCOCK. With Plate VI. 

5. Account of the Phasmidae with notes on the eggs. By D. SHARP, M.A., F.R.S. 

With Plates VII.— IX. 

6. Scorpions, Pedipalpi and Spiders. By R. I. PocoCK. With Plates X. and XI. 

PART II. 1899. 

7. Report on the specimens of the genus Millepora. By Sydney J. HiCKSON, M.A., D.Sc, F.R.S. 

With Plates XII.— XVI. 

8. Report on the Echinoderms (other than Holothurians). By F. Jeffrey Bell, M.A. 

With figures on Plate XVII. and one figure in the text. 

9. Holothurians. By F. P. Bedford, B.A. With figures on Plate XVII. 

10. Report on the Sipunculoidea. By Arthur E. Shipley, M.A. With Plate XVIII. 

11. On the Solitary Corals. By J. STANLEY GARDINER, M.A. With figures on Plates XIX. and XX. 

12. On the postembryonic development of Cycloseris. By J. STANLEY GARDINER, M.A. 

With figures on Plates XIX. and XX. 

13. On a collection of Earthworms. By FRANK E. Beddard, M.A., F.R.S. With Plate XXI. 

14. The Gorgonacea. By Isa L. Hiles, B.Sc. With Plates XXII. and XXIII. 

PART III. 1899. 

15. Orthogenetic variation in the shells of Chelonia. By HANS Gadow, M.A., Ph.D., F.R.S. 

With Plates XXIV.— XXV. and one text-figure. 

16. Enteropneusta from the South Pacific, with notes on the West Indian Species. By ARTHUR 

Willey, D.Sc, Hon. M.A. Cantab. With Plates XXVI.— XXXII. and seven text-figures. 

17. On a collection of Echiurids from the Loyalty Islands, New Britain and China Straits, with an attempt 

to revise the group and to determine its geographical range. By ARTHUR E. Shipley, M.A. 
With Plate XXXIII. 

PART IV. 1900. 

18. On the anatomy of a supposed new species of Coenopsainmia from Lifu. By J. STANLEY 

Gardiner, M.A., Fellow of Gonville and Caius College, Cambridge. With one plate. 

19. On the Insects from New Britain. By D. Sharp, M.A., M.B., F.R.S. With one plate. 

20. Report on the Stomatopoda and Macrura. By L. A. BORRADAILE, M.A., Lecturer of Selwyn 

College, Cambridge. With four plates. 

21. Report on the Slugs. By WALTER E. COLLINGE, F.Z.S., Mason College, Birmingham 

With two plates. 

22. Report on the Polyzoa. By E. G. Philipps, Newnham College, Cambridge. With two plates. 

23. The Hydroid Zoophytes. By LAURA ROSCOE Thornely, University College, Liverpool. 

With one plate. 

24. Astrosclera zvil/ejana, the Type of a new Family of Calcareous Sponges. By J. J. Lister, M.A., 

St John's College, Cambridge. With five plates. 

25. A contribution towards our knowledge of the Pterylography of the Megapodii. By W. P. 

Pycraft, A.L.S., British Museum. With one plate. 

26. The Stolonifera and Alcyonacea. By Sydney J. HiCKSON, M.A., D.Sc, F.R.S., Beyer Professor 

of Zoology in the Owens College, and ISA L. HiLES, B.Sc, Owens College, Manchester. 
With two plates. 

27. Report on the Xeniidae. By J. H. ASHWORTH, B.Sc, Demonstrator in Zoology, Owens College, 

Manchester. With two plates. 

PART V. 1900. 

28. A Description of the Entozoa collected by Dr Willey during his sojourn in the Western 

Pacific. By ARTHUR E. SHIPLEY, M.A. With Plates LIV.— LVI. 

29. On some South Pacific Nemertines collected by Dr Willey. By R. C. PUNNETT. 

With Plates LVII.— LXI. 

30. On the Young of the Robber Crab. By L. A. Borradaile, M.A. With figures in the text. 

31. Anatomy of Neohelia porcellana (Moseley). By Edith M. Pratt, M.Sc. 

With Plates LXI I. and LXI 1 1. 

32. On a new Blind Snake from Lifu, Loyalty Islands. By G. A. BoULENGER, F.R.S. 

With figures in the text. 

33. On Crustacea brought by Dr Willey from the South Seas. By the Rev. T. R. R. Stebbing, F.R.S. 

With Plates LXIV.— LXXIV. 

PART VI. 1902. 

34. Contribution to the Natural History of the Pearly Nautilus. By ARTHUR WiLLEY, D.Sc, F.R.S. 

I. Personal Narrative. II. Special Contribution. 
With Plates LXXV.— LXXXIII., a map and fifteen text-figures. 

CAMBRIDGE UNIVERSITY PRESS 

aonOon; FETTER LANE, E.G. 

C. F. CLAY, Manager 



INDEX 

TO ALL THE THREE VOLUMES 

The arable numerals refer to the page. The roman numerals i, ii and iii refer to the Vol., but roman numerals 

beyond iii refer to the page of the Introduction 
Names of families are in clarendon type, and in the alphabetical arrangement the taxonomical termination idae 

is treated as non-existent 



Abanchogastra Perk, ii 679 
debilis Perk, ii 680 

Ablepharus Fitz. i 368 

boutonii Wieg. i 368 

Acalles Schonh. ii 131 

augusticollis Sharp ii 135 
callichroma sp. n. ii 132 
chlorolepis sp. n. ii 136 
decoratus Blackb. ii 135 
duplex Sharp ii 135 
frater sp. n. ii 133 
humeralis sp. n. ii 132 
ignotus Blackb. ii 136 
innotabilis sp. n. ii 134 
koae sp. n. ii 133 
lanaiensis sp. n. ii 136 
lateralis Sharp ii 131 
leptothorax sp. n. ii 134 
mauiensis Blackb. ii 135 
melanolepis sp. n. ii 132 
nionticola sp. n. ii 138 
nigripennis sp. n. ii 136 
oahuensis sp. n. ii 138 
pallidicollis sp. n. ii 137 
pusillissimus sp. n. iii 653 
tuberculatus sp. n. ii 137 

Acanthaspidae Leth. & Sev. iii 150 

Acanthiidae iii 146, cciii 

Acanthia Fabr. iii 146, xlix 

exulans White iii 146, ii 554 
humifera Kirk, ii 554 
lectularia Fabr. iii 129 
nubigena Kirk, ii 554 
oahuensis Blackb. iii 146, ii 
procellaris Kirk, ii 554 

Acanthocephala ii 434 

Acanthocinides ii 113 

Acarina iii 702 

Achatina accincta Migh. ii 356 
adusta Gould ii 373 
bicolor Jay ii 373 
chrysallis Pfeiff. ii 358 
cochlea Reeve ii 373 
decora Fer. ii 304 
fuliginea Pfeifif. ii 373 
gravida F6r. ii 349 
lorata F^r. ii 303 
luteola F^r. ? ii 346 
newcombi Pfeiff li 374 
oahuensis Green ii 346 
obeliscus Reeve ii 374 

F. H. 



554 



Achatina sandwicensis Pfeiff. ii 368 

sculpta Pfeiff. ii 368 

spirizona Fdr. ii 344 

turricula Migh. ii 374 

turritella F^r. li 346 
Achatinellidae ii 296, xxx, Ixiii, Ixv, Ixx 
Achatinella ii 298 

abbreviata Reeve {^Bulimella) ii 305 

accincta Gould ii 356 

acuminata Gould ii 357 

acuta Newc. ii 349 

acuta Swains, ii 344 

adamsi Newc. ii 314 

adusta Reeve ii 327 

affinis Newc. ii 333 

alba Nutt. ii 303 

albescens Gul. ii 323 

albospira Smith ii 298 

alexandri Newc. ii 348 

amcena Pfeiff. {Frickeila) ii 379 

ampla Newc. {Buliinella) ii 305 

ampulla Gul. ii 318 

anceyana Baldw. {Partuli)ui) ii 311 

anthonii Newc. ii 353 

apexfulva Dixon ii 298 

apicata Pfeiff. ii 299 

apicata alba var. n. ii 299 

aplustre Newc. ii 321 

aptycha Pfeiff. (Newcombia) ii 311 

aptycha Pfeiff. {Partulina) ii 311 

assimilis Newc. ii 334 

augusta Smith {Achat inellastriim) ii 321 

bacca Reeve ii 305 

baileyana Gul. ii 318 

baldwinii Newc. ii 339 

bella Ree\'e {Acliatincllasfruvi) ii 321 

bellula Smith ii 325 

bicolor Pfeiff. ii 299 

bilineata Reeve {Achatitiellastrum) ii 321 

biplicata Newc. ii 334 

boetica Migh. ii 344 

brevis Pfeiff. ii 353 

buddii Newc. {Achati/iellasfntin) ii 321 

bulbosa Gul. ii 351 

bulimoides Swains. {Bulimella) ii 306, 309 

byronii Wood {Bulimella) ii 306 

callosa Pfeiff. {Labiella) ii 358 

canaliculata Baldw. {Newcombia) ii 331 

casta Newc. {Achatinellastrum) ii 322 

castanea Reeve ii 327 

cerea Pfeiff. ii 376 



FAUNA HAWAIIENSIS 



Achatinella cervina Gul. {Achatinellastrum) ii 322 
cestus Newc. ii 300 
chamissoi Pfeiff. {Auriciilella) ii 376 
cinerea var. n. ii 305 
cinerosa Pfeiff. {Buliinelld) ii 302 
cingula Migh. ii 359 
cinnamomea Pfeiff. {Newcombia) ii 331 
citrina Pfeiff. ii 348 
clara Pfeiff. ii 370 

Clementina Pfeiff. {Achatincllasirum) ii 305 
coesia Gul. ii 321 
cognata Gul. ii 322 

colorata Reeve {Achatinellastrum) ii 322 
compta Pease {Pariulina) ii 311 
concavospira Pfeiff. ii 303 
concidens Gul. ii 301 
concinna Newc. ii 348 
concolor Smith {Achatinellastnun) ii 322 
confusa nom. n. {Partiilina) ii 312 
consanguinea Smith ii 325 
contracta Gul. ii 323 
cookei Baldw. {Apex) ii 300 
cornea Newc. ii 335 
corneola Pfeiff. ii 360 
corrugata Gul. ii 309 
corusca Gul. ii 319 
costulata Gul. ii 360 
crassa Newc. {Partiilina) ii 312 
crassidentata Pfeiff. {Achatinellastrum) ii 32 
crassilabrum Newc. ii 335 
crocea Gul. ii 314 
crystallina Gul. ii 361 
cucumis Gul. {Achatinellastnon) ii 322 
cumingi Newc. ii 331 
cuneus Pfeiff. {Achatinellastnun) ii 323 
curta Newc. {Achatinellastrum) ii 323 
cylindrica Newc. ii 335 
decepta C. B. Adams ii 347 
decipiens Newc. {Bulimella) ii 307 
decora Fer. ii 301, 304 
delta Gul. {Achatinellastrum) ii 323 
dentata Pfeiff. ii 365 
deshayesii Mor. ii 334 
diluta Smith {Achatinellastrum) ii 323 
dimidiata Pfeiff. {Leptachatina) ii 359 
dimondi C. B. Adams ii 350 
dimorpha Gul. ii 323 
dolei Baldw. {Partulina) ii 312 
dolium Pfeiff. {Bulimella) ii 302 
dubia Newc. {Partulina) ii 312 
dunkeri Pfeiff. ii 326 
dwightii Newc. {Partulina) ii 313 
eburnea Gul. ii 318 
elegans Newc. {Bulimella) ii 308 
elevata Pfeiff. ii 363 
ellipsoidea Gould ii 336 
elongata Newc. ii 349 
emmersonii Newc. ii 325 
ernestina Baldw. {Achatinellastrum) ii 324 
exilis Gul. ii 361 
extincta Pfeiff. {Laminella) ii 336 
faba Pfeiff. {Bulimella) ii 307 
fasciata Gul. ii 318 
flavescens Newc. ii 337 
forbesiana Pfeiff. ii 300 
formosa Gul. {Achatinellastrum) ii 324 
fragilis Gul. ii 363 
fricki Pfeiff. ii 307 

fulgens Newc. {Achaiinellastrum) ii 324 
fuliginosa (iould. ii 346 
fulva Pfeiff. ii 319 



Achatinella fumosa Newc. ii 362 
fusca Newc. ii 362 
fuscobasis Smith ii 310 
fuscolineala Smith ii 327 
fuscozona Smith ii 321 
fuscula Gul. ii 362 
fusiformis Pfeiff. {Laminella) ii 340 
fusoidea Newc. {Partulina) ii 313 
gemma Pfeiff. {Newcombia) ii 331 
germana Newc. {Achatinellastrum) ii 324 
gigantea Newc. ii 347 
glabra Newc. {Bulimella) ii 307 
glauca Gul. ii 329 

globosa Pfeiff. {Achatinellastrum) ii 304 
glutinosa Pfeiff. {Laminella) ii 363 
goniostoma Pfeiff. {Laminella) ii 333 
gouldi Newc. {Partulina) ii 313 
gracilis Pfeiff. {Achatinellastrum) ii 363 
grana Newc. ii 363 
granifera Gul. ii 356 
gravida F^r. ii 349 
grayana Pfeiff. {Laminella) ii 337 
grisea Newc. {Partulina) ii 313 
grossa Pfeiff. {Laminella) ii 341 
gummea Gul. ii 363 
guttula Gould ii 363 
hanleyana Pfeiff. ii 302 
havvaiiensis Baldw. {Partulina) ii 316 
hayseldeni Baldw. {Partulina) ii 313 
helena Newc. ii 330 
helvina Baldw. {Laminella) ii 350 
herbacea Gul. ii 307 
horneri Baldw. {Partulina) ii 313 
humilis Newc. ii 337 
hybrida Newc. ii 326 
induta Gul. ii 314 
inflata Pfeiff. {Laminella) ii 337 
innotabilis Smith ii 301 
inornata Migh. ii 346 
intermedia Newc. ii 338 
irregularis Pfeiff. ii 338 
johnsoni Newc. ii 321 
juddii Baldw. {Achatinellastrum) ii 324 
juncea Gul. ii 322 
kauaiensis Newc. {Carinella) ii 355 
labiata Newc. ii 365 
lacrima Gul. ii 363 
lactea Gul. ii 319 
lagena Gul. ii 365 

lehuiensis Smith {Achatinellastrum) ii 324 
leucochila Gul. ii 365 
leucophaea Gul. ii 301 
leucorraphe Gul. ii 302 
ligata Smith {Achatinellastrum) ii 325 
lignaria Gul. {Partulina) ii 314 
liliacea Pfeiff. {Achatinellastrum) ii 328 
limbata Gul. ii 306 
lineolata Newc. ii 338, 365 
livida Pfeiff. ii 327 
livida Swa. {Achatinellastrum) ii 325 
longispira Smith {Achatinellastrum) ii 325 
lorata Fer. ii 303 
lorata Reeve ii 308 
luctuosa Pfeiff. {Laminella) ii 338 
lurida Pfeiff. {Auriculella) ii 376 
luteostoma Baldw. {Bulimella) ii 308 
lymaniana Baldw. {Bulimella) ii 308 
lyonsiana Baldw. {Bulimella) ii 308 
magna C. B. Adams ii 339 
maliogani Gul. li 306 
manoaensis Newc. {Amastra) ii 347 



INDEX 



Achatinella margarita Pfeiff. {Leptachatina) ii 356 
niarginata Gul. ii 366 
marniorata Gould {Pariuliita) ii 314 
mastersi Newc. ii 339 
mauiensis Pfeiff. ii 330 
melampoides Pfeiff. ii 347 
melanosis Newc. ii 339 
melanostoma Newc. ii 306 
niicans Pfeiff. {Laminella) ii 350 
microstoma Gould ii 345 
mighelsiana Pfeiff. (Parlii/ina) ii 314 
modesta C. B. Adams ii 339 
moesta Newc. ii 340 
monacha Pfeiff. ii 303 
morbida Pfeiff. {Bulimella) ii 315 
morbida Pfeiff. \PartuIiiui) ii 315 
mucida Baldw. {Pa>-tiili)ta) ii 315 
mucronata Newc. ii. 340 
multicolor Pfeiff. ii 309 
multilineata Newc. ii 303 

multizonata Baldw. {Achatinellastrum) ii 325 
mustelina Migh. ii 301 
myrrhea Gul. ii 313 
napus Pfeiff. {Achatiiiellastruui) ii 303 
natii Baldw. cS; Hartm. {Achattnellastrum) ii 326 
neglecta Smith ii 301 
nigra Newc. ii 340 
nittida Newc. ii 366 
nivea Baldw. {PartiiHna) ii 315 
nivosa Newc. ii 305 
nobilis Pfeiff. ii 303 
nubilosa Migh. ii 341 
nucleola Gould ii 353 
nucleola Reeve ii 333 
nympha Gul. ii 306 
obclavata Pfeiff. (Leptacliatina) ii 368 
obeliscus Pfeiff. ii 377 
obesa Newc. ii 354 
obesa var. agglutinans Newc. ii 353 
obliqua Gul. ii 306 
obscura Newc. ii 340 
obtusa Newc. ii 366 
octogyrata Gul. ii 366 
olivacea Reeve {AihatineUastrum) ii 326 
ooniorpha Gul. ii 306 
ornata Newc. ii 330 
or>za Pfeiff. {Leptacliatina) ii 367 
ovata Newc. (Bittimetla) ii 308 
oviformis Pfeiff. {BiiUinella) ii 309 
ovum Pfeiff. {Achatinellastrunt) ii 303 
pallida Nutt. ii 303 

papyracea Gul. (Achatinellastrtim) ii 326 
parvula Gul. ii 372 
perdi.\ Reeve {Partulina) ii 315 
perforata Gul. ii 319 
perversa Swains, ii 301 
petila Gul. ii 367 
petricola Newc. ii 341 
pexa Gul. ii 321 
pfeifferi Newc. ii 332 
phaeozona Gul. ii 308 
physa Newc. {Partulina) ii 312, 316 
pica Swains, ii 298 
picta Migh. ii 350 
planospira Pfeiff. ii 307 
platystyla Gul. ii 307 
plicata Pfeiff. ii 332 
plumata Gul. ii 321 
plumbea Gul. {Partulina) ii 316 
polita Newc. {Achatinellastriun) ii 326 
polymorpha Gul. ii 299 



Achatinella porcellana Newc. {Partulina) ii 316 
porphyria Newc. ii 341 
producta Reeve {Achatinellastruni) ii 326 
proxima Pease {Partulina) ii 316 
pulchella Pfeiff. {Achatinellastruin) ii 304 
pulcherrima Reeve ii 328 
pulcherrima Swains, ii 306 
pulla Pfeiff. ii 342 
pumila Gul. ii 339 
pupoidea Newc. ii 342 
pusilla Newc. ii 342 
pygmjea Smith ii 323 
pyramidalis Gul. {Partulina) ii 317 
pyramis Pfeiff. ii 367 
radiata Gould {Partulina) ii 317 
radiata Pfeiff. ii 310 
recta Newc. ii 306 

redfieldi Newc. (Partulina) ii 317, Ixvii 
reevei C. B. Adams li 325 
remyi Newc. ii 351 
resinula Gul. ii 368 
reticulata Newc. ii 343 
rhodoraphe Smith ii 323 
rosea Swains. {Bulimella) ii 309 
rotunda Gul. {Bulimella) ii 309 
rubens Gould ii 343 
rubens Pfeiff. ii 339 
rubiginosa Newc. ii 310 
rudis Pfeiff. ii 344 
rufa Newc. {Partulina) ii 318 
rugosa Newc. {Bulimella) ii 309 
rutila Newc. ii 311 

saccata Pfeiff. {Achatinellastrum) ii 327 
saccula Hartm. {Liptachatina) ii. 368 
sanguinea Newc. ii 351 
saxatilis Gul. ii 368 
scitula Gul. ii 307 
semicarinata Newc. ii 320 
semicostata Pfeiff. {Leptachatina) ii 369 
sericea Pfeiff. {Laminelld) ii 343 
simulans Reeve ii 304 
solida Gul. ii 318 

solitaria Newc. {Achatinellastrum) ii 327 
sordidum Newc. ii 304 
soror Newc. ii 351 
sowerbyana Pfeiff. {Bulimella) ii 310 
spadicea Gul. ii 308 
splendida Newc. {Partulina) ii 31S 
spirizona Fer. ii 344 
stewarti Green ii 328 
straminea Reeve ii 352 
striatella Gul. ii 370 
striatula Gould ii 370 
subrostrata Pfeiff. {Laminella) ii 345 
subula Gul. ii 363 
subvirens Newc. ii 311 
succincta Newc. ii 370 
suffusa Reeve ii 350 
sulcata Pfeiff. {Newcombia) ii 332 
swainsoni PfeitT. {Bulimella) ii 304 
swiftii Newc. ii 300 
taeniolata Pfeiff. {Bulimella) ii 310 
talpina Gul. ii 313 

tappaniana C. B. Adams {Partulina) ii 3 if 
terebra Newc. {Partulina) ii 319 
terebralis Gul. ii 371 
teres Pfeiff. {Leptachatina) ii 371 
tessellata Newc. {Partulina) ii 319, Ixvi 
tetrao Newc. ii 352 
te.xtilis F<fr. ii 345 
theodorei Baldw. {Partulina) ii 330, Ixvii 

A 2 



FAUNA HAWAIIENSIS 



Achatinella torrida Gul. ii 309 

transversalis Pfeiff. {Laminella) ii 345 
tricolor Smith ii 32S 
trilineata Gul. {Achatinellastrutn) ii 327 
tristis Fifr. ii 346 
triticea Gul. ii 367 
tuba Gul. ii 318 
turgida Newc. ii 300 
turrita Gul. ii 372 
turritella F^r. ii 346 
umbilicata Pfeiff. {Laminella) ii 341 
undosa Gul. ii 315 
undulata Newc. ii 323 
ustulata Gul. {Partulina) ii 319 _ 
valida Pfeiff. {Achatinellastrum) ii 300 
variabilis Newc. {Par/ulina) ii 319 
variegata Pfeiff. ii 347 
ventrosa Pfeiff. ii 303 
ventulus Fdr. ii 345, 347 
venulata Newc. ii 326 
venusta Migh. ii 352 
versicolor Gul. ii 301 

versipellis Gul. {AchatineUasfruiii) ii 327 
vespertina Baldw. ii 299 
vestita Migh. ii 304 
vidua Pfeiff. {BiilimcUa) ii 310 
violacea Newc. ii 347 
virens Gul. ii 327 
virgulata Migh. [Parttdhia) ii 320 
viridans Migh. (Bulimelhi) ii 310 
viridans Pfeiff. ii 307 
viridans Reeve ii 325 
vittata Reeve ii 304 
vitrea Newc. ii 372 
vitreola Gul. ii 372 
vulpitia Fdr. {AchntineUastruvi) ii 327 
wailuaensis sp. n. {Achatinellastruni) ii 32 
wheatleyi Newc. ii 307 
zebra Newc. ii 330 
zonata Gul. {Achatinellastrum) ii 329 
Achatinellastrum Pfeiff. ii 320 
conspersa Pfeiff. ii 343 
olesonii Baldw. ii 329 
Achorutidae iii 302 
Acompse suavis Kocli. ii 511 
Acridiodea ii 8, 687, ccxiv 
Acritochaeta gen. n. iii 41, 84 

pulvinata sp. n. iii 42 
Acritus Lee. iii 511 

angustisternum sp. n. iii 528 

angustus sp. n. iii 531 

basalis Lee. iii 522 

basalis var. Sharp iii 522 

concentricus Sharp iii 531 

facilis Sharp iii 517 

facilis var. hawaiiensis var. n. iii 517 

flavitarsis Lew. iii 532 

germanus sp. n. iii 523 

longipes var. haleakalae var. n. iii 530 

insolitus sp. n. iii 532 

insularis Sharp iii 515 

kauaiensis sp. n. iii 51S 

kukuiae sp. n. iii 519 

longipes Sharp iii 529 

makaweliae sp. n. iii 521 

mauiae sp. n. iii 519 

minor sp. n. iii 516 

molokaiae sp. n. iii 518 

monticola Hlackb. iii 527 

nepos sp. n. iii 526, 527 

ornatus sp. n. iii 516 



Acritus parvulus sp. n. iii 533 
perkinsi sp. n. iii 529 
pulchellus sp. n. iii 528 
punctatus sp. n. iii 520 
sculptus sp. n. iii 520 
similis sp. n. iii 527 
solitarius sp. (?) n. iii 532 
subalatus sp. n. iii 530 
subbasalis sp. n. iii 523 
subrotundus sp. n. iii 533 
waianaae sp. n. iii 522 
Acrodrepanis Perk, iii 355 

megalophylla Meyr. iii 355 
nesiotis Perk, iii 355 
Acrolepia Crt. i 726 

aureonigrella sp. n. i 727 
autumnitella Crt. i 726 
Acrosticta Loew. iii 44i,85 

pallipes sp. n. iii 44, 85 
Acrulocercus Cab. i 440, xx.\i 
apicalis Gould i 445 
bishopi Rothsch. i 445 
braccatus Cass, i 445 
nobilis Merr. i 445 
Actidium Matth. iii 535 

sharpianum Matth. iii 535 
Adelencyrtus Ashm. i 323 

kaalae sp. n. i 323 
Adelocera Latr. iii 368 

modesta Boisd. iii 368 
Adenoneura gen. n. i 677 

falsifalcellum sp. n. i 677 
latifemoris sp. n. i 679 
marcidellum sp. n. i 678 
montanum sp. n. i 679 
plicatum sp. n. i 67S 
rufipennis Btl. i 680 
Adoretus Cast, iii 402 

tenuimaculatus Waterh. iii 402 
vitiensis Nonfr. iii 402 
Adrastidia gen. n. ii 503 

longula sp. n. ii 504 
nebulosa sp. n. ii 503 
stigniatica sp. n. ii 503 
stigmatica Sim. iii 343 
Aedia Dp. i 507 
Aegosoma Serv. ii 96 
hirtus iii 645 
reflexum Karsch. ii 96, 
Aeletes Horn iii 511 

concentricus Sharp iii 531 
facilis Sharp iii 517 
flavitarsis Lew. iii 532 
longipes Sharp iii 529 
monticola Blackb. iii 527 
Aeolus cinnamomeus Boisd. iii 369 
Aeschnina ii 62 
Ageleuidae ii 505 
Aglaotoma Forst. i 301 

molokaiensis sp. n. i 301 
rufiventris sp. n. i 301 
Agnostochthona Kirk, iii 694 

alienigera Kirk, iii 694 
Agonismus gen. n. i 512 

argentiferus sp. n. i 513 
coruscans sp. n. i 513 
flavipalpis sp. n. i 512 
Agrilus Steph. iii 400, cxxxi 

sp. iii 400 
Agriolimax Mcirch ii 276 

benvenoti Coll. ii 276 



645 



INDEX 



Agriolimax globosus Coll. ii 276 
laevis Mull, ii 276 
perkinsi Coll. ii 276 
sandwichiensis Soul, ii 276 

Agrionidae ii 693, xxxii 

Agrionina ii 63 

Agrion Fabr. ii 63 

adytum sp. n. ii 69 
adytum var. tillyardi n. ii 695 
amaurodytum sp. n. ii 66 
asteliae sp. n. ii 66 
blackburni M'Lachl. ii 76 
calliphya M'-Lachl. ii 71 
calverti sp. n. ii 694 
deceptor M'^Lachl. ii 74 
eudytum sp. n. ii 68 
hawaiiense M'Lachl. ii 64 
heterogamias sp. n. ii 77 
jugorum sp. n. ii 72 
kauaiense sp. n. ii 75 
koelense Blackb. ii 65 
leptodemas sp. n. ii 70 
microdemas var. n. ii 71 
molokaiense sp. n. ii 73 
nesiotes sp. n. ii 72 
nigro-hamatum Blackb. ii 65, .\.\xii 
oahuense Blackb. ii 74 
oceanicum M'Lachl. ii 76 
oresitrophum sp. n. ii 69 
orobates sp. n. ii 70 
pacificum M^Lachl. ii 64 
satelles Blackb. ii 74 
vagabundum sp. n. ii 75 
williamsoni sp. n. ii 696 
xanthomelas Selys. ii 64 

Agromyzidae iii 74 

Agromyza F"all. iii 74 

Agrotis Ochs. i 142, iii 346 

arenivolanus Butl. i 144 
aulacias sp. n. i 145 
austalea sp. n. i 152 
baliopa sp. n. i 149 
ceramophaea sp. n. i 151 
chersotoides Butl. i 151 
cinctipennis Butl. i 150 
cremata Butl. i 145 
crinigera Butl. i 148, iii 346 
dislocata Walk, i 146, iii 346 
epicremna sp. n. i 149 
hephaestaea sp. n. iii 346 
melanoneura sp. n. i 146 
mesotoxa sp. n. i 148 
microreas sp. n. i 143 
neurogramma sp. n. i 149 
panoplias sp. n. i 144 
perigramma sp. n. i 145 
photophila Butl. i 147, iii 346 
psammophaea sp. n. i 151 
saucia Hubn. i 143 
selenias sp. n. i 150 
spoderopa sp. n. i 148 
tephrias sp. n. i 147 
xiphias sp. n. i 146 
ypsilon Rott. i 143, iii 346 

Agrypnus modestus Boisd. iii 368 

Alaptus Hal. ii 661 

immaturus Peik. ii 661 

Aleocharini iii 551 

Aleyrodidae ii 599 

Alitargus Casey iii 418 

balteatus Cas. iii 418 



Alloeocranum Reut. iii 150 

biannulipes M. & S. iii 150 
AUolobophora Sav. ii 414 

caliginosa Sav. ii 414 

foetida Sav. ii 414 

limicola Mich, ii 414 

nordenskioldii Eisen. ii 414 

putris Hoffm. ii 414 

rosea Sav. ii 414 
Alloniscus Dana ii 524 

floresianus Dollf. ii 524 
Aloha Kirk, ii 579 

ipomoeae Kirk, ii 581 

lehuae sp. n. ii 581 

inyoporicola sp. n. ii 581 

naniicola sp. n. ii 580 

oceanides sp. n. ii 580 

ohiae sp. n. ii 581 

pacifica sp. n. ii 581 
Alphitobius Staph, ii 253 

diaperinus Panz. ii 253 

lateralis Boh. ii 253 

piceus 01. ii 253 
Alucita Z. (Z.) i 477 
Alysiidae i 358, ex 
Amalia M.-T. ii 275 

babori Coll. ii 275 

gagates Urap. ii 276 
Amastra H. & A. Adams ii 333, xix, xlvi 

affinis Newc. ii 333 

alata Pfeiff. {Kai/aia) ii 355 

albolabris Newc. ii 333 

alexandri Newc. {Laininella) ii 348 

amicta Smith ii 333 

anthonii Newc. {Ainastrella) ii 353 

antiqua Baldw. [Aiiuis/rel/a) ii 353 

assimilis Newc. ii 334 

aurostoma Baldw. ii 334 

badia Baldw. ii 334 

biplicata Newc. ii 334 

breviata Baldw. ii 334 

bulbosa Gul. ii 351 

carinata Gul. {Amastrella) ii 353 

citrea Sykes ii 335 

citrina Pfeiff. {Laminelld) ii 348 

concinna Newc. {Lamiitella) ii 348 

conicospira Smith ii 335 

conifera Smith ii 333 

cornea Newc. ii 335 

crassilabrum Newc. ii 335 

cyclostoma Baldw. {Axiastrella) ii 353 

cylindrica Newc. ii 335 

decorticata Gul. ii 336 

depicta Baldw. {Laiiiinelhi) ii 348 

durandi Ancey ii 336 

ellipsoidea Gould ii 336 

elliptica Gul. ii 336 

elongata Newc. (Laininella) ii 349 

erecta Pe.ise {Laininella) ii 349 

extincta Pfeiff. ii 336 

farcimen Pfeiff. [Laininella) ii 349 

flavescens Newc. ii 337 

fratema Sykes (Laininella) ii 349 

frosti Ancey ii 337 

gravida Fer. {Laininella) ii 349 

grayana Pfeiff. ii 337 

heliciformis .Ancey {Kauaia) ii 355 

helvina Baldw. {Laininella) ii 350 

humilis Newc. ii 337 

hutchinsonii Pease {Laininella) ii 350 

inflata Pfeiff. ii 337 



FAUNA HAWAIIENSIS 



Amastra intermedia Newc. ii 338 

irregularis Pfeiff. ii 338 

kauaiensis Newc. {Kauaia) ii 355 

knudseni Baldw. {Kauaia) ii 356 

lineolata Newc. ii 338 

longa Sykes ii 338 

luctuosa Pfeiff. ii 338 

magna C. B. Adams ii 339 

nialleata Smith ii 339 

mastersi Newc. ii 339 

melanosis Newc. ii 339 

micans Pfeiff. {Laminclla) ii 350 

modesta C. B. Adams ii 339 

moesta Newc. ii 340 

mucronata Newc. ii 340 

nana Baldw. ii 340 

nigra Newc. ii 340 

nigrolabris Smith ii 344 

nubilosa Migh. ii 341 

nucleola Gould {A mast reiki) ii 353 

nucula Smith ii 341 

obesa Newc. {AmastreUa) ii 354 

peasei Smith ii 341 

pellucida Baldw. ii 341 

petricola Newc. ii 341 

picta Migh. {Laminella) ii 350 

porphyria Newc. ii 341 

porphyrostoma Pease ii 342 

pullata Baldw. ii 342 

pupoidea Newc. ii 342 

pusilla Newc. ii 342 

rernyi Newc. {Lamindla) ii 351 

reticulata Newc. ii 343 

roseotincta Sykes ii 344 

rubens Gould ii 343 

rubens Reeve ii 347 

rubicunda Baldw. ii 343 

rubida Gul. ii 343 

rudis Pfeiff. ii 344 

rugulosa Pease {Amastrella) ii 354 

rustica Gul. ii 333 

sanguinea Newc. {Laininella) ii 351 

sericea Pfeiff. ii 343 

siniilaris Hartm. ii 343 

similaris Pease ii 354 

simularis Hartm. ii 343 

solida Pease ii 344 

soror Newc. {Lamiiiella) ii 351 

sphaerica Pease {Amasirella) ii 354 

spirizona Fi^r. ii 344 

straminea Reeve {LainineUa) ii 352 

subrostrata Pfeiff. ii 345 

tenuilabris Gul. ii 345 

tenuispira Baldw. ii 345 

tetrao Newc. {Laviinella) ii 352 

te.\tilis Fer. ii 345 

transversalis Pfeifif. ii 345 

tristis Fdr. ii 346 

turritella Fdr. ii 346 

umbilicata Pfeiff. ii 341 

umbrosa Baldw. ii 342 

undata Baldw. ii 346 

uniplicata Hartm. ii 346 

variegata Pfeiff. ii 347 

ventulus Fdr. ii 347 

venusta Migh. {Laininella) ii 352 

vetusta Baldw. {Amastrella) ii 354 

villosa Sykes {Laminclla) ii 352 

violacea Newc. ii 347 
Amastrella subg. n. Amastra ii 352 
Amblyolpium Sim. ii 519 



Amblyolpium longiventer Keyserl. ii 519 

Amblyptilia Hiibn. i 472 

Amblyptilus Wlgrn. i 472 

Amyntas Kinb. ii 414 

hawayanus Rosa ii 420 
hesperidum Bedd. ii 417 
heterochaeta Mich, ii 416 
peregrinus Fletch. ii 414 
schmardae Horst. ii 426 

Anacampsis sarcitea Crt. i 649 
sarcitella Stph. i 649 

Anagyrus How. ii 652 

fraternus sp. n. ii 653 
laeviceps sp. n. ii 654 
major sp. n. ii 652 
nigricans sp. n. ii 653 
tantaleus sp. n. ii 654 
xanthogaster sp. n. ii 653 

Ananca collaris Sharp ii 247 

Anastatus Motsch. i 320 

koebelei sp. n. i 320 

Anas L. i 458 

boscas Linn, i 458 
wyvilliana Scl. i 458 

Anax Leach ii 62 

Junius Drury ii 62 
strenuus Hagen ii 62 

Ancaeus Fauv. iii 539 

laevigatus Kraatz iii 539 

Anchomenides iii 190 

Anchomenini liv 

Anchomenus bardus Blackb. iii 240 
corruscus Er. iii 206 
cuneipennis Blackb. iii 241 
epicurus Blackb. iii 235 
erro Blackb. iii 216 
fossipennis Blackb. iii 239 
fraternus Blackb. iii 236 
fugitivus Blackb. iii 240 
incendiarius Blackb. iii 208 
lucipetens Blackb. iii 210 
meticulosus Blackb. iii 234 
musicola Blackb. iii 223 
oceanicus Blackb. iii 240 
protervus Blackb. iii 240 
putealis Blackb. iii 231 
rupicola Blackb. iii 208 
scrupulosus Blackb. iii 235 
sharpi Blackb. iii 209 

Anchonymus gen. n. iii 199 

agonoides sp. n. iii 199 

Anchotefflus gen. n. iii 195 

elegans sp. n. iii 196 
gracilis sp. n. iii 195 

Ancylostoma duodenale Dub. ii 428 

Ancylus Geoffrey ii 394 

sharpi sp. n. ii 394 

Anemosa aurora Butl. i 225 

Anesychia Hb. i 507 

Anisodactylus cuneatus Karsch iii 193 

Anisolabis Fieb. ii 4 

annulipes Luc. ii 4 
littorea White ii 4 
maritima Bon. ii 4 
paciiica Erichs. ii 4 

Anisomeristes Matth. iii 416 
basalis Sharp iii 416 

Anisops Spin, iii 148 

sp? Kirk, ii 555 

Anobiidae iii 581, cxxviii 

Anobium iii 613 



INDEX 



Anobium paniceum L. iii 613 

Anomalochrysa M^Lachl. ii 47 
angulicosta sp. n. ii 50 
biseriata sp. n. ii 58 
cognata sp. n. ii 50 
debilis sp. n. ii 49 
deceptor sp. n. ii 54 
frater sp. n. ii 52 
fulvescens sp. n. ii 60 
gayi sp. n. ii 56 
haematura sp. n. ii 58 
hepatica M'-Lachl. ii 59 
longipennis sp. n. ii 53 
maclachlani Blackb. ii 54 
molokaiensis sp. n. ii 48 
montana Blackb. ii 49 
nana sp. ti. ii 52 
ornatipennis Blackb. ii 59 
paurosticta sp. n. ii 53 
peles sp. n. ii 49 
princeps sp. n. ii 47 
proteus sp. n. ii 59 
raphidioides sp. n. ii 57 
reticulata sp. n. ii 57, 691 
rhododora sp. n. ii 60 
rufescens M'Lachl. ii 50, 691 
simillima sp. n. ii 55 
soror sp. n. ii 51 
sylvicola sp. n. ii 48 
viridis sp. n. ii 51 
xerophylla van n. ii 61 
zoe sp. n. ii 56 

Anosia Hiibn. i 193 

erippus Cram, i 193 

Anotheorus Blackb. ii 173 

ignavus Blackb. ii 173 
montanus Blackb. ii 173 
robiistus sp. n. ii 173 

Anoura Gerv. iii 302 

Anous Leach "Noio" i 464 

hawaiiensis Rothsch. i 464 
stolidus L. i 464 

Anser Briss. i 457 

albifrons Hart, i 457 

Anteris Forst. i 297, ii 623 

hawaiiensis sp. n. i 298 
montana sp. n. ii 623 
nigricornis sp. n. i 297 
oahuensis sp. n. ii 624 
perkinsi sp. n. i 298 
tarsalis sp. n. i 298 

Anthicidae ii 247, cx.wi 

Anthictis Payk. ii 247 

mundulus Sharp ii 247 
oceanicus Laf. ii 247 

Anthocorinae iii 125 

Anthocoridae ii 551, cxcix 

Anthomyidae iii 29, 84, clxxxvii 

Anthomyia aenescens Wied. iii 30 
spp. iii 43 

Anthothrips Uzel iii 695 

usitatus sp. n. iii 695 

Anthribidae ii 182, cxxiv 

Antilissus Sharp iii 429 

aper Sharp iii 430 

Apamea cinctipennis Butl. i 150 

Apanteles Forst. ii 684 
spp. ii 684 

Apate lifauna Mont, iii 643 

castanoptera Fairm. iii 643 

Apetasimus gen. n. iii 458 



Apetasimus involucer sp. n. iii 458 

Apetinus gen. n. iii 458 

brevis Sharp iii 460 
explanatus Sharp iii 460 
macrothorax sp. n. iii 459 
medius sp. n. iii 459 
pumilio sp. n. iii 460 

Apex albofasciatus Smith ii 304 
coniformis Gul. ii 301 
flavidus Gul. ii 300 
guHcki Smith ii 300 
innotabilis Smith ii 301 
leucophaeus Gul. ii 301 
leucorraphe Gul. ii 302 
leucozonus Gul. ii 303 
liliaceus Gul. ii 300 
neglectus Smith ii 301 
polymorpha Gul. ii 299 
tumefactus Gul. ii 304 
versicolor Gul. ii 301 
tuberans Gul. ii 300 
turbiniformis Gul. ii 300 

Aphaereta Forst. i 358 

muscae Ashm. i 358 

Aphelia Stph. i 687 

Aphidae ii 599, ccx 

Aphidencyrtus Ashm. ii 655 

Aphodius 111. iii 401 

lividus Oliv. iii 401 

Aphonogryllus gen. n. ii 26 
apteryx sp. n. ii 26 

Aphthonetus gen. n. i 517 

albocinera sp. n. 1 527 
aspersa Btl. i 529 
bitincta sp. n. i 521 
columbella sp. n. i 521 
confusa sp. n. i 523 
corticicolor sp. n. i 519 
diffusa sp. n. i 527 
digressa sp. n. i 522 
divergens sp. n. i 520 
elegans sp. n. i 530 
eleuthera sp. n. i 522 
exsul sp. n. i 526 
fluctuosa sp. n. i 519 
fugitiva sp. n. i 518 
hirsuta sp. n. i 528 
humerella sp. n. i 528 
kauaiensis sp. n. i 518 
lichenalis sp. n. i 522 
mediocris sp. n. i 517 
nemo sp. n. i 526 
passerella sp. n. i 520 
plumbifer sp. n. i 524 
polia sp. n. i 525 
puncticiliata sp. n. i 529 
sagittata sp. n. i 530 
spurcata sp. n. i 523 
subocellata sp. n. i 526 
trichophora sp. n. i 524 
veterella sp. n. i 525 

Apis mellifica i 115 

Apodia Hein. i 478 

Apoidea ii 601 

Apomecyna Serv. ii 114 

pertigera Thorns, ii 114 

Aporodes micacea Butl. i 212 

Aporodesminus gen. n. iii 327 
wallacei sp. n. iii 328 

Apororhynchus hemignathi Shipl. ii 434 

Apterocis gen. n. ii 265 



FAUNA HAWAIIENSIS 



Apterocis ephistemoides Sharp ii 
hawaiiensis sp. n. ii 267 



!65 



hystrix sp. n. 11 
impunctatus sp. 
lanaiensis sp. n. 
minor var. n. ii 
montanus sp. n. 
ornaiipennis sp. 
rufo-notatus sp. 
strigosus sp. n. 
subaeneus sp. n. 



270 
n. ii 268 

ii 266 

266 

ii 266 
n. ii 269 
n. ii 268 
ii 269 

ii 270 



266 



405 



182 



vagepunctatus Blackb. 11 

variabilis sp. n. ii 267 

variegatus sp. n. ii 268 

Apterocyclus Waterh. iii 403, cxxxi 
adpropinquans sp. n. iii 404 
deceptor sp. n. iii 405 
feminalis sp. n. iii 405 
honoluluensis Waterh. iii 
munroi sp. n. iii 403 
varians sp. n. iii 404 
waterhousei sp. n. iii 403 

Apterogasterinae iii 704 

Apteromesus gen. n. iii 211 

maculatus sp. n. iii 211 

Aptus Stal. iii 153 

Arachnida ii 443, iii 339, ccxxi 

Araeocerus Schonh. ii 182 

constans sp. n. ii 182 
fasciculatus De Gear ii 

Aramigus fuUeri Horn, iii 653 

Aranea oculata Walck. ii 478 

Araneae ii 444 

Araneus Clertk ii 482 

domesticus Clerck ii 505 
emmae sp. n. ii 482 
emmae Sim. iii 341 
kapiolaniae sp. n. ii 483, iii 34 
nauticus Koch ii 482 

Archips Hiibn. i 690, 736 

basialbana Wl^r. i 690 
capucinus Wlsm. i 736 
consociana Wkr. i 690 
dotatana Wkr. i 690 
foedana Wkr. i 690 
fuscoviridis sp. n. i 693 
leopardelkis sp. n. i 692 
lichenoides sp. n. i 694 
longiplicatus sp. n. i 691, 736 
postvittanus Wkr. i 690, 736 
punctiferanus sp. n. i 692 
retractana Wkr. i 690 
reversana Wkr. i 690 
scitulana Wkr. i 690 
secretana Wlcr. i 690 
secundana Wkr. i 690 
sobriana Wlcr. i 690 
subsenescens sp. n. i 695 
trocliilidanus sp. n. i 693 

Arctocorisa blackburni White ii 

Ardea Linn, i 456 
sacra Gm. 

Argiopidae ii 460 

Argiopinae ii 476 

Argiope Aud. ii 476 

avara Thor. ii 476, iii 
avara kauaiensis subsp. 

Argocerus gen. n. iii 411 

similaris sp. n. iii 41 1 
subguttatus sp. n. iii 412 

Argyresthia (?) aurisquamosa Btl. i 71 



456 



5S4 



341 
n. ii 



477 



Argyresthia zebrina Btl. i 715 
Argyritis Hein. i 478 
Argyrodes Sim. ii 446 

argyrodes Walck. ii 446 

epeirae Sim. ii 446 

hawaiiensis sp. n. ii 447 
Ariadna Aud. ii 445 

perknisi sp. n. ii 445 
Ariamnes Thor. ii 447 

corniger sp. n. ii 447, iii 339 
Aristotelia Hiibn. i 478 

arcuata sp. n. i 482 

elegantior sp. n. i 481 

epermeniella sp. n. i 480 

ichthyochroa sp. n. i 479 

lanaiensis sp. n. i 481 

maculaticornis sp. n. i 478 

mendax sp. n. i 481 

nigriciliella sp. n. i 479 

notata sp. n. i 480 
Arma pacifica Stftl. iii 171 

patruelis StAl. iii 171 
Armadillo Latr. ii 521 

albospinosus sp. n. ii 521 

australis Budde-Lund ii 521 

bidens Budde-Lund ii 521 

danae sp. n. ii 522 

hawaiensis Dana ii 521 

perkinsi sp. n. ii 522 

sharpi sp. n. ii 523 
Artema Walck. ii 446 

sisyphoides Dol. ii 446 
.Ascaris lumbricoides Lutz ii 428 
Asio Briss. "Pueo" i 448 

accipitrinus Pall, i 448 
Asiracidae ii 576, ccv 
Asiracinae iii 124 
Asopia gerontialis Walk, i 
Asopus griseus Burm. i" 
Aspidiotus Bouchc iii 

aurantii Mask. 

camelliae Sign 

citricola Pack. 

cydoniae Comst. iii 107 

cydoniae var. tecta Mask, iii 107 

duplex Cock, iii 108 

greenii Cock, iii 108 

hederae \'all. iii 109 

lataniae Green iii 108 

longispina Mask, iii 107 

maskelli Cock. {Morgaticlla) iii 107 

maskelli Cock, iii 107 

napax Nevvst. iii 108 

nerii Bouche iii 109 

perniciosus Comst. iii 108 

perscarum Cock, iii 107 

pmnaeformis Bouchd iii no 

pomorum Bouche iii 1 1 1 

proteus Mask, iii no 

rapax Comst. iii 108 

transparens Green iii 108 
Aspilota Fcirst. i 359 
Astacopinae iii 162 
Asteiidae iii 73 
Asteia Meig. iii 73 

apicalis sp. n. iii 73 

hawaiiensis sp. n. iii 73 

sp. iii 74 
Astemma Lep. & Serv. iii 158 

peruvianus Gu(fr. iii 159 
Asterolecanium Targ. iii 104 



. 1 


275 


i 


71 


107 




iii 


107 


iii 


108 


iii 


no 



INDEX 



Asterolecanium pustulans Cock, iii 104 

Astichus Foist, i 326 

cyaneus sp. n. i 326 

Astrimus Sharp ii 96 

hirtus Fairm. ii 96 
obscurus Sharp ii 96 

Ataenius Har. iii 401 

pacihcus Sharp iii 401 
peregrinator Har. iii 401 
stercorator Fabr. iii 401 

Atelidiuni gen. n. iii 284 

munroi sp. n. iii 285 

Atelothora.x gen. n. iii 269 

optatus sp. n. iii 269 

Atelothrus gen. n. iii 216 

constrictus sp. n. iii 217 
depressus sp. n. iii 220 
dyscoleus sp. n. iii 220 
erro Blackb. iii 216 
filipes sp. n. iii 218 
gracilis sp. n. iii 219 
hawaiiensis sp. n. iii 219 
insociabihs Blackb. iii 220 
limbatus sp. n. iii 217 
longicollis sp. n. iii 217 
longulus sp. n. iii 218 
platynoides sp. n. iii 221 
politus sp. n. iii 216 
stenopus sp. n. iii 218 
transiens sp. n. iii 221 

Atheta Thorns, iii 576 

coriaria Kraatz iii 577 
olaae sp. n. iii 576 

Athyreodon Ashm. ii 679 

debilis Perk, ii 680 
hawaiiensis sp. n. i 343 
hawaiiensis Ashm. ii 679 

Athysanus Burm. ii 575 

Atrachycnemis Blackb. iii 193 
koebelei sp. n. iii 194 
perkinsi sp. n. iii 194 
sharpi Blackb. iii 193 

Atractomorpha Sauss. ii 687 
crenaticeps Hi. ii 687 

Atrometus Forst. i 350, ii 681 
citricinctus sp. n. i 353 
delicatus sp. n. i 354 
flavifrons sp. n. 1 352 
haleakalae sp. n. i 353 
hawaiiensis sp. n. i 354 
molokaiensis sp. n. i 353 
satelles sp. n. ii 683 
sociabilis sp. n. ii 682 
solitarius sp. n. ii 683 
tarsatus sp. n. i 352 
tephrias sp. n. ii 681 

Attagenus Latr. iii 413 

plebeius Sharp iii 413 

Attidae ii 511 

Attus adansoni tardigradus Aud. ii 511 
africanus Vins. ii 512 
capito Luc. ii 511 
ligo Walck. ii 512 
nigrofuscus Vins. ii 511 
paykulli Aud. ii 512 

Auchenorrhyncha iii 114 

Aulacaspis Cock, iii 109 
rosae iii 109 

Auricula owaihiensis Cham, ii 375 
sinistrorsa Cham, ii 375 

Auriculella Pfeiff. ii 375 



Auricullela ambusta Pease ii 375 

auricula F^r. ii 375 

brunnea Smith ii 375 

cerea Pfeiff. ii 376 

chamissoi PfeilT. ii 376 

crassula Smith ii 376 

diaphana Smith ii 376 

expansa Pease ii 376 

jucunda Smith ii 379 

lurida Pfeiff ii 376 

newcombi Pfeitf. ii 377 

obliqua Ancey ii 377 

patula Smith ii 376 

perkinsi sp. n. ii 377 

perpusilla Smith ii 378 

petitiana Pfeiff. ii 377 

ponderosa Ancey ii 376 

pulchra Pease ii 378 

solida Gul. ii 379 

solidissima Smith ii 379 

tenella Ancey ii 378 

tenuis Smith ii 378 

triplicata Pease ii 378 

umbilicata Ancey ii 383 

uniplicata Pease ii 378 

vvesterlundiana Ancey ii 379 
Automola pelodes Meyr. i 486 
Autosticha Meyr. i 486 

pelodes Meyr. i 487 
Aves i 368, xli, Ixxii 
.Azinis hilarella Btl. i 507 

Bacanius Lee. iii 510 

atomarius Sharp iii 510 

confusus Blackb. iii 511 
Bactra Stph. i 687 

straminea Btl. i 687 
Baeus Hal. ii 622 

persordidus sp. n. ii 622 
Balea newcombi Pfeifif. ii 377 
Banchogastra Ashm. i 343, ii 680 

nigra sp. n. i 343 

vitreipennis sp. n. ii 680 
Banza xxx 
Baracus gen. n. iii 143, ii 553 

hawaiiensis sp. n. iii 143, ii 553 
Baryneus gen. n. iii 209 

sharpi Blackb. iii 209 
Barypristus Sharp iii 208 

incendiarius Blackb. iii 208 

rupicola Blackb. iii 208 

sharpi iii 209 
Bassus Fall, i 340 

laetatorius Fab. i 340 
Batodes Gn. i 703 
Batrachedra Stn. i 508, 734 

bedelliella sp. n. i 509 

ephelus sp. n. i 509 

lomentella sp. n. i 511 

microstigma sp. n. i 510 

ruficiliata sp. n. i 510 

sophroniella sp. n. i 5H 

supercincta sp. n. i 510 

syrraphella sp. n. i 509, 734 
Bavia Sim. ii 511 

aericeps Sim. ii 511 
Bedellia Stn. i 723 

convolvuliella i 723 

minor Busch. i 724 

orchilella sp. n. i 725 

orpheella Stn. i 723 



F. H. 



lO 



FAUNA HAWAIIENSIS 



Bedellia soinnulentella Z. i 723 

staintoniella Clem, i 723 

struthionella sp. n. i 725 
Bembidiides iii 275 
Bembidium Latr. iii 278 

advena sp. n. {Synechostictus) iii 278 

ignicola Blackb. {Emphanes) iii 279 

molokaiense sp. n. {Emphanes) iii 279 

pacificum Blackb. {Emphanes) iii 279 

spurcum Blackb. iii 277 

teres Blackb. {Synechostictiis) iii 278 
Beris servillei Macq. iii 79 
Bernicla Steph. Nene. i 457 

sandvicensis Vig. i 458 
Bethylidae i 282, ii 612, xcix 
Blabophanes Z. i 727 

heringi Rdsn. i 728 

hyalinella Snell. i 728 

lombardica Snell. i 728 

longella Btl. i 727 

monachella Meyr. i 727 

obumbrata Btl. i 728 

rusticella Btl. i 729 
Blackburnia Sharp iii 191 

frigida Blackb. iii 200 

insignis Sharp iii 191 

kaalensis var. n. iii 191 
Blastobasis Z. i 648 

inana Btl. i 648 
Blattidae ii 5, 690, ccxiii 
Blepharipeza monticola Big. iii 20 
Blepyrus How. ii 655 

marsdeni How. ii 655 
Bolitochara impacta Blackb. iii 571 

testacea Kraatz iii 571 
Borboridae iii 75 
Borborus Meig. iii 75 

bilineatus sp. n. iii 75 

venalicius Ost-Sack. iii 75 
Boreophila minuscula Butl. i 233 
Bostrychidae iii 642, cxxvii 
Bostrichus migrator Sharp iii 642 
Bothriothorax Ratz. i 321 

insularis Cam. i 321 
Botys accepta Butl. i 204 

blackburni Butl. i 202 

continuatalis Butl. i 205 

demaratalis Walk, i 205 

localis Butl. i 206 
Brachydeutera Loew iii 49, 85 

argentata Walk, iii 49, 85 

dimidiata Loew iii 49 
Brachymetopa Redt. ii 8, 687, xxx 

affinis sp. n. ii 11 

blackburni Borm. ii 9, 687 

deplanata Brunn. ii 12 

discolor Redt. ii 9, 687 

kauaiensis sp. n. ii 10 

mauiensis sp. n. ii 12 

molokaiensis sp. n. ii 12 

nitida Brunn. ii 9 

parvula sp. n. ii 11 

unica sp. n. ii 10 
Brachypeplus affinis Sharp iii 475 

aper Sharp iii 470 

apertus Sharp iii 498 

bicolor Blackb. iii 465 

bidens Sharp iii 483 

blackburni .Sharp iii 503 

brevis Sharp iii 460 

celatus Sharp iii 498 



Brachypeplus discedens Sharp iii 494 

discedens var. kauaienbis Blackb. iii 495 

expers Blackb. iii 455 

explanatus Sharp iii 460 

floricola Blackb. iii 480 

guttatus Sharp iii 454 

impressus Sharp iii 463 

inaequalis Sharp iii 463 

inauratus Sharp iii 475 

infimus Sharp iii 506 

koelensis Blackb. iii 488 

lanaiensis Blackb. iii 504 

metallescens Sharp iii 493 

obsoletus Sharp iii 467 

olinda Blackb. iii 477 

omalioides Sharp iii 469 

quadraticollis Blackb. iii 501 

paralleUis Blackb. iii 497 

protinoides Sharp iii 486 

puncticeps Sharp iii 494 

reitteri .Sharp iii 505 

robustus Sharp iii 452 

sordidus Sharp iii 455 

spretus Blackb. iii 46S 

striatus Sharp iii 465 

tinctus Sharp iii 493 

torvus Blackb. iii 485 

varius Sharp iii 496 

vestitus Sharp iii 497 
Braconidae i 359, ii 683, 684, ex 
Bracon Fabr. ii 684 
Branta Scop, i 458 

canadensis Ridgw. i 458 

nigricans Lawr. i 458 
Brontolaemus Sharp iii 423 

agilis sp. n. iii 425 

currax sp. n. iii 424 

currax var. lanaiensis var. n. iii 

currax var. mauiensis var. n. iii 

elegans Sharp iii 423 

elegans var. koebelei var. n. iii 424 

nudicornis sp. n. i 
Brosconymus gen. n. iii I 

optatus sp. n. iii 
Bruchidae ii 95, cxv 
Bruchus L. ii 95 

sp. ii 95 
Bubaloceras gen. n. i 

subeburneum 

subeburneum 
Buchananiella Rent, iii 

sodalis White iii 
Buenoa Kirk, ii 554 

pallipes Fabr. ii 
Bulimella PfeifT. ii 305 

attenuata Pfeiff. 

Candida Pfeiff. ii 

forbesiana Pfeiff. 

fuscobasis Smith 11 310 

macrostoma Pfeiff. ii 310 

multicolor Pfeiff. ii 306 

rosea Swains, ii 309 
Bulimus armatus Migh. ii 375 

clausinus Migh. ii 359 

gouldi Pfeiff. ii 317 

insignis Migh. ii 320 

junceus Gould ii 383 

kauaiensis Pfeiff. ii 399 

lactifluus Pfeiff. ii 399 

liratus Pfeiff. ii 332 

nevvcombianus Pfeiff. ii 3 



424 
424 



425 



548 
Wlsm. 
sp. 



128 

128 

555 

ii 319 
308 
ii 300 
3' 



548 
549 



INDEX 



1.1 



Bulimus pumicatus Migh. ii 379 

pyrgiscus Pfeiff. ii 3S4 

rohri Pfeiff. ii 320 

scutilus Migh. ii 368 
Buhveria Bon. i 462 

buhveri J. & S. i 463 
Buprestidae iii 400 
Buprestis adjecta Lee. iii 400 
Buteo Cuv. "lo" i 446 

solitarius Peale i 447 
Bythoscopinae iii 114 
Bythoscopus Germ. Kirk, iii 114 

kaiamamao sp. n. iii 115, ii 557 

kukanaroa sp. n. iii 114, ii 557 

peregrinus StAl. iii 115 

viduus Stal. iii 1 16 

Caccodes Sharp iii 368 

debilis Sharp iii 368, cx.xix 
Cachura Wkr. i 712 

objectella Wkr. i 712 
Cacodminae Kirk, iii 129, ii 552 
Cacoecia postvittana Meyr. i 690 
Caecilianella baldwini Ancey ii 384 
Caecilioides Herrm. ii 384 

baldwini Ancey ii 384 
Cafius Steph. iii 548 

nauticus Fairm. iii 548 
Calandrini ii 139 
Calandra Clairv. ii 139 

hnearis var. striata Thunb. ii 139 

obscura Boisd. ii 139 

oryzae L. ii 139 

remota Sharp ii 139 
Calidris Cuv. i 451 

arenaria L. i 451 
Callidiopsides ii 97 
CalUdium pilicorne Fab. ii 97 
Calliphora Desv. iii 27 

azurea iii 27 

vomitoria Linn, iii 27 

spp. iii 28 
CalHthmysus Sharp ii 113 

cristatus Sharp iii 650 

hirtipes var. n. ii 113 

koebelei Perk, iii 649 

microgaster Sharp ii 113, iii 649 
Caloteleia Westw. ii 624 

elegans sp. n. ii 624 

exul sp. n. ii 625 
Calotermes Hag. ii 88 

marginipennis Latr. ii 88 
Cambalidae iii 329 
CamplNoneuraria Kirk, iii 138 
Campodeidae iii 293 
Campodea Westw. iii 293 
Camponotus maculatus cii 
Camponotus maculatus niitis hawaiensis i 122 
Campsicnemus Walk, iii 13, 80 

calcaratus sp. n. iii 14 

distortipes sp. n. iii 14 

fimbriatus sp. n. iii 13 

patellifer sp. n. iii 80 

sp. iii 15 
Capsidae cc 
Capsaria Reut. iii 142 
Capsus pellucidus Stal. iii 143 
Capua Stph. i 703 

castaneana sp. n. i 705 

flavocincta sp. n. i 704 

ilavopicta sp. n. i 703 



Capua fulva sp. n. i 708 

glaucoviridana sp. n. i 706 

ochreocuprea sp. n. i 708 

picta sp. n. i 703 

pleonectes sp. n. i 705 

trigonifer sp. n. i 704 

variabilis sp. n. i 706 
Carabidae iii 190, liv, cxxxix 
Caraboidea iii 175 
Carabus pallens Fabr. iii 288 
Caradrinidae iii 345, cxlv 
Caradrina Ochs. i 153 

e.xanimis sp. n. i 153 

venosa But!, i 153 
Carcinops Mars, iii 510 

quatuordecimstriata Steph. iii 510 
Cardiastethus mundulus White iii 126 

sodalis White iii 128 
Cardiocondyla nuda minutior i 120 

wroughtonii hawaiensis i 119 
Carelia H. & A. Adams ii 373, xix, Ixxi, Ixxii 

bicolor Jay ii 373 

cochlea Reeve ii 373 

cumingiana Pfeifl". ii 373 

dolei Ancey ii 374 

glutinosa Ancey ii 374 

olivacea Pease ii 374 

paradoxa Pfeiff. ii 374 

sinclairi Ancey ii 374 

turricula Migh. ii 374 

variabilis Pease ii 374 
Carinella Pfeiff. ii 355 
Carpophilus Steph. iii 507 

dimidiatus Fabr. iii 507 

hemipterus Linn, iii 507 

humeralis Fabr. iii 507 

maculatus Murr. iii 507 
Carposinidae i 654, 735, clxvii 
Catamempsis gen. n. i 491 

decipiens sp. n. i 491 
Catapicephala limbipennis Thorns, iii 22 
Cathartus Reiche iii 428 

advena Waltl iii 428 
Catorama mexicana Chev. iii 614 

pusilla Sharp iii 614 
Cerambycidae ii 95, iii 645, cxv 
Cerambycini ii 96 
Cerambyx globosus Oliv. ii 96 
Ceraphronidae ii 616, cii 
Ceraphron Jur. ii 616 

abnormis sp. n. ii 617 

plebeius sp. n. ii 616 
Ceratopogon Meig. iii 5 

sp. iii 5 
Ceratotaxia gen. n. iii 399 

tristis sp. n. iii 400 
Ceresium Newm. ii 97 

simplex Gyll. ii 97 
Ceroplastes Gray iii 104 

ceriferus And. iii 105 

floridensis Comst. iii 105 

rubens Mask, iii 104 
Cerostoma maculipennis Crt. i 652 
Cestoda ii 429 
Chaenosternum Blackb. ii 138 

konanum Blackb. ii 138 
Chaetogaedia Br. & Berg, iii 20, 83 

monticola Big. iii 20, 83 
Chaetoptila Peale i 445 

angustipluma Peale i 445 
Chalcidoidea ii 630 



B 2 



12 



FAUNA HAWAIIENSIS 



Cbalcididae i 305, civ 
Chalcis i 306 

obscurata Walk, i 307 

polynesialis Cam. i 306 
Chalcolepidius Eschsch. iii 368 

erythroloma Cand. iii 368 
Chalcomenus gen. n. iii 206 

corruscus Er. iii 206 

costatus sp. n. iii 207 

molokaiensis sp. n. iii 207 
Chalybe Dp. i 507 
Charadrius L. i 449 

fulvus L. i 450 

squatarola L. i 450 
Charitonetta Stejn. i 461 

albeola L. i 461 
Charopa baldwini Ancey ii 289 
Chasiempis xxxi, clxii 

cabanis "Elepaio" i 379 

gayi Wils. i 381 

sandvicensis (imel. i 381 

sclateri Ridgw. i 381 
Chaulelasmus Bon. i 461 

streperus Linn, i 461 
Cheliferidae ii 517 
Chelifer Geoffr. ii 517 

bifissus Sim. ii 517 

hawaiiensis sp. n. ii 518 
Chelisoches Scudd. ii 4 

morio Fab. ii 4 
Chelodynerus Perk, ii 607 

chelifer Perk, ii 607 
Chelonus Jin", i 359 

blackburni Cam. i 359 

cameronii D. T. i 359 

carinatus Cam. i 359 
Chen Boie i 457 

hyperborea Pall, i 457 
Chermidae ii 598, ccx 
Chermes oleae Bern, iii 106 
Chernetes ii 517 
Chiloides i 687 

straminea Btl. i 687 
Chiloneurus maculatipennis Prov. i 321 
Chilopoda iii 324 
Chionaspis biclavis Comst. iii 112 

biclavis var. detecta Mask, iii 112 

eugeniae Mask, iii 112 

prunicola Mask, iii 112 
CMronomidae iii 47, clxxx 
Chironomus Meig. iii 4 

hawaiiensis sp. n. iii 4 
Chlamydataria Kirk, iii 131 
Chloridops Wils. i 439, Ixxii 

kona Wils. i 440 
Chlorodrepanis i 408, Ixxii 

chloridoides Wils. i 411 

chloris Cab. i 411 

kalaana Wils. i 411 

parva i 409, 4 1 1 

stejnegeri Wils. i 411 

virens Gmel. i 411 

vvilsoni Rothsch. i 411 
Cholovocera Motsch. iii 422 
Chrestotes dryas Btl. i 726 
Chrysoclista haleakalae Btl. i 573 

tigrina Btl. i 533 
Chrysoesthia Hb. i 478 
Chrysomelidae ii 95 
Chrysomyza Fall, iii 85 
Chrysopides ii 47, 691 



Chrysopa Leach ii 61, xvii 

microphya M'Lachl. ii 61 

Chrysopora Clem, i 478 

Chrysotus Meig. iii 1 5 

hawaiiensis sp. n. iii 16 
saxatilis sp. n. iii 16 
spiniger sp. n. iii 15 

Cicadidae iii 114 

Cillaeopeplus gen. n. iii 505 
dubius sp. n. iii 506 
infimus Sharp iii 506 
perkinsi sp. n. iii 506 

Cimicidae iii 171, ii 533, cxci 

Cimex lectularius Linnd iii 129 

rubrofasciatus de Geer ii 550 

Circus Lac. i 447 

hudsonius Linn, i 447 

Ciridops Wils. Ula-ai-hawane i 405, Ixxii 
anna Dole i 405 

Cioidae ii 253, xlix, cxxvii 

Cis Latr. ii 255 

alieniis Sharp ii 255 
angustiformis sp. n. ii 265 
apicalis Sharp ii 260 
attenuatus Sharp ii 261 
bicolor Sharp ii 256 
bimaculatus Sharp ii 257 
breviformis sp. n. ii 255 
calidus Sharp ii 260 
chloroticus Sharp ii 259 
concolor Sharp ii 260 
cognatissimus sp. n. ii 256 
diminutivus Sharp ii 264 
ephistemoides Sharp ii 265 
evanescens Sharp ii 264 
fallax sp. n. ii 263 
haleakale sp. n. ii 262 
insularis Sharp ii 258 
kauaiensis sp. n. ii 261 
laeticulus Sharp ii 264 
longipennis Blackb. ii 265 
mimus sp. n. ii 263 
mirabilis sp. n. ii 262 
molokaiensis sp. n. ii 262 
nesiotes sp. n. ii 256 
nigrofasciatus Blackb. ii 257 
pacificus Sharp ii 255 
porcatus Sharp ii 258 
roridus Sharp ii 261 
setarius Sharp ii 260 
signatus Sharp ii 261 
simulator sp. n. ii 259 
tabidus Sharp ii 259 
unicus sp. n. ii 258 
vagepunctatus Blackb. ii 266 

Cistelidae ii 247, cxxvi 

Cistela Fabr. ii 248 

apicalis sp. n. ii 249 
crassicornis Sharp ii 248 
kauaiensis sp. n. ii 248 
konae sp. n. ii 250 
montana sp. n. ii 249 
nigricollis sp. n. ii 248 
subaenescens sp. n. ii 249 

Cixiaria iii 117 

Clambus Fisch. iii 535 

Clerada Sign, iii 160 

apicicornis Sign, iii 160, ii 545 

Cleridae iii 367 

Clinidium Kirby iii 429 

liratum Chevr. iii 429 



INDEX 



Clinocoridae ii 552 
Clinocoris Fall, ii 552 
Clubionidae ii 504 
Clytides ii 97 
Clytarlus Sharp ii 98, 104 

abnormis sp. n. ii 102 

annectens sp. n. ii 104 

blackburni Sharp ii 1 1 1 

claviger sp. n. ii loi 

cristatus Sharp ii 113 

debilis sp. n. li 99 

filipes Sharp ii 99 

finschi Har. ii 106 

fragilis Sharp ii 99, iii 645 

laticollis sp. n. ii loi 

longipes sp. n. ii 103 

niediocris sp. n. ii 99 

modestus Sharp ii loi 

nodifer sp. n. ii 102 

obscurus sp. n. ii 100 

pennatus Sharp ii 102 

pulvillatus Karsch ii 1 1 1 

robustus Sharp ii 107 

ultimus sp. n. iii 645 
Clytus Laich. ii 97 

crinicornis Chevr. ii 97 
Cnaemidophorus Wlgrn. i 472 
Cnephalocotes Sim. ii 461 

simpliciceps sp. n. ii 462, iii 340 
Coccinellidae iii 414, c.\,\xii 
Coccinella abdoniinalis Say. iii 414 
Coccophagus Westw. i 327 

lecanii Sni. i 328 

ochraceus How. i 328 
Coccidae iii 102, ii 399 
Coccus Linnc iii 105 

acuminatum Sign, iii 105 

adonidum Linne iii 103 

beckii Newm. iii 1 10 

ceriferus And. iii 105 

citri Risso iii 103 

coffeae Walk, iii 105 

gloverii Pack, iii 1 1 1 

hederae Vail, iii 109 

hesperidum Linnd iii 105 

longulum Dougl. iii 106 

niori Sign, iii 106 

nigrum Nietn. iii 106 

oleae Bern, iii 106 

perforatum Newst. iii 106 

tessellatum Sign, iii 106 

zizyphus Luc. iii no 
Cochlogena decora Vix. {Helicteres) ii 301 

lugubris Chemn. {Helicteres) ii 298 
Coenosia Meig. iii 32 

biseta sp. n. iii 39 

dexioides sp. n. iii 33 

dispar sp. n. iii 35 

flavobasalis sp. n. iii 32 

ingens sp. n. iii 40 

kauaiensis sp. n. iii 37 

latimana sp. n. iii 35 

longipes sp. n. iii 38 

rudis sp. n. iii 40 

seminigra sp. n. iii 2,2, 

striata sp. n. iii 38 

triangulifera sp. n. iii 36 

valida sp. n. iii 34 

spp. iii 41 
Coleoptera ii 91, iii 367, xxxii, xlix, cxii 
Coleotichus White iii 172 



n 



•72, n 534- cxcii 



314 



Coleotichus blackburniae White iii 
Collembola iii 299, ccxx 
Colobicus Latr. iii 430 

conformis Pasc. iii 430 
parilis Pasc. iii 430 
Colour groups (Wasps) xciii 
Colovocera Motsch. iii 422 
Colpocaccus gen. n. iii 213 

apicalis sp. n. iii 215 

hawaiiensis sp. n. iii 214 
lanaiensis sp. n. iii 215 

marginatus sp. n. iii 215 

posticatus sp. n. iii 215 

tantalus Blackb. iii 214 
Colpocephalum brachysomum sp. n. iii 

conspicuum sp. n. iii 315 

discrepans sp. n. iii 316 

tpiphanes sp. n. iii 313 

kilauensis sp. n. iii 312 
Colpodes mysticus Blackb. iii 213 

octocellatus Karsch iii 209 
Colpodiscus Sharp iii 209, 213 

lahainensis sp. n. iii 210 

lucipetens Blackb. iii 210 

tantalus Sharp iii 214 
Colydiidae iii 429, cxxxiv 
Colymbetes pacificus Boisd. iii 289 

parvulus Boisd. iii 288 
Comys fusca How. i 321 
Conocephaloides gen. n. ii 13 

hawaiiensis sp. n. ii 13 
Conocephalus blackburni Borm. ii 9 
Conops calcitrans Linn, iii 28 
Conorhinus Lap. ii 550 
Copelatus Erichs. iii 288 

mauiensis Blackb. iii 288 

parvulus Boisd. iii 288 
Coptops Serv. iii 650 

aedificator Fabr. iii 650 
Corixidae iii 148, cciii 
Coriscinae Kirk, iii 169 
Corixa Geoffr. iii 148 

blackburni iii 148, ii 554 
Corizus Sign, iii 170 
Corticaria dentata iVIarsh iii 429 
Corvus Linn, i 372 

hawaiiensis Peale i 373 
Corylophidae iii 415, cxxxii 
Corylophodes Matth. iii 417 

rotundus Sharp iii 417 

suturalis Sharp iii 417 
Corylophus Sharp iii 417 

rotundus Sharp iii 417 

suturalis Sharp iii 417 
Cosilidae ii 616 
Cosmophila Boisd. i 157, iii 348 

noctivolans Butl. i 158, iii 348 

sabulifera Gueii. i 158, iii 348 
Cosmopteryx argyrogrammos Z. i 514 
Cossonini ii 139 
Cothonaspis Hart, ii 668, ii 669 

abnormis sp. n. {Hypodii-aiichis) ii 673 

ashnieadi sp. n. (NcsocUranchis) ii 668 

debilis sp. n. ii 672 

debilis van similis \ar. n. ii 672 

debilis var. subdebilis van n. ii 673 

declivis sp. n. (Hypodiranchis) ii 671 

dichroma sp. n. {Hypodiranchis) ii 674 

dubiosa sp. n. {Hypodiranchis) ii 673 

intermedia sp. n. (Hypodiruiichis) ii 670 

naias sp. n. {Hypodiranchis) ii 672 



14 



FAUNA HAWAIIENSIS 



Cothonaspis pele sp. n. {Hypodiranchis) ii 669 
strigosa sp. n. ii 67 1 
tantali sp. n. {Hypodiranchis) ii 670 
Crabronidae ii 606, Ixxxv 
Crabro i 15 

abnormis sp. n. i 25 
adspectans Bl. & C. i 29 
affinis i 19 
atripennis sp. n. i 24 
curtipes sp. n. i 24 
fulvicrus sp. n. i 22 
hawaiiensis sp. n. i 17 
mandibularis i 21 
mauiensis i 17 
molokaiensis sp. n. i 16 
nionticola sp. n. i 15 
notostictus sp. n. i 20 
polynesialis i 22 
rubrocaudatus Bl. & C. i 27 
stygius Kirby i 29 
tumidoventris sp. n. i 18 
unicolor i 23 
Crambidae iii 356, clvii 
Creophilus Mann, iii 548 

maxillosus L. iii 548 
Crioceris parvula Fabr. ii 95 
Crocidosema Z. i 675, 736 

blackburnii Btl. i 675 
plebeiana Z. i 675, 736 
Crossotarsus Chap, ii 182 

externedentatus Fairm. ii 182 
Crustacea amphipoda ii 527 
Crustacea isopoda ii 521 
Cryptamorpha WoU. iii 428 

desjardinsi Gudr. iii 428 
Cryptinae ii 676 
Cryptodesmidae iii 327 
Cryptophagidae iii 422 
Cryptophagus serratus Gyll. iii 423 
Cryptophilus Reitt. iii 422 

integer Heer iii 422 
Cryptophlebia carpophaga Wlsm. i 680 
illepida Btl. i 681 
illepida fulva var. n. i 
illepida suffusa var. n. 
tetrao sp. n. i 683 
vulpes sp. n. 1 683 
Cryptorhopalum GutJr.-Mifn. iii 
brevicorne .Sharp iii 
terminale Sharp iii 
Cryptorhynchini ii 131 
Cucujidae iii 423, cxxxiii 
Cucujus pusillus Schonh. iii 
minutus Oliv. iii 4: 
Culicidae iii 6, clxxxi 
Culex Linn, iii 6 

taeniatus Wied. iii 6 
Curculionidae ii 117, iii 650, cxix 
Curtomerus Steph. ii 97 

luteus Steph. ii 97 
pilicornis Fab. ii 97 
Cycloneda Crotch iii 414 

abdominalis Say iii 414 
Cyclonotum Erichs. iii 579 

extraneuni sp. n. iii 579 
subquadratum Fairm. iii 578 
Cyclosa Meng. ii 477 

albisternis Sim. ii 478, iii 341 
cucurbitula sp. n. ii 482 
oculata Walck. ii 478 
olorina sp. n. ii 481 



681 
i 682 



111 413 

413 
4>3 



425 

25 



Cyclosa perkinsi sp. n. ii 479 

simplicicauda sp. n. ii 479 
simplicicauda rufescens ii 480 
turbinata Walck. ii 478 
walckenaeri McCo. ii 478 
xanthomelas sp. n. ii 480 

Cyclothorax angusticoUis Blackb. iii 246 
bembidioides Blackb. iii 262 
brevis Blackb. iii 267 
cordaticollis Blackb. iii 259 
deverilli Blackb. iii 270 
inaequalis Blackb. iii 249 
karschi Blackb. iii 266 
laetus Blackb. iii 262 
micans Blackb. iii 244 
montivagus Blackb. iii 253 
multipunctatus Blackb. iii 252 
nubicola Blackb. iii 244 
oahuensis Blackb. iii 274 
obscuricolor Blackb. iii 266 
paradoxus Blackb. iii 263 
pele Blackb. iii 254 
robustus Blackb. iii 268 
rupicola Sharp & Blackb. iii 244 
scaritoides Blackb. iii 272 
simiolus Blackb. iii 274 
unctus Blackb. iii 257 
vulcanus Blackb. iii 249 

Cydninae iii 172, ii 534 

Cylas Latr. ii 131 

turcipennis Boh. ii 131 

Cyminae iii 161 

Cymus calvus White iii 162 

criniger White iii 161 

Cynipoidea ii 667 

Cyrtopeltis Fieb. iii 138 

hawaiiensis sp. n. iii 138, ii 553 

Cyrtostolus gen. n. iii 457 

subalatus sp. n. iii 457 

Dacnitus gen. n. iii 384 

currax sp. n. iii 385 
Dactylopius iii 103 

adonidum Mask, iii 103 

albizziae Mask, iii 103 

calceolariae Mask, iii 103 

vastator Mask, iii 103 

virgatus Cock, iii 103 
Dactylosternum Woll. iii 578 

abdominale Fabr. iii 579 

subquadratum Fairm. iii 578 
Dacus Meig. iii 44 

cucurbitae Coq. iii 45 
Dafila Steph. i 461 

acuta L. i 461 
Dasyuris Guen. i 168 

holombra sp. n. i 168, xxxix 
Decadarchis Meyr. i 714 

melanastra Meyr. i 714 

simulans Btl. i 714 
Degeeria Nic. iii 300 
Deilephila Ochs. i 191 

calida Butl. i 192 

lineata Fabr. (livornica Esp.) i 192 

pyrias sp. n. i 191 

smaragditis sp. n. i 191 

wilsoni Roths, i 192, xxix 
Deinocossonus gen. n. ii 147 

nesiotes sp. n. ii 148 
Deinomimesa gen. n. i 11 

cognata sp. n. i 13 



INDEX 



15 



Deinotnimesa ferox sp. n. i 12 

haleakalae sp. n. i 14 

hawaiiensis sp. n. i 13 

punae sp. n. i 13 
Delphax maidis Ashm. ii 577 

psylloides Leth. ii 577 

pulchra Stal. ii 586 
Deltocephalus hospes Kish. ii 576 
Dendrophilus quatuordecimstriata Steph. iii 510 
Depressaria aigentea Btl. i 496 

convictella Btl. i 487 

gigas Btl. i 493 

gossypiella Sndrs. i 731 

indecora Btl. i 497 

lactea Btl. i 497 

usitata Btl. i 504 
Dermaptera ii 4, 690 
Dermestidae iii 406, cxxxi 
Uermestes Linn, iii 413 

cadaverinus Fabr. iii 413 

fumatus Linn, iii 419 

hemipterus Linn, iii 507 

surinamensis Linn, iii 428 

unidentatus Fabr. iii 428 

vulpinus Fabr. iii 413 
Dermothrips gen. n. iii 677 

hawaiiensis sp. n. iii 678 
Derobroscus gen. n. iii 197 

micans sp. n. iii 197 

politus sp. n. iii 198, cxl 

solitarius sp. n. iii 198 
Derolathrus gen. n. iii 430, cxxxiv 

atomus sp. n. iii 431 
Deroligota subg. n. Oligota iii 555 
Deropristus gen. n. iii 192 

blaptoides Blackb. iii 192 

deroderus sp. n. iii 192 

puncticeps sp. n. iii 192 
Diaea Thor. ii 490 

insulana Keys, ii 490, iii 342 

vitellina sp. n. ii 491 
Diapriidae ii 626, xlix, ciii 
Diapria Latr. ii 629 

drosophilae sp. n. ii 629 

xenica sp. n. ii 629 
Diaspis boisduvalii Sign, iii 109 

costa iii 109 

fioriniae Targ. iii 112 

patelliforniis Sasaki iii no 

rosae Mask, iii 109 
Diceratothrips Bagn. iii 696 

brevicornis sp. n. iii 697 
Dichelia Gn. i 709 

foedana Wkr. i 690 

reversana Wkr. i 690 

sobriana Wkr. i 690 
Dicranomyia Steph. iii 7 

apicalis sp. n. iii 7 

brunnea sp. n. iii 8 

hawaiiensis sp. n. iii 7 

kauaiensis sp. n. iii 8 

latifrons sp. n. iii g 

variabilis sp. n. iii 8 
Dictyophorodelphax Swez. ii 598 
Diestota Muls. & Rey. iii 566 

aberrans sp. n. iii 571 

angustifrons sp. n. iii 567 

athetiformis sp. n. iii 566 

carinata Sharp iii 569 

clavicornis sp. n. iii 575 

crassicornis sp. n. iii 574 



Diestota currax sp. n. iii 566 
frontalis sp. n. iii 570 
incognita Blackb. iii 569 
kauaiensis sp. n. iii 571 
lanaiensis sp. n. iii 573 
latifrons Sharp iii 574 
latiuscula sp. n. iii 572 
lurida sp. n. iii 572 
rnauiensis sp. n. iii 573 
mayeti Muls. & Rey. iii 571 
molokaiensis sp. n. iii 567 
montana Blackb. iii 568 
occidentalis sp. n. iii 569 
palpalis Sharp iii 574 
parva Sharp iii 574 
plana Sharp iii 575 
puncticeps .Sharp iii 571 
robusta sp. n. iii 569 
rufescens Sharp iii 574 
sculpturata sp. n. iii 568 
sordida sp. n. iii 570 
subplagiata sp. n. Hi 570 
testacea Kraatz iii 571 
trogophloeoides sp. n. iii 575 

Dilasia Reut. iii 126 

decolor White iii 126, ii 551 
denigrata White iii 126 

Dilophogaster How. i 324 

californica How. i 324 

Dimera iii 113 

Dimerogonus Attems iii 330 
beddardi sp. n. iii 333 
carpenteri sp. n. iii 332 
harnieri sp. n. iii 336 
koebelei sp. n. iii 338 
lankesteri sp. n. iii 336 
perkinsi sp. n. iii 337 
pococki sp. n. iii 334 
sedgwicki sp. n. iii 334 
sharpi sp. n. iii 330 
sharpi var. iii 331 
shipleyi sp. n. iii 332 
sinclairi sp. n. iii 335 

Dimidiatella Z. i 712 

Dinoderus Steph. iii 643 

minutus Fab. iii 643 

Diomedea L. i 464 

iinmutabilis Roths, i 464 
nigripes Aud. i 464 

Diphysa spinigera Walk, iii 79 

Diplocheta iii 329 

Diploiulus Bed. iii 338 

luscus Mein. iii 338 

Diplopoda iii 327 

Diplosara Meyr. i 646 

lignivora Btl. i 647 

Diptera ii 697, iii I, 79, xlviii, clxxx 

Diptera pupipara iii 86 

Dipterina Meyr. i 697 

fulvosericea sp. n. i 697 
imbriferana Meyr. i 697 

Diranchis Forst. i 302 

monticola sp. n. i 302 
rufipes sp. n. i 302 

Discolomidae iii 431 

Disenochus iii 200, 210 

agilis sp. n. iii 202 
anomalus Blackb. iii 201 
aterriinus sp. n. iii 203 
brevipes sp. n. iii 201 
cephalotes sp. n. iii 201 



i6 



FAUNA HAWAIIENSIS 



Disenochus curtipes sp. n. iii 202 
erythropus sp. n. iii 204 
flavitarsis sp. n. iii 202 
fractus sp. n. iii 205 
longipes sp. n. iii 205 
micantipennis sp. n. iii 206 
sulcipennis sp. n. iii 204 
lerebratus Blackb. iii 211 
Disthymnia Hb. i 507 
Distoma clavatum Rud. ii 429 
hepaticum Lutz ii 429 
Distribution of animals xlvi 
Disuse 1 

Ditoma rugicoUis Walk, iii 430 
Decophorus communis Nitz. iii 306 
macgregori sp. n. iii 306 
fuliginosus hawaiiensis var. n. iii 307 
Dolerotbrips gen. n. iii 682 

angusticeps sp. n. iii 68S 
barbatus sp. n. iii 683 
bicolor sp. n. iii 688 
dubius sp. n. iii 691 
tlavipes sp. n. iii 685 
intermedius sp. n. iii 689 
lanaiensis sp. n. iii 690 
ovatus sp. n. iii 686 
perkinsi sp. n. iii 687 
sp. iii 692 
Dolichopodidae iii 11, 80, cl.\.K.\iii 
Dolichopus Latr. iii 80 
Doryonychus gen. n. ii 465 
raptor sp. n. ii 466 
Doryphora Hein. i 478 

Urepanidotaenia hemignathi Shipl. ii 429, 434 
Drepanididae i 381, .\.\.\i 
Drepanis Temm. i 398, l.xii 

pacifica Gmel. i 401 
Drepanorhaniphus Rothsch. i 401 

funereus Newt, i 402 
Dromaeolus Kiesenw. iii 386 

agriotoides sp. n. iii 388 
arduus sp. n. iii 388 
bonvouloiri Sharp iii 387 
brachycerus sp. n. iii 394 
cephalotes sp. n. iii 397 
collaris sp. n. iii 393 
compressus sp. n. iii 390 
concolor sp. n. iii 396 
cuneus sp. n. iii 389 
elateroides sp. n. iii 394 
germanus sp. n. iii 387 
grandicollis sp. n. iii 398 
hawaiiensis sp. n. iii 397 
kauaiensis var. n. iii 398 
konensis sp. n. iii 390 
mauiensis sp. n. iii 392 
mi.xtus sp. n. iii 391 
molokaiensis sp. n. iii 395 
obscurus sp. n. iii 390 
obtusus Blackb. iii 393 
pachyderes sp. n. iii 398 
parallelus Blackb. iii 392 
perkinsi sp. n. iii 386 
piger sp. n. iii 396 
punticeps sp. n. iii 397 
pumilio sp. n. iii 399 
puncticollis sp. n. iii 391 
sculpluratus ISlackb. iii 392 
solitarius sp. n. iii 392 
sordidus sp. n. iii 389 
sp. iii 394 



Dromaeolus sputator sp. n. ni 395 
subtilis sp. n. iii 395 

Drosophilidae ii 699, iii 50, 86, clxx.\viii 

Drosophila Fall, iii 55, 86 

anomalipes sp. n. iii 62 
carinata sp. n. iii 70 
cognata sp. n. iii 69 
conspicua sp. n. iii 59 
crassifemur sp. n. iii 66 
crucigera iii 86 
exigua sp. n. iii 72 
flaviceps sp. n. iii 63 
haleakalae sp. n. iii 64 
hawaiiensis sp. n. iii 60, 86 
humeralis sp. n. iii 64 
inaequalis sp. n. iii 69 
infuscata sp. n. iii 63 
lanaiensis sp. n. iii 60 
longiseta sp. n. iii 68 
mauiensis sp. n. iii 67 
melanosoma sp. n. iii 68 
molokaiensis sp. n. iii 67 
monticola sp. n. iii 69 
nasalis sp. n. iii 66 
nigra sp. n. iii 62 
obscuricornis sp. n. iii 71 
obscurifrons sp. n. iii 72 
ochracea sp. n. iii 61 
olaae sp. n. iii 66 
parva sp. n. iii 65 
paucipuncta sp. n. iii 62 
perkinsi sp. n. iii 59 
picticornis sp. n. iii 57 
pilimana sp. n. iii 61, 86 
plumosa sp. n. iii 72 
polita sp. n. iii 71 
pusilla sp. n. iii 70 
setiger sp. n. iii 64 
sharpi sp. n. iii 65 
sordidapex sp. n. iii 63 
sp. iii 68 

undulata sp. n. iii 58 
variegata sp. n. iii 57 
varifrons sp. n. iii 71 
xanthosoma sp. n. iii 68 

Dryinidae c 

Dryophthorus Schonh. ii 139, xxviii, Ixiii 
brevipennis sp. n. ii 144 
crassus Sharp ii 141 
declivis Sharp ii 142 
distinguendus sp. n. ii 140 
fuscescens sp. n. ii 145 
gravidus Sharp ii 141 
homoeorhynchus sp. n. ii 142 
insignis Sharp ii 144 
insignoides sp. n. ii 144 
kauaiensis sp. n. ii 143 
modestus Sharp ii 142 
nesiotes sp. n. ii 141 
oahuensis sp. n. ii 143 
peles sp. n. ii 140 
pusillus Sharp ii 143 
squalidus Sharp ii 139 
verticalis sp. n. ii 145 
Dryotribus mimeticus xlvii, cxxii 
Dyscolus Blackb. iii 213 

caliginosus Blackb. iii 233 
mutabilis Blackb. iii 232 
palmae Blackb. iii 232 
tantalus Blackb. iii 214 
Dyscritobaeus gen. n. ii 621 



INDEX 



17 



Dyscritomyia comitans sp. n. ii 622 

gen. n. iii 21, 83 

affinis sp. n. iii 23 

claripennis sp. n. iii 23 

hawaiiensis sp. n. iii 22 

fulgens sp. n. iii 23 

limbipennis Thorns, iii 22 

sp. iii 24 
Dysderidae li 444 
Uysdera Latr. ii 444 

crocata Koch ii 444 

interrita Hentz ii 444 

rubicunda Blackw. ii 444 
Dysdercus Am. .Serv. iii 158 

peruvianus cxciii 
Dysomma gen. n. ii 151 

sylvicola sp. n. ii 152 
Dysphoria gen. n. i 547 

semicolon sp. n. i 548 
Dytiscidae iii 288, cx.\xviii 

Eccoptocera gen. n. i 673, 735 

foetorivorans Btl. i 674, 735 
Echinococcus ii 434 

Echinorhynchus campanulatus Dies, ii 441 
Echthromorpha Holingr. i 336 

flavo-orbitahs Cam. i 336 

maculipennis Holmgr. i 336 
Ecphylopsis Ashm. i 363 

nigra sp. n. i 363 
Eidoreus Sharp iii 415 

minutus .Sharp iii 415 
Elachistidae Meyr. + Plutellidae Meyr. i 508 
Elachista Tr. i 513 

longisquamella sp. n. i 514 

spilota sp. n. i 513 
Elateridae iii 368, cxxix 

flightless iii 384, cxxxi 
Elater humeralis Karsch iii 372 

melanocephalus Thunb. iii 369 
Elenchiis melanias sp. n. iii 667 

melanias var. silvestris var. n. iii 667 
Eleiitheroda Brunn. ii 7 

dytiscoides Serv. ii 7 
Elimaea .Stal. ii 8 

appendiculata Brunn. ii 8 
Ehpsocus Hag. ii 83 

criniger sp. n. ii 85 

debilis sp. n. ii 85 

erythrostictus sp. n. ii 86 

fngidus sp. n. ii 87 

inaequifuscus sp. n. ii 86 

inconstans sp. n. ii 84 

micramaurus sp. n. ii 87 

montanus sp. n. ii 83 

psylloides sp. n. ii 85 

vinosus M'Lachl. ii 86 
Embiidae ii 88 
Emesiidae iii 151, xUx 
Emoia cyanura Stejn. i 367 
Emperoptera gen. n. iii 81, xlviii 

mirabilis sp. n. iii 81 
Emporius Ganglb. iii 429 
Enarmonia Hb. i 683, 736 

conspicua sp. n. i 684 

crassicornis sp. n. 1 685 

obliqua sp. n. i 686 

storeella sp. n. i 686 

walsinghami Btl. i 684, 736 
Encyrtidae i 314, li 630, cv 
Encyrtus Latr. i 321 

F. H. 



Encyrtus fiiscus How. i 321 

insularis Cam. i 321 
Endemic fauna xlvi 
Endodonta alata Pfeiff. (Picrodiscus) ii 292 

apiculata Ancey ii 287 

baldwini Ancey ii 289 

binaria Pfeiff. ii 289 

capillata Pease ii 289 

contorta Yir. ( Thaumatodon) ii 288 

decussatula Pease {Ne.wphild) ii 290 

distans Pease [Nesophila) ii 290 

elisae .4ncey (Nesophila) ii 290 

hystricella Pfeiff. {Thauiiiatodon) ii 288 

hystrix .\Iigh. {Nesophila) ii 290 

jugosa Migh. {Nesophila) ii 290 

lamcllosa F^r. ii 287 

laminata Pease ii 287 

lanaiensis Sykes {Nesophihi) ii 291 

nuda Ancey {Thaumatodon) ii 288 

paucicostata Pease {Nesophila) ii 291 

ringens Sykes {Thaumatodon) ii 288 

rugata Pease ( Thaumatodon) ii 289 

sp. {Nesophila) ii 291 

stellula Gould {Nesophila) ii 291 

tiara Migh. {Nesophila) ii 291 

wesleyi Sykes {Pterodiscus) ii 292 
Endrosis Hb. i 648 

betulinella Hb. i 649 

fenestrella Stn. i 649 

kennicottella Clms. i 649 

lacteella Stgr.-Wkr. i 649 

sarcitea Hw. i 649 

sarcitella Hw. i 649 
Enicospilus Curt, i 345, ii 677 

capnodes sp. n. ii 679 

castaneus 5p. n. i 349 

dimidiatus Perk, ii 679 

dispilus Perk, ii 679 

henshawi sp. n. i 349 

kaalae sp. n. i 347, ii 678 

longicornis sp. n. i 350 

mauicola sp. n. i 347 

molokaiensis sp. n. i 349 
.^ nigrolineatus sp. n. i 348 

semirufus Perk, ii 678 

tyrannus sp. n. ii 678 

variegatus sp. n. i 348 

waimeae sp. n. i 348 
Entedon hagenowii Ratz. i 329 
Entomobryidae iii 300 
Entomobrya Rond. iii 300 

insularis sp. n. iii 301 

kalakaua sp. n. iii 301 
Entozoa ii 427 
Eocerus gen. n. iii 412 

depressus sp. n. iii 412 
Eopenthes .Sharp iii 370 

ambiguus Blackb. iii 380 

antennatus sp. n. iii 378 

arduus sp. n. iii 378 

auratus sp. n. iii 371 

basalis Sharp iii 371 

caeruleus sp. n. iii 370 

celatus sp. n. iii 376 

cognatus sp. n. iii 375 

debilis Sharp iii 380 

deceptor sp. n. iii 374 

divisus sp. n. iii 374 

funebris sp. n. iii 377 

germanus sp. n. iii 373 

gracilis sp. n. iii 376 



FAUNA HAWAIIENSIS 



Eopenthes humeralis Karsch. iii ; 
kauaiensis sp. n. iii 373 
konae Blackb. iii 375 
longicollis sp. n. iii 371 
marginatus sp. n. iii 381 
mauiensis sp. n. iii 376 
muticus sp. n. iii 3S0 



172 



oahuensis sp. 



378 



obscurus Sharp iii 372 
pallipes sp. n. iii 373 
parvulus sp. n. iii 381 
perkinsi sp. n. iii 374 
plebeius sp. n. iii 377 
politus sp. n. iii 373 
satelles ISlackb. iii 379 
tarsalis sp. n. iii 381 
tinctus sp. n. iii 379 
unicolor sp. n. iii 377 
varians sp. n. iii 379 

Epagoge Hb. i 709 

argentinotata sp. n. i 711 
infaustana sp. n. i 709 
pernitida sp. n. i 710 
xanthogona sp. n. i 710 

Epeira caudata Hentz ii 478 
nautica Koch ii 482 
oculata Walck. ii 478 
pullata Thor. ii 482 
strangulata Koch ii 478 
turbinata Walck. ii 478 
walckenaeri Keys, ii 478 

Ephestia Guen. i 196 

albosparsa Butl. i 196 
desuetella Walk, i ig6 
elutella Hub. i 196 
ficulella Barr. i 196 
humeralis Butl. i 196 

Ephestiodes Rag. i 196 

erythrella Rag. i ig6 
gilvescentella Rag. i 196 
infimella Rag. i 196 

Ephydridae iii 49, 85, cl.xxxviii 

Epitragus Latr. ii 252 

diremptus Karsch ii 252 

Epitranus Walk, i 305 

lacteipennis Cam. i 305 

Epitrix Foudr. ii 95 

parvula Fabr. ii 95, cxiv 

Epyris Westw. i 286 

hawaiiensis sp. n. i 286 

Eremotylus Forst. i 345 

orbitalis sp. n. i 345 

Ereunetis Meyr. i 714, 737 

flavistriata sp. n. i 716 
iuloptera Meyr. i 714 
melanastra Meyr. i 715 
minuscula Wlsm. i 716, 737 
simulans Btl. i 715, 737 
zebrina Btl. i 715 

Ergatis Hein. i 478 

Erigone Aud. ii 460 

litoralis Koch ii 461 
vagans Aud. ii 460 

Erinna A. Adams ii 393 

newcombi A. Adams ii 393 

Eriococcus Targ. iii 102 

araucariae Mask, iii 102 

Eris niveipalpis Gerst. ii 511 

Eristalis Latr. iii 19, 82 

punctulatus Macq. iii 82 
tenax Linn, iii 19 



Erotylidae iii 415, cxxxii 
Ethmia Hb. i 507 

colonella sp. n. i 507 

hilarella Btl. i 507 
Euchiradia Hb. i 477 
Euchromius Guen. iii 356 

bella Butl. non Hiib. i 197 

ocelleus Haw. i 197, iii 356 
Eucnemidophorus Wlgrn. i 472 
Eucnemini iii 385 
Eucoilinae ii 667 
Eucoila Westw. ii 674 

hygrophila sp. n. {Psichacrd) ii 675 

orcias sp. n. {Psichacrd) ii 674 

orobates sp. n. {Psichacra) ii 675 
Eucoilidea micromorpha sp. n. ii 676 
Eucymatoge Hiib. i 159, iii 349 

craterias sp. n. i 163, iii 349 

dryinombra sp. n. i 161 

monticolans Butl. i 164, iii 349 

niphoreas sp. n. i 162 

orichloris sp. n. i 163 

phaeocausta sp. n. i 160 

prasinombra sp. n. i 162 

rhodopyra sp. n. i 162 

scoriodes sp. n. i 160 

staurophragma sp. n. i 161 
Eudiestota gen. n. iii 565 

grandis sp. n. iii 565 
Eulachus hispidus Blackb. iii 430 
Eulophidae i 326, ii 657, cvii 
Eulophus Geolifr. i 332 

citripes sp. n. i 332 
Eulota Hart, ii 293 

similaris F^r. ii 293 
Eumenidae ii 607, Ixxxix 
Eumerus Meig. iii 82 

marginatus sp. n. iii 82 
Eunitidula gen. n. iii 451 

sublaevis sp. n. iii 
Eucphrys delibuta Koch li 
Eupelminae ii 630 
Eupelminus D. T. i 315 

subapterus sp. n. i 
Eupelmus Dalm. i 315, ii 

achreiodes sp. n. ii 

amaurodes sp. n. ii 647 

aporostichus sp. n. ii 648 

asthenes sp. n. ii 641 

axestias sp. n. ii 636 

axestops sp. n. ii 638 

basileius sp. n. ii 651 

caerulophantes sp. n. ii 646 

chalcoprepes sp. n. ii 645 

chloropus sp. n. ii 643 

chrysopinus sp. n. ii 640 

dryas sp. n. ii 643 

dysombrias sp. n. ii 637 

dysoplias sp. n. ii 652 

epilamprops sp. n. ii 642 

epimelas sp. n. ii 646 

euoplias sp. n. ii 644 

euprepes sp. n. ii 639 

eustichus sp. n. ii 649 

flavipes Cam. i 319 

hawaiiensis sp. n. i 319 

hemixanthus sp. n. ii 649 

heterosomus sp. n. ii 647 

konae sp. n. i 317 

leptophyas sp. n. ii 642 

leucothrix sp. n. ii 650 



452 
512 



315 
630 
648 



INDEX 



19 



Eupelmus melanacrias sp. n. ii 645 

melanotarsus sp. n. ii 652 

molokaiensis sp. n. i 318 

monas sp. n. ii 641 

niger sp. n. i 316 

ombrias sp. n. ii 636 

oreias sp. n. ii 640 

oribates sp. n. ii 639 

paraleucothri.x sp. n. ii 651 

parasthenes sp. n. ii 641 

para.xestops sp. n. ii 637 

parombrias sp. n. ii 636 

pauroxanthus sp. n. ii 650 

peles sp. n. ii 644 

pelodes sp. n. ii 649 

pelopus sp. n. ii 645 

rhodias sp, n. ii 638 

rhododorus sp. n. ii 647 

rhyncogoni Perk, ii 635 

setigei' sp. n. ii 634 

splendissimus sp. n. i 317 

subsetiger sp. n. ii 635 

vulgaris sp. n. i 318 

xanthodonis sp. n. ii 639 

xanthopus sp. n. i 319 

xanthotarsus sp. n. ii 651 

xestias sp. n. ii 635 

xestops sp. n. ii 637 
Euperissus Btl. i 643, 735 

cristatus Btl. i 643, 735 
Eupetinus gen. n. iii 461 

aper Sharp iii 470 

bicolor Blackb. iii 465 

brevicollis sp. n. iii 469 

brevicornis sp. n. iii 472 

curtus sp. n. iii 466 

derasus sp. n. iii 462 

dubius sp. n. iii 472 

dubius var. iii 472 

hawaiiensis sp. n. iii 464 

impressus Sharp iii 463 

insignis sp. n. iii 461 

laevigatas sp. n. iii 473 

laevigatas var. molokaiensis var. n. iii 474 

lanaiensis sp. n. iii 471 

lanaiensis var. mauiensis var. n. iii 471 

latimargo sp. n. iii 473 

niarginatus sp. n. iii 471 

obscurus sp. n. iii 462 

obsoletus Sharp iii 467 

omalioides Sharp iii 469 

priscus sp. n. iii 465 

sculptus sp. n. iii 467 

sculplus var. parcus var. n. iii 468 

spretus Blackb. iii 468 

striatus Sharp iii 465 

subaper sp. n. iii 463 

sulcatus sp. n. iii 466 

tardus sp. n. iii 470 
Eupithecia monticolens Butl. i 164 
Eurycreon litorea Meyr. i 224 
Eurymelini ii 576 
Eurytomidae ii 630, cv 
Eurytoma 111. ii 630 
Eusipalia gen. n. iii 576 

brachyptera sp. n. iii 576 
Eutettix perkinsi Kirk, ii 559 
Euthyrrhapha Burm. ii 7 

pacifica Coqueb. ii 7 
Eu.xesta Loew iii 44, 85 

annonae Fabr. iii 44, 85 



Euxestus Well, iii 415 

minor Sharp iii 415 
Evagora Clem, i 478 
Evaniidae i 333, cviii 
Evania Fabricius i 333 

appendigaster Linne i 334 

sericea Cam. i 333 
Evaspidiotus hederae Leon, iii 109 
Evergestis Hiib. iii 361 

anastomosalis Guen. iii 361 
Eysarcoris Hahn iii 171 

insularis Dall. iii 172 
Exochus femoralis Grav. i 340 

Falagria currax Sharp iii 578 
Fallia Sharp iii 433 

elongata sp. n. iii 433 
Fauna xxiv 
Figitidae ci\-, ii 667 
Filaria immitis Leidy ii 428 
Fiorinia Sign, iii 112 

camelliae Comst. iii 112 

fioriniae Green iii 112 

pellucida Targ. iii 112 
Flightlessness xlviii 
Flora xxiv, Ivii 
Flowers, inconspicuous xxx 
Forficuloidea ccxii 
Forficula hawaiiensis Brunn. ii 5 
Formicaleo perjurus Walk, ii 61 

wilsoni M^Lachl. ii 62 
Fornax bonvouloiri Sharp iii 387 

obtusus Blackb. iii 393 
Fregata Cuv. "Iwa" i 461 

aquila L. i 461 
Frickella Pfeiff. ii 379 

amoena Pfeiff. ii 379 
Frost xxiii 

Fulgoridae cciv, iii 116 
Fulgormae iii 117 
Fulica L. i 455 

alai Peale i 456 
Fuligula Staph, i 461 

marila L. i 461 

vallisneria A. Wils. i 461 

Gallinago Steph. i 451 

delicata Ord. i 45 1 
Gallinula Briss. i 454 

galeata Bon. i 455 
Garypidae ii 518 
Gary pus Koch ii 51S 

personatus sp. n. ii 51S 
Gehyra mutilata Wieg. i 367 
Gelechiadae clxiii, i 478, 731 
Gelerhia Hb. i 731 

adreptella Wkr. i 654 

gossypiella Sndrs. i 731 

incertella Wkr. i 515 

operculella (> Bryotropha) i 483 
Genophantis Meyr. i 195 

iodora Meyr. i 195, iii 355 
Geoligia DoUf. ii 525 

perkinsi sp. n. ii 525 
Geophilidae iii 326 
Geophilomorpha iii 326 
Geotomus Muls. & Rey. iii 172 

jucundus White iii 172 

pygmaeus Uall. ii 534, iii 172 

subtristis White iii 172 
Gerridae cxcviii, iii 157 



C 2 



20 



FAUNA HAWAIIENSIS 



Glyptogastra Ashm. i 338 

ashmeadi sp. n. ii 677 
hawaiiensis sp. n. i 338 

Clyptoma Erichs. iii 538 

blackburni Sharp iii 538 
brevipenne Sharp iii 539 

Gnamptopsilopus Aldr. iii 11, 80 
palHdicornis sp. n. iii 12 
patellifer Thorns, iii 11, 80 

Gnathocerus Thunb. ii 252 

cornutus Fabr. ii 252 

Gnathohgota subg. n. OHgota iii 556 

Gnatholymnaeum gen. n. iii 276 
blackburni sp. n. iii 276 

Godwinia gen. n. ii 277 

caperata Gould ii 277 
tenella Gould ii 27S 

Gonatocerinae ii 661 

Gonatopus Ljungh i 293 

haleakalae sp. n. i 293 
perkinsi sp. n. i 293 

Goniocotes chinensis sp. n. iii 311 

Gonioryctus Sharp iii 440 

acuminatus sp. n. iii 441 
anticatus sp. n. iii 448 
arduus sp. n. iii 447 
bifarius sp. n. iii 447 
blackburni Sharp iii 449 
calvus sp. n. iii 443 
dissimilis sp. n. iii 446 
elegans sp. n. iii 443 
extraneus sp. n. iii 449 
fugitivus Blackb. iii 438 
haleakalae sp. n. iii 443 
kauaiensis sp. n. iii 440 
koae sp. n. iii 441 
lanaiensis sp. n. iii 446 
latus Sharp iii 445 
latus var. dubius var. n. iii 445 
mauiensis sp. n. iii 440 
molokaiensis sp. n. iii 441 
monticola Sharp iii 451 
oahuensis sp. n. iii 444 
oppositus sp. n. iii 450 
pusillus sp. n. iii 449 
similis Blackburn iii 445 
sp.? iii 443, 444 
suavis sp. n. iii 442 
vicinus sp. n. iii 447 

Goniothorax gen. n. iii 436 

conicicoUis sp. n. iii 436 
cuneatus sp. n. iii 437 
elongatus sp. n. iii 437 
ereniitus sp. n. iii 439 
foveatus sp. n. iii 439 
fugitivus Blackb. iii 438 
inaequalis sp. n. iii 438 
plebeius sp. n. iii 437 
perkinsi sp. n. iii 438 

Gonitis sabulifera Guen. i 158 

Gracilaria Hw. i 721 

convolvuliella Mn. i 723 
epibathra sp. n. i 722 
marginestrigata sp. n. i 721 
praeangusta Hw. i 508 

Gryllodea ii 14, ccxvi, ii 688 

(iryllodes Sauss. ii 15 

poeyi Sauss. ii 15 

Gryllotalpa Lair, ii 14 

africana Fabr. ii 14 

Gryllus ii 14, 688 



Gryllus innotabilis Walk, ii 14 
Gygis Wag. i 464 

alba Sp. i 464 
Gypsonoma Meyr. i 676, 736 

leprarum sp. n. i 676, 736 

Habrobracon Ashm. ii 684 

Haematobia Desv. iii 29 

serrata Desv. iii 29 

Halobates Eschsch. iii 158 

sericeus Eschsch. iii 158 

Haloxenus gen. n. ii 148 

immigrans sp. n. ii 149 

Halticaria Kirk, iii 139 

Hapa White iii 126 

Haptoncus Murr. iii 507 

mundus Sharp iii 508 
tetragonus Murr. iii 508 

Harpalini iii 190 

Hasarius Sim. ii 511 

adansoni Aud. ii 511, iii 344 
garetti Keys, ii 511 
paykuUi Sim. ii 512 

Hastula Mill, i 709 

hyerana Mill, i 709 

Hawaiian group, age of Ix 

Hawaii, island xxi, Ixxii 

Hednota hydrophila Meyr. i 199 
oxyptera Meyr. i 199 

Helcogaster Boh. iii 368 

pectinatus Sharp iii 368, cxxix 

Helegonatopus Perk, ii 655 

Helicinidae ii 396 

Helicina Lam. ii 396 

antoni Pfeiff. ii 397 
bronniana Phil, ii 397 
constricta Pfeiff. ii 396 
crassilabris Phil, ii 397 
fulgora Gould ii 397 
laciniosa Migh. ii 396 
magdalenae Ancey ii 396 
pisum Phil, ii 397 
rotelloidea Migh. ii 397 
sandwichiensis Soul, ii 397 
uberta Gould ii 397 

Helicter hutchinsonii Pease ii 350 
proximus Pease ii 316 

Heliothis Ochs. i 152, iii 346 

armigera HUb. i 152, iii 346 

Heliothrips Hal. iii 698 

haemorrhoidalis Bouch^ iii 698 
rubrocinctus Giard iii 699 

Helicidae ii 292 

Helix alata Pfeiff. ii 355 

barnaclei Smith {Merope) ii 292 
barrakporensis Pfr. ii 285 
binaria Pfeiff. ii 289 
byronii Wood ii 306 
capillata Pease ii 289 
chamissoi Pfeiff. ii 280 
cicercula Gould ii 283 
contorta Ft5r. ii 288 
cryptoportica Gould ii 279 
decora Fer. ii 304 
decussatula Pease ii 290 
disculus Pfeiff. ii 283 
distans Pease ii 290 
exaequata Gould ii 283 
exserta Pfeiff. ii 293 
filocostata Pease ii 291 
fornicata Gould ii 293 



INDEX 



21 



Helix fricki Pfeiff. ii 287 

gravida Fer. ii 349 

hawaiensis Pfeiff. ii 279 

hystricella Pfeiff. ii 28S 

hystrix Pfeiff. ii 290 

intercarinata Migh. ii 288 

jugosa Mijjh. ii 290 

lamellosa Fi^r. ii 287 

laminata Pease ii 287 

lorata F^r. {Cochlo£;t'iia) ii 303 

luteola Yix. {Cochlogena) ii 346 

nevvcombi Pfeiff. ii 277 

obtusangula Pfeiff. ii 283 

paucicostata Pease ii 291 

pauxillus Gould ii 279 

pusillus Gould ii 279 

rubiginosa Gould ii 290 

rugata Pease ii 289 

sandwichensis Pfeiff. ii 293 

setigera Gould ii 290 

similaris Fer. ii 293 

spirizona F^r. {CocJilogena) ii 344 

stellula Gould ii 291 

subrutila Migh. ii 285 

subtilissima Gould ii 285 

textilis Fer. {Helicteres) ii 345 

tiara Migh. ii 291 

tristis Fdr. {Helicteres) ii 346 

turritella Fdr. {Coc/ilogena) ii 346 

ventulus Yir. (Helicteres) ii 347 

vulpina Fdr. ii 327 
Hellula Gucn. i 227, iii 361 

undalis Fabr. i 227, iii 361 
Helops piceus Ol. ii 253 
Hemerobiidae ii 36, 691, clxxi 
Hemidactylus Gray i 366 

garnotii D. & B. i 366 
Hemignathus Licht. i 421 

lanaiensis Rothsch. i 425 

lichtensteini Wils. i 425 

obscurus Gmel. i 425 

procerus Cab. i 426 
Hemiphyllodactylus leucostictus Stejn. i 367 
Hemiptarsenus Westw. i 330 

hawaiiensis sp. n. i 330 
Hemiptera iii 93, ii 531, clxxxix 
Hemitelini i 334 
Hemiteles Grav. i 334 

melitaeae Ashm. i 335 

tenellus Say i 335 

variegatus Ashm. i 334 
Henicopidae iii 325 
Henoticus Thorns, iii 423 

serratus Gyll. iii 423 
Hesperobaenus Lee. iii 434 

capito Fairm. iii 434, cxxxiv 
Hesperophanides ii 96 
Heteractitis .Stejn. "Ulili" i 450 

incana Gmel. i 451 
Heteramphus Sharp ii 152, cxxiii 

cylindricus Sharp ii 154 

filicum sp. n. ii 152 

foveatus Sharp ii 152 

frater sp. n. ii i 53 

haleakalae sp. n. ii 153 

hirtellus Sharp ii 154 

kauaiensis sp. n. ii 154 

molokaiensis sp. n. ii 153 

nivicola sp. n. ii 154 

wollastoni Sharp ii 152 
Heterocrossa Meyr. i 654, 735 



Heterocrossa achroana Meyr. i 671 

atronotata sp. n. i 669 

bicincta sp. n. i 661 

cervinella sp. n. i 667 

corticella sp. n. i 662 

corticella \ar. n. i 664 

crinifera sp. n. i 657 

dispar sp. n. i 661 

distincta sp. n. i 666 

divaricata sp. n. i 665 

ferruginea sp. n. i 664 

gemmata sp. n. i 660 

gracillima sp. n. i 672 

graminicolor sp. n. i 654 

graminis sp. n. i 658 

herbarum sp. n. i 658, 735 

inscripta sp. n. i 669 

irrorata sp. n. i 668 

latifasciata sp. n. i 662 

mauii sp. n. i 668 

nigromaculata sp. n. i 666, 735 

nigronotata sp. n. i 656 

olivaceonitens sp. n. i S^j- 

pipeiatella van n. i 663 

plumbeonitida sp. n. i 054 

puuctulata sp. n. i 671 

pusilla sp. n. i 670, 735 

pygiiiaeella sp. n. i 662 

semitogata var. n. i 662 

solutella sp. n. i 672 

subolivacea sp. n. i 655 

subumbrata sp. n. i 660, 735 

tincta sp. n. i 659 

togata sp. n. i 665 

trigononotata sp. n. i 670 

viridis sp. n. i 656 
Heterophaga lateralis Boh. ii 253 

pandanicola Fairm. ii 253 
Heteropoda Latr. ii 504 

regia Fabr. iii 343, ii 504 

venatoria ii 504 
Heteroptera iii 125, ii 533 
Heterorhynchus Lafr. i 426 

affinis Rothsch. i 430 

hanapepe Wils. i 430 

lucidus Licht. i 430 

wilsoni Rothsch. i 430 
Heterotomaria Kirk, iii 132 
Hevaheva gen. n. iii 113, ii 598 

monticola Kirk, ii 598 

perkinsi sp. n. iii 113 

silvestris Kirk, ii 598 
Hexaplasta Forst. i 304 

konensis sp. n. i 304 
Hiniantopus Briss. i 452 

knudseni Stejn. i 453 
Himatione Cab. i 407 

sanguinea Gmel. i 408 
Hippoboscidae iii 77 
Histeridae iii 598, cxx.xvi 
Hodegia gen. n. i 488 

apatela sp. n. i 4S8 
Holcobius Sharp iii 581 

affinis sp. n. iii 5S3 

diversus sp. n. iii 583 

frater sp. n. iii 585 

glabricollis Sharp iii 584 

granulatus Sharp iii 584 

haleakalae sp. n. iii 582 

haleakalae var. chrysodytus n. iii 583 

hawaiiensis sp. n. iii 583 



22 



FAUNA HAWAIIENSIS 



Holcobius insignis sp. n. iii 582 

major sp. n. iii 582 

minor sp. n. iii 584 

simplex sp. n. iii 584 

simulans sp. n. iii 582 
Holepyris Kieff. ii 615 

hospes sp. n. ii 615 
Holobus Solier iii 555 
Holochila blackburni Tuely i 194 
Holochlora Stal. ii 687 

venosa ii 687 
Holocorynus gen. n. iii 550 

subdepressus sp. n. iii 551 
Homalomyia Bouche iii 30, 84 

canicularis Linn, iii 30 

femorata iii 84 
Homalota coriaria Kraatz iii 577 
Homoeosoma Curt, i 196 

amphibola sp. n. i 197, iii 356 

humeralis Butl. i ig6, iii 355 
Homona Wkr. i 703 
Homoptera iii 102, ii 555 
Hoploderma = Hoplophora Kocli iii 704 

dasypus Dug&s iii 704 
Hoplogaster Rond. iii 43 

dubia sp. n. iii 43 
Hormius Nees ii 685 

peregrinus sp. n. ii 685, c,\i 
Howardia Berl. & Leon, iii 1 1 1 

biclavis Comst. iii 112 

eugeniae Mask, iii 112 

prunicola Mask, iii 112 
Hyalinia baldwini Ancey ii 279 
Hyalopeplus Stftl. iii 143 

pellucidus St41. iii 143, ii 553 
Hydriomenidae iii 349, c.\lix 
Hydriomena Hi.ib. i 164 

aphoristis sp. n. i 165 
Hydrobius Leach iii 57S 

nesiticus sp. n. iii 578 

seniicylindricus Eschsch. iii 578 
Hydroessa Burni. iii 157 
Hydrophilidae iii 578, cxxxviii 
Hydrophilus semicvlindricus Eschsch. iii 578 
Hydrotaea Desv. iii 29 
Hymenia Hub. i 206, iii 358 

recurvahs Fab. (fascialis Cram.) i 206, iii 358 
Hymenoptera ii 600, Ixxiii 
Hymenoptera Aculeata i i 
Hymenoptera Parasitica i 277 
Hypena obsoleta Butl. i 157 
Hypenodes Guen. i 153, iii 347 

altivolans Butl. i 155, iii 347 

arrhecta sp. n. iii 347 

cyanias sp. n. i 154 

epichalca sp. n. i 154 

oxygramma sp. n. i 154 

sarothrura sp. n. i 155, iii 347 
Hypenomyia gen. n. iii 53 

varipennis sp. n. iii 54 
Hyperdasys gen. n. i 640 

arundinicolor sp. n. i 641 

cryptogamiellus sp. n. i 642 

semiustus sp. n. i 640 

unicolor sp. n. i 642 
Hyperectis gen. n. iii 356 

dioctias sp. n. iii 357 
Hyperoniorpha Blackb. ii 139 

squamosa ii 139 
Hypocala Guen. i 159 

andremona Cram, i 159, iii 348 



Hypodiranchis gen. n. i 303 

havvaiiensis sp. n. i 303 
Hyponomeutidae i 508, 734, clxv 
Hyposmocoma Btl. i 549, 734 

abjecta Btl. i 559, 734 

adelphella sp. n. i 615 

adjacens var. n. i 567 

admirationis sp. n. i 580 

adolescens sp. n i 587 

advena sp. n. i 569 

albifrontella sp. n. i 565 

albonivea sp. n. i 599 

alliterata sp. n. i 600, 734 

arenella sp. n. i 627 

argentea sp. n. i 596, 734 

atrovittella sp. n. i 555 

auripennis Btl. i 571 

auroargentea sp. n. i 570 

auropurpurea sp. n. i 614 

bacillella sp. n. i 601 

barbata sp. n. i 635 

bella sp. n. i 614 

belophora sp. n. i 631 

bilineata sp. n. i 573 

blackburnii Btl. i 549, 559, 561, 734 

brevistrigata sp. n. i 633 

butalidella sp. n. i 578 

calva sp. n. i 617 

candidella var. n. i 564 

canella sp. n. i 602 

carbonenotata sp. n. i 599 

carnea sp. n. i 610 

centralis sp. n. i 636 

chilonella sp. n. i 637 

chilonella var. n. i 637 

cincta sp. n. i 566 

cinereosparsa sp. n. i 603 

commensella sp. n. i 558 

conditella sp. n. i 620 

continuella sp. n. i 577 

costimaculata sp. n. i 572 

cupreoniaculata sp. n. i 561 

discella sp. n. i 553 

discolor sp. n. i 633 

divisa sp. n. i 554 

domicolens Btl. i 562, suffusella i 564 

dorsella sp. n. i 605, 734 

dubia var. n. i 590 

emendata sp. n. i 587 

enixa sp. n. i 586 

ensifer sp. n. i 588 

epicharis sp. n. i 639 

evanescens sp. n. i 574 

exornata sp. n. i 550 

fallacella sp. n. i 605 

falsimella sp. n. i 606 

ferricolor sp. n. i 575 

fervida sp. n. i 626 

flavicosta var. n. i 551 

fractinubella sp. n. i 592 

fractistriata sp. n. i 635 

fractivittella sp. n. i 593 

fulvida sp. n. i 608 

fulvocervina sp. n. i 610 

fuscopurpurea sp. n. i 582 

fuscotogata sp. n. i 634 

geminella sp. n. i 594 

genitalis sp. n. i 592 

haleakalae Btl. i 573 

humerovittella sp. n. i 566 

illuminata sp. n. i 570 



INDEX 



23 



Hyposmocoma impunctata sp. n. i 575 
indicella ap. n. i 577 
inflexa sp. n. i 632 
inttrniixta sp. n. i 593 
inversella sp. n. i 615 
iodes sp. n. i 621 
irregularis sp. n. i 626 
labetella sp. n. i 557 
lacertella sp. n. i 582 
lactea sp. n. i 597 
lacticretella sp. n. i 597 
leporella sp. n. i 558 
limata sp. n. i 568 
lineata sp. n. i 579 
liturata sp. n. i 622 
li.xiviella sp. n. i 623 
longitudinalis sp. n. i 636 
lucifer sp. n. i 574 
ludificata sp. n. i 579 
lugens sp. n. i 586 
lunifer sp. n. i 584 
lupella sp. n. i 563 
lupella + suffusella Wlsni. i 734 
maestella sp. n. i 610 
malornata sp. n. i 619 
marginenotata sp. n. i 585 
mediella sp. n. i 565, 734 
metallica sp. n. i 576 
metrosiderella sp. n. i 616 
minienia sp. n. i 613 
mimica sp. n. i 622 
modesta sp. n. i 604 
montivolans Btl. i 620 
nebulifera sp. n. i 555, 734 
nebulifera sp. n. i 628, 735 
niger sp. n. i 582 
nigralbida sp. n. i 551 
nigrescens sp. n. i 581 
nigrodentata sp. n. i 600 
niveiceps sp. n. i 583 
nividorsella sp. n. i 552 
notabilis sp. n. i 556 
numida sp. n. i 581 
obliterata sp. n. i 601 
obscura sp. n. i 609 
ocellata sp. n. i 586 
ochreocervina sp. n. i 612 
ochreociliata sp. n. i 618 
ochreovittella sp. n. i 606 
oculifera sp. n. i 551 
ossea sp. n. i 595 
pallidipalpis sp. n. i 591 
paradoxa sp. n. i 613 
parda Btl. i 623 
partita sp. n. i 554 
patriciella sp. n. i 577 
percondita var. n. i 638 
persimilis sp. n. i 598 
phalacra sp. n. i 625 
phantasmatella sp. n. i 595 
picticornis sp. n. i 594 
progressa sp. n. i 561 
propinqua sp. n. i 580 
pseudolita sp. n. i 625 
pucciniella sp. n. i 589 
punctifiimella sp. n. i 584 
punctiplicata sp. n. i 584 
quadripunctata sp. n. i 607 
quadristriata sp. n. i 581 
quinquemaculata sp. n. i 619 
radiatella sp. n. i 634, 735 



Hyposmocoma rhabdophora sp. n. i 571 
roseofulva sp. n. i 611 
rubescens sp. n. i 628 
rusius sp. n. i 614 
sabulella sp. n. i 565 
saccophora sp. n. i 604 
saliaris sp. n. i 624 
scandeus sp. n. i 609 
scepticella sp. n. i 590 
schismatica sp. n. i 603 
scolopax sp. n. i 629 
semifuscata sp. n. i 589 
sideritis sp. n. i 608 
similis sp. n. i 617 
somatodes sp. n. i 630 



spp..? 1 



624 



stigmatella sp. n. i 591 
straminella sp. n. i 576 
subargentea sp. n. i 596 
subcitrella sp. n. i 632 
subflavidella sp. n. i 569 
sublimata sp. n. i 568 
subnitida sp. n. i 639 
subscolopax sp. n. i 629 
subsericea sp. n. i 611 
sudorella sp. n. i 607 
suffusa var. n. i 563 
syrrhaptes sp. n. i 612 
tarsimaculata sp. n. i 618, 734 
tenuipalpis sp. n. i 557 
tetraonella sp. n. i 630 
thoracella sp. n. i 568 
tomentosa sp. n. i 605 
torella sp. n. i 627 
torquata sp. n. i 556 
tricincta sp. n. i 588 
trilunella sp. n. i 583 
trimaculata sp. n. i 598 
triocellata var. n. i 637 
tripartita sp. n. i 567 
trossulella sp. n. i 554, 734 
turdella sp. n. i 631 
unistriata sp. n. i 578 
venosa var. n. i 638 
vermiculata sp. n. i 585 
vicina sp. n. i 607 
vinitolor sp. n. i 615 
virgata sp. n. i 572 
Hypothenemiis Westw. ii 180 
griseus Blackb. ii 180 
insularis sp. n. ii 181 
maculicollis Sharp ii 180 
ruliceps sp. n. ii iSi 
sylvicola sp. n. ii 181 

Icerya Sign, iii 102 

purchasi Mask, iii 102 

Ichneumonidae cviii, i 334, ii 676 

Ichneumon appendigaster Lin. i 334 
femoralis Fourcr. i 340 

Idechthis Forst. i 355 

oahuensis sp. n. i 355 

Idia pleuralis Thoms. iii 28 

Idiomyia gen. n. iii 50 

heteroneura sp. n. ii 699 
oahuensis sp. n. iii 52 
obscaripes sp. n. iii 52 
perkinsi sp. n. iii 51 
picta sp. n. iii 53 
silvestris sp. n. ii 700 

Immigrant species xliv 



?4 



FAUNA HAWAIIENSIS 



Imported species xliv 
Incilaria bilineata Bens, ii 286 
Inostemnia Hal. ii 626 

abnormis sp. n. ii 626 
Interbreeding xli 
lolania gen. n. iii 1 18 

perkinsi sp. n. iii i ig 
Ischiodontus Cand. iii 369 

hawaiiensis Cand. iii 369 
Ischiogonus Wesm. i 362, ii 684 

palliatiis Cam. i 362 

pallidiceps sp. n. ii 684 
Isometrus macuiatus de Geer ii 517 
Isoptera clxxiv 
Isosoma Walk, ii 630 
Isotonia Bourl. iii 302 

perkinsi sp. n. iii 302 
Ithamar gen. n. iii 169 

hawaiiensis sp. n. iii 170, ii 534, cxciii 
Itodacnus Sharp iii 382 

blackburnianus sp. n. iii 383 

chloroticus sp. n. iii 384 

collaris sp. n. iii 384 

coruscus Karsch iii 382 

gracilis Sharp iii 382 

kauaiensis sp. n. iii 383 

major sp. n. iii 382 

sordidiis sp. n. iii 383 

Japygidae iii 293 
Japyx Hal. iii 293 

sharpi sp. n. iii 293 
Jassidae iii 1 16 

Jotus microphthalmus Koch ii 512 
Julidae iii ^;8 



Kahoolawe xvi 
Kalania Kirk, ii 553 

hawaiiensis ii 553 
Kaliella Blandf. ii 285 

konaensis Sykes ii 286 



Kamehameha gen. n. 



137 



lunalilo sp. n. iii 
Kauai xviii 
Kauaia nom. n. ii 355 
Kelisia Fieb. ii 578 

paludLim sp. n. ii 579 

sporobolicola sp. n. ii 578 

swezeyi sp. n. ii 578 
Kermicus Newst. iii 104 

bambusae Mask, iii 104 
Klinophilos Kirk, iii 129, ii 552 

lectularius Linne iii 129 
Koanoa gen. n. iii 136 

hawaiiensis sp. n. iii 136, ii 554 

Labdia deliciosella Wkr. i 514 
Labeo hawaiiensis sp. n. i 294 
Labetis Waterh. ii 250 

comitans sp. n. ii 251 

hawaiiensis sp. n. ii 250 

sp..' ii 252 

tibialis Waterh. ii 251 
Labia pygidiata Dubr. ii 4 
Labidura icterica Serv. ii 690 
Labiella conipacta Pease ii 359 

pachystonia Pease ii 367 

tenebrosa Pease ii 37 i 

turgidula Pease ii 371 



Labrocerus Sharp iii 406 
affinis sp. n. 
concolor Sharp iii 
curticornis sp. n 
dasytoides sp. n. 
flavicornis sp. n. 



407 
iii 407 
iii 409 
410 



ni 



Labulla 



gravidas sp. n. ni 407 
jaynei Sharp iii 408 
laticornis sp. n. iii 408 
moerens sp. n. iii 406 
obscurus Blackb. iii 408 
obsoletus sp. n. iii 409 
pallipes sp. n. iii 410 
quadrisignatus sp. n. iii 
setosus sp. n. iii 410 
simplex iii 407 
sp. iii 409 

suffusus sp. n. iii 41 1 
vestitus sp. n. iii 409 
Sim. ii 462 
graphica sp. n. ii 46; 



411 



"3 
13 



torosa sp. n. ii 464, iii 341 

Laemophloeus Cast, iii 425 

aeneus Sharp iii 426 
minutus Oliv. iii 425 

Lagocheirus Thorns, ii 113 
araneiformis Sharp ii 
obsoletus Thoms. ii 

Laniiini ii 113 

Lamia aedificator Fabr. iii 650 
bankii Fabr. ii 114 
nutator Fabr. ii 114 

Laminella Pfeiff. ii 348 

albida Pfeiff. ii 344 
depicta Baldw. ii 348 
electa Pease ii 349 
ferussaci Pfeiff. ii 351 
luctuosa Pfeiff. ii 338 
luteola Fdr. ii 346 
mastersi Newc. ii 339 

Lamnonyx spissus Wood iii 326 

Lamprotes Hein. i 478 

Lamyctes fulvicornis Mein. 
hawaiiensis var. n. 
heterotarsus sp. n. 

Lanai xx 

Larus barrovianus Ridg. i 
californicus Lawr. 
delawarensis Ord. 
franklini Sw. & R, 



32s 
325 

325 



464 
1 465 
1 465 
465 



glaucus Br. i 464 
Philadelphia Ord. i 465 
Lasiochilus Reut. iii 126 

decolor White ii 551 
denigrata White iii 126, 
montivagus Kirk, ii 552 
nubigenus Kirk, ii 552 
silvicola Kirk, ii 552 
Lasioderma serricorne F. iii 614 
Lathridiidae iii 421 
Lathridius nodifer Westw. iii 421 
Lathrostizus insularis sp. n. i 355 
Laurentia insularis Butl. i 166 
Laverna abjecta Btl. i 559 
aspersa Btl. i 529 
corvina Btl. i 553 
domicolens Btl. i 562 
parda Btl. i 623, 620 
sarcitella Crt. i 649 
Laysan xvii 
Lebiides iii 288 



n 551 



INDEX 



25 



Lecanium Biirm. iii 105 

acuminatum Sign, iii 105 
chiriniollae Mask, iii 106 
coffeae Walk, iii 105 
depressum Targ. iii 106 
hemisphaericum Targ. iii 105 
hesperidum Mask, iii 105 
hibernaculorum Boisd. iii 105 
longulum Dougl. iii 106 
mori Sign, iii 106 
nigrum Nietn. iii 106 
oleae Sign, iii 106 
perforatum Newst. iii 106 
tessellatum Sign, iii 106 

Leimacis peregrina sp. n. ii 661 

Leiolopisma noctua Stejn. i 367 

Lelapinae i 311 

Lepidocyrtus Bourl. iii 300 

Lepidodactylus crepuscularis Bav. i 367 
heterophthalmus sp. n. iii 300 
lugubris D. & B. i 367 

Lepidoptera i 123, iii 345 
Review of cxliv 

Lepidoptera (micro) i 469 

Lepidosaphes tlava Targ. iii III 
gloverii Pack, iii 1 1 1 
pallida Mask, iii III 
pinnaeformis Bouchd iii iio 
poniorum Boucht? iii lii 

Lepisma hawaiiensis sp. n. iii 296 

Leptachatina Gould ii 356 

accincta Migh. ii 356 
acuminata Gould ii 357 
antiqua Pease ii 357 
antic[uata Pease ii 357 
approximans Ancey ii 357 
arborea sp. n. ii 357 
balteata Pease ii 358 
brevicula Pease ii 35S 
callosa PfeifT. (Lahiclla) ii 358 
cerealis Gould ii 358 
chrysallis Pfeiff. ii 358 
cingula Migh. ii 359 
clausiana Migh. ii 359 
clausina Migh. ii 359 
columna Ancey ii 359 
compacta Pease ii 359 
conicoides sp. n. ii 359 
convexiuscula sp. n. ii 360 
corneola Pfeiff. ii 360 
coruscans Hartm. ii 360 
costulata Gul. ii 360 
costulosa Pease ii 361 
crystallina Gul. ii 361 
cylindrata Pease ii 361 
emerita sp. n. ii 361 
exilis Gul. ii 361 
extensa Pease ii 362 
fumida Gul. ii 362 
fumosa Newc. ii 362 
fusca Newc. ii 362 
fuscula Gul. ii 362 
glutinosa Pfeiff. ii 363 
gracilis Pfeiff. ii 363 
grana Newc. ii 363 
guttula Gould ii 363 
hartmani Hartm. ii 336 
imitatrix sp. n. ii 364 
impressa Sykes ii 364 
isthmica Ancey ii 364 
konaensis sp. n. ii 364 

F. H. 



Leptachatina labiata Newc. {L<Mclla) ii 365 

laevis Pease ii 365 

leucochila (ail. ii 365 

lineolata Newc. ii 365 

lucida Pease ii 365 

marginata Gul. ii 366 

nitida Newc. ii 366 

obsoleta Pfeiff. ii 366 

obtusa Pfeiff. ii 366 

octavula Paet. ii 368 

octogyrata Gul. ii 366 

oryza Pfeiff. ii 367 

pachystoma Pease {Labiclla) ii 367 

petila Gul. ii 367 

perkinsi Sykes ii 367 

pyramis Pfeiff. ii 367 

resinula Gul. ii 368 

saccula Hartm. ii 368 

sandwicensis Pfeiff. ii 368 • 

saxatilis Gul. ii 368 

saxitilus Gul. ii 368 

sculpta Pfeiff. ii 368 

scutilus Migh. ii 368 

semicostata Pfeiff. ii 369 

semipicta Sykes ii 369 

simplex Pease ii 369 

smithi Sykes ii 369 

striata Newc. ii 370 

striatella Gul. ii 370 

striatula Gould ii 370 

stiria Gul. ii 369 

succincta Newc. ii 370 

supracostata ii 370 

tenebrosa Pease ii 371 

tenuicostata Pease ii 371 

terebralis Gul. ii 371 

teres Pfeiff. ii 371 

turgidula Pease ii 371 

turrita Gul. ii 372 

vana sp. n. ii 372 

vitrea Newc. ii 372 

vitreola Gul. ii 372 
Leptacinus flavipennis Kraatz iii 551 
Lepthemis blackburni M'Lachl. ii 62 
Leptia Gn. i 687 
Leptogenys falcigera i 118 

insularis Sm. i 118 
Leptogryllus gen. n. ii 28, 688 

apicalis sp. n. ii 689 

cylindricus sp. n. ii 688 

deceptor sp. n. ii 689 

elongatus sp. n. ii 29 

forficularis Brunn. ii 29 

fusconotatus sp. n. ii 29 

kauaiensis sp. n. ii 29 

nigrolineatus sp. n. ii 28 

nigromaculatus sp. n. ii 28 

siinilis sp. n. ii 28 

simillimus sp. n. ii 30 
Leucania Ochs. i 138, iii 345 

amblycasis sp. n. i 141, iii 346 

anthracias sp. n. i 139 

compsias sp. n. i 139 

dislocata Walk, i 146 

euclidias sp. n. i 140, iii 345, c.xlv 

macrosaris sp. n. i 140 

niphadopa sp. n. i 140 

photophila Butl. i 147 

pyrrhias sp. n. i 141 

typhlodes sp. n. i 141 

unipuncta flaw, i 142 



D 



26 



FAUNA HAWAIIENSIS 



Leucophaea surinamensis Fab. ii 7 
Leucostonia analis Meig. iii 20 
Leurocorynus gen. n. iii 548 

cephalotes sp. n. iii 549 
Liancalus nietalliciis sp. n. iii 13 
Libellulina ii 62 
Liburnia psylloides Dist. ii 577 
Lilia dilecta White iii 128, cc 
Limacidae ii 275 
Limax gagates Drap. ii 276 

laevis Miill. ii 276 

sandwichiensis Soul, ii 276 
Liranaea Lam. ii 390 

aSfinis Soul, ii 392 

ambiyiia Pease ii 391 

aulacospira Ancey ii 391 

binominis nom. n. ii 391 

compacta Pease ii 391 

hartmanni Cless. ii 392 

moreletiana Cless. ii 392 

natlcoides Cless. ii 392 

oahuensis Soul, ii 392 

peasei Cless. ii 393 

reticulata Gould ii 393 

rubella Lea ii 393 

sandwichensis Phil, ii 391 

turgidula Pease ii 393 

volutata Gould ii 392 
Lininaecia phragmitella Stn. i 485 
Limnaeus sandwichensis Phil, ii 392 
Limneria Holmgr. i 356 

polynesialis Cam. i 356 
Limnerium Ushm. i 356 

blackburni Cam. i 357 

hawaiiense Cam. i 357 

polynesiale Cam. i 356 
Liumobiidae iii 6, clx.xxii 
Limnobia perkinsi sp. n. iii 6 
Limosa baueri i 451 
Limosina aequalis sp. n. iii 76 

venalicia Ost.-Sack. iii 75 
Limothrips Hal. iii 701 

avenae Hinds iii 701 

cerealium Hal. iii 701 
Lineodes Gn. i 469, 731 

ochrea sp. n. i 469, 731 

subextincta sp. n. i 470 
Linyphia argyrodes Walck. ii 446 
Liophaena Sharp iii 564 

centralis sp. n. iii 564 

fiaviceps Sharp iii 564 

gracilipes Sharp iii 564 

oahuensis sp. n. iii 565 
Lipeurus docophoroides minhaensis var. n. iii 
Lipogomphus Berg iii 168 
Lispe argenteifacies sp. n. iii 30 

cupreigena sp. n. iii 31 

spp. iii 31 
Lispinodes Sharp iii 539 

crassus sp. n. iii 542 

explicandus Sharp iii 540 

germanus sp. n. iii 542 

hawaiiensis sp. n. iii 541 

kauaiensis sp. n. iii 541 

mauiensis sp. n. iii 541 

molokaiensis sp. n. iii 540 

obscurus sp. n. iii 541 

oxytelinus sp. n. iii 540 

pallescens Blackb. iii 539 

pallidus sp. n. iii 540 

quadralus Blackb. iii 539 



Lispinus impressicollis Motsch. iii 539 

laevigatus Kraatz iii 539 
Lissonota bellator Grav. i 336 
Litargellus Cas. iii 419 
Litargus balteatus Lee. iii 418 

vestitus Sharp iii 419 
Lithobiidae iii 324 

Lithobius hawaiiensis sp. n. iii 324 
Lithocharis fuscipennis Kraatz iii 544 

incompta Sharp iii 545 

vilis Kraatz iii 544 
Lithurgus sp. .'' ii 605 
Lobioptera lacteipennis Loew iii 74 
Loboptera extranea sp. n. ii 6 
Locastra monticolens Butl. i 211 
Locustoidea ii 8, 687, ccxi\ 
Loxioides Oust, i 435 

bailleui Oust, i 436 
Loxops Cab. i 417 

caeruleirostris Wils. i 421 

coccinea Gmel. i 420 

ochracea Rothsch. i 421 

rufa Blox. i 421 
Loxostege conisalias sp. n. i 226 

helio.xantha sp. n. i 226 

melanopis sp. n. i 226 
Lozostoma flavofasciata Stn. i 712 
Lucanidae iii 403 
Lucilia Uesv. iii 28, 84 

caesar Linn, iii 28 
Luteva insolida White ii 550, iii 152 

insulicola Kirk, ii 551 
Lycaena baetica Linn, i 194, iii 355 

blackburni Tuely i 194, iii 355 
Lycaenidae clvi, iii 355 
Lyctidae cxxviii, iii 644 
Lyctus Fabr. iii 644 

brunneus Steph. iii 644 

sp. ? iii 644 
Lycosidae ii 505 
Lycosa Latr. ii 505 

hawaiiensis Sim. ii 506 

kalukanai sp. n. {Trochosa) ii 506 

likelikeae sp. n. ii 506 

oahuensis Keys, ii 505 

perkinsi sp. n. iii 343 

stygialis Sim. ii 505 
Lycosella annulata sp. n. ii 508, iii 344 

spinipes sp. n. ii 507 
Lygaeidae ii 534, iii 169, cxciii 
Lygaeus hyalinus Fabr. iii 170 

peruvianus Guer. iii 159 
Lygosoma cyanurum Less, i 367 

noctua Less, i 367 
Lymnaeus rubellus Lea ii 393 
Lyonetia orpheella Stn. i 723 

somnulentella 7^. i 723 

Machilidae iii 294 

Machilis heteropus sp. n. iii 295 

perkinsi sp. n. iii 294 
Macranillus gen. n. iii 287 

coecus sp. n. iii 287 
Macrochlamys perkinsi Sykes ii 284 
Macrodyctium omiodixorum Terry ii 684 
Macropis Auctt. iii 114 
Macrosteles kilaueae sp. n. ii 575 
Malacodermidae iii 368 
Mallophaga iii 30; 

Hosts of, iii 320 
Mammalia i 465 



INDEX 



27 



Mantodea ii 7, 689, ccxiii 
jSIapsidius gen. n. i 650 

auspicata sp. n. i 650 

iridescens sp. n. i 651 

quadridenta sp. n. i 651 
Mareca americana Gni. i 460 
Margarodes e.xaula Meyr. i i.X) 
Margaronia Hiib. i 200, iii 357 

cyanomichla sp. n. i 201, iii 337 

exaula Meyr. i 2CX3, iii 357 

glauculalis Ikitl. i 200 
Maui XX 

Mauia satelles lilackb. ii 183 
Mauna Blackb. iii 200 

frigida Blackb. iii 200 
Mecaphesa gen. n. ii 495 

cincta sp. n. ii 495 

perkinsi sp. n. iii 342 

seniispinosa sp. n. ii 496, iii 343 
Mecistocephaliis Wood iii 326 
Mecomenus gen. n. iii 230 

koelielei sp. n. iii 230 

putealis Blackb. iii 231 
Mecostomus gen. n. iii 229 

perkinsi sp. n. iii 229 
Mecyclothorax gen. n. iii 243 

aeneus sp. n. iii 255 

amaroides sp. n. iii 256 

anguslicollis Blackb. iii 246 

bicolor sp. n. iii 246 

bradycellinus sp. n. iii 247 

cognatus sp. n. iii 255 

cymindicus sp. n. iii 248 

daptinus sp. n. iii 249 

inaeqnalis Blackb. iii 249 

interruptus sp. n. iii 252 

iteratus sp. n. iii 250 

konanus sp. n. iii 248 

laetus sp. n. iii 247 

longulus , 1. n. iii 25 1 

micans Bla^^kb. iii 244 

niicrops sp. n. iii 245 

montivagus Blackb. iii 253 

multipunctatus Blackb. iii 252 

nubicola Blackb. iii 244 

obscuricornis sp. n. iii 245 

occultus sp. n. iii 256 

oculatus sp. n. iii 251 

ovipennis sp. n. iii 250 

pele Blackb. iii 254 

pusillus sp. n. iii 243 

robustus sp. n. iii 255 

rusticus sp. n. iii 244 

sobrinus sp. n. iii 253 

vLilcanus Blackb. iii 249 
Mecyna -Steph. i 225, iii 360 

aurora Butl. i 225, iii 360 

ennycliioides Butl i 216 

exigua Butl. i 245 

nigrescens Butl. i 211 

virescens Butl. i 225 
Medon celebensis Fauv. iii 545 

debilicornis WoU. iii 545 

debilicornis Ganglb. {Hypomcaon) iii 545 
Megachilldae Ixxxiii, ii 605 
Megachile diligens i 113 

palmarum sp. n. i 114 

schauinslandi i 114 
Megalomus Ramb. ii 36 
Megalomus hospes sp. n. ii 36 
Megascolex schmardae Horst ii 426 



Melaniidae ii 394 
Melania Lam. ii 394 

baldwini Ancey ii ^94 

contigua Pease ii 395 

indefinita Lea ii 394 

kauaiensis Pease ii 395 

mauiensis Lea ii 395 

newcombii Lea ii 394, 395 

verrauiana Lea ii 395 

verreauxiana Lea ii 395 
Melanoleuca Stph. i 507 
Melanomecyna ennychioides i 216 

nigrescens i 211 

stellata Butl. iii 359 
Melanoxanthus Eschsch. iii 369 

melanocephalus Thunb. iii 369 
Meliphora grisella Fab. i 197 
Melittobia hawaiiensis Perk, ii 658 
Membracidae ccix, ii 555 
Menemerus paykulli Thor. ii 512 
Menopon cyrtostigum sp. n. iii 318 

hawaiiensis sp. n. iii 317 

hilensis sp. n. iii 319 

invadens sp. n. iii 320 
Mergus serrator L. i 461 
Merininetria gen. n. i 482 

flaviterniinella sp. n. i 4S2 
Merragata hebroides White ii 546, iii 168 
Mesolelaps gen. n. i 313 

cyaneiveiitris sp. n. i 313 . 
Mesothriscus gen. n. iii 222 

abax sp. n. iii 226 

alternans sp. n. iii 228 

coUaris sp. n. iii 225 

hawaiiensis sp. n. iii 224 

kauaiensis sp. n. iii 226 

lanaiensis sp. n. iii 223 

microps sp. n. iii 226 

muscicola Blackb. iii 223 

opacus sp. n. iii 227 

optimus sp. n. iii 227 

prognathus sp. n. iii 223 

tricolor sp. n. iii 224 

truncatus sp. n. iii 225 

vagans sp. n. iii 222 
.Mestolobes Butl. i 228, iii 361 

abnormis Butl. i 239, iii 361 

amethystias sp. n. i 229 

antichora sp. n. iii 361 

aphrias sp. n. i 237 

arctura sp. n. i 231 

autodoxa sp. n. i 239, iii 361 

banausa sp. n. i 233 

chimonias sp. n. i 236 

chlorolychna sp. n. i 237 

chrysomolybda sp. n. i 230 

crypsichrysa sp. n. i 230 

droseropa sp. n. i 232 

epidelta sp. n. i 231 

erinnys sp. n. i 231 

eurylyca sp. n. i 235 
• homalopa sp. n. i 240 

iochrysa sp. n. i 232 

mesacma sp. n. 1 237 

minuscula Butl. i 233, iii 361 

ochrias sp. n. i 234 

oenone i 239 

ombrias sp. n. i 233 

orthrias sp. n. i 235 

perixantha sp. n. i 23S 

pessias sp. n. i 236 



D2 



28 



FAUNA HAWAIIENSIS 



Mestolobes pragniatica sp. n. i 239 
scleropis sp. n. 1 234 
semiochrea Bull, i 238, iii 361 
simaethina i 233 
sirina sp. n. i 231 
xanthoscia sp. 11. i 230, iii 361 
Metacoelus femoralis Fourcr. i 340 
Metasia abnonnis Bull, i 239 
Methana ligata Brunn. ii 6 
Metrarga iii 165, ii 535 

contracta Blackb. iii 166 
nuda iii 165, ii 535 
obscura iii 166, ii 535 
villosa White iii 167 
Metrarginae subf. nov. iii 164 
Metrocidium gen. n. iii 285 

admirandum sp. 11. iii 286 
brevicolle sp, n. iii 285 
Metromenus Sharp iii 231 

aequahs sp. n. iii 233 
angustifrons sp. n. iii 234 
bardus Blackb. iii 240 
calathoides sp. n. iii 242 
cahginosus Blackb. iii 233 
cinctus sp. n. iii 239 
cuneipennis Blackb. iii 241 
epicurus Blackb. iii 235 
fossipennis Blackb. iii 239 
fugitivus Blackb. iii 240 
fraternus Blackb. iii 236 
fraudator sp. n. iii 237 
meticulosus Blackb. iii 234 
moerens sp. n. iii 237 
mutabilis Blackb. iii 232 
mysticus Sharp iii 213 
latifrons sp. n. iii 242 
lentus sp. n. iii 235 
limbatus sp. n. iii 241 
oceanicus Blackb. iii 240 
palmae Blackb. iii 232 
pavidus sp. n. iii 238 
perpolitus sp. n. iii 241 
protervus Blackb. iii 240 
scrupulosus Blackb. iii 235 
sphodriformis sp. n. iii 238 
velo.\ sp. -n. iii 236 
Metrothorax gen. n. iii 269 

blackburni sp. n. iii 272 
crassus sp. n. iii 274 
curtipes iii 273 
debilis sp. n. iii 275 
de\erilli Blackb. iii 270 
discedens sp. n. iii 275 
extimus sp. n. iii 273 
haleakalae sp. n. iii 271 
laticollis sp. n. iii 271 
macrops sp. n. iii 270 
molops sp. n. iii 269 
oahuensis Blackb. iii 274 
perkinsianus sp. n. iii 270 
rotundicollis sp. n. iii 273 
scaritoides Blackb. iii 272 
simiolus Blackb. iii 274 
Micracantha insularis Pascoe ii 114 

nutans Sharp ii 114 
Microcleptes Stal. iii 150 
Microcystina Morch. ii 279 

cryptoportica Gould ii 279 
Microcystis Beck ii 279 

abeillei Ancey ii 280 
baldwini Ancey ii 281 



Microcystis boettgeriana Ancey ii 283 
chamissoi Pfeiff. ii 280 
depressiuscula Ancey ii 284 
indefinita Ancey ii 284 
lymanniana Ancey ii 284 
oahuensis Ancey ii 284 
perlucens Ancey ii 284 
platyla Ancey ii 2S4 
plicosa Ancey ii 285 
sericans Ancey ii 285 
turgida Ancey ii 285 
Microdus hawaiicola sp. n. i 361 
Microgasterinae ii 684 
Microneta insulana sp. n. ii 461 
Microterys Thorn, i 322, ii 655 
flavus How. ii 655 
molokaiensis sp. n. i 322 
Microvelia Westw. iii 157 

vagans White iii 158 
Midway island xvii 
Milu Kirk, ii 549 

kerasphoron Kirk, ii 549 
rubritinctus Kirk, ii 549 
Miniesa antennata Sni. i 10 
Minthea rugicollis Walk, iii 430 
Miraria Reut. iii 144 
Miridae iii 125, ii 552, cc 
Mirinae ii 552, 652, iii 129 
Mirosternus Sharp iii 614 

acutus Blackb. iii 639 
affinis sp. n. iii 621 
affinis var. suturalis n. iii 622 
amatus sp. n. iii 632 
amaurodes sp. n. iii 636 
angulatus sp. n. iii 631 
bicolor Sharp iii 629 
blackburni sp. n. iii 621 
blackburnioides sp. n. iii 621 
carinatus Sharp iii 638 
cognatus sp. n. iii 632 
debilis Sharp iii 640 
denudatus sp. n. iii 624 
dimidiatus sp. n. iii 641 
discolor sp. n. iii 634 
dubiosus sp. n. iii 637 
duplex sp. n. iii 620 
elongatulus sp. n. iii 636 
epichrysus sp. n. iii 619 
euceras sp. n. iii 620 
eutheorus sp. n. iii 628 
excelsior sp. n. iii 617 
eximius sp. n. iii 635 
fractus sp. n. iii 625 
frigidus sp. n. iii 619 
glabripennis Sharp iii 630 
hawaiiensis sp. n. iii 619 
hirsutulus sp. n. iii 640 
hypocoelus sp. n. iii 633 
ignotus sp. n. iii 626 
irregularis sp. n. iii 624 
kauaiensis sp. n. iii 626 
konanus sp. n. iii 631 
laevis sp. n. iii 625 
lanaiensis sp. n. iii 620 
latifrons sp. n. iii 624 
lugubris sp. n. iii 627 
marginatus sp. n. iii 634 
maurus sp. n. iii 627 
molokaiensis sp. n. iii 626 
montauus sp. n. iii 623 
muticus Sharp iii 634 



INDEX 



29 



Mirobternus nigrocastaneus sp. n. iii 629 

obscurus Sharp iii 642 

oculatus sp. n. iii 616 

pallidicornis sp. n. iii 630 

parcus sp. n. iii 632 

parvulus sp. n. iii 628 

peles sp. n. iii 630 

plebeius sp. n. iii 636 

punctatissimus sp. n. iii 617 

punctatus Sharp iii 635 

pusillus sp. n. iii 624 

pyrophilus sp. n. iii 618 

rufescens sp. n. iii 637 

rugipennis sp. n. iii 623 

sciilptus sp. n. iii 622 

simplex sp. n. iii 623 

solidus sp. n. iii 642 

solitarius sp. n. iii 635 

sordidus sp. n. iii 618 

stenarthrus sp. n. iii 641 

subparcus sp. n. iii 633 

testaceus sp. n. iii 617 

tetragonus sp. n. iii 637 

tristis sp. n. iii 639 

varicolor sp. n. iii 622 

varius sp. n. iii 640 

vestitus sp. n. iii 639 

xanthostictus sp. n. iii 629 
Miscogasteridae cv, i 307 
Misumena anguhventris sp. n. ii 488 

anguhventris Sim. ii 486, iii 34 r 

cretacea sp. n. ii 487 

nesiotes Sim. ii 485, iii 341 

nigrofrenata sp. n. ii 486 

oreades sp. n. ii 484, 485, iii 341 

velata sp. n. ii 489 
Mollica microphthalmus Koch ii 512 
Mollusca ii 271 
Mollusca, Review of cc.\.\iii 
Molokai, island xi.\ 
Monanus Sharp iii 429 

brevicornis Blacklj. iii 429 

crenatus Sharp iii 429 
Monochroa Hein. i 478 
Monocrepidiiis exsul Sharp iii 369 
Monodonta seminigra Lam. ii 298 
Monolexis ? palliatus Cam. i 362 
Monomera iii 102 
Monomorium floricola i 119 

minutum liliuokalanii i lig 

vastator i 119 
Monopis Hb. i 727, 737 

crocicapitella Clms. i 728, 737 

ferruginella Wlsm. i 728 

hering"i Rdsn. i 728 

hyahnella Stgr. i 728 

lombardica Hrng. i 728 

longella Wkr. i 727 

mediclla F. i 727 

monachella Hb. i 727 

obumbrata Btl. i 728 
Monotomidae iii 434 
Mopsus Keyserl. ii 512 
Moranila testaceiceps Cam. i 324 
Muscidae clxxxvi, iii 27, 83 
Musca annonae Fabr. iii 44 

azurea Fhi. iii 27 

caesar Linn, iii 28 

canicularis Linn, iii 30 

casei Linn, iii 48 

domestica Linn, iii 29 



Musca flavinervis Thoms. iii 29 

putris Linn, iii 48 

regina Meig. iii 28 

tenax Linn, iii 19 

vomitoria Linn, iii 27 
Mycetaeidae iii 421 
Mycetaea hirta Marsh iii 421 
Mycetophagidae ( = Tritomidae) iii 418 
Mycetophilidae clxxx, iii 2 
Myllaena Erichs. iii 551 

apetina sp. n. iii 553 

cognata sp. n. iii 553 

curtipes Sharp iii 552 

discedens Sharp iii 551 

familiaris Sharp iii 553 

haleakalae sp. n. iii 552 

oxypodina sp. n. iii 553 

pacifica Blackb. iii 553 

rufescens sp. n. iii 552 

sp..' iii 551 

vicina Sharp iii 552 
Mymaridae civ, ii 660 
Myodochidae cxciii^ ii 535 
Mysticomenus gen. n. iii 212 

mysticus Blackb. iii 213 

tibialis sp. n. iii 212 
Mytilaspis citricola Green iii iio 

flava Targ. iii 1 1 1 

flava var. hawaiiensis Mask, iii iii 

gloverii Mask, iii 1 1 1 

gloverii var. pallida Green iii in 

pallida Mask, iii 1 1 1 

pinnaeformis Nevvst. iii no 

pomorum Comst. iii n i 
Myriopoda iii 323 

Review of ccxxi 
Myrmecophila quadrispina sp. n. ii 14 
Myrmeleonidae clxxxiii 
Myrmeleon perjurus Walk, ii 61 

violentus Walk, ii 61 
Myrmicidae ci 



Nabidae iii 153, ii 546, cxcv 

Nabis Leth. & Sev. iii 153 

blackburni White iii 155 
capsiformis Germ, ii 546 
curtipennis Blackb. iii 157, ii 549 
innotatus White iii 154 
koelensis Blackb. ii 547, iii 156 
lusciosus White ii 548, iii 157 
oscillans Blackb. ii 547, iii 156 
rubritinctus Blackb. ii 549, iii 157 
subrufus White iii 156 

Naeogeidae iii 168 

Nanina discus Pfeiff. ii 283 
rinkii Morch. ii 279 

Nannodia Hein. i 478 

Nauphoeta bivittata Burm. ii 7 

Nausibius dentatus Marsh iii 429 

Neanura MacGill. iii 302 

citronella sp. n. iii 303 

Necremnus maculatipennis sp. n. i 331 

Necrobia ruficoUis Fabr. iii 367 
rufipes Fabr. iii 367 

Neda abdominalis iii 414 

oculata Fabr. iii 414 

Neelysia gen. n. i 532 

agnetella sp. n. i 538 
anthinella sp. n. i 535 
argyresthiella sp. n. i 544 



FAUNA HAWAIIENSIS 



Neelysia basivittata sp. n. i 539 
cleodorella sp. n. i 535 
complanella sp. n. i 546 
cuprea sp. n. i 532 
e.\altata sp. n. i 542 
fuscodentata sp. n. i 534 
fuscofusa sp. n. i 538 
incongrua sp. n. i 546 
lignicolor sp. n. i 533 
mactella sp. n. i 545 
municeps sp. n. i 537 
nemoricola sp. n. i 544 
ningorella sp. n. i 540, 543 
paltodorella sp. n. i 537 
petalifera sp. n. i 541 
pluviella sp. n. i 540 
poeciloceras sp. n. i 542 
psaroderma sp. n. i 536 
rediviva sp. n. i 543 
repandella sp. n. i 536 
rotifer sp. n. i 539 
sciurella sp. n. i 546 
semifusa sp. n. i 547 
subaurata sp. n. i 541 
terminella sp. n. i 547 
tigrina Btl. i 533 
tischeriella sp. n. i 545 

Nematoda ii 428 

Neoclytus Horn ii 104 

Neoe.xaireta spinigera Wied. iii 79 

Neolelaps gen. n. i 312 

flavipes sp. n. i 313 
hawaiiensis sp. n. i 313 

Neoliodes=Liodes Heyd. iii 704 

theleproctus Herm. iii 704 

Nereine spinosa Cambr. ii 460 

Neripteron gigas Less, ii 39S 

Neritidae ii 398 

Neritina Lam. ii 39S 

cariosa Gray ii 398 
conve.xa Nutt. ii 398 
granosa Sow. ii 398 
lugubris Phil, ii 398 
neglecta Pease ii 399 
nuttalli Reel, ii 398 
papillosa Jay ii 398 
sandwichensis Uesh. ii 398, 399 
solidissima .Sow. ii 398 
vespertina Nutt. ii 399 

Nesamiptis obsoleta Bull, i 157 
plagiota sp. n. i 156 

Nesapterus gen. n. iii 450 
exilis sp. n. iii 451 
monticola Sharp iii 451 

Nesidiocheilus, gen. n. iii 127 

hawaiiensis sp. n. iii 127 

Nesidiolestes gen. n. iii 152 
insularis Kirk, ii 551 
sehum sp. n. iii 153 

Nesidiorchestes gen. n. iii 139 

hawaiiensis sp. n. iii 139 

Nesiomiris gen. n. iii 144 

hawaiiensis sp. n. iii 145, ii 554 

Nesochhde epi.xantha Perk, iii 349 

Nesocidium gen. n. iii 280 

atomarium sp. n. iii 284 
. corticarium sp. n. iii 283 
fulgens sp. n. iii 282 
koebelei sp. n. iii 282 
laeticulum sp. n. iii 280 
lahainense sp. n. iii 281 



Nesocidium perkinsi sp. n. iii 282 

rude sp. n. iii 283 

scydmaenoides sp. n. iii 284 

smaragdinum sp. n. iii 282 
NesocHmacias contracta ii 535 

lanaiensis ii 535 
Nesocrabro gen. n. i 25, ii 606 

adspectans Blackb. i 29, ii 606 

bidecoratus sp. n. i 27, ii 606 

compactus sp. n. i 25 

daemonius sp. n. i 28, ii 606 

rubrocaudatus Blackb. i 27, ii 606 

stygius Bl. & C. i 29 
Nesocryptias villosa ii 535 
Nesocymus Kirk, ii 535 

calvus White ii 536 
Nesodiranchis subgen. n. ii 668 
Nesodryas Kirk, ii 596 

dryope sp. n. ii 597 

elaeocarpi Kirk, ii 596 

eugeniae Kirk, ii 597 

gififardi Kirk, ii 597 
Nesodynerus Perk, ii 607 

conifer Perk, ii 607 

cooki Perk, ii 60S 

dilatatipes Perk, ii 608 

eupteryx Perk, ii 608 

freycinetiae Kirk, ii 596 

oblitus Perk, ii 608 

optabilis Perk, ii 608 

paractias Perk, ii 608 

rudolphi D.T. ii 607 

vittativentris Perk, ii 608 
Nesogryllus gen. n. ii 26 

stridulans sp. n. ii 27 
Nesoligota subg. n. Oligota iii 557 
Nesolymnaeum gen. n. iii 277 

spurcum Blackb. iii 277 
Nesomartis psammophila Kirk, ii 535 
Nesomedon gen. n. iii 546 

brunnescens iii 546 

oahuensis sp. n. iii 547 

quadratus sp. n. iii 547 
Nesomicromus gen. n. ii 37 

angularis sp. n. ii 691 

angustipennis sp. n. ii 38 

belkilus sp. n. ii 40 

brunnescens sp. n. ii 43 

distinctus sp. n. ii 44 

drepanoides sp. n. ii ' 39 

forcipatus sp. n. ii 44 

fulvescens sp. n. ii 39 

haleakalae sp. n. ii 42 

infumatus sp. n. ii 41 

latipennis sp. n. ii 38 

longispinosus sp. n. ii 42 

minimus sp. n. ii 45 

minor sp. n. ii 41 

molokaiensis sp. n. ii '41 

ombrias sp. n. ii 692 

parado.xus sp. n. ii 39 

phaeostictus sp. n. ii 692 

rubrinervis sp. n. ii 43 

stenopteryx sp. n. ii 45 

subochraceus sp. n. ii 44 

\agus sp. n. ii yj 
Nesomicrops gen. n. iii 286 

kauaiensis sp. n. iii 286 
Nesominiesa gen. n. i 8 

antennata i 10 

hawaiiensis i 1 1 



INDEX 



ly 



Nesomiinesa kauaiensis sp. n. i 9 

nitida i 1 1 

sciopteryx i 10 
Nesopeplus gen. n. iii 474 

abnormalis sp. n. iii 4S1 

anticatus sp. n. iii 476 

ater sp. n. iii 491 

bidens Sharp iii 483 

cognatus sp. n. iii 487 

coUaris sp. n. iii 476 

confertus sp. n. iii 490 

cunithorax sp. n. iii 476 

fallax sp. n. iii 479 

floricola Blackb. iii 480 

inauiatus Sliarp iii 475 

insolitus sp. n. iii 482 

koelensis iii 487 

lambiamis sp. 11. iii 484 

latiusculus sp. n. iii 488 

molokaiensis sp. n. iii 478 

nigricans sp. n. iii 487 

obscurans sp. n. iii 486 

olindae Blackb. iii 477 

pictus sp. n. iii 480 

protinoides Sharp iii 486 

puncticollis sp. n. iii 490 

roridus sp. n. iii 479 

segnis sp. n. iii 477 

serratus sp. n. iii 4S2 

simiHs sp. n. iii 489 

sinuatus sp. n. iii 491 

solitarius sp. n. iii 481 

sp. ? iii 488 

testaceipes sp. n. iii 484 

torvus Blackb. iii 485 

vagepictus sp. n. iii 478 
Nesopetinus gen. n. iii 492 

apertus .Sharp iii 498 

blackburni Sharp iii 503 

blackburni var. ianaiensis Blackb. iii 504 

blackburni var. mauiensis var. n. iii 504 

blackbiirnianus sp. n. iii 504 

celatus Sharp iii 498 

concolor sp. n. iii 500 

discedens Sharp iii 494 

eremitus sp. n. iii 497 

filipes sp. n. iii 501 

gonioryctoides sp. n. iii 493 

intermedins sp. n. iii 503 

kauaiensis Blackb. iii 495 

metallescens Sharp iii 493 

omissus sp. n. iii 495 

pallidus sp. n. iii 500 

parallelus Blackb. iii 497 

perkinsi sp. n. iii 502 

pusillus sp. n. iii 496 

quadraticoUis Blackb. iii 501 

rudis sp. n. iii 502 

scottianus sp. n. iii 499 

sp. .■■ iii 496, 500 

tinctus Sharp iii 493 

varius Sharp iii 496 

vestitus Sharp iii 497 
Nesophila Pilsbry ii 289 
Nesophrosyne Kirk, ii 556, 558 

arcadiicola sp. n. ii 571 

bobeae sp. n. ii 564 

caeHcola sp. n. ii 566 

craterigena sp. n. ii 571 

ehu sp. n. ii 569 

filicicola Kirk, ii 556 



Nesophrosyne giffardi sp. n. ii 563 

gouldiae sp. n. ii 560 

haleakala sp. n. ii 567 

halemanu sp. n. ii 559 

ignigena sp. n. ii 570 

imbricola sp. n. ii 566 

insularis sp. n. (Nesoreias) ii 573 

kaiamamao Kirk, ii 557 

koleae sp. n. ii 562 

kukanaroa Kirk, ii 557 

maritima sp. n. ii 560 

microlepiae sp. n. ii 557 

milu sp. n. ii 565 

monticola sp. n. ii 562 

montium sp. n. ii 569 

montivaga sp. n. ii 569 

myrsines sp. n. ii 568 

nigrolineata var. n. ii 563 

nimbicola sp. n. ii 565 

nimbigena sp. n. ii 567 

nubigena sp. n. ii 567 

nuenue sp. n. ii 572 

oceanides sp. n. {Nesoreias) ii 573 

oneanea sp. n. ii 566 

opalescens sp. n. ii 561 

oreadis sp. n. ii 569 

paludicola sp. n. ii 564 

palustris sp. n. li 564 

pele sp. n. ii 570 

perkinsi Kirk, ii 559 

pipturi sp. n. ii 560 

pluvialis sp. n. ii 568 

ponapona sp. n. ii 561 

procellaris sp. n. ii 565 

silvicola sp. n. ii 570 

silvigena sp. n. ii 559 

sp. ? ii 562 

ulaula sp. n. ii 563 

umbratilis sp. n. ii 558 

umbricola sp. n. ii 565 

unibrigena sp. n. ii 571 
Nesopleias gen. n. ii 582 

dubautiae sp. n. ii 583 

nimbata sp. n. ii 582 
Nesoprosopis gen. n. i 75, ii 601, l.\iv 

andrcnoides sp. n. i in 

angustula sp. n. i 95 

anomala sp. n. i 112 

anthracina i 100 

assimulans sp. n. i loi 

assimulans oahuensis i 102 

blackburni i 97 

caeruleipennis sp. n. i 107 

chlorosticta sp. n. i 78 

comes sp. n. i 90 

coniceps i 91 

connectens sp. n. i 85 

crabronoides sp. n. i 94 

ditficilis sp. n. i 80 

dimidiata sp. n. i 96 

dumetorum sp. n. i 92 

erythrodemas sp. n. i 112 

facilis i 77 

finitima sp. n. i 100 

flavifrons i loi 

flavipes i 99, ii 605 

fuscipennis i 106 

fuscipennis obscuripes i 107 

haleakalae sp. n. i 87 

hilaris Sm. i 103 

hirsutula sp. n. i 79 



FA UNA HA J I AIIENSIS 



Nesoprosopis hostilis sp. n. i 104 
inquilina sp. n. i 102 
insignis sp. n. i no 
kauaiensis sp. n. i 90 
koae sp. n. i 85 
kona i 82 
laeta sp. n. i 81 
laticeps sp. n. i 88 
longiceps sp. n. i 98 
mauiensis sp. n. i 94 
melanothrix sp. n. i 86 
mutata sp. n. i 93 
neglecta sp. n. i 89 
nivalis sp. n. i 83 
obscurata sp. n. i 99 
ombrias sp. n. ii 604 
paradoxica sp. n. i III 
perspicua sp. n. i 109 
pubescens sp. n. i 107 
rugulosa sp. n. i 84 
satelles i 109 
setosifrons sp. n. i 108 
simplex sp. n. i 79 
specularis sp. n. i 93 
sphecodoides sp. n. i 105 
unica sp. n. i 88 
vicina sp. n. i 84 
volatilis i 105 
volcanica sp. n. i 83 

Nesoreias subg. n. ii 573 

Nesorestias Kirk, ii 583 

filicicola Kirk, ii 583 

Nesosteles Kirk, ii 573 

hebe Kirk, ii 574 
peregrina sp. n. ii 575 
plutonis sp. n. ii 574 
volcanicola sp. n. ii 574 

Nesosydne Kirk, ii 583 

argyroxiphii Kirk, ii 590 
chambersi Kirk, ii 590 
cyathodis sp. n. ii 589 
gouldiae sp. n. ii 586 
haleakala sp. n. ii 587 
halia Kirk, ii 584 
hamadryas sp. n. ii 587 
imbricola sp. n. ii 590 
ipomoeicola Kirk, ii 5S6 
koae Kirk, ii 583 
monticola sp. n. ii 591 
nephelias sp. n. ii 588 
nephrolepidis Kirk, ii 586 
nubigena sp. n. ii 589 
palustris Kirk, ii 589 
pele sp. n. ii 585 
pipturi Kirk, ii 584 
procellaris sp. n. ii 588 
raillardiae Kirk, ii 590 
rubescens Kirk, ii 584 
umbratica sp. n. ii 585 

Nesothauma gen. n. ii 46, xlix 
haleakalae sp. n. ii 47 

Nesothoe Kirk, ii 591 

bobeae Kirk, ii 593 
flatus Kirk, ii 592 
frigidula Kirk, ii 593 
hula Kirk, ii 592 
laka Kirk, ii 594 
perkinsi Kirk, ii 593 
pluvialis Kirk, ii 595 
pulani Kirk, ii 594 
silvestris Kirk, ii 595 



Nesothoe terryi Kirk, ii 594 
Nesothrips Kirk, ii 681 

oahuensis Kirk, ii 682 
Nesotocus gen. n. ii 150 

giffardi sp. n. iii 654 

kauaiensis sp. n. ii 151 

munroi sp. n. ii 150 

newelli sp. n. ii 151 
Nesotyphlias Kirk, ii 548 

lusciosus Kirk, ii 548 
Neuroptera ii 31, 691 

Review of clxx 

flightless xlix 
Newcombia Pfeifif. ii 331 

canaliculata Bald, ii 331 

chlorotica Pfeiff. ii 344 

cinnamomea Pfeiff. ii 331 

cumingi Newc. ii 331 

gemma Pfeiff. ii 331 

perkinsi Sykes ii 332 

pfeifferi Newc. ii 332 

philippiana Pfeiff. ii 332 

plicata Pfeiff. ii 332 

sulcata Pfeiff. ii 332 

zebrina Pfeiff. ii 331 
Niihau, island xvi 
Niihoa, Bird island xvii 
Niphonides ii 1 14 
Nirmus diaprepes sp. n. iii 309 

gloriosus emarginatus var. n. iii 310 

minhaensis sp. n. iii 307 

orarius hawaiiensis var. n. iii 310 

stenozonus sp. n. iii 308 
Nitidulidae cxxxiv, iii 435 
Nitidula dimidiata Fabr. iii 507 

humeralis Fabr. iii 507 
Nomophila Hiib. i 227, iii 361 

noctuella Schiff. i 227, iii 361, xlvii 
Nomia Clem, i 478 
Notaspis lucorum Koch, iii 704 
Noteraula straminea Meyr. i 687 
Notiphila argentata Walk, iii 49 

insularis sp. n. iii 49 
Notonectidae cciii, iii 148 
Notonecta pallipes Fabr. ii 555 
Notopeplus gen. n. iii 505 

reitteri Sharp iii 505 
Number of species xli 
Numenius Briss. i 451 

tahitiensis Gni. i 452 
Nycticorax Raf i 456 

griseus Linn, i 456 
Nymphalidae cliv 
Nymphula Schranck. i 200, iii 356 

fluctuosalis Zell. (linealis Guen.) i 200, iii 356 
Nysius Dall. ii 536, iii 162 

arboricola White ii 538 

atralis var. n. ii 541 

blackburni White ii 540 

brunnealis var. n. ii 541 

coenosulus StAl. ii 540 

dallasi White ii 538 

delectus White ii 538 

hylaeus sp. n. ii 539 

insulicola sp. n. ii 541 

insulivagus sp. n. ii 544 

kamehameha sp. n. iii 164, ii 539 

lichenicola sp. n. ii 540 

longicolHs Blackb. ii 538 

mauiensis Blackb. ii 538 

monticola sp. n. ii 544 



INDEX 



33 



Nysius montivagus sp. n. ii 544 

nemorivagus White ii 543 
nimbatus sp. n. ii 543 
nitidus White ii 543 
nubicola sp. n. ii 542 
ochriasis sp. n. iii 162, ii 541 
oresitrophus sp. n. ii 542 
oribasus sp. n. ii 544 
pteridicola White ii 543 
rubescens \\'hite ii 543 
saundersianus sp. n. iii 163, ii 537 
silvestris sp. n. ii 541 
vuIcan White ii 543 
whitei Blackb. ii 538 



Oahu, island .wii 

Ocean currents xlvii 

Oceanodroma cryptoleucura Ridg. i 463 

Odonata ii 62, ii 693, clxxv 

Odynerus i 30, xc, clx 

acoelogaster sp. n. i 65, ii 6og 
aeyanus Perk, ii 611 
agilis Sni. i 41 
aprepes Perk, ii 611 
axestes sp. n. i 66 
blackburni i 58 
brevicostatus sp. n. i 46 
caenosus sp. n. i 35 
camelinus sp. n. i 44 
cardinahs Bl. i 46 
cephalostictus sp. n. i 66 
chelifer sp. n. i 70, ii 607 
conifer sp. n. i 38 
congruiis i 69, ii 607 
cooki sp. n. i 64, ii 608 
cyanopteryx sp. n. i 60, ii 612 
cyphotes sp. n. i ^li 
cypris Perk, ii 612 
deinogaster sp. n. i 48 
dilatatipes sp. n. i 37, ii 608 
diversus Bl. i 45 
dromedarius i 45 
dryas sp. n. i 53 
dubiosus i 62 
dyserythrias sp. n. i 60 
ecostatus sp. n. i 42 
egens sp. n. i 61, ii 609 
eludens sp. n. i 36 
epipseustes Perk, ii 61 1 
erro sp. n. i 31 
erythrognathus sp. n. i 32 
erythrostactes sp. n. i 53 
eucharis Perk, ii 610 
eupteryx sp. n. i 47, ii 608 
eutretus Perk, ii 612 
flosculus sp. n. i 64 
frater D.T. i 59 
heterochronius sp. n. i 49 
hiloensis sp. n. i 34 
holomelas Perk, ii 612 
homochromus sp. n. i 37 
homoeogaster sp. n. i 49 
homoeophanes Perk, ii 610 
illudens Perk, ii 609 
infaustus sp. n. i 73, ii 609 
instabilis sp. n. i 48 
insulicola i 67 
iopteryx sp. n. i 63, ii 609 
kauaiensis sp. n. i 38 
kirbyi i 37 

F. H. 



Odynerus konanus sp. n. i 34 

laevisLilcatus sp. n. i 51 

lanaiensis sp. n. i 68 

leiodemas sp. n. i 51 

leucozonias sp. n. i 36 

lipocharis sp. n. ii 610 

lithophilus Perk, ii 609 

localis i 71 

niaurus Sm. i 70 

melanognathus sp. n. i 33 

mesospilus Perk, ii 612 

microdemas sp. n. i 55 

mimus sp. n. i 50 

molokaiensis sp. n. i 41 

monas Perk, ii 611 

monobius sp. n. i 55 

montanus i 43 

montivagus sp. n. i 54 

naiadum sp. n. i 32 

nautarum sp. n. i 72 

nautarum D.T. i 67 

nesiotes sp. n. i 61 

nesotrephes sp. n. i 68 

newelli Perk, ii 612 

nigripennis i 70 

nivicola sp. n. i 52 

nubicola sp. n. i 42 

oahuensis i 45 

oblitus sp. n. i 74, ii 608 

obscurepunctatus i 59 

orbus sp. n. i 35 

pacificus Bl. i 59 

paludicola Perk, ii 6ii 

paranaias Perk, ii 611 

peles sp. n. i 39, ii 610 

petrobius sp. n. i 57 

potamophilus sp. n. i 56 

pseudochromoides Perk, ii 611 

pseudochromus sp. n. i 50 

pterocheiloides sp. n. i 69, ii 607 

pterophaennes sp. n. i 63 

purpurifer sp. n. i 47 

radula i 71 

relictus sp. n. i 74, ii 609 

rubritinctus i 56, 58 

rubropustulatus i 57 

rudolphi i 46 

rudolphi D.T. ii 607 

sandvvichensis sp. n. i 73 

sandwichensis D.T. i 56 

scoriaceus sp. n. i 40 

smithii i 41 

sociabilis sp. n. i 39 

sorer sp. n. i 58 

subegens sp. n. ii 610 

tempe Perk, ii 611 

thersites sp. n. ii 609 

threnodes sp. n. i 62 

unicus sp. n. i 43 

venator sp. n. i 31 

vittativentris sp. n. i 65, ii 608 

vulcanus i 34 

waianaeanus sp. n. i 64 

xanthorhoes sp. n. i 52 

xerophilus Perk, ii 611 
Oechalia ii 533, iii 171 

grisea Burni. ii 533, in 171 

patruelis iii 171 

pacifica White iii 171 
Oecia maculata Wlsm. i 649 
Oecophoridae Meyr. i 507, clxiv 



34 



FAUNA HAWAIIENSIS 



Oecophora hydeniella Fr. i 514 

phycidella Z. (Scythris) i 648 

Oedemeridae ii 247 

Oedemothrips gen. n. iii 680 
laticeps sp. n. iii 680 

Oemides ii 96 

Oenemona Motsch. ii 104 

Oestrelata i 462 

phaeopygia Salv. i 462 

Oestridae iii 20 
. Oestrus ovis Linn, iii 20 

Olethreutinae i 673 

Olfersia acarta sp. n. iii 87 

Oliarus StAl iii 119 

hevaheva sp. n. iii 122 
kanakanus sp. n. iii 121 
koanoa sp. n. iii 124 
opuna sp. n. iii 122 
orono sp. n. iii 124 
tamehameha sp. n. iii 120 
tarai sp. n. iii 123 

Oligochaeta ii 413 

Oligostigina curta Bull, i 200 

Oligota Mann, iii 554, iii 560 

adpropinquans sp. n.- iii 561 

aethiops sp. n. iii 563 

anomalocera sp. n. {Gnatholigota) iii 556 

bicolor sp. n. {Nesoligota) iii 560 

brevicollis sp. n. i^GnathoUgotal) iii 556 

clavicornis Sharp [Holobus) iii 555 

cribripennis sp. n. {Nesoligota) iii 559 

currax sp. n. {Nesoligctn) iii 559 

extranea sp. n. [Gnatholigota) iii 557 

frontalis sp. n. iii 561 

glabra Sharp (Nesoligota) iii 559 

gyninusa sp. n. iii 561 

haleakalae sp. n. iii 562 

insolita sp. n. iii 561 

kauaiensis Blackb. {Nesoligota) iii 558 

latifrons sp. n. {Giiatlioligota) iii 556 

latipennis sp. n. {Nesoligota) iii 558 

longipennis Blackb. iii 563 

niutanda .Sharp iii 562 

oahuensis sp. n. iii 562 

parca sp. n. {Gnatholigota) iii 557 

pernigra sp. n. iii 563 

polita Sharp {Nesoligota) iii 55S 

prolixa Sharp {Deroligota) iii 555 

scripta sp. n. iii 560 

scripta var. laetior var. n. iii 560 

simulans Blackb. iii 562 

tenuicornis sp. n. iii 563 

variegata Blackb. iii 563 

Oligotoma insularis McLachl. ii 88, clxxiv 

Olisthopus insularis Karsch. iii 253 

Omicrus brevipes .Sharp iii 579 

Omiodes Guen. i 201, iii 357 

accepta Butl. i 204, iii 358 
anastrepta sp. n. i 204, iii 358 
antidoxa sp. n. iii 358 
asaphombra sp. n. i 202 
blackburni Butl. i 202, iii 357 
continuatalis Wall, i 205, iii 358 
demaratalis Walk, i 205, iii 358 
epicentra sp. n. i 203, iii 357 
euryprora sp. n. i 202 
iridias sp. n. i 203 
localis Butl. i 206, iii 358 
monogona Meyr. i 206 
monogramma sp. n. i 205, iii 35S 
telegrapha sp. n. i 204 



Omphale metallicus sp. n. i 327 

Oncophorus advena Kell. iii 311 

Oniscosoma pallida Brunn. ii 7 

Oodemas Bohem. ii 155 

aenescens Bohem. ii 169 
aeolosoma sp. n. ii 169 
aequale Blackb. ii 161 
affine sp. n. ii 162 
angustum Blackb. ii 169 
apionoides sp. n. ii 161 
borrei Blackb. ii 170 
brunneuni sp. n. ii 159 
chrysodorum sp. n. ii 158 
corticis sp. n. ii 168 
costatum sp. n. ii 157 
crassicorne Blackb. ii 160 
cupreum sp. n. ii 159 
dilatatipes sp. n. ii 156 
dubiosum sp. n. ii 164 
flexirostre sp. n. ii 164 
gracilifonne sp. n. ii 163 
grande sp. n. ii 167 
haleakalae sp. n. ii 160 
halticoides Blackb. ii 169, iii 654 
hauaiiense sp. n. iii 655 
infernum Blackb. ii 172 
insulare Blackb. ii 170 
konanum sp. n. ii 171 
leithorax sp. n. ii 164 
longicorne sp. n. ii 157 
longirostre sp. n. ii 155 
mauiense Blackb. ii 170 
molokaiense sp. n. ii 15S 
montanum sp. n. ii 160 
multiforme sp. n. ii 171 
nitidissimum sp. n. ii 170 
nivicola Blackb. ii 156 
oblongum sp. n. ii 167 
obscurum Blackb. ii 161 
olindae Blackb. ii 155 
pachysoma sp. n. ii 168 
parallelum sp. n. ii 162, iii 654 
pulchrum sp. n. ii 166 
puncticoUe sp. n. ii 165 
punctulatissimum sp. n. ii 157 
purpurascens sp. n. ii 16^ 
ramulorum sp. n. ii 163 
robustum Blackb. ii 169, iii 654 
sculpturatum Blackb. ii 170 
soliclum sp. n. iii 654 
striatum sp. n. ii 165 
substrictum Blackb. ii 161 
tardum Blackb. ii i5i 

Oopsis nutator Fabr. ii 114 

Opatrum seriatum Boisd. ii 252 

Opeas junceus Gould ii 383 
prestoni Sykes ii 384 
pyrgiscus Pfeifif. ii 384 
striolata Pease ii 384 

Ophelinus mauiensis sp. n. i 329 
hawaiiensis sp. n. i 330 

Ophioniedon Sharp iii 545 

incomptus Sharp iii 545 
subtilis sp. n. iii 545 

Ophioninae ii 677 

Ophion lineatus Cam. i 341 
nigricans Cam. i 341 

Ophthalmomyia lacteipennis Loew iii 74 

Ophyra aenescens Wied. iii 30 
leucostoma Wied. iii 30 

Opisthacantha dubiosa sp. n. ii 623 



INDEX 



35 



Opogona aurisqiiamosa Btl. i 713, 737 

diniidiatella Z. i 712 
Opostega dives sp. n. i 711 

maculata sp. n. i 711 
Qpsicoetus biannulipes Montr. & Sign, iii 150 
Opuna gen. n. iii 140 

hawaiiensis sp. n. iii 140, ii 553 
Orchestia hawaiensis Dana ii 529 

pickeringii Dana ii 528 

platensis Kroy. ii 527 
Oreomyza Stejn. i 413 

bairdi Stejn. i 416 

flammea Wils. {Paroreomyza) i 417 

maculata Cab. {Parorcomyza) i 417 

niana Wils. i 416 

montana Wils. {ParorMiitysa) i 417 

newtoni Rothsch. {Paroreomyza) i 417 

perkinsi Rothsch. i 417 
Oribatidae iii 702 
Oribala alata Herm. iii 703 

globula Xic. iii 702 

lapidaria Luc. iii 703 

oriformis sp. n. iii 703 

ovalis Nic. iii 703 
Origin of fauna Iii 
Orneodidae i 477, 731, clxiii 



Orneodes Ltr. 



477. 731 



objurgatella sp. n. i 477, 731 

Ornithoica confluenta Say. iii 91 
peroneura var. n. iii gi 

Ornithomyia varipes Walk, iii 89 

Oronomiris gen. n. iii 144 
hawaiiensis iii 144 

Orothreptes gen. n. ii 147 

callithrix sp. n. ii 147, iii 655 

Ortalidae iii 44, iii 85, clx.wvii 

Ortalis annonae Wied. iii 44 

Orthocladius sp. iii 5 

Orthodera prasina Burm. ii 7 

Orthoea Dall. iii 159 

nigriceps Dall. iii 159 
pacifica Kirk, ii 545 
periplanios Kirk, ii 545 
\incta Say ii 545 

Orthoniecyna Butl. i 240, iii 362 
albicaudata Butl. i 243 
alloptila sp. n. i 243 
aniphilyca sp. n. i 245 
aphanopsis Meyr. i 244 
chrysophanes sp. n. i 243 
crossias sp. n. i 242, iii 362 
cupreipennis Butl. i 245 
epicausta sp. n. i 244 
exigua Meyr. i 245 
heterodyas sp. n. i 242 
niesochasma sp. n. i 244, iii 362 
nietalycia sp. n. i 241 
phaeophanes sp. n. i 241 
pi erodes sp. n. i 244 

Orthomorpha gracilis Koch iii 328 

Orthoperiis aequalis Sharp iii 417 

Orthoptera ii i, ii 687 
Review of ccxi 

Orthostolus gen. n. iii 452 

atratus sp. n. iii 456 
expers Blackb. iii 455 
germanus sp. n. iii 454 
guttatus Sharp iii 454 
kauaiensis sp. n. iii 456 
nepos sp. n. iii 453 
prosternalis sp. n. iii 454 



Orthostolus robustus Sharp iii 452 

robustus var. lanaiensis var. n. iii 453 

sordidus Sharp iii 455 
Orthotylus Fieb. iii 132, ii 552 

azalais sp. n. iii 136 

daphne sp. n. iii 135 

daphne var. kassandra Kirk, ii 553, iii 135 

iolani sp. n. iii 133 

kanakanus sp. n. iii 134 

kekele sp. n. iii 134 

perkinsi sp. n. iii 133 
Otiorhynchini ii 122 
Oxacis coUaris Sharp ii 247, cxxvi 
Oxya velox Fab. ii 8 
Oxytelini iii 543 
Oxytelus advena Sharp iii 543 

bledioides Blackb. iii 543 

ferrugineus Kraatz iii 543 

pygmaeus Kraatz iii 543 

sp. iii 543 
Oxyurus vermicularis Lutz ii 428 

Pachymerinae iii 159 
Pachyneuron Walk, ii 656 
Paederini iii 544 
Paedisca immersana Wkr. i 690 
Pagiopalus gen. n. ii 499 . 

atomarius sp. n. ii 500, iii 343 

nigriventris sp. n. ii 502 

personatus sp. n. ii 501 

semipunctatus sp. n. ii 501 
Palmeria i 405, Ixxii 

dolii Wils. i 407 
Paludestrinidae ii 396 
Paludestrina porrecta Migh. ii 396 
Paludina porrecta Migh. ii 396 
I'amera Leth. & Sev. iii 159 

nigriceps St;\l iii 159 

vincta Stal ii 545 
Panaphelix gen. n. i 695 

marmorata sp. n. i 696 

sp. ? i 697 
Panchlora maderae F. ii 690 
Pandemis consociana Wkr. i 690 

secundana Wkr. i 690 
Pandion haliaetus Linn, i 447 
Pantala flavescens Fab. ii 62 
Pantomorus fuUeri Horn, iii 653, cxx 

olindae sp. n. ii 130, iii 653 
Papuina barnaclei Smith ii 292 
Parandra puncticeps Sharp ii 95 
Parandrita aenea Sharp iii 426 

gracilis sp. n. iii 427 

konae sp. n. iii 426 

liturata sp. n. iii 427 

molokaiae sp. n. iii 427 

perkinsi sp. n. iii 426 
Paranemobius schauinslandi Alfk. ii 688 
Paraphasis gen. n. i 730 

perkinsi sp. n. i 730 
Paraphelinus xiphidii Perk, ii 658 
Pararrhaptica gen. n. i 689 

perkinsana sp. n. i 689 
Paratenodera sinensis Sauss. ii 689 
Paratrigonidium Brunn. ii 15, xxviii, xxx, xlix, Ixviii 

atroferrugineum Brunn. ii 17 

attenuatum sp. n. ii 20 

crepitans sp. n. ii 19 

debile sp. n. ii 21 

exiguum sp. n. ii 21 

filicum sp. n. ii 17 

E 2 



FA UNA HA WAIIENSIS 



Paratrigonidium freycinetiae sp. n. ii i6 

grande sp. n. ii 19 

molokaiense sp. 11. ii 20 

pacificum Scudd. ii 22 

robustum sp. n. ii 20 

roseum sp. n. ii 16 

saltator sp. n. ii 16 

subroseum sp. n. ii 17 

varians sp. n. ii 18 

viridescens sp. n. ii 18 
Parlatoria proteus Run iii iio 

proteus van pergandii Comst. iii no 

ziiiyphi Newst. iii iio 
Paroi'chestia hawaiensis Dana ii 529 
Partula auricula Fdn ii 375 

densilineata Reeve ii 317 

dumartroyi Soul, ii 375 

pusilla Gould ii 379 

terrestris Pease ii 399 

virgulata Migh. ii 320 
Partulina Pfeiff. ii 311 

liayseldeni Bakhv. ii 313 

pioxinia Pease ii 316 
Patula digonophora Ancey ii 292 
Paucity of individuals xx.\viii 
Pelopaeus caementarius i 8 
Pemphredonidae ii 605, Ixxxiv 
Pennula Dole i 453 

ecaudata King i 454 
Pentaithron flavuni sp. n. ii 660 

semifumatum sp. n. ii 659 
Pentarthrum blackburni Sharp ii 147, iii 655 

obscurum Sharp ii 146 

prolixum Sharp ii 146 
Pentatoma insularis Dall. iii 172 
Perdicclla Pease ii 329 

fulgurans sp. n. ii 329 

helena Newc. ii 330 

mauiensis Pfeiff. ii 330 

minuscula Pfeiff. ii 330 

ornata Newc. ii 330 

theodorei Baldw. ii 330 

zebra Newc. ii 330 

zebrina Pfeiff. ii 331 
Peregrinus maidis Ashm. ii 577 
Perichaeta amazonica Rosa ii 420 

bermudensis Bedd. ii 420 

cupulifera Fed. ii 420 

floweri Benh. ii 414 

hawayana Rosa it 420 

hesperidum Bedd. ii 417 

heterochaeta Mich, ii 416 

indica Mich, ii 416 

mandliorensis Mich, ii 420 

mauritiana Bedd. ii 420 

morrisi Bedd. ii 420 

pallida Mich, ii 420 

peregrina Fletcli. ii 414 

sandvicensis Id. ii 417 

trityphla Bedd. ii 426 

vesiculata Goto lit Hat. ii 426 
Periplaneta americana L. ii 6 

australasiae Fab. ii 7 
Perkinsiella saccharicida Kirk, ii 578 
Peropus mutilatus Stejn. i 367 
Phaenopria ambulator sp. n. ii 627 

frater sp. n. ii 626 

hawaiiensis sp. n. i 296 

hylaea sp. n. ii 628 

niontana sp. n. ii 628 

soror sp. n. ii 627 



Phaenopria subtilis sp. n. ii 627 
Phaeogramnia gen. n. iii 47 

vittipennis sp. n. iii 48 
Phaeornis Sclater i 377, xxxi 

lanaiensis Wils. i 378 

myiadestina Stejn. i 378 

oahuensis Blox. i 378 

obscura Gniel. i 378 

palnieri Rothsch. i 379 
Phaethou lepturus L. & D. i 462 

rubricauda Bodd. i 462 
Phalaena curtisella Don. (coenobitella) i 651 
Phalainesthes Kirk, iii 117 

schauinslandi Kirk, iii 117 
Phalaropus fulicarius L. i 453 

lobatus Linn, i 453 
Phanerotoma hawaiiensis sp. n. i 360 
Pheidole megacephala Fab. i 118, xxxix, xli, ci 
Philodoria gen. n. i 717, 737 

auromagnifica sp. n. i 718 

basalis sp. n. i 720 

floscula sp. n. i 718 

niicropetala sp. n. i 719 

nigrella sp. n. i 721 

splendida sp. n. i 719, 737 

succedanea sp. n. i 717 
Philodroniinae ii 496 
Philomycus bilineatus Bens, ii 286 
Philonesia gen. n. ii 280 

abeillei Ancey ii 280 

baldvvini Ancey ii 281 

boettgeriana Ancey ii 283 

cicercula Gould ii 283 

exaequata Gould ii 283 

hartnianni Ancey ii 283 

indefinita Ancey ii 284 

lymanniana Ancey ii 284 

oahuensis Ancey ii 284 

perkinsi Sykes ii 284 

perlucens Ancey ii 284 

platyla Ancey ii 284 

plicosa Ancey ii 285 

sericans Ancey ii 285 

subrutila Migh. ii 285 

subtilissima Gould ii 285 

turgida Ancey ii 285 
Philonthus discoideus Grav. iii 548 

nauticus Fairm. iii 548 

nigritulus Grav. iii 548 

seybalarius Nordm. iii 547 

turbidus Gra\'. iii 548 
Philoscia angusticauda Budde-Lund ii 525 
Phloeophagosonia tenuis Gemm. ii 149 
Phloeopora cingulata Sharp iii 577 

diluta Sharp iii 577 
Phloeothripidae Hal. iii 677 
Phlyctaenia Hub. i 206, iii 359 

argoscelis Meyr. i 217 

calliastra sp. n. i 207 

caminopis sp. n. i 215 

chalcophanes sp. n. i 209 

chytropa sp. n. i 210 

despecta Bull, i 217, iii 360 

endopyra sp. n. i 219, iii 360 

ennychioides Butl. i 216, iii 359 

ephippias sp. n. i 210 

eucrena Meyr. i 213 

heterodoxa sp. n. i 220 

hyacinthis sp. n. i 208 

iocrossa sp. n. i 212 

lanipadias sp. n. iii 359 



INDEX 



Plilyctaenia liopis sp. n. i 218 

metasenia sp. n. i 214 

niicacea Butl. i 212, iii 359 

monticolans Butl. i 211, iii 359 

nigrescens Butl. i 211, iii 359 

ommatias sp. n. i 215 

pachygramnia sp. n. i 216 

platyleuca sp. n. i 214 

pyranthes sp. n. i 220 

rhodias sp. n. i 219 

stellata Butl. iii 359 

synastra sp. n. i 20S, iii 359 
Pholcidae ii 446 
Pholcus distinctus Cambr. ii 446 

elongatus Vins. ii 446 

margarita Workm. ii 446 

phalangioides Dol. ii 446 

tipuloides Koch ii 446 
Phora sp. n. iii 76 
Phormia regina Meig. iii 28 
Phoxopteris rufipennis Btl. i 680 
Phrynomorphini iii 555 
Phrynomorphus hospes Kirk, ii 576 
Phthoriniaea operculella Z. i 483, 485, 731 
Phycitidae iii 355, clvi 
Phyllodromia gernianica L. ii 5 

hieroglyphica Brunn. ii 5 

hospes sp. n. ii 5 

obtusta Brunn. ii 5 
Physa flavida Cless. ii 391 

hartmanni Cless. ii 392 

moreletiana Cless. ii 392 

naticoides Cless. ii 392 

peasei Cless. ii 393 

reticulata Gould ii 393 

sandwichensis Cless. ii 391 
Physopleurella mundula White ii 551, iii 126 
Physopus rubrocincta Giard iii 699 
Pieris rapae cliv 
Piestini iii 538 

Pilinothrix bicolor sp. n. i 299 
Pimpla Fabr. i 339, ii 676 

hawaiiensis Cam. i 339, ii 676 
Piophila casei Linn, iii 48 
Pipunculidae ii 697, iii 17, clxxxiii 
Pipunculus acrothrix sp. n. ii 698 

hawaiiensis Perk, ii 699 

holomelas sp. n. ii 698 

juvator Perk, ii 699 

molokaiensis sp. n. iii 17 

nigrotarsatus sp. n. iii 18 

oahuensis Perk, ii 699 

pyrophilus sp. n. ii 698 

rotundipennis sp. n. iii 18 

sp. iii 19 

sweezeyi Perk, ii 699 

terryi Perk, ii 699 

vulcanus sp. n. ii 697 
Pison hospes i 14 

iridipennis i 14 

sp. li 606 
Pitys elisae Ancey ii 290 
Plagithmysides ii 98 
Plagithmysus Motsch. ii 104, xxvi, cxvi 

aequalis Sharp ii 112 

aestivus Sharp ii 111 

arachnipes Sharp ii 112 

bilineatus Sharp ii 108 

bishopi Sharp ii 108, Ixviii 

blackburni Sharp ii 1 1 1 

coUaris sp. n. ii 107 



Plagithmysus concolor Sharp ii 106 

cristatus Sharp ii 113, iii 650 
cuneatus Sharp ii 106 
darwinianus Sharp ii 109, Ixvii 
diana sp. n. ii 107 
elegans sp. n. iii 648 
finschi Har. ii 106 
fractus sp. n. iii 646 
funebris Sharp ii 106 
giffardi Perk, iii 646 
immundus sp. n. iii 646 
laniarckianus sp. n. ii 1 10, Ixvii 
lanaiensis Sharp ii 108 
munroi sp. n. ii 112 
newelli Sharp ii 105 
perkinsi Sharp ii 109 
permundus sp. n. ii 105 
pulverulentus Motsch. ii 107 
pulvillatus Karsch ii 1 1 1 
simplicicollis sp. n. iii 648 
solitarius Sharp ii 106, iii 647 
speculifer Sharp ii in 
sulphurescens Sharp ii iii 
varians Sharp ii 109 
varians Sharp var. ii 1 10 
vicinus Sharp ii 108, Ixviii 
vitticollis Sharp ii 105 
vitticollis var. longulus ii 105 

Planchonia pustulans Cock, iii 104 

Plants, introduced xxv 

Platydenia obscurum Sharp ii 252 

Platygasteridae ii 626, ciii 

Platygaster lecanii Fitch i 328 

Platyhelminthes ii 429 

Platymischoides gen. n. i 296 

molokaiensis sp. n. i 296 

Platynus Bon. iii 228 

ambiens sp. n. iii 228 
calathiformis sp. n. iii 229 
planus Karsch iii 216 

Platypini ii 182 

Platyptiha Hb. 1 472, 731 

brachymorpha Meyr. i 474 
cosmodaetyla Btl. i 472 
fuscicornig Z. i 472, 731 
insularis sp. n. i 475 
littoralis Btl. i 476 
repletalis Btl. i 475 
rhyncophora Meyr. i 475 

Platyura fuscocostata sp. n. iii 2 
hawaiiensis sp. n. iii 3 
insularis sp. n. iii 4 

Plegadis guarauna L. i 456 

Pleuroneurophion hawaiiensis sp. n. i 342 

Plexippus ardelio Thor. ii 511 
ligo Koch ii 512 
paykuUi Aud. ii 512 

Plochionus bonfilsii Dej. iii 28S 
pallens Fabr. iii 288 

Plodia interpunctella Hiib. i 195 

Ploiiariinae iii 151 

Ploiariodes White iii 151 

pulchra Blackb. iii 152 
rubromaculata Blackb. iii 151 
whitei White iii 151 

Ploiariola Reut. iii 151 

Plusiadae i 153, iii 347, cxlviii 

Plusia Ochs. i 159, iii 348 
biloba Steph. i 159 
chalcites Esp. iii 349 
chalcites (verticillata) i 159 



?8 



FA UNA HA WA I I EN SIS 



Plusia pterylota sp. n. iii 348 
Plutella albovenosa sp. n. i 653 

annulatellus Wd. i 652 

brassicella Fitch i 652 

cicerella Rndi. i 652 

cruciferarum Z. i 652 

dubiosella Bent, i 652 

galealella Mab. i 652 

limbipennella Clins. i 652 

maculipennis Crt. i 652 

niollipedella Clms. i 652 

xylostella Hb. i 652 
Poekillopteridae iii 116, cciv 
Poeciloptera acuta Walk, iii 117 
Pogonoiiiyrme.x occidentalis i lig 
Polistes aurifer i 29 

hebraeus i 30 
Polyainma Kri. i 336 
Polynema Hal. i 332, ii 661 

apicalis sp. n. ii 665 

ciliata sp. n. ii 666 

gigas sp. n. ii 661 

havvaiiensis sp. n. i 332 

jassidarum sp. n. ii 666 

nana sp. n. ii 667 

oahuensis sp. n. ii 664 

perforator sp. n. ii 665 

pyrophila sp. n. ii 662 

reduvioli Perk, ii 667 

rubri\entris sp. n. ii 663 

scrutator sp. n. ii 663 

tantalea sp. n. ii -664 

terrestris sp. n. ii 662 

triscia sp. n. ii 664 
Poly.xenus hawaiiensis sp. n. iii 327 
Polyzosteria soror Brunn. ii 6 
Ponera gleadovvii decipiens i ii8 

kalakauae sp. n. i 116 

perkinsi sp. n. i 117 
Pontoscolex hawaiiensis Bedd. ii 4-14 
Porcellio laevis Latr. ii 524 

scaber Latr. ii 524 
Prays Hb. i 651, 735 

fulvocanellus sp. n. i 652, 735 
Prenolepis bourbonica hawaiiensis i 120 

longicornis i 120 

obscura i 120 

sharpii i 121, xlii 
Prionini ii 95 

Prionopteryx exonoma sp. n. i 198 
Priperia gen. n. iii 340 

bicolor sp. n. iii 340 
Pristomerus hawaiiensis sp. n. ii 680 

hilaris sp. n. ii 681 
Proctotrupoidea ii 616, ciii 
Proctotrypes hawaiiensis sp. n. i 294 
Prodisenochus gen. n. iii 210 

terebratus Blackb. iii 211 
Proernus nom. n. ii 497 

aculeatus sp. n. ii 498 

castaneus sp. n. ii 498 

schauinslandi Sim. ii 497, iii 343 

velox sp. n. ii 499, iii 343 
Prognathogryllides ii 22 
Prognathogryllus Brunn. ii 24, ii 689, xlix 

alatus Brunn. ii 24 

elongatus sp. n. ii 25 

forficularis Brunn. ii 29 

inexspectatus sp. n. ii 26 

robustus sp. n. ii 25 

oahuensis sp. n. ii 25 



Prognostola gen. n. i 167 

creninopis sp. n. i 167 
Promecoderus fossulatus Karsch iii 211 
Promylaea gen. n. i 227 

pyropa sp. n. i 227 
Propalticus Sharp iii 420 

oculatus .Sharp iii 421, cxxxiii 
Prosopidae ii 601, Ixxviii 
Prosopis anthracina Sm. i 100 

flavifrons i 10 1 

hilaris Sm. i 103 

kona Bl. i 82 

rugiventris Bl. & C. i 100 

volatilis Sm. i 105 
Prosoplus banksii Fabr. ii 114 
Prosthetochaeta gen. n. iii 24 

fasciata sp. n. iii 25 

lucilioides sp. n. iii 25 

obscura sp. n. iii 25 

robusta sp. n. iii 24 

spp. iii 26 
Protapanteles hawaiiensis i 362 
Protaulacistis gen. n. i 246 

cataphaea sp. n. i 246 
Proteopteryx blackburnii Btl. i 675 

walsinghamii Btl. i 684 
Proterhinidae ii 183, iii 656, cxxv 
Proterhinus adelus sp. n. ii 202, iii 657 

adelus var. chrysadelus var. n. iii 658 

adelus var. constricticeps var. n. iii 658 

affinis sp. n. ii 238 

alyxiae sp. n. ii 244 

amaurodes sp. n. ii 190 

analcis sp. n. ii 230 



angularis Sharp ii 243 



659 



angustiformis sp. n. 11 197 
angustior sp. n. ii 233 
anthracias sp. n. ii 185 
antiquus sp. n. ii 193 
archaeus sp. n. ii 209, iii 666 
arhopalus sp. n. ii 219 
basalis Sharp ii 187 
binotatus sp. n. ii 191 
blackburni Sharp ii 246, iii 666 
brevifonnis sp. n. ii 229 
brevipennis sp. n. ii 218 
calliphyas sp. n. ii 224 
cognatus sp. n. ii 197 
collaris Sharp ii 190 
comes sp. n. ii 213 
compactus sp. n. ii 203 
convexiusculus sp. n. ii 232 
crassicornis sp. n. ii 1S5 
debilis Sharp ii 245 
deceptor sp. n. ii 245, iii 664, l\iv 
deinops sp. n. ii 201, iii 657 
denudatus sp. n. ii 203, iii 661 
desquamatus sp. n. ii 240 
detritus Sharp ii 228 
difficilis sp. n. ii 188 
dispar Sharp ii 243, iii 663 
dubiosus sp. n. ii 187 
echidna sp. n. iii 658 
echinoides sp. n. ii 232 
epichlorus sp. n. ii 230 
epichrysus sp. n. ii 218 
epimelas sp. n. ii 226 
epitrachys sp. n. ii 220 
epitretus sp. n. ii 229 
erythrodes sp. n. ii 234 
eugonias sp. n. ii 186 



INDEX 



39 



Proterhinus eulepis sp. n. ii i88 
eurhopaius sp. n. ii 196 
eurhynchus sp. n. ii 239 
excrucians sp. n. iii 662 
facilis sp. n. iii 663 
ferrugineus sp. n. ii 241 
gigas sp. n. ii 185 
gracilis Sharp ii 238 
haleakalae sp. n. ii 219 
hawaiiensis sp. n. ii 236 
hemichlorus sp. n. ii 217 
homoeochromus sp. n. ii 197 
heterostictus sp. n. iii 661 
heterotarsus sp. n. ii 234 
humeralis Sharp ii 217 
hypotretus sp. n. ii 240 
ineptus Sharp ii 228 
innotabilis sp. n. ii 242 
insignis .Sharp ii 227 
integer Sharp ii 228 
kaalae sp. n. ii 2(39 
kamptarthrus sp. n. ii 199 
lanaiensis sp. n. ii 227 
laticolHs Blackb. ii 211 
laticornis sp. n. ii 196 
lecontei Sharp ii 213 
leiorhynchus sp. n. ii 200, iii 656 
leptophyas sp. n. ii 198 
leptorhynchus sp. n. ii 222 
leptothrix sp. n. ii 207 
leucothorax sp. n. ii 233 
linearis Blackb. ii 198 
longicornis Sharp ii 227 
longulus Sharp ii 208, iii 660 
maculatus sp. n. ii 221 
maculifer sp. n. ii 198 
maurus sp. n. iii 658 
megalotarsus sp. n. ii 216 
microtarsus sp. n. ii 215 
minimus sp. n. iii 666 
mirabilis sp. n. ii 223 
molokaiensis sp. n. ii 231 
myrsineoides sp. n. iii 659 
myrsineus sp. n. iii 659 
navita sp. n. ii 244 
neglectus sp. n. ii i8g 
nigricans Sharp ii 191 
nivicola sp. n. ii 225 
oahuensis sp. n. ii 208 
obscuricolor sp. n. ii 202, iii 660 
obscurus Sharp ii 210, iii 663 
ombrophilus sp. n. ii 235 
oscillans Sharp ii 210, iii 664 
osculans sp. n. ii 222 
oxygonias sp. n. ii 206 
pachycnemis sp. n. ii 211, iii 664 
paradoxus Sharp ii 208 
peles sp. n. ii 237 
persimilis sp. n. ii 224 
pipturi sp. n. iii 665 
platygonias sp. n. ii 204 
platygiiifiioides sp. n. iii 661 
podagricus sp. n. iii 656 
pteridis sp. n. ii 235 
punctipennis Sharp ii 225 
pusillus Sharp ii 212, iii 665 
pusillus var. subpusillus van n. iii 665 
robustus Blackb. ii 205 
rufescens sp. n. ii 237 
ruficollis sp. n. ii 226 
ruficornis sp. n. ii 200, iii 657 



Proterhinus scutatus Blackb. ii 192 
separandus sp. n. ii 221 
serricornis sp. n. ii 195 
seticoUis sp. n. ii 207, iii 661 
setiger sp. n. ii 193 
setulosus sp. n. ii 192 
sharpi sp. n. ii 213 
similis Blackb. ii 241 
simplex Sharp ii 206 
solitarius sp. n. ii 193 
squalidus sp. n. ii 194 
squamicollis sp. n. ii 201, iii 657 
sternalioides sp. n. ii 215 
sternalis Sharp ii 215 
subangularis sp. n. iii 660 
subdeceptor sp. n. iii 664 
subplanatus sp. n. ii 205, iii 660 
tarsalis Blackb. ii 239 
transversalis sp. n. iii 662 
tuberculiceps sp. n. ii 214 
validus Sharp ii 212 
vestitus Sharp ii 205, iii 662 
vicinus sp. n. ii 212, iii 665 
vulcanus sp. n. ii 236 
wikstroemiae sp. n. ii 195 

Proterocosma triplanetis Meyr. i 514 

Protocolletis constricta Meyr. i 225 

Protoparce blackburni Butl. i 193 

Psallus sharpianus sp. n. iii 131 

Psamniodius Serv. iii 402 

Psammoechus Latr. iii 427 

desjardinsii Gudr. iii 428 
insularis Sharp iii 428 
pallidipennis Blackb. iii 428 

Psecadia Hb. i 507 

Pseudobaeus gen. n. ii 620 

peregrinus sp. n. ii 621 

Pseudobroscus gen. n. iii 196 
lentus sp. n. iii 196 

Pseudoclerada gen. n. iii 140 
kilaueae Kirk, ii 553 
niorai sp. n. iii 141, ii 553 

Pseudococcus adonidum Linne iii 103 
albizziae Mask, iii 103 
calceolariae Mask, iii 103 
citri Risso iii 103 
vastator Mask, iii 103 
virgatus Cock, iii 103 

Pseudocoremia paludicola Butl. i 179 

Pseudocorylophidae Matth. iii 431 

Pseudohyalina kauaiensis PfeifF. ii 279 

Pseudolus hospes sp. n. ii 149, iii 656 
longulus Boh. ii 149 

Pseudonestor Rothsch. i 431, Ixxii 
xanthophrys Rothsch. i 432 

Pseudopsectra gen. n. ii 46, xlix 
lobipennis sp. n. ii 46 

Pseudopterocheilus ii 607 

Pseudopterocheilus congruus Sm. ii 607 
pterocheiloides Perk, ii 607 
relictus Perk, ii 607 

Psilopus patellifer Thorns, iii 11 

Psittacirostra Temni. i 433, Ixii 
psittacea Gmel. i 435 

Psocidae ii 77 

Psocoptera clxxv 

Psocus distinguendus sp. n. ii So 
haleakalae sp. n. ii 77 
heterogamias sp. n. ii 82 
hualalai sp. n. ii 79 
kauaiensis sp. n. ii 79 



40 



FAUNA HAWAIIENSIS 



8i 

ii 

82 

81 

78 

81 



490 
i 490 



11 497 



Psocus konae sp. n. ii 79 
lanaiensis sp. n. ii 
molokaiensis sp. n. 
monticola sp. n. ii 
oahuensis sp. n. ii 
simulator sp. n. ii 
sylvestris sp. n. ii 
unicus sp. n. ii 78 
vittipennis sp. n. ii 

Psychra gen. n. i 489 

brevipalpis sp. n. i 
phycidifonnis sp. n. 

Psychodidae iii 6, cl.xxxi 

Psychoda inornata sp. n. iii 

Psyllidae iii 113 

Pterelas schauinslandi Sim. 

Pteritoptides ii 114 

Pterodiscus digonophorus Ancey ii 
petasus Aucey ii 292 
wesleyi Sykes ii 292 

Pteromalidae i 323, ii 655, cvi 

Pteromalus fuscitarsis sp. n. i 323 

Pterophorina i 471, 731, clxii 

Pterostichides iii 243, Ivi 

Ptiliodes blackburni Matth. iii 536 
insignis sp. n. iii 536 
pulchellus sp. n. iii 537 

Ptinella pacifica Matth. iii 537 

Ptychothrix gen. n. i 489 

vagans sp. n. i 489 

Puffinus cuneattis Salv. i 463 
newelli Hensh. i 463 

Pulvinaria mammeae Mask, ii 
psidii Mask, iii 104 

Pundaluoya simplicia Dist. ii 

Pupidae ii 293 

Pupa acanthinula Ancey ii 294 
admodesta Migh. ii 294 
bacca Pease ii 294 
costata Pease ii 294 
costulosa Pease ii 295 
iyonsiana Ancey ii 294 
lyrata Gould ii 294 
magdalenae Ancey ii 294 
mirabilis Ancey ii 295 
nacca Gould ii 295 
newcombi Pfeiffer ii 295 
pediculus Shuttl. ii 295 
peponum Gould ii 380 
peponum Gould ii 382 
perlonga Pease ii 295 
seminulum Boettg. ii 295 
striatula Pease ii 294 

Pycnophion fuscipennis sp. n. ii 680 
kauaiensis sp. n. i 344 
molokaiensis sp. n. i 344 

Pyralididae i 275, iii 365 

Pyralidina i 731 



104 



577 



Pyralis Linn. 
Boisd. 



365 



365 



achatina Butl. 1 275 
grotiana F. i 709 
manihotalis Guen. i 275 
mauritialis Boisd. i 275 
Pyrameis atalanta etc. xxix 

Cordelia Doubl. i 193 
Pyraustidae iii 356, clvii 
Pyrausta Schranck i 220, iii 360 
brontias sp. n. i 223 
bryochloris sp. n. i 221 
chloropis sp. n. i 222 



224 
223 



549 

547 

548 



Pyrausta constricta Butl. i 224, iii 360 

dracontias sp. n. i 221 

dryadopa sp. n. i 222 

litorea Butl. i 224, xxv 

phaethontia sp. n. i 225 

psychropa sp. n. i 

thermantis sp. n. i 
Pyroderces HS. i 514 
Pyrrhocoridae iii 158 

Querquedula carolinensis Linn, i 461 

Rainfall xxiii 

Reclada moesta White iii 160 

Recurvaria sarcitea Hw. i 649 

Reduviidae iii 149, ii 550, cxcviii 

Reduviinae iii 150 

Reduviolus Kirby iii 153, ii 546, xlix, Iwiii cxc\' 

arrogans Kirk, ii 547 

blackburni White iii 155, ii 546 

capsiformis ii 546, cxcv 

curtipennis Blackb. ii 549 

innotatus iii 154, ii 546 

kahavalu Kirk, ii 546 

kaonohiula Kirk, ii 548 

kerasphoros Kirk, ii 

koelensis Blackb. ii 

lolupe Kirk, ii 549 

lusciosus iii 157, ii 

milu ii 549 

monticola Kir-k. ii 548 

montivagus Kirk, ii 548 

morai sp. n. iii 155, ii 547 

nubigenus Kirk, ii 547 

oscillans Blackb. ii 547 

paludicola Kirk, ii 549 

procellaris Kirk, ii 548 

rubritinctus iii 157, ii 549 

sharpianus sp. n. iii 156 

silvestris Kirk, ii 549 

silvicola Kirk, ii 548 

subrufus iii 156, ii 547 

tarai sp. n. iii 154 

truculentus Kirk, ii 547 

volcanicola Kirk, ii 548 
Reduvius laniger Butl. iii 150 
Reptilia i 365 

Rhantus pacificus Sharp iii 289 
Rhinia testacea Desv. iii 83 
Rhinomactrum gen. n. i 531 

rutilellum sp. n. i 531 

scapulellum sp. n. i 531 
Rhizophagus capito Fairm. iii 434 
Rhodacanthis Rothsch. i 436, Ixxii 

flaviceps Rothsch. i 438 

palmeri Rothsch. i 438 
Rhodaria despecta Butl. i 217 
Rhopalus Schill. iii 170 

hyalinus Fabr. iii 170, cxciii 
Rhygchium nigripennis Holm, i 70 
Rhyncogonus blackburni Sharp ii 126 

depressus sp. n. ii 128 «^ 

dubius sp. n. ii 125 

extraneus sp. n. iii 651 

freycinetiae sp. n. ii 126 

funereus sp. n. ii 123 

fuscus sp. n. iii 652 

kauaiensis sp. n. ii 127 

koebelei sp. n. ii 126, iii 653 

lahainae sp. n. ii 128 

lanaiensis sp. n. ii 128 



INDEX 



41 



Rhyncogonus minor sp. n. ii 124 

niolokaiensis sp. n. ii 125 

nitidus sp. n. ii 122 

oleae sp. n. iii 652 

sharpi sp. n. iii 650 

simplex sp. n. iii 651 

sordidus sp. n. ii 127 

squamiger sp. n. ii 123 

stygius sp. n. ii 124 

sylvicola sp. n. ii 130 

tuberculatus sp. n. ii 129 

vestitus Sharp ii 130, \xv - 

vittatus sp. n. ii 129 
Rhyncolus gracilis Boh. ii 149 

longulus Boh. ii 149 

tenuis (}enini. ii 149 
Rhynchopimpla Kriechb. i 336 
Rhyparochromus nigriceps Dall. iii 159 
Rhysodidae iii 429 
Rhyzodes liratiis Chevr. iii 429 
Rhyzopertha dominica Fabr. iii 643 

pusilla .Steph. iii 643 
Riphidophora Hb. i 477 

.Sacium angusticolle sp. n. iii 416 
.Salbia continuatalis Wall, i 205 
.Sakla exulans White iii 146 

oahnensis Blackb. iii 146 
Salticus ■acutus Blackvv. ii 511 

citus Canibr. ii 511 

ruficapillus Dol. ii 511 

scabellatus Butl. ii 511 

striatus Luc. ii 511 

vaillanti Luc. ii 512 
.Sandalodes Keyserl. ii 512 

albociliatus sp. n. ii 514 

canosus sp. n. ii 515 

cruciatus sp. n. ii 516 

navatus sp. n. ii 515 

pubens sp. n. ii 513 

seniculus sp. n. ii 517 

validus sp. n. ii 514, iii 344 

verecundus sp. n. ii 516, iii 344 
Saprinus lugens iii 510 

oregonensis Lee. iii 510 
Sapromyza sp. iii 85, cl.\.\xviii 
Saprosites pygmaeus Har. iii 402 
Sarcophagidae iii 21, 83 
.Sarcophaga barbata Thorns, iii 26 

dux Thoms. iii 27 

pallinervis Thoms. iii 26 
Sargus sp. iii 1 1 
Sarona gen. n. iii 142 

adonias sp. n. iii 142 
Saronychium inconspicuum Blackb. iii 288 
Scaphisonia perkinsi sp. n. iii 534 
Scarabeidae iii 401, cxxxi 
Scardia lignivora Btl. i 646, 647 
.Scatella hawaiiensis sp. n. iii 49, 85 
Scelionidae ii 618, cii 
Scenopinus niger Mg. iii 11, clxxxii 
Schistoceros cornutus Pall, iii 642 
Sciara niolokaiensis sp. n. iii 2 
Scieropepla typhicola Meyr. i 485 
Sciomyzidae clxxxvii 
.Sciomyza hawaiiensis sp. n. iii 84 

sp. iii 43 
Sciophagus pandanicola Fairm. ii 253 
Scleroderma breviventre sp. n. i 286 

chlorodes sp. n. ii 613 

euprepes sp. n. ii 612 

F. H. 



Scleroderma kaalae sp. n. i 2S5 
lanaiense sp. n. i 2S5 
nigriventre sp. n. i 285 
perkinsi sp. n. i 284 
poecilodes sp. n. ii 613 
polynesiale Saund. i 284 
sophorae sp. n. ii 614 
Sclerostomum armatum Dies, ii 428 
Scolothrips 6-maculatus Perg. iii 700 
Scolytidae ii 173, cxxiii 
Scoparia Haw. i 246, iii 362 
actias sp. n. i 256 
aeolias sp. n. i 262 
altivolans Butl. i 155 
amphicypella sp. n. i 250 
antimacha sp. n. i 252 
balanopis sp. n. i 250, iii 362 
bucolica sp. n. i 263, iii 364 
catactis sp. n. iii 363 
clonodes sp. n. i 265 
coarctata i 271 
crataea sp. n. i 259 
cryerodes sp. n. i 259 
dactyliopa sp. n. i 268 
demodes Meyr. i 268, iii 364 
empeda sp. n. i 267 
epimystis sp. n. i 269 
erebochalea sp. n. i 264 
formosa Butl. i 257 
frigida Butl. i 270, 272 
geraea sp. n. i 269, iii 365 
gonodecta sp. n. iii 362 
halirrhoa sp. n. i 255 
halmaea sp. n. i 272 
halmaea Meyr. iii 365 
hawaiensis Butl. i 267, iii 364 
ianthes sp. n. i 261, iii 364 
ischnias Meyr. i 271 
jucunda Butl. i 257 
loxocentra sp. n. i 255 
niacrophanes Meyr. i 271 
marmarias sp. n. i 261, iii 364 
nielanocephala sp. n. i 256 
melanopis Meyr. i 271, iii 365 
melichlora sp. n. i 266 
meristis sp. n. i 272, iii 365 
mesoleuca sp. n. i 252, iii 363 
niiantis sp. n. i 252 
montana Butl. i 271 
nectarias sp. n. i 262 
nyctombra sp. n. i 264 
oenopis sp. n. i 264 
ombrodes Meyr. i 260, iii 364 
omichlopis sp. n. i 274 
orthoria sp. n. i 251 
oxythyma sp. n. i 250 
pachysema Meyr. i 254 
parachlora sp. n. i 261 
passalota sp. n. i 251 
pentaspila sp. n. i 253 
peronetis sp. n. i 270 
platyscia sp. n. i 273 
probolaea sp. n. i 273 
pyrseutis sp. n. i 263, iii 364 
religiosa sp. n. iii 365 
rhombias sp. n. i 249, iii 362 
siderina sp. n. i 260 
struthias sp. n. i 257 
tetranesa sp. n. i 249 
thalamias sp. n. i 254 
thyellopis sp. n. i 265 



42 



FAUNA HAWAIIENSIS 



Scoparia triacma sp. n. i 258 

tyraiila sp. n. i 253 

venosa Butl. i 274 

zophochlora sp. n. i 266 
Scoptonoma Z. i 469 
Scopula argoscelis Meyr. i 217 

constricta Butl. i 224 

despecta Meyr. i 217 

ennychioides Meyr. i 216 

eucrena Meyr. i 213 

exigua i 217 

litorea Butl. i 224 

micacea Meyr. i 212 

monticolans Meyr. i 211 

nigrescens Meyr. i 211 
Scorpiones ii 517 
Scotomera hydrophila Butl. i 199 
Scotorythra Butl. i 170, .\xvi 

anagraptis sp. n. i 178 

arboricolans Butl. i 173, iii 350 

artemidora sp. n. i 186 

aruraea sp. n. i 176 

brachytarsa sp. n. i 187, iii 354 

capnopa sp. n. i 183 

caryopis sp. n. i 173, iii 350 

corticea Butl. i 174 

demetrias sp. n. i 180 

diceraunia Meyr. iii 354 

dissotis sp. n. iii 351 

epicyma sp. n. i 174 

euryphaea sp. n. i 188, iii 354 

goniastis sp. n. i 180 

hecataea sp. n. i 177, iii 351 

homotrias sp. n. i 176, iii 351 

hyparcha sp. n. i 189 

idolias sp. n. i 178 

isospora sp. n. i 175 

leptias sp. n. iii 354 

macrosoma sp. n. i 185 

megalophylla sp. n. i 189, iii 355 

metacrossa sp. n. iii 352 

nephelosticta sp. n. i 183, iii 353 

ortharcha sp. n. i 181 

oxyphractis sp. n. i 181, iii 352 

pachyspila sp. n. i 185, iii 353 

paludicola Butl. i 179 

paratactis sp. n. iii 353 

platycapna sp. n. i 183 

rara Butl. i 187, iii 354 

syngonopa sp. 11. i 172 

trachyopis sp. n. i 184 

trapezias sp. n. i 177 

triscia sp. n. i 182, iii 353 
Scotosia rara Butl. i 187 

corticea Butl. i 174 
Scymnus discedens Sharp iii 414 

ocellatus Sharp iii 414 

vi\idus .Sharp iii 414 
Scytodes marmorata Koch ii 444 
Selidosemidae iii 349, cl. 
Seniiotoscelis Reut. iii 126 
Semisulphurea Stn. i 712 
Semnoprepia gen. n. i 644 

fulvogrisea sp. n. i 644 

margella sp. n. i 645 

petroptilota sp. n. i 645 
Sephora gen. n. iii 161 

calvus White iii 162, ii 536 

criniger White iii 161, ii 535 
Sericoderus basalis Sharp iii 416 

pubipennis Sharp iii 417 



Setomorpha dryas sp. n. i 726 

rutella Z. i 726 

Sexual characters Ixv 
Sierola Cam. i 286, ii 614 

collaris sp. n. i 292 

dichroma sp. n. ii 614 

flavocoUaris sp. n. i 291 

kauaiensis sp. n. i 292 

leuconeura Cam. i 289 

molokaiensis sp. n. i 290 

monticola Cam. i 289 

oahuensis sp. n. i 290 

testaceipes Cam. i 291 
Sierolomorpha hospes sp. n. ii 616 
Silpha hirta Marsh, iii 421 
Silvanus surinamensis Linn, iii 428 

unidentatiis Fabr. iii 428 
Simodactylus cinnamomeus Boisd. iii 369 
Simplicia caeneusalis i 153 

robustalis Guen. i 153, iii 347 
Sinella Brooke iii 300 
Sinoxylon conigerum Gerst. iii 643 
Siphanta acuta Walk, iii 117 
Sisyrophyta gen. n. i 168 

gomphias sp. n. i 169, iii 349 
% ochetias sp. n. i 169 
Smeringopus elongatus Vins. ii 446 
Snow xxiii 

Solanella Bd\ . i 4S3 
Solenopsis geminata rufa i 119 
Solenius i 15 

Solindenia picticornis Cam. i 314 
Sotenus setiger Sharp, ii 97 
Spaelotis crinigera Butl. i 148 
Spalangia cameroni sp. n. ii 656 

hirta Hal. i 325 

lanaiensis sp. n. i 325 

simplex sp. n. ii 657 
Spathius perdebilis sp. n. ii 685 
Spatula clypeata L. i 461 
Species-formation Ixi 
Sphaeridium abdominale Fabr. iii 579 
Sphaerococcus bambusae Mask, iii 104 
Sphegidae Ixxxix 

Sphenophorus obscurus Boisd. ii 139 
Sphingolabis hawaiiensis Borm. ii 5 
Sphingidae cliii 
Spliinx conxoh'uli Linn, (cingulata Fab.) i 193 

celeus Hub. i 193 
Spiraxis cumingiana Pfeiff. ii 373 

obsoleta FfeilT. ii 366 

paradoxa Pfeiff. ii 374 

sandwicensis Pfeiff. ii 399 
Spodoptera exigua Hiibn. i 153 

flavimaculata i 153 

mauritia Boisd. i 152, iii 347 
Stagmatophora honorariella sp. n. i 515 

incertulella Wkr. {Proterocosma) i 515 

quadrifasciata sp. n. i 516 

sordidella sp. n. i 516 

tridigitella sp. n. {Protcrocosma) i 515 
Stagmopinipla Sauss. i 336 
Staphylinidae iii 538, cxxxvii 
Staphylinus discoideus Giav. iii 548 

maxlllosus L. iii 548 

ni^^ritulus Gra\-. iii 548 
Stasilea curvicomis Karsch. ii 114 
Steganoptycha foetori\-orans Btl. i 673 
Stcnagria currax Sharp iii 578 
Stenamma longiceps Sm. {hchiuvnyriiicx) i 118 
.Stenocorus simplex Gyll. ii 97 



INDEX 



43 



Stenogyridae ii 383 

Stenopsociis pulchripennis sp. 11. ii S3 
Stenopterus pulverulentus Mot5ch. ii 107 
Sterna fuliginosa Gniel. i 464 
liinata Peale i 464 
nielanauchen Temm. i 464 
Stictomischus haleakalae sp. n. i 311 
Stigmus inordinatiis Fox ii 605 
Stoeberhinus Htl. i 485 

testaceiis Btl. i 485, 733 
Stomorhina Rond. iii 28 

pleuralis Thorns, iii 28 
Stomoxys Geoffr. iii 28 

calcitrans Linn, iii 28 
Stratiomyidae iii 11, 79, clxxxii 
Strepsilas inteipres Linn, i 449 
Strepsiptera iii 667 
Stiomatium hirtuni Fairm. ii 96 
Strongyloides intestinalis Bav. ii 428 
Stylopyga decoiata Brunn. ii 6 
.Styringomyia didyma sp. n. iii 10 
Succinea Drap. ii 384 

aperta Lea ii 390 
apicalis Ancey ii 390 
appro.Nimata ShutU. ii 390 
aurulenta Ancey ii 385 
baldwini .Ancey ii 385 
bicoloiata Ancey ii 385 
caduca Migh. ii 385 
canella Gould ii 385 
casta Ancey ii 386 
cepulla (ioiild ii 386 
cinnamomea Ancey ii 386 



delicata Ancey ii 



386 



elongata Pease ii 386 
explanata Gould ii 387 
fragilis Soul, ii 386 
garrettiana Ancey ii 387 
inconspicua Ancey ii 387 
konaensis Sykes ii 387 
latulenta Ancey ii 387 
lumbalis Gould ii 387 
niauiensis Ancey ii 388 
newconibi Pfeiff. ii 389 
newcombiana Garr. ii 388 
patula Migh. ii 389 
protracta sp. n. ii 388 
pudorina Gould ii 390 
punctata Pfeiff. ii 388 
rotundata Gould ii 389 
rubella Pease ii 389 
rubida Pease ii 389 
socilcyeti Ancey ii 386 
tenerrima Ancey ii 390 
thaanurni Ancey ii 389 
venusta Gould ii 389 
vesicalis (joukl ii 390 
waianaensis Ancey ii 390 

Sula tiber L. i 462 

piscator L. i 462 

Sulamita gen. n. iii 129 

dryas Kirk, ii 552 

lunalilo sp. n. iii 130, ii 552 

opuna sp. n. iii 131 

oreias Kirk, ii 552 

Sulamitaria Div. n. iii 129 

Sympetrum blackburni McLachL ii 62 

Sympiesis konae sp. n. i 331 

Synaerna Sim. ii 492 

dimidiatipes sp. n. ii 493 
fronto sp. n. ii 493, iii 342 



Synaenia impotens sp. n. ii 494 
naevigerum sp. n. ii 4^4 
rutntliorax sp. n. iii 342 

Synoniotis i 492 

epicapna Meyr. i 500 

Syritta oceanica Macq. iii 19 

Syroloma jjen. n. ii 509 

major sp. n. ii 509, iii 344 
minor sp. n. ii 510 

Syrphidae iii 19, iii 82, clxxxiv 
.Syrphus grandicornis Macc|. iii 19 
obesus Fabr. iii 19 

Tachinidae iii 20, 83, clxxiv 

Tachyusa pumila Sharp iii 577 

Tachys arcanicola Blackb. iii 287 
atomus Blackb. iii 287 
mucescens Blackb. iii 288 
oahuensis Blackb. iii 287 

Taenia crassicoUis Rud. ii 434 
solium Lutz. ii 434 

Talis floricolans Bull, i 200, iii 356 
geralea sp. n. i ig8 
homoroda sp. n. i 199, iii 356 
hyacinthena sp. n. i 200 
hydrophila Butl. i 199, iii 356 
oxyptera Meyr. i 199 

Talitridae ii 527 

Tanytarsus lacieiclavus sp. n. iii 5 

Tapinoma melanoceplialum i 120, cii 

Tarsostenus univittatus Rossi iii 367 

Tatare familiaris xvii 

Tebennophorus australis Bergh ii 286 
bilineatus Bens, ii 286 
striatus Hass. ii 286 

Tegenaria civilis Walck. ii 505 
dehrani Thor. ii 505 
domestica Clerck ii 505 

Tegeocranus pustulatus sp. n. iii 704 

Telenomus adelphus sp. n. ii 6ig 
despiciendus sp. n. ii 618 
rhopali sp. n. ii 61 8, cxciii 
paractias sp. n. ii 619 
vulcanus sp. n. ii 619 

Telephanus insularis Sharp iii 428 

pallidipennis Blackb. iii 428 

Telmatophilus debilis Sharp iii 422 

Temperature xxiii 

Tenebrio diaperinus Panz. ii 253 
mauritanicus Linn, iii 434 

Tenebrionidae ii 252, cxxvii 

Tenebroides Pill. & Mitterp. iii 434 

Tephritis annonae Fabr. iii 44 
crassipes iii 45 
cratericola sp. n. iii 46 
limpidapex sp. n. iii 46 

Teras basialbana Wkr. i 690 
dotalana Wkr. i 690 
illepida Btl. i 681 
postvittana Wkr. i 690 
retractana Wkr. i 690 
secretana Wkr. i 690 
scitulana Wkr. i 690 

Teratodes Gn. i 703 

Termitidae ii 88 

Tetigoniidae or Jassidae iii 114, ccix 

Tetigonia Geotlr. iii 116 

varicolor Sign, iii 116 

Tetragnatha Latr. ii 466 

cuneiventris sp. n. ii 475 
hawaiensis sp. n. ii 470, iii 341 



F 2 



44 



FAUNA HAWAIIENSIS 



Tetragnatha kauaiensis ii 472 

leptognatha Thor. ii 468 
mandibulata Walck. ii 468 
minatoria Sim. ii 468 
netrix sp. n. ii 468 
perkinsi sp. n. ii 470 
restricta sp. n. ii 473 
sobrina sp. n. ii 471 
Lincifera sp. n. ii 474 

Tetramoriuiii guineense Fabr. i iiS 

Tetrasticlius hagenowii Ratz. i 329 

Tettigoniidae ii 555 

Teutana grossa Koch ii 460 

Thaanumia omphalodes Ancey ii 373 

Thallatodora gen. n. ii 146 

insignis sp. n. ii 146 

Thauniatodon Pilsbry ii 288 

Thauniatogiylkis gen. n. ii 27 
vai'iegatus sp. n. ii 27 

Theridion Walck. ii 448 

acutitarse sp. n. ii 456, iii 340 
bajiilans Koch ii 450 
borbonicum Vins. ii 450 
canipestratum sp. n. ii 450 
flavoaurantiacum Sim. ii 450 
grallator sp. n. ii 456 
grossum Koch ii 460 
haleakalense sp. n. ii 453 
kauaiense sp. n. ii 455 
longipes Hass. ii 450 
luteipes Cambr. ii 450 
mauiense sp. n. ii 452 
mehnum sp. n. ii 457 
perkinsi sp. n. ii 454, iii 340 
posticatum sp. n. ii 458 
praetexturn sp. n. ii 450 
praetextum concolor ii 451 
rufifes Luc. ii 450 
tepidariorum Koch ii 449 
trigonium Hentz ii 446 
vulgare Hentz ii 449 

Thoracophorus blackburni Sharp iii 538 
brevipennis Sharp iii 539 

Thrips 6-maculata Perg. iii 700 
multispinus sp. n. iii 699 
pallida Beach iii 700 

Thriscothorax gen. n. iii 257 
apicalis sp. n. iii 264 
argutor sp. n. iii 268 
bembidioides Blackb. iii 262 
bradyderus sp. n. iii 267 
, bre\is Blackb. iii 267 
chalcosiis sp. n. iii 264 
constrictus sp. n. iii 261 
cordaticollis Blackb. iii 259 
discedens sp. n. iii 263 
ducalis sp. n. iii 266 
filipes sp. n. iii 257 
gracilis sp. 11. iii 258 
insolitus sp. n. iii 261 
karschi Blackb. iii 266 
laetiis Blackb. iii 262 
laticoUis sp. n. iii 267 
modestiis sp. n. iii 259 
molokaiae sp. n. iii 260 
mundanus sp. n. iii 261 
obscuricolor Blackb. iii 266 
palustris sp. n. iii 258 
paradoxus Blackb. iii 263 
perkinsi sp. n. iii 265 
perstriatus sp. n. iii 260 



Thriscothorax platysminus sp. n. iii 268 
robustus Blackb. iii 268 
subconstrictus sp. n. iii 259 
unctus Blackb. iii 257 
variipes sp. n. iii 265 

Thyene Sim. ii 512 

Thyreocoridae cxcii 

Thyrocopa .Meyr. i 492, 733, xxvi 

abusa sp. n. i 492, 504, 733 
adumbrata sp. n. i 503 
albonubila sp. n. i 495 
alterna sp. n. i 501 
argentea Btl. i 496, 733 
cinercUa sp. n. i 494 
depressariella sp. n. i 498 
epicapna Meyr. i 500 
fraudulenlella sp. n. i 502 
geminipuncta sp. n. i 506 
gigas Btl. i 493 
immutata sp. n. i 501 
indecora Btl. i 497 
inerrnis sp. n. i 503 
lactea Btl. i 497 
leonina sp. n. i 505 
mediomaculata sp. n. i 506 
megas sp. n. i 495 
minor sp. n. i 496 
nubifer sp. n. i 499 
pallida sp. n. i 502, 733 
pulverulenta sp. n. i 500 
usitata Btl. i 504, TlZ 
usitata Meyr. i 492, 504 
seminatella sp. n. i 499 
sp. Btl. i 499 
subahenea sp. n. i 505 
tessellatella sp. n. i 498 
viduella sp. n. i 494 

Thysanoptera iii 669, ccxi 

Thysanura iii 293, ccxx 

Tichorhinus Fieb. ii 552 

kassandra Kirk, ii 553 

Tinea L. i 729 

aurifluella Hb. i 507 
crocicapitella Clnis. i 73S 
decunella Hb. i 478 
ferruginella Wlsm. i 72S 
fuscipunctella Hw. i 729 
hyalinella Stgr. i 728 
lactella SchilT. i 648, 649 
pellionella L. i 729 
pyrausta Hb. i 507 
rusticella Hb. i 727 
simulans Btl. i 714 
sp. ? i 729 
syringella F. i 721 

Tineidae i 711, 737. clxix 

Tineina i 478, 731 

Tomocera How. i 324, ii 655 
californica How. i 324 
ceroplastis Perk, ii 655 

Tornatellina Beck ii 380 

bacillaris Mouss. ii 382 
baldwini Ancey ii 380 
castanea Ffeiff. ii 376 
compacta sp. n. ii 380 
confusa sp. n. ii 380 
cylindrica sp. n. ii 381 
dentata Pease ii 381 
euryomphala Ancey ii 381 
extincta Ancey ii 381 
gracilis Pease ii 381 



INDEX 



45 



ToniatcUina newconibi Pfeiff. ii 382 

oblongata Pease ii 3S2 

peponum Gould ii 380, 382 

perkinsi sp. n. ii 382 

petitiana Pfeiff. ii 377 

striata Newc. ii 370 

trochoides sp. n. ii 383 

umbilicata Ancey ii 383 
Tortricidae i 673, 735, clxvii 
Toitrix L. i 698 

capucina sp. n. i 701, 736 

chlorocalla sp. n. i 699 

faleriana sp. n. i 701 

lanceolana Hb. i 687 

metallurgica sp. n. i 699 

piceana L. i 6go 

rubiginis sp. 11. i 702 

thoracina sp. n. i 700 

viridana L. i 6g8 

woeberiana Schiff i 683 
Toxcunia Walk, i 307 

affinis sp. n. i 309 

fenugineipes sp. n. i 309 

hawaiiensis sp. n. i 308 

nigrocyinea sp. n. i 311 

nubilipennis sp. n. i 310 

tarsata sp. n. i 309 
Toxocampa noctivolans Butl. i i 58 
Tramea lacerata Hagen ii 62 
Treniatoda ii 429 

Triatoma riibrofasciata de Geer ii 550 
Tribolium ferrugineum Fabr. ii 253 
Trichoceplialus dispar Rud. ii 428 
Trichogrammidae ii 658, cviii 
Trichopterygidae iii 535, cxxxvi 
Trichopteryx Kirby & Spence iii 538 
Trichoptilus W'lsm. i 471 

centetes Meyr. i 471 

hawaiiensis Btl. i 471 

oxydactylus Wkr. i 471 

pygmaeus Wlsm. i 471 
Tricliothrips laticornis sp. n. iii 692 

nigricans sp. n. iii 693 
Trigonidium paciticum Scudd. ii 22 
Trimicra lateralis sp. n. iii 9 
Tringa acuminata Horsf. i 451 

maculata Vieill. i 451 
Trioza iolani sp. n. iii 114 
Triphleps Fieb. iii 125 

persequens White ii 551, iii 125 
Tritocleis gen. n. i 190 

microphylla sp. n. i 190 
Trogophloeus abdoniinalis Sharp iii 544 

fontinalis Sharp iii 544 

senilis Sharp iii 543 
Trogosita cornuta Fabr. ii 252 
Trox scaber Linn, iii 401 
Trypetidae iii 44, clxxxvii 
Trypeta crassipes Thorns, iii 45 
Trybliographa hawaiiensis sp. n. i 300 
Trypoxylon bicolor Sm. ii 606 
Trypoxylonidae ii 606, Ixxxviii 
Turbo ape.xfulva Dixon ii 29S 

lugubris Chenin. ii 298 

lugubris sinistrorsus Chenin. ii 301 
Typhaea fumata Linn, iii 419 



Ulesanis oahuensis sp. n. ii 460 

Urocryptus Westw. i 315 

L^rophora ciuadrivittata Macq. iii 44 



\'anessa atalanta Linn, i 193 

cardui Linn, i 194 

huntera Fab. i 194 ^ 

tammeamea Esch. i 193, civ 
Variability Ixviii, Ixix 
Veliomorpha de Carl, iii 157 
Vermes ccxxviii 
Vertigo nacca Gould ii 295 
Vespidae xcix 
Vestiaria Flem. i 402, xxi 

coccinea Forst. i 405 
Viridonia Rothsch. i 412, Ixxii 

sagittirostris Rothsch. i 413 
Vitrea Fitz. ii 278 

lanaiensis Sykes ii 278 

molokaiensis Sykes ii 278 

pauxillus Gould ii 279 
Vitrina caperata Gould ii 277 

tenella Gould ii 27S 
Vo ucella obesa Fabr. iii 19 

Westwoodella hilaris sp. n. ii 658 

Xanthocorynus gen. n. iii 549 

deceptor iii 550 
Xanthogramma grandicornis Macq. iii 19 
Xanthorhoe Hiib. i 165 

caustoscia sp. n. i 167 

insularis Butl. i 166 

ioxantha sp. n. i 165 

leucoxyla sp. n. i 166 
Xenusa pumila Sharp iii 577 
Xeroscopa demodes Meyr. i 268 

formosa Meyr. i 257 

hawaiensis Meyr. i 267 
■ jucunda Meyr. i 258 

melanopis Meyr. i 271 

pachysema Meyr. i 254 

venosa Meyr. i 274 
Xiphidium fuscum Fab. ii 14 
Xyleborus agamus Blackb. ii 178 

confusus Eichhoff ii 177 

dubiosus sp n. ii 177 

exsectus sp. n. ii 179 

frigidus Blackb. ii 178 

hawaiiensis sp. n. ii 175 

ignobilis sp. n. ii 180 

inimaturus Blackb. ii 178 

insularis Sharp ii I77 

kauaiensis sp. n. ii 174 

lanaiensis sp. n. ii 176 

littoralis sp. n. ii I79 

mauiensis sp. n. ii ,175 

oahuensis sp. n. ii I77 

obliquus Sharp ii 176 

rugatus Blackb. ii 178 

simillimus sp. n. ii 176 

truncatus Sharp ii 175 

vulcanus sp. n. ii 179 
Xyletobius Sharp iii 585 

affinis Sharp iii 595 

aleuritis sp. n. iii 595 

ashmeadi sp. n. iii 598 

aurifer sp. n. iii 609 

beddardi sp. n. iii 599 

blackburni sp. n. iii 598 

blackburni van scutellaris n. iii 599 

blackburni var. simplex n. iii 599 

blackburni var. suturalis n. iii 599 

brimneri sp. n. iii 597 

carpenteri sp. n. iii 596 



46 



FAUNA HAWAIIENSIS 



iii 6og 

604 
603 
589 

587 
594 



Xyletobius chryseis sp. n. 
collingei sp. n. iii 
cyphus sp. n. iii 
dollfusi sp. n. iii 
durranti sp. n. iii 
euceras sp. n. iii 
euops sp. n. iii 601 
euphorbiae sp. n. iii 602 
flosculus sp. n. iii 610 
forelii sp. n. iii 6cx3 
fraternus sp. n. iii 606 
fraternus var. laetior n. iii 
grimshawi sp. n. iii 588 
hawaiiensis sp. n. iii 613 
insignis Blackb. iii 602 
kirkaldyi sp. n. iii 600 
lasiodes sp. n. iii 612 
linealus Sharp iii 611 
lineatus var apicalis n. iii 
lineatus var. holomelas n. i 
lineatus var. humeralis n. i 
marmoratus Sharp iii 589 
megalops sp. n. iii 601 
mesochlorus sp. n. iii 592 
meyrickii sp. n. iii 589 
nigrinus Sharp iii 603 
mimus sp. n. iii 594 
niolokaiensis ii 174 
nionas sp. n. iii 603 
numdus sp. n. iii 608 
nuptus sp. n. iii 592 
nuptus var. kauaiensis n 
nudus sp. n. iii 604 
oculatus Sharp iii 595 
pele sp. n. iii 593 
praeceps sp. n. iii 608 
proteus sp. n. iii 590 
proteus var. apicalis n. i 



606 



611 



611 
612 



HI 592 



59 



Xyletobius proteus var. dorsalis n. iii 591 
proteus var. hastatus n. iii 591 
proteus var. maurus n. iii 591 
proteus var. simplex n. iii 590 
roridus sp. n. iii 606 
scotti sp. n. iii 609 
serricornis Blackb. iii 612 
sharpi sp. n. iii 605 
silvestrii sp. n. iii 588 
simoni sp. n. iii O05 
speiseri sp. n. iii 604 
stebbingi sp. n. iii 610 
stebbingi var. notatus n. iii 610 
submimus sp. n. iii 594 
suboculatus sp. n. iii 596 
sulcatus sp. n. iii 613 
sulcatus var. apicalis n. iii 613 
sykesii sp. n. iii 607 
sykesii var. molokaiensis n. iii 607 
walsinghamii sp. n. iii 587 
Xylocopa aeneipennis i 113, lx.\xiii 
Xylophagus spiniger Wied iii 79 
Xylopsocus castanoptera Fairm. iii 643 
Xylopertha religiosa Boisd, iii 643 
Xylothrips religiosa Boisd. iii 643 
Xylotrogus brunneus Steph. iii 644 
Xystophora Hein. i 478 
Xystrocera globosa Olivier ii 96 

Zacranium gen. n. i 295 

oahuense sp. n. i 295 
Zelinae iii 149 
Zelus Fabr. iii 149 

laevicollis Champ, ii 550 

peregrinus sp. n. iii 149, ii 550 

renardi Kol. ii 550 
Zonitidae ii 277 
Z)gocheta iii 338 



CAMBRIDGE: PRINTED BY JOHN CLAY, M.A. .\T THE UNIVERSITY PRESS 



DESCRIPTION OF PLATE I. (VOL. II.) 



ORTHOPTERA. 



Fig. 


I. 


Fig. 


2. 


Fig. 


3- 




is 


Fig. 


4- 


Fig. 


S- 


Fig. 


6. 


Fig- 


7- 


Fig. 


8. 


Fig. 


9- 


Fig. 


lO. 


Fig. 


II. 


Fig. 


12. 


Fig. 


13- 


Fig- 


14. 


Fig. 


IS- 


Fig. 


16 


Fig- 


17- 


Fig. 


18 


Fig. 


19 



Brachymetopa discolor $ ; \ a front view of head of the same. 
B. blackburni ? . 

B. nitida <? ; 3 a variety of the ? ; 3 1^ example in the attitude assumed when the insect 
molested. 

B. unica $ . 

B. inaiiiensis ? . 

B. pannila $ ; 6 a var. hrumiea c? - 

B. deplanata $, . 

Conocephaloides hawaiiensis ? . 

Paratrigonidium roseiim $ - 

P. atroferrugineum $. 

P. viridescefts i . 

P. grande $ . 

P. atteniiatum <$ . 

PrognathogiyUus robi/stus ? . 

P. elongatus 9 . 

Thaumatogryllus variegatus ? . 

Leptogryllus nigro-maculatus ? . 

L. elongatus 9 ■ 

L. simillimus ? . 



FAUNA HAY/AIIE^ISiS. 



VOL. II, PL ATE 1. 




m m 



i 



Perkins Orthoptera. 



^fi^/mn, liih Caml»ialffe 



DESCRIPTION OF PLATE II. (VOL. II.) 

ORTHOPTERA. 

Fig. I. Brachymetopa discolor, apex of abdomen showing cerci in dorsal view; \a the same in 

apical view. 
Fig. 2. B. nitida, cerci in dorsal view ; 2 a apical view of the same. 
Fig. 3. B. deplanata, cerci in dorsal view ; 3 a apical view of the same. 
Fig. 4. B. mmdefisis, cerci in dorsal view ; 4 a apical view of the same. 

Fig. 5. Paratrigonidium attettuatum, var. tni/ior, before the last ecdysis, showing wings and tegmina 
about equally developed. The former entirely disappear at the last moult. 

Fig. 6. Lateral field of superior tegmen of P. filicum ; 6 a the same of P. pacificum. 

Fig. 7. Myrmecophila quadrispina, posterior tibia with four spines on its inner margin. 

Fig. 8. Prognathogryllus alattis, posterior tibia in dorsal aspect ; 8 a apex of tibia viewed outwardly ; 
%b the same on inner side {s, s\ the apical spine of the two series, c' and c' the upper and 
lower calcar in each figure). 

Fig. 9. Aphonogryllus aptery.x ; g a meso- and metathorax in lateral aspect; qb outer, and <)c inner 
aspect of apex of posterior tibia (the lettering as before). 

Fig. 10. Thaumatogryllus variegatus, mesothorax, metathorax and first abdominal segment of $, 
showing the scale-like tegmina; \o a the same parts in the ?, the tegmina less developed {teg. 
tegmina conceahng more or less of the meso- and metathorax, ab i the first abdominal segment) ; 
10/; and loi' the two lateral views of apex of posterior tibia, the lettering as before; \od dorsal 
aspect of apex of posterior tibia of another example, in which the apical spine of either series 
is very far removed from the calcaria. Only the apex of one of these spines is shown, that 
on the other side being situated still further towards the base. 

Fig. II. Nesogryllus stridulans; 11 a superior tegmen of the same enlarged. 

Fig. 12. Leptogryllus nigromaculatus, S tegmina; 12a the same in the ? (lettering as in fig. 10 
and 10 a). 

Fig. 13. Z. similis $, mesothorax with very small tegmina, and metathorax. 

Fig. 14. L. forficularis, posterior tibia in dorsal aspect; 14a and 14;^ outer and inner aspects of 
apex of the same (the lettering as in fig. 8). 

Fig. 15. L. simillimus, posterior tibia in dorsal aspect; 15a and \ib outer and inner aspects of 
apex of tibia of a variety of the same, in which the apical spine of either series is very 
remote from the calcaria (the lettering as in fig. 8). 



FAUNA HAWAII t:HSIS 



VOL 1! PLATE II 




I5b 



l.'i 



£W:^sori,i.it,\ Camirtdt/e. 



Perkins Orthoptera 



DESCRIPTION OF PLATE III. (VOL. II.) 

NEUROPTERA. CHRYSOPIDES. 

Fig. I. Anoinalochrysa prmceps $. 

Fig. 2. A. sylvicola $. 

Fig. 3. A. angidicosta $. 

Fig. 4. A. viridis $. 

Fig. 5. A. lotigipeiuiis ?. 

Fig. 6. A. dcceptor $. 

Fig. 7. A. gayi $. 

Fig. 8. A. rap/iidioides $ . 

Fig. 9. A. biseriata $ . 

Fig. 10. A. hepatica $. 

Fig. II. A. proteus $. 

Fig. 12. A. proteus $ . 

Fig. 13. A. proteus 9 var. 

Fig. 14. A. fulvesceus $. 

Fig. 15. ^. fulvesceus $ var. 

Fig. 16. ^. fulvesceus 9 var. 



FAUNA HAWAIiRN.-:;; 



VOL. 11. PLATE III. 




cmbridge 



Perkins. Neuroptert 



a. 



I 



DESCRIPTION OF PLATE IV. (VOL. II.) 



NEUROPTERA. HEMEROBIIDES and CHRYSOPIDES. 



Fig- 
Fig. 
Fig. 
Fig. 

Fig- 
Fig. 

Fig. 
Fig. 
Fig. 
Fig. 
Fig. 
Fig. 

Fig- 
in dorsal 

Fig- 
Fig. 
Fig- 
Fig. 
Fig. 
Fig. 
Fig. 



Megalomus hospes. 
M. Iiospcs vai'. 
Nesomicromus vagus. 
N. latipennis. 
N. angiistipennis. 
N. drepanoides. 
N. paradoxus. 
N. bellulus. 
JSI. brunnescens. 
N. subochraceics. 
N. minimus. 
Pseudopsectra lobipennis. 
Nesotliauma haleakalae ; 



I. 

2. 

3- 

4- 
5- 
6. 

7- 
8. 

9- 

10. 

n. 

12. 

13- 

aspect. 

14. Nesomicromus forcipatus, $ terminal segments in dorsal aspect; 14(7, the same in lateral view. 

15. N. loiigispinosus, $ terminal segments in dorsal aspect. 

16. Megaknnus hospes, $ terminal segments in lateral view ; i6(7, the same seen from the ape.\. 

17. Anomalochrysa hepatica, apical segments of abdomen of $ in lateral view. 

18. A.frafer, apical segments of abdomen of $ in lateral view. 

19. A. deceptor, apical segments of abdomen of $ in dorsal aspect; 19a, the same in lateral view. 

20. A. princeps, apical segments of abdomen of $ in dorsal aspect; 20a, the same in lateral view. 



13(7, the same in profile; 131^, terminal segment and appendices 



FAUNA HAWAIIENSIS 



VOL.n.PLATK.IV. 




,y 



A 



>amm 



'-^ 




12 






13a. 



■* 




13b 



Perkins. Neuroptera 



Hdwin W'jlscnjith. Cambridge . 



DESCRIPTION OF PLATE V. (VOL. II.) 

ODONATA. 

Fig. I. Agrion xanthomelas $. 

Fig. 2. A. jugorum $ . 

Fig. 3. A. heterogamias $. 

Fig. 4. Terminal segment and appendices of Agrion xanthomelas i in dorsal ; 4a, in 
external lateral view. 

Figs. 5 & 5^. The same parts in A. nigrohamafum. 

Figs. 6 & i>a. The same in A. pacifiniin. 

Figs. 7 & 7(7. The same in A. koelense. 

Figs. 8 & 8«. The same in A. oresitroplmm. 

Figs. 9 & 9(7. The same in ^. calliphya. 

Figs. 10 & io(7. The same in A. nesiotes. 

Figs. II & 11(7. The same in A. jugorum. 

Figs. 12 & 12(7. The same in A. oahuense. 

Figs. 13 & 13(7. The same in ^. vagabundum. 

Figs. 14 & 14(7. The same in A. oceanicum. 

Figs. 15 & 15(7. The same in A. blackburni. 



FAUNA HAWAliEN^lS. 



PLATE Y. 







4a 





Perkins. Neuropter 



a. 



DESCRIPTION OF PLATE VI. (VOL. II.) 

COLEOPTERA. CERAMBYCIDAE. 

Fig. I. Clytarlus mediocris $. 

Fig. I a. Terminal ventral segments of Clytarlus filipes $ . 

Fig. I b. „ „ " mediocris c? . 

Fig. 2. Clytarlus pennaius i. 

Fig. 3- '. » ?• 

Fig. 4. C. longipes $ . 

Fig. 5. FlagitluiiysHS vitticollis $ . 

Fig. 6. P. permundus. 

Fig. 7. P- newelli ? . 

Fig. 8. P. concolor 5 • 

Fig. 9. P- ciineatus ? . 

Fig. 10. P. funebris $. 

Fig. II. P. diana. 

Fig. 12. P. bishopi. 

Fig. 13. /". collaris $. 

Fig. 14. jP. bilineatus ?. 

Fig. 15. /'. perkinsi ?. 

Fig. 16. /*. danviniamis $. 

Fig. 17. /". sulphurescens 6- 

Fig. 18. Z'. albcrtisi i. 

Fig. 19. /'. inunroi ? . 

Fig. 20. P. arachnipes ? . 

Fig. 21. P. cristatus $. 



I'^AUNA HAWAlIfiNSIS. 




Sharp. Cole opt era. 



Fig. 


I 


Fig- 


2 


Fig. 


3 


Fig. 


4 


Fig. 


5 


Fig. 


6. 


Fig. 


7 


Fig. 


8 


Fig. 


9 


Fig. 


lO 


Fig- 


II. 


Fig. 


12 


Fig. 


13 


Fig- 


14 


Fig. 


15 


Fig. 


16 


Fig- 


17- 


Fig. 


18. 


Fig. 


19 


Fig. 


20 



DESCRIPTION OF PLATE VII. (VOL. II.) 

COLEOPTERA. CURCULIONIDAE. 

Rhyncogonus nitidus. 

R. squatniger. 

R. stygius. (N.B. Although in the figure this species appears very like No. i, it is really 

very different, being opaque, whereas R. nitidus has the surface shining.) 
R. freycinetiae. 
R. koebelei. 
R. depressus. 
R. vittatus. 
R. vestitus. 

Acalles lateralis $ (Kauai form). 
A. callichroma. 
A. melanolepis. 
A. leptothorax. 
A. duplex $. 
A. duplex 9 . 
A. nigripennis. 
Dryophthorus nesiotes. 

D. homorhynchus ; 1 7 (7, head and rostrum of 5 ; 171^, head and rostrum of $ . 
D. declivis. 
D. insignis. 
D. verticalis. 



FAUNA HAWAIIRNSIS, 



VOL. [I. PLATE YII. 




Perkins Coleoptera 



K Wilson. !ith Camiirjugc 



DESCRIPTION OF PLATE VIII. (VOL. II.) 

COLEOPTERA. CURCULIONIDAE. 

Thalattodora insignis ; i a, antenna of the same. 
Orothreptcs callithrix ; 2 a, antenna of the same. 
Deinocossoiius nesioUs var. hawaiiensis ; 3 a, antenna of the same. 
Haloxenus immigrant. 
Nesolocus kauaiensis. 
Dysomina sxlvicola. 
Heteraviphiis filiami. 
H. molokaiensis. 
Figs. 9 & 10. H. cylindricus (large and small form). 
H. kauaiensis. 
Oodemas olindae 9 var. 

O. longicorne $; 13(7, anterior tarsi of the same. 
O. molokaiense ; 14 a, anterior tarsi of the same. 
O. chrysodorHm $ ; 1 5;?, anterior tarsi of $ ; 15/', anterior tarsi of 
O. graciliforme. 
O. leiothorax. 
O. pukhnim. 
0. oblongum. 
O. grande. 
0. corticis. 
O. pachysoma. 
O. aencscens. 
O. sculpiuratum. 
O. mauietise. 
O. striatum. 
Anothcorus robust us. 



Fig. 


I. 


Fig. 


2. 


Fig. 


3- 


Fig. 


4- 


Fig- 


5- 


Fig. 


6. 


Fig- 


7- 


Fig. 


8. 


9 & 


10. 


Fig. 


II. 


Fig. 


12. 


Fig. 


13- 


Fig. 


14. 


Fig. 


15- 


Fig. 


16. 


Fig. 


17- 


Fig. 


18. 


Fig. 


19. 


Fig. 


20. 


Fig. 


21. 


Fig. 


22. 


Fig. 


23- 


Fig. 


24. 


Fig. 


25- 


Fig. 


26. 


Fig. 


27- 



FAUNA HAWAI.'BNSIS. 



VOL. [I. PLATE VIII. 







Perkins. Coleoptera. 



Edwin Wi/sonjith. Cambridge . 



DESCRIPTION OF PLATE IX. (VOL. II.) 

COLEOPTERA. PROTERHINIDAE. 

Fig. I. Proterhinus gigas $. 

Fig. 2. P. eugonias 5 . 

Fig. 3. P. etilepis i. 

Fig. 4. P. wiksiroe?niae i. 

Fig. 5. P. laticornis $ . 

Fig. 6. P. scrricornis, antenna $ . 

Fig. 7. P. eurhopalus, antenna $ . 

Fig. 8. p. kptophyas ?. 

Fig. 9. P. maculifer $. 

Fig. 10. P. kamptarthrus $. 

Fig. II. P. deinops ?. 

Fig. 12. P. platygonias $. 

Fig. 13. P. leptothrix $. 

Fig. 14. /". pachycnemis cJ. 

Fig. 15. -P. laticoUis $ ; 15 «, front tarsi of the same. 

Fig. 16. P. validus $. 

Fig. 17. P. sharpi $. 

Fig. 18. P. tubercuUceps $. 

Fig. 1 9. Z'. sternalis $ ; ig a, front tarsi of the same. 

Fig. 20. Z'. microtarsus $ ; 20 a, front tarsi of the same (drawn on same scale as 19^). 

Fig. 21. P. arhopalus $. 

Fig. 22. P. mirahilis $. 

Fig. 23. P. alyxiae $. 

Fig. 24. P. pteridis $. 



FAUNA HAWAIIENSIS. 



VOL. [[.PLATE IX. 




Perkins. Coleoptera 



i'Mi/mn, ilA. Camdndgr 



DESCRIPTION OF PLATE X. (VOL. II.) 
COLEOPTERA. PROTERHINIDAE, SCOLYTIDAE, CISTELIDAE, CIOIDAE. 



Pig- 


I. 


Proterhiinis detritus ? . 


Fig. 


2. 


P. epitrdiis ? . 


Fig- 


3- 


P. kaalac $. 


Fig. 


4- 


P. oahuensis $ . 


Fig. 


5- 


P. b/ackburiii var. bisignatus i. 


Fig- 


6. 


P. osculans ? . 


Fig. 


7- 


P. persimilis i . 


Fig. 


8. 


P. hawaiiensis ? . 


Fig- 


9- 


P. curhynchus $, front of head. 


Fig. 


10. 


Xyklwrus molokaiensis '} . 


Fig- 


II. 


X. immaturus $ ; 11 a, $ . 


Fig. 


12. 


X. exsectus i. 


Fig. 


13- 


X. vulcaniis $ . 


Fig- 


14. 


Apterocis la7iaiensis. 


Fig. 


15- 


A. 07-naiipennis. 


Fig- 


16. 


A. sirigosus. 


Fig. 


17- 


Cis haleakalae. 


Fig. 


18. 


C. nigro-fasciatui. 


Fig. 


19. 


C. insidaris. 


Fig. 


20. 


C. signatus ; 20 a, zob, 20 c, ott 


Fig. 


21. 


Labetis coinitans. 


Fig. 


22. 


Cistela kauaiensis. 



FAUNA HAWAIi: 



Vi)i..Il,ri.ATK X. 




SOa 



SOh 



^Oc 



■erkins, Goleopiera. 



r^ 



^-v 



DESCRIPTION OF PLATE XI. (VOL. II.) 



MOLLUSCA. 



Fig. 



Fig. 


1 . 
2. 


Fig. 


3- 


Fig. 


4- 


Fig. 


S- 


Figs 


6,7 


Fig. 


8. 


Fig- 


9- 


Fig. 


10. 


Fig. 


II. 


Fig. 


12. 


Fig. 


•3- 


Fig. 


14. 


Figs. IS 


16. 


Figs. 17 


18. 


Figs. 19 


20. 


Fig. 


21. 


Fig. 


22. 


Fig. 


23- 


Fig. 


24. 


Fig. 


25- 


Fig. 


26. 


Fig. 


27. 


Fig. 


28. 


Fig. 


29. 


Fig. 


3°- 


Fig. 


31- 


Fig. 


32. 


Fig. 


33- 


Fig. 


34- 


Fig. 


35- 


Fig. 


36. 


Figs. 37, 


38. 


Figs. 39, 


40. 


Figs. 41, 


42. 


Figs. 43. 


44- 


Figs. 45. 


46. 



Tornatellina compacta, sp. nov. (p. 380). 

Achatinella {Fartulina) hayseldeni Baldwin (p. 313). 

Amastra simularis Hartm., van roseoiincta Sykes (p. 344). 

Amastra citrea Sykes (p. 335). 

Perdicella fulgurans, sp. nov. (p. 329). 

Achatinella cestus Newcomb (p. 300). 

Leptachatitia impressa Sykes (p. 364). 

Lcptachatina imitatrix, sp. nov. (p. 364). 

Leptachafina emerita, sp. nov. (p. 361). 

Leptachatina convexiuscula, sp. nov. (p. 360). 

Leptachatlna setnipida Sykes (p. 369). 

Leptachatina konaensis, sp. nov. (p. 384). 

Tornatellina perkinsi, sp. nov. (p. 382). 

Achatinella {Partiili?ia) redfieldi Newcomb (p. 317). 

Atiriculella perkinsi, sp. nov. (p. 377). 

Achatinella {Achatittellastrum) wailuaensis, sp. nov. (p. 328). 

Leptachatina arhorea, sp. nov. (p. 357). 

Leptachatina sup7-acostata, sp. nov. (p. 370). 

Amastra {Laminella) fraterfia Sykes (p. 349). 

Amastra {Laminella) villosa Sykes (p. 352). 

Siiccinea protracta, sp. nov. (p. 388). 

Leptachatina conicoides, sp. nov. (p. 359). 

Leptachatina vana, sp. nov. (p. 372). 

Tornatellina cylindrica, sp. nov. (p. 381). 

Leptachatina smithi Sykes (p. 369). 

Leptachatina perkinsi Sykes (p. 367). 

Tornatellina trochoides, sp. nov. (p. 383). 

Siiccinea cinnamomea Ancey (p. 386). 

Kaliella konaensis Sykes (p. 286). 

Siiccinea konaensis Sykes (p. 387). 

Amastra longa Sykes (p. 338). 

Newcombia perkinsi Sykes (p. 332). 

Endodonta {Nesophila) lanaiensis Sykes (p. 291). 

Eiidodonta {Thaiimatodon) ringens Sykes (p. 288). 

Philo)iesia perkinsi Sykes (p. 284). 

Vitrea lanaiensis Sykes (p. 278). 

Vitrea molokaiensis Sykes (p. 278). 



FAUNA HAWAII EN SIS. 



VOL 11. PLATE XI. 













10 



ji 





u 









73 



16 



/7 



/* 



/.V 








30 








26 



25 



28 



32 







2i) 



33 



.?# 



37 






:U} 



SH 



n 




35 




/.-i 



45 





J Green, del eihth-. 



38 




^ 



40 



44 



4o 



Mintern Bros, imp . 



Sykes,Mollusca. 



DESCRIPTION OF PLATE XII. (VOL. II.) 



MOLLUSCA. 



Fig. I. 



Fig. 


2. 


Fig. 


2 a. 


Fig. 


3- 


Fig. 


3(7. 


Fig. 


4- 


Fig. 


5- 


Fig. 


5 a. 


Fig. 


S'''- 


Fig. 


5^- 



Philonesia baldwini Ancey (p. 281). 

Mantle margin, with shell and dorsal lobes, a, denotes white spots on integument covering 

the branchial cavity. 
Extremity of foot from left side. 
Portion of foot showing sole. 
Genitalia, not quite complete. 

Male organs enlarged, to show the retractor muscle and coiled vas deferens. 
Jaw. 

Centre and two side teeth of radula. 
Median teeth, 7th — nth. 
Lateral teeth, 19th — 22nd. 
Eleven of the outermost, or marginal, teeth. 



Fig. 
Fig. 
Fig. 
Fig. 
Fig. 
Fig. 
Fig- 
Fig. 
Fig. 



GODWINIA CAPERATA Gould (p. 277). 

6. Animal with shell removed, showing dorsal lobes. 

7. Portion of mantle zone, near respiratory orifice. 

8. Extremity of foot. 

g. Buccal mass, and salivary gland, &c. 

10. Sole of foot. 

11. Jaw. 

12. Central tooth of radula. 

1 2 a. Fifth intermediate, and following lateral teeth. 

\2b. Outermost, or marginal, teeth. 



Fig. 13. Leptachatina acuminata Gould. Central teeth of radula (p. 357) 
Fig. 13(7. Intermediate and marginal teeth. 
Figs. 14, 14 «. Ancyliis sharpi, sp. nov. (p. 394). 



EXPLANATION OF THE LETTERING. 



Al. Gd. albumen gland. 

B. 711. muscle of buccal mass. 

Geti. ap. generative aperture. 

h. d. hermaphrodite duct. 

/. intestine. 

/. d. I. left dorsal lobe. 

/. s. I. left shell lobe. 

P. male organ. 



pr. 


prostate. 


r. d I. 


right dorsal lobe. 


r. m. P. 


retractor muscle of penis 


r. s. I. 


right shell lobe. 


Sal. gld. 


salivary gland. 


St. 


stomach. 


V. d. 


vas deferens. 



The numerical digits in smaller type indicate the tooth figured, reckoning from o, the central tooth. 
All figures (except 14, \\a) are from dissections and drawings made by Lt.-Col. H. H. Godwin-Austen, F.R.S. 



FAUNA HAV/AII5NSiS. 



'OL.;; PLATE All 




V^^fy^/J^jy 4 




e .? 




\JI 



^ 



'// 



/^<'X4 




lo 



Sykes.MoUusca. 



'^. 



i' ^m 



ai.gid.: 



J'' 

I 



MniemBrcs .r/ir 



V. 



DESCRIPTION OF PLATE XIII. (VOL. II.) 
ENTOZOA. 

LIST OF ABBREVIATIONS. 



b. 


Brain. 


71. s. 


Central nerve ganglion. 


c. m. 


Circular muscles. 


11. 


The amoeboid nuclei of the skin and the 


c. 


Cuticle. 




lemnisci. 


d. e. c. 


Dorsal excretory canal. 


0. 


Ovary. 


c. 


Ectoderm. 


P- 


Nucleus of parenchymatous cell. 


e. m. 


Masses of ova. 


p.c. 


Parenchyma cell. 


f.c. 


? Flame-cell. 


p.e. 


Knob-like ends of ectoderm cells under 


g.d. 


Genital duct. 




cuticle. 


g-P- 


The external opening of the duct. 


r. s. 


Receptaculum seminis. 


I. <t la. 


The lacunae in the skin. 


r. 


Rostellum. 


k. 


The lemnisci. 


s. 


Coagulated masses of spermatozoa in the 


li. 


The ligament. 




body-cavity of the female. 


I. I. 


The large lateral lacunae of the trunk. 


t. 


Testis. 


I. m. 


Longitudinal muscles. 


u. 


Uterus. 


I. H. 


Lateral nerve. 


V. e. c. 


Ventral excretory canal. 


m. 


The muscles from which the ligament arises. 


V. s. 


Vesicula seminalis. 


n. c. 


Nerve cell. 







Fie. I. 



Fig. 


2 


Fig- 


3 


Fig. 


4 


Fig. 


5 



Fig. 6. 



Fig. 7- 

Fig. 8. 

Fig. 9. 

Fig. 10. 



A view of Drepanidotaenia hemignathi, x 15. The dark patches in the anterior two-thirds of 

the body are caused by the generative organs ; in the posterior third they represent the eggs 

in the uterus. 
An isolated hook from the rostellum, x 500. 
A longitudinal section through the head, x 100. The rostellum, r., is retracted. The point 

of fusion of the two lateral nerves is shewn at n. s. The section passes between the 

suckers. 
A transverse section through a mature proglottis, x 70. 
A longitudinal section, somewhat oblique, showing the regular arrangement of the longitudinal 

muscles, x 50. 
A longitudinal section through several mature proglottides, x 50. This shows the transverse 

connection between the two ventral longitudinal excretory canals and the transverse lines 

formed by the concentration of the parenchyma at the posterior end of each proglottis. 
A view of the head in an expanded, flattened-out state, x 60. 
A view of another head in a contracted, bunched-up condition, x 40. 
A transverse section through the head, showing the ten hooks on the rostellum and the 

four suckers. 
A portion of a proglottis, highly magnified to show the minute anatomy, x 450. 



r^AUNA HAV/AIIENSIS, 



VOL. n. PLATE. X!il. 




..fiO. 



S Irn. 



SMpley. Entozoa. 



'^mbridge- 



DESCRIPTION OF PLATE XIV. (VOL. II.) 

ENTOZOA. 

For explanation of letters see description of Plate XIII. 

Figs. II, 12, and 13. Three views of three different specimens of ArhyncJius hemigiiafki. Each x 20. 

The division of the body into three regions is well marked. The details are shown in Fig. 1 1. 

Figs. 12 and 13 are rough sketches. 
Fig. 14. A transverse section through the head of a female, crowded with ova and egg-masses; the 

ligament is shown in section, x 40. 
Fig. 15. A transverse section through the same, just below the edge of the collar. In the centre is the 

neck, which fuses with the collar a few sections further back. The big circular canal of the 

collar is shown at /., x 40. 
Fig. 16. A transverse section through the trunk of the same. The uppermost lemniscus is cut in two 

places. The ovary is double, and shows egg-masses as well as eggs ; some coagulated 

masses of spermatozoa are lying in the body-cavity, x 40. 
A surface view of the external opening of the genital duct, x 40. 
Some developing ova, highly magnified. 
A transverse section through the trunk near the genital pore, taken from the same series as 

Figs. 14, 15, and 16. It shows part of the funnel-shaped internal opening of the genital 

duct, g. d., X 40. 
Fig. 20. A transverse section from another specimen taken behind the opening of the genital duct. 

This shows the arrangement of the lacunae and their communications with the lateral 

lacunae, /. /. 
Fig. 21. A longitudinal section through the central part of the skin of the head, showing the origin of 

the ligament and the ganglion cells of the brain, lying in a mass of ova and egg-masses. 
Fig. 22. A small portion of the skin in section, showing the single layer of circular muscle-fibres, x 40. 
Fig. 23. A median longitudinal section through a female. The whole body-cavity full of ova and 

egg-masses. The ligament is seen in the head, and the genital duct near its opening in the 

trunk. The left lemniscus, cut twice, is alone seen, x 30. 
Fig. 24. A transverse section through the trunk of a male, showing one of the testes. This section 

shows also the longitudinal muscles on the lemnisci and the large lateral lacunae. Inf. lac, 

X 40. 
Fig. 25. A transverse section through the head of a male, showing the brain in the ligament, and the 

longitudinal muscle-fibres very well developed, x 40. 



Fig. 


17- 


Fig- 


18. 


Fig. 


19- 



FAUNA HAWAIIENSIS. 



VUL. 11. PL ATE XIV. 







Shipley. Entozoa. 



£c/><m Wilscn.Jith. Cambridge. 



DESCRIPTION OF PLATE XV. (VOL. II.) 

ARANEAE. 

Fig. I. Ariad/ia perkinsi "K. Sim. $ ; la, maris pes-maxillaris ; i/^, ejusdem pes primi paris. 

Fig. 2. Ariamnes corniger E. Sim. $ . 

Fig. 3. Theridion cainpestratHin E. Sim. ? . 

Fig. 4. T. praetiwtum E. Sim. ; 4«, $ pes-maxillaris. 

Fig. 5. T. haleakalense E. Sim. ; 5^, i pes-maxillaris. 

Fig. 6. T. perkinsi E. Sim. ? ; 6(7, $ pes-maxillaris. 

Fig. 7. T. kauaicnse E. Sim. 5 • 

Fig. 8. T. grallator E. Sim. $ (puUus) ; 8(7, (J pes-maxillaris. 

Fig. 9. T. actttitarse E. Sim. 9 ; 9^7, c? pes-maxillaris. 

Fig. 10. T. me/ill II 111 E. Sim. ? , loa, c? pes-maxillaris. 

Fig. II. T. maiiiense E. Sim., maris pes-maxillaris. 

Fig. 12. T. posticatiim E. Sim., maris pes-maxillaris. 



FAUNA HAWAIIENSIS 



VOL, II. PL ATP: XV. 




Edwin Wilson.hth. Cambridge 



E. Simon, Araneae. 



DESCRIPTION OF PLATE XVI. (VOL. II.) 

ARANEAE. 

Fig. I. Labulla torosa E. Sim. $; la, pes-maxillaris ; i/>, ejusdem tarsus pronus ; ic, chele maris 
supina; id, chele feminae supina. 

Fig. 2. Z. graphica E. Sim. $; 2a, pes-maxillaris; 2/', ejusdem tarsus pronus; 2c, epigynum feminae; 
2d, chele feminae supina ; 2e, chele maris supina. 

Fig. 3. Cyclosa xanthomelas E. Sim. ? . 

C. simp/icicauda E. Sim. 5 • 

C. oloriiia E. Sim. ? . 

Aranetis emmae E. Sim. 9 ; 6^, epigynum feminae ; db, ejusdem plagula inferior. 

A. kapiolaniae E. Sim. ? ; "ja, epigynum feminae. 

Doryonychus raptor E. Sim. $. 

Lycosclla spiiupes E. Sim. 9 • 

L. annulata E. Sim. 5 . 

Ulesanis oahiiensis E. Sim. $ ; cephalothorax abdomenque a latere exhibita. 



•- '5- 

Fig. 


5- 


Fig. 


6. 


Fig- 


7- 


Fig. 


8. 


Fig. 


9- 


Fig. 


10. 


Fig. 


1 1. 



/^AUNA HAWAlIBNSiS, 



VOL. II, PL ATE XYI. 




E Simo/i. Araneae. 



DESCRIPTION OF PLATE XVII. (VOL. II.) 

ARANEAE. 

Fig. I. Misumena nigrofrenata E. Sim. 9. 

Fig. 2. Syiiaema iiaevigertim Y,. Sim. ¥; 2a, pes-maxillaris maris. 

Fig. 3. 6'. dimidiatipes E. Sim. $ . 

Fig. 4. Mecaphesa semispinosa E. Sim. ? ; 4a, plagula genitalis. 

Fig. 5. Proernus velox Yj. 'i>VLi\. $; 5^7, processus tibialis pedis-maxillaris maris. 

Fig. 6. P. schaitinslandi E. Sim. $ ; 6(7, oculi proni ; 6/;, chela supina ; dc, pes-maxillaris maris ; 
(id, ejusdem processus tibialis a latere exhibitus. 

Fig. 7. Pagiopaliis atotnarius Y,. 'awsx. i; 7a, pes-maxillaris maris ; 7/^, ejusdem processus tibialis. 

Fig. 8. P. persoiiatus E. Sim. ? . 

Fig. 9. Adrastidia longula E. Sim. $ . 

Fig. 10. A. stigmaiica E. Sim. $. 

Fig. II. Misumena anguliventris E. Sim., maris pes-maxillaris; \\a, feminae plagula genitalis. 

Fig. 12. M. ve/ata E. Sim., maris pes-maxillaris; 12a, feminae plagula genitalis. 

Fig. 13. Z'/ire'a /«W««a Keyserl., processus tibialis pedis-maxillaris maris ; 13(7, id. varietas. 

Fig. 14. D. vitellina E. Sim., processus tibialis pedis-maxillaris maris. 

Fig. 15. Pagiopalus semipunctatiis E. Sim., pes-maxillaris maris; 153, ejusdem processus tibialis. 



FAUNA HAWAI1EN31S 



vol.[[.platf/>:yii. 




E.Simon. Araneae. 



Sdwin V/'ilsonJsth. Cambridge ■ 



DESCRIPTION OF PLATE XVIII. (VOL. II.) 
ARANEAE. 

Syroloma major E. Sim. $ ; irt, ungues tarsorum ; \b, plagula genitalis. 

S. tninor E. Sim. $ . 

Lycosa likelikeae E. Sim. $ ; 3(7, plagula genitalis. 

L. kalukanai E. Sim., plagula genitalis. 

Lycosella spinipes E. Sim., plagula genitalis. 

Sandalodes validiis E. Sim. $. 

S. pubens E. Sim. $ ; 7a, pes-maxillaris maris. 

6'. albociliatus E. Sim. $ . 

6'. navatus E. Sim. $ . 

S. vereciindiis E. Sim. ? . 

S. cruciatus E. Sim. $. 

S. senicuhis E. Sim. $ . 

S. canosus E. Sim. 9 ■ 
Fig. 14. Cnephalocotes simplkiceps E. Sim., pes-maxillaris maris. 
Fig. 15. Alicnmeta i/isiila/ia E. Sim., chele maris prona ; 15a, pes-maxillaris. 



Fig. 


I. 


Fig. 


2. 


Fig. 


3- 


Fig. 


4- 


Fig. 


5- 


Fig. 


6. 


Fig. 


7- 


Fig. 


8. 


Fig- 


9- 


Fig. 


10. 


Fig. 


II. 


Fig. 


12. 


Fig. 


13- 



FAUNA HAV/AIIEKSIS. 



VOL.11, PLATE. XYlll 




Edwin I'/' 



E Simon. Araneae 



DESCRIPTION OF PLATE XIX. (VOL. II.) 

ARANEAE. 

Fig. I. Doryonychus raptor E. Sim.; \a, chele maris prona ; i/', id. supina ; ic, processus tarsalis 
maris; \d, chele feminae prona; ic, id. supina; \f, ungues tarsorum i' paris intus ; 
i^, id. extus. 

Fig. 2. Tetragiiatlia uncifcra E. Sim.; 2(7, chele maris prona; 2b, id. varietas ; 2C, id. supina; 2d, chele 
feminae prona ; 2f, id. supina. 

Fig. 3. T. ameiventris E. Sim. ; 3^, chele maris prona ; 2)^, id. supina ; 3<r, abdomen a latere 
exhibitum. 

Fig. 4. T. hawaieiisis E. Sim. ; 4^, chele feminae prona ; 4/', id. supina. 

Fig. 5. T. perkinsi E. Sim. ; 5rt, chele maris prona; 5/^, id. supina; 5c, chele feminae prona ; 5;/, id. 
supina. 

Fig. 6. T. netrix E. Sim. ; 6(7, chele maris prona ; 6/', id. supina ; 6c, chele feminae prona ; dd, id. 
supina ; 6c, processus tarsalis maris. 

Fig. 7. T. maiidihulata Walck. ; 7(7, chele maris prona ; 7^, id. supina ; 7c, chele feminae prona ; 
id, maris processus tarsalis. 

F'ig. 8. T. sobrina E. Sim. ; 8(?, chele maris prona ; 8/', id. supina ; 8c, processus tarsalis maris. 

Fig. 9. T. kaiiaiensis E. Sim.; 9(?, chele maris supina; 91^, id. supina; 9c, chele feminae prona; 
()d, id. supina. 

Fig. 10. T. restricta E. Sim.; \oa, chele maris prona; loi^, id. supina; loc, chele feminae prona; 
\od, id. supina. 



FAUNA HAWAHRNSIS, 



VO;..n.PLATRXlX. 




Edwin IV/7son,Jir/i. Cambridge . 



E. Simon ^ Araneae. 
(Tetragnatha,.) 



DESCRIPTION OF PLATE XX. (VOL. II.) 
CRUSTACEA. ISO POD A. 

Figs, la & li. Annadillo hawaiiensis Xixc\-3.; kt, clypeus $; i/', clypeus ?. 

Figs. 2a — 2h. A. albospinosus, sp. nov. ; 2a, cephalon et premier segment pereial (c?); 2b, cephalon 
(vii en dessous) {$); 2c, extremite de I'antenne ; 2d, partie laterale (coxale) des deux premiers 
segments pereiaux (vue en dessous) ; 2t', cinquieme segment pleonal, pleotelson et uropodes 
( c? ) ; 2/j premier pleopode {i); 2g, uropodes et pleotelson (vus en dessous) {$); 2//, cephalon 
et premier segment pereial ( ? ). 

Figs. 3rt — 3/ A. perki)isi, sp. nov. $; T,a, cephalon et premier segment pereial ; T,b, cephalon (vu en 
dessous); y, extremite de I'antenne; 31/, partie late'rale (coxale) des deux premiers segments 
pereiaux (vue en dessous) ; y, cinquieme segment pleonal, pleotelson et uropodes ; 3/, premier 
pleopode. 

Figs. A,a — 4^'-. A. danae, sp. nov. $ ; 4a, cephalon et premier segment pleonal ; 4/', cephalon (vu en 
dessous) ; ^c, extremite de I'antenne ; 4^, partie laterale (coxale) des deux premiers segments 
pereiaux (vue en dessous) ; 41', cinquieme segment pleonal, pleotelson et uropodes ; 4/, premier 
pleopode ; 4^, uropode et pleotelson. 

Figs. 5(7 & c^b. A. sharpi, sp. nov. 5 ; S'^ cephalon et preinier segment pereial ; 5/', cinquieme segment 
pleonal, pleotelson et uropodes. 

Figs. 6(? — 6/. Geoligia perkinsi, sp. nov. $ ; da, cephalon et deux premiers segments pereiaux ; 
6/', cephalon (vu en dessous) ; 6c, antenne ; dd, pleotelson et uropode ; de, levre inferieure 
6//', mandibules ; 6^'-, premiere maxille ; dh, deuxieme maxille ; 6/, maxillipede. 



FAUNA HAWAHENSIS 



VOL II. PLATE XX. 




Edmn Wilsonjith. Cambridge . 



A-Dollfus. Crustacea, Isopoda. 



DESCRIPTION OF PLATE XXI. (VOL. II.) 
CRUSTACEA. AMPHIPODA. 

A. 

Orchcstia plataisis Kroyer. 

At the top ; lateral view of head and gn athopods, with lateral view of pleon immediately below. 
a. s. First antenna. 

gn. 1, 2. First and second gnathopods of the male, with portions more highly magnified. 
j/r/>. I, 3, First and third uropods. 

mx/>. Maxillipeds, with palp of one side detached. These are from a second (male) specimen. 
gn. I ? , gn. 2 ? . First and second gnathopods of a female specimen, with terminal part of gn. i more 
highly magnified. 

B. 

Orchcstia pickeringh Dana. 
O. P. gn. 2. Second gnathopod of male. 

C. 

Paroichestia /imcaiensis Dana, $. 

At the top ; lateral view of head and gnathopods. 
/. s. Upper lip. 
nixp. Maxillipeds. 

gn. I, 2. First and second gnathopods, with portions more highly magnified. 
tcrp. I, 2, 3. First, second, and third uropods, third also more highly magnified. 

T. Telson, to the same scale as the uropods, and also more highly magnified, to the same scale as 
upper lip and maxillipeds. 



FAUNA HAWAIIBNSIS 



-/OL. 11. PLATE XXI. 




T R R,S Del 



2d:vjn I;}'- - '■■'- ^'a.inbridge 



Slebbmg. Crust ac e a , Amphipoda. 



^ 



ZOOLOGICAL RESULTS based on material from New Britain, •" Guinea, Loya 
Islands and elsewhere, collected during the years 1895, i >, and 1897, 
Arthur Willey, D.Sc. Lond., Hon. M.A. Cantab., Late Bait Student of t 
University of Cambridge. Demy 410. Parts I, II, and III. -.e 12s. 6d. eai 

Parts IV and V. Price 21s. each. Part VI (completing the work) Price 12s. 6d. 

PART I. 1898. 

1. The anatomy and development of Peripatus novae-britanniae. By Arthur A^illey, M.A., D. 

With Plates I. — IV. and 7 figures in the text. 

2. Metaprotella saiidaknsis, n. sp. [Caprellidae]. By Dr Paul Mayer. With 6 figures in the te; 

3. On a little-known Sea-snake from the South Pacific. By G. A. BOULENGER, F.R.S. 

With Plate V. 

4. Report on the Centipedes and Millipedes. By R. I. POCOCK. With Plate VI. 

5. Account of the Phasmidae with notes on the eggs. By D. Sharp, M.A., F.R.S 

With Plates VII.— IX. 

6. Scorpions, Pedipalpi and Spiders. By R. I. PocoCK. With Plates X. and XI. 

PART II. 1899. 

7. Report on the specimens of the genus Millepora. By SYDNEY J. HiCKSON, M... D.Sc, F.R 

With Plates XII.— XVI. 

8. Report on the Echinoderms (other than Holothurians). By F. JEFFREY BELL, \ 

With figures on Plate XVII. and one figure in the text. 

9. Holothurians. By F. P. Bedford, B.A. With figures on Plate XVII. 

10. Report on the Sipunculoidea. By Arthur E. Shipley, M.A. With Plate XVIi. 

11. On the Solitary Corals. By J. STANLEY GARDINER, M.A. With figures on Plates X. and ' 

12. On the postembryonic development of Cydoseris. By J. Stanley Gardiner, M.. 

With figures on Plates XIX. and XX. 

13. On a collection of Earthworms. By FRANK E. Beddard, M.A., F.R.S. With Pla.. XXI. 

14. The Gorgonacea. By ISA L. HiLES, B.Sc. With Plates XXII. and XXIII. 

PART III. 1899. 

15. Orthogenetic variation in the shells of Chelonia. By HANS Gadow, M.A., Ph.D., P S. 

With Plates XXIV.— XXV. and one text-figure. 

16. Enteropneusta from the South Pacific, with notes on the West Indian Species. .y Arti 

Willey, D.Sc, Hon. M.A. Cantab. With Plates XXVI.— XXXII. and seven text-figures 

17. On a collection of Echiurids from the Loyalty Islands, New Britain and China Straits, wit' an attei 

to revise the grouji and to determine its geographical range. By ARTHUR E. Shipll ', M.A. 
With Plate XXXIII. 

PART IV. 1900. 

18. On the anatomy of a supposed new species of Coenopsmnmia from Lifu. By J. jTAN: 

Gardiner, M.A., Fellow of Gonville and Caius College, Cambridge. With one plat.-. 

19. On the Insects from New Britain. By D. Sharp, M.A., M.B., F.R.S. With one plate. 

20. Report on the Stomatopoda and Macrura. By L. A. Borradaile, M.A., Lecturer of Seh 

College, Cambridge. With four plates. 

21. Report on the Slugs. By Walter E. Collinge, F.Z.S., Mason College, Birmingham 

With two plates. 

22. Report on the Polyzoa. By E. G. Philipps, Newnham College, Cambridge. With two ates 

23. The Hydroid Zoophytes. By Laura Roscoe Thornely, University College, Liverpoo 

With one plate. 

24. Astrosclera willeyana, the Type of a new Family of Calcareous Sponges. By J. J. LlSTl , M 

St John's College, Cambridge. With five plates. 

25. A contribution towards our knowledge of the Pterylography of the Megapodii. By W. 

Pycraft, A.L.S., British Museum. With one plate. 

26. The Stolonifera and Alcyonacea. By Sydney J. HiCKSON, M.A., D.Sc, F.R.S., Beyer Pi ofes 

of Zoology in the Owens College, and ISA L. HiLES, B.Sc, Owens College, Manchester. 
With two plates. 

27. Report on the Xeniidae. By J. H. ASHWORTH, B.Sc, Demonstrator in Zoology, Owens ColU 

Manchester. With two plates. 

PART V. 1900. 

28. A Description of the Entozoa collected by Dr Willey during his sojourn in the West 

Pacific. By Arthur E. Shipley, M.A. With Plates LIV.— LVI. 

29. On some South Pacific Nemertines collected by Dr Willey. By R. C. PUNNETT. 

With Plates LVII.— LXI. 

30. On the Young of the Robber Crab. By L. A. BORRADAILE, M.A. With figures in the text. 

31. Anatomy of Neohelia porcellana (Moseley). By Edith M. Pratt, M.Sc. 

With Plates LXI I. and LXI 1 1. 

32. On a new Blind Snake from Lifu, Loyalty Islands. By G. A. BouLENGER, F.R.S. 

With figures in the text. 

33. On Crustacea brought by Dr Willey from the South Seas. By the Rev. T. R. R. StebbinG, F.F 

With Plates LXIV.— LXXIV. 

PART VI. 1902. 

34. Contribution to the Natural History of the Pearly Nautilus. By Arthur Willey, D.Sc, F.E 

I. Personal Narrative. II. Special Contribution. 
With Plates LXXV.— LXXXIII., a map and fifteen text-figures. 

CAMBRIDGE UNIVERSITY PRESS 
''^^OA 7@ aonbon: FETTER LANE, E.G. 

C. F. CLAY, Manager 



y 



w 



SMITHSONIAN INSTITUTION LIBRARIES 




3 'IDflfi DD^^5DSfl 3 

nhent qQL345 H3F2 
V 2 Fauna hawaiiensis,