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GRASSLAND 
RANGES 

in  the  southern  interior 
of  British  Columbia 


CANADA  DEPARTMENT  OF  AGRICULTURE      PUBLICATION  1319     1968 


p.  /  $  a 
/tot 


JUL    25  1SW     s 


GRASSLAND  RANGES 

In  the  Southern  Interior  of  British  Columbia 

ALASTAIR  MCLEAN  AND  LEONARD  MARCHAND      k       * 
Research  Station,  Kamloops,  B.C. 

GUIDE  TO  TYPES  AND  CONDITIONS  OF  RANGES 

A  successful  range  manager  should 

•  be  able  to  recognize  the  types  of  forage  on  his  range ; 

•  know  something  of  the  quality  and  quantity  of  forage  each  of  his  range 
types  is  capable  of  producing; 

•  be  able  to  appraise  the  condition  of  the  range  in  relation  to  its  potential. 
A  range  site,  in  terms  of  range  management,  is  an  area  of  rangeland  that  has 

a  certain  potential  for  growing  forage.  The  potential  depends  on  the  soil, 
climate,  and  slope,  and  is  achieved  by  the  climax  plant  community  characteristic 
of  the  site. 

Range  condition  is  the  state  or  health  of  a  range  in  relation  to  its  potential. 
It  depends  on  the  responses  of  forage  cover  and  soil  to  grazing  practices.  It  is 
determined  by  observing  the  species,  the  relative  abundance  and  vigor  of  the 
plants,  and  the  amount  of  litter  or  mulch  on  the  ground,  and  the  degree  of 
erosion.  In  British  Columbia  the  condition  is  a  measure  of  forage  production 
in  relation  to  the  potential  under  ungrazed  or  climax  vegetation  or  ideal  manage- 
ment. 

A  climax  plant  community  is  one  in  which  the  plant  species  have  reached  a 
state  of  equilibrium  with  the  climate  and  soil. 

Plant  succession  is  the  replacement  of  one  plant  species  or  group  of  species 
by  others.  The  changes  may  result  from  disturbances  caused  by  animals  or  man, 
changes  in  the  climate  or  soil,  or  changes  wrought  by  the  plants  themselves. 
Plant  succession  may  be  detrimental  or  beneficial,  depending  on  the  land  use. 
There  are  many  stages  of  succession  and  these  may  vary  depending  on  what  is 
causing  the  change. 

Range  plants  can  be  classified  according  to  their  response  to  grazing.  The 
classes,  as  defined  by  the  United  States  Department  of  Agriculture  (Agricultural 
Handbook  235),  are  decreasers,  increasers,  and  invaders.  In  rating  ranges  as  to 
condition  the  following  classes  are  standard:  excellent,  good,  fair,  and  poor. 
These  classes  reflect  the  relative  amounts  of  desirable  or  climax  forage  plants 
(decreasers)  compared  with  less  desirable  species  (increasers)  and  weeds  (in- 
vaders), mostly  annuals. 

Decreasers  are  species  that  decrease  under  prolonged,  excessive  grazing. 
Usually  they  are  perennials  that  produce  much  high-quality  forage,  and  dominate 
the  climax  community.  In  determining  the  range  condition  class,  the  total  of  all 
such  species  present  is  tallied. 


Increasers  are  species  that  increase  with  moderate  overuse;  they  decrease 
with  continued  heavy  overuse.  Increasers  commonly  are  the  shorter,  less  pro- 
ductive, subdominant  species  in  the  climax  plant  community.  Their  forage  value 
ranges  from  low  to  high.  Under  grazing,  those  of  low  forage  value  tend  to 
increase  more  rapidly  than  those  of  high  value. 

Invaders  are  species  that  invade  the  plant  community  as  a  range  deteriorates. 
They  are  not  members  of  the  climax  plant  community  for  the  range  site  since 
they  cannot  withstand  the  competition  for  moisture,  nutrients,  and  light  in  the 
climax  vegetation.  They  come  in  and  grow  along  with  the  increasers  after  the 
preferred  species  have  been  destroyed  by  grazing.  Invaders  need  not  be  exotics 
since  they  may  be  members  of  the  climax  plant  communities  on  other  range 
sites  in  the  same  locality.  Some  are  annual  weeds  and  herbaceous  perennials 
and  some  are  woody  shrubs;  many  have  some  grazing  value  but  others  have 
little. 

Excellent  range  has  at  least  75  percent  of  the  forage  made  up  of  decreasers. 
The  plants  are  vigorous,  have  good  root  systems,  and  set  a  good  crop  of  viable 
seed  in  most  years.  There  is  enough  mulch  on  the  ground  to  prevent  erosion, 
conserve  soil  moisture,  and  prevent  winter-killing  of  plants.  The  total  amount  of 
mulch  varies  considerably  with  the  site,  but  the  relative  amount  for  each  class 
generally  does  not.  Rain  soaks  in  rapidly.  Erosion  is  not  useful  as  a  criterion  of 
excellent  range  on  nearly  flat  land  or  on  soils  that  resist  erosion. 

Good  range  has  50  to  75  percent  of  the  forage  made  up  of  decreasers.  The 
plants  are  vigorous  and  set  an  adequate  crop  of  viable  seed  in  most  years.  There 
is  enough  mulch  to  allow  little  erosion.  Rain  soaks  in  fairly  rapidly. 

Fair  range  has  from  25  to  50  percent  of  the  forage  made  up  of  decreasers, 
the  rest  made  up  of  increasers,  many  of  which  are  unpalatable.  Some  invaders 
are  usually  present.  The  decreaser  plants  lack  vigor,  have  some  killing  at  the 
crowns,  and  set  adequate  seed  crops  in  fewer  than  half  the  years.  There  is  not 
usually  enough  mulch  to  prevent  erosion.  The  amount  of  mulch  is  not  a  useful 
criterion  of  fair  range  if  the  range  has  had  good  management  for  a  few  years 
but  not  long  enough  to  change  the  plant  species  present.  Rainfall  soaks  in 
poorly,  causing  considerable  runoff  and  some  sheet  erosion. 

Poor  range  has  less  than  25  percent  of  its  forage  made  up  of  decreasers,  and 
those  present  are  generally  in  a  weakened  condition.  Invaders  and  unpalatable 
increasers  are  abundant.  Very  little  mulch  is  present,  water  runoff  is  frequent, 
and  the  topsoil  is  generally  compacted.  Sheet  and  rill  erosion  is  usually  marked. 

Stocking  rates  in  this  publication  have  been  estimated  on  the  basis  of  acres 
per  animal-unit  month  (AUM),  this  being  the  number  of  acres  needed  to  graze 
a  1000-pound  cow  for  1  month.  They  cannot  be  determined  directly  from  the 
estimated  total  forage  yields.  The  amount  of  available  forage  has  to  be  calculated, 
a  45  percent  carryover  being  allowed  to  avoid  damage  to  the  range.  On  the  aver- 
age, about  660  pounds  of  available  forage  is  needed  per  AUM.  Other  factors 
such  as  stocking  history,  livestock  grazing  habits,  season  of  grazing,  kind  of 


livestock,  distance  of  water,  steepness  of  slope,  rockiness,  and  palatability  or 
nutritive  value  of  the  forage  must  also  be  considered. 

The  following  are  commonly  accepted  animal-unit  equivalents:  yearling, 
0.6;  bull,  1.3;  horse,  1.5;  ewe,  0.2. 

Range  trend,  that  is,  whether  the  range  is  improving  or  deteriorating,  should 
be  determined  along  with  range  condition.  The  following  criteria  should  be 
used :  the  degree  of  use,  or  percentage  carryover  of  forage ;  the  plant  species  that 
are  reproducing  and  becoming  established;  the  relative  vigor  of  desirable  and 
undesirable  species;  and  erosion  signs,  healing  over  of  gullies,  evidence  of  soil 
puddling,  and  accumulation  of  mulch. 

The  botanical  and  common  names  of  plants  are  listed  in  Tables  1,  2,  3,  and  4. 

Caution:  The  classifications  listed  in  this  publication  are  to  some  degree  an 
oversimplification  of  the  plant  communities  as  you  will  find  them.  For  example, 
you  will  find  many  situations  that  are  intermediate  between  the  types  and  classes 
recorded.  Other  sites,  because  of  a  peculiar  climate  or  a  soil  that  is  especially 
shallow,  sandy,  salty,  or  rocky,  will  not  conform  to  the  pattern  or  will  be  unable 
to  reach  the  potential  indicated.  There  is  no  substitution  for  careful  observation 
and  sound  reasoning  and  judgment  in  the  application  of  the  principles  given 
in  this  publication. 

Big  Sagebrush  —  Bluebunch  Wheatgrass  Site 

ELEVATION 

1100  to  2000  feet  near  Kamloops;  often  different  in  other  areas. 

PRECIPITATION 

Nine  to  10  inches  per  year,  slightly  more  than  half  falling  from  April  to 
the  end  of  October. 

TEMPERATURE 

From  a  high  monthly  mean  of  73°F  (July)  to  a  low  mean  of  23°F  (January); 
yearly  mean  is  48°F. 

The  summers  are  hot  and  dry  with  occasional  storms.  The  winters  do  not 
usually  have  prolonged  cold  spells;  snowfall  is  rather  light,  and  occasional 
chinook  winds  bare  the  ground  and  may  be  followed  by  drying  winds.  March 
and  April  are  usually  dry  and  windy. 

TOPOGRAPHY 

From  nearly  level  to  rolling  and  very  steep  with  small  benches  and  deep 
gullies.  Most  of  these  grasslands  are  on  outwash  plains  and  terraces  on  the 
drier  valley  slopes. 

SOILS 

Brown,  medium  textured,  neutral  to  slightly  saline.  Lime  is  generally  found 
at  about  1  foot. 


GRAZING  USE 

Early  spring,  late  fall,  and  winter  range. 

GROWING  SEASON 

Mid-March  to  end  of  June.  In  most  years  there  is  only  a  small  amount  of 
regrowth  in  the  fall. 

NATIVE  VEGETATION 

Decreasers  Increasers  Invaders 

Bluebunch  wheatgrass  Big  sagebrush  Tansy  mustard 

Needle-and-thread  Dwarf  pussytoes  Downy  brome 

(some  soils)  Needle-and-thread  Russian  thistle 

Rabbitbrush 

Sand  dropseed  (sandy  soils) 

Sandberg's  bluegrass 

These  grasslands  are  distinguished  by  the  presence  of  big  sagebrush,  well- 
spaced  bunchgrasses  with  bare  ground  between  plants,  and  relatively  few  plant 
species.  With  overgrazing,  bluebunch  wheatgrass  is  replaced  by  Sandberg's 
bluegrass  and  big  sagebrush. 

Under  prolonged  misuse,  annuals  become  abundant.  On  some  dry  sites, 
needle-and-thread  may  be  an  important  increaser  until  grazing  becomes  severe 
and  it  is  forced  out.  It  occurs  in  varying  proportions  with  bluebunch  wheatgrass. 

Needle-and-thread  forms  a  climax  and  with  big  sagebrush  becomes  a  de- 
creaser  on  some  coarse-textured  soils.  On  the  basis  of  forage  yield  and  plant 
composition,  however,  such  sites  still  fit  into  the  condition  classes  as  established. 
Range  dominated  by  needle-and-thread  is  not  considered  excellent  range  under 
any  conditions;  this  grass  yields  less  than  bluebunch  wheatgrass  and  may  injure 
the  mouths  of  grazing  animals  because  of  the  barbed  seeds  and  long  awns  if  it  is 
grazed  after  seed  is  formed  and  before  it  is  shed.  As  big  sagebrush  —  needle- 
and-thread  range  deteriorates,  sand  dropseed  and  big  sagebrush  increase  at  the 
expense  of  the  needle-and-thread.  Since  the  needle-and-thread  climax  sites  are 
usually  inherently  less  productive  than  the  bluebunch  wheatgrass  sites,  the 
amount  of  big  sagebrush  present  is  usually  less  for  the  same  condition  class. 

CONDITION  CLASSES 

The  percentages  of  ground  cover  for  the  main  plants  by  classes  are: 


Plants 

Excellent 

Good 

Fair 

Poor 

Decreasers 

60 

60-40 

40-20 

20-0 

Increasers 

Big  sagebrush 

0-15 

15-30 

30-45 

45-60 

Total 

0-30 

30-50 

50-70 

70 

(Amounts  of  big  sagebrush  are  usually  about  10  percent  less  for  the  same 
condition  class  on  needle-and-thread  sites) 


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FORAGE  YIELDS 

The  yields  (pounds  per  acre)  and  average  stocking  rates  (acres  per  AUM) 
for  the  four  classes  of  condition  are: 


Excellent 

Good 

Fair 

Poor 

Yield,  average 

550 

400 

250 

100 

Yield,  range 

900-400 

700-250 

400-150 

250-50 

Percentage  of  potential 

100 

73 

46 

18 

Stocking  rate 

2.2 

3.0 

4.8 

12.0 

Bluebunch  Wheatgrass  —  Sandberg's  Bluegrass  Site 

ELEVATION 

2000  to  2600  feet  near  Kamloops ;  often  different  in  other  areas. 

PRECIPITATION 

Ten  to  11.5  inches  per  year,  slightly  more  than  half  falling  from  April  to 
the  end  of  October. 

TEMPERATURE 

From  a  high  monthly  mean  of  70°F  (July)  to  a  low  mean  of  22°F  (January); 
yearly  mean  is  47°F. 

As  the  elevation  increases,  mean  temperatures  and  evaporation  decrease 
and  moisture  effectiveness  increases.  Winter  cold  spells  are  more  prolonged  and 
drying  winter  winds  more  common  than  in  the  big  sagebrush  —  bluebunch 
wheatgrass  grasslands. 

TOPOGRAPHY 

From  nearly  level  to  rolling  and  steep,  often  with  hummocky  microtopog- 
raphy. 

SOILS 

Dark  Brown,  medium  light  textured,  neutral  to  slightly  saline.  Lime  is 
generally  found  at  about  1.5  feet. 

GRAZING  USE 

Spring  and  fall  (early  May  and  October)  range. 

GROWING  SEASON 

Late  March  to  early  July.  In  most  years  there  is  some  fall  regrowth. 

CONDITION  CLASSES 

The  percentages  of  ground  cover  for  the  main  plants  by  classes  are: 
Plants  Excellent  Good  Fair  Poor 

Decreasers  60  60-40  40-20  20-0 

Increasers  0-30  30-50  50-70  70 


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FORAGE  YIELDS 

The  yields  (pounds  per  acre)  and  average  stocking  rates  (acres  per  AUM) 
for  the  four  classes  of  condition  are: 

Excellent  Good  Fair  Poor 

Yield,  average  500  420  325  250 

Stocking  rate  2.4  2.9  3.7  4.8 


NATIVE  VEGETATION 

Decreasers 

Bluebunch  wheatgrass 
Needle-and-thread 
(some  soils) 


Invaders 
Downy  brome 
Dandelion 
Sixweeks  fescue 
Woolly  plantain 


Increasers 

Dwarf  pussytoes 

Junegrass 

Needle-and-thread 

Pasture  sage 

Rabbitbrush 

Sand  dropseed  (sandy  soils) 

Sandberg's  bluegrass 

The  absence  of  big  sagebrush  and  closer  spacing  of  the  plants  distinguish 
this  site  from  the  big  sagebrush— bluebunch  wheatgrass  site;  the  dominance  of 
bunchgrasses  as  compared  with  forbs,  the  absence  of  aspen  clumps,  and  the 
presence  of  only  a  few  species  distinguish  it  from  the  bluebunch  wheatgrass  — 
rough  fescue  site. 

Under  misuse,  needle-and-thread  usually  becomes  dominant  but  is  replaced 
in  turn  by  downy  brome  and  low-growing  weeds.  Needle-and-thread  may  form 
a  climax  and  become  a  decreaser  on  certain  coarse-textured  soils.  However, 
on  the  basis  of  forage  yield  and  species  composition  such  sites  fit  into  the  con- 
dition classes  as  established.  Range  dominated  by  needle-and-thread  is  not 
usually  considered  excellent  range;  this  grass  yields  less  than  bluebunch  wheat- 
grass  and  may  injure  the  mouths  of  grazing  animals  because  of  the  barbed 
seeds  and  long  awns  if  it  is  grazed  after  seed  is  formed  and  before  it  is  shed. 


Bluebunch  Wheatgrass  —  Rough  Fescue  Site 

ELEVATION 

2600  to  3000  feet  near  Kamloops;  often  different  in  other  areas. 

PRECIPITATION 

Eleven  to  13  inches  per  year,  slightly  more  than  half  falling  from  April  to 
the  end  of  October. 

TEMPERATURE 

From  a  high  monthly  mean  of  66°F  (July)  to  a  low  mean  of  20°F  (January); 
yearly  mean  is  45°F.  As  the  elevation  increases,  mean  temperatures  and  evapora- 


10 


tion  decrease  and  moisture  effectiveness  increases.  Winter  cold  spells  are  more 
prolonged  and  drying  winds  less  common  than  in  the  bluebunch  wheatgrass  — 
Sandberg's  bluegrass  site.  The  ground  is  seldom  bared  of  snow  during  the  winter. 

TOPOGRAPHY 

From  nearly  level  to  rolling,  sometimes  steep. 

SOILS 

Dark  Brown  to  Black,  usually  neutral  in  reaction  and  medium  textured. 
Lime  is  generally  found  at  about  2  feet. 

GRAZING  USE 

Spring  and  fall  (late  May,  June,  and  October)  range. 

GROWING  SEASON 

First  week  in  April  to  mid-July.  In  most  years  there  is  also  considerable 
regrowth  in  September. 

CONDITION  CLASSES 

The  percentages  of  ground  cover  for  the  main  plants  by  classes  are: 
Plants  Excellent  Good  Fair  Poor 

Decreasers  60  60-40  40-20  20-0 

Increasers  0-30  30-50  50-70  70 

FORAGE  YIELDS 

The  yields  (pounds  per  acre)  and  average  stocking  rates  (acres  per  AUM) 
for  the  four  classes  of  condition  are: 

Excellent  Good  Fair  Poor 

Yield,  average                         1000  800  700  500 

Yield,  range                        426-2278  412-1866  420-1339  368-854 

Percentage  of  potential           100  80  70  50 

Stocking  rate                               1.2  1.5  1.7  2.2 

NATIVE  VEGETATION 


Decreasers 

Bluebunch  wheatgrass 
Idaho  fescue 
Rough  fescue 


Invaders 
Downy  brome 
Dandelion 
Common  mullein 
Compound 
fleabane 


Increasers 

Columbia  needlegrass 
Junegrass 

Kentucky  bluegrass 
Pasture  sage 
Pussytoes 
Sandberg's  bluegrass 
Silky  lupine 
Timber  milk-vetch 
Western  yarrow 

This  site  is  distinguished  by  more  species  and  greater  abundance  of  forbs, 
increased  height  growth  of  grasses,  and  more  dense  plant  cover  than  the  other 


11 


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two  zones.  Clumps  of  aspen  occur  frequently  in  draws  and  swales.  With  over- 
grazing the  decreasers  are  replaced  normally  by  Sandberg's  bluegrass  and 
weedy  species  such  as  compound  fleabane,  pasture  sage,  pussytoes,  western 
yarrow,  dandelion,  and  common  mullein.  Downy  brome  and  other  annuals 
increase  greatly  on  range  in  poor  condition.  The  zone  lies  adjacent  to  the  Douglas 
fir  forest.  On  certain  range  sites  in  fair  condition  needle-and-thread  and  Co- 
lumbia needlegrass  increase  but  decrease  again  with  prolonged  misuse. 

Idaho  fescue  replaces  rough  fescue  to  a  great  extent  in  the  Similkameen  and 
Kettle  valleys  and  southern  part  of  the  Rocky  Mountain  Trench.  The  fescues 
generally  increase  on  the  more  moist  upland  sites  within  the  zone  as  compared 
with  bluebunch  wheatgrass,  and  are  confined  largely  to  the  north-  and  east- 
facing  slopes  in  the  lower  parts  of  the  zone.  Kentucky  bluegrass  generally 
dominates  on  deep,  fine-textured  soils  and  swales  but  frequently  increases  in 
the  bluebunch  wheatgrass  —  rough  fescue  community  as  the  latter  deteriorates 
with  grazing  pressure. 

Ponderosa  Pine  Site 

ELEVATION 

2000  to  3000  feet  near  Kamloops;  often  different  in  other  areas. 

PRECIPITATION 

Eleven  to  13  inches,  slightly  over  half  falling  from  April  to  the  end  of 
October. 

TEMPERATURE 

Approximately  the  same  climatic  regime  as  the  middle  grasslands  and  lower 
portion  of  the  bluebunch  wheatgrass  —  rough  fescue  grasslands. 

TOPOGRAPHY 

From  nearly  level  to  rolling,  sometimes  steep. 
SOILS 

Brown  Wooded  or  Dark  Gray,  neutral  in  reaction,  and  generally  light 
textured.  Often  on  duny  sands  and  outwash  gravel  terraces. 

GRAZING  USE 

Spring  and  fall  (May,  June,  and  October). 

GROWING  SEASON 

Late  March  to  end  of  June.  In  most  years  there  is  considerable  regrowth 
in  September. 

CONDITION  CLASSES 

The  percentages  of  ground  cover  for  the  main  plants  by  classes  are: 
Plants  Excellent  Good  Fair  Poor 

Decreasers  60  60-40  40-20  20-0 

Increasers  0-30  30-50  50-70  70 

(Text  Continued  on  Page  24)  13 


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Big  sagebrush  -  bluebunch  wheatgrass  condition  classes.  Top,  excellent.  Bottom,  good. 


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Big  sagebrush  -  bluebunch  wheatgrass  condition  classes.  Top,  fair.  Bottom,  poor. 


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Bluebunch  whcatgrass  -  Sandberg's  bluegrass  condition  classes. 

Top,  excellent.  Bottom,  good. 


17 


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Bluebunch  wheatgrass  -  Sandberg's  bluegrass  condition  classes. 
Top,  fair.  Bottom,  poor. 


18 


Bluebunch  wheatgrass  -  rough  fescue  condition  classes.  Top,  excellent.  Bottom,  good. 


19 


Bluebunch  wheatgrass  -  rough  fescue  condition  classes.  Top,  fair. 
Bottom,  fair  (needle-and-thread). 


20 


Top,  bluebunch  wheatgrass  -  rough  fescue  condition  class,  poor. 
Bottom,  ponderosa  pine  condition  class,  excellent. 


21 


Ponderosa  pine  condition  classes.  Top,  good.  Bottom,  fair. 


22 


Ponderosa  pine  condition  classes.  Top,  poor.  Bottom,  poor  (big  sagebrush). 


23 


NATIVE  VEGETATION 


Decreasers 

Bluebunch  wheatgrass 
Idaho  fescue 
Needle-and-thread 

(some  soils) 
Rough  fescue 


Increasers  Invaders 

Balsamroot  Annual  phlox 

Big  sagebrush  (some  soils)  Downy  brome 

Dwarf  pussytoes  Dandelion 

Junegrass  Woolly  plantain 

Needle-and-thread 
Rabbitbrush 
Sandberg's  bluegrass 
Silky  lupine 
Western  yarrow 
In  many  cases  this  site  parallels  the  bluebunch  wheatgrass  —  Sandberg's 
bluegrass  and  bluebunch  wheatgrass  —  rough  fescue  grasslands.  It  is  character- 
ized by  the  presence  of  ponderosa  pine  in  open  savannalike  stands.  Hawks- 
beard,  rabbit  brush,  and  balsamroot  are  more  common    here   than   in   the 
other  sites.  The  community  occurs,  most  commonly,  on  the  lower  valley  slopes. 
The  productivity  of  this  site  is  more  variable  than  that  of  the  other  sites 
since  the  yields  in  each  condition  class  vary  greatly  depending  on  the  density 
of  the  tree  cover.  Reestablishment  of  trees  is  usually  slow  after  removal  because 
of  the  relatively  dry  soil  conditions.  This  zone  needs  more  study  before  reliable 
estimates  of  productivity  can  be  made.  The  range  of  forage  yields  between 
condition  classes  overlapped  considerably  depending  on  plants  and  tree-canopy 
density.  The  maximum  range  of  all  sites  was  184  to  890  pounds  per  acre;  the 
average  yield  was  428  pounds.  The  average  stocking  rate  would  therefore  be 
about  3  acres  per  AUM. 


MANAGING  YOUR  RANGES 


You  can  produce  more  meat  per  acre  and  make  more  profit  per  ranch  by 
using  your  grassland  range  in  the  right  season  and  to  a  safe  degree.  To  do  this, 
however,  you  must  know  the  growth  characteristics  and  growth  requirements 
of  the  main  forage  plants.  This  information  will  help  you  to  graze  your  range 
so  that  forage  production  will  be  maintained. 

PLANT  GROWTH  CURVE 

To  illustrate  the  principles  involved  in  grass  growth  the  familiar  growth 
curve  is  used.  This  curve  shows  the  rate  of  growth  of  the  grass  plant  during 
the  growing  season.  It  is  characterized  by  slow  development  in  the  spring, 
followed  by  a  period  of  rapidly  increasing  growth,  and  then  a  gradual  tapering 
off  in  the  early  summer. 

The  period  of  slow  development  in  the  spring  may  cover  several  weeks, 
depending  on  the  weather.  Because  the  soil  temperatures  are  too  cold  for  root 


24 


activity  at  the  beginning  of  this  period,  the  food  for  plant  growth  comes  from 
that  which  was  stored  in  the  roots  by  the  plant  the  previous  summer  and  fall. 

When  the  air  and  soil  temperatures  become  warmer,  nutrients  are  absorbed 
from  the  soil  and  transported  to  the  green  leaves  where,  in  the  warm  sunlight, 
they  are  made  into  food  for  plant  growth.  The  increase  of  leaf  growth  at  this 
stage  in  reality  increases  the  capacity  of  the  plant  to  produce  food.  This  is 
where  the  growth  curve  starts  to  rise  very  rapidly. 

The  rapid  rate  of  top  growth  continues  until  the  grass  begins  to  form  seed 
stalks.  It  then  begins  to  taper  off,  partly  because  of  the  increasing  demand  of 
seed  stalks,  flowers,  and  seed  for  food.  And,  as  this  demand  increases  and  the 
rate  of  top  growth  decreases,  the  curve  flattens  out.  The  onset  of  dry,  hot 
weather  also  slows  top  growth. 

Not  all  of  the  plant  food  manufactured  in  the  leaves  of  the  grass  goes 
toward  the  production  of  top  growth,  however.  About  the  time  the  growth 
curve  starts  its  rapid  rise,  the  grass  begins  to  build  up  its  store  of  reserve  plant 
food  in  the  roots  and  crowns  for  use  early  the  next  year.  Then  during  the  period 
of  rapid  top  growth  and  seed  production,  only  a  little  of  the  plant  food  goes 
back  into  storage.  During  and  after  seed  maturity,  the  storage  process  increases 
again  and  continues  until  growth  stops. 

In  addition  to  being  stored,  some  of  the  plant  food  returned  to  the  roots 
during  the  growing  season  each  year  goes  into  the  growth  of  new  grass  roots, 
which  replace  the  part  of  the  root  system  that  dies. 

As  well  as  producing  more  forage,  grass  with  adequate  stored  food  survives 
drought,  hard  winters,  and  insect  damage  better  than  starved  plants. 


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25 


GROWTH  CHARACTERISTICS  OF  GRASS  SPECIES 

Different  species  of  range  grasses  have  different  growth  habits,  season  of 
growth,  forage  yields,  chemical  compositions,  reaction  to  grazing,  and  other 
characteristics. 

The  different  grasses  have  different  periods  of  growth  and  best  seasons  of 
use.  For  example,  bluebunch  wheatgrass  starts  growth  early  in  the  spring, 
while  grasses  like  sand  dropseed  have  later  seasons  of  use. 

Bluebunch  wheatgrass  is  very  susceptible  to  grazing  damage  at  certain 
times.  Analyses  at  the  Research  Station  at  Kamloops  show  that  the  plant  has 
its  lowest  root  reserves  when  the  tops  are  about  7  inches  high.  From  then  on, 
the  reserves  are  built  up  to  provide  stored  food  to  start  the  next  year's  spring 
growth.  The  sugar  contents  in  the  roots  were  6  percent  at  2  inches,  2  percent  at  7 
inches,  8  percent  at  full  bloom,  and  12  percent  at  seed  shed. 

Bluebunch  wheatgrass  should  not  be  grazed  before  it  is  7  inches  high  if  the 
animals  are  to  be  left  on  until  the  grass  matures.  Grazing  may  be  started  when 
the  grass  is  only  4  inches  high,  but  the  animals  must  be  removed  when  the  plants 
have  put  on  no  more  than  half  their  season's  growth.  The  plants  will  then  have 
time  to  grow  and  store  food  in  their  roots  for  regrowth  in  the  fall  and  the  next 
spring. 

In  one  test  at  Kamloops  in  which  bluebunch  wheatgrass  was  clipped  at 
different  intensities  to  simulate  grazing,  all  spring  clippings  regardless  of  in- 
tensity caused  some  damage.  Plants  clipped  not  later  than  early  May  showed 
the  least  damage,  whereas  over  80  percent  of  the  plants  clipped  at  the  end  of 
May  were  killed  in  one  season.  The  grazing  heights  mentioned  apply  only  to 
bluebunch  wheatgrass;  most  other  species  react  differently. 

The  growth  habits  of  a  grass  species  are  important  in  determining  safe  degree 
of  use.  For  example,  bluebunch  wheatgrass  has  an  erect  growth  habit  whereas 
needle-and-thread  has  a  spreading  growth  habit.  As  a  result,  close  grazing 
removes  many  more  leaves  of  bluebunch  wheatgrass  than  needle-and-thread, 
and  fewer  green  leaves  are  left  to  continue  manufacturing  food.  Measurements 
from  ranges  near  Kamloops  show  that  clipping  bluebunch  wheatgrass  5  inches 
above  the  ground  when  fully  grown  removes  about  70  percent  of  the  plant, 
whereas  clipping  needle-and-thread  at  the  same  height  removes  only  about 
45  percent.  For  most  grasses  it  is  best  to  leave  at  least  half  of  the  total  season's 
growth  to  ensure  adequate  food  reserves  in  the  plant. 

RANGE  READINESS  AND  PROPER  USE 

For  proper  seasonal  use  delay  grazing  in  the  spring  until  about  the  time  the 
grass  begins  to  grow  rapidly.  At  this  stage  the  grass  is  no  longer  dependent 
mainly  on  the  limited  supply  of  plant  food  stored  in  the  roots.  It  is  now  able  to 
obtain  raw  materials  from  the  soil.  If  grazing  begins  at  an  earlier  date  and  the 
tops  are  kept  eaten  off,  the  natural  growth  cycle  of  the  grass  plant  is  delayed 
and  the  plant  may  never  be  able  to  produce  food  for  storage. 


26 


Grassland  range  in  the  dry  belt  is  ready  for  grazing  by  the  time  the  bluebunch 
wheatgrass  is  about  7  inches  high.  In  the  Kamloops  area  this  varies  from  about 
April  8  on  the  lower  ranges  (1100  to  1700  feet)  to  about  May  10  on  the  higher 
grasslands  (2300  to  2800  feet). 

By  the  time  the  bluebunch  wheatgrass  is  6  to  7  inches  tall,  rough  fescue  is 
at  the  same  height,  needle-and-thread  is  about  3  inches  high,  Sandberg's  blue- 
grass  and  balsamroot  are  starting  to  head  out,  and  the  Johnny-jump-up  flowers 
are  withered. 

Some  top  growth  should  be  left  at  the  end  of  the  grazing  period.  During 
the  growing  season  this  gives  the  plant  some  leaf  growth  to  continue  food 
manufacture  while  moisture  is  still  available  in  the  soil.  In  the  fall  old  leaves 
and  seed  stalks  that  remain  protect  the  soil  and  conserve  moisture.  The  amount 
of  ungrazed  material  required  to  accomplish  this  varies  with  such  factors  as 
type  of  soil  and  rainfall  pattern. 

To  repeatedly  violate  these  requirements  of  grass  growth  by  grazing  too 
early  and  keeping  the  grass  grazed  down  too  closely  during  the  growing  season 
will  result  in  a  weakening  of  the  plants  and  deterioration  of  site  condition  and 
forage  production. 

RANGE  CONDITION 

When  these  principles  are  ignored,  the  good  forage  species  such  as  blue- 
bunch wheatgrass  and  rough  fescue  disappear  and  are  replaced  by  other  plants 
such  as  Sandberg's  bluegrass,  downy  brome,  pasture  sage,  big  sagebrush,  and 
pussytoes.  The  range  then  goes  down  in  condition. 

Fence-line  differences  often  show  these  changes  dramatically.  One  site 
studied  contained  50  percent  wheatgrass  and  7  percent  big  sagebrush  and  yielded 
1500  pounds  of  grass  per  acre,  whereas  the  adjacent  grazed  range  had  5  percent 
wheatgrass  and  35  percent  big  sagebrush  and  yielded  only  140  pounds  per  acre. 
Another  area  that  has  had  only  incidental  grazing  at  any  time  had  70  percent 
bluebunch  wheatgrass  and  13  percent  big  sagebrush  and  produced  700  pounds 
of  feed  per  acre.  On  the  adjacent  heavily  grazed  side  of  the  fence  there  was  only 
7  percent  bluebunch  wheatgrass  but  25  percent  big  sagebrush,  which  yielded 
only  90  pounds  per  acre.  A  site  that  may  never  have  been  grazed  by  stock 
contained  80  percent  bluebunch  wheatgrass  and  practically  no  cactus,  big 
sagebrush,  or  downy  brome;  it  yielded  nearly  550  pounds  of  grass  per  acre. 
But  across  the  fence  on  an  abandoned  field  there  was  6  percent  bluebunch 
wheatgrass,  47  percent  downy  brome,  22  percent  cactus,  and  1 1  percent  sage- 
brush, which  produced  only  50  pounds  of  grass  per  acre. 

RANGE  RECOVERY  TIME 

Some  areas  recover  quickly  from  overgrazing  while  others  may  take  genera- 
tions. One  area  fenced  in  1923  near  Riske  Creek  has  shown  a  recovery  of  blue- 
bunch wheatgrass  from  4  percent  to  13  percent,  and  a  decrease  in  pasture  sage 


27 


from  36  percent  to  13  percent  with  relatively  little  improvement  in  yield. 
Another  area  near  Tranquille  fenced  in  1936  has  shown  a  recovery  of  blue- 
bunch  wheatgrass  from  28  percent  to  58  percent  and  a  decrease  in  needle-and- 
thread  from  17  percent  to  1  percent.  The  average  yield  has  increased  from 
164  pounds  of  forage  per  acre  to  360  pounds.  A  third  area  on  the  Skookum- 
chuck  prairie  fenced  in  1952  has  shown  an  increase  in  bluebunch  wheatgrass 
from  7  percent  to  25  percent  and  a  decrease  in  pussytoes  from  26  percent  to 
10  percent. 

The  feed  value  of  range  forage  is  an  important  consideration  in  season  of 
use.  For  example,  it  is  sometimes  possible  to  graze  a  range  high  in  broad- 
leaved  plants  early  in  the  season  to  take  advantage  of  the  feed  produced  by 
these  species  before  they  dry  up  and  disappear.  More  information  is  given  in 
the  publications  Chemical  Composition  of  Native  Forage  Plants  in  British 
Columbia  in  Relation  to  Grazing  Practices,  and  Handbook  on  Grazing  Values 
of  Range  Plants  of  British  Columbia,  available  from  the  Research  Station  at 
Kamloops. 

POINTS  TO  REMEMBER 

For  proper  range  management 

•  graze  moderately  during  the  growing  season,  leaving  enough  top  growth 
for  manufacture  and  storage  of  food,  and  protection  for  the  plant  crowns ; 

•  rest  the  range  after  close  grazing  at  any  time  in  the  growing  season  to 
allow  growth  to  resume  and  plant  food  to  be  stored  in  the  roots. 


28 


.■If, 


Copies  of  this  publication  may  be  obtained  from: 
INFORMATION  DIVISION 

CANADA  DEPARTMENT  OF  AGRICULTURE 

OTTAWA 


ROGER  DUHAMEL.  F.R.S.C.  QUEEN'S   PRINTER    AND  CONTROLLER  OF  STATIONERY,  OTTAWA,  1968 

6M  —33964—    6:58  Cat.  No.:  A53-1319