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A HAND-BOOK 


TO THE 


eeeorA OF CEYLON. 


A HAND-BOOK 


TO THE 


mEORA OF CEYLON 


CONTAINING 
DESCRIPTIONS OF ALL THE SPECIES OF FLOWERING PLANTS 
INDIGENOUS TO THE ISLAND, 


AND 


NOTES ON THEIR HISTORY, DISTRIBUTION, AND USES. 


BY 


mewn Y FTRIMEN, M.B. (Lonp.), F.R.S., 


DIRECTOR OF THE ROYAL BOTANIC GARDENS, CEYLON. 


Wiitb an Ztlas of Plates 
ILLUSTRATING SOME OF THE MORE INTERESTING SPECIES. 


Part: i, 
CONNARACEA—RUBIACE#. 
LIBRARY 
WITH PLATES XXVI-L. NEW YORK 
AND INDEX TO PARTS I, AND U1. BOTANICAL 
GARDEN 
PUBLISHED UNDER THE AUTHORITY OF THE GOVERNMENT OF CEYLON, 


LONDON : 
DULAU & CO., 37 Sono Square, W. 
1594. 


LONDON: 
PRINTED BY STRANGEWAYS AND SONS, 
Tower Street, Cambridge Circus, W.C. 


LIBRARY 
NEW YORK 
BOTANICAL 

GARDEN 


HANDBOOK TO THE FLORA OF CEYLON. 


T has been found desirable to make an alteration in the 
mode of publication of the text of this book. It was my 
ntention that the whole should have formed Two Volumes 
(each of about 800 pages), the first Two Parts together making 
up Vol. I.; but, owing to the thick paper used, I find that such 
volumes would be too bulky and heavy for convenient use. 

Part II. ts therefore now tssued as a volume of itself, with 
Sresh pagination; and the subsequent Parts will be treated in 
the same way. 

An Index to the contents of Parts I, and II. is, however, 


now published with the present one. 
is yall 


MAY 1 - 1905 


CORRECTIONS IN PART I. 


Page 7, line 14, for ‘ 7hwazteriz’ read Thwattesit. 


12, 
21, 
31, 


33; 


15, for ‘ Kurawita’ vead Kuruwita. 

17 from bottom, for ‘Ua’ read Unona. 

11 from bottom, for ‘ Karanita’ vead Karawita. 

7, for ‘ Sagenea’ read Sagerea. 
2 from bottom, for ‘tomatose’ vead tomentose. 
8, for ‘ supra-tomatose’ vead rufous-tomentose. 
10, for ‘2720’ read 2, 720. 
13 from bottom, for ‘ 2000’ vead 4000. 
5, for ‘ neelghenense’ read neelgherrense. 
3, for ‘ Pedum’ zead Peduru. 
9, for ‘ Mont.’ vead Mant. 
II, after ‘ Kokatiya’ add, S. 
19 from bottom, for ‘and’ vead And. 
6, after ‘imbricate’ add Cotyledons equal. 
7, after ‘imbricate’ add Cotyledons very unequal. 
8, after ‘reflexed’ add Cotyledons unequal. 

10, omit ‘near Pelmadulla (F. Lewis).’ 

16, 17, for ‘paler beneath and pellucid-punctate with trans- 
mitted light,’ ead veinlets minutely reticulate, pellu- 
cid, with free dilated ends. 

20 from bottom, before ‘montane’ zzser¢ upper. 

omit line 9. 

17, for ‘ Macdowell’ read Macdowall. 

11, for ‘are’ read one. 

8, for ‘leaf’ read vein. 
5 from bottom, for ‘ Devadaram’ read Tévataram. 

11, for ‘aukenda’ read ankenda. 

3, after ‘seeded’ add red. 
12, for ‘ Hanasgiriya’ read Hunasgiriya. 
3, omit * Balu-nakuta, S.’ 

16, for ‘McDowall’ read Macdowall. 

21 from bottom, for ‘ERIOCLADA’ vead INDICA. 

12 from bottom, for ‘ Rata’ read Ratu. 

1, for ‘ Schmedelia’ read Schmidelia. 
3 from bottom, for ‘ Kadaganuwa’ read Kadaganawa. 


XLITI.—CONNARACEE. 


SHRUBS or trees often scandent; |. imparipinnate (in E/i- 
panthus apparently simple), without stip.; fl. regular, bisexual, 
small; cal. deeply 5-fid., segm. equal, imbricate or valvate ; 
pet. 5, equal; stam. 5 or 10 (5 shorter), inserted at base of 
or on cal.-tube, fil. slightly connate at base; carp. 5 or 
solitary, with 2 collateral ovules in each; fruit a solitary 
{rarely 2) follicle (rarely 2-valved) ; seed large, solitary, erect, 
surrounded by a large or small aril, without endosperm. 


L. pinnate ; stam. Io. 


Stam. all perfect . : : 5 : ; . I. ROUREA. 
Five stam. with abortive anth. F : : . 2. CONNARUS. 
L. 1-foliolate ; stam. 5 : : : ; : . 3. ELLIPANTHUS, 


Of our four species, Comnarus monocarpus occurs chiefly in the dry 
region; the others only in the wet, C. Championiit and Ellipanthus (both 
endemic) extending into the lower montane zone. 


I ROUREA, Audi. 

A small, semi-scandent tree, 1. imparipinnate, fl. small, 
regular; cal. very deeply 5-fid, segm. strongly imbricate, 
much enlarged in fruit; pet. 5 ; stam. 10, inserted on base of 
cal., fil. connate at base, the 5 opp. the pet. shorter than the 
other 5; carp. 5, distinct, styles slender; fruit a solitary 
curved follicle (rarely two) ; seed solitary, erect, with a small 
basal aril, no endosperm.—Sp 50; 12 in FZ. B. /nd. 


R. santaloides, |. & A. Prod. 144 (1834). Mirindi-wel, S. 

Herm. Mus. to. Fl. Zeyl. n. 408 (Santaloides). dgiceras minus, Gaertn. 
Fruct., i. 216. Connarus santalotdes, Vahl Symb. ili. 87. Moon Cat. 49. 
Thw. Enum. 80. C. P. 749. 

Fl. B. Ind. ii. 47. Gaertn. Fruct. i. t. 46 (fruit only). Bedd. Fl. Sylv. 
t. 11 (fl. only). 

A scrambling shrub or small tree, branchlets slender, bark 
shining, purplish, buds pubescent; 1.-rachis 3-6 in., slender, 
curved, glabrous, Ifits. 5 or 7 (2 or 3 pairs and a terminal one), 
very shortly stalked, 2-3 in., oval or lanceolate, rounded at 
base, caudate- acuminate, glabrous, shining above, with pro- 
minent reticulate venation beneath ; fl. small, } in., on slender, 
jointed ped., arranged in lax, slender, erect or ascending, 
glabrous, racernose panicles, several of which arise from leaf- 
axil, bracts minute, hairy ; cal.-segm. orbicular, very obtuse, 

PART I. I 


2 Connaracee. [ Connarus. 


glabrous, very strongly imbricate ; pet. much exceeding cal., 
oval-oblong, spreading ; stam. erect, styles spreading ; follicle 
# in. or a little more, cylindrical, falcately curved, tapering to 
point, apiculate, striate, surrounded at base by a cup formed 
of the much enlarged and imbricated sep., dehiscing ventrally; 
seed about 3 in., oblong-ovoid, cotyledons plane-convex. 

Moist low country, up to 3000 ft.; rather common. FI. April. 

Also on the Malabar coast of India. 


Very strong ropes for tying buffaloes and strengthening fences are made 
by twisting the stems. 


2. CONNARUS, J. 


Shrubs often scandent; 1. imparipinnate ; fl. small, regular, 
in pyramidal panicles; cal. deeply 5-fid., segm. imbricate, not 
enlarged in fruit; pet. 5, distinct, erect ; stam. 10, inserted in 
cal.-tube, fil. slightly connate at base, the 5 opp. sep. long, 
with perfect anth., the 5 opp. pet. short with abortive anth.; 
carp. 5 (4 often abortive), stigma small; fruit a follicle or 
2-valved ; seed solitary, with a large aril, cotyledons plane- 
convex, no endosperm.—Sp. 55; 13 in /2. B. Jud. 


Fruit distinctly stalked, not striate. : . I. C. MONOCARPUS. 
Fruit not stalked, striate. : ; : . 2. C. CHAMPIONII. 


1. G. monocarpus, Z. Sf. Pl. 675 (1753). Radaliya, S. 
Chettupulukodi, 7. 

Herm. Mus. 39, 54. Burm. Thes. 199. FI. Zeyl. n. 248. Omphalo- 
bium indicum, Gaertn. Fruct. 1.217. C.astaticus, Willd. and C. pinnatus, 
Lam., Moon Cat. 49. Thw. Enum. 80. C. P. 328. 

Fl. B. Ind. ii. 50.. Burm: Thes. t. 890: ‘Gaertn. Fruct. into yess 
(fruit). 

A much-branched bush, young parts rusty-puberulous, 
l.-rachis 14-24 in., pubescent, thickened at base, Iflts. 3 or 5 
(1 or 2 pairs and a terminal one), on short, swollen, channelled 
stalks, 2-3 in., ovate-oblong or ovate-oval, rounded at base, 
obtuse, often twisted at apex, glabrous and somewhat shining 
above; fl. about } in., articulated on short, stout, densely 
pubescent ped., crowded in erect, irregularly pyramidal, 
pubescent, paniculate cymes terminating the branchlets ; cal.- 
segm. oblong-lanceolate, subacute, densely pubescent; pet. 
much longer than cal., oblong-linear, more or less pubescent 
outside ; follicle 24 in., fusiform, slightly falcate, apiculate, 
tapering below into a narrow stalk, surrounded at base by the 
persistent but not enlarged cal. and pet., smooth, brilliant 
scarlet, splitting into 2 valves which are smooth within ; seed 


Ellipanthus.] Connaracee. > 


I in., ovoid, slightly compressed, shining, purplish-black, sur- 
rounded at base by a large, pulpy, yellow aril. 

Dry region, very common; moist low country in sandy places rather 
rare, e.g., Colombo, Ratnapura, Deyandera, S. Prov. Fl. Sept.—Feb. 

Also in Southern India. 

Sir J. Hooker, in FI. B. Ind., refers Burmann’s figure to Rourea santa- 
loides; but, though not very good, it clearly represents C. monocarpus. 
The ripe follicles with the seed are very pretty objects, hanging in clusters 
from the ends of the branches. 


2. ©. Championii,* 7iw. Enum. 80(1858). Wel-radaliya, S. 
C. P. 2399. 

Fl. B. Ind. ii. 52. 

A climbing shrub, bark brown, rough with lenticels, young 
parts rufous-puberulous ; l-rachis 4-6 in., much tumid and 
wrinkled at base, cylindrical, smooth, Iflts. 5-7, on short 
swollen stalks, 3-4 in., oval, rounded at base, caudate-acumi- 
nate, glabrous, shining above, veins prominent beneath; fl. 
under } in., on very short rufous-hairy ped., arranged on 
spicate branches of spreading or pyramidal, rufous, axillary 
and terminal panicles, bracts small, rufous-woolly ; cal.-segm. 
linear-lanceolate, acute, covered with red pubescence; follicle 
1} in., obovoid, very blunt and truncate, tapering to narrow 
base (but not stalked), which is surrounded by persistent cal., 
nearly straight along upper edge, very gibbous along ventral 
suture, glabrous, striate with oblique ridges, opening by ventral 
suture only, scarlet; seed ?in., broadly ovoid, nearly black, 
aril as in the last. 

Moist region, up to 4000 ft.; rather rare. Hantane; Hunasgiria. 


Fl. Jan.; cream-coloured. 
Endemic. 


3. ELLIPANTHUS, //i./. 

Tree, |. reduced to a single Iflt.; fl. small, regular, often 
unisexual, racemose; cal. deeply 5-fid., segm. valvate, per- 
sistent but not enlarged in fruit; pet. 5, spreading ; stam. 5, 
alternating with 5 small, tooth-like fil. without anth., slightly 
connate at base; carp. solitary, style short, stigma dilated ; 
fruit a follicle ; seed solitary, with a large pulpy aril, cotyle- 
dons plane-convex, no endosperm.—Sp. 5; all in 7. B. Jnd. 


BZ. Thwaitesii, ///. f in Fi. B. Ind. ii. 55 (1876). 

Connarus unifoliatus, Thw. Enum. 80. £. unifoliatus, Hk. f., 1d. 410. 
C. P. 2443. 

Fl. B. Ind. ii. 55. Bedd. Fl. Sylv. t. 170 (2. unifoliatus). 


* Commemorates Capt. (afterwards Col.) J. G. Champion, 95th Regt., 
who was stationed in Ceylon 1838-47, and paid much attention to its 
Botany. Killed at Inkermann, in 1854. 


4 Leguminose. 


A small tree, 20-30 ft., twigs smooth, purple, young parts 
puberulous; 1. simple (unifoliolate), articulated on the short 
petiole, 2-3 in., oval, rounded at base, acuminate or slightly 
caudate, glabrous, rather coriaceous, reticulate venation pro- 
minent beneath ; fl. 2in., articulated on short pubescent ped., 
3-5 together, arranged in small axillary racemes less than 
1in. long; cal.-segm. linear-lanceolate, acute, rigid, pubescent; 
pet. longer than cal., obtuse, finely and densely pubescent ; 
fil. hairy at base ; ov. pubescent ; follicle 1-14 in., much falcate 
when young, afterwards nearly straight, tapering at base into 
a narrow stalk surrounded by persistent cal., sharply pointed, 
densely covered with yellow tomentum ; seed ? in., blue black, 
shining, covered half way up by the dark pink pulpy aril, 
embryo green. 


Moist region, 2000-4000 ft., rather rare. Deltota; Hunasgiriya; 
Gangala; Hantane; Maskeliya. Fl. Jan.; pale green. 
Endemic. 


E. unifoliatus is the older name; but, as it is applicable to the whole 
genus, Sir J. Hooker has rightly given a new one. 


XLIV.—LEGUMINOSE-. 


HERBS, shrubs, or trees, often twining or climbing ; l. alt., 
usually compound, with stip., lflts. always entire, often with 
stipels ; fl. bisexual (rarely unisexual), irregular or regular ; 
cal. more or less deeply cut, sometimes into separate sep., 
segm. usually 5, often unequal, or forming two lips; pet. 5, 
unequal or equal, or fewer, rarely connate or 0; stam. usually 
10, often fewer, sometimes indefinite, inserted on base of cal.- 
tube, often declinate, fil. frequently diadelphous (9 and 1) or 
monadelphous, rarely distinct; ov. superior, with I—many 
ovules on ventral suture, style simple, often declinate with the 
stam.; fruit a dry pod (legume), dehiscent along both 
sutures, or breaking up into joints, or indehiscent, rarely 
fleshy and drupe-like ; seeds without (rarely with) endosperm, 
cotyledons large, generally plane-convex. 

It is necessary to divide this vast Order into 3 Sub-orders, 
as follows :— 


I. Papilionacez. Pet. unequal, papilionaceous, imbricate, the 
upper one (standard) outside in bud, the two lateral (wings) and two 


Leguminose. 5 


lower (keel) in pairs; stam. 10 or less, usually di- or monadelphous. 
(Genera I-47.) 


Il. Ceesalpiniese. Pet. slightly unequal, imbricate, the upper one 
inside in bud; stam. 10 or less, distinct. (Genera 48-57.) 


III. Mimosez. Pet. equal, more or less connate into a corolla, 
segm. valvate in bud; stam. often indef., distinct or connate. (Genera 
58-64.) 


The largest Order in the Ceylon Flora, containing 207 species, or 
nearly 7 per cent. of the whole. The majority of these, however, are of 
wide distribution, and we have only 12 endemic species, Perzcopszs being 
the only endemic genus. 


I. Papilionacee. 


Stam. mon- or diadelphous. 
Pod not jointed. 
Pod dehiscent (exc. 6 and 7). 
Stems not twining. 
Stam. monadelphous (Gem/éste@). 


Keel-pet. nearly free, anth. uniform . I, ROTHIA. 
Keel-pet. connate, anth. dimorphous. 
Pod flat ; 3 : : . 2. HEYLANDIA. 
Pod inflated ; . 3. CROTALARIA 
Stam. diade]phous (exc. 6 and 7). 
L. 3-foliolate ( 77zfoliec) . 4. PAROCHETUS. 


L. pinnate (rarely simple), (Galegec). 
Anth. apiculate ; hairs attached by 
centre . 4 ‘ : : . 5. INDIGOFERA, 
Anth. blunt ; hairs basifixed. 
Pod indehiscent. 


Pod 1-seeded. Herb 6. PSORALEA. 
Pod several-seeded. Shrub — 7. MUNDULEA. 
Pod dehiscent. 
Without partitions between seeds 8. TEPHROSIA. 
With partitions ; 9. SESBANIA, 
Stems usually twining (exceptions in 27, 30, 
37, 39; 40, 41) 
L. pinnate ( Viciez) ; . 21. ABRUS. 
L. 3- (rarely 1-) foliolate (Phaseolea). 
Lfits. not gland-dotted. 
Style not bearded. 
Nodes of infl. not tumid. 
Pet. about equal. 
Stam. diadelphous. 
Cal. segm. distinct , . 22, SHUTERIA. 
Cal. truncate ; ‘ . 23. DUMASIA. 
Stam. monadelphous. 
Anth. all fertile ; . 24, GLYCINE, 
Alt. anth. abortive e . 25. TERAMNUS. 
Pet. very unequal. 
Keel longest. : ; . 26, MUCUNA, 
Standard longest (trees) . 27. ERYTHRINA, 


Keel and standard longer than 
wings ‘ P . . 28. STRONGYLODON. 


6 ; Leguminose. 


Nodes more or less tumid. 
Stam. diadelphous. 
Herbaceous twiner 
Erect tree 
Stam. monadelphous. 
Seeds 5-I0 ‘ c : 
Seeds 2 or 3 2 : 3 
Style bearded. 
Stigma oblique. 
Keel curved into a ring : 
Keel not or much less curved 
Stigma terminal. 
Pet. unequal . 
Pet. equal 
Lfits. gland-dotted. 
Seeds 3-6. 
Pod with deep lines between seeds 
Pod with faint lines 2 : 
Seeds I or 2. 
Funicle attached to end of hilum 
Funicle attached to centre of hilum. 
L. pinnately 3-foliolate 
L. palmately 3-foliolate : 
Pod indehiscent (Dalbergiea). 
Lfits. alternate 
Fl. white or pink 
Fl. yellow 
Lfits. opposite. 
Erect tree ; fruit not winged 
Climbers ; fruit winged on back 
Pod breaking up into 1-seeded joints 
(Hedysaree), (exc. 16 and 17). 
Lfits. without stipels. 
Stam. monadelphous. 
Joints muricated 
Joints wrinkled ‘ 
Stam. in two bundles of 5. 
Pod inclosed in cal. 
Pod exserted. 
Cal. 2-lipped 
Cal. with 5 segm. 
Lfits. with stipels. 
Pod 1-jointed and 1-seeded 
Pod not distinctly jointed, several- 
seeded. 
Pod turgid 
Pod flat 
Pod distinctly jointed. 
Joints folded on one another 
Joints in a line, turgid 
Joints in a line, flat 
Stam. free (Sophoree). 
Pod moniliform 
Pod oblong 


. GALACTIA. 
. BUTEA. 


. CANAVALIA. 
. DIOCLEA. 

. PHASEOLUS. 
. VIGNA. 

. CLITORIA. 

. DOLICHOS. 
. ATYLOSIA. 

. DUNBARIA. 
. ERIOSEMA. 
. RHYNCHOSIA. 
. FLEMINGIA. 
. DALBERGIA. 


. PTEROCARPUS. 


. PONGAMIA. 
. DERRIS. 


. ZORNIA. 
. STYLOSANTHES. 


. SMITHIA. 


. AXSSCHYNOMENE. 
. ORMOCARPON. 


. ELEIOTIS. 

. PYCNOSPORA. 

. PSEUDARTHRIA. 
. URARIA. 

. ALYSSICARPUS. 


. DESMODIUM. 


. SOPHORA. 
. PERICOBPSIS. 


Heylandia.] Leguminose. 7 


Of the 159 species in this Sub-order, as many as 131 are found 
only in the low country, where many are common weeds. Confined 
entirely to the dry region are 51 species, including Io species of /#digo- 
fera, and all those of the following genera :—Mundulea, E-schynomene, 
Ormocarpon, Strongylodon, Butea, Pterocarpus, Rothia, Heylandia, 
Psoralea, Stylosanthes, and Clitorta. Limited to the moist low country 
are 25 species, including the genera Déoclea, Vigna, Pseudarthria and 
Pericopsis. The remaining 28 species are found in the montane region, 
but only 17 are confined to it, including the genera Parochetus, Shuteria, 
and Dumasia, 4 species of Crotalaria, 6 of Desmodium, and 2 of Atylosia. 

The great majority are widely distributed plants, and only 9 are 
endemic to Ceylon. 


1. ROTHIA, /ers. 


Annual herb, |. 3-foliolate; fl. very small, in axillary 
racemes ; cal.-segm. as long as tube, 2 upper broader ; pet. 
nearly equal, clawed; keel-pet. very slightly coherent ; 
stam. monadelphous, anth. uniform ; style short, stigma capi- 
tate; pod linear, many-seeded, not septate within.—Sp. 2; 
tin FZ. B. Ind. 


R. trifoliata, Pers. Syn. P/. ii. 302 and 659 (1807). 

Herm. Mus. 33. FI. Zeyl.n. 285. Z7¢gonella indica, L. Sp. Pl. 778. 
Moon Cat. 55. Thw. Enum. 82. C. P. 1452. 

FI. B. Ind. ii. 63. Wight, Ic. t. 199. 

A much-branched, spreading, annual herb, stem slightly 
‘woody at base, branches slender, hairy (as is whole plant) ; 
l. small, rachis about }in., slender, Iflts. about 4 in., sessile, 
narrowly lanceolate, subacute, minutely mucronate, hairy on 
both sides, stip. small, linear-oblong, acute, persistent ; fl. on 
very short ped., solitary or 2-4 in very short axillary racemes ; 
cal.-segm. lanceolate, acuminate, silky-hairy; pet. twice as long 
as cal.; pod 1}-—1}in.; narrowly linear, nearly straight, 
slender, silky-hairy, tipped with very slender style; seeds 
numerous, closely packed. 


Dry region ; common in sandy places. Fl. Feb. &c.; pale pink. 
Throughout India, also in N. Australia. 


Ulex europaeus, L. The English Furze or Gorse is a conspicuous 
plant at Nuwara Eliya, where it is quite naturalised and seeds freely. It 
does not, however, spread much. The date of its introduction 1s not 
known to me. 


2, HEYLANDIA, WC. 
Annual herb, |. simple, without stip.; fl. small, solitary, 
axillary ; cal. deeply cut, upper segm. slightly coherent ; wing- 
pet. much shorter than standard, keel-pet. connate; stam. 


8 Leguminose. [Crotalaria 


monadelphous, anth. dimorphous, 5 large and 5 small; style 
long, sharply bent at base; pod small, flat, 1-2-seeded.— 
Monotypic. 

H. latebrosa, DC. Mem. Leg. 201 (1825). 

Thw. Enum. 81. C. P. 1275. 

Fl]. B. Ind. ii. 65. DC. Le. t. 34 (A. hebecarpa). 

An annual herb, with numerous prostrate spreading 
branches 6-24 in. long, much branched, clothed with lone 
spreading hair; 1. numerous, small, close, nearly sessile,, 
simple, broadly ovate, cordate and often unequal at base, 
apiculate, sparingly hairy on both sides, minutely punctate,, 
stip. 0; fl. nearly sessile in the axils; cal.-segm. acute; pet. 
much exserted ; pod very small, § in., surrounded at base by 
persistent cal., ovate-oblong, compressed, tipped by long style,. 
smooth or with a few long hairs, pale brown. 

Dry region; common. Extending also into the intermediate region. 
as Galagama, Nalande, Etampitiya, &c. Fl. Nov.-April; yellow. 

Also throughout India. 


There are specimens from Moon, 1819, in Mus. Brit., but it is not in— 
cluded in his Cat., unless his Crotalaria huméfusa, Cat. 52, be this. 


3. CROTALARIA, J. 


Herbaceous or semi-shrubby ; |. simple or 3- (very rarely 
5-) foliolate, with or without stip.; fl. in terminal or leaf- 
opposed racemes ; cal.-tube campanulate, short, segm. nearly 
equal, the upper 2 often more or less connate; wing-pet. 
shorter than standard, keel-pet. equalling wings, completely 
connate and prolonged into a sharp curved beak; stam. 
monadelphous, anth. dimorphous; style long, abruptly curved. 
upwards at base, stigma minute; pod sessile or stalked, 
straight, turgid or inflated, usually many-seeded, rarely 
I-seeded.—Sp. about 300; 77 in F/, B. Ind. 
_Leaves simple. 


Racemes solitary ; seeds numerous. 
Stems prostrate or ascending, diffuse, herbaceous. 


Stip. o. 
Fl. 1-4 in raceme. 
Pod globular 2 5 : 3 - | f.) CA BIFLORA: 
Pod oblong . 5 5 5 : . 2. C. PROSTRATA. 
Fl. 4-20 in raceme. 
L. linear-spathulate . : . 9g. C. ALBIDA. 
L. oval-oblong ; raceme capitate. . 10, C. NANA. 
L. obovate-oblong ; raceme lax . . II, C, LINIFOLIA. 


Crotalaria.) Leguminosae. 9 


Stip. present. 
Pod glabrous. 


L. oval; stip. foliaceous 3. C. FERRUGINEA. 
L. linear ; stip. small . 7. C. MYSORENSIS. 
Pod hairy. 
Stem nearly cylindrical 4. C. EVOLVULOIDES. 
Stem strongly triquetrous 8. C. TRIQUETRA. 
Stem erect, robust, often semi- shrubby. 
Stip. large, decurrent . : 6. C. RUBIGINOSA. 
Stip. small, not decurrent. 
Fl. yellow. 
Pod glabrous. 
Plant very hairy . : : ig. CAEYCINA, 
Plant nearly glabrous. : : 4. Cy RETUSA 
Pod puberulous . : - : . 16. C. WALKERI. 
Pod densely conte - - - . 5. C. MULTIFLORA. 
Fl. purple = ; : : . 15. C. VERRUCOSA. 
Stip. o. 
Pod glabrous . : : : : veh TECTA, 
Pod hairy A , . 17. C. JUNCEA. 
Racemes panicled ; seed solitary - ; . 18. C. LUNULATA. 
Leaves compound. 
Pod small, subglobose, 1-2-seeded . : . 19. C. MEDICAGINEA. 
Pod large, oblong, with many seeds. 
Lfits. oval. 
Pod with a short stalk 
Pod silky, 10-12-seeded . : : . 20. C. CLAVATA., 
Pod glabrous, 20-30-seeded . . . 2. OC STRIATA, 
Pod with a very long stalk. : : . 22. C. LABURNIFOLIA. 
Lfits. linear-oblong ; - : : . 23. C. QUINQUEFOLIA. 


1. ©. biflora, 2. Mant. Alt. 570 (1771). 

C. nummudlaria Willd., Moon Cat. 52. Thw. Enum. 81 and (C. g/odosa) 
410. C.P. 3325, 3736. 

Fl. B. Ind. ii. 66. 

Herbaceous, annual (?), with copious, trailing, very much- 
branched, elongated, silky or hairy stems; |. small, }—} in, 
nearly sessile, simple, ovate or roundish, mucronate; hairy 
on both sides, without stip.; fl. solitary or 2 together at end 
of slender hairy ped. twice as long as 1.; cal.-segm. long- 
acuminate, hairy ; pet. scarcely longer than cal.; ; pod about 
3 in., shortly stalked, nearly globular, pericarp soft, compres- 
sible, set with scattered hairs ; seeds about 20, smooth, brown. 

Dry oa rare. Jaffna, abundant; Trincomalie (Glenie). Gardner's 
specimens (C. P. 3325) have no locality. Fl. Feb., March; yellow. 

Also in Peninsular India and Java. 

The true C. globosa, W. and A., has not been collected in Ceylon. 


The Jaffna specimens are less slender than the Trincomalie ones, and 
rather shaggy-woolly than silky. 


2. C. prostrata, Koxl. Hort. Beng. 54 (1814). 
Thw. Basan. 81. C. P. 1266. 
Fl. B. Ind. ii. 67. 


IO Leguminose. [ Crotalaria. 


Annual, with many slender, ascending, hairy branches, |. 
sessile, $—-1 in., oblong, obtuse, usually unequal and subcordate 
at base, hairy on both sides especially beneath, without stip. ; 
fl. small, pedicellate, in racemes of 2-4, peduncle slender, 
divaricate, hairy, very much exceeding |.; cal.-segm. very hairy; 
pet. scarcely exceeding cal.; pod small, scarcely 4 in., oblong, 
inflated, glabrous; seeds about 12, small, highly polished. 

Dry region; rather rare; extending into intermediate region. Uma- 
oya Etampitiya; Batticaloa; Ekiriyankumbura. FI. March, April ; 

ellow. 
: Also in India and Java. 


3. C. ferruginea, Grah. in Wall. Cat. 5398 (1828). 

hw Enum or.. (Cb. 1265: 

Fl. B. Ind. ii. 68. 

A tall herb with long ascending branches covered with short, 
fulvous or pale yellow hair; 1. 14-24 in., distant, very shortly 
stalked, oval or oblong or obovate-oval, acute at base, obtuse 
and mucronate at apex, hairy on both surfaces, paler beneath, 
stip. conspicuous, }in.or more, foliaceous, acuminate, spreading, 
persistent; fl. rather small, on very silky ped., in lax, elongated, 
very silky, erect, few-fld., leaf-opposed racemes, bracts very 
acute, persistent; cal.-segm. all elongated, very acute, densely 
silky ; pet. scarcely as long as cal.; pod about 1 in., oblong, 
inflated, tapering to base but not stalked, tipped with short, 
stiff, recurved style, smooth, veined, black ; seeds about 30. 

Moist region, from 2000 to 4000 ft., rather rare. Near Kandy ; Ram- 


boda ; Maturata ; Lunugala; Ella. Fl. all the year; pale yellow. 
Also in E. Bengal, Burma, Java, and the Philippines. 


4. S. evolvuloides, Wight in Wall. Cat. n. 5410 (1828). 

Arn. Pug. 11. Thw. Enum. 81. C. P. 3593. 

Fl. B. Ind. ii. 68. 

A much-branched straggling herb, semi-woody at base, 
branches elongated, divaricate, slightly angular, hairy- 
pubescent ; |. #-14 in., on very short, hairy petioles, varying 
from rotundate to oval or oblong-lanceolate, obtuse, mucronate, 
rounded at base, glabrous above, shortly hairy beneath, stip. 
small, linear ; fl. rather small, on short pubescent ped., laxly 
arranged in elongated, curved, pubescent, few-fid., secund, 
axillary racemes greatly exceeding 1., bracts small; cal.-segm. 
acute, pubescent ; pet. rather longer than cal.; pod small, 
scarcely 3 in., broadly ovoid, inflated, not stalked, tipped with 
strong base of style, densely hairy, pale yellowish-brown ; 
seeds about 12. 

Dry region; rare. Uma-oya; Seven Korales district; Ekiriyan- 
kumbura, Uva. FI. Jan.-April ; yellow. 

Also in S. India. 


Crotalaria. Leguminose. Il 


5. ©. multifiora, Benth. zn Lond. Journ. Bot. ii. 478 (1843). 
C. bifaria, var. (?) multifiora, Arn. Pug. 11. Thw. Enum. 81. C. P. 


1268. 
Fl. B. Ind. ii. 69. 


A robust herb, not much branched, branches stout, cylin- 
drical, curved, densely and softly hairy; 1. on very short hairy 
petioles, numerous, rather closely placed, very variable in form 
and size, 1—4in., rotundate or broadly or narrowly ovate or ovate- 
lanceolate or linear, rounded at base, acute and apiculate at 
apex, shortly hairy on both sides, pale and with prominent 
reticulation beneath, stip. linear, small, divaricate or reflexed ; 
fl. very large, 2 in., on stout nodding ped., 2-6 arranged in 
racemes at end of long axillary (or apparently terminal) 
peduncles, bracts small, reflexed ; cal. 1 in., segm. foliaceous, 
lanceolate, acute, pubescent ; pet. much exserted ; pod I} in., 
oblong-ovoid, not stalked, cylindrical, tipped with long style, 
densely clothed with fulvous or brown silky hair, pale green 
blotched with purple ; seeds about 12. 

Montane zone, up to 6000 ft., on the patanas; rather common. 
Dolosbagie ; Elk Plains ; Hewahette; Bandarawella; Haputale. FI. 
Sept., Oct.; sulphur-yellow with purple veins. 

Endemic. 

The most ornamental of our species, with very large showy flowers ; 
but the name mu/tzflora is scarcely justified. 


[C. bifaria, L.f. is given for Ceylon in Fl. B. Ind. ii. 69, on the faith of 
a specimen so labelled by Mr. Bentham in Hb. Kew, without collector’s 
name. An error is probable, and the specimen really from S. India.] 


6. ©. rubiginosa, Wi//d. Sp. P1. iii. 973 (1800). 

C. Wightiana, Grah., Thw. Enum. 81. C. P. 2772. 

Fl. B. Ind. ii. 69. Wight, Ic. t. 885. 

A robust herb, erect or suberect and spreading, usually 
much branched, 1-2 ft. high, stem softly hairy; 1. very 
variable, 1-2 in., very shortly stalked, broadly oval or oblong- 
linear, very obtuse, apiculate, more or less densely silky or 
velvety with golden or grey hair, lat. veins strongly marked 
beneath; stip. large, decurrent on stem so as to form wings 
with the upper part dilated falcate and spreading; fi. 
moderate-sized, on curved, silky ped. 1—3 in., in long racemes, 
peduncle leaf-opposed, spreading or ascending, stiff, silky, 
bracts ovate, acute, bracteoles immediately beneath cal. 
lanceolate ; cal.-segm. large, nearly } in., acuminate, densely 
silky with golden or fulvous hair; pet. scarcely longer than 
cal.; pod 14-1} in., shortly stalked, narrowly oblong-ovoid, 
turgid, tipped with sharp hooked stiff base of the style, 
glabrous, pale brown, often marked with white streaks ; 
seeds 20-30. 


12 Leguminose. [ Crotalaria. 


Montane zone, up to 6000 ft. on the patanas ; rather common, especi- 
ally in the Uva Province, where it also occurs at lower elevations, as at. 
Bibile. Fl. Aug.; pale yellow. 

Also on the Nilgiris. 

Easily known by the stipular wings to the stem. Our form (C. 
Wightiana, Grah.) is usually more robust than the type, but it occurs also. 
on the S. Indian hills, and there are intermediate forms. 


7. ©. mysorensis, Roth Nov. Pi. 338 (1821). 
Thw. Enum. 82. C. P. 3594. 
Fl. B. Ind. ii. 70 (not given for Ceylon). 


An erect herb, 2-3 ft., with numerous long ascending 
branches, densely hairy, with long, spreading hair; |. 14-23 in., 
numerous, ascending, nearly sessile, linear or strap-shaped, 
obtuse, strongly apiculate, covered, especially beneath, with 
long, white, spreading hair, stip. acicular-linear, strongly api- 
culate, 2in., spreading ; fl. moderate-sized, on 4 in. ped., in lax, 
erect, terminal racemes; cal.-segm. 2 in., very acuminate, 
upper ones lanceolate, lower linear, all clothed with long, 
spreading hair; pet. slightly longer than cal.; pod 1} in., 
oblong-ovoid, narrowed at base but scarcely stalked, truncate 
at apex, which is tipped by the very strongly deflexed base 
of style, glabrous, pale brown; seeds about 30. 

Dry region; rare. Batticaloa; Nilgala; Uma-oya. FI]. Jan.—April;, 
yellow. 

Throughout Peninsular India. 


A handsome plant, covered with long silvery hair, which becomes. 
fulvous in drying. : 


8. * GC. triquetra, Dalzell in Kew Journ. Bot. ii. 34 (1850). 
Thw. Enum. 410. C. P. 3832. 
Fl, B. Ind. ii. 71. 


A spreading under-shrub, 1-2 ft. high, with numerous, 
slender, ascending, strongly triquetrous, slightly hairy branches; 
l. 14-2 in., very shortly stalked, oblong-oval, subcordate at 
base, obtuse, glabrous, thin, stip. small, lanceolate; fl. small, 
in very long lax racemes, bracts minute ; cal.-segm. linear- 
lanceolate, acute, silky ; pet. slightly longer than cal.; pod not 
seen (‘? in., oblong, cylindrical, shortly silky, with 15-20 
seeds’), 

Probably an introduction. The only specimens I have seen are the 
very poor C.P. ones. Thw. (l.c.) gives the locality ‘Peradenia, rare.” 


Fl. pale yellow. 
Also in Southern India. 


9g. ©. albida, Heyne in Roth. Nov. Pl. 333 (1821). 
Thw. Enum. 82. C. P. 269. 
BESBs lndaintene 


Crotalaria.) Leguminose. 13 


A perennial herb or small under-shrub, with a short, 
woody base and very numerous ascending, slender branches 
6 in.—2 ft. long, covered with short, adpressed hair; 1. numerous, 
_ rather close, small, 4}—-}in., shortly stalked, linear-spathulate, 
acute at base, obtuse, truncate or retuse at apex, apiculate, 
glabrous, and minutely punctate above, silky with adpressed 
hair beneath, without stip.; fl. small, numerous, on slender 
ped., secund, in lax, terminal racemes, bracts minute, often 
on the cal.-tube; cal.-segm. gin., the upper linear-oblong, 
very obtuse, the lower linear-lanceolate, acuminate, all very 
finely silky; pet. about as long as cal.; pod 4 in., oblong-ovoid, 
not stalked, tipped with hooked base of style, glabrous, brown; 
seeds 6-12. 

Lower montane zone; rather rare. Abundant in the Uva hill- 
ceansy on the patanas; Maturata; Galagama. FI. Jan., Feb.; bright 

eulow. 
Throughout India and Malaya, also in Philippines and China. 


10. ©. nana, Burm. Fl. Ind. 156 (1768). 
Moon Cat. 52. Thw. Enum. 82. C. P. 3301. 
Fl. B. Ind. 11.71. Burm. FI. Ind. t. 48, f. 2. 


A small herb, 6-12 in. high, with numerous short branches 
from the base of the stem, slender, prostrate, clothed with long 
hair; 1. }-2in., very shortly stalked, oval-oblong, obtuse at 
both ends, apiculate, hairy on both sides, minutely pitted 
above, without stip.; fl. small, on short, hairy ped., 2-6 in a 
crowded capitate terminal raceme; cal.-segm. shaggy with 
long hair, upper connate, lower linear, acuminate; pod }in,, 
scarcely exceeding cal., globular-ovoid, tipped with short, 
reflexed style, glabrous, rough, black; seeds 10-12. 

a B,umbellata. C. umbellata, Wight, Thw. Enum. 82. C.P. 
1276. 

More robust, often 2 ft., much less branched below, but 
with several stout, stiff, ascending branches above which ex- 
ceed the central one; fl. more numerous, 4-8 in a terminal 
umbel ; seeds 6-8, rather larger. 


Low country; rare. Katukanda, Negombo (Thwaites). Moon gives 
Colombo. Var. £, lower montane zone, 3000-4000 ft.; rather rare. 
Ramboda; Maturata; Passara. FI. July, Sept., Feb.; yellow. 

Also in Peninsular India. 


11. ©. linifolia, Z./ Supp. P/. 322 (1781). 
Thw. Enum. 82. CF. 1277. 
Fl. B. Ind. ii. 72. 


Herbaceous, stems numerous, prostrate, spreading, slender, 
densely silky with adpressed hair; |. small, under }in., on 


14 Leguminose. [ Crotalaria. 


short silky petioles, obovate-oblong, obtuse at both ends, 
glabrous above, silky beneath, without stip.; fl. small, numerous, 
on curved silky ped., arranged in erect, lax, terminal racemes; 
upper cal.-segm. connate, lower lanceolate, acuminate, all 
densely hairy ; pod not seen (‘small, as long as cal., ovoid- 
oblong, glabrous, black, 8—10- seeded ’). 

Moist low country; very rare. The only specimens are the C. P. 
ones collected near Galle by Gardner, and they are imperfect and barely 
sufficient for identification. 

Has a wide distribution throughout the E. Tropics. 


12. C. tecta, Heyne in Roth. Nov. Sp. 334 (1821). 
Trimen in Journ. Bot. xxvii. 162. 

Fl. B. Ind. ii. 72 (not given for Ceylon). 

Semi-shrubby at the base, stems 1 ft. or more, numerous,, 
ascending, stiff and wiry, much branched, clothed with ad- 
pressed white hairs; |. small, $-2in., shortly stalked, oval- 
oblong, obtuse at both ends, glabrous above, adpressed-hairy 
beneath, without stip.; fl. small, on curved hairy ped., in short, 
lax, terminal, racemes; cal. very hairy, upper segm. connate, 
lower much acuminate; pod rather over }in., scarcely ex- 
ceeding cal., oblong, cylindrical, not stalked, truncate at end, 
tipped with short style, glabrous, grey; seeds about 10. 

Dry country ; very rare. Collected in dry paddy-fields at Mineri,. 
N.C. Prov., 1885. Fl. Sept.; pale yellow. 

Also in S. India. 
Perhaps scarcely more than a suffruticose state of C. /znzfolia. 


13. C. calycina, Schrank, Pl. Rar. Monac. t. 12 (1819). 

C. anthylloides, W. and A. Prod. 131 (non Lam.). Thw. Enum. 82. 
CARA526. 

HMB Indi 72: 

An erect herb, 1-2 ft. or more, stem stout, with long, 
ascending branches, densely clothed with fulvous hair ; 1. very 
variable, nearly sessile, 14-6in., from oblong-oval to linear, 
subacute at base, obtuse or acute and mucronate at apex, 
conduplicate when young, glabrous or sparingly hairy above, 
more or less hairy or shaggy beneath, stip. subulate, hairy; fi. 
large, on stout, nodding ped. (which become deflexed in fruit), 
4-8 in lax terminal racemes, bracts in pairs immediately beneath 
each fl., large, lanceolate, acuminate, shaggy with long hair ; 
cal. large, 2-1in., densely clothed outside with long, shaggy, 
fulvous hair, glabrous within, upper segm. shortly connate at 
base, obovate-oblong, obtuse, lower ones lanceolate-linear, 
acuminate ; pet. equalling cal.; pod rather shorter than cal., 
not stalked, glabrous, dark brown; seeds about 20, pale brown, 
polished. 


Crotalaria] Leguminose. 15 


Montane region, descending a little into upper zone of low country, on 
patanas; common. Also at Nilgala, Uva. Fl. Jan., May—Sept.; yellow. 

Throughout the E. Tropics and in Trop. Africa. 

On the mountains at high elevations this is a dwarf plant, 6 in. high, 
with smaller fl. and small linear leaves. 


14. C. retusa, /. Sf. P/.715 (1753). Maha-andana-hirlya, S. 
Kilukiluppai, 7. 

Burm. Thes. 80. Herm. Mus. 20. FI. Zeyl.n. 276. Moon Cat. 52. 
Thw. Enum. 81. C. P. 1274. 

Fl. B. Ind. ii. 75. Bot. Reg. t. 253. Herm. Hort. Lugd.-Bat. t. 201. 

A stout, semi-shrubby herb, with a few long, erect, striate 
branches, glabrous, or with adpressed pubescence; 1. 14-2} in., 
on very short, pubescent ped., narrowly obovate-oblong, 
tapering to base, very obtuse, often emarginate, glabrous 
above, finely pubescent beneath, stip. very small, subulate ; 
fl, numerous, large, 1}in. on stout, curved, pubescent ped., 
closely placed in erect, terminal racemes 6-12 in. long, bracts 
shorter than ped.; cal. large, nearly glabrous, tube wide, cam- 
panulate, segm. triangular, acute; pet. much exserted; pod 
14-1} in., oblong, cylindrical, tapering at base to a short, 
broad stalk, tipped with slightly curved style-base, glabrous, 
purplish-green ; seeds 15-20. 

Var. 6, maritima, 7rim. Syst. Cat. Ceyl. Pl. 22. 


Stems prostrate, densely pubescent; fl. more crowded in 
shorter racemes. 

Low country, especially the dry region, up to 2000 ft., in waste places. 
Var. 8 on the seashore. FI. Aug.-October; bright sulphur-yellow. 

Found throughout the Tropics. The Fl. B. Ind. states this to be 


‘often cultivated,’ and it has a tough fibre, but it is not grown in Ceylon 
on that account. 


Hermann (Mus. 20) gives the name “Jacberija’ (Yakbériya) for this, 
and explains its meaning (see C. /aburnifolia). 


15. ©. verrucosa, /. Sf. ?/.715 (1753). Nil-andana-hiriya, 
S. Kilukiluppai, 7: 

Burm. Thes. 81. Herm. Mus. 65. Fl. Zeyl. n. 277. Moon Cat. 52. 
Thw. Enum. 81. C. P. 1273. 


Fl. B. Ind. ii. 77. Burm. Thes. t. 34. Herm. Hort. Lugd.-Bat. t. 199. 
Wight, Ic. t. 200. 

A large, much-branched herb, 2—3 ft., branches divaricate, 
irregularly quadrangular, finely puberulous, buds silky; 1. 
1}-1j in., shortly petiolate, spreading, ovate-rhomboid or 
ovate-deltoid, tapering to base, obtuse, undulate, with minute 
scattered pubescence on both surfaces, paler beneath, with 
prominent veins, somewhat bullate above, stip. rather large, 
foliaceous, 4-ovate or }-cordate, acute, persistent; fl. rather 
large, j-1in., on short, curved, downy ped., about 6-12in., 


16 Leg UMUNOSE. [ Crotalaria. 


erect, leaf-opposed (apparently terminal), slightly curved 
racemes, bracts linear, acuminate, shorter than ped.; cal. 
minutely pubescent, segm. nenrouly triangular, acuminate, 
the upper rather shorter; pod 14-14in., oblong, cylindrical, 
rather dilated at end, not stalked, hairy, nearly black; seeds 
about 12, yellow. 


Low country; a common weed in open ground. FI. all the year ; 
bluish-purple, rarely white. 

Found throughout the Tropics. 

The white-flowered form is occasionally found; but I have never 
seen it with yellow fl. as given in ‘ Fl. B. Ind.’ 


16. ©. Walkeri, Arn. in Act. Acad. Nat. Cur. xviii. 328 (1836.) 
[PLATE XXVI.] 

C. semperfiorens, Vent., Thw. Enum. 81. C. P. 12. 

Fl]. B. Ind. ii. 78 (C. semperflorens var.). 


A large perennial herb, often semi-shrubby at base, stems 
elongated, sarmentose, semi-scandent, with many spreading 
branches, slightly striate, glabrous or nearly so, young parts 
very finely pubescent; 1. 2-4 in.,on short, stout petioles, oval or 
lanceolate-oval, obtuse or acute, finely mucronate, glabrous 
above, usually puberulous beneath, thin, stip. variable, usually 
very small, either subulate, linear, or linear-lanceolate and 
very acute, or often falcate or semi-lunar, rarely foliaceous 
and larger; fl. rather large, 1 in., on rather slender, short ped., 
4—I0 in erect, loose, terminal racemes, with two minute bracts 
immediately beneath cal.; cal. very finely puberulous, tube 
broadly campanulate, segm. narrowly triangular, acuminate ; 
pod 2-24 in., tapering to distinct stalk at base, oblong, much 
inflated, especially at truncate apex, tipped by short, broad 
style-base, very finely puberulous ; seeds 7—I0. 

Montane zone, especially at the higher elevations; rather common, 
scrambling over bushes like a climber. FI. all the year, especially in 
December and January; bright golden yellow. 

Endemic (?). 

A very ornamental plant, well worth cultivation. Differs from C. 
semperfiorens, Vent. (C. Wallichiana, W. and A.) of the Nilgiri Hills in its 
muchless pubescent leaves and inflorescence, andits much smaller stipules, 
but possibly only a variety. C. Wallichiana is figured in Wight, Ic. t. 982. 
A dwarf variety occurs on the Horton Plains patanas, with short 
prostrate stems, small leaves, 3-3 in. long, and smaller pods less inflated 
at the ends. 


17. ©. juncea, L. Sp. P/. 714 (1753). Hama, S. 
Burm. Thes. $2. Moon Cat. 52. Thw. Enum. 81. C. P. 1264. 
Fl. B. Ind. ii. 79. Roxb. Cor. t. 193. Duthie Field-crops, t. 21. 


An erect annual, 2-4 ft. or more, branches numerous, stiff, 


Crotalaria.| Zs Cgune 220SE. 1 F/ 


ascending, slender, striate, finely pubescent; 1. 14-4 in., dis- 
tant, on very short, thickened petioles, lanceolate-oblong or 
linear, acute at base, subacute, apiculate at apex, pubescent on 
both sides with adpressed silvery hair, stip. none or very 
minute; fl. large, 1f in. or more, on stout, curved, pubescent 
ped., 3-8 in erect, lax, terminal racemes, bracts very small, 
2 on cal.-tube; cal. nearly 1 in., shining with wants hair, segm. 
very deep, lanceolate- linear, acuminate ; pod 1-1} in., oblong, 
cylindrical, not stalked, tipped with ‘short, nearly straight 
style-base, brown, densely covered with short, stiff white hair; 
seeds about 12. 

Low country, principally in the dry region ; rather common. Doubt- 
fully native in the moist region. Fl. Aug.—Oct.; brilliant yellow. 

Found in E. Tropics generally, but much cultivated. 

This affords the San or Sun Hemp of commerce ; it is grown in small 
quantity by the Sinhalese as about Chilaw, and by the Tamils at Jaffna. 
It is generally known here simply as ‘ Hemp.’ 


C. fulva, Roxb., is given for Ceylon in FI. B. Ind. on the faith of 
specimens in Kew Herbarium (Gardner, 199, and G. Thomson from 
Kandy). They may have been cultivated in the Gardens. 


18. ©. lunulata, Heyne in Wall. Cat. n. 5378 (1828). 

faweeoum. 81. C.P. 1270. 

Fl. B. Ind. ii. 80. Wight, Ic. t. 480. 

An erect, stiff herb, 1-2 ft. high, branched in upper part, 
stem and branches copiously clothed with soft, spreading 
white hairs (becoming brown when dried); 1. closely placed, 
4-24 in.,on very short petioles, oval or obovate-oblong, sub- 
acute at base, obtuse or rarely acute, apiculate, densely clothed 
with silky hair on both surfaces, stip. small, broadly oval, 
obtuse, amplexicaul, recurved, glabrous and shining (often 
absent); fl. rather small, # in., numerous, on rather long, 
slender, hairy ped., arranged in drooping racemes which 
terminate divaricate or spreading branches, the whole forming 
a large paniculate inflor., bracts numerous, conspicuous, 
rotundate-cordate, acute, reflexed, hairy beneath, glabrous and 
shining (like the stip.) above, many on the lower part of the 
racemes often empty; cal. very deeply cut, segm. about 4 in., 
equal, linear, very acuminate, hairy outside, glabrous within ; 
pet. slightly longer; pod small, not exceeding cal., Zin., not 
stalked, oval-oblong, compressed, tipped by very long style- 
base, sparingly clothed with long hair; seed solitary. 

Dry region; rare. Jaffna; Batticaloa; a few miles N. of Kurune- 
gala; Kantalai. FI. Aug.—Oct., Feb.-March; yellow. 

Also in S. India. 

Very unlike our other species. 


PART I. Cc 


18 Leguminose. [ Crotalaria. 


19. ©. medicaginea, Lam. Encyc. Meth. ii. 201 (1786). 

Thw. Enum. 82. C. P. 1278. 

Fl. B. Ind. ii. 81. 

A small perennial herb, with a thick woody branched 
rootstock, and numerous, slender, filiform, spreading, much- 
branched, prostrate stems with scanty adpressed hair ; 1. very 
small, 3-foliolate, petiole ~,-% in., stip. minute, acicular, lflts. a 
little longer than petiole, stalked, obovate or oval, acute, 
obtuse or truncate, with adpressed hair on both surfaces, paler 
beneath ; fl. very small, on slender ped., 2-5 in leaf-opposed 
racemes; cal. with adpressed hairs, segm. oblong-linear, pet. 
about twice as long as cal.; pod pisiform, 4 in., pointed or 
beaked; clothed with adpressed hair; seed solitary. 


Var. 6, luxurians, Baker in Fl. B. Ind., l.c. 


Stem erect, 1-2 ft., more strongly pubescent; lfits. larger, 
4-3 in., glabrous above; fl. 6-10 in raceme; pods in a crowded 
head. 

Dry region, chiefly on seacoast ; rather rare (?). Near Jaffna, abun- 
dant; Trincomalie; Batticaloa. Var. 8, Intermediate region. Haragama; 
Ella Pass, Uva. Fl. Sept.—Feb., dull yellow. 

General in the E. Tropics ; also in China and Afghanistan. 


C. Willdenowtana, DC. was found once or twice about Colombo by 
W. Ferguson, but is not native (Thw. Enum. 441). It is C. P. 3853. 
The specimens are poor. 


C. incana, L. A weed in the Botanic Garden, Peradeniya and neigh- 
bourhood for many years past, but no doubt introduced ; Thw. Enum. 82. 
C. P. 1269. It seems to be the plant recorded in Moon Cat. 52 as 
C. capensis, and is becoming common in waste places. A general Tropical 
weed, probably originally American. 


20. ©. striata, DC. Prod. ii. 131 (1825). 
Thw. Enum. 82, 410. C. P. 3810. 
Fl. B. Ind. ii. 84. Bot. Mag. t. 2300. 


A tall herb, 2-3 ft., stem stout, somewhat striate, with few 
long spreading branches, very minutely puberulous, young 
parts silky; 1. large, 3-foliolate, petiole 2—3 in., stout, spreading, 
stiff, swollen at base, finely pubescent, stip. o, lflts. longer 
than petiole, on short, swollen, pubescent stalks, broadly oval, 
acute at base, obtuse, often emarginate, apiculate, thin, 
glabrous above, minutely puberulous and glaucous beneath ; 
fl. numerous, moderately large, on short deflexed ped., densely 
arranged in narrow erect spicate racemes 6-12 in. long and 
terminating stem and branches, bracts filiform, early deciduous; 
cal. minutely pilose, truncate at base, segm. equal, lanceolate- 
linear; pet. much exserted, standard narrow, strongly bent 
upwards ; pod 14 in. or rather more, much deflexed, linear- 


Crotalaria.] Leguminose. 19 


oblong, cylindrical, with a very short stalk, very minutely 
puberulous (apparently glabrous), pale cinnamon-brown ; 
seeds about 25, dark brown. 

Var. 6, acutifolia, Zyrim. Cat. Ceyl. Pl. 22. Thw. Enum. 4lIo. 
C. P. 3608. 

Lfits. large, acute; pods longer, with more numerous seeds. 


Low country in waste ground common ; var. 6, about Kandy. 
Fl. Dec.—Feb., dull yellow, veined with purple. 
Found throughout the Tropics. 


21. G. clavata, W. and A. PRUE 194 (1834). 

FI. B. Ind. ti. 83. 

A small semi-shrubby herb, branches with fine adpressed 
hair, |. 3-foliolate, lflts. small, 4—?in., obovate-oblong, tapering 
to base, often retuse at apex, glabrous above, slightly downy 
beneath, rather thick, pale green, stip. minute, setaceous, re- 
curved, pubescent ; fl. rather small, on short pubescent ped., 
in short erect racemes, bracts deflexed, setaceous; cal. 
pubescent, segm. narrowly triangular, acute ; pod about 1 in. 
deflexed and somewhat curved, slightly widened at end, finely 
silky ; 10—12-seeded. 

Dry country ; very rare, only found as yet near the Giant’s Tank, 


Mannar. FI. Feb.; yellow, not striped with red. 
Also in S. India. 


22. ©. laburnifolia, Z. Sp. P/. 715 (1753). Wak-bériya, S. 
e = Thes. 82. FI. Zeyl. n. 278. Moon Cat. 52. Thw. Enum. 82. 

Fl. oa Ind. ii. 84. Burm, Thes. t. 35. Herm. Hort. Lugd.-Bat. t. 197. 

A large, erect, semi-shrubby herb, 2-4 ft., stem and branches 
glabrous, young parts sparingly pubescent; 1. 3-foliolate, 
petiole 14-24 in., glabrous, stip. 0, lflts. on very short stalks, 
14-1 in., oval, acute at base, obtuse and mucronate at apex, 
thin, glabrous on both sides; fl. large, on ped. $—# in. long, 
distantly arranged in very lax terminal racemes, bracts small, 
acicular; cal. glabrous, segm. narrowly triangular, acuminate ; 
pet. much exserted, the keel very long-beaked ; pod about 2 
in. on a stiff slender stalk 1 in. long, oblong, cylindrical, tipped 
with hooked style-base, glabrous, brown; seeds 20 or more. 

Low country, very common. FI. Feb.-May; bright pale yellow, keel 
tinged with purple. 

Also in S. India, Malaya, and the Philippines. 

Readily known by the very long stalk to the pod. The name “ Yak- 


bériya” is applied to all the species with the pods much inflated, and 
which therefore “ pop” when squeezed. 


23. ©. quinquefolia, /. SP. P/. 716 (1753). 
Thw. Enum. 82. C. P. 1272. 
Fl. B. Ind. ii. 84. Rheede, Hort. Malab. ix. t. 28. 


20 Legumtnose. [Parochetus. 


An erect annual herb, 1-2 ft., stem slightly branched, 
striate, glabrous; |. 3- or 5-foliolate, petiole short, }-? in., 
glabrous, stip. very small, linear, lflts. 1-14 in., subsessile, 
oblong-linear, tapering to base, obtuse, thin, glabrous, paler 
beneath; fl. moderate-sized, on erect slender ped., arranged in 
very lax terminal racemes of variable length, bracts small, 
lanceolate, acuminate; cal. glabrous, segm. narrowly triangular, 
acute; pet. not exserted, about twice as long as cal.; pod 
about 2 in., spreading, oblong-ovoid, tapering into short stalk, 
tipped with nearly straight style-base, glabrous, veiny; seeds 
25 or more. 


Dry region, in swampy places; rare. Bintenne; Chilaw; near 
Jaffna. FI. Jan., Feb.; yellow. 
Also in India, Burma, Malaya, and the Philippines. 


4. PAROCHETUS, Ham. 


A prostrate, creeping herb, |. 3-foliolate, fl. solitary, 
axillary; cal.-tube campanulate, upper segm. connate; wing 
and keel-pet. equal, shorter than standard; stam. diadelphous, 
anth. uniform; ov. sessile, style glabrous, stigma terminal; 
pod straight, many-seeded.—Monotypic. 


P. communis, Hamilt. in Don Prod. Fl. Nep. 240 (1825). 

Thw. Enum. 82. C. P. 24009. 

Fl. B. Ind. 1. 86. Wight, Ic. t. 483 (P. major). 

A creeping herb, stem very long, prostrate, slender and 
thread-like, slightly branched, glabrous or nearly so, rooting 
at the distant nodes; 1. distant, 3-foliolate, petiole erect, 3-6 
in. long, very slender, with scattered hairs, stip. ovate, acute, 
membranous, lflts. $-? in., very shortly stalked, broadly 
obovate, cuneate at base, truncate and emarginate at apex, 
thin, glabrous above, slightly hairy and paler beneath; fi. 
solitary, 2 in., peduncle axillary, about as long as petiole or 
longer, erect, slender, jointed about #in. below the fl.; cal. }in., 
slightly hairy, segm. acuminate; pod ? in., oblong-linear, 
tapering to apex, compressed, glabrous, dark brown; seeds 
about 15, dull, dark brown. 

Montane zone, from 4000 ft. upwards, common in wet places. . 

Fl. Jan., Feb. ; violet-blue. 

Also in mountains of India, Burma, Java, and E. Trop. Africa. 


A very pretty little plant; the leaves are remarkably like those of an 
Oxalis. The only native Trefoil. 


Trifolium repens, L., the common white or Dutch clover, is completely 
established on roadsides about Nuwara Eliya; and Z. smznus, Sm., the 
small yellow Trefoil, and 7. arvense, L., the Haresfoot Trefoil, also occur. 
No doubt all were introduced with grass seed. 


Indigofera.] Leguminose. 21 


Meililotus parvifiora, Desf., is also met with as a casual, probably 
introduced with grain. There is a drawing of this in Hermann’s collec- 
tion, and it is n. 552 of the Fl. Zeylanica and the 77zfolium Melilotus 
indica of Linnzus. 


5. INDIGOFERA, J. 

Herbs or shrubs, usually more or less clothed with 
adpressed silvery hairs attached by their centre; |. usually 
imparipinnate, rarely simple; fl. small, in axillary racemes, 
Bemeertaccly “Solitary; call small, tube more or less 
campanulate, segm. nearly equal or the lower ones rather 
longer, usually narrow; pet. nearly equal, keel-pet. connate, 
produced at base into a spur, not beaked; stam. diadelphous, 
anth. uniform; style short, stigma capitate; pod usually linear 
(rarely oblong, falcate or globose) cylindrical, usually many- 
seeded.—Sp. about 300; 40 in FV. B. Ind. 


Leaves simple; pod 1-seeded. 


Pod sickle-shaped, spiny 1. I. ECHINATA. 
Pod globose, not spiny E 2, I. LINIFOLIA. 
L. imparipinnate; pod 2- or more- seeded. 
Pod 2-seeded_ . : : : é : . 3. I. ENNEAPHYLLA. 
Pod 6-12-seeded. 
Fl. solitary P ; : é : . 4. I. ASPALATHOIDES, 
Fl. in greaty racemes. 
Lfts. 3-1 
Fl. few (2-8) in raceme. 
Pftss.5, |: ; : : 5 : . 5. I. GLABRA. 
Lfts. 7 or more. 
Branches nearly glabrous : . 6. I. TENUIFOLIA. 
Branches covered with glandular hairs 7. I. VISCOSA. 
BATS: 3°: 5 iene: I, TRIFOLIATA. 
F]. numerous in raceme. 
Lfits. 3 or 5. 
Pod not torulose. 
Pod straight, quadrangular . Oo) LeTRIDA: 
Pod curved, cylindrical B 10. I. SUBULATA. 
Pod torulose. 5 = - II. I, PAUCIFOLIA. 
Lits, 7,.9, OL PI. 
Pod not torulose, 8-12-seeded. 
Pod very hairy . ; : 12. I. HIRSUTA. 
Pod nearly glabrous. , 13. I. TINCTORIA. 
Pod torulose 3-6-seeded . : 14. I. CONSTRICTA. 
Lfits. 15-23. 
Lfits. 15-19, thick; pod 1} in.. : 15. I. WIGHTII. 
Lflts. 19-23, thin; pod2in. . ; 16. I. GALEGOIDES. 


I. I. echinata, W7//d. Sp. Pi. iii. 1222 (1800). 

Herm. Mus. 31. Burm. Thes. 113. FI. Zeyl. n. 288. Hedysarum 
nummularifolium, L. Sp. Pl. 746.  Acanthonotus echinatus, Benth., 
Thw. Enuin. 83. C.P. 1453. 

Fl. B, Ind. ii. 92. Wight, Ic. t. 316. 


22 Leguminosae. [Undigofera. 


Annual, stems numerous, prostrate, spreading, 6-12 in. 
long, slender, 2-edged, branched, nearly glabrous; 1. simple, 
4-3 in., very shortly stalked, obovate-rotund, very obtuse, api- 
culate, with a few small scattered white hairs on both sides, 
stip. linear, filiform ; fl. very small, on very short ped., 3-8 
together in axillary racemes slightly exceeding I|., bracts aci- 
cular; cal.-segm. setaceous, bristly; pod + in., compressed, 
semilunar, tipped with sharp style-base, the upper margin 

_ slightly curved, the lower very much so, and set with several 
rows of divaricate hooked spines, 1-seeded. 
: ees region, in rocky and sandy places; common. Fl. Dec._March ; 
Ink. 
H Also in Peninsular India and in W. Trop. Africa. 


Linnzus’ name is often erroneously referred to Alysstcarpus nummu- 
larifolius, DC. (as in Fl. B. Ind. ii. 158, following other writers). 


2. I. linifolia, Retz. Obs. Bot. iv. 29 (1786). 

Thw. Enum. 83. C. P. 3514. 

BIBS ind. i592. sWirhbelest.313: 

Annual, stems very numerous, 1-2 ft. long, prostrate or 
ascending, much branched, slender, 2- salpec, white with ad- 
pressed silvery hair ; 1. numerous, simple, 4 4-3 in., linear, acute 
at both ends, strongly mucronate, silvery with adpressed hair 
on both surfaces, stip. setaceous ; fl. very small, on short ped., 
3-8 together in very numerous short axillary racemes; cal. 
very deeply cut, segm. setaceous ; pod very small, size of a 
mustard-seed, globular, apiculate, white and glistening with 
adpressed hair, 1-seeded. 

Dry country; rare. Kurundu-oya, on the banks of the Mahaweli 


River; Kantalai. Fl. May, August; very pale pink. 
Throughout the E. Tropics, also in Afghanistan and Abyssinia. 


3. I. enneaphylia, L. Mani. ii. 571 (1771). Cheppunerinchi, 7. 

Moon Cat. 54. Thw. Enum. 83. C.P. 2775. 

Fl. B. Ind. 11.94. Wight, Ic. t. 403 (bad). 

Annual or perennial, root-stock often woody; stems 
numerous, trailing, prostrate but not rooting, much ee 
sparingly strigose with white adpressed hair ; |. small, $—#in., 
imparipinnate, stip. rather large, lanceolate, acuminate, Iflts. 
7-9 (rarely 11), nearly sessile, alternate, minute, narrowly 
obovate, cuneate at base, retuse and apiculate at apex, stiff; 
fl. very small, nearly sessile, 10-20 crowded in a short close 
spicate head, peduncle short, bracts lanceolate, acuminate, 
scarious ; cal. set with long stiff white hairs, segm. very long, 
setaceous ; pod very small, } in., ovoid-oblong, with a sharp 
apiculus, more or less clothed with white adpressed hair, 
2-seeded with a partition between the seeds. 


Indigofera. Leguminosae. 2 
> fo 


Dry districts and on the seashore; common. FI. all the year ; bright 
red. 

Throughout the E. Tropics and in Angola, W. Trop. Africa. 

Found often in the hottest and most barren places, where it becomes 
curiously compact and stunted. 


4. I. aspalathoides, Vahl, ex DC. Prod. ii. 231 (1825). Chiv- 
anarvempu, 7. Rat-kohomba, 5. 

Herm. Mus. 34. Burm. Thes. 89. Fl. Zeyl. n. 271. Aspalathus in- 
dicus, L. Sp. Pl. 712; Moon Cat. 52. Thw. Enum. 83. C. P. 1455. 

Fl, B. Ind. ii. 94. Wight, Ic. t. 332 (not good). 

A low shrub, much branched, the branches divaricately 
spreading, rigid, the young twigs snowy white with a felt of 
copious short hair which is deciduous and shows the branches 
glabrous, shining, and purple; |. minute, sessile, crowded on 
the twigs, | but soon caducous, sessile, digitate, lflts. 1-5, usually 
3, about +5 in. long, sessile, linear, oer involute, with a ny 
large, white, adpressed hairs, rather fleshy ; fl. solitary, 4 
on slender axillary ped. longer than 1.; cal.-segm. linear ; oe 
about $in., narrowly linear, apiculate, straight, turgid, nearly 
glabrous, pale brown; seeds about 8 with partitions between 
them. 

Dry region; rare. Jaffna, very abundant on sandy flats ; Batticaloa. 
Fl. Nov.—Feb.; dark pink. 

Also in the Carnatic plains of S. India. 

Used for making brooms at Jaffna. It is also a favourite medicine 
among the Tamils. The S. name given is that of the drug as sold in the 
bazaars. Hermann (Mus. 34) gives it, and probably obtained his speci- 
mens from that source, as it grows nowhere in the S. of the island. 

Very different in habit from all our other species ; the Iflts. look like 
simple leaves in fascicles. 


5. Z. glabra, LZ. SZ. Pl. 751 (1753). 

Herm. Mus. 34. Burm. ae 39. FI. Zeyl. n. 274. L. pentaphylla, 
ce fuw. Enum. 411. C. P. 

"FL B. Ind. ii. 95 U. Fa TN Wight, Ic. t. 385. 


Annual, 1-3 ft., with numerous, very slender, ascending, gla- 
brous branches; |. imparipinnate, rachis #-1 in., stip. setaceous, 
persistent, Iflts. usually 5 (2 pair and end one), very shortly 
stalked, 4—? in., obovate-oval, obtuse, thin, slightly hairy ; fl. 
small, on slender ped., 2-4 in. short axillary racemes; cal.- 
segm. long, setaceous; pod about # in., narrowly linear, 
straight, cylindrical, apiculate, glabrous ; seeds about 12 with 
partitions between. 

Moist low country; very rare. I have seen, besides Hermann’s, only 
the C. P. specimens, collected at Colombo by Ferguson in 1859, which 
are very imperfect. fl. red. 

Occurs throughout India and in Trop. Africa. 


Of Linnzus’ duplicate names /. glabra has 21 years’ priority over 
I. pentaphylla, but neither name is very appropriate. 


24 Leguminosae. [ Indigofera. 


6. X. tenuifolia, ARott/. ex W. and A. Prod. 200 (1834). 

Thw. Enum. 83. C. P. 1462. 

Fl. B. Ind. it. 95. 

Annual, 6-18 in. high, with numerous spreading or as- 
cending, slender, slightly strigose branches ; |. imparipinnate, 
rachis 4-2 in., strigose, stip. small, setaceous, Iflts. usually 9 
(4 pair and end one), very shortly stalked, obovate-linear, with 
few coarse adpressed hairs on both sides, stipel minute, 
setaceous ; fl. small, shortly stalked, 3-8 in axillary racemes 
usually longer than lL, cal.-segm. setaceous ; pod nearly 1 in., 
linear, compressed, straight, with scattered adpressed hair, 
rather torulose, about 10-seeded. 

Dry region; rare. Trincomalie; Bintenne; Bibile, Uva. Fl. Jan.; 


red. 
Also in S. India. 


7. £. viscosa, Lam. Encycl. iil. 247 (1789). 

Thw. Enum. 83. C. P. 1459. : 

Fl. B. Ind. ii. 95. Wight, Ic. 404 (not good). 

Annual, but often semi-woody below, 1-3 ft., branches 
numerous, spreading, densely clothed with gland-tipped hairs, 
so as to be very viscous to the touch; |. imparipinnate, rachis 
about 1in., slender, very viscous, with glandular hair, stip. 
filiform-setaceous, Iflts. 7-13 (3-6 pair and end one), usually 
11, obovate-oval, obtuse, apiculate, nearly sessile, finely hairy 
above, with larger adpressed hairs beneath ; fl. small, very 
shortly stalked, 3-6 in glandular racemes rather shorter than 
1.; cal.-segm. filiform, setaceous ; pod 4—#in., linear, straight, 
cylindrical, torulose, apiculate, clothed with both adpressed 
white hairs and copious gland-tipped ones; seeds 10-12. 

Dry region ; very common in open places. Fl. May-Sept.; pink. 

Throughout the Tropics of Asia, Africa, and Australia. 


8. I. trifoliata, Z. Amen. Acad. iv. 327 (1759). 

Thw. Enum. 83. C. P. 3592. 

1Pik, 1835 Iievel, ti, Clee Winednie, Mes te, Suits 

Herbaceous, 1-2 ft., branches few, slender, elongated, with 
scattered, adpressed white hairs; |. 3-foliolate, petiole £3 in., 
slender, stip. very small, setaceous, lflts. $-1in., shortly stalked, 
narrowly obovate-oval, tapering to base, obtuse, apiculate, 
nearly glabrous above, glaucous with white adpressed hair 
and with scattered glandular dots beneath ; fl. small, on very 
short, curved ped., 3-6 crowded in very short sessile racemes; 
cal.-segm. lanceolate, acuminate; pod 4—#in., deflexed, linear, 
straight, somewhat quadrangular, with 4 narrow longitudinal 
wings, one on either side of each suture, not torulose, with a 
few scattered white hairs, 6—8-seeded. 


Indigofera.) Leguminose. 25 


Dry or intermediate regions; rare. Mineri; near Badulla. Fl. Dec.— 
March ; dark pink. 

Throughout India, and in Java, China, N. Australia. 

There are specimens in Hb. Perad. without locality which are probably 
I. vestita, Baker, of S. India. 

g. £. trita, LZ. 6 Supp. Pl. 335 (1781). Wal-awari, S. 

Thw. Enum. 83. C.P 1463, 

Fl. B. Ind. ii. 96. Wight, Ic. tt. 315, 386. 

A low, coarse oedensiaae. 1-2 ft. high, branches numerous, 
rigid, divaricate, hoary with adpressed white hair; 1. 3-folio- 
late, petiole 4-3 in, stip. small, setaceous, Iflts. narrowly 
obovate-oblong, the end one :stalked, the lat. ones nearly 
sessile, usually slightly emarginate, dark green and glabrous 
above, whitish and finely hairy beneath, stipels minute; fl. 
rather small, 3 in., numerous, nearly sessile, in spicate racemes 
shorter than 1|.; cal.-segm. narrow, acuminate; wing- and keel- 
pet. very long, the former emarginate at apex; pod 1-1}in., 
linear, divaricate, rigid, straight, quadrangular, spine-pointed, 
not torulose, purplish-silvery, with adpressed hairs ; seeds 8. 

Dry region ; rather common in damp waste ground. Trincomalie ; 
Batticaloa; several. places bet. Anuradhapura and Trincomalie; Ata- 
kalan Korale. Fl. May; salmon-red. 

In most countries of Trop. Asia, N. Australia. 

The fi. at the top of the racemes do not produce pods, and the end of 
the rachis in fruit becomes rigid and spine-pointed. 


Io. I. subulata, Por. Encycl. Meth. Supp. iii. 150 (1813). 

tL. flaccida, Koen., Thw. Enum. 83. C. P. 1460. 

Fi. B. Ind. ii. 96. Wight, Ic. t. 387 (Z flaccida). 

Perennial, with very long, slender, sub-scandent stems, 
branches numerous, ascending or spreading, striate, nearly 
glabrous, when young with white adpressed hair ; |. impari- 
pinnate, rachis 1-2 in., slender, hoary, stip. filiform, lflts. 5 (2 
pair and end one), stalked, readily disarticulating, oval, obtuse, 
apiculate, hairy on both sides, paler beneath; fl. small, on 
short, strongly curved ped., numerous, in very long-stalked, 
slender, spicate racemes, exceeding the 1.; cal.-segm. setaceous ; 
pod 14-1#in., linear, slender, deflexed at base, curved out- 
wards, and divaricate, sharp-pointed, subquadrangular, nearly 
glabrous, not torulose; seeds 8-12. 

Dry region, scrambling over bushes; rather rare. Trincomalie; 
Haragama ; Anuradhapura. Fl. December-Feb.; pale purplish-red. 

Found in S. India and in Trop. Africa and America. 

Easily known by its straggling sub-scandent habit. The racemes 


greatly elongate after flowering, frequently reaching 9 in., the pods being 
mostly found i in the lower half. 


il. I. paucifolia, Del. Descr. Egypte, 251 (1812). Nanti, 7. 
Thw. Enum. 83. C. P. 1454. 
Fl, B. Ind. ii. 97. Wight, Ic. t. 331. 


26 Leguminose. (Indigofera. 


A small shrub, 1-3 ft., with numerous stout, woody, as- 
cending branches, more or less densely hoary with small, 
white adpressed hair, young parts white, silky; | imparipin- 
nate, rachis 4-4in., stip. lanceolate, acuminate, Iflts. 3-5, 
rarely 7, alternate, very shortly stalked, 2-1in., narrowly 
oblong-lanceolate, shortly acuminate, slightly hairy above, 
white with dense pubescence beneath, conduplicate when 
young, terminal one largest; fl. small, numerous, on short, 
curved ped., racemes spicate, rather close, longer than 1. cal.- 
segm. lanceolate, cuspidate ; pods numerous on whole length 
of raceme, deflexed, 4—# in., slightly curved outwards, apiculate, 
torulose, hoary; seeds about 6. 

Dry and desert regions; rather common. Jaffna, abundant; Aripo ; 
Kalpitiya and Karativu I., abundant; Matalan, N. Prov. (Nevill). Fl. 
Feb., March, August ; red. 


Also occurs in Trop. Africa, Arabia, Beluchistan, Sind, and in Java. 
Mr. Nevill gives me as the Tamil name of this ‘ Kuttukarasamatti.’ 


I. parviflora, Heyne. A scrap in the Peradeniya collection collected 
by Ferguson in 1865 at Colombo may be this species, but is too imper- 
fect for certain identification. 


12. £. hirsuta, Z. Sf. P/. 751 (1753). 

Herm. Mus. to. Burm. Thes. 37. FI. Zeyl. n. 272. Moon Cat. 54. 
Thw. Enum. 83. C. P. 1456. 

Fl. B. Ind. 11. 98. Burm. Thes. t. 14. 

Annual, 1-3 ft., erect, slightly branched, branches hori- 
zontal, covered (as is whole plant) with spreading reddish 
hair ; 1. imparipinnate, rachis 14-2 in., stip. 2 in., filiform, con- 
spicuous, lflts. 7 (3 pair and end one), shortly stalked, #-1 in., 
oblong-oval, very hairy on both sides, especially beneath ; fi. 
numerous, on short curved ped., racemes long-stalked, rather 
dense, greatly exceeding 1. cal.-segm. long, filiform, very 
hairy; pods 2in., numerous, deflexed, overlapping, oblong- 
linear, somewhat quadrangular, apiculate, densely clothed with 
spreading hair; seeds 8. 

Dry and intermediate regions; very common. Fl. March-Aug.; 
bright pink. 

In nearly all Tropical countries. 

Hermann’s name for this is ‘ Ghasundupjali’ (Gas-undupiyali). 


13. * I. tinctoria, Z. Sd. P/. 751 (1753). Nil-awari, S. 

Herm. Mus. 32. Burm. Thes. 69. FI. Zeyl. n. 273. Moon Cat. 54. 
Thw. Enum. 411. C. P. 3591. 

Fl. B. Ind. ii. 99. Wight, Ic. t. 365. Bentl. and Trim. Med. Pl. t. 72. 


A low shrub, 2-4 ft.. with numerous virgate, slightly 
angular branches, more or less covered with white adpressed 
hairs ; |. imparipinnate, rachis 14-2 in., adpressed-pilose, stip. 


Indigofera.] Leguminose. De) 


small, linear-cuspidate, lflts. 7-11 (3-5 pair and end one), 
shortly stalked with minute stipels, about #in., obovate- 
oblong, acute at base, rounded, often retuse, apiculate, glabrous 
above, silvery-silky beneath, bright green, but drying a 
blackish-grey, readily disarticulating; fl. small, on short, 
slender ped., in erect, rather lax, tapering spicate racemes, 
bracts small, cuspidate ; cal. very shallow, silvery-hairy, segm. 
lanceolate, acute; pod 1-1} in., linear, straight or slightly 
curved, cylindrical, strongly apiculate, thickened at both 
sutures, nearly glabrous, slightly or not at all constricted 
between the seeds; seeds 8-12. 


Low country, by roadsides and waste places ; rather common, especi- 
ally in the dry region, but very doubtfully indigenous. FI. all the year ; 
pink, standard green outside. 

Found also in India and Tropical Africa, but scarcely known as a 
wild plant anywhere, unless in West Trop. Africa. 

This is the Indigo plant, so largely cultivated in India. Here there 
is no cultivation, but the natives of Jaffna use the leaves of the wild plant 
in obtaining a black dye for cloth. 


I. Anil, L., is occasionally met with as a roadside weed. It is another 
indigo-producing species, and is generally believed to be of American 
origin. 


14. Z. constricta, 77zm. Cat. Ceylon Pl. 23 (1885). 
I. flaccida, var. B, constricta, Thw. Enum. 411. C. P. 3811. 
Fl. B. Ind. ii. 99 (note under (JZ. ¢2cforia). 


A small erect shrub, with few cylindrical branches, silvery 
with adpressed hair ; |. imparipinnate, rachis about 2 in., stip. 
short, stiffly setaceous, Iflts. 9, shortly stalked, about #in., oval 
or oblong-oval, acute at base, obtusely apiculate, finely hairy 
on both sides, especially beneath, easily falling ; fl, numerous, 
racemes about as long as l1.; cal. silky, tube campanulate, 
segm. short, triangular, acute; pod about 14in., linear, slightly 
curved, somewhat 4-angled, pointed, much constricted be- 
tween the seeds, sparingly silky; seeds 3-6, larger than in 
I. tinctoria. 

Montane zone ; very rare. Etanwelle, Matale East (Thwaites). Fl. 


June ; pet. not seen. 
Also in Peninsular India (N. Canara, Talbot n. 320 in Herb. Calcutta). 


15. I. Wightii, Grak. in Wall. Cat. n. 5488 (1828). 

I. tnamena, Thw. Enum. 83. C. P. 3513. 

Fl. B. Ind. ii. 99. 

A small shrub, with numerous spreading, furrowed, grey- 
silvery branches, young parts densely covered with adpressed 
silvery hair; |. imparipinnate, rachis 2-3 in., silvery, stip. very 
small, cuspidate, Iflts. 15-19 (7-9 pair and end one), small, 


28 Leguminosae. [ Psoralea. 


3in., shortly stalked, lanceolate-oblong, acute at base, obtuse, 
with a strongly curved mucro at apex, coriaceous, densely 
covered with adpressed white hair on both sides; fl.on slender 
ped., in nearly sessile crowded racemes much shorter than L; 
cal.-segm. lanceolate-linear; pet. very hairy outside; pod 
I}+in., stout, linear, cylindrical, mucronate, hairy, not torulose; 
seeds, 10-12. 

Low country very rare. Kalupahane near Haldummulla, at about 


2000 ft. (Thwaites). Fl. April, red. 
Also in S. India. 


16. I. galegoides, DC. Prod. ii. 225 (1825). 

Avro, 120, mi, ~ Ili, Iran, (C, IPs waleyt, 

Fl. B. Ind. ii. too. 

A tall shrub or little tree, twigs angular, glabrous, young 
parts silvery-pubescent; |. imparipinnate, rachis 3-44 in., 
channelled on upper side, pubescent, stip. filiform, lflts. 19-23 
(Q-II pair and end one), distinctly stalked, oblong, acute at 
base, obtuse, finely mucronate, silky on both sides, paler 
beneath, thin, stipels minute; fl. on short ped., in dense, erect, 
spicate racemes; cal. hairy, segm. short, triangular, acute ; 
pod 2in., linear, straight, with a long beak, nearly glabrous ; 
seeds 8-10, very truncate at ends. 

Moist low country; rather rare. Colombo (Ferguson) ; Welipenna, 
Pasdun Korale; Kaduganawa; Matale (Gardner). Fl. April-August ; 
pale pink. 

Also in E. Bengal, Malaya, and S. China. 

Often planted for ornament. 


6. PSORALEA, J. 


Annual herbs, dotted with small dark glands; |. simple; 
fl, in short, dense, axillary racemes; cal. cut about 4 way 
down, lowest segm. longest, keel-pet. obtuse, not beaked ; 
stam. diadelphous, but the upper one often partially connate 
with rest, anth. nearly uniform; style long; pod small, inde- 
hiscent, 1-seeded.—Sp. about 100; 2 in FZ. B. Lnd. 


P. corylifolia, L. SZ. Pl. 764 (1753). Bodi, S. 

Moon Cat. 55. Arn. Pug. 11. Thw. Enum. 84. C. P. 1443. 

Fl. B. Ind. 11. 103. Bot. Mag. t. 665. 

Annual, 1-3 ft., stem erect, with scattered hairs or nearly 
glabrous, dotted with glands, slightly branched ; 1. simple, on 
erect petioles 4-1 in. long, 14-2} in., roundish or broadly 
ovate, rounded at base, acute or subacute, irregularly repand- 
dentate, sparingly hairy on both sides, copiously sprinkled 


Mundulea.| Leguminose@. 29 


with black, glandular dots, veins prominent beneath, stip. 
triangular, acuminate; fl. small, on short ped., crowded in 
a dense spicate raceme #in. long, terminating a stiff, erect, 
axillary peduncle about 2 in. long, bracts membranous, acute, 
gland-dotted; cal. hairy, gland-dotted, segm. obtuse; pod 
zo in., enclosed in cal. and scarcely exceeding it, broadly 
ovoid, glabrous, black, indehiscent. 

Dry region; ratherrare. Anuradhapura; Nalande; Delft I. (Gardner); 
Mannar. Fl. Feb., Aug., Nov.; purple. 


Throughout Peninsular India. 
Arnott (I. c.) states that Ceylon specimens are sometimes 3-foliolate. 


Ceylon has no species of A/z//ettéa, of which 24 species are included in 
Fl. B. Ind. 


7, MUNDULEA, DC. 


Small tree; |. imparipinnate; fl. large, in terminal racemes; 
cal. deeply campanulate, segm. short, two upper connate; pet. 
about equal, standard with long claw, wing-pet. coherent with 
keel near the base; stam. monadelphous, anth. uniform; style 
long, curved; pod linear, flat, scarcely dehiscent, few-seeded.— 
Speswat in fY. Bb. Ind. 


IM. suberosa, Benth. Pl. Jungh. iii. 248 (18 54). 
Tephrosia suberosa, DC. Thw. Enum. 84. C. P. 1486. 
Fl. B. Ind. ii. 110. Wight, Il. t. 82. 


A small slender tree or erect shrub, 10-15 ft. high, with 
pale corky bark, young twigs densely pilose-velvety ; 1. 
numerous, rachis 4-6in., densely silky-pubescent, stip. small, 
linear, Iflts. 13-23 (6-11 pair and end one), the lower ones 
often alternate, shortly stalked, ?-14in., narrowly ovate-lan- 
ceolate, obtuse, apiculate, glabrous and bright green above, 
silky and silvery beneath; fl. large, on pilose ped. din. long, 
in rather dense terminal racemes; cal. silky, segm. broadly 
triangular, acute; pet. over #in., standard silky outside; pod 
2-3} in., linear, straight, rostrate, strongly compressed, densely 
velvety, with short, golden-brown hair, thickened at sutures ; 
seeds 2-7, about }in., yellowish-brown. 

Rocky hills in the dry region; rare. Dambulla Hill, abundant ; 
rocks near Habarane. FI. March, Sept., Nov.; bright pinkish-violet. 

Also in S. India, Trop. Africa, Madagascar. 

It is just possible that this pretty shrub may have been originally in- 


troduced to its few Ceylon localities. It is sometimes cultivated in 
gardens. 


30 Leguminose. [ Tephrosia. 


8. TEPHROSIA, ers. 


Perennial herbs or low shrubs, |. imparipinnate, with 
setaceous (rarely spinous) stip. and oblong Ifits., fl. in leaf- 
opposed racemes or few axillary; cal. campanulate, segm. 
nearly equal, acuminate; keel-pet. not beaked ; stam. diadel- 
phous, anth. uniform; style much curved, filiform or flat- 
tened ; pod linear, flat, dehiscent, many-seeded, without septa 
between the seeds.—Sp. 100; 10 in FZ. B. Ind. 


Stipules spinous’. : : : 5 : . 1, TL SPEINOSA: 
Stipules not spinous. 
Pod slightly pilose or silky. 
Lfits. unequal. 


FI. axillary 2. T. SENTICOSA. 
Fl. in racemes 3. T.. TINCLORIA 
Lfits. nearly equal. 
Seeds 5 0r6. 4. T. PURPUREA. 
Seeds 11-14 . 5. T. MAXIMA. 
Pod densely hairy. 
Pod about 2 in., nearly straight 6. T. HOOKERIANA. 
Pod. 17 in., distinctly falcate . 7. T. VILLOSA. 


1. T. spinosa, Pers. Syn. Pl. ii. 330 (1807). 
Thw. Enum. 411. 
FI. B. Ind. ii. 112. Wight, Ic. t. 372. 


A small spreading shrub with rigid divaricate branches, 
silvery-grey with adpressed hair; 1. small, rachis 4-2 in, 
pubescent, stip. sharply spinous, persistent, becoming woody, 
Iflts. 2-4 pair and end one, very shortly stalked, nearly equal, 
obovate-oblong, strongly mucronate, silky on both surfaces, 
more or less conduplicate ; fl. rather small, on slender ped., 
solitary or 2 or 3 in axils of 1.; cal. hairy, segm. sharply 
acuminate; pod 1-14 in, linear, slightly falcate, pendulous, 
mucronate, thinly silky, 5-7-seeded. 

Dry region ; very rare. Near Giant’s Tank, Mannar District, 1890. 
The locality given in Thw. Enum. is ‘ North of the Island, Mr. W. 
Ferguson ;’ and Mr. F. informed me that the specimens were sent to him 
from Aripo in 1859 by the late Dr. Charsley. FI. Jan._Feb.; pink. 

Also in S. India and Java. 

A common bazaar drug under the Tamil name of ‘ Mukavallivér.’ 


2. 'T.senticosa. Pers. Syn. Pl. ii. 330 (1807). 

Herm. Mus. 12 and 14. FI. Zeyl. n. 303. Cvacca senticosa, L. Sp. Pl. 
752. W.and A. Prod. 212. Trim. in Journ. Linn. Soc. xxiv. 148. 

Fl B. Ind.ai. 112. Wight, Ic. t. 370. 

A low shrub with numerous divaricate hoary branches ; 
Iflts. 1-3. pair and an end one (which is much the largest), 
I-2 in., obovate-oblong, emarginate, nearly glabrous above, 


Tephrosia.] Leguuinose. BT 


clothed with adpressed soft white hair beneath, stip. subulate ; 
fl. shortly stalked, solitary or 2 in the axils; pod 14-2 in, 
spreading, linear, much curved, sparingly pilose, 6—8-seeded. 

Very rare. I have seen no specimens but those in Hermann’s 
Herbarium; the pods of these are very sparingly pilose, but there are no 
flowers. The plant is perhaps only a cultivated state of 7. spzzosa ; some 
of Hermann’s specimens have 7-9 Ifits. 

Also in S. India. 


The above description mainly from Roxburgh (Galega pentaphylla, 
Fl. Ind. iii. 384). 


3. T. tinctoria, Pers. Syn. P/. ii. 329 (1807). Alu-pila, S. 

Herm. Mus. 34. FI. Zeyl. n. 302. Cvacca tinctoria, L. Sp. Pl. 752. 
Galega tinctoria, L., Moon Cat. 55. Thw. Enum. 84. C. P. 1449. 

Fl. B. Ind. ii. 111. Wight, Ic. t. 388. 


Perennial, somewhat shrubby below, 14-3 ft., stem and 
ascending branches stiff, slightly angled, more or less clothed 
with yellow or brownish pubescence ; 1.-rachis 1-2 in., pubes- 
cent, stip. small, triangular, acuminate, lflts. 7-11 (3-5 pair 
and an end one), unequal, decreasing in size towards base of 
], very variable in width, obtuse at both ends, often retuse at 
apex, mucronate, glabrous above, more or less densely silvery- 
silky beneath, lat. veins numerous, parallel, conspicuous ; fl. 
rather small, on slender, short, hairy ped., in short, erect 
racemes, on stiff hairy peduncles exceeding the 1. bracts 
setaceous, hairy; cal.-segm. long, setaceous, hairy; pod 
2-24 in., linear, very slightly curved upwards at end, apiculate, 
more or less densely covered with soft silky hair, seeds 7-10, 
distant. 


Var 6, pulcherrima, Laker in FI. B. Ind, 7. c. 


Lfits. 1-5, very unequal, the terminal one much the longest 
(3 in. or more) and often the only one present, more densely 
silky beneath and with lat. veins more prominent, stip. larger; 
pods more thickly velvety. 


Upper zone of moist low country and lower mountain zone, rather 
common; abundant in the patanas of Uva. Var. 6B, Matale. Fl. Jan., 
Feb.; bright pink. 

Also in Peninsular India. 

Very variable in foliage; when the terminal Iflt. is not larger than the 
rest, it is 7. zwtermedia, Grah. An inferior Indigo is obtained from this, 
and is or was in use by the Sinhalese (see Herm. Mus. 1. c.) 


Sa zi purpurea, Pers. Syn. Pi. ii. 329 (1807). Pila, S. Kolin- 
chi, 7. 
Burm. Thes. 77. Fl. Zeyl. n. 301. Cvracca purpurea, L. Sp. Plant. 
752. Galega purpurea, L., Moon Cat. 55. Thw. Enum. 84. C. P. 1445. 
Fl. B. Ind. ii. 112. Burm. Thes. t. 32. 


32 Leguminose. [ Tephrosia. 


A perennial herb, 1-2 ft. high, with numerous wide- 
spreading, glabrous or sparingly pilose branches; |. numerous, 
rachis 14-3 in., slender, flexuose, channelled above, pilose, 
stip. setaceous, lflts. 5-10 pair and end one, on pilose stalks, 
equal, narrowly cuneate-oblong, tapering to base, obtuse or 
retuse, mucronate, glabrous above, finely adpressed-silky be- 
neath ; fl, numerous, rather small, on slender ped., 2 or 3 
together in small, lax, narrow, erect, terminal racemes; cal. 
thinly silky, segm. long, acuminate; pod 14-1? in,, linear, 
slightly curved, mucronate, very thinly hairy, 5-6-seeded. 

Low country in both moist and dry regions, a very common weed. 
Fl. July, Nov., Dec.; purplish-pink. 

Found throughout the Tropics. 

There is a form with very small flowers, which seem to be that figured 


by Burman. 
A common village medicine for children. 


5. T. maxima, Pers. Syn. Pi. ii. 329 (1807). [PLATE XXVII.] 

Burm. Thes. 228. Fl. Zeyl. n. 300. Cvacca maxima, L. Sp. Pl. 752. 
Thw. Enum. 34. Galega maxima, L., Moon Cat. 55. C. P. 1444. 

BE Be nde rises purpurea, var.). Burm. Thes. t. 108, f. 2 (bad). 

Perennial, often prostrate, with many long, slender branches 
set with rather long spreading hair; |. distant, rachis 24-6 in., 
channelled above, with more or less long spreading hair, stip. 
cuspidate, deflexed or spreading, rigid, conspicuous, Iflts. 
stalked, in 4-6 pair and an end one, nearly equal, narrowly 
obovate-oblong, usually strongly emarginate, mucronate, nearly 
glabrous on both sides, lat. veins prominent beneath; fl. large, 
few, on slender ped., in lax, long-stalked, apparently terminal 
(? really axillary) racemes exceeding the |, cal. pubescent, 
segm. narrow, acuminate; pet. #in.; pod about 2% in., nar- 
rowly linear, slightly falcate, beaked, scantily pubescent or 
nearly glabrous ; seeds 11-14. 

Dry region in sandy ground liable to be overflowed ; rather common. 
Jaffna; Mineri; Kantalai. Fl. March—-Aug.; bright pale pink. 

Also in S. India. 

A very pretty plant with large rose-coloured flowers. I cannot agree 
with Baker, in Fl. B. Ind., in making this a variety of 7: purpurea. 

The fl. are sometimes solitary in the leaf axils. The stip. are de- 
veloped before the |., and are very conspicuous in the buds. 


6. T. Hookeriana, W. and A. Prod. 212 (1834). 

LT. hirta, Thw. Enum. 84. C. P. 2776. 

BEB Sinden 13: 

Perennial, erect, with ascending pubescent branches; 1. 
large, rachis about 3 in., pubescent, stip. linear, filiform, Iflts. 
in 7-9 pair and an end one, 1-14 in., shortly stalked, oblong- 


Sesbania. | Leguminosae. 33 


linear, obiuse, retuse, mucronate, glabrous above, white and 
shining with adpressed hair beneath; fl. rather large, racemes 
very long, lax, erect, spicate, terminal ; cal. very hairy, upper 
segm. broader ; style flattened ; pod 1#-2 in., linear, slightly 
falcate, weakly mucronate, very densely clothed with copious 
fulvous hair; seeds 6-10. 


Moist low country; very rare. The C. P. plants are labelled 
‘Colombo (Macrae);’ and Thw. also gives ‘Hantane (Gardner).’ FI. 
purplish-white. 

Also in India and Malacca. 


7. T. villosa, Pers. Syn. Pi. ii. 329 (1807). Bu-pila, S. 

Burm. Thes. 78. Fl. Zeyl. n. 299. Cvracca villosa, L. Sp. Pl. 752. 
Galega villosa, L., Moon Cat. 55. Thw. Enum. 84. C. P. 1447, 1446. 

Bee ind. ii. 113. Burm. Thes. t. 33. 


Perennial, semi-shrubby below, branches numerous, spread - 
ing, white with adpressed hair; 1. numerous, rachis 1-2 in., 
silky, stip. rigid, sometimes almost spinous, spreading, Ifits, in 
5-9 pair and an end one, distinctly stalked, narrowly obovate- 
oblong, obtuse, mucronate, nearly glabrous above, finely silky 
beneath; fl. rather small, on hairy ped., in clusters of 2 or 3, 
racemes lax, terminal, on short peduncles, bracts linear ; cal. 
densely covered with long soft hair, teeth setaceous, plumose; 
standard hairy on the back; pod about 14 in., broadly linear, 
distinctly falcate, bluntly apiculate, very thickly covered with 
dense spreading brownish hair; seeds 6-8. 

Dry region; very common. FI. all the year; pink, rarely white. 
Also in Peninsular India, Trop. Africa, and Mauritius. 

C. P. 1446 (from Jaffna) has the young leaves and inflorescence much 

more densely clothed with silvery hair; it is var. avgentfea, Thw. Enum. 


(T argentea, Pers.). Burman’s figure (I. c.) represents a white-flowered 
variety. 


9. SESBANTA, Pers. 


Annual or perennial, semi-woody, |. long, abruptly pinnate 
with numerous pairs of Iflts., fl. in axillary racemes; cal. 
campanulate, segm. shallow, equal; pet. equal, long-clawed, 
keel obtuse, not beaked ; stam. diadelphous, anth. uniform, 
style curved upwards, glabrous; pod very long and narrow, 
dehiscent, with distinct partitions between the numerous 
seeds.—Sp. 20; 4 in FV. B. Ind. 


Lfits. 20-30 ; pods pendulous, twisted he . I. S, HGYPTIACA, 
Lfits. 40-100 ; pods spreading or erect, not twisted , 2. S, ACULEATA. 


PART II, D 


34 Leguminose. [Sesbania. 


1. *S. egyptiaca, Pers. Syn. P/. ii. 316(1807). Chittakatti, 7. 
Moon Cat. 53. Thw. Enum. 84. C. P. 1512. 
PE Belnd) iris) Wightalest.32" 


A weedy tree-like herb 6-10 ft. high, with numerous 
‘spreading branches, nearly glabrous, without prickles, young 
parts silky; 1. numerous, rachis 24-4 in., slightly pilose, without 
prickles, furrowed above, swollen at base, stip. filiform, 
membranous, falling early; Iflts. in 10-15 pair, very shortly 
stalked, $—? in., oblong-linear, obtuse, apiculate, glabrous or 
nearly so, rather glaucous; fl. few, rather large, on very slender 
ped., in very lax, slender, pendulous, axillary, stalked racemes; 
cal. glabrous, segm. very shallow, acute; pod 8-9 in., pendulous, 
very narrowly linear, sharply beaked, usually somewhat 
twisted, weak, slightly torulose, seeds 30 or more, greenish- 
grey. 

Dry country; rare. Habarane (Gardner); Trincomalie (Glenie). 
Doubtfully native, but often seen in gardens. Fl. Nov., Dec., June; 


pale yellow, the standard dotted on the back with purple. 
Throughout the Tropics, but often cultivated. 


2. S. aculeata, Pers. Synz. P/. ii. 316 (1807). 

Burm. Thes. 93. Herm. Mus. 55. Fl. Zeyl. n. 2098, var. 6 (in part). 
LA schynomene Sesban, L. Sp. Pl. 714 (part). Moon Cat. 53. Thw. 
Enum. 84. C. P. 1513. 

FI. B. Ind. ii. 114. Burm. Thes. t. 41 (poor). 

A semi-woody annual, 1-3 ft. high, with numerous, more 
or less prickly, glabrous, ascending branches ; 1. ascending, 
rachis very long, 6-10 in., branch-like, stout, cylindrical, more 
or less set beneath with small hooked prickles, glabrous, stip. 
membranous, acuminate, soon falling, Iflts. very numerous, 
20-50 pair, shortly stalked, $-? in., linear or oblong-linear, 
obtuse, mucronate, glabrous; fl. as in the last but rather 
smaller; pod 6-9 in., spreading or erect, narrowly linear, 
slightly falcate, beaked, stiff not twisted, glabrous, slightly 
torulose. 


Var. B, sericea, Benth. in Thw. Enum. 441. C. P. 3850. 


Branches, rachis and Iflts. permanently silky beneath ; 
racemes with shorter peduncles. 


Low country especially in dry region; rather common. Var. £B, 
‘Colombo. Often a bad weed in paddy fields. FI. Jan., July; yellow, 
standard with red dots on the back. 

Throughout the E. Tropics. 

An unarmed variety is common in paddy fields with shorter erect 
straight pods always under 4 in. long and solitary and sessile in the 
axils, which is probably &. uliginosa, Roxb., or perhaps . procumbens, 
"Roxb. 

Under his 4. Sesban, Linnzus doubtless included also S. egyptiaca. 


Zornia.| Leguminose@. a5 


S. grandifiora, Pers. (Agati, Desv.) is often seen in native gardens, 
its large white or red flowers being very conspicuous. It is called 
Katuru-murunga in S., Akattz in T., and is figured in Bedd. FI. Sylv. 
t. 86. The leaves and flowers are eaten. 


Io. ZORNIA, Gel. 


Annual, |. compound, Iflts. 2, dotted; fl. in long lax spikes, 
each enclosed in a pair of large flat bracts; cal. minute, 
paleaceous, upper segm. connate; pet. much exserted ; stam. 
monadelphous, anth. dimorphic; pod jointed, easily separable 
into flattened, circular, muriculate, indehiscent, 1-seeded joints. 
—Sp. 10; 1 in FZ. B. Ind. 


Z. diphylla, Pers. Syn. P/. ii. 318 (1807). 
Herm. Mus. 18. Burm. Thes. 114. Fl. Zeyl. n. 291. Aedysarum 
diphyllum, L. Sp. Pl. 747. Z. angustifolia, Sm., Thw. Enum. 84. C. P. 


3598. 
Sete adi. 147. Rheede, Hort, Malab. ix te. 82, 83 

Annual, with many prostrate or ascending, glabrous or 
puberulous, wiry branches, 6-12 in. long; |. small, rachis 4-3 in., 
stip. large, lanceolate, acuminate, produced below into an 
acuminate appendage, Ifits. 2 (a single pair) very shortly stalked, 
linear-lanceolate or lanceolate, acute, often unequal-sided, 
glabrous, rigid, dotted with black glands beneath; fl. small, 
sessile, distant, each enclosed within a pair of ovate, acute, 
flat, reticulate-veined, leaf-like, persistent bracts, often ciliate, 
dotted like the 1., arranged in stiff, erect, very lax axillary 
spikes; pod either quite enclosed between or slightly exceeding 
bracts, about 4 in., composed of 2—5 compressed indehiscent 
joints set with numerous short straight spines. 

Var. B, conjugata, 77rim. Hedysarum conjugatum, Willd. Sp. PI. iii. 


1178. Z. zeylanensis, Pers. Syn. Pl. ii. 213. Moon Cat. 54. Z. conju- 
gata, Sm., Thw. Enum. 85. Burm. Thes. t. 50, f. 1. C. P. 3600. 


Lfits. 4 in., ovate ; pods larger, twice as long as bracts. 

Var. y, Walkeri, Laker. Z. Walkeri, Arn. Pug. 12; Thw. Enum. 85. 
C. P. 3599. 

Lfits. }—? in.; pods scarcely longer than bracts, reticulate- 
veined, without spines. 

Low country, especially dry region; common. Var. 6, Colombo. 
Var. y, Colombo ; Jaffna ; Badulla; Galagama; Hewahette. FI. all the 
year ; yellow. 

Found throughout the Tropics. Var. y is endemic, and has perhaps 
good claims to be kept up as a species. 

Herm. gives ‘Mahasweenna’ (Maha-aswenna) as the S. name. 

[Geissaspis cristata, W. and A., is given for Ceylon in FI. B. Ind. ii. 
141, but I know of no specimens from here in any Herb.] 


36 Leguminose. [ Smathia. 


11. STYLOSANTEHES, Sw. 


A dwarf, rigid under-shrub, |. pinnately 3-foliolate, with 
large stip., fl. few, in dense heads; cal. tubular, the lowest 
segm. longest; keel-pet. slightly beaked; stam. monadelphous, 
dimorphic ; style long, straight; pod of 1 or 2 flattened, 
wrinkled, indehiscent, 1-seeded joints—Sp. 15; 1in FZ B. Ind. 


S. mucronata, Wild. Sp. Pi. iii. 1166 (1800). Wal-nanu, S. 

Herm. Mus. 42. Burm. Thes. 226. Fl. Zeyl. n. 553. AHedysarum 
hamatum. L. Sp. Pl. ed. 2, 1057. Arachis fruticosa, Retz. Obs. v. 26. 
Moon Cat. 54. Thw. Enum. 84. C. P. 1451. 

Fl. B. Ind. ii. 148. Burm. Thes. t. 106, f. 2. Bedd. Ic. t. 294. 


A much-branched, dwarf, woody perennial, branches stiff, 
wiry, ascending, hairy; |. distant or crowded, rachis }—3in., 
stip. large, membranous, adnate to rachis for half its length 
and forming a sheath round stem and terminating in 2 
acicular spreading sharply spiny teeth, lflts. 3, shortly stalked, 
34 in., the terminal rather the largest, lanceolate, strongly 
mucronate, rigid, glabrous above, silky on the prominent 
veins below and marked with groups of conspicuous black 
dots ; fl. rather small, few, sessile in axils of leaf-like stipulate 
bracts and forming small close terminal heads or spikes; 
cal. membranous, splitting into laciniz in fruit; pod very 
short, concealed by persistent bracts, tipped by long curved 
beak, joints pubescent, with raised veins. 

Dry and desert region ; common in the most arid places. Fl. March, 
August; yellow. 

Also in Peninsular India, and in Malay Islands and Trop. and S. 


Africa. 
I doubt the Sinhalese name above given. 


12, SMITHIA,* Azton. 


Annual or perennial herbs, |. abruptly pinnate, the rachis 
ending in a bristle, with large stip., fl. axillary, racemose, or 
paniculate, with 2 persistent scarious bracts adpressed to cal.; 
cal. deeply divided into 2 lips which are either entire or lobed; 
stam. in 2 bundles of 5 each, anth. uniform; pod very small, 
enclosed in persistent cal., of 2-6, easily separable, seed-like, 
1-seeded joints.—Sp. about 30; 12 in Fd. B. Ind. 


* Commemorates Sir James Edward Smith, the purchaser of Linnzeus’ 
collections and founder and first NESS CSE of the Linnean Society. 
Died 1828. 


Smithia.] Leguminose. 37 


Cal. rigid, the lips entire, acute. 


Fl. in small axillary racemes 5 . : . I. S. SENSITIVA. 
Fl. solitary or2,in axils. . 2. S. GEMINIFLORA, 
Cal. membranous, the lips lobed ; fl. in panicles =) Go. BLANDA: 


I. S. sensitiva, Azt. Hort. Kew. iii. 496 (1789). 

C. P. 3946. 

Fl. B. Ind. 11. 148 (not given for Ceylon). Ait. Hort. Kew. t. 13. 

Annual with very slender much-branched, glabrous stems, 
1-14 ft. long; 1. small, rachis under 4 in., set with bristly 
hairs, stip. lanceolate, acute, Iflts. 3-6 pair, oblong-linear, 
obtuse, bristle-pointed, glabrous above, with rather long 
bristles on margin and midrib beneath; fl. small, on slender 
ped., 2-6 crowded at ends of small, stalked, erect, axillary 
racemes rather longer than 1.; cal. striate, somewhat bristly, 
the lips equal, acute, entire; joints of pod 4-6, papillose, pale 
brown. 


Low country in intermediate region; very rare. Kaluganga, Lagalla 
(Beckett) ; 3 wet places near Lenadore. Fl. Feb.; bright yellow. 

Also in India, Java, China, Trop. Africa, Madagascar. 

The leaves are excessively sensitive. 


2. S. conferta, S77. zz Rees Cyclop. xxxiil. (1816). 

Thw. Enum. 85. C. P. 2777. 

Fl. B. Ind. ii. 149. (S. gemznzflora, Roth. var.) 

Annual, 2-4 ft., branches long, virgate, straggling, smooth, 
with very long internodes below, |. small, rachis #-}in., bristly, 
stip. very large, scarious, brownish- red, learesclais. acute, with 
a long deflexed acuminate or lacerate auricle at base, Ifits. 4-8 
pair, $-4} in. oblong, obtuse, mucronate, glabrous above, 
strongly ciliate on margin and midrib beneath, fl. rather large, 
conspicuous, with 2 scarious bracts close beneath cal., solitary 
or 2, on slender ped., from axils of upper |. which are 
crowded at ends of branches, so as to form a congested head ; 
cal. scarious, rigid, lips entire, acute; pod very small, joints 3 
or 4, minute, wrinkled. 


Moist low country up to 3000 ft.; in wet places, rather common. 
Pasdun Korale; Hantane; Maturata; Dikwelle; Ella. Fl. Jan., July, 
September ; pale yellow. 

Also in India, Java, and Trop. Australia. 

This is doubtless the S. sensitiva of Moon Cat. 54. The leaves 
become deflexed and the leaflets close up quickly after gathering, or on 
being struck. 


3. S. blanda, Wall. Cat. n. 5669 (1828). 

S. paniculata, Arn. Pug. 12. Thw. Enum. 85. C. P. 58. 

FI. B. Ind. ii. 151. Wight, Ic. t. 986 (?). 

A perennial herb, 1-2 ft., erect or ascending, stems slightly 
branched, usually hispid, more rarely glabrous; 1. distant 


28 Leg UMLNOSE. [4 schynomene. 


rachis 4-1 in., generally hispid, stip. as in the last, lflts. 2-6 
pair, overlapping, nearly sessile, oval-oblong, obtuse, mucro- 
nate, glabrous on both sides; fl. rather large, on slender hispid 
ped., in large, lax, spreading, terminal panicles, bracts beneath 
cal. short, obtuse; cal. membranous, hispid, the upper lip 
broad, with 2 rounded lobes, the lower narrower with 3 segm., 
the central one acute; joints of pod 1-4, nearly globular, 
reticulated. 

Montane zone, 47000 ft., on patanas, common. 

ie September, December ; bright yellow, standard with two red spots 
at base. 

Also in the mountains of India. 

This is a variable plant, but I cannot distinguish any well-marked 
constant varieties. Wight’s figure above quoted is very unlike our plant. 

The lfits. are sensitive, and close up on being struck or gathered. 


13. HSCHYNOMENE, J. 


Large herbs; |. pinnate with very numerous Ifits., fl. few, 
in axillary clusters, with 2 bracts below each; cal. deeply 2- 
lipped, lips nearly entire or lobed ; pet. soon falling, keel not 
beaked; stam. in 2 bundles of 5 each, anth. uniform; style 
curved upwards; pod long-stalked, linear, compressed, shortly 
beaked, composed of 6-9 readily separable i-seeded in- 
dehiscent joints.—Sp. 30; 2 in FZ. B. Ind. 


Cal. glabrous ; joints of pod smooth . : . 1. ZL AINDIGR 
Cal. hispid; joints of pod strongly muriculate . 2. AZ. ASPERA. 


1. HE. indica, Z. Sp. P/. 713 (1753). Diya-siyambala, S. | 

Herm. Mus. 40. Burm. Thes. 120. Fl. Zeyl. in 551. . pumila, 
L. Sp. Pl. ed. 2, 1061. Moon Cat. 54. Thw. Enum. 85. C. P. 1515. 

SB yincliiets peat Gat 405. 

Annual, 1-2 ft., stem erect with ascending, cylindrical, 
slender, glabrous branches; 1. very shortly stalked, rachis 
stout 14-4 in., glabrous, stip. linear-lanceolate, acute, auricled 
at base, Ifits. very numerous, 40-70, alternate, sessile, small, 
zo-% in., set obliquely on the rachis and overlapping, linear, 
unequal-sided, obtuse, apiculate, glabrous, thin, the upper ones 
much the smallest; fl. few, on slender glandular ped., small, 
I-3, on axillary peduncles (or branches) much shorter than 
l.; cal. glabrous, lips very shortly toothed; pod 1-14 in, 
narrow, slightly curved, glabrous or very slightly muriculate, 
7-9-jointed. 

Dry region, in wet places, especially near tanks, common. FI. July— 


Nov., yellow. 
Throughout the Tropics of the Old World and in Japan. 


Ormocarpum.] Leguminose. 39 


Variable. The stem, petioles, and pedicels are often covered with 
stalked glands. .@. Jumila, L. is a form with acute leaflets, and more 
hispid pods. Mr. Nevill sends from Puttalam a dwarf prostrate state, no 


doubt from dry ground. 
The base of the stem is often swollen and pith-like, but to no such. 


extent as in @. aspera, 


2. HE. aspera, Z. SP. Pl. 713 (1753). Maha-diya-siyambala, 
S. Attuneddi, 7. 
Herm. Mus. 24, 41. Burm. Thes. 119. Fl. Zeyl. n. 298. Moon Cat. 


. Thw. Enum. 85. C. P. 1514. 

Fl. B. Ind. ii. 152. Wight, Ic. t. 299. 

Perennial, stem very stout, often horizontal below and 
floating, then erect 2-4 ft., glabrous, scarcely branched, usually 
with a hollow pith-cavity, structure very light and spongy; 1. 
large, rachis 4-6 in., stout, glabrous, stip. over 4 in., linear, 
acuminate with a rounded membranous ciliate auricle at base, 
Iflts. as in the last but still more numerous; fi. rather large, on 
hispid ped., 2-6 in axillary corymbose panicles much shorter 
than 1, bracts ovate, hispid; cal. hispid, upper lip rounded, 
lower 3-fid; pet. hairy outside; pod 3-4 in., over } in wide, 
nearly straight, woody, glabrous, 6- or 7-jointed, each joint 
with a group of sharp asperities in the centre. 
as region, in tanks and paddy fields, common. FI. December ; 
ellow. ; 
z Also in India, Malaya, and Trop. Africa. 

This is remarkable for the light, spongy character of the tissue of the 
stem (which in large plants attains 3 in. in diameter), whence it is often 
called the ‘ pith-plant’ by the English. It is the ‘Shola’ of Bengal, and 
from slices of the stem the well-known pith hats are made. A manufac- 
ture of similar hats was started in 1885 in the jail at Hambantota, in the 
neighbourhood of which place the plant is plentiful. 


14. ORMOCARPUM, Leauv. 


Shrubs, |. imparipinnate; fl. few, in axillary racemes; cal.- 
tube campanulate, upper segm. broader than lower ; keel-pet. 
much incurved, not beaked; stam. in 2 bundles of 5 each, 
anth. uniform; style much curved upwards; pod long-stalked, 
composed of few narrow, indehiscent, oval-oblong joints 
readily disarticulating.—Sp. 6; 1 in Fv. B. Lund. 

O. sennoides, DC. Prod. ii. 315 (1825). 

Hedysarum sennoides, Willd., Moon Cat. 54. Thw. Enum. 85. C. P. 


1438. 
Fl. B. Ind. ii. 152. Wight, Ic. t. 297. 


A small, rather straggling, branching shrub, with smooth 
pale bark and slender twigs; |. spreading, rachis 14-2} in., 
usually rough with minute prickles, stip. small, triangular, 


40 Leguminose. [Pycnospora. 


sharply acuminate, striate, persistent, lflts. 9-15, usually alter- 
nate, shortly stalked, oval-oblong, rounded at both ends, 
apiculate, glabrous, pale beneath; fl. few, rather small, 
nodding on slender, hispid ped., about 3 in small hispid and 
glandular racemes, much shorter than |.; cal. somewhat hispid- 
glandular, or nearly glabrous, segm. acute; pod 1-2 in., long- 
stalked, moniliform, beaked, much contracted between the 
oblong-ovoid joints, joints 1-4 longitudinally striate, muri- 
culate and viscous with glands. 

Dry region, rather common. Anuradhapura; Kekirawa; Tissa- 
maharama; Bibile. Moon’s locality is Matale; there are specimens from 
him in Herb. Mus. Brit., collected in 1819. FI. Dec.-April; dull yellow. 

Also in Peninsular India, Siam, Philippines, Polynesia, and Trop. 
Africa. 


A form with a very prickly pod was collected at Vigitapura, near 
Kalawewa Tank. 


15. ELELOTIS, DC. 


Annual, 1. 1-foliolate; fl. minute, in long axillary racemes; 
cal.-segm. long, setaceous; keel-pet. adherent to wing-pet.; 
stam. diadelphous, anth. uniform; pod of a single boat-shaped 
membranous joint, 1-seeded.—Monotypic. 


&. sororia, DC. Prod. ii. 348 (1825). 
Thw. Enum. 412. C. P. 3765. 
Fl. B. Ind. ii. 153. Burm. FI. Ind. t. 50, f. 2 (Glycine monophylla). 


A prostrate annual, stems long, slender, triquetrous, 
glabrous, very slightly winged at the angles; 1. simple, Ifit. 
small, rounded, much broader than long, deeply emarginate, 
glabrous above, hairy beneath, with two minute stipelle at 
base; petiole 4-2 in., stip. triangular, acute, striate, mem- 
branous; fl. shortly stalked, in numerous very long, filiform, 
pubescent, trailing axillary racemes; pod very small, of 1 joint 
zin. long, compressed, boat-shaped, glabrous, veined. 

Moist low country; very rare. Only collected near Colombo by 
Ferguson in 1862; but probably overlooked elsewhere. 

Also in Peninsular India. 

I have but little material for description. 


16. PYCNOSPORA, ?. Lr. 


Perennial, semi-shrubby, |. pinnately 3-foliolate; fl. small, 
in terminal racemes; cal. very deeply cut, upper segm. con- 
nate, lower longer; keel-pet. obtuse; stam. diadelphous, anth. 


Pseudarthria.] Leguminose. Al 


uniform; pod ovoid, not jointed, several-seeded, without 
partitions.—Monotypic. 


P. hedysaroides, 47. 7x W. and A. Prod. 197 (1834). 

Flemingia polysperma, Moon Cat. 54 (?). &. nervosa, W. and A,, 
Thw. Enum. 92. C. P. 279. 

BE B. Ind. 1.153. 

Perennial, with a woody base and numerous slender 
prostrate or ascending stems with adpressed pubescence; 
l-rachis about } in., stip. lanceolate, long-acuminate, hairy, 
deciduous, lfits. shortly stalked, with filiform stipels, }-1 in., 
the terminal the largest, obovate-oval, obtuse at base, rounded 
at apex, pubescent on both sides, especially beneath, which is 
paler and with prominent veins; fl. small, on slender ped., in 
lax, narrow, erect, terminal racemes, bracts ovate, strongly 
acuminate, densely silky, falling before buds open; cal. hairy, 
segm. acuminate; pod small, }-in., ovoid, straight along 
dorsal side, turgid, apiculate, glabrous, veined, dark brown; 
6—10-seeded. 

Low country and up to 4ooo ft.; rather common, abundant in the 
lower hills of Uva; absent in much of the dry region. Fl. Feb.—May, 
pale purplish. 

Also in India, China, Philippines, and North Australia. 


17, PSEUDARTHRIA, VW. and A. 


Herbaceous, |. pinnately 3-foliolate; fl. in elongated race- 
mose panicles; cal. minute, upper segm. shorter, subconnate; 
keel-pet. obtuse; stam. diadelphous, anth. uniform; pod 
linear-oblong, compressed, indehiscent, 4-6-seeded, without 
partitions, not jointed.—Sp. 5; 1 in FZ. B. Ind. 

Scarcely distinct from Desmodium. 


PR. viscida, WV. and A. Prod. 209 (1834). Gas-gonika, 5S. 

Herm. Mus. 36. Burm. Thes. 187. Fl. Zeyl. n. 295. Hedysarum 
viscidum, L. Sp. Pl. 747. Glycine viscida, Willd., Moon Cat. 53; Thw. 
Enum. 87. C. P. 1432. 

Fl. B. Ind. ii. 154. Burm. Thes. t. 84, f. 1. Wight, Ic. t. 286. 

A large perennial (?) herb, 2-4 ft., erect, with numerous, 
more or less hairy branches; |. long-stalked, rachis 2-4 in., 
pubescent or hairy, stip. linear, Iflts, on short stalks, stipellate, 
the terminal 2—3}4in., the lateral 1-24in. and somewhat un- 
equal-sided, all rhomboid-oval, acute, pubescent above, densely 
white-silky beneath; fl. small, very numerous, on slender, 
spreading ped., arranged in fascicles on numerous lax, elon- 
gated and axillary racemes or panicles, bracts acicular, shorter 


42 Leguminose. [ Uraria. 


than ped.; cal. hairy, segm. acute; pod linear-oblong, about 
2in., truncate, apiculate, strongly compressed, flat, viscous- 
pubescent, ie: seeded, slightly indented along the edges 
between the seeds. 


Moist low country to 3000 ft.,a common weed. FI. Jan., Feb.; bright 
pink. 

Also in Peninsular India and Timor. 

The flat viscid pods attach themselves to clothing or animals, and are 
thus transferred to other places. 


18. URARIA, Desv. 


Semi-shrubby perennials, 1. imparipinnate, sometimes I- 
foliolate; fl. numerous, in long terminal racemes; cal. very 
deeply cleft, segm. nearly equal or lower longer; keel-pet. 
obtuse; stam. diadelphous, anth. uniform; pod composed of 
2-7 very small, indehiscent, 1-seeded joints, usually folded 
face to face on one another.—Sp. 8; all in FZ B. Lnd. 


Leaflets 5-7, linear . ; i ‘ : : . UU PIGEs 
Leaflets 1-3, broadly oval : : : : 5 . 2. U. HAMOSA. 


I. U. picta, Desy. Journ. Bot. iii, 123 (1813). 

Ghw.Enume 5.) 9251437. 

FL B. Ind. ii. 155. 

A small erect, semi-shrubby perennial, stem slightly 
branched, harshly pubescent; 1. rather large, rachis 3-5 in., pu- 
bescent, stip. $—-} in., triangular, acuminate, striate, lflts. 5-7 (2 
or 3 pair and end one), very shortly stalked, with filiform stipels, 
24-4 in., linear, rounded at base, slightly tapering to acute 
apex, glabrous above, and usually mottled with white along 
both sides of midrib, scabrous-pubescent, paler and with pro- 
minent veins beneath; fl. small, on hispid ped., arranged in 
fascicles in a dense, erect, spicate raceme 6-Qin. long, bracts 
large, lanceolate, acuminate, striate, strongly ciliate, completely 
concealing the buds but soon falling; cal.-segm. plumose with 
long hair, lower acuminate; pod of 2-4 ovoid, turgid, white, 
polished, seed-like joints. 

Dry region in grassy places; ratherrare. Mineri and Topare Tanks; 
Kalawewa; Batticaloa district; near Bibile. Fl. Dec., March. 

Also in India, Malay Is., Philippines, and Trop. Africa. 

In fruit the pedicels are stiffly curved upwards and inwards, so that 


the pods are in contact with the rachis, and the persistent calyx-segm. 
become rigid. 


[U. crinita, Desv. is given for Ceylon in FI. B. Ind. It is only here 
under cultivation in gardens. ] 


Alyssicarpus.] Leguminose. 43 


2. U. hamosa, Wall. Caz. n. 5681 B. (1828). 

Thw. Enum. 85. C.P. 3590. 

Fl. B. Ind. ii. 156. Wight, Ic. t. 284. 

A small semi-shrubby perennial with slightly pubescent 
branches ; |.-rachis about I in., hairy, stip. acuminate, hairy, 
Ifits. I or 3, the terminal 2-33 in., the lateral (when present) 
much smaller, all stipellate, shortly stalked, broadly oblong- 
oval or subrotund, nearly glabrous above, paler and silky with 
prominent veins beneath ; fl. smal], on shaggy ped., arranged 
in very lax, elongated, spicate, terminal racemes, bracts ovate, 
acuminate, woolly; cal. pubescent, lower segm. not elongated ; 
pod of 4-7 flattened, glabrous, slightly wrinkled, greyish- 
brown joints. 


Low country; rare. Badulla; Nilgala; Gampola. FI]. Feb., March. 
Also in India, Burma, and the Malay Archipelago. 


19. ALYSSICARPUWUS, Weck. 


Annual, semi-woody at base with many wiry diffuse stems; 
|. 1-foliolate, thick, with large scarious stip., fl. in leaf-opposed 
or terminal racemes or spikes ; cal. deeply cleft with narrow 
glumaceous segm.; keel-pet. obtuse, adnate to wings ; stam. 
diadelphous, anth. uniform, pod turgid, cylindrical, usually 
moniliform, composed of few indehiscent 1I-seeded joints.— 
Sp. 15; 9 in FZ. B. Ind. 


Pod pubescent, moniliform . : : . I. A. MONILIFER. 
Pod glabrous, or very nearly so. 
Pod not at all moniliform . ; 3 . 2. A. VAGINALIS. 


Pod more or less moniliform. 


oints quite even : . A. BUPLEURIFOLIUS. 
pees h 3 
oints reticulate-veined . A. LONGIFOLIUS. 
. ; 4 
Joints transversely ribbed . 5. A. RUGOSUS. 


1. A. monilifer, DC. Prod. ii. 353 (1825). 

Hledysarum moniliferum, L., Moon Cat. 54. Thw. Enum. 412. 
C. P. 3613. 

Fl. B. Ind. ii. 157 (not given for Ceylon). Burm. FI. Ind. t. 52, f. 3. 


Stems numerous, slender, prostrate, much branched, spar- 
ingly clothed with spreading deciduous bristly hair, 1. small, 
Z-2 in., on petiole about half as long, broadly oval, often cor- 
date at base, very obtuse, glabrous above, paler, with promi- 
nent venation and slightly bristly on the veins beneath, stip. 
long, membranous, very acute; fl. shortly stalked, about 
4-10 in rather close, erect, shortly pedunculate racemes ; cal. 
with scattered bristles, segm. very narrow ; pod. erect, monili- 


44 Leguminose. [Alyssicarpus. 


form, with 2-6 turgid, globose joints, pubescent, not reticulate, 
cal. a little longer than lowest joint. 

Low country; rare. Colombo (Ferguson); Trincomalie (Glenie) ; 
Maturata (Moon). 

Also in India, Burma, and Abyssinia. 

I have not met with this. 


2. A. vaginalis, DC. Prod. ii. 353 (1825). Aswenna, S. 

Burm. Thes. 104. Fl. Zeyl. n. 287. AHedysarum vaginale, L. Sp. 
Pl. ie Moon Cat. 54. Thw. Enum. 87. C.P. 1428. 

PE Bind i153) Burm hes. t240, 1. 


Stems numerous, long, 2-4 ft., ascending, branched, wiry, 
glabrous, often rooting at base; |. variable, 4-2 in., on petioles 
about a third as long, varying from linear-lanceolate to 
broadly oval, often cordate at base, acute, apiculate, glabrous 
above, whitish with adpressed hair and reticulately veined 
beneath, stip. scarious, as long as petiole, closely pressed to 
stem, very acuminate straw-coloured, persistent; fl. very 
shortly stalked, clustered, racemes short on short peduncles ; 
cal. slightly pilescens segm. setaceous, with a few long 
bristly hairs ; pod 4—# in. of 4-7 joints, erect, cylindrical not 
moniliform, apiculate, glabrous or nearly so, reticulate-venose, 
cal. nearly as long as two lowest joints. 

Low country; common, especially in the dry region. Fl. Dec., 
January; pinkish violet. 

Throughout the Tropics of the Old World. 

The habit is just that of Polygonum aviculare of Europe. A common 
form with short rounded leaves is A. xummularifolius, DC., to which 
has been erroneously referred Hedysarum nummularifolium, L. (for which 


see Indigofera echinata). Another form collected at Nilgala, in Uva: has 
very rigid linear leaves and a more shrubby habit. 


3. A. bupleurifolius, DC. Prod. ii. 352 (1825). Mutiraivali, 7. 
Thw. Enum. 87. C. P. 1427. 
PS BainG ai t5o. ikoxbs Corelatastone 


Stems long, 1-3 ft., stiff, wiry, nearly glabrous with long 
internodes ; |. linear but varying to oblong or even broadly 
oval below, very shortly stalked, acute, rigid, glabrous on both 
sides, stip. thin, chaffy, soon falling; fl. on very short ped., 
usually 2 together, distantly placed, racemes very long, erect, 
rigid, lax, spicate ; cal.-segm. lanceolate, acute, ciliate, much 
imbricated ; pod shortly stalked, 4-2 in., of 3-6 readily sepa- 
rating joints, very slightly moniliform, strongly apiculate, 
glabrous, without reticulations, cal. as long as lowest joint. 

Dry country; rare (?). Batticaloa; Bintenne; Mannar. Fl. Dec., 
Jan.; pink. 

Throughout the E. Tropics, also in China and Mauritius. 


Desmodium.]| MB egu MLNOSE. 45 


In the Mannar District the seeds are eaten as a grain, and have the 
same name as Panicum frumentaceum. 


4. A. longifolius, W. and A. Prod. 233 (1834). 

HhweEnums 412) Gy PY 3812. 

Fl. B. Ind. ii. 158 (not given for Ceylon). Wight, Ic. t. 251. 

Stems stout, ascending, 4-5 ft. glabrous; 1. on short 
petioles, 14-2 in., lanceolate-oblong, slightly cordate at base, 
obtuse, glabrous on both sides, stip. as long as petiole; fl. 
numerous, on short hairy erect ped., racemes rather dense, 
very long, spicate, bracts very conspicuous, concealing buds, 
ovate, acuminate, silky, falling before fl. open; cal. hairy, 
segm. oblong, acute, ciliate ; pod. 3-4 in., very shortly stalked, 
terete-compressed, strongly apiculate, of 3-6 joints, slightly 
moniliform, puberulous, reticulate-veined, cal. as long as 2 
lowest joints. 


Low country; very rare (?). Near Colombo (Ferguson). 
Throughout Peninsular India. 


5. A. rugosus, DC. Prod. ii. 353 (1825). 

A. scariosus, Grah., Thw. Enum. 88. C. P. 3589. 

Fl. B. Ind. ii. 150. 

Branches erect or prostrate, rather stout, more or less 
covered with spreading bristly hairs ; 1. small, on short hairy 
petioles, 4-1 in., varying from broadly oval to oblong or 
oblong-linear, somewhat cordate at base, obtuse, apiculate, 
glabrous above, slightly bristly beneath, stip. longer than petioles, 
acute, glabrous ; fl. nearly sessile, densely crowded in short 
nearly sessile heads or spikes, bracts large, chaffy, acuminate, 
strongly ciliate, persistent; cal.-segm. lanceolate, acute, 
strongly ciliate ; pod short, stalked, of 2—5 joints, moniliform, 
turgid, glabrous, strongly transversely ribbed, nearly all 
included in cal. 

Var. 8B, Heyneanus, Laker. A. Heyneanus, Wight, Thw. Enum. 88. 


Much larger, more hairy, especially |. beneath; fl. in longer 
spikes. 

Low country, in the dry region; rare. Mineri Tank; Mantai, &c., in 
the Mannar Dist. abundant. Var. 8, Colombo (Ferguson). Fl. Feb., 
March; bright dark pink, with the standard yellow, conspicuous. 

Also throughout the Tropics of the Old World, and in W. Indies and 
S. Africa. 

Our plant as usually met with is very dwarf and stunted, and is var. 2 
Slyractfolius of F1. B. Ind. 


20. DESMODIUM, Lesv. 


Small shrubs or perennial herbs, |. 3- or 1-foliolate, Ifits. 
shortly stalked, stipellate, fl. small, usually in lax axillary or 


46 Leguminose. [Desmodium. 


terminal racemes or panicles; cal. campanulate or cup-shaped, 
segm. usually nearly equal, upper often more or less connate ; 
keel-pet. obtuse, stam. diadelphous ; style curved upwards ; 
pod much compressed, composed of several 1-seeded joints, 
usually indehiscent (very rarely opening along ventral suture). 
—Sp. 120; 49 in FZ. B. Ind. 


Joints of pod well marked, as long as or longer than broad (except 
sometimes No. 6). 
Joints indehiscent. 
Stems more or less erect. 
FI. in axillary umbels or heads (Dendrolobium). 
Branches cylindrical, Iflts. obtuse . I. D. UMBELLATUM. 
Branches triquetrous, lflts. acute . 2. D. CEPHALOTES. 
Fl. in terminal or axillary racemes. 
Each cluster of fl. concealed between 
a pair of roundish Ifits. (Pyllodium) 3. D. PULCHELLUM. 
Fl. not concealed. 
Pod under 3 in., of 2 joints (Dicerma) 4. D. BIARTICULATUM. 
Pod over 3 in. 
Branches triquetrous, |. 1-foliolate 
(Pteroloma) ‘ : . 6. D. TRIQUETRUM. 
Branches not triquetrous. 
Joints at least twice as long as 
broad. 
Pod 3-4 in., joints 5 times as 
long as broad . : 7. D. ORMOCARPOIDES. 
Pod 2-3 in., joints 24 times as 
long as broad (Catenarza). 5. D. LABURNIFOLIUM. 
Pod 1-13 in., joints twice as 
long as broad. 
Pod long-stalked, joints 


obliquely truncate . . 8. D. GARDNERI. 
Pod sessile, joints semi- : 
circular . : . 12. D. THWAITESIHL. 
Joints less than twice as long as 
broad. 
Pod shortly stalked , - 9, D. SCALPE: 


Pod sessile. 
L. 1-foliolate. 
Lflt. broad-oval, acute . 10. D. GANGETICUM. 
Lfit. ovate-deltoid, obtuse 11. D. LATIFOLIUM. 
L. 3-foliolate. 
Bracts small, setaceous . 13. D. WIGHTII. 
Bracts large, ovate or 
lanceolate. 
Pod of 5-8 joints in- 
dented on ventral 


margin. 
Pods spreading . 14. D. RUFESCENS. 
Pods erect : . 15. D. HETEROCARPUM. 


Pod of 3 or 4 joints, 
not indented- . . 16. D. JUCUNDUM. 


Desmodium. | ib egum 2720S. 47 


Stems numerous, prostrate, slender. 
Fl. all axillary, without a peduncle . 17. D. TRIFLORUM. 
Some fl. ona peduncle . : . 18. D. HETEROPHYLLUM. 
Joints dehiscent; stems prostrate, slender . 19. D. PARVIFLORUM. 
Joints of pod inconspicuous, shorter than broad, 


dehiscent. 
Pod very slightly hairy . : : a . 20. D. GYRANS. 
Pod with long shaggy hair . : : . 21. D. GYROIDES. 


I. D. umbellatum, DC. Prod. ii. 325 (1825). 

Herm. Mus. 21. Burm. Thes. 115. Fl. Zeyl. n. 293. Hedysarum 
umbellatum, Sp. Pl. 747; Moon Cat. 54.  Dendrolobium umbellatum, 
Benth., Thw. Enum. 86. C. P. 1436. 

Fl. B. Ind. ii. 161. Burm. Thes. t. 51. 


A shrub, 5-6 ft., much branched, young parts with dense, 
adpressed pubescence ; 1. 3-foliolate, rachis 1-14 in. downy, 
stip. small, acicular, quickly falling, Iflts. 14-3 in. (the terminal 
much the largest) on short silky stalks, oval, obtuse at both 
ends, glabrous above, pubescent and whitish beneath, lat. veins 
prominent, parallel, curved, running to margin; fl. on silky 
ped. as long as or longer than cal., about 4-10 m stalked axil- 
lary umbels, peduncles 4-1 in.; cal. densely silky; pod 
1-14 in., slightly curved, of 3-6 broadly oblong joints, slightly 
indented between each, reticulate-veined, adpressed-hairy. 


Low country; common. : 
Also in Burma, Malaya, Polynesia, and the Mascarene Is. 


2. D. Cephalotes, Wall. Cat. n. 5721 (1828). 
Dendrolobium Cephatotes, Benth. Thw. Enum. 86. C. P. 2780. 
Fl. B. Ind. ii. 161. Wight, Ic. tt. 373 and 209 (D. congestum). 


A shrub, 4-5 ft., branches ascending, acutely triangular, 
densely adpressed-silky; 1. 3-foliolate,rachis #-1+in., triquetrous, 
channelled above, stip. lanceolate, bristle-pointed, glabrous, 
persistent, lflts. shortly stalked, with filiform conspicuous 
stipels, 2-34 in. (terminal one rather the larger), very acute, 
glabrous above, silky and white beneath when young, veins 
conspicuous beneath, parallel, reaching to margin; fl. very 
numerous, very shortly stalked, densely crowded in roundish 
axillary heads, peduncle very short 4-} in., cal. densely 
white-silky ; pod #-1 in., slightly curved, of 3-6 joints which 
are as broad as long, indented between the joints, slightly 
pubescent. 

Low country up to 3000 ft.; very common. FI. Jan.; bright red. 

Also in India, Malaya, and China. 


48 Leguminosae. (Desmodium. 


3. D. pulchellum, Benth. in Fl. B. Ind. ii. 162 (1876). Ham- 
pilla, S. 

Herm. Mus. 36, 58. Burm. Thes. 116. FI. Zeyl. n. 292. Aedysarum 
pulchellum, L. Sp. Pl. 747; Moon Cat. 54. Phyllodium pulchellum, Desv., 
Thw. Enum. 86. C. P. 1270. 

Fl. B. Ind. ii. 162. Burm. Thes. t. 52. Wight, Ic. t. 418. 


A small shrub about 4 ft., branches numerous, pubescent ; 
1. 3-foliolate, rachis short, about I in., pubescent, stip. small, 
acuminate, lfits. shortly stalked, the terminal nearly twice as 
large as the lat. ones, oblong-lanceolate, rounded at base, sub- 
acute or obtuse at apex, repand or undulate at margin, nearly 
glabrous above, densely silky-pubescent and pale beneath ; 
fl. small on short ped., arranged in clusters of 3 or 4 in long 
erect terminal and axillary inflor., each cluster entirely con- 
cealed by a pair of rotundate Iflts. ( bracts’) placed back to 
back on a very short common petiole and persistent ; cal. 
pubescent; pod small, 2in., joints 2, a little longer than broad, 
glabrous, reticulate-veined. 


Low country up to 3000 ft.; common, especially in the dry region. 
Fl. March. 

Also in India, Malaya, and China. 

The Iflts. enclosing the little umbels of flowers and fruit are obviously 
the lateral ones of the ordinary foliage leaves modified; the large terminal 
Iflt. is suppressed, but when young the rachis is present as a long-pointed 
bristle. 

Hermann gives ‘ Palzghas’ as the S. name for this. 


4. D. biarticulatum, Genth. in Fl. B. Ind. ii. 163 (1876). 

Herm. Mus. 5, 31. Burm. Thes. 114. FI. Zeyl. n. 296. Hedysarum 
biarticulatum, L. Sp. Pl. 747. Moon Cat. 54. Dicerma biarticulatum 
DC., Thw. Enum. 86. C. P. 1450. 

Fl. B. Ind. ii. 163. Burm. Thes. t. 50, f. 2. Wight, Ic. t. 419. 

A small spreading shrub, with slender elongated branches 
covered with adpressed silky hair ; 1. small, 3-foliolate, rachis 
4-{ in., hairy, stip. connate at back, laciniate, striate, glabrous, 
brown, Iflts. on very short hairy stalks, oblong-lanceolate, 
obtuse, minutely apiculate, conduplicate, glabrous above, more 
or less silky beneath ; fl. on very short ped., in small clusters, 
laxly arranged in a long, narrow, spicate, terminal raceme, 
bracts small, setaceous ; cal.-segm. longer than tube, slightly 
silky ; pod small, 2 in., of 2 rounded joints, very deeply in- 
dented between them, strongly pubescent. 


Low country; common, especially in the dry region. Fl. Jan.—March. 
Also in India, Burma, Malay Islands, and N. Australia. 


5. D. laburnifolium, DC. Prod. ii. 337 (1825). 

D. cateniferum, Arn. Pug. 13. Catenaria laburndtfolia, Benth., Thw. 
Enum. 86. C. P. 1435. 

Fl. B. Ind. ii. 163. 


Desmodium.) Leguminose. 
as 49 


A slender shrub with nearly glabrous branches, |. 3-folio- 
late, rachis 14~13 in., channelled, nearly glabrous, stip. small, 
acuminate, lfits. 3-4 in. (the terminal rather the largest), 
lanceolate, acute, glabrous above, very slightly pubescent 
beneath, stiff ; fl. medium-sized, on slender spreading ped.a 
little longer than cal., racemes slender, axillary, about as long 
as l., bracts setaceous ; cal. slightly pilose, the lowest segm. 
much the longest ; pod 2-2? in., linear, indented between the 
joints, joints 3-6, more than twice as long as broad, rough with 
minute hooked hairs. 

Upper zone of moist low country, rather common (?). Hantane; 


Dolosbagie. Fl. Dec. h 
Also in E. Bengal, Malay Is., China, and Japan. 


6. D. triquetrum, DC. Prod. ii. 326 (1825). Baloliya, S. 

Herm. Mus. 35. Burm. Thes. 176. Fl. Zeyl.n. 286. Hedysarum 
triguetrum, L. Sp. Pl. 746. Moon Cat. 54. Péteroloma triquetrum, 
Benth., Thw. Enum. 86. C. P. 1431. 

Fl. B. Ind. ii. 163. Burm. Thes. t. 81 (leaves only). 

An erect shrub, branches few, erect, triquetrous, glabrous 
or hairy on the angles only; 1. 1-foliolate, petiole 3-1} in., 
leaflike, with a broad wing on either side, stip. 4-2 in., erect, 
lanceolate-linear, very acute, glabrous, brown, Ifit. variable in 
size, 3-6 in. (the uppermost ones often smaller), linear- 
lanceolate (the uppermost almost linear), subcordate at base, 
gradually tapering to acute apex, glabrous above, hairy on 
midrib beneath ; fl. small, on short slender ped., 1-3 in a 
cluster, arranged rather laxly in slender, erect, spike-like 
terminal and axillary racemes 6-12 in. long, bracts setaceous; 
cal. with few bristly hairs; pod about 1 in., erect, linear- 
oblong with 4-7 joints, continuous and scarcely indented 
joints as broad as or broader than long, hairy. 

Moist low country up to 3000 ft.; very common. FI. Sept.; bright 
violet, often white. 

Also in India, Burma, China, Philippine and Seychelle Is. 


Burmann’s figure was made from a living plant in the Amsterdam 
Hortus Medicus, grown from seed sent from Ceylon by Hartog. 


7. D. ormocarpoides, DC. Prod. ii. 327 (1825). 

Thw. Enum. 87, 441. C. P. 3812. 

Fl. B. Ind. ii. 164. 

A small slender under-shrub, the stem procumbent and 
rooting at the base, branches hairy; 1. 1-foliolate, striate, 
glabrous, 3-4 in., ovate, somewhat acuminate, mucronate, 
rounded at base, somewhat crenate at margin, glabrous above, 
paler and silky-pubescent especially on veins beneath, petiole 
4—4 in., stip. acuminate, stipels filiform, conspicuous ; fl. few, 

PART II. E 


50 Leg UNULNOSE. [_ Desmodium. 


on filiform spreading ped., much longer than cal., in very lax 
terminal racemes, bracts small; cal. slightly hairy; pod 3-4 in. 
long, spreading or pendulous, linear, somewhat curved, dis- 
tinctly constricted between the joints, joints 4—6, five times as. 
long as broad, longitudinally striate, densely covered with 
minute hooked hairs. 

Montane zone; very rare. Matale East (Thwaites). F1. June; pale 


violet. 
Also in the hills of S. India. 


68. D. Gardneri, Benth. Pl. Jungh. 226 (1853). 
’ D. podocarpum, Thw. Enum. 87 (non DC.).  C. P. 1434. 
FI. B. Ind. ii. 165. 


A slender, slightly branched, erect perennial, 14-3 ft., 
woody at the base, stem angled, slightly hairy, purplish; 1. 
large, mostly from the base of stem, 3-foliolate, rachis long, 
2-6 in., stip. acute, striate, glabrous, lflts. 2-5 in. (the terminal 
the largest), ovate, acuminate, obtuse, glabrous on both sides, 
the lat. nes meually unequal-sided ; fl. medium-sized, on 
slender ped. 3-2 in. long, in small clusters of BVOr IA distantly 
placed ona nee erect terminal raceme or panicle much ex- 
ceeding the 1.; cal. cup-shaped, segm. very short and broad ; 
pod on a stalk 4-2 in. long, joints 1-3, about 4 in. long, each 
one bent down at right angles with the last or with the stalk, 
wider at the end which is rounded and obliquely truncate, 
tapering to very narrow base, finely hairy. 

Moist region; 1000-4000 ft.in shady places; rather common. Matale; 
Hantane; Ramboda; Hunasgiria; Rangala; Nitre Cave. FI. April, 
Sept., Oct.; pinkish white. 


Also i in ics India and in Japan. 
A pretty and curious plant; very distinct from our other species. 


g. D. Scalpe, DC. Prod. ii. 334 (1825). 
D. strangulatum, W. and A., Thw. Enum. 87. C. P. 1433. 
FI. B. Ind. ii. 165. Wight, Ic. t. 985 (D. strangulatum). 


A slender, erect perennial, woody at base, stem slightly 
angled, pubescent, |. 3-foliolate, spreading, rachis 14-24 in., 
hairy, stip. large 4-% in., persistent, brown, Iflts. 14-24 in., 
owate, rounded at base, acute, slightly hairy above, densely 
and finely pubescent beneath, the terminal one rather larger 
and somewhat rhomboidal; fl. rather large on slender as- 
cending ped. 1-14 in. long, arranged in pairs on branches of 
very lax slender pubescent terminal panicles much over- 
topping the 1, bracts small, ovate, caducous; cal. hairy, 
deeply cut, segm. acuminate; pod very shortly stalked, 
nearly straight, straight on dorsal, deeply indented on ventral 


Desmodium.) Leg UMLNOSCE. 51 


margin, joints 3 or 4, boat-shaped, 14 as long as broad, finely 
pubescent. 

Montane zone; rather common. F/I. Oct._Jan.; salmon-red or pale 
orange. 

Also in S. India, Malay Is., Mascarene Is., and Trop. Africa. 

A graceful plant, worth cultivation for the peculiar colour of its flowers. 


Io. D. gangeticum, DC. Prod. ii. 327 (1825). 

Burm. Thes. 113. Medysarum gangeticum, L. Sp. Pl. 746. Thw. 
Enum. 411. C. P. 3813. 

Fl. B. Ind. ii. 168. Burm. Thes. t. 49, f. 2. Wight, Ic. tt. 271 and 
(D. latifolium) 272 (not 270). 

A slender under-shrub, stems cylindrical, nearly glabrous, 
young parts hairy; 1. 1-foliolate, petiole long, #-14 in., hairy, 
stip. setaceous, lfit. broadly oval or rotundate, rounded at base, 
subacute, glabrous above, silky and whitish beneath; fl. very 
small, on short erect hairy ped. in lax erect terminal spicate 
racemes, bracts setaceous; cal. hairy, segm. lanceolate; pod 
small, 4-1 in., usually somewhat curved, nearly straight on 
dorsal, very deeply indented on ventral margin, joints 4-8, as 
broad as long, rounded on ventral margin, set with minute 
hooked hairs. 

Montane zone, apparently very rare, but probably overlooked. Matale 


East (Thwaites). Fl. May. 
Throughout Trop. Asia and Africa, &c. 


11. D. latifolium, DC. Prod. ii. 327 (1825). 
Thw. Enum. 87. C. P. 3588. 
FI. B. Ind. ii. 168. Wight, Ic. t. 270 (not 272). Bot. Reg. t. 355. 


A small shrub, 3-5 ft., branches densely velvety-pubescent, 
1. 1-foliolate, petiole short, about } in., stip. broad, suddenly 
acuminate, lfit. 24-3 in., deltoid-ovate, truncate or subcordate 
at base, obtuse or subacute, repand at margin, finely hairy 
above, densely velvety beneath; fl. small, numerous, on ped. 
shorter than cal., rather crowded on the branches of copious 
axillary and terminal ascending panicles; cal. hairy, segm. 
linear-lanceolate ; pod about # in., straight, slightly indented on 
dorsal, much more so on ventral margin, joints 4-6, as long as 
broad, hairy. 

Dry region; rather rare. Bibile; Mineri. There are also specimens 
labelled Matale East, probably from the low country. Fl. Feb., June, 
August; bright red-purple. 

Throughout the Tropics of the Old World. 


12, D. Thwaitesii, Laker in Fi. B. Ind. ii. 169 (1876). 
D. strangulatum, var. minor, Thw. Enum. 87. C. P. 3327. 
Fl. B. Ind. ti. 169. 


52 Leguminose. [Desmodium. 


A very small perennial, slightly woody at base; stem 
prostrate and rooting at base, scarcely branched, very hairy; 
1, 3-foliolate, rachis #-14 in. hairy, stip. narrow, very 
acuminate, with long white hairs, lflts. small ?-14 in., oval, 
obtuse or subacute, nearly glabrous above, pale with silky 
hair beneath, the terminal one the largest; fl. few, very small, 
on slender ascending ped. 4 in. long, in pairs in slender, erect, 
very lax, terminal racemes, bracts caducous; cal. nearly 
glabrous, segm. lanceolate, acute; pod ?-1 in., straight, scarcely 
indented along dorsal, much so along ventral edge, joints 2-4, 
twice as long as broad, semicircular, covered with fine hooked 
hairs. 

Lower montane zone; rare. Ambagamuwa. FI. Oct., Nov. 

Endemic. 


This appears to be closely allied to D. Scalfe, of which Thwaites, 
indeed, considered it a variety. 


13. D. Wightii, Grak. in Wall. Cat. n. 5718 (1828). 

D. Walkeri, Arn. Pug. 13. Thw. Enum. 87. C. P. 31209. 

Bly B. Ind. it, 169: 

A tall perennial herb, 2-4 ft., branches slender, striate, 
slightly hairy; 1. 3-foliolate, rachis 14-2} in., stip. setaceous, 
soon falling, lfits. 2-34 in., lanceolate-oval, obtuse or subacute, 
mucronate, somewhat undulate at margin, glabrous or slightly 
hairy above, finely silky beneath; fl. small, on very slender 
ped. 2-4 times as long as cal., in pairs in very lax narrow, 
elongated, spicate, terminal and axillary racemes, bracts 
setaceous as long as ped., conspicuous; cal. very hairy, segm. 
lanceolate, acute; pod rather under 1 in., slender slightly 
falcately curved, scarcely indented on dorsal, deeply so on 
ventral edge, joints 6 or 7, as broad as long, transversely 
wrinkled, slightly rough with minute bristles. 

Low country; very common. Fl. Jan.—April. 

Also in S. India. 


D. Watkeri, Arn. (founded on specimens of Walker’s n. 205) seems to 
be merely a sub-glabrous state; but C. P. 2973, referred to this by Thw. 
(1. c. 87), is D. cajanefolium DC.—an introduced weed in the Bot. 
Gardens, and a Tropical American species. To this also, or to the 
closely allied D. /ecocarfpum, Don, of Brazil, is to be referred C. P. 3544 
(from Maturata) given by Thwaites (l.c. 411) as Hedysarum mucronatum, 
Bl., no doubt also an introduction. 


D. diffusum, DC. Another weed in the Peradeniya Gardens and 
round about appears to be this common Indian species. 


14. D. rufescens, DC. Prod. ii. 335 (1825). 
D. ferrugineum, Wall., Thw. Enum. 87. C. P. 1426. 
BIS Belinda 1 Wwacht let mcutOoAs 


Desmodium. Leruminose. a2 
: 58) 


An erect much-branched under-shrub, branches terete or 
nearly so, densely clothed with short spreading yellow or 
orange hair; |. 3-foliolate, rachis 1-14 in., pilose, stip. quickly 
caducous, lflts. 1-2 in., obovate, rounded, truncate or retuse at 
apex, strongly mucronate, glabrous above, densely silky and 
silvery beneath; fl. large on erect hairy ped. longer than cal., 
in rather dense terminal and axillary racemes, bracts large, 
ovate, acuminate, very silky, soon falling; cal. puberulous, 
segm. lanceolate, acute, ciliate; pod 1-14in., linear, falcately 
curved, not at all indented on dorsal and only slightly so on 
ventral margin, joints 5-8, longer than broad, slightly 
pubescent. 

Moist region up to 4ooo ft.; common. Fl. Jan.-April; bright purple. 

Also in S. India. 

The young spikes are completely covered by the large overlapping 
bracts. Varies very much in amount of hairiness, sometimes almost 
glabrous. Occasionally cultivated for its pretty flowers. 


[D. capitatum, DC. Given for Ceylon in Fl. B. Ind. ii. 171, on 
authority of the younger Burman. That author, however (FI. Ind. 167), 
does not give Ceylon as a locality, but merely quotes as a synonym a 
name in Burm. Thes. 115. This from the citation of Hermann given is 
only a variety of D. heterocarpum. But the plant is likely enough to 
occur here. ] 


15. D. heterocarpum, DC. Prod. ii. 337 (1825). Et-undu- 
DPiyali, S. 

Herm. Mus. 22. Burm. Thes. 117. FI. Zeyl. n. 294. Hedysarum 
heterocarpum, L. Sp. Pl. 747; Moon Cat. 54. D. polycarpum, DC., Thw. 
Enum. 86. C. P. 1425. 

Fl. B. Ind. ii. 171. Burm. Thes. t. 53 f. 1. Wight, Ic. t. 406 (D. foly- 
carpumy). , 

A suberect perennial, somewhat woody and often rooting 
below, branches very numerous, elongated, cylindrical, finely 
downy with adpressed hair; 1. 3-foliolate, rachis 1-1} in., stip. 
large, narrow, filiform-acuminate, persistent, Iflts. variable, 
1—24 in., shortly stalked, lanceolate or oval, obtuse or subacute, 
glabrous above, more or less silky with adpressed hair beneath, 
stipels filiform, conspicuous; fl. rather small, on slender erect 
ped. longer than cal., crowded in short, dense, erect, terminal 
and axillary racemes, bracts large, lanceolate, very acuminate, 
ciliate; cal. nearly glabrous; pods erect, crowded, about 1 in., 
straight, not indented on dorsal, moderately so on ventral 
margin, joints 5-7, about as broad as long, with raised 
reticulation, glabrous or hairy. 

Var. B, trichocaulon, Laker in Fl. B. Ind.l.c. C. P. 3512. 


Stem and branches with white spreading hair; racemes 
more lax, bracts less acuminate; pods hairy. 


54 Leguminose. [Desmodium. 


Moist low country and lower montane zone; very common; var. 8 in 
Maturata, Kalupahane, &c. Fl. May-Sept., Nov.; purple, sometimes 
white. 

Throughout the Eastern Tropics, China, and Japan. 

As DC.’s two names are of the same date, | have preferred to maintain 
that which preserves Linnzeus’s specificname. This specially belongs to 
the Ceylon plant, though the peculiarity to which it refers, z.e., the lower 
pods with only one joint, is by no means always or even often present. 


16. D. jucundum, 7iw. Enum. 411 (1864). [PLATE XXVIII] 
C. P. 3778. 
TPH 183, revel, Tul, 9/724, 


A shrub, 3 or 4 ft. high, branches stout, purplish, the 
young shoots white with dense silky hair; 1. rather crowded, 
3-foliolate, rachis 1-1} in. velvety, stip. ovate, acuminate 
slightly silky, crimson, lfits. 1-2 in. (the terminal rather the 
larger), oblong or obovate-oval, very obtuse, minutely mucro- 
nate, glabrous above, white with a dense covering of silky 
hair beneath and with very prominent lat. veins; fl. large on 
slender erect ped., longer than cal., usually in pairs; arranged 
in short, erect, pubescent, terminal racemes, bracts large, ovate, 
acute, silky, red, soon falling; cal. nearly glabrous, segm. longer 
than tube, narrowly lanceolate, acuminate, ciliate; pod #—1 in., 
straight, shortly beaked, not at all indented on dorsal and 
scarcely on ventral margin, joints 3 or 4, half as long again 
as broad, faintly reticulate, slightly pilose. 

Lower montane zone; very rare. Half way up Lagalla, Matale East; 
first discovered by Mr. A. O. Brodie in Oct. 1862. 

Fl. June, Sept., Oct.; rich mauve-purple. 

Endemic. 

A showy species, well suited for garden cultivation. 


17. D. triflorum, DC. Prod. 334 (1825). Hin-undupiyali, S. 

Herm. Mus. 2, 38. Burm. Thes. 118. Fl. Zeyl. n. 297. Hedysarum 
triftorum, L. Sp. Pl. 747 (pro max. parte); Moon Cat. 54. Thw. Enum. 
0; (Ca 1, 2/710) 

BBs Inds 1737) Burm. dihes: t 54) fo) Wiehe lento 


A very small perennial herb, with numerous long slender 
prostrate branches rooting at the nodes, clothed with white 
spreading hairs; 1. small, 3-foliolate (lower ones often 1-folio- 
late), rachis under 4$in. stip. ovate, acuminate, persistent, 
lfits. 4-2 in., broadly oval or obovate or obcordate, glabrous 
above, slightly silky beneath; fl. very small, on long very 
slender hairy ped., 1-5 (usually 3) together in the axils of L; 
cal. with few bristly hairs, segm. long, setaceous; pod 4-4 in., 
nearly straight, not indented on dorsal, slightly so on ventral 
margin, joints 2-4, as long as broad, reticulate, glabrous. 


Desmodium. Leg umuwnose. 55 


Low country up to 2000 ft. or more; very common. FI. June—Oct.; 
bright purple. 

Throughout the Tropics. 

An abundant and valuable constituent of turf in Ceylon, standing 
drought well. This and D. heterophyllum are used as a medicine in cases 
of fever. 


18. D. heterophyllum, DC. Prod. ii. 334 (1825). Maha- 
undupiyali, S. 

Burm. Thes. 118. Fl. Zeyl. n. 297 6. Hedysarum heterophyllum, 
Willd., Moon Cat. 54. Thw. Enum. 86. C. P. 2778. 

Peis ind. i. 175.) burm. Lhes. t.54, f 1. Wicht; Ie: t. 201. 


A perennial herb, stems numerous, prostrate, slender, 
rooting at lower nodes, densely covered with spreading hairs ; 
1. 3-foliolate, rachis $—? in., hairy, stip. broadly lanceolate, 
acute, ciliate, persistent, lfits. $-1 in. (term. one the largest), 
oval or obovate-oval, rounded at both ends, glabrous above, 
sparingly hairy beneath; fl.small, on very slender ped., 4-1 in. 
long, 2-4 together, axillary racemose or umbellate, bracts 
inconspicuous; cal. very hairy, segm. narrow, setaceous; 
pod. ?-1 in. straight, not at all indented on dorsal, but 
considerably so on ventral margin, joints 3-6, as broad as 
long, faintly reticulate, slightly hairy. 

Low country up to 2000ft.; very common. FI. pale purple. 


Also throughout India and Malaya, and in China, the Philippines and 
the Mascarene Is. 


19. D. parvifolium, DC. Prod. ii. 334 (1825). 

Thw. Enum. 68. C. P. 1429. 

F]. B. Ind. i. 174. 

A low under-shrub, with very numerous slender branches 
trailing to a great length, with many short lateral twigs, when 
young triangular, hairy; Il. very numerous, crowded, small, 
3- (or often 1-) foliolate, rachis short, filiform, stip. linear- 
lanceolate, acuminate, persistent, Iflts. }-}? in. (the terminal 
much the largest), lanceolate or oval, obtuse or subacute, 
mucronate, glabrous above, slightly hairy beneath; fl. very 
small, on long, filiform, hairy ped., arranged in very lax 
spreading racemes, which are axillary or terminal on the 
lateral twigs, much exceeding l., bracts inconspicuous ; cal. 
covered with long silky hairs, segm. long, setaceous ; pod 
under 4} in., straight, indented on both margins, dehiscent 
along ventral suture and everted, joints 1-4, roundish, about 
as long as broad, rough with minute hairs. 

Montane zone to over 7000 ft.; common in patana land. I have 
wag seen it in the low country. Fl. Feb., April, September ; bright 
JINK. 

; Also in India, Malaya, China, and Japan. 


56 UL eg UmunOse. [ Desmodium... 


Each joint of the pod is completely: dehiscent along the ventral 
suture, but the valves remain united along the dorsal margin and are- 
stiffly everted. 


20. D. gyrans, DC. Prod. ii. 326 (1825). Chanchala, 5S. 
Fledysarum gyrans, L.f., Moon Cat. 54. Thw. Enum. $7. C. P. 1430. 
Fl. B. Ind. u. 174. Wight, Ic. t. 294. 


An erect perennial, branches few, cylindrical, glabrous, 
young shoots roughly pubescent; 1. 3- (often 1-) foliolate, rachis. 
#1 in., glabrous, stip. linear-lanceolate, acuminate soon falling; 
Iflts. on very short densely pubescent stalks, very unequal, ter- 
minal one 2-3 in., drooping, linear-lanceolate, obtuse, glabrous. 
above, silky-pubescent beneath, lat. ones about 4 in., broadly 
linear, tapering to base; fl. of medium size, on short ped. 
arranged in rather long, erect, lax, terminal and axillary 
racemes, bracts broadly ovate, acuminate, glabrous, soon 
falling ; cal. cup-shaped, nearly glabrous, segm. deltoid, 
short, subacute ; pod 14-13 in., linear, slightly falcate, dehis- 
cent along ventral suture which is slightly indented, slightly 
rough with short scattered hair, joints 5-10, broader than 
long, inconspicuous. 

Moist and intermediate low country to 3000 ft.; rather rare. Han- 
tane ; Galagama ; about Badulla, frequent. Moon’s locality is Dumbara 
Fl. Feb.-April.; pale pink. 

Throughout India, Malaya, and the Philippines. 


This is the ‘Telegraph,’ or rather ‘Semaphore Plant,’ so named from. 
the spontaneous jerky movements of the pair of small leaflets. 


21. *D. gyroides, DC. Prod. ii. 326 (1825). 
Thw. Enum. 87. C. P. 3326. 
IM) 18), levels re W9/Is. 


A shrub, 4-6 ft. high, twigs pubescent; |. 3- or more. 
usually 1-foliolate, rachis #—1 in., slender, often deflexed, stip. 
linear-lanceolate, long-acuminate, terminal Iflt. 14-2 in., ob- 
long-oval, rounded at both ends, glabrous above, slightly 
pilose beneath, lat. ones 4-3? in.; fl. of moderate size, om 
spreading hairy ped., crowded in short, pedunculate, terminal 
or axillary racemes, bracts broadly ovate, apiculate, ciliate, 
caducous; cal. campanulate, nearly glabrous, segm. short, 
triangular, subacute ; pod 1-1} in., nearly straight, dehiscent. 
along ventral suture which is very slightly indented, densely 
shaggy with short fulvous hair, joints 5-9, broader than: 
long. 

Moist low country, probably introduced. Hantane (Hb. Perad.). L 


have only seen it as a cultivated plant here. FI. April; purple. 
Also in E. Bengal, Burma, and the Malay Islands. 


Abrus.] Leguuinose. 


On 
N 


21. ABRUS, J. 


Twining perennials, 1. abruptly pinnate with deciduous 
stipellate lfits., fl. on short swollen branchlets of axillary 
racemose panicles; cal. small, campanulate, truncate ; keel- 
pet. completely connate, slightly curved longer than wings 
and standard; stam. 9 (tenth one absent), connate into a 
tube split above, adnate at base to standard; style glabrous, 
stigma capitate ; pod flat, oblong or linear, 3-9-seeded with 
papery septa between the seeds.—Sp. 5; 3 in #2. B. Ind. 


Infl. shorter than 1.; pod oblong . : : eA PRECATORIUS: 
Infl. much longer than 1.; podlinear . ‘ 5 By deo PUY GIST EOSS 


1. A. precatorius, Z. Syst. Nat. ed. 12, 472 (1767). Olinda- 
wel, S|. Kuntumani, 7. 

Herm. Mus. 16. Burm. Thes.177. Fl. Zeyl. n. 284. Moon Cat. 52. 
Thw. Enum. 91. C. P. 1466. 

Pies. ine 1.175. Bentl. and Trim. Med. Pl. i. t. 77. 

A perennial twiner, stems numerous, scarcely woody, 
slender, glabrous, with long internodes; |. spreading, pinnate, 
rachis 2-4 in., thickened at base, very slender, channelled, 
nearly glabrous, stip. minute, setaceous, Ilflts. 20-24 (10-12 
pair) or more, opposite, very shortly stalked, quickly de- 
ciduous, about 2 in., oblong, obtuse at both ends, minutely 
apiculate, glabrous above, slightly hairy beneath, thin; fl. 
rather small, shortly stalked, several together on very short 
swollen knob-like branches crowded at end of stout, curved, 
swollen peduncles, usually shorter than 1|.; cal. glabrous or 
very slightly silky ; pod. 14-1? in. by 4 in. wide, flat, oblong, 
truncate, with a sharp deflexed beak, finely silky ; seeds 3-5, 
rather over } in., ovoid, usually bright scarlet with a black 
patch on top, polished. 

: ail country ; common, especially in the dry region. Fl. Feb.; pale 
violet. 

Every Tropical country. 

The pretty scarlet and black seeds are well known, and are used as 
weights by jewellers. The flowers are sometimes white when the seeds 
are of the same colour; there is also a form with the seeds wholly black. 

The root is the well-known ‘Indian Liquorice,’ Glycyrrhiza indica of 
the older writers, and is used both internally and externally in native 
medicine. The seeds are an acrid poison unless cooked. 


2. A. pulchellus, Wa/l/. Ca/. n. 5819. 

Thw. Enum. 91. C.P. 1467. 

Fl. B. Ind. ii. 175. 

Much like the last, but stems slightly hairy ; Iflts. 10-20 
(5-10 pair), 1-1} in., narrowly oblong; fl. less crowded in 


58 Leguminosae. [Dumasia. 


more lax racemes with more elongated branches, and pe- 
duncles much exceeding 1.; pods 2-24 in. by 2 in. broad, 
linear, slightly curved upwards, shortly beaked, thinly hairy ; 
seeds 9 or 10, oblong-ovoid, somewhat compressed, black. 


Dry region; rather common. Fl. December—Feb.; pale violet. 
Also in E. Bengal, Burma, Malay Islands, Trop. and S. Africa. 


22, SHUTERIA,* W. and A. 


Twiner, |. pinnately 3-foliolate, stipels conspicuous, f1. 
in short racemes ; cal. tubular-campanulate, with short teeth ; 
standard exceeding other pet.; stam. diadelphous; style 
beardless, stigma capitate; pod linear, flat, slightly curved 
upwards, with several seeds.——Sp. 5; 4 in Fl. B. Lud. 


S. vestita, W. and A. Prod. 207 (1834). 
Thw. Enum. 88. C. P. 661. 

PIB. Inds ii 181) Wight, Ter t-1165: 

Stems slender, twining, and branched, very hairy ; |.-rachis 
14-24 in., hairy, stip. lanceolate, acute, ciliate, lflts. shortly 
stalked, broadly oval or rhomboid-oval, 1-12 in., rounded at 
both ends, apiculate, hairy on both sides, stipels long filiform ; 
fl. small, on hairy ped., in erect axillary racemes shorter than 
l., bracts lanceolate, hairy, persistent; cal. densely covered 
with long hairs, teeth acuminate ; pod 1-1} in., linear, nearly 
straight, compressed, thin, softly hairy ; seeds 5-7, ovoid, com- 
pressed, brown. 

Montane zone, 4000-7000 ft.; rather rare. N. Eliya; Maturata; 


Ramboda; Passara. Fl. August, January. 
Also in the Indian hills. 


23. DUIMASTIA, DC. 


Twining herb, 1. 3-foliolate, fl. in axillary racemes; cal. 
tubular, gibbous at base with an obliquely truncate mouth; 
pet. equal, wings and keel adherent, with very long claws; 
stam. diadelphous; ov. stalked; pod linear, torulose, 2- or 
3-seeded.—Sp. 3; 2 in FZ B. Ind. 


D. villosa, DC. M/ém. Leg. 257 (1825), var. leiocarpa, Benth (sp.). 
D. pubescens, DC., Arn. Pug. 12. Thw. Enum. 88. C. P. 663. 
Fl. B. Ind. ii. 183. Wight, Ic. t. 445 (D. congesta). 


* Named after Dr. Shuter, ‘ Naturalist on the Madras Establishment,’ 
to whom the genus Shutereza (Convolvulacez) had, however, been dedi- 
cated the previous year (1833) by Choisy. 


Glycine.) Leguminose. 59 


A twining herb, stems slender with long internodes, 
glabrous or with few adpressed hairs; |.-rachis 2—4 in., swollen 
at base, stip. small, linear, lflts. shortly stalked, 14-2 in., ovate, 
very obtuse, apiculate, glabrous on both sides or with very 
few hairs beneath; fl. moderate-sized, on slender ped., 
arranged in racemes which are either of 3 or 4 fl. and much 
shorter than 1., or of 12-20 fl., and equalling 1; cal. quite 
glabrous, persistent; pod about Ijin., stalked and exserted 
beyond persistent cal., linear, straight, sharply beaked, toru- 
lose, perfectly glabrous; seeds 2 or 3, ovoid, fin., black and 
shining. 

Upper montane zone; common. FI. Feb., March; pale yellow. 

Also in the Indian mountains, and in Java, Madagascar, and Natal. 


The ordinary form of this species has the stem densely hairy, and the 
leaves very pubescent beneath; but this has not been found in Ceylon. 


24. GLYCINE, Z. 


Herbaceous twiner, |. 3-foliolate, with stipels, fl. small, 
numerous, in narrow racemes; cal. with a campanulate tube, 
segm. long, filiform; pet. about equal,not longer than cal.-segm.; 
stam. monadelphous, anth. uniform, all fertile; style beardless, 
stigma capitate; pod linear, straight, 2-4 seeded, with septa 
between seeds.—Sp. 12; 3 in FZ. B. Ind. 


G. javanica, LZ. Sp. P/. 754 (1753). Goradiya, S. 

Moon Cat. 53. Benth. in Journ. Linn. Soc. viii. 266. Soya Wighizz, 
Grah., Thw. Enum. 88. G. P. 1477. 

Fl. B. Ind. ii. 183. 


An herbaceous twiner, stems somewhat angular, clothed 
with deflexed hair, young shoots very silky; 1.-rachis 2—3 in., 
with deflexed hair, stip. small, triangular, acute, lflts. shortly 
stalked, 11-2} in., broadly oval (the lat. ones unequal-sided), 
subacute, strongly apiculate, softly hairy on both sides, paler 
beneath with the lat. veins strongly marked; fl. small, 
numerous, on short, slender, hairy ped., crowded in narrow, 
rather dense, shortly stalked, axillary racemes 1-6 in. long, 
bracts long, filiform, hairy; cal.-segm. setaceous, plumose; pod 
nearly 1 in., linear, compressed, pointed, densely hairy, some- 
what compressed between seeds, 

Low country; common. FI. Nov., Dec. 


Also in India, Java, and Trop. Africa. 
Appears to be sometimes cultivated as a pulse. 


60 Leg UMINOSE. [Mucuna. 


25, TERAMNUS, Sw. 


Herbaceous twiner, |. pinnately 3-foliolate, fl. small, in 
very lax racemes; cal. tubular-campanulate, segm. linear; 
pet. nearly equal, slightly exserted; stam. monadelphous, alter- 
nate ones small, abortive; style beardless, stigma capitate; 
pod linear, with a strongly hooked point, 8—10-seeded, septate 
between seeds.—Sp: 4; 2 in FZ. B. Lnd. 


T. labialis, Srene. Syst. iii. 235 (1836). 

Glycine parvifiora, Lam., Moon Cat. 53 (?). Glycine labialis, L. f.,, 
Thw. Enum. 88. Glycine mollis, W. and A., Am. Pug. 12. TZ. mollis, 
Benth. in Journ. Linn. Soc. viii. 265. C. P. 1471. 

Fl. B. Ind. ii. 184. Wight, Ic. t. 168. 

A twining herb, stems slender, much-branched, glabrous or 
with deflexed hair; l.-rachis 1-24 in., glabrous or hairy like 
the stem, stip. minute, deciduous, Iflts. 14-3 in., lanceolate or 
oval-lanceolate, subacute, apiculate, glabrous on both sides or 
softly pubescent beneath, lat. ones unequal-sided; fl. small, on. 
short slender ped., arranged in small clusters in usually very 
lax elongated slender axillary racemes, from 2-9 in. long, 
bracts small, setaceous; cal. hairy, segm. setaceous; pod 
14-2 in., narrowly linear, straight or slightly curved upwards, 
tipped with a short stout beak or hook abruptly curved 
upwards, hairy when young but nearly glabrous when mature; 
seeds 8-10, cylindrical, truncate at ends, dark brown. 

Low country; common. FI. Nov.—Dec.; pinkish or white. 

Found throughout the Tropics. 


A very variable plant, perhaps including more than one species. The 
hairy form is 7. mollzs, Benth. 


26. MUCUNA, Adazns. 


Woody or herbaceous twiners, |. 3-foliolate, with stipels, f1.. 
large, in drooping racemes; cal. campanulate, 2 upper segm.— 
connate, lowest one longest; pet. exserted, very unequal, wings. 
twice as long as standard, keel rather longer than wings,. 
curved into a stiff beak at apex; stam. diadelphous; style 
beardless, stigma capitate; pod 1-few-seeded covered with 
sharp needle-like bristles, slightly compressed, turgid; seed 
large, with an oblong or linear hilum.—Sp. 20; 10 in FZ. B. [nd. 


Pod winged on both sutures and sides. 
Cal. truncate with small teeth. 5 : . I. M. MONOSPERMA. 
Cal. 2-lipped, with large lower segm. M. ATROPURPUREA.. 
Pod winged at sutures only. M. GIGANTEA. 
Pod without wings (Stézolobium). M. PRURIENS. 


RWS 


Mucuna.] Leguminose. 61 


1. WZ. monosperma, DC. Prod. ii. 406 (1825). 

Thw. Enum. 89. C. P. 2662. 

Fl. B. Ind. ii. 185. Wight, Ic. t. 35 (2 plates). 

A large woody twiner, the younger branches clothed with 
rufous, deciduous tomentum; |. large, rachis 24-44 in., with 
red deciduous pubescence, stip. deciduous, Iflts. on short swollen 
stalks, 2-4 in., rotundate or broadly oval, shortly acuminate, 
smooth above, more or less densely pubescent beneath, lat. 
ones unequal-sided; fl. large 1% in., on divaricate ped. $ in. 
long, 6-10 in a lax pubescent raceme (or panicle) shorter than 
1.; cal. sparingly clothed with red bristles, upper segm. wanting; 
standard often with a few bristles on the back; pod 2 in, 
broadly ovate-ovoid, shortly stalked, somewhat curved, with a 
short decurved beak, with a broad double horizontal wing 
along both sutures, and several (about 6) broad erect distant 
wings extending from them at right angles nearly half way 
down the sides, the whole covered with orange-red deciduous 
wool mingled with irritant red bristles; seed solitary, nearly 
circular, compressed, dark brown, hilum linear, extending 
round # of the edge. 

Upper zone of moist low country; rather rare. Kitulgala; Hunas- 
giria; Lunugala; Uva. Fl. Jan.; bright violet, keel green. 

Hills of India and Burma. 

This is probably Stzolobium rugosum, Moon Cat. 53, from Kandy. 
Our plant has the sutures of the pod very distinctly double-winged, 
and thus does not well agree with published descriptions of JZ. mono- 
sperma. 


2. MZ. atropurpurea, DC. Prod. ii. 406 (1825). Punnakalichi, 
Phandatullai, 7. 

Thw. Enum. 89. C. P. 1483. 

Fl. B. Ind. ii. 186. 


A large woody twiner, young branches slender, glabrous; 
l.-rachis 2-3 in., glabrous, stip. deciduous, lflts. shortly stalked, 
24-4 in., lanceolate or oval, acuminate, acute, glabrous on 
both sides, lat. veins very oblique; fl. very large, 24-23 in., 
on drooping ped. 4-7 in., about 10-15 in a short racemose 
panicle; cal. large, densely downy, with a very few orange 
bristles, 3 lower segm. triangular, equal; pod 3-4 in., oblong- 
ovoid, scarcely stalked, blunt, doubly winged along sutures, 
and with numerous (10-14) deep, close, parallel, flat wings or 
plaits running obliquely from each across sides, and meeting 
along centre, slightly pubescent and clothed with copious red 
irritant bristles; seeds 2 (or 1), 1in., broadly ovoid, slightly 
compressed, grey mottled with brownish-black, hilum ex- 
tending 4 round edge. 


62 Leguminose. [Mucuna. 


Dry region; rather rare. N. Province (A. Clark); Batticaloa 
(Gardner); Hunguranketa; Trincomalie (C. La Brooy); Bintenne Dis- 
trict. Fl. Feb.; rich blue-purple. 

Also in Peninsular India and at Malacca. 

The same remark as to the pod applies to this as to the last. The 
sutures lie at the bottom of a very deep furrow formed by the wings, 
which are j in. or more high. 


3. M. gigantea, DC. Prod. ii. 405 (1825). 
Stizolobium giganteum, Pers., Moon Cat. 53. Thw. Enum. 89. C. P. 


3587. 
Fl]. B. Ind. ii. 186 (not given for Ceylon). Hk. Bot. Misc. ii. t. 


supp. 14. 

A large woody twiner, with glabrous branches; Ifits. ovate, 
acute, glabrous when mature; fl.on long slender ped., almost 
umbellate at end of long peduncles; pod 4-6 in., stalked, 
linear-oblong, apiculate, with a broad double wing along both 
sutures, but without wings or plaits on the sides, densely 
covered with long adpressed chestnut-brown, irritant bristles. 

Dry country; very rare. I have seen only the C. P. specimen (a pod 
merely), which was collected by Gardner, and is labelled ‘ Batticaloa’ (?). 
Moon’s locality is Bentota. It grows in the Bot. Gardens, Peradeniya. 
F]. Jan.; sulphur-yellow. 

Also in Peninsular India, Malaya, Polynesia, &c. 


4. M. pruriens, DC. Prod. ii. 405 (1825). Achariya-pala, S. 

Burm. @hes. 191. Herm. Mus. 67; EI Zeyl. n. 5305 eves 
pruriens, L. Sp. Pl. i. 1019. Stzzolobtum fruriens, Pers., Moon Cat. 53. 
M. prurita, Hook., Thw. Enum. 89. C. P. 3150. 

Fl. B. Ind. ii. 187. Hk. Bot. Misc. ii. t. suppl. 13. Bentl. and Trim. 
Med. Pl. i. t. 78. 

An annual (?), semi-woody twiner, branches slender, usually 
clothed with short white deflexed hairs; 1. large, rachis 3-5 in., 
sparingly deflexed-hairy, stip. linear-setaceous, hairy, lflts. 3-4 
in.,on short thick hairy stalks, terminal one smallest and rhom- 
boid-oval, lat. ones very unequal with the lower half greatly 
dilated, all acute, mucronate, pubescent above, densely covered 
with shining silvery adpressed hair beneath; fl. numerous, 
14-1? in. long, on short pubescent ped., usually 2 or 3 together 
at intervals in a slender pubescent raceme 6-12 in long, bracts 
4 in. lanceolate, hairy, soon falling; cal. densely silky, 2 upper 
segm. completely connate, lowest much longest; pod 24-3 in. 
by about 4 in. broad, linear, blunt, falcately curved at both 
ends, with a longitudinal rib along whole length of each valve 
but without wings, densely covered with close rather weak 
orange-brown irritant bristles pointing backward and readily 
detached, 4-6-seeded with partitions between them; seed 
ovoid, + in., compressed, brownish mottled with black, hilum 
oblong, not half the length of seed. 


Erythrina.] Leguuinose. 63 


Var 6, biflora, 7vzm. Cat. Ceyl. P/. 25. 


L. smaller, more hairy above; peduncles about 1 in., erect, 
2-flowered. 

Dry and intermediate regions; rather common. Var. , Batticaloa. 
Fl. Jan., Feb.; dull dark purple, the keel yellowish-green. 

Found throughout the Tropics. 

This is the ‘ Cowage’ or ‘Cowitch’ plant. Moon’s S. name is ‘ Wel- 
damaniya.’ 


27. ERYTHRINA, J. 


Trees with prickles on the bark, 1. pinnately trifoliolate, 
with stipels, deciduous, fl. large in secund racemes; cal. tubular, 
splitting irregularly; pet. very unequal, standard much longer 
than wings or keel, keel- pet. distinct; stam.diadelphous, but the 
upper one often connate with the rest half way up; style beard- 
less, stigma capitate; pod cylindrical torulose, 3-10-seeded ; 
seed large, with a large oval hilum.—Sp. 30; 7 in FZ. B. Ind. 


Lfits. roundish-ovate, green on both sides ‘ Se Lee INDICA. 
Lfits. oval, white beneath . : ; ‘ ‘ . 2. E. OVALIFOLIA. 


1. BE. indica, Zam. Enc. Méth. ii. 391 (1786). Brabadu, S. 
Mullu-murukku, 7. 

Herm. Mus. 49. FI. Zeyl.n. 275. £. corallodendrum, L. Sp. Pl. i. 706 
(in part). Moon Cat. 52. Thw. Enum. 89. C. P. 1464. 

Fl. B. Ind. 11. 188. Wight, Ic. t. 58. 


A tree attaining to a large size, bark thin, smooth, grey, 
young twigs thick, set with small straight, horizontal, broad- 
based sharp black prickles, leaf-scars conspicuous; |. very 
large, deciduous, rachis 6-12 in., smooth, dilated at base, stip. 
none or very early caducous, Iflts. 4-6 in. on short swollen 
stalks, readily disarticulating, roundish-ovate, acute, glabrous 
and green on both sides, the term. one the largest, stipels thick 
roundish persistent; fl. numerous large, on stout puberulous 
horizontally spreading ped. about 4 in. long, arranged in 2’s or 
3's, and closely crowded on the upper half of very stout, rigid 
puberulous racemes 6-12 in. long, 2-4 of which diverge 
horizontally from the summit of the branches; cal. (before ex- 
pansion of fl.) tubular, 14 in., covered with deciduous tomen- 
tum, upper segm. subulate, sharp but not rigid, two lat. similar 
but smaller, lowest one longer, doubled over the rest to form 
a blunt point to the bud, soon splitting along the back 
(between the upper teeth) to the base and the whole turned 
down like a spathe; standard nearly 3 in., wings less than 
1in., keel-pet. # in.; stam. much exserted and projecting in 


64 Leguminose. [ Szrongylodon. 


front of fl., 22in.; pod 5-6 in., cylindrical, distinctly torulose 
with a sharp curved beak; seeds 3-8, bean-like, about I in. long. 


Low country ; very common, and much planted. Fl. Feb.; scarlet, the 
wings and keel crimson. 

Also throughout India and Malaya, Java, and Polynesia. 

When well grown this is a handsome tree; but as usually seen— 
planted for hedges or as a support for climbing plants—it is a shabby 
stunted bush. The brilliant flowers are produced when the branches are 
bare of leaves, and give the plant the name of ‘ Coral-tree.’ 

A variety with pale pink flowers is figured by Moon in his collection 
of drawings in the Brit. Mus. labelled ‘ Ela-erabadu.’ 

The indurated incurved tip of the lowest segm. of calyx prevents the 
bud from opening, and the growing standard forces its point through the 
cal. between the two upper teeth, and gradually splits it down the back. 


2. E. ovalifolia, oxt. Hort. Beng. 53 (1814). YWak-erabadu, S. 

E. picta, Moon Cat. 52 (?L.). Thw. Enum. 89. C. P. 3499. 

His Indie co.  eWicht. les t2777 

A moderate-sized tree with wide-spreading branches, 
trunk covered with large pyramidal, corky, prickle-tipped 
thorns, young branches as in last, but prickles brown, smaller 
and sharper; buds pubescent; |.-rachis 4-5 in., glabrous, 
stip. small, roundish, adpressed to rachis, soon disappearing, 
Iflts. 34-44 in., broadly oval, on swollen stalks readily disarti- 
culating, obtuse, glabrous on both sides, silvery-white beneath, 
stipels wart-like, persistent; fl. rather large, arranged as in 
£.. indica, with 2 small ovate bracts immediately beneath cal., 
cal.campanulate completely closed and ending in a curved beak 
in bud, afterwards splitting irregularly into several unequal 
divisions, pod 6-10 in., shortly stalked, with a sharp beak, 
curved, more or less torulose, very finely but densely downy, 
6—10-seeded with partitions between them ; seed #in., oblong, 
cylindrical, truncate at ends, black. 

Low country, by the banks of rivers and streams ; rather common. 


Fl. January; standard dark crimson, wings and keel purple. 

Also in E. Bengal, Burma, Malaya, and Polynesia. 

[Z. suberosa, Roxb., is given for Ceylon in Fl. B. Ind. ii. 190, but I do 
not know on what authority. There are no specimens at Kew from 
Ceylon. | 

E. lithosperma, Bl., from Java (‘ Dadap’ of the Malays) ; &. umbrosa, 
H. B.K., from Central America (‘Madre de Cacao’); and £. velutina, 
Willd., of the W. Indies are all frequently met with on estates, especially 
of Cacao, planted as shade-trees. 


28. STRONGYLODON, /2.. 


Herbaceous twiner, |. pinnately 3-foliolate, with stipels, fl. 
in lax racemes; cal. campanulate with very short rounded im- 


Galactia.] . Leguminose. 65 


bricate segm.; standard nearly 3 times as long as wings, keel 
longer than wings, prolonged into a beak ; stam. diadelphous, 
anth. uniform; ovary stalked, style beardless, stigma capitate; 
pod ovoid, beaked, stalked, turgid, indehiscent ; seeds 1 or 
2, large, with a carunculate hilum extending more than half 
way round the margin.—Sp. 3; 1 in FZ B. Ind. 

S. ruber, Vogel in Linnea, x. 585 (1836). 

Thw. Enum. 89. C. P. 1465. 

Fl. B. Ind. ii. 191. A. Gray, Bot. Amer. Expl. Exp. t. 48. 

A herbaceous twiner, stems much branched, glabrous; L.- 
rachis 24-4 in., slender, stip. small, triangular, lfits. stalked, 
24-4 in., ovate-lanceolate, acuminate, apiculate, glabrous on 
both sides, thin, stipels linear; fl. about 1 in. long, on slender 
spreading ped. 4—? in. long, numerous, forming a lax pyra- 
midal raceme 3-5 in. long at end of long stout axillary 
peduncles, bracts small, orbicular, one at base of ped. and 2 
immediately beneath fl.; cal. glabrous, gibbous at base above; 
pod rather long-stalked, about 3 in., ovoid, inflated, with a 
sharp curved beak and compressed margin, papery, yellow ; 
seeds I or 2, 1 in., globular-ovoid, slightly compressed, smooth, 
black, caruncle prominent, wing-like, red. 

_ Low country in intermediate region; very rare. Galagedara 
(Gardner); Kurunegala (Thwaites). Fl. June; reddish-pink. 


Elsewhere found only in the Polynesian Islands, and recently in S. 
Andaman I. 


29. GALACTIA, P. Br. 


Very slender twining herbs, |. pinnately 3-foliolate, with 
stipels, fl. small, in lax racemes with very slightly tumid nodes; 
cal. with short campanulate tube and linear segm., the 2 upper 
completely connate, the lowest largest; pet. nearly equal ; 
keel nearly straight; stam. diadelphous; style beardless, 
stigma capitate, pod linear, flat, recurved.—Sp. 45; 2 in 
Fi. B. [nd. 

G. tenuifiora, W. and A. Prod. 206 (1834). 


Glycine tenuiflora, Willd., Moon Cat. 53. Thw. Enum. 88. C. P. 1468. 

Fl. B. Ind. ii. 192. 

A herbaceous twiner, stems filiform, with scanty adpressed 
hairs; l.-rachis #—-1}4 in., slender, pubescent, stip. minute, lflts. 
small, 1-1% in., lanceolate or oval-lanceolate, subacute, apicu- 
late, glabrous above, paler and silky-pubescent beneath ; fl. 
small, few, shortly stalked, at end of very slender, lax, pubes- 
cent axillary racemes, bracts small, caducous; cal. slightly 


PART II. F 


66 Leguminosae. [Butea. 


hairy ; pod 14-2.in. by } in. broad, slightly curved upwards, 
apiculate, at first silky, becoming glabrous ; seeds 6-9, small, 
compressed, black. 

Upper zone of low country, rather common; Kotmalie; Hantane ; 
Matale. FI. pink. 

Also in India, Siam, Malay Is., Australia, and E. Trop. Africa. 


The nodes of the inflorescence can scarcely be said to be tumid in 
the Ceylon species. 


[Spatholobus Roxburghit, Benth., is represented in Herb. Kew by 
specimens of Walker’s (n. 1331) professing to be from Ceylon, and the 
plant is entered for this country in Fl. B. Ind. ii. 193. They are not 
further localised, and, as no one else has met with this conspicuous woody 
climber, some error is probable. The species, however, occurs in India 
and Burma; it is figured in Wight, Ic. t. 210 (as Butea parvifiora).| 


30. BUTEA, ox. 


An erect tree, 1. pinnately 3-foliolate, with stipels, fl. large, 
fascicled, in racemes with swollen nodes; cal. shallowly cam- 
panulate, segm. shallow, 2 upper connate, 3 lower equal; pet. 
nearly equal, keel-pet. completely connate, much curved ; 
stam. diadelphous, the free fil. shorter than the rest; pod 
large, flat, linear-oblong, leathery, the lower ? without seeds 
and indehiscent, the terminal } containing the solitary seed 
and dehiscent round the edge ; seed with a small hilum.—Sp. 
2 rallye a7. 


B. frondosa, Koen. ex Roxb. in Asiat. Researches, iii. 469 (1792). 
Gas-kéla, S. Parasu, 7. 

Moon Cat. 52. Thw. Enum, 89. C. P. 1465. 

Fl. B. Ind. 11. 194. Bedd. FI. Sylv. t. 176. Bentl. and Trim. Med. PI. 
t. 79 (from a Ceylon specimen). 

A moderate-sized or small tree, with a crooked trunk and 
irregular branches, bark thick, rough, ash-coloured, fibrous, 
exfoliating in small pieces, young shoots densely pubescent ; 
l. large, rachis 5-8 in., slender, pubescent when young, swollen 
at base, stip. small, linear-lanceolate, deciduous, Ifits, 4—8 in., 
unequal, the terminal the largest and rhomboid-orbicular, the 
lat. ones obovate-oval, dilated in lower half, all very obtuse, 
glabrous above when mature, closely and finely tomentose 
and with much raised reticulation beneath ; fl. large 14-2 in., 
on velvety drooping ped. ?-1 in. long, 2 or 3 together from 
the swollen nodes of rigid stout racemes coming off from 
woody tuberosities, bracts small, deciduous; cal. finely velvety 
outside, lined with white silvery hair, segm, acute; keel very 
deeply boat-shaped acute; pod pendulous, 5-8 in., by about # 
in wide, on a densely woolly stalk ?in. long, obtuse, thickened 


Canavalia. Leguminose. 67 


at the sutures, leathery, transversely veined, densely but finely 
pubescent especially at end; seed Ifin., flat broadly oval, 
smooth, reddish brown. 

Dry region in open country; rather rare, but locally abundant. 
Jaffna; Batticaloa; Bintenne; about Bibile gregariously. Moon’s locality 
is Wellasse. FI. July; orange-scarlet, very silvery outside with silky 
hair, so that the buds are white. 

Also in India and Burma. 

A dark red astringent juice exudes from the bark, which hardens into 
a brittle, ruby-coloured, resinous-looking gum, the ‘Bengal Kino’ of 
commerce. Wood whitish, soft, light, not durable. 

[Moon l.c. gives also 4. sugerba, Roxb., with the name ‘ Wel-Kéla’ 
and the locality “ Kandy.’] 


31. CANAVALTIA, DC. 


Perennial herbs, |. pinnately 3-foliolate with stipels, f1. ire: 
in erect racemes with large swollen nodes; cal. deeply 
campanulate, 2 upper segm. connate to form a rounded 
projecting lip, 3 lower small, equal, triangular; standard rather 
longer than wings, keel curved, obtuse, equalling wings; stam. 
monadelphous, style beardless, stigma terminal; pod large, 
linear or oblong, somewhat compressed, with a prominent rib 
on each side close to dorsal suture; seeds several, with a short 
linear hilum.—Sp. 12; 2 in FZ. B. Ind. 


Lfits. oval, thin : F : , . I. C. ENSIFORMIS. 
Lfits. broadly obovate, thick. : : 5 , 2, C, OBUUSMROMINN, 


1. ©. ensiformis, DC. Prod. i. 404 (1825). Wal-awara, S. 
Koli-avarai, 7. 

Dolichos gladiatus, Jacq., and D. rotundifolius, Moon Cat. 53. C. 
£ladiata, DC., Thw. Enum. 88. C. P. 1481. 

Fl. B. Ind. 11. 196. Wight, Ic. t. 753. 


A stout perennial twiner, stems long, glabrous or nearly 
so, young shoots pubescent; l.-rachis about 3 in., channelled 
above, stip. inconspicuous, deciduous, Iflts. about 3 in., shortly 
stalked, oval, rounded at base, very shortly acuminate, 
apiculate, glabrous on both sides, thin, reticulately veined; fl. 
numerous, large, 1in. long, on very short ped., generally i in 
pairs from the tumid nodes of stout, curved, erect long- 
stalked racemes, bracts minute, caducous ; cal. very finely 
pubescent, upper lip with 2 rounded lobes ; pod shortly 
stalked, about 6 in. by nearly 1 in wide, straight or very 
slightly curved upwards, shortly pointed, smooth; seeds 8-10. 


Var. 6, virosa, Laker. Dolichos virosus, Roxb., Moon Cat. 53. C. 
virosa, W. and A., Thw. Enum. 89. C.P. 2782. 


68 Leguminosae. _ [Dioclea. 


Lfits. broader, more acuminate; pod shorter and broader, 
4in. by 14 in wide, oblong, 5-or 6-seeded. 

Dry region; rather common. Jaffna; Anuradhapura; Kekirawa; 
Naula. Var. 8, moist low country up to 3000ft.; common. Fl. Dec., 
Feb., August; bright pink or nearly white. 

Throughout the Tropics, wild or cultivated. 

This seems truly native in Ceylon, though the type-form is also largely 
grown asa pulse. The English call it the ‘ Sabre-bean.’ 

The stam. are figured as diadelphous in Wight, Ic., but I find them 
always monadelphous in flowers examined. 


2. ©. obtusifolia, DC. Prod. ii. 404 (1825). Mudu-awara, S. 
Thw. Enum. 88. .C. P. 1484. 
Fl. B. Ind. 1. 196. 


Perennial (? biennial), stems spreading, prostrate, rarely 
twining, glabrous, young shoots slightly silky; 1. rather small, 
rachis 2-34 in., stip. small, caducous, lfits. 1-2 in. broadly 
obovate or roundish, quite glabrous, thick; fl. rather smaller 
than in the last, few, in erect racemes longer than 1.; cal. 
pubescent; pod about 4 in., taperimg at base into long stalk, 
* in. wide, shortly beaked, nearly straight, glabrous; seeds 5 
or 6, ovoid, compressed, over 4 in., black when dry. 


Sandy seashores; common. FI. Jan., July, August; bright pink. 

Throughout the Tropics. 

A truly seashore plant. The flowers are sometimes white. The seeds 
are much eaten, boiled, at Batticaloa and elsewhere. 

I doubt if the figure of Rheede, Hort. Malab. viii. t. 43, can be referred 
to our plant, but the species is very variable. It is possibly the original 
wild state of C. enszformzs. 


32. DIOCLEA, Z. 2.4K. 


Woody twiner, |. pinnately 3-foliolate, with stipels; f1. 
moderate-sized, on short thickened branchlets of racemose 
panicles; cal. deeply campanulate, upper segm. connate for 
nearly whole length, lower ones narrow-triangular the lowest 
the longest ; pet. about equal, keel not beaked ; stam. mona- 
delphous, the top one free below; style beardless, stigma 
capitate; pod large, oblong-falcate, compressed, flattened on 
dorsal suture with a narrow horizontal wing along each side; 
seed large, flat, hilum narrow, linear, extending round more 
than 2 of the edge.—Sp. 16; 1 in FZ. B. Lud. 


D. reflexa, “2. 7. in Niger Flora, 306 (1849). 
D. Fergusoniz, Thw. Enum. 412. C. P. 3817. 
Fl. B. Ind. 11. 1096. 


Phaseolus.] Leguminose. 69 


A stout woody twiner, branches wide-spreading, the young 
ones covered with dense soft fulvous hair; |. large, rachis 
3-4 in., fulvous-hairy, stip. small, roundish attached by their 
centre, lfits. 4-6 in., ovate or oval, rounded at base, somewhat 
attenuate, subacute, apiculate, glabrous above when mature, 
softly silky beneath ; fl. about #in., numerous, on short ped., 
2 or 3 together on very short incurved swollen branchlets of a 
narrow elongated racemose panicle 12-18 in. long, bracts 
below ped. much longer than them, linear, recurved, bracts 
beneath fl. 2, small, roundish; cal. fulvous-pubescent; pod 
about 5 in. by 2 in. wide, slightly falcate, at first hairy, glabrous 
when mature; seeds 2 or 3, separated by spongy partitions. 
orbicular, I-14 in., much compressed, black shining. 

Moist low country; very rare. About Kaduwela near Colombo, the 
only known locality ; found by W. Ferguson in 1863. FI. pale violet. 

Also in E. Bengal, Burma, Malaya, and Trop. Africa, and America, 


but not in the Indian Peninsula. 
The seed is very like that of a Mucuna. 


33. PHASEOLUS, L. 


Erect, prostrate or twining herbs, 1. 3-foliolate, with stipels, 
fl. in axillary racemes, bracts conspicuous; cal. campanulate, 
2 upper segm. connate, nearly equal or lowest one longest; 
keel-pet. extended into a long stiff beak curved round into a 
circle; stam. diadelphous, anth. uniform; style curved round 
with the keel, conspicuously bearded along the inner side below 
the very oblique stigma; pod linear, somewhat compressed or 
nearly cylindrical, with partitions between the numerous 
seeds.—Sp. 60; 15 in Fl. B. [nd. 


Stip. attached by their base ; fl. not yellow. 


Twining. 
Fl. pink . 1. P. ADENANTHUS. 
Fl. violet-purple 2. P. GRAHAMIANUS. 
Erect ; fl.red . 3. P. SEMIERECTUS. 
Stip. peltately attached ; fl. yellow. 
Lfits. more or less 3- -lobed Ce also 7). 
Stip. large, oval ; AA ea URILOBUS, 
Stip. small, oblong. : ; . §. P. ACONITIFOLIUS. 
Lfits. not lobed (except sometimes 7) 
Pod with long silky hair : ; 3 06s bs VLA, 
Pod with short fulvous hair . , ; . 7. P, TRINERVIUS. 
Pod glabrous . : ; : 3 } oy I CALCAR AI US: 


Two species are commonly cultivated in native gardens under the 
name of ‘ Dambala’ or more usually the Dutch name ‘Bonchi,’ P. /wnatus, 
L. and P. vulgaris, L., the former under several varieties. Both are 


7O Leg UMIMNOSE. [ Phaseolus. 


probably of American origin but now common throughout the Tropics, 
and the latter, the French or haricot bean, ‘ Potu-bonchi,’ S., also in 
temperate regions. 


Le adenanthus, Meyer, Prim. Fl. Esseg. 239 (1818). Wal- 
mé, S. 
P. traxillensis, H.B.K., Thw. Enum. 89. C. P. 1480. 

Fl. B. Ind. 11. 200. Wight, Ic. t. 34 (P. vostratus). 

A perennial herbaceous twiner with glabrous stems; 1. 
distant, rachis 3-4 in., stout, stip. small, oval, lflts. 2-44 in., 
rhomboid-oval, subacute, apiculate, glabrous or slightly 
pubescent, the lat. ones dilated in lower half; fl. large, over 
I in., few, on short ped. with a large round gland at base, 
solitary or in pairs, closely placed at end of an axillary 
peduncle about as long as 1., bractlets 2, immediately beneath 
fl., ovate, acute, striate, cal. finely pilose; keel very long, 
twisted into a complete spiral; pod 5-6 in. by 2 in. wide, 
compressed, shortly beaked, falcately curved, glabrous ; seeds 
12-16, nearly circular, } in., flat, black. 

Moist region, up to 4000 ft.; rather common. FI. Dec.; dark pink. 

Throughout the Tropics. 

The flowers are very large and handsome, and the plant well worth 


cultivation for ornament. This may be the plant called P. Caracalla by 
Moon (Cat. 52), with the English name ‘ Snail-flower.’ 


2. P. Grahamianus, VV. and A. Prod. 244 (1834). 

Thw. Enum. go. C. P. 3586. 

IML 18 Mask, i, Zo, 

A perennial twining herb, with slender glabrous much- 
branched stems; |. small, rachis 14-2 in., stip. small, oval, 
spreading or reflexed, lflts. 14-2 in., rhomboid-oval, obtuse, 
glabrous, readily disarticulating ; fl. medium-sized, on short 
slender ped., 1-3 together on thickened nodes, racemes 
longer than 1.; cal. glabrous; standard very large, keel 
curved into a complete circle; pod 3% in. by ¢ in. broad, 
narrowly linear, beaked, slightly curved upwards at end, 
sutures thickened ; seeds 8-10, oblong, truncate, slightly 
compressed. 

Low country; rather rare. Batticaloa; Galle; Dambulla; Nalande. 
Fl. Jan. Feb.; pale bright violet-purple (not ‘reddish,’ as given in Fl. B. 


Ind.). 
Also in Southern India. 


3.* BP. semierectus, LZ. Mami. i. 100 (1767). 

Thw. Enum. go. C. P. 3609. 

IDL, 1B), line. Wi, Boi,  WWikelnte, Ne, ft BAO), 

An erect semi-shrubby annual, 2-3 ft. high, with few 
cylindrical ascending glabrous branches; l.-rachis 2-24 in., 


Phaseolus.] Leguminose. 7 


stip. linear, subulate, persistent, lfits. 11-2} in., oblong-ovate 
or oblong-lanceolate, acute, glabrous on both sides, readily 
disarticulating ; fl. over I in., nearly sessile, usually in pairs 
distantly placed on elongated erect, rigid, spike-like racemes 
12-18 in. long, bracts long, setaceous; cal. pubescent, segm. 
aristate shorter than tube; standard shorter than wings ; pod 
3-4 in. by = in., nearly cylindrical, straight, sharply beaked, 
somewhat silvery with adpressed white hairs ; seeds about 20 
small, quadrate-reniform, dull dark brown. 

Low country, common; a weed in waste ground. FI. Jan., July, &c.; 
dull purplish-red. 


Also in India, Malaya, and Trop. America. 
This looks like an introduction in Ceylon. 


b) 


4. P. trilobus, Azz. Hort. Kew. iii. 30 (1789). Bin-mé, S. 
Narippayaru, 7. 

Moon Cat. 52. Thw. Enum. go. C. P. 1474 (pt.). 

Fl. B. Ind. 11. 201. Wight, Ic. t. 94. 

Perennial with a woody rootstock giving off very numerous, 
long, prostrate, wiry, slender, glabrous stems, not at all 
twining; 1. small, rachis 1-14 in, stip. peltately attached, 
large, oval, acute or obtuse, Iflts. +? in., stalked, rhomboid- 
rotundate, obtuse, apiculate, glabrous, often 3-lobed with the 
middle lobe broader and spathulate; fl. small, very shortly 
stalked, 2 or 3 together at the end of stiff slender peduncle 
2—5 in. long and much exceeding 1., bracts ovate, acute; cal.- 
segm. short, deltoid; pod small, 1-1$ in., narrowly linear, 
slightly compressed, straight, glabrous; seeds 6-12, very 
small, cylindrical, truncate at ends, dark brown. 

Dry region, common; also on the seashore. Fl. Feb., Sept., &c.; 
yellow. 


Also throughout India, Malay Is., and E. Trop. Africa. 
This may be cultivated as a pulse in the dry districts to a small extent. 


5. P. aconitifolius, Jacg. Obs. Bot. iti. 2 (1768). 


C. P. 1474 (part). 0 
F]. B. Ind. ii. 202. Duthie, Crops N.W. Prov. t. 1. 


Annual or perennial, stems as in the last, but usually 
longer and copiously covered with spreading or reflexed hairs ; 
Ifits. larger, always deeply cut into 3 narrow lobes, hairy 
beneeth, stip. small, oblong, obtuse; fl. as in P. ¢rilobus, 
bracts lanceolate-oblong, acute, setaceous; pod 1-2 in, 
usually rough with few adpressed hairs; seeds 8-14, rather 
larger than in the last. 

Dry region; rather common. Jaffna; Batticaloa; Bintenne; Kanta- 
lai; Bibile. Fl. Jan., August, &c., yellow. 

Also in Peninsular India. 


72 Leguminose. [Phaseolus: 


Dr. Thwaites did not distinguish this from P. ¢vz/odus, and it is scarcely 
separable as a species. I have never seen this in cultivation in Ceylon, 
but suspect it is sometimes grown in the North. 


6. P. Max,* Z. Sf. Pl. 725 (1753). Mun, Mun-éta, Bu-mé,, 
S. Chiruppayaru, 7. 

Herm. Mus. 22. Fl. Zeyl: n. 280, Moon Cat: 52.” Aigo 
Moon Cat. 52. 

F]. B. Ind. ii. 203 (P. Mungo). Duthie, Crops N.W. Prov. t. 9. 


An annual herb, stem erect, about 2 ft., branching, an- 
gular, clothed with spreading or deflexed hairs; |.-rachis long, 
hairy like the stem, stip. peltately attached near the base, 
ovate, acuminate, lfits. 2-3 in., ovate, acute, more or less hairy 
on both sides, thin, the lat. ones dilated on lower side; fl. about 
4 in. shortly stalked, 4-6 crowded at end of stout hairy 
peduncles shorter than |.; cal. silky, lowest segm. lanceolate;. 
standard much broader than long, keel curved into more than 
a circle; pod 2-24 in., linear, nearly cylindrical, shortly beaked, 
more or less hairy with spreading hair; seeds 8-12, small 
4-7 in., oblong-ovoid, truncate or blunt at ends, green or 
blackish. 


Var. 6, radiatus, Baker in Fl. B. Ind. \.c. UWlundu, S. 
Uluntu, 7. Herm. Mus. 47. Fl. Zeyl. n. 281. P. radiatus, L. Sp.. 
Pl. 725; Moon Cat. 52; Thw. Enum. 89, 412. C. P. 3622. 

Dill. Hort. Elth. t. 235, f. 304. Duthie, Crops N.W. Prov. t. Io. 


Larger, stems more trailing or slightly twining, usually 
more hairy, peduncles longer, generally exceeding |.; seeds 
longer, + in., dark brown or black. 


Commonly cultivated, and occasionally found apparently wild in the 
dry region. Fl. Nov.—Feb.; pale yellow. 

Throughout the Tropics, but usually cultivated. 

The seeds are known as ‘Green Gram’ and ‘ Black Gram’ respect-. 
ively; there is also a variety with yellow seeds. 

P. Maz is the older of Linnzeus’s two names, but authors usually have 
preferred P. Mungo for the collective species, and restrict P. ax to the: 
black-seeded form. Very variable in habit and amount of hair. Her- 
mann’s specimens of P. (Zax have very woolly leaves. 


7. B. trinervius, Heyne in Wall. Cat. n. 5603 (1828). 
Thw. Enum. 90. C. P. 1476. 
Fl. B. Ind. ii. 203. 


A perennial twining herb, stems clothed with deflexed! 
fulvous hair, internodes very long, |. rather large, rachis 2—34in., 
hairy like the stem, stip. peltately attached, oblong-lanceolate,. 
acute, hairy, lflts. 2-34 in., term. one rhomboid-oval often some- 


* Mav is the Spanish name in Mexico, as given by Hernandez. 


Vigna. Leguminose. Te 


what 3-lobed, conspicuously 3-nerved from base, lat. ones ovate 
or lanceolate, much dilated in lower side, all acute, silky-hairy on 
both sides; fl. $in., very shortly stalked, in short close racemes 
with swollen nodes, at end of stiff hairy peduncles much 
exceeding 1.; cal. nearly glabrous, segm. very short nearly 
equal; keel curved into nearly a circle; pod 2-2? in., cylindrical, 
straight, bluntly pointed, rather densely covered with brown 
hair ; seeds 8-12, about $in., oblong-oval with truncate ends, 
dark brown. 

Moist region, I-4000 ft. in grassy places; common. FI. March, 
Aug., Nov.; yellow. 

Also in India and Malaya. 


8. BP. calcaratus, Aoxb. Hort. Beng. 54 (1814). 
Gaweenum: 412. C. P. 1473, 
FI. B. Ind. 1. 203. 


A small perennial herb, stems flexuose, prostrate at base 
and often rooting at nodes, or twining, with scanty reflexed 
hairs; 1. distant, rachis 14-2 in., with deflexed hairs, stip. 
peltately attached, linear-lanceolate, Iflts. usually 14-24 in. 
(rarely 34), ovate, acuminate, acute, hairy on both sides, very 
thin, lat. ones dilated on lower side; fl. small, 4in., on slender 
crowded ped., racemes at first very short but lengthening to 
14 in., at end of long slender peduncles greatly exceeding l., 
bracts linear, longer than cal.; glabrous, segm. triangular ; 
pod 14-24 in., linear, straight, beaked, glabrous; seeds 8-12, 
oblong, narrower than in last . 

Moist region, I-4000 ft.; common in grass. FI]. Sept.-Nov.; yellow. 


Throughout India and Malaya. 
Neither this nor P. ¢77mervius are ever cultivated in Ceylon. 


34. VIGNA, Savi. 


Prostrate or twining perennial herbs, |. 3-foliolate with 
stipels, fl. in short axillary racemes; cal. campanulate, segm. 
nearly equal, upper not connate; keel either truncate or 
curved and beaked; pod much as in Phaseolus ; from which this 
Genus is scarcely distinct.—Sp. 40-50; 10 in FZ, L. Lund. 


Fl. yellow, keel not beaked . ; ; ; : 3 oe , LUDROLA, 
Fl. pink, keel beaked . ; ; j ; : . 2. V. VEXILLATA. 


1. V.luteola, Lenth. in Thw. Enum. 90 (1859). 
ae luteus, Moon Cat. 53 (non Sw.). Thw. Enum. go. C. P. 
1482. 
Fl. B. Ind. ii. 205. Mart. Fl. Bras. xv. t. 50, f. 2. 


74 Leguminose. [Clitorda. 


A perennial herb, branches trailing, prostrate, quite 
glabrous; l|.-rachis 2-3 in., swollen at base, stip. small, 
lanceolate; Ifits. stalked, equal, 14-24 in., broadly oval or 
somewhat obovate, narrowed at base, rounded at apex, quite 
glabrous; fl. about 4 in., on slender ped. longer than cal., in 
short racemes at end of stiff axillary peduncles usually exceed- 
ing |.; cal.-segm. short, broad ; keel not beaked; pod 2-24in., 
rather broadly linear, subcylindrical, turgid, with thickened 
sutures, slightly curved, apiculate, glabrous or nearly so; 
seeds 5-8, $in., nearly globose, black. 

Moist region, on or near the coast; rare. Galle (Gardner); Kalutara 


(Moon); Colombo. FI. July; yellow. 
Found in Bengal, Burma, and the Tropics generally. 


V. Catiang, Endl., Wil-mé and Lz-mé, S., Kodippayam, T., and its 
twining variety, Y. szwenszs, Endl., Mé-karal and Wanduru-mé, S., are 
commonly grown in native gardens. Both were known to Hermann. For 
figures see Duthie, Crops N.W. Prov. tt. 29, 30. The pods of V. sznenszs 
are often 16 in. to 2 ft. long, the large flowers pale purple and white, and 
the seeds either white or pale reddish-brown. 


2. V. vexillata, Benth. in Thw. Enum. 90 (1859). 
Thw. Enum. 90. C. P. 2781. 

Fl. B. Ind. ii. 206. Dill. Hort. Elth. t. 234, f. 302. Wight, Ic. t. 202 
(V. pulnetensis). 

A perennial herbaceous twiner, stem long, stout, glabrous, 
the young parts with deflexed hair; l.-rachis 1-2 in. slightly 
hairy, stip. small, lanceolate, acute, hairy, Iflts. 14-24, narrowly 
ovate-lanceolate, rounded at base, gradually tapering to acute 
cuspidate apex, finely hairy on both sides; fl. large, over 1 in., 
on very short ped., few, crowded at end of axillary peduncles 
longer than 1, cal. puberulous, segm. triangular with long 
setaceous joints; standard very large, keel obliquely curved 
into # of a circle, beaked ; pod (not seen) “ 3-4 in. by ¢ in., 
slender, thinly silky or glabrescent, 10-15 seeded.” 


Moist region, I-4000 ft.; rather common. Fl. August; rose-coloured. 
Cosmopolitan in the Tropics. 


35. CLITORIA, Z. 


Perennial twiner, 1. imparipinnate, with stipels, fl. large, 
axillary, solitary; cal. long tubular, segm. nearly equal; 
standard much exceeding other pet., keel slightly curved ; 
stam. diadelphous; style flattened, bearded along upper side, 
stigma terminal ; pod linear, flat, many-seeded.—Sp. 27; 5 in 
FUSE. Ind. 


Dolichos.] Leguuinose. 75 


Cc. ternatea, ZL. Sf. Pi. 753 (1753). Nil-katarodu, Kata- 
rodu-wel, S. Karuttappu, 7. 

Herm. Mus.10. Burm. Thes. 100, tor. FI. Zeyl. n. 283. Moon Cat. 
53. lhw. Enum. 88. C. P. 1485. 

Fl. B. Ind. ii. 208. Bot. Mag. t. 1542. 


A perennial twining herb, stems long, cylindrical, with 
scanty adpressed hair; 1. numerous, rachis 1$—-2 in. stip. very 
small, linear, lflts. 5 or 7 (2 or 3 pair and end one), 3-14 in., 
oval or oblong-oval, shortly stalked, obtuse often emarginate, 
glabrous above, slightly hairy beneath; fl. large, over I in. 
long, on a drooping, short, jointed axillary ped., with 2 large 
erect rotundate bracts immediately beneath it; cal. ? in., thin, 
nearly glabrous, segm. long-triangular, acuminate; standard 
very large, somewhat reflexed, bifid; pod 4-5 in. by 3 in. 
wide, nearly straight, sharply beaked, with a few adpressed 
hairs; seeds 8-12, 4 in., oblong-ovoid, greenish mottled with 
brown. 

Dry region, common, and perfectly wild; in moist region usually the 
result of cultivation. Fl. Dec., Jan.; intensely bright blue. 

Throughout the Tropics, but often cultivated. 


A lovely climber; the flowers are often much larger in cultivation, and 
are sometimes white. A double-flowered variety also is cultivated. 


Centroseama Plumierd, Benth., is quite naturalised in the moist low 
country in many places, as about Colombo. It is a native of Tropical 
America. 


Periandra Berteriana, Benth., is another’ weed not unfrequent about 
Kandy; a native of the W. Indies. 


36. DOLICHOS, Z. 


Perennial twining herbs, 1. 3-foliolate with stipels, fl. solitary 
or racemose; cal. campanulate, upper segm. connate, lower 
longer, nearly equal; pet. nearly equal, keel not curved into a 
ring, beaked or not; stam. diadelphous; style either bearded 
along inner side or with a tuft of hair round terminal stigma; 
pod more or less curved, usually much flattened, few-seeded.— 
Sp. 20; 6in Fl. B. Ind. 

Style flattened, strongly bearded along inner edge . 1. D. LABLAB. 
Style filiform, not bearded on inner edge. 
Fl. yellow. 
L. not ciliate 


L. strongly ciliate 
Fl. crimson 


. D. UNIFLORUS. 
PIO, fey ny AN is}, 
. D. FALCATUS. 


WN 


76 Leguminosae. [Dolichos.. 


1. D. Lablab,* ZL. Sp. Pl. 725 (1753). Tatta -payaru,, 
Minni, 7. 

Herm. Mus. 60. Burm. Thes. 191. WD. albus and D. purpureus, 
Moon Cat. 53. Lablab vulgaris, Savi, Thw. Enum. go. C. P. 1470. 

Fl. B. Ind. 11. 209. Wight, Ic. tt. 57, 203. Duthie, Crops N.W. Prov.. 
tt. 34, A. and B. 


A perennial twining herb (annual under cultivation), stems. 
stout, wide-spreading, glabrous; |.-rachis 1-2 in., slender, stip. 
small, triangular, Iflts. 1-14 in. broadly ovate-rhomboid, 
broader than long, very acute, glabrous on both sides; fl. 
medium-sized, ped. shorter than cal., several together on short. 
tumid branches of the racemose axillary panicles, peduncle 
long, stout, erect, curved, glabrous; cal. glabrous, 3 lower segm. 
lanceolate; standard with a horizontal thickened claw, keel 
with a straight beak turned up at right angles with the rest; 
style very large, much laterally flattened, also bent up at right 
angles, strongly bearded; pod 14-2 in., by 4—? wide, straight 
along dorsal suture, much curved along ventral edge, suddenly 
truncate, tipped with long deflexed beak, nearly glabrous ;. 
seeds 2 or 3, oblong-oval, compressed, hilum long, linear. 

Low country; probably wild in the dry region, but much cultivated 
throughout the island. Fl. Jan., Feb.; reddish-purple or white. 

Throughout the Tropics of the Old World. 

The above description is from the apparently wild form, but the plant is. 
extremely variable under cultivation in size and shape of pod, and colour 
of flowers and seed. The commonest form in the moist region (ZL. cutra- 
tus, DC.) has purplish-pink fl., a curved, linear, rather inflated tapering pod 
about 3 or 4 in. long, with crisped or warted sutures and white seeds. It 
is the green pod that is used as a vegetable, not the ripe seed. I find no 
recognised S. name. Hermann gives ‘ Hindamini,’ and Moon ‘ Peti- 
dambala,’ and I have had ‘ Pal-avarai’ given me for the wild form, and. 
‘ Dambala’ for the cultivated varieties. 


2. D. uniflorus, Lam. Encycl. Meéth. ii. 299 (1798), var. glaber;, 
Thw. ms. 

Gihw. Enum./90, GC. P. 1475. 

Fl. B. Ind. 11. 210 (under D. dzflorus). 


A perennial twining herb with a woody base, stems slender,. 
with very long internodes, almost completely glabrous; |. small, 
distant, rachis ?# in., slender, stip. oval, acute, ribbed, persistent, 
Iflts. #-1 in., ovate-lanceolate or lanceolate, acute, quite gla- 
brous, thin; fl. ¢in., on very short ped., solitary, axillary, bracts. 
acuminate as long as cal.-tube; cal. widely campanulate,. 
slightly hairy, segm. all acuminate; pod 24in. by + in. wide, 
much flattened, somewhat curved, with a long beak, glabrous; 


* The Egyptian name. 


Dolichos.| Leguntinose. 7] 


seeds 5 or 6, small, narrowly oblong-oval, much compressed, 
brown. : 

Low country to 2000 ft.; in rocky places, rather common. FI. Dec.- 
March; pale yellow. 

Probably general in Eastern Tropics. _ 

This is never cultivated in Ceylon. Whether it be rightly referred to 
D. unifiorus is doubtful; I follow Thwaites. It is certainly D. axillaris, 
E. Mey., of Mauritius and Trop. Africa. 


D. bifiorus, L., is an annual cultivated form of D. unzflorus, or an 
allied species, erect and very hairy. It is largely grown in S. India, and 
the seed is well known in Ceylon as ‘ Kollu’ or ‘Madras Gram.’ It is 
also cultivated here in a few places in the dry region, but to a very slight 
extent. Moon gives it as ‘cult.’ (Cat. 53) under the name of Glycine 
willosa, There seems to be no published figure of this well-known pulse. 


3. D. ciliatus, Klein in Willd. Sp. Pi. iti. 1049 (1800). 

Thw. Enum. 90. C. P. 1460. 

BE. Ind: 11.°210. 

A perennial twining herb with a tuberous root, stems 
slender, glabrous or nearly so, buds silky; 1.-rachis 1-14 in, 
tumid at base, hairy, stip. ovate, striate, rigid, persistent, Ifits. 
1-24 in., oblong-oval, rounded at base, obtuse, apiculate ; 
glabrous on both sides when mature, densely ciliate with 
white hairs on margin, veins conspicuous beneath; fl. on very 
short ped., 3-6 together in extremely short cymose racemes, 
bracts oval, striate; cal. pubescent, lower segm. lanceolate, 
setaceous; pod about 14in. by }in. wide, much compressed, 
slightly curved, ventral margin much curved, suddenly 
narrowed to curved slender beak, glabrous, slightly veiny; 
seeds 2-4, 4 in., oval, compressed, dark brown. 

ag region; rare. Elephant Pass (Gardner); Jaffna. Fl. Jan.; pale 
jfelLiow. 
: Also in Southern India. 


D. falcatus, Klein in Willd. Sp. Pl. i. 1047 (1800). Wal- 
dambala, 5S. 

Thw. Enum. go. C. P. 1472. 

F]. B. Ind. ii. 211. 


A small twining perennial, roots tuberous, stems very slender, 
slightly hairy; 1. numerous, rachis 1-2 in., slender; stip. 
narrowly triangular, acute, lfits. very variable in size, rhomboid- 
ovate, acute, usually more or less 3-lobed with the lat. lobes 
obtuse, glabrous on both sides; fl. #in., ped. as long as cal., 
1-3 on slender axillary peduncles shorter than 1.; cal. nearly 
glabrous, lower segm. short-triangular, acute; pod 2-24 in. by 
+ in. wide, distinctly curved, with a very short point, glabrous; 
seeds 5 or 6, } in., oval-reniform, compressed, dark brown. 


78 Leguminosae. [Atylosia. 


Low country to 2000 ft.; especially in the dry region. Batticaloa; 
Hantane; Peradeniya; Jaffna. FI. Jan., Feb.; bright light crimson-pink 
(not ‘ yellow,’ as stated in FI. B. Ind.). 

Also in India. 


37. ATVLOSIA, W. and A. 


Erect bush or twining herbs, |. 3-foliolate, lflts. without 
stipels, dotted beneath, fl. in axillary racemes; cal. with upper 
segm. completely connate, lower ones nearly equal or lowest 
longest; pet. persistent, keel not beaked ; stam. diadelphous ; 
pod compressed, straight, few-seeded, strongly compressed 
between seeds by transverse lines; seed with a small white 
2-lipped aril round hilum.—Sp. 20; 16in FV. B. Ind. 

IBKeCEn ae : ; : ; ; . 1. A, CANDOLLEI. 
Twining 
Cal.-segm. triangular . 
Cal.-segm. linear acuminate 


Pod velvety ‘ 
Pod with long soft hair . 


A. ALBICANS. 


N 


A, RUGOSA. 
A. SCARABAOIDES. 


Simice 


1. A. Gandollei, W. and A. Prod. 257 (1834). Et-tora, S. 

abla, lamina, Or, (Co JP, BS: 

1B) 18), Harel mh, Bie \Woaerotm, GS (ts 9/8. 

An erect much-branched bush, 2-4 ft., branches stout 
Lionel, densely clothed with silky hair; 1. numerous, rachis 

—2in., densely silky, stip. very small, caducous, Ifits. very 
a stalked, 1-24 in., linear- lanceolate, acute, finely pu- 
bescent above, densely felted with white or yellow _ silky 
tomentum beneath, thick with reticulate veins very prominent 
beneath, terminal one the largest; fl. rather large, over ? in., 
ped. silky, 4 in. long, solitary or 2 articulated on top of erect 
short axillary peduncle, and enclosed in bud by two large 
rounded, silky bracts which fall before fl. open; cal. very silky, 
segm. linear-lanceolate, lowest longest ; pod 14-14 in., by 3 in. 
wide, straight, with a short deflexed beak, 3—5-seeded, covered 
with dense spreading velvety golden-brown pubescence; seeds 
slightly compressed, black. 

Montane zone on the patanas 3-6000 ft.; very common. FI. all the 
year; bright gamboge yellow, standard red on back. 

Also on the Nilgiris. 


An ornamental shrub, occurring (like broom or furze) gregariously on 
the open patanas. 


2. A. albicans, Leth. Plant. J/ungh. 243 (1853). 
Thw. Enum. 91. C. P. 2783. 
IIE 185 Ikeh th BLS. 


A tylosia.] Leguminose. 79 


A large twining perennial herb, stems long, much-branched, 
angular, finely pubescent; 1. on the main stems much larger 
than on the lat. flowering branches, rachis (of the former) 
3-1} in., divaricate, puberulous, stip. inconspicuous, Iflts. #-14 
in., obovate-rotundate, obtuse, glabrous above, silvery-white 
with minute pubescence beneath, venation not prominent; fl. 
copious, }in. on rather long slender ped., 4-8 in short 
corymbose racemes rather exceeding small 1. of the lat. 
branchlets, often 2 from an axil, bracts roundish, soon falling ; 
cal. puberulous, lower segm. equal, triangular, acute; pet. much 
exserted; pod (not seen) “1-14 in., oblong-linear, beaked, 5-6- 
seeded.” 


Low country to about 2000 ft.; rare. Hantane; about Bibile. FI. 
Jan.; yellow. 
Also in S. India. 


3. A. rugosa, W. and A. Prod. 257 (1834). Wal-kollu, S. 

PawesP nam. O11) CP. TAT: 

FI]. B. Ind. ii. 215. 

A straggling semi-twining perennial herb, stems furrowed, 
finely woolly; |. variable in size, rachis (of the larger ones on 
the main stems) 13-2 in. woolly, stip. ovate, obtuse, incon- 
spicuous, Ifits. 1}-14, obovate-rotundate, usually very obtuse, 
softly velvety above, densely tomentose-velvety and white 
beneath, with prominent reticulate venation, rather thick ; fl. 
about 4 in., ped. pubescent, shorter than cal., 1-3 together at 
end of axillary peduncles about equalling 1., bracts small ; cal. 
densely pubescent, segm. lanceolate-linear, acuminate; pet. 
not much exserted ; pod 1 in. by 4 in wide, with a very short 
point, densely velvety-pubescent ; 3—4-seeded. 

Montane zone, on dry patana land, 3-6000ft.; very common. FI. 
Sept.; yellow. 

Also in S. India. 


4. A. scarabeoides, Benth. Pl. Jungh. iii. 243 (1854). Wal- 
kollu, 5S. 


Herm. Mus. 6. _ FI. Zeyl. n. 282. Dolichos scarabeotdes, L. Sp. PI. 
726. Moon Cat. 53. Thw. Enum. g1. C.P. 1442. 

Fl. B. Ind. ii. 215. Pluk. Phytogr., t. 52, f. 3. 

Annual (?) with a suberect stem and numerous slender, 
cylindrical, twining pubescent branches; |. small, rachis 4-1 in., 
pubescent, stip. very small, Iflts. 4-1} in., oblong-lanceolate or 
oblong-oval, obtuse, finely velvety above, softly pubescent and 
grey beneath, with rather prominent venation, 3-nerved at 
base, thick; fl. small, 4in., ped. shorter than cal., 2 or 3 
together on very short axillary peduncles ; cal. densely hairy, 


80 Leguminosae. [Dunbaria. 


segm. linear-acuminate ; pet. little exserted; pod 3-?in., by 
4in, wide, apiculate, covered with long soft brownish hair, 
4—5-seeded, very strongly lineate between the seeds. 

Low country in both dry and moist regions; rather common. FI. 
December ; yellow. 

Throughout India, Malaya, China, and Mascarene Is. 

Hermann gives the Sinhalese names ‘ Weel-undu’ and ‘ Weel-undu- 
wel’ for this. 


Cajanus indicus, Spreng. is commonly cultivated in gardens. It is 
‘Rata-tora,’ S., ‘ Thavarai,’ T., and ‘Chick-Pea’ of the English ; and is 
mn, 279 of El, Zeyl. feured in Burm. Dhes. t. 37. It has@thes@ars 
number 720, but is nowhere wild in Ceylon. 


38. DUNBARIA,* VW. and A. 


Woody twiners, |. pinnately 3-foliolate, lfits. with stipels, 
conspicuously gland-dotted, fl. in axillary panicles; cal. with 
lowest seem. much longest; pet. persistent; stam. diadelphous ; 
style beardless, stigma capitate; pod linear, more or less 
compressed, few-seeded, marked with lines between seeds ; 
seed with a ring-like small aril—Sp. 10; 7 in FZ B. Ind. 


Lfits. silky beneath, stip. caducous . 6 ; . I, D, FERRUGINEA. 
Lfits. nearly glabrous beneath ; stip. persistent . 2. D. HEYNEI. 


1. D. ferruginea, W. and A. Prod. 258 (1834). 

C, 12s Boo 

TEL 185 Wine, ii, U7. 

A woody twiner, with stout, cylindrical, hairy branches ; 
l. rather large, rachis 14-2 in., hairy, stip. small, lflts. 2-3 in., 
ovate-lanceolate,very acute, glabrous above when mature, finely 
hairy beneath, the lat. ones dilated in lower half; fl. $in., ped. 
slender, +in., hairy, numerous, in erect axillary panicles longer 
than 1., bracts large, ovate, caducous; cal. covered with 
ferrugineous hair, lowest segm. as long as tube; pod over 
Ii in. by 3 wide, straight, witha short deflexed beak, covered 
with long, silky, yellowish hair; seeds 4, nearly circular, 
compressed, black. 

Dry region; apparently very rare. Dambulla, 1868 (Thwaites) ; 


near Bibile, Uva. Fl. Jan., Feb. 
Also in S. India. 


2. D. Heynei, W. and A. Prod. 258 (1834). 
D. oblonga, Arn. Pug. 15. Thw. Enum. go. C.P. 1478. 
Bie Bard ei. 208: 


* Dedicated to Prof. George Dunbar, of Edinburgh. 


Rihynchosia.] Leguminose. SI 


A woody twiner, with stout much-branched stems, more 
or less covered with spreading hair, petiole 4-14 in., hairy, 
stip. lanceolate, acuminate, persistent, lflts. 14-3 in., rhomboid- 
oval, acuminate, acute, glabrous or nearly so above, more or 
less hairy on veins beneath, thin; fl. 8 in. or more, ped. slender, 
hairy, longer than cal., arranged in stalked axillary panicles 
exceeding |., bracts small, ovate, caducous ; cal. roughly pube- 
rulous, lowest segm. very much longest; pod about 2 in. by 
2 wide, nearly straight, with a rather long beak, turgid, covered 
with long weak spreading glandular hair, with faintly marked 
lines between the seeds; seed 4 or 5, ovoid-reniform, mottled 
black and brown. 


Low country; rather common up to 2000 ft. Fl. Jan.; purple. 
Also in S. India. 


39. ERIOSEMA, DC. 


Small erect shrub, |. 1-or 3-foliolate, fl. axillary; cal. cam- 
panulate, segm. equalling tube; keel-pet. shorter than wings; 
stam. diadelphous; style not bearded, stigma capitate; pod 
oblong, turgid; seeds 1 or 2, oblique with linear hilum.— 
sp 50; iin FZ B. [nd. 


E. chinense, Vogel in Meyen Obs. Bot. 31 (1843). 
Fl. B. Ind. ii. 219. 


A small erect slightly branched shrub with a tuberous root; 
lfits. 1-2 in., linear-ligulate, with a few adpressed hairs above, 
glaucous and densely pubescent beneath, stip. linear, per- 
sistent; fl. 1 or 2, on short axillary ped.; cal. densely pilose; 
pet. twice length of cal., pod 2in., oblong, densely pubescent. 

Inserted on the authority of specimens in Hb. Kew from Col. Walker, 
but without special locality. 

Also in E. Bengal, Burma, Malacca, China, Philippines, and Trop. 
Australia, but not in Peninsular India. 

I have never met with this, which should be searched for. The 
description is taken from the Fl. B. Ind. Walker’s specimens are very 
dwarf, and look as if grown in dry patana land. It is possible they may 
have come from E. Bengal. 


40. RHYNCHOSIA, Lour. 


Twining, trailing, or rarely erect, 1. pinnately 3-foliolate, 
lfits. usually with stipels, gland-dotted beneath, fl. in axillary 
racemes ; cal. with upper segm. more or less connate; keel- 
pet. not beaked ; stam. diadelphous ; pod short, flattened or 

PART I. G 


82 Leguminosae. [Rhynchosia. 


turgid ; seeds I or 2, with or without a small white aril round. 
oblong hilum.—Sp. 80; 22 in FV. B. Lnd. 


Seeds with a distinct aril. 
Cal.-segm. oblong, obtuse . 5 ; é . I. R. RUFESCENS. 
Cal.-segm. acuminate or setaceous. 
Pod circular, 1-seeded ; annual (Vomzsmia) . 2. R. NUMMULARIA.. 
Pod 2-seeded ; shrubs. 


Pod with a distinct partition between seeds 3. R. SUAVEOLENS. 
Pod without a partition . 5 : . 4. R. CANA. 

Seeds without an aril. : 
Cal.-segm. oblong, obtuse (Cyanosfermum) . 5. R. CYANOSPERMA.. 
Cal.-segm. acute. 

Racemes elongated. 
Lfits. obtuse : : : : : . 6. R. MINIMA. 
Lfits. acute. 
Ped. very short 7. R. VISCOSA. 
Ped. longer than cal. 5 . 8. R. ACUTISSIMA. 
Racemes capitulate > ; A : . 9. R. DENSIFLORA. 


I. R. rufescens, DC. Prod. ii. 387 (1825). 

Lespedeza (?) indica, Spr., Arn. Pug. 14. Thw. Enum. 91. C.P. 1424. 

HSB s indir 220) eblookea Ke Pin treo! 

An erect, annual (?) herb, semi-woody below, with long, 
slightly twining, densely pubescent branches; |.-rachis I-14 in., 
pubescent, stip. minute, caducous, Ifits. #-14 in., lanceolate or 
oval, acute, finely silky on both sides, with raised reticulate 
venation beneath; fl. rather small, shortly pedicellate, 1-4, 
distant in very slender elongated racemes, bracts minute ;. 
cal. finely pubescent, segm. very much longer than tube, 
oblong, obtuse, imbricate, somewhat enlarged in fruit; pet. 
rather shorter than cal.; pod }-3in., roundish with a short 
beak, pubescent; seed solitary, black, shining, with a succulent. 
2-lipped aril. _ 

Dry region; rather rare. Batticaloa District, common (Thwaites).. 


Fl. March; yellow. 
Also in India and Java. 


2. R.nummularia, DC. Prod. ii. 386 (1825). 

Nomismia nummularia, W. and A.,Thw. Enum. 91. C. P. 1439. 

Fl. B. Ind. ii. 221. Wight, Ic. t. 283. 

Annual, stems long, prostrate, trailing, hairy; 1. variable 
in size, rachis 1-3 in., pubescent, stip. small, ovate, acuminate, 
Iflts. 1-2 in. wide, shorter than broad, truncate or very obtuse,, 
thick, finely pilose on both sides, the lat. ones much dilated 
in lower half; fl. small, on short ped., 4-8 in distant pairs, 
racemes stalked, usually shorter than 1.; cal. hairy, segm. 
lanceolate, acuminate ; pod circular, 4 in. diam., tipped with a. 
sharp point, strongly wrinkled with veins on both sides, 


Rhynchosia.] Leguminosae. 8 
clothed with a few long hairs; seed solitary, }in., with a 
rather large 2-lipped aril. 

Low country; rare. Jaffna (Gardner); Colombo (Ferguson). 

Also in S. India. 


R. aurea, DC., is given for Ceylon in Fl. B. Ind. on the faith of a 
specimen in Hb. Kew, labelled by Bentham ‘ Ceylon, Walker.’ It is not 
unlikely to occur, but has not been noticed by any one else. Figured (as 
Nomismia capitata) in Wight, Ic. t. 295. 


3. R. suaveolens, DC. Prod. ii. 387 (1825). 

Thw. Enum. 441. C. P. 3851. 

Fl. B. Ind. 11. 221. 

An erect undershrub, stems covered with a fine viscous 
pubescence, branches numerous, spreading, slender, flexuose ; 
l.-rachis 1-1} in., slender, pubescent, stip. linear, persistent, 
lfits. 15-2} in., lanceolate or oval, acuminate, acute, very thin, 
finely pubescent on both sides; fl. small, on short ped., soli- 
tary or 2 at end of stiff slender peduncles shorter than 1.; cal. 
hairy, segm. twice as long as tube, linear, setaceous ; pod 2 in. 
by 4 in. wide, with a very short decurved point, finely pubes- 
cent; seeds 2 with a distinct external line between them, 
small, with well-marked aril. 

Dry region; very rare (?). Trincomalie (Glenie). 

Also in Peninsular India. 


4. R. cana, DC. Prod. ii. 386 (1825). 

Flemingia biflora, Moon Cat. 53. Thw. Enum. 91. C. P. 1280. 

Fl. B. Ind. i. 222. 

A small erect shrub, with very numerous ascending stems 
with short slender lateral branches, all covered with short 
dense spreading hair; 1. numerous, much larger on the stems 
than on lat. branches, rachis (of former) I in., hairy, stip. very 
small, acicular, lflts. 1-2 in., oval, acute (on lat. branches much 
smaller, lanceolate or linear-lanceolate), slightly pubescent 
above, softly velvety - pubescent, paler and with prominent 
reticulate venation beneath; fl. small, on short slender nod- 
ding ped., 2 together at end of peduncles shorter than 1.; cal. 
hairy, segm. linear, setaceous, longer than tube; pod about 
4 in. by + in. wide, with a minute apiculus, turgid, very finely 
pubescent ; seeds 2 without any partition between them, 
ovoid, aril small. 

Low country; rather common, growing gregariously. Near Kalu- 
tara; Badulla; Nilgala and Lunugala; Jaffna District. Fl. Dec.-March; 
yellow. 

Also in Peninsular India. 

Moon gives the S. name ‘ Gas-kollu’ for this. 


84 Leguminos. ae. [Rhynchosia. 


5. R. cyanosperma, Lenth. in Fl. Trop. Africa, ii. 218 (1871). 
Cylista tomentosa, Roxb., Moon Cat. 53. Cyanospermum tomentosum 
W. and A., Thw. Enum. 92. C. P.2471 


Fl, B. Ind. ii. 222. Wight, Ill. t. 84 (Cyanospermum). 


A stout woody twiner, with thick, very pubescent branches; 
1. large, rachis 14-4 in., softly rufous-hairy, stip. inconspicuous, 
caducous, lflts. 2-34 in., broadly ovate, shortly acuminate, 
finely velvety on both sides, venation conspicuous beneath, 
stipels large, setaceous, hairy ; fl. rather large, # in., on ped. 
shorter than cal., numerous, in rather lax softly pubescent, 
erect or ascending, stout, stalked racemes longer than 1, 
bracts very large, ovate, acute, velvety, quickly falling; cal. 
large, densely covered with long soft silvery hair, segm. longer 
than tube, oblong, very obtuse; pet. scarcely exceeding cal., 
persistent ; pod 4 in. by about + wide, slightly exceeding cal., 
with a short curved point, densely velvety with pale green 
hair; seeds 2, with a constriction between them, broadly 
ovoid or subglobular, very slightly compressed, shining dark 
violet-blue, without an aril, remaining attached to the valves 
of pod after dehiscence. 


Upper zone of moist and intermediate low country; rather rare. 
Ambagamuwa; Allagala; Hanguranketa; Etampitiya. Moon’s locality 
is Four Korales. Fl. Jan., Feb.; pale yellowish-white. 

Also in Peninsular India, E. Trop. Africa and the Mascarene Is. 

Remarkable for the deep metallic blue of the seeds. Moon gives 
‘ Hin-garadiya’ as the S. name. 


6. R. minima, DC. Prod. ii. 385 (1825). 


Burm. Thes. 188.  Dolichos medicagineus, Lam., Moon Gale 53- 
Thw. Enum. gt. C. P. 1440. 


Fl. B. Ind. 1. 223. Burm. Thes. t. 84, fi 2. 


Annual, stems numerous prostrate, trailing and slightly 
twining, sien much branched, slightly GUIDES EE 1, small, 
rachis 4—? in., stip. lanceolate, setaceous, lfits. 3-2 in. , broadly 
oval or rhomboid - oval, obtuse, apiculate, glabrous above, 
pubescent on veins beneath and conspicuously black-dotted, 
terminal one often broader than long; fl. very small, 4 in., on 
very short ped., 3-6, racemes lax, slender, exceeding the 1; 
cal. pubescent, segm. longer than tube, lanceolate, acuminate ; 
pod rather over 4in. by 4 in. wide, somewhat compressed 
but slightly turgid, a little curved upwards, finely puberulous, 
seeds 2, compressed without an aril. 

Low country; common in the dry region, less so in the moist. FI. all 
the year; yellow. 

Found throughout the Tropics and at the Cape and in United States. 


Moon calls this * Maha-wal-kollu.’ Burman’s description is from speci- 
mens in Hartog’s collections. 


Rhynchosia.] Leguminose. 85 


7. R. viscosa, DC. Prod. ii. p. 387. 

flemingia viscosa, Moon Cat. 54(?). &. véllosula, Thw. Enum, 412. 
C. P. 3790, 3895. 

Fl. B. Ind. ii. 225. 

Perennial, semi-woody at base, stems stout, twining, closely 
set with short glandular hairs ; |. rather large, rachis 2-23 in., 
viscous-hairy, stip. narrowly triangular, acute, Iflts. 14-21 in., 
broadly trapezoid-ovate, truncate or subcordate at base, acute, 
viscous-hairy on both surfaces, thin; fl. 4 in., on very short 
ped., numerous, racemes close or lax, stalked, exceeding l., 
bracts ovate acuminate, soon falling; cal. viscous-hairy, segm. 
acute, upper ones connate for less than half way, lowest one 
much the longest; pod about 1} in. by 2 wide, narrowed at 
base, tapering to long point, somewhat compressed, finely 
puberulous, when young covered with long yellowish hair 
afterwards deciduous; seeds 2, nearly circular, } in. diam., 
with a very small aril. 

Low country; rare (?). Trincomalie (Glenie); Colombo (Ferguson). 
Fl, Jan.—March; pale yellow. 

Also in India, Malaya, Trop. Africa, and Mascarene Is. 


8. R. acutissima, 7iw. Enum. 413 (1864). 
Thw. Enum. 413. C. P. 3445. 

Fl. B. Ind. ii. 226. 

Perennial, branches stout, twining, covered (as in whole 
plant) with short, soft, fulvous hair ; 1. large, rachis 14-23 in., 
stip. lanceolate, deciduous, Iflts. 24-5 in. trapezoid-ovate, 
tapering to very long acuminate acute apex, fulvous-pubescent 
on both sides; fl. 4 in., ped. in pairs, rather longer than cal., 
racemes equal to or shorter than 1., bracts lanceolate, acute, 
soon falling ; cal. pubescent, upper segm. connate nearly to 
end, lowest conspicuously longest ; pod not seen. 

Moist region; very rare. I have only seen the C. P. specimens which 
were collected at 3000 ft. on Hantane in Dec. 1859. FI. pale yellow, with 


red lines. 
Also in Travancore. 


g. R. densiflora, DC. Prod. ii. 386 (1825). 
Trimen in Journ. Bot. xxiii. 144. 
Fl. B. Ind. ii. 226 (not given for Ceylon). 


Perennial, stems slender, twining, clothed with deflexed 
silvery hair; l.-rachis 14-2 in., hairy, stip. small triangular, 
acute, Iflts. 14-2 in., rhomboid-ovate, acute, finely hairy on 
both sides, thin, conspicuously black-dotted ; fl. 2 in., on short 
hairy ped., numerous, crowded in a dense, ovoid, nearly sessile, 
capitulate raceme, bracts linear, acuminate, hairy, persistent, 


86 Leguminosae. [Flemingia. 


dotted with red glands, tube very short, segm. linear attenuate, 
two upper connate for less than half way; pod ( immature) 
nearly 2 in., tapering to point, clothed with long hair, gland- 
dotted, 2-seeded. 


Dry region; very rare. As yet, only found at Tissamaharama near 
Hambantota. 

Fl. December. 

Also in Peninsular India and in W. Trop. Africa. 


41. FLEMINGIA,* 7ox2. 


Erect shrubs, |. 1- or palmately 3-foliolate, fl. small; cal. 
with a short tube and long narrow segm., 2 upper not connate, 
lowest one the longest; pet. equal, keel obtuse or slightly 
beaked ; stam. diadelphous, style beardless, stigma capitate; 
pod small, inflated, usually 2-seeded ; seed without an aril,— 
Sp 4 set lpinty bea. 


L. unifoliolate. 

L. 3-foliolate 
Fl. in spreading panicles. 
Fl. in dense racemes 


. F. STROBILIFERA. 


Ll 


. |, LINEATA: 
. F. CONGESTA. 


1. FE. strobilifera, Br. in Azt. Hort. Kew., ed. 2, iv. 350 (1812). 
Hampinna, S. 

Herm. Mus. 53. FI. Zeyl.n. 289. Hedysarum strobiliferum, L. Sp. Pl. 
746. Moon Cat. 54. Thw. Enum. 92. C. P. 697. 

Fl. B. Ind. ii. 227. Fl. Zeyl. t. 3. Wight, Ic. t. 267. 

A shrub, 4-6 ft. high, bark smooth, grey, branches densely 
pubescent with adpressed hair; 1. 1-foliolate, petiole 4 in, 
hairy, stip. small, caducous, lflts. 9-5 in., ovate, often cordate 
at base, acuminate, subacute, glabrous above, finely pubescent 
and paler beneath, with lat. veins parallel and very prominent; 
fl. small, on very short ped., 2-4 together in axils of and 
completely concealed by very large conduplicate bracts which 
are roundish, rather broader than long, #-1 in. by 1-1} in., 
deeply cordate at base, obscurely cuspidate, membranous, 
reticulately veined, finely pubescent outside, arranged dis- 
tichously and much overlapping on either side of a slender 
zigzag hairy rachis (but all turned upwards) at the end of 
short lateral branches; cal. pubescent; pod 4in. by }in. wide, 
apiculate, densely downy, completely concealed in the per- 
sistent hooded bracts; seeds 2 or 1, small, mottled. 


Low country; rather common and locally abundant. FI. July, August; 
white or purple. 


Go N 


* Commemorates Dr. John Fleming, F.R.S., ‘ Physician-General in 
Bengal.’ Died 1815. 


Flemingia.) Lecuminose. 87 


Also in the Himalayas, E. Bengal, Burma, Malacca, Malay Is., and 
Philippines. 

The heads of persistent membranous pale-brown bracts are very 
suggestive of hop-cones or those of Carf~zmus. The plant has a first-sight 
resemblance to Desmodium pulchellum, and the fl. are arranged in much 
the same way. Intermediate stages between the ordinary foliage leaves 
and the hood-like bracts are often met with at the base of the flowering 
‘branchlets. 


2, F. lineata, Roxb. Hort. Beng. 56 (1814). 

Thw. Enum. 92. C. P. 1423. 

FI. B. Ind. ii. 228. Wight, Ic. t. 327. 

A small erect shrub, young branches with adpressed 
pubescence ; |. 3-foliolate, rachis about 4 in. much furrowed 
above, stip. conspicuous, long-acuminate, scarious, subper- 
sistent, lfits. very shortly stalked, 1$-2 in., lanceolate-oblong, 
narrowed at base, subacute, glabrous above, paler and very 
finely silky especially on the very oblique veins beneath, 
rigid ; fl. small, on very short ped., in small clusters, arranged 
in pedunculate erect spreading axillary panicles shorter than 
leaves, cal. puberulous, segm. linear-lanceolate; pod 3 in., 
oblong-ovoid, compressed, blunt, minutely apiculate, sparingly 
pubescent. 

Dry region; rare. Batticaloa (Thwaites); N. Prov. (Gardner). Fl. 


Feb. 
Also in India, Malaya, and N. Australia. 


3. F. congesta, Roxb. Hort. Beng. 56 (1814). 

Thw. Enum. 92. C. P. 2663. 

Fl. B. Ind. ii 228. Wight, Ic. t. 390. 

An erect shrub, 3 or 4 ft, young branches angular, 
adpressed-hairy, 1. 3-foliolate, large, rachis 3-4 in., sulcate 
above, stip. caducous, Iflts. on short densely hairy stalks, 5-7 in., 
narrowly rhomboid-lanceolate, rounded at base, tapering to 
long acute apex, glabrous above, sparingly adpressed-hairy 
beneath ; fl. on very short ped., crowded in dense, oblong, 
sessile racemes often 2 or 3 from same axil, rachis silky, bracts 
small, lanceolate, acuminate, silky, soon falling; cal. densely 
adpressed-hairy, segm. linear-setaceous; pod 3 in., oblong- 
ovoid, blunt, inflated, apiculate, very finely pubescent. 

Var. 6, semialata, Paker. F. semialata, Roxb., Moon Cat. 54; 
Thw. Enum. 92. C. P. 256, 3584. Wight, Ic. t. 326. 

Rachis margined with a narrow leafy wing; racemes 
elongated. 

Low country; rather common. The type in Ambagamuwa, near the 
Maskeliya River. Var. 8, Colombo; Badulla; Batticaloa, Fl. Feb, 
March; pale pink. 


88 Leguminose. [Dalbergia 


Also in India, Malaya, Philippines, China. 

Moon gives ‘ Wal-undu’ as the S. name. C. P. 3584, which has the: 
leaves narrower, obtuse and densely pubescent beneath, is probably the 
variety W7zghtiana of F1. B. Ind. 


42. DALBERGIA, L. 7. 


Trees or climbing shrubs, |. pinnate, with alternate lfits.,. 
fl. small, in axillary panicles; cal.-tube campanulate, segm. 
short, the upper 2 not connate; keel-pet. connate at top only;. 
stam. 9, monadelphous, or 10 in two bundles of 5 ; style short,. 
stigma capitate; pod indehiscent, much compressed, usually 
thin, 1-4-seeded.—Sp. 60-70; 28 in FZ. B, Ind. 

Stam. 9, monadelphous; climber . ; . . I. D. CHAMPIONII. 
Stam. 10, in 2 bundles of 5 each 


Tree, pod straight . F - 3 ; . 2. D.LANCEOLARIA. 
Climber, pod falcate : 5 : : . 3. D. MONOSPERMA.. 


I. D. ppemlont? Thw. Enum. 94 (1859). Bambara-wel, S. 
GaP Or 
Fl. B. Ind. ii. 232. 


A scandent shrub, climbing by means of hooked lateral 
twigs, bark light brown, young parts clothed with very fine: 
yellow pubescence ; |. rather large, rachis 14-3 in., cylindrical, 
glabrous, stip. small, oblong, obtuse, falling before 1. expand, 
lflts. I-4, on stout wrinkled stalks, oval, caudate-acuminate, 
quite glabrous on both sides; fi. rather small, 2 in., on slender 
ped. nearly as long as cal., panicles shorter than 1.; cal. very 
finely pubescent, upper segm. very short, broad, obtuse, lower 
acute, pet. with very long claws; stam. 9, monadelphous; pod 
distinctly stalked, 24-22in., by 3 in. wide, straight, very flat,. 
obtuse, finely pubescent, with reticulate venation over seed 3. 
seed solitary, 1fin., oblong, much flattened. 

Moist region, 2000-4000 ft.; rather rare. Deltota; Hantane; Dolos- 
bagie; Ella. Fl. April, May; white. 

Found also in Borneo. 

If D. pseudo-Sissoo, Miq., of Java, should prove to be the same plant,. 
that name has priority over Thwaites’. 


_D. latifolia, Roxb., has occurred in one or two places, but I do not 
think it wild. Trees have been grown for very many years in the Botanic 
Gardens, and seedlings distributed. It is figured in Wight, Ic. t. 1156, 
and Bedd. Fl. Sylv. t. 24. 


2. D. lanceolaria, L. f/f. Suppl. Pl. 316 (1781). Weluruvai, 7. 

D. zeylanica, Roxb. Hort. Beng. 53. D. frondosa, Roxb., Thw. Enum- 
94; Trim. Syst. Cat. 27. C. P. 1496. 

Fl. B. Ind. ii. 235. Wight, Ic. t. 266. 


Dalbergia.) Leguminosae. 89 


A large tree, bark smooth, greyish-white, young shoots 
glabrous ; |. rather large, rachis 5-7 in., slender, stip. minute, 
soon falling, Iflts. 10-15, on stalks }-+in. long, 1-14 in, 
obovate-oval, acute at base, emarginate, glabrous above, 
whitish or glaucous beneath with a very fine mealy puberu- 
lence, lat. veins very oblique, numerous, parallel, conspicuous ; 
fi. small, 2 in., panicles short, very numerous from axils of 
young l., ped. shorter than cal., cal. hairy, segm. lanceolate, 
subacute, the lowest conspicuously longest; stam. in two 
bundles of 5 each; pod 24in. by } in. wide, straight, long- 
stalked, tapering to acute apex and gradually to base, thin and 
flexible, nearly glabrous, reticulate-veined over seeds; seeds 
1-4, +—2 in., oval, compressed. 

Low country, chiefly in the dry region; rather rare. Jaffna; Elephant 
Pass; Vavuniya; Mulliativu; Trincomalie; Deltota. Fl. May-—Aug.; 
pale violet, cal. brownish-purple. 

Also throughout Peninsular India. 

A beautiful tree when profusely covered with flowers and young leaves. 

Koenig’s full description of the Ceylon plant in ‘Supp. Pl.’ does not 
quite agree with ours, but his Indian specimens (from Tranquebar) in 
Mus. Brit. are certainly the same species. The specific name applies to 
the shape of the pod. 

D. zeylanica, Roxb., sent to the Calcutta Bot. Garden by Gen. Hay 
Macdowall in 1801 under the Sinhalese name ‘Belloo-labba,’ is very 
briefly described in F1. B. Ind. iii. 228, but is probably this. 


D. volubilis, Roxb., is recorded from Ceylon in FI. B. Ind. ii. 235, on 
the faith of specimens in Hb. Kew, collected by Mrs. Walker, but there is 
no other record. Should be looked for; figured in Roxb. Pl. Cor. t. 191. 


3. D. monosperma, Dalz. in Kew Journ. Bot. ii. 36 (1850). 

Thw. Enum. 94. C. P. 243. 

Fl. B. Ind. ii. 237. 

A stout scandent shrub, scrambling by means of its 
divaricate lateral twigs often hooked at the ends, bark 
purplish-black, smooth; 1. closely placed, rachis 14-24 in., 
cylindrical, nearly glabrous, Iflts. 4-6, shortly stalked, 14-14 in., 
oval or obovate-oval, obtuse, often emarginate, generally acute 
at base, glabrous above, paler and very finely pubescent 
beneath ; fl. small, under } in., ped shorter than cal., panicles 
very short, under 1 in., crowded, bracts beneath cal. oval, 
obtuse; cal. nearly glabrous, segm. short, obtuse; stam. in two 
bundles of 5; pod tin. by 3 in. wide, very shortly stalked, 
falcately curved, acute, glabrous, 1-seeded. 

Near the seacoast, especially in Mangrove swamps, scrambling over 
bushes; rather rare. Panadure; Trincomalie; Koddiyar. Fl. Aug.- 
Nov.; dull pink. 
pee on shores of India, Malaya, Philippines, N. Australia, and 

ina, 


90 Leguminose. [Pongamia. 


43. PTEROCARPUS, J. 


Tree, 1]. pinnate with alternate lfits., without stip., fl. 
moderate-sized in terminal panicles; cal. urceolate, curved, 
with very short equal segm.; pet. with long claws, margins 
crisped, keel-pet. very slightly coherent ; stam. 10, monadel- 
phous, fil. unequal; ov. with 2 ovules, stigma terminal, pod 
indehiscent, stalked, flat, broadly winged, 1-seeded.—Sp. 15 ; 
Ain FL. B. L[nd. 


P. Marsupium, orb. Cor. Pl. ii. 9 (1798). Gammalu, S. 
WVenkai, 7. 

P. bilobus, Roxb., Moon Cat. 52. Thw. Enum. 92. C. P. 1495. 

Fl. B. Ind. ii. 239. Bedd. Fl. Sylv. t. 21. Bentl. and Trim. Med. 
Pl. t. 81 (from a Ceylon specimen). 


A tree with a stout crooked stem and wide-spreading 
branches, bark thick, yellowish-grey, young parts glabrous; 1. 
large, rachis 3-4 1in., cylindrical, thickened at base, glabrous, 
Iflts. 4-6, alternate, on +4 in. stalks, 24-34 in., acute at base, 
more or less bilobed at apex, glabrous on both sides, shining 
above, thick, lat. veins very numerous, parallel and prominent; 
fl. £in., articulated on short ped., panicles 3-6 in., rather lax, 
bracts beneath cal. small, deciduous; cal. finely velvety-pilose, 
segm. broadly triangular; pod indehiscent, nearly circular, 
about 1in. diam., very much curved upwards, the small beaked 
apex being almost in contact with the base, surrounded by a 
very broad, rigid undulate veined wing, glabrous; seed small, 
kidney-shaped. 


Low country, chiefly in the dry and intermediate regions up to 3oo0oft., 
rather common; in the moist region on open grass land, rarer. Common 
about Navalapitiya. FI. July—Sept.; bright yellow. 

Also in Southern India. 

Good specimens are in Brit. Mus. collected by Keenig in 1781. 

A dark red gum-resin (Kino) exudes from the bark, and is used as a 
medicine and for outward application. Affords a fine timber, very hard, 
heavy, dark reddish-brown, durable, containing a red resin. 


44. PONGAMIA,* Vezt. 


Tree, 1. imparipinnate, fl. in axillary racemes; cal. campa- 
nulate, almost truncate at mouth; pet. much exserted, standard 
very broad, keel-pet. coherent near tip; stam. 10, monadel- 
phous, but the uppermost one free for half way down ; stigma 


* From Pougam, the name given in Rheede, Hort. Malab. for the tree. 


Dervis. Leguminosae. gl 


capitate; pod woody, indehiscent, much compressed, not 
winged, 1-seeded.—Monotypic. 


P. glabra, Vent. Jard. Malm. t. 28 (1803). Magul-karanda, S. 
Punku, 7. 

Dalbergia arborea, Willd., Moon Cat. 51. Thw. Enum. 92. C.P. 1489. 

Fl. B. Ind. ii. 240. Wight, Ic. t. 59. Bedd. Fl. Sylv. t. 177. 

A large tree, bark soft, grey, not cracked or furrowed, buds 
slightly puberulous ; |. large, rachis about 5 in., glabrous, stip. 
small, oblong, obtuse, soon falling, Iflts. 5-9 (2-4 pairs and an 
end one), 3—5 in., on thick stalks, oval or lanceolate, acute at 
base, acuminate, glabrous and shining on both sides, thin, 
bright green; fl. gin., ped. rather long, slender, swollen at 
base, articulated, often in pairs, racemes often 2 together, 
elongated, about equalling 1., cal. minutely puberulous, pod 
about 2 in. by 14 in. wide, scarcely stalked, pointed, oval-oblong, 
glabrous. 


Low country, banks of streams and rivers, especially near the coast; 
common. FI. April; greenish-pink or white, with cal. purplish-brown. 

Throughout Tropical Asia, and in Seychelles. 

Wood moderately hard, yellowish-white, not durable. The seeds 
afford an oil which is used in skin diseases. 


45. DERRIS, Lour. 


Climbing shrubs, |. imparipinnate, lflts. without stipels, f1. 
very numerous, in axillary or terminal panicles or racemes ; 
cal. shallowly campanulate, segm. very short; pet. much 
exserted, keel-pet. slightly connate; stam. monadelphous or 
diadelphous, stigma capitate; pod indehiscent, flat, with a 
narrow wing along dorsal suture, 1-4-seeded.—Sp. 40; 23 in 
FI. B. Ind. 


Stam. monadelphous or imperfectly diadelphous. 
Standard without calli at base ; Iflts. 3-9. 
Lfits. 14-2 in. 


Fls. in clusters, in long elongated panicles 1. D. SCANDENS. 
Fls. 2 or 3 together, in short panicles 2, D. PARVIFLORA. 
Lfits. 2-3 in. 
Fl. in simple racemes ; 3. D. ULIGINOSA. 
FJ. in much-branched panicles ; 4. D. PANICULATA. 
Standard with 2 callosities at base; Iflts. 9-13 5. D. OBLONGA. 
Stam. distinctly diadelphous ; ; ; 6. D. SINUATA. 


1. D. scandens, Penth. in Journ. Linn. Soc. iv., Suppl. 103 (1860). 
Kala-wel, 5. Tekil, 7: 

Thw. Enum. 413 and (Brachyplerum scandens, W. and A.) 93. C. P. 
1492. 

Fl. B. Ind. ii. 240. Wight, Ic. t. 275 (Brachyplerum). 


92 Leguminosae. [Derris:. 


A large far-reaching climber, stems smooth, dark purplish,, 
young parts pubescent; |.-rachis about 3 in., deeply channelled, 
stip. small, soon falling ; lfits. usually 9 (4 pair and end one) 
14-2 in. long, on short stalks, oblong-lanceolate, acute at base, 
obtuse often emarginate at apex, quite glabrous when mature 
on both sides, rather rigid; fl. small, 3 in., very numerous, on 
rather long slender ped., with 2 very small obtuse bracts just 
below cal., arranged in clusters on the very short nodular 
branches of elongated narrow, pendulous, terminal and axillary 
racemose panicles 10-18 in. long; cal. finely pubescent; stam. 
monadelphous ; pod 17-24 in., by about 4 in. wide, oblong- 
lanceolate, tapering to both ends, pointed, straight, very finely 
hairy, 1-3-seeded. 

Dry region; very common. FI. Aug.—Sept.; white. 

Also in India, Burma, Malaya, Trop. Australia, and China. 

Very beautiful in the driest season when little else is in blossom im 


the dry forest country; its great masses of pure white flowers drooping: 
from the tops of the trees. 


2. D. parviflora, Benth. in Journ. Linn. Soc. iv., Suppl. 105 (1860). 
Thw. Enum. 413 and (Brachypterum elegans, Thw.) 93. C. P. 2508. 
F]. B. Ind. ii. 240. 


A climbing shrub, perfectly glabrous throughout; 1.-rachis. 
3-4 in., lfits. usually 7 (3 pair and end one), 14-2 in., acute at 
base, shortly acuminate, quite glabrous, paler beneath and 
when young very finely reticulate; fl. as in D. scandens, but 
only 2 or 3 on each node and with panicles much shorter, 
scarcely as long as 1. and more branched; cal. quite glabrous, 
segm. acute; pod 14-2? in., by scarcely 4 in. wide, narrowly 
oblong, obtuse at both ends, glabrous, 1-3-seeded. 

Low country, principally in the dry region; rather rare. Deltota ; 


Uma-oya; near Tirapane, N.C. Province. Fl. June-August, pale purple. 
Endemic. 
The specific name is unfortunate, as the flowers are as large as in 
D. scandens, but Thwaites’ one is preoccupied by D. elegans, Benth. of 
Burma. The young leaves turn black in drying. 


[D. robusta, Benth. (Dalbergia Krowee, Roxb.) is given for Ceylon in 
Fl. B. Ind., and has been sent from here by many collectors (Gardner, 
Walker, Macrae, &c.). It is a large tree, not a climber, and has long been 
cultivated in the Bot. Gardens, &c., but has no claim to be considered a 
native. Wight, Ic. t. 244.] 


3. D. uliginosa, Benth. Pl. Jungh., 3, 252 (1854). Bala-wel, S. 

Herm. Mus. 21. Fl. Zeyl.n. 417. Thw. Enum. 92. C. P. 1494. 

Fl. B. Ind. ii. 241. Hook. Bot. Misc. iii. t. Suppl. 41 (Pongamia 
triphylla). 

A strong woody climber, bark dark grey, with large, scat- 
tered lenticels ; 1. rather large, rachis 3-4 in., Iflts. 3-7 (1-3 


Dervis. Leguminose. 93 


pair and end one), 14-3 in., oval or ovate-oval, rounded at 
base, obtusely acuminate, quite glabrous, rather thick, minutely 
reticulate ; fl. on short ped. arranged in simple axillary 
racemes often from the old wood and 4-6 in. long; cal. finely 
pilose, truncate with obscure segm., stam. monadelphous or 
diadelphous ; pod rather over 1 in., as broad as long, nearly 
circular, shortly apiculate, glabrous, veined; seed solitary, 
+4 in., kidney-shaped, compressed. 

Low country near the seacoast, chiefly in the dry region ; common. 
Fi. March, August; pink. 


Found also in India, China, Polynesia, Trop. Australia, E. Trop. 
Africa, and Madagascar. 


Though included in his FI. Zeyl., where he referred it to the genus 
Pterocarpus, Linnzeus never subsequently named this species; he has 
confused all subsequent writers by having quoted for it an old figure and 
description of a Trop. American plant, Pzerocarpus Draco. 

Hermann notices the use of this plant as a fish-poison, for which the 
roots are still employed. The leaves turn yellow in drying. Ropes and 
fishing-lines are made of the strong fibre from the stem. 


4. D. paniculata, Benth. in Journ. Linn. Soc. iv., Suppl. 105 (1860). 

Brachypterum Benthamiz, Thw. Enum. 93. D. Benthamiz, Thw. 
Enum. 412. C. P. 2925. 

Fl. B. Ind. ii. 242. 

A woody climber, with slender dark-coloured lenticellate 
branches, young parts with fine fulvous pubescence; 1-rachis 
3-4 in., cylindrical, stiff, lflts. 3-7, 1-3 in., stalked, lanceolate- 
oblong, obtuse, acuminate, twisted at apex, rounded at base, 
glabrous, coriaceous ; fl, numerous, ped. about as long as cal. 
panicles terminal, 4-8 in. long, much-branched, drooping, 
fulvous-pubescent ; cal. pubescent, upper segm. absent, lower 
triangular, obtuse, stam. monadelphous but the uppermost 
partially free; pod 1-1#in., oblong, compressed, apiculate, 
finely fulvous-tomentose, I—2-seeded. 

Dry region ; very rare. Uma-oya (Thwaites). It was also collected 


by Walker, but the locality is not known. FI. July; very pale pink, 
cal. red. 


Endemic. 


5. D. oblonga, Lenth. in Journ. Linn. Soc. iv., Suppl. 112 (1860). 
{PLATE XXIX. ] 


Thw. Enum. 413 and (D. ovalifolia, Benth. var.) 92. C. P. 1493. 

Fl. B. Ind. ii. 242. 

A strong woody climber, bark dark grey, smooth, with 
large distant lenticels, young parts puberulous ; |. large, rachis 
4-6 in., glabrous, Iflts. 9-13 (4-6 pair and end one), stalked, 
2-24 in., lanceolate-oblong, acute at base, obtuse often emar- 
ginate at apex, glabrous, paler beneath, stip. conspicuous with 


94 Leguminosae. [ Sophora. 


young 1. rotundate, adnate to stem, covered with orange 
pubescence; fl. fully 4in, ped. as long as cal., racemes 
numerous, sessile, coming from the base of the shoots below 
the young 1. 3-4 in. long; cal. slightly pilose, upper segm. 
nearly absent, lower ones shallow, broadly triangular; standard 
with 2 callosities at base; stam. monadelphous but upper one 
partly free ; pod 14-2 in. by over ? in. wide, much compressed, 
acutely pointed, glabrous, 1-seeded. 

Moist region, 2000-4000 ft.; rare. Sabaragamuwa; Kitulgala; Han- 
tane. Fl. March; white, tipped with pink, or all pale violet-pink, cal. 
coppery-pink. 

Also in Western India. 


A very ornamental climber; the young leaves, twigs, &c., are of the 
same coppery-pink colour as the calyx. 


6. D. sinuata, Benth. in Thw. Enum. 93 (1859). 

Herm. Mus. 16. Fl. Zeyl. n. 164. Sophora heptaphylla, L. Sp. Pl. 
373 (non Auct. post.). Thw. Enum. 93. C. P. 1491. 

Fl. B. Ind. i. 246. 

A large scandent shrub with pale bark; 1.-rachis 4—7 in. 
smooth, Iflts. 7 (3 pair and end one), on thick petioles + in. 
long, about 3 in., oval or lanceolate, rounded at base, obtuse 
and emarginate at apex, somewhat conduplicate, perfectly 
glabrous, coriaceous; fl. $ in., articulated on hairy ped. shorter 
than cal., panicles large elongated terminal; cal. pilose, nearly 
truncate; standard without callosities at base; stam. distinctly 
diadelphous; pod 24-5 in. by 1 in. wide, tapering to base, 
usually beaked, glabrous, shining, veined, 1—4-seeded, often. 
contracted at the places where seeds have not developed. 

Low country, chiefly on the coast; rather rare. Colombo; Kalutara;: 
Weligama; Batticaloa. Fl. Feb., September; pale pink. 

Also in Malaya. 

Hermann’s excellent specimens, which are in flower but without fruit, 
show Linnzeus’ Sofhora heptaphylla to be nies ; but in his later books 
Linnzus has quoted Rumph. Amboin. iv. t. 22 (which looks like a true 
Sophora), and has thus confused subsequent es (see under Sophora 
zeylanica). 


Hermann gives the name ‘ Radaliya’ for this, which belongs rightly to. 
Connarus monocarpus. 


46. SOPHORA, JL. 


Trees or shrubs, |. imparipinnate, Iflts. without stipels, fl. 
numerous in terminal or leaf-opposed racemes; cal. cam- 
panulate or cup-shaped, oblique, more or less truncate with 
very short segm.; pet. about equal, with long claws, keel-pet. 
obtuse; stam. 10, free to base or nearly so; stigma capitate ; 


Sophora.| Leguminose. 95 


pod moniliform, indehiscent, few-seeded.—Sp. 20; 10 in FZ. 
B. Ind. 


Lfits. 15-23. 
Fl. yellow : : : 4 : ye Pale - I. S. TOMENTOSA. 
Fl. violet . : ‘ : : : : 2a OM VLOLAGCHA: 
Lfits. 7-11; fl. white . : 3 ; ‘ F 5 35 Sb ZBYILAINTICA, 


1. S. tomentosa, Z. Sf. P/. 373 (1753). Mudu-murunga, S. 
é oe Mus. 11. Fl. Zeyl. n. 163. Moon Cat. 33. Thw. Enum. 94. 

ae LAO. 

Big) Hicen. Hort. Lusd- Batt 172: 

A small tree, the branches and whole plant covered with 
short, soft, grey velvety pubescence; |. large, shortly stalked, 
rachis 4-7 in., cylindrical, tumid at base, Iflts. 15-17 (7 or 8 
pair and end one) often alternate, shortly stalked, 1-14 in., 
broadly oval, obtuse and rounded at both ends, the margin 
somewhat reflexed, rather thick, the lower surface with more 
dense pubescence; fl. rather large #-1 in., ped. as long as cal.,. 
jointed near the top, rather closely arranged in stout, erect 
stalked racemes about 6 in. long, bracts acicular, shorter than 
ped., deciduous; cal. somewhat inflated, densely velvety, segm. 
very small, tooth-like; pod long-stalked, 2-6in., moniliform, 
the joints separated by narrow necks as long as themselves, 
sharply pointed, covered with velvety down ; seeds 1-8, 2 in. 
diam., nearly globular, pale brown. 

On the seashore; rather rare. Bentota; Galle; Batticaloa; Jaffna. 
Fl. July, Aug.; sulphur-yellow. 

On Tropical shores throughout the world. 


2. S. violacea, Thw. Enum. 94 (1859). 

CAP: 3546. 

FL. B. Ind. ii. 249. 

A small undershrub, bark pale yellowish-grey, young parts 
finely pubescent, |. numerous, closely placed, spreading, rachis 
5-8 in., slender, cylindrical, nearly glabrous, lflts. 17-23 (8-11 
pair and end one) very shortly stalked, oblong-oval #-1 in., 
obtuse at both ends, glabrous, thin, glaucous beneath; fl. 
numerous, over #in., ped. as long as cal., pubescent, racemes 
erect, 6-8 in., bracts minute; cal. tubular-campanulate, set. 
obliquely on ped., slightly pilose, segm. very shallow, rounded ; 
pod 2-3 in., stalked, moniliform with a broad flattened neck 
between the joints and shorter than them, long-beaked, seeds 
1-4, ovoid, compressed, red, afterwards brown. 

Low country; very rare. Near Dewinuwara bet. Matara and Dik- 


welle, S. Proy. (Thwaites); near Trincomalie (W. Morris). Fl. April, 
Sept.; violet. 


96 Leguminosae. [Pericopsis. 


Endemic. : 
Discovered by Thwaites in 1857. A pretty plant, now much grown in 
Ceylon gardens. 


3. S. zeylanica, 77im. [PLATE XXX.] 

S. heptaphylla, Arn. Pug. 10 (non L.); Thw. Enum. 94; Trim. Syst. 
Gat 27/3 Gaba 570: 

Fl. B. Ind. ii. 250. (S. heptaphylla.) 


A much-branched bush to-12 ft. high, young branches finely 
tomentose with fulvous hair, buds densely brown-velvety; 1. 
numerous, spreading, shortly stalked, rachis 4-6 in., narrowly 
furrowed above, fulvous-tomentose, stip. minute, acicular, lfits. 
7-11 (3-5 pair and end one), often alternate, 15-24 in., ovate- 
oval, acuminate, obtuse, glabrous on both sides, shining above, 
dull and paler beneath; fl. rather small, numerous, articulated 
on divaricate pilose ped. about as long as cal., racemes 4-6 in., 
lax, spreading; cal. finely fulvous-pilose, funnel-shaped and 
thickened at base, above expanding into broadly cup-shaped 
limb, obliquely truncate, segm. very small, tooth-like; pod 
14-2 in., tapering to short stalk, strongly beaked, much 
constricted bet. joints, slightly compressed, thin, veiny, 
glabrous when mature, when young densely covered with 
brown silk ; seeds 1 or 2, 2in., broadly ovoid, bright dark red. 


Montane zone, 4000-7000 ft.; rare. Hunasgiriya (Thwaites); about 
Nuwara Eliya. Fl. Jan., Feb.; white. 

Endemic. The locality ‘China’ given in Fl. B. Ind. is an error, the 
plant there being a species of Cladrastis (see Hemsley in Journ. Linn. 
Soc. xxiii. 201). 

Walker-Arnott (1. c.) seems to have been the first to apply Linnzeus’ 
name S. heptaphylla to this montane plant. The name belongs to Derris 
sinuata (q. v.): Hermann, of course, had no opportunity of collecting 
any hill species. In FI. B. Ind. the flowers are erroneously described 
as yellow. 

S. Wightit, Baker, of the Nilgiri Hills seems to differ chiefly in its 
much larger flowers; this is S. heptaphylla, Wight (non Arn.), figured 
jn cht LS 5. 


47. PERICOPSIS, 7iw. 


A large tree, ]. pinnate with large alternate lfits., fl. large, 
in terminal and axillary panicles; cal. campanulate, attenuated 
below, segm. deeply cut, the upper ones connate; pet. nearly 
equal, keel-pet. slightly coherent ; stam. 10, quite free; stigma 
small, oblique; pod large oblong, compressed, coriaceous, 
indehiscent, few-seeded ; seed compressed, with a very small 
hilum.—Monotypic and endemic. 


Leguminose. 97 


P. Mooniana, 7iw. Enum. 414 (1864). Nédun, S. [PLATE 
BOCK | 

Dalbergia lanceolaria, Moon Cat. 51 (non L. f.). D. Mooniana, Thw. 
Enum. 93. C.P 439. 

Fl. B. Ind. ii. 252. Bedd. Fl. Sylv. t. 187. 


A large tree with thick trunk, bark smooth, pale brownish- 
pink on trunk and flaking off in thin flat pieces, reddish-grey 
on twigs, buds slightly silky; 1. large, rachis 6-10 in., cylin- 
drical, stout, glabrous, much swollen at base, lflts. 6-8, always 
alternate, shortly stalked 24-3 in., oval, rounded at base, 
shortly acuminate, subacute, glabrous on both sides, shining 
above, rather paler beneath; fl. nearly 1 in., ped. stout, nearly 
as long as cal. and jointed immediately beneath it, with 2 
minute bractlets, panicles broad, lax; cal. nearly glabrous, 
thick, tube narrowed below, segm. broadly lanceolate, acute, 
as long as tube; pod 3-6 in. by 14 in. wide, shortly stalked, 
pointed, flattened, coriaceous, glabrous, thickened at the 
sutures, veiny, usually straight, occasionally constricted at 
places where seed has not developed; seeds 1-6, oval, over 
4 in., flattened, smooth, pale orange-red. 


Moist low country to 1000 ft., especially by river banks; formerly 
common, but becoming rarer. Moon’s locality was Kalutara. Fl. Dec.— 
March; dark blackish-purple. 

Endemic. 

The timber is much used for household furniture, and is one of our 
best woods for the purpose, pale orange-brown varied with darker lines, 
smooth and shining, moderately hard, but easily worked. The pod is 
indehiscent, but ultimately breaks up into one-seeded joints. 


II. CASALPINIEA. 


Leaves bipinnate. 


Neither suture of pod winged ; ; . 48. CASALPINIA. 
Both sutures winged . : ‘ | ; . 49. PELTOPHORUM. 
Dorsal suture only winged . , : : . 50. MEZONEURUM. 


Leaves pinnate. 
Cal. without a tube. 


Pet. 5. 
Pod many-seeded . ; , ; : . 51. CASSIA. 
Pod 1-seeded : , , A : . 52. CYNOMETRA. 
Pet. o. 
Stam. 2 5 r 2 5 ; : . 53. DIALIUM. 
Stam. 10 : ; , , ; ‘ . 54. CRUDIA. 
Cal. with a conspicuous tube. 
Pet. o : ; ; ; ; , ‘ - 55. SARACA, 
Pet. 5 ; 2 ; - A ; 7 . 50. HUMBOLDTIA, 
Leaves simple, 2-lobed_ . i 7 i 3 . 57. BAUHINIA. 


PART II. i 


98 Leguuiinose. [Cesalpinia. 


Our 26 species of this sub-order are all low-country plants, only one, 
Cassia mimosoides, reaching a little way up into the montane zone. 
Peltophorum, Dialium and Bauhinia are nearly confined to the dry 
region; Mezoneurum, Crudia and Humboldtia to the moist country ; 
many species of Cassza are very common weeds. There are but two 
endemic species, Dialium ovoideum, and Crudia zeylanica. 


48. CHESALPINIA,* Z. 


Prickly woody climbers, |. abruptly bipinnate, fl. large, in 
terminal or axillary racemes or panicles; cal. with an ex- 
panded flattened base, segm. very deep, imbricate, the lowest 
much the largest and covering the bud like a hood; pet. 
clawed, spreading, the uppermost much the smallest; stam. Io, 
distinct, declinate, 5 often longer than others; pod flattened, 
slowly dehiscent, 1- or few-seeded.—Sp. 40; 10 in FZ. B. nd. 


Pod very spiny (Guzlandina) . : : : . I. C. BONDUC. 
Pod not spiny. 
Lflts. 2-3 pairin each pinna . : ° ° . 2. C. NUGA 
Lfits. 6-9 pair . : é : : ‘ : . 3. C. SEPIARIA. 
Lflts. 10-12 pair. ; : : : : . 4. C. DIGYNA. 


1. ©. Bondue, ford. Hort. Beng. 32 (1814). Kumburu-wel, S. 
Punaikkalaichchi, 7. 


Guilandina Bonduc, L., Moon Cat. 34. Thw. Enum. 94. C. P. 1524. 

Fl. B. Ind. i. 255. Rumph. Herb. Amb. v. t. 48. 

A stout climbing shrub, stems with fine grey pubescence 
and scattered usually straight small prickles; 1. very large, 
main rachis 5—7 in., cylindrical, finely woolly-pubescent, pinnz 
6-8 pair, 14-24 in., spreading nearly at right angles, all with 
numerous hooked prickles, usually in pairs on the under 
surface, no stip., lflts. numerous, 10-14 (5-7 pair) to each 
pinna, very shortly stalked, oval or oblong-oval, rounded at 
base, obtuse, strongly mucronate, glabrous above, pubescent 
on midrib and margin beneath; fl. moderate-sized, articulated 
with ped., ped. as long as cal., pubescent, bracts linear, longer 
than cal., caducous, racemes long-stalked, pyramidal; cal. 
densely tomentose-pubescent ; pod shortly stalked, 24-34 in. 
by 14 in. wide, ovoid, compressed, pubescent and covered with 
very numerous long sharp spines, dehiscent; seeds 1-3, nearly 
globular, $in., with a long funicle, smooth, greenish-grey. 


Low country, especially near the seacoast; rather common, clamber- 
ing over trees and bushes. Fl. Feb.; bright yellow. 


* Commemorates Andrea Czesalpini of Arezzo, the founder of system- 
atic botany. Died 1603. 


Casalpinia.] Leguminose. 99 


Also in India, Malaya, Polynesia, and the W. Indies. 
The root has a reputation as a tonic medicine. 


C. Bonducella, Flem. (Guzlandina, L.). There are specimens of'this 
in Hermann’s Herb., and it is n. 156 of Fl. Zeylan. Moon also (Cat. 34) 
gives it for Kalutara. It is common in nearly all Tropical countries, and 
probably occurs here, but I have not met with it. Distinguished from 
C. Bonduc by its smaller leaflets and the possession of stipules; and 
figured in Rumph. Herb. Amb. v. t. 49, and in Bentl. and Trim. Med. 
PL. t.: $5. 


2. ©. Nuga,* Azz. Hort. Kew. ed. 2, ii. 32 (1811). Diya-wawul- 
étiya, S. 

Fl. Zeyl.n. 157, C. paniculata, Roxb., Moon Cat. 34; Thw. Enum. 
Op nGE! 4525. 

Fl. B. Ind. ii. 255. Wight, Ic. t. 36 (C. paniculata). 

A large woody climber, bark rough, fibrous, yellowish- 
grey, young branches glabrous, dark green, distantly set with 
extremely sharp hooked black prickles which are persistent 
and become ultimately raised on a mamilliform corky base ; 
1. large, main rachis about 5 in., pinne 2 pair, all smooth 
and polished, cylindrical, with very small, sharp, hooked 
prickles beneath at the insertion of the Iflts., lflts. 4 :arely 6 
(2 or 3 pair) distinctly stalked, distant, 2-24 in., ovate-oval, 
obtuse, glabrous, paler beneath; fl. on rather long ped., racemes 
or panicles terminal and axillary, buds globular; cal. glabrous, 
segm. very obtuse; fil. hairy in lower half; pod 24-24 in. by 
1} in. wide, flattened, tapering at base to very short stalk, 
sharply beaked, glabrous, dark brown, leathery-woody, slowly 
dehiscent ; seed solitary, oval-rotund, compressed, over # in., 
smooth, purplish-brown. 

Low country up to 3000 ft.; rather common. FI. August; yellow, 
sweet-scented. 

Aliso throughout the Eastern Tropics. 

The specimen in Hermann’s Herbarium isa very bad one, but certainly 
C. Nuga; it was first called by Linneus Cesalpinia Crista in Sp. PI. 
380, and afterwards, in Mant. ii. 378, referred to G. Bonduc, and he did 
not recognise it as his G. Muga. 

C. Sappan, L., Pattangi, S., the ‘Sappan Wood,’ is a very ancient 
introduction to Ceylon, and is frequently met with in a semi-wild state, 
or planted as afence. There are specimens in Hermann’s Herb. (FI. Zeyl. 
n. 158), and it was formerly much cultivated as a dye-wood, and exported 
to a large amount, but is now little grown. It is native to India and 
Malaya, and is figured in Bedd. FI. Sylv. t. go. 


C. pulcherrima, Sw. (Poinciana, L.), the ‘Peacock-flower,’ is very 
common in native gardens. It is found throughout the Tropics, but not 
known in the wild state. There are specimens in Hermann’s Herb. (FI. 
Zeyl. n. 159). 


* Nuge silvarum is Rumphius’ name for this in Herb. Amboin. v. 
P- 94, t. 50. 


100 Leguminosae. [Caesalpinia: 


3. ©. sepiaria, Roxb. Hort. Beng. 32 (1814). 

Thw. Enum. 95. C. P. 3602. 

Fl. B. Ind. ii. 256. Wight, Ic. t. 37 (poor). 

A stout woody climber, branches finely downy with few 
small yellow prickles ; |. large, main rachis 5—7 in. with 4-6 
pair, pinnz pubescent, with small hooked prickles beneath, 
Iflts. 12-18 (6-9 pair), very shortly stalked, $-3in., oblong, 
rounded at both ends, pilose on both sides, paler beneath; fl. 
large, 14 in., ped. 1-1} in., stout, hairy, ascending, racemes 
terminal, large 6-12 in. long, erect, bracts long, lanceolate,, 
acuminate, caducous; cal. finely but densely pubescent, segm. 
very obtuse, reflexed, pet. shortly clawed, reflexed; stam. 
much exserted, fil. woolly for more than lower half; pod 4 in. 
by 1% wide, not stalked, sharply beaked, pubescent with 
deciduous hairs, orange-brown, somewhat veiny, tardily 
dehiscent ; seeds 6-8, oblong, 2in., greenish, mottled. 


Var. 8, auricoma, 77277. 


More robust, whole plant including prickles densely 
covered with a velvet of soft spreading bright golden-yellow | 
hair; |. larger 9-12in., with 8-10 pair of pinnz; prickles. 
larger and more numerous. 

Low country, and up to 4500 ft., especially in the dry and intermediate 
regions. Var. 8, lower montane region to 5000 ft. Matale; Maturata;, 
Uda Pusselawe. Fl. March, April, September; bright chrome-yellow, 
filaments crimson. 

Also in India, Malaya, China, and Japan. 

Var. 8 is a very handsome plant, which further examination may, show 
to be a distinct species. I have not seen the fruit. 


[C. mzmosozdes, Lam., is recorded for Ceylon in Fl. B. Ind. ii. 256, but 


I am not aware upon what authority. It is native to Peninsular India. 
and Burma. | 


4. ©. digyna, Rottl. in Neue Schrift. Naturf. Freunde Berlin, iv.. 
198 (1803). 

Thw. Enum. 95. C. P. 1527. 

Benda 25 Os iNottlnlicy i113: 

A woody climber, branches glabrous, purple, with hooked 
brown prickles, young parts rufous-pubescent; |. narrow, 
main rachis 6-8 in., with 9-12 pair of pinnz, with rachis 14-2 
in., pubescent, lflts. 20-24 (10-12 pair), 2 in., sessile, closely 
placed, overlapping, oblong, very obtuse, pubescent on both 
sides, thin ; fl. $in., ped. 1 in., horizontally divaricate, slender,. 
racemes stalked, axillary, 6-8 in., bracts setaceous, falling very 
early; cal. glabrous, segm. separating from the flat base; stam. 
little exserted, fil. very woolly for more than basal half; pod 
(not seen ripe) ‘oblong, rather fleshy, indehiscent, 24 in. by 


Peltophorum.] Leguminose. IOI 


I in. wide, smooth, torulose, 2—4-seeded,’ seeds ovoid, ‘size of 
a large pea.’ 
Dry country; very rare. Kurunegala; Doluwe Kande. 


Fl. September; yellow. 
Also in India and Malaya. 


C. coriarta, Thunb., the ‘ Divi-divi’ tree, a native of Central America, 
thas been long since introduced, and planted trees are now not unfre- 
quently met with in Rest-house compounds and similar places in the dry 
districts and called Vaznz, T. 


49. PELTOPHORUM, Vogel. 


A large tree, |. abruptly bipinnate, fl. large, in terminal 
and axillary panicles; cal. cut more than half way down, segm. 
nearly equal; pet. large, with a broad claw; stam. Io, declinate, 
anth. uniform; stigma large, peltate; pod very flat, thin, 
indehiscent, with a broad flat wing along both sutures, 1-3- 
seeded.—Sp. 6; 2 in #2. £. Lnd. 


P. ferrugineum, Zenith. Fl. Austral. ii. 279 (1864). Tya-vakai, 
Z. [PLATE XXXII.] 

Cesalpinia Gleniez, Thw. Enum. 414. C. P. 3815. 

FL B. Ind. 11. 257. 

A very large tree, much branched above, with rather smooth 
grey bark, young parts covered with orange-brown velvety 
tomentum; |. large, main rachis 6-12in., secondary ones 
3-4 in., all swollen at base, narrowly channelled above, rusty- 
pubescent, stip. minute, early caducous, pinne 6-10 pair, 
spreading horizontally, lflts. 20-24 (10-12 pair), sessile, 4—? in., 
oblong, emarginate, unequal-sided, the upper half the larger, 
glabrous above, rusty-pubescent on midrib beneath, firm; fl. 
large, 14 in,, on pubescent viscous ped. shorter than cal., in 
large erect panicles with rusty viscous pubescence; cal.-segm, 
obtuse or subacute, more or less covered with ferrugineous hair, 
strongly reflexed, pet. with similar hair on the claw and on 
back, much crisped, somewhat reflexed, filaments with tufts of 
silky hair at base; pod 2-2 in. by 1 in. wide, tapering to very 
short stalk and to obtuse or subacute apex, glabrous, striated 
with longitudinal veins, reddish-brown, wings broad sharp, 
rigid ; seed oblong, flattened, 2 in., smooth, yellowish-grey. 

Dry region; rather rare. Trincomalie (Glenie); Batticaloa (Capt. A, 
M. Walker). Fl. May-September; bright dark yellow, sweet-scented. 

Also found in the Andaman Is., Malacca, Singapore, Malay Islands, 
and N. Australia, but not in Peninsular India. 


Has been much planted for ornament, and is a magnificent sight when 
in full blossom. 


102 Leguminosae. [Cassia 


50. MEZONEURUM, Des/. 


A prickly woody climber like Cesalpinza in habit and fl.; 
lowest cal. segm. very much enlarged and hooded in bud ; 
stigma small, capitate; pod large, thin and papery, very flat, 
indehiscent, with a broad papery wing along the dorsal suture; 
seeds few, small.—Sp. 10; 5 in FZ. B. [nd. 


IZ. enneaphylilum, VW. and A. Prod. 283 (1834). Goda- 
wawulétiya, 5S. 

Mhwa tnume Ardy 1G. by s60r- 

Fl. B. Ind. ii. 258. Miq. Fl. Ind. Bat. i. 1, t. ii. f. B (flower and fruit). 


A climbing shrub, stems finely pubescent or glabrous, set 
with scattered decurved prickles, 1. large, main rachis 7-12 in., 
cylindrical, branch-like secondary ones 14-3 in., horizontal or 
deflexed, with pairs of hooked prickles beneath at the origin, 
pinne 8-12 pair, distant, lfits. 10-16 (5-8 pair) shortly stalked, 
oval or obovate-oblong, rounded or omnes. glabrous or 
pubescent, much paler beneath; fl. $in., pod 2 in., “horizontal, 
pubescent, in axillary or supra- elles? long- stalked racemes 
6-12 in. long; cal. densely pubescent; fil. very hairy in lower 
half; pod 5-7 in, by 14-14 wide, pendulous, not stalked,. 
tapering to a somewhat hooked apex, very thin, the wing 
4 in. wide, z.e., nearly half the width of pod so as to cause the 
dorsal suture to look like a vein running along its centre, 
brilliant pinkish-crimson; seeds 3-5, very smail. 


Hoist low country up to 2000 ft.; rather common. FI. July See 
ber; yellow. 


Also in Burma and Malay Archipelago. 


The thin, papery, crimson pods hanging in large cheers are very con- 
spicuous. 


It is possible that the above includes JZ. pudescens, Desf., which is 
given for Ceylon in Fl. B. Ind. ii. 259. 


Parkinsontia aculeata, L.,a native of Trop. America, isa not uncommon 
small tree in the dry districts, especially about Jaffna; usually planted,. 
but sometimes self-sown on borders of paddy-fields, &c. 


51. CASSTA, Z. 


Trees, shrubs or more rarely herbs, |. abruptly pinnate 
often with glands on the rachis, fl. usually large, in axillary 
racemes or terminal panicles; cal. cut almost to the base, 
segm. (nearly separate sep.) much imbricate; pet. usually 
nearly equal; stam. 10 (very rarely 5), either all perfect 
or more usually with the 3 upper barren, anth. opening by 


Cassia.] Leguminose. 103 


pores (rarely by slits); pod various, flat or cylindrical, 
dehiscent or indehiscent, with partitions between the usually 
numerous seeds; seed with more or less abundant endosperm, 
embryo with flat cotyledons.—Sp. about 340; 18 in FZ. B. Ind. 


Pod terete, very large. 
Lfits. 4-6 . : ‘ : 5 ; é > la (Gy IPTSIROILA, 


Lfits. 20-30 : : : : : ., 2. C. MARGINATA. 
Pod more or less flattened. 
Three upper stam. barren. 
Petiole with one gland at base. 


Lfits. 8-Io . 3. C. OCCIDENTALIS. 
Lfits. 12-20 4. C. SOPHERA. 
Petiole with glands between one or more 
pairs of lfits. 
Lfits. 6 5. C. TORA. 
Lfits. 16-24 ; : 6. C. AURICULATA. 
Petiole without glands. 
Lfits. 8-12 ; pod 14 in. 7. C. OBTUSA. 
Lfits. 12-20; pod 8-12 in.. 8. C. SIAMEA. 
Lfits. 20-30 ; pod 4 in. g. C. TIMORIENSIS. 
All stam. perfect. : 
Stam. 5; Iflts. 4 . : : 2 5 Mus (Cy ANSISUS. 
Stam. 10; Iflts. very numerous. 
Gland on petiole distinctly stalked. 5 tilts (G5 Iie arari 
Gland on petiole sessile . . ‘ . 12. C. MIMOSOIDES. 


I. C. Fistula,* Z. Sp. P/. 377 (1753). Bhéla, S. Tirukkontai, 


Ravani, 7. 
Burm. Thes. 56. Fl. Zeyl.n.149. Cathartocarpus Fistula, Pers., 


Moon Cat. 34. Thw. Enum.g95. C. P. 1502. 
Fl. B. Ind. ii. 261. Wight, Ic. t. 269 (C. rhombifolia). Bentl. and 


Trim. Med. PI. ii. t. 87. 


A small or moderate-sized tree, with a straight trunk and 
slender branches, bark smooth and pale grey when young, 
dark brown and rough when old; 1. large, rachis 6-12 in., 
glabrous, stip. minute, setaceous, lflts. 4-12 (2-6 pair) distinctly 
stalked, articulated with rachis, 2-6in., ovate, acute at base, 
obtusely acuminate, quite glabrous, stiff, pale yellowish-grey, 
veiny; fl. large 14-2in., on slender glabrous spreading ped. 
about 14 in. long, in very lax long-stalked pendulous axillary 
racemes longer than 1, bracts lanceolate, caducous; cal. 
glabrous, segm. concave, very obtuse; pet. concave, veined ; 
3 lowest stam. with very long doubly curved exserted fil. and 
oblong anth. dehiscing longitudinally, 4 lat. ones with short 
straight fil. and versatile anth. opening by pores at the base, 


* * Cassia fistula’ was the medizeval pharmacists’ name for the pods, 
having been transferred to them from some variety of Cinnamon Bark. 


104 Leguminose. [Cassta. 


3 uppermost ones much smaller, erect, with indehiscent abortive 
anth.; pod 1-2 ft., pendulous, shortly stalked, cylindrical 
#-1 in. diam., nearly straight, hard, smooth, shining, purplish- 
black, not torulose, indehiscent, with very numerous, 25-100, 
horizontal seeds immersed in dark-coloured pulp and com- 
pletely separated by thin transverse partitions ; seed small, 
4in., ovoid, slightly compressed, with a raised raphe down 
whole of one side, hard, smooth, shining, pale orange-brown, 
cotyledons flat, foliaceous, endosperm abundant, hard and 
horny. 
Rene of the dry region; very common. Fl. July-August; pale 
ellow. 
Also in India, China, and Malaya. 
A conspicuous tree in the dry country whether covered with its droop- 
ing racemes of bright flowers or its pendulous black cylindrical pods. 
The flowers are used in temple ceremonies; the astringent bark for 
tanning and as a medicine; and the pulp of the fruit is a well-known 


purgative. Heart-wood very hard and heavy, dark red, durable, but of 
no great size. 


2, ©. marginata, Roxb. Hort. Beng. 31 (1814). Ratu-wa, S. 
Vakai, 7. 


Cathartocarpus rosea, Moon Cat. 34. C. Roxburghit, DC., Thw. 
Enum OG C-eb. ai5O5. 


Fl. B. Ind. 11. 262. Wight, Ill. t. 83 (C. Roxburghiz), copied in Bedd. 
F]. Sylv. t. 80. 


A small tree, with slender drooping branches, bark brown, 
deeply cracked longitudinally, young parts densely pubescent 
or tomentose; |.-rachis 6-10 in., slender, furrowed above, pubes- 
cent, stip. large, falcate, pubescent ; quickly deciduous, Ifits. 
20-30 (10-15 pair), shortly stalked, 1-1} in., oblong, obtuse, 
emarginate and apiculate at apex, unequal at base, glabrous 
above, pubescent beneath ; fl. about ? in., on pubescent ped. 
# in. long, in close pedunculate supra-axillary racemes much 
shorter than |, bracts ovate, cuspidate, soon falling; cal. densely 

ilky, segm. subacute; two lower pet. larger than others; 3 
owest stam. with long doubly curved exserted fil. and large 
anth., other 7 very much shorter with smaller anth. all fertile; 
pod 8-12 in., cylindrical, straight, about 2 in. diam., semi- 
woody, somewhat torulose, with very numerous seeds separated 
by partitions, glabrous, black; seed about 4 in. compressed, 
pale brown, with abundant endosperm. 

Dry region; very common. FI. July, August; pink or salmon- 
coloured. : 

Aliso in Southern India. 


Very ornamental when covered with its abundantly produced flowers. 
Heart-wood heavy, very hard, reddish brown, small. 


Cassia. Leguminosae. 105 


3. ©. occidentalis, Z. Sf. Pl. 377 (1753).  Peni-tora, S. 
Ponnantakarai, 7. 

C. Tagera, Moon Cat. 33 (? Linn). Thw. Enum. 95. C. P. 1262. 

Fl. B. Ind. ii. 262. Bot. Reg. t. 83. 

A coarse annual, 1-3 ft. high, often semi-woody at base, 
stam. slightly branched, nearly glabrous, furrowed, purplish ; 
1. rather large, rachis stout, 44-6 in., glabrous, swollen at base, 
where on the upper side is a sessile purple gland, stip. large, 
semi-sagittate, acuminate, caducous, lflts. 8 or 10 (4 or 5 pair), 
on very short stalks, 2-4 in. (the end pair largest) narrowly 
Ovate-lanceolate, rounded and unequal at base, gradually 
attenuate to very acute apex, glabrous, very thin, dark-green 
above, slightly glaucous beneath; fl. large, 1} in. or more, ped. 
about 4 in., usually in pairs in the leaf-axils; cal.-segm. obtuse, 
nearly glabrous; 7 stam. with perfect anth. opening by pores, 
2 or 3 lowest longer and declinate, other 3 erect with barren 
anth.; pod 34-41 in. by } in. wide, erect, stiff, linear, slightly 
curved, bluntly apiculate, flattened, slightly compressed 
between the seeds, with thickened sutures; seeds 20-30, com- 
pressed-ovoid, pointed at one end, shining, very hard, olive- 
green. 

Low country in waste places; a verycommon weed. Fl. June, March, 
&c.; bright orange-yellow. 

Throughout the Tropics, and perhaps originally from the New World. 
The colour of the flowers in FI. B. Ind. is wrongly given as ‘pale 
lilac’ The plant often strongly tinged with purple. 


Very fetid when bruised. Both this and C. Zora are much used as 
alterative medicines. 


é a Sophera,* Z. Sp. Pl. 379 (1753). Uru-tora, S. Taka- 
rai, 7. 


EL Zeyl n. 150, Moon Cat. 33 Thw. Enum: 97. C. P. 1507. 

Fl. B. Ind. ii. 262. Burm. Thes. t. 98. 

A large annual, more or less woody below, stem slightly 
branched, glabrous, 1. large, rachis 5-7 in., furrowed above, 
nearly glabrous, swollen at base and provided there with a 
large solitary gland, lflts. 12-20 (6-10 pair) closely placed, 
shortly stalked, 14-2} in., lanceolate, rounded at base, acute, 
finely mucronate, thin; fl. as in the last, but 4-7 in an axillary 
raceme; pod about 34in. by 2 wide, nearly straight, somewhat 
turgid, glabrous, acute, sutures broad, thickened, with very 
numerous small seeds separated by partitions. 

Low country, especially in the dry region; common. FI. Feb. and 
March; pale yellow. 

Found throughout the Tropics. 


* Sophera, a name taken from Prosper Alpinus, and applied by him 
to some Egyptian plant, perhaps this. 


106 Leguminose. [Cassia 


5. ©. Tora, LZ. Sd. Pl. 376 (1753). Wéti-tora, S. VWaddu- 
takarai, 7. 

Burm. Thes. 58. Herm. Hort. Lugd.-Bat. 557. FI. Zeyl. n. 152. 
Moon Cat. 33. Thw. Enum. 96. C. P. 2785. 

HSB indi 2635) ill Elort leith: 63. 

An annual herb, 1-2 ft. high, with spreading glabrous. 
branches; |. small, numerous, long-stalked, rachis 1-2 in., 
stout, furrowed, slightly pubescent, with a linear yellow gland 
above between the Iflts. of the lower one or two pair, stip. 
long, setaceous, Iflts. 6 (3 pair), very shortly stalked, 1-14 in., 
(the terminal pair largest), obovate, tapering to unequal base, 
apiculate, glabrous above, very finely silky beneath, thin and 
flaccid; fl. about 2in., solitary or in pairs, axillary, ped. 
slender, 3? in., cal. glabrous, 2 segm. larger than others ; upper 
pet. 2-lobed, the others entire; barren stam. minute; pod very 
long, 6-8 in. by less than } in. wide, linear, much curved when 
young, afterwards less so, quadrangular (the sutures being 
very wide), glabrous, with about 30 seeds and very slight 
partitions between them, thin, pale brown; seed not com- 
pressed, pointed at one end, rather truncate at the other, very 
hard, rough with scurfy hair, pale brown. 

Throughout the low country by roadsides and waste ground; a very 
common weed, especially in the dry region. Fl. Jan., August, &c.; rather 
pale orange-yellow. 

Found throughout the Tropics. 

Hermann, followed by other writers, gives the native name ‘ Tala’ for 
this (Mus. 69); probably a misprint for Tora. Has a peculiar fetid scent. 
when bruised. The seeds are sometimes eaten. 


C. omentosa, L., Thw. Enum. 95, C. P. 2410, is a Tropical American 
species, cultivated and naturalised about Nuwara Eliya and elsewhere in 
the mountains. 


C. hirsuta, L., is a frequent weed about Kandy and elsewhere. It is 
also an introduction from Trop. America. 


C. levigata, Willd., also introduced from Trop. America, is completely 
naturalised, and a very common weed throughout the montane zone, in 
the Planting Districts. 


6. ©. auriculata, ZL. Sf. Pl. 379 (1753). Ranawara, S. 
Avarai, 7. [PLATE XXXIII.] 

Herm. Mus. 56. Fl. Zeyl.n. 151. Moon Cat. 33. Thw. Enum. 96. 
C. P. 1503. 

Bee lidiie263" 

A large much-branched bush, bark smooth, cinnamon- 
brown, branchlets closely pubescent; 1. very numerous, closely 
placed, rachis 34-5 in., narrowly furrowed, slender, pubescent, 
with an erect linear gland between the Iflts. of each pair, stip. 
very large, reniform-rotund, produced at base on side next 
petiole into a filiform point, persistent, lflts. 16-24 (8-12 pair), 


Cassia.] Legsuminose. 107 


very shortly stalked, slightly overlapping, oval-oblong, obtuse at 
both ends, mucronate, glabrous or minutely downy, dull green, 
paler beneath ; fl. large, nearly 2 in., ped. glabrous, 1 in., 
racemes few-flowered, short, erect, crowded in axils of upper 1. 
so as to form a large terminal inflor. (the |, with the exception 
of their large stip., being suppressed at the upper nodes); cal. 
glabrous, segm. very unequal, concave, membranous; pet. with 
rather long claws, crisped on margin; pod short, 3-44 in. by 
2in. wide, oblong, obtuse, tipped with long style-base, flat, 
thin, papery, undulately crumpled, pilose, pale brown; seeds 
12-20, distant, each in a separate cavity. 

Low country, especially in the dry region and near the coast; common;. 
locally gregarious and abundant. FI. Feb., March; bright yellow. 

Also in Peninsular India. 

A great ornament to the barren coast regions. 

The leaves of this form ‘Ceylon Tea’ or ‘ Matara Tea,’ and the in- 
fusion is drunk by the natives on several parts of the coast; when 
properly cured, rolled, and dried, it is very like true tea in appearance, 
and somewhat aromatic,* but slightly purgative. The bark is rich in 
tannin, and is largely employed for tanning leather in Colombo. 


7. ©. obtusa, Roxb. Hort. Calc. 31 (1814). Nilavakai, 7. 

Thw. Enum. 441. C. P. 3843. 

Fl. B. Ind. ii. 264 (C. obovata; not given for Ceylon). Wight, Ic. t. 
757. Bentl. and Trim. Med. PI. ii. t. 89 (C. odovata). 

Annual (or perennial?) 1-2 ft., somewhat woody below, 
with spreading, glabrous, angular branches; |. spreading, 
rachis 2—24in., glabrous, without glands, stip. small, linear- 
lanceolate, acute, rigid, persistent, lflts. 8-12 (4-6 pair), 3-1 in., 
very shortly stalked often somewhat overlapping, broadly 
oblong or obovate-oblong, obtuse at both ends, mucronate, 
unequal-sided at base, glabrous or nearly so, rather stiff, pale 
green, glaucous beneath; fl. about ? in., shortly stalled, 
racemes stalked, erect, lax, axillary, short but lengthening 
with fruit and ultimately exceeding 1.; cal.-segm. very obtuse, 
membranous, veined; perfect stam. with short fil.and unequal 
anth., abortive ones very small; pod 1} in. or a little more by 
% wide, very shortly stalked, oblong, much curved upwards, 
rounded at both ends, tipped with small curved style-base, 
very flat, parchment -like, flexible, glabrous or nearly so,. 
greyish-brown, veiny, with a line of prominent rounded crests 
on the sides over the seeds; seeds 8-10, separated by very 


other nitrogenous substances, with 41°14 per cent. of gum, tannin, and 
saccharine matters. 


108 Leguminose. [ Cassia. 


compressed, with a large hilum at the pointed end, endosperm 
scanty. 


Dry region; rare. About Jaffna, frequent; Trincomalie (Glenie) ; 
Mannar (Crawford). Fl. Dec.—Jan.; yellow. 

Also in desert parts of India, E. Trop. Africa, and Arabia. 

One of the true medicinal sennas; I think wild at Jaffna, where, how- 
ever, it is also cultivated for use as a drug. 


C. alata, L., ‘ Rata-tora,’ S., a stately handsome plant, is common by 
paddy-fields and other wet places in the low country. It does not any- 
where here look like a native, but is found in all Tropical countries. Well 
-worth cultivation. 


8. G. siamea, Lam. Encycl. i. 648 (1783). Wa, Aramana, S. 
Vakai, T. 

C. sumatrana, Roxb. Moon Cat. 33. C. florida, Vahl, Thw. Enum. 
06. C. P. 3485. 

Fl. B. Ind. 11. 264. Bedd. FI. Sylv. t. 179. 

A moderate-sized or small tree, bark grey, nearly smooth, 
shoots glabrous; |. rather large, spreading or deflexed, rachis 
6-10 in., stout, stiff, swollen at base, shallowly channelled 
above, nearly glabrous, stip. caducous, lflts. 12-20 (6-10 pair), 
stalked, 24-3in., oblong-lanceolate, rounded at both ends, 
tipped with minute sharp mucro, glabrous on both sides, 
glaucous beneath, rather thick, stiff; fl. 11-14 in, ped. about 
I in., downy, crowded at ends of stout pedunculate axillary 
and termina] corymbose racemes or panicles, the whole forming 
together a very large pyramidal terminal inflor. 2 ft. or more 
long, bracts curved, linear, stiff, fleshy, much shorter than ped., 
cal.-segm. slightly pubescent; pet. distant, distinctly clawed ; 
fil. of 2 of the fertile stam. twice as long as the rest; pod very 
long, 8-12in. by 4-2 in. wide, stalked, flat, beaked, flexible, 
very finely and closely pubescent, thickened at sutures, 8-15- 
seeded. 

Low country to 2000 ft., chiefly in the moist region; common, especi- 
ally near rivers and streams. Fl. March, November; rather pale yellow. 

Also in India and Malaya. 


Heart wood very hard, blackish-brown, durable ; a beautiful and use- 
ful wood for small articles. 


9. ©. timoriensis, DC. Prod. ii. 499 (1825). 

Thw. Enum. 96. C. P. 1504. 

HBS Inds 2652 

A small tree, twigs pubescent; |. large, rachis 7—I0 in., 
cylindrical, channelled above, pubescent, stip. foliaceous, 
broad, with a filiform tail, semi-persistent, lflts. 20-30 (10-15 
pair) shortly stalked, 14-12 in. narrowly oblong, obtuse, 
strongly apiculate, pubescent on both sides especially beneath ; 
fl, If in., on 7 in. ped. in short numerous corymbose panicles 


Cassia.] Leguminose. 10 


from axils of young 1, the whole forming a large terminal 
inflor.; cal.-segm. pubescent; pet. rather narrow, fil. of perfect 
stam. all very short; pod about 4 in. by 4 in. wide, very 
shortly stalked, pointed, very flat, the sutures not thickened, 
glabrous and shining, thin and flexible, 14-20-seeded with 
raised transverse lines marking the partitions between them, 
brownish-black. 

Moist low country; rather rare. Heneratgoda; Sitawaka; Four 
Korales Dist. Fl. September; yellow. 

Also found in Burma and the Malay Islands, the Philippines, and N. 
Australia, but not in Peninsular India. 

C. glauca, Lam., I know here only as a garden plant. There are 
specimens from Moon (C. P. 1526) in Herb. Perad. and in his Cat. (p. 33) 
he gives Colombo as locality. See also Thw. Enum. 96. There are also: 
specimens from Macrae in Herb. Kew. 

Some leaves from Trincomalie, collected by Glenie in 1864, are in 
Herb. Perad. labelled by Thwaites ‘(?) C. suf/ruticosa, Koenig, but are 
scarcely determinable. 


10. ©. Absus,* Z. SA. Pl. 376 (1753). Bu-tora, S. 

Herm. Mus. 15. Burm. Thes. 212. Fl. Zeyl.n. 153. Moon Cat. 33. 
Thw. Enum. 96. C. P. 1506. 

Fl. B. Ind. ii. 265. Burm. Thes. t. 97. 

An erect annual, 6-18 in. high, with few horizontal 
branches densely covered with spreading glandular hair; 1. 
small, spreading or deflexed, rachis 3-1 in., swollen at base, 
covered like the stem with viscous hairs, with an erect minute 
linear gland between the lfits. of every pair, stip. small, linear, 
Ifits. 4 (2 pair) very shortly stalked, closely placed at end of 
rachis, broadly oval or obovate-oval, unequal at base, rounded 
with a minute mucro at apex, glabrous above, slightly hairy 
but not glandular beneath, the terminal pair the larger; fl. 
small, a little over 4 in., on very viscous ped. longer than cal.,. 
in leaf-opposed erect racemes longer than 1, cal. slightly 
glandular-hairy, segm. oblong; pet. with long claws, veiny; 
stam. 5, equal, all fertile, opening by slits not pores; ov.densely 
bristly; pod 1-2 in. by }-3 in. wide, nearly straight, oblique, 
tipped with short style-base, compressed, with long, scattered, 
white, bristly hairs, 6-8-seeded, with depressed lines outside 
at the partitions; seed trapezoid, compressed, dark brown, 
brilliantly shining. 

Low country, especially in the dry region; common. Fl. Dec., Jan.; 
yellowish-pink. 

Found throughout the Tropics of the Old World. 

A very distinct species. Hermann gives the name ‘ Telamandukola’ 
for this. The seeds are employed in medicine. 


* Absus is the name of Prosper Alpinus for this, probably taken from 
some Egyptian one. 


110 Leguminose. [Cassda. 


11. ©. Rleinii, W. and A. Prod. 293 (1834). Bin-siyambala, 5. 

Thw. Enum. 96. C. P. 1508. 

EB: Ind: 11.1266: 

A perennial herb with a woody base, 1-3 ft. high, much- 
branched, spreading, rigid, glabrous or pilose; 1. numerous, 
rachis 1-14 in., pilose, with a small, distinctly stalked peltate 
gland just below lowest pair of Iflts., stip. linear-acuminate, 
striate, acute, rigid, persistent, lflts. 16-40 (8-20 pair) sessile, 
crowded, overlapping, about 4in., strongly mucronate, oblong, 
oblique, the midrib close to upper edge; fl. $ in., on slender 
ped. $ in. long, 1-3 in. the axils; cal.-segm. very acute; stam. 
10, fil. unequal; pod 1?-2 in. by tin. wide, slightly curved 
upwards, abruptly pointed, sparingly pilose especially on 
suture, 10O—-12-seeded, dark brown. 

Var. B, pilosa, Tiw.7.c.97. 


Branches and Iflts. densely clothed with short spreading 
hair. 
Low country, in moist and dry regions; very common. Var. £, 


Hantane. Fl. July, August; yellow. 
Also in Peninsular India and Java. 


C. pumila, L., is given for Ceylon in Fl. B. Ind. I have seen no 
specimens. It appears to differ from C. A/ezwzz principally in having 
5 stamens instead of Io. 


12. ©. mimosoides, LZ. Sf. P/. 379 (1753). Bin-siyambala, 5S. 

Herm. Mus. 18. Burm. Thes. 160. Fl. Zeyl. n. 154. Moon Cat. 33. 
C. angustissima, Lam., W. and A. Prod. 292. Thw. Enum. 96. C. P. 
1510. 

: IMG 185 liaGh Mh, AOS 

Annual but often woody at base, usually erect, 2-3 ft. high, 
with long ascending stiff pilose branches ; 1.-rachis 24—3 in., 
very slender, curved, glabrous or hairy, with a flat, sessile 
gland between the lfits. of the basal pair, stip. linear, setaceous, 
persistent ; lflts. minute, § in., sessile, very crowded and over- 
lapping, 60-100 (30-50 pair) or more, linear, very unequal- 
sided, glabrous or somewhat ciliate ; fl. on slender hairy ped. 
over 4in. long and with 2 small bracts beneath fl. 2 or 3 
together in axils; sep. acute, pilose; pod 2-21 in. by ? in. 
wide, linear, slightly curved, sparingly pilose, about 20-seeded, 
slightly constricted at the partitions. 

ae 8, auricoma, Grah. (sp.). Var. vzllosula, Thw. |.c. 96. C. P. 

3903. 

Whole plant sparingly covered with spreading yellow hair; 
Iflts. larger, aristate. 

Var. y, Wallichiana, Baker. C. Wallichiana, DC., Thw. Enum. 
Sh (Ga. 125 BBO: 


Cynometra. Leguminosae. Ill 


Stems stouter, gland on rachis more prominent but not 
stalked, placed rather below the lowest pair of Ifits., lfits. fewer, 
much larger, }-3in., aristate; pods smaller, 1}-1# in. 

Low country and lower montane zone to 4000 ft. in open grassy 
places; common. Var. 8, Hantane; Maturata. Var. y, the commonest 
variety. Fl. April, August; yellow. 

Found throughout the Tropics. . 

The leaves are very sensitive. I know of no figure of this common 
Pee A. 

Var. y, I should prefer, with Thwaites, to regard as a distinct species ; 
it is a common weed about Kandy, and very distinct in appearance, 
having red-purple stems and blue-green foliage. 


52. CYNOMETRA, J. 


Trees, |. abruptly pinnate with 2 or 4 lfits, fl. small in 
axillary racemes; cal. very deeply cleft almost to base; pet. 5, 
equal, narrow; stam. 10, equal, spreading ; ov. with 2 ovules; 
pod leathery-woody, indehiscent, obliquely ovoid, 1-seeded ; 
seed large, without endosperm, cotyledons plane-convex.— 
eo, 5 in 77. B. Ind, 


Cc. ramifiora, LZ. Sf. Pl. 382 (1753). Gal-mendora, S. 
Attukaddupuli, 7. 


Moon Cat. 33. Thw. Enum. 97 (var. a). C. P. 3604. 

Fl. B. Ind. 11. 267. Rumph. Herb. Amb. 1 t. 63. 

A small or moderate-sized much-branched tree with rather 
smooth, brownish-grey bark, twigs slender, pendulous, young 
parts glabrous, |. composed of I pair of sessile Iflts., articulated 
on a short thick wrinkled rachis }—3 in. long, Iflts. 44-6 in., 
approximated not spreading, linear-lanceolate, very unequal- 
sided at base, tapering to long acuminate obtuse apex, stiff, 
glabrous on both sides, with rather conspicuous but not 
prominent venation; fl. small, on very slender ped. }-2 in. 
long, numerous, in short, spreading, sessile, axillary, corymbose 
racemes barely 1 in. long, bracts small, broad, pubescent, cal.- 
segment glabrous; pet. narrowly lanceolate, acute; stam. twice 
as long as pet., spreading ; pod not seen (but see under Crudia 
zeylanica, p. 113). 

Var. 8, heterophylla, 7iw./.c. C.P.1500. Bedd. Fl. Sylv. t. 315 
(from Ceylon specimens). 

Lfits. very variable, usually 4 (2 pair), of which the lower 
are much the smaller often not more than 3 in., the upper 
reaching 4 in., oval, obtuse or emarginate ; pod from 4-1 in., 
compressed-turgid, nearly straight along dorsal suture, very 
gibbous on ventral, often deeper than long, shortly but dis- 
tinctly beaked, very deeply rugose. 


ion Leg UMLNOSE. [ Dialium.. 


Forests of the low country, in the dry region; rare. Only seen from 
Uva. Var. 6 much commoner, and in both moist and dry regions. 
Fl. Jan.; white. 

Found also in Peninsular India, Malay Islands, and N. Australia. 

There is no specimen in Hermann’s Herb., and his drawing is 
indeterminable. 

I suspect there are here two distinct species, but I have no fruit of 
the type. A very small-leaved form of var. 8 from the coast at Puttalam 
and Trincomalie may be C. mzmosozdes, Wall. The young foliage of the 
type form is pendulous, flaccid, and white. The S. name is unfortunate, 
as this tree has no similarity to the true ‘ Mendora’ trees, which are all 
Dipterocarps. 


C. caulifiora, L., is cultivated in gardens, and is included in Linnzeus” 
Fl. Zeyl. n. 166. It is known by its Malay name, ‘ Nam-nam,’ and has no. 
claim to be considered native here. 


53. DIALIUM, Z. 


Trees, 1. imparipinnate, fl. small, in terminal panicles; cal. 
divided nearly to base into 5 much-imbricated petaloid seem. ; 
pet. 0; stam. 2, anth. large, extrorse; ov. with 2 ovules; pod 
small, indehiscent, ovoid, densely covered with velvety pu- 
bescence; seed solitary; embryo surrounded by endosperm.— 
Sp. 10; 5 in FZ B. Ind. 


D. ovoideum, Z7iw. Enum. 97 (1859). Gal-siyambala, S. 
Kaddupuli, 7. 

thw Enum O75) €. bs 3TA0: 

Fl. B. Ind. ii. 269. Bedd. Fl. Sylv. t. 181. 

A tall tree, bark rather smooth, twigs slender; 1. 
rather small, rachis 14-24 in., slender, cylindrical, glabrous, 
Iflts. usually 5 (2 pair and end one) generally alternate, shortly 
stalked, lanceolate or ovate-lanceolate, rounded at base, 
bluntly acuminate, glabrous and shining on both sides, dark 
green; fl. small, numerous, in erect, much-branched, pyra- 
midal, terminal panicles, ped. shorter than cal., grey with 
adpressed pubescence, buds linear, pointed; cal.-segm. lan- 
ceolate, subacute, faintly ciliate ; stam. as long as sep. erect ; 
pod about #in., broadly ovoid, slightly compressed, densely 
covered with fine brown velvety hairs, with a TERY thin and 
brittle shell and a spongy interior ; seed nearly in. pale 
brown, embryo green. 

Low country in the dry or intermediate region ; rather rare. Hara- 
gama and Billahul-oya, east of Kandy; several places in Eastern 
Province. Fl. April; white. 

Endemic. 

The fruit is known as the ‘ wild Tamarind’ or ‘velvet Tamarind,’ and 
has a pleasant acid flavour ; the endocarp in which the taste resides is of 
a spongy-powdery, not pulpy, consistence and buff-coloured. Not un- 


Saraca.] Leguminose. 113 


frequently the fruit is attacked by an insect, and then forms a large soft 
lobed spherical gall with numerous small cavities. 
Heart-wood hard, heavy, dark red, strong, and handsome. 


54. CRUDIA, Schred. (?) 


A tree, |. pinnate, Iflts. alternate, fl. in terminal racemes ; 
cal.-tube very short, segm. 4 or 5,rotundate; pet. 0; stam. Io, 
free, the alternate ones shorter; ov. I-celled with 3 or 4 
ovules ; pod 1-seeded, seed without endosperm. 


C. zeylanica, Benth. in Trans. Linn. Soc. xxv. 314 (1865). 
Detarium zeylanicum, Thw. Enum. 414. C. P. 3714. 
Fl. B. Ind. ii. 271. Bedd. FI. Sylv. t. 190 (poor). 


A large tree; |.-rachis 14-2 in., glabrous, lflts. 2-6, alternate, 
usually with a terminal one but sometimes ending with a pair, 
on short, thick stalks, 2-5 in., oval or ovate-oval, unequal and 
rounded at base, shortly and obtusely acuminate, glabrous, 
rather thick, glaucous beneath, minutely reticulate; fl. small, 
on short ped., arranged in terminal spicate racemes with a 
stout rachis 2—5 in. long; cal.-segm. roundish, obtuse, persistent; 
pet. 0; ov. oblong, tomentose; pod (very immature) very 
shortly stalked, oblong, compressed, slightly falcate, apiculate, 
densely tomentose. 


Moist low country; very rare. I have only seen the C. P. specimens 
collected at Galpata, near Kalutara, in 1861 and 1863. FI. January. 

Endemic. 

The material is insufficient for a full description, or even for certainty 
as tothe genus. I have followed Thwaites in points not shown in the 
specimens. He describes 3 or 4 ovules in the ov.; hence Bentham has 
transferred the plant from Detarzwm to Crudza, the fruit being unknown 
to both. 

In Jan., 1891, I obtained from the Prov. of Uva, under the name of 
*Okuru’ (which properly belongs to Clezdion javanicum), some ripe pods 
which are unknown to me, which may perhaps be the fruit of the 
present species. They are 1-1} in. long, and nearly as much in diam., 
not compressed, slightly curved along dorsal suture, very gibbous and 
rounded below, very shortly apiculate, pericarp thin, brittle-leathery, 
rough but glabrous outside, yellowish-grey, indehiscent with a tendency 
to split along dorsal suture; seed large, solitary, with thick plano-convex 
cotyledons, without endosperm. Leaves, said to be from the same tree, 
are those of the plant here given as Cynometra ramiflora (type). Further 
investigation is needed. 


55 SARACA, Z. 


Small tree, 1. abruptly pinnate, fl. in dense terminal or 
axillary panicles; cal. petaloid, with a long tube and 4 
PART II. I 


Ie al Leguminose@. [Saraca.. 


spreading segm., pet.o; stam. 7 (rarely 8, the uppermost being 
nearly always suppressed or combined with ov.) inserted on a 
fleshy lobed ring at mouth of cal.-tube; ov. stalked, occupying 
the position of the uppermost stam., pubescent, style short ;. 
pod large, flat, dehiscent, 2- or 3-seeded; seed without endo- 
sperm.—Sp. 6; 4 in FZ. B. Lund. 


S. indica, Z. Mant. 98 (1767). Diya-ratmal, Diya-ratam- 
bala, S. 

Jonesia pinnata, Willd., Moon Cat. 30. /onesta* Asoka,+ Roxb.,. 
Dhwaksnums O75) (EePro53 

HE Ba indsii 2705  Wicht. lest, 200) es bote Vidcat ore: 


A rather small, much-branched spreading tree with 
reddish-grey cracked bark; |. large, spreading horizontally, 
rachis 6—9 in., stout, much thickened, corky and flexible at 
base, stip. large 4 in., intrapetiolar, ovate, stiff, brown, deciduous, 
Iflts. 8-12 (4-6 pair) distant, the lowest pair close to base, on 
short, very stout, flexible stalks, 4-9 in. (the middle ones. 
longest) by 14-14 in. wide, linear, tapering to very acute apex, 
perfectly glabrous, rather stiff, venation conspicuous beneath ; 
fl. numerous, about I in. diam., ped. $ in., bracts 2, at summit 
of ped., small, ovate, looking like a cal., panicles large, sessile,. 
much-branched, corymbose, dense, axillary and terminal; cal.- 
tube #-1 in., solid at lower part, segm. 4in., oval, obtuse 3. 
stam. with very long fil, much exserted, spreading; style 
strongly curved into a ring; pod large, 5-9 in. by nearly 2 in. 
wide, tapering to both ends, much compressed, rigidly leathery, 
glabrous, veiny; seed 13-1? in., ovoid, slightly compressed. 

By streams in forests in the low country, especially in the drier 
districts; rather common. Fl. Jan—March, &c.; pale orange-scarlet, 
filaments purple, anth. black. 

Also in India and Malaya. 

The flowers are very sweet-scented, especially in the evening; they 
are at first yellow, and become orange, and finally red, and are profusely 
produced for many months. The young leaves are pendulous and 
coloured. The plant is much cultivated for its beauty and sweet scent. 
Wood light, pale red, rather soit. 


Tamarindus indica, L., Styambala, S., Pulz, T.; the Tamarind is a 
very commonly planted tree, especially in the dry districts. There are 
specimens in Hermann’s Herb., but it is nowhere wild here. In the dry 
region of the island it is very conspicuous, every village having several. 
and often magnificent specimens. Trees are occasionally found appa- 
rently wild in the jungle, but always on the site of abandoned villages. 


* Named in honour of Sir William Jones, the Oriental scholar and. 
botanist, who first fully described this tree. 
+ Asoka is the Sanskrit name. 


Bauhinia. Leguminose. T15 


56. HUMBOLDTIA,* Vai/. 


Small tree, |. abruptly pinnate with very large double stip., 
fl. in erect axillary racemes; cal. with short tube and 5 
petaloid segm.; pet. 5, clawed ; stam. normally 5, often 3 or 4, 
inserted at mouth of cal.-tube and alternating with 5 small 
staminodes, anth. versatile; ov. shortly stalked, style very 
long; pod flat, dehiscent ; seed without endosperm.—Sp. 5 ; 
4 in FZ. B. Ind. 


H. laurifolia, Vai, Sym. Bot. iii. 106 (1797). Gal-karanda, S. 

Moon Cat. 17. Thw. Enum.97. C. P. 328. 

Fl. B. Ind. ii. 273. Vahl, Symb. i. t. 56. Wight, Ic. t. 1605. 

A small tree or a shrub with numerous horizontally 
spreading or rather drooping branches, the young ones with 
swollen hollow internodes; |. numerous, spreading distichously, 
‘rachis 4-6 in., cylindrical, glabrous, thickened and flexible at 
base, stip. apparently double, really peltate with two separate 
halves, the longer erect, 14 in., ovate, obtuse, stiff, the smaller 
spreading horizontally, broad, auricled towards petiole, very 
acute at other end, rigid, lflts. 8 or 10 (4 or 5 pair), very 
shortly stalked, drooping, 24-34 in., ovate, slightly unequal at 
base, caudate-acuminate, undulate, glabrous, rather stiff; f1. 
about #in., on short finely pubescent ped., arranged in shortly 
stalked, rigidly erect, axillary racemes about 3-5 in. long, 
bracts at base of ped. rounded, green, persistent, those at 
summit oval, pink, enclosing the bud; cal.-segm. oval, obtuse, 
recurved, thin; pet. clawed, spreading; stam. erect, twice as 
long as pet.; ov. pilose; pod 3-4 in. by 14 in. wide, scarcely 
stalked, pointed, much compressed, glabrous, veiny. 

Damp places in forests in the moist low country; common. FI. Feb.; 
white, cal. pink; honey-scented. i 

Also in Malabar. 

The hollow, swollen internodes of the branchlets are inhabited by 
small ants, which gain access by a little perforation situated between the 
stipules of the leaf above. 

The racemes are stiff and quite erect as figured by Vahl, not drooping 
as stated in FI. B. Ind. 


7. BAUHINIA, /. 


Trees or woody climbers, 1. apparently simple, usually 
bifid at top, fl. in racemes; cal. either with a short tube and large 
limb which splits down one side in a spathe-like manner or 


* Named in honour of F. A. von Humboldt, author of ‘ Flora Fribur- 
gensis. Vahl first named the plant Batschza (1. c. 39). 


116 Leguminosae. [Bauhinia. 


campanulate with 5 segm., pet. 5; stam. 10 (3 in B. anguzna), 
distinct; ov. stalked; pod dehiscent or indehiscent, linear, 
many-seeded ; seed with endosperm.—Sp. 130; 36 in FZ. B. 
Lhd. 


The simple bilobed leaf is obviously a pair of connate Ifits. 
Trees or shrubs. 


Fl. large, bright yellow . : 5 ‘ : . I. B. TOMENTOSA. 
Fl. small, nearly white . : : c c . 2. B. RACEMOSA. 
Woody climber; fl. minute . A é : . 3. B. ANGUINA. 


1. B. tomentosa, L. Sf. P/. 375 (1753). Petan, Kaha-petan, 
Tiruvatti, 7. 


’ Burm. Thes. 44. Fl. Zeyl.n.147. Moon Cat. 33. Thw. Enum. 98. 
CAE TAGS: 


Fl. B. Ind. ii. 275. Burm. Thes. t. 18 (poor); Bot. Mag. t. 5560. 


A large erect bush with slender branches, bark yellowish- 
erey longitudinally furrowed; 1. 14-3 in., conduplicate, deflexed 
on petiole, divided more than 4 down into 2 oval rounded 
lobes with a setaceous mucro between, truncate at base, 
glabrous above, densely pubescent and whitish beneath, 7- 
nerved from the base, petiole $—-? in., thickened at both ends; 
fl. large, nodding, over 2 in., ped short, stout, racemes small, 
terminal, bracts linear; cal. about 1 in., bifid at top, pubescent, 
pet. very much imbricated, broad, obovate, rounded, very 
shortly clawed; style long, stigma terminal, peltate, pod 
4-6 in. by 2 in. wide, shortly stalked, narrowed to base, pointed, 
finely pubescent, 8—12-seeded, dehiscent; seed 4 in., ovoid, 
shining. 

Dry region; very common. FI. Jan., Feb., August; sulphur-yellow, 
the upper pet. with a dark-purple blotch at base. 

Throughout India and Malaya, and in China and Tropical Africa. 

Heart-wood hard, very tough, shining, dark red, moderately heavy, 
an excellent wood, but too small to be of much value. The colouring of 
the fl. is precisely that of several species of Azézscus,; the petals never 


spread, but remain imbricated, forming a deep bell-shaped corolla, which 
is very conspicuous. 


[B. acuminata, L., is given for Ceylon in Fl. B. Ind. It is, however, 
only known here in gardens, where it is an old introduction; there are 


specimens in Hermann’s Herb., and he gives (Mus. 8) the native name 
‘ Mayila’ for them, which rightly belongs to the next. | 


2. B. racemosa, Lam. Ency. Meth. i. 390 (1783), (non Vahl). 
Mayila, S. Atti, 7. 

B. parviflora, Vahl, Moon Cat. 33. Piliostigma racemosum, Benth., 
Thw. Enum. 98. C. P, 1497. 

Hy Binds 7 6s bedd Hla Sylvat 132) 


A small tree with a thick knotted trunk, much branched, 
bark blackish, very rough and much furrowed; 1: small, con- 


Bauhinia.| Leguminose. Ly, 


duplicate, 11-14 in., broader than long, truncate at base, cut 
scarcely 4 down into 2 broadly oval rounded lobes, glabrous 
on both sides, whitish and 9-nerved from the base beneath, 
with fine reticulate venation, rigid, petiole 4—-}in.; fl. small, 
under 1in. ped. shorter than cal, in erect lax racemes 
terminating the new growth, buds pointed ; cal. reflexed 2 or 
3-toothed at end; pet. distant, small, strap-shaped, acute, 
spreading ; stam. a little longer than pet.; style none; pod 
shortly stalked, very long, 8-10 in. by ?in. wide, pendulous, 
tapering to base, blunt at apex, somewhat falcate, leathery, 
indehiscent, glabrous, 12—20-seeded. 


Dry region; very common. FI. April, June-September; yellowish 
white. 


India, Malaya, and China. 
Wood rather heavy, hard, pale brownish-grey; attains a fair size. Very 
strong fibre is obtained from the inner bark, used for rope-making. 


[B. purpurea, L.,is commonly grown in gardens. There are specimens 
in Hb. Kew, ‘Peradeniya, Macrae;’ hence it is given for Ceylon in 
Fl. B. Ind., but it has no claim to nativity here. | 


3. *B. anguina, Roxb. Cor. Pi. iii. 82 (1819). 

Trimen in Journ. Bot. xxiii. 144. 

Fl. B. Ind. ii. 284. Hort. Mal. viii. tt. 29-31. Roxb. Cor. Pl. t. 285. 

A very wide-spreading woody creeper, with the older stems 
singularly compressed, and strongly curved in alternate direc- 
tions between each node, so as to be serpentine, young shoots 
glabrous with short lateral twigs, each one provided at about 
14 in. from the base with a pair of opposite, stiff, flattened, 
circinately curved, prehensile tendrils pubescent on upper 
surface ; 1. numerous, rather large 5-6 in., ovate, more or less 
cordate at base, acuminate, quite entire or more or less bifid 
or bilobed at apex, glabrous and shining, dark green, petiole 
1-3 in., stiff, slender, stip. very small, linear, obtuse, reflexed, 
soon falling; fl. and pod not seen (fl. minute in copiously 
panicled racemes; stam. 3; pod 14-2 in., oblong, indehiscent, 
glabrous). 

Low country in the dry and intermediate regions; very rare, and 
probably merely introduced. Foot of Doluwe Kande, Kurunegala Dis- 
trict; near Dambulla (Bower), 

Native of India and Malaya. 

I have seen no flowers. This great creeper has been long cultivated 
in the Gardens, and is included in Moon’s Catalogue, 1824. Asthe stems 


readily root at the joints, it is easily propagated, and freely establishes 
itself. 


118 Leguminose. [Neptunia. 


III. Mimose. 


Stam. Io. 
Aquatic herb. : ; : : : . 58. NEPTUNIA. 
Woody climber . : : ; 5 ‘ . 59. ENTADA. 
Trees or shrubs 
Not spinous . : : : . 60. ADENANTHERA. 
With spinous branchlets : : : . 61. DICHROSTACHYS. 
Stam. indefinite 
Stam. free, = i : 5 .) 62: 7 ACAGIN 
Stam. monadelphous below 
Pod straight, flat . : ; 63. ALBIZZIRe 
Pod curved into a circle, often twisted . 64. PITHECOLOBIUM. 


All our 22 species, with the exception of PieoIaHD subcoriaceum, 
are plants of the low country, but P. dzgemznum also extends up into the 
lower montane zone. Veptunza, Dichrostachys, and 7 species of Acacia 
are restricted to the dry region, and Adenanthera bicolor, Albizzia 
stipulata, and Pithecolobium bigeminum are confined to the moist. 
P. geminatum is the only endemic species. 


58. NEPTUNTA, Lour. 


Aquatic annual with floating stems, |. abruptly bipinnate ; 
fl. very small, in heads, the lower ones barren; cal. cam- 
panulate, 5-toothed ; pet. 5, connate near base; stam. 10 with 
a gland above the anth. when young, staminodes in barren fl. 
petaloid ; ov. stalked, absent in barren fl.; pod small, dry, early 
dehiscent along dorsal suture, much flattened, with several 
seeds.—Sp. 8; 2 in FZ B. Ind. 


N. oleracea, Lour. Fl. Cochinch. p. 654 (1790). Diya-nidi- 
kumba, 5S. 


Thw. Enum. 99. C. P. 1516. 

Fl. B. Ind. ii. 285. Roxb. Cor. Pl. t. 119 (AZémosa natans). 

Annual, floating in water, stem elongated, scarcely 
branched, with numerous plumose roots pendulous from the 
nodes, stout, often swollen, purple, often provided on the out- 
side with floats of white spongy tissue; |. bipinnate, main 
rachis 13-2 in. at right angles with stem, glabrous, stip. 
membranous, ovate, acute, persistent, pinne 2 or 3 pair, op- 
posite, ee stalked, ?-1 in. long, Iflts. 18-28 (9-14 pair), 
sessile, ;-gin., linear, obtuse, glabrous, glaucous beneath; fl. 
sessile in an oblong head 4-1 in. long, at end of stout, erect, 
tapering, glabrous, axillary peduncle 3-5 in. long, the barren 
ones in lower part of head; pet. strap-shaped ; stam. much 
exserted, staminodes longer, g in., strap-shaped, acute; pod 
long-stalked, deflexed, I “in. by 2 2 in. wide, broadly oblong, 
beaked, clabrous, 4-6- ‘seeded. 


Entada.] Leguminose. 119 


Shallow pools and stagnant water in the dry region; rather common. 
Fl. all the year; yellow. 

Found throughout the Tropics. 

The plant is completely aquatic; its stems float by means of white 
spongy masses developed on the outside of the cortex. 


lV. plena, Benth., a Tropical American species, is said in F1. B. Ind. to 
be introduced to Ceylon. There are specimens in Hb. Kew labelled 
“Ceylon, Walker n. 1417.’ 


59. ENTADA, Adazns. 


A very large woody climber, |. bipinnate, ending in a 
tendril, fl. very small, polygamous (male and bisexual) in long 
narrow spikes; cal. campanulate with minute teeth ; pet. 5, 
equal, valvate, slightly connate at base, stam. I0, free, twice 
as long as pet., anth. with globular gland on top; pod very 
large, woody, composed of indehiscent joints; seeds very 
large, flat—Sp. 10; 1 in FZ. B. Ind. 


eer ondene, Benth. in Hook. Journ. Bot. iv. 332 (1842). Yus- 
wel, S. 
Henn. Mus. -44. Burm. Thes. 139. FI. Zeyl. n. 219. Mzmosa 
Entada, L. Sp. Pl. 518. Acacia scandens, W., Moon Cat. 73, Thw. Enum. 
98. C. P. 2661. 

Fl. B. Ind. ii. 287. Rheede, Hort. Mal. ix. t. 77, and viii. tt. 32-4. 

An immense woody climber, with a thick trunk and long 
twisted snakelike branches; |. large, main rachis 6-10 in., 
stiff and semi-woody ending in a bifid spiral tendril by which 
the plant climbs, pinnz 2 or 3 pair with stiff glabrous rachis 
3 or 4 in. long and much thickened at base, Iflts. 6 or 8 (3 or 
4 pair), 14-2 in., obovate-oblong, acute at base, rounded, often 
emarginate at apex, glabrous, paler beneath, with the lat. 
veins conspicuous; fl. sessile, crowded in long, narrow, pe- 
dunculate axillary spikes 6-10 in. long; pod 14-2 ft. long, by 
24-3 in. wide, curved, often twisted, compressed, woody, in- 
dented on both sutures between the seeds, joints 6-12, turgid, 
1-seeded, indehiscent, glabrous, brown; seed very large, 
17-2 ‘n., circular, compressed, smooth and shining, brown, 
testa thick and woody. 

_ country up to 2000 ft.; common. FI. Feb.-April; greenish- 

white. 

Throughout the Tropics. 

The joints composing the immense pod break away from one another, 
and also from the thickened sutures which remain as an empty frame. 

The juice of the wood and bark is used as an external application to 
ulcers; also, I am told, to poison fish. 


120 Leguminose. [Adenanthera. 


60. ADENANTHERA, LZ. 


Trees, not spinous, |. bipinnate, without stip., lflts. alternate, 
fl. very small in axillary spicate racemes; cal. campanulate,. 
shallowly toothed; pet. 5, equal, valvate, very slightly coherent 
at base; stam. 10, free, equalling pet., anth. with an ovoid. 
gland on summit; pod narrow, falcate or twisted, thin, torulose,. 
with several seeds.—Sp. 4; 2 in FZ. B. Ind. 


Lfits. rounded; seed all red . ; : . I. A. PAVONINA.. 
Lfits. acuminate; seed half red, half black : F . 2. A. BICOLOR. 


1. A. pavonina, LZ. SZ. Pi. 384 (1753). Madatiya, S. Anai- 
kuntumani, 7. 


Herm. Mus. 21. Burm. Thes. 79. Fl. Zeyl. n. 160. Moon Cat. 34. 
Thw. Enum. 98. C. P. 3605. 


FI. B. Ind. ii. 287. Bedd. Fl. Sylv. t. 46. Wight, Ill. t. 80. 

A tall tree with erect trunk, and pale grey rather smooth: 
bark, young parts quite glabrous; |. large, wide-spreading, 
main rachis 8-12 in., glabrous, furrowed above, striate, pinne 
3 or 4 pair with rachis horizontally spreading and thickened at 
base, lfits. 8-12, alternate, stalked, distant, 14 in. or more, 
oval-oblong, rounded at both ends, unequal at base, glabrous, 
thin, dark green, paler beneath; fil. on short horizontal 
elabrous ped., racemes stalked, lax, shorter than |., buds blunt, 
bracts minute, very early deciduous; cal. glabrous; pet. linear- 
lanceolate, acute; pod 6-8 in. by nearly ? in., tapering to base, 
pointed, falcately curved, glabrous, valves becoming spirally 
twisted after dehiscence; seeds 8-10, 2 in., globose-lenticular 
with blunt keel, brilliant scarlet, shining. 

Da country; common, but generally planted. Fl. April; Re ioe. 
white. 

Also in India, Malaya, Philippines, and China. 

The bright scarlet seeds are well known; they are called ‘ Mad-éta,’ 


and are the principal weights used by gold and silversmiths, weighing 
each somewhat less than 4 grains. 


Heart-wood hard, moderately heavy, close-grained, pale brownish-red. 


2. A. bicolor, Joon Cat. Ceyl. Pl. 34 (1824). Meas-mora, S. 
[PLATE XXXIV.] 


Thw. Enum. 98. C. P. 324. 

Be liad 2a 77 

A moderate-sized tree with wide-spreading branches, 
young shoots glabrous, entirely of a dull vinous-purple 
colour; |. numerous, spreading, main rachis 6-8 in., glabrous, 
shining, purplish, scarcely channelled, pinnze 3 or 4 pair or 
often alternate with slender purple rachis thickened at base, 
Iflts. 5-7, alternate, very distant, shortly stalked, disarticulat- 


Dichrostachys.] Leguminose. 12f 


ing, It-I{ in., broadly oval, acute at base, shortly and bluntly 
acuminate, glabrous, paler beneath; fl. as in the last; pet. 
lanceolate; pod 3-5 in. by about 2 wide, usually much twisted; 
seeds 6-10, 3-4 in., shining, the basal half red, the upper half 
black. 


Moist low country; rather rare. Kalutara (Moon); Madulla; Mapala- 
gama. Fl. December, April; greenish-pink. 

Also found at Malacca. 

Moon quotes for this Rumph. Herb. Amboin. iii. t. 212, a doubtful: 
plant not yet clearly identified. 


61. DICHROSTACHYS, VW. and A. 


A bush with spinous branchlets, |. abruptly bipinnate, f1. 
very small, in drooping spikes, the lower ones barren; cal. 
campanulate, segm. short, triangular, pet. connate for ? length 
from base; stam. 10, free, represented in barren fl. by long 
staminodes, anth. with a stalked gland on summit, pod linear,. 
flat, indehiscent, somewhat jointed, much twisted up spirally 
when ripe, 6-10-seeded.—Sp. about 5; 1 in FZ. B. Ld. 


(aaa W. and A. Prod. 271 (1834). Andara, S. Vidat- 
ta : 

Herm. Mus. 24. Burm. Thes. 3. FI. Zeyl.n. 215. MJmosa cinerea, 
L. Sp. Pl. 520. Desmanthus cinereus, Willd., Moon Cat. 73.. Thw. 
Enum. 99. C. P. 1523. 

Fl. B. Ind. ii. 288. Burm. Thes. t. 2. Bedd. Fl. Sylv. t. 184. Wight,. 
fc 6357- 

A shrub or small tree with numerous divaricate branches, 
bark yellowish, fibrous, vertically furrowed, ultimate twigs. 
sharply spinous at end, many reduced to short, woody, axil- 
lary spines, bark yellowish-grey, young parts pubescent; 1. 
small, mostly from the suppressed branchlets, rachis 1-14 in., 
slender, hairy, with a small erect gland between the bases of 
each pair of pinne, pinne 8-16 pair, 4-2 in., sessile, hairy, 
Ifits. minute, numerous 24-40 (12-20 pair), closely placed, 
sessile, linear, oblique, subacute; fl. numerous, sessile, crowded 
in dense spikes about 1 in. long on drooping peduncles 
coming off from the suppressed branchlets below the 1, bracts 
minute setaceous ; cal. pubescent, stam, much exceeding cor., 
staminodes much longer (nearly } in.) filiform; ov. hairy; pod 
2-3 in. by # in. wide, glabrous, dark brown. 

Dry region; very common; and in the most barren places. Fl. Dec.- 
March, Aug., Sept.; barren fl. bright purplish-pink, fertile ones yellow. 

Also in India, Malay Islands, N. Australia, and (?) Trop. Africa. 

The tassel-like, particoloured, drooping flower-spikes are very pretty, 
and a noticeable feature in the stunted jungle in desert places. 


22 Leguuiinose. [ Acacia. 


The heart-wood is dark red, extremely hard and strong, and said to 
‘be the toughest in Ceylon; but it is too small to be of much use. 


Desmanthus virgatus, Willd., C. P. 3606, is a not uncommon weed 
in cultivated ground, but certainly not indigenous, here. It was collected 
by Hermann, “and is Fl. Zeyl. n. 505 (AZzmosa virgata, L.). 


Leucena glauca, Benth., is naturalised in many places in the low 
country. A native “of N orth America. 


Mimosa pudica, L., is a very abundant and most troublesome weed 
throughout the low country. It is said by Gardner to have been intro- 
duced here by Moon, and it is included in his Catalogue (1824), p. 73, 
but it seems to have been here earlier. Its native country appears to be 
Brazil. Like other sensitive plants, it is called ‘ Nidi-kumba’ by the 
Sinhalese. The root is considered an antidote for cobra bite. 


62, ACACIA, Wild. 


Trees or shrubs often scandent by prickles, |. bipinnate, 
often with sharp persistent stipular spines, fl. very small, in 
heads or spikes, often polygamous; cal. campanulate or funnel- 
shaped, segm. more or less triangular ; pet. connate into a cor. 
with 5 lobes, stam. indef. free, much longer than cor., anth. 
not gland-tipped; pod dry, usually compressed, dehiscent 
(rarely indehiscent); seeds several, on long funicles.—Sp. 43 
(mostly Australian); 18 in FZ B. Lnd. 


Trees or shrubs. 
Fl. in globular heads. 
Heads axillary. 


Pod strongly constricted between seeds . . 1. A. ARABICA: 
Pod not constricted. 
Pod cylindrical . : : : ; . 2. A, PLANIFRONS. 
Pod fiat. 
Lfits. 5-8 pair, glabrous 3. A. EBURNEA. 
Lflts. 8-16 pair, tomentose 4. A. TOMENTOSA. 
Heads in panicles 5. A. LEUCOPHLEA. 
F]. in spikes. 
Pinnze 9-18 pair 6. A. SUNDRA. 
Pinnz 4-6 pair 7. A, FERRUGINEA. 
Woody climbers. 
Lfits. 10-22 pair, linear-oblong 8. A. CAHSIA. 
Lflts. 40-50 pair, narrowly linear. 9. A. PENNATA. 


. Several Australian species, especially A. ellie, Link, and varieties 
of A. decurrens, Willd., have been extensively planted in the montane 
region; A. Melanoxylon, Br., is also common about Nuwara Eliya. 

A. Farnesiana, Willd., is quite naturalised in many parts of the low 
country, and is C. P. 1522. It appears to be indigenous to Western Trop. 
_America and Australia, and perhaps to S. Africa. 


1. * A. arabica, Willd. Sp. P/. iv. 1085 (1805). Karuvel, 7. 
Thw. Enum. 415. C. P. 3640. 
FL. B. id= 152635 Bedds Bly Sylv: t: 47: 


Acacia.) Leguminose. 1 


A shrub or small tree, with dark bark and slender terete 
straight branchlets, young parts pubescent; |. small, rachis 
2-24 in., pubescent, stipular spines variable from small and 
inconspicuous to 2 in., slender, spreading white, sometimes 
quite absent, pinnz 3-5 pair, 1-14 in., distant, lflts. 24-30, 
(12-15 pair), 4 in., very shortly stalked, strap-shaped ; fl. in 
dense globular heads, 2-5 together from the axils, peduncles 
4-2 in., pubescent with 2 bractlets above the middle; cor. 
twice as long as cal.; pod 4-6 in., straight, 8—12-seeded, 
strongly constricted at sutures between the seeds, densely 
covered with fine grey down. 

Dry region; very rare. Collected at Trincomalie by Glenie, but 
perhaps only introduced there; Jaffna, apparently wild, but probably 
originally planted. 

Native in N.W. India, Arabia, Egypt, Tropical Africa. This is the 
well-known ‘Babul’ so common in N. India; but Beddome has never 
seen it truly wild in the forests of the Peninsula, and it is unlikely to be 
native to Ceylon. 

On the upper part of the branches bearing the flower-heads the leaves 
are often suppressed. 


2. A. planifrons, W. and A. Prod. 276 (1834). Odai, Udai, 7. 
[PLATE XXXV.] 

Trimen in Journ. Bot. xxiii. 144. 

Fl. B. Ind. ii. 293. 

A small tree, usually with an erect stem and an umbrella- 
like head, branches spreading ina plane (like those of Cedar), 
bark very thick, dark grey, rather smooth, on the young 
branchlets purplish, stipular spines of two kinds mixed, either 
very short and curved downwards, or 14-2 in. long, straight, 
very divaricate, slender, scarcely tapering, white with a brown 
polished point; 1. very small, in fascicles from arrested 
branchlets, rachis about 1 in., very slender, flattened above, 
slightly hairy, pinne 3 or 4 pair, closely placed near end of 
rachis, 4-2 in. long, Iflts. very minute, 10-20 (5—Io pair) strap- 
shaped, obtuse; fl. minute, sessile in dense globular heads 
Zin. diam., peduncles 4—? in. very slender, with the bracts 
below the middle, coming off with the 1. from the arrested 
branchlets ; pod about 2 in. by } in. wide, cylindrical, turgid, 
acute, curved nearly into a ring, glabrous. 

Dry region; very rare. Confined to the island of Mannar (and a 
very small part of the mainland opposite), where it is extremely abundant. 
Fl, Oct.-March ; pale creamy-white. 


Also in S. India and in Ramisseram Island. j 
This is the ‘Umbrella Tree’ of the English ;* and very characteristic 


* The Rev. J. Cordiner, who visited Mannar in 1804, gives an account 
of the ‘ Umbrella Tree’ in his ‘ Ceylon,’ vol. ii. pp. 8 and 32. 


124 Leguminosae. [ Acacia. 


of Mannar, where it composes almost the whole forest, and from the 
horizontal arrangement of the branches gives it a very peculiar aspect. 
The trees, when covered in Feb. with sheets of white blossom, look like 
old hawthorn trees. It is cultivated in a few Colombo gardens for orna- 
ment, and has occasionally appeared on the shore there, no doubt brought. 
with ballast from Tuticorin, where it is also abundant. 

The wood is very hard and heavy, pale yellow, and when fresh cut has 
a faint alliaceous smell. It is much used for firewood, and there is a 
large export from Mannar to S. India (Tuticorin and Negapatam): 
amounting in 1890 to 3,446,422 logs, value Rs. 68,243 (as I am informed 
by Mr. M. S. Crawford). 


The pods are largely used as food for sheep, goats, and cattle in the 
dry season. 


The description under this name in F1. B. Ind. is partly made up from 
some other species. 


3. A. eburnea, Willd. Sp. Pi. iv. 1081 (1805). Udai-vél, Kal- 
udai, 7. 

Ghw. Enum. 99. “G. P1521. 

Fl. B. Ind. ii. 293. Roxb. Cor. PI. t. 199 (not good). 


A large bush or small tree, much branched, bark of young 
branches purplish-brown, smooth, stipular thorns always 
straight, varying in size from } to 2} in. gradually tapering 
from a broad base toa very sharp point, ivory-white, polished ; 
I. small, arranged as in the last, pinne 2-7 pair, 4-2 in, 
glabrous, lflts. minute 10-16 (5-8 pair) oblong, obtuse; fl. as. 
in the last, heads } in. diam., bracts about middle of peduncle ; 
pod 3-4 in. by 2 in. wide, shortly stalked, compressed, more 
or less falcately curved, pointed, glabrous, veiny, 6—9-seeded, 
slightly compressed between the seeds. 

Dry and desert regions, especially near the coast; rather common. 
Trincomalie; Hambantota District, abundant; Jaffna, and the whole west 
coast to Puttalam. Fl. Dec.-Feb.; bright dark yellow. 

Also in India, Affghanistan, and Arabia. 

Cordiner (‘ Ceylon,’ ii. 5) noticed this in 1804 on the coast north of 


Aripo, and calls it the ‘ Cockspur Thorn,’ a very appropriate name. 
The wood is small but very hard, yellowish-white. 


4. A. tomentosa, Willd. Sf. Pi. iv. 1087 (1805). Anaimulli, 7. 

Thw. Enum. 99. _ C. P. 744. 

Fl. B. Ind. ii. 294. 

A small bushy tree, bark yellowish-grey with large lenticels, 
purplish on twigs, young parts tomentose, stipular thorns 
straight, broad at base and rapidly tapering to sharp point, 
varying in size, the larger ones reaching 34 in. and over $ in. 
wide at base, inflated, pubescent when young, afterwards 
glabrous, grey becoming black, very sharply pointed; 1. longer 
than in last, rachis 14-33 in., flat, very pubescent, pinnze 6-10 
pair, 1-1} in., the lower ones shortest, lflts. 16-32 (8-16 pair), 


Acacia. Leguminosae. 125 


opposite, sessile, }-lin., broadly oblong, tomentose-pubescent 
on both sides; fl.-heads globular, over } in. diam., peduncles 
about Iin., pubescent with the bracts a little above the 
middle; pod 4-5 in. by $ in. wide, compressed, strongly 
falcate, bluntly pointed, 5—8-seeded, somewhat torulose, at first 
pubescent and grey afterwards glabrous and reddish; seed 
2 in., ovoid, compressed, olive green with a ring of paler colour 
on either face. 

Dry region; rare. Kalpitiya (Gardner) ; Jaffna; Ilupaikadavai, Mannar 
Dist. Fl. Feb—April; pure white (not ‘purple’ as given in Fl. B. Ind.). 

Also in Bengal, Java, and perhaps Southern India. 

The enormous black or greyish-brown thorns, whence its name of 


‘Elephant Thorn,’ are very characteristic. Tennent (‘ Ceylon,’ i. 108) says 
they are called the ‘Jungle Nail’ by Europeans. 


5. A. leucophlea, Willd. Sp. Pi. iv. 1083 (1805). Maha- 
andara, Katu-andara, S. Velvél, 7. 

Thw. Enum. 99. C. P. 1520. 

Fl. B. Ind. ii. 294. Bedd. FI. Sylv. t. 48. 

A large tree, bark thick, pale yellowish-white, branches 
dense, spreading, young branches pubescent, stipular thorns 
small, straight; 1. numerous, closely placed, rachis’ 24-3 in., 
densely pubescent, with a small red sessile gland between 
bases of each pair of pinnz, pinnz 5-15 pair, lflts. very small, 
sessile, crowded 30—40 (15-20 pair), }-+in., linear-strap-shaped, 
subacute, Bore or slightly hairy; fl.-heads numerous, 
globular, small, }—3 in. diam., peduncle 2in., woolly, the bracts 
about the middle, in large axillary tomentose panicles with 
slender branches; pod 3-5 in. by } in. wide, linear, very slightly 
curved, flexible, pendulous, subdehiscent, finely but densely 
tomentose, seeds numerous, 

Dry region; common. FI. Aug.—Sept.; bright pale yellow. 

Also in India, Burma, Malay Islands. 

The thick yellowish-white bark is known as ‘Velam’ bark, and is 
exported to India for the arrack distillers. It is used in Ceylon for dyeing 
sails, mats, &c., and for tanning, and also affords a coarse fibre. The 
heart-wood is very hard, heavy, and brownish-red, tough, and strong. 


6. A. Sundra,* DC. Prod. 458 (1825). Rat-Kihiri, S. Kodali- 
murunkai, 7. 

A. Catechu, Thw. Enum. 99 (non Willd.). C. P. 3547. 

Fl. B. Ind. ii. 295. Bedd. Fl. Sylv. t. 50. 

A small or middle-sized tree, bark dark brown, young 
shoots purple, quite glabrous; stipular thorns small, decurved, 
hooked, black, often absent; l.-rachis 4-5 in., tapering, 
elabrous, with a large sessile gland betwen the bases of each 


% ities or Chundra is the Telegu name. 


126 Leguminose. [ Acacia. 


pair of pinne, slightly channelled above, often with recurved 
prickles beneath, pinnz 9-18 pair, 1-14 in,, lflts. 40-60 (20-30: 
pair), small, tin., sessile, crowded, easily falling, linear, obtuse, 
quite glabrous; fl. small, sessile, in rather lax, shortly stalked, 
erect, axillary spikes 3-4 in. long; cal. quite glabrous; cor. 
three times as long as cal., lobes linear-lanceolate; pod 

4-5 in. by 2 wide, stalked, nearly straight, tapering to each 
end, very thin, margin often indented between the seeds, quite 
glabrous, veiny, dark brown; seeds 4-6, broadly oval, much 
compressed, pale greenish-brown. 

Dry region; rare. Bintenne county below Balangoda, 1857 (Thwaites) ;. 
Northern Prov. (Vincent); between Mannar and Vavuniya (Capt. 
Walker). The late H. Meade recorded it from Patchilapillai, near Jaffna, 
in 1859. Fl. September; pale yellow or pinkish. 

In Peninsular India and Burma ; also in Trop. Africa (?). 

The heart-wood is red and extremely hard and heavy, whence the T.. 
name (‘axe-breaker’). It is also bitter, and the Sinhalese make drinking- 
cups from it. 

This is so closely allied to A. Catechu, Willd., as to be combined with 
it by many botanists. That tree, however, has more numerous pinne 
and Ifits. and the calyx is tomentose. It appears to be confined to 
Northern India and Burma, and is figured in Roxb. Cor. Pl. t. 175 and 
Bentl. and Trim. Med. Pl. ii. t. 95. From the wood of A. Catechu is 
prepared the astringent extract called Cutch in India, but I have no 
knowledge of anything of the kind being obtained from our Ceylon tree, 
though it probably could be. 

A. Suma, Kurz., figured (as A. Catechu) in Bedd. FI. Sylv. t. 49, is 
given as a Ceylon species in Fl. B. Ind. ii. 294. The specimen in Herb. 
Kew is from Macrae, and was in all probability gathered from the old 
trees of this species in Peradeniya Gardens. But it may perhaps occur 
in the Northern Prov., and is easily known by its white papery bark. It 
also affords Cutch. 


7. A. ferruginea, DC. Prod. ii. 458 (1825). 

Trimen in Journ. Bot. xxiii. 145. 

Fl. B. Ind. ii. 295 (not given for Ceylon). Bedd. Fl. Sylv. t. 51. 

A moderate-sized tree, bark rust-coloured, young parts. 
glabrous, stipular thorns small, straight, often absent; |.-rachis 
2-34 in., glabrous, with a sessile gland midway between base 
and lowest pair of pinnze and another between the bases of 
uppermost pair, pinne 4-6 pair, rather distant, 14-2 in., lfits. 
30-40 (15-20 pair) very shortly stalked, 4-1 in., linear-oblong, 
unequal-sided, obtuse, glaucous-green paler beneath, rigid ; f1. 
sessile, in slender axillary erect spikes 3-4 in. long; cal. 
elabrous, segm. triangular, acute; cor. twice as long as cal., 
lobes lanceolate; pod (not seen) much as in the last. 

Dry region; very rare. At present collected only at Anemaduwe in 
the North-west Province between Puttalam and Kurunegala. Fl. Aug.; 
yellowish. 

Also in Southern India. 


Acacia. Leguminosae. ay 


8. A. czesia, Willd. Sp. Pi. iv. 1090 (1805). Hinguru, 5S. 
Herm. Mus. 34. Burm. Thes. 3. Fl. Zeyl.n. 217. Mimosa cesta, 
L. Sp. Pl. 522. Moon Cat. 73. A. Jnésza, Willd., Thw. Enum. 99. C. P 


Fl. B. Ind. ii. 297 (A. Jntsia). Pluk. Phyt. t. 330, f. 3. Rheede, 
Hort. Mal. vi. t. 4. 
A large woody climber, stems attaining ? in. diam., slightly 
‘rough but not pubescent, greyish or yellowish, copiously set 
with small, very sharp, hooked, decurved prickles, young parts 
minutely pilose; 1. numerous, rachis 5-6 in., swollen at base, 
nearly glabrous when mature, slightly channelled above, with 
a large sessile gland near base and smaller ones just below 
the bases of one or two uppermost pairs of pinne, the under 
side set with decurved hooked prickles, stip. cordate, acute, 
early caducous, pinne 5-8 pair, with a flattened rachis 2-3 in. 
long, lfits. 20-40 (10-20 pair), nearly sessile, crowded and 
slightly overlapping, 3-3, very unequal-sided, the midrib near 
the upper margin, linear-oblong, truncate at base, abruptly 
acute or mucronate, glabrous on both sides; fl.-heads globular, 
very numerous, small 2in., on finely woolly peduncles 4-2 in. 
long and 2-4 at a node, arranged in large pubescent pyramidal 
terminal panicles, bracts linear, inconspicuous ; cal. funnel- 
shaped, pubescent, lobes acute; pod large, about 6 in. by 14 in. 
wide with a short stalk, bluntly pointed, straight, flat and 
thin, glabrous, slightly veined, 6—-10-seeded, dehiscent. 
Low country; common. FI. Sept.; yellow. 
Also in India, Malaya, and Philippines. 


A variable plant, horribly prickly; the Iflts. soon fold together after 
gathering. A. /ntsia cannot be separated from A. c@sza even as a variety. 


[A. concinna, DC., is included in Thw. Enum. 99, and following this 
is given for Ceylon in FI. B. Ind. ii. 297 and Trim. Syst. Cat. 30. But all 
the specimens of C. P. 1519 that I have seen are referable to A. ce@sia, 
with which Thwaites clearly confounded it. There are no Ceylon speci- 
mens in Herb. Kew or Brit. Mus.] 


g. A. pennata, Willd. Sp. P/.iv. 1090 (1805). Goda-hinguru, S. 

Herm. Mus. 30. Burm. Thes. 2. Fl. Zeyl.n. 216. Mimosa pennata, 
L. Sp. Pl. 522. Moon Cat. 73. Thw. Enum. 99. C. P. 3300. 

Fl. B. Ind. ii. 297. Burm. Thes. t. i. (good). 

A stout woody climber, stems much as in the last, but with 
smaller and straighter prickles (often quite absent), young 
branchlets pubescent ; l|.-rachis 4-6 in. densely pubescent, 
with a large sessile gland above near its base, and also 
between the base of the upper 2 or 3 pair of pinne, under 
surface with recurved prickles, pinne 10-18 pair, 14-14 in., 
usually curved, lflts. very numerous, 80-100 (40-50 pair) very 
closely placed, slightly overlapping, very small, 4 in., narrowly 


128 Leguminose. [Albizzia. 


linear, somewhat falcate, mucronate, very unequal-sided, the 
midrib close to the upper edge, glabrous or Sen hairy ; 
fi-heads globular, ¢—} in. diam.,-on pubescent ped. # in. long, 
2-4 together in axils of 1 or bracts, the whole (oem a long 
racemose panicle; cal. glabrous ; pod 44—5 in. by Z in. wide, 
stalked, apiculate, flat, glabrous, not veiny; seeds 10-14, 
closely placed, oval-oblong, compressed, dark brown. 

Low country; rare. Kalutara (Moon); Haragama; Lagalla. 

Fl. July, August; yellow, purple in bud. 

Throughout India, Malaya, and Trop. Africa. 


Readily known by the very numerous overlapping narrow Ilfits., 
‘making each pinna like the feather of a bird. 


63. ALBIZZIA,* Durazz. 


Trees, without thorns or prickles, |. bipinnate, fl. small in 
small heads or umbels, not polygamous; cal. funnel-shaped or 
tubular, segm. very short; pet. connate half way up into a 
cor.; stam. indef., slightly monadelphous at base, much longer 
than cor., anth. not gland-tipped; pod large, flat, thin, 
straight, dehiscent, seeds several on long funicles.—Sp. 30; 
10 in FZ. B. Lund. 

Pinnz 2-4 pair. 
Lfits. 5-9 pair; fl. stalked . 
Lfits. 10-14 pair; fl. sessile 
Pinnze 10-16 pair. 


Lfits. with midrib close to upper edge : .” 3. AS SEIPURAGS 
Lfits. with midrib nearly central 5 5 . 4. A. AMARA. - 


. A. LEBBEK. 
A. ODORATISSIMA. 


i) 


N 


1. A. Lebbek,t Benth. in Lond. Journ. Bot. ii. 87 (1844). Mara, S. 
Kona, Vakai, 7. 

Thw. Enum. 99._ C. P. 3130. 

Fl. B. Ind. ii. 298. Bedd. Fl. Sylv. t. 53. 

A large tree, with cracked, grey bark and wide-spreading 
branches, young parts elabrous, buds finely rufous-pubescent ; 
1. large, main rachis 7-9 in., swollen at base, glabrous, and 
shining, with a large circular gland close to base, and another 
at apex, pinne 2-3 pair, rather distant, 5-6in., rachis spread- 
ing horizontally, swollen at base, glabrous, with 2 small glands 
near the base, and 1 below the insertion of each pair of lfits. 
except the basal ones, lflts. 10-18 (5-9 pair) shortly stalked, 
large, 14-2 in., oval-oblong, rounded and retuse at apex, very 
unequal-sided, glabrous, rather paler beneath, fl. on slender 


* Dedicated to the old and noble family of the Albizzi, of Florence. 
+ From the Egyptian name, which, however, very doubtfully belongs 
really to this tree. 


Albizzia.] Leguninose. 129 


ped. 14-2 in., 1-4 together, the whole forming a lax corymbose 
terminal panicle; cal. nearly glabrous, tubular-funnel-shaped, 
segm. short, triangular; cor. twice as long as cal.; stam. 
forming a tassel { in. long ; pod large, 9-12 in. by 1# in. wide, 
bluntly pointed, thin, pale yellow; seeds about 12, small, com- 
pressed. 

Dry region; rather rare. Anuradhapura; Uma-oya. Often planted. 
F]. April; greenish-white. 

Throughout Tropical and Sub-tropical Asia and Africa, but usually a 
planted tree. sal ‘ 

Heart-wood brown, hard, shining, with large pores; a fine wood, does 
not warp or crack. 


2. A. odoratissima, Senth. zm Lond. Journ. Bot. iii. 88 (1844). 
Suriya-mara, Huriyi, S. Ponnaimurankai, 7. 

Mimosa odoratissima, L. f. Suppl. Plant. 437; Moon Cat. 73. Thw. 
Enum. too. C. P. 1529. 

Fl. B. Ind. ii. 299. Bedd. FI. Sylv. t. 54. 

A very large tree, with thick grey rough bark and spreading 
branches, young parts pubescent; |. rather large, rachis 4-6 in..,. 
pubescent, with a single large sessile gland near the base, and 
generally one between bases of terminal pair of pinne, pinnz 
generally 4 pair, rather distant, about 4in., pubescent ; Iflts. 
20-28 (10-14 pair), sessile, rather large, #-1}in., oblong, 
rounded, apiculate, very unequal-sided, the midrib near the 
upper margin, glabrous above, finely downy and glaucous 
beneath, stiff; fl. sessile, few, heads $—? in. diam., peduncles 
2in., pubescent, 1-4 together, arranged in terminal panicles, 
cal. densely pubescent,segm. short, broadly deltoid; cor. pubes- 
cent outside, cut into 4 acute segm.; stam. twice as long as 
cor.; pod 4~7 in. by 1-14 in. wide, very shortly stalked, abruptly 
pointed, roughish but glabrous, slightly veiny; seeds 6-12, 
small, much flattened. 

Low country in both moist and dry regions, especially the latter; 
common. Fl. May—September; pure white, sweet-scented. 

Also in India and Malacca. 

Heart-wood dark brown, very hard, works and seasons well. 

3. A. stipulata, Bow. in Encycl. XIX* Sivcle, ii. 33 (1834 ?). 
Kabal-mara, Hulan-mara, 5S. 

Thw. Enum. too. C. P. 1517. 

FI. B. Ind. ii. 300. Bedd. Fi. Sylv. t. 55. 

A tall tree, with pale grey, thin, rather smooth bark, 
young branchlets and buds covered with fine golden pu- 
bescence; |. large, rachis 8-14 in., stout, at first pubescent 
afterwards glabrous, angled above, with a prominent gland 
half way between base and lowest pair of pinnez, and also 
between the bases of each pair, pinnae 10-15 pair, 34—5 
in., thickened at base, pubescent, Iflts. very numerous, 60-80 

PART II. K 


130 Leguminosae. [Albizzia. 


(30-40 pair), crowded, subsessile, about %in., $-ovate, the 
midrib close to upper margin, truncate-rounded at base, acute, 
slightly pilose beneath, stip. very large but soon falling, 14 in., 
unequally cordate, acuminate, with a filiform tail, base much 
dilated on one side, membranous, crisped, pubescent, pinkish- 
orange ; fl. on very short ped., in small lax heads, peduncles 
1? in., divaricate, pubescent, 1-4 together, arranged in rather 
small terminal or axillary panicles; cal. tubular, puberulous, 
very slightly toothed ; cor. twice as long as cal., with acute 
recurved segm.; stam. I} in., fully 4 times as long as cor.; pod 
6-8 in. by 14 in. wide, very shortly stalked, tapering to base, 
blunt, roughish but glabrous, pale brown, 8—12-seeded. 

Moist low country up to 2000 ft.; common. FI. April, May; white, 
the end half of the filaments pink; sweet-scented. 

Also in E. Bengal, Burma, and Malaya. 

Readily recognised by the golden-orange colouring of the young 
foliage. The great pink stipules are very characteristic of this beautiful 
tree, but they very soon fall away (not ‘persistent’ as given in FI. B. Ind.), 
leaving no trace beyond a small oval scar. The Iflts. quickly fold together 
after being gathered. 

A quick-growing tree, often planted for shade. Heart-wood pale 
brown, soft, light, recommended for tea-boxes. 


4. A. amara, Bozv. 7n Encycl. X/X° Siecle, i. 34 (1834 ?). UWyil, 7. 

Mhwa oun toon Cee sensro: 

Bibs indi. 300s ebeddil i oylvat. O12 

A rather small much-branched tree, young shoots 
densely pubescent; 1. rather small, rachis 3-4 in., densely 
pubescent, with a small gland midway between base and 
lowest pair of pinne and another between the bases of that 
pair, pinnz 10-16 pair, 1-14 in., densely pubescent, lfits. 20-60 
(10-30 pair), closely placed and slightly overlapping, very 
small, }in., sessile, linear, subacute, the midrib nearly in the 
centre, pubescent, glaucous beneath, articulated and easily 
detached; fl. shortly stalked in rather dense heads, 1-14 in. 
diam., peduncles solitary or 2 or 3, ascending, from axils of 
young |. and not forming a panicle; cal. funnel-shaped, pube- 
rulous, segm. short, ciliate; cor. pubescent outside, 3 times as 
long as cal.; stam. over $in.; pod 4-5 in. by 1 in. wide, shortly 
stalked, apiculate, very thin, veiny, 6-8-seeded. 

Dry and desert regions; rare. North of the Island (Gardner); Kalpi- 
tiya; Mannar, abundant; near Nalande (Alexander). FI. August; 
pinkish. 

Also in Peninsular India and E. Trop. Africa. 

This is, no doubt, the ‘ Weel-tree’ which Cordiner (I. c. 34) records 
from Mannar in 1804. 

Wood extremely hard and strong, heavy, and close-grained, dark 
rown. 


Pithecolobium.| Leg UI21N0SE. 31 


A. moluccana, Miq., a native of Java and other Malayan Islands, has 
been a good deal planted as a shade-tree during the last ten or twelve 
years, having been introduced in 1880. _ Its growth is extraordinarily 
rapid; a tree in the Bot. Garden at Peradeniya 63 years old, cut down in 
1893, was 89 ft. high, and girthed 63 ft. at a yard above the ground. 


64. PETHECOLOBIUM, Ja77. 


Trees, sometimes thorny, |. bipinnate, with or without stip., 
fil. small in rounded heads or umbels, not polygamous; cal. 
campanulate or tubular, segm. very short ; pet. connate half 
Way up into a cor.; stam. indef., monadelphous at base, much 
longer than cor., not gland-tipped; pod flat, usually curved 
into a circle and often twisted, usually dehiscent, sometimes 
jointed, several-seeded.—Sp. 100; 15 in A’. B. Ind. 


Pinnze I pair : : : i : 3 . I. P. GEMINATUM. 
Pinnz 2-4 pair. 
Lfits. 5-8 pair 2. P. UMBELLATUM. 
Lfits. 1-3 pair 3. P. BIGEMINUM. 
Lfits. 13-20 pair 4. P. SUBCORIACEUM. 


1. P. geminatum, Zenth. in Lond. Journ. Bot. iii. 202 (1844). 

Thw. Enum. 100. Calliandra (?) geminata, Benth. in Trans. Linn. 
spcyaex. 546. GC. P. 1531. 

Bie. Ind: ii. 303. Hook. Ic. Plant. t. 1510. 

A small bushy tree with smooth grey bark, young branches 
slender, pubescent ; |. small, stip. spinous, persistent, at length 
#in., straight, very sharp, dark brown, pubescent, rachis very 
short 4-4 in., pubescent, with stalked erect gland at end, 
pinne I pair, about I in., slender, pubescent, lflts. 2-10 (1-5 
pair), sessile, unequal, 4—? in., oblong, unequal at base, obtuse, 
mucronate, dark green, glabrous and shining, with conspicuous 
veins; fl. sessile, crowded, heads (with stam.) 14 in. wide, 
peduncles slender, pilose, 1-3 from axils of 1.; cal. shallowly 
campanulate, glabrous, segm. widely triangular; cor. cam- 
panulate, segm. ovate acute; stam. exserted }—}in. beyond 
cor.; pod 2-4 in. by 2 in. wide, shortly stalked, much curved, 
often twisted, pointed, flat, usually much constricted (almost 
moniliform) between the seeds, dark brown, glabrous, shining, 
opening by ventral suture only, seeds about 8. 

Dry region; common. Fl. Sept.—Nov. 

Endemic. The FI. Brit. Ind., following Beddome, gives also Peninsular 
India, but I have seen no specimens thence. 

First collected by Koenig. Bentham has referred this latterly, with 
a doubt, to Calliandra, not having seen the pod, which is, however, 
quite that of Pzthecolobium. 


P. dulce; Benth., the ‘Madras Thorn’ of the English in Ceylon, is 
much planted for hedges and as a shade-tree in towns, and was, doubt- 


ie2 iL, egu minose. [P2thecolobium. 


less, a very early introduction. It is figured in Bedd. FI. Sylv.t. 188, and 
is a native of Tropical America. There are large old trees in Colombo 
and Galle forts. 


2. P. umbellatum, Benth. ti Lond. Journ. Bot. iii. 202 (1844). 
Eyamalai, Ichavalai, 7. 

Mimosa umbellata, Vahl, Symb. 11. 103. Thw. Enum. too. C. P. 1530. 

Fl. B. Ind. ii. 303. 

A small tree with rather smooth grey bark, lateral twigs. 
often ending in a spinous point, young parts glabrous; 1. 
rather small, rachis about ?in., glabrous or pubescent with a 
gland between insertion of each pair of pinnae, stip. short, 
acute sometimes spinous, often absent, pinne 2 pair, 14-14 in., 
lflts. 10-16 (5-8 pair), sessile, closely placed, $—2 in., oblong, 
very obtuse, glabrous on both sides, the terminal pair broader 
and approximated ; fl. on glabrous ped., umbels on peduncles. 
about 1in. long; cal. tubular-funnel-shaped, glabrous, segm. 
very short, triangular; cor. twice as long as cal., puberulous, 
segm. lanceolate, acute; stam. exserted $ in. beyond cor., pod 
4-5 in. by 2 in. wide, shortly stalked, sucaredl nearly into a 
circle, not twisted, strongly constricted between seeds (almost 
moniliform), clabrous and shining, ultimately breaking up 
into 1-seeded indehiscent joints; seeds 7-10, very flat, nearly 
circular, 

Dry and intermediate country; rare. Jaffna; Arippu; Mannar; 
Teldeniya. Fl. Sept—Nov.; creamy white. 

Also in Southern India and Malay Islands. 

First collected by Kcenig. Turns black in drying. Readily dis- 
tinguished by its jointed indehiscent pod. Mimosa nitida, Vahl, is 


probably according to Benth. (l.c. 583) this species; the specimen in 
Vahl’s Herb. is from Ceylon. 


P. Saman, Benth. mga Saman, Willd.), a native of Central and 
Tropical S. America, whence it was introduced into Jamaica, where 
it is called ‘Guango,’ has been very much planted as a shade-tree by 
roads and in public places. It was introduced to Ceylon about 1851, and 
there are very large trees in Peradeniya planted in that year. The pod 
is straight and indehiscent, with a sweet pulp round the seeds, and it 
scarcely fits well in the genus Pzthecolobium. It is often called the ‘ Rain- 
tree,’ and grows with great rapidity to its full size. 

3. BP. bigeminum, Senth. in Lond. Journ. Bot. iii. 206 (1844). 
Kalatiya, S. 

Fl. Zeyl. n. 218. Mimosa bigemina, L. Sp. Pl. 517. Thw. Enum. too. 
C. P. 644. 

Fl. B. Ind. ii. 303. Hort. Malab. vi. t. 12. 

A moderate-sized tree, with thin, smooth brown bark, 
young parts glabrous; |. rather large without stip., main 
rachis 4—6 in., glabrous, channelled above, with an oval gland 
near the base and another between bases of upper pair of 
pinne, pinne 2 (rarely 3) pair, unequal, lowest pair very much 


Pithecolobium.] Leguminose. AS 


the smallest, distant, horizontal, rachis thickened at base, with 
an oval gland just below insertion of each pair of Iflts. except 
the lowest, Ifits. large, shortly stalked in lowest pinnz usually 
2 (I pair) rarely 4 or 6 (2 or 3 pair), 24-3 in., in upper pinne 
usually 6 (3 pair) 4-5 in. the terminal pair much the largest, 
all oval or lanceolate, acute at base, caudate-acuminate, 
glabrous, thin, dark green; fl. sessile, 2-4 together in little 
pedunculate clusters, arranged on long branches of slender 
pubescent panicles rather shorter than 1.; cal. campanulate, 
pubescent, segm. short, deltoid; cor. 3 or 4 times as long as 
cal., segm. lanceolate, acute; stam. tassel-like, exserted for 
x in. beyond cal.; pod 3-5 in. by ? in. wide, flat, strongly 
curved into a ring or spiral, bluntly pointed, rough but 
glabrous, not indented bet. seed, reddish-brown, bright red 
within, 5—8-seeded. 

Moist region up to 4ooo ft. or more; common. FI. June; cream- 
coloured. 

Also in India. 

The S. name given to his specimen by Hermann is ‘ Ulhanda.’ 

ne wood is poor and light, the heart-wood scanty or wanting alto- 
gether. 


4. P. subcoriaceum, 7iw. Enum. i100 (1859). Mimini- 
mara, S. 

fibwe Enum: too: GC. P. 337. 

Fl. B. Ind. ii. 305. Bedd. Fl. Sylyv. t. 189 (P. anamallayanum). 

A large tree, with smooth thin grey bark and spreading 
branches, branchlets covered with very fine brown pubescence; 
1. numerous, crowded, rachis 2-3 in., stout, pubescent, with 2 
distant sessile glands below lowest pair of pinnz and one 
below each of the upper pairs; pinnz 2-4, usually 3 pair, 
3-4 in., the uppermost the longest; lflts. 26-40 (13-20 pair), 
sessile, }-2 in., obtuse, oblique, the upper side dilated at base, 
margin reflexed, glabrous above, pilose beneath, rather thick 
and stiff; fl. sessile, heads lax on short peduncles, arranged in 
pyramidal terminal panicles, bracts small, fulvous-pubescent ; 
cal. tubular, densely pubescent, segm. short triangular; cor. 
twice as long as cal., pod shortly stalked, curved into a circle, 
puberulous, orange-yellow within, seeds 5-7, jet-black. 

Montane zone, 4000-6ooo ft.; common. Fl. March, August; greenish- 
white. 

Also in mountains of S. India; if P. anamallayanum, Bedd., be the 
Same species, as Bentham considers. 

Turns black in drying. Wood yellowish, light, soft, smooth. 


134 Rosacee. [Pygeume.. 


XLV.—ROSACE/. 


HERBS, shrubs or trees, 1. alt., simple or compound, with 
stip.; fl. bisexual (rarely polygamous) regular ; cal.-tube free 
from (in Photinia adnate to) ov., segm. 4-6, sometimes with 
small adnate bractlets (epicalyx) outside; pet. 4-6 or 0; stam. 
def. or indef., perigynous; carp. 1-3 or indef., usually distinct 
and free though often enclosed in cal.-tube (adnate to it in 
Photinia), style often basal, ovules 1 or 2; fruit various, in- 
dehiscent, usually of dry (in Rubus pulpy) achenes, in Photznza 
berry-like ; seed without endosperm, cotyledons plane-convex. 
Carp. not included in cal.-tube. 


Carp. solitary; trees (Prune) . , : : . I. PYGEUM. 
Carp. numerous. 
Carp. drupaceous; shrubs . : : ; - 2, RUBUS: 
Carp. dry (achenes); herbs . : . 3. POTENTILLA. 
Carp. included in cal-tube, but not adnate to it 
(Poteriea). 
Peta: 
L. simple . 4. ALCHEMILLA. 
L. pinnate . 5. POTERIUM. 
Bet 5 . 6. AGRIMONIA. 
Carp. adnate to cal.-tube (inferior) (Pomee) . 7. PHOTINIA. 


This is an Order mainly of temperate distribution, and all our 12 
species, with the exception of Pygeum zeylanicum, are montane plants. 
Rubus moluccanus also extends downwards into the moist region as low 
as 1000 ft. No species occurs in the dry region. All the genera are 
British, except Photiénza and Pygeum. 


I. PYGEHUM, Gaerin. 


Trees, |. simple, entire, with 2 glands at base, stip. minute, 
fl. small, in axillary racemes; cal.-tube obconical, segm. 5 or 63, 
pet. very small, 5 or 6 tomentose; stam. 12-20 or more, 
perigynous, spreading; carp. solitary, free, ovules 2, pendulous, 
style terminal, stigma capitate; fruit transversely oval, 2-lobed,, 
indehiscent, pericarp thin, dry; seed solitary with large 
hemispherical cotyledons.—Sp. 20; 16 in Fl. B. Ind. 
L. quite glabrous ; stam. 20 or more; ov. glabrous. 1. P. WIGHTIANUM 
L. hairy on veins beneath ; stam. 12; ov. hairy . 2. P. ZEYLANICUM. 


1. BP. Wightianum, B/. del. Botan. 2 (1855). 

Polyodontia zeylanica, Wight, Ill. i. 203. Thw. Enum. 102. C. P. 638. 

Fl. B. Ind. ii. 319. Wight, Ic. t. 256 (Polyodontia zeylanica) from a 
Ceylon specimen. Bedd. FI. Sylv. t. 59 (P. zeylanicum). 


Rubus.) Rosacee. ars 


A moderate-sized tree, bark rather rough, pale brown, 
young shoots slightly adpressed-pubescent; 1. 3-6 in., broadly 
oblong-oval, rounded at base, obtuse, entire, glabrous on both 
sides, thick, with 2 large round or oval immersed glands one 
on either side of midrib near its base, petiole about 4 in., stout, 
furrowed, fl. about $ in. wide, ped. longer than cal., spreading, 
racemes numerous, erect, narrow, stout, 3—4 in. long, finely 
pubescent, cal. pubescent, segm. usually 5; pet. oval, villous on 
Margins, stam. 20 or more, much exceeding pet.; ov. glabrous ; 
fruit about 2 in. wide by 2 in. long, somewhat didymous, apicu- 
late, glabrous, pericarp thinly coriaceous. 


Var. 6, parvifolium, 7iw. C. P. 1596. 


L, 2-3 in., more rigid, margins often recurved, sometimes 
lanceolate, acute; racemes 1-14 in., fl. smaller, on shorter ped. 
Montane zone, above 4000 ft.; common; and occasionally at some- 


what lower elevations. F]. August-October; white, sweet-scented. 
Also on the hills of S. India. 


2. P. zeylanicum, Gaerin. Fruct. i. 218 (1788). Golu-mora, 
HRankumbal-kétiya, S. 

Polyodontia Walkeri, Wight, Ill. i. 203. Pygeum Watlkeriz, Bl. Mel. 
Beem. Low. Enum. to2.. C. P. 1532. 

Fl. B. Ind. 11. 321. Gaertn. Fruct. 1. t. 46, f. 4 (fruit only). 

A large tree, with smooth grey bark, branchlets with 
numerous large linear lenticels, young shoots densely velvety 
with yellow silky hair; 1. rather large 5-7 in., ovate-oval, 
rounded at base, acuminate, subacute, glabrous above, hairy 
on the prominent veins beneath, entire, stiff, with 2 basal 
glands as in the last, petiole short, stout ; fl. about 4in., ped. 
woolly, scarcely longer than cal., racemes numerous, 2-4 in., 
erect, narrow, rufous-tomentose; cal. woolly-pubescent, segm. 
usually 6, shortly acuminate; pet. 6, very woolly; stam. 
usually 12; ov. hairy; fruit 2-1 in. wide by 2—# in. long, not 
apiculate, rusty-pubescent becoming glabrous. 


Moist low country, rather common. FI]. Feb.; creamy white. 

Endemic. 

The pet. are difficult to distinguish from the cal.-segm. Wood close- 
grainea, rather heavy; yellow. ‘The seeds when bruised have a strong 
odour of prussic acid. 

In drying, the leaves of both species turn a dark coppery brown. 


z, RUBUS, /. 


Prickly shrubs with long sarmentose stems, 1. simple or 
compound, stip. petiolar or cauline, fl. in axillary and terminal 


136 Rosacee. [Rubus. 


corymbose panicles; cal. persistent, segm. 5, large, valvate; 
pet. 5, imbricate; stam. indefinite; carp. numerous on a convex 
receptacle, style subterminal; fruit of numerous small crowded 
drupes, stone hard, woody; seed pendulous.—Species va- 
riously estimated, say 200; 41 in FZ. B. Ind. 
L. simple. 
Pet. 4 length of cal.-segm : 
Pet. more than 2 2 length of cal.- -segm : 


L. 3-foliolate 
L. imparipinnate 


R. GLOMERATUS. 
R. MOLUCCANUS. 
R. ELLIPTICUS. 

R. LASIOCARPUS. 


SS) 


1. R. glomeratus, L/. Bijdr. 1111 (1826). 

R. micropetalus, Gardn. in Calc. Journ. Nat. Hist. viii. 6. 2. rugosus, 
var. 6, Thw. Enum. to1. Trim. Syst. Cat. 30. C. P. 1536, 1537. 

FI. B. Ind. ii. 328. 

Stems slender with few small hooked prickles, at first 
covered with cinnamomeous tomentum, afterwards glabrous, 
1. simple, 3-5 in., rather narrower than long, deeply cordate at 
base, acute, 3- or obscurely 5-lobed at base, with usually acute 
lobes, irregularly dentate-serrate, hairy on the veins above, 
densely felted with fine pale cinnamon-coloured tomentum 
beneath and with the reticulate venation very prominent, 
petiole 14-24 in., stip. deeply pectinate with filiform segm.; fl. 
crowded in small short racemose clusters on short stout 
peduncles in axils of upper 1. and of pectinate bracts, the 
whole forming an elongated tomentose panicle; cal. densely 
covered with cinnamomeous hair, segm. entire or slightly 
toothed near apex, pubescent within; pet. very small, about 4 
length of cal.-segm., rounded; fruit very small, purplish-red. 

Upper montane zone; rather common. FI. Jan.-June; white. 
Also in Malaya, but not recorded from Indian Peninsula. 
Blume’s description is very brief, and the pet. are not described, but 


it appears to apply well to our plant, which is, however, perhaps but a 
variety of A. moluccanus. 


2. R. moluccanus, L. Sf. P/. 1197 (1753). Wel-buté, S. 
Moon Cat. 4o. &. rugosus, Sm., Thw. Enum. tor. C. P. 2765. 
ABs Inds 10330). Wisiht, lext225. 
Stems stout, densely covered with woolly grey or yellowish 
hair and set with numerous strong hooked prickles; 1. simple, 
3-5 in., usually about as broad as long, cordate at base, 
acute, more or less deeply 5- (or 7-) lobed with obtuse or sub- 
acute lobes, unequally dentate-serrate, glabrous or hairy on 
veins and bright green above, very hairy and more or less 
yellowish or grey beneath with prominent reticulate venation 
and often with prickles on the main veins, petiole long, 


Rubus.) Rosacee. AG) 


14-24 in., very hairy, with prickles beneath, stip. large, ovate, 
deeply pectinate, very silky, enclosing the buds, caducous ; 
fl. in elongated terminal panicles, on long stout ped., bracts 
oval, toothed or pectinate at end only; cal. densely silky- 
hairy, segm. entire or pectinately toothed at end; pet. fully half 
as long as cal.-segm.; fruit bright red or dull purplish-red. 

Var. 8, Fairholmianus,* Gard. 1.c. 7 (sp.). A. vrugosus, var. y. 
hye bc GC. -P..135: 

Stems with few nearly straight prickles, tomentum nearly 
white; 1. very rugose above; fruit of very numerous carp., 
dark purple. 

Var. y, macrocarpus, Gardan. |. c. 7 (sp.). A. rugosus, var. 6. Thw. 
Pee, P1535. 

Prickles nearly straight, tomentum cottony, cinnamo- 
meous; l. very rugose above; fruit of numerous very large 
juicy carp., black. 

Moist country at all elevations above 1000 ft.; very common ; vars. 8 
and y only in upper montane zone. FI. allthe year; white; vars. 6 and 
y, bright pink. 

Common throughout India and Malaya. 

This is a very variable species in the hill districts ; and the two 
varieties given were considered separate species by Gardner. The fruit 


of var. y is the only real “ B/ackberry” in Ceylon, it is large and juicy, 
and when quite ripe has a good flavour. 


3. R. ellipticus, Sy. 27 Rees’ Cyclop. xxx. (1815?). 

R. flavus, Ham., Thw. Enum. to1. FR. Gowreephul, Roxb., W. and 
A. Prod. 298. -C. P. 25096. 

Fl. B. Ind. ii. 336. Wight, Ic. t. 230 (2. Gowreephul). 

Stems flexuose, finely pubescent and also clothed with 
numerous long red horizontal hairs and scattered stout hooked 
prickles; 1. pinnately 3-foliolate, rachis 2-24 in., cylindrical, 
armed as the stem, stip. obviously petiolar, 4 in., linear-filiform, 
erect, Iflts. very shortly stalked, 2-3 in., broadly oblong-oval or 
rotundate, obtuse or abruptly acute, sharply serrate, glabrous 
above, densely and finely white-pubescent beneath with the 
lat. veins prominent ; fl. rather small, on long ped., in copious, 
dense, pubescent, axillary and terminal panicles, bracts linear; 
cal.-segm. acute, mucronate, densely pubescent ; pet. a little 
longer than cal.-segm., obovate; fruit globular, of very 
numerous carp. which are covered with a few long hairs, deep 
yellow, slightly juicy, stone rugose. 


* Dedicated to W. Fairholme, Esq., who accompanied Mr. Gardner 
on a journey to Adam’s Peak in 1846, when they were lost for five days in 
the forest. 


138 Rosacee. [Rubus.. 


Montane region; common. FI. all the year; pure white, anthers. 
cream-colour. 

Also in Western Himalayas, Burma, &c. 

This may be the &. Janzculatus of Moon Cat. 40, from Maturata. 

Though this species is very similar to the Blackberries of Europe, its. 
deep yellow finely hairy fruit is scarcely edible. 


4. R. lasiocarpus, Sm. zz Rees’ Cyclop. xxx. (1815). 
R. parvifolius, Moon Cat. 40. Thw. Enum. tor. C. P. 1533. 
MB. Indi 339, Wight, Ic. t. 232: 


Stems erect, glabrous but covered with a fine white pinkish 
or blueish “bloom” easily rubbed off, armed with large nearly 
straight compressed prickles ; |. imparipinnate, rachis 34—5 in., 
stout, horizontal, flexuose, with numerous hooked prickles, 
furrowed above, stip. rather small, linear, silky, lflts. 5-9 
(2-4 pair and end one), shortly stalked, lanceolate-oval or 
ovate, acute or rounded or cordate at base, acute at apex, 
coarsely and irregularly serrate, glabrous above, very white 
with fine cottony tomentum beneath; fl. in long pubescent 
ped. in small short axillary and terminal corymbs, bracts. 
filiform; cal.-segm. very acute, aristate, woolly on both 
sides; pet. rounded; fruit rather large, $ in. or more, carp. 
numerous, fleshy, pink, densely covered with fine white 
pubescence. 


Var. B, subglaber, 7hw. /.c. R&R. leucocarpus, Arn. Pug. 16. 
(Coes TseuL 


Lfits. smaller, broadly ovate, not white beneath but 
glabrous save for scanty hair on the veins. 


Montane region, 3000-6000 ft. or more; common. Var. 6, Nuwara. 
Eliya. Fl. throughout the year, pink. 

Also throughout India and in Java. 

There are two forms, large and small-flowered, the former being met 
with at the higher elevations only. 

This is known as the ‘ Wild Raspberry,’ which it much resembles ;. 
the fruit, however, has no flavour, and is besides covered with a woolly 
coat, which renders it inedible. 


Fragaria vesca, L. Walker-Arnott, in Pug. 16, say she has seem 
badly dried specimens from Ceylon gathered at 6000 ft. And about 
Nuwara Eliya patches of so-called ‘wild’ strawberries are to be found 
occasionally, but only, | think, as escapes from cultivation.* About 
Ootacamund in the Nilgiris, however, a wild strawberry, /. mdlgerrensis, 
Schldl., is said to be very common, and is figured (as /. e/atzor ) in Wight,, 
Ic. t. 988 ; but our plant does not agree with this, but with the common 
wild strawberry of England. 


* T am informed that the first wild strawberries noticed at N. Eliya 
came up in a Cinchona clearing made in 1864, but Arnott’s must have: 
been collected at a much earlier date. 


Potentilla.] Rosacec. 139 


3. POTENTILLA, J. 


Herbs, |. compound, stip. adnate to petiole, fl. in cymose 
panicles; cal. with 5 segm., persistent, with 5 adnate bractlets 
outside (epicalyx); pet. 5; stam. indef.; carp. indef., distinct, 
style basal or terminal, ovule solitary, pendulous; fruit com- 
posed of numerous very small dry indehiscent achenes.— 
Sp. 120; 39 in. Fl. B. Ind. 


L. imparipinnate. : : : : ‘ . I. P. MOONIANA. 
L. palmately 5- foliolate : . ‘ é : 2 By Les IRIN TUMSUAS 


1. P. Mooniana, Wight, Jc. 1. t. 233 (1840). 

Thw. Enum. tor. C. De 2767. 

Fl. B. Ind. ii. 349. Wight, l. c. (no petals): from a Ceylon specimen. 

A perennial herb with a stout woody rootstock and long 
stout, prostrate, very hairy branches often rooting at the 
nodes; 1. large, in a rosette at top of rootstock, and distant on 
the branches, imparipinnate, rachis stout, very hairy, usually 
3-4 in. (sometimes much longer), stip. acuminate, # in., mem- 
branous, lflts. numerous, sessile, very unequal in size, usually 
in alternately larger and smaller pairs, larger ones 4-1 in., 
oval, obtuse, coarsely serrate, with bristle-tipped serrations, 
nearly glabrous above, silvnesley on the veins beneath, 
plicate, smaller ones about 4 in., rotundate; flowering branches 
erect from leaf-axils, 6-12 in. , bearing the fl. on long, slender, 
hairy, stiff ped. in a lax terminal irregularly dichotomous 
panicle; cal.-segm. oval, acute, hairy, bractlets similar but 
rather shorter, pet. orbicular, longer than cal.-segm.; achenes 
very numerous, smooth, on a hairy receptacle. 

Upper montane zone; rather rare. Nuwara Eliya; Totapella ; 


Horton Plains; Pedurutalagala. FI. July, August; yellow. 
Also in the Himalaya, but not in S. Indian mountains. 


2. P. Kleiniana, W. and A. Prod. 300 (1834). 

Thw. Enum. 102. C. P. 2768. 

FI. B. Ind. ii. 359. Wight, Ill. t. 85. 

Annual, stems numerous, prostrate but not rooting, slen- 
der, hairy; |. palmately 5-foliolate, the radical ones on very 
long ee the stem ones on much shorter hairy petioles, 
stip. 4 in., lanceolate, acute, glabrous, Iflts. unequal, nearly 
sessile, Se in. (terminal the largest), oval or spathulate-oval, 
acute at base, obtuse, deeply toothed, glabrous above, silky 
on veins beneath ; fl. smaller, but arranged much as in the last, 
cal.-segm. and bractlets with a few long hairs; pet. scarcely ex- 
ceeding cal., achenes on a glabrous receptacle, deeply wrinkled. 


140 Rosacee. [Poterium. 


Upper montane zone; rather common. FI. April, September, 


October ; yellow. a ; 
Also in the Himalaya and Nilgiris; and in Yunan, Java, and Japan. 


4. ALCHEMILLA, Z. 


Perennial herb with prostrate stems, |. simple, with large 
petiolar stip., fl. minute, in corymbose cymes; cal. campanulate 
with 4 adnate bractlets outside, segm. 4; pet.o; stam. 4; disk 
at mouth of cal.-tube large and thick; carp.1, free, style basal ; 
fruit a single achene enclosed in persistent cal.-tube—Sp. 30 ; 
2 in Fl. B. Ind. 


A. indica, Gardn. in Calc. Journ. Nat. Hist. viii. 8 (1847). 

A. vulgaris, Arn. Pug. 16 (non L.). A. vulgaris, L., var. sarmentosa, 
Aline Ids, WO, (C, 12, Ao. 

Fl. B. Ind. ii. 361. Wight, Ic. t. 229 (A. vulgaris). 

A perennial herb with a semi-woody erect rootstock 
clothed with remains of sheathing stip., stems numerous, very 
long, prostrate, with abundant spreading hair; 1. numerous, 
suborbicular, the basal ones 14-2} in., the stem ones I-14 in., 
cut into 7-9, shallow, rounded lobes, sharply crenate-serrate, 
silky-hairy on both sides but especially beneath which is 
silvery white, plicate in bud, petiole variable in length, very 
long in basal |., covered with long spreading hair, stip. very 
large, connate below into a tube surrounding the stem, the 
free part oblong-lanceolate, acute, often toothed at the end, 
hairy, membranous or herbaceous; fl. very small, on short ped., 
cymes small, rather dense, on erect hairy peduncles from axils 
of stem-l., bracts numerous, ovate, often toothed ; cal. villous 
with white hairs. 

Var. 6, sibthorpioides, /7Z. 7.1. c. 361. 

“Very slender, 1. $- in. diam., cymes panicled.” 


Upper montane zone ; rather common. I have not seen var. 8, which 
was collected by Gardner. Fl. April; yellowish-green. 

Occurs also in the Nilgiris Pulney Hills. 

When very luxuriant the leaves are 9-11-lobed, with the lobes over- 
lapping. Wight, followed in FI. Brit. Ind., quotes ‘A. zeylanica, Moon’ 
for this, but there is no such name in his Cat., and this high mountain 
plant could scarcely have been known to him. 


5. POTERIUM, Z. 


Erect perennial herb, 1. imparipinnate, stip. petiolar, fi. 
small, polygamous, nearly sessile in small dense terminal 
heads ; cal. with a top-shaped persistent tube, and 4 petaloid 


Agrimonia.] Rosacee. I4I 


deciduous segm.; pet. 0; stam. 4 or 2; carp. I-3; ripe carp. 
solitary, enclosed in hardened cal.-tube.—Sp. 20; 5 in FZ. B. 
Ind., but no species is known from the Indian Peninsula. 


P. indicum, Garvan. in Calc. Journ. Nat. Hist. viii. 10 (1847). 

fibws Emume 1025 G2 Po 2771. 

Wi Bb. Indi. 363. 

An erect perennial herb, 2-3 ft., stems glabrous, striate ; 
], pinnate, rachis 3-6 in. glabrous, sheathing at base, Ifits. 
shortly stalked, about 4 in. broadly oval, with a few large 
teeth, nearly glabrous; head of fruit small, 2 in., globular, on 
very long erect peduncle, bracts rounded ciliate, ripe carp. 
enclosed in persistent cal.-tube which is about }in., trapezoid- 
ovoid, hard, with 4 narrow wings. 

Montane zone; very rare. This has apparently been twice gathered, 
first on ‘Adam’s Peak, 1835,’ by Mr. H. de Alwis, from whose specimens 
in Hb. Perad. (now C. P. 2771) Gardner described the species; and 
afterwards, in 1839, by Sir J. A. Stewart Mackenzie,* Governor of Ceylon, 
whose excellent specimens, without definite locality, are now in Hb. Kew. 
Subsequent search for it has been vain. 

Endemic. 

There is some doubt as to the locality of these specimens, and I sus- 
pect all are really from the same gathering. My material in Hb. Perad. 
is very scanty, and I am unable to give a full description. 


Rosa. There is no wild species of Rose in Ceylon. 


6. AGRIMONTIA, J. 


Erect perennial herb, |. interruptedly imparipinnate, stip. 
petiolar, fl. small, in spikelike racemes; cal. persistent and 
enlarged in fruit, tube turbinate, with 3 whorls of hooked spines 
round the top, segm. 5; pet. 5; stam. usually 5; disk thick, 
nearly closing mouth of cal.-tube; carp. 2 at base of cal.-tube, 
styles terminal; fruit of 2 (or 1) achenes enclosed in hardened 
enlarged cal.-tube.—Sp. 8; 3 in “7. B. Ind., but no species 
in Peninsular India. 


A. zeylanica, J/00n Cat. Ceyl. Pl. 37 (1824). 
A. LEupatorium, Thw. Enum. 102; Arn. Pug. 16. C. P. 2769. 
Fl. B. Ind. ii. 362. Wight, Ic. t. 224 (A. Eupatorium) (bad). 


A perennial herb about 3 ft. high, stem erect, slightly 
branched, villous with coarse shaggy hair; 1. numerous, the 
radical ones often 8-10in., the stem ones smaller and 
gradually passing into bracts, rachis stout, villous-hairy, stip. 


* Printed as ‘Sir J. G. McKenzie’ in FI. B. Ind. 


142 Rosacee. [ Photinia. 


large, leafy, hairy, adnate to petiole below, free part semi- 
hastate, deeply toothed or gashed, Iflts. very unequal, the 
larger ones #-14 in., very broadly oval or subrotundate, obtuse, 
coarsely toothed, silky-hairy on both sides, especially beneath, 
the smaller ones often minute; fl. very shortly stalked, distant, 
in very long erect terminal spikes, each in axil of toothed 
leafy bract; cal.-segm. acuminate; pet. oval-oblong; styles 
exserted, stigma bilobed ; cal.-tube in fruit strongly grooved 
hairy, spines erect, glabrous, red. 

Montane zone, 4000-7000 ft., in grassy or bushy places ; rather rare, 
though locally plentiful, especially in Uva Province. Fl. Feb.—April ; 
yellow. 

Endemic. 

This is very close to the common A. Lufatorium of Europe, but is a 
sufficiently well-marked local variety todeserveaname. It is remarkable 
that it does not occur in the S. Indian hills. 


7, PHOTINIA, Lindl. 


Trees, 1. simple, with subulate deciduous stip., fl. small, in 
terminal corymbs; cal.-tube adnate to ov., segm. 5; pet. 5; 
stam. I5 or 20; ov. inferior, 2-celled, with 2 ascending ovules 
in each cell, styles distinct, stigmas dilated; fruit a small 
berry-like pome, flesh scanty, endocarp membranous ; seeds 2 
with a leathery testa.—Sp. 8; 5 in AZ Bb. Ind... 


P. Notoniana, W. and A. Prod. 302 (1834). 

Thw. Enum. too. C. P. 136. 

Fl. B. Ind. ii. 380. Wight, Ill. i. t. 86; Ic. t.991. Bedd. FI. Sylv. 
t. 192. 

A small tree with wrinkled bark, young parts glabrous ; 
]. 4-6 in., oval or lanceolate, tapering to or rounded at base, 
acuminate, acute, entire, somewhat undulate, glabrous on 
both sides, thin, rather stiff, shining above, paler beneath, 
petiole long, 1-14 in.; fl. small, 4 in., numerous, on glabrous 
ped., corymbs terminal, much branched, quite glabrous; cal. 
glabrous, segm. triangular; pet. broadly oval or rotundate, 
distant, clawed; fruit less than }in., capped with remains 
of cal.-teeth, glabrous, red-purple; seeds compressed, pointed 
at both ends. 


Upper montane zone, to highest elevations ; ratherrare. Fl. March, 
April; pinkish-white, slightly scented. 

Also in the Nilgiris, and mountains of Khasia, Martaban, and Java. 

The only Ceylon member of the apple tribe ; the little berry-like fruit 
has a harsh, bitter flavour. 


Crassulacee. 143 


MEV SAXNIPRAGACE Ze. 


HERBACEOUS, |. opp., without stip. fl. small, regular ; cal.- 
tube completely adnate to ov., segm. 5; pet. 5; stam. 5, 
epigynous; ov. completely inferior, 1-celled, with 2 pendulous 
placentas, ovules numerous, styles 2, distinct; fruit a capsule, 
dehiscing septicidally from apex; seeds numerous, very small, 
with endosperm. 

A single annual herb very badly represents this large Order. There is 


no species of Sazzfraga in our hills or in the Nilgiris, but 35 are given in 
FI. B. Ind. for the Himalaya range. 


VAHLIA, 7hun). 
For characters, see Order.—Sp. 33; 2 in Fv. B. Ind. 


V. oldenlandioides, ox. F7. Ind. ii. 89 (1832). 

Thw. Enum. 130, 418. C. P. 1686, 1687. 

Fl. B. Ind. ii. 399. Wight, Ic. t. 562. 

Annual herb with a long tap root and a short erect central 
stem and many spreading or prostrate ones, glabrous or very 
finely glandular-pubescent ; 1. small, #-14 in., sessile, linear- 
strap-shaped, tapering at both ends, acute, glabrous or 
puberulous ; fl. small, numerous, on short slender ped., usually 
2 at end of slender glandular-pubescent axillary and terminal 
peduncles as long as 1.; cal. finely glandular-pubescent, segm. 
short triangular, persistent, pet. oblong, about as long as cal.- 
segm.; capsule 4 in., subglobose, crowned with cal.-segm. ; 
seeds very minute and numerous. 
eed region ; apparently very rare. Jaffna, frequent. Fl. Feb., March; 
white. 

Dry or desert districts of India, Persia, and Trop. Africa. 

Easily mistaken for an Oldenlandia. 


[V. viscosa, Roxb., given for Ceylon in Fl. B. Ind., does not occur 
here (see Thw. Enum. 418).] 


XLVII.—CRASSULACEE. 
PERENNIAL herbs, |. opp., simple, succulent, without stip.; fl. 
regular, bisexual, in paniculate cymes; sep. 4, very slightly 
connate at base; pet. completely combined into a cor. with a 


144 Crassulacee. [Kalanchoe. 


long inflated tube, lobes 4, spreading ; stam. 8, inserted in 2 
rows on the tube of cor. (epzpetalous); carp. 4, slightly adnate 
to base of cor.-tube, distinct, each with a linear scale at the 
base, styles long; fruit of 4 follicles; seeds very numerous, 
with endosperm. 


RALANCHOE, 4aazis. 
For characters, see Order.—Sp. 25; 6 in FZ. B. Ind. 


Leaves oval, crenate. : ‘ : : : . I. K. FLORIBUNDA.. 
Leaves pinnatifid. . : : : 3 ; . 2. K. LACINIATA. 


1. K. floribunda, W. aud A. Prod. 359 (1834), var. glabras, 
Clarke in Fl. B. Ind. 1. c. 

Thw. Enum. 129. C. P. 540. 

Fl. B. Ind. ii. 414. 

A large perennial herb, 2-3 ft. high, perfectly glabrous. 
throughout, stem erect, not branched, hollow; 1. all opposite, 
articulated, very succulent, the lowest 4 in., gradually becoming” 
smaller upwards, the uppermost about 14in., spathulate-oval, 
rounded or obtuse at apex, gradually tapering to long base- 
(? petiole), coarsely serrate-crenate, pale yellowish green; fl. 
numerous, ped. a little shorter than sep., arranged in large, lax 
flat-topped trichotomous corymbose cymes, bracts linear; sep. 
narrowly lanceolate, very acuminate; cor. 2 in., tube inflated 
below, segm. }in., very acute; follicles 2in., erect, narrow,, 
beaked, at first enclosed in the white scarious persistent base 
of cor.-tube. 


Montane zone, on rocks; rare. Hantane; Pundalu-oya (E. Green) ;. 
Padanawella, Uva (Nock) ; face of the hill opposite Hakgala towards. 
Fort Macdonald (Nock). Fl. Nov.—Jan.; orange-yellow. 

Also in the Indian Peninsula. 


2. K. laciniata, DC. P/. Grasses, ili. t. 100 (1799). 

Moon Cat. 32. Thw. Enum. 129,417. C. P. 3836. 

Fl. B. Ind. ii. 415 (not given for Ceylon). Wight, Ic. t. 1158. 

A large succulent herb, perfectly glabrous throughout, 
stems very stout, slightly branched, pale glaucous green ;, 
]. numerous, large, very succulent, 3—4 in., on flattened fleshy 
petiole 1-13 in. long, very variable, the lowest deeply gashed, 
the middle ones more or less deeply pinnatifid, the uppermost 
with narrow almost cylindrical segm., all shining, pale 
glaucous-green ; fl. rather large, ped. glabrous about as long 
as cal., cymes large, lax, paniculate not flat-topped; sep. 
lanceolate, acute; cor.-tube 4+in., segm. }in., oval, acute. 


Lower montane zone; very rare. Matale East (Thwaites). There- 
are no specimens of C. P. 3836 in Hb. Perad. nor any drawing. Moon. 


Drosera. | Dyroseracee. IA5 


gives ‘Kandy’ as locality, where it was also found by H. de Alwis. 
There are plants in the Botanic Gardens (of unknown origin), and from 
these the above description is taken. Fl. March—May ; bright chrome 
yellow. 

Also in India, Malaya, and Trop. Africa. 


Bryophyllum calycinum, Salisb. (C. P. 541) is a common plant on 
bare rocky places throughout the low and lower montane country, and 
has all the look of a native. It occurs in similar places throughout the 
Tropics, but is believed to be originally from Trop. Africa. Mrs. Walker, 
in her tour in 1837 (Hook. Journ. Bot. ii. 242), says it had been then in- 
troduced only ‘a few years ago.’ It is called ‘Akkapana,’ or ‘ Rata- 
gowa’ by the Singhalese, and is recorded by Moon in 1824 (Cat. 32) as 
Calanchoe pinnata, with Colombo as the locality. 


XLVIII.—DROSERACE. 


SMALL perennial herbs, |. radical or cauline fringed with 
glandular tentacles, stip. adherent to petioles or absent; fl. 
small, regular, bisexual; cal. free, persistent, segm. 5, deep; 
pet 5, persistent; stam. 5, Ayfogynous; ov. superior, 1-celled 
with 3 or 5 parietal placenta, ovules very numerous, styles 3 
or 5; fruit a dry capsule, loculicidally 3- or 5-valved ; seeds 
very numerous, minute, black, with raised reticulations, with 
endosperm. 


The flowers have quite hypogynous pet. and stam., and the Order is 
often placed near Vzolacea. 


Of our three species one is montane, one confined to the low country, 
and one generally distributed. 
DROSERA, Z. 
For characters, see Order.—Sp. 100; 3 in FY, B. Ind. 


L. radical, round-spathulate 
L. cauline, linear. 
L. cauline, semilunate . 


. D. BURMANNI. 
PLO ENDING Ay 
DP ETAT AY 


GN = 


1.D. Burmanni, Vahl, Symb. Bol. iii. 50 (1794). Wata- 
essa, 5S. 

Herm. Mus. 18. Burm. Thes. 207. YD. rotundifolia, L., Sp. Pl. 281 
(part). Fl. Zeyl. n. 120. Moon Cat. 23. Thw. Enum. 21. C. P. 1080. 

Fl. B. Ind. ii. 424. Burm. Thes. t. 94, f. 2. Wight, Ill. t. 20, A.; Ic. t. 


944 (not good). 

A small stemless herb, |. all radical forming a close rosette 
on the ground, 4-4 in., round-spathulate, tapering into a flat 
petiole, upper surface with numerous gland-tipped tentacles, 

PART II. ie 


146 Droseracec. [Drosera 


_longest at the margin, stip. adnate to petiole, scarious, cut into 
long filiform lacinie; fl. about } in. on long slender ped.., 
secund, about 8-20 at end of erect slender, glabrous scapes 
4-6 in. high, 1-3 in number, from axils of inner leaves; cal. 
minutely papillose, segm. linear-oblong, obtuse; pet. rather 
longer than cal., spathulate; styles 5, undivided; capsule 
5-celled, seeds reticulate. 

Wet places throughout the island; common at all elevations. FI. 
March, April; white. 

Found throughout the Tropics of the Eastern Hemisphere. 

Closely allied to the common Sundew (DV. rotundifolia) of England. 


2. D. indica, L. Sf. P/. 282 (1753). Mandul-essa, S. 

Herm. Mus. 63.° Burm. Thes. 207" Bl Zeyl) n. 127. )MooniGatwzs" 
Thw. Enum. 21. C. P. 1088. 

Fl B. Ind: 1.424. Burm: Thes. t. 94, £1. Wight, Ul istacoumnc 

Stem weak, decumbent, slightly branched, glandular- 
pubescent; 1. numerous, cauline, alternate, 1}-13in., narrowly 
linear, strongly circinate in vernation, deflexed but again 
curved upwards at end when old, lower half (petiole) glabrous, 
rest copiously fringed with very fine gland-tipped tentacles ; 
fl. over 4in. on long ascending ped., 6-8 in rather stout 
axillary and terminal glandular racemes longer than 1.; cal.- 
segm. oblong-oval, subobtuse, entire, glandular-pubescent ;, 
pet. narrow, spathulate, obtuse; styles 3; seeds minute, ovoid, 
strongly ridged, with raised reticulation. 

Low country; rather rare. Pasdun Korale; Negombo; Kurunegala; 
Nilgala; Jaffna (dwarf form). Fl. July; white. 


Also in S. India, Burma, Malaya, and China, and in Trop. Australia. 
and Africa. 


3. D. peltata, Sm. in Willd. Sp. Pl. i. 1546 (1797). 

D. lunata, Ham., Thw. Enum. 22; Moon Cat. 23. C. P. 2553. 

Fl Bind, i. 424. Wight, [1 t. 20, & Dy Elcok: eeeianaass 
(D. lunata). ais 

Stem erect, 6-10 in. high, occasionally with a branch or 
two near the top, glabrous, arising from a globose, solid, 
smooth, red bulbil about 4in. diam. (a few rootlets come from 
base of stem above the bulbil); 1. numerous, alternate, rather 
distant along whole length of stem, petiole very slender, 
spreading, curved, blade vertical, peltately attached, semi- 
circular with the upper edge straight or slightly concave and 
prolonged at the corners into long acuminate horns giving the 
whole a semilunate outline, tentacles very numerous, the 
marginal ones long and spreading, the rest short; fl. nearly 
3 in., on long ascending, slightly pilose ped., about 8 in a raceme 
which is at first nodding, afterwards erect; cal.-segm. ovate, 


Serpicula.] Flaloragee. 147 


acute, laciniate or ciliate at margin, glabrous; pet. truncate 
much longer than sep.; ov. globose, furrowed, glabrous, with 
5 placentas, styles 3, much divided into blunt processes. 


Wet places in upper montane zone; rather common. FI. April, 
August-November ; white. 


Also in India, Malaya, and Australia. 

Turns black in drying, and makes a vinous-purple stain on the paper 

When old the plant has exhausted the bulbil, which then become 
shrunken, a new one being formed at its side. 


XLIX.-HALORAGE. 


SMALL aquatic or semi-aquatic herbs, |. opp. or alt. or 
verticillate, fl. minute, unisexual, moncecious, axillary; male 
fl.:—cal. 4-fid or 0, pet. 4 or 0, stam. I or 4-8; fem. fl. :—cal.- 
tube adnate to ov. segm. 4 or 0, pet. 0, ov. inferior, I- 2- or 
4-celled, ovules 4, pendulous, styles or stigmas 2 or 4; fruit 
minute, indehiscent or separating into constituent carp., seeds 
1-4, embryo in axis of fleshy endosperm. 


Pet. of male fl. 4 

Pet. o. 
Submerged water plant 
Creeping herb 


I. SERPICULA. 


2. MYRIOPHYLLUM. 
3. CALLITRICHE. 


Myriophyllum is found in ie ae tas country, Serpzcula and Callz- 
triche in the montane zone. 

1. SERPICULA, Z. 

Small decumbent much-branched herbs, 1. opp. or alt., fl. 
minute, unisexual, moncecious, axillary, male fl. stalked, female 
sessile; male fl.:—cal. with short tube, segm. 4, pet. 4, stam. 8, 
ov. 0; fem. fl.:—cal.-tube adnate to ov. segm. 4, pet. and stam. 
oO, ov. inferior, 1-celled, with 4 pendulous ovules, styles 4, short, 
with large rayed stigmas; fruit minute, indehiscent, hard, 
t-seeded.—Sp. 4; 2 in #2. B. Ind. 


Fruit glabrous, strongly ribbed and tubercled_ . Le Oa ZEVUANICA, 
Fruit pubescent, not ribbed or tubercled 2. S. HIRSUTA. 


1. S. zeylanica, Arn. in F/. B. Ind. ii. 431 (1878). 

Haloragis oligantha, Arn, Pug. 18 (non Wight). 
Thw. Enum. 123. C. P. 146, 451, 447. 

Fl. B. Ind. ii. 431. 


Stems sometimes slightly woody at base, much branched, 
decumbent, ascending or even erect, stout, glabrous; 1. very 


S. indica (part), 


148 FI alorag Ce. [Wyriophyllum. 


numerous, close, usually alt., articulated with stem and leaving 
a prominent scar, usually narrowly linear, acute, entire or 
with a few large sharp teeth in upper part, glabrous, some- 
times cuneate-obovate, with minute teeth and ciliate; ped. of 
male fl. not longer than 1., very slender, glabrous; fruit minute, 
oblong-oval, narrowed above, glabrous, strongly 8-ribbed, 
with the ribs tuberculated. 


Var. 6, minor, Clarke in Fl. B. Ind. 1. c. 

A minute plant, 13-3 in., with filiform stems, 1. opposite, 
very small. 

Upper montane zone ; very rare. On Adam’s Peak, where it is abun- 
dant near the base of the uppermost cone; Maturata. Var. 6, Watte- 
kelle, 1864. FI. March, &c. 

Endemic. 

Thwaites did not distinguish this from the next, and called the com- 
bined plants S. zzdica,; and this is perhaps the best treatment. C. P. 
451, with the fruit of S. zeylanzca, has the leaves of S. Azvsuta, and looks 
quite intermediate ; var. 8 seems to be merely a dwarf state of this form. 


2. S. hirsuta, W. and A. Prod. 338 (1834). 

S. brevipes, W. and A., Arn. Pug. 18. S. zzdica, Thw. Enum. 123 
(part); Trim. Syst. Cat. 31. C.P. 1545. 

Fl. B. Ind. ii. 431 (S. zza@zca). Wight, Ic. t. toot. 

Stems prostrate, wide-creeping, often very long, much 
branched, rooting at the nodes, slender, slightly pilose, red ; 1. 
numerous, opp. or alt., nearly sessile, usually under 4 in., 
tapering to cuneate base, oval or obovate-oval, with a few 
large teeth in upper half, glabrous on both sides, ciliate on 
margin, often red beneath; male fl. few, on very slender, 
pubescent ped. longer than 1. (4-3 in.), fem. fl. sessile, in 
axillary clusters; male fl.:—cal.-segm. acute, pet. oblong, 
concave, pubescent outside, soon falling, anth. large, cream- 
coloured ; fem. fl.:—cal.-segm. triangular, pubescent, stigmas 
spreading ; fruit very minute, broadly ovoid or subglobular, 
not ribbed, pubescent. 

Moist open ground, especially near streams in the montane zone above 
4ooo ft.; very common. Fl.all the year; pinkish. 

Also in the mountains of S. India. 

As S. hirsuta and S. brevipes of Wight and Arnott are not distinguish- 
able as species, and as the names are of the same date, I adopt that most 
suitable to our form. 


2. MYRIOPHYLLUM, Z. 


Submerged aquatic, |. verticillate, fl. very small, axillary, 
unisexual, moncecious (rarely bisexual); male fl.:—cal. minute 
4-fid, pet. 4, stam. 4-8; fem. fl.:—cal.-tube connate with ov., 


Callitriche.] flaloragec. I49 


segm. 4, very small, pet. 0, ov. quite inferior, 4-celled, stigmas 
4, plumose ; fr. of 4 nearly distinct carp—Sp. 15; 5 in FZ B. 
Ind. 

M. indicum, /V7//d. Sp. Pi. iv. 407 (1805). 

Moon Cat. 64. Haloragis oligantha, Wight, Ic. iii. pt. 4, 3. Thw. 
num 123. C.P: 1549. 

Fl. B. Ind. ii. 433. Wight, Ic. t. 1061. 

A submerged water-herb, stems long, much branched, stout, 
glabrous, the upper portion often floating on the surface; 1. 
very numerous, in whorls of 2-5 (usually 4), $-#in., the 
uppermost ones linear, attenuate to base, acute, nearly entire, 
glabrous, the intermediate ones more or less toothed or 
pectinate, the lowest ones entirely cut into narrow filiform 
segm. looking like branched rootlets; fl. sessile, male in the 
upper axils, female in the lower ones, and a few bisexual ones 
between them; stam. generally 6, as long as pet.; fruit 
depressed of 4 nearly globular lobes (carp.), glabrous, bluntly 
muriculate. 


Canals, ponds, and tanks in the dry region; rathercommon. Jafina; 
Bintenne; Chilaw; Kalawewa. FI. all the year. 

Also in the Carnatic, Bengal, and Assam. 

I have never seen anything like a spicate arrangement of the inflo- 
rescence in our plant. When left growing on mud uncovered by the 
water the much-divided 1. are not produced. Wight’s figure above quoted 
is good for this form, but Clarke (in F]. B. Ind.) refers it, and also Wight, 
Herb. n. 942, to MW. zntermedium DC., which is, however, probably not 
distinguishable as a species. 

Willdenow, in the original description, gives Ceylon as a locality. 


3. CALLITRICHE, L. 


Seema annual herb, |. opp., ff. minute; unisexual, 
moncecious; cal. and pet. o (achlamydeous), male fl. of 1 
stam., female fl. of a 4-celled ov. with 1 pendulous ovule in each 
cell, styles 2 long; fruit indehiscent of 4 carp. combined in 2 
pair, ultimately separating.—Sp. about 10; 2 in FZ. B. Lud. 


C. stagnalis, Scop. Fl. Carniol. ed. 2, ii. 251 (1772). 

C. verna, Thw. Enum. 290 (non L.). C. P. 303, 3636. 

Fl. B. Ind. ii. 434. Wight, Ic. t. 1947 (C. Wightiana). 

A very small annual flaccid herb, with slender prostrate 
glabrous stems rooting at the nodes; |. small, opposite, 4-—# in., 
spathulate-oval, tapering to petiole, obtuse, glabrous; fl. 
axillary sessile, a male and a female usually in each axil, 
bracts large, boat-shaped, white, soon falling ; fruit very small, 
compressed, circular, glabrous, each carp. with a narrow wing 
round the edge. 


150 Rhizophoracee. [Rhizophora. 


Upper montane zone ; a common weed on damp ground or mud. FI. 
all the year. 

Found in most parts of the Old World, temperate and Tropical. The 
Ceylon plant is somewhat smaller than the common English form, and 
has a rather narrower wing to the fruit. 


L.—RHIZOPHORACE/E. 


TREES or shrubs, |. opp., with large interpetiolar stip. (alt. and 
without stip. in Azzsophyllea), entire, glabrous; fl. regular, 
bisexual; cal. more or less adnate to ov. (free in Wezhea), 
segm. 4, 5, or 8-14, valvate; pet. same number as cal.-segm., 
often cut or laciniate; stam. double number of pet. (in Wezhea 
more) perigynous or epigynous (hypogynous in Wezhea); disk 
annular and lobed or absent; ov. inferior or half-inferior 
(superior in Wezhea), 2-4-celled, with 2 (1 in Azzsophyllea) 
pendulous ovules in each cell; fruit leathery, indehiscent, 
1-seeded (dehiscent and with 4-6 seeds in Wezhea); seed 
usually with a large macropodous embryo without endosperm, 
sometimes with a small embryo in axis of copious endosperm. 


L. opp., with interpetiolar stip. 
Seed without endosperm, embryo germinating in 
the fruit (Rhizophorea). 


Pet. 4, ov. = 4-inferior I. RHIZOPHORA. 

Pet. 5, ov. $-inferior 2. CERIOPS. 

Pet. 8-14, a, quite inferior s 3. BRUGUIERIA. 
Seed with fleshy endospenn (Legnotidea), 

Ov. inferior. 4. CARALLIA. 

Ov. superior 5. WEIHEA. 


L. alt., without stip. Seed without endosperm (Ani- 
"soph ry lle@) : ‘ : : . 6. ANISOPHYLLEA. 


The tribe AAzzophoree are the true Mangroves, and our 6 species are 
found in suitable places all round the coast of the island. Of the rest, 
the endo Carallia calycina is nearly restricted to the lower montane 


zone; Wezhea and Anisophyllea are low-country plants, and the latter is 
Berne to the moist region. 


1. RHIZOPHORA, Z. 


Moderate-sized trees, sending down numerous aérial roots 
from the stem and branches, |. opp. entire, thick, with large 
interpetiolar deciduous stip. enclosing the buds, fl. in pairs, 
each supported by 2 hard thick fused bracts; cal. very deeply 


Rhizophora.] Rhizophoracee. 151 


divided, segm. 4, persistent, enlarged in fruit; pet. 4; stam. 8 
or II (or 12), anth. nearly sessile, with numerous cells; ov. 
half-inferior, 2-celled, with 2 ovules in each cell; fruit conical, 
leathery, indehiscent, 1-celled, 1-seeded, perforated at apex by 
the large cylindrical radicle of the germinating seed.—Sp. 5 ; 
2 in Fl. B. Ind. 


Fl. on short ped., peduncle 13 in. . : . . I. R. MUCRONATA. 
Fl. sessile, peduncle j in.. : : : : . 2. R. CANDELARIA. 


1. R. mucronata, Lam. Encyc. vi. 169 (1804). WMadol, S. 
KEandal, 7. 

R. Candel, Moon Cat. 36 (non L.). Arn. in Ann. Nat. Hist. i. 362. 
Wight, Ill. i. 209. Thw. Enum. 120. C. P. 1960. 

Fl. B. Ind. ii. 435. Wight, Ic. t. 238. 

A moderate-sized tree with a spreading head, branchlets 
thick, marked with close scars of fallen J. and stip., young 
parts glabrous, purple; 1. 44-5 in., oval, acute at both ends 
and with a strong brown point nearly } in. long at apex, 
perfectly glabrous, thick, bright green, pale beneath and dotted 
with minute red spots, petiole 4—#in., stout, stip. 2 in., glabrous, 
soon falling; fl. moderate-sized, on short, very thick ped., 2 
pairs together at end of stout somewhat drooping peduncles 
about 14 in. long from axils of 1. of same year; cal.-segm. 
about 4 in., oblong-lanceolate, thick, glabrous, keeled within ; 
pet. about as long as sep., narrow, obtuse, curved, thick, 
induplicate, hairy within; stam. 8, anth. linear, very acute; 
fruit 14-2 in., ovate-conical, pendulous, slightly rough, dark 
brown ; embryo attaining 18in.,or even more, before falling 
from fruit. 

Salt-water estuaries and backwaters round the whole coast. FI. 
Aug., Sept.; white, sep. pale yellow. 

Seashores throughout the Eastern Tropics. 

A good picture of the habit of this mangrove is given in t. i. of 
Karsten’s ‘ Mangrove-Vegetation’ (Biblioth. Botan. pt. xxi. 1891). Moon 
calls this ‘Ela-Kadol, S. In drying it turns a dark purplish-brown. 
Heart-wood reddish, very hard. The fishermen soak their nets in the sap 
to make them durable. 


2,.R. Candelaria, DC. Prod. iii. 32 (1828). Kadol, S. 
Kandal, 7. 

?. Mangle, Moon Cat. 36 (L. in part). 2. conjugata, Arn. 1. c. 363; 
‘Thw. Enum. 120 (non L.). C. P. 1968. 

Fl. B. Ind. ii. 436 (2. conjugata). Rheede, Hort. Malab. vi. t. 34. 
mare, 1. c. t. 2, £..1 

A rather smaller tree than &. mucronata, but otherwise 
very similar, 1. 5-6 in., lanceolate-oval, acute at base, tapering 
to acute apex and very shortly mucronate, paler and slightly 


152 Rhizophoracez. [Cerdops.. 


dotted beneath, petiole a7! in., stip. 2-3 in.; fl. sessile 
peduncles wee) short, about 4 in., stout, from axils of fallen 1.; 
cal.-segm. 2—# in., ovate-oblong, acute, striate externally ; pet. 
linear, flat, ‘glabrous, soon falling; stam. 11 (or 12); fruit about 
it Was? ‘otherwise as in the last. 

With A. mucronata, but rather less common. FI. Jan., Feb.; white, 
sep. pale green, brown outside. 

All Tropical seashores of Asia and Africa. 

By common consent this has been accepted as R&. conjugata of 
Linnzeus, but it is not the plant of his Fl. Zeyl. n. 181 (see Bruguzera) nor 
of his Herbarium. He included the present species under A. Manele, 
which principally referred to an American one, to which the name is now 
restricted. Rheede’s plate above quoted certainly represents the present 
species rather than 2. mucronata, for which it is quoted in Fl. B. Ind. 


Though easily distinguished when living, the confusion in books between 
the two is very great. 


These two Ahzzophoras form, with Bruguzera gymunorhiza, the bulk 
of the mangrove forests of the coast. The bark of all is used for tanning” 
in Ceylon, and is exported to India in small quantity. 


2, CERIOPS, Arn. 


Small trees or shrubs, sending down aérial roots as in: 
Rhizophora, and with |. and stip. of that genus; cal. deeply 
divided, persistent, segm. 5; pet. 5, auaieinels: stam. I0O,. 
inserted between the lobes of an annular disk; ov. fully half-. 
inferior, 3-celled, with 2 ovules in each cell, style simple, fruit. 
as in Rhzzophora, but the radicle of the germinating embryo- 
sharply angled.—_Sp. 3; 2 in FZ B. Ind. 


Pet. flat, glabrous 5 : A . I. C. CANDOLLEANA. 
Pet. concave, ciliate towards apex : ‘ . 2. C. ROXBURGHIANA. 


1. ©. Candolleana, Arz. iz Ann. Nat. Hist. i. 364 (1838). 
Chirukandal, 7. 

Thw. Enum. 120. C. P. 1966. 

HB inden 436. aNWichtele. t4210: 

A shrub or small tree with numerous opposite branches,,. 
twigs marked with scars of |. and stip., young parts glabrous 3. 
1, 2-3 in., obovate-oblong, tapering to base, rounded at apex, 
glabrous, thick, pale green; fl. small, sessile in clusters of 2. 
(or 3) arranged in small lax umbels on stout rather deflexed 
glabrous peduncles from axils of upper |. }-4 in. long; cal. 
glabrous, seem. oblong, subacute; pet. shorter than cal.-segm. 
flat, emarginate, tipped with 2 or 3 minute clavate processes,,. 
glabrous; fruit pendulous, under 1 in., cylindrical, tapering,, 
rough, reddish-brown; embryo attaining 8-10 in. before: 
falling. 


Bruguiera.] Lehizophoracee. 153 


Mangrove forests on the coast, common; Negombo; Chilaw; Kalpitiya; 
Trincomalie. Fl. Sept.; pet. white, sep. green. 
Tropical seashores of the Old World. 


2. ©. Roxburghiana, Arz. in Ann. Nat. Hist. i. 363 (1838). 

BSB! Ing. 1.436. WKarsten,.l.c: 't./3, f. 1. 

Scarcely differs from the last in habit and foliage, but fl. in 
a more dense capitate cyme; cal.-segm. subovate, rather 
obtusc; pet. concave, setose-ciliate towards apex; stam. often 
12; fruit less tapering and embryo shorter. 

In the tidal estuary of the Mahawéli River at Kottiyar, 1890 (Karsten). 

Also in the Sunderbuns, Andaman Islands, and coast of Burma. 

Prof. Karsten informs me that he found this little tree abundantly in 


the above locality with the other species. I have seen no specimens. 
The two appear very closely allied. 


[Kandelia Rheedez, W. and A., is given for Ceylon in FI. B. Ind. on 
the faith of a doubtful specimen in Herb. Kew. It is figured in Rheede, 
Hort. Malab. vi. t. 35 and in Wight, Ill. i. t. 89, and may possibly 
occur ; but I have seen no specimens. | 


3. BRUGUIERA, Lam. 


Trees or shrubs with habit and 1. of RAzzophora, cal.-tube 
connate with ov. below and free above it for some distance, 
deeply cut into 8-14 persistent segm.; pet. 8-14, concave, 
deeply bilobed, with terminal appendages; stam. 16-28, two 
enclosed in each pet.; ov. wholly inferior, 2—3-celled, with 2 
ovules in each cell; fruit crowned by the persistent cal.-limb; 
radicle of germinating embryo ace or nearly so.—Sp. 8; 
5 in Fl. B. Ind. 


Fl. solitary ; cal.-segm. 10-14. 5 : . I. B. GYMNORHIZA. 
Fl. in threes ; cal.-segm.8 . ; : . 2. B. CARYOPHYLLOIDES. 


1. B. gymnorhiza, Lam. Encycl. iv. 696 (1797). 

Fl. Zeyl. n. 181. Rhizophora conjugata, L. Sp. Pl. 443. 2. gymno- 
rhiza, L., Moon Cat. 36. Thw. Enum. 120 (including &. Rheedzz. Miq.). 
CrP. 3611, 3612. 

Fl. B. Ind. ii. 437. Rheede, Hort. Malab. vi. tt. 31, 32. Wight, Ic. t. 
Zag imatsten, lc. t. 2; f, 2; 

A tree, young branches smooth, with pale bark and 
prominent scars of |. and stip.; 1. 34-5 in., oval, tapering to 
base, slightly acuminate, acute, not mucronate, glabrous, rather 
thick, bright green, paler beneath, with the midrib very 
peomitiertt, petiole 4-1 in.; fl. rather large, solitary, drooping, 
peduncle 4 in., curved, from axils of upper 1.; cal, coriaceous, 
glabrous, tube striate, segm. 10-14 (usually rt), 4—# in., linear, 
very acute, erect; pet. 10-14, very deeply bifid, hairy at the 


154 Lehizophoracec. [Carallia. 


base, with 2 bristles at the top of each lobe and I in the 
notch, stam. 20-28 (usually 22); ov. quite inferior, 3-celled, 
style long ; fruit about 1 in., obconical, crowned by the some- 
what enlarged cal.-segm., smooth, scarlet ; embryo attaining 
Q in. before falling, radicle stout, fusiform, blunt, very slightly 
angled. 

Mangrove forests; common all round the coast. Fl. December, Jan.; 
cal. orange-yellow. 

Tropical shores throughout the Old World. 

Heart-wood red, very hard and strong, but of small size. 

There is no specimen in Hermann’s Herb., but his drawing is unmis- 
takably this species, and it is the whole foundation for} Linnzus’s Rhzzo- 
phora conjugata, which name has been since always applied to another 
plant, R. Candelaria, DC.,to which this bears a strong resemblance in foliage. 

An excellent figure of the curious interlacement of the aérial roots of 
this species is given by Karsten (I. c. t. 10, f& 121). 


2. B. caryophylloides, &/. Exum. Plant. Jav. i. 93 (1830). 

Kanilia caryophylloides, B\., Thw. Enum. 120. C. P. 1965. 

Fl. B. Ind. ii. 438. Rheede, Hort. Malab. vi. t. 33. 

A bush or small tree branched from the base; 1. 3-4 in., 
readily disarticulating, lanceolate-oval, tapering below, shortly 
acuminate, subacute, glabrous, rather thin, midrib very pro- 
minent beneath, petiole #-1 in.; fl. small, on very short ped., 
three together at end of slender peduncle about # in. long 
from axils of upper 1.; cal. glabrous, segm. usually 8, linear, 
subacute; pet. 8, erect, concave, hairy outside, deeply bifid, 
with several long bristly hairs at end of each lobe; stam. 16; 
ov. 2-celled ; fruit small, $in., oblong-ovoid, smooth, capped 
with reflexed cal-segm. about half its length; embryo 
attaining 6 in. before falling, slender, cylindrical. - 

Mangrove forests ; rare. Trincomalie (Gardner); Kalpitiya. FI. 
July ; white. 

Also in S. India and Malaya. 


4. CARALLIA,* foxd. 


Trees, 1. opp. with large interpetiolar stip., fl. in axillary 
cymes; cal.-tube campanulate, connate with ov., segm. 4-8 ; 
pet. 4-8, cut or fimbriate; stam. 8-16, inserted (with pet.) 
outside a prominent annular crenulated disk; ov. inferior, 
4-celled, with 2 ovules in each cell, stigma 4-cleft; fruit fleshy, 
indehiscent, 1-seeded ; seed globose-reniform, with a cylindrical, 
curved, green embryo in axis of copious endosperm.—Sp. 7; 
ee iba IHG Ib Mao: 


* Carallia, from the Telugu name, Karalli. 


Carallia.] Rhizophoracee. 155 


Fl. sessile ; pet. 7 0r8 . ‘ : ; : . I. C. INTEGERRIMA. 
Fl. stalked ; pet.4 . : ; ‘ : : . 2. C. CALYCINA. 


1. GC. integerrima, DC. Prod. ili. 33 (1828). Dawata, S. 

C. zeylanica, Arn. Pug. 16. Thw. Enum. 120. C. P. 1963, 1964. 

Fl. B. Ind. ii. 439. Wight, Ill. t. 90 (C. zeylanica). Bedd. Fl. Sylv. 
t. 193. 

A rather large tree with erect trunk and wide-spreading 
head, bark rather smooth, grey, young parts glabrous; 1. on 
very short petioles, 34-43 in., obovate-oval, tapering to base, 
rounded or very bluntly acuminate at apex, entire or obscurely 
serrulate at top, margin somewhat revolute, glabrous and 
somewhat shining on both sides, very dark dull green above, 
pale and with obscure glandular dots beneath, rather thick, 
veins inconspicuous, stip. large, #in., interpetiolar, enclosing 
terminal bud, caducous; fl. small, sessile, few, in small heads, 
on stout branches of axillary trichotomous cymes much 
shorter than 1.; cal. glabrous, segm. 7 or 8, lanceolate, acute; 
pet. small, 7 or 8, clawed, roundish, coarsely undulate-dentate 
at margin; stam. twice as many as pet., scarcely longer than 
cal.; style stout; fruit small, about 4 in., berry-like, smooth, 
red ; seed with thick orange testa, lobulate. 

Moist low country ; rather common. FI. Feb.; creamy-white. 

Also in India, Burma, Malaya, China, and Trop. Australia. 

Heart-wood very hard, pale orange-coloured, tough, shining, with 
‘conspicuous medullary rays, not durable. A gum exudes from the fl.-buds 
and glues the fl. together in the heads. This tree very obviously shows 
its relationship to the mangroves by the large tufts of aérial roots it 
sends forth from the trunk and branches: these are admirably seen in 
many of the old wild trees in Victoria Park, Colombo. 

The leaves show on their under surface a line or fold at some distance 
on either side of the midrib precisely similar to those in Erythroxylon 
Coca and due to the same cause, the mode of packing in the bud. 


2. ©. calycina, Benth. in Journ. Linn. Soc. iii. 75 (1859). UWh- 
bériya, S. [PLATE XXXVI.] 

Thw. Enum. 121. C. P. 3458, 2588. 

Fl. B. Ind. ii. 439. 

A large tree, with thin rough grey bark, |. on very short 
petioles, 3-3} in., broadly oval, tapering at base, shortly 
acuminate and twisted at apex, entire, glabrous, stiff, bright 
green, gland-dotted beneath, lat. veins numerous, parallel, 
conspicuous; fl. few, nearly 4 in., on stout ped., in scanty erect 
dichotomous axillary cymes half as long as 1., bracts rounded 
mucronate, caducous; cal. glabrous, segm. 4, acute, as long as 
tube, pet. 4, clawed, incised-fimbriate; stam. 8; fruit about 3 in., 
pyriform, crowned with erect connivent cal.-segm., red, 
glabrous. 


156 Lhizophoracee. [ Wethea.. 


Forests in the moist region; rare in the low country, and only re- 
corded from the Singhe Rajah Forest ; common in the lower montane. 
zone and even extending above 5000 ft. Fl. Jan.—April; white. 

Endemic. 

The montane form has the leaves more coriaceous, smaller, 3 in., 
and proportionately broader ; but does not deserve to be made a variety. 

Wood red, rather heavy, hard but liable to split, medullary rays- 
extremely wide, giving a very characteristic appearance. This beautiful 
wood is often called ‘ Dawata’ by the carpenters. 


5. WEIHEA, Sireng. 


A large shrub, |. opp., with interpetiolar stip., fl. axillary ; 
cal. free, cut nearly to the base into 5 segm.; pet. 5, Zypogynous,. 
much cut; stam. about 30, Zypogynous,; ov. quite superior, 
- 3-celled with 2 pendulous ovules in each cell, style long, 
persistent, stigma 3-lobed; fruit a fleshy capsule, 3-celled, 
ultimately dehiscent septicidally; seeds 4-6, with a small aril, 
embryo straight in axis of endosperm.—Sp. 5; 1 in FZ. B. Lud. 

The structure of the flower is completely thalamifloral, hence Gardner 
and others have referred it to 7z/zacee (Eleocarpee) and Moon to £/@o- 


carpus itself (see also Arnott in Ann. Nat. Hist. ili. 23) which its pet. so- 
much resemble. 


W. zeylanica, Baill. Adans. iii. 38 (1862). Kannu, 7. Pana, S. 

Eleocarpus integrifolius, Moon Cat. 42 (non Lam.). <Azstrutheria* 
geylanica, Gardn. in Calc. Journ. Nat. Hist. vi. 344. Thw. Enum. 121. 
CoP SuLiG: 


Fl. B. Ind.i1. 441. Cale. Journ. Nat: Hist. vi. t.4. -Bedd. Flisylve 
t. 194. 

A large shrub, very much branched from the base, young 
parts glabrous; |. 12-3 in. (or more) on short erect curved. 
petioles, broadly oval, rounded at base, obtusely acuminate,, 
entire, glabrous, shining above, bright apple green, rather 
paler beneath, stip. interpetiolar, small, lanceolate, acute,. 
ciliate ; fl. # in., on short curved ped., solitary or rarely 2 or 3. 
in axils, buds small, globular, completely enclosed for a long 
period before expansion in 2 thick, cup-shaped, rounded, 
ciliate bracts; cal.-segm. oblong-lanceolate, acute, densely 
silky outside, persistent, ultimately deflexed ; pet. with a long 
claw, very deeply cut into linear segm., soon falling; fruit. 
nearly globose, tipped with long style, at length dehiscent by 
3 valves. 


Low country, principally in the dry districts, where it is common ;; 


* In honour of Philip Anstruther, Colonial Secretary of Ceylon from. 
1846 to 1850, a gentleman much interested in Agriculture, Horticulture, 
and Botany. 


Anisophyllea. Rhizophoracec. 157 


rarer in the moist region, e.g. Kalutara; Galle; Kandy. Fl. July- 
September ; white. 

Also in Southern India. 

Specimens of this are in Wight’s Herb. labelled ‘ Ceylon, 1796, Klein,’ 
‘but there are none in Rottler’s collection at Kew. 

A very remarkable plant; the stip. are like those of Rudzacea. 


6. ANISOPHYLLEA, 37. 

Tree, 1. alt., dimorphic, without stip., fl, small, in supra- 
axillary racemes; cal.-tube adnate to ov.,segm. 4; pet. 4, 
laciniate ; stam. 8, epigynous; ov. inferior, 4-celled, with a 
solitary pendulous ovule in each cell; fruit coriaceous, inde- 
hiscent, I-seeded; seed without endosperm, embryo large, 
without cotyledons.—Sp. 5; 4in FZ. B. Ind. 

A very singular plant; the embryo, as remarked by Thwaites, is 


precisely like that of Barringtonza and Careya, along with which genera 
he places it. 


A. zeylanica, Benth. in Niger Flora, 575 (1849). Wéli-penna, 
‘Weli-piyana, S. 

Tetracrypta cinnamomoides, Gardn. and Champ. in Kew Journ. Bot. 1. 
314, and v. 378. 

Thw. Enum. 119. C. P. 2205. 

Fl. B. Ind. ii. 442. Kew Journ. Bot. v. t.5. Bedd. Fl. Sylv. t. 195. 

A moderate-sized tree, with dark brown bark, branches 
horizontal or drooping, young twigs pubescent; 1. of two kinds, 
normal large and persistent ones, and very small usually 
deciduous ones on the younger branches, normal |. on a very 
short, broad, flattened petiole, 4-5 in., ovate-oblong, rounded 
or tapering to unequal base, caudate-acuminate, obtuse or 
subacute, entire, glabrous, strongly 5-nerved from base with 
the nerves very prominent beneath, dark green, paler beneath, 
the smaller |. alternating with the normal ones, sessile, 
4-2 in., lanceolate, acute, green, stipuliform ; fl. small, on short 
slender pilose ped., arranged in 2 or 3 very short racemes, 
superposed and supra-axillary; cal.-tube pubescent, deeply 
8-furrowed, segm. triangular; pet. cuneate, deeply cut into 5 
linear laciniz; fruit about 1in., fusiform, crowned with per- 
sistent cal.-segm., deeply 8-furrowed, leathery, glabrous. 

Moist low country, up to 3000 ft., rather common. Moon’s specimens 
are from Kalutara. 

Fl. April-October ; greenish-white. 

Endemic. 

When young the branches are whorled, and spread horizontally. The 
larger leaves have much resemblance to Cinnamon, but are often very 
unequal-sided. Wood greyish-yellow, rather hard, with very conspicuous 


medullary rays and pores, the former connected by transverse bars of 
soft tissue. 


15 8 Combretacee. [ Terminalia. 


LI.—COMBRETACE. 


TREES or shrubs, sometimes semi-scandent, |. alt. or opp., 
simple, without stip.; fl. small, regular, bisexual or poly- 
gamous or unisexual ; cal.-tube adnate to ov. and prolonged 
above it, segm. 4-7 (2 in Gyvocarpus), pet. 4 or 5, or more 
usually 0; stam. 8 or 10 (4 in Gyrocarpus), alternating with 4 
staminodes), inserted on cal.-limb; ov. inferior, 1-celled, with 
2-5 (1 in Gyrocarpus) pendulous ovules; fruit indehiscent, 
dry, fleshy or drupe-like; seed solitary, cotyledons large, 
plaited or convolute, no endosperm. 


Anth. opening by slits 


Pet. o. 
Fl. in spikes, 5 3 : : : . I. TERMINALIA. 
Fl. in globular heads . s 5 : . 2. ANOGEISSUS. 
Pet. 4 or 5. 
L. alternate. : : ; F . 3. LUMNITZERA. 
L. opposite . 4. COMBRETUM. 
Anth. opening by recurved valves (Gyrocarpee) . 5. GYROCARPUS. 


Eleven species, none extending into the montane zone. Axogetssus 
and Gyrocarpus are dry-country trees, Terminalia belerica and T. parut- 
Jjiora confined to the moist region, ‘and Lummnitzera an munelecaa of 
mangrove swamps. 


I. TERMINALIA, Z. 


Trees, |. alt. or opp., without stip., fl. polygamous, sessile, 
in axillary or terminal paniculate spikes; cal.-tube adnate to 
ov., limb free, campanulate, cut into 4 or 5 triangular segm.; 
pet. 0; stam. 8 or 10, 5 often longer, inserted on cal.-tube 
outside an annular epigynous hairy disk; ov. inferior, I- 
celled, with 2 or 3 pendulous ovules; fruit an indehiscent 
drupe, flesh hard or fibrous, stone woody or bony; seed 
solitary, without endosperm.—Sp. 80; 12 in FZ. B. Ind. 

Drupe not angled or winged 

Stone of drupe very obscurely angled 
L. glabrous, drupe pubescent . T. BELERICA. 
L. pubescent, drupe glabrous . 0. CHEBULA. 


I 
Stone of drupe 5-rayed. , 3. T. PARVIFLORA. 
Drupe with 5 broad wings (Pentaptera) 4. IT. GLABRA. 


Terminalia. Combretacee. 159 


I. T. belerica,* Roxb. P?/. Cor. ii. 54 (1798). Bulu, S. Tanti, 7. 

Myrobalanus bellerica, Gaertn. Fruct. ii. 90. Moon Cat. 73. Thw. 
Enum. 103. C. P. 1605. 

Fl. B. Ind. ii. 445. Wight, Ill. t.91. Bedd. FI. Sylv. t. 19. 


A large tree, with straight, often buttressed trunk, and 
long horizontal branches, bark vertically furrowed, brown, 
young parts glabrous; |. alternate, closely placed at ends of 
branchlets, 4-6 in., obovate-oval, tapering to base, rounded or 
very shortly acuminate at apex, entire, with a pellucid margin, 
glabrous and shining on both sides, paler beneath, stiff, 
venation finely reticulate, pellucid, main veins prominent 
beneath, petiole 14-2 in., cylindrical; fl. small, the male 
shortly pedicellate, the bisexual nearly sessile, numerous in 
axillary spikes or racemes shorter than l., often crowded at 
ends of branchlets without |. so as to form a terminal panicle; 
peduncles pubescent, bracts minute, caducous; cal. tomentose 
outside, the limb with long crisp hair within; drupe 14-1} in., 
ovoid, often somewhat pyriform, narrowed at base, irregular 
on surface, but not at all angled, covered with a fine, close, 
brownish-yellow tomentum, stone large, woody, very obscurely 
angled, with a large seed-cavity. 

Low country, moist and intermediate regions, not common. Kurune- 
gala (Gardner). Often planted. Fl. February; pale greenish-yellow, 
strongly scented. 

Also in India and Malaya. 

The Ceylon tree is the variety /aurinotdes, Mig. usually, but it varies 
considerably in the form of the leaves. It never possesses the two glands 
at the top of the petiole figured and described by Roxburgh. 

The fruits are the ‘ Beleric Myrobalans’ of the old pharmacists, and 


are much used in native medicine. Wood greyish-yellow, hard but not 
durable, moderately heavy. 


T. Catappa, L., the Kottamba of the Singhalese and the ‘country 
almond’ of the English, is largely planted throughout the low country. 
It is native to Malaya, but is grownin most Tropical countries. Hermann 
does not seem to have noticed it in Ceylon. 


2. T. chebula,* (Retz. Obs. Bot. v. 31 (1789). Aralu, S. Ka- 
dukkay, 7. 

Moon Cat. 73. Thw. Enum. 103. C. P. 1604. 

Fl. B. Ind. ii. 246. Bedd. FI. Sylv. t. 27. 

A moderate-sized tree, trunk crooked, with many spreading 
branches, bark very thick, grey-brown, young parts pubescent; 
l. alt., 3-5 in., broadly oblong-oval, rounded or cordate at 
base, very obtuse at apex, pubescent on both sides especially 


* The names belerica and chebula are Latinisations of the Arabic 
names for these two kinds of myrobalans by which they were first known 
in Europe. 


160 Combretacece. [ Zerminalia. 


when young, thick and rigid; petiole 4—?in., pubescent, with 
2 prominent glands at the top just beneath |.; fl. as in the 
last, but spikes all terminal, bracts linear, hairy, conspicuous 
in bud; cal.-limb glabrous outside, with long hair within; 
drupe pendulous, about 1$in., broadly ovoid, glabrous, not at 
all angled, yellowish-green, stone oblong, bony, very thick, 
obscurely angled, seed cavity very small. 

Low country up to 2500 ft., principally in the dry districts, rather rare, 
but locally abundant. Jaffna (Gardner); Watagoda (Thwaites); Panwila; 
Uva Province, frequent; abundant about Nilgala, where it is gregarious. 
Fl. April; greenish-white. 

Also in India, Burma, and Malaya. 

The fruits are called ‘Ink-nuts’ or ‘Gall-nuts’ by the English, and 
are collected for sale to the Moormen by the country people. In Uva 
the collection is leased, and the sale in 1892 fetched R. 7500. They are 
the ‘Chebulic Myrobalans’ of the old pharmacists, and are here a valued 
medicine, both when young and in a mature condition. The flesh is very 
rich in tannin. When dried they show five blunt obscure angles. Wood 
heavy, very hard, dark brown with a purplish tinge, close-grained, durable. 


3. T. parviflora, 7iw. Enum. 103 (1859). Hanpalanda, S. 

T. zeylanica, Van Heurck and Mull.-Arg. Obs. Bot. 220. C. P. 618. 

Fl. B. Ind. ii. 446 (var. of Z. Chebula). 

A moderate-sized tree, with smooth, thin, pale brown bark, 
young shoots quite glabrous; |. numerous, alt. or sub-opposite, 
34-44 in., ovate-oval, rounded or narrowed and often unequal 
at base, tapering to acuminate obtuse apex, entire but some- 
what undulate, perfectly glabrous on both sides, rather thin, 
venation pellucid, petiole 3-2 long, flat above, without glands; 
fl. sessile, very small, spikes in terminal and axillary panicles, 
very finely pilose, about equalling |., bracts minute, linear, 
pubescent; cal. pubescent outside, disk covered with long 
hair, segm. 4 (rarely 5), triangular, acute; stam. 8 (rarely 10); 
drupe 1-1} in. by ~ wide, oblong-ovoid, blunt, cylindrical, 
not at all ribbed, smooth, pale olive green with white dots, 
stone bony, with 5 deep wide grooves, and with 5 sharp angles. 
so as to be star-shaped in section. 


Moist region in forests up to 4000 ft., rather rare. Ramboda (Gardner); 
Ambagamuwa; Hantane; Heneratgoda; Karuwita Korale. 

Fl. April-June; pale sulphur yellow. 

Endemic. 

Clarke, in Fl. B. Ind., places this as a variety under 7. chebula; he 
cannot have examined adequate specimens. 

Heart-wood orange-brown, moderately heavy, close-grained, smooth. 


4.7%. glabra, W. & A. Prod. 314 (1834). BKumbuk, 5S. 
Marutu, 7. 

T. alata, Moon Cat. 73. Thw. Enum. to4. C. P. 1603. 

Fl. B. Ind. ii. 447 (Z. Avjuna). Beda. FI. Sylv. t. 28 (7. Avjuna) not 
good. 


Anogeissus.] Combretacee. 161 


A very large tree with an extremely thick trunk and 
horizontally spreading branches, bark very thick, smooth, 
pinkish- or greenish-white, flaking off in large flat pieces, 
young twigs finely pubescent ; 1. opp. or sub-opposite, 3-6 in., 
oblong or oval-oblong, rounded at both ends, obscurely apicu- 
late, very shallowly serrate-crenate in upper part, glabrous 
‘but not shining on both sides, pale dull green, veins pellucid, 
petiole very short, 1-3 in., with 2 (or 1) prominent glands at 
top immediately beneath 1.; fl. sessile, spikes rather lax, short, 
axillary or in small terminal panicles; cal.-limb nearly glabrous 
-on both sides, disk with a few long white hairs ; drupe 14-2 in., 
obovate-ovoid, somewhat narrowed at base, bluntly pointed, 
glabrous, fibrous-woody, with 5 stiff hard projecting wings 
becoming wider upwards, and striated with very numerous 
much-curved veins, dark brown. 

Banks of streams and rivers in the low country; very common in the 
-dry region ; rare in the moist districts. Fl. April, May; greenish-white, 
strongly honey-scented. 

Also in India. 

The name 7. Avjwna, Bedd., seems a quite unnecessary synonym. 

Grows to an enormous size in the beds of tanks and rivers in the dry 
region, and is, generally speaking, the most noticeable feature of such 
scenery. Cordiner recorded specimens at Yala, north of Hambantota, 
in 1800 (which is the earliest notice I find of it for Ceylon), with trunks 
23% ft. in circumference at 5 ft. from the ground. A well-known one at 
Colombo measured, in 1879, 45 ft. in circumference round the base, and 
243 ft. at 8 ft. above ground. 

The astringent bark is used in medicine. It is remarkable for the 
immense amount of lime it contains, and it is largely burnt as a source of 
lime for chewing with betel, the copious ash almost entirely consisting 
of pure calcium carbonate. 

Wood greyish-brown, with bands of darker colour, very hard and 
heavy, smooth, strong. 


[ 7. tomentosa, W. and A., is recorded for Ceylon in Fl. B. Ind. The 
specimen is in Herb. Kew., labelled by Gardner as from ‘ Jaffna, Dyke.’ 
It was, doubtless, planted there by Mr. Dyke. The species is very doubt- 
fully distinct from 7. glabra.] 


2, ANOGEISSUS, Wall. 


Tree, |. alt. or sub-opp., no stip., fl. small in globular heads; 
cal. with a long solid tube or neck extending above ov. and 
persistent, limb campanulate with 5 segm.; pet. 0; stam. 10, 
inserted on cal.-limb in 2 rows; ov. inferior, compressed, 
1-celled, with 2 pendulous ovules; fruit small, indehiscent, 
winged, capped by a beak formed of the persistent neck of 
cal.; seed solitary, cotyledons convolute.—Sp. 5; 4 in. “7 B. 
Ind. 

PART II. M 


162 Combretacee. [_Lumnitzera.. 


A. latifolia, Wall. Cat. x. 4014 (1828). Dawu, S. VekkKali, 7. 
Conocarpus latifolia, Roxb., Thw. Enum. 103. C. P. 1220. 
Fl, B. Ind. ii. 450. Wight, Ic. t. 994. Bedd. Fl. Sylv. t. 15. 


A small tree, trunk erect, bark very smooth, whitish-grey, 
young parts glabrous; |. alt. or sub-opp., 2-3 in., broadly 
oblong-oval or sub-rotundate, rounded or subcordate at base, 
obtuse, often apiculate at apex, entire, more or less undulate,. 
quite glabrous, pale dull glaucous green, pink when young, 
often conduplicate, venation pellucid, midrib prominent be- 
neath, pink, petiole very short, channelled above; fi. sessile in 
small dense heads on slender slighly pubescent peduncles 
scarcely longer than petioles; cal. glabrous, segm. short, 
broad ; fruit very small, crowded, in a small globular head, 
plane-convex, with a rather broad wing along the edges, and. 
beaked with the long persistent upper part of cal.-tube, 
brown. 

Open grass-lands in the dry country; rare, but locally abundant. 
Haragama; Bintenne; Nilgala, gregarious. Fl. Jan. Feb.; pale 
greenish-yellow. 


Also in Peninsular India. 
Heart-wood extremely hard and tough, but small. 


3. LUMNITZERA, Willd. 


Large shrub, |. alt., fleshy, without stip., fl. small, in axillary 
spikes ; cal.-tube produced much beyond ov., with 2 adnate 
bracteoles near the base, limb with 5 shallow segm.; pet. 5, 
small; stam. 10; ov. inferior, 1-celled, with 2-5 pendulous 
ovules ; fruit indehiscent, capped with the persistent cal.-tube 
and limb; seed 1, cotyledons convolute.—Sp. 2, both in AZ. 
BL. Ind. 


&.racemosa, Wlld. in Neue Schrift. Berl.iv.(1803). Bériya, S. 
hye Enum s1Osk CaP aTso4 
Fl. B. Ind. ii. 452. Rheede, Hort. Mal. vi. t. 37. 


A shrub or small tree, with smooth purplish bark, young 
parts quite glabrous, buds pointed ; |. sessile, 2-3 in., readily 
disarticulating, spathulate-oblong, much tapering to base, 
rounded at apex, more or less shallowly crenate in upper 
part or entire, quite glabrous, shining, fleshy, veins not visible 
except midrib; fl. small, sessile, laxly arranged in short 
spikes from the |.-axils; cal.-tube fusiform, quite glabrous, 
segm. shallow, obtuse, finely ciliate ; pet. oblong, acute ; fruit 
3-# in., oblong-ovoid, compressed, capped by persistent cal., 
glabrous, hard-fleshy, longitudinally striate when dry. 


Combretum.] Combretacee. Oz 


Mangrove swamps round the coast; rather common. Colombo; 
Chilaw; Puttalam; Kalpitiya; Panadure; Kalutara; Jaffna. FI. Feb.; 
white. 

Shores of the Eastern Tropics generally. 

This forms part of the vegetation of tidal swamps, and is reckoned as 
amangrove. Turns black in drying. Wood pale reddish-grey, smooth 
and shining, rather heavy, very strong and durable, but of small size. 


4. COMBRETUM, L. 


Semi-scandent or straggling shrubs, |. opp., entire, fl. poly- 
gamous, in axillary spikes or racemes; cal.-tube constricted 
above ov. and more or less produced beyond it, limb cam- 
panulate cup-shaped or funnel-shaped, segm. 4; pet. 4, very 
small ; stam. 8, inserted with pet. on cal.-limb; ov. inferior, 
narrow and stalk-like, 1—-celled, with 2-5 pendulous ovules ; 
fruit dry or fleshy, indehiscent, with 4 prominent wings ; seed 
solitary, cotyledons large, plaited.—Sp. 120; 18 in FZ. B. Ind. 


Fruit with thick fleshy wings . : ; ; . I. C, ACUMINATUM. 
Fruit with thin papery wings 
Cal.-limb cup-shaped . 5 : é : . 2. C. OVALIFOLIUM. 
Cal.-limb long funnel-shaped : ‘ . 3. C. EXTENSUM. 


I. C. acuminatum, foxd. Hort. Beng. 28 (1814). 

C. sarcopterum, Thw. Enum. 415. C. P. 3715. 

Fl. B. Ind. ii. 455. 

A low straggling bush, with long branches, not climbing, 
young shoots with rough scaly ferrugineous pubescence ; 
|. 5-7 in., oval or ovate-oval, rounded at base, more or less 
acuminate, sub-obtuse, undulate, glabrous, densely covered 
beneath with minute prominent glands, venation prominent 
beneath, petiole very short, 4-4 in.; fl. on very short ped., 
racemes spike-like, usually solitary, shorter than 1.; cal. 
glabrous, minutely glandular, limb campanulate ; pet. minute, 
spathulate ; stam. twice or thrice as long as cal.; fruit nearly 
2 in., ovoid-oblong, narrowed at both ends, fleshy, minutely 
glandular, brown, wings thick and blunt. 

Moist low country; rare. Colombo (Ferguson); Horana, Reigam 
a Hiniduma; Kalutara; Heneratgoda. FI. August-October; 
E€llow. 

: Also in Assam, Burma, Malaya, and the Philippines. 


2. C.ovalifolium, ox). Hort. Beng. 28 (1814). Kaduruketiya- 
wel, 5S. 

C. Wightianum, Thw. Enum. 103 (in part). C. Thwazttestanum, Van 
Heurck and Mull. Arg. Obs. Bot. 238. C. P. 1601. 

Fl. B. Ind. ii. 458. Hook. Bot. Miscell.'iii. t. supp. 22 (C. Hleyneanum). 


164 | Combretacee. [Gyrocarpus. 


A large straggling or climbing shrub, with numerous 
divaricate branches thickened at the nodes, bark smooth 
yellowish-grey, young parts glabrous; |. opposite, 3-44 in., 
oval, acute at base, acuminate, obtuse, glabrous on both sides, 
petiole % 4-1 in. long; fl. numerous on very short glabrous ped., 
in show, rather dense, pedunculate racemes (often paniculate) 
axillary from |. of previous year and 3-6 (occasionally reaching 
8) in. long, buds globose, apiculate, bracts minute; cal.-tube 
glabrous outside, constricted into a short neck above ov., then 
suddenly expanded into a cup-shaped limb, hairy within, 
segm. triangular, acute, reflexed ; pet. about as long as cal.- 
segm., oblong; fruit 1-14 in., pale shining golden-brown, 
wings very large, $ in. in widest part, papery, stiff, glabrous, 
finely transversely striate. 

Dry country; rather common. Tiripane; Kekirawa; Haragama; 


Kurunegala. Fl. July, August; white. 
Also in the Indian Peninsula. 


3. C. extensum, oxb. Hort. Beng. 28 (1814). 


C. Wightianum, Wall., Thw. Enum. 103 ay C. platyphyllum, 
Van Heurck and Mull. Arg. Obs. Bot. 242. C. P. 1602. 


FI]. B. Ind. ii. 458. Wight, Ic. t. 227 (C. Wightianume). 

A straggling semi-scandent shrub with the habit and |. of 
the last ; but petioles shorter, under }in.; fl. rather larger but 
similarly arranged, buds ovoid, sharp-pointed; cal.-tube much 
longer, contracted above ov. and then at once expanding into 
a long funnel-shaped limb hairy within, segm. narrowly tri- 
angular, acuminate, very acute, reflexed; pet. oblong, truncate ; 
fruit (not seen) much as in C. ovalzfolium. 


Low country to 3000 ft.; rather rare. On Dambulla Hill, trailing over 
bare rock; Hantane (Gardner). Fl. May; yellowish white. 
Also in India and Malaya. 


Thwaites combined 2 and 3, but they are, though much alike, readily 
distinguished by the calyx. 


5. GYROCARPUS, /acg. 


A tree, |. alt., clustered at ends of branches, no stip., fl. 
very small, unisexual and moncecious or polygamous, in large 
bractless cymes; male fl.:—cal. with 4-7 irregular segm., pet. 0, 
stam. usually 4, alternating with 4 small fleshy staminodes, 
_anth. opening upwards by 2 valves; fem. or bisexual fl.:— 
cal.-tube adnate to ov., segm. 2, persistent and greatly enlarged 
in fruit, pet. 0, ov. inferior, 1-celled with a single pendulous 
ovule, style short, stigma capitate; fruit drupe-like, crowned 


Myrtacee. 165 


with the long wing-like persistent cal.-segm., stone bony ; 
seed with large convolute cotyledons. 
Placed by Thwaites at end of Lauracee, to which its stamens are so 


remarkably similar. The fruit is constantly mistaken for that of a 
Dipterocarp. 


- pasacaria Roxb. Cor. Pi. 1.,1. (1795). Hima, S. Tanak- 
u, 7. 
@ astaticus, Willd., Moon Cat. 73; Thw. Enum. 258. C. P. 2202. 

Fl. B. Ind. ii. 461 (not given for Ceylon). Bedd. Fl. Sylv. t. 196. 

A moderate-sized or large tree, with an erect trunk and 
rather small spreading head, bark smooth, shining, greenish- 
white, young branches stout, marked with leaf-scars, young 
parts densely pubescent; 1. alt. rather crowded, spreading, 
4-6 in. and nearly as wide, very broadly ovoid or somewhat 
rhomboid, tapering to base, acuminate, acute, glabrous above, 
densely soft-pubescent with stellate hair beneath, strongly 
3-veined from base, petiole very long, 2-5 in., slender; fl. small, 
the male very numerous, the female few, all on slender ped. 
arranged in globular clusters in large much-branched cymes 
on long pubescent peduncles, several of which arise from the 
ends of the twigs above the new 1.; cal. densely hairy- 
pubescent ; fruit (without the wings) about 2 in., ovate-ovoid, 
furrowed in upper part, wrinkled below, pubescent, wings 
24-3} in. long, linear-spathulate, obtuse, thinly leathery, 
striate, pubescent, greyish-brown, stone ovoid, bony, furrowed. 

Dry and intermediate country; rather common. Moon’s locality is 
Kattregam. FI]. Feb., March, April; cream-coloured. 


Throughout the Tropics, G. americanus, Jacq., not being distinguish- 
able. 

The tree is quite bare of leaves for a short time. Wood very light, 
soft, greyish white; much used in the north of the island for catamarans 
(rafts) and the outriggers of boats. Lac is found on this in the Uva 
country. 


LII.—MYRTACE/E. 


TREES or shrubs, |. opp. or alt., simple, without or with very 
minute stip., fl. regular, bisexual ; cal.-tube adnate to ov. and 
rarely produced beyond it, segm. 4 or 5 (rarely 2 or 3), usually 
persistent in fruit; pet. 4 or 5 (very rarely more), distinct, o1 
slightly connate at base or united into a deciduous cap; stam 
indef., epigynous, distinct or connate below; ov. inferior, 2-4- 
celled, with many ovules in each cell, style simple; fruit 


166 Myrtacee. [Rhodomyrtus. 


indehiscent, berry-like, pulpy or hard, with 1, few, or numerous 
seeds; seed without endosperm, embryo with small cotyledons, 
or quite without them (macropodous). 


L. opposite ; stam. distinct. 


Ovary 3-celled; 1.3-nerved .. 1. RHODOMYRTUS. 


Ovary 2-celled ; |. with an intramarginal vein . 2. EUGENIA. 

L. alternate ; stam. connate below (Barrzngtoniec). 
Seed solitary . ; : : : ‘ : . 3. BARRINGTONIA. 
Seeds numerous . j : : c 2 4s \CARTEYAS 


Our species are mainly inhabitants of the moist region and the hills. 
Rhodomyrtus and 15 species of Augenza are confined to the montane 
region, and 22 species of Eugenza and 3 of Barringtonia to the moist low 
country, Carveya extends up from the one into the other. Only 4 species 
of Eugenia are found in the dry region, 1 only, &. bracteata, being con- 
fined to it, as is also Barringtonia acutangula. 


Eucalyptus. A considerable number of species of this large 
Australian genus have been introduced into the hill-country and much 
planted on estates for fuel and shelter. The species most frequently 
met with are £. Globulus (Blue Gum), £. diversicolor (Karri), &. Leu- 
coxylon (Iron Bark), £. robusta (Swamp Mahogany), £. marginata 
(Jarrah) ; but there are many others. 


I. RHODOMYRTUS, DC. 


Large shrub, |. opp. 3-nerved, fl. axillary ; cal.tube tur- 
binate, adnate to ov., and not produced beyond it, segm. 5, 
persistent ; pet. 5, spreading, stam. indef., distinct ; ov. inferior, 
3-celled, with numerous ovules in each ceil superposed in a 
double row, style simple, stigma capitate; fruit a berry, 3- 
celled, with numerous horizontal seeds; seed compressed, 
embryo curved with a long radicle and small cotyledons, no 
endosperm.—Sp. 5: 1 in FZ. B. Ind. 


R. tomentosa, Wight, Spicil. Nezlgh. i. 60 (1846). 

Myrtus tomentosa, Ait., Moon Cat. 39; Thw. Enum.114. C.P. 1591. 

HB nd ne46e. SWaeht. | entazi- 

A large shrub or small, very much-branched tree, bark 
yellowish, vertically fissured, exfoliating, young branches 
finely tomentose; 1. 2-24 in., oval or oblong-oval, acute at 
base, obtuse or acute at apex, glabrous (when mature) above, 
densely covered with fine grey or yellowish tomentum beneath, 
stiff (very rigid when old), strongly 3-nerved (besides the 
marginal one), margin slightly recurved, petiole 4-3 in.; fl. 
on stout pubescent ped., about I in. diam., solitary or 3 in a 
peduncled cyme, with 2 small linear bracts beneath each ; 
cal. densely tomentose, segm. rotundate, spreading; pet. 


Eugenia.) Myrtacee. 167 


broadly obovate, tomentose on the back, spreading or re- 
flexed ; fruit nearly globular, capped with cal.-segm., tomen- 
tose, soft, 3-celled, with a double row of seeds in each cell; 
seed triangular-reniform. 

Upper montane zone; common. FI. April—July; pet. white, tinged 
outside with purplish-pink, fil. flesh-coloured, darker at base. 

Also in S. India, Malaya, and China. 

The fruit is edible and pleasant. I have heard it called ‘wild guava’ 
by the English at Nuwara Eliya; in the Nilgiris it goes by the name of 
“ Hill Gooseberry.’ Moon gives the S. name ‘ Sudu-kotala ;’ his locality 
is Uva. 

At Malacca I found this abundantly on the sandy seashore, but in 
Ceylon it is entirely a montane plant. 


Psidium Guyava, L. The Guava must have very quickly become 
naturalised here. There are specimens in Hermann’s Herbarium, and he 
expressly states (Mus. 3) that it was brought by the Portuguese. There 
is no true Singhalese name, ‘ Péra,’ the name usually employed, being 
Portuguese for Pear. P. pumilum, Vahl, Symb. ii. 56, is based on 
Ceylon specimens collected by Koenig. The Guava is native in Mexico, 
and perhaps other parts of Trop. America, but has now all the look of a 
wild plant in Ceylon. 


2. EUGENTA,* Z. 


Trees or shrubs, |. opp., entire, with a more or less con- 
spicuous marginal vein, usually gland-dotted, without stip., 
fl. in terminal or axillary paniculate cymes or solitary in axils 
of 1. or scales below the 1. often on suppressed branchlets, 
thus appearing to be fasciculate or racemose; cal.-tube from 
cup-shaped to tubular, adnate to ov. and sometimes produced 
beyond it, segm. 4 (rarely 5); pet. 4 (rarely 5 or more), either 
distinct and spreading or connate into a cap (calyptrate) 
which falls off on expansion of fl.; stam. indef., usually quite 
distinct, in many rows, anth. small, versatile; ov. inferior, 
2-celled with several ovules in each cell, style simple; fruit 
usually a juicy berry, rarely dry, globular or ovoid, crowned 
with persistent cal.-limb or segm.; seeds I or very few, 
embryo with thick often connate cotyledons, no endosperm.— 
Sp. about 700; 131 in FZ. B. Jnd. 

Our largest genus (except Cyperus), forming a main constituent of the 
forest of the moist and montane regions, to which the brilliant pink, 


orange or crimson tints of the young foliage are a great ornament. No 
less than 29 of our species are endemic. 


* Given by Micheli in 1728, in honour of the illustrious Prince Eugene 
of Savoy, to the plant now called £. Micheliz, Lam. 


168 Myrtacee. [ Zugenia,. 


Though the species are not difficult to recognise, the differences are 
hard to describe in few words, and the following clavis needs checking by” 
the fuller descriptions. 
rl. in cymes. 

Pet. large, distinct ; cal. with large staminal 
disk ; fruit over 2 in. ( /ambosa). 
Cal.-tube funnel-shaped or turbinate. 


Lat. veins of 1. few, distant 1, E. AQUEA. 

Lat. veins of |. numerous, close 2, E. GRANDIS. 
Cal.-tube hemispherical 3. E. HEMISPHARICA.. 
Cal.-tube cylindric, long 4. E. CYLINDRICA. 

Pet. small, usually combined and falling ‘off as 

a cap (calyptrate) (exc. 12, 13, and 22) ; 

no staminal disk; fruit "under Sine 

(Syzygium). 

Cymes terminal or in axils of present 1. 

Cal.-tube long, funnel-shaped. 

Fruit globose, white ‘ 5. E. SPICATA. 
Fruit ovoid- turbinate, tapering to base 6, E. LANCEOLATA. 


Fruit ovoid-urceolate, crowned with 
long cal.-limb . : 7. E. FERGUSONI. 
Cal.-tube short, cup-shaped or turbinate. 
L. oval or lanceolate. 
Branchlets quadrangular. 


L. over 2 1n., acuminate : . 8. E. LISSOPHYLLA. 
L. under 2 in., rounded : . g. E. SUBAVENIS. 
Branchlets cylindrical, often com- 
pressed. 
L. petiolate. 
Cal. truncate. 
Tree, |. oval . 2 ¢ . Io. E. GARDNERI. 
Bush, |. obovate-oval . . 11, E. CORYMBOSA. 
Cal.-segm. obvious. 
Pet usually distinct. 
L. not revolute, fl. on ped. 12. E. MICRANTHA. 
L. revolute, fl. sessile. . 13. E, REVOLUTA. 
Ree calyptrate. 
Fl. small, ¢ in. diam. . 14. E. SYLVESTRIS. 
Fl. large, 3 ‘in. diam. . . 15. E. ASSIMILIS. 
L. sessile. 
L. very broadly oval é . 16. E. CORDIFOLIA. 
L. lanceolate-oblong : . 17. E. NEESIANA. 
L. rotundate. 
L. over 2 in., branchlets cylindrical 18. E. CYCLOPHYLLA.. 
L. under 1 in., branchlets quad- 
rangular. , 
L. petiolate . : : : . I9. E. ROTUNDIFOLIA.. 
La sessile 2 ° ; : . 20. E. SCLEROPHYLLA.. 
L. obovate (see also 11); fl. few, 
sessile, capitate . : : . 21. E. OLIGANTHA. 
L. spathulate; fl. in cymes, pedi- 
cellate . ; : ; . 22. BE. OLIVIFOLIA. 
Cymes in axils of fallen leaves. 
Lat. veins few : : . 23. E. OPERCULATA.. 
Lat. veins very numerous é . 24. E, JAMBOLANA. 


Eugenia.] Myrtacee. 169 


Fl. solitary or in fascicles ; pet. distinct (see 
also 21 and 22). 
Young parts pubescent (very minutely so 
in 33). 
Fl. pedunculate. 
Fl. over I in. diam. 


L. rotundate, revolute . ; S25 UCIDA: 
L. oblong-ovate, cordate 26. E. HACKELIANA. 
Fl. under 1 in. diam. 
L. oval or lanceolate, over 2 in. 
Fruit over 4 in. diam. 
Peduncles short; 1. over 3 in. 
Fruit over #1 in. . 27. E. TERPNOPHYLLA. 
Fruit 3 in. , orange- -yellow . 28. E. XANTHOCARPA. 
Peduncles long ; l.under 3in, 29. E. BRACTEATA. 
Fruit + in. diam. . 30. E. RUFOFULVA. 
IL linear- lanceolate, under bine =. 31. E. PHILLYRAOIDES.. 
Fl. sessile or nearly so. 
F]. about 1 in. diam. 
L. lanceolate-oblong . : . 32. E. FLOCCIFERA. 
L. oval or ovate-oval 33. E. RIVULORUM. 
F]. under # in. diam. ; 
L. under 5 in. : : : Sestily 1d TNOIENH NG 
L. over 6 in. 35. E. INSIGNIS. 
Young parts glabrous or nearly 50. 
Fl. sessile. 
L. 4-5 in. : ; : 5 . 36.. E. DECORA. 
L, 1-3 in. 7 : : ; . 37. E. ROTUNDATA 
Fl. pedunculate. 
Peduncles under } in. 
L. nearly sessile, oval-spathulate . 38. E. MABAOIDES. 
L. distinctly petiolate, rotundate . 39. E. APRICA. 


Peduncles over + in. 
sessile . : : : . 40, E. AMGENA. 
L. petiolate. 
L. under 3 in. 


Peduncles over}in. . . 41. E. PEDUNCULATA. 
Peduncles under } in. . . 42, E. MOONIANA. 
L. over 3 in: ; ; ee SAS EL WALCE SIU, 


1. BE. aquea, Lurm. Fil. Ind. 114 (1768). Wal-jambu, S. 

Herm. Mus. 67. Burm. Thes. 125. £&. sylvestris, Moon Cat. 38 (non 
Wight). /ambosa aquea, DC., Thw. Enum. 115. C. P. 418. 

Fl. B. Ind. ii. 473. Wight, Ic. tt. 216, 550. 

A small or middle-sized tree with smooth grey bark, 
much branched, twigs slightly quadrangular, glabrous; 1. 
very shortly stalked or nearly sessile, variable, 2-8 in., oval 
or oblong-oval, usually rounded or subcordate at base, obtuse 
or rounded at apex, coriaceous, shining, glands minute, black, 
lat. veins distant, rather prominent beneath; cymes 3-I0- 
flowered, on short, stout, quadrangular peduncles, terminal 
and from upper axils, ped. shorter than cal.-tube; cal.-tube | 
tubular-funnel-shaped, somewhat variable in length and 


170 Myrtacee. [Zugenia. 


breadth, segm. large, usually broader than long, with mem- 
branous margins, spreading; pet. large, roundish, spreading ; 
fruit #-1 in., globose, crowned with prolonged neck-like cal.- 
limb and segm. 


Moist region, especially between 2000 and 5000 ft.; common; often 
planted. Fl. March—May; white or purplish rose-coloured. 

Also in East Bengal and Burma. 

At the higher elevations this puts on a very different appearance, 
with smaller (13-3 in.) more oblong and more coriaceous leaves ; but it is 
connected with the type by intermediate forms. 

There is a cultivated spreading tree, very handsome in May when 
covered with its copious brilliant deep rose-coloured flowers, which 
must, I suppose, be referred to this species. But I have not seen it wild, 
and its origin is doubtful. It bears a large urceolate pale-green fruit 
crowned with the spreading cal.-segm. 

E. malaccensis, L., the Malay apple, is cultivated under several 
varieties, as is also Z. /ambos, L., the Jambu or rose-apple. Both are 
Malayan species, early introduced here, and both noticed by Hermann. 
The former, with large rose-red flowers and red pear-shaped fruits, is well 
figured in Wight, Ill. t. 98; the latter, with white flowers and narrow 
leaves, in Wight, Ic. t. 435. These fruits are pleasantly subacid and 
refreshing. 


2. BE. grandis, Wight, /1/. ii. 17 (1850). 

e ee jfirmum, Thw. Enum. 417 (S. montanum, Thw. Enum. 116). 

Fl. B ind. u. 476. Wight, Ic. t. 614. 

A large tree, bark smooth, whitish, shoots glabrous, 1. 
large, 4-8 in., obovate-oval, tapering to base, rounded often 
retuse or very abruptly acuminate at apex, margin revolute, 
very coriaceous, thick and shining, lat. veins numerous, close, 
parallel, petiole about 4 in., very stout ; cymes terminal and 
axillary, large, about as long as 1. fl. almost sessile, in 35; 
cal.-tube nearly 4 in., turbinate, attenuate below, segm. 
rounded, 2 larger than others ; pet. rounded ; fruit not seen, 
‘globose or somewhat pyriform, 1-14 in., crowned with the 
large cup-like cal.-limb.’ 

Montane zone at about 4000 ft.; very rare. Ambagamuwa District. 
Fl. Sept.; white. 

Also in Burma, Malay Pen., and Borneo. 

Our plant differs slightly from this common Malayan species in its 


longer and laxer panicles, with fewer and larger fl., and its more obovate 
leaves. 


3. &. hemispherica, W777i, J//. ii. 14 (1850). 

Strongylocalyx hemisphericus, Bl, Thw. Enum. 116. C. P. 2450, 
3438, 2540. 

BBS sid i477 NV etamlient 525. bedd bi Sylvaitzoge 

A medium-sized tree, with smooth, yellowish-grey bark, 
young twigs sub-tetragonal ; 1. 34-5 in., lanceolate, tapering 


Eugenia.) Myrtacee. iG 


to base, caudate-acuminate, subacute, stiff, smooth and shin- 
ing, petiole about 4 in., slender ; cymes terminal and axillary, 
shorter than l., fl. large, 1} in., ped. long, articulated below f1., 
‘buds large, broad; cal. tube short, 4-3 in., hemispherical, 
segm. large, rounded, white, with membranous margins, 
‘spreading ; pet. large, ‘rounded, very concave, reflexed ; fruit 
globose, #-1 in., green, crowned with cal.-segm. 

Moist region up to 6000 ft., chiefly in the lower montane zone; rather 
rare. Deltota; Ambagamuwa; Kukul Korale; between Haputale and 
Galagama, abundant. Fl. Feb._May; cream- coloured, sweet-scented. 

Also in Southern India. 

An ornamental plant when in flower. The fruit resembles a small 


apple. C. P. 2540is a form from Elk Plains (over 6000 ft.), with 1. only 
13-23 in. long, and small fl. 


4. E. cylindrica, Wight, ///. ii. 14 (1850). 
Jambosa cylindrica, Thw. Enum. 115. C. P. 601. 
Fl. B. Ind. ii. 479. Wight, Ic. t. 527. Bedd. Fl. Sylv. t. 201. 


A medium-sized tree, with smooth, yellowish-grey bark, 
twigs cylindrical ; 1. 34-54 in., narrowly lanceolate, tapering 
to base, caudate-acuminate, subacute, pale beneath with 
prominent venation, lat. veins very few, uniting with the 
marginal vein at a considerable distance from the edge, 
petiole 4 in., stout; cymes terminal and from upper axils, 
shorter than 1., peduncles long, slender, compressed-quadran- 
gular, ped. divaricate, slender; cal.-tube 4-3 in., slender, 
cylindrical, tapering to base, slightly narrowed at summit, 
limb expanded cup-shaped, segm. pounds i triangular, sub- 
acute ; pet. large, concave; fruit about # in., nearly clobose, 
crowned by the short neck- ‘like cal.-tube aa reflexed segm. 


Moist low country to 3000 ft.; rather rare. Ambagamuwa District ; 
Hantane; Kalawane; Kukul Korale. Fl. Feb., March; pinkish-white. 

Endemic. 

A beautiful species, easily distinguished when in flower by the long 
slender, white calyx-tube; the buds look like large white cloves. When 
not in flower the peculiar venation of the leaves is characteristic. 


5. EB. spicata, Lam. Encycl. Meth, iii. 201 (1789). Maran, 
Maranda, 5S. Marungi, 7: 

Herm. Mus. 9. Burm. Thes. 166. FI. Zeyl.n. 182. Myrtus zeylanica, 
L. Sp. Pl. 472; Moon Cat. 39. £. zeylanica, Wight, Ill. 11.15. Acmena 
zeylanica, Thw. Enum. 118. C. P. 56, 380. 

Fl, B. Ind. ii. 485 (£. zeylanica). Wight, Ic. t. 73. Bedd. Fl. Sylv. t. 
202 

A large shrub or tree, with smooth or furrowed pale 
prow bark, and slender, shining, purplish twigs; 1. variable, 
14-44 in., from ovate-oval to lanceolate- linear, usually 
caudate-acuminate, obtuse, margin slightly revolute, smooth 


172 Myrtacee. [Eugenia. 


and shining on both sides, lat. veins numerous but incon- 
spicuous, petiole about }in.; cymes axillary and terminal, 
shorter than 1. fl. small, numerous, rather crowded, ped. very 
short, glandular ; cal.-tube under } in., funnel-shaped, densely 
glandular, segm. rotundate, obtuse or subacute, erect ; pet. 
Salpemncey calyptrate; fruit globose or nearly so, a little 
under }in., pure white. 

Dry a eerie country up to 2000 ft. or higher; very common. 
Fl. March and April; white. 

Also in S. India and Malaya. 

The pure white berries are ornamental and conspicuous. The leaves. 
are very fragrant; they vary much; the narrow-leaved form is the com- 
monest in the dry districts. C. P. 380, from Palagama, has small thick 
leaves and larger flowers. 

Wood brown, heavy, liable to split. 


6. E. lanceolata, Lam. Encycl. Meth. iii. 200 (1789). 

Wight, Ill. ii. 15. <Acmena lanceolata, Thw. Enum. 119. C. P. 2863. 

Fl. B. Ind. ii. 285 (Z. Wightiana). Wieht, Ic. tt. 529 (Z. Wightiana): 
and 530. 

A rather small tree with grey bark and slender cylindrical 
twigs; |. 34-5 in., broadly or narrowly lanceolate, tapering 
below, acuminate-caudate, acute, glabrous, lat. veins very fine, 
numerous, faintly marked, midrib impressed above, petiole 
4 in. or less; cymes short, usually few-fld., axillary often 
from axils of fallen 1., fl. nearly sessile, articulated ; cal.-tube 
3-3 in., tubular-funnel-shaped, tapering below, segm. 4 or 5, 
very shallow, rounded ; pet. numerous, sometimes as many as. 
12, but usually calyptrate; style long, persistent ; fruit about 
7 in., erect, ovoid-turbinate, tapering to long base, capped 
with spreading cal.-segm. and long erect style, vertically 
striate. 

Low country to 2000 ft. in the moist and intermediate regions, 
especially by streams; rather rare. Near Matara; Pasdun Korale;, 
Singhe Rajah Forest; Ambagamuwa; Kurunegala. FI. March; white. 

Also in South India. 

I think there are two plants confounded here, as I have collected in 
the Pasdun Korale specimens with thicker, broader, nearly sessile leaves, 
with more marked venation and with a shorter calyx; but I cannot fit our 
plants to £. lanceolata and £. Wightiana as separate species. 


7. &. Fergusoni, 77772. 


A bush or small much-branched tree, bark reddish-grey,. 
twigs stout, quadrangular, thickened at the nodes, orange- 
coloured, ieaf-scars prominent ; |. 14-3 in., crowded, very nearly 
sessile, ovate-oval, acute or obtuse at apex, rounded or subcor-- 
date at base, margin generally slightly curved, thinly coriaceous, 
stiff, lat. veins numerous, faintly marked beneath; fl. large, 


Eugenia.| Myrtacee. 17% 


sessile, cymes strictly terminal, with very short branches, so 
as to be corymbose or umbellate; cal. (when in fl.) very long, 
3-14 in., tubular-funnel-shaped, gradually tapering to slender 
stalk-like base, mouth 3in. diam., almost truncate, with 
shallow rounded segm.; pet. calyptrate (or sometimes expand- 
ing), large, reddish outside ; fruit broadly ovoid, about ? in., 
somewhat urceolate, crowned with rather long cal.-limb. 

Var. 8 minor, 7rim. Syzygium calophylifolium, Thw. Enum. 118 
{non £. calophylizjolia, Wight). C. P. 160. 

L. much smaller, not exceeding 1 in., very shortly stalked, 
fruit about 4in., nearly globular, purple. 

Upper montane zone; in open sunny places. The type only in the 
North-eastern mountain block, Wattekelle (Ferguson), Knuckles Hills 
and Rangala Ridge. Var. 8, Adam’s Peak (Gardner and Thwaites). FI. 
April; pinkish-white. 

Endemic. 

Var. 8 seems to differ altogether from £. calophylizfolia (to which 
it is still referred in Fl. B. Ind.) in its leaf-venation as well as its 


remarkably long calyx. The C. P. specimens are in fruit only, and I 
have not seen the flowers. 


8. E. lissophylla, Duth. in Fl. B. Ind. ii. 488 (1878). Maha- 
kuretiya, 5S. 


Syzygium lissophyllum, Thw. Enum. 117. C. P. 2452. 

Fl. B. Ind. ii. 488. 

A moderate-sized tree, with thin, rather smooth pale- 
brown bark, young twigs quadrangular; 1. 2-3 in., oval, 
tapering to base, caudate-acuminate, obtuse, glabrous and 
shining above, paler beneath, lat. veins very fine and nume- 
rous ; fl. sessile or nearly so, in 3s at ends of quadrangular 
branches of terminal, and axillary cymes generally exceeding 
].; cal. shortly campanulate, segm. short, rounded ; pet. 
calyptrate; fruit globose (not seen ripe). 


Forests in the lower montane zone; rathercommon. FI. April, May ; 
pinkish-white. 

Also in the hills of Southern India. 

Wood rather heavy, hard, yellow. 


9g. &. subavenis, Duh. in Fl. B. Ind. ii. 489 (1878). 

Syzygium umbrosum, Thw. Enum. 118 (non Berg). C. P. 2539. 

Fl. B. Ind. ii. 489. 

A moderate-sized much-branched tree with brownish-grey 
bark, young twigs quadrangular, smooth, purple; |. small, 
14-17 in., broadly obovate-oval or oval, tapering to base, 
obtuse or rounded and often emarginate at apex, usually 
revolute at margin, thick, lat. veins numerous, fine, incon- 
spicuous, fl. as in Z. “ssophylla,; fruit 3 in. diam., nearly 
globose. 


174 Myrtaceae. [Eugenia 


Forests of the montane zone, 4000 to 6000 ft.; rather common. FI.. 
April; white, calyx crimson. 

Endemic. 

The up- country carpenters call the wood of this tree ‘ Weli-damba,’ or 
‘Hin-damba.’ It is softer and more open than that of most species. 


Io. B. Gardneri, Duth. in Fl. B. Ind. ii. 489 (1878). Dambu,* 5.. 
Nir-naval, 7. 

Syzygium Gardnert, Thw. Enum. 117. C. P. 2946. 

Fl. B. Ind. 11. 489. 


A moderate-sized or large tree, with smooth pale-grey 
bark, much branched, young twigs cylindrical, often some- 
what compressed ; |. 3-34 in., oval or broadly oval, acute at 
base, caudate-acuminate, obtuse, smooth and shining, rather 
thin, bright apple-green, lat. veins very Mumeroms, fine, con- 
spicuous, pellucid, petiole 21 in.; fl. small, numerous, 
sessile, cymes copious, axillary and terminal, shorter etal L i 
cal. campanulate, truncate with obscure segm.; fruit 3-4 in., 
nearly globular and without any crown of cal.-segm. 


Moist region between 2000 and 4ooo ft.; common.~ FI. July-Sept.;. 
white. 

Also in Southern India. 

Wood rather hard and heavy, smooth, greyish-yellow. 


11. &. corymbosa, Lam. Encycl. Meth. iii. 199 (1789). Dan, 
Hin-dan, S: 

Herm. Mus. 3,14. Burm. Thes. 57. FI. Zeyl.n. 183. yrtus cary- 
ophyllata, LL. Sp. Pl. 472. Calyptranthes caryophyllata, Pers., Moon Cat.. 
38. Vays zum caryophylleum, Gaertn. Fruct. i. 166; Thw. Enum. <n 
CoE bos: 

Fl. B. Ind. ii. 490 (Z. caryophyllea). Wight, Ic. t. 540 (Id). 


A bush or shrubby tree, with smooth grey bark, young 
twigs slightly compressed, scurfy, orange-brown ; |. 24-34 in., 
obovate-oval, acute at base, rounded, obtuse or very shortly 
and bluntly acuminate, glabrous, shining and bright apple- 
green above, rather pale beneath, lat. veins numerous, fine 
but rather conspicuous beneath, petiole 4 in.; fl. small, 
numerous, nearly sessile, cymes terminal, corymbose, tricho- 
tomous, buds ovoid- clobose ; cal. campanulate, STRESS 
seem. scarcely perceptible ; pet. calyptrate; fruit about @ in., 
depressed-globose, inky-purple or nearly black, shining, j juicy. 


Low country in both moist and dry regions ; very common, especially 
in open sandy places. Fl. Feb.-May ; white. 

Also in Southern India and Borneo. 

The small black fruit is edible. 


* The name Dambu or Damba is applied to many other species of 
Eugenia. 


Eugenia. | Myrtacee. 175, 


12. E. micrantha, Duth. in Fl. B. Ind. ii. 483 (1878). 

Syzygium micranthum, Thw. Enum. 117. C. P. 1580. 

Fl. B. Ind. ii. 483. 

A moderate-sized tree with smooth, greyish-brown bark,. 
branchlets slender, cylindrical; 1. crowded, 2-3 in. or rather 
more, broadly oval, abruptly narrowed to base, conspicuously 
caudate-acuminate, obtuse at apex, margin often revolute, 
quite glabrous, lat. veins numerous, rather close, straight, 
petiole +4 in., slender; fl. very small, on short ped., cymes 
numerous, axillary, shorter than 1.; cal.-tube short, broadly 
turbinate, segm. distant, very short, acute; pet. very small; 
fruit very small, under 4 in., globose, capped by cal.-segm., 
pale purple. 

Forests in lower montane zone; common. FI. Feb., March, and’ 


October ; greenish-white. 
Endemic. 


13. BE. revoluta, Wight, ///. ii. 17 (1850). 

Syzygium revolutum, Thw. Enum. 117. C. P. 2521. 

Fl. B. Ind. ii. 492. Wight, Ic. t. 534. 

A moderate-sized tree with pale brown, finely cracked, 
bark, twigs somewhat compressed ; |. 14-4 in., more or less 
obovate-oval, tapering to base, obtuse, rounded, often emar- 
ginate at apex, more or less revolute at margin, thick, stiffly 
coriaceous, shining above, usually glaucous beneath, lat. veins 
prominent especially beneath, petiole very short; fl. sessile 
in clusters of 3 or more, cymes numerous, pedunculate, 
axillary and terminal, shorter than |, sometimes from the 
old wood, cal.-segm. short, pet. usually distinct, small; fruit 
about 3} in., globose, fleshy, dark purplish-red. 

Forests of the montane zone up to 7000 ft.; common. FI. Feb., 
March, and September ; pinkish-white. 

Also in the Nilgiris. 

At the higher elevations this has smaller leaves, 1-14 in. long only. 

The wood-cutters at Nuwara Eliya call this ‘ Karon-damba,’ the wood 
is very hard and strong, but liable to split. 


14. BE. sylvestris, Wight, ///. ii. 15 (1850) non Moon. Alubo, 5S. 

Calyptranthes Jambolana, Moon Cat. 39. Syzygtum sylvestre, Thw. 
Enum. 116. C. P. 2862. 

Fl. B. Ind. ii. 493. Wight, Ic. t. 532. 

A large tree with smooth yellowish-grey bark, twigs 
cylindrical; 1. 34-5 in., oblong-oval, tapering to base, shortly 
acuminate, obtuse and generally twisted at apex, margin 
slightly recurved, coriaceous, lat. veins numerous, fine, rather 
prominent, petiole }—4 in., fl. sessile, small, in dense clusters, 
cymes nearly all terminal, on long peduncles, crowded ; pet. 
calyptrate ; fruit about } in., globose, purplish-black. 


176 Myrtacee. [Zugenia. 


Moist and intermediate low country to 3000 ft.; rather common. FI. 
April-June ; white. 

Endemic. 

Wight gave Moon’s name sy/vestrzs to this in error ; Moon’s plant is 
£. aquea, as Wight’s description ‘copied from Moon’s notes’ shows ; 
but Wight’s figure represents the present species, and there is thus no 
need to alter the accepted nomenclature. 

Wood reddish-grey, smooth, durable. 


15. B. assimilis, Duh. in Fl. B. Ind. ii. 493 (1878). 
Sysygtum asstmile, Thw. Enum. 116. C. P. 46, 2449. 
Fl. B. Ind. 11. 493. 


A moderate-sized tree, twigs stout, sometimes subquad- 
rangular; 1. variable, 13-5 in., oval or broadly oval or 
obovate-oval, tapering to base, more or less acuminate, 
obtuse and often twisted at apex, margin often revolute, 
stiffly coriaceous, lat. veins numerous, rather conspicuous, 
petiole +-4 in. stout; fl. rather large, 4 in., sessile, cymes on 
stout peduncles, crowded, mostly terminal; cal.-tube cup- 
shaped ; pet. usually calyptrate ; fruit $—2 in., globose, green, 
capped with large cal.-segm. 


Moist region, from low country up to 6000 ft.; common. FI. March, 
April ; cream-coloured. 

Endemic. 

Distinguished from £. sylvestrzs by its much larger flowers and more 
prominent lateral leaf-veins. 

Very variable in size and shape of leaf, and possibly more than one 
species are included under the name. C. P. 2449 (from Hunasgiriya) has 
almost rotund |., and in specimens from Dimbula they are narrowly 
oblong-lanceolate. This is the tree to which the name ‘Damba’ seems 
most usually applied by wood-cutters in the hill districts ; the wood is 
much used. 


16. &. cordifolia, Wight, 7/7. ii. 16 (1850). 

Herm. Mus. 24. Fl. Zeyl.n. 184. Myrtus androsemoides, L. Sp. Pl. 
472. Calyptranthes cordifolia, Moon Cat. 39. Syzygium cordifolium, 
‘Thw. Enum. 116. &. androsemoides, Bedd. Fl. Sylv. Anal. Gen. 107. 
C4 12, ihe), ACD, 

Fl. B. Ind. ii. 491. Wight, Ic. t. 544. 

A moderate-sized tree, twigs very stout, cylindrical, 
glabrous; |. very large, 4-8 in., sessile, very broadly oval, 
cordate and somewhat amplexicaul at base, rounded or very 
shortly and bluntly acuminate at apex, often much re- 
volute at margin, thick and leathery, shining above, mid- 
rib depressed above, very prominent beneath, lat. veins 
numerous, conspicuous; fl. nearly sessile, rather large, few, 
cymes terminal and axillary, usually much shorter than 1|.; 
cal.-tube shortly turbinate, much expanded at mouth, thick, 
segm. broadly triangular, obtuse; pet calyptrate ; fruit large, 


Eugenia. Myrtacee. yp 


2in., globular, crowned with the broad neck-like cal.-tube 
and spreading segm. 

Moist region, extending from the low country up to 4ooo ft. or more ; 
rather rare. Kalutara (Moon); Colombo (Ferguson) ; Ambagamuwa ; 
Adam’s Peak ; Ratnapura; Pasdun Korale. Fl. March-June ; white. 

Endemic. 

A very distinct species, certainly AZyrtus androsemoides of Linnzus. 
The plants from Pasdun Korale and (C. P. 2622) Adam’s Peak have 
longer and more pointed leaves than the type, and smaller flowers. 

Moon gives the native name ‘ Panu-keera,’ which really applies to 
£. Neesiana ; Hermann gives no native name. 


17. E. Neesiana, Wig/t, ///. ii. 15 (1850). Panukéra, 5S. 
Syzygium Neestanum, Arn. Pug. 17; Thw. Enum. 117. C. P. 735, 


4013. 

FI. B. Ind. ii. 493. Wight, Ic. t. 533. 

A very large tree, bark smooth, reddish-brown coming off 
in very thin flakes, branchlets terete; 1. 2}—4 in., nearly sessile, 
lanceolate-oblong, rounded or somewhat cordate at base, shortly 
acuminate, obtuse, glabrous and shining on both sides, lat. 
veins numerous parallel; fl.small, cymes terminal and axillary, 
small, lax, with divaricate angular branches, ped. 4 as long as 
cal.; cal.-tube turbinate; pet. calyptrate; fruit about } in, 
nearly globular. 

Moist low country ; common. FI. March, April; white. 

Endemic. 


Rather variable in width of 1.; C. P. 4o13 is a broader-leaved form 
with somewhat larger flowers. Wood useful. 


18. E. cyclophylla, 7hw. zx Fl. B. Ind. ii. 494 (1878). 

CP, 3015. 

Fl. B. Ind. ii. 494. 

A small tree, bark flaking off in small pieces, twigs stout, 
cylindrical; |. 2-3 in., sub-sessile, rotundate, as broad as long, 
rounded at base and apex, margin usually very strongly 
revolute or cucullate (the end being also recurved), very thick, 
stiffly coriaceous, midrib very broad and prominent beneath, 
lat. veins coarse, conspicuous, marginal vein doubled, glands 
black, conspicuous; fl. sessile, in clusters, cymes terminal, 
shorter than |., peduncle very stout, quadrangular ; fruit small, 
about } in., globose, crowned with cal.-limb. 

Upper montane zone ; very rare. Only found on ascent to Adam’s 


Peak from Maskeliya; first collected in 1866. FI. March; white. 
Endemic. 


19. B. rotundifolia, Wight, ///. ii. 17 (1850). 

Syzygium rolundifolium, Arn. Pug. 17; Thw. Enum. 118. C, P. 1587, 
Fl. B. Ind. ii. 494. 
PART II. N 


178 Myrtacee. [Zugenia. 


A small tree or shrub, with smooth reddish-grey bark, 
much branched, twigs very numerous, quadrangular; 1. 
numerous, persistent, closely placed, small, $—? in., orbicular, 
often retuse at apex, stiff and coriaceous, shining above, paler 
and with the numerous lat. veins slightly prominent beneath, 
petiole under $in., slender; fl. sessile, 2 or 3 together, cymes 
small, crowded, terminal, scarcely exceeding |.; fruit nearly 
4 in., nearly globose, dull reddish-purple. 


Upper montane zone down to 6000 ft.; very common. FI. April; 
pinkish-white. 

Endemic. 

Known readily from Z£. sclerophylla by its distinctly though shortly 
stalked leaves. This and the next, with their dense flat-topped heads of 
evergreen foliage, are very characteristic of our highest hill jungle. 


20. BE. sclerophylla, Duth. in Fl. B. Ind. ii. 494 (1878). 

Syzygium sclerophyllum, Thw. Enum. 118. C. P. 274. 

Fl. B. Ind. ii. 494. 

A small or moderate-sized much-branched tree, twigs very 
numerous, quadrangular, with prominent leaf-scars; |. very 
numerous and very persistent, sessile, 3-1 in., broadly oval or 
rotundate, rounded or very slightly cordate at base, rounded at 
apex, smooth and shining above, paler and with the numerous 
lat. veins prominent beneath; fl. sessile in threes, cymes ter- 
minal, slightly exceeding 1., branches stout, quadrangular ; 
fruit about 4 in., ovoid-globular, crowned with cal.-limb. 


Upper montane zone down to 6000 ft.; common. FI. Feb.—April ; 
white. 


Endemic. 


21. BE. oligantha, Duth. in Fl. B. Ind. 11. 494 (1878). 

Syzygium oliganthum, Thw. Enum. 118. C. P. 452. 

Fl. B. Ind. 11. 494. 

A bush or small much-branched tree, twigs very numerous, 
slender, quadrangular ; |. ?-1} in., obovate, tapering to base, 
rounded and often emarginate at apex, shining above, con- 
spicuously dotted beneath with the lat. veins scarcely visible, 
petiole g in.; fl. few, very small, sessile in twos or threes on 
terminal or rarely axillary peduncles much shorter than 1.; 
cal.-tube pyriform, segm. ovate, acute; fruit about 4in., globose, 
dark red. 


Lower montane zone; rather rare. Ambagamuwa; Hantane ; 
Maskeliya. Fl. March, April; white. 
Endemic. 


22. E. olivifolia, Duth. in Fl. B. Ind. ii. 495 (1878). 


Syzygium spathulatum, Thw. Enum. 118 (non Berg). C. P. 2493. 
Fl. B. Ind. ii. 495. 


Lugenia.| Myrtacee. 179 


A small much-branched tree, twigs numerous, slender, 
quadrangular ; |. 1-17 in., spathulate-oval or spathulate-lan- 
ceolate, much tapering at base, shortly acuminate, obtuse at 
apex, smooth and shining above, paler and somewhat glau- 
cous beneath and with oblique rather conspicuous lat. veins, 
petiole 4 in. or more; fl. very small, on slender ped. as long 
as cal., in copious, erect, slender, lax, axillary and terminal 
cymes about as long as |.; cal.-limb cup-shaped, seem. shallow ; 
pet. sometimes expanding ; fruit 4 in., globose, purplish-red. 

Montane zone, rather common ; at lower elevations, rare. Deltota ; 


top of Nillowe Kande, Pasdun Korale. Fl. July—Sept.; pinkish-white. 
Endemic. 


23. E. operculata, ox). Hort. Beng. 37 (1813). Bata-damba, 
Kobo-mal, S. 

Calyptranthes caryophyliifolia, Moon Cat. 39. Syzygium nervosum, 
DC., Thw. Enum. 417, 116. C. P. 2801. 

Fl. B. Ind. ii. 498. Wight, Ic. t. 552. 

A large tree with spreading branches, bark on trunk rather 
thick, grey and exfoliating, on branches smooth and nearly 
white, twigs cylindrical, slightly thickened at nodes, somewhat 
compressed ; |. 4-6 in., oval or broadly lanceolate, tapering to 
base, shortly acuminate, obtuse, stiff, shining on both sides, 
bright green above, paler beneath, lat. veins few, curved, rather 
prominent, pellucid, petiole about }in.; fl. sessile in threes, 
cymes large, lax, pedunculate, spreading, 3-4 in. long, coming 
from axils of fallen 1.; cal. ovoid-turbinate, becoming cam- 
panulate, smooth and shining, mouth truncate; pet. calyp- 
trate, the cap pointed; fruit 2in., nearly globular, purple, 
juicy. 

Moist region up to 4000 ft.; common. FI. Feb.-April; cream- 
coloured. 

Also in Himalaya, Assam, Burma, S. China, Malay Islands. 

Wood greyish-yellow, moderately heavy and hard, durable. The 
leaves when bruised have an agreeable aromatic odour. ‘The fruit is 
not eaten in Ceylon. 


24. HE. Jambolana,* Lam. Dict. iii. 198 (1789). MWaha-dan, 
Ma-dan, 5. Naval, Peru naval, 7. 

Herm. Mus. 8. Burm. Thes. 197. FI. Zeyl.n.185. AZyrtus Cuminz, 
L. Sp. Pl. 471. Calyptranthes Cumini, Moon Cat. 39. Syzygium Jambo- 
lanum, DC. Thw. Enum. 417, 116. C. P. 3644. 

Fl. B. Ind. 11. 499. Wight, Ic. t. 535. Bedd. Fl. Sylv. t. 197. 

A large or very large tree, bark grey or nearly white, thick, 
rough, exfoliating, twigs cylindrical-compressed ; |. 3-34 in., 


* Jambolana, the Portuguese name for the fruit, from /amun, tle 
Indian one. 


180 Myrtacee. [Eugenia.. 


oval or oblong-oval, tapering to base, acute or subacute at 
apex, glabrous, not shining, rather thin, light bright green, 
lat. veins very numerous, faintly marked, petiole {-4 in.; fl. 
small, on short ped., cymes about 3 in., lax, pedunculate, with 
slender divaricate branches, coming off from axils of fallen 1., 
often at some distance down the branches ; cal. shortly turbi- 
nate, limb cup-shaped, truncate or with very obscure segm.;. 
pet. calyptrate, cap not pointed; fruit 3-4 in., usually ovoid,. 
often lop-sided, crowned with truncate cal.-limb. 


Var. 6B, microcarpa, 7hw. /.c. Var. caryophyllifolia, Duth. in. 
HESBeind eas Wishtalert 55S 0s Camel 5on 


L. ovate-oval, acuminate, berries smaller, globose. 


Low country, in both wet and especially in dry regions, attaining to: 
3000 ft. in former; var. 8 the commoner form. Fl. May—August; white, 
honey-scented. 

Throughout the Tropics of Asia. 

I do not think this is ever cultivated in Ceylon ; but the fruit, which is 
generally no bigger than a large pea, is much eaten. We do not appear 
to have in Ceylon any of the large-fruited sorts found in India. 

The leaves have a pleasant aromatic scent when bruised. Wood 
reddish-grey, not heavy, moderately hard, durable, much used, often 
called ‘ Méni-damba.’ 

Jambolifera pedunculata, Gaertn. Fruct. i. 178 (non Vahl) is a form of 
this with oblong fruit, also figured in Wight, Ic. t. 620 (Z£. obtusdfolia, 
Roxb.). On the confusion by Linnzeus and subsequent writers between. 
this species and Acronychia laurifolia see under that plant (Pt. I. p. 216). 


25. E. lucida, Lam. Dict. iii. 205 (1789). [PLATE XXXVII.] 

£. hypoleuca, Thwaites in Bedd. For. Man. cxii. (name only). 
Drim Syst. Cat; 33.- > C, P3865: 

IMs dss Uae ih, Feige: 

A much-branched bush with rough silvery grey bark, 
twigs cylindrical, compressed, young parts hairy-pubescent ; 
I, 13-25 in., rotundate or broadly oval, tapering at base, 
rounded at apex, usually very strongly revolute at margin, 
dark green and polished above, purple when young, yellow- 
white ‘beneath, very stiff and coriaceous, petiole 1-3 in., stout; 
fl. large, 14 in. few, solitary, usually from the base. of the 
young shoot below the l., peduncle 4-14 in., stiff, pubescent, 
with 2 small oblong bracts immediately beneath f1.; cal.- 
tube shallow, cup-shaped, pubescent outside, segm. very large,, 
orbicular-ovate, finely ciliate, unequal, imbricate; pet. large, 
rounded, 1-14in.; stam. inserted on very broad disk; fruit 
over 1 in., globular, crowned with large erect cal.-segm., seeds. 
2 or 3, very large, brownish-white. 

Montane zone, 4500-6000 ft., in open sunny places; rare. Only in 


the North-eastern mountain mass. Knuckles ; Wattakelle ; Kobonilla ;, 
Rangala Hills. Fl. April and September ; white. 


LEugenia.| MM ‘yrtacee. 181 


Also found in Mauritius and Bourbon, but nowhere in Asia. 

The fruits are ripe at the same time as the flowers are expanded ; 
they are precisely like small rosy-cheeked apples. The flowers are 
handsome, and the largest in the genus. 

I follow the Fl. B. Ind. in referring this to Z. Zucéda, but Lamarck 
describes his plant as having sessile flowers. 


26. E. Heckeliana,* 777m. in Journ. Bot. xxiii. 207 (1885). 


A large bush or small tree, bark reddish, young shoots 
cylindrical compressed, floccose with fulvous tomentum ; 
1. large, 4-6 in., oblong-ovate, more or less cordate at base, 
subacute at apex, margin somewhat undulate, when young 
densely clothed on both sides with yellow wool which wears 
off with age leaving adult |. quite glabrous above and slightly 
floccose near midrib beneath, lat. veins rather distant, pro- 
minent, petiole very short, stout; fl. large, 14 in., solitary, on 
short, stout fulvous-woolly ped. with two lanceolate acute 
bracts at summit, coming off from axils of two opposite bracts 
(scale-leaves) at base of shoots below the new I.; cal.-tube 
campanulate-turbinate, as long as ped., densely fulvous-hairy, 
segm. large, ovate, subacute, somewhat recurved ; pet. rather 
longer than cal.-segm., oval; staminal disk large, square, 
densely hairy ; fruit (not seen ripe) globose, crowned by 
erect cal.-segm. 

Moist low country; very rare. Only found among rocks by the sea- 
eure at Wéligama in the extreme south of the island. Fl. Dec.; pinkish- 
white. 

Endemic. 


The leaves have a very strong midrib, which is usually somewhat 
curved, and the leaf is often stiffly conduplicate. 


CO el alata Thw. Enum. 114 (1859). 
Ee Pfapish 
Fl. B. Ind ii. 503. Bedd. Ic. t. 283. 

A moderate-sized tree, branchlets slender, cylindrical, 
young parts with ferrugineous tomentum ; 1. 3-4 in., varying 
from narrowly lanceolate to broadly oval, acute at base, 
caudate-acuminate, obtuse at apex, glabrous, thin, lat. veins 
few, fine, conspicuous beneath, uniting at a considerable 
distance within the margin, petiole }—? in.; fl. on very short 
rusty-pubescent ped., solitary or in small axillary clusters ; 
cal,-tube turbinate, rusty-pubescent, segm. large, ovate-oblong, 
obtuse; pet. rather longer than cal.-segm., oblong, obtuse ; 
fruit rather over ? in., globose, crowned with spreading cal.- 
segm., rusty-pubescent when young. 

* Commemorates Prof. Ernst Haeckel of Jena, who spent six weeks 
at Wéligama shortly before I collected this plant there. 


182 Myrtacee. [Zugenia.. 


Moist low country ; rare. Ratnapura and Reigam Korale (Thwaites) ;. 
Ambagamuwa. Fl. Jan.-March; greenish-white. 
Endemic. 


28. E. xanthocarpa, 7hiw. Enum. 416 (1864). 

C2 IP, Asay. 

Fl. B. Ind. 11. 503. 

A large bush or very small tree, with grey bark, twigs: 
cylindrical, young parts with rusty-brown pubescence ; 1. 3-5 
in., oblong-oval, acute at base, more or less acuminate, obtuse. 
at apex, glabrous, light green above, pale greenish-white 
beneath, lat. veins numerous, rather conspicuous, uniting near 
the margin; fl. on short rusty-hairy ped. with two small 
linear bracts at the top, solitary or in small axillary cymes of 
three or five; cal.-tube canipanulate, rusty-hairy, segm. large, 
oval, obtuse; pet. much longer than cal.-segm., obovate- 
oblong, obtuse ; fruit } in., depressed-globose, often somewhat. 
lobed, crowned with erect cal.-segm., finely pilose, bright 
orange. 

Moist low country ; very rare. Only found near Bentota, on the 
S.W. coast. Fl. November—March ; white. 

Endemic. 

The flowers have a singular creosote-like scent. 


29. E. bracteata, Rox). Hort. Beng. 37 (1813). Tembiliya, S: 
Kaya, Venkalikaya, 7. 

£). zeylanica, Roxb. (non Willd.), Moon Cat. 38; Arn. Pug. 18. Z. 
Willdenoviz, Wight, Thw. Enum. 114 (non DC.).  C. P. 1586, 1590. 

FE By Ind. i 502, Hook: Journ Bot. 1) t) 124) (2) Roxizz2na)s 
Wight, Ic. t. 545 (2. Willdenoviz). 

A much-branched shrub or small tree, with very smooth. 
yellowish-grey bark, young twigs cylindrical, densely covered 
with rusty pubescence ; |. 1?-3 in., oblong-lanceolate or oval, 
tapering to base, obtuse at apex, often somewhat acuminate, 
glabrous, paler beneath, veins inconspicuous, petiole 1-2 in.;. 
fl. usually solitary, often 2-7 in corymbose cymes, ped. rather 
long, stiff, rufous-pubescent, with two small linear bracts at 
summit; cal.-tube campanulate, rufous-pubescent, segm. 
oblong-ovate, obtuse; pet. large, ovate-oblong, pubescent out- 
side, ciliate, somewhat reflexed ; fruit 4-2 in., usually globose, 
sometimes pear-shaped, crowned with cal.-segm., orange- 
yellow or red. 
ae region, especially near the coast ; verycommon. FI. Feb.—May;. 
white. : 

Also in Southern India. 

A showy species when in full blossom; variable in size of leaf and 


flower. I fail to find any means of clearly distinguishing, even as a 
variety, . Roxburghiz, DC. (£. zeylanica Roxb., E. Willdenovit, Wight). 


Eugenia.) Myrtaceae. 183 


By the original descriptions the flowers should be more numerous than in 
£. bracteata, but there is much confusion in later authors. Further 
critical examination is required. 

A specimen from Colombo (Ferguson) has much longer leaves, and 
bears a note stating that it has globular red fruit ‘quite different’ from 
poe of ordinary £. dracteata, but it is too incomplete for certain identi- 

cation. 

The foliage is very like that of a MJemecylon, and the Tamils give it 
the same name as J. wmbellatum, with which it often grows. The leaves 
are aromatic when bruised. Wood greyish-yellow, hard, close-grained. 


30. &. rufo-fulva, 7iw. Enum. 416 (1864). 

©. Py 3835: 

Fl. B. Ind. ii. 503. 

A moderate-sized tree, bark pale brown or nearly white, 
young shoots cylindrical, slender, elongated, rufous-pubescent ; 
]. 24-43 in., lanceolate or linear-lanceolate, acute or tapering 
at base, very long caudate-acuminate, acute at apex, smooth 
and shining above, densely covered with very fine brownish- 
red tomentum beneath (becoming glabrous when old), petiole 
4-2 in., slender, curved ; fl. small, ped. slender, 4-4 in., pubes- 
cent, I-3 in leaf-axils ; cal.-segm. longer than tube, oblong, 
obtuse, tomentose outside; pet. oblong, obtuse, longer than 
cal.-segm.; fruit globular, under } in., pubescent, crowned 
with spreading cal.-segm. 

Moist low country; rare. Gattehatte, near Avisawella; Reigam 
Korale; Nillowe Kande. Fl]. March—June. 

Endemic. 

Varies with very narrow leaves. 


31. E. phillyrzoides, 772m. in Journ. Bot. xxiii. 207 (1885). 


A much-branched dense twiggy shrub, with roughish pale- 
brown bark, young parts with white hair; 1. very small, barely 
I in., numerous, crowded, erect, linear-lanceolate, tapering to 
base, acuminate, obtuse, margin slightly revolute, dark green 
above, paler beneath, lat. veins inconspicuous, petiole short ; 
fl. (not seen) solitary, axillary, peduncle shorter than 1.; fruit 
2-1 in., depressed-globose, flat-topped, crowned by wide sta- 
minal disk, and large rather unequal spreading cal.-segm., 
glabrous when mature (pubescent when young), red. 

Montane zone; very rare. Only found at summit of Kalupahane 
Kande, Lagalla, East Matale (about 5000 ft.). Not seen in flower. 

Endemic. 

Approaches £. Rottleriana, W. and A., figured (in flower only) in 
Wight, Ic. t. 100, of the hills of S. India, with which it requires careful 
comparison. 


32. B. floccifera, 7hiw. Enum. 115 (1859). 
C. P,; 463; 
Fl. B. Ind. ii. 504. 


134 Myrtacez. | [Zugenia 


A small tree, with brown cracked bark, branchlets terete, 
densely covered with soft brown tomentum ; 1. large, 4-7 in., 
lanceolate-oblong, acute at base, long-acuminate, acute at 
apex, densely brown-tomentose when young, becoming gla- 
brous when old, lat. veins fine, not conspicuous, petiole 2 in.; 
fl. rather large, nearly 1in., on very short tomentose ped., 
solitary or in fascicles of 2-4, axillary; cal. densely fulvous- 
tomentose, segm. over }in., linear-lanceolate, acute, reflexed ; 
pet. about as long, oval, obtuse ; fruit not seen. 

Moist low country ; very rare. Reigam Korale (Thwaites). FI. bluish- 


white. 
Endemic. 


33. B. rivulorum, 7iw. Enum. 115 (1859). 

C, IPs Suu. 

Fl. B. Ind. ii. 504. 

A small tree, bark smooth, quite white, branchlets cylin- 
drical, glabrous, young parts with a very minute purplish 
tomentum ; |. very large, 6-12 in., oval or ovate-oval, rounded 
at base, acuminate, bright green above, pale yellowish-green 
beneath, thick and leathery, veins strongly marked, indented 
on upper surface, prominent beneath, especially the intra- 
marginal one, petiole 4-3 in., very thick; fl. large, over 1 in., 
sessile, few in axillary fascicles, chiefly on old wood; cal.- 
tube campanulate, faintly pilose, segm. broad, rotundate, 
spreading ; pet. nearly }in., oblong; fruit large, 1 in., irregu- 
larly globose, pericarp thick fleshy, dark brown, wrinkled. 

Wet forests in the moist low country; very rare. Singhe Rajah Forest; 
yous Forest. Fl. March, April; pet. pale violet-pink, stam. white. 

ndemic. 


The large flowers are often borne even at the very base of the old 
stems, close to the ground, giving the plant a singular appearance. 


34. E. fulva, 7hw. Enum. 115 (1859). 

CAE. 3008: 

Fl. B. Ind. ii. 504. 

A small tree, young shoots cylindrical, compressed, clothed 
with fine orange tomentum ; 1. 4-5 in., oval, acute or rounded 
at base, shortly acuminate, glabrous and shining above, finely 
yellow-tomentose, becoming glabrous beneath, lat. veins few, 
conspicuous, uniting much within the margin, petiole 4in., 
channelled ; fl. 2 in. wide, sessile, solitary or 2 or 3 together 
in axils of present or fallen 1.; cal. fulvous-hairy, seem. ovate- 
triangular, acute; fruit globose, fulvous-tomentose, crowned 
with enlarged cal.-segm. 

Moist low country; very rare. Hewesse; Morowak Korale. FI. 


Sept.—Dec.; white. 
Endemic. 


Eugenia.) Myrtacee. 185 


35. BE. insignis, 7iw. Enum. 416 (1864). 

© P. 3677. 

Fl. B. Ind. ii. 504. 

A small tree, bark pale, twigs cylindrical, compressed, 
young parts covered with white tomentum, 1. distant, large, 
6-8 in., oblong-oval, rounded at base, suddenly acuminate, 
obtuse, glabrous on both sides, pale beneath, lat. veins few, 
distant, prominent beneath, uniting with the very strongly 
marked intramarginal vein at some distance from edge; fi. 
sessile, about ? in.; cal. densely covered with white hair, segm. 
broadly oval ; pet. longer than cal.-segm., oblong-oval, ciliate; 
fruit over I in., nearly globular, densely covered with a felt of 
white hair, capped with cal.-segm. 

Moist low country; very rare. Galpatta, Reigam Korale, is the 
pe the C. P. specimens, and I have seen no others. FI. white. 

ndemic. 


The intramarginal veins are so strongly marked and so distant from 
the edge that the leaf looks almost 3-nerved. 


36. E. decora, 7iw. Enum. 115 (1859). 

C. P. 3545. _ 

FI]. B. Ind. ii. 504. 

A small tree with smooth grey bark, twigs cylindrical, 
thickened at nodes, young parts glabrous; |. 4-5 in., oval, 
rounded at base, shortly acuminate, obtuse, glabrous on both 
sides, shining above, paler beneath, lat. veins not close, rather 
prominent, marginal veins double; fl. quite sessile, in small 
axillary clusters; cal.-segm. rounded, pilose ; fruit depressed, 
broader than long, # in. wide, 2-seeded. 

Moist low country; very rare. Only found near Galle. FI. Sept.; 
white. 

Endemic. 

This is somewhat obscure. I have seen only the C. P. specimens, 
which are perhaps of two different species, and quite insufficient. The 
above description applies to those which Thwaites seems to have 
intended by the name ; the others have almost precisely similar leaves, 
but the fl. are smaller and on slender pedicels has long. There is but 
a single separate fruit. 


37. B. rotundata, 777. 

£. Mooniana, var. B, Thw. Enum. 114. £. amena, Thw., var. 
voltundata, Trim. Syst. Cat. 33. C. P., 2803. 

A much-branched bush with whitish bark, twigs numerous, 
cylindrical, glabrous ; 1. small, 1-2 (rarely 3) in., usually rotun- 
date or very broadly oval, rarely narrowly oval, rounded at 
base, very obtusely acuminate, stiff and rigid, paler and copi- 
ously dotted beneath, veins inconspicuous, petiole very short, 


186 Myrtaceae. [_Hugenia. 


thick ; fl. small, sessile, in very small, axillary, and terminal 
fascicles, often on old wood ; cal. slightly pilose, segm. rounded; 
pet. oval, much longer than cal.-segm., fruit 2-4 in., globular, 
scarlet. 


Rocky hills in the low country, especially in the intermediate region. 
Deltota; Uma-oya; Doluwe Kande; Lagalla; summit of Ritigala, 
abundant. Fl. May-September ; pinkish-white. 

Endemic. 

This is combined with Z. amena in F1. B. Ind., but it seems nearer 
EE. Mooniana, under which Thwaites puts it. 


38. BE. mabzeoides, Wight, 11. ii. 13 (1850). 

Thw. Enum. 114. C. P. 445, 1588. 

Fl. B. Ind. ii. 505. 

A shrub, very much branched, bark brownish-grey, twigs 
very numerous, cylindrical, compressed, young parts glabrous; 
1, $-14 in., broadly or narrowly oval-spathulate, tapering to 
base, rounded at apex, very thick, stiff and rigid, dark green 
and shining above, paler beneath, petiole very short; fl. very 
small on usually short slender ped. solitary or in small 
axillary or terminal clusters; cal. very glandular, segm. 
rounded, finely ciliate; fruit 2-} in., globular or somewhat 
ovoid, capped with small cal.-segm. 

Montane zone, 4000-7000 ft.; rather common. FI. Sept., Oct.; pale 
green. 

Endemic. 

There are specimens in Herb. Kew, from ‘J. Watson, 1834’ (Hb. 
Griffith n. 2362). The leaves are often very small and C. P. 1588 is this 
form, which is found at the higher elevations. 


39. E. aprica, 7777. 

A much-branched bush, twigs cylindrical compressed, 
young parts glabrous ; 1. 1-2} in., rotundate or very broadly 
oval, suddenly and shortly tapering to base, rounded or very 
obtuse at apex, margin more or less revolute, rigid, pale 
beneath, and with rather prominent veins, petiole about ¢ in.; 
fl. small, 3-4 in., on short ped., usually solitary from base of a 
contracted branchlet, and appearing extra-axillary ; cal.-segm. 
obtuse; fruit 2-5 in., ovoid, capped with small cal.-segm., 
bright red. 


Open places in the montane zone; very rare. Apparently confined to 
the N.E. mountain mass. Knuckles Mountains (Ferguson); summit of 
Rangala Ridge. Fl. April; white. 

Endemic. 

I regret making another species, especially as my specimens are. 
somewhat insufficient; but I cannot fit them into any existing one. 


40. &. amoena, Zhiw. Anum. 114 (1859). 
C. P. 3439. _ : 
Fl. B. Ind. 11. 505 (in part). 


Eugenia] Myrtacee. 187 


A. small tree, branchlets terete glabrous; 1. large, 34-6 in., 
quite sessile, ovate-oblong, cordate at base, tapering into long 
acuminate, obtuse apex, father thick, veins inconspicuous ; fl. 
about # in., on very slender clabrous ped. about # in. long, in 
clusters of 2-10 between the terminal pair of 1., bracts be- 
neath the fl. very small; cal. glabrous, segm. broadly oval, 
obtuse ; pet. ovate, acute, scarcely twice length of cal.; fruit 
not seen. 

Moist low country; very rare. Hapugodde, Kukul Korale; Rakwane; 
Dolosbagie up to 1500 ft. (Thwaites). Fl. April and September; white. 


Endemic. 
A singular species with the habit of Safrosma indicum. 


41. EB. pedunculata, 777m. 72 Journ. Bot. xxvii. 162 (1889). 

A shrub, much branched, twigs cylindrical, young parts 
glabrous, |. 14-2}in., broadly oval, shortly tapering to base, 
shortly and very obtusely acuminate, thick bright green, paler 
beneath, lat. veins rather SURIEFOU, more conspicuous above 
than beneath, petiole 4-1in.; fl. rather large, ?in. or a little 
more, few, solitary, on straight, slender, slightly pilose ped. 
3-24 in. long, coming off from base of young shoots below the 
l. (and at first appearing as if in clusters); cal.-tube ovoid- 
turbinate, pubescent, segm. ovoid, obtuse, finely ciliate, re- 
flexed ; pet. large, spreading or reflexed; fruit not seen. 

Montane zone; very rare. As yet only found on summit of the 


Rangala Ridge at about 5000 ft. Fl. Sept.; white. 
Endemic. 


42. &. Mooniana, Wight, Lil, ii. 13 (1850), zon Gardn.* Pini- 
baru, 5. 

faw. Enum, 114. C. P. 11, '365. 

Fl. B. Ind. ii. 505. Wight, Ic. t. 551 (not good). 

A shrub or small tree, twigs numerous, slender, cylindrical ; 
glabrous ; 1. variable in size and shape 29] in. usually oval or 
rhomboid- oval, tapering to base, more or less acuminate, 
obtuse at apex, thin, green on both Oe, veins fine, incon- 
spicuous, petiole short, slender ; fl. 2-4 in., on very slender 
ped. about } in. long, in fascicles Se base of (often sup- 
pressed) branchlets (apparently axillary and terminal) ; cal.- 
tube fusiform, constricted at top, segm. lanceolate, acute or 
obtuse ; pet. oval-oblong, obtuse, often reflexed ; fruit }—} in., 
usually ovoid, rarely globular, pendulous, scarlet or crimson. 

Moist and intermediate country up to 4000 ft.; common. Fl. March 
and April, and September; white. 

Also in Southern India. 


Gardner’s £. Mooniana is a Brazilian species ‘dedicated to Mrs. 
Moon, of Rio Janeiro.’ 


188 Myrtacez. [Barringtonia. 


Though collected by Moon, he did not name or include it in his 
Catalogue. C. P. 365 from Medamahanuwara has larger and thicker 
leaves, and sometimes nearly sessile flowers, and approaches £. votundata. 

Wood yellow, smooth, hard, durable. From a large-leaved var. called 
‘ Géta-kaha,’ S. (found at Haputale), the very hard and heavy black sticks 
universally carried by Tamil coolies are obtained. They call them 
Karumbadikambu and Karutamurutukambu, and the black colour is 
produced by burial in mud for some months. 


E. Micheliz, Lam. (MV yrtus brasiliana, L., E. zeylanica, Willd. (non 
Roxb., nec Wight), £. Wzl/denoviz, DC. (non Wight), is occasionally met 
with in gardens, and known as the ‘ Brazil cherry,’ or ‘ Rata-jambu.’ 
Linneeus’s name, £. wvzflora, which included both this and a form of 
£. mataccensis, should be abandoned (see Journ. Linn. Soc. xxiv. 142). 
This is the original species of Ezgenza, it is a very ancient introduction 
to the East from the New World. 


43. &. Thwaitesii, Duth. in Fl. B. Ind. ii. 506 (1879). 
E.. concinna, Thw. Enum. 416 (non Phil.). C. P. 2802. 
Fl. B. Ind. ii. 506. 


A tree with whitish bark, twigs cylindrical, young parts 
pilose ; 1. 3-5 in., ovate-oval, slightly narrowed at base, cau- 
date-acuminate, obtuse at apex, glabrous on both sides when 
mature, lat. veins numerous, rather conspicuous, petiole under 
din., stout; fl. small, on slender ped. #-1 in. long, in apparent 
fascicles (really on suppressed branchlets), fruit about ? in., 
globular, crimson. 

Moist country; very rare. Only from Ambagamuwa. FI. Jan. 

Endemic. 

3. BARRINGTONIA,* Forsz. 

Trees, |. alt., stip. very minute, caducous, fl. in long ra- 
-cemes; cal.-tube adnate to ov., not produced beyond it, segm. 
2-4; pet. 4 (or 5), imbricate, usually slightly connate at base 
and adnate to stam., stam. very numerous in several rows, 
epigynous, connate at base into a thick tube, fil. very long ; 
ov. inferior, 2—4-celled, with several pendulous ovules in each 
cell, style long, simple ; fruit fibrous or leathery, indehiscent, 
1-celled ; embryo without obvious cotyledons (macropodous) 
showing on transverse section central and peripheral portions, 
no endosperm.—Sp. 20; 10.in FZ. B. Ind. 


‘Cal. quite closed in bud, splitting irregularly into 2 
or 3 segm. 
L. entire - : : 5 5 : : ; 1. 1B. SPECIOSAS 
L. finely serrate. 
Fruit broadly ovoid . 
Fruit narrowly oblong 
Cal. not closed in bud, segm. 4 


. B. RACEMOSA. 
. B. ZEYLANICA. 
B. ACUTANGULA. 


wh 


* Commemorates the Hon. Daines Barrington, F.R.S. Died 1800. 


Barringonia.] ¢ Myrtacee. 189: 


I. B. speciosa, forst. Char. Gen. Plant. 76 (1776). WMtudilla, S. 

Moon Cat. pt. 2, 29. Thw. Enum. 119. Agasta indica, Miers in 
fags. inn, Soc: Ser. 2,1. 61. * 

Fl. B. Ind. ii. 507. Wight, Ic. 547 (not good). Trans. Linn. Soc. 
l. c. t. 12 (Agasta indica). 


A rather small or moderate-sized tree with a close round 
head, bark pale grey, branchlets very stout, marked with promi- 
nent leaf-scars, young parts glabrous; 1. very large, 10-12in,, 
crowded, sessile, obovate, tapering to base, rounded at apex, 
entire, perfectly glabrous, polished and shining on both sides, 
midrib very stout and broad; fl. very large, about 7 in. diam., 
on very stout glabrous ped. 3-4 in. long, in a terminal erect 
raceme, buds nearly globular, apiculate ; cal.-tube about } in., 
bluntly quadrangular, glabrous, limb veiny, splitting into 2 or 
3 concave segm., pet. usually 4 (rarely 5), very slightly con- 
nate, about 24in., broadly oval, concave; staminal ring 
slightly adnate to base of pet., fil. 3-4 in., erect ; ov. inferior, 
4-celled, ovules 6-8 in each cell, style as long as stam.; fruit 
large, depressed, about 4in. high, quadrangular, bluntly 
pointed, crowned with persistent cal.-lobes, angles usually 
acute, pericarp very thick, smooth and shining, pale brownish- 
yellow, texture light fibrous-spongy, with strong fibres round 
the seed ; seed over 2 in., ovoid. 


Seashores; very rare. On the south coast from Galle to Matara, 
possibly native; certainly planted elsewhere. FI. May, creamy-white, 
odorous, filaments and style pink at top. 

Also on the coasts of the Andamans, at Singapore, and generally in 
the Malay Archipelago and Polynesia to N. Queensland. 

Thwaites gives C. P. 3610 for this, but in Hb. Perad. that number is 
LB. racemosa, and there is no specimen of B. sfeciosa. A beautiful tree, 
much planted for ornament on the coast. If this were an old inhabitant 
here, it could scarcely have escaped notice by Hermann ; as, however, 
it is one of those trees the seeds of which are carried by sea-currents, it 
may well have been brought to our southern shores by this natural 
agency. (See Tennent, ‘Ceylon,’ ii. 100.) 

First collected in the Pacific Is., and Miers (I. c. t. 10) figures the 
original plant of which the fruit is broadly ovate-ovoid and very slightly 
quadrangular. 


2. B. racemosa, 2/. ix DC. Prod. iii. 288 (1828). Diya- 
midella, 5S. 

Fl. Zeyl.n. 191. Lugenia racemosa, L. Sp. Pl. 471. Stravadia rubra, 
Moon Cat. 39. Thw. Enum. 119. Sutonica racemosa, Juss., Miers, 
Lc. 66. C€..P, 3610, 

Fl. B. Ind. ii. 507 (not given for Ceylon). Wight, Ic. t. 152. 


A small tree with long drooping branches, bark grey with 
prominent leaf-scars; |. large, crowded at ends of branches, 
sub-sessile, 6-12 in., obovate-oval, tapering to base, acute, 


190 Myrtacee. | Barringtonia. 


finely crenate-serrate, glabrous and shining on both sides, 
somewhat bullate between the strongly marked veins; ff. 
large, 2} in. diam., on stout minutely puberulous divaricate 
ped. +2 in. long, laxly arranged in flexible pendulous 
racemes I-2 ft. long, and terminal or from axils on the old 
wood, buds ovoid, bluntly pointed; cal.-tube turbinate, 
puberulous, limb splitting irregularly into 2 or 3 unequal segm., 
one of which often carries with it the top of the cal. , leaving 
the others truncate ; pet. connate at base, about I in., oval or 
oblong-oval, spreading ; fil. rather over I in., erect spreading ; 
ovules 3 or 4 in each ov.-cell ; fruit 2-24 in. by 1? in. wide, 
ovoid or oblong-ovoid, crowned with persistent cal.-segm., 
circular on section or very bluntly 4-lobed, pericarp leathery, 
thick, brownish-crimson; seed 1# in. 


Moist low country, especially near the coast, on the shores of back- 
waters, lakes, &c., but also inland; common. Conspicuous round 
Colombo Lake. FI. nearly all the year; cream-coloured, fil. pink or 
crimson, very slightly scented. 

Also on Malabar Coast and in Malaya and Polynesia. 

There are two varieties of flower in this, one with the filaments 
pinkish-cream-coloured, the other with them crimson, the darker-flowered 
plant has also usually smaller leaves. 

Hermann has a good figure (but no specimen), and from this Linnzeus 
took his description. 


3. B. zeylanica, Gardn. in Fl. B. Ind. ii. 508 (1879). Goda-= 
midella, 5S. 

B. racemosa, var. 8, Thw. Enum. 119. Butonica zeylanica, Miers in 
dans) Winns SOG. useiZ2./7-un Ow aZOB2" 

Fl. B. Ind. ii. 508. 


Bark grey, rough, marked with prominent leaf-scars; 1. 
smaller than in 3B. vacemosa, 4-7 in., sub-sessile, obovate- 
lanceolate, much tapering to base, acuminate, acute, Due 
serrate, clabrous, rather thin; f1. about 4 in. diam., ped. 3 4-2 
in., rather slender, divaricate, buds nearly globose ; cal.-tube 
turbinate, limb irregularly splitting into 2-4 unequal segm.; 
fil, about # in.; fruit about 3 in. by less than ? in. wide, 
narrowly oblong, tapering to base, crowned with cal.-segm., 
obtusely 4-angled ; seed about 14 in. long. 


Moist low country; rare. Galle, 1844 (Gardner); Ratnapura ; 
Gilimalle. Fl. March. 

Endemic. 

I have never met with this in a living state, and only know it from the 
C. P. specimens from which the above description is taken ; it seems a 
good species. It is probably Stravadia integrifolia, Moon Cat. 39, from 
Kalutara, but of that I have not seen authentic specimens. 

The astringent bark is used in medicine. 


Careya.] Myrtacee. IQI 


4. B. acutangula, Gaertn. Fruct. ii. 97 (1791). Gla-midella, S. 
Adampu, 7: 

Fl. Zeyl.n. 190. Eugenia acutangula, L. Sp. Pl. 471. Stravadium 
obtusangulum, Bl. in Fl. des Serres, vii. (1851) 24. Thw. Enum. 11g. 
Po rhg2: 

Fl. B. Ind. ii. 508. Gaertn, J. c. t. 101 (fruit only). Bedd. Fl. Sylv. 
+. 204. Trans. Linn. Soc., series 2, i. t. 17 (Stravadium acutangulum). 

A small tree with thick trunk, bark brownish grey, rough, 
young parts glabrous; 1. 3-5 in., obovate- or oblong-oval, 
tapering to base, subacute at apex, crenate-serrate, jptg nt 
green, not shining, venation reticulate pellucid, petiole {3 in.; 
fl. about 1 in. diam. on short spreading ped., racemes Q— 12in., 
terminal, pendulous ; cal. not closed in bud, tube very short, 
segm. 4 regular, broadly oval, rounded, finely ciliate, pet. 
small, about + in., fil. about 2 in: fruit ee in., oblong-ovoid 
somewhat narrowed to base, truncate at both ends, bluntly 
quadrangular, capped with small cal.-segm. 

Dry region, round margins of tanks and similar places; rather 
common. Jaffna; Mannar; Trincomalie; Kantalai; Batticaloa. FI. 
June-August; pet. cream-coloured, stam. dark bright crimson. 

Also in Peninsular India, Malaya, and N. Australia. 

Wood white, moderately heavy, even-grained. The fruit is chewed as 
a remedy for sore throat; it is also considered efficacious in rheumatism. 

Miers (I.c. 81) maintains as a separate species S. obtusangulum, Bl., 
and quotes for it the C. P. number above given. I can find nothing to 
warrant the separation; the angles of the fruit are always obtuse, as 
shown in the excellent original figure of Gaertner. 


4. CAREYA,* Roxb. 


Tree, |. alternate, fl. in crowded erect spikes; cal.-tube 
adnate to ov. and not produced beyond it, segm. 4, imbricate; 
pet. 4, imbricate ; stam. very numerous, epigynous, in many 
rows, connate at base, fil. very long; ov. inferior, 4-celled, 
with numerous ovules in 2 rows on axile placentas, style long, 
simple; fruit large, fleshy, indehiscent, with numerous seeds 
immersed in the flesh ; embryo without cotyledons (macropo- 
dous), no endosperm.—Sp. 3; all in “7. B. Ind. 


C. arborea, fox. Hort. Beng. 54 (1814). Kahata, S. Ka- 
chaddai, 7’. 

Thw. Enum. 119. C, P. 3169. 

Fl. B, Ind. ii. 510 (not given for Ceylon). Wight, Ill. tt. 99, 100, 
copied in Bedd. FI. Sylv. A z05 (poor). 


% Named. after Dr. William ‘Carey, the missionary, superintendent of 
the Botanic Garden at Serampore, and editor of Roxburgh’s ora 
Indica. Died 1834. 


192 Melastomacee. 


A small or middle-sized tree, with very thick, rough, dark- 
grey bark and a small rounded head, young branches with 
-very prominent leaf-scars, growing parts quite glabrous; 1. large,. 
6-12 in., usually sessile, broadly obovate, much tapering to 
base, very obtuse or rounded at apex, denticulate-crenate,. 
thick, smooth and shining on both sides, pale green, veins 
pellucid ; fl. large, 34-4 in. diam., sessile, crowded in very 
thick, swollen, hard terminal spikes, each with a central oval 
bract and two lateral linear ones ; cal.-tube about I in., cam- 
panulate, glabrous, segm. rounded, stiff, erect; pet. 2-24 in.,., 
ovate, obtuse or acute, margin often revolute; fil. about as. 
long as pet., spreading ; style a little longer than stam. ; fruit 
24-3 in., globular, apple-like, green, glabrous, crowned with 
persistent cal.-segm. and style, solid, with the cells nearly 
obliterated, seeds several immersed in the flesh. 

Moist region, especially on exposed patana land, up to 5000 ft.; very 
common. Also rarely in the dry region, ¢.g., Mulliativu. FI. Nov.— 
March; pet. pale green, fil. pale pink. 

This is the ‘Patana Oak’ of the English, and almost the only tree 
(besides Phyllanthus Emblica) on the barren grassy patanas. The 
remark in Fl. B. Ind., ‘No example from Ceylon’ (which, doubtless, has. 
reference merely to the Kew Herbarium), is calculated to mislead, as this 
is one of our most familiar trees, 

Moon has this only under the Sinhalese name (Cat. pt. 2, 8). I find 
no earlier record for it. 

Heart-wood dark reddish-brown, heavy, moderately hard, even-grained,, 
very durable. The bark is very astringent, and is much used in medicine: 
as well as for tanning. The inner bark gives a strong fibre. 


LITI—MELASTOMACE. 


TREES, shrubs, or herbs, |. opp., fl. regular, bisexual ; cal.-tube 
adnate to whole or base of ov., and more or less prolonged 
above it, segm. 3, 4, or 5; pet. same number as cal.-segm., 
contorted in bud; stam. 3, 8, or 10 inserted at edge of cal.- 
limb, anth. large, opening by 1 or 2 terminal pores, connec- 
tive often with appendages at base; ov. wholly or half in- 
ferior, 3- 4- or 5-celled (1-celled in Wemecylon) with numerous 
ovules on axile (or free central) placentas, style simple; fruit 
a dehiscent capsule or an indehiscent fleshy or dry berry, 
many-seeded (1-seeded in MWemecylon); seed without endo- 
sperm, cotyledons small (larger and much folded in 
Memecylon). 


Osbeckia.] Melastomacec. 193 


Seeds numerous. 
Fruit capsular. 
Pet. 4 or 5. 
Shrubs or herbs. 


Stam. equal I. OSBECKIA, 
Stam. unequal 2. MELASTOMA. 
ae climber 3. KENDRICKIA. 
Pee 4. SONERILA. 
Fruit baccate 5. MEDINILLA. 
‘Seed solitary (Memecylee) 6. MEMECYLON. 


The most recent account of Ris leeee Order ‘(which is mainly 
American) is by Cogniaux in Monograph. Phanerog. vi. (1891). 

Of our 52 species (of which more than half belong to one genus, 
Memecylon) only one, Osbeckia zeylanica, is found to be restricted to the 
dry region; the rest are all natives of the low country or montane zone of 
the moist region, many species of Osdeckia and Sonerzla being very 
characteristic of the higher hills; and only Osdeckia aspera and 
Memecylon umbellatum and M. capitellatum extend also into the dry 
country. No less than 35 of our species (including 20 of Memecylon) 
appear to be endemic. 


1. OSBECKIA,* Z. 


Shrubs or herbs, generally bristly-hairy; 1. opp., 3-7- 
nerved, fl. usually large, in small terminal cymes; cal.-tube 
usually campanulate, adnate to lower half of ov. and pro- 
longed above it, generally covered with hairs, scales or stellate 
processes, segm. 4 or 5, usually deciduous; generally with 
small bristle-pointed teeth alternating with them ; stam. 8 or 
10, equal, anth. large, obtuse or attenuate or beaked, con- 
nective not produced at base, but often with 2 tubercles there; 
ov. 4-inferior, 4-5-celled, with very numerous ovules in each 
cell ; fruit capsular, enclosed in cal.-tube and partially adnate 
to it below, dehiscent loculicidally from apex by pores or 
short valves, seeds very numerous, kidney-shaped——Sp. 51 
(Cogniaux); 26 in Fl. B. Ind. 

Pet. 4. 
Anth. not beaked. 
Stems prostrate, softly hairy 
Erect shrub, scabrous 
Anth. beaked ; annual herb 
Pet. 5. 
Cal.-tube with hairs, scales, or bristles. 
Cal. with simple hair, or hairy or fringed scales. 
L. hairy above. 


O. ERYTHROCEPHALA. 
O. RHEEDII. 
O. ZEYLANICA. 


WN eS 


L. 5-nerved ; : ; : . 4. O, ASPERA. 
L. 3-nerved - ; 7 ; . 5. O. WALKERL. 


L. glabrous above P . ; rede OF BUXIFOLIA, 


* In honour of Peter . Osbeck, who travelled and collected in 1 Japan 
and China in 1751. 


PART Ul. O 


194 Melastomacea. [ Osbeckia. 


Cal. with stellate stalked processes. 


Much-branched bush . : : . 7. O. RUBICUNDA. 
Prostrate herb. : : : . 6. Os MOONmE 
Cal.-tube nearly glabrous . : E . 9. O. OCTANDRA. 


1. O. erythrocephala, Naud. in Ann. Sc. Nat., ser. 3, xiv. 58 (1850). 
Bowitiya, S. 

O. truncata, Arn. (non Don) and O. parvifolia, Arn. in Hook. Comp. 
B. Mag. 11. 308. O. Leschenaultiana, Thw. Enum. 104 (non DC.). Cogn. 
Mon. 328. C. P. 1576, 284. 

Fl. B. Ind. ii. 514 (O. cupularis, Don, var.) and 515 (O. parvifolia). 
Wight, Ic. t. 996 (O. Leschenaultiana). 

Stems numerous, prostrate or ascending, long, cylindrical 
below, quadrangular above, slightly branched, reddish-purple, 
covered with long spreading hair ; |. variable in size, 4-14 in., 
broadly oval, rounded at base, acute at apex, finely ciliate- 
serrate, densely and softly hairy on both sides, 5-nerved, 
petiole Lin. hairy; fl. 1-14 in., on short ped., cymes sessile, 
dense, capitulate; cal.-tube urceolate, thickly set with simple 
bristly hair below and with stalked stellate processes above ; 
cal.-segm. 4, triangular, ciliate, with a large stellate tuft at 
apex ; pet. 4, roundish ; stam. 8, anth. blunt; ov. with coarse 
erect bristles on top. 

Montane region, especially in upper zone; very common. FI. all the 
year; pale pink. 

Also in S. India ? 

An abundant plant on the open patanas and variable in the amount of 
hair on stem and leaves. A small form, O. darvifolia, Arn. (originally 
based on a specimen of Walker, n. 332) is figured in Bot. Reg. t. 565 
under the name O. zeylanica, and with the fl. pure white. C. P. 284 
(from N. Eliya) has much longer anthers, and the flowers occasionally 
5-merous; it was regarded by Thwaites as probably a hybrid with 
O. rubicunda or O. aspera. 

This never turns black in drying, as stated in the books. 

I have thought it best to retain Naudin’s specific name, which refers 
certainly to the Ceylon plant; as this has almost invariably 4-merous f1., 
whilst O. cupularzs, Don, of S. India, is described (W. and A. Prod. 323) 
as 5-merous. 


2. O. Rheedii, Gardn. in Thw. Enum. 104 (1859). 
thw. Enum od, Cogn. lve: 329, 2) P1120: 
1BIE 18}, Iiravel, iis ‘Siti. 


A small erect bush with numerous ascending branches, 
branchlets quadrangular, very scabrous with prickly hairs ; 
I. #-14in., lanceolate or oval, tapering to both ends, scabrous 
above when young, afterwards nearly smooth, scabrous 
beneath with small prickles, especially on nerves beneath, 
stiff and rigid, pale beneath, very prominently 3-nerved, 
petiole short, scabrous; fl. small, about #in., nearly sessile, in 


Osbeckia.] Melastomacee. 195 


threes ; cal. densely covered with simple coarse hair, segm. 
triangular, acute, about as long as tube, persistent ; pet. 4, 
broadly ovate, acute; anth. obtuse, with a single pore ; cap- 
sule small, depressed, dehiscent by 4 valves from apex. 

Upper montane zone; rare. Adam’s Peak (Gardner) ; Ambagamuwa 
(Thwaites); bed of upper Maskeliya River, abundant. Fl. March ; 
pinkish-white. 

Endemic. 

When dried, the leaves become coppery-brown above and golden- 


yellow beneath. 


3. O. zeylanica, L. f Suppl. Plant. 215 (1781). 

Moon Cat ar.) @hw: Enum. 104. Cogn. | ¢.326. C. P:1575- 

FL. B. Ind. ii. 517. Plukenet, Phytog. t. 173, f. 4. 

Annual erect herb, 3-12 in. high, with long slender ascend- 
ing quadrangular branches with a few adpressed bristly hairs; 
|. -14 in., oblong-lanceolate, acute at both ends, finely ciliate- 
serrate, with a few long scattered hairs on both surfaces, 
3-nerved, petiole $-} in., slender, hairy; fl. small, 4-2 in., on 
slender bristly ped. about half as long as cal.-tube, 3-8 to- 
gether in umbellate terminal cymes; cal.-tube long-campanu- 
late, slightly contracted above, ribbed, shaggy, with numerous 
large stellate bristly hairs with very long rays; segm. 4, linear- 
lanceolate, more than half the length of tube, membranous, 
keeled, ciliate, tipped with stellate bristles, persistent, reflexed 
in fruit; pet. 4; anth. terminating in a long slender beak. 

Dry region in grass; rather rare. Jaffna; Trincomalie; Batticaloa ; 
Bintenne (Nevill). Fl. March; mauve-coloured. 

Also in S. India. 


4. O. aspera, B/. in Flora, xxiv. 474 (1831). Bowitiya, S. 

Fl. Zeyl. n. 172. Herm. Mus. 14. Melastoma asperum, L. Sp. Pl. 
391; Moon Cat. 35. Thw. Enum. 105. C. P. 1568. 

FL. B. Ind. ii. 519. Burm. Thes. tt. 72, 73. Wight, Ic. t. 377 (small 


form). 

A small shrub, with numerous slender branchlets, twigs 
quadrangular, hispid, purple; 1. 14-2 in., oblong-lanceolate, 
rounded at base, acute or subacute, often slightly twisted at 
apex, finely dentate-crenate, slightly hispid on both sides, 
especially on veins beneath, 5-nerved, the lat. ones faint, mar- 
ginal, petiole 4-+ in.; fl. 14-1%in., cymes short; cal.-tube 
covered with simple bristly hairs, segm. triangular, about half 
as long as tube, obtuse, bristly at summit, recurved ; pet. 5, 
ovate-deltoid, truncate; anth. with attenuate end; capsule 
over J in., ovoid, truncate, scabrous. 

Var. 8, minor, 7riana, /. c. 55 (sp.). C. P. 1569 (part). 

L. very small, under % in., crowded, very hairy on both 


196 Melastomacec. [ Osbeckia. 


sides ; cal. with many stellate or pectinate hairs mixed with 
the simple ones, especially at top of segm. 

Var. y, Kleinii, Arz. /. c. 309 (sp.). C. P. 1569 (506). 

L. broader than in type; cal. usually densely covered with 
long bristles mixed with stellate hairs, and at the top with 
usually a few scales, terminating in a tuft of long hair; fi. 
generally larger, often 24 in. 


Var. 6, Wightiana, Benth. in Wall. Cat. 4060 (s.p.). Wight, Ic. t. 
998. €. P. 1570. 

L. oval, obtuse, more hairy, marginal veins stronger; upper 
part of cal. densely covered with scales, fringed by long 
bristles, and terminal by stellate bristly hairs ; fl. larger, 2 in. 
Or MOFe. 

Low country, and up to 4000 ft. very common. Var. £, very rare, 
East Matale only, collected in 1863. Var. y, principally in the dry 
country ; summit of Ritigala. Var. 6, lower montane zone; common. 
Fl. all the year; mauve colour. 

Also in Peninsular India. 

This is a very variable species, and O. Wzghtiana is kept as distinct 
in Fl]. B. Ind. C. P.506 connects it with the others. O. mnor, Triana, is 
based on specimens of C. P. 1569 which were called ‘forma minor’ by 
Thwaites; these Clarke does not appear to haveseen. Cogniaux (1. c. 315) 
keeps it up as a species, as he does also O. K/ezniz (l.c. 316) and O. 
Wightiana (1. c. 318). 

O. glauca, Benth., is recorded from Trincomalie in Wall. Cat. n. 4073 
(see Fl. B. Ind. ii. 519). It was cultivated at Kew, and is figured in Bot. 
Mag. t. 5085 (as O. aspera); the specimens in Kew Herb. from the Gardens 
are said to be originally from Ceylon. They have the leaves densely 
silky-hairy on both surfaces, but are scarcely more than a variety of 
O. aspera. 


5. O. Walkeri, Arn. in Hk. Comp. Bot. Mag. ii. 309 (1836). - 

Phy Enumtoss Ce eens 716 

Fl. B. Ind. i. 519. 

A. small shrub, twigs cylindrical, densely silky; 1. #-1 in., 
oval or lanceolate-oval, tapering to base, subacute, edges very 
slightly revolute or flat, closely adpressed-hairy above, silky, 
especially on nerves beneath, 3-nerved, petiole short, silky ; 
fl. 14-17 in., on short ped., solitary or in threes; cal. covered 
with long simple silky adpressed hair, segm. linear, erect, as 
long as tube ; pet. 5. 


Var B, Beckettii, 7iw. zn Triana, |. c. 54 (Lecheté by error), C. P. 
3908. L. smaller, 7-3 in., broadly oval, stiff, coriaceous. 


L. smaller, 4-4, broadly oval, stiff, coriaceous. 


Upper montane zone; rare. Horton Plains; Nuwara Eliya. Var £, 
Wattakelle Hill (Beckett); Knuckles Mountain. FI. November—Feb. ; 
mauve. 


Osbeckia.] Melastomacee. 197 


Endemic. 
Triana |. c. combines with var. 8 O. duxzfolia, var. mznor, to which the 
leaves have much resemblance. 


6. O. buxifolia, Arz. iz Hk. Comp. Bot. Mag. ii. 309 (1836). 
Melastoma buxifolia, Moon Cat. 35. Thw. Enum. 105. C. P. 1572. 
Fl. B. Ind. ii. 518. 

A flat-topped very much-branched bush 5-8 ft. high, 
branchlets densely covered with copious ferrugineous wool ; 
]. very numerous, crowded, small, 2-3 in., rotundate or broadly 
oval, rounded or subcordate at base, emarginate at apex, 
margin and apex revolute, upper surface striate with raised 
lines, nearly glabrous when mature, densely ferrugineous- 
woolly beneath, coriaceous, rigid, strongly 5- (or 7-) nerved, 
the nerves much depressed on upper surface, petioles short, 
with long rusty hair; fi. large 21 in., sessile, solitary or in 
threes, terminal on every twig, bracts leaf-like, close beneath 
and adpressed to cal.; cal. surrounded at base by long rufous 
sete, tube completely covered by scales bearing tufts of long 
shaggy rufous hair, seem. narrowly triangular, pectinate on 
margin, shaggy with hair outside, glabrous within, with dense 
tuft of very long bristles on top; pet. 5, finely woolly-ciliate. 


Var 6, minor, 7hw. /.c. (not O. minor, Triana), C. P. 2618. 


L. smaller, quite glabrous above with the veins less in- 
dented; cal.-tube densely covered with very long simple 
rufous woolly hair. 

Upper montane zone above 6000 ft.; rather common. Adam’s Peak 
(Moon); Pedurutalagala. Var. 8, Nuwara Eliya; Totapella; Adam’s. 
Peak. Fl. March, April, September; rich mauve colour. 

Endemic. 

There are tufts of persistent hair at each node between the leaf-bases, 
apparently stipular in nature. A most beautiful shrub, and very profuse 
in flowers. 


7. O. rubicunda, Arn. in Hk. Comp. Bot. Mag. ii. 309 (1836). 

Thw. Enum. 105.. C. P. 52. 

Fl. B. Ind. ii. 520. 

A large much-branched bush, bark pale yellowish-grey. 
twigs sub-quadrangular, very hispid with spreading scale-like 
hair; 1. 1-3 in., variable, oblong-oval oblong-lanceolate or 
somewhat ovate, rounded or subcordate at base, acute or sub- 
acute at apex, minutely spinous-serrate, scabrous-hairy on 
both sides, pale beneath, rather thick but not stiff, 5-nerved, 
petiole 4—# in., very hispid ; fl. very large, on very short ped., 
3-8 together crowded in terminal clusters ; cal.-tube thickly 
studded with large stalked processes with the summit ex- 
panded peltate long ciliate or stellate, segm. a little shorter 


198 Melastomacee. [ Osbeckia. 


than tube, triangular-lanceolate, acute, ciliate, tipped with a 
stellate process; pet. 5, 1-14 in. finely ciliate; fruit-cal. 
almost prickly with the hardened persistent bases of the 
stellate processes. 


Upper montane zone to 7000 ft.; common. About Nuwara Eliya 
abundant; Ramboda; Matale; Galagama; Elk Plains. Fl. Jan.—April, 
October; brilliant purplish-crimson. 

Endemic. 

A very showy plant, the flowers often over 23 in. across, and produced 
in great abundance. Variable in form and size of leaf. When withering 
the leaves turn a bright orange-scarlet on upper surface. 

O. elliptica, Naud, 1. c. 62, is referred to this by Thwaites and Triana, 
but Cogniaux (1. c. 320) maintains it as distinct, distinguished by 1. rounded 
at both ends and 3-nerved, and broadly triangular cal.-segm. 


8. O. Moonii, 7iw. Enum. 105 (1859). 

Melastoma repens, Moon Cat. 35. Thw. Enum. 105. Cogn. 1.c. 319. 
C. P. 2800. 

Fl Be Ind: i. 520: 

Herbaceous, stems long, prostrate, rooting at the nodes, 
bristly-hairy ; 1. oval 14-24 in., acute at both ends, entire, 
coarsely hairy on both sides, 5-nerved, petiole 4-3 in.; fi. 
rather small, about 1 in. on slender hairy ped., usually in 
threes, terminating short erect axillary branches from the 
prostrate stems, bracts small, rotundate; cal.-tube shortly 
campanulate, thickly studded with long spreading processes, 
which are bristly on the sides and expand at summit into a 
peltate head with long radiating bristles, segm. linear-lanceo- 
late nearly equalling tube, strongly ciliate and bristly, inter- 
mediate teeth conspicuous, very bristly ; pet. 5, ciliate; anth. 
moderately attenuate. 

Low moist country; rare. Colombo (Moon); Sittawaka; Paregodde; 


Kuruwita Korale. Fl. Sept._Jan.; purplish-pink. 
-Endemic. 


9. O. octandra, DC. Prod. iii. 142 (1828). 

Herm. Mus. 36. Fl. Zeyl. n. 173. MWelastoma octandra, L. Sp. Pl. 
391. O. polycephala, Naud. in Ann. Sc. Nat., ser. 3, xiv. 67. O. virgata, 
Don, Thw. Enum. 105. C. P. 507, 26109. 

Fl. B. Ind. ii. 521. Wight, Ic. t. 376 (not good). 

A small erect shrub, with many erect virgate branches, 
bark pale reddish-brown, flaking off in fibrous pieces, young 
shoots quadrangular, hispid with adpressed bristly hairs ; 
1. narrowly lanceolate or oblong-lanceolate, shortly tapering 
to base, subacute, very faintly serrate, ciliate, almost entirely 
glabrous above, glaucous green and with scattered long hairs 
beneath, 3-nerved, petiole  in., flat, hairy; fl. about 14 in., on 
stout hairy ped. about half length of cal.-tube, numerous in 
close corymbose or paniculate terminal cymes ; cal.-tube long- 


Melastoma.] Melastomacee. 199 


campanulate, faintly ribbed, usually perfectly glabrous, occa- 
sionally with scattered simple or stellate bristly hairs, segm. 
narrowly triangular, not half length of tube, ciliate, tipped 
with a bristle or large stellate hair, intermediate teeth large ; 
pet. 5, rounded, ciliate ; anth. attenuate ; ov. with a tuft of 
long hairs on summit. 

Moist low country and extending up to 5000 ft.; common. FI. all the 
year; pale purplish-pink, often nearly or quite white. 

Also in South India. 

O. polycephala, Naud., is based on C. P. 507 in Hb. Kew, and is 
maintained by Cogniaux (I. c. 314). I fail to see any difference from 
Spal O. octandra beyond the presence of a few stellate hairs on the 
cal.-tube. 

It is difficult to understand why Linnzeus should have given the name 
octandra to this 1o-stamened plant; Don’s later one is very much more 
appropriate. 


2. MELASTOMA,* Z. 


A large shrub with the young shoots bristly-hairy, 1. opp., 
fi. large in terminal cymes; cal.-tube adnate to base of ov. 
and prolonged beyond it, covered with simple scaly hairs, 
segm. 5, deciduous with minute bristle-pointed teeth between 
them ; pet. 5; stam. 10, unequal, those opp. cal.-segm. with 
purple anth. and the connective much produced below and 
ending in two blunt projections, those opp. pet. with yellow 
anth. and the connective very short with 2 small projecting 
lobes ; ov. 4-inferior, 5-celled with very numerous ovules on 
large axile placentas, fruit small, irregularly dehiscent, with 
very numerous seeds; seed small, kidney-shaped, embryo 
curved, no endosperm.—Sp. 37 (Cogniaux) ; 6 in FZ. B. Ind. 

M. malabathricum,t Z. SZ. Plant. 390 (1753). Maha- 
bowitiya, S. 

Herm. Mus. 10. Burm. Thes. 155. Fl. Zeyl.n. 171. Moon Cat. 35. 


Thw. Enum. 106. Cogn. Mon. 349. C.P. 1574. 
Fl. B. Ind. ii. 523. Wight, IIl. t. 95. 


A much-branched bush, with cylindrical branchlets at first 
covered with red bristly forward-pointing hairs, afterwards 
with thin yellowish fibrillose bark; |. 24-6 in., lanceolate, 


* From péhac, black, and o7éua, mouth, from the staining of the lips, 
which results from eating the fruit. The name first given by Burman, 
who took it from Hermann, who says the plant was called ‘ Bocca preto’ 
by the Portuguese for this reason. 

+ Linnzus seems to have thought this to be the source of the /olza 
malabathri, at one time much used as a medicine in Europe. This name 
of the old pharmacists was a corruption of the Indian one, Tamalapattra, 
and the leaves really those of Ciznamomum Tamala, trees. 


200 Melastomacee. [Kendrickia.. 


acute at both ends, faintly ciliate- or spinous-denticulate, 
slightly scabrous above with short adpressed hairs, hispid om 
the veins beneath, finely softly hairy between them, 5-nerved, 
the outer ones fine and close to margin, petiole 2-3 in., fl. 
large, 14-3 in., ped. about as long as cal.-tube, clothed with 
hair-scales, in trichotomous, few-flowered, terminal paniculate 
cymes; cal.-tube broadly campanulate, completely covered 
with adpressed, lanceolate, silvery hair-scales, segm. variable, 
linear- to broad-lanceolate usually about as long as tube, 
spreading, glabrous within; pet. narrowed to base, rounded 
or truncate; connectives of long anth. curved over the short. 
ones; ov. bristly-hairy on summit, style erect, longer thar 
stam. and standing over them; fruit softly coriaceous, about 
4in., broadly ovoid, bursting irregularly, usually in a circle, 
and exposing the 5 large purplish-black pulpy placentas 
covered with numerous minute seeds. 

Moist low country up to 3000 ft.; very common. FI. all the year;. 
violet-mauve-coloured. 

Throughout the Indian Peninsula. 

The pulp of the ripe fruit is edible, and Moon calls the plant the 
‘Black Strawberry tree.’ 


M. ellipticum, Naud. in Ann. Sc. Nat., ser. 3, xiii. 291, is from Ceylon, 
collected by Leschenault. From the description I cannot see how it. 
differs from JZ. malabathricum. M. Royenit, Bl. in Flora ii. 483, also. 
from Ceylon, is referred by Thwaites also to the present species, which, 
indeed, does not vary much, save in the cal. segments. 


3. RENDRICKIA,* ZZ. 7 

An epiphytic climbing shrub, 1. opp., fleshy, fl. large, im 
terminal umbels; cal. very fleshy, tube adherent to ov. and 
slightly prolonged beyond it, segm. 4; pet. 4, large, spreading, 
fleshy ; stam. 8, equal, anth. opening by a single pore, connec- 
tive produced at base into a short horn-like process; ov. 
inferior, 4-celled, with very numerous ovules, style simple, 
stout ; fruit a capsule, 1-celled by absorption of septa, split- 
ting from above into 4 valves; seeds very numerous, closely 
packed, prismatic.—Monotypic. 

i. Walkeri, (2. f in Benth., and Hk. f, Gen. Plant. i. 752 (1865). 
__ Medinilla (?) Walkeriz, Wight, Il. i. 217; Gardn. in Calc. Journ. 
us I ve Pachycentria Walkert, Thw. Enum. 107. Cogn. Mon. 469. 

Fl. B. Ind. i. 526. Bedd. Ic. t. 271 (very poor). 

* Named in honour of G. H. Kendrick Thwaites, Director of the 


Royal Botanical Gardens, Peradeniya, from 1849 to 1880. The name 


Thwattesia had been previously given in 1846 to a genus of Algez. Died 
II Sept. 1882. 


Sonerila.] Melastomacee. 201 


An epiphytic scandent shrub, the young stems climbing to. 
a great height by copious slender adventitious roots, branches 
cylindrical or young ones quadrangular, stout, bark pale 
yellow, smooth, shining, flaking off in rectangular pieces, 
young parts scurfy ; 1. on the young climbing branches small, 
I-14 in. lanceolate, acute, ciliate-dentate, on the flowering 
shoots larger, 2-24 in. oval or oblong, acute at base, obtuse, 
entire, glabrous, thick, 5-nerved, the lat. nerves joining the 
midrib, bright green and shining above, pale and dull beneath, 
petiole very short, stout; fl. large, 24-3 in. 3-8 together in 
terminal umbels, ped. 1-14 in., stout, bracts small, oblong, 
obtuse; cal.-tube turbinate, ovoid, glabrous, limb short 
spreading, seem. short, fleshy, pointed; pet. ovate-rotundate, 
fleshy, spreading ; top of ov. 4-lobed, glabrous, style longer 
than stam. 

Climbing over trunks of trees in the forests of the lower montane 
zone, and extending down to 2000 ft. in the Kukul Korale; common. 
Fl. January to May; bright clear rose colour, cal. red. 

Also found in the Anamallay Hills, S. India. 

A very lovely plant when in flower, and a great ornament to the lower 
hill-forests, reaching to the tops of high trees. The mode of climbing is 
precisely like that of ivy, the young shoots and leaves being pressed close 
to the bark and clinging to it by numerous rootlets, whilst the flowering 
branches stand away from the support. 

The leaves vary in form, and are sometimes suborbicular and rounded 
at the base. 


4. SONERILA,* Rox. 


Herbs often woody at base, |. opp., fl. in scorpioid ter- 
minal racemes; cal.-tube connate below with ov. and pro- 
longed beyond it, segm. 3, small, usually deciduous ; pet. 3, 
ovate, acute; stam. 3, anth. blunt or pointed or rostrate, 
opening by 2 pores, cells more or less divergent at base; ov. 
inferior, 3-celled, with numerous horizontal ovules; fruit a 
trigonous capsule, dehiscing at summit by 3 short valves, 
seeds very numerous, usually muriculate, with minute 
tubercles.—Sp. 70 (Cogniaux) ; 43 in /%. B. Ind. 

The species have been unduly multiplied by botanists; for Ceylon 
there are 15 given by Thwaites and 16 in FI. B. Ind., and Cogniaux 
makes as many as 21. In a recent paper by Stapf (Ann. Bot. vi. 291, 
1892), many of these are reduced to S. zeylanica and S. Wightiana, and 
he has clearly shown that the length of the anthers does not afford a 


sufficient character for separating species. 
It is p bxopaie: that natural hybrids not peace nnenthy) occur in this genus, 


* Sonert- ila, the native Malabar name as given by Rheede, 


202 Melastomacec. [ Sonerila. 


Stem erect. 
L. 3-7-nerved. 
Nerves from the base of leaf. 
L. stalked. 
L. glabrous beneath. 
Capsule small, as long as or shorter than ped. 


L. 5-nerved, unequal at base I. S. ZEYLANICA. 
Ik 3-nerved, equal at base. 6 2. S. RHOMBIFOLIA. 
Capsule large, much longer than ped . 3. S. BRUNONIS. 
L. more or less hairy beneath. 
L. glabrous above. 4. S. ARNOTTIANA. 
L. hairy above. 
L. oval : : 4 . 5. 5. WIGHTIANA. 
L. ovate-cordate ; ; : ._ 6. S. HERSUa UREA 
L. sessile or nearly so. 
L. hairy above. 
Capsule broadly ovoid . 7. S. GARDNERI. 
Capsule campanulate : : . 8. S. ROBUSTA. 
L. quite glabrous . : . 9. S. LANCEOLATA. 
Nerves from the midrib (penninerved) : . 10:9) PILOSWiene 
L. 1-nerved, linear. : : . II. S. LINEARIS. 
‘Stem creeping, I. penninerved ; , : . 12, S. PEDUNCULOSA. 


1. S. zeylanica, W. and A. Prod. 322 (1834). 

Arn. in Comp. Bot. Mag. ii. 307. Thw. Enum. 109. C. P. 349, 3345. 

Fl. B. Ind. ii. 530. 

An erect annual herb, 3-12 in. high, much branched above, 
branches quadrangular, glabrous; 1. 4-1 in., ovate-lanceolate, 
acute, sharply dentate-serrate, with conspicuous scattered 
bristly hairs above, glabrous and paler beneath, usually very 
unequal at base, 5-nerved, thin, petiole at least half as long 
as 1. slender; racemes 2-6-fld., pet. very acute; anth. short, 
blunt, or pointed; fruit with tapering ped. 3in., ped. less than 
half the length, narrowly turbinate, prominently 6-ribbed, 
glabrous. 

Var. B, pumila, Clarke. S. pumila, Thw. Enum. 109. Cogn. 1. c. 
Aon, (Cy IPs AOU, 

Smaller, branched from the base ; 1. broader, usually with 
white dots above and purple beneath; anth. very blunt; cap- 
sule with ribs less prominent. 


Var. y, cordifolia, 7hw. ms. S. cordifolia, Cogn. 1.c. 502. C. P. 
2893. 

L. cordate at DES, petioles longer than 1.; anth. pointed, 
capsule very short, 4 in., campanulate, shorter than ped. 

Var. 6, affinis, 77727. S. affinis, Arn. 1. c. 307. Thw. Enum. 109. 
C2 P5353) HISB inde iims 33: 

L. ovate, more coarsely toothed with spreading serrations; 
cal. slightly hairy; anth. attenuate; capsule campanulate, 


Sonerila.] Melastomacee. 203 


4 in., considerably shorter than curved long ped., the ribs not 
prominent. 


Var. «, rostrata, Clarke. S. rostrata, Thw. Enum. 108. C.P. 2644. 


L. less sharply serrate, anth. rostrate, long-acuminate, 
capsule campanulate under 2 in., rounded at base, rather 


shorter than ped., ribs slightly marked. 


Montane zone, and extending down into moist low country ; common. 
Var 8, very common at higher elevations. Var. y, only found as yet in 
Singhe Raja Forest. Var. 6, Watagoda. Var. «, Ambagamuwa; Hini- 
duma; Reigam Korale. FI. all the year; pink, var. 6 white. 

Endemic. 

Var. y, with its very short capsule, has some claim to specific rank. 
Clarke (1. c. 531) has another var., Walkere, differing from the type in its 
seeds being smooth without the usual minute elevations. 


2. S. rhombifolia, 7Zw. Enum. 108 (1859). 

S. glaberrima, Arn. in Comp. Bot. Mag. ii. 307 (ex Clarke). C. P. 2799. 

Fl, B. Ind. ii. 533. 

Stem 6-12 in., stout, quite glabrous, branches obscurely 
quadrangular ; |. #-14 in., broadly oval-ovate, often somewhat 
rhomboid, acute at both ends, shallowly crenate-serrate, 
upper portion quite glabrous, 3-nerved, equal at base, pale or 
purplish beneath, petiole half as long as |. or more; cal.-segm. 
large, triangular, acute; anth. acuminate; capsule with 
tapering ped. barely 2 in., of which ped. is about half, with 
strong ribs. 

Var. B, angustata, 7hw. ms. S. angustata, Triana, l.c. 408. 
Fl, B. Ind. 1. 536. 

L. lanceolate-oblong, the lat. nerves faint; cal.-segm. 
narrow, very acute. 

Moist low country; rare. Kukul Korale; Hiniduma; Adam’s Peak 
(Moon); Var. 8, Galle district (Fl. B. Ind.). Fl. Dec.; deep pink. 

Endemic. 

Triana, followed by Clarke, Cogniaux, and Stapf, considers var. 6 as 
a quite distinct species; but by the latter S. yhomébzfolza itself is regarded 
as merely robust S. zey/anica. Arnott’s name is the older, and, if certainly 


the same plant, should be adopted. Cogniaux (1. c. 499) keeps S. glaber- 
vima as distinct. 


3. S. Brunonis, W. and A. Prod. 321 (1834). 
Cogn. l. c. 493. C. P. 662 (in Hb. Kew). 
FI,.i3. Ind. 11. 531. Wight, Ill. t. 94; Ic. t. 1059. 


Stem 12 in., rather stout, branches quadrangular, nearly 
glabrous, |]. ovate on long petioles, bristly-serrate, 5—7-nerved 
at base; pet. lanceolate, anth. short, pointed; capsule, in- 
cluding much tapering ped., fully 4 in., of which ped. is about 
a third, very strongly ribbed. 


204 Melastomacec. [ Sonerila. 


Upper montane zone. ‘ Nuwara Eliya, 1846’ (Gardner). FI. ‘Jan 

Also in S. India. 

I do not know this species. The Kew specimens of C. P. 662, which 
are those presumably referred to in Fl. B. Ind., have very large capsules. 
Thw. afterwards merged C. P. 662 in C. P. 349 (5S. seylanica), and 
Cogniaux quotes the latter number, and also C. P. 353 (.S. afinzs) in part 
for S. Brunonzs. 

Wight (Ill. 1. c.) colours the petals a bright light blue! 


4. S. Arnottiana, 7iw. Enum. 108 (1859). 

(C31 AGIs 

DleBalnd i sar bedd slewing Orst. 400: 

Stem 1-2 ft., branched above, more or less glandular- 
hairy ; 1. 13-3 in., lanceolate, tapering to both ends, acute or 
obtuse at apex, more or less unequal at base, finely spinous- 
serrate, glabrous above, paler and with a few coarse hairs. 
on veins beneath, 3- or 5-nerved, petiole half as long as l., 
glandular-pilose ; racemes glandular-hairy, often forked ; cal.- 
tube glandular-hairy ; anth. short, pointed; capsule short, 
strongly ribbed, asperous. 

Var. 6, tomentella, Z7im. S. tomentella, Thw. Enum. 109. 
(Gy 1S Doo, 

Stem shorter, clothed with short spreading hair, not glan- 
dular; 1. smaller, very unequal at base, almost entire, with a 
few scattered hairs above, tomentose on the veins beneath ; 
racemes and cal. not glandular but densely and finely hairy ; 
anth. not pointed. 

Montane zone, 4000-7000 ft.; rather rare. Horton Plains; Bopata- 
lawa; Ambagamuwa; Maskeliya. Var. 6, Agrawatte; Ambagamuwa. 


Fl. March and April; pale pink. 
Also in Anamallays, S. India (?). 


5. S. Wightiana, Arz. in Comp. Bot. Mag. ii. 307 (1836). 

Cognwlc105 aa Cub 39072 

MM i, levels wis I 

Stems 1 ft. or more high, much branched, somewhat 
flexuose, rough with coarse rufous bristly hair; 1. 1-14 in, 
oval, very unequal at base, acute or subacute, sharply serrate, 
rough with scattered short bristly hairs on both sides, 5-nerved, 
petiole about 4 in., roughly hairy ; fl. rather large, ? in., ped. 
with strigose hair; cal.-tube very hairy, segm. oval, acute; 
pet. long-clawed, very acute; anth. short, very blunt; capsule 
4 in., strongly ribbed, scabrous. 


Var. 8, Hookeriana, 77im. S. Hookeriana, Arn. 1.c. 308. Thw. 
Iso, WO, (CG, 12, 17/3. 


Larger and semi-woody at base, stems more villose with 
longer hair; 1. less unequal at base, rounded or obtuse at 


Sonerila.] Melastomacee. 205 


apex ; cal.-tube slightly hairy, segm. often strongly ciliate 
anth. bluntly pointed. 

Montane zone; rare. Adam’s Peak; Dolosbagie; Var. 6 only at the 
foot of the cone of Adam’s Peak on the southern side. Fl. March and 
April; pale pink. - 

Endemic. 

I follow Stapf in combining these under one species. Thwaites con- 
sidered some of his specimens to be hybrids between the two. 


6. S. hirsutula, Arz. 7x Comp. Bot. Mag. ii. 307 (1836). [PLATE 
DOES] 

Thw. Enum. 108. Cogn. 1. c. 503. C. P. 276. 

Fl. B. Ind. ii. 534. 

Stems 2 ft. or more high, semi-woody at base, much 
branched, densely covered, especially at nodes, with long, stiff, 
spreading hair; 1. 1-14 in., broadly ovate, rounded or cordate 
at base, obtuse at apex, spinous-serrate, very hairy on both 
sides, usually 7- (sometimes 5-) nerved, petiole variable, often 
longer than 1., very hairy ; racemes short, few-flowered, hairy, 
fil. large, about I in.; cal. hairy ; anth. elongated, rostrate, the 
cells divaricate at base; capsule over + in. nearly smooth, 
ribs prominent. 


Upper montane zone; rare. Pedurutalagala; foot of Totapella; 
Horton Plains. Fl. Jan—April; deep rosy-pink or pale purplish. 

Endemic. 

A beautiful plant, the flowers nearly as large as those of S. Gardnerz. 


7 S. Gardneri, 7iw. Enum. 107 (1859). 

Boom lL e..496), GC. P2623. 

Bio, tad i. 532, Bedds lexind; Or ta208: 

Stem 1-14 ft. high, woody below, very slightly branched, 
densely covered with short, dark red, glandular hairs; 1. #-14 
in., very shortly stalked, broadly ovate, rounded or subcordate 
at the equal base, acute or obtuse at apex, finely spinous- 
serrate, hairy on both sides, paler beneath, 5- or 7-nerved ; 
raceme lax, glandular-hairy, fl. very large, over I in., cal. 
glandular-hairy ; anth. shortly acuminate, not beaked; cap- 
sule 2 in., broadly ovoid, faintly ribbed, glandular-pubescent. 

Var. B, firma, Thw. ex Triana, l.c. 75 (sp.). C. P. 3873. 

L. under } in., crowded and overlapping, quite sessile; fl. 
smaller, fruit not seen. 

Montane zone at about 5000 ft.; very rare. On the southern slope of 


Horton Plains near Galagama, first found by Gardner. Var. 6, on 
Wattekelle Hill; on Adam’s Peak. Fl. Feb., March; dark pink. 
Endemic. 


This and S. Azrsutula have the largest flowers of the Ceylon species. 
Very distinct by its capsule. A plant from Dumbanagala Hill, Rangala, 
seems to be this, but is not in fl. or fruit. 


206 Melastomacee. [ Sonerdla.. 


8. S. robusta, Arn. in Comp. Bot. Mag. ii. 308 (1836). 

Thw. Enum. 107. C. P. 426.* 

Fl. B. Ind. ii. 533 (including S. Harveyz). 

Stem arising from a short rootstock with thick fleshy 
scales, 1-2 ft. high, branched above, shaggy with long, 
spreading, stiff, rufous hair; 1. numerous, closely placed, often 
overlapping, 4-1 in., very shortly stalked or nearly sessile, 
broadly oval, rounded or subcordate at equal base, obtuse, 
finely serrate-ciliate, rufous-hairy above with long hairs, 
glaucous with scattered hairs beneath, strongly 5-nerved, 
often with two other fainter ones; raceme short, subumbellate, 
with long rufous hair; fl. rather large, under 1 in., anth. long- 
acuminate, capsule campanulate, strongly muriculated. 


Var. B, glabricaulis, 7hw. iz Fl. B. Ind.\.c. C. P. 3955. 


More slender, stems glabrous; 1. more rotundate, quite 
glabrous beneath, conspicuously 7-nerved, fl. smaller. 


Var. y Harveyi.t S. Harveyz, Thw. Enum. 107. S. Hartwegz, 
Triana, |. c. (errore typogr.) C. P. 2974. 


As in var. (3, but anth. short, apiculate or somewhat 


acuminate. 

Upper montane zone ; veryrare. Only found on Adam’s Peak, where 
it is abundant near the base of the cone. Var. 8, Bopatalawa. Var. y, 
between Maturata and Ramboda, near Palagala at about 6000 ft. FI. 
March and October; bright pink, calyx purple. 

Endemic. 

The type has some resemblance to 5S. Hookeriana, from which its. 
sessile leaves readily distinguish it. The remarkable scaly rootstock, in 
appearance like that of Lathrea, does not seem to be always present. 
Triana combines glabricaulis with S. ‘Hartwegi’ (= Harveyz), and they 
are scarcely distinguishable. 


9g. S. lanceolata, 7iw. Enum. 107 (1859). 

C, IP, Aeoy- 

lS Balnd si 535-)) bedd. le) IndVOr t 200: 

Stems 2-4 ft. with divaricate branches, quadrangular, quite 
glabrous ; 1. large, 2-34 in., sessile, or very nearly so, narrowly 
lanceolate, more or less cordate at base, attenuate and sub- 
acute at apex, faintly serrate in upper part, perfectly glabrous 
on both sides, 3-nerved ; fl. large, about I in., racemes long- 
stalked ; cal. glabrous, segm. triangular, acute, anth. acuminate, 
not rostrate ; capsule over } in., long-funnel-shaped, glabrous, 
with broad thick ribs. 


* Some of C. P. 426 is said in FI. B. Ind. to be S. Hookeriana. 

+ Commemorates W. H. Harvey, Professor of Botany in Dublin 
University, and a distinguished Algologist. He spent three months in 
Ceylon in 1853 on his way to Australia, and was with Thwaites when 
this plant was gathered. 


Sonerila.] Melastomacee. DOV 


Moist low country; very rare. Karawita Kande; Kukul Korale; 
Panadure. Fl. March and September; rose-coloured. 

Endemic. 

A handsome large plant, suitable for cultivation; very unlike the 


other species. 


10. S. pilosula, 7iw. Enum. 108. 

Gosn. Ic. 506. C. P.. 2687. 

Fl. B. Ind. ii. 534. 

Stem 6-12 in. high, with subterete pubescent branches ; 
], large, 2-34in., ovate-lanceolate, very unequal-sided at base, 
acuminate, obtuse, serrate, pubescent on both sides, flaccid, 
often purple beneath, 7-9-nerved, the lat. nerves coming off 
from the midrib above the base (and thus penninerved), 
petiole long, hairy; fl. about #in., cal. hairy; anth. long, acu- 
minate; capsule funnel-shaped, muriculate. 

Moist low country; very rare. Only found at Palabadala, Kuruwita 
Korale. Fl. March; dark rose-coloured. 


Endemic. 
The only Ceylon species with penninerved leaves except S. Jedunculosa. 


11. S. linearis, Hook. f. ex Triana in Trans. Linn. Soc. xxviii. 76 


(1871). 
S. Guneratnez,* Trim. in Journ. Bot. xxii. 208; Cogn. l.c. 498. C. P. 


o12. 

Fl. B. Ind. ii. 536 (not given for Ceylon). 

Stems erect, 12-15 in. high, woody below, cylindrical, with 
very numerous weak, slender, spreading branches, quadran- 
gular, quite glabrous, purplish red; 1. apparently in whorls of 
4, sub-sessile, $—? in., linear-strap-shaped, tapering at base, 
obtuse at apex, entire below, with a few distant sharp serra- 
tions in upper third, glabrous, thick, pale beneath, with very 
prominent midrib, no side nerves; fl. in smaJl very short 
recemes or solitary, about in, diam., ped. slender, glabrous, 
segm. fleshy, oblong-oval, obtuse; anth. acuminate but not 
rostrate; capsule small, under }in., funnel-shaped, much 
tapering at base, quite glabrous, strongly ribbed. 

Moist low country; very rare. Near Hewesse, Pasdun Korale (1878), 
ee Kande, Kukul Korale (1891). Fl. August, Sept.; bright mauve- 

Ink, 
e Also in Burma. 

The apparently whorled leaves consist of 2 pairs very closely inserted 
and decussate. When describing this in 1885 I had not had an oppor- 
tunity of comparing it with S./zearés. Since examining Lobb’s specimens 
in Herb. Kew from Moulmein, I am, however, convinced it is the same 
species. Cogniaux, however, maintains it as separate. 


* Commemorates Don Seman de Silva Guneratne, for many years 
plant-collector for the Peradeniya Botanic Gardens, and the discoverer of 


many novelties. He died in 1886. 


208 Melastomacee. [Medinilla. 


12. S. pedunculosa, 7iw. Enum. 109 (1859). 

Cognmincw403)) C2 P= 3001, 

Fl. B. Ind. 11. 531. 

Stems prostrate, rooting at the nodes, with long inter- 
nodes, glabrous ; 1. 14-2} in., broadly ovate, rounded or sub- 
cordate at base, obtuse or rounded at apex, finely crenate- 
serrate, with numerous scattered coarse hairs above, nearly 
glabrous beneath, penninerved, flaccid, often purplish beneath; 
racemes small, at end of long, erect, slightly glandular-pilose 
peduncles terminating short axillary branches; fl. small, $ in. 
diam.; cal. with scattered glandular hairs ; anth. short, acute; 
capsule small, under % in., turbinate, smooth, faintly ribbed. 

Moist low country below Iooo ft.; rare. Hiniduma; Mapalagama; 
Pasdun Korale. FI. Aug., Sept.; pinkish-violet. 

Also in Travancore. 


In a young state the leaves are crowded, and form a sort of rosette 
on the ground, and the peduncles look like scapes. 


[Marumia zeylanica, Bl.,is said to have been sent from Ceylon by 
Koenig and Van Royen. If so, the plant must have been cultivated here, 
but it is more probable that the specimens were collected at Malacca 
or in Java.] 


5. MEDINILGA, Gaud. 


Epiphytic shrubs, 1. opp., fleshy, 3- or 5-nerved, fi. few, 
axillary; cal.-tube campanulate, adnate to ov., fleshy, limb 
produced, segm. 4, obscure; pet. 4; stam. 8, equal, anth. with 
two short projections at base and a short spur behind ; ov. 
inferior, 4-celled, with numerous ovules, style simple; fruit 
soft, crowned with cal.-limb.; seeds numerous, ovoid.—Sp. 100 
(Cogniaux); 11 in FZ B. Ind. : 


Branches cylindrical, |. 5-nerved . ‘ 4 . 1. M. FUCHSIOIDES. 
Branches quadrangular, |. 3-nerved : : . 2. M. MACULATA. 


1. NM. fuchsioides, Gardun.in Calc. Journ. Nat. Hist. viii. 12 (1847). 

Thw. Enum. 106. C. P. 138. 

Fl. B. Ind. ii. 548. 

An epiphytic shrub, dichotomously branched, branches 
stout, cylindrical, marked with leaf-scars, bark smooth, greyish 
white ; 1. 2-3 in., rhomboid-lanceolate, narrowed to both ends, 
but rounded at the actual base, obtuse or subacute at apex, 
glabrous, paler and with very minute scattered glands be- 
neath, 5-nerved, the marginal pair faint, the others strongly 
marked, petiole very short, stout, crimson; fl. rather large, 
#in. long, solitary or 2 or 3 on a common peduncle, from 
axils of fallen 1., pendulous, ped. as long as cal., much thick- 


Memecylon.] Melastomacee. 209 


ened beneath fl, buds pointed; cal.-tube glabrous, segm. 
small, tooth-like; pet. erect, much imbricate, rotundate ; 
anth. acute; fruit about }in., subglobose, dark purple. 

On trees in the upper montane zone; rather rare. Nuwara Eliya; 
Horton Plains; Hakgala. Fl. Feb.; bright purplish-pink. 

Endemic. 


2. IM. maculata, Gardn. in Calc. Journ. Nat. Hist. viii. 13 (1847). 
[PLATE XL.] 

Thw. Enum. 106. C. P. 1578. 

Fl. B. Ind. ii. 548. 

An epiphytic shrub with a climbing stem, very thick at 
base, branches opposite, very numerous, quadrangular, the 
angles often with a narrow undulated wing, bark grey, on 
young shoots purplish; |. 1-2 in., variable, more or less oval, 
rounded or subcordate at base, obtuse, rounded or emargi- 
nate at apex, shallowly crenate, glabrous, dark green with 
purple blotches above, paler and with very minute glandular 
hairs beneath, somewhat fleshy, 3-nerved, petiole very short ; 
fi. small, arranged as in last; cal.-tube urceolate, glabrous, 
limb spreading, almost truncate, fleshy ; pet. reflexed ; fruit 
small, under 4 in., subglobose, truncate, red. 

Var. 6B, cuneata, 7hiw. Enum. l.c. Var. Thwattestz, Clarke in 
Fl. B. Ind. C. P. 3443. 

L. larger, 2-3 in. rotundate, tapering to a long cuneate 
base, abruptly and shortly acuminate or retuse; fl. and fruit 
not seen. 

Epiphytic on trees in lower montane zone; rathercommon. Hantane; 
Ambagamuwa; Ramboda; Matale; Hunasgiriya; Knuckles. Var. £, 
moist low country; very rare. Hewesse, Pasdun Korale. Fl. Nov.; 
pale pink. 

Endemic. 


Has much the appearance of a Loranthus. Var. B is very little known, 
and perhaps a different species. 


6. MEMECYLON,* Z. 


Shrubs or trees, |. opp., entire, glabrous; fl. small, in 
axillary simple or compound umbellate cymes; cal.-tube 
campanulate, adnate to ov., limb more or less dilated, segm. 4, 
often ill-defined; pet. 4; stam. 8, equal, anth. short, opening 
by slits, connective produced into a large horn below; ov. 
quite inferior, 1-celled, surmounted by a depressed epigynous 
disk usually marked with 8 radiating ridges, style simple, 


* The Greek name for the fruit of the Avdutus. It is not evident why 
Linnzus applied it to our ‘ Wélikaha’ (1. capztellatum). 


PART II. te 


DO) Melastomaceez. [Memecylon. 


ovules 8-16, on a free central placenta; fruit a small dry 
berry, more or less globose, crowned with cal.-limb, 1-seeded ; 
seed large, with a shell-like testa, cotyledons much folded, no 
endosperm.—Sp. 126 (Cogniaux); 4o in FZ. B. Ind. 


This is one of the most difficult genera in our flora. There are 
probably too many species in the following enumeration, and I had hoped 
to materially reduce them on a critical revision. This, however, I find it 
impossible to do on any satisfactory principle, and the plan followed in 
Fl. B. Ind. of placing a number as varieties of JZ. ede in no way simpli- 
fies matters. I find Thwaites’ method of grouping the Ceylon species 
much better than more recent ones, and have followed ithere. Many of 
the forms he collected I have not met with alive, and they are still 
represented only by the C. P. specimens and drawings. 

I have also several apparently undescribed species, but hesitate to 
publish further names in this genus. 


Leaves prominently 3-nerved. 
Twigs cylindrical E 5 5 . I. M. ARNOTTIANA. 
Twigs quadrangular . 2. M. GARDNERI. 
Leaves penninerved, sometimes slightly 
3-nerved at base. 
Epigynous disk smooth or with radiating 
ridges which are not winged. 
Twigs usually 4-winged. 
L. over 8 in., veins very conspicuous 3. M. HOOKERI. 
M 


L. under 4 in., veins obscure . 2 4- Vi \WitGHaDir 
Twigs 4-angled. 

Pander. ; , : . 5. M. PARVIFOLIUM. 

L. over I in. 


L. acute at base, cymes stalked. 


L. thick, cymes dense 6. M. VARIANS. 
L. thin, cymes few-flowered 7. M. ELEGANTULUM. 
L. obtuse at base, cymes sessile. 8. M. ELLIPTICUM. 


Twigs cylindrical (occasionally sub- 
quadrangular). 
L. cordate at base (see also vars. 
of 3 and 4). 
L. over 6 in., lanceolate-oblong 9. M. MACROPHYLLUM. 
L. under 4 in., rotundate. 
L. revolute at margin. 


Cymes sessile . i . Io. M. OVOIDEUM. 
Cymes pedunculate . . 1. M. REVOLUTUM. 
L. not revolute . : . 12. M. ORBICULARE. 
L. not cordate. 
L. over I in. 


L. rounded at base, over6in. 13. M. PROCERUM. 
L. acute or tapering at base 
(sometimes rounded in 15). 
L. rounded or obtuse at apex. 
L. obovate, sessile. . 14. M. CUNEATUM. 
L. oval, shortly stalked. 15. M. UMBELLATUM. 


Memecylon.| Melastomacee. 211 


L. acuminate or caudate. 
L. over 4 in., fruit yellow. 


L. lanceolate, thin . 16. M. CLARKEaNUM. 
L. narrowly linear- 
lanceolate thick . 17. M. DISCOLOR. 


L. under 3 in., fruit purple. 
L. ovate or oval. 
Ped.with 2 bractlets 
beneath fl. . . 18. M. FUSCESCENS. 
Ped. without bretlts. 19. M. ROSTRATUM. 
L. linear or lanceolate- 
line arene : . 20. M 
L. under 3 in., very crowded 
and numerous . ; S Big WML 
Epigynous disk with the ridges strongly 
winged (except sometimes in 26). 
Twigs quadrangular. 


. ANGUSTIFOLIUM. 


. PHYLLANTHIFOLIUM. 


L. over 3 in., fl. blue : : . 22. M. RHINOPHYLLUM. 
L. under 3 in, fl. white . : . 23. M. LEUCANTHUM. 
Twigs cylindrical. 
Fruit 2in. diam. . F : . 24. M. MACROCARPUM. 
Fruit under 2 in. diam. 
L. not 3-nerved at base, fl. stalked. 
Fl. large, buds pointed . . 25. M. LEVIGATUM. ~- 
Fl. small, buds globose . . 26. M. GRANDE. 
L. 3-nerved at base, fl. sessile . 27. M. CAPITELLATUM. 


1. M. Arnottianum, Wight in Thw. Enum. 113 (1859). 

Eugenia (?) capitellata, Arn. Pug. 17. C. P. 1589. 

FI. B. Ind. ii. 553. 

A small bush, bark pale brown, smooth, branchlets slender, 
cylindrical; 1. 14-2 in., ovate-oval, subacute or rounded at 
base, long-caudate-acuminate, obtuse at apex, pale beneath, 
conspicuously 3-nerved, purplish when young, petiole very 
short ; fl. very small, sessile, solitary or very few at apex of 
slender, erect, axillary peduncles #?-1 in. long; cal.-segm. 
large, triangular, acute; fruit about }in., globular, blue-black. 

Forests of moist low country ; rather rare. Pasdun Korale; Pelawatte, 
abundant ; Gilimale; Mapalagama. Fl. December-May; white. 

Endemic. 

The specimens from Gilimale have much larger leaves, 3-4 in. (var. 
grandifolia, Cogn. Mon. 1133). 

Called ‘ Pinibaru’ in Pasdun Korale, the name properly of Zugenza 
Moonii, which it somewhat resembles. 


2. M. Gardneri, 7iw. Enum. 113 (1859). 

M. Thwaitesiiz, Cogn. Mon. 1134. C. P. 2387. 

Fl. B. Ind. ii. 553. 

A small tree, bark yellowish-grey, smooth, branches much 
thickened at nodes, twigs quadrangular, slightly winged at 
angles ; |. 14-24 in., very shortly stalked or subsessile, usually 
broadly oval, but sometimes rhomboid-lanceolate or rotundate, 


2P Melastomacee. [Memecylon. 


narrowed to base, usually shortly acuminate, obtuse, thick, 
pellucid-punctate, very prominently 3-nerved; fl. very small, 
sessile, few, in axillary fascicles, peduncle very short or 0; 
cal.-tube campanulate, segm. broad; pet. oval, distant, disk 
without radiating ridges; fruit + in., purplish-black. 

Moist region, principally in the lower montane zone, but going down 
to 2000 ft.; rather rare. Dimbula; Maskeliya; Ambagamuwa; near 
Ratnapura. Fl. Jan.-April; white. 

Endemic. 

Cogniaux distinguishes his @. Thwazteszz by broader and more obtuse 
leaves, more coriaceous and drying of a pale colour, which characters 
appear to indicate the form at the higher elevations. 


3. M. Hookeri, 7iw. Enum. 113 (1859). Wevitiya-kéra, 5. 


(var. £). 
CocnmiNeniia7. GP 2680: 


Fl. B. Ind. ii. 554. 

A large shrub, 6-12 ft. slightly branched, bark grey, 
flaking off in square pieces, branchlets stout, strongly 4-angled 
with narrow wings on the angles; |. very large, 8-12 in, 
sessile, cordate at base, oblong-lanceolate, tapering to 
acuminate subacute apex, penninerved, veins very prominent 
beneath, lat. ones numerous, joining a very strong intra- 
marginal one at some distance from the edge; fl. rather large 
(for genus), numerous, ped. slender, twice as long as cal., sur- 
rounded at base with a little involucre of acute white bractlets 
and brown hair-like scales, very densely crowded in large 
axillary clusters on the old wood, buds sharp-pointed ; cal.- 
segm. broadly oval, acute; fruit large, 2 in. blackish-purple. 


Var. 3, exalatum, 771m. Syst. Cat. Ceyl. 35. C. P. 2686.* 


Branchlets perfectly cylindrical, not winged; ped. rather 
shorter. 

Moist low country; rare. By streams in Ambagamuwa at 1000 ft. 
Var. 8 about Colmobo; common. FI. July-November; purplish-blue, 
buds pink. 

Endemic. 

Clarke refers var. 6 to 17. macrophyllum, but the texture and venation 
of the leaves are entirely different in that species. 


4. M. Wightii, Zw. Enum. 113 (1859). 

M. depressum, Benth., Triana, l.c. 158. Cogn. 1.c. 1145. C. P. 2570. 

ID) 1835 Wawel, we SiS 

A small tree, branchlets slender, quadrangular, winged, 
the wings broader beneath each node; |. 24-44 in., sessile, or 
very nearly so, cordate at base, lanceolate-oblong, tapering 
to acuminate obtuse apex, margin more or less revolute, 
thick, shining above, penninerved, lat. veins obscure; fl. on 


Memecylon.] Melastomacee. oe 


slender ped. crowded in fascicles on axillary tubercles usually 
on the old wood, no peduncle; cal.-limb nearly truncate ; 
disk without rays; fruit small and conspicuously crowned 
with cal.-limb. 

Var. 6, cylindricum, 777. Wight, Ic. t. 279 (AZ. amplexicaule). 

Branchlets quite cylindrical, not winged. 

Lower montane zone; rather rare. Hewaheta; Maturata; Uda 
Pusselawe. Var. 6, Central Prov. (Thwaites). Fl. March and September; 
bluish-white. 

Also in S. India. 

Var. 8 appears to be precisely Wight’s JZ. amplexicaule, which, 
according to FJ. B. Ind., p. 559, is a distinct species. If that work be 
correct in saying that this is also JZ. amplexicaule of Roxburgh, that 
name should take precedence. 


5. M. parvifolium, 7/w. Enum. 113 (1859). 

M. varians, var. parvifolium, Trim. Syst. Cat. Ceyl. 35. C. P. 2955, 

8. 
fs Fl. B. Ind. ii. 556. 
A small tree, sometimes reaching 30 ft., with very numerous 
branchlets and grey finely cracked bark, twigs quadrangular; 
|. 3-1 in., broadly oval or obovate-oval, tapering to base, 
obtuse or rounded at apex, not at all acuminate, coriaceous, 
quite veinless (except midrib), with a very narrow cartilaginous 
margin, dark green and shining above, pale and dull beneath, 
petiole short but distinct; fl. few, small, ped. about as long 
as cal.-tube, cymes very small, nearly sessile ; cal.-tube short, 
limb spreading, cup-shaped, segm. broad, shallow, acute; rays 
of disk very indistinct; fruit about 4 in., tipped with very 
small cal.-limb, black. 


Forests of the upper montane zone; rather common. F]. May; white. 

Endemic. 

This is combined with JZ, varzans in F1. B. Ind., but seems distinct. 
Thwaites (I. c. 416) distinguishes C. P. 758 (from Ramboda) as a large- 
leafed form (17. Thomsoni, Thw. ms.) = var. Thomsonz, Triana. 

Much resembles some of the small-leaved montane Eugenias; the 
young foliage also, as in them, is red. Wood yellow, hard, heavy. 


6. M. varians, 7iw. Enum. 112 (1859). 

Cogn. Lc. 1164. C. P. 2927. 

Fl. B. Ind. ii. 556 (in part). 

A small tree, much branched, twigs quadrangular, slender; 
|, 2-34 in., ovate-lanceolate, acute or tapering at base, more or 
less caudate-acuminate, acute or obtuse at base, dull not 
shining, rather thick, veins very obscure, petiole very short ; 
fl. nearly sessile, in small axillary heads, peduncles very short ; 


2A: Melastomacec. [Wemecylon. 


cal.-limb shallow, truncate, disk-rays distinct; fruit 4 in., 
capped with small cal.-rim. 

Var. 8, rotundatum, 7iw./.c. M. rotundatum, Cogn. |. c. 1164. 
CHESZO50: 

L. sma.ler, I-2in., obtuse at both ends, often revolute at 
margin. 

Lower montane zone, and extending downwards to 2000 ft.; apparently 
rather common. Var. 68 at the higher elevations. Fl. April and Sep- 
tember; pale blue or white. 

Endemic (?). 

I do not well understand this species, and follow Thwaites in dividing 
it into two varieties which seem distinct; Cogniaux separates them 
as species. Both often have the branchlets cylindrical, except at the 
summit of the internodes. The leaves turn a pale bright greenish-orange 
when dried. 


7. M. elegantulum, 7/w. Enum. 112 (1859). 

C. P. 2684. 

PIS Being sis 55: 

Branchlets very slender, elongated, quadrangular; 1. on 
very short petioles, 2-3in., ovate-lanceolate, acute at base, 
with a long caudate acute apex, $—-?in. long, thin, shining 
above, the veins obscure ; fl. very few or solitary, small, sessile, 
at end of filiform axillary peduncle; fruit pendulous, some- 
what ovoid, purple. 

Moist low country; very rare. Karawita Kande; Gilimale. Fl. May; 
white (?). 

Endemic. 

I have very scanty specimens of this plant. 


8. M. ellipticum, 7/w. Enum. 112 (1859). 
Cogn lic 1160 ENP 3470: 
EBs ind yiis56: 


A small tree, twigs slender, quadrangular; 1. oval or 
oblong-lanceolate, 24-3 in., slightly rounded at base, suddenly 
caudate, with a narrow obtuse tail 3 in. long, veins incon- 
spicuous, petiole very short ; fl. on short ped., clusters small, 
often in axils of fallen 1., sessile ; cal. nearly truncate, shallow; 
disk-rays very obscure ; fruit not seen. 


Moist low country; very rare. Singhe Raja Forest only. FI. Sept.; 
white. 

Endemic. 

I have seen none but the C. P. specimens. C. P. 1557 referred to 
this by Clarke, and Cogniaux I have not seen. Thwaites considered it 
the same as C. P. 1561 (see WZ. umbellatum). 


9. M.macrophyllum, 7iw. Enum. 111 (i859). 
Cosi, Ib CG Live, (Cs ik, Delos. 
PBs Inds 5505 


Memecyton.] Melastomacee. 215 


A small tree, branches thickened at nodes, twigs stout, 
cylindrical ; 1. very large, 6-12 in., narrowly lanceolate-oblong, 
very nearly sessile, more or less cordate or rounded at base, 
acuminate, acute, veins indistinct; fl. on very short ped. in 
dense axillary sessile clusters on the old wood; disk not 
rayed ; fruit rather over } in., crowned with wide but shallow 
cal.-rim. 

Moist low country; rather rare. Hewesse; Gangolla. Fl. white or 
very pale blue. 


Endemic. 
Very like JZ. Wightzz, but the leaves much larger. 


10. M.ovoideum, 7iw. Enum. 112 (1859). 

GP. 2621. 

Fl. B. Ind. ii. 556 (in part). 

A small tree with grey bark, twigs thick, cylindrical ; 1. 
large, 24-4 in., nearly sessile, oval-rotundate, often retuse at 
apex, margin often revolute, very coriaceous, pale beneath, 
veins inconspicuous; fl. rather large, on very short ped., 
fascicles sessile, crowded, axillary, on old wood; disk without 
rays; fruit 4-3 in. by 2 wide, ovoid, purple. 

Montane zone; very rare. Ambagamuwa district; Dimbula. FI. 
Feb., March. 

Endemic. 

Clarke in Fl. B. Ind. appears to combine the very different AZ. #acro- 
carpum with this, in which he is followed by Cogniaux; but the former 
quotes for ovozdewm C. P. 2622 (which is Lugenia cordifolia), and says 
also that it is JZ. revolutum, so that the whole is of doubtful meaning. 


11. M. revolutum, 7/w. Enum. 111 (1859). 

Gera, tb c.1147.. C. P. 29075. 

FI. B. Ind. ii. 560. 

A small tree with thick branches, twigs stout, cylindrical ; 
|. 2-3 in., nearly sessile, rotundate, subcordate at base, very 
obtuse, much revolute at sides and apex, coriaceous, somewhat 
bullate, lat.-veins few, coarse, rather conspicuous beneath ; 
fl. small, ped. twice as long as cal., in rather large peduncu- 
late panicles from axils of fallen 1.; cal.-limb expanded, segm. 
broad, acute; disk with faint rays; fruit about 3in., clobose, 
purplish-black. 

Montane zone, apparently very rare. Palagala, near Ramboda, at 
about 5000 ft., is the locality of the C. P. specimens. Fl. Sept.; white. 

Endemic. 

C. P. 751, from Ambagamuwa district, appears to be also this species, 
which has, doubtless, been overlooked from its foliage closely resembling 
several other montane trees. 


12. M. orbiculare, 7/w. Enum. 112 (1859). 
Cogn. lc, 1163. C. P. 3441. 
Fl. B. Ind. ii, 112. 


216 Melastomacee. [Wemecylon. 


A small tree, twigs cylindrical; 1. sessile or nearly so, 
14-2 in., orbicular, often rather wider than long, subcordate 
at base, retuse, occasionally very obscurely acuminate, paler 
beneath, veins inconspicuous; fl. in dense axillary fascicles, 
ped. equalling cal., peduncle very short or none; disk not 
rayed ; fruit under }in. 

Moist low country; very rare. Only found on Kalabowitiya Kande, 


Hiniduma Pattu. Fl. April; bright blue. 
Endemic. 


13. IM. procerum, 7iw. Enum. 415 (1864). 
Goons lice iis t= |G. P3347. 
Fl. B. Ind. ii. 556 


A moderate-sized tree, 30-40 ft., twigs cylindrical ; 1. very 
large, 6-10 in., ovate or lanceolate-ovate, rounded at base, 
gradually acuminate, pale beneath, petiole 2in., stout, chan- 
nelled above; fl. (not seen) nearly sessile, with large bracts, 
cymes sessile; disk (in fruit) faintly rayed; fruit large, 2in., 
crowned with large erect cal.-limb. 


Moist low country; rare. Kalutara; Leenakande, Pasdun Korale. 
Endemic. 


14. ME. cuneatum, 7/w. Enum. 112 (1859). 

Cogn 116250 Cy Patz: 

Fl. B. Ind. 11. 564 (JZ. edule, var. 7). 

A small tree, much branched, branchlets thickened at 
nodes, twigs cylindrical or sub-quadrangular; 1. 14-2 in., 
sessile or very shortly stalked, from obovate-oblong to almost 
rotundate, tapering to cuneate base, rounded at apex, coriace- 
ous, veins invisible; fl. small, ped. shorter than cal., cymes 
sessile in axils on old wood, buds acutely pointed ; disk with- 
out rays ; fruit tin. 

Lower montane zone; rare. Maturata; Mattettugama. FI. August; 
purplish-white. 

Endemic. 


15. WZ. umbellatum, Survm.f. Fl. /nd. 87 (1768). Kora-kaha, 
S. Kaya, Kurrekaya, Pandikaya, 7. 

Herm. Mus. 4o. Burm. Thes. 76. Samara leta, L. Mant. ii.; Moon 
Cat. 11. MW. ramiflorum, Lam. Enc. Meth. iv. 88. Thw. Enum, 111. 
C. P. 764, 1561, 1562. 

F. B. Ind. i. 563 (JZ. edule, vars. 1 and2?). Burm. Thes.t. 31. Wight, 
Ill. t. 93 (AZ. ténctorium). 


A large bush with many branches, twigs cylindrical; 1. 
14-24 in., oval or ovate-oval or oblong-oval, acute (rarely 


rounded) at base. often slightly acuminate, obtuse at apex, 
dull not shining, lat. veins almost invisible (in fresh speci- 


Memecylon.] Melastomacee. 217 


mens), the intramarginal ones joining the midrib at base and 
giving a slightly 3-nerved appearance, petiole under +}in.; f1. 
small, numerous, on slender ped. as long as or longer than 
cal.. cymes umbellate, sessile or on a very short peduncle, 
copious, closely placed in axils of fallen 1. on the old wood, 
buds pyriform-globular, blunt ; cal. rather shallowly campanu- 
late, puberulous, segm. broadly triangular, acute, a little shorter 
than tube; disk quite without or with very faint rays; pet. 
small, distant, reflexed; fruit 4 in., with inconspicuous cal.-limb. 


Var. 8, Thwaitesii, Clarke, /. c. (MW. edule, var. 10). M. urceolatum, 
Coens) c.2156: ~C. P. 387. 


L. 3-4in., acuminate, obtuse; cal. large, glabrous, tube 
deeply campanulate, segm. very acute, not half as long as 
tube. 


Var. y, rubro-czruleum, Clarke, /. c. (WV. edule, var. 6). M.rubro- 
ceruleum, Thw. Enum. 415. C. P. 3809. 


A small tree of 20ft.; 1. tapering to base, lat. veins quite 
invisible ; peduncle about +in.; cal.-limb longer, nearly trun- 
cate. 


Low country in both moist and dry regions; common. Var. 6 only 
at Haldummulla. Fl. March-September; brilliant blue (cal. red in 
var. 7). 

Also in S. India. 

A very attractive shrub when in flower, from the quantity of brilliant 
bosses of flowers along the whole length of the branches. Var. 6 looks 
distinct, but I suspect the enlarged cal. is really due to hypertrophy from 
the attacks of some insect. 

There is no specimen or drawing in Hermann’s Herb., so that Linnzeus 
had only Burman’s figure to go upon. This is excellent; but in the 
separate flower only 4 stamens are shown. Hence Linnzus’s genus, 
Samara, based wholly on that figure. Burm. f.’s name was published 
without description; if it be set aside on that account, JZ. ramzflorum, 
Lam. (1796) takes its place. I donot know what relation JZ. edule, Roxb. 
(1795), bears to it, though Roxburgh quotes Burman’s figure, and both 
Clarke and Cogniaux make it the same (see under No. 26). 

Wood whitish, smooth, rather light. The leaves and young shoots 
afford a yellow dye, chiefly used for colouring mats. 


16. M. Clarkeanum, Cogn. Mon. Melast. 1141 (1891). 
M. jambosoides, Thw. Enum, 112 (non Wight) part. C. P. 2468. 
Fl. B. Ind. ii. 560 (7, Heyneanum, var. latifolia). 


A small tree, twigs elongated, cylindrical ; 1. large, 4-7 in., 
long-lanceolate, much tapering at base, acuminate, acute at 
apex, thin, veins fine, inconspicuous, petiole very short or 
none; fl. sessile or nearly so, cymes sessile, small, buds 
pointed; cal. shallow, truncate, disk not rayed, fruit 4 in., 
yellow. 


218 Melastomacee. [Memecylon. 


Moist country; rare. Deltota; Ambagamuwa; Maskeliya. FI. 
Sept.; white. 

Endemic. 

Cogniaux keeps 17. Heyneanum, Benth. (M. jambosoides, Wight, Ic. t. 
277), of S. India, as a distinct species from our Ceylon one. 


17. M. discolor, Cogn. Mon. Melast. 1149 (1891). 

M. jambosoides, Thw. Enum. 112 (non Wight) part. C. P. 1555. 

Fl. B. Ind. ii. 560 (AZ. Heyneanum, var. angustifolia). 

A small tree, branchlets elongated, thickened at nodes, 
twigs cylindrical; 1. large, 5—7 in., narrowly linear-lanceolate, 
tapering to base, very long-acuminate, acute at apex, thick, 
midrib very prominent beneath, lat. veins almost invisible 
(when dried, upper surface rich brown, under surface light 
golden greenish-yellow), petiole very short, stout ; fl. on very 
short ped., cymes sessile, buds pointed; cal. nearly truncate ; 
pet. reflexed ; fruit nearly 4 in., yellow. 

Moist low country; very rare. Ambagamuwa; Palabadala, Kuru- 
wita Korale. Fl. Sept.; white with pink calyx. 

Also in S. India. 

I have seen only leaves of this, but there are good drawings of the 
flowers and fruit. It is, though very different in appearance when ary, 


probably not distinct from JZ. Clarkeanum, but I have seen no fresh 
specimens. 


18. IM. fuscescens. 7hw. Enum. 111 (1859). 

Coon, esii40) 2@ P2387, 

PBS indie 262: 

A bush, branchlets slender, twigs cylindrical or sub- 
quadrangular; |. 2i-2#in., nearly sessile, ovate, acute at base, 
suddenly caudate-acuminate with a tail 4—# in. long, sub- 
acute at apex, membranous, shining above, veins inconspicu- 
ous; fl. few, ped. short, slender, with two bractlets at summit, 
cymes sessile, very small; cal.-segm, acute, disk not rayed ; 
fruit large, over 2 in. 

Moist low country; rare. Kukul Korale, abundant; Lihinigalla, 
Pasdun Korale. Fl. Aug., Sept. 

Endemic. 

Clarke and Cogniaux give C. P. 2926 for this, which I have not seen. 


19. M.rostratum, 7iw. Enum. 111 (1859). KMurétiya, Hin- 
kurétiya, 5. 

Cognac mi6oN ne Ps 1560; 

HIN B: Inds i. 262: 


A small tree, branchlets very numerous, ascending, twigs 
compressed or sub-quadrangular, very slender; I. small, 
(4-23 in., lanceolate or oval-lanceolate, tapering to base, cau- 
date-acuminate, with a tail in. long, obtuse at apex, veins 


faintly marked, petiole 3-{in.; fl. very small, ped. at least 3 


Memecylon.] Melastomacee. 219 


times as long as cal., slender, umbels on a short stout peduncle 
in axils of both past and present I|., buds pointed; cal. shallow, 
nearly truncate, disk distinctly rayed ; fruit small, ¢ in., glo- 
bose. 

Lower montane zone 3000-4000 ft.; rather common. Hantane; 
Deltota; Medamahanuwara. Fl. May; very pale blue. 

Endemic. 

A fairly distinct species by its very small flowers in simple stalked 
umbels. Wood yeliow, hard, smooth. I have it from the Pasdun Korale 
under the name of ‘ Gal-pinibaru.’ 


20. M. angustifolium, Wighi, ///. i. 215 (1840). 

Cogn. lc. 1157. Thw. Enum. 111. C. P. 1556, 3346. 

Fl. B. Ind. ii. 262. Wight, Ic. t. 276. 

A small shrub, 6-8 ft., with many erect slender branches, 
twigs sub-quadrangular; |. 2-3 in., linear or lanceolate-linear, 
almost sessile, much tapering to base, caudate-acuminate or 
rounded at apex, stiff, shining above, paler and dull beneath, 
midrib depressed on upper surface, lat. veins invisible; fl. on 
slender ped., 2 or 3 times as long as cal., umbellate or pani- 
culate, peduncles under 4in., disk without rays; fruit small, 
about + in., black-purple. 

Low country in the beds of streams and banks of rivers up to 2000 ft.; 
rather common. Sigiri (Gardner); Uma-oya; Ambagamuwa; Sabara- 
gamuwa; Labugama; Knuckles (Ferguson). Fl. Feb.-July; bright 
purplish-blue. 

Also in Southern India. 

C. P. 1556 (from Uma-oya) has obtuse non-acuminate |. and consider- 
ably larger fl. in a larger compound pedunculate inflorescence. 


21. M. phyllanthifolium, 7hw. ex Trianain Trans. Linn. Soc. 
XXVili. 159 (name only) (1871). 

Gorn ic: 1165. C. B. 3001. 

Fl. B. Ind. ii. 563. 

An erect stiff shrub, with numerous erect nodular branches, 
twigs cylindrical; |. very numerous, closely placed, small, under 
4 in., nearly sessile, erect, rigid, linear-oblong, tapering to base, 
rounded at apex, margin somewhat recurved, lat. veins in- 
visible; fl. nearly sessile, very small, 2 or 3 together at summit 
of very short peduncle, cal.-limb spreading, segm. acute; fruit 
small, scarcely 4in., crowned with conspicuous cal.-limb. 

Upper montane zone; very rare. Bogawantalawa at about 5000 ft.; 
first collected in 1866. Fl. March. 

Endemic. 


Quite unlike the rest of the genus, with the habit of a shrubby 
Phyllanthus. 


22. M. rhinophyllum, 7/w. “num. 110 (1859). 
Cogn. |. c. 1166. C. P. 2928. 
Fl. B. Ind. ii. 562. 


220 Melastomacee. [Memecylon. 


A small tree, twigs slender, quadrangular; 1. sessile or 
very nearly, 3-41in., ovate-oval, very acute at base, long-acu- 
minate, obtuse at apex, dull not shining, paler beneath, veins 
invisible; fl. numerous, ped. twice as long as cal., slender, in 
rather large umbellate-corymbose cymes on short flattened 
peduncles; cal.-segm. broad, shallow; disk with winged rays ; 
fruit ¢ in., purplish-black. 

Moist low country to 3000 ft.; rather common. Ratnapura; Alagala; 
Hantane. Fl. April; pale violet-blue. 

Endemic. 

Clarke, followed by Cogniaux, quotes for this C. P. 2572 in error; 
that number is an Overonia. 


23. M. leucanthum, 7/w. Exum. 110. 

M. edule, var., Cogn. Mon. 1156. C. P. 1559. 

Fl. B. Ind. ii. 564 (47. edule, var. 6). 

A small tree, twigs obscurely quadrangular ; |. 14-24 in., 
broadly oval to rotundate, acute at base, shortly acuminate, 
obtuse at apex, lat. veins rather conspicuous beneath, petiole 
very short; ped. as long as or longer than cal., divaricate, 
cymes racemose-paniculate, lax; cal.-tube campanulate, segm. 
short, erect, obtuse; pet. clawed, somewhat lacerate, strongly 
reflexed, disk-rays distinctly winged; fruit small, under } in., 
crowned with large cal.-limb. 

Lower montane zone; rather rare. Hantane; Watagoda; Dimbula. 
Fl. April; pure white. 

Endemic. 


24. MZ. macrocarpum, 7/w. E7um. 110 (1859). 

M. ovotdeum (non. Thw.), Cogn. 1. c. 1147. C. P. 2798. 

Fl. B. Ind. ii. 556 (with JZ. ovotdeum). 

A small tree with straight erect trunk, branches thickened 
at nodes, twigs stout, cylindrical; |. rather large, 3-4 in., ro- 
tundate, acute at base, obtuse or rounded at apex, very thick 
and coriaceous, margin slightly revolute, pale beneath, lat. 
veins quite invisible, petiole very short and thick; fl. (not 
seen) from large axillary bosses on the old wood, ped. short ; 
fruit large, #in. or rather more, globose, suddenly contracted 
at summit into a short neck, crowned by truncate cal.-limb., 
disk conspicuously winged. 

Moist region; very rare. I have only seen the C. P. specimens from 
Ambagamuwa, which are in fruit. 


Endemic. A 
A very distinct species, curiously confused with J/. ovozdeum. 


25. M. levigatum, B87. Mus. Bot. i. 358 (1851). 
M. sylvaticum, Thw. Enum. 110. C. P. 1558, 1567, 3302. 
IIE 183, Teal, is Ou 


Memecylon.] Melastomacee. 221 


A small tree, twigs cylindrical; 1. 2}-5 in. sub-sessile, 
broadly oval or ovate-oval, tapering to base, rather abruptly 
acuminate, obtuse at apex, shining on both sides, pale be- 
neath, coriaceous, veins inconspicuous; fl. large (for genus), 
on very short thick ped. or nearly sessile, cymes small, crowded 
in axils of fallen 1, peduncles usually very short, buds 
large, pointed; cal.-limb cup-shaped, truncate, mealy-pubes- 
cent outside, segm. obscure, disk-rays prominently winged ; 
pet. broader than long, very obtuse, recurved; fruit about 4 in., 
often somewhat ovoid, crowned with truncate cal.-limb, dark 
purple. 

Forests from 2000-4000 ft. or higher; common. FI. Jan.-April; deep 
violet-blue, cal. white or pink. 

Also in Malay Peninsula and Islands. 

Known from JZ. grande by its larger, nearly sessile flowers. Cogniaux 
(1. c. 1150) keeps up Thwaites’ AZ. sylvaticum as distinct from JZ. leviga- 


tum. At the higher elevations the leaves are smaller (13-23 in.) and 
more coriaceous. 


26. MZ. grande, fefz. Obs. Bot. iv. 26 (1786). Dodan-wenna, 
Dedi-kaha, S. 

M., edule, var. 8, Thw. Enum. 110. Cogn. 1.c.1153. C. P. 3442. 

Fl. B. Ind. 11. 557. 


A large shrub or small tree with smooth grey bark, twigs 
cylindrical; |. large, 4-6in., broadly oval or ovate-oval, acute 
at base, caudate-acuminate, often twisted, obtuse at apex, 
thick, dark green above, veins inconspicuous, petiole 4-4 in.; 
ped. about as long as cal.; cymes large, often 2 in. lax, 
paniculate, spreading, from axils of present and fallen 1. 
peduncles 4~2 in., flattened, buds globose; cal.-limb truncate, 
segm. obscure; disk-rays winged; fruit 2 in., crowned with 
small erect cal.-limb, dark purple, rather juicy. 


Var. 8, ovatum, Clarke (M. edule, var. 4). M. edule, Roxb., Moon 
Cat. 32; Thw. Enum. I1o (vars. aand y). C. P. 1563, 1566. 


L. smaller, 24-4 in., often somewhat oblong, less acute at 
base and less acuminate; ped. a little longer, cymes smaller, 
umbellate-corymbose, disk-rays usually winged (but some- 
times not or even faintly marked), fruit smaller, 4} in. 


Moist region in the low country, especially near the coast; rather 
common. Fl. Sept.-April; pale purplish-blue; var. 8, darker. 

Also in India and Malaya. 

I follow Thwaites in combining these. The type of Retzius’ species 
(from Koenig) in the British Museum is identical with C. P. 3442. As to 
var. B, it may be Roxburgh’s . edule; but his figure (Cor. Pl. t. 82) is 
unsatisfactory, and the description insufficient, and the name appears 
to be now very vaguely applied. In Fl. B. Ind., for instance, all the 
‘enormous mass’ of specimens in Kew Herb. that ‘appear difficult to 


222 Lythracee. 


separate ’ from it (including many fairly distinct species) are placed under 
the name. On the whole, it seems safer not to employ it for any 
Ceylon species. 7. Royenzz, Bl. Mus. Bot. i. 360, from Ceylon, probably 
comes here. 
Wood hard, strong, and rather heavy. The fruit is eaten when cooked. 
The rays of the disk are sometimes quite inconspicuous in this species. 


27. M. capitellatum, Z. S#. Pl. 349 (1753). Wéli-kaha, 
Dodan-kaha, S. Kattikaya, Venkalikaya, Pavaddai- 
kaya, 7. [PLATE XLI.) 

Herm. Mus. 4. Burm. Thes. 76 Fl. Zeyl. n. 136. Moon Cat. 32. 
Thw. Enum. 110. Cogn. 1.c. 1142. C. P. 1564. 

Fl. B. Ind. ii. 564 (WZ. edule, vars. 3 and 5). Burm. Thes. t. 30. Bedd. 
Fl]. Sylv. t. 206, fig. A. 

A shrub with brown bark, twigs cylindrical; 1. very shortly 
stalked, oblong or slightly rhomboid-oval, acute at base, ob- 
tuse at apex, sometimes slightly acuminate, thick, shining on © 
both sides, bright light green, paler beneath, lat. veins incon- 
spicuous, the intramarginal one joining the midrib at base 
and making it distinctly 3-nerved (very obvious when dry); 
fil. rather large, quite sessile, few together in small heads at 
ends of stiff peduncles 4-14 in. long, from axils of present 
and fallen |.; cal. shallow, seem. conspicuous, fleshy, erect, 
bluntly pointed, disk-rays strongly winged; fruit 2in. or a 
little more, crowned with large cal.-segm., black purple when 
quite ripe. 

Low country up to 1000 ft. in both moist and dry regions; common, 
especially in the northern forests. F]. May, June; violet-blue. 

Endemic. (?) 

This is the original type of the genus, the only species described by 
Linneeus. JZ. ¢éuctorium, Koen. in Willd. Sp. Pl. 11. 347, is referred, as a 
variety, to this by Blume, who has seen Kecenig’s specimen which was 
collected at Galle. Clarke quotes for this Saszara /eta, Moon, which 
belongs to MZ. umbellatum (No. 15). 

One of our best-marked species, abundantly distinct from J. edule, 
Roxb. (as a variety of which it is considered in FI. B. Ind.), and easily 
recognised by the foliage alone. Wood hard, strong. The leaves turn 
orange-yellow when dried; they afford a yellow dye. 


LIV.—LYTHRACE/. 


TREES, shrubs, or herbs, |. opp., entire, no stip., fl. regular, 
bisexual, variously arranged ; cal. persistent, free or slightly 
adnate to base of ov. (in Azzzandra completely adnate), 
segm. 3-6; pet. 3-6 or 0; stam. double the number (rarely 
equal to number) of pet., or indef., inserted on cal.-tube; ov. 


Ammannia.| Lythracee. 228 


superior (in A1zzandra inferior), 1-6-celled, with numerous (1 in 
Axinandra) ovules in each cell, style simple; fruit a capsule, 
often included in cal.-tube, variously dehiscent, rarely inde- 
hiscent; seeds numerous, sometimes winged, embryo straight, 
rarely bent, no endosperm. 


HAeEDS) * . 5 : : : : : . I. AMMANNIA. 
Shrubs or trees. 
Stam. 12. 
Capsule irregularly dehiscent . 2. WOODFORDIA. 
Capsule eras Py. a oe 3. PEMPHIS. 
Stam. 8 A 4. LAWSONIA. 
Stam. indef. 
Fruit a 6-valved capsule . : : : 5. LAGERSTREMIA. 
Fruit fleshy, indehiscent . 6. SONNERATIA. 
Stam. 10 (Azomalous Genus: ov. inferior) . 7. AXINANDRA. 


Most of our 15 species are low-country plants; indeed, Woodfordia is 
the only hill species. The Amsmannias are paddy-field weeds. Lawsonia 
is confined to the dry region, and Lagerstremia and the anomalous 
Axinandra to the moist. The Sozneratias inhabit mangrove-swamps, 
and Pemphis is a seashore plant. 


1. AMMANNIA, /. 


Annual herbs, |. opp. (rarely alt.) or whorled, fl. small, 
axillary; cal. more or less deeply campanulate, segm. 3-5, 
often with intermediate teeth ; pet. 3-5 or 0, inserted between 
cal.-segm.; stam. 2, 4, 5, or 8; ov. superior, 1—5-celled, the - 
septa often becoming absorbed, ovules very numerous on 
axile placentas; capsule membranous, enclosed in’ (though 
free from) persistent cal., dehiscing by valves or irregularly 
circumscissile, seeds minute, very numerous.—Sp. 30; 18 in 
Fl. B. Ind. 

F], solitary in axils alah: 


Stam. 4. 1. A. PEPLOIDES. 
Stam. 3. 2. A. ROTALA. 
Stam. 5. : 3. A. PENTANDRA. 
F]. in axillary cymes or clusters. 
Stam. 4. 
L. oblong-linear, obtuse 4. A. BACCIFERA, 
L. oval, cordate at base ; ; 5. A, CORDATA. 
it spathulate- oval, ig to base . 6. A. LANCEOLATA, 
Stam. 8. ; ? 7. A. OCTANDRA. 


A. peploides, Spreng. Syst. Veg. i. 444 (1825). 
Mipeletin indica, DC., Thw. Enum. 122. C. P. 1546. 
FI. B, Ind. ii. 566. Wight, eet 257, 
Stems 6-12 in., usually decumbent at base and rooting at 
the lower nodes, with numerous ascending quadrangular 


224 IL yth vacee. [Ammannia. 


branches ; |. opp., on main stem and branches 4—? in. but on 
floriferous branchlets very much smaller and bract-like, nearly 
sessile, rounded at apex, spathulate-oval, narrowed at base, 
glabrous, veins prominent beneath; fl. very small, solitary 
in axils of the small 1., which are more or less approximated 
so that the branchlets look like spikes, bracts 2, linear, 
scarious; cal.-tube tubular, campanulate, glabrous, segm. 4, 
narrowly triangular, very acute, intermediate teeth minute ; 
pet. 4, very small or 0; stam. 4; capsule ovoid, somewhat 
compressed, opening by 2 valves from the summit; seeds 
very minute, oblong, chestnut brown. 

Low country up to 3000 ft., in paddy-fields, tanks, and wet places; 
very common. FI. all the year. 


Throughout India and Malaya, China, Persia. 
Has somewhat the habit and appearance of an Amarantacea. 


2. A. Rotala, /. Muell. Fragm. iii. 108 (1863). 

Rotala verticillaris, L., Thw. Enum. 122. C. P. 1547. 

Fl. B. Ind. ii. 567. Wight, Ic. t. 260, A (A. verticzllaris). 

Stems 3-6in., often procumbent and rooting at very base, 
then erect, slightly branched, glabrous, striate; 1. in whorls of 
4-8, about }in., narrowly linear, truncate or bifid at apex, 
1-nerved ; fl. sessile, solitary, whorled; cal.-tube tubular, very 
thin, segm. 3, short, broadly triangular, acute; pet. 3 or 0; 
stam. 3; ov. 3-celled; capsule ovoid, completely 3-valved. 

Moist low country; rather common. FI. all the year. 

Also in India, Australia, Trop. Africa. 


Thwaites says this is ‘very abundant.’ I have not found it so in my 
experience, but it is readily overlooked. 


3. A. pentandra, fox. Fl. Ind. i. 425 (1832). ; 
A. debtlis, Ait. Moon Cat. 11. Thw. Enum. 122. C. P. 1548. 
Fl. B. Ind. il. 568. Wight, Ic. t. 260, A (Rotala Roxburghiana). 


Stem stout, quadrangular, rooting at the base, 6-12 in.; with 
numerous horizontal branches; |. opp., on the stem $4 in., but 
on the branches much smaller and bract-like, sessile, oblong, 
subacute; fl. sessile, solitary, bractlets subulate; cal. shortly 
campanulate, segm. 5, triangular, very acute, intermediate 
teeth long, subulate; pet. 5; stam. 5; capsule depressed- 
globular, completely 3-valved; seeds hemispherical. 


Low country in wet places; common. FI. all the year. 
Throughout the Tropics of the Old World. 


4. A. baccifera, Z. SZ. P/. 120 (1753). 
Moon Cat, 11. A. zndica, DC., Thw. Enum. 121. C. P. 1542 
Fl. B. Ind. ii. 569. Blume, Mus. Bot. ii. t. 46, A (A. ¢zdica). 


Ammannia.] Lythrac Ce. 225 


Stems 8—24 in., erect, quadrangular, glabrous, with nu- 
merous horizontal or ascending branches from the very base, 
eradually becoming shorter upwards so as to give the plant 
a pyramidal form; 1. opp. (rarely alternate), on main stem 
I-I? in. but smaller on branches, sessile, oblong-linear, much 
tapering to base, obtuse at apex, flaccid; fl. shortly pedicellate, 
in small loose cymes, forming whorls in the axils; cal. shortly 
campanulate, segm. 4, broad, acute, intermediate teeth slightly 
marked; pet.0; stam. 4 (or 2); capsule exceeding cal., de- 
pressed-globose, red, irregularly circumscissile ; seeds plane- 
convex, the plane face excavated. 

Moist sandy ground in the low country, especially in the dry region 


and near the coast; common. Fl. Jan.—April. 
Throughout the Tropics of the Old World. 


5. A. cordata, Wight and Arn. Prod. 304 (1834). 

A. salicifolia, var. B, Thw. Enum. 121. C. P. 3296. 

Fl. B. Ind. ii. 570. 

Stem 6-12 in., quadrangular, glabrous, branched from the 
base; 1. 2-14 in., sessile, ovate-oval, cordate and somewhat 
amplexicaul at base, with a narrow cartilaginous margin; fl. 
very shortly pedicellate, in small opp. clusters; cal.-tube cam- 
panulate, segm. 4, large, broadly triangular, acute, interme- 
diate teeth conspicuous; pet. and stam. 4; capsule depressed- 
globose, almost covered by the inflexed cal.-segm. 


Dry country; rare. Batticaloa; Haragama. Fl. March—June. 
Also in India. 


6. A. lanceolata, Heyne in Wall. Cat. 2106 (1828). 
A. salicifolia, Thw. Enum. 121 (non Monti). C. P. 2796. 

Fl. B. Ind. ii. 570. 

Stem 3-9 in., slightly branched, quadrangular; |. 3-1 in., 
sessile, spathulate-oval, tapering to base; pet. 0; otherwise as 
in A. cordata. 

Low country; rather common. Kalutara; Uma-oya; Batticaloa. 
Fl. March, April. 

Also in Southern India. 

This is described in Fl. B. Ind. as having petals considerably exceed- 

ing the cal. Our specimens seem to have none. 


7. A. octandra, L. /. Supp. Pl. 127 (1781). 

Moon Cat. 11. Thw. Enum. 121. C. P. 2797. 

Fl. B. Ind. ii. 571. Roxb. Cor. Pl. t. 133. 

Stem 1-2 ft., erect, stiff, quadrangular, with numerous 
and ascending branches; 1. on main stem 14-24 in. but 
smaller on the branches, sessile, linear-lanceolate, auriculate 
at base, much tapering to acute apex; fl. rather large, on very 

PART II. Q 


226 Lythracee. [ Woodfordia. 


short ped., in threes at end of short scabrous peduncles; cal. 
long-campanulate, quadrangular, the angles winged, segm. 4, 
short; pet. 4, rotundate, spreading; stam. 8; capsule eranely 
ovoid, enclosed in persistent 4-winged cal. 

Dry region; rather common. Jaffna (Moon); Dambulla; Matale; 
near Kurunegala. Fl. March, April; bright pink. 

Also in S. India and Malaya. 

Much the most showy of our species. 


2. WOODFORDIA,* Saiisd. 


A spreading shrub, 1. opp., fl. in copious axillary panicu- 
late cymes; cal.-tube widely tubular, oblique at mouth, segm. 
6; pet. very small, 6, inserted between cal.-segm.; stam. 12, 
inserted at base of cal.-tube; ov. superior, enclosed in but free 
from the campanulate base of cal.-tube, 2-celled, with very 
numerous ovules; fruit a membranous capsule included in 
cal.-tube, with very numerous seeds.—Monotypic. 


W. floribunda, Sa/isb. Parad. Lond. t. 42 (1806). Malitta, S. 
Grislea tomentosa, Willd., Moon Cat. 31. Thw. Enum. 122. C. P. 1552. 
BEB wlnd si. 5723) skoxbs Cor) Plots sit bot. Macat 19a: 

A straggling shrub, with many long arching branches, 
bark cinnamon-brown, shredding off in fibres, shoots covered 
with fine white pubescence, cylindrical; 1. sessile, 3-4 in., 
narrowly ovate-lanceolate, cordate at base, tapering to acute 
apex, entire, finely velvety on both sides, veins pellucid, pro- 
minent beneath, the lat. ones uniting within the margin, paler 
or whitish beneath, dotted with very minute orange glands; 
fl. on pubescent ped., arranged in numerous short, divaricate, 
cymose panicles from the axils of fallen 1. on the old wood, 
and rarely from those of present |.; cal. over } in., with a small 
campanulate base, and a long, slightly curved, somewhat in- 
flated funnel-shaped tube, pubescent, red, segm. very short, 
triangular, acute; pet. very small, about as long as cal.-segm.; 
stam. declinate, much exserted, persistent; capsule enclosed 
in persistent cal.-tube which becomes vertically split, about 
2 in., pericarp thin, membranous, irregularly dehiscent; seeds 
very numerous, oblong-wedge-shaped, brown, smooth. 

Open sunny places in the lower montane zone; rare. Maturata; Uva 
patanas locally abundant, as on the Badulla road about Wilson’s bungalow, 


Bandarawella, &c., but. scarcely in the Central Prov. mountains. FI. 
March, April; bright brick-red. 


* In honour of E. J, A. Woodford, Esq., a great cultivator of rare 
plants in London in the early part of this century. 


Lawsonia.| ie yt hracee. DDG 


Also in India, China, Trop. Africa, and Madagascar. 
The stamens are trimorphic as in Lythrume. 


Nes@a trifiora, Kunth., is C. P. 1541, and recorded in Thw. Enum. 417. 
The specimens were collected by Walker, and were no doubt introduced. 
The plant is a native of Mauritius. 


3. PEMPHIS, forsz. 


A maritime shrub, |. opp., fleshy; fl. axillary, solitary; 
cal.-tube campanulate, segm. 6, short, with intermediate teeth; 
pet. 6, inserted between cal.-segm., spreading; stam. 12, in- 
serted near the base of cal.-tube; ov. superior, 3-celled with 
numerous ovules, stigma large, capitate; fruit a capsule, en- 
closed for 2 of its length in cal.-tube, pericarp thin in enclosed 
portion, thicker in projecting blunt summit, which separates 
as a cap; seeds large, numerous, cuneate, angular, smooth.— 
Monotypic. 


P. acidula, /orst. Gen. 67 (1776). 

meen Pemphis, L. f. Suppl. Pl. 249. Moon Cat. 37. Thw. Enum. 
Pome P 1553. 

FI. B. Indi 573. Wight, Ic. t. 1996 (Waclellandia Griffithiana). 

A shrub, 3-5 ft. high, with very numerous ascending vir- 
gate branches densely clothed with fine white pubescence, 
especially round the nodes; |. small, numerous, sessile, often 
very closely placed, 4-14 in., readily disarticulating, lanceolate- 
oblong, acute at base, obtuse at apex, densely silvery-pubes- 
cent on both sides, fleshy; fl. on pubescent ped. as long as 
cal., solitary, axillary; cal. 12-ribbed, densely pubescent, segm. 
very short, triangular, acute, intermediate teeth similar but 
smaller; pet. rather large, as long as cal., oblong, undulate; 
capsule } in. long, tipped by persistent style. 

Seacoast; rather common. Jaffna; Trincomalie; Galle; Dondra 
Head; Kalpitiya; Chilaw. Fl. July, August; pink. 

Tropical shores of the Old World generally. 

First collected by Koenig. 


4. LAWSONTIA,* 7. 


A shrub, 1. opp. entire; fl. small,in large terminal panicles; 
cal.-tube very short, segm. 4; pet. 4, inserted at summit of 
cal.-tube; stam. 8, inserted on cal.-tube in pairs; ov. superior, 
4-celled, ovules numerous, style simple; fruit a small capsule, 
with thin pericarp, dehiscing irregularly; seeds numerous, 
closely packed, smooth, angular.—Monotypic. 


* Commemorates Dr. Isaac Lawson, a Scotch friend of Linnzeus. 


228 Lythracee. [Lagerstramia. 


L. alba, Lam. Enc. Meth. iii. 106 (1789). Marutonti, 7. 

Burm. Thes. 142. FI. Zeyl. nn. 134 and 135. JL. sfzmosa and L. 
ners, le. op. L. 349; Moon Cat. 31. Thw. Enum. 1225) Geraaassn 

Fl. B. Ind ii. 573. Wight, Ill. t. 87. 

A much-branched shrub, the lat. branchlets often ending 
in a sharp spinous point; |. small, $-14 in., very nearly sessile, 
oval or lanceolate, tapering to base, acute or obtuse at apex, 
often apiculate, entire, glabrous; fl. numerous, small, under 
4 in., ped. slender, glabrous, pink, arranged in rather long 
axillary and terminal panicles, the whole forming a large 
pyramidal inflor.; cal. glabrous, tube very short, segm. oval, 
acute; pet. oblong, undulate, spreading or reflexed; stam. 
spreading in pairs; capsule not + in., globose, supported on 
persistent cal., tipped with style. 

Dry and desert regions, especially near the seacoast; rather rare. 
Batticaloa; near Chilaw; Mannar; Jaffna, abundant by the salt lakes. 
In the moist region cultivated only. Fl. Feb., March; pet. cream-yellow, 
sep. pinkish; very sweet-scented. 

ee in Western India, Kabul, and Persia, and much cultivated else- 
where. 

Called ‘Tree-Mignonette’ in cultivation in Colombo. This is the 
‘Henna;’ the leaves are used for staining the finger-nails and teeth, and 
- also for applying to the head when bathing. 


5. LAGERSTREMIA, L. 


arse tree; opps entire; my very larce; inietennaimal 
panicles; cal.-tube campanulate, fleshy; segm. 6, shorter than 
tube, intermediate teeth very small; pet. 6, clawed, inserted 
at brim of cal.-tube; stam. very numerous, in several rows 
inserted near base of cal.-tube, connective dilated; ov. superior, 
6-celled, with numerous ovules, style simple; fruit a woody 
capsule, surrounded at base by persistent, semi-woody cal. 
loculicidally dehiscent into 6 valves; seeds numerous, flat, 
winged at one end.—Sp. 18; 12 in FZ. B. Ind. 


L. Flos-reginze, Retz. Obs. Bot. v. 25 (1789). Muruta, S. 

Herm. Mus. 61. Burm. Thes. 137. Moon Cat. 42. Thw. Enum. 122. 
CPE U5 54. 

Fl. B. Ind. 11. 577. Wight, Ic. t. 413. Bedd. FI. Sylv. t. 29. 

A large tree with wide-spreading branches, bark pale, 
rather smooth, flaking off in irregular pieces; |. large, 6-10 in., 
oval or oblong-lanceolate, rounded at base, subacute, entire 
but somewhat repand at margin, perfectly glabrous on both 
sides, paler beneath, with lat. veins very prominent, petiole 
4-4 in., stout; fl. very large, 2-4 in., on very stout, pubescent 


Sonneratia.] Lythracee. 229 


spreading ped. thickened upwards and articulated below the 
middle, where are 2 small opp. bractlets, panicle large, 14$-2 ft., 
stout, erect, with spreading branches, bracts small, buds pyri- 
form, truncate; cal.-tube with 12 vertical grooves, mealy- 
pubescent, segm. long-triangular, acute, spreading, rather 
shorter than tube; pet. I-1#in., rotundate, leaf-like, with a 
stout petiole-like claw, veiny, much undulate and crumpled, 
spreading, margin slightly erose; fil. under 1 in.; ov. glabrous; 
fruit 2 in., broadly ovoid, about 4 concealed by persistent, 
cup-like cal., very hard and woody, apiculate, smooth, valves 
remaining connected by their bases; seed with wing } in., 
glabrous, pale brown. 

Moist low country up to 2000 ft., especially by rivers and streams ; 
rather common. FI. April-July; deep or light mauve, or bright rose- 
coloured. 

Also in India, Malaya, and China, but often cultivated. 

A magnificent tree when in full flower; it is deciduous and bare of 
leaves for a very short time. Wood light brownish-red, hard, not heavy, 
very useful. 

Though Hermann knew this tree, he preserved no specimen in his 
herbarium ; hence it was left unnamed by Linnzeus among the ‘ Barbare’ 
(FL Zeyl. n. 533). 


6. SONNERATIA,* Z. 7 


Shrubs or trees; |. opp., entire, thick; fl. large, solitary 
or 3, terminal; cal. coriaceous, tube campanulate, slightly 
adnate to base of ov., segm. 4 or 6; pet. 6 or 0; stam. very 
numerous, inserted within rim of cal.-tube, fil. very long; 
ov. almost entirely superior, many-celled, with numerous 
ovules in each cell, style long, stigma capitate; fruit large, 
hard-fleshy, indehiscent, supported on persistent cal., many- 
seeded; seeds small, angular, embryo bent.—Sp. 5; 3 in 
Fl. B. Ind. 


Pet. o. 
Tree; cal.-segm. 4 : ; , ‘ ; . I. 5. APETALA. 
Shrub; cal.-segm.6 . : , , : : 2 e ALBA: 
Pet. 6 : 2 : A ; ‘ j 5 6 2) 3) ScACIDA: 


1. S. apetala, Ham. in Symes, Emb. Ava, iii. 313 (1800), 

Trim. in Journ. Bot. xxvii. 163. 

Fl. B. Ind. ii. 579 (not given for Ceylon). Symes, Emb. Ava, t. 25. 

An erect tree, attaining 40 ft., with slender drooping 
branches; 1. 24-44 in., narrowly oblong-lanceolate, much 


* Commemorates P. Sonnerat, who figures S. ac¢da in his ‘ Voyage a 
la Nouv. Guinée,’ 1776. 


220 Lythracec. [Sonneratea. 


tapering at base into short petiole, obtuse at apex, glabrous, 
rather thick; fl. solitary, terminal, large, articulated on rather 
long ped.; cal.-tube cup-shaped, segm. 4, longer than tube, 
oblong-ovate, acute, deflexed; pet. 0; fruit about # in. wide, 
depressed-globular, tipped with style, smooth, white. 

Mangrove swamps; very rare. Only found at Koddiyar, near Trin- 
comalie, Aug. 1885, in fruit. Fl. June (?). 

Also in Bengal and Burma. 

This possesses upright root-processes like S. aczda. 


2. S. alba, Smith in Rees Cyclop. xxxiii. (1816). 
Trim. in Journ. Bot. xxiii. 171. 
Fl. B. Ind. ii. 580 (not given for Ceylon). Rumph. Herb. Amb. iii. t. 73. 


A bush or small tree, with smooth grey bark, twigs 
cylindrical, marked with prominent leaf-scars; 1. 3-4 in., 
oval-rotundate, slightly tapering at base, rounded at apex, 
very thick and fleshy, petiole very short; fl. large, on very 
short thick ped., buds 6-angled, pointed; cal.-tube with 
6 prominent ridges extending from between the segm. to the 
base, segm. 6, narrowly lanceolate, acute, longer than tube; 
pet.o; style 14-2 in., stigma large; fruit supported on the 
much-enlarged cal. which forms a cup round the base with 
spreading segm., turbinate, about I? in. wide, much depressed, 
flat on top, with the apiculate short style-base in the centre. 

Mangrove swamps; very rare. Only found as yet at Chilaw, where it 
was first collected by Mr. Nevill in 1881. Fl. Nov.; filaments white. 


Also in Malay Peninsula and Islands, Tropical Australia and Africa, 
but not in Peninsular India. 


3. S. acida, LZ. f Supp. Pi. 252 (1781). HKirilla, S. Kinnai, 7. 

IMony, ein, 123, (C, IP, ws. 

Fl. B. Ind. ii. 579. Wight, Ic. t. 340. 

A shrub or small tree, twigs quadrangular; |. nearly 
sessile, 2{-3 in., oblong-oval or roundish, tapering to base, 
obtuse, sometimes apiculate at apex, slightly fleshy; fl. large, 
3 in. long, terminal, solitary, on very short thick ped., buds 
not ribbed, blunt but apiculate at apex; cal.-tube quite with- 
out ribs, segm. 6, lanceolate, acute, longer than tube; pet. 6, 
linear, about I in. rather longer than cal.-segm., persistent; 
stam. about 2 in. erect; style considerably exceeding stam. ; 
fruit broadly ovoid or subglobular, supported on enlarged 
cal. which forms a shallow cup, about 13 in. wide, rounded 
and slightly depressed on top, with the persistent tapering 
style-base forming a point I in. long in the centre. 


Tidal estuaries with the mangroves; rather common. Chilaw; 
Negombo; Panadure; Kalutara. Fl. Sept.; dark rose-red. 


Axinandra.] Lythracee. DOI 


Also in Bengal, Burma, Siam, Java. 

Wood white, soft, light, but fine-grained. 

The curious erect root-branches are figured by Karsten in his ‘ Man- 
grove Vegetation,’ t. 10, f. 139; they were first described by Rumph (Herb. 
Amb. iii. 112). As they attain 18 in. to 3 ft. in height, and 3 in. in 
diameter, and have a soft, firm, even texture, they form a fine substitute 
for cork, and are cut into slices and used for entomologists’ boxes and 
other purposes. Their wood is very much lighter than that of the stem. 
(See Dr. Templeton, of Ceylon, in Trans. Ent. Soc. ili. 302.) 


7. AXINANDRA, 7iw. 


A tree; 1. opp., entire; fl. very small in short racemes ; 
cal.-tube shallowly campanulate, completely adnate to ov., 
limb spreading, segm. 5; pet. 5, connate into a deciduous cap 
(calyptra); stam. 10, inserted in two rows at mouth of cal.- 
tube, fil. very short, dilated, anth. with a very large connective 
much produced behind; ov. completely inferior, 6-celled, with 
I ovule in each cell, style very short ; fruit a woody capsule, 
the lower half fused with enlarged cal.-tube, loculicidally 
dehiscent in the upper part by 2-4 valves; seeds oblong, 
much compressed, with a long wing at upper end.—Sp. 5; 
2 in FZ. B. /nd., but none in Peninsular India. 


This anomalous genus is included by Baillon in MMelastomacee 
(Wemecylez), and he is followed by Cogniaux. It has points of alliance 
also with Rizzophoracee (Legnotidee). 


A. zeylanica, 7hw.in Kew Journ. Bot. vi.66 (1854). Wékiri- 
wara, S. 

Cogn. Mon. Melast. 1113. Thw. Enum. 122. C. P. 2668. 

Fl. B. Ind. ii. 581. Kew Journ. Bot. vi.t. Ic. Bedd. Fl. Sylv. t. 207. 

A tree reaching a large size, but usually about 30 ft., with 
a very straight trunk and numerous short drooping branches 
almost to the base, bark smooth, pale brown, branchlets 
thickened at nodes, nearly cylindrical, twigs with 4 stipular 
wings broad and leafy at the summit of each internode 
decurrent and gradually narrowing to base of internode 
which is quite cylindrical; 1. large, 7-10 in., on a very short 
stout petiole, lanceolate- or ovate-oblong, subcordate at base, 
acuminate, acute, rather thick and coriaceous, glabrous, dark 
green above, paler beneath, veins very strongly marked, 
pellucid, depressed above, prominent beneath, lat. ones very 
numerous, uniting with a strong intramarginal one at a short 
distance from the edge; fl. very small, ped. about as long as 
cal., in lax, slender, stalked axillary and terminal racemes 
shorter than |. and sometimes paniculate with opposite 
divaricate branches, bracts linear, as long as ped.; cal. gla- 


DAD Onagracee. [ Jussiea. 


brous, segm. broadly triangular, acute; pet. connate by their 
tips and falling on expansion of fl.; stam. erect, anth. almost 
sessile ; capsule about I in., ovoid, woody, blackish-brown ; 
seed with oblong obtuse wing, nearly 1 in. 

Moist low country in forest below 1500 ft.; rare. Raitunwelle; 
Ambagamuwa (Thwaites); Udagama (Ferguson); Nillowe Kande; Kuru- 
wita Korale. FI. Feb.-June; yellowish-green, pet. white. 

Endemic. 

The flowers have a very disagreeable odour. 


A very remarkable plant; the leaves have much resemblance to those 
of Memecylon Hookert. ; 


LV.—ONAGRACE/E. 


HERBS, I. opp. or alt., with or without stip., fl. axillary, regular, 
bisexual; cal.-tube adnate to whole or half of ov., segm. 4 or 
5; pet. 4 or 5; stam. equal to or double the number of pet., 
epigynous; ov. inferior or half-inferior, 4~-5-celled, with 
numerous ovules, or 2-celled with a solitary ovule in each 
cell, style simple; fruit a membranous capsule or a hard in- 
dehiscent nut; seeds without endosperm. 


Seeds very numerous. 


Stam. 8 or Io : ; : : : : ; . I. JUSSIAA. 
Stam. 4 . ; : 5 : 2 : : ; . 2, LUDWIGIA. 
Seed solitary . : : ; 4 ; : : . (37 SERBS 


Our few species are all low-country plants, mostly in wet places. 
Trapais completely aquatic. 


1. JUSSIZA, Z. 


Herbaceous or semi-shrubby perennials, |. alt., entire, fl. 
axillary, solitary; cal.-tube completely adnate to ov., and not 
produced beyond it, segm. 4 or 5; pet. 4 or 5; stam. 8 or Io, 
epigynous; ov. inferior, 4- or 5-celled, with very numerous 
ovules in each cell, style very short, stigma large; fruit a 
capsule, irregularly dehiscent between the vertical ribs; seeds 
very numerous.—Sp. 30; 2 in FZ. B. Ind. 


Stems creeping or floating; pet. 5. 


. J. REPENS. 
Stems erect ; pet. 4. 


I 
2. J. SUFFRUTICOSA. 


Jussiza.) Onagracee. 23 


Oo 


1. J. repens, Z. SA. P/. 388 (1753). Béru-diyanilla, S. 
Fl. Zeyl. n. 16g. Moon Cat. 35. Thw. Enum. 123. C. P. 2793. 
Fl. B. Ind. ii. 587. Hook. Bot. Misc. iii. t. supp. 4o. 


Stem herbaceous, elongated, prostrate or creeping or 
floating on water, rooting at nodes, much branched, glabrous, 
succulent, young shoots often pubescent; |. 14-2 in., linear- 
spathulate, much tapering to base, rounded at apex, quite 
glabrous or slightly pubescent, petiole short, slender; fl. about 
2 in., ped. shorter than cal., glabrous, without bracts; cal.- 
tube about 4 in., cylindrical, glabrous or rarely pubescent, 
segm. 5, narrowly lanceolate, acuminate, much shorter than 
tube; pet. 5, shortly clawed, obovate; stam. 10, fil. moderately 
long; fruit about # in., cylindrical, narrow but slightly inflated, 
glabrous or pubescent. 

Borders of ponds and tanks in the low country; common. Fl. May- 
September ; pale yellow or white. 

Throughout the world in hot countries. 

The stems float on the water by means of white spongy bodies 


(modified stipules ?) springing in tufts from each side of the base of the 
petioles. 


2. J. suffruticosa, Z. Sp. P/. 388 (1753). 

Moon Cat. 35. /. villosa, Lam., Thw. Enum. 123. C. P. 2794, 3297. 

FI. B. Ind. ii. 587. 

A semi-shrubby perennial, 2-4 ft., woody below, much 
branched, branches stiff, erect, cylindrical, striate, thickly 
clothed with short spreading hair; 1. 2-4 in., nearly sessile, 
varying from linear to broadly oval but usually lanceolate, 
tapering to base, acute, entire, hairy on both sides, lat. veins 
numerous, prominent beneath; fl. large, 14-1? in., ped. shorter 
than cal.-tube, with 2 lanceolate bracts at summit; cal. hairy, 
tube quadrangular, segm. 4, ovate, attenuate, acute, longer 
than tube; pet. 4, rotundate, shortly clawed, often emarginate, 
pinnately veined; stam. 8, erect, fil. very short; style very 
short, stigma large, quadrate-pyramidal; capsule about 1 in., 
quadrangular, truncate, tapering downwards, hairy, 8-ribbed, 
thin; seeds minute, ovoid, brown, polished. 


Var. 8, subglabra, 7hiw. /.c. FI. Zeyl.n. 170. /. evecta, L. Sp. Pl. 
388 ; Moon Cat. 35. C. P. 1540. 
Lam. Ill. Gen. t. 28, f. 3 (J. angustifolia). 


L. linear, nearly or quite glabrous; fl. smaller, under 1 in.; 
capsule longer, much narrower and not tapering, glabrous. 


Wet places in the moist low country to 3000 ft.; common. Fl. May- 
October ; bright chrome-yellow. 

Throughout the world in warm countries. 

Extremes of var. 3 and the type look so different that I am much in- 


(234 Onagracee. [Ludwigia 


clined to follow Linneus and make 2 species here ; but they are con- 
nected by intermediates. The large broad-leaved woolly plant with 
large flowers is handsome enough for garden cultivation. 


2, LUDWIGIA, Z. 

Annual herbs, 1. alt., entire, fl. solitary, axillary, small; cal.- 
tube adnate to ov. and not produced above it, segm. 4, per- 
sistent; pet. 4; stam. 4, epigynous; ov. inferior, 4-celled, with 
very numerous ovules; capsule as in /wssz@a.—Sp. 20; 2 in 
Fil. B. L[nd. 


Capsule oblong, somewhat inflated . : é . I. L. PARVIFLORA. 
Capsule very narrow, linear . : : : . 2, L. PROSTRATA. 


1. L. parviflora, Roxb. Hort. Beng. 11 (1813). 

Herm. Mus. 17. Burm. Thes. 146. Fl. Zeyl. n. 66. L. opposztzfolia, 
L.. Syst. xii. 135; Moon Cat. 11. Thw. Enum. 123. C. P.1530)13298; 
3299, 3510. ; 

Fl. B. Ind. 11. 588. Wight, Ill. t. ror. 

An erect annual, 6-18 in., stem much branched, often 
angular, glabrous; |. very variable in size, 1-34 in., usually 
about 14 in., lanceolate-linear to lanceolate, much tapering to 
base, subacute, glabrous, thin, petiole obscure; fl. small, on 
very short ped.; cal.-tube cylindrical, oblong, glabrous or 
slightly pubescent, segm. short, acute; pet. very small, not 
longer than cal.-segm.; capsule short, 4-2 in., cylindrical, 
somewhat inflated, glabrous, 4-veined ; seeds very numerous 
in several rows in each cell. 

Low country, especially in the dry region; very common. FI. 
December—May; yellow. 

Throughout India, Malaya, and in Trop. Australia and Africa. 

There is some doubt as to what this should be called. Linnzeus first 
named it L. ferennis, and afterwards L. offosztifolia, both of which 
names must be rejected as contrary to fact. ZL. zeylanica, Pers. (1805) 
would have the next claim, but it was not properly published till 1825 
(in Spreng. Syst. Veg. i. 444). 

Hermann gives a number of native names for this, including 
‘ Kékirinda,’ and says it is used as an eye-medicine. 


2. L. prostrata, foxb. Hort. Beng. 11 (1813). 

Jussiea parvifiora, Moon Cat. 35. Thw. Enum. 123. C. P. 2795. 

PPB elindaia soo. Woshtles ti762. 

Annual, stem over 1 ft. glabrous, flaccid; 1. 2-34 in., 
lanceolate, tapering to both ends, glabrous, thin, petiole 
obscure; fl. very small, nearly sessile; cal.-tube 2 in., linear, 
very slender, segm. lanceolate-linear, short, acute; pet. very 
small, linear; capsule #in., very narrow and slender, linear, 
with very thin transparent walls; seeds in a single row in 
each cell. 


Trapa.] Ouagracee. 35 


Moist low country ; very rare. I have only seen Moon’s specimens 
collected at Kalutara. 
Also in India, Malaya, Japan. 


Gnothera fruticosa, L. A form of this common N. American plant 
has become semi-naturalised by roadsides in the Uva hill-country, having 
probably originated from Hakgala Garden. Another species, perhaps 
(. odorata, is also found in similar localities. 


3. TRAPA, J. 


Aquatic herb; 1. alt., floating, stipulate; fl. solitary, 
axillary; cal.-tube adnate to lower half of ov., segm. 4, in 
2 pair, outer pair persistent and becoming spinous in fruit ; 
pet. 4; stam. 4, inserted outside lobes of epigynous disk ; 
ov. half-inferior, 2-celled, with a solitary pendulous ovule in 
each cell; style slender, surrounded at base with a disk of 
S rounded lobes, stigma capitate; fruit large, bony, in- 
dehiscent, horned, angular, 1-seeded ; seed large, completely 
filling fruit, cotyledons extremely unequal, one very large and 
thick, the other merely a very small scale, radicle superior, no 
endosperm.—Sp. 2; both in F72. B. Jud. 


T. bispinosa, Roxb. Hort. Beng. 11 (1813). Tkiliya, S. 
Thw. Enum. 124. C. P. 2932. 
BIB: ind. ii. 590. Roxb. Pl. Cor: t. 234. 


An aquatic, stems long, flexuose, ascending in the water, 
cylindrical, smooth, the more submerged part thickly set with 
pairs of green pectinate spreading organs (? roots) coming off 
from immediately below the position of stip. of fallen lL, and 
1-1} in. long; 1. crowded in upper part of stem, petiole at 
first short, but growing as the internodes lengthen to 4 in., 
cylindrical, upper portion roughly hairy, dilated at a short 
distance below the end into a large fusiform, spongy float, 
lower portion glabrous, blade floating on surface of water, 
I} in. by 1} in. wide, rhomboid-triangular, truncate and entire 
at base, irregularly incise-serrate above, acute, very glabrous 
and shining, dark green mottled with brown above, densely 
pubescent and reddish-purple beneath, stip. linear, mem- 
branous, caducous; fl. few, solitary, axillary, on short, stout, 
hairy ped.; sep. subacute, pubescent outside; pet. ovate, 
delicate; fruit ripening under water, 14 in. wide, turbinate, 
somewhat compressed, extended above at each end into a 
spreading, flattened, spinous horn, the top tipped in centre by 
a short, sharply pointed, conical beak, each side at the base 


236 S) amyaac é@. [Casearia. 


with a projecting lobe, brownish-black; radicle immediately 
beneath the central beak, whence it protrudes in germination. 


In tanks in the dry region; rather common. Also in the moist low 
country; very rare, ¢.g., Heneratgoda. Fl. Feb.; white. 

Also in India, Malaya, Trop. Africa. 7. dccornis, L. f. of China, where 
the seeds are much used for food, is perhaps a mere variety. 

The pectinate submerged organs cannot be considered as leaves (as 
in Fl. B. Ind., following Wight); their position suggests a stipular nature, 
and they are so called by Roxburgh, who has well figured and described 
them (1. c.), but there are real stipules also present with the young leaves. 


LVL—SAMYDACEE. 


TREES or shrubs, |. simple, alt., with very small stip.; fl. small, 
regular, bisexual; cal. free or adnate to lower part of ov., 
segm. 5 (or 4); pet. 5 (or 4), or 0, imbricate, stam. 4-10, peri- 
gynous, alternating and often connate with large staminodes ; 
ov. superior or half-inferior, 1-celled, ovules parietal, style 
simple or 3 or 4; fruit a soft capsule, loculicidally dehiscent 
by 2 or 3 valves; seeds usually few, surrounded by a fleshy 
aril, embryo seaigihe | in axis of endosperm. 
Has strong affinities with Bzracee. 


Peto: 
Fl. in axillary fascicles . : : : : . I. CASEARIA. 
F]. in terminal panicles 5 4 : : : . 2. OSMELIA. 
Pet. 4 or 5 : 4 : : . 3. HOMALIUM. 


Osmelia and Figpiiiies are eonanel to the moist region, the former 
ascending a little into montane zone, to which Casearia cortacea is con- 
fined. C. zomentosa occurs only in the dry region, and C. escudenza in all. 


I. CASEARIA, /acg. 


Trees or shrubs, |. alt., stip. minute, fl. small, in axillary 
elusters; cal. free, deeply cut into 5 segm.; pet. 0; stam. 6 or 
8, with alternating staminodes, all slightly connate at base, 
hypogynous; ov. superior, 1-celled with several ovules, style 
simple ; fruit half-fleshy, dehiscent by 2 or 3 valves; seeds 
few, surrounded by a red aril, embryo straight, in axis of 
endosperm.—Sp. 80; 14 in FZ. B. Lud. 


L. glabrous. 
L. lanceolate or oval 
L. obovate 

L. pubescent beneath. 


C. ESCULENTA. 
C.:CORIACEA. 
C. TOMENTOSA. 


Go Noe 


Casearia.] Samydacee. a7 


1. C. esculenta, Rox. F7. Ind. ii. 422 (1832). Wal-waraka, S. 
KRakkaipalai, 7. 

C. ovata, Moon Cat. 35 (? Willd.). Vavreca* zeylanica, Gaertn. Fruct. 
i. 290. C. zeylanica, C. Championit, C. varians, vars. a, ovata, and 8, 
minor, Thw. Enum. 19. C. P. 415, 2604, 2608, 2657, 3365. 

Fl. B. Ind. ii. 592. Bedd. Fl. Sylv. t. 208 (C. vavzans). 

A shrub or tree with slender branchlets, bark yellowish- 
white, smooth, young parts glabrous, |. 2-5 in., from narrow- 
lanceolate to oval, tapering to base, shortly acuminate, obtuse 
or subacute, entire or faintly serrate in upper part, perfectly 
glabrous, rather thick, petiole short, stip. persistent; pet. 
stout, longer than cal., and articulated at the base which is 
surrounded by numerous very small bracts, few or many on a 
raised boss in axils of past and present |.; cal.-segm. ovate, 
obtuse, persistent ; stam. 6 or usually 8, staminodes as long as 
fil, oblong, ciliate or hairy; fruit about #in., broadly ovoid, 
apiculate, glabrous or very slightly pubescent, orange-yellow, 
dehiscing by 3 (or 2), thick valves; seeds several, almost en- 
tirely covered by the large fleshy lacerate scarlet aril. 


Both moist and dry regions, and extending up to 5000 ft. Fl. Feb. 
April; greenish; fruit orange. 

Also in South India and the Malay Peninsula. 

Moon gives the native name ‘ Wal-munamal.’ Possibly this is S. 
ovata, Willd., based on Rheede, Hort. Mal. iv. t. 49, in which case 
that name would take priority over S. esculenta. I agree with Clarke in 
combining the species of Thwaites. The Fl. B. Ind. quotes C. P. 2603 
as var. azgusta, with narrow lanceolar leaves, which number is referred 
by Thwaites to his C. zeylanica. 

Wood pale yellow, moderately hard and heavy. The wood and leaves 
are used medicinally; the fruit is eaten. 


2. ©. coriacea, 7hw. Enuzi. 20 (1858). 

C. varians, var. y, obovata, Thw. Enum. 20. C. P. 465, 1217. 

Fl. B. Ind. 11. 592. 

A moderate-sized tree, twigs glabrous; |. 2-3 in., obovate- 
oval or obovate, tapering to base, obtuse or rounded at apex, 
entire, glabrous, more or less coriaceous, venation conspicu- 
ous, reticulate, petiole very short; ped. short, slender, glabrous, 
arranged as in the last; cal.-segm. rounded, stam. 6 (or 8), 
staminodes about half as long as fil., ciliate, ov. glabrous; 
fruit about #in., ovoid, apiculate, smooth, 2-valved, orange. 

Upper montane zone above 6000 ft.; rare. Adam’s Peak; Nuwara 
Eliya. Fl. Jan.; green. 

Endemic. 

C. P. 465, from Adam’s Peak, has the 1. more coriaceous, and with a 
more copious reticulation. 


* Vareca, from the Sinhalese name. 


238 Samydacee. [Homalium. 


3. C. tomentosa, Roxb). Fl. Ind. ii. 421 (1832). 

hy Enum. to, 46. 221248; 

Fl. B. Ind. 11. 593. Brandis, For. Fl. t. 31. 

A shrub or small tree, twigs pubescent; |. 3-4 in., lanceo- 
late-oblong, acute at base, acute or obtuse at apex, shallowly 
crenate-serrate, nearly glabrous above, pubescent beneath, 
petiole 1? in., pubescent; fl. not seen; fruit about ? in., ovoid, 
apiculate, bluntly 6-angled, 3-valved. 

Dry region; very rare. Jaffna and Sigiri (Gardner). 

Also in India, Malaya, N. Australia. 

I have seen only Gardner’s specimens. 


2, OSMELIA, 7iw. 


Tree, |. alt., with minute deciduous stip.; fl. small, in long 
terminal panicles; cal. free, with 5 deep segm.; pet. 0; stam. 
10, 5 alternating with large hairy bifid scale-like stamin- 
odes and 5 coming off from them between their lobes; ov. 
superior, I-celled, with 3 placentas and few ovules on each, 
styles 3, short; fruit a subcoriaceous 3-valved capsule; seeds 
1-4, surrounded by a fleshy red aril, embryo straight, in axis 
of endosperm.—Sp. 3; 1 in FZ. B. Ind. (the other 2 from the 
Philippine Is.). 

O. Gardneri, 7/w. Enum. 20 (1858). 


CE I216: 
Fl. B. Ind. 11. 592 (O. zeylanica). Bedd. Fl. Sylv. t. 209. 


A moderate-sized tree, with smooth grey bark, twigs 
cylindrical, pilose; 1. 24-4 in., lanceolate, acute at base, sud- 
denly and shortly acuminate, obtuse, entire, glabrous, veins 
slender, conspicuous, petiole }-3in.; fl. numerous, almost ses- 
sile, surrounded at base by a small cup-like bractlet, in inter- 
rupted spicate racemes arranged in long drooping terminal 
panicles; cal.-segm. rounded ; ov. very hairy; fruit about $in., 
subglobose, densely pubescent, purplish. 

Moist region at 2000-4000 ft.; rare. Hantane (Gardner); Uduwelle; 
Alagala; Potupitiya, Kukul Korale. Fl. July, August; greenish-white, 
tinged with pink. 

Endemic. 

3. HOMALIUM, /acg. 


Large tree, ]. alt., simple, fl. small, in long axillary race- 
mose panicles; cal.-tube adnate to lower half of ov., segm. 4 
or 5; pet. 4 or 5; stam. 4 or 5, opp. pet., alternating with 
rounded, pubescent, bilobed staminodes; ov. 4-inferior, 1-celled, 


Passifloracee. 239 


with 4 clusters of ovules pendulous from near top, styles 4, 
long; fruit capsular.—Sp. 30; 11 in FZ. B. Ind. 


Sometimes regarded as a separate order, and often placed near 
Chailletiacee. 


H, zeylanicum, Senzh. in Journ. Linn. Soc. iv. 35 (1860). Liyan, 
Liyangu, S. 

Blackwellia zeylanica, Gardn. in Calc. Journ. Nat. Hist. vii. 452. 
Thw. Enum. 79, 410. C. P. 388. 

Fl. B. Ind. ii. 597. Wight, Ic. t. 1851 (B. ¢etrandra). Bedd. FI. Sylv. 
t. 210. 

A very large tree, with a straight trunk, bark white, rather 
rough, breaking off in irregular pieces; 1. 24-5 in., oval, 
narrowed to base, shortly acuminate, acute, serrate-crenate, 
glabrous and shining on both sides, rather thick, veins some- 
what prominent and purplish-red beneath, petiole very short; 
fi. very numerous, articulated on short ped., closely arranged 
in numerous dense corymbose clusters, on a long, narrow, 
interrupted pendulous panicle, 6-9 in. long; cal.-tube glabrous, 
segm. oblong, obtuse, pubescent; pet. spathulate-oblong, ob- 
tuse, twice as long as cal.-segm., pubescent, ciliate; stam. 
longer than pet.; fruit not seen. 

Moist low country to 3000 ft.; rather common. FI. March-June; 
greenish-white. 

Also in Malabar. 

The young leaves are a fine crimson colour. In both Wight’s and 
Beddome’s figures the panicles are erroneously represented as erect. The 
cal.-segm. and pet. are precisely alike except in size. I have never been 
able to find a fruit, nor is it described (in this or any of the Indian species) 
in the books I have consulted. 

Wood pale brown, the heart-wood darker, heavy, very hard and close- 
grained, durable. 


Turnera ulmifolia, L. (Order Turneracee), a West Indian and Trop 
American species, is a very common weed of roadsides and waste ground 
about Colombo and other places in the low country. The flowers are 
bright yellow, and the leaves have a musky odour. There is a figure in 
Bot. Mag. t. 281. 


LVII.—PASSIFLORACE. 


PERENNIAL herbs climbing by tendrils of the inflor.; fl. 
regular, unisexual, dicecious, in axillary cymes; male fl.:— ~ 
cal. campanulate, segm. 5, imbricate; pet. 5, distant, included 
in cal.-tube; disk (corona) a row of white cilia arising from 
base of cal.-tube, and 5 erect glands opp. segm.; stam. 5, peri- 


240 Passifloracee. [Modecca. 


eynous, inserted at base of cal.-tube, distinct or connate ; fem. 
fl.:—cal. and pet.as in male; staminodes 5, erect, surrounding 
ov.; Ov. superior, stalked, 1-celled, with 3 parietal placentas, 
ovules numerous, stigmas 3, feathery or fringed; fruit a cap- 
sule, dehiscing loculicidally by 3 valves; seeds several, en- 
veloped in a pulpy aril, flattened, muriculate, embryo straight 
in fleshy endosperm. 


Our single genus is not typical for the Order, belonging to the tribe 
Modeccee. Both our species are low-country plants. 


MODECCA,* Lam. 
For characters, see Order.—Sp. about 25; 6in FZ. B. Ind. 


Fl. small, } in. long, stigmas sessile . ; ; . I. M. WIGHTIANA., 
Fl. large, 3 in. long, styles long : : : . 2. M. PALMATA. 


1. M. Wightiana, Wall, Cat. n. 6764 (1828). 

Thw. Enum, 128. €, P. 1621. 

Fl. B. Ind. ii. 601. Wight, Ic. t. 170. 

Small perennial herb, stems long, flexuose, wide-spreading, 
very slender, glabrous; |. distant, few, readily disarticulating, 
2-3 in., variable in form, cordate-ovate or triangular or more 
or less 3-lobed, cordate at base, acute at apex, entire or faintly 
dentate, glabrous, very thin, often minutely brown-dotted on 
upper surface, petiole 3-1 in., dilated at summit, where isa large 
prominent gland; fl. small, 4in., on slender articulated ped., 
in two short opp. cymes near the end of a long slender axillary 
curved peduncle, which is continued beyond as a much-coiled 
circinate tendril; cal.-tube campanulate-turbinate, segm. 
about as long as tube, ovate, acute ; pet. rather shorter than 
cal.-segm., ovate, inserted about middle of cal.-tube, disk 
(corona), a ring of hair arising from cal.-tube just below pet.; 
male fl.:—stam. connate at base, anth. ovoid; fem. fl.:—ov. very 
shortly stalked, surrounded by 5 erect staminodes, ovoid, 
tapering, style o, stigmas fringed; fruit very shortly stalked, 
the stalk surrounded by persistent cal, about I%in., not 
pointed, valves chartaceous, smooth, brownish-yellow; seeds 
about 8, + in., surrounded by the aril, ovoid, compressed, 
strongly muriculate on sides. 

Dry country; ratherrare. Jaffna; Puttalam; Karativu; Trincomalie; 


Kekirawa. Fl. Aug.—Nov. 
Also in S. India. 


* Modecca is given as the Malabar name by Rheede, Hort. Mal. iv. 39. 


Modecca. Passifloracee. 2A1 
4 


2. M. palmata, Zam. Encycil. iv. 209 (1796). Hondala, Potu- 
honda, 5S. 

Herm. Mus. 41. Burm. Thes. 49. FI. Zeyl. n. 353 (in part). Bryonda 
palmata, L. Sp. Pl. 1012. M. tuberosa, Roxb., Moon Cat. 48. Thw. 
Enum. 128. C.P 1627. 

Fl. B. Ind. ii. 603. Wight, Ic. t. 201. 


A large perennial herb, becoming woody at base, stems 
long, thickened at nodes, cylindrical, smooth and _ shining, 
mottled with purple and covered with a ‘bloom,’ slightly 
branched; 1. large, 4-5 in., broader than long, usually very 
deeply palmately 5-lobed (rarely 3-lobed or undivided), very 
glabrous and shining, especially beneath, lobes oval, narrowed 
at base, shortly acuminate, acute, entire, veins conspicuous, 
vinous-red, prominent beneath, where are often dots of red 
colour, and between the bases of main veins 4 circular, flat, 
pellucid glands, as well as 2 larger ones on the outer side of 
the lateral veins, petiole about 2 in., stout, curved, with 2 short 
setaceous stip. at base; fl. rather large, on short articulated 
ped. in two opposite cymes of three, on a long axillary 
peduncle, which is continued beyond them as a long simple 
tendril; cal. 2 in., truncate at fleshy base ,broadly and squarely 
campanulate, glabrous, segm. triangular, acute, slightly 
spreading; pet. small, distant, linear, inserted at base of cal.- 
tube, rather larger in male fl, more or less hairy, disk wide, 
with 5 circular pits opp. segm., with white cilia (corona) on 
their outer side, and a short blunt, erect process within each; 
male fl.:—stam. distinct, fil. very short, anth. linear, ov. rudi- 
mentary; fem. fl.:—staminodes 5, erect, immediately sur- 
rounding ov.,small, acute, ov. shortly stalked, globose, smooth, 
tapering into 3 long styles, stigmas much divided, feathery ; 
fruit nearly 2 in. on a short stalk, globose, apiculate, 
smooth, orange, splitting into 3 fleshy valves; seeds on long 
stalks, nearly 4 in., muriculate and pitted, black, each enclosed 
in large pulpy aril. 

Moist low country; common. FI. May-August; greenish-white, 
tinged with pink. 

Also in the Indian Peninsula. 

The only specimen now in Hermanw’s Herb. representing Bryonia 
palmata, L.is a very bad one, and no doubt is B. dacinzosa ; but Linnzeus 
seems to have had specimens of the present plant also before him in 
writing his description. 

The fruit is poisonous, and has caused death. The root is used in 
medicine. 


Passiflora suberosa, L., with small green flowers and purple fruit like a 
small grape, is occasionally found as a weed in the moist low country. It 
is figured as a Ceylon plant under the name of ?. Walkerie in Wight, 
Ill. t, 108. Native of the W. Indies. 

PART Il R 


2A2 Cucurbitacee. 


P. fetida, L., is extremely common, and has spread into forest in the 
dry country, where it has in many places quite the look ofa native. It 
is also indigenous to Tropical America, and is easily to be known by the 
beautifully pectinate and moss-like involucre. 


P. edulis, Sims, has escaped from cultivation in the hill country. It 
has an edible fruit which goes by the name of ‘ Sweet-cup’ in Jamaica. 


P. stipulata, Aubl., P. glauca, Ait. (non Kunth), is common by road- 
sides in the montane zone as an escape from cultivation, and in places 
quite naturalised. Native of Trop. S. America. 

We have no native species of Passzflora. 


LVIII—CUCURBITACEE. 


HERBACEOUS, climbing by spirally twining tendrils arising 
solitarily at the nodes by the base of the 1.; 1. alt., simple 
(rarely compound), often palmately divided; fl. regular, 
unisexual, moncecious or dicecious; male fl.:—cal.-tube usually 
short. campanulate, segm. 5, small; pet. 5, usually more or 
less connate (rarely free), inserted on cal.-tube, valvate or 
imbricate; stam. 3 (rarely 5), inserted on cal.-tube, anth. 
distinct or connate, one I-celled, two 2-celled, cells linear, 
conduplicate or sigmoid or nearly straight; fem. fl.:—cal.-tube 
adnate to ov. and often produced beyond it, segm. as in male; 
pet. as in male; staminodes 3 or 5 or none, ov. completely 
inferior, 1-celled, with 3 large fleshy parietal placentas often 
meeting in centre (hence apparently 3-celled), ovules usually 
numerous, horizontal, rarely few and pendulous, stigmas 3, 
large; fruit fleshy, generally indehiscent; seeds usually 
numerous, surrounded with pulp, no endosperm. 

The most recent account of this difficult Family is that by Cogniaux 
in Monogr. Phanerog. ili. (1881). The cal. and pet. are often so fused 
below that the cal.-segm. look: like appendages on the outside of a cor.- 
tube. 

Of the 26 Ceylon species the greater part are plants of the low 
country, and many occur in both moist and dry regions. But the latter 
has the greater number, and Cz¢vullus, Rhynchocarpa, Corallocarpus, 
Ctenolepis, and Cucumdzs trigonus are confined to it, whilst Zanonza and 
Zehneria hastata are restricted to the moist region. Seven of the low- 
country species extend into the montane zone, and the following are found 


only there: Cerastocarpum, Gynostemma, Trichosanthes integrifolia, and 
Mukia leiosperma. Only 3 specimens are endemic. 


Trichosanthes.] Cucurbttacee. 24 


(SS) 


Stam. 3 (sometimes 5); fem. fl. solitary. 
Anth.-cells conduplicate or sigmoid. 
Pet. strongly fimbriate . : 5 : . I. TRICHOSANTHES. 
Pet. (or cor.-lobes) not fimbriate 


Cal.-tube of male fl. long, tubular 2. GYMNOPETALUM. 
Cal.-tube of male fl. short, campanulate. 
Anth. more or less connate. 
Cor. campanulate, cut about half way 
down : . 3. CEPHALANDRA. 
Cor. cut nearly to the base. 
Fruit tubercled. 4. MOMORDICA. 
Fruit smooth 5. CUCUMIS. 
Anth. distinct. 
Male fl. in racemes 6. LUFFA. 


Male fl. solitary or in clusters. 
Fruit with hard pericarp . ; 5 ge CIABRIGILILITIS, 
Fruit a soft berry F 8. BRYONIA. 
Anth.-cells straight or slightly curved. 
Fruit globose or ovoid, not beaked. 


Male fl. in clusters ‘ : : . 9. MUKIA. 
Male fl.in racemes. i : . 10. ZEHNERIA. 
Fruit beaked, indehiscent. 
Fruit glabrous . : : : UE LOE RTA. 
Fruit very hairy . 12. RHYNCHOCARPA. 


Fruit beaked, dehiscent round the base. . 13. CORALLOCARPUS. 
Fruit depressed, not beaked. 


Depressed-globose, seeds 3-6. . 14, CERASIOCARPUM. 
Depressed-turbinate, seeds 1-2 . 15. CTENOLEPIS. 
Stam. 5 ; fem. fl. in racemes or panicles. 
L.compound . : : ; : : . 16. GYNOSTEMMA. 
L. simple . : 5 : ; : ; . 17. ZANONIA. 


1. TRICHOSANTHES, /. 


Annual or perennial, tendrils branched or simple, 1. pal- 
mately lobed or entire, fl. dicecious, rarely moncecious, male 
usually in racemes (rarely solitary), fem. solitary ; cal.-tube 
very long, segm. large or small, pet. distinct or nearly so, 
fimbriate; male fl.:—stam. 3, inserted in cal.-tube, fil. short, 
anth. connate, linear-oblong, cells conduplicate; fem. fl.:— 
ov. with numerous horizontal or pendulous ovules on 3 parietal 
placentas, style simple, stigmas 3; fruit large, globose or 
narrow, red; seeds numerous, compressed, each enclosed in an 
envelope of coloured pulp.—Sp. 40 (Cogn.); 12 in #2 B. Lund. 


Male fl. with large sheathing bracts ; ; . I. T. PALMATA. 
Male fl. without or with very small bracts. 
Male fl. in racemes. 
L. much longer than broad 
L. broader than long 
Male fl. solitary . 


T. NERVIFOLIA. 
T. CUCUMERINA 
T. INVEGRIFOLIA. 


EYP 


244 Cucurbttacee. [ Trichosanthes. 


I1..T. palmata, ford. FV. [nd. iii. 704 (1832). Titta-hondala, S. 

Lryonia palmata, Moon Cat. 67. Thw. Enum. 127. TZ. bracteata, 
Voight, Cogn. Mon. 375. C. P. 1626. 

JIL 18}. lial, jie COS, Wate, ING THis iy, KOs, 


Stems long, woody below, wide-climbing, angular, slightly 
scabrous, tendrils 3- or 2-fid; |. 4-5 in. and about equally 
wide, variable, more or less deeply palmately 3- or 5-lobed, 
lobes acute, more or less dentate-serrate, glabrous, often 
scabrous with small scales above and on the veins’ beneath, 
petiole about I in., scaly-scabrous; racemes of male fl. 6-9 in., 
drooping, fl. large, over 2 in. nearly sessile, distant, each in 
the axil of a very large broadly wedge-shaped, glabrous or 
pubescent, lacerate, persistent bract 1 in. or more long, often 
set with broad flat glands; cal.-segm. leafy, laciniate, pet. 
rather longer than cal.-segm., I in. wedge-shaped, with many 
and long filiform laciniz; fem. fl. shortly stalked; cal.-tube 
nearly 2 in., segm. acuminate, pet. shorter and narrower than 
in male, style long; fruit large, 2-24 in., globose with a blunt 
nipple, smooth, brilliant scarlet-crimson, pericarp thick; seeds 
densely packed, about 4 in. oblong, compressed, smooth, 
brownish-grey, each enveloped in dark-green pulp. 


Var. 8, tomentosa, feyne ex Fl. B. Ind. 1. c. 
L. divided less than half way down, tomentose beneath. 


Low country ; rather common, especially in dry region. Var. #, rare. 
Peradeniya. Fl. Feb.—June; bright pink. 

Throughout the Eastern Tropics. 

The large scarlet fruits are very conspicuous and ornamental. The 
pounded fruit is employed as an application to ulcers. 

Cogniaux (Mon. Phan. iii. 376) quotes C. P. 1302 for this, which is a 
Fern. 


2. T. nervifolia, LZ. Sf. P/. 1008 (1753). 

T. caudata, Willd., Moon Cat. 66. Thw. Enum. 127. C. P. 1608. 

Fl. B. Ind. i. 609. Rheede, Hort. Malab. viii. tt. 16 (fem.) and 17 
(male). 

A large perennial, stems somewhat woody below, flexible, 
thickened at nodes, much branched, branches slender, striate, 
glabrous, tendrils bifid; 1. 24-34 in., ovate-lanceolate, cordate 
or rarely lobed at base, very acute, mucronate, distantly 
denticulate, glabrous, dark green, paler beneath with promi- 
nent reticulate venation, petiole 4-1 in.; male fl. on short 
ped., 4-12 in pedunculate corymbose racemes, bracts small, 
caducous; cal.-tube #-1}in., very narrow, inflated above, 
segm. minute, setaceous, pet. oblong, acuminate, with the fim- 
briza much branched and very long at end of pet. but shorter 
on the sides, doubled inwards in the bud; fem. fl. shortly 


Trichosanthes. | Cucurbitacec. 245 


stalked, solitary, cal.-tube nearly 2 in., very much produced 
above ov., segm. longer than in male fl.; fruit 14 in., ovoid, 
shortly beaked, smooth, scarlet, pericarp thin; seeds few with 
very long stalks, ovoid, compressed, thickened at margins, 
each enclosed in an envelope of scarlet pulp. 

Moist low country and lower montane zone; rather common. FI. 
Jan.—July ; white. 

Also in Western India. 


3. T. cucumerina, Z. Sp. P/. 1008 (1753). Dummeélla, S. 
Pudal, 7. 

Moon Cat. 66. Thw. Enum. 126. C. P. 2806. 

Fl. B. Ind. ii. 609. Rheede, Hort. Malab. viii. t. 15. 


Annual, stems slender, glabrous or slightly hairy, tendrils 
3-fid; |. 2-4 in. long and considerably broader, very cordate, 
with an excavated sinus at base, more or less deeply pal- 
mately 3- or 5- or 7-lobed usually about half way down, lobes 
rounded or subacute, distantly denticulate, glabrous or nearly 
so above, more or less pubescent beneath, thin; fl. often 
moncecious, male in long-stalked racemes, female often in 
same axil, solitary, shortly stalked; male fl.:—cal.-tube about 
1 in., dilated at top, segm. narrowly lanceolate, acute, spread- 
ing, pet. 2 in., lanceolate-oblong, with a tuft of curled lacinize 
at apex; fem. fl.:—cal.-tube gradually dilated upwards, lower 
part often hairy; fruit 14-2 in., ovoid-fusiform, tapering at 
both ends, with a long sharp beak, smooth, at first striped 
with white, afterwards scarlet, pericarp thin ; seeds compressed, 
surrounded with red pulp. 


Var. B, laciniosa, 7hw./.c. C. P. 1620. 


L. deeply cut nearly to the base, segm. oblong, densely 
pubescent beneath. 


Low country, common; var. #, especially in the dry region. FI. 
Sept.-December ; white. 

Also in India, Malaya, and N. Australia. 

Var. 8 is connected with the type by intermediate forms. 

The plant is much used as a febrifuge medicine ; the bitter and pur- 
gative fruit is also eaten after being cooked. 


T. anguina, L., the ‘ Snake-gourd’ of the English (Patola, S., Podivi- 
langu, T.) is probably only a cultivated variety of the last. The long 
fruit is variously twisted and contorted. It is well figured in Duthie, 
Field and Garden Crops, t. 46, and is only known as a cultivated plant, 
the fruit being a favourite vegetable. 


4. 'T. integrifolia, 7hw. Enum. 127 (1859) non Kurz. [PLATE 
XLI1.] 

T. Thwaitesiiz, Cogn. Mon. Phan. iii. 387. C. P. 1629. 

Fl. B. Ind. ii. 610, 


246 Cucurbitacee. [Gymnopetalum. 


Stems angular, glabrous, tendrils long, simple; |. 24-4 in., 
ovate-oval, rounded or subcordate at base, acuminate, often 
twisted at apex, acute, entire, glabrous, subcoriaceous, strongly 
3-nerved from the base, venation prominent beneath; male 
and fem. fl. both rather large, solitary, axillary, ped. #-1 in.; 
fruit globose, 2 in. diam., smooth, crimson; seeds oblong, 
compressed, smooth, grey, each enclosed in an envelope of 
deep green pulp. 


Lower montane zone; rare. Uduwela, Hantane; Dolosbagie. FI. 
July ; pale yellow. 

Endemic. 

The leaves are like those of S7zz/ax, the crimson fruit as large as an 
apple and very handsome, the pulp surrounding the seeds is a dull, deep, 
metallic green. 

There is no necessity for Cogniaux’s new name, as Kurz’s T. zutegri- 
folia was not published till 1877. 


2. GYMNOPETALUM, 477. 


Perennial herb, tendrils simple, 1. slightly lobed, fi. small, 
dicecious, male in close racemes, fem. solitary; cal.-tube long, 
contracted at top, segm. setaceous; pet. spreading, not fim- 
briate; male fl.:—stam. 3, inserted above middle of cal.-tube 
and included in it, anth. connate, cells conduplicate; fem. fl.:— 
staminodes 3, very small, ov. very hairy, style long, stigmas 3, 
long; fruit fusiform, red; seeds numerous, ovoid, compressed, 
each enclosed in coloured pulp.—Sp. 6 (Cogniaux); 4 in 
TEU Bs SOL, 


+e. Wightii, Arz.in Hook. Journ. Bot. iii. 278 (1841). [PLATE 

al Imani, 75 (CLI, ep, 

PSB inde itor: 

Stems slender, furrowed, bristly, young parts hairy; 
]. 2-4 in. cordate-ovate, with deep narrow sinus at base, 
acuminate, very acute, more or less (usually slightly) 3- or 
5-lobed, distantly denticulate, rough with short coarse bristly 
hairs above, more softly hairy beneath, petiole hispid; racemes 
of male fl. stalked, short, umbellate, crowded, ped. slender, 
bracts small, setaceous; cal.-tube #-1 in., more or less hispid 
or hairy, segm. setaceous; pet. about }in., oblong-ovate, | 
acute; fruit 14-2 in. tapering at both ends, tipped with 
withered cal., scarlet, slightly hispid; seeds 4in., smooth, 
blackish-brown, immersed in pale-red pulp. 

Var. 8, Zeylanicum, A477. /.c. (sp.). Bryonia tubifiora, W. and A. 
Prod. 347. 


Cephaiandra.] Cucurbttacee. 247 


L. more strongly 5-lobed; cal.-tube glabrous or nearly so. 


Low country in both moist and dry regions and lower montane zone ; 
rather common. Abundant about Anuradhapura. Var. 6, Trincomalie, 
1796 (Rottler). Fl. April, October; white. 

Also in S. India. 

Var. 6 is probably not worth distinction, but Cogniaux reaps it up as 
a species under the name G. ¢udzflorum ,; Rottler’s specimens (now in 
Herb. Kew) are the type of 3. ¢ubiflora, W. and A. 

The leaves form an ingredient in curries in the dry country. 


Lagenaria vulgaris, Ser., the ‘ Bottle-Gourd,’ Diya-labu, S., Churat, T., 
is commonly cultivated. Iti is figured in Duthie, Crops, t. "48. mhers ane 
several varieties in the form of the fruit besides the usual bottle- -shape. 


3. CEPHALANDRA, Schirad. 


Perennial, tendrils simple, fl. dicecious, rather large, solitary, 
axillary; cal.-segm. linear; pet. combined about half way up 
to form a 5-lobed campanulate corolla; male fl.:—stam. 3, 
anth. connate, cells conduplicate; fem. fl. —ov. solid, pla- 
centas 3, style short, stigmas very large; fruit ovoid, cylin- 
drical, smooth, red; seeds numerous, ovoid, compressed, each 
surrounded by envelope of coloured pulp.—Sp. 12; 1 in /2 B. 
Ind. (the rest African). 


C. indica, Naud. in Ann. Sc. Nat. ser.5,v. 16 (1866). Kowakka, 
S. Kovvai, 7. 

Bryonia grandis, Willd., Moon Cat. 67. Coccinea indica, W. and A., 
Thw. Enum. 128. C. P. 1069. 

feeelid. 1.621. Burm. Thes: t; 10,1. 1,2. Hook. Ic: Pl. t. 138: 

Stems slender, cylindrical, quite glabrous; 1. 2-4 in., 
variable in form, usually broadly cordate-ovate in outline, 
with the basal sinus triangular, obtuse, apiculate, more or less 
5-lobed, distantly dentate, quite glabrous on both sides, 
rather succulent, dull green above, glaucous beneath, finely 
punctate, provided with several large flat circular glands near 
the axils of the veins beneath, especially at the base; peduncle 
14-2 in.; cal. glabrous, segm. linear, reflexed; cor. 1} in. diam, 
widely campanulate, segm. broad, acute, recurved, pubescent 
inside, veiny; male fl.:—fil. free; fem. fl.:—ov. fusiform, 
glabrous, stigmas long; fruit 1-14 in., fusiform-ovoid, cylin- 
drical, slightly beaked, smooth, bright scarlet when fully ripe; 
seeds oblong-ovoid, much compressed, smooth, yellowish-grey. 

Low country, especially in the dry region in bushy places ; common. 
Fl. June-September ; white, with green veins. 

Also throughout India, Malaya, and Trop. Africa. 

The large white bell-flowers and brilliant scarlet fruits render this a 
very pretty little creeper. The leaves vary greatly in the depth of the 


248 Cucurbitacec. [ Momordica. 


partitions, when very deeply cut nearly to the base it is var. alceefolia, 
Cogn. 1. c. 531, occasionally met with. The structure of the fruit is 
peculiar, the seeds are truly parietal in attachment, but each is enclosed 
in a separate compartment and surrounded by a reddish pulp; thus a 
vertical section looks like that of a small pomegranate. 

I refrain from quoting FI. Zeyl. n. 354 and Lryonza cordifolia, L. for 
this, as Hermann’s specimens are undoubtedly (/ukza scabrella.. Cogniaux 
(I. c. 529), however, adopts Linnzeus’s specific name, and calls this plant 
Coccinia cordifolia. 


4. MOMORDICA, J. 


Annual or perennial, tendrils simple, fl. moncecious or 
dicecious, male in racemes or solitary, fem. solitary; cal.-segm. 
5, oval or narrow; pet. very slightly connate at base; stam. 3, 
anth. slightly connate, cells conduplicate; ov. muricate or 
papillose, style short, stigmas large; fruit ovoid or fusiform, 
beaked, coarsely tuberculate, many-seeded; seeds rather large, 
ovoid, in a pulpy envelope.—Sp. 35 (Cogniaux); 6 in FZ B. 
L[nd. 

F]. moncecious . ; 3 : : : ‘ . I. M. CHARANTIA. 
FI. dicecious. 


Male fl. solitary, with a large hooded bract beneath 2. M. DIOICA. 
Male fl. usually in racemes, without bracts beneath 3. M. DENUDATA. 


1. *M. Charantia, Z. Sf. P/. 1009 (1753). Karivila, Batu- 
karivila, S. Pakal, Nutipakal, 7. 


Herm. Mus. 39. Burm. Thes. 161. Moon Cat. 66. Thw. Enum, 126. 
C2 IP, Wott, 


F]. B. Ind. 11. 616. Wight, Ic. t. 504. Duthie, Crops, t. 64. 


Annual, stems somewhat twining, much branched, sharply 
5-angled, roughly pubescent, young parts hairy; 1. 24-5 in, 
almost circular in outline, very deeply cordate at base, 
palmately cut to beyond the middle into 7 or g lobes, lobes 
acute, apiculate, coarsely spinous-dentate, pubescent on the 
veins on both sides, petiole 1-2 in., channelled above and 
narrowly bordered with decurrent leaf-base; fl. moncecious, 
solitary, axillary, on slender peduncles 2 or 3 in. long, usually 
with a large, sessile, rotundate, entire bract on the lower half, 
and often close to the base; cal.-segm. oval, subacute, pu- 
bescent; pet. twice as long as cal.-segm., rounded, veiny; 
fem. fl.:—ov. finely pubescent, stigmas bilobed; fruit large, 
3-6 in., pendulous, fusiform, usually pointed or beaked, closely 
tubercled and also bluntly muricated, orange-coloured; seeds 
ovoid, large, about 4 in., compressed, each enclosed in red 
pulpy envelope. 


Low country up to 3000 ft., but cultivated or escaped from cultivation 
always. Fl. June-August; lemon-yellow. 


Momordica. Cucurbitacee. 249 


Throughout the Tropics. 

Several varieties are grown, and much used for curries by the natives. 
M. zeylanica, Mill., is a form with shorter fruit. Leaves very bitter to 
the taste (hence the plant was called ‘ Maragosa’ by the Portuguese) with 
a disagreeable smoky odour when bruised. 


2. M. dioica, Roxb. in Willd. Sp. Pl. iv. 605 (1805). Tumba- 
karivila, S. Tumpai, Paluppakal, 7. 

Moon Cat. 66. Thw. Enum. 126. C. P. 197. 

Fl. B. Ind. ii. 617. Wight, Ic. tt. 505 (male), 506 (female). 


Perennial, with tuberous roots, stems somewhat com- 
pressed and 2-edged, striate, glabrous and shining; |. variable, 
2-4 in., broadly ovate in outline, very cordate at base, acute, 
more or less deeply cut into 3 or 5 lobes, distantly dentate or 
denticulate, thin, quite glabrous and shining on both sides, 
minutely punctate beneath, petiole 1-14 in., pubescent, chan- 
nelled above; fi. dicecious, solitary, peduncle about 2 in, 
slender, glabrous or finely pubescent, in the male with a large 
hooded bract a little below the fl. and enclosing it, in the fem. 
with a minute bract below the middle; cal.-segm. distant, 
linear; pet. 3-1 in., lanceolate, acuminate, slightly pubescent; 
fem. fl.:—ov. densely covered with long soft papille, stigmas 
bifid with erect horns; fruit about 2 in., oblong-ovoid, beaked, 
glabrous, evenly covered with equal-pointed papille; seeds 
broadly oblong, compressed, nearly smooth, pulpy covering 
red. 

Low country, especially in the dry region; common. FI. June- 
August; pale lemon-yellow. 

Throughout India and Malaya. 

The leaves vary extremely in the depth of the lobes; they are not 


bitter to the taste. The fruit is bitter, but that of the cultivated forms is 
much less so and is eaten as a vegetable. 


3. M. denudata, Clarke in Fl. Brit. Ind. ii. 618 (1879). 

M. dioica, var. denudata, Thw. Enum. 126. Cogn.l.c. 448. C. P. 1615. 

Fl. B. Ind. ii. 618. 

Annual (?) stems compressed, 2-edged, furrowed, glabrous, 
young parts glabrous; |. 2-4 in., cordate-ovate in outline, very 
acute, more or less deeply 3- or 5-lobed at base, distantly 
denticulate, glabrous, punctate beneath, petiole channelled 
above, slightly pubescent; fl. dicecious, male in a small raceme 
of 4-8 and a separate basal one (often alone produced), ped. 
about 4 in., puberulous, bracts minute, fem. solitary; cal.- 
segm. linear-lanceolate or linear, acuminate, longer in fem. ; 
pet. }-1 in., lanceolate, acute, pubescent, narrower in fem.; ov. 
densely covered with long acute papille; fruit about 14 in., 
ovoid, usually lop-sided, suddenly narrowed into a strong 


250 Cucurbitacec. [ Cucumis. 


blunt beak, covered with short pointed processes; seeds large, 
over }in., ovoid, not compressed, tuberculate. 

Moist region and lower montane zone, 1500-4000 ft.; rather common. 
Matale (Gardner); Hunasgiriya; Maturata; Ramboda; Dimbula; about 
Kandy. Fl. July, Nov., Dec.; lemon-yellow. 

Endemic. 

The leaves are very variable; the central lobe is, however, always the 
largest, and is often elongated and very acuminate. 


5. CUCUMIS, Z. 

Annual (or perennial ?), tendrils simple; fl. rather small, 
monoecious, male in small clusters, fem. solitary; cal.-limb 
campanulate, segm: 5, setaceous; pet; 5) connate sar base; 
male fl.:—stam. 3, fil. very short, anth. connate (or distinct ?), 
connective produced into a lobed appendage; fem. fl.:—style 
short, slender, stigmas very large; fruit fleshy, spherical or 
ovoid, solid, smooth; seeds very numerous, oblong-oval, com- 
pressed, smooth.—Sp. about 25; 4 in FZ. B. Ind. 


L. rather deeply cut into 5 obtuse lobes. : . 1, CsrRiconus: 
L. shallowly cut into 3 or 5 acute lobes. ‘ . 2. C. PUBESCENS. 


1. ©. trigonus, Roxb. Fl. Ind. iii. 722 (1832). Metukku, 7. 
Thw. Enum 127. C. P. 1618. 
Fl. B. Ind. ii. 619. Wight, Ic. t. 497. 


Annual (2), stems numerous, prostrate, more or less hispid; 
l. rather small, 14-2 in., broadly cordate-ovate, more or less 
deeply 5-lobed, lobes rounded, dentate, scabrous on both sides, 
petioles usually longer than 1.; fl. as in C. pubescens, fruit 
not seen. 

Dry region; rather rare. Aripo and Batticaloa (Gardner); Mannar; 
Minneri. 

Throughout India, Affghanistan, Persia. 

I am quite doubtful as to the name of this dry-country plant, of which 
I have not seen the fruit. I suspect our specimens are often merely 
degenerated examples of the common native cucumber, ‘ Rata-kékiri’ 
(a torm of C. saz¢evis, L.), which is much cultivated. 


Wight, |. c., figures the fruit as globular, less than 1 in. diam., and with 
Io stripes down the sides. 


Cc. pubescens, Willd. Sp. Pl. iv. 614 (1805). Gon-kékiri, 
Kékiri, S. 


C. maderaspatanus, Moon Cat. 67 (non L.). Thw. Enum. 127. C. P. 


3534- 
Fl. B. Ind. 11. 619 (under C. ¢7Zgonus). Wight, Ic. t. 496 (?). Royle, 
Ill. Him. t. 47 f. 3 (C. Hardwickiz). 


Stems long, bluntly angular, rough with hooked prickly 
hairs on ridges; |. 3-4 in., broadly cordate-ovate, with the 


Luffa.] Cucurbrtacee. 251 


basal lobes rounded, usually shallowly cut into 3 or 5 acute 
lobes, slightly dentate, very roughly hairy on both sides, 
petiole stout, often as long as l., deeply sulcate above, often 
much twisted, very harsh with prickly hairs; fl. shortly stalked, 
in. wide; cal.-limb campanulate, very hairy, segm. setaceous; 
pet. united for about } up, rounded; male fl.:—anth. small, 
connate; fem. fl.:—cal.-tube constricted above ov., ov. ovoid, 
with scattered, bulbous-based, bristly, deciduous hairs; fruit 
ovoid-globose, slightly trigonous on section, about 14 in. by 14 
wide, quite glabrous, obscurely striped with dark and light 
green, solid, fleshy, with 3 partitions; seeds very numerous, 
horizontal, narrowly ovoid, compressed, smooth. 

Waste ground in the low country; common. FI. Aug., Sept.; yellow. 

Throughout India and Malaya. 

The name of this is very doubtful; the fruit is always quite glabrous. 
It appears to be a native, and may be the wild original of C. sazcvus, L., 
the cucumber, or C. M/e/o, L., the melon. Cogniaux refers C. P. 3534 to 
the former (but the fruit is at all stages quite without spines or tubercles) ; 
but he puts C. pubescens, Willd., to the latter. (Under C. A7/e/o he quotes 
by error C. P. 1104, which is a 77vzanthema.) Good figures of some of 
the Indian cultivated forms of C. Me/o and C. sativus will be found in 
Duthie, Crops, tt. 49-54. 

The fruit is too bitter to be eaten, but is used in medicine. The plant 
is never cultivated. 


6. LUPFA,* Cav. 


Large annual herbs, stems angled, tendrils 3-fid, 1. pal- 
mately lobed, fl. large, moncecious, male in long racemes, fem. 
solitary from same axil; cal.-limb cup-shaped, segm. 5, lan- 
ceolate, acuminate; pet. connate for more than half way up, 
obovate-oblong; male fl.:—stam. 3 or 5, anth. distinct, cells 
sigmoid; fem. fl.:—cal. contracted above ov., ov. long-oblong, 
style short, thick, stigmas large; fruit large, ovoid-oblong, 
fibrous when ripe, not very succulent, 3-celled, indehiscent; 
seeds numerous, ovoid, compressed.—Sp. 10; 5 in /V. B. Lnd. 


Stam. 5; fruit not ridged . ; : : , tno eG MP ACA: 
Stam. 3; fruit sharply 10-ridged__.. : : . 2. L. ACULANGULA. 


1. *L. egyptiaca, M7/7//. Vict. ed. 8 (1768). Niyan-wéta-kolu, 
S. Pikku, Pichukku, 7. 

L. cylindrica, Roem., Moon Cat. 66. L. pentandra, Roxb., Thw. 
Enum. 126. C. P. 2805. 

At, B. Ind. ii. 614. Wight, Ic. t. 499 (ZL. pentandra). Duthie, Crops, 
t. 63. 


Stems very stout, 5-angled, twisted, slightly pubescent, 


* Luffa, from the Ex gyptian or Arabian name ‘often spelt ‘ Loofah.? 


252 Cucurbitacec. [Luffa. 


young parts more so; l. large, 4-8 in., orbicular in outline, 
often broader than long, very cordate at base, usually more or 
less 7-lobed, lobes acute, distantly denticulate, finely scabrous 
on both sides, petiole 1-3 in., angular, slightly scabrous; male 
41. numerous in racemes 4-8 in. long, 14 in. wide, ped. 4-4 in., 
pubescent, with a small fleshy bract near the base bearing 3 
or 4 large immersed glands, buds pointed, fem. fl. larger, 
over 2 in., solitary, on peduncle 3-6 in. long; cal.-segm. lan- 
ceolate, acute; pet. obovate-oblong; male fl.:—stam. 5, dis- 
tinct; fem. fl.:—cal.-tube produced a little beyond ov., ov. 
oblong, cylindrical, glabrous or pubescent; fruit very large, 
6-12 in. or more, cylindrical or somewhat trigonous, not 
ribbed, blunt at end; seed nearly 4 in., oval, much compressed, 
narrowly winged, rough on sides, grey. 

Low country to 3000 ft.; common in native gardens, but scarcely a 
wild plant. Fl. July; yellow. 


Throughout the Tropics, cultivated. 
The young fruits are eaten as a vegetable. 


2. L. acutangula, fort. Hort. Beng. 70 (1813). Weta-kolu, 
Dara-weta-kolu, S. Peypichukku, 7. 

Herm. Hort. Lugd.-Bat. Cat. 482. Fl. Zeyl. n. 352 (part). Momordica 
Luffa, L. Sp. Pl. 1009 (part); Moon Cat. 66. Thw. Enum. 126. C. P. 1624. 

Fl. B. Ind. ii.615. Duthie, Crops, t. 62. 


Very close to L. egyptiaca, from which it differs in its 
much more slender stems with sharper angles, rather smaller 
fl, 3 stams., strongly ridged ov., and in the fruit, which is 
6-12 in., clavate-oblong, tapering to base, very obtuse, smooth, 
longitudinally ribbed or almost winged with 10 sharp angles 
or ridges, and with the seeds over 4in., not winged, oblong- 
ovoid, much compressed, slightly corrugated on sides, black. 


Var. 6B, amara, Foxd. (sp.). Thw. Enum. 417. C. P. 1623. 
Seeds smaller, # in., more corrugated, black or white. 


Largely cultivated in native gardens in the low country, but nowhere 
wild; var. 6 in the dry region, possibly a native. Fl. yellow; July. 

Throughout the Tropics, but introduced in the New World. 

The fruit is much used when half ripe and still succulent for curries 
and as a vegetable, and is preferred here to L. egyftzaca, but that of 
var. 8 is too bitter for use. 

Hermann’s figure (there is no specimen), called Womordica cylindrica 
by Linn., represents this species, not Z. egypiiaca. 


Benincasa cerifera, Savi, the ‘Ash-Pumpkin’ (4/u-puhul, S., Puchint, 
T.), is much cultivated by the natives for the fruit, used as a vegetable; 
is also candied as a sweetmeat. It is grown throughout the Eastern 
Tropics, and in China and Japan. Figured in Duthie, Crops, t. 45. 


Bryonia.| Cucurbitacec. 253 


7. CITRULLUS, Schrad. 


Annual or perennial, tendrils 2-3-fid., fl. rather large, 
moneecious, all solitary in the axils; cal.-limb campanulate, 
segm. 5, narrow; pet. 5, united nearly half way up to form a 
cup-shaped cor.; male fl.:—stam. 3, small, anth. distinct, cells 
conduplicate; fem. fl.:—ov. broadly oval, style stout, with 3 
small staminodes around it, stigmas 3; fruit rather large, 
globose, smooth, pericarp hard ; seeds numerous, immersed 
in solid white pulp, oval, compressed, smooth, not winged. 
—Sp. 2; both in FZ B. Lnd. 


Cc. Colocynthis, Schrad. in Linnea, xii. 414 (1838). Wake 
komadu, S. Peykkomaddi, 7. 

Moon Cat. 67. Thw. Enum. 126. C. P. 1607. 

Fl. B. Ind. ii. 620. Wight, Ic. t. 498. Bentl. and Trim. Med. Pl. 
EATEA 

Stems long, prostrate, slender, angular, scabrous or some- 
times hairy; |. 14-34 in., variable, usually deltoid in outline, 
very deeply trifid, the middle lobe the longest, each lobe 
deeply pinnatifid, with the segm. lobed or dentate, margin 
often involute, nearly glabrous above, very scabrous with 
short, swollen, prickly bristles beneath, petiole about half 
length of |., bristly or hairy; fl. rather large, peduncle about 
3 in., rough or hairy; cal. hairy, segm. linear-lanceolate ; cor.- 
segm. oval, subacute; ov. globular-ovoid, hairy, placentas 
very large, filling cavity, style short; fruit globular, slightly 
depressed, 2 in. diam., smooth, dark green mottled with 10 
lines of paler irregular spots, pericarp thin, completely filled 
with white spongy pulp divided by 6 spurious partitions ; 
seeds horizontal, scarcely + in., pale brown. 

Dry and desert regions ; ratherrare. Batticaloa (Gardner); Puttalam 
(Moon); Jaffna. Fl. January, Feb.; pale yellow. 

In India, Arabia, W. Asia, N. and Trop. Africa, and the Mediterranean. 
The plant is not cultivated in Ceylon, and the fruit above described is 
that of the wild plant, and differs much in size from the Colocynth 
obtained from the Mediterranean for pharmaceutical purposes. 


C. vulgaris, Schrad., the Water Melon (Komadu. S.), indigenous to 
Trop. Africa, is cultivated in the dry districts. There are figures of 3 
varieties in Duthie, Crops, tt. 47, 55, 56. 


8. BRYONTA, /. 


Perennial, tendrils bifid, fl. rather small, monoecious, male 
and fem. in small clusters, in same axils; cal.-limb cup-shaped, 
segm. 5; pet. 5, united for about } way up; male fl.:—stam. 3, 


254 Cucurbttacee. [Uukia. 


quite distinct, cells sigmoid ; fem. fl.:—ov. globose, glabrous, 
style slender, 3-fid.; fruit a globose berry, spuriously 3-celled ; 
seeds not numerous, each surrounded by a pulpy coat, slightly 
compressed.— Sp. 12; 1 in HZ B. Ind. 


B. laciniosa, Z. Sp. P/. 1013 (1753). 

Herm. Hort. Lugd.-Bat. 95. Burm. Thes. 50. Fl. Zeyl. n. 355. Moon 
Cat G72) dhwa enum 1265) Garba Koos 

BEB: Ind! i1,,622; Burm) ce7t: 97... Wight, lest. 500! 

Root large, tuberous, stems very slender, glabrous, often 
spotted with darker green, internodes very long; |. 3-5 in., 
ovate-rotundate in outline, very deeply cordate at base, cut 
nearly to base into 5 lanceolate or linear acute, coarsely ser- 
rate segm., the two basal ones deeply pedate, glabrous, thin, 
the upper surface slightly rough with minute scattered scales, 
petiole 1-14 in.; fl. in small clusters of 3-6, on short ped.; 
cal.-segm. linear, filiform ; cor.-segm. oval-oblong, acute, pu- 
bescent ; fruit globose, over # in. diam., smooth, bluish-green, 
with broad, white, vertical stripes ; seeds gibbous at the sides, 
with a prominent raised band running round the edge. 

Moist country up to 6000 ft.; common. FI. Aug.—Oct.; very pale 
yellow. 

Throughout the Eastern Tropics. 

There is no specimen in Hermann’s Herb. I have no Sinhalese 


name for this, Burman gives ‘ Basuagilli.’ Hermann l. c. says it is used 
as a cathartic. 


9. MUKIA, Arn. 

Perennial, tendrils simple, fl. small, moncecious, male and 
fem. together in axillary clusters; cal.-limb campanulate, with 
5 linear segm., pet. very slightly connate; male fl.:—stam. 3, 
distinct, anth.-cell straight ; fem. fl.:—ov. ovoid, style thick, 
surrounded at base by annular disk, placentas usually 2; 
fruit a globose berry, spuriously 2-celled; seeds ovoid, some- 
what compressed.—Sp. 2; both in FZ. B. Jnd. 


Cogniaux places this under J/elothria. 


Ibe scabrous beneath ; seeds tuberculated on sides. 1. M. SCABRELLA. 
L. softly hairy beneath ; seeds smooth . ‘ . 2. M. LEIOSPERMA. 


I. M. scabrella, Arz. zn Hook. Journ. Bot. iii. 276 (1841). HMin- 
kékiri, S|. Mochumochukkai, 7. 

FI. Zeyl. n. 356. Cucumis maderaspatanus, L., Sp. Pl. 1012. Thw. 
Enum 125-0 © eb lone: 

Fl. B. Ind. ii. 623. Wight, Ic. t. 501. 


Stems long, slender, much branched, angular, very hispid 
with spreading bristly hairs, young parts covered densely with 


Zehneria.| Cucurbttacee. 255 


white hair; |. variable in size, usually 3-4 in., but often only 
I in. or less, deltoid-ovate, very deeply cordate at base, with a 
wide sinus, and the rounded lobes often overlapping, acute or 
obtuse at apex, rather shallowly 5-lobed, coarsely dentate- 
serrate, usually scabrous with stiff hairs on both sides, petiole 
fully half as long as 1., cylindrical, very hispid ; fl. very small, 
male on pilose ped. as long as cal., fem. nearly sessile ; cal. 
hairy, segm. linear, pet. ovate, ciliate, a little longer than cal.- 
segm.; berry about 4in., broadly ovoid, apiculate, with a few 
scattered hairs, scarlet ; seeds horizontal, closely packed, oval, 
compressed, bluntly muriculate on the sides, enveloped in a 
pulpy envelope. 

Low country up to 3000 ft.; common. FI. all the year; yellow. 
Fruit scarlet. 

Tropics of the Old World generally. 

In the dry region this is often extremely harsh and hispid, but Her- 
mann’s original specimens have glabrous leaves, as described by 
Linnzus ; the scabrous-leaved specimens were calle | Bryonta cordifolia 
by him (see under Cephalandra). Leaves used as an expectorant. 


M. leiosperma, Wight, ex Roem. Synops. iii. 46 (1846). 

@hw. Enum. 125. C. P. 2700. 

FLB. Ind. 11. 623. 

Stems slender, hispid, young parts densely and softly 
villous-hairy; 1. 14-3 in., broadly cordate-ovate, acute, slightly 
3- or 5-lobed, strongly dentate, harshly hairy or scabrous 
above, densely and softly hairy beneath, petiole short, {-? in., 
hispid-hairy; fl. on slender hairy ped.; cal. very shaggy with 
long hair; pet. hairy outside; ov. thickly covered with very 
long hair; fruit under 4in., globose, very slightly hairy; seed 
rather larger than in JZ. scabrella, less compressed, smooth on 
the sides, conspicuously bordered. 

Montane zone, 4000-5000 ft.; very rare(?) Only found in Maturata. 
Fl, Aug. 

Also in S. India. 

I have not met with this living. It is very near JZ. scabred/a. 


10. ZEHNERIA, £7xd/. 


Annual or perennial, tendrils simple, fl. moncecious or 
dicecious, the male in racemes, the fem. solitary; cal.-limb 
campanulate, segm. 5, small; pet. 5, very slightly connate at 
base; male fl.:—stam. 3, inserted low down in cal., anth. dis- 
tinct, small, cells slightly curved; fem. fl.:—cal.-tube much 
constricted above ov. into a narrow neck, placentas 3, style 
surrounded at base by a lobed disk and 3 erect staminodes; 


256 Cucurbitacee. [Zehneria. 


fruit globose or ovoid, pulpy; seeds numerous, ovoid, smooth. 
—Sp. 15; 3 in #/. B. Ind. 


Cogniaux includes this under JZelothria. 


L. slightly cordate ; fl. moncecious ; fruit globular. 1. Z. HOOKERIANA. 
L. very deeply cordate at base; fl. dicecious ; fruit 
narrowly ovoid. : B : : 5 . 12), Zo HAS TAMAS 


1. Z. Hookeriana, 47x. ix Hook. Journ. Bot. iii. 275 (1841). 
Melothria perpusilla, Cogn. 1. c. 607. Thw. Enum. 125. C. P. 1613. 
Fl. B. Ind. ii. 624. Wight, Ic. t. 758 (Bxyonia maysorensis). 

Stems slender, cylindrical, glabrous, |. 12-24 in., broadly 
ovate, subcordate or nearly truncate at base, acuminate, acute 
at apex, distantly dentate, thin, slightly rough above with 
minute flat scales, paler and quite glabrous beneath, petiole 
#—I1in.; fl. moneecious, male 3-6, pedicellate, in short stalked 
racemes, fem. shortly stalked from same axils; cal.-segm. very 
short, setaceous; pet. ovate, acute, slightly hairy within; ov. 
globose, style 3-fid; fruit 2 in., globose, smooth, crimson ; 
seeds 30-40, ovoid, very much flattened, smooth. 

Moist region, 2000-4000 ft.; common. FI. Jan.; pale yellow. 


Also in India and Java. 
Cogniaux calls the Ceylon plant var. szbtruncata. 


2. Z. hastata, J/g. F/. [nd.-Bai. i. pt. 1, 656 (1855). Rawudu- 
kekiri, S. Peyppudal, 7. 

Bryonia umbellata, Moon Cat. 67. Z. umbellata, Thw. Enum. 125 ; 
ibe Sy, Cai 35, (Cz 12s 11, B05), 

Fl. B. Ind. 11. 625 (Z. umzbellata)., Rheede, Hort. Malab. viii. t. 26. 


Root with pendulous tubers, stems very long, slender, 
angular, glabrous, tendrils very long, young parts puberulous; 
l. 3-4 in., very variable, usually triangular- ovate, with a 
cordato-hastate base, the lobes often overlapping, but some- 
times prolonged into oblong or even linear divaricate lobes, 
acute, distantly denticulate, slightly rough with scales and 
bright green above, smooth and glaucous beneath, scabrous 
on margin, petiole $in., cylindrical, twisted; fl. dioecious, 
male on slender ped., numerous, crowded in a close, umbellate, 
pedunculate corymb much shorter than 1. fem. solitary; 
cal.-segm. minute; pet. very short, triangular; ov. glabrous, 
10-ribbed, style stout, stigmas very large; fruit about 1} in., 
oblong-ovoid, cylindrical, tapering to a point but not 
beaked, smooth, red; seeds ovoid, scarcely compressed, 
smooth, white. 

Moist low country; very common. FI. June, July; yellowish-white. 


Throughout Tropical Asia. 
Extremely variable in the shape of the leaves. Cogniaux distinguishes 


Rhynchocarpa.| Cucurbitacee. 257 


12 forms. In C. P. 3506 they are very deeply trifid with oblong-linear 
diverging segments. ; : 
The name ‘ Telaberiya,’ S., has also been given me for this. 


11. MELOTHRIA, Z. 


Tendrils simple, fl. moncecious, male and fem. often from 
same axils, cal.-limb campanulate, segm. 5, setaceous, pet. 5, 
very slightly connected; male fl.:—stam. 3, inserted in middle 
of cal.-limb, fil.short, anth. distinct, connective wide, produced 
into a point, cells nearly straight ; fem. fl..—ov. with 3 pla- 
centas, style rather long, surrounded at base with annular 
disk and 3 staminodes; fruit fusiform, beaked ; seeds numerous, 
small, much compressed, smooth.—Sp. 25; 5 in AZ. B. Jnd. 


M. zeylanica, Clarke in Fl. B. Ind. ii. 626 (1879). 

Bryonia deltoidea, Arn. Pug. 19. chmandra deltoidea, Arn. in Hook. 
Journ. Bot. i. 274. JW. deltoizdea, Thw. Enum. 124 (non Benth.). Cogn. 
Besse ee. E2501, 3128, 1610. 

Fl. B. Ind. ii. 626. 

Stems very slender, striate, glabrous; 1. 14-3 in., deltoid 
or triangular, with a wide, truncate and often hastate base, 
without other lobes, acuminate, distantly denticulate, more or 
less scabrous with white scales above, smooth or slightly hairy 
on veins beneath, petiole 4—# in., slender; fl. on long slender 
ped., 1-4 from axil; cal. very hairy at the throat, segm. 
recurved; pet. oval, puberulous, spreading; fem. fl.:—ov. fusi- 
form, the cal.-tube prolonged above it into a very narrow long 
neck; fruit 14-14 in., much tapering to both ends, cylindrical, 
with a sharp point or beak, and 6 slight vertical ridges, green; 
seeds horizontal, very much compressed, smooth, whitish. 

Moist region ; rare in the low country, common in the montane zone 


up to 7000 ft. Fl. Jan.-April; cream-coloured. 
Endemic. 


12, RHYNCHOCARPA, Schrad. 


Annual, tendrils simple, fl. moncecious, very small, male in 
slender racemes, fem. solitary; cal.-limb campanulate, segm. 5, 
subulate ; pet. 5, spreading, connate into a rotate cor.; male 
fil.:—stam. 3, inserted in cal.-limb, anth. distinct, connective 
slightly prolonged, cells straight; fem. fl.:—ov. ovoid, beaked, 
no disk, stigmas 2 or 3; fruit ovoid, strongly beaked; seeds 
few, smooth, narrowly winged.—Sp. 5; 1 in Fv. B. Ind. 


Cogniaux sinks this Genus under Kedrostis, Medik. 
PART II. 5 


258 Cucurbitacee. [Corallocarpus. 


R. rostrata, Vaud. in Ann. Sc. Nat. 4, xvi. 177 (1862). 

FI. B. Ind. il. 627 (A. fetida: not given for Ceylon). 

Stems prostrate, hairy; 1. small, 1-14 in., orbicular-ovate, 
cordate at base, acute, coarsely dentate, closely pubescent on 
both sides, petiole } in., hairy; male fl. very small, on long 
filiform ped., racemes corymbose, much shorter than 1.; fem. 
nearly sessile; cal. and pet. pubescent; fruit # in. or rather 
more, ovoid, tapering into a long narrow beak, red, densely 
covered with long white hair; seeds ovoid, gibbous on sides, 
with a narrow sharp wing, smooth, brown. 

Dry region; very rare. Collected in 1890 near the Giant’s Tank, 
Mannar District. FI]. Feb. 

Also in S, India. 


Combined with R&. fetida, Schrad., a common African plant, in 
Ik, 1B, Wael 


13. CORALLOCARPUS, Welw. 


Perennial, tendrils simple, fl. small, moncecious, male in 
small long-stalked corymbs, fem. solitary or two together ; 
cal.-limb campanulate, segm. very small; pet. 5, very slightly 
connected at base; male fl.:—stam. 3, inserted at mouth of 
cal.-limb, fil. very short, anth. distinct, cells straight, con- 
nective not produced ; fem. fl.:—ov. ovoid, usually 2-celled, 
style long, stigma large, 2-lobed ; fruit ovoid, beaked, succu- 
lent, circumscissile near the base; seeds few, ovoid, slightly 
compressed.—Sp. 15 (Cogniaux); 3 in FZ. B. Ind. 


CG. epigzeus, Clarke in Fl. B. Ind. ii. 628 (1879). Gopalanga, S. 

Atchmandra epigea, Arn., Thw. Enum. 125. C. P. 1622. 

Fl. B. Ind. ii. 628. Wight, Ic. t. 503. 

Root very large, turnip-shaped, stem rather succulent, 
prostrate, zigzag, glabrous, glaucous; |. rather small, 1-14 in., 
rotundate, very cordate at base, more or less 3- or 5-lobed, 
with obtuse or acute, irregularly dentate lobes, shortly and 
roughly pubescent on both surfaces, rather thick, petiole 
more than half length of 1, stout, glabrous; male fl. very 
small, on short ped., peduncle 14-24 in., straight, stiff, fem. 
larger; fruit #-1in., ovoid, with a blunt beak, smooth, scarlet 
in middle, with base and beak green, dehiscent by a circular 
line at junction of red and green portions near base; seeds 
about 9, in orange-coloured pulp, very slightly compressed, 
pale brown, adpressed-pilose. 


Dry country; rather common. Uma-oya; Dambulla; Mannar. FI. 
Jan.—March; yellowish-green. 
Also in dry parts of India. 


Ctenolepis.] Cucurbitacee. 259 


14. CERASIOCARFUM,* Hz. / 


Perennial, tendrils simple, fl. moncecious, male in small 
racemes, fem. solitary, often from same axil; cal.-limb cam- 
panulate, segm. 5, very small; pet. 5, slightly connate at base; 
male fl.:—stam. 3, inserted at mouth of cal., distant, anth. 
dorsifixed, cells straight, those of the 2-celled ones far apart, 
connective broad, not produced; fem. fl.:—ov. with 2 placentas 
and 4-6 ovules, style long, stigma 3-lobed, no staminodes ; 
fruit depressed-globose, pulpy; seeds 3-6, large, ovoid, with a 
narrow border, surrounded by orange pulp.—Monotypic. 


o Cc. zeylanicum, Clarke in Fl. B. Ind. ii. 628 (1879). [PLATE 
LIV. 

ae seylanica, Thw. Enum. 125. C. Bennettz, Cogn. |. c. 
7292. E 3002, 3500. 

Fl. B. Ind. 11. 628. 

Stems cylindrical, slender, glabrous; 1. 3-4 in., ovate- 
oblong, cordate at base, often hastate or with short spreading 
basal lobes, shortly acuminate, margin entire or coarsely 
sinuate, distantly denticulate, rough. above with few distant 
coarse prickly hairs, paler and glabrous beneath, thin, petiole 
about $in.; fl. on short slender ped.; cal.-segm. minute, 
setaceous; pet. ovate, acute or obtuse, pubescent; male fl. :— 
fil. extremely short, hairy; fem. fl.:—ov. globose, glabrous ; 
fruit 2 in. wide, depressed-globose, smooth, bright orange red; 
seeds } in. or more, ovoid, scarcely compressed, with a blunt 
crest at one end and a narrow blunt rim round the edge, 
finely pilose, brown or white. 

Lower montane zone; rather rare. Hantane; Medamahanuwara; 
Watagoda. Fl. November; pale yellow. 

Also in Java and Sumatra, and perhaps in S. India. 

The leaves are very like those of Zrichosanthes nervifolia. 


15. CTENOLEPIS, ZH. 7 


Annual, tendrils simple, fl. moncecious, minute, male in 
very short racemes, fem. solitary from the same axils; cal.- 
limb campanulate, short, segm. 5, subulate; pet. 5, connate 
below; male fl.:—stam. 3, anth. distinct, cells straight ; fem. 
fl.:—ov. globose, slightly depressed, style long, no disk; fruit 
small, depressed-top-shaped, 1- or 2-seeded; seed oblong- 
ovoid, compressed, bordered.—Sp. 2; both in FZ. B. Lund. 


* From the cherry-like fruit. 


260 Cucurbitacee. [Gynostemma. 


CG. Garcini,* Clarke in Fl. B. Ind. ii. 629 (1879). Mochu- 
mochukay, 7. 

Sicyos Garcint, Burm. f. Fl. Ind. 211. Zehneria Garcint, Stocks, Thw. 
Enum. 125. Blastania Garcint, Cogn. |. c. a2 (185 WO, 

Fl. B. Ind. 11.629. Burm. f. FI. Ind. t. 57, f. 3 (bad). 

Small, prostrate, stems slender, much branched, glabrous; 
]. rotundate in outline, small, 1-2 in., palmately cut more than 
3 down into 5 obovate or wedge-shaped obtuse lobes, strongly 
dentate-serrate, slightly scabrous-hairy on both sides, petiole 
4-1 in., bearing in its axil a stipule-like, large, cordate or 
reniform bract fringed with very long filiform cilia; fl. ex- 
tremely small, male racemes very short, concealed by the 
bract, pubescent; pet. oblong, reflexed; fruit 3-3 in., wide, 
flat-topped, smooth, orange; seeds slightly cnened. 

Dry country; rather rare. Jaffna; Matalan, E. Prov. (Nevill); 
Tissamaharama, S. Prov. Fl. Dec.—February; white. 

Also in Southern India. 

The original locality where Garcin collected this seems to have been 
Tuticorin, not Ceylon, though ‘Zeylona’ is the locality given. Burm. f. 
mistook the pectinate bracts for fruit. 

Used by the Tamils as a medicine in quinsy and other affections of 
the throat. 


16. GYNOSTEMMA, 2. 


Perennial, tendrils bifid, 1. compound, fl. dicecious, very 
small, in lax panicles; cal.-segm. 5, small; pet. 5, very 
acuminate; male fl.:—stam. 5, fil. connate into a column, 
anth. quite distinct, cells straight; fem. fl.:—ov. globose, 
3-celled, with 2 pendulous ovules in each cell, styles 3, 
spreading, bifid; fruit a small berry, 1-3-seeded ; seed large, 
worted, furrowed round edge, not compressed.—Sp. 5 (Cog- 
niaux); 1 in FZ. B. Ind. 


G. laxa, Cogn. Monog. Phan. iii. 914 (1881). [PLATE XLV.] 
Zanonia Wishtiana, Arn, Pug. 386. Pestalozzia laxa, Thw. Enum. 
iw, G, pedata, Trim. Syst. Cat. Cob IIb ss (@lIw, (Gs IF, e532. 

Fl. B. Ind. 11. 633 (G. fedata). 

Stems very long, sometimes slightly woody below, slender, 
cylindrical, glabrous and shining ; 1. 3-foliolate, petiole 1-14 in, 
stout, furrowed above, often twisted and sub-prehensile, 
glabrous, Iflts. 2-4 in. shortly stalked, lanceolate, acute, 
coarsely serrate, glabrous, rather thick, the middle one rather 
the largest, the lat. ones either dilated on lower side or 


* Laurent Garcin collected in S. India at the beginning of the 18th 
century, and his plants are in Burman’s Herbarium. He may have also 
yisited the North of Ceylon. Died 1752. 


Zanonia. Cucurbitacee. 261 


divided pedately into another Iflt.; fl. very small (male a little 
larger), very numerous, on slender ped., arranged in peduncled, 
elongated, pyramidal, lax, spreading, pubescent panicles, in 
the fem. about as long as |., in the male often very much 
longer ; cal.-segm. ovate, acute; pet. linear-lanceolate, much 
attenuate-acuminate, spreading; fruit about 41in., globose, 
smooth, dull black; seeds 1-3, in dark green pulp, rather 
large, }-+in., globose-trigonous, slightly compressed, furrowed 
on edge, wrinkled and worted on sides, dark chestnut-brown. 

Montane zone, 3000-6000 ft.; rathercommon. Abundant at Hakgala. 
Fl. Feb., Oct.; yellowish-green. 

Also in Assam, Khasia, Sumatra, Java. 

Clarke combines this with G. fedata, Bl., but Cogniaux distinguishes 
it by its narrower pet. and trifoliolate ].; but the latter are occasionally 
pedate in our plant. 


17. ZANONTA, Z. 


Perennial, semi-woody, tendrils simple, fl. dicecious, male 
in panicles, fem. in racemes; cal.-segm. 3, valvate (no cal.- 
limb); pet. 5, connate below to form a rotate cor., imbricate; 
male fl.:—stam. 5, inserted on central disk, distinct, anth. 
I-celled, opening transversely; fem. fl.:—staminodes 5, short, 
ov. narrowly turbinate, cylindrical, at first 3-celled (afterwards 
1-celled), with 2 pendulous ovules on each cell, styles 3, 
spreading, bifid; fruit a truncate capsule, opening on summit 
by 3 valves; seeds very few (often 6), large, ovoid, much 
compressed, surrounded by a very wide wing much prolonged 
at each end, embryo large, with flat fleshy cotyledons.—Sp. 2; 
1in F/. B. Lnd. 


Z. indica, Z. S. P/. ii. 1457 (1768). Wal-rasakinda, S. 

Moon Cat. 68. Thw. Enum. 124, 442. C. P. 1628. 

Fl. B. Ind. ii. 634. Wight, Ill. t. 103. 

Stems stout, cylindrical, semi-woody, grey, glabrous ; 
l. large, 3-6 in., deciduous, leaving a very prominent circular 
scar, broadly oval to lanceolate, rounded or cordate at base, 
somewhat acuminate, apiculate, quite entire, glabrous, rather 
thick, reticulate venation rather conspicuous beneath; fl. 
rather small (fem. much the larger), on short ped., male 
panicles 6-12 in. long, branched chiefly at base, with fl. in 
small clusters, fem. racemes longer, with fl. solitary, distant; 
cal.-segm. rotundate, concave, glabrous; pet. ovate, acuminate, 
obtuse, with incurved points; male fl.:—fil. short, broad, 
spreading ; fem. fl.:—ov. 4 in., glabrous, styles rather long ; 


262 Begoniacee. [ Begonia. 


capsule 1-1} in. or more, cylindrical, rounded at base, truncate 
at apex, glabrous, pale yellowish-brown ; seed with wing as 
long as fruit, very flat, glabrous, yellow, wing rounded at the 
ends. 


Moist low country up to 2000 ft.; ratherrare. Kaduganawa ; Kurune- 
gala; Ambagamuwa. Moon’s locality is Kalutara. Fl. June; greenish- 
yellow. 

Also in India and Malaya. ee 

Has much the aspect of a Mendspermacea, as its S. name indicates. 


LIX.—BEGONIACEZ. 


PERENNIAL succulent herbs, 1. alt., with stip., fl. unisexual, 
moncecious ; male fl.:—perianth-segm. 2 or 4, in pairs, outer 
pair valvate, inner pair (when present) smaller; stam. indef. 
(30-50), usually somewhat monadelphous; fem. fl.:—peri- 
anth-segm. 2-5; ov. quite inferior, 3-celled with very 
numerous ovules crowded on axile placentas, styles 3, distinct 
or slightly connate; fruit a triangular papery capsule, winged 
at the angles, opening by an elliptical slit on each face ; seeds 
minute, very numerous, with scanty endosperm. 


This curious Family is very poorly represented in Ceylon. Of our 
species, 2 are confined to the moist low country and 2 to the montane 
zone, whilst one occurs in both. 


BEGONTIA, Z. 
For characters, see Order.—Sp. 398 ; 64 in FZ. B. Ind. 


Stemless. 
Placentas 2-lobed. F : ‘ : : . I. B. CORDIFOLIA. 
Placentas undivided. 
L. rounded or subacute at apex : 5 . 2. B. TENERA. 


L. acuminate. 
Scapes and ov. very hairy : 5 . 6. Be Dewan sie 
Scapes and ov. nearly glabrous . : . 4. B. SUBPELTATA. 
Stem stout, erect A : : ; 5. B. MALABARICA. 


1. B. cordifolia, 7iw. Enum. 129 (1859). Gal-ambala, 5. 

C. P. 3584. 

Fl. B. Ind. i. 641. Wight, Ic. t. 1816 (Dzploclinium cordifolium). 

A perennial herb with a short horizontal rootstock, sending 
up leaves and flower-stalks; 1. erect or spreading, petiole 
3-7 in., stout, succulent, with spreading hair, pink, blade 


Begonia.] Begoniacee. 263 


3-6 in., ovate-rotundate, strongly cordate with the lobes nearly 
equal at base, acute or obtuse at apex, very slightly lobed, 
crenate-dentate, unequal-sided, thin, sparingly hairy on both 
sides, bright light green, stip. ovate-lanceolate, acuminate, 
glabrous ; scapes 4-6 in., erect, succulent, hairy, pink, slightly 
bifurcate at top; fl. 3-14 in., male and fem. usually on sepa- 
rate scapes, ped. of male fl. $-1in.,, filiform, of fem. fl. shorter 
and stouter, bracts small, ovate, acute; male fl.:—per.-segm. 
4, the outer pair rotundate, cordate at base, inner narrowly 
oblong, stam. about 30, fil. monadelphous below; fem. fl.:— 
per.-segm. 3-5 (usually 5), the 2 outer rotundate, the 3 inner 
oval-oblong, ov. with bifid placentas, shortly 3-winged, 
glabrous, style deeply divided into 3 stout curved branches, 
each bilobed at end ; capsule very thin and papery, dehiscent 
on the three sides, wings nearly equal, variable in width, 
upper margin rounded; seeds broadly ovoid, ridged, dark 
brown. 

Low country to about 1500 ft.; very rare. In plenty in crevices of 
wet rocks near Lunugala, Bibile, and Nilgala, Uva Prov. Fl. Jan— 
March; white. 


Also in S. India. 
A. DC. (Prod. xv. pt. I, 329) calls the Ceylon plant var. zzsu/aris. 


2. B. tenera, Dryand. in Trans. Linn. Soc. i. 169 (1791). 

B. rupestris, Moon Cat. 64(?). 8. Thwatteszz, Hk., Thw. Enum. 128 
(part). C. P. 3952, 2808 (part). 

Fl. B. Ind. ii. 652. Trans. Linn. Soc. i. t. 16. 

Stemless, with a stout prostrate rhizome; 1, 24—4 in., rotun- 
date, very strongly cordate and slightly unequal at base, 
rounded or subacute at apex, faintly serrate-crenate, hairy on 
both sides, thin, petiole 3-6 in., hairy, succulent, stip. ovate, 
acuminate, somewhat fleshy; scapes 3-9 in., erect, stiff, hairy ; 
fi, few, umbellately corymbose, on slender ped., the male erect, 
the fem. drooping, bracts ovate, acute; male fl.:—per.-segm. 
4, 2 outer rotundate, cordate, hairy outside, 2 inner oval- 
oblong, stam. 40-50, fil. connate at base; fem. fl.:—2 outer 
per.-segm. broad, 3 inner narrow, ov. very hairy, placentas 
undivided, styles 3, distinct almost to base, stigmas bilobed ; 
capsule 2 in. by 2 in. wide (with wings), glabrous, wings nearly 
equal, their upper margins nearly straight, forming a sharp 
angle (or slightly rounded) with sides. 

Moist low country; rare. First collected by Koenig, but he does not 
give any locality. Near Bulat-Singhale, Pasdun Korale (Thwaites); 
Wellawaya (F. Lewis). Fl. Sept.; white. 

Endemic. 

Dryander gives in full Koenig’s description from his MSS.; and the 
figure is taken from his specimen in the Mus. Brit. 


264 Begoniacee. [ Begonia. 


3. B. Thwaitesii, Hook. Bot. Mag. 4692 (1853). 

keichenheimia Thwaitesiz, Klotzsch, Begon. 55. Thw. Enum. 128 
(part). C. P. 3953, 2808 (part). 

Fl. B. Ind. 11. 652 (under &. zenera). Bot. Mag. t. 4692 ; Klotzsch, 1. c. 
La. 

Differs from B. zenera in having |. much more unequal at 
base, acuminate and more acute at apex, more or less mottled 
with purple above and with the veins dark purple instead of 
white; petioles and scapes much more hairy, even shaggy, 
with red hair; capsules longer, $—3 in. 

Lower montane zone, 3000-4000 ft.; rare. Kotmale (Gardner) ; 
Matale ; Hunasgiriya. Fl]. Sept._January; pale cream-coloured. 

Endemic. 

The leaves are usually beautifully mottled and variegated with 
coppery-red, purple and green above, and entirely purple beneath. 


4. B. subpeltata, Wight, lc. v. 2,9 (1852). 
Thw. Enum. 128. C. P. 2597. 
INL 1B, Ike moss, Wiel Met Insie 


Differs from B. zenera in having |. more acuminate, some- 
what lobed and more finely hairy ; scapes longer and nearly 
glabrous ; flowers larger; ov. quite glabrous. 

Lower montane zone; very rare. Ambagamuwa (Thwaites). FI. 
March ; white. 

Also in S. India. 

The leaves are not at all subpeltate in Ceylon specimens. 

The last two species and this are all very closely allied, and are 
probably only varieties of one. C. P. 346 (see Thw. Enum. 129) from 
Hiniduma (without flowers), with leaves covered with dense red 
pubescence, is probably another species or variety of the same series, 
but it has a larger rootstock and the leaves are less unequal at the base. 


5. B. malabarica, Lam. Encyc/. i. 393 (1783). Hakambala, S. 

Moon Cat. 64. &. adifetala, Graham in Bot. Mag. t. 2849. Thw. 
Enum. 128. C. P. 2807, 3949. 

Fl.-B. Ind. n. 655. Bot. Mag. t. 2849. Wight, Ic. t. 1813: 


Stem erect, 2-3 ft., cylindrical, stout, zigzag, thickened at 
the nodes, glabrous, branched ; 1. 4-6 in., numerous, extremely 
unequal-sided, the large side deeply 4-cordate at base, the 
other simply rounded, very acuminate, acute, sharply spinous- 
serrate and sometirnes coarsely incised, slightly scabrous 
above or glabrous, paler or purplish, shining and glabrous 
beneath, petiole succulent, cylindrical, stip. over 4 in., lanceo- 
late, acute, membranous, deciduous; fl. rather large, on long, 
slender, glabrous, drooping ped.; per.-segm. 2, orbicular, much 
broader than long; male fl.:—stam. above 4o, fil. connate at 
very base ; fem. fl.:—ov. quite glabrous, placentas undivided, 
styles 3, dilated and undulate at ends; fruit very light and 


Tetrameles.] Datiscacee. 265 


papery, pendulous, near I in., pale brown, shining, wings 
slightly unequal, broad, with upper margin either straight, or 
completely rounded off without an angle. 

Moist region, up to 5000 ft.; rather common in wet rocky places, 
especially above 3000ft. Ritigala, N.C. Prov. (Bell). Fl. all the year ; 
rose-pink or nearly white. 

Also in Malabar. 

Variable, but not divisible into distinct varieties. I find the angled 
and rounded wings in fruits on the same plant. C. P. 3949 (from Matale) 
has narrow rounded wings and deeply gashed leaves. 


LX.—DATISCACE/:. 


TREE, |. alt., simple, without stip., fl. small, unisexual, dice- 
cious in panicles, apetalous; male fl.:—cal. deeply cut into 4 
segm.; pet. 0; stam. 4, opp. cal.-segm., inserted round a flat 
central disk; fem. fl.:—cal.-tube connate with ov., segm. 4, 
short; pet. 0; ov. quite inferior, 1-celled, with numerous 
ovules on 4 parietal placentas, styles 4, distinct, persistent ; 
capsule small, ovoid, opening at summit; seeds minute, 
flattened, with a very loose testa, with endosperm. 
There are only 2 other genera in this small Order. 


TETRAMELES, 3,. 


For characters, see Order.—Monotypic. 


T. nudiflora, 2. Zr. 7x Lenn. Pi. Jav. Rar. 79 (1838). Nigunu, 
Mugunu, 5S. 

aw. enum.252., 'C. Pian: 

Fl. B. Ind. ii. 657. Wight, Ic. t. 1956 (7. Grahamiana). Bedd. Fi. 
Sylv. t. 212. 

A very large tree, with a straight trunk, unbranched below 
and buttressed at the base, bark smooth, grey, fibrous, twigs 
marked with scars of past leaves, young parts puberulous; 
l, alt. large, 6-7 in., ovate, strongly cordate at base, acumi- 
nate, acute, irregularly dentate, in upper part, bright green and 
glabrous above, paler and with prominent, yellow, softly 
hairy, reticulate venation beneath, petiole 4-6 in., cylindrical, 
pubescent ; fl. numerous, panicles many, from the base of the 
new growth below the young |., 6-10 in., stalked, slender, 
hairy ; male with spreading branches and fl. on slender ped., 


266 Cactacea. [Rhipsalés. 


fem. with erect branches and sessile fl.; male fl.:—cal.-segm. 
linear-oblong, glabrous, stam. spreading, fil. much longer than 
cal.; fem. fl.:—ov. faintly 8-ribbed, slightly pubescent, styles 
much exserted, stigmas club-shaped ; fruit a little over jin, 
urceolate, truncate, 8-ribbed, slightly rough. 

Forests in low country to 3000 ft.; rare. Ambagamuwa and 
Maskeliya (Thwaites); Maha-oya, Batticaloa Dist. (A. Walker); Kegalla; 
Lagalla Dist.; Allagalla. FI. March; September; yellowish-green. 

Also in W. India, Sikkim, Burma, Andamans, Java. 

Attains 14 ft. in circumference of trunk, and is completely deciduous 
and quite bare when in flower. Wood very soft and light, pale yellow. 


IC AVE TAC IB 45, 


SUCCULENT, leafless perennial, fl. bisexual, regular ; cal.-tube 
adnate to ov., segm. 4 or 5; pet. 5 or 6; stam. about 20, free, 
inserted at mouth of cal.; ov. quite inferior, 1-celled (incom- 
pletely 3- or 4-celled), with numerous ovules on parietal 
placentas, style simple, stigma 3- or 4-lobed ; fruit a succulent 
berry ; seeds few, oblong, without endosperm. 


A large Order confined to the New World with the exception of a few 
species of Rizpsalzs. R. Cassytha alone reaches Asia. 


RHIPSALIS, Gaerin. 


For characters, see Order.—Species about 40; 1 in Fé. 
B. Ind. 

R. Cassytha,* Gaertn. Fruct.i. 137 (1788). Wal-nawahandi, S. 

Cactus pendulus, Willd., Moon Cat. 38. Thw. Enum. 129. C. P.2976. 

Fl. B. Ind. 11. 658. Bot. Mag. t. 3080. 

Perennial, slightly woody at very base, stems fleshy, 
slender, 4-6 ft. long, pendulous, cylindrical, much dichoto- 
mously (or 3- or more) branched, very flexible, glabrous, dull 
green, |. quite absent but represented on the young shoots by 
very minute, distant, apiculate scales; fl. small, sessile, soli- 
tary ; cal.-segm. unequal, broad, acute; pet. oblong, obtuse, 
spreading ; ov. glabrous, style longer than stam.; berry small, 
4in., ovoid, white, smooth, translucent, at first crowned with 
persistent cal. and pet.; seeds 8-16, in watery pulp, very 
small, oblong-ovoid, faintly striate. 


_ * From its resemblance to that genus of Lauracee, to which, indeed, 
Aiton and Miller had referred it. 


Sesuvium.| Ficoidee. 26 7 


Epiphytic on the trunks of trees, also on rocks, in the moist region up 
to 4000 ft.; common. FI. Aug., Sept.; greenish-white. 

Also in Mauritius, Madagascar, Trop. Africa, W. Indies, and Trop. 
America ; but nowhere else in Asia. It is, however, without doubt a 
native in Ceylon. 

Sometimes called ‘ Mistletoe’ by the English in Ceylon, from a slight 
resemblance in general appearance and berries to Viscum album of 
Europe; but it is not truly parasitic, having very long fibrous roots, 
which spread over the surface of the wood or rock on which the plant 
grows. When quite young, the plant is set with stellate tufts of bristles. 


Opuntia Dillenit, Haw., is the common ‘Cactus’ met with on waste 
ground and by roadsides in the low country, rarely in the moist region, 
but in great abundance near the coast in the dry; called ‘ Katu-patuk’ by 
the Sinhalese and ‘ Naka-kalli’ by the Tamils. It is figured in Wight, 
Ill. t. 114, and is an old introduction, now quite naturalised, and indeed a 
great nuisance in parts of the Northern Prov. The plant is S. Americar 
in origin. No other species is naturalised here. 


LXII.—FICOIDE/E. 


ANNUAL or perennial herbs, |. simple, opp. or alt., without 
stip., fl. small, regular, bisexual, usually apetalous, sep. 5, 
distinct or more or less connate (in Seswvzum slightly adnate 
to ov.), pet.o (rarely 5); stam. def. or indef., perigynous or 
hypogynous ; ov. superior or nearly so, 2-5-celled, with few 
or numerous ovules in each cell (in Gzsekza of 5 separate carp. 
each with 1 ovule); fruit a thin capsule, transversely or locu- 
licidally dehiscent ‘in Gzsekia separate cocci); seed more or 
less reniform, embryo curved round the farinaceous endo- 


sperm. 


A somewhat ill-defined Order. The 2 first genera are allied to Portu- 
lacacee ; Mollugo to Caryophyllacee ; and Gisekia to Phytolaccacee. 


Stam. inserted on cal.-tube (Azzozdec). 
Styles3 . ; : : 3 A , : . I. SESUVIUM. 
Stylestor2 . ; ; ; 2. TRIANTHEMA. 
Starn. hypogynous (Molluginez) 
Fruit a loculicidal capsule , Z ; : . 3. MOLLUGO. 
Fruit of separate carpels . ; ; : ; 2 4, GISEKIA. 


Eleven species, all found in low country only, and usually weeds. 
Sesuvium is a seashore plant. 


1. SESUVIUM, Z. 


A fleshy prostrate herb, 1. opp., without stip., fl. solitary, 
axillary; cal.-tube very short, adnate to base of ov., segm. 5, 


268 Ficoidee. [Zrianthema. 


coloured, persistent; pet. 0; stam. indef. (about 30), inserted 
on cal.-tube; ov. nearly superior, 3-celled, with several ovules 
in each cell, styles 3, distinct; fruit a membranous, 3-celled 
capsule, circumscissile near the base; seeds few, reniform, 
smooth, embryo curved into a ring.—Sp. 4; 1 in Fd. B. Ind. 


S. Portulacastrum, Z. Syst. Veg. ed. 10 (1759). Wankiru- 
valai, 7. 

S. pedunculatum, Pers., Moon Cat. 39. S. repens, Rottl., Thw. Enum. 
az, (C, IP, 106%. 

Fl. B. Ind. ii. 659 (not given for Ceylon). Hk. Comp. Bot. Mag. i. t. 
23 (S. repens). 

Herbaceous, stems prostrate or buried in the sand, 
copiously rooting at the nodes, stout, glabrous, succulent, 
much branched; 1. numerous, opp., I-14 in., spathulate- 
oblong or linear, very fleshy, tapering to base, obtuse, 
glabrous, petiole short, much dilated and amplexicaul at base 
with broad scarious margins (adnate stip. ?); fl. nearly 4 in., 
solitary, ped. as long as cal., glabrous; cal. glabrous, segm. 
lanceolate, cuspidate; stam. shorter than cal.; capsule shorter 
than and enclosed in persistent cal.-segm., pointed; seeds 
smooth and polished, black. 

Sandy seashores; rather rare. Trincomalie (Gardner); Mannar; 
Karativu; Colombo (Moon); Chilaw; Tangalle. Fl. Jan. Feb.; pink. 

Most Tropical and Sub-tropical coasts. Moon gives the Sinhalese name 
as ‘Maha-sarana.’ The leaves are eaten in curries. 


The figure quoted (of the Carnatic plant) shows much shorter leaves, 
and is generally smaller than our plant. 


2. TRIANTHEMDIA, J. 


Herbs with prostrate branches, 1. opp. or alt., without stip. 
but petioles dilated at base, fl. small, axillary; cal.-tube long 
or short, not connate with ov., segm. 5; pet. 0; stam. 5, 10, 
or 20 inserted at top of cal.-tube; ov. quite superior, I or 2- 
celled, with few basal ovules, style 1 or 2; fruit a capsule, 
membranous below, with a hard thick cap which is detached 
by a transverse dehiscence and carries away with it one or 
more seeds; seeds few, reniform, embryo curved into a ring. 


—Sp. 10; 5 in FZ. B. Ind. 


Style 1. 
L. obovate; capsule about 8-seeded : : . I. T. MONOGYNA. 
L. linear-lanceolate; capsule 2-seeded . : . 2. T. TRIQURTEAS 


Styles 2; capsule 4-seeded. : : : : . 3. T. DECANDRA. 


Trianthema.] Ficoidee. 2 69 


I. T. monogyna, LZ. Mazi. 69 (1767). Hin-sarana, S. 
T. obcordata, Roxb., Thw. Enum. 23. C. P. r1o2. 
Fl. B. Ind. ii. 660. Wight, Ic. t. 288 (Z. obcordata). 


A prostrate, somewhat succulent herb, stems rather 
angular, glabrous or slightly pubescent, much branched; 1. 
obliquely opp., very unequal, the upper one larger, 2-1 in, 
the lower smaller, +-} in., obovate, tapering to base, rounded, 
often apiculate at apex, petiole 4 in., connate, very much 
dilated and membranous at base, especially of the smaller 1. 
which forms a deep triangular axillary pouch containing the 
solitary sessile fl.; cal.-segm. ovate, acute; stam. 10-20; ov. 
truncate, style simple; capsule small, almost concealed in the 
stipular pouch, about 8-seeded, cap exserted, truncate, carrying 
away with it 3 seeds; seeds reniform, black, dull, muriculate. 

Low country, especially on the coast and in the dry region near tanks. 


Colombo; Kantalai; Anuradhapura. 
Throughout Tropical regions. 


2. T. triquetra, Rotil. and Willd. in Neue Schrift. Natur. Fr. 
Berlin, iv. 181 (1803). Chiruppadikkirai, 7. 
Fl. B. Ind. ii. 660 (7. crystallina). 


A diffuse herb with a woody rootstock, branches numerous, 
prostrate, divaricate, covered with minute papille; |. opp. 
or alt., 4-2 in., linear or linear-lanceolate, obtuse, fleshy, 
petiole short, sheathing; fl. sessile, numerous, several in an 
axil, not enclosed in sheath of petiole; cal.-tube thick, with 
numerous slender ribs, segm. broad, triangular, acute, per- 
sistent, folded over top of ripe capsule; stam. 5; style simple; 
fruit enclosed in cal.-tube, 2-seeded, cap flat but much de- 
pressed in centre, carrying away with it one seed; seeds rather 
large, strongly striate, dark brown. 


Var. 8, Rottleri, 777m. T. crystallina, Thw. Enum. 23; Trim. 
Syst. Cat. 39 (non Vahl). C. P. 3552. 


Less woody at base or quite herbaceous, branches more 
slender, often tufted and congested; 1. smaller, {-3in., less 
fleshy; fl. smaller, I or 2 in an axil; cal.-segm. more shortly 
triangular, less acute, not folded over top of ripe capsule. 


Dry region; rather rare. The type only at Mantai, N. Prov. Var. 8 
more common; Trincomalie; Gunners’ Quoin; Batticaloa; Tissamaha- 
rama; Mannar. FI. Nov.—Feb. 

Also in the dry regions of Afghanistan, Arabia, and India. 

I was inclined to consider these as distinct species, but Dr. Stapf, of 
Kew, informs me that in that Herbarium there are connecting forms. 
TI. crystallina, Vahl, to which this is referred by Thwaites and in FI. B. 
Ind., is a quite different species from Arabia and Nubia only. 

Eaten in curries at Mannar. 


270 Ficowdec. [Mollugo. 


3. T. decandra, LZ. Mant. 70 (1767). Maha-sarana, 5S, 
Charanai, 7. 

Wen, Jair, 23 (CIPS iO ik, 

Fl. B. Ind. 11. 661. Wight, Ic. t. 296. 

Stems long, prostrate, slightly branched, angular, glabrous, 
with long internodes; |. opp., pair rather unequal, ?-14 in., 
oblong-oval, rounded and apiculate at apex, slightly fleshy, 
petiole }-4 in., much dilated and amplexicaul at base; fl. on 
short ped., in dense umbellate, shortly stalked,. axillary 
clusters, bracts scarious, acute; cal.-tube short, segm. much 
longer, oval, acute, with membranous margins; stam. 10; 
styles 2; capsule not included in cal.-tube, 4-seeded, cap large, 
solid, very truncate, carrying away with it 2 seeds; seeds 
finely wrinkled, black. 

Low country, chiefly in the dry region; rather rare. Batticaloa and 
Jaffna (Gardner); Trincomalie (Glenie); Colombo (Thwaites); Tissama- 
harama; Mantai. FI. Dec.; bright pink. 

Also in India, Burma, and Timor. 

The flowers are very pretty when fully expanded. 


3. MOLLUGO, Z. 


Small, often annual, herbs, |. apparently whorled or radical, © 
without stip., fl. small, axillary or in terminal cymes; sep. 5, 
distinct; pet. 0 or 5; stam. 3, 5, or 10, hypogynous; ov. 
superior, 3- or 5-celled with numerous ovules, styles 3 or 5 ; 
capsule membranous, enclosed in persistent sep., 3- or 5-celled, 
loculicidally dehiscent; seeds reniform, embryo more or less 
curved.—Sp. 12; 6 in FZ. B. Ind. 


Fl. axillary; seeds with a filiform anpeneee. 
. HIRTA. 


Densely stellate-hairy : 3 on alse VE 
Glabrous . M. OPPOSITIFOLIA. 
Fl. in terminal cymes; seeds not appendaged. 
L. in whorls at nodes. 
Seeds with raised tubercles . 3. M. PENTAPHYLLA. 
Seeds smooth or nearly so. 
Stems numerous, glabrous. 4. M. CERVIANA. 
Stem solitary, glandular pubescent 5. MM. DISMICEESas 
L. all radical E ; : 6. M. NUDICAULIS. 


1. M. hirta, 7hunb. Fl. Cap. 120 (1818). 

Glinus dictamnoides, L., Thw. Enum. 24. C. P. 293 

Fl. B. Ind. t. 662. Burm. f, FI. Ind. t. Bos 1 1 (Ga Totoides). 

Annual, stems numerous, prostrate, spreading from a 
centre, 6-12 in., not much branched, densely stellate-hairy, 
|. opp. or several at a node, unequal, largest 4—? in., rotundate 
or broadly oval, tapering to base, very obtuse at apex, stellate- 


Mollugo.] Ficoidee. 271 


downy on both sides, petiole 4in., slender, stellate-hairy ; fl. 
axillary, 1-6 at a node, ped. unequal, }-2 in., stiff, divaricate ; 
sep. + in., lanceolate, acute, stellate-hairy outside; pet. filiform, 
about as long as sep., deeply bifid; stam. 10; ov. ovoid, gla- 
brous, 5-lobed, 5-celled, with very numerous ovules, stigmas 
5, sessile, spreading ; seeds ovoid, pitted, dark brown, shining, 
with a small white scale at the hilum extended into a long 
filiform process which curves over the top of the seed. 


Dry region; rather common. Bintenne; Anemaduwa, N.W. Prov. 
Fi. July, August; pinkish-white. _ 
In all Tropical and warm countries. 


2. M. oppositifolia, Z. S~. Pl. 89 (1753). Hin-pala, 5S. 
Kachchantiray, 7. 

Herm. Mus. 4. Burm. Thes. 13. Fl. Zeyl. n. 52. Moon Cat. 9. 
Pharnaceum Mollugo, ., Moon Cat. 23. MW. Spergula, L., Thw. Enum. 
Meee bemasmyst. Cat. 39. C. P. 1095. 

Fl. B. Ind. ii. 662 (17. Sfergula). Burm. Thes. t. 7. 


Annual or perennial, stems numerous, prostrate, much 
branched, glabrous, with long internodes; |. in whorls of 
4 or 5, unequal, 2 usually larger than the others, reaching 
?in., from linear to oval, often somewhat spathulate, much 
tapering to base, rounded or acute, apiculate at apex, petiole 
obscure; fl. axillary, 1-5 at a node, ped. +-2in., very slender; 
sep. oblong, subacute, with membranous margins; pet. gene- 
rally 0; stam. 3; capsule surrounded by persistent sep., 
oblong-ovoid, 3-celled; seeds very numerous, tuberculate, 
brown, with a similar appendage to WW. /urta. 


Low country, especially in the dry region; a common weed. FI. 
all the year; white. 

Also in India, Trop. Africa, and Australia. 

Varies considerably in habit and in width of leaves. Linnzeus’s name, 
M. Spergula, has been usually applied to this, but is later by six years 
than his 17. ofpositifolia. 

Wight and Arnott state (Prod. 44) that all the Indian specimens have 
5-forked pet. and 5 or 10 stam. 

Used as a pot-herb. ‘Jean-pala’ is Hermann’s name for this. 


3. M. pentaphylia, LZ. Sf. P/. 89 (1753). 

Herm. Mus. 33. Surm. Thes. 12 and 13. FI. Zeyl. n. 51. Moon 
Cat.9. Thw. Enum. 24. M. stricta, L., Thw. Enum. 24; Trim. Syst. 
Cat. 39. C. P. 1096, 1097. 

Fl. &. Ind. ii. 663 (47. stricta). Burm. Thes. t. 6, f. 2 and t. 8, upper 
figure. 

Annual, with many spreading, diffuse, dichotomously 
branched, slender, glabrous, quadrangular stems; 1 2-6 in 
a whorl, 4-1 in., from lanceolate-linear to obovate, much 


tapering to base, apiculate, petiole obscure; fl. numerous, on 


272 Ficoidee. [Mollugo. 


filiform ped., arranged in lax, corymbose, terminal cymes ; 
sep. rotundate-oval, obtuse; capsule globose, walls very thin, 
seeds numerous, closely covered with minute raised tubercles. 
Low country, especially in the dry region; common. FI. June, July; 
white. 

Throughout India, Malaya, China, Japan, &c. 

M. pentaphyllais the older by nine years of Linnzus’s two names, and 
is therefore here adopted. JZ. stricta has narrower leaves, not at all 
obovate, and is kept up as separate by Thw. 

Hermann erroneously gives the S. name ‘ Wisnagarandi’ (=‘ Vis- 
nukiranti’ which belongs to Zvo/vulus) for this. 


4. M. Cerviana, Ser. ix DC. Prod. i. 392 (1824). Patpada-= 
kam, 7. 


Pharnaceum triflorum, Moon Cat. 23. Thw. Enum. 24. C. P. 1092. 
Fl-B. Ind. i. 663. Lam: Ill, t. 2145 f. 1. Pluk, Phyt) eege2enee 


(good). 

Annual, with very numerous, erect, very slender stems 
5-8 in. high, SRAC LES umbellate, nodes thickened; |. 2-6 or 
more in a whorl, 4-%in., linear, very narrow, apiculate; fl. 
numerous, on long, stiff filiform ped., usually in 3’s at end of 
long filiform axillary and terminal peduncles; sep. oval- 
oblong,obtuse, with membranous margins; seeds quite smooth, 
red-brown. 

Dry region; rathercommon. Jaffna and Kalpitiya (Gardner). 


Also in dry districts of India and rest of Asia, Africa, and Australia. 
Much used as a medicine in fever. 


5. M. disticha, Ser. ix DC. Prod. 1. 392 (1824). 


Burm. Thes. 161. Pharnaceum distichum, L., Moon Cat. 23. Thw. 
Enum: 245) ©. P1004 


Fl. B. Ind. ii. 663. Burm. Thes.t. 71, fig. 2. Wight, Ic. t. 3(2). 


Stem solitary, erect, angular, finely glandular-pubescent, 
dichotomous with ascending branches; |. 3-6 in. a whorl, 
#-1 in., lanceolate-linear, tapering to base, subacute, glabrous, 
petiole obscure; fl. on short glandular-pubescent ped., arranged 
in corymbose cymes which afterwards lengthen out into stiff 
elongated racemes with distant fl.; sep. oval, obtuse, mem- 
branous ; capsule broadly ovoid, seeds very nearly smooth, 
shining, red-brown. 

Near the seacoast; rare. Colombo and Jaffna (Gardner). FI. white. 

Also in S. India. 


Wight’s figure represents a large hairy plant, unlike Ceylon examples, 
and Burmann’s is not very characteristic. Thw. says this is ‘not un- 
common’ on the coast, but there are no specimens collected by him 
among the C. P. plants. 


6. M. nudicaulis, Zam. Encycl. iv. 234 (1796). 
Thw. Enum. 24. Burm. Thes. 14. C. P. 2809. 
F]. B. Ind. ii. 664. Burm. Thes. t. 8, lower fig. 


Gisekia.] Ficoidee. 27a 


Annual, slender, |. all radical, crowded, about 14 in., oblong- 
spathulate, rounded at apex, much tapering into ill-defined 
petiole, glabrous, entire; fl.-stems several, erect, 4-6 in., 
slender, glabrous, leafless, trichotomously branched above, fl. 
on stiff, glabrous ped., arranged in lax trichotomous cymes ; 
sep. oval, very obtuse, with membranous margins, capsule 
nearly globose ; seeds minute, reticulate, dull black. 

Low country; rare: probably overlooked. Colombo (Thwaites). 


Fl. Dec.; white. 
Also in dry parts of India, Trop. Africa, New Caledonia, Cuba. 


4. GISEKIA,* Z. 


Annual, |. opp., without stip., fl. small, in axillary clusters ; 
sep. 5, slightly connate at base, persistent; pet. 0; stam. 5, 
hypogynous, fil. dilated below; carp. 5, distinct, ovule solitary 
in each carp., basal; ripe carp. enclosed in persistent sep., 
membranous, seed ovoid, slightly compressed, filling the carp., 
embryo curved into a ring.—Sp. 5 or 6; 1 in #2. B. Ind. 


G. pharnaceoides, ZL. Mant. 562 (1771). Manali, 7. 

Moon Cat. 23. Thw. Enum. 250. C. P. 1093. 

Fl. B. Ind. ii. 664. Wight, Ic. t. 1167. 

A rather succulent herb, Branches prostrate or ascending, 
elongated, glabrous; |. opp., $—-# in., oblong-spathulate, taper- 
ing at base to short petiole, obtuse or rounded and apiculate 
at apex, glabrous, rather thick, glaucous; fl. very small, 
numerous on slender ped., in dense, nearly sessile axillary 
umbels; sep. oblong-oval, obtuse, with membranous borders; 
ripe carp. membranous, surrounded by sep., seed solitary, 
rounded on back, black, with scattered short white promi- 
nences (glands ?). 

Low country in damp sandy places; rather common. Jaffna; Trin- 
comalie; seashore, Mt. Lavinia; Mannar. FI. Feb. 

Drier parts of India, Afghanistan, Africa. 

There are specimens of this in Hermann’s Herb. (i. fol. 19); and in 
his Mus. (p. 4) he says it is eaten in salads, as is still the case. Linnzeus 
seems to have included it under FI. Zeyl. n. 52 (Wollugo oppositifolia), 
Hermann having given the same S. name for it. 

The !eaves and other parts abound in large raphides. 


* Commemorates P. D. Giseke, M.D. of Hamburg, Editor of Linnzeus’s 
Prelectiones on the Natural Orders, published in 1792. 


PART IL. al 


274 U; mbelliferce. [ Hydrocotyle. 


LXIITI.—UMBELLIFER/A. 


ANNUAL or perennial herbs, 1. alt. simple or compound, 
without stip. (except in Hydrocotyle) but with petiole sheath- 
ing; fl. usually bisexual, often slightly irregular, in simple or 
compound umbels; cal.-tube adnate to ov.,segm. 5 or 0 (so 
that there seems to be no cal.); pet. 5, distinct, often bifid 
with an inflexed apex, imbricate (sometimes valvate in Hydro- 
cotyle); stam. 5, epigynous; ov. inferior, crowned with large 
epigynous disk, 2-celled with one pendulous ovule in each 
cell, styles 2; fruit of 2 separable carp. (mericarps) attached 
by summit to slender central axis, indehiscent, with 5 ridges 
down back and sides and often with sunken oil cavities (vittz) 
between them ; seed pendulous, with a small embryo in carti- 
laginous endosperm. 


F]. in heads or simple umbels 


Leaves entire I. HYDROCOTYLE. 


Leaves palmatisect . : 5 : : . 2. SANICULA. 
Fl. in compound umbels 
Fruit rounded or slightly compressed laterally 
Fl. yellow, 1. simple : : é . 3. BUPLEURUM. 
Fl. white 
Primary umbels with an involucre 4. CARUM. 
Primary umbels without an involucre 5. PIMPINELLA. 
Fruit much dorsally compres ee flat 
Glabrous : 5 6. PEUCEDANUM. 
Hairy 7. HERACLEUM. 


Only Io species represent have this vast Order of temperate climates. 
Of these, 6 are restricted to the hill country and 3 more extend into it, 
Carum stictocarpum being the only species confined to the low country. 


1. HYDROCOTYLE, /. 


Prostrate perennial herbs, rooting at the nodes, |. orbicular 
or reniform, with stip., fl. in very small heads or umbels opp. 
the 1.; cal.-segm. minute or 0; pet. entire, valvate or imbricate; 
fruit¥rather fleshy, laterally compressed, commissure narrow, 
mericarps with the primary and sometimes the secondary ribs 
visible, vittza o.—Sp. about 70; 5 in FZ. B. nd. 

Stip. cauline; carp. with primary ribs only 

L. 3-4 1n.; peduncles clustered . : - . I. H. JAVANICA. 

L. under 1 in.; ; peduncles solitary . 2. H. ROTUNDIFOLIA. 
Stip petiolar ; carp. with both pamary and se- 


condary ribs 3. H. ASIATICA. 


Hydrocotyle. OUmbellifere. Dis 


I. H. javanica, 7iund. Diss. ii. 415 (1800), Maha-gotukola, 5S. 

Hf. capitata, Moon Cat. 22. A. zeylanica, DC. Prod. iv. 6, 7. Z#. 
nepalensis, Hk., Arn. Pug. 19. Thw. Enum. 130. C. P. 1634, 2812. 

Fl. B. Ind. ii. 667. Wight, Ic. t. 1003 (4. folycephala). 


Stems very long, succulent, slightly pubescent especially 
beneath the nodes, sending up erect, stout, flexuose branches; 
. 1. 3-4 in, orbicular in outline, rather broader than long, very 
deeply cordate, palmately lobed less than half way down, very 
coarsely crenate, thin, very shining on both sides, with a few 
weak bristly hairs on the veins, petiole of |. from prostrate 
stem, erect, 6—8 in., of those of upright branches much shorter, 
cylindrical, pubescent, stip. large, cauline, rotundate, mem- 
branous ; fl. very small, sessile, 20 or more crowded in dense 
globular heads on short hairy peduncles 2-6 together from 
nodes of the upright branches; cal.-segm. 0; pet. lanceolate, 
acute, valvate; ov. much compressed, glabrous, style spreading ; 
fruit under 4 in., much compressed, smooth, mericarps with 
primary ridges. 

Moist places in grass under trees; very common in the moist region 
up to 7000 ft. Fl. May—October; pale green. 

Also in India, Malaya, Trop. Australia, and E. Trop. Asia. 

The large stip. completely enclose the end of the growing branch and 
young leaf. At the higher elevations the petioles and peduncles are more 
woolly, and the flowers on longish pedicels, var. B, Zhw. (C. P. 2812). 
H!. zeylanica, DC., was collected by Leschenault about Kandy, and is 
merely a luxuriant form. 

Used medicinally in the same way as H. aszatica. Mr. E. Green, of 


Pundaluoya, states (Taprobanian, ili. 7) that the Tamil coolies employ it 
as a fish-poison. 


2. H. rotundifolia, Rox). Hort. Beng. 21 (1813). 
H. nitidula, Hk., Thw. Enum. 130. C. P. 1635. 
Fl. B. Ind. ii. 668. Wight, Ic. t. 564. 


Very small, stems filiform, copiously rooting, glabrous ; 
1. $-? in., nearly circular in outline, the basal lobes almost in 
contact, but not overlapping, palmately cut half way down 
into 5 or 7 broad obtuse lobes, crenate, glabrous and shining 
above, with few coarse hairs beneath, petiole erect, 4-14 in., 
glabrous, stip. membranous, obtuse; fl. minute, almost sessile, 
4-8 in a head, peduncle very short, slender, solitary, bracts 
oblong, as long as fl.; pet. lanceolate, acute, valvate; styles 
spreading; fruit extremely small, rather broader than long, 
compressed, mericarps with primary ribs blunt, glabrous. 

Upper montane zone, in moist places rather common. Nuwara Eliya; 


Hakgala; Horton Plains. FI. all the year; pinkish. 
Also in India, Malaya, W. Trop. Africa. 


276 Umbellifere. [Sanicula, 


3. H. asiatica, LZ. Sf. Pl. 234 (1753). Win-gotukola, S. 
Vallarai, 7. 

Herm. Par Bat; 238) Burm.) Thes. 122. Fli(Zeyl. ni 11ce\aen 
Catr22-ebhwa enum 130% Pa2o10: 

Fl. B. Ind. ii. 669. Herm. Par. Bat. t. 238. Wight, Ic.t. 565. Bent. 
and Trim. Med. PI. t. 117. 

Stems long, prostrate, given off from l.-axils of a short 
vertical rootstock, cord-like, glabrous, with very long inter- 
nodes; 1. several from the rootstock and 1 to 2 from each 
node of the runners, petiole 3-6 in., erect, glabrous, furrowed 
above, stip. short, adnate to petiole and forming a sheathing 
base, blade horizontal, more or less cupped, 14$—2 in., reniform, 
rather broader than long, the rounded basal lobes often over- 
lapping, shallowly dentate-crenate, glabrous and shining on 
both sides; fl. nearly sessile, usually 3 together at end of short 
erect pubescent peduncles, 1-3 from the nodes, opp. the lL, 


bracts 2, close beneath umbel, ovate, obtuse; cal.-segm. 0; 


pet. minute, ovate, acute, slightly imbricate (?); ov. very much 
compressed, slightly hairy, styles very short, erect; fruit 
about 4in., ovoid, hard, pericarps thickened, mericarps with 
the primary and secondary ribs very obscure or vein-like. 

Waste grassy places from sea-level to the highest elevations; a very 
common weed. FI. May to October; dark pink. 

In all Tropical and Sub-tropical countries. 

Much used as an alterative tonic, especially for children, and has aleo 
a reputation for skin complaints. The powdered dry leaves are an official 
drug in the Indian Pharmacopceia. 


2. SANICULA, Z. 


A perennial, erect herb, 1. deeply palmatisect; fl. minute, 
in small involucrate heads arranged in cymes; cal.-segm. 
lanceolate ; pet. with a long incurved point; fruit ovoid, not 
compressed, thickly covered with hooked spiny prickles, ribs 
obscure.—Sp. 10; 1 in AZ. B. Ind. 


S. europea, L. Sp. Pl. 235 (1753). 

Thw. Enum. 130. C. P. 2 2813, 

Fl. B. Ind. ii. 670. Wight, Ic. t. 1004 (S. e/aza). 

Rootstock stout, ascending; 1. mostly radical, on erect 
petioles 4-6 in. long and glabrous and sheathing at the base, 
14-2 in, very broadly ovate in outline, cordate at base, 
palmately cut nearly to the bottom into 3 acute, spinous- 
serrate lobes often again deeply 3-fid, glabrous on both sides, 
dark green above, paler beneath; flowering stem 9-18 in. 
high, slightly dichotomously branched above, with a short- 
petioled leaf at each node, striate, glabrous ; fl. sessile, 3 or 4 


wee TS 
2a ah he Bt ee ee EI 


Bupleurum.] Unibell. afer a. 2777) 


in little heads on long peduncles arranged in trichotomous 
cymes, bracts of involucre about 9, short, linear, acute ; pet. 
minute, bifid, with a long incurved acute point; ov. thickly 
covered with long, erect, soft, white, hooked prickles, styles 
long, spreading; fruit about 3in., ovoid, not compressed, 
thickly set with stiff spreading hooked prickles, mericarps 
semicircular on section, the ribs very obscure. 

Upper montane zone in woods, very rare. N. Eliya (Nock); Elk 
Plains; between Maturata and Nuwara Eliya. Fl]. April-June; white. 

Also in the Himalaya and Nilgiris, and in Temp. Asia and Africa and 
Europe. An English species. 


3. BUPLEURUM, L. 


Herbaceous, |. quite entire, fl. in compound umbels with 
both general and partial involucres, small, yellow; cal.-segm. 
0; pet. broad, notched point much inflexed ; disk broad, flat ; 
fruit not compressed, mericarps somewhat pentagonal, ridges 
very distinct, with 1-3 vitte between them.—Sp. 60; 13 in 
Fl. B. Ind. 


B. virgatum, VW. and A. Prod. 370 (1834). Wal-enduru, S. 

B. nervosum, Moon Cat. 22. JS. falcatum, L., var., Thw. Enum. 131. 
C, Biger; 

Fl. B. Ind. ii. 676 (B. mucronatum, var.). Wight, Ic. t. 1007 (B. 
YAMOSISSLNIUUML). 


Annual (or perennial ?), stem erect, 1-3 ft., with many 
ascending branches, cylindrical, striate, glabrous; |. grass-like, 
sessile, 24-44 in., amplexicaul but not sheathing at base, 
linear, acuminate, mucronate at apex, glabrous, glaucous green, 
with a very narrow horny margin, veins slender, nearly parallel, 
distinct and unbranched; general umbels long-stalked, nu- 
merous, arranged in lax, elongated, terminal, somewhat 
corymbose panicles, rays 4-6, general involucre of 3-6 lanceo- 
late-linear very acuminate bracts, partial umbels of 6-8 fl, ped. 
short, partial involucre of 4 or 5 bracts rather exceeding f1.; 
pet. distant, broad, point much inflexed; fruit rather over } in., 
ovoid, brown, shining, mericarps rounded on back with the 
primary ribs prominent, and with a broad, conspicuous dark 
vitta in each furrow, often with a smaller one on either side. 

Rocky open places and patanas in the montane zone, 3000-7000 ft.; 
common on the Eastern (Uva) slopes, but scarcely found on the Western 
‘Kandy) side. FI). Feb.-August; lemon-yellow. 

Also in the Nilgiri Mts. 
The Ceylon plant is not at all variable except in size, but the Fl. B. 


Dy 8 GT; mbelitferce. [| Pzmpinella. 


Ind. is no doubt right in combining Wight’s three species, B. vzrgatum, 
mucronatum, and ramosisstmunz, into one. Moon’s name is the oldest, 
but was published (as were all of his names) without description. 


4. CARUM, L. 


Herbaceous, |. compound; fl. in compound umbels with 
both general and partial involucres; cal.-segm. very small ; 
pet. broad, obtuse; fruit ovoid, scarcely compressed, meri- 
carps sub-pentagonal, ridges slender, with a single vitta in the 
furrows.—Sp. about 50; 10 in /Z. B. Lnd. 


C. stictocarpum, Clarke in 27. B. Ind. ii. 688 (1879). 


Annual, 2-5 in. high, much branched, stem glabrous; 
lower |. pinnate or bipinnate, lflts. deeply cut into linear- 
lanceolate apiculate segm., glabrous petiole with broad ciliate 
sheaths ; primary umbels with 3-5 rays, involucre of 3 seta- 
ceous bracts, partial umbel of 6-10 fl., involucre of 3-6 lan- 
ceolate-linear bracts with broad white margins; ov. finely 
pubescent; pet. broad, obtuse; fruit very small, glabrous, 
brown, mericarps not compressed, the ridges slender, filiform, 
rather faint. 

Dry country, very rare. In sheltered crevices of the rocky boulders 
of coral-stone forming the headland at Kirimalai near Kankesanturai, 
Jaffna, abundant. There are also specimens in Herb. Perad. from 
W. Ferguson, without locality, labelled ‘ Pzmpinella involucrata (?)’ 
Fl. Feb., white. 


Also from Concan, Western India, and Mt. Abu, N.W. Provinces, 
India. 

The numerous specimens in Hb. Kew are all much more luxuriant 
than ours, which are, doubtless, much dwarfed by the very hot and dry 
locality. It is possibly an accidental introduction. 


[C. nothum, Clarke, is recorded with doubt for Ceylon in FI. B. Ind. ii. 
681. There are specimens in Hb. Kew so localised by Dr. T. Thomson, 
but an error is probable. Another specimen is from Mysore. | 


C. Roxburghianum, Benth. (Ptychotis, DC.), cultivated in native 
gardens, is an occasional weed. It is figured in Wight, Ic. t. 567. 


5. PIMPINELLA, L. 


Annual or perennial, |. simple or compound, fl. in com- 
pound umbels, with partial but no general involucre; pet. 
with long inflexed points ; fruit ovoid or subglobose, slightly 
compressed, mericarp subterete, but flat or concave at com- 
missure, ridges prominent or obscure, vittze 2 or 3 in each 
furrow.—Sp. about 70; 22 in FZ. B. Ind. 


Leaves trifoliolate . P . I. P. HEYNEANA. 
Leaves simple. : : . 2. P. LESCHENAULTII. 


Peucedanunt | Unibellifere. 27.9 


I. P. Heyneana, Wall. Caz. 7.566 (1828). Wal-asamodagan, S. 

Stum triternatum, Moon Cat. 22. Aelosciadium Heyneanumt., DC., 
Thw. Enum. 130. Arn. Pug. 19. C. P. 182. 

Fl. B. Ind. ii. 684. 

Annual, stem 13-3 ft., with ascending branches, glabrous; 
|. trifoliolate, smaller upwards, rachis 14-3 in., dilated with 
ciliate margin at base, lfits. stalked, lanceolate or linear- 
lanceolate, tapering to base and apex, acute, coarsely serrate, 
glabrous, the lat. ones often deeply 3-lobed; primary umbels 
6—12-rayed, rays slender, unequal; partial umbels 4—8-flowered, 
with long spreading unequal ped., bractlets 1 or 2, linear, or 0; 
styles short; fruit very small, 7; in., almost globose, slightly 
didymous, primary ridges slender, prominent. 

Waste ground and roadsides, rather rare. Four Korales (Moon); 
Uva, very abundant up to 4ooo ft. (Thwaites); Morowak Korale; Anurad- 
hapura. 

Also in India and Burma. 

In Morowak Korale this was growing in great abundance with 
Kurakkan, and looked as if purposely sown. 


2. P. Leschenaultii, DC. Prod. iv. 122 (1830). 

Thw. Enum. 131. C. P. 515. 

FI. B. Ind. ii. 687. Wight, Ic. t. 1005. 

Perennial with a branched rootstock ; |. nearly all radical, 
petioles erect, from 2 to 12 in., sparingly hairy, much dilated 
and sheathing at base, blade 14-4 in., rotundate-ovate, very 
deeply cordate at base, with a narrow sinus or with the 
rounded lobes slightly overlapping, coarsely crenate-dentate, 
glabrous above, more or less hairy beneath and on edges, 
rather thick, veins conspicuous beneath, stem-l. few, much 
smaller and on shorter petioles, the uppermost ones cut into 
lanceolate segm.; primary umbels with 12-20 equal pubescent 
rays, partial ones of 15-20 fl., ped. pubescent, bractlets few, 
setaceous, hairy; pet. with a long inflexed point, the outer 
ones of the umbels somewhat radiant; styles rather long, 
erect, ov. pubescent; fruit about ¢in., ovoid, rather tapering 
upwards, crowned with cal.-segm. and conical disk, mericarps 
very slightly compressed, concave on commissure, ribs obscure, 
vittz small, 2 in each furrow, and 2 on commissure. 

In wet places in the patanas, upper montane zone, very rare. Only 


on Horton Plains and on the descent thence to Galagama. FI. Feb.; white. 
Also in the Nilgiris. 


6. PEUCEDANUM, J. 


Herbaceous, |. triternate, umbels compound, with no 
general and a small partial involucre; pet. obovate, bifid ; 


280 Ombellifere. | Heracleum. 


fruit oblong-oval, very much compressed, mericarps flat, the 3 
dorsal ridges slender and narrow, the 2 lat. ones expanded 
into narrow wings, vitte solitary in the furrows.—Sp. about 
100; 9 in Fd. B. Ind. 


P. zeylanicum, Gardn. in Calc. Journ. Nat. Hist. viii. 14 (1847). 

Palimbia ramosissima, Thw. Enum. 131 (non DC.). C. P. 135. 

Fl. B. Ind. 11. 710. 

Annual (?), stem erect, 3-5 ft., branched, striate, glabrous, 
hollow ; radical 1. about 1 ft. long, triternate, rachis slender, 
glabrous, sheathing at base, lflts. very deeply 3-sect, the lobes 
linear, acuminate, very acute, glabrous, glaucous; uppermost 
1. reduced to sheaths with a few small filiform segm.; primary 
umbels with 10-12 rays, no involucre, partial of 10-12 fl. on 
very long ped., bractlets few, setaceous; fruit {-% in., very 
much compressed, tipped with conical disk, mericarps very 
thin, the dorsal ridges slightly raised. 

Upper montane zone, very rare. Found by Gardner, in 1846, in open 
grassy rocky places on the descent from Horton Plains to Galagama, but 
not since collected. Fl. Feb., March; white. 

Gardner says that the plant when bruised gives an odour like that of 
Fennel, and that it was called ‘ Wal-enduru’ by the Sinhalese. 


7, HERACLEUM, Z. 


Perennial herb, 1. mostly radical, pinnate, umbels com- 
pound, many-rayed, with both general and partial involucres ; 
cal.-segm, acute; pet. with long inflexed point; fruit oval, very 
much compressed, mericarps flat, with 3 dorsal ribs flat and 
broad and 2 lateral ones expanded into broad wings, vittz 
conspicuous, one in each furrow and. several on .commissure, 
carpophore bipartite—Sp. 50; 21 in FZ. B. Ind. 


H. zeylanicum, Gardn. in Fi. B. Ind. ii. 716 (1879). 

Szum lobatum, Moon Cat. 22. HA. nepalense, Dor, Arn. Pug. 19. 7. 
Sprengelianum, Thw. Enum. 131 (non W. and A.). C. P. 145. 

Fl. B. Ind. ii. 716. 

Rootstock stout, ascending, nodose, tough; |. mostly 
radical, spreading, pinnate, rachis 4-10 in., stout, cylindrical, 
hairy, with broad ciliate stip. adnate to lower part and forming 
a sheathing base, lflts. 3-7 (1-3 pair and an end one), 2-3 in., 
the lowest pair generally stalked, broadly ovate in outline, 
but very deeply pinnately cut (often almost to the base) into 
broadly oblong, acute, coarsely spinous-serrate lobes, hairy on 
both sides, especially on the raised reticulate venation beneath; 


Polyscias.| Arahacee. 281 


flowering stem 1-24 ft., erect, slightly branched, striate, finely 
hairy, stem-l. few, gradually smaller upwards, and at top re- 
duced to the broad sheathing petioles without Iflts., umbels 
few, on long stalks, general with about 12-20 rays and an 
involucre of 2 or 3 inconspicuous obtuse bracts, partial with 
about 20 fl. on rather long ped. and an involucre of few un- 
equal lanceolate bractlets; pet. broad, a long, attenuate 
incurved apex; styles short, curved ; fruit 3-4 in., very much 
compressed, oval or oblong-oval, rounded at both ends, often 
slightly obovate, mericarps much dorsally flattened, clabrous 
and shining, wings semi-transparent, vitte black. 

Upper montane zone, rather common. FI. Feb.—July ; greenish-white, 
anth. purple. 

Endemic. 

In Fl. B. Ind. ii. 715, some specimens of C. P. 145 collected by 
Gardner on Adam’s Peak are referred to AH. rvzgens, Wall. figured in 
Wight, Ic. t. 1059. I do not, however, think we have more than a single 
species, and, though probably it is conspecific with the Nilgiri forms, 
it does not quite agree with any of Wight’s figures or descriptions. 
That figured in Wight, Ill. ii. t. 116 (as Pastinaca ligustictfolia), seems 
nearest. The form of the lower leaves is very variable, and on Horton 
Plains occurs a luxuriant form with very much cut leaves, which are nearly 
glabrous. 


LXIV.—ARALIACE/E. 


TREES or climbing shrubs, |. compound, alt., with stip. adnate 
to petiole; fl. regular, bisexual; cal.-tube adnate to ov., 
segm. very small or 0; pet. 5-9, distinct, valvate ; stam. 5-9, 
epigynous; ov. quite (rarely only 4-) inferior, crowned with 
epigynous disk, 4-9-celled, with one pendulous ovule in each 
cell, styles as many as cells, distinct or connate or 0; fruit 
small, berry-like, indehiscent, pyrenes several, compressed, 
embryo minute in copious endosperm. 


L. pinnate ; : ; : 5 , : MIE LOLVSCIAS: 
L. digitate : ; 5 : ‘ : . 2, HEPTAPLEURUM. 


Heptapleurum allan and emarginatum inhabit the moist low 
country, Polyscias and the other species of Heptapleurum the montane 
zone, into which also #7. emarginatum extends. 


I. POLYSCIAS, /orst. 


Tree, |. imparipinnate, with adnate stip., fl. jointed on the 
ped., in panicles; cal.-limb with 5 tooth-like segm. pet. 5, 


282 Aratiacee. [Heptapleurum. 


thick, valvate ; stam. 5, small, epigynous; disk flat ; ov. quite 
inferior, 4- or 5-celled, with one pendulous ovule in each cell, 
styles 4 or 5, distinct; fruit broadly ovoid, ribbed, pyrenes 
compressed, endosperm not ruminate.—Sp. 8; 2 in FZ. B. Lud. 


P. acuminata, Seem. in Journ. Bot. iii. 181 (1865). 

Hedera acuminata, Wight, Thw. Enum. 131. C. P. 4. 

Fl. B. Ind. ii. 727. Wight, Ic. t. 1062. Bedd. Fl. Sylv. t. 213. 

A small tree, quite glabrous; |. large, pinnate, rachis 
8-9 in., stout, cylindrical, somewhat thickened at base and 
amplexicaul, Iflts. 5 or 7 (2 or 3 pairs and end one) shortly 
stalked, 34-44 in., lanceolate, acute at base, attenuate, very 
acute at apex, entire, with narrowly recurved margin, glabrous, 
rather thick; fl. on short stout ped. articulated at each end, 
3-5 in little umbels terminating short branches, which come 
off from upper parts of thick erect axes, 2-5 of which are 
umbellately arranged at end of stout peduncle opp. the lL, 
bracts linear, setaceous ; cal. glabrous, segm. acute; pet. very 
thick, concave, apex inflexed, acute; disk broad, fleshy ; 
styles short, erect, blunt; fruit (not seen ripe) crowned with 
cal.-limb and enlarged spreading styles, obscurely ribbed, 
glabrous. 


Montane zone; very rare. Banks of streams, Gangalla, near Adam’s 
Peak, between 4000 and 5000 ft. (Thwaites); Bogawantalawa. FI. Feb.- 
May; pale green. 

Also in the mountains of S. India. 


| Pentapanax Leschenaultz:, Seem., is given for Ceylon in FI. B. Ind 
ii. 724, but I do not know on what authority. There are no specimens. ] 


Panax fruticosum, L., is a very common cultivated hedge-plant. Itis 
probably a native of Malaya, but is nowhere now found wild. 


2. HEPTAPLEURUM, Gaertn. 


Trees or climbing shrubs, |. digitate, with stip. adnate to 
petiole or intrapetiolar, fl. in umbels or racemes arranged in 
panicles ; cal.-limb small or absent; pet. 5-9, valvate, stam. 
same number as pet., epigynous, disk flat or prominent, ov. 
quite or half inferior, cells as many as pet. and stam., with a 
single pendulous ovule in each cell, styles usually o or very 
short or combined into a cone; fruit small, ovoid, often bluntly 
ribbed, endosperm not ruminate.—Sp. 50; 15 in FZ. B. Ind. 


Lfits. 4-9. 
Styles combined intoacone . 5 : . I. H. RACEMOSUM. 
Styles o. 
Climbing shrub; disk flat. : s . 2. H. STEDUATIOE 
Tree ; disk conical ; : 5 ‘ . 3. H. EXALTATUM. 


Lfits. solitary or 3. : : j : : . 4. H. EMARGINATUM. 


Fleptapleurum.| Araliacee. 28 3 


I. H. racemosum, edd. Fl. Sylv. t. 214 (1873 °?). 

Hledera racemosa, Wight, Thw. Enum. 132. C. P. 549. 

Fl. B. Ind. ii. 729. Wight, Ic. t. 1015. Bedd. FI. Sylv. t. 214 (froma 
Ceylon specimen). 

A large tree, much branched, branchlets stout, marked 
with scars of fallen l., young parts covered with dense, pale 
orange, readily detached scurf; 1. palmate, petiole 4-5 in., 
cylindrical, thickened and stem-clasping at base, Iflts. 5-9, 
spreading, on stalks about 1 in. long, oblong-ovate, 3-4 in., 
rounded at base, acuminate, usually twisted at apex, often 
conduplicate, quite glabrous when mature, dark green above, 
paler beneath, rather coriaceous, venation reticulate, pellucid ; 
fl. numerous, on short, stout, furfuraceous ped., arranged in 
umbellate stalked racemes spreading divaricately from stout 
rachis of large, erect, or drooping terminal or extra-axillary 
panicle which is over 12 in. long, buds small, ovoid, blunt ; 
cal. furfuraceous, limb very slightly developed, truncate ; pet. 
usually 5, ovate, subacute ; disk nearly flat; ov. quite inferior, 
styles combined into a short blunt cone; fruit about } in, 
ovoid, tipped with a beak, with 5-9 blunt, vertical ribs. 

Forests of montane zone, 3000-6000 ft.; rather common. FI. April, 
May; pale green. 

Also in the mountains of S. India. 


The leaflets of young plants are often deeply gashed, with unequal 
narrow very acute segment. 


2. H. stellatum, Gaertn. Fruct. ii. 472 (1791). Itta, Itta- 
wel, S. 

Herm. Mus. 50. Hedera terebinthacea, Vahl, Symb. i. 42; Moon 
Cat. 18. Paratropia terebinthacea, Arn. Pug. 20. Hedera Vahliz, Thw. 
Bosn.132.  C. P. 1632. 

Fl. B. Ind. ii. 730. Wight, Ic. t. 1011 (Hedera osovata). 


A large, scandent, epiphytic shrub, branchlets stout, 
glabrous, marked with leaf-scars ; 1. palmate, petiole 24-4 in., 
cylindrical, glabrous, thickened and with an amplexicaul 
sheath at base, lflts. 4-7, on petioles 4-1 in., oval, 24-33 in., 
acute at both ends, rather thick, glabrous; fl. over § in., on 
rather long slender glabrous ped. articulated at base, 3-8 in 
stalked umbels arranged on branches of large terminal 
panicles, bracts membranous, acuminate, soon falling, buds 
very blunt; cal. truncate without a limb; pet. 6-9 (usually 8) 
acute, with an inflexed point; stam. shorter than pet.; disk 
large but not prominent; ov. 6-9-celled ; styles 0; fruit over 
} in., clavate-ovoid, faintly ribbed, yellow. 

Moist low country to 3000ft. or higher; common. FI. June; yellow. 

Also in S. India. 

The leaves used as a cattle medicine. 


284 Arahacee. | Heptapleurum. 


3. H. exaltatum, Seem. 77 Journ. Bot. iii. 80 (1865). 

Fledera exaltata, Thw. Enum. 132. C. P. 1633. 

Fl. B. Ind. i. 730 (4. Wallichianum). 

A large tree, glabrous; |. palmate, petiole 8-9 in., stout, 
cylindrical, glabrous, Iflts. 5-9, on stalks 14-2 in. long, large, 
5-8 in., oblong-lanceolate, rounded at base, acuminate, entire, 
glabrous, thickly coriaceous ; fl. nearly 4 in., ped. § in., 12-20 
in long-stalked umbels, arranged on a stout rachis over 1 ft. 
long, no bracts, buds large, nearly globular; cal. glabrous, 
limb truncate; pet. 7-0, linear-oblong, acute, thick; stam. 7-9, 
spreading, fil. flat, narrow; disk very large, conical, blunt, 
ribbed ; ov. 4-inferior, styles 0; fruit 4 in., globose, the upper 
half above the rim of the cal.-limb, with 7-9 blunt vertical 
ribs, glabrous. 

Montane zone, 4000-6000 ft.; rather rare. Hunasgiriya; Passelawe; 


Ramboda. FI. June. 
Also in the mountains of S. India. 


4. H. emarginatum, Seem. 72 Journ. Bot. i. 80 (1865). [PLATE 
SAV: 

Woe emarginata, Moon Cat. 18; Thw. Enum. 132. C. P. 655. 

Fl. B. Ind. 11. 729. 

An epiphytic climbing shrub, but often dwarf and bushy, 
glabrous, branches stout, virgate, with numerous short lateral 
branchlets, bark silvery-grey ; 1. numerous, closely placed, 1- 
(rarely 3-) foliolate, petiole #-1 in., slender, thickened and 
amplexicaul at base, Iflts. nearly sessile, articulated on petiole 
14-14 in., cuneate or oblong-cuneate, tapering to base, truncate 
rounded and deeply emarginate or 2-lobed at apex, very thick 
(veins prominent when dried); fl. rather small, on rather long 
stiff ped., 2-4 together in small umbels at end of divaricate, 
slender branches of very lax panicles coming off opp. the 1. 
of the lat. branchlets, no bracts; cal.-limb 0, pet. 5, ovate, 
acute, very soon falling; stam. longer than pet.; disk rather 
prominent; ov. quite inferior, 5- (rarely 4-) celled, styles 5 
(or 4), very short, distinct; fruit small, 3 in., broadly ovoid, 
truncate, smooth, pulpy, dark purple. 

Moist country up to 4000 ft.; rather rare. Sabaragamuwa (Moon); 
Hunasgiriya (Thw.); Kukul Korale; near Peradeniya; Kuruwita Korale. 
Fl. May, June; greenish-pink. 

Endemic. : 

This curious species often appears as an ordinary bush, but it is able 
to climb the largest trees. 


Alangium.| Cornacee. 285 


LXV.—CORNACE. 


TREES or shrubs, I. alt., without stip.; cal.-tube adnate to ov., 
segm. 4 or 5; pet. 4-10, valvate; stam. def. or indef., epigynous; 
ov. quite inferior, crowned with epigynous disk, 1-celled with 
a single pendulous ovule, style simple; fruit a drupe, epicarp 
fleshy, stone woody; seed oblong, embryo in fleshy endosperm. 
Stam. I5 or more; pet. very long . : ‘ : . I. ALANGIUM. 
Stam. 4 or 5; pet. short . : 3 ; ; : a 25 MUAGTIDSTAL 


Alangium Lamarcki is a dry-country plant, MJastixia arborea is 
montane, the other two species are found in the moist region, extending 
into the hills. 


1. ALANGIUM,* Zam. 


Trees or shrubs, 1. alt., no stip., fl. in axillary clusters ; 
cal.-tube adnate to ov., limb saucer-shaped, segm. 5, small ; 
pet. usually 6 (4-10) long, valvate; stam. 15-indef., epigynous, 
fil. short, with a tuft of hair near base, anth. very long, linear; 
ov. inferior, 1-celled, with a single pendulous ovule, style very 
long, surrounded at base by epigynous disk, stigma large, 
4-lobed; drupe externally fleshy, crowned with cal.-limb ; 
seed solitary, embryo straight in axis of slightly ruminate 
endosperm, cotyledons foliaceous.—Sp. 2 or 3; 1in FZ B. Lud. 


Small erect tree; fruit I in. . : : ; . I. A. LAMARCKIL. 
Subscandent shrub; fruit 2Zin.. . : ‘ . 2, A. GLANDULOSUM. 


1. A. Lamarckhii, 7/iw. Enum. 133 (1859). Mul-anninchil, 7. 

A. decapetalum and A. hexapetalum, Lam., Moon Cat. 42. Thw. 
Enum. 133. C. P. 760. 

Fl. B. Ind. ii. 741. Wight, Ic. t. 194 (A. decapetalum), Ill. t. 96 (A. 
hexapetalum). 

An erect tree of small size, occasionally with short, sharp, 
spinous branchlets, bark whitish, young parts pubescent ; 
|. variable, 24-6 in., oblong-oval or oblong-lanceolate, acute or 
rounded at base, more or less acuminate, obtuse at apex, 
entire, glabrous above, pubescent on veins beneath and 
often with small tufts of hair in their axils, venation rather 
prominent beneath, the lat. veins very oblique, petiole short, 
pubescent; fl. large, on densely pubescent ped. 3 in. long and 
jointed near the top, 1-3 in the axils; cal. silky-pubescent, 


* Alangi is given by Rheede as the Malabar name. 


286 Cornacee. [ Wastixia. 


segm. very shallow, acute; pet. usually 6, about 2—? in., 
narrowly linear, densely pubescent outside, reflexed; stam. 
numerous (over 20), nearly as long as pet., erect; style as 
long as stam., stigma very large; fruit nearly globular when 
fully ripe (previously oblong-oval), near I in. diam., capped 
with persistent cal.-limb, finely pubescent, not ribbed, pur- 
plish-red. 

Dry and intermediate regions; rather common. Jaffna; Bintenne; 
Polonarua; Uma-oya; Lagalla District; Hanguranketa; Dikwelle Uva; 
Kurunegala. Also about Colombo, but not wild there. Fl. June-July; 
white. 

Also throughout India and Malaya, S. China, Philippines, and E. 
Africa. 

Moon gives native names, ‘ Ipetta’ and ‘ Katapetta,’ for this, and 
‘ Kiri-anguna’ has been given me, but I doubt if it has a real Sinhalese 
name. Specimens from Polonarua have narrowly oblong-linear leaves. 

The bark is bitter. Heart-wood hard, close-grained, dark yellow. 


2. A. glandulosum, 7iw. Enum. 133 (1859). 
Ceo IPs Ail 
Fl. B. Ind. 11. 742 (A. Lamarckiz, var.).. 


A subscandent shrub, climbing by its horizontal branchlets, 
bark brown, young parts furfuraceous, dark red; 1. 24-5 in., 
oblong-oval or oblong-lanceolate, acute at base, conspicuously 
caudate-acuminate, glabrous above, very minutely glandular- 
puberulent beneath, lat. veins with small glandular pits in 
their axils; fl. smaller than in the last, ped. glandular- 
pubescent, articulated close to the top with 3 or 4 small 
bractlets beneath the joint, 1-4 in the axils; cal. glandular- 
pubescent, segm. small, very acute; pet. usually 6, barely $in., 
linear ; stam. 15-20; fruit 2 in. diam., nearly globose, smooth, 
purplish-red. 

Moist region, 2000-4000 ft.; rather rare. Dolosbagie; Alagala; 


Hantane; Ambagamuwa; Adam’s Peak. FI]. March; white, calyx red. 
Endemic. 


This is regarded as a variety of A. Lamarckiz in FI. B. Ind.; it does 
not appear ever to have thorns. 


2, MASTIXIA, 3/. 


Large trees, |. alt., without stip., fl. in terminal panicles; cal.- 
tube adnate to ov., segm. 4 or 5, small; pet. 4 or 5, valvate; 
stam. 4 or 5, epigynous, anth. short; disk large, fleshy ; ov. 
quite inferior, 1-celled, with a single pendulous ovule, style 
simple ; drupe with a hard fleshy epicarp, and a bony endo- 
carp, which is deeply intruded down one side; seed ovoid, 


Mastixia.] Cornacee. 237 


embryo narrow, in copious fleshy endosperm, radicle very long. 
—Sp. 8; 6in Fl. B. lnd. 


Placed in Olacacee by some authors. 


Stam. and pet.4 . : . : ° 6 . I. M. TETRANDRA. 
Stam. and pet.5 . 3 : : : : . 2. M. ARBOREA. 


1. M. tetranda, Clarke ix Fl. B. Ind. ii. 745 (1879). Maha- 
tawara, S. [PLATE XLVII.] 

Bursinopetalum tetrandrum, Wight, Thw. Enum. 42. C. P. 2441. 

FI]. B. Ind. ii. 745. 

A large tree, bark smooth, pale, twigs numerous, stout, 
puberulous ; |. 3-4 in., oval or lanceolate-oval, acute at base, 
caudate-acuminate, obtuse at apex, entire, glabrous, rather 
stiff, dark green, pale beneath, petiole 4 in., channelled above ; 
fi. small, about i in., on very short pubescent ped., in lax, 
terminal, corymbose panicles ; cal.-segm. broad, shallow; pet. 
4, ovate, acute; stam, 4, fil. very short ; style very short; fruit 
2 or = in., ovoid-oblong, crowned with small cal.-teeth and 


style, cylindrical, smooth. 


Var. 8, Thwaitesii, Clarke, /.c. Diya-taleya, S.C. P. 2542. 


Young parts glabrous, |. crowded, smaller, 13—2 in., erect, 
rounded or very obtuse at apex, rather coriaceous; panicles 
small, few-flowered, glabrous. 

Moist region up to 4ooo ft.; rather rare. Deltota; Ambagamuwa. 
Var. 8, montane zone, 4000-7000 ft.; rather common. Fl. March—-May; 
pale green. 

Endemic. ce 

Var. 8 ought perhaps to be regarded as a distinct species. Wood red, 
rather heavy, liable to split. The leaves have a resinous odour and dry 
black, and the bark yields a scented resin. The inflorescence is some- 
times monstrous, the flowers being converted into short twigs, set with 
minute imbricated bracts. 


2. M. arborea, Clarke in Fl. B. Ind. ii. 746 (1879). 

Bursinopetalum arboreum, Wight, Thw. Enum. 42. C. P. 637, 2440. 

Fl. B. Ind. ii. 746. Wight, Ic. t. 956. 

A large tree, twigs very stout, glabrous, marked with leaf- 
scars; 1. 4-8 in., oval or obovate-oval, tapering to base, rounded 
or very shortly acuminate at apex, margins often revolute, 
glabrous, coriaceous, petiole 4-14 in., broad, flattened above ; 
fl. in. wide, on very short ped., in threes at ends of branches 
of stout, terminal corymbose panicles, bracts small, lanceolate, 
acuminate; cal. glabrous, segm. triangular, acute; pet. 5, 
ovate-oblong, acute, with an inflexed thickened apex; stam. 
5; disk large, prominent; style short, conical; fruit about 
1} in., oblong-ovoid, cylindrical, truncate, smooth, purplish- 
green. 


288 G, apref. oltaceg. [Viturnum. 


Montane zone, 4000-7000 ft., rather rare. Hantane; Dimbula; 
Ambagamuwa; Nuwara Eliya. FI. March, April; green, disk yellow. 

Also in Cachar and the Nilgiris. 

The leaves dry black. C. P. 2440 has leaves with larger petioles and 
a broader fruit. 


LXVI.—CAPRIFOLIACE. 


SHRUBS or small trees, 1. opp., without stip., fl. regular, bi- 
sexual, in paniculate cymes; cal.-tube adnate to ov., limb 
small, segm. 5; cor. tubular-campanulate, lobes 5, equal ; 
stam. 5, inserted on cor.-tube; ov. inferior, 1-3-celled, with a 
single ovule in each cell, style very short, stigma large, slightly 
3-lobed; fruit a drupe, with a thin hard endocarp, 1-seeded; 
seed compressed, embryo very small in fleshy endosperm. 


Chiefly distinguished from Rudzacee by the absence of stipules. Both 
our species occur only in the higher mountain region. 


VIBURNUM, J. 


For characters, see Order.—Sp. 80; 17 in FV. B. nd. 


L. entire; fruit black : : : 4 : . I. V. CORIACEUM. 
L. serrate; fruit red . ‘ : : : : . 2. V. ERUBESCENS. 


1. V. coriaceum, &/. Bijd., 656 (1825). 

V. zeylanicum, Gardn. ex Hk. f. and Th. in Journ. Linn. Soc. ii. 179. 
V. hebanthum, Thw. Enum. 136 (non W. and A.). C. P. 39. 

BiB. Indean: 5: : 

A large shrub, young parts glandular-pilose; 1. 4-5 in.; 
oval-lanceolate, subacute at base, shortly acuminate, acute at 
apex, entire, somewhat undulate, glabrous above and very 
nearly so beneath; rather thick, lat. veins about 6 on each 
side, oblique, prominent beneath, petiole ?-1 in., channelled 
above; fl. sessile, in threes at ends of branches of terminal, 
stalked, umbellate-corymbose cymes, finely glandular-pubes- 
cent; cal.-limb very short, segm. very small; cor. nearly 
glabrous, lobes very small, acute, spreading; stam. slightly 
exserted ; fruit } in., oblong-ovoid, compressed, tipped with 
small cal.-limb, glabrous, black, endocarp 2-grooved on dorsal, 
I-grooved on ventral surface. 


Var. 6, capitellata, Clarke. JW’. cafitellata, Wight, Ic. t. 1022. 
Fruit rather smaller and more compressed. | 


Rubtacee. 289 


Upper montane zone; common(?). Horton Plains; Maturata; 
Adam’s Peak. Var. 8, Adam’s Peak (Fl. B. Ind.). Fl. April, white. 
Also in the mountains of India and Malaya. 


2. V. erubescens, Wall. P/. Astat. Rar. ii. 29 (1830). 

Thw. Enum. 136. C. P. 1636. 

Fl. B. Ind. iii. 7. Wight, Ic. t. 1024 (V. Wiehtianum). Wall. 1. c. 
t. 134. 

A small slender tree with few virgate branches, bark 
smooth, yellowish, buds stellate-pubescent; 1. 2-4 in, oval, 
acute or rounded at base, acuminate, acute at apex, sharply 
dentate-serrate in upper half, glabrous above, stellate-hairy 
on veins beneath; thin, lat. veins 4 or 5 on each side, very 
oblique, conspicuous, petiole $—? in., sparsely stellate-hairy; 
fl. on short ped., in small, slightly drooping, racemose cymes, 
terminating short lateral branchlets; cal. glabrous, segm. 
lanceolate, cor.-lobes rounded, spreading or recurved; stam. 
not exserted; fruit 4 in., oblong-ovoid, much. compressed ; 
glabrous, red. 


Upper montane zone; rare. Sita Eliya, near Hakgala; Elephant 
Plains (Gardner); Horton Plains (Thwaites). Fl. May; pale pinkish- 
white or cream-coloured. 

Also in the Himalaya and Nilgiri Mountains. 

A very pretty shrub when in blossom. The leaves have a strong 
pungent odour when bruised. The wood is said to be very hard and close- 
grained. 


We have no species of Lowicera in our mountains, though 2 occur in 
the Nilgiris. 


LXVII.—RUBIACE/, 


HERBS, shrubs or trees, rarely climbing, |. opposite or whorled, 
simple, entire (sometimes crenate in WMeurocalyx), with inter- 
petiolar stip. often connate and forming a sheath (absent in 
Galieg); fl. regular, bisexual or rarely unisexual, often 
dimorphic ; cal.-tube adnate to ov., free limb usually more or 
less developed, rarely 0; cor. varying from long-tubular to 
rotate, lobes 4-9, usually 5 (2-lipped in Dzchzlanthe), valvate 
or imbricate or twisted in estivation; stam. as many as cor.- 
lobes, inserted in throat or tube of cor.; anth. distinct (very 
rarely connate); ov. inferior, 1-10- usually 2-celled, crowned 
with epigynous disk, ovules solitary or many (rarely 2) in each 
cell, erect or horizontal or pendulous, style usually simple, 
PART IL. 


290 Rubtacee. 


stigma entire or lobed; fruit a capsule, a berry or a drupe, 
2—10-celled, or of 2-4 dehiscent or indehiscent cocci; seeds 


2—numerous, embryo various, in fleshy or horny endosperm. 


A very large Tropical Family of which 138 species are here included ; 
particularly characteristic of the wet region. In the dry country only 31 
species occur, and of these 11 only are restricted to it; Adama and 
Scyphiphora (the latter a seashore plant) being the only genera so circum- 
scribed. In our moist region, on the other hand, are found 107 species, 
and 77 of these do not extend beyond it. These ‘latter are pretty equally 
dispersed over the low country and the hills, and 42 are confined to the 
former and 35 to the latter. To the moist low country are restricted the 
genera Uncaria, Fergusonia, Schizostigma, Byrsophyllum, Nargedia, 
Scyphostachys, Guettarda, Dichtilanthe, and Geophila, and to the 
montane zone the genera Al/wophania and Acranthera. 

Nearly half of our species, as many as 68, are endemic, and four 
genera are peculiar to Ceylon: Lezcocodon, Schis zostigma, Nargedia, and 
Scyphostachys. 

Ovules more than 2, usually numerous, in each cell. 
Fl. crowded in dense globose heads (Vauclea). 


Cal.-tubes fused into a fleshy mass I. SARCOCEPHALUS. 
Cal.-tubes distinct. 
Trees. 
Cal.-segm. linear 2. ANTHOCEPHALUS. 
Cal.-segm. clavate 3. ADINA. 
Cal.-segm.o . ; : : 4. STEPHEGYNE. 
Cal.-segm. oblong, appendaged 5. NAUCLEA. 
Climber with hooked peduncles . 6. UNCARIA. 
Fl]. not in dense globose heads. 
Fruit a capsule. 
Cor.-lobes twisted in bud . A ‘ . 7. WENDLANDIA. 
Cor.-lobes valvate in bud. 
Cor.-lobes with a tooth on each side =) 0. DEN mEIAe 
Cor.-lobes entire. 
Cal.-segm. 5. 
Cal.-segm. large, membranous; cap- 
sule dry, not compressed. . 9. NEUROCALYX. 
Cal.-segm. small; capsule coriaceous, 
much compressed . ; : . 15. OPHIORRHIZA. 
Cal.-segm. 4, small 
Cal.-segm. contiguous. : . 12. HEDYOTIS. 
Cal.-segm. distant. 
Seeds numerous . é : . 13. OLDENLANDIA. 
Seeds few  . ; : : . 14. ANOTIS. 
Fruit a fleshy berry. 
Cor.-lobes valvate (Zussendea). 
Inflorescence terminal, cymose. 
One cal.-segm. often developed into a 
large leaf, rest deciduous . : . 16. MUSSENDA. 
Cal.-segm. equal, persistent . : 17. ACRANTHERA. 
Inf. terminal, capitate, surrounded by 
large involucre 0 ‘ . 18. LEUCOCODON. 
Infl. axillary; ov. s-celled. 
Shrubs . ; 5 ; j ; . 19. UROPHYLLUM. 


Creeping herb 5.) 20 SCHIzosmremnas 


Rubiacee. 


Cor.-lobes twisted (Gardenia). 
Inf. terminal or leaf opposed. 
Fl. dicecious . d 
Fl. bisexual. 
Ov. 2-celled. 
Seeds few 
Seeds numerous 
Ov. 1-celled 
Infl. axillary (also sometimes in n 23). 
Endosperm ruminate 
Endosperm not ruminate. 
Fl. in bracteate spikes 
Fl. in clusters 
Ovules 2 in each cell (see also 12 and 27) 
Ovules solitary in each cell (see also 14). 
L. with stipules. 
Fruit indehiscent, usually fleshy. 
Cor.-lobes valvate (or rarely imbricate). 


Oy. 2-celled (spuriously 4-celled in 37). 


Corolla 2-lipped 
Corolla regular. 
Ovules pendulous 
Ovules on the dissepiment 
Fl. in a confluent head . 
Fl. not confluent 
Ovules basilar, erect. 

Trees or shrubs; infl. cymose. 
Cor.-tube short, straight 
Cor.-tube rather long, curved 
Cor. funnel-shaped 

Herb; infl. umbellate 

Oy. 4- or more-celled. 
Cor.-lobes imbricate 
Cor.-lobes valvate. 

Ov. 12-celled 
Ov. 4-celled; stip. connate 


Ov. 4- (or 5- -) celled ; stip. not connate 


Cor.-lobes twisted (Loree). 
Fl. in copious cymes. 


Style longer than cor.; stigma 2-lobed 
Style twice as long as cor.; stigmaentire 


Fl. solitary or in threes 
Fruit aul dehiscent septicidally. 


Carp. 2, very small, spares from central 


axis, indehiscent 
CArp.. 2, 
dehiscent . 


Carp. 2, large, much compressed, tardily 


separating 
Carp. 4, indehiscent 
L, without stip., whorled (Galea), 
Corolla campanulate 
Corolla rotate . 


very small, one or both ventrally 


oie 
5 BB 


5 Bike 
4 Role 


=) 39: 
. 40. 
5 ANG, 
SAE 


20: 


3 oh 
“| 10: 


42. 


34. 
35. 
5 By6y 


e 291 


. BYRSOPHYLLUM. 


. WEBERA. 
. RANDIA. 
. GARDENIA. 


. NARGEDIA. 
. SCYPHOSTACHYS. 


. DIPLOSPORA. 
. SCYPHIPHORA. 


DICHILANTHE. 
CANTHIUM. 


MORINDA. 
PRISMATOMERIS. 


PSYCHOTRIA. 
CHASALIA. 
SAPROSMA. 
GEOPHILA. 


GUETTARDA. 


TIMONIUS. 
ALLG@OPHANIA. 
LASIANTHUS. 


IXORA. 
PAVETTA. 
COFFEA. 


. KNOXIA. 
. SPERMACOCE. 


. HYDROPHYLAX. 
. FERGUSONIA. 


. RUBIA. 
. GALIUM. 


292 ; Rubiacee. [Anthocephalus. 


1. SARCOCEPHALUS, 4 /zclius. 


Tree, stip. large, fl. small, sessile, in dense globose terminal 
heads, without bractlets, the cal.-tubes fused into a fleshy 
mass; cal.-segm. 4 or 5, clavate, persistent ; cor.-tube long, 
lobes 4 or 5, imbricate; stam. 4 or 5, inserted in mouth of 
cor.; ov. long, 2-celled, with numerous imbricated ovules; 
fruits sunk in a fleshy mass, 2-celled, seeds few, small, 
oblong.—Sp. 8; 3 in FZ. B. Ind. 


S. cordatus, Mig. Fl. Ind. Bat. ii. 133 (1856). Bakmi, S. 


Vammi, 7. 
Fl. Zeyl. n. 53. Vauclea orientalis, L. Sp. Pl. ed. 2, 243. WV. macro- 


phylla, Roxb. (?), Moon Cat. 14. lV. cordata, Roxb. Fl. Ind.i. 509. JV. 
coadunata, Roxb., Thw. Enum. 137. C. P. 1658, 781. 

FI. B. Ind. iii. 22. Bedd. Fl. Sylv. t. 318. 

A small tree, bark silvery-grey, smooth, branches marked 
with prominent leaf-scars, young parts minutely stellate- 
pubescent; 1. large, 4-8 in., broadly ovate-oval, slightly caudate 
at base, rounded or very obtuse at apex, entire, undulate, 
glabrous above, glabrous or finely pubescent beneath, petiole 
3-1 in., stip. $-1 in., obovate-rotundate, slightly connate, 
deciduous ; fl. very numerous, fused together by their fleshy 
cal.-tubes, heads 2-1 in. diam., on stout peduncles; cal.-segm. 
small, club-shaped, cor.-tube long, lobes acute ; stam. included; 
style very long, much exserted, stigma large, clavate ; head 
of fruit about 1 in., globose, solid, fleshy ; seeds slightly rough, 


black. 
Low country in both moist and dry regions; rather common. FI. 


May, June; pale yellow, styles white. 

Also in Malaya, Philippine Is., and perhaps N. Australia, but not in 
Peninsular India. 

Hermann’s drawing (there is no specimen) is certainly this ; which 
fixes Linneus’s Wauclea orientalis as originally the same, but he also 
quotes Hort. Malab. iii. t. 33, which rather represents Anthocephalus 
Cadamba. Roxburgh’s lV. cordata was raised from Ceylon seed sent to 
Calcutta in 1802. 

The leaves and flower-heads of the dry country form (C. P. 781) are 
smaller than those from the moist region, and the 1. more pubescent 
beneath. The head of fruit is succulent, and is eaten, hence the tree 
is often called ‘Kana-bakmi.’ Wood yellow, light, rather soft. 


2. ANTHOCEPHAULUS, 4. fich. 


Tree, stip. large, fl. sessile, in dense globose terminal heads, 
the cal.-tubes closely pressed together but not fused, without 
bractlets; cal.-segm. 5, linear, persistent; cor.-tube funnel- 


Adina.] Leubracee. 204 


shaped, long, lobes 5, imbricate; stam. 5, inserted at mouth 
of cor.; ov. 2-celled below, 4-celled above, with numerous 
ovules in each cell, style long, stigma large, ovoid, fruit cap- 
sular, narrowly turbinate, truncate, pericarp thin below but 
hard and 4-lobed above ; seeds numerous, very small, angular, 
not winged.—Sp. 3 or 4; 1in Fé. B. Ind. 


A. Cadamba, M79. F?. Ind. Bat. ii. 135 (1856). Bmbul-bakmi, 
S. Wellaikadampa, 7. 

Nauclea orientalis, Moon Cat. 14 (non L). Vauclea Cadamba, Roxb., 
Thw. Enum. 137. C. P. 1659. 

Hilee iad. i-23. Bedd. FI. Sylv. t. 35. 


A large tree with erect trunk and horizontal branches, 
young parts pubescent; |. large, 6-11 in., oblong-lanceolate, 
acute or rounded at base, acuminate, acute at apex, glabrous 
and shining above, pubescent on the veins beneath, petiole 
I-I}in., stip. about 4in., narrowly lanceolate, acute; heads 
14-1}in.,on short stout peduncles; cal.-segm. obtuse, pilose 
at ends; cor.-lobes erect, lanceolate, acute; capsule about 
4in., compressed, angular, closely packed but scarcely con- 
fluent, crowned by long persistent cal.-segm.; pericarp trans- 
parent in lower part; seeds muriculate. 

Low country up to 2000 ft., Thwaites says ‘common,’ but I have only 
seen specimens from Kurunegala, Alagalla and Pusselawa. F1.September ; 
orange-yellow, styles white, fragrant. 

Also in India, burma, Sumatra, Borneo. 

The little capsular fruits are not confluent as generally described, 
though in very close contact with one another. 

Thw. gives ‘Hélamba’ as S. name for this ; it more properly belongs 
to Stephegyne tubulosa, but may be used for both. 


3. ADINA, Salisé. 


Large deciduous tree, stip. large, fl. small, sessile, crowded, 
not fused, in globose, axillary heads, with bractlets; cal.-segm. 
5, clavate; cor.-tube long, lobes 5, short, valvate; stam. 5, 
inserted at mouth of cor.; style very slender, stigma globose; 
capsule small, 2-celled, truncate, readily dehiscent into 2 
valves; seeds numerous, tailed at both ends.—Sp. 6; 4 in 
Fl. B. Ind. 


4. cordifolia, Hk. f, in Gen. Pi. ii. 30 (1873). WBMolon, S. 
Manchalkadampa, 7. 

Neuclea cordifolia, Moon Cat. 14; Thw. Enum. 137. C. P. 1660. 

Fl. B. Ind. iii. 24. Bedd. Fl. Sylv. t. 33 (Vauclea). 


A very large tree with erect trunk and horizontal branches, 


204 Rubwacee. [ Stephegyne. 


bark brownish-grey, thick, soft, furrowed, young parts pu- 
bescent; 1. closely placed, 34-44 in., very broadly ovate, 
cordate at base, acuminate, acute at apex, glabrous above, 
more or less softly pubescent beneath especially when young, 
minutely reticulate, petiole long, 2-4 in., pubescent, stip. 
oblong-oval, obtuse, pubescent ; heads about ?in., peduncles 
long, slender, usually in threes from axils of young L., jointed 
a little below the heads, fl. very numerous, bractlets clavate; 
cal. hairy inside and out, segm. club-shaped; cor.-tube slender, 
dilated above, pilose, lobes subacute, recurved; style much 
exserted ; capsule very small, } in., turbinate, very hairy. 

Dry region; rather common, extending into intermediate region, é.g., 
Kurunegala. Fl. July; yellow 

Also in India and Burma. 

This is quite deciduous for a short time. A form with thinner leaves, 
more cordate and with fewer longer hairs beneath, occurs at Anuradha- 
pura. Wood rather hard and heavy, smooth, pale yellow, durable. 


4. STEPHEGYNE, Korth. 


Trees, stip. large, fl. sessile, crowded but not fused in 
terminal globose heads; cal.-limb tubular, truncate, segm. 0; 
cor. tubular, lobes 5, valvate, acute, recurved ; stam. inserted at 
mouth of cor.; ov. 2-celled, with numerous ovules in each cell, 
style long, stigma large, oblong-ovoid; capsule .2-celled, 
septicidally dehiscent; seeds numerous, small, winged.—Sp- 
LOR esuin lBel a7: 


IL. under 3 in.; cal.-limb short . 3 - E . I. S. PARVIFOLIA. 
L. over 4 in.; cal.-limb long and tubular . , . 2. S. TUBULOSA. 


1. S. parvifolia, Korth. in Verh. Gesch. Nat. Bot. 161 (1843 ?). 
Chélampai, Nirkadampa, 7. 


Nauclea parvifolia, Roxb., Thw. Enum. 137. C. P. 780. 

F). B. Ind. iii. 25. Bedd. Fl. Sylv. t. 34. Wight, Ill. t. 123. 

A moderate-sized tree, bark rather smooth, whitish-grey 
thick, flaking off, young parts glabrous; 1. 23-3 in., rotundate 
or broadly obovate-oval, acute or rounded at base, very 
obtuse or rounded at apex, entire, glabrous on both sides 
save for small tufts of hair in vein-axils beneath, thin, bright 
apple-green, pale beneath and minutely pellucid-reticulate, 
petiole under din., stip. $in., oblong-spathulate, very obtuse 
pink; heads #in., shortly pedunculate, cal.-limb very short ; 
cor.-tube three times as long as acute recurved lobes, head of 
fruit 2in., capsules crowded but quite separate, oblong with 


blunt rounded tops marked with line of the cal.-limb, with 
10 blunt vertical ribs. 


Nauclea.] Ru biacee. 205 


Dry region; rather rare. Trincomalie; Mineri; Anuradhapura; 
Puttalam. FI. April, July, August; greenish-yellow, scented, styles white. 

Also in India and Burma. 

Wood hard, heavy, smooth, pinkish-yellow, durable. 


2. S. tubulosa, Hk. f 77 Gen. Plant. 11. 31 (1873). Wélamba, S. 

Nauclea tubulosa, Arn. in Thw. Enum. 137. JV. parvifolia, Moon 
Gat 14 (non Roxb.). C. P./1657. 

Fl}, B. Ind. iii. 25. Bedd. Ic. Fl. Ind. or t. 18. 


A small tree, bark smooth, pale brown, young parts slightly 
pubescent; |. 4-6 in., broadly ovate, rounded or subcordate at 
base, acuminate, acute at apex, glabrous on both sides or 
pubescent on veins nea thin, lat. veins numerous, very 
prominent beneath, intermediate venation minutely reticulate, 
pellucid, petiole 4—{in., stip. very large, 1 in., broadly oval, 
very obtuse, veined, strongly keeled; heads usually terminal 
and sessile, sometimes axillary and pedunculate, 14 in., fl. not 
densely crowded, bractlets numerous, spathulate; cal.-limb 
long tubular; cor.-tube about twice as long as cal.-limb, hairy 
within, lobes half as long as tube, linear-oblong, acute; stvle 
much exserted, stigma oblong, truncate; capsule 3 in., ovoid- 
oblong, crowned by persistent cal.-limb, slightly puberulous, 
10-ribbed; seeds long. 


Var. 8, minor, 7iw.l.c. C. P. 1656. 
L. smaller, 2-3 in.; fl.-heads smaller, ? in. 


Low country in the moist region; rare. Colombo; Kalutara (Moon) ; 
Hiniduma; Kukul Korale, abundant (Thwaites). Var. 6 in the dry 
country. Jaffna (Gardner); Anuradhapura (Brodie); Haragama. FI. 
May and (var. 8) Feb.; pink. 

Also at Cochin, S. India. 

Wood as in the last. 


5. NAUCLEA,* ‘Z.’ 


Tree, stip. large, fl. sessile, crowded in globose terminal 
heads, not fused; cal.-segm. 5, oblong, tipped with a small 
process; cor, tubular-funnel-shaped, lobes 5, imbricate, re- 
curved, acute; stam. inserted at mouth of cor.; ov. 2-celled 
with numerous ovules in each cell on pendulous placentas, 
style long, exserted, stigma globular; capsule 2-celled, dehis- 
cent; seeds numerous, winged.—Sp. 30; 5 in Fv. B. Jud. 


‘4 


* Linnzeus’s genus /Vauclea consisted of one species only, JV. orzentalis, 
which, as above shown, is Sarcocephalus cordatus. \t does not appear 


upon what principle it can be now employed for a different one. 
° 


29 6 Rubtwacee. [ Uncaria. 


N. zeylanica, 2. 7. zz Fl. B. Ind. iii. 26 (1880). 

NV. triflora, Moon Cat. 14. NW. peduncudaris, Thw. Enum. 137 (non 
Wall). C. P..2820. 

Fl. B. Ind. ii1. 26, Bedd. Ic. Pl. Ind. or t. 235. 

A moderate-sized tree, branchlets slender, young parts 
elabrous; |. 3-5 in., lanceolate, tapering to both ends, some- 
what acuminate, obtuse or subacute at apex, glabrous on 
both sides, petiole short, under } in., stip. oblong, rounded at 
apex; heads $—?in., peduncles 1-14 in. long, usually in threes 
at end of branchlets, slender; cal.-segm. obtuse, pubescent, 
with a small appendage at apex; cor. silky-pubescent, lobes 
oval, apiculate; style twice the length of cor.; capsule (not 
seen) short, many-seeded. 

Moist low country; rare. Adam’s Peak (Moon); Sabaragamuwa; 
Balangoda, abundant (Thwaites) ; Uma-oya. Fl. May, June; dull orange. 

Endemic. 

This is very close to WV. peduncularis, Wall. of Sumatra, to which 
Thw. refers it. If distinct, Moon’s name is the oldest, and, though with- 
out description, should perhaps be adopted in preference to Hk. f.’s. 


6. UNCARIA, Schred. 


A shrub, climbing by hooked peduncles, stip. large, fl. 
small, in dense pedunculate axillary heads; cal.-limb very 
short, segm. 5, triangular, obtuse; cor.-tube very slender, 
lobes 5, valvate; stam. 5, inserted at mouth of cor.; ov. 
2-celled, with numerous ovules on large placentas, style long, 
exserted, stigma large, oblong; fruit an elongated capsule, 
septicidally dehiscent, valves 2, chartaceous; seeds minute, 
numerous, compressed, with very long membranous tails at 
each end.—Sp. about 30; 19 in FAY. B. Jud. 


U. dasyoneura, Korth. Verh. Nat. Ges. Bot. 169 (1839). Var. 
Thwaitesii, Wk. f 7x Fl. B. Ind. |. c. 

U. Gambier, Thw. Enum. 138 (non Roxb.). C. P. 1661. 

Fl. B. Ind? m1. 31., Bentl, and Trim. Med) Plot: 130, fa7eangee 
(fruit and seed only. U. Gambier). 

A very large wide-climbing shrub; stems stout, bluntly 
quadrangular, compressed below each node when young, 
glabrous, dark green, internodes long, young parts glabrous; 
l. very numerous, persistent, spreading or deflexed, 3$—44 in., 
narrowly ovate or oblong-oval or oblong-lanceolate, rounded 
at base, suddenly caudate-acuminate, often twisted at apex, 
glabrous and shining above and beneath except for little tufts 
of hair in axils of lat. veins, coriaceous, dark green above, paler 
beneath, lat. veins 5-7 on each side, slender, conspicuous, 


Wendlandia.]| Rubtacee. 297 


unbranched, prominent beneath, and connected by transverse 
venation, petiole 2-3 in., stip. about 4in., oblong-oval, obtuse, 
entire; fl. numerous, nearly sessile, head #-1 in., globose, 
receptacle hairy, peduncle of two parts separated by a pair of 
bractlets, upper part slender, cylindrical, pubescent, lower 
part hard and woody, laterally compressed, tapering, curved, 
glabrous, becoming after the fall of the fruit much enlarged, 
falcately hooked and very rigid; cal. woolly, segm. very short; 
cor.-tube 2in., hairy outside, lobes obtuse, glabrous within ; 
capsule #in., stalked, narrowly ovoid, much tapering at both 
ends, crowned with persistent cal.-limb, slightly hairy, valves 
chartaceous, flat, separating from cal., which remains entire 
at top; seeds with very long membranous tails, 2 at one end 
and I at the other. 

Moist low country up to 3000 ft.; rather rare. Kandy; Hantane; 
Deltota; Ambagamuwa; Morowak Korale; Kaduganawa; Colombo. 
Fl. March; yellowish-green. 

The type in the Malay Peninsula, but our var. endemic. 

This differs from U. Gambier, Roxb., of Malaya, which affords the 
Gambir or ‘Terra japonica’ of commerce, in its shorter cal.-segm. and 
the want of tufts of hair inside the cor.-lobes: I have also ascertained 
that the extract to be obtained by boiling the leaves of our plant does not 
resemble the commercial product from that species (which is figured in 


* Medicinal Plants,’ t. 139 exclusive of f.7 and 8); but they are very closely 
allied. 


The leaves when dried are a dark coppery-brown. 


7. WENDLANDIA, Lari. 


A small tree, |. often in whorls of 3, stip. small; fl. small 
in large terminal panicles; cal.-segm 5, small; cor. tubular 
below, then recurved, shortly 5-lobed, lobes twisted in bud; 
stam. 5, inserted between cor.-lobes, anth. almost sessile; ov. 
globose, 2-celled, with numerous ovules on large placentas, 
style long, stigma large, bilobed, slightly exserted; fruit cap- 
sular, very small, globose, 2-celled, 2-valved; seeds numerous, 
very minute, flat—Sp. 16; 14 in FZ. B. Ind. 

W. Notoniana, Wall. in W. and A. Prod. 403 (1834). Rawan- 
idala, S. 

W. bicuspidata, W. and A. Prod. 403. Thw. Enum. 159. C. P. 315. 

Fl. B. Ind. iii. 40. Wight, Ic. t. 1033. Bedd. Fl. Sylv. t. 224. 


A small tree or large shrub with erect or ascending 
branches, bark orange-red coming off in fibrous strips, 
branchlets densely pubescent, buds very silky; 1. usually 3 
at a node, 4-5 in., lanceolate-oval, much tapering to base, 
acuminate, acute at apex, slightly pilose above, pubescent 


298 Rubracee. [ Dentella. 


beneath, undulate, rather thick, bright light green, paler 
beneath, with the prominent: veins pink, petiole very short, 
pubescent, stip. barely + in., linear-lanceolate, acute, con- 
duplicate, sometimes bifid; fl. sessile or nearly so, very 
numerous, rather close on the branches of dense pubescent, 
pyramidal terminal panicles 6-12 in. long, bracts small, seta- 
ceous; cal. glabrous or nearly, segm. acute; cor.-tube about 
; in., hairy within; capsule capped with cal.-segm., splitting 
from above loculicidally about half way down. 
Var. 8, zeylanica, (7h. f. in Fl. B. Ind., ¢.c. 


L. nearly glabrous; fl. pedicellate, distant, panicles lax, 
drooping; cor.-tube longer. 

Open places in the moist region from sea-level up to all elevations; very 
common. Var. @, rare. Fl. Feb., March; white. 

Also in Peninsular India. 

The young leaves are a delicate purplish-pink. The calyx is usually 
glabrous or nearly so in Ceylon. Neither of the figures quoted are 
characteristic of this very common plant, which, curiously, does not seem 
to have been seen by Hermann. 

Wood rather heavy, hard, strong, brownish-red. 


8. DENTELLA, forest. 


A small prostrate annual, fl. solitary, axillary ; cal-limb 
tubular, segm. 5; cor. funnel-shaped, lobes 5, short, toothed 
on each side, valvate; stam. 5, inserted low down in tube, 
included; ov. globose, 2-celled, with numerous ovules on large 
placentas, style very short, stigmas 2, long, filiform; fruit 
small, globular, dry, indehiscent, 2-celled ; seeds numerous, 
minute, angular.—Monotypic. 


D. repens, forst. Char. Gen. 26 (1776). 

Thw. Enum. 144. C. P. 1683. 

Fl. B. Ind. ii. 42. Rumph. Herb. Amb. v. t. 170, f. 4 (poor). 

A slender prostrate herb, stems long, slender, much 
branched, rooting at the nodes; |. very small, 2 in. at most, 
lanceolate or subspathulate, tapering to base, acute, glabrous 
or with a few bristly hairs on margin, petiole obscure, stip. 
truncate, membranous; fl. solitary often in the forks of the 
stem, nearly sessile, cal.-limb persistent, segm. very acuminate ; 
cor.-lobes ovate, acute, with a tooth on each side; fil. very 
short; fruit about $in., crowned by large persistent cal.-limb, 
densely villous with coarse hair. 

Low country, a weed in moist places; rather rare. Colombo; Hara- 


gama; Peradeniya. Fl. Aug., Sept.; white. 
Throughout the Tropics of Asia, Polynesia, and N. Australia. 


Neurocalyx.| Rubtacee. 299 


9. NEUROCALYWX, Hook. 


Small perennial herbs, |. crowded, stip. large, usually 
divided; fl. rather large, on drooping ped. in axillary racemes; 
cal.-segm. 5, large, coloured, membranous, reticulately veined, 
persistent ; cor. small; rotate without a tube, lobes 5, valvate; 
stam. 5, fil. very short, anth. connate into a conical tube; ov. 
2-celled with very numerous ovules in each, style simple, 
stigma terminal; fruit crowned with large persistent cal.- 
segm., dry, bursting irregularly; seeds very numerous, globose, 
muriculate——Sp. about 8; 5 in FZ, 5. Lnd. 


L. glabrous above. 
Stip. divided into setaceous segm. 
Stip. ovate, often bifid . 

L. hairy on both sides. 
Cor.-lobes 4 as long as cal. . 
Cor.-lobes as long as cal. 


. N. ZEYLANICUS. 
. N. WIGHTII. 


| 


iS) 


. N. GARDNERI. 
. N. CHAMPIONII. 


a SSS) 


1. N. zeylanicus, Hook. Jc. Plant. t. 174 (1837). 

Am. in Ann. Nat. Hist. iii. 21. Thw. Enum. 138. C. P. 286. 

Hib. ind. ii. 47. le. Plant. t. 174. 

Stem 3-4 in., slender, erect, ringed with stipular scars; 
1. closely placed, 4-7 in., linear-lanceolate, much tapering 
below, acuminate, acute, slightly undulate, crenate, glabrous 
above, paler and pubescent on the prominent veins beneath, 
petiole 4—$in., pubescent, purple, stip. green, divided almost 
to base into setaceous, rather rigid, spreading segm.; fl. on 
rather long pubescent ped., racemes, erect, few-flowered, lax, 
shorter than 1|., bracts linear-setaceous, rather shorter than 
ped., cal.-segm. 4in., lanceolate, acuminate; cor.-lobes about 
half as long as cal.-segm., acute. 


Moist region up to 4000 ft.; rare. Diyabetene, Sabaragamuwa ; 
Opatte, S. Prov.; Adam’s Peak (Walker). Fl. March; white. 
Endemic. 


2. N. Wightii, Avz. in Ann. Nat. Hist. iii. 22 (1839). 
Thw. Enum. 138. C. P. 595. 


FI. B. Ind. iii. 47. Wight, Ic. t. 52 (VV. Hookeriana). f 

Stems 3-8 in., semi-woody below, stout, marked with leaf- 
scars, the internodes often developed; 1. large, 6-12 in., lance- 
olate, very much tapering to base, acuminate, acute, entire, 
glabrous above, paler and pubescent on the veins beneath, 
petiole obscure, stip. very large, foliaceous, ovate, acuminate; 
often with a filiform point, sometimes separated into their 
constituent pairs; fl. on pubescent ped., crowded, racemes 
dense, very much shorter than l., bracts large ovate, acumin- 


300 Rubtacee. [Meurocalyx. 


ate; cal.-segm. about 4 in. (enlarged in fruit), ovate, acuminate; 
cor.-lobes rather more than half as long, very acute; fruit 
pubescent. 

Damp shady places in forests in moist low country up to 2000 ft.; 
rather common. Kukul Korale; Hiniduma; Ambagamuwa; Deltota; 
Dolosbagie. Fl. March, April; very pale pinkish-violet. 

Also in S. India. 

The above description covers /V. capztata, Benth. in Fl. B. Ind. ili. 47 
(where it is considered a distinct species). This appears to be the 
commonest form, but cannot be separated. JV. Hookerzana, Wight, is a 
small form merely. 


3. N. Gardneri, 7iw. Enum. 139 (1859). 

Ghw-) Enum ngo) (Cee sroz1. 

Fl. B. Ind. iii. 47. 

Stem 2-4 in., stout, densely hairy; |. 6-9 in., oval or 
obovate-oval, tapering to base, subacute or very shortly 
acuminate, entire, thin, hairy on both sides, petiole very 
short, stip. split, the halves ovate, acuminate, densely pubes- 
cent; fl. on short ped., racemes villous-hairy, bracts lanceolate, 
acuminate, 2-toothed near base; cal.-segm. broadly ovate, 
acuminate, hairy outside, cor.-lobes about 4 length of cal.- 
segm., oblong-lanceolate; fruit villous-hairy. 

Damp forests in moist low country; very rare. Pasdun Korale 
(Gardner); Hewesse (Thwaites). Fl. Aug. 

Endemic. 

I have not met with this living. Much the most hairy of our species. 


4. N. Championii, enth. in Thw. Enum. 139 (1859). 

Thw. Enum. 139. C. P. 3417, 3480. 

Fl. B. Ind. iii. 47. 

Stem very short, 1-3 in. slender, closely ringed with 
stipular scars; 1. very closely placed, 3-6 in., obovate-oblong, 
tapering to base, rounded or obtuse at apex, often somewhat 
crenate, hairy on both sides (becoming glabrous above), 
usually very bullate above, thin, petiole #-14in., stip. deeply 
cut into 2 lanceolate acuminate segm.; fl. on slender hairy 
ped., racemes lax, few-flowered, bracts lanceolate, as long as 
ped., with 2 very small lateral lobes; cal.-segm., about 4 in., 
ovate, acute; cor.-lobes equal to cal.-segm., linear-oblong, 
fruit hairy. 

Damp forests in moist low region; rather rare. Kukul Korale; 
Singhe Raja Forest; Nillowe Kande; Bambarabotuwa; Hewesse. FI. 
Mei April, and September ; calyx bright pink or white, corolla white. 

ndemic. 


Very like an Acrotrema in habit and foliage. Best distinguished 
by its bullate leaves. 


Alleophania. | Rubtacee. 301 


10. ALLGOPHANIA, 7iw. 


A weedy under-shrub, stip. large, connate, membranous, 
fl. small, dimorphic, crowded in dense axillary whorls; cal.- 
limb tubular, membranous, segm. 4, linear-setaceous, persistent ; 
cor. funnel-shaped, lobes 4, valvate; stam. 4; ov. 4-celled, 
with 1 erect ovule in each cell, style simple, stigmas 4, long; 
fruit small, turbinate, enclosing 4 small bony 1-seeded 
pyrenes.—Sp. 2 or 3; 2 in FZ. B. Jud, but none in Indian 
Peninsula, 


This genus is difficult to place in the Order; it has affinities with 
Lasianthus as well as with Hedyotzs. 


A. decipiens, 7/w. Enum. 147 (1859). [PLATE XLVIII.] 

C. P. 3093. 

Fl. B. Ind. ili. 48. 

A coarse rigid perennial, stems numerous, 2-4 ft. high, 
semi-woody below, much divaricately branched, sub-quad- 
rangular, adpressed-pubescent; |. 2-24 in., lanceolate or oval- 
lanceolate, acute at base, acuminate, acute, glabrous or nearly 
so and bullate above, pilose silky or pubescent on the very 
prominent veins beneath, especially when young, petiole 1-3 
in., stip. connate, forming a tube, truncate, membranous, with 
branched bristly hairs, at length splitting up and deciduous ; 
fi. sessile, very crowded in dense opp. axillary clusters forming 
whorls, cal.-limb long, segm. shorter than tube, linear, acu- 
minate, bristly-hairy; cor. hairy at the mouth, lobes more 
than half as long as tube, lanceolate, acute, recurved, stam. 
erect; style hidden or exserted, stigma 4-fid; fruit small, 
flat-topped, crowned with cal.-limb and often connivent segm., 
pyrenes very small, triangular, smooth, yellow. 


Var. 6, lavescens, 7iw. Enum. l.c. C. P. 3094. 


L. oval or subrotundate, rounded or subcordate at base, 
the margin strongly revolute, veins much more hairy, often 
villous beneath. 


Var. y, Arnottii, Hz. f ix Fl. B. Ind. \.c. (sp.). Hedyotis nodulosa, 
var. a, Thw. Enum. 143. C. P. 87. 


L. larger, acuminate at both ends, nearly glabrous, on 
longer petioles. 

Montane zone from 3000 ft. upwards, especially at the higher eleva- 
tions; very common. FI. Feb.—Aug.; white or bluish, anth. violet. 

Endemic. 

This bears so strong a resemblance to Hedyotzs nodulosa (with which, 
indeed, it was mixed even by Thwaites) that it can be distinguished with 
certainty only by examining the ovary or the fruit-structure. It is very 


302 Rubiaceae. [ Fergusonia. 


variable in respect to size, hairiness, width of leaves, and length of 
petiole, but I cannot see a second species among the forms, as in Fl. 
B. Ind. 

The flowers are very liable to attack by some insect, which causes the 
corolla to become an enlarged hard inflated gall. 


1. FERGUSONIA,* HZ. 7 


A prostrate herb, stip. rigid, ciliate; fl. small, solitary 
sessile, axillary; cal.-segm. 4, acuminate; cor.-tube tubular- 
funnel-shaped, with a ring of hair within, lobes 4, valvate; 
stam. 4, inserted between cor.-lobes, fil. very short ; epigynous 
disk of 4 prominent lobes; ov. 4-celled wzth 1 erect ovule in 
each cell, style simple, slightly exserted, stigmas 2, filiform ; 
fruit of 4 separable membranous indehiscent cocci—Monotypic. 


A very anomalous plant of doubtful position in the Order. 


FE. zeylanica, 72. 7. in Ic. Plant. xii. 23 (1876). 

Borreria tetracocca, Thw. Enum. 442. /. Thwatteszz, Hk. f. in Gen. 
Plant. 1129 C3 O50: 

Fl. B: Ind. i. 48." Hook. Ic. Plit. 1124. Bedd. Ic Ply Indontrge 
(Borreria tetracocca). Bork, "S? 


Annual herb, stems procumbent or ascending, rooting at 
the lower nodes, 1-24 ft. long, slightly branched, finely ciliate 
on the angles, pinkish; 1. #-14in., narrowly oblong-lanceolate, 
subsessile, tapering to base, acute, glabrous above, sparingly 
bristly-hairy beneath, finely spinous-ciliate on edge, stip. 2 in., 
lanceolate-linear, setaceous, rigid, strongly ciliate, persistent ; 
fl. very small, sessile, solitary; cal.-segm. lanceolate, acute, 
ciliate with coarse bristly hairs; cor.-lobes oval, acute, with a 
few hairs on the back; fruit-cocci very small, pyriform, bristly 
on top, readily separating, and each bearing on top a single 
persistent cal.-segm. 


Moist low country in dry places; apparently rather rare. Near 

Colombo (Ferguson); Labugama; Karawita. FI. Sept.—Jan.; very pale 
ink. 

, Also found in Coorg. 

Of Sir J. Hooker’s two names, /. 7hwazteszZ is apparently the older, 
but he uses /. zey/anica in F1. B. Ind. 

This insignificant but curious plant much resembles Hedyotis nztida 
and also Spermacoce hispida and S. stricta in habit. 


* Commemorates the late W. Ferguson, F.L.S., of Colombo, for many 
years a diligent and very successful investigator of the flora of Ceylon, 
who died July 31st, 1887. 


Hedyotis.| Rubiacee. 303 


12, HEDYOTIS,* Z. 


Shrubs or under-shrubs, rarely herbs, stip. conspicuous, 
often connate, forming a sheath; fl. small, in terminal or axil- 
lary, dense or lax cymes; cal.-segm. 4, usually close; cor. 
funnel-shaped or tubular, often hairy, lobes 4, valvate; stam. 4, 
inserted in tube or throat of cor.; ov. 2-celled, ovules numerous 
on large placentas, style simple, stigmas 2; fruit a small cap- 
sule, usually septicidally dehiscent into 2 carp., each of which 
splits down ventral surface, or rarely loculicidal on the crown 
only, or indehiscent; seeds 2-many, angular or plane-convex. 
—Sp. about 80; 57 in Fi. B. Ind. 


Of our 21 species as many as 17 are probably endemic. The flowers 
of many species are dimorphic. 


Capsule splitting septicidally into 2 vertically dehiscent carp. 
Shrubs or erect perennials. 
Capsule not protruded beyond the cal. 
Stip. neither pectinate nor sheathing. 
Cymes paniculate ; 1. over I in. 
Cymes large, terminal. 


L. glabrous. 
Cal.-segm. shorter than tube . 1. H. FRUTICOSA. 
Cal.-segm. longer than tube 3. H. CYMOSA. 
L. softly pubescent : 5 Als Jal, IMUNGRZaI, 
Cymes small, often axillary . . Io, H, CINEREO-VIRIDIS. 
Cymes cory mmbose ; 1. under I in. 2. H. EVENIA. 
Stip. pectinate, not sheathing ; cae axillary. 
Cymes pedunculate : 5. H. OBSCURA. 
Cymes sessile, forming whorls. 
L. glabrous . ; 5 : . 7. H. MEMBRANACEA. 
L. hairy . : 8. H. THWAITESII. 
Stip. pectinate, sheathing ; ; cymes axillary. 
Cymes pedunculate, lax . ‘ 6. H. COPROSMOIDES. 
Cymes sessile, dense, forming whorls g. H. NODULOSA. 
Stip. not or slightly pectinate, very large, 
sheathing, cymes mostly terminal. 
L. not crowded into a rosette. 
L. over I in. 
Lat. veins obscure. . I1. H. RHINOPHYLLA. 
Lat. veins very prominent . . 12. H, LESSERTIANA. 
L. under 1 in., 5-nerved. F 13. H. QUINQUENERVIA. 
L. crowded forming a rosette at end 
of stem . ; . 14. H. GARDNERI. 
Capsule protruded beyond the cal. . . 15. H. LAWSONIA, 
Large stoloniferous herb. 16. H. VERTICILLARIS. 
Capsule opening loculicidally on the € top i annuals. 
L. linear, aristate . ; ; ; . 17. H. CERULEA. 
L. lanceolate, acute . Z Z ; . 18. H. NITIDA. 


* Hedyotis from the use of H. Auricularia in ear-complaints. 


304 Rubiacee. | Hedyotis. 


Capsule indehiscent. 
Capsule crustaceous. 


Prostrate annual; |. under 2 in. : . 19. H. AURICULARIA. 
Erect shrub; 1. over 3 in. P : . 20. H. INAMENA. 
Capsule membranous . 2 , : . 21. Hi. CYANESCHINSE 


1. H. fruticosa, Z. Sf. P/. iol (1753). Wéraniya, S. 


Burm. Thes. 227. Fl. Zeyl. n.63.. Moon Cat. 10. Thw. Enum. 142. 
Gy Laos: 


Fl. B. Ind. iii. 49. Burm. Thes. t. 107 (not good). 

- A much-branched shrub, 4-6 ft. high, bark white, smooth, 
twigs quadrangular with very blunt angles, glabrous, shining, 
whitish; |. 3-4 in., narrowly lanceolate, tapering to base, long- 
acuminate, acute at apex, glabrous, dark green above, pale 
whitish-green beneath, rather thick, lat. veins fine, very oblique, 
pellucid, petiole very short, about { in., stip. slightly connate, 
broadly triangular, acute, thick, stiff, finely serrate with dark 
brown glands in the serratures and in plenty on the inner 
surface; fl. sessile, numerous, cymes both terminal and in axils 
of upper |., much branched, paniculate, branches quadrangular, 
often slightly pubescent, bracts short, truncate, gland-ciliate; 
cal.-limb broadly campanulate, often pubescent, segm. narrow, 
acute, curved outwards, with shaggy white hair within, rather 
shorter than tube; capsule about 4 in., the cal.-segm. covering 
the top, oblong-oval, pubescent, septicidally dehiscent, carp. 
opening ventrally; seeds few. 

Moist region in the low country up to 3000 ft. and perhaps higher ; 


very common. FI. April-August; white. 
Also in Travancore. 


2. H. evenia, 7iw. Enum. 140 (1859). 
CARo3: 
Fl. B. Ind. 111. 50. 


A small shrub about 3 ft. high, branches numerous, eres 
the old ones cylindrical, striate, thickened at the closely 
placed nodes, the young ones sharply quadrangular, glabrous; 
]. very numerous, closely placed, decussate, ?-1 in., lanceolate, 
subacute, margin revolute, quite glabrous on both sides, shining 
above, paler beneath, thick and rigid, lat. veins invisible, 
petiole 4 in., stip. connate, triangular, acute, glabrous, stiff, 
persistent, slightly glandular-denticulate; fl. on short glabrous 
ped., cymes small, corymbose, terminal; cal.-limb glabrous, 
segm. short, narrowly triangular, obtuse; cor. glabrous, tube 
short; capsule ¢in., broadly ovoid or subglobose, septicidally 
dehiscent into 2 ventrally splitting carp. 


Upper montane zone; very rare. Adam’s Peak (Gardner) ; among 
rocks in the river, Maskeliya. Fl. March; white. 


Hedyotis.] Rubiacee. 305 


Endemic. 
Pale golden-green when dry. 


3. H. cymosa, 7iw. Enum. 142 (1859). 

CG] Pe 342T. 

Fl. B. Ind. iii. 50. Bedd. Ic. Pl. Ind. Or. t. 35. 

A small shrub, with numerous spreading, flexuose, nearly 
cylindrical, glabrous branches; 1. 12-3 in., linear-lanceolate, 
sessile, tapering at both ends, subacute, glabrous, lat. veins 
inconspicuous, stip. triangular, long-acuminate, glandular- 
serrate, keeled, persistent ; fl. on short ped., cymes terminal, 
large, very lax, regularly dichotomous, puberulous, bracts 
small, leaflike ; cal. glabrous, segm. longer than tube, linear- 
lanceolate, spreading, recurved; cor. hairy at throat, lobes 
linear-lanceolate, shorter than tube; capsule about ¢ in., nearly 
globose, crowned with the long cal.-segm., dehiscent in 2 
ventrally splitting carp., seeds three in each carp., finely 
muriculate. 

Moist low country; very rare. Pasdun and Reigam Korales 
(Thwaites); summit of Nillowe Kande about 1000 ft. Fl. March, April, 
and September; white. 

Endemic. 


Turns black in drying. I do not find the cal.-segm. longer than ripe 
fruit, as stated in Fl. B. Ind. 


4. H. Macreei, Hz. f. in Fl. B. Ind. iii. 50 (1880). 


‘Branches obscurely 4-angled, softly pubescent ; 1. 3—4 in., 
lanceolate, narrowed at base, acuminate, recurved and com- 
plicate, rather scabridly pubescent above, softly tomentose 
beneath, many nerved, petiole 4-4 in., stip. broader than long, 
abruptly cuspidate, eglandular, tomentose; fl. small, 4 in., 
cymes large, panicled with whorled spreading branches, 
rounded, terminal; cal. pubescent; cor.-lobes pubescent 
within, capsule small, ellipsoid, cal.-teeth shorter than it, cells 
many-seeded.’ 

Based on a specimen collected by Macrae (ex Herb. Hort. Soc. 
Lond.) in Herb. Kew, without definite locality. 

The above description is copied from Fl. B. Ind. 


5. H. obscura, 7iw. Enum. 141 (1859). 

CG FP. '94. 

Fl. B. Ind. iii. 51. 

A low shrub, stems stout, compressed, glabrous; 1. 2-34 in., 
from oval to narrowly lanceolate, tapering to base, acute at 
apex, quite glabrous, rather thick, lat. veins obscure, petiole 
1-4 in., stip. very large, slightly connate, ovate, very deeply 
pectinate with long linear ciliate teeth, soon caducous ; fl. on 

PART II. x 


306 Rubtacee. [Hedyotis. 


very short ped., cymes axillary on rather long glabrous 
peduncles, bracts small, leaflike; cal. glabrous, segm. narrowly 
triangular, subacute; cor. rather large, very hairy at mouth, 
lobes oblong-oval, as long as tube; capsule about {in., tur- 
binate-ovoid, dehiscing into 2 carp. 

Var. 8B, minor, 7hw. in Fl. B. Ind. iii. 52. 


L. smaller, 3-1 in., nearly sessile, oblong-lanceolate, 
rounded at base; fruit much smaller. 

Upper montane zone; rare. Adam’s Peak (Gardner); Maskeliya; 
Horton Plains at foot of Totapella; Wattekelle Hill. Fl. Feb.—April; 
white. 


Endemic. 
Dries black. 


6. H.coprosmoides, 7777. Cat. Ceyl. Pl. 41 (1885). 

Hi. obscura, var. B, Thw. Enum. 141. C. P. 91, Iot. 

An erect shrub, about 3 ft. high, branches divaricate, supra- 
axillary, slightly quadrangular or compressed or nearly 
cylindrical, glabrous; |. varying from 4-2in., ovate-oval or 
ovate, acute at both ends, glabrous above, pilose on veins 
beneath, stiff, lat. veins curved, very conspicuous beneath, 
petiole short, distinct, stip. connate, forming a loose truncate 
sheath with few or many long filiform teeth ; fl. few, pedicel- 
late, cymes supra-axillary, pedunculate, lax; cal. glabrous, 
segm. narrow, acuminate; cor. rather large, lobes as long as 
tube, oblong, hairy over whole surface; capsule nearly glo- 
bose, crowned with erect cal.-segm. 

Upper montane zone in wet shady places; rare. Adam’s Peak 
(Gardner); Horton Plains; Pedurutalagala. Fl. Feb.-April; pinkish- 
white. 

Endemic. 

Besides the characters given, this differs from ZH. obscura in drying 
green instead of black. The two C. P. numbers represent two forms; 


C. P. 91 has larger leaves and flowers and less pectinate stipules than 
GSE 108. 


7. H. membranacea, 7hiw. Enum. 143 (1859). 

(Gs 1B Bion 

Fl. B. Ind. iti. 54. 

Stem simple, stout, bluntly quadrangular, glabrous; 1. 5-6 
in., narrowly oblong-lanceolate, gradually tapering to base, 
long-acuminate, glabrous, pale beneath, lat. veins fine, incon- 
spicuous, petiole 4—? in., stip. not connate, pectinate with 
linear teeth, cymes capitate, sessile, axillary; fl. and fr. not seen. 

Moist low country; rare. Pasdun Korale; Ambagamuwa. First 
pias by Moon, but his specimens are without ‘locality. FI. Sept.; 
white. ; 

Endemic. 


Hedyotis.] Rubiacae. 307 


I have seen only the C. P. specimens, which have the flowers abortive 
and represented by chaffy scales. The whole plant appears to be 
diseased or abnormal. 


8. H. Thwaitesii, 4. f. 22 F7. B. Ind. iii. 54 (1880). 

HZ. macrophylla, Thw. Enum. 142 (non Wall.). C. P. 104, 466. 

FL. B. Ind. iii. 54. 

Stem stout, slightly branched, with long internodes, 
compressed, glabrous, young parts hairy; |. large, 4-6 in, 
lanceolate-oval, tapering at both ends, acute, harshly hairy 
on both sides and on margin, lat. veins conspicuous, petiole 
4? in., pubescent, channelled above, stip. large, slightly con- 
nate, pubescent, pectinate, with long hairy teeth, deciduous; 
fl. sessile, crowded in dense capitate axillary and terminal 
sessile cymes, bracts (often wanting) at base of heads, large, 
ovate; cal. hairy, seem. large, much longer than ov., oblong, 
apiculate; cor. pubescent inside and out; capsule small, ;’p in., 
crowned by long cal.-segm., hairy, splitting into 2 ventrally 
dehiscent carp. 

Moist region; rare. Adam’s Peak (Gardner); Reigam Korale 
(Thwaites). Fl. March, Sept.; white. 


Endemic. 
HH, Thwaitesiz, Hance, is a Chinese plant, now referred to Azotzs. 


9. H. nodulosa, Arn. Pug. 22 (1836). 

Thw. Enum. 143 (excl. var. a). C. P. 88, 3542, 3095. 

Fl. B. Ind. i. 54. 

An undershrub, 3-4 ft., erect, much branched, old stems 
near cylindrical, rough, yellowish-grey, brittle, young ones 
sub-quadrangular, densely covered with spreading hair, buds 
silky-woolly; 1. 2-3} in., ovate-lanceolate or oval, rounded or 
acute at base, rather acuminate, acute at apex, velvety-hairy 
on both sides, veins prominent beneath, petiole very short, 
stip. connate, forming a short tube, truncate at top but with 
several setaceous hairy teeth; fl. sessile, cymes very dense, 
axillary, forming globose whorls; cal.-limb short, broad, segm. 
long, linear-setaceous, covered with long bristly hairs; cor. 
hairy outside, and very hairy in the throat, lobes acute, much 
recurved, nearly as long as tube; capsule oblong-turbinate, 
crowned with long cal.-segm., hairy, splitting into 2 hard 
cocci, each containing a few small seeds. 


Var. 3, Walkeri, Hk. f HH. Walkeri, Arn. Pug. 21. 
Leaves glabrous above. 


Moist region; in low country rare, in montane zone very common up 
to 7000 ft. Var. 8 Ihave not seen. FI. Sept. to April; white or yellowish. 
Endemic. 


308 Rubiacee. [ Hedyotis. 


Easily confounded with Allzophania decipiens unless the fruit be 
examined; but the two stigmas (instead of 4) will also distinguish it, as 
well as the softer character of the hair. Very obviously dimorphic. 


10. EH. cinereo-viridis, 7/w. Exum. 419 (1864). 

CePA 95: 

Fl. B. Ind. iii. 55. 

A shrub, branches slender, spreading, supra-axillary, when 
young acutely quadrangular afterward cylindrical, glabrous; 
1. 4-3 in., long-lanceolate, acute at base, acuminate, acute, 
glabrous, paler beneath, with few lat. veins very fine but 
rather prominent, stip. not connate, narrowly triangular, pro- 
longed into a beak, 2-ribbed, pilose; fl. on short ped., cymes 
axillary and terminal, shortly pedunculate; cal. glabrous, 
limb large, tubular, segm. linear, acute, as long as limb; cor.- 
tube long, slender, lobes as long as tube, oblong-linear, acute, 
glabrous ; ripe fruit not seen. 


Var. 6, subverticillata, 777m. Syst. Cat. Ceyl. 42. C. P. 2639. 


L. larger and broader; cymes all axillary, sessile. Dries 
green. 


Var. y, fumata, 7hw. in Trim.\.c. C. P. 3909. 
Fl. pedicellate, cymes sessile, lax. Dries black. 


Moist region; rare. The type from Matale East. Var. 6 from Am- 
bagamuwa and Nillowe Kande. Var. y from Adam’s Peak. Fl. March— 
June; white. 

Endemic. 

Var. y ought perhaps rather to be considered a distinct species. Var. 
truncata, Trim. |. c., is probably rather a monstrosity (gall) with the 
calyx-limb almost without lobes ; it does not dry black. This species and 
its various forms require further examination. 


af a HZ. rhinophylla, T7hw. ex Trim. in Journ. Bot. xxiii. 208 
I b 
Cp. 3984. 

A small much-branched shrub, branches slender, cylin- 
drical, finely puberulous when young; |. 14-3 in., narrowly 
lanceolate, much tapering to base, very caudate-acuminate, 
acute, glabrous, pale green, whitish beneath; lat. veins obscure, 
petiole short, puberulous, stip. connate, forming a loose sheath, 
puberulous, truncate with a few short deciduous teeth, per- 
sistent ; fl. on long slender ped., cymes terminal, paniculate, 
very lax, bracts small, leaflike; cal.-limb large, campanulate, 
glabrous, segm. short, triangular, subacute; cor.-tube narrowly 
funnel-shaped, lobes rather short, puberulous within; capsule 
$4 in., broadly ovoid, crowned with cal.-limb, readily splitting 
nto 2 ventrally dehiscent carp.; seeds numerous, muriculate. 


Hedyotis.] Rubwacee. 309 


Upper montane zone; very rare, and only in the N.E. mountain block. 
Wattekelle Hill; Summit of Rangala Hill. Fl. Sept.; white. 
Endemic. 


12. H. Lessertiana, 4rz. Pug. 21 (1836). [PLATE XLIX.] 

Thw. Enum. 141. C. P. 96, 99. 

Fl. B. Ind. iii. 52. 

Erect shrub, often almost a small tree, very variable, stems 
stout, cylindrical, glabrous, the younger ones compressed ; 
1. varying much in size, 24-10 in. (or even more), lanceolate 
or oval-lanceolate, tapering at base, gradually acuminate, 
acute at apex, glabrous, rather stiff, strongly plicate, lat. veins 
numerous, very oblique, strong, impressed above, very pro- 
minent beneath, petiole }-1}in., stip. connate, quite free from 
petiole, sheathing, loose, often 14 in. long (in the larger forms) 
and covering whole internode or overlapping, glabrous, trun- 
cate, with a few deciduous setaceous teeth, persistent and 
becoming white and scarious; cymes numerous, terminal and 
from upper l.-axils, in large spreading panicles, ped. as long 
as or longer than cal., cal.-limb campanulate, segm. triangular 
or linear, acute, deciduous in fruit; cor.-lobes narrowly oblong, 
as long as tube, hairy within; capsule nearly jin., broadly 
oblong-ovoid, splitting into 2 ventrally dehiscent carp. 

Var. B, pilosa, Thiw./.c. C. P. 2810. 


Stems hairy; 1. linear-lanceolate, often very narrow, more 
or less hairy beneath; cal.-segm. very short. 


Var. y, confertiflora, 7/w./.c. C. P. 103. 


L. coriaceous; cymes rather dense; fl. larger; cal.-segm. 
triangular. 


Var. 6, marginata, 7hw. ex Fl. B. Ind. l.c. C. P. 3935. 


L. finely ciliate; cymes few, terminal, globose, very dense; 
cal.-segm. long, linear. 


Var. e, lavescens, 7hw./.c.(sp.). C. P. 86, 126. 


Very stout and woody; 1. oval or ovate-lanceolate, very 
coriaceous, petiole dilated; fl. very large, cymes dense; cal.- 
segm. long-lanceolate, acute, persistent; capsule over } in, 
turbinate-ovoid. 


Throughout the montane zone; verycommon. Var. 6, Kotiya Kande, 
Dickoya. Var. e, Maskeliya; Adam’s Peak; Galagama. Fl. March- 
June; white. 

Endemic (?). 

Extraordinarily variable, and might be divided into several species. 
The commonest form at the higher elevations is var. wzajor, Thw. (C. P. 
96). Var. e turns yellow in drying. The flowers are remarkably dimor- 
phic, as noticed by Arnott. I refrain from quoting Bedd. Ic. t. 31, which 
scarcely appears to represent any form of this species. 


310 Rubiacee. [Hedyotis. 


13. H. quinquenervia, 7/w. Exum. 141 (1859). 
CERNIZ0: 
ED S elincesttiws 2a beddwiewbl slid Om to. 


A much-branched bush, older branches cylindrical, smooth, 
whitish, younger ones compressed, glabrous, sheathed with 
persistent stip.; 1. numerous, closely placed, small, 4—# in., 
nearly sessile, broadly ovate, acute, the apex and margin 
strongly recurved, glabrous, thick and leathery, strongly 
5-nerved from the base, veins depressed above, prominent 
beneath, stip. large, connate, forming a cup-shaped lax sheath, 
truncate at mouth, with several linear hairy teeth, scarious, 
white, setose, persistent; fl. on very short ped., cymes sessile, 
capitate, terminal; cal.-segm. linear, subacute, pilose at tip; 
cor.-tube hairy at mouth, lobes strongly recurved, pubescent 
outside at the ends; capsule small, tin., ovoid, separating 
into 2 ventrally dehiscent carp. 


Upper montane zone; very rare. Abundant on Pedurutalagala 
between 7000 and 8000 ft. Fl. April; white, purple outside, anth. purple. 
Endemic. 


14. H. Gardneri, 7iw. Enum. 142 (1859). 

CHE UIG: 

Fl. B. Ind. iii. 53. 

A shrub, stems very stout, woody, 4in. diam., cylindrical, 
slightly branched, completely enveloped in the very large, 
persistent and permanent, closely placed, imbricated, white, 
chartaceous stip., which increase the apparent thickness of the 
stem to about I in.; i. very closely placed, forming a large 
rosette at end of stems, 3-4 in., lanceolate, tapering at base 
into short broad petiole, attenuate-acuminate, acute, glabrous, 
very coriaceous, thick, rigid, somewhat conduplicate, falcately 
recurved, veins inconspicuous, stip. very large, rounded-ovate, 
acute, glabrous, papery, white, overlapping ; cymes paniculate, 
axillary, nearly 1 ft. high, erect, with numerous opp. tricho- 
tomous branches, bracts large, sheathing; cal.-segm. triangular, 
acute, glabrous; fruit not seen. 


Upper montane zone; very rare. By the stone steps on the steep 
part of the ascent to the cone of Adam’s Peak on the Maskeliya side. 
First found by Gardner. FI. Feb. 

Endemic. 

A singular-looking plant resembling a woody Dracena in foliage. I 
regret it was not in flower when I collected it. 


15. H. Lawsoniz,* W. and A. Prodr. 407 (1836). 
Thw. Enum. 140. C. P. 13, 1697. 
BIS Bs lads ainso: 


* The name was bestowed in error. Our plant is not Wendlandia 
Lawsonie, DC. (Lawsonia purpurea, Lam.), which is A. fruticosa, L. 


Hedyotis.] Rubiacee. Ali 


A small shrub, 3-6 ft. high, with very numerous ascending 
branches, stems stout, very bluntly 4-angled, glabrous and 
shining, often purplish, branches supra-axillary, young parts 
glabrous; |. varying from I to over 4 in., but usually 2-3 in., 
oval or lanceolate, acute at both ends, glabrous, rather thick, 
petiole under }in., stip. short, broad, obtuse, rather thick, 
keeled down the centre, deciduous; fl. pedicellate, rather 
large (for genus), cymes very numerous, small, stalked, supra- 
axillary and terminal, forming large compound inflor. usually 
exceeding leaves; cal. glabrous, segm. short, triangular, acute; 
cor.-tube broad, lobes ovate, acute, as long as tube, recurved, 
woolly inside; disk tumid; fruit } in., ovoid-globose, rounded 
and blunt at the top, which projects beyond the erect 
persistent cal.-segm., splitting from above into 2 ventrally 
dehiscent carp. 

Upper montane zone; common, often growing gregariously. FI, 
March, September, and October; white, anth. violet. 

Endemic. 

This beautiful profusely fowering shrub is not variable, but the leaves 
of some specimens from Adam’s Peak are very small with strongly 
revolute margins. It is one of the great ornaments of the higher hill- 
forests, and is well worthy of cultivation in gardens. 

Sir J. Hooker has pointed out that Wight, Ic. t. 1026, cannot repre- 
sentthis species. Neither is it Rheede, Hort. Malab. iv. t. 57, quoted by 
W.and A. The flowers are markedly dimorphic. 


16. H. verticillaris, W. and A. Prodr. 409 (1834). 

FT, plantaginifolia, Arn. Pug. 22. Thw. Enum. 142. C. P. 107. 

FI. B. Ind. iii. 56. Wight, Ic. t. 1020. 

A large stemless perennial herb, with a long vertical 
tap-root and short erect rootstock giving off from the base 
long stout suckers; radical 1. very numerous, very close, erect, 
forming a large rosette, 8-16 in., sessile, much sheathing at 
base, linear-lanceolate, ensiform, acuminate, acute, stiff, quite 
glabrous, bright light green, paler beneath, with 3, 5, or 7 
very strong, thick, longitudinal ribs from base to apex, pro- 
minent and almost winged beneath, channelled above, stip. 
large, adnate to sides of 1.-bases, 3 in. long, narrow, linear, 
acuminate, somewhat laciniate at top, ciliate, with short 
fleshy rootlike papillz especially abundant at the base; scapes 
numerous, axillary, erect, rather shorter than 1, much com- 
pressed, glabrous, fl. sessile, in dense globular stalked heads, 
arranged trichotomously or in whorls in terminal cymose 
panicles, bracts at base of branches large, leaflike, smaller 
upwards; cal. glabrous, segm. narrowly lanceolate, acute, 
cor.-tube hairy within, shorter than cal., lobes acute, recurved ; 
fruit turbinate, crowned with cal, and prominent convex disk, 


Bie heubtacee. [Hedyotis. 


which is perforate in centre; seeds numerous, finely muri- 
culate, dark brown. 

Upper montane zone on the patanas above 6000 ft.; very common. 
Fl. April, May; white. 

Also in the Nilgiri Mts. 

A very curious species, totally unlike all the others in habit. The 
large stipules are completely concealed in the sheathing leaf-bases, and 
are constantly immersed in the water that fills them. 


17. H. ceerulea, WV. and A. Prod. 412 (1834). 

Thw. Enum. 144. C. P. 1674. 

Hl Balindsni oo. bedds tere a ind lOrat-sso: 

Annual, stem erect, 3-12 in., very slender, usually much 
branched, cylindrical, rough with small harsh prominences; 
1, about }in., narrowly linear, sessile, sharply aristate at apex, 
minutely scabrous, margins reflexed, stip. adnate to l.-base, 
pectinate, with long filiform bristle-like teeth ; cymes small, 
few-flowered, sessile, terminal or axillary, fl. sessile, often 
solitary in the bifurcations; cal-segm. lanceolate, rigid, 
tapering to sharp spinous apex; capsule globular-ovoid, 
crowned with erect cal.-limb nearly equalling it in length, 
pubescent, loculicidally dehiscent on the top only. 

Sandy places near the coast, especially in the dry region ; rather 
common. Colombo; Chilaw; Mannar; Jaffna; Batticaloa. FI. Dec.— 


_Feb.; blue. 
Also in S. India. 


18. H. nitida, W. and A. Prod. 412 (1834). Pita-sudu-pala, S 

Hi. Neestana, Arn. Pug. 23. Thw. Enum. 143. C. P. 122. 

Fl. B. Ind. iii. 61. Bedd. Ic. Pl. Ind. Or. t. 36 ? (&. glabella). 

Annual, branches numerous, prostrate or ascending, slender, 
quadrangular, slightly rough on the angles; |. 1-1#in., ovate- 
or oblong- or linear-lanceolate, acute or rounded at base, 
acute at apex, margin rough, often slightly revolute, bright 
light green, glabrous and shining above, scabrous at end, 
whitish and with the midrib often slightly rough beneath, 
stiff and rigid, petiole very short, stip. of long filiform ciliate 
bristles; fl. axillary, solitary or 2 or 3 together, sessile; cal.- 
segm. large, broadly lanceolate, rigid, spinous-ciliate, recurved; 
cor.-lobes longer than tube; capsule globose, crowned with 
large cal.-segm., glabrous, veiny, dehiscent on summit only. 

Lower montane zone; rather rare. Ambagamuwa; Ramboda; near 
Bilahul-oya. Thwaites says ‘common;’ the Fl. B. Ind. queries its 
occurrence here. Fl. March; white. 


Also in Peninsular India. : 
Thwaites says that the leaves of this are chopped up, boiled, and 


eaten with rice. 


Hedyotis.] Rubiacee. ane 


19. H. Auricularia,* Z. SZ. P/. 1o1 (1753). Géta-Kola, S. 

Burm. Thes. 227. Fl. Zeyl.n.64. Moon Cat. 10. Thw. Enum. 142. 
GLP. 1694. 

ieee nd ii. So-9) bur. thes, t 108, f) 1) Bedd. Ic. Pl. Ind. 
OR i127. 


Annual, branches numerous, prostrate, spreading, not 
rooting at nodes, nearly cylindrical, stout, hairy especially 
at nodes and along two lines on opp. sides, dark purple; 
]. 1-2 in., oval, lanceolate or linear-lanceolate, slightly rounded 
at base, acute, very minutely spinous on edge, slightly hairy 
above, more so beneath, lat. veins much curved, impressed 
above, prominent beneath, petiole usually very short, stip. 
short, not connate, very slightly adnate to base of petiole, 
membranous, hairy, with 3 or 5 unequal filiform bristly teeth; 
fl. on very short glabrous ped., crowded in small very dense 
axillary cymes; cal.-segm. small, triangular, acuminate, hairy, 
recurved ; cor.-tube very short, lobes oblong, obtuse, recurved, 
hairy at base; capsule small, nearly globular, hairy, hard, 
indehiscent. 


Low couniry; a common weed. FI. June-September; white. 
Also in most parts of the Eastern Tropics. 


20. H. inamoena, 7hiw. Enum. 143 and 419 (1864). 

C. P. 3543. 

Fl. B. Ind. ii. 61. 

A slender shrub, erect, with few spreading branches, stems 
sub-quadrangular, glabrous; |. 34-44 in., oblong-lanceolate, 
attenuate at base, acuminate, acute at apex, quite glabrous, 
lat. veins inconspicuous, petiole short, stip. connate below, 
triangular, obtuse, 2-ribbed, glandular-pubescent, soon falling; 
fl. few, small, nearly sessile, axillary, within the stipular 
sheath; cal. pilose, segm. ovate, acute ; cor. small, pilose, tube 
rather long; capsule nearly globose, almost glabrous, crowned 
with short cal.-segm., indehiscent. 

Moist region; veryrare. Pitigala Kande, near Balangoda (Thwaites). 
The Fl. B. Ind. gives Adam’s Peak. FI. Sept. 

Endemic. 

Dries black. Thwaites formerly included his H. cénereo-viridis under 


this species, and var. B of that species is very like, but can be easily dis- 
tinguished by the different calyx. 


21. H. cyanescens, 7iw. Lnum. 143 (1859). 
C, P. 2384. 
Fl. B. Ind. iii. 62. 


* Auricularia, from its use as a cure for deafness; a name of the old 
pharmacists. 


314 Rubiacee. | Oldenlandia. 


A much-branched undershrub, the lower branches often 
prostrate and rooting at the nodes, stems acutely quadrangular, 
glabrous, |. 14-24 in., oval, tapering to base, acute at apex, 
rough, with scattered harsh hairs on both sides and margins, 
petiole 4-4 in. stip. short, connate, triangular, very deeply 
pectinate with long subulate teeth, deciduous; cal.-segm. 
linear, ciliate, purple; cor.-lobes ovate, acute; capsule very 
small, narrowly turbinate, membranous, crowned by the cal.- 
segm. which are many times longer, indehiscent. 

Moist low country; very rare. Murutu (Gardner); Ambagamuwa. 
Fl. December; pinkish-white. 

Endemic. 


The whole plant is often of an intense purple colour, especially the 
under sides of the leaves. 


13. OLDENLANDIA, L. 


Herbs, usually annual, stip. various, acute or truncate, 
fl. small, solitary or in small cymes, axillary; cal.-segm. 4, 
erect, distant in fruit; cor.-lobes 4, obtuse, valvate; stam. 4, 
ov. 2-celled, ovules numerous in each cell, on large placentas, 
style simple, stigmas 2; fruit a small capsule, usually some- 
what didymous, loculicidally dehiscent on the crown; seeds 
numerous, smooth or pitted.—Sp. 70; 23 in FZ. B. L[nd. 


Ov. terete. 
Capsule as broad as long. 
L. linear. 
Fl. solitary or in pairs. 
Top of capsule flat, not protruded. 


Fl. pedicellate, in pairs 1. O. CORYMBOSA 

Fl. nearly sessile, solitary : 2. O. DIFFUSA. 
Top of capsule rounded, preideg beyond 

Cala 5 3. O. HERBACEA. 

Fl. 3-10 together, umbellate 4. O. UMBELLATA. 

-L. ovate or rotundate . : 5. O. TRINERVIA. 
Capsule twice as long as broad 6. O. STRICTA. 
Ov. quadrangular 7. O. BIFLORA. 


1. O. corymbosa, ZL. SZ. P/. 119 (1753). Wal-patpadagam, 5S. 

Burm. Thes. 22 (in part). Hedyotis Burmanniana, Br., Thw. Enum. 
144. O. difiora, Moon Cat. 11 (non L.). C. P. 1676, 2638 (part). 

Fl. B. Ind. iii. 64. Burm. Thes. t. 11 (good). 


Annual, small, stems numerous, slender, erect or ascending 
or Sec 6-12 in., glabrous or slightly hispid, cylindrical; 
|. 4-1 in., linear, sessile, apiculate, margin usually scabrous, 
stip. short, truncate, pectinate with few teeth; fi. on filiform 
ped. longer than cal., in pairs (rarely solitary), peduncles very 


Oldenlandia.] Rubwacee. 315 


slender, shorter than 1.; cal.-segm. small, narrowly triangular, 
mucronate; capsule nearly globose, often slightly pyriform, 
truncate, somewhat didymous, glabrous. 

Var. 8, racemosa, 7iw. Enum. 419. C. P. 3837, 2638. 

Fl. 3-6 together, racemose or umbellate, peduncle shorter. 

Low country; a common weed. Var. 8, Colombo. FI. July—Sept.; 
white. 

Throughout the Tropics. Linnzeus founded the species on the 
American plant. Some species seem intermediate in characters between 
this and O. diffusa (e. g. C. P. 2638). 

Very much used as a medicine in remittent fevers. 


2. O. diffusa, Rox. Hort. Beng. 11 (1814). 

Hedyotis Burmanniana, var. brachypoda, Thw. Enum. 419. C. P. 
3838. 

FL. B. Ind. iii. 65. 

Stems numerous, prostrate, often rooting at nodes, slender, 
glabrous; |. $-14in., linear or linear-lanceolate, acute, stip. 
short, slightly pectinate; fl. solitary (rarely 2 together), on 
very short ped.; cal.-segm. rather long, narrow, cuspidate, 
finely ciliate, very distant in fruit; capsule larger than in 
corymbosa, broader than long, very truncate and flat on top. 

A weed in waste ground in low country; less common than O. corym- 
bosa. ¥F\. June-September; white. 

Tropical Asia generally. 


3. O. herbacea, Roxb. Fl. Ind. ed. Wall. i. 445 (1820). 

Fl. Zeyl. n. 65. Hedyotis herbacea, L. Sp. Pl. 102. Moon Cat. Io. 
Hi. Heyniz, Br., Thw. Enum. 144. C. P. 1673. 

Fl. B. Ind. ii. 65 (O. Heyuiz). Rheede, Hort. Mal. x. tt. 23 and 35. 

An erect, rigid, much-branched annual herb with dicho- 
tomous, slender, quadrangular, glabrous, divaricate branches 
with long internodes; |. #-14 in. linear (the lower rather 
broader), sessile, acute, glabrous, margin often recurved, stip. 
very short, inconspicuous; fl. solitary, on stiff, divaricate, 
axillary ped., longer or shorter than 1.; cal.-teeth lanceolate, 
acute, short ; capsule ovate-ovoid, with the top rounded and 
protruded beyond the erect cal.-segm. 

Low country; very common, and up to 4000 ft., in dry sandy places. 
All the year ; white. 

Also in Peninsular India, Malay Islands, and Trop. Africa. 

Hermann’s specimens are the whole foundation for Linnzeus’s 4. 
herbacea, and they are clearly this species, which is generally called 
O. Hleyniz, though Roxburgh had retained Linnzeus’s specific name under 


Oldenlandia, ‘This does not vary in Ceylon, and is an easily distinguished 
species. 


O. crystallina, Roxb., an Indian species, has been occasionally found 
as a weed in gardens, doubtless introduced with other plants in soil. 


3216 Rubwacee. [ Oldenlandia. 


4. O. umbellata, Z. SP. Pl. 119 (1753). Saya, S. Chaya, 7. 

Herm. Mus. 22. Burm. Thes. 208. Fl. Zeyl.n.67. Moon Cat. 11. 
Fledyotis umbellata, Lam., Thw. Enum. 144. C. P. 125. 

Pi beelndenityoon. koxbs Cory bint: 

Annual but semi-woody at base, stems very numerous 
prostrate or ascending, 6-10in., quadrangular or compressed 
glabrous or rough; |. numerous, rather close, sessile, 4-1 in., 
linear, very acute, margin setaceous and often reflexed, stip. 
very short, pectinate; fl. on short ped., 3-10 together in very 
numerous irregular small umbels on stiff erect peduncles ; 
cal. glabrous, segm. triangular, cuspidate, spinous-ciliate ; 
capsule nearly globose, broader than long, top slightly pro- 
truded above cal. 

Sandy ground, especially near the coast and in the dry region; 
common. All the year; white. 

Also in S. India and N. Burma. 

This affords the ‘ Chay-root,’ at one time an important dyeing material 
and a monopoly of Government.* The root was collected especially in 
the north of the island, and is still gathered to a small extent in Mannar 
I., where is a village wholly occupied by a caste who dye cloths with it. 
The colour is a dull pinkish-purple, and very durable. There was formerly 
a considerable export to India. 


5. O. trinervia, Retz. Obs. Bot. iv. 23 (1786). 

fledyotis trinervia, R. and S., Thw. Enum. 144. C. P. 1681. 

Fl. B. Ind. ii. 66. " Bedd. Ie. Pl. Indy Orit2o: 

A small annual, stems weak, flaccid, slender, branched, 
slightly hairy; 1 numerous, broadly ovate or rotundate, 
tapering to base or obscure petiole, nearly glabrous, thin, 
3-veined at base ; fl. on very short ped., in small clusters; cal. 
hairy, segm. narrowly triangular, acute; fruit very small, 
globose, very hairy, flat-topped, opening ‘by a wide cireular 
mouth. 

Low country in damp places; very rare (?). Jaffna(Gardner); Reigam 
Korale (Thwaites). Fl. September. 


Also in India, Burma, Malay Islands, and Trop. Africa. 
Has the habit and appearance of an Alsinaceous plant. 


6. O. stricta, LZ. Mant. 200 (1781). 

Hledyotis maritima, Moon Cat. 10; Thw. Enum. 144 (? Linn. f.). 
C. P. 1678. 

Fl. B. Ind. iii. 68. Pluk. Phytogr. t. 332, f. 3 (bad). 

Annual or perennial with a woody base, and numerous, 
erect, slender, wiry, sub-quadrangular, glabrous, dichotomously 
branched stems, often 2 ft. high; 1. 1-14 in., sessile, linear, 


* Extracts from the Dutch Records referring to the collection of this _ 
plant for Government are given by the late Colonial Surgeon Ondaatje 
in the appendix to the Ceylon Almanac for 1853, pp. 14-16. 


Anotis.| Rubiacee. Bi, 


very acute, glabrous, stip. adnate to base of |. and with them 
forming a close sheath round the stem, mouth truncate, ciliate; 
fl. on slender, erect ped., few, distant, in slender erect elon- 
gated racemose cymes; cal.-segm. short, lanceolate, subulate; 
cor.-lobes oblong, obtuse, longer than tube; capsule 3 in., 
twice as long as broad, oblong, crowned with tooth-like cal.- 
segm., glabrous, top slightly rounded. 

Sandy seashores ; rather common, especially in the north of the island. 
Fl. Dec.—March; pink. 


Also in S. India. 
Remarkably like a Sz/ene or Tunzca in habit. 


7. O. biflora, Z. SA. Pi. 119 (1753). 

E Fl. Zeyl. n. 68. AHedyotis racemosa, Lam., Thw. Enum. 144. C. P. 
1679. 
PL B. Ind. iii. 69 (O. paniculata) and 70. Burm. f. Fl. Ind. t. 15, f. 1. 
(bad) (O. paniculata). Wight, Ic. t. 312. 

Annual, stems 4-12 in., erect or ascending, rather stout, 
quadrangular, glabrous, slightly branched; |. 4-14 in., oblong- 
oval, tapering at base into obscure petiole, subacute, apicu- 
late, glabrous, rather thick, stip. short, broadly triangular, 
acute; fl. rather large (for genus), pedicellate, 2 or 3 together 
on short axillary peduncles, or, the 1. being small and bract- 
like, together forming a short racemose panicle; cal.-segm. 
lanceolate, acute, glabrous; ov. distinctly quadrangular, 
glabrous; capsule turbinate-ovoid, strongly 4-ribbed, truncate, 
crowned with distant cal.-segm.; seeds pitted. 

Low country, chiefly near the coast; rather common. Galle; 
Colombo; Batticaloa; Trincomalie; Mannar. All the year; white. 

Throughout the Tropics of Asia. 

I cannot distinguish two species here, as is done in Fl. B. Ind. 
Hermann’s type specimen of O. dz/lora, L., is a luxuriant one, and would 
be now generally named O. faniculata. The brief description of it in F1. 
Zeyl. is good. O. paniculata, L., is, moreover, quite doubtful; it is 
entirely based on a figure of Burman in Thes. Zeyl. t. 71, f. 2, which is 
apparently a J/o//ugo (certainly not an Oldenlandia). But O. paniculata, 
Burm.f. (the figure quoted above from his Fl. Ind.), is the present species. 


14. ANOTIS, D.C. 


Annual or perennial herbs, stip. pectinate or bristly or 
nearly absent, fl. small, nearly sessile, in small axillary or 
terminal cymes; cal.-segm. 4, distant in fruit; cor.-lobes 4, 
shorter than tube, valvate; stam. 4; ov. 2- or 4-celled, ovules 
few or I in each cell, placentas basal, style simple, stigmas 2 
or 4, linear; fruit a small capsule, dehiscent on the top or 
indehiscent; seeds very few, pitted.—Sp. 25; 18 in FZ. B. Lund. 


318 Rubtacee. [Anotis. 


Capsule 4-celled . : : : : . I. A. QUADRILOCULARIS. 
Capsule 2-celled. 
Capsule flat, orbicular, indehiscent. 


Lat. veins of 1. obvious . : : . 2. A. NUMMULARIA. 
Lat. veins of |. invisible . : : . 3. A. NUMMULARIFORMIS. 
Capsule ovoid-turbinate, septicidal . . 4. A. RICHARDIANA. 


1. A. quadrilocularis, 2. f in Fl. B. Ind. iii. 74 (1880). 

Hedyotis guadrilocularis, Thw. Enum. 144. C. P. 3132. 

Fl. B. Ind. iii. 74. 

Annual, 6-18 in., with many flaccid, spreading, glabrous, 
cylindrical branches; |. ?-14in., ovate-oval, acute at both 
ends, slightly hairy, very thin, pale beneath, petiole 4 in., stip. 
short, truncate with a few bristles; fl. nearly sessile, solitary 
or 2-4 together in the axils (those on short branchlets with a 
single pair of l. appearing terminal); cal.-segm. broadly 
oblong; cor.-lobes ovate, bristly on back; ov. 4- (rarely 3-) 
celled, stigmas 4, linear, drooping; capsule depressed, broader 
than long, glabrous, 4-lobed, capped with large cal.-segm., 
4-celled; seeds 3 in each cell; nearly globose, deeply pitted. 

Low country; rare, on wet rocks. Bintenne District; Kindegodde, 


Badulla Dist. (Thw.); near Lunugala, Uva. Fl. Jan—April; white. 
Also in S. India. 


2. A. nummularia, “2. f. F/. B. Ind. iii. 75 (1880). 
; Fledyotis nummutlaria, Arn. Pug. 23; Thw. Enum. 142. C. P. 1692, 
2817. 
FL B. Ind. iii. 75. . 

A perennial stoloniferous herb, stems prostrate at base 
and rooting at the nodes, then erect, about 1 ft. high, stiff, 
quadrangular, slightly winged, pubescent or nearly glabrous, 
rather succulent, very slightly branched at top; |. sessile, 4-3 
in., rotundate, subacute, usually hairy on both sides, some- 
times glabrous, rather thick, lat. veins obvious, stip. 0 or a few 
bristles; fl. numerous, sessile, cymes terminal on rather long 
peduncles, with 2 leaf-like bracts at base, dense, capitate; cal.- 
segm. shorter than ov., lanceolate, acute, glabrous, cor.-tube 
somewhat inflated, lobes lanceolate, acute, recurved, with a 
few bristly hairs outside; stam. erect, exserted; style short, 
stigmas 2, plumose, disk annular; capsule about fin. flat, 
orbicular 2-celled, indehiscent, smooth, with 2 ridges on each 
face, seeds I or 2 in each cell. 

Upper montane zone in wet places; common. FI. Feb. and August; 
pale violet—blue. 

Endemic. 


Plant slightly fetid when bruised. Nearly black when dried. C P. 
2817 is a glabrous form, but different from the next. 


Anotis.] Rubriacee. 319 


3. A. nummulariformis, 77im. Syst. Cat. Ceyl. 42 (1885). 

Hledyotis nummulariformis, Arn. Pug. 23. Hf. nummularta, var. 
glabra, Thw. Enum. 142 (in part). C. P. 113. 

Fl. B. Ind. iii. 75 (A. smmularia, var.). 

Perennial (?), stems 6-18 in., branched, weak, ascending, 
bare below, strongly quadrangular, slightly winged, usually 
glabrous (sometimes slightly hairy), shining and polished; 
1. very shortly stalked, $-?in., narrowly ovate, usually 
rounded at base, acute, margins recurved, glabrous above, 
very pale and sometimes hairy beneath, rather thick, midrib 
prominent beneath but lat. veins invisible, stip. pectinate with 
long linear teeth fimbriate at apex; fl.nearly sessile, crowded in 
small capitate cymes on long peduncles both terminal and 
axillary; cal.-segm. triangular, acuminate; cor.-lobes short, 
acute, recurved; capsule as in the last. 

Upper montane zone in wet places; rather common. N. Eliya; 
Maturata; Adam’s Peak; Knuckles Mts. Fl. Feb., and Aug., Sept.; 
pale violet, 

Endemic. 

I do not think this should be combined with O. xzaummularia; as is 
done by Thwaites. 


4. A. Richardiana, Zé. f. in Fl. B. Ind. iii. 75 (1880). 

Hedyotis Richardiana, Arn. Pug. 22. A. monosperma, W. and A., 
var. subglabra. Thw. Enum. 142. C. P. 97. 

Fl. B. Ind. iii. 75. 

A perennial herb, stems prostrate, rooting at the nodes, 
spreading, much branched, flowering shoots ascending, cylin- 
drical, hairy; |. usually small, $-?in., broadly ovate, rounded 
or slightly cordate at base, acute, hairy on both sides, rather 
thick, with the numerous lat. veins prominent beneath, petiole 
very short, stip. large, hairy, cut half way down into linear 
teeth; fl. sessile, cymes small, lax, terminal, on short unequal 
peduncles; cal. oblong-lanceolate, acuminate, hairy, cor.-tube 
narrow, hairy within, lobes nearly as long as tube, spreading, 
stam. not exserted, style long, stigmas 2, plumose; capsule 
4 in., ovoid-turbinate, slightly compressed with a convex top 
exceeding the persistent cal.-segm., glabrous or hairy, 2-celled, 
with a single seed in each cell, at length splitting septicidally. 

Upper montane zone; very common, and occurring in large masses. 
Fl. August, Sept.; bright violet. 

Endemic. 

A very pretty little plant, forming large beds in the hills, but with a 
disgusting odour when bruised as by walking over it. 

The above description is that of the ordinary form, but the 1. are often 
much larger, 3-4 in., on longer petioles and nearly glabrous, the stems 
erect and much taller, and the cymes very lax. I suspect some of ours is 
rather A. monosperma, W. and A. (Wight, Ic. t. 1031), to which Thwaites 
referred it. 


320 Rubiacee. [ Ophiorrhiza. 


15. OPHIORRHIZA,* L. 


Herbs or undershrubs, stip. not conspicuous, fl. small, 
erect, secund on branches of dichotomous cymes, on ter- 
minal peduncles, with or without bracts; cal.-limb 0, segm. 5 ; 
cor. tubular, lobes 5, valvate; stam. 5, inserted in tube, anth. 
linear; disk of 2 very large, prominent erect lobes which 
become much enlarged in fr.; ov. 2-celled, placentas arising 
from base of dissepiment, with many ovules; fruit a capsule, 
very strongly compressed, crowned with enlarged disk, which 
protrudes beyond cal.-segm., coriaceous, much broader than 
long, loculicidally dehiscent on the top, seeds numerous, 
angular.—Sp. about 50; 31 in FZ. B. Ind—The species are 
obscurely defined. 


Bracts absent or early caducous. 
Stem erect. 


Herbaceous; cal. segm. short, triangular 1. O. MUNGOS. 
Shrubby; cal.-segm. narrow, very acute 2. O. HARRISIANA. 
Stem prostrate, rooting below. 3. O. RADICANS. 


Bracts persistent with fruit. 
Bracts glabrous. 


L. glabrous . 4. O. PECTINATA. 
L. hairy 5. O. PALLIDA. 
Bracts ciliate . 6. O. GLECHOMIFOLIA. 


1.0. Mungos,t Z. Sp. Pl. 150 (1753). Dat-kétiya, Wal- 
ékaweriya, 5. 

Herm. Mus. 37. Fl. Zeyl. n. 402. Moon Cat. 19. Thw. Enum. 139. 
C. P. 1704. 

Fl. B. Ind. ii. 77. Gaertn. Fruct. i. t. 55 (fruit only). 

Herbaceous, never shrubby, stem 14-2 ft., erect, cylin- 
drical, slightly branched, glabrous or very finely puberulous; 
l. rather large, 4-8 in., lanceolate, much tapering at base and 
decurrent on short petiole, shortly acuminate, subacute, 
glabrous above, finely puberulous on veins beneath, thin, 
bright green above, pale beneath, lat. veins very numerous, 
depressed above, prominent beneath, stip. very short; cymes 
several, horizontally spreading, arranged sub-umbellately in a 
flat-topped terminal infl. quite without bracts, fl. on short 


* From the alleged specific action of the roots against the poison of 
serpents, Linnzeus following Kaempfer in this. 

+ Kaempfer (1712) is the first to connect this plant with the Mungoose 
by calling it Radix Mungo and quoting Garcia de Orta’s account of 
‘Lignum colubrinum’ for it. I think, however, from the description 
given by the latter, that the plant meant was rather Rauwolfia serpentina 
(see under that species). 


Ophiorrhiza.] Rubiacee. 321 


ped.; cal.-segm. very short, triangular, cor.-tube inflated at 
base, broad, with a ring of long silky hair at mouth, lobes 
much shorter than tube; disk very prominent; capsule 2 in. 
wide, coriaceous, 3 times as wide as long, very much com- 
pressed, protruded beyond cal.; seeds very numerous, 
angular, pale brown. 

Var. 8, nemorosa, /7k. f. O.emorosa, Thw. Enum. 139. C. P. 205. 


L. narrower, stip. with 1 or 2 long setaceous bristles; fl. 
much larger; cor.-tube longer. 


Var. y, angustifolia, 7k. f O. angustifolia, Thw. Enum. f4o. 
C. P. 431. 


L. smaller, oblong-lanceolate, stip. with setaceous bristles, 
cymes small, with small linear bracts, tomentose. 


Shady places in the moist region from sea-level up to 6000 ft.; 
common. Var. 8, Medamahanuwara; Galagama. Var. y, Ambaga- 
muwa. All the year; white, in var. 8, pinkish. 

Also in India, Burma, Malay Islands; the varieties endemic. 

The root is said to have a reputation for snake-bites in India; but not, 
I think, here. The leaves are bitter, and a remedy for toothache. There 
is no published figure of this familiar plant. 

Linnzeus refers to this also the ‘ Naghawalli’ of Herm. Mus. 55, which 
the latter says is brought from Adam’s Peak, the leaves being a specific 
cure for the bite of the cobra. They are described as spotted, and it is 
likely some other plant was intended. Moon, indeed, gives this name for 
Justicia picta (Graptophyllum hortense), a common garden plant. 


2. O. Harrisiana, Heyne in Wall. Cat. n. 6236 (1828). 

O. decumbens, Gardn. in Thw. Enum. 419. C. P. 3656. 

Fl. B. Ind. iii. 78. Wight, Ic. t. 1162 (O Harrisoniz). 

A small undershrub, with many ascending slender branches, 
often rooting at the base, young twigs glabrous or pubescent; 
l. 14-34 in., ovate-oval or lanceolate, tapering to both ends, 
acute, pale and pubescent on veins beneath, stip., with a few 
setaceous bristles; cymes few, small, lax, pubescent, on long 
slender peduncles, bracts setaceous, soon falling; cal.-segm. 
very acute, capsule small, triangular, with the ends prolonged, 
pubescent. 

Moist shady places in lower montane zone; common. FI. June- 
September; white. 

Also in India and Burma. 

Scarcely more than a variety of O. Mungos, but its shrubby habit gives 

it a different look. 


3. O. radicans, Gardn. in Thw. Enum. 139 (1859). 
©, P..1706. 
Fl. B. Ind. iii. 80. 
Stems prostrate, filiform, copiously rooting, with ascending 
branches and ends, finely woolly-pubescent; |. small, }—1 in., 
PART IL. Y 


322 Rubiacee. [Ophiorrhiza. 


numerous, ovate or rotund-ovate, obtuse or subacute, glabrous 
except on veins beneath, petiole about half as long, pubescent, 
stip. inconspicuous; fl. few, shortly stalked, in small bifid 
- cymes on erect slender apparently terminal peduncles much 
exceeding the l., bracts subulate, caducous; cal.-segm. acute; 
capsule very small, Lin. wide, 3 times as broad as long, hairy. 

Moist region to 4000 ft.; rather common (?) Hantane; Kukul Korale, 
&c. Fl. June; white. 

Endemic. 


I have a very Swnileaved creeping form from Kalubowitiya Kande, 
Hiniduma. 


4. O. pectinata, Arz. Pug. 20 (1836). 

Thw. Enum. 140. C€. P. 712; 750. 

IML, 13}, Tea iis ie 

Stem erect, often semi-shrubby below, slightly branched, 
glabrous or pubescent; |. 3-6 in, oval to linear-lanceolate, 
much tapering below into obscure petiole, acuminate, acute, 
glabrous on both sides, thin, midrib very broad, lat. veins 
numerous, curved, stip. over 4in., broad at base, then filiform; 
fl. very shortly stalked, closely placed in dense cymes um- 
bellately arranged at end of stiff peduncle, and forming an 
almost globose head, bracts conspicuous, much exceeding 
cal., linear-lanceolate, acuminate, glabrous, persistent in fruit; 
cal.-segm. small, subulate; cor.-tube 4 in., lobes short, hairy 
within; capsules crowded and overlapping, 2 in. wide. 

Lower montane zone in forests; very common. Fl. March—May; 
white or greenish. 

Also in S. India. 


C. P. 750 has the flowers very closely packed in the corymbs, and is 
var. 6 of Thw. (O. zmbricata, Gardn.). 


5. O. pallida, 7iw. Enum. 140 (1859). 

C. P. 711, 3344. 

Fl. B. Ind. i. 81. 

Stems more or less procumbent and often rooting below, 
roughly tomentose; |. 14-34 in., lanceolate or ovate-lanceolate, 
acute or rounded at base, acuminate, acute, hairy on both 
sides, petiole rather long, hairy, stip. simple setaceous; cymes 
small, subglobose, glabrous, on long peduncles, bracts rather 
long, linear; cor.-lobes very short; capsule about }in. wide, 
glabrous. 


Moist region; very rare. Murutu (Gardner); Ambagamuwa. FI. 
December; white. 


Endemic. 


C. P. 3344 (from Ambagamuwa) is a very small form more creeping, 
and with broader, smaller leaves. 


‘Mussenda.| Rubracee. 


Oo 
Oo 


6. O. glechomifolia, 7iw. L7w7. 140 (1859). 

C. P. 1708. 

Fl. B. Ind. iti. 84. 

Stems completely prostrate, rooting a the nodes, covered 
with crisp hairs, rather stout; 1. small, 3 L—3 in., broadly ovate, 
rounded or slightly cordate at ‘base, acute or subacute at apex, 
coarsely hairy on both sides, petiole rather long, hairy, stip. 
inconspicuous; cymes small, bifurcate, few-flowered, peduncles 
glabrous or pubescent, bracts linear, ciliate, persistent; cal.- 
segm. large, lanceolate-oblong, acute, strongly ciliate; SO 
lobes as long as tube, hairy within; capsule very small, yz in., 
crowned with large cal, -segm. , hairy. 

Moist low country to 3000 ft.; rare. Allagala; Ramboda; Sabara- 
gamuwa Dist. Fl. Feb., July, Sept.; white. 

Endemic. 

Very like O. radicans in habit, but distinguished by its much more 
hairy leaves and larger ciliate cal.-segm. 


16. MUSSZENDA,* Linz. 


Scandent, shrubby, stip. small, sometimes bifid, fl. in ter- 
minal cymes; cal.-segm. 5, narrow, deciduous, one sometimes 
developed into a large petioled white leaf; cor.-tube with a 
dense lining of hair at mouth, lobes short, valvate, with 
reflexed margins; stam. 5, inserted about 4 down and enclosed 
in cor.-tube, fil. 0, anth. linear; ov. 2-celled, ovules numerous, 
on large fleshy placentas; fruit soft, indehiscent, 2-celled ; 
seeds many, minute, pitted.—Sp. 4o; 11 in FV. B. [nd. 


MM. frondosa, ZL. Sf. Pl. 177 (1753). Wussenda, Wel- 
butsarana, 5S. 

Hern. Mus. 36. Burm. Thes. 165. Fl. Zeyl.n. 84. Moon Cat. 15. 
Thw. Enum. 138. C. P. 1664. 

Fl. B. Ind. iii. 89. Burm. Thes. t. 76. Wight, Ill. t. 124. 

A scrambling shrub, climbing by its long, flexuose, divari- 
cate branches, which are cylindrical and finely velvety; 1. 3-5 
in., ovate-lanceolate, rounded at base, shortly acuminate, sub- 
acute, finely velvety on both sides, paler and with prominent 
venation beneath, petiole 1-4in., pubescent, stip. rather small, 
triangular, usually more or less bifid, membranous, pubescent, 
soon falling; fl. on stout pubescent ped. as long as ov., in 
rather small, contracted, paniculate, terminal cymes, bracts 
small, subulate, caducous, buds blunt, strongly 5-angled; cal.- 
segm, nearly 4 4 in., narrowly linear, pubescent, deciduous, the 


* The Sinhalese name unaltered. 


324 Rubiacee. [Acranthera. 


enlarged one (only developed on the outer side of a few fi.) 
in all respects save colour resembling the foliage-l, but rather 
broader, velvety on both sides, creamy white, persistent; cor.- 
tube Zin., with a dense lining of short yellow hairs completely 
blocking the mouth and extending 4 way down, lobes short, 
broadly oval, apiculate, convex above, spreading; anth. small; 
fruit 2 in., nearly globose, faintly 2-lobed, slightly scabrous. 


Var. B, glabrata, kz. f. in Fl. B. Ind. 


L. smaller, 2-3 in., narrower, tapering to base, quite 
glabrous on both sides; buds acute, not 5-angled; cal.-segm. 
very short, acuminate, glabrous; cor.-lobes acuminate; fruit 
glabrous. 

Moist region up to 4000 ft.; verycommon. Var. in the low country. 
Fl. April-August; brilliant orange. 

Also in India, Andaman Is., and Malay Archipelago. 

A very conspicuous plant everywhere, from the white leaf-like enlarged 
calyx-lobes and brilliant, though not large, flowers. The hairs on the 
leaves are sometimes pink. Extremely variable in amount of hairiness, 
often villous in the hills. In Fl. B. Ind. all the Ceylon plant is called 
var. zeylanica, which is, however, restricted by the description to the 
velvety or tomentose form, which is the type as described by Linnzeus. 
Var. 6, which looks quite distinct, is perhaps the JZ. corymbosa of Roxb., 
which he states (Fl. Ind. i. 556) to be a native of Ceylon, or possibly 1. 
glabra, Vahl, which has a wide Malaya and Burma distribution but is 
not recorded for Peninsular India in FI]. B. Ind. 

A decoction is used externally in inflammations. 


17, ACRANTHERA, Arn. 


Perennial herb, 1. large, stip. triangular; fl. large in ter- 
minal cymes; cal.-segm. 5, long, persistent ; cor. tubular-funnel- 
shaped, lobes 5, valvate; stam. 5, inserted near base of cor.- 
tube, anth. connate at their points; ov. 1-ce//ed, but apparently 
2-celled, the two parietal placentas completely meeting in the 
centre and reflexed, stigma clavate; fruit oblong, indehiscent, 
2-celled; seeds minute, very numerous.—Sp. 10; 6 in FZ. B. 
Ind. 


A. zeylanica, Arn. ix Ann. Nat. Hist. iii. 21 (1839). 

Thw. Enum. 138. C. P. 334. 

Fl. B. Ind. iii. 92. Bedd. Ic. Pl. Ind. Or. t. 24 (not good). 

A low perennial herb, rootstock thick, flexuose, stem. 2-8 
in., often prostrate and rooting at base, then erect, stout, com- 
pressed, densely silky-hairy; 1. large, 4-6 in., closely placed, 
obovate-oblong or oblong-lanceolate, tapering to base, rounded 
or subacute at apex, sparingly hairy above, and densely so on 


Leucocodon.| Rubiaceae. 225 


veins beneath, strongly ciliate, thin, very bullate above, paler 
with prominent coarse reticulate venation beneath, petiole 
short, densely silky-hairy, stip. large, triangular, acuminate, 
hairy; fl. large, nearly sessile, 3-12 together in rather dense, 
trichotomous cymes, on stout, very hairy, erect (deflexed in 
fruit) terminal peduncle, bracts numerous, very hairy; cal. 
hairy, segm. linear, long, acuminate; cor.-tube 3 in., much 
exceeding cal.-segm., narrow below, dilated in upper part, 
lobes nearly 4in., oblong, obtuse, spreading; stam. included, 
fil. apiculate; style included ; fruit 3 in., crowned with cal.- 
segm., oblong-fusiform, membranous, very hairy; seeds nearly 
globose, reticulate. 

Lower montane zone, in damp, shady places; rather rare. Adam’s 
Peak; Sabaragamuwa; Ambagamuwa; Hunasgiriya; Rangala Hill. 
Fl. April-June; bright pale violet-blue, faintly sweet-scented. 


Endemic. 
First collected in 1836 by Wight and Walker. 


18. LEUCOCODON, Gardn. 


An epiphytic climbing shrub, stip. large, connate at base; 
fl. rather small, in a terminal head, surrounded by a large 
campanulate involucre; cal. adnate to lower 3 of ov., limb 
shortly tubular, irregularly truncate, segm. 0; cor.-tube funnel- 
shaped, lobes 5, valvate; stam. 5, inserted at mouth of cor.- 
tube; ov. only partially inferior, crowned with an annular disk, 
2-celled, with numerous ovules, stigma bilobed; fruit fleshy, 
indehiscent, 2-celled; seeds numerous, compressed.—Mono- 
typic and endemic. 


L. reticulatum, Gardn. in Calc. Journ. Nat. Fist. vii. 5 (1846). 

Thw. Enum. 138. C. P. 347. 

Fl. B. Ind. iii. 94. Bedd. Ic. Pl. Ind. Or. t. 94. 

A woody epiphyte, sending off (like Ivy) adventitious 
rootlets by which it climbs, stems cylindrical, marked with 
leaf-scars and stipular rings, young parts glabrous; |. 4-6in., 
lanceolate or linear-lanceolate, acute or attenuate at base, 
shortly acuminate, acute at apex, glabrous on both sides, 
petiole about 4in., stip. 1 in. long, broadly oblong, connate at 
base, bidentate at apex, glabrous, membranous, soon falling; 
fil. rather small, sessile, capitate, on a short terminal peduncle, 
surrounded by a very large involucre (formed of connate 
bracts) which is campanulate, 14-17 in. wide, with 8 broad, 
obtuse, recurved lobes, strongly 8-ribbed, glabrous, greenish- 
white; cal.-limb short, truncate; cor.-tube narrow, lobes 


226 Rubiacee. [ Urophyllum. 


lanceolate, acute, shorter ian tube; fil. very short, anth. 
exserted; fruit (not seen) 4 in., obovate- oblong, compressed, 
fleshy, white: seeds striate, “yormaish: black. 

Wet forests up to 4000 ft.; rare. Ambagamuwa (Gardn.); Ramboda; 
Adam’s Peak; Maskeliya; Singhe Raja Forest; Kottawa Forest, S. Prov. 
Fl. January-May; greenish-white. 

Endemic. 

A remarkable plant; the involucre resembles a large corolla. 


19. UROPHYLIUM, Wail. 


Shrubs, stip. undivided ; fl.small, often unisexual, in axillary 
cymes or umbels; cal.-limb large, campanulate, thick, segm. 5, 
shallow; cor.-tube short, broad, hairy at mouth, lobes 5, longer, 
thick, valvate; stam. 5, inserted in cor.-tube, fil. short; ov. | 
capped with thick prominent disk, 5-celled, with numerous 
ovules in each; style short, stigmas 5; fruit a fleshy globose 
berry; seeds minute, numerous, tubercled.—Sp. 35; 9 in FZ. 
B. Ind. 

Both our species are endemic. 


Stip. large, oblong; cymes nearly sessile : . I. U. ELLIPHICUM 
Stip. small, triangular; cymes pedunculate . . 2. U. ZEYLANICUM. 


1. U. ellipticum, 7iw. Enum. 419 (1864). 

Axzanthes elliptica, Wight in Calc. Journ. Nat. Hist. vii. 147. Thw. 
Enum 1305 /€. 2.350: 

Fl. B. Ind. i. 97. Wight, Ic. t. 1164 (A xanthes). 


A large shrub, with stiff branches, young ones cylindrical 
and glabrous below, sub-quadrangular and pubescent above; 
]. large, 6-10 in., lanceolate, acute at base, acuminate, acute, 
glabrous above, often pubescent on wea beneath, rather 
thick, veins prominent beneath, petiole 1-2 in. , slightly hairy, 
stip. ¢in., linear-oblong, acute, thick, sig tL * deciduous: cymes 
small, nearly sessile; fl. crowded, on very short elabrous ped. ; 
cal. glabrous, segm. very broad and shallow, apiculate ; cor. 
with long coarse hair within, lobes ovate, acute; berry small, 
dull orange-yellow. 

Moist region from 1000-6000 ft.; rather rare. Adam’s Peak; N. Eliya; 


Deltota; Labugama. fl]. Jan._May; white. 
Endemic. First collected by Wight in 1836. 


2. U. zeylanicum, 7iw. Enum. 418 (1864). 

Axanthes zeylanica, Wight in Calc. Journ. Nat. Hist. vil. 146. Thw. 
Enum. 139. C. P. 449. 

IME 184) Iavel, soil, Git, 1 Wane, WWE. tes WlOsy 


Webera.] Rubiacee, 327 


A shrub, with long spreading, cylindrical glabrous branches; 
1. 3-5 in., lanceolate, acute at base, acuminate, subacute, quite 
glabrous on both sides, venation reticulate, pellucid, prominent 
beneath, petiole $—#in., stip. small, narrowly triangular, thick, 
persistent; cymes irregularly umbellate, peduncles about as 
long as petioles, ped. rather long, puberulous; cal. and cor. 
asin U. ellipticum,; berry 4—2 in., depressed-globose, crowned 
by rim of truncate, prominent, fleshy cal.-limb, smooth, pale 
waxy, orange-yellow; seeds ovoid, compressed, pale brown. 

Montane zone, 3000-7000 ft.; rather common. FI. Feb.—April, and 
Aug.—Sept.; pale yellow. 

Endemic. 

The Sinhalese carpenters at N. Eliya call this ‘Wal-handun.’ 


20. SCHIZOSTIGMA, Arn. 


A creeping, very hairy herb, stip. ovate, persistent; fl. rather 
large, axillary; cal.-limb short, segm. large, unequal; cor.-tube 
funnel-shaped, lobes 5, narrow, valvate; stam. 5, inserted in 
upper part of cor.-tube, anth. slightly exserted; ov. 5-celled, 
with numerous ovules, stigmas 5, spreading; fruit a hairy berry, 
5-celled, many-seeded.—Monotypic and endemic. 


S. hirsutum, 4rz. in Ann. Nat. Hist. iii. 20 (1839). 

Thw. Enum. 139. C. P. 268. ‘ 

Mie ind. 11. 100,, Bedd:, le: Pl, Ind: Or. t. 95: 

Stems prostrate, long, rooting at nodes, cylindrical, very 
hairy, with short erect branches; |. 2-3 in., lanceolate-oval, 
acute at both ends, hairy on both sides, especially on veins 
beneath, thin, petiole 4-1 in., very hairy, stip. rather large, 
ovate, acute, nearly glabrous, persistent; fl. solitary or 2 or 3 
together, on hairy axillary ped.; cal.-segm. 3 in., both broadly 
and narrowly lanceolate, acute, slightly hairy; cor.-lobes as 
long as tube, oblong, acuminate, hairy; berry crowned with 
large cal.-segm., about $in., nearly globular, densely covered 
with long hair. 

Moist low country up to 2000 ft. or higher; rather common. FI. Feb., 
March; white. 

Endemic. Apparently first collected by Finlayson (Wall. Cat. n. 8463), 
who visited Ceylon in 1817-8. 


21. WEBERA, Schred. 


Shrub or small tree, stip. simple, subpersistent, fl. moderate- 
sized, in terminal paniculate cymes; cal.-limb short, segm. 5 
cor.-tube short, hairy in throat, lobes 5, twisted in bud 


Be 8 Rubiacee. [Byrsophyllum. 


stam. 5, inserted at mouth of cor., fil. short, anth. narrow; 
ov. 2-celled, with few ovules in each cell half immersed in 
fleshy placentas; fruit a hard 2-celled berry, seeds few.— 
Sp. about 40; 20 in FZ, B. Ind. 


W. corymbosa, W7lid. Sp. Pl. i. 1224 (1797). Tarana, S 
Karanai, 7. 

Fl. Zeyl. n. 80. Rondeletia astatica, L. Sp. Pl. 182. Tarenna zeylanica, 
Gaertn. Fruct. t. 139. Moon Cat. 19. Stylocoryne Webera, A. Rich., 
Thw. Enum. 158. C. P. 185. 

Fl. B. Ind. iii. 102. Wight, Ic. tt. 309, 584. 


A small tree or large bush, bark smooth, pale grey, twigs. 
compressed, glabrous, buds exuding a waxy secretion; |. 24-6 
in., oblong-lanceolate, acute at both ends, glabrous, polished 
above, rather thick, lat. veins prominent beneath, usually with 
small pits in their axils, petiole about 4in., stip. narrowly 
triangular, acute, closely pressed to stem, and at first glued 
to it by waxy secretion, subpersistent ; fl. on short nearly 
glabrous ped., about 4 in. diam., cymes trichotomous; cal.- 
segm. oblong, obtuse, faintly pilose; cor. hairy at mouth, 
lobes longer than tube, oblong, acute, reflexed; stam. ex- 
serted; stigma and portion of style exserted; berry 4-4 in., 
more or less globose, glabrous, dull green, becoming black, 
usually without cal.-segm. on top. 

Var. 6B, montana, 7hw. /. c. Webera cerifera, Moon Cat. 19. 
CaP. s1620: 

Inflor., buds and stipular nodes exuding copious resinous 
wax; fl. larger; cal.-segm. acute, persistent, and crowning the 
fruit. 

From sea-level up to 7oooft.; very common. Var. 6 in upper montane 
zone; common. Fl. Feb.—April,and September; white. In var. 8, yellow. 

Also in Peninsular India and Malay Is. 

Very variable. In the dry country the 1. are narrower and the fruit 
smaller. Var. 8 perhaps distinct. There is no specimen or drawing of 
this in Hermann’s collection. 

Wood hard, heavy, yellowish, very strong, and, being also durable, is 


used (in Northern Prov.) for constructing granaries, which are said to 
last for fifty years. 


22. BYRSOPHYLLUM, #7. /. 


A small tree, stip. large, deciduous, fl. large, dicecious, in 
small terminal cymes; cal.-limb short, truncate; cor.-tube 
long, contracted and villous at mouth, lobes 5, twisted in 
bud; stam. 5, inserted in cor.-tube, anth. sessile, included; ov. 
2-celled, with numerous ovules immersed in fleshy placentas; 


Randia.] Rubtiacee. 320) 


fruit a large 2-celled berry; seeds few, flattened, immersed in 
pulp.—Sp. 2; the other one in S. India. 


B. ellipticum, Bedd. Forest. Man. 134-2 (1875 ?). 
Stylocoryne elliptica, Thaw. Enum. 421 and 154 (Coffea ?). C. P. 3482. 
Fl. B. Ind. iii. 107. Bedd. Ic. Pl. Ind. Or. t. 55 SID (not good). 


A small tree, 10-12 ft., branchlets stout, marked with close 
leaf-scars, young parts glabrous; |. closely placed, large, 4-9 
in., oval, tapering to base, shortly and abruptly acuminate, 
obtuse, quite glabrous on both sides, rather coriaceous, paler 
beneath, lat.-veins inconspicuous, petiole about I in., stout, 
stip. I in., oblong, acuminate, glabrous, deciduous; fl. iW, 
large, in sessile, trichotomous, terminal cymes, ped. stout, 1 4-4 
in.; cal.-limb ‘glabrous, truncate, without Poa) cor.-tube 
over I in., slightly wider upwards, lobes 3-3 in., oval-oblong, 
obtuse; berry pyriform-globose, 1} in., rather tapering to 
base, smooth; seeds few, large. 

Moist low country; very rare. Hiniduma Pattu (Thwaites). FI. 
Sept.; white. 

Endemic. 


I have not met with this, and have nothing but the C. P. specimens, 
which include no fem. flowers. Turns black in drying. 


23. RANDIA,* Z. 


Shrubs or small trees, rarely scandent, one leaf of opp. pairs 
often absent, stip. small, triangular; fl. rather large, solitary 
and terminal or in leaf-opposed cymes; cal.-limb short, wide, 
segm. large and leafy or very small; cor.-lobes 5 (rarely 6 or 
7), longer than tube; anth. nearly sessile, long, exserted; ov. 
2-celled, with many or several ovules in each cell sunk in 
fleshy placentas, stigma large, clavate or fusiform, 2-fid.; fruit 
a berry, 2-celled, with numerous seeds immersed in pulp.— 
Sp. 90; 18 in FZ. B. Ind. 

Spiny. 
Fl. 1 or 3, terminal; cal.-limb persistent. 

Fl. over 14 in.; fr. over 2 in. 

Fl. and fr. both under 1 in. 

Fl. in cymes; cal.-limb deciduous 
Without spines. 

L. narrow-lanceolate, cymes axillary 

L. oval, cymes terminal or pabepivecd 


R. ULIGINOSA. 
. R. DUMETORUM. 
. R. MALABARICA. 


R. GARDNERI. 
R. RUGULOSA. 


Vik wndH 


* Isaac Rand was Curator of the Apothecaries’ Garden at Chelsea 
from 1724 till his death in 1743. 


330 Rubiacee. [Randia. 


1. R. uliginosa, DC. Prod. iv. 386 (1830). Bt-kukuruman, 
Wadiga, S. 

Gardenia uliginosa, Retz., Moon Cat. 19. Thw. Enum. 159. C. P. 2936. 

Fl. B. Ind. il. 110 (not given for Ceylon). Wight, Ic. t. 397. 


A small tree, much branched, branches thick, horizontal, 
many of them short and terminating in 2 pair of short sharp 
spreading spines, bark rough, shredding off in fibres, reddish- 
brown; |. fascicled on suppressed branchlets, 3-5 in., spathu- 
late-oblong or lanceolate, tapering to base, obtuse, glabrous 
on both sides or slightly pubescent beneath, thin, reticulate- 
veined, petiole very short, stip. acute; fl. large, 14-2 in. diam., 
solitary at end of suppressed branches, ped. short; cal.-limb 
broadly tubular, segm. very shallow, rounded, ciliate; cor. 
glabrous outside, tube with ring of hairs at mouth, lobes 5-7, 
orbicular, much overlapping; fruit large, 24-3 in., pyriform 
ovoid, smooth, pale or orange-brown, 2-celled, pericarp thick; 
seeds numerous, closely packed in pulp. 

Margins of tanks, &c., in the dry region; rather rare. Trincomalie; 
Kantalai; Dambulla. Fl. Aug.—Sept.; white. 

Also in India and Burma. 

The flesh of the large yellow fruit is used in curries, but it is astringent. 


2. R. dumetorum, Lam. ///. Genres, 11.227 (1793). Zukuruman, 
S. Karai, 7. 

Moon Gat. 19: Thw. Enum: 159, GC. BP. 1652! 

Fl. B. Ind. iii. 110. Roxb. Cor. Pl. t. 136 (Gardenia dumetorum). 
Wight, Ic. t. 580. 

A shrub or small tree, branches horizontal, rigid, many of 
the lat. ones suppressed and very short, spines in opp. pairs 
coming off immediately above the branchlets, 4-1 in. long, 
horizontal, woody, strong, very sharp; 1. usually fasciculate 
on the suppressed branchlets, nearly sessile, 1-2 in., obovate- 
oval or spathulate, tapering to base, obtuse, apiculate, glabrous 
or slightly pubescent, thin, reticulate-veined, stip. acuminate; 
fl. 1 in. diam., 1-3 at ends of suppressed branchlets, ped. short; 
cal.-limb broadly tubular, from nearly glabrous to very hairy, 
seem. leafy, ovate, acute, imbricate, glabrous or slightly hairy ; 
cor. hairy outside; tube as long as cal., lobes rounded, spread- 
ing; fruit globose or broadly ovoid, about # in., crowned with 
large cal.-limb, pilose, yellow, 2-celled, pericarp thick; seeds 
flat, surrounded with pulp. 

Dry region; very common, especially in sandy places near the coast. 
F]. August; yellowish white. 

Also in Peninsular India, Burma, Java, S. China, E. Trop. Africa. 

Usually a mere spiny bush. Fruit said by Nevill to be used as a fish- 


poison by the Veddas. The pulp of the fruit is a safe and much-used 
emetic, and the root is given in bowel complaints. 


Randia.] Rubtacee. Bel 


3. R. malabarica, Zam. ///. Genres, ii. 226 (1793). Pudan, 7, 

Griffithia fragrans, W. and A., Thw. Enum. 158. Gardenia fragrans, 
Moon Cat. 19. C. P. 1234. 

Fl. B. Ind. iii. 111. Wight, Ic: t. 310. 


An erect, much-branched shrub, with many branchlets 
reduced to short sharp spreading axillary spines; 1. 14-3 in., 
lanceolate or oblong-lanceolate, tapering to base, subacute, 
quite glabrous, petiole }-3 in., stip. acute, quickly caducous ; 
fi. 2-1 in. diam., in few-flowered, spreading, glabrous, leaf- 
opposed cymes, bracts small, connate in pairs, and so cup- 
shaped; cal. glabrous, limb wide, cupular, deciduous, segm. 5 
or 6, narrowly triangular, acute; cor. hairy at mouth, lobes 
lanceolate, acute; anth. very long, exserted; fruit globose, 
4-1 in, marked on top with ring-scar of cal.-limb; seeds 
about $ in each cell, closely packed. 

Dry and intermediate region; common. FI. April, September; 
lemon-yellow. 

Also in S. India. 

Moon gives ‘ Maha-géta-kulu’ as the S. name for this. A dwarf, very 
spinous state, with small rotundate leaves, which rarely flowers (much 
like a Canthium in appearance) is very common. 


4. R. Gardneri, 2. 7. in Gen. P/. ii. 88 (1873). 

Griffithia Gardneri, Thw. Enum. 158. C. P. 657. 

Fl. B. Ind. iii. 112. Bedd. Ic. Pl. Ind. Or. t. 38 (Grzfithza). 

A rather small tree, with very many slender, drooping, 
cylindrical branchlets, quite without spines, young parts 
glabrous; 1. 34-4 in., narrowly oblong-lanceolate, acute at 
base, attenuate-caudate, acute at apex, often somewhat fal- 
cate, glabrous, shining above, paler beneath, petiole very 
short, stip. very short, apiculate; fl. 4-2 in. diam., on long 
ped., cymes rather lax, spreading, from axils of past and 
present l., bracts opp., connate, small; cal. pilose, limb short, 
almost truncate, segm. tooth-like ; cor.-tube hairy at mouth, 
lobes linear- aera acute ; fruit pyriform-globose or broadly 
ovoid, nearly } in., seeds numerous. 

Moist region up 5 3000 ft.; rather rare. Colombo; Karawita; Udu- 
gama; Hantane; Deltota. Fl. April; white or pale yellow. 


Endemic. 
Closely related to 2. densiflora, Benth. of Burma and Malaya. 


R. rugulosa, ///. f. in Fl. B. Ind. iii. 113 (1880). 
t iffithia rugulosa, Thw. Enum. 159. C. P. 2 
Fl. B. Ind. iii. 113. Bedd. Ic. Pl. Ind. Or. t. 37 (Griffithia Spectosa). 
A climbing shrub, with many long drooping branches 
quite without spines; 1. 3-44 in., oval, acute at base, shortly 
acuminate, acute at apex, glabrous, with tufts of hair in axils 
of veins beneath, rather thick, veins impressed above, pro- 


B22 Rubiacee. [ Gardenia. 


minent beneath, petiole }-} in., stip. small, cuspidate; fl. over 
4 in. diam., ped. pubescent, rather longer than cal., cymes 
paniculate, spreading, either terminating short branchlets or 
leaf-opposed (one |. of the opp. pair being suppressed), bracts 
numerous, connate, cuspidate; cal. adpressed-pilose, segm. 
tooth-like, distant; cor.-lobes oblong, acute; fruit 4 in, 
globose. 


Moist low country below tooo ft.; rather rare. Galle; Kottawa; 
Ratnapura ; Ambagamuwa. Fl. March and April, white. 
Also in S. India. 


24. GARDENTA,* /. 


Small trees, stip. large or small, usually connate and 
sheathing; fl. very large, solitary; cal.-limb long, tubular, 
segm. various or 0; cor.-tube long, lobes 6-9, much twisted in 
bud; stam. as many as cor.-lobes, anth. nearly sessile ; ov. 
1-celied with numerous ovules on 3 or 5 parietal placentas, 
stigma large, clavate, lobed; fruit large, outer coat fleshy, 
endocarp bony, splitting into 3 or 5 valves along the placentas ; 
seeds very numerous, compressed, packed in pulp.—Sp. 60; 
14 in FZ. B. Ind. 


Placentas 3. 


Fruit not ribbed . 2 : 4 ‘ . I. G. LATIFOLIA. 
Fruit ribbed . : : A 5 : 3 . 2. G. CORONARIA. 
Placentas 5. ; : : : < . . . 3. Ga DURE 


1. G. latifolia, Az¢. Hort. Kew. i. 294 (1789) non Roxb. Galis, S. 

Moon'Gat 195) fhwe Enum 421. C2 Eso 

Fl. B. Ind. iii. 116, 120. Wight, Ic. t. 759 (not good). 

A small, many-stemmed tree or large shrub, bark smooth, 
pale-grey, flaking off in small rounded pieces, young parts 
glabrous but coated with a resinous exudation; |. crowded 
at ends of branches, 4—7 in., obovate-oval, slightly tapering to 
base, rounded or very shortly acuminate and obtuse at apex, 
slightly scabrous on both sides, especially on the veins (often 
pubescent) beneath, stiff, thin, bright apple-green, paler 
beneath, lat. veins 16-20 each side, parallel, very prominent 
beneath, petiole very short, stip. adnate to base of petiole, 
connate forming a tube which is thin, truncate, at first en- 
closing young bud, but after fall of leaf separating at base, 
and forming a loose ring round the stem; fl. very large, soli- 
tary, apparently terminal (really leaf-opposed), on short stout 


* Commemorates Alexander Garden, M.D., F.R.S., who practised as 
a physician in Carolina, and corresponded with Linnzus. Died 1791. 


Gardenia.) Rubiacee. 212) 


peduncle; cal.-limb 2in., broadly tubular, slightly dilated at 
top, either truncate (without segm.) or segm. 7-0, acute, con- 
duplicate, recurved, usually with 7-9 prominent ridges or 
wings, margin ciliate; cor.-tube 2-3 in., straight, lobes 7-9, 
13-2 in., horizontal, obovate-oblong, obtuse; anth. half exserted; 
stigma very large, exserted, 3-lobed; fruit nearly globose, 
about 14 in. diam., quite without ribs, crowned with the large 
erect, persistent, rigid, enlarged, winged cal.-limb which is 
#in. long, slightly pubescent, pale green, epicarp dry-fleshy, 
valves of endocarp 3, bony, yellow, polished within; seeds 
small, very numerous, horizontal, flattened, pale brown, pulp 
purplish-grey filling the fruit. 

Moist low country in damp places; rare. Colombo (Ferguson); 
Kalutara; Pelawatte; Pasdun Korale, in several places. FI. March; 
pale lemon-yellow, becoming orange when fading. 

Also in W. and S. India. 

This can scarcely be G. latifolia of Roxburgh as figured in Cor. PI. t. 
134, which shows a more ovoid fruit without a persistent cal.-limb. The 
Fl. B. Ind. seems to be doubtful as to its proper name. 

The buds and flowers exude a copious yellow resin, which when dry 
becomes powdery, and covers the branches, leaves, and fruit with a sort 
of =cut Wood hard, close-grained, rather heavy, yellowish, easily 
Worked. 


2. G. coronaria, Ham. in Symes, Emb. to Ava, iii. 307 (1800) ? 

G. carinata, Thw. Enum. 159 (? Wall.). Trim. in Journ. Bot. 163. 
CPi 3018. 

Meeeesne. 3. 317,120. Ham. |: c. t.22: 

Much like G. latifolia, from which it differs thus: 1. 
smaller, 3-4 in. (rarely longer), slightly pubescent above, and 
usually densely so beneath (sometimes nearly glabrous) ; 
cal.-limb about 4in., angled, pubescent; cor.-tube 3-4 in, 
Narrow, pubescent, lobes 7, I-14 in.; fruit 14 in., ovoid, 
tapering to base, crowned with cal.-limb, and with its angles 
(usually 6) continued down the fruit as long, blunt, vertical 
ribs, pubescent, valves of endocarp as in the last, seeds 
numerous, pulp yellow, filling the fruit. 

Exposed rocks in the dry region; rare. Batticaloa; about Bibile 
and Nilgala; Trincomalie (Nevill). Fl. June; pale yellow. 

Also in Burma. 

This plant is very closely allied to G. /atifolia, and may be merely a 
dry-country variety, the longer narrower cor.- tube and shorter lobes, with 
the distinctly ribbed fruit, being the chief differences. It does not quite 
agree either with G. carinata or G. coronaria, but I hesitate to make 
another species where there seem already too many. 


3. G. turgida, Roxb. Hort. Beng. 15 (1814). 
Trim. in Journ. Bot. xxvii. 163. 
Fl. B. Ind. iii. 118. Wight, Ic. t. 579. 


334 Rubracee. [ Scyphostachys. 


A small tree, with thick branches and numerous stout, 
short, lat. twigs, bark smooth, reddish-grey ; |. very crowded 
at ends of branchlets, sessile, 3-4 in., obovate-oval, much 
tapering to base, subacute, glabrous on both sides, rather 
coriaceous, stip. small, triangular, slightly connate, cuspidate, 
caducous; fl. not seen; fruit large, 1}-1#in., nearly globose, 
often bluntly pointed but not beaked, glabrous, rough, with 
small worts, pale greyish-brown, epicarp thick, fleshy, valves 
of endocarp 5, bony; seeds very numerous. 

Dry country; very rare. Only met with as yet on rocky ground at 
Nilgala, Uva; in fruit Jan. 1888. Fl. white. 

Also in India and Burma. 


Not having seen the flowers, some doubt as to the name of this may 
be felt. 


25. NARGEDIA,* Bedd. 


A large shrub, stip. acute, deciduous, fl. few, axillary; cal.- 
limb very short, segm. 4 (rarely 5), very short; cor.-tube 
short, with lines of hair within, lobes 4 (rarely 5) shorter than 
tube, twisted in bud; stam. inserted in cor.-tube, anth. nearly 
sessile, linear; ov. 2-celled with 2 ovules in each cell, stigma 
clavate, bilobed; fruit a dry berry, 1-celled; seeds usually 2, 
endosperm deeply ruminate.—Monotypic and endemic. 


Iv. macrocarpa, Ledd. For. Man. cxxxiv. 2 (1873 ?). 

Lflyptianthera macrocarpa, Thw. Enum. 157. C. P. 2709. 

DB lind si 1224 5 Beddy El Sylvats 320! 

A shrub, 8-10 ft., bark smooth, polished, yellow, young 
parts glabrous; 1. large, 54-64 in., oblong-lanceolate, acute at 
base, acuminate, acute, quite glabrous, petiole 4 in., stip. 
narrowly triangular; fl. rather small, 1-3 in the axils, ped. 
shorter than petiole, smooth; cal.-limb glabrous, short, segm. 
very shallow, cor.-tube broad, lobes short, rounded, hairy in 
the middle; fruit # in., broadly ovoid, slightly tapering at 
bothvends-)seedsp 2) oi 3° 

Moist low country; very rare. Raitunwelle; Ambagamuwa; Singhe 
Raja Forest. Fl. Feb.; cream-coloured. 

Endemic. 

I have not seen this living. 


26. SCVPHOSTACHYS, Jiw. 


Shrubs, stip. connate, fl. small, in axillary or supra- 
axillary bracteate spikes or clusters; cal.-limb short, segm. 4; 


* An anagram of Gardenia. 


Scyphostachys.| Rubiacee. 33 
cor.-tube short, hairy within, lobes 4, small, twisted in bud; 
stam. 4, anth. nearly sessile, included; ov. 2-celled, with 4 
pendulous ovules in each cell, stigmas 2, pilose; fruit a berry, 
1-celled, with I or 2 seeds.—Endemic Genus, the following 
the only species. 


Stip. hairy within; berries purple F : . I. S. PEDUNCULATUS. 
Stip. glabrous within ; berries scarlet . i . 2. S. COFFZOIDES. 


1. S. pedunculatus, 7hw. Enum. 157 (1859). 

Es ee 

Fl. B. Ind. iii. 122. 

A shrub, branches slender, cylindrical, slightly rough; 1. 
34-54 in., lanceolate-oblong, abruptly narrowed at base, 
shortly acuminate, acute at apex, glabrous above, pubescent 
on midrib beneath, petiole } in., stip. rather large, connate, 
ovate, acute, lined with long silky hair, striate outside; fl. 
rather small, sessile, few, in small supra-axillary, stalked 
spikes, peduncle $—? in., pubescent, deflexed, bracts rather 
large in several opposite pairs, connate, sheathing and over- 
lapping, pubescent, forming a short imbricated catkin; cor.- 
lobes short, rotundate, berry tipped with cal.-segm., nearly 
2 in., ovoid, very soft, dark purple; seed usually solitary, 
narrowly oblong. 


Moist low country; very rare. Only found at Hewesse, Pasdun 
Korale. Fl]. April; white. 


Endemic. 


2. S. coffeeoides, 7Thw. Enum. 157 (1859). Wal-kopi, S. 
er. 27 10: 


Fl. B. Ind. iii. 122. Bedd. FL. Sylv. Anal. Gen. t. 16, f. 6; Ic. Pl. Ind. 
Or. t. 240. 

A shrub with spreading virgate branches, bark smooth, 
shining, yellow; 1. 34-5 in., lanceolate, acute at base, acuminate, 
acute at apex, quite glabrous, somewhat undulate at margin, 
subcoriaceous, petiole very short, stip. connate, acute, puberu- 
lous; fl. small, sessile in small dense axillary clusters or 
solitary, each surrounded at base by an involucre of 2 connate 
puberulous bracts; cal.-limb very short, hairy within, lobes 
rotundate, recurved; berry about 2 in., oblong-ovoid, sup- 
ported by persistent bracts, truncate at top, scarlet; seeds 2, 
oblong-ovoid. 

Moist low country; rare. About Ratnapura; Badamua, S. Prov. Fl 
March, April; white. 

Endemic. 

The native name, ‘Wild Coffee,’ is very appropriate as far as the 
foliage goes. 


226 Rubiacee. [Diplospora. 


27, DIPLOSPORA, DC. 


Trees, stip. triangular, persistent; fl. small, polygamo- 
dicecious, in sessile axillary clusters; cal.-limb very short, 
segm. 4; cor.-tube short, lobes 4 (rarely 5) twisted in bud; 
stam. 4 (rarely 5) inserted at mouth of cor., anth. exserted ; 
ov. 2-celled, with 1-5 ovules in each cell; fruit a 2-celled 
berry, with few rather large seeds.—Sp. 10; all in FZ. B. Ind. 


I. D. Dalzellii,* Hk. f, Fl. B. Ind. iii. 123 (1880). Wella, 7. 
FPLATE L.] 

Discospermum Dalzelliz, Thw. Enum. 158. D. spherocarpa, Bedd. 
Hom Manter3497) (CPs sor 

HIS Bs Ind ni-123: 

A small pyramidal tree with numerous ascending branches, 
bark very smooth, white or yellowish, twigs compressed, 
thickened at nodes; 1. 3-4 in., lanceolate or oval, acute at 
base, acuminate, obtuse at apex, glabrous, shining above, 
paler beneath, subcoriaceous, stiff, petiole + in., furrowed 
above, stip. small, broadly triangular with a long terminal 
bristle, sub-persistent; fl. small, nearly sessile, in dense 
cymose clusters; cal. glabrous, segm. triangular, subacute ; 
cor.-tube short and broad in fem. fl. longer and narrower in 
male fl., lobes nearly as long as tube, oval-oblong, recurved ; 
berry 3-3 in. or rather more, nearly globose, smooth, dark 
green, crowned with cal.-segm., with several seeds in each cell. 

Low country, principally in the dry region; rather common. FI. 
April; greenish-white. 

Endemic, but very close to D. spherocarpa of S. India. 

When young, this is very like JZemecylon in habit and foliage. Wood 
hard, heavy, fine-grained, white, smooth. 


2. D. erythrospora, edd. For. Man. cxxxiv. 3 (1873 ?). 
Discospermum erythrosporum, Thw. Enum. 158. C. P. 2510, 3481. 
FI. B. Ind. ii. 124. 

A much-branched tree, branchlets stout, much thickened 
at nodes, twigs compressed, glabrous; |. 24—4 in., oval, acute 
at base, usually rounded sometimes bluntly acuminate at 
apex, margin usually recurved, glabrous, coriaceous, with 
glandular pits in axils of veins beneath, petiole very short, 
stip. triangular, thick, sharply acuminate, the basal portion 
persistent; fl. as in 1,male rather the larger; cal.-limb. almost 
truncate, the segm. minute, tooth-like ; cor.-lobes oblong, 
obtuse, much longer than tube; berry less than } in., ovoid or 


* After N. A. Dalzell, Conservator of Forests, Bombay, and author 
(with A. Gibson) of ‘The Bombay Flora’ (1861). Died 1878. 


Guettarda.] Rubiacee. 227i 


subglobose ; seeds I-5 in each cell, oblong, bright red, im- 
mersed in purple pulp. 

Lower montane zone; very rare. Ambagamuwa Dist. at about 3000 
ft. (Thwaites); Haputale at about 5000 ft. Fl. Feb., Sept.; pale greenish- 
yellow. 

Endemic. 


28. SCYPHIPHORA, Gaertn. 


A small tree, stip. short and broad, fl. small, in short 
axillary cymes; cal.-limb cup-shaped, persistent, segm. very 
small, tooth-like ; cor.-tube short, lobes 4 (or 5), imbricate ; 
stam. 4 (or 5), inserted at mouth of cor.; anth. sagittate at 
base; ov. long, 2-celled, with 2 ovules in each cell, one erect, 
the other pendulous; fruit a drupe, with 2 connate pyrenes, 
each with 2 seeds.—Monotypic. 


S. hydrophylacea, Gaevin. f. Fruct. iii. 91 (1805). 

Epithinia malayana, Jack, Thw. Enum. 157. C. P. 1665. 

Fl. B. Ind. iii. 125. Griff. Ic. Pl. Asiat. t. 478 (repeated at t. 644 A). 

Small tree or shrub, branchlets stout, somewhat thickened 
at nodes, sub-quadrangular, glabrous, young parts exuding a 
resinous secretion; |]. 2-24 in., obovate-oval, tapering to base, 
rounded at apex, glabrous, rather fleshy, obscurely veined, 
petiole $in., readily disarticulating, stip. short, rounded; fl. 
small, on very short ped., cymes small, on short peduncles, 
cal.-limb glabrous, almost truncate; cor.-tube hairy within, 
lobes shorter than tube, ovate, acute; anth. exserted; drupe 
small, about 4in., crowned with persistent cal.-limb, obovate- 
ovoid, slightly compressed, with 8 (or 10?) thick, blunt, ver- 
tical ridges, glabrous, pyrenes crustaceous. 


Mangrove swamps; rare. Only seen from Jaffna (Gardner). FI. 
white. 


Occurs also on the Carnatic coast, Malaya, Andamans, N. Australia, 
and New Caledonia. 
Remarkably like Lusnnitzera, with which it grows. 


29. GUETTARDA, Z. 


Small tree, stip. ovate, caducous; fl. large, unisexual or poly- 
gamous, in long-stalked axillary cymes; cal.-limb tubular-cup- 
shaped, truncate, deciducus; cor. densely pubescent, tube long, 
lobes 5-9, imbricate; stam. same number as pet., inserted in 
cor.-tube, fil. 0; ov. 5-g-celled with a single pendulous ovule 
in each cell, stigma capitate; fruit a drupe, stone woody, with 
5-9 sharp angles, and as many cells.—Sp. 45, all, but this one, 
American only. 

PART Il. Z 


33 8 Rubiacee. [ Zmoncus. 


G. speciosa, L. SZ. P/. 991 (1753). Nil-pitcha, S. Panir, 7. 

Moon Cat. 14. Thw. Enum. 153. C. P. 2990. 

iP), 1B, Ibavel, i, TAG,  Warelnre, WE AO, Biol INGE Th, U3IOKs 

A small tree, with stout pubescent branchlets, marked with 
l.-scars; 1. rather crowded at ends of branchlets, large, 5-8 in., 
obovate-rotundate, somewhat cordate at base, usually rounded 
at apex, pubescent on both sides especially beneath, pink 
when young; petiole 1-14 in., pubescent, stip. ovate, acute ; 
fl. nearly sessile, few, crowded on the short arms of a bifid 
cyme, peduncle long, pubescent; cal. silky-pubescent; cor. 
glabrous within, lobes shorter than tube, obtuse, spreading ; 
anth. sessile, small, included; drupe hard, depressed-globose, 
under I in., somewhat lobed, slightly pubescent. 

On the coast; rare. Galle; Kalutara; and often cultivated in gardens 
near the sea. Fl. Nov.—Jan.; white. 


Tropical shores of both worlds. 
The large jasmine-like flowers are extremely sweet-scented. 


30. TIMONTUS, DC. 


Small tree or shrub, stip. triangular, caducous; fl. rather 
small, polygamo-dicecious, axillary, fem. often solitary, male 
in bifid cymes; cal.-limb small, segm. 4; cor. tubular, rather 
longer in male, lobes 4, valvate; stam. 4 inserted in cor.-tube, 
fil.o; ov. usually 12-celled with 1 pendulous ovule in each 
cell, style with 4 or more branches; fruit a drupe with 10-12 
gompressed pyrenes.—Sp. 20; 3 in FZ. B. Ind. 


T. Jambosella, Zw. Enum. 153 (1859). Peddimella, 
Angana, S. 

Nelitris Jambosella, Gaertn. Fruct. i. 134. Eupyrena glabra, W. & 
Ae COG W423 -m Como 

Fl. B. Ind. iii. 127. Bedd. Ic. Pl. Ind. or t. 190. 


A small tree-like shrub, 8-10 ft., branchlets slender, cylin- 
drical, glabrous, marked with leaf-scars; |. 2-5 in., rather 
closely placed, oval or lanceolate, acute at both ends, glabrous 
but often pubescent on veins beneath, and with tufts of hair 
in their axils, venation pellucid, margin sometimes recurved, 
petiole {-?in., stip. triangular, acuminate; fl. sessile, with 2 
very small bractlets beneath, male on short spreading branches 
of cyme, fem. solitary or three together, peduncle about I in., 
cal. silky; cor. pubescent outside, smooth within, fleshy, lobes 
shorter than tube, recurved; anth. sessile, linear, included ; 
drupe 4-4 in., nearly globose, lobed or ribbed over pyrenes, 
smooth, dark green, tipped with cal.-segm. 


Dichilanthe.| Rubiacee. 330 


Moist region, rarely in the low country, principally in the lower mon- 
tane zone, but extending up to 6000 ft.; common. FI. March, April, 
September; yellow. 

Also in the Andamans, Malaya, and Mauritius, but not in Peninsular 
India. 

The genus Zimonzus dates only from 1830, and should rightly be 
superseded by Ve/¢¢rzs (1788) ; for Gaertner’s figure of the fruit (t. 27, f. 5) 
shows that this was the plant intended. He has in the text, however, 
confused it with some Awgenza, the specimens having been named ‘ Wal- 
jambu.’ The name thus became applied by Decandolle to a genus of 
Myrtacez (properly Decasfermumt, Forst.). 


31. DICHILANTHE, 7hw. 


A tree, stip. connate into a ring, persistent; fl. large, in 
terminal heads; cal.-limb tubular, curved, with 5 hairy scales 
at the mouth, segm. 5, longer than tube, slightly unequal; cor. 
irregular, curved, tubular below, funnel-shaped above, 2-lipped, 
upper lip 2-toothed, lower lip 3-toothed; stam. 5, inserted near 
mouth of cor., disk large, annular, fleshy; ov. 2-celled with 
I pendulous ovule in each cell, style very long, stigma clavate, 
bilobed ; “fruit obovoid, gibbous, 2-celled, pericarp thick.”— 
There is a second species in Borneo. 


This genus with its irregular cor. might well stand in Cafrzfoliacee, 
where Thw. and Beddome, and at one time J. D. Hooker, placed it. 


D. zeylanica, 7iw.in Kew Journ. Bot. viii. 270 (1856). 

Thw. Enum. 136. C. P. 3422. 

Fl. B. Ind. iii. 128. Kew Journ. Bot. viii. t. 8,f. a. Bedd. Fl. Sylv. 
Anal. Gen. t. 15, f. 4. 


A tree, usually small, branchlets stout, swollen at nodes, 
bark smooth, grey; |. 4-6 in., lanceolate, acute at base, shortly 
acuminate at apex, quite glabrous, coriaceous (with fine raised 
reticulation when dry), petiole very short, stout, stip. short, 
truncate; fl. sessile, about 6-8 together in a very shortly 
stalked terminal head and the cal.-tubes glued together with 
resinous exudation; cal.-segm. linear-lanceolate, acuminate, 
spreading; cor. 1}-I} in., strongly curved downwards, silky 
outside, lips short; fil. short, anth. small, oblong; style much 
exserted; fruit not seen. 


Moist low country; very rare. On the summits of Hiniduma Kande 
and Nillowe Kande; Bambarabotuwa, Sabaragamuwa. FI. March- 
June; scarlet. 

Endemic. ; 

A very remarkable and anomalous plant; the persistent ring-like stip. 
become coated with resin, and form nodosities on the branches. 


340 Rubtacee. [Knoxia. 


32. RNOXIA,* Linz. 


Annual or perennial herbs or undershrubs, stip. with rigid 
bristle-like teeth ; fl. sessile or nearly so, small, dimorphic, 
numerous, in terminal cymes or spikes; cal.-segm. 4, one often 
greatly enlarged; cor.-tube hairy at mouth, lobes 4, short, 
valvate; stam. 4; ov. 2-celled, with 1 pendulous ovule in each 
cell, stigmas 2; fruit very small, dry, of 2 indehiscent carp. 
separating from a central persistent columella and usually also 
from one another.—Sp. 8; all in AZ. B. Ind. 


Cal.-segm. nearly equal. 
L. sessile, linear-lanceolate . 5 : 5 . I. K. CORYMBOSA. 
E. stalked, oval . : : : . 2. K. MOEUIS: 

One cal.-segm. greatly enlarged. 

Fl. in long spikes ; fruit not compressed 
Fl. in capitate cymes; fruit compressed 


The fruit is remarkably like that of an Umbellifer. 


I. K. corymbosa, Willd. Sp. Pi. i. 582 (1797). 

Moon Cat. 10. KX. stricta, Thw. Enum. 152 (? Gaertn.). C. P. 3620. 

Fl. B. Ind. iii. 128. Wight, Ill. t. 128. 

An erect annual, 2-4 ft. high, almost unbranched, stem 
rather stout, with elongated internodes, densely and softly 
hairy; 1. few, distant, 3-44 in., sessile or very nearly, linear- 
lanceolate or linear, much attenuate at both ends, acute, 
usually hairy on both sides, lat. veins very oblique, stip. with 
3-5 filiform hairy bristles; fl. very small, numerous, on sides © 
of slender branches of large lax corymbose cymes; cal.-segm. 
minute, triangular, none enlarged; cor.-lobes obtuse; fruit 
zs in., oblong-ovoid, scarcely compressed, splitting from the 
base into two carp., carp. plano-convex, glabrous, not ribbed 
on back. 

Low country and up to 4000 ft.; rather common. Bintenne; Ugal- 
duva Valley; Ramboda. Moon’s locality is Walapane. FI. Feb.; violet. 

Also in India, Malay Islands, and Trop. Australia. 

The mericarps readily separate from the columella when ripe with the 
slightest pressure. 


2. K. mollis, W. & A. Prod. 439 (1834). 

K. corymbosa, Thw. Enum. 151 (non Willd.). C. P. 3619. 

F]. B. Ind. iii. 129. 

Annual, 2-4 ft., branched, stem erect, pilose or nearly 
glabrous; |. 24-44 in., oval or ovate-oval, tapering at both 
ends, acute, slightly hairy on both sides, petiole 4-1 in., stip. 


K. ZEYLANICA. 
K. PLATYCARPA. 


Bo 


* Commemorates Robert Knox, who was in captivity among the ~ 
Kandians from 1659-79, and wrote an ‘ Historical Relation of Ceylon,’ 
published in 1681, giving an account of his experiences and escape. 


Knoxia.] Rubiacee. 341 


as in the last; fl. in lax corymbose cymes; cal.-segm. 
triangular, acute, nearly equal or one sometimes slightly 
larger; cor.-tube tubular-funnel-shaped ; fruit about + in., 
broadly ovoid, compressed, readily separable from the colu- 
mella but without splitting into separate carp., smooth, 
black. 

Montane zone to 6000 ft.; rare. Uva (Thwaites); Maskeliya; Hak- 
gala. Fl. May, June, Sept.; white. 

Also in mountains of S. India. 

Fl. B. Ind. refers to this with doubt X. stricta, Gaertn. (Fruct. i. 122, 
t. 25,f. 8). Gaertner’s specimens were collected in Ceylon by Keenig, 
but could scarcely be this montane species. 

Ceylon specimens are often nearly glabrous. 


3. K. zeylanica, Z. Sf. Pl. 105 (1753). 

Herm. Mus. 11. Burm. Thes. 144. FI. Zeyl. n. 400. Moon Cat. Io. 
Thw. Enum. 152. W.andA. Prod. 440. C. P. 1690. 

FL. B. Ind. iii. 130. Burm. FI. Ind. t. 13, f. 2. 


A small undershrub, 1-2 ft., with a large, stout tap-root 
and ascending usually much-branched stems, branches erect, 
cylindrical, stiff, nearly glabrous, shining, purple; 1. numerous, 
1-2 in., broadly or narrowly ovate-oblong, nearly sessile, acute 
at both ends, glabrous, finely ciliate on margin, stip. adnate 
to base of leaf, short, with 3 or 5 filiform teeth bristly at end; 
fl. in terminal spikes which become greatly elongated with 
fruit; cal.-segm. acute, the outermost one enlarged into a 
stiff horn twice as long as ov., persistent ; cor.-tube 2 in., very 
slender, very woolly at mouth, lobes 4 as long as tube, 
broadly oval ; fruit about ;4; in., ovoid- globular, crowned with 
cal.-segm. (the large one as ‘long as fr.), glabrous, pale brown, 
carp. readily separating, plano-convex, not compressed, 
rounded and keeled down the back. 


Open ground in the moist low country; very common. FI. Feb.; 
white or pinkish. 

Endemic. 

Hermann gives ‘ Wissaduli’ as the native name for this, but no doubt 
incorrectly (see footnote, Part I. p. 146, under W7ssadula zeylanica). 
Moon has ‘ Ela-ratmul,’ but I do not find this generally used. 


4. K. platycarpa, Arn. Pug. 26 (1836). [PLATE LI.] 

Thw. Enum. 152. C. P. 1689. 

FL. B. Ind. iii. 131. 

A dichotomously branched, small shrub, 1-3 ft., branches 
erect, cylindrical, glabrous, twigs with two lines of pubescence; 
1. sessile, erect, 1-24 in., narrowly oblong-lanceolate or linear, 
subacute, glabrous, sometimes ciliate on margin, rather thick, 
stip. of 3 or 5, rigid, often hairy bristles, the middle one the 


342 Rubiacee. [ Canthium. 


longest ; fl. crowded in small heads terminating the branches 
of trifid terminal cymes, no bracts; cal. glabrous, segm. very 
small, linear-lanceolate, acute, one much enlarged, apiculate ; 
cor. glabrous outside, tube 3 in., slender, lobes half as long, 
oblong-lanceolate, subacute, thick; fruit 3-§ in., broadly 
ovoid, crowned with cal.-segm., slabrous, carp. readily sepa- 
rating, compressed, plano-convex, with a dorsal rib, dark 
brown. 

Var. B, hirsuta, 7hw./l.c. K. hirsuta, Arn. Pug. 26. C. P. 1688. 


Whole plant (except upper surface of 1.), including cor. 
outside, more or less hairy; fl. rather larger. 


Var. y, foliosa, 7iw. Enum. l.c. C.P. 51. 


Hairy ; 1. broader, oval, stip. with longer bristles; bracts: 
foliaceous ; cal.-segm. much longer ; cor. hairy outside. 


Var. 6, spicata, Thw. in Trim. Syst. Cat. 44. C. P. 3996. 


L. quite glabrous; cymes lengthening out into spikes ; 
fruit distant on the spikes, larger, more elongated, hairy. 

Montane zone in the patanas; very common. Var. y asylvan variety. 
Var. Orare. On False Pedro Mountain. FI. all the year; pale violet- 
pink or nearly white. 

Endemic. 

An extremely variable plant. The type, and, still more, var. 4, 
approach K. zeylanica. The dimorphism of the flowers (long-styled and 
short-styled) is striking—in one the style, and the other the stam., being 
exserted. 


33. CANTHIUM,* Lam. 


Trees or shrubs, with or without woody spines, stip. 
broadly triangular, acute, fl. small, in axillary cymes or 
fascicles ; cal.-limb very short, segm. 4 or 5, usually tooth- 
like, acute ; cor.-tube campanulate or globose, with a ring of 
deflexed hair within, lobes 4 or 5, valvate, reflexed; stam. 
inserted at mouth of cor.; ov. 2-celled with 1 pendulous 
ovule in each cell, stigma large, capitate, fruit a drupe, more 
or less didymous, with 2 hard pyrenes.—Sp. 70; 17 in #72. B. 
Ind. 


Branches without spines (also sometimes in 7). 
L. lanceolate, over 3 in. . : . . I. C. DIDYMUM, 
IL rotundate, under 3 in. : ; ; . 2. C. MONTANUM. 


ss * From ‘ Canti,’ the Malabar name for C. Rheediz, as given in Hort. 
alab. 


Canthium. Rubiacee. 343 


Branches with spines. 
Fl. usually 5-merous. 
Cor.-tube not globose. 
L. under I in., acute at base. : . 3. C. PUBERULUM. 
L. over I in., rounded at base. 
Fruit under 3 in. 
Beant Tin.’ . : 
Cor.-tube nearly globose 
Fl]. always 4-merous, very small 


1. ©. didymum, Gaertn. f. Fruct.iii.94 (1805). Pana-karawu 
Gal-karanda, Pandaru, S$. Vatchikuran, Yerkoli, 7. 
Psydrax dicoccos, Gaertn. Fruct. i. 125. Thw. Enum. 152. C. P. 


3419, 2607, 1667, 3511. : 
Fl. B. Ind. iii. 132. Wight, Ic. t. 1034 (C. wmbellatum). Bedd. FI. 


Sylv. t. 221. Gaertn. Fruct. i. t. 26, f. 2, and iii. t. 196, f. 5 (fruit only). 

A moderate-sized tree, bark smooth or vertically furrowed, 
whitish, branchlets numerous, drooping, thickened at nodes, 
glabrous, somewhat compressed, without spines; |. 34-44 in., 
lanceolate, acute at base, acuminate, obtuse, very glabrous, 
bright green above, paler beneath, subcoriaceous, rather 
undulate, lat. veins few, very oblique, pellucid, petiole 1-2 in., 
stip. triangular, acute, usually caducous; fl. small, on rather 
long ped., numerous, cymes lax, pedunculate, corymbose, 
bracts minute or none; cal. truncate or with 5 teeth, glabrous 
or pubescent; cor. campanulate, lobes 5, shorter than tube, 
lanceolate, acute; drupe }-4in., nearly globose, somewhat 
didymous when dry, purple, pyrenes kidney-shaped, very 
much warted, black. 

Var. 6, lanceolatum, 7hw. Enum.l.c. Webera lanceolata, Moon 
Cat. 19. C. lanceolatum, Arn. Pug. 24. C. P. 2513. 

L. narrowly oblong, stip. acuminate ; fruit smaller. 

Low country and up to 4ooo ft.; common. Var. 6, Kalutara (Moon) ; 
Kandy; Hunasgiriya. FI]. April; white. 

Also in India, Malaya, and S. China. 

A variable plant. C. P. 3419 is var. grandifolium, Thw. (from Cara- 
wita Kande), and C. P. 2607 (from Ratnapura) is var. vostratum, Thw.; 
but neither seem worth separate names. Gaertner’s Psydrax is described 
and figured from Ceylon specimens (in the Leyden Museum) collected by 
Hermann, but the plant does not seem to be noticed by other authors on 
Hermann’s collections. Another genus of Gaertner’s, Caranda pedun- 
culata, is based on this species. This is described in Fruct. ii. 17, and 
figured at t. 83. The supposed fruit is a large, hard, beaked leaf-gall, 
very frequently found, sometimes clustered, on the young branches of 
this tree. They are called ‘fruits’ even in FI. B. Ind. 

The wood is very hard, fine-grained, greyish, heavy, and has been 
called ‘ Ceylon Box-wood.’ The native names of this tree are not well 
ascertained. Moon gives ‘ Gal-séru’ for var. danceolatum. 


. C. RHEEDII. 

. C. MACROCARPUM. 
C. CAMPANULATUM. 
. C. PARVIFLORUM. 


NOU 


2. ©.montanum, 7/w. Enum. 152 (1859). 
C. P. 2610, 2653. 
Fl. B. Ind. iii. 132. 


344 Rubwacee. [Canthium. 


A large or moderate-sized tree, bark thin, blackish, flaking 
off in small pieces, branchlets numerous, stout, thickened at 
nodes, without spines, twigs quadrangular-compressed, gla- 
brous; 1. 14-24 in., rotundate, rounded at both ends, more or 
less revolute at margin, very stiff and coriaceous, quite gla- 
brous, veins few, reticulate, prominent beneath, petiole 
extremely short and stout, stip. short, broad, with a stout 
mucronate apex ; fl. on long ped., cymes umbellate on stout 
peduncles ; cal.-segm. short, broad, apiculate, cor.-lobes 5, as 
long as tube, acute, reflexed; stam. exserted; drupe 2 in., 
ovoid, very slightly didymous. 

Var. 8, minus, 7hw./.c. C. P. 1685. 


L. smaller, somewhat oval; fl. a little larger, ped. strongly 
reflexed after flowering. 

Forests of the montane zone from 4000-7000 ft.; rather common. 
Fl. Dec. to April; green. 

Endemic. : 

The wood is very hard, heavy, and fine-grained, and, like others in 
the hills, is called, incorrectly, ‘Wal-burutu’ by the up-country sawyers. 
It has been used for railway sleepers. 


3. ©. puberulum, 7/w. zz Fl. B. Ind. iii. 134 (1880). 

Gy 3995: 

A shrub, with virgate branches armed with sharp, supra- 
axillary spreading spines, pubescent when young, and with 
numerous short, suppressed branchlets; 1. small, 4-1 in., 
fasciculate on the suppressed branchlets, oval or oblong-oval, 
acute at both ends, slightly pubescent above, more so beneath, 
petiole short, slender, stip. small, hairy, with a filiform apex ; 
fl. very small, on rather long pubescent ped. with 2 bractlets 
near the base, I-3 on very short peduncles; cal. hairy, segm. 
long, linear-lanceolate, very acute, recurved ; cor.-tube shorter 
than cal., pubescent, lobes acute, recurved ; fruit not seen. 

‘Central Province’ (Fl. B. Ind.). There is no locality attached to the 


specimens in the Peradeniya Herbarium, which were collected in 1869. 
I have seen no others. 


4. ©. Rheedii, DC. Prod. iv. 474 (1830). 

Thw. Enum. 153. W.and A. Prod. 426. C. P. 1666. 

Fl. B. Ind. iii. 134. Rheede, Hort. Malab. v. t. 37. 

Asmall,much-branched bush, branchlets divaricate, roughly 
pubescent, usually armed with straight or slightly curved, sharp, 
supra-axillary spines 4-4 in. long; |. 1-2 in., ovate, cordate or 
rounded at base, acuminate, obtuse or subacute at apex, 
glabrous, or with a few bristly hairs beneath on veins; thin, 
lat. veins rather prominent beneath, with glands in the axils; 
fi. small, solitary or few in small sessile cymes; cal.-limb 


Canthium.] Rubiacee. 345 
spreading with 5 distant teeth, cor.-tube short and wide, the 
rings of hair within stiff and bristly, lobes 5, ovate, acute; 
anth. small, exserted; stigma bilobed; drupe broader than 
long, about 2 in. wide, didymous, compressed, bilobed at top, 
pyrenes strongly worted. 


Var. 8, minus, 7hiw./.c. C. P. 3420. 


L. smaller, $-1 in., spines very slender, straight; cor.-lobes 
long, attenuate. 

Moist low country up to 3000 ft.; rather common. Kalutara (Moon); 
Galle; Hunasgiriya. Wight (I. c.) records specimens from Rottler’s 
Herb. collected at Colombo April, 1806. Var. 8, Singhe Raja Forest. 


Fl. Nov.-Feb.; pale greenish-yellow. 
Also in S. India. 


5. ©. macrocarpum, 7hw. Enum. 152 (1859). 

Plectronia macrocarpa, Bedd. Ic. Pl. Ind. Or. 57. C. P. 2816. 

ieee. ii. 135. Bedd. Ic. Pl. Ind. Or. t. 238: 

A half-climbing shrub, branchlets long, cylindrical, rufous, 
at first pubescent, afterwards glabrous, spines supra-axillary, 
stout, compressed, either straight and $-#in., or shorter and 
strongly hooked, very sharp; |. 2-24 in. oval or lanceolate, 
rounded at base, very acuminate, acute, rough on both sides 
with minute, scattered, rufous, prickly hairs, petiole 4 in,, 
rufous-pubescent, stip. broad, hairy, with a long filiform point; 
fl. rather small, on short ped., cymes small, pedunculate, 
pubescent; cor.-tube campanulate, lobes 4 or 5, oblong, acute, 
longer than tube; stam. and style exserted, stigma very large, 
cubical; drupe 1 in. wide, broader than long, 2-lobed at top, 
compressed, didymous but with one lobe often suppressed ; 
pyrenes rugose. 

Very rare, and only found on the Hantane Range at about 3000 ft. 
Fl. Feb.; pale greenish-white. 

Endemic. First collected by Champion. 


6. ©. campanulatum, 7hw. Enum. 153 (1859). 

ak. 2472. 

Fl. B. Ind. iii. 135. 

A small semi-scandent shrub, with many very slender 
branches, adpressed-hairy when young, spines ? in., supra- 
axillary, numerous, slender, quite straight, very sharp; 1. often 
fasciculate on suppressed branchlets, small, {-14 in., ovate or 
oval, acute or rounded or rather cordate at base, obtuse but 
apiculate at apex, glabrous, petiole very short, stip. sharply 
pointed ; fl. usually solitary, or 2 or 3 together, ped. very 
slender, glabrous, jointed near the base, buds much pointed; 
cal.-limb saucer-shaped, segm. very acute; cor. thick, tube 


346 Rubiacee. [Zvora. 


inflated, nearly globose, lobes 5, rather longer than tube, 
lanceolate, with long acuminate points, reflexed; fl. very 
short; style rather exserted, stigma very large, truncate 3, 
drupe 3 in. wide, didymous, compressed, tapering to base. 

Low country to 3000 ft.; rare. Deltota; pe eS Madulsima. 
(Marshall Ward). Fl. Aug.; cream-coloured. 

Endemic. 

This is the only plant besides species of Cofa upon which the 


destructive parasitic fungus, Wemzleia vastatrix, was detected by Marshall 
Ward in Ceylon. 


7. ©. parvifiorum, Lam. Encycl. Meth. i. 602 (1783). Mara, S. 
Karai, 7. 

Moon Cat.10. Thw. Enum. 152. C. P. 1662. 

Fl. B. Ind. iii. 136. Rheede, Hort. Malab. v. t. 36. Roxb. Cor. Pl. t. 51. 


A much-branched shrub or small tree, branches usually 
stout and rigid, with the supra-axillary spines numerous, 
thick, woody, over 1 in. long, divaricate, often 3 in a whorl, 
but sometimes the branches are less rigid and quite with- 
out spines; I. often fascicled, variable, #?-24 in., rotundate or 
oval or oblong-oval, obtuse, glabrous, whitish beneath with 
conspicuous reticulate venation, petiole short, stip. short, with 
a long point; fl. very small, on long slender ped., cymes. 
many-flowered, sessile or pedunculate; cal.-segm. 4, acute ; 
cor.-tube inflated, globose, lobes 4, as long as tube, ovate, 
acute; drupe 3 in., ovoid, compressed, slightly didymous. 

Low country, especially in dry region; common: and up to 4000 ft.,. 
rarer. Fl. Nov.—March; yellowish. 

Also in S. India. 

The form in the moist region, especially at upper elevations, is much 
less rigid and often without spines, and has larger leaves, looking quite a 
different plant. 


The leaves are used in curries in Northern Prov. Wood very hard 
and close-grained. 


34. TZRORA,* Linn. 


Shrubs or small trees, stip. cuspidate, more or less per- 
sistent, fl. numerous, in terminal corymbose or paniculate 
cymes; cal.-limb undeveloped, segm. 4, small; cor. with very 
long slender tube, lobes 4, twisted in bud; stam. 4, inserted 
at mouth of cor., fil. very short, anth. small, linear; ov. 2- 
celled, with a single ovule in each cell attached to septum, 


* Taken by Linnzeus from Rheede, who states that Baldzeus in his. 
account of Ceylon says that the flowers (of /. coccinea) are offered in. 
sacrifices to ‘the god Ixora’ (= ? Isvara). 


Ixora.]| Rubtacee. 347 


style exserted, stigmas 2; fruit globose or sub-didymous, with 
2 plano-convex pyrenes.—Sp. 100; 37 in FV. B. Ind. 


Cal.-segm. as long as or longer than ov. 


L. under 3 in., cal.-segm. twice as long as ov. el CARVCINAY 
L. over 3 in., cal.- -segm. as long as ov. . : ee le MOEN ATTESIT 
Cal.-segm. shorter than ov. 
Cor. “tube under 3 Zin. : : : : . 3. I. PARVIFLORA. 
Cor.-tube over I in. 
L. over 4 in., acute at base. j : : > 4. 1 JUCUNDA, 
L. under 4 in., rounded at base . 5 5 . 5. 1. COCCINEA. 


1. I. calycina, 7iw. Enum. 155 (1859). 
One a Vt-F 
Fl. B. Ind. iii. 138. Bedd. Ic. Pl. Ind. Or. t. 97. 


A bush or small tree, much branched, branchlets nodulose 
with closely placed and very prominent |.-scars, young parts 
glabrous; |. 14-3 in., oval or oblong-oval, slightly cordate or 
rounded or acute at base, acute, margin often recurved, 
glabrous, rather glaucous beneath, rigid, petiole extremely 
short, stip. short with an almost spinous point, persistent; 
fi. nearly sessile, cymes rather small, trichotomous, glabrous, 
bracts linear; cal.-segm. more than twice as long as ov., linear- 
lanceolate, acute, glabrous; cor.-tube # in., ee slender, lobes 
short, broad, ovate, acute, recurved; fruit } in. broadly ovoid, 
crowned with long cal.-segm.., purple. 

Montane zone 4000-7000 ft.; rather common. Fl. Feb., March, and 
September ; pale pink. 

Endemic. 

2. I. Thwaitesii, Hz. f in Fi. B. Ind. iii. 138 (1880). 

I. acuminata, Thw. Enum. 155 (non Roxb.). C. P. 748. 

FL. B. Ind. iii. 138. Bot. Mag. t. 5197 (1. jucunda). 

A small slender tree, bark thick, exfoliating in oval pieces, 
branchlets compressed ; |. rather large, 44-7 in., lanceolate or 
linear-oblong, rounded or acute or tapering at base, shortly 
acuminate, acute, lat. veins rather prominent, petiole very 
short, stip. with a long cuspidate point, persistent; fl. rather 
large, nearly sessile, crowded, cymes large, very corymbose 
and flat-topped; cal.-segm. about as long as ov., narrowly 
lanceolate, acute; cor.-tube 1-1} in, lobes 4-4 in., spreading; 
fruit 4 in., nearly globose, somewhat didymous, red. 

Var. 3, velutina. 

Twigs and |. on both sides densely and finely velvety. 

Low country in both moist and dry regions; rather common. Var. B 
at Uma-oya. Fl. Jan.-May; cream-coloured. 

Endemic. 

Wood very hard and closed-grained, pale brownish-yellow. The 
flowers vary much in size. The figure in the Botanical Magazine is very 
good for this species, but is unfortunately named /. jucunda. 


348 , Rubiacee. [Zxora. 


3. ZT. parviflora, Vahl, Symb. Bot. ii. 11 (1794). Maha- 
ratambala, S. Karankutti, Painkuray, 7. 

Moon Cat. 10. Thw. Enum.155. C. P. 2575. 

HBS indy 143.) Wight, Ie"t) 711 Bedd) Ely Sylvs tazz2 

A small, much-branched tree, bark thick, reddish-brown, 
exfoliating in irregular pieces, branchlets somewhat com- 
pressed, glabrous ; 1. 3-5 in., oval-oblong or somewhat obo- 
vate, rounded at base, suddenly and shortly acuminate, acute, 
glabrous, coriaceous, dark green and shining above, paler 
beneath, venation pellucid, petiole very short, stout, stip. 
short, cuspidate, subpersistent ; fl, very small and numerous, 
on short ped., cymes trichotomous, stalked, laxly arranged in 
lax cermiing! inflor.; cal.-segm: very shallow, tooth-like ; cor.- 
tube 4—4 in., very slender, Welbes less than half as long as tube, 
oblong, reflexed ; fruit small, 1 in., globose. 

Wave, (6, iene Ik. f. in Fl. B. Ind. l. c. JI. jucunda, var. y, 
bhwe Enumens 5.1 C.) by 2: 

L. larger, 4-7 in., broader, more acuminate, acute at base, 
thinner; branches of inflor. longer ; fl. rather larger ; fruit 3 in 

Low country, especially in the dry region; common. Var. 6 in lower 
montane zone, Adam’s Peak; Medamahanuwara; Hunasgiria. FI. 
April-July; white, var. 8, pinkish, very Sweet- scented, 


Also in India and Burma; var. 8, endemic. 
Wood very hard and heavy, fine-grained, brownish-yellow. 


4. I.jucunda, Zhw. Enum. 155 (1859) (excl. var. y). [PLATE mai 

C. PR. 702, 3424. 

Fl. B. Ind. iii. 144. 

A small tree, 10-12 ft., branchlets stout, thickened at the 
nodes ; 1. large, 4-6 in., oval or lanceolate, acute at both ends, 
glabrous, rather coriaceous, petiole 4-4 in., stip. acuminate; 
fl. on rather long ped., numerous, cymes large, rather lax, 
trichotomous ; cal.-segm. oval, obtuse, shorter than ov.; cor.- 
tube 1} £ tae lobes oblong- oval, less than 4 length of ‘tube ; 
fruit 2-4 in, nearly globose. 

In fees ae region up to 4000 me rather common. FI. most of 
the year; white or pink, sweet-scented. 

Endemic. 

A very beautiful shrub. C. P. 3424 is var. 8 of Thwaites, which differs 
in having narrower, often linear-lanceolate, leaves, cor. with smaller, 
shorter, strongly reflexed lobes and acute cal.-segm., but seems to be 
connected with type by intermediates. The cal. and whole inflorescence 
are often bright coral-red. 


Bo ke coccinea, L. Sp. Pl. 10 (1753). Ratambala, S. 
Vedchi, 7. 

Herm. Mus. 15, 36. Burm. Thes. 125. FI. Zeyl.n. 54 and 55. Z alba, 
L. Sp. Pl. 110. Moon Cat. 10. Thw. Enum. 154. C. P. 190. 

FI. B. Ind. iii. 145. Burm. Thes. t. 57. Wight, Ic. t. 153. Bot. Mag. 
t. 154 (1. grandiflora). 


Pavetta.] Rubiacee. 349 


A shrub with long branches, twigs compressed, thickened 
at the nodes; |. small, 2-3 in., obovate or oval-oblong, 
rounded or even subcordate at base, acute, often cuspidate 
at apex, glabrous and shining, rather rigid, lat. veins some- 
what conspicuous, pellucid, petiole extremely short, stout, 
stip. with a long, rigid bristle, subpersistent ; fi. rather large, 
shortly stalked, cymes lax, trichotomous; cal.-segm. either 
short with toothed margin, or longer and acute, shorter than 
ov.; cor.-tube I-14 in., very slender, lobes oblong-oval, acute 
or obtuse, about 4 as long as tube, spreading; fruit 3 in., 
nearly globose, purple. 

In the low country; very common everywhere up to 2000 ft. FI. all 
the year; brilliant scarlet. 

Also in India and Burma. 

One of our commonest and most conspicuous shrubs. The flowers 
vary much in size, and are occasionally found pure lemon-yellow, also 
sometimes pale cream-coloured with a tinge of red. The latter form has 
a found about Galle and Kalutara, and is probably /. ada, L., and FI. 

eyi. n. 

‘The ae and bark are used as a remedy for diarrhoea, and the 
former also externally to ulcers, &c. Wood very hard and heavy, but of 
no S1Zze. 


35. PAVETTA,* Zinn. 


Large shrubs, stip. often connate, deciduous or persistent, 
fl, numerous, in terminal or axillary cymes; cal.-segm. 4, 
short or long ; cor. with a usually long tube, lobes 4, twisted 
in bud; style very long, at least twice as long as cor.-tube, 
very much exserted, stigma entire; otherwise as in /rora.— 
Sp. 60; 10 in FZ. B. Ind. 


Cymes corymbose, stalked. 
Cal.-segm. shorter than ov. 
L. lanceolate or obovate. 


L. glabrous ; cor.-tube 4 in. 1. P. INDICA. 
L, pubescent beneath; cor.-tube over. 3in,. 2. P. HISPIDULA. 
L. linear or ‘eel glabrous cor.-tube over 
i ae 3. P. ANGUSTIFOLIA. 
Cal.-segm. twice as long as ov. 4. P. GLENIEI. 
Cymes capitulate, sessile 5. P. INVOLUCRATA. 


The first 3 species are very closely allied. 


P. indica, L, SP. Pl. 110(1753). Pawatta,S. Pavaddai, 7. 
‘a Mus. 3. Burm. Thes. 209. Fl. Zeyl. n. 56. Moon Cat. Io. 
Thw Enum. 155. C. P. 1663, 2611, 2546, 3616. 

Fl. B. Ind. iii. 150. Wight, Ic. t. 148. 
A bush or small tree with numerous spreading branches, 
bark smooth, yellowish-white, twigs cylindrical, glabrous; 1. 


* The Sinhalese name scarcely altered. 


350 Rubwacee. [Pavetta. 


3-6 in.; lanceolate, tapering to base, acuminate, subacute, 
glabrous on both sides, dark green and shining above, with 
scattered large, thickened, hard warts, more prominent above 
than beneath, petiole 4-4 in., stip. connate, triangular, acute, 
thin, deciduous ; fl. very numerous, on ped. longer than cal., 
cymes copious, lax, corymbose, terminal, glabrous, often with 
bracts below the branches ; cal.-segm. very small, tooth-like ; 
cor.-tube about $ in., lobes oblong, obtuse, about half length 
of tube; style exserted for fully ?in., very slender, stigma 
slightly clavate; fruit }-3? diam., nearly globose, black, 
polished. 

Var. B, montana, 7iw. in Fl. B. Ind.\.c. C. P. 2512. 

L. narrower, obtuse; fl. larger; cor.-tube 4} in. long. 

Low country; common. Var. 8 extending in the hills to 4oooft., or 
higher. Fl. Dec._May; white. Var. 8, greenish. 

Also in India, Burma, Malaya, S. China, N. Australia. 

This common shrub is very variable in the foliage; C. P. 3616 is a 
pubescent form from Batticaloa, and C. P. 2611 a very narrow-leaved 
form with a few larger flowers on very slender long ped. 

2. P. hispidula, W. & A. Prod. 431 (1834). 


Thw. Enum. 156. C. P. 742, 3924 (°?). 

Fl. B. Ind. iii. 151 (excl. var. 4). 

A large bush, branchlets somewhat compressed, usually 
pubescent; |. large, 4-7 in., variable, usually obovate-oval, 
acute at both ends, shortly acuminate, usually glabrous 
above but more or less pubescent beneath, petiole short, 
stip. rounded, often pubescent; fl. large, numerous on short 
ped., cymes very large, corymbose, terminal, more or less 
pubescent; cal. pubescent, segm. narrowly triangular, acute, 
shorter than ov.; cor.-tube from #~1 in. or more long, slightly 
widened upwards, lobes oblong, obtuse, apiculate, less than 
half length of tube; style exserted for 14-2 in., gradually 
tapering, stigma scarcely at all fusiform; fruit 2-4 in., 
broader than long, somewhat didymous, metallic grey- 
purple, shining. 

Moist region up to 4oooft.; rather rare. Kukul Korale; Hantane; 
Pusselawe; Hunasgiria. Fl. June; greenish-white. 

Also in S. India. 

A variable species. C. P. 3924 was thought by Thwaites to be a 
natural hybrid between this and P. zzdica. The FI. B. Ind. distinguishes 
as var. zeylanica a larger-flowered form, equally common with the 
smaller-flowered one. 

3. BP. angustifolia, 7iw. Enum. 156 (1859). 


Gees2677 
Fl. B. Ind. iil. 151 (P. Azspzdula, var. 4). Bedd. Ic. FI. Ind. Or. t. 99. 


A shrub, with smooth, white bark, branches cylindrical, 
often nodular with thickened, closely placed nodes, twigs 


Pavetta.| Rubiacee. 351 


compressed, glabrous; |. rather crowded, 4—7 in., linear or 
lanceolate-linear, much tapering to either end, subacute, 
often somewhat undulate, glabrous, deep green and polished 
above, paler beneath, with similar warts to P. zudica, stip. 
connate into a tube, with a terminal bristle, usually splitting 
down one side, subpersistent, fl. on short ped., cymes corym- 
bose, rather lax, axillary or terminal; cal. glabrous, segm. 
distant, linear-lanceolate, acute, rather shorter than ov.; cor.- 
tube 2-i in., lobes linear-oblong, obtuse, more than half as 
long as tube; anth. very long, linear; style rather stout, 
tapering to very slightly fusiform stigma, exserted for 14-2 
in.; fruit 4 in., capped with cal.-limb, sub-didymous, broader 
than long, greenish-purple. 

Moist low country by rivers, rather rare. About Ratnapura; by the 
‘Gindura near Wattagalle. Fl. March, April; cream-coloured or pale 
lemon-yellow. 


Endemic. 
Considered a variety of P. hésfzdula in FI. B. Ind. 


- 4- = Gleniei, 7hw. iz Fl. B. Ind. iii. 152 (1880). Wetpavad- 
dai, 7. 
P. tomentosa, Thw. Enum. 156 (non Sm.). C. P. 2815. 

Fl. B. Ind. iii. 152. 

A large shrub, bark pale cinnamon-brown, smooth, twigs 
compressed, pubescent; |. 2-4 in., oblong-oval or oblong- 
lanceolate, acute at base and apex, pubescent on both sides, 
densely so beneath, with scattered warts, venation pellucid, 
petiole about $ in., stip. slightly connate, triangular, acute; f1. 
small, in lax, trichotomous, hairy, axillary cymes shorter than 
l.; cal.-segm. linear, fully twice as long as ov., hairy, persistent ; 
cor.-tube about } in., lobes narrowly oblong, obtuse, more than 
half as long as tube; style exserted for about }in., stigma 
slightly clavate, pubescent; fruit small, }in., nearly globose, 
crowned with long erect cal.-segm., hairy. 

Dry region; common. Puttalam; Trincomalie; Batticaloa; N. Prov.; 
Anuradhapura; Dambulla; Doluwa Kande, near Kurunegala. FI]. Nov.- 
March; white. 

Endemic. 

This common dry-country plant is very constant. I have not found 
the glabrous form recorded in FI. B. Ind. 


5. P.involucrata, 7hw. num. 156 (1859). 

C. P. 428. 

Fl. B. Ind. iii. 152. Bedd. Ic. Pl. Ind. Or. t. 100, and FI. Sylv. Anal. 
t. 29. 

A shrub, 10-12 ft. high, branchlets thickened at nodes with 
prominent L-scars, twigs much compressed, glabrous; |. 3-44 


352 Rubiacee. [ Coffea. 


in., often crowded, obovate-oval, tapering to base, subacute, 
shortly acuminate, glabrous on both sides, with scattered 
warts, petiole 4 in. stip. large, connate, acuminate, tube 
splitting down one side, deciduous; fl. sessile, crowded in 
dense, sessile heads at ends of short axillary branches, usually 
bearing a single pair of 1. immediately beneath the head, bracts 
very large, ovate, membranous, forming an involucre; cal.- 
segm. linear, acute, about as long as ov.; cor.-tube short, 2 in.,. 
broad, lobes lanceolate, acuminate, erect; style thick, tapering, 
exserted for $—3in.; fruit 4in., globose, with a crown of cal.- 
segm. 

Forests of the upper montane zone; rather rare. Adam’s Peak; 
Dimbula; N. Eliya. Fl. March, April; greenish-white. 

Endemic. 

Very unlike the other species in the inflorescence. Becomes black 
when dried. 


36. COPFEA, Linz. 


Small shrubs, stip. small; fl. solitary or in threes, axillary 
(often apparently terminal), cal.-limb many-toothed or none 3. 
cor.-tube long, lobes 5, spreading, twisted in bud; stam. in- 
serted in throat of cor., fil. o, anth. linear; ov. 2-celled, with 
a single ovule in each cell attached by centre to septum, 
stigmas 2, linear; fruit containing 2 cartilaginous pyrenes ; 
seeds with a deep ventral groove, endosperm horny.—Sp. 20; 
6 in FZ. B. Ind. 

Stip. persistent, spinescent; cal.-limb many- 


toothed. j : : 5 : . I. C. WIGHTIANA. 
Stip. caducous; cal.-limb.o . : 5 . 2. C. TRAVANCORENSIS. 


1. ©. Wightiana, Wal/. Cat. x. 6246 (1828). Kaddumallikai, 7. 

Thw. Enum. 154. C. P. 1654. 

Fl. B. Ind. ii. 154. Wight, Ic. t. 1598 (not good). 

A small bush, with white bark and many rigid, stout, 
divaricate branches (often suppressed and very short), young 
shoots puberulous; |. small, 4-14 in., nearly sessile, oval, 
tapering to base, obtuse, glabrous, stip. short, rigid, sharply 
pointed, persistent and becoming small curved prickles, 
especially on the short lat. branches; fl. solitary, sessile, on 
the short lat. branches ; cal.-limb with many teeth; cor.-tube 
about $in., lobes oblong-oval, obtuse, rather more than half 
as long as tube; fruit not seen, “34in., much broader than long, 
didymous.” 


Dry region; rather rare. Jaffna; Mihintale; Uma-oya; Atakalan 
Korale. Fl. Aug., Sept.; white, very sweet-scented. 


Morinda.) Rubiacee. 253 


Also in S. India. 
A resinous gum envelops the calyx. This seems to me abundantly 
distinct from the next. 


2. €. travancorensis, WV. & A. Prod. 435 (1834). [PLATE LIII.] 
Coffea trifiora, Moon Cat. 15 (non Forst.). Thw. Enum. 154. C. P. 2458. 
FI. B. Ind. iii. 154. Bot. Mag. t. 6746. 

A small shrub, branchlets stiff, slender, thickened at nodes, 
twigs flattened, slightly pubescent; |. small, 14-3 in., oval or 
lanceolate, acute at base, acuminate, acute or obtuse at apex, 
quite glabrous, rather thin, petiole very short, stip. small, 
long-triangular, acute, soon falling; fl. solitary or in threes, 
very shortly stalked, each with two minute, linear bracts at 
base ; cal. very small, puberulous, limb quite absent; cor.-tube 
2 in., glabrous within, lobes ovate-oblong, acute, rather shorter 
than tube; ov. surmounted by thick disk surrounding style, 
stigmas large, erect; fruit 3 in. diam., distinctly didymous, 
broader than long, slightly puberulous when young, black. 

Moist and intermediate region to 3000 ft.; rather rare. Kalutara 
(Moon); Kurunegala (Gardner); Negombo; Deltota; Doluwe Kande. 
Fl. April-June; white, very sweet-scented. 

Also in Travancore. 

Moon’s S. name for this is ‘ Gas-pitcha,’ and the flowers much resemble 
those of a jasmine. 


C. arabica, L., the common Arabian coffee, introduced to Ceylon 
from Java by the Dutch about 1690, and largely cultivated, is often found 
about the sites of former native gardens or plantations, but has shown no 
tendency to become naturalised. 


37. MORINDA,* Zinn. 

Trees or climbing shrubs, stip. various; fl. in dense stalked 
heads, the peduncle often taking the place of one |. of the pair 
and thus leaf-opposed but within the stip., or terminal; cal.- 
tube more or less fused with adjacent ones in the head, limb 
short or 0, segm. usually 0; cor. tubular, lobes 4 or 5, valvate, 
hairy at mouth; stam. 4 or 5, anth. oblong; ov. 2 or spuriously 
4-celled with 2 or 4 ovules, stigmas 2, linear; fruit in each 
head concrete into a fleshy globose mass, pyrenes 3 or 4 to 
each berry, 1-seeded.—Sp. 40; 7 in /Z. B. Lund. 


Trees; cor.-lobes 5. 


L. under 6 in., dull , ; ; ; : . I. M. TINCTORIA. 
L. over 6Gin., shining . { , ; 5 . 2. M. CITRIFOLIA. 
Climbing shrub: cor.-lobes 4. i ; ; . 3. M. UMBELLATA. 


F]. conspicuously dimorphic. 


* Morinda, a name given by Vaillant; said to be a contraction of Morus 
indica, and doubtless alluding to the compound fruit. 
PART IIL. AA 


354 Rubtacee. (Morinda. 


1. M. tinectoria, Roxd. Hort. Beng. 15 (1814). Ahu, S. Man- 
chavanna, 7. 

M. exserta, Roxb., Thw. Enum. 145. C. P. 1668. 

Fl. B. Ind. iii. 156. Bedd. FI. Sylv. t. 220 (Z. cétréfolia). 


A usually small tree, bark thick, spongy, with deep longi- 
tudinal furrows, branchlets thickened at nodes, l.-scars pro- 
minent, twigs compressed, glabrous; |. 3-5 in., lanceolate, 
tapering to base, acuminate, acute, glabrous save for tufts of 
hair in axils of lat. veins beneath, dull green not shining, 
petiole very short, stip. broadly triangular, acute, deciduous ; 
fl. rather large, heads globose, peduncles long, solitary or 
rarely 2-3 together ; cal.-tube about 4 in., lobes 5, narrowly 
oblong, more than half as long as tube; head of fruit about 
#in. diam., globose, fleshy, pyrenes usually 4, oblong. 

Dry region; rather common. Jaffna; Trincomalie; Anuradhapura; 
Badulla District (Thwaites). Fl. Sept., Oct.; white, very sweet-scented. 

Also in India and Malaya. 

Wood moderately hard and heavy, brownish-orange, durable. The 
reddish-brown bark of the root affords a red dye, much used for dyeing 
cloth, but the tree is not cultivated in Ceylon. The leaves are a remedy 
for diarrhoea, and an external application to wounds. The plant dries a 
dark metallic brown. 


2. M. citrifolia, 2. SP. P/. 176 (1753). Ahu, S. 

Fl. Zeyl. n. 82. Moon Cat.15. J. dvacteata, Roxb., Thw. Enum. 
144 1G Pa765: 

Fl. B. Ind. iii. 155. Wight, Ill. t. 126 (JZ. dvacteata). Rheede, Hort. 
Malab. i. t. 52. 

A small tree, with yellowish-white bark, branchlets stout, 
marked with l|.-scars, twigs glabrous; |. closely placed, large, 
6-8 in., oval, attenuate at base, shortly acuminate, acute, 
glabrous, |. shining, with glandular pits in axils of lat. veins 
beneath, rather thick, bright apple-green, veins strongly 
marked, prominent beneath, pellucid, petiole 4 in., very stout, 
stip. short, broad, obtuse, persistent; fl. as in the last, but 
smaller; cor.-lobes 5; head of fruit ovoid, over 1 in. long, very 
fleshy, white, some of the enlarged cal. of the lowest fruits 
developing large and leafy bract-like segm.; pyrenes ovoid, 
compressed, winged on one edge, convex on one surface, con- 
cave on the other. 

Near the coast in the moist region; rather rare, often cultivated. 
Colombo; Galle; Tangalle. Fl. October; white. 

Also in Andaman Is., Malaya, Pacific Is., N. Australia; doubtfully 
wild in Peninsular India. 

There are only drawings—no specimens—of this in Hermann’s col- 
lection. The bract-like leafy appendages to some of the lower calyces 
are persistent in the head of fruit, and very characteristic. 

This is a Malayan and Pacific maritime plant, and, like several others 


Prismatomeris.] Rubiacee. Bo 


of that group, native here only in a few spots on our south-western 
coast, but it is much cultivated in-gardens. It is, I believe, quite distinct 
from the dry country 4. “uctorta, of which it has been considered 
as a cultivated variety. 

The root-bark affords a red dye as in MZ. “enctoria. 

3. M. umbellata, Z. Sf. P/. 176 (1753). Miri-wel, Maha- 
kiri-wel, S. 

Fl. Zeyl. n. 81. MZ. scandens, Roxb., Moon Cat. 15. Thw. Enum. 
145. C. P. 1669. i 

Fl. B. Ind. iii. 157. Rheede, Hort. Malab. vii. t. 27. 

A scrambling shrub, climbing by very long, slender, sar- 
mentose, divaricate branches, bark grey, striate, internodes 
very long, young parts puberulous; |. rather small, 2-4 in., 
usually oblong-lanceolate, but sometimes broadly oval, tapering 
or acute at base, shortly acuminate, acute, glabrous, thin, dark 
green, venation finely reticulate, pellucid, petiole 4 in., often 
twisted, stip. connate, sheathing, membranous, subpersistent ; 
fi. few together, heads small, less than } in. diam., globose, 
stalked, peduncles 4-10 together in terminal umbels; cal.- 
limb short, perfectly truncate; cor.-tube very short, lobes 4, 
oblong-oval, much longer than tube; head of fruit small, 
about 4 in. diam., lobulated, smooth, scarlet. 


Moist region extending up to 4000 ft. or higher; common. 

Fl. Feb.; white or greenish. 

Throughout the Tropics of Asia. 

At the higher elevations this often forms a small bush with no ten- 
dency to climb, and with much broader and thicker leaves. 


38. PRISMATOMERITIS, 7iw. 


Shrub, stip. shortly connate, often split; fl. few, terminal, 
unisexual; cal.-limb cup-shaped, truncate with 5 tooth-like 
segm.; cor.-tube glabrous within, lobes 5, very thick and 
prismatic on section, valvate; stam. 5, inserted in tube, small; 
ov. 2-celled, with a single ovule in each cell, stigmas 2, 
spreading; fruit a berry completely filled by 2 large seeds; 
seed rounded on back, deeply excavated on ventral surface, 
endosperm horny.—Sp. 3; 2 in Fi. B. Ind. 

P. albidifiora, 7/iw. in Kew Journ. Bot. viii. 269 (1856). 

Thw. Enum. 154. C. P. 728. 

Fl. B. Ind. iii. 159. Kew Journ. Bot. viii. t. 7, f. A. Bedd. Ic. Pl. Or. 
t. 93, and FI. Sylv. Anal. Gen. t. 29, f. 4 (not good). 

A shrub 10-12 ft., much divaricately branched, bark grey, 
branchlets sharply quadrangular compressed, thickened at 
nodes, yellow, polished; 1. 24—5 in., oval or lanceolate-oval, 
tapering to base, shortly acuminate, subacute, glabrous, 
venation conspicuous, reticulate, petiole very short, stip. 


356 Rubwacee. [Psychotria. 


forming a short tube; fl. moderate in size, 1-4, usually in 
pairs at end of twigs, ped. 1-14in., slender, divaricate; cal. 
glabrous; cor.-lobes oblong, acute, excavated at base, about 
as long as tube; fil. very short; fruit 2 in., nearly globose, 
broader than long and very slightly didymous, purple. 


Var. 6, Fergusonii, 77vim. P. Fergusoniz, Thw.* ex. Bedd. FI. 
Sylv. Fl. Man. 134, 10. FI. B. Ind. iii. 159. 

Pedicels 5-10, in terminal usually stalked umbels, corolla- 
segments less fleshy. 

Moist region, rare, extending up to 4oooft. Between Negombo and 
Kurunegale; Ambagamuwa; Dolosbagie ; Maskeliya. Var. 8, Colombo 
(Ferguson). Fl. March, April; yellowish-white, sweet-scented. 

Also in Khasia, Burma, Malaya; not in Peninsular India. 

Var. 6 is maintained as a species in FI. B. Ind. on the ground of 
Beddome’s statement that the corolla-lobes are imbricate. This, is, 
however, not the case; the zestivation is strictly valvate, the prismatic 
lobes forming an angular bud. Indeed, their extremely fleshy nature 
necessitates a valvate or reduplicate-valvate zestivation. The stipules. 
exude a resinous gum. I have never seen 4-merous flowers. 


39. PS¥CHOTRIA,* Z. (part). 


Shrubs, rarely climbing, branchlets usually compressed 
and glabrous, stip. various, often connate, fl. small, in terminal 
cymes usually without bracts; cal.-limb long or short, segm. 
broad or narrow, sometimes caducous; cor.-tube short, straight, 
hairy or glabrous in throat, lobes 5 (rarely 4), valvate; stam. 
inserted in throat of cor.; ov. 2-celled, with one erect ovule in 
each cell, stigmas 2; fruit small, berry-like, with 2 plano- 
convex, I-seeded pyrenes; endosperm plane or ruminate.— 
Sp 50s 52 10 Bela: 

Nine of our 13 species are endemic. 


Pyrenes without furrows on either surface ; endosperm ruminate(Gvw7u7z/eq). 
Branches of cyme whorled. 
Leaves glabrous. 
Lat. veins invisible : . 3 . 1. P. STENOBEWEWAS 
Lat. veins conspicuous. 
Berry in. long. 


Cal.-limb long 2. P. GLANDULIFERA. 
Cal.-limb very short 3. P. GARDNERI. 
Berry near 3 in. long . 4. P. THWAITESII. 
Leaves hairy beneath 5. P. WIGHTIANA. 
Branches of cyme in pairs 6. P. ELONGATA. 


* Thwaites informed me that he had no recollection of bestowing 
this as a specific name, and that he never considered the plane specifically 
distinct from P. albidiflora. 

+ Psychotria altered from Psychotrophum, the name given to a W. 
Indian plant by P. Browne, now Psychotria Browne, Spreng. 


Psychotria.] Rubiacee. 357 


Pyrenes with 4 furrows on back and 2 shallow 

ones on face; endosperm not ruminate SAR MENT OSA: 
Pyrenes with 2 broad furrows on back, none on 

face ; endosperm not ruminate. 


Fl. sessile. 
L. over 6in., stip. ciliate . : : Sao ee VLOONET. 
L. under 4 in., stip. not ciliate . ; Oe SORDIDAS 
Fl. pedicellate. 
Petiole over Tin. . : : ; . Io. P. LONGEPETIOLATA. 
Petiole under 3 in. 
Lat. veins numerous. 
Cal.-segm. linear-lanceolate  . . Ii. P. PLURIVENIA. 
Cal.-limb nearly truncate . : oe 2s PETRI ES: 
Lat. veins few; cal.-segm. triangular . 13. P. BISULCATA. 


I. P. stenophylia, 7. f~ zz Fl. B. Ind. iii. 161 (1880). 

Grumilea stenophylla, Thw. Enum. 147. C. P. 2609. 

Fl. B. Ind. iii. 161. 

A much dichotomously branched bush, branchlets thickened 
at the nodes, smooth and shining; |. 3-5 in., linear or linear- 
lanceolate, tapering at both ends, acute, quite glabrous, midrib 
broad, conspicuous, lat. veins scarcely visible, rather coriaceous, 
margins somewhat recurved, pale green, petiole very short or 
none with a tuft of brown hair in axil, stip. large, connate, 
ovate, obtuse, glabrous, caducous; fl. sessile, surrounded with 
ferrugineous hair, in small globose, stalked clusters, usually in 
a whorl of 4, and a terminal one forming a small pedunculate, 
pyramidal cyme (occasionally axillary); cal.-segm. very shallow, 
cor.-tube very short, pilose in throat, lobes 4, ovate, acute; 
berry 4 in., broadly ovoid, shining, bright purple; pyrenes 
without furrows, endosperm ruminate. 


Moist low country to 2000 ft.; rare. Nillowe Kande; Ratnapura; 
Reigam Korale; Ambagamuwa. FI. Feb., March; green. 
Endemic. 


2. P. glandulifera, 7iw. ex Hk. f. Fl. B. Ind. iii. 161 (1880). 

rr. 3011. 

Fl. B. Ind. iii. 161. 

Shrub, branchlets glabrous; 1. 4-7 in., oblong-lanceolate, 
acute at base, suddenly acuminate, acute, glabrous, pale be- 
neath, with the lat. veins conspicuous, and with large and 
deep glandular pits in their axils, petiole 4-2 in., stip. as in 
the last; fl. few, nearly sessile, in short, sessile, trichotomous 
cymes; cal.-limb long, segm. very shallow, acute; berry } in., 
ovoid, crowned with large cal.-limb. 

Only found near Adam’s Peak, where the C. P. specimens (in bud and 
fruit) were collected, March, 1866. 

Endemic. 

I have seen only the C. P. specimens, and unfortunately the fruit is 
not sufficiently ripe to show the character of the seed. 


35 8 Rubiacee. [Psychotria. 


3. BP. Gardneri, He. f. in Fl. B. Ind. iii. 161 (1880). 

Grumilea Gardneri, Thw. Enum. 147. C. P. 1718. 

FI. B. Ind. ii. 161. 

A shrub, twigs glabrous; |. large, 6-9 in., obovate-oblong,. 
very much tapering to base, very abruptly narrowed into a 
long tail-like end, obtuse at apex, margins narrowly recurved, 
glabrous, lat. veins numerous, fine, distinct, petiole obscure, 
stip. ovate, obtuse, quickly falling; fl. sessile, in small heads, 
cymes large, paniculate with whorled branches, bracts con- 
spicuous, short, broad, rounded ; cal.-limb very short, segm. 
obscure; berry small, 4+ in., ovoid, black, pyrenes without 
furrows, endosperm ruminate. 

Lower montane zone; rare. Ramboda (Gardner); about Balangoda, 


abundant. FI. white. 
Endemic. 


4. B. Thwaitesii, Hz. ~ F/. B. Ind. iii. 162 (1880). 

Grumilea nigra, Gaertn.(?), Arn. Pug. 25. G. nudiflora, Thw. Enum. 
147 (non W. and A.).  C. P. 18. 

FI. B. Ind. iii. 162. 

A shrub, branchlets stout, glabrous, |. rather large, 4-9 in., 
oblong-lanceolate, often somewhat obovate, acute at base, 
acuminate, acute at apex, glabrous, midrib very strong, lat. 
veins distinct, petiole }-4 in., stip. broadly oval, obtuse; fl. 
nearly sessile, with brown hair at their base, in small round 
heads, cymes small, long-stalked, pyramidal, with whorled 
branches; cal.-limb deciduous, segm. obscure, broad; cor.- 
tube very short; berry rather large, 2-2 in., broadly ovoid, 
without cal.-limb, pyrenes without furrows, endosperm much 
ruminated. 

Var. 6 coronata, ZZ. f. /.c. 


L. smaller and broader ; cal.-segm. ovate, acute, persistent 
and crowning the fruit. 


Moist region, from about 1000 to 6000 ft.; very common in damp 
forests. Var. 8 at the higher elevations. FI. Jan._March; greenish. 

Also in S. India. 

This is almost certainly Grumzlea nigra, Gaertn. Fruct. i. 138 and t. 
28, and the type of that genus (now combined with Psychotria). He 
gives ‘Kogdala’ as the Sinhalese name. 

Var. 6 is very close to P. congesta, W. and A., of the Nilgiri Hills. 


5. BP. Wightiana, Zz. f F7. B. Ind. ii. 167 (1880). 

Lastanthus ? dubius, Wight in Calc. Journ. Nat. Hist. vi. 516. Gru- 
milea Wighttana, Thw. Enum. 148. C. P. 586. 

Fl. B. Ind. ii. 167. 

A small shrub, branchlets stout, roughly hairy ; |. large, 
5-9 in., oval or lanceolate, acute at base, acuminate, acute, 


Psychotria.] Rubiacee. 359 


glabrous above, softly hairy beneath, petiole }—{ in., very 
hairy, stip. large, connate, oblong, obtuse, pilose; fl. sessile in 
small round heads, cymes large, peduncled or sessile, lax, 
very hairy, branches long, whorled, divaricate ; cal.-limb large, 
erect, very hairy, segm. short, obtuse, glabrous within; berry 
2 in. broadly ovoid, crowned with cal.-limb, very hairy, 


8 
pyrenes without furrows, endosperm ruminate. 


Var. 6 affinis, Wk. 7. /.c. Grumilea affinis, Thw.\.c. C. P. 2706. 


L. smaller, linear-lanceolate, less hairy beneath; cal.- 
segm. acute. 

Moist low country by streams; rare. Ambagamuwa; Ratnapura; 
var. 8, Hewesse; Palabadala; Kitulgala. Fl. Feb—April; green. 

Endemic. 

Wight’s specimens, which were without fruit and hence misled him 
as to the genus, were communicated to him by Col. Thacker (and also 
collected by himself) in April, 1836. 


6. P. elongata, Hk. f. F/. B. Ind. iii. 163 (1880). 

Grumilea elongata, Wight, Thw. Enum. 147. C. P. 1719. 

Fl. B. Ind. iii. 163. Wight, Ic. t. 1036. 

A shrub, twigs very smooth; |. 3-44 in., obovate-oval, 
obtuse at base, suddenly and shortly acuminate, acute, 
glabrous, lat. veins numerous, 12-16 on each side, parallel, 
prominent, usually with deep glandular pits in the axils 
beneath, petiole }in., stout, stip. short, pubescent, partially 
persistent ; fl. nearly sessile, small, in dense clusters, cymes 
pedunculate, narrow, branches short, opposite, peduncle with 
2 large sheathing stipular bracts at base; cal.-segm. tri- 
angular, acute, caducous; cor.-lobes rather shorter than wide 
tube; berry 4 in., ovoid or slightly pyriform, without a cap 
of cal.-limb, faintly vertically 1o-ribbed, pyrene without fur 
rows, endosperm ruminate. 


Montane zone to 7000 ft.; common. FI. Feb.—April; white. 
Also in mountains of S. India. 


7. P. sarmentosa, 4/. Lijdr. 964 (1826). Wal-gonika, S. 

P. scandens, Moon Cat. 14. Thw. Enum. 148. C. P. 1712. 

Fl. B. Ind. iii. 165. Wight, Ic. t. 1038. 

A shrub, climbing by adventitious rootlets, stems sarmen- 
tose, cylindrical, smooth, with long internodes; |. rather 
small, 24-3 in., oval or oval-lanceolate, rounded or acute at 
base, shortly acuminate, subacute, glabrous, lat. veins few, 
faintly marked, petiole } in., stip. connate and sheathing, 
acute; fl. small, shortly stalked, cymes lax, corymbose, 
branches opposite, divaricate; cal. pilose, segm. broadly 
triangular, acute ; cor.-tube with a ring of dense yellow hairs 


2M 


360 Rubiacee. [Psychotria. 


in mouth, lobes oval-oblong, obtuse; berry nearly 4 in., 
globular, without a cal.-crown, smooth, white, pyrenes with 
4 furrows and 5 ribs down the back and 2 shallow furrows 
on ventral surface, endosperm not ruminate. 

Low country up to 3000 ft. or more; common. FI. Jan—March; 
green. ' 
~ “Also in India and Malaya. 

The fruit is a white globular berry, not capped with calyx teeth, and 
when fresh is quite without ribs or furrows, but, when dried and the pulp 
shrivelled up, it appears strongly 1o-ribbed, as in Wight’s figure above 
quoted. 


8. P. Moonii, 72. 7, F/. B. (nd. ii. 170 (1880). 

Grumilea Mooniz, Thw. Enum. 148. C. P. 585. 

Fl. B. Ind. iil. 170. 

A large shrub, branchlets glabrous; 1. 6-8 in., oblong-oval 
or slightly obovate, tapering to base, abruptly acuminate, 
acute, glabrous on both sides, thin, lat.-veins numerous, 
curved, often with glandular pits in axils beneath, petiole 4-1 
in., stip. large, oblong, connate, obtuse, ciliate; fl. sessile, in 
dense heads, with numerous large rotundate, strongly ciliate, 
persistent bracts, cymes glabrous, with long opp. branches, 
the lowest pair strongly deflexed; cal.-limb large, tubular- 
cup-shaped, segm. obtuse, strongly ciliate; berry 3-4 in., 
oblong, crowned with large cal.-limb, smooth, pyrenes flat on 
ventral surface, with 2 broad shallow furrows on back, en- 
dosperm plane. 

Moist country below tooo ft.; rare. Reigam Korale (Thwaites) ; 


Pahingale, Pasdun Korale. Fl. June, September; white. 
Endemic. 


9g. P. sordida, 7iw. Enum. 149 (1859). 

C. P. 333 (355, 1716). 

Fl. B. Ind. 1. 170. 

A. much-branched bush, branchlets slender, glabrous or 
minutely pubescent; |. rather small, 2—4 in., lanceolate-oblong, 
tapering to base, long-acuminate, very acute, glabrous, rather 
thick, lat. veins horizontal, usually inconspicuous, petiole 
4-2 in., stip. broad, connate, acute or obtuse; fl. sessile in 
clusters of 3, cyme sessile, branches usually 3, short, slender 
divaricate or deflexed, no bracts; cal.-limb very short or none, 
segm. small, linear, acute; cor.-tube inflated, glabrous within; 
disk very large and prominent; berry 4—3in., oblong, capped 
with small cal., pyrenes with 2 deep and wide furrows on back, 
endosperm plane. 


Upper montane zone; rather common. FI. Feb.-April; greenish- 
white. 


Psychotria.| Rubwacee. 361 


Endemic. 

Varies a good deal in shape of leaves; at the higher elevations they 
are often smaller, broadly oval, less acuminate, coriaceous with recurved 
margins. 


10. P. longepetiolata, 7iw. Enum. 149 (1859). 

© B2707. 

FL. B. Ind. iii. 170. 

A shrub, branchlets glabrous; |. 33-6in., oval or oblong- 
oval, rounded and often unequal or acute at base, abruptly 
acuminate, acute, glabrous and shining, petiole very long, 
14-2 in., stip. oblong, acute, deciduous; fl. on short ped., 
cymes pedunculate, pyramidal, branches whorled ; cal.-limb 
short, spreading, segm. broad, obtuse; cor.-tube very short, 
with a ring of hair in throat; disk large; berry 4 in., oblong- 
ovoid, pyrenes very widely and deeply 2-grooved on back. 

Lower montane zone, 3000-4000 ft.; very rare. Diyabetane, Sabara- 
gamuwa; Pitigala-Kande, near Balangoda. Fl. March and Sept.; green. 


Endemic. 
The leaves dry yellow. 


I1. P. plurivenia, 7iw. Enum. 149 (1859). 

= 2052. 

Fl. B. Ind. iu. 170. 

A shrub, with slender divaricate glabrous branchlets; |.small, 
2-4 in., oblong, acute at both ends, glabrous, lat. veins numer- 
ous, fine, nearly horizontal, united by an intramarginal one, 
petiole short, stip. tapering into long filiform point, deciduous; 
fi. on short, slender ped., usually in pairs, cymes very lax, with 
long filiform spreading branches, few-fld.; cal.-limb short, 
segm. rather long, linear-lanceolate, acute; cor.-tube hairy at 
mouth, lobes long, oblong, acute; berry nearly 4 in., oblong, 
crowned with erect cal.-segm., pyrenes broadly 2-furrowed on 
back. 

Lower montane zone, 3000-4000 ft.; very rare. I have only seen the 


C. P. specimens collected in Hewahette Dist. in 1852. FI. Oct.; white. 
Endemic. 


12. PB. filipes, Hz. f. F7. B. Ind. iii. 170 (1880). 

P. bisulcata, var. 8B, Thw. Enum. 148. C. P. 1720. 

Fl. B. Ind. iii. 170. 

A shrub, branchlets glabrous, |. 24-64 in., linear-lanceolate, 
tapering to base, acuminate, acute, glabrous, not thick, lat. 
veins numerous, 10-20 or more, petiole very short, stip. broad, 
deciduous; fl. on slender ped., cymes small, lax, with slender 
irregular branches; cal.-limb cup-shaped, nearly truncate ; 
berry 2-%in., oblong, truncate, pyrenes with 2 deep furrows 
on back. 


362 Rubwacee. [Chasalia. 


Moist region from 2000-4000 ft.; rare. Hantane (Gardner); Sabara- 
gamuwa (Thwaites). Fl. March. 
Endemic. 


13. P. bisulcata, W. and A. Prod. 434 (1834). [PLATE LIV.] 

e ss ae Moon Cat. 14 (non L.*). Thw. Enum. 148 (excl. var. 6). 
o ky THO: 

Fl. B. Ind. iii. 170. Wight, Ic. t. 1039. 

A shrub, with rather stout cylindrical glabrous and shining 
branches; |. 13-4 in., lanceolate or obovate-lanceolate or oval, 
tapering to base, shortly acuminate, acute or subacute, glabrous, 
paler and rather silvery beneath, rather thick, lat. veins 7-10, 
depressed above, rather prominent beneath, often with very 
deep pits in their axils, which appear as warts on upper surface, 
petiole 4-4 in., stip. $in., oblong, acute, membranous, soon 
falling; fl. on short ped., rather large (for genus), cymes 
usually shorter than 1., lax with divaricate opp. branches; cal.-. 
limb cup-shaped, segm. short, triangular, acute ; berry 4 in.,. 
ovoid-oblong, crowned with large cal.-limb, smooth, pyrenes 
with 2 deep grooves on back. 

Upper montane zone; very common. Fl. March—May; green. 

Also in the Nilgiri Mountains. 


P. vaginans, DC. Prod. iv. 520 (Ophioxylon arboreum, Koenig in Hb. 
Royen), from Ceylon, is Gaertnuera Kenigiz, Wight (Loganiacez). 


40. CHASALIA, Comm. 


A small shrub, stip. usually bifid. subpersistent; fl. rather 
small, dimorphic, in terminal cymes; cal.-limb very short, 
segm. 5 (or 6), short; cor.-tube slightly dilated above, some- 
what curved, glabrous within or with a few hairs at insertion 
of stam., lobes 5, much shorter than tube, valvate; anth. long, 
sagittate at base; ov. 2-celled, with one ascending ovule in 
each cell, styles long, spreading ; disk very large and pro- 
minent ; fruit a berry, pyrenes 2, very thin, very concave on 
ventral surface, cup-shaped (crescent-shaped on section), en- 
dosperm not ruminate.—Sp. 10; 2 in FZ. B. Ind. 


CG. curvifiora, 7iw. Enum. 150 (1859). 

(Ce 12, uit, 

Fl. B. Ind. iii. 176. Wight, Ill. t. 127 (Psychotria ambigua). 

A slender shrub, twigs compressed, glabrous; |. 3-9 in., 
from oval to linear-lanceolate (usually 3-5 in. and lanceolate), 


* Linnzeus’s P. aséatica, the basis of the genus, is really a W. Indian 
plant, now P. Brownez, Spr. 


Geophila.| Rubiacee. 363 


tapering to base, acuminate, acute or obtuse, glabrous, thin, 
lat. veins curved, anastomosing within the margin, rather pro- 
minent beneath, petiole obscure, 4-1 in., stip. short ; fl. sessile, 
usually in threes, cymes small, stalked, terminal, trichotomous, 
peduncle 1 in., glabrous, pink or white, bracts linear, small ; 
cal. glabrous, segm. broad, acute or PASH) cor. thick, tube 
rather slender, lobes oblong, obtuse; berry 4in., depressed- 
globose, tipped with withered cal. -segm., pulpy, inky-purple, 
pyrenes dorsally compressed. 

Moist region up to 5000 ft., or higher; rather common, especially 
above 3000 ft. Fl. Jan—April and July; pinkish-white, calyx purple. 

Also in India, Burma, Malaya. 

C. lurida, Miq. F1. Ind. Bat. ii. 282 (1856), is an earlier name. The 
petioles and midrib beneath are usually stained with dark purple. Var. 


longifolia, Hk. f.. has longer more oblong |. with the lat. veins more 
numerous and horizontal; we have it from Reigam Korale. 


41. GEOPHILA, Von. 


A small creeping herb, stip. broad, subpersistent ; fl. few, 
moderate-sized, terminal; cal.-segm. 5, long, linear-lanceolate; 
cor.-tube long, lobes 5, valvate; stam. inserted low down in 
tube, anth. sagittate at base; ov. 2-celled, with 1 erect basal 
ovule in each cell, stigma bilobed; fruit a berry, pyrenes 2, 
plano-convex, not grooved, endosperm plane.—Sp. 8-10; 1 in 


fl. B. Ind. 


G. reniformis, LD. Don, Prod. Fl. Nepal. 136 (1825). Agu= 
karni, S. 
Psychotria herbacea, L., Moon Cat. 14. Thw. Enum. 150. C. P. 1705. 

Fl. B. Ind. 111.178. Wight, Ic. t. 54. 

Small, herbaceous, stems prostrate, rooting at the nodes, 
filiform, puberulous, with short erect branches; |. small, 4-1 
in., ovate-reniform, obtuse, slightly pubescent or glabrous, 
paler beneath, petiole erect, 4-1 in., pubescent, stip. broadly 
ovate, obtuse; fl. on short ped., 1-3 together, with 2 linear 
bracts beneath each, terminal; cal.-segm. much longer than 
tube, acuminate, ciliate; cor. slightly pubescent outside, tube 
dilated upwards, lobes oblong, acute, shorter than tube; berry 
2in., globose, crowned with large cal.-limb, smooth, pulpy, 
red, pyrenes small, flat and smooth on ventral, convex, rough 
and bluntly keeled on dorsal surface. 

Moist low country to 3000 ft.; rather common (according to Thwaites), 
but I have met with it but rarely. FJ. May and June. 

In most parts of Tropics of Asia, Africa, and America. 

Moon’s locality is Kalutara; he also gives as S. name ‘ Koturu-bedda.’ 


364 Rubiacee. [Lastanthus. 


42. LASIANTHUS, /ack. 


Shrubs, |. usually much acuminate, with arching lat.-veins 
connected by transverse veinlets, stip. more or less triangular, 
fl. small, sessile or nearly so, in axillary, usually sessile heads 
or cymes; cal.-segm. 4 (or 5), persistent; cor.-tube funnel- 
shaped, villous in throat, lobes 4 (or 5), hairy; stam. inserted 
in cor.-throat, fil. short, anth. oblong; ov. 4- (or 5-) celled, 
crowned with large fleshy disk, with 1 erect ovule in each cell, 
stigma 4- (or 5-) lobed; fruit a small berry, containing 4 or 5 
bony triquetrous pyrenes.—Sp. 80; 52 in AZ. B. Lud. 


I have mainly followed the Fl. B. Ind. in this difficult genus, but feel 
no certainty as to the limitations or characters of Sp. 2-8. These dingy, 
unattractive shrubs form an undergrowth in shady forests, and require 
further investigation; I do not find the cal.-segm. or number of lat. veins 
very constant. All our species are considered to be endemic, but a 
critical comparison with those of S. India is needed. 


Fl. surrounded with long bracts; cymes peduncled 1. L. MOONII. 
Fl. without bracts. 
Cymes sessile. 
Cal.-segm. linear-lanceolate. 


L. thick, hairy beneath . : : . 2. L. THWAITESII. § 4 
L. thin, glabrous beneath . ‘ s . 3. L. RHINOPHYLLUS. 
Cal.-segm. triangular. 
Cal.-segm. obtuse . : ; : . 4. L. WALKERIANUS. 
Cal.-segm. acute. 
L. not cordate at base. 
L. hairy beneath; fl. in clusters . 5. L. GARDNERI. 
L. glabrous; fl. usually solitary . 6. L. OLIGANTHUS. 
L. cordate at base . 5 , 7. L. OBLIQUUS. 
Cal.-segm. tooth-like (limb truncate) 8. L. STRIGOSUS. 
Cymes on a short peduncle g. L. VARIANS. 


1. &. Moonii, Wight in Calc. Journ. Nat. Hist. vi. 504 (1845). 

Mephitidia Mooniz, Thw. Enum. 145, 420. C. P. 92, 3997. 

Fl]. B. Ind. iti. 180. 

A slightly branched shrub, branches stout, usually very 
hispid with shaggy hair; 1. 3-6in., oval or lanceolate, acute 
or rounded at base, acuminate, subacute, more or less hispid 
on both sides, especially beneath, often bullate above, lat. 
veins 7 or 8, very prominent beneath, petiole $in., with thick 
spreading coarse hair, stip. ovate, obtuse, hairy, caducous; 
cyines dense, capitate on short, stout, hispid peduncles or 
nearly sessile, several in each axil, bracts numerous, as long 
as fl., the outer ones broady ovate, the others linear, very 
strongly hispid or ciliate with coarse hair; cal.-segm. long, 
linear, acuminate, hispid; cor.-lobes short; berry }in., broadly 
ovoid, hispid, scarlet. 


Lasianthus.] Rubwacee. 365 


Moist low country up to 3000 ft.; ratherrare. Sabaragamuwa Dist. fre- 
quent ; Morowak Korale ; Hewesse ; below Adam’s Peak. F1]. March; white. 

Endemic. 

Occasionally, as in C. P. 3997, the stems and upper surface of the 
leaves are glabrous, as was the case with the original specimen described 
by Wight; in the Hewesse specimens when living the hairs were a 
brilliant pink in colour, giving the plant a very ornamental appearance. 


2, L. Thwaitesii, H%. f Fl. B. Ind. iii. 185 (1880). 

Mephitidia tomentosa, Thw. Enum. 145. L. strigosus, Thw. Enum, 
420 (part). C. P. 2711. 

Fl. B. Ind. iii. 185. 

A shrub, branchlets slightly compressed when young, 
densely tomentose with brown hair; |. rather small, 2-34 in., 
narrowly lanceolate, acute at base, tapering into long attenuate 
acute apex, glabrous and shining above, tomentose-hairy 
beneath, rather thick, lat. veins 5, much curved, prominent 
beneath, petiole 4in., stip. pubescent; fl. solitary or few; cal. 
hairy, segm. usually linear-lanceolate, as long as cor.-tube; 
cor. hairy outside; berry 4in., crowned with cal.-segm., hairy, 
black, pyrenes oblong, channelled down ventral side. 

Var. 6, nitidus, 7hiw. zz Fl. B. Ind. iii. 189. C. P. 3910. 

L. rather smaller, glabrous and shining on both surfaces, 
or puberulous on veins beneath. 

Moist region at about 3000ft.; very rare. Only at Laxapanagala in 
Ambagamuwa in 1853. Var. @ more common; Adam’s Peak; Hakgala. 
Fl. Feb., March; white. 

Endemic. 

The Fl. B. Ind. refers var. 6 to the L. stvZgosus of that work, with 
which it does not at all agree, but is connected with the present type 
by pubescent forms. Many fl. are abortive. 


3. L. rhinophyllus, 7iw. Enum. 420 (1864). 

Mephitidia rhinophyllus, Thw. Enum. 145 (by error vizzophyllus). 
C.-P. 361. 

Fl. B. Ind. iii. 185 (ZL. rhzzophyllus). 

A shrub, branchlets elongated, slender, compressed when 
young, adpressed-pubescent; |. rather small, 2—3 in., lanceolate, 
acute at base, long-acuminate, acute, glabrous on both sides, 
thin, lat. veins 5 or 6, oblique, delicate; fl. solitary or few; 
cal. adpressed-hairy, segm. linear-lanceolate; fruit not seen. 

Moist region; very rare. Adam’s Peak (Gardner); Kukul Korale. 
Fl, June; white. 

Endemic. 

I have but scanty material for this, which is very near the last. 


4. lL. Walkerianus, Wight in Calc. Journ. Nat. Hist. vi. 512 
(1845). 

Mephitidia Walkeriana, Thw. Enum, 146. JL. protractus, Thw. (part) 
Enum. 429. C. P. 289 (340). 

FI. B. Ind. iii. 186. 


366 Rubiacee. [Lasianthus. 


Branches puberulous or nearly glabrous, compressed when 
young; |. 3-5 in., lanceolate or linear-lanceolate, tapering to 
base, long-acuminate, very acute almost mucronate at apex, 
coriaceous, glabrous on both sides or midrib beneath slightly 
pubescent, lat. veins 6-8, slender, petiole very short; fl. in 
crowded fascicles without bracts; cal. pubescent, segm. short, 
triangular, rather obtuse; berry small, 4—}in., nearly globose, 
truncate at top, tipped with tooth-like cal.-segm., glabrous. 

Moist region up to 5000 ft.; rather common. Fl. Dec._March; white. 

Endemic. 

A narrow-leaved form (var. /anceolatus, Hk. f.) is as frequent as the 
type. The C. P. numbers 80 and 1728 are quoted for this in Fl. B. Ind., 
but are again quoted for LZ. strzgosus, and C. P. 290 is also given there for 
L. Walkerianus. These numbers are not now represented in Herb. 
Perad., but were originally included by Thwaites under ZL. varzans. 


s, G. Gardneri, 12. f Fl. B. Ind. 186 (1880). 

Mephitidia Gardnert, Thw. Enum. 145. JL. strigosus, Thw. Enum. 
420 (part). C. P. 348, 3985 (?). 

Fl. B. Ind. ii. 186. 

Branches stout, much compressed when young, more or 
less covered with brown hair, becoming glabrous; 1. rather 
large, 4-7 in., lanceolate, acute at both ends, shortly acuminate, 
glabrous above, more or less hairy on veins beneath, coria- 
ceous, lat. veins 5-7, strong, very prominent beneath, petiole 
jin., stip. narrowly triangular, acute, hairy; fl. sessile, in small, 
dense, crowded clusters; cal. hairy, segm. narrowly triangular, 
acute; berry not seen. 

Upper montane zone; rare. Adam’s Peak (Gardner); Nuwara . 


Eliya; Morowak Korale. Fl. July. 
Endemic. 
Thwaites ultimately made one species of this and his J. tomentosa 


(L. Thwatteszz), and considered it to be L. strigosus, Wight. They are 
certainly very closely allied. 


6. G. oliganthus, 7iw. Enum. 420 (1864). 

Mephitidia oligantha, Thw. Enum. 146 (excl. var. 8). C. P. 357. 

Fl. B. Ind. 111. 186. 

Branches slender, slightly rough with adpressed hair; 1. 
2-4 in., acute or obtuse at base, acuminate, acute, with long 
acuminate apex, quite glabrous, rather thin, lat. veins 4 or 5, 
stip. small, narrow; fl. solitary (or 2 or 3), very small, sessile; 
cal.-segm. narrowly triangular, acute; berry very small, about 
4 in., depressed, truncate at top. 

Moist region, 1000-4000 ft.; common. MHantane (Gardner); about 


Peradeniya; Palabadala; Morowak Korale. Fl. March. 
Endemic. 


Seems to flower but rarely. 


Lasianthus.] Rubtracee. 367 


7. L. obliquus, 7iw. Exum. 420 (1864). 

Mephitidia oligantha, var. obligua, Thw. Enum. 146. C. P. 3418. 

Fl. B. Ind. ili. 192. 

Stems slender, nearly cylindrical, much branched, more 
or less strigose with adpressed hair, buds hairy; 1. 2}-53 in., 
nearly sessile, ovate or oblong-ovate, cordate and unequal- 
sided at base, acuminate, acute, glabrous and very shining 
above, pale and with the veins finely pubescent beneath, 
rather thin but stiff, lat. veins 4 or 5, much curved, fine and 
slender but prominent, stip. very small, soon falling ; fl. very 
small, sessile, solitary (or 2 or 3); cal.-segm. small, triangular, 
acute; cor.-lobes shorter than tube; berry 4 in., depressed- 
globose, crowned with tooth-like cal.-segm., smooth, very soft, 
black, pyrenes nodular on surface. 

Moist region up to 3000 ft.; rather rare. Singhe Raja Forest; near 
Balangoda; Morowak Korale; Madagatte, S. Prov.; Kitulgala. FI. 
March, April; white. 

Endemic. 


8. L. strigosus, Wight in Calc. Journ. Nat. Hist. vi. 512 (1845). 

Mephitidia strigosa, Thw. Enum. 146. L. Walkertanus, Thw. 
Enum. 420. C. P. 1725. 

Fl. B. Ind. iii. 189. 

Stems more or less strigose with adpressed hair, cylindrical, 
stout; |. large, 3-6 in., oblong-lanceolate, acute at base, acu- 
minate, mucronate (the midrib protruded), glabrous and shining 
above, paler and strongly pubescent on veins beneath, lat. 
veins 7-9 (usually 8), spreading, strongly marked, as are also 
the connecting veinlets, petiole about }in., stip. small, trian- 
gular, acuminate; fl. largest in the genus, sessile, crowded ; 
cal. slightly pubescent, limb broadly tubular, truncate, ciliate, 
segm. small, tooth-like; cor. hairy outside, tube 1 in., lobes 
shorter than tube; berry 4in., nearly globose, crowned with 
tubular cal.-limb (which is sometimes deciduous). 

Var. B, protractus, zk. f. Mephitidia protracta, Thw. Enum. 146. 
L. protractus, Thw. Enum. 420 (in part). C. P. 354. 

L. 6-8 in., very much attenuate to apex, lat. veins very 
oblique. 

Moist and intermediate country from 1000-5000 ft.; common, and 
often gregarious. Fl. all the year; white. 

Endemic. 

This is the only species that extends beyond the moist region. In 
woods north of Kurunegala and on the slopes of the isolated Ritigala 
Mountain, it forms the principal undergrowth. Var. £8 was collected by 
Gardner, but the locality is doubtful; the specimens are too imperfect 
to enable it to be referred to this species with any certainty. Often (like 
other Rudiacez) called ‘ Wal-kopi’ by the Sinhalese. The upper surface 
of the leaves is sometimes beautifully variegated with yellow. 


368 Rubracee. [Saprosma. 


9. L. varians, 7hw. Enum. 420 (1864). 

Mephitidia varians, Thw. Enum. 146. C. P. 57. 

Fl. B. Ind. iii. 191. 

A much-branched shrub, branchlets compressed when 
young, thickened at nodes, glabrous; |. usually small, 1-3 in.,. 
oval or lanceolate or oblong-lanceolate, acute at base, mu- 
cronate with excurrent midrib at apex, margins often recurved,, 
quite glabrous, very coriaceous, lat. veins scarcely distinguish- 
able from the general reticulate prominent venation; fl. sessile 
or on very short ped., in small clusters, borne on short 
peduncles sometimes nearly 4in. long; cal.-limb cup-shaped,. 
segm. usually Tenge ae acute ; cor. glabrous outside, tube 
short ; berry 4 t-# in., crowned with erect tooth-like cal. -segm.,. 
broadly oven. place 

Upper montane zone; common. Nuwara Eliya; Pedurutalagala; 
Horton Plains; Elk Plains; Adam’s Peak. Fl. Feb.-May; white. 

Endemic. 

The FI. B. Ind. would apparently restrict this to specimens with the. 


inflorescence borne on a peduncle. Easily distinguished from the rest by 
the quite different venation of the leaves. 


43. SAPROSMA, Llume. 


Shrubs, stip. connate, with stiff bristles on margin, deci- 
duous; fl. moderate-sized, solitary or in threes, axillary or 
terminal; cal.-limb cup-shaped, segm. 4, small; cor.-tube short, 
hairy at mouth, lobes 4, induplicate-valvate in bud; stam. 4, 
anth. oblong; ov. 2-celled, with 1 erect ovule in each cell, 
stigmas 2, fruit a berry, containing 2 thin crustaceous pyrenes. 
—Sp. 8; 7 in Fl. B. Ind. 


- 


L. sessile, subcordate at base . : : : . I. S. INDICUM. 

L. petiolate, acute at base. 
Branchlets scabrous : : : : 4 . 2. S. SCABRIDUM. 
Branchlets smooth. : : : : : . 3. S. ZEYLANICUM. 


S. indicum, Dalz. in Kew Journ. Bot. iii. 37 (1851). 

Seriisa Gardnert, Thw. Enum. 150. C. P. 82. 

Fl. B. Ind. ii. 192. Bedd. Ic. Pl. Ind. One 16 (Serissa glomerata). 

A bush, branches slender, dichotomous, widened and flat 
below each node, yellowish-brown, smoothand shining; 1. 2-4 
in., nearly sessile, ovate, rounded or subcordate at base, 
acuminate, acute at apex, glabrous, stip. small with 3 bristles; 
fl. rather small, on short ped., I or 3 at apex of twigs between 
last pair of 1.; cal.-limb short, segm. narrowly triangular, 
acuminate; cor.-lobes ovate, acute as long as tube; berry 
nearly 4in., broadly ovoid, shining, blue-black. 


Saprosma.| Rubtacee. 369. 


Montane zone; very rare. Only collected in forests near Galagama 
below Horton Plains, where it was first found by Gardner. Fl. April, 
May; white. 

Also in S. India. 

The Ceylon plant has broader and more cordate-based leaves than 
the Indian one, and is called var. Gardner7 in FI. B. Ind. 


2. S. scabridum, Zedd. Fl. Sylv. For. Man. 134, 12 (1873 °). 

Serissa scabrida, Thw. Enum. 151. C. P. 3306. 

FI. B. Ind. iii. 193. 

Branchlets rough, with short stiff hair; 1. 2-3 in., oval, 
acute at base, slightly acuminate, acute, glabrous above, 
pilose on veins beneath, petiole }in., stip. scabrous, with long 
bristles; fl. solitary, terminal; cal-segm. lanceolate-linear ; 
berry (unripe) about 4 in., globose. 

Moist low country; very rare. An imperfect specimen in Hb. Perad., 
with a single nearly ripe fruit and no flowers, marked ‘ Near Ratnapura, 
March, 1853,’ is all the material for this species. 

Endemic. 


3. S. zeylanicum, Bedi. Fl. Sylv. For. Man. 136, 12 (1873 ?). 

Dysodidendron* zeylanicum, Gardn. in Cale. Journ. Nat. Hist. vii. 2. 
Serissa zeylanica, Thw. Enum. iso. C. P. 261. 

Fl. B. Ind. iii. 193. Bedd. Ic. Pl. Ind. Or. t. 15 (Serzssa Wighitiz). 

A shrub, 8-12 ft., with slender erect branches, branchlets 
very slender, brittle, much flattened and dilated below the 
nodes, bark pale brownish-yellow, smooth, twigs glabrous ; 
]. usually 2-4 in., lanceolate or oval, acute at base, acuminate, 
acute, often very shallowly and irregularly crenate near the 
end, glabrous, shining above, paler beneath, lat. veins much 
curved, prominent beneath, with deep glandular pits in axils, 
petiole very short, stip. with short bristles; fl. rather large, 
axillary (often from axils of fallen 1.) or terminal, ped. 4—$ in., 
slender, glabrous, with 2 small linear bracts below the middle, 
and often also beneath the fl.; cal. glabrous, segm. broad, 
shallow, apiculate; cor.-tube funnel-shaped, lobes much longer 
than tube, broadly ovate, acute, spreading, hairy above, thick, 
the broad margin thinner, turned upwards and crisped (in 
zstivation strongly induplicate, the bud being bluntly 
quadrangular); fil. very short; stigmas linear, spreading ; 
berry over ? in., obovate-ovoid, tipped with small cal.-segm., 
smooth and shining, bright blue. 

Montane zone in forests 4000-7000 ft.; common. FI. March, April, 
and August; pale sulphur-yellow. 


Also in S. Indian hills. 
The leaves and flowers are extremely fetid, especially when bruised or 


* From the very fetid odour of the plant. 
PART II. BB 


370 Rubiacee. [Spermacoce. 


withering. The colour of the fruit is an intense metallic lapis-lazuli blue. 
A small, narrow-leaved form occurs in the higher mountains. 


(44. HYDROPHYLAX, Linz. 7 


A prostrate perennial herb, stip. large, adnate to 1., forming 
a sheath; fl. rather large, solitary, axillary; cal.-segm. 4, long, 
persistent; cor.-tube broadly funnel-shaped, with a ring of 
hair at mouth, lobes 4, valvate, stam. 4, inserted in throat of 
cor., fil. slender, exserted ; ov. 2-celled, with a single ovule in 
each cell peltately attached to septum, stigma 2-lobed ; fruit 
dry, corky, compressed with sharp edges, 2-celled, 2-seeded.— 
Spr 371 i ee 7a. 


H. maritima, L. 7 Suppl. Plant.126 (1781). Mudu-gétakola, S. 

Sarissus anceps, Gaertn. Fruct.i.118. Moon Cat. 10. Thw. Enum. 
T5ile pen Come lO 7.26 

Fl. B. Ind. i. 199. Wight, Ic. t. 760. 


Perennial herb, stems numerous, prostrate, rooting at 
moles, much-branched, cylindrical, stout, succulent, glabrous ; 
1, }-# in., sessile, more or less recurved, spathulate- oval, acute, 
very fleshy, smooth or slightly papillose, ‘stip. large, forming 
with adnate l-bases a cup round the stem, truncate, slightly 
ciliate, persistent; fl. sessile; cal-segm. linear-lanceolate, 
acute, rather shorter than ov.; cor.-lobes ovate, acute, much 
shorter than tube; ov. 4-angled, glabrous; fruit 4 in. , oblong- 
ovoid, much compressed, crowned with large sai -segm., with 
3 prominent ribs on each side, glabrous and shining. 


Sandy seashores; common round the island. FI. all the year; 
pinkish-lilac. 

Also in S. India. 

Gaertner gives as the Sinhalese name for this, ‘ Hin-taccada.’ 


45. SPERMACOCE, Liuz. 


Annual herbs, stems quadrangular, stip. adnate to l.-base, 
forming a cup round the stem, strongly ciliate or pectinate ; 
fl. small, sessile in axillary clusters, forming whorls; cal.-segm. 
4, linear; cor.-tube long, lobes 4, valvate; stam. 4, inserted in 
throat of cor., ov. 2-celled, with a single ovule in each cell ; 
fruit a small dry capsule, septicidally dehiscent into 2 carp., 
one or both of which again dehisces ventrally; seeds oblong, 
with a groove down ventral surface-—Sp. 150; 4 in FZ. B. Ind. 


Capsule with both carp. ventrally dehiscent. 
L. linear ; capsule 5 sin. , . : - : = I) S. SERICES 
L. oval; capsule +} jsin. . . 2. S. OCYMOIDES. 
Capsule with only one carp. ventrally ‘dehiscent . . 36.5. HISPIDAS 


Spermacoce.| Rubiacee. a7 


1. S. stricta, Z. ~ Suppl. Plant. 120 (1781). 

C. P. 1695 (part). : 

Fl. B. Ind. iii. 200. 

Annual, stems slender, thickened at nodes, erect, quadran- 
gular, slightly branched, angles slightly hispid, internodes 
very long; |. I-2in., sessile but much tapering to narrow 
base, linear, acute, mucronate, margin often revolute, more or 
less scabrous on both sides, bristles of stip. numerous, long, 
filiform; fl. few, clusters dense; cal.-segm. filiform, hairy; cor. 
minute; capsule }in., broadly ovoid, crowned with long cal.- 
segm., hairy, dehiscent from above, each carpel dehiscing 
ventrally; seed narrowly oblong, black, dull. 

i country; apparently rare. Colombo (Ferguson). FI. August; 
white. 


Also in India and Trop. Asia and Africa generally. 
Thwaites did not distinguish this from S. hzspzda. 


2. S. ocymoides, Burm. f. Fl. Ind. 34 (1768). 

Borreria ocymoides, DC., Thw. Enum. 151. C. P. 1684. 

Fl. B. Ind. iii. 200. Burm. f. Fl. Ind. t. 13, f. 1. 

Annual herb, stem slender, 6-12 in. high, branched at 
base, quadrangular, glabrous or nearly so; |. sessile or shortly 
petioled, $—# in., oval, tapering at base, acute, nearly glabrous, 
thin, stip. short, scarcely forming a cup, bristles long; fl. very 
small, cymes dense, globose; cal.-segm. linear, apiculate, 
nearly glabrous; capsule very small, 7g in., oblong-globose, 
glabrous, thin, white, both carp. ventrally dehiscent ; seeds 
black. 


Moist low country; rare. Kalutara (Moon); Galle (Gardner) ; 
Colombo (Ferguson). 

Also in India, Malaya, Trop. Africa, and Mauritius. 

Has the habit of Ammannia. I have not met with this. 


3. S. hispida, Z. SZ. P/. 102 (1753). Hin-geta-kola, S. Yar, 
Nattaichchuri, 7. 

Herm. Mus. 3. Burm. Thes. 103. Fl. Zeyl. n. 62. Moon Cat. to. 
Thw. Enum. 151. C. P. 1695 (part), 1696. 

Fl. B. Ind. iii. 200. Burm. Thes. t. 20, f. 3. 

Annual (sometimes perennial?), stems prostrate, long, 
sharply quadrangular, roughly hairy on the angles, stiff, inter- 
nodes generally long; 1. sessile, 4-1 in., oblong-oval, acute, 
slightly pubescent on both sides, scabrous-ciliate on margin, 
rather thick, veins inconspicuous, stip. short, membranous, 
bristles few, long; fl. few, within the stipular cup, bracts long, 
filiform, hyaline; cal.-segm. twice as long as ov., linear, ciliate ; 
cor.-tube slender, lobes short, bristly on back; capsule } in., 
oblong-ovoid, hairy, crowned with long cal.-segm., one carp. 


372 Rubacez. [Rubéa. 


only ventrally dehiscent, the other closed by the septum 
remaining attached to it; seeds smooth, brown. 
Low country in dry waste places; verycommon. FI. March-August; 


pale violet. 

Throughout India, Malaya, and S. China. 

A maritime state grows in the sands at Kalpitiya; it is almost shrubby, 
with closely placed, small, thick, very bristly leaves with cartilaginous 


margins, and a larger fruit. 
Hermann gives the S. name for this ‘ Aswanna.’ 


46. RUBIA, Linz. 


Perennial, 1. in whorls of 4; fl. small, in axillary cymes; 
cal.-limb 0; cor.-tube campanulate, lobes 4, valvate; stam. 4, 
very small, inserted in tube; ov. 2-celled with 1 erect ovule in 
each cell, styles 2, stigmas globose; fruit didymous, fleshy; 
seeds globose——Sp. 30; 8 in FZ. B. Ind. 


R. cordifolia, Z. Syst. Nat. ed. 12, iii. (1768). Manda-madini- 
wel, Yogana-wel, S. 

R. secunda, Moon Cat. 10. Thw. Enum. 151. C. P. 2705. 

FI. B. Ind. iii. 202. Wight, Ill. t. 128 bis, f. 1. 

Perennial, roots very long, cylindrical, flexuose, with a 
thick red cortex, stems very long (often many yards), becom- 
ing slightly woody at base, flexible, tough, cylindrical, bark 
white, branches scandent by means of the very numerous 
divaricate or deflexed branchlets and petioles, quadrangular, 
sometimes prickly on the angles, smooth, shining; 1.4 ina 
whorl, #-14 in., ovate, cordate at base, tapering to acute 
apex, scabrous above and on the veins beneath, and 
especially on the margin, with small, white recurved prickles, 
5- or 7-veined from the base, stiff, often convex above, with 
the apex hooked, sometimes pubescent beneath, petiole about 
twice as long as I., usually deflexed, tapering, stiff and branch- 
like, deeply channelled above, triangular, with many sharp 
recurved prickles on the angles; fl. on short glabrous ped., 
cymes lax, trichotomous, glabrous; cor.-tube thick, very short, 
lobes ovate, obtuse; fruit about 4 in., very didymous, the carp. 
almost distinct, smooth, shining, purplish-black. © 

Bushy places in montane zone from 3000-7000 ft.; verycommon. FI. 
August, September; greenish-white. 
ne Also in the mountains of India and Java, and in N. E. Asia and Trop. 

T1Ca. 

Our Ceylon plant usually has the stem almost destitute of prickles. 
Mocn seems to have first noticed this plant ; his locality is Walapane. I 
pee net been able to get a satisfactory S. name; those above given are 

OUDTIUL. 


Galium.] Rubiacee, 373 


The roots afford an excellent red dye, but it does not appear to be 
used here. 


47. GALIUM, L777. 


Slender herb, |. 4-6 in a whorl; fi. minute, sessile, axillary; 
cal.-segm. 0; cor. rotate, without a tube, lobes 4, cut almost to 
base, valvate; stam. 4, inserted at mouth of cor., anth. globose; 
ov. 2-celled, style bifid, stigmas globose; fruit very didymous, 
tipped with short styles—Sp. 150; 20 in FZ. B. Ind. 

A genus of Temperate climates; one other species occurs in the 
Nilgiris. 

G. asperifolium, Wall. in Roxb. Fl. Ind. (ed. Wall.) 1. 381 (1820). 

Thw. Enum. 151. C. P. 1682. 

Fl. B. Ind. ili. 207 (G. Mollugo). 

Stems very slender, prostrate, rooting at the nodes below, 
2 or 3 ft. long, weak, much-branched, 4-angled, glabrous or 
very slightly rough on the angles; ]. 4-6 in a whorl, small, 
4-4 in., sessile, lanceolate or linear-oblong, narrowed to base, 
obtuse and cuspidate at apex, glabrous, with a few coarse 
recurved prickles on midrib beneath, and on slightly recurved 
margin; fl. solitary in axils of minute bracts, forming a small 
dichotomous leafy inflor.; cor.-lobes ovate, sharply acuminate; 
stam. much shorter than cor., erect; fruit glabrous, covered 
with blunt warts all over. 

Upper montane zone to 7ooo ft.; common. FI. August, September; 
pale greenish-white. 


Also in mountains of India. 
In the FI. B. Ind. this is referred to the widely spread species (common 


in England) G. Mollugo, L. Dries black. 


Reba ee 


tas 


INDEX TO PARTS I. AND II. 


BOTANICAL NAMES. 


[Natural Orders are printed in small capitals; synonyms, introductions, errors, &c., 


Abelmoschus angulosus, Thw. 


i. 156 
Jiculneus, W. and A. 155 
moschatus, Moench. 156 
Aberia Gardneri, Clos. i. 74 
Abrus precatorius, Z. ii. 57 
pulchellus, Wall. 57 
Abutilon asiaticum, G. Doz, i. 


T44 
crispum, G. Don, 146 
graveolens, W. and A. 145 
indicum, G. Don, 145 
muticum, G. Doz, 145 
polyandrum, W.and A. 144 
Acacia arabica, W7d/d. ii. 122 
ceesia, Wzlld. 127 
Catechu, Thw. 125 
concinna, DC. 127 
eburnea, W7//d. 114 
Farnesiana, Willd. 122 
ferruginea, DC. 126 
Intsza, Willd. 127 
leucophleea, W7d/d. 125 
pennata, W2d/d. 127 
planifrons, W. and A. 123 
scandens, Willd. 119 
Suma, Kurz. 126 
Sundra, DC. 125 
tomentosa, W2d/d. 124 
Acanthonotus echinatus, 
Benth. ii. 21 
Achyranthes corymbosa, L.i. 88 
Acmena lanceolata, Thw. ii. 
172 
zeylanica, Thw. 171 
Acranthera zeylanica, Arz. ii. 


324 
Acronychia laurifolia, BZ. i. 216 
pedunculata, Walp. 216 
Acrotrema bullatum, Zw. i. 7 
dissectum, Thw. 8 
Gardneri, Thw. 8 
intermedium, 7hw. 8 
lanceolatum, //Z, 8 
lyratum, Thw. 9 
sylvaticum, Thw. 8 
Thwaitesii, /4. f. and Th. 9 
uniflorum, //4. 7 
Walkeri, Wight, 8 


in italic type. | 


Adenanthera bicolor, 1/07, ii. 
120 
pavonina, Z. 120 
Adina cordifolia, Hz. f. ii. 203 
a lasiopetala, Chozs. 


Echnanive deltoidea, Arn. ii. 
257 
epigea, Arn. 258 
seylanica, Thw. 259 
AB giceras minus, Gaertn. ii. 1 
Egle Marmelos, Corr. i. 229 
eschynomene aspera, ZL. ii. 39 
indica, Z. 38 
pumila, L. 38 
Sesban, L. 34. 
Agasta indica, Miers, ii. 189 
Aglaia apiocarpa, Azern, i. 
245 
odorata, Lour. 247 
Roxburghiana, 142g. 246 
Agrimonia Lupatorcum, Thw. 
ii. 141 
zeylanica, Joon, 141 
Ailantus excelsa, Roxb. i. 230 
malabarica, DC. 230 
Alangium decapetalum, Lam. 
ii. 285 
glandulosum, Thw. 286 
hexapetalum, Lam. 285 
Lamarckii, Thw. 285 
Albizzia amara, Bozv. ii. 130 
Lebbek, Benth. 128 
moluccana, Miq. 131 
odoratissima, Bexth, 129 
stipulata, Bozv. 129 
Alchemilla indica, Gardn. ii. 
140 
vulgaris, Arn. 140 
Allccophania Arnottiz, Hk. f. 
ii. 30% 
decipiens, Tw. 301 
Allophylus Cobbe, 27. i. 303 
hispidus, Zvzm. 303 
zeylanicus, LZ, 302 
Alphonsea lutea, Hk, f. and 
Th. i. 36 
sclerocarpa, Thw. 37 
zeylanica, 1k. f. and Th. 36 


Adansonia digitata, Li. 159 | Alsine nervosum, Moon, i. 87 


Alsodeia decora, Trzm. i. 69 
virgata, Hk. f. and Th. 69 
zeylanica, Thw. 68 

Alyssicarpus bupleurifolius, 

DC. ii. 44 
Heyneanus, Wight, 45 
longifolius, W. and A. 45 
monilifer, DC. 43 
nummularifolius, DC. 44 
rugosus, DC. 45 
scariosus, Grah. 45 
vaginalis, DC. 44 

Ameletia indica, DC. ii. 223 

Ammannia baccifera, Z. ii. 224 
cordata, W. and A, 225 
debilis, Ait. 224 
lanceolata, Heyze, 225 
octandra, LZ. f. 225 
pentandra, Roxd, 224 
peploides, Spreng. 223 
Rotala, /. 7, 224 
salicifolia, Thw. 225 

Amoora Rohituka, W and A. 

i, 249 

Ampelocissus erioclada, 

Planch. i. 289 
tomentosa, Planch, 288 

Amyris zeylanica, Retz. i. 239 

ANACARDIACE, 1. 316 

Anacardium occidentale, L., i. 


SL hee 
Anagallis esculenta, Mooh, i. 


53 a 
Anamirta paniculata, Coed. i. 
40 
toxifera, Miers, 40 
Anaxagorea luxonensis, Gray, 


i. 27 
seylanica, Hk. f. and Th. 27 
ANCISTROCLADE#, i, 138 
Ancistrocladus Vahlii, Av. i. 
138 
Anemone rivularis, Zam. i. 3 
Anisophyllea zeylanica, Benth. 
li, 157 
Anoda hastata, Cav. i. 141 
Anogeissus latifolia, Wadd. ii. 
162 
Anona astatica, \..\. 32 
muricata, Dun, 32 


376 


Anona reticulata, L. i. 32 
squamosa, Lu. 32 
ANONACEZ, 1. 16 
Anotis nummularia, 2. f. ii. 
318 
nummulariformis, 7vzm. 319 
quadrilocularis, Hz. 7. 318 
Richardiana, A. 7. 319 
Anstrutheria zeylanica,Gardn. 
li. 156 
Anthocephalus Cadamba, AZ. 
ll, 293 
Apodytes Gardneriana, /zers, 
i. 262 
Aquilicia Otillis, Gaertn. i. 296 
Arachis fruticosa, Retz. ii. 36 
ARALIACEA, ii. 281 
Argemone mexicana, L. i. 52 
Artabotrys odoratissimus, B7. 
ih Br 
zeylanicus, Hk. f. and Th. 22 
Arthromischus armatus, Thw. 
i. 22 
Aspalathus indicus, VL. ii. 23 
Atalantia Missionis, Olzv. i. 227 
monophylla, Corr. 226 
racemosa, W. and A. 226 
zeylanica, Olzv. 227 
Atragene zeylanica, L. i. 2 
Atylosia albicans, Benth. ii. 78 
Candollei, W. and A. 78 
tugosa, W. and A. 79 
scarabzeoides, Benth. 79 
Averrhoa Bilimé?, L. i. 200 
Carambola, LL. 200 
Axanthes elliptica, Wight, ii. 
326 
seylanica, Wight, 326 
Axinandra zeylanica, Tw. 231 
Azadirachta indica, A. Juss. i. 


244 


Balanocarpus zeylanicus, 
Trim. i. 130 
Balanopteris Tothila, Gaertn. 
i, 167 
Balsamodendrum Berryi, Arz. 
i. 237 
caudatum, March. 236 
Bannisteria bengalenszs, L. i. 
LOST 
Barringtonia acutangula, 
Gaertn. il. 191 
racemosa, B/. 189 
speciosa, Forst. 189 
zeylanica, Gardz. 190 
Bauhinia acuminata, L. ii. 116 
anguina, Roxb. 117 
parviflora, Vahl, 116 
purpurea, L. 117 
racemosa; Lam. 116 
tomentosa. Z. 116 
Begonia cordifolia, Tw. ii, 262 
dipetala, Grah. 264 
malabarica, Lam. 264 
rupestris, Moon, 263 


Begonia subpeltata, Wzghz, ii. 
264. 
tenera, Dryand. 263 
Thwaitesii, Hook. 264 
BEGONIACEZ;, ii. 262 
Benincasa cerifera, Savi, ii. 252 
BERBERIDEA, i. 48 
Berberis aristata, DC. i. 48 
Berchemia parvifiora, Thw. i. 
284 
Bergera Koenigit, L, i. 220 
nttida, Thw. 221 
Bergiaammanioides, Roxd.i.g2 
verticillata, W2z2/d. 92 
Berrya Ammonilla, Roxé. i. 173 
Bigamea, Koen. i. 138 
Biophytum Candolleanum, 
Wight, i. 198 
intermedium, Wzght, 199 
nervifolium, Tw. 198 
nudum, Wzght, 198 
proliferum, Wzghz, 199 
sensitivum, DC. 197 
Reinwardtiz, Hk. f. 198 
Bixa Orellana, L. i. 70 
BIXACEZ, i. 69 
Blackwellia zeylanica, Gardn. 
iil. 239 
Blastania Garcini,Cogn.ii.260 
Bocagea coriacea, Hk. f. and 
Th. i. 34 
obliqua Hz. f. and Th. 33 
Thwaitesii, ZZ. f, and Th. 33 
Bombax, Cezda, L. i. 160 
gossypinum, L.. 70 
malabaricum, DC. 160 
pentandrum, L. 161% 
Borreria ocymoides, DC. ii. 371 
tetracocca, Thw. 302 
Boswellia glabra, Roxb. i. 238 
Brachypterum Benthamiz, 
Thw. ii. 93 
elegans, Thw. 92 
scandens, WW. and A. 91 
Brassica juncea, Ak. f. and 
Wile, i At 
Bruceasumatrana, Roxb. i. 231 
Bruguiera caryophylloides, BZ. 
ii. 154 
gymnorhiza, Lam. 153 
Bryonia cordifolia, L. ii. 248 
deltoidea, Arn. 257 
grandis, Willd. 247 
laciniosa, Z. 254 © 
palmata, L. 241 
palmata, Moon, 244 
tubifiora, W. and A. 246 
umbellata, Moon, 256 
Bryophyllum calycinum, Sal. 
li. 145 
Buchananiaangustifolia, Rox. 
i. 316 
zeylanica, Bl. 317 
Bupleurum falcatum, L. ii. 277 
nervosum, Moon, 277 
virgatum, W. and A. 277 


Ludex to Botanical Names. 


BURSERACEZ&, i. 235 
Bursinopetalum arboreum, 
Wight, ii. 287 

tetrandrum, Wight, 287 

Butea frondosa, Koez. ii. 66 
superba, Roxb. 67 

Butonica racemosa, Juss. ii. 189 
zeylanica, Miers, 190 


Byrsophyllum ellipticum, Bedd. 


li. 329 


CACTACEZ, ii. 266 
Cactus pendulus, Willd. ii. 266 
Cadaba indica, Lam. i. 60 
trifoliata, W. and A. 59 
Cezesalpinia Bonduc, Rox. 11. 98 
Bonducella, Flem. 99 
Corzaria, Thunb, 10r 
digyna, Fott/. 100 
Gleniez, Thw. Lor 
mimosordes, Lam. 100 
Nuga, Azz. 99 
paniculata, Roxb. 99 
pulcherrima, Sw. 99 
Sappan, i. 99 
seplaria, Roxb. 100 
Cajanus indicus, Spreng, ii. 80 
Calliandra (?) geminata, 
Benth. ii. 131 
Callitriche stagnalis, Scop. ii. 
149 
verna, Thw. 149 
Calophyllum acuminatum, 
Moon, i. 102 
angustifolium, Roxb. Loz 
bracteatum, 7w. 102 
Burmanni, Wzght, 99 
Calaba, L. 99 
cordato-oblongum, Zw, 103 
cuneifolium, Zw. to2 
decipiens, Thw. 102 
elatum, Bedd. tor 
Inophyllum, Z. too 
Moonzz, Wight, 99 
pulcherrimum, Wal/. 100 
spectabile, Wzl/d. 99 
Thwaitesii, P27. and Tri. 102 
tomentosum, Wzghé, rot 
trapezifolium, Tw. 103 
Walkerii, Wzght, 104 
Calyptranthes caryophyliata, 
Pers. ii. 174 
caryophyllifolia, Moon, 179 
cordtfolia, Moon, 176 
Cuminz, Moon, 179 
Jambolana, Moon, 175 
Cambogia Gutta, L. 1. 96 
Camellia Thea, Link. i, 112 
Campnosperma zeylanicum, 
Thw. i. 326 
Cananga odorata, Hk. f. and 
Th. i, 22 
Canavalia ensiformis, DC. ii. 67 
gladiata, DC. 67 
obtusifolia, DC. 68 
virosa, W. and A. 67 


Index to Botanical Names. 


Canarium dalsamiferum, 
Moon, i. 239 
brunneum, Bedd, 238 
commune, L. 240 
zeylanicum, B/. 239 
Cansjera Rheedii, Gmel. i. 259 
scandens, Roxb. 259 
Canthium campanulatum, 
Thw. ii. 345 
didymum, Gaertn. f. 343 
lanceolatum, An. 343 
macrocarpum. Thw. 345 
montanum, 7hw. 343 
parviflorum, Lam. 346 
puberulum, 7hw. 344 
Rheedii, DC. 344 
CAPPARIDEA, i. 54 
Capparis drevispina, Thw. i. 61 
divaricata, Lam. 61 
floribunda, Wzght, 64 
grandis, LZ. f 63 
Heyneama, Wadd. 61 
horrida, Z. f. 64 
Moonii, Wzght, 62 
pedunculosa, Wail. 63 
retusella, Thw. 64 
Roxburghii, DC. 62 
sepiaria, Z. 64 
tener, Dalz. 65 
tetrasperma, Thw. 65 
zeylanica, LZ, 61 
CAPRIFOLIACE2, ii. 288 
Capsella Bursa-Pastoris, 
Moench. i. 54 
Carallia calycina, Benth. ii. 155 
integerrima, DC. 155 
seylanica, Arn. 155 
Caranda pedunculata, Gaertn. 


i. 343 
Carapa moluccensis, Lam. i. 
251 
Cardamine africana, L. i. 53 
subumbellata, Hz. f. 53 
Cardiospermum canescens, 
Wall. i. 300 
Corindum, ZL. 300 
Helicacabum, ZL. 299 
Careya arborea, Roxb. ii. 191 
Carria speciosa, Chois. i. 111 
Carum zothum, Cl. ii. 278 
Roxburghianum, Benth. 278 
Stictocarpum, C/. 278 
Caryolobis indica, Gaertn. i. 
119 
CARYOPHYLLACE®, i. 84 
Casearia Championii, Thw. ii. 
237 
coriacea, Thw. 237 
esculenta, Roxb. 237 
qvata, Moon, 237 
tomentosa, Koxb, 238 
varians, Thw. 237 
zeylanica, Thw. 237 
Cassia Absus, L. ii, 109 
alata, Li, 108 
angustissima, Lam, 110 


Cassia auriculata, Z. ii. 106 
Fistula, Z. 103 
florida, Vahl, 108 
glauca, Lam. 109 
hirsuta, L. 106 
Kleinii, 1’. avd A. 110 
levigata, Willd. 106 
marginata, Roxb. 104 
mimosoides, Z. 110 
obtusa, Roxb. 107 
occidentalis, Z. 105 
pumila, LL. 110 
Roxburghiz, DC. 104 
siamea, Lam. 108 
Sophera, Z. 105 
sumatrana, Roxh. 108 
Tagera, Moon, 105 
timoriensis, DC. 108 
tomentosa, Li. 106 
Tora, Z.106 
Wallichiana, DC. 110 
Catenaria laburnifolia, Benth. 
li. 48 
Catha emarginata, G. Don, i. 
273 
fruticosa, Thw. 273 
Cathartocarpus Fistula, Pers. 
lil. 103 
rosea, Moon, 104 
Ceanothus asiaticus, L. 1. 285 
seylanicus, Heyne, 284 
Cedrela serrata, Royle, i. 252 
Toona, Roxb. 252 
CELASTRACE, i. 266 
Celastrus glaucus, Vahl, 271 
paniculatus, Wzd/d. 272 
Centrosema Plumiert, Benth. 


ii. 75 
Cephalandra indica, Naud. ii. 
247 
Cerasiocarpum Bennetiiz, 
Cogn. ii. 259 
zeylanicum, CZ, 259 
Cerastium indicum, W. and 
A, i. 85 
vulgatum, Z. 85 
Ceriops Candolleana, Ava. ii. 
152 
Roxburghiana, Arn. 153 
Chailletia sumatrana, J/zg. i. 
254 , 
CHAILLETIACEZ i, 253 
Chasaliacurviflora, Thw. ii. 362 
Chickrassia tabularis, A. Juss, 
1. 252 
velutina, Roem, 252 
Chloroxylon Swietenia, DC, i. 
a eee 
Cipadessa fruticosa, BZ, i. 245 
Cissampelos convolvulacea, 
Moon, i. 41 
delicatula, Miers, 47 
grallatoria, Miers, 47 
hernandifolia, Willd. 45 
Pareira, LZ. 46 
subpeltata, Miers, 47 


37f 


Cissus acuminata, Thw. i. 292 
adnata, Roxb. 290 
angulata, Lam. 29% 
carnosa, Lam. 294 
edulis, Dalz. 289 
Gardnerz, Thw. 293 
glauca, Thw. 292 
glycosmoides, Planch. 297 
elyptocarpa, Thw. 289 
lanceolaria, Roxb. 296 
latifolia, Moon, 292 
lonchiphylla, Thw. 290 
pallida, Planch, 291 
pedata, Lam. 295 
guadrangularis, L. 289 
repens, Thw. 291 
reticulata, Thw. 294 
retinervia, Planch. 294 
seto-a, Roxb, 296 
tenuctfolza, Heyne, 295 
Thwaitesiz, Planch. 290 
trilobata, Lam. 293 
verrucosa, Moon, 297 
vetigened, Li. 291 

Cistus lobatus, Moon, i. 70 

Citrullus Colocynthis, Schrod, 

li. 253 
vulgarés, Schrod. 253 

Citrus Hystrix, DC. i. 228 
Limonellus, Hassk. 228 
tuberoides, J. W. Benn. 228 

Clausena indica, Olzv. i. 22 
Willdenowii, W. axd A. 222 

Clematis Gouriana, Roxdé. i. 2 
smilacifolia, Wadd. x 

Cleome aspera, oem. i. 56 
Burmanni, W. and A. 56 
Chelidonii, Z. 7, 56 
dodecandra, L, 56 
felina, L. f, 56 
gynandra, Li. 57 
zcosandra, lL. 57 
monophylla, Z. 55 
pentaphylla, L. 57 
tenella, L. f. 55 
viscosa, L. 57 i 

Cleyera emarginata, Gardn. 1. 

108 
eymnanthera,W.and A. 107 
lastopetala, Wight, 108 

Clitoria ternatea, Z. ii. 75 

Coccinea indica, W. and A. ii. 

247 : 

Cocculus Burmanni, DC. i. 47 
orbiculatus, Moon, 46 
macrocarpus, W, and A. 43 
villosus, DC. 44. 

Cochlospermum 

DC. i. 70 

Coffea arabica, L, ii. 353 
travancorensis, W. and A. 

352 
triflora, Moon, 352 
Wightiana, Wall. 353 

Colubrina asiatica, Brongn, i. 

285 


Gossypium, 


378 


COMBRETACEZ, ii. 158 
Combretum acuminatum, 
Roxb. ii. 163 
extensum, Roxd. 164 
ovalifolium, Roxd. 163 
platyphyllum, Van Heurck, 


164 
sarcopterum, Thw. 163 
Thwattesianum, Van 
Heurck, 163 


Wightianum, Thw. 163, 164 
Commiphora caudata, Engl. i. 
236 
CONNARACE#, ii. 1 
Connarusaszaticus, Willd. ii. 2 
Championii, Tw. 3 
monocarpus, Z. 2 
pinnatus, Lam. 2 
santalotdes, Vahl, 1 
untfolzatus, Thw. 3 
Conocarpus latifolia, Roxb, ii. 
162 
Corallocarpus epigzeus, 
Clarke, ii. 258 
Corchorus acutangulus, Lam. 
i. 183 
capsularis, Z. 181 
fascicularis, Zam. 183 
olitorius, Z. 182 
tridens, Z. 183 
urticeefolius, W. and A. 182 
CORNACEZ, ii. 285 
Coscinium  fenestratum, 
Colebr. i. 41 
Cracca maxima, L. ii. 32 
purpurea, L. 3 
senticosa, L. 30 
tinctorza, L. 31 
villosa, L. 33 
CRASSULACES, ii. 143 
Cratzeva Roxburghii, Br. i. 59 
Crotalaria albida, Heyze, ii. 12 
anthylloides, \W. and A. 14 
bifaria, Lf. 11 
biflora, Z. 9 
calycina, Schrank, 14 
clavata, W. and A. 19 
evolvuloides, Wight, 10 
ferruginea, Grah. 10 
fulva, Roxb. 17 
zncana, L. 18 
juncea, Z. 16 
laburnifolia, Z. 19 
linifolia, Z. f. 13 
lunulata, Heyne, 17 
medicaginea, Lam. 18 
multiflora, Bexth. 11 
mysorensis, Roth. 12 
nana, Burm. 13 
nummularia, Willd. 9 
prostrata, Roxb. 9 
quinquefolia, Z. 19 
Tetusa, Z. 1 
rubiginosa, W2Z/d. 11 
semperfiorens, Vent. 16 
striata, DC. 18 


Crotalaria tecta, Heyne, ii. 14 
triquetra, Dalz. 12 
umbellata, Wight, 13 
verrucosa, Z. 15 
Walkeri, Arn. 16 
Wightiana, Grah. 1% 
Willdenowitana, DC. 18 

CRUCIFERZ&, 1. 52 

Crudia zeylanica, Benth. ii. 113 

Ctenolepis Garcini, Clarke, ii. 

260 

Cucumis Lufa, L. ii. 252 
pubescens, W2d/d. 250 
maderaspatanus, Moon, 250 
maderaspatanus, 1. 254 
trigonus, Roxb. 250 

CUCURBITACEA, il. 242 

Cullenia excelsa, Wight, i. 162 

Cyanospermum tomentosum, 

W. and A. ii. 84 

Cyathocalyx zeylanicus, 

Champ. i. 20 

Cyclea Burmanni, Jzevs, i. 47 
peltata, Hk. f. and Th. 47 

Cylista tomentosa, Roxb. ii. 84 

Cyminosma Ankenda, Gaertn. 

i. 216 
pedunculata, DC. 216 

Cynometraramifiora, Z.1i. III 

caulifiora, 1. 112 


Dalbergia arborea, Willd. ii. 


QI 
Championii, Tw. ii. 88 
Srondosa, Roxb. 88 
lanceolaria, LZ. f. 88 
lanceolarza, Moon, 97 
latifolia, Roxb. 88 
monosperma, Dalz. 89 
Mooniana, Thw. 97 
volubtlis, Roxb. 89 
seylanica, Roxb. 88 
DATISCACE, ii. 265 
Delima sarmentosa, Z. i. 5 
Dendrolobium Cephalotes, 
Benth. ii. 47 
umbellatum, Benth. 47 
Dentella repens, Fus¢. ii. 298 
Derris Benthamii, Thw. ii. 93 
oblonga, Benth. 93 
paniculata, Benth. 93 
parviflora, Benth. 92 
robusta, Benth. 92 
scandens, Benxth. 91 
sinuata, Bezth. 94 
uliginosa, Bezth, 92 
Desmanthus cinereus, Willd. ii. 
121 
virgatus, Willd. 122 
Desmodium biarticulatum, 
Benth. ii. 48 
cajanefolium, DC. 52 
capitatum, DC. 53 
catentferum, Arn. 48 
Cephalotes, Wad/. 47 
diffusum, DC. 52 


Index to Botanical Names. 


Desmodium ferrugineum, 
Wall. ii. 52 
gangeticum, DC. 51 
Gardneri, Benth. 50 
gyrans, DC. 56 
gyroides, DC. 56 
heterocarpum, DC. 53 
heterophyllum, DC. 55 
jucundum, 7Thw. 54 
laburnifolium, DC. 48 
latifolium, DC. 51 
letocarpum, Don, 52 
ormocarpoides, DC. 49 
parviflorum, DC. 55 
podocarpum, Thw. 50 
polycarpum, DC. 53 
pulchellum, Bezzth. 48 
rufescens, DC, 52 
Scalpe, DC. 50 
strangulatum, W. and A. 
50, 51 
Thwaitesii, Baker, 51 
triflorum, DC. 54 
triquetrum, DC. 49 
umbellatum, DC. 47 
Wightii, Grah. 52 
Weatlkeri, Arn. 52 
Detaritum zeylanicum, Thw. 
ii, 113 
Dialium ovoideum, Yhw. ii. 
Ii2 
Dicellostyles axillaris, Benth. 
i, I50 
Dicerma biarticulatum, DC. 
ii. 48 
Dichilanthe zeylanica, TAw. ii. 


339 
Dichrostachys cinerea, W. and 
A. il, 121 
Dillenia aguatica, Moon, i. 11 
dentata, Thunb, 11 
indica, Z. 12 
integra, Thunb. 13 
retusa, Thunb. 13 
DILLENIACE4, i. 5 © 
Dimocarpus pupilla, Moon, i. 
309 
Dioclea Fergusoniz, Thw. ii. 69 
teflexa, Hk. 7. 69 
Diploclisia inclyta, Miers, i. 43 
Diplospora Dalzellii, 2. 7. il. 
336 
erythrospora, Bedd. 336 
DIPTEROCARPACE4, 1. 112 
Dipterocarpus glandulosus, 
Thw. 1. 115 
hispidus, Zzw. 114 
insignis, Zw. 116 
oblongifoltus, Thw. 114 
oblongus, A. DC. 114 
scabridus, Zw. 115 
turbinatus, Goertn. f. 115 
turbinatus, Moon, 114. 
zeylanicus, Thw. 114 
Discospermum Dalszellzz, Thw. 
ii. 336 


Index to Botanical Names. 


Discospermum erythrosporum, 
Thw. 336 
Spherocarpa, Bedd. 336 
Discostigma acuminatum, FI. 
and Tri. i. 98 
seylanicum, F\. and Tri. 97 
Dittelasma Rarak, Hk. f. i. 300 
Dodonzxa Burmanniana, DC. 
i. 312 
viscosa, Z. 312 
Dolichos a/éus, Moon, ii 77 
biforus, L. 77 
ciliatus, K7ezz, 77 
falcatus, K/ezn, 77 
gladiatus, Jacq. 67 
Lablab, Z. 76 
luteus, Moon, 73 
medicagineus, Lam. 84 
prurzens, L. 62 
purpureus, Moon, 76 
rotundifoltus, Moon, 67 
scarabeoides, L. 79 
unifiorus, Lam. 76 
vtrosus, Roxb. 67 
Doona affinis, Tw. i. 120 
congestifiora, Thw. 122 
cordifolia, Tw. 122 
Gardneri, Tw. 121 
macrophylla, Thw. 124 
nervosa, 7hw. 121 
oblonga, Thw. 123 
ovalifolia, Thw. 123 
trapezifolia, Tw. 121 
venulosa, Tw. 123 
zeylanica, Thw. 119 
Drosera Burmanni, Vahl, ii.145 
indica, Z. 146 
lunata, Ham. 146 
peltata, Sm. 146 
rotundifolia, L. 145 
DROSERACE:, ii. 145 
Drymaria cordata, W22/d. i. 87 
Dumasia pubescens, DC. ii. 58 
villosa, DC. 58 
Dunbaria ferruginea, W. and 
A. ii. 80 
Heynei, W. and .4. 80 
oblonga, Arn. 80 
Durio zeylanicus, Gardn. i. 162 
zibethinus, Moon, 162 
Dysodidendron zeylanicum, 
Thw. ii. 369 
Dysoxylum_ binectariferum, 
Hk. f. i. 247 
Championii, Hk. f. & Th. 248 
macrocarpum, Thw. 247 


Lccremanthus eximius, Thw. 
i. 310 
Elzocarpus amoenus, 7hw. i. 
185 
copalliferus, Retz. 13% 
coriaceus, Uk. 186 
cuncatus, Willd. 185 
glandulifer, Mast. 187 
integrifolius, Lam. 185 


Elzeocarpus zzdegrzfolius, 
Moon, ii. 156 
montanus, 7hw. i. 186 
obovatus, A7vz. 186 
serratus, Z. 184 
subvillosus, Arz. 186 
zeylanicus, Mast. 187 
Elzodendron glaucum, Pers. 
127.0 
ELATINE&, i. 92 
Eleiotis sororia, DC. 40 
Ellipanthus Thwaitesii, Hz. 7. 


li. 3 

untfoliatus, Hk. f. 3 
Entada scandens, Ben¢h. ii. 119 
Lpithinia malayana, Jack, ii. 


337 
Eriodendron anfractuosum, 
DC. i. 161 
orientale, Steud. 161 
Eriosema chinense, Vog. ii. 81 
Erythrina corallodendrum, L. 
ii. 63 
indica, Lam. 63 
lithosperma, Bl. 64. 
ovalifolia, Rox. 64 
picta, Moon, 64 
suberosa, Roxb. 64 
Erythrospermum phytolac- 
coides, Gardn. i. 72 
Erythroxylon lanceolatum, 2. 
f. i. 19 
lucidum, JZoon, 191 
monogynum, fox). 190 
obtusifolium, Hz. f, 192 
Eucalyptus, ii. 166 
Eugenia acutangula, L. ii. 191 
amoena, Zw. 186 
androsemoides, Bedd. 176 
aprica, Z77im. 186 
aquea, Burm. 169 
assimilis, Duth. 176 
bracteata, Roxb. 182 
capitellata, Arm. 211 
concinna, Thw. 188 
cordifolia, Wzght, 176 
corymbosa, Lam. 174 
cyclophylla, Zw. 177 
cylindrica, Wight, 171 
decora, Thw. 185 
Fergusoni, 77im. 172 
floccifera, Thw. 183 
fulva, Thw. 184 
grandis, Wzght, 170 
Gardneri, Duth, 174 
Heeckeliana, 7rim. 181 
hemisphzerica, Wight, 170 
hypoleuca, Thw. 180 
insignis, Thw. 185 
Jambolana, Lam. 179 
Jambos, 1. 170 
lanceolata, Lam, 172 
lissophylla, Duth. 173 
lucida, Lam. 180 
mabzeoides, Wight, 186 
matlaccensis, 1. 170 


379 


Eugenia Michelli, Zam. ii. 188. 
micrantha, Duth. 175 
Mooniana, Wight, 187 
Neesiana, Wzght, 177 
oligantha, Duzh. 178 
olivifolia, Duth. 178 
operculata, Roxb. 179 
pedunculata, 772m. 187 
phillyreeoides, Trzm. 183 
racemosa, 1. 189 
revoluta, Wzght, 175 
rivulorum, Zw. 184 
rotundata, 7rzm. 185 
rotundifolia, Wzght, 177 
tufofulva, Thw. 183 
sclerophylla, Duzth. 178 
spicata, Lam. 171 
subavenis, Duth. 172 
sylvestris, Moon, 169 
sylvestris, Weght, 175 
terpnophylla, Zw. 181 
Thwaitesii, Duth. 188 
Willdenoviz, Wight, 182 
xanthocarpa, Tw. 182 
zeylanica, Wight, 171 
zeylanica, Willd. 188 
seylanica, Roxb. 182 

Euodia Roxburghiana, Bexth. 

i. 214 
triphylla, Thw. 214 
Euonymus dichotomus, Thw. 
i. 267 
revolutus, Wight, 267 
Thwaitesii, Laws. 267 
Walkeri, Wight, 267 
seylanicus, Moon, 267 
Euphoria Gardneri, Thw. i 
309 
Longana, Lam. 309 
Eupyrena glabra, W. and A. 
li. 338 

Eurya acuminata, DC. i. 110 
chinensis, By. 109 
elliptica, Gardn. 109 
japonica, Thunb. 109 
lasiopetala, Gardn. 108 
parvifolia, Gardn. 109 
membranacea, Gardn. L1o 
zeylanica, Wight, 109 

Evia amara, Comm. i. 327 

FagaraLunu-ankenda, Gaertn, 

i. 214 
triphylla, Roxb. 214 
Fergusonia 7waztesiz, Hk. f 
ii. 302 
zeylanica, /Tk. f. 302 
Feronia elephantum, Cory, i. 
228 

FICOIDE, il. 267 

Filicium decipiens, 7iw. i. 24 

Firmiana colorata, Br. i. 166 

Flacourtia Cataphracta, Roxb, 

1. 73 
inermis, Roxb. 73 
Ramontchi, Sher. 73 
sepiaria, Koxd. 73 


380 


Flemingia 6zfora, Moon, ii. 83 
congesta, Roxb. 87 
lineata, Roxb. 87 
polysperma, Moon, 41 
semzialata, Roxb. 87 
strobilifera, Br. 86 
viscosa, Moon, 85 
Fragaria vesca, L. 138 
193 
Gaertnera racemosa, Roxb. i. 
Galactia tenuiflora, W. and 
Al, itis OR 
Galega maxima, L. i. 32 
purpurea, Li. 31 
tinctoria, Li. 31 
villosa, V4. 33 
Galium asperifolium, W/Z. ii. 


373 
Garcinia Cambogia, Desr. i. 95 
echinocarpa, Tw. 96 
Morella, Desv. 96 
Papilla, Wight, 95 
guesita, Piene, 95 
spicata, Hk. f. 98 
terpnophylla, Tw. 97 
Thwaztesiz, Piene, 98 
seylanica, Roxb. 95, 96 
Gardenia carinata, Thw. ii. 333 
coronaria, Ham. 333 
latifolia, Azt. 332 
Jragrans, Moon, 331 
turgida, Roxb. 333 
uliginosa, Retz. 330 
Garuga brunnea, March. i. 238 
Geissaspis cristata, W. and A. 
rth, 
Eeauiee reniformis, D. Dox, 
ii. 363 
GERANIACEZ, i. 195 
Geranium nepalense, Sweet, 1. 
195 
Gisekia pharnaceoides, L. 11.273 
Gleniea zeylanica, 2. 7. i. 305 
Glinus dictamnoides, L. 11. 270 
Glycine javanica, LZ. li. 59 
labialis, L. f. 60 
mollis, WW. and A. 60 
parviflora, Lam. 60 
tenutfiora, Willd. 65 
villosa, Moon, 77 
viscida, Willd. 41 
Glycosmis arborea, DC. i. 217 
bilocularis, Tw. 218 
pentaphylla, Corr. 217 
Glycycarpus racemosa, Dalz. i. 
325 
Glyptopetalum zeylanicum, 
Thw. i. 268 
Gomphandra axillaris, WadZ. 
i, 261 
coriacea, Wight, 261 
Gomphia angustifolia, Va, i. 
235 
seylanica, DC. 235 
Goniothalamus Gardneri, HA. 
J: and Th. i. 30 


Goniothalamus Hookeri, 7zw. 
i. 30 
reticulatus, Zw. 31 
salicinus, Hk, f. and Th. 31 
Thomsoni, Zw. 31 
Thwaitesi, Hz. f.and Th. 29 
Walkeri, zk. f. and Th. 30 
Gordonia elliptica, Gardz, 1. 
Lit 
speciosa, Chozs. 11 
zeylanica, Wzght, 110 
Gossypium barbadense, Li. 159 
herbaceum, LL. 159 
obtustfolium, Roxb. 159 
Gouania microcarpa, DC, i. 
286 
Grewia asiatica, Z. i. 174 
bracteata, Hleyne, 175 
columnaris, S77. 174 
diptocarpa, Thw. 175 
helicterifolia, Thw. 177 
hirsuta, Vahl, 176 
microcos, L. 177 
orbiculata, Rottl. 175 
orientalis, Z 176 
polygama, Aoxd. 177 
populifoiia, Vah/, 178 
rotundifolia, Thw. 174. 
tilizefolia, Vahl, 175, 
Grifithia fragrans, W. and 
fe Thy BEI 
Gardneri, Thw. 331 
rugulosa, Thw. 331 
Grislea tomentosa, Willd. ii. 226 
Grumilea affints, Thw. ii. 359 
elongata, Wight, 359 
Gardneri, Thw. 358 
Mooniz, Thw. 360 
nigra, Gaertn. 358 
nudipfiora, Thw. 358 
stenophyila, 357 
Wightiana, Hk. f. 358 
Guatteria coffeoides, Thw. i. 
24 
Korintz, Dun. 25 
longifolia, Thw. 24 
montana, Moon, 34 
persicifolia, Hk. f. and Th. 
26 
suberosa, Dun. 25 
Guazuma tomentosa, Kunth. 
hy Typ 
Guettarda speciosa, ZL. i. 338 
Gutlandina Bouduc, L. ii. 98 
GUTTIFERA, i. 94 
Gymnopetalum ¢ubzflorum, 
Cogn. ii. 247 
Wightii, Arz. 246 
Gymnosporia emarginata, HA. 
ff i. 273 
fruticosa, Hk. f. 273 
Gynandropsis pentaphylla, 
Gai, 
speciosa, DC. 58 


Gynostemma laxa, Cog. ii. 260 


pedata, Trim. 260 


Index to Botanical Names. 


Gyrocarpus aszaticus, Willd, 
li. 165 
Jacquini, Roxb, 165 


HALORAGEAZ!,, ii. 147 
Hlaloragis oligantha, Arn. ii. 


147 
oligantha, Wight, 149 
Harpalosia Leflingiw, Wall. i. 
8 


7 
Harpullia imbricata, Thw. i. 
311 
Hebradendron cambogiozdes, 
Grah. i. 96 
Fledera acuminata, Wight, ii. 
282 
emarginata, Moon, 284 
exaltata, Thw. 284. 
racemosa, Wight, 283 
terebinthacea, Vahl, 283 
Vahtliz, Thw. 283 
Hedyotis Auricularia, Z.ii. 313 
Burmanniana, Br. 314, 315 
cerulea, W. and A. 312 
cinereo-viridis, Tw. 308 
coprosmoides, 772m. 306 
cyanescens, Zw. 313 
cymosa, Zw. 305 
evenia, Zw. 304 
fruticosa, LZ. 304 
Gardneri, Tw. 310 
herbacea, Lu. 315 
Heynit, Br. 315 
inamoena, Thw, 313 
Lawsoniz, W. and A. 310 
Lessertiana, A772. 309 
Macrei, Hz. f. 305 
macrophylla, Thw. 307 
maritima, Moon, 316 
membranacea, Tw. 306 
monosperma, W. and A. 319 
Neestana, Arn. 312 
nitida, W. and A. 312 
nodulosa, Arz. 307 | 
nummularia, Arn. 318 
nummulariformts, AM. 319 
obscura, Zw. 305, 306 
paniculata, L. 317 
plantaginifolza, Arn. 311 
guadrilocularis, Thw. 318 
quinquenervia, 7Zw. 310 
racemosa, Lam. 317 
rhinophylla, TZw. 308 
Richardiana, Hk. f, 319 
Thwaitesii, Wk. 7, 307 
trinervia, R. and S$. 316 
umbellata, Lam. 316 
verticillaris, W. awd A. 311 
Walkeri, Arn. 307 
Hedysarum biarticulatum, L. 
ii. 48 
conjugatum, Willd, ii. 35 
diphyllum, Li. 35 
gyrans, L. f. 56 
gangeticum, Iu. 53 
hamatum, L. 36 


Index to Botanical Names. 


Hledysarum heterocarpum, L. 


3 
pay, Willd. 55 
montliferum, L. ii. 43 
mucronatum, Bl. 52 
nummularifolium, L. 21 
pulchellum, L. 48 
sennozdes, Willd. 39 
strobiliferum, L. 86 
triqguetrum, L. 49 
triflorum, L. 54 
umbellatum, L. 47 
vaginale, L. 44 
viscidum, L. 41 
Helicteres Isora, Z. i. 168 


Flelosciadium  Heyneanum, 
DC. 279 
Hemigyrosa canescens, Thw. 


i, 301 
deficiens, Bedd. 302 
trichocarpa, Thw. 301 

Heptapleurum emarginatum, 

Seem. ii. 284 
exaltatum, Seem. 284 
racemosum, Bedd. 283 
stellatum, Gaertn, 283 

Heracleum zepalense, Don, ii. 

280 
Sprengelianum, Thw. 280 
zeylanicum, Gardz. 280 

Heritiera littoralis, Dryand, i. 


167 
Heylandia latcbrosa, DC. ii. 
8 


Hibiscus Abelmoschus, Z. i. 
156 
angulosus, Mast. 156 
cannabinus, L. 154 
collinus, Roxd. 152 
ficulneus, Z. 155 
furcatus, Roxb. 152 
Lampas, Cav. 158 
lunariifolius, Wzd/d. 153 
micranthus, Z. f. 153 
pandurzeformis, Burm. 154 
populneus, L. 158 
vigidus, L. f. 153 
Sabdariffa, 1.. 154 
Solandra, ZL’ Herit. 155 
surattensis, Z, 152 
tiliaceus, ZL. 157 
vitifolius, Z. 154 
zeylanicus, L. 149 
Hippocratea Arnottiana, Wz. 
i. 275 
indica, Willd. 276 
obtusifolia, Roxb. 275 
“iriditora, Moon, 275 
Hiptage Madablota, Gaertn. 
i. 193 
parvifolia, W. and A. 193 
Hugonia ferruginea, W. and 
A. i. 1g0 
Mystax, L. 189 
Homalium zeylanicum, Benth. 
ii. 239 


Hopea cordifolia, Zrzm. i. 
126 
discolor, Tw. 125 
jucunda, 7hw. 125 
Humboldtia laurifolia, Vak, 
li. 115 
Hydnocarpus alpina, Wéghd, 
i. 76 


ib GA 
znebrians, Vahl, 75 
octandra, Thw. 76 
venenata, Gaertn. 75 
Hydrocera angustifolia, BZ, i. 
212 
trifiora, W. and A, 212 
Hydrocotyle usiatica, L. ii. 276 
capitata, Moon, 275 
nepalensis, Ak. 275 
nittidula, Ak. 275 
javanica, Thunb, 275 
rotundifolia, Roxb. 275 
seylanica, DC. 275 
Hydrophylax maritima, Z. 7. 
li. 370 
HYPERICACE#, i. 93 
Hypericum a@urztum, Moon, 
1. 93 
canpestre, Moon, 93 
humifusum, L. 94 
japonicum, Thunb. 93 
mysorense, H/eye, 93 
Hlypserpa cuspidata, Miers, i. 


42 
paucifiora, Miers, 43 
uniflora, Miers, 43 

Lfyptianthera macrocarpa, 

Thw. ii. 334 


Tex denticulata, Wall. i. 265 
Walkeri, Wight and 
Gardn. 264 
Wightiana, Wall. 265 
ILICINEA, i. 264 
Impatiens acaulis, Arz. i. 201 
appendiculata, Arz. 209 
Arnottii, Tw. 211 
Belsamina, Z. 203 
biglandulosa, Moon, 208 
bipartita, Arn. 203 
brevicornu, Arn. 205 
bulbosa, Moon, 201 
cornigera, Arn, 210 
cornuta, 1, 203 
cuspidata, W. and A, 202 
elongata, Avm. 210 
fasciculata, Lum. 201 
flaccida, Arn. 206 
gibbosa, Arn. 205 
glandulifera, Arn. 203 
andis, Heyne, 209 
enslowiana, Arn, 207 
Hookeriana, Arn. 208 
janthina, Zw. 207 
latifolia, L. 202, 203, 206 
leptopoda, Arn. 205 
Leschenaultiz, Wall. 206 
leucantha, Thw. 209 


38% 


Impatiens linearis, A772. 209 
macrophylla, Gardz. 204 
oppositifolia, Z. 202 
repens, Joon, 205 
rosmarinifolia, Retz. 202 
serrata, Moon, 202 
subcordata, Arz. 208 
tenella, Heyne, 202 
trijiora, L. 211 
truncata, Thw. 206 
Walkeri, Hz. 211 

Indigofera Aznz/, L. ii. 27 
aspalathoides, Vahl, 23 
constricta, Trim. 27 
echinata, W2dld. 21 
enneaphylla, Z. 22 
flaccida, Koen. 25 
galegoides, DC. 28 
glabra, Z. 23 
hirsuta, Z. 26 
znamena, Thw. 27 
linifolia, Refz. 22 
parviflora, Heyne, 26 
paucifolia, Ded. 25 
pentaphylla, L. 23 
subulata, Pozr. 25 
tenuifolia, Roftl. 24 
tinctoria, Z. 26 
trifoliata, Z. 24 
trita, Z. ff 25 
viscosa, Lam. 24 
Wightii, Grah. 27 

Tonidium ramosissimum, 7%. 

i. 68 
suffruticosum, Gz2g..67 
Lsauxis Roxburghiana, Thw. 
i, 128 
Ixora acuminata, Thw. ii. 


347 
alba, L. 348 
calycina, Thw. 347 
coccinea, Z. 348 
jucunda, Zw. 348 
parviflora, Vahd, 348 
Thwaitesii, Hz. f. 347 


Jambolifera pedunculata, in 
7, 216; il. 180 

Jambosa aguea, DC, ii. 169 
cylindrica, Thw. 17% 


Jonesia Asoka, Roxb. ii. 114 


pinnata, Willd. 114 
Julostylis angustifolia, Tw. i, 
150 
Jussizea erecta, L. il. 233 
parviflora, Moon, 234. 
repens, LZ. 233 
suffruticosa, L. 233 
villosa, Lam, 233 


Kadsura Wightiana, Avm.i.16 
Kalanchoe floribunda, W, and 
A, ii. 144 
laciniata, DC. 144 
Kandelia Rheedez, W, and A, 


il. 153 


382 Index to Botanical Names. 


Kanilia caryophylloides, Bl. 
ii. 154 
Kayea cuspidata, Pl. and Tri. 
i, 104 
stylosa, Tw. 104 
Kendrickia Walkeri, #2. 7. ii. 
200 
Kleinhovia Hospita, \. 1. 167 
Knoxia corymbosa, Thw. ii. 
34° é 
corymbosa, W2d/d. 340 
hirsuta, Arm. 342 
mollis, W. and A. 340 
platycarpa, Ar7.341 
stricta, Gaertn. 340, 341 
zeylanica, L. 341 
Koon, Gaertn. i. 304 
Kokoona zeylanica, Zhw. i. 
270 
Kurrimia zeylanica, Arz, i. 
2/5 an 
Kydia angustifolia, Arn. i. 150 
axillaris, Thw. 150 


Lablab vulgaris, Savi, ii. 76 

Lagenaria vulgaris, Ser. ii. 
247 

Lagerstreemia Flos-reginz, 
Retz, ii. 228 

Lagunea lobata, Willd. i. 155 

Lasianthera apicalis, Zw. 1. 
260 

Lasianthus dubzus, Wight, ii. 


35S ae 
Gardneri, Hk. f. 366 
Moonii, Wzght, 364 
obliquus, 7w. 367 
oliganthus, Zw. 366 
protractus, Thw. 365, 367 
rhinophyllus, Zw. 365 
strigosus, Thw. 365, 366 
strigosus, Wzght, 367 
Thwaitesii, Hz. 7. 365 
varians, Tw. 368 
Walkerianus, Thw. 367 
Walkerianus, Wight, 365 
Lavallea zeylanica, Baill, i. 
257 : 
Lawsonia alba, Lam. ii. 228 
znermts, 1. 228 
spinosa, L. 228 
Leea Ozzllis, DC. i. 297 
sambucina, W2d/d. 297 
Staphylea, Roxb. 297 
LEGUMINOS4A, ii. 4 
Lespedeza indica, Spreng. ii. 32 
Leucocodon reticulatum, 
Gardn. ii. 325 
Leucena glauca, Benth. ii. 122 
Limacia cuspidata, Hz. f. and 


Th. i. 42 
Limonia aczdissima, Auct. i, 
223 


acidissima, L, 228 
alata, W. and A. 223 
arborea, Roxb, 217 


LimoniaCzzzamomum, Moon, 
i. 223 
citrifolta, Moon, 227 
crenulata, Roxb. 223 
misstonts, Wall. 227 
monophylla, 1. 227 
pentaphylla, Roxb. 217 
pusilla, Gaertn. 71 
LINACEZ, i. 188 
Linum mysorense, Heyne, i. 
188 
Liriodendrum lilitfera, Moon, 
nh i 
Ludvigia opposztzfolza, L. ii. 
234 
parviflora, Roxb, 234 
prostrata, Roxb. 234 
Luffa acutangula, Po-xd. ii. 252 
zegyptiaca, AZi//. 251 
cylindrica, Roem. 251 
pentandra, Roxb. 251 
Lumnitzera racemosa, W2d/d. 
li, 162 
Luvunga eleutherandra, Dads. 
i, 224 
scandens, Thw. 224 
LYTHRACE#, ii. 222 
Lythrum Pemphis, 1. f. ii. 227 


Meerua arenaria, Hk. f. and 
Th. i. 58 
oblongifolia, Thw. 58 
MAGNOLIACE&, i. 14 
Mallea Rothit, A. Juss. i. 245 
MALPIGHIACE, i. 192 
Malva tomentosa, L. 1. 143 
MALVACE4&, i. 140 
Malvastrum  tricuspidatum, 
A. Gray, i. 140 
Mangifera zzdzca, L. i. 318 
indica, Thw. 317 
zeylanica, HR. f. 317 
Mangostana Cambogia, 
Gaertn. i. 95 
Morella, Gaertn. 96 
Mappia Championiana, Miers, 
1. 263 
fetida, Thw. 262 
Gardneriana, Miers, 262 
ovata, Mzers, 262 
Marumia zeylanica, Bl. ii. 208 
Mastixia arborea, Clarke, ii. 
287 
tetrandra, Clarke, 287 
Medinilla fuchsioides, Gardz. 
li. 208 
maculata, Gardn. 209 
Meesta serrata, Gaertn. i. 235 
Melastoma asperum, L. ii. 195 
buxtfolta, Moon, 197 
ellipticum, Naud. 200 
Malabathricum, Z. 199 
octandra, L.. 199 
repens, Moon, 198 
Royenzz, Bl. 200 
MELASTOMACEAZ,, ii. 192 


Melia Azadirachta, L. i. 


244 
Aszedarach, L. 244 
Azedarach, Moon, 243 
compostta, Willd. 243 
dubia, Cav. 243 
parviflora, Moon, 245 
pumila, Moon, 242 
MELIACEA, i. 241 
Melilotus parviflora, Desf. ii. 
ar 
Meliosma Arnottiana, Walp. i. 
315 
pungens, Thw. 314 
simplicifolia, Walp. 315 
Wight, Planch. 314 
Melochia concatenata, L. i. 
170 
corchorifolia, Z. 170 
pyramidata, 1. 170 
Melothria deltotdea, Thw. ii. 
257 
perpusilla, Cogn. 256 
zeylanica, Clarke, 257 
Memecylon angustifolium, 
Wight, ii. 219 
Arnottiana, Wzght, 211 
capitellatum, Z. 222 
Clarkeanum, Cogn. 217 
cuneatum, 7hw. 216 
depressum, Benth. il. 212 
discolor, Cogn. 218 
edule, Roxb. 217, 220, 221, 
222 
elegantulum, Tw. 214 
ellipticum, Zw. 214 
fuscescens, Thw. 218 
Gardneri, Tw. 211 
grande, Retz. 221 
Hookeri, 7hw. 212 
jambosoides, Thw. 217, 218 
leevigatum, BZ. 220 
leucanthum, Zw. 220 
macrocarpum, Zw. 220 
macrophyllum, Tw. 215 
orbiculare, Zw. 216 
ovoideum, Cogn. 220 
ovoideum, Zhw. 215 
parvifolium, Zw. 213 
phyllanthifolium, Zw. 219 
procerum, Zw. 216 
ramifiorum, Lam. 216 
revolutum, Zhw. 215 
rhinophyllum, Tw. 219 
rostratum, Zw. 218 
rotundatum, Cogn. 214 
Royenit, Bl. 222 
rubro-ceruleum, Thw. 217 
timctorium, Koen. 222 
umbellatum, Burm. 7. 216 
urceolatum, Cogn. 217 
varians, Thw. 213 
Wightii, 7zw. 212 
MENISPERMACEZ, i. 37 
Menispermum fenestratum, 
Gaertn, 1. 41 


Index to Botanical Names. 


Mephitidia Gardneri, Thw. ii. 
66 


3 
Moonzi, Thw. 364 
oligantha, Thw. 366, 367 
protracta, Thw. 367 
rhinophylla, Thw. 365 
strigosa, Thw. 367 
tomentosa, Thw. 365 
varians, Thw. 368 
Walkeriana, Wight, 365 
Mesua coromandelina, Thw. i. 
106 
ferrea, Z. 105 
Nagaha, Gardn. 105 
pulchella, Pl. and Tri. 106 
salicina, Pl. and Tri. 105 
sclercephylla, Thw. 106 
speciosa, Thw. 106 
Thwaitesii, PZ, and Trz. 106 
Walkeriana, Pl. and Tri. 
105 
Mezoneurum enneaphyllum, 
W. and A. ii. tor 
Michelia Champaca, L. i. 15 
glauca, Wight, 14 
nilagirica, Zenk. 14 
ovalifolia, Wight, 15 
Walkeriz, Wight, 14 
Microcos paniculata, L. 1. 177 
Micromelum pubescens, BZ/, i. 
218 
Microtropis ramiflora, Wight, 
i. 269 
Wallichiana, Wigh7, 269 
Miliusa indica, Lesch. 34 
montana, Gardn. 34 
zeylanica, Gardn. 35 
Millingtonia Arnottiana, 
Wight, i. 315 
Milnea apiocarpa, Thw. i. 245 
Roxburghiana, W. and A. 


246 
Mimosa bigemina, L. ii. 132 
cinerea, L, 121 
cesia, L. 127 
LEntada, L. 119 
odoratissima, 1. 129 
pennata, L. 127 
pudica, Li. 122 
umbellata, Vahl, 132 
virgata, L. 122 
Mitrephora Heyneana, 7hw. 
1s2 
Moacurra gelonioides, Roxb. 
i. 254 
Modecca palmata, Lam. ii. 241 
tuberosa, Roxb, 241 
Wightiana, Wall, 240 
Mollugo Cerviana, Ser. ii. 272 
disticha, Ser. 272 
hirta, Thunb. 270 
nudicaulis, Lam. 272 
oppositifolia, Z. 271 
pentaphylla, Z. 271 
Spergula, L, 271 
stricta, Li, 271 


Momordica Charantia, Z. ii 


248 
denudata, Clarke, 249 
dioica, Roxb. 249 
seylanica, Mill, 249 
Monocera glandulifera, Hk. i. 


167 
Wealkeriz, Wight, 187 
seylanica, Arn. 187 
Monoporandra cordifolia, 
Thw. i. 137 
elegans, Thw. 138 
lancifolia, Thw. 136 
More, Gaertn. i. 309 
Morinda érvacteata, Roxb. ii. 


_ 354 
citrifolia, Z. 354 
exserta, Roxb. 354 
scandens, Roxb. 355 
tinctoria, Roxé. 354 
umbellata, Z. 355 
Moringa pterygospernia, 
Gaertn. i. 327 
zeylanica, Pers. 327 
Mucuna atropurpurea, DC, ii. 
61 
gigantea, DC. 62 
monosperma, DC. 61 
pruriens, DC. 62 
prurita, Hk. 62 
Mukia leiosperma, Wight, ii. 


255 
scabrella, Arn. 254 
Mundulea suberosa, Benth. ii. 


29 
Munronia pumila, Wighi, i. 
242 
Murraya exotica, Z. i. 219 
Gleniei, Tw. 220 
Keenigii, Spreng. 220 
Musszenda corymbosa, Roxb. 
li. 324 
frondosa, Z. 323 
Myriophyllum indicum, 
Willd, ii. 149 
Myrobalanus bellerica,Gaertn. 
il. 159 
MYRTACEZ&, ii. 165 
Myrtus androsemoides, L, ii. 
176 
brasiliana, L. 188 
caryophyllata, L.. 174 
Cumini, L. 179 
tomentosa, Ait. 166 
zeylanica, L. 171 


Naravelia zeylanica, DC. i. 2 
Nargedia macrocarpa, Bedd. 
ii. 
Nasturtium indicum, DC, i. 52 
Oficinale, Br. 53 
Nauclea coadunata, Roxb. ii, 
292 
cordata, Roxb, 292 
cordifolia, Moon, 293 
macrophylla, Moon, 292 


383 


Nauclea orzentalis, L. 292 
orzentalis, Moon, 293 
parvifolia, Moon, 295 
parvifolia, Roxb. 294 
tubulosa, Arn. 295 
zeylanica, Hk. f. 296 

Nelttris Jamboselta, Gaertn. 

ii. 338 
Nelumbium speciosum, 
Wetld. i. 51 
Nephelium d2zfolatum, Thw. i. 
397 
erectum, Thw. 308 
eximtum, Thw. 310 
fuscatum, Thw. 305 
Gardneri, Zw. 309 
Longana, Camb. 309 
pupillum, Wight, 309 
stmplicifolium, Thw. 308 
Neptunia oleracea, Lour. ii. 
118 
Plena, Benth. 119 
Nesea trifora, Kunth, ii. 227 
Neurocalyx capztata, Benth. 
li. 300 
Championii, Benth. 300 
Gardneri, Zw. 300 
Wightii, Arz. 299 
zeylanicus, Hk. 299 
Nomismia nummularia, W. 
and A. ii. 82 
Nothopegia Colebrookiana, 
Bi. ii. 325 

Nymphza ae L. i. 49 
Nelumbo, L 
stellata, Willd. 50 

NYMPHAACES, i. 49 


Ochna Zuczda, Lam. i. 233 
Moonii, Thw. 234 
nitida, "Thunb. 234 
rufescens, 
squarrosa, L. 233 
Wealkeriz, Planch. 233 
Wightiana, Wall. 233 
seylanica, Lam, 235 
OCHNACEZ, i. 232 
Odina Wodier, Roxd. i. 318 
Gnothera fruticosa, L. ii. 235 
OLACINE#, i. 254 
Olax psittacorum, Vahl, i. 256 
scandens, foxb. 256 
Wightiana, Wad/. 257 
zeylanica, L. 257 
Oldenlandia biflora, Z. ii. 317 
biflora, Moon, 314. 
corymbosa, Z. 314 
crystallina, Roxb. 315 
diffusa, Roxb. 315 
herbacea, Roxb, 315 
Stricta, ZL. 316 
trinervia, /ee/z. 316 
umbellata, Z. 316 
Omphalobium indicum, 
Gaertn. ii. 2 
ONAGRACE4Z:, ii. 232 


384 


Ophiorrhiza angustifolia, 
Thw, il. 321 
decumbens, Gardn. 321 
glechomifolia, Tw. 323 
Harrisiana, Heyne, 321 
Mungos, Z. 320 
nemorosa, Thw. 321 
pallida, Tzw. 322 
pectinata, Arz. 322 
radicans, Gardz. 321 
Opilia amentacea, Hoxd. i. 258 
Opuntia Dillenit, Haw. ii. 267 
Ormocarpum sennoides, DC. 


li. 39 
Oruztrophe allophylus, Pers. i. 
302 
Cobbe, Willd. 303 
serrata, Roxb. 303 
Orophea coriacea, Thw. i. 34 
fleyneana, Hk. f. and Th. 32 
obligua, Ak. f. and Th. 33 
zeylanica, Hk. f. and Th. 35 
Osbeckia aspera, BZ. ii. 195 
buxifolia, Avz. 197 
cupularis, Don, 195 
elliptica, Naud. 198 
erythrocephala, Vaud. 194 
glauca, Benth. 196 
Leschenaultiana, Thw,. 194 
Moonii, Tw. 198 
octandra, DC. 198 
parvifolia, Arm. 194 
polycephala, Naud. 198 
Rheedii, Gardn. 194 
rubicunda, Arz. 197 
truncata, Arm. 194 
virgata, Don, 198 
Walkeri, Arz. 196 
zeylanica, L. f. 195 
Osmelia Gardneri, 7Aw. ii. 238 
Oxalis corniculata, Z. i. 196 
nuda, Arn. 198 
prolifera, Arn. 199 
sensitiva, L. 197 
violacea, L. 197 


Pachycentria Walkerz,Thw. ii. 
200 
Pachygone adversa, Miers, i. 45 
ovata, Mzers, 45 
Plukenetiz, Miers, 45 
Palimbia ramosissima, Thw. 
ii. 280 
Panax fruticosum, L. ii. 282 
Paramignyaarmata, O/zv. i.225 
monophylla, Wzghz, 224 
Paratropia terebinthacea, Arn. 
ii. 283 
Paritium tiliaceum, Juss. 1.157 
Parkinsonia aculeata, 1. ii. 102 
Parochetus communis, Ham. 
il. 20 
Passifiora edulis, Sims, ii, 242 
fetida lL. 242 
stipulata, Aubl. 242 
suberosa, Li. 241 


Passifiora Walkerie, Wight, 
il, 241 
PASSIFLORACES, il. 239 
Pavonia glechomifolia, A.Rzch. 
i. 148 
odorata, W2l/d. 149 
zeylanica, Cav. 149 
Patonzia parvifolia, Wight, i.28 
Paullinia aszatica, L. 1. 215 
Pavetta angustifolia, Zw. ii. 
359 
Gleniei, Tw. 351 
hispidula, W, and A. 350 
indica, ZL. 349 
involucrata, Tw. 351 
tomentosa, Thw. 351 
Pectinea zeylanica, Gaertn. i. 72 
Peltophorum ferrugineum, 
Benth. ii. tot 
Pemphis acidula, /orst. ii. 227 
Pentaloba zeylanica, Arn. ii. 68 
Pentapanax Leschenautltiz, 
Seem. ii: 282 
Pentapetes phcenicea, Z. i. 169 
subertfolta, LL. 169 
Periandra Bertertana, Benth. 
th, FAS 
Pericopsis Mooniana, 7hw. ii. 


+ SY 
Pestalozzia laxa, Thw. ii. 260 
Peucedanum zeylanicum, 
Gardn. ii. 280 
Phantis, Linn. i. 227 
Pharnaceum distichum, L, ii. 
272 
Mollugo, L.. 27% 
triflorum, Moon, 272 
Phaseolus aconitifolius, Jacq. 
We GAs 
adenanthus, Mey. 70 
ealcaratus, Roxb. 73 
Grahamianus, W. axd A.7o 
Max, Z. 72 
Mungo, . 72 
radiatus, L. 72 
semierectus, ZL. 70 
trilobus, Az¢. 71 
trinervius, Heyze, 72 
truxtllensis, H. B. K. 70 
Phoberos Gaertnerz, Thw. i. 71 
flookerianus, Wight, 71 
Photinia Notoniana, W. and 
A. ii. 142 
Phyllodizm pulchellum, Desv. 
li. 48 
Piliostigmaracemosum, Benth. 
ii, 116 _ 
Pimpinella Heyneana, Wadd. 
li, 279 
Leschenaultii, DC. 279 
Pithecolobium bigeminum, 
Benth, ii. 132 
dulce, Benth. 131 
geminatum, Bezth. 131 
Saman, Benth. 132 
subcoriaceum, Tw. 133 


I[udex to Botanical Names. 


Pithecoleobium umbellatum, 
Benth. ii. 132. 
PITTOSPORACEA, i. 77 
Pittosporum neelgherrense, 
W., and A. i. 78 
beeen W. and A. 


reneaicann Wight, 78 
Pityranthe verrucosa, Tw. i. 
172 
Platea axillaris, Thw. i. 261 
cortacea, Thw. 261 
Plectronia macrocarpa, Bedd. 
li. 345 ; 
Pleurostylia Wightii, W. axd 
A. 1. 271 
Polanista viscosa, Thw. i. 57 
Polyalthia acuminata, Tw, i. 


25 
coffceoides, B. and Hk, f. 24. 
Korinti, B. and Hk. f. 25 
longifolia, B. and Hk. f. 24 
Moonii, Tw. 26 
persicifolia, B. and HR. f. 26 
suberosa, B. and HR. f. 25 
Polycarpoea corymbosa, Zam. 
i, 88 
Polycarpon Leeflingize, B, and 
Hb. f. i. 87 
Polygala arillata, Ham. i. 79 
arvensis, Willd. 80, 81 
chinensis, Z. 81 
czlzata, L. 83 
elongata, Klein, 81 
glaucoides, Z. 80 
Llomeratu, Lour. 82 
hirsutula, Arn. 8i 
javana, DC. 80 
leptalea, DC. 80 
rosmarinifolia, W. and A. 82 
sibirica, Z. 82 
telephioides, Wzl/d. 82 
Thea, Burm. f. 68 
triflora, 1. 81 
POLYGALACE&, i. 78 
Polyodontia Walkeriz, Wight, 
li. 135 
zeylanica, Wight, 134 
Polyscias acuminata, Seem. ii. 
282 
Pometia eximia, //Z. f. i. 310 
Pongamia glabra, Vezz. ii. 91 
Portulaca oleracea, LZ. i. 89 
quadrifida, Z. 90 
suffruticosa, Wzght, 90 
tuberosa, Roxb. 90 
Wightiana, WadZ, 89 
PORTULACACE, i. 88 
Potentilla Kleiniana, W. and 
A, il. 139 
Mooniana, W2zght, 139 
Poterium indicum, Gavdz. ii. 
I4I 
Phar albidiflora, 
Thw. ii. 355 
Fergusonii, Thw. 356 


Index to Botanical Names. 


Protium caudatum,\W.and A. | Rhamnus /izjuda, L. 280 


i. 236 

Pseudarthria viscida, W. and 
A, ii. 41 

Pseudocarapa Championii, 
Hlemsi. i. 248 

Psidium Guyava, L. ii. 167 

pumilum, Vahl, 167 
Psoralea corylifolia, Z. ii. 28 


Psychotria aszatica, Moon, ii. 


362 
bisulcata, W. and A. 362 
elongata, Hz. f. 359 
filipes, Hk. f. 361 
Gardneri, Hz. f. 358 
glandulifera, 7hw. 357 
herbacea, L. 363 
longepetiolata, Thw. 361 
Moonii, 2. 7. 360 
plurivenia, Zw. 361 
sarmentosa, B/. 359 
scandens, Moon, 359 
sordida, Tw. 360 
stenophylla, Hz. f. 357 
Thwaitesii, Zz. 7. 358 
vaginans, DC. 362 
Wightiana, 2. f. 358 
Psydrax dicoccos, Gaertn. ii. 


343 

Pteridophyllum decipiens, 
Thw. i. 240 

Pterocarpus 42/odzs, Roxb. ii. 


go 
Marsupium, Roxb. 90 

Pteroloma triquetrum, Benth, 
ll. 49 

Pterospermum  suberifolium, 
Lam, i. 169 

Pterygota alata, Thw. i. 166 

Pycnospora hedysaroides, Br. 


ii. 41 
Pygeum Wadkeriz, Bl. ii. 135 
Wightianum, B/. 134 
zeylanicum, Gaertn. 135 
Pyrenacantha volubilis, Hook. 
i. 263 


Randia dumetorum, Lam. ii. 


330 
Gardneri, 2. f. 331 
malabarica, Lam. 331 
rugulosa, Hz. f. 331 
uliginosa, DC. 330 
RANUNCULACEZ i. 1 
Ranunculus, sastatus, Walk. i. 


4 
pinnatus, Arn. 4 
ittifolius, 72. 4 
allichianus, W. and A. 4 
zeylanicus, Moon, 4 
Reichenheimia Thwaitesii, K1, 
ii. 264 
RHAMNACE#, i, 278 
Rhamnus Arnottianus, Gaertn, 
i. 283 
circumscissus, Li. f, 284 
PART II, 


Napeca, L. 281 

Gnoplia, L. 280 

Wightii, W. and A. 283 
Rhipsalis Cassytha, Gaertz. ii. 


266 
Rhizophora Candelaria, DC. ii. 


ISI 
Candel, Moon, 151 
conjugata, Arn. 151 
conjugata, L. 153 
gymnorhiza, L. 153 
mangle, Moon, 151 
mucronata, Z. 151 

RHIZOPHORACE#, ii. 150 

Rhodomyrtus tomentosa, 
Wight, ii. 166 

Rhus Cobbe, L. i. 303 

decipiens, Wight, 240 

Rhynchocarpa rostrata, Vaud, 
ii. 258 

Rhynchosia acutissima, Tw, 


li. 85 
aurea, DC. 83 
cana, DC. 83 


cyanosperma, Benth. 14 
densiflora, DC. 85 
minima, DC. 84 
nummularia, DC. 82 
rufescens, DC. 82 
suaveolens, DC. 83 
villosula, Thw. 85 
viscosa, DC. 85 
Rissoa zeylanica, Arn. i. 227 
Rondeletia asiatica, L, ii. 328 
ROSACE&, ii. 134 
Rotala verticillaris, L. ii. 224 
Rothia trifoliata, Pers. ii. 7 
Roumea hebecarpa, Gardn. 1.74 
Rourea santaloides, W.andA. 


ii. I 

Rubus ellipticus, Sz. ii. 137 
flavus, Ham. 137 
glomeratus, B/, 136 
Gowreephul, Roxb, 137 
lasiocarpus, Sm, 138 
leucocarpus, Arn. 138 
micropetalus, Gardn. 136 
moluccanus, ZL. 136 
parvifolius, Moon, 138 
rugosus, Sm, 136 

Rubia cordifolia, Z. ii. 372 
secunda, Moon, 372 

RUBIACE#, ii. 289 

RUTACEZ, i. 213 


SABIACE#, i. 314 
Sagerea Thwaitesit, 
and Th. i. 33 
Sageretia afinis, Thw. i, 284 

costata, Wig. 284 
Sagina procumbens, 1. i. 85 
Salacia diandra, Thw. i. 277 
oblonga, Wadl, 277 
prinoides, DC. 276 
reticulata, Wight, 277 


Hike i 


385 


Salacia zerminalis, Thw. 275 
Salomonia cordata, Avz. i. 83 
ciliata, DC. 83 
oblongifolia, DC. 83 
Samadera indica, Gaertz. i. 
231 
Samara leta, L. ii. 216 
SAMYDACEA, ii. 236 
Sanicula europza, Z. ii, 276 
SAPINDACE2, i. 298 
Sapindus bifoliatus, Azerz, i. 


397 
dejficiens, Wight, 302 
emarginatus, Vak/, 307 
erectus, Azerz, 308 
Glenzez, Thw. 305 
laurifolius, Vah/, 306 
tetraphylla, Vahl, 301 
Thwaitesii, Hzern, 308 
trifoliatus, L. 305 
uniugus, Thw. 305 
Saprosma indicum, Dad/z. ii. 
368 
scabridum, Bedd. 369 
zeylanicum, Bedd. 369 
Saraca indica, Z. li. 114 
Sarcocephalus cordatus, J/zq. 
li, 292 
Sarissus anceps, Gaertn. ii. 370 
Sarosanthera lasiopetala, Thw. 
i, 108 
SAXIFRAGACEZ, ii. 143 
Schizostigma hirsutum, Avrz. 


ii, 327 

Schleichera trijuga, W7d/d. i. 
304 

Schmidelia allophylla, DC. i. 


302 
Cobbe, DC. 303 
hispida, Thw. 303 
villosa, Wight, 304 
Schrebera albens, Retz. i. 271 
Schumacheria alnifolia, 2. 7. 
and Th, i. 10 
angustifolia, Hk. f. and Th. 
Il 


castanezefolia, VahZ, to 
Sclerostylis Arnottiana, 
Wight, i. 227 
zeylanica, Wight, 227 
Scolopia acuminata, C/os, i. 70 
Arnottianus, Thw. 71 
chinensis, Clos, 71 
crassipes, Clos, 71 
Gaertneri, 7hw. 71 
pusilla, Willd. 7% 
Scopolia aculeata, Sm. i. 215 
Scutia indica, Brongn. i. 284 
Scutinanthe brunnea, Thw. i. 


238 
Scyphellandra virgata, Thw. 


i, 69 
Scyphiphora hydrophylacea, 
Gaert. f. il. 337 
Scyphostachys coffoeoides, 
Thw. ii. 335 
CC 


386 


Scyphostachys pedunculatus, | Smithia gaxzculata, Arn. 37 


Thw. ii. 335 


sensitiva, Azt, 37 


Semecarpus acuminata, Zw. | Sonerila afinzs, Arn. ii. 202 


rome) 
coriacea, Tw. 321 
cuneata, Engl. 324 
Gardneri, Tw. 322 
leevigata, Thw. 325 
marginata, 7hw. 319 
Moon, Thw. 321 
nigro-viridis, Tw. 323 
oblongifolia, Thw. 323, 324 
obovata, AZoon, 321 
obscura, Thw. 320, 324 
parvifolia, Tw. 324 
pubescens, Zw. 320 
subpeltata, Tw. 320 
Thwaitesiz, Ak. f. 321 
Walkeri, Hz. f. 322 
Serissa Gardneri, Thw. ii. 368 
scabrida, Thw. 369 
zeylanica, Thw. 369 
Serpicula Jreviges, W. and A. 
ii. 148 
hirsuta, W/. and A. 148 
zndica, Thw. 147, 148 
zeylanica, Arn. 147 
Sesbania aculeata, Pers. il. 34 
zegyptiaca, Pers. 34 
grandifiora, Pers. 35 
Sesuvium pedunculatum, 
Pers. ii. 268 
Portulacastrum, Z. 268 
repens, Rottl. 268 
Sethia acuminata, Arn. i. 19% 
indica, DC. 190 
lanceolata, Arn. 191 
obtusifolia, Thw. 192 
Shorea dvevipetiolaris, Thw. i. 
r18 
Dyerii, Tw. 117 
lissophylla, Tw. 117 
oblongifolia, Tw. 116 
reticulata, Thw. 117 
stipularis, Tzw. 118 
Shuteria vestita, W. and A. 
ii. 58 
Sicyos Garcini, Burm. f. ii. 260 
periplocifolia, L. 147 
Sida acuta, Burm. i. 142 
alba, L. 142 
alnifolia, L. 142 
astatica, L. 144 
carpinifolia, L. f. 142 
cordifolia, Z. 143 
humilis, Cav. 141 
mysorensis, W. and A, 142 
persica, Brum. 145 
vetusa, Li. 143 
rhombifolia, £Z. 143 
spinosa, Z. 142 
SIMARUBACE, 1, 229 
Szum lobatum, Moon, ii. 280 
triternatum, Moon, 279 
Smithia blanda, Wad/, ii. 37 
conferta, S72. 37 


Arnottiana, Thw. 204 
angustata, Triana, 203 
Brunonis, W. and A. 203 
cordifolia, Cogn. 202 
firma, Thw. 205 
Gardneri, Thw. 205 
glaberrima, Arn. 203 
Guneratnez, Trim. 207 
Harveyz, Thw. 206 
hirsutula, Arn. 205 
Flookeriana, Arn. 204 
lanceolata, Thw. 206 
linearis, Hz. f. 207 
pedunculosa, 7/zw. 208 
pilosula, Thzw. 207 
pumila, Thw. 202 
thombifolia, Zzw.203 
robusta, Av. 206 
rostrata, Thw. 203 
tomentella, Thw. 204 
Wightiana, Arz. 204 
zeylanica, W. and A. 202 
Sonneratia acida, L. f. ii. 230 
alba, Smzth, 230 
apetala, Ham, 229 
Sophora heptaphylla, Arn. ii. 
6 


9 
heptaphylla, L. 94 
tomentosa, Z. 95 
violacea, Thw. 95 
zeylanica, T77m. 96. 
Soya Wighti, Grah. il. 59 
Soymida “jebrifuga, A. Juss. i. 


251 
Spatholobus Roxburghitz, 
Benth. ii. 65 
Spergula arvensis, L i. 85 
Spermacoce hispida, Z. ii. 371 
ocymoides, Burm. f. ii. 371 
stricta, Z. f. 371 


Spondias Mangifera, W/zd/d. 
1. 327 

Stalagmitis cambogioides, 
Murr. i. 96 


Stellaria drymarioides, Thw. 


1. 25 
media, W2th. 85 
Stemonoporus acuminatus, 
Bedd. i. 133 
affinis, Tw. 134 
canaliculatus, Tw. 135 
Gardneri, Thw. 133 
lanceolatus, Thw. 134 
Moonii, Tw. 137 
nervosus, Zw. 136 
nitidus, Tw. 136 
oblongifolius, Tw. 135 
petiolaris, Tw. 135 
reticulatus, Tw. 136 
rigidus, Tw. 134 
Wightii, Zw. 132 
Stemonurus apicalis, Thw. 1. 
260 


Index to Botanical Names. 


Stemonurus Heyneanus, Miers, 
i, 261 
polymorpha, Wight, 265 
Walkeri, Miers, 261 
seylanicus, Miers, 261 
Stephania hernandifolia, 
Waép, i. 45 
tntertexta, Miers, 45 
Stephegyne parvifolia, Auth. 
ii. 294 
tubulosa, Hz. f. 295 
Sterculia Balanghas, L. i. 165 
colorata, Roxb. 166 
foetida, Z. 164 
guttata, Roxb. 165 
Thwaitesii, AZas¢. 166 
urens, Roxb. 164 
STERCULIACEA, i. 163 
Stizolobium giganteum, Pers. 
li. 62 
pruriens, Pers. 62 
rugosum, Moon, 65 
Stravadia integrifolia, Moon, 
ii. I9g0 
“rubra, Moon, 189 
Stravadium obtusangulum, 
Bl. ii. 19 
Streptostigma viridiflorum, 
Thw. i. 311 
Strombosia javanica, Thw. i. 
257 
zeylanica, Gardn. 257 
Strongylocalyx CLLDEE ES, 
BI. ii. 170 
Strongylodon ruber, Vog. ii. 65 
Oe elliptica, Thw. ii. 


Wie, A. Rich. 328 
Stylosanthes mucronata, 


Willd. ii. 36 
Sunaptea disticha, Zvzm,. i. 
127 


scabriuscula, 772m. 126 
Suriana maritima, Z. i. 232 
Sweetenia Chloroxylon, Roxb. 

1. 253 
Syzygium assimile, Thw. ii. 
I76 

calophyllifolium, Thw. 173 

caryophylleum, Gaertn. 174 

cordifolium, Thw. 176 

jirmum, Thw. 170 

Gardnerzi, Thw. 174. 

Jambolanum, DC. 179 

lissophyllum, Thw. 173 

micranthum, Thw. 175 

montanum, Thw. 170 

Neestanum, Arn. 177 

nervosum, DC. 179 

oliganthum, Thw. 178 

revolutum, Thw. 175 

rotundtfolium, Arm. 177 

sclerophylium, Thw. 178 

spathulatum, Thw. 178 

syluestre, Thw. 175 

umbrosum, Thw. 173 


Index to Botanical Names. 


387 


Tamarindus indica, L. ii. 114 | Tribulus /anuginosus, L.i. 194 | Vateria acuminata, Hayne, i. 


TAMARISCINES, i. ot 


terrestris, Z. 194 


Tamarix erzcozdes, Rottb. i. gt | Trichadenia zeylanica, Thw. i. 


gallica, Z. 91 
éndica, Willd. 91 


Tarenna szeylanica, Gaertn. ii. 


328 


Teramnus labialis, Spreng. ii. 


60 
mollis, Benth. 60 
Tephrosia #zrta, Thw. ii. 32 


Hookeriana, W. azd A. 32 


maxima, fers. 32 
purpurea, Pers. 31 
senticosa, Pers. 30 
spinosa, Pers. 30 
suberosa, DC. 29 
tinctoria, Pers. 31 
villosa, Pers. 33 


Terminalia a/ata, Moon, ii. 160 


belerica, Roxb. 159 
chebula, Retz. 159 
glabra, W. and A. 160 
parviflora, Tw. 160 
tomentosa, W. and A. 161 


seylanica, Van H. and Mull.- 


Arg. 160 

Ternstroemia emarginata, 
Choits, i. 108 

japonica, Thunb. 107 

Terpnophyllum zeylanicum, 
Thw. i. 97 

Tetracera lzvis, Vahl, i. 6 

Tetracrypta cinnamomoides, 


Gardn. and Champ. ii. 157 
Tetrameles nudiflora, By. ii. 


265 
Tetrastigma lanceolarium, 
Planch. i. 296 


Thalictrum glyptocarpum, W. 


and A. i. 3 
javanicum, B7/. 3 


Thespesia Lampas, Dalz. and 


Gibs, i. 158 
populnea, SoZ, 158 
TILIACEZ# i. 171 


Tiliacora cuspidiformis, Miers, 


i. 42 
Sraternaria, Miers, 42 
racemosa, Colebr. 42 


Timonius Jambosella, Thw. ii. 


38 


3 
Tinospora cordifolia, W7zers, i. 


39 
crispa, Miers, 39 
malabarica, Miers, 38 
tomentosa, Miers, 39 


Toddalia aculeata, Pers. i. 215 
Toxicodendron Cobbe, Gaertn. 


i, 303 


Trapa bispinosa, Roxb. ii. 235 


ZS 
Trichosanthes azgwzna, L, il. 


245 
bracteata, Voight, 244 
caudata, Willd. 244 
cucumerina, Z. 245 
integrifolia, Tw. 245 
nervifolia, ZL. 244 
palmata, Roxb. 244 
Thwattestz, Cogn. 245 
Trifolium repens, Li. ii. 20 
Trigonella indica, L. ii. 7 
Tristillateta australasica, A. 
Rich. i. 194 


Triumfetta anzgzlata, Lam. i. 


Bartramia, L. 179 
consticua, Trim. 180 
glabra, Rof¢/, 180 
neglecta, W. and A. 181 
pilosa, Roth. 179 
pilosula, Thw. 181 
rhomboidea, /acg. 179 
tomentosa, Boj. 179 


Turpinia zepalensis, Wall.i. 313 


pomifera, DC. 313 


Turrea villosa, Benn, 243 


Ulex europeus, L. i. 7 


UMBELLIFERA, li. 274 
Uncaria dasyoneura, Korth. ii. 


296 
Gambier, Thw. 296 


Unona discolor, Vahl, i. 23 


elegans, Thw. 23 
tripetaloidea, Moon, 28 
uncinata, Dun. 21 


zeylanica, Hk. f. and Th, 23 
Urandra apicalis, Thw. i. 260 
Uraria crinita, Desv. ii. 42 


hamosa, Wail. 43 
picta, Desv. 42 


Urena lobata, Z. i. 147 


sinuata, Z. 148 


Urophyllum ellipticum, Thw. 


1. 326 
zeylanicum, 7hw, 326 


Uvaria macrophylla, Roxd. 1.18 


macropoda, //z. f. and Th. 


19 
Narum, Wadi. 19 
purpurea, B/. 18 


semecarpifolia, /7k. f. and 


Th. 19 
sphenocarpa, 7k. 7, and Th. 
18 


zeylanica, L, 20 


Trianthema crystallina, Thw. | Vahlia oldenlandioides, Roxd. 


ii. 
decandra, L. 270 
monogyna, L. 


triquetra, Kottl. and W. 269 


ii. 143 
viscosa, Roxb. 143 
Vareca zeylanica, Gaertn, ii. 


237 


131 
acuminata, Thw. 133 
afinis, Thw. 134 
canaliculata, ‘Vhw. 135 
cordifolia, Thw, 137 
disticha, ‘Thw. 127 
elegans, Thw. 138 
Gardneri, Thw. 133 
indica, Bl. 131 
indica, L. 132 
jucunda, Thw. 133 
lanceolata, Thw. 134. 
lancifolia, Thw. 136 
Mooniz, Thw. 137 
nitida, Thw. 136 
oblongifolia, Thw. 135 
petiolaris, Thw. 135 
reticulata, Thw. 136 
rigida, Thw. 134 
scabriuscula, Thw. 126 
seylanica, Wight, 133 
Vatica affinis, 7hw. i. 128 
cordifolia, Thw. 126 
disticha, A. DC. 127 
obscura, 777m. 129 
Roxburghiana, &Z. 128 
scabriuscula, A. DC. 126 
Thwattesiz, A. DC. 134 
Ventilago maderaspatana, 
Gaertn. 1. 279 
Viburnum coriaceum, #4. ii. 
288 
erubescens, Wad/. 289 
hebanthum, Thw. 288 
zeylanicum, Gardn. 288 
Vigna Catiang, Endl. ii. 74 
luteola, Benth. 73 
sznensis, Endl. 74. 
vexillata, Benth. 74 
Viola crenata, Moon, 67 
distans, Wadd. 66 
enneasperma, Li, 67 
hastata, Moon, 66 
Patrinii, DC. 66 
serpens, Wall. 67 
suffruticosa, lL. 67 
Walkeriz, Wight, 66 
Wightiana, Thw. 66, 67 
VIOLACE#, i. 65 
Vitis acuminata, 772m. i. 292 
adnata, Wad/. 290 
earnosa, Wad. 294 
Gardneri, Laws. 293 
glyptocarpa, Laws, 289 
Heyneana, Wad/. 292 
indica, LZ. 288 
lanceolaria, Vad/, 296 
Linnezei, Wall. 29% 
lonchiphylla, Laws. 290 
pallida, Trim, 292 
pallida, W. and A. 29% 
pedata, Vahl, 295 
quadrangularis, Wa//. 289 
repanda, W. and A, 292 
reticulata, Laws, 294 


388 L[udex to Sinhalese Names. 


Vitis Rheedii, W. and A. | Woodfordia floribunda, Sad. | Zanthoxylum zeylanicum,DC. 


293 ii. 226 i. 214 
setosa, Wall. 206 Wormia hamata, Vahl, i. 139 | Zehneria Garcinz, Stocks, ii. 
tenuifolia, W. and A. 295 retusa, Hk. f. and Th. 12 250 
tomentosa, Heyze, 288 triquetra, fot¢/. 11. hastata, Wzg. 256 
trifolia, Li. 294 Hookeriana, Avz. 256 
Vitmannia elliptica, Vahl, i. | Xanthochymus ovalifolius, umbellata, Thw. 256 
231 Roxb. i. 98 Zizyphus Jujuba, Lam. i. 280 
Xanthophyllum flavescens, Linnei, Laws. 285 
Walkeria serrata, Moon, i. Roxb. 1. 84 lucida, Moon, 28r 
235 virens, Roxb. 84 Napeca, W2l/d. 281 
Walsura Gardneri, 7zw. i. 250 | Ximenia americana, W2/d. i. Nummularia, W. and A. 
Piscidia, Roxb. 250 255 280 
Thwattesit, Cas. DC. 250 | Xylocarpus Granatum, Koen. Cnoplia, A@z27. 280 
Waltheria indica, Z. i. 171 iL DEE rugosa, Lam. 282 
Webera cerzfera, Moon, ii. 328 | Xylopia Championii, HZ. 7. xylopyra, W2zl/d. 282 
corymbosa, W2zd/d. 328 and Th. i. 28 Zornia angustifolia, Sm. ti. 35 
lanceolata, Moon, 343 nigricans, Hk. f. and Th. 28| conjugata, Sm. 35 
Weihea zeylanica, Bazl/. ii. parvifolia, Wk. f. and Th. 28| diphylla, Pers. 35 
156 Watkeri, Arn. 35 
Wendlandia ézcuspzdata, W. | Zanonia indica, L. ii. 261 seylanensis, Pers. 35 
and A. ii. 297 Wightiana, Arn. 260 ZYGOPHYLLACES, i. 194 
Notoniana, Wadl?. 297 Zanthoxylum Rhetsa, DC. i. 
Wissadula Leschenaultiana, 215 
Mast. i. 147 tetraspermum, W, aud A. 
periplocifolia, Thw. 146 215 
zeylanica, MZedzk. 146 triphyllum, Juss. 214 


SINHALESE NAMES. 


Aba, i. 54 Bévila, i. 141 Diwul, i. 228 
Achariya-pala, ii. 62 Bin-béru, i. 7 Diya-kirindiwel, i. 275 
Agu-karni, ii. 363 Bin-mé, ii. 71 Diya-kudalu, i. 2t2 
Ahu, ii. 354 Bin-kohomba, i. 242 Diya-labu, ii. 247 
Akkapana, li. 145 Bin-siyambala, ii. 110 Diya-midella, ii. 189 
Alu-bo, ii. 175 Bodi, ii. 28 Diya-mitta, i. 46 
Alu-pila, ii. 31 Bo-kéra, i. 233, 235 Diya-na, i. 106 
Alu-puhul, ii. 252 Bora-daminiya, i. 177 Diya-nidikumba, ii. £18 
Andara, ii. 121 Bowitiya, ii. 194, 195 Diya-para, i. 11 
Andun-wenna, i. 265 Bu-gétiya, i. 189 Diya-ratambala, ii. 114 
Angana, ii. 338 Bu-hora, i. 114 Diya-ratmal, il. 114. 
Ankenda, 1. 216 Bu-kinda, i. 38 Diya-siyambala, ii. 38 
Anoda, i, 144, 145 Bu-kobbe, i. 303 Diya-taleya, il. 287 
Aralu, ii. 159 Bulu, ii. 159 ~ Diya-wawulétiya, ii. 99 
Aramana, ii. 108 Bulu-mora, i. 310 Dodan-kaha, li. 222 
Aridda, i. 326 Bu-mé, ii. 72 Dodan-pana, i. 217 
Aswenna, il. 44, 372 Bu-pila, ii. 33 Dodan-wenna, li. 22T 
Atu-kétiya, i. 28 Burulla, i. 297 Domba, i. 100 
Buruta, i. 253 Domba-kina, i. 99 
Badulla, i. 321, 322, 324 Bu-tora, ii. To9 Dorana, 1. 115 
Bakmi, ii. 292 Duhudu, i. 272 
Bala, i. 325 Chanchala, ii. 56 Dummella, ii. 245 
Baloliya, ii. 49 Dun, i. 117, 119, 125 
Bambara-wel, ii. 88 Dambu or Damba, ii. 174, 176 
Bata-kirilla, i. 191 Daminiya, i. 175 Ehéla, ii. 103 
Batu-damba, ii. 179 Dan, ii. 174 Ela-gokatu, 1. 98 
Batu-karivila, ii. 248 Dara-vétakolu, ii. 252 Ela-imbul, i. 70 
Béli, i. 229 Dat-kétiya, i. 28; ii. 320 Ela-kadol, ii. 151 
Béli-patta, i. 157 Dawata, ii. 155 Ela-midella, ii. 191 
Beraliya, i. 122 Dawu, ii. 162 Ela-ratmul, ii. 341 
Bériya, ii. 162 Dedi-kaha, ii. 22 Elbedda, 1. 315 


Beru-diyanilla, ii. 233 Dikwenna, i. 172 Embarella, i. 327 


Embul-bakmi, ii. 293 
Ensalu, i. 237 

Epula, i. 179 
Erabadu, ii. 63 
Eta-hirilla, i. 313 
Et-amba, i. 317 
Eta-mura, i. 259 
Eta-werella, i. 312 
Etkara-bembiya, i. 222 
Et-korasa-wel, i. 6 
Et-kukuruman, ii. 330 
Et-olu, i. 49 

Ettériya, i. 219 
Et-tora, ii. 78 

Etuna, i. 167 
Et-undupiyali, ii. 53 


Gal-ambala, ii. 262 
Gal-demata, i. 205 
Galis, ii. 332 
Galkaranda, ii. 115, 343 
Gal-kura, i. 170 
Gal-mendora, li. IIE 
Gal-mora, i. 310 
Gal-siyambala, ii. 112 
Gammalu, ii. go 
Gandapana, i. 262 
Gas-bévila, i. 142 
Gas-gonika, ii. 41 
Gas-kéla, ii. 66 
Gas-kollu, ii. 83 
Gas-nidikumba, i. 197 
Genda-kola, i. 89 
Geta-kaha, ii. 188 
Giriwadi-bévila, i. 142 
Godapara, i. 13 
Goda-hinguru, ii. 127 
Goda-midella, ii. 190 
Goda-wawulétiya, ii. 102 
Gokatu, i. 96 

Golu mora, ii. 135 
Gomma, i. 76 
Gonapana, i. 248, 98 
Gona-wel, i. 139 
Gon- kékiri, ii, 250 
Gopalanga, i ii. 258 
Goradiya, ii. 59 
Goraka, i. 95 
Gurulla, i. 297 
Guru-kina, i. 99 


Hak-ambala, ii. 264 
Hal, i. 131 
Hal-bembiya, i. 245 


Hal-mendora, i. 128, 132 


Hal-milla, i. 173 
Harpilla, ii. 48 
Hampinna, ii, 86 
Hana, ii. 16 
Hanpalanda, ii. 160 
ae 15 
gage “ » 29 
Hik, i. ituade 
Hima, ii. Ra 
Himbutu-wel, i. 277 
Hin-anoda, i, 143 


Index to Stnhalese Names. 


Hin-dan, ii. 174 
Hin-embulembiliya, i. 196 
Hin-épala, i. 148 
Hin-eraminiya, i. 280 
Hin-garadiya, ii. 84 
Hin-gendakola, i. 90 
Hin-gétakola, ii. 371 
Hin-gotukola, ii. 276 
Hingul, i. 249 
Hingutru, ii. 127 
Hin-himbutuwel, i. 276 
Hin-kékiri, ii. 254 
Hin-kina, i. 99 
Hin-kurétiya, ii. 218 
Hin-napiritta, i. 152 
Hin-pala, ii. 271 
Hin-sarana, ii. 269 
Hin-undupiyali, ii. 54 
Hiressa, i. 289 
Honda-beraliya, i. 124 
Hondala, ii. 241 
Hondapara, i. 12 
Hora, i. 114 
Hulanhik, i. 252 
Hulan-mara, i iil. 129 
Hurihi, ii, 129 


Ikiliya, ii. 235 
Imbul, i. 161 
Ipetta, i. 20 
Itta, ii. 283 
Itta-wel, ii. 283 


Jaladara, i. 184 
Jambu, ii. 170 


Kabal-mara, ii. 129 
Kadol, ii. 151 
Kaduruketiya-wel, ii. 163 
Kaha, i. 70 
Kaha-andanahiriya, ii, 15 
Kaha-penela, i. 306 
Kaha-petan, ii. 116 
Kahata, ii, 191 

Kaju, i. 317 

Kakum, i. 282 
Kakkuta-pala, i. 53 
Kalati, i. 25 

Kalatiya, ii. 132 
Kala-wel, ii. 91, 92 
Kalu-badulla, 1. 322 
Kalu-kéra, i. 29 
Kandul-essa, ii. 146 
Kana-goraka, i. 96 
Kankumbala, i. 313 
Kankumbal-kétiya, ii, 135 
Kapukinissa, i. 156 
Kapuru, i. 30 

Kara, il. 346 
Karapincha, i. 220 
Karivila, ii. 248 
Katarodu-wel, ii. 75 
Katu-andara, ii. 125 
Katu-boda, i. 162 
Katu-imbul, i. 160 
Katukenda, i. 70 


Katu-kina, i. 215 
Katukurundu, i. 7 
Katuru-murunga, ii, 35” 
Katu-patuk, ii. 267 
Kawudu-kékin, ii. 256 
Kéhipittan, i. 47 
Kékala, i. 20 

Kékiri, ii. 250 


389 


Kekiri-wara, i, 10; ii, 231 


Keku, i. 34 
Kékuna, i. 239 
Keéliya, i. 177 
Kénawila, i. 166 
Kétambilla, i. 74 
Kétiya, i. 78 
Kevitiya-kera, ii. 2T2 
Kina, i. 101, 104 
Kinihiriya, i. 70 
Kiri-henda, i. 88 
Kiri-kaju, i. 146 
Kiri-kon, i. 246, 250 
Kirilla, ii. 230 
Kirindi-wel, ii. r 
Kiri-wel, ii. 355 
Kobbe, i. 303 
Kobo-mal, ii. 179 
Kohomba, ii. 244 
Kohukirilla, i. 177 
Kokatiya, i. 96, 97 
Kokun, i. 251, 270 
Kola-wakka, i. 279 
Kollu, ii. 77 

Kolon, ii. 293 
Komadu, ii. 253 
Kon, i. 304 
Korasa-wel, i. 5 
Kora-kaha, li. 216 
Kospana, i. 260 
Kottamba, ii. 159 
Kotikan-beraliya, i. 121 
Kotikan-bévila, i. 143 
Kowakka, ii. 247 
Kudalu- dehi, i, 228 
Kudalu-kola, i, 203 
Kudalu-mal, i. 206 
Kudu-miris, i. 215 
Kukulu-pala, i. 87 
Kukuruman, ii. 330 
Kumbalu, i. 230 
Kumbuk, ii. 160 
Kumburu-wel, ii. 98 
Kurétiya, ii. 218 


Lima-dehi, i. 228 
Li-mé, ii. 74 

Liniya, 1 168 

Liyan, i. 258 ; il. 239 
Lunu- ankenda, i, 214. 
Lunukétiya-wel, i. 45 
Lunu-midella, i, 243 
Lunu-warana, i. 59 


Madatiya, ii. 120 
Madol, i. 96 
Magul-karanda, ii. 91 
Maha-andara, ii. 125 


390 Index to Sinhalese Names. : 


Maha-badulla, i. 320 
Maha-bowitiya, li. 199 
Maha-bulumora, i. 238 
Maha-dan, ii. 179 
Maha-debara, i. 280 
Maha-diya-siyambala, ii. 39 
Maha-eraminiya, i. 282 
Maha-gétiya, 1. 189 
Maha-gotukola, ii. 275 
Maha-kiri-wel, il. 355 
Maha-kurétiya, li. 173 
Maha-ratambala, ii. 348 
Maha-sarana, ii. 270 
Maha-tawara, i. 287 
Maha-undupiyali, i. 55 
Maha-wal-kollu, ii. 84 
Makulu, i. 75 

Malitta, ii. 226 
Mal-kéra, i. 233 
Malmora, i. 118, 125 
Manda-madini-wel, li. 372 
Manel, i. 50 

Mara, il. 128 

Maran. ii. 17I 
Maranda, ii. 171 
Masbedde, i. 237 
Mas-mora, ii. T2¢ 
Mayila, ii. 117 
Médiya-wel, i. 295 
Mé-karal, ii. 74 

Mella, i. 257 

Mendora, i. 126, 128 
Méni-damba, ii. 180 
Migon-karapincha, i. 22T 
Mibhiriya, i. III 
Mimini-mara, il. 133 
Mi-wenna, i. 25 
Molpetta, i. 250 

Mora, i. 309 

Mudilla, ii. 189 
Mudu-awara, ii. 68 
Mudu-gétakola, ii. 370 
Mudu-murunga, ii. 95 
Mudu-nelun, i. 251 
Mugunu, ii. 265 

Mun, ii. 72 

Mun-éta, ii. 72 
Murunga, i. 327 
Muruta, ii. 228 
Mussenda, ii. 323 


Na, i. I05 
Naghawalli, ii. 321 
Na-imbul, i. 311 
Na-mendora, i. 126 
Napiritta, i. 152 
Nara-wel, i. I, 2 


Nava, i. 165 
Nédun, ii. 97 
Nelun, i. 51 


Neralu, i. 271 
Netawu, i. 28 
Neya-dasse, i. 109 
Nidikumba, ii. 123 
Nigunu, ii. 265 
Nika-dawulu, 1. 315 


Nil-andana-hiriya, ii. 15 
Nil-awari, ii. 26 
Nil-katarodu, ii. 75 
Nil-mé, ii. 74 
Nil-pitcha, ii. 338 
Niri-wel, i. 42 
Niyan-vétakolu, ii. 251 


Olinda-wel, ii. 57 
Olu, i. 49 
Omara, i. 24 


Palala, i. 84 
Palanga, i. 20 
Palen, i, 274 
Palu-kan, i. 20 
Pamburu, i. 227 
Pana, ii. 156 
Panaka, i. 271 
Pana-karawu, il. 343 
Pandaru, li. 343 
Panukéra, ii. 177 
Patabambara, 1. 279 
Patola, ii. 245 
Patta-épala, 1. 147 
Pattangi, il. 99 
Pawatta, li. 349 
Peddimella, 11. 338 
Pehimbiya, i. 240 
Pena-mihiriya, 1. 108 
Penela, i. 306, 312 
Penela-wel, i. 299 
Peni-tora, il. 105 
Péra, ii. 167 

Petan, ii. 116 
Petika-wel, i. 22 
Péti-tora, il. 106 
Pila, ii. 31 

Pinibaru, ii. 187 
Pini-beraliya, i. 123, 125 
Pita-sudu-pala, ii. 312 
Piyari, i. 271 
Potu-honda, ii. 241 
Pubbériya, i. 258 
Pundalu, i. 311 
Puswel, ii. 119 
Puwak-gediya-wel, 193 


Radaliya, ii. 2 
Ranawara, ii. 106 
Rasa-kinda, i. 39 
Ran-manissa, 1. 57 
Rasamora, i. 309 
Rata-balat-wel, 1. 288 
Rata-gowa, ll. 145 
Rata-jambu, ii. 188 
Rata-kékiri, ii. 250 
Ratambala, i. 348 
Rata sapu, i. 22 
Rata-tora, ii. 80, 108 
Rata-uguressa, i. 73 
Rat-beraliya, 1. 125 
Rat-kihiri, ii. 125 
Rat-kohomba, il. 23 
Rattota, i. 108 
Ratu-mihiriya, i. 109 


Ratu-tiya, i. 108 
Ratu-wa, ll. 105 
Rawan-idala, i. 297 


Samadara, i. 231 
Sapu, 1. 15 

Saya, li. 316 
Sembu-nerinchi, i. 194 
Siviya, i. 237 
Siyambala, ii. 114 
Suriya, ii. 158 
Suriya-mara, ll. 129 
Suwanda, i. 104 


Tarana, ll. 328 
Télaberiya, il. 257 
Telambu, i. 164 
Tel-domba, i. 100 
Tel-hiriya, 1. 285 
Tembiliya, ii. 182 
Tilo-guru, i. 80 
Tiniya, i. 122 

Titta, 1. 75 
Titta-hondala, ii. 243 
Titta-kinda, i. 39 
Titta-wel, 1. 40 
Titta-weralu, 1, 185 
Tolol, i. 75 

To-wel, i. 288 
Tumba-karivila, ii. 249 
Tumpat-kurundu, i. 223 


Ubbériya, ii. £55 
Uguressa, i. 73 
Ulundu, ii. 72 
Uru-éta, i. 260 
Urugenda, i. 90 
Uruhonda, i. 260, 274 
Urukanu, i. 260 
Utiu-tora, ii. 105 


Wa, il. 108 

Wadiga, ii. 330 
Wal-aba, i. 57 
Wal-amba, i. 317 
Wal-awara, il. 67 
Wal-awari, li. 25 
Wal-bevila, i. 143 
Wal-bilin, i. 230 
Wal-asamodagan, ii. 279 
Wal-buruta, 1. 15 
Wal-diyalabu, i. 292 
Wal-ékaweriya, il. 320 
Wal-endaru, ii. 277, 280 
Wal-gonika, ii. 359 
Wal-karapincha, i. 218 
Wal-kinda, i. 38 
Wal-kobbe, i. 302 
Wal-kollu, ii. 79 
Walkopi i. 335 
Wal-kudulu, i. 212 
Wal-mé, ii. 70 
Wal-mora, 1. 305 
Wal-munamal, ii. 237 
Wal-nanu, il. 36 
Wal-nawahandi, ii. 266 


Wal-niviti, i. 291 
Wal-patpadagam, ii. 314 
Wal-rasakinda, ii. 261 
Wal-rat-diyaiabu, i. 294 
Wal-sapu, i. 14 
Walu-kina, i. 102 
Wal-waraka, ii. 237 
Wampara, i. 12 
Wam-bodiya-wel, 1. 279 
Wana-potu, i. 270 
Wana-sapu, i. 22 
Wanduru-mé, ii. 74 
Wata-essa, li. 145 
Wéda-pana, i. 222 


Achchu, i. 178 
Adampu, ii. 19 
Akatti, ii. 35 

Amarai, i. 303 
Ampallai, i. 327 
Anaikuntumani, ii, 120 
Anaimullai, ii. 124 
Anaittadichchal, i, 296 
Atti, ii. 116 
Attuchankulai, i. 76 
Attukaddupuli, ii. rrr 
Attuneddi, ii. 39 
Avarai, ii. 106 


Chadachchi, i. 175 
Chaddavakku, i. 250 
Charanai, ii. 270 
Chavandalai, i. 173 
Chaya, ii. 316 
Chélampai, ii. 294 
Cheppunerinchi, ii. 22 
Chemmanatti, i. 190 
Chettupulukodi, ii. x 
Chevakanpudu, i. 143 
Chilanti, i. 233 

Chintil, i. 39 
Chivanarvempu, ii. 23 
Chiruchemmanatti, i. 191 
Chirukandal, ii. 152 
Chirillantai, i. 255 
Chirunerinchi, i. 194 
Chiruppayaru, ii. 72 
Chirupiyari, i. 271 
Chiruppadikkirai, ii. 269 
Chirupunnai, i. 99 
Chittamaddi, i. 143 
Chittakatti, ii. 34 
Chomuntiri, i. 167 
Chundan, i. 277 
Churai, i. 280, 282; 


Ichavalai, ii. 132 


il, 247 


Index to Tamil Names. 


Weéla, i. 57 

Welanga, i. 169 
Welangiriya, i. 64, 224 
Wel-buté, ii. 136 
Wel-butsarana, ii. 323 
Wel-damaniya, ii. 63 
Weli-kaha, ii. 222 
Wel-kéliya, i. 176 
Wéelipenna, ii. 157 
Wel-mediya, i. 176 
Wel-radaliya, ii. 3 
Weni-wel, i. 41 
Weralu, i. 184. 
Weraniya, ii. 304 


TAMIL* NAMES. 


Tlantai, i. 280 
Tyamalai, il, 132 
Iyavakai, ii. 10 


Kachaddai, ii. 19 
Kachchantiray, ii. 271 
Kadalranchi, i, 256 
Kadukkay, ii. 159 
Kadukkodi, i. 45 
Kadduma, i. 317 
Kaddumallikai, ii. 352 
Kaddumuntiri, i. 291 
Kaddupuli, ii. 112 
Kakaipalai, i. 218; ii, 237 
Kaludai, ii. 124 
Kandai, i. 215 
Kandal, ii. 151 
Kannakompu, i, 246 
Kannu, il. 156 

Karai, ii. 330, 346 
Karanai, li, 328 
Karankutti, 1, 348 
Karivempu, i. 220 
Karukkuvachchi, 1, 272 
Karunchurai, i, 64 
Karuttappu, ii. 75 
Karuvel, ii. 122 
Katkarai, i. 233 
Kattikaya, il. 222 
Kattoddi, i. 61 
Katukali, i. 73 
Katuperatti, i, 178 
Kavali, i. 164 

Kavani, ii. 103 
Kavarachu, i. 158 
Kaya, ii. 182 

Kilivai, i, 236 
Kilukiluppai, il, 15 
Kinnai, ii. 230 

Kiri, i. 91 
Kodalimurunkai, ii, 125 
Kodippayaru, il. 74 


392 


Weétakolu, ii. 252 
Wisaduli, i. 146 footnote 


Yakada-wel, i. 22, 279 
Yakahalu, i. 121, 123 
Yak-bériya, ii. 19 
Yak-erabadu, ii. 64 
Yak-eraminiya, i. 281 
Yakinaran, i. 227 
Yak-komadu, ii. 253 
Yogana-wel, ii. 372 


Koditaviddai, i. 174 
Kokottai, i. 98 
Koliavarai, ii. 67 
Kolinchi, ii. 31 
Kona, ii. 128 
Kongu, i. 70 
Konnai, i. 275 
Korakkaipuli, i. 95 
Kovvai, ii. 247 
Kula, i. 304 
Kuntumani, ii. 57 
Kuruntu, i. 227 
Kutiraivali, ii. 44 


Makal, i. 75 
Malaiparutti, i. 166 
Malaivempu, i. 243 
Manali, ii. 273. 
Malanninchil, ii. 285 
Manchalkadampa, li. 203 
Manchavanna, li. 354 
Mara-illupai, i, 24 
Marungi, ii. 171 
Marutonti, ii. 228 
Marutu, ii. 160 
Mayladikkuruntu, ii, 228 
Metukku, ii, 250 

Minni, ii. 76 
Mochumochukkai, ii. 254, 260 
Motirakanni, i. 189 
Mudkondai, i. 63 
Mutirai, i. 253 
Mulanninchil, i. 73 
Mulkilivai, i. 237 
Mullu-murukku, ii. 63 


Naka, i. 105 
Nakakalli, ii. 267 
Nanti, ii. 25 


Narilantai, i. 282 
Narippayaru, ii. 71 
Nattaichchuri, ii. 37% 


* Lam ignorant of the correct spelling of many of these names, and have been unable to 
give the proper English transliteration in such cases, 


392 


Naval, ii. 179 
Navala, i. 59 
Navilankai, i. 59 
Nedunarai, i. 24 
Neykkoddan, i. 307 
Nirkadampa, ii. 294 
Nirnaval, ii. 174 
Nurai, i. 309 
Nutipakal, li, 248 


Odai, ii. 123 
Odi, i. 318 
Oritadtamarai, i. 67 


Painkuray, ii. 348 
Pakal, ii. 248 
Pakkilipal, i. 239 
Palampadu, i. 145 
Paluppakal, ii. 249 
Panalai, i. 307 
Pannai, i. 22T 

Panir, ii. 338 
Papparappuli, i. 159 
Parasu, il. 66 
Parutti, i. 160 
Patpadakam, ii. 272 
Pavaddai, ii. 349 
Pavaddaikaya, li. 222 
Perilantai, i. 280 
Perukka, i. 159 
Perunkuruntu, i. 226 
Perumaddi, i. 153 
Perunaval, ii. 179 
Perunpiyari, i. 271 
Peruntutti, i. 145 
Peykkomaddi, ii. 253 
Peykuruntu, i. 227 
Peyppichukku, ii. 252 
Peyppudal, ii. 256 


Phandatullai, ii. 6z 
Pichukku, ii. 251 
Pichchuvilatti, 1. 63 
Pikku, ii. 251 
Pirandai, i. 289 
Piyari, i. 271 
Podivilangu, ii. 245 
Ponnaimurankai, ii. 129 
Ponnantakarai, ii. 105 
Puchini, ii. 252 
Pudal, ii. 245 

Pudan, ii. 331 

Puli, ii. 115 
Pulikkirai, i. 89 
Punku, ii. 91 
Punaikkalaichchi, ii. 98, 61 
Punnai, i. 100 
Punnikki, i. 171 
Purankainari, i. 221 
Puvarachu, i. 158 
Puvu, i. 304 


Takarai, ii. 105 
Tamarai, i. 51 
Tanakku, i li. 165 
Tanti, i. 159 
Tattappayaru, ii. 76 
Tavarai ii. 80 
Taviddai, i. 176 
Tayirvalai, i. 57 
Tekil, ii. 9 
Tévataram, i. 190 
Tirukkontai, ii. 103 
Tiruvatti, ii. 116 
Tommakottai, i. 100 
Tumpai, ii. 249 
Tumpalai, i. 129 
Tuvadi, i. 284 


LONDON: 


Index to Tamil Names. 


Udai, ii. 123 
Udaivel, ii. 124 
Uluntu, ii. 72 
Uluvintai, i. 25 
Uyil, ii. r30 


Vaddutakarai, ii. 106 
Vaddattutti, i. 145, 154 
Vakai, ii. 104, 108, 128 
Vallarai, ii. 276 
Vallampuri, i, 168 
Vammi, ii. 292 
Vandakkay, i. 156 
Vankiruvalai, 11.268 
Vanni, ii. Tor 
Vatchikuran, 11. 343 
Vedchi, ii. 348 
Vekkali, ii. 162 

Vella, ii. 336 
Vellaikadampa, ii. 293 
Veluruvai, ii. 88 
Velvel, ii. 125 
Vempadam, i. 279 
Vempu, i. 244 
Venachchi, i. 61 
Venkai, ii. 90 
Venkalikaya, ii. 182, 222 
Vet-pavaddai, ii. 351 
Vidattal, ii. r2r 
Vidpani, i. 172 

Vila, i. 228 

Vildtti, i i, 228 

Vili, i. 60 

Vilvam, i. 229 
Vinanku, i. 169 
Virali, i. 312 


Yar, ii. 371 
Yerkoli, ii. 343 


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