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IE CRAOUATEO SEBtCS OF ftEAOINC-BOOKS.
.^^i.-ii'"'/'
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HANDBOOK
OF
ZOOLOGY.
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o
HANDBOOK
ZOOLOGY
BY
X VAN DER HOEVEN,
FHIL. ITAT. BT M.D. PS0FBS80R OF ZOOUMY IN THK ITKITBBSriT OF LBTDKN. CriGHT OF THB OEDBK
OF THB DUTCH UON AND OF THE SWEDISH ORDER OF THB POLAR STAR. MEMBER OF THB ROYAL
AGADMMT OF SCIEKCBB, OF THE DUTCH SOCIErT OF BdENCBB AT HAARLEM. OF THB IMPERIAL
UBOPOLDO-CAROUNE ACADEMY. OF THB IMPERIAL 80CIBTT OF NATURAUSTS AT MOSCOW. OOR-
BMBPOITDINO MEMKER OF THB BUTISH ASSOCIATION. OF THB ROYAL ACADEMY OF SCIBNCBB AT
TUKIK. OF THB ROYAL ACADEMY AT PARIS. BTO.
Trsdo qiue pofcni.
IN TWO VOLUMES.
VOLUME THB FIRST.
( INVgETBBRATM ANIMALS. )
TRANSLATED FROM THE SECOND DUTCH EDITION
THE REV. WILLIAM CLARK, M.D. F.R.S. Ac.
LATB m.LOW or TBiiriTT ooLUox, Am rsoraaoB or axatomt
IK THC oirrrsiiaiTT or OAVBRriHix.
CDambtflige:
PRINTED AT THE UNIVERSITY PRESS,
roK
LONGMAN, BROWN, GREEN, LONGMANS, AND ROBERTS,
LONDON.
1856.
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Ki ;iT^(o'^o (^^)
/
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PREFACE.
Ths first Edition of Professor Van dxr Hoeykn's manual
was published in parts between the years 1827 and 1835.
He undertook the labour, as he informs us, not with any
desire to add one more to the numerous works with a
similar title already in existence, which should be neither
better nor worse than these. On the contrary, if he had
found any one of them to be a sufficient guide for his
public teaching, without great alterations, he would have
abstained from his contemplated task. Accordingly, the
plan of his work differed from that of most other manuals
in beginning with the simpler forms of animals, and pro-
ceeding upwards to the highest : and from that of all of
ihem, in embodying a much larger amount of anatomical
information. His work was received with marked appro-
bation not confined to the limits of his own coimtry.
During the lapse of nineteen years, which intervened
between the first and second editions, the acquisitions
both of Zoology and Zootomy had been greatly enlarged,
so that in many departments the former science had
assumed an entirely new aspect. Consequently the second
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VI PKEFACE.
edition, similar in plan to the first but greatly different in
its contents, was almost entirely re-written. This edition
has, like the first, been published in parts between the
years 1846 and 1855, inclusive. From the high terms in
which it has already been alluded to firom time to time,
in the writings of various active cultivators of different
departments of Zoology on the continent, it is obvious
that the general estimation of his work will still be such
as might be expected in the case of an author of vast
erudition, of appropriate tastes^ talents and genius, and
whose office it has been for nearly thirty years, as Pro-
fessor of Zoology in the University of Leyden, to bring
the value and import of the new acquisitions of Ana-
tomy and Zoology (many of them the result of his own
labours) from time to time before his auditors.
The University of Cambridge, a few years ago, directed
in a more marked manner the attention of our students to
the Moral and Natural Sciences, by proposing honorary
distinctions to those who might excel in certain depart-
ments of those sciences respectively; and by requiring
proof of satisfactory attention to some one at least of such
departments on the part of all candidates for the degree
of Bachelor of Arts, who were not aspirants for Mathe-
matical honours. Amongst the departments of Natural
Science, Comparative Anatomy and Physiology were
indicated, with special regard (as is presumed) to Zoology.
It thus became a part of my office to place within reach
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PREFACE. Vll
of our students the best assistance I could recommend for
their studies in this direction. In fulfilment of it I ap-
plied to Professor Van der Hoeven for his permission
to translate his Work, in which I found all that could
be required^ He had the kindness not only to grant this
permission, but also to enrich the English translation with
numerous references to works too recent for notice in his
own second edition. It is to be much regretted that his
other engagements did not allow him, as I requested,
to weave the new matter in his own terse and pleasing
style into his introductions to the classes and elsewhere.
Consequently such additions, in this respect, as are in-
cluded within square brackets are mine.
The study of Zoology is now in such general favour
with cultivated persons in this country, that I believe the
present work, from its scientific value and the interest of
its historical and other notices, as well as from the con-
tinuous references to the works of the original discoverers,
will secure for itself, beyond the walls of Universities, a
reception not unworthy of its Author's great name.
W. C.
Cambbidox, July i, 1856.
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CONTENTS
OF
VOLUME THE FIRST.
»4«B
iHTBODUCnOV 1^36
Organic and Inorganic Bodies 3 — 4
Flaots and Animals 4-* 7
Zoology 7—8
The 1188068 of Ammals 9—19
The Vital FanctionB of Animals 19 — 34
Derdopment of Animals 94 — 26
On the Alt of dasBifying (ITaKifMMMfa) 36 — 36
Glass I.— Infusobhs. {Infd99na). . . 37—59
Spermaiogoii, Bo-caBed seminal animalcules 43 — 47
Systematic Arrangement of Inlnsories 45 — ^59
Older L Inftuoria nmplieisnma 4j
7am. I. Vibrionida A.
Order II. Hkkopoda 45— 50
Fam. II. Amctbaa 46
„ m. Aredlina 4^—6^
Order m. Airidia 50—51
Fam. lY. Manadina 50
„ ▼. (kypiamonadina A.
„ Yl. Vohocina t&.
„ Tn. Attatia 53
„ Yin. Peripkryffoma f6.
Order IV. EpUricka 53—59
Fam. IX. Peridinaa 53
„ X. Triekodina 53—55
„ XI. Oxiftrickina 55
„ xn. EupUfUk 55— 5<5
„ xm. Voriicdlina *fi — 59
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X CONTENTS.
rxoK
ClaBB II.— POLTPB. {Pdl^i.) .... 60—97
Systematic Amngement of Polyps « 73~-^
Section I. Anihozoa 73 — 84
Order L Mydtifonma 73—77
Fam. I. Hydrina 73 — 74
„ n. Seiivlcmna 75 — 77
Order II. OcUuAinia 77—84
Fam. m. Xenina 77 — 78
„ IV. Haleyonina 78 — 80
,y ▼. PennatfUina 80—81
„ Ti. Tubiparina 8s
„ VII. Cortieata 83 — 84
Order III. PUyckdinia 84 — 93
Fam. vni. Madrtporina 85
IX. OcelUfia 86
X. CfyroM 86—87
XI. Funffina 87 — 89
xn. ZoafUhina 89 — 90
xm. AcHfUna 90—91
Section n. Brywtoa 93—97
Order rV. Bryotoa 93—97
Fam. xrv. SUhntUopoda 93 — 96
„ XV. Zophopoda 96 — 97
ClasbIII. — Sba-Nbttlbb. (Acalephce.) . 98 — 136
Systematic Arrangement of Sea-netUes 108 — 136
Order I. Siphonoph4)r<B 108 — 119
Fam. I. VeUUidm 108—111
„ II. PhyttopkoridcB iii — 116
,, ni. ffippopodida 116
,, IV. IHpkyidm 116 — 119
Order IL Ctenopkora 119 — i3i
Fam. V. Btr<iKdea no — i3i
Order III. DiicophonB 133
Fam. VI. Qtryonida 133—133
„ VII. JZAtzoitomtcZes 133
„ vui. Medusidea 133 — 134
„ IX. OcMnida 134 — 135
„ X. jEquoridcB 135 — 136
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OONTBim. XI
tAUB
Class IV.— Eohxhodhiib, (Sekinodtrmaia). . 197—163
SyBtemuitic Amngement of Echinodenns. 143 — 16a
Older I. Eekmodu iwufa pedieUlaia {PedkulaU Sekm,) 143—159
Fam. I. OnmeSdm 143—146
„ n. Atteridea 146 — 149
„ lu. Sehinidea 150—156
„ IV. HoUthuridea 156 — 159
Order n. Apoda 159 — 161
Fam. ▼. SynaptintB 159 — x6o
„ Ti. SipuHculaeea 160—163
Glass Y.— IifTisnirAL Wobmb. {EnUaoa). . 163 — 193
Sjsiematio Arrangement of the Intestinal Worms 178 — 193
Older I. Sterdminiha 178—188
Fam. I. Cettwdea 178—184
„ n. Aeanihocephala 184
„ m. TrenuUoda 184—188
Order II. Oodelmintka 18S— 193
Fam. IT. NematoUUa 188—193
Appendix to the Class of Intestinal Worms 193 — 194
Class VI. — ^Whskl-Ahdcalculbs. {Rotatoria). 194—306
Systematic Arrangement of Wheel-animalcmlBS 199 — 3o6
Order sing^le. MUatoria 199—306
Fam. I. Flotetdarke . 199 — 300
„ n. Mdicertina 300—301
„ IIL Braehioncea 30i — 303
„ lY. Hydaiinoea 303 — 305
„ ▼. PkUodinan 305 — 306
Class VIL—RiXGiD Worms. (Annidata). . 307—346
Systematic Arrangement of Binged- worms 319 — 346
Order I. TurbeUaria 319 — 334
Fam. I. Planariea 319 — 333
„ II. Nemerttni 333 — 334
Order IL SueUma 335—338
Fam. m. Birudinea 335 — 338
Order HI. SeUgera 338
Fam. IT. LumMeini 338 — 333
„ T. Maldama 333 — 334
„ TI. AmphiirUa «34— «37
„ Tu. ArmieoUg 337
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Xll
CONTENTS.
Fam. vin. Ch£Btoptenna 437 — 438
IX. Peripatina 938
X. Aricia «38— «39
XI. Nemda 139— 14a
xn. EumceiB 34^ — ^43
xm. AmphinomaeecB '243 — ^44
XIV. ApkrodUaeeiB 344 — M^
Class Vm.—lK8B0T8. {Inteda), . «47— 555
Systematic Anrangement of Insects 988 — ^555
Order I. Myriapoda a88— 296
Fam. I. Jidida «89— 193
„ 11. Seolopendriiks 993 — 996
Order II. Thysanwra 196—300
Fam. m. Zepitmena ^97 — *9^
„ IV. PodurOla 198—300
Order m. ParatiHea 300—303
Fam. V. ffcemaiopina 300 — 301
„ VI. MaUopkaga 301—303
Order rV. Sudoria 303 — 305
Fam. vn. PuUHcUb 303—305
Order V. Shreptiptera 305—308
Fam. vni. Sirepnplera 307 — 308
Order VI. JHptera 308—346
Fam. IX. Pupiparce Z^^—ZH
„ X. Athericera 314— 3«8
„ XI. TanysUmata 3^8—335
„ xn. Notaeaniha 335—339
„ xjn. Nemoeera 339 — 346
Order VII. ffymenopiera 346—389
Fam. XIV. Mdlifera 350— 359
XV. DiplopUryga 359— 36«
XVI. Ederogyna 363 — 367
xvn. Fouora 367 — 371
XVIII. ChrysidideB 37i— 37«
XIX. (hyura 373—375
XX. Cfhalcidia 375—378
XXI. Ichneumontdes 378 — 383
xxn. Cynip^ea 383—385
xxm. Urocerata 385—386
XXIV. TetUhredinda 387—389
Order VIII. Lqddoptera 389—4"
Fam. XXV. Noctuma 393— 404
„ XXVI. Creptuctdaria 404 — 406
,, XXVII. JHuma 407—411
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CONTENTS. XIU
WABM
Order IX« NetsropUra 413 — ^497
Fam. zxnn. Pkrffffanidea 413—415
99 xzix. PanarpaUB 415 — 416
„ XXX. ffemerobmi 416 — ^410
„ XXXI. LibeUmlimai 490—433
„ xxxn. Bphewterina 433 — 435
„ xxxm. Perlaria 435
„ xxxiY. TerwMfUB 436—437
Order X. JlewiipUra 437 — 448
Fam. xxxv. Coccma 430 — 433
„ XXXYI. Aphidii 439^435
„ xxxvn. CieadainKB 435—439
9, XXXVIII. Hydncorim ...... 439 — 441
9, XXTTT. Oeoeoriw 443 — ^448
Order XL Orthopiera 448—464
Fam. XL. ChryiUdei 451—458
„ xu. ManHdet 458—461
„ xui. JBhUtaria 461—463
„ xuii. Fot:fieultm€B 463 — ^464
Appendix to the Orikoptera, Tkiftcmopfera, Qtrnxm Thript ... 464
Order XII. CoitopUra ^^—553
Fam. xuv. CoeemeUida 467—468
,9 XLV. Fvmgkfi^ 468—469
99 XLVi. CUvripdlpi 469—470
„ XLvn. Oydica 471—474
,y XLTiii. Eupoda 474 — 476
,, xux. Maeroeerata ...... 476—480
,9 ^ . Seo^fkuria 480—483
„ u. Bhynckopkora 483—489
„ m. Stendfira 49^—493
„ un. Taaieomei 493—495
9, uv. MeUuamata 495 — ^499
„ LV. (kmiharidia 499—504
„ LVL JkmdUeorma 504 — ^519
y, LYiL XyUphaga 530—533
9, Lvm. SerricofnUa 533 — ^537
f> ux. Braehdytra 538—533
,9 LX. Clavicomia 533 — 540
„ LXL PalpicortUa 540 — 543
„ LXn. Bjfdroeainiharma 543 — ^544
„ Lxm. CarabicifM 545—555
CLiJSS IX.— Abachnids. (AraehnoHdea). . . 556—597
Systematic Arrangement of Aradmids 571 — 597
Order I. Polysfonapoda 57i__573
Fam. I. Pfcnoffonida 571 — ^573
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XIV CONTENTS.
rAox
Order II. Cdopoda 574—575
Fam. 11. Arctiaca 574—575
Ord^rlll. Acarina 575— 58^
Fam. in. Acarea 575— 57<5
„ IV. Nolaapidea 57^5—577
„ V. Ixodea 577
VI. Ganuuea 577—579
„ VII. ffydramchnidia 579 — 581
„ vin. BdeUea . . . ' 581
„ IX. TrombidifM 581 — 582
Orderly. PhalanffUa 58a— 584
Fam. X. PheUoMffita 582—584
Order V, Pseudoacorpiones 584
Fam. XI. Pteudoaeorpiones 584
Older VI. Solifugm 584—585
Fam. XII. CfaUodea 584 — 585
Order VII. Pedipalpi 585—586
Fam. xni. Phrynide* 585 — 586
„ XIV. Scorpiona 586—587
Order Vm. Aramddea . 588—597
Fam. XV. ArcmMea 588 — 597
Class X.-— Crustaobahs. {Orwitacea) . . 598 — 679
Systematic Arrangement of Crustaceans 622 — 679
Order I. Pcecihpoda 622—^24
Fam. I. Xiphotwra 622 — 624
Order II. lehihyopJUhira 624 — 631
Fam. II. Lemaouxa 624 — 625
„ III. Lemaopoda 625 — 627
„ IV. JSrgatUina 627 — 629
„ V. CaUffina 629 — 631
„ VI. Argfdina 631
Order III. Lophyropoda 631—^33
Fam. vn. Copepoda 632—^33
„ Vin. OOracoda 633
Order IV. (Hrripedia 633 — 640
Fam. IX. Bcdanoidea 636 — 637
„ X. Ltpadicea 638—^40
Order V. Cladoeera 640 — 641
Fam. XI. Daphnidea 640^641
Order VI. PhyUcpoda 641--646
Fam. XII. JSranckiopoda 641 — 642
„ xm. Atpidephora 642 — 644
Trilobitee r. Paladades 644—646
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CONTENTS. XV
TkOU
Order VII. Iwpoda 646—655
Fam. xiT. Bpieandm 646 — 647
„ XT. CymMkoadea 647 — 649
,» XTi. Sphwnmiida 649
„ XYn. Praniddea 649 — 650
„ XYin. Onitdda 650—659
,y XIX. AadUfUk 65a — 654
„ XX. Idoteidea 654—655
Order VIII. Ampkijpoda 655—660
Fam. XXI. Lomodipoda 655—657
,, xxn. ffyperinaa, Uropiera 657 — 658
„ xxin. Qatntnarina 658 — 660
Order IX. SUmatopoda 660—664
Fain.xxnr. UnipdkOa 660—661
„ xxv. BipeUaia 661—661
„ XXVI. Oariduadea «. SMupoda 662— 66$
Oumaeea (Funily of uncertMB pomiioD) 663
Order X. Jkcapoda 664 — 679
Fam. xxvn. Caridina 664 — 666
„ xxvni. Aataeina 667 — 669
,, XXIX. Iiorieala 669 — 670
„ XXX. Anomwn 670—671
„ XXXI. Noiapoda 679—673
„ xxxn. Oxyttonusta 673—674
y, xxxm. Majacea 674—^76
„ xxxiy. Canerina 676—679
On MoHubcs in general 680—690
Clasb XI.— TuiriCATXS. (Tunieak^ . 691—696
Systematic Anangemeot of Tanicatee 697 — 707
Order I. ThaUaoea 697 — 700
Fam. I. StJpma 697 — 700
Order n. TdhfoMUa 701—707
Fam. n. Lueia 709
„ m. A$ciduB 709 — 707
Class XII.— CknroHinats or Bivalvss. {Omckifera) . . 708—757
Systematic Arrangement of Conchiferi 719 — 757
Order I. Pattiobranekiata 719 — 793
Fam. I. BraekUtpoda , 715^793
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XVI
CONTENTS.
Order II. LamdlibranMUa 7^3^757
Fam. II. Ottracea 7«5— 7^7
in. PeeUnidea 7^7—718
IV. MaUeacea 729 — 731
V. Mytilaeea 731—733
VI. Arcaeea 733—734
vn. Trigwiiaeea 734
VIII. NaXadea 734 — 737
iz. CardUaeea 737
X. Chamaoea 737—738
Rudida 738—739
ZI. Tridacnacea 739 — 740
XII. Cardkicea 740—741
xin. Veneraoea 741 — 743
xnr. Oydadea 743—744
XV, Lueinaoea 744 — 745
XVI. Saxicavina 745 — 746
xvn. TelUnacea 746 — 748
xviu. Macfiraoea 748 — 750
XIX. Myacea 750 — 751
XX. PholadMiiyacta 753 — 753
XXI. SoUnacea 753—754
xxn. Phoiadacea 754— 75^
xxm. Tvbicola 75^—757
Clabs XIII.— Molluscs. {MolluMa)
758-831
SyBtematio Arrftngement of Molluscs 774 — 831
Ordar I. Ptoropoda 77«— 77^
Fam. I. HyoiUacea 773—775
„ II. CUo/idea 775— 77<>
Order II. Oatiaropoda 776—817
Fam. m. Meteropoda 776—778
„ IV. Ikrmatobranchiata 778 — 784
,, V. ffypobranehiaia 784 — 785
„ VI. Pleurobranchiata 785—788
„ vn. Oydobranckiala 788 — 791
f, vin. Aipiddltranckiata 791 — 793
„ IX. AuldbranchicUa 793
„ X. Cten6hr€mchiaia 793 — 811
„ XI. Pneumonioa 81 r — 817
Order III. Cephaiopoda 817— 83T
Fam. xn. NtnUilacea 835 — 828
„ xin. Sepiacea 828 — 830
„ XIV. OOopoda 830—831
Explanation of Plates 833—844
Index of Generic Names . 845 — 853
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ERRATA AND 00RRBCTI0N8.
VAOI
LJKS
vom
aBA9
5
»5
SOHWAMMKBDAX
SwAmasDAX.
i6
arterial
TenotiB.
«3
Soemmering
ScenmieiTing.
^7
13
called yariettet.
constitute varieties.
3«
19
worse
now more usual.
67
4 uToin Dottom
oonseqnenoe of
after the.
79
5 from bottom
yniHiip,
metre.
87
'9
Fungina Lam.
FungiaJuAM.
«9
'3, 14
might periiaps, fte.
seems to require new observations
propriety.
9I«
6
rMkjnm^im
vertebrates.
«57
lafinm bottom
take Bcaroely any
^63,
note 9.
Yauouvltv
VAUQITILIir.
«99
10
aaUfUhMrus.
SmyiUhurtu.
4i«
8 from bottom
pupa
larva.
4^
13 from bottom
494
7
J7eZetif
ffeUsui.
672
JO from bottom
Drxwia
Dromia.
Hie following, not considered distinct genera^ but divisions of the genus imme-
diately above each respectively, ought to have been printed in the smaller type used :
CMwmia Ehbbhb. .
58
ChraUivm TjAU.
83
ifetitea Lax. .
83
/mLam. ....
83
FuMgiaJiAU. .
87
«38
OphdiaSKy. .
138
«44
«44
CfhloeiaQjLY
«44
CmioeeraMAOQ.
3^3
Awophda MxiG. .
346
rAes
Apii Latb.
. 35«
MeUUwrgaloLrB. .
. 356
PofiltetLATB. .
. 363
Labidui JuBori .
. 366
Bethyhu Latb. .
. 375
BUopia TsBsrecBKB,
397
£%elidia Ochbbkh. .
• 399
Phuia OOHSEMU. .
• 399
ffdceut JjAXR. .
. 494
. 651
Trichonueut BRAinn
. 651
Orouwtu Rathu .
654
The following distinct genus ought to have been printed in the laiger type :
Sipkonaria Sow. p. 795.
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INTRODUCTION.
We often hear the word Nature used in such a way that it is
diffictdt to know what meaning is attached to it. Some denote
thereby the system of all the forces to which matter is subjected,
and thus distinguish between Nature and the Universe; under-
standing by the last the entire complex of created bodie."*.
But such a distinction is quite arbitrary. The word Nature,
introduced into modem languages from the Latin, is derived from
nasciy to be born, to come into being^. In this sense we call the
aggregate of all that comes, or has come, into being, and is for us
an object of observation either by external sense or internal percep-
tion, Nature — the material world and the spiritual world — ^Nature
in space and Nature in thought. Finally we oppose Nature to Art,
understanding by the last whatever change the intellect of man has
induced upon the products of Creation, in order to satisfy his wants,
or to enhance his enjoyments.
However different these and other meanings may be, we may
admit that to be the most general which defines nature as the
material world, the world of matter, all that is created or has being,
together with the forces inherent in the matter, and the laws
according to which they act. The knowledge of this whole, so
stupendously vast, the ancients named jghysica : and considered to
he a part of the philosophy which they termed a science of divine
and himian things and of their causes. But though this science, like
nature its object, be one, yet its great extent on the one hand and
the narrowness of the human intellect on the other, has rendered
the subdivision of it necessary. Yet the limits of the different
natural sciences can scarcely, on account of their mutual relations,
^ So the Greek ^(nnt from 4>^'
VOL. I.
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2 INTRODUCTION.
be defined with precision: and into whatever path we chance to
strike, difficulties from the very nature of the case are unavoidable.
The Natural Sciences relate either to Nature and her several
products considered by themselves : or they teach us so to apply
those products as to contribute to our service, or to satisfy our
wants. The latter are called practical natural sciences, the former
iheorettcaL To the practical natural sciences belong especially
Agriculture and Technology: and they are founded upon the
theoretical; of which the truths are applied only in a degree propor-
tioned to the particular object that is had in view. They may
therefore be called Applied Natural Sciences. Of the pure, or
theoretical Natural Sciences there are several. To them belong^
Phaenomenal Doctrine, Chemistry, and Natural History. What
characterises such sciences and distinguishes them from each other
lies less in the objects which belong to the province of each, than
in the manner of considering them, and in the different direction
of the enquiry. Metals, salts, earths belong as much to the province
of Chemistry as to that of Natural History : but the chemist, in
all these things, investigates only the matter and its properties, its
affinities and combinations : the mineralogist is busied with their
form, their natural occurrence, their classification. The chemist,
moreover, investigates those elements which occur in nature only
in combination with other matters : such elementary substances are
excluded from the province of Natural History.
Whilst Physics investigate the common properties of bodies, and
the motions by which a temporary change is effected in their
condition. Chemistry enquires into their component parts, the
special properties of each elementary substance, and its various
combinations with other elementary substances. Natural History,
finally, arranges the bodies occurring in Nature according to form.
In a certain sense, therefore, it may be termed a special Phenomenal
Doctrine : but its essence lies in describing and classifying. It is
ordinarily limited to the bodies which occur upon the surface of
our earth, or at small depths below and accessible by mining : but
it is by no means necessary thus to limit it. It depends upon the
* [Natur-lehre, The VMt body of observed facts throughout nature " bound together
under the form of laws and principles." Vid. Whewkll'b History of the Inductive
Science*, and his PkUoeophy ^ ike Inductive Seiencee, passim.] TV.
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PRIMARY DIVISION OF BODIES. 3
manner of treating it alone, to include within its province not only
the planet which we inhabit, but even the whole visible universe ^
The certainty to which we can attain in this science is grounded
upon the testimony of our own senses, on that of other observers,
and upon conclusions, drawn from the combination of particular
observations (Induction) : of which conclusions the security is
increased in proportion to the number of observed phaBnomena
which tend to their establishment ; for observation is the principal
foundation upon which this science is raised.
Organic and Inorganic Bodies.
A primary division of the bodies of our earth is that according
to which they are separated into organic {organtca), and inorganic
{anorganica). Inorganic bodies can grow, or increase in bulk, only
by external addition of homogeneous parts ; they possess no hetero-
geneous parts, though they may be composed of several chemical
elements. In their perfect condition they ordinarily present regular
forms, which are bounded by planes and straight lines. The
knowledge of them is the object of Mineralogy.
The remaining bodies are called organic, because they consist
of different parts, of fibres, vessels, cells, &c., the combination of
which is called organisation. In these bodies there prevails that
mutual dependence between all the parts, of which, in the inorganic,
we recognise no trace. In these last, each of the parts exists for
itself, and when separated from the whole does not cease to be the
same that the whole was before. As to form, the boundaries which
drcumscribe plants and animals are very generally round surfaces
and carved lines, very rarely straight lines and planes.
Organic beings present phaenomena which are called 'Vital
Phenomena,' of which the most general consist in an incessant
susception of new matters, in the formation of new parts and
organs (Growth, Development, Reproduction), and in the production
of similar beings (Propagation). The separation of those constitu-
ents of food that are unfit for nutriment, and of matters that have
been changed through the action of life, and are no longer fitted for
Cotmograiphia, HiM/oria Mvndi.
1—2
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4 INTRODUCTION.
its support, supplies the means hj which the peculiar chemical
composition, characteristic of each individual being, is preserved;
at death, on the cessation of this interchange of matter, the organic
substance passes into solution or putrefaction. Instead of those
complex combinations of elements, which form the proximate con-
stituents of organic bodies, simpler combinations arise, which being
taken up by the air or the earth, become anew the vital stimulants
and the nutriment of that vegetable world, on whose existence the
life of animals is dependent. Thus we perceive here an interchange
of matter on a large scale, as we do in every organic being on a
small one : and perishableness becomes the means whereby new life
and fresh youth are efiFdsed over the whole of nature ^
Plants and Animals,
The above may suffice to give a general notion of organised
bodies. It scarcely requires notice that the term includes plants
and animala.
At first sight it seems easy to distinguish an animal from a
plant: and even the most unskilled person thinks he has a clear
notion of the difference. Yet it is just his want of knowledge that
causes the difference to appear so prominent : whilst he overlooks
the intermediate links, and thinks, for instance, of a dog and a
peax-tree. There are two sorts of judgment with conviction. Such
a judgment may arise either from want of knowledge, or from pro-
found insight, the result of long and accurate investigation. Who-
ever seeks after truth must learn to sacrifice the first, even though
he may never attain to the second.
Animals are usually considered as more composite and more
perfect than plants. Yet when we compare the simple substance of
which Inftisory Animals and Polypi are composed with the orderly
and beautifril structure of the higher plants, we become satisfied
that this proposition is far from having a general value.
It is said, plants are rooted in the ground, and by this token
are sufficiently distinguished from animals. But it is here over-
looked, that there are free-swimming water-plants, just as there are
* " Ut opui naturof perenni Jlore rideat.*' LiKNiiUS.
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PLANTS AND ANIMALS. 5
animals that live in water &st fixed to a given spot. Again, it was
imagined that plants might be distinguished from animals by their
possessing only at certain times the parts which serve for the main*
tenance of the species. But all animals have not organs of propa-
gation during the whole of their existence : insects acquire them
only in the latest period of their lives, like plants that only flower
once. It is true that Anatomy points out the rudiments of these
future organs in the Larvae of Insects : but tliis does not prevent
the Larv» of Insects from being termed, in a certain sense, sex-less.
Moreover many plants and animals are propagated only by spon-
taneous fission, or by buds, without possessing proper parts sub-
servient to propagation. To me the difference of Nutrition appears
of more importance. It has been observed that Plants live on inor-
ganic matters, Animals, on the other hand, on organic. Some
animals, it is true, seem to live on earth. Schwammebdam believes
mud to be the nutrient matter of the worm or larva of the Ephemera^
and never found any other food in its intestinal canal. Pallas could
find nothing but fine sand in the intestine of Uialassema echiurus.
I might allege several other examples, but will only add that man
himself sometimes lives upon earth. At least the celebrated Hum-
boldt tells of a people on the banks of the Oronoco and Meia that,
when the waters are low, live upon fish and turtle: but as soon
as the streams begin to swell and fishing to become laborious,
devour, during a season which lasts for two or three months, enor-
mous quantities of earth. The earth which these people eat is a
&t soft clay, which they knead into lumps and bum on the outside
at a slow fire, and again moisten when required for use. This
observation however, now that microscopic investigation has dis-
covered in different deposits and kinds of earth entire strata of
living or fossil organic beings, admits of another explanation ^
That earth may have contained organic constituents, as was the
case with the earth that, mixed with meal and leaves of trees, was
baked for bread on the failure of the crops in the year 1832 in the
north of Skandinavia, and in which Ketzius discovered nineteen
different forms of Infusories, or rather of the fossil remains of these
animals. In the same way, the mud and sand, found in the intes-
^ Gomp. Ehbevbero, Dm umiehtbar wrhende orgamische Leben. Leipzig, 1843.
pp. 41, 42.
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6 INTBODUCJTION.
tinal canal of insects and worms, can scarcely be devoid of organic
components. In general it cannot be contested that the vegetable
kingdom prepares from inorganic nature those substances which
serve for nutriment to the animal kingdom. Albumen and Fibrin,
the principal constituents of the blood of animals, are already
present in the parts of plants which they consume. But still, on the
other hand, there are plants which grow on other plants, and thus
apply to their own development the matters which have been pre-
pared by those organic bodies.
Scruples arising from such considerations must not mislead us
to consider plants and animals as belonging to one and the same
kingdom. Perhaps the following remarks may help to distinguish
them from each other.
If we consider the nutrition, we perceive that animals convey
their food by one or more apertures into a common cavity, the
stomach or intestinal canal, from which the prepared matters are
absorbed and applied to the nutrition of the whole body. Thus the
intestinal canal is for animals what the soil and air are for plants.
The plant is consequently so constructed that its surface has the
greatest possible extension : in the animal all is contrived for union
round a center. Moreover the plant, which receives nutriment by
means of its surface and the parts there situated, (pores, hairs, &c.)
has no need to seek for food : it lives in the midst of its food : when
this is deficient it cannot move and must consequently die. The
animal, on the contrary, is destined to seek its food, which it must
conduct into its intestinal canal : it moves therefore when nutriment
is deficient. Let it not be here objected that plants move towards
the light, and send larger roots towards the side where moisture is
more abundant — ^for this would be to confound growth with motion.
The stimulants (light, moisture, &c.) act upon the plant, and there-
fore its growth is more vigorous in that direction. The animal has
independent motion which is excited by internal stimuli. Hence
sensation is ascribed to animals. In the higher animals it is known
that the contraction of the muscles is under the influence of the
nervous system : that the stimuli, of whatever kind, if they pro-
duce motion, act upon the nerves and through these upon the
muscles. Comparative Anatomy, it is true, has, in some animals,
hitherto failed to demonstrate a nervous system ; but it does not
therefore follow that these animals do not possess sensation, any
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ZOOLOGY. 7
more than it followa that their want of muflcular fibre juBtifies us
in denying to them yolnntaty motion, which the unprejudiced
obseiver may recogniae even in the most simple animals.
As to chemicid composition — ^Nitrogen, it is trae, is not ex-
clusively an animal principle: but still it occurs as an ultimate
constituent of animal organisation in much greater quantify than
in plants. In plants, on the other hand, Carbon is predominant
Not long ago, it was believed that a transition from Plant-Life
to Animal-Life had been observed ^ L. G. Treyiranus had
remarked that the spores of Confervas move like Lifusories^ A
few years since Unger described this phenomenon in Vaucheria
clavaiaj and thought that he had surprised plants at the very
moment of their becoming animals'. These aporidia move 1^
means of dlia, but cannot on that account be considered animali'^.
Rather ought we, from such instances, to conclude that cilia are no
exclusive character of the animal kingdom. The same may be said
of a similar motion which Grant observed in sponges, which, as
little, on that accoimt, are animals.
As we have already remarked above, the difference between
plants and animals will always be more apparent as the organisa-
tion becomes more perfect and more complicated: the difficulty
occurs in the case of the most simple vegetable and animal forms;
and here may be applied what Ovid says of the change of colour in
the rainbow.
Usque adeo quod tangit idem est, taraen ultima distant —
Ma. vi. 67.
Zoology.
There are three parts of Natural History, as there are three
great divisions of the bodies which occur on our Earth. We sepa-
rate Natural History into Mineralogy, Botany and Zoology, as we
1 See the eadier obeervatioiis on this point in G. B. Tsxvibanub* Bidogie, oder
PkOotopkie der Ubendm Natur. 8to. II. p. 344.
» BeUrd^ mr PJUinMnphj^nologie, Gottingen, 181 1. Svo. pp. 78, 79.
' Die JPJUmze im Mamente der Thierwerdung. Wien. 1843. Svo.
* C. Bi SiEBOLD, Dies, de jMm nUer regMtm atUmaU ti vegeUMe conttkuendii,
1844. 4to.
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8 INTRODUCTION.
divide nature into three kingdoms, that of minerals, of plants, and
of animals.
As the history of mankind does not result from a collection of
biographies, so Natural History is not formed by the description of
animal species. Therefore Zoology describes not merely the sepa-
rate animals {m(mograph%cally) according to external parts and
internal structure, but it comprises the entire kingdom of animals,
denotes their mutual relations, and assigns to each animal its rank
and position.
Zoology falls into different parts. First, it is divided into
Description and History, Description of animals {zoographid) sup-
plies precise descriptions of the separate internal and external parts
of the animal body, and thus of the entire animal. In a narrower
sense, it makes us acquainted with the external parts and the ex-
ternal form of the animal, and with the distribution into classes
and orders. When it makes us acquainted with the internal struc-
ture of animals, as well in respect of form and position [structura)
as of tissue {textura)^ it is called the Anatomy of Animals {Zootomia),
which has been especially cultivated of late years, and is generally
named Comparative Anatomy {Anatomia Comparata). But this ap-
pellation has not exactly the same meaning as the first: it denotes,
rather, a philosophical science, which, not content with the simple
knowledge of the different forms, investigates, by comparison of the
anatomy of all animals and also of the human body, the general
laws of animal organisation and its unity.
The History of Animals {Ristoria Animalium) comprehends a
comparative history of the nature and intellect of animals : it illus-
trates the phasnomena of life, and their obedience to law in the
animal economy. It may be also termed General Physiology
(Biology). The knowledge of the geographical and physical dis-
tribution of animals over the surface of the earth, the knowledge
of the series of forms which in earlier periods inhabited our planet,
and of which the remains have been found in beds and strata of
rocks deposited from water, also belong to the History of the Ani-
mal Kingdom.
These subdivisions cannot dispense with mutual assistance.
Conjointly they form only one science which we term Zoology.
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ANIMAL TI88UES. 9
The TiMues of Animals.
Nature, in the structme of animalB, has exhibited inconceivable
resources of art. Not only is the body as a whole, not only are its
grosser parts, but even tlie smallest parts of the organs are ma-
chinery; and the knife of the anatomist exhibits to us, even to the
simplest fibre, nothing but parts constructed for a purpose. The
whole body consists of many members : the members themselves
consist of muscles, vessels, and nerves; the muscles again of
tissues, vessels, fibres, &c. Such a body may well be called
organised.
In order to form an idea of the texture of animals, we must go
to work as the chemist does. He divides the parts which compose
bodies into proximate and ultimate constituents, and terms those
constituents ultimate, or elements (elementa), which by his art he
can separate no farther into heterogeneous parts. In the same way
we find in the textures of animals proximate and ultimate con-
stituents. Of the last, or the organic elements {elementa organica)^
we shall speak hereafter : the first question is — ^what are the proxi-
mate constituents? {partes constituentes proxim^je). They are called
Tissues {Teloe).
BiCHAT was the foimder of the science of the organic tissues in
man, and named it General Anatomy. In the investigation of the
tissues he had recourse to chemical reagents, to maceration and
partial decomposition in water. He described each tissue according
to its physical and chemical properties, its physiological phseno-
mena and its morbid changes. After him this science was ad-
vanced by Meckel, Heusinqer, and BicLARD^ in the same spirit.
Within the last ten years it has received a totally different direction
through microscopic research. By means of it General Anatomy
has become, for the most part, Microscopic Anatomy. Our immortal
Leeuwenhoegk, about a century and a half ago, had collected
much material which remained almost unused; until, in our day,
with the improvement of the compound microscope, a more general
interest in such enquiries has been excited, and the importance of
the knowledge of the minutest organic constituents to accurate
^ Here it may suffice to refer to the well-written aod suocinct manual of the last
named author : EUmtM (TAnatomde giniraU (!■« Mit. Paris, 1827. 8vo.)
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10 INTRODUCTION.
Physiology been gradually recognised. We cannot omit mention-
ing the names of Erause, Valentin, Schwann, Henle in pre-
ference to many others whom we could willingly refer to in this
field of enquiry*. We must limit ourselves to a short survey,
and will rather enumerate than describe the different Tissues. A
complete description, however compressed, would demand more
space than is consistent with the nature of this manual.
I. Conjunctive Tissue {tela conjunctiva)^ ordinarily Cellular
Membrane or Areolar Tissue^ called also by others Formative
Tissue, occupies- almost everywhere the space between the
different parts, and forms, according to Cuvier's well-selected
comparison, a kind of sponge, which has the same shape as
the whole body which it contributes to form. By long
boiling it is dissolved into glue. It consists of bundles of
threads, and has no resemblance to the cellular tissue of
plants which consists of hexagonal cells. The threads are
long, have a somewhat tortuous course and an extreme
tenuity (about ^^3 millim.)*.
II. Adipose Tissue {tela adiposa). Formerly this constituent
was not distinguished from the former; it was considered
to be cellular tissue containing fat. But it is quite neces-
sary to distinguish between them. This tissue consists of
vesicles or cells, which, compared with other elementary
parts of the animal body, are pretty large (about S'-^n i^il"
lim.). Fat is, with the exception of some constituents of
milk in the class mammalia, the only non-azotized substance
of the animal body and, like most vegetable constituents,
rich in Carbon. Fat, except in situations where it occurs
as a distinct membrane, forms in the combined state a con-
stituent of different tissues and of many animal fluids.
III. Vascular Tissue {tela vascularis). Vessels are hollow cylin-
ders, which contain nutrient fluids that circulate through
the body. To these belong the Lymphatics as well as the
blood-vessels. The larger blood-vessels are composed of
^ Here too it mAy suffice to refer to a single work of eminence. J. Hsrlb, Alff^-
meine Anatomk, Lehre von der Miickun^s-und Formbettandtheilen des memcldichen
K9rpen. Leipzig, 1841. Bvo.
[> A millimeter is about half an English line, or the i4ih part of an inch.]
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ANIMAL TIS8U£S. 11
different layers : the finest vessels (called Capillaries) of a
single uniform membrane, without fibres or other tissue,
but in which lie round or oblong microscopic corpuscles
(cell-nuclei). In the larger blood-vessels, and also in the
lymphatics, the innermost coat is formed of cells, which,
as in the Cuticle, lie side hj side, pavement-fashion. This
coat is covered by others which present fibres partly longi-
tudinal, partly circular. Then comes the most external
layer of conjunctive tissue, which connects the vessels with
the neighbouring parts.
lY. Nervous Tissue (tela nervea). To this belongs in the higher
animals, the brain, the spinal cord, the ganglia and the
nerves: inferior animals have only nerves and ganglia,
which last take the place of the central parts of the nervous
system. The chemical constituents of this tissue are
Albumen and a species of Fat containing Phosphorus.
The nerve-stems and the bundles of which they consist,
are suiroonded with coats of conjunctive tissue, called
Neurilema: dilute muriatic acid dissolves the neurilema:
alkaline solutions, on the contrary, cause the nervous
medulla to disappear, the neurilema remaining alone. The
nerves consist of fine threads, which neither subdivide, nor
anastomose with each other. They are of very unequal
thickness, ^...to ^ millim. and less, especially in the
nerves of sense. Besides these threads there are found
corpuscles with nuclei: these present themselves in the
ganglia and in the grey substance of the brain and spinal
cord. These ganglion-corpuscles are very dissimilar in
form and size, mostly ^...^ millim.
V. Homy Tissue {tela cornea). The parts consisting of this
tissue have neither blood-vessels nor nerves. Cuticle, nails,
hair, feathers, horns and scales belong hereto. They lie on
the surface of the body, whilst a covering {epithelium), resem-
bling Cuticle, lines the inner surface of the mucous membranes
(as of the stomach) and also of the internal closed cavities
and sacs, as well as of the vessels (see above, lU. Vase.
Tis,). The Cuticle, or Epidermis, consists of microscopic flat
cells joimng on to each other like a pavement, and of which
each contains a nucleus. Water swells up the epidermis.
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12 INTRODUCTION.
boiling leaves it unchanged, by concentrated sulphuric acid
it is dissolved gradually, by alkalis readily. Scales, nails,
&c., which consist of this tissue, are secreted by a highly
vascular bed {mainix) in layers. The Epithelium is formed
in part, like the Epidermis, of flat cells : in other situations
these cells are cylindrical, or conical, and stand perpen-
dicularly, side by side, like fibres. In many situations (as
the nasal cavities, the respiratory organs of mammalia, birds
and reptiles, the gills of bivalve molluscs) these conical cells
carry cilia, whose motions had been seen on the surface of
the body of many of the lower animals by the earlier
observers, but were distinctly recognised by Purkinje and
Valentin as a very general phaenomenon of the animal
kingdom only a few years ago.
VI, Cartilaginous Tissu/e [tela cartilaginea) is semi-transparent,
elastic, and mostly of a bluish-white colour. On section it
presents a very smooth surface and looks like a substance of
uniform density. But under the microscope, small, granular,
round or oblong corpuscles are seen in the clearer trans-
parent principal mass. The glue which is obtained fi:om
cartilage by boiling differs in many lespects from the glue
of bone, and was called by Mueller, who first called atten-
tion to the difference, Chandrine {cartilage-glue). This glue
is also obtained from the cornea of the eye, which is com-
posed of many thin layers or plates formed of fibres that
cross one another in all directions. Certain yellow highly
flexible and elastic cartilages contain numerous fibres {carti-
laginea fibro80B)i to this division belongs ex, gr. the cartilage
of the external ear in man and mammalia. Cartilage con-
tains two-thirds of its weight of water. In the ash are
found carb. soda, sulph. soda, and carb. lime as the chief
constituents. Here belongs :
Osseous Tissue {tela ossea). The tissue of bone is hard
and opaque, and of a laminated structure. The chief con-
stituents are cartilage, which on boiling passes entirely into
gelatin or common glue: and bone-earth, of which the
quantity increases with the age. The last consists princi-
pally of phosphate of lime, which has a great affinity with
the colouring matter of madder, so that the bones of animals
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ANIMAL TISSUES. 13
fed on it acquire a red colour. Bone may be freed of its
lime by acids (ex. gr. dilute muriatic acid). The cartilage
which remains has in general the structare of permanent
cartilage : the bones also, in the first period of life, corre-
spond to cartilage, and previous to ossification (t. «. before
the bone is hardened by the phosphate of lime) the glue
which they contain is also Chondrine, which is precipitated
by alum, acetic acid and the sulphate of alumina. In the
bones are found small medullary canals communicating
with one another (^...A millim.) which are connected with
the medullary cavities, or the cellular spaces in the middle
of the bone, and give to the bone a streaky or fibrous
appearance visible to the naked eye. These canals are
surrounded by several layers, which lie included between
the other layers or plates that, in the flat bones, are arranged
in the direction of their surface, and in the long bones in a
circular form round their internal medullary cavity. These
medullary canals contain fat and minute blood-vessels.
Between the layers are found microscopically small oval
corpuscles, resembling cartilage-corpuscles, and from which
extremely fine tubules, partly branching, proceed. These
parts, when treated with acids, become quite transparent,
and their granular content is consequently bone-earth.
Vll. MuscuhiT Tissue {tela muscularis). Muscles consist of
bimdles of fibres : the primitive bundles, which consist of
some hundreds of fibres, are by means of conjunctive tissue
(cellular tissue) collected into larger bundles, and these again
into still larger. Muscular tissue belongs to the albuminous
substances. Flesh becomes harder by boiling : on cooling
the decoction becomes gelatinous from the glue into which
the cellular tissue has been changed. If finely-divided
flesh be pressed, a red acid fluid is obtained, which contains
albumen, the colouring matter of blood, lactic acid, salts,
and ozmazom. The red colour of muscles (in animals that
breathe by lungs) is heightened by exposure to light ; some
ascribe this solely to the blood. It is not a common
character of this tissue: in fishes the flesh is white: the
muscles of many articulata are brownish, yellow, or light
red. Muscles are distinguished into two kinds. There are
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14 INTRODUCTION.
muscles with varioose stmctore of the primitive fibres, and
fine transverse stripes of the primitive bnndles. This is
the structure of the muscles of voluntary motion amongst
vertebrate and articulate animals, and of the muscles of the
heart. These fibres are very fine, ^o millim. and less, and
are amongst the finest parts of the animal organism. Other
muscles consist of coarser fibres !55...95o millim. which are
not jointed or varicose, and which correspond to the primi-
tive bundles of the former. In these therefore no transverse
stripes are seen. To these belong the muscular fibres of the
intestinal canal, also the red fibres of the muscular stomach
of birds. Usually these muscles of organic life have a
pale and somewhat yellow colour. Muscular tissue has the
property of contracting, upon the application of a stimulus,
in the direction of its fibres. This irritability [irrttabilitcLs)
is a vital property, and is distinct from the elastic contrac-
tility, which other parts of the body retain even after death.
VIII. Elastic Tissue [tela elastica). This tissue has much resem-
blance to conjunctive tissue, and holds, as it were, an inter-
mediate position between it and muscular tissue. The fibres
are of unequal size (ro'-sSo millim.) and have a serpentine
course : they divide frequently and unite at many points with
branches from other fibres, whence a reticulate distribution
arises. The colour of this tissue is yellow: it retains its
elasticity imaltered by keeping in spirit of wine, or by
boiling for several days. After long boiling it gives a small
quantity of a peculiar glue which in some points agrees with
cartilage-glue. The cervical ligament of mammals is com-
posed of this tissue : also in arteries a layer of elastic fibres
lies between the circular fibrous coat and the external coat
of cellular tissue : in large trunks this layer may be clearly
distinguished as a continuous membrane. The yellow
ligaments on the arches of the vertebrae, and the ligaments
of the trachea, also consist of elastic tissue. But not merely
as separate ligaments or membranes, but also mingled with
other tissues, elastic fibres are met with in different situa-
tions, as for instance, in serous membranes and in skin.
We have here spoken only of those tissues which occur most
generally. We have not noticed Dental Tissue because, whilst we
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ANIMAL TISSUES. 15
treat of the entire animal kingdom, it occurs only partially. Other
tissaes, which ordinanlj leceiTe special notification, may be re-
duced to one or other of the foregoing. Tendinous Tissue belongs
to Conjunctive Tissue, as does that of Skin proper (cortum): to this
also belongs in part Mucous Tissue (in the intestinal canal, &c.)
The Serous Membranes merit a special mention. They serve to
line cavities in the interior of the body, and ordinarily form sacs
which are closed on every side. They, too, belong to Conjunctive
Tissue, and are smooth only on their free surface, which is covered
with an epithelium. This smooth surfEuse secretes a serous fluid.
We cannot admit a proper Glandular Tissue {tda glandulasa), as
most authors do. Under the term Gland Anatomists arrange very
different parts, of which the consideration belongs to special and
descriptive Anatomy. Lymph-glands {glanduhB fymphatica s. con^
globaice) which are found only in higher animals, are round or
oblong bodies of different size, in which one or more lymphatics are
distributed; these tortuous branches are again collected into larger
vessels, which pass out on the opposite side of the gland, to pursue
their course onward: numerous blood-vessels, whose fineness ex-
ceeds that of the lymphatics, surround all these branches. Conse-
quently lymph-glands are only vascular networks, and may be put
on a level with the so-called Retia Mirabilia of the blood-vessels.
In the dass of Glands, moreover, are reckoned different parts of
the animal body which, apart fix>m their coverings, consist of con-
junctive tissue, blood-vessels and nerves, and for the most part have
an internal closed cavity which is filled with a granular fluid.
Such are the Supra-renal Capsules, the Thyroid gland, the Spleen,
the Thymus gland. These ate the parts which Heusinger com-
prises under the name of parenchymatous tissue — under which,
however, he also classes other parts, as the Lymphatics and the
Ovaries. Other authors style these parts Blood-glands (ganglia
sanguineo-va&culosa), comparing them with the lymph-glands {gan--
glia IgmphaHco-vasculosa) ; but since these parts are not distin-
guished fix>m others by their blood-vessels, the comparison is arbi-
trary. Finally, in a more special manner, the term gland is applied
to those parts of the animal body which secrete a fluid that does not
return into the current of the blood. These, in addition to lym-
phatics blood-vessels nerves and conjunctive tissue, have an efferent
canal {dttctus excretaritui) formed of mucous membrane, for the
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16 IXTRODUCTION.
passage of the secreted fluid, which is conveyed into the intestinal
canal or to the surface of the body. This efferent canal receives,
like an arterial trunk, the finer canals which effect the secretion,
and which are covered with epithelium. To such belong the
kidneys, the liver, the salivary glands, &c.
From what has been said, it is obvious that we cannot adopt
that division of the Tissues which an esteemed writer^ has pro-
posed: into simple, constituent, and compound tissues. Doubtless
eveiy muscle contains nerves and blood-vessels, but nerves and
blood-vessels are not on that account constituents of muscular
tissue. According to our view, every tissue is simple, but it may,
either by itself, form special parts, or only in combination with
other parts. The corneous tissue is the only one which comes
under the first head : all other tissues form this or that part, only
in combination with one another : nervous tissue, for instance, does
not by itself form a nerve, but only in combination with conjunc-
tive tissue and blood-vessels. Some of these compound tissues are
distributed generally throughout the whole body, others are limited
to certain parts. To the generally distributed belong conjunctive
tissue, vascular tissue, and nervous tissue: the other tissues are
appropriated to determinate parts of the body and have a greater
self-subsistence, as cartilage tissue, muscular tissue, elastic tissue.
This was the division formerly adopted by Bichat. Other di-
visions of the tissues, founded on chemical research, as into gelati-
nous and albuminous tissues, may have their use in Physiology, but
are not to be considered as anatomical divisions.
The above tissues, then, build up the proximate organic con-
stituents of the animal body. Formerly, when less weight was
allowed to microscopic enquiry in general anatomy, the ultimate
organic constituents in these tissues were neglected : but now their
description forms a part of the description of the tissues themselves.
In this way we have learnt to recognise in conjunctive tissue, in
nerves, in muscles, &c. fibres as the ultimate elements of microscopic
analysis : in cartilage, round or oblong corpuscles : in corneous and
adipose tissues, cells. It maybe asked, whether these organic elements
can be deduced firom one another ; or, in other words, whether all the
^ £. H. Wbbeb in the 4tli edition of F. Hildbbbandt'b ffandbuch der Anatomie
des Mensckm revised by him. Braunschweig, 1830. 8. s. 169 — 178.
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ANIMAL TISSUES. 17
tissues proceed originally from homogeneous elements. Fontana,
and afterwards Tkeyiranus, busied themselves with this enquiry ;
Trevikanus belieTed that he was borne out in adopting a sameness
of organic elements in all parts of the animal body, viz. globules
and thin cylinders (elementary or primitive cylinders) ^. According
to others, these cylinders were by no means primitive, but consisted
of globules arranged in a row: so that only globules, or round
vesicles, remained for the elementary particles out of which, in fine,
all the animal tissues were composed and formed. Subsequent
enquiries proved, as indeed had been already surmised, that these
vesicles were due merely to optical illusion'. Every one, who in-
vestigates the tissues with the excellent microscopes of the present
time, will easily convince himself, that such parts no where exist
as ultimate elements of organic animal matter.
Within the last few years, since regard has been paid in the
investigation of the tissues to their origin and to their development,
the problem has received quite a different treatment. That the
tissues consist of different elementary parts, fibres, granules, cells,
is plain from what has been said above ; but it is another question
whether these parts did not originally proceed from some common
ftmdamental form, of which they are subsequent developments and
modifications. Much had been already effected by scattered obser-
vations, but to Schwann is the distinction due of having esta-
blished the original cellular structure of the different tissues, and, at
the same time, the great similarity between the microscopic struc-
ture of Plants and Animals, of which Dutkochet and Baspail
had already a general notion': our limits do not allow us to pro-
pound his views, to which the name of CelJr Theory has been given,
in detail. We will give an outline of them, in a few words, with
a notice of the modifications which, from later researches, they
would seem to require.
The first elements of organic beings are cells. They have theit
^ See Vermiachie Schrifien cmcUomi9ehen und phjftiologiachen InkaUs von G. R. und
L. G. T&EVISAKUS. 4to. I. Gotfemgen, 1816. 8. 1x7 — 144. Ueber die organiacU Elenunie
der tkiaretcken Kdrper,
> MiLMB Edwabiml Stdkerchei nUeroBcopiqua iur la Mruetwre iiUime de9 tiaut
orffoniqua de» Animavx, Annak$ dea Sc. natur. IX. 1826, p. 362 — 394. PL 50.
> Mtkroakopiache Uhtertwhungm iiber die Uebereinstimmung in der StnUstur und
<frm WackMkum der Thiere und Pjktmen wn Db Th. Sohwavn. Berlin, 1839. 8vo.
VOL. I. 2
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18 INTRODUCTION.
origin in a formless matter {Cytoblastema, germ-substance of cells);
what afterwards remains of this substance may be distinguished as
Intercellular substance (suistantia intercellularis). The cells are
vesicles, and consist of a fine membrane which encloses a fluid often
containing granules. For the most part these cells have a so-called
nucleus, a small dark-coloured corpuscle, lying on the wall of the
cell. In this nucleus a round spot has been distinguished and
termed nucleolus. The formation of these cells seems to proceed not
always in the same manner. According to Schwann a nucleolus
arises first, round this a nucleus is formed as its enyelope, by the
aggregation of granules in the fluid germ-substanoe : at a slight
distance firom this nucleus there coagulates, as it were, a thin mem-
brane, the Cell-wall, which at first is raised, like a watch-glass, on
one side of the nucleus, and afterwards encloses it all round. On
this account the nucleus is considered to be the germ of the cell
{Gtftohlastus) ; when the cell is formed, the nucleus, according to
Schwann, has discharged its office : it is detached and disappears.
The researches of Henle* have shewn that such is not universally
the case, but that in fibrous tissues formed from cells, the cell--
nucleus is changed into peculiar fibres.
Cells when once formed are multiplied by fission, or by the
formation of new cells within those already formed. The parts
then of those tissues, with which we have become acquainted
above, are either cells or fibres which have been formed fix>m cells.
(1) In some tissues the cells, which have been plainly isolated, are
present as elements at a later period, as in adipose tissue and
cuticle ; (2) in other tissues the walls of the cells become thick-
ened, and coalesce with one another and with the intercellular sub-
stance, whilst the cavities remain separate, as in cartilage ; (3) in
others, again, the cavities coalesce, whilst the walls of the cells
that mutually touch, are destroyed or absorbed. Finally, other
tissues, still, exhibit as elementary parts little plates without
cavities, which may probably have existed at an earlier period.
These either join one another in a plane, or range themselves
lengthwise in a row, as in the fibres of organic muscles and of
Conjunctive Tissue. Other fibres may, according to Henle, be con-
sidered as compound cells, i.e. those whose nucleus was originally
^ Hbxlv, AUgemeine AnaUnnie^ s. 188—9.
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THE VITAL FUNCTIONS OF ANIMATE. 19
a cell that has become enclosed by a wall or envelope of later forma-
tion. Muscles, according to Schwann, consist at first of nucleated
cells which range themselves in a row ; the nuclei adhere to the wall, -
and within the tube (of the primitive bundle) are formed the proper
primitive fibres. According to Valentin and Henle, on the other
hand, the primitive fibres are arranged around the row of cells
which occupies the middle of the primitive bundle, and the external
covering of this bundle is a sheath formed afterwards. But these
and other diverging views we cannot here develope more minutely.
If once the fundamental truth of Schwann's doctrine be ac-
cepted, that cells are the original form of animal and vegetable
tissues, then is it of subordinate importance whether this or that
Wew in the case of particular tissues be adopted, and we may sup-
pose, as, for example, in parts which are formed of plates in which
there is no distinction of wall and cavity, that the cells have not
been perfectly formed firom the amorphous blastema, but were
joined together before they possessed a cavity ^
We must here add a-word concerning the blood-corpuscles« They
are flat vesicles, filled with the colouring matter of the blood:
having in mammalia a round, in birds, reptiles, and most fishes, an
oval outline. In man, the mean diamet<sr is about ^ millim. In
reptiles, especially in those without scales, they are larger. In the
firog, for instance, they have the length of three and the breadth of
two human blood-corpuscles. Here a nucleus is present, of wliich the
existence in mammalia is doubted by some writers. The blood-
corpuscles, therefore, are cells : and we may consider the fluid, so
rich in albumen and fibrin, in which they swim and with which,
during life, they circulate {liquor sangmnts)^ as a liquid intercellular
substance of the blood-cells.
The Vital Functions of Animals.
In order to complete the general idea which we ought to form
of the animal body^ we must not stop at the membranes, but must
also look at the structure of the principal organs. We unite organs
Henls, AUg, An4Mt. s. i88, 189.
2—2
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20 INTRODUCTION.
and functions in our rapid sketch : and hasten to place before our
readers a view of the whole.
The functions performed by animals may be brought into two
chief classes. One class comprises the yegetative, the other the
animal functions. The first are so called because they occur
equally in plants, and are also, on that account, called organic
fanctions. The last are peculiar to animals, and therefore are
called animal ftmctions.
To the organic Actions belong Nutrition in the widest sense,
and Propagation. To nutrition belong three systems : namely, that
of Circulation, that of Assimilation, and that of Secretion. Respi-
ration is a part of the system of secretion : for the object of respi-
ration, like that of secretion, is the elimination of effete matter, its
volatilization, or its separation in a more fixed form : and both, in
this way, support that unceasing interchange of matter by which
the circle of vital phsenomena is characterised.
By means of these fdnctions, which together are comprehended
by the name of Nutrition, the life of the individual is secured and
provided for. Other fimctions have reference to the life of the
species, and ensure its existence after the death of the individual.
These functions constitute Propagation, of which a part are dis-
charged by the male individual, viz. the secretion of the impreg^
nating fluid {semen) ^ and its conveyance to germs capable ot
development. These germs are prepared and protected by the
female individual, and on the union of these fdnctions depends the
being of the Embryo, the development of which is the final purpose
of propagation.
To the animal functions also belong three systems: viz. the
nervous system, that of the organs of sense, and that of the organs
of motion.
The food, when solid, is comminuted by means of the jaws and
teeth, or, when fluid, is imbibed. It is then conveyed into the in-
testinal canal, which ordinarily has an expansion called the stomach.
Here and at other parts of the intestinal canal different solvent
fluids are secreted for assisting the conversion of the food. The
nutrient part of the food is thus separated firom the rest and taken
up by the surface of the inner wall of the canal consisting of
formative tissue : the remainder is rejected afl unfit for the support
of the creature.
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THE VITAL FUNCTIONS OF ANIMALS. 21
The tube in which this first process of nutrition is effected, is a
continuation of the skin. In some very simple animals, where the
whole bodj is composed of a homogeneous mass (ex. gr. in Polyps)
there is properly no special intestinal canal. The body is simply
excavated, and the internal surface has the same structure as the
external. Such creatures may be turned inside out, like the finger
of a glove, without dying in consequence : nutrition can proceed
undisturbed. Such animals are entirely intestinal canal, independ-
ently vital stomachs. The external skin also corresponds in func-
tion with the surface of the canal. The skin has the function of
Imbibition, which may be compared with absorption by the intesti-
nal tube : and on the entire internal surface of the intestinal canal
there is evaporation, which corresponds to that of the skin, and
with the diminution of this increases.
In some very simple kinds of animal there is in the intestinal
canal only a single opening, which allows the food to enter and
the refuse to escape. In the rest the two openings are separate.
The Chyle, or nutrient juice which has been produced by
digestion, is in many animals immediately poured into the forma-
tive tissue of the entire body, and so serves for the nutrition of the
different parts. In others it is mixed with a nutrient fluid of higher
rank, the blood, which circulates in a system of vessels; this
motion. is called Circulation. The vessels which carry the blood
towards the parts are called Arteries : those which carry back the
blood firom the parts towards the center of the circulation are called
Veins. This motion is ordinarily assisted and regulated by one or
more muscular organs, called Heart. But the chyle is not suflicient
to renew the venous blood and render it fit for the nutrition of the
parts. It must be brought in contact with atmospheric air, and so
be submitted to change before passing into the arterial stream*
This fimction is called Bespiration, and the mechanism for it is in
different creatures so variously contrived, that it is oftien difficult to
harmonise such variety with the poverty of our language, accus-
tomed to include every form under GfiUs and Lungs. In the case
of Lungs, the medium that serves for respiration, mostly air, pene-
trates the cavities whose external surface is bathed with blood. In
the case of GUIs, the medium, here mostly water, does not pene-
trate within the tissue, but only bathes the surface on which the
blood-vessels are spread out. Gills have very different forms, as of
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22 INTRODUCTIOX.
Plates, Leaves, Threads, Twigs, &e. Many simple and imperfect
animals breathe by means of the skin. Others, which have either
no circulation, or none that is perceptible, have Air-Canals, i.e.
such respiratory organs as convey the air through the entire body
to the nutrient fluid.
The nutrient fluid which has thus been separated from the food
and changed by means of respiration, is now fit for the nutrition
of the parts. How that nutrition is effected, so that every part
receives from the common fluid that which is requisite for its
support, is not known. Here we can only conjecture : and if any
one chooses to call it a chemical affinity he is at liberty to do so,
if he merely means that he is contemplating living creatures whose
organism has a determinate chemical composition, and so does not
forget that he has given a name to the process, but has not ex-
plained it.
Besides the glands which separate from the blood fluids for the
internal economy, as the Liver, &c., there are others which separate
constituents that must quit the blood in order that it may become
more pure, or in order that the due proportion of its constituents
may be preserved. Thus the kidneys secrete urine, the skin watery
vapour, &c. Sometimes a secretion is a means of defence, as is the
case with the Ink of the Cuttle-fish, and with the offensive exhala-
tions of many animals, which thus repel their enemies or are avoided
by them. Bightly to estimate all these secretions we must never
forget that an animal is a whole, and that the secretion of this or
that fluid, though it may be performed by an individual organ, is
still under the control of all the other organs, and of life, which
combines them all.
Propagation, which also belongs to the vegetative life, has the
following organs for its instruments : the ovary {ovarium), by which
we imderstand the site and the coverings ^of the eggs and the eggs
themselves, conjointly ; the oviduct (oviducfvs) or the tube, through
which the eggs, that have been detached from the ovary, pass
onwards : the tUerus, a residence for the eggs during their develop-
ment, and the vagina along which they pass to leave the body of
the mother. Li the case of two sexes, the male (by means of
glands named teaticult) secretes the seed {sperma) which fertilizes
the germs, and effdcts their development. Penis is the name of the
part, which, in some animals, conducts the seed into the vagina of
the female.
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THE VITAL FUNCTIONS OF ANIMALS. 23
With respect to the animal life — a perfect sensation appears to
be possible only through a Nenxms 8tf9tem. This nervous system,
in the higher, or more perfect animals, consists principally of the
brain and the spinal cord. The larger the mass of the brain is in
proportion to the nerves, the more perfect appears to be the develop-
ment of the intelligence and mental fecolties of the animal — a law
that was first discovered by* the celebrated Scemmebino. In pro-
portion as we descend to the lower animals, the nervons masses are
more dispersed and removed from one another, and in the lowest
femiUes of the animal kingdom no traces of a special nervous
system remain.
The Head is that part of the body which includes the brain and
the chief organs of sense. There are five senses, of which Touch
{taehut) appears to be the most widely difiused through the whole
animal kingdom. The seat of touch is the skin, the general
covering of the body, which is everywhere interwoven with nerves.
The nerves of the skin are lost, with their little twigs, in its
middlemost and very dense layer. The ends of the cuticular nerves
are covered and protected by the cuticle, and in many places by
other external insensible parts, as scales, hair, &c. In the ijrgan €f
Taste, the twigs of the nerves of taste pass into the soft papill» of
the tongue, and end there. The twigs of the Olfactory Nerve are
spread out upon a mucous membrane (the membrana Schneideriana) :
the continuation of the medulla of the Optic Nerve forms the Betina,
which Physiologists determine to be the seat of vision. Lastly, the
most simple form of the Auditory organ is that of a sac filled with
fluid, in which there float, as it were, the soft and delicate termina-
tions of the auditory nerve. From all this it appears, that the
general form {fypus) of an organ of sense is to be sought for in a
nerve whose terminations form a delicate mass suited to the recep-
tion of external impressions. But in each particular organ of sense
the proper nerve of sense is only capable of a determinate action*
The auditory nerve is only susceptible of sound, or rather, every
stimulus which affects it is perceived only as sound: the optic
nerve recognises no other impressions than those of light Such,
at least, is the case with man and the higher animals: and one
o^an of sense can never 8iq>ply the proper office of another. Im-
I»essions are conveyed, by means of the nerves, to the brain or any
other nervons center.
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24 iNTRODUcrnoN.
Thus nerves are the messengers by which the mind receives
information of the external world {nuntii rerum).
But the nerves are equally the ministers of the will, which by
their assistance is able to act upon the muscles. By Muscles are
understood those active organs of motion {organa motua activa)
which are fixed to other parts, as their point of resistance, and
these last are called passive organs of motion {organa vnotus paasiva).
The harder fibres, which serve for the insertion of muscles, form
Tendons, of which the colour in animals with red fiesh, as in man,
is white. In many animals the muscles are inserted into the skin,
or into certain hard portions of the skin, as in Insects, whose hard
and often homy coverings supply the place of a skeleton in that
respect. A skeleton is, properly, a connected whole of internal
passive organs of motion — cartilaginous or bony, and these serve
not only for motion, but moreover, and indeed especially, for the
protection of the most important parts of the nervous system, the
Brain and Spinal Cord. The skull (for the protection of the Brain)
and the Vertebral Colimm (which encloses the Spinal Cord) must
therefore be considered as the principal parts of the skeleton, of
which ribs and limbs are only appendages: in this simple condi-
tion is the skeleton met with in the Larva, for example, of the
Frog.
Development of Animals.
How the expression imperfect Animal is to be understood.
We have attempted to give a general idea of the organs which
compose the animal body. But these organs are by no means
found in all animals. Only in the more perfect animals is the
structure thus complicated. When from these we descend in the
animal scale, we perceive in the long series one instrument after
another gradually decrease in magnitude and development, and at
last entirely disappear. In Polyps {hydrce) nothing remains but
the Intestinal Canal. The entire animal forms a blind sac com-
posed of a single tissue, and all the vital functions which it
performs are effected through one and the same gelatinous mass.
Finally, in some Infusories we no longer perceive even an intestinal
canal — nothing remains but an homogeneous gelatinous body, whose
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DEVELOPMENT OF ANIMALS. 25
surface appears to discharge the fnnctions of absorption and nu-
trition.
A gradual course of development, similar to that observed in
the animal series, is also pursued by the embiyo of the more
perfect animals. The whole Life is Metamorphosis ; and there are
animals in which the change of form is so great and so remarkable,
that it does not escape even the eye of the multitude. Thus, for
instance, a Caterpillar is changed into a Butterfly : a creeping, dull,
voracious creature into one that flies and runs, and scarcely takes
any food. In the same way the metamorphoses of Frogs are
notorious. But there are other animals whose metamorphoses do
not occur in so striking a^ manner, but are principally limited to the
earliest periods of life. Every animal is slowly developed, and
becomes more perfect as new organs are added to those already
present. But this idea must not be so apprehended as if a Mam*
mal, for instance, had been at first an Infusoiy, then a Polyp, a
Medusa, afterwards an Insect, a Fish, a Bird, &c., as some express
themselves ^ This would be as extravagant as it is unfounded:
but properly, as we conceive, many modems assume that all the
organs in different periods of life do pass through a development
and metamorphosis, and that the structure of a perfect animal, in
its foetal state, is more simple, and corresponds with that of the
lower animals of the same Type to which itself belongs. Thus the
first rudiments of all vertebrate animals are similar, and the history
of the development of the Chick may illustrate that of Mammals in
the first periods. This is more than a phrase without proof: rather
is it the result of very numerous observations — for instance, those
on the Brain and the Heart in the human embryo — and we shall
find it confirmed by firequent instances in the course of the present
work.
We have already on various occasions made use of the terms
'imperfect' and 'perfect' animals, and shall have to use them
often. But since every animal is perfect in its kind, the term
^ How this gradual progress of the embryo iliroagh the diflforent gnwiatioiis of the
animal kingdom is to be understood cannot here be further particolarized. Compare
hereon C. F. Knuam, Ueber du VerhHUnUie der argamteken Krdfie wnter emander
in der Baht dor venehiedenen Organiaationen. Tubingen, 1814. 8vo. s. 38. The differ-
ent works of Cabub, Tisdemahit and J. F. Msokxl supply many examples of the
apj^cation of this position.
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26 INTRODUCTION.
requires some explanation. By perfect animals we understand those
that, in the number and importance of their functions, and in the
complicated structure of their organs, make an approach to Man :
whilst those are called imperfect' whose simple organisation, and
less numerous fanctions, remove them from that perfection of which
Man supplies the pattern." In this sense, as I conceive, the expres-
sion may be well defended. Aristotle says that in all other things
we must proceed just as we do in the investigation of coins, com-
paring them individually with those which are best known to us :
but man is necessarily the best known to us of all animals^. Let it
be added, that Man is in fact the center of organisation to which
the animals, like rays, may be considered to converge — ^and so is the
union of what is most perfect and most beautiful in them all'.
Hence animals which have a resemblance to man are, not without
reason, styled perfect.
On the art of Classifyiry (Taxinomia).
Such conceptions become still clearer by unfolding the art of
Classifying. Classification and systematic division are indispensable
in Natural History. How innumerable are the species of animals
which are scattered over the surface of the earth ! Each of these
species has its country, its determinate form, its peculiar properties.
How shall we attain to all this knowledge: how shall we turn
to account the observations of earlier writers, how learn to what
species they refer? how can we, in fine, communicate our own
observations to others, xmless we make use of a classification?
Classifications then are as old as the study of Natural History, and
their difference is to be sought in their more or less scientific found-
ation and plan. — By means of its systematic arrangement the study
of Natural History obtains a more extensive influence upon our
entire scientific cultivation, and in this respect it cannot be suffi-
ciently recommended to young persons, in order that they may
* "Qawtp yiifi wofda-futra irfi6t t6 oAtois Hojttw yviapifuixrarw ^Ki/Jtdi'oua'tPf ovrw Hi
Kol A» Totf dXXocf. '0 9* HarOpiinrot rtaif {ibiaif yinapifiuiyraTw iffup i^ d^ymft ^9rip,
> See J. G. Hebdsb'8 Idem zarPhUonphie der OuekichU der Mentchen, Carliruhe,
1794. 1 Tbl. B. 100 — 108.
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ON THE AKT OP CLASSIFYING. 27
accnsliiBi fimnMolYes to strict order in all their other branchea of
stady.
The foundation of all Zoological division is the Species. By this
is understood the assemblage of all the individuals which have more
conformity to each other, than to other similar creatures; which, by
means of mutual impregnation, can generate prolific individuals and
propagate themselves by generation, so that it can be inferred from
analogy that they all sprung from a single pair. By specific cha^
racier is understood the collection of all the characteristics which
are shewn to be permanent. Those characteristics, on the other
hand, by which different individuals of a species vary amongst
themselves, and which are attributable to deviation from species, are
called varieiies.
The causes of varieties consist in the influence of external cir-
cumstances, and in the mixture of other similar species. Differences
from this last cause are called Hybrids. The form is here a com-
bination of the two parents.
Such Hybrids appear to be limited, £»bulous stories apart, to
those species which have great mutual resemblance. They are in
general not prolific — not able to continue their race. They occur
therefore, beyond doubt, in a state of nature extremely rarely, and
are rather the consequence of the constrained state of servitude in
which our domestic animals exist. This cause therefore is not of a
kind to disturb the regular course of nature and to endanger the pre-
servation of the species. On. the other hand, varieties produced by
the influence of external circumstances, by climate, difference of food
and mode of life, are able to engender young that are prolific. But
they do not suggest any doubt that we ought, perchance, to receive
them for species. It must moreover be remarked, that those varieties
of ordinary species which on account of the pliancy of their organi-
sation and their tenaciousness of life are able to live in every climate,
and appear, for the most part, to have followed man over the entire
surface of the earth, are the most striking and the most numerous.
By Genus is understood a second group formed by the xmion of
like species, as the species was formed by that of like individuals.
Species which in general have a striking resemblance in their orga-
nisation, form a genus. The idea of genus is so natural that we
meet with traces of it even in the language of children. Still all
genera are not natural. Many of them have been formed upon some
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28 INTEODUCTION.
resemblances of species in one or anotber characteristic arbitrarily
selected in disregard of the general impression of the external form,
and in neglect of the precept of the immortal Linnseus that ' Charac-
ter does not make Genus ^' When a species deviates very much
from all the others, even firom those most resembling it, then a sepa-
rate genus must be made of it. Hence there are genera which con-
tain only a single species. The characters of a genus must be
common to all the species contained in it, and can only be drawn
from a comparative study of all those species.
This is the place to say a word concerning the Nomenclature of
animals. LiNN-ffiUS was the first who gave to every object in
nature a double name : thus the Lion, for instance, is termed Felis
Leo, the Dog Cants famiUaris. The first of the two names (Jelis,
cants) is that of the genus, and therefore common [nomen genericum)
to all the species which belong to that genus. It must be a noun
substantive. Different rules have been laid down for the formation
of names: but to expound them would lead us too far away. Of late
years, after the example of the Botanists, the names of persons have
been adopted for the generic name, as BoneUia, BoUenia, Dorthesia,
Desoria ; but this is much more usual in botany. The second name
is the specific name, as Leo, jumiliaris ; it is either a substantive or
an adjective, and in the latter case must agree in gender with the
generic name. By itself it has no meaning, and indicates nothing
until joined with the generic name^. This double name has thus
an intimate connexion with the Idea of G«nus.
Grenera again, after a similar manner, are grouped together and
formed into Orders, and these again into Classes. We may
reverse the proposition and say that the Animal Kingdom is first
divided into Classes, then into Orders and Genera, which last
contain the Species.
We have now been taught to recognise the chief divisions.
An arrangement which teaches us to find with ease the names of
animals is called a System : which, according to Cuvier's apt com-
parison, is a dictionary, but with this difference, that here the
* "Character non £sMdt genus.**
' It 18 the same with the family names and the prsenomens of persons. Hie first
indicate a family, the last acquaints us with a particular suhject of the family ; only
their order is reversed : i. e. the baptismal name is placed first, and after it the family
name.
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ON THE ART OF CLASSIFYING. 29
pioperties serve us for finding out the name, whereas in ordinary
dictionaries the known name serves to acquaint us with the pro-
perties. That a system may serve its purpose, and supply an easy
means of finding the name, it must be artificial, i. e. it must be
taken firom a single system of organs and their differences. The
characters should be easy to find out, and be borrowed from ex-
ternal parts. An example of such an artificial system is the
sexual system of Linnseas. In the animal kingdom we have no
such artificial system: most of the systems are mixed; neither
entirely artificial nor entirely natural.
For there is yet another kind of systems, called NoOural systems
(Method): of which the chief object is, not so much to find the
names readily, as to unite in an unconstrained manner those
natural products which, in the greatest number of respects, corre-
spond. They are founded, not on a single organ or system of
organs, but on the whole structure. K an object be seen only on
one side, on the north or south, east or west, just so many partial
representations of it will be obtained as there are points of view :
but he only who observes it in all directions is able to form a
judgment of its nature and being. This is the advantage of a
natural method over artificial systems : it does not forget the center
in the circumference, but comprising all the parts and properties
of animals in its estimate, it allots to them a place in the arrange-
ment according to their structure and to the importance which
belongs to them in the economy of nature, and so combines them
in a great organic tohole^.
A perfectly natural classification has not yet been discovered:
but we must continue to search after it, and to collect its scattered
firagments. It is, according to LiNNiEUS, the first object and the
last of the hopes of the Botanist : it ought to be no less so of the
Zoologist*. We please ourselves with the reflexion that we have
approached nearer to this goal, now that men, especially in our
century, have begun to investigate the internal structure of animals
with the same curiosity and the same zeal with which, in the last
century, after the example of Linnaeus chiefly, they studied the
1 See J. Spix, Oeachiehte und BewiJmUmg aUer Sffttme in derZootoffie. Nttraberg,
i8ir. 8vo. 8. 8 — II.
* PkiUmophia hoUmira, § 77.
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30 INTRODUCTION.
external fonn. Even Linn^us himself has declared that a natural
classification of animals is indicated by their internal structured
But if this natural system were quite perfect, it would not merely
be a register of animals or a large lexicon, but a true image of the
animal kingdom and a short survey of the entire science. The
more nearly the science approaches this end, the greater will be its
perfection.
We must here, in few words, mention some of the systems
which have been proposed in Zoology.
Animals may be divided into Glasses in different ways, and the
differences amongst individual zoological systems are very remark*
able. Aristotle divided animals into those that liave blood
(Imufia) and those that have not blood (jbuifui), and distributed these
two chief divisions into lesser ones. Pliny founded his division
upon the different elements in which animals reside, and distin-
guished these as Terrestrialy Aquatic, and Volatile animals. It
would carry us too far firom our object to enter more fiilly into
these and other early attempts at classification. But we must not
omit to notice the system of Linnaeus, who threw a new light on
every department of Natural History^.
In his primary division of animals LiNN-fius was a fol-
lower of Aristotle : he named however those animals which
Aristotle called bloodless, white-blooded : whilst of the rest the
blood is red. The basis of his further division is taken firom the
Circulation of the Blood. Here follows a sketch of his system.
Heart with 2 ventricles and i yiviparoiis I. Mammalia.
2 auricles ; warm, red blood I oviparous II. Birds,
Heart with i ventricle and , with lungs III. Amphibia.
I auricle ; cold, red blood ) with gills IV. Fitkes.
Heart with i ventricle, no i with antennae .... V. Ifuects.
auricle ; cold, white blood \ with tentacula . . . VI. Warms.
1 "Divisio naturalis animalium ab interna structura indicatur.*' Systtm. Nalur.
Tom. I. p. 19. Ed. XII.
" Carolus Linn^ub, bom at Stenbrohult in Southern Sweden, 1707, died 1778.
Comp. on his momentous life Riohaed Pultkney, A general view of the Writings of
LinwEus, 2nd edit. London, 1781. D. H. Stoyrr^s Leben des Hitters Cari von JAnn£,
2 Thle. 8. Hamburg, 1793. Egenhdndiga Anteekningar of Carl Link^db om ng
Vdf, med anmerkningar och tOldg, Upsala, 1823. 4to. (this is transited into German:
C. VON Ltnne iAer sick sfihnt, &c.) The first edition of his Systema Naturae appeaml
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ON THE ART OF CLASSIFYING. 31
Thus LiNN^us adopts six classes, of which the differences,
thongh founded on internal structure, have still reference to the
function of a part which is hj no means common to all animals.
The nnmber of LiNNJSUs' Insects and Worms that have no heart
at all, is in fact at least as great as the number of those in which
it is present. The so-called Zoophytes, and the Intestinal Worms,
have no heart : in many animals vessels are already present before
a heart can be found: insects which undergo metamorphosis have
only a doubtful rudiment of a heart (the so-called dorsal vessel).
The physiological importance of the heart is consequently not of
that kind that it should be considered indispensable in the animal
economy, as appears to be tacitly assumed in this arrangement.
Moreover, in Amphibia the heart is provided not with one auricle
only, but with two, and many of the Worms have not merely a
ventricle, but also an auricle. Still the first four classes are so
truly characterised and so firmly founded in nature, that we may
well wonder that they were not in all times recognised, and not
earlier formed. That clear smd accurate insight which charms
every one, and that simplicity to which all flatter themselves they
can attain in their own province, are the true characteristics of
genius. Linnjsus was less happy in his two last classes, as we
will shew more particularly. Moreover, it is not commendable
that the division is founded upon a single organ or system of
organs, as, for instance, those of circulation. In this way, indeed,
artificial divisions may be obtained, but not a natural method.
(See above, p. 29.)
The arrangement of LiNN^us, in these last times, has under-
gone various alterations, when men began, after the example of
Camper, Pallas, Poli, and others, to investigate the internal struc-
ture of animals : in which investigation Cuvier*, the first anato-
mist of our age, by his incomparable achievements was especially
at Leyden in large folio in 1 735 : it consists of three tables, each containing one of the
three kingdoms of nature with some notices appended. He begins with the Mineral
Kingdom and ends with the Animal Kingdom.
^ George Leopold CHBiTiSK Fredsbio Daoobert Cuyieb, b. at Montb^ard
14 Ang. 1769, d. at Paris 13 May 1832. See my biography of this celebrated contem-
porary in the B^dragen tot de Natuurhundige Wdentchappeny vitgegeven door H. O.
VAN Hall, W. Vbolick^ en G. J. Mulder, vn. 1832. p. -298 — 333. Very important
for the history of CuYiEB's education are the letters addressed by him to C. H. Ppaff,
the friend of his youth, lately published at Kiel by ProfesRor Behn.
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32 INTRODUCTION.
distinguished. CuviER and Lamarck at first divided animals
into two large primaiy classes: those which have an internal
skeleton, and those without a skeleton. They called the first, in
as much as the principal part of the skeleton is the Tertebral
column (see above, p. 24), Vertebrate animala^ the last Invertebrnte
ammah {Aninumx verUhris et animaux sans vertibres). Vertebrates
have moreover red blood, whilst the division of Invertebrates have
either no blood or blood that is white. This i&ct was known to Aris-
totle, who ascribed a spine to all animals that have bloods Having
made these great divisions, they adopted four classes of vertebrate
animals, the same as those already established by Linn^us, except
that they called his amphibia Reptiles^ or creeping animals, a worse
appellation. But the number of classes of invertebrate animals has
so greatly increased that instead of the two classes. Insects and
Worms, first five*, and afterwards by Lamarck', twelve classes,
were adopted.
It would carry us too far away to discuss particularly all these
divisions, and the modifications which were introduced into them.
It is sufficient for our purpose to remark, that attention was now
no longer directed to one organ, or one system of organs, but
to the whole organisation. We propose to follow in our work
Cuvier's last arrangement with some modifications, but in a
reversed order: whilst he descends fi-om man to the less perfect
animals, we, after the example of Lamarck and others, will begin
with the least perfect animals, and ascending gradually to the
more composite, will conclude with man. This course is the
most advantageous for Physiology. If we would understand the
physiological value of an organ, we must investigate its gradual
development,
CuviER, in his latest works, has followed a general division of
the animal kingdom, which takes the place of the two great
primaiy classes of Vertebrate and Invertebrate animals^. He per-
^ Hirra t& Jim, &ra heufiA icrw, Ixci ^x^' ^ dorwji;, i^ ix9M6Coiii, ffitt. Anim,
Lib. III. cap. 7.
* See G. CuvnER, Tableau SUvMntaire de VHUUnre naturdle des Animanx, 8vo.
Paris, an. 6 (1793).
> Db Lamabok, Hid, Nat, de$ Animaux sant verUbr9$. 8vo. vn. Tom. Paris, 1815
—1831.
* Sur un nouveau ^rapprochmerU d itahUr Ub danu qui eompotetU U R^gne animal ;
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ON THE ART OP CLAflSIPTING. 33
ceived tiiat the invertebrate division had only negative charac-
ters: these animals so greatly differ amongst themselves^ that in
order to say any thing precise of their organisation, they mnst be
spoken of specially as Soft Animals, Insects, or Zoophytes. He
adopted, therefore, in the place of the division of invertebrate ani-
mals three others, and thns divided the whole animal kingdom into
fonr large gronps, whose characters we here snbjoin.
I. Vertebrate Antmala {Antmalia vertebraia). The central parts
of the nervons system, the Spinal cord and the brain, in these
animals lie on the dorsal surface, in a bony or cartilaginous
cavity formed by the vertebral column, of which the more
developed anterior portion is called the skull. The limbs,
when present, are never more than four in number. The
muscles cover the bones, and are inserted into them. The
mouth has two jaws placed horizontally.
n. MoUuscs {Animalta moUusca). The central parts of the ner-
vous system consist of ganglia, of which usually one, that
from which the nerves of sense arise, lies above the oeso-
phagus, whilst the rest are differently distributed on the
abdominal surface. This nervous system lies in the same
cavity with the viscera, enclosed by the soft skin into which
the muscles are inserted.
III. Arttculate8 {Animalia articulata). The central parts of the
nervous system consist of ganglia, of which one lies in the
head above the oesophagus : the rest in a row in the middle
of the body, on the abdominal sur&ce, and are connected by
two nervous cords. Tie covering of the body is divided
into rings, and has different degrees of hardness : the muscles
are placed within these rings, and are affbied to them.
When limbs or feet are present, they are usually six, fre-
quently more : when the mouth is armed with jaws, they are
placed sideways, and arranged in pairs.
rV. Badiatea {Antmalia radiata). A special nervous system is
not always present : when found, it appears as a ring, near
the mouth, surrounding the beginning of the intestine,
Ann. du Museum xiz. iSis, p. 73—84. Afterwards in his classical work entitled,
Le BigM animal ditlribuS d^aprU $on arganitaiion, iv. yoL 8yo^ Paris, 181 7, (and md
edition v. roL Paris, 1839, 1830).
VOL. I. 3
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34 iNTRODUC?rroN.
and from this ring the nerves proceed, like rays, to the
periphery of the body. The entire body presents a radiant
form, for the similar parts are not arranged, as in the Articu-
lates, behind one another in rings, but beside one another in
a plane. When muscles are present, they are attached to
the external, sometimes calcareous, covering of the body.
Let it not be imagined that this modification is insignificant :
and that the invertebrate animals might very properly be opposed
to the great division of vertebrate animals, and be afterwards split
up into three sub-divisions. Such symmetrical separations are
usually deceptive, and can only be of real service when the two
groups are of equal rank, and are distinguished by positive charac-
ters. Beyond doubt, all natural bodies, for instance, are either
AnimalB or Non-^nimala : but who, on that account, would think
of separating these bodies into an Animal Kingdom, and a Non-
animal Kingdom? and the Nonranimals again into Plants and
Minerals ? Of like value was, in my opinion, the separation of the
Animal Kingdom into Vertebrate Animals and Invertebrate Ani-
mals : the latter division meaning only " other than vertebrate
animals;^' it is an indefinite appendage to a defined group, and
contains no general idea that can be contrasted with another general
idea.
But what is especially to be attended to in these four great
divisions of the animal kingdom is this : that they are not so much
distinguished by greater or less perfection of organisation, as by
general form, and by the manner in which the parts respectively
are related to one another. A great variety of tissues, of organs
and of subordinate parts, makes the organisation more complicated
or perfect ; but that must be distinguished from the general form,
from the plan of the organisation. CuviER did not overlook this
truth : and even the name of Fundamental Forms ( Tgpes)^ which he
is careful to use for these four great divisions, indicates the guiding
idea which led him to adopt them. In each type there is a gradual
rise and fall of organisation : we descend, says Cuvier, in the type
of the Molluscs, from the syna to the oyster, as in that of the verte-
brates, from man to the fish. But it must not be overlooked, that
Cuvier did not always sufficiently distinguish the two ideas (the
Type and the Perfection of organisation), and to this it is to be
ascribed that his division of radiates comprises many animals which
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ON THE ABT OF CLASSIFTINO. 36
are not radiant, but are placed in this tjrpe on acoomt of the
imperfection of their organisation alone.
It is to Von Baer especially that we are indebted for a clear
insight into this distinction ^ He considers the classes as sub-
divisions of the fundamental form or type, which differ in respect of
the higher or lower grade of organisation. We may conceive the
matter differently, thus : that every animal class is determined by
two factors, of which one is the type of organisation, the other the
perfisctness of structure. The highest perfectness is in general
linked to the type of the vertebrates : yet how imperfect, how poor
(if I may use the term) the organisation may be, even in a verte^
brate animal, the anatomical investigation of amphioocua lanceolatua
in recent times has taught us. That this little fish is in complexity
of organisation far surpassed by many insects and molluscs, cannot,
on unprejudiced examination, be denied.
In the arrangement, therefore, of classes, we shall lay down as
our basis this idea of type of organisation, and shall thus avoid
separating that which is connected by natural transitions. We
begin with the Badiata, because in this type organisation standis on
the lowest grade, and even the most perfectly organised radiates
are surpassed by a great, nay the greatest, number of Annulates
and Molluscs in complexity of organisation, variety of functions,
and multifarious enjoyment of life. Whether, after these, we pro-
ceed with the Annulata or the MoUusca, is in a certain sense indif-
ferent. Nature has not formed the creatures in a single ascending
rank. Were this the case, then would a single type necessarily
prevail in them all : yet even in the divisions (classes, orders and
families) belonging to a single type, we are not able to discover
such a simple ascent of organisation. The most perfect fish is not,
by proximate aflSnity, joined on to the least perfect of the reptiles :
nor the most composite bird to the most imperfect mammal. It
was a pleasing dream of the amiable Bonnet^, that all creatures
» BeUr&ffe zur Kentniu der niedem Thiere, von Dr. K. A, V. Ba«b, Nov. Act. Oce$.
X. C. Nai. OurioB. Vol. ini. P. n. 1817. a. 513— J62, especially 8. 739—759; ^^^^^
JSa^nekdvngsgaeku^ie der Thiere. Beobachiung und RefiexUm. 40. i. KonigBberg, i8a8.
0. 207 — 419. In France also similar ideas were afterwards advanced by MiLNX
E^DWABiys. See ese. ^. his remarks in the new edition of Lamabok : ffiMoire NaL des
Ani. a. veH. L Paris, 1835, PP* 335— 337> revised by him and Dbshatbs.
s OontempUaicn de la Nature, (Euvm d^Hiat, NaturtUe et de PkOotophie de Ch.
BovKET. Tome VII. Neufchatel, 1781, 8vo. pp. 51—55, and other places.
3—2
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3d intkoduction.
form an imbroken chain ; that without anj sudden leaps, all hang
together by insensible transitions. But the breaks in the chain
have not been filled up by later observation, rather have new and
previously unrecognised deviations firom it been discovered. It is
not a ladder with uniformly ascending rounds, but rather a net
which may afford us a conception of the multifarious connexions
and the various affinities according to which nature has arranged
her products.
We have already remarked that the vertebrate animals ascend
to the highest grade of perfection of organisation : of them, there-
fore, we may properly consider the different classes last.
In treating of the Animal Kingdom we shall not make use of
Cuvieb's distribution of it into four divisions, fiDu*ther than as a
guiding idea. The Infiisories (exclusive of the Rotatories and
others, which were joined to them on account of their minuteness
alone) appear to form a distinct group, or at least do not indicate
the radiating form by which Polyps and others of the lower animals
are distinguished. We make, therefore, for these simplest animal
existences a distinct Division, naming them, after the example of
other authors. Protozoa. Their form is round or oblong, often not
rigorously determined, but variable during life^.
^ JFvrt great divisionB of the animal kingdom might be establiflhed, and named:
Pr<i*<m<i, Aetinotoa, EeUnoeoa, Malaeoeoa, Spondi/loBoa, We are too indi£Eiarent to the
introduction of new names to propose these except in a note. Under Sctinctoa (from
iierebw, exlendo) we understand those animals in whose oiganisation the elongated type
preYails: they nearly agree with the Artietdata. The other names have in part been
used already, and require no further explanation.
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CLASS I.
INFUSORIES (INFUSORIA).*
These animals are called Injusorxea^^ becauae they are to be
found in infiisions of every kind of organic matter, as well yegetable
as animal. MoreoTer thej live in all stagnant marshy waters, and
even in ronning waters, salt and fresh.
These animals, which on acconnt of their minute size can only
be discovered, or at all events examined, by means of magnifying
glasses, were unknown to the ancients. Our Leeuwenhoeck iSrst
saw them in infusions towards the end of the seventeenth century
(1675). After Leeuwenhoeck, many species of these animals
were observed and described in the last century by Roesel, Leder-
MUELLES, VON Gleichen and others : but more especially O. F.
Mueller, the Danish naturalist, in a work which appeared after
his death, figured many species, and gave a systematic arrangement
of the class. In the present century, Ehrenberg has contributed
most largely to our knowledge of inftisories, and since 1828 has
from time to time published his investigations in the Transactions
of the Berlin Academy of Sciences, and more recently has collected
them into a body in a large illustrated work.
As the discoveries of astronomers allow us to cast a glance into
the illiinitable extent of space of the universe, so the eye, artificially
strengthened, affords conviction of the illimitable distribution of
^ The chief works rdatmg to this class are :
O. F. MuXLLiB, Ammtdcula inftuoria JluviatSia d ma/rwa, H«mife, 1786. 4to.
nie Plates are copied in the BneydopSdie mitkodique,
C. G. Ehbsnbibo, Die If^udontUUerchmg als voQkommene Organmnen, NebH einem
Adas von 64 eolorirten Kupfertafdn, Leipzig, 1838 folio. (G. Yalihtin gave in his
Beperiorwm fBr Anat, und Physiol, iv. Bd. Jahrgang, 1839, '^ detailed abstract of
this work, containing the characters of all the genera and spedes. s. 136 — 18 f.)
JSistoire naiurdle des ZoopkyUs, Ifrfusoires, eomprenasd la Physiologie d la Class\fi-
eaUon de cea AnimoMx, par F. Dujabdik, Paris, 1841. 8yo.
Die Inftuumdhiere auf ikre Enitoidediungsgesekdehte untenuehi wm Db Fbibdbich
Snnjr, ndt 6 Kupfertafeln. Leipzig, 1854. 4to.
> First by Lkdebhuxllxb, according to Ehbbnbbbo.
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38 CLASS I.
life in the direction of the smallest space. Tliere are infdsories
(monads) having a mean diameter of iso'-isio of a line, which yet
live so closely together, that the intervening space scarcely exceeds
their diameter ; a single drop of water measuring a cubic line, if
only one-fourth of its space were filled with such animals, would
still contain 500 millions of them.
This minuteness has misled some authors to designate infasories
as microscopic animals. We cannot allow to this appellation a
preference to that of infdsories : magnitude ought not to supply the
character of a class of the animal kingdom, or a ground of division.
By such an appellation, the union of diminutive species of higher
classes of animals with infusories, often practised by older authors,
would be justified.
In determining what is to be understood by infusories, we must
look to the whole of their organisation : it requires not many words
to shew that the investigation of the organisation of creatures so
minute has difficulties to contend with, which even the best optical
expedients of our time have only partially removed. For, although
every species of infusories be not so small as to escape the naked
eye, yet even these are not bigger than two or three millimeters^.
Of the minute animals that are usually comprehended under the
term inAisories, Ehrenberg's investigations led him to distinguish
two classes, which he named Polygastrica and Botatoria^. The
incontestably greater complexity of structure in the last, the sym-
metry of their form and their resemblance to the type of the Articur
lata, suggested to us, as early as 1834, the propriety of separating
them entirely firom the others — a proceeding now approved of by
almost all. zoologists.
Consequently, we comprise in the class which now occupies us
only those animals which Ehrenberg calls Polygastrica^. We
have not, however, adopted that name, for it rests on the opinion
that the cavities observable in the interior of these animals are
stomachs, which is doubted by many writers: but even if that were
admitted, numbers remain in which no such stomachal cavities are
to be seen. The class, thus limited, contains animalsof very simple
1 A millimeter is about half a line, or ^ of an En^liBh inch.
^ For the literature vid. Sibbold and Stahnidb, Leh/rb. der verffldck. AnaUmie, I.
Abtheilung. Berlin. 1845. ^^o. 8. 7.
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IXFU80SIES. 39
fltmcture. Some, in which no oral^ aperture can be found, must
leoeive nntriment by means of absorption through the external
surfiM^ alone. Others have a mouth, usually surrounded by alia,
by the motion of which a current is produced in the water, carrying
the food which it may contain to the mouth. The subsequent
transmission of nutriment throughout the body is effected in spaces
of a vesicular form — ^which contract again when their contents have
been expelled : they are pushed on by others that hare more lately
come into view, and have motions that cannot be reconciled with
the notion that they possess proper walls. We must rather suppose
therefore that these vesicular spaces are excavations in the gelatinous
tissue of the body. The undigested portion of the food is, in many
ci these creatures, cast off by another aperture* of the body : in
others it escapes through the same opening by which it was received.
Special organs of Respiration have not been recognised. The
external surface of the body appears to be the seat of that fimction.
Still less are there any blood-vessels : perhaps those spaces, which
in many species are seen to contract and expand, and which are
various in form and number, may serve for moving and transmitting
the nutrient fluid which supplies the place of the blood'.
Although no muscular fibres^ are present, these animals possess
the power of motion in a great degree. Some move slowly, others
very nimbly. As organs of motion the greater number have cilia.
In some are produced by contraction all sorts of changes in the
^ [Stbih coDfiiden all ciliated infuBories without a mouth to be larval or emhryotic
forms of other creatures. IHe Infusitnuth, s. i8i.]
' [According to Stkn there is no ready-formed anal opening in any infusory. In
those families which do not reject their excrement by the mouth, it is forcibly pressed
through the integument of a determinate r^on of the body, more or less extensive, and
not sharply defined. After the exclusion, the rupture is completely dosed again. It
is allowable, therefore, to speak of an anal region, not of an anal aperture. Die J^fu-
tiontikiere. p. 34.]
' EmtEKBBBa holds these contractile spaces, or vesicles (?), to be veiicukB temmaUt,
The Uaiadut is with him a structure whose connexion with these vesicles he has not
demonstrated, and which y. Sisbold regards as a nuofeus, whilst he compares the
entire infusory to an organic celL
^ In the hollow pedicle of Vorticdla, and other similar genera, there is a longitudinal
musde which by its contraction effects the npiral retraction of this part. Ehsbitb. Die
Infiuiontth. s. 270. [GzBBMAOK denies that the contractile streak in the canal of the
stem is a muscle : see Sibbold and Kollikbb's Zeitschrtft fOr wieeenecht^Uiche Zooh-
gie, IT. 443. Also see Stun, op. cU. p. 81.]
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40 CLASS I.
form of the bodj and of the short digitiform elongations, by means
of which thej move : others again have filiform, branching append-
ages, that can be drawn in and out. There is no doubt that these
motions are to be ascribed to an internal power : they have alto-
gether the character of volition, for the creatures sometimes retard
their motions, or suddenly stop and again as suddenly swim quickly
away. Infusories make no distinction of day and night : they are
incessantly in motion, and no indication of sleep can be detected^.
Coloured (red) spots have been supposed to be eyes, without
any particular proof: but neither a nervous system nor any distinct
organ of sense is to be found.
The multiplication of these animals is by spontaneous fission of
the body, generally in the direction of the length. In this way
they can increase incredibly in a short time. In a very few multi-
plication by buds has also been observed'.
^ Ehbbnbbbg, Dm Itrfuriontthiereken, b. 539.
* Ez.gr, ID. VarUcdla (early observed by Spallaitzani), we Ehbknb. op. cU. Tab,
ZXV. fig. m. 3 ; in EpUtylia, kc.
[Besides the multiplication of infusories by longitudinal and transverse fission, and
that by external gems, other modes have been brought to light by the labours of CoHir
and BiBiir. They are forms of the encytttng-procas, which SraiK sees reason to believe
to be common to all true infusories. Cdpoda eucuUut does not undeigo fission, but
becomes endosed in a cyst, which in all cases is the secretion of the animal's surface.
Within this it multiplies by successive division, so that a progeny usually four in
number, occasionally eight, arises. Each of these is a special cyst, with its own
external membrane. The original cyst bursts, and the special cysts repeat the same
process, often several times, until at length the content of each special cyst escapes
through the ruptured external membrane as youug Oolpodas. Stsut, op. cU. pp. 15 —
45. Tab. m. fig. i — ^31.
In VorHcdUnei (besides the generation by buds or germs, and by longitudinal fission),
the animal, becoming encysted, is changed into a spherical mass, in which none of the
original oigans can be perceived except the ribbon-shaped nucleus, and a clear space,
which, however, does not pulsate. Processes are sent through the thin covering at the
upper exremity, and the form becomes some one of "EBXRVBVBkQ'BAcineUB, according to
the different genera^ as Podophrya or AcHnophrya. The nucleus, or rather part of it,
is then transformed into an internal embryo that rotates actively, and closely resembles
the germ-progeny of a young Vorticelline of the same species. This process is frequently
repeated in the same Acineta. The progeny may either encyst itself anew, and go
through the same process, or may at once secrete a pedicle, and become an oidinaiy
Vorticelline of the species.
Such is the process in a young Vorticelline. In a full-grown large one it is different.
Here the encysted lx)dy is transformed into a homogeneous mass, the nucleus falls down
into a nimiber (more than thirty) of disci-form bodies, which derive nutriment from a
portion of the liquefied substance of the mother-cyst : another portion of the maternal
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INFUSOttlES. 41
If these animalfl are not propagated by means of eggs, then
neither can they have come into being firom eggs, and the general
law that all that lives proceeds firom an egg {omne vwum ex ovo)
does not apply in this case : a wider meaning, then, than the ordi-
nary one, must be given to the word egg, which denotes ^ part
that requires impregnation before it can be developed, and contains
a yolk with a germ-veside smronnded by an external covering.
To explain the origin of these creatures in infusions, if equivocal
generation be denied, nothing remains but to suppose that the air
conveys infnsories as it does minute particles of duBt, and that the
organic matter, which served for the infusion, fitvoured merely as an
abundant nutriment the multiplication of the creatures^. The
advocates of equivocal generation have not been able to deny the
possibility of this explanation: and that possibility destroys the
force of all their proofs and argumentations : especially when it is
remembered that no space can be so perfectly closed that air cannot
penetrate it, and that even boiling does not destroy every kind of
infusory : for their opponents themselves could not absolutely deny
that infusories were found in boiled infusions which were stopped^
Indeed the constancy of form in the species, which had been over-
looked by earlier observers in their experiments, or not understood,
is irrecondleable with the view that these animal forms are produced
by external forces as a mere sport of chance : but it is not by any
sabstanoe MsomeB & gelatinoas fonn in which the embiyM swim, and by which thay
are Bomninded when the cyst ifi buiBi. When this substMice has been diasolved in
the soTTOunding water. they swim freely away, and change the monad fonn for the
▼ortioeUine. Snnr, op. cU, pp. 50 — 64 and p. 146. Tab. in. fig. $2 — ^41. AJbo vid. GoHir,
Ztkackrifi far Wi»$, ZooL m. b. 360—179. Tab. vn. fig. i^-ii.]
^ [It ifl well known that Infoaoriee are oonveyed by the air : EHsnrBBBa fonnd them
in the dnst borne by the trade-wind : Svxur disooyered cysts of Cfoipoda, of Pk3odina
roaeota^ and of MacrMotm Hufdamdii, on the tenninal twigs of beech-trees that grow
aooo feet aboTe the level of the sea. PkUodnui roseola is the rotifer which gives the red
colour to snow. Yid. Stuh, op. cU. p. 35. He quotes Raohxkhob8T*8 assertion that
if a slip of j^ass be moistened by the breath, and moved about in the confined space of
an i^wrtment, infusories may be seen upon it. ScHMmr's/aAr-fttie^, 1850, Bd. Lxvni.
B. 383.]
> [The experiments of Sohwahn, PocK}Bin)OBrF*8 Annalm, Bd. xu. s. 184, and of
HxLMHOLTX, MuBLUB's ArMv, 1843, B. 453, have satisfiustorily shown that an infusion
boiled so long as to kill any germs previously existing in it> is never visited by infuso-
ries if only such atmospheric air be allowed access as has passed through a red-hot tube,
or sulphuric add, or caustic potass.]
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42 CLASS I.
means neeessaxy to coimect such a conception as this with the tenn
equivocal generation. As long as it is not pretended bj this term
to afford an explanation, but only to indicate that there are some
animal and vegetable species that arise not firom eggs, but, in a
way thai toe are unable to explain, firom the decomposition of organic
matter, so long do we belieye that the expression cannot at present
be dispensed with in Physiology^. The formation of Infusories is
no primary production of organic matter*. Their immediate origin
firom the organic matter of Infusions has never, as we believe, been
observed at the very instant of its occurrence, and probably never
will be. Even in the development firom the egg we never see the
forming, but only the thing already formed. In the case of the
intestinal worms' the same obscuri^ recurs, and the difficulty of
applying the proposition that all living creatures come firom eggs is
but too obvious firom the very constrained and improbable explana-
tions which have been resorted to. The reason why organisable
matter assumes those determinate forms that are distinguished as
genera and species, is altogether unknown : and Physiology is, in
the same degree, unable to explain how it is that in a complexly
organised creature developed firom cells, in one part muscular fibre
should arise, in another nerves, and cartilage in another.
The knowledge which we possess of the geographical distribu-
tion of Infiisories is due to the investigations of Ehrenberq. His
travels in Asia and in Africa have taught us that in different coun-
tries different species, nay different genera of these animals are
found. The species which have the widest geographical distribu-
tion in the northern hemisphere are Monas termo, UveUa glaucoina.
^ Vide note a, iiage 40.
» "Eagi^^ heine Erfakrung, diefiir eine EnUlehung Idtmder K&rper out Stoffa^ der
kbham NatiMr tprdche." G. R. Tbeviranus, Biologie, u, s. a66. In this work may be
found a full account of the earlier observations on this subject, to which the author has
added many investigations of his own, s. 264—353. Although more than forty yean
have since elapsed, the Ubours of Tbbvihanub on this point still retain a great value.
Aa to the green matter of Pbiestlbt, in which transformations of infusories are sup-
posed to occur, this is not exclusively of a vegetable nature, but consists, according to
the exact investigations of later enquirers, of a collection of dead, and in part still living
IirfvMorie»t Chlamidomat pulvUcvltu (Ehbenb. L. i. p. 64), Eugkna viridis (Ehkknb.
p. 1 10), &c.
' [The presence of Entogoa in situations where it was thought impossible they could
be introduced from without is now explained : tfid, notes on that cUss.]
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INFUSORIES. 43
ParamoBcwm ehrymJdSy Colpoda cucuUm: &e last appean to occur
evetywheie {Comnopolite). Ehsenberq found Monas termo both
in stagnant water at the Bed Sea, and in spring-water on Mount
Sinai. Colpoda cucuUus is one of the most constant animals in
artificial In&sions, and was formerly observed hj Leeuwenhoeck
in Infusions of pepper.
APPENDIX
TO THE CLASS OF INFUSORIES.
ON (THE SO-OALLED) SEMINAL ANIMALCULEa
{Spermat'Ozoa.)
Von Baeb first (1827) introduced the name Spermatozoa {Nov.
Act. Acad. CcBs. Leop. Gar. Vol. ziii) : earlier names, as animalia
spermatica, Ceroaria seminis^ &c. hare merely an historical value.
The bodies, for the most part filiform, which have been observed
in the fecundating fluid of animals of almost every class, have as
microscopic creatures been ranked with Infusories: other writers
class them with the Entozoa, a proceeding which can only be justi-
fied in a degree by the locality where they occur ; while, on the
other hand, it has introduced several wrong views. An independ-
ent organisation, in virtue of which they might be considered as
real animals^ has not been discovered. The cause of their motion
is altogether enigmatical.
Leeuwknhobck, the discoverer of the Infusoiies, was the first
also who made obseryations upon these small corpuscles that in
appearance perform voluntary motion in male spermatic fluid.
Tlieir discovery is due to a medical student, named Hajc, who an-
nounced them to Leeuwenhokck in August, 1677 {Phil. Trans.
1678, Na 142 ; comp. Letter 113 in the Sevende Verfolgder Bri^
ven, 1702, p. 65 . . ., or in the Latin edition, CofUinuaUo Arccmomm
NcUura, Operum Torno ui. p. 60 sqq.). Leeuwenhoeck named
these bodies animals, and observed them consecutively in diflerent
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44 SPERMATOZOA.
species of various classes of animala It was his opinion that they
foimed the embryo, and that the female's share in the work of pro-
pagation was simply the reception and nutrition of the male pro-
duct This view of Leeuwenhoeck's as to the office of spermar
tozoa in propagation was afterwards entirely rejected : until, in our
own century, Dumas maintained that they form in ftTiiTnft3a the
foundation of the nervous system (the brain and spinal cord) : to
which view he was led by a certain resemblance of the first rudi-
ment of the embryo (the so-called primitive streak) to a spermato-
zoon (picL Glassique (THistovre ruUtM-eUe, T. viL 1825. p. 221,
article GSniraiion, Annalea des Sc not xii. 1827. p. 443-454).
But it is not founded on observation, and is moreover sufficiently
refuted by th^ fitct that some animab have spermatozoa closely
resembling those of mammals, whilst their nervous system has a
totally different form from theirs.
According to Wagner's investigations, these active molecules are
formed in cells, singly or in bundles : firom w^ch, on bursting of
the cell-wall, they are set free. In insects they are found as fine
threads without a head, or thicker portion: but in most other
creatures they consist of a thicker part, the head, and a very fine
thread, or tail The head is, in diffiarent animals, of a different
shape.
[The spermatozoon of the Batrctchia has an extremely fine mem-
brane attached to its tail in the direction of its axis and throughout
its whole length by one of the sides, the other being firee and wavy.
Thus a delicate undulating border is formed. It was discovered by
Amigi and rightly described by him, and afterwards by Poulet. By
others it was mistaken for a thread surrounding the tail with a loose
spiral coiL Yid. J. N. Czerieak, ZeUsch. /, tmasenach. zooL R ii.
350-355, also von Siebold, ifnd. pp. 356-364.]
The different memoirs and treatisee upon this Bubject with whose history, aa
Ehbenbbbg says, whole Tolumee might be filled, are not noticed by us
that we may not incur a diffusenees unsuitable to the limits of this manual.
B. Wagnbb's Lehrbuch dor tpeaeUm Zoohgie, 2^ Auflage, Leipsig, 1843,
8vo, 8. 10—30 maybe consulted with advantage. It g^vee a full account of
the most important discoyeries of the author and of other contemporary
observers. [This work has been translated into English by Dr Willis.
Comp. the later work of B. Waokeb and R Lsuckabt, Aride Semen in
Todd's O^fdop, o/Anat. and PkysioL YoL iv. p. 849.]
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SYSTEMATIC
ARRANGEMENT OF INFUSORIES.
CLASS L
INFUSORIA-
MiCBOBOOPiC animalculea, often of mutable form, without
nervous Efjstem and yessels, mostly with internal globose cavities,
moving hj means of vibratile cilia, or exsertile processes, without
tme articulated feet.
From a definition all that is uncertain and hypothetical must be
excluded as &r as possible^ Hence we do not call the internal cavities,
which are seen in most Infosoiies, stomacha Our dass, limited
as above, contains several of the Pclype9 gdat/moMx of Cuvisb,
and agrees with the Pclygasirica of Ehbenbebo. Many genera^ as
BiJbcXttafrukf NaniouJUk^ Clogterium, are omitted, because it is highly
probable that th^ belong, as many writers have admitted, to the
vegetable kingdont
OsDEB L Simpltciaaima.
Naked, extremelj minute, external organs none, form persistent.
Family X VtbrionidcB. Body filiform.
Vibrio MuELL. (exclusive of many species.) Gknera: Bcuieriumj
Vibrio, Spirillum Ehrenb.
Sp. Vibrio Uneola MuxLL., Ehbshb. (and M<ma$ termo Muill. t) Muxllbb,
7i^. Tab. VI. fig. I ; Ehbsnbkbo, It^kuunuthierchm. Tab. v. fig. 4 ; in
difierent infamons, fte.
Ordeb II. Ehizcpoda DuJARD. {Pseudopoda Ehrenb. in part).
Animalcules with mutable form, moving by means of multiform
exsertile processes, without vibratile cilia or other external organs.
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46 CLASS I.
Family 11. AmoelxBa. Animalcules naked, emitting and retract*
ing irregular, mutable lobes by continuous motion*
Amceba Ehbenb. {Proteus MuELL.)
Sp. Amaba cUfiuetu Ehbsnb., Volvox CkaoM It,, Proteut diffluens Muell. ;
B0B6XL, Ina. iiL Tab. oi. fig. a— T ;i Muell., Infiu. Tab. n. fig. i . . . i« ;
Ehbenb., Infuaioruih. Tab. viii. fig. zii. A gelatinouB mass, of rounded
fonn, if the entire animal contracts itself on disturbance of the water ;
when the water becomes quite at rest the body extends itself variously
into lobes and processes, which are drawn in again. Bobsbl observed
these parts to be even torn asunder by extension, so that there arose two
anJTnalw (Prop^hgation by spontaneous division). The name Prateua had
been previously given (by Laubenti) to a genus of Reptiles, and was on
that account changed into AnUba* by BoBT, and into Anuj^ by Ehben-
BEBQ. [Perhaps Afnoeba is a temporary state of other forms, as of the
shelled Bhizopods, &c. Yid. LiEBERKUEHir, in Mubllbb's AriMv. 1854,
B. 17, and Ck)HK in Sibbold and Eobllikeb's, ZeU$ckf. Wiatenaehtrft, Zool,
Bd. IT. B. 262,]
Family HI. Arcellina. Animalcules enclosed in a membranous
lorica or calcareous test, partly exsertile from their coyering, and
emitting processes sometimes filiform and branched.
They are small calcareous forms (sheila) divided into cells, found
in sea-sand and in a fossil state in tibe Chalk-formation, and espe-
cially in the coarse tertiary limestone. These miscroscopic crea-
tures occur in incredible numbers, 6000 of them having been
counted in an ounce of sand from the Adriatic sea, whilst an
ounce from the shore of the Antilles contains, by computation, near
four millions. They were investigated at the end of the last cen-
tury by 80LDANI, and in the present by Fichtel and Moll, and
afterwards especially by D*Obbiont, who defined more than 1600
species of them. Until within a few years these bodies were refer-
red to the Molluscous Division, genus Na^iUtus L. {Cephalopoda,
vid the first edition of this Manual, il pp. 107, 108). Beoent
observations, however, consign these Folf/thalamia or Ctllulacea to
a much lower position, near the genus Proteus of MueUer, Although
D'Obbigky has been satisfied by the investigations of Dujabdin
that these animals do not belong to the Molluscs, he still believes
that they ought to be considered as a distinct class of the animal
kingdom (standing between the Polyps and Echinoderms), and
caUs them Fora/mm^feraj the same name under which he formerly
^ BoBT DB St Yxkcent and Dujakdib refer these figures to another species, sup-
posed to differ from Proteus diffhtens by its greater size.
3 DicHonn. dass, d^Hitl, ncctur. i. 1821. p. 261.
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INFUSORIA. 47
described them as MoUiisc& From the end of the last cell of the
shell by one or more openings, or from numerous pores on the sur-
fiuse of the shell, thin contractile threads are extended which serve
for motion. [On dissolving the calcareous matter from the shell of
Hving species, there always remains an organic base of the exact
fbim of the shell with all its pores and passages. This is a secre-
tion effected by the contained animal mass. Sgsultze*.] Ehben-
Biaio thinks that these aniTnals are allied to the Bryozoa (the so-
called Polyps of Flustra, &c) ; his principal reasons against their
arrangement in the class of the Infusories are, that they have no
polygastric intestinal canal, and that there is no other instance of
calcareous shells amongst Infiisories.
On this division compare, amongst other works ; D'Orbioitt, TabUau
methodiqiie de la Claate des Cephalopode», Afkn. det Sc. naL premUre SSrie,
Tom. Vn. i8a6. p. «45— 315. PL 10—17.
DuJABDiK, ObservaHoM nouveUet mr let CSphalopodet nUero$eopiquet.
Ann. des Sci. not,, ieetmde S^rie. Tom. III. 1835. ZooUtgU, p. xo8, X09 ; and
Beekerekee aur les Organiimee tj^^rieurs. 1. Sur la Oromia ovtformit H mtr
let JRhkopodet m gininl. ibid. Tom. IV. Zoologia, p. 343 — 35a. PL 9.
Ehbsnbbbo, UAer die BUdung derKreidrfebm u, detKreidemerfftit dmxK
untichibare Otyanitmen, Ahhandl. der h&nigUek, Ahademie der Wittentek,
tu Berlin, Aut dem Jahre 1838. s. 59—147. Also, Ueber nock tekr
tahlreich Idtende Thierarten der Kreidebildung und den Organitmnt der
Pokfikodamien, &c. Aut dem Jahre 1839. s. 81— 1 74, espedally s. xo6— i la
A. D'OsBiGirr, Article Fonmimfiret, Didumnaire umvertd tTSiet, not,
parCR. D'OBBiomr, Tom.V. 1845, P* 661—671.
Glabk, On recent PoramMfera, Ann, of Nai. Sid, Sec. Series, 1850.
VoL V. p. 161 — 171.
Cabfentkb, Microtcopie Structure 0/ NummuUna, &c. Journal cf the
Qeot, Soc. tfLondony 1850.
WmJAMBON, On the minute ttruOwre of ealoareout ^eUt, fto. Trant,
Mierotc, Society of Zondon, YoL III. 185 1, and Quarierlp Joum. of
Mierote. Science, 1853, No. TV. p. 87.
Ehbeitbkbo, I>at wrhen det untiehUKuren Jddnen Lebent anf der Brde,
1854, Tab. XIX— xxxn. Leipsig, 1854.
Max. Sigmund Schultzs, U^)er den Orffonitmut der Polpthal. ftc.
md 7 UUtminirten Tctfdn, foL Leipag, 1854.
On NoctUuca miUarit, which appears to belong to this diyision, though
it does not emit expansions esctemally but has a moveable appendage
attached, see QuATBiYAais Ann, det tc. not, tit, Sirie Zoai. xre. p. ia6 —
135 PL 5, Kbohn in Archiv f Naturgetch, 185a s. 77 — 81, Taf. 3, fig. «.
It is a chief cause of the phosphorescence in se»-water.
To this femily belong also some fresh-water species.
^ SCHULTZE; op. cU. p. 7.
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48 CLASS I.
A. Body simple {Monoategia D'Orb.)
* Loriea membranoui ur homy.
ArceUa Ehbenb. Loriea scutellate, globose, or hemispherical,
sometimes angnlate, open beneath: the animal emitting processes
variable, plane, obtuse, through the aperture.
These anhnals Uve in fresh water. See Figores in Ehbbitbibo'b In-
fimonHkienhen, Tab. ix. fig. v— Tm ; Dujard, Ii^m, PL ii. fig. 3—5.
Dtffluffia Leclerc. Loriea globose or oval (sometimes sub-
spiral ?), emitting from the terminal aperture processes of the animal
variable, multifidous.
LsoLEBO first discovered these fonns (1815) ; see Note aw la Diffugie,
MSm.duMii$6fim, U. p. 474—478, PL 17. Bp.J>iffi.prote{fortni8, fig. 3. 3 ;
Ehbbnb. If^uticntlh. Tab. re fig. i. The shell, according to Lbclebg,
is spiral, what later observers did not perceive ; it is covered with minnte
grains of sand. — JHjfl. gkbulota Ddja&dik, Ann. de$ Se. not., te Sirie.
Tom. VII. 1837. Zodogie, p. 310, 313. PL ix. fig. i.
Oromia DujARD. Loriea globose, membranous, emitting pro-
cesses of the animal variable, slender, of great length, from a round
aperture.
8p. OronUa omfcrmu Dujabd. Ann. de» Se. nai. 2e Serie, Tom. IV. Zodogk,
PL IX. fig. I. 1, in salt water, amongst marine plants ; — Oromia JUma-
tiUt Dujabd. ibid. Tom. VIII. Zoologie PL 9. ig. « ;—Orwn. omfcrmU
DuJ. SoHULTZB, op. cU. Tab. i. fig. i.
** Test calcareous.
Genera : Orhulina, Oclma and AmphorioM^ D'Osa*
B. Body composed of several segments. Test calcareous, divided
hy sqpta into cells.
* Odlt imple, arranged on an aaie, etraigM, or MghOy curved.
{Stichostegid).
Genera : Nodosaria Lam. (Sp. Nodosaria IcmeUosa D'Obb.) Anr^
des Se. not 1826, Toul vn. Tab. z. fig. 4-6.— 6%z9u^tna D'Ob&
(Sp. GlanduL ksvigata ib. fig. 1-^), Orthocerirui, Bmtalma D'Ona,
Frondieularia Dsfr., LrngtUma, Eimulma, Vagmtdina, MwtgvnMr
Una, (Jomdma, Pcbvonma, Webbina D*Oim.
^ Since tliese small bodies are separated by D'Obbignt according to characters
especially derired fi^m the shell, we baye thought it sufiBicient, for the sake of breyity,
to indicate the names of the genera.
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INFUSORIA. 49
** CdU nmple, arranged in a apiral
{Uelicostegia),
Genera: CriOeUa/ria Lax., Flabdlina, Robulina D'Orr (Sp.
RolmUna orbicularis D*Obb. 1. 1. Tab. xy. fig. 8, 9), Fundina Fis-
CHEB, Nonicnina, Ntimmvlina D*Orb. {NummvliUa and LenUculitea
liAM. '), Assilina, SidercUna^ Hauerina, Operctdina (Sp. OperciU,
eamplanata D'Orr L L Tab. xiv. fig. 7-10), Vertelfralma D'Orb.,
Polyatamella Lak. [Sp. PolyH, strigiUata D*Orbl Schultze op. cit
Tab. IV. fig. 1], Peneroplis Lajl, Lendritina D'Orb. (Sp. Dendr,
arbtiscula D'Orr L L Tab. xv. fig. 6, 7), Spirolina Lam., Cydolina
D'Orr, Littwla Lajl, Orbicidina Lail (Sp. Orbic, numismcUis
D'Orb. 1. 1. Tab. xvn. fig. 8-10), AlveoUna D'Orb. (Sp. Alved.
Quoit D'Orr 1. L Tab. xvii. fig. 11-13), JRotalina [Sp. EotaL veneta,
E. Freyeri Schultze op, cil. Tab. ui. fig. 1-7], GMngerina, PUmor-
bulina, Truncatulinay Anomalifia D'Orb. (Sp. AnoTfk pwnctiulata
D'Orr 1. L PL xv. fig. 1), Rosalhia D'Orb. (Sp. Roaal, gldfmUmM
D'Obb. L L PL xni. ^. 1-4), Vahmiinay Vemeulina, Btdimina,
Uvigerina D'Orb. (Sp. Uvig, pygmaa D'Orb. L L Tab. xii. fig. 8,
9), PyruHna, Faujcuvna, Ccmdeina, Chrysalidina, Clavtdina D'Orb.
(Sp. Clavul. angtdaris D'Orb. L L PL xil fig. 7), Gaydryna D'Orr
*** CdU aUemoHng diipofed on two cuoef, and arranged in a tpire
{JEtUomosteffta),
Genera: Eobertinay Asterigerina, Amphistegina, Heterostegina,
Caasidulina D'Orb. (Sp. CassiduL Icevigata D'Ork L L Tab. xv.
fig. 4. 5).
**** CdU aUemating, diapoud in two or three rotes, not forming a spire
(JEnaUostegia),
Genera : Dimorphina, GvUvlina, PolymorjMnay Virgulina, Bige-
nerina D'Orr (Sp. Bigen, nodosaria D'Orr L L Tab. xi. fig. 9 — 12),
Gemmidina D'Orr, Textularia Defrance (Sp. Textvl. acicutata
D'Orr L L Tab. xi. fig. 1 — 4), Vuhulina D'Orr (Sp. Vuhul. capre-
olus D'Orr L L Tab. xi. fig. 5 — 8), Bolvvinay Sagria, Ctmeolina,
D'Orr
^ PJutekeSy ZmtieuHtes or LeniH-sUmes. These petn&ctionfl are found in some
loeaHties in such great abundance as to form eztensire deposits affording good
bnilding-stoneB. In Egypt many monuments are constructed of them. Confer Blu-
HBNBAOH, AhbUdwngen naturhist. Oegenstdnde, Ko. 40. According to Debhatks there
is found in most of the stone of which Paris is built as much oiMiUola (vid. p. 46) as of
sand-grains — and it may be said, without exaggeration, that Paris is built of MUioUe.
Ehbxkbebg, Abhandl. der Ahad. zti Berlin, 1838, p. 65.
VOL. I. 4
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50 CLASS I.
' CklU HmpUf dutUred round an aaeu, each tnahing Kaif a wpirt
{Agathistegia).
Genera: UwUoculma, BUaculina D'Osa (Sp. BUoeul, BvUoidet
D'Obb. L 1. Tab. xvi. ^, 1—3), Fabulaaia Defbancb (Sp. FabuL
discolithea D'Orb. L L Tab. rm. Fig. 14-17), JSpiroloculina, Trilo-
eulvna D*Orr (Sp. TrUoc trigomda D'Orb. L L Tab. xvi. fig. 5-9),
OnicUoculinOy ArtilociUina, Sphwroidina, QuinqvslocvUna D'ORa
(Sp. QmnqwHoc. aaxorum D'Orb. L 1. Tab. xyl ^, 10-14), Addonna
D'Orb.
The last division coincides for the most part with the genus
MUiola Lasc. Dujabdin has described and figured a living species
of this genus with its capillaiy processes or extensions under the
name oiMiliola wlgaaia; Infus, PL i. fig. 14.
Order III. Atricha.
Animalcules without a distinct mouth, fiimished with one or
more flagelliform filaments for motion, form persistent or mutable.
Family IV. Monadina, Body not loricated, gelatinous, pellucid.
Monaa Muell. (exclusive of several species). Body oblong or
round, with a single flagelliform filament.
See figures in Ehbenb. Infia, Tth. L To ihiB genus belong animalcules
of Tooo ^% hi which the highest magnifying power shews no oiganisation,
and which even at the present day, with the assistance of the best
microscopes, dannot be otherwise characterised than as punctiform bodies —
the character given by MuBLLBB to his genus Mwuu.
Uvellu BoRY, Ehrenb. [Monadines associated in clusters in
form of a mulberry or of grapes revolving in all directions.]
Cercomonas DuJARD., Bodo Ehrenb. (in part). Body caudate.
Family V. On/ptomonadina. Body loricated, with a mem-
branous flexible test.
Cryptomonas Ehrenb. {Cryptoglena ejusd., with an eye-
point).
Family VI. Vohodna. Several animalcules contained in a com-
mon envelope, or furnished each with its own envelope, which is
confluent into one mass.
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INFUSORIA. 51
Pandcrina BoRT (in part), Ehrenb. Animal without eye-point
and tail, famished with a vibrating fiagellmn, a simple niceolate
lorica, bj spontaneous internal division resembling a mulbeny.
Sp. Pandorina morum, Vcivox morum MuxLL. It^uB. Tab. m. fig. 14 — 16,
Ehbxnb., IfrfkaiontUL Tab. IL fig. 33.
Oontum MuELLu Animals without eye-point and tail, by spon-
taneous division conjoined in a common quadrangular flat envelope.
Sp. Oomum peetomU Muxll. Infut. Tab. xvi. fig. 9 — 11 ; Bh&ihb. Tab,
ni. fig. I.
[81/nura Ehrenb. (Tab. III. fig. 9.) an imcertain genus].
ChlamidomofMs Ehrenb.' Animal with eye-point and double
flagellum, without tail, included in an urceolate envelope, either
simple or multiple from spontaneous division within the common
covering.
Sp. Chtmidomonat ptdvUculus, Mona» pnMMculus Mtjxll. Itrfvi. Tab. i. ^,
5. 6; Ehbsnb. If^furiontth. Tab. iii. fig. x : represented by authors as in
part the green matter of Pbixstlbt. These animalcules were long ago
observed by LsxnwxNHOSOK ; see Sevende vervolg der Brieven, 1709. No.
142, p. 403.
Volvox L. (exclusive of several species) Animalcules with eye-
point and single or double flagellum, included in the surCace of
a globular envelope which rolls on its axis : there are often smaller
globules {ffemnue) within the large one^.
9p. Volvox globator L., Lebuwbnh. Sevende vervclg der Brieven, No. 122^
p. 156, fig. 9 ; BoKBKL, Int. in. Tab. d. fig. 1-3 ; Musll. Ir^u», Tab. m.
fig. 11-13; I^HIUUB. Infttsunuth. Tab. IT. ^. i. DUJABDIK, I^f. PI. iv.
fig. 30. OUbe-animalcule ; a small green globule, as much as J line in
size, and henoe yisible to the naked eye as a fine grain of sand ; in
marshy water. This form was first disooTered by LueuwKHHOlOK. On the
surface of the globule minute warty points are seen ; these are the individual
animalcules or monads of ~ line. Within the globule* smaller globules
are devdoped, which occasionally rotate within the large one until it
bursts and (ties away'..
> [See F.Ck>HN*8 paper in Siibold and Kobllikeb's ZeiU^riflfiir wUtenaehafdiche
Zcdogie, Band lY. p. 77, ftc. for reasons why the VoU>ocina ought perhaps, as Von
SiSBOLD and others believe, to be classed amongst Ahjm.']
[* For an account of the development and encysting of individual monads of the
colony, of a size nearly as large as that of a young colony, see Stkik's I^futionsthiere,
Ac pp. 45—46. When the full size has been attained the cyst thickens into regular
oonical processes, giving the form which Ehrenberg has described as a distinct species,
Vohox tttOattu, These large encysted volvoces are for the continuation of the species
after the ordinary individuals of the colony have perished.]
4—2
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52 CLASS I.
Family VII. Astasice. Body not loricated, caudate or ecaudate,
form mutable.
Astasia Ehrenb. Animal free, caudate, without an eye-point
Sp. See figoree in Ehbknbebg's Infutiondh. Tab. YU. fig. I.-IV. DuJABDnr
Ir^, Tftb. ▼. fig. II.
Euglena Ehrenb. (and Amblyophts ejusd.) Animal free, with
an eye-point.
* Body ecaudate.
AmhlyophU Ehbbnb.
•• Body caudate.
Sp. Eugiena viriditf Cercaria viridii Muell. Jf\fitM, l^b. XIX. ^. 6-13;
Ehbbkb. Infuticndh, Tab. vn. fig. iz ; Dujabd. Infui. PI ▼. ^. 9, 10.
Thifl species also belongs to Prii8TLXT*8 green matter ; another species can
occasionally by its red colour g^ye to water a blood-red appearance.
Family VIII. Periphrygana (.E^Aa^ta Ehrenb. in part). Body
orbicular, surrounded with setaceous tentacles, without vibratile
cilia.
Ehbenbebo ascribes an oral aperture to Actinophn/s, which Du-
JARDIN could not perceive There are no cilia, but there are appen-
dages or cvrrhu
Actinophrys Ehrenb., Peritricha BoRT. Body rough with
tentacles radiating in all directions.
Sp. AitinophryB iol Ehbenb., Trichoda sol MnXLL. Infia. Tab. xxm. fig.
43-45* Ehbenb. IfrfusUmtih. Tab. xxxi. fig. vi. Dujabd. It\fut, Tab. ni.
fig. 3. in fresh water.
Subgenus Podophrya Ehbenb. Body with a transparent appen-
dage resembling a pedicle.
Sp. Podophrya cotn^ta, TrichodafioM MuxLL. ^
Trichodiscus Ehrenb. Body radiating with tentacles at the margin
only.
17 The pedicle of Podophrya is very variable in length. In some of the forms it
almost disappears, so as to render it difficult to determine whether an adimophryB or
podophrya be under examination. Srsiir thinks there is no specific difierence between
the two, certainly no generic. From his observations it appears, indeed, that both
AdinopihiryB and Podophrya are Acinda — forms derived fnym encysted VortieeUa micro-
itoma. Die Infugiotuihieref p. 138, &c.]
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INFUSORIA. 53
Order IV, B^nHcha.
Animalcales moying by means of vibratile cilia.
Section I. Aatoma^,
Family IX. PeridmoBa. Loricated, with a coronet, or a trans-
verse belt of cilia.
Pendinium Ehrenb. (Species of Ceroarta Muell.)
Sp. Peridintum tripos Ehbenb. {Cerearia tripot MuxLL., Ifrfui, Tab. xix.
fig. 93 ; Ehbehb., Ii^uiiontth. Tab. xxn. fig. xyni. ; the lorica termmatea
in three points ; two anterior cfonred backwards, and one posterior,
which is straight. The aiumacule attauas a length of ^ line ; it is foond in
the Baltic.
MiCHAXLis obsenred a phosphoresoenoe in this and some other species of.
this genns, and thus prored, what had been suspected before, tiiat Infu-
sories contribute to the illiunination of the sea. Ueber da$ Lewihim der
Odaee, Hamburg, 1830; oomp. Ehsznbebo, Dai Leuchien det Meeres. Ein
in der K&iUgU Akademie der Wiiaeiuckti^ftem gehaltener Vorirag. Berlin,
1835, 4to.
To this genus also probably belong as fossil species some oiganio
remains which Ehbimbibo disoorered in the ehalk-fiannatioii in fire-
stones.
Dinophys%8 Ehrenb. {Ahhandl. d. Konigl. Akad. d^ Wiss. zu
Berlin, a.d. Jahre 1839, s. 124.)
Section II. Btomatoda. (Animalcules with mouth and oeso-
phagus leading into the parenchjme of the body. One or more
round, contractile cavities, pulsating rhythmically, situated beneath
the integument at the surface of the body.)
Family X. Trtchodina {Tracheltna and Colpodea Ehrenb.)
Body oval, with vibratile dlia, without cirri or styli, not loricated,
Trichoda MuELL. (in part, T}ri€hoda DUJARD. and Trachdiua
ScHRANE, Ehrenb., Dujard.) An oblique row of large cilia at
the mouth.
Phiodma BoRT, Ehrenb.
^ I consider this desoription as merely provinonal. As to the presence of a mouth
in particular genera of Infusories much yarietj of opinion exists, and that the point
is not easy to determine wiU be obvious to ereiy one who obserres for himselfl This
character, then, in the present state of our knoiHedge, scaroelj deserves a promineut
position* [Vid. note I, p. 39.]
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54 CLASS I.
Efnchdya MuELL. (exclusive of several species^), Aoomia
DujARD., OastrochosUi ejnsd., AlyBcum ejnsd., Unmoma ejiiscL
Btirsaria MuELL. (in paxt), DuJARD, {BursariaBxA Spirostomum
Ehbekb. in part.) Body everywhere ciliated, often dilated pos-
teriorly ; mouth oblique surrounded by cilia arranged spirally.
Sp. BvTdaria truneatdla MuXLL.^ Ir^, Tth, zvn. figs. 1—4 ; EHfiBNB.
Infusunuth. Tab. zxxiY. fig. 5.
To this division Ehbxnbsbo refers the Opalina Manarum of Pubkin jb and
Valbnttk, first discovered and figured by Lbxuwxnhoxok : OtUUdingm
en Onidekkingm 1685, p. 13, fig. 3, A. Dujabdik and VOK Sibbold [and
Stbin.] do not admit the presence of a mouth in this species, the first two
retain the genus Opalina. [Stbht. suspects the Opalina to be larvie of
worms. The different species have very different structure. Die It^fit-
iiontth, s. 181 — 187.]
Ophryoglena Ehbenb. Body rough with cilia disposed in lon-
gitudinal rows, ovate, with eye-point black or red.
See Fig. in Ehbbkb. Tab. XL. figs. 6-8.
Spirostomum Eurexb. in part, Dujabd.
Glaucoma Ehbenb. Body everywhere ciliated, mouth un-
armed, with a tremulous valve like a longitudinal lip.
Sp. Glaucoma icintUlam Ehbbnb., If^uiiontth. l^b. zxzvi. £g, v., Dujabd.
J^fM. Tab. VL fig. 13.
Chilodon Ehbenb. Body oval, with a lateral sinus forwards,
cilia all over disposed in longitudinal rows, mouth inclosing a cylin-
drical fisusciculus of little rods {teelJi),
Sp. CkHodon cucuUvhu, Kolpoda eueuUvku Mubll., Ehbbztb. If^futioMih,
Tab. xzxvi. ^, VI., Dujabd. Ifrfui. Tib. vi. ^. vi. [Sibih. Jf^uiioneth,
Tib. HL 51.]
Nassula Ehbenb.
Lacrymaria Ehbenb. (and Trachelocerca ejusd.)
Sp. Lacrymaria dor Ehbbnb., yibrio dor MuBLL., /t^. Tab. z. figs. I3-I5,
Ehbbnb. Ii^vowmO^ Tab. xxxvm. %%. vn.
Colpoda Ehbenb. (Species from the genus Colpoda Muell.)
Body laterally emarginate or sinuous, reniform, with cilia disposed
in rows, mouth lateral unarmed.
Sp. Colpoda cucuUut MuBLL.| Ii^, Tib. ziv. figs. 7-14^ Ehbbnb. Tab.
XXXIX. figs. V. &c.
Paramecium Muell. (exclusive of species), Ehbenb* (in part)^
Dujabd.
^ Nothing can be more ci^ricious than the use by modem writers of this generic
name of Mubllbb. See DujabdiVi Mitt, not, det Infus, pp. 385, 386*
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INFUSORIA. 55
Amph%lqp(u8 Ehbenb. (Amphileptua and LoxaphyUum DuJASD.)
Sp, AmfkU^pitu mdeagrii, Kolpoda mdeagru MuiLL., /i0it. Tab. ZIV. figs.
I-6» ZV. figs. 1-5, EhBXBB. 1^. ZZZYXU. fig. 4..
Family XI. Oxytrichtna. Body mostly plane or depressed,
armed with yibratile cilia and set», and hooks or styles not vibra-
tile, not loricated.
This fiunily agrees with the genus Kerona of Mueller. Besides
the usual fine cilia^ the animals have other organs for creeping and
for the support of the body in climbing, and which are distinguished
by Ehbeetbebo as bristles («eto), styles {stylC) and hooks (tcnctm) ;
Infudcmstk. & 338.
Genera : Kerona Muell. in part^ Dujabb. {Stylonyehda and Kerona
Ehsknbl, Ceraiidvum ejnsd) — Oxf/irieha Boby {Oxf^irieha and
Urottyla Ehhenb) ; HtUteria Dujakd.
Sp. Kenma mytOus (and Ker. hoMteOum) Mvill., I^fui. Tab. zzxiT. figs.
1-4, Ehrenb. I^futiontth, Tab. XLI. fig. ix.^ Dujabd. Ii^fui, Tab. xm.
figs. 7, 3 ; very oomznon in fresh water, size ^ to 1 line. If this form be
oompared with monads, Vibriot and the animals of Volvox globatar, it will
be seen that in respect of size there is as great a diffiBrence amongst Info-
sories as amongst Mammalia.
Family Xn. .E^2bto(.£^2oto and ^^tc2ik»na Ehrenb.) Body
oval, depressed, loricated* Yibratile cilia around the month ; besides
cilia, often styles and hooks for motion.
Euphtes Ehrenb. {Phlascmia Bobt, Dujard.) Lorica oval or
snborbicnlar, longitudinally ribbed or striated; body with styles
and hooks.
Sp. Euplota patdla, Kerona patella Mxjvll,, I^fue, Tab. xxxni. figs. 14-18,
Ehbbnb. IftfuticndK, Tab. zin. fig. ul, Dujabd. Itrfue, Tab. vm. figs.
1-4.
Ghlamtdodan Ehbenb. Styles and hooks none. Slender rigid
rods arranged in a cylindrical fascicnlus aronnd the mouth (teeth).
Sp. ChUmUdodan mnemoeyne Ehbihb,, Irrfuaiontih, Tab. XLn. fig. Tnx ; in
the Baltic.
Dtcpkrys Dujard.
ffimantophorus Ehrenb.
Aspidiaca Ehrenb. Lorica produced beyond the body forwards,
hyaline ; longitudinal flexible setss on the ventral side^ for stepping
and creeping.
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56 CLAfiS 1.
Sp. Aipidiica l^neetu, Trickoda fyneeui Musll., /V«t. Tab. XXQL figs, i, i,
Ehbeitb. Inftuiontth. Tab. XTXTX. ^. i.
Loxodes DuJABD.y not Ehbenb. (jonng indiyidiials of Chilodon
cucullulus with imperfectly developed infdndibulum. Stein. Iti/us.
p. 131.]
Ervilia DuJABD. (Species from the genus Euphtes Ehrenb.)
Trochilia DUJASD.
Family Xm. VorticelUna ( Vorticellina and Ophrydina Ehbenb.)
Body campanulate or infimdibuliform, with large vibratile cilia at
the margin of the aperture. Mouth and anus approximate, situated
in a pit of the margin.
Formerly these «-Tiifntt.la were classed amongst the Polyps as BeU-
Polypa, BastardrPclypSy dw. (See the first Dutch edition of this
Hamdbooky l p. Q^,) It was belieyed that the infundibular or bell-
shaped body is the cecal stomach, and the large opening the
mouth. The true oral aperture, howeTer, is placed on the edge of
the hollowed body. The food describes a circle in the parenchyme
(according to Ehbenbkbo in a special intestinal canal* with several
lateral dilatations) and is again ejected near the oral aperture (hence
the names cydoccda and cmopisthia given by Ehbenbebg). The
analogy with the molluscan type, even if an intestinal canal be not
admitted, cannot be overlooked, and probably these aniTnals will be
ranked by future writers, as imperfect forms, with the Bryazock
The cilia on the edge of the bell-shaped body cause in the water
an eddy which hurries onwards minute oorpucles whether dead or
alive, and conveys them towards the cavity. If some early and
also later observers (amongst others even Aoardh, Nov. Act, Acad,
CcMo/r. Leop. Carol, Nalwr. CtMriosor. x. 1821, pp. 127 — 137, Tab. vn.
iL) have seen in this a power of fiusdnation, the &ct must be ascribed
to the circumstance, that they did not notice the cilia.
Phalanx I. Body not pedunculated.
A. Naked.
Stentor Oken, Ehrenb. (Species of Varticella Muell.)
Body conical, from its contractility polymorphous, everywhere
^ See FoKKS's obserFatioziB on Stentor, wbich led him to doubt bo eariy the exist-
enoe of a special intestinal canal. Okbh's Int, 1836. s. 785, 786.
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INFUSORIA. 57
covered with small cilia, besides a coronet of larger cilia, firee or
temporarily sessile.
SteiUor Mudieri Ehhxkb., Hydra ttenUma L., Bossn Int. m. Supfil, Tab.
XdV. figB. 1, %, MUBLL. InfuM, Tab. zuii. figs. 6-13, Bhbxnb. InfvMtmMk.
Tab. xxin. fig. i, Dujabd. Ii^ub, Tab. xv. fig. i ; on the ander suHaoe
of Lemna. When swimmiog the animal has an OTal f<Mnn, and moyes in
Berpentine and differently carred Unee ; when at rest or attached, it has an
extended trumpet-Hke form.
Urceolarta Lam. (in part), DuJARD. {Trickodina Ehrenb.)
Bodj globose or nrceolate, not ciliated throughout
Urceolariadeaina, Trichodinapedietdvs'RBXXSB., Oydtdium pedieuUuMvELL.
and VorticeUa ttellina and discina ejiud., MUKLL. If^fiu. Tab. xxxvm. figs.
3-5, Eebkhb. I^fui. Tab. xxiv. fig. iv. The ammalcole creeps on finedi-
water polyps and other bodies by means of moveable hooks on the dofsal
Bor&oe (or on the extremity opposite to the opening) ^ : in swimming it
rotates rapidly.
Urocentrum NiTZSCH, Ehrenb.
B. Loricatedy or contained in a geUxtinous envelope.
Ophrydium Ehrenb. {Ophrydia BoRT in part.) Aggregated
animalcules, contained in a common gelatinous body {infusoror
rium ?)•.
8p. Opkiydimfk venaOle Ehbkhb., Ittfm. Tab. xxz. fig. i. Green globnlar
mnssii! of i— 5 inch, in diameter, forming as it were the infnsory-stock or
hiye of an animal whidi Mujillbb first described and figured as VorticeUa
versaiUia. Earlier and later authors have taken this Polypaiy for a species
of plant, and have described and figured it under the name of Ulva prufU"
formis, Fvcu§ tubglobotus, CoceoeMoris atagtnna, &a The analogy of
the Yortioellines with the JBryoeoa and Molluscs (compound Aaeidiof}
slluded to aboTe, derives confirmation from this form also.
Vagtnicola Lam. (in part) Ehrenb. Animal solitary with ur-
ceolar lorica, body and lorica sessile.
8p. Vaginieola cryslaUina Ehrsnb., /^tu. Tab. xxx. fig. v., Dujaed.
It^, Tab. 16 bis, fig. 6 ; formerly observed by Lexuwskhosox ; they
propagate by longitudinal fission within the transparent sheath.
^ [For an elaborate description see Stun. JHe I^fus. s. 175.]
' [The geUtinous infusoiy-stock or hive is a product of secretion from the base of
the body of each animalottle, and so is the homologue of the pediceb in V orticelUnes.
Vide SniH. op. eU» p. 946.]
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58 CLABS I.
Caihimiia Ehbekb. Animal solitary, Bessile, with niceolar
loiica, pedimcolate.
(According ta Dujabdht not raffioiently diatinct from the preceding genns.)
Phalanx IL Body (in the first period of life) pediculated.
A. Naked.
VorttceUa Muell. (exclusiye of several species). Animal cam-
panolate, with a flexible pedicle spirally contractile*
These animals adhere to water-plants, water^insects, small Crustacea (C^dops),
&0. At a certain period they part from the stem, and then, as freely
moving forms, are provided with cilia near the posterior extremity by
means of which they move forward, whilst the coronet of cilia at the edge
of the opening is entirely retracted*
a) wkh rimple pedioU,
Vorticdla Ehbemb. Sp. Vorticdla cowoaUaria L., VcrUeeUa 7iAfd\fera
Ehbenb., Robssl, /ns. m. Suppl, flgs. a, 4-7, Muzll. Iitftu, Tab. xlt.
fig. I, Ehbsitb. Tab. zxt. fig. i. — A yeiy similar species occurs in artifidal
infusions, which on contraction exhibits transrerse rings, and which
Khrkntibrq distinguishes as VorUc. convaOariet,
h) vUhbranMngpedick,
CardMtm Ehbemb. Sp. VoHiceUa poli/pina L., Mubll. /i0tf. Tab.
XLVi. figs. 7-9 ; Ehbekb. /«^. Tab. zxvi. ^g. 5 ; polypa d hovqwt
Tbbhblbt ; resembles an umbeDifBrous shrub, of about one line in sixe^ in
fiiesh and also in sea-water ; see Bi.8TBB, Naiviwrk. UiUpcoMtingtn i. Tab.
m. fig. I, 0 ; Slabber, Natunvrk, VerkuUgingfen 1778. TM>. ▼. fig. a.
[ZoothamniuM^ Ehbemb.
Sp. ZootkamniwH^ airhueula Ehbemb., Ifrfuiianilh. Tab. xxnr. Bg. «.]
P The stem of spirally flexible Vorttcellines consists of a wall and an internal
canal contuning a thread, or streak, which does not exactly fill it. "When a stem or
branch divides, the structure of the divided parts is not exactly the same in Carekesium
and Zoothamnium, In Carckerium the canal and streak of a branch have no connexion
with the corresponding parts of the stem on which it stands. After each fission
one only of the fission-progeny occupies the apex of the already existing stem, and
continues to prolong it by secretion of new matter, the canal and the thread suffering
no breach of continuity. The other individual secretes at first a short portion of stem
which is quite solid, and is in connexion with the outer wall only of the stem pre-
viously existing. It is after this commencement that a new canal and a new streak
begin to be seen. The same occurs at every subsequent division : the individual at
the apex has the canal and streak of its stem in continuity with the similar parts
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IHFUSORIA. 59
I^fiatylis Ehbenb. (and Opercaloaria ejnsd.) The aniinal coni-
cal or campanulate, with rigid pedicle, suDple, or branched from
imperfect spontaneona diyision.
Sp. Ejpiutiflu JUimeam»'Sbsaxsh.,Vo^^ xsu
SmppL Tab. 0. ;— QpemilarHi ofiteufato Bbbivb., BoniLy ib. TU>. zcnrm.
figB. 5, 6, &c.
B. LortcatecL
TmUnmus Ehbenb.
Sp. I^MmMM Mijtttf mici, VagmMa mgpiUma Lam.
previooBly existing, whilst tlie other has the begiimmg of its stem solid, and afterwards
a canal and streak not oontinaoiis with thoae prerionsly existing.
In ZooCAamaifHM a continuoos canal nms throng the stem and all the hnnohes of
the colony, and the streak also divides at every fork, so that all the streaks and canals
are in oonneaion, Sixar* op. cd. pp. 8a — 84.]
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CLASS n.
POLYPa (POLYPI)'.
Polyps are gelatinous, oblong or conical animals with a con-
tractile body, an intestinal cavity and an oral aperture, which is
surrounded by a circlet of arms or tentacles.
Besides these arms there are no special organs of sense, at least
in the greater number of Polyps, though all appear to be very sensi-
ble of the stimulus of light. Propagation is effected partly by eggs,
partly by germs or buds : in many instances the last are not de-
tached firom the parent stem, and thus there arise compound animals^
different individuals being connected.
Our Polyps were, for the most part, unknown to the ancients :
and under this name entirely unknown. By it they understood
naked molluscs of the form of sepia, especially that genus which
is now called Octopus^ by Zoologists. From analogy, and from
some resemblance of form, BiiAUMUR and JussiEU first gave the
1 Of the numerovui woiiu on tliiB dan we tan content to qnote the following :
A. TsEMBLiT, MinunreBpowr mvir d VHidoire d^une genre de Pclypet d^eau dauee,
d hra$ en forme de eomet. Leide, 1744^ 4to.
J. Ellis, An Eteay towards a Natural HulUiry ef tkt CbraOtnef os^d ffAer JIfarine
Produetumi, &o. Lond. 1755, 4to, with plates.
J. Ellis and D. Solahdbb, 7%e Natwral HiMory ef many eurunte and uneammmi
Zoophytes, with 62 plates. London, 1786, 4to.
P. S. Pallas^ Elenchue Zoophytontm. Hag« Oomitnm, 1766.
F. Cavolini, Memorie per eervire alia ttoria cfe* PoUpi Marini. Kapoli, 1785, 4to.
E. J. C. EsPBB, Die Pjlansenthiere in Ahlnldungen naek der Natwr. m. Thle.
Nlimberg, 1761 — 1797 (with two Bupplements).
W. Rapp, UAer diePolypen im AUgem/einen tmd dieAeUnien inebesondere. Weimar^
iSap, m. 3 color. Kupfertafeb, 4to.
0. G. Ehbshbkbo, JDie CoraUetUhiere dee rothen Meeree, PhyeQeaUecKe Ahhand-
lungen der KdnigL Ahad, der Wieeeneek^ften' tu Berlin aus dem Jahre 1833, s. 315
— 380. (Also pnbliflhed separately, Berlin, 1834, 4to.)
G. JoBHBTOV, Hidcry of BrUieA Zoophytes, Seeond edition, with numerous tSttf-
irations on copper and vfood. 1 vob. 8to, 1847.
' The French name Pouipe now giren to this animal is merely a oorraption of the
ancient name Potypms,
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POLYPS. 61
name Polyp to those fresh-water animals that had been described
bj Trembley, and which are provided with a circlet of arms.
To this class belong many marine animals, which at first sight
rather resemble plants than animals. Formerly these so-called
SearPlants were, on accomit of the hardness of the calcareous sub-
stance of which they consist, referred to the mineral kingdom : and
Corals were compared to branching crystallisations {Arbor Dtanoe)
and stalactites. Hence the name {Lithophytay Lithodendra) stone-
plants. The ancients believed that Corals were soft whilst in the
sea, and only became hard in air :
Nunc ^[uoque euraUis eadem natwra remantit,
Duritiem taeto capiant ui ab aere, qiiodque
Vknen in cequore erat, pU tuprs cejuora muMim,
Ovid, M^, iv. 750—752.
Even amongst later authors traces may be found of the same
opinion, founded on defective observation, or on conftision of soft
species with similar hard ones. Up to the middle of the last cen-
tury, it was the prevailing view that these Corals belonged to the
vegetable kingdom. Mabsigli, in 1706, observed on* the shore of
the Mediterranean some of these products {Alcyimium^ CoraRxwrn^
Aniipathesjy and found in their pores little bodies that contracted
when the stem was removed from the water. Such bodies or buds
he took to be flowers, and so believed that at length the view was
definitively established which consigned these marine products to
the vegetable kingdom. But still the animal odour, that was ob-
served, opposed this view, as well as the chemical investigations
of Geoffeoy, of Lemery, and of Marsigli himself, which demon-
strated ammoniacal constituents in these supposed sea-plants, just
as in animal substances. Peysonkel, a physician of Marseilles,
observed at that place (1723) the Blood-Coral, and afterwards on
the coast of Nortibem Africa examined different Madrepores and
Millepores : the result was that he found Mab8IGLI*s Plants to be
Animals, and named them Orttes Corallines. He imparted his dis-
covery to K^UMUR : to whom the notion seemed so improbable,
that in a short notice of it which he gave in the MSmoires of the
Academy of Sciences at Paris 1727, he felt bound to suppress the
discoverer's name. Shortly afterwards, when Peysonnel's dis-
covery had been forgotten, Trembley found in our country the
feesh-water Polyp, and communicated his observations to Keaumur,
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62 GLASS II.
In the two following years Bernard de Jussieu, the celebrated
Botanist, investigated AJUyaniwn, {JLobtdaria)^ Flustra and Hdn^
laria on the coast of Normandy, and confirmed Peysonnel's dis-
covery: whilst B^AUMUR also adopted his views, Linn^us,
accordingly, transferred the Corals and stone-plants to the animal
kingdom: and thus more than half a century was required to
effect the adoption by Science, as a firm truth, of that view which
Ferrante Imperato had announced at the beginning of the
16th century ^ Ellis, Pallas, Cavolini and other authors,
in the latter half of the past century, extended and multiplied our
acquaintance with these interesting marine animals, of which the
investigation still affords to S9avants of the present day a rich
material for new and important discoveries.
Polyps are either naked, or are provided with a body more or
less hard, which they surround like a bark, or by which they are
surrounded. To the naked Polyps belong the well-known Armed
Polyp of fresh-water {HydraL.^ Polype ^eau dauoe, h bras ^n forme de
cornea). The body of this animal is hollow within, and terminates
in a little cylindrical stalk that is without any opening. There is
a single row of tentacles round the mouth which can be extended
like long rays, or be contracted into little conical swellings.
These tentacles are not all formed at once, but at different times :
their number is therefore indeterminate, and frequently varies in the
same species. Grenerally there are not more than six tentacles
present : rarely more than twelve. By their assistance the fi:esh-
water Polyp can creep along upon water-plants or upon the bottom,
overpower its prey, and convey it to the mouth. These Polyps are
very voracious, and feed upon minute Crustaceans {Cypria^ Bcephnia^
Monocuhia, &c.), and upon worms {Stylaria pahidoaa Lam. NalSy
Tuhifsx, &c.), which frequently surpass them in bulk. Accordingly
^ To complete this oompressed hiBtorical reyiew, we refer to B. Dx Jusanu, Examen
de qud^uea productiona marirtea, &c. MSm. de VAcad, rcyale des Sciencet, 1742. pp. 290
— 302 ; Kbauhub, Mimaireg pour aervir d VHistoire de$ TnaecteSf Tom. vi. 1741. Pre-
face, pp. 49 — 80 ; Pallas, Mench. Zoophytor. pp. 13 — 20; LAVOiaNON Malbshxrbss,
ObaervoHont 9wr VHidtcire mUur. de Buffon et de Daubxbton. Paris, 1798, n. pp. 154
— 206 ; Ehrsnbxbo, IHe Corallenthiere dee rcthen Meeres, pp. 4, 5 ; Milne Edwabds,
Ann. det Sc. Nahir. sec. S^rie, Tom. vi. Zoohgie, 1836. pp. 5 — 9; Flourkns, Analyse
<Pun auvroffe mantuerU intUulS, Tra^ du CoraU dx. par Ds Pbtbonnkl, Ann. dee Se.
Nai. 8ec. S4r. Tom. ix. Zoaloffie, 1838, pp. 334 — 351.
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POLYPS. 63
their mouth admits of much expansion : and the body can be dis-
tended. The food is moved to and firo in the cavity of the bodj,
and in a short time (often within a quarter of an hour) is converted
into a pap. The undigested residue is rejected through the mouth.
Propagation is effected usually by buds. A minute swelling rises
on the surface of the Polyp; it grows, loses its conical form, be-
comes tubular, acquires tentacles, and is then a new Polyp. The
yoimg animal continues to sit on the body of the parent, and thus
receives the same nutrition in common. Upon this young one
other young buds may be developed. Thus a branching arises.
At length the young Polyp separates itself from the parent stem
(in summer frequently after four days, in winter later), assumes an
independent state, and new buds are formed, or those already
present are multiplied.
Thus these Polyps may form compound animals. Many indi-
viduals of the same species are united so as to make up a single
body. All the animals thus combined gain their nutrition in com-
mon,— ^have a common life. It is not the animal kingdom only
that affords us instances of compound living bodies : the vegetable
kingdom presents many such^. By an individual, in the vegetable
and animal kingdoms, may be understood a body that cannot be
divided into two or more similar portions, without the idea of a whole
being lost, and whose vital fimctions pass through a determinate
cycle of periods*. The development of the fruit is the final function
in vegetable life: when this is accomplished the plant may die.
Many plants bear fruit only once — whether in one year, or in two
or more years ; such plants die after fructifying, and are true indi-
viduals. There are other plants again, which leave a determinate
portion after the fructification, that continues to live, and, after a
time, bears firuit anew. The portion that thus remains may con-
sist of root alone, or of root and stem. Such plants are, in reality,
compotmd.
A tree therefore is not an individual, not a single plani. The
buds of the tree are new plants : they are developed, grow, possess
an independent life, which is passed in determinate stages. Hence
> See Lamabok, Ifisi, Nai, dn onm. mwm vtrt, I. p. 69, fta (and and edit. p. 65,
Ac). CoimpaniKm da Ammawe eampoaSt atvec de$ vigitaux pttreilUment C9mpo§i9,
3 See SoHLBiDBN in Mubllbr'b Arckiv, 1838. b. 168.
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64 CLASS II.
old trees may afford an emblem of pereimial youth : every spring
they are covered again with leaves as fresh as those they had fifty
years before. The stem alone is old, the leaves are still yoimg
again.
We might be able, from the branching of the fresh-water Polyps
from their living stem, to explain the plant-like forms of Corals
and other such marine products. When a Polyp does not consist
of a single soft mass, but contains a harder substance, or is sur-
rounded by a calcareous sheath, then from the union of many such
a body may arise which resists decomposition, and as such after the
death of the Polyps, may be preserved in our collections for a
length of time, as for ages they have been preserved in the cal-
careous strata of our mountains, formed at the bottom of the sea in
a former epoch of the world. This common mass is named a PoZ^-
pary or Polypstock {Polyparium) ^. After the Polyps had been dis-
covered, these stone-plants, as they had been called, were supposed
to be the work of the animals that dwelt in them, and were com-
pared to the cells of bees. This view of the matter does not now
require confdtation. That of Lamabgk and others agrees more
closely with the true nature of the process; they consider the
polypary to be a secretion upon the surface of the Polyps, and com-
pare it with the shells of Molluscs (Snail or Mussel-shell). As
there are Snails both naked and with shells, in like manner there
are Polyps that are naked, and that are shut up in tubes : and the
Polypstock is the union of the shells caused by the connexion of
the Polyps that lived in them. Thus the Polypary would be, on
this view, a dead substance, deposited in layers like a mussel-shell.
Though this be nearer the truth than the earlier idea according
to which the Polyps built their houses, still it does not entirely
accord with the true nature of the process. Observation proves
that this part, at least in many species, has a proper life, that it is
nourished, grows, and is the seat of that gemmation whence new
^ It appears that Rii&uicuB first invented this appellation, now in common use ;
"AuroU-on pupr^wir. , . . que cet carpt qui tembloient avoir vigiU dam la mer, itment
pour les poifpet ce que lea guipier* 9orU pour lea guipea; qu'on ne devoU plui leur lai8$er
le nom depUmtea et que pour lew en imposer un qui exprUndt exactetaeiU ee qu'Ue soni, on
devoU lee appeUer dee pciffpierst" Mim. pour eervir d Pffiet. dea Inaectea. Tom. vi.
Preface, p. 69.
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POLYPS. 66
Polyps have their heing. It is a covering which, like a dermal
skeleton ^ may become homy or calcareous.
The hard, stone-like Polypstocka which form coral-banks, are
particularly deserving of notice. But the part they play in altering
the earth's surface has been much exaggerated by Forsteb, Peron,
and other voyagers. The numerous coral islands of the southern
Pacific having an annular form with banks steep on the outside
and shelving gently down to the trough or the included water, are
clearly of volcanic origin. They are covered with Corals, but do
not consist of Corals. Poljrps cannot live at great depths, but the
Corals rest on shallows or on mountain-ridges in the sea, similar to
the rocks parallel to the coast of the Bed Sea. Hence Corals may
contribute to the formation of islands, or may prevent the washing
away of the shores of islands already formed, just as plants that
grow on sandy coasts protect the hillocks from being blown away'.
After these general remarks on Polyps and Polypstocks, we must
dwell for a little on the particulars of structure of the different
animals that belong to this class. It would be a defective and
erroneous idea, to suppose that Tkembley's fiesh-water Polyps are
to be considered as the Type of the class. That we drew the atten-
tion of our readers, in the first instance, to the firesh-water Polyp, is
merely to be attributed to the historic form which, in introducing
this class, we thought useful fur the right understanding of it. The
animals which live in Polyparies have in several respects a much
^ See MnjTB EDWABDfly ObmrvatianM 9wr la natwre H U mode decroittanee det Poly-
pien, Ann, des Sc. NcOnr. Seconde S^e, Tom. x. 1838. Zooloffie, pp. 321 — 334.
Lamabck appears to me in some degree to contradict himself, when in one place he
caOa the polypary a common hody possessing an independent life, and producing new
indiyiduals upon its surfkoe, which die and are again replaced by new ones, and con-
tinuing its Hfe afanost unobserved as long as it is surrounded by water alone {ITiM. Nat,
des AfUm. earn Vert. i. p. 63^ new edition) ; and in another place denies to the poly-
paiy aU Ufe, and compares it with the shells of molluscs, ibid, n. pp. 86—99. Before
this, liiHHiBtTS, Pallas and others had recognised in the polypaiy a proper life, but of
late yean this opinion, on the authority of Lakarck, has been almost generally
reBnquiehed.
' Comp. J. B. F0R8TSB, Bemerhtngen avf eeine Seiee um die Welt, Wien. 1787.
Sto. b. ISO, 131 ; A. TON Kotzbbux, ErUdeekunge-reiee in die SikUee, iii. Weimar 182 r,
B. 187 ; QuoT et Gaimabd, Mimmre twr VaecniaemenJt dee Polypee eoneidM gSdo-
^qimemeni, Ann, dee 8e. Nat, vi, 1815. pp. 9^3 — 390 ; Ehbxnbkro, Ueber die Natur und
Bildmngder OoraUenbanhe dee rotken Meeree, Phyeik. Ahhandlungen der Akad, der Wie-
senscA. ru JBerUn, 1833. b. 381 — 438.
VOL. I. 5
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66 CLASS II.
nearer alliance to SeorAnemonies {Actinics), which Chamisso and
EiSENHARDT had properly classed with Polyps^, although Cuvier
joined them to the Medusae {Acalephw), Lamarck and Schweioger
to the Star-fishes {Eckinodermata). These Actintof have a tubular
form, or resemble truncated cones. By their diacoidal base they
adhere to rocks, marine shells, and other bodies; but are able to
loosen their hold, and to consign themselves to the motions of the
water. They can also creep by means of that base, as the belly--
footed molluscs {Gasteropoda) do by means of their ventral disc.
But ordinarily the motions of these animals are restricted to a
greater or less expansion of the oral aperture, and to a contraction
of the hollow tentacles which surround the mouth in a variable
number, but always greater than twelve. These Actiniae are naked
Polyps, rather of a coriaceous than a gelatinous consistence ; they
were not imknown to the ancients, and are noticed by Aristotle'
as Acalephw^ and by Pliny* as Urticw. Such Polyps with Polyp-
stocks are the genera Fungiay CaryophyUa^ Astrofa^ Mopandrina.
The Polyps of other Polyparies, as Ins, Ahyonvum {Lobularia),
Tuiijfora, &c. have eight tentacles, which are flat and notched on
the edges or have lateral {»*olongations. In all these the intestinal
canal is a blind sac. But there are other Polyps which, by their
more perfect organisation, approach the Molluscs. Their intestinal
canal is reflected upwards, and terminates by an opening close to
the mouth. AuDOUiN and Milne Edwards observed this struc-
ture (1828) in Polyps of the genus Flvstra^: at the same time
Ehrenberq published his earlier observations to the same efiect,
and gave to Polyps, with this organisation, the name of Bryozoa;
which has been received into the systematic works of zoologists*,
^ Nov. Act, Acad, Ccuar, Leop, Carol, Natur, Owiosor, x, p. 354, 355.
' "E^Ti 8^ Kttl rd rOif dxaXi^^cai' yiifos (Stor ' irpocTi^M di rcut Tirpats, iStnrep hia
tQv 6arpaKod4pfuav' dToKAerai 6' Mart, O^k ^ec 8^ 6<rTpaK0P, dXXd ffapKQ6€s t8p
i^Ttp a^ov, jc.r.X. De Anim, Hid, r7. c. 6. These words, in my opinion, apply to
Actiniie alone.
' Hiat. Nai, Lib. ix. c. 68. But that Medutce also (our Actdephci) were by the
ancients designated under this name, I will by no means deny.
^ Raumi det recherehet nur lea Animaux 9an$ VerUbret faitei aux Ue$ Chaiuaey, Awn,
dea Sc, not, Tom. XT. iSaS. pp. X9, 13.
' Symhola phyaioa $eu Ieone$ ei De&eripiionet AtUmaUum everUbratorum ex itinere
F. G. Hbmpbicfh et C. G. Ehbsnbkbo, Dea I. Berolini, 1828. folio. Polypi, p. 1. Also
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POLYPS- 67
and is now commonly used* Milne Edwabdb makes a separate
class of these creatures, and places it amongst the Molluscs, after
the naked Acephala^ or Tunicata, Their arms are heset with cilia,
which however are also remarked in some other Poljps, and conse-
quently are not a distinctive character of Bryozoa, as would seem
to be implied by the namis Ciliobrachiata. To these Bryozoa belong
the genera AlcyaneUa^ Eschara^ and Flustra. The remaining
Polyps we shall name, with Ehrenbeisg, Antkozoa.
The simple stomach of the Anthozoa is, for the most part, sepa-
rated from the cavity of the body. In Actinia this space is divided
into many prismatic cells by partitions that stretch perpendicularly
from the outer surface of the stomach to the innermost surface of
the covering of the body. Similar partitions are found in many
Anthozoa, though in much smaller number. One or more openings
conduct from the bottom of the stomach to the cavity of the body,
or -to the general common cavity of the Polypstock^ In Hydra
there is no special cavity of the body, but the cavity of the stomach
is in immediate connexion with the walls of the body. It was
formerly erroneously believed that the stomach of this animal is
simply an excavation of its body, and that the structure of both
surfaces is the same. The internal surface is coated with conical
cells whose points are directed inwards^. The external sur&ce, on
the contrary, is formed of flat cells, and contains oval vesicles, from
which a long delicate thread can be projected, {AngeUcrgan of the
Grermans'). Tremblet, amongst his many experiments on the
reproductive power of the fresh-water Polyp, even turned the body
inside out, like the reversed finger of a glove. Nevertheless, the
creature continued to live, and took food. This may be explained
by a change of structure, the consequence of the violence of the
experiment. In Bryozoa, the intestinal canal is freely suspended in
the cavity of the body: a longer or shorter oesophagus leads to a
muscular stomach, lined, in some cases, with homy teeth, closely
Graxt, 16 etfly as 1837, had obserFed the reverting inteetiiud cMial, and the vibrating
dha on the arms oiFluaira, Edmb. New PkOoi, Jawm, in. pp. 107 — 337.
^ LiSTKB, PkHoi, Trcmaae, 1834, p. 371, PI. vm. fig. 3 in SerUOa/ria, Milnb
Edwasdb in the new illustrated edition of Cutub, lUgne Amm, Zooph. PL lxxz. in
Jn»nobSi$t fta
* See CoBDA, Nov. Act. Aead. OcBsar. Leap. Carol, Natwr. Curiotor. Tom. xym.
Ann, daSe, natur. Tome vin. Zoologie, p. 363.
5-2
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68 CLASS II.
set in rhomboidal rows : then comes a second stomach ending below
in a blind sac that is continued at the upper part into an intestine,
which ascends by the side of the stomach and oesophagus, and
near the mouth, or a little below it, terminates in the antis.
Brown follicles cover the external wall of the stomach, and seem to
represent the liver^. In some Polyps a circulating system has
been observed, or at least vessels, which probably arise as branches
from the intestinal tube, anastomose with one another in the Polyp-
stock, and effect a communication between the different individuals'.
In many, moreover, a stream of water is found, which penetrates
by the mouth into the canals of the Polypary. It has been observed
that the flow is caused by cilia on the walls of the canals. Probably
this motion is in connexion with the fan(;tion of Bespiration.
Lister saw in the stem of Plumularia pluma Lam. the stream in
the same canal moving alternately in opposite directions'.
We have seen above that propagation in Polyps is usually
effected by buds. In Hydra^ after being developed, they are sepa-
rated: in others they remain attached to the parent-stem. But
besides this mode of propagation, a sexual generation has been
observed in this class. In Hydra^ about winter-time, a periodical
development of eggs on the inferior part of the body has been ob-
served. The thin membrane, surrounding the egg as it projects
from the body, bursts, and the egg attaches itself to some object or
other in the water. In some species the yolk-membrane is covered
with cloven ramiform processes, as if with spines. After two or
three months the young one is visible. The conical excrescences
which arise higher up on the body at the base of the arms, and
which are perforated at the point, contain spermatozoa ; and may, to
a certain extent, be considered to be external testes*. These genital
^ See A. Fabbe, Obtervatioiu on the nUnute Mruelvre qf ike higher farmt of Polypi,
PhU, Trana, 1837, pp. 387— 4«6. PI. xx— xxvn.
> MUiNS Edwabds, Ann. dea Sc, not, sec. S^rie, iv. Zool. p. 338.
• PkU. Trans. 1834, p. 369.
^ The egg of Bydra was figured long ago by Bobsbl, Suppl. Tab. lxxxol fig. x a
and fig. 9. See also the figures of EHBXiraxBa and £bdl in Waonbr, Icon. Zootom.
Tab. xxziY. figs. 8 and 10, and of Laubbkt, Rackmikea 9wr VHydre et VEponge deau
dowe, Paris (1844), PI. n. Here figs. 9—14 the exclusion from the egg^ is figured,
which had also been observed by Pallas : " Ovtda avtumno generare Hydras 6baerwau'm
etc.. . polypi compendium per kyemem dv/raturum conHnentia. — Hone par ovula propa-
gationem bia meit oenlit petfeckm observan.** Mench. Zoophytor. p. iS.
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POLYPS. 69
organs may exist at the same time, and in variable number, in one
and the same individual. And many other Polyps are also henna-
phrodite. In others again the sexes are separate: whether both male
and female individuals occur on one and the same stem {Monaeciaj
a^ in Plants), or one Polypstock bears only males, another only
females {DicBcid). The last is the case of VereUllum. In the
Bryozoa^ Mancecia appears to prevail imiversally, yet so that (to
judge from the investigations of Nordmann in Tendra zastericola^
and of Van Beneden in AlcyoneUa) the cells which contain Polyps
with eggs are more numerous than those with spermatozoa. These
l)eculiar constituents of the seed (vid. above, p. 43), of which the
motions are so striking under the microscope, have, of late years,
caused the important discovery of the sexual propagation of Polyps ;
but for them, ovaries alone would now, as twenty years ago, be
ascribed to this class, especially as the seed-secreting organs {(estea)
are not to be distinguished in it, as to external appearance, from
those that prepare the germs (o^?arta)^ In those Anthozoa that
have, like ^e Actinue, a cavily of the body distinct from that of
the stomach, they are situated between or upon the partitions that
divide that cavity into cells (see above, p. 67). In Sertularia and
Campanularia most of the Polyps are without sex, whilst cells with
ova are developed in the axillae of the branches.
Propagation by spontaneous division does not occur in most
Polyps. In Caryophylla there is a complete longitudinal fission,
occasioning the dichotomous form of the Polypary, since two Polyps
come from one, four from two, &c. K the longitudinal fission be
incomplete, cells of irregular form arise, as in Mceandrina.
In most Polyps the power of reproduction is very great. Trem-
blet's exj)eriments on the fresh-water Polyp are well known : he
divided them longitudinally and transversely, and every piece
formed a new animal'. Koesel found that even the tentacles or
^ Sach 18 the cue also in MoUutea, nay eren in some fishes ; and in general the
sexual oi^gans in the animal kingdom possess a similarity in the two sexes, which was
observed by the ancients, and occasioned many fimciful appeUatiqns and comparisons.
' Hence Livnjbus borrowed the name Hydra for this animal genus, from a com-
parison with the ffydra of mythology ;
. . »ab ipto
DucU opes animumque ferro.
HORAT. Od, IV. 6o.
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70 CLASS II.
arms, when separated, grew into new Polyps ; an experiment which
did not succeed with Trehbley. The power of reproduction is
in ActmioB just as great ; they regenerate parts that have been cut
away, and admit of propagation by artificial division, as Dicquemake
especially has shewn by his experiments. ^
Before leaving the propagation of Polyps, we must notice those
late observations which indicate so close an affinity between MedtufCB
and certain Polyps, that in time probably a great revolution will
be made in the systematic arrangement of the animal kingdom.
In Syncoryrhe, for instance, and Coryne {Clava), and certain Cam-
panularim bell-shaped appendages or off-shoots have been noticed,
which at length are separated from the stem, and resemble minute
Medusas. Conversely, also, the observations of Sabs and of Yon
Siebold have shewn that Medtisce come from the egg under an
oblong form resembling that of infusories beset with cilia : these
move freely, at first, then fix themselves, lose their cilia, become
clavate, acquire arms, and perfectly resemble Hydra. These hydra^
like forms divide by transverse indentations, and separate into rings
from which Medvsw arise.
It is possible, therefore, that all Aydfra^form Polyps maybe only
imperfect forms of Medtisce. And if so, those animals which
K^AUMUB first named Polyps, would no longer belong to this class.
But on this supposition it is wonderfdl that Spermatozoa should be
observed in Hydra and Coryne : a fact that may cause us to hesitate
before we conclude, with Dujardin, that the eggs, described above
(p. 68), are BulbiUu At all events the perfect form of ^<2ra would
then be unknown ^
^ We cannot detail these obflenratioiu more particularly, without being difiVue
beyond our object. Let it suffice to refer the reader to LoTi^K Stoekk. Vetentk. A had.
ffcmdl. 1836 ; Wibomank'b ArchivfOr Naiwrpetch, v. 1837, s. 219—161, s. 311—316;
Ann, dea Se» not. tec. SSrie, Tom. XV. Zool. pp. 157 — 176. {Obaervationi mr le
diviloppemtni et lea metamorpKotee dea ffenrea Campawulaire d Syncoryne,)
Sabs, BeakrieveUer og JagtUtgdaer over nogle maerhdige eller n^je i ffavet ved der
Bergenahe Kyai levende Dyr, Bei^^en, 1835. *
C. Th. V. SiKBOLD, NmtetU Schrifien der natwf, Geadlaeh. in Daiaaig m. 1 Heft
1839, s. 16—35.
Sabs, Mim, aur le divfloppement de la Meduaa auritn et de la Oyanea eapiUaia.
Ann, dea Sc. not. aec. Sfrie. Tom. XTi. Zodogie, pp. 311 — 348.
Stssnstbup om Fortplantninff og l/dmkling gjennem vexUnde QeneratUmaraekker,
Kjobenham, 1841. 40. (Translated by 6. Bubk for the Ray Soe, from the German
Translat. 1845, On the Alternation of Oeneratuma.)
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POLYPS. 71
Of the nenroiiB system of Poljps little is known ; it has not
been described as a conneeted whole, but different obsenrers have
imagined that they had discovered ganglia or a neryons ring near
the mouth. From analogy it is probable that the nervous system,
where it exists, does form a ring ronnd the month ; and that the
threads, which Spix described in the pedal disc of Actiniof as
nerves, can on this acconnt scarcely be considered to be such* As
organs of sense, four colonred spots at the edge of the disc have
been observed in the Medusorform products of Byncoryne and Coryns
frttiHana of Steenstrup, which entirely resemble the parts that
Ehrenbesg considers to be eyes in Medusa. Still more distinct is
this oi^an in a form described by Quatbefaqes, which he names
EleuAeria dichotoma* Here this author found six eyes with a
hemispherical lens, a granular pigment of a red colour, and a
spherical projection of the integument closing the eye like a cornea^.
In Bryozoa the muscular system is most largely developed, and
serves principally to retract the animal within its cell. It protrudes
itself partly by straightening the alimentary canal, partly by means
of transverse muscular fibres which contract the diameter of the
visceral cavity, and in that way elongate it*. Traces of a muscular
system have also been met with in other Polyps.
As to the geographic distribution of Polyps, we have only
imperfect notices. Fresh-water Polyps, Hydra^ Alcytmellaj Phi'-
mateHaj have, as far as I know, only been observed in Europe ; but
from this to conclude that they do not occur in tropical regions,
would probably be premature. Still Ehbenbebq found no Hydrw
in Africa and Arabia. AcHnim are met with in all seas. The
Polyps with polyparies, which are almost all inhabitants of the sea,
are richest in species in warm regions; Fltutra, amongst the
Bryozoay is it seems an exception to this : the European species of
this genus are about as numerous as the foreign : these last come
principally from New Holland; Oorgonia occurs in all seas, but
Van BiKEDkN, MSm. mr le$ Campanulairet delaeUe tTOttende, Mim. de VAcad.
Rofole de BrtuDdlet, xrn. 1845. ^»*>* ^ 3c, futtur., tee. Sirie, Tom. xx. Zool, pp.
35<^— 375.
DuJAKDiH, MhMire mr U dMhppement dee Midusee tt dee Polypee hfdrairee, Ann,
dee. 8e, Nal, y SSrie, Tom. IV. 1845. Zooloffie, pp. ^57 — 281. PL xiv. XV.
^ Ann. dee Sc. Naiur. eee. Sirie. Tome xviii. p. 180. PL vin. fig. 6.
• MiDQtely detailed by Fabbk in BowerbafHsia, Phil, Traneact, 1837. pp. 393—39^-
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72 CLASS II.
nearly one-half of the numerous species is found in America. Of
swimming polyparies (Penfiatulai)^ many species are found in the
Mediterranean, the Indian Ocean and the North Sea, and the large
UrnheUarta of Greenland, which attains a length of six feet, is
remarkable. Of the stony polyparies, the greater number occur in
the seas of hot countries, as ex. gr. Madrepora, especially Astrwa,
Garyophylla, Fungia^ Mceandrina. Most of the species of the last
genus are found in the Indian Ocean and South Pacific. The Bed
Sea also has very many species from the division of these stony
l>olyparies. More than one-fourth of the known species of this
class were found there by Ehrexberg. Notwithstanding the
proximity, it seems that the Bed Sea, with the exception of some
species oi Actinia, has no species in common with the Mediterranean.
laia nobilis {Gorallium rvirum) appears to occur in the Mediter-
ranean alone.
Of many genera numerous fossil species are found, especially of
those whose species now live in hot regions. Thus the genus
Astrxa numbers more fossil than living species : and these petrified
remains belong principally to the Jura- and chalk-formations. Still
more numerous are the fossil species, as compared with those now
living, in the genus Turhirwlia. A genus which appears to connect
Fwngia and TurhiTiolia, CyatAophyUum of GoLDFUSS, has quite died
out, and occurs in' transition limestone. In the same way fossil
species alone are found of the genera Cericpora, Favaaitis, Ptistu-
lapora, Hetercpora, Catinipora and Aulcpora. The fossil Polyparies
prove, like all other remains of an earlier vegetable and animal
world, that the surface of our earth had in earlier times a higher
mean temperature than it has now.
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SYSTEMATIC
ARRANGEMENT OF POLYPS.
CLASS 11.
POLYPI.
AnimaL£I contractile, having an intestinal cavity, with distinct
terminal mouth, surrounded bj tentacles or radiating lobes, seldom
free, oftener affixed, aquatic ; UBuall j secreting a hard calcareous
or homy body {j?olypary)y and adhering to it.
Propagation is by eggs, gems, stolons. Usually compound ani-
malB are formed of many individuals cohering.
Section I. ^n/Aoisoa Ehrenb.
Aperture of the nutrient canal single. Tentacles contractile,
mostly without vibratile cilia.
Order I. Hydriformta.
Tentacles of variable number. Nutrient canal excavated in the
parenchyme of the body, not surrounded by an abdominal cavity.
Family I. Hydrtna. Naked, free, spontaneously affixed, with
deciduous progeny.
Hydra L. Body cylindrical, narrowed into a pedicle, simple or
ramose from the progeny not having been detached. Mouth
crowned with tentacles, round, contractile, in a single whorl.
Armed-Polyp: Sp Hydra viridis, Tbemb. PI. i. fig.j Robs. InB, iii.
Polyp, Tab. 88, 89. This green species was the first which Tbkkbley
found ; it is rarer and smaller than the others. — H Gruea, Trehb.
PL I. fig. 2 ; — H /tucay Tremb. PL i. fig. 3, 4, polype dt hngs bras.
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74 CLASS II.
BoES. InB. III. PdyjK Tab. 84, 85, 87 ; Tbemblbt performed most
of his experiments on these hust two species. Oomp. what is said
at pp. 67, 69.
On this genua compare the work of Tbkmblet, noted in p. 6o, and of
Laurent, p. 68. The first discoverer of the genus was Lxkuwsnhoick, who
probably observed Hydra grUea. He announced this animal in a letter to
the BoySd Society of London, of Dec. 35, 1701, observed its propagation
by gemmation, figuring it with two young ones, and especially described
the great contractility of the arms. PhU, Trans. No. ^83. VoL xxm. for
the years 1701, 1703, pp. 1304 — 1311. This description, however, had
been forgotten when Tbemblbt discovered the animal a second time.
Clava Gmel. Coryne of authors, not of Gaebtn. Body cla-
vate. Ten&cles scattered.
These animals are marine, and adhere to different bodies. Propa-
gation is effected by buds of a round or bell-shaped form, which
contain ova or spermatozoa, and which occasionally detach them-
selves from the stem on which they were developed, swim freely
about, and resemble small Medusse.
Sp. Clava parasitica Gm., Hydra mtdtieomiSf FobskIl, Icon. Rerum naiur,
HawnicBf 1 776. Tab. xzvi. fig. b, B. Cor. squamata Mukll. Zoci. Dame. Tab.
IV. about three lines, according to Ratheb, }inch long. Rathks dis-
covered Spermatotoa in the swellings called ^Si^iiaiiMe by MuBLLUt; vid.
Wiiomann'b and Erichson'b Arckiv. f. Naturgesck. 1844. s. 155—165 ;
Awn. des 8c, Nat. ^^"^SSric n. 1844. ^^- PP> -^^ — ^'o; Waonbb found
eggs in these swellings, Icon. ZooUm. Tab. xxxiv. fig. 16. Thus the sexes
appear to be distinct.
Eleutheria QuATREPAGES. Body hemispherical, hollow. Ten-
tacles six, divided into two retractile branches, terminating in a
round head. Six eyes placed at the base of the tentacles.
Sp. SUmkeria dccAotomo, Da Quathstaois, MiiMre sur VSleuAme dieho-
tome, Ann. des 8c, not, see, Sirie. Tom. xvn. 184a. Zoci. pp. 373 — 288,
PI. VIII. Discovered in the Atlantic ocean, on the shore of the Islands
ChoMsey, Dep. La Mameke, \ millim. in diameter. There were eggs in the
posterior part of the body. This form is probably a free bud for the propa-
gation of some species of drryne, Sertidaria or Tubtdaria. See Y. Bensdsh,
Bulletin de VAcad. Roydle de BrtoDeUes, Tom. xi. No. 10. Quatrkfaoks, ib.
Tom. xn. No. «.
[Genus PedtceHarta MuELL. delendum^]
^ Pediculated organs with three valves are found between the spines and tentacles,
or suckers, of the sea-urchin, which MusLLSB took to be parasitic Polyps, and of which
he formed the genus PediceUaria.
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FOUTFL 75
Family 11. Seriularina. P0I7P8 affixed hy a membranaceonB
paUium secreting the PolTpary, or loiicated, induded in a tubnle
or cell.
Hydractinia Van Bened., Echinochorium HA8SALL, Dysmor--
phoaa Phiuppi, Synhydra Quatrefages. Polypfl of two sorts,
sterile and fertile, set on an incrosting polyparj. Tentacles nnmer-
ons. The fertile polyps without month.
Sp. Hydradmea lactea, IHanwrphoaa eonchieolay Phiuppi, Sf/nkgdra paratUm
QUATRIP. Ann, da 8c. mU. Me. SSrie. Zod, Tom. xx. PL Tm. ix., Hab-
SALL^ Ann. and Mag. of Nat. ffitk VoL TiL PL x. ftg. $, PmuFF^
Whom. u. Ebiohs. Arekh, Tm. 1841. Ttl. i. fig. 5. Tiirbiii«te sheila are
Offten found incnufted with a greyish-brown finn oovering, difficult to remove:
this IB the common body to which the several polype, some miUimeten long,
are attachedj which have of late years been/described by diflerent observers.
QuATBSFAOKS foond his specimens on the shells of Turbo and Buocmumf in
which the hermit-crab (Poffurui) had taken up his abode ; so aJso BAflTBB
previously, natwurk. UiUp. I. Tab. in. fig. 5. According to Quatbsfaois,
it is by no means established that all the above-mentioned names refer, as
Yak Behkden determines, to the same animaL
Coryne Gaertn., Stipula Sars, Syncoryna Ehrenb. Polyps
with scattered tentacles, nodiferous or globose at the tip. Polypaiy
papyraceous, suhramose, with polyps at the extremities of the
tubes, not retractile within the tubes.
The name Coryne of Gaebtneb, Pallas Spicileg. Z06L Fasc. x.
1774, pp. 36, 40, 41, has by later writers been perversely given to
other kinds, and the original typical species been named Syncoryne.
Sp. Coryne punlla GtASBTv., Coryne ^nduio»a Pall. SpieU. Zool. x. Tab.
rv. fig. 8, &c Compare on this genus Lovi^K, Konff. Vdauk. Akad. Sand-
ling. 1835; Wdegm. Arckw, m. 1837. pp. 31 1—316, Ann. dea Se. Nat.
a SMe. Zool. XV. p. 170, PL vni. JoBsmos, Bid. of Brit. Zoophytes, pp.
3^-41, PL n.
Tuhtdaria L. (in part). Polyps with a double coronet or whorl
of tentacles, the upper surrounding the mouth. Polypary affixed
by the base, tubular, gelatinous or membranaceous, with polyps
terminal, not retractile.
a) Simple TtdndarioL
Sp. Tubularia indwi»a L., Tubvl. calamaris Pall., Jussdeu, M4m. deVAoad.
Jtoyaie det Sc. 174a. p. 196. PL x. fig. 2. A, B; Ellis, CoraU. PL xvi. c ;
LiSTKB, PhU. TroMoct. 1834. p. 166. PL vin. % i, &c. ; Johnston, Hist.
Br. Zooph. p. 48. pL m.
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76 CLASS II.
h) Branched TubularuB, [Gemm ^wiendrium Kosl]
Sp. TmtmL nmam L^ Kll Cbmff . FL m. fig. a. PL xm. a» A, fta
Corymarpka Bams.
Comp. 8aB8» Sakrimbe eg Jm^tLangdatr oter tugi^ m/mahdi^ dUr mije i
Htmi ved der Bergentke KfM lemmde Dfr. 60)^01. 1855- PL i. fig. 3.
FoBsn and GooDS^^ Om ike Coiymorpkm mmkm$, Amm. mmd Mag. nf NaL
BiM. T. 1840. pp. 909—315 ; JoEnrov, B*M. Br. Zoapk, pp. 54— 5^- Pl-
Tn. figa. 3—6.
Pennaria GrOLDP. Polyps clayate, the club with scattered ten*
tacles globose at the extremity, and with a whorl of longer tentacles
at the base. Polypary ramose, with branches alternate polypi-
ferons on one side (with Polyps second.)
Sp. Pemmaria Cavolmm, SatmUmia petmmria Gatolivi, pp. 134—159. Tbb. v.
Campamdana Lah. {SerhilaruB species L., Ehr.) Polyps fim-
nel-shaped, with month situated at the extremity of a retractile
conical tubercle. A whorl of tentacles, numerous, warty, with
dart-cells at the base of the tubercle. The Polypaiy corneous,
tubular, branched, with cells campanulated, pedunculate, the pedicle
long, continuous with the stem. The terminal cells sterile, the
axillary oviferous.
Comp Lister, FhiL Trans. 1834, p 372 Ac. ; LovAf, Kong.
Vetensk Akad. HandL, Wibgmanh's Arehiv, iil b. 249—262. Ann.
des Sc. not 2e sericy Zod. xv. p 151 ; Yan Bbnedbn, Mem. swr les
Campamdaires de la cdfe d'Ostende. Bmxelles, 1843, 4^ {Mem. de
VAcad. de Bruaeelles, xm.), Ann, des Se, not. 2e serUy Zod. xx pp 350
—369. PL 13 (Extract).
Sp. Cam^aimilima diekoioma Lam., S&rUdaria geikiculeOa L., Elub, CoraU.
PL xn. No. iS A, a. C, c. PL xxxym. fig. 5, fto.
BerHilaria L. (exclusive of several species). Polyps funnel-
shaped; tentacles hispid, numerous, arranged in a whorl at the
base of the mouth. Polypary corneous, simple or ramose, with
cup-shaped cells sessile, or subpedunculate, with short pedicle,
distinct from the stalk. Polypiferous cells sterile ; others fertile,
oviferous, scattered over the stalk and branches, situated usually
near the base.
The Polype of this genus do not differ from those of the former,
but the cells are non-pediculate, or the very short pedicle is dis-
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POLYPI. 77
tinctly inaerted into an indent of the stenL The last fonn the
genus Laomedea Laxoubouz. Those which have cells entirely
without pedicle may be thus divided :
a) with oeUs hifiurious or scattered {Serhtlaria Lax.)
L., Ell. Corall, PL v. No. 8. fig. a, A, Liarra, PkU. Tntnt. 1834. PI. viii.
fig. s» &C. JoHNBTON, JBr. Zoopk. p. 66, PL zi. figi. 3, 4, and p. 75, PL
xin. fig. I.
6) with cells yerticillate {Anlennularia Lax.)
Sp. Sertttlaria animnina L., Bll. Corall. PL IZ a.
c) with cells secund {Plumularia Lax.)
Ftp. Sertularia plttma Ell. Corall, PL vii. fig. b, B, to, Jomxtnon, Br,
Zoopk, p. 9«, PI. zzni. figs. 1—3.*
Order IL OctacHnia.
Tentacles eight, pinnate. Nutrient canal contained in a distinct
abdominal cavity^ connected with it by interposed lamellae.
Family III. Xenina. Common body, fleshy or membraneous,
affixed by the base. Polyps not retractile, with pinnate tentacles.
Xenta Sav. Common body growing upwards into stems divided
at the top, branches short. Polyps &sciculate, collected at the
extremities of the branches into globose heads, or umbels.
Sp. Xenia umbdlala Saviont, D6$eripHon de VEgypte, Polypea, Tab. I. tig.
3 ; ScHWSiGOKR*a Beobaehtungen a/af natwrhitt, Jteisen, Tab. y. fig. 48 ; in
the Bed Sea.
Anthelia Sav. Common body, membraneous, plane, spread over
marine bodies, stoloniferous. Polyps standing out, erect, crowded,
at the surface of the membrane.
Sp. Anihdia glauea SATiaHT, I>S9cr. de VEgypte, Polypa, Tab. i. fig. 7.
Note, C^nus RM/zooxnia Ehbenb. is founded on a figure of
Zoamtha thalasacmtha of Lesson in the zoological plates of Du-
perbt's voyage (Voyage mUov/r du Monde swr la Corvette la
Coqttille, pendant lee annees 1822 — 1825). The common body is
^ Many species which have been refened to SerhUaria, belong to the Bryotoa,
Anumgat them an thoee which Lamabok has brought together under the genus Serio*
huria. See Yak dkb Hoitbn's ffandb. der JHerkunde, first edition, i. p. 76.
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78 CLASS II.
made up of stolons, connecting tubes erect, ventricose, striated, each
containing a Polyp. Wliether the Polyps ara retractile or not, does
not appear. Oomp. genus Evagora Phiuppi (p. 79).
Family IV. Hahyonina. Polypary fleshy, spongy, perforated
by many canals, and crowded with microscopic calcareous spicula.
The Polyps associated in the polypary, retractile, with tentacles
pinnate.
This fiimily has its name from the genus Alcyoniwra L. called in
Holland Zeeachmm or Zeekv/rk (sea^foam or sea-cork). Under the
genus Ahyonivmh of Linn.£US were comprised species which, like
Ahsyonium Schlosseri, belong to the Molluscs (Asddice), as Sa-
VI6NY has shewn ; other species are Bryozoa {Alcycn. gdcUinosum),
The genus Alcyonium of Lamarck contains plant-like forms without
polyps. To these spongy plants belongs also the genus AlcyoneUiMn
QuoY (EuplecteUa Owen). Consequently there remain for this
family those species alone which Laicabck has united under the
genus Lobvlarioy and probably it would be well, according to the
suggestion of Schweigoeb, to reject the name Alcyonvum altogether,
for the sake of avoiding confusion.
On the sponges comp. SoHWSiaoKB, ffcmdb, der Naturg€8ck, d&r skeUttr
loten ungegliederUn Thiere, s. 370^374, R. £. Obakt, in Edinb. PhUot.
Joum. VoL xin. p. 333, H. F. Link, Ueber PJUtnzenthiere Uberhavpt und
die dagu gere(^neten ChiodcKse bemmden. Phynk, Abhandl, der Ahad. der
Wiseenech, eu Berlin a. d. Jahre 1830. s. 109 — 123.
Ahyonidia Milne Edw. Polypary simple or ramose, with a
basal portion coriaceous, a terminal polypiferous, soft, retractile by
invagination within the former. Polyps with tentacles having
pinnae, which are hollow, in a single row at the margin, retractile
separately.
Sp. Alcyonidda elegane Milmb Edwabds, Ann, dee. 8c. not. w Sir, iv, 1835.
pp. 333 — 333. PL f a, 13. la the Mediterranean at Algiers.
Nephtoki Sav. (according to Ehrenb. to be written Nephthya).
Polypary ramulose or shrubby, with Polyps retractile within warts
armed with spicula.
Sp. Nephtaa inMmUnaia Blaikv., Nephthifa Savignyii Ehbknb., AmmoihM
Chabrolii AuDouiN, DUcr, de VEgypU, Polypes, PL n. fig. 5. To the flame
genus belongs Spkongodee cdona LssBON, IlltutraHons de Zociogie, PL xxi.
which seems scaroely different from Alcyonium pni/dfum Esfsb, Alcytm,
Tab. xvi.
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POLTPI. 79
AmmoAea Say. Fcdjpmy ramnloee or shmbby, with Poljps
retractile into unarmed warts, clustered on the last branchlets.
Sp. Ammolhm viraceM Satight, l>^fer. de VBgypU^ Polypet, PL n. fig. 6.
Sympodtum Ehrenb. Common bodj, membraneous, efiuse, with
Polyps retractile into unarmed warts that project only slightly,
without stem.
Sp. Sifm/pod, fuUginotum Ehbinb., Anihelia species Audouih, Dfyer. de
r^ypte. Polyp, PL L fig, 6.
Some AiUkdia have retractile Polype. Thej cover ytaioxta marine bodies
as an incrostatioiL One species of this genus was described by Pallab as
the orost of a Oorffonia: Oorg. eoraUoidei, Blench, Zoophytor, p. 199,
EflFSB Gorgon. Tah, xxzn.
Evagora Philippi. Polypary incrusting, formed of stolons
conjoining the several Polyps. Polyps with a basal portion harder,
coriaceous, a terminal retractile, soft.
%p, Evagora roeea Prilipfi, Wibomanh n. Ebichson Arekw f, Natwrgeeck,
▼m. 1843, L s. 36. Taf. I, f. 2, c. — Zoawtha ihalastantha Lbssok (see above,
p. 77) appears to be another laiger species of this genus.
Alcyonium Cuv. Milne Edw. {Lobularta Lam. Alcyanii
species L.) Body fleshy, turgid, usually inciso-lobate, covered with
Polyps scattered.
The separate polyps are entirely retractile within tiie common
body, formed by the union or concretion of the external covering
of the polyp& This is thick and spongy, and contains a great
quantity of small irr^ular crystals of carbonate of lime. PropagSr
tion is by eggs and buds (^emmce). The form and size may be very
different in one and the same species, so that the distinction of
some of the species proposed by authors is imcertain.
Sp. AlcyoKtum lobaiwn Pall., Ale, digUattm L., Juasnu, Mim. de VAcad,
des 8c.de Parie, 1742. PL IX. f. i. A— J ; Ell. Corall. PL xxxin. fig. a, A;
Sfix, Ann, du Mas, xm. 1809. PL xxxiu. fig. 8 (named Ale, exoi); La-
MODBOUX, HieL dee Polj/piers JUxtbUa, PL xiz. fig. 4, PI. xm. PL xnr.
^, I, Johnston, ffiet. Br. Zooph. 174. Pll. zzxnr. zxxiv*. This species,
called by the Dutch fishennen, according to Pallas, dooden manehand or
duimen (deadman's hand or thumb), occurs in the North Sea^ and attains a
size of 0.14 — 0.1 miUim. ; the form is very irregular, which, as it seems to me,
the name given to PaUas well indicates; the colour is brown-yellow. — Ale,
palmaium Pall. Ale. eacoa L,, Bohadsch De quibued. anim. mar. Tab. ix.
f. 6, 7. EsFBB Alcgon, Tab. n. &c. This species occurs in the Mediterra-
nean ; it has the form of a little tree cr shrub, and tbe branches are coloured
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80 CLASS II.
red. It has been specially inveetigsted by Milkb Edwabdb^ mmI very
beaatifbOy figured in his Obtervatiims sur U» Alcyon*. Ann, da 8e. not. le
Sir. Tom. iv. Zool. pp. 333— J43- PL I4» '5-
Family V. PennatuKna, Stem free, fleshy, containing inter-
nally an axis stony or homy. Polyps naked, aggregated on the
common stem, with tentacles pinnate or pennatifid.
Sea-Feathers (Polypi naiarUes & PenncB marinm). The opinion,
that these polypstocks swim about in the sea, appears to be un-
fomided. The stem is fixed in the ooze at the bottom of the sea,
or the polypary lies on the bottom ; it is only when the waves or
the fishermen's nets have broken the Pewnahda loose, that it swims
free in the water. Comp. W. Rapp Ueber Pclypen tc Actinien
s. 8, 34. Costa in Fbobief^s neueyoHzeny Bd. xxl Feb. 1842, s. 154.
Many species are phosphorescent: Pennahda phasj^uyrea, Pen.
griaeoy Pen. rubra {P. granulosa Lail), VereHUum cynomorium finom
the Mediterranean and Pen, argentea from the Indian sea.
The genus EncrinuSy placed by Lamarck amongst the searfeathers,
belongs to the Echinoderms, and is, as Ellis long ago remarked, a
species of star-fish with a stem. Nai. Hist ofCoraiL
A. Shaft pinnated in scales at the upper part, pinnae polypiferous.
Pennaiula L. (exclusive of species). Shaft fleshy, at the lower
part naked, at the upper pinnate, axis stony. Pinnae two-ranked,
patent, plicate, dentate on the upper margin.
Sp. Pennatula ffri$ea L., Pennaiula apinosa Lam., Albiki Annot. Acad, Lib.
I. Tab. vi. figs. I, a, Bohadsgh i>e quHmtd. animalib. mar. Tab. ix.
figs. I — 3, EsPBB PJUmeentk, Pennat, Tab. i. Pen. rvbra L., Pen, granu-
loma Lam., Albih. L 1. figs. 3, 4, Esfkb PjUvngmOh. Pennat, Tab. n. both
firom the MediierraQean.
Virgularia Lam. Shaft elongate, slender, naked below, pin-
nated above, with sub-stony axis. Pinnae small, unarmed.
Sp. Virgularia mirahiUt, Pennat, mirahiUa MuiLL. (not L.) Zooil, Danie.
Tab. XI. Cuv. M, Ant, Mi, Uludr,, Zoopkyt, PL xoi. fig. a.
B. Shaft simple, with polypiferous warts or papillae at the upper part
Funiculina Lam. {Pavonarta and Sdrparia Cuv.) Shaft elon-
gate, filiform, with axis homy or sub-stony. Polyps arranged in
series, secund or alternate.
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POLYPI. 81
a) Polyps secund {Pawmaria Cnv.)
Sp. Funieu^inaamtemiiHa,PemiatiUaquadrai^^
L., B0HAD6CH De guOnud. Anim, mar. Tab, IX. fig. 4 ; iq the Meditem-
nean, more tluin two feet long. [Found near Oban, Argyleshire, forty-eight
inches in length, by Ptof. YoKBMB. Yid. JoHVBTON, HiM, Br, Zooph. p. 165,
PI. XXZI.]
5) Polyps alternate {Scirparia Cur.)
Sp. Pennatvia mirdlnlit h.^
C. Shaft simple, Polyps scattered, sessile.
Veretilhim Cuv. Body cylindrical, fleshy, upwards polypife-
rous, with large Polyps. Axis like a ligament or osseous, short.
^. VtrMhim efMmarwm, PenntUula Cjfn/omorktm Pall. Mi»e, Zoei. Tab.
zm. t 1—4, Raff, Nov. Act. Acad. Camur. Leop. Carol. Mrfur. Ourioi.
xiT. 9. 1829. Tub. zxxvni. fig. i. Ebdl in Waonsb leon. ZooUm. Tab.
xxziv. fig. I. Mediterranean, Ac.
D. Shaft simple, polypiferous at the extremity only, polype grouped in
an umbeL
UmheUularta Lam. Body elongate, slender, with a long osseous
axis. Polyps large, terminal.
&p, UmbeUaria gnmdandieay PennahUa enermui Pall. ; Ell. CoraU, Tab.
xxxvn. taken in very deep water at 79* N. L.
K Shaft short, cylindrical, dilated into a flattened reniform expansion,
which is polypiferous on one side,
ReniUa Lam.
%). RenMa americana Lam., Pennatula renfformis Pall., Schwbioobs JM.
auf naturhut. Reiten. Tab. n. fig. 10 ; — BeniUa vkiacea Quot et GADf abd
Voyage de VUranie, Zoologie. PI. Lxxxvi. fig. 5—7, Cuv. R. Anim. 4dU.
iUuMr^e, Zoophfft. PI. xci. fig. 3.
1 Scirparia or Seirpearia Ou/v. is said to be distinguished by Polyps plaoed alternate
on the two sides. This genus is founded on Pcnnaiida mirdlnlu. Polypus mMubili$
Limr. Mui. Adolph. Prederici Regie, Hohnise, 1754. FoL Tab. xix. %g. 4. p. 96. It
is very possible that Limr iEUS afterwards mistook a foreign species (from China, see
AmaenU. Acad. iv. p. 157) for one from the North Sea {Fowna Sueciea, p. 543, "Aa5t-
lot in oceano Norvegico**), and this last may be VirgtUaria mMubUie. To me the genus
Seifparia appears very doubtful. The type which served for Linnaub' description,
was not known at Stockholm, as my friend Prof. Sundbvall wrote to me (4 July,
1846).
VOL. I. 6
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82 CLASS II.
Family VI. Tubiparina. Polypary calcareous, of parallel
tabes, close set, conjoined by transverse partitions. Polyps tubular,
the neck retractile, soft, the lower part indurated, forming the poly-
pary. Tentacles in single or double row at the margin, retractile by
involution.
Tviipora L. (exclusive of several species).
Sp. Tubipora mutica L., Tubularia TouBNXr. Inttit. Bei het^ricB Tab. 343
(the Polypary) ; for this animal and its organisation compare especially the
beaatlAil plate in Fbbtoinbt, Voyage de VUranie^ Zool. PL 88. The
Orffan-Ooral consiBts of cylindrical, hollow tubes, standing perpendicular
with transverse partitions. These last arise from a horizontal expansion,
which at the top of the tube surrounds its droumference radially. Hie
expansions connect the tubes together, and become partitions when the
tubes above them b^gin to grow. From this elongation of the tubes
their jointed form arises, and when the growth ceases, they form a new
transverse expansion round the wall of their aperture. Ehbekbbbo has
distinguished the spedes of this genus more accurately : they are usually
comprised under the collective name of Tubipara fMuica, The Polypary in
all the species is purple-red ; in the Indian species which Peboh {Voyage
avac terree AuatraUe I. p. 146), and QuoTand Gaimabd {Voyage de VUranie,
Zoologie, pp. 634 — 641 and PL 88) observed, the Polyps are green, in others
they are whitish or light red, as in those which Chahibso described (Nov.
Ad, Acad. Leap. Carol, N. C. Tom. x. p. 370, Tab. xxzin. fig. 2), and in
TMpora rubeola Quot {Voyage de FAstrolahe, Zool. TV. pp. 357 — 359),
GusBiN Iconographie, Zooph. PI. xzn. fig. i, where the fin-like indents
at the edge of the tetvUieula stand in a single row, as in Tubip. ffempriehii
Ehbenb., whilst in Tubipora musica Ehbxnb. to which Fbetcihbt's plate
quoted above refers, they form a double row.
To Tubipora fossil Polyparies appear to belong, Catenipora {egcharoidea)
and Syringopora Goldf. from the oldest limestone (mountain-lime).
Family VII. Corticata. Polypary fixed, ramose, its bark soft,
supplied with calcareous spicula or granules, polypiferous, its axis
hardish stony or homy. Polyps retractile, with tentacles having
a single row of small conical appendages at the margin, gemmi-
parous and oviparous, conjoined by canals creeping through the
bark.
The hofrhed^oraU (corticiftrei) of Lamabck form a division veiy
nearly allied to Alct/onvum and FenncUulina. The polypary is here
in its origin and mode of structure vety different from that of the
TtMporina, but on the other hand resembles that of the Fefma-
tulvna. The hard axis, which alone is usually preserved in collec-
tions, may be compared with that of the FenruUtdina; they are, in
a word, fixed FenruihUincB.
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POLYPI. 83
A. Axis Hany {Isidea Ehbenr)
litis L.
Corallium Lah. Shaft uniform, rigid, finely striated longi-
tudinallj.
Sp. CcToUium nidnuii Lam., /#» nobiUi L. Toubkbp. InatU. Bei herbarice.
Tab. ooGZXziz. (Axis), Espbb, I^fUnuenth. Ind. Tab. yn, vm. ; CAVOLiiri
PoUpi, pp. 33^47, Tab. n. Cutub R, Anim, 4dU, iOutirie, Zoofh, PL
80. Blood-eoral; in the Mediterranean, espeoiaUy on the African coast. It
is exported to the East Indies, and is aJso much used in Europe for neck-
ornaments. It grows on all sorts of marine bodies, even on other oorab,
and not only downwards, but in all direotioDS, increasing yery slowly ; it is
seldom more than a foot long. Hie streaks risible on the unpolished aids
are the impressions of vessels which run in the bark, and form a oommuni-
cation between the different polype.
MeKUea Lam. Shaft knotty, genicula tumid, ramiferoos.
Sp. MdUoea oehracea, Itis oehraeea L., PaIiL., Natuurl. Hid, der plamidieren,
door BoDDAXBT, Tab. vn. Mxuiv, Beite urn die Brde, m. Zool, Tab. zxxiz.
in the Indian Ocean.
Isis Lam. Shaft with jointed axis, nodes stony, striated, rami-
feroos, intemodia homy.
^. Isi$ kippwris L. ; EsPEB l^merOh, Ind, Tab. I— m.
Mopsea Lamour., Ehrenb. Shaft with jointed axis, nodes
homy, ramiferons, intemodia stony.
Sp. Mopiea dkhoiama, Ins diekokma L. ; ESFKB PJlanun^, Ind. Tkb. v.
Note. Here also belongs Ina donffoia, Espeb I^nzenth, Ind. Tab. vi,
according to two specimens brought by the noble V. Sixbold firom Japan,
which are preserved in the Leyden Museum, and agree with Espxb'b
figure. Is the lame species also found in the Mediterranean, as Philippi
supposes, who refers to it Moptea Mediterranea R188O ? See Wibomann
u. Ebichsok'b Archiv. Tin. 1843. s. 38.
B. Axis homy {Cerato-coraUia or Gorgoma Ehrenb.)
Gorgonia L. (exclusive of species of Antipathes). Stem with
axis homy, distinct: the crust polypiferous, fibroso-calcareoiis, per-
sistent.
Seorshrvh, HomrplarU. These horn-plants grow with stem and
branches upwards ; the latter are usually situated in a plane, and
often coalesca Many earlier and later writers have believed the
stem to be a plant, on which Polyps had fixed themselves. (Ds
naiwra vegetabiU Oorgoniofrum, auctore G. L. C. Gravenhobst,
Obxh's Isis 1823. & 724. Beetle Aoademia delle scienze di Torino
6—2
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84 CLASS .II.
T. xxvL) The species are very numerous, and many might perhaps
by closer investigation be better defined Laicoubotjx and Ehben-
BEBG haye formed different genera, which by the last especially have
been distinguished by the arrangement of the Polyps.
Subgenera: Prymnoa Lamour., Ehrenb. Mwricea Lamour,
Ehbenb. Etmicea Lamoub., Ehbenb. Plexaura Lahoub., Ehrekb.,
Gorgonia Lahoxtr., Ehrenb., Fterogorgia Ehrenb. — ^A new genus
Bebryee Philippi appears to be distinguished by non-retractile
Polyps.
Sp. ChrgoniaJUibdUim L., Ell. ChraU. PL zxvi. fig. A — O. Sea-fan, MermauTt
fan, in different seas.
AnttpcUhes Pall. {Oorgonice Spec. L.) Stem with axis homy,
distinct, covered usually with minute spines, with bark polypiferous,
gelatinous, deciduous.
Sec^shrub, The bark which is gelatinous, not calcareous or
fibrous, is missing in specimens taken from the sea : hence, when
preserved in collections, they resemble branches of dead wood.
EHRENBERa thiuks AfUipcUhes ought not to be joined to Gorgonia,
and that it probably belongs to the Bryozoa. He refers to later
communications, which have not yet, as &r as I know, been pub-
lished {pie CoroMevUhiere dea rothen Meeres, s. 113 in a note.)
Milne Edwards does not hold this opinion, Lamarck ffisL naL
dea Am. s, v, ii. p. 684. According to Gray the Polyps of Anti-
pathee, which he investigated in a specimen referred by him to
ArU, dichotoma Pall, have six arms, but, with the exception of
this strange anomaly, agree with those of Gorgonia, Proceedings of
the ZooL Soc. of London. 1832. p. 41, 42.
Sp. AnHpatket spiralU Pall., Esfsb PJlamenth, Antip. Tab. vni., Pallas
Plantdieren by Boddaebt, Tab. vi. fig. s—Antipath. myriophyOa Pall.,
EsPiB L L Tab. z» GuimN Iconogr. Zoophyl. PI. xxin. fig. i. ftc.
Order III.
Polydctinia {ZoocoraUta polyactinia, PhytocoraUia polyactinta,
and PhytocoraUia dodecactinia Ehrenberg).
Polyps with twelve or more non-pinnate tentacles, simple or
aggregate. Nutrient canal suspended in the cavity of the body, by
means of lamellsB forming partitions. Aperture of the nutrient
canal single, external, supplying the office of mouth and of anus.
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POLYPI. 85
Section I.
Tentacles twelve. {PhytocoraUia dodecactinia Eurenb.)
Family VIII. Madreporina {Madreporina and MiUeporina
Ehrenb.) Polypary secreted by the Polyps, stony, supplied vdth
polypiferous cells, usually ramose or expanded, lobate. Tentacles
short.
Madreporalt. (exclusive of many species), Lam. (Portia Ejusd.
Eeterapora and Madrepora Ehrenb.) Polypary stony with cells
circumscribed, lamellose, often prominent, with porous interstices.
Sp. Madrepora palmaUt, Ifeteropora palmata Ehbinb., Madrepora mmieaia,
w. EsFKB PfiameiUh. Madrtpor, Tab. u. On the animal of this ipeciea
oomp. Lebubub, Mim. du Mm, vi. pp. 190, 191, PL ZYn. fig. 18. Madre-
pora ahrotanoidei, Madrepora muricata Pall., Quot and Gaimabd Vo^offe
de VUran, PL zcvi., GUBKIK Iconogr, Zooph. PL xzin. fig. io,^Madre-
pora podEUfera Lam. &c.
PodUipora Lam. Polypary stony, ramose, with cells of slight
depth not lamellose, contiguous.
Sp. PociOipora damkomdg Lam., Espsb P/aneenth., Madrtp. Tab. zlvi.
and XLYi A. &o.
Genus NvJlipwa Lail Syst^me des Ani. 8, vert^bres 1801. p. 374.
{MiUepores with pores not evident Hist not des Anu s. verUbres IL
p. 31 1) according to Ehsenberg is in port to be brought here.
SeriaUypcra Lam. (in part). Polypary stony, ramose, with cells
disposed in longitudinal rows, with margin slightly prominent.
Polyps with the structure of the dodecactinia, destitute of tentacles.
Sp. Seriatopora eubulata, MUUpora lineata h,, Espbb PJlameiUh, MiUep,
Tab. ziz.
MiUepara L. (exclusive of species) Polypary stony, ramose, with
cells deep, obsoletely or not at all lamellose, separate, scattered.
Sp. MUUpora alcieomii L., Espbb I^ntenth, MiUep. Tab. v, vn, xxvi, &a
(Here also the Polyps appear not always to possess arms). Many species
which were formerly placed amongst the MUUpora are now ranked in other
genera. — MUlepora truneata, the genus Trunctdaria Wibomaitn {ffandb,
der ZoU.), Myriopora Blaikv. belongs to the Bryozoa, Ehbxnb. Die Coral-
lenihiere des rothen Mteree, ss. 126, 154, MiuvB Edwabds in and edition of
Lahabck H'ui, Nai, des Am, s. v, p. 306.
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86 CLASS II.
Section II.
Tentacles numerous, exceeding twelve.
A. Polyps secreting a stony Polypaiy , by which they are
affixed {PhytocoraUia polyacHnea Ehrenb.)
Family IX. OceUina Ehrenb. (and Dcedalina ejusd. in part).
Cells circumscribed.
Genera: GcMryophyUia Lail, Oculina Lam., ExpUmcvria Lail,
Cladocora Hempb. and ^EssESB,,AnthophyUum Schweiggeb, Ehbenr,
AstroMb Gil {Astrea Lam.)
Sp. CaryophyUia ramta Lak., Madrepora ramea L., OcuUna ramea Ehbsnb.,
TouBKBFOBT Inttit. BH kerbaricB, Tab. COOXL, Madrepora, EsPiB PJhn-
zerah,, Madr^por. Tab. ix. x a., Milne Edwards in Cuvikb J2. AtU, id,
lUutMe, ZoopK PI. Lxxxm. fig. i, i a, i b (with the animals). — Cwyopk.
eaUcularis, Cladocora calyctda/ria Ehbknb., Cavolihi Polipi mairmL Tab.
m, fig. I — 5, pp. 48—58, Milne Edwabds in Cuv. R, Ani. id. iUtaMe,
PL Lxxxin. fig. 2,
Hate, The too numerous genera in this ^mily, severed from the
genus Madrepora Linn., might perhaps be properly referred to two
genera, Oculina and Astrcea, Add Monomycea Ehrenb. with a
solitary star.
Family X. Oyroea {DcBdalina Ehrenb. in part). Cells con-
fluent into sinuous furrows, on both sides lamellose.
t Stars oonccbve.
Moeandrina Lam. Mceandra Oken^ Ehrenb. Polypary
stony, hemispherical, on the convex surface stars winding, con-
tiguous, lamellose.
Sp. M(Ba/ndrina eetdtriformM, ^anglicb Brain-tUme; — Mceandr, labyrintkwh
Mut, Bederian, 1716. Tab. xxvi. fig. i, Madrepore, SAVioinr Dieer, de
VEgypU, Zoopkyt, PI. y. fig. 4, ftc. In this Polypary the confluent stara or
cells {ambulacra), with their transverse plates, resemble the monntain-
chains as usually engraved on geographical maps.
Agaricia Lam. Polypary foliaceo-lobate, on one side only fur-
nished with furrows or stars in lamellas.
Sp. Agaricia depkoMahte Ehbsnb., Esfbb PfioMmJtk, Madrep. Tab. zvm.
Ac.
Pavonia Lam. Polypary foliaceo-lobate, with leaves com-
pressed, on both sides stelliferous.
* Lehh, der NtAurtftaek, iii. i. s. 70. 18 15.
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POLYPI. 87
&p. Pavonia agturicUe$, Madttpon agtmeket L., JSenM PjUmamik,^ Madrep,
Tab xz, CvY. Jl. Afd. 4dU. iOiutr,, Zoopk, PL LXZXiv. fig. i;— Pov.
lactuca, Madrep, laehiea Tall,, Ebpkb lyUuumtk,, Madrep. Tab. xzxni.
A, B^ QuoT et Gaim. Voyage de tAairolabe, PL xviu. fig, i, copied in Cur.
IL Am, id. UlMtir,, Zoopk. PL Luuuv. ^. i. The aninuJ figured and
deMribed by QnOT and Gaucasd has round the moath tnberolefl and no
arms, is veiy flat^ and reaemblea an Aetima; Ebbxvbibo places this ipeeiea
with Maandrina pectinaia, Meeandr, areotaia, and some others under a
new genus Mameina, The singularly flat and thin leayes of this Polypary
have given occasion to the name of Bndwe-Oorol (Zoctwea).
ft Stars convex,
Monticularia Lam.
B. Polyps secreting internally a hard body (Polypary stony,
not affixed).
Family XI. Fungina Ehrenb.
The stony polypary is here an internal induration of the animal,
and is by Ehrekbebo compared with the calcareous plate of Cephalo-
poda (the back-bone of the Sepia).
Fungina Lam. Polypary free, orbiculate or oblong, hemi-
spherical or conical, above convex and lamellose, with an oblong
central lacuna or gap, below concave and ragged. Star single,
occupying the upper surface with lameUe denticulate or rough on
the margin.
SecMnuskroom. The numerous plates, running from the center to
the drcumferenoe, give this Polypary some resemblance to a mush-
room, in which however the plates are situated beneath the cap.
Some have an elongated form, and hence, in the names they bear, are
compared to moles or slugs.
The Fungioe lie in clefts of rocks and cavities of coral-reefis,
surrounded by branched corals, so that the force of the current is
broken whilst the access of eearwater is not precluded. The older
specimens are quite free : but younger ones are seated on a stem,
on rocks, or sometimes are fixed to the dead remains of other
FwnguB ; in the pedunculate state they resemble the genus Caryo-
phyHia Laic. The stem is at first hollow, and is afterwards filled
with calcareous coral-substance; the disc becomes laiger, and at
last the stem entirely disappears. S. Stutchbubt, An Account of
the Mode of Growth of Yoimg CorcUs of the genue Fungia, Trane-^
aH. of the Linnean Society of London, voL xvi. 3. p. 493—498.
1833.
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88 CLASS II.
In most the entire Poljpary belongs to a single Polyp. In some
species no tentacles or arms are distinguishable ; but in others there
are numerous thick, conical arms, irregularly scattered; in the
middle the large, transverse oral aperture is seen. The animal
surrounds the Polypary as well beneath as above. See the figure
of Ftmgia crassUentactUata QuoY and Gaihabd, Voyage de r Astro-
labCy Zooph, PL xiv. £ 3, 4, also transferred into the illustrated
edition of Cuvieb, R, Ani, ZoophyteSy PL lxxxii. ^g, 1. Guj^rin,
Iconogr, Zoophytes^ PL xxiii. fig. 6. In other species, according to
the observations of Eschscholtz, Quoy and Gaimabd and others,
many animals are grown together ; the oral apertures, here without
tentacles, lie partly in the oblong median depression of the Polypary,
partly between the plates. These form the genus PclyphyUia Quoy
and Gaimabd, and HerpolUha Esghsch. {fferpetolUha Leuck.),
HaUgloBsa Hehpb. and Ehrenb.
See on this genus F. S. Leuckabt, Obwroai, Zool. dt ZoopkytU Coral-
liis, ttpeeioHm de genere Fungia. Cum TabuUs IV. ofri incias. Friboigi
BiiBigavorum. 184 1. 4to.
Sp. Fvngia agariciformis Lax., Madnpcra fungitea L., Mm, Bederum.
Tab. XXVI. fig. 3. FoasK. Icon, Rer. natural. Tab. xlh., Esfib PJUm-
zenih, MadATtp, Tab. I. Lbuokabt 1. 1. Tab. iv. fig. i — 3, round, with fine
toothed laminsB ; the animal had been observed before by FobbkIl, and
varies in colour ; QuoT and Gaiicabd have figured it entirely red, if indeed
their figure refers (as Ehbbnbebo concludes) to this spedee. Voyage de VUror
niCf Zool, PI. xovi. fig. I, 1. — Fungia limacina Lax., HaUgloua limebdna
Ehuhb., Ebpsr PJUmzmtk, Madrep, Tab. Lxm. ;— Fungia talpa. Poly-
phylUa talpa, kc,
G«nus CydolUhda {Cydolites Lail) Polypary stony, orbiculate,
with center sub-lacunose (monostoma), above lamellose, with dicho-
tomous lamellae, beneath with plane surface, with concentric rings.
Fossil species from the oolitic and chalk formations, allied to
FungicB with which Goldfuss joins them.
Sp. Cycl, h^mitphcerica Lam., Bbonn Unodtliche PJUtmenihiere 1825, foL
Tab. V. fig. 11; Cycl. cancelUUa Lax., Faujab db Sadtt Fond, ffiet. not.
de la mont. de Saint Pierre, PI. xxxvni. fig. 8, 9, ko,
TurbmaUa {Turbtnolta Lam.) Ehrenb. Poljpary conical, with
base acuininate, cell single, terminal, lamelloso-stellate. (Is this its
place?)
Sp. Turbin. rubra QuoT and Gaix. Voyage de VAttrolahe, ZooL Tom. iv.
p, 188, PI. XIV. fig. 5—9. GuEBiK, Iconogr, Zoophyt, PI. xxni. fig. 7,
Cuv. R, Ani. id UluMr., Zooph. PI. Lxxxii. fig. 5. This species, drawn up at
New Zealand from a depth of twenty-five fathoms, &8tened to a shell, has
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POLYPI. 89
an aniiiuJ mnch resembling an Actinia, wiih a large oval oral-i^)ertare
BUTTOunded by nnmeroufl, yery large, transparent tuberoolated rays. Hie
other speciee upon which Lamabok has founded this genus are only known
in the fossil state. It was thought that they were not affixed, and oon-
sequently they were referred to this fiunily ; the discovery, howeTer, of the
voysgerB quoted above, shews that the species now Uving £u- rather belongs
to the family of the OedUna, and probably ought to be joined to if ono-
myeet Ehsbnb. It may be suspected, perhaps, from Stutohbubt's
observations, that here younger forms of Fungia have crept in.
To Thtrbinalia the genus JHploctenium GrOLDFUSS, FldbeUum Lbbbov
may be added. See Flaibdlum pavofUiMim Lxss. IUutiration$ de Zool,
PL XIV.
J^ote, The genus Lithactinia Lesson related to the Fungue, might
perhaps from recent inyestigations be established with propriety.
Comp. Lesson lUustrcUiona de Zodogie, PL vi.
C. Polyps with the whole body sofk or subcoriaceons.
Family XII. Zoanthina. Polyps affixed, never detached
spontaneously, rarely solitary, more frequently gregarious, gemmi-
parous or oviparous, never dividing spontaneously.
Zoanihus Cuv. Bodies fleshy, subcylindrical, below slender, at
the top clavate, gregarious, adhering by filiform gemmiferous stolons
of the base. Mouth terminal, crowned with tentacles filiform or
clavate.
Sp. Zwadhu ElUm, Actinia mciata Ellis, Phil. Trantad. 57, Tab. xix.
fig. I, a. Encydop. nUth. PL LXX. fig. i, GuEBiw Iconogr., Zoopk, PL XX.
fig. 4. Zoanih. Bertkdtdii Ehbskb., Po^eAoa -ScreAotow Audodih, Saviovt
JHter. de VEgypU, Pobfpet, Tab. ii. fig. 3. Zoa/nthui CouckU JomrsTOK,
BiM, Br, Zooph. p. 101. PL xxxv. fig. 9.
MamiUtJera Lesueur, CavoUnia ScHWEiGG. Bodies cylindrical
or clavate, gregarious, conjoined by a membranous base, not retrac-
tile.
Sp. MammilHfera CavoUnii, Madrepora denudata CAYOLiNn, PoUpimarini,
Tab. m. fig. 6-8, pp. 57, 58.
Palythoa Lamoub., Ehrenb., Gorticifera Lesueur. Bodies
gregarious, connate, dilated into a coriaceous expansion, with the
little apertures slightly emergent.
Sp. Palyth4)a ocdUUa, Aleyomum ocdUUum Ellis and Sol.
Hughea Lamour., Ehrenb. Polyps solitary, oviparous, with-
out any stolons.
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90 CLASS II.
8p. ffvffkM Sairignfi, Pal^tkoa Sturignyi AuDOUur, ZMmt. de VEffypU^ PUy-
pa, Tab. n. ^, i.
Family XIII* AcHnina. Polyps affixing themselves by the
part opposite to the mouth, loosening spontaneously and creeping
or swimming, solitary, oviparous or viviparous, never dividing
spontaneously, rarely gemmiparous.
Actinia L. Body conical or cylindrical, with mouth at the top
simple, surrounded by tentacles numerous, cylindrical, radiant in
one or several rows, with base discoidal.
Sea Anemonies, Comp. on these ammaLs, Basteb Naiuwrhwndige
uilapanningeThy L 1762, bL 138 — 142 ; Dicqueacase, Essay towards
the dtusidaUng of the history of the SecnmemonieSy Fhilos. Transact,
1773, p. 361, 1775, p. 207, 1777, p. 56 ; Rapp, Ueber die Folypen
im Algemeinen Whd die Actinien insbesonckre, 1829 ; A. A. Bebt-
HOLD, Zergliederung der see-anemonen tmd namentlich der Actinia
coriacea in BeUrage zwr Anatonvie, Zootomie und Physiol, Gottingen,
1831. 8vo. & 1 — 19 ; J. F. Bbandt, Prodronms JDeseriptionis Ani-
maUttm ah H. Mebteksio in orbis terroflr. circumnavigatione observa-
torum Fasc. i. Petropoli 1835, 4:to. pp. 9 — 17 Ac
The Anemonies live on Crustaoeay conchifera &a, swallow even
occasionally large mussels, reject the shell, when the fleshy part has
been extracted and consumed, by the mouth, and evert for this
purpose their body, which they do Ukewise whenever they feel
hunger. Their reproductive power is almost as great as that of
Hydra ; if they be divided transversely, new tentacles after a few
weeks are seen on the inferior portion, and each half becomes a
perfect creature ; thus they may be propagated by fission, but
propagation by spontaneous fission does not appear to occur
naturally amongst Actinice: usually it is effected by ova which get
into the stomach from the ovaries and are there developed ; when
the young ones come out of the egg they are rejected by the mouth.
That the actinice are viviparous was formerly observed by Basteb.
The young have at first fewer arms or tentacles than are after-
wards present.
These animals, with their coronet of tentacles, resemble com-
pound or double flowers; at the same time many also attract by
their lively colours. Most of them are very sensitive of the
stimulus of Hght, and the brighter the day spread their tentacles
the more. Of Actinea depressa Bapp observed that it immediately
contracted when sun-light fell upon it.
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POLYPI* 91
The cylindrical body is formed of a thick skin of which the
innennoBt layer consists of longitudinal and transverse mnsclesL
The tentacles are hollow. The stomach is a folded blind sac. The
space between the stomach and the skin is divided hy numerous
partitions ; the ovaries^ whose efferent canals open into the base of
the stomachy lie in the chambers thus formed'.
ActimcB are marine animals; they occur in the temperate and
torrid zonea Some specieB are brought to market by the Italians
and are eaten.
A. WUh UUeral pores {Cribrina Hempb. and Ehbkkb.)
8p. AetkUa rfata L., Bastkb L Tab. ziv. fig. 3, Raff, L L Tab. n. fig. 4. —
AeUmia coriaeea Cuv., AcUnia mMis L., Bastib 1. Tab. xni. fig. «. Bapt,
L L Tsb. I. fig. 3, 4, Lnsov Ilkutr, de Zcologief PL uv.
B. WUhout lateral pores {Actinia Ehbenr)
Sp. Aetmia vmdU OtUSL., Priapiu viridii Fobsk. Icon. Rar, natural. Tab.
xxvn. fig. B, b. Actinia Cerem Bapf, I 1. Tab. n. fig. 3; this species is
eaten ia the soath of Franoe» and is known by the name OrCte or Ortiffue; —
Actinia tapetum Hxmfb. and Ehbhtb. with short and numerous Uniaeida;
this species in the contracted state occasioned the establishment of a new
genus, supposed to be distinguished by the absence of tentadee : DiMcot(ma
BuBFPKL and Lxucx., Neue wirbdUm Tkiere dea rathen Meeret, Frankf. a*
Main. 1828, Tab. i. fig. i.*
Actinodendron QuoY and Gaimabd. Tentacles ramose (or
provided with vesicles lateral, fiEUM^iculate, Ehrenb.)
ThalassiaMhus BuEPP. and Leuck., Epichdia EuRENB. Ten-
tacles pectinate.
Minyaa Cuv. Actinecta Less. Body free, globose, ribbed.
Mouth snrronnded bj tentacles in many rows, which are sometimes
lobate. Disc opposite the month supplied with aeriferous canals,
serving to suspend the animal in water.
1 Besides the works of Bbrthold and Raff referred to, that of Btmib Jokbs,
General (hdUne of ike Animal Kingdom and Mamud of Oomfarative Anatomy, London,
184 1, pp. 39 — 44, also contains a detailed anatomy of Actinia.
* Comp. also the descriptions of sevenJ Mediterranean species of Actinia given by
A. F. Grubs, Actinien, Echinodemien und Wilmur dee AdrioHschen und Mittdmeere,
Kdnigsbeig, 1840, 4to. ; amongst the new species is one remarkable for its change of
colour. Act. Chamaieon Qbubb.
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92 CLASS II.
Sp. Minyas cinerea Cuv. IL Ani, i^ edit. PI. zv. fig. 8, Lesson Centurie
Zocl, PL LXli. fig. I, in the Atlantic Ooean. This genus is referred by
CuYiEB to the EMnodermcBta apoda; Lbsukub, who has made known some
other species of it, gives it a place near Actinia. An accidental, not a
natural opening in the disc, opposite the mouth, was taken by Cuyieb for
anus. See the md edition of Lamabok, HUt. not. da Anm, mhm veriibres
ni. pp. 417—429.
LiLcemarta MuELL. Body gelatinous, radiate, the rays tentap-
culiferous at the tip, above flattish, with mouth central, fiinnel-
shaped, protracted, below elongated into a pedicle disciform at
the extremity.
Sp. Lueemaria guadricomis ZoU. danic. Tab. TXXTX. Johnston, ffitt. Br.
Zooph. pp. 244—151. fig. 3—7.
Comp. on this genus Lauoubouz, M4m. du Museum, u. pp. 460 — 471.
PI. xvi. Does it belong here t Lahabok refers this genus to the AoaUpha.
Edwardsia QuATREF. Body firee, cylindrical, rounded behind.
The middle portion of the body with thicker epidermis, opaque ;
the anterior and posterior pellucid, retractile within the middle.
Mouth furnished with tentacles, hollow, arranged in single or
double row.
Sp. Edwardna BtauiemptU Quatbbtaobb, Ann. det 8e. nOt. i« Sirk. Tom.
xvni. Zod. PL I. fig. I, &».
These remarkable animals, discovered by Quatbbfaobs, live on the sea-
shore in the sand, like Sipunadiu and some Annulata. The tentacles are
not perforate at the extremity, as little as they are so in Actinia, in which
preceding authors (Raff, Rtxeb Jonbs and others) admit a reception of
water through the presumed apertures.
Order IV. Bryozoa.
Nutrient canal supplied with double aperture (mouth and anus),
replicate, the posterior portion ascending by the side of the anterior.
Tentacles long, furnished with vibratile cilia, surrounding the
mouth. The anterior part of the polyp soft, retractile within the
posterior by inversion.
Ehrentberg was the first to separate with precision these animals
from the other forms of the Polyps — see the Introduction to this
oiass. Milne Edwards makes of them, in company with the
AcephcUa nuda^ a division of the type of the MoUuaca under the
name of Mollvsc&ides. As in onr first order of Polyps we see a
resemblance to AccUephas or Medusee, in the second recognise the
proper type of the Polyps, and in the third perceive a transition to
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POLYPI. 93
the BMnodermaici, so in this last order we cannot mistake the
affinity to the Mdkuoa; this affinity is even so close that we hold
the nnion of it with the Molluscs to be almost the more natural
one.
Family XIY. Stelmatapoda nob. Tentacles disposed in a zone
around the month.
A.) Cell (the posterior harder portion of the animal) covered by
a moveable operculum. (Tentacles numerous, 16 or more.)
EBchara Lam. (Species of genus Eschara Pall., of MtUepora
L.) Polypary of aggregate cells substcmy, foliaceous, ramose.
Both surfaces of the polypary covered with opposed cells.
Onui-Ooral. Sp. Eteham foUaeea Lam. (not Pall.) Ell. CoraU. xxz. fig.
a, A, By C i—Eichara cervieomU Lax., Cuv. IL Ani. idU. iU., Zoopk, PI.
96, kG.
Gomp. on this genns Mn.NB Edwabds, Beck. amaUmiquei, phenol. H
zooL 9ur Us Biekapet, Awud, da Se, nalL 7e Sirie. VI. 1836, ZooL pp. 5 —
53, PI. I— V ; Obtervatitmi aur let p€hfpier$ fouU/u du genre Bsehairt, ibid,
pp. 3«i — 345. PI. rx — ^xn. These fossils occur partly in the chmlk-fomia-
tion, partly in the tertiaiy straU. M*COT, Jkecrip. Brit, PaUeoe. Foet. in the
OeoL Jftcf. €fthe Umv, of Cambridge. Gamb. 185 1. 4to. Pt n. pp. 44 — 47.
H. fig. 14—17.
Mdvcerita Milne-Edwakds. Fossil genus. Comp. Ann. des Sc,
not. 2e /SS^. vl Zod, pp. 345—347.
Seiepora Lah. Polypary reticulato-ramose or perforated reticu-
lately, calcareous. Cells of the Polyps situated on one side only of
the polypary.
Sp. Bdepora cdlvlota, MiOepora ceUtdoea L., Ell. CoraU. Tab. xzv. ^g. d,
D, F ; EsFXB, PJlanssenth., MiOep. Tab. I. ; Gavoliki, PcUpi marini. Tab.
m. fig. 12, 13. This polypaiy resembles a piece of fine lace, hence the
firench name dentdle de mer or manchette de Neptune (/)
Adeana Lamour., Lam. Polypary frondescent or fan-shaped,
on l)oth surfaces celluliferous, calcareous, supported by a stem sub-
articulate, not polypifeious.
Sp. Adema foUifera Lam., Schwsigoeb Beob. auf naiurh. BeU. Tab. i;
Cuv. B. AnL idU. UL Zooph. PI. 88, fig. i ;—Adeona erOmformU Lam.,
SoHWEiGaxB L L Tab. n. fig. 5, Cuv. L 1. ^. 1. In this species the stem
bean a flattened expansion, perforated like a sieve or a net, from coalescence
of the branches, and permanence of the intervening spaces.
Of this genus the Polyps, as fiu* as I know, have not been observed, but
it IB placed here from the agreement of the Polypary ; an idea may be
formed of it by supposbg an Etchaira to be placed npon a jointed stem.
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94 CLASS u.
Flustra L. Polypary of aggregate cells, membranaceous, firon-
descent, lobate or expanded into a crust, celluliferous on one or
both sides. The cells are often aculeate on the anterior margin,
their opening transverse, semicircular, or lunate.
Sea-OrutiL Sp. FlttMra foUaeea L., Eichara fdiacta Pall., Dk Jussieu
Mim, de VAoad, royale des Se, AdH^ 1741. PL ix. fig. 3 ; Ell. CoraU. PI.
XXIX. fig. a. A, B, G, £ ; Guv. R, Ani. idit. UL PI. lxxviu. fig. i ; John-
ston, ffitt. Br. Zooph, pp. 343, 343. PI. Lxn. fig. i, 1 ; — Fhutra comuta
Mnjni Edw., Guy. R Ani, idU, iU, PL Lxxvin. L L fig. i, kc.
Eucmtea Lahoub., (in part). Milnb Edw. Ann. dea Sa nai., 2e
SertBy IX. Zoolog. pag. 204, PL viii.
B.) Cell (the posterior harder part of the animal) without oper-
culum; a setose collar or a crown of set» in the anterior part of the
body in manj, or a muscular ring in others, in place of operculum.
Tendra NoRDM.
Sp. Tfniira zotterioola, CeUepora potUica EiOHW.
TuiuUj>ora. Polypary calcareous made up of crowded tubular
cells, parasitic or incrusting. Aperture of the cells orbicular. Ten-
tacles 12.
Sp. TubuUpora verrueoia MUiNB Edw., TubuUpora arbieului Lah. (syiio-
nymee excluded); Guv. R, AtU, SdU, iUfutr., Zoophyt, PL LXX. fta
Gomp. on this genus MUiNB Edwasds, Ann, des Sc. not. ie SSrie, vni.
Zoologie 1837. pp. 311—338. PL xxn— xxiv. Johnston, Hist. Br. Zooph,
pp. 165—174. PL XLVI. fig. 3, 4.
Subgen. Diastopora Lail
Cellularia Pall., Gellaria Lam. Polypary ramose, composed
of cells arranged in a single or double row or verticillate, tubular,
calcareous, with orbicular aperture.
Sp. Cdlidairia e&unMo, Crisia ebwmen Lahoub., Seri^daria ebumea L., Ell.
Oorail. XXI. fig. a, A ; Guy. R, Ani, idU. HI. Zoophyt, PL Lxxm. fig. 1 ;
Van Bbnsdbn, Nouv, Mim. de VAcad. de Brvx, xvm. PL m. ^. 11—
16, &c.
Gomp. MiLNi Edwards, Mim, eur lea Orisi«$, lea Bomhta et pluaieura
autrea peljfpea vivana <m foaaUea dont rorganiaation eat analogtie d edle dea
Tubfdtporea, Ann, dea 8c. not, ie S6rie, Zoolog, Tom. IX. 1838. pp. 193 —
138, PL VI— XVI ; Van Bxnedbn, Reckerehea awr lea Bryoaoairea, Now,
Mim. de VAcad. de Brux. Tom. xvin. 1845. pp. 11 — 19.
On the Polypary of this and some other genera shear-like organs
are seen that have some resemblance to a bird's head ; they have jomts
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POLYPI. 95
which admit of motion like the jointed feet of afHeuhta; their motions
persist even when the animal has been for some time dead. Their us? is
not known. See figures in Ellis CoraU, PL xz. fig. 2 A. (of CeUidaria
avieularia Lam. : of Fludra anguatUoba Lam.); oomp. Yam Bihkdir
Beekatkes mr lu Bryoz. L I pp. 14—95, and NomDMAirv and KxOHN
died by SiSBOLD LAfrb. der ver^^dch, Anai. i. p. 53 : also Dabwih Voyaget
cf AdvetUwre and BtagU, voL m. pp. 359—61 quoted, with figures, by
JoHKffiOK, Hkt, Br, Zoopk, pp. 319 — 331.
VaUceria Fleming, Farre.
Sp. Valheria euietUa, Sertularia eu$euta L. ; Ell. CforaU. PI. Ziv, C, C.
Fabbb, PhiL Trana. 1837. PI. zxm.
Vestcularia Thompson, Farre.
Sp. Vetieularia apinota, Seriidaria apinata L. ; Ell. CforaU. PI. Xi. fig. 17
b, B, C, D ; Fabbk, L L PI. xxn.
Serialaria Lam. Polypary homy, ramose, composed of cells
cylindrical, parallel, cohering in rows.
Sp. Serialaria lendigera, Sertul. Undigera L. ; Ell. OoraU. PI. XV. No. 34 b,
B; Catoliki, Polipi mar. PI. IX. fig. i, 1 ; the cells stand close together,
as in a Pan*8-pipe. Johkston, L 1. p. 369.
Angumaria Lam. Cells elongate, subclavate, perforated by a
lateral aperture, set on a creeping fistulous stolon, erect, distant.
Sp. AngtMmwria spalidata, Serivl. anguina L. ; Ell. Corall. PL xxii. fig.
% c, G ; LiSTXB PhiL Trans. 1834. P^- ^^» %• 4 > Johnston, Hitt. Br.
Zooph. p. 39a PL L. fig. 8, 9.
Bawerbankia Fabre.
Sp. BowerbanJna dema Fabbb, 1. 1. Tab. xxi. xxn.
Lttgrmcula Yak Beneden, LageneUa Farre.
Sp. Zaguncula rtpem Fabbb, L L Tab. xxiv.
Note. According to A. S. Hassall Bowerbcmkia densa is a
younger state of VcUkeria vmbricata ; Annals and Magaz. of noL
Hist. vn. 1841, p. 363, 364. But in Valkeria and Veaicularia
firom the observations of Farre and Yan Beneden there are
8 tentacles not spinous; in Bowerhankia and Laguncula 10 — 12
tentacles, besides vibratile cilia^ supplied with inunoveable setae or
spines. — Lusia Milne Enw. (in a note to Lahaeck Hiat not. dea
AnL a. vert. 2e ^ii. n. p. 72) is it Lagtmcula-f
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96 CLASS II.
Halodactylus Farrb. Polyparj fleshj, gelatinous, pellucid.
Polyp with tentacles 12 — 16, often longer on one side.
Sp. HalodaetyUu dtaphanut Fabbk, Alcycn, ffdaUfumm L., Ell. OairUl.
PL xzzn. fig. d, EaPBB, J^metUh, Aleyon, Tab. zmi., Fabiui Phil.
Tran», 1837. PL xxv, xxvi. ; Van BbnedxIt Beck, wwr let Bryozoairet 1. 1.
PL V. fig. I, 2 ; ocean on our coast (Dutch), adhering to marine plants
and sheik resembling a gelatinons, transparent, tabular, and irregularly
branched sea-weed.
Family XV. Lophopoda Dumortier s. Cristatellina. Tentacles
set pectinately on two arms, numerous.
Cristatella Cuv. Polypary free, disciform, polypiferous on tlie
margin.
Sp. OritUUeUa muoedo Guy., CriMai, voffona Lax., Boss. ni. SuppL Tab.
XOI ; in fresh water ; three, four, or more Polyps are seated in a freely-
swimming Polypary. Gbbyais and Tubptn have figured the egg, which is
provided with tubular spines terminating in two or more hooks ; it bursts
into two yalves, when the young animal is bom. See Ann. det Sc. not. ze
SSrie, VII. Zool, pp. 65 — 93. PL n. PL m. A. Johnston, HiM» Br, Zooph.
p. 389.
PlumcUella nob. Polypary affixed, tubular, with extremities of
tubules retractile, polypiferous.
Flume-Polyps {Polypes h pannaehe Tremr) Comp. Memovre sur
rAnatamie et la Phydologie des Polypiers composes cTeau douce
nommes Lophopodes, par B. C. Dumobtieb. Toumay 1836. 8vo.
(publifihed in part at an earlier date in the BuUetin de VAcad des
Sc de Bruxelles 1835, p. 422 <&a) Propagation occurs by eggs and
buds. Tremblet also observed spontaneous fission of the Polypary
in PlumcUella cristata,
Plvmatella Laic. Naisa Lahoub. Stem branched or lobate.
Sp. Plumatella critiakt Lam. ; Trsxb. Polyp. Tab. z. f. 8, 9 ; the body
transparent, i'" long, the plume nearly of the same length ; about rixty
tentacles ; it lives in fresh water beneath Lemna.
Pltm. eampanulata Lam. Robs. Int. T. m. Suppl. Tab. 73—75. (Lin-
NJBUS united this species with the former, under the name of Ttibularia
campanulaia.) Probably Plumatella repent Lam., Sohasffes Armpo-
lypen 1754 (and edit. 1763). Tab. i. fig. i, a, Eiohhobn Watte/iihiere,
1 78 1. Tab. IT. p. 43 ((2er Polyp mU dem Federhutch), is only a yariety
of this. According to Norbmann, the tube continues to grow for some
time after the death of the Polyp. When full-grown it has up to sixty ten*
tacles, but in young animals they are less numerous, shorter and tixicker.
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POLYPI. 97
FrederidUa Gebvais.
Sp. PlumaieUa tuliana, Tnbularia tultana BLUnvB. ffandb. d, fMrf. Hid,
PL I. fig. 9.
Alcyondta Lail Polyparj incrusting, irregular, multiform, com-
posed of tubules aggr^ate, cylindrical
Sp. PlumateUa fungoM nob., AUyoneUa tiagnorum Lam., Encyd. mMkod,
Vers, PL 472. fig. 3, a, b, c, d, Rabpail HiU, naiurelU de VAlcffomdief/vk-
viaHle, M^moirea de la Soe. dPHiat. not. de Paris, it. i8a8. pp. 75 — 130,
pL 12 — 16. To this species probably Bakxb'b figure belongs, Employ-
ment for the oUeroteope, PL xu. fig. 13 — 22 {Bdl-Jlower animal). See
Jossssov, Hi$t. Br. Zooph. pp. 391 — 395.
Rabpail is of opinion that this animal is merely a form of PlamateUa
eampanulata altered by age, from which he does not distinguish PUimat,
erittata ; as soon as the animal breaks the egg, it moves freely about, and
should then be described as CrisUUeUa, so that all of them belong to one and
the same species at different periods of Hfe. But amongst other objections
to the union with PlunuUdla eritUOa and eampanuUUa is the number of
the tentacles which in Alcycnella is, according to R^lbpail, only forty-four
(according to Ehbsnbxbo forty-two), whilst here, if it were an older state
of PlumaieUa campanulata, the number might rather be expected to be
greater. Also in CridateUa there are more tentacles. However, it is
possible that OriaiaieUa may be a younger fonn of another spedee of
PlumateUa, and so at some future time to be excluded from the list of
genera.
Pallab, as it seems, first discovered the Alcyondla in our country in the
well-known lake of Rockanje, in the island Voom, and described it by the
name of Ihibularia fungoea, Nov. Oommentar. Acad. Seient. Pelropol. xii.
1768. The name AlcyoneUa should be rejected, for it was borrowed from
a supposed resemblance with Alq/onium, which on further investigation
was found to be totally unfounded.
Van Bkkeden has observed the sexes to be distinct in AlcgondUt, male
and female individuals occurring in the same Polypary. Bullet, de VAcad.
dee 8c. de Brux. Tom. vi. 1841. p. 2^6.
FalttdiceUa Gebvais.
Sp. AlcyoneUa aMnUata Ehbkitb. Symbol, f Comp. Yak Bkvidbn BuUet. ffe
VAcad. deaScde BnuceUen vi. L 1.
VOL. I.
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CLASS III.
SEA-NETTLES {AGALEPHJS)\
Bt Sea-nettles are understood radiate animals of a gelatinous,
transparent substance, which swim freely, and of which the organisa-
tion in parts that axe frequently repeated usually indicates the number
four or its double. They have no intestinal canal, but ordinarily a
blind stomach from which numerous tubes, mostly divided into
branches, run like rays to the circumference of the body (aquiferous
vessels, respiratory organs?). Li some the sexes axe separate:
others appear to be bi-sexual. Traces of a nervous system are
present at least in some; yet organs of sense, except appendages
which serve for feeling, are absent in many, whilst some have in
addition parts which by certain writers are considered to be organs
of hearing, by others organs of sight.
Of the name Seornettlea, AcalephcB^ UrticcB marinoe^ we have
treated above. With Lamarck they form an order of the class of
radiate animals {Badiaires), under which this writer also groups
1 On this olaas may be compared especially :
F. EaoHBOHOLTZ, <^ttem derAcalephm, Mit i6 Kupfertafeln. Berlin, 1839, 4to.
C. O. Ehbutbibo, JXe Acalephen des rothm Mwru unci der Organismtu der
Jlfeduaen der Ostsee, Mit 8 colorirte Kupfertafeln. (Reprinted firom the Ph^fiihal.
Abhandi. der Kdnigl. Ahad, der WieeenchafUnzu Berlin aiue dem Jahre 1835).
Hittoire natureUe dea Zoophytes. AcaUphee par B. P. Lbbson, ay. pL Paris,
1843, 8vo-
J. G. Will, fforce TergetHma^ oder Beschreibung und Anatomie der m fferbste 1843
hei Trieet heobachieten Ahalephen, Mit 1 Tafeln, Leipzig, 1844.
£dw. Forbbs, a Monograph of the BriHeh Naked-eyed MedaetB^ miikfiguree of all
the epeciee. London, printed for the Kay Society, 1848, fol.
HnxLBT, On the afiiUHee of the MedMos, Phil. Traneact. London, 1849, Pt. il.
p. 413 Ac.
On the Siphonophora :
A. KoELLiKBB, Die Sehmmmpdypen, oder Sipkonopkoren von Meuina, Mit 12
Tafeln, Leipzig, 1853, ^1-
B. LiuoKABT, Die Sipkonopkoren, eme Zooloyiecke Untereuehung, Giessen, 1853, 4to.
B. Lbuokabt, Zur ndhem KerwUnite der Sipkonopkoren von Nima, Mit 3 Kupfertaf.
Berlin, 1854, ^^o*
0. YoaT, Sur lee Sipkonopkoree de la Mir de Nice. Gen^e, 1854.
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SEA-NETTLES. 99
those with a spiny skin {E(^tnoderm(Ua) as a second order ^ They
were first described as a distinct class of animals bj CuviER in the
first edition of the B^gne Animal^ under the name of AcalephcBK
The name of gelatinous animals is more appropriate to this class
than to some others of the Radiates of Cuyies. The kinds most
generally known bear the name of Zeekwallen in Holland, [Quallen
in Grermany, Sqnalders* or sea-jelly or sea-blubber in England.]
When thrown by the sea upon the strand, they lie motionless
during the ebb ; for they do not creep, but can only move in the
water by contraction and expansion.
The pungent and burning pain like that caused by stinging
nettles, which many species inflict on being touched, was generally
considered in former times to be the effect of a mucus secreted by
the skin of these creatures. It is only since 1841, fix)m Wagner's
microscopic investigations, that minute threads situated on the
surface of the skin have been recognised as connected with this
phsenomenon, since in Acalephes which cause no such pain (as in
Camopea) they were not discovered. Each of these threads lies
rolled up in a little oval vesicle or cell, firom which, on pressure or
irritation of the skin, it is forced out by eversion ; they are readily
detached with the vesicle to which they are fixed by a tubercle,
and are always present in the secreted mucus that produces a bum-
ing pain. Yet the cause of this ought not to be considered as
entirely mechanical ; it is probable that some acrid fluid, secreted
by the cells, adheres to the threads ^ Nevertheless an accomplished
^ RadiaMr» moUatse$, Lamabok SjfU. denmim. «. verUhres, 1801, pp. 341, 359, and
in his later works, Extrcut du C<nirs de ZodUgU sur let anim. 8. v. 18 12, and Hiit. Nat.
daAnim, s. veri. 1815.
* CuviEB in his TaUeau iUmentaire (179S) and in the tables at the end of the first
part of his Lefon» tFAnat, eomp, had comprised all the animals, which he afterwards
named radiated animals, in a single cIms, under the name of Zoophytes. Of this the
Ortieg de Mer make the second order, which agrees with his later class of the Aealepho!,
Here also he placed the AcUnicB, which however in the second edition of his lUgne
Anaaal he separated from it again, in order to unite them, as had already been done
by others, with the Polyps.
* [Sm T. Bbowke's Works edited by WnKiNS, Vol. iv. p. 333, quoted by F0BBE8
Brit. Staff, p. 87.]
* B. Waonxb Hber mvOkmoMcke NtMehtymne der Medu$en, Wibohann u. Ebich-
SOH Arekiv. f. Naturg. 1841, I. s. 38-;-43; ^efter den JBau der Pelagia nodUvca.
Lexpdg, 1841, fclL Jam. Zoot. Tab. xxxin. — Subsequepily theoe parts were also inves-
tigated by Ehbenbebo, Philuti, Will, Milnb Edwabd^ kc
7—2
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100 CLASS III.
observer, Dr F. Will, found these tlireads in Emharia (amongst
the Beroecea) without perceiving any irritation on toucliing it
In many there is only a single oral aperture, placed in the center
on the inferior surface of the body. In others many suctorial tentacles
are seen, or the arms have apertures conducting to tubes, which,
like vessels, fall into larger stems and finally open into a common
cavity, the stomach (Rhizoatoma Cuv.). From the stomach arise
water-canals, which are provided internally with cilia. By some
writers these have been regarded as blood-vessels : but far rather
ought they to be considered as respiratory organs, since in part they
open freely on rtie surface of the body. But, in addition, blood-
vessels have been found, which, at least in Beroe^y lie round about
the water-canals, surrounding them like a sheath. Here nucleated
corpuscles have been observed (blood-corpuscles?) which however
move only very slowly and irregularly.
The sexual organs are distinct in the disciform Acalephes, but
have in both sexes the same form. In jEquorea they lie in form of
folded plates on each side of the water-canals which spring from the
stomach, towards the inferior surface of the disc. In the eared
Medusa (Aurelia or Medusa aurita) there are four cavities, opening
below at the disc and which have been taken for respiratory cavi-
ties, in which lies a folded organ, that is, an ovary or a testis,
according as it contains ova or spermatozoa ; in most Acalephes the
spermatozoa have the ordinary cercarial form. In other Acalephes,
as Beroiy ovaries and testes are united in the same individual:
here they lie along what are called the ribs, beneath the skin.
The metamorphoses, of which we have already spoken above,
are remarkable in young Meduece, The eggs, that pass from the
ovaries along the canal of the arms to their folds, are collected here
and carried about, for a time, by the mother in saccules which
afterwards disappear*. The young animals quit these receptacles
in the form of ciliated Infrisories resembling Leucophrys or Bursaria.
These swim freely about, but after a short time (two or three days
according to Siebold) become fixed by their thicker anterior ex-
tremity which has a sucker. Next, the body becomes cylindrical,
transparent, and at the free end, which thickens, an oral aperture is
1 [The blood veaselB described by Will could not be seen by Fobbss, Hdzlit,
Leuckabt, &C.]
« See the figures of Ehukbebo Die AkaUphen &c. Tab. m. fig. i . i. Tab. vm. fig. i .
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SEA-\ETTLES. 101
seen, round which at first two, then other two projections appear
tliat become lengthened out into arms. Soon there grow out four
other similar arms, and the animal has then the form of an eight-
armed Polyp. These arms can shorten and lengthen themselves
greatly. In this state the animal continues several months, and
can multiply itself by buds and offsets. Subsequently there come
into view transverse indentations, by which the animal is divided
into several rings resting u|)on one another, round each of which
eight rays arise. These rings are like young medusae piled on one
another : they become separated from each other, from before back-
ward ; and then swim freely away as young medusae ^ How long a
period is requisite for their full growth is unknown. The genital
organs could not be distinguished imtil they were more than an
inch in size. Hence it is clearly an error when some ascribe to
these creatures in general a very short duration of life and a rapid
growth — or even name them annual animals. Their power of re-
production seems to be very small. Some species are able to bear
the loss of parts, but these when removed do not continue to grow.
In the genus Ceatum^ however, Mertens was led to believe, from
some observations which he was unable to complete, that propaga-
tion by spontaneous division may be admitted. [And this has been
lately confirmed by Koelliker*, in Stomobrachtum mtrabile, which
he believes to be a larval form oiMesonema C€erule8cen8, a new species
of Medusa discovered by him at Messina. The fission begins in the
stomach — many individuals having been found with their round
disc somewhat elongated and with two stomachs more or less com-
pletely separate by constriction, but still lying side by side. Next,
a meridional groove is seen on the outside of the disc between the
stomachs: it deepens gradually until the acaleph is separated into
two distinct individuals. The whole process may be completed in
eight to twelve hours. The multiplication however is not yet
^ Sabs BeskriveUer og Jagttagdaer &c. (extract from it in Wixomann's ArcUv,
1836, 8. 197 &c.) ; and in Wiegm. Archiv. 1837, B. ii. s. 406 ; C. T. VoN SiEBOLD
Bekroffe tur NcAurgeach, der fpvrbdtosm Tkiere, Dantzig, 1839, 4to. 8. 26—35. Sabs
in WixoMAHir «nd Ebioh80M*b Archiv. 1841, 8. 9 — 34. MSvuoire 9wr U d^vdoppement
de la Mediua aurita etdela Oyanaa eapiUata, Ann. dea »e. not. 8ec s^rie xvi. Zoclogie,
pp. 311 — 348, PI. 15 A— 17. 1 84 1. J. Steenstrup Om Foriplanining og UdvUding
gjainem vexlende GeneraiumtraJl^ker, Kjbbenhayn, 1841, 4to.
' SiEBOLD and Koellikeb's Zeittckriftf. witsefitehcfi. Zod. iv. pp. 335, 327.
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102 CLASS III.
finished: the halves again divide in a direction perpendicular to
the former line of division : but here the fission does not always
begin in the stomach. How often the process may be repeated is
not known. No trace of fission was observed in Mesonema^ though
it was often seen to occur in the larval forms, even when ova were
already distinctly visible on the vessels.]
[The SipkancphorcB have been shewn, by the investigations of
Huxley, Leuckart, Koelliker, Gegenbaueb and Vogt, to be
compound animals, or colonies, connecting the hydroid polyps with
the acalephs. They are named by Koelliker, in consequence.
Swimming Polyps {polypi nechalei). They consist in general of a
stem, usually cylindrical and long {Diphyes)^ sometimes shortened
and sacciform (Physalia), sometimes disciform {Veldla), to which
appendages are attached which diflfer remarkably in form and func-
tion. Some of these are suctorial tubes or stomachs, others motive
organs, others feelers and prehensile organs, others again protective
laminae (bracts), and sexual capsules. Great differences prevail with
respect to the number, arrangement, and development of these parts,
in the different families : those which are constant in all siphono-
phors are the stomachs, the prehensile apparatus, and the sexual
capsules. The stem is muscular, and hollow — ^the interior forming a
canal in which the nutrient fluid moves with rapidity. The swim-
ming apparatus is either passive or active — ^when passive it is a
hydrostatic apparatus consisting of a bladder filled with air which is
always placed at the upper extremity of the common stem : when
active it consists of swimming-bells, which are also placed at the
upper extremity of the stem, and are variously grouped, and in
variable number in different genera : the swimming-bells may exist
conjointly with the air-sac or without it. These bells are, in
general, formed on the plan of a Medusa, consisting of an elastic
bell-shaped mantle, very various in form, with an internal mus-
cular layer which surrounds the swimming-sac. On the outer sur-
face of the latter, there is a system of four radiating vessels, which,
at the circumference of the aperture, fall into a circular vessel, and
at the summit of the bell arise fi-om a single vessel, which passes
through the pedicle of the bell and falls into the cavity of the com-
mon stem. All the other appendages of the stem have also a more
or less perfect system of vessels, which communicate with the inter-
nal cavity of the stem in a similar way. The only communications
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8EA-NETTLE8. 103
from without with the cavity of the stem are by the mouths of
the digestive tubes, which answer to the bodies of polyps. The
food digested by these stomachal polyps is conveyed from their
extremity into the cavity of the stem, fix)m whence it is carried
through the vessels of all the appendages, partly by the contractility
of the walls of the stem, partly by the action of the cilia which
line the vessels of the appendages.
The polyps, or suctorial tubes, or stomachs, have no tentacles
round the mouth. They consist of three portions; the external,
very variable in form, the proboscis and mouth : the middle swollen
portion, the digestive stomach, with dark streaks containing bile-
cells : the terminal rounded portion, with thick cellular walls. At
the base of the stomach, or sometimes immediately on the common
stem, is the prehensile apparatus for the capture of prey. This
usually consists, for each polyp, of a single long and thin thread
with lateral subdivisions, which do not branch ; more rarely of sim-
ple threads or shorter cylinders. This apparatus is always supplied
with midtitudes of thread-cells, which in the case of lateral acces-
sories are grouped in very regular and constant forms, and are con-
spicuouB from their bright yellow colour. The sexual appendages
have large swimming bells of the general medusan form. They
consist of a bell-shaped mantle and vessels — and a nucleus, more or
less conspicuous, which contains in its substance the sexual ele-
ments, and is dependent from the vertex like the clapper of a bell.
In some cases the medusan form of the mantle is in great measure
suppressed, whilst in others it is quite complete, and here the sexual
appendage is detached at an early period, as in certain hydroid
polyps, and the sexual elements are developed afterwards; where
the medusan form is not thus perfect, the contents of the sexual
capsules, when detached, are found to be mature. The Diphyidm
are, according to Leuckart, all uni-sexual, but the observations
of Gegenbauer [Zeitachr. f. toissensch. Zool, V. p. 313) shew that
some at least have the organs of different sexes on different groups
of the same stem: the PhyBophoridce are all bi-sexual, in some
{Stephmomia) the organs of the two sexes being on different pedi-
cles, in others (Fhysalia) on the same pedicle.
The organs of less general occurrence are the Bracts, Lamin»
or Covers, and the Feelers. The Bracts or Covers, more solid
than the other organs, are for their protection : they contribute little
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104 CLASS in-
to the motion of the colony, but sway gently to and fro with the
contractions of the stem. The Feelers are cylindrical or vermiform
structures having much resemblance to the polyps, but without an
external opening. In some cases they are in constant motion,
feeling about in all directions: in others they are more sluggish
and are loaded with the general nutrient fluid. The latter fact
would seem to suggest a respiratory function. Some writers have
considered them to be receptacles for the fluid forced from the
interior of the prehensile apparatus during its violent contractions :
and hence the name, sometimes given to them, of fluid-holders.
But it has been objected to this explanation, that the two sets of
organs are often at great distances from each other, and that their
alternate action has not been established.
The composition of all the organs in the Siphonophors cor-
responds to that of the other acalephs. As in these, the specific
gravity of the mass diflfers little from that of sea-water. The shell
found in some families, {veleUa, porptta) is the thickened and hard-
ened wall of the air-sac*.]
On the Nervous System of Acalephs, the observations are
hitherto imperfect. In the Medusas Ehrenbebg observed, at the
base of each of the eight marginal corpuscles which he takes to be
eyes, a part which he considers to be a ganglion. Each of these
ganglia is double, or consists of two limbs that diverge towards the
marginal corpuscle. Besides these a row of ganglia lies near the
tentacles at the margin of the disc ; every ganglion divides into two
twigs, each of them for one of two adjoining tentacles: so that
every tentacle receives two nervous twigs coming from diflerent
ganglia. This ring of ganglia round the margin is interrupted by
the larger double ganglia of the marginal corpuscles. Moreover,
Ehrenbehg saw four groups of ganglia lying in the cavities for
the four genital organs and in connexion with the tentacles of these
cavities. Ehbenberg could not detect a nervous ring round the
mouth, the usual form of the nervous system in Radials^. Some
writers are of opinion that it by no means follows from these obser-
vations that the parts so described are really nerves. Grant de-
scribed in Cydippe pileus a nervous ring with eight ganglia, each
* CJomp. ItEVCKAUT Zooloffiacke UrUersuchungen, 8. 3 — 41 ; Koellikkr in ZeU. f.
Wissenaeh. Zool. iv. 8. 306 — 315.
' Ehbbnbsbo DU Akalephen &e. 8. 25, 36.
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SEA-NETTLES. 105
giving off three nerves, of which the largest mn lengthwise between
two bands of cilia* close to the external surface of the body; but a
later observer, F. Will, was not so fortunate as to find this system
of nerves. On the contrary he detected a conical ganglion above
the fdnnel-shaped structure from whence the water-canals arise,
consequently opposite to the mouth. From this ganglion many
fine threads arise, to be distributed to the substance of the body
and its different parts. In the same situation Milne Edwakds also
saw a ganglion in the genus Lesueuria, from which numerous
nerves, collected into four bundles, arose*.
Ehrenbebg considers the eight prominent organs, situated in
Medusw at the edge of the disc, to be eyes : they contain a sandy
or stony concrement, a quantity of minute hexagonal prismatic
ciystals composed of carbonate of lime. Such calcareous concre-
ments are often found in the animal organism in the neighbourhood
of nerves ; as for instance in Frogs by the sides of the vertebral
column near the exit of the spinal nerves. A red pigment which
moreover generally distinguishes these marginal corpuscles, (it is
sometimes wanting,) caused Eurenberg to conjecture that they
were eyes*. There is more probability in Koeluker's idea, that
they are to be considered as auditory organs*. Even in vertebrates
little stones or grit of carbonate of lime are found in the auditory
sac or vestibule of the membranous Labyrinth. In Beroe and the
allied genera only a single organ of the sort is found, a pedunculated
vesicle with calcareous crystals at that end of the body which is
opposite to the mouth*.
The apparatus for motion consists of transparent muscular
fibres, having sometimes a longitudinal, sometimes a circular
course : they exhibit the same microscopic transverse stripes which
are characteristic of the voluntary muscles of higher creatures*.
Several of the Animals belonging to this Class are phospho-
rescent. According to Ehrenberq the Arabians on the Red Sea
1 Traru. of the Zooloff. Soc. London, i. 1833, P- 'O.
* Attn. desK. not. le S^rie, Tom. xvi. Zoolog. p. 206.
* Die Aiaiepken, b. 14.
^ Fbobiep'8 Neue Nctken, xxv. Bd. (Januar. 1843) s. 81 — 84.
< MiLXS Edwabds, 1. ]., Will, Sone Tergest. s. 45, 46.
* Wagnkb Ban der Pdagia nodiluea; his Tab. Zoot. zxxiil. fig. 30 ; Will, 1. 1.
■• 4^—49-
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106 CLASS III.
name the entire family of Medusae Seor Candles {Kandil el BaJir^).
Bosc, and other writers after him, went too far when they main-
tained that all Med%isw, nay all Acalephes (Eschscholtz) are
phosphorescent. Still, not .Medusae alone, but other Acalephes also
do possess this property: the phaenomenon has been observed in
species of Berde ( Cydippe pileus^ Eucharis multicomis, '&c.) : Ste-
phanomia also diffuses a lively light by night. This phosphorescence
is a vital phaenomenon, and ceases on the death of the animals :
though some of them, like other organic substances, are luminous
even after death ; but that light is of a different nature from phos-
phorescence during life. Thus Will, for instance, saw Beroe rw-
fescens emit a light after death, which differed by its bluish-green
colour from the yellowish-red irradiations of the living animal.
Dead Acahphoe^ or mucus arising from decomposition of animal sub-
stances, can contribute little or nothing to the gorgeous spectacle
of the illumination of the sea, of which so many voyagers have
given striking descriptions : the chief cause of the brilliant sparks
resides in minute marine animals, especially Meduace^ like the
species which SuRiRAY named Noctiluca miliartSy and which, being
not larger than a pin's head, looks like a globule of mucus to the
naked eye^.
Acalephes are met with in all seas. A very large number of
species occur in the Mediterranean belonging to the most different
forms. In the seas of the cold and temperate zones scarcely any
Siphonophotw are found, at least not in the northern hemisphere ;
however the currents may occasionally bring with them southern
forms from a distance, as is proved by the fact that Owen, on the
south-west coast of England, observed Velella and Porpita, and
Hyndman, on the coast of Ireland, Dtphyes^. Some species are
widely diffused, asAurelia auriia, and Gyancea capiUata: the first was
^ Ehbehbebo das Leuchten dea Meerea, a. 146. Comp. espedally on Uus subject
Uie work already quoted p. 53, so instructiye as well from the author's own observa-
tions as from the extensive use he has made of earlier works.
* [Van Bbniden refers Noctiluca mUiaria not to the Acalephes but rather to
the Bhizopoda; see note by Dr Sohleosl in the german translation of this work,
p. 106.]
' Owen Lecturei on the comp, Anat, of the invert^. Animals, tS4^, p. 101 ; Htnd-
KAN Note on the occurrence of ike Oenus Diphya on the coast of Inland, Ann, qf Nat,
Hist, vn. 184 1, p. 164.
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SEA-NETTLE3. 107
found hj Ehrenberg in the Red Sea, and he could not discover
any difference between it and that found in the northern and the
Baltic seas. Oftentimes manj species of Medusae are collected in
such quantities at certain places, that they form as it were banks
in the sea, which it requires days to sail through \ In fine calm
weather Mednaas come to the surface : during storms they seek the
quieter waters of the depths.
1 PiBOir 0( Lbsusub AnnaXu Sm. Mum, xiv. p. 221 ; rooh a bank of M0du9a aurita
m the Baltic at the month of the Weichael was also described by V. Sibbold, Beitr. z,
Naturge$eh. der wirbeUoten Thiere, s. 5.
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SYSTEMATIC
ARRANGEMENT OF ACALEPHiE.
CLASS III.
ACALEPH^
Gelatinous animals, swimming freely. Stomach included in
the parenchyme of the body, without an abdominal cavity ; canals
arising from the stomach, filled with water. Ovaries and testes in
one and the same individual or the sexes distinct without organs
of copulation. Vestiges of a nervous system not always distinct.
Arrangement of parts usually quatemarian.
Order I. Siphonophorw.
[Swimming Polyps without tentacles round the mouth, attached
to a common stem of variable length, and moving freely by means
of special swimming apparatus, with prehensile filaments, feelers,
and protective covers or bracts, or some only of these organs,
attached mediately or immediately to the same common stem.]
This first order includes the Acdl^phea hydrostaUqties of Cuvier
and a part of his AcaUphes sirnplea.
Family I. VelelltdcB or Chondrophorce. Common body, sup-
ported by a cartilaginous* lamina, which is cellular internally.
The part of the body which faces upward is supported by a
disc, which in Porpita is even in some degree calcareous, and con-
\} The disc contains horay substance, not cartilage, according to Leuokart.]
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ACALEPHiE. 109
tarns cells which are full of air. Above, this disc is covered by the
integument alone; below, it sustains all the parts of which the
[compound] animal consists.
[The shell of Vddla with its horizontal and perpendicular plates
consists of a single piece. The thicker horizontal portion is formed
of two laminse connected by perpendicular concentric pieces, so that
annular canals are formed which are filled with air. These canals
communicate with each other in Velella, but not in Porpita : in both
they open externally by many minute pores on the upper surfieu^e.
The soil parts constitute a mantle which covers the sheU and projects
beyond its edge by a free border. At the inferior excavated portion
of the shell, the mantle has on its outside the attached polyps and
appendages^ on the inside the large liver.
The polyps are of two sorts, a single large and central polyp, and
many small ones disposed around it in irregular rows. They have
been designated " stomachs " and " suctorial tubes." But observers
do not agree respecting their function. Lessok attributes to both sets
a digestive power. Voyage de la CoquiUe, pp. 49 — 56^ and Acalip^iea
p. 561 : whilst Y. Siebold Vergl. AruU. a 63^ note, thinks that the
smaller polyps alone discharge the office of digestion, and consigns
the large one to the respiratory Gfystem : and Hollaed Ann. des Sc.
NcU, T. iiL 1845/ p. 250, says that the large central pouch is the
stomach and the small ones canaiix aquiftres, Koelukeb however
assures us "that he has found small Crustacea both in the large and
the small tubes, and has seen the residue of digestion pass from
them all indifferently. Consequently we conclude with him and
others, that the VdeUidcB are colonies or compound animals.
The liver m a large brownish mass placed above the central
stomach : it fills the inferior cavity of the horizontal plate. It is a
collection of fine canals formed of homogeneous membrane lined
with brown cells. A certain number of the canals branch firom two
openings in the base of the central polyp : they frequently anastomose
and form a network on the surface of the liver frx>m which fine
vessels pass to the perpendicular plate and to the margin of the
horizontal plate {Velella). These vessels, then^ would seem to have
received the nutriment which has passed from the central stomach
into the liver^canals, for the purpose of redistribution to the soft
parts when it has been modified by the biliary secretion. Of the
smaller polype a few, which hang beneath that part of the liver which
projects beyond the large polyp, open into liver-canals: but the
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110 CLASS III.
greater part of them have no oonnezion with these canals, or with
the central polyp, but lateral branches of the Toeeb open into their
pedicles, so that they at once give the product of digestioa to the
vascular system. In Porjnta the lesser polyps open into liyer^janala
and not into vessels.
The generative organs are seated, as clusters of minute bodies, on
the pedi<des of the smaller polyps. They become transparent and
pyramidal, and having gradually assumed the mediisan form are
detached. They were first noticed by DEiiLE Chiaje, Descriz, iv.
p. 107, Tav. 146, fig. 10, 12. The sexual germs are formed on the
wall of the radiating vessels. Huxley, Gegenbauer, L L
Hie prehensive organs are placed around the lesser polyps on the
horizontal margin of the mantle. They are hollow and open into a
vessel like the lesser polyps. They have no special nettle-nodes, but
numerous scattered thread-cells.
The air-canals were discovered by Ksohn ; they are minute vessels
which pass firom the innermost air-spaces of the horizontal cartilage,
perforate the mass of the liver, and reach the walls of the polyps
where they appear to terminate by closed extremities. They are
most numerous in Porpvta. See Kokt.t.tkkr Die Siphonoph. pp. 46 —
64.]
Velella Lam. A semi-orbicular crest, compressed, containing a
cartilage within, placed obliquely above the disc. Marginal ten-
tacles simple.
Sp. Vddla »piran8, Meduia vdeUa L., Holoth. tpirana Fobsk. Icon. Ber,
nalwr. Tab. xxvi. fig. k, ArmeiMtarium veUUa Costa Ann, da 8c. not,
sec. adrie, Tom. xvl PI. 13, fig. 3, (figure of the vessels from the stomachs
on the inferior surface of the cartilaginous disc), in the Mediterranean.
Aooording to FobskXl the French sailors call the animal VaUeUe: they
eat it fried with flour and batter. The name VeUUa appears to be derived
from vdvm and from the crest^ which like a full-spread sail, adorns the
upper sur£EU». The beautiful blue colour of the animal is impaited to the
water in which it is examined, but disappears in spirit of wine. During
life the creakure is not unattractive {**non tnvemuta eat quantum vemU
Ueet," Fobsk. Deacr. Animal, p. 105) ; see the coloured figure of Lbsueub
in TiBOV, Voyage attx terret auatr, PI. xxx. fig. 6. (This species is from the
Tropical Seas, VeUUa ecapkidia PiboN). For the other species, not easily
to be distinguished, of this genus, consult chiefly Esohboholtz Syat. der
AcaUphen, s. 168 — 175.
Subgen. JRcUaria Ebchsch. Crest membranous, placed longitudi-
nally on the disa
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ACALEPHiE. Ill
NcU. If the figures in FobskJU, Tab. xxvl fig. k3, k4, k5, belong to
a }f(mng VeleUa, aa appears from the explanation of the plate, this genus
must be suppressed ; which is Blaikvillk*b opinion.
Porpita Lam. The Lamina cartilaginous (?), circular, marked
with concentric strias decussated radially. Marginal tentacles
appendiculate.
Sp. PorpUa medUerranea EaOHSOH., Porp. Fonkalii, Dx Bjlav, ffol, denu-
data FoBBL Icon, Bar. waL Tab. xxvi. fig. L., in the Mediterranean ; —
Porp, umbeUa Esghsgob., Porp. giffomUa PiBON, Voy. aux ferret auttr,
PL zxxi. fig. 6, in the Tropical Seas ; Porp. ckryeocoma Liaa., GUJE&IN
Iconogr.y Zoophytet, PL xviu. f. 2. — {Medusa PorpUa L. is merely the
cartilaginous disc of some species of this genus.)
Family II. PhyasophoridoB {Hydrostatica Cuv.) Body sus-
pended in the water by means of a swim-bladder or of receptacles
filled with air.
Btadder-bearers. The opinion that these animals are able to
expel the air from the air-hladder at will was rendered doubtful, as
a general rule, by Olfebs, who could find no opening in the large
bladder of Physalia. [Subsequent observations however have deter-
mined that Physalia is the only one of the Physsophoridce whose
bladder does really communicate with the external air. But^ though
there be no such communication in the rest, Leuckart states that
in many of them (and he believes it to be true of all) the air may
be readily caused to pass from the cavity of the bladder into that of
the common stem, by the expansion of the upper extremity of which
the air-'bladder is in all cases surrounded.
a) with short stem or axis without swimming bells.
Physalia Lam. Swimming bladder very large, crested above,
with an aperture at one extremity : the whole of the common stem
expanded so as to form a receptacle for it: from the inferior surface
of the eitpanded stem the polyps are suspended together with
feelers and prehensile organs, of dififerent thickness and of great
length.]
SeaMadder. The colony swims constantly on the sur&oe of the
sea^ and for that purpose makes use of the crest on the top of the
bladder as a saiL Hence its name, het bezaantje, the Partuffuese
man of loor, la petite galh'e, &c If in the nomenclature we ought
strictly to hold to priority, then this genus ought to be named
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112 CLASS III.
Salacia, for thus Lu7N.£UB annoonoed it in the earlier editions of his
Syst N(U, ; in the tenth and following editions it is no longer met
^ with, and Linn.£US afterwards arranged the species known to him
under the genus HohthwrioL
Yon Olfebs especially threw much light upon the organisation of
this genus by the investigation of Phyacdia caravella Eschsch. {Phys,
wreikusa Tiles.) A PhyaaUa has two bladders, the internal is
filled with air, and was described by Olfebs as perfectly closed; the
external has an aperture situated at one extremity and surrounded
by a sphincter.
[QuATREFAGES has desciibed the action of this sphincter muscle,
and the connexion of both bladders with the aperture; he also
caused the air contained in the interior bladder to be analysed, and
found that it contained less of oxygen than atmospheric air by about
3 per cent. : the animal appeared to be able to expel the air volunta-
rily at intervals, and to distend the bladder again after a short time:
it would therefore seem to be a respiratory organ for the colony : the
air-bladder is surrounded on all sides by the external bladder or
envelope, which is in fact the expanded stem of the colony : with the
under surface of this the various appendages are connected, and into
its cavity the cavities of them all open more or less directly : the
bladder in Phyaalia did not appear to Quatrefages^ to be merely a
passive organ, for besides the power of emptying and distending it
the animal seemed to be able to direct the fluid contained in the
cavity of the appendages into this or that bundle of them at will,
and so to alter the position of the center of gravity of the bladder,
and by thus bringing different regions of it to the surfiice to steer
its course.]
The larger and smaller tentacles are capable of extension and
contraction, and serve probably for feeling and seizing. Small
clumps of red corpuscles, which are situated between the larger
tentacles, are, according to Olfebs, eggs : but the sexual organs of
the Pkysaophoridce require further investigation.
See V. Olfbbs in Physikal. Ahhandl, der KSnigl. Akademie der Wisaemck,
zu BerUna, d, Jahre 1831, Berlin 1832, 8. 155 — aoo.
Comp. also on iliis genus J. G. Van Hasselt in Algem. Kund. en LeUer-
bode i8«8, No. 44, 45 ; F. W. Etsbkhardt, Nov. Act. Acad. Oca. Leap.
^ Ann. det Sc. wiL %e S^rie, Tom. Ii. p. 115.
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ACALEPHJE. 113
Ca/roL Tom. x. a. 410 — 416, Tab. xxxv. fig. 49 ; Esohboholtz in o. t.
KoTZKBUB*s Entdeckwng9^ceiae m. 1821, a. 193, and Syti. der Acalephen, s.
157—164. LsucKART in Zeiitchr. fOr Wieuentch, Zooloffie in. 1 89 — 2 1 3.
AthoryUa EsCHSCH. {Rhodophyaa Blatnv.) [The motor organ
of the colony a coronet of solid bracts, or covers, fixed to the stem
immediately beneath the air-bladder. Polyps, feelers and prehen-
sile filaments attached to the very short remainder of the stem.
Sp. AthoT^fia rotaeea Esohbch. Koellikxb Die Siphonoph. Tab. vn. The
Polyps are not nearly as numerous as the bracts. In large colonies Kokl-
LIKXB could not count more than eight of them whilst the bracts numbered
10 — 40. They are seated in t^e space covered by the bracts, and with their
points project somewhat beyond them when the coronet opens, but lie entirely
concealed when it closes. The feelers are more numerous than the polyps
(11 — ?o), long and nearly filiform, and play between the bracts when they
open. The lateral subdivisions of the prehennle organs terminate by two
filaments.
b) with short axis or stem and swimming bells.
Physsophora FoBSK. Several swimming bells disposed verticil-
lately roimd the common stem. The polyps with feelers and
prehensile organs, but without bracts (Gegenbauer), attached to
the remainder of the stem immediately below the swimming
column.]
Phyasophora (fix)m (putra or ^ua-a-a), literally bladder-becMrer, wa«
compared by Fobskael to the so-called Cartesian Imp {situs animalts
hydrostdUictis mMaJtnis pulmone extra corpus, ad formamh machines
quami Diaholwm GaHesicmwm appeUamms), Descr, Animal, p. 112.
Sp. Pkys8ophora kydroatatica F0B8K. Icon. Rer, not.. Tab. zxxni. fig. E, in
the Mediterranean ; Physaoph. muzonema Pkbon, Voy. aux terret ausfrtUe*,
PI ^9, fig. 4; Lesson AeaJ^phen, PL 9, fig. 1, in the Atlantic, &c.
The species are not sufficiently distinguished ; Fobskael figured an im-
perfect specimen, which for the most part had lost the suctorial tubes and
other appendages. The figure given by Phili^pi not long ago represents
the animal in an uninjured state ; this writer supposes that the specimen
found by him in the Mediterranean belongs to another species, which he
names Phyuophora tOrtuticha. There are four rows of swimming bells, and
in each row four. See his Memoir in Mukllxb's Archiv. 1843, <«. 58 — 67,
Taf. v. [KoELLiKEB statos that thia.of Philifpi is identical with the Physto-
phora rotacea of t)BLLA Chatja, */>flj<T. d^U anim. invertehr. TV. pag. 119,
•Tav. 33, fig. ^f]'- ' ^ ^
VOL. T. 8
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114 CLASS III.
[See KOEiiLiKEB'8 description of a new species (1. 1 p. 19, Tab. t.) found by
him at Messina, and which he calls Phyn. PkiUppi. The type of PhysBophara
IB distinguished by the very small length of the axis below the swimming
column. The column is as usual tenninated by a small bladder above the belln
filled with air. Beneath it the polyps, feelers, prehensile and sexual organs
are all compressed into a small space. The feelers surround the axis immedi-
ately beneath the bells in a continuous coronet, like the calyx of a flower.
They are described by Kosllikkr as exceedingly sensitive, in constant
motion, and even laying hold of prey. Within the circlet of feelers arise
the Polyps, each with its prehensile filament The nettle-node has an
exceptional formation. There are no bracts or covers, their protective office
being supplied by the near neighbourhood of the swimming column. The
sexual organs are seated in bunches on the stem dose to the Polype, a pair
of different sex at the base of every Polyp.
c) with long axis without swimming bells.
Rhizophysa P^RON. No swimming bells: the polyps with
their prehensile organs lateral, usually secund : bracts and feelers
wanting.
Sp. Jthizophysa Percnii Esohsch. AeaL Ta£ xm. fig. 3. Rhizophysa JUifor-
mia Lax., ZdUehriftfQ/r msaenach. Zoologie, v. s. 324—330. Taf. xvin. fig.
5—".
d) with long axis or stem and swimming bells at the upper
part of the common stem.
Stephanomia P^KON, EsCHSCH. Swimming bells numerous,
forming a conical column which surrounds the stem with many
spiral turns. Polyps set on the stem by a long, slender, contractile
pedicle. Feelers pediculate aflSxed to the stem, usually in threes
between two successive Polyps. Bracts or covers not confined to
the stem, but also surrounding the base of the Polyps as the calyx
a flower. Prehensile filaments very long with lateral branches
at regular intervals bearing a node and terminating in a single
thread. Sexual organs in bunches close set on the feelers fi-om
the stem.
The Stephanomia uvaria of Lesueub does not, according to
KoELLiKER, difiTer from Apolemia Esohsch.
In a specimen of Stephanomia four feet in length Leuckart
counted no less than 20 spiral turns in the swimming column, with
10 — 12 bells in each turn. The three feelers from the stem are two
on a common pedicle and one sessile. The male and female organs
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ACALEPHiE. 115
are in dose proximity to each other at the base of the double feelers.
The male are the least numerous^ the form oval, slightlj medusan,
^the nucleus as it ripens passing from red to yellow : there are four
radial canals and a circular canal, the mantle lying close to the
nucleus and having an opening. The female appendages are smaller
and round, but of similar structure, except that the central vessel of
the nucleus is not developed, for each appendage contains only a
single ^g. Ck)mpare the interesting observations and figures of
Milne Edwabds Anru des Sc. NcU. 2e S6rie Tom. xvi. Zool. pp.
217—229, PL 7—10. Also R. Anim. Cuv. ^d. iUustr. Zooph. PL
59. See Leuckabt Zodog. Untersuch. Erster Heft. s. 38.
Forskalia Koelliilbb.
8p. FankcUia Bdwardtii Koell. This new genus and species differs little^ if
at aD, from Stephanomia according to Liuokabt. See the description and
beantifal figures in Koelltkkb Die Siphcnophoren v. Metsina, s. i — lo.
Taf. I. II.
Agalmopsis Sabs. Swimming colimm formed of two rows of
bells alternating. Below it, the stem gives origin to single Polyps,
feelers, prehensile filaments and sexual organs with numerous
transparent bracts or covers.
Sp. Agalmoptis degofOB Sabs, Fauna litter. Norvegia i. p. 36, Tab. v. fig. 7, 8.
The prehensile filaments give off branches which again subdivide to terminate
in two threads : a contractile bladder is seated below the point of last divi-
sion, and immediately before this is a spiral nettle-node covered by a beU-
shaped duplicature from the filament on which it is placed. AgalmoptU
puneUUa Kosll., a new species, differs from the last i|i the branches of the
prehensile organs having the node without a cover, and in terminating
without further division: also the feelers have special prehensile organs
which are knotted.
Agalma EscHSCH. Swimming column with bells in two rows :
feelers scattered. Polyps with bracts and prehensile organs of
which the lateral branches are provided with a large nettle-node
and subdivide to terminate in two threads having a contractile
vesicle between them at the division.]
Sp. Agalma OJceni Eschsoh. Itis xvi. 1825, Tab. 5, Sytt. der Acaleph. p. 151,
Tkb. xin. fig. I.
[ApolenUa EscHSCH. Swimming column composed of two rows
of bells of quadrangular form with rounded angles. Feelers firom
two to four set on the stem between successive pairs of bells.
8—2
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116 CLASS III.
Polyps numerous, sessile, prehensile organs with simply spiral
nettle-nodes. Bracts claviform with special prehensile organs small,
knotted. At regular distances below the swimming column a
collection of polyps with all these appendages surround the stem.
Note, — ^In no other genus of Phyiaophorida lune feelers met with on the
part of the stem which supporta the swimming bells.
Steplumomia twaria LX880N belongs to this genus : Kokllikss, Die
Schmmmpolypen, s. i8. See Gboenbaubb's description of a complete
specimen of it, and figure, ZeUechrift f&r vnssensch. Zool. v. s. 319 — 324.
PL xvni. fig. I.
Family III. HippopodidoB, Colonies of swimming Polyps,
without swimming bladder, with short common stem, the swim-
ming column not formed of bells.
Hippopodius QuOY and Gaim. Eschsch. The swimming
column formed of bracts in two rows, and covering one another
imbricately, with filiform short stem, to which the polyps with
their prehensive and sexual organs are attached.
Sp. ffippop, Ivtens, Ann. dee Sc, not. x. 1827, s. 172, 173, PI. iv. A.
GnxBiK leonogr., Zooph. PL xix. fig. 4. — ffippopod. neapolitanfie Kokll.
Die Siphon, pp. 28 — 31. Tab. vi. figs, i — ^5.
Vofftia KOELL.
Sp. Vogtia pentacaniha Eobll. Die Siph, von Meesina, s. 31, 32. Tab.
Family IV. DiphyidcB. Locomotive apparatus of the colony
two distinct cartilagineo-gelatinous transparent pieces aflfixed to the
upper part of a thin cylindrical common stem. The stem begin-
ning in the substance of the anterior piece passes in a groove of the
posterior between the two, and then gives attachment to groups
consisting of a single polyp and its appendages.]
This fiEumly includes certain marine animals, transparent as glass,
which swim by means of the contraction of hollow organs filled
with water ; it has the genus Diphyes for its type, which was first
formed by Cuvieb in the first edition of his R^gne Animal, iv. p. 61.
This genus rested on a species discovered by Bort be Sadtt-
Vincent at the beginning of this century (1801), in the South
Atlantic Ocean, and described mider the name of Salpa bipartita;
see his Voycbge dans les qucUre principaUea ties des Mera dA/Hquej
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ACALEFHiEl. 117
I. 1804, p. 134, PL VL fig. 3, A, B, C. The two pieces provided with
swimming cavities, nearly similar in form, were afterwards, by
CuviEB {B^gne Anvm. sec, ed, iii. p. 288) and other writers,
erroneously taken to be two animals which had become attached
to each other, an opinion occasioned by observing that they
were readily separated. This separation, or spontaneous detach-
ment of different parts, has often been remarked in the entire
order — as in Fhyssophora, Kkiaephyaay StephanomM. In fact,
JXphyes is much more nearly allied to these genera than might
be suspected from many, and sometimes very confused, descriptions
of it. I may remark, that the part which, in our description,
we have indicated as anterior, is called posterior by the first
discoverer, Bory, and by many others after him.
QuoY and Gaimabd, who discovered many new species of this
family and formed new genera from them, {Ann, dee Sc not.
Tom. X. 1827, p. 5 — 21,) determined subsequently to bring them
all \mder the single genus Diphyes, {Voyage de dicouvertea de
FAstrdahe, Zoolog. Tom. iv. 1833, p. 81). [But more accurate
observations of late years have shewn that this proceeding is not
advisable. In Praya the swimming beUs are similar in form,
and are placed, more or less, side by side, and their cavities
open on opposite sides of the stem. In Diphyes the beUs are
placed behind one another and open backwards.
The common stem b^ins within the substance of the anterior
bell, or piece, in a more expanded portion, which is lined with
large epithelial cells, and has very different form in different genera.
This expanded portion often contains a globule of oily matter.
Beneath it the stem gives origin to the canals of the swimming
pieces, and then is prolonged to become the common axis of the
colony.
The polyps with their different appendages are fixed to the stem
at regular intervals. Those nearest to the swimming pieces are
quite undeveloped and without appendages. Those at the other
extremity of the stem are the oldest and most perfect, and have
their appendages most complete. Each group consists of a Polyp,
a set of prehensile organs, and the generative organ, which partakes
more or less of the medusan form — ^the whole being covered by
a protecting bract. Such a group either persists in adhering to the
common stem, {Diphyes^ Praya), and then only certain parts are
detached, as the medusiform capsules of the sexual organs ; or it is
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118 CLASS II r.
capable, to a certain degree, of independent existence, and when
fully developed, separates itself from the stem of the colony {Ahf/lay
In the Praytdas, the coTers are bounded by round sur&ces above,
and have a cavity below, like a helmet, to receive the other members
of the group : in Diphyea they surround the stem like a rolled leaf,
(differing, however, in form in the different species) and adhere to
it by the narrow part In Ahyla they cover the members of the
group imperfectly, are almost solid, yet with a cavity in the interior
which is in connexion with that of the stem, and which is lined by
large cells. In all these groups the sexual buds are more or less
medusiform, and may be developed into a swimming belL They
sprout from the base of the Polyp.]
Comp. on this family, besides the above work of QuoT and Gaikabd,
especially Esohsoholtz, Syst, der AhaUphen, s. 172, 133, Lbuckart Zodlo-
giache Untersuchungen, i. s. 41 — 49, Koelliksr Die Sckwimmpolyp. s. 36 —
46 : also Will, nora TergestincR, s. 76 — 83, Gboenbaubb in Zeiitchr, f,
wi88, zool, V. 8. 197 — 300.
[Diphyes Cuv. The posterior swimming piece received into a
cavity of the anterior : the groups on the common stem protected by
a bract or cover in form of a rolled leaf.
Sp. Dipkyes angustata EsoHSCH. Tab. xn. fig. 6 (the species of Bobt seems
to belong here;) — IHph. campawdifira Esohsoh., Quot and Gaihabd
Ann. dea Sc. not. x. 1827, PL I. ; IHph. gracUis Gbgkkbaueb ZeiUckr. far
wiateMch. Zoologie, v. s. 309—315- Taf. xvi. fig. 5— 7.— />tpA. Siebddei
KOELL. Die Siphon, v. Messina, s. 36 — ^41. Taf. xi. fig. i — 8. — Diph. gra-
cUis Gegbnb. Zeitsehr. /. w. Zool. v. s. 313 — 315. Taf. xvi. fig. 5 — 7. —
Diph. Kochii Will Hot, Tergest. Tab. 11. f. 21—16 ; figured without the
posterior portion which Will did not meet with in any one of the six speci-
mens examined by him^.
Ahyla EsCHSCH. QuOY and Gaimard [Calpe of the same).
The two swimming bells of very diflFerent size, the anterior mnch
the smaller. The bracts cover the members of the groups imper-
fectly, are massive, and have a cavity communicating with the
stem.
Sp. Ahyla, pentagona Esohsch., Koellikbb Die Siph. v. Mess. s. 41 46,
Tab. X. ; here the single polyps have no coTers : see a complete specimen
described and figured in Lecokabt Zod. Untersuch., i. s. 56—61, Taf. nr.
fig. I— 10. The bracts, or covers, which are not visible on the polyps at
^ Compare also on this genus Lesson Centurie Zoologiqtte, 1830, p. 16 t — 183,
1*1. 55—57.
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ACALEPH^. 119
the upper pari of ihe item, undeiigo remarkable metamorphooes after Uieir
first appearance aa buds until they attiun the cubical form, when the group
of which it fonnB a part exactly reaembles a young Eudana cvboidea, QuOT
and Gaim. See also Geoinbaueb ZeUach. fur tns$. Zool. v. s. 191—195.
Fratfa Lesson. The two swimming pieces of the colony nearly
similar and equal, the covers of the developed groups bounded by
rotmd surfaces above and concave beneath.
Sp. Praya diphyea Lsss., Koellikeb Die Siphon, von Me$». s. 33— 3<5, Taf.
a,—Pngya maaeUna Gegekb. Zeitach. f, wiaamich. ZooL v. a. 3o»— 309» Taf-
xvn. fig. I — 6.
The genera Endoxia, Ersasa, Agktisma Esohsch., which have
only a single polyp, have been termed monogastric diphyida^ by
Huxley j but it is almost certain that they are not independent
genera. It has been noted above, when treating of Ahyla petUa-
gona, that a single group of this compound diphyea exaxstly resem-
bles Eudooda cuhoides; and here the groups have been seen to
detach themselves from the colony both by Leuckabt and by Geqen-
BAUEB — as indeed the same &ct had previously been observed by
Sabs in his Diphyeg tnmccUa, Eudoxia campantUata is believed by
Leuckabt to be a group of Diph. acuminata, a new species observed
by him at Nice ; whilst he has found that Aglaisma pentag<mum is
not a monogastric diphyea , but an imperfectly developed Ahyla pen-
tagma, see Zoologiach, Unterauch. s. 54. Eracea is suspected by
Leuckabt to be a detached group of Diphyea Koch, Wilu
Evdoxia (&c.) consists of a cover or bract, a polyp with its pre-
hensile organs, a swimming bell (sexual capsule), and usually a
smaller bell sprouting firom the base of the polyp, which is destined
to replace the larger when this has been detached. These ports
are all connected by their canals to a portion of common stem.]
Sp. EnoBa pyramidaUs Will, i. I fig. 27 > &c- ;— Comp. Leuckabt Zocl,
Untenuch. 1, s. 43—61, Gegbnbaubb Zeiisch. f. mn, Zool. v. 185—296.
Ordeb II. CtenophorcB, or Beroecea.
Mouth simple, stomach situated in the axis of the body. Vibra-
tile cilia disposed in rows on the surface of the body. Swimming
bladders none.
The Beroecious animals are Acalephs of very different form,
which, however, are distinguished from the former order by the
absence of swimming bladders [bells] and cartilaginous laminae, as
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120 CLASS III*
well ajB suctorial mouths : [they are single animals, in short, and not
colonies.] The projecting edges, usually named ribs (coattB) which
are beset with cilia, especially characterise this family : whence the
German name BippenqtMUen. Whether these vibratile cilia, which
occasionally are so arranged as to form vibrating laminse, do really
cause the progression of these animals, as is usually assumed, is in
consequence of the objections raised by Mertens and by Will
{ITorcB Tergest b. 8 — 13) exceedingly doubtful.
The name Beroe given by Beown {N<U. Hist of Jamaica) to the
animal discovered by him in the middle of the last century, is
borrowed from Mythology; it is that of one of the numerous
daughters of Oceanua :
CUoque a Beroi soror, OceaniUdei amho, —
ViBOiL, Gtorgic, Lib. iv. 341.
Comp. on this order : Bang, hablitaemaa de la FamUU des BSroidet et
description de deux genres nouveaux qui lui appartiennerU ; MSinoires de la
Soc. d'ffist, not, de Paris, Tom. iv. i8a8, pp. 166 — 173, PL 19, ao.
Mebtenb Beobachtungen und Untersuchungen Uber die heroearligen Aca-
lephen, MSm, de VAead, imp, des sc. de St. Petenixmrg, sc. physiq, Bixibme
s^rie, Tom. 11. 1838, pp. 479—543, Taf. i. — im. (A copious extract may
be found in Oken'b /<nf, 1836, b. 311 — 321.) Lesson, Mim, aur la
famille des Biroides, Ann. des Sc. not. 1* s^rie, Tom. vi. Zool. 1836,
PP- «3S— «^-
Family V. Beroidea. (The characters of the order are those
of the single family.)
A) Stomach smalL
Cesium Lesueur. Body transverse, elongate, gelatinous, with
ciliated margins.
Sp. Cestum Veneris Lesueub Nouv. BuUet. de la soc. philom. Join, 18 13,
PI. V. (Recus. in Oken's Isis, 1817, s. 1505—1508, Tab. xn.) Gdi^bin,
Iconogr. Zooph, PI. 18, fig. i. (after a drawing by LAUBHiLABD) in the
Mediterranean. This girdle of Venus has the form of a band of more
than fiv^ feet long, and full two inches high. In the thinner inferior edge
is situated the oral aperture (opposite to the phbce assigned to it by
Lesueur in the thicker superior edge). In Cestum Najadis Esohsoh.
Acal. Tab. i. fig. i, from the South-Sea, near the Line, two long tentaeula
beset with fine threads ore present, which in the species from the
Mediterranean are often, and in Cestum Amphiirites Mebtenb (1. L Tab. i.)
are (always ?) wanting.
The genus Lemniscus QuoY and Gaim. is probably founded on a detached
piece of Cestttm.
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ACALEPH^. 121
CaUtanira P^RON. Body lobate or supplied with lateral
wings.
Subgenera : Eucharia Eschsch., Leucotkea Mebtens, Mnemia
EscH. {Alcinoe Eang), Leaueuria Milne Ebw., Calymma Eschbch.
{Ocyroe Rang), CaUianira Peron, Eschsch.
In bringing these numerous genera together^ and giving greater
extension to the name CcUlianiray than has been done by former
writers, my sole object is to &cilitate the review, and at the
same time to indicate the affinity of these animala Beyond doubt
the genera are too numerous here. The genus Bucephalon of
Lesson {CaUianira bucephalon Eeynaud, Less. Cenlwr, zool. PL 28)
also belongs here, and probably does not differ from Calymma
Trevirani
Sp. CaUianira heaoagona Eschbch., CaUian. SlahUri Db Haan, Natutir'
hundige Bijd/ragen II. 1827, pp. 150 — 153; this species has been con-
founded with Bero6 keaoagonus of BBUOUiaBBS (found at Madagascar).
In the genus CaUianira proper, there are two filiform branched tentacles ;
the other subgenera have mostly four conical or triangular ciliated
tentacles.
Cydippe EsCHSCH. {Beroe Freminville, Mertens.) Body
globose or ovate, with eight longitudinal, ciliated ribs. Tentacles
two, retractile within two subcutaneous vesicles,
Sp. Cifdippe pHeus, Bero€ piUus Muell., Volvox hicaudaltu L. ; L. Th.
Gbonovius in UUgezocfUe Verhandelingen Amsterd. 1758, in. p. 464,
PL 16, f. 1—5 ; Baster Natuwrh, UUsp. i. PI. xiv. fig. vi. vii. ; Cuv. R,
Ani, id. iU.f Zoopk, PL 56, fig. a, &c. (Since the vesicles, in which the
tentacles he, open towards that extremity of the body which is opposite to
the mouth, the nervous ring (?) described by Grant, cannot surround the
mouth, (see above, p. X04). The tentacles are able to lengthen themselves
greatly ; why Eschbcholtz describes them as incUvisa, is not apparent ; at
least in most species they are much divided.
B) Stomach large A circle of vessels (aqueducts) round the oral
aperture.
Beroe EscHSCH. (spec, of the genus Beroe Gronov., Muell. and
others) Idi/a Freminv., Oken, Mertens. Body oval, ribbed,
with large circular aperture beneath.
Sp. Bero€ ovata Browk, not. Hist, of Jamaica, PL xrv. fig. ^ ; (this animal
was the first named Beroe ; LiNMiEUS named it in the tenth edit, of the
Sytt, Nat., Medusa Berog, in the twelfth (1767) Volvox Beroe) ^—Beroi
Forskalii, Medusa Beroe Forde., Milnb Edw. Ann. des sc. not. 0? s^rie,
Tom. XVI. Zool. PL 5, 6 ; Cuv. B^gne Anim. id. iUustr., Zooph. PL 56,
fig. I, &c.
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122 CLASS III.
Order III. Discophorce,
Body disciform or campanulate, above naked, below usually-
provided with arms or tentacles.
The Meduacnda or SecMvhber. They have a gelatinous disc^ on
the upper surface more or less spherical, which from its form has
been compared to an umbrella or a hood ; the form has some re-
semblance to toad-stools {agarici). These animals move themselves
by expansion and contraction of the hood. The mouth, or the suc-
torial organs which take the place of the mouth, are situated in the
center of the inferior surface, sometimes elongated into a pedicle
and provided with different tentacles. On this difference are
foimded the numerous genera which modem writers have felt jus-
tified in adopting.
Compare on this order : P^bon et Lesusub, Tableau des earac-
Ures ghiiriqiLes et specifiqtie3 de toutea lea eap^cea de Miduaea can-
nueajuaqtCd, cejour. Annal. du Muaeum xiv. 1809, pp. 325 — 366.
J. F. BRAinxr, Auafiih/rliche Beachreihung der van C. H. Mebtens
aufaeiner WeUumaegdung heobdckteten SchirmquaUen ; mU 34 Tneist
colorirten Tafeln. St Petersbui^, 1838, 4to (firom the Mem. de
VAcad, dea Sc de SL Feterabua-Qy vi*. Serie, Tom. iv.)
A) Many oscules.
Family VI. Oeryonidce. A peduncle from the center of the
inferior surface of the disciform body, with the free extremity
lobate, or furnished with arms. The border of the body mostly
tentaculate. {Genua Diancea Lam.)
It is not so completely established that all the forms here brought
together are really characterised by the absence of a simple mouth.
Will at least, in the animals placed by him in the genus Geryonia,
found a mouth surrounded by four lobes. In some the pedicle is
supplied at its extremity with a folded appendage (Gerf/onia), in
others at its base, or at its extremity, it is beset with threads :
Favonia, Zi/mnorea, &c.
Genera : Geryonia Pj^ron, Froboacidactyla Brandt, Diancea,
Linuche EscHscH., Saphenia Eschsch., Eirene Eschsch., Limnorcea
Peron, Fa/vonia Peron.
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ACALEPHJi:. 123
Bp, Oerffonia proboieidalii, Medtua proboicidalit FoBSK. lean. rer. not, TWb.
56, fig. I ; GniBiir Iconogr,, Zooph. PL 16, fig. 9 ; CcviiBy Jt, Anim,, idU.
Ukutr., Zooph,, PL 51, fig. 3. This species from the Mediiernuiean, with
BIZ threads or tentacles at the margin of the disc, may be considered as
the type of this divisioD.
Family VII. Ehtzoatamidce. Anna ramose, with manj suc-
torial oscules. Margin of the hody without tentacles* Disc with
four avaria or testes, sometimes (in Cassiqpea) eight.
Rhtzostoma CuY. Tentacles amongst the arms none; arms
confluent into one pedicle inserted in the disc.
Sp. Bhuottoma OuvierU, Rbauvub Mim. de VAcad. dei »e, de Paris, 17 10,
PL XI. fig. 17, 18 ; CuviBB Journ, de Phynque Tom. XLIX. p. 436 ; Cuv.
R. Anim, Sdit, HI. Zooph. PL 49. This species sometimes attains to
a great size. The Jthizostoma has four pairs of suctorial anns, which are
provided with absorbent vessels ; by these it receives its nutriment, which
consequently consists of minute animalcules, or of animal matters in
solution. These absorbent vessels and their branches coalesce into four
stems, which run along the pedicle and end in the stomach. From the
stomach run laterally vessels through the hood. Surrounding the stomach
are four cavities, with very wide opening below, in which the organs
of propagation are seated. The uppermost portion of the hood consists of
a substance more firm than the rest of the body. See K. W. Etsbvhabdt,
ZUr AntUomie und N'aturgeachichie der QucdUn, Nov. Ad, Acad, CctM^
Leop. Carol. Nat. Curios. T. X. pp. 375, &c. with figures. RKieoatama
Aldrovandi Pi£bon, Guebin Iconoffr., Zooph. PL 15, fig. i, &o.
Casswpea Peron. Tentacles amongst the arms none. Arms
eight or ten, very much branched, not conjoined at the base into a
peduncle, famished with yesicular appendages.
Sp. Cassiopea frondosa, Mediua frondo$a Pall., Spic. Zool. X. Tab. n. ^,
I — 3 ; Ccunopea horhonica Delle Chiaje, Mimorie wUa ttoria e notonUa
degli Animali $ema vertdn^ del Regno di Napcli, I. 1823. Tab. m. ; Gu&iN
Icon. Zooph., PL 15, fig. 1 ; Cuv. R. Anim. 4dU. HluttT., Zooph, PL 51,
fig. 1, &c. (See other figures of Tilesiub Nov. Act. Acad. Cau. Leop, Car,
Natur, Cwrios. Tom. XV. 2. 1831, pp. 247—288, Tab. 6g — 73.
Cephea Peron. Large cirri amongst the arms.
Sp. Cephea eyclophora P^bon. Meduta cephea Fobsx. Icon, rer. nai. Tab.
XXIX. fig. (copied in Ccv. R. Anim. 6dU. ilhutr., Zooph. PL 51, fig. 4) &c.
B) Mouth single central.
Family VIII. Medustdea. Mouth teti-agonal central. Arms
four, mostly very distinct, very rarely none. Four lateral cavities
in the disc, open beneath, inclosing the genital organs.
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124 CLASS III.
This ^milj nearly coincides with the genus Cyancea Cut. The
four openings beneath the disc, conducting to the cavities which
contain the organs for propagation, were by Pebon and Lamarck
incorrectly considered to be four mouths.
Cyancea Cuv. (and species of the genus Pelagia ejusd.)
Grenera : Stiienonia Eschsch., FhaceUophora Bb., Cyancsa
EscHSCH., Aurdia Peron; Pelagia Peron, Chrysaora Peron,
Ephyra Eschsch. {Buryale and Epkyra Peron.)
Sp. Cyancea awrita, Medusa aurita L. ; Mubllbb Zoclog. danic. Tth. 76, 77 ;
Ehbenb. Ahhandl. der AJcad. zu Berlin, phynk. Klasse 1835 ; CuY. R.
Anim. 6dit, ill., ZoopK The four anns are considerably longer in old than
in younger specimens ; these arms consist of two laminae crumpled at the
edges, which during life face each other in such a way as to form a canal ;
afler death they are flaccid and parted asunder. The disc is not quite
circular, but in some degree divided by indentations of the margin into
eight lobes. The four arms unite at the center of the body to form a
circular aperture : this mouth leads to the stomach, which has four lateral
cavities. From the stomach there run sixteen vessels to the margin of the
duic, of which eight, divided into branches, alternate with eight others un-
divided and open at the margin. In addition, there are eight corpuscles at
the margin, which Ehbsnbebg considers to be eyes, and which were^ noticed
above. This species is found in the North Sea and the Baltic. Comp.
H. M. Gaedb Beitrdge zur Anatomie und Phynologie der Medtuen, mil
2 Kupferiafdn, Berlin, 18 16, 8vo ; Baeb Ueber Medusa aurila, Mbckel*S
Arckiv fiir die Physiol, vin. 1823, s. 369 — 391, with fig. ; F. Rosenthal
Beitrag zwr ATtaJtonde der Quallcn, £eitschrift fiir Physiol., herausgegeben
von F. TiEDEMANN, G. R. und L. C. Tbevibanus, i. 1, 1825, s. 318 — 330,
with fig.
Cyanaa capillaia, Medusa capillata, Basteb Natuurk. Uiisp. il.. Tab. v.
fig. I.
Pelagia noctUuca EsOHSOH., Medusa noctUuca F0B8K., Waqneb Bau der
Pf^. noctUuca and Icon, Zool, Tab. xxxiii. ; in the Mediterranean, &c
Ephyra EscHSOH., probably rests on young forms of Cyancea; comp.
Will ffor, Tergest. Tab. n. fig. 10, and Sabs in Ebichson's Archiv, 1841,
Tab. II.
Family IX. Oceanidce, Disc without lateral cavities to in-
close generative organs. Body campanulate. Mouth and oeso-
phagus often elongated into a proboscis. Anna conspicuouB or
lobes around the mouth. Canals proceeding from the stomach
elongate.
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ACALEPH^. 125
Oceania Peron (with the addition of several species, and
other genera).
Subgenera: Ocecmia Peron, Circe Mebtbns, Cants Brandt,
CcUlirhoe Peron, ThaujnanHas Eschsch., Tima £schsch., Mdicer-
twn Oken, Cytasis Eschsch., Phorcynia P^bon.
Sp. Oceania martupialis EscHSCH., Medtua manupialis L. ; Plano. de
Conch, min. not. Tab. TV. fig. 5 ; MiLNB Edwabdb, Ann. de$ Sc. not. xxvni.
1833, pp. 248 — 266, PI. II — 13, Mediterranean; — CaUirhoi Bcuteriana
PfooN, Babtxb NcOuurh. ViUp. 11. Tab. v. fig. 2, 3, &c.
Family X. JEquoridce, Disc without lateral cavities, inclosing
organs of generation. Body depresso-campanulate or plane. Mouth
and oesophagus not elongated into a proboscis. Arms none or
little evolved. Stomach with sacculated appendages or canals
radiating, elongate, numerous.
JEquorea Peron, Cuv.
Subgenera : jEqtiorea P^ron, Stomohrachium Brandt, Mesonema
EscHSCH., Ev/ryhia Eschsch., FoLyxena Eschsch.
Sp. Mqyorta PorgkaUrta Eschsoh., Medusa aquorea F0B8K. Icon. rer. not.
Tab. xzxii. ;—jEquorea violacea MiLNB Edwabdb Ann. da Sc. not. le
B^rie, Tom. xvi. Zool. pp. 193 — 199 ; Cuv. R. Ani. id. HI., Zooph. PL 71 ;
the maigin has many conical cirri, the mouth Ls wide and round ; from
the stomach proceed about eighty long undivided rays (water-<»mals), which
run towards the mai^gin, and appear to open on a small conical point
between two drri. The genital organs are situated below on the disc, on
each side of every ray as folded borders, but they do not extend as far as
the margin. (Esohboholtz divided the Diseophorce into Oryptocarpce and
Phanerocarpce, Sytt. der Acal. p. 41 ; to the last, which have the sexual
organs placed crucially in the disc and attracting observation by their
colour, belong the JthieottomidcB and Medusidce ; to the first the Oeryonidce,
Oceaaiidce and ^quoridcB ; in all of these the sexual oi^gans have not yet
been detected, but they will probably be found at the under side of the
disc, and since in uBquorea vidaeea, according to the observations of
MiLNB Edwabds, they strike the eye on this surface, and are also dis-
ting^hed by their violet colour, we cannot accept the name Crypto-
carpce.
[The Cryptocarpse of Esohscholtz include the naked-eyed Medutce of Fobbbs.]
Note to the Diseophorce. There remain certain genera ofauthors,
in which a mouth has not been found, namely Evdora and Berenice
PisoN. (Sp. Berenice rosea, Cuvieria Perok Voyage attx terr,
austr. PL 30, £ 2 ; Gu^rin Iconogr. Zooph. PI. 16, fig. 1.) These
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126 CLASS III.
genera, to which maj be added Stav/rophora Bbandt, form a family
in the system of Eschscholtz^ to which he has given the name
BerenicidoB, This fiunilj appears to me to be doubtfiil, as it does to
Brandt and others ; we must leave the decision to time, I prefer
to wait the resnlt of new observations, rather than to attribute to
Acalephs nutrition by superficial absorption.
For the rest, several genera of Acalephs are founded on figures of
authors alone— and these sometimes imperfect and faulty. Hence
no part of Zoology is more uncertain than this.
Accordingly there are many genera which I have not recorded,
and possibly more might have been omitted. For here excess of
timidity is better than dearth of prudence.
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CLASS IV.
ECHINODERMS {ECHINODERMATA)\
The name Echinodermata was used by J. T. Klein rather
more than a centuiy ago, to denote the shells of the sea-urchins,
called Echini. *Extiw, with the Greeks, denoted both an urchin
and a sea-urchin. Afterwards BruguiAres gave the name of
Echinodermata to a division of the animal kingdom that comprised
at once the sea-urchins and the star-fishes. Cuvier retained that
name for this division, but added to it animals having a coriaceous
skin without spines or quills. Although the name is no longer
applicable to all the species of this class, yet the class itself must
be looked upon as an extremely natural division of the Animal
Kingdom. These animals are distinguished by an intestinal canal
hanging firee in the cavity of the body by single threads or mem-
branes, usually long and tortuous, or provided with lateral append-
ages if it be short: blood vessels and respiratory organs are present;
the genital organs, in both sexes of the same external form, are
without any special apparatus for copulation ; the fertilisation being
1 Consult on this class :
L. Agassiz, Monoffraphdes d'Echinodermes vivans etfoasiUa. Neuch^atel, 1838 and
foU. 4to. ^e ])«rt8 thus far have reference only to certain figures of Echmi. The
fourth, pubHflhed in 1843, contains the anatomical description of Echinus Uvidus, by
Valbhtih.)
£. F0BBB8, A HUUynf of BrUUh Star-JUhes and other animaU of the doss Echino-
denuMkt, lUuttraUd fty woodeuU. London, 184T, 8to.
A capital work on the anatomy of this class is :
F. TrgOKMAify, AnaionUe der Jt&hrm-ffolothurie, de$ pomeranz-farbigen Seettems
und Stein-SeeigeU. Landshut, 18 16, folio; a prize Essay crowned by the French
Institnt in i8i3.
W. Shasfet, The article EchinodemuOa in Todd's Cydopadia of Anai. and Phy-
Mog. n. pp. 30—46 (1836).
J. MnsLLSB, Veber den aUgememe Plan der Entmckektng der Eckmodermen, Mit 8
Kupfertaf, Berlin, 1853, 4to, and previous papers in the Ahh. d, Ahad. der Wisten-
icke^. tu BerUn 1849, '^5^
Q. L. DuTXBNOT, M^, 8ur VAnalogie de CompotUicn d eur qvdguee poinU de
TOrga$id$aHon dea Eehinoderms, Mim. de VAead. det Seienceij XX. Paris, 1848.
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128 CLASS IV.
eflFected by means of the sea-water in which these animals live.
When parts are multiple, the number five prevails as remarkably
in this class, as did the number four in the preceding: and the
quintuple organisation is often obvious, as in star-fishes and sea-
urchins, in the external form of the body. This is round or some-
what pentagonal in the sea-urchins ; flat and spread out in rays in
the star-fishes, with the mouth on the inferior surface. The Holo-
ihurm haye, on the contrary, a cylindrical body.
In the star-fishes, of which the body is flat, the mouth conducts
to a wide stomach that fills the disc of the body. It was supposed
formerly that this stomach was in all of them a blind sac, as
TiEDEiiANN has described it in Asterias aurantiaca {Astropecten
aurantiacus MuELL. and Tr.) Afterwards Meckel detected in
Comatuke a second opening of the intestinal canal, that lies on the
same surface of the body with the mouth \ Only lately it has
become apparent from Mueller's investigations, that in most of
the proper Asterioe an anvs exists, and that the structure in Ast.
aurantidca is to be considered as the exception rather than the
rule. But this second opening does not lie, as in the Camatuke,
on the same surface as the mouth, but opposite to it, on the back
of the disc. In the Ophturce and Euryale it is wanting: conse-
quently these, with some star-fishes (the genera Astropecten, Cteno-
dtscics, and Lutdia of the modems), and some Crinoids are the
only Echinoderms in which the intestinal canal forms a blind sac,
as in the Anthozoa. In the Astertce proper, the intestinal canal has
csBcal appendages, which divide into branches, and fill the rays of
the body ; in those species whose intestinal canal is a blind sac,
the appendages proceed laterally firom the stomach, at whose base
on the dorsal surface there are usually two caecal appendages in
addition. In those Asteriee which have an anal opening, the
stomach is divided by a circular fold firom a second compartment,
to which the c«Bcal appendages of the rays are attached ; to this
succeeds a third compartment, the rectum, a short straight tube,
which has also caecal appendages ; sometimes they are placed round
the intestine in rays, like the appendages of the rays, and fill up
^ Archiv fUr die Physiologie vni. 1833. s. 470 — 477. The same observation was
made by Leuokabt and HEtiBiNOKB ; see the not very clear description by the last-
named in Mboksl's Archivf. Anat. u. Phynol. 1826. s. 317 — 324.
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ECHINODERAIS. 129
the spaces between the rays. In the disciform genus Guldtay the
appendages of the rectum are greatly developed, five in number,
each divided into two branches, and clustered ^ In Ophiura and
Euryale the csdcal stomach has lateral recesses, or even branched
blind appendages, mostly ten in number, which, however, do not
penetrate the rays. In Comatula the intestinal canal is tubular,
and winds round a spongy structure in the axis of the disc ; firom
this an edge projects, that penetrates into the canal and forms a
valve*. In the sea-urchins {Echinus) the intestinal canal is very
long. The oesophagus is tortuous, narrow, and beset with numerous
follicles. Where it passes into the much wider intestinal canal,
there is a csecal appendage. The walls of the canal are very thin :
its course is close to the shell in five afcs directed outwards ; when
it has returned nearly to the point firom whence it began, it bends
round and follows a similar route in an opposite direction, until at
last, having become somewhat narrower, it mounts up to the anus
(at the uppermost part of the shell). In the Holothurice the intes-
tinal canal is nearly of the same width throughout. It proceeds
firom the mouth along one side of the body to the lower extremity,
thrai bends back to the anterior part, and finally descends along the
other side to the cloaca, into which the respiratoiy organs also
open. In Eckiurus the intestinal canal is, in like manner, much
longer than the body, and makes many convolutions: it has numer-
ous cystiform widenings, and very thin walls. In Sipunculus,
where the anus is placed not at the end, but in the anterior half of
the body, the intestinal canal, with its threefold bending, is nearly
four times the length of the body. In Bynapta^ on the other hand,
it is nearly straight, and about the length of the body, the anus
being at the posterior extremity'. In the star-fishes probably the
radiating appendages are to be considered as organs for preparing
bile (liver) : they are filled in Ast. rvhem with a yellow turbid
fluid*.
* J. MusLLEB und F. H. TROiCfHBLL^ Sydem der Aateriden. Braunschweig, 1842,
4to, B. 131. Taf. zii. fig. T.
* J.MiJVLLB&,Abhandl,derBerl. Akad. a, d. J, 1841, Phygik.Kl Tab. v. f. 7—10.
' QUATBBFAGBS, Ann. de$ Se. not, sec. s^rie xvii., Zoologie, p. 51.
* Other writers oonsider the blind appendages at the bottom of the stomach or at
th6 rectom as a rudiment of Uver. Owen, Led, on the Camp. A not. of invertebr.
AmmalB, 1843, P- 1 15- In these appendages a rudimentary form of kidney might also
be repognised, an opinion, however, which does not rest on chemical investigation.
VOL. I. 9
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130 CLASS IV.
Notwithstanding much careful investigation, there still exists
great obscurity about the circulation of the blood in Echinoderma.
TiEDEHANN and Delle Chiaje give very conflicting descriptions
of the vascular system — ^the diflference being founded in the inter-
pretation of the dermal vessels, which are connected with the
organs of motion. The first of these authors considers the motion
of fluid observed in these vessels to be altogether distinct from the
circulation, whilst, according to the other, they are a part of the
system of blood-vessels. In Aaterias Tiedemann found on the
inner surface of the skin of the back, a vascular ring, which he
considers to be venous. The vessels which run upon the surface
of the visceral appendages of the rays open into this ring. From
it a canal arises, which performs the office of a heart, lying near
the so-called lime-canal which is found there. The canal runs
into a vascular circle surrounding the mouth, which Tiedemann
holds to be arterial, and from which branches proceed to the
intestines. Besides these two vascular rings (one on the dorsal
and one on the abdominal surface), there is a third ring of an
orange-yellow colour found on the inferior surface beneath the skin.
Tiedemann was not able to discover any conununication between
this ring and the rest of the vascular system. In Echinus vascular
rings occur, in like manner, round the mouth and the anus, on each
surface two, of which one is to be considered arterial, the other
venous. The heart is oblong, divided into many cells, and lying
on the oesophagus\ In Hohthuria there is a circulating system
without a heart, or rather the heart has the form of a contractile
vessel, that runs at the outside on the surface of the intestine.
At the anterior extremity of the intestinal canal this vessel forms a
vascular circle, whence very fine branches arise; when near the
anus it has become small, having given off a multitude of fine
branches, which run on the surface of the intestine. There is a
transverse vessel which connects the longitudinal trunk on the first
loop of intestine with that on the second. Many intestinal veins,
which seem at the same time to perform the part of absorbents or
^ Comp. the descriptioiifl and figures of Yalkntik, AnatonUe du geiire JBchimu,
pp. 89—96. Tab. vn. fig. 119, 115, 127. Tab. vm. fig. 144—159, &c. There U a
figure also of Uie heart and part of the blood-vessels in Spaiangut in CuviBBll JUgne
Anim. id. iUutlrie, ZoophyUt, PL f i bis.
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ECHINODERMS. 131
lymphatics, run upon the mesentery to form a stem, having a
carved course, firom which other vessels arise to run to the respira*
toiy organ and so may be named pulmonary arteries. With these
pulmonaiy arteries the pulmonary veins are in connexion, from
whose union a longitudinal stem arises from which branches proceed
to the arterial vessel with which we began our description *.
Besides the blood-vessels already described there are other
vessels which in Echinoderms provided with suckers or feet are in
connexion with these organs of motion. The integument of the
body is perforated by numerous pores arranged regularly in rows ;
in the sea-urchins the rows have been called, on account of their
regularity, Ambulacra, from a comparison with orderly rows of
trees and garden-walks. Through these pores membraneous cylin-
drical feelers (the feet) pass out, each terminating in a minute
suctorial disc According to the investigations of Valentin these
feelers are in Echini perforated at their extremity by a fine aperture.
Within the integument there are vesicles in connexion with them.
The feelers, hollow within, are filled with a fluid, usually sea-
water, which the animal can press at will from the vesicles, or, by
contraction of the former, can cause to flow back. In this way the
animals move their body, the numerous feet contracting and elon-
gating, and adhering by means of the suckers. There are vessels
corresponding to the rows of feet or feelers, from which lateral
branches proceed to the vesicles of the feelers. The ordinary
number of these longitudinal vessels of the integument is five ; in
the star-fishes their number corresponds with the number of the
rays of the body. These lymphatics fall into an annular vessel
surronnding the mouth. In Holothuria the appendages of the
feelers which surrotrnd the mouth proceed from this annular vessel :
and from it there arise also five other vessels that descend along the
commeacement of the intestinal tube, where they terminate in
another annular vessel from which one or two oblong csBcal vesicles
depend {AmpuUa PoliaTui), that are in like manner filled with
watery fluid*.
The change of the blood from venous to arterial, the proper
1 See T»D«MAKir, Anai. der HOhrm-ffotoOiftrk, s. 15—18, Tab. in. ; oomp. also
CuvuB, lUgne Anim., €diU HI., Zoophytet, PI. 18.
* See the figures in TniDiitAKN, Tab. in. fig. 4, 6.
9—2
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132 CLASS IV.
object of respiration, may be effected in any part of the animal
organism, where the finest branches of the blood-vessels (the
Capillaries) are bathed in the medium in which the animal lives.
Hence it is easy to conceive that this function is not always neces-
sarily connected with determinate parts.
In almost all Echinoderma the sea-water penetrates into the
cavity of the body, and bathes as well the internal surface of the
integument as the outer surface of the intestines. Where no
especial respiratory organ exists, the function of such an organ, the
change of the blood, may be effected in the fine vessels which run
on the surface of the intestinal tube.
In the Star-fishes the sea-water penetrates to the cavity of the
body by means of fine tubules in the integument, which are found
in great numbers on the dorsal surface ^. In Ophiura there are on
the abdominal surface in each of the five fields between two rays,
two or four fissures leading into the cavity of the body.
In the Echini it is not known with certainty in what way the
water penetrates the cavity of the body. The ten branched organs
round the mouth, which Tiedemann considers to be tubules to
convey the water in and out, have, according to Valentin, no
external apertures *.
As little is known hitherto of the course which the water takes
in most Holothurice to reach their cavity. In those which have no
special respiratory organ, the genus Synapta has between the ten-
tacles that surround the mouth four or five small papilliform
eminences, having an opening at the apex and conducting to as
many tubules that open between the muscles of the mouth. The
openings are beset with cilia, like the tubules of the integument in
Star-fishes*. In other JSolothuricBy as in those which Tiedemann
investigated, there are special respiratory organs. From the Cloaca
in which the intestinal canal terminates, there proceeds upwards
a short tube, that soon divides into two very long principal branches
which run as far as the anterior part of the intestinal canal. From
these smaller tubes arise which subdivide into twigs which termi-
^ [This is Tiedemakn'b opinion with respect to star-fishes, but the obsenrations of
Shabfbt, Ehkenbbbo and Mdkllbb, are opposed to it ; they saw the streams of
water from within turn bacl^ when they reached the extremity of the tabules.]
* Valentin, op. cit. p. 83.
* QUATBXFAOBS, Op. dt. p. 65. PI. 5, fig. f, /.
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ECHINODEBMS. 133
nate in csecal vesicles, or pulmonary cells. The right branch is
intimately connected with the intestinal veins ; the left branch of
the respiratory organ is connected, by means of muscular fibres,
with the internal surface of the integument. The form of this
respiratory organ agrees with that of Lung, although HolothurtoB
breathe water and not air. These parts are very contractile : in
a Holothuria that was opened alive they did not cease, as long as
life lasted, to force the water in and out by alternate contraction and
expansion. But in respiration it is not the contraction of the
muscular membrane alone of these branches that acts, but the
contractility of the common integument of the body also. This con-
tractility of the skin is so great, that occasionally, when the creature
is irritated, a portion of the intestines together with the right
branch of the respiratory organ is forcibly ejected from the Cloaca.
In the Sea-urchins Valentin considers the ten branched organs
surrounding the mouth, first described by Tiedemann (and noticed
above, vid. p. 132), to be external gills. As internal gills Krohn*
and Valentin consider the foliated vesicles, which, in the interior
of the shell, are in connexion with the ambulacral tubules : and
which have a closely-woven vascular net-work. Valentin found,
as has been stated, the ambulacral tubules perforated at the extre-
mity in Sea-urchins. Through these openings the water penetrates
into the vesicles, and the general opinion that the fluid is urged
into the tubules from the vesicles and so distends them is not valid,
according to Valentin, in the case of Sea-urchins*. In that of
the Star-fishes and Hblothurtce, where the tubules appear to be im-
perforate, it has not been satisfactorily made out to what extent, if
at all, the attached vesicles contribute to the respiratory act.
The organs for propagation are in difierent families of this class
of a different form, but still, as was stated above, have, in the two
sexes x)f the same species, exactly the same form. Hence, it
appears that the discovery of the different sexes belongs exclusively
to the latest scientific period, since formerly it was believed that all
the individuals were of the same sex, either really bisexual or solely
female '.
^ MuELLKB^s Arehiv. 1S41, 8. 5, 6.
* [This observation of Valentin is contradicted by Muelleb, Arehiv. 1850, p. i^B-]
' Wagneb first discovered the difference of sex in HoUtihwria tubulota; then Petebs,
1840, in EchxMU, Kathee in Opiwwra and Sea-stars, &o.
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134 CLASS IV.
Tet, without microscopic inyestigation, even the colour of the
organs of propagation is sufficient to point out the difference of the
sexes; the testes are distinguished by a milk-white, the avaria by
a yellowish-brown or red colour. In ninety-eight specimens of
Eckinua Petebs found that forty-three were males and fififcy-five
females, so that the two sexes are nearly equal in numbers. In
Comatula also, Muelleb found the sexes distinct ; the avaria and
testes are here situated on the ptnnulce ^ In Ophiura, on the other
hand, they lie in the disc, roimd the stomach as ten structures
composed of lobes and blind pouches, that run into a pedicle; in
the spaces between every two rays two such structures are placed
close together so as to form five pairs '. In the Star-fishes they lie
in the angles between the rays, and have the form of bunches of
beaded strings: in some species they extend into the rays; their
number is double that of the rays. On the dorsal surface, in some
species, in each inter-radial space, two spots are found, which are
perforated, sieve-like, by numerous closely arranged pores ; these
openings allow the ova or the seed to escape. In other species the
products of the genital organs, which have been poured into the
cavity of the body are probably allowed to pass out by the respira-
tory tubules on the back. In the Sea-urchins five ovaries or testes
lie on the inner surface of the shell, and fill the spaces between the
ambulacral plates. They are of an oblong flattened form, and con-
sist of numerous csecal pouches, which open into an excretory duct
running through the midst of the organ. The duct then runs finely
like a footstalk, by which the testis or the ovary is attached to the
upper surface of the shell where it opens. There are thus five such
openings, in five pentagonal calcareous plates around the anus. In
Holothurioe the ovary or testis is a bundle of branched tubes ; these
hang by their blind extremities downwards, and open above into
a single excretory duct, being fiuitened to it like a brush. The
oviduct or the efferent vessel lies along the anterior portion of the
intestinal canal, and terminates near the anterior extremity of the
body by a distinct opening on the dorsal surface. Near this canal
^ They are figured in the Ahh, der Bed, Akad. Bau des PeniaerifiUt, Taf. v.
fig. 17. 18.
* H. Bathkb BeUrdge sur vergL Anal, u, PhytioL,, BeiaenboMrkungen aut
Shandinavien, Danzig, 1841, 4to, s. 116, 117. Tab. n. ^. 3 — 7.
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EOHIKODERMS. 135
are sitoated pear-shaped vesicles, collected in some species into
bunches, which Tiedemann conceives to be male genital organs,
an opinion which falls to the ground now that the sexes are known
to be distinct Moreover they do not lead into this canal, but are
in connexion with the oesophagus. It must be noted as a remark-
able exception that in Synapta^ according to the investigations of
QuATREFAGES, a complete hermaphroditism prevails. The genital
organs have the form of long strings, whose internal sur&ce is beset
with conical structures containing Bpennatozoa^ whilst the inner*
most cavitj is filled with a pulpy substance in which the eggs are
found. These eggs, as they grow, are pressed against the conical
structures and so fertilized; and then the germ-spot, which was
before visible, disappears. As the development of the eggs pro-
ceeds, the testes which adhered to the inner wall of the string and
surrounded the eggs, are so much compressed that they become
atrophied and disappear. This periodical development is a very
remarkable physiological phsenomenon ^.
With the exception of some interesting observations of Sabs
little was known respecting the development of Echinoderms until
the last few years. [To the distinguished and unremitted labours of
MuEU-ER we are principally indebted for nearly complete informa-
tion respecting the very curious and unexpected processes which
occur in different species. Very remarkable differences are observed
according as more or less of the development is effected within the
body of the parent, or according to the locality where the embryo
is deposited on leaving the egg, or according to the different modes
in which it is destined to acquire its food. As a general rule it
may be stated that in littoral species when the embryo escapes at
an early period from the egg the series of metamorphoses is less
numerous: but that in pelagic species, where the embryo has to
seek its food by swimming on the surface, the necessity for provi-
sional organs of a complicated nature renders the changes very
1 Awn, det 9e. wU., sec s^r. xvn. 1843. Zool. pp. ^, 73. A talented obsdrrer, whom
we have already quoted when treating of Polyps, (p. 70.) SnxNSTBUP Las with miioh
acuteness endeavonred to reject Hermaphrodititm altogether, and is of opinion, that
even here Quatbxfaobs has taken oeUs of spermatozoa for eggs without the gennin»l
spot. UndertoegeUer over Serma^radiHsmM TilvaereUe % Natwen, Kjdbenhayn,
1845, 4^> PP* ^h ^4- (81BBOLD also surmises the same mistake of Quatbivagbs.) We
shall perhaps recur subsequently to Stbbnstbtjf^b opinion.
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136 CLASS IV.
remarkable. Amongst the Oph%ur<By Ophiohpis gqtiamata is vivi-
parous. The jovLng, about ten in number, are developed between
the integument and the wall of the stomach of the parent, in the
inter-radial spaces, each in its own compartment, formed hy mem-
brane extended between the wall of the body and the stomach and
suspended by a ligament attached near one of the angles of its disc.
When fully formed it passes out by one of the genital fissures \]
In Echtnaater sanffuinolerUvs the embryo according to the observa-
tions of Sass^, on its escape from the egg is of an oval form and
covered with cilia. Presently excrescences, club-shaped processes,
arise at one extremity by which it adheres to the inferior surface of
the disc of its parent, now converted by the infolding of the rays
into a brooding cavity. When the arms begin to shoot forth these
processes disappear, and feet or tentacles, few in number but pro-
portionally vBry long, serve for the creeping and adhesion of the
creature. The whole development occupies six or seven weeks.
When the clavate processes are about to disappear they are near the
edge of one of the inter-radial spaces of the disc of the Echinoderm.
Of ComatvloB it had been discovered by Thompson* that during an
early period of their life they are fixed to a stem and then resemble
Pentacrinij in other words, that the form which in Pentacrini is
permanent, is in them transitory. But their previous metamorphoses
were unknown. [BusCH has observed these changes from the egg
until the period when the embryo is about to be attached. The egg
having passed from the parent by an aperture at the side of the
pinnules, remains attached to the pinnula by an abundant mucus,
from spherical becoming oval, and the embryo may be seen rotating
within the egg by means of its general covering of cilia. When the
egg falls from the pinnula the embryo escapes : its oval form is
elongated, the straight sides assume a gently undulating contour: on
the tops of the undulations transverse bands of larger cilia are seen
in place of the general ciliated covering : the bands are at first three
in number, afterwards four, surrounding the body in parallel circles:
the longitudinal axis of the body now becomes gently curved, and a
mouth is seen on the concave surface : the bands of cilia disappear
1 Kbohn in MnxLLEB's Arc/Uv. 185 1, s. 338 — 343.
* Sabs in Wibomann's Archiv. x. b. 169.
» Thompson, Edinb, New Pkihs, Joumaly xx. p. 195.
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ECHINODERMS. 137
and the creature having lost its apparatns for motion sinks to the
bottom : tentacles in the meanwhile, five on each side of the mid*
line, have been developed, and hooks are seen at the two extremities
of the body which shew by their peculiar form that those extremities
correspond to the ends of the arms : the embryo now cup-shaped
from the increased convexity of its dorsal surface attaches itself to
the bottom by this surface from which it secretes its pedicle. The
absence of symmetry in the relation of the Echinoderm to its larva
is indicated by the stem of the Echinoderm being placed at right
angles to the axis of the larva, and the tentacles and mouth on the
opposite 9ur£eicei.
In by far the greater number of Echinoderms the embiyos pass
gradually into forms which, however remarkably they may diiSer,
are all laterally symmetrical. The axis becomes bent and on the
ventral surface (that where the mouth opens) is a depression bounded
above and below by transverse bands of cilia which are continuations
of the lateral bands which bound the dorsal surface. They all have
a complete digestive tube consisting of mouth, oesophagus, stomach,
intestine and anus. This tube is placed in the median plane, the
mouth in the ventral depression described above, and the tube
curves from it to terminate beyond the transverse band of cilia
above the mouth on the same ventral surface. Also they have all
an aquiferous system, a tube terminating externally in a dorsal pore
and internally in a sac. When Mueller observed the singular
forms of the larvae of Ophturce and Echini with their long processes
supported by slender rods of carbonate of lime he named them
Pluteus from their general resemblance to a painter's easel with his
work upon it. In AstericB and Hohthurvje the larvae have a more
flattened form, like a coat of arms with its surrounding ornaments.
The process of development in these^ different larval forms is two-
fold. In the first case the body of the Echinoderm is formed by
gemmation round the stomach of the larva, which continues to be
its stomach, and when it is formed, all that remains of the larva,
with the exception of certain structures in connexion with the aqui-
ferous system is gradually ( Ophiura and Echinvs) or simultaneously
[Bipinnaria asterigera) rejected. In the second case the symmetrical
^ [Comp. Beobach. vber Anatomie u, Enttnchdung eintger wirbeUotm Sedhiere von
I>B. W. BuscH. foL Berlin, 1851, s. 83—88.]
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138 CLABS IV.
larva with its bilateral ciliated band passes into the radial type, as
in the larva of Gomatala: it has a cylindrical form with five trans-
verse bands of cilia. From this pupa^-state the Echinoderm is
developed without the rejection of any part of the pupa. It is either
formed upon a part of the pupa, and the rest is absorbed by it
{Tonnaria of certain Aaterioe), or the entire pupa is simultaneously
metamorphosed into the Echinoderm {HolothuricB), In both these
cases, but especially the first, the axis of the Echinoderm does not
coincide with the axis of the larva, but crosses it at a large angle.
Csecal processes are formed round the stomach of the larva in con-
nexion with the vesicle of its aquiferous system: they are the
rudiments of the water-canal around the mouth of the Echinoderm
and of the vessels which proceed to the tentacles and feet. Where
the afferent water-tube meets the surface of the Echinoderm the
madrepore-plate is formed, and the portion of the tube within the
body of the Echinoderm becomes its stone- or lime-canal. In OphiuriB
the madrepore-plate has been discovered by Mueller*, its pores do
not open externally being covered by a plate of the inferior surface
of the disc. In Hohihurice it is within the abdominal cavity, in
Asterice on the dorsal surface, in Echini it coincides with one of the
genital plates. In all these cases a tube, whether it be hardened
by carbonate of lime (stone-canal) or not, runs firom the madre-
pore-sac or plate to the circular aquiferous canal surrounding the
oesophagus.]
The power of reproduction in Echinoderms is very great. Star-
fishes are firequently seen with one or more small arms or rays, that
have been formed anew in consequence of the loss of the parts.
GuETTARD and Bernard de Jtjssieu confirmed the reproduction
by many experiments'. In Hblothurice even viscera may be lost
and formed anew; and some species are propagated by spontaneous
division*.
^ MvJOAXR Archiv, 1850, p. 121.
* Comp. Mueller JHe larven u, mdamorphos. der Ophiwen u. Seetffd. Berlin, 1848,
1849 and 1852. Mueller Uher die larven u. metamorph. der ffolothurien u, Asterien,
Berlin, 185a Mueller l/eher den aUgemeinen Plan in der Enttoiehdung der RMnoder-
men. Berlin, 1853.
• BiAUMUR Mem. pour tervir d VHiet, dea Ins. VI. Preface, pp. 61, 62.
4 According to the observations of Sir J. Qrahav Dalzbll, quoted by Forbes,
Bid. of Britiah tStar-flshea, pp. 199, lOo.
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ECHINODERMS. 139
The nervous system of EcliinodermB was first described and
figured bj Tiedehann in Asteriaa auraniiaoa {AstropecUa) ^ There
is fonnd around the month a nervons ring without ganglia, whence
is giren off a fine thread for each raj, and running along it. On each
side of this thread is another, which descends into the cavity of the
body. In searurchins also and in star-fishes, in which Tiedehann
could only detect obscure traces of a nervous system, Kbohn dis-
covered a few years ago a distribution similar to that of AsteruB. In
Echinus the ring surrounds the mouth within the apparatus usually
named Aristotle's Lantern (see below in the systematic arrange-
ment) : in Hohihuria in the calcareous ring to which the longitudinal
muscles are attached. Five principal nervous stems run with the
vessels that are in connexion with the ambulacral apparatus*.
Little is known of special organs of sense in Echinoderms. In
star-fishes Ehbenbeeg discovered at the point of the rays on the
abdominal surface, a small red spot, surrounded by a ring of cal-
careous tubercles, which he considers to be an eye. In specimens
preserved in spirit the pigment disappears, and so the existence of
the spots cannot be recognized. Moreover they are wanting in
many species ^ Forbes discovered five similar spots in sea-urchins,
on the upper surface, situated upon as many pentagonal plates that
alternate with five larger plates on which the oviducts open. Both
in the star-fish and sea-urchin each of the five principal nerves runs
as far as one of these spots and ends beneath it^. But in neither of
these animals has a lenticular transparent body been discovered.
The ambidacral tubes and the feelers around the mouth may, as
highly sensitive parts, be ranked amongst the organs of touch.
To the motive apparatus of Echinoderms belong the little feet
or tentacles, abready noticed, the ambulacral tubules by means of
which the animals creep. They have muscular fibres on their walls.
In Echmu8 Valentin found in them both transverse and longi-
tudinal bundles, and radiating fibres in the suckers at their termina-
tion. He conceives that the motions of the ambulacral tubes are
^ In Mbckil'b Arehiv /. die PkyiioL i. 18x5, 8. 161, ftc. and in his often quoted
prize essay.
« Mueller's ilfc^tv. 184 1, pp. i— 13, Tab. i.
* Die Ahalepken dee rothen Meeree, s. 39 — 34, Tab. Tin. fig. 11, 12,
* Comp. YALSNTiir, op. dt. pp. xi, 100, Tab. n. fig. la, Tab. iz./. i8J3 — 190.
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140 CLASS lY.
effected hj means of tliese muscnlar fibres, and in no degree by
forcing fluid into and out of them. To how great an extent these
parts can be elongated may be judged by the length of the spines
of the shell in sea-urchins: they must have the power of extending
beyond the longest spines. Li specimens that have been kept in
spirits these tubules are found to be short and mutually entangled.
The teeth, and the curious apparatus that encloses them (Aristotle's
Lantern) have a number of proper muscles. As to the general
muscular system, that which mores the whole body, it is foxmd
especially developed in HohthuruB, Here ten muscles run the whole
lengtli of the body, beneath the skin: they are arranged in pairs,
which are separated by spaces in which transverse circular fibres are
found that cover the entire inner surface of the skin. By means of
the longitudinal muscles the body can be shortened and bent: by
the transverse fibres it is contracted transversely, and so attenuated
and elongated. The longitudinal muscles are inserted into a cal-
careous ring composed of five large and five smaller pieces, and
surrounding the commencement of the intestinal canal.
All these animals inhabit the sea. They are found in all parts
of the world: yet, most of the species only in limited regions. Since
we have only a partial acquaintance with foreign species, many
more of which will doubtless be hereafter discovered, we are not in
a condition to give a survey of the geographical distribution of the
genera. Unlimited confidence is not to be reposed in the accounts
given by some writers of the localities in which the species are
native. Of the Asterice nearly one-third of the known species are
found in the East-Indian seas, one-fifth in the seas of Europe. Of
OphiuroB, on the other hand, the European and Afirican species
are more numerous than the East-Indian. The western hemisphere
has, on the whole, fewer star-fishes than the eastern : in America
there are no species of Set/taster, as, for instance, of Culcita^ Astro--
gonium, Stellaster ; Echtnaster, on the other hand, is peculiarly
American. The species of EuryaU are chiefly found in the Arctic
and in the Indian seas. To the species which are very generally
distributed belongs that which occurs on the coast of Holland,
Asterias {Asteracanthion) ruietis. The Red sea so rich in species of
the class of Polyps, has only a comparatively small number of star-
fishes, and, in this respect is much behind the Mediterranean, which
is especially rich in species oi Astropecten. The Baltic appears to be
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ECHINODEKMS. 141
avoided by Star-fishes ^ Of Sea-urchins about one-third of the
known species are found in the East-Indian seas : here especially
are found CidarUes and ScuteUa. Of Hohthurim more species
appear to be met with in the southern Pacific than in other seas.
The western hemisphere is as much behind the eastern in respect
of Echini and HolothuritB as of Star-fishes.
^ Gomp. MuELLKB u. Tbosohsll Ud)tr die geographiickt Verbreitwng der Atleridm
in WiEOMAmr'B u. Ebichson's Arch./. Natwrgetch, x. 1843, 8. 113—130. We have
borrowed the above short notices on the geographical distribution of the Eehi/nodemuxta
from the Syttem der Atteriden of these writers, and from Lamabck Hid, not, de$ Arwn.
iam vefUbret, (compared with the specimens in the Leydan Museum).
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SYSTEMATIC
ARRANGEMENT OF ECHINODERMS.
CLASS IV.
ECHINODEKMATA.
Animals with integument coriaceous, often calcareous; with
distinct nutrient canal, freely suspended in an abdominal cavity.
Organs of circulation and generation conspicuous; sexes nearly
always distinct. Disposition of the organs most frequently quinary,
with body mostly radiate or globose, in some cylindrical. Distinct
vestiges of a nervous system, a ring for the most part surrounding
the mouth and sending off nerves radially.
Okder I. Pediculate Echinodenm.
Tentacles numerous, membraneous, contractile, terminated by a
suctorial disc, and issuing from minute apertures in the integument.
Family I. Onnmdea. Integument calcareous (external skele-
ton). Bays articulate, supplied with a central canal, absent in
some. Mostly two apertures of the nutrient canal.
The name Crindidea, given by Miller to this division of the ani-
mal kingdom, is derived from Kpivov^ a lily. At the beginning of the
last century the name seorlUi/, stone-lily was given to the Encrinus
moniliformis, or liliiformis, a remarkable petrifistction of the Mvsckd-
kaiUc Most of them are set upon a stem ; the non-pediculate {GovMir-
ttUa Lail) in the young state, according to the observations of
Thompson, are also fixed to a pedicle. The non-pediculate specieB
known to Lnnr^sus were placed by him in the genus Asterias {Aste-
ricu pectinata, Ast, muUiradiaia) ; the pediculate species in the
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ECHINODERHATA. 143
genua /m, oonfoimding the foflul Bnerintu and the FefUacrintu
capui Medusa o£ the existing creation under the name of lets
AaiericL This singular union of an Echinoderm with a genus of
Polyps had doubtless an influence upon the later arrangement of
LAMABGKy who placed Encrinus (see above, p. 80) amongst the sea-
feathers. ScHWEiGGEB and CuYiEB restored it to the proper place,
ahready assigned to it in the middle of the previous century by
GuETTAKD, who first described Pentacrinus caput Medusae This
whole fiunily belongs rather to the former period of the history of
our globe, than to the present. The species now living in our seas
are almost all non-pediculate, whilst geological investigation has made
us acquainted with numerous forms of pediculate sea-lilies. What is
now a youthful mutable form of life was then the prevalent and
permanent. The same thing may be observed in other classes also
with respect to the fossil representatives of genera that are living
at the present time.
The chief work on this fiunily is :
J. S. MiLLEB, Natmral Hidory of the Orinaidea, Bristol, 1821, 4to.'
A. Onnotds affixed,
a) SeesUe.
Holopus D'Obbigny. Calyx affixed, hollow, undivided, with
scattered tubercles. Four calcareous pentagonal parts at the upper
margin of the calyx, sustaining four pairs of articulate and pinnate
aims.
Sp. BoUypui Rangii, D'OaBiairr, Gusbin Magatin de ZooL 1837, CL x. PL 3;
firom the Caribbean Sea at Martinique. Both in the want of a stem and
the number of the anna this genua differs from the other Grinolda.
h) Pedtculata. An articulate column sustaining the calyx.
* Tesselata. Calyx non-articulate.
a) Kays or arms none.
Grenera: Sphceranites HisiNGER, PefUatrematttes Say {PerUremites
GrOLDF.), Echtnoaphcerttes Wahl., Hemicoemites Gray, Sycocy elites
V. BUCH.
FoasO genera from the Transition- and Mountain-limestone. Comp. Bbonk,
LdhoM ffeoffnottica 1835, Tab. IT. fig. i, &c, Anoording to the opinion of
some these were pediculate Eddni.
P) With rays.
1 See also W. Bugklakd, Otology amd MmenHogg, London, 1835, pp. 416— 44*.
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144 CLASS IV.
Grenera : Caryocrinites Say, Plaiycrinites Mill., Actinocrinites
Mill., Bhodocrinites Mill., Cyadiocrinites Mill., Poteriocrinites
Mill., Melocrinites Goldf.
For these fossil genera and others unnoticed we must for want
of space refer to Miller's work already cited, to Goldfuss di^
Petrefacten DeutchUmda, and other geological works. Comp. also
Goldfuss Ud>er foaaUe Crinoideen, Nov. Act. Acad. Cass. Nat
Cv/rios. XIX. I. 1839, p. 329—352, and L. v. Buch on Ca/ryocrinvs
orruUus in his work Ueher Gystideen, Berlin, 1845, 4:to.
In these and the following pediculate Crinoids, the cup-like part
at the extremity of the stem and base of the arms is named Calyx.
The bottom of this part, which is pentangular, and composed of five,
four, or sometimes three plates, is named by Miller pelvis ; Joh.
Mueller names the plates haaaUa; at the margin of these plates
are the basal-pieces of the arms, forming the uppermost part of the
calyx; there are two or three rows, and the uppermost bears the
arms. Miller gives to this part the name of scapula; the two
pieces situated below are costm. Joh. Mueller names these pieces
radiaUa (radiale primmny r. sectmdttm and r. aasUlare; his radiale
axUUvre is the scapula of the English author). In those now
named Orirmdea tesseUxta these parts are joined together without
articulation. The fossil species of this division are found in the
transition-limestone and the grey-wacke.
* * ArtiGulaUi. The rays free directly from the pelvis of the
calyx, the first radial conjoined to the second, and the second
to the third by articulation.
Apiocrtnites Mill. Column incrassated towards the calyx,
pyriform.
Sp. Apiocrinites rotundua Mill. Tab. i— vn. Apiocrin. Parhintonii Bronn,
LethoM Tab. xvn. fig. 15, (Millkb*8 figure) fossil from the oolite forma-
tion, like other species of this genus.
Encrinus GuETTARD (in part). Column round, not incrassated
towards the top.
Sp. jEncrinu* UUiformU Lam., EncrinUa monUiformU Milueb, pp. 37 — ^44,
Tab. I— m. ; Ellis OoraU. Tab. 37, fig. k, &c. One of the most character-
istic fossils of the MiuekeOsalk. The head, on account of the numerous
articulations of the arms that lie side by side, resembles an ear of Turkish
wheat {Zea Maps) ; the joints of the stem, sometimes found in inoradible
numbers, changed into calcareous spar, arja named Troch4te$\
1 QUENSTBDT {Ud)er die Enkrimien des MutehdhaUcs, Wibomann's ^rdUv. 1835, "•
s. 123—328, Taf. IV.) describes a specieB with different division of the arms, under the
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ECHINODERMATA. 145
PerUacrinus MiLLBB. Column not incrassated towal'ds the top^
pentagonal with joints having a pent^petalons mark. Cirri adher*
ing to the column in whorls.
The fossil species of this genus helong to the oolite period, as
ex. gr, PevUacrinua hriaretu Miller, pp. 56 — 58, Tab. l il Guy. B.
Anu ed. illiLgtr,, ZoopL PL 7, fig. 1, from the lias-schist
The living species of this genus, FerUcuyrinua Capui MeduacB^
is found in the Caribbean Sea, and was first described by Guettabd
Mem. deVAcad. Ray. des Se. 1755 (Paris, 1 761) pp. 224, Ac PI. &— 10 ;
another, more mutilated specimen, foimd on the coast of Barbadoes,
was described bjELUS in 1761, FhU. Transact. toL 52, p. 357, PL xiii.
Specimens of this species are rare in Museums, seven only, as fiir
as is known, existing in the different Cabinets. The most complete
description was given not long ago by J. Mueller ( Ueber den Bau
des Pentacrinus CapiU Medvsoi, Berlin, 1843, folio). The stem of
Fentacrinus has no musdes, but is merely passively motile or flexible
by means of fibrous bundles and an elastic tissue between the jointa
The arms and pinntUce are moveable by muscles without transverse
stripes ; these muscles are situated on the side corresponding to the
mouth, and can only flex the parts : extension, or motion outwards,
seems to be effected merely by the elasticity of the parts. The
growth of the joints of the stem occurs in the part at the top
nearest to the calyx, which corresponds to what is observed in the
growth of the joints in worms and entozoct.
B. Free Crinmda.
a) Tesaelate.
Marsupites Mantell. Marsupiocrinitea Blatnv, Calyx of
parts calcareous pentagonal striated ; arms ?
Sp. Marsup. omatut Mill. Crinoid p. 134, with figure, Bbonn, Lelh. Tab.
XXiz. f. 13 ; (Tab. zxxrv. fig. 9, with oonjectaral restoration of the arms
alter the figure of Mamtkll) ; fossil from the chalk-period.
b) Articulate.
Comatula Lam, {Aledo Leach, Muell. Cirri dorsal articu-
late, around a pentagonal disc. Kadials mostly without basals
name of Encrinita Schottheimu of which H. Y. Mbtbb has formed a new genus,
Ckdoerinva. See the paper of the latter ; Tsocrimta und Chdocrinus, Zwei neue T^pen
u. i, w., Mtaeum Senkenberffian., n. p. 249.
VOL. I. 10
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146 CLASS IV.
composing the calyx. Bays pinnate ; pinnae sulcate, the farrows
confluent with the longitudinal farrow of the rajs ; tentacula situ-
ated in the furrows. Mouth and anus on the ventral side ; mouth
central in the bottom of the calyx ; anus lateral, tabular.
See J. MuELLEB, V^)eT die GaUungen tmd Arten der ConuUtden^
Emchson Archiv Jur NaJtwrgtscL 1841, a 139 — 148, and Nem
BeUrdge zwr Kewntniaa der Arten der ComcUtUeny ibid. 1843,
a 131—136.
* Jtay$ of Calyx bifid {ten-raffed).
8p. Comattda rosacea, AUdo ewropaa Lkaoh, Fobbxs Brit, Sta/rf. p. 5. To
this species belong the obsenrations of Thompson spoken of above^
(ppu 136 and 143). Memoir on the PeiOacrinm eurojxgut, Cork, 1827.
Oomat, eairiiuOa Lam., GuAb. Iconogr, Zooph. PL i. fig. a.
* * Bays of Cfalyx mfUt\fid (mamy'Tayed).
Sp. Comatula multifida Musll., Com. muUiradiata Lam. ftc
The species, which like Pentacrinus have & pelvis, form the genus Comaster
AoASS., MuELL. Here belongs Comattda mvUiradiata Gold, (not Lam.)
Fossil species of this genus are foimd in lithographic stone.
Family IL Astendea. Body depressed, free (not pediculate),
multangular or radiate, with integument coriaceous or calcareous.
Row of joints calcareous internal, running along the middle of
the rays and taking their origin from the mouth. Mouth central,
inferior ; anus dorsal or none.
A capital work on this division was published some years back,
System der Asteriden von J. Mueller und F. H. Tboschell, mit 12
Kupfertafeln, Braunschweig, 1842, 4to. As plates for this and the
preceding family may be recommended : J. H. Linckii De SteUis
marinia Liber singtdariSy Lipsise, 1733, foL
Phalanx I. Ophiurce. Disc distinct from the arms ; with arms
non-sulcate. AnuJg none.
Eurycde Lam. {Gorgonocephalus Leach). Arms prehensile,
contortile towards the mouth, not scutate, sub-rotund, flattish be-
neath. Disc tumid, sub-globose, with five obtuse angles.
According to the division of the arms the species of this tribe
are arranged in three different genera by Mueller and Teoschell.
They are undivided in AsteronyXy divided dichotomoualy and only
towards the end in Trichaster Agass., divided from the baae, first
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ECHINODERMATA. 147
dichotomously and afterwards into many branches, in Antrophyton
LiKCK. To the last division belongs :
Sp. EwryaU verrueosum Lam., Adentu Caput Medusa L. (in part) RuiCPHinSy
Amb. JRarUeithamer Tab. XYI. Guy. R Ani., Sdit. iOus., Zooph. PL 5, from
the Indian Sea ; a yery Bunilar species is found in the North Seas, and
distinguished by MuiLLSB and Tboschsll as Attrophyton JAnekU; LnrcK
de Stell. mar. Tab. 29, fig. 48. These Meduta-Keadt belong to the most
singular and beautifid forms of radiate animals. Yid. FoRBis Br, Star-
M^f pp. 67—70.
Ophiura Lam. Arms five, undivided, serving for creeping, scu-
tate, articulate. Disc plane, with two or four genital fissures in
each interbrachial area on the ventral side.
The name Ophiuray from o^ic, nrpent, and wpdy tail, denotes very
appropriately the form of the arms by which these Seorskbrs are
distinguished, and which are often so long as to exceed five or six
times (nay in Ophinra longipeda even twenty times) the diameter
of the disc.
Sub-genera : Ophiocoma Agass., Ophiolepis, Ophia/rctchna, Ophior
eantha, Ophiomastix, Ophvomyxa, 0phio8colex, OphiothriXy Ophio-
cnemisy Ophioclerma, Muell. and Tbosch.
Sp. Ophiura textwrata Lam., Atleriaa ophiura L. (in part), Ophiol^ni cUiala
MuxLL. and Tsosoh., Lutck de Stdl. mar. Tab. 11. ^g, 4, Encydop. PL 113,
fig. %, 3. FOBBXS Briti$h Starf. p. 2a, ftc. in the Mediterranean, the
North Sea, Ac.
Phalanx II. Aatence, Body depressed, angulate or stellate
the angles being produced, with tentaculiferous furrows below, ex-
tending as far as the point of the angles. Anus dorsal in most,
surrounded by a mound of calcareous papillae.
Asterta Lam. (Most are species firom the genus Astertas L.)
The Seor-gtars, The form is very various, so that in some species
the entire body seems to consist only of anns, ex^ffr.in Ophidicuter,
in others only of a pentagonal disc. But the arms are never
sharply separate from the disc as in the OphiwraSy but are an
immediate continuation of it. In moat of the species there are ^yq
rays, however in these sometimes four or six occur as exceptions ;
six arms as the normal number are found in Astertas gdcUinoaa, in
EchiruMier eridanellay six or seven in Asteriscua Diesingiiy seven to
nine in the sub-genus Luidiay eight to ten, mostly nine, in Solaster
endeca, eleven to fourteen, generally twelve, in Solaster papposus,
twelve or thirteen in Astericts aster , fift,een in Agteriscus rasacetiSy
10—2
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148 CLASS IV.
fourteen to one-and-twenty in EcMnaster aaUmB; finally, in Aeteruu
hdiaaUhv>8 the rays axe found up to thirty and more. The greater
the number that any species possesses, the less is it constant. On the
dorsal surQice is placed a calcareous star-formed plate between two
rays of the disc (yerraca doraiy tubercyle madreparifarmef Mctdrepore-
plate), which in Ophiwra is wanting, and in Eurycde lies on the
oral surfaca Aoassiz who endeavours, with great acuteness, to prove
a lateral symmetry in the Echinodemuxta, considers the ray that is
opposite to this plate to be the first ray of the body. A tortuous
tube proceeds from this dorsal plate downwards as &r as the mouth,
and is filled internally with a calcareous matter (see above, p. 130).
This tube wm named by TiSDSic akn {Anat. der SoehrenSoloth.f &c. 53,
54) Stone-oamU or Sand-eanal; it terminatefl, becoming narrower, in the
circular yeesel suirounding the mouth and filled with watery fluid ; see
above, p. 131. Siebold has closely investigated the calcareous balk,
consisting of several joints and internally hollow, which occupies this
canal and described its complicated structure ; Muelleb*s Archiv. 1836,
s. 291, &c. [Also Shabpkt, in Todd's Cydop. of Anat. arid Phyi. n.
pp. 35, ^., describes in the interior of the jointed calcareous tube a lamina
attached longitudinally, which passes inwardly a certain way and then
separates into two which are rolled in opposite directions, something after
the manner of the inferior turbinated bone of the ox.]
The Sea-stars can bend their rays towards each other, which is
serviceable in moving through narrow fissures and between stones.
They do not swim, but creep by means of their tentacles with
mouth downwards. They feed principally upon Molluscs. Though
the genus Asteriaa of Lamabck, by the exclusion of Comatula^
Ophiwra and EwryaJlSy be much more narrowly limited than the
same genus in the Sf/stema NaJtM/rcR of Linn^us, still the species are
too numerous and the forms too various not to be regarded rather
as a natural group which ought to be divided into several genera
or 8ub-genera^ This has been done by Linck, and more lately
especially by Aoassiz and Mueller and Tboschbll, to whose works
we refer. The primary division of the group by Mueller and
Troschell is founded on the Tentacles, which in most of them are
placed in two rows in every furrow, but in others in four rows.
A. Ventral fwrrowBy with two rows of tentacles.
* Amis none.
Astropecten LiNCK, {Aatropecten and Ctenodiacua Muell, and
Trosoh.), Luidia Forbes.
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ECHINODERHATA. 149
8p. AMtnptdm attrmUiaeui, AtUriat aranciaea L., LurcK, dc SleU, mar.
Tab. 5, fig. 6, Tah. 6, fig. 6, TiKDMiANir Anat. Tab. 5, 6, Forbu^. Starf.
p. 130 ; in the Mediterranean and Atlantic Ocean.
** Antu ceniral, or wh-cenlral, dorsoL
a) Body disciform, toith short rays.
Asteropsis Mu£ll. and Tbobch., Stelkuter Grat, Ganiodiscus
MuELL. and Tbosch., Astrogomu/m Muell. and Tbosch., Asieriscus
MuELL. and Tbosch., Pteraster Muelu and Tbosch., Ctdeita
AOAfiStZ.
^. Oulciia dtKofdea, Atterieu cUacoidea Lam., Sohmidsl in Naiiwrforsck. xyl
1 781, Tab. I., BLAimr. A(tmol. Tab. 13, fl i ; Atteriscw palmUpea, Aderiat
mmbranaeea BiTz, Gmu.., Likck, de StdL nuir. Tab. i, fig. ^, BLAOrv.,
AainoL Tab. 93, fig. 7, Forbes Brit. SUarf. pp. 116, &c.
6) jB(xfy steUaUy unth rays distinct, surpassing the disc three
times or more,
Archaster Muell. and Tbosch., Oreaster Muell. and Tbosch.,
Scytaster Muell. and Trosch., Ophidiaster Aoass., Chcetaster
Muell. and Trosch.^ Solaster Forbes, Schinaster Muell. and
Trosch., (and Echinites in Erigh& Archiv, 1844).
Sp. SdUuUr pappotus, Atteria$ pappota L., LlircK de SteU, mar. Tab. 39,
^I^- S4f %• 54> FOBB. Brit. Starf. p. iii, in the Atlantic, on the
coast of England and Scotland, &c. ; when fresh of a beautiful red colour
above, of a yellowish brown below ; Echinatter aolaria, EchinUe* tolarit,
AtUriou EehifUteB "Lam., Naturf orach. 3JCYII. 1793, Tab. i, u. from the East
Indian Seas, 9ui.
B. Ventral furrows with four rows of tentacles. Body with distinct
rays, with integument coriaceous, aculeate. Anus dorsal, sub^
central.
Asterias Gray, (species of the genus Asterias L., Asteracan"
ihian Muell. and Trosch.)
Sp. Aateritu rubens L., Linok de SteU. mar. Tab. 36, fig. 61, Bastbb
Natuurk. Uitap. i. Tab. xii. fig. 1—4, Fobbbs Brit. SUtff. p. 83 : the com-
mon Sea-star, Gross-fish; so common on some coasts as to be used for
manuring the land in the neighbourhood. Aatericu hdda/nihu* Lax.,
Enqfdop. PL 108, 109, Blainy. AdinoL Tab. 23, fig. 5, in the Pacific and
at Chili ; one of the most remarkable and most beautiful species, &c.^
^ For a knowledge of the numerous species of A^eridea comp. also J. E. Gbat,
Sjfnopeit of the Genera and Species of the cUua Bypoetoma (Atteriae L.) in the Annate
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150 CLASS IV.
Family III, Echinidea. Body sub-globose or depressed with-
out radiant lobes. Mouth and anus distinct. Mouth inferior. Inte-
gument calcareous, beset with moveable spines.
Seor Urchins. Compare on this family (besides the Monographiea
dEcJdnodermes of Agassiz cited above) Jac. Theod. Klein Natvr
rails dispositio EchmoderrnoUum cum tab. Gedani 1734, 4:to. Ordre
naturel des Oursins de Mer et fossUes par M. Theodore Klein,
Paris 1754, 8vo. av. ^g. (Many of Klein's figures are copied in the
Encyclopedia metliodiqtie, Vers,) M Van Phelsum, Erie/ an C.
NozEHAN over de gewelv-slekken ov Zee-^gelen. Met 3 pL, Rotter-
dam, 1774, 8vo. Ch. Desmoulins, Etudes sv/r lea EchinideSy
Bordeaux, 1835—1837, 8vo.
The shell of these animals consists of an arrangement of plates
having a pent- or hexangular form. They compose ten girdles, each
made up of two row^ of such plate& Five of the girdles, commonly
narrower than the others, have two rows of small apertures and
alternate with thesa The rows of apertures are named artihvlaGra :
they either entirely surroimd the periphery {ambuiUtcra perfecta), or
are found only on the uppermost part, resembling in their arrange-
ment a star or five-petalled flower {cmdyidacra circwmscriptcty By
these apertures the tentacles or ambvlacrail tubes are exserted,
of which we have treated above. The Sea-Urchins effect their
movements by means of these tubes', they appear to have a
great power of elongation, to be able to stretch fiirther than the
extremities of the rigid spines, which in certain species are some
inches in lengtL Around the anus are five larger apertures (in
some genera only four) which are the outlets of the oviducts or
efferent vessels; they are situated in as many pentagonal calcareous
plates, with the point directed outwards, of which one, larger than
the rest and of a different structure, corresponds to the calcareous
plate {madrepore-plate) of the Searstars, as Basteb had previously
of Natural Eittory vi. 1841, pp. 175 — 184, pp. 475—290. Want of space prevents
our noticing the numerous generic names of Gbat ; some genera agree with those
of Mueller and Tbobchxll, of which a more detailed notice by Agassiz may
be found in the prefiuse to the second number of his Monographiei dCEchmodermes,
pp. 5, 6.
^ GANDOLraB QikdUi tatU leijambet dee Owrtmt t Mim. de VAoad. dee Se. de Parte
pour 1709, ffietoire, p. 33. With his observations those of Bastsr, Tiedbmann and
others completely agree ; AOASSiz, who at one time considered the spines to be organs
of motion, and doubted that such was the office of the ambulacral tubes, has since
renounced that opinion.
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ECHINODEBMATA. 151
obserred'. Between these five genital piatea, lie five smaller (the
octUa/r plaies), also pCTitagonal but with the point turned inwards, so
that the plates are wedged into the spaces between the points of
the first fiva There are still other smaller plates, variable in
number, that lie within the ring formed by these ten, and immedi-
ately surround the anus.
The chemical composition of the calcareous shell is remarkable
on account of the extremely small proportion of organic matter ; it
consists almost entirely of carbonate of lime. The growth of the
shell ia effected by enlaigement of the plates, and by addition to
their number upwards ; hence younger individuals are flatter than
older ones, and the form of itself alone becomes a deceptive criterion
in determining speciea
The moveable spines, of which the number increases with the
age, have at their base a small cavity, by which, as by an articular
surface, they are connected with the tubercle of the shelL These
tubercles are placed in rows, like the apertures of the ambulacra,
and are found both on the ambulacral and on the interambulacral
fields.
The mouth is provided vrith five teeth, whose points are sharp
and hard, and meet in a pentagon at the inferior aperture. These
teeth are long rods, which become soft and transparent inwards.
They perforate five triangular pyramids, which by their mutual
arrangement form a conical apparatus with the broad base facing
inwards, and to which base still other small calcareous pieces are
united. This very complicated apparatus, with its provision of
muscles, bears the singular name of Aristotle's l<miem\
Sea-urchins cannot swim, but only creep along the bottom of the
sea. There are species that sometimes live in cavities which they
have formed in the rocks'.
The food of the Searurchin consists, according to the microscopic
investigation of the excretions in JEehinus Iwidtu by Yalentik,
^ Naluurik, UUsp, i. bL 132.
* ThiB apparatus is described m detail by Cuvns Zepons cPAnai. wmpa/rie, Paris
1805, m. pp. 3«9-~33S> »nd ae ^t. Paris 1837, vi. pp. 377— 382 ; oomp. also the
woriLB of Tudbiiasn and Yalxntih abready cited, and B. Joins, OvUine of the Anim.
Kingd. pp. 166 — 169.
* Eehinns Utkophagui of Lxaoh, which does not appear to diflfer from B^inu*
Umdm, is often thus found on the western coast of Irdaod ; E. T. Bxnkitt Litm,
Tran$acfian9, xv. 1817. pp. 74 — 77.
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152 CLASS IV.
principellj, if not exclusively, of marine plants (Ftid, Confervas) ;
according to others it would seem to live on Molluscs, but the
fragments of shell, often found with considerable quantity of sand
in the intestinal canal, may have been contained in the sea-water
swallowed, and need not by any means to be looked on as the
remains of shell-fish that had been consumed.
Petrified shells of Searurchins are found in great numbers in
secondary strata, particularly in the chalk-formation, the interior
being usually filled with silicious earth.
* Arms eccefrUric superior or inferior,
a) MofuJth eccentric, {Genital pores 4.) Sptxkmgoidea Agass.
Spatangus Klein, Lam, Ambulacra circumscript, five or only
four, the odd one (the anterior) being either little distinct or
wanting. Test ovate or cordate, often at the fore-part famished
with a furrow proceeding from the summit.
Genera Holaster, Heinipneustes, Micraster, SpcOangus, Amphi-
detus, Brissus, Schizdster Agassiz.
Some species of this division are fossil, and occur especially
in the chalk-formation, others in tertiary deposits. Amongst the
species now living, which chiefly belong to the genus Brissus Agasb.
we note :
Spatang. verUricotxu, RUMPH. Amh, Jtariteitk. Tab. xiv. No. i ; this
foreign species attains a very large size. To the proper genus SpcUanffug
AoABS. belongs i^pat. purpureus, Blatmv., AcHnol. PI. xiv., Fobbbs Brit,
Staff, p. 1 83, in the North Sea and Mediterranean. The form is heart-
shaped. Of this species Milne Edwabds has ^ven an anatomical figure
in CuvTEB R. Anim. 4d, iUustrie, Zoophytes, PL xi. bis. See also some
notices on the Anatomy of Spatangus in SoHWBiaOBB^s Handh. der Natur-
ffeschickie der skelelU. ungeglied, Tkiere, s. 538, 539.
The mouth is in this genus without teeth. There are only four
ovaria or testes present, as also only four pori gemtales. Philippi
not long ago described three American species in which only three
genital pores existed, and which he united under the name of
Tripylvs, See Erichson's Archiv f, NaJtwrgescLy 1845, s. 344,
&C. Tab. XI.
Arumohytes Lam. (exclusive of some species), Agass. Ambu-
lacra radiating and diverging from the vertex to the margin, not
interrupted. Body irregular, oval or conoid ; test without a furrow
to the anterior ambulacrum.
Sp. Ananckytes ovata Guv. and Bbonon. Ltscr, gM, des environs de Paris in
CuTiBB Reck, s. I ossem. foss, 11. 2 PI. v. fig. 7, Bbonn LeUL giogn. Tab. 29,
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ECHINODERMATA. 153
fig. 28. Ananck. hemupkofiiea Cur. and Bbovoit. 1. 1. fig. 8, Ac. All th«
species of this genus are fossil and belong to the chalk-formation. Hie
casts of the internal cayity in Fire-stone {nuclett$, numU inUriewt) shew
minute eleyated points in place of the ambulacral pores ; such a moule is
the AnoHchyUt puMidosa Lam*
Dysaster Agajss. {CoUyrUes Desmoul. in part, species of Spa-
tangus and Ananchytes Auctor.) Test rotund or cordate, with
ambulacra not interrupted, mouth sometimes almost central not
transverse. Ambulacra above radiating firom a double vertex, the
three anterior confluent in the central part of the test above, the
posterior ones above the anus.
Sp. Ihpuuter eofiinabiu, Spatangtu earinatui Bbonn Letk. giogi^. Tab. ZTIL
fig. 7, in the Jur»-foimation like most of the species of this ganns. Comp.
DssoB in the 3d number of Aoasbiz Monogr. d'Echinoderm., who describes
QO species, of which 17 are from the Jura-fonnation and 3 from the chalk-
period. No species either of this genus or of Ananchyta has been
discoyered belonging to the present animal world; AOASaiz places this
genus in the following division, that of the Clypecutr&idea; its place cannot
be determined with preosion, before it is known whether teeth be present ;
still we are of opinion that it ought not to be separated frt>m Ananekj/tei,
but forms the transition between that genus and the folloiiring division.
b) Month central or stdhcentral. {Clypeastraidea Aqass.)
Spines lUUe evolved^ sometimes very smcUl, resembling hairs.
Galerites Lam. Body conoid or sub-oval, plane beneath. Am-
bulacra in ten fiirrows in pairs from the vertex to the base, radiately
inscribed, not interrupted. Anus in the margin, or beneath and
near the margin. Genital pores four.
Genera: Glypeus E1i£in, Agassiz, Discmdea EIlein, Agassiz,
Galerites AxiABA.
8p. GaleriteB albo-galeru$ "Lam., Guv. M. Ani., idit. tS. Zooph, PI. xiv. fig. 4,
Bbonk Leih. ff^ogn. Tab. xux. fig. 18, Aoass. Monogr, i* Livr. Tab. i.
f. 4 — II, Tab. xin. f. 7, &c. This genus consists of fossil species of the
Jura and especially of the chalk-period. Comp. on this and some other
genera DxsoB in the 3d number of AOAsaiz Monographiu d^Eehinod.
already referred to,
Pygaster Agass. Body depressed, with outline orbicular or
sub-angular, beneath concave. Anus large, superior, situated in
a broad furrow. Grenital pores four.
All the species of this genua are fossil; most of them bebng to the
Jura-formation.
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154 CLASS lY*
NucUoUtes Lam. Body ovate or cordate, convex, sub-iiregolar.
Ambulacra five, extending radially from the vertex to the base, not
interrupted. Anus superior, near the margin.
All the species are fossil, partly from the seoondaxy, partly frt>m the
tertiary formation.
Casstdulus Lam. Body irregular, elliptic, ovate, or subcordate.
Ambulacra five, circumscript, resembling a pentapetalous flower.
Anus superior, near the margin.
This genus also consists of fossil species from different formations ; ex.
gr. from the chalk formation is Ctusidulus lapu eaneri Lam., Fadjaa db
St Foin>, Beach, van den St. Pidenberg, PL 30, fig. i ; Bbonh Ldkon
giogn. Tab. xxix. fig. 10.
Fibularia Lam. Echinoct/amus Leske. Body sub-globose, with
outline oval or orbicular. Ambulacra five short, circumscript. Anus
inferior near the mouth, or median between the mouth and the
posterior margin.
8p. Fibularia ovuLum Lam., GruiRnr Iconogr. Zooph. PL in. fig. 5, EcMnocy-
amuapunllui Mukll., Fobbes Brit. Starf. pp. 175, &c. From the Atlantic
Ocean, of the size of a pea. Amongst the foreign species, aooording to
Yak Phelsuh, mostly American, ^we note Fibularia cranhlarii, V.
Phblbum Gewehdekken Tab. i. f. 16 — 35. Also in the chalk-formation
and in tertiary deposits species of this genns occur.
Echirumeus Leske, Lam. Body sub-ovate or orbicular, sub-
depressed. Ambulacra inscribed radially in ten fiirrows firom the
summit to the base, not interrupted. Anus inferior, oblong, near
the mouth. Genital pores four.
Sp. Eckinoneut semilunarii Lam., QviR. Iconogr. Zoopk. PL m. fig. 2, Cuv.
B. Anim. id. ill., Zoophgtes, PL xiv. fig. x, from the West Indian Sea, ftc.
Of this genus no fossil species are known.
Clypeaster Lam. Body oval or sub-angular. Ambulacra cir-
cumscript radiating from the summit, resembling a pentapetalous
flower. Anus inferior, near the margin, or in it. Grenital pores
mostly five.
Sp. ClgpeatUr rotaeeua, Echinua roioeeui, L. Klbin Diapot. Bckinod. Tah.
xvn. fig. A, Tab. xviii. fig. B, BLAiKyn.LB AcUnol. PL xvn. ; from the
Indian and Japan Seas. The shell is thick, and parted intemaUy by
pillan; oomp. the figures in Klbik L L Tab. XXTOL ZZIZ. FoeaU
species also from the tertiary formations are known.
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ECHINODEBMATA. 155
EcMnoiampaM Okat, AoAsa Ambulacra cutnunacript, very wide
at the summit^ nannower towards the margin, not oonjoined Duo
submarginate forward& Genital pores four.
^. Clypeatier eecenirieus, Echinolampaa Kleinii BsONV Leth. g^ogn. Tab.
XXXYI. fig. lo : fossil from the tertiary formations, &c.
Scutella Lah. Body flattened, with margin thin, sab-acute.
Ambulacra short, circumscript, resembling a pentapetalous flower.
Anus inferior between the mouth and the margin. Genital pores
four or five.
From this genus AoASSiz at first separated the sub-genus Echinr
anrcuihnius {Prodrome cTune Monogr. des Radiarea, <!bc.,) afterwards
he divided the genus ScuUUa of Lamarck into thirteen genera
{Monogr, dEchinod,, 2e Livraie, lea Scutellea) ; RotvZa Klein,
Agass., Runa Aoass., MellUa Kleik, Encope Agass., Lobophara
Agass., Amphiope AoAsa, Scutella Agass., Echinarachmtu Y.
Phel&, Arachnoides Kuoir, ScuUllina Agass., Lagannm Kleik,
Echmocyamfue Y. Phels., MotUinia Agass.
Sp. Scutdla teaforis Lam., MeUUa kexapara Agass., GirfRiK leonogr, Zooph,
PL 3, fig. 4, Gov. R. Ant, €d, 01., ZoopK PL 15, fig. i. Aoabs. Monogr,
Livr. a, PL iv. fig. 4—7, PL nr». fig. 11, i« ; from the shores of the Mo-
lucca Islands, &c. — Scutdla denUUa Lam., Jtottda SumphU Kuuir, Agass.,
BuMFH. Amb. Sariiekk, Tab. ziv. fig. i, &c. (Linnjeus comprehended all
the spedes known to him under the name of Sekinua orbictdarU), Except
a very small number from the chalk-formation, all the fossil species of this
genus are from the tertiary formations.
** Atms superior centred, opposite to the mouth. Test regular,
AmbuUusray perfect. Genital pores, Jive, {Cidaridea Gray, Agabs.)
Echinus Lam. (spec, of genus Echinus L.) Ambulacra wider
towards the middle of the test, divergent, contracted at the summit
and towards the mouth. Spines placed upon imperforate tubercles,
sometimes very long.
Sp. Echinus Uvidua Lam. {Echin, sexaHlts Tiedxm. 1. L), YALEirriir in Agass.
Monogr. SEchinod., Liv. 4, PL i, £abtbb Natuurh, Uitip. i. Tab. xi.
fig. I — 8, F0BBS8 BrU. Starf. p. 167. In the Mediterranean and North
Sea, ftc. For the synonomy of the European species of Echinus oomp.
Agassiz in the preface to the fourth number of his Mcnograpkia.
Arbacia Gray, Agass., Echinome^ra Bbmjts, Gka.t, Agass.
Sp. Eehmui mamiOatui L., Eekinomdra mamSlata. Bumph. Anib. JRariteitk.
Tab. xm. £g. x, 9, Cut. R, Ani, 4dU. iOuttr., Zooph, PL xin. fig. i, East
Indian Sea, &c.
Salenia Gray, Agass. {Goniopygus, PeUaster, Goniophorus Agass.)
Gomp. Aoassiz, Monogr. cPEckmod, Livr. r.
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156 CLASS IV.
Cidaris Klein, Cidaritea Lam. {Gidarisy Biadema, Astropyga
Grat, Agass.) Ambulacra parallel. Tubercles sustaining tibe
spines not perforate, often remarkable for their size.
Sp. Cidaris verticUlaia, Gu^B. Iconogr. Zooph. PL 3, fig. i ; in the Indian
Sea, on the coasts of Timor, &c. Of this genus many fossil species also
are known.
Family IV. Holoihuridea, Body free, mostly cylindrical,
covered with a coriaceous skin, furnished with calcareous particles
scattered, reticulate. Mouth surrounded by retractile tentacles.
Anus terminal, opposite to the mouth (Grenus Hblothuria L. exclu-
sive of several species).
Comp. on this family :
G. J. Jaeger De ffolothv/riia, Dissertatio inaug, Turici, 1833, 4to.
cum tab. J. F. Brandt Prodrorrvus descriptionis animalivm ah
H. Mertensio in arhis terrarum circumnavigattone observaiorum^
Fasc. I. Petropoli, 1835, 4to. pp. 42 — 62, Grubb Actinien, Echino-
dermen und Wurmer des Adriatiachen und MiUlemeers, Konigsb.
1840, 4to. pp 33 — 42. There are many figures of European
Uolothv/ncB in the Zoologia danica and of foreign species in Lesson
Centurie Zoologique, Paris 1830, 8vo.
We have spoken above (p. 140) of the caloareous ring which may
be considered as a vestige of a skeleton, and which serves for the
attachment of the longitudinal musdes. The pieces of which this
ring consists are toothed above, but they do not discharge the office
of teeth, the food, as far as it undergoes separation and mastication
previous to deglutition, being thus effected by the callous skin
surrounding the mouth alone. HoloUmrias feed upon conchifera and
other marine animals ; Tiedemann frequently found shells entire and
tmiujured in the intestinal canal of Holothwria tfubuloaay so that the
molluscs appeared to have been dissolved in the shell and digested.
The bits of shell and the other matters unfit for use and undigested
are rejected from the doaca with the water in expiration. We
have indicated above the chief particulars respecting the internal
structure.
See, besides, Tudbmann 1. 1., also the beautiful engraving from a prepara-
tion by HuiTFBB in Oataiogue of the Phytiolog, Series qf Qmpar. Anat,
contained in the Museum of the Eoyol College of Surgeons, Vol. L London,
1833, PL m. pp. «50~«54.
The numerous species of this fiunily are separated according
to the form of the Tentacles (Laicabck, Grube), the position of the
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ECHINODERMATA. 157
feet or ambulacral tubes (Cuvier), or the union of both these
characters (Brandt) into many genera. The arrangement of
Brandt is drcnmstantialy and here and there rather artificial than
natural Regard ought also to be had to the Habitua, If the
position of the Tentacles be made the chief point in the arrange-
ment, animals will be separated which in their habitui are nearly
allied. These tentacles are either branched like a tree, fingered, or
shield-like, (jpeltaU), i. e. they consist of a pedicle with an expansion
which is disdfonn, often indented or arborescent. Their number is
usuaUy ten, or twice ten. Where eight tentacles seem to be present,
sometimes on closer inspection two others, smaller and undeveloped,
are fotind, (the sub-genera Anaperug and Colochirus of Troschell').
In some no special internal respiratory organs are found (the genus
OncUdbes of Brandt ; in most the arborescent respiratory organ
exists which we described above in ffoloihurta ttibulotcL
Peniada GrOLDF. {Clcidodactyla, Dactyhta, Onctnolabes^ Aspido^
chir Brandt). Feet equal, disposed in five, or, more rarely, in six
rows longitudinal, parallel. Body cylindrical or quinquelateral,
attenuated at both ends. Tentacles mostly ten, sometimes twelve
or fifteen.
a) Tentades peltate,
Aapidochir Bbakixp. A genua anknown to me. 8p. Aspidochir
MertensU, in the island Sitcha (Tentacles la).
b) TevOacUa ramose.
Cladodactyla BRASvr (Tentacles lo).
Sp. Pentada Pentaetet, ffoloth. pentadet Mdkll. Zool. dan. Tab. 31, fig. 8,
FoBBBS BrU. Staff, p. 413 ; PerUacta frondosa, Hdloth. frondoM L., ffoloth.
pentaetes Abildg. Zool. dan. Tab. 108, figs, i, 2, Tab. 114. Fobb. £r. Star/.
p. 209 ; in the European seas. Pentada doliolum, Actinia doliolum Pall.
Spie. Zool. Tab. xi. figs. 10— i a. Cur. JL Anim. id. ilL Zooph. PI. 20,
fig. 4. These species are called^ from their external resemblance, iSSea-
Cuewmiben.
c) Teni€kde8 digiitUe or pinnate.
Dadplota BBAinyi.
Sp. Pentada pellueida, Boloth. pdlucida, Zool. dan. Tab. 135, fig. i.
{Note. — ^Here also is to be referred the genus Ocnue Fobbks, Brit. Starf.
P- ««9-)
* Neue ffolathunengattungen in Ebichboh's Archiv f, Natwrgeach. 1846, s.
60—66.
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158 CLASS IV.
Oncinolahea Bbandt. Body covered all over wiih reonrved
booklets. (Differs from FerUacta hj defect of respiratory organs.)
Thyone Oken, MuUeria Flem. (not Jaeg.) Feet equal, nume-
rous, scattered through the whole body. Body cylindrical.
a) Tentadea ramose, often unequal {PhyUoporus Gbubb, Anapenu
Tboboh., OrcttZa Trosoh.)
Sp. Thyone papiUoaa, Hoi. paptUoia, Zool. dan. Tab. xo8^ fig. 5, FoBB. BrU,
Sta/rf, p. a'33 ; in the North Sea, &a
h) Tentada pdUUe (Sporadipus Bbandt).
8p. Sporadipue Ualanenni Bb. &c.
Holothurta noK (Species of genus Holothurta L.) Feet of two-
fold structure and figure, some cylindrical, dilated at the tip,
usually occurring in the abdomen only, others situated on the back,
not dilated at the tip, emerging from warts on the back. Body
cylindrical, or flattened in the abdomen.
a) TerUacUa peltate or umheUate usually 10 [Stickopttt Bb., Diploperi-
derie Bb., ffohthuria Bb., Bohadechia Jabo., MuUeria Jabo., Tr^ang
J ABO.)
Sp. ffolotkuriatuhvlosa, Fietularia tvhidosa JjAU.; ffolothuria tremula Gmbl.,
Cuv., BoHADSCH de quibusd. Animal, marin, 1761, Tab. Yi., Tibdbmann
Anai, Tab. i., Cuv. R, Ani, id. HI. Zooph. PL ao, fig. 3 ; one of the largest
of the species in the Mediterranean. Twenty tentacles, placed alternately
in two circles, surround the mouth. Jffoloth. argus, Bokadschia argue
Jabgeb de ffoloth. Tab. n. fig. i, on the coasts of Celebes and Timor.
Iloloth. edvXis, Trepang eduUe Jaeg., Less. Cent. Zocl, PI. 46, fig. 2,
GuiBiN Iconogr, Zooph, PI. 4, fig. 7 ; this species lives on the cond-reefii of
the Molucca, Philippine and Carolina islands, and is fished up in large
quantities with other species under the name of Trq)ang, to be dried and
smoked and prepared with condiments for the use of the Chinese and
Malays, who consider it an exceedingly nutritive and stimula^g {aphro-
dieiacum) article.
I) Tentacles ramose.
Genera : Cladolabes Brandt (tentacles 20), Cohchirus Trosch.
(Tentacles ten, two smaller).
Psolm Oken (in part). Feet equal, disposed on a ventral flat
disc, on the back none. Tentacles ramose.
Psoitis Jaeg., Br. Body attenuated backwards, covered with
skin rugose or scaly. Three rows of feet in the ventral disa
Sp. Psolus phaniapus, ffoloth. phantapus L., Zool. dan. Tab. in, FoBB. Br.
Starf. p. 403, GUKR. Iconogr. Zooph. PI. iv. fig. i ; in the North Sea.
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ECHIKODERICATA. 159
Ouvieria PisoN. Body often oon^ex, oovered vith calcareoiu
imbricate^ scales, plane beneath, beset with nmnerons feet
Psoiiu tquamaiui MuxLL. Zool. dame. Tah. X. &g. 1—3^ Gu^iK Jeonoffr.
Zooph, PL 4, fig. 2, &c.
Order n. Apoda.
Bodj covered with a coriaceous, sometimes soft, skin, destitute
of ambulacral feet.
Family V. Synapttnce. Body cylindrical, elongate. A cal-
careous ring around the oesophagus. Mouth crowned with tentacles.
Most writers unite these animals with the HolothuricB. But
beyond doubt they form a distinct family, of which, however, the
structure has been made known to us only in a single genus by the
investigations of Quatbefages.
Ltosama Brandt. Tentacles peltate. Eespiratory organs
arborescent.
Bp. Lioaoma SUchaente Bb.
Chirodota EscHSCH. Tentacles digitate at the extremity.
(Respiratory organs ?) Skin thickish. Body vermiform.
Sp. Chirodota discolor, EscHSOH. Zooloffiaher Adas n. folio, Berlin, 1829,
p. 12, Tab. X. fig. 2 ; CMrod. verrucosa Eschsch. ib. fig. 3, both from the
North-west coast of America ; these animals live in the sand and under
stones op the shore, in situations which are not quite dry at ebb-tide.
Here belongs also Bohth. purpurea Less., CetUur. Zool, PL 52, fig. 2, and
Jffolothuria digUata Moktaoit, Lkin, Transact, xi. Tab. iv. fig. 6. Accord-
ing to Bbandt there are situated on the mesenteiy small cylindrical bodies
divided at the extremity, which are subservient to respiration.
Sjfnapta EsCHSCH., Tiedemannia Leuck. Tentacles pinnatifid.
No arborescent respiratory organs. Skin very delicate, rough from
booklets extremely minute, calcareous.
Ebohboholtz named this genus (from tf-iWrrw, adoiteeto), on account of its
adhering to the skin by means of small booklets, which he compares to the
appendages of the calyx of the Burdock {Amtivm lappa). He found
Sjfnapta mamtOosa on the coast of 0tah4nii, Zool, Ad, n. Tab. x. fig. i.
To this genus belongs sJbaoHoloth, oceanica Lbss. Centur. Zool, PI. 35 ; HoUAh,
radiosa, ibid, PL 15 ; Fistidaria vittata Forsk. Icon. Iter. Nat. Tab. xxxvn.
tg. 2, Ac. QuATSBFAOBS discoYered a species in the sand at the Ghausey
T«1a»^ which, in my opinion, has much resemblance to the last-named species
of F0B8KAL, and gave a detailed description of it. M^moire sur le Synapte
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160 CLASS IV.
de Duvemoy {Synapta DuvemcBa), Ann. des Sc. Nat, ae S&ne xvn. Zool.
pp. 19 — 93, PI. 2 — 5. Bespiration is effected in the abdominal cavity,
into which the water passee by five apertures between the tentacles. The
hooklets, shaped like anchors, one-tenth niillim. in length and less, are set
on oval eminences of the skin, which are visible under the microscope alone.
Other similar eminences are covered with cells which can evolve an
extremely fine thread (nettle-organ?). These anfmals appear to endure
wounding and extreme mutilation without being destroyed.
Note. — Ought the genus Haplodactyla Grube to be placed here 1
Tentacles cylindrical^ simpla The author of the genus says nothing
about feet. Five respiratory organs, laciniate, sub-arboresoent Gbube
IL p. 42.
Molpadia Cuv. Body tending to cylindrical or sub-pentagonal,
coriaceous, attenuated backwards. Tentacles short, cylindrical,
simple. Mouth armed with an apparatus of calcareous particles.
Sp. MoU>pctdia holothwrioldes Guy. ; Mciop. muscultu Bisso, Eitrop. tnSrid.
Tom. y. figs. 31, 3a. Comp. Cuv. R. Am., m. p. 241, Blainv. Man.
dPAcUnU. p. 651.
Family VI. Sipunculacea. Body cylindrical, elongate. No
calcareous ring around the oesophagus. Mouth provided with
a retractile proboscis.
Sipuncuhis L. Body round, elongate, annulate. A papillated
retractile proboscis, with incised tentacular border or coronet of
simple tentacles surrounding the mouth. Anus a lateral aperture
of the body, situated towards the anterior part.
Sp. SipunculvM nvdus L., Syrinx Bohadboh, de qwbutd. animal, marin.
Tab. vn. fig. 6, 7, Fobbsb BrU. Stanf. p. 245, Leuokabt Brevet animalitm
quorundam DeecripHoneef Heidelbergse, i8a8, 4to. fig. 3, and especially
Gbube in Mubllbb*s Arehiv. 1837, Tab. x. fig. i ; in the seas of Europe,
especially the Mediterranean. This animal can attain a length of more
than one foot, exclusive of the proboscis, which is nsuaUy retracted. It is
very contractile and sometimes shortens itself one-half. Longitudinal and
circular bundles of muscle, visible through the skin, give it a latticed
appearance. The intestinal canal is very long, making two bends back-
wards and forwards ; the last ascending portion is wound spirally round the
preceding piece. Two brown vesicles, connected with the propagation,
terminate with fine apertoires before and by the side of the anus. The
nervous system presents two rings round the oesophagus, and a cord
running on the abdominal surface, that gives off numerous branches, and is
surrounded by a blood-vessel as by an envelope ; see Kbohn in Mublleb's
Arehiv. 1839, ^' 34^- From this species Lumineui phaUoidet Pall.
SpieUeg. Zool. x. Tab. i. f. 8, seems not to differ. Sipunctdu* edulu,
Lumbr. edulit Pall. ib. fig. 7, is a species eaten by the Chinese.
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ECHINODERHATA. 161
FoBBBS givM the name of Syrmx to the spedei which have a short
proboacb with an indented tentacular fold around the mouth, like Sipune.
tmdut. In other species the proboscis is longer, and there is a ring of
simple lancet-shaped tentacles round the mouth. Of these he forms his
genus SijnmeuluB, which appears to agree with PMaaeolommM LiuoK.
Sp. PhaseoUmma granuUshim Lbvok. L 1. fig. 5, from the Mediterranean,
probably not different from Ascomnna Blumenbaehii, Und. fig. 5 ; SipunculuB
eapUahu Bathkb, JVbr. Ad, Aead, Leop. Car, xx. i, Tab. vi. fig. 20, 21 ;
Sip, Bemhardut Fobb. Brit, Starf, p. 951 ; it Hves, like the Hermit Crab,
in the empty shells of Molluscs (Stromhw pei pdeeani, LittoriuM, TurrOeUa,
DaUaUum, Ac.)
J^riaptUtui Lam. Bod^ (r^lindrical truncated posteriorly. Pro-
boscis retractile; no tentacles around the mouth. A branched
pyramidal appendage hanging from the posterior part.
Sp. Priaptihu eaudatui, HdUtikima priapui L., Zool. damica, Tab. xcfvi.
fig. I, and OZXXY. f^. 2, Fobbbb Brit. Starf, p. 156. Gu^BiN Iconogr,
Zoaph. PL 5, fig. I. It is supposed that the bundle of threads at the
extremity of the body serves for respiration. This singular animal fives in
the North Sea.
Banellta Rolando. Body oval, contractile, with a long pro-
tensile proboscis, divided at the extremity into two lacinise. Anus
posterior, terminal.
See BOLAHDO in Jaum. de Phytique, de Chim., ^ffid, not, Ac. Tom.
xcv. Juillet, 1813, pp. 49, 59, av. fig. (from the Mim, de TAead. de Turin,
Tom. XXVI. pp. 357, Ac., Tab. XIV. xv.) The long proboscis was taken for
a tail by this writer, the anus described as mouth ; near the aiMte two
trussed organs are situated (salivaiy organs according to Bolabdo), which
correspond to the arborescent respiratory organs of HoUdkuria, The
intestinal canal is narrow and long, with two loops and many undulations.
Sp. Bondlia viridii L L fig. i, Guj^bin leonogr, Zooph, PL 6, fig. i ; B<m,
fuliginoea. Theaa two species were found by Kolaitdo on the shore of
Sardinia.
Thala88ema Cuv. Body cylindrical contractile. Proboscis short,
with spoon-shaped appendage, or sulcated lacinia, undivided. Two
shining uncinate setsB in the anterior part of the body, behind the
proboscis.
* Body smooth posteriorly. Thalasiema Gaebtneil
Sp. ThaUueema N^uni Gaibtn., Pall., Spicil. Zool, x. Tab. i. f. 6, GufBiv
Iconogr. Zooph. PI. 6, ^, 2, FOBBSS Brit, Starf, p. 159. (Here also seems
to belong the genus OcsMostoma Lbuok., Sp. OcKd. eryC^ro^nMimon, AULae
zu der Reiee YoN Ed. BuFFXLli, Neue wirbeUoie Thdere dee rolken Meeree,
1818, Tab. n. ^, 3, HoMh, eaouari Lxas. CenL Zool. and GuiB. Iconogr,
Zooph. PL 4, fig. 6.
VOL. I. 11
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162 CLASS IV.
** Body armed posteriorly with zones of homy setae. Echiurus
Cuv. (Echiwwrua),
Sp. Thcila89ema eehiurtu, Lumbricua echiuna Pall., Misc. Zool. x. Tab. i,
fig. I — 5, FoBBES, Brit, 8t<vrf, p. 263, Gu^ Iconogr, Zooph, PL 6, fig. 3 ;
on the coasts of the North Sea ; this species is used by the fishermen for
bait; it is flesh-ooloured, here and there translucent blue and red; the
spines are shining and yellow, as far as they project from the body. The
orange-coloured intestinal canal is long, with many undulations and
vesicular expansions. There are two long, brown, somewhat tortuous,
csBcal tubes near the anus, and four white vesicles at the anterior part
of the body, which belong to the procreative apparatus. The nervous
system consists of a thread running along the intestinal cavity. The skin
has much muscularity, sp that when wounded it spirts out the contained
water like a fountain, and then the intestines are forced out.
**• Body furnished anteriorly and posteriorly with zones of
setae ; with belly in the middle plane and in front of the amvs
sciUate, (Setae at the sides of the scute; anaJ tubule oonical,
retraotila) Stemaspis Otto.
Sp. Thakusema wuUxtua Rakzaiti, Skmaapit thalaaaenundes Otto, Nov,
Act, Acad, Leop. Cwrd, Tom. x. Tab. 50, GuiBiN Iconogr, Zooph, PI. 6,
fig. 4, from the Adriatic. Otto took the conical extremity incorrectiy for
the mouth ; see the communication relative to the structure of this animal
by Kboeet in Mublleb's Arclm. 1842, s. \i6, Rakzani, who described it
before Otto, rightiy distinguished the two extremities of the body, and also
figured the spoon-shaped prdbotcis, in which it resembles EcKiwrut. Op%i»GoU
tcient^fiee i. Bologna, 18x7, 4to. pp. iia — 116, Tab. iv. f. 10, 11.
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CLASS V.
INTESTINAL WORMS {ENTOZOA)\
Bt Intestinal Worms are understood worms that live in other ani-
mals, (not necessarily in their viscera). Though not more correctly
in point of language, they might, perhaps, be better named Internal
Worms, which would be a literal translation of the word Entozoa.
This class is by no means natural, for the animals included in it not
only differ remarkably from each other in external form, but also
in internal structure. The efforts that have been made hitherto to dis-
tribute the different orders and families of the Entozoa amongst the
different classes of invertebrate animals have not been satisfactory.
In this, as in so many other attempts to determine the affinities of
animals, a certain external resemblance has been mistaken for
correspondence of internal stmcture and of type of organisation, by
which alone true affinities can be established. An unprejudiced
inquirer will scarcely attach more value to a comparison of Echino-
rhynchua with Sipunculua, of Cyaticercus with bladder-bearing
Acalephs, &c. than to a comparison of bats with birds. Consequently
we are compelled to retain the class Entozoa, however we acknow-
^ Compaore on thiB class :
C. A. RuDOLPHi^ Entozoorum sive Vermium irUesUnaUum ffistoria naturalu, m.
Totm, c. tab. aen. Amstebedami, 1808 — ro, 8vo.
C. A. KuDOLPHi, Enioeocrum SynoptU, eui cLccedunl Mantissa duplex et indices
loevpleUssimif c. tab. m. ten. Berolioi, 1819, 8yo.
Db. Brbmsbb, Ud>er lebende WUrtner im lebenden Mensehm, Mit iv. Ulum. Kupfeit.
Wien, 1819, 4to.
Bbzmssbi Icones Hdminthum. m. Fasciculi. Yiennse, 1823, folib.
J. Cloqubt, Anatomie des Vers ifUesHna/ux Ascaride hmbrieoide et Eehinorynqus
g€a^, Avec 8 pi. Paris, 1814, 4to.
A. Yon Nobdhann, Afihrographische Beitrage zur NaturgeschicJUe der wirbeUosen
Thieve, Istes Heft. Mit ro Kupfertaf. Berlin, 1832, 4to.
F. DuJABDiN^ Histoire nalurdU des ffdminthes ou vers intestinavx. Avec 12 pi.
Paris, 1845, 8to.
Other important, more special works of Mbblis^ Cbbflut, Lbxtoeabt, G. Th. Yon
SiBBOLD, will be indicated farther forward. See also the article Bntosoa of Owen in
ToDD*B Oydcpadia n. pp. 110 — 144 (1837).
11—2
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164 CLASS V.
ledge it to be unnatural. Rudolphi, to whose investigation this
part of Zoology is so greatly indebted, compares the class of the
intestinal worms to a Fauna, a collection of animals that live in a
certain region. Their country is the living body of other animals.
Becent observations have, however, taught us that some species in
the cycle of their developments, leave their country for a time. The
radial type, which we observed in the former classes, particularly
in the Polyps and Acalephs, is here wanting. All Entozoa approxi-
mate to the elongated form of the articulate worms, although in
some of them that form, on account of a bladder to which they are
affixed by their extremity, is not at first sight apparent. Special
respiratory organs are wanting. Some are entirely without sex,
and are propagated by gemmation ; others are bisexual ; in others
the sexes are distinct.
We must premise something as to the division of the Entozoa,
EuDOLPHi adopts five principal sections, which had been already
distinguished by Goetze and Zeder before his time. The first is
that of Cystic worms {Cystica firom rworw, a bladder). The body is
compressed or cylindrical, the posterior part of it passing into a
bladder ; sometimes many such enJtozoa are fixed on one bladder.
The head has suckers, a circlet of hooks, or iom prdbosddes armed
with hooks. No genital organs have been discovered; nor any
intestinal canal. [These have been shewn to be imperfectly deve-
loped Tcmice.] The second division contains the Tape-worms
{Cestotdeay fix>m KtaT6t, a girdle, a band, ti^, form). They have an
elongated, compressed or flat body that is mostly jointed. The head
has suckers ; all the individuals are bisexual ; an intestinal canal is
wanting. The third division is formed by the Suctorial Worms
{Trematodes, rfnuAankbiiu perforate). The body is soft, compressed
or roundish, and provided with one or more suctorial pores. The
intestinal canal has in general only one opening, and is usually
branched ; it lies in the tissue {parenchyma) of the body, not in a
firee space. The genital organs of both sexes are united in the
same individual. The fourth division is that of the Thorn-headed
Worms {Acavihocephala, from oKovOa, a thorn, and xc^iy, head) ;
they have a cylindrical, pouch-shaped body. The sexes are
distinct ; an intestinal canal is wanting. The fifth division includes
those intestinal worms which have an intestinal canal, with both
mouth and anus, suspended fii-eely in the cavity of the body. The
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INTESTINAL WORMS. 165
body is cylindrical. The sexes are distinct. They are named Thread-
worms or Bound'toarms {NemcUoidea, from i^fia, a thread, and ccdof).
CuYiER makes a distinct order of this division, that of InUsttnaux
caviiatres; all the remaining entozoa he unites in a second order
nnder the name of Inteatinaux parenchymateux. OwEN has denoted
these two principal divisions of CuviEB by the names Sterelminiha
and C(ElelminlJia^. In onr compressed description of the structure
of the intestinal worms we shall avail ourselves of the names
which we have explained.
In the cystic worms, the tape and thorn-headed worms, there
is neither mouth nor intestinal canal; the nutriment is introduced
by absorption of the skin. In the thorn-headed worms two parts
are met with by the side of the sheath of the proboscis, usually of
a flattened form, and very small anteriorly. These parts, called
Lemmsciy are, according to Rudolphi, subservient to nutrition.
They contain, according to the description of Yon Siebold, a finely
granular parenchyma, and are very vascular.
In the suctorial worms the intestinal canal commences with
an cesaphagus^ more or less long, which, at its fore part {pharynx)
is surrounded by muscular walls. In those that have a sucker at
the anterior extremity of the body, the oral aperture is situated at
its bottom. The intestinal canal divides, below the (xsophagus, into
two branches, which have csecal terminations backwards. In many
species these branches do not subdivide^; frequently they expand at
their termination. In other species these principal branches give off
other branches. This is especially the case in the Liver-fluke of
sheep {Distoma hepaticum), where the branches divide to such a de-
gree that the whole canal has an arborescent appearance. The oral
aperture performs also the office of anus; at least the Distomes,
which are taken alive from the liver, and exposed to air or placed
in water, reject by the mouth, entirely or in part, the brown fluid
with which the intestinal canal was filled*. The genus Pmtastoma
or Lingtuttula^ differs from all the other suctorial worms in the
1 Todd's Cjfdop<ed%a n. p. iii.
* Ab in Ditkma perlatum Nobdmakk, h eii. Tab. iz., Dittoma ro$aeeum, ib. Tab.
ym., IHfioBlomata foimd in the vitreone humour of the eye of the Perch, i6.
Tab. n. nL, and in many others. See also BIaubb BeUr&ge sur AnaUmie der Entazoen,
Bonn, 184T, 4to.
* Comp. E. MzHLiB Ofmrv<Uume$ analomuxe de ZHttOTnaie hepoLico et lanceoUUo,
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166 CLASS V.
structcire of its intestinal canal as well as in other respects. On that
account Cuvier and Owen have with propriety removed it from this
division in which Rudolphi placed it. The intestinal canal lies in
a free cavity of the body, though covered by the coik of the oviduct,
and ends with a distinct anus ^ The position and form of the canal
agrees with the same in the Roundrworms; only in the oesophagus
is there some difference, since this tube in the NemaUndea runs
from the mouth at the anterior extremity of the body backwards in
the same plane with the intestinal canal, whilst in PentasUmia it
ascends obliquely because the mouth is situated on the abdominal
surface. In the NemaUndea the cesophagus is muscular, and in
many species wider at its termination. The intestinal canal that
succeeds it is straight, and its whole course continues nearly of the
same width. In Aacaris lumbrtcotdes pedunculated pyriform vesicles
axe foimd, which adhere to the internal surface of the integum^it,
and occupy the space between the skin and the intestinal canal.
A vascular system has been discovered in many entozoa.
[Amongst the NemcUotdea Blanchard has described in Aacaris
megalocepJuda Cloquet two longitudinal vessels lodged in each of
the lateral canals within the integument, which extend from one
extremity of the body to the other. At about the depth of one
third of the oesophagus, the two, supposed to be arteries, leave their
tubes to form an arch behind the oesophagus ; on the arch a small
ampulla is seen which is supposed to supply the office of a heart.
The two arteries descend in the tubes throughout the whole length
of the body, and communicate with the two other longitudinal
vessels supposed to be veins K In the tcenta the longitudinal canals,
four or six in number, communicate by transverse branches, and
open in the last joint into a pulsatile vesicle, which expels their
contents in drops at intervals. In the suctorial worms the fine
vascular network, hitherto considered to be a circulating system,
has been shewn by Van Beneden to be an appendage of the
tubular system, which terminates in a vesicle that opens externally
by 9k foramen caudale. The apparatus in the last two families of
1 See Owen Transact, ZaU. Soc. I. 1835, PL 41, fig. 12 ; DiEBiSQAnn, dea Wiener
Museums, i. 1856, Tab. 11. fig. 2 ; comp. Tab. i. fig. 20, of PetUasloma proboseideum.
• Blanchard Ann, des Sc, not. $e S^rie, Zool. Vol. xi. pp. 146, 147, and Cutixb
R. Anim. idU. iUus, Zoaph. Fl. 26, fig. i c.
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INTESTINAL WORMS. 167
worms appears to be similar, and for the purpose of secretion; a
renal secretion, as Van Beneden supposes \] In the thorn-headed
worms there are two lateral canab situated beneath the skin that
nm the whole l^igth of the body.
We have already remarked that special respiratory organs are
wanting. So far as any action occurs between the air of the
medium in which these animals live and their nutrient fluid, it must
be efiPected by means of the skin. But Entozoa live, for the most
part, in situations where the atmosphere exists in a condition very
impure and unfit for respiration ; or where no air at all can enter,
as in the liver, brain, kidney, &c. It is therefore probable that
they derive firom the fluids absorbed from the animals in which they
live, the quantity of oxygen necessary for their life, and that they
experience the influence of this gas only mediately through the
animals in which they live *.
With respect to propagation : no genital organs, as noticed
above, have been detected in Cystic worms. What many writers
have described as eggs in these worms are calcareous corpuscles
beneath the skin, which also occur in Tape-worms. Their multi-
plication is eflected by gemmation. In Ccenurus there arise on the
bladder on which the worm, or that extremity of it that bears
the head, is seated, little buds which again develop other buds ; in
Echinococcus new bladders are formed within the parent bladder,
like cells within cells, in which young Ediinococci are developed
that continue hanging by a thread for a time, after the containing
envelope is ruptured, and then fall into the cavity of the parent
bladd^^ In Cysticercua the mode of propagation is yet unknown.
In T^remaioda there is found on the abdominal surfsice, generally
nearer to the anterior than the posterior extremity, an opening
common to the organs of both sexes. From this a penis, usually
named drnu*, can be evolved; near this part the vagina opens.
1 Yah Bbnxdsn LeUre relative d Vlfist. de» ver§ cetlcides, Ann. des Se.nat, 30 S^rie,
Zool, VoL XVII. pp. a I — 30.
' Comp. OD the respiratioii of intestinai worms, BuDOLPHi Hid. not. £ntoxoor, i.
pp. 139 — 944, uid Cloqubt An<tt» des vers vntetUn$, pp. 42 — ^44.
' MuxLLSB in the Jahretbericht for 1835, Arckiv. s. ovn. OVIU. ; V. Sibbold in
Buxdaob's Phynol, ite Aufiage n. s. 183—185.
^ See the fig. of JOittoma hepaUcum in Mehlib, fige. 8, 9, 11. In fig. 8 is seen
near the cirrua the opening of the vagina, thi-ough which a bristle has been passed to
distinguish it.
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168 CLASS V.
The testes have mostly a rounded form ; in Amphistoma subtriqw-
trurn and gigatUeumy thej are finger-shaped and branched^. From
these testes, nsually two in ntunber, efferent vessels proceed to a
seminal vesicle lying at the base of the penis in the Cirrus-sac
{recq^tacidum Penis) ; firom which a canal arises that runs to the penis.
But besides this, one of the testes gives still a third vas deferens to
a seminal vesicle lying further behind [vesicula seminaUs interior) ^
and from which arises a short tube connected with the oviduct^.
Here self-impregnation may be effected : in which case the second
vesicula seminaUs and the external genital organs are difficult of
explanation: unless we suppose that both self-impregnation and
copulation are possible. In the female organs of the suctorial worms
we would direct particular attention to that remarkable arrangement
by which, according to V, Siebold, the yolk and the germ {vesicula
germinaiiva) are not produced in the same organ: — ^that here we
must consequently, instead of ovary, distinguish a germ-stock and a
yolk-stock. The lateral parts, usually dendritic or botruoidal, which
were formerly believed to be ovaries, are the yolk-stocks: the germ-
stock is placed in the middle of the body, and has a roundish form.
In the Tape-tDorms the two sexes are also united : and V. Siebold
suspects that in this case, also, the germ and yolk-stocks are dis-
tinct organs. In the jointed Tape-worms {Tomia, Bothriocq^hahis)
the sexual organs are situated in every joint, only the anterior more
recent joints do not yet indicate them, because they become more
perfectly developed in proportion as the joints are more posterior.
These animals, therefore, during their growth present us with a
successive repetition of the same organisation. Some authors think
that eveiy individual joint is to be considered as a suctorial worm,
and the Tape-worm as a compound suctorial worm. The sexual
openings are situated in every joint, either on the edge or in the
middle. In Bothriocephalus lotus, for instance, on the abdominal
surface of the body there is a fold of skin in the middle of each
1 AmphiiUma triquHrum, Bojanub liit, 1B21, truuferred to Sohmalz TdlnU.
Anatomiam JBrUoeoar. illuitr, Dresda, 185 1, Tab. Tm. figs. 7 — 9 ; AmpkuUma gigtm-
teum, DiEsnra in Wiener AnntUen i. Tab. xxii. figs, g, 14, 15.
* V. 81XBOLD found this arrangement in many apeeiei of JHiUma, and suspects that
it oocun in all. See BuBXSiSTXB on Didotna ghbiporum in WnOM. Archiv, i. 1835,
B. 187 ; V. SiXBOLD, ibid. n. 1836, s. 117, Tab. vi., and in Mubllbb's ArMv. 1836,
B. S35 — 137, IHstoma noduloeum, Tab. z. fig. i.
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INTESTINAL WORMS. 169
joint, jet nearer the anterior edge, with two openings : through the
anterior and larger the penis is evolyed; the posterior smaller
is the female sexual opening: round both of them are minute
white points which Eschbicht^ concluded, under high powers of
the microscope, to be follicles (mucous crypts of the skin). The
eggs of Boihriocephalu8 have a hard shell, as in the Distomes^ of a
brown or brownish-yellow colour, and seem like them to spring
open with a sort of hood. In the thorn-headed and round-worms
the sexes are distinct, and may be often recognised externally
by their different form and size. In the thorn-headed worms the
sexual organs iill the greater part of the cavity of the body. From
the sheath that surrounds the proboscis there runs backward in the
axis of the body a band-like structure, which has been erroneously
supposed to be a canal, but which lb for the support of the organs
that prepare the germ or the seed {Ugamentum sutpeiwarium) ; YoN
SiEBOLD supposes' that even the ovaries are developed in this organ.
These ovaries are found free in the abdominal cavity, as masses of
oblong-roimd eggs : the eggs become detached as they advance in
development. The muscular oviduct terminates in a very small
opening, scarcely visible at the posterior part of the body : it has at
its anterior extremity an infundibular expansion which alternately
widens and contracts, and takes up the eggs that were floating freely
in the cavity of the body and moves them onwards* to the oviduct.
This arrangement, in virtue of which the oviduct opens freely into
the cavity of the abdomen and is not an immediate continuation of
the ovary, is found in most vertebrates, with the exception of the
osseous Fishes, but has not hitherto been observed in invertebrates
except in Echinorhynclius. In male thorn-headed worms there are
usually two testes lying one behind the other. The penis lies in a
sac having a conical appendage that can be everted from the body
in the form of a little bell*
Amongst the Thread-worms the males are less frequent than the
females : they are smaller and more slender, and may frequently be
1 See EflomnoHT AnaU)mUck-pkiftioU>giK^ UiUermckungtn uAer die Btdhruh
eepkalen; Act, Acad, Cam, Leap. Carol, Nat, Curia, Vol. ziz. Snpplem. n. 1840.
* VoK SiSBOLD m Buboagh'b Physiologie, n. b. 197. See » figure in Buftow
Bekinorhyncki itrumori Anatome, Diss. 2jOotom. Rogiomonti, 1836, 8yo. &g, ig. fig. 6 ;
oomp. DujASDm op. cit. p. 494, PI. vn. fig. 7, D 5. {Bchinorhynehut anlhwrit, a species
from the freshwater SaUinander).
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170 CLASS V.
recognised by their curved extremity or by the sexual organ exter-
nally visible ; this penis is in most species double, in Triooc^Aalus
and Trtchasoma single. The organ which prepares the germ or
seed has the form of a slender convoluted canal. It is single in the
males : in the females, with few exceptions, it is double. These
canals are of great length: according to Cloquet, in the male
Ascaris lumbrtootdes when quite unravelled it measures from 2^^ — 3
feet, and in the female each of them measures 4 — 5 feet. The dif-
ferent divisions of the canal may be considered to be ovary or testis,
and vas deferens or oviduct with lUertis. The terminal portion in
the female is distinguished as uterus by its greater circumference
and its vigorous peristaltic motion. A very wide, longish sao-
shaped structure at the termination of the canal in the male is to be
considered as vesicula seminalis. The external sexual opening is, in
the male, always situated at the posterior extremity of the body ;
in the female ordinarily further forward, and in some species in the
middle, or towards the anterior extremity ^ Non-sexual Nematoids
are met with* ; Creplin gives it as a general rule that a Nematoid
living in a perfectly closed cyst, or shut up on every side by a mem-
brane, never has sexual organs. It has been suspected tibat these
species are in an incomplete state, and can attain their perfect deve-
lopment only in other localities*.
We here approach what, until very recently, was one of the
most obscujre problems in the economy of the Entozoa: and
MiESCHEB was fiilly justified in his remark that many of the
observations relating to their development are riddles of Natural
History*. [With respect to the suctorial worms our information is
in fact only fragmentary : yet since the observations refer to veiy
different periods of their development in different Trematodes, we
are able from analogy to collect a tolerably connected history of the
whole process in any one of the class.] It is well known that the
^ In Ascaris lumhric&idea, these parts are figured in the work of Gloquet already
cited, PI. n. figs. 8— TO, PL rv.
• Von Sibbold, Wibgm. Arekivf, Naturgtick, w. i. 1838, s. 301—311 ; GBXFLiir,
ibid, s. 373.
' M1B8OHEB, however, has observed OYaries in Fiiaria PiteiMm, Wisomakn's
Archiv, 1 84 1, II. s. 301.
* F. MiEBCHSB JSeachreibuny und Uniersuchung da MonoMoma h^ugum. Basel,
1838, 4to. 8. 14.
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INTESTINAL WORMS, 171
Danish zoologist Muelles classed amongst his Infbsories minute
worms with tails, to which he gave the generic name of Cercaria.
The aocnrate observations of NrrzsCH taught us to define the genus
more completely, and supplied a lively picture of the form and
movements of a worm that to the naked eye seems like a moving
point ^. He compared these animals to Distomes that have had a
tail-like appendage attached to them, and their motions to those of
a Vibrio: each of these structures, the body and the tail, had a
separate motion : when the body moved on by creeping and sucking
the tail was at rest; aad, conversely, when the tail tmdulated rapidly
it forced along with it the body, which seemed now to have no
independent motion of its own. NiTZSCH further saw that Cercarios
cast thdr tail, and observed in Cercaria ephemera that it fixes
itself, covers itself with a shell that presently hardens, and thus
becomes, as it were, a pupa. Such pup« remain unchanged for
months: what was to succeed remained m[iknown. Afterwards
similar observations were made by others, particularly by VoN
Si£BOLD. Steenstbup' shewed that the pupss change into Di-
stomes after the lapse of several months. Cercarice live as parasites
within the bodies of different Molluscs, as Lymnasus and Flanorbis.
Bnt this does not terminate the surprising series of these changes.
CercaricB, the larvsB of Distomes, do not arise immediately from the
eggs of these last. Within the above-named Molluscs, and in some
others, little bags of an oblong form {germ-pouches) are found, in
which a peculiar organisation and motion may sometimes be traced,
but which in other species are motionless, and contain CercaricB in
a more or less advanced state of development. Considered as para-
sites of the germ-pouches, these have been named parasites of the
second order: yet they are not parasites, but the progeny of these
vermiform germ-pouches*. How these last originate fix)m the young
of Distomes has not been absolutely ascertained. The young animals.
1 C. L. NrrzaoH BeUrag zur Ii^u»orienhmde, Halle, 1817, 8yo.
' Stxbnbtbup AUenvUion of Oeneration, Translated by Bubk for Raj Soc.
' BojAKUS, who diflcoyered such yermiform germ-sacs of a yellow colour in lAm-
fUguM tUtgntUu, named them Kin^s ydlow leomu; Im, iSiS, s. 799. The celebrated
y. Baeb published many similar obseryations in Nov. Act, Cass. L. C, Not, Cur.
VoL xni. P. 2, pp. 605---659, Taf. xxxi. ; as also the €ftr-fiuned inyestigator of the
lower animal forms V. Siebold in Bubdacr'b Phytiol, 2^ Ausgabe, s. 186, ftc.
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172 CLASS V.
that proceed from eggs of Distomes, move rapidlj hj means of
cilia with which their body is covered like that of many Infusories,
In such a young animal Yon Sieblod saw in Monostoma mtUabile
a parasite lodged which agreed in form with that of the germ-pouch
of Cercaria echinata, [He concludes that the germ-pouch is thus
shewn to be the descendant of a Trematode. The Monostoma mutor
bile is a parasite of water-birds and lives in cavities of their body
which communicate with the external world. K a MonoaUyma mtU.
has produced an embiyo, this can readily escape from the abode of
its parent into the water, and may by means of its cilia find in-
stinctively the animal suited to supply a fit residence for the further
development of the germ-sac which it contains. It may pass into
the interior of that animal by some one of the natural openings.
Having now ftilfilled its office of a living envelope to the germ-
pouch, the ciliated embryo will die, and the germ-pouch seeks by
perforation that situation within the body of its host which is suited
to its further growth, and to the supply of due nutriment for the
Cercariar-brood which it contains. The greater number of suctorial
worms are, when they have gained their sexual organs, parasites of
the higher vertebrates. How then can Cercaria effect an entrance
into the body of such vertebrates as never come near the waters in
which they live? VoN Siebold has given a probable answer to
this enigma. He very commonly found in the aquatic larvae of
EphemeridcB, PhryganidoB, LtbeUultdce, &c., as well as in the perfect
insects, encysted Cercartce which had divested themselves of their
tail, and which in fact were larval trematodes. In order to effect a
passive migration into the vertebrate in which their development is
to be completed, they must wait until their temporary host is
swallowed by some insectivorous land bird or mammal ^] Though
there still remains an ample field for conjecture and fancy, thus
much may be concluded from the observations hitherto made, that
in these animals (as in Medusae, vid. pp. 100, 101) a succession of
alternate generations occurs : that the first series does not resemble
the parent, but that from it young ones proceed which return to the
original form. The germ-pouches constitute the first series, the
nurses (nutrices) ; they are the starting-point of animals to be bom as
* Comp. Von Sibbold Die Band u, Bkuen-wOrmer, Leipeig, 8vo. 1854, pp. 21— 31.
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INTESTINAL WORMS. 173
larvse, CercaricB^ from the larval state to be changed into pnpsB, and
from this finally to become Distomes^
Besides these changes of Distomes, of which the entire series, in
its chief features at least, has been investigated, there are others
relating to Tcenim which [until lately] were only observed firag-
mentaxily. Such was that of Leblond, who found in the peritoneum
of Murcena conger a worm enclosed in a cyst and containing a
young Tetrarhynchtis. He described this larva of Tetrarhynchtia as a
species of Amphistoma and the Tetrarhynchua as its parasite*. Mies-
CHER also made similar observations. Sometimes the successions of
development appear to be possible only on change of abode. The
simple LigidoB of fishes are found, according to Budolphi, in birds
that feed on fishes in their more perfect form and furnished with
developed sexual organs: the worms of th& [supposed] genus
Soolex, that live in Fleuronectce, are probably changed in the bodies
of Kays and Sharks into Bothriocephalic and the Bothriocephalus
9olidu8y that lives in Qasterosteua pungititiSy is changed, according to
Abildgard* and Creplin, into the Bothriocephalus nodoaus of water-
birds {Merfftts, Colymlms, &c.), which feed on that fish. That worms
should thus continue to live in other animals becomes less surprising
when we consider their tenaciousness of life; Liguloe have been
found alive in under-cooked fish ; Rudolphi found individuals of
Ascaris speculigera stiff and hard in the gullet and stomach of a
Cormorant that had been kept for eleven days in spirit of wine
which returned to life in warm water : and MiRAM saw individuals
of Ascaris acus firom the Pike dry and sticking to a board revived
by water, and in some instances moving a part which had imbibed
the fluid whilst the rest continued shrivelled up and adhering im-
moveably to the board*.
These observations prove, by the way, that it is not necessary
to have recourse to equivocal generation in all cases of Entozoa
where their existence appears to be inexplicable by the ordinary
mode of propagation.
[Tape-worms attain their full development and mature sexual
^ See SnsHSTBUF AUematum qf CfmeraHon,
' Awn. de» 8c. nai. aec. S^rie. Tom. vi. Zool. pp. 289 — 195, pi. 16, f. x-
* Rddolfhi SnUuoor. Hid. Nat. n. P. u. pp. 60, 61.
* WnEOMAHs's ArrMvf. Noburgack. 1840, I. b. 35—37.
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174 CLASS V.
organs only in the intestinal canal of vertebrate animals: when
found in other viscera of these animals, or in the interior of inferior
creatures, they are always immature. The ova, however, of Tape-
worms are never developed in the intestine of the animals which
harbour the parent worm: still the embryo is so far advanced within
the ovom contained in mature joints when discharged from the
intestine that its form may be distinguished. In all instances the
armature of the embryo is the same, however different it may be in
the heads of fully developed worms of different species. Thus the
embryos of Tcmia and of Bothriocephalus have both of them six
booklets, though the head of a developed Tcmia is armed with a
coronet of numerous booklets and that of Bothriocephalus is unarmed.
These six booklets are not all of the same form : the pair in the
middle are not curved at the extremity like the others, they are
straight, very finely pointed, thinner throughout and also longer
than the other four, which are also disposed in pairs. The middle
pair are for penetrating soft tissues, and the rest for helping the
embryo forward when it has once penetrated them. Stein* saw
these embryos free within the intestinal canal of larvae of Tenebrio
molitor and encysted on the outside of the canal, and justly con-
cluded that the latter had perforated the canal from the interior,
having entered by the mouth. The ftiture tape-worm does not
appear to arise from the embryo by metamorphosis, but to be formed
within it by gemmation, whilst the six teeth of the embryo are
rejected when they have performed their oflSce and are found
dispersed on its outer surface. A bud is seen within the embryo,
which gradually assumes the special form of the head and neck of
the ftiture Tape-worm. As the development proceeds the head and
neck would be permanently enclosed within the embryo in which it
is being formed, were it not that at the same time a canal from the
exterior is formed around them, so far as to allow the head and
neck to be produced when the larva is freed fix)m its cyst. It is
then found that the neck of the larva is continuous with the body
of the embryo, which forms a vesicle at its extremity. To this larva
of a Toenia the name of Scolex, proposed by Van Beneden, is now
appropriated by the common consent of Helminthologists. K now
1 Stkin, in SiBBOLD and Koxllikeb's Zeiitehr. f. wittetachaft, ZooL iv. 1853,
8. 407.
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INTESTINAL WORMS. 175
the Scolex can gain a passive migration into the intestinal canal of
an animal suitable for its deyelopment, that development will
proceed; the vesicle will be cast off: joints will be formed suc-
cessively beneath the neck, and in these joints the genital organs
will be developed, the joints first formed, or nearest the posterior
extremity of the body, being the first to become mature. Thus the
Scolex is changed into a Tape-worm. Many naturalists now con-
sider the Taeniae to be compound animals, (which indeed was the
opinion long ago of Yalisnieri and Coulet and afterwards of
Blumenbaoh,) colonies, like certain Polyps: the head and neck
corresponding to the Polyp-stock, and the joints, under the name of
Proghttis, to the single Polyps. By such observations as these
Von Siebold has been enabled to interpret justly those of Leblond
and Miescher alluded to above. The Amphutoma of Leblond
was the embryo of the Tape-worm, now the receptaculum Soolecis^
the Tetrarhynchus the Scolex of a Rhyruxibothrius. When the
minuteness of these embryos is considered (they are not more in
volume than the blood-disc of the firog) it is not difficult, as Van
Benedbn ^ says, to comprehend that they may perforate the walls of
the intestine to encyst themselves beneath the peritoneum, or to
penetrate the vessels and distribute themselves with the blood in
different viscera of the body, not excepting the brain itself, or the
humours of the eye. Dr Haubner of Dresden caused six young
lambs to swallow the living and mature joints of Tamta serraUu
They all died of the peculiar vertiginous disease produced by
Ocenurus cerebraUs. The Ckenm^/a vesicles were found in the brain,
and the heart, lungs, and voluntary muscles abounded with encysted
broods of TcBnia^. It would seem from this that the different forms
of Ocenurm and Gysiicercus assumed by the larva depend upon the
locality occupied by the embryos, and the quantity and nature of
the nutriment which they obtain there. And this conclusion is
confirmed by the previous and converse experiments of Von Sie-
bold. On causing young dogs to swallow Oystic. jnstformts from
the liver of the hare. Cist tenuicoUea from the mesentery of the
sheep, Cist. ceUulosa from the muscles of the swine, Ccenurus cere-
hralta fix)m the brain of the sheep, the same form of Tcenia^ viz.
^ Vau Bbhxdbh Ann, de$ Se, tuUur. Sdrie m. Zooloff. Vol. xx. pp. 390, 331.
' Von Sisbold Band u, BUuen-wOrmer, 8yo. Leipeig, 1854, p. 106.
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176 CLASS V.
Tomta serrataj which in its developed state resides in the small
intestine of the dog, was found in all cases. The experiments were
performed so frequently, and with such precautions as to render the
results incontestable\ The cjst is a secretion of the infested animal,
and derives its blood-vessels from it.
There is reason to conclude that a nervous system exists in
most Entozoa. In Bothriocephalus Blaxchard describes a small
ganglion on each side of the head (probably connected by a trans-
verse band), which sends a fine thread forwards and backwards ; but
he was not able to detect the distribution of these*.] No nerves
have been discovered in the incomplete cystic forms. In Distomes,
and various other suctorial worms, two small ganglia have been
seen near the mouth, which are united by a transverse band that
runs over the oesophagus. From these two threads arise that have
a parallel course through the body, become finer backwards, and
give off some lateral branches'. In the thorn-headed worms there
is, according to the investigations of Von Siebold, in different
species of Ediinorhynchus^ a nervous mass at the base of the pro-
boscis, from whence threads radiate in all directions, whose course
is not easy to follow. In Pentastoma a large nervous ganglion lies
beneath the oesophagus, from which a nervous ring arises that runs
transversely above the oesophagus; small branches go from the
ganglion to the muscles of the mouth and the surrounding parts,
and two threads proceed longitudinally, one on each side, back-
wards, giving off fine branches, and, at last, losing themselves in
the muscular tissue^. In Stranffylus gigaa the nervous system forms
a flattened ring round the commencement of the oesophagus, and a
string running backwards in the middle of the abdominal surface,
which ends in a ganglion close by the anus. From the string arise
at short and nearly equal distances transverse nervous threads.
Special organs of sense are not found in the intestinal worms, with
^ See Band u. BUuen-w. b. 8o — 95.
> Blanohabd Ann, de$ 8c. not. 30 S^rie, Zool. Vol. XI. pp. 113, 114^ Cuvieb
^dU, iUiu. Zooph, pi. 36. fig. id.
* MsHLis Ofmrvat. anat. de DiaUmate; DiSBiNO in AmpMitoma gigawtewn, Wiener
Amuden i. p. 146, Tab. xxn. fig. 16, kc.
* OwBN TranMct. of the Zool. Soc. i. pi. 41. fig. 13 ; Todd Cfytlop. 1. L p. 130,
fig. 78 in PenUutama UenuMes; Dibbino Wiener AnnaL. i. Tab. il. figs. ^, 19, in
PenUuL proboeeideum.
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INTESTINAL WORMS. 177
the exception of those of touch about the mouth in certain thread
and suctorial worms, such as threads, papillae, or tentacles, which
can he extended by eversion.
A special muscnlar system is usually present, but the muscular
fibres are not always united into bundles to form distinct muscles.
In the suctorial worms the muscular fibres in their entire course are
completely imited and interwoven with the rest of the mass of the
body^. In the tape-worms fibres are visible beneath the skin
running longitudinally. In the thread and thomheaded-worms two
layers of muscular fibres may be distinguished beneath the skin,
the one longitudinal, the other transverse. The proboscis of Echi-
norhynchus has special muscles for its inversion and production.
The Trematodes make use of their suckers, especially the pos-
terior, for fixing themselves. Other intestinal worms have spines
or hooks for that purpose, which are movable, and often possess
considerable hardness. It is probable that these hooks cause by
their irritation an increased afflux of fluids, whereby the nutrition
of the worm within the animal that harbours it is £Eicilitated.
Entozoa are inhabitants of very different classes of animals; the
most numerous are the species which occur in birds and fishes.
Amongst the tape-worms especially are genera, which are found
exclusively or principally in fishes ; nearly all the species of the
genus Bothriocephalus occur in fishes; of Tcsnia, on the other
hand, more than half the numerous species occur in birds, very
many in mammals, in fishes very few. Of the genera Distoma and
EcJiinorhynchvs so rich in species, by far the most of these are found
in fishes and birds. The genus Cysticercus appears to occur almost
exclusively in mammals, as also Echinococcus and Ccenurus. Of
the genus Ascarut all the classes of Vertebrates have many species;
of Strongylus especially the mammals. The four genera, Distoma^
Tcemay Ascaris and Echinorhynchtis, contain the greatest number of
the known species of intestinal worms, especially the three first.
Entozoa live in all parts of the body, but mostly on mucous mem-
branes, in the intestinal canal and the lungs. Even in the heart
and the blood-vessels some species occur.
^ See D1X8INO4 op. dt. Tab. xxi. figs. 4 — 8 in Amphittoma gigaiUeum,
VOL. I. 12
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SYSTEMATIC
AKKANGEMENT OF ENTOZOA.
CLASS V,
ENTOZOA.
Animals mostly elongate, without peculiar organs of respira-
tion, parasitic, occurring in various internal parts of other living
animals, sometimes destitute of nervous system, or having a nervous
ring surrounding the mouth and a single ventral or double lateral
cord.
Order I. Sterehniniha s. Parenchymatasa,
Intestinal canal wanting in some, vascular in others, surrounded
hy the parenchyme of the body.
Family !• CesUndea. Body elongate, depressed, soft, conti-
nuous or articulate. Mouth none; head usually furnished with
fossettes or suctorial oscules. All the individuals hermaphrodite.
GaryophyUoBua Gmel., Caryophyllua Bloch. Body continuous
(not divided into segments), depressed, with head dilate, lobate or
laciniate, mutable.
8p. Oatyoph, mulabtUs Rud., FrUozoor, HiM. nai. Tab. vm. figs. i6 — 18,
BSEMBBB, Icon, Hdmimth. Tab. xi. figs, i— 8 ; this spedes liyes in fr«rii-
water fishes (Cyprinui, CobitU), CaryopkyUceat is distinguished from the
rest of the tape-worms by the oocurrenoe of the genital organs only once,
whilst in the rest^ on the contrary, they occur repeatedly behind each other
in the length of the body.
Ltgula Bloch. Body continuous, depressed, extremely long,
in the imperfect state with neither head nor genitals conspicuous,
with a median or two lateral furrows nmning lengthwise ; in the
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ENTOZOA. 179
developed state with head fturnished on both sides with an ex-
tremely simple fossette, and with single or double rows of ovaries.
Sp. Liffula sifRpUcimfna Bbsmb., Icon. Hdminik, Tab. xn. fig. i ; in
different species of fresh-water fishes in the abdominal cavity (Patciola
iniesHnalis Jm, Der Jiiemenwilrm, Fiaekrieme^ Strap-worm, It is difficult
in this state to distinguish the species: in Oifprinua carataiui, C&IPUN
found a species with two longitudinal streaks, Ligvla digramma. The
Ligvke met with in birds have one row or two of gemtal organs : Liffvla
uniaerialit RuD., Entoeoor. Hid. not. Tab. ix. fig. i. [In £tct Lig. gimpli-
eimma of fishes is the undeveloped state of Lig. sparta or Lig. terialit of
water-fowls. VoK SiSBOLD Band u. Bkuen-wi^rmer, s. 41.]
Bothrtocephalus EuD. Body elongate, depressed, articulate.
Head somewhat tumid, oval or sub-quadrangular, with two or four
opposite hoihria or fossettes.
Ck>mp. F. 8. LxuCKAST, Zoologi$ehe Brudulttkke i. Hefanstadt> 18 19,
4to. mit 1 Kupfert. D. F. Ebchsicht Anatomiiehrphgnolog, Untenueh'
ungen fiber die Bolhrioc^Jialen, mit 3 Kupfer. 1840, 4to. (a reprint from
the Act. Leop. Carol. Vol. xix. Supplem.)
Sp. Bothriocephahu laiut {Tcmia lata of Authors). Brxmsxr Ueb, leb.
W&rmer, Tib. n. f. i — la, the hroad Tape-worm; this species lives in the
small intestines of man^ and attains sometimes a length of twenty feet ; it
is especially met with in Russia and Switzerland, in Germany and Holland
less frequently. Gomp. below, on T(mia tolium,
Bothriocephal. punctatus Bud., Lsuckast, ZoqI, BnichdOche I. Tab. i.
f. j6, Tab. n. fig. 40, Ebchb. 1. 1. Tab. m. figs. 18 — ^8 ; in the Turbot and
other species of Pleuroneetet, and in other marine fishes, especially in
Cottus Korpio. The several joints are multiplied by transverse partition,
just as a multiplication of individuals takes place in Nait by growth. The
transverse partition commences before the genital organs are developed.
The young animals consist of a head and a small number only of joints.
Probably each animal performs annually a determinate circuit of develop-
ment. When it has cast off its joints mature and full of eggs in the
sunmier or autumn, new joints begin to grow ; in winter no eggs are found,
and even occasionally in large individuals no developed genitals. Probably
a similar renewal of the animal occurs also in BothriocepJudus latut (and in
Tmiia), when the new joints are developed in the part that succeeds the
head, the so-called neck, which becomes marked off in joints. In this
way may be explained what Esohbioht observed in a sufferer from Boikr.
IcUuB, that amongst the pieces cast off, the subsequent piece did not fit on
to that which had preceded, but on the contrary was similar to it ; narrower
and more imperfect joints had in the meantime been developed into broader.
Sub"gen. Schistocephahu Cbepl. Head triangular, obtuse, bifid at
the extremity.
Sp. Schidocephaliu dimorphus, Bothriocephaltu solidtis, Tania gasteroitei
Abildgard, Shrirter of naturh. SeUkabd i. 1790, Tab. v. fig. I, Bbbmbkr,
12—2
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180 CTASS V.
lean, Hdm. Tab. xin. f. lo, x i ; Lbuokabt^ L I. Tab. n. fig. 17, (m the abdomi-
nal cavity (not in, but on the outdde of the intestinal canal) in QoMerotUnu,
In water-fowls this worm changes its form and is then named Bothrioceph.
nodomu: [its joints and genital organs become gradually developed in the
intestinal canal of its new host» which had swallowed and digested the
fftuterotteut. See Yon Subold, Band «. Blatm^wQrmer, s. 40.]
Sub-geiL TricBTiopluynis Bub. JointB sub-indistinct; head bila-
biate, armed on both sides with two tricuspid hooks (rpiatpa, tridens).
See figures in Leuck. L L Tab. 11. £ 34 — 36, Bbeicseb Icon. Helm.
Tab.m, £4—16.
Note, — Qenus Scolex Muell. appears to be founded on imperfect
species of BtOhrwcephaU, The body is depressed, continuous as
in Ligulch The head supplied with four fossettes. It is found in
marine fishes, especially of the genus Plewronectes,
Tcenia L. (exclusive of many species). Body elongate, de-
pressed, articulate. Head with four suctorial oscules, and mostly
with a Tostellum median, imperforate, retractile, armed most fre-
quently with a coronet of booklets, especially in the young state.
Sp. Tania aoliwm L. Qn part), Bremssb, Ueb. leb. WUrm. Tab. m. f. i— 14,
GUIS., leonogr,, Zooph. PL 12, f. 2, Der Kv/rbmrtlrm, der KeltenwUrtn,
Ver foUtotrs, U Tcenia d longB antneamx. It is a mistake, that in jthe same
person only one worm of this species is invariably to be met with, as the
French name ver solitaire indicates. This species Hycs in the small
intestine, and is in Holland, Grermany, and England, the ordinary and
perhaps the only species of this family which oocors in the hiunan body.
Only very few cases are known where a person had both a Tcenia toliwn
and a Sothrioc^phahis latius\ These two tape-worms are distinguished not
by the head alone, but also by the greater or less breadth of the body. In
Tcenia tolivm the middlemost joints are longer than they are broad ;
in Bcthrioeephalue lotus the joints throughout the entire body have more
breadth than length, and in the middle of each joint are two apertures of
which the anterior is the larger and more readily perceptible ; from it the
penis occasionally hangs everted ; in Tcenia solium the apertures are at the
edge of the joints and alternate irregularly, i.e. they are situated sometimes
on the left, sometimes on the right side, without determinate order of
succession {foramina marginalia vagi altema). Since these worms are
often rejected in fragments alone, the knowledge of these characters is for
the Physician not without interest.
^ A case of this kind, the only one known to him, is given by Rudolfhi Orundriss
der Physiol, n. 1, s. 139, and another by W. VROLnt, Bijdragen tot de natwurh.
Wetenseh. m. 1828. BoeUcbeschouving, bl. 292.
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ENTOZOA. 181
For the anrangement of the numerous species of this gentts, Rudolfbi
Availed himself slso of the cfaarscter of the hookkts on the hesd, sad
distingnished imtrma and orsMtfa. But sinoe this chsncter is inoonsCsnt^
and many of the mersief of Budolphi have hookkts in the joimger period
of their life^, it cannot be reoommeDded fior this purpose. Among the
species oocorring in our domestic animals Taema fUetia BOD., Tcmia
tMgna Abildo., ZooL dam. Tab. no, fig. i, BuDfaiB, /eon. ffdtm. Tkb. 2Y.
fig. I, deserves to be recorded for the great siae of its tetngonal head, which
surpasses that of all other species. It Uves in the small intestine of the
horse.
Ditkrydium Bud. Unoertain genua. Ckonp. Bimoi^Bi Entotooor.
Synopa. p. 559, YAUorcisinrBS Ann. des Sc not de EKrie, n. ZooL
1844, p. 24a [Von Siebold, ibid. YoL xt. p. 201, aaya it Ib ft larval
form of a Tcmia without joints and sexual oi^gana.]
[RuDOiiPHi's first fiunilj of Eniozoa is not included in the
systematic arrangement of the daas in this edition of the Hand-
book, because it has been satisfiMstorilj proved by YoK Siebold,
Yak BjENEDBir, Dujabdih, Blanchasd, dkc., that it conaastB of
larval forms of TcenioBy usually encysted in situations unfitted for
their further development, and in which they become distended
with fluid. But from the great interest that attaches to them on
account of their occurrence in the human body as well as in that of
other vertebrates, we subjoin the description of them, with a
reference to the literature contained in the 2nd Edit of Yak deb
HoEVE3f 8 Handbook.']
Cystica. Body depressed or roundish, terminating posteriorly in
a vesicle fall of fluid and proper to individual entozoa, or common to
several. Sexnal and digestive organs none. Head furnished with
a coronet of booklets and four suctorial oscules.
Comp. on cystic worms, Ad. Tbghudi, Die BUuen-wHrmer. Sin numO'
graphiaeher Vertueh, Freibuig im Breisgau, 1837, 4to. mit 3 Kupfert.
Echtnococ(m8 RuD* Vesicle either single or enclosed in an ex-
ternal capsule formed by the organ in which it is contained. On
the interior surface are set many entozoa, extremely minute, resem-
bling a grain of sand, with body obovate.
Worms in this state have been ordinarily named Hydatids^ a name
which has also been extended to the rest of the cystic worms
indiscriminately, as well as to serous vesicles, the consequence
of a morbid nutrition, that contain no intestinal woi*ms. Laennec
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182 CLASS V.
named these pathological prodacte and BMnocoeeua also Acephalo-
cystea.
True echinococd propagate themselves hj means of cells or vesicles
within the parent vesicle. This last consists of several concentric,
thin, albuminous layers ; see Yon Siebold's figure in Yogel Icon,
EistologicB paifiologicoB, Tab. xiL fig. 11. Frequently the worms die
in the living body and the vesicles are changed into a gelatinous,
yellow-green mass.
Comp. Rendtobff de Hydatidihu$ in corpore humane, pneserHm in
certbro repertia, Berolini, i%2i, 8to. ; Kuhk RecherekeB aur la AcSphaJUh
cyateB, M6m. de la Soe, dPHist. not. de SlroAourg, i. 2, (1833) ; ^^ trans-
ferred to the Ann, det Sc, not, Tom. xxix. pp. 173 — 300. (The Author
distinguishes AcepluUocysUa endogena and A. exogena; some Echinococd
appeared to multiply themselves by forming new yesicles on the outside of
the parent vesicles ; such vesides, he says, occur, especially in the sheep, in
the lungs and liver ; VoN SiBBOLD has not remarked this mode of propaga-
tion ; WiBOM. and Ebiohsok's AnMv /. NalurgcMch, 1845, 2 Bd. s. 241.)
Glugb Note 8wr la atructure nUcrotcopique de$ Hydatidea, BuUd. de
VAcad. roycUe de JBnmeUea, 4 Nov. 1838, Ann. dea Se, naL ae S^rie,
Tom. vm. Zool, pp. 314 — 317.
Sp. Echinococctta erratieua mihi, Eehinoeoecua veterinorum Run., Hiat, not.
Entoi. Tab. xi. fig. 4, Bbsksbb, Icon. Hdminih, Tab. xvm. figs. 3 — 13;
in many domestic animals, especially in their liver. The Echinococeua
hominia Run., is probably not a different spedes from this. It has been
met with in the abdominal cavity, in the liver, in the heart, also in the
voluntary musdes, and in the cavities of the brain {ventriaUi cerd>n),
Ccenurus KuD. Vesicle single, on which are seated several
worms, retractile, depressed, rugose.
Sp. Comurua cerebrcUia Run., Bial. not. Eniozoor. Tab. xi. fig. 3, Bbemseb,
Icon. Jffdminth. Tab. xvin. figs, i, 7. There is only one species known,
which occurs in the brain and spinal cord of sheep, occasionally also in
cattle, in a species of antelope, and according to Roubsbau in rabbits.
The Vertigo of aheep (le toumia, daa Drehen) is a consequence of these
worms ; the symptoms vary according to the situation occupied by the
worms ; the general characters of the disease are, that the sheep at first are
somnolent, then &11 into convulsions, run up and down and die of exhaus-
tion. The vesicles, filled with water, by their expansion compress the
brain and distend the ventricles, and this sometimes to such an extent, that
the bones of the skull are affected and become extremely thin.
Gyaticercua RuD. Worm solitary with depressed and roundish
body passing into a caudal vesicle. Another vesicle external, in-
cluding the worm.
Sp. Cyaticercua edhdotcB Run.. Hydatia finna Blumenb. Abb. naturhiat.
Oegenatdnde, Tab. 39 (copied in Gui^ Iconogr. Zooph. PL 13, fig. 5),
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ENTOZOA. 183
Brkmbsb Ueb, Ub, WUrm, Tab. iv. figs. i8 — 46. In man, in oxen,
and espedally in domesticated swine, in which this worm sometimes occurs
in great numbers, and even in the heart and eyes (SflUCMlBBUro found this
species once in the anterior chamber of the eye in man : since then it has
been met with a few times in the eo^juncUva) ; mostly in the muscles of
▼olnntary motion, sometimes in the brain ^.
Chfslicercus faaeiolaris RuD. EnUn. ffiat, nai. Tab. xi. fig. i, Bbbmbbs
Icon. Tib. rvii. figs. 3 — 9 ; this species lives in the liver of rodents,
especially of mice and rats. The jointed body is very long and the vesicle
at its extremity small, so that the entire worm has the appearance of
a Taenia. It has been surmised, that the Tania cramcMs of cats proceeds
from the cffgtuxrctu /(uciolaris of rats.
Note. — The body which Sulzeb described as an ErUozoon and
named IHtrctchyceras rude (IHceras Rubolphi), preyiously placed
amongst the cystica and then amongst the anthocephala, is nothing
else than a carpel of Moras nigra, macerated and deprived of its
colour by the action of digestion.
Tetrarhynchus B.UD. ( Gymnorhynchtis ejosd. ; Anihocephalus
ejusd. ; Flcricepa Cuv. ; Rhyncobothriua Blainv., Dujard.) Head
bilobed, emitting four uncinate proboscides.
a) With body articulate. (Species of BothriocephctltbS Bud.,
Bhyneobotkriua Dujard.
Sp. Tdrarhynchut paleaeetu, Bcthrioc. iMcep% LsuoK. op. cit. Tab. i. fig. i ;
Tdrarh. hicolor, Bcthrioc. bicolor KoBDM. Microgr. BeUr. i. Tab. vn.
figs. 6—10, &c.
h) With body continuous, elongate (Vymrwrhynchva RuD.)
Sp. Tetrarhynchut reptans, ScciUx gigas CuY., Bbehs. Icon. Hdm. Tab. XI.
figs. 10 — 13, Tab. xvn. figs, i, 1.
c) With body short, clavate, supplied with two bipartite both/ria
{Tetrarhynchus RuD.)
Sp. Tararhynchtu megaeephalut Bud. Entoxoor, Syn. Tab. n. figs. 7, 8 ;
TOrarh. diicophortu Bbbms. Icon. Hdminih. Tab. xi. figs. 14, 15, &c.
d) With body terminated posteriorly by a bladder, and in-
cluded in a cyst {ArUhocephaikLs RuD., Floriceps Cuv.)
1^. Anithoceph. dongaku Our. R. Ani. {<6dit, I.) PI. xv. figs, i, 2, Bud. ErUO"
zoor. Syn. Tab. ni. figs. 12 — 17, Bothrioeeph. pcOulua Lbuok. LI. Tab. 11.
figs. 29, 30. Lives in the mesentery of AgathorUem mola.
^ Comp. I. C. Stbdybuoh, De Tcenta hydatigena anomdla. Erlangie, 1801, 8.
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184 CLASS V*
[Note, — The different species of T^cvrhynchus are, according to
Y. SiEBOLD, nothing else than imperfect and sexless forms of tape-
worms, which in the perfect and developed condition belong to the
genus RhyTicchothriua of Rudolphi. These last are found only in
the intestinal canal of Bays and Sharks. The embryos of Bhynco-
hothrii make use of the bodies of other marine animals on which
Bays and Sharks feed, as a temporary residence, until they attain
to the intestinal canal of the latter by being swallowed together
with their host. Such animals are Flat-fish, the Turbot, Barbel,
Haddock, Gurnard, Conger-eel, Sepia ; in all of which young Tetror
rhynchi have been found. That in such situations they find only
a temporary abode is proved by their being so frequently encysted,
and by their restless state, for they perforate the flesh, the walls of
the stomach, and the substance of the difierent organs, digging and
boring with their four proboscides that can be forced in and out,
and are covered with innumerable recurved booklets. Von Siebold
Band und Blasen-tounner, pp. 43, 44.]
Family II. Acanthocephala. Body utricular, roundish, marked
with transverse rugae. Mouth none ; probosciB retractile, covered
with recurved hooks. Sexes distinct.
Uchinorhynckus MuELL,, EuD.
Spiny-anouted worm. This genus (the only one of the family
and of which the characters, therefore, coincide with those of the
family) abounds in species which occur especially in the intestinal
canal of vertebrate animals (mostly of birds and fishes).
The largest species lives in the small intesiiiie of the wild and tame hog,
Echinorh. gigaa Gloqubt AncBt, de$ vers inlestinavXf PL 5 — 8, Brimskb
Icon. Hdm, Tab. vi. figs, i — 4. See for the anatomy Cloqubt's work ;
comp. Busow Echinorkifnchi Hrumosi Anatome, Dm, Zootam, Begiomonti,
1836, 8vo. This Echinorh, Btrwmotua lives in the small intestine of
different species of Phoca,
Family HI. Trematoda. Body depressed or roundish, soil.
Suctorial pores. Mouth distinct; nutrient canal divided, mostly
ramose. All the individuals hermaphrodite.
Distoma Retz., Zed. {Fasciola L. in part). Body soft, depressed
or roundish. Two suctorial acetabula ; one terminal anterior, with
perforated base leading to the mouth, the other ventral, situated not
far from the former, impervious.
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ENTOZOA. 185
Sp. DUloma kepaHatm, and DitUma laneeolatum MiHL. PoBciola k^paiiea L.,
Bbembeb Ud>. Ub, Wirmer, Tib. vr. figs. 1 1 — 14, Mbhub Obaervai. anat.
de JHdomate hepatico H lameeolato, Gk>ttmg»y 1835, foL ; the Uver^worm,
JUJx, la douve, Ltherwwrm, Sckac^fwurm, oocutb in the gall-bladder of
man, but more frequently in ruminating animalB, the ox, the deer, and
especially the aheep. (MsHUS has shewn that here two species have been
confounded ; the worm figured by Brkmbbb, op. dt., is Dittama laneeolatum.)
DuU, globiporum Bud. in different species of the genus Cfj/prinus. Gomp.
H. BuBMBiSTEB in WaaMAJB[ii*6 Arckiv. 1835, n. s. 187 ; Y. Subold, ibid,
1836, I. s. 917—133, T^b. Yl. ; Ditt. appendiculaium Rin>. EnUm. Hid, %ai.
Tab. Y. f. I, 4 ; Matbb Bekr&ge zur AruU. der BtUoz. 1841, pp. 18, 19; in
the intestines of Clupea aloaa, &c.
Dutotna BerogsWihL, in Ebichson'b Arehiv. 1844, s. 343—345, Taf. 10,
figs. 10—13.
Dtplostomum NoRDM. Body soft, depressed, oval or roundish,
elongate. Month anterior, elliptic ; suctorial acetabula two ventral,
the anterior the smaller, situated nearly in the middle of the body.
(Small animals living gregariously in the eye of fishes.)
Sp. IHploat. voheru Nobdm . Mikrogr, BeUr, i. Tab. n. ; found by NosDM ANN
in the vitreous humour and in the lens of different fresh-water fishes (Perea
JluviatUis, Chidus hta, &c.) It may be the cause of a species of Cataract ;
see the figures at Tab. i. figs, i, 1.
lifote, — Genus Cercaria Muell. (comp. above, p. 171,) contains
larvBB of DistomcUcb. Similar larvae of TremcUoda have caused the
formation of other genera by authors. Here are to be referred
Bucephaltis Y. Baeb, and LeucoMoriditim Cab.
Amphigtoma RuD. {Sirigea Abildg.) Body soft, roundish.
Single anterior and posterior pore.
Comp. C. M. D1K8INO Monographie der Cfatkmgm Amphtdoma and
Diplodiscus, Ann. da Wiener Mutetuns, I, 1836, pp. 435 — 260 ; also his
Nachtrag zwr Monographie der AmphiBtomen, ibid. u. 1839, ^ ^35 — ^5^<
Genera ffohstamum Nitzsoh, DipU>discu8 Dies.
Sp. Amph. comtOwm RuD. JEnt. Hid. not. Tab. y. figs, i—y, (H<^odomum),
in the intee^uea of Charadriuspluvialia; most of the species of Amphidoma,
particularly of the sub-genus Holodonmm, live in birds ; Amph. tuhdavatum
{Diplodiscut nibelamdus Dnss.), Bbehsbb Icon. Hdm. Tab. Yin. figs. 30, 31,
Dn»iNG Wiener Awn. i. Tab. xxiv. f. 19—249 u fi^m the intestine of iZana
and £ufo; Amph. conicum Muell. NcAwforacherXTLiL Tab. in. f. ii, DiE-
SINO ]. 1. Tab. zxiii. figs. T — 4, is from the paunch of the ox and other rumi-
nating animals ; comp. Lauser De Amphidomate conico, Gryphise, 1831.
Monosioma Zed., Rud, {Festucaria Schrank, Cuv.) Body
soft, roundish or depressed. Single anterior pore, aperture inferior
or anterior.
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186 CLASS V.
Sp. MonoftomafaJba Sobmalz Tab, anatomiam eMtotiocr, iUudr. 1831, Tab. vi.,
M1E8OHIB Betekrab, und UrOermtch. der MonotUma hijugum, 1838, 4to.
In some species of the genus Fringilla this Eniozoan has been found
oocaaionally in membraneous sacs immediately beneath the skin, usuallj of
the abdomen, back, or even of the thig^ and resembling round ebistic
tum<mra of the size of a pea. In every sac lie two worms (of the length
of I J or 2 Hues, and of the same breadth) in a little fluid. Monott. plieatum
CssPLiH, Nov, Act, Acad, Ccu. Leap, Car, xiv. 9, 1839, Tab. 52, in the
BaL roatrata, &c.
Aspidogaater Y, Bazb.
Tnstoma Cuv., KuD. {Capsala BOSC, Phyllim Oken, Nitzschia
V. Baer). Body depressed. Mouth anterior between two acetabula
simple, marginal ; third acetabulum posterior, large, circular, and
marked internally with projecting lines ; these are disposed fre-
quently in the form of a wheel or a star.
Comp. D1E8IKG Nov, Act, Acad, Leop. Car, Tom. xvin. i. Tab. i. (and
in French M&nographie du ffcnre Trittoma, Ann, det Sc, not. ie S^rie, ex.
Zool. 1838, pp. 77 — 89, PL l) These worms live on the gills or on the
skin of different fishes, and thus are not entozoa in the proper sense of the
term, but rather external parasites. Sp. Tridoma coccineum Cuv; Jt, ArU,
181 7, PI. xv. fig. 3, RuDOLPHi Entozoor. Synopt, Tab. i. figs. 7, 8, Bbbmssb
Icon. Bdm. Tab.X. figs. 12, 13, on the gills of Ortha^oritcut mola and other
fishes. Tritl, mactdatum RuD. Voyage de LA Pebouss 17. pp. 79, 80,
PL 50, figs. 4, 5, on a species of Diodon of CaHfomia. — TrUioma kamaitum
Rathkb, Hirvdo hippoglotii Mubll., Bast. Natuurk. UUtp. n. Tab. Yin.
fkg. XI. p. 154, Zocl, danica, Tab. 54, figs, i — \ {ynfri^, G. Johnston Ann.
of Nai, HiM. I. 1838, p. 431, PL xv. figs, i — 3, Rathkb Nov, Act. Acad.
Leop, Car, IX, 1843, Beiirage tur Fauna Noneegen's, pp. a 38 — 24a,
Tab. XII. figs. 9 — 1 1 ; on the Halibut, &o.
Polystoma RuD. Body roundish or depressed, narrowed for-
wards, with terminal mouth ; in the posterior dilated portion fur-
nished with six acetabula muscular, supported by horny parts,
uncinate.
Genera ffexaootyle De la Roche, ffeocabothrvum NoBDMAim.
Sp. Polyttoma integerrimum Bjtd., Bbems. Icon, Hdminih, Tab. x. figs. 25,
26 ; the urinary bladder of the frog, &c.
Note. — Genus Diplobothrium Leuck. (Sp. Diplohothr, amuUum in
the gills of Acipenaer stellatus) is said to differ from Polystoma by
its six (mterioT acetabtUa; comp. Leuckabt Zoologische BruchetUcke
III. Freiburg, 1842, 4to. pp. 13 — 18, Tab. i. fig. 6. Nordmann con-
siders these acetabula to be posterior, and does not separate the
species from the Polystomata, but calls it ffexacotyle elegans; La
Marcr ffiat not. des ani a, v. 2e 6dit. iii. 1840, p. 600.
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ENTOZOA. 187
Octobotkrtum Leuck. {McLzocraes Herm., OctOBUma Kuhn).
Body soft, elongate, depieBsed, fonushed posteriorly on each side
with four bivalve acetabula. Mouth anterior, simple. (Mostly two
anterior acetabula lateral, small.)
Comp. HsBMANN Naturform^er xvn. 178a, pp. 180 — 181, Tab. iv.
figs. 13 — 15 ; Leuckabt Brevet animal. Deter. Heidelb. 1828^ p. 18, Zool,
BruehMUeke ni. 1842, pp. 18 — 33, Kuhn Deteripticn d*un nouveau genre
de Vordre det Douvet, Mim. du MutSum xvm. 1829, pp. 357 — 362. PL 1 7 biB.
These species live od the gills of fishes. The most commoa is the species
that lives on the shad {Clupea alota L.) : Octobothrium lanceolatum Leuck.,
Brevet anim. Deter. Tib. i, fig. 7 a, 5, Kuhn Mim. da Mut. 1. 1. figs, i — 3,
Matsb BeUr. zwr Anat. der Bnioz. pp. 19 — 25, Tab. ui. figs. i. — x.
Diplozoon NoRDM. Body cruciate, as though formed of two
worms adhering together. Posteriorly four prehensile organs (suc-
torial acetabula) adhere to each limb on both sides, set upon a
common disc.
Sp. Diplotoon paradamtm NoBDif. Mikrogr. BeU. i. Tab. v. vi. (and Ann,
det Se. not. Tom. xxx. PL 20). This singular animal was discovered by
KoBOMANN on the gills of the Bream {Cyprinut Irama) ; it is 3 — 5 lines
long, and presents a body as if two specimens of Octobothrium had grown
together in the middle, like the Siamese twins. Other obseryers also have
met with this animal on the gills of other species of the genus Oyprinut.
DuJASDiN found very small entoeoa on the gills which resembled a half
Diplozoon, and formed thereof the genus Diporpa; he leaves it undeter-
mined whether they are young and separate individuals of Diplozoon.
[This question has been determined in the affirmative by V. Sixbold. He
discovered in the middle of the posterior portion of the body two slender
booklets which had been overlooked by Dujardht in Diporpa and by
NoBDMAKN in Diplozoon : they are bent back at an acute angle. Diporpa
is without sex, and always much smaller than Diplozoon; it has, moreover,
behind the middle of the body, at that part where the two bodies of
Diplozoon coalesce, a sucker. The prehensile organs are much simpler in
Diporpa than in Diplozoon; but SiXBOLD found instances of every inter-
mediate stage of complexity in them in different pairs of Diporpa which
had coalesced, so that in some the resemblance to Diplozoon was in aU
respects exact. After this conjugation or copulation, the generative organs
appear in the united individuals, and eggs are produced. See 0. Th. V.
Sebbold Ueber die Conjugation det ZHplozoon paradoxum, nebtt Bemerhungen
ud>er die Conjugatumt-Proeett der Protozoen, In Zeittch. fUr Wittenteh,
Zoologie, in. 185 1, pp. 6a— 68.]
The motion of fluid which Nobdm ANN thought he perceived in the
vessels and their branches (in each half of the animal there are on each side
two principal stems) is according to later investigations to be ascribed to
vibratile dlia which exist on the inner surface of these vessels and produce
the appearance of a very rapid current. (Ehbenbxbo, Wieghann's
Arekiv. 1835, ii. s. 118, Mayer Beitr» z. Anat, der Entoz. s. 93, 14.
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188 CLASS v.
Note, — Doubtful genera: Gyrodactyhia Nobdil, Mikrogr. Beitr,
I. p. 195, Hedocotyhia Cuv. Arm. dee So. not Tom. xtiil 1829, p.
147, Tab. XL A, Fhcsnicu/nu Rud. {Vertunmtts Otto, Nov. Act,
Acad. Leop. Car. xl 2, p. 294, Tab. xll fig. 1). Comp. Dujabdin
Hist Nat des Hdminthes, pp. 480—482, and 640.
Order II. CoRlelmtntha s. Utrtctdaria.
Entozoa with nutrient canal suspended in a distinct abdominal
cavity, supplied with mouth and anus. Sexes distinct.
Family IV. Nematotdea. Body round, elastic, often attenuated,
filiform.
Phalanx I. Acanthotheca DiESiNG. Mouth inferior between
two pores on each side which emit a single or double booklet.
Body roundish or depressed, transyersely annulate.
Pentastoma BuD., Linguatula Frcelich, Lam.
The worms of this genus were arranged by Rudolphi with the
TrerruUcda, but they differ from these by their internal structure ;
in external form some species resemble the Geatoidea; they form a
small group which ought to be separated from the pi*oper Nematoidea,
but still belongs to the Codebnintha. Comp. on this genus G. M.
DiESiNG VeraiLch einer Monographie der GaUung Fentaatom^jt, An-
nalen des Wiener Museums l 1835, & 1 — 32, Tab. l — iv.
Sp. PenUutoma tcmioldeB RuD., LinguaMa toenioldet tiAM., Cuv., OwsK,
RUD. JSrUozoor. Hid. naJt. Tab. xii. figs. 8 — 1 1, Bbeksbb Icon. Hdm. T^b. z.
figs. 14—16, DuESiNa L 1. Tab. m. figs. 1—5, Owsn Tra/M. of Zocl. 80c. i.
4. 1835, pp. 3^5 — 330. PL 4, f. 10 — 16, MiRAM, Beilrag zu einer Anaiomie
de$ Pent, keniotd. Nov. Act. Acad. Ocn. Leop. Car. Tom. xvn. 1835, pp.
623 — 646, Tab. 46, Ann. dei Sc. nai. «• B^rie, Tom. vi. 1836, Zool. p. 135,
PL 8 ; in the frontal Binus of the dog and the wolf, also in the laiynx of
these animals, and, according to some observations, in the frontal sinuses
of the horse and the ass; the male is four times smaller than the female,
whicb attuns a length of three inches and more.
PenUutoma monUiforme Diebino L 1. Tab. it. figs. 11 — 13 ; in the lungs
of the Indian serpent (Python).
The name Pentcutoma is to be rejected, because the four lateral openings
near the mouth are not mouths, and because by its lesemblanoe to similar
names of genera of Trematoda it may easily mislead to the idea of an
union with this division. It is, however, so generally received, that it can
scarcely be altered without needless confusion. The name LingucUtUa of
Frcelich as the older would deserve the preference, but it applies properly
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ENTOZOA, 189
only to some species from MmnniJili^ wMch ha^e a flat tongue-like form,
and especially to that found in the lungs of the hare by Fbouoh, and not
since re-discovered, LrngtiaiuUt temia. See J. A. Frcbuch, BetckteOmng
einiger neuer BinffewetdewHrmer, Natwffoneker zxnr. 1789. a. 148 — 150.
Tab. IV. f. 14, 15.
[T. D. ScHUBSBT (Letter to Y. Sisbold Zekickr./. vntMemtho^ zool, iy.
1851. s. 117, 1x8) concludes from his observations on the development of
Penioiloma in the egg that it ought to be placed amongst the Aearina or
Jjemteacea. The embiyo has two booklets at the anterior part» or head,
two pairs of lateral appendages or feet, each foot furnished with two daws,
and a tail as long as the rest of the body, into which the intestinal canal
is continued.]
Phalanx 11. Strongyldidea nob. Mouth terminal, or sub-ter-
minal anterior, not surrounded with retractile hooks. Body round,
elongate, elastic.
a) Mouth anl^or, not terminal.
Rictuhria Froblich, Dujard.
Comp. Fboeuch Naturfortcher xxix. 1803. s. 9. Tab. i. f. 1^3; Du-
JABDur ffdminth. p. aSo.
Ophiostoma BuD.
Genera Bochmius, Dctcnilea Duj.
b) Mouth anterior, terminaL
CucuUanus MuELL. Body elongate, posteriorly attenuated.
Head broad, with bivalve apparatus for manducation. Mouth a
longitudinal, vertical fissure.
Sp. CuevUanut eUffam Zedsb, Kud., EiUm. HiM. not. Tab. m. figs. 1—3,
Bbbmsbb Icon. ffdm. Tab. n. figs. 10—14 ; in the intestinal canal, the
stomach and the pyloric appendages of the perch and other fresh-water
fishes ; almost all the other species of this genus live likewise in the intes-
tinal canal of fishes.
Eeierochetliu DiESlNG.
Strongylus MuELL. Body round, sometimes filiform, very long,
acuminated anteriorly. Mouth orbicular or triangular. Apex of
the tail terminated in the male by a bursa emitting a double or
single penis.
Sp. ArmgyUu ffigat RuD. Bntozoor, ffiM. not. Tab. ii. figs, i — 4 ; Bbehsbb
l^df, Ub. Wiirm. Tab. 3-— 5, in the kidneys of man and of different mam-
malia ; this worm can attain the length of more than a foot, the female of
three feet. The colour is red, as in many other species of this genus.
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190 CLASS V.
Genera FseudalifM Dujabd., Sderostoma Duj., Stenurus Dujabd.
Sp. StrongyL i^fiexus, Stenunu injUxnu Duj. ; found by W. Vbouk in large
numbers in the pulmonary arteries and veins of the Ddphiniu phocoena.
See Bijdragm M de naiuurk. Wdentch, I. 1836. bl. 77—84.
StepJianurua DiESlNG.
Spiroptera RuD. (and Physahptera RuD.) Body attenuated an-
teriorly or at both ends. Mouth orbicular, sometimes surrounded
by papillae. Tail of male mostly rolled spirally or deflected, sup-
plied with a lateral expansion or bladder inferior, not terminal ; tail
of female conical, straight.
Most of the species of this genus live in mammals and birds between
the ooats of the oesophagus or of the stomach. Sp. Spiroptera Mrongylina
Bud., Bbsmbsb lean. HdmirUk. Tab. n. figs. 15 — iS^in the wild and tame
hog; — Spiropl. strumoaa, Ascaris ttrumota Fbcklich, Natv/rfortcher xxv.
Tab. ni. fig. 15, C. L. Nitzboh Spiropteroe etrunOt, Detcriptio. Halee, 1819.
4to, cum Tabula ; in the stomach of the mole, &c.
DispJiarctgus Duj. (Species of Spiroptera Run.) Head terminated
by two papillee surroimding the mouth.
Sp. Spiropt. cytUdieola RuD., Cyttidicola G. Fischeb, Reil's Archiv. in.
1799. s. 95 — 100. Tab. II. ; in the swim-bladder of trout.
Odontobius EoussEL DE Vauzi&me. (Is this its place ?)
Ascarts L. (in part), RuD. {Ascaria and Heterakis Duj.) Body
acuminate at each extremity. Head trivalved. Male genital
organ a double spiculimi.
Most of the species live in the intestinal canal of vertebrate animals.
Sp. Atcaria lumhricoidet h., A. Valisnisbi Opere finco-mediche, Venezia,
*733» !• PP- *7i — «8^' Tab. 34, 35, Bbembeb Ueb. Ub. Wurmer, Tab. i.
fig8. 13—17, Icon, ffdtn. Tab. iv. figs. 10, 11 ; round roorm, U lombriCf der
Spid'Wwrm, &c. This spedes lives in the intestinal canal of man, and
attains the length of 15 inches ; with this is ueually united a similar worm
fix>m the horse, which, however, according to Cloqubt and GURLT, differs
firom it {AtcarU megaloc^hala). Also the round* worm which occurs in the
swine, is, according to Dujabdin, specifically different (Aaearis suilia Duj.)
Comp. on the structure of the round- worm the work of Cloqubt indicated
above.
Oxyuria EuD., Bkems. Body cylindrical or fusiform, the pos-
terior part in the female attenuate, subulate. Mouth orbicular or
triangular. Penis vaginate simple, with a small posterior accessory
part. (Small worms, the females much bigger than the males.)
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ENTOZOA. 191
Sp. OoBgwru vermietUarii, Atcarii vermicuktm L., Bexmbkr Ueb, leb. WUrm.
Tab. I. figs. 6 — 13; Der Meutumrm, Sprinffwurm; it lives in the largo
intestine of man (especially in children), and causes a very troublesome
itching and occasionally various nenrous symptoms. The male was first
discovered by B&kmbkb in 1815, in a specimen sent to him by SdmraBRPfQ,
(see S. Th. v. SauooEBBuro's Leben u. Verhehr mU mnen Zeitgenosten von
B. Waoksb. Leipsig, 1844- i- o- 340) previously the much larger female
alone was known.
Tricocephalus GrOEZE. Body filiform, elongate anteriorly capil-
lary, passing, suddenly into the more ample posterior part. Male
genital organ a simple spiculum, long, vaginate.
Sp. TricocqphalmdUpar'RTn},, Bbxhsbb UA, leb. WUrtner. Tab. i. figs, i — 5 ;
this species has frequently been met with in the intestinal canal of man,
especially in the etecum, first by Mobgaoki, afterwards by BomsBBB, &c.,
in bodies of persons dying of typhus (Rokitakskt Handh. d. path. Anat.) ;
frequently in cholera-subjects in Italy by Dellb Chiaje (Itis, 1843,
p. 557).
Trichosoma RuD.
Gomp. Bud. Entozoor. Synopa. p. 13, Dujabdin Ann. de$ Se. not. le
B^rie, XX. 1843, Zoologie, p. 333, pi. 14.
Genera : Th4ym,inaCy JEucoleus, Calodium Dujasd.
Filaria MuELL. Body very long, filiform, subequal. Mouth
orbicular. Male genital organ a long spiculum with a contorted
accessory part.
Sp. FUaria medinensit, Oorditu medinemii It,, Bbembeb {7e&. M>. Wilrm,
Tab. IV. fig. I. DracunctdfU, Vena medinentie, the Aatr-«wm, ffiunea-worm,
le dragonneau, &c. This worm lives in man under the skin, in the cellular
tissue, especially in the legs, and may attain a length of ten feet ; male
individuals of this species do not seem to have been observed hitherto.
Sometimes this worm occasions severe pain ; it is met with in hot countries
especially of the old world, less frequently in America, except in the
island of Cura9ao, where it is endemic, although the worm-sickness does
not always prevul there with the same intensity. See the still interesting
notices of B. Hussem in the Vekr. van M Zeeutpsch. OmooUch. n. 1771,
443 — 464. The thread-worm is viviparous, and the young differ in form
from the mother. See Jacx)BBON and Bb Blainvillk inAnn.du MuUum,
nouvdU Sirie III. pp. 80 — 85.
Liorhynchus EuD. Body round. Head without valve, with
tubule of mouth emissile, smooth. (Doubtful genus.)
Sp. Liorhynchiu denHeulatus Bud., Bbbmb. Icon. Hdm. Tab. v. figs. 19 — 32 ;
in the stomach of Murcena angwUa.
Cheiracanthus DiES. Body annulate, posteriorly attenuate, an-
teriorly armed with palmate or dentate spinules, whicli in the middle
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192 CLASS V,
of the body are simple, in the posterior part evanescent. Head sub-
globose, beset with simple spinules. Mouth bivalved naked. Tail
of male spiral, with genital spiculum elongate, simple.
Sp. CheiraeamJth, rcbuthu Doss. Ann. des Wien. Mm. n. 1840, Tab. xv.
figs. I — 7 ; in the stomach of different species of Gat. This animal has
four long sacs near the oesophagus which recal the Umniaci of the Acan-
Ihocfephala. According to DiSBiNO the genus QncOhodoma Owen, {Pro-
ceeding* of the Zool. Society iv. 1836, pp. 113—116), a worm found in the
walls of the stomach of a tiger, is not distinguishable from this ; the mouth
however is differently described by Owen.
LecanocephaliLS DiESlNG. Body anteriorly obtuse, with head
expanded in form of a platter, and mouth trilabiate. Simple
spines surrounding the body in zones. Tail of male inflected, with
double spiculum.
Sp. Lecanoc, epinvlosm DiESiNO, Ann. des Wien. Mue. n. Tab. xiv. figs.
12 — 20.
AncyracarUhas DiESiNG. Body acuminate at both ends. Mouth
orbicular, armed with four spinules pinnatifid, disposed in a cross.
Tail of male inflected, with double spiculimi.
Sp. Aneyr. pinnaHftdus Diesing, Ann. dea Wien. Mu9. n. Tab. r7. figs.
31 — a 7 ; in the stomach and small intestine of South American tortoise.
Here also there are four long caxud sacs near the ce3ophagus, as in Ckdra-
canthm.
Note. — ^To the NemaMidea are also referred some filiform eniozoa, included
in a yeside. In the peritoneum of various fishes, between the coats of
ibe intestines and elsewhere a white worm of this sort, convoluted spirally,
is found, which Linnaus called Oordius marinus, Rddolphi FHaria
pitcium. Comp. SiflBOLD in WiEOM. Archiv. iv. 1838, pp. 305, kc.
Here also belongs a microscopic worm found by Owen in the muscles of
man, and called Trichina apiraUs. See Transact, of the Zool. Soc. 1. 4to.
1835, pp. 315 — 324, Tab. 41, figs. T — 9. Is it a Nematoid in an imperfect
state, the rest of whose fortunes are unknown T [This is V. SiEBOLD*a
opinion. The encysted Trichina is sexless and does not increase in size.
Sometimes the liver of different marine fishes is beset with cysts containing
round worms which have grown to an inch or more in length ; they have
been named Aacaria capaularia, Filaria piaciwn, &c. Siebold could never
discover in them sexual organs, but stiU they have so remarkable a
resemblance to Aacaria ostnUata, apiculigera, angulata, &c., worms with
developed sexual organs which live in the intestinal canal of the Seal, the
Cormorant, the Diver, the Gull, and predaceous fishes, that they may be
suspected to be rekted to them. Siebold believes that the encysted sexless
worms only attain their perfect development in the intestine of the verte-
brates which have swallowed their temporary hosts. VoN Siebold Band
it. Bl<iaen-tcUrm. s. 31, 33.
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APPENDIX
TO THE CLASS OF INTESTINAL WORMS.
These are certain worms which do not live in other animals,
but reside in water, or in moist earth, or in vegetable substances
undergoing acetous fermentation, and which, nevertheless, since in
form and internal structure they correspond with Ascaria, Oxyurisy
or Filaria, appear to belong to the order of thread-worms. Some
of them were by former writers arranged amongst the Infusories, as
species of the genus Vibrio. To these belong the minute animals
which LiKN^US brought together under the name of Chaos redivi-
vum, and which were described and figured by Mueller as varieties
of one species. Vibrio anguiUula {Animalcula infusoria^ pp. 63-68),
although he doubted whether they ought not to be regarded as
different species of a genus for which he had altready proposed the
name Anguilhilay by naming them AnguiUula aceti, Aug, ghuinisy
Ang. Jiuviatilis, and Ang. marina. The genus Anguilhda was
aftOTyards adopted by Ehbenbebg to distinguish these animals
from Vibrio^. DuJABDiN named the same genus Bhahditia, but
assigned to it somewhat different characters.
AnguiUula Ehrexb. [Rhahditia Duj.) Body filiform, pellucid.
Mouth round, terminal, naked. Anus before the posterior extre-
mity, sub-terminal. The male with tail naked or amplified by a
membrane (alate). External genital organ a double spiculum.
Tail of the female conical, acute.
Sp. AnguHMa aceU Gotzb Natwrfortcher xvni. Tab. m. figs. i«— 18;
DnoKS Ann, cUsSc not. ix. 1836, PI. 47, fig. 2 ; from i — 1 miUim. in sue ;
these aninialB maj be frozen without dying, whibit occaBionaUy on the other
hand a slightly increased temperature affects them mortally. Another
species, AnguiUula gluUnis, lives in sour paste (MuiLL. It^us, Tab. IX.
1 SifmboUe phffticai, Pk^ftotoa, and OrganiioUon, tjfttematiJk und geoffrapkitckn
VerMUmiM der If^fudomlhierehen, Berlin, 1830, s. 68, 105. Okkn in his Lekrb. der
Nahtrgeach, in. 1, 181 5, s. 191, places these animals under the genus Oordius, yet
in the index he keeps AnguiUula as the name of a genus, (see s. 847).
VOL. I. 13
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194 CLASS V.
figs. I — 4) ; thii is killed by vinegar. A third species that lives in the
grains of blighted ears of com may be revived, after lying dry for months
and years, by moistening, (Needham and Baueb).
Gomp. on these species Gceze Nocbirfoncher i. 1774, s. i — 53, ix. 1776,
s. 177 — i8a, xvin. 1784, s. 36 — 65, Baueb PkUos. Trana, 1823, p. i,
PL I, i, {Ann. det 8c. not. Tom. 11. 1824, pp. 154—167, PL 7, 8), Buaia
Ann. des Sc. not. Tom. ix. i8a6, pp. 115—^51, PL 47, 48.
Also in the intestinal canal of insects minute worms have been observed
and conmionly considered to be Atcarides, which belong to this division^.
In other species the mouth is provided internally with three
unciform structures or jaws. They may be included in the genus
JEnoplus DUJARD. {Enopltis, Oncholaimus DuJARD., Amhlyura
Ehrenb.?) They live in fresh and salt water.
Finally, certain small worms that live in water and in moist
earth cannot well be placed otherwise than in the neighbourhood of
the Nematotdea; they are included in the genus Oordius L. (the
Filartce excepted). They are, however, distinguished from the
Nematoids by their structure, and especially by the absence of
a posterior aperture in the intestinal canal. Dujardin and V. SiE-
BOLD have shewn that these animals in the early period of their
life live parasitically in insects.
Family Oordtacea. Body filiform, extremely slender, elastic.
Anus none ; sexes distinct.
G(yrd%u8 L. (in part). Head rotund, mouth none, or not distinct.
Tail of male bifid, of female rounded.
Sp. Qwdma aqyyaiicuA L., Bfnc^do^. Ven. PL 39, fig. i. Seven to ten inches
long, scarcely half a line thick ; comp. Chabvet Nowf. Ann. du Mus. lu.
1834, pp. 37 — 46 ; Bebthold Uib. den Ba/u des WcuBerkalbes, Gottingen,
184a, 4to ; V. SiEBOLD Eniomol. ZeUwng, 1843, s. 77, Ebiohson's Archiv.
1843, n. s. 30a— 308.
Mermis Dujard. Mouth terminal. In female the vulva for-
ward, transverse.
Comp. Ann. des Sc. not. le S^rie, Tom. xvni. 184a, pp. 119, ftc., PL 6.
^ Here, too, may be placed Oxyuria gryUo4alpce, L^K DuFOUB Ann. des Sc. not.
2e B4sne, Tom. vni. Zool. PL I. fig. 2, and perhaps the genus AnguiRina of Hammbb-
BOHiODT not described in detail {AnguiUina monUis in Aphodius conspurcatus), Okbk's
Isis, 1838, p. 318, which however more probably belongs to Mermis Dujaedin.
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CLASS VI.
WHEEL-ANIMALCULES {ROTATORIA)\
We return from the consideration of different animals whose
bodies amongst the Invertebrates may be styled large to that world,
invisible to the naked eye, with which in the class of the Infosories
we began to treat of the animal kingdom. And in the classes
that follow, however some species may be found that are scarcely
perceptible to the imassisted eye, no one of them consists entirely
of creatures so small as Infusories and Wheel-animalcules. Wheel-
animalcules, as a whole, surpass Infosories in size ; still they are
very minute animal forms, mostly between \ — ^ millimeter. Leeu-
WEKHOECK, who discovered the Infiisories, was also the first who
observed some species of Wheel-animalcules.
The name of Wheel-animalcules is borrowed from the vibratile
cilia which at the anterior extremity of the body are set upon the
margin of a disc capable of eversion and inversion. In species,
where that margin is not divided or indented, an optical illusion is
caused by the motion of the cilia, as though a toothed wheel were
revolving with great velocity in a circle, and so Leeuwenhoeck
thought such was really the case, who compared the rotatory organ
with the wheel of a watch-work". Every one who has observed
the phenomenon of vibrating cilia is aware that the deceptive
appearance of a rapid motion or current in a given direction is pro-
duced : if, then, vibrating cilia be met with on the smooth margin
of a circular structure, the appearance of a rotating wheel will
follow of course. It is to be remarked, however, that the motion is
^ See on this dass the worka refeired to (p. ^7) at the dasa of InfuMria of
MusLLBB, EHSENBKBa Bod DuJASDiN. Also may be compared O. Schmidt Veinu4:h
emtr DardeUvng der Orgamsatian der Baderthierchen, in Ebiohson's Archiv /. Natur-
yeaehiekle, 1S46, a. 67— St, Taf. m.
' Send-brieven, 17 18, vn. Brief, bl. 67. Dutboghst haa attempted to exphun the
phenomenon by mnscahur motion ; according to him the wheel ia merely a cmmlar,
mnacnlar string, which by its contraction causes other parts of the gelatinona substance
to project alternately in the form of conioal papille, whence a drcahir motion appears
to arise. Awn, du Mut^um, zx. 18 13, pp. 469 — 473.
1.3—2
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196 CLASS VI.
subjected to the will of the animal, for otherwise the vibratile cilia
would be in a constant motion, which ceases onlj on death.
The Wheel-animalcnles are capable of contraction in a remark-
able manner, many of them assuming thereby an oval form. This
fiwjulty of contraction gave occasion to the name SystolideSy by
which DUJABDIN wishes to distinguish this class of animals, but
which probably will not supersede that of Botatorta. In some the
integument is hard and rigid, so as to form a shield or a shell
{BrachionuSf Anurcea, &c.). In most there is a caudiform appendage
on the abdominal surface (Ehrenbebg names it processus ][>ediformi3
or pseudopodium), which can be drawn in and out annularly like
a telescope, and ends in a suctorial disc or in a forceps; by it the
Rotatories fix the posterior extremity of the body, whenever, being
at rest, they set the wheel-organ in motion.
The intestinal canal is straight, in by far the greatest number
of species, and the antis is found at the hinder end, at the base of
the tail. At the commencement of the intestinal canal, behind the
oral aperture, is a muscular organ of cylindrical form armed with two
lateral homy jaws. Leeuwenhoeck, Baker and Fontana took
this structure for a heart, and its motions of grasping and opening,
as the first of these authors so aptly describes them^ for the con-
traction and expansion of the heart; whereon FoNTANA expresses
his surprise that such motions should be dependent upon the will
of the animal. The lateral jaws indicate a similarity of form with
articulate animals, the insects and cmstaceans, and some writers
have even supposed that the Wheel-animalcules may be regarded
as veiy simply organised crustaceans'. On the whole, by inserting
these animals between the intestinal and the articulate worms, the
nearest affinities and natural place of the class are not indicated;
but in an arrangement that gives the classes in succession, there
must always be much that is arbitrary, for the affinities cannot be
represented by a single ascending series.
The lateral jaws present themselves under two forms. In the
greater number they consist of two pieces ; the posterior serves as
a pedicle, for the attachment of the muscles of mastication; the
anterior passes transversely inwards at a right or obtuse angle, and
^ Sevende vervolg der Brievm, Delft, 1 70a, I448te Jftsnvtf, bl. 405.
' Such WM the determination of NrrzBCH in 1844 on the genus Brachiomu,
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WHEEL-ANIMALCULES. 197
ends in a single point, or in several teeth when the part becomes
broad and indented in form of fingers. In other Rotatories the
jaws have the form of two stirrups, with the bases turned towards
each other, on which lie two or more teeth transverselj, which
arise firom the outermost arch^.
[Where the oesophagus opens into the stomach, or lower down,
are two or more oral glandular pouches, which EHRENBEBa com-
pared to the jMzncreas. The stomach is large and sacculated, and
in the saccules are large nucleated cells, or coeca are appended to
them. The cells and cceca are supposed to supplj the office of a
liver. The intestine narrower, and of variable length, but generally
short, opens into a cloaca, of which the outlet is on the dorsal sur-
&ce at the extremity of the body. But sometimes the intestine
and anal outlet are wanting, and then the residue of digestion is
returned by the mouth^ The stomach and intestine are covered
with fine vibratile cilia.
This description applies only to the females; for, in the year
1849, the veiy interesting discovery of the male of Notommata
anglica was made by Brightwell* of Norwich, and in it the entire
intestinal tract was absent ; there were neither pharynx, jaws, oeso-
phagus, nor digestive tube, and the mouth was closed.
There is no circulating system. The nutrient fluid fills the
cavity of the body, and bathes all the contained organs. The re-
spiratory organ is supposed to be represented by tortuous tubes,
which are seen at each side of the body. A highly contractile
transparent vesicle opens into the cloaca, and firom this vesicle the
tubes in question arise. To the tube on each side, minute pedi-
culated structures, various in number, with vibratile leaflets, are
attached.
The female organs consist of an ovary situated under the diges^
tive tube, generally of an oval form, or like a horse-shoe, of which
the efferent duct opens into the cloaca.
The ova are of two different kinds, summer- and winter-eggs, thin- or thick-shelled.
The summer-eggs are developed within the parent body, and the animal is then vivi-
parous. The winter-eggs have been described by Ehuenbebo, by Huxlet, and by
liKTDia, in many different species: their thicker external covering is granular, or
tuberculated, or beset with hairs. The winter-eggs are always laid, or are attached
1 See EHBBirBEBa Zur Erkenntnisa der OrgtmiaaUon in der Bicktung det Jdeiniten
», Berfin, 183a, s. 46—51, Tab. iv.
' [Daiathpu Jktcript. of on InfvMry Anim. aUUd to Notommata, PhU, Trans,
x^49> P" 333*] * [iliMio;* of Nat. Bitt, Sept. 1848.]
*
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198 CLASS VI.
to ihe mother and carried about by her {Bradtiomu, &c.); and thus, in the cold Beaaon
of the jear, these aaimala are oviparous.
Beodes the male of Notommata anfflica, discovered by Bbtohtwill, that of N.
Sieboldii has been observed by Lbtdio, who further gives reasons for believing that
SiUeropUa hydoHna Ehb. is the male of Hydatina iewta, Notommata granularis the
male of Ncftom. Brachionus, and DigUna granularis the male of Dig, catdHMa,
GossE also has ascertuned that the sexes are distinct in many others^. The males
are leas than the females, and also differ in form in most cases. The dioecious character
of the class may thus be considered to be established.
The generative organs of the males consist of a white and round
bladder or testis, filled with spermatozoa, and an efferent duct
{penis Dalrtmple) ciliated in the interior, which opens close to
the outlet of the respiratory vesicle. All the males observed are
entirely destitute of digestive tract; they possess the respiratory
organs of their species, whose fimction seems to suffice for the
maintenance of their short life, employed exclusively in impreg-
nating the females.]
The nervous system has been discovered by Ehrenberg in
different genera, and he described, as central portion, different ganglia
{ganglia cephalica seu cerebralia)^ situated close to the wheel-organ,
from which distinct nerves arise. In Hydatina aenta, according to
the investigations of the same observer, two threads also arise from
them that run downwards on the abdominal surface, and unite to form
a ganglion from which a single nervous string with many small
ganglia or swellings arise*. As organs of sense, in most of them red
eye-spots (generally two, sometimes one or three, seldom more than
four) have been perceived; sometimes these exist in young individuals
alone, and disappear on full growth, as in the genus Floscularia.
Besides the muscles of the special parts, there are found in many species thin
bundles of muscles running longitudinally, one oa the dorsal surface, one on the
abdominal surface, and two lateral.
With respect to the geographic distribution of Wheel-animalcules nothing deter-
minate can yet be specified. Only do we know, from the observations and notices of
EHBBNBEBa, that, besides Europe, they are found in northern and western Asia, in
the north of Africa and in North America. Beyond doubt they occur in all quartera
of the world. The physiological peculiarity of life suspended for a length of time, to
be again awakened by the vital stimxdus of fluid, has given a special celebrity to these
animals. On this subject we refer to what will be offered below when we notice
lUxtrftT vitlgaris, in which this phenomenon has been chiefly observed.
1 [GossB On Uu dioecious eharact, of the Rotifera, Proceedings of iks Soyal Soe.
Vol. vra. pp. 66, 68.]
* Die Infusionsthierchen, s. 416. Somewhat differently ordered is the nervous
system jp Notommaia (s. 415) and in Diglena (s. 443), but in all there Ues a principal
mass, as the coUection of nervous ganglia, on the dorsal surface of the anterior
extremity of the body.
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SYSTEMATIC
ARRANGEMENT OF ROTATORIA.
CLASS VI.
ROTATORIA.
MiCBOSCOPiC animals, contractile, crowned with vibratile cilia at
the anterior part of the body, which by their motion often resemble
a wheel revolving rapidly. Intestine distinct, terminated at one
extremity by a mouth, at the other by an anus ; generation ovipa-
rous, sometimes (periodically) viviparous.
Order Single. Botatoria.
(The characters of the class are those of the single order.)
Family I. Floscularice. Tentacles or lobes around the mouth
(with rotatory organ deeply cloven Ehrenb.), furnished with cilia.
Body affixed by a pedicle.
The hairs of this wheel-animalcule are, according to Dujabdin,
Peltier and other observers, not vibratile cilia, but are capable
individually of expansion and contraction ; Ehrenberg, who admits
that these hairs may for a long time continue at rest and be flaccid,
still maintains that they occasionally vibrate, and refers to Eichhobn
who perceived the same thing in his croton-polyp, Stephcmoceros
{BeUrdge zwr NixbMrgeach, der Jdemsten Wasaerthiere, s. 21).
Floscularia Oken, Ehrenb. Body clavate, or campanulate,
anteriorly expanded, five or six lobes sustaining a fasciculus of long
cilia. A vagina transparent, cylindrical, often covering the solitary
animal.
Sp. FUwmlaria omata Ehbbkb., Ihr F&nger Eiohhorn 1. 1. Tab. m. figs.
0 — L, p. 39 ; Ehbenb. Organisation in der Richt. des U, Raum. 3tf«r. Beitr,
Tab. vm. fig. a ; InfimonHh, Tab. xlvi. f. a ; Dujard. Tnfutoir. PI. 19,
figs. 7, &c.
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200 CLASS VI.
St^hanoceros Ehrenb. Body campanulate, surrounded by a
transparent vagina. Tentacles five around the mouth, covered with
cilia in whorls.
Sp. Stephanoeeroi EUMomii Ehbxhb., Ikr Kron-Polffp Eiohh. 1. L Tab. i.
fig. i; Ehbkitb. OrganU in d. Richl. d. Jd. Maum, ^fUr Beitrag, Tab. zi.
fig. I, Infuiurndk. Tab. zly. fig. a.
Family EL Melicerttna. Rotatory organ simple, with margin
entire or lobate. Two stapediform maxillas, with teeth transversely
incumbent. Body aflSxed by a pedicle.
Ptygura DuJARD. (Ptygura^ (EcisteSy Canochirus Ehrexb.)
Lactnularia Oken, Schweigg. [Megahtrocha Ehr. and Laci-
lunaria ejusd.) Rotatory organ large, incised on one side, hence
bilobed or reniform. Animals often social, and sometimes covered
by a gelatinous envelope.
Sp. Lacinvlaria sociaUi SoHWEioa., Hydra toeiaUs L., Brachianui gocialu
Pall., VorticeUa tocialis Mubll., Iirftuor. Tab. XLm. figs. 13 — 15, (and
Vortie, Jlotculosa Muxll. ibid, figs. 16 — io), IUbsbl, Ins. m. Supp. Tab.
94, figs. I — 6; Ehbekb. Infusumtth. Tab. xliy. fig. 4. They form
minute, white, conical bodies, which adhere to the roots of water-plants
(Lemna, CeraiophyUum, Chora, ftc), and consist of fifty or more such
wheel-animalcules, whose extremities are all directed to the centre. After
a time the young ones separate themselves from this connexion, moye
away and adhere to different plants, to form new colonies. Megahtrocka
aJho-fiamcani Ehb., Koisel Im. ni. Suppl. Tab. 95, 96, {Megalotr, alba
Ehrknb., Zur ErhenTiimsa d. Organi», in der Riekhing det Jdeinslen Raumea.
Iter Beitroff, Tab. m. f. 15, intestinal canal), Ehb. Infitsitmslh. Tab. ZLiv.
fig. 3, is distinguished from the former species by the absence of an
envelope, though united with it by former writers.
Tvbicolaria Lam. (in part), Ehrekb. Body clavate, with rota-
tory organ four-lobed, and respiratory tube double, included in a
gelatinous vagina.
Sp. Tubicdaria nqjcu Ehbbnb., Rotifer dlbo-vettUut Dxttboohet, Ann. dw
Mu8, Vol. xrz. PL 18, figs. 9, 10; Ehbskb. If^funontth. Tab. ZLV. fig. i.
Melicerta Schrank, Oken. Body clavate, with rotatory organ
four-lobed, and double respiratory tube, retractile within a vagina
conico-tubular, granulose, opaque. Two ocelli in the younger age.
Sp. MeUeeria ringent Sohbakk, Sabdla ringena L., 8y^, not. id, xn. Lexu-
wximoxcK, PhU. Transact, 1704, Vol jltv. p. 1784, figs. 3, 4 ; Sendbrieven,
Delft, 1 718, vii*. Brirf, bl. 6$, &c.; SoHuIFFEB Die Blumenpalypen der
eOaaen Waeser. Mit 3 Kupfert. Begensbui^g, 1755, 4to; Rotifer guadrieir'
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ROTATORIA- 201
cmlarU Dutboohit Ann. du Mu$, Vol. zix.'PL i8, figs. 1—8; Ehbinb.
Iirfuriondk, Tab. XLVi. fig. 3. Tliese animalculei seated in a cue that
adheres to duck-'weed, belong to the forma which were first disooyered by
LKKU VtMN UOSCK.
idmntas ScHRANK, Ehrenb. Body clavate, with rotatory
organ bilobed, and respiratory tubule none, solitary, retractile into
an opaque envelope. Ocelli two.
Sp. Xmnttu ceraiophffUi, Ehbsxb. Ifrfutiomth. Tab. zltl fig. 4.
Family III. BrachiofUBa. Animals swimming freely, covered
with a membraneous scute univalve or bivalve, furnished with rota-
tory organ double or multiple {zygotrocha or polytrocha Ehrenb.)
Pterodina Ehrenb. Shield orbicular or oblong. Botatory
organ double. Two ocelliform points. Tail cylindrical, transversely
rugose, terminated by a suctorial disc which is often ciliated.
Sp. Pterodina patina Ehbenb., Brackiowui patina MuxLL., If^fuaor. Tab. 48,
figs. 6 — 10, Ehbsnb. It^uaiomth. Tab. lxit. figs. 4, &c.
Brachtanus HiLL, MuELL. (in part). Scute urceolar, open in
front and behind, with anterior aperture or both denticulate. Rota-
tory organs two. Madllss digitate.
I. Tail aifieulate, forked at the paini.
a) With ooelliform point above the maxille. (Genus BrachionuB
Ehbxnb.)
Sp. Braehionus ureeolari$ Musll., Ii^u$ot. Tab. L. figs. 15 — -si. Ehbkkb.
Organic, in d. Rieht. det Id, Raumet, ^/tter Btitrag, Tab. ix. fig. m.,
Iirftuiomth. Tab. LXin. figs. 3, &c.
b) Without ooelliform point. (Genus Noteut Ehbxnb.)
Sp. Noteui guadricomia Ehbxnb., InfuaUmdk, Tab. mi. fig. n.
IL TaiU none, (Grenus Awwrcsa Ehbxnb.)
Sp. Braehitmiu aquamtda Muxll., Anurcsa aquamula Ehbxnb., Muxll.
Ii^fusor. Tab. 47, figs. 4—7, &c.
Lepadella BoRT (spec, of Brochionus MuELL.) Scute oval,
convex above, flattish beneath, open at both ends. Rotatory organ
divided into several lobes. Tail triarticulate, forked at the extre-
mity. Maxillae naked, terminated by a single point, or by two or
three teeth.
(Genera : Lepadella, Metopidia, Stephanops, and SquameUa Ehrenb.
Sp. Lepaddla (Stephanope Ehbxnb.) lameUaria, Braekionui lamdlarit Muxll.,
Iftfut, Tab. 47, figs. 8— II ; Ehbxnb. Jtrfusiontth. Tab. ux. figs. 13, &c.
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202 CLASS VI.
Euohlania Ehrenb. (spec, of Cercaria MuELL.)
Sp. Buchl. luna, Cercaria luna MusLL., Fureocerea luna Lam., Muell.
Iirftuor. Tab. xx. figs. 8, 9 ; 'Eebxsb, Infu^ionsth, Tab. LXii. figs. 10, &c.
Dinocharis Ehrenb.
Salpina Ehrenb.
Colurm Ehrenb.
Monura Ehrenb.
Rattulus Lam. [Mastigocerca and Monocerca Ehrenb.) Body
oval, covered with a scute carinate, narrowed posteriorly. Botatory
organ divided into several lobes. Tail styliform, long, rigid.
Ocelliform point single.
Sp. Ratttdus cariruUus Lam., Triehoda rattus MuKLL., EiOHHOBH Waiter
thiere, Tab. ii. fig. 0, die WasaerraUe MuKLL., Ivfutor, Tab. xxix. figs.
5 — 7. (Ehbenbebo distinguishes here two species and two genera:
Mastigocerca carinata Muell. I. 1, fig. 7, Infuaumsthier. Tab. Lvn. fig. 7,
which has a shell, and Monocerca rattus. Tab. XLvm. fig. y, to which
Eichhobn'b drawing and the first two figures of Mueller 1. L belong,
which wants the shell, whilst he however remarks, that both are very
similar ; Dujabdik is of opinion that only one species should be adopted.)
Rattulus Ehrenb. With two ocelliform points, and tail styli-
form, inflected. (Animal naked? Is this its place?)
Sp. JtatttUus lunaris, Trichoda htnaris Muell., I^fus, Tab. zxix. figs, i — 3,
Ehbbnb. If^usiotuth, Tab. lti. fig. i.
Polyarihra Ehrenb.
Triarthra Ehrenb.
Sp. Triarthra longiseta Ehbbnb., Eichhoen Wasserihiere, Tab. i. fig. 7,
Ehbenb., Organisat, in d, Jiicht. des Id. Baumes, $ttcr Beitrag, Tab. vui.
fig. I, Tnfiisionsth. Tab. LV. figs. 7, &c.
Family IV. Uydatincea {Furcularina DujARD.) Animals swim-
ming freely, naked, with integument contractile, flexible, often
marked by parallel rugae. Tail forked.
A. Rotatory organ single, continuous, not lobed at the margin.
{IcJithydina Ehrenb. in part).
Ichthydium Ehrenb. (species of Cercaria MuELL.) Body
smooth.
Sp. Ichihyd. podura Ehbenb., I^fusUmsth, Tab..XLin. fig. 3.
Chcetonotus Ehrenb. (species of Trichoda Muell.) Body
hairy.
Sp. ChcBtcnotus larus, Trichoda larus Muell., Irrfusor. Tab. 31, figs. 5 — 7;
Ehbenb. If\fusionsth. Tab. xliii. figs. 4, ftc.
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ROTATORIA. 203
[DuJARDiN {Injus. p. 268, 1841) gave reasons for the exclusion of
ChcBtonotua and Ichthydium firom the Rotatoria^ and placed them
provisionally in his order Infusoirea symitriques. Afterwards
C. YoGT referred them to the Turbellaria. Sohultze discovered in
the sea-sand at Guzhaven (1853) a new genus and species closely
allied in anatomical structure to our Ichthydina, which he named
Twrbanella hycdina. Neither in Ickthydivm nor Chcdtonottuf do the
dlia form a true wheel-organ round the mouth capable of protrusion
and retraction. In Ichthydium^ they are spread over the entire
abdominal surface j in Chcetonotua over the whole of the anterior half
of the abdomen, and at the margin of the posterior half form a band
which surrounds the closely set, stiff hairs, which are much finer than
the spines on the dorsal sur&ce and directed backwards to cover the
non-ciliated portion of the abdomen. The tail is forked, but not
jointed. The intestine is straight, the anal opening at the fork of
the tail. Neither vessels, nerves, nor muscles can be seen, except
that the oesophagus is muscular. The sexual organs are situated
between the intestinal canal and the integument of the back, the
testes consisting of a loose cluster of vesicles in front of the ovary.
ScHULTZE concludes that the Ichthydina of Ehrenbebg (exclusive of
his genera Ptygv/ra and Glenaphora which are true rotatories) must
be excluded from the Botaloria, and that they belong more nearly
to the TtcrbeUaria than to any other order of worms. There are
however remarkable differences of structure between them and
any &mily of TwrheUa/ria, For in those Turbellaria which have a
straight intestine with anal opening, the sexes are distinct ; whilst
in those which are hermaphrodite the intestine has no anal opening.
ScHULTZE contends however that in worms the characteristic derived
from the form of the intestinal canal is of greater systematic value
than that derived from the formation of the sexual organs : and
recommends that the Ichthydhia, limited as above, be placed pro-
visionally amongst the Microstoma of the order Turbellaria, which
will then require to be subdivided into the dioecious and the monoe-
cioius (JchthydincC). See Schultze in Mueller's Archiv. 1853.
s. 241-253. Ta£ vl]
B. Rotatory organ multilobed or parted {Uydaiincea Ehrenb.)
Oioglena Ehrenb; Maxillse none. Ocelliform points three, the
middle one sessile, the two lateral pediculate.
Enteroplea Ehrenb. Body oval, oblong, anteriorly truncate.
Maxillad none. Ocelliform points none.
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204 CLASS vr.
Sp. EnieropUa hydcAina Ehbenb. Tab. XLvn. fig. i, Dujabd. Infvaoirea,
PI. XIX. fig. a.
Hydatina Ehrenb. Body oval, anteriorly truncate. Maxillse
digitate at the extremity, terminated by teeth five in number, firee.
Ocelliform points none.
Sp. ffydatina mitct, VarticeUa tenia Muell., ImfuKT, Tab. xu. figs. 8 — 14 ;
Ehbenb. Organiaation, Sygtematik, &c. 1830, Tab. viii. TnfusUmdh. XLVn.
fig. 1 ; this is the animal in which Ehbxhbbbq first demonstrated the com-
posite structure of the Rotatoria,
Notommata Ehrenb. (in part). Body oval or oblong, posteriorly
narrower, anteriorly truncate. Maxillse digitate, with several teeth
at the extremity, Ocelliform point single, forward, dorsal, or
several points clustered.
Sp. NUommaia clavulata Ehbenb., OrffatUaation in der lUehtung d. Jd,
Baumei, 2iUer Beitrag, Tab. x. fig. i, I^funcntthierchm, Tab. L. figs. 5, &c.
OydogUna lupvs Ehbenb., /7^tuio9u<A. Tab. XYi. fig. 10, (the form of
the maxilla* not yet accurately known).
SyncJuBta Ehrenb. Body broad anteriorly. Rotatory organ
armed with styles. Ocelliform point single, anterior, dorsal.
Sp. SyvichoBta haltica Ehbenb., Infunoruth. Tab. uii. fig. 5 ; phosphorescent^
in the Baltic, &c.
Furmdarta Lam. (in part), Dujard. Maxillas forcipate, with
extremity usually imdivided and acuminate, or bidentate, protrac-
tile as far as the margin of the rotatory organ.
a) Ocelliform point none. Pleurotrocha Ehbenb.
Sp. Pleurotrocha contLricta Ehbenb., Ififimbneth, Tab. XLVin. figs, i, fto*
h) OceUiform point single. Furcfdaria and Scaridium Ehbenb.
Sp. Fwrcvlariagibha'EwQXS-R,, InfutionOh, Tab. XLViii. figs. 3, ko,—Fwrcul,
Umgicauda Lam., Trickoda longicauda Muell., Infiuor, Tab. xxxi. figs.
8 — 10, Scandium lonfficaudum Ehbenb., InftitioTuth. Tab. liy. fig. i,
with a hook in front on the wheel-organ and a very long tail, by which the
animal progresses in the water by leaps.
c) Ocelliform points two (JHgUna and JHtAemma Ehbenb. exclusive of
Dtetemma marinum ejusd.)
Sp. Furcularia forcip€Ua, Cercaria forcipata and vermicularii Muell., It^Vr
$or. Tab. xx. figs. 18—23 ; Ehbenb. Infutiwuth. Tkb. lv. fig. i, Dekinia
vermictdariB, Mobben Bijdragen tot de natmtrk. Wetensch. v. pp. 337, ftc.
d) Ocelliform points three (Triopkthalmtu, Eosphora Ehbenb.)
e) Ocelliform points numerous, disposed in two clusters (Theonu
Ehbenb.)
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ROTATORIA. 205
Lindia DuJARD.?
I^ote, — Genus AJbertia Dujard. is distrngoished by a shield in
front of the rotatory oi^gan and by a tail which is conical, short, un-
divided. Body cylindrical, elongate. It lives parasitically, in the
intestinal tnbe of earth-worms and slugs. Comp. Ann. dea Sc not,
2e s6rie, Tom. x. pi 175, Tab. ii.
Family V. PMlodinoBa Ehrenb. {Rotifera Dujard.) Animala
swimming freely or creeping (after the manner of geometric cater-
pillars). Body elongate, fusiform, contractile into a ball. Rotatory
organ double. Tail articulate, frimished posteriorly with little
hom§ or styles. The stapediform maxillas with two parallel teeth,
sometimes with three.
Rotifar Cuv.i (The characters of the family.)
o) Without proboscis.
TyphUna Ehrenb (and Hydrias ejusd. Ocelliform points none).
MonoUtbiM Ehrenb. Ocelliform points two.
h) Anterior process retractile, proboscidean.
*) Ocelliform points none.
CaUadma Ehrenr
**) Ocellifoim points two.
Philodina. The ocelliform points situated behind the proboscis.
Sp. PhUodina erythrophthalma Ehbsnb., OrganiaoHon, Syttemaiik, ke,
Berlin, 1830, Tab. vu. fig. 2; Infiuioruih, Tab. LXi. fig. 4; by former
writers confounded with Bottfer vulgarii, DuJABDUf names this species,
with which he thinks some other species of Philodina Ehhshb. ought to
be united. Rotifer inJUUua, Infusmres, PI. 17, fig. a.
Rotifer Ehrenb. (and AcAinwrus ejusd). Ocelliform points situated
at the anterior part of the proboscis.
Bp. JZMi/er vulgaris Sohbank, Ehbsnb., Purctdaria rtdtviva Lam., VortictUa
rotatoria Mubll., Infutor. Tab. xui. figs. 11 — 16; Eebbitb. Organimt,,
Sytlematik, &c. 1830. Tab. vii. fig. i, Ittfunonith. Tab. LX. fig. 4; Dujab-
MH, Ififutoiret, PL 17, fig. i.
This Wheel-animalcule was first discribed and figured by Lbiuwbkhokok
in 1701. {Sevmde vervolg der Brievm, i^iteMisBive,h\^^o^. He observed
1 Tableau iUmentaire, 1798, p. 653.
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206 CLASS VI.
thAt this animal, which he had found in a leaden gutter of his house, after
it had been dried with the sand and other matters that adhered to it,
revived again, when after two days he poured water upon it, which, having
been previously boiled, could contain no living animalcule. He afterwards
found that the same phenomenon occurred after a lapse of five months
(bl. 413). Different observers repeated these experiments. Font ana
found Wheel-animalcules that had been dried for two years and a half
revive on being moistened {TraitS aur U renin de la Vipdre I., Florence,
1 781, 4to. pp. 90', 92), and Spallanzani saw the waking from slumber
occur even after four years {Opuscules de Physique, traduits par J. Senebisb,
Geneve, 1777, u. p. 310). The last-named observer saw the same pheno-
menon many times in succession ; nay, even eleven times he saw alternately
apparent death and life. A few minutes are often sufficient to revive the
creatures ; but such alone as were surrounded with sand and other matter,
not those which lying quite bare had been dried, were revived. Observations
in the present century also are not wanting, by Dutroch£T, C. Sohultze
(according to Ehbenbebo on Philodina), and others.
This phenomenon does not stand quite alone. We have spoken above of
Anguillula (p. 194), and Sfallamzani observed the same phenomenon in a
microscopic aquatic animal, which he named Tardiffrcidef and which has
been called Arctiscon by other writers. Dujardin brings this and other
similar animals as well as the Wheel-animalcules into the same class of
Systolides; but we are of opinion that they ought rather to be placed, as
very imperfect forms of ArachnoHdea, with the Acari. It is on these iar-
digrades that Dot^bb not long ago performed his very interesting experi-
ments, and also, after most perfect desiccation of the animals when quite
uncovered, succeeded in reviving them (Ann. des 8e. not. 2« s^rie, 1841.
Tom. xviii. Zooloffie, pp. 5— 35)-
To these observations no exception can well be taken ; the fitcts must
ttther be stoutly denied or be accepted as we find them. Spallakzani
asserted incorrectly that life was quite gone, and that a real revival
occurred (1. 1, p. 3^1). Lkeuwenhoeck expressed himself more cautiously.
Bonnet too speaks of a seeming death, and says that life is not quite
extinguished (Consid. swr les corps organisis, (Euvres, Neuchatel, 1779,
8vo. VI. p. 214, Contemplation de la nature, ibid. Tom. vin. p. 762), VoN
Humboldt calls the state of apparent death in these animaLs one of sleep,
or of suspended life {Versuche iiber die gereizte Muskd-und Nerver\faser,
1797. 8vo. I. s. 196). In this desiccated state life is potentially present,
but does not announce itself by actual phenomena. If ws choose to name
this life latent, we must not call death itself a latent life; certainly these
animals are not dead, but their life is brought to a stand by the want of
one of the most common and most necessary of vital stimuli, by the want
of water.
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CLASS VII.
RINGED WORMS {ANNULATA)K
Linn JBUS placed (vid. p. 30) all animals that are destitute of a
proper intemsd skeleton in two classes, that of Insects and that of
Worms. In reviewing the principal modifications which subsequent
writers have introduced into the general classification of the animal
kingdom, we find that thej relate chiefly to the animals placed bj
LiNNiEUS in the latter class. All those classes which we have
hitherto treated of have been formed by separation from the LiN-
N.£AN class of worms ; all the animals which we shall describe in
the sequel as Molluscs formed collectively a portion of the same
great division. Amongst creatures so numerous and of such variety
of form, there are some which in type, or plan of organisation,
approximate towards insects; they are, like insects, articulate
animals, but differ firom them by the absence of articulate feet. As
early as the end of the last century Cuvier made of these a distinct
division of the animal kingdom under the name of Worms, and, at
1 Compare on this class : —
O. F. MuELLEB, Vermium. terrestrium et fiuviatilium teu AnimoUium infusoriorum,
kdminthiGirum et tedaeeorum, non marinorum, wccincta HUtoria, Haunise et Lipeie,
1773, 1774, n Volamina 4to. (This work describes the Worms of Liknaus, that is,
Uie invertebrate inarticolate animals and the ringed Worms.)
O. F. MuELLEBj NaturgtschichU einiger WurmarUn det tiUienundnlziffen Wauert,
mit Knpf., Kopenhagen, I77i> 4to. (new edit. 1800). Descriptions and observations
referring chiefly to the genera Nais, Nereis and Aphrodile.
J. G. Satiokt, Sjfdlme dee Annelidet, DeteripUon de VEgypt€y Tom. xxvi. Paris,
i8a6, (pp. 3^5—472).
AUDOUIN et MiLNB Edwabds, Clataificaiion dee Anndides et Ikecription de eeUcn,
qui kabUent lea cdtea de la Frajice, Annalet dee Sc. natur. Tom. xxvii. 1833, pp. 337 —
447, Tom. xxvra. 1833, pp. 187—147, xxix. pp. 195—269, pp. 388—411, xxx.
pp. 411— 4«5-
MlLNS EDWABoa^ Anndida in Todd'b Oydapoedia I. 1835, pp. 164—173.
A. S. Obbsied, Qrceidanda Anmdata doreibranchiata in Kongl. Daneke videneka-
hemee SeUhahe NatwrvidenMbeliffe og mathematieke AfhandUnger t. 181 3, pp. 153—
116, with figures.
A. E. Gbubs, IHe FamUien der Anneliden, Bin S^ematierhea Venuch. Berlin,
1851.
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208 CLASS VI.
the same time, added to it the Entozaa^. Some jears afterwards
CuYiER discovered, that manj of these animals have red-coloured
blood, and thought that a name implying this was justifiable {vers
h sang rouge), whilst Lamarck, on account of the rings into which
their body is divided, named them Anndides.
It was necessary to premise these historical notices in order to
make it clear why we have given to this class of animals the name
'^Binged Worms;" and, notwithstanding, include in it animals
whose body is not divided into rings. The name may be defended
by similar iostances from other classes of animals, where names do
not always suit all the iadividuals in them*. But, further, we
prefer this name to that of " Worms," because this last is too
indeterminate, and, as has been alleged above, has a double meaning.
One of the best writers on the Anrmlata is 0. F, Mueller, to
whom the whole of Zoology is indebted for distinguished services.
The immortal Pallas, also, described many ringed worms, and
investigated them anatomically. In the present century they have
been especially iuvestigated by Savignt, Milne Edwards, GtRUBE
and Oersted. Ehrenbero has placed some of these worms,
on accoimt of the vibratile cilia with which their integument is
beset, in a separate class, under the name TurbeUaria. But, besides
that we think too great a multiplicity of classes is to be avoided,
some of these TurbeUaria approach far too nearly to other natural
divisions of the Annulata to allow us to place them in a distinct
class of the animal kingdom '.
Cuvier and Lamarck placed the Annulata higher in the animal
kingdom than the rest of the articulates ; above the Crustaceans. It
is true that the last-named Zoologist considered the Crustaceans to be
the more highly organised, but believing that the AnnuIcOa ought to
stand above the Insects, and that it was inexpedient to break the con-
nected series formed by the Insects, Arachnoids, and Crustaceans,
^ Tableau OSmeiU. de VHitt. not. dea Anima/ux, 1798, p. 644 ; Le^. d^Anat. con^
parSe I. ivi^me Tableau, Lakabok adopted the same class in his Sjftt^me dee Anim,
eane vertibree, i8or, p. 315.
* The class of the Acalephce for instance, the order of the henUptera, a name which
is properly applicable to the division of the heteroptera alone.
* In the foUowing general yiew of the internal stmcture of this class we shaU
especially fix our regards on those animals which indicate most clearly the articulate
type; for the rest we refer to the special notices in the Systematic Arrangement.
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RINGED-WORMS. 209
by the AnmUatay he preferred to place these last at the top. We
are quite as ready to acknowledge that Articulates with articulate feet
form a single connected series^ and would not therefore separate
them from each other; but we place the AnnuUUa below the Insects,
not above the Crustaceans. This arrangement, formerly adopted by
us when it was less common, appears now to be generally received;
even by Frenchmen, as, for instance, Milne Edwards.
The body of ringed-worms is generally much elongated and
cylindrical; in some instances it is broader and oval. It is divided
by transverse folds into rings or girdles, which, in most species, are
very numerous, and in one and the same species may vary greatly
in number, at least when that number is very great. The common
Leech has abont 100 such, Eunice gigarUea above 400 ; in Phyllo-
doce laminoaa Sav., Audouin and Milne Edwards found nearly
500 rings, whilst in other individuals of the same species there
were sometimes only 300. The integument is always soft, not
corneous, but some of them live in sheaths or shells, sometimes
compacted with bits of shell or grains of sand into a mosaic work-
of considerable strength, and sometimes consisting of calcareous
matter, as in the genus Serpula,
In some the head is not distinct from the succeeding rings of
the body. In others it is distinguished from the trunk by its
different form, and is provided with eyes and even with threads,
which many anthors name Antennse, after the so-named parts in
Insects and Crustaceans; but they differ from these, and can be
pushed in and out like the horns or feelers on the head of snails.
The number of these feelers differs; there are rarely more than five,
and some species have only a single thread of the kind. ,
On the rings of the body spines or hairs are usually set, which
however may be entirely wanting in some, as in the leech. In
most the hairs or spines are placed upon minute lateral tubercles,
which may be considered as rudiments of feet. These rudimentary
feet are, however, never jointed as in insects. They are usually
divided into two parts, which may be named oars or fins; one on
the dorsal surfja.ce, another on the ventral siuface {rame dorsale et
rame venirale Savigny). On each of these two projections a
bundle of hairs {aetcB) is set, of very different form; and, besides this,
each projection has, as the rule, a conical spine that can be re-
tracted into its sheath and is called needle (acus). Moreover, at the
VOL. I. 14
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210 CLASS VII.
base, or foot-piece of each of these oars there is usually placed
a filiform appendage {cirrus). In the Dorsibranchiates there are
found, in addition, on the dorsal surface towards the sides and near
the oars, or upon them, the external respiratory organs, Oilis, of
various forms ; sometimes divided like a comb, or branched like a
tree, sometimes composed of simple filiform appendages resembling
the cirri of the oars. In other ringed-worms the gills are situated
at the most anterior part of the body. In the Leech, the Earth-
worm, and allied genera, no respiratoiy organs are visible externally.
In those ringed-worms that have not a distinct head, the mouth
is usually found quite at the anterior extremity of the body ; in the
rest it is situated on the inferior surfsu^, and usually a muscular
proboscis can be everted (FhyUodoce Nereisy &c,). In'these, more-
over, the mouth is ordinarily armed with homy jaws, placed late-
rally, differing in number in the different genera. Occasionally the
number is not the same on the two opposite sides. Thus the genera
CEnone and Aglaura Sav. have four jaws on the right, five on the
l^ft ; Lysidice and Leamce three on the right, and four on the left.
The intestinal canal is, for the most part, straight, yet there are
exceptions. In SabeUa ventilabrum the canal makes a great num-
ber of transverse flexures, lying upon one another, and winding
sometimes to the right^ sometimes to the left: the first portion
alone, the oesophagus, is straight^; Grube observed the same dis-
position in Cirratulus^, In Amphtctene (Amphitrite auricoma bel-
gica Cuv.) the intestinal canal makes two flexures, as in Hohihurtay
first proceeding backwards, then straight forwards, and then back-
wards again with a narrower portion between the two others ^ In
the remainder, where the canal is straight, there are usually lateral
appendages, or it is as though divided into. cells by transverse
constrictions. In the Earth-worm a short round stomach succeeds
to the oesophagus, and then there is another muscular stomach. In
Arentcola the middle wider portion of the intestinal canal has very
thin walls, and is covered with very regular vascular reticulations
1 Meckel, SyaUm der Vergl, Anal, iv. 1829, s. 71, R. Waoneb in Oken's his,
i«3«, B. 657, Tab. X. fig. 13.
' A. E. Gbube, Zw Anatomic und Physidifgie der Kiemenmirmcr, Konigsberg,
1838, 4to. 8. 34.
* Pallas, Misc. Zoolog. p. 129, Tab. ix. figs. 12, 13.
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RINGED-WORMS. 211
dividing it, as it were, into cells. Two conical yellow ccecal pouches
are placed at the commencement of this portion of intestine : thej
may probably be considered to be radiments of the liver. In
the common Leech, the short oesophagus, of an oval form, wider
towards the middle of its length, passes into a long stomach, which
is divided by transverse walls into eleven portions: on each side are
seen ten ccecal appendages to the stomach, the last of these being
the longest ; the inferior opening of the stomach {Pyhrus) extends,
like a funnel, into the intestine by a narrow opening. In other
genera of Himdineay ex. gr. in HcBTnopsisy the intestinal canal is
more simple, having only two cSecal appendages i. In Aphrodtta
there succeeds to a very muscular cylindrical tube, which Pallas
described as stomach, a thin intestinal canal of considerable width
with about twenty coecal appendages on each side*. These append-
ages are narrow at their insertion into the intestine, wider in their
middle, where they are provided with branched lappets, and termi-
nate in longish coecal sacs. This structure recalls the disposition
of the intestinal canal in FlanaricB and DiatomcUa, and the blind
branched appendages of the intestinal canal in Star-fishes may be
compared with it. They are filled, as these are, with yellow fluid,
and may be compared to rudiments of liver. In other animals again
the liver appears as a protrusion of the intestinal canal.
The system of Blood-vessels presents very many modifications
in this class. As to the blood itself, we have seen above, that
CuviER believed it to be red in all the ringed-worms. Such is
really the case in by far the greater number, as Hvrudo^ Imnbricus^
Arenioola^ Nereisy TerebeUa, Serpulaf &c.: in others it is nearly
colourless, sa in Aphrodite: yellow, as in PolyruoH^ and Phyllodoce, or
even green, as Milne Edwards found it m a species of SabeUa. The
general arrangement of the circulating apparatus is as follows: there
are two main stems, one on the dorsal surface, the other on the
ventral surface, which run in the midst through the whole length
of the body, and as fiur as the course of the blood could be deter-
mined in the living body — (for which investigation small indivi-
duals are frequently more fitted than large ones, on account of their
> See a figure in Brandt und KxTZKBURa, Medieinitehe Zoologie, ii. Bd. 1833,
Tab. xzix. B. fig. is.
■ Pallas, 1. 1. Tab. vn. fig. 10 d,d, fig. 11 <7,5r. G. B. Treviranus in Zeiiwhrift
ftkr Phyai'oUigie Til. 1819, s. 159—161, Tab. xn. fig. 9.
14—2
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212 CLASS Til.
transparency), the blood moTes in llie dorsal vessel from l)ehind for-
wards, in the abdominal vessel from before backwards^. In the Earth-
worm {Lumbrtcua) the two trunks are united in the anterior part of
the body by five or more (7 — ^9) arches widened like strings of
pearls. (It is almost impossible not to recall here the vascular arches
which in the embryos of mammals run along the branchial fissures.)
In others the connexion forwards is effected by vascular plexuses
{retia mirahilic^\ The dorsal vessel is usually considered to be
arterial, the abdominal venous : and in most of the ringed-worms
this opinion is not without ground, as might indeed have been con-
cluded fr^m analogy with other aiticulates. Sometimes the anterior
part of the dorsal vessel becomes wider, resembling a rudiment of a
heart, which then is in most cases an arterial heart like that of
spiders and crustaceans. The exception however observed by
Milne Edwards must not be forgotten; in TerebeUa the heart
drives the blood to the gills, and must therefore be considered to be
a venous heart, analogous to that of fishes.
Other less important modifications of the vascular system consist
in the breaking up of the two main stems into several, which are
sometimes quite separate from each other, though placed in proxi-
mity (Nephthysy Eunice)^ or in the presence of lateral longitudinal
stems. In Phiane carunculcOa there are as many as seven longitu-
dinal stems: Jour on the ventral surface, of which the middle ones
are small and lie at the sides of the nervous system, and the two
outer which are larger and give twigs to the gills, and three on the
dorsal surface, of which the two lateral receive the blood from the
gills, and are connected by transverse branches with the third or
median trunk '. In the Leech there are four principal stems, one
dorsal, one abdominal, and two, larger than these, lateral.
1 In ibis simple fundameDtal form the vascular system presents itself in NaU,
where an arched vessel at the anterior extremity of the body unites the two longitudinal
vessels. Gruithuisxn, Anai. der gegOngelim NiOde, Nov. Act. Acad. Cm, Leop.
Tom. XI. p. 133. And Ueber die Nal$ diaphana, ibid. Tom. xiv. pp. 407, ftc
' In Nereit : see H. Rathks, de Bopyro d Nereide €ommentaticne$ dua, 1837, 4to.
who calls these parts orffona rdicuUOa. Milkb Edwabdb, Ann. dea Sc. nai. ae S^rie,
Tom. X. Zocl. 1838, PI. 13, fig. I. Similar vascular plexuses exist also in PUione
caTunculaia, see G. R. Tbeyibakub, Beobachtungm aut der Zool. u. Phytiol. Bremen,
1839, ^ S4f ai^d A. £. Grubb, Jk Pleione cartmctUata Dm. ZooUm. Regiomonti
Prussor. 1837, p. 19.
' Gbubs, De PUione caruncuUUa, pp. 18, 19. On the circulation in the ringed-
worms I. MUKLLKB in Bubdaoh's Phyaiologie nr. 1831, s. 143— 149, may be also
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RINOED-WORMS. 218
Bespiration is effected hj the skin, or hj external gills of very
different form, or by vesicles on the sides of the bod j. In the Leech
there are found about seventeen such vesicles on each side, which
open on the abdominal sur&ce. The openings are extremelj minute,
and between two of them on each side there are four rings or seg-
ments of the bodj without such openings. A white convoluted
structure is connected with these vesicles by means of a thin pedicle,
and contains (according to DuGi^) a blood-vessel in its interior.
That these vesicles secrete mucus, is no proof that they are not
respiratory organs ; some writers think that it is their sole function
to supply that secretion; and Brandt believes that respiration in
the Leech is effected by the skin. At all events, though these
vesicles receive and return blood-vessels, they have not a perfectly
separate circulation of blood in them, and the respiratoiy organs
would seem to receive in this case, as in that of Beptiles, a portion
only of the venous blood. Li the Earth-worm there are more than a
hundred such vesicles ; their openings are on the abdominal surface,
according to Leo and DuG^, whilst Meckel and Morren think
that they are connected with a single series of apertures on the
dorsal surface, which Willis formerly described and compared to
the spiracles of Insects*.
The ringed-worms, until within the last few years, were sup-
posed, almost universally, to be bisexual. It was only in the
AphroditcB that a separation of the sexes was, with some hesitation,
accepted, when Pallas had shewn that certain individuals were full
of eggs at the same time that in others the cavity of the abdomen
contained a tenacious milky fluid*. Afterwards Rathke also found
a separation of the sexes in Amphitrite^^ and Quatrefages observed
the same in a large number of marine ringed-worms {tubicoloB and
erraniiay. The observations of Steenstrup on Lepidonote, Phylh-
doce^ Nereis^ Nephthys, Terebella, and Serpula are to the same effect:
in the last genus the sexual distinction may be recognised by the
oonsoHedy and espedaUy Milni Edwards, Awn, dea Se, not, see. S^rie, Tom. z.
pp. 193—941, Fi. 10, It. (These figures are also transferred to the new edition of
GuTiBB, Riffne AnimtU, Anndidee, PI. i, Ac.)
^ J)e Amma BrtUcrum, AmsteLodami, 1674, Svo. pp. 34, 35, Tab. iv. fig. 3.
' Mi$e, ZoU, p. 90.
■ BeUrage zwr vergl, Anat. u, Physiol, Danag, 184a, s. 66 — 68.
* MiLHB Edwabds, Bajijpcrt tur une Sirie de Mimoires de M. A. DK Quatrk-
rAOKSy Ann, de$ 8c, nak 3ibme S^rie i. p. 21.
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214 CLASS VII.
colour shining through the skin, which is white in male individuals
and reddish in female. Other differences of external appearance
in the two sexes are not known, unless we except an observation of
Oersted, which however is not altogether free from doubt, accord-
ing to which in a new genus very nearly allied to SyUia^ which he
names Exogone, the male individuals axe distinguished by longer
hairs, as they are in the genus Nats^. There still remain the
genera of the Hirudinea and Lumbricim, in which Steenstrup in-
deed adopts separation of sex under similar external form of the
parts : but this requires confirmation after accurate investigation, for
it is in conflict with earlier observations, whilst by later it is in part
contradicted*. On the whole, no common type can be assigned for
the genital organs : for the most part, there are some pairs of vesicles
{ovaria, testes) in the fore part of the body. In some Anmdata aetyera,
apertures at the base of the foot-swellings have been seen, through
which passes seed or eggs : but in many of them such an outlet is still
unknown. For the most part, external genital organs are deficient :
neither does copulation occur, except in Lumbricini and Hintdinea.
The development of the egg has been investigated only in a
few species. Here also that remarkable cleaving and successive
division of the yelk has been observed, which RuscONi and VoN
Baer first detected in the eggs of frogs and of fishes. The deve-
lopment of the embiyo begins on the abdominal surfeuse, and the
yelk lies on the dorsal surface, as in Crustaceans and Insects : two
abdominal streaks are observed at the commencement of develop-
ment, which recall the dorsal plates of vertebrate animals^.
The most recent times have made us acquainted with some
remarkable metamorphoses in the course of the development of
ringed-worms. Loven found the first stage in a worm of the
family of the Nereids (probably a species of PhyUodoce) to resemble
^ Eriohson's Archivf, Naturgetch. 1845, i. b. 10—23.
* See F. Mueller on the Hermaphroditism of the Hirudinea, in the German
transhition of Steenstrup'b work cited above (p. 135) Untertuchungen ueber das
Vorhcmmen det ffermaphrodititmus in der Natur, Greifswald, 1846, 8. no — 114.
* Most of the observations refer to ffirvdinea. Such are the following works : —
E. H. Weber, Ueb. die Entvnckdung dea medicin. BhUegda, Meckel's Arckiv. 1828,
8. 366—418, Taf. X. zi. ; R. Waoner, BruchaMicke atu der ErUwickdung des gemeinm
Mutegels, Hirudo vulgaris L., Nephdis ieasdaJta Sav. Oken*s Isis, 183a, s. 398 — 408,
Taf. iv.; A. E. Geubb, Untersuchungen ueber die JSntwickdung der Cleptinen, Mit 3
Kupfert. Konigsberg, 1844. We may expect several observations on marine amwlola
from QuATREFAOES. See Arm, des Sc, not. ^ihm.B S6ne, Zoologie i..p. 3i.
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RIKGED-WORMS. 215
a hemispherical or conical body of about ^ milllm. terminating in a
ciliated disc on whose edge the mouth seemed to be placed. At
the pole of the hemisphere was the anus. This conical body
increased gradually in length and became divided into rings gra-
dually more numerous, the last formed ring being that next the
disc (just as in Esc^hricht's observations on Baihriocephalua the
new rings were formed in the anterior part of the body). Each ring
originally consisted of four pieces: an anterior and a posterior
piece being larger, almost a semicircle, and a smaller piece on each
side connecting them. The disc with its vibrating cilia diminished
gradually and became changed into two fin-like appendages to the
head, from which the feelers probably proceed^. Sars saw the
incipient form of Polynbe cirrala as a short, oval, inarticulate body
with a transverse circle of vibratUe cilia round the middle*. It
may be confidently asserted therefore that there is a melamorplums;
parts are present which afterwards disappear (the vibratile cilia),
others are deficient which are afterwards developed, and the entire
form is changed.
The [Reproductive force is, in some animals of this class very
great, in others small, although worms that have been cut through
transversely continue to live for a long time, as has been observed
in the leech, and by O. F. Mueller in Nereis vernoolor. Treh-
bley's experiments on the Fresh-water Polyp induced Bonnet to
repeat them on Fresh-water Worms (Naules), and he found that
the pieces he had cut off grew into new worms^. Mueller also
succeeded in similar experiments^. It has been thought also that
they have succeeded in the Earth-worm, but here they have con-
stantly failed with other experimenters. According, however, to
the experiments of DuG^ a few rings at the anterior part of the
body may be reproduced and gradually changed into a head^
' S. Loven, Zooloffiska Bidrag; MeUtmorpkoB hos en Anndid {Aftryck ur K,
Voentk-Alcadtm, Handlingar, 1840) ; translated into most of the zoological journals :
Ann, de$ Sc. not. 2e S^r. xvni. p. 288.
■ 'EniCBBOTx'B Archiv. 1845, i. s. 11—19, Tab. i.
' ObtervtUioTU mr qudquet e$pice» de Yen d'eau douce; (Ewvret (Aiit. 8vo.) pp. 167,
Ac. Especially in LumMctu variegaius Muell. {Lumbricvlut variegalue Gbubb) is
this reproductive power great, in which Bonnet saw the amputated head renewed eight
times in two months.
^ Von Wilmum dcs mtsen u. sakigcn WasterSf s. 43, Si, &c.
• Ann. des Sc. not. XV. 1828, pp. 317, 318.
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216 CLASS VII.
The Nervous System in the AnnuUxta proper consists, as in
Insects, of ganglia connected bj two cords and placed behind each
other in a series in the middle of the body on the abdominal sur-
face. Originally each ganglion consists of two lateral portions, as
is proved by the process of development : on the regeneration also
of parts that have been cut away the nervous system appears to be
formed of two lateral portions. A larger ganglion lies in the head,
and is connected, by two nervous threads that form a ring around
the oesophagus, with, the first ganglion of the abdominal chain.
But the Nervous System presents much variety in different genera,
as well in the number as in the greater or less development of the
ganglia and in the nerves that spring from them ; whilst in the
earth-worm, for instance, the numerous ganglia of the abdominal
chain almost touch each other, in the leech they are only twenty-
four or twenty-five in number, and are placed far asunder, especially
in the middle. In Pleione carunculata the Nervous System con-
sists, according to Grube, besides the middle chain, of two lateral
cords, also with ganglia, which are connected with the former by
transverse threads ^ In Eunice sanguinea Quatrefages found
minute ganglia at the base of the rudimentary feet, which however
were not connected, as a chain, by longitudinal filaments. In ad-
dition to this nervous apparatus a special nervous system has been
detected in many instances, agreeing with that portion of the
nervous system in Insects which has been compared to the Nenms
aympathicus of the higher animals : of which we shall treat more
at large at the class of Insects. In Hirudo medtcinalis Brandt
discovered three minute ganglia in the head, which are united by
threads with the cerebral ganglion, and from which the maxillary
nerves arise ; with the middlemost of the three ganglia a nerve is
probably in connexion, which runs beneath the stomach in a longi-
tudinal direction and finally divides into two branches ; but this
nerve differs from the sympathetic of insects in respect of its
position on the inferior surface. In Eunice sanguinea and some
1 IH88. zootom. de Pleione carimc, p, 9, figs, x, 5. Stannius (/w, 1831) observed
the same thing in another species of Pleione (Amphinome rostrcUa), It is as though there
were a repetition of the form of the vascular system on the dorsal surface, which here
consists of three stems; see above (p. an). Perhaps this arrangement occurs in
several Annulata; at least Wagneb describes it also in PowUMdla fMiricata, Lekrb.
der vergl, AncU. 1835, s- 38 r.
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RINOED-WORMS. 217
NereidiB QuATREFAQES fonnd this system to be composed of differ-
ent ganglia, and named it, on account of its position on the pro-
boscis, sysdme atisoBacphagien, or probasddten supirteur^.
With respect to organs of sense, with the exception of tentacles
and other appendages subservient to a finer sense of touch, for sight
there are found in most species only coloured spots, usually black,
of variable number, as special organs. According to I. Mueller's
investigations in a Nereis, the eyes of ringed-worms contain no
transparent parts, but are merely swellings of the visual nerves sur-
rounded by black pigment. They are endowed with sensibility for
light, and the worms can distinguish between light and darkness :
but what is properly named aiffht, perception of the form of objects,
such eyes cannot afford. In Alciopa lepidota Krohn however found
a lens and a vitreous body. A special auditory organ has not been
detected ; the first portion of the oesophagus is supposed to be the
seat of taste.
The organs of motion are in some more complicated than in
others. In all muscular fibres are found beneath the skin, which
may be separated more or less completely into layers : the external
layer has a circular, the internal a longitudinal course. In some, as
Aphrodite, these fibres are imited to form distinct bundles. By means
of the layers or bundles the body can be moved, contracted, extended,
bent. Besides this general muscular system, motion of the body
in the Leech can be also effected by means of a suctorial disc at its
posterior part, in which there are circular and radiating fibres. The
proboscis, which is capable of eversion and retraction, has proper
muscles for these purposes. Concerning the bristles and hairs,
which are found on many, we have already spoken above. These
parts, springing firom the sides, supply fixed points for the motions
of the body, like the spines of the Echini: they are retracted, ex-
tended or moved sideways by proper muscles.
Many species of this class diffuse a phosphoric light. It is
^ There lies also a small ganglion in firont of the brain {ffan^ion cervical Quatbs-
FAGKs), and from the lateral parts of the brain a thread arises, which with that of the
other side surrounds the mouth (Sydime mmHUOfhagien labial au prohoscidien
infiriewr); this last portion of the nervous system may be compared with the arrange-
ment in the MoUtuca gcuUntpoda, See on the nervous system of the Anndidt
a memoir of Quatbefaobs illustrated with beautiful figures, Ann. def Sc. not. 3e S^rie,
Tom. n. Zoologie, 1844, pp. 81 — 104,
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218 CLASS VII.
asserted that this phenomenon has been occasionallj observed in
the earth-worm {Lumbricus terrestris L.) It is quite certain that it
has been seen to occur in very many marine Annelides: and hence
these are creatures also which contribute to the illumination of the
sea. NereidcB are especially noted in this respect : DuGiis observed
the phenomenon in a Mediterranean species 4!' long, Syllis fidgun
rans^. QuATREFAGES made the important discovery that, in certain
minute marine Annelides (species of Syllis and Poli/noe), the seat of
the phenomenon is at the base of the feet-tubercles — ^in fact in the
muscles: it was only when the muscles contracted that the light
appeared like an electric spark ^.
Binged-worms are found in all countries and seas : but it is im-
possible to present a view of the geographical distribution of the
species hitherto known, for this class is perhaps more generally
neglected by voyagers than any other, and we are acquainted with
few marine annelides except those from the Atlantic ocean, the
Mediterranean, and the Red sea. Of the genus Palmyra only one
species is known, which was found at the Mauritius. From the
Indian sea some large and beautiful species are known, as Laodicea
gigantea; Serpula gigantea is from the West Indies : and, in general
the large and beautifdl species are most numerous in warm regions.
Some species appear to have a very wide geographical distribution,
as Hestane splendtda, found by Savignt in the Red sea, and brought
by Matthieu from the Island of Mauritius : and Pleione carunculata,
which, according to Pallas and Savigny occurs in the American
seas, according to Seba in the Indian sea, and was found by Grube
in the Mediterranean at Sicily. The Hirudtnea and Lumlmcini
have scarcely been investigated elsewhere than in Europe.
^ Ann, dea Sc. not, Tom. xxix. p. iig. The NereU noctUuca L. is a Bmall animal-
cule, probably the same species as Nereu cirrigera o{ Viviani. P6lyno€ ffd^rans, at
most half a line long, was observed and figured by Ehbembbbo, Leuchten det Meera,
Tab. I. fig. I.
' Ann. des Sc. mU. sec. S^rie, xix, 1843, Zooloffie, pp. 183 — 192.
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SYSTEMATIC
DISTRIBUTION OF ANNULATA.
CLASS vn.
ANNULATA-
Animals elongate, living in waters or moist earth, not parasiti-
cally in other animals, mostly articulate, without jointed feet, but often
in place of feet supplied with setse or setiferous tubercles which are
retractile. Kespiration effected either by external branchiaa or in-
ternal sacs or by the skin itself. Organs of circulation in most
distinct ; contractile vessels instead of heart. The nervous system
composed of a cephalic ganglion single or double, and most fre-
quently of a double ventral cord with ganglia at intervals.
Order I. Turhellaria.
Body cylindrical or depressed, most frequently inarticulate, or
ringed by transverse rugae, beset with vibratile cilia.
Family I. PlanariecB. Nutrient canal with one distinct aperture
alone, anus none. Body inarticulate.
This &mily was originally formed from the genus Plcmaria of
O. F. Mueller, which was divided by later writers into other genera,
and round which in consequence of new discoveries other different
genera were arranged. It appears to us to be inconsistent with the
idea of a class, to raise this group to that rank, as Yok Siebold has
done, who has formed his class of the Tttrbellaria of it alona The
name TwheUaria was first, though in a more comprehensive sense,
used by Ehbenbebg* (see above, p. 208). The phenomenon of
rotatory motion in the water surrounding these animals, which gave
> SymiMa phynae Anim. eveiiebraia exduiit ifUectia, i. Berolini, 183 1, fol.
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220 CLASS VII.
ocoacdon to the name, was first, as it seems, observed by Ducds in
FlcmcMrice, although he did not refer it to cilia (Ann. des Sc. tuU.
Tom- XV. p. 165), whilst Von Baeb observed at the same time an-
other phenomenon which could only be an effect of these cilia,
that when a portion of these animals is cut off it continues to rotate
circularly in the water {N'ov, Act, Acad. Cceaar. Leap. Carol. Tom-
XIII. P. 2, p. 711>
The internal structure of these creatures was first recognised with
precision in this century, especially through the investigations of
"DuGite, V. Baeb, Fokke, Oebsted, and Quatbefages; it presents
important differences in different genera whilst the external habUua
is similar. We must confine ourselves to the notice of a few general
traits.
Besides the cilia on the external sur&ce, the external integument
in many is distinguished by the presence of cells with nettle-threads,
like as we stated in Acalephse. Beneath the integument there is
a layer of transparent, homogeneous tissue, which, according to
QuATKEFAOES, suppUes, as it seems, the place of muscle, and by its
contraction effects the movement of the body. The motion occurs
by swimming in the water, the lateral margins beating to and fro,
by creeping with bending and straightening of the body, or by equably
gliding, much as the gasteropod molluscs move with their so-named
foot.
The oral aperture is sometimes more forward, sometimes more in
the middle of the body on the ventral sur&ce. The intestinal canal
is in some straight, and extends itself, when the mouth ia not placed
quite forward, anteriorly as well as posteriorly, with blind terminar
tions in both directions In others the intestinal canal is like a
tree divided into branches ; in our fresh-water species one branch is
seen to run forwards, on both sides provided with blind appendages,
and two stems or main branches backwards at the sides of the body
(in Pkmaria lactea the branches may be readily distinguished
externally by their dark colour). In other species from sea-water
the division of the branches is somewhat different ; sometimes quite
retiform. In the cavity of the mouth is situated a part that can
be extended by eversion, serving for the seizure of food, and various
in form. It is able, when severed from the living creature, to move
independently for some time whilst it swallows greedily sun"ounding
substances which are seen to pass out by the posterior open ex-
tremity as through a ^nnel. With respect to the vascular system
little is known ; that which is described as such by some writers.
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ANNULATA. 221
belongs probably to the nervous cfystem. In some species with
straight intestinal canal, there have been observed at the sides two
tortuous canals running longitudinally, which, without giving off
lateral branches, bend round in a loop at the back part Respira-
tion is probably effected by the skin itself and the water on the
sur&ce is constantly renewed by the vibratory motion.
As nervotis system in many a double nervous ganglion has been
observed, which lies at the anterior end, and from which many
branches arise. The eyes, which are in some instances very nume-
rous, present in many a transparent body, corpus vitreum or lens
crystaUina.
The reproductive power is very great, and severed parts grow, as
appears from the observations especially of Duoes and J. R Johnson,
to new aniuLals. In some propagation occurs by spontaneous division.
The sexual organs have one common or two separate openings behind
the mouth; in the latter case the anterior opening belongs to the
male organs of copulation. Two long tubes supply the office of
testes, and end as vasa deferentia in a seminal vesicle, with which a
penis of various form is connected. The spermatozoa have been
observed by Quatrefages and Yon Siebold. A double oviduct
leads to a spacious vagina, into which two special hollow bodies also
opecL The ^gs lie dispersed in the parenchyme of the body,
between the ooecal branches of the intestinal canal (Quatrefages),
where probably they are contained in special ramified tubes
(ovaries).
Compare on this family :
VoN Baeb, Ueber Plana/rien. Nov, AcL Acad, Cess, L, G, not
cur, VoL XIII. P. 2, pp. 690—730.
DuGis, Eecher<^es su/r Vorgamisat, et les moev/rs des Flanariees,
Arm, des Sc, not xv. 1828, pp. 139 — 187 ; Observations rumv. siur
les FUmaires, Und. xxi. 1830, pp. 72—92.
A. S. Oersted, Erdwy/rf einer systematischen Eintheilung und
speciellen BerschreU/trng der Plattmirmer. MU HoUz-schnUtenund
3 Tafdn^ Copenhagen, 1844, 8vo.
A. Db Quatrefages, Memovre sur qaelques FUmariees marines,
Arm, des Sc. not. 3me S6rie, Tom. iv. 1845, Zodog, pp. 129—184.
PL 3— 8.
M. S. ScHULTZE, Beitrdge sur Natmgeschichte der TwrbelUm^n.
Erste abtheUwng, Hit Kwpfertaf. Greifswald, 1851, 4to. also in
Wibgicann*8 Ar<Mv, 1849. § 290.
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222 CLASS YII.
Phalanx I. Ehabdocoela. Intestine simple, cylindrical, not ex-
sertile from the mouth. Body elongate, roundish or depressed.
Prostoma Oerst. (not DuGES), Oyrator Ehrenb. Oral aperture
anterior terminal.
Sp. ProtUma lineare Oebst., Oyrator hermaphrodUus Ehbskb., Ahhandl.
der Ahad, der Wisaeruck, zu Berlin, 1835, l^b. I. fig. 2.
Vortex Ehrenb.
Sp. Vortex truneatus, Plancma truncatck, Zool, dcmie. Tab. 106, fig. i^ a, b,
Ehbskb. 1. L figs. 3, &c.
Deroatoma Oerst. (DuGieis in part).
Mesostoma DuGES, Oerst. Body depressed, transparent; oral
aperture annular, rotund, situated a little in front of or in the middle
of the body. Eyes two anterior.
Sp. MeBOttoma EhrmbergU, PUmmria tetragona Muxll., PatcioUi quadrangu-
larU Pall., SpicU. Zool, x. Tab. L fig. 13; Zool. danic. Tab. 106,
figs. I — 5 ; FOKKX, Aim.det Wiener Museume, L 1836, pp. 191 — 306, Tab.
xym. Thia species has been elaborately inyestigated by the last-nained
writer ; it undergoes various changes of form ; from the flat form a qua<f-
rangular arises, as though the animal were about to divide itself in the
length. Pallas had already observed this Planaria more than 70 years
earlier at Sorgdiet, and it has lately been found again at Leyden by Hkbb
HXBKLOTS.
Strongylosioma Oerst.
Tj/phhplana Ehrenb.
Mcuyrostoma Oerst.
Microstoma Oerst. (Vid. Schultze quoted p. 203.)
Convoluta Oerst.
Phalanx II. Dendrocoela. Nutrient tube branched. Body
depressed. {Dendrocoda and Gryptoccela Oerst.)
a) Appendages nwmerouSy t/ubular or papiUce on 1M hack,
Thysanozoon Grube, Eolidtceros Quatrep. Eyes sessile, nu-
merous ; body excised anteriorly and cloven into two tentacles.
Sp. Thytanozoon BroechU, Eclid. Brocckii Quatsbf. L 1. PI. 5, fig. i, (perhaps
the same species as ThysanozMn Dietingii Gbube, Actimen, Echinodermen
u, Wilrmer, fig. 9) ; in the Mediterranean. The intestinal canal has here a
retiform division.
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ANHUUITA.
h) Body smoodi
SttfhchuB Ehsenb. Eyes numerous, all or most of them sup-
ported hj dorsal tentacles.
Ijeptoplana Eh&enb.
PUmaria Ehrenb. (Species of genus Planartd MuELL.) Fla-
naria and Dendroccdum Oerst. Eyes two or a row of many eyes
in the anterior margin of the body. Oral aperture in tlie middle of
the body.
Sp. PUinaria laetea MuxLL., Zool. dan. Tab. T09, figs, i, 1, Planaria torva
MuKLL., ibid, figs. 5, 6 ; Planaria nigra Musll., fig«. 3, 4, all in freah-
water.
Polycelis Ehrenb. (and Prosthioatomum QuATREF.)
Tetracdis Ehrenb.
Tricdis Ehrenb.
Monocelis Ehrenb.
Note, — On these, and some other genera all of which are not yet
sufficiently limited, consult Ehrenberg Symb. phya. AniTn, evertebr,
exduais inaectia, i., and Oersted L 1.
Family II. Nemertim, Nutrient tube simple, with double
aperture, anus terminal. Body elongate, extremely contractile,
roundish, or depressed, indistinctly annulate.
It is not without hesitation that, after Oersted, we have given
these characters of the fiunily of the Nemertini, whilst amongst the
different writers, with respect to the true nature of the distinct
partSy a remarkable variety of opinion prevails, so that it is un-
certain whether the aperture, considered as cmua, really belongs to
the intestinal canaL Beneath the skin in these worms muscular
fibres are seen, of which the external layer runs longitudinally, the
innermost annularly or transversely. A canal of uniform width, by
many supposed to be the intestine, runs straight through the body
(Delle Chiaje, Huschke, Kathke). On its dorsal surface lies a
canal, which is closed at its termination backwards, becomes narrower
forwards and ends in a long proboscis. This part is, according to
QuATREFAGES, the proper intestinal canal, which consequently has
no amis. Huschke supposed it to be an organ of propagation
(testis f) and the proboscis an external copidative oigan ; hence the
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224 CLASS YII.
name Notoapertnvs, which he gave to the worm examined by him.
The Nemertini appear, according to Kathke and Quatbefaoes, to
have the sexes distinct, and the organs of propagation (testes, ovaHa)
consist of blind saccules, which lie beneath the integument longitu-
dinallj, upon the wide canal already described. There are three
blood-vessels running longitudinally, two on the sides and more
towards the ventral surface, and one on the dorsal surface, which
divides anteriorly into two branches which pass into the lateral
vessels. The nervous system consists of two head-ganglia united by
a transverse cord, from which (besides other nervous branches) two
very notable nerves arise, which run longitudinally backwards,
along the sides of the body.
Compare for the anatomy of this family, besides Oersted and the other
writers cited above, Delle Chiaje, Memorie n. pp. 406 — 409 and 427,
(extract by R. Waoneb in Oken's Ina, 1832, s. 555, 556, 8.647—649) ;
HuBOHKE, Okek*s Im, 1830, 8. 68 1— 683, Tab. vn. figB. i — 6 ; Rathke,
Beitrdge zur vergL Anat. u. Physiol, Danzig, 1842, s. 93 — 104 ; Quatbe-
faoes, VJnstUut, Journal universd, &c. No. 660, 1846, p. 286, and a figure
in the new illustrated edition of Cuyieb, B^gn^ Animal, Zoophytes, PI. 34
(and also a M^moire sur la famUle des Nimeriiens, Ann. des Sc. not. ^ihme
S^rie, Tom. vi. Zool. pp. 173 — 303).
Nemertes Cuv., Borlasia Oken. Several eyes (often indistinct).
Two pits (respiratory?) at the sides of the head, surrounded with
vibratile cilia.
Sp. Nemertes Borlasii, Borlasia Angtias Oken, Borlabe, Nat, BisL, 0/
Cornwall, fol. 1758, PL xxvi. fig. xm. (cited by Cuvibb), Quatbefaoes in
Cuv. B. Ani, idU, HI,, Zooph, PL 33 ; this worm becomes more than four
feet long ; &c.
Note, — Oersted cites as synonyms of this genus Notaspermus
HuscHKE, Meckelia Leuck., Ophiocephaltu Quoy and Gaiil
Oersted has given the name Borlasia to species with constricted
head, without respiratory fissures, with indistinct eyes. Add several
genera, of which the synonyms cannot be made out except by com-
parison of the specimens : Cephalothrix Oerst., Astemma Oerst.,
Tetrastenvma Ehrenr, Polia Delle Chiaje, Polystemma, Omr
TnatopUoy Amphiporus Ehrekk, Cerebrat^us Renieri, Amphiportis
Oerst. (not Ehrekr), Serpentaria Goodsir*.
^ Ikscriptions of some gigantic forms of invtrtebraie animals. Annals and Magazine
of NaL Hid. VoL XV. 1845, P- 377, PL "•
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ANNULATA. 225
Order II. Suctana.
Body annnlate, without setae, terminated by a prehensile cavity
posteriorly or at both extremities. External branchiee none.
Family III. Hirudtnea. (The characters of the order are also
those of the single family.)
The &mily of the blood-suckers (leeches) is formed fix>m the genus
Hvrudo L. These animals are able to convert the anterior extremity
of the body into a suctorial cavity, or have there, as at the posterior
extremity, a round suctorial disa They creep along the ground, by
Affixing this sucking apparatus and by alternately contracting and
extending the body They swim with a serpentine and sinuous
bending of the body, which is effected with much velocity.
Comp. on this funily MoQunr-TAirsov, Mimographie de» Bintdiniet,
nouv. Sdit, av, pi, color. Paris, 1846, 8vo.
A. Head made up of several segments of the body, slightly or
not at all distinct &om the rest of the body, capable of change into
a suctorial acetabulum by its own motions.
Clqmne Sav. Body depressed. Mouth unarmed, furnished
with a proboscis exsertile, tubular. Eyes 2 — 6 (sometimes eight?).
Sp. Clepsine kyaUna, Eirudo hyalina L., Trkmblst, Polyp, PL vu. 6g. 7 ;
Cleptme complanaia, &c.
Tins animal lays its eggs on water-plants {StraUotet aiolda) andoontmues
to sit thereon. The eggs also attain their deyelopment even when the mother
is driven away, but are then frequently affected and spoilt by oonfervae. lliese
^;g8 are thin-skinned vesicles in which numerous yelk-spheres, 15 to 30,
are contained, and from which a corresponding number of young are
developed.
Comp. on the species of this genus F. Mublleb, Jk ffirudinibui eirea
Berolinvm httcuaque obtervatiSf Berolini, 1844, 8vo, and T. Budob, Cleptine
hioculata. Mit 2 Taf. Bonn, 1849.
Under this genus F. Mubllbb also places JSvrudo nuurffinata and Hir,
ietwlata of O. F. Mxtblleb, though the last has eight eyes, whilst in
Cleptine the number does not exceed six ; the arrangement of the eyes is
in two rows longitudinally which meet forwards, just as in the six-eyed
CUpaineB; the blood also is white.
Nephelis Sav. Body elongate, posteriorly incrassated, obtuse,
with acetabulum obliquely terminal. Mouth unarmed. Eyes eight,
disposed in a series semicircular, transverse.
VOL. I. 15
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226 CLASS m.
Sp. NtfUk w^gtriM, Bwmdo PrtifrfHi I^ £mrfd4^ aiA^h^ Ttn. FL51,
fig*. 5—7; Smoi, iXnteU. A«m ti. 4 Heft; Jonnov, PAa7. IVm.
1817, PL IT. (reprintod in bk FmriUr ObmnaL m, tke Lmk, 1825); thk
species swallows HBafl wonniL
Troeheta DcnocHn, Trotketia Lam. {GeobdeUa Blust. in put).
Branckiobddla Odiek. Body somewhat depresaed, with laige
rings, not nnmerons. Two homy jaws. Ejes none.
8p. BramekiobdeUa adan, OmBB, Mimk. de Im Sk. ^BuL m(. d^ Pflrif I.
i^^if PP- ^ — 7^9 PI- !▼• talreadyolmiiul and figured bj B«»bl» /at. IIL
PL LOL figs, ig—^1^); Bmek, pflrsnCa, eomp. Hsm, Tcft. ii«e giiff—y
BranckiobddU, Mukllkr's ilfritr. 18^ s. 574, Jkc TwL xrr.
Htrudo L. (exclnsiTe of several species). Bodj oblcmg, sub-
depressed, with nnmerons rings. Three homj jaws. Eyes nsoallj
ten.
Bdetta Say. Jaws not denticulate. Eyes eight.
Sp. BddLoL nUcHca Sat., Guiani, lamoffr., Annil. PL 4, fig. to.
HamopU Say. Maxillap? aimed with a double row of denticles not
numeroua Eyes ten.
Sp. Hamopu unguisorha, Hirmdo mmguimtga L., Emofdap, mitk., Ven.
PL 51, figs. 3, 4, Uacky grey-green beneath; this speoes is larger than the
common Leech. Conmionly two species haTe been here confounded,
which MoQuur-TAjn)0!r pUuses in two different genera: AwUutoma {ffir.
mnguituga MuKLL., Hir, (hUo BaAUV, SruaM, iXewtodU. Famna ti. 2) nnd
Heemopu {Bit, mtmguiguga Bbbgic, L.)
Scmguisuga Say. Jaws armed with a double row of denticles
very slender and crowded Eyes ten.
Sp. Htrudo medieinalit L., Hirudo vetuaeetor Brack, Sturm, DeuttekL
Pavna Ti. a Heft ; BRAVnr u. RAnsBURO, Medu. Zool. 11. Taf. xxtiii.
figs. 5, 4 ; GvtaiM, loonogr., AnmiL PL 10, fig. 3 ; aboTe, blackish-green,
with six long stripes spotted reddish and black, beneath diTe-ooloured with
many black spots (four to ^ye inches long). Another species, Hir, ojfcinalis
Sat., is also used for medicinal porposes, which is yellowish beneath with
a broad black edge, without spots, see Brandt n. Ratzxb. L L Taf. xxx.
fig. I ; J. J. KnoLZ, Naturkui, Abkandl. ueber die Bluiegd, Wien, 1820,
1 This genus must by no means be confounded with £ranchiobdeUion Run., Bran-
cJidlion Satiqnt, which like Cleptine and NepkdU has no jaws, but only three project-
ing points ; if the semicircular little plates on the same part of the body be really gills,
as Sationt says, (Cutixr doubts and Moquin-Tandon denies it^) then it does not
belong to this order. Latrullb places it near the genus Areaioaia.
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ANNULATA. 227
8to. Tftb. I. fig. a ; aooording to Kholz it is thia specieB especiaUy which
is used in Vienna and brought there from Hungary^.
Hirudo medicinalis is the most useful species of Leech {ianfftue, leech,
BhOeffd), which almost everywhere in Europe lives in fresh water, especially
in ponds, marshes and canalsy and in winter, rolled up annularly, conceals
itself in the mud. This animal lives on the blood of animalfl {verUinrale and
uivertebnOe) exclusively ; the jaws serve to wound and to penetrate the skin .
The first segment of the body, which also is occasionally parted by a trans-
verse stripe, has a semilimar form and is not closed beneath. It can extend
itself as an upper lip for feeling or bend itself downwards to cover the
mouth. The ten black eye-spots are arranged in form of a horse-shoe on
the back-side of the head ; the first on the first segment, the two next on
the third, and the two last on the sixth ring of the body. The organs of
propagation of the leech are by different writers determined very differently,
whilst, however, the latest investigations (especially of H. Mecksl,
Mubllkb's Archiv. 1844, s. 476 — 480) bring us back to the generally
received opinion of former times. According to it, nine purs of round
vesicles of a white colour are teslee, (Tbeyiranus thought they must be
held to be ovaries, ZeiUchr. fUr Phytiol. rv. 2, 1832, s. 159—167). By
means of short transverse tubules these vesicles are connected with a com-
mon canal which runs at each side of the body ; this canal goes forward
into a structure which is white and consists of many convolutions (the
epididynUs or the seminal vesicle). From each of these two seminal
vefddes arises a short vessel (vas ejcumlatorium), wMch runs to the spherically
widened sheath of the penis : the penis can be everted outwards through
an opening in the twenty-fourth ring of the body. In the fifth ring
behind this is seen the second sexual opening, that of the female parts ; it
leads to a wide vagina (uterus, according to BoJAirus) which, by means of
a tube that divides forwards into two branches, is connected with two
smaU ovaries or vesicles filled with granular bodies. These two ovaries lie
between the seminal vesicles and the vagina. The impregnation in Leeches
is mutual. The Leech lays eggs, or rather capsules, in which eggs
are contained, 5 — 16 in number. These capsules or cocoons are three-
fourths of an inch long, oval and surrounded with a spongy or fiiothy
substance, and filled with a brown albuminous fluid. The germs appear as
round discs; these minute yelks grow by means of the surrounding
albumen, which w absorbed by a structure which closely resembles a funnel-
shaped oesophagus, and is already visible on the germ when only half a line
in size (E. H. Wkbeb in Mbckel'b Archiv, i8a8, s. 366—418, Muxllbb'b
Archiv. 1846, s. 428 — ^434).
Comp. on the Leech amongst others : JoHVSOV, TreaHae on the Medicinal
Leech, London, 18x6, 8vo, and by the same. Further Obeervat, on the Med,
Leech, With engravings. London, 1825, 8vo ; Kuntzmaitn, Anatomieche
Physiol, UfUersuchungen liber den Blutegel, m. 5 Kupfert. ; Bojanub in
^ Other species still, which have been discovered, may be used for drawing blood,
as the large black species spotted with white which was discovered in Sweden some
years ago by WAHLBK&a, and named Hirvdo albopunctata.
15—2
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228 CLASS VII.
Okbn'8 ItU, 1817, 8. 881 (with fig.)> the Bame 18 18, b. ao8o; Knolz
(seep. 116) ; BBAimr Mediz, Zool. n. 1833, s. 230 — 297 ; Moquik-Tavdok
Monographic dct Hirudiniea, &c.
Amongst the foreign species we note Jffirudo zeylaniect, found in the
Island of Ceylon ; its poisonous bite is followed by very tedious uloen.
Tttlsb, JSdi9ib. new PhUot. Joum. 1826, p. 375.
B. Acetabulum of the mouth of a single segment, distinct
from the rest of the body by stricture.
HoBmocharis Sav., Pisdcola Blainv., Lam. Body cylindrical,
attenuated forward, with few rings, little distinct. Anterior aceta-
bulum slightly excavate, with mouth triangular, edentulous, placed
in the bottom towards the inferior margin; posterior acetabulum
large, obliquely terminal.
Sp. Mcanocharis piaeium, Hirudo gwmttra "L., RossKL In», m. Tab. xxxn. ;
Lxo, Ud>er einige attsgezeichneU anatomiscKe und phygiologiwhe VerhdUniam
derPiscieola geometra, Mueller's Archiv, 1835, s. 419— 427, Taf. xi. This
species lives in fresh-water and adheres very firmly to Carp, Tench, ftc.
It moyes like a geometrical caterpillar ; on the back-side of the cephalic
disc are four black eye-spots ; copulation occurs in the upright position, in
which the ftntmnla Bupport themselves on the ventral disc and embrace in
form of an X. They lay eggs of a yellow-brown colour, three-fifbhs of a
line long.
Piscicola respirani Tbobchxll, new species, Archiv f. Ntthtrgetch, xvi.
1850.
PorUobdella Leach, Lam., Albiane Sav. Body cylindraceo-
conical, attenuated forwards, with unequal rings. Acetabula very
concave; mouth small unarmed, placed at the bottom of the anterior
acetabulum ; posterior acetabulum exactly terminal.
These animals live in the sea and adhere to different fishes, especially to
Rays. Most of the species are beset with nodes or with warts on the
rings, which are flatter in Pontdbd, verrucosa, Babtbb Natuwrh. Uiiapan-
ningen i. Tab. x. fig. n., more pointed in Pontobd. muricata Lbach. In
others these nodes are entirely absent, as in Pontobd. lubrica Gbubb ; eyes
seem not to be present.
Order III. Settgera.
Body annulate, provided with setae or with setigerous rudiments
of feet. External branchiae in most.
A. No external organs of respiration {Abranchia).
Family IV. Lumbridni. Branchiae none. Body provided with
setae, without rudiments of feet.
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ANNULATA. 229
ChoBtoffoster V. Ba£B. Eyea none. Fasciculi of setae ventral.
Rings sliglitlj distinct.
Sp. ChastoffOMter limnm V. Babb, Nw. AcL Acad. On, L. C, Nai. OwrioB,
Tol. zm. PL 7, pp. 6ii — 615, IVb. xxix. fig. 43 ; Duais Ann, dn 8e.
not. aec. S^. vni. Zocl, PI. i, f. 14.
JEjolosoma Ehrenb. Eyes none. Body distinctly articulated-
lateral fasciculi of setae in each joint. Mouth anterior inferior sur-
mounted by a lip dilated, produced.
Sp. ^do9oma ffemprichii EHBBirB. Symb, phy, Phytoaoa, Tah. T. fig. 7.
Pristina Ehrenb. Eyes none. Upper lip produced into a soft
bearded proboscis. Setae lateral.
Sp. Priabma hmguda Ehbenb. Symb. phydc, eveiiAr, Dec. I. Ac.
Nais MuELL. (exclus. sev. spec.) Eyes two. Setae lateral, long:
fasciculi of short setae on the belly.
Sub-gen. Stylaria Lail Pvohoscis frontal, stylifonu, soft.
Sp. NalM prohoicidea, Nereis lacuetrie L., Tbemblbt Mim, rar Ua Potfyptf
FL 6, fig. I, {MiUepied d dard) ; Robs. Ina, m. Tftb. 7S, figs. 15 a,
x6, 17, 18, g, K i, i; Tab. 79, fig. i ; MuBLLEB Ntaurffcteh, emiger
Wurmarien, b. 14 — 73, Tab. i; GRUiTHUiaBir Nov, Ad. Acad. Leap. Car.
Naiur. Curioe. Tom. xi. pp. 233— 148, Tab. xxxv.
Sub-gen. Ndis Lam. Proboscis none.
Sp. NaU mrpentina Gmbl., B0B8. Ins. in. Tab. xon. ; Mubllbb Notwrgeeek.
einiger Wwrmarten, b. 84, Tab. TV. &c
On the propagation of iVaii Bee B. Lbuokabt UngeeckleMiehe Vermekmng
hei NaU proboseidea, ArchivfOr Naturgeteh. 1851. Sohultzb on the same
subject, ibid. 1852, a, 3 — 7.
The genera noted above form a small natural group of vorms, for
the most part living in fresh-water, the Ifaidina of Ehbebtbebo.
Ck>mp. on these and some other genera P. Gebvais, JNote stir la
disposition syst^maUqns des Arm^lides chHopodes de la /amiUe de
NdU, BriUet, de VAcad, r. de Bruxelles, Tom. v. no. 1 y O. Schkibt
BeUrdge zwr AruU. u. Physiol, der Ndiden; Mueller's ^rcAtt?. 1846,
8. 406, <fea Besides the propagation by eggs, these fl.Tn'Tin^,|i^ are also
multiplied by spontaneous division. The most complete observations
on this point relate to Ndis proboseidea. In the last joint of a
simple Nais (which Mueller calls Jung-/er Natd, Virgin Nais) a
young Nais with eyenspots is gradually developed; it grows and
remains connected with the mother : sometimes on this a second or
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230 CLASS VII.
third daughter is found to be developed, which always arises more
forward (the last is the oldest, that which first came into being),
and ordinarily the first daughter already possesses the rudiment of
a little daughter before she separates herself; the vessels, the in-
testinal canal, the nervous cord run uninterruptedly through these
united animals ; at length the united or compound animal is broken,
and the eldest daughter (herself already a mother) separated herself,
after the mothe]>nais has made frequent strokes to and fro with
her tail On the eggs of the NdidcR see Duges Ann. des Sc, ncU. xv.
pp. 322 — 324. Six or seven eggs are enclosed in a common capsule,
a grey-coloured vesicle of j line in diameter.
[From later observations, as those of Leuckabt and Sghultzb
referred to above, the process of non-sexual multiplication does not
appear to be quite so simple as here described. The first and all-
important step is the development of a bud between two rings
nearly in the middle of the length of the body ; so that this now
consists of three portions, the anterior, the posterior, and the inter-
vening bud. All the three become distinct individuals, the first,
by developing its tail, the last its head, and the bud the head-
segments and anal portion in the same order of succession as in
development from the egg. Previous to the separation of these
three worms a new bud is usually formed in front of the middle
worm, and in front of it a third bud, &c., so that sometimes a chain
of many connected individuals is met with which all receive nutri-
ment (introduced by the mouth of the anterior member of the
chain) frx>m the intestinal canal common to them all. This process
appears to have been observed in other &milies also {AmphUrUcB,
Nereides), but would seem in all to be limited to the period pre-
ceding the sexual development]
EfttchytrcBua Henle. Mouth inferior, sub-terminal. Sexual ori-
fice in the eleventh ring of the body. Four fasciculi of usually three
setae short and uncinate in each ring. Body round, anteriorly
acuminate, posteriorly truncated.
Sp. Enehytraus albidtu Henlb, Mublleb*s Arehiv. 1837, b. 74 — 90, Tab. vi ;
a white worm two to six lines long, it lives in the earth and is especially
found in flower-pots.
Tubtfex Lam., Tvhilumhrictis Blainv. Body filiform, transpa^-
rent, doubly aculeate, attenuated at both ends, inclosed in a tube
composed of particles of mud and open at both ends.
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AXNULATA. 231
Sp. Tubifex rimdonm, LumMeu$ tubifex MuiLL. ; Tbimblbt Polfpes, PL 7 ;
fig. a ; Eneydop. nUth,, Veri. PI. 34, figs. 4—7 ; MuKLL. Zacl. dan. Tftb. 84,
figs. I, 2. This reddiBh litUe worm Utm at the bottom of ponds taxd becks ;
by the onion of many accnmnlated worms of this speeies red spots are
caused at the bottom of the water, which, on being touched^ immediately
vanish, for the wonns hide themselyes in the ground.
Samurts HoFFMEiSTER. Upper lip exsert, spoon-flhaped, Cli-
tellom little, distmct. Four fasciculi of five to eight Bet» in each
ring.
Comp. HoFFioiBTKR De$ vernnUnu quUnudam ad gemu Lumbrieomm
yerUnentibut, 4to. BeroUni, 1841.
Lumbricidus Grube. Body ronnd, with fonr rows of double
aciculse. Mouth inferior; a lobule resembling an upper lip, not
distinct from the following segment. Cingulum none. Segments
of the body numerous.
8p. LumbriculvM variegattu {LumMcui vari^atuB MuXLL.t) Gbubi in
Ebiohbon's Archivf. natwrgtich, 1844, s. aoo— 307, Taf. Yii. fig. 1 ; about
two inches in length ; through the transparent skin the motions of the
dorsal vessel fall of red blood, and of its blind digitifonn lateral appendages
which contract and expand in every segment, may be 1
Sub-genus Euaaoea Grube {EhynchekrUa Hofeil). First segment
(head) elongate, sometimes produced into a long thread
Sp. Euaacea JUirodru Gbube, Ebiohbon's Archw. 1844, pp. S04— 307, Taf.
yn. fig. I, in fresh-water like the former ; 3} inches long. Euaxet oUuri-
rattrii Mbkqe, £bichson*s Archiv. 1845, l^* m. fig. i.
Lumbricua L. (exclusive of species). Unterion and HypogcBon
Say. Body cylindraceous, attenuated at both extremities, obtuse
posteriorly. Mouth sub-terminal, under the exsert upper lip.
Sets not retractile, disposed in longitudinal rows. Clitellum or
cingulum, t. e, a tumid fleshy glandular zone mostly composed of a
Tarioufi number of rings in the anterior part of the body (saddle or
girdle).
Of this genus there are different species in Europe which were
formerly confounded under the name of Lumbricua terreHris. See
Sayignt Anodyae dPun Mhn, sur lea Lambriea, Comptea rendua dea
travaux de rinatiUut. 1820; Duges Ann. dea Sc. noL xv. 1828,
pp 289 — 294,tWrf. sea sfer. Zool. vra. 1837, pp. 18 — 25; Fitzinokr,
in Oxen's laia, 1833, pp. 549 — 553 ; Hoffmeisteb IHaa. de Vermibua
quibvadam ad genua Iwmhricarum pertinenUhua. Berolini, 1842,
(Ebighson's AriMv /. Nabwrgeach, 1843, p. 183) ; the same : Die
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232 CLASS VII.
bekan/rUe aHen avs der Fa/miiie der EegenuHirmer, mii Zeicfmungen
nach dem Leben, Braimscliweig, 1845, 4to.
On the anatomy oompare MoirritoBE ObaervatUma swr les Lorn-
hrica ou vera de terrey Menu du Museum, i. 1815, pp. 242 — ^248, PL
12 j J. Leo De etfmctwrd Lumbrici terrestris, Regiomonti, 1820,
4to, cum Tab, cen.; C. F. A. Mobben CkntimientcUio de strucbwra
anaiomica et historia ncUmrcdi LtumJbrici vulgaris sive terrestris
{A7inal. Acad. Ganda/vensis), Gandavi, 1829, cum talmlis, dec
The setee are short and rigid, in every ring 8, on each side two
pairs, so that eight rows run longitudinally on the body, four laterally,
and four beneath; in Hypogoeon Say. there is moreover another
row of single hairs in the middle of the back. The intestinal canal
is straight, with a membraneous pyriform proventriculus and a round
or spherical muscular stomach ; behind the stomach it is divided by
many transverse folds into blind pouches, which further back are
less developed, where also the intestinal canal becomes smaller
though on the whole it is wide throughout. In the interior of the
canal on the dorsal side is a band, which begins a little behind the
stomach, at this antenor end, as also at the posterior, runs to a
point, and consists of two membranes, of which the external is
yellow, the internal white ; irUestinum in irUestino Willis, typhla-
sole MoRBEN. This enigmatical part is probably a duplication of
the membrane of the intestine, an internal mesentery (Morren) ; it
may be compared with the valvular membrane of certain sharks ^
To the sexual organs belong in the first place three pairs of grey-
yellow saccules which are situated in the anterior part of the body
(in the common large earth-worm, LuTnhricus a^gricola Hoffmeisteb,
in the eleventh, twelfth, and thirteenth ring), and of which the
posterior pair is the largest. These parts are usually considered to
be ovaries, but Steekstrup, who here also denies Hermaphroditism,
supposes them to be testes in which the seed is formed with the
spermatozoa in cells, that may be readily mistaken for eggs.
H. Meckel maintains that these organs are in all individuals testeSy
and says, that the ovariay intimately conjoined with them, lie like
a brown-yellow lobe on each of these saccules. Four small vetdcles,
resembling barley-corns, placed more laterally (two on each side),
contain in the pairing season a white fluid with spermatozoa free
and developed : by most writers they have been signalised as the
^ Perhaps also it is fumiflhed with a yessel (Vena iMsmteriea interior); see
DuvKBirOT in the second edition of CaviSB, Zep. d'Anat, comp, Tom. V. 1817, p. 335.
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ANNULATA. 233
testes; Stebnstbup <hi the contrary thinks that they ought not to be
considered to be the parts where the seed is formed, but where it is
collected (as seminal vesicles in the male subject, as bursa coptUatricee
in the female). The efferent ducts of these vesicles open externally,
according to Savigny ; but later writers have failed to discover the
openings ; rather are they in connexion with the efferent ducts of
the yellow saccules ; these ducts fidl at length into a common canal
on each side backwards and end with two openings at the fifteenth
or sixteenth ring of the body. ^ At the origin of these two canab lie
two small irregular saccules, covered by a thin and gUstening mem-
brane, which according to DuG^ and Steenstrup are filled with
many convolutions of the efferent canal and form the passage of the
yellow saccules to the straight part of the canal which runs back-
wards \ Earth-worms are oviparous, not viviparous ; they pair
during the whole Summer, especially by night, when they creep
from the earth ; but how impregnation is effected, is not yet suffi-
ciently explained, since the apertures of the sexual oigans are not
brought immediately Together. The anterior portions of the two
worms lie next each other, but with the heads in opposite directions
(see in Morbek L 1. Tab. xxvu). Thus the part named by Willis
CliteUtim (saddle) in each of the two worms lies towards the place
where the sexual openings of the other worm are found. This di-
tellum is a roimd swelling of the body which occupies from six to
nine rings (in LwmJbrums agricola from the 29th to the 36th or from
the 31st to the 38th ring), and which during the time of copulation
is more strongly developed, and in young individuals ia entirely
wanting.
Sp. Lumbricus agricola Hoffm., LumbricuB ierrestris L. (in pftrt)i HoFF-
MEIBTEB Die bekannte ArUn aus der Fam, der Regenw. fig. i ; the largest
species in northern Europe, fix>m eight inches to more than a foot in
length.
Family V. Maldantce Say. Branchi» none. Mouth bilabiate,
inferior. Rudiments of feet provided with setae ; the three anterior
pairs without ventral pinna, the rest with a transverse tubercle,
supplied with uncinate setse, in place of a ventral pinna.
Ch/mene Sav. Body cylindrical, with few elongate segments,
^ Hie best description and figure of the organs of propagation in Lumbricut were
given by G. B. Tbbyibanus, ZtUtek. fOr PhytioL v. s. 154 — 166, Tab. m. ; see also
GmxmavEXjr, JffermapkrodiHtmui TUvoerdte, pp. 35 — 40, Tab. i. figs. 3—7, and H. Mxo-
KELin MiTBLLEB's ArehUfo, 1844, s. 480 — 483.
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234 CLASS VII.
the posterior extremity infundiboliform with margin usually denti-
culate. A membraneous tube covered with fragments of shells, open
at both ends, including the animal.
Sp. Clymene ampAueoma Sat. Dtter, deVEgypU, AnniL PI. i. fig. i; Gn^BiBr,
lowogr., AnniL PL lo, fig. i, from the Bed Sea. See fig. of other species,
Cuv. R. AnL H. iU., Annil. PL i«.
B. External organs of respiration.
* Tubulate.
Note, — The CephdUhrcmtMaUy or tubiccloue £dw. Annxdata, are
more imperfect than the roving or notobranchiate. It seems right
therefore to introduce them here, although the affinity by which
ArenvccHa is connected with the Lumbricmi points to a different
arraDgement.
Family VI. AmpkitritcB Sav. Head not distinct, eyes none,
body usually encased in a tube. ^
A. Branchiae anterior, more or less composite, with one, two
or three pairs.
SiphonosUyma Otto. Two larger tentacles (branchisB ?) and seve-
ral soft cirri around the mouth. Fasciculi of setae in double pairs
in every segment; the setae in the anterior segments extremely long,
directed forwards, glistening with gold. The worm not included in
a tube.
Sp. SipKonodofiva diploehaitut Otto Nov. Act, Acad, Naiur. Ourio§. Tom. x.
i, I Si I, p. 62S, Tab. 5t, in the Mediterranean at Naples; other species
have been described by KiLNX Eowabds, Gbubb and Rathkk ; see
Rathks BeUr. zur Fauna Norwegem, Nov, Act. Acad. Nahir. Ouriot,
Tom. XX. I, 1843, pp. 311— 219, Tab. xi.
To the same division also appears to belong the worm described by
Abildoabd in the Zoolog. domic. Tab. 90, as Afnpkitrite plumoaa, but
which differs from AmphUr. plumota of O. Fabbioius (Fauna grcenL
p. 188) ; Oksn formed from it the genus Pkertua^, (Lekrh. d, Zoolog. i.
B. 377) : Siphonottoma phbmosum Rathkx JBeitr. zur vertfi^ Anal, u, Physiol.
184a, p. 84, Tab. VI. figs. I — 7, BeiJtr, zur Fauna Norwegena, p. 3o8, Tab.
XI. f. I, 1.
^ The name Pheruaa was also given by Laxouboux to a genus of the class of
Polyps, of which the polypaiy alone is known and to which FUutra tvJMoM belongs ;
Hid. de$ Palypien JUxSble$, 18 16, p. 117 ; G. Johnston formed from Ampk, plumoM
the genus Flemingia, from which his genus Trophonia does not difo-. An», qf NaL
Hist. XYii. p. 394.
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ANNULATA. 235
NaU, — Chlorcema Dujabd. Ann. des So. not. sec. S^. Tom. xi.
1839, Zool. p. 288, Tab. 7, fig. 1, is a species of Siph<mo$toma beset
with Tilli secreting mucus ; comp. Siphon. vUlonun Rathke Faun,
Norweg, L L In SiphonoaUyma pltMnosum also the blood has a green
colour, Rathks L L p. 211.
Amphitrtte Cuv. (in part), Amphictene Sav. Mouth surrounded
\>y numerous tentacles, and covered by a denticulate velum. Setae
glistening with gold, in a double row in the anterior segment of the
body. Branchiae on both sides, two in the third and fourth segment
of the body, incurved, pectinate. The worm included in a thin
oblongo-conical tube made of sand cemented together by gelatinous
substance.
Sp. AmphUriie auriwma,^ SaMla granulaia L., Pectinaria Mgica Lam. ;
Pallas MUc, ZoU. Tab. ix. figs. 3—5 ; Ratilu, Beitr, z. vergl, Anat, u,
Pkytiol. Tab. y. : these worms, whose tube is known by fishermen under
the name of Sand-guiver, is met with on our coast Amph, cegyptim Say.
Ikaerip. de VEgypU, AnniL H, i, fig. 4, GulbuN Iconogr,, AnnH, PI. 1,
figs. 3* &C.
TerebeUa Cuv. (spec, of genus Terebella Gm.) Mouth bilabiate,
transverse; upper lip produced, surrounded by numerous long tenta-
cles. Budiments of feet with a double row of uncinate setae at the
ventral pinna, except the first pair. Branchiae ramose in the anterior
segments, which are without rudiments of feet. The worm included
in a tube composed of sand and fragments of shell cemented together.
8p. Terebdla conehUegaj Nereis conchiUga Pall., Mite, Zool, pp. 131 — 138,
Tab. IX. figs. 14 — 22 ; very common on our coast, where whole heaps of
the cases or bouses (generally empty) of these animals are met with. Tere-
beUa medusa Sav., GuiSbik leonogr., AnnUl, PL 2, figs. 2, ftc.
TerebeUidea Sabs. Four pectinate branchise.
Sp. Terebdlides Straemii Sabs, JBeskrivdser og JagttageUer wer nye % Hav^
ved den Bergeneke hyet levende Dyr, 1835, Tab. 13, fig. 31.
Sabella Cuv., Sav., Amphitrtte Lam. (sp. of SabeUa L.) Mouth
transverse, not tentaculated, situated amongst the branchiae. Bran-
chiae two flabellate, infimdibuliform or pectinate, spiral, large, with
bearded laciniae and a soft cylindrical filament at the base in the
iSrst segment of the body, which is without rudiments of feet. The
anterior pediform tubercles with uncinate setae at the ventral pinna,
with a fasciculus of subulate setae at the dorsal pinna; the posterior
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236 CLASS vii.
tubercles supplied with uncinate setse at the dorsal pinna, with a
fasciculus of subulate setae at the ventral pinna. The worm included
in a gelatinous tube covered with sand.
Sp. SaheUa pavanina Say. ; AmphUriie peniciUut Lam., Babteb NcAuwrh.
UUsp, I. p. 88, Tab. IX. fig. i ; Tubularia perUeiUuB, Zool. dan. Tab. 89, figs.
I, 4, in the North Sea; Sabdla magnifica Sav. ; TvhuUuia magmfca Shaw
lAnn. Transact, v. p. 128, Tab. IX. ; Sabdla {amphUrite) taurica Kathki
Fauna der Krym, Mini, dea Sav. ttrangen de VAeaH. imp, de Saini-Petenb,
Tom. m. 1837, p. 416, Tab. vni. figs. 8—15, fto.
Serpula L. Mouth situated between the branchiae, not tentacu-
late, transverse. Branchiae two, large, pectinate, flabellate, with
bearded laciniae and a cylindrical filament at the base of different
length in each branchia, the longer sustaining an orbicular disc or
infundibuliform operculum. Feet as in the preceding genus.
Calcareous tilbe procumbent, twisted or convoluted into a spire,
including the animal.
Sp. Serpula contortupliccUa L., Gufanr Iconogr., Annil. PL i, fig. i, (the
animal) ; Ellis CoraUinet, Tab. 38, fig. 2 ; Serpula vermieularia L., ZoU.
danie. Tab. 86, figs. 7 — 9, &o.
Comp. on this genus, which is somewhat differently determined and into
which SabeUa protula Guv. is also brought, A. Phiufpi in Ebiohson's
Archiv, 1844, "• iS^ — 19^*
Spworbia Lail
Sp. Serpula epirorhie, Spirorhie nautilcides Lam., ZooL danie. Tab. 86, figs.
I — 6 ; GuiBiN Iconogr., Annil. PL i. fig. 6.
B. Branchiae dorsal numerous.
Hermella Sav. {Ainphitrtte Cuv. in part), Sabellaria Lam.
First segment of the bodj supplied on both sides with a triple series
of very glistering tufts, the external very patent, the internal close.
The rudiments of the feet, in addition to setae, supplied ¥rith a
cirrus elongate, adhering above to the base, performing the office
of branchiae. Animals living gregariously, included in tubules
made of sand and fragments of shells, conjoined to form a common
honey-combed mass.
Sp. Ifermdlaaheolata, SabeUa alveolata L., Elus CoralUnee, Tab. xzxn. ; on
the English and French coasts. Formerly the bundles of threads beneath
the first segment were supposed to be gills. Milnx Rdwabds was the first
who indicated the true gills, on account of which this animal belongs to the
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ANNULATA. 237
AmUltdea donibrant^ of CuviSB, whUsty however, in a nfttunl anrange-
ment it might better remam with the AmpkitritO!, Ann. de» Se. not, sec.
S^. X. Zool, p. 108.
•• Naked, roving.
(Commonly NotobranchtcUe^ Dars^branches Cuv.)
Family YII. Arentcoke {Teletktufm Say.) Rudiments of feet
of a dorsal fasciculus of setss and a ventral transverse tubercle with
setae veiy minute, plane, incurved. Branchiae arborescent in the
middle of the body, with a double row at the sides of fasciculi of
dorsal setae. Head not distinct; eyes and jaws none.
Aremoola Lam. Body elongate, with segments subdivided by
transverse folds, incrassated forwards, becoming smaller backwards,
without setae or other appendages behind the last pair of branchin.
Mouth terminal supplied ¥rith a proboscis retractile, papillose.
Sp. Armieola pitcatorum, Lumbricus marinus L., NereitlumhrtcokUi, Pallab
Nov, Aa, Petrop. IL 1788, p. 123, Tab. v. f. 19, 19*; Hon PkU.
Transact. 1817, Pt. I. Tab. 3 ; Oksk, Isis, 181 7, p. 469, with fig. ; AuDOUiN
and MiLNS Edwasdb, Ann. des 8c. not. Tom. 30, 1833, PI. 22, fig. 8.
This specieB has thirteen pairs of gills. It lires in deep canals excavated in
the sea-sand, which the worm forms with its head, whilst the sand is
swallowed and passed through the intestinal canal; this worm is flesh-
ooloored, sometimes blackish (Arenicola carhonaria Liach), and exndes a
yellow fluid on being touched. Fishermen use it as bait to catch shell-fish
with the hook. Arenieola branehiaHs AuD. and £dw. 1. 1. fig. 13, has nine-
teen or twenty pairs of gills, and is smaller than the former. Arenieola
BoBckii Rathkk, Favna Norweffens, p. 181, Tab. vin. f. 19 — 22, differs from
the former species by the much more numerous gQHB and by the rings,
which lie behind the last pair of gills, possessing bundles of hairs ; also the
anterior part of the body is not incrassated like the former species. It
seems, therefore, that this species should form a sub-^nus.
It seems that the genus Scalibregma Rathke ought to be added
to the Areuicolse : it has four pairs of arborescent branchisd (in the
fourth, fifth, sixth and seventh segment) with a proboscis not warty;
L L p. 182, Tab. ix. figs. 15—21.
Family VIII. Ch<Btopterina. Anterior and posterior rudiments
of feet with a fasciculus of dorsal setse, without uncinate setoB, the
middle feet with a dorsal appendage, membraneous, large (bran-
chial?). Head not distinct; maxillae none.
Ckcetcpterus Cuv. (Worm elongate, included in a coriaceous
tube).
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238 CLASS VII.
Sp. ChoBtopteruB pergamentacew Cut., Milni Edwards Ann, det Se. not, Tom.
XXX. PL 22, fig. I, Cuv. JB. Ant. id. ill., Annil. PL ao, fig. a ; in the West
Indian Sea. A species of this genus occurs also in the Mediterranean.
Family IX. Peripaiina. The rudiments of feet are conical
tubercles, supplied with a fasciculus of thinly set setaB at the point.
Head distinct, provided with two cirri (antennaB) annulate, large, a
short proboscis, and two jaws.
Peripatus Lansdown Guilding. Body with few segments
subdivided by annulate folds, obtuse at both extremities, gibbous
above, plane beneath.
Sp. Peripatus iulifomut Lajnsdown GuiLDiNa, Zool. Jowmal, ii. PL xiv.
fig. I ; AuDOUiN and Milnb Edw. Ann. dea Sc. nal. Tom. XXX. pp. 41a —
414, PI. 22, figs. 5 — 7 ; West Indies. This animal would seem, according
to some, to he a myriapod insect ; the English author who first made it
known, considered it to he a mollusc. Milnb Edwabds, on anatomical
grounds, defends its reception into this class ; Ann. des Sc, not, sec. S^r.
xvni. ZodUtg. pp. 126 — 128. Branchice are not present, if the conical
rudiments of feet are not to he considered as respiratory organs.
Family X. Aricice. Body cylindrical, attenuated at both ex-
tremities, with head little distinct. Proboscis short, without jaws.
Rudiments of feet with cirrus usually single; branchiae either none
distinct or resembling cirri or lobes adhering to the base of the feet.
This small group, regarded by Audouik and Miuns Edwards as
a distinct &mily, contains imperfect ringed worms, which in part
belong to the Nereids of former writer& Where no special gills are
present, the cirri appear to serve for respiration.
CirrcOulvs Lam. Body elongate, roimd, with few dorsal and
ventral set® remote, and long dorsal cirri. The branchiae very long
cirri in the antenor part of the body.
Sp. Cirratvlua horealis, Lumbricva cirratua Mdell., O. FaBBIOH Fauna
gronU. pp. 281 — 283, fig. 5, Encydop, milk.. Vert, Pi. 34, figs. 10 — 12 ;
CirraL Lamarckii Am), and Edw. Ann. det Sc. naL xxvn. PL xr. figs.
I — 4, XXIX. pp. 410, 411 ; Grubs Kiemenwiirmer, 1838, pp. 32, 33.
Ammotrypana Bathke {Beitr, z. Fauna Norweg.) A genus
related to the preceding: it differs by defect of the long cirri (bran-
chiae) in the anterior part of the body.
Sp. AfMuoirypana aulogoiter Bathkb, L L p. 188, Tab. X. figs, i — 3, kc
Ophelia Sav. Comp. Edw. and AuD. Ann. dea 8c. not. xxix.
pp. 403 — 407. According to Sars the animal is so described that
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ANNULATA, 239
the dorsal surface is taken for the yentral, the anterior part for the
posterior; the author, Ann. des 8c. not. sec. s^r. vii. Zoologte, p. 247,
counts it amongst the Nereids.
Aonis Sav.
Gomp. AuD. and Edw. Arm. dei Se. nai, Tom. xxvii. pp. 400 — ^403,
PL xvm. figs. 9 — 13.
Artcia Sav. Body elongate, attenuated at both ends, with
conical head. The ventral oar of the anterior feet with a transverse
incised crest, of the posterior with a conical setiferous tubercle and
small cirrus, with soft branchial appendage. Dorsal cirri triangular,
plane.
Sp. Arieia Cuvierii AuD. and Eow. Ann, des Se. wU. xxix. p. 397, xzvir.
PL 15, f.5— 13» *<5-
Scolophs Blainv.
Comp. Obbstid Chifnl. Annml, doraibr, p. 199. Here also Beems to
belong the genua TravUia JoHHSTOir, Ann. of Nat. Hid. it. p. 373, PL xi.
f. 11—18.
Spto O. Fabr. Head ¥rith two very long antenniform tentacles.
Mouth inferior or subterminal, little exsertile and without jaws.
Body elongate, slender. Superior pinna with uncinate or capillary
setae, inferior with capillary setae; branchiae ligulate, dorsal, in the
anterior part of the body very large, in the posterior evanescent.
Sp. Spio seticomU O. Fabr., Bastkb Nat. Uittp. n. pp. 149, 150, Tab. xn.
fig. 3.
O. Fabkiotus, van dem Spio-OeseMecht, Seriften der Berliner Oeedheh.
naturf. Prmnde vi. p. 156. Kathkx, Beitr. zur Fauna der Krym, Tab.
vm. figs. I — 6, p. 421 (Spio IcevieomiB), Osbstbd Qnad, Annul, pp.
TOi, 103.
The genua Malaeoeeroe QuATBSTAaBB is distinguished by the defect of
eyes, Gviiaiv, Magae. de Zocl. 1843.
Family XI. Nereidce. Body elongate, slender, with head
distinct, supplied with tentacles (antennae) and eyes. Rudiments
of feet similar throughout the whole body. Branchiae not distinct
from the feet or small appendages of the feet, like lobes or tu-
bercles. Proboscis large, often armed with two homy jaws.
Goniada AuD. and Edw. Head conical ; with pinnae of segments
remote, each of them supplied with an acns and setae with conical
lobes or cirri. Proboscis large, famished beneath with a double row
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240 CLASS VII.
of homy denticles, without maxillae or armed at the point with two
maxillae.
Sp. Omiada etnerUa Aud. and Edw., Ann, des Sc. not, xxix. PL 15, figs.
1—4.
Ephesta Eathke. Head conical; with dorsal pinnae of the
segments mammillate, setiferous, the ventral supplied with a fasci-
culus of short setae. Proboscis large, clavate, smooth.
Sp. JBphena gracilU Rathkb, BeiiT, zu/r Fauna Norwegent, pp. 174 — 176,
Tab. vn. figs. 5—8.
Olycera Sav. Head conical, at the extremity with four tentacles
small, subulate, arranged in a cross. Dorsal and ventral pinnae
approximate, inserted in a common tubercle, supplied with acus and
a fasciculus of few setae. Cirrus at the base of each pinna; branchial
appendage simple or bifid in every segment, except only the anterior
and posterior. Proboscis large, usually with four jaws.
Sp. Cflycera Meckelii Aud. and Edw., Ann. det Sc. n<U. xxix. p. 363, xxvii.
PI. zrv. figs. I — 4 ; Glyeera albct, NereU alba Muellir, Zool. dan. Tab.
LZii. fig. 6. (Comp. Johnston, Ann. of Not. Hid. zv. p. 148, Rathkk,
Beiir. swr Fawna Narweg. p. 173.)
Pollicita Johnston {Behryce Thompson). Comp. Ann, of Nal,
Hist XVI. pp. 4 — 6.
Nephthya Cuv. Head truncated anteriorly, supplied with four
small tentacles. Dorsal and ventral pinnae remote, setiferous, in-
creased by a membraneous lobe. Branchiae ligulate at the dorsal
pinnae. Proboscis large, furnished with conical tentacles and two
maxillae not exsert. Body linear, elongate, with terminal style.
Sp. Nephthya Homhergii Cuv., Aud. and Edw., Ann. det Sc. not. zxix. PL
XVII. figs. 1—6, Cuv. jB. Ani. id, ill., Ann€l. PL XV. fig. 1 ; Neph. UmgiMe-
tota O1B8T. GromL Annul, p. 195, Tab. vi. figs. 75, 76, (perhaps the same
as NepUi, cUiaJta Rathkb, Beitr. z. Fawna Norwegent, p. 1 70).
PhyUodoce Sav. (Ranzani). Head small, supplied with two
eyes, and four or five tentacles, the fifth unequal, very small, remote.
Tentacular cirri in the anterior segments. Setigerous tubercles
undivided, with dorsal and ventral appendage lamellose, branchial.
Body terminated by two styles. Proboscis thick with small tentacles
at the orifice, without jaws.
Sp. PhyUodoce lanUnoaa Sav., Aud. and Edw. Ann. dee Sc, not. xxix. p. 344,
PL 16, figs. I — 8; PkyOod. davigera, Nereis viridWMuELL.^SulaUaf Bav.,
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ANNULATA. 241
AuD. and Edw. L L p. 9484 PL 16, figs. 9—13 ; PkyUod. aaxicola Quatsb-
rAOBB, GuiBDr Maga§. de Zod. 1843^ AniUl, p. i, PI. i, fto.
Fsamathe Jobnst.
I<nda JoHKST.
Cknnp. JoHKSTON, ^nf». 0/ iVol. JTtif. iv. pp. ia<^— 131.
Myricma Say.
Alciopa AuD. and Edw. Eyee large, lateral Tubercles lobate
(glandular) at the base of the pediform tubercle& Other characters
a» m Fhyllodooe.
Sp. Ale. ReynaudU AuD. and Edw. Ann. de$ Sc, fiat. xxix. pp. 136 — 338,
T9h. TV, figs. 6— II ; Comp. A. EJtOHK, ZooL wnd amat. Semer^bmngm
nAet die Aleiopm, EaiCHSOv's AtMv, 1845, 8. 171— 184, Tab. n. Bendea
the deacsriptbn of aome new apedea thia memoir oontaina alao anatomical
details, amongst which, especially those upon the eyes are worthy of notice
(see above, p. 317). The glandular appendages of the rudimentary feet
AuDOunr and Milkb Edwabdb consider to be gills.
JBesione Say. Head broad, truncated, furnished with four lateral
eyes and four small tentacles. Long tentacular cirri at the sides of
the head. Setigerous tubercles of the segments undiYided, with
dorsal and ventral cirrus filiform, the dorsal long. Body oblong,
with segments not numerous. Proboscis large, without jaws.
Sp. Metione tplendida Say. Dsmt. de VEffifpU, Annil. PI. in. fig. 3, Ou<B.
Ictmogr. Annil. PI. 8, fig. 3.
Nate. — Genus Halvmede Rathke is distinguished from Hedone
by three branchiA (lobed appendages) at each of the pediform
tubercles {BeUr. z. Fofuna Nww. pp. 166 — 169).
SyUis Say. Head bUobed, anteriorly emarginate, with four
eyes placed in transverse row, and three tentacles, thin, moniliform.
Setigerous tubercles of the segments undivided, with dorsal cirrus
long, moniliform. Probosds without jaws. Body elongate, slender,
with numerous segments.
Sp. Sfi,u nwMlane Sat. Deicr. de VEffyple, Awn4l. PL lY. fig. 3, Guia.
Icanogr, Anna. PI. 8, fig. i ; SifU. Maeuloea Edw., Cuv. jB. Ani. id. iU.,
Annil. PI. 15, figs. I, &c.
Nereis CuY. (spec, of gen. Nereis L., Lycoris Sav. and Lycastis
Sav.) Head anteriorly attenuated, with four eyes arranged in two
series and four short tentacles, the external larger,conical. Subulate
VOL. I. 16
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242 CLAas VII.
tentacular cirri at the base of the head in the first segment of the
body; two cirri in each segment. Proboscis thick, cylindrical,
armed with two homy exserted jaws. Body elongate, with nume-
rous segments.
A. The dorsal pinna of the feet confluent with the ventral or
not distinct, without branchial appendages. Lycaatia Sav., Aud.
and Edw.
B. The dorsal pinna distinct from the ventral, with aciculus
and bundle of setsB at the extremity of each, and appendages or
lacinias supplying the office of gills. Lycoris Sav., Nereis of
Authors.
Sp. Nereit wmtia, I/yeor, nwidia Say. Deter, de rigypte, AfmSl. FL iv. fig. 3,
On^RiN leonogr.f AnnSL PI. 7, from the Bed Sea ; NereU peiagica L.,
Basteb Natuurk, Uitap, ii. Tab. ti. fig. 6, OERffriD OrcnU. Annul, p. T75,
Tab. IV. figs. 53, &c.
Heteronerda Oerst.
Sp. Heierfmereu arcUca OxBST. L I. Tab. iv. fig. 51; NtreU gra/ndifolia
Rathks, Beitr, z. Fawna Ncrweffma, pp. 155, &o.
Family XII. BunicecB, Body elongate, with numerous seg-
ments. Rudiments of feet supplied with a single pinna, a terminal
setiferous tubercle, and two cirri. Proboscis armed with seven,
eight or nine homy jaws. Branchiae in some none (cirri supplying
the office of branchiae), in others above the dorsal cirrus adhering to
many segments of the body, pectinate.
A, BranchiaB not distinct £rom the cirri.
• Head covered by the first segment of the body. Jaws nine.
Genera Aglaura Sav., (Enone Sav.
Comp. Gu^iN Iconogr, AmUl, PL 6.
*♦ Head not covered by the first segment of the body.
Lmnbrineru Blain., Aud. and Edw. Head obtusely oonical,
with tentacles either none or two inserted into small tubercles at
the posterior margin. Jaws eight.
Sp. Lufnhrineri$ d*OHtygn{i Aud. and Edw. Ann. des Sc, not. Tom. xxyii.
PI. 12, figs. 9 — 12 ; Lumbrineria pedinifera Quatbsf.^ GuilB. MiMffOM, de
Zool. 1843, Annil. pp. 6—8, PI. n. figs. 3—8, &o.
Lysidioe Sav. Head broad, small, with three short tentacles.
Jaws seven.
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ANNULATA. 243
Ck>mp. ArD. mod Edw. Ann. de$ Se, not, XTin. pp. 353—^57, Tom.
xxvn. figs. I — 8.
B. BrancliM distinct. Jaws seven.
DiopcOra AuD. and Edw. Head small, with nine tentacles.
Filameoda of branchisB nnmerotis, placed on a petiole twisted
spirally.
Sp. Diopatra amloimmma Avd. and Edw. Ann, def ^. not. xxvin. pp. 329,
330, PI. X. figs. 6—8.
OnupAis AuD. and Edw. Head small, ftimislied with seven
tentacles. The first two pairs of pinn« larger, directed forwards.
Branchiae in every segment, except the first two, the anterior of a
simple filament, the posterior pectinate.
Sp. Onuphrii tremiia AuD. and Edw. Jim. dt$ So* not. zxnn. pp. 936,
317, PI. X. figs. I — 5; OnuphrU Etchrichtii Oibst. OnmU. Annul, p. 173,
Tab. ui. figs. 33 — 4I, fig. 45. These worms live in cases ; thejr are some-
times ooTered with bits of shell, like that of TerAeUa, as in the last-named
species, from which I suspect that Onuphria conchiUga Sabs, BeArivtUeTf
1835, pp. 61 — 63, PI. X. fig. 38, does not differ.
Eunice Cuv., AuD. and Edw. {Leodice Sav.) Head distinct,
round or lobate, with five tentacles. Pectinate branchiss above the
dorsal cirrus in most of the segments, or in the anterior part of the
body.
Sp. Eunice ffigantea. Nereis apkrodiiois, Pall. Nov, Act. Pehy>pol, Tom. ii.
pp. 339, 330, Tab. ▼. figs. 1—7, Cut. jR. Ani. id, ill,, AnnSl, PL 10 ; this
species becomes more than four feet long : — Eunice sanguinea, Nereii
tanffiUnea Montagu Trane. of Linn, Soc, Tom. xr. pp. 30, 31, Tab. 3,
figs. I — 3 ; Eunice cmtennaia Sav., Deecrip, de VEgypte, AnnSl, PI. y. fig. i,
Gu&iN, Iconogr., Annil, PL y. figs, i, &c.
Family XIII. Amphirwmacece, Body depressed, oblong. Head
furnished with two or four eyes and mostly five tentacles. The
pediform tubercles supplied with setae only, not with aciculae.
Branchise arborescent or fasciculate in all the segments of the body,
the three or four anterior excepted, placed at the sides of the back.
Proboscis without jaws.
Amphtnome BeuguiAre. (Spec, of Aphrodita Pall., of Terthella
Gm.)
A. Pediform tubercles with undivided pinna and single drrus.
Tentacles five in the head; caruncles behind the base of the middle
tentacle none. Branchifle ramosa
16—2
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244 CLASS YII.
ffipponoe AuD. and Edw.
Sp. ffipponoi C/audichaudii AuD. and Edw. Ann. det Se, nai. Tom. xx. 1830,
pp. 156—159, PL ni. figs. 1—5, Gn^BiH Iconoffr., AnnSl. PI. 4 bis. fig. 3,
in New HoUand (Port Jackson).
£. Pediform tubercles with double pinna, remote, each provided
with a cirru& Caruncles behind the base of the intermediate tentacle
at the dorsal surflEUse in the head and anterior part of the bodj.
JEktphrosyne Sav. Head with single subulate tentacle, and two
eyes. Branchise made of many branched appendages, arranged in
a row between the dorsal and yentral pinna.
Sp. Suphrotyne loMrtaia Sat. I>etcr. de VEgypU, Annil. PL n. fig. i,
GuiBiK Iconogr., Annil. PL 4 bis. figs, i, &c.
AmpMnome AuD. and Edw. Fleione Sav. Head with five
short tentacles, and four eyes. Branchiss ramose, or fasciculate at
the base of the dorsal pinnss.
Sp. Am^pikinome nOtrata Tall. Miae. Zool. Tab. via. figs. 14 — 18, from the
Indian Ocean; Ampkinome ca/nmeulata Pall. fig. 11 ; oomp. A. E. Gbubb,
De Pleione eanmeulata Diss. Zoot. e¥m tab. cm. Begiomontani, 1837, 8to.
Chloeia Sav. Head supplied with five tentacles and two eyes.
Branchise like a tripinnatifid leaf, placed on the back, remote firom
the base of the pinnse. Two terminal styles at the posterior part of
the body.
Sp. Amphinome capiUata, ApkrodUa Jlava Pall. MUe. Zool. Tab. Tin. figs.
7 — II ; Guy. M. Ani. id. iU. Annil. PL 9 ; in the Indian Ooean, from
Amhoyna, fta The bundles of aetce are yellow, the gills, which nearly
resemble leaves of Fern, are pmple- coloured.
Family XIV. Aphroditacece. Body in most depressed, oval.
Head supplied with tentacles usually five (2 — 5) and with four eyes.
Dorsal and ventral pinnae distinct, furnished with acus, a fascicle of
setse and a ventral cirrus. Squamss {Elytra Sav.) in most, in place
of cirri, placed on the dorsal pinnsB that alternate with dorsal pinn»
supplied with a cirrus. Proboscis usually armed with four jaws.
Palmyra Sav. Dorsal squamsB none.
Sp. Palmyra awrifera Say., Aud. and Edw. Ann.duSe. fiat. Tom. xxyii.
PP* 445i AA^t PL X. figs. I — 6, from the island ifoicri^tw.
Genus Spinther Johxstok ; is it of this &mily ) Body oval,
with back convex, abdomen plana Head indistinct Eljtra none.
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ANNULATA. 245
Tubercles of the feet aunilar in all the segments^ supplied with
a ventral cirrus only.
€k>mp. JoHKBTOH, Awn, of Nat, Hi$t. xn, 1845, pp. ft— 10, Spmther
imuooHdei, PL n. figs. 7 — 14.
Sigalion AuD., Edw. Body depressed, elongate, with numerous
segments. Dorsal squamss together with dorsal cirrus in most of
the segments; the anterior segments that are without squams alter-
nating with squamiferous segments.
Sp. SigaUon Mathilda AuD. mod Edw. J mi. det Se. not. Tom. xxm,
pp. 441—443, PI. IX. figs. I— 10 ; SigaUon boa JomrSTOV, Aim. nf Nai,
Hiit. n. pp. 439 — ^441, PI. ZXin. figi. 6—15, (probably the SMne ipecies m
SigaUon Iduna Rathke, BeUr, twr Fauna Norweg. pp. 150—155, Tab. n.
figs. 1—8 ; Gomp. also Sigalion EtteUa Gu£b. Magat. de ZodL 1833, AnmU,
PL 3).
Aconites AuD. and Edw. Body elongate, with numerous seg-
ments. Branchial tubercles at the base of the pediform tubercles
in all of them, dorsal squam» large, the squamiferous segments,
without dorsal cirrus, alternating with segments supplied with dorsal
cirrus. A coriaceous tube longer than the bodj concealing the
worm.
Sp. AcoiU$ Ploei AuD. and Edw. Jim. da Se. not, zzvn. p. 437, PI. x.
figa.7— 14.
Foh/odofUes Benieri, Aud. and Edw. {Phyllodoce Banzani).
Head small, with three tentacles and two pedunculate eyes. Dorsal
cirri in the segments destitute of elytra, which alternate with squa-
miferous segments; ventral cirri short, subulate in all the segments;
distinct branchiae none. Proboscis with two subulate cirri and warty
margin, with four large denticulate jaws*
PoljfodonUg JRanzanU, Phyllodoce maanllota Ranzaht, Opft$eoU $cieniiifiei,
Bologna, 181 7, Tom. I. pp. 105 — 109, Tab. rr. figa. 4—9, MedhenraiMan.
Polynoe Sav. Body in some oblong or oval, in others slender,
elongate. Head with four eyes set on tubercles, and five, sometimes
four, tentacles. Jaws large, homy. Branchiae and dorsal cirri
in pediform tubercles destitute of squama, which alternate with
squamiferous feet.
Sp. Polynoi $gmmaia, AphrodUa squamaia L., BAsna, NakmHk. UUap. n.
Tikb. TL fig. ▼. Pallas Miaeell. Zool. p. 91, Tab. vn. fig. 14 ; PUgnoi
kgvie, Aud. and Edw. Ann. dee Se. naL xxvn. p. 491, PI. ix. figs. 11—19,
GuiBiK, Iconogr. Annil. PL 9, figi. 4, Ac.
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246 CLASS VII.
Aphrodtta L. (exclusive of some species), Halithea Sav. Head
concealed under squamas or setae, furnished with two pedunculate
eyes and three tentacles. Jaws small or none. Branchiae and
dorsal cirri in feet destitute of squama, which alternate with squa-
miferous feet. Body oval, depressed.
* DoTBtd iqwima xakBA.
Sp. AphfrodUa kyOrix, HalUkea kpOrw Say., Aud. and Edw., Ann. det Se,
not. xxvn. PI. 7, figs, i — 9. .
* * Dorsal iquama coTored by » stratum of vUlote deke,
Sp. Aphrodita actUeata L., Basteb^ Naiuwk, Uittp. Tab. vi. figs, i, 3, Pall.
Miic, Zool. pp. 77, &o. Tab. vii. figs, i— 13 ; Phytalus Swammebd. Bibl.
not, Sea-Mouse, Ordinarily five or six inches long and an inch and a half
broad. The hairs on the two sides of the body are glistering, green and
red, or playing with all the colours of the rainbow ; the back is clothed
with a felty coTering, that consists of interwoven hairs. When this
covering is opened, five nearly circular plates {fqwm/a, dytra) are seen on
each side, which partially cover each other, and of which the middlemost
are the largest. If two consecutive plates be separated, there are seen on
the ling that lies between them, small longitudinal nodes, which are parted
by a pit, and are provided outwards and backwards with pectinated
appendages as though torn at the margin (the gills). In the Atlantic
Ocean, the Meditenranean, &c.
Note. — The genus Sagitta Slabber, Quoy and Gaim., whose
place is uncertain, seems to approximate closely to the Annvlata.
Body not annulate, elongate, pointed at both extremities, supplied
with lateral pinnae and a terminal truncated pinna; head distinct
from the body by a neck, with two eyes, and homy teeth on both
sides. All the individuals hermaphrodite.
Comp. A. Kbohn Beobackhmffen ueber die Sagitta bipunckOa, Hamburg,
1844, 4to; R. Wilms, ObeervoHonee de SagiUa, Berolini, 1846, 4to.
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CLASS vni.
INSECTS {INSEGTA)\
LinnjEUS united all those invertebrate animals, whose body is
divided into rings and which have feet consisting of different joints,
into a single class, that of Insects. Together with the class of
Ringed-Worms they make up one of the four principal groups into
* There i« no class cyf aDtmals on which more hu been written than that of Insects ;
comp. Latbullx, who has given a sketch of the history of Entomology (Mim. dM
Muteum, Tin. 1823, pp. 461 — ^481, also J. N. Eisxlt, Oeaehiehte, Sydemaiik vmd
lAUerohtr dtr Imedenkunde, Leipzig, 1836, 8yo, and J. PlBcaEOEiiOK, BtUiograpkie
Enicmoloffiqu^f Paris, i^yj, 1 vols. 8to). As observers deserve to be named especially
R^UMUB {M^moires powr tervir d mist. de$ Inaeetes, Paris, 1734 — 1749, 6 vols. 4to},
A. J. ROSSBL {Inteettn-Bdngtigving, Ntimberg, 1746 — 1761, 4 vols. 4to, with exceUent
eolonred figwes) and G. Db Gbxb {Mim powr Mervir d VHist. det Inteetet, Stookhofan,
1751 — 1778, 7 vols. 4to). The anatomy was excellently treated by our great country-
maa, SwAMMKaDAM {Biblia naturte, Leiden, 1737, i vols. foL), and afterwards by
P. Ltoket {Trail4 cmatomique de la Chenille qui ronge le baii de Saule, la Haye, 1769,
4to). In this part also in the present century Ramdohb, Tbivibanus, ' Hmou),
BBAimr, Jj£ov Dufoub, Stbaus, Dbuokhbih, Blahohabd, Nkwfobt, ko. have
published many important investigations. Comp. the Articles, Inaeetet, by Audouin,
in the JHdionnaire clcuaique d^Hiti, nai. Tom. vm. 1825, pp. 559 — 579, a^d /nMcto,
in ToDD*B Cyclopcsdia of Anat. and Phynol, n. 1839, PP> ^73 — 994-
As general works, introductions and systematic handbooks, the following, amongst
ethcrrs, may be used with advantage :
J. G. Fabbicii, PhUoaophia Enkmoloffica, Hambuigii et Kilonii, 1778, 8vo.
Ejusd. EfUomologia SystemaHca, Hafniae, 1799, iv. Tom. (6 vols.) 8vo; Index
alph/xbettcw, in J. C. Fabbicii, Erdomol. Syatam, Hafoise, 1796, 8vo. SupplemeiUwn
BtUomei. SydemoHcaiy Hafisin, 1779, 8vo.
P. A. Latbbillb, HiM. naiur des Cnutaeiet et dee Inaectei (suite anx CEuvres de
BuFFON, par SoKNivi), Paris, i8oa, 1803, 14 vols. 8vo, avec fig.
Ejusd. Genera Cnutaceorwn et Insectorum, Parifdis et Argentorati, 1806, 1807,
IV. Tom. 8vo.
W. KiBBT and W. Sfshos, Introduction to Enionudogy, 5th Edit. London, 1818,
4 vols. 8vo.
H. BuBiOEiBTBB, ffandbuck der Entomoloffie, i Bd., Alffemeine Entomologie, Mit
16 Steindrflcken, 4to. Berlin, 1833, 8vo.
J. O. Wkstwood, An Introduction to the modem Cflamjleation of Ineedi. Lon-
daBf 1840, 1 vols. 8vo (with many woodcuts).
As engravings we may, besides Roxbxl, especially notice the figures of the Icono-
graphic du lUgne animal, by Gu^iK, and those of the Dictiomn. dee Sciences nalureUea,
which are also to be found in A. M. G. Duk^il, Coneiderations ginSralea eur la Claue
det Inaeetes, Paris, 1833, 8vo.
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248 CLASS yiii.
which Cuvier, as stated above (p. 33), divided the entire Animal
Kingdom. We allude to the Type of Articulate Animals, which
may be separated into two divisions.- The first division includes
articulate animals without jointed feet {apoda, the AnmdaUi)^ the
other articulate animals with feet {Gondyhpoda). Again, articulate
animals with limbs (Insects of LiNNiBUs) are divided into three
classes, of which the first, immediately to be treated of, retains the
name of Insects; the two other classes, those of the Arachnids and
of the Onistaceans, comprehend those animals which LiNNiEUS con-
sidered as ¥ringless Insects.
Insects (in the narrower meaning of modem sjstematic Zoology)
are mostly provided with wings; but the presence of these organs
of motion does not constitute the character of the class. That is to
be sought for in the head distinct firom the trunk, to which two
antennsB are attached, and in the Respiration by means of air-canals
distributed internally through the body and generally divided into
very fine branches. The first of these characters distinguishes the
Insects firom the Arachnids in which the head and thorax form a
single piece, and which have no antennae, the other distinguishes
them firom the Crustaceans, whose respiratory organs are gills or
other external appendages.
The names Insecta, notched animals, and in Greek JWofia, have
all the same meaning. From the last is derived the word Entomo-
logy (Insect-Science).
The species belonging to this class are very numerous: in this
respect no other class can be compared with Insects. In treating
therefore of this class we must keep within strict limits, that we may
not too greatly extend our work in the estimation of those who take
less interest in this special part.
We will first describe the external structure of these creatures a
little more precisely The body of the six-footed Insects, which
make up by far the largest portion of this class, is separated into
three parts : head^ trunk, and abdomen. On the head, besides the
parts about the mouth, the antennae and eyes are distinguished.
By Antennm are understood moveable jointed threads, which,
unconnected with the oral organs, are attached to the head, usually
close to the eyes. The number of joints is very difierent; in But-
terflies, for instance, very large, in Beetles, mostly eleven, &c. All
insects in the perfect condition or last period of life have two
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IN8ECT8. 249
antennsB. Their relative size is very different. Sometimes as in
Locu8t€By they exceed the length of ttie body, in others again they
are veiy short and almost concealed beneath the eyes. The fonn
likewise is different : the antenn» are said to he^t/brm when they
are thin and of the same thickness throughout: clavate when they
baye a knob at the end formed of thicker joints, as in butterflies
{Papiltones), &c.
The eyes (oculi) are either simple or compound. The simple
eyes are named eye-points {ocelli, atenmiata) : they look like smooth
shining points placed usually in a triangle behind the larger eyes ;
they are seen in Bees, Wasps, &c. The larger eyes are composed
of numerous six-sided fa^ettes, and are occasionally of such magni-
tude (as in Dipteraj LtbeUuloB), as to meet, the head seeming to
consist almost entirely of these two eyes. In some instances the
number of fa^ettes is surprisingly great: Leeuwenhoeck counted
8000 in the eye of a fly, Straus nearly 8820 in that of a cockchafer ^
The oral apparatus {prgtma cibaruxy Trophi) consist of six
principal parts, of which four are in pairs and move transversely,
whilst two face each other above and below. Of these last the
uppermost is the upper lip {labrum) : it is homy and fixed by a
joint transversely to the most anterior part of the head. The part
of the head to which the upper lip is fastened is named head^shidd
{dypeas, in French chaperon). The undermost part, facing the other,
is named the under lip {labium)', it closes the mouth below: is
composed of two parts of which the inferior and more rigid is named
diin {merUum), and the superior, generally membraneous, tongue
{ligula). Sometimes the liguJu has two lateral lobes {paro/glossoB).
The remaining four parts are known as upper and under jaws. The
upper jaws {fnandibuke) are two, placed immediately beneath the
upper lip: they move transversely from within outwards, and are
often very hard. The under jaws {maxilloB) are ordinarily softer:
are placed beneath the mandibles, and also move laterally, but are
less serviceable for cutting the food small than for holding it in the
mouth and conveying it to the gullet to be swallowed. In the
Orthcptera there is a membraneous valve which is fastened to the
maxilla. It is called the helmet {galea) of the lower jaw.
In addition to these principal parts there are also fselers {palpi,
^ See pUte XX. of Swammbbdam's B^id der natmir, where the nmple and com-
jMimd eyei of a bee are figured.
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250 CLA^ Vlll.
aniennuhB)^ jointed threads, attached to the under lip {palpi labicUeSy
s, posteriores)^ and to the nnder jaw {palpi maxiHares), The upper
jaws in Insects are not provided with palps.
In masticating Insects, as Beetles, Locusts, &c., the parts of the
month, that haye been described, maj be best and most readily
observed. In those which feed by sucking fluids the structure is in
appearance very different; yet even here it may be observed that
nature remains true to her plan, and that she has provided the
suckers not with different but with modified oral parts. We are
indebted to the illustrious Savigny for the knowledge of that plan^
The sucking Insects possess oral organs which are named Tanguet
Beaky Sucker and Snout.
The Butterflies {Oloseaia Fabr.) afford an example of what has
been called tongue, or spiral tongue {lingua, lingua apiralia). It is
a canal, occasionally of great length, composed of two laminss which
are corneous or membraneous, on the inside excavated and round
externally. When at rest it is rolled up and concealed between two
palps. This was almost the entire amomit of what was known of
the oral parts of Butterflies. But Savignt pointed out in addition
two minute upper jaws, placed at some distance from each other, and
little, if at all, adapted for motion or mastication. The upper lip is
small and membraneous. The lamin» of the tongue, as Latreille'
had already shewn, are in fact nothing else than greatly elongated
and extended lower jaws. Their base is united to the head and upper
lip, and bears a palp composed of two or three joints. The two larger
palps which include the tongue and conceal it when at rest are
seated upon a triangular homy under lip.
The case is similar with the suctorial apparatus of the Hemiptera^
{Bugs, Gicadoe, &c.) named beak {rostrum). It consists of a homy
sheath {vagivui) in which setae are contained {seUe rostelli), that at
first sight appear to be three in number. The two lateral setae are
elongated upper jaws: the hair in the middle is double, and consists
of two similarly elongated and united under jaws: the under lip,
usually jointed, forms the sheath. In the same way in Diptera
{Flies, &c.) the under lip forms the snout {proboscis). In its interior
^ J. C. SAVioifT, Mimoire9 9ur la Animaux mm verUhres, Paris, 1816, 8to. li^re
fiMcicule.
* Latbbilli, ffitUfire tuUurdle det Onuku^ €t de$ In$ecle$, An. zn. 8to. T. ii.
p. 140.
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INSECT8. 261
are aetsB, like as in the beak, which fona the sucker {haustdlum).
A triangalar upper lip covers the basal piece of the beak in Hemt-
ptera, as it does that of the snout in Dij^era.
To the head succeeds the trunk or thorax. This part consists
of three pieces, of which each bears a pair of feet. The first ring is
namf d Prothorax, the second Mesotkorax, the third Metathorax. In
four-winged Insects the anterior wings are placed on the middle
piece, the posterior wings on the hinder piece. The wings of Di-
piera are placed on the mesothorax. The inferior surface of the
trunk is called breast {pectus), on which there is sometimes fixed a
pointed elongated appendage, the breast-bane {sternum). The
shield {scuteUum) is a part found on the upper part of the thorax
behind (at the mesothorax) stretching between the wings ^
The feet are attached on the inferior surface of the body : in the
hexapod Insects eyery ring of the thorax carries a pair. Between
the sternum and epwieran is an articular cavity (acetabulum). The
first joint is termed hip {coxa, eondylus) ; sometimes there is a small
and yerj moveable piece between the epimeron and coxa {trochanr
terium, trochafUin Audouin), but it is usually wanting or has coa-
lesced with the coxa. The second joint is termed Trochanter, it is
very small and mostly annular. Then comes the thigh {femur), the
stoutest, and often also the longest joint of the leg. To it succeeds
ihib shank {tibia) more slender, and in general flattened laterally.
Last is the foot {tarsus), consisting of many joints placed in a line
like the small bones of our fingers. The number of these joints is
different in different families; occasionally, in certain coleopterous
' The upper surface of the thorax {donum of AuDOUnr) may be named notum, the
under sm-fiKK sternum, and juBt as the entire thorax is divided into three rings, so abo
sk prono^tmKod prottemvm, metoncium and meMottenwm, metanotwn and meUutemum
may be distinguished. Moreover, each ring of the thorax consists of definite special parts,
which, however, are not distinctly seen in every ring, whilst some coalesce with others,
or by the greater development of others are suppressed ; these parts aie a tiemum on
the under surfaoe, on either side an epidemum as a chief part, and behind this an
epimeron; and, finally, on the upper four pieces placed behind each other, to which
AuBOUiK gives the names of prcescutwm, tcuivm, tcutdlwn and pott-sctOeUum; thus
there are properly three seuUUa, but what is usually named tevleBum is a part of the
memmatum; at the sides of the scutum the wings are attached. Gomp. on this interesting
sulject AuBOunr, Rdi^erehe$ amatomiquei aur U ihcrax ites Antmatm oHictdSi, Ann, des
8c. not, I. 1824, pp. 97— 135» 4i<5— 432, W. 8. Mao-Lkat, Comp, AntU. of thoraa in
wnged InteeU, Zoolog, Journal, No. 18, or Ann, dea 8c, not. xxv. 1832, pp. 95 — 151,
with remarks by AusouiK and Nbwfobt, Todd's Cfyclopeedta, n. pp. 911 — 924.
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252 CLASS YIII.
insects it is not the same in the first two pairs of feet and in the last
pair, yet in most Insects' the number is five. In some coleopterous
insects, the penultimate joint is extremely short, and was in conse-
quence overlooked formerly. The last joint of the foot usually ends
with two hooklets, or claws : in addition, its inferior surfiace is often
covered with fine hair, to attach it to small inequalities which even
the smoothest objects'present. Sometimes these hairs are set on
two or three delicate membraneous appendages {cushions, jndviUi)
which the Insects mould to the surfaces over which they run. In
this way flies can move upwards on mirrors, or with head down-
wards on smooth ceilings, as is seen daily ^.
Besides the feet, wings also are placed on the thorax of volant
insects : on the meso- and meta-thorax, as stated above, when there
are four : when only two, on the meso-thorax. They are set on the
dorsal surface, and may be compared with the elytra or squamos in
Aphrodita : with the wings of vertebrate animals (Birds, Bats), which
are only modifications of the anterior limbs, they have only similarity
of use : they are not modified feet : they exist contemporaneously
with feet and are independent of them*. Wings are membraneous,
arid, usually transparent, composed of two laminae grown together
at the edges ; these laminas are expansions of the skin like the
parachute extended between the fingers of Bats and between the
ribs of flying Lizards {Draco). Canals (improperly named Veins
or Nerves) run between the laminse, and are more or less numerous,
more or less branched. These veins are branches of the air-tubes,
which lie between two wide homy semicanals of the upper and
under laminae that compose the wing. In some species the males
alone have wings. Bees, Wasps, Butterflies, &c. have four wings.
In the Diptera, besides the wings there are two parts which may
be considered as traces of hind-wings, called poisers {haUeres) ;
they consist of a little button with a pedicle, and are often covered by
a membraneous scale {squanM haUerum)\ The anterior wings are
^ Blaokwbll, Semarht <m thepuMUi of InmcU, Trantad, of theLinn, 8oe. Vol. zvi.
Pt. 3, pp. 487— 49«-
' Oksv names the wings of insects gills; the djftra of Goleopten he oonsiden, less
happily, to be giU-coven ; they must have the same anatomical interpretation, (Bodett-
tung), as the under-wings. LehH>ueh der Naiimrphiilo9opkie, ni. 181 1, s« 971; tho
same work entirely revised. 1843, "• 3'^-
> See Aupouiir, Diet, dan. ^HiM, mtt, n. pp. 140—149, at the word JBoIoiicwrt,
and Kkwfobt, 1. 1. p. 916.
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INSECTS. 253
in some insects harder, homy and opaque; thej are then called wing^
covers (elytra), and the nnder-wings, nsually larger, are when at
jrest folded transversely beneath the covers and concealed (as in
Beetles, Golecptera). In other instances the under wings disappear,
and the wing-covers coalesce by their inner edges (elytra coadtt-
nata), Hemdytra is the name given to the anterior wings, when
homy or coriaceous at the base but membraneous towards the apex
(in Hemtptera, as Water "SCorptOTia, Nepa cinerea, &c.)
The hinder-body (abdomen) constitutes the third portion of the
body of Insects, and usually consists of nine rings, of which
however the last are in some instances so much concealed, and in
others so small or so fused with the preceding, that they seem to
be entirely wanting. As the organs of sense have their seat in
the head, and those of motion in the thorax, so do the principal
organs of vegetative or organic life reside in the abdomen.
The digestive organs present differences according to the
Orders and Families. Here the comparative length of the intes-
tinal canal does not always depend, as in vertebrate animals, upon
the nature of the food, and many species that live on animal
substances have a longer and more convoluted canal than others
which live on plants; in Grasshoppers for instance (Orylltj
Locustce) it is almost straight, though these insects live exclusively
on vegetable food. In those Insects whose body consists of
unifom rings (as the myriapods) and in vermiform larvas of Insects
with a complete Metamorphosis, the intestinal canal is straight, or
makes only few and inconspicuous curves. The intestine has the
greatest length in proportion to the body in certain Coleoptera and
Hemtptera. In the last it is at least twice, often four or five times
the length of the body (ex. gr. in LygcBue apterua Fabr.) ; in
Guxida omt the intestinal canal is about ten times as long as the
body*. Amongst Coleoptera the Scarabceidea, to which the common
cockchafer belongs, are remarkable for their very long and tortuous
intestinal canal, which in Copris lunaris measures ten or twelve
times the length of the body.
The membranes or coats of the intestinal canal are, first, a thin
covering, which without sufficient reason has been compared with
1 Uov DuFoxTB, JReeherehet antU. et jihyncl. tur let MlmipUrtt {Extrait du Mhn,
de$ javant Hrangen, Tom. iv.) Paris, 1833, 4to. p. 99, Fl. vm. f^, 95.
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254 uLAias VII r.
tiie peritoneal covering of the intestines in yertebralg ndmab;
next, a muscular coat of longitudinal and transverse fibres ; then s
white, smooth membrane, a layer of areolar tissue probably corre-
sponding to the tunica propria of the intestine in vertebrates, but
which is often beset with minute glands in transverse rows : and
lastly the innermost membrane, an Epithelium, that occasionally, as
in the muscular stomach of the Orthoptera, is found hard and homy,
forming the teeth or sharp plates with which the stomach is armed.
In the intestinal canal of Insects several parts are to be
distinguished : but it is much to be wished that writers in the
names given to them had been careful to preserve greater uni-
formity. The first part is the oesophagus, it has often an expan-
sion named crop {inglumes) ; next follows a muscular stomach
{veniriculus musculosuSy der Kaumagen^ le gSsier, the gizzard) ; it is
found in the Orthoptera and amongst the Goleoptera in the genera
Staphylinusy Dytiscus, and the family of the Carabid^, and is
remarkable for the great development of the innermost coat, for
the projecting plates, teeth or booklets of corneous tissue which
serve for bruising the food ; it is usually folded and has a round,
more or less spherical, form*. Then comes a long cylindrical
stomach in which the proper digestion proceeds. Leon Dufour
names it ventricule chylifiqijLe ; Ramdohu calls it simply the sto-
mach, which name appears to me to be sufficient and preferable to
the other. This organ is always present, and beneath its termina-
tion the vasa urinaria (of which hereafter) are always inserted •.
To this succeeds a longer or shorter, sometimes (as in the Hemi-
ptera) a very short canal, the sm^ll intestine {intestinum tenue),
which is continued into the short large intestine {infest crassum)y
having occasionally a ca^ium or expanded portion when the con-
nexion takes place obliquely and at the side*.
^ Ll^N Dufour has ako discovered a muscalar stomach in Tomicui typographut.
Ann. det Sc, not. iv. p. io8.
' Raudohr names it FeUtenmagen (plicated $tomaeh, amatut), a very ill-ehosen name.
* Mabokl BR SxRRRS considered the stomach to be duodenum ; in that case many
insects must have no stomach at alL The name of Crop (fahot succeniurU) by whi<^
Straus denotes this part in the Cockchafer is not explicable.
^ We are indebted to L^N Dufour for most of the investigations of the intes-
tinal canal in Insects. They were preceded by those of Ramdohr, who published a
work on the subject, (Ahhandhmgen Hher die Verdauungtwerheuge der Inaeden, mit
30 Kupfertafeln, Halle, 181 1, 4to).
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INSECTS. 25a
The intestinal canal of Insects is connected to the other parts
of the body partly by a large quantity of fat (the adipose body, of
which below), and partly by numerous branches of air-tdbes, aad
so retained in its place.
In veiy many Insects Scdivary Olands ate pieflent; they are
placed at the commencement of the integtiaal canal. In CcleopUra^
for the most part, they are wanti]^; Ramdohb found them in
Curculio {Cryptorhynchus) lapaiki, L^ON DuFOUR, besides in other
Curctdumiday also in Bloft^ Diaperis, Mordella and some other
Coleoptera; moreover in the other orders of Insects they are
present in by far the greater number of Families, probably in all
Orthoptera^ Hymenopteray Lepidopteray Dtptera and Myriapoda.
Amongst the Neuropiera they are wanting in LibeUula and Ephe-
meray amongst the Hemtptera in Aphides, It is veiy remarkable,
and not easily explicable, that in Panorpa amongst the Neurcptera
die female has no salivary glands, or more correctly only small
rudiments of them, whilst the male has them largely developed^.
They have here the form of long convoluted canals (three on each
side), which towards the end are turned upwards, and becoming
thinner terminate by blind extremities. This form of blind con-
voluted canals occurs also in the salivary glands of some other
insects, ex. gr. of the Lepidoptera; but it is by no means general,
for in the Hymencptera and Orthoptera these organs appear commonly
as blind sacs grouped in clusters. Microscopic investigation has
demonstrated in these salivary vessels aad glands, as in other
glands, a layer of epithelial cells with i\uclei^.
Below the inferior orifice of the stomach in Insects very fine
vessels are implanted, the so«called Malpighian vesselsy which in
former times were generally looked upon as organs for the prepara-
tion of bile {vasa h&patica) — an opinion still maintained by LiON
DuFOUB, Owen' and other writers. It is, on the other hand, the
^ Our meritorious countiyman BaAirrs first made this interesting obeerration,
Ti^dschr. voor not. Oeaeh, en Physiol, vi. 1859, ^^* '73 — '9^- ^^ ^^ afterwards also
made known by L^n Dufoub {Mimoirea prisenUa d VAcad. mydU de» Sc, yn. 1841,
PP* S^^f 583, PI. II, fig. 1^9) who overlooked, however, the nidimentaiy salivaiy
glands in the female.
' See the beautiful investigations of H. Mkokxl, MvzLhKB^BArchiv. 1846, s. 7$ — 35.
* [It is not to be inferred that OwKN holds this opinion now : his Lectures were
published many years ago, and a new edition of them is now in the press.]
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256 CLASS vin.
opinion of most writers of the present day, that they correspond to
the kidneys of the higher animals, whence the term nsed above
p. 264, {vasa urinaria) for these vessels. Besides other grounds for
this opinion, it is supported by chemical investigation^. These
vessels appear to be present in all Insects, with the exception of
Aphidii amongst the Hemipteray where it has not been possible to
find a trace of them. Their number is very different, and seems
to be on the whole inversely proportional to their length ; they are
short and very numerous, more than twenty, in the Hymencptera and
Orihoptera, and in LibeUula and Ephemera amongst the Neuroptera.
Here they are arranged in a ring round the intestinal canal which
they perforate, whilst at the free extremity they terminate coecally.
In Ghyllotalpa said AchetaFABRAhey {all into a common canal before
opening into the intestine. In the remaining Insects there are
usually only four or two of them present {Diptera, Hemtptera,
many Colecptera), or six, as in other Coleoptera (the Heteromerdtay
Tetramerttta, and Trimerata), When there are only two, they
form a loop on each side of the intestinal canal, which seems to
arise from the fusion of two vessels ; and so open by four termi-
nations into the canal. In those Coleoptera which have six, they
are also attached to the inferior extremity of the intestinal canal
(the Rectum) y but do not open into it there; they run upwards as
very fine vessels between the coats of the intestine and terminate
blindly*.
If we consider these organs as Kidneys it becomes uncertain
whether Insects have a Livjer ; for the idea that these vessels may
represent at once both Kidneys and Liver (whence it has been
proposed to name them vasa urino-itliaria) is not, as appears to
me, the result of comparative investigation either anatomical or
1 See Bkhookb's Ph^ndogitche Untenuchunffen Uber dU thierische ffauakaUung der
Imeden, Tubingen, 1817, 8vo. Gomp. Wubzeb, Chemische Unterauekung da Surffes,
v)eieher nch in den aoffenanrUen Oallenffifdtten dea SehmeUerlifigt der Seidenrauste
b^findet in Meckel's ^rc^tv. iv. 18 18, b. 113 — 915. Also Chbvbbul found in the
matter of these vessels potass, ammonia and uric add; see Stbaub ContideratUmM
gMralea eur VAnatonde dee Anim, orHctdie, auxqueUei on a Joint VAnat, detaipHve du
MdoUnOha vfdgarie. Paris, 1818, 4to, p. 951. In a £«caiifw little stones have been
found in these canals consisting of uric acid, AvDovnt Ann. dee 8c, not, le S^r.
Tom. V. 1836, p. 199. 0. Vehlobbk found in lanrsB of Lq^idoptera {Sphinx Uguehri)
no uric acid in these yessels, but hippwric acid, as he informed me by letter in 1843.
' L^N DuFOUR, M6m, ewr lee vaieeeavec hiliairee dee Ineedee, Ann, dee 8e, not.
ae S^rie, Tom. xix. 1843, PP- i45— >8i, PI. & — 9.
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INSECTS. 257
physiological, and would never have been entertained but for the
attempt to reconcile two conflicting views, and which ought always
to be distrusted when it interferes with more extended enquiij.
But if we suppose an organ answering to the liver to be alto-
gether wanting in insects, then it must be proved that the separa-
tion of bile is more important in the animal economy than the
excretion of urea, before an argument can be borrowed therefrom
against the function ascribed to the Malpighian vessels* We do
not forget that by respiration and the elaboration of bile the
quantity of carbon in the living body is diminished, and that from
the large development of the respiratoiy organs in insects the
-excretory office of the liver is in a great measure dropped ^
Nevertheless it is still highly probable that parts, whose function
agrees with that of a liver, are not altogether absent in Insects.
In the first place we might here refer to the great quantity of
fat — the adipose body — ^situated between the skin and the intea*
tine, which invests every organ and is of very great extent, more
especially in larvae whose respiration is less perfect ; the carbon
and hydrogen which in other instances is combined with oxygen to
quit ^e body by respiration, here forms that provision of com-
bustible matter so necessary in the animal economy for the support
of respiration, especially in the case of Insects, which as Nymphs
take scarcely any food. Since then this production of fat exerts
the same influence on the composition of the fluids as the separa-
tion of bile, it is not to be considered as a proceeding entirely
arbitrary if some recognise in the adipose body an analogon of
the liver^« The adipose body consists of a multitude of minute
sacs or vesicles bound together by air-tubes which spread them-
selves as a fine network on their surface. In the second place, coecal
appendages are seen below the muscular stomach in the Orthoptere
(eight in ManttSy six in Qryllus^ two mAcheta) which involuntarily
call to mind the appevidices pyhriccB of osseous fishes : they pro*
bably secrete a fluid that performs the office of the bile in diges-
tion^ In other insects, finally, as in the Carabici among the
1 BUBHBI0TKB Momdb. der EwUmal. i. p. 403.
> OuBN Ukrb, der NnlwjpkUinopkie, m. 181 1, n, 170 (atte Avfa^, a. 435).
* ThatiheBe bUnd appendages ariae from an immodiate extension (protrusion) of the
intestinal canal is no proof, as LioK DuForR suppose?, that they cannot be secretoiy
VOL. T. 17
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•258 CLASS Tin.
Colscptera the entire stomach {ventrumlechylifique'DvFOJJB) is beset
-with numberless conical or filiform saccules, giving a floccolent
•aspect to its external surface. It may be, that these parts, whose
office was formerly supposed erroneously to be the absorption of
nutrient fluid from the intestine, prepare the bile : but it seems
more probable that they serve to separate the gastric juiced
The Heart of Insects has the form of a long vessel that
terminates behind by a blind extremity and lies above the intesr
tinal canal on the dorsal surface. This daraal vessel becomes
narrower forwards, after it has curved slightly downwards. The
smaller part may be considered to be an arteiy, whilst the wider
posterior portion answers more closely to the heart of other crear
tures. In this posterior part are different lateral openings, mostly
eight or nine pairs : and in front of each opening is a valve formed
by a duplicature inwards of the wall. In the diastole of the heart
the blood flows into it between two sets of valves, of which the
posterior pair come into apposition, whilst the anterior lie folded
against the wall and so permit the onward moticm of the blood.
Systole and diastole succeed each other alternately, moving along
the length of the dorsal vessel from behind forwards. SwammeBt
DAM long ago, and Straus in more recent times noticed irr the
dorsal vessel longitudinal and transverse muscular fibres, the latter
forming the innermost layer. Surrounding the heart is a space which
some writers consider to be a sintts venostis; it is covered by lateral
muscles, flat, and of triangular form, which have their broad base
towards the heart and fix it in its position {les ailes du Omar of
Ltonet). From behind, the blood flows through the lateral
openings into the heart, and moves forwards ; from before, it flows
from the aorta between the organs, especially along the course of
orgsnfl. Mim. pri$enUt, vn. p. 302. In LeutoptU also unongst the Bymmojitra two
•uch bHnd saccules are met with ; L^ir Dufour, ibid, p. 544.
^ The great unoertainty which prevails concerning the interpretation of the
secretoiy organs in the lower animals, Ib a necessary consequence of the fact that the
selftame secretion, as we learn from comparative anatomy, may be e£Pected by very
differently formed glands ; see J. Muxlleb's Handb, der Pkntujl., n. Buch, Ahaekn, %
(i. Bd., s. 457, 3tte Aufi,) Chemical investigation alone can here afford light^ and
a beginning of the enquiry has been made in invertebrate animals in these last yean.
C. SoHXiDT*s Investigations: Zur vtrgUichendm PhyHologie der leirbeUoien Thiere,
Braunschweig, 1835, deserve^ therefore, our thanks, and make us hope for further com-
munications.
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die air-tabes, in regular streame backwards. These stzeams of
blood on the outside of the heart were first observed by Cabus
twenty years ago in the three fin-shaped caudal processes of the
larva of Agrwn ; afterwards he observed a similar motion of fluid
in the imperfect wings of the Nymphs ; the blood-globules (accord-
ing to many, rather according to V^BLOREN, the fieit-globules), which
swimming in a clear fluid indicate the direction of the current.
The later observations of various authors, on transparent larve
principally, have ascertained the phenomenon in Insects of every
order — and it may therefore be ccwifidently accepted as general. A
question which requires further investigation for its solution is
this, whether the circulation is effected in vessels, as ex. gr. New-
POBT and BowERBANK believe, or in free spaces between the
organs, without special walls. The writers who maintain the
latter opinion, allege that the Aorta has an open termination in
the Head. In the Myruypoda^ besides the dorsal vessel, there are
Btill others present; amongst which a trunk that lies upon the
nervous cord in the abdomen, ought to be mentioned. In the
Butterflies also Treveranus discovered on the ventral surface a
vessel, lying on the nervous cord and running longitudinally, from
which on each side numerous transverse branches arise ^. New-
port found this vessel in the genus Sphinx^ and thinks that the
blood flows in it backwards, as it does forwards in the aorta. This
last author discovered in this same genus, and in certain Ccleoptera
branches firom the aorta in the head, but was not able, on account
of the delicacy of the parts, to follow their further course*.
The Bespixatory organs of Insects are their air-canals {tracheeR)^
* ZeitBdvt, far Phenol, iv. «, iSs^, 8. i8i — 184, T»f. xiv. fig. 13.
' Comp. on the donal voflsel and the dreulatioii of ioiectfl Ltokst, Trakl AwU. de
la Chenille, pp. 413, Ac.; on the fluid contained in it, ihid, pp. 496, 437; Hxbold,
Pkytiol. UnUrsuchungen iiher das JiUckengrfast der Insecten, Marburg, 1893, ^^o- i
Stbaus Anal. camp, dee Anim, articuUa, pp. 345 — 358 ; J. Muklleb, Nov. Act, Acad,
Ccee. Leop. Car. Tom. xn. 2^ 1835 (on a connexion between the dorsal vessel and tlie
ovariev) ; C. G. Cabus, Enideckung eines eif^achen vom Jffenen ave hesehleunigten
JShUkreisUntfeB in den Larven neUfiUglieher Imeden. Mit 3 Kupfert. Leipzig, 1897,
4to ; Waoveb, BeolHtehlungen Hb, d. Kreielauf det BhUei «. d. Ban dee B&ehengrfdetee
hei den Inteeten, Okxn's Iris, 1839, s. 310—331, Taf. n. ; Kkwpobt, Todd's Etkcydop,
n. pp. 975-— 981. The tieatwe of our ezceUent M. C. Vbbloben, crowned by the
Branels Academy of Scienoes in 1844 (1) is impatitntly waited for ; I have made wa
of tho obsenrations he had the goodness to communicate to me when treating of the
dorsal TSflseL
17—2
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S60 CLASS vxtr.
which are usually filled with air bj external openings {a^mata).
These canals have three coats: an external, loose, transparent
memhrane, in which fibres and scattered points (cell-nuclei) may
be distinguished ; a middle, composed of a flat, homy, sometimes
brown or yellow elastic thread rolled spirally : and an inner coat
which is composed of chittne, a continuation of the exteraal skin,
«nd is thrown off at every moult^. Through the elasticity of the
spiral thread the air-canals are duly kept open : its turns lie close
to each other, and so the appearance of rings is produced, as in the
wind-pipe of mammals (this the representation of the trachesB of
PediculiAs in Swammerdam, Bibl natur. Tab. I. fig. VII. resem-
bles too closely) ; but the similarity is only in appearance ; there
are no absolute rings, but only the turns of a single uninterrupted
thread. Each branch, arising from a stem, has a new thread,
whether the branch proceeds laterally firom the stem, or two branches
arise at the end of the stem ; this thread is finer than that of the
stem, and in the terminal branches is only visible when very
highly magnified. From being full of air, the canals, when
Insects are dissected under water, have a silvery splendour, and
present on account of the extreme fineness of their branching a
very beautiful appearance to the observer*. Usually the air-canals
divide, like arteries, into continually finer branches. In some
Insects however there spring firom a large stem on every side
throughout a greater or less extent extremely fine and numerous
branches (as ex. gr. according to Leon Dufour, in PtionuB^ firom
the double stem which lies between the last atigma of the thorax,
and the first of the abdomen). In Nepa and tUmaira saccules are
seen in the cavity of the thorax, between which similar fine
branches {j'eiia mirabtlia) of the air-canals lie, and which are sur-
rounded by a muscular coat'. Care must be taken to distinguish
these saccules from the sacculated dilatations of the air-canals them-
selves, which are met with in flying Insects in the last period of
^ It has not been uuide oat, tshr tsl know, whether the innermost membnuie of
the MT-tubee is present in those insects also which have no stigmata^ bat gill-plateSy as
the UrYVd of Ephemera, for instance.
* M. MALPiam, who first made use of the names of tnu^^a and Migmata, sajs,
^* TaaHa edfruatifiealio korvm vasonim, torn mira implicatianet ut nilpfdchriui ecmpiei
'pomi." Jh BomJbyce, p. la. Opera am, Tom. ii. Londini^ 1687, fol.
' JAov DUFODR, Reck, tur let Eimiptirte, p. 153, PL XTiii.
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INSECTS. 261
iSttm life, and which beyond doubt are of serrice in diminishing
weight during flight These dilatations are oval or pear-shaped, and
occasionally a tabular trachea proceeds anew from their further
side. In the ApiaruB amongst the Hymenoptera^ the two lateral
main trunks of the air-canals in the abdomen are in this way
converted into large reservoirs of air.
The stigmata are present in different numbers in the hexapod
Insects, but it is rare to find more than nine pairs of them; in
Ihftiscua amongst the Coleoptera and in Locusta amongst the
Orthoptera there are ten pairs (Burmeisteb Handhuch der Ento-
mologie I. p. 175). Also in Chryllotalpa I foimd ten pairs, three in
the thorax and seven in the abdomen. These air-slits axe small^
generally oblong fissures (like button-holes), often surrounded by a
homy ring (jperttrema) with a cavity behind them which again, by
a second fissure whose posterior half can be retracted by muscles,
leads to the air-canals. In other instances there is no peritrema^
but the stigma is formed by a fissure between two lips, whose
edges are beset with hairs. Sometimes there are in the cavity of
the st^ima special moveable homy plates {epiglottides Straus),
which can close the entrance of the air-canal that proceeds fix>m it.
By means of the oblique position of the lips, of which one often
projects over the other, by means of the narrow opening, and of
the hair or down on their edge, the entrance of dust or other
small bodies into the stigmata is prevented, whilst the air alone ia
admitted as through a sieve. From every air-slit, or its cavity
(vestibule) there arises an air-canal {trtzchie cTorigine Stbaus).
which divides into nxmierous branches (in Scolopendra), or proceeds
transversely after having given off one or two lateral main-stems.
These main-stems running along the length of the body, (in most
Insects there is only one on each side,) receive all the canals that
spring from the air-slits or fissures, and connect them together.
They give off the numerous branches which spread through every
part of the body. The distribution of the air-canals after the
manner of vessels is interesting; by such a disposition of the
respiratory organs in Insects, the atmospheric air has access in
equal degree to every part of their body ^ But it is too much to
' " In no^ a 6on^mUibu9 tangwmi hmmm pulmomt petit . .^mwaeeUi nen Ma
mn^fmni$ m/oim in pubnonm eovifiuH, $ed invena vi^ jndmonet ip$i, VMorum Hto, 4n.
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363 ciiASS vin.
conclude from hence that the circulation of blood is unnecessary m
Insects, and consequently does not exist. The circulation of blood
has not respect to respiration alone, it is not merely for the con-
version of venous blood into arterial ; it is necessary that arterial
blood should circulate that it may serve for nutrition and secretion.
Many Insects live in water : but of these the greater number
breathe atmospheric air ; like whales amongst mammals some come
to the surface of the water for that purpose. But usually there are
special arrangements for conducting the air, so that the Insect can
remain under water. This is seen ex. gr. in the larvae of Diptera^
which live under water; those of Cuhx have at the posterior
part of the body a lateral canal with fine hairs at the orifice ; the
larvas of Stratiomys have a canal at the end of the abdomen, whose
orifice is fringed with a circlet of plumose hairs ; the genera N^>a
and Ranaira have a tail composed of two filaments at whose base
are two air-slits*. These water-insects die in a few hours if the air
has no access to the water. Other Insects breathe in the water itself,
that is, they breathe the air that is diffused through the water, as
fishes do by their gills. Such Insects have no air-slits : the air
must therefore penetrate the walls of the trachecB, which to that
end are spread out either in filiform or capillary appendages (in the
larvae of Chfrtnus, of Semblts, the nymphce of Chtrtmamus) or in
leaf-like plates at one side of the body [Ephemera) y or at the
extremity of the abdomen {Agrion). These parts have been termed
Otlh*/ they are not found in perfect Insects. Gills of this sort,
firom which blind air«tubes arise, occur in the rectum of the larva
of LibeUula as five rows of plumose incised leaflets. From them
arise six longitudinal stems, of which two, larger than the rest,
univenum eorpvs ditperguntur, wic itf nngida partes o/eria particfdat per ptdmonet el
eangtUnie pwiionei per arterioi recipicU,*' Malpiohi Anatome plmUearum, Op* om. t.
p. *5.
^ Figures of CkJem in Swaiocsbdaic, J^iiN. fuK. Tkb. xxzi. figs. 4, $; <iii SttvUamifB,
ihid. Tab. zxxiz. ; of Nepa, in Dufoub, 1. L The abdomen of N^pa and Banatm
has besides three pair of conspicuous, but dosed, auMilits, in which very large branches
df air- tubes terminate with blind enda.
* This nomendaiore is only in part oorrect The proper respiratory argxBM of
Insects, the air-tubes, belong to the category of lungs, whether the air penetrates by
external apertures (eUgmata), or the tubes be filled with air from endosmotic action.
The air in iket is in the inside^ and the stream of blood (akmg the trachea) on the
outside^ and this relation is just tBe reverse of that which preyaili in gills.
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IK8ECT8. 368
"beoonie aftenraids the main tnuikB ci the fexSdci Lisect which art
in. connexion urith the air-«lits^.
Obserrations have shewn that Respiration in Insects effects die
same chemical changes of the air, as in higher creatores ; respijra-
tion is more active, the need of air greater and the production of
carbonic acid moie abundant in the perfect Insect than in the
larva. In the perfect Insect, moreover, respiration is performed
principally bj the air-slits of the thorax, which are larger than
tliose of the abdomen, whilst in the larva that function is distri-
buted more equally amongst all the stigmata. This fact is in
connexion, with the development of tke thorax and with the
mechanism for motion affixed there in the perfect Insect Accurate
investigations have shewn that Insects, at least under certain cir«
cumstances, have a proper toarmih, and that thej can raise the
temperature of their body remarkably by motion, or by volimtary
acceleration of respiration^
The sexes are distinct in all Insects, and the eggs are not
fertilized, as in fishes, after they are laid, but union of the sexes
must precede the laying of the eggs if they are to prove fruitful.
A remarkable peculiarity has been observed in Plant liee {Aphide$)^
^ Comp. on the rospiratoiy organa of insects, besides MalfiohIi SWAmuBDAX,
Ltokit, Stbaub and other writers already cited, C. Spbinokl CommenJUirivM dt
parHbuM, fuibus Inteeta ipiriium ducuni, LipsiA, 1815, 4to. cum tdMi$; SuoEOW,
JUtpit%Uiom> der Jnmdm, wtbesondtre Hber die Ihrwtrupiraiion der Ae$ckiw ffrandU,
HxuanrosB'B Zeitsch. /. die organ, Phynk, 11. 1828, s. 44 — 29 ; BuBMiisns Eandh,
der SnUnwd. i, s. 169 — 194 (a very careful reyision of the observations of others and of
his own) and Newpobt, PhU, Tram, f 836, Pt «, pp. 519—566 (or in ToDD*B Oydop,
n, pp. 984-— 990). We refer also to the beautiful figures in hiova TraiU an, deia
CkmUUj PL XXI., and Straus AnaL des anim, artie. Fl. 7, in order to give an idea of
the minute division of the air-tubes. Mabokl db Sibbxs has figured the trachee and
alr^sacs in some Orlhopiera {TruxaUi, Mantis) in Mim, dm Mushtm, lY. PL 15, 16.
' Already in 1793 Vauouklin had made experiments on the respiration of Insects
(LoeuHa viridimima), Comp. also G. tt. Trsyibavub, Vmwehe Hber doe AikemhoUn
derniedem Hkiere, Zeiieckr. f. Phenol, iv. 183 1, s. i— S9i and Nxwtobt, PhiL Trane,
L 1.9 for the speeifio wanath, which was formerly denied by J. Davt, against whose
observKtkms Nobili and Mbixohi had already advanoed objections (Ann, de Chim, ei
de Phffwique, 1831, Octofare, pp. 307—910). All animals, LntBlft justly observes, are
warm-Uooded, but only is sudli as brsathe by lungs (better, in mammals and \Mb), is
the specifie warmth entirely independent of the external temperature. IHe ifrgan,
Ckemie in ikrer Anwendung amf Ph^friol, «. Pathol. 1849, s. so. Comp. also Bbbtbold,
Neue V€r$nehe Hb. d. Temperatw der hkUMtige TKieirt, Oottingen, 1815* "- 35* 3^i
s. 43.
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264 CLASS vnt.
where a single impregnation suffices for many families in sncoes*
sion ; the males are not observed until the end of summer or in
autumn ; they impregnate the last family, consisting of wingless
females, which without copulation would be barren. Their eggs
remain during the winter on branches of trees, and in spring
produce only female plant-lice which without copulation are prolific
and viviparous. Bonxet, to whom we owe this discovery, found
that in the space of three months nine successive generations were
produced without copulation^.
Amongst the Myriapoda the Chilopods have only a single ovary,
in form of a long sac situated beneath the intestine. In the
remaining Insects there are two ovaries. Sometimes they have the
same form of tubes or sacs {Forficulaj Ephemera^ Stratiomys) ; in
some flies the sac is very long, rolled spirally like a watch-spring,
and separated by many transverse partitions into cells^ In most
Insects each ovary consists of a number of tubes {ffoinea evt^rea
Leon Dufour). Sometimes these are situated around a sacciform
dilatation from which the oviduct arises {ovaria bacoata), as in
Melo^ L. and Lycua {colecpteray. Or these tubes are situated
lengthwise along the origin of oviduct {ovaria ramaaa)^ as in
Cicada^/ sometimes on one side only, like the teeth of a comb, as
in Pha87na and TerUhredo {Atkalia). But in by fer the greatest
number of cases, these tubes are situated at the beginning of the
oviduct like the leaflets of a digitated leaf, at the end of a common
stalk {ovaria digitaiay Jhadculata), Such ovaries are seen in the
X^epidoptera, where each of them consists of four tubes. The
number of these tabes is however very different, not only in the
different orders, but even in the same order, and occasionally in the
same natmral family ; whilst, ex. gr. Bombyx and Xylooopa {Hymen--
cptera) have four, in the Honey-Bee are more than one hundred*. In
^ 0. BoinnR', TraUi iFIntedoloffie, i. Obaervatumi wr les Puemmi, Psaris^ 1845,
lamo. G^wfret i. 1771, 8to. Buyau has obtained eren eleven suooesriye generations
wtthoat oopnlaiion ; Ann, da Se, nat, v. 1835, p. 334. There are also some examples
of the same phenomenon in insects of other orders. Bubmbistib, 1. L s. 336, 337.
* IUauvub, Mhn. pour 9ervir A VSitt. da Int. iv. PL 49, f. 7 and 8.
* Bbavdt and EAncBUBa, Medmn, Zoologie n. Tab. rm. fig. 1 k, Mdoe wrU-
fftOui, Tth. zix. figs. II, 15, LyUa vetieaiana; L^v Dufoub, Aim, da. Se. noL vi.
PL 18, ^. I, LffC¥$ rm/lpenmi.
* LiSoB DuFOUB, ffimipUra, PI. 17, fig. 189.
' LiOB DuFOUB, Mim, prSanUM, Tom. vit. p. 408. According to Swamkkrdam,
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INSECTS. 265
many Hemy^tera heteroptera there are seyen, as also in many Carof
hid ; the Cockchafer and other LamMicomia have six, the Stag*
Beetle {Lucanua oervus) twelve, &c« The length of these tabes is
different, but on the whole is more marked in proportion as the
number is fewer, as in the Butterflies ; thej contain the eggs in a
string ; the largest and most developed are at the lower end, the
smaller above. Here the ovarial tubes run out into a fine thread
which Leox Dufour terms Suspensory Ligament, whilst J«
HuELLEB considers the parts to be vessels which connect the ovaries
with the dorsal vessel. In most instances the threads unite on
each side to form a cord; in others (in Phasma ex. gr.) they
proceed separately to the dorsal vessel.
From the inferior termination of the ovaries proceed two oviducts
{tubiB)^ which coalesce to form a common tube beneath the rectum :
it is ordinarily much shorter than the tuba; in the cockchafer, on the
eontrary, it is longer than these. Different homy plates surround the
dilated inferior termination of this common tube; it has a sphincter
muflde to contract it, as well as several others^. Generally it falls,
with the rectum, into a common cloaca^ or it opens beneath and
IB fix>nt of the antu. Sometimes the external sexual organs of the
female, generally seated in the ninth ring of the abdomen, which is
included and hidden in the eighth, are prolonged into an appendage
externally. Here belongs the tubular vagina of Flies {vagina tubi-
JhrmiB)j in Chrysia, &c.', which is formed of the last abdominal
rings that can be drawn within each other like an opera-glass. In
others the vagina is Uoo-valved {vagina bivalvis)^ as in Locusts
{Locu8Ub)j and projects beyond the last segment of the abdomen as
an ensiform compressed prolongation. In others there is a per-
forator borer {terebra) or a sting {aculeus); here, besides the bi-
valved vagina, there is a sharp organ for puncturing, with serrated
edges, and composed of one or of two homy threads ; when at rest
the sting is concealed within the abdomen ; it is connected with a
poison-gland*.
who hM gi^en a highly magnified figure of these |Mrto, each avariium oonsuts in tbe
Honey-Bee of 150 tahes, BiU, natw, p. 471, Tab. zix. fig. 3.
^ See Stbaub, Anai. dm anim. ant, p. 399, and the figures of the Oockehafer,
tUd. PI. 5, figt. 4, 5 m, PI. 6, ^, 1 h, If.
' Aleo in MffOenu cwetdMa amongst the CoUcfUrOf Ann. det Sc, not. Tom. tt.
PI. 19. fig. 5.
' Comp. here especially Burmbistib, Hdndh. der Brtiom. 1. 1. 109— 9i5» Taf. 11.
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266 CLASS ym.
Diffisrent appendages belong to the common oviduct, or to the
ragina. Of these one is more constant than the rest, and opens into
the common ovidnct, close to the uppermost part, where this is
formed fifom the imion of the two tubes. Previous to copulation it
is empty, but after that act is filled with a white fluid — ^which is the
seed, as microscopic investigation has demonstrated beyond doubt,
from the presence of the hair-like spermatozoa in motion. This part,
g^[ierall7 single, may therefore be called receptaculum seminw.
Frequently it has an appendage {glandula appendictdaria). In
many insects there is another vesicle present which, during copula*-
tion, receives the penis {bursa copuhtbrix^poche oopuUxtrice Audouin),
and which in the cockchafer is a large bladder beneath the oviduct.
In the Butterflies this organ opens externally, and not into the ovi-
duct, so that there are two sexual orifices, whilst a canal leads from
the bursa copulatrix to the oviduct, and conducts the seed into the
receptaculum seminis situated above. There are other vesicles, or
glands, generally in pairs and situated more behind for the purpose
of covering the eggs with an adhesive fluid. In the Butterflies
these are seen as two pyriform vesicles laid transversely with their
broad bases opposed, which at the other end pass into a very long,
contorted, blind canal. In a few Insects still other secretory organs
have been observed, which probably secrete a peculiar odorous
matter to attract the male^
Ob the borer (terebra) in the Cicada, see DoTiBi in Ann, de$ Se, not, le S^rie» vn.
Zoologie, pp. 193 — 199, PI. 8, the middle brutle {U pdn^on Dotsbe) works liko
a wedge.
1 It 18 difSctilt to be brief on a subject which has inference to such an important
difference of organisation, and which, on account of the various Tiews of observers,
possesses an historical interest Malpiohi (de BcmJbyc^ long ago recognised the
veticula eopuUUrix as the oiigan which receives the penis and gave it the name of
ntervM; often the penis or a part of it is broken off, and remains here after copulation.
The penis is figured in this part in Spkinx Itguttri from a preparation by HninsB in
the Catalogfie of the Phytklogical Seriu of the Muteum of the Cottege rf Siu(rg0om»,
VoL V. London, 1S40, Pi. 67, fig. 8. It was with fluid from this vesicle that Huktxb
impregnated artificially the eggs of other butterflies. Phil, Trant, 179a, p. 175 O^^
BomJtyi/x mari^ ; an experiment already devised by MALnoHi, but attempted without
success. Spallamzahi, before Huhtxb had effected the artificial impregnation of the
eggs of the silkworm, but with the seed taken directly from the male butterfly, so that
his experiment does not belong to our present subject (Eic^»inence$ pour tervir d VHui,
de la GSnfyr, Qen^e, 1785, 8. p. 313). AuDOUiir has the honour of having been the
first in our century to direct attention to this subject, whilst previously all these
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iKSBcnu 267
The male indiTidnals amongst Insects have nsnallj two teHeSj
tikough there be occasionally only one, jnst as in the female thore
may be only one ovarium, Sfnch is the case with lAihchiua where
the part has the form of a long tortuons canal. In the Scolcpendra
proper {Seal, mcrsitans^ &c.) axe different oval testes, much extended
in lengthy (described improperly by Kutoboa as epididymides)
-which at each pointed extremity send off an efferent canal; all these
canals coalesce to form a single canal which is very tortuous and
widens into a sac below {testiculus Kutobga'). In Scutigera there
are two rery tortuous canals {tesHculif) present, which begin with
an oval sacciferm expansion, and then pass into a single fine tube,
of great length and winding right and left with close curves ; this
tube opens into a canal, which as an arc connects the two efferent
canals each of which dilates twice into an oval vesicle '• In Julua
there are two long blind tubes, which, connected by transverse
canals, have the form of a ladder, and to which laterally blind sacs
are appended ; these sacs may be considered to be testes and the two
longitudinal canals to be va/sa deferential.
In the hexapod Insects the parts which prepare the seed are
always in pairs* There is found indeed in most Lepidcptera and in
certain Cohqptera {ex. gr. in Ophonus and Harpalus, genera of the
family of the Carabici) a single testis^ but since two efferent canals
arise from its lower edge, it is obviously formed by the union of two
appendages had bean confounded together as secreting organs, Ann, det 8c. not, n. 1834,
p. 381. We owe to C. Th. Vov Sibbold the most complete investigation of this
mbject; see his Femere Beobaehtungen Uber die Spermatozoa der wirheUoBen Tkiere, in
MuslixbIb ArMo, 1837, s. 393—433. If* m Von Siibold assures ns, the verieula
copulatrix only seldom contains Spermatozoa, and then usually dead ones, it is less to
be wondered at that the experiments of MALPiaHi failed than that those of Huimni
succeeded; they ought to be repeated with better success with the fluid from the
reeqfUieulum aeminis, LioN DurovR still persists in considering all these i^pendages
zm giandei $&fi/lqttei.
^ S. Kutoboa, Scolopendrtz monUantu Anatome^ PetropoL', 1834, 4ta pp. 10, 11,
Tab. n. figs. 3—5 ; Rtioib Joras in Todd^b Oj^dop. n. p. 413* Ag' ^i*
* liios DuFOUB, who has giren a description and figure of tibese parts, considers
the first pair of these Tesicalar expansions as tette$; the tortuouB canals as veiienkz
mminale$f Ann, dez Se. nai, n. 18^4, p. 97* PL v. fig. 3.
* See figures in ToDD*a Ojfdopad, in. p. 551, (article Myriapoda, by BnocB
Joms) and by Smir in Mubllbb's Arekw, 1843, Ta£ xm. figs. 17, 18.
^ L^9 Duvoub, Ann, da 8c, not, n. p. 133* Xab. Tl. fig. 8 of ffarpalif rm/feomie
(copied in Wagvbb's Icon, PkynoL Tab. xxz. fig. 8).
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268 CLASS nil.
wliich were originally distinct, as at least in Butterflies is placed
beyond all doubt by the history of deirelopment. In many Hymm-^
optera the two testes lie aide by side in a common covering {scrotum
Dufour). Sometimes these organs, usually white, are distinguished
by lively colours (deep red in Paptlio Irassicce^ and in some Hemir
ptera^ yellow or orange-coloured in some GoUoptera) which depend
upon the investing membrane. Moreover the structure of the tesieB
is veiy manifold, and, as in glands generally, nature has here solved
the problem, in a small given space to increase as much as possible
the secretory surface, in very different ways. The simplest form is
that of a single blind canal, which is sometimes veiy tortuous^. In
other cases this blind canal has more the form of a sac, ex. gr. in
Scutellera, Edessa. Yet they are not always constructed in this
simple way, when they have externally the form of a single blind
sac and have also been so described by some writers ; in Libellula^
for instance, this sac contains a number of small round vesicles'. In
by far the greatest number of Insects each testis consists of a collec-
tion of different, sometimes very numerous, vesicles, or cylindrical
canals {capsules siminijiques L^ON Dufour) terminating blindly,
which are united in form of a fan, of a star, of an umbel, or in
bunches, and from which canals arise that afterwards tenninate in
a single efferent canal'. This efferent canal forms sometimes at its
commencement numerous tortuosities, to which the name of ept*
didymis has been given (as in many Carabidy in Meloloniha^y in
Nepa, &c.). The lowest part has often an expansion* to which the
name of vesicula seminalis has been fitly given. Far less propriety
is there in giving this name to different blind canals which are met
^ In jyytueut marginalia the entire canal when unwound appean to saipass the
length of the animal twenty times, Heoetsohwbilxb I>e Inaeetar. ffmiioL p. 19.
• L^H DUFOUB, Mim. pr€$enU8, vii. p. 57a.
> For a methodical review of all these forms an arrangement is requisite in which,
at the same time, there are not too many divisions. Comp. JoH. Muillxb, J)e glan"
dvlarum tecemaiUum ttrudura penitiori, 1850. fol p. 103 ; Bubicbistkb, Jfandb, der
MtU^m, I. s. « 1 7— 119 ; Waonxr'8 Lekrb, der vergi, Anat, 1834, s. 319—331, and the
flgures chiefly bonx>wed from htov Dufoub's nomoous investigations in Muxllsb,
1. L Tab. XVI. figs. 1—19, and in Waobbb, /eonei Pkgnol. 1839^ ^<^^ 3^^* %*• i— «<$•
^ Stbaub, 1. L Tab. vi. fig. 1, c, c,
' For instance, in HydnpkHu$y in ApU mdUfioa, in Genu and Vdia (JL6om
DUFOUB, JK0C&. «. I, ffimipt. Tab. XX. figsi 138, 139), in Coitui mmrginatu» (Liov
PUFOUR, ih. Tab. X. fig. 1 27).
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INSECTS. 269
^ith in most Insects and which unite with the efferent canal. That
these parts rather serve to effect special secretions and correspond
with the fTostate and Cowper's glands in higher creatures is pro-
bable even from their composite structure and great development.
Hence in some Insects they have been even described as tesHculi
and the true testes as glands of an unknown use, as for instance, by
SwAHMEKDAM in Oryctes nasicomta^. In this and other LamMi-
comxa {MdolorUh^ij Cetonia) these glands are two tortuous canals,
which resemble the testicuU of the Carahici. In Hydrophihis pioeus
these parts exceed the testes in circumference. Ordinarily there is
one pair': often also there are two pairs of these accessory organs
present; in some are found three or even more pairs. They are
entirely absent only in few Insects {Oersisy Velia, Banaira). In
lAbelhila they are also wanting'; yet I think it is nearer the
trutii to say that here they are not connected with the testicuU
and are present in an unusual situation, which is in agreement with
the copulation of these animals^. From this also it is apparent that
these organs are not vesiados seminales.
For determining the purpose of the different parts connected
with the organs of propagation microscopic investigation iu the
recent state ia of great service. C. Th. Von Siebold has never
met with spermcUozoa in the parts which we consider subservient to
special secretions, which however they ought to contain if they
were really seminal vesicles as Leon Dufoub and other writers
suppose. The spermatozoa of Insects are like hairs, and are often
found in the testes united in bundles and surrounded by a transpa-
rent covering*.
We must notice, in addition, that in many species of Insects a
great similarity of form has been observed between the organs of
propagation in the two sexes. We see this resemblance in certain
^ B^bd der natmur. Tab. zxx. fig. 8, m m; these are the true tesU$. I oonld ahaost
sappose that LifoN Dufour bo iDdelatigably laborioiis in the anatomy of Inaects and
80 rich in experience, has made the same mistake in Pdoffonut; see the J2m4. 9ur tea
ffimipt. PI. XI. fig. 137 A., where I consider the spiral organs i 6 to be tetiet,
s As in the ffymenoptera, for instance.
» LioK Dufoub, Mhn, priteniis, vii. p. 572.
* See below, in the systematic arrangement, in the fiunily of the JAbeUvUna.
' See Von Socbold, Ueb, dU Spermaiozoen der Onuiaeem, iMeeten, QatUropodm
«. einiffer anderm mrbelloten Tkiere, Muxlleb'8 Archiv, 1836, s. 10—43, '^^^^ ^^*
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270 CLASS YUI.
beedes, but especiallj in seTend Hemtptera^ as well in the form of
the accessory glands, as in that of the testes and ovariuy in the nam**
ber of the oviducts in the last and of the spermatic ducts in the
former, &c. We cannot however admit that this similarity has
the value of a general rule; the Lepidoptera, for instance, not to
speak of other losects, exhibit an entirely di£ferent type in the two
sexes.
The external sexual organs lie, as in the females, at the hind-
most part of the abdomen^. The penis has a very different form
and substance. Ordinarily it is surrounded by two homy plates,
and encbsed in a membraneous sac in the retracted condition; in the
Coleoptera the penis is covered by a homy case, and supported by
two homy threads*.
Amongst the malformations of Insects hermaphroditic individuals
occasionally occur, in which one half of the body is male, the oth^
female, like the Androgynes in A£rica, of whom the ancients &bled,
and who had a female breast on the left side, and a male on the
right'. This lateral bisexualily is most frequently seen in Butter-
flies, in which it strikes the eye more readily from the form of the
antennsB or the colour of the wings ^; yet some instances of it are
known in other orders also^.
Before we turn from the consideration of the sexual organs of
Insects we must shortly notice another peculiarity observed in bees
and other Hymenoptera living \xl societies. Amongst these many
^ The Chilognaiha (Jvlua) are an exception to this ; the partB, in both aexee, are
here situated veiy far forward, at a short distance from the head. They are also
double (two vulva, two penes), as in the enuiacea.
' See the figures of Sthaus (op. dt.) in the Cockchafer, Fl. n. figs, ii, ^t, PI. vi.
fig. I. Waonxr compares these horny threads with the astictUum perns, found in
many mammals. On the sexual organs of insects, in addition to the works cited above,
two monographs (both, however, of somewhat old date) may be consulted, viz. J. J.
Hbgitsgewbiler, Diss, de Insedorum genitaltbus; cum Tab. Turici, 1830, 4to. and
Oedechtsorgane der Inseden von Dr Svokow in HEVsnrosR'B Zeitsehr. f. organ, Pkystk,
n. Eisenach, 1818, a. 331—764, and further, F. Srsur, Die weiUu^ (kseUeektsorgam
der K&ftT, Hit ix. Kupfertaf, Berlin, 1847, 4to.
" C. Pltnu, HiA, not. Lib. vii. cap. a.
* For instance, in Bombyx dispar by Sorarffer, in BomJb, eraic^, by EsPRR
{Seobachtungen an einer neu entdeekten ZuntterpkaUme, Erlangen, 1778, 4to.) in Vemessa .
wiices, by Rapp (Orrk's Isis, 1833, s. 235), Ac.
' As in SeoUa maenlata, by Rovand, Ann. des 8e, entomol. TV, 1835, P* I9<> ^
Zucanui cervus, figunxl in ASMUSS, Mot$siruosiiaUs ColeopUrer, Bigm, 1855, T^. z.
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IKSKCIB. 271
indlTidxials occur, which being incapable of propagating have been
commonly considered to be sexless (the so-called neuters, working-
bees, &c) Even the external form indicates that they resemble
rather the female than the male individnals, as the same is also
indicated by their mode of Ufe and their instinct directed to the care
of the yonng ones. Anatomical investigation has confirmed this
conclusion by demonstrating in^the working^bees imperfect oyaries^
These individuab thus remain imperfect females, nurses, foster-
mothers*
Observations are not wanting with respect to the development
of Insects in the egg, although hitherto this subject has not been
sufficiently investigated to allow a general representation of it to be
offered. Li eggs that are just laid, nay in those which lie at the
lowest part of the oviduct and are the most mature, the germinal
vesicle has disappeared ; in eggs situated higher up in the oviduct
it may be seen clearly with the germinal spot^. On the yelk is
formed, from a union of cells, a layer as germinal membrane {bl<ut<h
derma) which continues to grow so as to surround the entire yelk.
The first rudiment of the embryo, the nota primitiva, lies on the
ventral surface ; the yelk lies on the dorsal surface, and becomes
enclosed by the constantly growiug ventral plates, without the for-
mation of a special umbilical- or yelk-sac by constriction. The
9tigm<Ua are developed only at a late period, and become open only
shortly before the escape from the egg'.
^ Madsm. Jubikb in Hubeb Nouv, ob»ervati<m$ tur le$AleSle9, ^e ^t. Pftrii et
Gen^e, 8vo. n. Tab. xi. fig. i, figure copied by Ratzxburo in his enquiries on this
nibject in JVov. Ad, Acad. Cm. Leop. Car. Vol. zvi. PI. n. Tab. 47.
* Bee the microeoopio r^raeentation of an egg-tube from Agri9f^ by B. WAaNXB,
Ahkand, der mathem. physic. Klaste der Ahademie in MUnchm, Bd. u. fig. i ; see
8. 558.
* On the development of insects in the egg there are some observations of SuoKOW,
AnaUmuekrpkydoloffUehe Untenueikimgen dor Inseoten mnd Krtitimtkitre, Heidelbeig,
1818, 4to. mit Kupf. s. 19, 33, 35 (eggs of BombjfX pini) ; also short, but interesting
communications by Rathks, on Blatta ffermaniea in MxoKSL*s Arehiv. 1844, s. 37 —
37, Taf. n. (here there is on each side of the abdomen in the embryo, behind the attach-
ment of the third pair of feet, a pediculated disciform organ which is, perhaps, a tem-
porary respiratoiy organ to be compared with the giUs of ]arv« of salamanders ; there
are only four Malpigfaian vessels, which are increased in number after birth), and
finally, by Kobllikkb (on CJdnmomut, SimuUa, Danacia) Obtervaiionet de prima
InmOarum gmeH ZHu. ina/ug, Adjeota muU in. Tabid. Turid, 1844, 4to. The numerous
plates of HiBOLD in his J>i9^iiitioit€$ dt AnimaUum verUMi curenUvm in 090
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272 GLASS VIII.
Most Insects quit the egg in a form entirely different from that
which thej afterwards possess. An impregnated female butterfly,
for instance, deposits eggs, from which caterpillars proceed, which
present not the slightest external resemblance with the mother.
Thej are vermiform creeping animals, provided with different pairs
of feet, which eat enormously, grow rapidly, cast their skin often,
and at the last casting change into quite another creature, with a
very hard and homy skin, which has no Hmbs, does not move from
its placed takes no food, and falls as into a death-sleep. There
may however be generally discerned in the seemingly formless
mass, on close observation, the external parts of the butterfly,
which folded and rolled together are concealed beneath the homy
shell, on whose surface they are traced out. After a longer or
shorter time, sometimes only after many months, the perfect insect,
the butterfly, quits its narrow cell. At first the wings are short,
moist and imfit for flying, but soon unfold themselres, become dry,
and then support the flapping Insect through the air, which soon
ftilfils its new destiny, the propagation of its kind, and dies'.
In these changes of form {metamorphoses) of Insects the first
form or first state is called that of the vimsk or larva^ and the
Insects are then named caterpillars, maggots, &c. The second state
is that of nymph or pupa (in day-butterflies called also chrysalis).
The third state is that of the perfect insect {insectum dedarojtum^
imago).
All Insects do not pass through this threefold state. The wing-
less hexapod Insects, with few exceptions, leave the egg in the
same form which they afterwards retain ; only the rings and the
feet become more numerous in the Myriapoda. These Insects
Latreille names Insects without metamorphosis. No winged
Insect, on the other hand, comes from the egg with wings ; but
evoiutione {De ffeneraiione Jnmdorum in cvo), Fnmcof. ad Moen. folio, FasdcuJi ii. (not
completed), relate principally to Mu$ca tomiioria and some LepieUfptara, bat do not
give 80 much information as might have been expected (rom the diligent and patient
inTestigations of the author.
^ If the pupa be however in such a situation that the perfect insect would not be
able to come out of it (in the branch of a tree, for instance), then it changes its place
towards the period of the last change, by pushing on its body by contraction, a motion
assisted in many cases by little hooks on the rings of the abdomen.
* Sometimes the perfect insect^ shortly after its coming forth, once more changes
its ooat» as is commonly known of the Epkenura,
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INSECTS. 273
amongst the winged there are many, which undergo no other meta^
morphosis than that they obtain wings. Their larvm resemble the
perfect Insect, but are quite without wings : the pup<B hare rudi*-
ments of wings and move themselves : in the last moulting these
wings become developed and perfect. These Insects undergo
accordingly an imperfact metamcrphosis {demumetamorpho^e La-
TBEILLE, metamorphosia incomplete^ ; this is the case, for instance,
with the grasshoppers. Most winged Insects, lastly, are subject
to a perfoct metamorphoais {metamorphosis completa), as we have
described it in butterflies ; the pupa takes no food, and remains in
a state of rest or slumber. The pupse of flies are entirely motion-
less, surrounded by a hard shell, and shew no limbs of the perfect
Insect concealed beneath it; this shell is formed by the dried
integument of the larva. Such a pupa is named pupa ooarctcUa.
In other dipterous Insects and in the Lepidcptera there is a hard
elastic membrane, surrounding the enclosed compressed external
parts of the Aiture perfect Insect, and so disposed that they can be
distinguished through the covering. Such a pupa is named pupa
obtecta; such pupsB move the rings of the abdomen. In still other
instances the wings and feet are free, without being surrounded by
a common covering, as in the pupsB of beetles and bees^.
These changes are not confined to the external parts; in the
internal structure also very remarkable changes occur. The intes-
tinal canal is in most larvsd straight, and consists principally of a
wide stomach. The oesophagus and the part of the intestinal canal
behind the stomach are longer and narrower in the pupa and in
the perfect Insect, since the stomach contracts and is more definitely
separated firom the rest of the intestine. The nervous system becomes
^ For pupce of the last kind the word nympha is someiiines specially used ; see
SWAMifEBDAM, jBtU, ncU, pp. lo, i6; Bladh, FundamentaZoolofficBf in Link. AmosnitcU.
Acad. Tom. vn. p. 151 ; Nswpobt in Todd's Oydop, 11. 879.
ItisjSMJJB names a pupa complete (pupa compleUi), which moves itself, and in all
respects resembles the perfect insect ; half-ccmpUte (temi-completa), that which is at rest
and takes no nutriment. Sytt, nal. Ed. ii, i. p. 534. Fabbioius transferred tiiese
names improperly fin>m the pupa to the metamorphoses, and thus named complete
metamoiphoais (metaiMrpKona completa), that which, in fiust, is no metamorphosis, as
in the myiiapods, the spiders, &c. The metamorphosts which Latbbillb names
oomplete {ex. gr, that of butterflies, beetles) Fabbioiub names incomplete (tneomp^eto) ;
the semi-metamorphosis bean with him the name of m/damorphotie iemi-completa. See
Fabbioiub, PkUoe, Entom, p. 56.
VOL. I. 18
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274 CLASS VIII.
tortaoQS in the pupa, and shorter in the perfect Insect ; the nervons
ganglia become less numerous, from some of them first approxi*
mating to each other and afi;erwards coalescing, whilst others
entirely disappear ; the first ganglion especially, which is situated
in the head, increases in amplitude. The sexual organs, of which
the germs already existed in the larvsB, become developed as well
in respect of complex structure as of size. In the pupa new organs
also come to view, of which before there was no trace, such as the
wings, which are seen folded and rolled together internally at the
thorax. The dotsal vessel undergoes less change than most of the
other organs.
There is in larvss, moreover, a peculiar fatty mass present, of
which we have already spoken above; the secretion of this taX
constantly increases, the nearer the larva approaches the condition
of pupa ; in this condition the &t is consumed again, and in that
of the perfect Insect, when it also takes food, fat is no longer
secreted.
The fatty secretion is thus obviously necessary to supply nutri-
ment to the pupa, and to afford the material for the development
of the organs of the perfect Insect. Yet the pupae of Insects
which undergo a complete metamorphosis, take, as stated above,
no food, and are in connexion with the external world through
respiration alone*. The condition of pupa therefore may be com-
pared with that of hybemating animals, which are very fat in the
autumn, during their sleep take nothing, and in the spring come
out of their retreats in a very emaciated condition. The larvae of
Insects eat more than is necessary for their own growth ; they are
therefore usually inactive; deficient motion and superfluous food
favour, as is known, the secretion of fat^. The reason why in
perfect Insects no more fat is secreted, is found as well in the
development of the sexual functions as in the greater activity of life
and rapidity of motion, which are peculiar to them.
^ On this aooount they lose in weight. Hub Iobb is, however, at first very smally
and only becomes remarkable on the approach of the last change. See Nswfobt in
Todd's Oydop, n. pp. 879, 880.
* That this fiit is not necessary for the life of the larva, is shewn by the examples
of caterpillars in which ichneumons have deposited their eggs ; the larvn from these
consume the fat of the former, which die from the robbery of their stock of food only
at the time they should change into pupn, or have changed.
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INSECTS. 275
We see therefore in the perfect Insect the manly period of their
life : in the larva the childish period. Between the two nature has
interposed a deep sleep of development. The marriageable period
is deadly for many. There are also many difficulties to be over-
come^. Some organs must for a time stand still, others (as for
instance, the silk-secreting tubes of caterpillars) must entirely
disappear. The development of the sexual organs is essential,
and for that everything must wait awhile ; these remain during the
larval state behind other organs ; now they repress in turn by their
development the activity of other organs. Finally, the perfect
Insect comes forth, in many respects a new creature. This is the
true object of the phsenomena, of which the metamorphosis is
composed, which is not so entirely unique in its kind, as might be
at first supposed. The perfect Insect lives for propagation, and
when it has attained that purpose of its being, it dies to make
room for others, and serves for food to birds and other animals.
Thus also an annual plant ceases to grow as soon as its bloom is
developed, and dies when the seed is come to maturity*.
^ Eyeiy casting of the skixi is oonnected with more or leas of danger; the moulting
is also a distressing season for birds ; bat especially the last shedding, "when caterpillars
are changed into pupae, is frequently fatal. Sometimes the casting is incomplete ; the
head of the caterpillar remains attached to the pupa. In this way may be explained
the occasional occurrence of butterflies with caterpillars' heads. See 0. F. Muellbb,
Ikacripikn (fun papShn dUtede ChenUle, MSm. prUemUi d VAcad. dea Sc,de Paris,
1774, VI. pp. 508, Ac, Natwrfonckery xvi. 1787, s. 203— « 13, Tab. iv. f. i, a ; Wes-
UAXL, Ann, det Sc. nai, sec. S^e. Tom. vin. 1837, Zoologie, pp. 191, 192; Bbuinsma,
huiUngewone afmjlnngen, wjuurfftnomen m de gedaantet^erwitseling det zijdeworma,
TydKkr., voor natuwrl. Oesch. en Phynd. vn. 1840, pp. 257 — 170, PI. iv. and my
Aanteekeninffen thereon, Hid. pp. 271 — 275. Somewhat different are other observations
of Majoli in Bomhtfx mori, in which the moths, without having first become pupae,
appear to have proceeded immediately from the caterpillars. 'M.eckki/b Archiv fOr die
Ph^cL n. 1816, s. 542.
* What is said here relates especially to the complete metamorphosis ; in the
incomplete the changes are less important. Ck>mp. on this subject RBNoasB's PhytuA.
UfOen. s. 49—87, and Hxboud's Bahnekdungaffeachiehie der Schmetterlinge, Casel
u. Marburg, 18 15, 4to, (one of the most excellent works on Natural History which
have been pubHshed in this century), in the numerous plates of which the development
may be followed without a break in the whole and in aU its steps. Comp. further, on
the changes which the intestinal canal undergoes on metamoiphosis, Dutboobxt,
J<mnuU de PkyHque, Tom. Lzxxvi. 18 18, p. 130, &c., and in Msokbl, Archiv f. d,
Pkyn/cl. IV. Bd. 1818, s. 285 — 293 ; and on the changes in the nervous system, New-
POBT, PhUoi, Trans. 1832, n. pp. 383—398, PL xn. xin.
18—2
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276 CLASS VIII.
The reproductive power in Insects which undergo metamorphosis
is wanting in their perfect state ; but if at an earlier period in
the state of larva they have lost a foot, it grows out again at the
next moulting, and is more or less perfectly restored. Also in the
Myriapoda excised antennas grow again ^
The nervous system of Insects has for central part a row of
ganglia of different number, which are usually connected with one
another by two threads that are often very intimately united. This
row of ganglia is situated on the ventral surface beneath the intes-
tinal canal in the mid region of the body ; the first ganglion however
lies in front of and above the oesophagus, and there arises, by reason
of the two threads which connect it with the second ganglion, a
ring which surrounds the oesophagus. The greatest number of
ganglia is found in the Myriapoda, eighteen in Lithobius {Scolopendra
foTJuxiJUi), twenty-three in Scolopendra morsttans. In the larvce of
butterflies thirteen are counted, but ordinarily they are less numerous
in the hexapod Insects. Large ganglia are situated in the thorax,
and in some there are none in the abdomen, but two nervous strings
alone, sometimes close together, sometimes separate from each other,
as in Nepa and Cicada, From the ganglion above the oesophagus
{ganglion cerebrale) arise the nerves of the eyes and antennse ; this
ganglion lies transversely on the oesophagus, formed of two oval or
somewhat conical lateral portions with their broad part turned to
each other; the inferior surface is somewhat concave, the upper
convex. The second ganglion, the first of those beneath the intestinal
canal, is by some writers compared to the cerebellum, by others, on
better grounds, with the medulla oblongata; the nerves that arise
from it proceed to the oral parts, and perhaps correspond to the
different branches of the fifth pair in vertebrate animals. Earlier
writers, as Ackermann, Reil and Bichat, thought that the abdo-
minal cord of Insects might be compared with the vermis sympa-
ihicua of vertebrate animals ; Cuvier and Gall, on the contrary,
have disowned and rejected this correspondence. It is necessary in
this inquiry to determine in tlie first place what character is to be
considered of sufficient value to distinguish the spinal cord from the
^ Newpobt nuhde experiments on Ivlua, lAUidbim and caterpillars of butterflies.
See Phil, TraiM, 1844, p. 183. In Phaama sometimes one foot is less than the rest,
being a new growth. I found this onoe aJso in Reduviv* penonaiu*.
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INSECTS. 277
system of the great sympathetic nerve. It is impossible to recognise
the position on the dorsal surface as such a character; for the reversed
position of the heart in the invertebrate animals might lead ns rather
to expect that that of the central nervoos system would be reversed
also. The pecnliarity of the sympathetic system, amongst other things,
consists in this, that it supplies nerves which are distributed to parts
not subject to the will. But since from the ganglionic cord in Insects
the nerves of the organs of sense and the nerves of the volimtary
muscles arise, there is no reason for comparing it to the sympathetic
nerve. The ganglia therefore of this cord are to be considered as
an union of the ganglia of the spinal nerves of the two sides*. This
view would seem to receive more support when we reflect, that the
spinal ganglia belong to the uppermost (the posterior) or the sensi-
tive roots of the spinal nerves, and that in Insects two strings have
been discovered in each connecting band between the ganglia, of
which the undermost alone is connected with the ganglia, whilst
the uppermost merely passes with its fibres over the ganglion*.
That here the uppermost and not the undermost string, as in the
spinal cord of vertebrate animals, is related to motion, is to be
explained by the reversed position of the nervous system. The
similarity becomes obvious when we thus consider the matter, that
in Insects as well as in vertebrate animals the motor strings are
placed towards the interior, the sensorial nearest to the surface. It
is, however, perhaps more prudent not to pursue this analogy too
far. With these uppermost strings we must not confound the
system of transverse nerves which Lyonet described long ago in the
caterpillar of the Willow-hawk under the name of brides Sptnth'es\
These are situated a little in front of each ganglion, pass transversely
over the straight muscles which lie lengthwise on the ventral
surface, and are distributed by their branches to the muscles and
especially to the air-tubes and the dorsal vessel. A longitudinal,
1 G. B. Tbxvuianub, Bidogie, v. s. 33 1, 333 ; E. H. Webkb, Anat, comparata
nervi tympathici, Lips. 18 17, p. 95.
• This 'important discovery of Newport, who was incited to it by C. Bell, the
celebrated discoverer of the distinction of the motor and sentient roots of the spinal
nerves, may be seen, illustrated by figures, in PhU, Trans, 1834, Pt. 2, pp. 406 — 410.
» TraiiS ancA, de la Ohm. pp. 98, 101, PI. IX. figs, i, 2, Newport has very
accurately investigated this nerve in Sphinx Liyustri, PhU. Trans. 1836, Pt. n.
pp. 544, 545, PI. xxxvn. (This figure is transferred to Todd's Cyclop. 11. p. 987.)
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278 CLASS VIII.
single nerve, lying above upon the strings of the ganglionic nervous
system, connects each plexus with the following one. In perfect
Insects this system is less distinctly visible, and often is completely
combined with the rest of the nervous system ^
There is found in addition still another nervous system in Insects,
destined especially for the organic life, which was made known in
part by the investigations of Swammerdam and Ltonet in former
times, and described by the last under the name of li/erf recurrerU^j
and to which in our century JoH. Mueller has by his investiga-
tions especially directed the attention of anatomists. It has been
compared by him and by most modem writers to the sympathetic
nerve of vertebrate animals, by others to the nervtta vagus. This
system of nerves consists of a single middle portion and of two lateral
portions. The single portion arises from one or more nervous
ganglia situated in the head, which are connected with the most
anterior part of the first (the cerebral) ganglion of the ganglionic
cord. From this single portion whilst situated in the head nerves
arise for the uppermost oral organs, and a thread which runs along
the oesophagus on the dorsal surface to the stomach, and at its
extremity terminates in a ganglion. In Pkasma firula Brandt
saw numerous branches arising transversely, and running in arches
over the oesophagus and stomach to form a fine nervous net. Per-
haps a similar distribution may be suspected in other Insects, where
the extreme delicacy of the nervous branches does not permit their
determination. In most Insects the middle single portion is the
most developed ; in Orylhtalpa and Oryllus, on the contrary, the
lateral portions are more developed than the single and middlemost.
The lateral portions consist ordinarily of two pair of ganglia that
lie close together behind the cerebral ganglion, of which the
anterior is connected with the cerebral ganglion by one or two fine
nervous threads. From these ganglia delicate nervous branches
arise which run to the oesophagus whilst they are also in connexion,
by fine threads with the single middle nerve that runs over the
oesophagus'.
^ Besides the authors cited comp. also especially an excellent paper on the nervous
system of Beetles by E. Blanohard, Ann, des Sc, NcUvr,, si^me S^rie, Tom. v.
Zooloffie, 1846, pp. 173— 379» PL 8— 15.
■ TradtS anat, de la Cken, pp. 413, 578, &c.
> Comp. JoH. Mueller, Uther tin eigenthMndUket dem Nervm vympatkicuM analog
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INSECTS. 279
Of the organs of sense of Insects the eyes are best known*
We have already spoken above (p. 249) of the distinction between
simple and compound eyes. The simple eyes have a crystalline
lens and a vitreous hnmonr. The cornea, on which the crystalline
lens lies, without being separated from it by an aqueous humour, is
formed by the common homy integument of the body, which at
that part is raised convexly and is more transparent. The vitreous
humour is surrounded by a black pigment of the choroid. The
compound eyes, always two in number, present a cornea which is
divided into many fa9ettes, ordinarily hexangular. Each of these
divisions has the form of a small, usually biconvex lens. Behind
them lie an equal number of transparent pyramids or conical
bodies which are turned by their base to the cornea and by their
apices approach each other inwards*. Lastly, there is a nerve at
the apex of every cone; the optic nerve in fact divides into as
large a nimiber of branches as there are divisions of the cornea.
A dark-coloured pigment, often violet or blackish brown, separates
the nervous fibres and the transparent cones, especially at their
pointed extremities, from each other. At the base of the cones,
beneath the cornea, there is frequently a pigment of a different and
more lively colour ; hence arises the metallic splendour of the eyes
in some Insects, as in Hemerobiua and Chrysopsj which however
disappears after death. No eyelids are present in Insects, but
between the fa9ettes of the cornea there are found in certain
Hymenoptera and Leptdoptera, here and there, some hairs, which
ward off substances from the eyes and defend them. Surrounding
Nenentydem der Eingeweide hei den Ituecten, Nov. Act. Acad. Oca. Zeop. Car, Tom.
XIV. P. T, 1828, pp. 71—108, Tab. vn. ix., and J. F. Bbandt, BeiMrkungen ueber die
Mvndmagen- oder Eingeweidnerven dcr Everiebraien, M6m. de VAcad, des Sc. de
St. Patent, (vi. S^rie, Tom. in. 2, Sciencea not.) published separately, Leipaag, 1835,
4to, with m. plates ; also in French in the Ann. des Sc. not. «e S^e, Tom. v. 183$,
Zod. pp. 81, kc. and 138.
^ Will considers these cones, which Muelleb compares to the yitreous hnmour,
for the most part as crystalline lenses, and supposes that behind them there is still a
▼itreoos body with concave anterior surface to be found. In Sphinx Atropot, where
these cones are very large, (I found them one-seventeenth Par. lin. long,) I have
several times observed the separation pointed out by Will at the posterior extremity
of the cone. In other insects the cones are so short, that the separation, even if it be
presenty cannot weU be perceived, whilst even on that account Tbevibakub thought
there was reason to suppose that in some insects the cones in question were absent in
their compound eyes. EneKeinungen u. Qttdze des orffan.Leb€n8 n. i, Bremen, 183a, s. 77.
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280 CLASS VIII.
the eyes are large air-sacs or wide air-tubes, from which fine
branches arise, which in part run to the pigment and connect its
granules, in part pass into blind cylindrical tubes situated between
the nerve-threads of the vitreous cones*.
De la Hire, who first discovered the simple eyes of Insects,
thought he might conclude from their presence, that the larger
(compound) eyes were not organs of vision. That they also serve
for vision the experiments of Swammerdam, who smeared them in
flies with black varnish, have proved. Reaumur also did the
same with bees. It is more difficult to determine exactly in what
respects the office of the compound and simple eyes differs, although
the last probably serve principally for seeing near objects. The
bees, in which Reaumur had smeared these eyes with a dark
varnish, whilst their compound eyes remained imcovered, could not
find their hives*; moreover all flying Insects are invariably pro-
vided with compound eyes. There are Insects which have simple
eyes alone, as the Myriapoda and Parasitica (also the larv® of the
Lepidoptera) ; few Insects are entirely without eyes, like a parasitic
Insect of bees {Br aula NiTZSCH*), and a new genus of the Garahici^
Anophthalmus of Schmidt^, and different Myricgpoda. In the
diurnal butterflies and most Coleoptera, there are two compound
eyes alone, without simple eyes ; simple eyes are also wanting in
certain Diptera^ in Forficula^ BlaUa and other Orthoptera, in many
Hemiptera ; where they occur in company with compound eyes,
usually three are present, sometimes, as in Oastnia, Sesia, Noctua^
Chryliotalpa^ two*.
* See on the compound eyes of insects amongst others Hooks, MicrograpKia,
Londini, 1667, Tab. 34, Swamxebdam, B%bl, not, pp. 487 — 498, Tab. XX., J. Mublleb,
Zur vergl. Phynol. dea OeaichUinnet, Leipzig, 1826, 8yo. s. 307 — 390 ; by the same,
Fortffe8etzt€ anatomtsche Vhtermchunf/en tuAer den Bau der Augm hei den Inteden u.
Cnutciceenf in Meckel's Archive 1819, s. 38 — 64, and Udyer die Augen de» Maihafen,
ibid. s. 177 — 181 ; F. Will, Beitrdge zv/r Anal, der zusammengetetzten Augen, Leipzig,
1840, if to; A. Bbaxtb on the air-tubes in the compound eyes of the AritcuiatOf Tyd-
schrif, voor not, Oetch. en Phydol. xn. 1845.
■ Mem. p. aervir d rffisi. des Ina, v. pp. aSj — 289.
* Gebmab, Magazin der Eniomol. in. 18 18, s. 314.
* See Jag. Stubm, DeuiacMand^a Inaecten xv. 1824, pp. 119 — 137, Taf. 303. Also
a genus of the Xylophagi, Anommatua terricoUif Robbbt, Aaid. toy. de BrtixeUea, 1836.
^ KLua, Ud>er daa VerhaUen der einf adie SUm vmd Sckeildaugen hei den Inteden
mU zuaammengea Augen. PhyaiJxd-Abhandlungen der KdnigL Ahad. der Wiuena^ zu
Berlin, aua den Jakre 1831, s. 301 — 312.
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iNSEcrrs. 281
' For touch, in some Insects the sucker serves as the organ ; in
others the palps which belong to the organs of the mouth, in many
the antennffi. Of taste, smell and hearing, little is known. Taste
has its seat in the internal surface of the mouth. In some Insects
there is a part present, which may be compared to a tongue^.
Respecting smell different opinions are offered. On theoretical
grounds, from presumed analogy with vertebrate animals in which
the first pair of cerebral nerves always goes to the olfactory organ,
Blainville has concluded, that the antennae, to which the* first
nerves from the cerebral ganglion proceed, must be the organs of
smell'. Baster, Beimarus, Dumi^ril and Straus place the
sense of smell in the air-slits {st^makijy which admit the external
air to the air-tubes. Treviranus however has with reason alleged
against this opinion, that the stigmcUaj inasmuch as they are
dispersed over the body, must be useless for determining the place
from which the odorous matter proceeds ; also that in Insects, which
have no stigmcUa and which respire by tracheal gills, it would be diffi-
cult thus to account for the sense of smell. Bosenthal discovered
in flesh-flies {Mu»ca camaria) a red-brown, folded membrane, which
is situated in the head beneath the setting on of the antennae'.
Treviranus thinks that in sucking Insects, which are especially
distinguished by their acute sense of smell, the seat of this sense
ought to be sought for in the sucking organ itself, or in the
oBSophagus. K these animals suck in air, then they may smell
with the same organ, by means of which, when they imbibe fluids,
For the theory of visioii with compotmd eyes it is necessary that the partial images
be erect; hence JOH. Muellkb (Zur vergl. Physiol, de$ Oetichiamnes) has concluded
that insects see with their compound eyes not by refraction of the rays, but by keeping
separate the rays of light that come from different points. Hence he denies to the
fEM^ttes of the cornea, which are true lenses, a refractive power ; yet that vision in
insects with compound eyes occurs by dioptric means, has been shewn by Dr. A.
Brants, and established by means of an instrument constructed on the plan of the
insect's eya TtQdtchr. vow naluurl. Qeschied, en PhytioL, XD., 1840, pp. 12 — $6,
PL I.
^ See this part figured and described in some hymenopterous insects by G. R.
TBBvniAinjs, Verm, Schriften u. s. 135, 131—133, Tab. xin. fig. i, L, fig. 4, 7 ; Tab.
!▼. fig.5, 7, 8> 9» I*' »"»d L.
* See Duois, PAyno^. compar, I. 1833, PP* '57 — ^^h ^^o endeavoured to establish
the same views by experiments, m also Lbfebvbe, Awn, de la Soc, entomol, de Ftnnce,
1838.
» Reil'b Archivf, die Phyeiol. x. s. 427 — 439.
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282 CLASS VIII.
they taste. An observation of Huber respecting bees pleads for
this opinion. Bees are very averse to oil of turpentine; these insects
were not, however, repelled when Hubeb brought a pencil dipped
in it to the air-slits and other parts of the body, but flew away as
soon as he approached it to the mouth. Lesser had already
noticed that flesh-flies, whose eyes had been smeared with oil of
turpentine, still flew to tainted meat, but not so when their snout
had been smeared with it^. In Insects that do not suck the seat
of smell is probably at the beginning of the oesophagus also*.
Just as uncertam are we respecting the auditory organ of
Insects, although it was known to the ancients that they have
hearing^ Of this sense also several writers, and lately Newport*,
have sought the seat in the antennae. Yet the experiment that
grasshoppers, when their antennae have been cut off, continue to
hear, is even less favourable to this opinion, than is the presence of
hearing in spiders that have, as is known, no antennae^. Bamdohr
thinks that bees hear by means of their mandibles ; Treviranus
thinks that he has discovered in BlaJtta orientalis*, in L^eUula and
in bees, and Blainville in Cicadas, a special auditory organ'.
When we remember that for an auditory organ in its simplest
form nothing more is required than a nerve specifically receptive of
undulations of sound and so expanded that such undulations may
be conducted to it by means of the hard covering of the body, it is
^ The secretion of Stapdiof, which resembles putrid flesh in smell, deceires flesh-flies
into laying their eggs on the flowers (see Boebbl, Int, n. Musoar, et CHdic, Tab. ix.) ;
a clear proof that the instinct of these animals is influenced more by smell than sight.
■ G. R.TEKVIRANU8, Verm, Schr.u, s. 146 — 155, Biologie, vi. s. 307 — 318, Eradiein-
ungen u. Qetetoe d, organ, Lth, n. s. 141.
* See for instance, on bees, Mliaxii de Anitrndium not., L. v. c. 13. Of the
modems Bbunelli amongst others has proved hearing in Crickets by interesting
experiments ; Comm. Acad. Bononieiu, vn. 1791, pp. 199, aoo.
* Todd's Ch/doped. n. pp. 891, 961.
' M. G. 0. Lehxavn, JDe Antennia Intedorum DitaeiiaUo poaterior, Londini et
Hamburgi, 1800, 8vo. pp. 45 — 47.
^ In BlaUa oriemtalU there is on each side of the head, behind the base of the
antennae, a white spot, formed by a round membrane, under which portions of the
first nervous ganglion are immediately situated, TBXViBAinJs, Annd, der FFctteravucAm
OtadUckafi, I. s. 169--171, Tnf. V. figs. 1—3. Bubmkistkb thinks these white spots
are rudiments of simple eyes.
' Comp. Tbsvibanus^ BMog, Ti. s. 358—360; BuoKYniLBy De torgamaatwm dea
Anim, 1B22, 1, p. 565, ke.
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INSECTS. w 283
readily seen, that it is not veij possible in all cases to determine
'by anatomical investigation the situation of this organ. Yon
SiEBOLD not long ago thought he had discovered in Orthcptera an
auditory organ which is not in the head. In Locusta there are on
the tibia of the first pair of feet two oval apertures, covered by a
tense membrane, which De Geer^ had already figured. Behind
this there is a vesicular expansion of the air-tube of the fore-feet
and at its anterior margin a nerve, which coming firom the first
thoracic ganglion, spreads out into a band-like swelling in which
oval, granular bodies, together with long pediculated, remarkable
rods, are contained. In Acridium and Truxalis there is situated in
the first segment of the abdomen, on each side above the third pair of
feet, a tense membrane, behind which there is a vesicle filled with
a clear fluid : this vesicle is surrounded by an air-sac, and to it
there nms a nerve firom the third thoracic ganglion, which is also
swollen, and in the swelling exhibits similar rod-like bodies to
those which in Locusta occur in the nervous swelling of the fore-
feet.
There still remains something to be said by us respecting the
organs of motion in Insects. The antennae of insects are attached
to the homy covering of the body, which forms an external firame-
work, a dermal skeleton. This ought not on that account, as has
sometimes happened in consequence of incorrect and confused notions,
to be put on a par with the skeleton of higher animals ; for the bones
or cartilages which form the fi-amework of vertebrate animals belong
for the most part to the neural skeleton, that is, the most essential
and central parts that compose the column of the vertebral skeleton
protect the spinal cord and brain and separate them firom the
rest of the body*. Yet there are parts present in Insects which
^ For a more detfiled description I refer to the obsenratioiifl of VoN SiSBOLD him-
Belf in £bioh80N*b ArchivfOr Naiurgesch, 1844, a. 53—81^ Ta£ i. With every con-
sideration for Siebold'b great merita in the anatomy of the lower animals, I yenture to
express modestly my doubts that in insects organs of sense can occur in such an
unusual situation. The eyes on the margin of the mantle in Peeten and Spondylua
afford little support to this -new, inasmuch as the type of the aeej^alous molluscs, haa
just as little claim to the posaession of a head, aa that of the Acal^[>ha and Echinoder-
mata,
' It ia a merit of Qaxo% well deaerving of acknowledgement^ that he recogniaed
and clearly defined the difference between the dermal, the yiaceral and the nervous
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284 «* CLASS YIII.
may be looked on as the mdiments of a neural skeleton. There is
in each division of the thorax a process, which often has the form of
the letter Y, supports the nervous cord, and by the expansion of
its two branches, which are directed upwards, partly covers it.
To this process Audouin has given the name of Eniotharax; it is
even found in the head and sometimes in the first abdominal ring.
These are the same processes which Treviranus had ahready com-
pared to vertebrae*. These vertebrsB, however, are not joined
together to form a spine, but are separated from each other by
certain spaces. The dermal skeleton of Insects consists of a pecu-
liar substance to which Odier gave the name of Chitine, Lassaione
that of Entomoline^j which occurs also in the integument of Arach'-
fwtdea and Crustaceay and which is not soluble in caustic potass,
neither is rendered yellow by nitric acid, like corneous tissue. It
bums without fusion or intumescence. It forms different layers of
which the most external is composed of irregular cells ^
The arrangement of the muscles is different in the different
orders of Insects, nay, in the same insect in its different states, if it
undergoes complete metamorphosis. The difference between the
muscles of the thorax and of the abdomen, which in the per-
fect insect is so marked, is absent in the vermiform elongated
larva, for instance, in caterpillars. Along the dorsal and ventral
surface riband-like muscles run longitudinally ; there are different
oblique muscles in addition. The muscles present in their bundles
transverse stripes, as in vertebrate animals^. They are usually
skeletoii ; see especially his excellent work Von den Ur-Theilen des Knochen- und Scha-
len-GertUt€8, Leipzig, 1826, folio.
^ Verm. Schriften, IV. s. 229, 230.
■ See Odibb, Mem, de la Soc. d'Hist. Nat, de Paris, i. 1823, pp. 29 — 42, and the
later Investigations of G. Sobmidt, Zur ver^chenden Physiologie der Thiere, Braun-
schweig, 8vo. 8. 33, 52. *
> Comp. H. Fbby and R Leuokabt in the new edition of B. Wagneb, Lehrb, d&r
Zootamie, revised by them, 1845, PP* 3 — 5 J also H. Mayeb in MusLLSB'ti Arckiv,
1842, 8. 12 — 16. In the skin of the Silkworms and their pupae (and also in other pupae
of lepidoptera) there are found tUUaU cells, which Plattnsb compares with the bone-
corpuscles in the osseouB tissues of vertebrates, Mublleb's Archiv, 1844, b. 4&, 47.
^ Since in every ring of the Larva't body the same arrangement of the muscles ia
observable, the number of the muscles, when those of all the rings are counted together,
is very great. Ltonnbt found in the larva of the Willow-hawk more than 4,000
muscles.
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INSECJTS, • 285
white, sometimes pale red or brown-yellow, and are not snrroimded
"by a fibrous membrane, so that, when detached from their inser-
tions, thej may be spread out like a pencil.
Many insects are distinguished bj special art-instincts, by their
cunning in overpowering their prey, by the care for their eggs or
young, by the construction of artificial habitations, &c Their
field of observation is greatly extended by the high development of
their visual organs. The fEMmlty of indicating beforehand changes
of the weather by certain actions, by which some insects are dis-
tinguished, rests probably on their finer sense of the difierent con-
ditions of the atmosphere, since the air penetrates their whole body
by the tracheae. In this respect, as in so many others, they resemble
birds amongst vertebrate creatures, whose air-sacs and hollow bones
are in connexion with the respiratory organs, and in which also
a perfect correspondence between the external atmosphere and the
internal parts of the body is thus maintained.
Manifold is the damage which Insects occasion to us, as well
by spoiling our luxuries, as by injuring or annihilatmg our pro-
perty. On the other hand they procure for us many advantages,
amongst which I need only name silk, wax and honey. But much
more important still is the use they supply in the great economy of
nature, and therefore indirectly to us*. The injury which Aey
sometimes cause us, is not only more than counterbalanced by
these benefits, but is for the most part only a consequence of the
beneficent action itself. It is these small animals that nature em-
ploys for her great purposes, and which effect by their numbers
what the largest animals working separately are unable to perform.
Hence they are less dependent on the will of man, which indeed
here and there may be able to destroy a species, but is unable to
exterminate it throughout entire districts, as it has annihilated
different mammals in lands which they formerly inhabited. Insects
maintain the due equilibrium in the vegetable kingdom, diminish
putrefection, and lastly afford to many other animals, especially
birds, an abundant and ever present nutriment.
The geographical distribution of Insects opens a wide field for in-
quiry, which however has only been lately entered. Many families,
^ On the benefit and the injury cansed by Inseots, see in detail KJABT and
Spbkob, IfOrodncUon to Eniamohgy, i. pp. 80—338.
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286 CLASS YIII.
nay, whole orders of insects, have been collected m other parts
of the world with more or less of inadvertency by travellers and
collectors, or at least not with such care as to allow us to deduce
from the species yet known any general rules. Thus, for instance,
if we compare the number of Diptera found out of Europe with the
European, and thus form a measure of the proportion which sub-
sists between exotic and European species, we shall arrive at a
conclusion which will certainly vaiy much from the truth. Some
genera are proper to the warm regions of the earth alone, and in
Europe are represented either not at all or only by a few species
from the southern part of our quarter of the earth, as the CicadcR
{Tettiff anted Fabr.) and the genus Phasma. On the whole our
knowledge of some orders of insects, especially of the Hemiptera
and Orthoptera, would be very confined, were we to limit ourselves
to European insects. The distribution of the same or very similar
species in cotmtries widely distant from each other, the remarkable
richness of the same natural group giving a special character to
FauncB, often depends upon the same quality of the soil and a
resemblance in the vegetation. Thus for instance the insects of the
sandy regions of Asia near the Caspian Sea correspond to those of
North Africa, nay even to those of the Colony at the Cape. A
similar remark may be made in relation to the class of Mammalia,
It is this remarkable abundance of certain forms which leads us
at first sight, and even without having determined a single species,
to distinguish a collection of insects from the Cape of Grood Hope,
for instance, from one from the Indian Archipelago; Mtflabrisy
Pimdia {Trcu^hi/notus, Sepidiurn)^ BracJiyceruSy Acrydtumy Mantisy
&c. in the first, Phasmay PenUUomay numerous resplendently coloured
Papilioma in the second, give to the two a totally difierent appear-
ance. Some species of insects are confined within very narrow
limits; others, as for instance, Papilio cardut, Pluaiagammay occur
in a considerable portion of the old world, and also in North
America ^ — The limits of vegetation on mountains, as well as near
1 On the geographical distribution of Insects comp. Latbullb, Inibrod/vifitiM^ d la
Olographic gSn^rale des Arachnides et des Imectea, Mim, du Mutium, iv. 1817, pp.
37 — 67; the same in JXct. CUus. d^ffiat, Nat, vn. 18^5, pp. 290— 296, and especially
Laoobdaibe, Jntrod. d VEniomol. n. 1833, pp. 528—619 (the best hitherto known on
this subject). See also C. G. Reioh, BeUrag zur Lekre von der geographiichm Ver^rei-
•
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INSECTS. 287
the poles, are also osnallj those of the distribution of Insects ; some
species live even on snow and ice, as a small black insect from the
order Thysanura^ which some years ago (1839) was first discovered
1)7 D^OR on the glacier of Monie Bosa, and after him is named
Desoria glacialts.
tung, der IntecUn, vMbewndere der Kitfer, Nov, Adt, Acad, Oca. Leop. Car, xn. i,
pp. 805— S40.
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SYSTEMATIC
ARRANGEMENT OF INSECTS.
CLASS VIII.
INSECTA.
Articulate animals, with articulate feet. Head distinct, fur-
nished with two antennae. Heart situated in the back, similar to
a longitudinal vessel. Organs of respiration tubular, branched,
distributed throughout the whole body (troucheci). Sexes distinct.
Section I. Apiropoda.
With numerous feet With thorax not separate from abdomen.
Order I. Myriapoda.
Wingless. Feet numerous (24 or more), disposed according to
the length of the body, terminated by a single claw. Two clusters
of simple eyes, in various number ; in some no eyes.
Myria/poda. Leach and other modem writers consider this order
as a class, and wish the name of Insects to be restricted to six-
footed articulate animals, of which the body consists of three prin-
cipal parts : head, thorax, and abdomen. Here there is no separar
tion between thorax and abdomen, but the whole body is parted
into rings. The reason why we have placed these insects at the
beginning of the class, is to be found in their resemblance to the
ringed worms, to which they are related, Dot in their external form
alone, but also in their internal structure ; for even the six-footed
insects, which imdergo complete metamorphosis, often in the larval
state approximate to the myriapods. We willingly admit, on the
other hand, that the myriapods accord with certain Crustaceans, and
even form an unconstrained transition to them. But this natural
transition is in some degree broken by other insects, which on the
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INSECTA. 289
contrary pass through the order of the Parasites to the Arachnids.
The Arachnids again condact us by another road (through the
genus Scorpio to ZimuliLs) to the Crustaceans. Thus is the entire
animal kingdom a net everywhere connected, and every attempt to
arrange ftnimfl.1g in a single ascending series must necessarily fail of
The oral organs consist in most of two mandibles, whidi are
toothed at their broad extremity, and of a fbur-lobed underlip whose
two lateral lobes represent the two low:er jaws (maxiUasy In some
the second pair of feet forms, by coalescence of their basal pieces,
a sort of second underlip, which covers the oral organs and the first
pair of feet from beneath. In certain species the jaws and the lip
are represented by pointed organs which coalesce to form a sucker;
but by far the greater number are manducating insects. Myriapods
in the first period of their life have fewer rings and only three
pairs of feet ; as they grow new rings arise and the number of feet
is augmented. In this respect also they resemble the ringed-worms,
whilst in the metamorphosis of Insects the homologous parts, rings,
segments, are not multiplied, but are developed unequally or are
united, to form the different divisions of the body in the peifect
Insect The number alBo of simple eyes increases during the
development of myriapoda
These Insects live in obscure places, under the bark of trees and
on the ground under fallen leaves, stones, <&c. .
Ck>mp. on this order amongst others : Leaoh, A tabvla/r View of the ex-
ternal Chcaraden of four Classes of Animals which Linn^ arranged under
Insecta, Transact, of the Lirm. Soe. xi. 1815, p. 306, &c. ^p. 376—386) ;
P^ Gkbyais, Stvdes pour servir d rifist. not. des Myriapodes, Ann, des Sc,
not. sec. ^rie, Tom. vn. 1837 ; Zool,, pp. 35 — 60 ; also, 30 S^rie,Tom. n.
1844, Zool., pp. 51—80; J. F. BRAVJyr, ReeueU de MSmoires reUoifs d
Vordre des Inaectes Myriapodes {extrait du BtUleUn publi^e par VAcad. des
Se, de St. Peter^^ourg, Tom. V.— ix.) 1841, 8vo ; A. F. Waoa, Observaiions
sur les Myriapodes, Bivue zool. publiiepar GufaiN, Mars 1839, pp. 76 — 90;
G. NewpoHt, List of Myriapoda in the British Museum, Ann. of Nat. Hist,
xm. 1844, pp. 94—101, pp. 363 — 270 ; 0. L. Koch, System der Myria-
poden, Begensburg, 1847, 8vo min.
Family I. Jultdce. {Chilognatha Latr.) Anterior feet not
changed into organs of mandncation ; rest of the feet in most of the
segments bigeminal, slender, short, of the two sides approximate,
inserted nearly at the middle of the abdomen. Antennae short,
filiform, with six or seven joints. Organs of copulation sitaated at
the anterior part of the body.
VOL. I. 19
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292 CLASS Vlll.
apertures, vhicli form a ix)w on each side of the body, to be air-slits
{stigmata). The true air-slits lie quite beneath, near the insertion
of the feet, (Savi op. cit. Tom. i. p. 334, Buemeioteb in Okek's
IsiSy 1834, B. 134 — 138. Ta£ i.) These animals can roll themselves
up spirally, with the head in the middle ; in which posture they
pass the winter. In copulating they bring the anterior part of the
body in which the sexual organs are situated (in tiie female in the
fourth, in the male in the seventh ring), perpendicularly upwards ;
the posterior part of the body rests tortuously on the ground In
the spring the female deposits her eggs in masses of sixty or
seventy in a hole excavated for the purpose under the groxmd; after
three weeks or more the young make tiieir appearance, but still
continue to adhere for some days by a string to the shell, which has
burst longitudinally, without motion, and surrounded by a proper
membrane; at that period they have no legs at all; as soon as
they have got three pairs of feet, they separate themselves from the
shell; they have now a great resemblance to the larvae of some
Coleoptera; soon the number of rings and feet begins to be increased
in that part of the body which is seated in front of the penultimate
ring.
8p, JutuB iobuUmu L., Koch in Panzbb n. Hebbioh Schjeffb^ DeutsekL
InB. Heft 163, No. 7. Some foreign species attain a length offive inches and
more, as Spirotrepttu Javanieua BBAinoT, and Spirobolus apinoswDB'HjLAXr,
Mm. Lugdan. &c. The last species, from New Zealand, is black, "with
different rows of spines running longitudinally.
Olomeris Latr. Body elongato-oval, gibbous above, plane or
concave below, contractile into a ball, with the first segment made
up of a small dorsp^l lamina, semicircular, the second broader than
the rest, the last semicircular. Antennae thick, with the sixth joint
the largest.
A, Eyes on both sides eight ; seven disposed in a curved Hue,
the eighth on the outside, out of rank. Joints of antennae seven,
the penultimate including the last. Sub-genus Glomeria B&AifDT.
Sp. Glomeria limbakt Latb., Ohm. marginata Leach, Bum^. Condd,
ghi6r. PL 57, fig. 3, Oniscus gonatus Paiheeb, DeuttehZ. Int., Heft 9,
No. 13, Bbandt u. Ratzebubo, Medizin. Zool. n. Tab. xin. figs. 7 — 10-
These animals resemble in external form some of the Onucitiea {Onigetu,
ArmadUla\ and are even met with ii^ apothecaries' shops, amongst the so-
called MiUipedcB, mixed up with ArmadiUo qfficinarwn. Gomp. on the
anatomy of this insect Brakdt in MvKLLNL*a Archiv, 1837, s. 320 — 327.
Taf. XII., and ReeueU ieMhnoirea, pp. 152 — 158.
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INSECTA. 293
B. Two oval clusters of numerous eyes (50 and more), trans-
verse^ situated in front of the antennse at the sides of the head
Sub-genus SplhCBropoeus Brandt. Joints of antennse six^ the last
large, truncated at the point.
Sp. SphoBntpoBUM intignii Brandt, Zephr<inia ovalU Gbat ; a large species
from JtkYA, figured in this ManuaL
Sub-genus Sphxrotherium BRAiayr. Joints of antenns seven, the
sixth oblong, the seventh the least.
Most of the species of this sab-diviBion are from the Cape of Good Hope.
The genera SpharopoevM and Spharothtrium are exotic, and seem to repre-
sent Crlomeris in warm regions.
Family II. Scohpendrtdce. {Chtlopoda Latr.) Second pair
of feet cheliform, terminated by a strong hook, which is perforate,
covering the first pair of feet and the organs of manducation
beneath, joined at the base, dilated, as if forming a second labium.
Body depressed, covered above and below with homy scutes, the
sides membraneous. Feet lateral, mostly a single pair in each
segment, the last longest, extended backwards. Antennae usually
more slender towards the extremity, with numerous joints (14—40
and more). Organs of copulation situated at the posterior extremity
of the body.
These animals live on animal food, insects, <&c. Their nippers
(feet of the second pair) contain the excretory duct of a poison-
gland, which secretes a fluid deadly to small animals, as De Geer^
and Latreille' observed in flies ; the bite of the large native
species may cause great pain in man and violent inflammation and
swelling I
Gomp. on the fiunily G. Nbwpokt, Monogratph cf the Clou Myriapoda,
Order Ghilopoda. Trantact, Linn. Soc, xix. p. 265.
A. Tarsi long, slender, midtiarticulate. Antennae setaceous, as
long as the body.
^ InteeL vn. p. 557, on the bite of Lithobiut forficatm,
' Hut da Cruet, et dee Ine. vil p. 88, on the bite of Scutigera araneoidee,
' LXBDWEITHOECK fiist obserred and figured the perforated nipper, Vervdff, der
Brieven &c pp. 138 — 140, fig. 10 (59th letter), and Sevende Vervotg. der Brievcn,
pp. 184 — 186 ,(ii4th letter).
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294 CLASS VIII.
BctUtffera Lam. Cermatia Illig. Feet elongate, especially the
last. Body behind the head covered with scutes above, the fourth
longer than the rest. Eyes two, compound.
Sp. ScuJtigera araneoides auctor. {Scolopendra coleoptrata L.?) DuM^. Cotu,
g4n. PI. 58, fig. 6 ; Gu^bin Iconogr,, Insed, PI. I. fig. 7 : this animal hu
15 pairs of long feet, which readily fall off as in gnats and harvest-spiders
(Phalangia) ; it is found in Fnwce and other parts of Europe. L^N
DuFOUB has communicated some anatomical details regarding it in Ann.
det Sc. not. u. 1824, pp. 92 — 98. The compound eyes of Scutigera may
be looked on as a special anomaly in this order; the cornea presents
hexangular fafettes, as already figured by Savxgnt, Deacr. de rJEgtfpte,
Myriapodes, PL i.*
There are still some other species in the warm regions of the old and
new world, but they appear to me to be not sufficiently determined. The
figure of Pallas {Jvlua araneoldea in his Spicilegia ZooL ix. Tab. iv. fig.
16), ordinarily considered as synonymous with Seutigera araneoides, is cer-
tainly a different species. The figure of Panzer, Deutschl, Insect, Heft
50, No. 12, under the name of Scolopendra coleoptrata, howeyw it be still
referred to by later writers, has no relation to Seutigera, but appears to
represent LUhobius forfica^ua.
B. Tarsi short, uniarticulate. Autennse shoi-ter than the body.
Liihohiua Leach. Superior scutes imbricate, unequal. Fifteen
pairs of feet behind the chelifonn feet. AntennaB with numerous
joints, in adults above 40. Two groups of eyes in the external
margin of the head behind the antennas, the hindmost eye larger
than the rest.
Sp. Lithobiiu forficatiu, Scolopendra fatjicata L., Gu£bin Icon., Ins, PL h,
fig. 6; Panzeb Deuttchl. Ins, Heft 50, No. 1$, Heft 190, No. 20; com-
mon in dunghills, under flower-pots, &c. ; 10 lines long, i^ lines broad.
See on its anatomy Tbevibanus, Verm. Schrifi, 11. 181 7, s. 18 — 33. Taf.
IV — vii., LfoN DuFOUB, Ann, dts So, not. Ii. pp. 81—^1. It has seyen
pairs of stigmata. Here also in young animals the number of rings of the
body and of the feet is smaller ; the augmentation, as the animal grows,
appears to occur in a manner different from that in Julus, so that new
segments and new feet appear not behind, but between those already
formed ; and thus it is explained that the smaller dorsal shields are between
the larger. GsBVAis, Ann, des Sc, nat., sec S^rie^ Tom. vn. Zool. pp.
57, 58.
Sub-genus Henicopa Newp.
^ In a species still unnamed firom Japan in the Leyden Museum, two Paris inches
in length, (the European species attains a length of only 8 or 10 lines), I found these
fa^ettes ^ miUim. in diameter.
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INSECTA. 295
Soohpefndra L. (exclusive of seyeral species). Superior scutes
plane, in some subequal, the posterior gradually larger in others
unequal, with larger and smaller almost alternate. More than 15
pairs of feet (almost always 21), behind the cheliform feet. An-
tennas with 17 — 20 joints. Eyes four on both sides, at the margin
of the head behind the base of the antennas.
To thifl genuB belong the largest species of this division. In these
Myriapoda there are 9 pairs of air-slits present, in the membraneous part
between the dorsal and ventral scutes (in the 3, 5, 8, 10, 12, 14, 16, iS,
and so ring), above and behind the insertion of the feet, (see my observa-
tions in the Tidtchr, voor not. Oach. en Pkynd. v. bl. 33a — 337. PI. vi.)
Comp. for the Anatomy Gaedb in Wiedemann's Zool, Magazin, 1. 1817.
B. 105 — 109 with fig., and Kutoboa, Seolopendra morntantis anaUme,
Petropoli, 1834, 4to. The species investigated by this writer is Scolopendra
cingtdarit Latb., which occurs in the south of Europe, in the Crimea and
in Egypt. In north and central Europe no species of this genus occur.
Formerly many species from different regions were confounded under the
name of Scol. morsitans L. ; the species figured by Kollab under this
name (BnmLitn*8 liuHge Insecten, Rate im innem v. BrcuUien von Db
PoHL, Wien, 1831, 4to, fig. 4) ought apparently to be referred to Scolopenr
dra tubspinipet Geby.
Cryptops Leach. Twenty-one pairs of feet behind the cheliform
feet. Eyes none. Antennae with 17 moniliform joints.
Sp. Oryptopi Savignii Leach, Seol. germanica Panzsb u. Hebbich
SoHiBPFEB, DetOtchl. Ins, Heft 143, No. a, &c. The species of this genus
are distinguished^ besides the absence of eyes, also by their much smaller
size from those of the former ^
Geophilus Leach. Pairs of feet numerous, 40 and more. Body
linear. Eyes none, antennse with 14 joints.
Add sub-genera MecUtocephaZuSj NecrophloBophagus, Gombreg-
mctiua Newport, and some others formed by Koch, Si/sL der
Myriapod. pp. 176—189.
Gomp. on this genus also, Gebvais, Gu^bin Maga», de Zoclogk 1835
(with a figure of a large species from France, QeopkUut WaUcencerii), and
Nbwpobt, Proceedingt of the Zool, Soc. 1842, pp. 178 — 181. Sp. Scolo-
pendra fiava Db Geeb^ Ine. vii. PL 35, figs. 1 7 — 30, GeophU, longicomU
Lbaoh, PA27ZEB u. Hebbioh Sohjsffeb^ DetUschl. Jne, Heft 142, No. 5.
Of thi» species Tbbvibakub has given an anatomy. Verm. Schr. u. s.
33—38.
^ Here also belongs the genus ScolopendroptU Bbandt, that appears to differ from
Cryptopa only by having two segments more.
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293 CLASS viir.
The phosphoric light which some species diffuse {ScU. dectrica phog-
phorea) is to he ascrihed to a fluid that passes upon the skin from open-
ings similar to those in the JulidcB {Waga), Numerous ohservations are
recorded that myriapods of this division, after having caused lingering
head-ache, have heen sneezed forth by men from the nose (F. Tibdemank,
Von lebenden WUrmem u, Jnsehten in den Gtruchsorganen des Menachenj
Mannheim, i844i 8vo, s. ii — 17, to which examples many others may
be added). To reject the observations, is certainly more easy than to
explain the continued life of these insects in such an unusual situation.
ScolopendreUa Gebv. Antennae with twenty joints, moniliform.
Ocelli two. Segments of the body sixteen. Mouth not chelate,
suctorial. Habit of Geophylus.
Comp. Ann. des Sc, not, troisibme S^r. Zool. Tom. n. p. 79, 8yo, PL 5,
figs. 15, 16. Is this its place 1 Are the sucking Scolopendnt to be com-
pared with the SiphonizaaUia chUognatha BSANDT ?
Section II. Hexapoda.
Feet SIX. Thorax separate from the abdomen.
Order II. Thysanura.
Hexapod, apterous, not undergoing metamorphosis, not para-
sitic. (Mouth with mandibles and maxillss. Two groups of simple
eyes. Setae or a bifid tail in most at the end of the body).
Gvadvovpot from Bitrapoif a frrngty a ttift, and ovpdf thus named
from some species which have jointed threads at the posterior
extremity ; in others there is a forked tail which in a state of rest
is bent forwards beneath the body, and on extension gives a blow to
the ground, which causes the body to spring upwards : hence they
may be named spring-tails. In some however the posterior part of
the body has only two small conical hairs (FodtiraJlmeUiria L.) or no
appendage at all. The body ia generally elongate. Some are very
small, and even the largest species are scarcely half an inch long.
They love moisture, and live on the ground, imder stones, in mould
or old fallen leaves, under the bark of trees, in dark corners of houses,
Ac. They undergo no metamorphosis, but change their skin often.
The intestinal canal is straight, the stomach wide. In Z^fnama
there are two, in JSmyrUhtMtLS, according to I^icolei, three vessels for
secreting urina The external sexual organs are placed at the
posterior extremity of the abdomen* In the female of Zepisma a
split tube or borer is found which serves for depositing the 6ggSL
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IN8ECTA. 297
The nervous system in SmyrUhurus presents only four ganglia, of
vhich the first and second lie very close together, the first above,
the second below the (esophagus (Nxoolet) ; in Lepiama Mccharina,
on the other hand, Treviranus found twelve ganglia. The two
eyes, which in most species are small, consist of a group of eight,
sometimes six or seven, in Fodura Jimetaria of fourteen simple
eyes (Nigolbt) ; in LepUma saccharina twelve simple eyes are
counted on each aide.
These Insects, however nearly alli<$d to the myriapods, approach
still more nearly to the Orthoptera, and especially to the genus of the
earwigs {Forficulas) ; see this Manual^ first edition, l 1828. pp. 204,
205. But if, on account of tins affinity, any one would place Lepiama^
or even the whole order of Tkysanuraf amongst the Orthoptera, I
think he would go too &r. Time, the touchstone of all things,
would reject the union. If Latreillb had united the Thysannra
with the Orthoptera in one order, zoologists of the present day
would for certain have perceived long ago that these Insects were
not in their place.
Comp. on ihiB order: TBKViRAinjs Verm, Sekr. n. b. ii — 17. Tab. 11.
ni. and iv. figs, i — 5 (on the anatomy of the genua Lepitma) ; Burmxistsb
in Oksk's Jits, 1834, s. 137, 138 (on the respiratory organs and the
digvMia of Lepisma, whose presence Trsvirakub had erroneously denied) ;
Latbkillb, J}e rOrganitaiion extSriewre et eomparie des Insecte$ de Vordre
det Thyaanourta, Nouv» Ann. du Mu$, i. 1832, pp. 161 — 187 ; B.Tbmfue-
*0K, TKyaanura ffibemio, or Deaeriptuma of auch apeciea of apring-taited
Inaeda, Podura and Lepisma, aa have been oBaerved in frdand, Tranaad.
of the EnUmol, 80c. Londm, 1836, i. pp. 89 — 98, PI. zi. xii.); H. Nioolst
Reeherchea pour aervir d VSiat, dea Podurellea, 88 pages et 9 planches
{NeueDenka^r, der dUg. Schtoekeriache Oeaellachtfi fOr die geaamnU. Nalw
wiaaenach, 3d, vi. Neucfaatel, 1841, 4to) ; Burmubtib, Handh, der Ento-
mologie, ii. 2, 1838, s. 443 — 458 ; Gkbvaib in Walkxsaxr, Stat, dea Ina.
apUrea (Suites h BurroN chez Bobbt), m. 1814, pp. S77—4S6, ko.
Family III. Leptsmeme. Body elongate, mostly covered with
minute scales. AntennaB setaceous, with joints numerous, very
short. Palps four, elongate. Abdomen of nine or ten distinct
^segments, besides other appendages supplied with three (or more
rarely with two) long setss, articulated towards the extremity.
Machilis Latr. Antennse inserted under large, contiguous
(compoxmd ?) eyes. Maxillary palps, exsert, filiform, long. Body
arcuate, with convex back, saltatory. Three setae at the extremity
of the abdomen, the middlemost larger than the lateral.
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298 CLASS YIII.
Sub-genera Petrobhu^ Machilia Leach.
Sp. MaekUu marUima Lath., M. polypoda Duifia. (not Latb.), Cons. gSn.
PI. 54, fig. a, ftc. Besides the jointed threadB (two on most of the s^-
ments of the abdomen), GufRiH has obaenred in MaekUu Teaides also on
the same segments, which he compares to gills ; Ann. deg Se. not. sec.
S^rie, Zool. V. pp. 374, 375, Iconogr., Int, PL n. ^. i t, These animals
remind us of lanrse of Ephemera.
Lepisma L. (in part). Antennae inserted amongst eyes, small,
remote. Palps moderate. Body depressed, squamose, not saltatory,
terminated by three subeqoal setae.
Sp. Lepisma saceharina L., Gu^BiH Iconogr., Int. PL 1, fig. i ; GsonmOT
Im. Pwr. U. PL XX. fig. 3 ; Hookb Micrographia 1667, Tab. 33, f. 3,
pp. 108 — 110. It is said that this insect was imported into Europe from
America^ where it lives in sugar: in Sweden it is rare, according to Db
Gbbb ; it is not spotted, covered with silvery scales, which under the
microscope are finely ribbed longitudinallj, and differ much in size and
form. This insect runs very rapidly, and with us is usually found in
clothes-chests, between damp books, &c.
Sub-genus Lepismina Geky. is distinguished by the body plane,
the thorax very broad
Sp. Lepwna awea L^v Dutoub, Ann. det Se. nai. Tom. XXU. p. 419. Tab.
xm. tg. I.
jNf'ote. — Genus NicoleHa Gekvais is distinguished by defidct of
eyes, and body not scaly ; Campodea Westwood, Ann. of NaL HiaL
Tom. z. 1842, p. 71, by the same characters and by the tail com-
posed of two setffi only. Is it Podwra cmbvlans L. %
Family FV. Podurelke. Body in some elongate, in others globose,
broad, covered usually with hairs, sometimes with scales also.
Antennas with few joints, or with four longer joints at the base, at
the apex only with short numerous joints. Palps inconspicuous.
Abdomen composed of only six or fewer segments, mostly termi-
nated by a forked tail, during rest bent forward beneath the
abdomen, serving for springing.
Podwra L.
Spring4ail (properly foot-tail, since the tail is used like feet for
motion). They are small insects, which live on trees, under bark,
under stones, on moist earth, &c. Some are met with on the sur&oe
of water. Since the tail, according to Waga, is at first wanting
in young animals (Ann, de la Soo. JEniom. de France, xi. quoted by
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INSECTA. 299
Ebichson, JakreaheridU, ArcLf, N<Uwrge9ck. 1843, a 270), it is poe-
aible that sometimes young specimens of Podura may be placed
under the genus Lipura (Anurophorus Nicol^) ; but this entire
genus is not on that account to be rejected, for it is very usual that a
young form in a natural fiunilj should be represented by a per-
manent generic form. The most remarkable deviation in this familj
is that of the genus Anura Geryais (AchartUes Nicolet), where in
place of jaws there is a suctorial mouth j see Nicolet op. dt. p. 33,
PL IV. fig. 5.
Sminthurua Latr. Thorax and abdomen conjoined into a
globose body. AntennaB geniculate; terminal portion of antenna
aetaoeons, mnlti-articnlate.
Gu^DT, leonoffr, Ins. PL i, fig. 4. (Add sub-gen. Diq^rUma Boubuet).
Podura Latb. Thorax separate from abdomen ; bodj elongate,
with abdomen narrow, oblong.
A. Mandibles and maxUlse none. Suctorial tube conicaL
Anwra Gerv. (Achorutes Nicolet not Templ.) No forked taiL
Abdomen rotundate behind.
B. Mandibles and mazillse.
a) AntennsB shorty with four or five joints.
Lipura BuBiL {Anurophorus Nicolet). Body not scaly. Abdo-
men, instead of a forked tail^ terminated bj two very small conical
diverging tubercles.
Sp. Podura jtmeUma L., Upwra ofmhulaiu GsBV. {ntU, Pod, amftuZoiM L.),
Bk Gsbb, vn. p. 33. PL 3, figs. 5, 6. Nioolbt op. dt. PL 5, fig. 1, ooni-
mon in gnrden-mould and in flower-bedsy ivoiy-white^ i lino long.
Achorviea Tekpleton^ Hypoga,atrwru8 Boxtblet, Podura Nicoletf.
Body not squamosa Tail forked, short
Sp. Podura aquaiica L., Db Gbkb, Int. vii. PL n. figs. 11— 13, Nioolkt
op. dt. PL 5, fig. 4 ; one line long, very common with us, forma large
black spots on the water, near which other white spots are floating, fonned
by the cast skins.
5) Antenme longer than head.
Genera Isotoma Boublet {Degeeria Nicolet, Desoria Nia),
Heterotojna Bourlet, OrcJiesella Templeton, Lepidocyriua Boublet
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300 CLASS VIII.
{Cyphodeirus Nicolet), Tamocerus Nicolet {MacroU>ma Boub*
LET.)
Sp. Detoria s^acialu Nicolet, L L PL 5, fig. 10; first found in 1849 on
Monte Soaa, afterwaixifl on the Unter-Aar spader; see AOASSIZ Geoloffitehe
Alpenreuen von D^SOB, BeuUdi yon C. Vogt, Fnmkt ft. Main, 1844. Sto,
8. 181, 182.
Order III. Parasitica.
Hexapod, apterous, not undergoing metamorphosis, parasitic.
Two simple eyes, sometimes none.
These animals, also named Epizoa (as opposed to Entozoa, our
fifth class, see above), cannot well be otherwise defined than by the
short account that we have given of them. The absence of a forked
tail or of setsB on the abdomen*, distinguishes them indeed from
most but not from all Insects of the former order. The flea and
some wingless species of the order Diptera are distinguished from
these parasites by their undergoing complete metamorphosis.
Comp. on thia division C. L. NrrzscH, Die Familien und OaHungen dcr
Tkieriniecten (insecta epizoica), in Gerxab u. Zutcksn, Magazin der Ea-
tomologie, m. Halle 18 18, s. «6i — 316. Here however the parasitic Dtp-
tera (Hippobotea, Nyderibia ftc.) are included in the same division. Netzsch
arranged the rest according to their oral organs, those in which they are
suctorial, amongst the ffemiplera, those in which they are manducatory,
amongst the Orlhoptera; two orders in which the inclination to abortion
of the wings is evident^ and which undergo an incomplete metamorphosis,
which therefore in the wingless genera can shew itsdf as cbazige of skin alone.
See also GuBLT U^>er die auf den ffata-Saiigetkiiren und Hwutvdgdn
lebenden 8chmarotzer'In$ecten und ArcKkniden, Magaain fOr die geaamte
ThierfieHkunde, viiL 1842. s. 411 — 433. Tab. iv. and iz. 1843. s. 1—24-
Tab. X. Some figures are also found in Ltonbt, Recherchee tur differentee
eapices d^Ineectes, (rnvrage paethume, Paris 1833, 4to ; DsinrT, Monographic
Anoplurorum Britanniee, or An Etaay on tke Britith Speeiet of Paratitea,
London, 1842 ; a work of detail which I received too late to make use of.
Family V. Hcematopitia s. PedicuUna, Mouth anterior, com-
posed of a rostellum, retractile, vaginate at the base. Tarsi uni-
articulate, with single arciLate claw^.
Pedicvlus L. (exclusiye of several species). AntennsB filiform,
quinquearticulate. Vagina of the rostrum aculeate at the point.
^ Hence the name Anoplwra Lsach. See his work On ike PamXlie$y Stirpee and
Genera of ike Order Anoplura, Zoological MieceUany, in. 181 7, pp. 64 — 67.
3 By some writers this hook is considered as the second joint of the tarsus.
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INSECTA. 301
The species of this genus occur only in man and some mammals,
whose blood they suck. Their motion is sluggisL
The louse of the human head has been investigated anatomically
by our Swammebdam. The intestinal canal is straight, with a large
stomach ; there are four vessels secreting urin& Each ovary con-
sists of five tubes. The nervous system consists, besides the cere-
bral ganglion, of three large ganglia in the thorax, so closely placed
behind each other as to touch ; from these ganglia the nerves of the
feet arise, and from the last ganglion arise in addition six nerves
which are distributed through the cavity of the abdomen.
See SWAMICSBDAV Bijbd der ntOuur. hi 63—^6. Tftb. I. u.^
Sub-genera Fhthirua, ffcBmatopinus, PediculuB Lsach.
Sp. Pedieidus humamis capitis, PedictUtu eerviealia Lkach, Db Gkib ln»,
vn. Tab. i. fig. 6, DumI^il Cotiaid. gH. «. L In$. PI. 53, figs, i, a,
Gu^BiN Icon, Im, PL a, fig. 5. (Comp. also Swammkbdam, and a gigaDtio
figure twenty inches long by Hooks Micrograph. Tab. 35). The larger
spedes, considered by LlNN^us as a variety, which lives upon the body
and amongst the garments, differs by the less deep indsures in the side of
the abdomen at each ring, by a thorax broader behind, and, as Guiaiir has
remarked, by longer antennae. Pedicnhu humamu corparia Db Gbbb,
Ins. L 1. fig. 5, {Pediculus humanus Lbaoh, Pedieulus vesUmenH Bubm.)
As a third parasitic species of man may be added Pedicidis pubis L.,
Phihirusinguinalis Jj^Aon, HsDiExper. circa generaHonem Insectortmi, Am-
stelodani, 1688, lamo. Tab. is, fig. superior, GnisiN op. dt. fig. 17.
Family VI. Mallophaga. Mouth supplied with mandibles
and maxillss. Tarsi biarticulate, with a single claw or with two.
On Mammalia, and especially on Birds, different parasitic Insects
are foimd, which wei-e placed by Linnjeus in the genus FedictUua,
but which differ from it by the presence of jaws on the under
surface of the head. De Geeb, who discovered this character,
justly held it to be so important and essential, that he placed these
animals in a distinct genus, to which he gave the name of Bicinua
^ liBBUWENBOBCK investigated the male louse (which is rarer and was unknown
to SwAMUBRDAM, op. cit. bl. 83) ; he found two testes on each side of the body.
This and other remarkable peculiarities in Pedicukta hum. corporis are found in
Lkeuwenhobck, sesde Vervolg. der Brieven, Delft, 1697, 988te Missive, pp. 187 — 217.
See also Vierde Vervolg. der Brieven 1694, 77ste Missive, pp. 587 — 591, where the
head is described and figured. The homy sheath of the penis L. described as a sting
at the posterior part of the body.
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302 CLASS viir.
{Mim. pov/r servir d rffut. d'Ins. vii. p. 69). J. F. Hermanit
changed this name, which had already been given to a genus of
plants, toiVirmtA9^; Nitzsch, who adopted this name for a sub-
genus alone, named these animals MaMophaga, from fjia\Xo^,Jleece,
since they live on hair, on scales of the epidermis andv feathers, and
not on blood. In birds there are no lice of the former family
known — all the bird-lioe belong to the MaMophaga. In some are
found small, moveable, oblong organs in front of the antenna, like
a second pair of imperfect antenna, which Nitzsch names trahe-
cvloB, The eyes are oflen difficult to distinguish : in some they are
entirely wanting.
A. Antennae filiform. Maxillary palps none, labial very short,
biarticulate.
Phihptems NiTZSCH. Antennee qninquearticulate. Claws of
tarsi two.
Sp. Ph&opterui haculut Nitzbch, Redi op. cit. Tab. n. Jigura wpericr,
Ltonibt op. dt. p. 41, pi. 5, fig. 10, GUBLT Magae. /. d. ge$, ThierheOk.
vm. Tab. IV. fig. 9, on Pigeons. All the species of this genos live upon
birds. Here belong the sub-genera Ooniodes, LipeuruB, Nirmus and Doe(h
phorue of NiTZSOH, together ^th Ooniocciea Buaic. and Omithobiut'DmwY.
Trichodecies NiTZSCH. Antennae triarticulate. Tarsi with a
single claw.
The species of this genus live only on Mammalia. Sp. Triehod, latHi
NiTZSCH, JtictHfu canU Db Gbbb^ Ins. vn. Tab. iv. fig. 16, Gublt op. dt.
IX. Taf. I. fig. I.
R Antennse clavate (four joints). Maxillary palps conspicuous.
I/iotheum NiZTSCH. Tarsi biunguiculate. Labial palps biar-
ticulate, short.
The spedes of this genus keep to birds, often of the same spedes on
which Phiiopteri also live*. Here belong the sub-genera Colpoe^ktUvm, Me-
nopon {Menopon and Nitachia Dbknt), TrinoHtm, Bureum, Loanobotkrium,
Phyaoatomum of Nitzsch. These insects move more quickly and creep
more nimbly beneath the feathers than the PhUopteri, Sp. Liotheum tub-
wqwde NiTZSCH, Ltoknbt 1. I, PI. 4, fig. 5, on crows, &o.
* MSviunre Aptirologique, 1804, p. 11.
■ Five different species of Mallophaga occur on fowls. Such names, therefore, as
PedieuUu gaUina may readily introduce confusion.
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INSECTA. 303
Oyropua NiTZSCH. Tarsi uninnguiculate. Labial palps none.
Of this genus only few species are known, wbich live on the Cavia and
the Sloth (Bradyjput tridactylua) t. e. on American specieB of Mammalia^
Order IV. Suctoria s. Siphonaptera,
Apterous, hexapod, undergoing perfect metamorphosis. Mouth
suctorial; rostellum formed of two serrated laminse and a single
more slender seta, included in a biyalye articulate sheath.
Family VII. Pulictdce (characters of the order are those of the
single family).
Pulex L. [Characters of the order.] Body compressed.
Antennae short, with three or four joints, the last large, flat,
serrated, received in a small cavity and covered with a scale.
Maxillary palps with four joints, porrect. Posterior feet saltatory.
Tarsi with five joints.
The order of sucking InsecU, which contains the genns Flea,
agrees in the complete metamorphosis with the Diptera ; but the
oral parts have little reBemblance. The flea is perfectly distin-
guished from the hemipterous sucking Insects not only by the
perfect metamorphosis, but also by the oral parts, by the presence
of palps for instance. Duces first drew attention to four smaU round
plates, which are situated on the last two segments of the thorax,
and of which the posterior pair is the largest These plates, which
in colour and substance do not differ fix>m the rest of the homy
covering of the body, may be looked on as rudiments of wings.
The resemblance with the metamorphosis of the Hymenoptera^
especially in the pupa^ together with the presence of these four
rudimentary wings, may serve to explain in some degree why an
inclination has prevailed to regard the suHoria as abortive hymen-
opterous Insects.
There are two simple eyes, which in Pvlex vespertUionis "Dvaia
are wanting. The sucker is inclosed by the flat labial palps con-
sisting of (three ?) joints, and is composed of two flat sete serrately
toothed on the edge and ribbed longitudinally in the middle (scalpeUa
KiBBT and Spence, mandibles), and a thin smooth thread of the
same length {UgiUa Sav.) There is no upper lip present, unless it be
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304 CLASS VIII.
represented by this ligula\ The maxUlcB are two small plates at
whose base are attached the feelers stretching forwards, and which
were formerly regarded as antennae ; they have four oblong joints,
of which the second and the fourth are the longest
The intestinal canal of the Flea is short and straight ; its stomach
is cylindrical^ the small intestine aa long aa the stomach, and rectum
short There are four short and wide luinary tubes, which are
implanted at the inferior orifice of the stomach. At both sides are
two saliyary vessels in fonn of round vesicles, whose efferent ducts
coalesce to form a canal on each side of the cesophagus^ which
moimts tortuously to the mouth.
See Ramdohb, Ahhandlwng Ob, d. Verdauungtwerkieuge der Ifueden,
8. 203. Tab. ZXIU. fig. 3.
Comp. on this £unily A. Duges, Becherches vwr les Caradh^
zodogiqvsa du genre Pulex, et 9ur la multiplicUe dea eep^ces qu'il
ren/erme, Ann, dee Sc. not xxviL 1832, pp. 165 — 175, PL ivi, and
P. F. BoucH^ Bemerhmgen iiber die GaUung Pulex, I^ov. Act
Acad. Ccee. Leap, Car, Tom. xvii. 1835, p. 501—508 (this writer
considers the parts indicated above aa mandibles to be lacinioe of
the under-lip, and gives the maxillary palps five joints).
Sp. Pulex irritam L. (in part), HoOKB Micrographia Tab. $4 (a lai^
fignre of full 15 Bhenish inches, or 4 decimeten in length), Duofts L L
fig. I (both figures represent the female). Ordinarily this species is con-
founded with others which occur on the dog and domestic cat, Pidex eanis
Duofts, Pulex cams and PvXexfdit BouCHi. To the last-named refer the
figures of EoBSBLy Ins, n. Muecar, atque Ctdicwn, Tab. n— IV. (Tab. m.
fig. 10, representing a male, refers according to BoucRti to Ptdex irritans).
Moreover these species can bear leaving then* natural habitat for a time,
and thus Pulex irritans may pass on to dogs, Pulex canis on to man. A
particular species also occurs on fowls, Pul, gaUinm, Schraitk, B0UOH&
All the species of this genus are small insects ; the laziest hitherto known,
Pulex gigas, has only a length of two lines, Fauna Bor, Americana of
J. BiCHABDSON, YoL TV, 1837. The hip (coxa) of the feet is as Urge as
the thigh {femur) and very thick ; the fore-feet are placed far forwards,
almost under the head. The abdomen is large, on the upper surface in the
male, concave ; in copulation the female is placed on the back of the little
mal& The eggs are white, oblong; sticky. In the sunmier season the
larvaa come forth after a lapse of six days, having no feet, and resembling
small, white worms. Lebuwrnhobck brought them up on dead flies,
BoBSBL on dead gnats and dried pigeon's blood. Blood that has fblloired
^ DuoBS, Ann, des Se, nai, sec. S^rie. Tom. vi. 1836. Zool, p. rao.
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INSECTA. 305
the prick of fleM and has dried into bUck globules, is often found dis-
persed by the flea near the eggs that are laid in the chinks of wooden floors,
in furniture, &c., and these globules form, according to the observations of
Db FiiANcn {Afui, dei Sc. not, i. 1814, pp. 440 — 443) the fayourite food of
the larve. These larvae within eleven days are full grown, then spin them-
selves up and change into pupee, from which after ten or eleven days the
perfect insects appear ; in this way a new greneration arises after the lapse
of only four weeks. Comp. Lkbuwbnhosck, Vierde Vervolg, van Brieven
1694. bL 537 — 573, 76ste Mitaive, and Kobsel, Inaeclm-Bd^$Uffung, u.
Mucken und Schnacken, s. 9—34.
In America^ particularly in the Brazils, there is a small spedes of flea,
whose proboscis is longer ; the fore-feet are not so far forward, and whilst
the labial palps are wanting, the three threads of the sucker are not sur-
rounded by a bivalved sheath. Consequently it has been proposed to
make a distinct genus of this species {SarcoptyUa Wbstwood, Dermaio-
phUui Gn^BDr). This species, which lives in the open air, and is often in
great numbers in sandy places, is PuUx penetrana L., Dum^bil, Com, gen.
9. lea Inaeda, PI. 53, figs. 4, 5 ; Gu^bin, Iconogr., InaecL PL a, fig. 9 ;
KOLLAB, JBraaUien'a vorzUglich Idatige Inaecten, ^. 5, s. 8, 9. The Portu-
guese call the animal Bicho, the Brazilians 7\tnga; it bears also the name
of Pique, Nigua^, &c. and of Chigoe amongst the English residents. This
insect penetrates beneath the skin of the feet, sometimes also of the hands
of man, and of the feet of dogs and other Tnftinmft.la ; the female, after she
has penetrated beneath the skin, expands astonishingly, whence malignant
ulcers arise, which sometimes occasion death. They infest principally newly-
arrived Europeans : see Y. Humboldt's Beiae in die ^quinociial-Cfegenden
dea neu/cn Continenta, TV, 1833, s. 90, J. J. VoN Tbohudi (who once had six
tumours thus caused on his right foot) Peru, Reiaeahizzen, i. 1846, s. 310,
311. A capuchin monk attempted to transplant a family of these insects
from St Domingo to Europe, but his zeal remained unrewarded, for his foot
in which he had harboiu^ the colony, was obliged to be amputated on thp
voyage (Kibby and Spenob, Introd. to Fntomd. i. p. 102). Comp. on this
little animal also Duais, who especially has illustrated the parts about the
month, Ann. dea 8c. not. sec. S^rie, Tom. Yi. 1836. Zoologie, pp. 139 — 134.
PL 7 b. a forked caudiform appendage, already figured by Catbsbt, and
compared by Jasvmtjb with the tail of Podura, is probably a male organ of
copulation (Gu^bik, Iconogr.); at least it does not occur in all individuals.
Order V. Strepsiptera a. Bhipiptera.
Heiapodal Insects ; (male) with four wings j anterior wings two
small moveable bodies, inserted into the sides of the thorax ; pos-
terior wings large, membranous, resembling a quadrant of a circle,
folded longitudinally like a fan. (Females apterous, apodous).
^ Nigua is the name also given to a species of Acarua (Ixodea americantu), which
also penetrates beneath the skin, and must not be confounded with this flea.
VOL. I. 20
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306 CLASS Till.
Metamorphosis complete. Mandibles two, forficate, narrow, sob-
arcoate. Palps two biarticnlate, very distant, inserted nnder the head.
(Lanrae and Pupae living paiasitically in differ^it Hymenopiera.)
This order, first distiiiguished by KraBY, cannot well be miited
with one of the othera^ and least of all with that of the Hemi-
pteroQS Insects^ as some would desire. The natmvl affinity is
difficult to determine; peihaps this order stands between the
Orthopteray Neuroptera, and HymeTioptercL. In this uncertainty
respecting the true place, we think we have some gnnmds for our
choice, to place it after the flea, in the perfect metamorphosis as
well as in the presence of four rudiments of wings in the genus
Fulex, However small, moreover, the number of inaedss of this
order may be, that can afford no sufficient reason either in an
artificial or in a natural system for rejecting it
The winged individuals, on which the characters of the order
are founded, are, as Bxjbmeisteb had sagaciously surmised, and the
complete investigations of the excellent 0. Th. von Siedold have
demonstrated, all males. In these, two large compound eyes are
present, which consLst of few fii^ ettes, separated from each other
by a raised margin. Simple eyes are wanting; the antennse are
composed of few joints, ordinarily split at the extremity into two
parts, or as if branched with lappet-like appendages. In front of
the wings are two small convoluted parts, named by Kibby Elytra,
but which do not cover the wings. It was formerly thought that
they were attached to the first ring of the thorax, and in that case
they could not in any sense be looked on as imperfect wings or
elytra; but more accurate investigation has proved that they belong
to the metaihoraXy and, therefore, may keep the name given to
them by Kirbt. They recal the short elytra of some PhaamatidiB.
The middle piece of the thorax is prolonged into a shield over the
abdomen. The tarsi have mostly four joints, in other species three
or two, and no claws at the extremity. The wings (hind wings)
are large, thin, whitish, opaque, and have some nervures which
radiate towards the circumference. The oral organs are somewhat
differently described by Savigny, whilst he regards as maanUce the
parts described by Kirbt as palpi^. The under lip has no palpi
(Westwood considers the palpi as belonging to the under lip, and
^ Saviont's oommunioationB to Lbach were publuhed by the latter in his Zoologi-
cal Mi»cdlany, m. 1817, p. 135.
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IKSECTA. 307
the mandMes aa Tnaxiilas. According to this view the mandibles
would be wanting.) These insects undergo complete metamorpho6i&
At j5rst the larva has six feet and two threads at the abdomen^
(somewhat the form of Lepisma); afterwards these feet disappear,
for the insect then lives in the abdomen of larvse of Hymenoptem,
In these HymeTioptera, also, the parasite is changed into a pupa,
which in the perfect Hymenoptera projects between the rings of the
abdomen. The wingless female remains in this situation, and is
impregnated there. It is viviparous. The six-footed larvie, which
in the genus Xenos jump like PodureUas^ were described by Kluq
and Westwood, and previously also by Yon Seebold, as parasites.
Comp. on this order W. ElmBT, Strepaipteroy a new order of
Insects, Linnasan Tramact. YoL zi. 1815. pp. 86—122, Tab. 8, 9 ;
AddendiMTiy pp. 233, 234 ; Lsach on the Rhipiptera of Latbeillb,
Zool. Miec UL pp. 133—136 ; W. R Pickerxng, ObeervcUions on
the Economy of the Strepeiptera, TramscbcL of the Entomol» Soc
London, i. 1836, pp. 163 — 172, PL xvii ; J. O. Westwood, Beecrip-
Hon of a new Strepeipieroue Insect, ibid. pp. 173, 174, PL xvil
fig. 15 ; C. Th. t. Siebold, Ueber Xenos Spheddarv/m und dessen
Sehmaroiaer, Beiirdge zwr Naturgeschichte der Wvrhettosen Thiere,
1839, B. 72—87, Tab. iii. figs. 62—74 ; by the same Ueber Strep-
siptera, in Ebichbon'b Archwf NcOwrgesch, ix 1843, & 137 — 162.
Tab. vn.
Family VIII. Strepsiptera s. Siylopidce. [Characters of the
order.]
* Tarsi with four joints.
Xenos^ Rossi. Antennee bipartite, single at the base, triarticu-
late, with first joint obconical, longer, with branches semi-round,
acuminate, not articulate.
Sp. Xenoi PedsU, Kzbbt, Lmn. Trant. Tib. 8, Tab. 9, fig. i, Kibbt and
Sfbnos, Inirod. to SnUm, i. PL n. fig. i; the larva infests Polytte fucala
(nor. Amer.) ; ftc. European species are Xenot Rouii and Xenoe Sphe-
cidarum. They are all small insects whose body is only about li line
long.
Styhps KiBBY. Antennae bipartite with stem single, biarticu-
late, upper branch triarticulate, (Antennae have six joints, the third
^ ^ivot or ^611^0$, a guest ; because harboured by other insects.
20—2
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308 CLASS VIII.
joint fonning a produced, lanceolate branch, almost equal in length
to the three last joints).
Sp. StylapamditUe, ftc. Icones : Wistwood, Introduct. Frontisp. VoL i. fig. 6;
Stylopt Speneii, Gn^. Iconoffr., Ins, PL 92, fig. i, %Z. Dalii, Lsaoh,
Zod, Miae, ui. Tab. 149, StyL Kirbii.
* * Tarsi with three or with two joints.
Halictophagus Curtis. Tarsi with three joints. Antennae with
seven joints, supplied externally with lamellae disposed pectinately.
EhncJius Curtis. Tarsi with two joints. Antennae bipartite,
with stem simple, biarticulate, and upper branch biarticulate,
elongate, slender.
Gomp. Curtis, BrUiah Entomology quoted by Westwood, Intro-
ducUon II. pp. 287, ko.
Order VI. Diptera.
Hexapod Insects with two wings, and two poisers. Mouth
suctorial, with labium not palpigerous produced into a proboscis, or
a sheath, which receives and incloses in a groove above an haus-
tellum formed of setae, various in number. Two palps (maxillary)
at the base of the proboscis. Metamorphosis complete.
Two-winged. Dvptera L., AntUa^ta Fabr. The principal works
on this order are the following : —
'J. 0. ScHELLEinBEBO, OoU/wngen der Fliegen, in 42 Kuj^erkifdn
entworfen. Zurich, 1803. 8to (with german and fii^nch text).
J. C. Fabricii, Systema AnUtcUorum. BrunsvigaB, 1805. 8vo.
J. W. Meioen, Systemaiische Beschreibung der bekcmnten Euro-
pdiachen zweijlugdigen Inaekten. Mit Kv^fertafehh. vii. TheUe, 8vo.
Aachen und Hamm, 1818—1838.
C. R. W. Wiedemann, Atisaerewopdische zwdfiUgdige Insekten.
MU Steintafel/ru 11. TheUe, 8vo. Hamm, 1828, 1830.
ffistoire ncOurelle dea Insectea JMpth'ea par Magquabt, av, pi,
n. Tomes, 8vo. Paris, 1834, 1835. (The work of Fallen, Diptera
Snecica, Lund 1814 — 1827, 2 vol., I hare not been able to inspect)
The general covering is, on the whole, very thin. The body
consists, as in the other hexapod Insects, of three principal parts :
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INSECTA. 309
the head, the trunk, and the ahdomen. On the head two compound
eyes are usuallj seen, which are Y&ry large, especially in the male ;
in the most, simple eyes or eyespots are also present, usually three,
sometimes only two in number. The antennae are placed on the
frons close to the eyes ; ordinarily they are, in comparison with
those of other orders, short Some sub-divisions of the genus
TiptUa L., make an exception to this, and the genera Mdcrocera
Meiq. and Megistocera Wiedemaks are especially distinguished by
their long antennae. The sucker consists of two, four, or six sharp
threads, which in the last case represent the upper lip (labrtMn'),
the tongue (ligula), the two upper and the two under jaws. At
the imder jaws (maxiUce) or, when these are wanting, at the base
of the sucker, two feelers are attached, which sometimes consist of
five, sometimes only of two joints, or of a single joint. The under
lip forms an imivalye sheath excavated above, in which the sharp
threads or stings reside by means of which these insects woimd ;
on this part there are no feelers. The first piece of the thorax is a
small ring, forming a neck-ring, but the middle piece (mesothorax)
is laige, and forms nearly the whole of the thorax. To this division
of the thorax the wings are attached. These are veined, usually
transparent as glass, and colourless, occasionally spotted ; they are
without that little homy point at the anterior or external margin
{punctum caUosu/m s. cubUale) which is seen on the anterior wings
of the Hymenoptera. Behind the wings there is mostly found a
small special appendage, a membranous scale (squaToa haUerwni),
which may be regarded as a part of the wing. To the metathorax
the poisers (haUeres) are attached, which consist of a thin pedicle
and a button. These haUeres are also foimd in species that have
no wings. They are to be considered as rudiments of hind-wings^.
The abdomen is often imited to the trunk by a small part alone of
the diameter of its base, and consists of from four to nine rings.
In the female it generally runs to a point at the posterior ex-
tremity ; if here fewer rings are foimd, it is because the last of
them form an oviposUor or case consisting of rings that can be
pushed in and out of each other, like a pocket telescope. The feet
are in most long and slender, and terminate always with tarsi
^ See above, p. 253 ; comp. aUo Wbstwood, Inlrod. to modem Clastific, 11. p. 500.
Latbeille regarded these parts not as rudimentary hind-wings, for he thought they
were attached to the abdomen.
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310 CLASS YIII.
oonsistiiig of five jointa. The last joint has two claws and two or
three plane elevations or cushions (see above^ p. 252).
The digestive organs of Diptera consist of a wide bent stomach
of moderate length, a small intestine, and an oval oblong large
intestina The salivary vessels vary in the different genera of this
order. The oesophagus has a dilatation (itigliwies, crop, see above,
p. 254); it is a bladder of various form, either simple, or pro-
longed into two or more divisions, and communicating with the
oesophagus by a long narrow tube (oflen with its lowest part close
above the stomach). In larvas the tube is shorter and inserted into
the oesophagus higher up. In by &r the greater number of Diptera
this bladder is present ', in the family of the Fupiparw it is wanting
(comp. Rahdohb, AbhamUung. vh, die Verdauungawerkzeuge d, Ins.
Tab. xn. — ^xxi, and pp. 170 — 185). Tbeyirakus named this organ,
which is also found in the HymeTioptera and Lepidaptera, sucking-
bladder ; he attributed to it a power of expanding, in consequence
of which the air in the oesophagus is rarified, whilst, to fill this
partial vacuum, the fluid in which the extremity of the sucker is
planted ascends as if pumped up, {Verm, Schr. n. & 110). The
Hemiptera and the PtdicidcB do not possess this bag ; on the other
hand, such a crop is present in the Orthoptera, which do not suck,
and according to Lkon Dufour in (Edemera amongst the Coleoptera
{Ann. des Sc. not. ul 1824, p. 484, PL 30, fig. 7). The name of
food-bag, which was given by Ramdohb to this part in Diptera, is
therefore more appropriate than that of sucking-bladder. When
flies that have long fasted suck their full of milk, according to the
investigations of HuiniER, milk penetrates into this bladder. By
prcssiu:e of the abdomen, and apparently also by contraction of the
muscular walls of the bladder itself, the food is afterwards brought
back from this diverticulv/m or reservoir of nutriment to the sto-
^ According to the anatomical investigatioDS of Schb<sdeb van deb Kolk it seema
to be wanting in the larva of CEstrus; but at the same point where ordinarily the
tube from the food-bag [the name given by Ramdohr to the sac] is inserted into the
GBSophagus, two tortuous canals are seen, which divide at their other extremity into
two branches, which are distributed to the adipose body. Minunre 8ur VAnaiomie
et la Pkysiologis du Gagtnu eqtU. Amsterdam, 1845, pp. ig, 30. PI. ui. fig. i, b, s, b.
Bamdohb has figured four such appendages above and near the stomach in the larva
of Muica vomitoria, which would seem at their other extremity to be implanted into the
salivary vessels, and in the perfect insect to disappear. Tab. XIX. fig. I. M, M, M, M«
Do these vessels form, perhaps, a second apparatus for secreting saliva ?
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INSECTA. 311
mach ; (see CcUalogtie of the Physiol. Series qfComp. AnaL contamed
in the Musevm of the Roy. Coll of ^/Surgeons, L 1833, pp. 189, 190).
The Diptera live long in the larva-state, but ordinarily very
briefly as perfect insects (flies however live long in this last state).
Their larvas have no feet, but some of them have appendages which
resemble them, or small hooks, which serve for motion or holding
fast, as for instance, the larva of (Estras. All these Insects undergo
complete metamorphosis. Some larvas cast their skin before chang-
ing into pupae, and some in addition spin themselves up. Others,
on the contrary, do not cast their skin, but this shrinks, hardens,
and forms for the pupa^ that resembles an egg, a kind of shell or
case {jpupa coarct€Ua, see above, p. 273). The internal parts separate
themselves from this shell, and the change into pupa occurs within
this integument, which at last is deserted by the perfect insect
* when it breaks off the uppermost part in the form of a lid.
Many of these animals are injurious to us by their puncture ;
others suck the blood of our domestic animals ; some spoil our food
by depositing their eggs on it, especially on flesh and cheese, where
the larvae (maggots) are developed. There is, on the other hand, no
single species of this order from which we immediately derive
advantage. Yet so much the greater is the utility they afford
us indirectly. Some limit the number of injurious caterpillars,
in which they lay their eggs, and which are fed on by the pupae.
Others free the air from pestilential exhalations by feeding on
carrion and putrescent matters^.
Family IX. Pupiparce, Haustellum of three unequal setae,
exsertile from an aperture at the lower part of the head ; at the
sides of the retractile haustellimi two laminae, inarticulate, pilose,
porrect. Antennae veiy short, biarticulate, or with a single pilose
joint. Head received behind in the emarginate thorax, or re-
sembling a tubercle set upon the thorax. Feet short, strong,
remote, furnished with two incurved claws. Wings divaricate,
sometimes very short ; in some, together with the poisers, entirely
wanting. Body depressed, covered with a hard and elastic skin.
PupiparouB insects suck the blood of mammals and birds. The
buccal organs pass as fine threads through a small opening (just like
^ It is however somewhafc hyperbolical, wben Ioiidibus Bays of Muaca vomitoria :
' Trt$ muscoe contunrnnl cadaver egtd, oeque cUo acleo.'* Syat. Natur. Ed. xii. i. p. 990.
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312 CLASS VIII.
a thread through the eye of a needle, Westwood). These threads
are very long in MdophUa and Omithomi/ia, shorter in ffippobosccu
Two oblong hairy plates project like a beak and cover the base of the
threads when they are exserted. Are these parts to be r^arded as
maxUkBf or as pcUpi maxUlares ? The mechanism of the sucker is
much more conformable to that of certain AccMrina than to the
proboscis of the rest of the Diptera.
These flies lay no eggs, but are viviparous. That which seems to
be an egg laid by these insects, and which is sometimes as big as the
abdomen of the mother, ought rather to be regarded as a pupa;
from it the perfect insect (imago) comes to view after an interval of
time, dependent upon the temperature to which the pupa is ex-
posed.
The intestinal canal of these insects is very long, and surpasses
the length of the body eight or nine times. This length is caused
principally by the stomach, or that part of the intestinal canal
which precedes the insertion of the vasa urinaria, and which presents
many tortuosities. The testes are two long and very tortuous
canals; the ovaries, two oval sacs; near the oviducts are two
secretory glands, consisting of very numerous branches, together
with two more simple receptacula aeminisy of which the form varies
in different species. The lowest part of the two ovaries opens into
a wide sac {tUerus, matrice L]£oN Dupour), in which the embryo
resides until it comes forth as a pupa. The nervous system has,
besides the cerebral ganglion, only a single round ganglion in the
thorax, from the posterior margin of which the nerves of the
abdomen arise.
Gomp. on the anatomy of this figunily, L^y Dufoub, Bech. anatomigues
9ur VHippdbosque, Ann. des Sc. not. vi. 1815, pp. 299 — 322, PL 13 : ako
his Etudes anatomiquea et physioloffiques tur lea Pupiparet, Ann. des Se, noL
troisibme S^r. Zool. Tom. in. 1845, pp. 49 — 95. PI. 2, 3.
Phalanx I. Nycteribiidce. Head small, placed at the upper
part of the thorax like an obconical tubercle. Thorax semi-
orbicular. Wings and poisers none. Feet long, with first joint
of tarsus very long, and last supplied with two claws incurved, den-
tigerous at the base, and with two oval appendages.
Nycterthia Latr.
Sp. Nycterthia vespertilitmis, Acarus vespertilumis L., Phthiridium vesperti-
lionis Herm., M^m. apterol. PI. y. fig. i ; Nycteribia LatreiUii Wbstw.,
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INSECTA. 313
Latbeillx SiH, wU, d, CruM, H des Iru. Tom. xiv. PL gi, f. 14, and other
burger species on foreign bats. Aocording to L^N DuFOUB there are only
two simple eyes, Ann. dea Se, not. Tom. zzii. p. 374 ; in foreign species
two simple eyes occur on each side. NirzflCH has observed that these
insects are true pttpiparas, like Mippobosca (Sohwbioobb's Jahrbueh der
Cfhemie «. Phytii, Bd. xti. 1826, 4. s. 436). Comp. on this genus J. O.
Webtwood, Trantact. of Zoolog. 80c, I. 1835, 4to, pp. 575—294. PI. 36.
All the species of this genus liye on bats.
Phalanx 13. Sippobosctdce {Cariacea Latr.) Head received in
the emarginate thorax. Wings divaricate or incumbent, in some
very small or none. Last joint of tarsus the longest of all.
Braula NiTZSCH. Eyes and ocelli none. Wings none.
Sp. Braula coeca Nitzsoh, Gibmab Magaz, der Entom, m. pp. 3x4, 315;
Ahbbkb, Pawn, Ins. Etirop, Fasc vi. Tab. 35 ; RiAUM. M^m. y. PI. 38,
figs. X — 4 ; this small insect liyes parasitically on bees ; the metamorphosis
is unknown. Is this its place ?
Melophila NiTZSCH, Melophagua Latr. Eyes small, ocelli none.
Wings and poisers none.
Sp. MdophUa ovina, fftppobosea ovina h,, Fbibch, Beschretb. von aUerl. Im,
y. 8. 40. 4. Tab. 18; Panzeb, DeuUchl. Ina. Heft 51, 14; Gublt, M<»gax.
f. d, ffesamnU. Thierheilk. 1843, l^- ^<^1^- ^> %• 15* The theep-louse is a
wingless fly ; Rahdohb has given a description and figure of its digestive
organs ; and LTONinrr in his posthumous works a careful and eUborate pic-
ture of its structure, especially as concerns the external parts ; Recherchet
aur VAnatomie et lea mOamorph. Ac. pp. i — 27, PI. i — 3. There are two
small oblong eyes each consisting of a hundred round fa9ettes remote
from each other, (groups of simple eyes t)
Sub-genus Lipoptena Nitzsch., (Spec of Melophagu8 Latr.,
Meio.) differs from the preceding by very short rudiments of wings,
by distinct poisers, by moderate eyes.
Sp. PediciUua cervi L., Fabb., Panzeb, DeutacM. Ina, Heft 51, Tab. 15.
Orniihomyia Latr., NiTZSCH. (Spec, of Hippobosca L., and
Fabr.) Eyes distinct; ocelli usually three in vertex. Wings
distinct. Tarsi with tridentate claws.
Sub-genera: Anapera Meig. (Oxypterum Leach.) Eyes none,
wings short, acuminate.
Stenopteryx Leach, Meig. With three ocelli, wings very narrow,
longer than the abdomen.
Omithomyia Leach, Meig. With three ocelli, wings incumbent,
obtuse.
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814 CLASS Tin.
Sp. OmUhomyia MrundinU, ffippob. Mnmdinii L., BUnopi, hinmdinU
Lbaoh, Msia., Gu^BiN leonogr., Insect, PI. 104, fig. 7 : on the oommon
Swift, Cypsdus muraritu,
StreUa Wiedemann. Eyes very small, triangular. Ocelli ? Wings
inciunbent, rotundate, longer than the abdomen, with parallel
yeiiiB.
Sp. Strata vetpetiUumis Wiedem., Auaaereurop, zweifiug, Ina. u. Tab. X.
fig. 13 ; on a bat of South America.
Hippohosca Latr. Nirmomyia NiTZSCH. (Species of Hippoboaca
L.) Eyes distinct, large ; ocelli none. Wings parallel, incumbent,
obtuse, multinervose. Tarsi with bidentate claws.
Sp. Hippoboaea eq^na L., Cuv. R, Ani. id. iUuatr,, Ins, PI. 182, fig. x;
Panzeb, Deutsckl. Ins, Faac 7, Tab. 13 ; OvRUS, Moffot, /. d. geaamnU.
Thierheilk. n. Tab. i, figit. 13, 14; moucAc Bretonne, moueke d^Espagne,
Pferde-laus, Forest-fy; thorax dark-brown, spotted with yellow ; abdomen
brown-grey, hairy. This species sucks the blood of hones, and attaches
itself especially to the belly and the inside of the hind legs.
If we were told that a bird laid an egg that produced a young one as
large as the mother, we should think the account fabulous and ridiculous ;
the fabulous part would not be diminished were the bird ever so small or
even a winged insect. Of this insect however the story is accurately true.
Let the reader consult the beautiful and circumstantial natural histoiy of
this fly which R^UMUR has recorded. Mfyn. pow servw d VHist, des Ins,
VI. pp. 569—608. PI. 48.
Sub-genus Ol/ersia Wikdem. {Feronia Leach.)
Family X. Athertcera. Antennae with two joints or three, the
last undivided, and presenting the form of a patella or capitulum,
and in most supplied with a seta or spicate appendage. Proboscis
retractile or slightly prominent, with haustellum composed mostly
of two, sometimes of four seta ; in some the mouth is closed, with
tubercles in place of proboscis. Pupa coarctate.
The name Athertcera (from dSfjpy spiea, ctriata), aristeUe or owned
cmt^ncBy expresses the character by which this family of IHptera is
distinguished. The larva has in this and the following fiunily a sofl,
ringed, somewhat conical, anteriorly pointed body. Progression is
effected by extension and contraction of the body, whose form is
very variable. On the whole the larva does not change its skin, and
has no feet ; in that of HdophUua and Eristcdia alone are there
seven pairs of membranous feet provided with small hooks, on the
body beneath, which is the only example of such appendages in this
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INSECTA. 315
oider^. The moet have no proper head, but a Tery moveable mouth
veil adapted for extension, and two homy hooks curved downwards.
The skin of the larva when it changes to a pupa is not cast, but
becomes hard, and is changed into the covering of the pupa ; the
anterior extremity becomes thicker and rounder, and the whole
recalls the form of an oval keg. The perfect insect, by moving its
head, which is extended forward like a bladder, breaks this shell at
its upper extremity, whilst a piece of it springs open like a lid.
Few genera of this family are in the perfect state carnivorous,
most of them living on flowers and plants.
Phalanx I. Proboscifl in some very short, in others none, in
place of proboscis and palps three tubercles [CEstrus L.)
Larvae parasitic, some living beneath the skin, others in the
frontal sinnses, or in the intestinal canal of mammalia.
a) Proboscis small
Genera: Cephenemyia Latb., CtUer^a Clask, Latb., Trypo-
derma Wiedehanx.
b) Proboscis none.
Genera : ffypocierma Clark, {(Estrus Meio.), (Edemagena Clark,
CephcUemyia Clark, Colax Wiedem., (Estrus Clark, {GasPnia
Meio.)
Note, — ^AntennfiB triarticulate, with seta naked in most, in Gvle-
rdyra plumose. Gasl/rus Meio. differs from (Estrtia Meig. by the
naked poisers, and wings without a transverse nervure at the apex.
Comp. Clask, Obaerv. on the gentu CEsinu, Trans, of the Linn, Sac. ni.
1796, p. 2Sg, &c. ; the same, An Essay on the Bots of fforses, &c., London,
1815, 4to, with fig. ; the same, On the Insects called OiBtros by the Ancients,
Trans. Linn. Soc. xix. 2. 1843, PP« 81—94-
A. NuMAN Waameminffen omtrent de horzdmaskers, welhe in de maag
van hetpaard hvdsvesten, Amsterdam, 1834, 4to, mit pi.
J. L. C. ScHB(EDEB VAN DES KoLK, Mimoirc 9ur VAnaUmie tt la Physiol,
du Qastrus equi, Amaterdam, 1^45) &v. pL
Sp. (Estrus equi Fabb., ^Oastrus equi Mno., Gu^aiK, Iconoffr., Ins. PI. loi,
fig. 5, Glabk, Essay on the Bots, PL i. figs. 13, 14, (Liksmvb described
this species under the name of (Estrus Bovis). About 5 lines long, body
^ In these BouOH^ has frequently observed a moulting ; Beiir&ffe mr Insektenkunde,
in Nov. Act. Acad. Cos. L. Oar. Tom. xviL i. 1835, p. 498.
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316 CLASS Till.
lieurj, jfBow, tboimx in tiie middle bhck, wings with m bfown-gicj, tnam-
rene itripe in the middle and two nmilar qwts at the point ; the lemale
has a long hiaek oripoeitor at the end of the abdomen, lliis lly htys her
yellow eggs in varioas ntoations on the h^ of the hcne, to which they
remain firmly attached by a glntinoas finid. Hie yoong larw come from
the eggSy which sfiring open by a lid, as rery long and actire little worms,
and are oonreyed by the lick of the horse's tongne into hii month and
gnlkt (with respect to those eggs whidi lie beyond tfaereafdi of the tongue,
we may soppoee, with NuXAir, that the larvae themsdres creep to other
situations nearer the head). Sabsequently the lanrs Htc in the siomadi of
the horse;, to which they hare become attarhed in rery great numbers
(several hundreds at once). Here they remain several months, from spring
tiP the beginning or middle of summer, then are detached, being expelled with
the excronent^ and change into pnpe, frtan which, after about five weeks^
the perfect insect comes to view. This species is ibund in the hone and
ass ; besides these, and sometimes simultaneously with them, larve also of
other species {GaMrtu kcpwMrrkoidalu for instance) live in the ssme result ;
the larvK of this last species are smaller and deep led ; see XuXAJr, FL n.
fig. I.
(Etinu bcvis Fabb., Msig., Gu^bdi, Icono^., Jul F1 ioi, ^. 3, Clabk,
L L PL n. figs. 8, 9, Cur. R. An., Id. iU., /ju. FL 176, fig. 1. The krva
of this species lives under the sldn of the bullock ; that of Gatnu {OepMaU'
myta) oru L., GuiBiH, lam. In*. PL loi, fig. 4, Mkioev, S^. Betekr. it.
^ 3^f figy '^« Ix^es in the frontal sinuses of the sheep^
Phalanx II. Proboscis distinct. Two 8et» of haostellmn.
A, MuscaruB (species from genus Musca L.) Proboscis distinct,
membranous, retractile, bilabiate at the point.
Though the FI7 genus {JfttMa), thus defined, be much smaller
than that of LiXNiEUSy it is still a very extensive group, in which
the modems distinguish many genera. Here maj be compared
RoBiNEAU Debtoidt, Essai 8ur les Jfyodaires, Mem. prtserUes d
rAcad. desScde VInstUtU de France, Tohl n. 1830, 4ta
1 A species is spoken of in man : CEMtus kominu (Gmbl., i%ae. not. Ed. 13, L
p. 1811) ; comp. KiBBT and Sfevcb, Introdw. to Enicmol. I. pp. 136, 137. Of later
observations IsiD. Gboffbot SAurr-Hn.AniB has given an account in the Ann, de
la Soc. Eniam. n. p. 518. That larvs of CEdrui bovu and of other Diptera may
sometimes five under the skin of man is probable fix»m some observations ; a few years
ago an insect was shewn me that had come out of a boil under the skin in a girl,
it was the pupa of a dipterum, and agreed very well with that of (Estnii. A
Uu-va which in many respects resembled that of a Tackina, but yet differed from all
the species of larvae of Diftera yet known, was observed by Dr Smut ; this larva
was pressed from a boil on the head of a giri 6^ yean old ; see J. J. Smrr and
C. J. SUXDEVALL, VeterA. Akad. Mandlingar, Stockhohn, 1840, pp. 63—68.
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INSECTA. 317
*Palps external. Nervnres of wings longitudinal only, none
transverse.
Phora Latk., Meiq. (previously Trineura Meig.) Antennas
inserted at the margin of the mouth, with elongate simple seta.
Posterior feet elongate. Wings rotundate, ciliate, with two thick
nervures at the outer margin, and three or four others nearly
parallel, running obliquely from the second marginal nervure to
the posterior margin of the wing. Foisers naked.
Fig. Mbiosn, Europ, tweif. Int, vi. Tab. 63, figs, i — 13 ; GuiBiK,
Iconogr,, Tru., PL 104, figs. 3, &c.
Add sub-genera : Gymnophora Macm^- and Canicera Meig.
**Palps inserted in the proboscis, retractile and capable of
being concealed with it. Nervures of wings longitudinal and trans-
verse. Antennse inserted in the frons.
a) First posterior cellule of the wings open, transverse apical
nervure nona
* Poisers naked.
1) SquamiB of the poisers small or nona Head elongato-globose
or broad, transverae, with eyes remote.
Teianocera DuMER., Latr., Meig. Antennsd longer than the
head, with second joint longest.
Sepedon Latb., Meig., Baccha Fabb.
Thecomyia Pestt., Macq.
Loxocera Meig., Latr. Antennse oblique, with third joint
longest. Abdomen elongate, sexannulate.
Sp. Loxocera ichneuftumea, Mutca ichneumonea L,, Panzeb, DeuUehl. In$ect.,
Heft 73, Tab. 24, SoHSLLENBKBO^ Tab. 7 &c.
Sub-genus : Platystyla Macq.
Condylura Fall., Meig., Latr. Antennse shorter than head.
Abdomen sexannulate, in males clavate at the apex.
a) Seta of antenxue plumose.
Sub-genera: Lissa Meiq., Merodma Macq., Teta/nwra Valu,
Chyliaa Fall, CordyUira Macq.
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318 CLASS vin.
(/S) Seta of auteniue umple (naked or pubeocent).
Sub-genera: Gleigastra Maoq., Myopma Bobin., Maoq. (species
from genus Gcmosia Meig.)
Scatcphaga Meig., Latr., Scatomyza Fall. Antennae shorter
than head. Head barbate beneath. Abdomen quinqueannulate.
Wings incumbent, parallel, extending far beyond abdomen.
Add sub-genera: Dryomyza Fall., Sapromyza Fall., Meig.,
Toaooneura Maoq., Sdomyza FaMmj Lucina Meig., Hdamyza Fall.,
BlepluMiptera Maoq., Heteromyza Faijl
Sp. SccUophctga ttercoraria, Musca stercoraria L., CuYiES, R. Am, id. HL, Ins.
PL 178 bifl, fig. 10, RiAUMUB, Bia, not. cUs Ins, TV. PI. 17, figs. 1—7 Ac
Comp. J. W. Zbttkbbtedt, Monographia Soaicpkagarum Scandinavia,
Ann. de la Soc. EnUmol. iv. 1835, pp. 175 — 189, Tab. iv. B.
Psilcmyia Latr. {Psila Meig.)
Add sub-genera: Oxygma Meig., Trigonometopus Macq., (species
from genus Tetanocera Meig.,) Ewrina Meig., Tetanops Fall.,
Pyrgota Wiedem., OtUea Latb., Macq., Platycephala Fall., Dorycera
Meig.
Ortalia Fall., Meig.
Sub-genera: Hermay Bobin., Macq. (Eichardia Robin., and
Revdlia Robin.,) Geroxya Macq., Cleitamia Macq., Amethysa Macq.,
UTakuxtrUhina Maoq., Ropalomera Wiedeil, Eurypalpua Macq.,
PlcUystama Latb., Loxoneu/ra Maoq.
Trypeta Meig., TephrUis Latr., Fabr.
Add sub-genera: JEnsmay Aciniay Terellia and Uropkora Robin.,
PetcUophora, Senopterina and Leploxyda Macq., Bactrocera Gueb.,
Dacus Meig.
Sp. Trypeta Arctii Meig., Db Gebb, Int. vi. Tab. 1, figs. 6—14, Panzer,
Deutschl. Int., Heft 103, Tab. 22 ; yellow-green body with yellow-brown
feet ; wings with four transverse brown stripes, which at the external or
anterior margin of the wings are united two and two. The larva lives in
the flowers and seeds of Arctium Lappa and other Synanthereoe ; each peri-
carp holds only a single larva, which is placed in it head downwards.
Other species live in excrescences (like those of gall-nuts) on thistles. The
species of this genus are very numerous. The head is broad ; the abdomen
has five segments, and ends in the female in an ovipositor extended to
a point. The wings during life are mostly in a quivering motion and
erect ; they are usually spotted or striped with darker bands.
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INSECTA. 319
Sepsis Fall., Meiq. {Cephalta Meio.) Antennffi shorter than
head. Head elongate. Eyes rotund. Abdomen quadriannulate,
narrow. Wings erect, vibrating.
Sub-genera: Chelig<uter Maoq., Nemopoda Robin., Michogcuter
Macm2.
Dtopsis L. Eyes very remote, the head being produced on both
sides into a transverse petiole, oculiferons at its apex, and before the
apex antenniferons. Antennse short, with three joints, the last
Buborbicnlar with a long naked seta. Scntellum bispinose, two or
four other spines at the sides of the thorax.
Comp. A. Dahl, pneside C. Limrjn) Bigce Intectorum 1775, Amaniiatet
Acad. vm. p. 303. PL vi. figa. i — 5 (reprinted in Fuebblt, Arehivei de
VHitt. det Im, pp, 19, to, Tab. 6) ; Balhak, Aa. Holm, 18x7, AnaUct,
ErUomoL No. i. (Okih'b liu iSio) ; J. O. Wbstwood, On Dioptia, Trara.
Linn, 8oe. xym. 1835, pp. 983 — 3x3, pL See also fignres of two species
0f this genus in Gu^Biir, Iconoffr., In$, pi. 103, figs. 8, 9.
Lmnrjsus described only one species of Diopaii {DiopM, icKnewmmta),
now about twenty are known. They are all exotic and from the old worid
(west coast of Africa^ India, Java) ; JHopiia brevicomii Sat, Wibdev., a
species from Pennsylvania^ seems not to belong to this genus. According
to WxsTWOOD there are four sets in the sucker in JHopM SykesU, as in
Syrphm, These small flies by their pediculated eyes remind us of Podophthal-
mm Latb., amongst the Onutaeea, and of Zygcena, amongst the fishes.
Calohata Meiq., Micropeza Late.
Micrcpeza Fall., Calohata Latr.
Tcmypeza Fall., Tcemaptera Macq., Ferius Fabr., Wiedeil
LoTigina Wiedem. AntennsB longer than head, with first joint
longest
Thyreophora Latk., Meig.
Adora Meig.
Ccelopa Meig., {Psalidomyia Doumerc).
Ck)mp. DOUMEBO, Mim. but U Psalydomyia fucicola, diptire vivanlt tur
Ua herds de la rner, Ann, de la 80c, EnUm, n. 1833, pp. 89—93. The
male has at the abdomen a forceps ahnost like that of the Porficvlmf but
with obtuse and hairy points.
UUdia Meig., MosiUua Latb.
Gymnopoda Macq., Lipara Meig., Tvmia Meio.
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320 CLASS VIII.
Latacania Latr.
Pachycerina Macq., Lonchasa Fall., Meig., Teremyia Maoq.,
FterochrUia G&at.
Celyphvs Dalm. Antennse of the length of the head. Scutel-
lum convex, entirely covering the abdomen.
Nottphila Fall., Meig.
Ochthera Latr., Dryxo Bobik., Dickceta Meig., HydreUia Robin.,
Discocerina Macq., Trimerina Macq., Diacomyza Meig., Ccenia
Robin.
Ptophila Fall., Meig.
Trichomyza Macq., Epkydra Fall., Meig., ArUsophysa Macq.,
Ochthiphila Fall., Campichceta Macq., Oitona Meig., DroaophUa
Fall., Stega/na Meig., Diastaia Meig., Leptopezina Macq., Opomyza
Fall., Meig., {Geomyza Fall.,) Graphomyzina Macq.
Sphcerocera Latr., Borhorus Meig.
Geroptera Macq., Grumomyia Macq., ffeteroptera Macq., Limosina
Macq., Apterina Macq., (wings none).
Sp. Borboru8 pedettris Mbig. Surop. tweifi, Ins. vi. PL 63, fig. 21, two lines
long, brilliant black, winglesB; this curious insect was discovered by
V. Wnrrami near Hamburg.
Oacinis Latr., Fabr., Ckloropa Meig.
Diasevna Macq., Avlacigaster Macq., Leptomyza Macq., Lettcopis
Meig., MUichia Meig., Gymnopa Fall., Meig., SiphoneUa !&Llcq.,
Homdlv/ra Meig., CneviMcamiha Meig., Heteronefwra 'Eajll., Meig.,
Therina Meig., Meramyza Meig., Chloropa Meig., Macq., Oscinis
Latb., Macq., Lewmyza Macq., Agromyza Fall., Meig., PhyUomyza
Fall., Aateia Meig., Elachiptera Macq., Myrmemorpha Dupoub.
2) Squamffi of poisers small or moderate. Head sub-globose,
with eyes in males (sometimes in both sexes) approximate. (Abdo-
men quadriannulate). Seta of antennse inarticulate, often plumose
{ArUhomyidas),
Gcerwsia Meig. Eyes distant, especially in females. Abdomen
of male clavate at apex. Wings incumbent.
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IN8ECTA. 321
Anthomyta Meiq. Eyes in each sex, or in males, approximate;
in last more frequently contiguous. Wings divaricate or incum-
bent.
Sp. Anthomyia pluviali$, Muaca plttvialii L., Gu^Biir leonogr, Iru. FL roa,
figB. 9, Ac. A very numerous genus.
Snb-genera: Aricia Bobin., SpUogaster Macq., ffydrophoria
BoBiN., HyUmyia RoBiK., ChortophUa Macq., Atomognster Macq.,
Eriphia Meig. (with eyes contiguous in male).
Drymeia Meig. lips of proboscis elongate, infLexed, forming a
terminal hamate capitulum.
Sp. Drymeia obtewra, Mmca hamata Pall., Mbo. Europ. Zwei/L Int. ▼.
'Tab. 44, figs, xo — 13.
•• Poisers obtect
Lispe Meig. Wings incumbent. Abdomen quadriannulate,
oval. Palps dilated at apex, cochleariform. (Habitus of ArUho'
myia. Seta of antennae plumose. Eyes distant).
6) Wings with a transverse apical nervure, closing the first
posterior cellule completely or for the most part Poisers covered
with large squamse. Seta of anteniuB with two joints or three
(CreaphiUB Late.)
Abdomen mostly quadrianntdate. Wings often divaricate.
1) Seta of cmtermcR piluTnoae.
Idia Meig., Wiedem. Head produced beneath into a rostrum
above the proboscis.
Achias Fabr. Head transverse, produced on both sides into a
thick oculiferous peduncle. AntennsB short, inserted in the frons.
Sp. Aehia9 ocuUUua Fabb., S^. antUator. p. 347, GuiSbin, Mctgas. de Zod. i.
1831, Ins. PI. 7, CuviEB R. Ani, id, iU., Ins. PL 178, fig. 5, Java, and
some other smaller species, on which oomp. G. B. G. Wikdemank, Achias,
Dipterorum genus iHustratwn novisque specidnu auctum, com Tab. 1
lithogr. Kili» HolBatorum, 1830, 8vo. This genus is distinguiflhed from
Diopsis by many characters, by the position of the antenns, and by the
habitus,
Musca nob. (Spec, of genus Mvsca L.) Head globose, not
produced either anteriorly or transversely.
a) Fed moderate. Seta of a/nienno! plum^ose to the apex.
Sub-genera; Cwrtonefwra Mac(j., Meaemhrvna Meig., FoUenia
Maoq., {FoUenia Robin, and Nilellia Robin.), Mvsca Maoq., CaUi-
VOL. I. 21
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322 CLASS viir.
pfu/ra Macq., {CaXUphora Bobin., Mufetia and Melinda Bobin.),
Lucilia Maoq., (Lucilia Kobin., Fharmia and PyreUia Bobin.),
Ochromyia Macq.
h) Pett moderate. Seta of antennce naked at the apex.
Sub-genera : Onesia Robin., Cynomyia Robin., Agria Robin.,
Macq. (Agrioy Gean^rla, Clyto Robin.), Sarcophaga Meig., PhriMO-
podia Macq.
c) Fed eUmgaie.
Sub-genera: Deasia Meig., Prosena Saint-Fabgeau and Sery.,
Dinera Robin., ScoHptera Macq., RubUia Robin.
To the diyision a) the common house-fly Mtuca domettica L., belongs,
ScHBLLEN. Tab. I., J. C. Kellbb Getchichte der germeinem Stvhen-JUege,
mit 4 Kupferiafdn. Nlirnberg, 1764, 4to. The larvae live especially in
horse-dung, and these insects are only found in the neighbourhood of human
habitations. In a month's time one generation succeeds another, for the
larva is full grown in fourteen days ; the fly also after fourteen days comes
from the pupa, and the ^g has only to lie a single day before the young
maggot creeps out. Hence it is that they multiply so astonishingly, and that
in warm summers, especially at the beginning of autumn, they may prove
so numerous. In the open country and on the roads various species of fly
are met with of the same size, which are usually confounded with them, as
MuKa corvina Fabb., Panzjer Deuischl. Ins., Heft 60, Tab. 13, Mutca
nigripea male. Heft 105, Tab. 13, Mueca ludifica fem. Also the unspotted,
gold-green glistering Musca casaar L., the blue flesh-fly Mueca vomiloria,
L., (CaUipkora ftUvibarbie Robin.), Roks. Ins. Ii. Mus. et Culic. Tabs. ix.
X., Panzbr Deutscld. Ins. Heft x. Tab. 19, which has a shining blue
abdomen with black stripes, the head black with red-brown palps. This
fly has a fine sense of smell, and readily penetrates into houses in summer
to lay its eggs on meat in kitchens and larders. The same applies to those
belonging to the
Division 6) Musca camaria L., {Sarcophaga eamaria Meig.,) R^auic.
Ins. IV. Tab. 18, figs. 1, 8, Db Gebb Ins. vi. Tab. 3, figs. 5—18, Cuv.
R. Ani. id. ill., Ins. PI. 178, fig. 2 ; still larger than the former, abdomen
grey vdth black spots ; black antennae and palps. It has been estimated
that the progeny of a single female of this species may in one summer
amount to more than 500 millions of flies ^.
2) Seta o/atUenncB downy or naked Meig. (Species of Thereva
Fabb.)
^ Mbiobm Sytlt. Besch, der Eur. ZvBmfivg. Ins. v. s. si, " Hatte die Natur also
Keine nUichtigunrhenden OegenanstaUen zwr VeHUgung gebroffen so w&rde dieses
fferr den Menschen woU wenig Fleisch Hbrig lassen, und die Fastentage wOrden steis an
der T<igesordnung sein/**
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INSECTA. 323
I^asia Latr. Antennae short, the third joint a little longer
than the second. Abdomen depressed, downy, qoinqueannulate.
Wings divaricate, broad, often coloured, thickish.
Fig. Schzllhtb. Tab. n. fig. a ; Guis. Iconogr. Im. PL loi, fig. 4.
Sub-genera : Hyalomyia, Alophoray Elomyia Robin.
THchopoda Latb.
Xysta Meig. Antennae short, two last joints sub-equal. Abdo-
men convex, oval with five rings or six, pubescent or sub-nude.
Wings lanceolate, divaricate.
Oymnowma Meig. Antennae moderate, with two last joints
sub-equal. Abdomen quadriannulate, globose, sub-nude.
Sub-genera : Cistogiuter Latb., Strongygaater Maoq.
Ocyptera Latb. Antennae moderate, with third joint longer
than second, linear, compressed. Abdomen elongate, subsetose,
quadriannulate.
Fhania Meig. Antennae moderate^ third joint elongate, linear,
compressed. Abdomen oval, subsetose, quadriannulate, inflexed
towards the extremity.
Lophosia Meig. Antennae moderate, third joint the largest,
compressed, obtrigonaU Abdomen cylindrical, qnadriannulate,
subsetose*
Curtocera Macq.
Tachina Meig., Echinomyta Dumek. Antennae moderate or
short, with second joint often long, sometimes exceeding the third ;
triarticulate seta at the base of the third joint. Abdomen conical
or oval, quadriannulate, setose.
A numeroiis gentu, which beyond any other gives us an idea of the
inexhaustible riches of nature in the fbnns and modifications of aidmal
organisation. MsiosH oonnts more than three hundred species whioh are
found in Europe, and of the foreign species, yet known so fragmentarily,
still more than a hundred have been described. In some, as for instance in
Tackina grtma, the second joint of the antennie is much larger than the
last (see Ds Geeb /m. vi. F1. i. fig. 3) ; to such species Latrbille
applies exclusively the name of JSIcAtnon^ui, to which accordingly TaMna
larvamm does not belong, and which only answers to a small part of the
Tachina of Mbtobn.
21—2
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324 CLASS Tin.
Tlie hrrm of these flies live paradticany in other insects, maoy spedes in
caterpiUarSy and, with the ichnenmons, are the most serviceable in pre-
serving a balance in the economy of nature, by restraining the excessive
multiplication of noxious insects. Many are viviparous. Comp. G. Th.
Voir SiKBOLD, U^ber die veihliche Getchleehisorgane der Tachinen, Wiso-
MANi!r*8 Arekiv f, Natvrffetch. 1838, s. 191 — loi. Some larve leave the
insect in which they Uve before changring into pups.
Sp. Tachina fera, Muscaferalt., Taszkr Deultdd. Itu, Heft 70, Tab. x8, &c.
Sub-genera: Echinomyia Dumer., Latb., Maoq., Micropalpus
Macq., Thryptocera Macq., Trixa Meig., Nemorcea Robin., Magq.,
Senometopia Macq., Eurigaster Macq., Masicera Macq., Metopia
(Meig. previously) Macq., LydeUa Robin., Macq., Tachina Macq.
Ghrysosoma Macq., Clysia Robin., MycHAa Robin., Macq., {ffebia^
Mdia, Myobia Robin.), Zophomyia Macq., Cassidcemyia Macq.,
Sericocera Robin., Macq., PkUocera Robin., Melanophara (Meig.
previously) Macq.
Gonia Meig. Antennse with third joint elongate, surpassing the
two preceding taken together. Seta of anteniue triarticulate, geni-
culate.
MtUogramma Meig. Antennae with third joint elongate, sur-
passing the two preceding taken together. Seta of antennie biarti-
culate, straight. Abdomen oval or conical, in some not setose,
downy.
To this genus the observations of O. Th. Voir SiSBOLD refer (Oftserv.
gwed, entomolofficas de Oxybelo uniglum^ aique Miltogramma eonica.
Erlangae, 1841, 4to); the female of Oxyhdus tmifflumis, a species of
hymenopterum, digs for each of her ^;g8 a hole in sandy ground, and
deposits near it some flies, her booty. In this work she is watched and
foUowed by MUiogramma conica, which lurks near the entrance of the hole
for an opportunity to sUp in with her as she enters, and to fix some young
larvBB on the booty, which afterwards penetrate from it into the larva of
Oxybdtu; the reason why the Oxybdat does not drag this MtUogramma
itself into the hole as food for the larva of her egg, is readily explained by
this dipterum being viviparous, for then she would have drawn in the
Trojan horse ; and hence these Tachinaria are avoided. Lkpklstisb de
Saikt-Fasoeau has made similar observations as well on Oxj^ui as on
Cerceris; Hist. NcU, des Hymen, 184 1, 11. pp. 567, 568, 573.
B,) GonopsaruB, Proboscis exsert, in most filiform, in some
cylindrical or conical.
• Wings imperfect, very short, unfit for flying:
Camtu NiTZSCH.
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IN8EGTA* 325
Sp. Oamus kmapUra$ Nitssgh, Gibmab Magtm, der SmUmol, m. pp. 305^
307, E. F. GsRMAB Favn, Inaedor. Ewrop. Fsso. ix. Tab. 34, 35. An
animalcale about the size of a flea ; it lives pararitically od StuimuM vulgariM,
Abdomen ia broader in the female, and terminated by an exsert style.
tt Wings adapted for flying.
Stomoxys Geofpr., Fabr. Antennae shorter than head, tri-
articulate, with third joint longer than rest, compressed, with
dorsal seta. Proboscis exsert, geniculate. Abdomen oval, quadri-
annulate. Wings divaricate.
BuoenUs Latb., Siphona Meio. Seta of antenniB triarticulate,
naked or downy at the apex. Proboscis geniculate at the base and
the middle.
Tlie lanrsB of this genus live, like those of the Tachins, parasitically, in
caterpiUan; Bx Gbsb /iu. yi. pp. 38, 39; Mtmehe eoudie, PL 2, flga,
19—13.
Stomoxya Msia, Latb. Seta of antennte biarticulate, naked or
plmnosa Proboscis horizontal, geniculate at the base alone.
8p. Stotnoxys calfiUrans, Conopa caleitrans L,, GEorFBOT Jnt, ParU, n. Tah,
zvni. fig. 2, ScHELLEKBBno, PL f 7, fig. I, Gu^RiK Iconogr. Itu., PL 10 1,
fig. 8 ; grey, with abdomen spotted black. This fly fixes itself on the legs
of cattle, and in rainy weather, especially at the end of summer, enters our
houses and attacks our legs ; the uninitiated fancy that the house-flies are
then pricking, and so confound this species with Mv$ea domaHca,
Myopa Fabr. Antennae shorter than head, with second joint
longer, or second and third sub-equal, the third ovate or globose,
stjlate at the back. Proboscis exsert, geniculate. Abdomen
elongate, inflected downwards. Wings parallel, incumbent. Ocelli
three.
Zodion Latb. Proboscis geniculate at the base alona
Myapa Latb. Proboscis geniculate at the base and the middla
Add sub-genera : Stachynia Macq. {Dalnumnia Robin.), Stylo-
gctster Macq.
Conopa L., (exclusive of species). Antenna porrect, of length
of head, with first joint cylindrical, second and third forming a
fusiform club. Style of antennae apical, biarticulate. Proboscis
exsert, geniculate at the base. Abdomen elongate, sexannulate,
inflected. Wings parallel, incumbent. Ocelli none.
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326 CLASS YIII.
Sp. Conopt JIavipei h., Dum^b. Contid, g4n. «. I, In$. PI. 46, fig. 4, Pakzkb
DeuUchl, Im., Heft 73, Tabs. 21, 2^, &o.
Phalanx III. Proboscis distinct, bilabiate at apex. Four
setas of haustellum, an upper (labrum) fornicate, emarginate at the
extremity, two lateral {maxtllw) linear, supplied with a palp
incrassated towards the extremity. Sji/rphidce,
These diptera, which for the most part were placed by Linnaeus
in his genus Musca, form the genus Syrpkua Fabb., and numerous
small genera of more modem writers.
A. Antennffi of the length of the head, or longer than the head.
t Antennse with apical style.
Ceria Fabr. AntennsB inserted on a common frontal pedicle.
Abdomen cylindrical.
Callicera Meig., Latr. Antennas inserted on a common
tubercle. Abdomen conical.
tt Antennae with dorsal seta.
Psarus Latr., Fabr., Meig. Antennas iifserted on a common
frontal pedicle.
Chrysotoxum Meig., Latr. Antennas inserted on a conical
frontal tubercle. Scutellum unarmed. Wings divaricate.
sp. ChrywtoaBum arewUwm, Mutca areuata L., Qit£rik Iconogr., In$. PI. 99,
figs. 6, fto.
Ceratophf/a Wiedeil
Microdon Meig., Aphritts Latr. Antennas inserted on a small
frontal tubercle, little distinct, with first joint elongate, cylindrical.
Scutellum bidentate.
Sp. Microd. apiformu, Mtuca aptformis De Gkeb, Gu&in leonogr. Im.
PI. 100, fig. I ; Micr. mtUahilU, Musca mutabUis avctor. (in part) ; the
UiT» of this species is, according to the investigations of SomiOTHAUSKB, the
same animal which Von Sfix and Yon Heijdbn have described as a sing,
under the name of ScutdUffera and Parmula, Oken's liu, 1S40, s. 993, 923.
Paragua Latr., Meig. Antennas inserted on the frons, approxi-
mate, of the length of the head. Abdomen elongato-quadrate,
depressed. Wings parallel, incumbent.
Sp. Paroffut bicdor, Syrphui htcolor Fabk., GuiRiN, Iconogr. Im. PI. 99,
figs. 5, kc.
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INfiECTA. 327
B. Antenne shorter than head.
t Proboscis moderate,
a) Wings congraous, parallel, incambent.
Milesia Latr., Fabr. Antennas with naked dorsal seta. Nasal
tubercle none. Abdomen mostly elongate, narrow.
Eumerua nob. (Eumtrua Meig., XyhOa ejusd., Aacia ejusd.,
Sphegina ejusd., Merodon ejusd.) Hinder thighs incrassated, spinose
beneath.
8p. MiUna (Xybta Msio.) pipiens, Mutea pipiem L., Pakub DeutaeJU. Im,
Heft 32, No. 99, SCHXLLKNBSBO, Tftb. z. figs. 3, kc,
Milesia Meig. (Fipiza, Fsil&ta ejusd.) Feet simple; posterior
thighs little or not at all thicker than anterior.
Triglj/phua LoEW. (Oxen's Ins, 1840, p. 565.)
Syrphus Latr. Antennae with dorsal seta naked or downy*
Nasal tubercle.
* With abdomen eloDgate, atteniuite anterioriy, clubbed at the apez.
Sub-genus : JBaccha Meig.
* * With abdomen oval, depressed.
Sub-genera : Chrysogasier Meig., and Syrphus ejusd.
8p. Stfrphfu pyrattri, Muaea pyrattri L., DuMiaiL Ccm, gin, 9, 1. Im, PI. 50,
fig. 10 ; Syrph. ribetii, Mtuea ribeaU L., Cur. B. Ani. id, HI,, Im. PL 174,
figs. 6, fto.
This pretty fiunily of flies, mostly adorned with yellow transyerse bands,
ofken continue hovering in the air in the same place, moving their wings
with very rapid vibrations, and producing a peculiar hum in a high
note. Their larvae, conical and pointed forwards, feed on plant-lice. See
GoBDiBBDT Metamorphosis naturalis, Mediobuigi, iimo, i. p. 99, Observ, 47,
R£auhub Im, ni. PI. 30, &c.
Note, — Sub-genus PhOyekevnu Sahtt-Fabo. and Sbby., with anterior tafsn
dilated in the male. Comp. Stjcokb in Kbotsb's Nat/ufrhisi, Tidashr, vr,
1843, P- 3«i-
Sericomyia Meig. Antennas with plumed dorsal seta. Nasal
tubercle.
6) Wings congruous, divaricate or deflected.
Eelophilus Meig. (with the addition of several species of genus
Enatalta ejusd. and MaUota ejusd.) Antennsa with dorsal seta
naked or downy. Abdomen oval or conical.
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328 CLASS VIII.
Sp. HdaphUui Uncm, Musca Umax L., Pahkkb Deuiickl. Im,, Heft 14, Nos.
33, 34, SoHSLLBNBEBO, Tab. g, fig. I ; common with us towards autumn,
and often mistaken by the uninformed for bees or wasps. The larra leaves
its biding place in August and September, to undergo its metamorphosis in
chinks of walls. The body of the larva of this and of other species ends
with a long tail ; hence the French name Vera d queue de rat; they live in
cavities of stems of trees in which water is ooUected, in cesspools and
necessaries, and breathe by this tail See Goedabdt Metam. Nat. i. Observ.
7, p. 20, Tab. II. ; Swammbbdam Bijb, d. Natuur, pp. 644—646, Tab. 38,
fig. 9; BiAUHUB Int. TV. PI. 20, &c.
VoluceUa Geoffb. Syrphus Fabr. Antennae with dorsal seta
plumed, mostly long. Abdomen oval or conical, thick, hirsute.
Sp. VoluceUa intriearia, Syrphtu intricariu$ (and Syrph. bonUfffUfarmdi
Fabb.), Miuea intricaria L., Panzsb Deutichl. In$,, Heft 59, Nos. ix,
13, &c.
The larva of one species, of which the metamorphosis is known, lives in
the nests of Bombus, and feeds on its larvae ; this is VoluceUa zonaria,
SyrphuB imam* Fabb., RiAUMUB In»ect. m. pp. 483 — ^485, PI. 33, figs.
15 — 19. The body of this larva is pointed before, broad behind, with six
rays disposed in a semicircle. They are also found in wasps' nests.
Pelioocera HoFFMANNSEGG, Meig. Antennae with short dorsal
seta, incrassate, triarticulate.
c) Wings more than twice the length of the abdomen (incum-
bent, parallel).
Brachycpa HOFFMANNSEGG, Meig.
ft Proboscis elongate (of the length of head and thorax). Head
porrect beneath into a conical beak.
Rhingia ScX)P., Fabr. Wings incumbent, parallel,
Sp. Kkingia rottrata, Conopt rostraJUi L., Pakzsb Deuiachl, In$., Heft 87,
No. 29, DUM^ C<ma. ff6n, t. I. Int, PL 47, fig. 7.
Family XI. Tanystomata. Antenna mostly with three joints,
the last setigerous. Proboscis exsert, containing an haustellum
composed of four or six setae.
Longbeaks. The larvse resemble oblong worms, and have no
feet; they have ordinarily a homy and hard, others a soft head,
but which is constantly provided with booklets or retractile append-
ages, of which they make use in gnawing or sucking their food
The most live undergrotmd. They moult before changing into
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IK8ECTA. 329
pupn ; the pupe8 are naked, and shew many parts of the perfect
insect Latkeille, Cut. B. Ani, sea €d. y. p. 455.
A. Setae of haustellum four.
Phalanx I. Proboscis shorter, terminated by two large labia.
Scencpintis Latr. Antennae triarticnlate, with third joint
elongate, sub-cylindrical, truncate, no seta. Wings incumbent,
parallel.
Sp. Scenopimu feneatralit, Qviaxs Iconogr, Int. PL 96, figs. 8, &c.
A genuB of unoertain place. The larva of Scenopinut uniUt which
Bouoni described, linear, supplied with few hairs, acuminate at both ends
like the larva of TherevOf and the naked nymph (not a coarctate pupa),
sufEiciently prove the genus to differ from the division of the Mutcce.
Naiurg. der /«. 1834, Tab. iv. figs, ai, 43.
Pipunculus Latr. Antennae triarticulate, the first joint the
least, the third ovate, compressed, with erect seta at the base.
Head globose. Wings large, incumbent, parallel.
Platypeza Meiq. Antennas triarticulate, with third joint ovate,
compressed, and a terminal seta. Head globose, with eyes in male
contiguous. Wings large, parallel, incumbent. Hind feet thicker.
The larvsp live in Agarici, See the larva and pupa of Plat, holosericea
figured by Lik>N DuFOUB, Ann. dea 8c. not. ^e S^rie, zni. PI. in. figs.
14 — 76.
GdUamyia Meio.
Dolichapus Latr. Antennss with third joint triangular or ovate,
and furnished with seta. Abdomen compressed, in males incurved
at the apex. Wings incumbent, parallel. Feet long, slender.
Sp. DoUchopua nobUUaius, &c.
Sub-genera : Ammobates Stannitjs, Sybistroma Meio., Rkaphium
Meio., Porphyrops Meio., Medeterus Fisch.
Comp. H. Stankius Die Ewropditchen Arien der zweifiugler Q<Utung Dolicho-
pus, Oeen's /«>, 1831, s. 28 — 68, 8. m — 144, s. 248 — 271 ; Stjbobb
Dantke Dolichopoder, Kboteb's Tidsakr. iv. 1843, PP* i — ^44*
Sub-genus: OrtochUe Latr. ProbosciB exsert, perpendicular,
with palps acute, incumbent.
Clinocera Meig. Antennae porrect, triarticulate, with terminal
incurved seta. Wings incumbent, parallel. Tarsi with three
pulvilli.
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330 CLAB8 VIII.
Oomp. Mkioev, SiftUm. B€$ekr. der aw. nte^. Im. n. p. ii^ Tab. i6, figs.
Leptts Fabil Antennae ponect, with third joint setiferouB.
Palps exsert Wings divaricate. Tarsi furnished with three pul-
yilli. Abdomen conical, elongate.
Sp. Leptu wedopaeea, Muaea aeolopacea L., Sghellivb. Tab. 31, Fig. i,
DuMiB. Com, gin, t. /. In». PI. 48, figs, i, &c Hie hurra (BoucH^ 1. L
p. 44, Tab. lY. fig. i) lives nnder ground, and is long and conical. That
of another species from France and the Sooth of Europe, Leptu vermileo,
Musea vermileo L., Schsllihb., 1. L fig. 2, digs, like the larva of the Lion-
ant, fnnnel-shaped pits in the sand to catch the insects that fidl in. See
RiAUMDB Mim, de VAead. royaU de$ Sc. de Paris, 1753, fig. 409, PL i ;
Db Gkbb Ins. n, pp. 168—183, PL X ; Bomahd Ann. de la Soe. EnUmcl.
n. 1833, pp. 498, 499, PI. 18 0.
Sub^nera; Atherix Meig., Pdolina SriBG., Zettebst.
Tkereva Latr., Meio. {Bibto Fabr.) Palps sheltered in the
cavity of the mouth. Antennas porrect, of the length of the head,
with third joint subulate or oblongo-conical, with a small biarticu-
late terminal style. Wings divaricate. Abdomen conical, tomentose.
Sp. Tkereva plebeia L., DuM&. C<mtid, gin, 1. 1 Ins. PL 48, fig. 4 ; l%er.
anilis, Musca anilis L., Pavzbb Deutschl. Ins. Heft 5, Kos. 93, &c.
PhUocephala Zettebst.
Mydas Fabr. {Midas Wiedem.) Antenna longer than head,
quinquearticulate, clavate. Wings incumbent. (Ocellus single, at
least in some, frontal, transverse, situated between two exsert
rugee.)
Sp. Midas JUata Fabb., Duic^b. Chns. gin. s, I. Ins. PL 48, fig. 8 ; Midas
giganteus Wikdeh., Cuy. J2. Ani. id. iU, Ins. PL 173, fig. 1, bothfirom
South America. The species, with the exception of a few from Portugal,
are all exotic and very large.
Comp. Wibdbmakk, Nov. Act. Acad. Cces. Leop. Carol, xv. 2, pp. 19 —
$6, Tab. n. — iv, 183 1, Wbbtwood Arcana entomologica, i. 1841, p. 49,
PL 13, 14. Respecting the place of this genus in the natural arrangement
there are different opinions. According to the observations of Habbis,
the larva and pupa agree with those of AsHus.
Note. — Qenus Gephcdooera Latb., related to Mydas, differs from
the other genera of the family by a long, porrect; slender proboseisy
yet it ought not to be severed from Mydas.
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INSECTA. 331
Phalanx II. Proboscis exaert, tubulose. Labia at the end of
the proboscis, sometimes scarcely discernible.
t Wings incumbent.
Astlus L. Proboscis straight, porrect. Antennae porrect, approxi-
mate, triarticulate, with third joint elongate. Body elongate.
Head transverse, above much broader than long, anteriorly barbate,
posteriorly separate from the thorax by stricture.
a) Tani without pulyilli.
Qonypea Latb., Leptogaster Meiq. Abdomen elongate, narrow.
Posterior feet elongate, with clavate tibiae. Antennie with biarti-
culate pilose style at the point.
Sp. Gonypet cylindrieut, Asilm tipuUadea Fabb., Schbllenb. Tub. xzx.
fig. I.
b) Tarsi with two pul villi.
*) With apical aeta of antennae distinct.
Ommaiivs Iluo., Wiedeic. Seta of antennsB plumose.
AsUua Meio. Seta of antennse oaked^ biarticolate, sometimes
clavate.
Sp. AtiluB erabraniformia L., Schbllbnb. Tab. zxiz, figs, i, i, DdmIk.
Cent. gSn. $. I. In*. PI. 46, figs. 10, &c.
Add sab-genus : Mallophora Macq.
* *) With style of antennn conical, short.
Dcuypogon Meio.
* * *) With style of antenne short, obtuse, biarticulate. Antenna
larger than head.
Sub-genera : CercUiurgus Wiedem., Dioctria Meio.
* * • *) Without apical style of antenns.
Laphria Meio.
Add sub-genera : Rhopalogtuter^ Xiphocera and Megapoda Macq.
Hyhos Meig. {Ocydromia HoFFMANNSEGG, Meig.) Antennae
porrect, triarticulate, with the two inferior joints small, often con-
joined, scarcely distinct. Proboscis horizontal, short. Head small,
globose. Thorax oval, gibbons. Wings large, longer than the
cylindrical abdomen.
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332 CLASS VIII.
Leptopeza Maoq. (spea of Ocydromia Meig.)
On the gynonomy of oertain specieB oomp. STiBOEB in Kboteb'b Tida-
akrifi, iv. pp. 93 — 101.
(Edalea Meig.
Empis L. Proboscis exsert, perpendicular, or inflected under
the body. Last joint of antennae terminated by a seta or style.
Head small, globose.
a) With ftntennsB biuticulate (the two inferior being confluent).
TcLchydromia Meig. {Si4su8 Latb.). Anterior thigbs incrassate.
Add sub-genera : Hemerodromia Hoffmansegg, Drapetis Me-
GERLE, PUUypalpus Maoq. (spec, of Tachydromia Meig.), Xiphi-
dicera Macq., Ardoptera and Elaphropeza Macq. (sp. of Hemero-
dromia).
Cyrtoma Meig. Is tliis its place I
b) With ftntennn triarticuUte.
Empis Meig. (Empis, Pachymerina Macq.), Rhamnphomyia Meig.,
ffil<iria Meig., Brachystoma Meig., Gloma Meig., Microphorus
Macq. {Trichina Meig.)
Sp. Empi» teueUata Fabb., EmpU opaea Fabb., &c.
ft Wings divaricate.
Cyrtua Latr. Proboscis inflected xmder the body. Antennae
approximate. Poisers small, covered by large squamae. Head
small, globose. Thorax ' gibbous. Abdomen inflated, vesiculose.
Tarsi with three pulvilli.
a) Proboscis short (sometimes not diaoernible in the dried insect).
*) With antennas biarticuUte, no style.
Pterodontia Gray.
* *) With antennjB biarticulate, and terminal style.
Henaps Illig., Faer,, Ogcodes (Oncodes) Latr., {Henops Meig.,
Acrocera Meig.)
Comp. Ebiohbon Archivf. Natwrgesch, 1846, p. 188.
Sp. Henopt giJbbowm, Muaca yHboio, L., GuiaiN lamogr., Itu, PL 94,
fig. 10.
* * *) With antenns triartioul&te, no style.
AstomeUa Dufour, Latr., Ocnea Erichs., Picdea Erichs.
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INSECTA. 333
Sp. At/UmuXla eurvherUrii Dur., Atlom. marginata Latr., L^H Dufoub,
Ann. de$ Se. not. xxz. 1833, pp. 3io. 9TI, PI. 17 A, fig. i, antennA.
Habitat Spain.
h) Proboscis elongate.
*) With antennflB biarticulate ; long terminal seta.
Oyrttis Latr., Meiq., Acrocera Meig., Latr., Psilodera Griff.,
ThyUis Erichs., Philopota Wiedem.
Sp. CyHtu ffibbtu Mbio., Cyrttu aeephahts Latb., Duk^b. Comid. gin. «. {.
Jm,, pi 48, fig. 7, V1LLEB8 Entom, Linn., Tab. z. fig. 21.
* *) With antennsB triarticulate, longer than the head, no style.
ParMps Lam., Latb., Lasia Wiedem.
Sp. PaMOp$ Baudini Lamabok, Ann, du Mum. m. 1804, pp. 963—^65,
PL xzn. fig. 3, habit, in New HoUand ; Panopi oeeUtger Wibdbm., Gu^nr
Iconogr., Ins. PL 94, fig. 9.
Comp. on these genera of Diptera Ebichson Entomograpkia, 1840,
pp. i35> *«•
Bambylius L. Antennad porrect, approximate. Proboscis por-
rect, slender, mostly elongate. Palps miiarticulate. Squama of
poisers small, not covering the poisers. Trunk gibbous. Feet
slender, elongate.
a) Abdomen elongate, narrow.
Phthiria Meio., Wiedebcann {Pkthiria and Megapalpvs Macq.),
Geron Hoffmannsegg, Syatropus Wiedem., Amicttis Wiedem., Apc^
tomyza Wiedem., Thlipaomyza Wiedem., CyUenia Latb., Meig.,
Toxophora Wiedem., Xestoniyza Wiedem.
Comp. Sytiropi generis Dipierorum Monographia, audore J. O. Wbbt-
wooD, Gu^Biir Magas. de ZooL 1843 ; Sgstr. eumenoldes Wbstw., L 1.
PL 90.
Toxophora CarcelU Gu^iN, Magas. de Zod. L 1831, Jns. PL 16.
5) Abdomen short. Body hinmte.
Ploa8 Latk, Meig. Proboscis of the length of the head. First
joint of antenn® longer than the rest, very thick.
Usia Latb., Meig., Bomhylius Meig., Latr. Proboscis longer
than the head (sometimes of the length of the body). Third joint
of antennoB longer than the rest.
Comp. J. C. MiKAN Monogrophia Bombyliorum Bohemia:, iconib. illustr.
Prage, 1796, 8vo.
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334 CLASS Till.
Sp. BamhgUmB msiim» L., Bombyl. discolor, MiK., Mottoffr. Tab. n. fig. i ;
Bombyl. tricolor Gvt^ lamogr. In». PL 95, fig. 4, from BeogaL
In the proper genus Bomibffttm the body is woolly ; they hover over
flowers, whilst they suck them, like hamming bees {Bowtbt). According to
Macleat the larvss hve upon larvie of these bees ; the pape are found
under ground. Wbstwood, Introduction, u. p. 543.
Nemestrina Latr. AntennaB porrect, remote, triaxticulate, with
style elongate, setiform, terminal. Proboscis very long, at rest
inflected under the body. Thorax not gibbous. Tarsi with three
pulvilli,
Sp. Nemettrina longirottrit WiSDKiL, Atutereurop. zweifl. Im. Tab. IL fig. 5,
GuiBiN Iconogr,, In». PI. 95, fig. 7 ; from the Cape of Good Hope. In
this species, and in most of the remaining, the point of the wing is di-
vided into many cells (retiform); this is not the ease in some others,
which make up the genus FaUenia Msio.
Anthrcuc ScoPOLi, Fabr. Antennae small, triarticulate, £ur-
nished with terminal style, mostly remote. Proboscis mostly short,
sometimes retracted.
a) Tarsi with three pulviDL
4% Hirmcneura Wdedeil, Meio. Proboscifl retracted Anterior
ocelltui remote.
5) Tarsi with two pulvilli, often little distinct.
*) Proboscis longer than head.
Genera : Mtdio Latb., Corsomyza Wiedex., Enica Macq.
**) Proboscis shorty concealed or subezsert
Genera: LonuUia Meio. (previoufily Stygia ejusd), Tcmomyza
WiEDEiL with antennn approximate.
Anthrax Meio., with antennae remote.
Sp. AnSkrax morio, Mutca morio L. ; Antkr. temiatra Hoffmanhsioo,
DuMi^B. Cons, gin. t. L Ins. PL 48, fig. 4 ; Cuv. R. AnL 4d. iU., Im. ¥L
168, fig. 3. Most of the species are exotic Schjbffeb figures the larva
and pupa of a species of dipterum that lives as a parasite in the nest of the
mason-bee (Megachile). • Die Mawvrhiene, Begensburg, 1 764, 4to. Tab. ▼.
figs. II, II. Webtwood quotes these figures under Anthrax.
B. SetsB of Haustellum six (in females).
Phalanx III. (To&anu Latr.)
Tahafius L. Antennsd porrect, triarticulate, with last joint
divided into several rings, without terminal seta or style. Wings
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IN8ECTA 335
divaricate in by far the most, in some parallel, deflected. Eyes
very large, contiguous in males. Tarsi with three pulvilli.
a) Proboscis of the length of the head or shorter thui the head, bila-
biate at apex.
*) AnteniuB longer than head.
Sub-genera: ffexatama Meig., HcRVMntopata Meio. Wings de-
flected, parallel Ocelli none.
Sp. ffcenuUopota pUmaUt, TabanvM pluvialu L., Panzbb, J>e¥tdeKl. Int,
Heft 13, no. 33; four lines long, blackish, thorax with white stripes,
wings clouded grey ; the female pricks sharply, especially in warm rainy
weather ; very common all oyer Europe.
Sub-genera: Chrywpa Meig., SUviua Meio. Wings divaricate.
Ocelli three.
Sp. (Jkry9op9 ooeeutient, Taban. ecseuUmt L., DuhISb. Cons, gin. «. {. Ins. PL
47, fig. 8.
** Antenna of length of head, (ocelli none, wings divaricate).
Subgenus : Tabantu Meio.
Sp. Tahanus hovinus L., Pakzsb, DeuisckL Ins. Heft 3, no. w ; Cuv. J2.
Ani. 4d, HI., Ins. PL lyt, fig. 3, &c. This spedes has naked eyes; in
other species the eyes are haired. Gomp. Zellbb in Oksn'b Isis, 1841, s.
8x3 — S33. The larvse live under ground : the pupa state lasts in Tdbanus
bovinus about four weeks. Db Gebb Ins. VI. pp. 314 — 319, PL 13,
figs. 6, 7.
h) Proboscis elongate (of length of head and thorax), acuminate. (Ocelli
mostly three, in some none.)
Sub^nera: Pangonia Latb., Meio. (previoucdj Tanyglosaa
Meig.), Rhinomyza Wiedem.
Family XII. Notacantha s. Odontomyiidm. AntennaB with
several joints, four or more being terminal, very often joined to
form an annulate body cylindrical or conical. Set» of haustellum
four. Palps small, clavate. Tarsi with three pulvilli. Scutellum
in many armed with spines or teeth, whence the family name.
Thomrhacks, armed Jlies, Beaumur gave to some species of this
family the name of matiches cbrmes, which Geoffbot (Hist, des In-
sectes qui se Prov/oerU aux environs de Paris, 11. 1762, p. 476) ren-
dered by Straiiomys, to which name well-founded objections may be
made (ZsLLEHy OEEifB Isis, 1842, p. 828), but since it was adopted
by FABBidUS, it is now in too general use to permit its rejection
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336 CLASS VIII.
without occasioning much confusion. LaNSMUS referred these flies
to the genus Mvsccl By their antennae they approach the last
family of the diptera, the Nemocera, in which there is constantly
found a great number of joints, whilst the rest of the families have
ordinarily three alone. At the same time the majority of writers
consider the antennae of the Notacantha to be three-jointed in like
manner, in which view the last joints are merely noted as rings in
the terminal joint. But there is much that is uncertain and arbi-
trary here. That the seta of the Athericera may be counted as a
joint of the antenna, and that it is not separated by any sharp
boundary from a stylus, which is itself often jointed also, will be
readily admitted by every one who has not studied nature from
books alone. The true place of the NotacoffUha in a natural system
cannot in any case be far from Tabanus, although some only agree
with Tabani in the metamorphosis, the genus Fachystorrma for in-
stance (Latreille Genera Crust et Ins, iv. pp. 286, 287), the properly
so-named Xylophoffi, and perhaps Ccmomyia (see Westwood, IrUrod.
to modem Chissif. of Insects, ii. p. 535). Most of the species, on the
other hand, the species of all the genera which establish the essen-
tial type of this family, do not cast their skin. Under the skin of
the larva, which however does not, as occurs in AthericerOy contract
itself to a ball, the pupa is formed. Some larvse live undergroimd,
others in decayed wood, others in water.
The antennae are mostly cylindrical or conical, sometimes club-
shaped, and seldom longer than the head ; this last is a semi-round,
of which the eyes in the male occupy almost the whole bulk; there
are three ocelli. The body is flat ; the wings are long and cross
one another, lying flat on the abdomen, and mostly leaving its sides
uncovered.
A. Antennse mostly with ten joints, the last eight confluent into
a single subulate body, style none.
t Antennae not longer than head.
Ccenomyia Latb. {Stcus Fabr.) Scutellum bidentate.
Sp. Cbmomyta ferrugmea Msio., Furop, noei/L In$, n. Tab. X4 ; Vxncta
Ooru, gSn, 1. 1. Int, PL 48, fig. 3.
Xylophagus Meig. Scutellum unarmed.
Xylophagus Westw. First joint of antennae elongate.
Sp. Xylophagut tOer Mxio., Europ, zwe^. Ins, n. Tab. 17, ^, 14 ; Empis
subuUOa pAirzBB, DeiOstM, Ins. Heft 54, no. 35.
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INSECTA. 337
The name Xylophagui is, aooording to the obserystions of "Dvkwbrv,
unsuitable, for the lanre do not feed on wood, but suck those of Tiptda and
Pyroehroa, which hare the same habitat (old trunks of trees). Ksotbr'r
TUUshr, TV. p. 103.
Note, — ^Here is to be referred genus Pachyitomus Late., with five
joints of antennee, the last three conjoined. Latbeille CreTi, Grustac.
et iTisector. iv. pp. 286, 287.
SvJbvla Megerle, Webtw. (spea of Xylophagua Meiqen). First
joint of antennae short
(A genus differing from the preceding in the metamorphosis, aooording to
the observations of Roseb, Hope and others ; oomp. Wbstwood ItUroduet
n. p. 534).
Beria Latr. Scutellnm armed with four, six or eight spines.
8p. Berit clavipet Pahzbb, Deuttehl, Ins. Heft 9, no. 19, ftc.
Acanthomera Weidem.
Raphiorhyrhcifms Weedem.
(This genus with Acanthomera is placed by Maoquabt amongst the
TabanU,)
ft Antennn longer than head.
a) Antennae simple.
Cyphomyia WiED. Scutellum bidentate.
Sp. Cyphomia awnjUmma Wudbm., GufRnr Iconogr., In$. Fl. 98, fig. 5.
Habit, in Brazil. All the species are American ; the habitus is that of
Stratum^, from which genus they seem to differ by artificial character
alone.
Hermetta Latr. Last joint of antennsB oval, elongate, set upon
the constricted apex of the preceding. Scutellum unarmed.
Species all exotic, mostly American.
6) Antennae flabellate.
PHlocera Wiedemann.
Sp. PtQocera qtiodriderUata, StraHomys qttadridentata Fabb., Wisdim ank
Au$ierewrop. eweijl. Ins, n. p. 59. Tab. vm. fig. 4. Habit, in islands
Sumatra and Java.
B. Antennae with joints not more than eight, with long seta
terminal or near the apex.
8argu8 Fabr., Meig. {Sargus and Ghrysomyia Macq.) Antennae
with last joint orbicular or elliptic. Scutellum imarmed. Wings
lanceolate, longer than abdomen.
VOL. I. 22
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338 CLASS VIII.
Sp. Sarfftu evpraaitu, Mutea eupraria h., Dux^iL Cons, g4n, s, L Int.
PI. 50, fig. 8 ; with tiB not rare, four lines long ; two white spots on the
head at the base of the antenn», breast blue-green, abdomen oopper-
coloured, towards the hinder part violet, glistening prettily. — Sargvafor-
motm, Chryaomiaformosa Maoq., &c.
Chryaochlora Latb.
Sp. Sa/rgm amelhyttinut Fabs., Out. R, Ani, id. tS., Ins. PI. 173, fig. 6;
on the island Maniitiiis.
Dicrarwphora Meiq. Scutellum with very long appendage
forked at the apex.
Sp. IHcra/no]phorafvyreifera, Sargus fureifer, Wikdbx., Gutinnr Iconogr., Ins.
PI. 98, fig. 12 ; from the Brazils.
C. Antennae with six or seven joints, furnished with conical
terminal style {Nemotelus Geoffr. in part).
Foppo Latr., Fabr., Pachygaster Meig.
t Proboscis long.
Nemotehta Meig. (Spec, of genus Nemotehis Geoffr., Fabr.)
Scutellum unarmed.
Sp. Nemotdus pantherinus Msio., Panzsb Deuitschl. Ins. Heft 46, No. si,
12. {Nemot. utiginosus and margtnattis).
+t Proboscis short.
JSphijijpium Latr., CUteUaria Meig. {Ephtppiumaxid. Oyclogaater
Macq.)
Oxycera Meig. Antennae sexarticulate, cylindrical at apex,
with style terminal or dorsal, biarticulate, slender. Scutellum
bispinose.
Sp. Oxgcera trilineata Mbio., Musca panlhenna L. {sxeUu. syon.), Paitzbb
Deuisehl. Ins. Heft I. n. 13 ; 3 lines long, yeUow-green, breast with three
black longitudinal stripes, abdomen with black transverse bands on the
dorsal surfStce, feet yeUow.
D. Antennad septarticulate, of length of head or longer than
head (with first joint elongate)^ no terminal seta.
Stratiomys Geoffr., Fabr. (exclusive of species.)
Note. — ^The genus Odaniomyia Mbigen, afterwards abolished by
himself but preserved by Latbeilub, differs by the shorter antenn»,
acuminate at the apex.
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INSECTA. 339
Sp. JStraUomyt ekamadion Fabb., Mxio., ifttwa chamalBon L., Ecbsel Ivm.
II. MuKar, et Oulie. Tab. v. Pankbb DeuUchL Ins, Heft 8, No. 24, 7 lines
long ; the acutellum, the feet and under surface of the abdomen yellow, the
upper surface of the abdomen black, with three yellow transverse stripes
interrupted in the middle, and yellow point. Here are to be referred the
observations and descriptions of Swamvebdam, ffiti, not. Ins, 1669,
p. 151, Tab. IV. (under the name of Tabanus), and Bijhd der Nat, pp.
649 — 694, Tab. 39 (under the name of AsiUu). The larva is elongate,
pointed at both extremities, with a star-shaped ring of more than twenty
feathered filaments at the end ; it moves very slowly on the surface of the
water.
Family XIII. Nemocera or TipularuB. Antennae filiform or
setaceous, with numerous joints, mostly fourteen or sixteen, never
fewer than six. Head small, globose, with large eyes. Proboscis
exsert, in some short, terminated by two large labia, in some pro-
duced into a rostrum. Palps two, external, inserted at the base of
the proboscis, filiform or setaceous.
Thorax large, gibbous. Wings oblong. Poisers naked, with
inconspicuous squamae. Abdomen elongate, composed mostly of
nine joints.
Feet long, slender. Pupa incomplete {nympha).
Tkrecul-^mtermcUe, Gnat4ike* Many, especially the smaller species,
fly in great troops dancing through the air. The females lay their
^gs on the wat^, some on plants, or on the ground. The larvae are
long and yermiform ; their body has twelve rings, besides the clearly
distinct homy head The head is provided with manducating oral
organs {mcmdihlea and maaslla). The stigmata are in number and
position yarioos. Tbese larvfB constantly cast their skin before
changing to pupae. In the pupa the parts of the insect may be
clearly recognised. Almost always these pupee lie uncovered in the
water or under l^e ground; only in some are they enclosed in a
case or web (Scia/ra, MycetophUa), Many of these pupee are pro-
vided with spines or horns, by means of which, about the time of
the last changing, they are able to work to the smr&oe of the earth.
This fiunily consists of the Linnaean genera Tipida and Gvlex,
If the genus of the flies of laNTHMun, the Athericera of the modems,
with short antennae and tun-shaped pupae, be considered to be the
proper type of the two-winged insects, then the insects before us
deviate the most from that type, and make the transition to othw
orders, to aome ^europtera {Phryganea) 9Xi6.Lepidoptera {PterophoruSy
22—2
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340 CLASS VIII.
Alucita), We begin, in eniunerating the geneira, with those that
approximate most nearly to the flies.
A. Proboscis short, thick, terminated by two large labia. Setse
of haustellum in many only two. Palps with four joints, sometimes
five, mostly incurved or uniarticulate, straight. Tipula L. {Tipu-
laruB or Tipulidce of the modems).
t Antennae scarcely longer than head (or at least shorter than
head and thorax together), mostly with eleven joints, filiform, moni-
liform or perfoliate. Wings broad, rounded at the apex.
Aspistes HoFFMANNSEGO, Meig. Antennae octarticulate, clavate
at the apex. Ocelli three.
Sp. Atpistet herdinentis, Msio. Europ. ZweijL. Im. I. p. 3 19, Tab. zi.fig. 16;
one line long, on the leaves of TuuUago petasUes and the flowers of Daucug
earotta in North-Germany.
Bibio Geoffr., Meig., {Hirtea Fabr.) Antennae novemarticu-
late, perfoliate. Ocelli three. Tarsi with three pulvilli. Anterior
tibiae armed with a spine.
Sp. Bibio Ma/rci, SSpula Marci L., Ri^UMCB Ins. y. PI. ; Pakzeb, DeuisM,
Iru. Heft 95, No. 20 ; known amongst ns by the name of bladk fy ; the
lanra has ten pajrs of air-slits ; it liyes underground, and passes the winter;
the pupa lies in an oblong round cavity of loosely compacted earth ; after
three or four weeks, early in the spring (in the last half of April), the per-
fect insect makes its appearance. Gomp. Ltonbt'b observations and
figures, Recherehes, &c., Ouvrage poUhume, pp. 58 — J7, PI. 7. That these
flies cause injury to the blossom of apple-trees is a common opinion, but
entirely without proof; it is quite untrue, at least, that they lay their egga
in the blossoms. The lary», which are found in apple-blossom, are those
of a small rostrated beetle, Anthonomui pomorum ; see P. H. van Berok,
VerhaTideUng over de zwarte vliegen, Haarlem, 1807, Svo.
Dilophus Meig. Antennae undecimarticolate, perfoliate. Ocelli
three. Thorax pectinate, with a doable row of denticles.
Sp. JHlophua vulgarii, THpvlafibrUii L. ; Msiosir Europ. tweijl. Ins. i. Tab.
XI. fig. I ; VUopk. coOarit QviB,, leonogr. Im. PL 93, fig. 7, South
America, &c.
Pleda H0FFMANN8EGG, Macq.
PerUhretna Meig., Latb. Antennsa undecimarticulate, per-
foliate. Ocelli three. Palps exsert, incurved, quadriarticulate. Feet
unarmed, long.
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IN8ECTA. 341
Scatopse Geoffb., Meig., Fabr. Antennae nndecimarticulate,
perfoliate. Ocelli three. Palps veiy small, with a single joint.
Sp. Seatopse noUOa, Tiptda notaia It,, Mbtoen Eump. zvoe^, In$, i. Tab. x.
fig. 13 ; Ds GsKB InB, VL Tab. 38, figs. 1—4, &c.
Simulia Meig., Simultum Latk. Antennae undecimarticulate,
moniliform, cylindrical or fusiform. Ocelli none. Palps quadri-
articnlate.
Small, but reiy troublesome species, with oral organs developed as in the
genus OuUx (according to the observations of Curtib, dted by Westwood,
Introd. n. p. 558), but shorter. With these they prick, which the TipuUe,
on the contrary, do not. In the south of Hungary, in the Banat, Simtdia
nMcuUxta, Musca columhtiSchcMia Gmel., is sometimes, from the enormous
numbers, very troublesome, and even dangerous. Here belongs also SimtUia
periinax Kollas, Brcmliens IdsHge Inaecten, fig. 14, which, under the
collective name of Musquitos {Motquitoa, Mauttigucs), is joined to the
gnats (Oidicea). According to Humboldt, in all the Spanish colonies these
last are not called Motquitoi, but Zancudoe. In North America it seems
to be the Culices which are named Moaquitos, whilst the SimMoB are dis-
tinguished firom them as 'black flies.'
tf Antennse longer than head, mostly of the length of head and
thorax together. Joints of antemisB yarious, mostly twelve or
sixteen.
Geddomyta Meig. Antennae with 12 or more joints, filiform,
porrect. Ocelli none. Wings incumbent.
The larvae of many species live in excrescences of plants, like the gall*
wasps. Here belong Cecydomia degtruetoTf the ffeanan Fly of the North
Americans^ and Gecidomyia tritici Kibbt in Linn, Trans* iv. p. 932, v.
p. 96, Tab. 4, fig. I. By such an insect, Cecidomyia aalicina, those ex-
crescences also are caused, which are sometimes seen in the form of double
roses on the top of willow-branches. Swamhbbdamm Bijbd der Natuw,
PP- 749> 750> Tab. XLiv. fig. 16 ; Db Geeb Ins, vi. pp. 412 — a^6, PI. a6,
figs. I — 7 ; Cecid. Pint. Comp. Ratzebubo Portit. Inseden, ni. 1844. Taf.
X. fig. 14; EBiCHSON's-irc^ti;/. Naturgesch. 1841, s. 233 — 247.Tftf. xi, &c.;
L&>N Dufx>UB HiaUiire des Metamorphoses des Oecidomies, &o. Ann. des
8c. not., sec. S^r. Tom. xvi. 1841, p. 357.
Psychoda Latr., Meig. (previously Trichoptera Meig.) Antennae
porrect, moniliform, pilose, multiarticulate. Palps exsert, with
four equal joints. Ocelli none. Wings broad, pilose, famished
with many longitudinal nervures.
Sp. Psychoda phalamoides, Tipula phaltenoides L., Db Geeb Ins. vi. p. 422.
PI. 27, figs. 6—9 ; Maoquabt DypUr. i. PL 4, fig. 12. This small (ij line)
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342 CLASS VIII.
•oil-gray inseoty that reBembles a imall moth. Lb found on walls in moist
situations ; it can torn itself about retry deyerly, and springs mors than
it flies. The larva is dirty yeDow, with a black-brown head and awl-shaped
homy tail, and lires in decaying vegetable matter. BouoH^ Natwyeteh,
d, Ins. Taf. n. figs. 90| 9i ; Ptjfek. ptUu&hii Mua. Guis. leonogr. Ins^
PI 9«, f, 5, &C,
Lasiqptera Meig. {Diomyza Meqerle.)
Sub-genus: Lcuiopteryx Steph., Westw. (spea of L<uioptera
Meio.)
Zygoneura Meig.
Sub-genus : Lestremia Maoq.
Mycetophila Meig. Antennae porrect, cjlindrical, sedecim-
articulate. Palps incurved, quadriarticulate. Ocelli two or three,
unequal, the middle one the least. Tibisd spurred at the apex, the
posterior with spiny sides.
Sub-genera : Leia Mmo., BoleUna St^sgeb, SdophUa Hoffxann-
SBGO, Gnoriste "Roffmaxkb.
Gomp. H. Staknius Semerkungm tuber Hnige Arten d&r zwdjUigUr
QaUungm Macrocera, Platjfura, SciqphUa, Leia, und Mycetophila, Okxh's
leis, 1830, pp. 752—758; Staobb, KnoiTSJBt'a Tidsthift, in. 1840,
pp. «a8 — 188.
Platyura Meig. (excl. PL tipuJmdes.)
CeroplatuB Bosc, Fabr., Latr.
Gomp. B08O Actee de la Soe, (fffid, not. de Pari$, Tom. l. p. 4a, ftc. ;
L^N DuFOUR lUvinon et Monograpkie du genre Ceroplatus, Ann. det Se,
not. see. S^rie. Tom. xi. 1839, ^^' PP* *93 — *i3» ^ 5-
Gcrdyla Meig., Latr. Antennae duodecimarticulate, com-
pressed, clavate. Ocelli none.
Sciara Meig. (Molohrus Latr.)
Campyhmyza Wiedem., Meig.
Mycetdbia Meig.
Macronefwra Macq.
Asindulum Latr.
Synapha Meig.
Rhyphtis Latr.
Macrocera Meiq. Antennae long, setaceous, with the two basi-
lar joints thick, the rest indistinct. Ocelli three. Wings obtuse,
parallel, incumbent.
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INȣCTA. 343
a) Antensa? longer than body. Ocelli three, disposed in a tri-
angle, Miicrocera MExa
h) Antennae shorter than body. Ocelli throe, placed in a trans-
Terse line. BoUtophUa Hoffillnnsbgo, Msa.
On the metftmorphosiB, comp. GuiBiN, Mimaire iw un Iraeele du genre
BoUtkvphUe, Ann, dee Se. not, z. 1837^ pp. 399 — 411. PL XYin.
Chionea Dalm. AntennsB setaceous, with ten joints. Palps
with four sub-equal joints. Ocelli none. Body apterous, with
poisers.
Sp. Ckicn, artmeoides, Daulut KongL Venieiuk. Acad, ffa/ndb. 1816, loa ;
Okjen's Isie, 1894, p. 419, Tab. Y.; OuiBor Icanogr., In§, PL 93, fig. 1 ;
found in Sweden on the snow.
Ardsomera Hoffmannsegg, Meig. (HeocaJt/oma ItATR^yNematocera
Meig.) Antennas very long, setaceous, with first joint cylindrical,
second short, cup-shaped, third elongate, filiform. Palps with four
equal joints. Wings incumbent.
Megistocera WiEDEM.
Dixa Meig.
Macropeza Meig.
Trichocera Meig.
Polymera WiEDEM.
Limnobia Meig. Antennae setiform, joints 15 — 17; first joint
cylindrical, second cup-shaped, rest oblong or globose. Palps with
four equal joints. Ocelli none. Wings incumbent, parallel, with
nervures naked or pilose {Eriopiera Meig.)
Add sub-genera : Symplecta Meig., Cylindrotoma Macq. (antennae
with thirteen cylindrical joints), LimnopkUa Macq., Idioptera
Macq., Bhamphidia Meio., Macq.
Sp. Limnobia picta, TiptUa picta Fabb., Sohillenbebo Tab. 38, fig. i ;
Gu^bin Ictmogr., Ins. PL 92, fig. 9 (named Limnobia oceUaris), &c. Comp.
T. E. SCHUUICSL, Besehreibung der in Schlesitn einheinUeehen Arten von
Limnobia, BeOragezur EnUmologie I. BresUu, 1829, Svo, a. 97 — 201. Tab.
I — 5 ; B. Staknius zur Verwandlungtgetch. der Limn, Xanthoptera, ibid, 8.
v>2 — 206.
Rhipidia Meig.
Ozodtcera Macq.
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344 CLASS YIII.
Tijmla (Sp. of Tijmla L.) Antennsd filiform or setaceous, with
13 joints, the first elongato-cylindrical, second cup-shaped, short,
ihe rest cylindrical, pilose. Palps incurved, with last joint longer
than the rest, cylindrical, annulate or nodose. Ocelli none. Wings
divaricate.
Sub-genera : Fachyrhina Macq., Tipida ejusd
Ctenophara Meig. (antennas pectinate in male).
In this diyiaion are found the largest species of Nemocera, for instance,
TiptUa prcepotens Wikdem. from the island of Java, whose body is i6 lines
long, and the slightly smaller European Tipula gigantea Schrank, Schel-
LXNBEBO, IHpt. Tab. 36, CuviEB B, Ani, id, UL, Int. PL 163, fig. 5 ;
Tiptda deraeea h., De Geeb Ins. vi. PI. 18, figs. 13, 1$; Tipula crocata
L.y YiLLSBS EnUmol. Linn. Tab. ix. fig. 2, dull black, with a yellow ring
behind the head, yellow spots on the thorax, and three orange-coloured
rings on the first part of the abdomen ; wings brownish, with a black spot
at the margin ; feet dark brown. With this species is often confounded
TipvlaJlaveokUa F., Clenopkora Jlaveolata Meio., R^umue Ins. v. Tab. i.
figs. 14—16; CaviEB K Ani. id. iU., Ins. PI. 162, fig. 2, which is easily
distinguishable by its thicker and yellow feet, by seren yellow rings on the
abdomen, of which one is at the base, and by its shining black, whilst the
male, moreover, has plumed antennae. Tlie larva of this species lives in
hollow stems of trees.
DicUnidia, Xiphtwa, Bbull^ Ann. de la Soc. Entam, de France, 1. pp.
105 — 309, PI. V. Species of Ctenophora,
Pedida Late.
Nephrotama Meig.
Ptychoptera Meio. Antennae with sixteen joints, the third joint
long, cylindrical. Last joint of palps very long, setaceous. Ocelli
none. Wings divaricate, folded on the posterior margin.
Sp. Ptych. cmtaminata, Tipula contaminata L., Cuv. B. Ani, id. ill., Ins.
PI. i6a, fig. 4.
Chironomus Meig. (with addition of some genera), Fabr.
Antennae plumose. Ocelli none.
CercUopogon Meio. Antennae with thirteen joints in both sexes,
pilose; the eight inferior joints in the male barbate outwards.
Wings parallel, incumbent
The larvie live under the moist bark of dead trees. The head excepted,
every ring has on the upper part two very long hairs, with a round knob at
the extremity, not transparent, which looks like a pearl.
See Gu&iN Ann, de la Soc. EnUm. de France, n. pp. 161 — 167, PI. vm.
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INSKCTA. 345
Corethra Meig. AnteniiK with fourteen joints in both sexes,
with hairs verticillate, veiy long in male. Wings incumbent.
Sp. Corethra pkmicamis MsiQ., Corethra lateralis Latb., Pakzeb DeuUcM,
Im,, Heft 109, No. 16; Ouvdeb JZ. Ant, id. iU., Int. PL 161, fig. 4 ; the
larva lives in fresh water, has forward on the head two curved hooklets,
and is very voradous ; R^UMUB In$, v. PL 6, f. 4 — 15 ; Slabbsb Na-
tuuri. VerkuHg. Tab. in. it.; Ltokxt, Owrage post. PL 17, figs. 14,
15, 19-
Chiranomus Meio., Tanypua ejusd. Wings deflected. Anterior
feet remote from the rest, inserted almost beneath the head, very
long (at rest porrect). Antennce filiform, with thirteen or fourteen
joints in both sexes or in males alone, in females sexarticulate {Ghir
ronomvs Meio.)
Sp. Chvronomu» plumotus, Tipula plumota L., Guv. M. Ani. id. iU., Ins,
PL 161, fig. 5. The larva is a blood-red worm, often met with in rain
reservoirs ; see B^UMUB Ins. v. PL 5, figs, i — 5. On the head are two
black eye* spots, and two short antennae consisting of one joint and two
threads at the point (these are wanting in R^umub's figure). The head
is alternately drawn into and pushed out of the next following joint by the
larva. The eggs of Chironomus, oval or navicular and united in strings,
were formerly taken for plants {Diatomacea:) : Gloinema Agabdh and
Echindla; see the observations of Bebkelbt Ann. of Nat. Hist. vii. 184 1,
pp. 449 — 451. PL xm. figs. I — 8 ; comp. Koellikxb Observ. de prima
Insect, ffcnesi, 1842.
B. Proboscis porrect, of the length of thorax, or longer than
thorax, made up of seven setae. Palps quinquearticulate, porrect.
Culex L. Antennae porrect, in male plumose, in female pilose.
Wings squamate, incumbent.
jEdes HoPFMANNSBGG. Palps in both sexes very short
Sp. uEdes cinereus Hoffmakn., Cuv. R. Ani. 6d. iU., Ins. PL 161, fig. 3.
Chdex Meig. Palps of male longer than proboscis, of female
short, with first two joints very short.
Sp. Culex pipiens L., ScHELLENBEBa Tab. 41, Cuv. R. Ani. id. ill., Ins.
PL 161, fig. I ; everywhere very common, especially in the neighbourhood
of turf-diggings, as in the province of Holland. The hum or song adds to
the inconvenience. The females alone sting ; the males, known by their
plumed antennse, little or not at alL Another species, with black-spotted
wings and white-ringed feet, Culex annuUUus Fabb., has been often met
with by me here in Leyden in vdnter and in the first days of spring, in
mild weather, in dwellings.
The gnat {Cousin, Schnacke, MUcke) is commonly known. The hirvae
live in water, and hang on the surface to breathe, with head downwards.
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346 CLAflS YIII.
On Uie back, at the ninth ring of the abdomfln, then is » tabe for rmpn^
turn. Tbeae hrwm swim expeditiously, change their akin a few timea, and
become pupfe, which alao move ainaotialy, but do not eai^ and advance in
the water head npwarda, it being kept in thia position by two littie tabes
or horns that stand above the thorax and serve for respiration. On
the htfi metamorphosis the skin splits between the tnbes, and the perfect
insect creeps into view throngh the opening thns effected. It drifti for
a time npon the cast-off skin as on a little boat^ nntil the wings are strong
enough, when the gnat leaves the water. These metamorphoses occur within
a period of three or four weeks. See Swakkbbdam B^. d. Natuvr. pp.
348 — 363. Tab. zxxi. xxxn. ; R^umub /m. iy. Tab. 43, 44 ; J. M. Bab-
THn, Ik Ouliee Duaertatio, Batisbomue, 1737, 4t0f c Tab. Ac;
Anopheles Meio. Palps in both sexes of the length of proboscis.
Sp. Anophdet btfurctUui, Ctdex bifu/reoOu* L., 61161. Iconogr. Int, PL 93,
Comp. on genus OuUx Bobivbau-Dbsvoidt, JSkaot sur la Tribu dm
Oidicidet, Mim. de la 8oe. d^ffitt. not, de Paris, in. 1837. pp. 390 — ^413
(new genera Sabaikei, Pnrophora, Megarhmtu).
Order Vn. Hymenoptera.
Hexapod Insects, with four membraneons wings, the inferior less
and with fewer veins. Maxillse elongate, mostly slender, encasing
the labium. Abdomen of females almost always terminated by a
terebra or aculefas (borer, sting). Metamorphosis complete.
HyTMnoptera L., Piezata Fabb. Amongst other works the fol-
lowing treat of this order :
J. L. Christ, Natu/rgeachichie, KlaanficaUon und Ifomendatw
der Insekten vom Bienen^ Wespen und AmeisengeschleehL MU 60
ausgemaUen Kupfert, Frankf. a Main, 1791, 4to.
J. C. Fabbicii, Sy sterna Fiezatarttm. Bnmsvigae, 1804, 8ro.
G. W. F. Panzer, ErUomologischer Versuch die Jurineschen Gal-
twngen der Hymenoptem nach dem Fabriziueschen system sea priifen.
Nlimberg, 1806, 8ya (also under the title of KrUische Revision der
Insekten/auna DeutscJdands lies Bdndchen)
(The work of Jurine, NouveUe MHhade de cUisser les Hymeruh
pUres et les DipUres, av. fig. Tom. i. Gendre, 1807, T have not
been able to meet with).
C. Dahlbok, Clams novi ffymenopterorum systemaMs adjecta
synopsi Lwrva/rwn Scandinav, cructformiwn. Cum Tab, liihog.
color, Lundae, 1835, 4to.
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IN8ECTA. 847
£J)u8(L Synopmi HyfmenopUrologim SeandinOvioB, Land, 1840,
4to. (Of this the firste part alone, which treats of the genus CrabOy is
known to me).
A. Lepelbtdsr db SAnravFABGEAU, ffiitoire NaUwrMt des Insectes
EynienopUrea. Paris, 1836—1846, 8yo, av. PL, it. Vol (the last
part by Brulli^).
This order is distinguished by four naked, membranous wings.
Geoffroy united the Nev/roptera with it ; LiNN.£ns, however, had
already (in the sixth edition of the SyHema Natv/rcB, 1748) distin-
guished this order ; and to the present day it has been preserved in
systematic arrangements. The Newroptera have usually retiform
wings, with numerous small cells; in the HymenopUra they are
merely veined, and the under wings are commonly smaller than the
upper. The lower jaws are mostly elongated, and form with the
under lip a kind of proboscis by which fluids are conducted to the
oesophagus. The under jaws serve not for manducation, but for the
gnawing off matters with which these insects construct their nests,
for the bearing of burdens, <kc There are three simple eyes pre-
sent; the compound eyes are large, especially in the bee& The
foot (Uvrsud) has constantly five joints. The abdomen of the
female is almost always armed at the extremity with a sting, or
with a borer for laying egg& Already had it been justly remarked
by Aristotle*, that the two-winged insects are distinguished by a
sting in front, and the four^winged by a sting behind ; the first
wound in order to feed, the last to defend or to avenge themselves.
In some species there are wingless individuals, of which more
hereafter. The hind wings have at the anterior margin, nearly in
the middle, a row of stiff hairs or booklets (hamvli), placed at equal
distances, and only visible when magnified, by which they are
fixed fast to the posterior or inner margin of the fore wings, and in
flying lie in the same plane with these. It is especially in this
order that in the determination of the genera use may be made of
the veins and cells of the wings. Jxtrine has for this purpose
devised a terminology, of which we must give a short account. His
names have all a reference to the fore wing. The first vein of the up-
per wing, that next to the anterior or outer margin, he names radi/usy
the second, that lies more inwards, cvbUuB. These two terminate
1 Terpdrrtpa . . . WureUe^pA trrf Btwrepa d* . . . i/ATp9tr$6Karrpa. AXon, Hid, Anim,
L. I. cap. 5, med.
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348 GLASS VIII.
in a thickened point (j^wrustfum alee 8, carpus) in the middle of the
outer margin of the wing. (It is, as may be readily supposed, without
any intention of indicating an analogy with the bones of the fore-
arm in vertebrate animals, that these names of rctditu, cubitus and
carpus have been selected.) The following veins, or nervures, which
like the radius and cubUtis arise from the base of the wing, he
names nervi hrachiales. These veins form by their branching and
mutual communication certain cells on the wing {ceUvloi s. areolaf).
The outermost vein, which runs from the pv/nctum alas to the apex
of the wing, is named nervus radialis, because it seems to be a con-
tinuation of the radius ; between it and the outer margin lies the
radial cell (cellula radialis). From the cubitus there arises the
nertms^cubitalis, continued in like manner frx)m the pu/nctitm alas;
the space which lies between this vein and the radial vein is named
that of the cubital cells (cdluloB cuhitales). Finally, there arise frt)m
the brachial veins nervi recu/rrentes, or such as form communications
with each other or with the cubitus, and thus form other cells, the
humeral cells (cellula humerales^).
These insects imdei^ a perfect metamorphosis. The larve of most
species are worms without feet ; in some species, however, the larvse
have six horny feet ; still other larvae have membranous feet;
the larger number of these feet (frt)m 12 to 16) distinguishes
them frt)m the caterpillars or larvae of butterflies, which in other
respects they resemble. The food of the larvae is various ; the per-
fect insect feeds especially on the juices of plants, or swallows the
^ ^ honey of flower& Many species also attack other insects, and thus
-^ appear to live on prey ; this prey, however, does not serve for their
own nutriment, but for that of the larvae ; they are the females
that bear it to their nest The hymsfnoptera on the whole do not
live longer than a year, from the egg to the last change. Many, as
the arUs, wasps and bees, live socially together in large bodies, and
form a regulated society.
The intestinal canal of the hymenoptera begins with an oesopha-
gus, narrow and ordinarily long, which runs straight through the
thorax. In the abdominal cavity the oesophagus usually forms
an oval expansion ; only in some {CrabrOy La/rray Trypoxylan) does
this expansion form a lateral crop. The muscular stomach is little
^ Compare the article A He dea Insectea, by Audooin in JHcUonn. dassique iJCHiA.
not., Paris, i8a2, 8vo. I. pp. 176—185, and JSncyd. mitliadiqnu, Hist, not. Ins. Tom. x.
1825, p. 264, or the article RadiaUy by Lspbletier de Saint-Fabobau and Sbbvillb.
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INSECTA. 349
deyeloped, and has above four membranous valves; below it is
narrowed in shape of a funnel, and usually is introduced into the
next stomach by invagination. This stomach is cylindrical, of vari-
ous length, and ordinarily divided by transverse folds as though
into rings. The small intestine (beneath the insertion of numerous
v<i8a urinaria) is narrower than the stomach, ordinarily not longer,
sometimes even shorter than it ; the rectum is wider again. The
whole intestinal canal has no very considerable length ; in many it
is little longer than the body. In the larvse of the wasps, according
to Bamdohb, there is nothing but a large blind stomach present ;
also in the pupse of the bees there is no amis; but these have an-
terior to the stomach a narrow oesophagus, and behind the stomach
an intestinal canal terminating blindly.
The air-tubes present in most hymenopterous insects sacciform
expansions. In the bees and wasps even the lateral primary stems
in the abdomen are widened into large air-sinuses. The nervous
system exhibits different modifications in the different fiunilies.
The first nervous ganglion, the cephalic ganglion, is usually large,
since the optic nerves especially are much developed. The second
ganglion under the oesophagus lies very close to the first In the
thorax there are ordinarily only two gangKa, of which, the posterior
is large ; in Athalia centifolioB Newport found three. The abdomen
has from four to seven gangKa, ordinarily, however, only five or
This order does not contain any particularly large species, although
in the mean they are somewhat larger than the Diptera, Only a
few species are bright coloured ; the colours most frequently occur-
ring are brown, black and yellow. The species are uncommonly
numerous, so that in this respect the order of Hymenoptera is per-
haps inferior to the Goleoptera alone. Most of them indicate a very
remarkable instinct, and many construct their nests artfully. There
is one species from which man derives a great and iihmediate advan-
tage, and which he has transported with himself to different regions
of the globe. We mean the honey-bee, of whose history we shall
shortly treat in the sequel.
In their metamorphosis these insects correspond with the beetles;
in this respect they differ entirely from most of the Nefvmyptera,
Some of them by their larvse approximate to the butterflies ; and
some butterflies {Sesia) shew a great similarity with hymenopte-
rous insects. However, beyond doubt, the ffymenoptera have the
greatest affinity with the two-winged insects, and we believe that.
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350 CLASS VIII.
in a natural arrangement, thej can take no other place than in the
immediate neighbourhood of thesa By inserting the Zepidotera
between the two, aa is done by Latreille, the natural transition is
interrupted.
Section I. Aculeata, Abdomen always petiolate, in females
(and neuters) armed with a puncturing sting that conducts, in many
at least, a poison, or containing glands that secrete and ejaculate an
acrid humour. Antennae mostly with 12 joints (in femaUa) or 13
(in males), Larvas apodous.
Stmg-bearers. The wings are constantly yeined. The larva have
a quantity of food sufficient for the entire state laid near the egg
by the mother, or are provided with it daily by the sexless nurses.
The latter is the case with those which live in society.*
The sting here takes the place of the ovipositor of other insects ;
it is connected with an apparatus for the secretion of poison, which,
in the bee, consists of two long blind tortuous tubes, which coalesce
at an acute angle to form a single tube that expands into an oval
bladder. From this bladder a fluid passes into the sting, just as
from the excretory duct of the poison-gland of the viper into the
hollow tootL The sting consists of a pointed case grooved on the
ventral sun&ce, in which groove two fine apicuUs drawn to a point
are placed. At the extremity these spiculaa are provided with sharp
teeth, having their points or barbs reverted, which are less power-
fully developed in the female (amongst bees in the queen) than in
the sexless individuals (the working bees) ; also in the former the
sting is longer and turned upwards, hollow on the ventral sin*&ce.
Certain homy plates cover the base of the sting. In the males
these parts are wanting. Comp. Swakmebdam Bijbel d. NaL bL
456 — i^Q, Tab, xviil figs, n — ^iv; Reaumur Ins, v. pp. 340 — 369,
PL 29 ; KuKZMAiTN in HuFELAim's Jowmal d, FraMischen Heil-
kunde, 1820, a 119 — 127. On the sexless individuals in the order
of fff/menoptera we have treated above, p. 271.
Family XIV. MeUifera s. Anthophila. All the individuals
winged. Wings expanded. First or basilar joint of posterior tarsi
{planta Kikby) large, compressed, elongato-quadrate or triangular.
Maxillad elongate, membranous, forming with the labium the pro-
boscis.
The larvse live on the pollen and honey of flowers. Most of the
species unite for a time, or for the duration of their life, to form a
large commimity. When the society is for life, there are constantly
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INSECTA. 351
many Bexleas indiyiduals whoee buamees it is to oonstruot the nest,
and to feed the laarvie nnintemiptedly^
Phalanx I. Aptarue. Median division of ligula filiform or
setaceous, of the same length as mentum^ or longer than mentmn,
inflected downwards, with the maxillae, quite firom the insertion of
the maxillary palps. Two joints of labial palps mostly conjoined,
forming a compressed homy seta or lateral lacinia of the ligula;
two succeeding joints very small, set laterally upon the acuminate
apex of former. Two short paraglosssd at the base of ligula.
The proboscis of the bee has been investigated by Swahmebdam
(JSiJb, d, Nat, bL 445 — 451), E£iuinjR (Jns, v. GOwie Mhnaire,
pp. 304 — 326), G. R Treviiianus ( Verm. Schnften von G. R u.
L. C. TREVIRANU& IL 1817 ; Ueber die Saugwerhzeuge der Insecten,
8. 112 — 130), and other writers; and although by these researches
we are able to learn its structure and its peculiarities even to
minuteness, yet there stiU renudn obscurities and conflicting opi-
nions, especially relating to the Auction of the parts. The maxillie
form an external case {demi-etuis exterieu/ra RfAUMtm) ; the labial
palps may be regarded as a second case, if the ligula alone be consi-
dered to be the proper proboscis. Concerning this ligula, Swammebt
DAM and TBEVi&AinTS adopted the opinion that it is perforated at
the extremity by a fine aperture, and imbibes honey by its internal
cavity. According to Trevibanus a canal runs firom the base of
the proboscis to the oesophagus, though he could not trace the canal to
it. In this case bees must have two mouths, which is contrary to all
analogy. The proper mouth lies, as Reaumttb correctly observed
(whilst Swamhebdak placed it in the supposed aperture of the pro-
boscis), in the ordinary situation, behind the maxillae, and above
the ligula; it is covered above by a little valve attached to the
labrum {epipharynx or epiglossa of Savigny*). According to R£a.u-
iftTB the bees lick honey with the proboscis, just as many mammals
drink by licking with the tongue. It is probable however that honey
is sucked up by the bee, and that the proboscis at the time, like the
sucker of a pump, is moved up and down between the maxillae. See
DvoiB Phyeiologie compar^ il pp. 317, 318, and especially the
^ We possess on this division a Monograph by the Nestor of modern Entomologists,
KiBBT Moncffraphia Apum Anglice, Ipswich, i8o9, Syo. 2 rols.
* We hare noticed this little valve above at p. 381. This part, already recognised
by R^UMUBy and consid6red by him to be the tongue, was also named by Trevibavus
in bees Zunge (tongue), in wasps vordere Zwnge (anterior tongue).
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352 CLASS vnr.
Bijdrcf^ge tot de hennis der mondrdvlen van eenige Hymenopteray by
our accurate and profound Brants, Tijdschrift voor ncU. Gesch. viii.
1841, bL 71—126.
a. Social, Males^ females and neuters or workers. Maxillary
palps short, uniarticulate. Posterior tibiae in neuters dilated out-
wardly towards the extremity. First joint of tarsi tomentose
externally, or famished with brushes.
1. Posterior tibiae with spurs either none or obsolete.
Apis Fabr., nob. (spec, from genus apis L.) Mandibles with
dorsum smooth.
Comp. Latreille, Ann, du Mvs, iv. pp. 383 — ^394, PL 69, v. pp.
161 — 171, PL 13 ; De Humboldt et BoNPLAin), ReaieU cTOhservor
turns de Zoologie et d'Anai, comp, i. 1811, 4to. pp. 270 — 297. PL 19 —
21 j Des AbeUles proprem&rU dites, et plus particuH^ement des insectes
de la meme /amU-le qui sont propres d VAmeriqv/e meridionale ; par
Latreille).
Metipona Illio., Latr., Trigona Jurike. Cubital cells two.
First joint of posterior tarsi obtrigonaL
Exotic species, almost all from South America, with mandibles dentica-
lato (TrigoTia Latb.), or edentulous (Mdipona Latb.) Comp. Latbbillb
LL ; M. Spinola, Obaervationa twr Us Apiaire$ Mdoponidea, Ann. da Sc.
not. 2eB6ne, Tom. zin. 1840. Zaol, pp. 116 — 140. PL 3 ; Blavohard,
l>ict. Univ. d^Hist, natwr. vin. 1847, PP- ^5 — 89, art. MdipoMia.
Sting none, or rather rudiments alone of sting, not adapted for puncturing.
Apis Latr, Cubital cells three. First joint of posterior tarsi
elongato-quadrate, in neuters covered with hairs disposed in trans-
verse rows, and produced anteriorly into an external tooth or
auricula ^.
Sp. Apu mdUfica L., Honey^tee, AbeOledomedtque, Mowhedmid, ffcLudnene,
Jlonigbiene, Bee; Ann. du Mus. v. PI. 13, fig. i — 3 ; DuM^n. Connd. g^.
8. 1. Ins. PL 2g, fig. 4; Brakdt u. Ratzkburg, Mediz. Zool. 11. Tab. 34 ;
blackish, yet apparently of a lighter colour from greyish hair, especially
on the thorax ; a transverse, woolly, gray stripe at the base of the third and
following rings of the abdomen ; length of the body about half an inch,
breadth of outspread wings ten lines ^ workers). This spedes is domes-
ticated in Europe, and has been transported into America. All the species
of the genus Apis Latb, belong originally to the Eastern hemisphere.
In one hire there are commonly 15,000 or 10,000 working bees, 600 —
800 males, named dnmes (the ancients named them ici70^f, fif^t ^nd
1 Comp. Awnal, du Mas. iv. PL 69, fig. 5.
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IN8ECTA. 353
uMaally one female, the queen, (the king of the ancients). The working
bees are smaller than the queen, which is also distinguished by a larger
abdomen. The drones are as Urge as the queen, or lai^er, (the wings
especially are larger) ; they have no sting, and the first joint of the tarsus
of the posterior feet is neither invested with a woolly ooyeriug, nor length-
ened into a point ; the eyes are Uiger and close together.
The working bees are, as was first disoorered by ScHlRACH, nothing else
than imperfectly developed females. If the larvse of workers in the first
three days after leaving the egg receive a more abundant and more fluid
nutriment, and be transferred to the larger royal cells, there proceed from
these, according to observations, which have been often distrusted, but, as
it seems, are not deceptive, fruitfdl females or queens. The instinct of the
working bees is consequently the instinct of female animals ; they accomplish
a part of the maternal duties and take care of the larvae, the progeny of
their more highly preferred sister. Some of the working bees have the
charge of collecting food and material for building ; others, apparently
weaker, remain in the hive, care for the feeding of the larvae, and fulfil
domestic duties.
These insects live oxiginaUy in hollow stems of trees. Our domestic bees
build in hives, to which different forms have been given. When a swarm
of bees first comes into a hive, they cover it internally with an adhesive,
resinous fluid, to keep out the cold air. This substance the ancients named
propolit; the bees obtain it from the clammy buds and young leaves of
willows, ehns, &c. Next they build with the wax (which was formerly
thought to be prepared from the pollen of flowers, but is a true secretion
from the honey ^) perpendicular flat cakes or combs, beginning from abova
These cakes consist of hexagonal cells, placed horizontaDy on each side, and
opposed to each other by their tops, which are formed of three rhombs that
meet in a solid angle. Each of these cells has 5^ millimeters in mean diameter,
and, the royal cells excepted, the rest are nearly of the same size. Between
the cakes they leave spaces, which serve as passages, and in which two bees
can creep at the same time. Some oeUs contain eggs, others larvse or
pupCB, others again honey or pollen. The cell for the future queen is more
spacious, almost cylindrical; its outer surface is rough, from impressed
angular cavities, resembling imperfect cells. The number of these royal
^ As early as the middle of the last century (1 774), a german priest (Hobkbobtbl),
under the name of MiLiTTOFHiLnB Thbobebastus, published observations on the sepa-
ration of wax, which however were rescued from oblivion by Trevibandb only twenty
years ago. The observations of John Huntbb, PhU, Trane, 1793, p. 143, are better
known. The secretion of wax occurs in very thin transparent little plates on the
abdominal sur£EM» of the working bees, and is collected in the folds between the rings.
See 6. R. T&svisanus in Fb. Tiedbmann, G. R. and L. G. Tbiyibakus ZeiUchr. /.
Phfeiol. ni. 1839, 8. 63 — 71 ; comp. on the chendcal question of the production of
wax, a note in Lixbio's Organ. Cfhem, s. 307—315, from W. F. Gukdlaoh's Natwr-
geech, der Bienen, Oassel, 1843, and the observations of Dumas and Milnb Edwabds,
supported by accurate weighing, conmiunicated to the Academy of Sciences at Paris,
Arm, dee Se, not. 7e S^rie, XX. Zool. pp. 174 — x8i.
VOL. I. 23
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354 CLAjSS Yin.
oellfl ifl from two to twenty^. They usually hang like stalactites at the
margin of the honey-combe. Mach wax is bestowed on these cells, which
sometimes weigh as much as 150 common cells.
Copulation, concerning which there has been much of fahle, seems to
occur duxing flight, and the lazy drones require to be excited to it by the
queen. According to Hubsb the penis remains in the queen, and the jnale
dies in consequence ; the rest of the males also, as well as the male laryie,
are put to death at the end of the summer, and cast forth from the hive.
The larysB leare the egg after three days. After five days they prepare for
changing. They surround themselves with a fine web, on which they are,
busied ij days, and three days afterwards change into pupse. From the
pupa after seven or eight days the perfect insect comes to view. This
metamorphosis requires a shorter time in the queen, and a longer in the
drones than we have here given for the workers (Hubbb). Tke working
bees, when the perfect insects make their appearance, purify the empty
cells, that they may be ready for the reception of new ^gs. If the queen,
as occasionally happens, should lay more than one egg in a cell, they carry
out the supernumerary one. In the first b^^ning of spring (from
February to April) no other eggs are laid by the queen than those which
are to produce working bees ; the eggs of the drones are laid at a later
period (April, May), and in succession ; shortly afterwards follow queen-
bee eggs. In this way the society increases, and then sends colonies forth.
The old queen is at the head of the colony, and leaves a daughter behind
in the kingdom which she is leaving. This is called swarming. Bees
swarm several times in the summer; sometimes three or four swarms
proceed from one hive; but the last swarms are small and commonly perish.
That a hive is about to swarm, may be known by a certain noise or song,
and an unusual movement within it. The swarm leaves the hive on a
&vourable day, and gathers on the branch of a tree, on which the bees
hang like a large bunch of grapes.
Respecting the age which bees attain, there is difference of opinion.
It is probable, however, as the experiments of Hubeb shew, that they do
not form an exception to other insects in this respect, and that, however a
beehive may last five, ten, or even thirty years, there is no cause for
believing that the bees themselves reach such an age as Abibtotlb sup-
XM)sed, according to whom they may be six or seven years old. The
queen lives longer than the working bees'.
Bees have many enemies, especially amongst birds and insects ; we shall
afterwards have occasion to mention some of them. They are also exposed
to many diseases.
Amongst the numerous works on bees we must limit ourselves to noting
some. The two chief authors on Natural History amongst the ancients.
^ Occasionally, even more numerous according to some observations (R^uhub once
saw 40), but if ten be met with in a hive it must be considered to be a great nimiber.
> F. G. Desbobouoh, On the durat. of Life in the Queen, Drone, and Worker of the
Honey 'hee. Tram, of the Entomclog. Soe, of London. New Series, London, 1S53, '^*
pp. 145— 17T.
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INSECTA. 355
Aristotlx and Plikt, must be used witli caution, (Abibt. Hid. Anim. v.
21, «« ; PuNii Hitt, NeU, Lib. xi. cap. v— xx).
Amongst the moderns our Swajimbbdaic made many observations on
bees, and bestowed especially much care on tbeir anatomy, Bijbd de
Natuur, bl. 369 — 550. The chief sources for knowledge of the economy of
bees are : R^umub, Mim.pour terv, d rffiM. not, dm Int, Y. pp. 207 — 738 ;
M. A. G. ScHiBAOH, ffiM, noL de la Beine deaAbeillet, la Haye, 1771, 8yo ;
HuBBB, Ncuvdle$ ObservtUions 9ur laAheiUet, 2 vols. 8vo. Paris et Geneve,
1814.
A review of the collected observationfl of different writers was given
first by Ch. Bonmxt, Coniempl. d, I. Nai. onziifM pariU, chap. 26, 37,
CSwora, Tom. ix. (M. 8vo. Neuchatel, 1781), pp. iii — 145, and afterwards
by KiBBT and Spbhob, IrUrod. to Eatomol, u. pp. 119 — 3x4, Letten 19, 90.
Seversl works are quoted and used with deep erudition in the extensive
article on the honey-bee, contained in the excellent work of Bbakdt und
Batzxbttbo, Median. Zooloffie, u. s. 177 — 305.
2. Posterior tibiae armed with two spines at the inferior and
inner part (Cubital cells three.)
Eughssa Latk. Body smooth, shining. Proboscis elongate.
Labnun qoadrate.
AgUaljBsrEUSsr.
Comp. Eiyeycl, method., ffist. Nat. ErUomol. Tom. x. 1835, p. 105.
(These insects appear to be parasitic, and differ from BuffUma as genus
Psithyrtu does from Bombui; see below.)
Bombvs Latr. Body hirsute. Proboscis moderate. Labium
transverse.
HufMMng-btea, These insects construct their nest with mosses under
the ground. Sp. Bombui ierrulris. Apis temttri$, L., B^UX. Mim. $. I.
In$. T. Yi. PI. III. fig. I, Panzbb, DeuUchl. Int. Heft i. Tab. 16 ; black,
with a yellow ring in front on the thorax and a yellow stripe at the base of
the abdomen, of which the extremity is white. Bombut lapidariiu, Api$
lapidaria L., B£aum . Ins. T. vi. pi. i. fig. i — 4, Ohbist, Tvb. 7. fig. i^
black, the extremity of abdomen orange or reddish. The species of this
genus are numerous. They live in small societies and in inartificial dwell-
ings, which bear the same relation to the thickly inhabited artistic habita-
tions of bees, that hamlets or villages do to large towns. Amongst the
females two varieties are found, of which the smaller alone lays eggs that
produce males ; so also in Apis meUifica workers are seen, that stand half
way between common workers and the queen, and which appear to proceed
from larv89 of workers, into whose cells some of the queen's food has
casually frJlen.
Some species have no neuters, and do not live in society, but parasiti-
cally in the nest of other Bombi, They ought, therefore, according to the
strict requirements of systematic division, to be arranged with the follow-
ing. Here belongs Apis eampeslris Pahzbb, DetUschl. Ins. Heft 74, Tab. if.
They form the genus :
2»— 2
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356 CLASS VIII.
Piithyrua Lepel. de St. Fabg., Apaihua NEWMAin^.
Gomp. Chb. Dbewsbh og J. SohioDTB, ForUgndae over Danake Arter of
Slaagteme Bombui og PsUhyru$; Kb6jbr*8 Tidaakr. n. a. 1838, pp. 105—
126. Tab. n.
13. SolUarioB. Males and females alone, without neuters.
Posterior feet with tibiae not excavated outwards, with first joint of
posterior tarsi not tomentose internally.
1. First joint of poeterior tarsi produced at the external angle
of the apex or slightly prominent : second joint placed in the oppo-
site or internal angle. Posterior feet often large, very hirsute.
t Paraglossse shorter than labial palps. (Cubital cells three.)
Epicharts Klug, Latr. Maxillaiy palps very short, with
single joint.
Aecmthopus Kluo.
CerUria Fabr. (in part). Maxillary palps with four joints.
Species exotic, American. Fig. GuisBiN, looitogr, Tn». PI. 74, fig. 6.
Comp. on this genua LEPlLiTiXBy Encyd. m&hod,. Hid. Nat, Ina, Tom. x.
p. 705.
Ancyhscdia Latr. {Tetrapedia Kluo), Ftilotopua Exuo.
Sarcpoda Latr. Maxillaiy palps with five joints.
Anihophora Latiu Maxillary palps with six joints.
8p. Antkopkora hAnuta Latb., Apiaplum^ Pall., Pallas, jS^. ZooL ix.
Tab. I. fig. 14 ;— Anihophora pairidma Latb., Atmal. du Mut, m. 1804,
pp. 351 — 259. TW). xxn. ^, I, A— D.
MeUiturga Latr,
ft ParaglosBse setaceous, of the length of labial palps or longer
than these. (Cubital cells in some two, in others three.)
Eucera ScoPOLi, Fabr., Latr. (and Macrocera Spinola).
Comp. J. A. SoOFOLi Annua quartua hiatorico-natwraUa, Lipeue, 1770,
Svo. Diaaert. da Apibua, pp. 8, 9.
Sp. Eucera longieomia, Apia longicomia L., Swammbrdam Bibl. nai. Tab.
XXVI. fig. 6; Panzbb, Deutaekl. Ina., Heft 64. Tab. 11. The antemue in
the male are somewhat longer than the body.
Afdiaaodes Latb.
2. First joint of posterior tarsi nearly of the same breadth or
narrowed gradually from base to apex, with external angle little
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INSECTA. 357
or not at all produoed; seoond joint inserted into the middle of
apex of former.
t Labial palps different in form fix>m maziUary, with two large
basal joints compressed, dilated
Nomada ScOPOLi, Fabr. Mandibles small, narrow, unidentate
or edentulous. Cubital cells in some three, in others two.
Sub-genera : Oxea ELlxto, Croeisa JuiinnE, Latr., Melecta Latk,
Fasites Jvbinb, Epeclua Latb., PhilerwMu Latr., AmmohcOeB
Latr.
Sp. Nomada tuccinda Pakz., Deutichl, Im,, Heft 55, Tab. ai ; Nomada
BoberjeoUana Takzksl, ibid. Heft 7a, Tkb. 19, Ac.
Comp. Hebbioh Sohjefrb, Auaekiandenetgung der europ&i$ehen Atien
tmigtr BienenffoUtmffm, in Gbbmab'b ZeUsckr. /. die StUomol. i. 3. 1839^
8. 367 — 388 ; the same in his continuation of Panzui'b DeuUehl, Im.
Heft t66 and 176.
Megachxle Latr. {A'n;Aophcra Fabr., PhyUoUma Dumer.)
Labrum elongate, inflected, porrect below under the mandibles.
Mandibles broad, dentate or narrow, porrect, bidentate at apex.
Cubital cells mostly two.
I. With two cMud cdU,
a) With abdomen mnootk or only downy, not poOinifferouM.
Sub-genera : Gcelioan/8 Latr, Stelis Panzer, Latr.
b) With abdcmen in femalet provided beneath with long $eta forming
brutheefor eoUecting poUen.
* With abdomen oval or triangular,
ArUhidium Fabr., Latr. Maxillary palps with single joint
Comp. Latskllb, Mimoire eur U genre d'Anthidie : Annale» dn Mu-
eium, xra. 1809, pp. ^9— 53 : pp. «o7--«34, PI. i.
Sab-genera: Osmia Pakzer {Trctchusa Jurine, in part, Omnia
and Anthocopa IsESiSLEt^^ LiUw/rgug Latr., Megachile Latr. {(Jhctlir
codama Bepelet.) Maxilliary palps with two to four joints.
Sp. Megachile eeniunculariSf Apis centunetdarie L., Duicfo. Oona, gin. s. I.
Ine. PL 19, fig. 3, Gu^anr Iconogr. Jn». PL 73, i^. 7. These bees cut
jneces off rose-leaves, which they stick together and fonn into cases rolled
artfully together in form of thimbles, lliey arrange several of these cases
behind one another in a row, and lay in each of them an egg and a supply
of food for the larva. See R^umub, Int. yi. M4m. w.— Megachile mu-
rarta, Xylocopa mwraria Fabb., RiAUM. %b. Mim. in. PI. 7, 8 ; G. C.
SoosFTKB, die MemrtfUene, 1764.
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358 CLASS VIII.
* * WUk ahdomea eUmgate, ejfUmdneaL
Sub-genera : fferiades Spikola, Latb., Chelagtoma Latr.
4. WUh three evbiial edU,
CeraUna Latr. Maxilliary palps with six joints.
Sp. Ceratima aXbUabrU, ProeopU aUnlabriM Vabr., Gxrmab and Ahbeks,
Fawn. Ine. Ewnp. Fate, ▼. Tab. 17, Cuv. IL Ani. id. ill., /fu. PL 126,
fig. 5 ; in the aoath of Europe : on the economy of this insect oomp. the
obaeryations of Sfhtola, Ann. dn Mue. x. 1807, pp. 936—348.
tt Labial palps similar to maxillary.
a) Cubital cells three.
Xylocopa Latr. Labium very hard, homy, transverse, with
anterior margin ciliate. Mandibles snlcate, with point bidentate.
Abdomen depressed, broad, with sides hirsute. Feet very hirsute.
Sp. Xylocopa violacea, Apt* viclaeta L., DuM^. Cone, gin. e. I. Ine. PI. 19,
fig. I ; Cuv. R. Ani, id. iU., Ine. PL f 16, fig. 4, in southern Europe, Ac.
Many Uige native species of remarkable size, with black or violet glister-
ing wings, belong to this diYisioa, as also Xj^ooopa morio from Java^ Ac.
Sub-genus : Mesoirichia Westw.
Systropha iLLia. Antennas of males convoluted into a spire at
the apex.
h) Cubital cells two.
Panurgua Panz., Latr., Eriopa Klug. Antennas clavate or
subclavate.
Ccunptapceum Spikola
Bophitea Spinola, Latr.
Phalanx II. Andrenetas, Median portion of ligula shorter
than mentum, lanceolate or cordate. Labial palps similar to maxil-
lary, quadriarticulate. (G^nus Melitta Kirbt.)
These bees are all solitaiy, and oonaist of male and female indivi-
duals alone. The females oollect pollen, not only witlf the hind feet
but with other haired parts of their body; they dig under ground
and place by their egg a provision of pollen and honey. Others,
not fonned for gathering pollen, lay their ^;gs in the nests of other
bees.
Andrena Fabr. Median division of ligula acuminate at the
apex, lanceolate or hastate (triangular, auriculate on each side).
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INSECfTA. 359
Sub-genera: Namia HalictUie, Spheeodea, Datypodoj Andrena
JjATBL
HylfBus Latr. (not Fabr.), Proaopia Fabr., Jurine, Lepelet.
Ligula dilated at apex, sub-emarginate. Cubital cells two. Body
smooth.
Sp. EyUxm ammilatvi, Apia anmUata lu, Sphex tignaia Pavzeb DeuUehl,
I'M, Heft 53, Tab. a ; Guv. R, ami. td. ill,, Im, PL 145, fig. i &o.
Colletes Latr. Ligula emarginate, with lobes divaricate (cor-
date). Cubital cells three. Body hirsute.
Sp. OoOsUsfodieni, Panzib, DeuUchl. Jna, Heft 105, Tab. ii, ii, ftc^
Family XV. Diplopteryga. All the individuals winged.
Anterior wings longitudinally duplicate, the insect being at rest.
First joint of posterior tarsi not dilated. Eyes emarginate (reni-
form). Antennae filiform or thicker towards the apex. Prothorax
produced laterally as far as the the origin of the posterior wings.
Body smooth. Females and neuters armed with a strong poisonous
sting.
Waaps, The inner edge of the upper wings is turned down bo
that their upper surface lies on the under wings'; hence the name
IHploptera (donhle-winged), which Kibbt, because the termination
ptera denotes the names of orders of insects, changed into Dip-
lopteryga (J. Bichabdson, Fcvuna Bareali-Amenocma, 17. 1837,
p. 263).
The wasps are mostly coloured yellow or red and black. The
pupsB are vermiform, and without feet. They are inclosed in separ
rate cells, where they find food placed by the mother with the egg
she has laid, or are fed daily by the mother or by the sterile
working-waspa
Phalanx I. Antenn» with only eight distinct joints, the rest
conjoined to form an obtuse or rounded club. Ligula with two
terminal laciniae, received in the tubular base. Cubital cells only
two complete.
1 Comp. Eniomologiea, anctore J. Imhoff, Okin's liii, 183s. s. 1198 — laoS; de-
Bcriptions of speoiM of the genera OoUetet, HyUew and Andrena^ with remarks on the
synonymy.
* The genoe Ceramiua eeems to be the only exception to thiB, which however is
denied by LsFiLiTiKBy Hiit. not, da HjfnUnopUrrs, n. p. 590.
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360 CLASS vni.
Mamria Fabr.
Masaria Latr. Antennss (of males) longer than head. Abdomen
elongata
Celonitea Latb. Antenns in both sexes very short, terminated
by a globose clab. Abdomen scarcely longer than thorax.
Sp. OeUmitea a^formis, Pamzib, BetdsM. /«. Heft 76, Tab. 19 ; Dum^bil,
CwM. gen, 9. 1, Int. PL 31, fig. 9 ; Cuv. R. ani. M, ill., Int. PI. IS3, fig. 9;
in Bouthem Europe.
Phalanx II. Antennae with all the joints distinct, 13 in males,
12 in females, with second joint geniculate, thicker from third
joint towards apex, acuminate at apex.
a) SclUa/ry^. Mandibiles elongated-triangular, connivent, like
a rostelluuL ligula narrow, elongate. Clypeus sub-oordiform or
oyal, with anterior margin produced and truncated.
t Cubital cells only two.
Ceramiua Latb., Klug. Labial palps longer than maxillary.
Sp, CbumtM Fonaoohmbii Latb., Gu£bin, Iconogr, Int. PL 73, fig. 4.
ft Cubital cells three.
Sjfnagris Latr., Fabr, Ligula produced into four very long
sub-plumose setae. Maxillary palps short, with four joints. Man-
dibles of males very large, porrect.
Sp. Synagrit caUda, Vetpa calida L., Gu^iir, leonogr. Int. PL 73, fig. 3 ; —
Synagr. camuta, Vetpa eomuta L., Bneyd. mkk., Int. PL 383, fig. la
Ck>xQp. on this genus LsPELETiKB in Encgcl. mtth. Int. Tom. X. pp. 509, 510.
Eumenes Latr,, Fabr. Ligula tripartite, with middle part
deeply incised, bifid. Four glandular points at the apices of ligula,
Maxillary palps with six joints.
a) With abdomen oval or conical, thicher at the hate.
Sub-genera : Pterochiku Klug (with labial palps plumose), Odf^-
nenis Latr. {Ehynekium Spikola).
Sp. Odynerut auctut, Vetpa aucta Fabb., Panzbb, Deuttehl. Int., Heft 81,
Tab. 17 ; Odyn. tpinipit, (fern. Od. murariut), Panzbb, Deuttehl. Int.
Heft 17, Tab. 18. (Here belong, according to Auoouur, the observations
of BiAUMUB, Int. VL pp. as I— 268. PI. xxvi. figa. 1— 10.)
^ H. F. DE Saussubb, Monographic det Guipet toliUUret, av. 11 pi. color. Paris et
Geneve, 185a, Svo.
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INSECTA. 361
Gomp. on this genua Wbstmakl, Monogr. det OdyrUra de la Bdgiqiie,
BnixelleB, 1833, 8yo, {Ann, des 8c, mU, xzx. 1833, PP* 4^^^ — 43^) ; Hkb-
BiCH-ScHJEFFBB, DeuUckl. TfU. Heft 173, 176 ; L^n-Dufoub, Mim, pour
mrvir d VHUt, de Vinduitrie el det vUtamorpkoaea det OdytUret, Ann, det,
Sc. not, le S^rie, Tom. zi. 1809. Zool, pp. 85—103. (Odynerut rvhicola) ;
AuDOUiF, Obtervationt tur let motwrt det Odfniret, ib. pp. 104 — 1 13.
b) WUkfrtt twojokUt 0/ abdomen coarctated to form a petiole, ihefirU narrow
cb-eonieal, the teoond tub-^ampantdaie.
Sub-genus : Eumenes Latb. (Zethtis Fabr., DUccdiuB Latb.)
Sp. Eumenet eoaretata, Vetpa coronata Panz., Gioffb. Int, n. PI. itl
fig. 7; Panzxb, Deuttchl, Int,, Heft 64, Tab. 116, and Vetpa coarctaia
Panz. ibid, 6$, Tab. 6; Vetpa pomtfomUt, Pamz. ib. 63, Tab. 7, fern. Ac.
b) Social. Mandibles subquadrate, obliquely truncated and
denticulate at apex. Lingula little elongate, tripartite, with middle
part bifid. Four glandular points at apices of lingula. Labial
palps quadriarticulate, maxillary sexarticulate. Cubital cells three.
Vespa (Species firom genus Vespa L., Vespay PoUatea Latr.)
In wasps the upper lip {lahrunC) is hidden behind the head-shield
(dypeua) and upper jaws (hvntere Zu/nge, Tbevirakus, V&rm. Schr.
IL Tab. XY. figs. 7, 8, 9, L'). Close under the upper-lip is found a
valve, which Saviony names epiphari/nx or epiglossa (vordere
Zunge Tbeviranits). The hypapharynx of Saviony {Klappe Tbb-
vmANUS, L L 8. 134, Tab. xv. figs. 6 — 8, letter v.) is a valve closing
a cavity which was discovered by BaAirrs, and named gluteT^-ccmty
(see his work cited above, p 452, Tijdschr. voor, NaJL Oesek, vm. bL
97) ; in this cavity, not unlike the buccal sacs of certain mAimrn^^^i^^
the wasps keep small pieces of wood for building their nests \ The
wasp's nest consists of a kind of paper, prepared firom small pieces
of old wood and bark of trees ; they gnaw off the pieces with their
upper jaws ; the cakes are usually horizontal ; the uppermost are
formed first and afterwards the lower ; they hang together by little
pedicles, and the uppermost of all is fixed to the common covering;
for most wasps form a common covering for their nests, whilst bees
merely hide their waxen dwellings in hollow trees or under the
groxmd. A wasp*s nest either hangs fireely in the air, or is formed
under ground or in cavities of trees. The cells are hexangular,
perpendicular, and generally with the opening facing downwards.
1 On the undar-lip (UgtUa) and the other parts of the mouth in wasps, oomp. the Bg,
in R^UMUB, Mfyn. 1. 1. Int, vi. PI. 16, fig. 2.
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362 CLASS VIII.
In bees we saw a monarchy with a queen at the head ; here the fonn of
government is a republic, the members of which are supported less by their
own industry than by rapine. Wasps are freebooters ; they are very eager
for the juice of fruits; they suck the fluid that adheres to meat in
slaughter-houses, cut pieces off, rob bees of their honey and murder them^
as well as other insects, not for their own use, but to feed their Uutsb with
them. This robber-state, however numerous its dtizens may be, owes its
origin to a single mother. She is fertilised in the autumn, and lives over
the winter, whilst the neuters and nudes die, and in the spring conmiences
the work alone. After a while she is assisted by sexless wasps which are
her first-born children. In autumn males and females are bom. At that
time some hundreds of the last are often found in a single nest, dwelling
in uninterrupted peace, whilst amongst bees only two or three females
are able to be of one mind together, for a short time. The working wasps
are smaller than the rest ; they all die from the cold of winter.
Gomp. B^UHUB, MSm. 8. 1, Ina, vi. M6m, vi. vn ; Boknkt, ConUmpla'
Hon de la Nature, XI. partie, chaps. 13 — 35 ; (Euvres cPJffitt. Nat, et de
Philos. Tom. IX. 8vo, pp. 99 — 100 ; Kibbt and Bfbkoe, IrUrod. to BrUo-
mol. n. pp. 107 — III.
Polistes Latr. Clypeus f)roduced anteriorly into a sharp tooth.
Abdomen in some adhering to the thorax by a long petiole.
Sp. Veapa nidnlans Fabb., Epipone chartaria Latb., ffiM. not. dea Onut. et
de& Jna, xin. Tab. 102, £ 6 ; Gu^bin, leonogr. Int. PI. 7a, fig. 7. This
South- American species makes very large nests, as though of pasteboard,
hung to a branch of a tree, Uke long sacks with a conical lower end, with
an opening in the middle. The cells are attached to different transverse
partitions, which are perforated in the middle ; this is the Ou^pe carionniire
of RliUMUB, InSi VI. M^. vn. p. 334, &c PI. 20 — 34, &c. To this division
also belongs the honey-gathering wasp of the Bracils, named XeeA^^uona ;
see Latbxillb, M^, du Mtuhtm, xi. pp. 313 — 320, and another spedes
which A. White names Myropetra sctUellaris, whose nest differs from that
of Vespa nidtdana by the conical knobs with which it is beset externally.
Ann. of not. Hitt. vn. 1841, pp. 315 — 312.
To the division Poliatea belong some European wasps whose nest has no
common covering, the cells lying bare. Swammebdam, Bybd der NaL
Tab. 16, fig. 15 ; Bobbbl, Im, n. Boinb. et Vesp. Tab. vn.
Veapa Latb. Clypeus truncated anterioriy^ emarginate.
Sp. Veapa crabro L., RiAUMUB, Ina. Yi. PL xvm.^ — Veapa vtdgaria L.,
R£&UM. ibid. PI. xiv. figs, r— 7, Panzeb, DeuUchl. Ina., Heft 49, Tab.
19, &c.
^ That this insect, the largest and most voracious wasp of Europe, may be to some
extent tamed, and then is not to be feared, appears from the observations of P. W. J.
MuELLEB ; see his amusingly written paper in Gebmab und hlJXCKBSf, Magadn
der Entomohgie, ni. 1818, s. 56 — 68.
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INSECTA. 363
Family XYI. Heterogyna. Females (amongst the solitary) or
neuters (amongst the social) apterous; males less than females.
Wings not plicate, with cells often few, incomplete. First joint of
posterior tarsi not dilated. Females and neuters in some furnished
with a sting, in others with anal glands that secrete a peculiar
acid. Ligula small, membranous, round, excavated or hooded.
AntennsB geniculate.
Phalanx I. Bodalia or Formicarim. Males females and neuters,
or abortive females. Neuters apterous, without ocelli, with head
very large, and labium large, descending under the mandibles.
Mandibles strong, often denticulate. AntennsB filiform or subin-
crassated towards the apex, with first joint very long, cylindrical
or obconical. Petiole of abdomen formed of one or two globose
nodes.
Formica L.
AfU9 {/(mrmia, Ameisen, mieren). The females have wings that
easily fall off, or are stript off by themselves after copulation. The
sexless individuals on the oofitniry are without wings, and without
simple eyes {oceUi) also ; they are, like the working bees, imperfect
females, as appears also from the observation of Hubeb, who fre-
quently saw males copulate with them, but the act always caused
the death of the neuters. The males and females are Ibund as per-
fect insects in the nests for a short time only, for they desert them
as soon as they have gotten their wings. The males are smaller
than the females, and have also a smaller head and smaller upper
jaws, but larger eyes. These ammals live together socially, construct
for themselves nests of earth, leaves, straw ^, <&c., in which other insects
and their larvie often reside, as those of Cetonia and other CoUoptera^
especially Brctchdytra, which in these last times has given occasion
to much inquiry among entomologists'. Above the place where
they dwell the ants raise small hillocks or round heaps of earth.
Other species Uve in hollow stems of trees. Along straight roads
^ Henoe there exists a species of vegetable manure and a high temperature in an
ant's nest> which continues even after it has been deserted by its inhabitants. BOBSBT,
Ann. da Se. not, sec. S^rie, XYm. Zool. 15S.
■ On insects living in ants* nests, see amongst others : MANmSHsnr, BulUl, de la
Soc, imp, de Moacou, rvi. 1843, pp. 70—78, Mmkjav, ibid. xix. 1846, pp. iS7— «87>
and espedally Fb. Mabrksl in Gbbmab'b ZeiUehr.f. d. BiU^moL m. 184 1, pp. 20—
22S,ibid, V. 1844, »• ^^93— »7i-
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364 CLASS VIII.
over the ground, often a hundred feet long, which all end like rays
at the dwelling, the ants pass to and fro ; irregular and tortuous
passages lead to the separate habitation of the future generation.
All the labour of building, of nursing and feeding the larvse <kc. is
discharged by the neuters. They live on fruits, insects and their
larvs, on dead birds and small Tnammala. They are very fond of
sugar, and follow the plant-lice in order to swallow the sweet sap (the
honey-dew) that drops from their body. They lay up no provision
for winter, as &r at least as relates to our native species, but pass
the winter in a state of torpor, taking no food at all in the severest
cold. The working ants bear the larv» and nymphs with the great-
est care between their jaws to the surface when the sun shines on
their dwelling, and down again when rain falls on the earth, and
they defend with iucredible courage the commonwealth which has
no other government but a true republic. The larvse and pupae are
commonly taken for eggs by the uninformed, and serve for food for
certain singing birds in cages : nightingales especially are fond of
them. In the last days of summer (August), in warm clear weather,
the winged males and female leave the nest in which they have
been brought up, fly in swanns through the air, copulate, and die
soon afterwards, being swallowed by birds, or drowned in water
and made food for fishes. The females that are left divest them-
selves with their feet of the wings that are now useless, and found
a new colony ; working ants, in whose neighbourhood they chance
to be, drag them to their nest to lay their eggs there; when that is
accomplished they are driven without mercy from the nest
Gomp. on Ants:
QwAXMXKDAiifJB^bd der not. bL 187 — 299; Ge. di Gxsb,/9w. xvmi^me
Jlitm. n. pp. 1042^-1 107 ; Boknbt, Ooniempl. de la Nature, Partie xi.
chap. 27, (Euvr, compl. %yo, ix. pp. 89 — 98 ; EIibbt and Sfknok, Inhrod.
to Entom, I. pp. 479 — ^484 ; n. pp. 45 — 106 ; Okek, AUffem, Natuiiyetek,
Vol 2, 1835. pp. 895—945.
Latbkills, ffui. nat, dea Fowrmu, i Vol. 8vo, av. fig. Paris, t8oi.
P. HuBSR, Bechtrchet mr let nusun de» FcwnM» indiffineB, i Vol. 8yo,
av. fig. Paris et Geneve, t8io.
Lund, Sur Us habiludea de qudqun Fmurmii du BretU. Awn, da 8e, nat.
xxm. 1831, pp. 113— 138.
A. Petiole of abdomen composed of two distinct nodes. Females
and neuters frimished with sting.
Myrmica Latr. (with addition of other genera).
Sub-genus : Atta Fabb., Latr., Maxillary palps short, with five
joints or fewer.
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INSECTA. 365
Sp. Atta cephaiotet Tabr., Formica cephalotei L., Db Gkbb, Ina. ill. PI. 31,
fig. II, Latb. Fowrmis, PL ix. fig. 57, Kollab, Branl, PoeOgL lad. Im.
fig. 10. The neuteiB are five lines long with » very large head, heart-
shaped, aimed behind with two sniall spinules, chestnut-brown all over.
These ants can strip whole trees of their leaves in a few hours. In Suri-
nam and Brazil^.
Add sub-genera : Gryptocerua Latb.^ Stenamma Westw., Myrme-
cina CuBns, Myrmecaria Saundebs, Ca/rehara Westw., Solenopna
Westw., Pheidole Westw.
Comp. J. O. Wbbtwood, Ducti^wm of ieveral exotic species cf Ants,
Ann, of not, Bist. VI. pp. 86—89.
Sub-genera Myrmica Latb. (and EcrUon ejusd.). Maxillary palps
long; with six joints.
Sp. Myrmita rubra, Formica rvbra L., Fabr., Swammbbdam, Bifid d. not.
Tab. XVI. figs. I — 13. Latbbillb, Fourmis, PL x. f. 62. Gomp. on this
ant, whose sting he has also figured, Lbbuwxnhobok, 58e, Missive von 9 Sept.
1687, Yervolg der Brieven, bl. 97 — 107.
B. Petiole of abdomen with a single joint.
Fanera Latr. Females and neuters aculeate.
Add sub-genera : OdoniOTJMchtia Latb. (Dacelon Pebtt does not
differ from Odantamachus, on Westwood's authority), Condylodon
Lund, TypJdopone Westw., Anamma Shuck.
Gomp. Shuokabd, Ann, of Nat, HisL v. pp. 396 — 338 ; Wbstwood,
ibid, VI. pp. 81 — 85.
Formica Latr. (spec, from gen. Formica L.) Sting none.
Add sub-genus : Polyergvs Latb.
Sp. Formica rvfa L., Latb., Fourmis, PL v. fig. 18, AB, (fern. FomUea
dorsata Panzbb, Deuischl, Ins, Heft 54, Tab. i). This species, without a
sting, aflPords the formic acid {Acidum formiearum s. formicicum, ctcidefor-
mique), a secretion from glands in the abdomen of the females and neuters.
This add is constantly fluid, colourless, of a pungent smell and sharp
taste. FouBOBOT and Vauqueun were of opinion that it consists of a
mixture of acetic and malic acid; but the experiments of Gehlen and
others have shewn that it is a peculiar acid.
Formica rtrfescens Latb., Foturmis, p. 186, PI. vn. fig. 38 ; this species
robs the nests of other species of ants of the larv» and pupn of neuters,
and carries them to its nest, where they are brought up with the young of
their robber by neuters which have proceeded from larvae and pupe stolen
^ Ants are very numerous iu South- America, and, by removing dead animals and
destroying other insects, perform the same office in the economy of nature with the
Cardbid and I>erme8tes and other Clavicomes. The natives also eat ants. Smoked
ants {Vachacos) are a favourite article of food with the Indians at the JUo-negro;
y. Humboldt's Beise in die JBqmnoctial-Qtgtnden, iv. s. 315.
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366 CLASS VIII.
at an earlier period. These ani-nesta are thus inhabited by two different
species, of which one alone works. They are the Amaztm-anU of HuBSB,
whose observations have been confirmed by Lat&killb {Mimaira aur
divert sujett, Paris, Svo, 1819, pp. 136 — 340), and by Hakhabt {WiMen-
Khafttieker ZeUtekr, von Lekrem der Baader ffochseh/ule, cited by Okebt,
Allg. Naturgeah, v. 2, s. 943 — 945).
Phalanx II. SoUtaria s. MutiUaricB. Males and females alone.
Males winged. Females apterous, without ocelli, furnished with
sting. AntennsB filiform, with first and third joints elongate.
BuBMEiSTER and Westwood place these insects in the neighhour-
hood of Scolia in the following division.
Dorylua Fabr., Latk. Antennae short, inserted near the mouth,
above the forcipate mandibles. Head small. Abdomen elongate,
cylindrical. Body, especially thorax, downy.
Insects of which the nudes alone are known, perhaps parasitic in ants'
nests. Sp. Borylm hdvolua, Mulilla hdvda L., DuM^. Cansid. gin, 9. 1,
Ins, PI. 11, fig. I (below) ; Cuv. R, Ani. id. HI., Ina. PI. 118, fig. i (the
feet are here badly depicted) ; habit. Cape of Good Hope. All the species
are exotic, from the eastern hemisphere, particularly Africa. (There is in
the Leyden Museum a species from Java and from Siberia f )
(Add sub-genera : Bhogrmu and JSnicttt4 Suuckard).
Labidus JuRlNE, Latr.
AH the species American. According to Shuokard, genus Typhlopone
Wbstw. should belong here and contain the females of the Labidi, on
which point see the opinion of Wsbtwood, Ann. of Nat, Hiat. vi.
But on Dorylua comp. by all means Shuokabd, Monograph of the Dory-
Udea, Awnala of Nat, Hiatory, v. 1840, pp. 188 — loi, pp. 258 — 271, pp.
315—39^.
Mutilla L. (exclusive of Mut, helvola.) Antennse inserted above
the anterior margin of the clypeus. Head transverse, broad. Abdo-
men oval or conical. Feet of females strong, with tibise spinous
ciliated.
A. Mutilla Latr. (spea of Mulilla L.) Thorax undivided.
Sp. MuUUa rvfipea Fabb., MtOiUa aellata Pansbb, 6 or 7 millim. in size,
thorax and feet red-brown, abdomen black with a white spot on the middle
and a transverse band of white hairs at the posterior extremity. The male
is Mulilla ephippium Fabb., Guy. R. Ani, id, iU., Ina, PI. 118, fig. 3.
Add sub-genus: Apterogyna Latr., Dalh. Antennae long, in
males almost of the length of body. Thoi-ax imdivided. Two
anterior segments of abdomen narrower, discrete.
Sp. Apterogyna Olivierii, IHctumn. rlaaa, d'Biat. not. Tab. 71, fig. 9, from
Arabia and Egypt &c.
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INSECTA. 367
Pacmmotherma Latb. Antemue (of make) bapectmata
Sp. Ptammothertna JldbeUaia, Guv. R, Ani, id, UL, Iiu. Fl. i iS, fig. 6.
B. Thorax (in females) divided or nodose. Sub-genera: M&-
thoca Latb. (male Tengyra Latb.), Myrmosa Latb., Myrtnecoda
ejusd. (male Thyivnui, ScotcBna).
Note, — That Tengyra \b the miJe of Mdhoca waa observed by Van dbb
liUmmf, Ann. det 8c. natvr. xvi. 1829, pp. 48, 49 ; on the others, comp.
Wbstwood, Introd. to Modem Clauif. 11. p. 215.
Genns Sderoderma Klug. Is this its place %
Family XVII. Foaaores s. Sphegina. Wings in both sexes
obvious, expanded. Tarsus of posterior feet simple. Aculeate, soli-
tary hymenoptera, {males 9Sidi females alone).
Diggers. This fiunily consists principally of the genus Sphex
(sand-wasp) of Linn.£U& The females of most of the species dig
in the ground nests for their young, and lay in these holes near
their eggs insects or larvee, sometimes spiders, as food for their larvae
when they leave the egg. The larvie have no feet, resemble mag-
gots, and spin themselves in, before changing into pupse. The per-
fect insect is usually very lively, and sucks with avidity the honey-
sap of flowers, on which (especially on the Umbelli/erce) it is
frequently met witL In many the lower jaws and under-lip are
prolonged and form a rostrum ; the ligula however is not filiform,
but commonly has a broad termination.
Grabro Fabr. Prothorax very short, linear, transverse, remiote
from the origin of anterior wings. Feet short or of moderate length.
Head large, quadrate above. Labrum concealed or scarcely exsert,
transverse. Abdomen obovate, constricted or clavate at base,
petiolate. Antennae often thicker towards the apex.
Sub-genera : Cerceris Latb., PhiUvrUhus Fabb., Latb., Fsen Latb.,
Jtjb. (Mimesa Shuk.), Alyson Jubike, Mellintis Fabb., Pemphredon
Latb., SHgmus Jubike, Crabro Fabb., Gorytes Latb., Trypoxylon
Latb., Fabb.
NoU. — Genus Crabro (In the stricter sense) is distinguished by antennss
geniculate, mandibles bifid at apex, a single cubital cell alone complete,
a radial cell appendiculate (another imperfect). Lbfelbtuib dk St. Fab-
OEAU divided it into several genera, of which for the sake of brevity I omit
the names. See his Hist. not. d. Hymin. ni. pp. 99, &c., and a critical
revision by Hbbbioh SoH^yFKB, Deutsckl. Ine. Heft 179 — 18 x. Comp.
also Dahlbom, Synope, HymenopUrol, Scandinav. i.
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368 CLASS VIII.
These insects are mostlj coloured black with yellow spots and
stripea The anterior part of the head {dypeua) is beset with fine
smooth hairs that often have a beautiful silvery or golden lustre.
They lay their eggs in holes, which they excavate with their fore-
feet, and place near them a provision of food (insects or spiders —
every species appears to select by preference a definite kind), which
they drag either with their jaws or their hind feet. Lefeletier
DE St. Faroeau was of opinion that certain species whose fore-feet
are not at all or only slightly haired, and hinder-feet without spines,
are unfit for digging, and that they lay their eggs in the holes of
other species, like the cuckoo in the class of birds. But later
observations oppose this; see Webtwood, IrUrod, to Mod, Classif. il
pp. 188—190.
Sp. Orahro cribrarius, Sphex cribraria L., DumAb. Ckma, gin, a. I. Ins. PL 31,
fig. 3; Panzkb, DeuUchl, Iw, Heft 15, Tab. 18,19 ; black ; a yellow traos-
verse streak forward on tbe thorax, as also a small double spot on the
middle of the thorax between the posterior wings and di£krent transrene
stripes, of which the two middlemost are interrupted in the middle ; the
tibioB and tarn of the same colour ; length 7 lines. The male of this, as of
some other species, has on the tibis of the forefeet a disciform expansion,
which on copulation serves to clasp the female. This disk has been taken
for a sieve (from whence the specific name is borrowed), and the property
of sifting the pollen of plants been recognised in the insect (Rolakdeb,
Venienak. Akad. ffandlinffor, Stockholm, 1751). This strange opinion owes
its origin to the erroneous notion that the light transparent points seen on
the disc are apertures ; it was refuted by Db Gebb (Mim. «. lea Jna. ii.
p. 818) and Goezb (Naiurrforacher n. 1774, s. 91 — 65).
Nyason Latr. Antennas filiform. Abdomen conico-ovate or
conical, broader at the base. Head moderate. The rest of the
characters of the preceding genus. Mandibles entire.
Sub-genera : Piaon Spinola, Latr., HUela Latr., Oxybdua Latr.,
JuRiKE, Nysaon Latr., Jurine, Astarte Latr. {JHmorpka Jurike.)
Sp. Oxybelua unightmia, Ordbro ufUglumia Panz., DeuUehl, Ina, Heft 64,
Tab. 44 ; Gu&nr, leonogr. Ins. PL 71, fig. 1, &c.
Larra Fabr. Prothorax short, transverse, linear, not extended
as far as the origin of anterior wings. Feet short or moderate.
Labrum concealed. Mandibles at the base deeply emarginate on
the outer side. Abdomen conical.
Dinetua Jurine, Miacophua Jurine, La/rra Fabr. (in part), Latr.,
Lyropa Illig., Latr. {Li/ria Fabr.), Falarua Latr, DryudeiUa Spi-
nola, Gaatroaericua ejusd. {Ann. de la Soc. entom. de France,')
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INSE(TA. 369
Sp. Larra ickneumoniformit Fabb., Panzeb, DeuUehl, Ins. Heft 76, Tab.
18, &c.
Bembex Fabr. Prothorax and feet as in the preceding genus.
Labrum exsert, often triangular, inflected. Mandibles narrow,
dentated on the internal side. Maxillae and labium often extremely
elongate, inflected. Body elongate, abdomen ovato-conical.
Sub-genus Bembex Latb., (and MonediUa ejusd.) Labrum trian-
gular. MaxiUsB and labium veiy long, linear, inflected, forming the
promu8ci&
Sp. Bembex roatrata, Api$ rotircUa L., Panzbb, DeuUchl. Int, Heft i. Tab.
10 ; DuiciB. Cons, g&n, s. I. Ins. PI. 30, fig. 10; Latb. Ann, duMus. xiv.
PI. 26, figs. 9 — 13; black, felty, with yeUow feet, and tight greenish
yellow transverse bands on the abdomen ; 9—10 lines long. The female
digs holes in the sand, and lays in each of them an egg with a sufficient
quantity of Diptera (especially EriMdUs tenax) for feeding the larva. Most
of the remaining species are at home in warm regions ; but the species
quoted occurs occasionally all over Europe, even in Sweden.
Sub-genus Stizus Lath., Jusine. Labrum small, semicircular.
MaxiUie and labium porrect, not inflected, nor elongato-linear. (In-
termediate lacinia of labium elongato-cordate. Maxillary palps
somewhat long, extended beyond the apex of maxilla.)
Sp. Bembex rt^/icomis Fabb., Bnt. syst., Larra ruficomis ^jusd. Sytt. Piez.,
Cuv. R. Ani. 6d. ill.. Ins, PI. xai, fig. 3 ; habit, in south of Europe and
Numidia, &c.
Sphex Lin. (exclusive of many species). Pro thorax continued
laterally as far as the anterior wings, narrowed forwards, resembling
a joint or node. Labrum scarcely or not at all exsert. Three com-
plete cubital cells. Antennae slender, with joints elongate, often,
at least in females, convolute or arcuate. Posterior feet very long,
with tibiae and tarsi spinose. Abdomen adhering to thorax by a
long petiole.
* With mandibles edentulous,
Sub-genera : Felopoeus Latb., Fabr., Podium Fabr., {Trigonop-
sis Perty), Fodium Latr. (not Fabr.), Ampulex Jur., Latr.
{Chlorum Fabr. in part).
* * With mandibles on inside dentate.
Sub-genera : Dolichi/ma Latr., Trirogma Westw., Aphlelotoma
VOL. 1. 24
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370 CLASS VIII.
Westw. *, Chlorion Latr. and Fabb. in part, Sphex Latb., Proncevs
IulTBl, Ammophila Kibby*.
Sp. Sphex iabulo$a L. {AmfnaphUa), Panzxb, Deutschl, In», Heft 65. Tab.
Pompilus Fabr. Prothorax produced as far as anterior wings,
subquadrate, not narrower forward. Abdomen with very short
petiole. Posterior feet long, ciliated on the inner margin. Cubital
cells three, or two, alone. Antennae as in the preceding genus.
Sub-genera : Aporus Spinola, Flaniceps Latb., PompUtut Latb.,
Geropales Latb., Fabb., Pepsia Fabb. (in part), Latb.
On these and other 8ub-genera oomp. Lkfelbt. Hymenop. in., and
J. SoHioDTB de tpeciebtu P&mpiUdarum in Dania obviii, Kbotsb's Tidsdcr.
I. 1837, pp. 313—354. Tab. IV.
Sp. PompUua viatieua Fabb., Sphex futca L., Panzkb, DetUschl, Int. Heft
65, Tab. 16; DiTX^ Com, gin, 9. 1 Int. PI. 33, fig. 3, ftc.
Sapyga Latr., Hellua Fabr. Prothorax transverse, produced
to anterior wings. Abdomen elongate, shortly petiolate. Labrum
not or scarcely exsert. Mandibles strong. Eyes emarginate (lunate).
Antennffi of length of head and thorax at the least, towards the
apex thicker or sub-clavate. Feet short, slender, with smooth
tibiffi.
Sp. Sapyga quinquepwnctata Latb., Oener, Cruti, et Ins, Tab. xiii. fig. 9,
(of which perhaps Sapyga ifaria, Encyd. tnHh. and Gu^BiN, Icanogr, Ins.
PI. 69, fig. II, IB merely, aa Lefeletibb suppoees, a variety), habit, in
France.
Note. — Genus Polochrum Sfikola, unknown to me, dififers from Sapyga
by its antennas filifonn.
Here also were referred formerly genera Thynnus Fabb. (in part), Latb.,
and Scokena Latb. Partly at least they belong to the MutQlaria, and contain
males of genus Myrmecoda. Comp. also genus Amblysoma Westw. and
Anodontyra ejusd., Gu^bin, Magas. de Zocl. 184 1, Ins. PI. 80, 81; females
are unknown.
Scolia Fabr. Prothorax produced laterally as far as wings, as
in the preceding genus. Eyes emarginate, reniform. Antennae
thick, filiform, in males almost of the length of head and thorax,
^ AwmIs and Magazine of nai. Hist. vn. 1841, p. 151.
■ Transact, of the Linn. Soc, Vol. iv. 1798, p. 195. For some other sub-genera we
may refer to Lbpeuetieb, Hymenop. m.
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iNSEcrrA. 371
in females shorter, arcuate. Labrum retracted. Mandibles strong,
cruciate. Abdomen elongate, with short petiole; body hirsute.
Sp. Scolia quadrimacuUUa ¥,, DuMiB. Connd. gen. s. L ln$, PI. 31, fig. 9,
habit, in Korth America, Ac. Most of the species exotic, some very large.
In the South of Europe occur Scolia hortorum Fabb., Scolia imubrica
(Scolia ifUerrupla) Panzkb, DeuUchl. Ins, Heft 61, Tab. 14, &c.
Note. — ^The males are distinguished by longer abdomen, trispinose at
apex (anus tridentate), whence the name of the genus appears to be derived
{(TKuiKos, tpina). Feet of females thicker, very hirsute.
Add sub-genera Meria Illio., Latr., M^/zine Latr, Tiphia Fabb.
Section II. Terebrantia, Abdomen in females furnished with
a borer or ovipositor, sessile in many. Antennae various, usually
with joints more or fewer than thirteen. Upper capitulum of femur
mostly distinct, as though forming a second trochanter.
Amongst these Hymenopteray which deviate more from the usual
type, there are many species whose larvse are provided with six
homy feet. Hartig first drew attention to the difTerence of articu-
lation between the hip and the thigh, and named these hymenoptera,
on that account, ditrocha, and those of the former division, on the
other hand, numotrocha. Sundevall has given a better explanation
of this disposition, which we have followed in our statement of
characters; ArsberdUelse om nyare zoologiska Arbeten 1837 — 1840.
Stockholm, 1841, pp. 324, 325, The genus Ckrysia according to
this character ought to belong to the first division.
A. Efntcmophaga {Pupivora Latr.) Abdomen petiolate. Larvae
apodous, mostly living parasitically in other insects.
Family XVIIL Chryaidides, Inferior wings with no cells,
but only some longitudinal veins ; superior with radial cell long,
single cubital cell imperfect. Antennae filiform, with thirteen joints
in both sexes. Abdomen joined to thorax by narrow, very short
petiole, below plane or vaulted, composed of only three or of four
conspicuous segments, dentate posteriorly in many. Integuments
of body hard, smooth. Borer of females inclosed in concealed
terminal segments of abdomen, receiving one another by inva-
gination, composed of three setse, the groove of one containing the
two others.
The golden^ujoaps. These insects wore thus named on account of
the shining metallic colours in which they glitter (the abdomen is
24—2
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372 CLASS VIII.
mostly gold-green or purple-red, sometimes blue, just as the head
and thorax usually are), and which have caused them to be compared
with humming-birds.
The first joint of the antennae is elongate ; at the second joint
they are bent geniculately. The females lay their eggs in the nests
of other hymenoptera, whose larvse are eaten by theirs. These
insects are protected firom the sting of bees and other hymenoptera,
their natural enemies, by their hard integument ; and besides this,
they have the power of bending the abdomen under towards the
thorax, and so, like the Armadillos amongst the mammalia, of
contracting themselves into a balL They form the transition
between the first and second divisions, and are joined to the first,
that of the aculeata, by Hartio, because they are without the part
which he considers to be a second trochanter.
Comp. on this famOy Lbfeletieb, Mim, xwr qudquea apices nouv. cTin'
aecU8 de la tection dea ffymenoplirta porl&4uyaux, ay. fig. col., Ann.du Mut.
vn. 1806, pp. 115 — 129.
The borer, improperly named a sting, is described and figured by Ds
Gekb, if^. p. VHid, d, Ins. 11. pp. 834—836, PI. 28, f. 19— a i, PL 29,
figs. I, 2.
Pamopes Latr. Maxillaa and labium very long, linear, form-
ing a kind of promuscis inflected beneath the thorax. Maxillary
and labial palps very short, biarticulate.
Sp. Pamopea eamea Latb., Duvia. (7o»«. gSn. 8. 1. Ins. PL 31, fig. 7 ; Ah-
BXNS (Gkrmab) Faun. Ins, Eur. Fasc. 11. Tab. 10. This species has its
habitat in the south of Europe, and ktys its eggs in the nest of Bembex ros-
trata; see Latbbillb, Ann. du Mus. d^Hitt.nat. xiv. p. 415.
Chrysis L. Labium not in form of a promuscis. Maxillary
palps with five joints, labial triarticulate.
a) Palps equal.
Sub-genus StiUmm Spinola, {StUhum and JUtichrc^us Latr.)
h) Maxillary palps longer than labiaL
Sub-genera: Gleptes Latr., Clvryda Spinola, Elampua Spinola,
Hedych/rwm Latr.
Sp. Chrysis ignita L., Fbisch, Ins, ix. Tab. x. fig. i, Sulzeb, Die Kenm, der
Ins. Tab. Xix. fig. xai, Cuv. R. Ani. Sd. ill., Ins. PL 1 16, fig. 6, glittering,
with thorax green, and abdomen golden irom above of a fire-red play,
and having at its extremity four denticles. Chrysis cyanea L., Panzkb,
Deutschl. Ins. Heft 51. Tab. 10, ftc.
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INSECTA. 373
Family XIX. Oxyura s. Proctotrupii Latb. {Codrini Dalman,
Nees von Essenbeck, with addition of other genera.) Inferior
wings without nervures, superior either in like manner without any
nervures, or with few and longitudinal only, destitute of cubital
cell and often of humeral likewise. AntennsB with 8 — 15 joints
(in most 10 — 12), filiform or thicker towards extremity, in males
mostly longer, in females sometimes clavate. Borer or ovipositor
at the extrQ^lity of abdomen, in some exsert, conical, in others re-
tractile within the abdomen^ containing three setse.
These taU-boring ichneumons are mostly very small, some only
1 or I line in size, or even smaller (Jchnefwmon atamua L.), so
that the investigation of the oral parts is rendered yeiy difficult,
and the characters derived from them are often insecure. The head
in most of them is broad, the thorax long, the abdomen oval or
conical In some there are no wings, or very imperfect wings
(especially in females) ; some have only four joints in the tarsus
{Iphib-achdu8 Haliday, Mymar, kc\ which has also been observed
in the fEuooily of the ChalddicB, a remarkable anomaly in the ffymen-
optercL The economy of the greater part is unknown ; but we may
conclude, from those whose metamorphosis is known, that their
larvce live parasitically in other insects, like the true Ichnewimms,
with which LiNifiEUS united the few species that were known to him.
The larvsB of Platygaster live as parasites in those of the genus
Ceddomyia {Diptera). Other species lay their eggs iu the eggs of
other insects (especially of Lepidoptera and some Jffemiptera), To
these belongs the Ichnevmum avulorum Ij., according to Haliday a
species of Myma^y and also the genus Teleas,
CkHnp. on this family C. G. Nbes von Essbkbieck, ffymenopUrorum Ich-
newnonibtu affinium, MonograpkuB, StuttgartisB et Tubingse, 1834, Syo. u.
pp. 31 1 — 397 ; and Wbstwood, Introd. to Modem (Muific. of Int. n.
pp. 167 — 173; as also the works of Halidat there referred to, and other
English entomologists.
Mymar Haliday. (Palps none ?) Antennae 13 — 10 jointed in
males, long, broken, 10 — 9 jointed, clavate in females. Wings
narrow, often linear, anterior broader at the extremity (spatulate),
ciliated on the margin. Abdomen iq some sessile, in others peti-
olate.
Sp. Mywar pulcheUus Walkeb, Gu^in, Iconogr, liu, PL 68, fig. 6 ; Heb-
BiCH-ScHJiFFBB, Deuitchl, Ins, Heft 184, Tab. 135, (fig. copied in Cubtis,
BrU. ErUom. Tab. 411).
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374 CLASS viir.
Nixte. — Here belong yarious sub-genera constituted by Halidat, on
which oomp. Wbbtwood, 1. 1. Omeric Synopsis, pp. 78, 79.
PlatygcLster Latr. (with the addition of genera Scelio and Teleas
ejnsd.) Palps short. Abdomen depressed, sessile or affixed by a
short petiole. Antennae broken, with ten or twelve joints, in females
incrassated towards the apex.
Sp. Tdeas IcBviuactUuM Ratzbbubo, Ford-Intektm, m. Tab. Yin. fig. 8 ; the
larva lives in the eggs of the Bombyx pint ; in those of Bomb, neuttria
lives the larva of TeUas terdnmu Ratzbbubo, Tdecu owlorum BouoH^.
Add genus Iphiirachdus Halid. (tarsi with four joints) ; other
sub-genera of this author are enimierated in Westwood, Generic
Synops, pp. 77, 78.
Sparaston Latr. Abdomen depressed, sessile. Antennae in-
serted below the frons, twelve-jointed. Maxillary palps long,
filiform, five-jointed, labial three-jointed. Wings almost without
nervures, with stigma distinct.
Sp. Spar<uion frontale ItATTL., Oaraphron comutut JuBiNB, HynUn. PI. 13,
fig* 44> CuviBB R. ani id. Uluttr., Ins. PL 116, fig. i. Hebbioh-
SOHJBFFEB, DeiUsehl. Ins. Heft 184, Tab. 25.
Ceraphron JuRlNE (in part). Abdomen subsessile, conico-acu-
minate. Antennas broken, with eleven or twelve joints, the first
elongate. Maxillary palps long, fonr-jointed, labial with two or
three joints. Wings without nervures, in some none.
Sub-genera : Megaapihis Westw., Micropa Haled., GaUiceras
Nbes.
Sp. Ceraphron sulcatus Jubinb, HynUn, PL 14.
Dryirma Latr., Gonatopua Klug., Dalm. Abdomen convex,
subsessile. Antennae ten-jointed, porrect, mostly short or moderate.
^ Mandibles somewhat prominent, acute, frequently dentate. Max-
illary palps elongate, five- or six-jointed. Anterior wings with
stigma pretty conspicuous, and radial cell incomplete, with two
brachial cells; posterior wings increased by a lobe, in some no
wings.
a) With anterior tarsi of females raptoriaL
Sub-genera ; Dryinua Latr., ArUeon Latr., Ohehgynua Haltd.,
Gonatopus Ljung.
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INSECTA. 376
Sp. Dryinui cursor Haud., QjjtBJtf, Icanogr,, Ins, PI. 68, fig. i, Hebbioh-
SoHJiFFSB, Deuischl, Ina, Heft 184, Tab. 31, (fig. oop. in CusTiB, Brit.
EnUm.); — Dryin, formicarita Latb., Cfener. Orud, H Int. I. Tab. zn,
fig. 5» &c.
h) With anterior tarsi of females simple.
Bethylus Latb., Omahis JuRiNE^ Nees. (Maxillarj palps sex-
articulate.)
8p. Bethylus cmopterus, Tiphia cenoptera Panzkb, Btutsch. Ins, Heft 81,
Tab. 14, kc.
Add sub-genera Aphdapus Dalm., Nkss, Myrmeeomor]^us,'^EniboUMus,
Epyris Westwood, 1. 1. p. 76,
Prociotrupea Latr., Codrvs JuRlNE, Nees. Abdomen conico-
petiolate, with anal segments attenuated to form a curred tubule
sheathing the borer. Antennae inserted in middle of frons, thir-
teen-jointed in both sexes, straight, porrect. Maxillary palps four-
jointed, much longer than labial. Superior wings with longitudinal
nervures and stigma distinct.
Sp. Procotrupes ccmipanulator, Bassus campanulaior Fabb,, Gebhab, Faun,
Ins. Burop. Faso. V. Tab. x6 ; Prodcirupes paUipes, Jubine, Bymin. PI.
13, fig. 46, &o.
Diapria Latb., Psiltis JuRiNE. Abdomen petiolate, campanu-
late. Antennae inserted in frons, with 12 — 15 joints. Maxillary
palps elongate, five-jointed. Wings often without nervures, and
with stigma little distinct or none.
Sp. Diapria verticiUata, Psilus degans, Jubinb, Hymin. PL 13, fig. 48;
Diapria comuta, Panzxb, DeulschL Ins, Heft 83, Tab. 11, &c.
Add sub-genera Helorus Lat&, Bdyia Latb. (Bdyta Jubine and
Cinctua ejusd.), and several genera of recent authors, principally
English, on which oomp. Westwood, L L pp. 75, 76.
Family XX. Chalctdue «. Chahides. Posterior wings without
nervures or with a single nervure submarginal, short ; anterior with
only one cubital cell, imperfect, radial cell mostly wanting. An-
tennae with joints various in number, not more than thirteen, with
first joint elongate, almost always geniculate, often thicker towards
the extremity. Head anteriorily bisulcate for receiving first joint
of antennae. Palps very short. Borer originating from a chink of
inferior surface of abdomen remote from apex^ mostly concealed or
exsert at the termination alone.
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376 CLASS VIII.
The ChalcidicB are small Insects^ mostly shining with metallic
lustre, many species of which are able to leap, though that is not
always the case where the ability might be inferred from the thick-
ness of the hind-feet. Their larvse live parasitically in those of
other insects, especially of Lepidoptera and Hymenoptera^ and also
in their eggs; they are small maggots without feet. The pupee
are mostly not included in a web. The number of species of this
division is extraordinarily great; in Great Britain alone nearly
1200 species have been found
Gomp. on this family amongst others^ Spinola, Estai cTune nowfdU
Clas9\fication de$ I>iploUpaire8, Ann, du Mus: xvn. 1811, pp. 138 — 15a ;
Nebs von Essbnbbok, ffymenopt, Ichneumonib. off, Monagr. n. pp. i — 310.
(Monographia PteromcUinorum, amongst which some families are arranged
that belong to the preceding family) ; the monographs of Dalman and
BoHEMAN, in the Trans, of the Swedish Acad, at Stockholm, for 1820 and
1833 ; BoTEB DE FoNSOOLOMBS, MonoffT. chalciditum, ftc., Ann. da Sc.
ntOur, XXVI. 1833, pp. 473 — 307, and Walker, Descriptions of the British
Ckalcidiies in Ann. 0/ Nat, ffisl. Vol. I— iv. 1838, 1839.
A. Prothorax narrower than mesothorax, attenuated towards
the head. Femora of posterior feet scarcely larger than the rest ;
tibisB straight.
Eulophus Geoffr., Latr., ErUodon Dalm. AntennsB with seven
or eight joints, very rarely with nine, in males sometimes ramose
(pectinate with three internal branches). Tarsi with four joints.
Sp. Eulophus pectinicomis, Ichneumon pecUnicomis L., Gn&Dr, Iconogr,
Int. PL 67, fig. 15, Ac.
Add sub-genus CirropsUtia "Westw., and several more, on which
comp. Westwood, Introd, to modem Ckusificaiion o/Ina, IL Generic
Syriopsis, pp. 73 — 75.
Encyrtus Latr. Antennae eleven- or twelve-jointed. Inter-
mediate feet longer than rest, with tibiae terminated internally at
the extremity by a strong spine.
Species numerous ; here belongs a figure of Schellenbebo, Tab. xiv. of
Mira mneora by name, placed wrongly amongst the Diptera.
On several sub-genera, to be referred to Encyrtus, comp. Westwood,
1. 1., pp. 73, 73*
Pteromalua SwEDER. (in part),LATR4, (species oi DiplohpisyFABK.)
Antennae eleven- to thirteen-jointed. Middle feet like the rest.
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IN8ECTA. 377
Sp. Pteromalut puparum, lehnetmon puparum L., Gocdjibdt, Metam, lutfur.
I. Tab. 77, p. 144, BxBSKL, Int. n. BomJbylior. et Vespar, Tab. m. ftc.
Add genera deonyrmu Latiu, FerUamptts ejusd. and nnmerous
sub-genera of more reoent authors ; on which oomp. Westwoop, L L
pp. 67—72.
Does genus Eucha/ris Latr. also belong here ? Prothorax short,
narrow, mesothorax gibbous, elevated Antennae moniliform,
eleven- or twelv^jointed, straight. Abdomen with long petiola
Feet slender, with posteiior femora not incrassated.
Sp. EucharU adacendent, Oynipt adscendeiu Panzbb, DeuUchl, Int. Heft 88,
Tab. 18, Ac.
0
B. Prothorax transversely quadrate.
1) Posterior feet not much differing from anterior, with tibiae
straight.
EuryUyma Illig.^ Latr. Antennae eleven- to thirteen-jointed,
in males longer, with joints distinct, nodose.
Add genus SpaUmgia Latr., and other sub-genera; on which see
Westwood, L L pp. 66, 67.
ToTdcantha Latb.
2) Posterior feet with femora veiy large, ovato-lenticnlar, with
tibiae arcuate.
Chalets Fabr., Latr. Prothorax much broader than long.
Antennae eleven- or thirteen-jointed, thicker towards extremity.
Borer concealed.
Sp. Chalcia minuta Fabb., CJuUcis femorata Dalm., Panzxb, Deuiaehl, Ins,
Heft 31, Tab. 6, DuM^. Cons, gin. $. I. Int. PL 34, ^. 1, &c. (Sub-
genus Brachymeria, Wbstw.)
Sub-genus Chi/rocera Latr (With antennas of males pectinate).
Dvrhinua Dalm.
Palmon Dalm.
Cowwra SpnrOLA. (With abdomen conical, acuminate.)
Sp. ConwrajUwieana Sfivola, Gu^rik, Magaxin de Zool. 1837, Int, PL 180 ;
hab. in Braail.
On aome other rab^enera oomp. Wbstwood, 1. 1. p. 66»
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378 CLASS YIII.
Leucospia Fabr. Thorax gibbous, prothorax transversely quad-
rate. Upper wings doubled longitudinally. Antennas with twelve
or fifteen joints, incrassated at apex. Borer reflected above the
back.
8p. Lew^oapU gigat Fabb., Kluo., Panzbb, Im. Heft 84, Tab. 17, 18, Cuv.
R. ani. id. ill., Ins, PL if 6, fig. 6. Leucospis intermedia Illio., Leue.
dortigera Panzeb, DeuUcU. Ins. Heft 15, Tab. 17, Dum^bil, Cans, gin,
s. I. Ins. PL 34, fig. 1, &C.
ThiB genus contains species of ^ inch and more, whilst the smallest
species are still more than 3 lines in size, so that it may be considered
gigantic in this family of dwar&. In Europe species of Leucosjns are
found in the southern regions alone, principally in Italy. To the extrica-
tion of the synonomy Illigbb, Kluo, and la^y Webtwood, have contri-
buted : see the monograph of the last-named author, Entomol. Magas. n.
p. 212, kc.
Family XXI. Ichneumonides. Wings four, veined, the anterior
always furnished with complete cells. Maxillary palps distinct^
elongate. Antennae mostly setaceous or filiform, long, with numer-
ous joints. Body elongate, slender. Ovipositor straight, often
exsert, bivalved, including a borer of three setse.
Slip-uxupSy IchnetMnona. We unite in this family the Ichneu^
monidea and Evcmiales of the systematic entomologists of recent
times. Of many species the economy is yet unknown, yet of all
whose mode of life has been observed the larvs are found in other
insects, and nearly always in the larvsB of these. They are especially
caterpillars, the larvsd of Lepidaptera, in which Iclmeiimona lay
their eggs j the laryee of Evania are parasites of the genus BlcUta,
and probably live, according to an observation of Mac Leat (related
by Wkstwood, Introd, to Tnod. Classif. of Ina. l. p. 422) in the
membrane of their eggs.
Comp. on this family amongst others:
J. J. TsEKTBFOHL, Mevisio critica generis Ichneumonis, Okkn'b Isis,
1816, pp. 55—87, pp. 293—308.
J. L. C. Gbavxnhobst, lehneumonologia europcea. Vratislaviie, 1829,
ni. Vol. 8vo.
Nbes ab EssENBBCKj ffymenopteroTum Ichneumanib, affin. Monogrct-
pkia. Volumen lum.
J. T. 0. Batzebubo, Die lehneumonen der FonHnsdiien. Berlin,
1844, 4to.
Phalanx I. Ickneumones. Abdomen inserted between the two
posterior feet. Antennce mostly setaceous or filiform, composed of
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INSECTA. 379
a great number of joints (sixteen, and many more*). Maxillary
palps^ mostly five, articulate.
A. Ickneumones adscitt, a. Braconides. A discoidal cell imder
the first cubital cell extended to the margin of wing, not divided
by a recurrent nervure. Second cubital cell firequently large,
Aphidius Nees. Head transverse, with vertex broad. Abdo-
men affixed by a short cylindrical petiole, incurvatile beneath the
thorax; Borer not exsert. Antennse with joints very distinct,
rather few (eleven to twenty-four). Maxillary palps shortish (five-
or four-jointed).
Sp. Ichneumon Aphidum L., Aphidiiu varius Neeb, Ds 6bbb, /rw. n. PL
30, f. 12, 13. (The fig. of Panzsb, DeuUehl, Ins. Heft 95, Tab. 13, also
belongs to this spedes according to NssB.) This smaU species lays its eggs
in Plant-lice, in each one a single egg. The pupa of the Ichneumon lies
curled up in the body of the Aphis. See Lbbuwekhobok, Sevende vervolg
van Brieven, bl. 135 — 194, I34e MissiTe Tan 26 Oct. 1700, (and the fig.
bl. 117 — 281), also Db Geer, L 1. pp. 866—875".
Sub-genera: Trioxys Halid., MonocUmua ejusd, ^oaurea Westw.
{Trionyx Haud.), Epheclrua Haud., Prcbon Haud.
Alysia Latr. Head broad. Abdomen sessile. Borer exsert.
Mandibles subquadrate, with apex tridentate, divaricate (even when
drawn together, distant). Maxillary palps sexarticidate. Antenn®
moderate or long, with more than twenty joints.
8p. Alysia manducator, Ichn. manducalor, Pahzeb, JhuUchl. Ins, Heft 73,
Tab. 4, Gu^RiK, Icofnogr. Int. PL 66. fig. 1 1, &c. The larva of many
species of this genus live in the pup» of Diptera, others in the larvie of
Scardbcsi.
Sub-genera : Ccelinius Nees (comp. Herbich-Sch^ffer DeuUch.
Ins. Heft 153, 154, 156), ChoErmsa, ChorebuSy Dacrvusa, (Enone,
Ckasmodon Hauday, (Westwood, Generic Synopa. £ %S), Copiaura
SCHIODTE.
^ Some species of the genus Aphidiua Nees, of which Haudat fonns the genus
JE^hedrus, make an exception to this, and have only eleven or twelve joints in the
antenne.
' These small parasites have their own in return : larvae of Oynip$, parMites of the
second order. See Goezb, NaturforKher, xii. 1778, s. 197 — 320.
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380 CLASS VIII.
Bracan Fabb. (in part), Latr. Clypeua exscinded, a hiatus
being left above the mandibles. Maxillary palps quihquearticulate.
Head transverse or subglobose.
Sp. Braccm impostor Nebs, Bracon denigrator Fabb. (exd. syn. L.)^
Panzsb, DeuUchl. Im. Hefb 45, Tab. 14, &c.
Eogcu Nebs. (See Westwood, L L p. 64 for other sub-genera).
8tgalj[)Ati8 JjATH. {Chelonus Jvm^E), Clypeus entire. Abdo-
men fornicate beneath, triannulate above, or continuous, no vestige
of incisures remaining, with all the segments united into one.
Maxillaiy palps sexarticulate, labial shorter, quadriarticulate.
Sp. Sigalphus irroratar, Cryptua irror<Uor Fabb., Db Gbeb, Ins, I. Tab. 36,
fig. la ; Gn^iN, Icognogr., Int. PL 68, fig. 9 (in this figure the diviaion
of the nervurea in the wings is incorrectly represented as though there
were a second recurrent nerve, as in the Ichneumonei genuini) ; 4J lines
long ; expanded wings 8 lines ; abdomen glistering at the extremity with
brownish green from fine smooth hairs, wings brownish with blacker exter-
nal margin and a white spot in the middle imder the radial cell. The
larya> according to Ds GSEB, lives in the caterpillar of Nodua psi, 1. 1.
p. 577.
Helcon Nees.
Microgaster Latb.
Note, — For other genera and Bub-genera^ here omitted on account
of our limited space, the works recommended above may be con-
sulted
B. Ichneumones genuini. Recurrent nerves two, one dividing
the area situated under the cubital cells. First cubital cell large,
confluent with the first discoidal cell ; second cubital cell rhombic,
pentagonal or trigonal, very small, in some none. Maxillary palps
with five joints, labial palps with four joints.
In this division no such small species occur as in the preceding
(ex. gr. the genus Aphiduui). The larvse live principally in cater-
pillars. Some species do not lay i^Jieir egga in the caterpillars, but
fitsten the eggs, which are provided with a pedicle for the purpose,
on the akin of the caterpillars. See Habtiq, Ueb. cL gestidten Eier
der Schlupftoespen, Wiegmakk's Archiv, 1837. s. 151 — 158. Taf iv.
Gbavxnhobst has described more than 1600 species of IchnenfMna gei^iUni ;
a number which will be continually increased by fresh observers.
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INSECTA. 381
t Genuine Ichneumons, with abdomen convex or depressed,
a) With abdomen petiolaie or tmh-pdiUaU.
Ichneumon L, (exclusive of many species). Head narrower than
thorax. Borer subexsert or concealed. Second cubital cell dis-
tinct, mostly pentagonal.
8p. Ichneumon aputaiar Fabb., Panzer, Deuikhl. Ins. Heft 19, Tab. 90;
— Ichn, Troscheli Batzbb., in the caterpillar of Nodua piniperda.
Tryphon Fall. Head narrower than thorax. Borer subexsert
or concealed. Second cubital cell almost obsolete, triangular.
Abdomen elongate.
On this genus, Which contains yery numerous species, oomp. Gbayxn-
BOB8T, Ichnewmol. ii. pp. i — 368.
Add sub-genus PolyUaatua Habtig, Schiodte.
Megastehis Schiodte.
Trogits Panzer, Gravenh. Head transverse. Borer concealed.
Second cubital cell triangular or quinquangular. Scutellum gib-
bous, prominent. Abdomen distinctly petiolate, oblong.
Tragus lutorius, Ichneum, ItOorius Fabr., Db Gebb, ii. PI. 19, fig. 9,
p. 848 ; one of the largest native hymenoptera, 10 lines to 1 inch long ;
thorax black, scutellum sulphur-yellow, feet and head yellow beneath,
abomen red-brown, at the extremity blackish. The larva lives in the cater-
pillar of Sphinx ocellcUa, Sph. pinaatri, &c.
Alomya Panzer, Gravenh.
Crypius Fabr. Head transverse. Abdomen oval, distinctly
petiolate. Borer exsert. /
Note, — Some species are distinguished by their small sisse, the defect of
wings, or by rudiments alone of wings : sub-genus Pezamxichtis Gbavenh.,
Sp. Vrypt. niffTihcindiu, Ichn. pedicularius Panzsb, DeuUchl. Ins. Heft
81, Tab. 13, &c.
Add genus Cylloceria Schiodte, see Gu£BIN, Magcta. de ZooL 1839,
Ins. PI. 9, 10.
Xorides Latr.
Accenittis Latr.
t /9) W'dh abdomen sessile (extremely short petiole).
Pimpla Fabr. Head transverse. Borer exsert, long. Antennae
long, filiform, slender, with numerous joints. Mandibles bifid at
apex.
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382 CLASS VIII.
a) With Beoond cubital cell eyanesoent.
Sub-genera : Glypta Gravenh., Folysphincta ejusd, Schizopyga
ejusd., CUstopyga ejusd.
h) With second cubital ceU distinct, mostly triangular.
Sub-genera: Rhyasa Gray., Trachyderma ejusd, EphiaUes
ScHBANK, Grav., Pimpla Grav., Lissonota Grav.*
Sp. Pimpla (EpkiaUes) manifestator, Ichneumon manifestator L., Paitzkb,
DetUachl. Int. Heft 19, Tab. 21, Duk&il, Cons, gin. b. I. In%. PL 32,
fig. I. Cut. R, ami., H. HI., Ins. PI. 1 10, fig. 8, Ac.
Metopius Panz., Gravenh., {Peltastes Illig.)
Sp. Ichneumon necatorius Fabb., Ichn. vesp&ldes Panz., Deuischl. Ins.
Heft 47, Tab. 19.
Bassus Fabr., Grav.
ft Cknuine Ichneumons with abdomen compressed.
Banchus Fabr. Abdomen sessile, or with veiy short petiole.
Ophion Fabr. Abdomen falcate, distinctly petiolate. Antennae
slender, filiform.
Sub-genera: Anomalon Jurine (in part), Gravenh., Ophion,
Pani8cu8y ko.
Sp. Ophion glaueopterus Fabb. ; — Ophion eircumJUxus, Ichneum. dreum-
JUxus L., Batzbb. Forst. Ins, m. Tab. yi. fig. 2, &c.
HeUwigia Gray. Abdomen petiolate. Antennae clavate.
Gomp. Gbaybnhobst, Hdlwigia, novum inseetorum genus ; Nw. Ad.
Acad. Cobs. Leop. Car. Natur. Cutiosor. xi. 1823, pp. 315 — 321, Tab. 43.
Phalanx II. Uvaniales. Abdomen inserted into thorax above,
before the origin of the two posterior feet. Antennae filiform
or setaceous, with thirteen or fourteen joints. Anterior wings with
distinct cells, posterior veined, destitute of cells. Maxillary palps
longer, sexarticulate, labial quadriarticulate. Posterior feet with
coxae long and strong, and femora often incrassate.
^ Of what small value this second cubital cell or areola is as a character in PimpUe,
appears from some species of Lissonota Gbaviekh., where it almost entirely disappears,
or is sometimes present on the right wing and 'wanting on the left Gbavenhobst,
1. 1. III.
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INSECTA. 383
A. Abdomen of moderate size or elongate.
f Borer exsert
Auhcaa JuRiNE. Abdomen compressed. Antennae setaceous.
Sp. Atdaeus ttriatua, Jubins, ffymSn, Tab. 7i fig. 15: habit, on the moun-
tainB of Switzerhmd.
Fcenus Fabr. Abdomen elongate, clavate at apex, exceeding
the length of head and thorax. Antennas filiform.
Sp. Faentu JaetUator, IchneumoH jaculator L., B4A.UMnit, /iu. iv. PL 10,
figs. 14, 15, Pahzeb, Veutschl. Ins. Heft 96, Tab. 16, DuviBiL, Cons,
gSn. i, I. Ins. PI. 31, fig. i, ftc.
ft Borer concealed
Pelecinus Latr., Fabr. Inferior wings almost without ner-
vures. Abdomen very long, filiform in females, moderate and
clavate in males.
Sp. Pdeciwiis polyeerator Latb., Gu^in, Iconoffr., Ins. PI. 65, habit, in
North and South America.
Comp. on this genus Lepeletieb and Sebville, Encyl. method., Ins.
Tom. X. T825, pp. 39, 30 ; De Romand, Note su/r U genre Pelecinus,
GuiBlN, Mag. de Zool., 1840, Ins. PI. 48, 49 ; ejusd. Notice s. I. genre
Pelec.; ibid. 1842, PI. 86 ; Klug, die Arlen der OaUung, Pelec., GsB-
mab's ZeUschr.f. d. Entom. m. 1841, s. 377 — 388, Tab. n. (A genus in
the opinion of this author related to the Oxyura, the genus Monomachus
Wbstw., forming the transition to the genus ProcMrupes.)
B. Abdomen very short, oyato-triangular, compressed, abruptly
petiolate, often inserted almost beneath the scutellum.
Evania Fabr., Latr.
Sp. Evania appendicigaster, E.IBBT and Spenob, Introd. to Entom. PI. iv.
fig, 2, DDMiB. Cons. gin. s. I. Ins. PI. 31, fig. 3, &c.
Sub-genera : Brachygaster Leaoh, Hyptia Illio. Gomp. on this genus
and the allied sub-genera Sfinola, Gu^in, Revue Zoolog. 1840, pp. 244 —
148, and Wbbtwood, Trans, of the Entom. Soc. of Lond. ill. 1843,
p. 237, Ac.
Family XXII. Cympsea 8, Gallicolw. Posterior wings with
no nervure or one only, anterior with radial cell, and two or three
cubital cells, the second triangular, third incomplete produced to
apex of wing. Antennae of the same thickness, or gradually
thicker towards the apex, with twelve to fifteen joints. Maxillary
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384 CTJkss VIII.
palps four- or five-jointed, labial with two or three joints. Thorax
gibbous, with mesothorax very large. Abdomen compressed.
Borer extremely slender, with three set», concealed^ rolled spirally,
between a bivalve sheath, exsertile from the last ventral chan-
nelled segment of abdomen.
GaiUrwaapB. The females of this family pierce different parts of
plants (leaves, leaf-^talks, buds, &c) and lay an egg in the wound
The irritation thus produced causes the sap to flow in greater abun-
dance to the wounded part, and thus different excrescences, often of
very singular kinds, arise, which serve the larva both for food and
habitation. The form of the excrescences is different for different
species, and may serve for recognising and distinguishing them.
The larviB, bent into a semicircle, lie as thick white maggots
in the cavity of these excrescencea Some species undergo their
metamorphosis in this situation ; others leave it before becoming
nymphs, and change under ground It is true that species also of
ChcUcidea are found in these excrescences, which were formerly
placed with species of Gynips in one genus, and to which Gboffboy
gave the name of Gynipa exclusively, which occasioned much
confusion in the nomenclature : they are ichneumons which have
taken the place of the natural inliabitant&
Gall-wasps, although living upon vegetable food, have neverthe-
less a great affinity with the Ichneumanides, and this is shewn more
• distinctly by the fact that some species {Allotria Wkstw.) really
live like ichneumons in insects {Aphides), without on that account
differing from the rest of the Gynipides by natural characters or
organisation (Westwood, Introd, to modem GlassificaL of Ins, u.
p. 132, Ratzebubo, Die Forst-Insecten, nr. p. 54).
To the excrescences, caused by gall-wasps, belong also the gall-
nuts or gall-apples, of which those that come from the East (from
Aleppo) are in most esteem. They consist, besides gallic acid, in
great measure of tannin, and are consequently very astringent. Hence
their use in medicine. Their property of forming a black pre-
cipitate with salts of oxyde of iron, causes these gall-nuts to be
employed in the preparation of writing-ink.
Gomp. on this family : Malpiohius de Oallu, in Anatomea plantarum
parte aUerd {Operum ed. Londin. i686, foL Tomo ii. pp. 17 — 38);
OuviEB, Encycl. mtth^ Hist, not. de$ Int. v. 1790, pp. 771 — 792,
Brakdt u. Batzeburo, Medkin. Zoolog. u. 8. 144—758; Botbb db
F0N800LOMBX, Detcriptiim des In*, de la fam, dee Ih'plolSpairea qui «e
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IN8ECTA. 385
trowfoU aux mvirom tPAix, Ann. de$ Se, ntU. xxvi. 1833, pp. 184 —
198 ; J. O. WxsTWOOD, Imaior. nonntUlor, e familia Cynipidarum
de$enpUonei; QviBUX, Magas. de Zool. 1837, Ins. PL 179; Habtio,
Ueb. die Familie der OaUwetpm; Girmab'b ZeiUchr. /. d. Bntom. n.
1840. B. 176—109, ra. 1841. 8. 3«a— 358, IV. 1843, 8. 395— 4««-
Cyntps L, (excluaive of some species), Diplolepis Geoffk.
Sub-genera : AUatria Wbbt., (Xystm Habtio), AnaduM/ru Dalm. {Meg^k-
pdm/ui Habtio), Leiopteron Pbbtt, Wxstw., Paras Westw., Figites Latb.,
Biorhyza WiSTW. (Apophjfius Habtio), Cynips Latb., Wbbtw., Ihalia
Latb., and others; on which see WisrwooD, Oenerie Synops. pp. 55, 56,
and Habtio, 1. 1.
Sp. C^ips OaUce tindoria Olivixb, Voyage dans Vempirs Othoman, Paris an
9, AUas, PL 15, Bbakdt u. Ratzbbubo, Mediz. Zool, n. Tab. xxi. fig.
1 1 — 13 ; this species lives on Quercus infectoria in Asia Minor, Syria, &c. ;
— Oynips QuercusfoUi L., Row. Ins. iii. SuppL Tab. 5 a, 53, f. 10, 11; — Oyn.
jRosa L., R^UMUB, Ins. m. PL 46, fig. 5-7, PL 47, fig. 1-4. Blankaabt,
StA^nibwrg, Tab. 16, fig. v-s, Bbabdt u. Ratzbbubo, L L Tab. xxi. fig.
5-7, on the wild or garden-rose, in which it causes mossy excrescences
named Dog-rose-sponge or Bedeguar.
Note. — ^Abdomen in most extremely shoii-petiolated. Some are
distdngiushed by their long petiole (sub-genera Anacharis, Leiop-
teran, Ac) Males are distinguished from females by their small
size, longer antenme^ mostly also by the third joint of their antennie
being sinuated outwards. Of some species the females alone are
known*
Family XXIII. UrooercUa {Siricidm CuKTis). Abdomen sessile,
continuoufl with thorax, covering the origin of posterior feet, cylin-
drical or oblong. Mandibles short, thick. Wings both anterior
and posterior with distinct cells. Tibia of first pair of feet with a
single terminal spine. Borer of females in some exsert, straight,
with three setse, received between two homy valves, in others
capUlaiy, contorted at the base, contained in abdomen. Larvae
furnished with six feet, phytophagous.
The larvse of the chief genus of this division Svrex live in wood,
especially fir and pine, some also in beech, poplar and birch. Of
others the larvee are still unknown. Dahlbom suspects that the
larva of Orysnu lives on gall-nuts. The opinion of Spikola and
Lepeletisb that the larvsB live parasitically in those of wood-
eating insects, like ichneimions, is an error, and rests on imperfect
observation.
VOL. I. 25
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386 CLASS VIII.
Gomp. on this family : F. Klug, Mcnographia Siricwn Cfermania, Tab.
SBD. Berolini, 1803, 4to^ and (on the following also) the exoellent work of
Th. Habtio, I>ie Familien der BlaUtoeapen und Molewespm, Mit Abbild.
Berlin, 1837, 8vo.
Phalanx I. Oryssides. Borer capillary, incurved at base^ con-
cealed. Radial cell one, cubital cells two.
0rr/88ri8 Late. Antennae short, with ten to twelve joints, in-
serted near the mouth. Maxillary palps long, five-jointed, labial
triarticulate. Anterior tarsi of females with three joints only.
Sp. Oryuus coroncUua Fabb., Panzbb, DetOschl, /tm. "tieft 59, Tab. 19
{Sirex vespertUio), Dum^b. Com. gin, s, I, Iru, PI. 35, fig. 4 ; in Germany,
France, &c.
Phalanx II. Uroceridce. Borer straight, exsert. Badial cells
two, cubital four. AntennsB with eighteen to twenty-five joints.
a) Wi^ meuoSlary palpi Umg, 6- or s-joinied,
Cephtis Latr., Fabb. Antennae incrassated towards the apex.
Abdomen compressed.
Sp. OepkuB tpinipei, Banehui ipi/nipa Pahzeb, Deuttchl, In$, Heft 73, Tab.
17, Ac.
(Speciee small. This genuB is placed in the foUowing family by La-
tbbhiLB and Wbstwood).
XipAydria Latr., Fabr. Antennae attenuated towards the apex,
setaceous.
Sp. Xiphydria eamduSf Sirex eamdus h,, Duh&il, Chnt. gin, $. I. Int,
PI* 3<^> fig* ^» Habtiq 1. L Tab. vm. tg, g, &c.
h) WUk maaaUary paXpt extremely ekoH, with only one or twojoinie,
Sirex L. (exclusive of species). Uroc&rua Geoppr. Antennae
setaceous or filiform, long. Maxillae united at the base.
. Sp. Svrex gigas !»., Ichneumon gigas, Sytt. not, Ed. x. fern. {Sirex marieeuM
L. ma.), B(BBBL, Int. Bomb, el Veep. Tab. vm. tL,, DuMiBiL, Cone. gin. $.
I. Int. PI. 36, fig. I, Batzkbubg, Forat-Int. m. Tab. iv. fig. 3F; head
blacky with a large yellow spot on each side behind the eyes ; the male
with stone-coloured abdomen, the last two rings black ; the female has the
abdomen at the base and apex yellow, in the middle dull black. This
insect is the largest native hymenoptenim ; the expanded wings measure
. 1 in. 1 lin., the body i in. 3 lin., and the borer 4 lin. ; but much smaller
specimens of the species are met with. The larva lives more, than a year
in the wood before it changes into a nymph ; in summer the insect comes
to view from the pupa after three weeks, but when the larva becomes
a pupa towards winter, it con^ues thus throughout the winter.
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INSECTA. 387
Family XXIV. Tenthredineta b. Serrijera. Abdomen sessile,
continuous with thorax, covering the origin of posterior feet, cylin-
drical or oyato-oblong. Mandibles large, homj, acuminate, in*
curved, mostly tridentate. Maxillary palps mostly sexarticulate,
labials four-jointed or quadriarticulate. Labium cloven into three
laciniss. Wings both anterior and posterior furnished with dis-
tinct cells. Borer almost always occult, included in a bivalve
sheath, compressed, cultrate, mostly serrate, composed usually of
four setsB (the upper one of other hymenoptera being here cloven to
the base). Tibise of anterior feet with two terminal spines. Larvffi
(similar to caterpillars) with mostly twenty-two or twenty feet,
feeding on leaves.
Leaf'waapa. The larvse mostly eat leaves like caterpiilars, some
live in gaU-exGrescences. These insects are often very destructive to
trees, and the knowledge of them is therefore veiy important to the
forester. Some also injure our potherbs.
On this family, besides the Monograph of Habtig Doted above and the
third part of the Fortt-Inncten of Batzebubo, may be oonsulted : Klug,
Die BlaUtoeipen der Fabrmschen SamnUung; Whdexann's Zoologiichei
Magazin, I. 3, 18 19, s. 84— 91, Tab. n., and by the same, Uebtrsickt der
TenihredindCB der {Berliner) Sammlung, in his JahrbUeker der Ituektet^
hunde, I. Bd. 1834, Svo, s. 133—253, Taf. n. figs. 5 — 10 ; 6. Dahlbom,
Clavia novi ffymenopterorum wytUmatia adjecta tynopti lairvaTum tocmdinoir
viccur, ertiq/brmttim, Lundn, 1835.
A. Borer exsert
Xyela Dalmann, Mastigocera Klug. Antennas thirteen-jointed,
with fourth joint longest (equalling or surpassing in length the
nine terminad jomts taken together). Borer of females of the
length of abdomen.
B. Borer occidt.
a) AntennsB with numerous joints^ (fifteen to thirty-six).
Lyda Fabe., Hartig, Pamphilivs Latr. Antennae setaceous
(nineteen- to thirty-six-jointed). Badial cells two, cubital four.
Posterior tibiae with three lateral spmes.
The larvn of this genus live together socially in a -web ; they have,
besides the riz homy feet on the thorado segments, only two propellen
directed outwards at the hind part of the body. Comp. Habtig 1.1.
Tab. VII. figs. I— 16, and Ratzbbubg, Forst-Intekten, m. Tib. i.
25—2
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388 CLASS vni.
Tarpa Fabr., Hartiq^ MdgalodonJtes Latr. Antennas shortly
pectinate on the inside (fifteen- to eighteen-jointed). lUdial cells
two, cubital four. Posterior tibiae with two lateral spines.
Comp. Kluo, EnUmol, Manogtriphien, Berlin 1824, 8yo, 8. 181 — 10.
Lophyrus Latr. Antennse in males pectinate, in females ser-
rate (seventeen- to thirty-jointed). KadiaJ cell single, cubital cells
four.
Sp. lophyrus Pini, Tentkredo Pini L., Batzebubo L I Tah. ii. fig. i ;
Lophfr, rvfuB, ftc. The larvsa have 11 feet^.
5) Antennae with mostly nine joints or fewer.
Tenthedro L. (exclusive of many species), Latr., AUantus
JURINE (with addition of some sub-genera). Antennae nine- to
eleven-jointed, simple. Badial cells mostly two, cubital four.
Labrum exsert.
Sub-genera: Macrophya Dahlb., TeTUhedro Habtig, AUcmiius
JuBiNE, Athalia Leach, SeUmdria Leach, Dmura Dahlb., Fhyl-
lotoma Fallen, Fenusa Leach, Emphytus Kluo, FdnuUopua
Hartio, Dclerua Jurine, Cryptocampiu Hartio, Nematua Jubine,
Clctditis Illio., and others which are recorded in Westwood,
Generic Synops. pp. 52 — 54.
(Areokd of wings and antenn» are used for subdivisions).
Sp. TefUkredo difarmit {Cladiua), Pakzxb, IkvUch. Ina, Heft 61, Tah, x.
(male with antenxuB pectinftte in middle) ; Tenthredo cenHfolia {Athalia),
Pakzbb, Deuisch. Ina. Heft 49, Tab. xvni.* ; TeiUhr, groaatdarice Dahlb.,
Blakkaabt, SchoubuiTff, Tab. n. figs. G — J ; Tenthredo gaUioola (Nematua)
WxsTW., SWAIOOBDAK, £ib, Nat. Tab. xuv., Bobsel, Ina. IL, Bombyl.
et Veap. Tib. x. figs, x— 4 {Ten^redo ffoUifex Haoxub. in MSa) fta The
lame have twenty or twenty-two feet.
Hylotoma Latr. (and Schtzocera ejusd.) Antennas triarticulate,
with third joint elongate, in males of some species forked {Schizo-
^ Comp. on thiB genus L. FumELMAKN, Zur Natwrgeaeh. einiger auf der KUftr
lAender Lophyren ; Nov. Ad. Ocea. Loop. Oar., Tom. xix. P. i, 1839, PP- *45— «8o*
Tab. XXY.
* On this insect^ whose Larva may be very destructive to turnips by eating the
young leaf, we have an excellent monograph by G. Nbwpobt, Obaerv. on the Anai.
and Economy of Athalia eenHfolicB; Prise Saaay of the Fniomoi. Soc. With a plate.
London, 1838, 8vo.
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INSECTA. 389
cera). Radial cell one, sometimes with accessory apical cell ; cubi-
tal cells four or three {Plilia Lepelet.)
&p, HylcUma rotarum Fabb. (not Tentkr,' ro§cB L.) BosiL, n. Bomb^ior,
€t Veapar, Tab. n., DumjIbil, Cont. g^n, «. L Ins. PL 35, fig. 6, Ac.
{Tenihr. rota L. is a speoies of Athalia.)
Cimbex Oiiv. Antennre clavate, five- to seven-jointed. Badial
cells two, cubital three.
Sp. Cimbex variabilis Klvq, Tenthredo lutea L. (and Tenthr, ftmarata
ejtud.), K(E8EL, Ins. u., Bomb, el Verp. Tab. zm., Batzebubo, Font-Ins.
lu. Tab III. fig. 10, &c. These species are the largest of this iamily ; the
larrn have twenty-two feet. The larva of Cimbex Uieorwn was not long
ago described and figured by Skxllxn yah YoLLEKBOTiir, Tijdsckr.
voor not. Gesch. 1. 1843. Tab. n.
Sub-genera : Abia Leach, Perga Leach, and others of this
author. Add sub-genus Pachylosticta S^UG, genus Syzygonia
ejusd. differing from all other cimbices in the cells of the wings,
but plainly resembling the Hyhtomam (Brazilian species.)
Okdek Vni. Lepidoptera.
Hexapod insects, with four membranous wings, covered with
minute coloured scales. Mouth with involute spiral tongue, com-
posed of protracted maxUlaa. Metamorphosis complete.
BvUerflies {Lepidaptera L., firom XfirU scale, and irrepov, Glossata
Fabr) The two chief works on the anatomy of this order, that of
Ltonet and of Herold, have been already cited (see above, pp. 247
and 275). To give a list approaching to completeness of the works
which treat of the arrangement of butterflies, or illustrate their
species by figures, would require too much space for our purpose.
We satisfy ourselves therefore with indicating some of the prin-
cipal sources for the knowledge of this order.
J. C. Sepf, Beachouwing der toanderen Gods in de minst geachte
Sch^Mden, of Nederlcmdsche Insekten, &c 4ta Amsterdam, 1765, and
folL Of this work, which is still being continued, 6 parts, each of
50 plates, have hitherto appeared.
P. Cbaheb, UUlandsche Kapellen, iv. parts, and Stoll*s Aankcmg-
sd, 4to. Amsterdam en Utrecht, 1779 — 1791. (With this may be
usefully consulted the academic prize-treatise of H. Yebloben,
CcUalogw systematicus ad CRAXERUMy Traj. ad Bhen. 1837, 8vo.)
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390 CLASS VIII.
SystemaHaches Verzeiehniss der SchmeUedmge der Wienergegend
herausgegeb, von einigen Lehrem am K. K. Theresiantim, Wien.
1776, 4to. (mit 3 col. Ta£)
Jac. HtJBNEB, Sammdung ewropdischer Schmetterlinge, nebat Fori-
aetzwng von C. Geter, gr. 4to. Augsburg, 1806 — 1841 ; — hj the
same, Geschichte ewropdiacher Schmetterlinge (Rcmpen, Fwppefn u,
FvMerpflcmzeTi)^. 4to. ibid. 1806 — 1841 ; — ^by the same, Sammhing
exotiacher Schmetterlinge, nebat Fortaetxung van C. Geyer, ill Bde,
gr. 4to. ibid. 1806 — 1841; and Zutrdge zwr •Samrrdung exotiacher
Schmetterlinge, ibid. 1818 — 1837. (I have not been able to consult
these comprehensive and costly works whilst preparing this order).
F. OcHSENHEiMEB, Die Sclvmetterlinge von Ewropa,fortgeaetzt durcJ^
Fb. Tbeitschke, jl Bde, 8va Leipzig, 1807 — 1835.
BoiSDUVAL, Speciea ghiercd dea Lepidopt^ea, Tom. i. av. pL Parisy
1836, 8vo. This excellent work, which makes a part of the well-
known SuUea dL Buffon, published by Bobet, appears to be discon-
tinued, to the great injury of science.
The scales, which cover the wings of these insects on both sides,
appear to the naked eye as dust, but when seen through the
microscope, are arranged in regular rows, like house-tiles ^ These
scales are implanted, by means of little pedicles, in short conical
tubules, whose openings are constantly directed to that margin
of the wing which is opposite to its base. Each scale consists
of two (or perhaps even of three) membranes or layer& On the
uppermost membrane lie granules of colouring matter. Elon-
gate, parallel stripes (ribs) run from the base to the free extremity,
which has sometimes a smooth margin, and sometimes ends in
certain points or lappets. The underside of the scale, which lies
next the wing, often presents a play of various colours*. When
the scales are removed, the wings are whitish and semi-transpa-
rent ; some butterflies have constantly such patches on the wings
where the scales are wanting ; in some the wings are almost quite
naked, whether because the scales are wanting from the first,
^ Numerous figures of this are to be seen in the works of microscopists, as
in Leeowenhobok, Derde vervclg van JBrieven, ye Missive, 24 Junij, 1691, bL 409,
fig. I ; see also Roiskl, Int. x. Tab. n. f. 5, 6, 7, Pap, PodaUriut, in. Tab. xuv.,
Pap. Iris, Ac.
* Bebnabd-Dbschamps, Recherches Microacopiquea sur Vorganisation dea aiUs
dans les LSpidopUres, An. dea tc. Nat., sec. s^rie iii. 1835. Zoologie, pp. iii — 137,
PI. 3, 4.
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INSECTA. 391
or becaoBO beixig fixed very loosely, they are lost on the first flighty
as has been obsenred in Sphiryx Jv4:ifor7M8.
Amongst the oral organs (compare above^ p. 249) is seen a small
upper-lip, often scarcely visible, which is of a triangular or conical
form ; the mandibles are small, immoveable, and remote firom each
other. These parts exist in a rudimentary state aJone. The prin*
cipal part of the organa ciba/ria is the spind tongue, formed of two
long threads (the maxiUai), which are excavated on the inside, else-
where even, and run out to a fine extremity; when these two
plates are laid towards each other, the two half canals form a com-
plete canal in the axis of the tongue. At the base of the maxilLe
are placed two very short paJps, consisting of one or two, at most of
three, joints. The under-lip (IMam) is triangular and flat, and
bears two large palps, which consist of three joints ; these palps are
covered with hairs, and mount with their points turned upwards on
both sides of the tongue, which rolled up in a spiral lies hidden
between them, whenever it is not in usa In some nocturnal lepi-
doptera the tongue is very short and not adapted for sucking.
The antennse of these insects differ in form, but always consist of
numerous joints. The two compound eyes are lai^e ; in many
species there are in addition two simple eyes present. The three
rings of the thorax are always connected immoveably with each
other; the middle piece {mesotharax) is the largest. The wings
are large and not folded ; in the females of some species they are
reduced to small rudiments, or are entu*ely wanting. There are five
joints in the tareua of all the feet. The abdomen consbts of six or
seven rings, without sting or borer, as in the preceding order.
The metamorphosis is complete. The larvse are called caterpUlara
(eruccBy chenillesy Bcvupen, rupsen). The body of caterpillars con-
sists of twelve rings exclusive of the head. There are on each side
nine air-slits; for the second, third and last ring are without them.
The normal number of feet in caterpillars is eight pairs ; the fourth,
fifth, tenth and eleventh ring have no feet. On the first three rings
three homy feet axe placed, which have a conical form, and consist
of joints; the last joint has the form of a bent nail These
six feet answer to those of the perfect insect. The remaining ten
feet (some species have only eight, six, or four) are membranous
and without joints ; they disappear in the perfect insect. On the
underside they have a flat sur&ce, which the insect is able to expand
and contract, and which is surrounded by a coronet of numerous
small hooks. The head is homy, and has six simple eyes on each side;
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392 CLASS VIIT.
moreover, there may be distinguished in it two short conical an-
tennae, two strong mandibles, two maxill» with small palps, and an
under-lip, which also has two small palps and terminates in a point,
under which the efferent canal of the matter with which the cater-
pillar prepares its web is situated. This substance is secreted as a
fluid by two long, blind, convoluted vessels, which lie at the sides
of the intestinal canaL Most caterpillars live on vegetable food,
especially leaves, and many are limited to a single species of vege-
tabla Others, however, eat leather, fur, fat, wax, &c., and these
belong especially to the family of the moths. Caterpillars usually
change the skin four or five times before turning into pupee.
The pupae of scaly-winged insects axe quiescent, and move their
abdomen alone when they are touched. They are oblong-ovate, and
covered with a homy skin {pupcs obtec'as, see above, p. 273). The
pupee of day-butterflies are usually not inclosed in a web, but merely
attached by some threads at their posterior extremity, and hang
freely with the head downward, or are fixed transversely to a branch,
or other object, by a ti*ansverse band, as if in a hoop. The pupes
of nocturnal butterflies either lie underground in a cavity that is
smooth and even within, and lined with web, or they are inclosed
in a cocoon (/oUiculus), which is fastened to a branch, or to a walL
The web is frequently silken, sometimes very closely woven, some-
times loosely ; sometimes it consists in part of finely gnawed fibres
of wood interwoven with the threads of web, or of other foreign
objects intermixed with the web, crumbs of earth, morsels of leaves,
<fec. These pupee have commonly a brown or black colour.
From the pupa of many species, especially of day-butterflies, the
perfect insect proceeds after the lapse of a few da3rs. Of such
species there are ordinarily two generations in a yeai*. Of other
species, however, the caterpillar or the pupa remains through the
winter, and then the perfect insect usually appears only once in the
year, in spring or in summer. Eggs that are laid in autumn are
mostly hatched in the following spring.
The intestinal canal of caterpillars is straight, and consists in
great measure of a wide cylindrical stomach. There are four very
long vessels for secretion of urina The perfect insect has a narrow
oesophagus with a lateral expansion or crop (the so-called sucking
bladder, see above, p. 310) ; the stomach has become shorter, the
rest of the intestinal canal longer. Lepidopterous insects in the
perfect state of butterflies either take no food at all, or suck the
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INSECTA. 393
sap of flowers ; they lick this up by means of their maxillip, pro-
longed into a tongua
This order is ^eiy numerous in species. Amongst these are many
which attract our eye by their brilliant colours, or the beautiful
design of the spots or streaks on the wings. It is as though nature
had given such large wings to butterflies in order to secure a wider
space for her pencil
Family XXV. Noctuma. Antennae setaceous or pectinate.
Wings horizontal for the most part or deflected, guarded by a
retinaculum, with few exceptions. Posterior tibise with double
internal spine.
NfxAivmal Lepidoptera, Moths, The species of this and of the
following family are distinguished by the so-called retinaculwnf
which they mostly possess. This part consists of a homy highly-
elastic hair, or of a little bundle of two or more hairs, which arise
on the anterior margin of the hind-wings close to their insertion.
A little flat ling on the undernsurface of the fore-wing allows it a
passage, and thus both wings are oonneeted, and similarly pressed
forwards and backwards, and in flight form only a single sur&ce^
Almost all these moths fly by night alone, or after sunset. The
females of some species are wingless, or have only minute rudiments
of wings (as Phaltena brumata, Bomhyx arUiqua, &a) The form of
the larva is various, and they have from 10 to 16 feet. Most of
them make themselves a web, in which they change to pupse.
This &mLly in the system of LiNViBUS forms only a single genus,
which he names PhdliBna.
Pterophorua Geofpr.. Fabr, Four wings or two posterior cloven,
with fringed digitations. Antennae long, setaceous. Body slender.
Feet elongate.
Sub-genus Fterophorua Latb. Labial palps smalL
Sp. Plavphortu peniadctctyhu, PhdUmia (AlueiUi) peniadadyla L. BcBSlL,
Ifu, I. C1m8 17. Pap, noetwm. Tab. v. Ac. The caterpillars of this species
have sixteen feet, are broad and hairy, and change without spinning them-
selves in ; the pupee hang by threads, like those of day-butterflies. The
perfect insect sits with outspread wings, and reminds us of TipuUt,
^ I have treated specially of this part, and figured it in the Naiuurh Bijdragen,
by H. C. Vak Hall, W. Vbolik and G. J. Muldeb, ii. 1817, bl. i73--«84.
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394 CLASS YIII*
Sab-genua Omeodea Latr. Labial palpB longer than head, with
seoond joint very scaly, the last somewhat nakeii, erect.
Sp. Pterophonu hexadactsfhu, Phakena {AlticUa) kexadadyla It., Bl&UMUB,
Ins, I. PI. 19, figs. 19— « I, DuMiBiL, Cons, gin, s, I, Ins, PL 43, fig. 8.
The sixteen-footed naked caterpillar feeds on the blossoms of KamperfoUa
(Lonicera), and spins itself up when about to change into a pupa. The per-
fect insect has each wing divided into six slips, and keeps the wings when
at rest deflected.
Tinea Fabe. (and Aludta ejnsd.) Anterior wings narrow^ or
horizontal, incumbent, or convolute round the body or deflected and
erect backwards. Labial palps erect, maxillaiy palps often distinct.
Antennae setaceous.
A. Labialpalps skoH, pilose. Antenna, especiaUy <;ff males, mtoeUf very
long, approximale, Eges subeontiguous posteriorly.
Sub-genus Ackla Latb. {AhusUa Ykbbl in part).
Sp. Tinea Degeerella li,, Db Gksb, Ins.i. PL 3a, fig. 13; Gu^Riir, Iconogr.
Ins, PL 91, fig. 7; FiB(msXi*a Ahifildungenzur Microl^idopierologie, 1854,
Tab. 66, 67, figs, a — e; P. Ltonbt, Ouvrage posUiume, PI. 19, figs. 17 —
95; wings gold brown with a yellow band on the fore-wing, which is
margined with violet. Expanded wings about 8"', aatenntB of male 14'"
long. In the pupa the antenna have special cases that project behind the
body.
B. Labial pcJps moderaU, very dMnct, EyesaandaadennartmUe.
a) Tongtte disLinct, dongaie.
Sub-genera (Ecophora Latb., Tponomeuia Latb., IlUkyia Latb.,
Omix Tbeitsohke.
Sp. THnea evonymeUa L. (Tponomeuta evon,), Rgssbl, Itis. i. Pap, TMetum.
Gl. TV. Tab. vin; the caterpillars live socially in a web, as do those of
Yponomeuia cognateUa, R(BS. 1. 1. Tab. vii. Ssfp vi. Tab. xxvn, that live
on thorn-bushes and fruit-trees, and also, what is often confounded with
it, TSn. padella L., Ssfp y. Tab. zxxn. whose caterpillar lives on wil-
low-trees.
6) Tongue very skorL {Cred on the head of Havre or scales,)
Sub-genus Tinea Latb.
Sp. Tinea pellioneUa L., Bobsbl, Ins. I. Pap, noctum, 01. TV. Tab. xyil the
fur-moth; shining brown upper wings with a black spot ; it shews itself in
the spring; according to TasiraoHKi there are two generations in a year;
— Tinea grandla L., Hoebel, ibid. Tab. xu. &c.
Crambm Fabr. Fore-wings narrow, much longer than broad.
Maxillary or upper palps very distinct, porrect above the base of
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IN8ECTA. 396
labial palps, conpexed with scales oar hairs. AntemuB mostlj simple.
(Caterpillars with sixteen feet).
a) ToDgae veiy short, scarcely distinct.
Sub-genus Phycia Fabr. {Phycis and Euplocoimus Latb., with
antenniB of males pectinate).
V) Tongue distinct.
Sub-genera OnvmJbua {Chiio Sommeb), AlucUa Latb.^ GaUeria
Fabb.
Sp. GaUeria certiUa, Tinea meUoneUa L., Bobsel, /tm. m. Pap, nod. CL TV,
Tab. XLi. Sbfp y. Tab. zlyi; with grey wings, the upper wings more
brownish, dark-spotted and excised on the outer margin in the male, in the
female obtuse. The caterpillar of this species lives in the hives of bees,
always protected and concealed under spun threads. The metamorphosis
occurs within the hive. The moths come from the pupa in the spring.
These enemies of bees were known to the ancients; YiBOiL calls them
dirvm tinea genu* (Qtorgic, Lib. iv. 346; comp. Plinius, Hid. nai. Lib.
XI. cap. xiz.)
Botya Latr. (and Hydrocampe ejusd.) Fore-wings triangular,
forming with the body a sub-horizontal triangle when the insect is
at rest. Four exsert palps. Tongue conspicuous. Antennae seta-
ceous.
Sp. Buys verticalis, PhaUma {Pyralis) wrticalia L., BofiXL, Ini, X. Pap,
nod, CI. IV. Tab. iv. Skpp v. Tab. xxiv. &c.
Aglossa Latb. Tongue inconspicuoua Habit and characters of
the preceding genus. (Species of genus Pyralis Tbeitschke, Cromr
hua'FjLEBL)
Sp. A^ana pinguinalis, PhdUma (Pyralia) pinguinalie L., De Gbsb, Im.
u. PI. VI. figs. 4— n, Skpp v. Tab. xx, &c
Ibrerw? Treitschke, Pyralis Fabr,, Latr. [Phalcena Tortrixlj.)
Wing of insect at rest representing the form of a roof much flattened
or subhorizontal, and with the body forming a triangle, short, broad,
anteriorily arcuate outwards, the external margin of the fore-wings
being produced to the base. Maxillary palps either none or short,
not exsert; labial palps with second joint thick, hirsute, in some
short, in others longer and in the anterior part of the head, pro-
duced like a beak. (Caterpillars with sixteen feet.)
Leaf-rollers. These have been thus named because the caterpillars
of many species roU up and spin together the leaves on which they
feed. Some tie up young buds and blossoms with their web; others
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396 CLASS VIII.
again live in fruits, as Tinea ponumdla L. (the genus Gao'pocapaa
Treitschke).
Sub-genera Cochylisy TeraSy Ca/rpaca/paa, ffcUiaay Pofdisca, Tbeit-
SCHKE and others, on which see Schmetterlmge van JSwropa, Tom.
VUL, and Westwood, Generic. Synopeis, pp. 106—110.
Sp. Totirix dUorana L.^ (ffcdiat Tbkitbohkx) Dum^ Ommd. ff^. s, 2. In$.
PI. 53, fig. 6, Skpp vl Tab. xin ; — Totirix vitana, Tortrix peUeriana,
Sydem. Verz. d. SchmeU. d, Wienergeg, p. ia6, Pyrale de la Vigne Boso.
This species, which causes great injuiy to vines, and so, in France especially,
from time to time produces serious damage, is the chief sul^ect of an exten-
sive and excellent work of Axtdouik, Sid, det Insectet nuinbUi d la Vigne,
Paris, 1843, 4(0.
Tortrix pomana, Tinea pomonella L., Rcbbkl, /fit. I. Pap, nod. CL iv.
Tab. xui. Sepp yi. Tab. x. &c
Herminia Latr., Hypena Schrank. Wings triangular, snb-
horizontal, deflected, forming with the body a triangle when the
insect is seated ; the anterior sub-falcate at the apex, with posterior
margin convex. Labial palps longer than head, compressed, with
last joint recurved. Ocelli two. Antennae of males ciliated or
sub-pectinate. (Caterpillars with fourteen feet).
Sp. Herminia proboeeidalie, Phalcena {PyraUe) proboteidaUa L., KuEEXANir,
Beytragt eu B<esil's In$. Tab. xxxn. Skpp n. 5e Stuk, Tab. n. the
brown snout-moth; on the sUnging-nettle; — fferm. rodralie, Phal. {Py-
ralie) roetraUe L., Rcbbel, In». 1. Pap. nod. 01. iv. Tab. VL ; on the
hop, also on stinging-nettles, &c. The caterpillars of these spedes have no
feet on the sixth ring of the body ; when touched they let themselves fall
to the ground, and leap like fishes drawn out of water.
Note. — Genus HyhUea Fabr. is joined with Herminia by La-
TREiLLE ; it contains exotic species, which whether they be all rightly
placed here, appears to me very doubtfid. ffyhkea litwrata Fabr.
from the Cape of Good Hope, Nabmforsch. xxix. Tab. iv. fig. 14,
appears to agree entirely with Herminia ; here also are to be referred
PJudcencR Servia and Sergilia Cram.
On genus ffyhkea oomp. Esfkb in Natturfor§eher, xxix. i8o3, pp. 191 —
ao6, Tab. ly.
Pkalcena Fabr. {Phakena geometrce L.) Wings broad, mostly
patent when the insect is at rest, with anterior only partly covering
posterior, sometimes erect. Antennss elongate, towards the extre-
mity attenuated, setaceous, or in males pectinate. Labial palps
moderate. Ocelli none. Body attenuated. Most of the caterpillars
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INSECTA. 397
with only ten feet, the rest with twelve, always with anal feet.
Pupa inclosed in a thin follicle.
The genus Fhaicsna or Geametra is principally characterised by
the caterpillars ; since these have no feet in the middle of the body
they move in creeping, as leeches do, by flexion and extension of the
body. Hence these caterpillai's are named swrveyora or sparircaler'
pilkbrs {eruccR geometrical, chenilles wrperUeiueSy Spanner,)
Comp. on this genus Tbhtbohki especially, JXe SehmeUerUngen von
Europa, of which the entire sixth part (Leipz. 1827, 1818) is set apart for
treating of the nomerous European species of this genus ; and further
WaxLCB-BcHMFTSB,, U^bentckt der Spanner, in his continuation of Paitzkr,
/«». Deutackl, Heft 165, 176, 179.
LnmiBUS has made a subdivision according to the antennse ; where they
are pectinate he gives to the specific name the termination -aria (as pini-
aria), when they are setaceous that of -ata (as ffrotndariata^). The cha-
racteristic of pectinate antennn, however, besides that it is proper to the
male alone, occasionaUy separates naturally allied species. As little can
the separation of these species, which have wingless females, of which
liATBBniUB fonns his sub-genus Hybemia, be commended.
a) CaterpiOars with twelve feet
EUopia Tbeitschke. (Antenna in males pectinate.)
Sp. PhaUma nuurfforitaria Fab., Pkakena fiuprgaritcia L. fern., Sbfp, Nederl,
I'M, n. NaehtU, n. Bende i Gezin, Tab. m., Paitz. DevtteU. In$. Heft 41,
Tab. 43, Heft 63, Tab. 13 ; %ht-green wings, the fore with two white
bands, the hinder with one only, which is a continuation of the most
external of the fore-wing. Amongst the Noet/WB also some species occur
with twelve-footed caterpillars, which however are not spanners.
h) CsterpiUars with ten feet.
£nnamos, Actena, Geametra, AspilaUs, CrocaUis, Gnophos, Boar-
mia, Amphidaeie, Psodos, Fidania, Cheeias, Cabera, Acidalia,
Larentia, Cida/ria, Zerene, Minoe, Idcea Tbeitschke (a name to
be rejected siace already given by Fabbioius to a genus of the
Dinmals). Ck>mp. on other genera^ here omitted, Westwood, Generic.
Sf/nops. pp. 98—104.
[Note, — ^The numerous species of PhakenoB are distinguished by variety
of form, and often present an analogy truly wonderful with other genera
^ Here we have an instance of the happy mnemotechnie of which LiKK^us in all
his writings made such rational use. The distinction however given by him is not in
every instance well-founded; Phal. tatnlucaria, for example, ought properly to be
etHikd iombueata.
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CLASS VIII.
of Lepidopttra, bo tha>t there may be obeenred, prinoipAlly amoDg the
exotic genen^ some that recall the genus Pieria, others PapUionet (or
Equites), not in habit alone, bat also in colour. Perhaps a parallel series
is formed by the Phalamce, comparable with most of the genera of diurnal
Lepidopiem. Other exotic Phakena approach more nearly to the genus
Urania, but yet amongst European species Phakena tambucaria presents
an analogy with it.]
8p. PhaUma hdtdaria L., Amphidoiii hettdaria Tbeftsohki, Skfp ii. 4e
Btuk, Tab. xxi. the hlaekiprinkled moth, Pakz. Ikutschl, Int. Heft 31,
Tab. 14; with long, small, round wings, outspread, 1" broad, length of body
usually 9"% the abdomen thicker and more unwieldy than in most species
of this division ; wings and body yellowish- white, with many black spots
and points ; — Phal. aanUmearia L., Aocena Mombucaria TaKxracHKE, Oum-
pteryx Mombucaria Lkaoh, Romkl, In9, I. Pap. Nodum, CL m. Tab. Yi.
Skfp, Nederl. Iru, i. 6e Stuk, Tab. i., one of the largest European species,
but of a totally different form, with broad wings, the anterior falcate at
the apex, the posterior excised at the margin with obtuse angles, of which
angles the third is produced into a short tail ; the general colour pale
sulphur-yellow, with two light brown bands on the fore-wings, and one
similar on the hind-wings, which is a continuation of the innennost
of the former. The eggs are prettily ribbed ; the caterpillar is a true
spanner, of a brown colour, resembling a dead twig. Phal. drfoliaria L.,
Pidonia dtfoHa/ria Tbxitbohks, Bcbskl in. Tab. xir. (the metamor-
phosis and the wingless female). Tab. XL. fig. 6 (the perfect male), SlFP,
NedM. I'M. n. 6 Stuk, Tab. vi. Batzibubo, Pont-InM. ni. Tab. zi.
fig. 5, Ac.
Platypteryx Lespetbes, Ochsenh., Drepana SCHRANK. Wings,
the insect at rest, patent, anterior broad, in some rotundate, in most
falcate. Palps short. Antennss in males pectinate, in females
setaceous or serrate or Tery shortly pectinate. Ocelli none. Cater-
pillars with fourteen feet, terminated by apex acute, erect, the anal
feet wanting.
8p. Platypteryx faievla, Phal. {Oeomdra) falcataria L., Ltoitbt, Oumrage
potth. PI. 35, figs. 6— 10 ; Plaiypt. hamtda, Phal. ftdeala Fabb., Skfp,
Nederl. Int. n. 40 Stuk, Tab. xyl The one4aUed eaterpiUara resemble in
some degree in miniature the two-tailed caterpillars {Bcmbyx vinula,
fwrculaj) whence some writers have placed them with the Bombyees
( VeneichniMt der Sehmdier. der Wiener gegend, p. 64, Hubkbb, Latbeillb) ;
LnfVJBXTS and Fabbioius, giving their attention exclusively to the per-
fect insect, placed the species known to them amongst the PhdUeiUB
geomdrce, witii which indeed they have a greater affinity. The point in
which the body of the caterpillar terminates behind forms a supemumeraiy
segment (a thirteenth ring), which represents the two hind-feet that are
wanting.
Noctiia Fabb. Tongue distinct. Palps in most moderate, with
third terminal joint more slender than the preceding or small.
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INSECTA. 399
Antennae mostly setaceous, in males of some species only pectinate
or ciliate. Ocelli, with a few exceptions, two. Wings apt for flight,
the anterior mostly triangular, deflected or incumbent. Thorax
large, very often crested. Abdomen elongato-conical. Larvae
solitary, mostly naked or seldom pilose, never without anal feet,
most with sixteen, some with only twelve feet. Pupa inclosed
in a follicle often lax.
a) L€Ut joint ofpalpa shorter than Mecand, scaly,
* CaterpxHoTS wiik twlvtfeet,
Euclidia OcHSENH. Palps short. Head small. Body slender,
with thorax not crested. (Anterior wings variegated with irregular
brown markings.)
Sp. Noctua Mi Im, Bspb, NederL Im. n. 50 Stuk^ Tkb. i., Ltonst, Ouvr.
poethwne, PL 39, figs, xo — 17 ; the caterfMllar lives on different Bpecies of
grass, on dover, &c,, and spins itself up between the leaves or in moss
when about to change into a pupa.
Plusia OCHSENH. Palps long. Thorax crested. (The fore-
wings often with spots or marks that shine with metallic splendour.)
Nochta gamma L., Basii^ Ins, 1., Pap, nodwm, CL m. Tab. T. Skfp,
Nederl. Ins, i. 50 Stuk, Tab. i. f. i — 6 ; the ffamma-moth ; body grey, fore-
wings reddish-grey with many stripes, in the middle and at the base dark-
brown, with a yeUowiah shining spot resembling the Ghi^eek letter 7, the
hind-wings yellowish-grey, with a blackish broad border ; length of the
body 3"' ; breadth of npper wings expanded 15'". The caterpillar of
this species sometimes occasions mnch damage to flax, hemp, cole-seed,
peas, pnlse, and all sorts of potherbs ; a visitation which afflicted different
districts of France in 1735, <^<^ ^^ province of Groningan in East-
Prussia in 1843 : see J. Jaoobson de Phal, noetua gamma Diss, Be-
giomonti, 1829, 8vo, and H. G. yait Hall, Oesch. van de verwoestingen
door de rupsen in het jaar 1829 aangerigt, Groningen, 1899, 8vo.
** CaterpiUarsmih sixteen feet.
Sub-genera: Brephos Ochsekh.^ Catocala Schrakk, Ochsenh.,
Ophiusa OcBSEtra,, Anarta Ochsenh.^ GucuUia Schrans, Ochsenh.
Noetua nob.
(Xylvna, Cerastis, Cosmia Hubbn., Xanthia HiriBN., Oortyna, Non-
agria, Leueania, Simyra, Caradrina, Orthoeia, Mythimna, Calpe, Thya-
tifrcky Mamestra, Apamea, Trachea, PUia Oohbenh., Misdia Hubbk.,
OcHSBHH., Badena Sohbahk, Mama Tbbit8GHKB, Amphipyra, Tri-
phono, Oraphipkora, Agrostis Ooebbith.)
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400 CLASS VIII.
JSpiaema Och8EKH.| Cymatophara Treitschke {Tethea Ochhemh.),
BryophUa Tbeitschke {Fcecilia Schbank, Ochsenh.), Diphthera
HuEBN.i OcHSSNH., AcTonycta Ochsenh.
Comp. Eneyd, miih., Hui. not., Ins, Tom. viiL 1811, pp. 21% — 360
(article NoctuelU), Tbkitsohkz, SchmeUeriinge von Europa, Yter Bd. 1825,
1826, WssTWOODy Oeneric Synoptis, pp. 93 — 98, &c.
Sp. Noetua nupia L., Sefp, Nedcrl, Ins, i. 46 Stuk, Tab. vn., Ltonbt
Ouvr, podk, PI. 45 ; body i" %'*' long, points of fore- wings in flight
distant 7," g"' or more from each other ; fore-wings above grey, passing
into bluish-green, with dark-brown flamed stripes, beneath white, with
three black bands; under-wings above carmine-red, with two broad
black bands, which are present on the under-side also, but on a white
ground, which towards the inner mai*gin passes into red. The cateipiUar
lives on the willow. A still larger, and with us a rarer species, has on the
under-wing a blue band on a bktck ground : Noctua fraxini L., B(B8EL,
Int. IV. Tab. «8, fig. i, Sbpp, Nederl, Ins, i. 40 St. Tab. xvin— xx.
— NoOua pronuba L., Triphama pronviba Tbbitbghke, Rcbsbl, Ins. iv.
Tab. 3a, fig. 6, Vkbhubll in Sbfp, Nederl. Ins. vi. Tab. 34, figs. 7, 9 ;
the upper-wings brown, dnnamon-coloured of di£Perent shades ; the under-
wings yellow, with black band dose along the maigin ; breadth of wings in
flight 7", length of body about 10'". A vezy common species, which in
the middle of summer is often seen in houses, being attracted towards
evening by the light. A lighter prothorax distinguishes it fitun Noftua
(Triphana) inwuha Tbeitschke, where the thorax is of a single colour;
this last has the fore-wings less flammate, often almost entirely of one
colour, and ordinarily a light coffee-colour {ettfi au laU) ; Bcbsbl, ibid.
figs- T> ^t 4> 5, Sepp Tab. 33, fig. 6, Tab. 34, figs. 8, 10. Livnjbub
united both spedee under his Noctua pronuba, and perhaps they are only
varieties. — Noctua piniperda, SVaehea piniperda Tbeitbohkb, Paitzeb,
DeuUchL Ins. Heft 83, Tab. 94, {Bombyx sprda Fabb., and Noctua
JUmmea, ^usd.,) Sepp, Nederl, Ins. in. Tab. 34, Batzebubo, Fortt-Ins,
II. Tab. X. fig. 4 ; half an inch long, flight 15 to 16'" broad, upper-
wings brown-red and grey, with two whitish spots in the middle, hind-
wings dark-grey, under-side of wings single shade of grey, shining ; the
oaterpiUar green and white striped longitudinally. This caterpillar occa-
sionally causes great mischief in forests ; see L08OHOE, Natwrgesch. der
ForUoder Kiifferravpe, Natufforseker, xxi. 1785, s. 27 — 65, Tab. m ;
as in Holland (particularly in the province of Gelderland) in 1808, and
especially in 1844 ; comp. hereon A. Bbahts in the Vaderl. Letierorfen*
ingen, 1844, Mengelwerk, bL 535 — 536 ; and on the destruction in the
pine-forests of the proviooe of Utrecht, H. Yeblobek in the Alg. Eunst-en
Zetter-bode, 1846, Kos. 13, 15, and 1847, Ko. 9.
5) Last Joint ef palps equal to second or longer than U, slender, sub*
nahed,
Erebus Latb. {Thyacmia Daijl)
Sp. Nodma Strix L., Fabb., Noetua Agrippma Cbameb, UiU. h^. i. Tib.
87, 88, fig. A ; CuviEB JJ. ani. id. iU., Ins. PI. 154, from Surinam. The
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INSECTA. 401
upper Bur&ce of wings white, with black lines forming many angles and
curves, Uie under surface brown-grey with white spots. This species is
one of the largest in this order ; the body is two inches long; but the fore-
wings, which are yety long, surpass those of all the rest in breadth
of flight, for the points are distant finom each other more than nine inches.
Another species also from South America, Nodua odora Gsaxeb, n. Tab.
169, f. A, B, was placed erroneously by LnrirjiUB amongst the AUaei
(Bombyx) ; it is smaller, brown-coloured, and has an eye-shaped spot on
the fore-wings.
Ltthosta Fabr. Tongne distinct, long, spiral. Labial palps
cylindrical, shorter than head, with third joint shorter than the
preceding, or coalesced with the second. AntennsB moderate, seta-
ceous, in males ciliated or subpectinate. Ocelli none. Anterior
wings narrow, horizontally incumbent. Caterpillars solitaiy, with
sixteen feet.
Sp. Lithoria guadraf PhoUcena {Noctua) quadra L., RcssiL, Ira, I., Pap,
noct, CL n. Tab. XVII. ; Skpp, NedeH. Ins, ra, 4e Stuk, Tab. VL ;
Duif&iL, Cone, gin, ». I. Int. PI. 42, fig. i bis; wings outspread 2",
yellow, the upper wings in the male grey-yellow, in the female yellow,
with two steel-blue or black spots ; the feet blue.
Sub-genus : NtidaHa Hawobth, Steph. (spec, of LUhoaia OcH-
SENH., species of CaUvmorpha Latr.)
Sp. Lithoe. mundana Oohsenh., Phakma mundana L., Hofficank. No-
twfortcker, xxvra. 1799, Tab. i. figs, i— -5.
Euprepia OcHSENH. {Arctta ScHRANK, Ckelonia GoDART,
BoiSDUV.) Tongue distinct. Antennae in males ciliated or pecti-
nate. Ocelli two. Wings deflected, variegated with colours often
lively, anterior triangular, posterior famished with retinaculum.
Larvae with sixteen feet, mostly hirsute with dense hairs.
CaUimorpha Late, (in part). Tongue elongata Antemue simple,
ciliated in males alone.
(Sub-genera: Emydia, Euehdia, and CaUimorpha Boisduv., species
of Euprepia and LUhosia Ochsenh.)
8p. Euprepia Jacdbaxe, Phal, (N'oel.) JaeobcuB L., K(BfiBL, Ina. i.. Pap. noct,
GL n. Tab. xliz. ; Sxpp, Nederl In$. 4e Stuk, Tab. xi; the upper wings
dark-bluish grey with two carmiue-red stripes along the anterior and
inner margin, and two round spots of the same colour ; the hind- wings
carmine-red with a narrow black border.
Arctia Sghrakk. Tongue short. Antennee, in males at least,
bipectinate. Abdomen thick.
VOL. I. 26
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402 CLASS VIIT.
Sp. Bupr^pia eaja, PhaL (Bomhyx) caja L., RoSBL, Int, I., Pap. nod.
GL n. Tkb. I.; Sepp, NederL Jn$. i. 40 Stuk, Tah. n ; winga outspread
^l" — $" broad ; fore-wings brown marbled with white, hind-wings vermilion-
red with blue-black round spots. Here belong many other species named
by ItUfiSMua Bombyces nobiles, Syst, not, ed. xn. i. p. 819.
Bsyche ScHRANK, Latr., Ochsenh. Tongue obsolete. AntennaB
in males pectinate. Ocelli two in many. Wings rarely scaly, sub-
pellucid, in females none. Caterpillars inclosed in a tube or sac of
conglutinated fragments of leayes and stalks.
Sub-genus : Oiketicus Guildino.
See Linn, Trant, TV. 1817, pp. 371 — 377. The apterous female does
not leave the follicle, and is there impregnated by the male.
Limacodes Latr., Heterogema Knoch, Treitschke. Palps
very short. Tongue obsolete. Antennae setaceous. Ocelli none.
Wings opaque; females winged as well as males. Caterpillars
broad, resembling slugs, creeping, with six homy feet very short,
membranous feet none, tubercles of the skin supplying their place.
Sp. LifMUiodet Teitudo, ffepialits Testudo Fabr., Sepp, Nedtrl. Int. n. 40
Stuk, Tab. IV. on oaks ; ffderogenea aaellana Tbeitsohsb, fftpialvM tueUm
Fabb., Knooh, BeUrdge zur Intectengetch. in. 1783, Tab. in.
Sericarta Latr. {Ltparts, Pygoera^ Endromts OcHSENH., Orgyta
Ochsenh., Latr.). Tongue short or obsolete. Antennae in males
or in both sexes pectinate. Ocelli none. Wings opaque, deflected,
posterior guarded by a retinaculum. Larvae with sixteen feet.
Sp. Sericaria ditpa/r, Phal. (Bomhyx) dispar L., B(ESEL, Ins. i., Pap. noet,
Gl. n. Tab. ni. ; Ratzbbubo, Font-Ins. n. Tab. ▼. fig. i. This species is
rery common, and sometimes very injurious to trees. The brown cater-
pillar with long bundles of hair has on each side of the body a row of
eleven round spots, of which the first five are bluish-white, the following
red. The female moths are a dirty white, with black bands on the upper
wings, and very sluggish; the male, much smaller, has brown wings.
In some species the females have merely short rudimentary wings {Orgyia
O0H8BNH.), Sericaria antiqua, Phal. {Bomb.) antiqua L., Robsbl, Ins. i.
Pap. noctum. CL n. Tab. xxxix. Tom. in. Tab. xm. &c.
NotodofUa Ochsenh., Latr.
Sp. Notodonta nczac, ^hal. {Bomb.) ziczac L., BcESBL, Ins. i., Pap. noctum.
CI. n. Tab. XX. ; Sepp, Nederl. Ins. 1. 40 Stuk, Tab. xii. &c.
Gerura ScHRANK, Latr., Harpyia Ochsenh. Tongue very
short or obsolete. Antennae in males or in both sexes bipectinate,
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INSECTA, 403
with teeth deflected, hairy, towards the extremity of antenn®
shorter or none. Ocelli none. Wings deflected, somewhat thin or
sub-pelludd, heneath pilose, the posterior bridled by retinaculum.
Body hirsute. Larv» smooth, with fourteen feet, and two conical
elongate tubercles in place of anal feet.
Sp. Cerwra mmda, Phal, {Bombifx) vinvla L., IUbsbl, Int. i,, Pap. noctwm.
CL n. Tab. XIX. ; SxFP, Nederi. Int. i. 40 Stuk, Tab. Y. ; Ltonbt, Owr.
potlh, PL 34, figs. I— 15. Putt Moth.
Bombyx nob. {Bonibyx and Lasiocampa ScHRANE, Latr., Qdi-
tropacha OcHSENH.) Tongue short or obsolete. Ocelli none. An-
tennae in both sexes bipectinate. Wings deflected, reversed, the
external margin of posterior produced beyond the margin of the
anterior ; retinaculum none. Body thick.
Sp. Bombyx mori, Phal. {Bombjfz) mori L., BassEL, Int. in. Tab. vn —
EX. the tUh^oorm; the caterpillar ia whitifih-grey, smooth, and has a smaU
horn at the hinder extremity on the back ; when fully grown, it reaches
quite 3" in length, and feeds on mulberry-leaves ; the moth is dirty-white,
and has on the fore-wings inconspicuous light brown stripes. The pupa is
brown, elongate, oyal and obtuse ; it lies inclosed in a close web (cocoon)
from which silk is procured. This species was first introduced into Europe
in the time of the Emperor Justinianus.
Bomhyx potatoria, Phal, (Bomb.) jwtatoria L., Bcbsbl, Int. 1., Pap.
nocium. CI. n. Tab. n. ; Skpp, Nederi. Int. i. 40 Stuk, Tab. vra. Drinker
moth.
Satumta ScHRANK, Latr. {Satumia and Aglia Ochsenh.)
Tongue obsolete. Antennae in males doubly pectinate, with two
teeth directed upwards and two downwards in every joint. Ocelli
none. Tfings when at rest expanded, the inferior without reti-
naculum, the disc adorned with ©cellar spot, which is most fre-
quently naked in the middle, pellucid.
Sp. Satumia carpini, Oohssnh., Phal. (Bomhyx) pawmia a) minor L.,
BcBSXL, I. Pap. nod. Gl. n. Tabs. iv. v. ; Skpp, Nederi. Int. i. 4e Stuk,
Tab. X. XI. Amongst the exotic species may be noted the East Indian
Salvmia atlcu (Gbaheb, UUl. Kap. i. Tab. 9, fig. a. Tab. 381, fig. c. Tab.
38a, fig. A.) on account of its size and beauty. All the species of this
genus are amongst the larger Lepidoptera.
Co88ti8 Fabr. {C088U8 Latr., Zeuzera ejusd.) Tongue obsolete.
Antennae denticulate or pectinate, in some towards the extremity
setaceous, simple. Ocelli none. Head small, deflected. Thorax
gibbous, large. Female with ovipositor exsert. Wings deflected,
26—2
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404 CLASS VIIl.
elongate, Totnndate, inferior furnished with retinaculum. Larvae
smooth^ with sixteen feet, living in wood.
Sp. Coutu ligniperda, Phal, (Bomhyx) Oomu L. ; Sbfp, Nederl. Int. III. Tab.
XLni. XLiY ; DuMiaiL, Cons. gin. «. I. Ins. PI. 45, fig. 5 ; Ratzebubq,
Forst-Ins. n. Tab. ni. fig. i. The expanded wings are more than 3"
broad ; the colour of the wings is ash-grey, with black interrupted stripes
that meet reticulately. The caterpillar reaches a length of three inches,
and is red-brown on the back; it lives more than two years before changing
into pupa, and infests not willows alone, but other trees also. It is this
species which was investigated by Ltonbt in his unrivalled TraiU anat.
de la Chenille, and is on that account so generally known. See also
his Recharches swr VAnat. et les Metamorphoses des Insectea, edited by D£
Haah, pp. 369 — 546, PI. 39 — 54 (on this insect in the state of pupa and
moth).
Zeuzera Latb. Antemifle in males pectinate at base, setaoeous at
apex, in females serrate.
Stf/ffia Dbapabn. (Species of Chimcera Ochsekh.)
Ifepicdua Fabr., Heptolus Illig. Tongue obsolete. Antennas
very short (shorter than thorax), filifoim or subserrate. Ocelli
none. Wings deflected, lanceolate, posterior mostly without reti-
naculum. Larvae smooth, with sixteen feet, subterranean, rhizo-
phagous.
Sp. Hepudvz humuli, Phal {Noetua) humuli L., Pakzeb, Deutschl. Ins. Heft
70, Tab. 23, 24; — Hepud. lupulinw, Phal. (Noetua) lupuUna L. ; Panzer,
Deutschl. Ins. Heft 94, Tkb. 1%, 14; Gu^in, Iconogr. Ins. PL 85,
fig. I, Ac.
Family XXVI. G^ejmacularia. Wings subhorizontal or de-
flected, bridled in almost all, the posterior furnished with -retinacu-
lum. Antennae prismatico-clavate or fusiform, simple or serrate, in
few pectinate. Tongue distinct, in some very long. Posterior
tibiae with double internal spine. Caterpillars with sixteen feet,
sometimes naked, furnished posteriorly with dorsal horn, sometimes
pilose or hirsute. Pupa smooth, in some foUiculated, in most
buried beneath the earth. Flight of imago matutine and ves-
pertine.
Hvening-matha, Tmlight-moths, Grepuacuiaa' moths. This fiumly
consists for the most part of the linnaean genus Sphinx,
I. ATiierior wings elongate, naarow, posterior small, much smaller
ikam anlerior. ^^
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INSECTA. 405
Sesia Fabb. (exclusive of species). Wings subcjlinclrical, in-
crassated before the apex, terminated by a fasciculns of rigid setffi.
Ocelli two. Wings hyaline.
Comp. J. B. Labpbtsbs, 8e»ia Furoptm ieonih. €t deacriptUmUmi iUui-
irakB, Berolini, 1801, 4to.
TAjfria Hoffmannsego.
The larvse of these two genera five in the wood or the pith of plants.
Zygcena Fabb., Anthrocera ScoPOLi. Antennn in both sexes
simple, flexuoso-clayate, with smooth apex. Ocelli two. Tongue
long, spiral. Wings deflected, liyely coloured, longer than body,
narrow.
Sp. Zygmafilaipmdvila, Sphinx flipmd/uki L., B<bsbl, In». 1., Pap. noetum,
CI. IL Tab. Lvn. ; Sbfp, NederL Iru, i. 4e Stuk, Tab. xxii : the body and
aDtennte are blackish-green, the upper wings dark-bluish green, shining,
with six carmine-red spots ; the under-wings cannine-red with a bluish-
black edge. The caterpillar lives on clover, &c. is yellow, thick, with
short hairs, without horn on the back, and makes a compact but thin web
on the stem of the plant when about to change into a pupa. Tliis genus
approaches Supnpia in the preceding family.
SynUonm Illio.
Comp. BoiEfDUYAL, Earn tw une Monographie da Zyg^nidea, Paris 1829,
8vo, av. pL color.
Olaucopis Fabb. (and Procris ejusd.) AntennsB in males or in
both sexes bipectinate. Ocelli two.
a) Tongue obsolete.
Aglaope Latb. (spec oi Atychia Ochsbnh.)
h) Tongue distinct.
Sub-genera : Glaucopia Fabb. (With antennse in both sexes bipec-
tinate), Procris Fabb. (Ino Leach), Atychia HoFPKANNa, Latr.
Sp. Sphinx Suaices L., Ssff, Nederl Ina, iv. Tkb. 40; Panzbb, DeuUehl. Ina,
Heft 32, Tab. 34.
Sphinx L. (in part). Antennae clavato-prismatic, incrassated
before the apex, mostly with uncus or short seta, recurved and
ciliated at the extremity. Labial palps broad, compressed, with
third joint indistinct. Ocelli none.
Smerinihua Latb. Tongue very short Antenn» serrated. An-
terior wings angulate.
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406 CLASS VIII.
Sp. Sphinx TUuB L., I^hinx Populi K, Sphinx ocellata L. Comp. Sefp,
Nederl. Ins.
Sphinx Latb. Tongue distinct, sometimes very long. Anterior
wings in some angulate, in others quite entire, rotundate.
a) Tongue shorter, {Acheraniia Oohsenh., Brachyglossa BoiSDUV.)
Sp. Sphinx Atropos, RoesKL, Ins. m. Tab. i. ii. ; Sxpf, Nederl, Ins, m. Tab.
XXII— xxYin. ; Panzeb, Deutschl, Ins. Heft 8, Tab. i6. The outspread
fore-wings have their tips more than 4" apart^ the body is 2" long. The
upper-wings are brown, grey, flamed black and yellow-white, with a small
whitish and round spot on the middle ; the hind- wings are yellow, with two
pale-black bands. The loud noise which these animals make (on the cause
of which there are many different opinions) has occasioned the superstitious
dread which has sometimes surrounded this insect ; to the yellow spot with
two black points, which is seen on the thorax, and which has some resem-
blance to a skull, it owes the name of Death's-head moth. The cateipillar
feeds on the leaf of the potato.
h) Tongue dongate.
Maeroglossa Oohsenh. {Macroglos^um Scop.) With eztrsmiiy of ab-
domen bearded. Tongue very long.
Sp. Sphinx stellatarum It,, B(ESKL, Ins. i., Pap. noelvm. CL i. Tab. vin. ;
Sefp, Nederl, Ins. ii. 30 Stuk, Tab. i. the ffumming-hvrd hawk.
Pterogon BoiSDUV. (Sp. ot Maeroglossa Oohbxnh.)
Sphinx BoiSDUV., {Deilephila and Sphinx Oohsxkh.) Abdomen conical,
with extremity not barbate.
Sp. Sphinx Ligustri L., Bcesel, Ins. us. Tab. v. ; Sxpp, Nederl, Ins. i. 30
Stuk, Tab. in. rv. : the flight is sometimes more than 4" broad, the body
is 2" long ; upper-wings brown with white at the point and the posterior
margin, hind-wings rose-red with grey-brown margin, and three black
transverse stripes ; abdomen red above with a brown mid-stripe and black
rings. The beautiful green smooth caterpillar, with oblique violet and
white stripes, on the syringas and privets, is rare in Holland.
II. Wings trtatiffular rotundate, the posterior congruous.
Castnta Fabr. Palps distinctly triarticulate. AntennsB clavate,
with club elongate, sub-fusiform, not bearded at the apex.
Sp. Pap. Licus Cramer, UiU. Kap. Tab. a 23, t%. a; Ccutoia PalUma
EsoHSOH. ; O. V. KoTZEBUB, Entdeckungs-Seise, ni. Tab. vx. fig. 47 ; —
Caslnia acrceoldes BoisDUV. ; GujSein, Iconogr. Ins. PL 83, fig. 4, ftc. All
the species hitherto known are from South America.
Comp. on this division J. W. Dalhak, Prodromus Monographic! CatinicB,
0. Tabula len. color. Holmse, 1815, 4to. These moths make the natural
transition to the following family, and are in their habitus rery similar
to some diurnal butterflies ; their wings however have a retinacalum.
Add sub-genera: Coronis Latb., AgoHsta Lsach and Cocytia
BoiSDUV. (Monogr. dea Zyghiides PI i. fig. 1.)
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INSECTA, 407
Family XXVII. Dtuma. Wings mostly erect when the in-
sect is seated, neyer bridled by a retinaculum. Antennae in by far
the most clavate, abruptly terminated by a capitulum, in a few
filiform or subsetaceous, with apex more slender, uncinate. Ocelli
none. The caterpillar always with sixteen feet. Chrysalis almost
always naked, angulate, attached posteriorly by threads, or sus-
pended vertically, or affixed by a transverse silken cord expanded •
above the middle of the body. Flight of Imago diurnal.
BuUer/liea (RhopcUocera BoiSDUV.) These insects have usually
clubbed antennse, which is the case with all our domestic species ;
when at rest they erect their wings, so that the upper surface of the
wing is turned inwards. The genus Papilio of LnryiEUS corresponds
to this &mily of later writers.
Comp. on thifl fiimily, Godabt, article PapiUon, making the entire ix.
part of the Hist, naiur., Inaectet, of the Encyclopidie nUthod. iBig, and E.
DovBLKD AY, TluOenera of diumalLepidoptera, iUudnUtd with colour, jplaia,
London, 1846 and foil. 4to.
Phalanx I. Posterior tibiae, as in the preceding families,
spinose not only at the extremity, but also on the inside before the
extremity. (Caterpillar very often living among leaves that have
been spun together. Chrysalis smooth, folliculate, or tied up by
a transverse thread.)
Urania Fabr. Antennae filiform, more slender at the apex,
and arcuate or uncinate. Labial palps triarticulate, elongate, slen-
der, with second joint greatly compressed, third slender, subcylin-
drical, almost naked. Wings broad, large.
Sp. Urania LeUw, Papilio (Eques) Leilus h., Klbbmak, BeytrOffe, Tab. n.
fig. I, South America; — Urania Boitduvalii Gu^iw, Uran. Femandinat
Mao Lkat, Gu^bik, Iconogr. Ins. PI. B2, fig. i ; the larva, the web and
the pupa figured in Trans, of the Zoolog. 80c. I. 2, 1834, pp. 179—189, PL
a6. The larva is thick, with a few hairs, in form not unlike a caterpillar of
Callimorpha, but with a very large head; the web is thin, so that the pupa
is visible through the meshes. These species belong t^ the genus Cfjfdimon
of Dalm AK, Pap. (Eques) Ortmtes L. (Cbamsb, UiU. Kap. Tab. LXXXin. figs.
A, b) to the genus Nyctalemon of the same.
Urania Ripheus Cbambb, Uid. Kap. Tab. cgclxxxv. fig. a, b ; 601s-
DUVAL, JVouv. Ann. du Museum, n. 1833, ^^> ^4^ ^' <> ^> ^^^ ^ spiny
caterpillar with the first four membranous feet short, so that it moves like
a geometric caterpillar. It does not spin itself up when about to change
to a pupa, but affixes itself by means of a thread stretched transversely over
the body, like the caterpillars of the genus Pieris, &c. This species^ placed
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408 CLASS VIIl.
by DaUIAH under Ojfdimon, fonns the iype of a sepanie sab-gsBW Urania
or Tkalmra Ddvoah.
The affinity with CtuHnia, espectaUy with Conmu and Affa^ritia^ cannot
be mistaken ; and some writen, amongst whom is Wbstwood, place UrasMa
in the former family. In those epecieSy howerer, which 1 have ezaminedy
I can discover no uHnacylwn,
Hesperia Fabr. (in part), Cuv., Latr. Head thick. Antennse
remote at the base, capitate. Palps shorty with second joint ex-
tremely scaly forwards, and third joint short. Upper wings erect
only when at rest, or expanded wings patent. Caterpillar often
living in leaves webbed together.
Sub-genera: £udamtt8 Boisduv., FamphUa Fabb., Hesperia
Fabsl, Syricthue Boisduv., Tkymele Fabb., Tcmaoe Boisduv.
Sp. Eetperia malwB Fabb., BosaxL^ /n«. i.. Pop. ditun^, CI. n. Tab. x. fto.
Phalanx 11. Posterior tibias spiny at the extremity or spurred
alone. Four wings erect when at rest. Antennae always capitate
or thicker towards the extremity. Caterpillar in the open air.
Chrysalis most frequently naked and angulate* (Antennae in aU
approximate at the insertion.)
I. Third or last joint of labial palps most distinct, naked or
slightly covered with scales or hairs. Claws of tarsi small Cater-
pillars oval, resembling (yiiiscL Pupie smooth, rotundate.
Polyommatas Latr., nob. Six ambulatory feet in both sexes.
a) Labial palps elongate, produced in front of head.
Myrvna Fabb.
6) Labial palps short or moderate.
Polyo7nm<Uu8 Latb., Ck>DABT. {Lyccena, Thecla, Hesperia Fabb.
Syst. OlossoL)
Sp. Polyommattu hetulcB, Pap. hetula L. Sepp, NederL Int. m. Tab. xn.;
Panzbb, DeutsclU. Ins, Heft ai. Tab. 20 i—Payominatua Phkdaa, Pap.
PUaat L., BcKBBL, Ina, m. Tab. 45, figs. 5, 6, &c.
Erycina Latr. {Erydna^ Nymphidium^ Heltcopisy Ernests Fabe,
Syst Glossat) Anterior feet in males imperfect, not ambulatory,
very short.
Sp. Erycina {Hdicopit) Oupido, Pap. Oupido L. Bxe&el, Ini. IV. Tab. m.
fig. 7 ; Gbameb, UUL Kap. Tab. 164, figs. i>>-a, from Surinam. All the
speciee of this genus are from South Amerioa.
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INSECTA. 409
NoU, — Hero, it would aeemy ought to be placed the anomalous
genus Ba/i^jioorms Godabt, with setaceous, ciliated antennae, uniting
the UranioB with the ErycmoB and PolyommtUL See fig. in Cuyieb
R, AnL ^d. iU., Ins. PL 143, fig. 4.
IL Third or kst joint of labial palps in some distinct, beset
with scales like the rost, in others small, subindistinct Claws of
tarsi distinct. Caterpillars oblong, cylindrical Pupa mostly angu-
late, anteriorly bimucronate, or acuminate.
a) ffexapodct. Six ambulatory feet, nearly similar in both
sexe& Chrysalis tied posteriorly and by a transverse string. Cen-
tral cell of posterior wings always closed. (Species of the SuceincU
BOISDUV.)
Papilio nob. {Papilianes Equiies and Heltconii L., exelusiye of
some species.) Posterior wings scolloped in the inner margin, not
covering the abdomen below. Anterior tibiaa furnished beneath,
with a spine in the middle.
Sub-genera : Leptodrcua Swainsok, Papilio Fabb., Latb., {Teinch
pdlpua Hope, Omithoptera Boisduval, Doubleday, PapiUo Bois-
Duv.), Ewrycua Boisn. {Cressida Swainson), Pamaaavua Latb.
1^19 Fabb.
Sp. Pap. Mackcum L. R(E8XL, In». I., Pap. dium. Gl. n. Tab. I. ; Skfp,
NederL Ins. i. a» Stuk, T^b. m. ; the queen-poige; the tips of the outspread
npper wings 4" apart; the Ixrgest domestic species of day-butterfly; the
wings yellow with black veinures, the fore-wings at the base black ) the
hind-wings tailed, with a margin of blue, half round spots on a black band,
and a red eye-spot bordered with blue and black at the inner side. The
green caterpillar has black rings with orange spots, and lives on the leaves
of carrots {Dauctu Carata, &c.) The genus of the Knights butterflies
belongs especially to warm countries, the East and West Indies.
Pierts nob. (Species of genus Pierts Schrank). Anterior tibisB
without middle spine. Posterior wings caniculate on inner margin^
receiving the abdomen beneath. (Wings mostly rotundate, in
many yellow or white or variegated with orange and yellow).
Sub-genera: Euterpe Swains., ZeptcUis Dalman, LetLcophcuia
Stephens, Poniia Boisnuv., Doubleday, Pieris Boisduv., Doublk-
DAY, Zegria Rambub, Nathalis Boisduv., Anthocharis Boisduv.,
Idmaia Boisduv., Theatiaa Boisduv., Eronia Huebn., Doubleday,
CkdUdri/aaBoiSDXJY.,DojJBLED., Gtmeptert/xliBAcn^ Doubled., {Ehdfh
docera Boisduv.), Cdias Boisduv., Teriaa Swain&, Boisduv.
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410 CLASS VIIL
Sp. Piarii hramieiB, Pitp. bramiea L., Bcbsbl, Int. L, Pap. diumor, OL n.
Tab. IV. ; Skfp, Nederl Int. i. ae Siuk, Tab. n., the great white hutterfy,'—
Pierit (Gon^eryx) Bhamni, PapUio Bhamni L. ; Sefp, Nederl, Int. TV.
Tab. xxzvn. the citron butterfly, &c.
b) Tetrapoda. Two anterior feet, in males at least, small and
recurved. Chrysalis tied posteriorly alone, suspended freely, with
head downwards. (Svspenri Boisduv.)
* Anterior feet in males very short, in females like the rest of
the feet.
Libyihea Fabb., Secaerge OcHSENH. Palps very long, porrect
in form of a pointed rostrum. Wings angulate, anterior falcate,
with apex truncated. Discoidal cell of posterior wings closed.
Sp. Libythea cdHt, Fuesslt, Archivet de VHitt. det Inteetee, Winterthour,
1794, 4to. Tab. 8, figs. 1—3. Tab. 14; Cuv. H. Ani. 6d. ttt., Int. PL 136,
figs. I, 1, in Southern France, the Tyrol and Italy.
** Anterior feet in both sexes very short.
BibUs Fabr. Palps longer than head, distant, with last joint
inflected. Antenna towards the end incrassated gradually into an
elongated club.
Sp. Btblit Thadana Godabt, Pap. Hyperia Gbambb, Uitl. Kap, PL 336,
figs. B, F ; GuvDEB, R. Ani. idU. HI., Int. PL 136, fig. 3, South America,
&C.
Satyrua Latr. Palps approximate, ascending, extremely hir-
sute. Nervures of upper wing often inflected at their origin. Dis-
coidal cell of posterior wings closed.
Sub-genera: Hippa/rchia "Fasel, Ochsenh., Hastera "Fasbl, and
some others of modem writers.
Sp. Saiyrut jBgeria, Pap. {nymphalit) JBgeria L., Sbpp, NederL Int. I. le
Stuk, Tab. VI.; Paxzbb, Deuttehl. Int. Heft 28, Tab. ai, &c
Morpho Fabr. Antennae long, gradually thicker towards apex,
almost filiform. Wings very large in proportion to the bulk of the
body.
Sub-genera : Pavonia Godabt, Amathusia Fabb.
Sp: Morpho Belenor Gbamib, UiU. Kap. Tab. 36, figs, a, b ; Guvdeb, jR. Ani.
id. iU., Int. PI. 140, South America &c.
Brassolis Fabr. Palps short, compressed, approximate, squa-
mate. Antennae terminated by an obconical club. Posterior wings
with discoidal cell closed, canaliculate on the outer margin for
receiving the abdomen. Body thick.
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INSECTA. 411
Sp. Bras9olU topkorof, Papilio {Danaut fuUvut) Bopkorce L., Klbbmanv,
Beytrdge, T»b. lY. figs, i, a; CuviZB, R. Ani. id. ilL, In$, PL 141^ fig.
1, Ac.
Nymphalta Latb. (With addition of other genera). Palps
approximate, erect, sqnamose. Posterior wings with discoidal cell
almost always open, with internal margin canaliculate to receiye
the abdomen.
a^ Antennae gradnallj passing into an elongate clnb.
Sub-genera : Limemtis Ochsenh. (LxTneniUa and Neptis Fabb.),
Ageronia Huebn., Doubleday, {Fertdramia Boisduv.), Apahira
Fabb., and some others of later writers.
Sp. NympKalis Iris, Papilio {Nymphalit) Iria It., Bcbsbl, Int. UL Tab. 49^
fig. a ; Panzbb, Deutichl. Ira. Heft 79, Tkb. 33, &o.
V) Antennae abruptly clavate. Eyes downy.
Sub-genus : Vcmesaa Ochsenh. ( Vcmesaa and Cynthia Fabr)
Sp. NymjphdlU polychloroi, Paip. (Nymph.) PolychUroi "L,, B/cessl, Im. I.
CI. I. Pap. Diwmor. Tab. n. ; Skpp, Nederl. Im. L le Stuk, Tab. 8; the
outspread wings 3 j" wide, above brown orange-ooloiuned, with black spots,
the hind- wings with a band of semilunar blue spots. The caterpillar of this
and other species is furnished with many spines. — ^Here belong NymphaUa
lo h., Sbfp, ibid. Tab. 7, DuM^ni, Com. yen. s. L Ins. PL 40, Nymphalis
UrHcoi li., SSFF, ibid. Tab. a, &c.
c) Antennae terminated abruptly by a short dub. Eyes naked.
Sub-genera : Melitcea Fabb., Argyrmia ejusd.
Sp. Nymphalis (Argynnis) Laihonia, Papilio (NymphdUs) Lathonia L.,
IUbssi^ Ins. m. Tab. x. &c.
Heliconiua Late. {Mechanites Fabr., Acrosa ejusd.), Hdtconia
GrODART. Palps short, remote. Abdomen slender, elongate. An-
terior wings elongate, oval, the posterior scarcely, or not at all, re-
ceiving the abdomen beneath. Discoidal cell of posterior wings
always closed.
Sub-genera : TUhoreay Heliconia, Lycorea, and others ; on which
comp. Doubleday, Genera pp. 99, ko. All the species are from
America, except sub-genus Ha/madrya8 Boisdttv.
Sp. ffdiconius Erato, Papilio {Htlicomus) Eraio L., Grajcbb, Uitl. Kap,
Tab. 177, fig. P; CuvuBB, R. Ani. 4d. HI., Ins. PI. 133, fig. 3.
Danats Latr. Palps remote. Antennae very long, in some
almost filiform, in others terminated by an elongate oval club.
Wings triangular, broad, rotundate at the margin ; discoidal cell of
posterior wings always closed.
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412 CLASS yiii*
Sub-genera : Eupkea Fabb., JkmoM "LkTBL^ Idea (ReHia) "Hvebs.
DOUBLEDAY.
Sp. Idea Agdia Godart, PafUio {Danmu candidm) Idea It,, GBAiCBBy UiU,
Kap. Tab. 193, figs. ▲, a; Doubledat L L Tab. 13, fig. i. Amboyna. Hie
species of the sub-genus Idea are large, and have white or grey-white wings
with black veins and spots. They all belong to the East Indian islands^
or to Kew Holland. Of the sub-genus Danaie, species are found both in
the Western and Eastern hemispheres.
Obder IX. Neurcptera,
Hexapod insects, with four wings membranous, mostly trans-
parent, reticulate, naked, most frequently equal. Mouth not suc-
torial, but in most formed for manducation, with mandibles in some
obsolete. Females never aculeate, very rarely famished with borer
or exsert oviduct. Metamorphosis in most incomplete, in others
complete. LarvsB hexapod.
Tendonrwinged. Newroptera L., from vevpov, tendon, and vrepov,
— OdoncUa and most SynisUxta of Fabbicius, Nevo'optera and Tricho^
ptera Kirby. On this order in general I am acquainted with no
other special work than that of P. Rambub, Histoi/re naJL dee In-
eectee NewropUree ; Ouvrage accompagnJ^ de pUmchee, Paris, 1842,
8yo, making a part of the Suitee d Buffon by Roret. Other
works on particular families we shall indicate below.
The Newroptera are insects with four naked, mostly transparent,
wings, of which the veinures form a fine net-work. Usually the
under-wings are as large as the upper ; in many species oiJSphemera,
however, they are very small, and in some entirely absent. The
females have no sting ; rarely only there exists a borer or an instru-
ment for depositing eggs. Hereby they are distinguished from the
HymeTioptera, as well as by the lower jaws with the under lip not ,
being produced into a sucker. The number of joints in the tarsus
is different in the different genera.
Many insects of this order are, in the state of pupa and perfect
insect, carnivorous and truly predaceous. The larvae for the most part
live in water {Phrygaaiea, SemJblie, Ephemera, Ltbelltda). Some larv»
spin themselves up when they change into pupse. In the larva of the
lion-ant the vessel which secretes the web, an oval sac, is situated in
the abdomen j it ends in a tube capable of extrusion ; by means of it
the larva spins together granules of sand to form a round ball, within
which it changes into a pupa. In the larvie of the genus of the
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INSECTA, 413
Ccue-nymphs {PhrygtmetB) there are, as in eateipillars, two long
Teasels, which secrete the silken material In their straight intesti-
nal canal, also, and in the small number of their v(Ma urinaria,
these larvae resemble caterpillars'; and on the whole the genus
Fhryganea, although still really belonging to this order, forms a
yery natural transition to the Lepidoptera, which does not rest
merely on general external resemblance or analogy.
In this order many species are met with in which the instinct
is surprising, and their economy extremely interesting; we name
merely the lion-ant with its crafty device to overpower its prey,
and the celebrated mines and buildings of the white anta^ or ter-
mUes, &0.
A. Neuroptera with metamorpkons complete.
Tarsi in all quinquearticulate.
Family XXVIII. PhryganidcB {Trtchoptera Kirbt). Wings
deflected, posterior often broader than anterior, folded longitudi-
nally, the anterior with numerous branched nervures, covered with
hairs, coloured. Mandibles obsolete or very small, remote, not
convergent. Maxillary palps long, with three to five joints,
labial palps short, three-jointed. Prothorax short. (Antennae with
numerous joints, long, mostly setaceous. Ocelli three.)
Gomp. C. F. PiOTBT, Recherchet pour tervir d VHUt, dt d VAncOonnie dea
Phryffanidet. Avec pi. color, i vol. 4to. Genbve, 1834.
Case-nymphsy Caddis-flies, The upper jaws are here veiy small
and little developed, as in the last order (Savioky, Mern. s. I. ani, s.
, vert. I. p. 29, PI. i. fig. 1). The larvse live in water, in cases open
at both ends, composed of small pieces of wood, of small fresh-water
shells (in which frequently the inhabitants are still resident), of
sand, of duck-weed, of fragments of leaves that have fiJlen into the
water, <&c. They fasten these substances together by means of their
web j the inner sur&bce of the case, which is very smooth, is also
1 Baudohb found only four, which agrees with the number in caterpillars, but
this requires to be confirmed by further investigation, since Ll^N DuFOUB, in the
species examined by him, always met with six. However, the Ephemerinae and
IdbeHuluuBf i,e, by £» the greatest number of the Neuroptera, have very numerous
(30 — 40) short va$a urinaria^ like the Orihoptera; those which undergo a perfect
metamorphosis only six or eight ; see LioN Dufoub, M^m. pr^senUe, Tom. vn., and
LoBW in Gbbmab's Zeilschrift far die ErUomologie iv. (1813), s. 4^3, &c. (Bemer'
hunffen Uber die anatonUachen VerhdUnme der Newropteren,)
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414 CLASS Till.
lined with the web. Most species drag the case with them, but
others fasten it to stones and creep in and out. These cases protect
the larrse from the x>er8ecution of fishes, which ar^ veiy fond of
them ; in England, where fishing with the line is brought to a much
greater state of perfection than perhaps anywhere else, these larvie
are much used for bait. The larvsB feed on vegetable as well as
animal substances. Most of them have white threads at the abdo-
men, external respiratory- organs, gills (see above pp. 262, 263).
The larvBB, when about to change into pupse, fix these oases firmly
and close the openings with crossing threads, that thus form a latticed
cover. The pupse, at first immoveable, quit this dwelling when
about to change into perfect insects ; it swells and bursts in order
to allow the perfect insect an exit. This last is generally brown-
coloured, and resembles a nocturnal butterfly. On an evening, at
country-houses, it is frequently allured by the light into apartments,
and flies round the lamp or candle until, its wings half-consumed, it
falls down with violence on the table. The eggs are joined together to
form a gelatinous mass, and are deposited by the female on leaves of
branches which hang over the water, so that the young larvie on
creeping out may easily fall into their natural abode, if the leaf by
falling previously has not already reached the water.
See on the economy of these Insects espedally RfiAUXUS, Mim, Tom. n.
pp. 497 — 581, and, on the arrangement, besides Raicbub and Wbstwood,
also especially Bubkbibteb, Handb. derEwUmoUu. 2, 1839, pp. 882 — 955.
Phryganea L. (exclusive of the PerlcB Geoffr.)
A. Antennffi filiform or pectinata
HydrcptUa Dalmai^. Maxillary palps with five joints. Wings
narrow ; posterior not plicate.
Sub-genus : Na/ryda, Stephens. With antennie pectinate.
B. AntennaB setaceous.
t Maxillary palps with five joints.
CKwMvrra, Leach. Posterior wings not plicate.
Mystadda Latb. Leptocerus Leach. Posterior wings plicate.
Maxillary palps long, pilose. Antennae very long.
Sp. Phryganea nigra L., Db Gbbb, n. PI. 15, figs. 11 — 23, ko.
Add sub-genera Odantocenu Lbaoh, Setodea Rambub, Ac
Hydrop8che Pictet.
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INSECTA. 415
Bhiaeophila Pictet.
Fat/chonda Latb.
B. ft Maxillary palps (of males) with two to four joints.
Sericostama Latk Maxillary palps in males mostly with two
joints, in some with three, dilated, yillose externally, often veiling
the head.
Sub-genera Pogcnostoma Ramb., Ikuifdonui ^H$d, : with nuudlkiy palps
of males three-jointed.
Sub-genera Trichotloma Piotbt, Z<uiottoma Ramb., Lepidodoma Bamb.,
Serieottoma Piotit : with maxiUaiy palps of males two-jointed.
Phryganea FiansT. Maxillary palps of males fonr-jointed, of
females five-jointed, longer than labial^ subpilose. Wings with
transverse nervures.
Sub-genera Xtmn^AtZaLsAOH, Manocentrct, Fnoicyla, OUgotrieha Rajcbub.
Noie. — For other recorded genera see Wbstwood, Oenerie SynoptU, pp.
49—5 1> Uld BUBMIISTBB, L 1.
Sp. Phrygtmea grandit L., Bcbsbl, Iru. u., Inteti. a/qwOU, CI. n. Tab.
xnn. ; Db Gbbb, Mim, n. PL 13, fig. i ; — Phryg. rhomJbica L., {Linne-
pkUa Lbaoh) ; Bgmbl, ibid. Tkb. xvi. ; Gbbxab, Faun, In$. Ewrop. Fasc.
IX. Tkb. 13, &c.
Family XXIX. Panorpatce. Posterior wings expanded. Man-
dibles distinct. Head with anterior process mostly rostriform;
ligula none ; labium and maxillss mostly connate at base. Maxillary
palps five-jointed, labial two-jointed. AntennsB filiform, long, mnl-
tiarticnlate.
A. Ocelli three frontal. Wings perfect, reticolate, with longi-
tudinal nervures numerous, transverse few.
BiUacus Latr. Feet elongate, slender, tarsi with single claw.
Abdomen cylindrical, long. (Habit of Tij[mla.)
Sp. BUtaew tipulariug, Panorpa tipularia Fabb., C. Db Yillbbs, Bntomol,
Linnean. Tab. 7, fig. 11; Sulzbb, Oeach. der Ins., Tab. zzv. figs. 7, 8;
GuIIbin, Icon,, Ins, PI. 61, fig. 2, habitat in south of Europe.
Panorpa L. (exclusive of P. hyemalia and Coa). Feet moderate,
tarsi with two claws, terminated by intermediate pulvillus. Abdo-
men of males with three posterior segments recurved, last oval,
globose, forcipate.
Sp. Panorpa wmnwms (and Pan, germanica) L., Pakzbb, DetUsckl. Ins,
Heft 50, Tab. 10; Dum^il, Cons. gin. s. I Ins, PI. 47, fig. 6. The
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416 CLASS viir.
Scorpion-fiy, A. Bbants has given an anatomical sunrey of this little
animal in the IHjdachr. voor not. Getch, en Physiol, vi. 1839, bl. 173 — 198,
PL vn. The pupa is concealed underground; it has been described and
figured by F. Stein, Wieqkaks'b Arekiv, 1838, s. 330. Taf. 7, figs. 10 — 13.
The larva, with its mode of life and its residence, is still unknown.
B. Ocelli none. Wings imperfect, in males subulate, in
females very short, rotundate.
Boreus Latr. (Female with triarticulate ovipositor, encased
beneath in a bilobed lamella.)
Sp. Boreui kyemalis, Panorpa hyemalit L., QryUut probotcideus Panzeb,
Deutaehl. Int. Heft a 3, Tab. 18 ; Gu^BiN, Iconoyr. Ins, PL 61, fig. 4.
Note. — Genus Choriata Klug, resembling Panorpa in habit and
characters, differs from it in the head not being produced into a
rostrum. Species from New Holland.
Comp. Kluo, Versuch einer sysUnuUischm Peststdlung der Insecten-fatnUie
PanorpatflB und A usanandenOzunff ihrer Oattungen tmd A Hen; A bhandlun-
gm der konigl, Akad. der Wissensch, zu Berlin, aus demJahre 1836. s. 81 —
108. c. fig.
Family XXX, Hemerobini. Head moderate or small, trans-
verse, with eyes prominent. Mandibles distinct. Mandibles deeply
bilobate, with external lobe often palpiform, articulate. Ligula
distinct. Labial palps triarticulate. Wings reticulate, with nume-
rous transverse nervures, mostly deflected. Antennse always com-
posed of many joints, in length very different.
Raphidia L. (in part). Palps short, filiform, maxillary quin-
quearticulate. Antennae filiform, short. Prothorax elongate, nar-
row, subcylindrical. Third joint of tarsi bilobed, receiving the
minute fourth joint between the lobes, fifth elongate, terminated by
double claw. Abdomen of female furnished with ovipositor, exsert,
flexible.
Camd'fy, These insects live in the state of pupa and nymph under the
bark of trees ; the nymph begins to move shortly before the last meta-
morphosis. Comp. A. Perchbbon, Mim, swr Us Paphidies in GuiaiK,
Mayasin de Zoologie, 1833, ^ i^* ^' ^l ^^ Stbin in Wibomani^'b
Archie, 1838, s. 316 — 330; 6. T. Sohnbidsb, Monoyrapkia gemeris
Paphidiat, adjectis tabul. vin. YratisbiTiaB, 1843, 4to.
LnTNiEUB united the European specimens known to him into one
species, Paphidia ophiopis. Modem writers distinguish different species ;
in most, three simple eyes are present ; in one species, first described by
SoHumcKL, Paph. crassieomis, they are wanting ; of this species, Sohnudkb,
op. cit., fonus a separate genus, InocdHa.
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INSEGTA. 417
MamUspa Illio., Latb. Anterior feet raptorial (Habit of
Mantis.) Ocelli mostly nona
Sp. ManHtpa pagana, Raphidia MarUUpa L., Gu&nt, leonogr., In». PI. 6$,
fig. I, ViLLEBS, Entom. Linn, Tab. vii. fig. 13, &c. Comp. Ebiohson,
Beitrage su oner Bfonogr. von Mantispa, GEB]CAfi*s ZeiUehr, f, Entom. I.
1838, 8. 147—173-
Semblis Fabr. (in part ; species of HemerMus and of Raphidia
L,) Maxillary palps filiform, more slender towards apex, with last
joint often shorter than the rest. Prothorax quadrate, longer than
mesothorax and metathorax. Anterior wings larger than posterior.
Antennae filiform or setaceous, long. Tarsi with two claws, without
intermediate pulvillus.
Sub-genus: SeTnUia (species of Semblis Fabb.), Sialis Latb.
Ooelli none. Penultimate joint of tarsus cordata Wings deflected.
Antennse simple, almost of length of body.
8p. Semhlts lutaria Fabb., ffemerobius lulariua L., R(BSBL, Int, n. Inteet.
aqaat, CI. n. Tab. xiu. ; Gu^bin, Iconogr,, Ins. PI. 62, fig. 6. The laira
lives in water, and has on .each side of the abdomen eight jointed threads,
which serve for respiration ; it leaves the water to change into a motionlesB
pupa in the ground. Gomp. on this genus SuoKOW in Hbusinoeb's
Zeittchr. f, organ. Phynk, 11. i8a8. s. 165 — i*ji, Tab. n., and Piotbt,
M4m. 9wr le genre SicUis de Latbbillx, Ann. det Se. ntU. sec. S^rie, v.
1836. Zoologie, pp. 69 — 80. PL 3.
Sub-genus : Cordalis Latr. Ocelli three. Penultimate joint of
tarsi undivided, small. Antennse simple, shorter than body. Head
broader than thorax, transverse. Mandibles exsert, in males veiy
long, at rest decussata
Sp. Semblia comuta, Raphidia comvla L., Ifemeroibiut oomutui Fabb., Dx
Gbsb, Im. lu. PI. 17, figs. I, a; habitat North America. (Mandibles
of females denticulate within, thicker ; of males, subulate;, with only one
tooth, small, internal before the apex.)
Sub-genus : Chattliodes Latb. Ocelli three. Penultimate joint
of tarsi undivided, small. Antennsa serrate or pectinate on the inside,
shorter than body. Head of breadth of thorax, angustate posteriorly.
Mandibles moderate, little or not exsert.
Sp. Semblia peetinieomiB Fabb., ffemerobiiu pectinieomia L., Db 6bbb, Ina.
in. Tab. 27, fig. 3, Cuv. R. Ani. id. Ul., Ins. PI. 105, fig. 2 ; habitat in
North America.
Note. — ^There is a species in the Leyden Museum from Java, with pro-
thorax orange, wings brown, with round transparent whitish spots, antennse
VOL. I. 27
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418 CLASS VIII.
serrated. To thia Db Haan gave the generic name Sialii. That genus
(Sialii Db H. not Fabb.) is the same as Neuromua KAKBUBf NeuropL
p. 441, which ought not to be separated from ChauUodet; the species which
I have indicated is Newr. ruficofUs of Rambub. Another genus of
Rambub, DUar, imknown to me, with pectinate antennte in males, appears
to be reUtted, but has a greater affinity, as I coi^jecture from the descrip-
tion and figure, with the ffemerobii,
Semerohiua L., Fabr. (exclusive of species). PaJps with last
joint thicker, elongo-ovate. Prothorax small, shorter than the
other segments of thorax. Wings ample, lanceolate, with costal
border broad. Antennae filiform or setaceous, mostly long. Tarsi
with two claws, and intermediate pulvillus.
a) Ocelli none.
Sub-genera : JlemeroUtu Leach, Drepanopteryx Leach, MegcUo-
mu8 RAHa, Siayra Burh., Chn/sopa Leach, PdysUxchotea Bubm.
Nymphes Leach. (With antennse short, wings narrow, elongate;
habit of lion-ant)
Sp. Hemer6bvM perla L., (Chrytopa perla Leaoh,)^ IUbs. Im, ui. Tsh, 21,
fig. 5, Cut. R. Ani, id. ilL, Ins. PI. 103, fig. 3; ffemerobita chrysopa L.
(Ohrytopa reticviaia), Roebel, Ins. ni. Tab. ai^ fig. 4 ; comp. N. Slabbsb,
Verh. der Haari. McuxUeha^j, x. a, bL 387 — 41a, with fig.
These broad-winged insects are provided with eyes shining with gold
dming life; the delicately latticed wings, as transparent as fine gauze,
present the colours of the rainbow. The larva lives on plants, and
resembles that of the lion-ant ; it feeds on leaf-lice, and is covered with a
mealy substance, to which the cast-off skins of the leaf-lice adhere, so that
the true form .of the insect is often quite irrecognisable ; the small, oval,
white eggs are seated on long thin pedicles upon twigs and leaves.
b) Ocelli three.
Osmyliu Latb.
Nemoptera Latr., Nematoptera BuRM. (species of Panorpa L.,
Fabr.) Head produced into a rostrum. Maxillary palps shorter
than maxillse, filiform, more slender towards the apex, with three
small terminal joints ; external lobe of maxillaB subulate, biarticu-
1 6. T. BoHNSiDBB, SymJbciUB ad Monographiam gtneris Chrysopa Lkaoh, Yratis-
lavisB, 1 85 1. Bvo. c. tab. pictis.
W. F. Evans, Monograph of ike BritUh Species of the genus Ckrysops, Trcensad.
of the BMomol. Soc. of London. 1847, Vol. v. p. 71, PL 9, 10,
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INSECTA. 419
late, palpiform. Ligula porrect, pilose. AntennaB filiform, long.
Ocelli none. Mesothorax large. Inferior wings narrow, veiy long,
sometimes resembling filaments.
Sp. Nemoplera extema Dumi^ru., Cons. g4n, 9. 1. Int. PI. ay, fig. 7, Gu^rik,
Iconogr., Ins. PI. 61, fig. i ; — Nemopt. lutUanica Klug, Panorpa hipennis
iLLio., Gbhmab, Fauna Ins. Ewrop, Fasc. i. 'Dftb. 16, &o. Species firom
Africa, aonthem Europe and western Asia.
Myrmdeon L. (in part), Fabr., Latk., Myrmecoleon BuRM.
Maxillary palps mostly shorter than labial ; external lobe of max-
illsB forming an internal Particulate palp. Antennas not longer
than thorax, gradually thickened into a club towards the apex,
arcuate. Ocelli iione. Prothorax short, transverse; mesothorax
large. Wings lanceolate, deflected. Abdomen elongate, cylindrical.
Feet short.
Sp. Myrmdeon formUaarvm L., R^umub, Ins. vi. PI. 34, fig. 7, B(BSBL,
Ins. ni. 'Dab. 20, figs. 95, a6, Pakzbb, DeiOsehl. Ins. Heft 95, Tab. 11.
The Utrya of this insect is known under the name of Lion-ant, and is
fiff-fiuned for its canning. It has six feet, very huge npper jaws, a flat
head, and a large oyal abdomen. It moves slowly, and almost always
backwards. Unable to pursue its prey rapidly, it seeks to compass it by
subtlety. To that end the lion-ant digs fimnel-shaped holes in the sand, by
moving constantly backwards in still smaller circles, and casting out the
sand with its head. During this work it is almost entirely concealed
beneath the sand, the head and jaws alone projecting above it. Should the
lion-ant in the meanwhile encounter a little pebble, too large for his jaws,
he creeps out of the sand, endeavours to push his abdomen under the
pebble, attempts in this way to creep with his load to the edge of the
funnel, and repeats the operation until he finally succeeds, although the
pebble repeatedly falls off behind him. When his work is completed, the
lion-ant lurks under the sand in the middle of the conical cavity, and
pushes his two upper jaws out of the sand like open shears, continually
during the night ; by day he does this more seldom, and only when an
insect falls into the hole. Any small insect that ventures incautiously to
the edge of this pitfall, and stumbles in the dry sand, is effectually griped
by the pincers ; should it try to escape, the lion-ant greets it with a shower
of sand, and causes it again to tumble down. If the lion-ant has clutched
the insect, he shakes and tosses it about, sucks it dry, and casts the
empty body as lumber out of his hole. The lion-ant may be most aptly
compared to a spider, and like as this weaves nets proportioned to its size,
so he makes his funnels larger the larger he grows. When full-grown, he
spins grains of sand together for an abode, almost perfectly spherical, in
which he changes to a pupa, and from which after four weeks he makes
his appearance as the perfect insect which flies by night. Comp. on the
economy of this insect R^UMUB, Ins. yi. pp. 336 — 386, PI. 3a — 34,
27—2
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420 CLASS VIII.
BoiSEL, Im, in. pp. 95—116, p. H4, Ac., Bonnet, (Euvrei, Tom. ix.
1781, 8vo. pp. 405—413-
This species occurs in Europe principally in southern Unds, yet even in
some parts of Holland also ; in England it seems not to be found. There
are still other European species of this genus (as Myrmdeon pichun Fabb.,
Gbbhab, Fauna Int. Eitrop, Fasa vni. Tab. 16, from Dalmatia and
southern Russia^ also from the south of France, Myrm, formica lynx L.,
BoiSBL, Ifu, m. Tab. 21, fig. 2, &c.), and many exotic species, amongst
which one is very large, with spotted wings, from the Gape of Good Hope^
Myrm. tpeciotut {Myrm, libelloidet L., et auctor. in part) &o.
Aacalaphus Fabb. Antennae long (longer than body), termi-
nated abruptly by an obconical capittdum. Abdomen of length of
thorax. Head hirsute. Eyes in most divided by a transverse
furrow. Remaining characters of the preceding genus.
Bp. Aacalaphtu bar^Kvnu Bubm., Ascal. iialicm Dum&il, Cons. gin. «. I,
Ins, PI. 26, fig. 2 i-^AscaL italicue Fabb., Panzbb, DeiUsc/U, Ins, Heft 3,
Tab. 2Z, &c.
Aseal. Napoleo, Lxfbbvbb, from New Holl. See Gu^Bnr, Magas, de
ZooL 1843, Ins, PL 93, where many sub-genera are proposed, partly also
received by Rahbub, which must be here omitted. Comp. Rambub,
Neuropt, pp. 341—364.
B. Neuroptera wUh meUanorphosis incomplde.
Tarsi with joints often fewer than three.
Fanuly XXXI. LibdluUrue. Antennae short, with few (7 — 8)
joints, the two basal thicker, passing into an acuminate seta com-
posed of the terminal joints. Wings reticulate, with numerous
transverse nervures. Mandibles and maxillae entirely covered by
the labia or margins of the exsertile head ; palps not exsert. La-
bium large, trilobed, with broad lobes. Eyes large ; ocelli three.
Tarsi triarticulate. Larvae aquatic.
LiheUcB or water-nymphs. These insects in all the states of their
life are very greedy and rapacious. The antennse of the perfect
insect have commonly from five to seven joints. The feet are short,
especially the middle pair, and bent forwards. Ooptdation occurs
in a remarkable manner, the extremity of the abdomen in the fe-
male being bent round like a ring towards the breast of the male.
Here are found, on the ventral sur&ce of the second ring of the
abdomen, the external organs of generation, a penis placed between
two homy booklets, which in jEshna and LUbeUvla is perforated by
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INSECTTA. 421
an efferent canal arising from two small yeaides filled with a yellow
fluid : these glands are wanting in Agrion\ The larvae aod nymphs
live in the water, and resemble the perfect insects. The mider-Up
has the form of a mask or hood, and oovers the mouth ; anteriorly it
terminates in two hooks'. It is bent round and set upon a long
flat pedicle, which the animal can move forward rapidly, whereon
the bending disappears ; thus the insect supplies itself with a move-
able forceps for seizing its prey unawares at a distance with the
hooks at the extremity of the under-lip. (See figures in Swamheb-
D^K, JBibl. not. Tab. xn. ^g. 5, B^aumub^ In8. vl PL 36, fig. 4m,
figs. 10—14, PL 37, fig& 4, 6, 7, PL 38, figs. 6—8, Ecesel, Ina. n.,
Ins, aqwU. CL II. Tab. nL figs. 7b, fig. 8c, Tab. ix. figs. 3, 4 j Lyonet,
(Efwr, posthmne, PL 18, figs. 13—15). The abdomen of the larva
ends either with five homy plates, which can be moved from and to
each other, or with three fin-like laminsB ; it was in these laminae
(in the genus Agrion) that Cabus first discovered the circulation of
the blood ; see above, p. 259.
With LIKN.SUS this &mily forms only a single genus, LibdhUa.
Comp. on this fcuxuly Van dsb Linden, Agrionea Bononiemet deteri^pUB.
OjnucoU 9cieiUifici, Bologna, 1833, Tom. IV. p. loi ; ejuscL jEtknce Bononi-
enset, ibid. p. 158, &c. ; ejusd. MonographicB LibdhUinarum Eurapaofvm
specimen. Bruxellis, 1835 ; Toussaint db Ghasfentibb, LibeUvUna
Suropace deacriptas d depictce, cum Tabulis 48 color. Lipsiss 1840, 4to ;
B. DE Selts Longohakps, MoHographie des LibellvUnes d^ Europe, Paris
et BruxeUes. 1840, 8yo; and the same author's different short oommuni-
cations in the 6th and 7th parts of the BtUletina de VAccuL de$ Se, de
BruxeUei,
A. Intermediate lobe of labium very small, lateral lobes very
broad, without articulate style,
Ldbdlula Fabr. Eyes large, often contiguous. Ocelli disposed
in a triangle. Wings horizontal, expanded, anterior different from
^ Fuller information concerning these parts may be found in H. Bathke, De
lAbeUvlaa^m partibui genUalibiu. Cum Tab. sen. iii. Regiomonti, 1833. 4to. Because
the penia is not connected with the testes, of which the vasa deferetUia terminate at the
ninth ring of the abdomen, Bathkb ¥ras of opinion that the union obeerred in the
Libellulince is not a true copulation, but only precedes it. VoN Siebold, however,
detected spermatozoa in the male at the base of the abdomen, so that the male, by
bending round the abdomen towards the thorax previous to' copulation, seems to deposit
the sperma there. Wiegmann's Archiv f, Naturgesch, 1838, p. 375.
' These nippers are the palpi of the under-lip ; see A. Bbull^ Observations sur
la houche des LibeUuUnes, Ann. dela Soc, Entomal, 11. 1833, pp. 343 — 351, PI. 13.
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422 CLASS VIII. .
posterior in the structure of the cells, supplied with basal cell
resembling a right-angled triangle inverted.
Sub-genera: LibelliUa, Epiikeca Chaspent. (ZibeUa Selys).
CorduUa Leach. (EpophthcUmia Bubmeisteb, and other genera of
Bambub ; see his Nevroptires, pp. 24 — 152.)
ff
Sp. lAbdkda depretta L., Rcbsbl, Ina, n., Tna. aguat. CL n. Tab. vi. fig. 4,
Tab. vn. %. 3, DuMiauL, Cons, g, 9. l. Ins. PI. aS, fig. 6;—LQfeUula
cenea L. {CorduUa cmea), Pakzeb, DeuUcM. Ins, Heft 88, Tab. lo.
To the right-angled triangle, at the base of the fore-wings, as character
of this genus, I directed attention in a short communication in the Bijdroffen
tot de fwtuiurk, WeUenschappen. in. bl. 335—339. PI. ni., and Atm. <ies Sc
not. TV, 1823, pp. 434 — 437. PL XI. h,
B. Intermediate lobe of labium broader than lateral ; lateral
lobes supplied with moveable appendix at apex, sometimes com-*
posed of many joints.
.^hna Fabk. Eyes large, approximate. Ocelli often disposed
in a transverse row. Wings expanded.
a) With eyes cantiguaua.
Sub-genera : Anax Leach, jMshna, ejusd, GyTw^artJtiha, Rambub.
Sp. JEskna grandis, LibeUula grandis L., B<s&KL, Ins, L 1. Tab. iv. &c.
h) With eyes approximcUef riot corUigumu, or remote; toith head
tromsverse.
Sub-genus : Lindenia De Haan, Y. D. Hoev. {DiaatcOomma
BXJBM.)
N^ote, — Here are to be referred sub-genera Gamphua, CorchUegcu-
ter and FetcUttara Leaoh, and some others proposed by Rahbxjb, opi
cit. p. 153.
Agrion Fabr. Eyes remote. Head transverse, much broader
than thorax. Wings narrow, erect when the insect is seated.
Middle lobe of labium deeply bifid. Abdomen slender, cylindrical,
often very long.
Sp. Agrion puella, LibeUvla pwsUa L. (is a coUectiye name for different
European species, and thus it is arbitrary to adopt it as a synonym), Agrum
furcatam Ghabfekt. ; Bossel, Ins. L L Tab. xi. fig. 7 ;— Agrion virgo,
Libdlula virgo L., B(ES. 1. 1. Tab. ix. figs. 5, 6, &c.
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INSECTA. 423
Sub-genera: Lesiea^ CalepUryXj Agrion Leaoh, Mecistogaster
Rambub, and some others of this author, L 1. p. 217.
Family XXXII. EphemerincB. Parts of mouth soft. Antennae
very short, subulate, triarticulate. Tarsi quinquearticulate, with
first joint mostly very short, scarcely distinguishable. Posterior
wings less than anterior, sometimes none. Abdomen terminated by
two or three filaments.
Ephefmera L. (Characters of the family.)
May-Flies, bcmk-hait. The larvsB and nymphs live at the bottom
of the water between stones, or in the ooze ; some dig horizontal
passages, in which they move very rapidly. In these two states
they live two or three years before changing into perfect insects ;
they proceed from very small eggs. The larva has always three
threads at the abdomen, of which the middle one in many species
disappears after the metamorphosis ; it is distinguished especially
from the perfect insect by longer antennae, by stronger upper jaws,
and by external respiratory organs at the first six rings of the abdo-
men ; the air-tubes appear as free threads or little plumes, or are
united by membrane to form a foliated lamina. Some larvsB have
on each side of every ring two such plates. The change fix)m
nymph to perfect insect occurs in Ephemera SvKMfnmerdamii {Palin-
genia longicauda Bubm.) so suddenly that there is almost at the
same moment a creeping and a flying creature. In other species, as
in those observed by Rcesel, the change takes place more slowly.
They moult once again in their perfect condition j and if we be
waUdng by the banks of the water over which they are flying we
ofien see this moulting performed on om* clothes.
In the middle of summer the May-Fly takes wing towards even-
ing. The time of flight diflers in diflerent species. That which
SwAHMERDAM described shews itseK nearly three days in succession,
about St John's day, in the month of May, and often in such quan-
tities that it is as difficult to count these little creatures as &Iling
snow-flakes. A great many fall into the water a prey to fishes, and
at that time, especially at Dordrecht, the roach is noted as being
peculiarly fiit and good Hence the name hamMaU, oeveraas (in
some parts of France la mmine). According to the relation of
Scopou they are collected by the country-people in Garniola to
manure the land {Entomol. CamioL p. 264).
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424 CLASS YIII.
SwAMMSBDAM, in the species deBcribed by him, did not ohaerve
the oopulation, and supposes that the eggs, as in fishes, are first laid,
and afterwards fertilised bj the male. RiAxnnTB doubted this,
because the eggs sink in water. But Bosbel, De Geeb, and later
writers, have actually observed the copulation in other species;
and thus it is improbable that the species of Swaxmebdam forms
an exception on this point.
EphemercB on the whole live as perfect insects but a short time,
some not even a single day. Once however I kept some specimens
of Ephemera dipiera (Cloe diptera Pictet) for some days alive under
a glass, two for more than a week
^^EphemeroB la/rvcB ruxUmt in aquU ; voUuUea facUs fruvmJbwr
gcvudio, tmo scepe eodemque die nuptias, puerperia et exaeqfdas
ceUbranUs'' I^njsmub, Syst, ruUwr, i. ed. zu. p. 906.
Gomp. on this family:
Bphemeri vita of Afbeddingk v<m.*» menachen leven door J. Swaxxxbdam .
Aznflterdam, 1675 kL Svo. m. afb. (For the fiMsts see B^bd der natuur, I.
bL ^S-f— 270, Tab. xm — ^xv.)
R^UMUB, Mhn. p. aervir d rHiMt. det Ins, VZ. pp. 457—523 ; Db GxxBy
M6m, n. pp. 617—660. PL x6— x8.
F. J. Piotbt, Hiti. not, det InaeOea N^vropt^ret, Seconde Monogrmphie,
FamiUe da Bphimerinei, Qenhve, 1845. 870. avec pL
A. Wings toith tramsverse nervwreafeWy posterior often obsolete,
i) Compowid eyes two JxAh in nudes andfemalei.
Sub-genera : Oligoneuria Pictet (Wings four), C(Bni8 Stephens,
Pictet (Wings two).
a) Eyetfcmr m males; the two accessory large, ptUdUUs.
Sub-genus Cloe Leach, Bubm., Pictet. Posterior wings very
small or obsolete. Caudal setn two.
B. Wings fowTy wUh trcmsverse nervures numerous,
i) TuH) accessory eyes in males, as in the preceding genus.
Sub-genus : Potamcmikus Ticr«r. Caudal setee threa
a) Compound eyes only two in both sexes.
Sub-genus : Bcstis Leach, Pictet. Caudal setce twa
Sub-genera : Ephemera, Falingenia Burjl, Pkttet. Caudal setso
in females three, in males the intermediate one sometimes shorter or
abortive.
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INSEGTA. 425
Sp. Ephemera Swammerdamia Latb., PaUnffenia Umgieauda PiomET, Swax-
MKBD. 1. .1. TBh. xm. ;— Ephemera wlgata 1,,, Houttutm, lieUmiH, ffii-
toire, xn« Stuk. PL 94, fig. i, Piotbt, L L PI. a, &c.
Family XXXDI. Perlarice. Mandibles distinct, denticulate.
Ma^cillffi with palps five-jointed, filiform or setaceous, and accessory
palp, mostly biarticulate (ffalea) ; labial palps triarticulate ; lingula
emarginate. Antennae long, setaceous, moltiarticulate. Wings
(insect at rest) cruciate, incumbent on the back horizontally;
inferior broader than superior, folded longitudinally like a fan.
Tarsi with three joints, the last with two claws, and intermediate
pulvillus.
Perla Geoffr. (Species of Fhryganea L., Semblia Fabb. in
part). Characters of family.
These insects were first distinguished by Geoffroy as a separate
genus, whilst LiKNiEUS placed them amongst the PhrygcmeoBy to
which they have some relation by the broad posterior wings. Till
within a few years it was generally supposed by writers that the
Perlas have a complete metamorphosis, in consequence of an imper-
fect observation of Nollet, recorded by Reaumuk. Pictbt however
has announced the metamorphosis to be incomplete ; the larvae do
not live in cases, but mostly in running water, by preference under
stones. Some have at the thorax external respiratoiy organs ; others
breathe through closed stigrnata, which have the same situation as
the future apertures of the perfect insect, and take in the air by
endasmose. They have constantly two threads at the hinder part of
the body, which sometimes disappear in the perfect insect, but in
many species persist at that period. The males of some species have
short wings, merely rvdimentari/,
Comp. on this family, of which more than a hundred species from all
parts of the world are now known (through later investigations and dis-
coveries), PiOTET, M4m. ewr lea larve$ de Nhnown, Ann, det Sc. not, zxvi.
1833, pp. 369 — 391. PL 14, 15; ejusd. M4m, iwr lea fMtamorphoeee dee
PerUa, Ann. dea Sc. not. xxvni. 1833, pp. 44—^5, PI. v. vi. and his Hist,
not, dea Inaectea Nivroptirea, Premiire Monograpkie, FamiUe dea Perlidea,
Gren^ye, 1841. 8vo.
Sub-genera: Perla Latr., mihi (Kollaria Pictet, Eusthenia
Westwood, Pterona/rcya Newm., Perla Pictet), Capnia Pictet.
Anemahi/ra mihi {NemaJUywra Latb., Pictet).
NoiU* — This fiunily with the following approaches by natural affi-
nity very nearly to the Orthoptera.
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426 CLASS VIII.
Family XXXIV. TermttimB. Mandibles homy, strong, den-
ticulate. Maxillae furnished with an appendage or lobe resembling
a galea, and with a palp four or five-jointed. Antennae filiform or
.setaceous, of the size of thorax at least, composed of more than ten
joints. Wings deflected or incumbent, with few transverse nervares,
sometimes absent; posterior wings not broader than anterior, in
many less than anterior. Tarsi with two to four joints.
Psocus Latr. Antennae setaceous, long, with joints cylindrical,
elongate. Maxillary palps four-jointed, labial none. Wings either
none or deflected, unequal, posterior less. Tarsi in most with only
two, in some with three joints.
Sp. Paoeus ptdaatoritu, Termes puUaiorium L., De Gkeb, Itu. vn. PI. iv.
fig. I, SuLZEB, Kenm. d. Int. Tab. xxii. fig. 144 ; the woodrUnut {fundlviM) ;
the body pale yeUow, with yellow eyes. This insect is commonly found on
wood, upon walls, in old papers, &c. ; it has superficially a resemblance to
a louse, but runs very rapidly. The Latin specific name is derived from
the idea that the ticking in old furniture and door-posts, which superstition
often receives as a bad omen, is caused by this insect ; on the contrary,
they are small beetles which cause the tapping sound. An anatomical
investigation of this little animal was published by NiTZSOH in Gsbmab u.
ZiKOKEN, Magaein der Entomol, iv. iSai, s. a76---a8a, Tab. 11.
Other winged species are met with on trees, as Psocua quadripwnoUUiU
Latb., Panzbb, IhuttehZ. Int. Heft 94, Tab. 33, &c.
Here belong sub-genera Apropos Leach (^Troctes Bubm., Airopas
and LachenUa Westw.), Thyrsophorua Bubbl, CoecUius Cubtis.
Termes L. (in part), Latr. Labium quadrifid, with palps three-
jointed. Antennae short, mostly moniliform. Wings narrow,
incumbent horizontally on the body, longer than body, deciduous,
sometimes none. Tarsi with four joints.
These insects bear, in different tongues, the name of fi>hite ants;
they live, like these, in society together, and in warm countries
cause incredible damage. Of the wood-work which they destroy, they
leave the surface untouched, so that every thing externally preserves
its form, but £bl11s to pieces on the slightest touch The hardest
wood cannot resist these animab ; glass, metal, and stones alone do
they leave uninjured. Some species live underground, others in
trees ; some erect pyramidal mounds above the sur£su;e, which in
Guinea and other regions of the African coast are raised to the
height of twelve feet, and are so stable that they may be mounted
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INSBCTA. 427
with safety {^wmeBfaUkiM L., Termea bellicams Skeathk.) Below,
in the middle of the building, is the residence of the female (mon-
strous is the dimension of the abdomen in a pregnant female which
lays perhaps 80,000 eggs) ; round about this residence are cells for
^6 ^S^ magazines for food, <bc., all orderly arranged. The upper-
most part of the hill-like dwelling is massive, to give more firm-
ness, and to guard the funily from the influence of weather and the
attacks of foes. It is generally supposed that the labour of building
in these societies is performed by the larvee, but perhaps it is by the
sexless individuals who have not got their wings ; they have no
eyes. There are still other wingless individuals, also without eyes,
but with larger jaws ; these are commonly called soldiera, because
they ward off attacks and always stand at guard. When these
insects attain their perfect state they leave the dwelling at night-
time j on the day following their dried wings fall oS, and they are
in great part eaten up by birds, lizards, and even by the negroes.
A fertilised female is borne off by the larvae to a dwelling, and there
she lays her eggs. According to Smeatbkan the workers would
seem to carry off a male and a female with them, in order to shut
them up and feed them for copulation.
Comp. Bm£athman in PhUowphical TravMLctiont, Vol. Lxzi. 1781. The
figures &re copied m part by Blumsnbaoh in Abbildungen nalturkitUmt-
cher GtgeMtande, Nob. 9, 10.
Add genus Embia Latr. Head broader than thorax. Wings
not longer than body. Tarsi triarticulate.
Order X. Hemiptera.
Hexapod Insects, with four wings, all membranous or anterior
coriaceous at base, thicker. Mouth with articulate rostrum (labium),
which ensheathes setse (mandibles and maxillae) ; palps none. Me-
tamorphosis, with few exceptions, incomplete.
Half-ioinged {Hemiptera from tifitaw half, and wrepoy). This
order originated indeed with Linn.£US, but is now limited differently,
since the Orthoptera have been separated from it as a distinct order.
Fab&icius, who also placed the flea here, named the half-winged
insects Rhyngota, which name Burheister properly changed into
Bhyrkcokta. On this order, amongst other works, we notice :
J. C. Fabricu Sy sterna Rhyngotorum, Brunsvigse, 1803, 8vo;
Hist. naU dea Insectea Hemipt^res, par G. J. B. Akyot et Audinet
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428 CLASS Till.
Sebville, Oworage (neoompagnJe de Planche$, Paris, 1843, 8vo (a part
of the SmUs d Buffon by Bobet).
As illustrated works : C. Stoll, deaden en WarUsen, Amsterdam,
1788, 4to (two parts with Dutch and French text) ; J. R Schellen-
BEBG, Cimicum in HeioetioR aquie et terns degentimn genue, Turici,
1800, 8vo (with 14 coloured plates) ; J. F. Wolff, AbhUdungen der
Wanzen, v. Hefte, Erlangen, 1800—1811, 4toj C. W. HAmr, Die
foanzenartige Ineecten, getreu nach der NaJbwr abgebildet tmd be-
schrieben, Niimberg, 1831, and subsequently, since 1836 continued
by HsBBICH-ScSiElFFEB.
Leon Dufoub treated of the anatomy of these insects in a
monograph, entitled JRecherches ancUomiquea et phyeioL efwr les
Hemipt^res, Paris, 1833, 4to av. 19 pi.
The Hemiptera are commonly provided with four wings, of whidi
the anterior are leathery at the base, thick, and not transparent, and
at the point membranous {hemelytra), or are membranous like the
posterior wings, but often stronger and larger than these. The mouth
consists of a sucker composed of threads and a case. The case is tubu-
lar, grooved above, and consists of joints ; it corresponds to the un-
der-lip of other insects. The small upper-lip becoming thinner for-
wards, covers the base of the sucker. In the groove formed above by
the turnover margins of the under-lip, there lie in appearance three
setse, but the middle one is double (the two imder jaws {mcKdUcB)
and the two lateral threads are the upper jaws'). The maxillary
palps are entirely absent ; so also are the labial palps, or these last
are only in quite a rudimentary state*. Thus the beak of the
hymenoptera is constructed for sucking. The fine threads (setw
hcmatelU, mandHmUB, maanUai) make a wound in the parts of plants
or animals, on the fluid or blood of which they feed, the fluid
ascending between the threads to the oesophagus above.
The antenna have commonly only four or five joints, extremely
seldom more than eleven. Many species have two or three simple
eyes. There are never more, but often fewer, than three joints in
^ G. B. Trevibanus was the first who distinguished the four threads in Oimex
t^/ipei (Awnalen der WeUerauischm GeteUtchafi, i Bd. a Heft, 1809. s. 171).
Saviont has shewn the analogy with the oral organs of other insects, and figured
the four filaments in Cimex nigricomia, and in a Nepa, M6m. », I, ani, 9. vert. I.
1816. PI. IV.
" SAViaHT, 1. oit. PI. IV. fig. 30, p. 37.
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INSECTA. 429
the feet The metamorphosis is incomplete, with only few excep-
tions (the males of the gall-insects and the genus Alewvdes),
With respect to the natural affinity of this order, and its place in
a natural arrangement of insects, we remark that it agrees with
the Orthoptera in the imperfect metamorphosis, but differs from this
order very greatly in the oral organs and in the internal structure.
The last &mily of the preceding order, the Perlof and Termites, on
the contrary, have a position very dose to the Orthoptera ; and it is
only with reluctance that we interrupt this natural transition by
interposing the Hemiptera between them. But the ffemiptera,
whatever position be allotted them, must always stand by them-
selves as a very deviating group of insect& The absence of
maxillary palps might perhaps be considered to be their most
important character, which elsewhere in the class of insects are
more constant in their presence than the labial palps \ Some
species of Hemiptera hamoptera (the genus FlcUa) have some
resemblance to the Lepidoptera, which however, is rather an
external similarity {cmalogy) than a true relationship {affinity).
The intestinal canal of the hemiptera is long, (see above, p. 253).
In many Cicada/rice {Cicaday Ledra, Cercopis, &c) there is an
arrangement, which was formerly considered to be a return of the
intestine into the stomach, but which has been since described by
DoTEBE in a more satisfactory manner; the tortuous intestinal
canal does turn back to the stomach and attaches itself to it, to
leave it subsequently at another point; but it does not open into
the stomach, but merely runs along its wall, between the coats of
the oigan*. In some GeocoriacB {Scutelleray Fentatoma, and some
species of Coreus) the inferior part of the stomach before the
insertion of the va^a wrinaria is tortuous, and consists of four
(in Coreus moflrginaJbis of two) grooves or semi-canals, which have
transverse white rings or folds of even width, between which there
is a very thin and transparent membrane'. Salivaiy glands are
ordinarily much developed; mostly more than one pair are pre-
sent; they are situated on each side of the intestinal canal, and
open into the oesophagus; in the leaf-lice they appear to be wanting.
' Batzebubo indeed describes in Cicada, a three-jointed maxillary palp (Afediz.
ZodL II. Tab. aj, figs. i6, Ty); but this part^ according to Bubmbibtkb, is a jointed
homy process, to which muscles are attached.
> Ann. des 8c, not, le S^rie. Tome xi. 1859, PP* 81—85. ^ i-
' Ramdohb has given to this part the name of Wamenmoffm,
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430 CLASS VIII.
The disagreeable smell, peculiar to most heteroptera, seems for their
protection, and is caused by a fluid that escapes through two aper-
tures, one on each side, beneath the thorax at the insertion of the
third pair of feet. According to Lj^on Dufoxtb this fluid is
secreted by a small round or oval pouch, commonly of a yellow
colour, which is situated in the abdomen; it is rarely double.
Many genera of hefmiptera are met with only in warm countries,
and the European species form but a small part of the numerous
order.
Section I. Homoptera. Elytra deflected, in most coriaceous or
membranous, similar to posterior wings, but larger and stronger.
Bostnun rising from the inferior part of head, inflected beneath the
breast between the bases of the feet. Thorax gibbous, with first
segment shorter.
In this division, which as well as the second, Westwood ranks as
a distinct order, the upper wings are constantly of a similar substance,
and not half-homy or leatheiy and half-membranous. The antenme
consist in many of more than six joints. The beak lies on the
breast {rostrum pectoraUy All live on juice of plants. The females
often have an instrument for laying their eggs, composed of three
filaments, toothed plates or saws, enclosed between two valves.
A. Metamorphosis (in males at least) complete.
Family XXXV. Goccina ( GaUimecta Reaum.) Males winged,
with mostly two wings, without rostrum. Wings without cells,
with only two longitudinal nervures. AntennsB mostly with nine
to eleven joints.
Females (one genus excepted) apterous, attaching themselves to
trees or herbs at the time of parturition, sometimes assuming the
form of a gall, their own vesicular dead bodies covering their eggs.
GoM-inaecta, These insects were thus named by Reauhub from a
superficial resemblance of the pregnant female to a gaU-nut. (See
above, p. 384). The joints of the tarsus have not been given by
us amongst the characters on account of the minuteness of the
objects and the prevalent uncertainty; for whilst many writers {ex,
gr, Latbeille and Westwood) ascribe one joint alone to this family,
and distinguish it principally by this, others assign three (Bouche
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INSECTA. 431
even foiir) joints. The character also of a single claw alone at the
end of the tarsus is not constant.
Comp. on this family, RiAUHUB, Mim, pour terv, d Fffist. des Ins. Tom.
IV. {Mhn, I et 2), pp. i — na; Batzsbcro, Medk, Zool. 11. pp. 214 — 218,
Font, Ins, ni. s. 188.
Coccus L. (in part). Wings two and often poisers in males;
females apterous. Abdomen in males with two terminal sete.
Rostrum of females short, with long exsertile setsB concealed and
inflected in abdomen.-
Sub-genera : Leccmvumi Iluo., Coccus Bubm., Pseudococcus West.
To this diviaion, and indeed especially to Leeanium, what we advanced
in the characters of the fiunily respecting the females has reference ; to this
genus alone the name QoUintecta is appropriate. To it belongs Coccus
Ilicis li,, Lecanium Hicia Illio., BIbaumtjh, op. cit. PL v. ; in the sonth of
Europe and in thd East, — the kermes of the Arabians, from which word the
appellation karmoz^, crimson, for a red colour, is derived. This insect,
formerly used in medicine, now only as a dye, has however lost much of
its value since the introduction of the CochenUU from America (Mexico),
which first came to Europe in 1526. The last-named species. Coccus cacti
L., lives upon the Nopal, Cactus coccineUifer (Optmtia coccindlifera
DiOAND.), cultivated with that view. See figures of the insect in Dum^bil,
Consid, gen, s, I. Ins, PI. 39, fig. 2, Brandt u. Batzeb. Medkdn, Zool. n.
Tab. a6, figs. 5—12, 16, 17, Bubmxibteb, Ha/ndb. der Entomol, 11. Taf.
n. fig. I. It is computed that 70,000 dried insects go to a pound of
' cocheniUe, and formerly 880,000 pounds of this dye were imported. Comp.
on the Cochenille, NaiuurUjke Historic van de CocheniUe, bewezen met
authentique Docum^enten {door M. Db Ruubsohbb), Amsterd. 1729, 8vo;
Thubbt de MiiroNViLLB, Traiti de la cuUure du Nopal et de V^ducation
de la CocheniUe, Av. pi. Cap fran9ai8, Paris et Bordeaux, 1787. 8vo.
On the introduction of this insect into our East Indian possessions, a
report may be found in the Alg. Konst-en Letterbode, 1839. No. 30.
Another spedee, Coccus fieus Fabb. (probably a species of Lecanium),
lives in Bengal on different species of Fieus and other plants; the. young
insects are seated dose together on the young shoots, and round about the
place where they have sucked themselves fast, there drops a thick fluid,
which hardens into a tough transparent substance, the gum-resin, gummi-
lacea. The dye of this substance is used under the name of lac (lac-dye,
Uk-IoJx), as a very beautiful substitute for cochenille, and the shdl-lae
deprived of the colouring matter as a component of lac- varnish, of sealing-
wax, and as an isolating body in electrical apparatus. See on this insect
Kebb, Phil, Tramsact, Vol. Lxxi. for the year 1781, pp. 374— 38«.
Coccus manniparus Ehbenb., Symbol, phys., Ins. Dec. i. Tab. 10; comp.
Geiobb'b Journal der Phannaeie, Juli 1830; on Tamarix mannifera in the
neighbourhood of Sinai, &c.
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432 CLASS YIIl.
McMrgarodes, See Lansbown Goldino, Trcmaact. of the Linn,
Soc XVI. 1.
Calypticus CosTA, Aspidiotus BouCHE, BuRM.
Mo7wphlebti8 Leach, Burm.
Porphyrophora Brandt.
Dorthesia Bosc, Wings two and poisers in males; females
apterous. Antennae of male nine-jointed, long, setaceous; of
female eight-jointed, short, subulate. Abdomen of male terminated
by a pencil of cilia.
Gomp. Boso, in Observations mr la Physique, &c., par KoziSB. 1781.
pp. 171 — 173, D0BTHB8, ibid. xzvi. 1786, pp. 107 — III, BuBinBiSTBB,
1. 1. figs. 6, II.
Aleurodes {Aleyrodes) Latr.
Note, — Genus Alewrodea {Aleyrodes) Latr., seems rightly to be
referred to this family by Burheisteb. Metamorphosis complete,
as in males of Cocci; pupa quiescent, folliculate. Also anterior
wings sufficiently similar to wings of Cocci, But the number of
wings (four), which are not wanting in females, the rostrum too, with
which, as it seems, the males also are furnished, seem to indicate
the separation of this genus from the Cocdna^ and accordingly
Latreille joined it with the Aphidii, and lately, Habtig with the
Payllw; Westwood, however, to avoid a difficulty, preferred creating
a small family from the single genus.
Sp. Aleyrodes Chdidonii Latb., Phalama {mnea) proleteUa L., RiAUiruB,
Ins, n. PL 25, figs. 1—7, Bubmkistrb, L 1. Tab. i. fig. la (larva), Tab. n.
fig. 7 (imago), Westwood, IrUrod. to mod. Class., in plate of Title to
YoL I., Haetig in Qebmae's ZeUscKr. /. Entcmil, in. p. 371, Tab. i.
^* y>i 31 (Antenna, wing).
B. Metamorphosis incomplete.
Family XXXVI. Aphidii s. Phytophthires* Wings four, with
nervures longitudinal alone, often ramose or bifid; in some no
wings. Tarsi with two joints, the last biunguiculate. Rostrum in
both sexes. Antennas mostly with six to ten joints, longer than head.
Aphis L. AntennsB with not more than six or seven joints.
First joint of tarsus very short. Fore-wings with marginal stigma.
Abdomen mostly supplied posteriorly with two conical tubercles.
In most species some imagines are apterous, others winged.
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IN8ECTA. 433
Leaf-licey FUmt-Ucej Fucerons. These animals live together on
different plants and trees, and mostly in veiy large numbers. They
do not leap, but run commonly very slowly. From the hind part
of the body there often drops a transparent honey-sweet fluid, of
which ants are very fond and on that account are often found
in the neighbourhood of the leaf-lice. These suck the sap of the
plants, and some species, by puncturing the leaves or leaf-stalks,
produce excrescences or swellings occasionally hollow, which ai'e
filled with a great number of these animals and often with a
considerable quantity of the sweet sap. The sticky fluid on leaves,
known by the name of honey-dew, is caused by leaf-Uce. Most
species are covered with a powdery substance or with white threads,
(a peculiar secretion of their body).
To counteract the excessive multiplication of leaf-Hce, in the great
economy of nature, a number of enemies are on the watch, not
merely ichneumons (see above, p. 378), but insects especially which
feed on them and devour large quantities, the pupse of some diptera,
of hemeroini (p. 418), of beetles (Goccinellce), &c.
Leeuwenhoeck had already noticed that plant-lice are viviparous,
that they also lay eggs was first discovered by Lyonet ; but the
succession of generations, the descendants of mothers and grand-
mothers which are viviparous and fruitful without copulation, was
first discovered by Bonnet ; see above, pp. 263, 264.
Comp. on this group of insects, as numerous in species a^ interesting in
their economy, E^UMUB, Ins. ni. Mim. a. pp. 481—350; C. Bonnet,
Traits cTInaectol. Tom. i. Paris, 1745 ; Db Gseb, Mim. p, aervir A rffiH,
d^Ins, in. pp. 19 — 129, and for the systematic arrangement, Habtig,
Verauch einer Eintkeilung der PflcmztidaiUe in Gbbhab's ZeUsckr, /. die
ErUomol, in. 1841, s. 359 — 376, and especially J. H. Kaltenbach, Mono-
graphie der Familien der PJlamenlaiUe, Mit Abbild. Aachen, 1843, 8vo;
also T. Waulsb, DeacripUons of Aphides, Ann. of NcA. Hist. sec. Ser. iv.
p. 202, V. 1850, pp. 14— a8, (169— a8i, 388—395, VI. pp. 41—48,
118 — ia«.
Some species live on the roots of planta. They have no wings ; BouoH^
however (according to Ratzbbvbo, Forst-Ins. in. s. a 16), discovered two
species of Kkitdbius that were winged. These species may be ooUected
provisionally under the name of :
Bhizopthiridiurn nob.
Here belongs the genus Bhizobius BUBIC (a name already given to a
genus of Ccieoptera), and the genera Pa/radduSf Trama and Forda Y.
Hetden^ EnUmol. BeUrdge in Abhandl. der Senckenb. CfeaeUseh. n. 1837,
s. 491 — -295, Rhuoterus Habtio, according to Kaltenbach, does not
differ from Forda V. Hetden.
VOL. I. 28
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434 CLASS VIII.
By hi the greatest number of species live above ground on branches
and leaves. Amongst these both winged and wingless individuals occur in
the same species. Hie winged individuals have constantly three simple
eyes (Kaltbnbaoh).
a) Wings horizontaL Abdomen without oomides or tubercles. (An-
tenn» short.)
Phylloxera Boteb de Fonsc., Habtig. Antemue triarticulatey
with last joint fusiform, elongate, emarginate, obscurely annulate.
Anterior wings with cubital nerve undivided.
VacuTM Y. Hetden, EIaltekb. Antenn» five-jointed. Anterior
wingB with cubital nerve bifid.
b) Wings deflected, obliquely erect above the body.
* Cubital nerve undivided.
FhlceapthMiditim mihi, Chermes Habtio. (Species from genus
Chermea L.) Antenn» shorty thickish, filiform, with five joints. Feet
short Abdomen without comiGle&
Note. — Cfhermet is a name by no means to be given to the modem genua,
which Fleeting all the other Linnsean species, contains one alone {C,
Ahidu) ; the diagnosis of Linhjbub himself teaches this. On Uus and
other allied species comp. especially Batzbbubo, FonA-Int, in. pp. 195 —
005, Tab. zn. lliese insects appear to be oviparous alone ; oopulation
has not yet been f
Pemphigus HABna. (Spea Chermea Bubil)
Tetira/newra Habtig. (Antennae with six joints.)
* Cubital nerve sending off one or two branches.
Schiz(mefwra Habtio, Kaltenb., Myssoooylus Blot {Erioaoma
Leach, in part). Cubital nerve bifid Antennas short, sex-articulate.
Comides of abdomen none or obsolete.
Sp. Aphis laniffera Hausmann, Briotoma mali Lbaoh, ftc.
Lctchnua Illio., Bubm. Cubital nerve trifid. Antennso sex-articu-
late, shorter than body. . Feet long. Short tubercles alone in
abdomen, in place of cornicles. Feet, especially the posterior, long.
Rostrum sometimes very long.
Sp. Laeknm fagi, ApkU fagi L., RiAUM. Ina. m. PI. 26, figs. 1—6. The
abdomen of this species |'" in size is covered with very long white wool ;
it looks like a bit of white flue ; this species is very often seen on beeches
in the groves at the Hague.
Aphia Iiiua., Bubil Cubital nerve bifid AntenncB sept-articulate,
longer than body. Feet long. Abdomen with two comides.
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INSECTA. 436
8p. Aphi§ rotas li., B£kuinni, L L PL ii, figs. 1—4 ; DuMiBiLy Com, ffSn.
ff. I, Ins, PI. 39, fig. 3, Ac
PsyUa Geopfr., Latr., Chermes L. (exclusive of Cherm. Abie-
tis). AntennsB with ten joints^ the last bisetose. Tarsi with two
equal joints. Wings deflected, ample, with nerves branched.
Imagines always winged.
Le(rf-mieher8, FaUe Uaf-Uee, Fantet pueerons. The larvn are ooyered
with a woolly ooaiing, just like leaf-lice. Hie perfect insect has the
kaJbitus of a small cicada^ the long antennss excepted. Gomp. lUUuM . Ina.
in. Mim. z. pp. 351 — 36a ; Db Gesb, M^m, m. pp. 130—157. Sp.
Ptylla pyri, Chermsi pyri li,, Ratzxbubo, Font-Ifueelen, m. Tab. xz.
fig. «, Ac
Livia Latb., Diraphia Ilug. Antennie short, thick. Head
anteriorly bifid
Sp. Lma juneorwn Latb., DnxiBiL, Cora, gin. t. I, Iw, PI. 39, fig. 5;
GlBMULB, Faun. Ins. Swrop, vi. Tab. ii.
Add sab-genus LiviUa Cubtib, Wsstw.
Family XXXVH CicadaricB. Wings four deflected, anterior
often coriaceous coloured elytra. Tarsi almost always triarticulate^
in a few biarticulate. Antenna short, with three or seven joints,
the last setaceous. Females furnished with serrated borer.
Comp. on this family Gbbmulb, Magtm. der EnUmol. m. pp. 177—997,
IV. pp. I — 106.
A. GicadaricB mutce. No musical organs. Antennae with three
joints. Eyes two or none. Tarsi triarticulate.
Phalanx I. CicadeUoB Latr. {Cicadellina and Membracina
Burmeister). Antennae inserted between the eyes. Posterior feet
saltatory.
A) Troihonx not produced posteriorly above the abdomen.
Scutellum distinct. Head horizontal, with frons mostly (Hrected
forwards.
t) Posterior tibiae spinoee or serrate.
a) Ocelli none.
Typhlocyba Germar. Body elongate. Posterior feet very long.
Oomp. especially Hbbbioh Sohaffbb, DeuUcM. Ins. Heft 164. Snudl
insects, leaping, mostly yellow or green. European species numerous,
amongst which are Cicada Ulmi and Cicada Rosa L.
28—2
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436 CLASS viii.
b) Ocelli two.
TeU^ania Latb., Cicada Fabr.
* With ocelli on yertex.
Sub-genera : Ceroopis Fabr. in part, ApkropharaGEEM., ClastopUra
Germ., Tettigania Germ., Burm., EuaccurUhus Germ., Ledra Fabr.,
Ledropsia White, Fenthimca Germ., Gypona Germ., Xerophlcsa
Germ.^
Sp. Oercopis tpwnaria Fabb., Aphro^j^ara tpumaria GxBM., Oic. spufMria
L., B(BSBL, Ins, n. Loetut. Tab. zxni. f. i — 4 ; the larva and nymph
liye in a white froth (vulg. cticJsoo-»pU) found in spring-time on different
plants, especially on willow-trees.
TeUigonia viridU, Cicada viridis L., &c.
** With ocelli in the maiigin of frona.
Jaama Fabr. (Add sub-genera Ccdidia, Fcwapia, Eupelia, Aoo-
cephalu8y Byihoacopug Germ.)
tt) Posterior tibioe unarmed.
Ulopa Fall., Germ. Head broad, with eyes somewhat pro-
minent.
JEtlialia Germ., Burm. {JEtalion Latr.) Head deflected an-
teriorly.
Sp. JBthalia reUcuUOa, Cicada reticulaia L., De Gebb, Int. in. PL 33,
figs. 15, 16, Latbiillb in Humboldt et Bokpland, Obterv, de Zool.
Hd^AnOt, comp. I. PI. 24, figs. J 2, 13. Habit, in South America; forms
the transition to the following section.
B) Prothoraz produced posteriorly above the abdomen, often
covering the scutellum, in some the wings also. Head deflected^
with frons inferior ; ocelli two in vertex.
Membracia Latr, {CentrotuSf Membracis [Ranatra Lesson*],
Damta Fabr.).
Add sub-genera; Lamproptera Germ., Bocydivm Latr., and
several others, on which comp. Burmeister, I. L il pp. 127, &c. and
Amyot ei Serville, Nefwr. pp. 532 — 553.
^ Besides these, many other genera have been added by Amtot et Sebvu:.ls,
Hid, not, de$ In$, lUfMpt, which frx>m our limited space we cannot record.
* This name was given long before to a genus of the water-bugs {Nepa linearis).
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INSECTA. 487
Spi J#<m5nicif fMOa Fabb., Oicada foUaia L., SvoLL» dead. Tab. r.
fig. i; DniciKiL, ConjL ^n. t. ;. /m. PL 38, fig. 3. Habit, in Nor b
America^ like many speciei of this aubdxvigion.
Phalanx II. FvlgoreUoB. Antennae inserted under the eyes.
Head compressed at the sides, marginate^ often elongated ante-
riorly, inflated. Ocelli^ in some none^ in others two, approximate.
A. Two palpiform appendages at the base of antennse.
Cclax Germas, Ottocerua KiBBY, BuRM. {ffynnia Bubm. in
part).
B. AntemuB simple.
Tettigometra Latr.
laaus Fabr.
Sub-genera: Eurybrachia GufeiK, Chrethrwra Hope, Ancyra
White.
Sp. Aneym appendieulaia Wmn, Wistwood, CaMntt of Orienial BiUomol,
PL 36, fig. 5.
Delphax Fabb.
Aacraca Latb. (and Ugyopa GuiRiNl)
Anotia KiBBT.
Derle Fabb.
Sub-genera : Mysidia, Lydda, Zeugma, Thraeioy Phcenioe, Faiara,
Cenchrea Wbstw.
Fkaa Fabb.
Sub-genera : FcecUopiera Latb., Ricania Geril, Cixia Latb. &c
Lystra Fabb.
Sub-genera : Apheena Git^bik, Poiocera Lapobtil
Fvlgora Fabb., Bubm.
Sp. Fvlgora latemaria L., Bcbsel, Ins, n. Loeud. Tab. ig, 30; SroLi^
Cicad. Tab. I. fig. i ; the Surinam lantern-hearer; the head terminates in
front in a bladder-like hollow procefls, the wings yellow-brown, with
brownish stripes meeting reticolately ; the under- wings with a large
yellow eye-like spot, bordered with blaok, with two round, white, oonfluent
rings in the middle. It is one of the larger insects of this order (the tips
of the fore-wings are f,",6'" asunder ; the body with the yesicular process
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438 CLASS VIII.
of the head a" 6'" long). Aooording to Meuak this ineecl at night-time
difiEusee a dear light ; which, however, is denied by later writers. Ab
little, according to Bowsma (Afin. qf not, HitA, xiv. p. 437), is the Chinese
species, Fulgora canddaria L., phosphorescent. In Europe, especially in
the southern parts, Fvlgora Ewopoea L., Pteudopluina ewropcea, is met
with. See a fig. in Panzeb, DeuUcM, Ing. Heft ao, Tab. 16, and NaUtr-
fancher, ix. Tab. 11.
B. Cicadce stridulantes {CtcadcB manmfsrce). Males ftimished
with a musical organ at the base of abdomen on each side.
Antennas with seven joints. Ocelli three. Tarsi in some (genns
Tibicen Latr.) biarticulate.
Gicada Oliv. (species from genus Cicada L., CicadcB manni-
fircB), Tetttgonia Fabr.
Cicadas. These insects are found in warm countries alone, in
woody districts especially. The females, when about to lay their
eggs, perforate the branches by means of two serrated homy plates^
enclosed between two jointed valves and supported at the back by a
grooved homy part, formed of two pieces which have coalesced \
The larvffi on leaving the egg quit the place, and betake themselves
underground, where they suck the roots. Here too they change
into nymphs.
The ancients made much of these animals and considered them to
be favourites of the Muses, whilst living on dew at the top of high
trees they heralded the approach of summer with their clear song,
and resembling the gods, as is sung in an Anacreontic ode, had like
them no blood in their bodies'.
The musical organ of the male is situated on each side at the base
of the abdomen and covered by a valve. A homy, triangular
partition corresponding to the eniothoraa^ separates the two lateral
cavities. Each of these, when viewed from the ventral aspect, presents
in front a white folded membrane, but down at the bottom, a stretched,
thin and transparent lanjina, which Reaumur calls le mirair, the
hoking-glasa {Mem, v. PL 17, figs. 1, 2, 3, 6 m). If the part be
opened from the dorsal sur&ce, another folded membrane is seen, the
^ R^UMUB, Mim. V. pp. 170 — 178, PL II, Westwood, Introd. n. p. 414;
DOTBRE ^ves a difiEerent explanation of the instrument, and thinks that the lateral
plates serve for fixing and that the middle piece by pushing penetrates the wood. See
Ann, det Sc, not. sec. S^rie, vn. 1837, Zool, pp. 193 — 199, PI. 8.
* See Abistoteles, de But, Animal, L. v. a 34 (Ed. ScHN.) vulgo c. 30.
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IN8SCTA. 439
drunk-membraoke {La iimbale, 1 1 figs. 5, 6, 9, 11 tyt), which is moved
by a yeiy strong muscle arisiiig from the partition mentioned above.
When these muscles contract powerfully on each side and then
suddenly relax, a vibratory motion arises, producing a sound which
is swelled by an air-vesicle, a vesicular trachea, that lies near the
drum. The sound is the so-called song of the cicada, which is
peculiar to the males, whence Xenabghus extolled the fortune of
these animals, whose wives are dumb.
The so-named manTta, not to be confounded with that which
sei-ved the Israelite^ for food in the Arabian desert, is an exudation
from ash-trees in consequence of the puncture of the Cicada omL
Sp. Cieada <ymi L., BoBSiL, Int. n. Zocuttar, Tab. xxv. figs, i, ^, Tab.
XXVI. figs. 3, 5 ; Brakdt and IUtzsbubo, Mediz. Zool. n. Tab. xxvi. figs.
1—4 ;— Cicada fraadrU, TeUigonta Praadni Tabr., B<B8BL, ibid. Tab. xxv.
fig. 4, XXVI. fig. 4, figs. 6 — 8, from the south of France^ Italy, &c.
Section II. Heteroptera. Elytra coriaceous at the base, mem-
branous at the apex. Eostrum frontal, rising from the anterior part
of the head.
The insects of this division are the only ones properly to which
the name of hemiptera is applicable. Some amongst them suck the
blood of other animals; others live, like those of the preceding
division, on the juices of plants. The anterior s^ment of the
thorax is much larger than the other two. The antennsB have
never more than five and ordinarily only four joints.
Comp. on tins division F. L. de la Pobtb, Ssaai dune ^lauyUaHon
systemaHque de Fardre dee HemipUrea; Gu^bin, Magae, de Zool, 1839, Ine.
PL 51—55-
Family XXXVIII. SydrocanacB. Antennse inserted under
the eyes, recondite, shorter than head or scarcely of length of head,
three- or four-jointed. Tarsi mostly biarticulate. Eyes mostly
large. Bostrum short.
WcUer-bii^s, They live in fresh-water. In the following &mily
also are some species which live in water, but which do not swim as
these do, with the exception of the genus Galffulus Latb., which
Westwood refers to the Geocarism,
I. Ocelli none.
A. Two anterior feet recurved downwards, with thighs not in-
crassated ; two posterior long, pilose, natatory, sometimes destitute
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440 CLASS VIII.
of tenninal claws* Body depresso-cylindrical or oblongo-ovatC;
thickish.
Notanecta L.
t Anterior tarsi uniartiodftte.
a) SouteUum not distincty oovered by prothorax.
CorioM Gboffr.^ Latb., (Sigara Fabb.)
Sp. Notanecta itriata L., Bcesel, /m. m. Tab. 99, fig. d; Sohsllenbsbo,
OinUc. Tab. ix. ;— (7or. Qtoffroyii Lsaoh, Cbr. imnetota Bubm., Bcbsxl, L L
figs. a» b, fto.
li) Scntelliun diBtinct.
Sigavra Leach (species of Sigoura Fabr.)
Sp. Sigara nUnuta Fabjl,, Notonecta minvUmma L.
ft Anterior tarsi triarticulate. Scatellttm distinct.
a) Posterior feet terminated by two daws.
Ploa Steph., BuBiL, Plea Lbach.
Sp. Notonecta minuHieifna Fabb., (not L.), Panzeb, Deutaehl, Ins. 1, Tab.
20 ; only 1"' long.
h) Posterior feet witb tarsi biarticulate, without daws.
Notonecta Fabr., (exclo. N, mimUisaima), Leach, Bubil
Sp. Notonecta glavca L., Bcbsel, Ins, m. Tab. 37; Sohbllenb. Cfimic.
Tab. X. ; Panzeb, Deutschl. Ins, Heft 3, Tab. 20, Water-bug; it swims on
its badE, and stings seyerely^.
B. Two anterior feet cheliform, the tibia and tarsus forming a
curved hook to be lodged upon the thighs, incrassated and grooved
on their anterior inner margin {N^)a L.).
Naucoris Geoffr., Fabb. AntennsB quadriarticulate. Head
almost as broad as thorax. Anterior tarsi uniarticulate. Bodj
ovate, depressed.
Sp. Na/ueoris cimicdides, Nepa cimicotdes li,, B<esel, Ins, m. Tab. aS;
Sohbllekb. Cimic, Tab. xii. ; Panzeb, DetUschl. Ins, Heft 95, Tab. 16, &c.
Betoatoma Latr. Antennae quadriarticulate. Head small, nar-
rower than thorax, triangular. Anterior tarsi biarticulate. Body
ovate, depressed.
^ Gomp. on this sub-genus Leach in Lin, Transact, Vol xii. 1818, pp. 10—18.
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INSECTA. 441
Sp. Bdottcma grande, Nepa grandu L., Bobskl, Ins. m. Tab. 26, from
South America; Bdostoma indieum Le Pslbtixb H Ssby., Stoll,
Wanteen, PL vn. fig. 4, from the West Indies. The largest species of this
order. Hie stiliform appendage at the abdomen is not, according to
SpnroitA, for respiration, and thus does not correspond to the filaments at
the abdomen of Nf^pa, Gir^iN, lUtme zooloffique^ 1839, -^^l^ P« n^*
Note, — Sub-genera Diphnycfma and SphoBrodema Lapobtb are
scarcely distinct.
Sp. Bdo9Uma rtuHeum, Nepa rwHca Fabs., Stoll, Wantzen, PL vn. ^, vi.
The females carry the eggs on their back, like PipcB amongst the
BatraeK&ldea.
Nepa mihi {Nepa and Ranatra Fabr.) AntennaB triarticiilate.
All the tarsi with only one distinct joint. Abdomen provided with
respiratory appendage formed of two setae.
* Body oval, depressed.
Sub-genus Nepa Fabr., Latr.
Sp. Nepa cmerta L., Swammbbdah, Bibl, not. Tab. m. fig. 4 ; Roessl, /n#.
m. Tab. la; Sohsllxkbebo, Cimie. Tab. XIT. ; DuM^. Com. gSn. 8, 1 /n#.
PL 37> ^* ^ ) Watencorpion, yery common with ns in ditches and fish-
ponds; the eggs, already described and figured by Swamickbdaic, have
seven threads at the end, which cause them to resemble the plumed seeds of
Syngenistte.
* * Body slender, elongate.
Sub-genus Ranatra Fabr., Latr.
Sp. Nepa linearis L., Swamicbbdah, L L fig. 9; Boksel, Ins, ni. Tab. 2$,
fto. ; rarer than the former species.
II. Ocelli two.
Oalgvhia Latr. Ocelli two, frontal.
Sp. Cfalguhu ocuUOus, Naucoris oeuiata Fabb., CuyiEB, R, Ani. 6d. HhuL,
Ins, PL 93, fig. 4.
MoTumyx Laporte.
Pelogonus Latr. Kostrum elongate, with sheath triarticulate,
second joint long. Ocelli in vertex.
Sp. PelogoMis marginatus Latb., Gebmab, Ftmn, Ins, Ewrop, Fasc. xi.
Tab. 33 ; Cut. R. Ani, id. iU. PL 93, fig. 3 ; on the coasts of the south of
France aad of Spain. This genus makes the transition to Salda, but can
scarcely, on account of the insertion and the shortness of the antenne, be
placed elsewhere than in this family.
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442 CLAJSS YIII.
Family XXXIX. OeocoriscB (Aurocorisa Westw.). Antennae
exsert, longer than head, inserted towards the inner margin of ejes.
Tarsi with three joints, with first joint very short in some. Ros-
trum often long, sometimes produced beyond the thorax.
Mostly terrestrial Hemiptera ; few aqnatic, living on the surfiace
of the water, or inhabiting the banks.
Land hugs. This numerous &mily forms with Liknjbus only one
genus {Cimex)y out of which, from the discovery of new fonns and
the more accurate distinction of the species formerly known, fiu*
more than 100 genera have already been formed by modem writers.
Phalanx I. Oculata {Acanthitdce Westw.) Sheath of rostrum
triarticulate. Feet long, slender, terminated by two long claws.
AntennsB quadriarticulate. Eyes large, protuberant. Ocelli two,
in vertex. Head without distinct neck, separated by a small stric-
ture from thorax.
SaJda Fabr., (in part) BuRM., Acanthia Latr. Antennse fili-
form, of the length of head and thorax.
Sp. Salda UUoralis Fabjl (and LygasuB saUatoriut Fabb.), Oimez liUoraUi
L. (and dm. Mltatoritu L.), Be Gikr, Mim, m. PL 14, figs. 17, 18;
Wolff, Waneen, Tab. vm. fig. 74 ; this species leaps like a frotli-Gicad&.
Leptopua Latr. Antennse setaceous, nearly of the length of
body, with third joint very long. Anterior feet spinose.
Sp. LeptopuB littoralii Latb., Cuv. R, AwL id, iU, PI. 95, ^g. i, ftc.
Phalanx II. Hydrodromica, {Hydrometridoe Westw., Ploteres
Latr.). Sheath of rostrum triarticulate. Head almost of the
length of thorax, without distinct neck. Ocelli sometimes none.
Four posterior feet longer than anterior, formed for going on the
water, inserted at the sides of thorax, remote from each other.
Head ovato-oblong or linear, covered beneath with a silky down.
AntennsB moderate, filiform, quadriarticulate, and either with a
single accessory joint, very short between the second and third, or
with three, interposed between each of the others. Tarsi with two
or three joints, the last joint shortest.
A. Claws of tarsi inserted in a fissure at the inside before the
extremity of last joint.
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INSECTA. 443
Oerris Latr., Fabr. previously, Hydrometra Fabr., Burm. (in
part). Four posterior feet very long, remote from anterior. First
joint of antennsB very long.
Sp. Qtrrit lacutirU, Cimex laeutHt L., Stoll, WmUeen, Tab. iz. fig. 63 ;
DvuiSLTL, Com. gin, s. l. Ins, PL 36, fig. 6. This species often oontinaes
wingless ; comp. Qerria eanalium, L^n Dufoub, Anat, dea ffSmipt, PL V.
fig. 59 ; — Oerria rufoscuteUata Latb., Stoll, L L Tab. 15, fig. 108 ; Cuv. R.
Ani, id, UL, Itu, PL g2, fig. 5, &c.
^0^060^ ESCHSCH.
Note, — Species small, apterous, or with little elytra, with abdomen short,
conical, in tropical seas and the Pacific ; comp. Esohsoholtz, Bntomogro'
phim, I* Lieferong, Berlin, iSas, 8yo. pp. 106 — iii. Tab. m. figs. 3 — ^5.
Are they larvas (and pnpaB) of species of genns Oerris t Comp. the figure,
much resembling a larva recently exduded from the egg, of L^N BuFOUB,
Anat, d, Hhwpt, PL xv. f. 178.
Velia Latr. Feet snb-equally distant, the middle rather longer
than the rest.
Velia West., Bubil First joint of antemite longer than rest.
Posterior femora inorassate, spinose.
Sp. Vdia riwlorum Latb., Qtrris ritmlorum Fabb., Ent. SyH,, Hyd/ro-
metra rvmdorwn Fabs., jS^. Shyng,, Cinr. R. Ani. %d, ill., Ins. PL 91,
fig. 4> &c«
Hydroessa Bubil, Mi4!r(yi>€Ua Westw. Last joint of antennie
longer than rest
Sp. Vdia pygmaxt, Jj6oTSf Dtjfoub, &c.
B. Claws of tarsi apical.
Hebrua Westw., Burm.
Hydrcyimtra Latr. (species of Hydrometra Fabr.), Lirnnobatea
BuRM. Body narrow, elongate ; head protracted beyond the eyes.
Antennas with third and fourth joints elongate, third very long.
Feet very slender, long.
Sp. Bydrometra stagnorum, Oimex stagnorum L., Sohellbitb. CHmM. Tab. IX.
fig. 3 ; DUM^B. Cons, gin, s. I. Ins. PI. 37, fig. 5. This small needle-shaped
insect has much resemblance to the genus Plotaria.
Phalanx III. NvdicoUia {Beduviidas West.). Sheath of ros-
trum triarticulate. Bostrum incurved. Head abruptly attenuated
towards the base, forming a distinct neck. Antenn» with four
joints, sometime annulate or divided into secondary joints. Feet
long, with tarsi short, triarticidate, with two terminal claws.
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444 CLASS VIII.
A. Anterior feet raptorial, short, with cox9e elongate, thicker
than thighs. (Four posterior feet very slender).
Ploiarta SCOPOLI, Latr. {Gerris Fabr., Burm.], Emesa Fabr.,
BuRM. (and Emesodema Spinola, Amtot). Antennae longer than
bodj, setaceous.
Sp. Ploiaria vagaburuUi, Oimex vagabwnduB L., Soexluekbebo, Cimie.
Tab. vm.; Cuvikb, jB. Ani. 4d. ill., Ins, PL 92, fig. 3; a J'" long, feet
ringed, yellow-white, and black ; this little oreatme baa a staggeiing gait,
like some long-legged gnats ^.
B. Anterior feet not raptorial, with coxcb short.
Zelu3 Fabr. Body linear, with very long feet.
Mytx/ria BuRM. Elytra entirely membranous. First joint of
tarsi indistinct, almost entirely retracted upon the tibia.
Beduviua Fabr. Body oblongo-oval.
Sub-genera : Ndlna Latr., and Eedtimus ejusd.
Sp. Beduvitu penonatus Fabb., Oimex perwmahu L., Dx Geeb, M^, m.
PL 15, figs. T— 9; SoHELLXNBXBG, Cimtc, Tab. vn. fig. i; Guv. R. Ani.
id. HI., In*. PL 92, fig. I ; black, nine lines long ; the lanra covered with
dust and flue, coated as it were, liyes in houses and feeds on insects; —
Bedumus amanua Gu^bin {Rcduv. hullattu Leyd. Mus.), Gu^. Iconogr.,
Im. PL 56, fig. 17; i". i'" long, abdomen vermilion red, concaye above,
cuticularly expanded with five black-blue round spots on each side ; Java.
Note. — ^Many sub-genera of more recent authors have been omitted, on
which comp. Btjbmeibteb (ffandb. der Sntamol. u. pp. 117 — 147), also
Amtot et Sebvillb {ffiet. not. dee Himipt. pp. 311 — 393 ; many of the
names barbarous, taken from the tongues of Indians, Hebrews and Chinese).
^ There is a still smaller native species, about i}"' long, which has darker wings,
and the black rings on the feet broader than the yellow-white, Ploiaria erraHca,
Oerrii erraticus KLua, Oimex culictformi* Db Geeb? m. PL 17, figs, i — 8. The
genus Ploiaria was established by J. A. SOOFOLI in his DelieuB Flora et Fauna Ineu-
hriea (Tidni, 1786, folio), and indeed upon a wingless species from which the genus
Emetodema was afterwards formed. This species, magnified, is several times figured in
SooPOLi*s work under the name of Ploiaria domestica, I. Tab. xxiv. figs, i, 3, n. Tab.
xxm. ; the Ploioria (data in the third part, Tab. zxv. fig. VI, figured by Count
Cabtiluonb, agrees perfectly with the species noted by me as Ploiaria erratica.
Lesson, in his lUushraHone de Zooloffie, PL 53, has figured, if I mistake not, the
Ploiaria dometiiea of Soopoli under the name of Ploiaria vagabumda.
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INSECTA. 445
Holcpttlus. Lepeleties and Sert. AntennsB long, hirsute.
Feet pilose, posterior tibiae very hirsute. Tarsi large. Body de-
pressed, with broad elytra.
Sp. ffolopHlua unut LnsL., Ds Latobtb, Quia., Uagat, de Zool. 1839,
Itii. PL 54, fig. I ; CuY. R, Anu, id, HI., Im. PL 99, fig. 9. In the hAbitaa
this genus approaches to Tingit,
Sub-genus; FtUocnenma Westwood.
Phalanx IV. Membranacea {CtmicidcB and Tinffidoi Westw.).
Sheath of rostrum triarticulate. Bostmm received in a sulcus be-
neath the head. Antennas quadriarticulate. Tarsi with three
joints or two. Body oval or suborbicular, depressed.
A. Antennae setaceous, with last two joints slender.
Acanthia Fabr., Cimex Latb.
Sp. CfkMx leOulariuB L., De Gxbb, Uim. m. PL 17, figs. ^—15 ; Sohkl-
LXirBBRO, Oimie. Tab. vi. fig. i ; DuKiB. 0ms, gin, a, I Im, FL 37, fig.
2 ; de weegluis, la punaUe, die JBetiwame, the hug; a}'" long, wingless,
brown-red, with short felty hairs, head small, thorax broad and short
This, too well-known, insect is able, according to the obsenrattons of Dl
Gub and L]foN Dufottb, to live for a long time without food^, and is not
killed by the severest oold of vnnter, but rerives on the first warmth firom
its torpor. This species would seem, according to Scx>FOLi, also to occur
winged; but here, in all probability, we have another species before us;,
which Schilling described as Oimex dometUau in 1833 (OKXB's/m, 1834,
s. 738, 739*). There are still other species (wingless) on pigeons, bats and
swallows, which JiNTNS has described and figured; Annali </ Nai, Hid,
m. 1839, p. a4i, PL I.
B. Antennae filiform or thicker towards the apex,
t Feet all similar, ambulatory.
Aradus Fabr. AntennsB filiform.
Sp. AraduM deprema Fabb., Sohxllbnbsbo, CinUe, Tab. y. f^, 1 (CbfVM
apiniger), Wolff, Wanunj Tab. zni. fig. 113; lives under the baik of
trees.
Ncie, — Here seems to belong the genus PKUa Latb.
Tingis Fabr. Antennae capitate.
1 Uknx DuFOtJB found three bugs, which were enclosed in a glass, to continue aUve
without food for a whole year.
' A species of Xylwsorit DvwoXTR, according to Bubmiibtxb.
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446 CLASS VIII.
8p. Tingis echU Fabb., Wolff, Waneen, Tab. xm. fig. 134, &c. ; they live
on leaves and flowers, and suck the sap of plants.
ft Anterior feet raptorial, with tibisB incraceated, canaliculate
beneath. Last joint of Antennn incrafisated.
%rti8 Fabr.
Sub-genera ; Phymata Latb., Mcbcroeephalm Swed., Latb.
Sp. ByrtU crauipea Fabb., Paxtzeb, DeuUchl. In$, Heft 33, Tab. 34; Guy.
It. AwL, id, ill,, Int. PL 91, fig. 4;— tS^fftf eroM) Oimezerotua Jj., Wolff,
Wanzen, Tib. ix. fig. 83; Sulzbb, Kmwi. der Im, Tab. XL f. 71, ftc.
On other sub-genera of this phalanx see the works cited above of
BuBMEiSTEB and AXTOT.
Phalanx Y. Longilahra. Sheath of roBtmm qnadriarticnlate.
Labrom elongate. Antennae with four or with five joints (in a few
three). Tarsi with three distinct joints, terminated by two claws
and two plantulsa (little membranous appendages).
A. Scntellum small or moderate, not produced as far as the
middle of abdomen.
t Antemus setaceous, with last joint very slender. (Ocelli
none.)
Capsus nob. {Capsus and Miria Fabb.)
a) First two joints of antenns thick; second very long, equalling or
surpassing in length the last two, which are very slender.
Heterotoma Latb.
Sp. Oapaus 9pUrie(>mis Fabb., Sohbllbkb. Oimie, Tab. m. fig. 4; PAKZBBy
IhiUachL Int. Heft 11. Tab. 16; Cut. JR. Ani., 4d. ill, Im. PL 91, fig. 3.
5) Second joint of antennn incrassated at the apex only.
Capsua Fabel Thorax broader posteriorly.
e) Second joint of antennn not incrassated.
Miria Fabb., Latb. {Miris, Fhytocoria Fall., Bubil) Thorax
broader posteriorly, trapeziform.
Aatemma Latb. {HaUicua Kahn, Bubil) Thorax transversely
quadrate.
ft Antennn filiform or capitate.
LygcBua Fabr.
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IKSECTA. 447
Sub-genus : Pyrrhochoria Fall., Bubm. Ooelli nona
Sp. L^gcnu apteruB Fabb., CHmex aptenu L., Wolff, Wansef^ Tab. tl fig.
loi; SroLL, Wcmtzm, PI. 15, fig. 103; 4}'" long, black with a red margin
round the thorax, red sbield-coYerB with a black point at the base, and a
larger ronnd spot of the same oolonr in the middle ; the membranous portion
of the shidd-ooTers is very short; wings commonly wanting.
Sub-genera furnished with eyes : Lyganu, Xylocoria LioK DuiouB
and others, on which oomp. Burmeisteb, L L pp. 288 — 299.
Coreus Fabr. Ocelli two. Body ovate or oblong. Antennae
straight.
Sub-genera: Alydua Fabb., Goriaua Fall, and others, for which
see BusMEiSTEB, L L pp. 301, <fec.
8p. Comu quadrahu Fabb., Wolff, Wanzen, Tab. vn. fig. 67 ; Hbbbioh
SoHiBFFBB DeuUcKl. Iiu, Heft 118, Tab. la, 9oc.
Leptocorisa Latr. {Oerria in part; add Berytus Fabr., Neidea
Fabr.) Ooelli two. Body elongate, linear. Antenna in some
straight, in others broken.
Sp. JBeryhu Hpularius Fabb., Oivnex HpuUmm L., DuxiBEL, Com. gH, $,
k Int. PI. 36, fig. 7; Wolff, Women, Tab. zx. fig. 198.
B. Scutellum large, produced as far as the middle of abdomen.
Cimex Fabr. Antennae mostly five-articulate {Pentatoma Oliv.,
Latr.), in some four-articulate {Tesserotoma Lepelet., Serv.)
NoU, — The generic name OimeK I judged formerly, with Latbbillb, to
be attributable to Cimeat leettdariua, aooordingto the rule of Linhjiub (PhU,
Baton, § 246), which says that if a reoeived genus ie to be sub-divided, the
preyioQsly common name ought to remain with the species most commonly
known (species Yulgatissima); but it seems advisable to give a received
name to numerous species rather than to a few. Besides, a new name
would be required, for that given by OuviXB, Pentoiama, is inapplicable to
several species of this genus, and the distinction taken from the number of
joints of the antemuB is artificial, often severing affinities. For many
genera of modem writers, not here recorded, see Bdbmbibtkb, L L p. 347,
Ac., and Axtot et Sbbvillb, Himipt. pp. 73 — i8a.
Qensok of Fabbictos, Cydnma, Hatya^ Bdeaaa, ^lia, Cimex.
Sp. €H,fMx haecorum L., Panzeb, Deuttchl, In$, Heft 33, Tab. 18 ; WoLrr,
Woneen, Tab. yl fig. 57; 4}'" long, the head and thorax red-brown with a
shade of green, and many impressed minute black points ; the scutellum
yellow at the point, the abdomen black above with yellow spots at the
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448 CLASS VIII.
margin, yellow-brown below; on numy plants in Holland, and common
everywhere in Germany ; I poesess a specimen from the Gape of Good Hope
little different. — Olmex rufipet L., Wolff, Wamen, Tab. i. fig. 9 ; Batzk-
BUBQ, Forst-Ins. ni. Tab. xi. fig. 3 ; 6'" long, the thorax with an ear-like
process on each side, scutellum and feet red ; — Cimex acuminaitu L., JSlia
acuminata Fabe., Panzbb, JkutscM, Ins, Heft 32, Tab. 17; Wolff,
Wamen, Tab. n. fig. 19, &c.
C. Scutellum produced to the apex of abdomen, sometimes
covering the wings entirely.
ScuteUera Lam. {Tetyra Fabr.)
For many genera of modem writers see Burmeisteb, L L pp. 382
— 396 and Amyot et Seryille, L L pp. 25 — 77. Comp. also Gebmar
in his Zeitachr.f, ErUom, L 1838, pp. 1—146, Tab. i.
PcecUocoria {PoicUochroma White previously) Dallas, Sketch of
the genua Foscilocoris, Trans, of ike Entomol, Sac. of London, *v.
1848, pp. 100—110, PL 13.
C^nus CanopiLs Fabb., with antennse four-articulate, is not to be
confounded with scutellera; the larvce only are known, apterous,
ocelli none.
Comp. J. W. Dalmav, JSphemerides entom. i. Holmie, 1824, 870.
pp. 34—36, and Lettre de M. Al. Db Lbfbbvbb d M, Audinbt Sebvillb
tivr le Comoptu obtectus de Fabb. ; Gu^ur, Magas, de Zocl. 1835. Ins,
PL 116.
Order XI. Ortkoptera.
Hexapod insects, with four wings, the upper coriaceous elytra,
the lower membranous and folded in their length radiately like a
fan. Mouth constructed for manducation, with strong mandibles ;
maxilldB furnished with galea cylindrical, vesicular (internal palp).
Metamorphosis incomplete.
Straighl-wmged, Olivier first separated theae insects under the
name of Orthoptera from the order of the Hemiptera of LiNK.£Us'y
and characterised this new order by the mode in which the under-
wings are folded and by the presence of a galea on the lower jaws.
In the oral parts they differ altogether from the heniiptera. But if
we stand, not upon the name of the order, but upon the distinction
* Eiusycl. mSth,^ Hist. 9tat. Tom. iv. Insect. Paris, 1789. IntroductUm, p. 16.
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INSECTA. 449
of it, then we cannot in any way look upon Olivier as its founder,
because Dk Gbeb had already collected into a distinct division, to
which he gave the name of Dermaptera, the self-same insects which
are now arranged in the order of the Orthoptera. Mem. pour sennr
d rHi8L de$ Ivs, 1773, p. 309. He characterised these insects
by their leathery shield-covers and the parts subservient to mas-
tication'.
Many orthopterous insects have two or three simple eyes. The
antennsB always consist of a great number of joints. The left upper-
jaw is on the whole more toothed than the right The under-lip is
divided into four lobes, of which the two exterior correspond with
the galeffi of the under-jaws.
The first thoracic segment is generally large. Ordinarily there are
four wings; sometimes, as in the mole-cricket and many species of
Fhatma, the anterior wings are very small, the posterior large; in
FhyUium, on the contrary, the hind-wings are entirely absent, and
there are only fore-wings or shield-covers; the instance of Perla-
marpha Cubtis is entirely peculiar, where shield-covers are wanting
and hind-wings alone are present. The joints of the tarsus are in
most fleshy or spongy beneath; the number of these joints is in
every species the same for all the feet, and varies from three to fiva
All the Orthoptera hitherto known are terrestrial, during the
different states of metamorphosis. Some are carnivorous or
omnivorous, but the greater number feed on plants. But some
species often appear in great numbers and are very ravening, and
may cause terrible devastation ; this is especially the case with the
locusts, a dreaded plague of the East.
The intestinal canal is on the whole short, and even in those
species in which it has the greatest relative length, it is to the
length of the body only as 1| or 2 : 1. The oesophagus has an
expansion which may be considered as a crop or fore-stomach ; this,
in the mole-cricket, is situated quite laterally. The muscular
stomach is small, orbicular, armed internally with homy teeth,
which are arranged in six rows. Hound the inferior orifice of this
stomach are blind appendages of variable number; the genus Acheta,
for instance, has only two such, ChryUus Fabr. {Acrydium Latr.)
six, Mcmtis and Blatta eight. Of the numerous vaaa urinaria in
this order we have spoken above (p. 256), as also of the presence of
^ Fabbioius named these inseets Ulonata,
VOL. I. 29
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450 CLASS YIII.
clustered salivary glands*. There are ten pairs of sHgmata, two on
the thorax and eight on the abdomen; the air-tubes are largely
developed In the species of the genus Acridium, which take such
long flights, there are vesicular expansions in the abdomen in which
the spiral thread is wanting. The nervous system consists of eight
to ten ganglia, and the nervous cord in the abdomen is often some-
what bent or tortuous.
Ck>mp. on this otder C. Stoll, Natwwrklijhe afbeddingen m Imckr^vmgm
der Spoken, wandelmde Bladen, ZabeUpringkanen em. Amsterdam 1787, en
yerv. 2 Deelen 4to.
AuDiNET Sebvillb, ffitL not. dee Ineectee, OrthopUree, Paris. 1839, ay.
PI. Sto. (a part of the Suites d Buffon, edited by Egret.)
For the anatomical peculiarities of this order, comp. L^N BuFOUB, Reek,
anatom. et phyeicl, ewr lee OrthopUree, lee ffymenopUree, et lee Newroptiree.
Mim, prieenUe, vii.
Besides the genenJ works of Buembistkb and Wistwood, oomp. also
Bbull^ in the work undertaken by him with AuDOunr, but not completed,
Jffiet, na/t, dee Ineectee, Paris, 1835. 8vo, Tom. ix. pp. i — 320, W. Dr
Haan, Bijdragen tot de kennie der Orthoptera in the Verhandelingen over
de Nalu/url. Oeechied, der Nederl. overzeeecke beeiuinffen, Leiden, 1839 —
1844, folio. Zoologie, Ineecta, bl. 45 — 148, and L. H. FiaoHBB, Orikoptera
ewropoea, AeeedtfMt tabida lapidi ineiem 18. Lipsie, 1854. 4to.
Section I. SaUaioria, Posterior feet saltatory, with large
thick thighs and tibiss, armed on the posterior margin with a
double row of spines. (The tibiae are received in a furrow on the
inferior surface of thighs, when folded forwards previous to leaping).
The insects of this division present^ more strongly than those of
the following, the typus of the order.
The production of a sound or song is peculiar to species only of
this division*. As in the CicadcB, so here it is only the males that
produce the sound.
Amongst the numerous writers on this subject^ it may suffioe to refer to
the work of GOBEAU, illustrated by many figures ; Eeeai eur la elfidulaUon
dee Ineectee, Annal, de la Soc, Enbomol. de France, vi. 1837, pp. 31 — 75.
^ L^N DuFOUB informs us, inoorrectly, that the salivary glands in the Orikoptera
were unnoticed until by himself, MSm. pr€e, vii. p. 297. But long previously, 6. B.
Tbevibanub had observed these parts in BlaUa {Biologie, rv. s. 3^3, 324), whose
opinion, that they occur in this genus of the order alone, was abundantly refuted by.
J. F. Meckel, who found them also in Mantie, Phaema, Acheta and Loeueta, though
less developed than in Blatta, System der verglekh, AnaJt. rv. s. 11 8, 119.
' Hence to the OrihopUra of this first division the name of Stridulontia might be
given, to the second that of if uto.
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INSECTA. 451
Family XL. Orylltdes nob. (Genus Orylhia L.) The charac-
ters of the section are also those of the single family.
A. Elytra and wings deflected.
Phalanz I. Acridii, AntennsB filiform or subclavate, some-
times depressed, acuminate towards the apex, mostly short or
moderate (not longer than half the body), with six to twenty-five
joints. Ocelli three. Tarsi with three joints.
Leaoh and Westwood (see Introduction to modem CUusif, of Im,
Westw. I. p. 438) name the insects of this group Locugtida, because
LiKN^UB had named the sub-division of his genus QryUuBf to which the
common grasshopper belongs, and which in great measure composes the
present division, LocvMa. It would certainly have been better if Geoffkot
had not given the name LocuMta to the sabred-graashopper with long
filiform antennsd (QryUug viridianmua L., &c.). But now that this generic
name has been adopted by Fabbioius, Latbeillb and all subsequent
writers, we think that a change of signification would rather increase than
diminish the confusion of which the above-named English authors complain.
The three simple eyes are placed in a triangle ; the two lateral,
between the compound eyes and the antennae, in the middle and
sometimes quite on that sur&ce of the head which is directed down-
wards.
Of the three joints of the tarsus the fii-st is long and appears to
consist of three joints that have coalesced ; so that to these insects
perhaps might very properly be ascribed five joints in the tarsus, as
by Latbeille in his Families naiur, du R^gne cmimal^ and by our-
selves in imitation of him in the first edition of this Handbook.
The sound produced by these insects is caused by a rapid friction
of the thighs of the hind-feet against the shield-covers; here the feet
act like the bow of a violin. The first abdominal segment is mostly
furnished with a dnun on each side, distinguished externally by
a circular or lunated membranous cover. Behind this membrane is
situated a small vesicle filled with fluid, and behind this again a lai^e
air-tube vesida Whilst some recognise in this an instrument for
soimd, like that of the CicaclcB, J. Muller and Y. Siebold consider
it to be an auditory apparatus; see above pp. 282, 283.
The borer for laying eggs doe? not project much, it consists
of four curved pieces. The eggs are mostly enveloped in a common
covering, adhering to each other by a tenacious frothy matter, and
in many species hidden beneath the eartL
29—2
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452 CLASS YXII.
L Posterior feet longer than body.
Tetrix Latb., Acrydium Fabr, Antennae short, with thirteen
to fifteen joints. Prothorax receiving a part of the head and
covering the oral organs. Pronotum produced into a scutellom
above the abdomen, sometimes beyond the abdomen. Plantola
between the claws of tarsi none.
Sp. Tetrix bipuneUUa, Acrydium bipunctalum Fabr., OryUuthiptmeUUut "L.,
Db Gbbb, m. PL 13, fig. 15 (magnified), DuM^iL, Cons, gin, «. /. Ins,
PL 25, fig. 5, &c, Zbttbbstbdt adopts different species here, which appear
to be only varieties. Gomp. on this species (whose scutellum is not longer
than the abdomen) and Tutrix siibtdata (whose scutellum extends with its
apex behind the abdomen) Philifpi Orihoptera Berolinensia (Berolini,
1830, 4to), pp. 41, 41.
Add genus Amcrphopua Sebv., imperfectly known, and HymoffioUg
Westw., ChoriphyUum Serville, with dorsum compressed into a
foliaceous crest erect above the body; habit of i/em6racufy SERViiiLE,
L L PL 8, fig. 5.
OryUua Fabr., Acridium Latk. Antennae of length of head
and thorax, with about twenty (twenty to twenly-four) joints.
Anterior extremity of prostemum not covering mouth. Plantula,
or pulvillus between the claws of tarsi.
Note. — LATBEUiLB Conjoined in sub-genus Podisma, species of which the
elytra and wings are short and unfit for flight. As Fabrioius had previously
written the genus Acrydium incorrectly, so also Latbbillb called it Acry-
dium, whom many imitated. It is, however, *kKpl^w ; and I would wil-
lingly hare rejected the diminutive form, and have given the name Acris
to the genus, since large, and even the largest species are contained in it.
It appeared, however, more advisable to give the name of Qryllus to Uus
the largest portion of the old Linnsean genus of the same name.
OryUua Fabb. AntenniB filiform or clavate. [Here belong
genera Ommexecha Sebv. (not Brull^, Gomphocerus Thunr,
(Edipoda Latr., Oxya Sebville, Monachidiwrn ejusd. and some
others; on which comp. Bubueister, Handh. n. 2, 1838, pp. 602,
<fea, and Sebville, HisL n. dea Orthopt.]
Sp. Qryllus migratorius L., B(ESBL, Ins. 11. Locust, xxiv. ; Blukbnbach,
Ahbild. naturk. Oegensi. No. 19 ; the thorax obtusely carinated, the jaws
bluish black« This insect is found in different regions of Europe and Asia,
and appears sometimes in great swarms, destroying everything on its road.
See on the devastations caused by this and other grasshoppers, EIibbt and
Spbkob, Introd, to Eniomol, i. pp. 215 — 226; Rittbb's Erdkunde also
contains many relations on that matter fit>m different parts of Asia. In
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INSECTA. 453
the year 1748 such a sort of all-destroying locusta spread itself o^er the
west of Europe, to the Netherlands, nay even to England. That they are
able to cross the sea, is placed beyond doubt by many accounts, according
to which people on ship-board, many miles from land, observed the swaims.
Some districts of the South of France are frequently damaged for several
years in succession by different species of grasshoppers, for the collection
of which large sums of money are sometimes bestowed. In 1844 at
SatTUet-Maries, in the neighbourhood of MctraeUlet, 15 18 oom-^acks were
filled with grasshoppers, and at Arlea 165 sacks; the expense amounted to
554a fr.; in 1833, in the first-mentioned place, 3808 kilogrammes of eggs
of these creatures were collected; the number of eggs that make up a
kilogramme may be computed at about 80,000. (See Ann, de la Soe.
BiUom, de France, il 1833, PP* 4^^ — 4^9 » these observations refer, how-
ever, Xo other species than OryUva migratoritu.) Different large species are
eaten by people of the East, as, for instance, OryUuB eriatatui L., Bocsbl,
1. 1. Tab. V. Already in Pliny examples are met with of locust-eating
nations {Hist. na>t. Lib. vi. c. 30 in fine, Lib. XI. o. 39 in fine), not to speak
of more modem accounts (Adanson, ffitt. de Senegal, pp. 88, 89, Salt,
Voya.ge to Ahytainia, London, 1814, p. 172).
GryUm camdescena L., Rssel, 1. L Tab. xi. fig. 4, Bum^bil, Cons, gin.
$, I. Ina. PI. 29, figs. 3, 4; thorax obtusely carinated, elytra brown-grey
with darker streaks, wings blue, at the point white, with a broad, black
margin.
GryUus UguUahu Chabpbnt., OryUue bigtmulue Panzbb (not L.), Pakz.
DeuUchl, In*. Heft 33, Tab. 6, Gssmab, Faun, Ina, Ewrop, Fmac. xx.
Tab. 22, 23, one of the smallest species of this genus, only $'" long ; the
male has knobbed antennae, &c.
Fhymatms Thunr, Serville. Anteniwe thick, longer than head
and thoiux, with joints distinct, the last longer, acuminate towards
the apex. Plantula large, orbicular between the daws of tarsL
(Ocelli little distinct or none.)
Sp. GryUm morbilloaui L., Bcbsel, Ins. n. Locust, Tab. 18, fig. 6, Stoll,
PI. n. b, figs. 3, 4, from the Gape of Good Hope.
Add sub-genera Pekuia Serv., Poscilocera ejusd., Rhovudea ejusd.,
and some others, here omitted.
Xiphicera Lam., Latr. (Xiphocera Burm.) Antennae depressed,
lanceolate or ensiform. Frons produced into a cone. Other cha-
racters as in GryUL
Sp. Xiphic, serrata, Gryllus serratus L., De Gesb, M6m. iii. PI. 42, fig. 2,
PI. 41, fig. 6 ; RcBSEL, II. Loc. Tab. 16, fig. 2, Stoll, PI. 19 b, fig. 71, PI.
a I, -fig. 81 ; — Xiphic. emarginata, Sebv., R(ESel, 1. 1. fig. 3, Guv. i2. Ani.
id. ill. PI. 85, fig. 2,
Add genus Trigonopteryx Charpent.
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454 CLASS YIIT.
TruaaUs Fabb. {OryUua Acrida L.). Antennie depreesed, ensi-
fonzL Head conical, longer than thorax. Bodj elongate. Elytra
narrow. Posterior feet very long, with tibia elongate, slender.
Sp. Truxalts nanUui Fabb., Orylhu namJhu L., Boesbl, Ins, n. Xoetuf.
ind, Tftb. v. (antenne badly drawn) ; \>\sit±BiL, Com. ffin. $. L Int. PL 14,
fig. 3; CUVDEB, JZ. Ani, id. Ulttttr. Ins, PL 84, fig. 8, in south of Europe
and in Africa ; — Trux, variabilis, Kluo, and others, very similar Bpecies ;
both the above-named species are also found at the Cape of Good Hope,
the first also at Japan ; see Db Haav, op, eit, pp. 141, 144. Comp. on
this genus Thvsbvrq, Nov. Act, Soc, UpsaUens, ix. 1837, pp. 76—88.
Proacopia Klug. Body elongate, apterous. Head elongate,
pyramidal, produced beyond the eyes ; antennse subulate, very short,
with six or seven joints. Prothorax very long.
Sp. Proscopia radula Klvo, Cuv. R, Ani, id, tU, Ins, PI. 85, fig. i, firom
Brazil, like most of the species (all from America). These animals
resemble Phasma in the habitus, and Mantis in the long prothorojR. I can-
not perceive a groove on the thighs for the reception of the tibiae^.
II. Posterior feet shorter than body.
Pneumora Thunb. Antennae filiform, with twenty-one to
twenty-five joints. Head short, with eyes remote. Prothorax pro-
duced into a membranous scutellum; thorax short. Abdomen In
males inflated or vesicular. Ghreatest part of elytra incumbent
horizontally on the back. Elytra and wings short in females ; in
some almost quite obsolete, covered by scutellum*
Sp. Pneumora variolosa, Latb., Oryllus variolosus, L., Fabb., Pneumora
marmorata, Thunb., Stoll, PI. w, fig. 78 ; Cuyibb, It. Ani, id, ill.. Ins,
PI. 84, fig. I ; from the Gape of Good Hope. The other species also of
this genus belong to the south of Africa.
Phalanx 11. Locustartce. Antennae setaceous, long (often
longer than body), with joints numerous. Ocelli in most none.
Tarsi with four joints. Females famished with ovipositor bivalve,
ensiform. In males the right, or more rarely both elytra, mostly
furnished with a basal portion, orbicular, transparent (musical
organ).
^ Chabpentieb records a winged species from Chili. Elytra were not present;
wings very small ; but what seems most to distinguish this species is the absence of a
ptdvUlus or plantuUt on the tarsus. Hence it might form a distinct genus Astroma
(Chabp.) Gebmab's Zeitschr. m. 184 1, s. 305.
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INSECTA. 455
Sabre-gnudkopperSyWiQi Linkjeds €hyUi Teaigomoy Sj^ Nat edL
xn. 1, p. 695; the name of TetHgomm^ however, is rightly given
bj later writers to the CuxuUb.
In many species the tibiae of the fore-feet have two oval apertores
covered by a tense membrane (see above, p. 2S3) ; comp. also
Laxsdown Guildikg, TranstMcL of tht Lirmaon Society^ xv. 1827,
pp. 153 — 155.
The wings are in some very short or entirely wanting {Hetrcdes^
some species of Bradyporua^ Sagct, Sic)
Locmta Geoffroy, Fabr., Latr.
Sub-genera: Brctdyportis Gharpent., ffe^rodes Fisch., Ephippi-
gera JjATB., Barhitistes Gharpent., Meconema Serv., Aeridopeza
GuER., BuRM., Fomatonota Burm., Jfecopoda Serv., Scapkura Ejbbt
(and Gymnocera Brulle), Phaneroptera Latr., Serv., PhyUoptera
BuRiL (Phylloptera, Ancylechay Steirodon Serv.), PhyUaphora
Thunr, Hyperomala Serv., Aspidonotus Brulle, Pterochroza Serv.,
CyrtophyUus Bueil, Pseudophyllus Serv., Aprion Serv., ThlibasceliB
Serv. {Platyphylltis Burm.), Meronddiua Serv., Acanlhodis Serv.,
Burm. (and PlatyphyUus Serv.), Copiophora Serv., Pseudorynchua
Serv., Conocephalus Thunb., Agrcecia Serv., Xiphidium Burm.
(Xiphidium Serv. and Orchelimum ejusd.), BticrcUes Burm., DecUcus
Serv., Bttrm., Locuata Serv. {Phasgonnra Westw.), Listroacdia
Serv., Sctga Gharpent., Schizodactylua Brulle {Acheta Burm.),
SteTiopelmattia Burm., Raphidophora Serv. {Phalangopaia Burm. in
part).
Sp. Loctuta jmpa^ Qrylhu puptu L. {ffderodea pupa, FisoH.), Stoll,
loeutt, Fl. xn. a, fig. 45, 46 ; Rcbsbl, Ins. u. Locust, Tab. vi. fig. 3 ; Da
GsEB, MSm. m. Fl. 39, fig. 5 ; winglees, thorax and abdomen armed with
spines. From the Gape of Good Hope.
Locutta viridisnmaYABU., Qryllutviridiaaimut'L., BacsiL, /f».n. Locutt,
Tab. X. XL ; Fanzeb, Veuttchl, Ins. Heft 89, Tab. 18, 19, DuMia. Cons. g6n,
s. I. Ins. Fl. 24, fig. I ; green, without spots, with green shield-covers, and
very long antenns. This species is not rare with us on pastures, in gardens,
ftc. The eggs laid in autumn, pass the winter, and the grasshoppers come
forth from them in May or later. After the third moult they shew rudiments
of wings and shield-covers (in July) ; towards the end of August they moult
for the last time, and become perfect insects. The copulation is repeated a few
times, and the eggs are placed in the earth by the female by means of her
sabre-shaped ovipositor, not all at once, but at intervals in different places.
These grasshoppers die towards the end of autumn. — Locusta varucivora
Fabr., QryUus vermeivorvs L., B(X8BL, Ins. n. Locust. Tab. vm. rx. Panzkb,
1. 1. Tab. 90, II ; this is somewhat smaller, has shorter antenme, and brown
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456 CLASS viir.
spots on the green elytra. The Swedish peasuits put this insect, aoooiding
to LiKNiBUB) upon their hands to bite off the warts. — LocuMtavaria¥ABR.f
Meconema varia Sebyillb, Panzbb, Deutschl. Int. Heft 33, Tab. i, &a
B. Elytra and wings horizontal, for the most part incumbent
on back.
Phalanx III. Achetidce {Qryllidea Latr.)- Labium with four
laciniae. Tarsi triarticulate, with joints slender, sometimes spinose.
The Crickets, gra/o&nde sprinkhanen (Grctbheuschreken) live under-
ground or in holes, and are mostly noctutnaL They form with
LiNK^sus the division Acheta of the genus GrylluSy or the genus
Gryllua of Geoffrot and Olivier and the genus Acheta of
Fabbicius.
Myrmecophtla Latr., Sphcertum Charpent., Burm. Antennae
setaceous, long. Head covered by prothorax. Ocelli none. Elytra
and wings none. Females with ovipositor exsert, bifid. Anterior
feet gressorial, posterior with thighs thick, very broad.
Sp. MyrmecophUa aeervorum, Blatta acervorum, Paitzbb, IkvttchX, Ins. Heft
68, Tab. 24; Gu&iK, Iconogr. Im. PL 54, fig. 6; Cuviek, A Ani, id. UL,
Ina. PL 8«, fig. a.
Acheta Fabr. (in part), OryUua Latr. Anterior feet gressorial.
Antennae setaceous, long. Ocelli mostly obsolete or none. Elytra
short, and in most two wings longer. Females provided with ovi-
positor exsert.
a) ProQotum transverse or quadrate.
Sub-genus: Acheta nob. {PlatyUemus Serv., Brachytrupea Serv.,
OryUua Serv., I^emobitis Serv., Trigonidium Serv.^ Podoacirtus
Serville, Sderopterus Hagenb., Eneoptera Buru.)
Sp. Acheta damesHca Fabr., OryUua domuticua L., RoiaEL, Ina. n.
Loeuat. Tab. iii. ; Panzeb, DeutacM. Ina. Heft 88, Tab. 6, 7 ; Cuv. R. Ant.
id. ill., Ina. PL 81, fig. 4 ; the hea/rth-crichet ; body yeUow-brown, wings
longer than shield-covers, terminate in a tail-like point. These ftwimala
live near the hearth and under stoves in an artificiaUy warm climate, and
chirp of an evening or by night, and especially when it is about to rain;
the sound caused by the friction of the shield-covers is in a very high note,
which some persons, otherwise not deaf, cannot perceive.
Acheta campeatria Fabb., QryUua eampeatria L., B<ESEL, I. 1. Tab. xin.
Panz. DeuUchl. Ina. Heft 88, Tab. 8, 9, Cuv. L 1. fig. 3 ; the Jidd-cricka,
This species is larger than the preceding, has longer shield-covers and a
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IKSECTA. 457
black body, without the elongated point at tlie wings, wbich in the
preceding species extends behind the elytra.
iVbte. — Eneoptera Bubm. (PlatydaOyluM Bbvll^ and Sibyillb in part).
The generic name devised by BuKMiiSTBK on account of the elytra similar
in both sexes of some species, haying no musical organ in males ; comp.
BuBMEiBTiB, H<mdb. derEnUmol. n. a, p. 10x5, and on the typical species,
De Haan, 1. L p. 431. The name of PUdfdaetyhu ought to be rejected,
since given long previously to a genus of Sauiians. In these AchtUx the body
is elongate , posterior feet very long ; ovipositor of females long, recurved
upwards, with valves divergent, clavate at apex. Sp. Ackda bnuUieiuia
Fabb., QrjfiUu twrinamtimi Di Gub, Mim, m. PI. 43, fig. i, GuiRiv,
leonogr,, Tn$, PI. 54, fig. i, (named ChyUua ServiUd).
h) Pronotum oblong, narrower forwards.
FhaUmgopais Sebtille. Feet elongate. Elytra short; wings
none, or yerj short rudiments of wings.
Sp. PhcUangopaii lonffipes Sbbvillb, ffi$i, not. da Orthopt., PL ii, fig. i.
Habit, in South America.
(Ecanthua Sebville. Feet elongate. Elytra and wings long.
Sp. Aekeia Ualica Fabb., (Eeantkut peUueem Sbbvillb, Pavzib, Z^vtscM.
Ifu. Heft M, Tab. 17.
Note. — Bbull^ and Sbbvillb assign four joints of tarsi in posterior
feet to these insects, which Bubxbisteb has shewn to be an error, ffcmdb,
der EiUomol. n. 2, p. 731.
Gryllotalpa Latr. (species of Acheta Fabr.). Anterior feet
fossorial, compressed, broad. AntennsB setaceous, shorter than
body. Ocelli two, placed between the compound eyes. Pronotum
elongate, oval, gibbous ; ovipositor of female none.
Sp. CfrylloUdpa mdgarU, GryUu$ OryUotalpa L., B<E8BL, Iru, n. Loeud. Tab.
XIV. zv. ; DdmIbil, Com. gin. «. L Ins. PL 2$, fig. 'j ; the mole-cridlcH,
li" long, brownish-black, yellow below, with grey-brown, darker- veined
shield-covers. The fore-feet work like saws, or as spades, with which these
insects burrow the ground. The female lays her eggs in a hole under
ground, to which a long curved passage leads, in and out of which she
often creeps ; at first the young ones live dose together, but soon disperse.
Horse-dung appears to attract these animals, swine-dung to repel them.
They live especially in garden-mould and turfy bog. On the anatomy see
KiDD, Philoa. Tram. 1835, Part n. pp. 203 — 246, and my AntedBtninffen
aver het inwendig maakeel dee veenmols, Bijdragm M de natuurh. Wetenech.
v. 1830, bl. 94 — loa. This species is spread throughout the whole of
Europe and of Asia, and other species of this genus are found in all parts
of the world.
Cylindrodea Gbat, Sebville. Body elongate. Yestiges alone of
wings and of elytra. Feet short, with tarsi didactylous.
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458 CLASS VIII.
Xya Illig. (Trtdadylua Oltv., Latr.) Anterior feet fossorial.
AntennsB short, filiform, with ten joints. Ocelli three. Posterior
feet elongate, saltatory, terminated hy lamellae in place of tarsi.
Xya Illig. Posterior tibisB Aimished with lamelkd elongate,
lanceolate, depressed.
Sp. Xyafotaor Bubm., TVidadyltu paradoaeut Latb. ; DuM^R. Com. gin, «. I,
Jtu, PL 2$, fig. 8 ; Oni^nr, leonogr, Int, PL 54, fig. 5. Hab. in Africa.
Xya variegata Illio., Bubic.^ Cdvibb, R, Ani, id. ill., Ins. PL 81, fig. a.
Hab. in south of France and in Spain. L^N DuFOUB is of opinion that
it belongs to the AcridU; oomp. his Reekerckes sur VHuL naL du Tridae-
tylepanachS, Ann. dei Sc. not. 2e S^rie, Tom. n. 1838, pp. 321 — 334.
Sub-genns : Khipipteryx Newman, BauLLi (species of Xya Buril,
Ehipidopteryx). Posterior tibiae destitute of lanceolate lamellse.
Section II. Curaoria. Feet ambulatory alone. Wings and
elytra horizontal, incumbent. (Musical organ present in none.)
Family XLI. Mantides nob. (genus MarUia L.) Tarsi with
five joints. Head not covered by thorax.
Phalanx I. Spectra [Phasmida JjE^kCll. Phasmodea BuRM.).
Fore-feet similar to the rest, not raptorial, very near the head,
remote firom the intermediate pair. Antennae mostly setaceous,
long, in a few short, filiform. Labrum deeply bifid. Mandibles
thick, strong; maxillsB hard at the apex, subdenticulate ; galea
plane, broad. Labium quadrifid, with laciniss unequal, the external
very long, plane, hatchet-shaped. Elytra almost always short,
wings large; sometimes wings and elytra none.
Pkasma LiCHTENST., Fabr. {Spectrum Stoll).
These insects are in great part confined to tropical countries;
many also are found in New Holland. They ]ive on vegetable food.
Some wingless species have the form of dried twigs, and others,
which are flat, with membranous and vein shield-covers, have a great
resemblance to leaves (the genus FhyUiujn.) Of some species the
males are much smaller than the females; the former have often
three simple eyes, which in the last are absent. In this division
the largest species of insects are foimd.
Comp. A. A. H. LiCHTKNBTEiN, Dissert, on ttoo ncU. genera hitherto con-
founded under the name of Mantis; Trans, of the Linnoean Soe. vi. 1801 ;
TouBSAiiTT VON Charpentisr's JBemerhungen zu Lichtbnstsin*8 Ahh.
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INSEGTA. 459
£5. die MatUU-Arten, in Gbrmab'b ZeUmHnt, /. d, JAilom. v. 1844, t.
1*11 — 311.
6. R. Gbat, Sfffwpsit of the tpecies of Insects heUmging to the family of
PhasmicUB, London, 1835, 8yo ; and his ErUom/ology of Audralia, Part I.
Monogr, of the gen, Phatma, London, 1833, 4to» with 8 ooL pL
Phasma Fabr. (Species of genus Phasma LiCHTENST.) Body
elongate. Mesothorax often very long.
a) Feet cylindrical or angulate, prismatic
* Both sexes apterous,
BacUlua Latr. Antennie short (of length of head), filiform or
conical.
Sp. Phasma Bossia Fabb., &ujppl, Entom. syst. (1798) p. 187; P. Rossif
Fauna Etrusca, Libnrni, 1790, Tab. vm. fig. 1 ; Cuv. R, Ani, €d. HL, Ins.
PI 79, fig. 1 ; Bkull4 Hist, not, d^Ins, ix. PI. 9, fig. 1 ;--Phasma tHpoU"
tantm, Be Haan, 1. 1. Tab.«xy. fig. 3.
Bacteria Latr. Antenna setaceous, longer than head and pro-
thorax.
Sp. Phasma fenda Fabb., Bacteria artmatia Gbat, R(BSBl, Ins. n.
Locust. Ind. Tab. xiz. fig. 10; Stoll, Speetr. PI. 13, fig. 51 ; Bacteria
sarmentosa Wbstw., Cabinet of Oriental BiUom, 1847, PL 33, fig. i, &c.
Note. — For Bereral other sub-genera see Bubhxistkb, L 1. pp. 568—610,
and Sebyhj.!, L L
* * Male winged, female apter<ms. (AntennsB setaoeons, long.)
Cladoa»ru8 Sebyille. {Cladomorphus Gray fern.)
Add sub-genus Monandroptera SebtHiLE, 1. 1. p. 244.
* * * Both sexes winged. (AntennsB setaceous, mostly long, in females of
some species shorter).
Phasma nob.
Sp. Phasma gigas F., Mantis gigas L., Stoll, Spedr. Tab. n. fig. 5 ; Cuv.
R. Ani. id. iU., Ins. PI. 80, fig. i ; De Haan, 1. 1. PI. 14, fig. 3. From
the Molucca Islands and Java.
Note. — Here belong Teiy many genera of modem authors, Diapherodes
Gbat, Eaplopus Gbat, Bubm., Oyphocrania Seby., Bubm., Necrosda
Sebv.
Aschiphasma Wbstw. {Perlamorpha Cubtib, Sebv.) is distinguished by
elytra none or very small, spiniform, with wings ample and anterior
margin obscure. De Haan records many species from the islands of
Java and Borneo, op. cit. pp. 113 — 1x6.
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460 CLASS VIII.
h) Feet winged or foliaoeous.
Sub-genera: Tropidoderus Gray, Ectatosoma Grat, Pritopui
Sebv., Gray.
Sp. Priaoptu Jlabelliformi$ Gray, Phanaa dracwnerdut Liohtbnst., Siou^
Spectr, PL i8, fig. 65, from Surinam. Priaop. HontokU Db Haan, 1. L
Tab. 12, fig. I ; hab. Gape of Good Hope. In the shorter body and entire
habit they somewhat approach the genus Phyllvum,
Phyllium Illig. (species of genus Mant%8 L. and Fabr., species
of Phasma LiCHTENST.) Abdomen oval, depressed. Prothorax
scarcely longer than mesothorax, sub-triangular, posteriorly nar-
rower. Antennae of males setaceous, pilose ; of females very short,
filiform. Elytra short and wings long in males ; in females elytra
large, membranous covering abdomen, wings very small. Thighs
dilated, alate.
Sp. PhyUium tiecifaLium, Mantis tiecifolia L., Fabb., R<bs., Ins. n. Loe,
ind. Tab. zvii. f. 4, 5 ; Stoll, Spectr, PI. Yir. f. 24, ^6 ; Dnio^. Oon$.
gin. s. L I'M, PI. 93, fig. 1 fem. &c. ; the wdking-Uaf, This species is
extended from the Sechelles over Java and Timor to New-Guinea. Some
allied species are from the same regions and from China; none are known
from the new world.
Phalanx II. MarUides. Anterior feet raptorial, with coxss long
and femora compressed, receiving the tibia beneath. Prothorax not
shorter than mesothorax, mostly exceeding the mesothorax in
length. Ocelli three. Antennae mostly setaceous, moderate. La-
bium entire, orbicular. Labium quadrifid, with lacinise equal.
Elytra and wings in all.
The anterior feet are situated close to the head, and are much
stronger than the rest, usually Yery thin; they haYe large com-
pressed thighs which are armed beneath with teeth or spines, and
haYe a channel in which the tibisa can be lodged like a clasp-knifa
With these fore-feet bent together and with head on high, these
insects often sit long in an immoYeable position. Henoe the eastern
people say that they are at prayers, and moreoYer, like true Ma-
hometans, with the face and hands tinned towards Mecca. With
this pious deportment howcYer, and the superstitious reYerence
which is in consequence conferred on these insects by different
nations, their cruel worldliness is at Yariance ; they Hyc on other
insects, and also mutually doYour one another. See Bluhenbach,
AbbilcL naiur-hiatoriech^ GegenHdndey No. 88.
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INSEGTA. 461
Stoll gives to these insects the name of waUanff-Uave$, by which how-
ever is usuaUy UDderatood the preceding genus PhyUium.
Mantis L. (in part), Fabr. (exclus. Mant. siccijblia).
a) Anitennm of males peeUnaie, of femalet setaceous. Head produced
fupwofrds inUo a horn,
Empaaa Illig.
Sp. ManHs gon^ylodes L. and Fabb. (fem. Mantis Jlabdlicomis Fabr. male) ;
BoiBSL, Ins, n. Loctui. Tab. vii. ; Stoll, Spee^. PI 16, figs. 58, 59. PI.
I7> fig* ^i ; Bengal, Ceylon; — MaiUis pauperata Fabk., Cuv. It, Ani. id,
ill., Ins. PI. 78, fig. 3; south of Europe, north of Africa, &c.
Sub-genus Blepharis Skbv.
b) AntemuB in both sexes simple ; head very often transverse, broad.
Mcmiia Illio.
8p. Mantis rdigiosa L., Boisbl, Ins, n. Locust, Tab. i. n. Tom. rv. Tkb.
xn.; Pakzeb, Deutsehl, Ins, Heft 50, Tab. 8; Cuv. R. Ani, 6d. iU., Ins,
PL 78, fig. I ; in south of Germany, in France, and other parts of the
south of Europe, and also in Africa; — Mantis oratoria L., Mantis beUa
Salzmak, GsBMABy Faun, Ins. Europ, Fasc. vi. Tab. 16, South of
Europe, &c.
Note. — Several sub-genera, founded on the form of the prothorax, the
eyes, the feet, and other characters, are recorded by Sebvillb and BuB-
MBIBTBB, not always well defined: BremophilaBv^u. {EremiapkUaliKFEB.),
MetalleiaicaWisafrw., Chateessa Bubh., Tarachodes Bubm., Tkeoclytes Sebv.
(Votes Bubm.), ffarpax Sbbv., Acanlhops Sebv., Schizocephala Sbbv., and
others; for which consult the authors cited above.
Mantis strumaria L. (Boissl, Ins. u. Locust. Tab. in.), and similar
species with prothorax dilated, partly covering head, form the transition to
the BlattcB,
Family XLII. BlattaruB, Tarsi with five joints. Head in-
ferior, covered by prothorax clypeiform.
BlaUa L. Body oval or orbicular, depressed. Antennas long,
setaceous. Ocelli obsolete. Maxillary palps elongate, with last
joint hatchet-shaped. Feet all similar, long, tibiae provided with
spinules moveable. Abdomen posteriorly supplied with two ap-
pendages conical, articulate.
a) Each sex apterous.
Sub-genuB : Polyzosteria Buem. (Species of BlaMa Serv.)
b) MoUe winged, female apterous.
Sub-genera: FerisphcBria Sbbv., Bubil, Heterogamia Buril
(Species of BlaUa Sebville.)
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CLASS VIII.
c) Each mx foinffed.
Corydia Serv., Btjrm. Elytra horny, with veinB indistinct (add
Phoraspia Seby., Bubm.)
Sp. Blaltta Petwenooka Fabb., Camda Pdiveriana h., Stoll, BlaU. Tab.
▼. d. figs. 11, 11, Blatta KeterodUa Pallas, Spic, ZooL ix. Tab. i, fig. 5,
ftc.
Sub-genus: Blatta nob. Elytra coriaceous^ with veina eminent;
posterior wings shorter, not reflected at apex.
Note, — ^Here belong several sub-genera of Burmeisteb and
Sebville; Thyrsocera Bubu. (Fseudoniops Sebv.), Ischnoptera
Bubm., Nyctibora Bubm. (species of Blatta Serv.), Feriplaneta
{Kakerlac Latb., Sebv.), EpUampra, PanMora^ Nauphoeta, Pros-
cratea, Zetobora Bubm., fformetica Bubm. {Brachycola Sebv.),
PaTiesthia Sebv., Bubm., Blabera Sebv., Bubm., (and Monachoda
Bubm.)
' Sp. Blatta orienialis L., PeriplanHa orientals Bubm., Db Gbkb, /m. m.
Tab. IS, fig. I ; Panzxb, J>eut9ckl, Ira. Heft 96, Tab. is ; 'ffemeene haker'
Idk, ffemdne Pfslerschabe, Blatte dea euitines ; Blatta lapcmea lu, Pakzkb,
ib. Tab. 13, &c.
The Kaherlact are noctumaL T^y are great deyonrera, and gnaw
even leather, shoes, &c. The name Blatta (from pXdwrcaf, to injure) oocurs
already amongst the ancients ; yet probably it signified other insects. The
female lays her eggs heaped into separate little masses distributed in
cells; see 6<bzb, Naturforscher, xvn. s. 183—189, Tab. rv. figs. 16 — 19.
Bathxe has communicated his observations on the development in .the egg
in BUata gerfnanica in MtcheCt Arehiv /. Anat, t*. PhyvM. VL 1833. s. 371
—378. Tab. rv.
Anaplecta Bubm. Elytra coriaceous, with veins eminent. Pos-
terior wings elongate, reflected at the apex.
Species small, American.
Family XLIII. Forficularice. Tarsi with three joints. Elytra
Bubcrustaceous, not reticulate, abbreviate, truncated, incumbent on
back, placed in contact by a straight suture. Posterior wings mem-
branous, partly folded in longitudinal rays, partly transversely,
covered by elytra.
The insects of this &niily make the transition from the Orthoptera
to the Coleoptera, LiNiTiBUS counted them with these last, as did
also Stbaus amongst more modem writers. Leach and Kibbt
form a distinct order of them, under the name of Dermaptera
{Transact, of the Lirm, Soc, Vol. XL 1815, p. 87); ihb name,
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INSECTA. 463
wMch BuBMEiSTER changed into DtmaUipi&ra^ had been previonsly
(see above, p. 449) given by De Geer to the order of the Orthopterck.
Idos DuFOTTB, who subseqaently proposed the same separation,
named these insects, in imitation of DuHiBiL, Lahidura; Webf-
WOOD EupUxoptera {Zool, Journal, 1831, Modem ClassificaUon, L
1839, p. 308). At all events these insects have greater agreement
with the Orthoptera than with the Coleopiera ; they differ from the
last by their incomplete metamorphosis and by many particularities
of internal structure. The great size of the nnder-wings in com-
parison with the elytra is very common in the Orthoptera (to refer
to PhasTna alone), and the reflexion of the point of the wing also is
not wanting in some other Orthopteray as, ex. gr, Anaplectay the last
of the sub-genera of BlaUa recorded abova
Forficula L. Body depressed, narrow, elongate, with prothorax
quadrate. Antennae filiform, a little longer than half the body, with
joints very distinct, mostly more than twelve (ten to forty). Ocelli
none. Labrnm entire, orbicular; mandibles denticulate; labium
bifid, with palps short. Abdomen with last segment forcipate, with
forceps homy, moveable, in males arcuate, or more open.
Comp. on the anatomy, Pobsbia, Dmert. tnaug. aittem taiiamina circa
Anatomiam ForJietUat aurictdarke L. iconib, iUuttr, Jens, i8oo. An
extract from it may be fomid in Wisdbmank, Archivf. ZooL u. ZooL i8oi,
L I, B. 130 — 334, with the corresponding figures, also Ii. a. PL in. with
the explanation by the author himself, s. 330—335.
L^N DuFOUB, Recherches Anat. wur lea Lahidourei, Atm. de$ Sc. noL
XIII. i8a8. pp. 337—3^^-
Sp. F<nfcula aurieularia L., Pakzbb, DeuUchl, Ina, Heft 87, Tab. 8, nude;
Gut. R, Ani, €d, HI,, Ina, PL 77, fig. i \—Fwiicula minor L. ; Panzbb,
L L Tab. 9, &C.
Barwiga, peree-oreUUa, Ohr-wG/rmer, It is said that they creep into the
ears, and hence this name ; this they do not more than other insects, which
by chance may get into the ears of persons asleep. They live in moist
places, and eat almost everything ; they are especially fond of fruits and
honey. The females sit on their eggs, and guard them; the young also
creep under their mother, like chickens under the hen, and she often rits
quietly for hours together over them. Da Gbbb, Mim, a, I. Ina. m.
p. 548.
Note. — Some are without hind-wings but provided with elytra^ or are
entirely apterous. According to difference of form and number of joints
in antennae, Lbach, Latbeillb, and especially Sebvillb, have formed
several sub-genera, of which it may here suffice to cite the names ; Apachya,
Meeamera, SparaUa, Diplah^, Forficula, Echinoaoma, Lobophora, Pyroffra,
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464 CLASS VIH.
Psalidophora Sebv., Pofficmia Latb., Ssby., Pyffidierana Siby. Comp.
Sbbvillb, ffitt, not. dea Orih. pp. 18 — 55.
LATBimiiLB conjoined apterous species imder the name of C%«^Mitt9ia, spec,
of genus Forficula Sxbville.
Appendix to the Order Orthoptera.
Thrtps L. Body linear, depressed. Head produced inferiorly
like a rostrum ; labrum elongato-conical ; mandibles setiform ; max-
illae triangular, acuminate, with short palp, of two joints or three,
without galea; labial palps very short, with two or three joints.
Antennae of the length of head and prothorax, filiform, mostly with
eight joints (five to nine). Wings mostly four, membranous, narrow,
incumbent, -with margin ciliated. Feet short, with tarsi biarticulate,
the second joint without claw, terminated by a vesicular plantula.
IJ^ote. — Small infiocts living in plants, especially in flowers, which
were first detected and described by De Geeb in the middle of the
last century, under the name of Phyaapua, He then discovered
palps ; and since the Hemiptera are destitute of these, they have been
referred improperly to that order bj several writers. Some think
that they compose an order of their own {Thyaanoptera Haltday,
Westwood). They differ from Orthoptera by the absence of the
galec^ The fiEunily is of uncertain position, intermediate amongst
the Neu/roptera, Orthoptera and Hemiptera. Metamorphosis incom-
plete.
In some the females are provided with a quadrivalve sting or borer,
(TerdiTainiia Hal.), in others there is not such a borer, but the abdomen
terminates in a tubule {TubuUfeni), Some species in both sections are
apterous.
Comp. Di Gkbb, Miim,. in. pp. i— 18. PL i ; Bubmsistbb, Mandb, der
BfUom, n. a, pp. 404 — 418; Westwood, Modem CUumfcaUon, n. pp. i —
5, Oeneric Sytwpnt, pp. 45, 46 ; liiov Dufoub, DeacripHon et figure cTune
nouveUe eepiee de Thript (Thripa aptera LioN J)UF., and PkUeotkripg
pedicularie H^Lro^T ?), Ann. dee Sc. not,, sec. S^rie, Tom. zi. 1839, PP-
341—324, PL 8, figs. 8—18; E. HsKOXB, BeUr&ge gur Naturyet^iehte der
Phyeapoden, Mit Kupfert. Aue dem Jahrgange 1852 der SUasunffeberickte
der Kaiaerl. Akad. der Wieeenech, zu Wien, Beeondere abgedruckt. 8vo.
Sub-genera Phlaxihripe, ffelicthripe, Sericothripe, Tkripe, Afdanthripe,
^dothrips Haudat, Entom. Magazine, Tom. ni. and iv.
Order XII. Coleoptera.
Hexapod insects, with four wings, the anterior hard, coriaceous,
covering the posterior membranous, folded back transversely before
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INSECTA. 465
the apex. Mouth formed for manducation, famished with mandi-
bles, maxillaB and palps both maxillary and labial. Metamorphosis
complete.
Shield-winged (Ooleoptera from KoXetk, shecUh, covervngy shieldy and
iTTepovy fving)y with Fabricius Eleutherata. We have on this
numerous order of insects amongst others the following works : —
J. R YoET, Gatalogua st/stemcUicus Goleopterorwnhy Hag. Com.
1769—1806, 4to, II parts. With (Ist part 55, 2d part 50) beautifully
coloured plates ; those of the first part mostly engraved by Kleekanh
and some by Koesel, and a very useless Latin, French, and Dutch
text. G. W. F. Panzer has supplied a german edition of it with
the addition of the synonymy. Erlangen, 1793—1798.
A. G. OuviEB, SrUomologie, ou Rist Nalwr, des Inaectes {Coleop-
t^es.) Paris, 1789—1808, vm Tomes, 4to, av. (363) pL enluminfes.
(The plates make up the last two parts). Stubm has given a less
costly edition of the plates, but not complete. Niimberg. 1802,
1803 (96 plates). Illioeb undertook a German edition of the
text
J. 0, FAfiBion Syatema deutkera/torvm. Kiliie, 1801, 8vo,
u TomL Index Entomologies in G. W. F. Panzer, Fcmn. Ins,
OerfMm, Part i. EleiUhercUa, Norimborgi^B, 1813.
O. J. ScHOENHERR, Synonymia Inaectontm, oder Versuch einer
Synonymie alter bisher bekannten Inaecten. Stockholm u. TJpsala^
1806—1817. 3 TUnUt Appendix, 8vo. nk iUtm. KupferU Ejusd.
Synon. InsectoruTrh. Tom. iv, also under the title of Genera et
Species Cwrcfidionidvm, Parisiis, 1833 — 1845, vin TomL
DijEAN, Species ghihral des Coleopt^es de la Collection de M,le
Comte Dejean. Paris, 1825—1839, 8vo. 6 Tom and 7 VoL
For the anatomy the above-cited Monograph on the Cockchafer
by Straus Durckheim may be consulted (see above, p. 266), and
also the numerous investigations of Leon Dufour, which extend
over the whole of this order. Ann, des Sc. naiur. Tom. n, nL 1824,
IV, V, VL 1825, vm. 1826.
This order is distinguished from the preceding by the complete
metamorphosis, as also by the absence of a galea, howsoever in some
two pairs of maxillary palps are present, and morphologically the
innermost palps do not differ from the galea of the Orthoptera,
VOL. I. 30
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466 CLASS VIII.
Simple eyes are not present in the generality of these insects; they
occur however in some; two are met with in many small Brachdytra,
species of ArUhophagus and OmcUiimi (see Klug, cited above,
p. 280), and in Pauasus hucephdlua; and according to Curtis one is
found in the middle of the head in soAe Dermeatidw (Westwood,
IrUrod to Mod, Classif, Vol. i. p. 35). The antennse have a very
different form and length, but consist commonly of eleven joints.
The anterior thoracic segment is large and moveable; the middle and
last segments of the thorax are entirely covered above by the shield-
covers. The imder wings are absent in some species, which natu-
rally also in that case are not able to fly, since the shield-covers are
not used for flight ; these last are never wanting, but are sometimes
very short. The integuments of the body are commonly hard, but
less so on the dorsal sur&ce, inasmuch as it is covered by the elytra.
These insects undergo a perfect metamorphosis. The larva resem-
bles a worm, and commonly has six homy feet*. Some larvse (Cwr-
culU>y CeramJbyx) have no feet, but membranous tubercles in their
stead.
Of the difl^on of this order an idea may be formed when we
reflect that, according to Latreille, more than fifteen years ago
about 30,000 species of beetles were known', and, according to
Westwood, in 1839, 28,000 were found in the British Museum
alone. And how many species have there not been since then, and
are there not almost daily discovered 1
The number of joints in the tarsus was first made use of bj
Geoffbot for the division of this order {Hist abregie des Ifiseetes, i.
pp. 50, 51), who was followed by Olivier {Ene. method,y Hist n. d.
Ins, Tom. v. 1791, p. 107), Latreille, Dukeril, and nearly all
modem writers; a character, of which some have exaggerated the
value (see ex, gr, Illiger in his Magazin, f, Ijisektenhmde, l 1802,
8. 285), but which has some exceptions'^ and moreover cannot
always be strictly applied, if we would not violate natiu*al aflinity^
The normal number of the joints of the tarsus is in this order, as in
most hexapod insects, Jwe^ and therefore by £at the most of the
^ Comp. Dx Haan, Mhnoirtt mr lea metamorphoaet des ColeopUret. NouveOeM
AnnaUt du MuUum, iv. 1835, pp. 125—164, PL 10—19.
* Revue Entom. par Silbebhann, i. 1833, p. 53.
" Ph. W. J. MiJLLEB, BemerhingenHb. die Fuugliedenahl evniger Ka/ergattungen,
in Illioer's Magaz.f. Ins, 17. 1805, 8. 197 — 219.
* I oommunicated some remarks on this subject id Okbn's Isit, 1824. s. 229, 223.
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INSECTA. 467
genera belong to the Pentamefra, The Heteromera are such as have
an unequal number of joints of the tarsus in different feet, namely,
five in the first four feet, four in the last two. The remaining
divisions, which have been formed, are Tetramera, Trimera, Dimera
and Moruymera; the last two, however, in consequence of later
observations, are obsolete', and there remain only the Trifnera and
TUramera; on which we may remark, that, according to more
accurate investigation, in the Trvmera and in many Tetramera a
small joint has been found situated at the base of the last joint, and
which, when this joint is in its ordinary position, is almost con-
cealed and invisible*. The Trimera are thus in reality Tetrafnera;
they, are named by Westwood, PaeudoPrim&ra (Cryptotetramera
BuBX.). The TetrcMnera of Latbeille are now named by Bub-
XEifirrEB Cryptopefntameray by Westwood Faeudotetramera, The
first three or four joints of the tarsus are commonly broad and
below flat, furnished with small inequalities and beset with little
hairs at the margin. The last joint (the last two joints taken
together in the Paeudotrimera and Paeudotetramera) is small,
inversely conical, elongate, homy and smooth, and terminates with
curved naUs or booklets.
A. Fseudotrimera.
Family XLIV. Coccinellulce {Aphidiphagi Latr.) Body
nearly semi-globose, above gibbous, below plane. Thorax short,
broad, lunate. Antennae inserted before the eyes, capable of con-
cealment under the head, shorter than thorax, terminated by a
capitulmn ob-triangular, compressed. Maxillary palps securiform
(with last joint large, depressed, triangular) ; labial palps filiform.
Feet short ; second joint of tarsus bilobed, broad.
Coccinella L.
Sub-genera: Coccidvla Megerle (Cacicula Steph.), Scymnua
KUGELANN, kG,
* ThuB the PieUxphii (supposed previously to be dimera) have, acoording to Lbaoh,
three joiDts, and Clambtu, the only genus of the so-named monamera, has four joints,
aooording to WxsTWOOD, Inlrod. to mod. Class, i. p. 37.
■ Ph. W. J. MtJLLEB observed this in CocciruUa, Illigeb's Mag. s. ai8. In
Oerambyx there are in this way five jomts. Gomp. also Kibbt and Sfenoe, Introd. to
EntomdL m. pp. 683, 684, and especially W. S. Mao Leat, On the Structure of the
Tarsus in the tdramerous and trimerous Coleoptera of the French Entomologists, Transact,
cfthe Linn. 80c. Vol. xv. 18^7, pp. 63 — 74.
30—2
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468 GLASS YIII.
Comp. L. Redtkkbaohib in Gkbmab's ZeiUckr. f. dieEmamol, y. 1844,
pp. m — 13a.
Sp. Ooecinella teptemptinetata L., Rgbsbl, Ins, n. Sear, ierratr. 01. m.
Tab. n, Pakzeb, DetOfchl. Ins, Heft 79, Tab. 3, Brandt u. Ratzkbubo,
Mediz. Zocl. n. Tab. XX. fig. i. This is throngbout Europe the oommonest
species of these small insects, which in the hmgoage of the people is called
Omen-lieoen-Heers-heettjes, Ma/rienkiifer, Vaehes d Dieu, Lady-birds, Hie
six-footed larvse are elongate, flat beneath, with a small head and twelve
rings, which become smaller backwards. They feed on plant-lice. The
perfect insects have commonly red or brown shield-covers with black spots,
or black shield-covers with red spots. Very many species belong here.
Gomp. SoHOSNHKBR, Synonym, i. 2, pp. 151 — 108, and Hkbbioh-Schjkffib,
Synopsis generis Coceinella, Deutsehl, Ins. Heft isS.
Clypeaster Andersch, (species of Cossyphus Gyllenh.) Body
clypeiform. Head concealed in a pit beneath the thorax. Thorax
of the breadth of abdomen. Antennae with nine joints. Tarsi with
four joints distinct.
8p. ClypeaMer pusiUus, Ahbenb et Gkrmab, Fa/una InseeL Bwrcp, Fasc.
vm. Tab. 10, Gu^BiN, Iconogr., Ins. PI. 51, fig. 5, &c.
Family XLV. Fungtcolce {Endomychidm Leach). Antenns
longer than head and thorax, clayate or incrassated towards the
apex. Maxillary palps filiform, thick. Body oval. Tarsi with
second joint bilobed.
Fvmgu9^inhabUcmtB. (So named from the abode of most of the
species; some live under the bark of trees).
Endomychus Pa YE. (in part), Latr. Antennae towards the apex
incrassated, filiform. Maxillary palps with last joint ob-triangular.
Thorax broader posteriorly, trapezoidal.
Sp. EndomyehiuM coccineus Patk.^, Ckrysomda eoceinea L., Pakzeb, Deutsehl.
Ins. Heft 44> Tab. 17, Dux^bil, Cons. g. s. I. Ins. PL 11, fig. 1, Cuv. R.
Ani., id. iU., Ins. PL 74 bis, fig. 3.
Lycoperdina Latr. Antennae filiform, incrassated towards apex.
Maxillary palps with last joint ovate. Thorax sub-quadrate.
Sp. Lycoperdina hovistce, Endomychus bovistcs Patk., Pakzkb, BeutscM.
Ins. Heft 8, Tab. 4, Cuvibb, R. AnL, id. ill., Ins. PL 74 bis, fig. 4.
^ 6. Patkull, Fauna Sueciea. Ins. 8yo. ii. 1798, p. rt9.
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iNSEC?rA. 469
Eumcyrphus Web. Anteim» clavate, with three terminal joints
broad, depressed. Maxillary palps thick, with last joint conical
Thorax trapezoidal, broader posteriorly.
Gomp. F. Websb, Obterval, entomol. Kilie, 1801. 8yo. pp. ^i, 59.
Sp. Eumorphua marginatut Fabb., DuiciBiii, Con9, gin. $, L Im. PI. 11,
fig. 1 (wrongly named Eumorphut Sumatra^, CuY. R, Ant., id, ill., Ins,
PL 74 bis, fig. I ; — Eumorph. SumatrcB Wbbeb, L I. p. 59, Erotylut quctd'
riguUcUtu Illio., Wiedbicann'b Archiv f. Zool. u. Zoat, i. 1, p. 124,
TaK I. fig. 4, &C.
Note, — Add genera Dapsam ZiKOL., Trockotdeum Webtwood, and some
oilier genera of this family here omitted.
B. Tetramera and {Pseudotetramera.)
Family XL VI. Chmpalpi. First three joints of tarsi spongy
beneath, fiimished with brushes of setaB, third bilobed. Antennas
filiform, or moniliform, capitate, short. Palps with last joint thick.
Body often gibbous, mostly oval, with thorax short, narrow an-
teriorly, with head small.
These insects (the Mb-palpd) are very closely allied to the pre-*
ceding, wherefore Westwood places them in the division of the
Pseudotrimera as a third &mily. They appear to live on Fungi
(Agcmaua, Boletus). The greatest number of species belongs to
South America.
t MaeeUlary palps nuUleiform, with UuA joint trwMvene, very large, re-
eembling a tegrnent of a circle,
a) Internal process of maadUce hiderdate at apex.
Erotyhi8 Fabr. (and JEgtthm ejusd.), Latk.
Comp. P. A. J. DuFOiroHBL, Monographie du genre Brotyle, Mim. du
Musivm, XII. 1825, pp. 30—61, PI. I. II. pp. 156 — 176, PI. in; Th.
Laoosdaibb, Monographie des Erotyliens, Paris, 184a. 8yo.
Sub-genera; Omoiotelus Hope, Erotylua nob. {Bcusis Chevbol.,
Hope, Lacobd., Priotehts Hope, Lacord., ScapJudomorphtLs Hope,
Lagobd., Euryca/rdiu8 Lacord., Z<mcuriu8 Hope, Lacobd., Erotyhis
Lacord.), jEgithus Fabr, Laoobd., Brachysphoenua Lacobd., Gocd-
morphus Hope, Lacobd., Cyclomorphvs Hope, Lacobd., EuphanisUa
Lacokd., Thoniua Lacord. (All from America).
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470 CLASS VIII.
Sp. ErUyUu gigantmB Fabb., Ch/ry9omda gigwdea 1m, Hoxtttuth, NatmtrL
Hut, I. ixe Stuk, p. 386, PL 74, fig. 3, Voet, CoUopt. n. Tab. 33, fig. iv,
DuFONOHSL, L L PL I. fig. ^-y—Erotylut gibbo8us'EAX&., ChrytomeUigibbota
L., HouTTUTK, L L fig. 5, VoKT, CoUopi, n. Tab. 44, tg, tl, DuiiiBiL^
Com, gin, $, I. Ins. PI. 20, fig. 13, &c.
AtUacocheilus Ohevbol.^ Lacord.
Sp. Avlaew^eUut vMaceui, Triplax violaeea Girkab, Famn. In$. Europ,
Fasc. ZIL Tab. 15. Habit, in Croatia. The remaining species firom the
Indian Archipelago and Western Africa.
b) Internal proceu of maxUlcB unidentate cU truncated apex.
Encatistea Dejean, Lacord. Body elongate, sub-parallel.
Elytra mostly dilated at the base, rather broader than thorax.
Species from East Indies, especially from Java. Comp. Laoorbaibe,
Monogr, pp. 33—44.
c) Internal process of maxUUB unarmed, mostly small, dUated or tub-
pubescent,
Trtplax Payk., Oliv., Latr., {Tnplax and Trttoma Fabk. not
Geoffr.)
Sp. Triplax nigripennis Fabb., SUpha russica L., Db Gxbb, Ins, v. PL 8,
figs, n— 15 (Anihribus ruber), Panzeb, Deutsehl. Ins. Heft 50, Tab. 7,
CuviBB, R, Ant,, id. iU., Ins, PL 74, fig. 3, &c.
Dacne Latr. (in part), Lacord. Tarsi with five joints.
Sp. Dacne fasciata, Bngis fasciata Pabb. Ac.
1 1 Maxillary palps vnth last joint elongate,
a) Tarsi with five Joints.
Episcapha DiSjean (in part), Lacord.
Trtplatoma Westw., Lacord.
Note, — Species of genus JBngis of authors.
b) Tarsi with four joints,
Languria Latr., Oliv. Body linear, narrow.
Fhalacms Payk. Body oval, gibbous. Antennae clavate, with
club elongate, triarticulate, last joint longer, conical.
Sp. Phalacrus corticalis, Anisotoma corticate Illio., Pakzbb, DeutsekL Ins,
Heft 37, Tab. la, Stubm, DeiOschl, Fauna, Ins. ii. Tab. xxx. Ac.
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INSECJTA. 471
Family XLVII. Cydica. Joints of tarsi as in the preceding
family. AntennsB filiform, or somewhat thicker towards the
end only, shorter than body. Maxillse without homy tooth, with
external division palpiform, often biarticulate. Ligula thick, sub-
qnadrate or oval, entire or little emarginate.
Gircula/rs. They are ooleopterous insecta of small or only moderate
size, which feed on plants, especially on leaves. The species here
belonging were collected by LiNKiEUS imder the genera Cassida^
Ghrysomela and HUpa,
Phalanx I. Ghrysomela {Ghrysomelidce Leach). Antennae re-
mote at the base, inserted before the eyes.
Chryaomela L. (exclusive of many species), Latr. (Cuv. R,
Am.), Head not covered by thorax, exsert.
A. Antennas short, moniliform, thicker towards the extremity.
Sub-genera: FrcuocvHs Latb. {Helodea Payil, Fabr.), Phoedon
Megerle, Ghrysomela L. (in part), Tim^a/rcha Meo. (apterous species),
Paropsis Oliv., Doryphora Illig., Trochalonota Westw., PhyUa-
charts Dalm., &c.
Comp. Hebbich-Sohaffeb, Auseinandersetzwig der OaUung Chrynrnda,
Deutschl. Ins. Heft 157.
Sp. Chrysomda Phdlandrii li., HdodespheUandrii Patk., Fabb., Duh^il,
<7(m#. gin. s. I. Ins. PL 19, fig. 4, Panzeb, Deutschl. Ins. Heft 83, Tab. 9 ;
— CJirysomela populi L,, Panzeb, Deutschl. Ins. Heft 100, Tab. 5, Batzb-
BUBO, Forst-Ins. i. Tab. XX, fig. 4; Ouvrage posth. de P. Ltonbt, PL 12,
figs. 36 — 4a ; this little beetle, 5'" long, is blue-black, with red-brown or
orange-coloured shield -covers, which have a black spot at the point; the
larvs eat the parenchyme of the leaves of poplars, so that the vascular net
is left like a skeleton leaf; see BATZBBUBa, 1. L Tab. xxi. fig. 6; — Chryso'
melafastuosa L., Pakzsb, Deutschl. Ins. Heft 44, Tab. la, &c.
B. AntenncB filtform, longer than half the body.
Sub-genera: Podontia Dalm., Colaspis Fabr.
Note. — Db La Pobts, Mim. sur les divisionji du genre Colaspis, Bevue
entomol. par Silbebmann, pp. 18 — 15, proposed several sub-genera.
Does genus Megamerus Mao Leat belong here 1 Habitus of PrumuSf
thorax oblongo-quadrate. Comp. Boisduval in Gu^BiK, Magas, de Zool.
1835, Ins. PL 124.
Gryptocephalus Geoffr. (in part), Fabr. Head vertical, almost
entirely intruded within the gibbous thorax.
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472 CLASS viii.
A. AntenwB longer than head and thorax.
Sub-genera: Evmolpus Fabr, Ewryope Daul, Choragus Kibby^
Cryptoceph(du8 anctor. kc
Sp. Orif^ptocepkalM tericeus Fabb., Chrytomda terieea li., Pahzeb, DeictecU.
Ins. Heft loi. Tab. 13, Cuv. R. Ami,, id HI,, Int. Tab. 7a, ^, 4, *»•
The buTflB of Oryptocephaku and of the rab-genus Clythra, are thick and
curved at the posterior extremity like those of the Cockchafer; they five
in cases, which they draw along with them, and which, according to Gbb^
are formed of their excrement. Ann, des Sc. naiur. ZX. 1830, p. 143.
SurFBiAB gave a revision of the Eoropean species not long ago, iMtnaa
entomUog. n. 1847. b. i — 194.
B. AntenniB short, aerraU or pectinate.
Snb-genera: Laanprosoma Kibbt, Chlaanya Kitoch, Clythra
Laichabting, Eabb.
Sp. Clythra quadripuntiata, Chryeomda quadripundata li., "Paxzbjl, DeutechL
Int. Heft 106, Tab. 10, &c.
Note. — ^Almost all the spedes of sab-genos Chlamyt are from Sonth
America; on which comp. Kltjo, Entomol. Monogr, 1844, pp. 85 — 150,
and Y. Kollbb, Monogr. Chlamydum. Gum Tab. sen. color, a. YiemuB
1844, folio. A new species irom the eastern hemisphere {Chi. indica) was
described by Guiaiir {Revue goal. 1840, p. 41).
Phalanx II. Galeructna {GalenusidoB Steph.). Antennas aap-
proximate at the base, inserted between the eyes near the mouth
(filiform, scarcely or not at all inerassated towards the apex, often
longer than half the body).
HaUtca Illig. [Altica Geoffr.). Posterior feet saltatory, with
thighs inerassated.
Sp. HdUicadUiracea, Chrytom/tHa oleraeea It., Panzbb, Deuttchl, Int, Heft ^i,
Tab. I, Batzkbubo, Fortt-Ins, i. Tab. zx. fig. 8 ; a"' long, blnish-green,
shining, with a transyerae furrow on the hindmost part of the thorax.
As larva and perfect insect this little beetle eats all kinds of leaves of
trees, garden-plants and pot-herbs, and may cause great damage if it
multiplies largely. It is known by the name of earth-Jlea, under which
however other species are also included.
Comp. on this genus Illigeb Magaz, Yi. and SoHOXNHXBB Synon» L, s,
pp. 300—311.
Sub-genera : Longitarsvs, Diholia, PayUiodea, (Edumychus Latb.
(Cuv. R. AnL 1829, v. pp. 154, 155), and some others proposed by
English authors, on which cons. Westwood InJtrodact. n. Otneric
Synopsis, pp. 42, 43.
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INSECTA. 473
OiJeruca Geoffb., Fabr. Posterior feet not saltatory.
Sub-genera: Luperua Gboffr., Gaieruca Gkoffr., FhyUobrotioa
Chrvb., AgdcuHca Cheyb., Adariwn Fabb., ice
Sp. Cfaleniea alni, Okrymmda abni L., PANUBy JDeutaehl. /««. Heft loa.
Tab. 3, Ratzkbubo, PoratrlnM. i. Tab. xz. fig. 6 ; 3'" long^ violet-blae,
sbining, dytn with scattered points, feet and antenna black ; this spedes
fiyes on Alnui gkOinota, and is yeiy common ; — Cfaleruea tanaeeU, Chryto-
mda tanaceti "L,, BounL, In$. n. Sear, tar, CI. m. Tab. v. Pansbb, 1. L
T^. 9, &a
XupenM fumOm Wsbtwood, Gu^bih Maga», de ZooL 1837, ^^^
PL 177.
Phalanx III. Casstdina {Caastdarim^ Cassidiadm Leach).
Antennae approximate at the base, remote from the mouthy inserted
in the upper part of the head (short, exsert, filiform or cylindrical,
sub-incrassated towards the extremity).
Htspa L. Body ovato-oblong, with head exsert. AntennsB en-
tirely exsert. Thorax trapezoidal or sub-quadrate.
Sub-genus Alvmua Eabb.
Sp. AVwmuM grouui Fabr., Yobt Ooleopt, n. Tab. 7g, fig. 9, Duv^bil, Chnt.
ffSn. «. I. Ins. PI. 19, fig. 3; — Alumm marginahu Thunb., Bitpe hardi
Latb., Cuv. B. Ani. 18x7, PL xm. fig. 5 (md edit. 1819, PL xm. fig. 5)
ko. Comp. Thunbbbo, Nov. Act. UpstU. ym. 182 1, pp. 199 — aoi,
GniBiN, Note monographique $ur le genre Alumus, Revue Zoolog. pp.
330-334.
Here also belong sub-genera Sotkrionota, Anieodera, CaUielola, and
Pnmecotheca Chxybol. and D^jbak.
Hispa L.
(Thorax and elytra frequently aculeata)
Sp. Hiapa tettcuxa L., Dtjh^bil Cone. gin. $. I. In$. PL 19, fig. 3 ; Ouv.
ErUom. No. 95, PL i. fig. 7; habit, in south of France, Italy, and
Barbary.
Comp. Sohobnhbbb Syrum. i. 3, pp. i — 7. Add sub-genus Chaiqnu
Thukb.
Cassida L., Late. Body clypeiform, plane beneath. Elytra
marginate.
Ckudda L. Head concealed beneath the plane thorax.
Sp. Cattida viridie L., Ba»BL, Ins. n. Scarab, terreebr. CI. m. Tab. Ti.,
Panzbb, DeiiUGhl. Im. Heft 96, Tab. 4 ; green shidd-heeUe, 3V" long*
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474 CLASS VII r.
green^ with a black body. The bura lives on thiBtles, is pale-green, pro-
vided with lateral spines, and is covered on the back by its own excrement.
The perfect insect^ when touched,' draws its feet and antennae under the
thorax, and remains so immoveable that it might be supposed to be dead.
Cassida equeHrit Fabb., Pakzib, DeuUchl Itu. Heft 96, Tab. 5 ; 4'" long;
differs from the preceding species by the yellow feet» and a yellow margin
at the abdomen. — Cauida nobilis, L., &c.
Gomp. for the species of this genus Sohoxnhbbb, Syiwn, L 2, pp. 109 —
330, which however have been since increased by many new discoveries.
F. W. HoPB possesses five hundred of them in his private collection ; and
knows, at the least, two hundred more. The greatest number (four-fifths)
of the known species are fomid in the new worid. See F. W. Hops, O&mt-
wUions on the Toriaue or ShiddrBtdlei, AnnaJU of Natural ffist. in. 1839,
pp. 91— 100, PL lY., where also some new genera are proposed as divisions
of this numerous group.
ImcUidiwm, Fabb. Head received in the thorax, emargiiutte an-
teriorly.
Family XL VIII. Eupoda s. Parameca {CrtocertdcB Leach).
Tarsi as in the preceding family. Body oblong. Antennse filiform
or somewhat thicker towards the extremity, inserted before the
eyes, scarcely longer than head and thorax together. Thorax
narrow, cylindrical or quadrate, receiving the head as far as the
eyes. Posterior feet in most large, with thickened thighs. Lobes
of maxilla membranous, with outer lobe dilated towards the apex,
not palpiform.
Phalanx I. Criocerina. Mandibles broad, with two or three
teeth at the apex. Ligula entire or scarcely emarginate
Megascelis Dej., Latr. Antennae almost of the length of body,
with joints slender, elongate. Mandibles truncated, thick.
Sp. American: Letna viUala Fabb., Cuv. R, Ani., id. HI., Iru. PL 71, fig. a;
Lema euprea Fabb. ; Lema nitidulu Fabb. ; Megatcdu Jkmpei Dij. ;
— Megatcdia cenea Stubm, CkUalog. meitier IfuedenaammL 1816, p. 80^ l^b.
IV. fig. 36, &c.
Crtocerts Geoffr., Lema Fabr. in part. Antennss scarcely
longer than head and thorax. Eyes emarginate.
Crioceris Latr. Posterior femora scarcely incrassated or «iTOilA.r
to the rest
Sp. Ci-iacerU merdigera Fabr., BiU. Syd., Lema merdigera Fabb. SyH,
Eteuih,, ChryMomda merdigera L., R^UH. Ins. m. PL 17, figs, i — 13,
VOBT Coleopt, 11. Tab. 29, fig. r, Panzeb, Deutschl. Int. Heft 45, Tab. «,
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INSECTA. 475
CuY. J2. Am,, 4d. iU,, Int. ¥1 70, fig. S; the red Ulf-beeOe^'^Orioeeni atpa-
ragi, Chryaomda Asparagi It,, iUiSBL, Ins. IL Sear, terr. GL m. Tftb. it.
VOBT, Tab. cii. fig. iv. Pakz. DevttM, Int, Heft 71, Twh. 1 ; a}"' long,
-with a black-blue abdomen, black head, brown-red thorax, with two black
epots, black elytra, each with three yellow spots and a yellow point. The
bora and perfect insect lire on the asparagus.
Petauristes Latb. Posterior femora incrassated
Donacia Fabb. Antenna scaxcelj larger than head and thorax.
Eyes entire.
Auchenia Thunb. Posterior femora not incrassated. Thorax
produced at the sides. Last joints of antenniB broader^ produced
inwardly.
Donada Fabr., Latb. (Donacia and ffcunonia Msa, Dejean).
Posterior femora incrassated Antennae filiform, with last joints
slender, elongata
Sp. Donada tagiUairia L., Panz. DeuMd, In$. Heft 7g, Tab. 7, CuY. M.
AfU., id ill, Ins. PL 70, fig. 5 ; Don, NymphwcB (and Don, ioricea) Fabb.,
Leptara aerigta L. &c. Lennaub enumerated the species known to him
under the genus Leptura, Gomp. on the genus Donacia, D. H. Horn,
Enumeraiio Intedorum dytratorum, circa Erlangam indigenarum, ErlangBo,
1795* 8vo. pp. 38 — 48, with 13 col. figs. They are small beetles, living on
water-plants, between 3 and 5'" long, mostly of a shining green, copper-
lustrous colour.
Phalanx 11. Sagrida. Mandibles triangular, with apex acute.
Ligula emarginate^
Sagra Fabr. Eyes reniform, emarginate at the base of an-
tennae. Posterior femora incrassated, tibisB incurved. Body narrow,
gibbous.
8p. Sagra femorata Fabb., Tenebrio femoratua Dbubt, TeneMo viridii
SuLZEB AbgekOrUe Getch, d. Ins, Tab. vii. fig. 8, Cuv. It. Ant., id, iU,,
Inf. PL 70, fig. a, Oliv. Coleopt. No. 90, PI. i. fig. i, &c. Gomp. on this
genus F. WxBiB, Obterv. EntomoL 1801, pp. 60 — 64. Add Sagr, JBuqtuttii,
Lesson; S, BoiaduvalUi Dupoirr, Lsssoir, lUudr. de ZoU. PL 30,
Gu^BiN, Magann de Zod, 1832, Ina, PL 32, &c.
Sub-genera: AmetaUa, Mecynodera, Hops; Megalopu* Fabb. Head
nodding ; eyes large, protuberant. Body oblong, somewhat broad.
^ To this the genus Poecilomorpka of Hops and Laoobdaibs forms an exception.
Perhaps it were better to distinguish both dirisions by the mandibles alone, or indeed
entirely to drop them.
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476 CLASS VIII.
Gomp. Mannxbheim, ObservaiioM sur U gmre Megcdoput, Mim, de
VAcad. im^r, dea Sc. de St Peteriboturg, Tom. x. (1816) p. ^93, &c., PI. 15.
Here belong sub-genera^ Mariostethus Laoobd., Hcmudoptenu Pkrtt,
AgaUMmeruB Laoobd., Temnoupia Laoobd., and PacUonwrpha Hops.
Orsodacne Latr, Eyes entire. Femora subequal. Maxillary
palps with terminal joint elongate, truncated.
Sp« Ordodacne MoroUca Latb. ; Orioceria Ceraai and Orioc. fulvieoUiM
Fabb. ; Panzbb, DeuUehl. Im. Heft 83, Tab. 8 ; Guv. JR. AnL, id. Ul.,
Ina, PL 70, fig. 3.
Family XLIX. Macroceraia s. Longicomia. Tarsi as in the
preceding family. Mandibles broad, triangular, with apex simple,
acute. Ligula mostly membranous, cordate, emarginate. Antenn®
filiform or setaceous, mostly long, almost of the length of the body
or longer than the body. Form of the body elongate, depressed.
Eyes in many emarginate, a sinus receiving the base of antennae.
The goat-beetles, or long-horns. These insects usually live a long
time in the state of larva and then reside in dififerent parts of plants,
especially in the inside in the wood, whence some species may become
very injurious to the growth of trees. The larvse have very short
feet, or are quite destitute of them, and are vermiform, with rings
strongly marked as though by constriction. The goat-beetles have
some affinity with the group of the Lucanidea amongst the IctmeHi-
eomia; this is however exaggerated, when some writers wish on that
account to place this fiEunily in the neighbourhood of Luccmus L.
Gomp. Audinbt Sxbville, NouvdU ClaaaificaJtion de la fanUUe dea
Longicomea, Ann. de la Soc. entomd. de France, i. 1831, pp. 18 — to, n,
1833, pp. 5«8— 573, IV. 18351 pp. 5—100, 197— ««3-
Phalanx I. L^tureta {Lepturidoe Leach, West wood). Eyes
mostly entire, in some sub-emarginate. Antennae moderate, inserted
before the eyes. Head nodding, broader posteriorly than the an-
terior margin of thorax. Thorax conico-truncated, broader pos-
teriorly. Elytra mostly attenuated towards the apex. Body often
arcuate, with feet long.
Leptura L. (in part), Dej., Latr. Antennas remote at the base.
Thorax mostly not produced into lateral tubercles, with margins
smooth.
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IN8ECTA. 477
Sp. Leptwra rubro-Utlacea Illio. (Leptura tetiaeea L. nuJe, And Zifi.
rubra L. fern.) Pahzbb I>euUchl. Int. Heft 69, T&b. ir, 17; Ratzsbubo
Fortt-Ins, I. Ta£ xvn. %8. 9, 10. Body, head, antennaa, and thighs
black, tibiae and tarsi ruddy, shield-oovera of male yellow-brown, of
female red-brown, as also the thorax, length of male 7'", female 8'".
Lepiura armata Pbetbl., Sohobnh., Strangalia armaia Sebyillb {Lep-
tura calcarata Fabb. male, Lept, tubapino$a Fabb. fem.), Yoet, Coleopi,
n. Tab. zxvi. fig. 5 ; Leptwra tomeniota Fabb., Out. Coleopi. No. 73,
PL 7, ^. 13, G. Duicfon. Cons, g4n, $, l. Int. PL 18, fig. 2, fto.
(Sab-genns ffeteropalput Buquet.. Mazillaiy palps with a pilose
appendage at the base of last joint. GuiBiN Magatin de ZoU,)
Bhagium Fabr. (with addit. of some species oi Lepiura ejasd.),
St&nocorua Geoffr., Oliy. Antennce approximate at the base,
inserted on two tubercles between the anterior margin of ejes.
Thorax on both sides tuberculated or spuiose.
Sp. Bhagium inquititor Fabb., OaramJb, inqutiitor L., Pakzbb DeuttcAZ.
Ins, Heft 8a, fig. 6; — Bhag. mordax Fabb., Yoet Coleopi, n. Tab. 95,
^, I, DuMiBiL Cons, gin, s. I. Ins, PL 18, fig. i, fto.
Vesperas Dej. Elytra of females short, soft, wings none. Thorax
not tuberculate. Insertion of antenne as in Bhoffium,
Sp. Vesperus Solieri D^jban, Gebxab Faun, Ins, Bump, xvm. Tab. 40 ;
CUY. B, Ani., id. ill.. Ins, PL 69, ^. a,
Desmocenia Dej.
Sp. Stenoeorus eyanenu Fabb., Cuv. B. Ani., M, iU,, Ins, Tab. dt. ^g, i,
Gu6siN leonogr., Ins. PL 46, ^, i. Hab. m North America.
NoU. — Several genera and sub-genera of more modem authors, as in this
whole family, must be here omitted for the sake of brevitj.
Phalanx II. Cera7nb{cina{Cerambycid<BJjEkCB). Eyes reniform,
smrronndiQg the base of antennae with an internal sinns. Antennae
long, sometimes longer than body. Head received in thorax as far
as the posterior margin of eyes ; thorax not narrower anteriorly, nor
separated from head by constriction. Labrum exsert, of the breadth
of the anterior margin of head. Maxillae with two distinct lobes.
Lamia Fabr., Latr. (with the addition of genns Saperda
Fabr. and Steiwcerus ejnsd. in part). Head vertical, inflected.
Palps filiform, with last joint oval or subcylindrical, attenuated to-
wards the apex.
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478 CLASS viii.
&) With thorcue unarmed (not tpinoae or tuberciUate).
Saperda Fabb. (with the addition of species of Stenooorus ejusd.).
Sp. Saperda earcharicu Fabb., Cerambyx c<xrcharia8 L., Pakzeb DeuUehl^
Ins. Heft 69, Tab. i, Cuvier R Ani., id. iU., Ins. PI. 68, fig. 9, Katzb-
bubo Forst-Ins. i. Tab. xm, fig. 4 ; i" to i" a'" long, antennae somewhat
shorter than body, ash-coloured or yellowish-grey, with many shining,
black, raised spots on the thorax and the elytra. The larva lives in
different species of poplars ; — Saperda populnea Fabb., Cerambyx popul-
neus L., Panzeb DetUscM. Ins. Heft 69, Tab. 7, Batzsbubo L L Fig. 5,
small 5"' to 6'" long, antenns of the length of body, brownish with
black spots and four yellow spots on each shield-cover ; three longitudinal
pale-yellow stripes on the thorax ; antennse ringed with white and black ;
Saperda oevlcAa, Cerambyx oculaius L., lives on Populus tremula, PAifzSB
DeutsM. Ins. Heft i. Tab. x8, Ac.
Note, — Sub-genera Adesmiu, Apomecyna, Colohaiheaf &c. proposed by
D^jban and others are here omitted. Sub-genus AgaparUkia Sebv. is
distinguished by antennse with twelve joints. Sp. Saperda Cardui Fabb. &a
b) With thorax spvnose or tvhereulate on both sides.
Lamia Fabb.
Sp. Lamia cedilis Fabb., CeramJ)yx csdUis L., Acanthocinus OBdHis MeoebIiB,
VOET C(deopt. n. Tab. iv. figs. 1—3, Ratzebubo Forst-Ins. i. Tab. xvi.
fig. a ; antennae in male four times longer than body, &c.
AorocvMis Illig., Macropus Thukb.
Sp. Cmmbyx longimanus L., Bosel Ins. n. Scar. terr. Prcrfat. CL n.
Tab. L fig. a, Vobt Coleopt. n. Tab. xii. xin. fig. 51 ; «A« carpetrbeetle,
VArlequin de Cayenne (lives in South America) ; a^" long ; the antemue
are 5" and the fore-feet more than 4" long ; the four other feet are much
shorter ; the thorax has on each side a moveable spine ; the shield-covers
are truncated at the extremity, and armed with two sharp, homy points.
Note. — In some species of this phalanx the eyes are divided into
two separate parts by the insertion of the antenn», the one situated
above the other below the base of the antennsa. It is thus in
Saperda prcBUsta Fabb., Leptu/ra prcmsta L. (genus Tetropa Ejbbt,
AncBtia Dij.) and in some exotic species of Lamick, 6. g. Lamia tor-
fuUor Fabb., which compose the genus :
Tdraopea Dalm. and Schoenh. (See Syrwn. Insect, l 3, pp. 401,
402.)
Obrium Meig., Diu.
Tjneststemus Latr.
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INSECTA. 479
Necydalxs L. Head exsert or nutant, not inflected vertically.
Palps filiform, tmncated at the apex. Elytra less than wings;
wings expanded in almost their whole length, replicate at the apex
alone. Abdomen elongate, narrow.
SteTiopterus Illio. (Species of Necydalia Fabb.). Elytra subu-
late, of length of abdomen.
Molorckaa Fabb. Elytra oval, short, not produced beyond the
base of abdomen.
Sp. Necpdaiis major L., Molorchut abbrtviatui Fabb., Gbokoy. Zoophylae,
Tab. XV. ^. s, DuxiBiL Cons. g4n. $. I. Itu. PL i8, fig. $, Pahzxb
JkuUchl. Ins, Heft 41, T&b. ao, &c. Comp. Hebbioh-Soosffeb DeutrnM,
I'M. Heft 120.
Note. — Stenopterus molorch&ides, & species from Chili, described by
Guf&iN, Magaa. de Zocl. 1838, Ins. PI. 333, ^. a, appears to form the
transition between the two genera.
Cerambyx L. (exclusive of many species). Head exsert, nutant.
Palps truncated at the apex, with last joint ob-triangular. Elytra
covering wings and abdomen.
CaUidium Fabb., Oliv. {Clytvs, CaUidmm Fabb. and CcMrtcUlum
Mbgeble, D£j.)
Sp. OeramJbyx he^us L., Pakzbb DeuUehl. Ins. Heft 70, Tab. i ; 10'" long;
the round, flat thorax felty, with two tubercles, broader than head ; the body
black-brown.
Cerambyx Fabb. (in part).
(Here belong many sub-genera of more modem authors, Gnoma
DijEAK, Sienocorus Dalu., Dej., ffammaticherus Mboeble, Calltr
chroma Latb. {Aromia Sebv. in part), Phosnicocerus Latb., Ctenodea
Oov., Kluo, Lophonocerua Latb., Trackyderea Dalm.', Dorcocerua
D£j., Mtgadema Dij., Lissanotua Dalil, iic)
Sp. Cerambyx moschatus Ja., VosT CoUopt. n. Tab. vi. fig. 14, RA.TZBBUBa
Fars^Ins. I. Tab. xvi. fig. i, &c.
Phalanx III. Priamna {Prtonidce Leach). Eyes emarginate.
Antennae of different length, inserted before the eyes. Labrum
small, occupying the middle part of the anterior margin of head
^ Gomp. on this genus Dttpont, Monographie des TracydSridea, GuiBiir, Magas.
de Z06L 1836, Ins. Tl. 141^146, and 1838, Ins. H. 186—100.
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480 CLASS VIII.
between the mandibles. Maxilte with only one lobe or with two,
internal very small ; mandibles strong, especially in males. Thorax
not separated from head by constriction anteriorly.
A. Antenna Umger than head and thorax together.
Primus Geoffr. Fabr. Body depressed. Thorax with sides
acute, often dentate or spinose. Antennae in some pectinate or
serrate, in others simple, attenuated towards the extremity. Man-
dibles of males sometimes exsert, long.
Sp. Priontu coriariue Fabb., Cerambyx coriariiu L., B<esbl Ins. n, Seartd^.
teir. CI. n. Tab. i. figs, i, a, Tab. n., Panzks DeutecM, Int. Heft 3, Tkb.
9 ; in Crermany and other parts of Europe.
To this genus belong some gigantic exotic spedes, e.g. PrionuM eervieomu,
Cerambyx cervicomie L., E<B8XL Ine. n, Scar. terr. CL n, Tab. i, fig. b ;
VoKT. CoUopt. n, Tab. n, figs. 4, 5 ; Cuv. M. Ani. Sd. ill., Int. PL 64,
fig. 5, from South America. Prionut ffayesii DOWMBB, HoPB Tranaae. of
the Zool. Sac. l, PI. 16, from the west coast of Africa near the Equator.
Some species, with short narrowed elytra, which occur in South America,
form the genus ATwcclut of Latbbillb ; other species, also firom South
America, haye besides no wings {Prumaptenu GuiBiH).
B. Antenna thort (of ike length of thorax). Head of the tame breadth at
thorax. Thorax unarmed.
Spondylis Fabr. Ligula membranous, cordate. Thorax glo-
bose. Body convex.
Sp. Spondylit huprettotdet Fabb., Attelabut hapretMfdet L., Dmcfan. Cont.
gin. 1. 1. Int. PL 17, fig. 6 ; Satsebubo Fortt-Int. i. Tab. zvn. ^. 13.
Parandra Latr. Ligula homy, Imiate, transverse. Third joint
of tarsus scarcely bifid, last longer than the rest taken together.
Sp. Sparanda hrevit Latb., Oen, Or%atac. et Int. Tab. 9, fig. 7 ; — Parandra
glabra Gtllenh., Attelabut glaber De Geeb Int. iv. Tab. 19, figs. 14 — 16,
SeariUa tettaeeut Fabb. The fom-th joint of the tarsus is here more appa-
rent than in the other Pteudotetramera, so that some count this genus
amongst the Pentamera.
The habitus of these insects resembles that of LuoANua Latbbillb
first placed this genus in the neighbourhood of Cfuet^ {ffieL not. det
Ortut. et det Int. Tom. xi. p. 352) and firom later writers also it received
the same place; the affinity however with Spondylit seems to assign to
Parandra a place amongst the Prionidof.
Family L. Scolt/tarta {ScolytidcR Westw., Xyhphagi Latr.
in part). Third joint of tarsus in some bilobed, in others entire.
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IN8ECTA. 481
Antennse shorter than thorax, with first joint elongate, terminal
joints forming a club. Head often protracted anteriorly for a while.
Palps small, conical.
A small family of small insects, called by the Germans Borken-
kd/er, hark-heeUeSy although many do not lire in the outer bark
{Barke), but in the hast or liber and some in the wood Some of
the species cause great injury to needle-leafed trees (fir, larch, <fec.)
The larvse have no feet, any more than those of the following fiunily,
with which the Scol^/tini, as already remarked by Westwood, might
almost be united Also the nervous system confirms the affinity.
In the ablomen there is no chain of ganglia, but they are all united
to form a single ganglion oblongo-conical, which is situated close to
the second thoracic ganglia (formed from the imion of those of the
meso- and metathorax). See the figure of the nervous system of
Scolytus pygmceus in the memoir of Blanghabd Arvn, des Sc. not,
3i^me S6rie, Tom. v. 1846, Zool, PL 13, fig. 8. Some affinity with
the Scarahandea cannot be mistaken.
Compare Ebtohson SydemaJtiaehe AuteinBmdenasung der Fcwniiie der
Borhaikafer (BostridudsB), Wiboxann'b Archiv /. NtOwrgetek. 1836,
8. 45—65.
Platypus 5eubst, Latr. Head exsert, broad, transverse. Tho-
rax elongato-cylindrical. Body cylindrico-linear. Antennae sexar-
ticulate, short, with last joint expanded into a club plane, ovate,
large. Tarsi with all the joints entire, first long, slender.
Sp. Platypus cylindrieut, Battrichut cyUndrus Fabb., Pakzsb DeuUchl. Ins.
Heft 15, Tab. 2, Ratzibubo Ford-Ins. Tbb. X. ^, 13^.
Tomicus Latr., Bastrichus Fabr. (in part). Head retracted,
vnth thorax produced anteriorly above it. Antennae inserted in the
sides of head between the base of the mandibles and the eyes.
Tarsi with all the joints entire.
Sp. TiinUeus typographuSf Dermestes typographus L., Pavzbb DeutseM, Ins.
Heft 15, Tab, 3, Ratzibubq Forst-Ins. I. Tab. it, fig. i ; forms in fir-
trees beneath the bark labyrinths and passages resembling letters or
characters ; hence the name of Typographer.
^ Here also seems to be the place of Tesserocerus Saundbbs, where the first
antennal joint (in the male) has interiorly a curved process^ and the tarsi five joints.
A BraoUan insect oooaaioned the formation of tUs genus, yiz. Tesserocerus insignts,
Dcmieerus agilis Spikola, Gu^bik Magas. de ZooL 1839, ^^- ^* 3*
VOL. I. 31
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482 CLAfiS viii.
Here belong sab-genera Xylotenu Ebichbobt {Trypodendron
Stepe.), Hopotenemut Webftw., £bigh&, &c.
8cohftu8 Geoffb. {Hyleainua Fabr.). Head partly received in
thorax, produced into a roBtmm short, thick. Tarsi with third
joint cordate or bilobed.
Sp. Scolyhu Ulmi DnxiB., ffylainus Sedytus Fabs., Gsoffbot ffiM. cTIns,
I. PL V. fig. 5 ; DnxiBiL Com. ffin. s. I, Jm. PL 17, fig. 3 ; Satzxbitbo
Font-Ina, Tab. z. fig. 4 ; thu ipaciea often deetroyi qnaiititiea of dnu.
Sub-genera: PhlceotrUnu Latb., Eceoptogaster Hebbst, CampUh
eerua D£r., Latb., Hylesinua Vabsl (in part), Latb., and Hylurgus
Latb., and some others proposed by Ebichsok, L L
Family LI. i?%7icAopA<>ra ((7urcw?ibniiie5ScHOENH.). Head an-
teriorly rostrate, with mouth at the extremity of rostrum. Antennae
inserted in rostrum, very frequently clavate at the extremity. Third
joint of tarsi almost always bilobed. Mandibles very firequently
small, strong.
Beaked-beeUe$. These are all herbivorous and gnawing insects.
Their larvaa have no feet or only tubercles in place of them.
A numerous £unily, of which above 7,000 species are known,
which in the last work of Sohobnhebb form more tfafen 630 genera \
Compare on thia fiunily, amongst othen :
[CLAiBvn<Li] Entomologieffelvetigue, VoL i. Zurie, 1798, 8to. av. fig. ooL
— G. P. TBJSVSKBQdeOoleopierii rtObratit commmtaUo, Nov. AcL Soe. UpeaL
VoL 7, 1815, pp. 104—115 in 0. J. SOHOSNHIBII OWeiiKoiitAim Di^o-
iiiio meihcdieci, Lipeis, 1S36 ; and Syn, imedor,, eee above, p. 465.—
Latskille JthyneophortM, in Dia. datt. d'BUi, not. Tom. ziv. 1818,
pp. 584 — 603. — ^The celebrated German entomdogirt Oibmab, by diffsrent
oommunications in the Magadn of Iluobb, and in that edited by himself
and ZivoKXN Magaein der BtUomoloffie (tv. Bd., iSsi, pp. 191 — 545,
Oenera quadam, Oirculioniium propa$Ua) and by descriptions of newspedes
and genera in his Coleopteror. tpecies nov. Hal», 1834, 8yo. pp. 174 — ^6t,
has conferred great seryices with respect to this fiunily.
A. Palpa very shorty conical. Labrum inconspicuous.
Phalanx I. Curculionita (genus Curculio L. in part). Antenn»
either arcuate or geniculate, with basilar joint or scapus much
elongated.
^ Thus, about 300 more than LnrKiinB, in the isth edition of his SyHma
adopted for the whole animal kingdom.
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IKSECTA. 488
t Bostrom elongate. Antennas genicnlate, inserted in the mid-
dle or in the base of rostrum.
a) AfUenna tmth anly ten joinU or fewer. Clvb buMrUctdate or
tuXheelid^ spongy cU the extremiiy.
Dryopkthortia Schuppel, Sohh. Tarsi with five joints. Antennae
sezarticulate, short. Body elongate.
Sp. Dr. Lymexykn, Lubum LyvMOBylon Fabb., Pakzxb DeiOteld. Int. Heft
i8. Tab. II, Gu&or Icpnoffr., Ins. PL 39 bis, fig. 8; under oak-bArk in
Sweden and Greimany.
Cosaonus Clairv. Antennae short, with eight joints before the
club ; club indistinctly articulate or sub-solid. Bostrum defected.
Body elongate, winged.
Sp. Coi$onu$ ferrugintm Claiby. (and linearis ^uad.) Eni/om. hdv. i.
Tab. I. p. 60, Panzxb JktUschl. Ins, Heft 18, Tkb. 7.
Sub-genera : Microxylchiua Chbvb., EhynchokLs Creutzeb, Amor-
phocerua Schosnh. and others of this author ; Gen^ et Spec Cwrcur
lion. IV. pp. 989—1087.
Calandra Fabr., Oliv., Rhynchophorus He&BST, Schoenh. An-
tennae with seven joints before the club, inserted towards the base
of rostrum. Elytra not covering the deflected apex of abdomen.
Body winged.
Sub-genera: CerddoceruBy Trigonotarstu GuiRiK, Mcicrochirtis
De Haan and several of Schoeitherr; comp. 1. L pp. 816 — ^988,
viiL 2, pp. 216—264.
Sp. Calandra palmarwn Fabb., Cfureulio palnuurum L., Voir CoUopt. n.
Tab. 37, fig. 45, Guvibb R. Ani. idU. HI,, Ins. PI. 60, ^g. 7 ; Sonth America ;
—CdUmdra ferruginea Fabb. , SuLZBB Oesck. d. Ins. Tab. 4, fig. 5, Cwreulio
henUptertis, Jaya, &c. ; large exotic species. In Europe, small i^eoies alone
are met with of this group ; everywhere dispersed, e.g. is Calandra granaria,
OurcuUo granarius L., Lbbuwbnhobok 57e Missive of 6 Ang. 1687 (Fer-
volg der Brieven, &c. bL 75 — 95, with a figure opposite to p. 84); Pabzbb
DeuisehL Ins. Heft 17, Tab. 11 ; the common weoU. This little beaked-
beetle may cause gpreat damage in granaries if the com be not stizred or
turned, just like the w>lf, Tinea graneOa, p. 394.
Bhtna Latr., Oliv. Antennae with geven joints or sometimes
six before the club, inserted towards the middle of rostrum. Elytra
covering the extremity of abdomen. Body winged. Feet, especially
anterior, very elongate.
31—2
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484 CLASS VIII.
Sp. Mina har^rotirUf Lixui barbirostrii Fabb., Sytt. Eleid,, Voir OcieopI,
n. PL 35, figs. 1, 3.
ffarpcusteruB, Sipaiku, SphcBrognaihui, Strombaeuruty Sohoenhebb,
L L IV. pp. 802 — 816.
I^oie. — ^Here also Latbxillb refers genera Anehonu8 Schoxnh.
and Orthochoftea Gericab (Styphlus Schoenh.), which are oonnted
amongst the Erirhinidea by Schoenhebb, L L Tom. UL ppi 507 —
512.
b) Antennoi with nine or ten joints, dub toUh three or four
joints.
Cionue Clairv.
Sp. Okmui teropkidairim, OwreuUo ScrophtOarUE L., VoR CfoUopt. n. Twh. 38,
fig«. 30, 3«.
Mecimts Osbmab, Nimophyes, GynrnUron Schoenh. L L Tom. nr.
pp. 743—789.
c) AnXmmm tot(& dwen or tunl/ve joints^ dub with four joints.
Oryptorhynohus Illig. Anterior feet distant at the base. Breast
channelled for receiving rostrum between the anterior feet.
Here belong many genera of Schoenhebb which must be omitted
for brevit/s saka See his Tom. iv. pp. 1 — ^721.
Sp. Oryptorhynchiu Lapatki, Ckuvulio Lapathi L., Von OdUopt. n. Tah. 39,
fig. 40, PAms Deia$eJd. Ins. Heft 49, Tab. 15, fto,
Baridius ScHOENH. (with addition of other genera). Anterior
feet distant, with breast indistinctly snlcate or plane.
Genera : Baris Gebh., D£r., Leptosehoinus Kluo and many others
proposed by ScHOSiraEBB; see his Tom. in. pp. 638—821, vin.
pp. JOl— 293.
Ahides Dalm.
Amerhinus Sahlb., Schoenh.
Cholus Germar, Schoenh.
On this and some other genera^ to be inserted here, comp.
Schoenhebb iil pp. 557 — ^636 and vm. pp. 1 — 101.
Orchestea iLLia. Anterior feet approximate at the base; pes-
terior feet saltatory, with femora thick.
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INSECTA. 466
Small leaping beaked-beetlM, of which the laiTBB feed on the paren-
chyme of leayee. Sp. OrehaUt Aim, Cwreulio AJmi L., Vokt OolUofi, TU
Tab. 40, fig. 51, DuMiBiL ConM, gin, s, I, Tn». FL 16, fig. 8; — Orehettei
populi, Bk^nchanm popuU F., Panub J}eut9ehl. /lu. Heft 18, Tab. 17 ;
Oreheda fagi, Mgnchanui fagi Gylluth., Batzbbubo Font-Ipg, i.
Tab. iv. fig. 14, &C.
Tach/yergea Schoenh.
BhyncJuBfiua Fabb. (in part), Enrhintdes ScHOENH. Anterior
feet approximate at the base ; posterior not saltatory,
Bala/nmu8 Gebmab. Rostnim slender, arcuate, yery long (longer
than half the body, sometimes than the whole body).
Sp. Bhynekanm mtctim, CwreuUo mteum L., Boesxl Tn», m. Suppl, Scar,
terr. CL rr. Tab. 67, figs. 1—6, Panzxb Ikuttchl, Tw. Heft 42, Tab. ai.
Anthonomui Gebmab. Rostrum of the length of head and
thorax, little arcuate. Feet somewhat long; anterior longer, with
thighs thick, dentate. Body sub-oval, pubescent.
Sp. Jthynehcmut pomorwn Fabb., Chirculio pomorum L., Voir CoUopt, n.
Tib. 35, fig. 8 ; Batzbbubg ForH-Ine, 1. Tab. v. fig. 8 ; V. Bbbck Ver-
hondding ten hewifze, dot niet de moaaU vliegen, maar de clifamtt-of tnuU-
tom^ de hederven tijn va/n de hkanen der vrwMboomen, Haarlem, 1817,
8to. (figure on the title-page.) Thu litUe beetle bores in the blossom-buds
of the apple a hole with its beak, and lays an egg in it ; the larva oonsnmes
the anthers and the pistil ; the blossoms become brown, and dose round
the little animal ; after four or five weeks the perfect insect makes its
appearanoe from a round aperture in the spoiled blossoms.
Add many genera or sub-genera which must be here omitted;
comp. ScHOENHEBB Tom. iiL pp. 154 — 489, vii. 2, pp. 27 — 369.
Lixua Fabb. Bostrum thickish, little arcuate. Antenn» short,
with club oblong, fusiform, four-jointed. Thorax conical, of the
breadth of elytra at the base. Body elongate, pubescent, covered
with dust (&rinose). Elytra sometimes dehiscent at the apex,
acuminate.
Sp. Lianu tvrbatua Gtllbkh., Cfurculio parapleeticu$ Pahzbb DeutscAZ. Iru.
Heft 6, Tab. 15, DuhIbil Cone. gin. 8. I. Ine. PI. 16, fig. 10, Eysbs-
UAifJK Bullet, de la 80c. Imp. de Motoou xvj. 1843, pp. 530^534^ ^^ "^ni.
f. 35 (metamorphosis). This species is often confoimded with Liaeue para-
plecHcut L., Ch^cuUo phdlandrii Db Gxxb Ine. v. Fl. 7, figs. 3—9. The
lary» of this beetle live in the hollow stalk of Phdlandrium aguaiicum and
other umbel-flowering plants.
Comp. on this and other allied genera Sobobkhbbb hi. pp. 1 — 154,
^ VII. I, pp. 418—479, 2, pp. I— «6.
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466 CLASS Till.
tt RoBtnun short, thick. Antennae inserted near the apex of
rostnun.
a) Antenna taUh eleven or twelve joifUSf masUy lon^
Ourculio L. (exclusive of many species), Fabr., Latb.
Cleanue Schoenh. Bostrum nutant or deflected, longish. Club
of aotemuB elongate, gradually inerassated Thorax postanorly
almost of breadth of elytra
Sp. Owculio glaucut (Fabb.!) Gtllbhh., Pakzu in Natwfoneker zxrir.
^ i> fig- 30, VoBT CUeopt, u. Tab. 40, fig. 51, fto.
NoU, — On tliu and many alUed sub-genera compare Sohoinhxbb L L
Tom. n. pp. 17 1 — 316, vi. 1, pp. i — 295, and on European species of
genus CUomkt Hjebbioh-So&btfeb DeuUchL Ina. Heft 173.
Molytee Schoenh.
Here belong sub-genera Ocmiatiu, Lepynu, PUnthuM, TVmytpAjfnif
GXBMAB, HyMmu Sohoxnh., and others, on which compare Schokbhibb
n. pp. 331^408, and VI. 1, pp. «95— 389-
Otiorhyn(^u8 Oebuab. Antenme long. Wings mostly nona
Here is to be referred sab-genus HyphofnlOvui Gbbmab, Schobnhkbb^ and
several other sub-genera of Sohobhhbbb, on which compare n. pp. 551 —
645, vn. pp. 957 — 417. 8p. Cwcvilio owUut L., Ourculio pabulima
Pakzbb DeuUdd. Ins. Heft 57, Tab. 19. Compare also, J. Waltob
Notet onikegenm OttorhynekfU, Aim, cf Nat. Bui. ZIX. 1847, pp. 445, &c.
CydoTims Schoenh.
Here refer sub-genera Amycterui Dalm., Epi$omui Schobith., and some
others proposed by Schobnhbbb, on which compare n. pp. 469—55 1, vn.
I, pp. 51— 156.
PhyUdfnue Schoenh.
Of this pUoe are sub-genera AfMyrhrnvM Sghobbh., Macropi Kibbt, and
others, on which see Sohobbhbbb ii. pp. 4^4 — ^469, and vn. i, pp. i — ^5a
8p. OuretUio pyri L., Ac., Pavzbb Devtackl. Int. Heft 107, 1M>. 4, &c.
Byreopa Schoenh.
Here is to be referred sub-genus LUhodua Gbbmab, and some others,
on which compare Sohobnhbbb n. pp. 408—434, vi. 4, pp. 389 — ^464.
BrcLchyderea Schoenh.
Here are to be referred ahnost a hundred sub-genera^ nearly all pro-
posed by Sohobnhbbb ; see i. pp. 515 — 655, and n. pp. i — 170^ v. p. 853,
at the end and vi.
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IN8ECTA. 487
PolydroiUi Qebm., Sohoshh.
Sp. Chtreulio $erieeu» Gtllekh. fto.
ChloropkoMU Vaim,
Sp. Ourculto vvridia L., Sulzeb Kennuiehm d, Im. Tab. m. fig. 14;
S0H.SFFIB, Elem. EfUomol. 1766, Tab. 108, {BMnomacer), I^ahzsb DeuttcM,
Ina, Hefb 107, Tab. 3 ; green, with longitadinal furrowB on the shield-
covera ; beneath, and also along the maigina of the ahield-ooTen, yellow ;
length, si"*; veiy oommon with lu.
Bra^^deres ScHOizra.
Sp. Curculio inccaius L., Pakzxb l>eutKkl. In$. Heft 19, Tab. 8, Batssb.
Porst-IuB. I. Tab. lY. fig. 4, &c. wmglew, greyish brown.
OMorMnm Sohoknh.
Sp. Owrculio geminatuM Fabb., 3'" long, with very short beak, very round
abdomen, wingless, grey brown, with white felt-hairs ; yery common on
our sandbanks.
Fachyrhj/nckua Germas.
Here are to be referred several sub-genera, most of them apterous, pro-
posed principally by Sohobnhebb. See his Tom. I. pp. 499—5159 ▼•
pp. 800 — 853.
ErUimua Germab.
Gonoemmg seyeral sub-genera to be referred here, see SOHOIHHXBB i.
pp. 444 — ^499, V. pp. 713 — 799 f apterous, most from Southern Africa (sub-
genus Hipporhinus Sohobbh., with seyeral species); winged, some fit)m
South America, especially BraziL Sp. Ourctdio imperialis Dbubt, Fabb.,
Lindbkbbbo Natuffancher x. p. 86, Tab. n. fig. i, Olivibb Ooleopi,
No. 83, PL I, fig. I, CuYiBB Jt, Ani, id. iU,, Ins, PL 57, fig. 6.*
b) Antetrnw wUh only nine cUaHnct jovnts, short, thick, arcuate,
Brachycerua Fabr. Body apterous, with elytra connate, gib-
bons. Feet strong, with tarsi not spongy below.
Sub-genera : Brachycerue Schoenh., Frotomantia Schoekh.,
Microcerua Gtllenh., Sghoenh. Comp. Schoenh. l pp 385 — 444,
V. pp. 605—730.
Sp. Brachycerua aptenu F., CwrcuUo apterua L., YoxT Coleopt, n. PI. 33,
^, A. — Brachycerut obeaua Fabb., Sulzbb Otich, d. Int, Tab. iv. ^. 10,
*
^ This insect is commonly called JHamond'hedle, and in gorgeous colour exceeds,
perhaps, all other natural objects.
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488 . CLASS VIII.
CuYiKB R, AnL id, iU., In*, PL 57, fig. 4 ; both frcnn tiie Gape of Good
Hope, like most species of this numerous genus ; a few oocur also in the
south of Europe, as Brackife, tnuialtts, Brackffch^ de Barbarie DuxiBiL
Com. gin, ». I. Int. PL 16, ^, 4.
Phalanx 11. AUelahita {Attdcibidea Westw., genus AtUlabua L.
in part). Antennae straight.
Attdabus L., Fabr. (in part), Bhinomacer Geoffr., Clairv.
(not Fabr.). Antennse eleven- or twelve-jointed, with three or
fonr last thicker, forming a club. Body oval, narrowed anteriorly.
Third joint of tarsus bilobed.
Sub-genera : Apian Herbst, Bhynchites Herbst, Attelalni9 Latb.,
ApoderuB Oliv., and others, on which oomp. Sohoenherr l
pp. 187—240, V. pp. 247—309.
On genus Apion compare EIibbt'B Monograph in lAnn, TroMocL Tom.
X. and XI.
Sp. AudabiU eoryU (and AUd, aveUanm) L., Sulzib Kmnz. der In*. Tab.
▼T. f. 2$, Pakzbb DetUtM, Ins. Heft 100, Tab. 8, Batzeb. Font-In». i.
Tab. ly. fig. 5 ; this little beetle rolls up the leayes of hazels and other trees
to form a well-dosed case, in which the Unra liyes. (See fig. in Bake-
BUBO, bl. 97.)
AUdabut frumenJtariua Fabb., Owreulio frumentmiu L., Claiby. Ent.
hdv. I. Tab. 13, f. I, 1, Panzbb DeutschL Ins. Heft 20, Tab. 14, &c.
Ramphvs Clairv. Posterior feet with incrassated femora^ sal-
tatory. *
Sp. Ram^hus jUivicomis Claibv., EnJUm. hdv. i. Tab. is (Latebillb
places this genus in the preceding diyirion next Orehestes).
Rhinotia KiRBT.
AuUtea Schoenh.
On these and some other genera, omitted here for the sake of brevity,
comp. Sohobnhbbb I. p. 943, and folL v. p. 345, and folL
Camarotus Germ.
See SOHOKKHBBB I. pp. 1 85— 1 87.
Cylaa Latr. Antennae with ten joints, last very long, thicker,
rest very short, transverse. Thorax constricted in the middle.
Compare Sohobnhbbb i. pp. 369 — 373.
Uhcerua SCHOENH.
BrenttLs Fabr., Latr. {BrerUhua Illig.). Antennae filiform, not
clavate, eleven-jointed. Last joint of tarsi bilobed. Rostrum
exflert, often very long. Body linear, elongate.
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IN8ECTA. 489
Sp. BreniuB anehoroffo, CwreuUo cmehorago L,, Voir Ooleopt, n. PL 34,
figs. I. n., DuMiBiL Cfont. gfn, $, I, Ins. PI. i6» fig. 11, South America ;
— Breml/vti (EvUrackeUa) TtmrnAnM Latb., from Jara ; ^, in IKcUonn,
vnvv. d^HiM, not., CU, PL 9, fig. i.
Note. — See other rab-genera described in SoHOBVH. I. pp. 313 — 369, v.
pp. 465—580.
B. Palps Jilifarm or thicker towards the apex, Labrum distinct.
Phalanx m. Bruchidea {Brtichidce Leach).
Anthribtts Fabb. (and Bhinofnacer ejnsd. in part). Antennas
clayate. Eyes in most entire. Third joint of tarsi small, often sub-
included in second.
Sub-genera : Pla/tyrhinus CLAntv., Tropideres Schoenh., Braehy-
ta/rsua Sghoekh. and several others; oomp. Sghoenhebb I. pp. 115
—185, V. pp. 147—277.
Sp. AnOmimi icabrotut F., Panzib I>euUehl, Im. Heft 15, Tab. 15, &e.
Mecoeerus Sghoenh. Antenuse in females passing gradually into
a club oblong, acuminate at the extremity, of the length of head
and thorax, in males longer than twice the body, attenuated at the
apex. Thorax in males armed beneath with a pair of spines.
Sp. Mecocerua gasdla Sohobkh., AecuUhothorax longieomU Gj&de, in GuiteiN
MagoM, de ZwA, 1833. Im, PL 15, habitat Jaya. Male resembles Lamia
tgdilis, in the length of antennae.
Bruchus L. Antennae filiform, gradually thicker, with short
joints, often serrate, sometimes sub-pectinate. Eyes emarginate.
Third joint of tarsi distinct, bilobed. Elytra not covering the extre-
mity of abdomen.
Sub-genera: Urodon Schoenh., Spermophagus Steven, Ca/rpo-
phagus Mac Leat. Oomp. Schoenh. i. pp. 31 — 114, v. pp. 1 — 147.
BruchuB piti h., Schoenh. ; Bruchua rtfimanus Schoknb., Pakzbb
DeuUcM. Ina, Heft 66, Tab. 14, &c.
Compare Hxbbioh-Sors7FKB AtueinanderaeUimg der Qattung Bruchua,
DetOachl, Ina, Heft 173.
C. Heteromera. Tarsi of four anterior feet five-jointed, of
posterior feet with four joints.
t Atrachelia,
Head sub-oval, posteriorly retractile into the thorax, towards
the base not abruptly nairowed, or without neck.
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490 CLASS Till.
Family LII. ^enelytra. Body with very few exceptions
winged. MaxillaB with internal process mostly unarmed. Mandibles
terminated by apex simple in some, in others bifid. Antennae fili-
form, in many longer than thorax and head together, in few only
sub-perfoliate. Penultimate joint of tarsi often bilobed.
Nao'Tow-ahielded. This denomination is not appropriate for all
the species, and this family of Latreille, in &ct, is neither very
natural nor sharply defined ; we retain it only because unwilling to
multiply too largely the number of fiunilies in our concise revision.
The larv» have six feet, but differ in other respects in the different
genera of this division. They live under the bark of trees or in
decaying wood.
Phalanx I. Rhynchostomata Latr., SdlpingidcB Leach, West-
WOOD. Head rostrate anteriorly.
Bhtnosimua Latr. {Salpingtta Illig., Anthribus Claiky., species
of Anthribtis Fabr.). Antennae moniliform, with three or five termi-
nal joints thicker, forming an elongate club. Thorax posteriorly
constricted, narrower than the base of elytra.
Sp. ShinontMU plardrostrii, AnihrQnu planirottrU Fabb., Pakzbb I>eui9ehL
Ina. Heft 15, Tab. 14.
Mycterua Clairv. Antennae filiform not clavate, with twelve
short joints. Thorax anteriorly narrower, posteriorly broader.
Sp. Mydenu curcuUoUHet, Bkinomacer cwrctdicides Fabb., Panzkb XhtOackl.
Itu. Heft I a, T&b. 8, Claibvilli ErUom. helv. i. Tab. xvl p. 134,
DuMiBiL C(m$, gH. $. I. Ina, PL 16, fig. 2, &c.
Gomp. OD this genua Olivieb Encyd. mith,, Int. Tom. vm. pp. 79, 80.
Stenostoma Latr, Antennae filiform, with joints elongate.
Body elongate. Thorax broader posteriorly.
Sp. SUnottofna rostratium Ohabpbnt., Leptwra rotlrata Fabb., Ga^Biir /con.,
In$. PL 33, fig. 8, &c.
Phalanx II. (Edemerites Latr. Head not produced into a
rostrom. Mandibles bifid. Posterior femora in males of many
species incrassated.
(Edemera Oliv., Latr, Antennae inserted between the eyes,
filiform or setaceous, sometimes serrate, mostly elongate, eleven-
jointed {oi males sometimes twelve-jointed). Eyes globose or reni-
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INSECTTA, 491
form. Last joint of mazillarj palps often large, triangular. Penul-
timate joint of tarsi bifid. Body mostly narrow, elongate. Eljtra
often soft and flexible, in many posteriorly aciiminate.
A) AfUermas short, nearlff equalling half the body. Claws of
tarsi bificL
Sub-genus Nothus Ziegleb, Out.
Comp. OuviBB Encyclop, mitk., EUt. not. Int. Tom. vm. pp. 583, 385.
B) Antenna elongate {almost of the length of body). Claws of
toflrsi entire.
Sub-genera : Calop%u ¥abbl, Sparedrus Megeble, Dytilus Fison.,
Nacerdes Stevek, (Edemera Oliy. (Species oiNecyddlis L., Fabb.,
Dryops Fabb.)
8p. tEdemara tanguinteolUi, Necydalia »angtdinecUi9 Fabb., Pakzbb
J>euUehl. Int. Heft 44, Tab. 18, named Necyd. JtaviooHit ,^^(Edem. notata,
Neeydalit noUUa Fabb., Naeerdet noiata Stiv., D^., Ac
Caleput terraticomit Fabb., Oerambyx terratieomit L. ; Duic^. C<mt.
gin. 1. 1. Int. PI. 12, fig. 4, Gu^BiK Iconogr., Int. PI. 33, ^. 5, Pakzkb
Deuttehl. Int. Heft 3, Tab. 15 ; habitat north of Europe.
Ck>mp. Olivibb Enq/d. mdh. 1. 1. pp. 437 — 450, and W. SoHmoT
Hevition der europaUchen (Edemeriden ; Liimaa entomciogiea I. 1846,
pp. I — 146, where alao some new genera are proposed.
Phalanx in. Serropalptdes Latb. {MelandryadoB Leach).
Head not produced into a rostrum. Mandibles emarginate or bifid
at the extremity. Posterior femora not incrassated. Claws of tarsi
simple. Penultimate joint of tarsi mostly bilobed. Maxillary palps
often serrate, with last joint large, securiform.
Serropalpus Hellenius, Payk.
Melandrya Fabr.
Sp. Melandrya terrata Fabb., Chrgtofnela caraboUUt L., Pakzbb DeuUchl.
Int. Heft 9, Tab. 3, CuY. R. Ani. id. ill., Int. PL 53, fig. i, Dum^bil
Cant. gin. 1. 1. Int. PL I3, fig. 3 T
Dtrcwa Fabr.
Add genera Conopalpus Gtll., ffypulus Payk., ffcdhmeivus
Payk., and some others.
Phalanx IV. Ctsteltdes Latr. {Cistelidce Kirby, Westw.,
Xystrojndes Solier). Head scarcely or not at all produced into a
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492 CLASS vjii.
roBtmin. Antenno at the base not obtect bj the margin of head.
Claws of taisi denticulate. Penultimate joint of tarsi mostly
entire, in few bilobed.
The luT» lire for the most pMi in hoUow trunks of treei. The perfect
insects sre mostly seen on flowers end hedges. Many of them have soft,
thin elytnL
Gomp. on this group of beetles SoLZSB Prodrome de la FamilU da
Xytl/ropidei, Ann^dela See. BiUom, iv. 1835, pp. 129 — 348.
a) MaxiUart/ palps with UutjowU large^ Hcwriform. {Mandibles
often bifid or tridentaU at the apex,')
Mtfcetochares Latb. {Mycetophtla Gtllekh., D^jsan, species
of Cistela Fabr.).
Sp. Myedoekatrts hairbata, Mdandrfa barbaia Fabb., Stdsm DeuUdU.
FawM,I%». u. Tab. 5a, Pavibb Jkutschl. /iw. Heft 105, Tsb. 5.
AUecula Fabs.
b) Maxillary palps filiformj incrassated towa/rds the apex,
{Mandibles entire.)
Cistela Fabr. (in part), Ctentopus Solier (and Megischia
ejusd.). Penultimate joint of tarsi entire. Head protracted before
the eyes,
Sp. Cidda mdfhvrta Fabb., Chrywmda Svlpkwrea L., Panzbb BetOacU.
Ina, Heft 106, Tab. 8, C5uv. R. Ani, 6d, iU., Ins. PL 5a, fig. 5 ; 4*^ long ;
the whole body, the elytra and feet sulphor-yellow ; black eyes ; in the
middle of summer common on umbellate and other flowers.
Phalanx V. HehpiiljArK, {Heioptdce Stephens, Westwood).
Head not produced into a rostrum. Antennae inserted before the
eyes under the exsert margin of head, covered at the base. Claws
of tarsi entire. Palps with last joint large, truncated. Penultimate
joint of tarsi mostly entire or not deeply bilobed.
a) Frostemum not produced posteriorly to form a pointed
projection,
* Body ehngatCy narrow.
Strongylium KiRBY (with addition of genus Stenochia ejusd.)
Stenotrackdus Latr.
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. INSECTA« 493
* * Body avalf chUmg,
Lcena Megerle, Dej. Thorax oboordato-tnmcate. AntennsB
filiform, incrassated towards the apex.
8p. Lcma pimdia, Hdops pmdia Fabb., Cuy. J2. AfU, 4d, ill., Jnt, PL 51,
fig. 10.
Helops Fabb. (in part). Thorax sub-quadrate, wings small,
imperfect.
Sp. H4op9 earabcXdeB Pakzbb, DetOkhl, Jnt. Heft 94, Tab. 3, Stubm
DevUchl. Fanunoy Int. n. Tab. 50, &e.
* * * Bodff oval, cocked above.
Spheniscua KiRBT. Antennas thicker towards the extremity,
with five last joints broader, forming a depressed club. Thorax
trapeziform. Body winged.
Sp. 8pkmi9cu» enOjfUadet Kibbt, Tram, qf the Linn. 8oe. zn. ^7, 4,
Cenlurie tPIntectet (Fr. edit Paris, 1834, 8to.) p. 47, PI. 3« fig. 4 ; from
South America, like the remaining spedea.
Adelium EliRBT. Anteimse filiform. Thorax short, broad.
Body apterous.
Sp. Addmm eakmmoidtt Kibbt, ibid. zxn. 4, Ceniwrie p. 45, PI. 3, ^, 3,
from N. HoUand.
Note. — Add genus Sphcerohu Kibbt, Aecmthopua Megkrue, Dij.,
and some others, on which consult Latreille, Cuyier E. Anu ed. 2,
nr. p. 38.
b) ProsUmtim produced pogterioriy to a pointy to he received in a
cavity 0/ meeoetemum.
Cnodalon Latr.
I^ntroffue Latr*
Compare Latb. Om, Onut. «t Tnted. n. pp. i8s, 183. Add genera
Caimptia and Camaria, Lspblbtikb et Sbbtillb, Enc. nUth., BitL nal.
Int. Tome x. pp. 454—456.
Family LIII. Taaicamea Latr. {Diaperidce Leach, West-
wood). Body winged. Maxilla on the inside mostly not unguicu-
late. Antennas not longer than head and thorax together, towards
the apex incrassated or terminated by a club, moniliform or perfo-
liate, mostly inserted under the exsert margin of head. Tarsi with
joints entire, terminated by claws simple.
Phalanx I. Cossyphtdes Westwood. Head immersed in
thorax or obtect. Maxillary palps with last joint broad, securi*
form. Elytra marginate.
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494 CLASS YIII.
Nilio Latr. Head small, received in thorax. Thorax broad,
very short. Body semiglobose. Antennas incrassated towards the
extremity.
Sp. NUio vUlows "Latr., jBgithtu marginatut Fabb., Latb., Otner. OrvM, a
/m. Tkb. X. fig. a. Guv. Jt. Ant, id. ilL, Ins. PI. 5, fig. 10 ; South
America. Thin insoct in ite Sdrb v^^nh^'^ OoccineUa,
Heleus Latr., Kirby. Head margined by thorax produced
forwards. Body plane. Antennas incrassated towards the ex-
tremity.
HdceuB jperfcnOus Latb., Guv. R. Ami id. L Tab. xn. fig. 6, id. n.
Tab. xvu. fig. 6 ; New Holland.
OosBypTma Oliv., Fabr. Head covered by clypeiform thorax.
Antenna terminated by a club perfoliate.
Sp. Costyphfu depreasui Fabb. (exd. Synon.), Oouyph. HofmafiiMffgU Hkbbbt,
DumiSbil Cims. ffhi. 1. 1. Im. PL 15, fig. 8, Guv. B. Ani. id. ill.. Int.
PI. 50, fig. 8; in Portugal and in North Afirica, fta Gomp. Sohobnhxbb
Synon. In$. i. 3, pp. 58, 59 ; — Con. tavricua FiscH., BuUeL delaSoe. Imp.
de Motcou, 1832, Tom. iv. Tab. v. t 1, a.
Phalanx II. Diaperiales. Head not received in thorax or
obtect. (Maxillary palps mostly filiform. Body immarginate.)
ElecUma Latb. (BolUophagus Fabb.)
Coxelus ZiEGL., Dej.
Lithophilus Fboelich.
Sp. Lithophilus ruficoUia J. A. Fboelioh Neue CkiUungen wad Arien von
KSfer, Naturforichar a8 St., 1799, Tab. i. fig. 11; Latbeillb placed this
genus with Coecinella.
Trachyacelts Latr.
Hypophlceus Fabr.
Sp. ffypophUffus eattanetu Fabb., Panzeb DeuUchl. Ina. Heft 11, Tab. 13,
DuMl^. Com. gin. ». I. Int. PI. 15, fig. 1.
Note. — Here ought, as it seems, to be inserted genus Nycteropus
Klug (with genus Dclichoderua ejusd) ascribed by GuiRiN to the
Helopii.
Gu^BiN Magcuin de Zocl. 1839, -^^^ ^^* ^03.
Dtaperia Geofpr. Maxillary palps filiform. Internal process
of maxilla unarmed. Antennae with fourth joint and following
transverse, dilated.
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IN8ECTA 496
Sp. ZHaperit MeH, Chrymmda hoUti L., Gioitb. HiM. d^Tnt. i. PL vi. fig. 3,
DuM^iL Cbn*. ghi. «. I Ins, PI 15, fig. 5 ; 3'" long, a"' broad, black, the
elytra with two orange transverBe bands, and an orange spot at the point.
Sub-genus : NeonUda Ziegl., Opiocephala Lapobte and Bbx7LIi£
Compare Db la Pobtb et BbuliJ, Monographie du gmre Diaperit,
Ann. det Se. not, Tom. xtttt. 1831, pp. ^2$ — ^410, PI. 10.
Add sub-genera^ in which the last joint of maxillaiy palps is
broad, truncated Platydema Lap. and B&y Ceropria and some
others! L
Fhaleria Latr. {Uhma, Phaleria Dej.). Internal process of
maxilla armed with a homy claw. Maxillary palps with last joint
incrassate, obtrigonal, truncated.
Sp. Phaleria eadaterina, Tenebrio eadavgriMti Fabb., Stubm Detdickl,
Fauna, Int. n. Tab. 47, fig. a, a, b, Guy. J2. Ani. Id, UL, Int. PI. 50,
fig. I, &e.
Famfly LTV. Melasomata Latr. Body often apterons. An-
tennae scarcely longer than head and thorax conjointly, inserted
under the exsert margin of head, mostly short, moniliform, with
second joint short, third in most elongate. Litemal lobe of maxilla
almost always armed with a homy tooth or claw. Mandibles bifid
or emarginate at the apex.
Most of these beetles have a blade colour ; others are brown or
grey, without contrasting spots ; they are shy of the light and
live abnoflt all on the ground, where they conceal themselves
in corners or xmder stones. The shield-covers of the wingless
species are grown together, and at the sides bent downwards.
Gompare on this fionily GuiBiN, MaUriaMX powr vine dautficatum de$
Mdaaomes, Magas. deZool. 1834, Ina. Pi loi — it 8.
Phalanx I. Tendn-tonita {TenebrtantdcB Leach, Westwood).
Body winged ; elytra loose. Maxillary palps with last joint broad,
truncated.
A) Body dong<Ue. Thorax sub-quadrale, AtUennoB momXliformy
graduaUy thicker towa/rda the apex.
Heteroiarstis Latr. Antepenultimate joint of tarsi deeply bilobed,
receiving the penultimate very small.
Sp. ffderotamu ienSMMet Latb., Guv. R. Ani. id. ill., Ina. PL 49, fig. 1 1,
from SenegaL
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496 CLASS VIII.
Tenebrio L. (in part). Pennltiinate joint of tarsi distinct, not
concealed in preceding.
Terubrio Fasbl, Upia Fabb., CcUcar Dij. (species of TroffOtUa
Fabr.)
Sp. Tendmc moliiar L., Panzkb Deutschl. Ins. Heft 43, Tab. 11, Stubm
DeuUchl. Fauna, Ins. n. Tkb. 46, DuifiBiL Cons, gSn, s. L Ins. PL 13,
^. 1 ; the meal-beetle ; abore f" long, black, brown beneath, elytra
striated. The larva (meal-worm), well-known in com- mills, granaries, and
bread-bakers' establishments, is elongate, cylindrical, white, and has twelve
rings besides the head.
B) Body dangate. Thorcw sub-quadrcUe, elangcUe. Antenma
davate or thick at the apeaa, fadform.
Toxtcum Latb. Antennae with four terminal joints forming a
club, last sub-orbicular, rest transverse. Tibias simple.
Add genus Boroa Hebbst.
Chiroscelia Lam., Latb. Antenna with last joint globose,
thick. Tibiae of anterior feet dilated, digitate. Thorax elongate,
posteriorly constricted, narrower than the base of elytra.
Sp. Ohirosedis lifenestrfOus, Lamabok Awnal, du Mushum in, 1804, p. 263,
963, PL 49, fig. 2 ; Gu^KiN leonogr., Ins, PL 30, fig. 5.
Sarrotrium Illig., Fabb. {Orthocerua Latb.), Antennae fusi-
form, thick, pilose.
Sp. Sarrcitrwm mwUewn, Hitpa mMtiea L., Pavzxb i^utodU. Ins. Heft i.
Tab. 8, DuMfBiL Cons. gin. s, I. Ins. PL 13, fig. 5 ; found in sand-pits.
CorUeus D^. Sp. Sarrotrium edUs Gsrm.
C) Body oval. Thorax transverse, trapemHdaly anteriorly nor-
rower y ^posteriorly of the breadth of elytra.
OpaJbrwin Fabb. (exclusive of many species). Clypeus emargi-
nate, including labrum. Antennae shorter than thorax* Thorax
of the breath of elytra, marginate.
Sp. Qpafnisi Mdvlofum Fabb., SSI/pha, saMosa L., Pahkkb DeutsdO, Ins.
Heft 3, Tab. 1, Stubm DeutschL Fawna, Ins. n. Tab. 40, Cur. J2. Ant,
id. ill., Ins. PL 49, fig. a^',^OpaUum tibiale Fabb. (IficrtwHm D£r.),
Pavzib Dtuischl. Ins. Heft 43, Tab. 10.
^ Of this species, as also of Sa/rrotrium muiieum, SoHlODTi has oommnnicated an
anatomical description in Kb5txb's Natuwrk. Tidskr. nr. 1843, pp. '904 — 9 11.
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INSECTA, 497
Crypttcus Latr. Body oval, convex, smooth. Clypeus not
emarginate. Antennas of the length of thorax. Tarsi with first
joint elongate.
Sp. Cryjiicui glaber Latb., Blapa glabra Fabb.; Cuv. J2. Ani. id. iU.
PL 49, fig. I ; the larva, which is met with in decayed willow-wood, is
filiform, cylindrical, and resembles that of ffdops BouoH^ Naiurgetch. d.
Int, s. 191.
Phalanx II. Blapsida {Blapstdcp Stephens, Westwood).
Body apterous, with elytra united together. Maxillary palps with
last joint larger, securiform or obtrigonal. (Gknus Blaps Fabb.)
A. Body oval, short. Thorax tranwerse, posteriorly broader, trapezoidal.
Elytra wUk tides Utile or not at aU it^/leeted under abdotnen. Tarsi of a/nU-
riorfeet in males dilated, often hirsute beneath.
Pedimts Latr.
t With clypeus entire : Sub-genus Platyseelis Latbbillb.
t+ WUh clypeus emarginate: Sub-genera Pedinus D^., Isoeerus
MsaBBLi, Pedonoctu Watebhoubb, Ewrynotus Kibbt, HdiophUus Dj^.,
Dendarus Mboeblb, Dj^. Compare Latbeillb in Cuv. R. Ani. sec. id.
▼. pp. 19, 70, Watebhoubb Arm. and Mag. of Nat. Hitt. xvi. pp.
Note. — Cokoptera allied to genus Opabrum, but apterous.
B. Body oblong. Elytra at the sides inflected downwards, posteriorly
often produced into an v/^fiected pointed process,
Blaps Fabr. (in part).
Sub-genera: Asida Latb., Scotinua Kibbt, MacMa Hebbst,
Beteroseelia Latb.
Gonopua Latb.
Blapa {Blaps and DUa FiscH.), Prosodes Eschsch.
Comp. FisoHEB BuUet. de la Soe. des notitwral. de Moscou, Tom. xvn.
1B44, PP* III — 118, Tab. n, Sp. Blaps kevicollis Gbbl.
Misclampus Latb., Hdiqfugus GuiBiN, Acomthomera Latb.,
Oxfwra Kibbt.
Sp. Blaps mortisaga Fabb., Tendnio mortisagus L., Houttutn Nat. Mist.
I. 10, Tab. 78, fig. 3, Panzbb Ikutschl. Ins. Heft 3, Tab. 3 ; 10'" or 11'"
long, dull black ; lives in cellars and other dark places. Some anatomical
notices on this species and on Blaps gages Fabb. (Pavzbb Deulschl. Ins.
Heft 96, Tab. i, Dum^bil Cotis. gin. s. I. Ins. PL 14, fig. i) were given
by J. F. Mbokil Beitrage z. vergl. Anat. i. 2. s. 123 — 136.
VOL, I. 32
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498 CLASS viii.
Phalanx III. Fimelarta. Body apterous, with elytra united
together. Maxillary palps filiform, with last joint scarcely larger
than the rest, subcylindrical.
A. Mentum aubguadraU, enHre, narrower at the hate.
Molurts Latr.
Sp. Pimelia striata Fabb., Yost Coleopt, ii. Tab. 50, fig. i.
Trachynotua Latr.
Septdmm Fabr.
8c<mru8 Fabr.
Tagenta Latr,
Note. — Several sab-genera of modem writers to be added, which are here
omitted for the sake of brevity. These are ooiinted amongst the BlapndeB
by the celebrated entomolo^t Wbstwood.
B. Menium very hroad, mostly not narrowed at the hate, covering maxtUa
beneath, aiUeriorly emarginaJtt or hiiobed, cordate,
t MeiUum anteriorly concaive, oordate. Antenna with only ten eUtHnet
joints.
Adehstoma DuP.
Eurychora Thunb., Fabr.
Sp. Eurychora eUiala Thukb., Yobt Coleopt. n. Tab. 43, fig. 23, Duic^ril
Oons, gin, s, I, Ins, PL 14, fig. 3 ; — Etirych, cimicoides QuKtf SEL, Sohoebh.
Syn, Ins. I. p. 137, Tab. n. fig. 5, Ac.
Steira Westw.
Bp. Steira costaia Wbstwood in Gu^bin Magas. de Zool. 1837, Ins, PI. 176.
ft MenUum bilobed, rotundate, AntenncB with deven joints.
Akis Herbst, Fabr. (exclusive of some species).
Tentyrta Latr.
Hegeter Latr.
Erodvus Fabr.
Sub-genera : Nyctelia, Zophoais, Erodiua Latr
NoU. — ^Watbbhoubb has described several new species of NydeUa, Pro-
ceedings of Zool. Soc, IX. 1841, pp. 105 — III.
Pimelia Fabr. (in part).
Sub-genera : OryptochiluSy Trachyderma Latb., and some others
here omitted.
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INSECTA. 499
Sp. Pimdia hipunetata Fabb., Dum^il Cons, gin, a. L Ins. PL 14, fig. 9,
Guy. R. Ani, id. ill., Ins. PI. 46, fig. i, &c. ; Tenebrio muricaUu L., a
congeneric species, is often confounded with the preceding. See Schoen-
HBBB Synon. Ins. I. p. 131.
^ote. — There is an affinity between this &mily and the ccbrdbici :
some genera agree in hahitus; thus genus Damcuter Kollab
amongst the CarcUnci almost presents the form of the Blaptidea,
but Mcmticara resembles Pimelia caronata. The anatomy also
proves this affinity, as Blanchard has lately shewn by the investi-
gation of the nervous system. Ann, des Sc Nat. 3idme S6rie v.
Zoologie, p. 350. The larvse, however, are altogether different.
tt Trachelia.
Head cordate, abruptly narrowed posteriorly and separated
from thorax by a stricture or neck.
Family LV. Gantharidia (Trachelides Latr., Cantharidi^
except Cistelidesy Blanch.).
Phalanx I. Meloeida {Canthartdtw Latr.). Antennae monili-
form. Head nutant. Maxillary palps filiform, with last joint
scarcely thicker, oval or obovate; labial palps with last joint
broader, truncated. Elytra soft. Joints of tarsi mostly entire.
Claws of tarsi bifid. (Genus Meloe L.)
t Wings none; dytra {at least in females, mostly also in males)
dbbrwiate.
Melof^ Fabr. Antennsa with eleven joints, filiform, sometimes
thicker in the middle, in few subincrassated at the apex.
Sp. Mdo€ proicarabceus L., Geoffbot ffist. des Ins. des env. de PariSf I.
Tab. VII. fig. IV., Bbandt u. Batzbbubo Med. Zool. n. Tab. xvi. figs.
4, 5 (Meloe tecta Hxllw., Panzeb DevUehl. Ins, Heft 10, Tab. 14, male,
Meloe punctidata Pavzbb, ibid. Tab. x6, male); black, commonly glancing
into steel-ooloured yiolet, with deeply impressed points on the head and
thorax; this species differs much in size. — Comp. on this genus Lbaoh
British species of the genus Mdoi, Tram, of the Linn. Soc. Vol. xi. 1815,
PP* 35—48, Tab. V. VI., and his Pwrther Observat. with the descr. of
six exotic species, ibid. pp. 242 — 151, Tab. xvin.; J. F. Bbakdt et W.
F. Ebiohson Monographia gener, Mel,, Nov, Act. Acad. Cobs. Leop. Car.
XVI. P. I. pp. loi — 147, Tab. vin. — ^Nbwpobt Nat, Hist, Anat. and
Developm. of the Oil-beetle, Trans. Linn, Soc. VoL XX. p. 197, Vol. xxi. P. 2,
p. 167..
ft Body winged. Ulytra not shorter them abdomen.
32—2
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500 CLASS VIII.
a) AntemuB with eleven jaifUs,
Lytta Fabr., Cantharts Geopfb,, Oliv., Latr, Antenna
straight, filiform, longer than head and thorax. Body oblong.
a) With penuUtjnate joint of torn enHrt.
*) With dytra (U the apex attenuaUd, tubulate, dehiscent,
SUa/ris Latb. {Apcdus Fabb. ^)
* *) With elytra covering the whole abdomen above, not aiUenuaied.
Sub-genera : Gnathium Kibbt, NemogncUha Illio., Latb., (with
external process of Tnaxillffi filiform, exsert), Zonitis Fabb., Tmesir
dera Westw., CarUharia Latb.
Sp. Ljftta veaicaUyria, Meloi vesicatorius It., GiOFFB. Ins, I. PL yl fig. v.,
DuM^BiL Cone. gin. $. I. Ins. PL lo, fig. 6, Bbandt u. Ratzbbubo
Mediz. Zocl. n. Tab. xvm. figs. 1—6, &c. Spamth Fly; green gold-
glancing, with black antennsB, a broad head, channelled above, thorax
quadrangular, broader forwards, length 10"'. The blistering power of this
insect renders it very important as a remedy in medicine. Other species
also of the Mdoiida possess the same property, which however, as is
asserted, is not perceived in Apalus. The female lays her eggs in the
ground ; the larvae make their appearance after about three weeks, like
those of Melog (and of Apalus, which are quite similar to them according
to GbniQ, hitherto known in the first period of their life alone, but not in
their further growth.
Compare C. A. Lbnaub, Meloi vesicatoriut, Diet., in C. LnrKin
AffuxnU. Acad. vi. pp. 132 — 147 ; Lobchob Beitrage zw Oet<^. cisr Span'
iachenFliege, Naturfortcher xJLiu. 1788, s. 37 — 48, Tab. i, figs, i — 8;~
Audouin Becherchet pour tervir d VHittoire not. det Caniharide*, Ann, det
8e. natur. ix. 1S26, pp. 31 — 61, PL 41, 43 (anatomical description) ;
Brakdt u. Ratzsbubo L L pp. 116 — 121 and PL xix ; Ratzebubo
Fortt'Int. X I. pp. 88—^2 &c.
P) With penultimate joint of tarti hilobed.
Tetraonyx Latb.
Comp. Latbbillb in Huvboldt et Bonpland Obterv. de ZooUtg. et
d^Anat. Ck>mp. L pp. 160 — 16«, PL xvi. fig. 7.
CEnas Latb. Antennas not longer than thorax or scarcely as
long, geniculate, with first joint elongate, obconical, the rest broad,
short, the last oval.
Bp. (Enat afer, Lytta afra Fabb., Latb. Oener. Cruet, et Int, Tab. x. fig. 10^
Cuv. R. Ani. id. ill.. Int. PL 54, fig. 9.
^ J. GBNi, MivMMre pour tervir d CHitt. not. de VApalue bimactdatut et det
Cantharidet en giniroL Ann, det 8e, not, zxiii. pp. 138 — 144. 1831.
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INSECTA. 501
Mylabris Fabb. AhtennsB short or scarcely of the length of
thorax, towards the extremity incrassated gradually into an elon-
gate club, arcuate.
Compare G. J. BiLLBBsa MofM^ra'phiok MyldbridAim, HoImi», 1813, 8fo \
SoHOiNHXBB Syn. Int, i. 3, pp. 30—43 ; F. Gsblxb Dei Mylabrides de
la Sib^rie &o. Nouveamx MSm, de la 80c, itnp^, de Motcou, I. 1829, pp. 147
—171.
Sp. Mylabrie cichorii Fabb., Mdoi Oiehorii L., Bbandt il Katzeb.
Mediz, ZooL n. Tab. ZYni. fig. 17, from China, East Indies. Other
species are often confounded with this, as Myldbris FuesUini Pahzxb
DeutechZ, Ins. Heft 31, Tab. 18, Mylabris voflriabUU Billb. &c
I/ydu» Mbgbblb (with external claws of tarsi pectinate). Sp. Mylab.
trimaculata Billb., Brandt u. Batzeb. Med. Zool, n. pp. 136, 127, Tab.
zym. fig. 16.
b) ArUemuB mith eight or nme joints, cHavate at apex, last joint
kurge, oval.
Hycleus Latb.
ArUhmema Oheyboi*
Sp. Arithtnema decemgiEUaia Chxvb., Gu^ein leonogr., Ins. PL 35, fig. 4,
pp. 131, 133.
Ceroooma Geoffr., ScH-fiFF,
Sp. Cerocoma Schafferi Fabb., Meloi Schc^eri "L,, Gboftb. Ins., I. Fl. 6,
fig. 9, SoHJSFTXB Mem. EnUm. Tab. 37, Dumi^bil Cons. gSn. s. L Ins, PL
10, ^. 7, Ac.
Phalanx 11. MordeUona {Mordellidce Leach). Antennae short,
very often serrated. Elytra hardish. Thorax gibbous. Body com-
pressed. All the tarsi or those of posterior feet with joints entire.
Claws of tarsi mostly bifid, with the superior divisions pectinate.
(Gknus Morddla L.)
Rkipiphorus Bosc, Fabr., Latr. Thorax produced posteriorly
between elytra ; scutellum concealed. Elytra abbreviate, or angus-
tato-acuminate, dehiscent. Antennae pectinato-serrate, in males
flabelliform. Palps subfiliform. Maxillae with external lacinia
long, linear.
Sp. Rkipiphorus paradoxus Fabb., Morddla paradoxa L., Panzkb D, Ins.
16, Tab. T4; — Bhipiphorus himaeulatvs Fabb., Dum^b. Cans, gin, s, I,
Ins. PL 1 1, fig. 4, &c. Comp. on some new species Fibcheb in BtUlet. de
la 8oe, impir. des Natwral. de Moscou, iv. 1831, p. 426, Tab. v.
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502 CLASS VIll.
Sub-genera : Myoditea liATR,, Ripidiua Thunb. {Sywbius Sundev.),
Macnmagon Hentz, Pelocotoma Fisch., Evamoeera Gu£bin (with
scutellum distinct).
Gomp. Hentz Trans, of the Amerie(m philos, 8oe., new Seriee, YoL m.
Pt. 2, pp. 463, 463. PI. XY. f. 3. Maerotiagon dimidi4aum {Jthipipkorus
dimidiatua Fabb.), Sundeval in Okbn's Ins, 183 1, s. J122 — 1328, Tab.
ym. Symhius Blattarum. (The larva lives in BlattcB after the manner of
Ichneumons.)
MordeUa Latr. Scutellum distinct. Elytra narrowed pos-
teriorly, not shorter than abdomen. Antennas filiform, subserrate.
Maxillary palps with last joint large, obconical.
^. Morddla aculeata L., Gkoffb. Ins, I. PL 6, fig. 7, Sohjeffeb Elem,
EnUm. Tab. 84, &c.
Anaapis Geoppr., Latr. Antennas thicker towards apex, sub-
filiform, not serrate. Penultimate joint of tarsi of four first feet
bilobed. Bemaining characters those of the preceding genus.
Sp. Anaapis frontalis, MordeUa frontalis L., Panzeb Deutschl, Ins. Heft
13, Tab. 13; variety Anaspis atra Dum&IL Oons. gin, s, I, Ins. PL 11,
fig. 6, Bm.
Phalanx III. Horialia {HoriidcB Westwood). Antennae fili-
form, moniliform or subserrate, short. Tarsi with joints undivided,
furnished with claws denticulate beneath, and with double filiform
appendage. Mandibles strong, exsert; palps filiform. Thorax
broad, transverso-quadrate. Head nutant. Feet short, posterior
with femora incrassated, especially in males. Elytra soft, dehiscent
at the apex.
Horia Fabr.
* Head WMrower than thorax. Antennas Umger iham iSumtet. Sub-genus
Cissites Latb. {Oener. Crust, et Ins. 2, p. a 13, previously Horia ejusd.
Hist, not, des Crust, et Ins. x. p. 364).
Sp. Horia testaeea Fabb., Dum^bil Cons. g$n, s, I. Ins, PI. 12, fig. 6, J. O.
Hubbkeb Beitr. z. Natturg. d. Ins. in NaturforscKer aS, p. 47, Tab. n.
figs. 14 — I •J, from East Indies.
** Head tumid hehind the eyes, of ihe breadth of thorax. Antenna
scarcely longer than thorax. Horia Latb. (previously Cissites ejusd.)
Sp. Horia maculata Fabr,, Cuv. R Ani. ed. HI,, Ins. PL 54, fig. 4, firom
S. Amer. The larva lives parasitically in the cells of the larvte of Xylo-
copa; on the metamorphosis cons. Lansdown Guildiho Trans, of the
Linn. Soc. Vol. xiv.
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INSBCTA. 603
Phalanx IV. P^ochrdidea nob. Antennae filifonn or gradu-
ally thicker towards the apex, subserrate, in some pectinate, longer
than head and thorax. Tarsi with penultimate joint bilobed, with
claws undivided, not denticulate. Maxillary palps with last joint
large, truncated, obtrigonal. Mandibles broad, short.
We unite in this group the Anthicides (NoiooddoR Stephens,
Westw.), the Pyrochrcndea and LagriarioB of Latbezlle, which are
all closely allied.
Notoxus Geoffb., Oliv. {ArUhicua Fabb.). Antennae filiform
with joints obconical, the last oval, longer. Maxillaiy palps with
last joint large, securiform. Head nutant, broader than thorax.
Thorax elongate^ broader anteriorly, often constricted in the
middle.
a) WUk thorax anteriorly porreet into a horn, {Notoxut Illio., ManO'
cerut Megeble, D&r.)
Sp. Notoxus manoderoSf Mdoi monooeroi 1,., Geoffb. Im, i. PL 6, fig. 8,
DuM^. Com. g6n. 9, 1, Ins. PL lo, fig. 5, Panzeb D, Ins. Heft 26, Tab.
b) WUh thorax unarmed.
Sp. Notoxus antherirvus, Mdoe antherinus It., Pakzbb D. Ins. Heft 1 1, Tab.
14, &c
Steropes Stev.
Scraptia Lats.
Note. — ^A genua to be referred perhaps to the Serropalpides (v. supra^
p. 491) : this u scarcely its place.
Pyrochroa Geoffr. Antennae rather long, in males serrate or
pectinate. Body elongate, narrower anteriorly, with thorax subor-
bicular or trapezoidal. Maxillary palps subserrate, terminated by
a longer obtrigonal joint ; labial palps filiform. Abdomen rounded
posteriorly.
Fogonoeerus Pisch. {DendrcHdes Latb.)
Fyroch/roa Latb.
Sp. Pyroekroa rv^bens ¥asbl., Lampyris rvbtns Sohalleb, Gmbl., Pahzeb
D. Ins. Heft 95, Tab. 5, Vobt Cdeopt. 11. Tab. 48, fig. i ; the body and
thorax black beneath, the head, the upper surface of the thorax, and the
elytra vermilion without spot ; the perfect insect is found on flowers ; the
larva lives in decayed willow- wood.
Note. — ^Here should be referred, according to Westwood, genus
Pyiho Latb., with body depressed^ thorax narrowed posteriorly,
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504 CLASS VIII.
truncated, toUh pemiUimate jaiaU of tarsi eniMre, tinleas it be pre-
ferred to count it with Latbeillb amongst the ffdopu, and to place
it before Strongylvwm, (see above, p. 492).
Lagria Fabr. Antennee inserted in a sinus of the eyes, filiform
or gradually thicker towards the apex, with last joint longer than
the rest. Palps both maxillary and labial thicker towards the
extremity, maxillary longer than labial, with last joint transverse,
truncated. Thorax narrow, cylindrical, subquadrate. Elytra much
broader than thorax, dilated towards the apex, rounded, soft. Body
very often villose or pubescent.
Sp. Lagria hvrta Fabb., Ckrysomela hirta L., Paitzeb D, Ins. Heft 107,
Tab. 2, Cuvikb R, Ani, Sd. HI., Ira, PL 53 bis, fig. i; the Urvas fonnd
by Ltonbt in winter under fallen oak-leaves, has yeryoonspicuons i^nt-fltma^*
projecting laterally; se^ Ltonbt Rechiertkn, pp. iii — 114. PL XL figs.
17— 3».
Statyra Latr,
Species all American, similar in habit to the CoirMeif with thorax OTa^
truncated. Sp. Statyra agroldes D6j., Cdv. R. Ani, 4d. HI., Ins. PL 53
bis, fig. 3. On this genus oomp. Saint- Faboeau et Sebvillb ^tcyd.
mStk.g Ins. Tom. ^. pp. 479, 480 ; Di/BAN enumerates 22 species of this
genus ; CatcUogue des CoUopi. 1837, p. 336.
D. Pentamera. Tarsi all with five joints.
We here employ this character only in so &r as it is available for the most
of the genera^ but not as absolutely universal and exclusive. Ahready
Latbbillb was obliged to admit some exceptions {Families nai. du Rigne
animal, 1835, p. 337); we subjoin still others, and do not separate, for
instance, Psdaphus from the rest of the Braehdytra, whilst, on the other
hand, we have placed the genera with five joints i)acne, Epueapika, and
TriflalUma (see above, p. 470) with the Tebramera in the family of the
Clavipalpi.
Family LVI. LamtUioornia. Antennas clavate, shc^i;, mostly
with nine or ten joints. Inserted under the exsert margin of head.
Club of antennsB lamellate, or serrate, or tunicate, with first joint of
club infimdibuliform, including the other joints. External margin
of the two anterior feet dentate. Mentum often large, covering the
ligula, and bearing palps.
Lcvmellicom beetles. The insects of this group form in the
system of Link^us the genera LiLcamu and Scarabagus. The larvsB
are thick, cylindrical, the posterior part of the body bent down-
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INSECTA. 505
wards j besides the head there are thirteen rings; no eyes are
present ; the feet have four joints and a strong daw at the end.
The pupfB, which lie in a hollow dod of earth, have the cases of
the wings projecting beyond the posterior margin of those of the
elytra. See the beautiful figures in the Mem, 9ur lea metamorphoses
ds8 CoUopUrea of W. De Haak, iiTouv. Ann. du Muahim iv. 1835,
pp. 125—164, pL 10—19,
Phalanx I. Lucantdea or Priocera. Antennas with ten joints,
club pectinate, with lamellae sub-parallel, and almost perpendicular
to the axis. (Elytra always covering the apex of abdomen. Ab-
domen with only five distinct segments underneath.)
The larvBB live on decayed wood and reside in hollow stems of
trees. This group is allied indeed to the following, but still differs
by some anatomical characters. The larv» have a nervous ef3rstem,
of which the ganglia of the thorax and the abdomen are separated
fix)m each other by considerable distances ; whilst in the following
group they lie veiy dose together, forming as it were a varicose
string. In the proper genus Lticamis the nervous system of the
perfect insect has quite a different form fix)m that of the Sccurahceida ;
the second and third thoracic ganglia are separated from each other,
and the abdomen has six distinct ganglia, whilst in the Sccvrabceida
the second and third thoradc ganglia are fused together, and in the
abdomen, in place of a chain of gangUa, a single central nervous mass
alone succeeds to this thoradc ganglion, from which mass the nerves
of the abdomen arise at acute angles (like the last spinal nerves in
the Ccmda equina of mammals).
See tha figures of Blanohabd Ann, dea Sc. not., $iime SSrie, Tom. v.
PL 8, fig. X, in Liieantu cervtu, and in Guvdeb iZL An. id, ill., Ins, PL 3,
in the cockchafer. Aa perfect insect PoBtalut follows the Scardbceida in
the form of the nenrous system, and therefore is referred by Blanohabd to
this group, and separated firom Lucanus.
A. lAgula mmJbrantmt, mostly bUobed and penieiUate, Antennas smooth,
mostly broken, with first Joint elonffate. Seutdlum between the base of dy6ra
at the beginning of the svture.
Lacanus L. (in part). Mandibles exsert beyond head, mostly
much larger in males, porrect. Labrum mostly conjoined with
clypeus or none. Maxillsa membranous. Antennae broken. Feet
often elongate, especially the anterior. Body somewhat de-
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506 CLAJBS VIII.
The Stag-beetles. The genus Lucanua, although firom the
characters giyenlEtbove much more limited here than with Linkjeus,
still contains different genera of modem authors which it is
impossible for us to set forth here. The under jaws are commonly
membranous ; in the females alone of some exotic species the
innermost lobe (mando) has a homy hooklet at the extremity \ In
some species the eyes are completely divided by a projecting flat
margin of the head into two, one situated on the upper and one on
the under side of the head.
Corap. C. P. Thukbsbg Lucani Monograpkta, MSm, de la 8oe. impir,
de Moseou, i. pp. 150 — 173. PI. ii; Mao Lsat, ffora enlamol, 1819 {An-
nulosa Javanica, idii. franQ. Paris, 1833, 8vo. pp. ii — 29) ; J. O. W«ST-
WOOD Descripiio gener. iwoor, e fam. Lucanid., Ann. det Sc. not. le S^rie,
I. 1834, pp. 113— 127, PL 7; BusMKiSTEB Handb, der EnUm. v. 1847,
8. 311—442.
Liicanvs Latb. (and Flatycerua ejusd in part).
Sp. iMcanut eervut L., Stag-beetle, cerf volant, Hirtek-Jcafer, SchrOler, kel
vliegend heri, kc, RossL Int. n. Scar. terr. CL i. Tab. IT. y., YoBT
OoUopt. I. Tab. 99, figs, i, a, DuMifiiL Cone, gin. 9. L Ins. PL 5> fig. i ;
dark-brown (like old mahogany) ; the male is larger than the female, baa
the head broader than the thorax, and branched upper jaws about the
length of head and thorax together, these included it is fall 2^" long; in
the female the upper jaws are less than the head. Hie size, however, yarieB
much ; a constantly smaOer yariety is Lucavtve capreolue Fabb. (not L.),
YoKT, I. L figs. 3, 4, SuLZKB Abgek. Oeech. d, Ine. Tab. n. fig. i, which
is found with the ordinary one. {Lticanue capreolue L. is a N. American
species, which Fabbioiub names Lucanus dama.) The metamorphosis of
the stag-beetle is described by Rcbssl. The larva is pale yellow, becomes
very large, and lives some years; it lives in decaying oak-trees. The
perfect insect makes its appearance in June and July.
Chiaaognathtta SxEPHENa Club not abruptly distinct, with six
lamellae. (Eyes always divided into two parts by the margin of
head, as by a septum.)
Sub-genera: ChioMognatkue Stephens (antennsB of males with a whorl of
hairs before the club), Sphaerognathue BuQDST. Mandibles of males in both
these sub-genera longer than thorax, sometimes of the length of body. Sp.
Chiasogn. Orantii, Tetropkthalma Chtloineie, Lesson Illuet, de ZooL
PL 14, Diet. wniv. cTBiel. not., Ine. Coliopt. PL 7, fig. ^i—Sphwrogn.
prionoides Buquet, Gu^in Magas. de Zool. 1839, ^^- ^^ ^'t Sphcarogn,
Feiethamelii Gu^ein ibid. 1840, PI. 39. Species from S. America.
Ryseonotue Mao L. Mandibles thick, with external margin unidentate.
Sp. Luc. neMoeue Kibbt, from New Holland.
* Westwood, Annate of Nat. Hielory, viii. 184^, pp. 1 11— 143.
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INSECTA. 607
Lamprima Latb., Schoekh. Labnim difitinct, ezsert Club of
antennae with three lamellaB. Internal lacinia of mazilliB in females
terminate hj a homy hook. (Head small, narrower than thorax.)
Sub-genera: Lamprima Latb. (Sp. Lampr, fulgida, Lampr. cenea
GuiBiN Iconogr., Ins, PL a;, fig. 2, Cuv. H, Ani. ed. iU., Int. PL 45 bis,
fig. 3 ; (anterior iibiie of males with tenninal spine broad, obtrigonal) ; —
PhcUdctus Mao Lsat. (Sp. Lamprima Sumboldti Gilleith., Schokkh.
Synon. I. 3, Append, p. 197, Gn^IN Iwnogr,, In$, PL 47, fig. 6, &c.
Syndesus Mac L. Mandibles exsert beyond the head. Labrujn
connate with clypens. Maxillae membranous. Antennae short, not
broken, club with six or seven joints. Eyes large, globose,
undivided. Posterior feet very remote from preceding. Body
cylindrical.
Sp. Syndenu eormOua, Sinodendrum eomutum Fabb., Cxrv. R, Am, id, iU,,
Ins. PL 45 bis, fig. 8 ; habit, in Van Diemen't Land,
iVbte.— Genus HaoapkyUum Gbat, Wbstw. {PtUodon Pbbtt) is rightly
joined with genus Syndenu by Bubmbistbb.
Ftffulus Mac L. (and Ntgidius ejusd.). Mandibles little exsert.
Internal lacinia of maxillae armed with a homy tooth. Labrum
exsert, mobile. Antennae broken, club with three lamellae. Lateral
margin of head extending beyond the eyes, and dividing them in
great part or entirely.
Sp. Fiffultu ttriatug Wbstwood, Blanch abd Bia. d. Ins, Paris, 1845,
PL 8, figs. I — 3 (imago, larva, nymph) ; hab. in island of Mauritius, &c.
NcU. — ^Add sub-genera Xiphodontus Wiarw. and Agnus Bubm.
Sinodendrum Fabr. Labrum exsert, free. Mandibles scarcely
produced beyond clypeus. Maxillae membranous, ciliated, with
internal process acuminate. Antennae scarcely broken, short, with
club three-leafed. Body cylindrical.
Sp. Sinodendrum cylindricum, Scarab, eylindrieut L., Ds Gbbb Ins. TV,
PL 10, figs. 9, 3, Panzbb D, Ins, Heft i. Tab. i. Heft i. Tab. 9,
Buv^BiL Cons, gin. s, I, Ins. PL 5, fig. 3 ; this insect ifi found in many
countries of Europe ; the larva lives in beeches, oaks, 9cc, The male has
a horn on the head, like many Scardbanda,
jEsalua Fabr. Labrum exsert, free. Mandibles produced be-
yond clypeus. Maxillae membranous. Antennae not broken, club
three-leafed. Body short. Feet moderate or short.
Sp. jBsalus scarabasoldes Fabb., Guisizr Iconogr., Ins, PL 27, fig. 2.
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608 CLASS Vltl.
Add genera CeratogruUkus Westw. (and MUophyUus Pabkt),
C&nidma Mac L., {Luetm, tenehrtncles Fabb., species of Flafycents
Latr)
B, LigvUa horny. Antmnos pubetcerU, arcuate. Thorax qwidraUj weptt-
rated by ttricture from abdomen. SeuieUvm in the pedunde of abdomemg
beyond the bate of elytra.
Pasaalua Fabr. (species of Lu/canus L.). Labrum transyersey
emarginate, ciliated. Mandibles broad, armed with thick teeth.
Maxillae homy, slender, with external lacinia incurved, acuminate,
internal shorter, bidentate. Body depressed, mostly oblong. Feet
short,
P. EsoHBCHOLTZ DtM. de Ooleoplerorum genere Paualut, Nimv. Mim. de
la Soe, imp. de Moecou, i. pp. 13 — t^, Ann. dee 8c. wU. xxn. pp. 333 —
336; Pbbohibok Monographic dee Paeealee, Paris, 1835, 8yo. ay. 7 pL
and his sapplementB to thia in Gu^Biv Magaein de ZooL 1841 and 1843.
Sp. Paeealve imterrvptue Fabb., Lueanue tnterruptue L., Yoet CoUopt,
I. Tab. 19, fig. I, 8. America; — Pace, pentaphyllua Gu^bin Iconogr., Ine.
PL 17, fig. 7, Ac. To this genuB belong very many species, nearly all
agreeing in form, and ooloured brown or black; they are in great part
found in S. America and the islands of the Indian Archipelago and South
Sea ; in Europe and North Airica none of them are found. The larva has
much resemblance to that of Lucanue, but the last pair of feet is replaced
by two small conical tubercles, as was first remariLcd by Psbcexbok.
BuBiCEiBTXB gave a more complete description of the lanra and the pupa in
his excellent Hwndbwh d. Entomol. v. s. 454, 459.
Note. — ^To this fimdly of insects genus TrictenoUyma Gbay was
formerly referred by English authors. Now, however, it is more
correctly referred to the Longicomia by Dupont and abo by West-
wood, where it is to be placed after genus Friontis (above, p. 480)
as an anomalous genus. By the number of joints of the tarsi it
belongs to the Heteromera. Antennae longer than head and thorax,
eleven-jointed, with first joint elongate, three last incrassate, forming
internally a sub-serrate dub
Sp. Trictenotoma CfhUdmU Gbat, Dufont, Gu^Biir Magae. de ZooL 1839,
Ine. PL 33, from Java. Westwood has lately added two other species
from the East Indies, Cahinet of oriental EnUmol. 1848, PL 15.
Phalanx 11. Scardbceidea or Petalocera* Club of antennae
lamellate or tunicate.
These insects form with LiNNiEUS the genus ScarabcBtts. They
compose one of the most numerous divisions of the Coleqpterct,
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IN8ECTA. 509
which amongst the Pentamera^ is surpassed by ^e Ccvrabici alone.
Manj large species belong to it, which are armed with knobs,
horns and other excrescences on the head or thorax. As the horns
amongst the ruminant mammals are often appropriated to the male
individuals, so here also these excrescences are mostly wanting in the
females, or much less developed. These insects live for the most part
on plants, some on decaying parts of v^etables or on animal dung.
The larvn live long; some change into nymphs only after three or
four years. The intestinal canal of the perfect insect is commonly
very long, and the stomach forms the chief part of it, (see above,
p. 253). The vessels for secreting urine are long, four in number,
and so much convoluted round the intestinal canal that their termi-
nation is found with difficulty. In the larvae the stomach is short,
wide, and surrounded by three rings of blind appendages, which
however are wanting in the larvie of Aphodiua, (Db Haak J^ouv.
Ann, da Mua. rv. pL 18, fig. 4). The small intestine is very short,
and the large extensive and wide, filling the whole space of the body
behind The air-tubes of the perfect insect form, especially in the
abdomen, many larger or smaller vesicular expansions, by which
these animals, notwithstanding their size, are often very lights which
increases their capacity for flying.
Compare on the diviaion of this group Mao Lbat Hora WUnndlogiotB
and BuBMEiBTBB Hamdb. der BnUnnol. m. pp. 95 — 117.
A. Melitophilct. Abdomen longer than head and thorax, with
last segment {pygidium) produced beyond the truncated elytra,
Lignla connate with mentum. Mandibles membranous internally.
AntennaB ten-jointed, with club three-leafed.
Gomp. H. GOBT and A. Pebohxbon Monographie des Citoines a ffenre$
voUins, Paris, 1833, ^^^' {mth 77 col. pi.), and H. B. Schaum Analeeta
entomologica. Cum tab. nnea. Halis Sajconum, 1841, Svo. pp. 31 — 49.
Cetonta Fabk. Elytra on the outer margin sinuate or excised.
A little triangular part on each side of thorax before elytra (epime-
ron of mesothorax eminent, conspicuous). Mesostemum termi-
nating anteriorly in a process between the base of middle feet.
Scutellum distinct, mostly conspicuous, large, triangular, in some
covered by a lobe of thorax.
^ In the third edition of the Cfataloffue dea CoUopUra de la CoUeetum deM.U
etc DijEAB are enumerated nearly 3300 species.
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510 CLASS yiii-
A very natural^ though numerous genua. The excision <^ the
shield-ooTers next the hind feet leaves generally a part of the
margin of the abdomen visible ; the wings, since the insect flies
with closed elytra, pass through the excision.
Sub-genera : Goliathus Lail (in part), GymnetU Mac L., Cetonia,
CremastochUus BInoch. (Add other sub-genera proposed by GrOBY
and Pebchebok, Bubmeisteb and others.)
Sp. Cetonia Cacicut Fabb., GolicUhui cacieus hUL, Yoet CoUapL i. Tkb.
«^» fig' i5i» Goby et Perch. Monogr. PI. 14, Diet. univ. ^Hitt. not.,
AUaa, CoUopt. PL 6; West coast of Africa, Sierra Leone, &c., as also the
rare and large species C^. goliata, Searabasus Ooliathut Dbubt, Exotic Ina.
Tom. ni. Pl. 40 (copied by Ouvibb CoUopt. i. No. 6, Tab. v. fig. 33,
Dutch edition of Stubic, Tab. 70), Cut. R. Ani. ed. iU,, Ins, PL 45, fig. 4.
{Cfymnetia Mao L.) Sp. Cetonia holoserieea Fabb., Yobt CoUopt. i.
Tab. II. fig. 10, S. America^ Surinam;— CWonta nitida, Scarab, nitidw "L,,
VoET I. Tab. 3, fig. aj. Gov. R. Ani. id, ill., In$. PL 45, fig. 5.
Cdtmia awrata Fabb., Scarab. OMraitu L., B(BSBL Int. n. Scarab, terr.
Gl. I. Tab. n. figs. 1—5, fig. 9, Pankeb D. Ins. Heft 41, Tab. 15; the
golden beetU, gold-green, elytra spotted white, 7'" to 9"' long ; common in
gardens upon roses, elder, Thalidrum, &c. The larva is often found in
ants' nests. — Ceton. cenea Gtllekb., Ceton. jUrioola Hebbst &c.
Trichiua Fabb. Elytra with external margin not excised;
epimera mostly not conspicuous in front of elytra. (Thorax sub-
orbicular in most ; scutellum often small.)
Gompare Lefelbtisb et Sbbyillb Eneyd. mSih., Ins. Tom. x. pp.
701—704.
Sp. TriMus ahdominalis D]^., Ebiohs. (and Trichdus gallicua DiJ.),
Trichivs fasciatus Latb., Gobt et Pebcheboit Monogr. PL 10, fig. i,
VoET CoUopt. I. Tab. v. fig. 43, Houttdtn Nat. Hist. 1. 90 Stuk, Tab. 71,
fig. 8, Guv. R. Am. id. Ul., Ins. PL 45, fig. i ; this species is diffSarent
from the true Trichius fasci^Aus L. (see Gyllsnhal in Append, to Sohoevh.
Syn. Tom. 3), which I have not observed here ; the Trichius fiuciatus L. is
TrichMbs succinctus Latb. (not Scar, succinctus Pall.), Gobt et Pebcheb.
Monogr. PL 10, fig. 7 ; — Trichius hemipterus Fabb., Scarab, hanipterus L.,
VoBT CoUopt. 1. Tab. x. figs. 88—^.
Add genus Inca Lepel. et Seby. (species of Cetonia Fabb.), and
some others of modem writers, ^ere also belongs, in Bubm£IST£b'8
opinion, genus Ev^hirus Kibby.
Sp. Euchirus longimanus, Scarab. Umgimanus Yoet CoUopt. 1. Tab. xi. fig.
97, HoDTTUTir Nat. Mist. i. pe Stuk, Tab. 71, fig. 3 ; habit, in East India
(Bengal). See other species figured in Wbstwood OriaUal Entomology,
Tab. I and 13.
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INSECTA. 511
B. Afiiltobia {Olaphyridoe Mac L.). Elytra shorter than abdo-
men, mostly dehiscent towards the apex. Last dorsal segment of
abdomen {pygidium) surpassing elytra. Antennae with nine or ten
joints, club three-leafed. Mandibles homy, membranous ctn the
inner side alone. Maxilla with process membranous, often elongate
and hirsute. Ligula free, membranous, bipartite, produced beyond
the homy mentum.
A smaJl group of beetles, of which Fabbioius brought the speoieB
luiown to him to the genus Melolontka, but which has just as much
affinity to Trichius, Most of the species are from the old world, especially
from the south of Africa, some are met with also in the south of Europe
(Italy, Portugal). Some species are veiy hairy ; in some the males are
distingruished by very long hind-feet with thick thighs; the booklets of
the four first feet are mostly bifid. Most of the species are small, between
3"' and 5"' ; only few are half an inch or more in size.
Pachycnema Lepel. et Serv.
Sp. Pachycnema crassipes, Mdol, crtusipes Fabb., Gu£bik Iconogr,, Im. PI.
a$ bis, fig. 6 ; habit. Gape of (rood Hope, &;c.
Anisonyx Latr.
Sp. Anisonyx vrstta, Mel. ursut Fabb., Yost Colec^L i. Tab. 5, fig. 35 ;
habit. Cape 0/ Good Hope.
Amphicoma Latr.
Sp. Amphicoma bomJn/liformM, Meld. hoiabyliformisVABR,, GutBin Iconogr.,
Ins, PL 15 bis, fig. a ; habit, in southern Russia, Ac.
Olaphyrus Latr.
Sp. Glaphyru8 m<mru8, Scar, maurus L. &c.
Add genera Lichnia and Gratoscelia Erichsok. Species from
South America (Chili) with eyes bipartite, claws of tarsi as in
Gktphynu and Amphicotna equal, antennee nine-jointed.
See Wibgmanu'b Archvo. 1835, i. pp. 267 — 270, Tab. ni. On other
genera of this subdivision consult Bubmxibteb Handh, der Entom, iv.
pp. 1—67.
C. PhyUophaga Latr. (with the addition of a part of Xyhphili
ejnsd., MelolonthidcB, Anophgnathidce and JRutelidoB Mac L., West-
WOOD). Elytra shorter than abdomen, with pygidium free. An-
tennae with eight to ten joints, club with three to seven. Man-
dibles homy as to the greatest part, frimished internally with a
lamina membranous, ciliated, with a large tooth at the base.
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612 CLASS VIII.
Ligola homy, immoveable, coimate with mentmn. Labmm mostly
exsert, homy.
These beetles in the perfect state live in great part on leaves, the
hCrvse on the roots of all kinds of plants.
Butela Latr. Labrom exsert, mostly descending. Mandibles
homy, exsert, with external margin often somewhat prominent,
crennlate or dentate. Maxill» with several teeth -at the apex.
Antennae with ten joints, club three-leafed. Feet thick, with claws
of tarsi unequal, incurved.
Areoda Leach, Chryaophora D£j., FeUdnota Mac L., Rutda
Mag L., Macraspia Mao L., Chcumodia Mao L., and some other
sub-genens on which see Bubmeisteb Entom, Hcmdb. iv. 1,
pp. 330 — 429.
Sp. Rtdda dvnftoehlora Latb., Chryiophora ehrygochlora D^,, Huxboldt
et BoNFLAND, Obterv, de Zool, el (TAnai, comp, i. PL xv. figs, i, a,
GuiBiN Iconogr,, Ins, PI. 34, fig. i ; Peni ; hind feet in male long, with
broad thighB, and tibiae terminating in a triangular process turned inwards.
Without having this process the hind feet in the male of Soarabatu macro-
ptu Fbanoilloit, Shaw, Ckryiophora maeropa D^., are in like manner
▼ery strongly developed.
Rutela Uneola Latb., Scarcibceui {SurinamM and) lineola L., OeUmia
Unecla Fabb., Yoxt CfoUopt. i. Tab. n. fig. 81 ; Surinam, Braals. — Jtutda
9plendida SOHOIVH., Cetonia iplendida F., VoKT Coleopl. I. Tab. vm.
fig. 69, kc
All the species of this genus are exotic ; the most are found in South
America ; they have usually lively colours or a metallic lustre.
Anomala Meqerle {Euchlora Mac L.). Mandibles with ex-
ternal margin rotundate, entire. Antennae with nine joints. (Other
characters of the preceding genus ; feet less thick, with external
claw of four anterior feet mostly bifid.)
Sp. Anomala Fritehii (Mdcl, FrUchii and MeL Julii Fabb.), Pahzbb
Deuttchl. Ins. Heft 97, Tab. 9, 10, Ratzkbubo JPont-Int. I. Tab. 3,
fig. 10 ; 4'" to 6'" long, green with usually yellowish brown elytra ; it
differSj however, much in colour ; see the accurate description and syno-
nymy in Ebiohsoh Natiurgesck, der Ins, DeiUsehl, CoUopt. m. i847>
B. 635 — 637. — Arwmala koriicola, Scarab, horiicola L., Mdol. horticoia F.,
{Anisoplia horiicola Meg., DiJ., Catal. des CoUopt. 3e Mit), Panzeb
DeutsckL Ins. Heft 47, Tab. 15, Ratzbbubo Foni-Ins. i. Tab. 3, fig. 9;
4" long, hairy, with brown-red shield-covers, steel-ooloured blue thorax ;
very common.
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INSECTA. 513
Amsoplia Mboerle, Dej. (in part). Clypeus angustato-produoed,
recurved.
Sp. Anitoplia fruHcola, Mdol. frvtiticdla F., Ratzebubo PonA-Im. I. 'Lth, 3,
fig. 7 ; in the easterD part of the middle of Europe.
Note, — Sub-genera Strigoderma D&r., Popilia Lbaoh, Mimela Kibbt,
Bhinyptia Di^. Compare Bubmeisteb Handh, der EnUmol, iv. i,
pp. 214—317.
ATwphffnathus Mac L. (with the addition of some other species).
Labrum porrect, descending, with middle portion acuminate, pro-
duced, touching mentum. Mandibles externally rotundate, con-
cealed beneath labrum. Antennss with ten or nine joints, club
trifoliate.
Sub-genus : Anojjlognathua Mao L. (and Repsimua Leach). Tarsi
simple, short, thick, with claws simple. Mesostemum elongate
mucronate. Antennae with ten joints.
Sp. Anoploffnathui vindi-ameus Mao L., Jtutela LatreiUU Sohorkh., Cuy. i2.
AfU, id, ill., Ins, PI. 42, fig. 3 ; from New Holland, like the other species
of this sub-genus.
Plabyccdia D£r.
Species American. Add sub-genus PhaUmffogonia BxTBir.
Brcuhyatemus Guerin (Amhlyterus MacL., Anoplaatemua Guerin
kc). Mesostemum not acuminate. Antennse with ten joints,
Adorehia Eschsch. {Trigonostoma Di^.)
Note, — On this and other sub-genera see BubmiisteB Mandb, der Eniom.
IV. I, pp. 437—477.
Geniates Kibbt. Tarsi dilated, densely pilose beneath (in some
species of anterior feet alone and only in males). Antennie more
often with nine, sometimes with ten joints. Mesostemum not
mucronate. Claws of tarsi imequal, one mostly bifid, the other
simpla Mentum in males mostly rough with close setse.
Sp. Oeniaiei harbaiui EjntBT Linn. Traruact. xii. PI. 21, fig. 8, OvtsLOS
leonogr., Ins. Fl. 94 bis, fig. 3 ; habit, in Brazil like several species (all are
American) of this sub'genus, to which are to be added the allied genera
Evanos IiATOVUE, Leucothyreus Mao L. and Bclax Fibchxb (and Lcacopyga
Westwood).
Melolontha Fabr. (exclusive of some species). Labrum trans-
verse, emarginate in the middle, mostly bilobed. Mentum homy ;
VOL. I. 33
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514 CLA88 VIII.
lignla mostlj homj, emarginate. Ma&dibles homy, tnangiilur.
Antenna with from seven to ten joints, club with three lamellc or
more, which are mostly longer in males.
HopUa lujG., Latb. Mandibles with broad membianons border
intemallj. AntemuB with nine or ten joints^ dnb trifoliata daws
of tarsi nneqnal, posterior feet often with daw sm^e, strong. Body
sqnamoseL
8p. BopUa phUafOkut Latb., ffoplia wrgadm Fabb., Pahzxb DotfidU. lin^
Heft aS, 1U>. i8, BATsnnnta /brrt-/iMi i. 1U>. ^ figs. 14, 15, Ac.
ilTofe. — ^Here iwe to be ineerted eerenl genera of more modem wiiben,
JfonodUIitf iLua., QfmimiUmA D&. ftc. ; on which conenh BnumBtiE
Hamdb, der EnUmol, vr, 1, pp. 84 — soa.
Serioa Mag L., Omaloplia Mfla, D^. Labrum connate with
cljx>eii& Mandibles with broad membranous border intenially.
Claws of tarsi equal, bifid
%». Serica hruimea, Seardbmu bnmnem L., MdU, hrwtmM Fabb., Von
CoUopt, I. Tab. YIL fig. 55, Pavzbb I>eyi$dU, Im. Heft 95, Tah. 7,
Ratzxb. Pcmt-I'M, I. Tab. 3, fig. la, Ac
NcU. — ^Add eub-genera AtUmOf Symimda, AtKUa Ebighbov, AVUben
'Dta,, and Bome oihen omitted here.
Melolantha Latb. (and Ehizotrogus ejuad.) Labrum distinct torn
clypeus, descending obliquely backwards. Mandibles with narrow
membranous border internally. Claws of tarsi equal, mostly
denticulate near the basa
a) Olub of antennae trifoliate. Claws of tarsi supplied with a sharp denti-
cle at the base. EhUotrogut Latb. Antenne with ten joints, AmpikimaUtu
Latb., Mulsaht. Antennte with nine joints. Sp. MeioL tolttkialit
Fabb., Scarab. BoUtUialit L., YoiT Ool. i. Tab. vi. fig. 51, Ratibbubg
Fort^Im. I. Taf. m. fig. 5 ; brown, hairy, 7'"— 8'" long.
b) Club of antennsB in males 5 — 7, in females 4 — 6 jointed. {Anoxia
Lapobtb, PclyphyUa Habbis, Ebiohbok, MeUdwUha Ebichboit.)
Sp. MddUmOM fuUo, Scarab, fuUo L., Bcbsbl In$, iv. Tab. XTi, Vobt
OoUopt, I. Tab. VI. figs. 48, 49, Dum^bil Com, gin, 9. I. Tnt, PL 4, fig. 6;
de diUnhever, le fouUm, grosser Julius hever, DUnenk^fer, the FuUer;
1$'" long ; antennsB of male with seven long leaves at the end, those of
female with only five shorter ; blackish brown and sometimes red-brown,
with white irregular spots on the shield-oovers, as thongh marbled;
scutellum almost entirely white.
Md, vulgaris, Scarahceus melolonlha L., Bcbsbl Ins. n. Sear, terr, CL i.
Voir Coleopl, i. Tab. Vi. figs. 45, 46 ; de meikever, le kaaneion, der
Maikitfer, the Cockchafer; nearly one inch long; the male with seven
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IKBECTA. 515
leaves at the chib of antenim, tbe female with six shorter ; thorax black
with a white felt ; stone-ooloured or red-brown Bhield-oovers. The larva
lives three years underground^ and destroys the roots of plants ; m the
autumn of the fourth year it changes into a nymph, from which the beetle,
that lives on leaves of all kinds of trees, after some weeks makes its
appearanee, but commonly continues under ground until May. This
insect in some years appears in large numbers, and causes dreadful damage,
as for instance, according to the newspapers, in 1836, around QuedUmburg,
where, in the middle of May, 133 Dutch mud (of four bushels) were col-
lected. Another species, somewhat smaller, with reddish-brown thorax, of
the same colour as the shield-covers. Meld, ffippoeattani Fabb. (Panzbb
DeuttcM. Ins, Heft 97, Tab. 6, Roebbl 1. L 6gs. 9, 10) is met with in this
country in some years as well as the former, and sometimes in no fewer
numbers. Compare on the cockchafer SuKOW Naturgeich, de» Mcahitfert
{aua dem, latm SUick der VerluMdl, det Baditchen Landwiaittk, Vtreiwt)
Carlsruhe 1834, (with an anatomical description of the larva and beetle),
and Stbaus Dubokhbim Cent, gin, kc,
D. Xyhphila {Xyhphili Latr. in part, DynasHdce Mac L.,
Westw.). Elytra shorter than abdomen, with pygidium free.
Antennse short, mostly with ten (more seldom eight or nine) joints,
with club lamellate, always with three joints. Mandibles princi-
pally homy, produced at the apex beyond the clypeus, armed with
a tooth at the base, mostly supplied internally with a border mem-
branous, ciliated, narrow. Ligula homy, connate with mentum.
Labrum covered by clypeus. Claws mostly equal (except those of
anterior tarsi in males of some species). Scutellum distinct, moderate
or small, broad, triangular, with apex rotundate.
These insects often present a great difference of form in the two
sexes, the head and thorax in the nude being armed with homy
excrescences, which in the female are less developed or entirely
absent They are usually brown or black in colour ; some exotic
species are amongst the largest of beetlea The larvsB live in decay-
ing wood and in garden mould.
t C%p«iM hroad, mosUy rotundaie or tubtimuUe. Bead of both $eM$
unarmedf or mpplied vnth a small tubercle, never homed,
Hexodon Oliv., Fabr. Mandibles not crenate exteriorly. Body
sub-orbicular. Elytra dilatato-marginate. Head small, with eyes
minute.
Sp. ffexodon reHcuUOum Oliv., Gu^nr leonogr., In$. PI. 23, fig. 6; babitat
Madagascar.
Pachyhis D£j., Bubh.
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516 CLASS VIII.
(hfdocephala Latr. Mandibles not crenate exteriorly. Body
oval. Elytra not dilated. Anterior tarsi in males mostly incras-
sated.
Species exotic, most from South America. FABBionrs placed those known to
him amongst the MdolotUha, Sp. Oydoc. mdanocq>hala, Md. mdcBno-
eephaHa Fabb., Yost Goleopt, i. Tab. 9, fig. 80 ; 0yd. undata, Mel.
tpiUyphihaJIma Hehbst, Vobt CoUopt. 1. Tab. 10, fig. 91.
jVb<0. — Add genus Cha^va Mao L. and some other sub-genera described
by BuBifEiBTBB Handb. d. Entom. v. pp. 18 — 86.
1 1 ClypeuM narrow, often acute; mandSblcB at the aides qf dypeue not
covered, with external margin often crenate.
Scarahmia Latr. (with the addit. of genera Phileurus ejusd.
and Oryctes Illig.), Geotrupes Fabr.
a) Head nU different in the eeooes, never homed but either emooth or
armed with one or two tuberdee, gtkiboue. Thoram in malet qften tuberculate
or homed.
Strategus Kibbt.
Sp. Scar. Alcnu L., Vosr Coleopt. i. Tab. xvm. fig. 139, Tab. ziz. fig. 138 ;
habitat South America.
b) Bead d^erent in the eexes, of male tuberculate or homed, qf female
obidetdy tul>erculate or entirdy unarmed.
Sub-genera: PhUewma Latb., AgaocephcUa Mannisrh., Oryctes
Illig., Sccurahceua Latr.
Note. — This section of the SearaboBi is indeed numerous, but the genera
of modem writers, most of them artificial, scarcely seem to throw any light
on the distinction of species and the natural airangement. Oryctes indeed
is distinguished by defect of teeth in the maxilla, but there are species
similar in habit with denticulate maxillsB, which cannot be separated firom
the Orydidoe (genera 8typotrupea Bubm., Xyloryctee Hops). Oharacter
non faeit genue.
Sp. Scarcibcene nasieorme L., Swamhsbdam BM. Nat. Tab. xxvn., RonxL
Ins. n. Soar. terr. CL i. Tab. vi.— ix., Panzbb Deutechl. Ins. Heft a8.
Tab. 3 ; the Bhinoceroe-beede; found especially under bark in hot-beds.
Scarah. HercvXea L., Vobt Coleopt i. Tab. xn. fig. 98, BocssL Int. n.
Scar. terr. Prsef. CI. i. Tab. a. i. Tom. iv. Tab. v. fig. 3, from the West
Indies, &c. These large beetles live on the sap that flows from wounded
trees or roots ; for that purpose they wound the bark with their honi, aa
the Liteani do with their jaws.
E. ArenicolcB [TrogidcB Mac L., Westwood, and Geotrupes
eorund.). Elytra covering the abdomen completely. Abdominal
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IN8ECTA. 517
stigmata situated in the membrane between the ventral and dorsal
segments, all covered by the elytra. Ligula not distinct from
mentum. Labmm homy, mostly exsert beyond clypeus. Mandi-
bles homy. Antennae with ten or eleven joints, club always
trifoliate.
A, VerUrcU segmefnta of ahdoTfienfioe,
TVox Fabr. Lignla covered by mentum. Antennae with ten
joints. Head small, nutant. Thorax transverse, with anterior
angles produced. Elytra hard, posteriorly involute, inflected,
rugose, often muricate in longitudinal rows.
Sp. Troz arencvritts Fabb., Silpha tcabra L., Paitzsb Deutsehl, Ins, Heft 97,
Tab. I ; — Trox aabuUmu It,, Scarab. aaimUmu L., Stubm Fawm, Iru. 11.
Tab. 38. These insects rarely fly. There are species that have no wings,
as Trox horridus ¥., from the Gape of Good Hope ; they form the sub-
genus Phobenu Mao L.
B. Ventral aegrmmta of abdomen six,
Hyho8oru8 Mac L. Ventral segments of abdomen connate, ex-
cept the last firee. Antennae ten-jointed.
Sub-genera: Phoeochroua Lap. {AtimuB Dfj.), ChceiochM West-
wood, Ac.
Orphnus Mac L.
Oeotrupes Latr. Ventral segments of abdomen free. Antennae
with eleven joints, club perfoliate. Eyes divided by the margin of
head. Mandibles homy, armed at the apex. Ligula bilobed, with
laciniae exsert beyond mentum. Body oval. Feet strong, with
tibiae spinose.
Sp. Qtotni^TypkoBM, Scarab. TyphoBush., Fabb., Vobt Odleopt. I. Tab. xix.
figs. 194, 195, Panzbb DetEtschl. Ins. Heft 2, Tab. 93 ; black, the male
with three horns on the thorax directed forwards. — OcUr, tiercorarius,
L., Fabb., Panzbb Deuttehl. Im. Heft 49, Tab. i, Stubm Fauna, Jna. i.
Tab. vi. fig. a. — Oeotr. vemalia. Scar, vemalii, &c.
Sub-genera: ButboceroiS Kibbt, Odontasus Meoeble and some
others of modem to be inserted here.
Lethrus ScoPOLi, Fabr. Ventral segments of abdomen free.
Antennae with eleven joints, club obliquely truncated, tuiicate.
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518 CLASS VIII.
with ninth joint including the two following. Eyes bipartite by
the margin of head. Mandibles homy, large, exsert, ending exter-
nally in a tooth incnrved, obtuse, with internal side serrate towards
the base. Thorax very broad. Body ovate, short. Elytra con-
nate ; only short rudiments of wings. Posterior feet remote.
Sp. LethruM cephalotes Fabb., Paitzsb Jkutsebl. Jna, Heft aS, Tftb. f,
Ddm^il Cons. g6n. «. Z. Tna, PI. 7, fig. i ; habit, in Austria and soath of
F. Goprcphaga {Aphodvidce Mac L., Westw., and ScarabceidoR
eorund., Copridea Leach, Ebichs.). Abdominal stigmata situated
in the membrane between ventral and dorsal segments, all covered
by elytra. Elytra entirely covering abdomen in some, truncated in
others, not concealing the posterior part of pygidium. Ligula
separate &om mentum. Labmm membranous, covered. Mandibles
membranous, ciliated. Antennae with eight or nine joints, club
always trifoliate.
Gopr%8 Geopfb. Antennae with eight or nine joints. Ventral
segments of abdomen connate. Posterior tibiae with single spur.
Scutellum mostly indistinct or very small. Intermediate feet more
remote from each other at their insertion than the rest, sometimes
very distant.
NaU, — lAxaj species are without anterior tarsi; in some there is a
sexual difference, so that this defect is peculiar to males. Compare
Bbull^ Ann. da Sc, wU, sec. S4rie, Tom. Tin. 1837, Zoolog. pp. 346 —
149. They form balls from dung, roll them very diligently, and having
indiosed an egg in each, hide them undeiground.
Ateuchu8 Webxb, Fabb., ScarabcBua Mac L. Posterior feet
elongate, with tibiee slender, tarsi filiform ; last joint of tarsi longer
than rest. Head not homed, with broad margin and clypeus often
both dentate. Eyes partly divided by exsert margin of head or
entirely bipartite,
Sp. Ooprii tacer, Scarab, iooer h., DuiciB. Cons, gin, s. I, In$, PL 4, fig. 4 ;
this species is found in southern Europe and in the north of Airioa ; it may
with the Atctichns jEgyptiorwn Latr., Gu^lK Iconogr,, Ina, PL 9r, fig. i,
be taken for the sacred beetle which the Egyptians figured on their monu-
ments, and imitated in different kinds of stones as ornaments for the dead
and amulets ; this beetle was by others taken for an emblem of the world
on account of the ball which it rolls ( . . . T\d(ra» a^oipotuHk wofiarK^
(Tcor r$ K6fffufi ffXV/*^ HoaAPOLL. BierogL I. 10).
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IN8ECTA. 519
See on this genus Wibtwood Deter, of new speeiei qf ioered beeUet,
Trans, of the Linncean Soc. n. pp. 155 — 163, PI. 99.
Sub-genera OymMopUurue Iluo., 8%»yphu$ Latb., OirceUium IlATR.,
CantKon Hofvmannb, {Coprobitu Latb.) and some others.
Copria Fabb. Posterior feet with tibisB dilated at the apex and
tarsi ciliated beneath. First joint of tarsi larger than the rest.
Head of males often homed (sometimes of females also).
a) With third Joint of labial piifpe very 9m<M
Subgenera: Euryttemug Dauc. {^tehrotea Mao L.), Dr^panoeerui
KiBBT, OMHoeOui Zibol., OtUhophaguM Latb.
Sp. Ooprie mtchicomUf Scarab, nuehioomis L. ; — Ooprit ecBHobita Fabb.,
Panzbb Deutachl, Ins. Heft 48, Tab. 6, Ac.
b) With ihird joint of labial palps distinct.
Sub-genera OniHs Fabb., PhancBus Mao L., Oopris Latb., Ontherus
Ebiohb., Pvnotus ejusd., Canlhidium ^usd., Macroderes WssTW., Soaio-
wmus Ebiohb., Ontocharis D^., ScoHmus EaiCHa, Charidium Lbpslkt.
et SxBV., Pedaria Lap., Ooptorhina HoFi.
Sp. Oopris mimasVAVB., Scarab. Mimas h., Rcbbxl Ins. n. Sear.CL i. Tab. b,
fig. J, VosT Chi, 1. Tab. xxm. fig. 4 ; habitat in South America, fta
Aphodivs iLLia., Fabr., Latb. AntennsB with nine joints.
Abdomen with ventral segments free. Posterior tibiae with two
terminal spines. Eyes undivided. Scutellnm distinct.
Sub-genera : ApSuod/wM^ Oxymm and some others proposed by Mulsaitt.
Sp. Aphodim fimetairius, Scar, fimelatius L., B<bsbl Ins. n. Soar, terregtr.
PntfaL O. L Tab. A. n. fig. 3, Voir CoL i. Tab. 11, ^, 147, Paihexb
Deutschl. Ins. Heft 31, Tab. 9 ; — Apkod. subterranetu. Scar, eubterraneus
L., Pahzib Deutsckl. Ins. Heft 98, Tab. 3, fto.
Compare Sghobnhsbb Syn. Insect, i. i, pp. 66 — 89, and SomaDT
Semeion der D&Osehen Aphodien-Arten, Gebmab's Zeitschr. f. d. £nUm. n.
8. 83—175.
I^ote. — Here are to be placed, according to Ebiohson^ genera
^ffialia Latb., and Chiron Mac L. ; thej differ however, in the
labrom and mandibles beiog homy; wherefore they ought to be
counted with the preceding section of the ArenicolcB as by Latbeillb.
jEgialia is enumerated by Westwood with the Trogidas; Chiron^
on account of a certain similarity of habit, is considered by many
to be allied to the FcusaH
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520 CLASS mi.
Familj LYIL Xylopkaga nob. (Genera firom the fiimilj of the
Xyhphagi Latr., Ptiniores and Xyloirogi ejnad.). Antennae fili-
form, sometimes serrate or pectinate (at least in males), in others
terminated abmptlj by larger joints. Mandibles homj, short, den-
tate. Maxillse bilobed. Palps filiform or thicker towards the
extremity, short. Body cylindrical or oraL Head, as to the
greatest part, mostly received in thorax.
Bostrichus Geoffb. (Apaie Fabr.). Antermae with ten joints,
club triarticolate, perfoliate. Tarsi with joints simple, first rery
small, scarcely distinguishable. Thorax gibbons.
Sp. Boibriekma eafmeiuut, Bermetta eapmeimmt L., Gworraar Itu. L Tib.
5, ftg. I, DuK^UL Cons. gtn. jl I. /m. PL 17, fig. i, lUnDUBfi Por^
In$. L Tab. Z. %g. 14, Ac
iVoa Fabb.
Dysides Perty.
Gis Latr.
Anobium Fabr. {IVnus L.). Antennae with eleven joints, the
three last larger, the ninth and tenth obconical, the eleventh ovaL
Body oblong, anteriorly and posteriorly rotondate. Tarsi short,
with five distinct joints.
Sp. Anobium ttriatum Iluo., Anobium pcrftnox Fabb., Pajizke JDlatftdU. Int.
Heft 66, Tab. 5; i^ — i'" long, black-brown with striated elytn. Tliey
live in old wood, and render our fdmitnre wormeaten, by small round holes,
from which a fine yellow powdered wood fells. The larva Uyes in these
cylindrical passages, and resembles^ like that of Apate, the larva of the
cockchafer in miniatue (see ftg. of the larra of Anob. teudlatum in Ratzi-
BUBO Forst-In9. i. Tab. n. fig. 19, of ApaU ilnd. Tab. XIY. ^. 33). Thia
little beetle causes a ticking sound in the wood, from which -vulgar super-
stition derives an nnfikvourable omen, as appears from the names TodUnnkr,
rharloge de la mart, Deatk-^wtOek. Another smaller, red-brown species,
Anobium paniceum, Dermates paniceut L., Paitzbb Deutschi. In$. Heft 66,
Tab. 6, lives in biscuit, wafen, kc., and sometimes gnaws through the
corks of bottles filled with white, sweet wine.
Comp. on the German species of this genus Stubm Fctuna, Int, XI, 1837,
s. 98 — 144. The intestinal canal of J 1106. ttriatum \b described and figured
by L^H DuBOUB, Ann, des Se, not. xiY, 1822, pp. 919 — sii, PL xn.
DryophUus Chbvbol. With three last joints of antennas cylindrical,
very long, slender.
Hp. Anobium putillum Gtllbbh., Dryopk, anobioikUi Chxvbol^ GuiBor
Magai. de Zool. 1833. CL ix. pL 3.
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1N8ECTA. 521
Dorcalama Herbst, Fabb. Antennse with first joint thick,
conical, the foUowing to the eighth small, the three last forming the
greatest part of antennae, the ninth and tenth serrated internally,
eleventh ovaL Body oval, abbreviate.
Sp. DorcaUma Dntdense Hjebbst, Panzxb DeuUehl, Ins. Heft 26, Tab. 10,
&c Ck>mp. Fiffwret in Stubm 1. L Tab. 144, 145.
JPtinus L. (in part). Antennae with eleven joints, filiform, long
(longer than half the body), with joints sub-equal. Thorax cylin-
drical, constricted posteriorly. Scutellum small, distinct. Feet
long, slender, with tarsi distinctly five-jointed, the first joint
elongate.
Sp. PtiMufur It., Panzkb DeuUehl. Iiu. Heft 99, Tab. 4, ma. (PtMvui davipes
Panz.), Gboffb. In». i. PL n. fig. 6 fern. ; i^ — 1'" long, the aatenn» of
the nude somewhat longer than body, the abdomen elongate, cylindrical,
BO that it resembles a small goat-beetle ; the female with the abdomen more
OTsl and wingless. This insect lives in old wood, in herbaria, in collections
of birds and insects. GoszB has described and figured the metamorphosis
drcumstantiaDy; Natwifoncher, vm. 1776. s. 76 — 89, Tab. n. figs, i — la.
Oihbium ScOPOLi. Antenna with eleven joints, filiform, longer
than half the body, with joints obconical, sub-equal, the last oval.
Thorax broader posteriorly. Head small, nutant. Elytra united ;
wings none. Scutellum none.
QMium acoHoi, Ptinua Scotica Fajbb., Panzxb DeuttchX. In$. Heft 5,
Tab. 8.
Add sub-genus Mezium Cubtis, Stubm Fcmna, In$. xn. pp. 54 — 58,
Tab. 148.
Ptilinua Geoffr., Fabr. Antennae with eleven joints, not
longer than half the body, serrate or (in males) pectinate. Thorax
posteriorly of the breadth of elytra. Wings and scutellum distinct.
Feet short, with tarsi distinctly five-jointed.
Sp. Ptilinua pectinicomis Fabb., Ptinus pectinicomis L., Pakzxb DeuUehX.
Ins, Heft 3, Tab. 7, Stubm J^avma,, Ins. xi. Tab. 336, &c. Add sub-genus
Xyletimu Latb., oomp. Stubm ibid. Tab. 237.
Lymexylon Fabr. Body elongate, narrow. Antenns of the
length of head and thorax, with eleven joints, filiform or sub-ser-
rate. Mandibles short, thick, dentate. Maxillary palps of males
with last joint ramose, flabellate. Head almost globose, nutant.
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522 CLASS VIII.
separated from thorax bj a neck. Eljtra shorter than abdomen,
sometimes very short. Tarsi distinctly five-jointed.
Sub-genera: Lymexylon Fabb., HyhcoBtfua Latb., Atrat^ooenu
Palisot-Beauvois.
Qenus Gupes Fabb. scarcelj belongs here.
Family LVIII. Serricornia s, Priocerata. Antennae mostly
short or moderate, more rarely incrassated towards the apex, filiform
or more slender at the apex, serrate or pectinate, especially in
males. Body elongate. Elytra mostly covering abdomen entirely.
Penultimate joint of tarsi often bilobed.
I. Prostemum not produced anteriorly beyond the mouth, nor
mucronate posteriorly (with few exceptions). Body mostly soft,
flexible {Midoboodernuxta Latr., Aproatemi Westw.).
Glerus Qeoffb. Palps, either all or at least labial, with last
joint truncated, mostly broad, obtrigonal or securiform. Mandibles
with apex acute, internal side emarginate or armed with another
tooth. Penultimate joint of tarsi bilobed, first in many very short
or indistinct. Head and thorax narrower than abdomen; head
posteriorly received in thorax, AntennsB filiform or subseirate, m
many clavate, with three terminal joints thicker.
Sub-genera : EnopUwn Latb., Necrcbia Latb., {Gorynetes Fabb.),
Lcuricobiiu Rosenhaueb, Glerus Geoffb., Latb. {Trtchodes Fabb.)
Thanasiimta Latb., EpicUnes Ohevb., OpUo Latb. (Notoxas Fabb.
not Geoffb.), Ewrypus Kibby, Axina Kibby, TUhts Oliv., GyUdrvs
Latb.
Sp. CleruB cUveaarntg, Tri^4)de$ alveariut Fajbb., Fakzsb DevUchl, Ins, Heft 31,
Tth, 14, Bj^aumub Ins. vi. Tab. 8, figs. 9, 10, pp. 8a, 83; the red larva
lires parasitically in the nests of wild bees (of the genus Osmia), — Olerus
viciaeeiu, I>ermestes violaoeus Jm, Necrobia violacea Latb., Panibb DeutsekL
Ins, Heft 5, Tab. 6, DuM^B. Cfons, gin. «. l. Ins. FL 17, fig. 4, &o.
Pdocophora Dej.
Melyria Fabr. Palps filiform, short. Mandibles emarginate
or bidentate at the apex, narrow, elongate. Thorax oblongo-quad-
rate, little convex or plane, covering the base alone of head. Joints
of tarsi entire ; claws unidentate or bordered internally at the base
by a membrane forming an appendage resembling a tootL Body
oblong, with dorsum plane. Antenn® filiform or serrate.
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IKSECTA. 523
Sub-genera : Daayiea Payk., Fabb., Melyrta Fabb., Z^gia Fabb.,
McUackiuB Fabb., Oliv.
Sp. Melyria eoandea Illio., Dcuyia coandeuB F., Panzer DeuUdd, Ifu, Heft
96, Tab. 10 i^Malaehwi ceneus, Cantharii amea L., Voet Cideopt, i. Tab.
4^> % 7> Fanzkb DeutschL /lu. Heft 10^ Tab. 2 ;--^Malach. bipuahdaitu
Fabb., ConCAar. hipuatulaia L., Yost Co2. i. Tab. 46, fig. 6, Fakzsb
DeiOteM. Int. Heft 10, Tab. 3, fta
MaUhinua Latr.
Telephoms Sch^ff., Latr., Caniharis Fabr., DbIj. (Species of
Cantharides L.). Palps securiform. Antennae filiform, with joints
elongate. Mandibles veiy acute, not emarginate. Penultimate joint
of tarsi bilobed ; claws simple. Body oblong, soft. Head deflected,
covered at the base by thorax.
Sp. Td^horw fiacus, Cantharii fuaea L., Yokt OoiUopi, i. Tab. 46, i^, 3 ;
— Tdeph. mdanumu, Oanthar, mdanura L., Yost ibid. fig. a, Panzxb
Ikuttchl. Ina. Heft 85, Tab. 6, &c. On the metamorphosiB of certain species
cons. Db Gxbb In$. m. Fl. 2, figs. 5 — 11, and Blakghabd, Gu^bin Mag,
de Zool. 1836, Im. PI. 168.
SiUa Meoeble, D£r.
Drtlus Oltv. Palps thick, conical. Mandibles bidentate. An-
tennsB serrate or pectinate in males (antennae of females with ten
joints obconical, subserrate). Penultimate joint of tarsi bilobed ;
claws simple. Females apterous, elongate, resembling larysB, far
larger than males.
Sp. DrUva fla/tfeteen§ Oltv., Gxoffb. Int. i. .PI. i, fig. 15, I>uifj^iL Cons,
gin, t, I. Ina, PL 9, fig. 5, male, GuiaiN Iconogr., Ina, PI. 14, figs. 4, 5.
Hie larva was discovered by the Count Miblzinskt living as a parasite in
ffeiix nemoralia, and the insect which proceeded from it (M. saw the female
only) conaideired to be a new genus, and named CochleoCUumta, Ann, dea
8c. not. I. 1824, pp. 67 — 77, PL 7. Dbsmabest first discovered that this
animal was the female, hitherto unknown, of Drihta flaveaceM ; Ann. dea
Se. not. n. pp. 157 — 170; oomp. also, ibid. p. 443, Ac., V. AuDonnr
Rech. amUomiguea aur lafemelle et le mdU du Drile jawndtre, PL 15.
Lampyris L. Antennae approximate at the base^ filiform^ with
eleven joints. Eyes in males very large. Thorax plane, semiorbi-
culate, covering head, concealing it. Mandibles entire. Penulti-
mate joint of tarsi bilobed; claws simple. Females mostly
apterous.
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524 CLASS VIII.
Sp. Lampyrit tplendidula L., Panzxb Deuischl. Ins, Heft 4, Tab. 8, m. and
fern., SoHiBFFEB, Elem. ErUom, Tab. 74 (nuJe), DuM^iL, Con9,g6n, 9. 1. Int.
PI. g, figs. I, 2 {de glimworm, Johannismirm, le ver luisant, the glouhworm).
This species, and some others {Lampyris noctUuca L., Lamp, hemiptera.
Lamp. UeUica, &c.), diffuse a phosphoric light by night; the males shine
less than the females, and with them the light ceases after the pairing; the
females, on the other hand, are very luminous when laying eggs. 6. B.
Tbevibaitub was of opinion that the light proceeded fix>m the internal
genital organs ; this subject is fully treated by him Biologic, V. s. 105 —
116, Verm, Sckrifien, L s. 87 — 93. The light becomes dull in carbonic
acid and hydrogen gases, and stronger in oxygen gas; see on these and
other particulars a letter of Mattxuooi to Duica.b in the Camptes rendnu,
14 Aug. 1843.
On the diffierent species of this genus oomp. Eucd cTimi^ reviaion du genre
Lampyre par Da La Pobtk Ann, de la Soe. entamoL u, 1833, pp. iss,
"3.
Phengodes Hofvmanns. Antennse with eleyen joints, bipectinate,
with cirri oonvoluted spirally. Elytra posteriorly narrowed.
Species American, as LawpyriM plumoea Fabb. &c.
Amydetea Hoffmanns.
Omalism Geoffr,
Dictyoptera Latr.
Lycvs Fabr., Oliy. Head small, concealed under the shield of
thorax, with mouth porrect, rostrate. Palps with last joint thicker.
AntennsB approximate, filiform or serrate. Elytra in many dilated,
broader than abdomen.
Gomp. on this genus Fabbioh ^^10^. Elevth. n. pp. no — 118, and
Dalman in SoHOENH. Synon, Ins, i. 3, Append, pp. 45 — 33, Tah. 5.
Cyphon Fabr., {Elodea Latr., Sdrtea Illio., Latr.). Mandi-
bles entire, with apex simple, small, scarcely exsert beyond
labrum. Palps filiform, labial furcate at the apex, with last joint
inserted before the apex of penultimate. AntennsB distant, longer
than thorax. Elytra rather soft. Posterior feet in many saltatoiy,
with femora incrassated.
Sp. Oyphon hemisphoerieus Fabb., Pakzsb DenOsehl, Ins, Heft 96, Tab. 7, &a
Add genera Evibria Zxeql., NycUua Latb.
DascilliM Latr.
PUlodactyla Illio.
Sub-genus : Colobodera Kluo*
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INSECTA. 625
Comp. Kluo Ueber ewei neue KdfergcMmgm^ Wiboxann's Archie /.
NaturffCKh. 1838. 8. 67 — 70.
Bhiptcera Latr., KiRBT.
Sandalua KnOCH.
CaUtrhipts Latr.
Anelastea KiRBT.
Cehrio Oliv., Fabr, (in part). Mandibles entire, with apex
simple, exsert, decussating. Palps filiform, labial with last joint
obconical, truncated. Antennae distant, longer than thorax (in fe-
males shorter, thick). Thorax broader posteriorly, with angles
produced. Prostemum mucronate posteriorly.
Sp. Cdnio giffos Fabb., Dux^ril Com. gSn, «. I, Iiu. PL 3, fig. i, Panzkb
Jkuiichl. Ins. Heft 5, Tab. 10; in south of Europe. The female lives under
ground, and continues there at pairing time, protruding the abdomen alone,
so that the nude does not see her. AuDOUiN et Bbull^ ffia^. not. det Int.
VI. 1837, pp. Ill, 272.
Phy9odactylu8 Fischer.
II. Prostemum produced anteriorly as far as the mouth or be-
yond the mouth, mucronate posteriorly. Elytra hard. {Stemoxi
Latr., Macrostemi Westw.)
Elater L. Antennae setaceous, serrate or pectinate. Mandibles
emarginate at the apex or bidentate. Palps with last joint larger,
obtrigonal, truncated. Mucro of prostemum received in a cavity of
mesostemum. Body elongate, with thorax oblong, narrower ante-
riorly, of the breadth of elytra posteriorly, with angles produced,
acuminate. Tarsi slender.
Spring-heetlet. When these animals are touched, they retract their feet
considerably. Since the feet are very short, these beetles, when laid on
their back, are not able to reach the groimd, and so could not right them-
selves again, if they could not by throwing themselves upwards attempt to
regain their feet. The animal, lying on its back, bends the thorax towards
the back or downwards, and thus causes the style of the prothorax to
emerge from the cavity in which it lies when at rest. Then bending the
body suddenly, so that the point again shoots into the cavity, the insect
with the back part of the thorax forcibly strikes the ground, and is thrown
upwards by the rebound.
The larvie live under the bark of trees, in decaying wood, or under
ground, where they gnaw roots. They have an elongate cylindrical body
with six feet and a very hard homy skin.
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526 CLASS viii.
Caanpylus Fisch., Exophthaimua Latb. Head behind the eyes
ezsert from thorax. Eyes large, globose. Body elongate, almost
linear.
Sp. Campylut linearU, Elaitr Unearii L. (and Slater fMiomdu9 ejnad.),
Fakzbb DeuUdU. /tu. Heft 8, Tab. ii.
Plater nob. Head reoeived in thorax as fiir as the posterior
margin of the eye&
Sub-genera: Ludiui Latb., Ctenieenu Latb., and many others proposed
by EsoHSOHOi/rz and Stephens. Gomp. Wxstwood Modem CUumfic,
Oeneric Synoptia, pp. 35 — 37, and Gebxab ZeUeehr, /. die Bntom. tL
pp. 141—378, iv. pp. 43—108* V- pp. 133— 19«» *c-
Sp. EUOer mwinus L., YoET Col, i. Tab. 44, fig. ^6\—EUA. ceneue L.,
YoET ibid. fig. 14 ; Elater aierrimus L., Panzeb Deutschl. Ins. Heft loi. Tab.
15, ftc The larvse of EUOer (Agriote$ Esohsoh.) aegetie, Slater Uneattu L.,
and other similarly- thin and elongated species are sometimes extremely
injurious ; they gnaw the roots of grasses and oom. See on these toireworme
KiBBT and Spenob Introd. to SiUom. I. pp. 180, 181, and Westwood,
Modem Classif. I. p. 338, and the authors there cited.
Amongst the numerous exotic species we note only Slater (^Pyropkorue
Illiq.) noctUuctu L., Yoet Coleopt. i. Tab. 43, fig. 16; it has on each
side of the thorax a smooth yellow spot ; by night it emits a clear lights
which radiates principally ^m the spots on the thorax (oomp. TBBTiBAinjB
BioilogU, V. s. 100 — 103, 475). This, with other phosphoreeoent species (see
KiBBT and Spenob Introd. n. pp. 410 — 412), is found in South America.
Gomp. Gebmab Zeittckr. /. Sntomol. in. 184 1, s. i — 76, J. Oubtib m
Zoai. Journal, ill. 1838, pp. 379 — 383.
[A fine specimen of Slater nocHlucue, which had been imported in acaigo
of logwood, came into Van deb Hcxvbn's possession after it had lived some
weeks in other hands. He kept it alire for a fortnight. The phosphorescent
spots were, in the dark, first blue, then emerald green, then incandescent.
Large printing could be read by the light, but it was necessary to cany the
insect along the lines to read continuously. In oxygen gas the luminosity
was not more splendid. Experiments to determine whether the tempera-
ture rose with the phosphoresoenoe were interrupted by the death of the
animaL On dissection, nothing peculiar was observed beneath the spots on
the thorax, except large or smaller masses of the adipose body connected
l>y numerous tracheee. The Professor thinks it not impossible that the
production of light may not be confined to a particular region, but may
occur wherever the adipose body occupies the spaces between the organs in
sufficient mass, and is adequately supplied with air-tubes. The two spots
on the thorax, though destitute of the brown colouring matter, are of the
same thickness as the neighbouring integument. Does this thickness, the
Professor asks, account for the transmisnon of the green light, whilst that
which proceeds occasionally from parts of the inferior surface, where the
membrane is thin, is yellow t However this may be, the adipose body must
be very unequally phosphorescent, otherwise the light from the under
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INSECTA. 627
BorfiMM would be always as strong as that from the upper. But it is
produced only rarely, and when the animal makes much exertion. See
Van dkb Hovbn in the Album der Natuwr, 1855, 7« Afley. bL 105.]
Cerophytum Latr.
Thrascua Latr.
Etumemis Ahrens.
Gomp. EucnemU, nuedorum ffentu, monographiee traOahtm iconibuaque
iUuatratwn a Banme Db Maknerbkiv, Petropoli, 18^3. 8ya recua. in
Afm, dei 8e. natur, ni. 1824, pp. 426—434. PL 17.
Galba Latr.
Melasis Oliy. Last joint of palps large, globose. Antennse of
males pectinate. Tarsi slender.
Note, — ^We omit here sereral sub-genera^ to be inserted between the
k Mater and Bupreitia,
Buprestis L. Antennaa inserted between the anterior part of
eyes, serrate, with short teeth. Mandibles short, triangular, with
apex simple. Palps filiform. Head retracted within thorax nearly
as far as the posterior margin of eyes. Thorax trapeziform, broader
posteriorly. Scutellum small or none. Tarsi with first four joints
dilated, bilobed.
Compare Solub Euoi mr lee BupretUdee, Aim, de la 8oe. entom. n. pp.
«6i--3i6.
Sp. Bvprettie gigamtea L., Voir Ooleopl. i. Tab. 48, fig. i, Sulzxb Kenm,
der Ine, Tab. vi. fig. 38 ; 2" 3'" long ; green copper-coloured ; the feet
steel-blue ; the shield-coverB terminating in two points ; South America.
— Bupr, futminans Fabb., i" 10"' long, green shining, the elytra striped
longitudinally with fine points, with ten fine teeth at the apex as though
serrated, and a red copper-coloured spot; very common in Java. In
southern Africa many species are found, which are b^et with little bundles
of hairs on the shield-coyers, as Bupreatit fatcicularie L. In Europe,
of this numerous genus ^ there are met with comparatiyely only few and
smaller species. Hie larvae live in bark and wood of trees ; they have no
feet» are elongate, flat and distinguished by the breadth and size of the
first ring of the body next the head.
Sub-genera : Trachya Fabb., Aphenisticua Latr., Buprestia Latb.
{AgrHus Megeble and many sub-genera proposed by Eschsgoboltz,
SoLiEB, De la Pobte, and Goby, kc)
1 DUFONOHXT announced a few years ago thaV he had found in different cabinets
more than 1500 spedes of this genus, — Diet, umiv. ^jffid, not. n. p. 779.
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528 • CLASS VIII.
Family LIX. Brachelytra {Microptera Gravenhorst). An-
tennae mostly moniliform, of the same thickness towards the apex,
or thicker at the apex. Elytra much shorter than abdomen, en-
tirely covering the wings which are long and replicate, not dehis-
cent. Body mostly narrow, elongate, arcuate upwards, the insect
running. Dorsal segments of abdomen all homy.
Most of these insects liye in offal, dung, or imder decaying plants.
They live on animal substances, especially other insects ; so also the
larvsB, which are elongate, with a large head and six strong feet.
The intestinal canal of these beetles is straight and short ; there are
four vcLsa urinaricL The similarity of these beetles with the genus
Forficula amongst the Orthoptera (see above, p. 462, 463) is rather
external and does not rest upon a natural affinity in the internal
structure.
The species of this family are very numerous, and in Europe fur
surpass even those of the CcurahicL However, this is quite other-
wise with the exotic species hitherto known, and hence some writers
are of opinion that this &mily \& eminently a form of the northern
hemisphere. Perhaps this may be a hasty conclusion, and the
rarity of StaphyUni from warm countries, in our coUections, may in
part be attributed to the inadvertence of travelling collectors.
Compare J. L. G. Gbavknhobst CoUoptera microptera, Bronsyigs, 1802,
Sto.; 'Eivmd. Af(mogr<tphiacoUopteror. mierapterorwa, Grottingie, 1806, 8 vo.;
Maknabhsim in Mint. presenUs d VAcad, des Sc. des Pdertbowrg, i. 185 1,
pp. 4 T 5— 461 ; G. F. Ebiohbon Qtniera et Sptcia SUtphyUnoruai, cum tab.
cen, V. Btiroliui, 1840, Svo.
Section I. Staphylinina, Abdomen with six or seven segments
distinct. Cutaneous papilla exsertile, pilose, on each side near
anus in both sexes.
Stapkyltnus L. (in part), Gravenhokst. Head entirely exsert,
separated from thorax by a narrower base as by a neck. Labrum
mostly bilobed. Palps mostly filiform, labial triarticulate, maxil-
lary quadriarticulate, with last joint in the former sub-equal. An-
tennae inserted in the anterior margin of frons, with eleven joints,
filiform, or in some incrassated towards the apex. Tarsi with five
joints.
Sub-genenL a) With antennee approximate at the base : PhUy-
prosopus Maitoebh., Othius Leach, HoUsua Erichson, Sterculia
Laporte, XanthMnua Dahl., &c.
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INSECTA. 629
b) With antennse distant at the base: StaphyUwus Ebichs.,
Ocyptis KisBY, FhUonthus Leach, Aeyhphorua Nordmaitn, Quedius
Leach, AatrapcBUs Gbay., Oxyporus Fabb., && Comp. Ebichsok,
LI. pp. 291— 559.
Sp. Staphylinua eryihroptenis Fabb., Staphyl. ccnareua Gbdeshjblm, Ebiohs.,
Fanzsb DevtHM. Tim. Heft 76, Tab. 4, Dum^bil Cons, gin, «. I. Ins, PI. 3,
fig. I ; 8'" to g"' long, black, with short, yellow, glisteriiig hain, shield-
coyera and feet red-brown, antennie yellow-brown, at the extremity mostly
blackish ; — Staphyl. ccutanopterus Gbat., Staph, erythroptertu L. &c. The
largest native species is Staphylinus oUns Fabb., Ocypus olens, Panzeb
1. L Tab. I, Batzbbubo Font-Ins, 1, Tab. i. fig. 14; 10'" to i" long, quite
black. Of this species the larva and nymph are figured by Blanchabd in
GuiBiK Magas, de Zod. 1836, Ins, PI. 165, and more accurately by
Oew. Hbbb {Observ, Entomol. Turici, 1836, 8vo. Tab. m. a) and Ratzb-
BUBa loc. dt.
Pcederua Fabb. Head exsert, aflSxed to thorax hj a slender
neck. Labrom with sides often membranous, bilobed, or slightly
emarginate in the middle. Maxillary palps elongate, with fourth
joint very small; labial palps short, triarticulate. Antennae inserted
tmder the margin of frons, with eleven joints, filiform. Tarsi with
five joints.
a) Fourth joint of tarsi simple.
Laihfrobivm Gravenh., Stilicus Latb., Ac
b) Fourth joint of tarsi bilobed or provided with a membranous lobe.
Pcederua Gbavenh., Swnius Leach.
On these and other sub-genera compare Ebiohboh L L pp. 560—667.
8p. Pcederus riparius Fabb., Panzeb DmUschl. Ins. Heft 9, Tab. 11 ; —
PoBderus ruficoUds Fabb,,, Panzeb DeutsM. Ins. Heft 37, Tab. 13, Gu^in
Iconogr. Ins. PI. 9, fig. 5, &c.
Prodmia Latb.
Pinophilus Gravenh.
On these and other genera and sub-genera, composing the tribe of the
Pinophilmi, consult Ebiohbon L L pp. 668 — 686.
Stenus Latr. Head exsert, inserted in thorax by a thick neck,
with eyes large, protuberant. Labrnm entire, homy. Labium
elongate, exsertile. Maxillary palps elongate, with fourth joint very
small; labial palps short, triarticulate. Antennas short, inserted
VOL. I. 34
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630 CLASS vin.
in the frons between the eyes, with eleven joints, last three joints
incrassated. Feet slender.
A. Tarsi with five joints. Labram entire.
Sub-genera : Dianoua Leach, Stenvs Latb.
B. Tarsi with four joints. Labrum denticulate or serrate.
Ev4JLe8the(ni8 Gravekh.
Sp. Stenm UgiMahu, StaphyUnua InguUahu L., Dum^ril Cam. g&n. «. 2.
In9. PL 3, fig. 4, Panzeb BeuUchl. Ins, Heft ii, Tab. 17, &c.
Compare Thion, Descrifivm des organes de la manducaiion ckez Ut
SUnes, Ann, de Ut Soe, EtUom. iv. 1835, pp. 153—168^ PL 3 B.
Oxytelm Gravenh. Head exsert, with base narrower. Labrum
entire, homy, mostly enlarged at the apex on both sides by a
membranous lacinia. Maxillary palps much shorter than head, with
fourth joint distinct, often subulate. Anterior tibiaB mostly spinose.
Head in many homed. Tarsi with last joint very long.
A. Tarsi with fire joints.
AfegcUops D£r. Antennse with ten joints, davate.
Osorivs Leach. Antennse with eleven joints, filiform.
B. Tarsi with three joints. Antennae with eleven joints.
Sub-genera : Oxytdus Grav. (in part), Bledius Leach, Platysthetua
Mannerh., GoprophUti8 Latr. and some others.
Compare Latbbillb Conndiraitions war la, irihvk de» DenUcruret, Nauv.
Ann. du MuUum, i. 1833, pp. 77 — 93 ; Ebiohson L L pp. 759 — 833.
Sp. OoB^ui triewnU Gbayekh., Panzeb DeuUchl. Im, Heft 66, Tab. 17 ; —
OayUiL piceus. Staph, piceut L., Panzeb DeuUchl. Int. Heft 17, Tab. 12 ;
Oaa/tdut {CoprophiUut) rttffoitu, GuiBiN Iconogr, Ins, PL x. fig. 1, Gbbmas
Faun, Ins. Europ. Fasc. XT. Tab. 3, &c.
Piestua Gravenh.
Prognatha Latr. {Siaganium Kirby), Leptachirus Germar, <&a
Compare Ebichsoi? L L pp. 813 — 841.
Omalium Gravenh. Head porrect, scarcely constricted at the
base. Labrum transverse, homy, simple or supplied with a mem-
branous border at the apex. Antennse with eleven joints, straight,
filiform. Elytra mostly longer than pectus, sometimes covering
a large part of abdomen. Tarsi with five joints. Body depressed.
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IN8ECTA, 531
A. Ocelli none.
Sub-genera : Olisthwrua Dej., PhUxoehania Mannerh.
B. Ocelli two.
Sub-genera: Anthophagvs Gray., Lesteva Latb., OmaUum
Gbayenh. (in part), Anihohivan Leach, and some others proposed by
Erichsok and others. Comp. Erichsok, L L pp. 846 — 900.
Sp. OmaUum rivuUtre Qkayvsk., Panzxb J>etUBckL In$, Heft S7, Tab. 15 ;
— Omaliuiii fattuUUum Ebiohs., Omal. ecBtum Gtll. ftc.
Prateinua Latil Head porrect, received posteriorly in thorax.
Antennse clavate or incrassated at the apex, mostly with eleven
joints, BometimeB with nine. Maxillary palps with fourth joint
narrower, acuminate. Ocelli none, or ocellus single, frontal. Body
depressed ; elytra longish. Tarsi mostly with five joints, in a few
with three joints.
Sub-genera: Prateinus Latb. (in part)^ Megarthms Kibby,
Phlcaobivm Dejean, Glyptoma Ebichs., Mioropeplus Latb.
Sp. Proteinui brachyptenu, Dermestn brachypienu Fabb., Cuvikb JL Am.,
id, iU., Im. PI. 31, fig. 8 ; Megarthrut dqtremu Ebiohb. kc
AleocJiara Grayenh. Head small, exsert, often deflected.
Labrum entire, homy. Maxillary palps with fourth joint subulate.
Antennae with eleven or ten joints, inserted in frons at the internal
margin of eyes, incrassated gradually at the apex, or clavate. Feet
slender, with tibias mostly unarmed, tarsi with five joints, some-
times with four, either the anterior only or all.
Sub-genera: Myrmedonia Ebichs., Aleockara Gbayenh. (in part),
BoUtachara Mankebh., Oxypoda Mannebh., Oligota Mannebh.,
Lomechuaa Gbayenh., Gyrophcena Mannebh., MyUasna Ebichs.,
Gymnriaa Kabsten, && Cons. Ebichson, L L, pp. 33 — 212.
8p. Aleochara canalictdala Gbay., Staphyl, canalieulatui Fabb., Pakzeb
DeuttcU, Int. Heft 37, Tab. 10, Gu:foiir Iconogr., Int. PL 10, fig. 5 ; —
Aleoch. futcipet Gbay., Gbbmab Faun. Int. Europ. Fasc. xui. Tab. 8, &c.
Tachypoma Grayenh. Head intruded posteriorly into thorax
almost as far as the eyes, not narrower at the base. Labrum entire,
simple. AntennsB below the eyes, inserted under the' lateral margin
of head, mostly filiform, with eleven joints, in a few clavate, with ten
34—2
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532 CLASS VIII.
joints. Feet with tibiae mostly spinose, tarsi with five joints, more
rarely with four. Body convex, ovaJ,
Sub-genera: HypocypbM Schuepp., Tackyporus Gbaveit., TachinuM
Gravenh., BoletMvs Leach, &c. Comp. Erichsok, pp. 213 — 289.
Sp. TackypoTU$ analu Gkav., SiapkyL oUuiut L., Pahzsb DeutaekL In*,
Heft 17, Tab. i6 ^—Taehyponu marginatut Gbat., Stapkyl. hypMnem
Fabb., Pakzsb Deuttehl. Int, Heft 37, Tab. 17, Gu^BXV leomogr.. Int.
PL f o, fig. 9, fto.
Section 11. Pselaphina. Abdomen with five segments, rotun-
date, obtuse. AntennsB clavate. Tarsi triarticulate.
Compare C. Aub^ PtdapAiorum Mcnographia, Gvfisnr, Moffotin de
Zool. 1833, Int. PL 78 — 94; H. H. Sohmtot Ditt. inamg. tool, de
Ptdapkit Fauna Pragentit cum anatamia Clavigeri, Pnge, 1836. Syo.
Thffy are smaU inaecto that live under stoneaj baric of troea, decaying
leavea, and in ante* nests.
A. AnUnna with Joinit indutind, or with tingle joint, thuk, tnmoaied.
ArticeruB Dalm.
B. AnlewMB with tix joinit.
Claviger Pretssler. Eyes none (ocelli two Schmidt). Claw
of tarsi simple.
Sp. Claviger teitocettf-PANZBB DevUchl. Int. Heft 9, Tab. 3, DuMiaiL Cont.
gin. 1. 1. Int. PL 11, fig. 5. Compare P. W. J. Mubllbb BeiMige turNtUur-
getch, der OaUung Claviger, Gbbmab's Magazin der JSwUmol, m. 18 18,
pp. 69— in.
C. AnienwB with eleven jointt.
Pselaphua Herbst. Maxillary palps mostly elongate, clavate.
Eyes two.
Sub-genera, a) With single claw of tarsi : PsdaphuSy Herbst
(in part), BryaxU Leaoh, Bythinua, Tychua ejusd, Euplectua ejuad.
— 5) With two claws of tarsi : Chennium Late., Tyrtia Aub^
^o^muaAuBi; whom see LL Comp. also Erichsok, 2)t6 JTq/er dar
Mark Brcmdmbwg, 1. pp 263 — 281.
Sp. C^£nmum hitubereulatum, DmciaiL Cont. gin. t, I. Int. PL 33, fig. 4, &c.
Family LX. Glavicomia {Glavicames Latr., Hiateridm^ Byr^
rhtdcSf JDermestidcB, SilphidoB^ &c. Leach). Antennas mostly with
eleven joints, terminated by a club perfoliate or solid, or gradually
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IN8ECTA. 533
incrassated towards the extremity, longer than maxillary palps,
with base little covered or quite naked. Integuments hard. Elytra
sometimes abbreviate, in all however covering the greatest part of
abdomen. Joints of tarsi mostly entire.
Scydmcenus Latr. Maxillary palps elongate, with fourth joint
subulate, small or very small, scarcely distinguishable. Antennss
moniliform, gradually incrassated towards the apex, or clavate.
Elytra mostly solute. Head exsert, mostly constricted posteriorly.
Sp. ScydfMBWM HeOmigii Latb., Nokmu mimiltu» Panzxb Deutadd. Ins,
Heft 13, Tsb. 6 ; Sqfdm, Oodatti Latr. Qtner. Orutt. et Im, Tab. vin.
fig. 6, &c. SeyenJ species are described and figured by SruSM Jkuttchi.
Famna, Im. xiu. pp. i— 51, Tab. ^59^964.
Clidtus Laporte.
Mastifftis Illig. Antennse filiform, scarcely incrassated at the
apex, with first joint very long. Maxillary palps elongate, clavate.
Elytra connate, wings none. Head exsert, constricted posteriorly.
Sp. MaatiguM pcdpalis Latr. 1. 1. fig. 5, Guv. M. Am,, id UL, Ins, PI. 34, fig.
10. For other species see Kluq EnJtomolog. Monogr. pp. 161 — 168.
Hister L. AnteimsB short, geniculate, with first joint elongate,
clavate, with capitulum oval or subglobose, solid, triarticulate. Man-
dibles arcuate at the apex, denticulate, mostly exsert. Maxill» with
two membranous lobes. Palps filiform. Head small, mostly re-
tractile within thorax. Feet contractile, anterior approximate at the
base, posterior remote at the insertion, with tarsi all five-jointed.
Elytra shorter than abdomen, truncated. Body smooth, mostly
subquadrate. Abdomen composed of five segments.
a) With head not retractile.
Hololepta Path.
b) With head retractile.
Sub-genera: Saprirma ERiCHa, Ahrasus Leach, Onthophilus
Leach, HUter Leach, Plalysoma Leach, kc
Sp. Hkter unicolor L., Sturm Deufschl, Int. 1. IVtb. 16 ;-— Hister macuUOus
L., Ac.
Necraphonta Fabr. Antennae short, clavate, with capitulum
subglobose. Maxillae unarmed. Palps short, filiform. Ligula
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534 CLASS viii.
bifid, with lacinias divergent, intemallj ciliated. Elytra shorter
than abdomen, troncated. Anterior tarsi broad, famished with
fasciculi of hairs,
Sp. Necrophortu veapUlo Vabh., SUpha vetpiUo L., Bcbsel /m. iv. Tab. i,
Pakzeb DeutscM, Iru. Heft a. Tab. 21 ; iJu grove-dig^ ; 6"'— 8"' long,
black, elytra with two orange-coloured transvene bands, the dub of an-
tenns brown. This beetle flies with elytra erect, the under-side turned
outwards ; it diffuses a strong smell resembling musk. Like Qtatrupei
ttereorctriuB and vemaUs it is sometimes infested by small acari. The
periods of larva and nymph together extend only to seven weeks. This
species, Uke the others of this genus, feeds on dead animal substances. By
an union of their forces these beetles dig away the ground from under the
carcases of small animaLs, moles, mice, &c. and so bury them beneath it.
They lay their eggs in the bodies thus interred, and the larvs feed on them.
— Necrophorua humaior Fabjl, Pakzeb DeuUM, In$, Heft 41, Tab. a, &a
Silpha L. (exclusive of some species, Fabr.) Antenn» longer
than head, with club elongate or gradually thicker towards the
apex. Maxillae armed internally with a homy hook. Head small.
Thorax broad, plane, marginate.
Sub-genera: Necrodea Wilkik, Oiceoptoma Leach, ThanatopkUut
Leach, Fhosphaga Leach.
Sp. Silpha UUoraUt L. {Necrodet), Sulzeb Otich, d. Int, Tab. n. fig. 14,
(SUpka clavipes), Panzbb DetOtehZ, Int, Heft 40, Tab. 15 ; — Silpha quadri-
punctata L., VoBT Coleopt. i. Tab. 4T, ^. 5, Panzbb DeiUachl, Int. Heft
40, Tab. 18. This beetle hunts after different kinds of caterpillars on oaks
and other trees.
NecrophUtu Latb. Mazilke unarmed, with two membranous
laciniae. Palps very short, thickisL Thorax broad, exsert an-
teriorly.
Sp. SUpha iubterrunea Illio., Sturv DeuttM, Fauna, Int. xm. Tab. ^70.
Agyrtes Froelich.
Consult Fboilioh Nahwf&ncher, xxvm. 1799, ■• '5— '9» Tab. i,
fig. n.
Catopa Fabr.
Choleva Latb., Spence.
Consult Monography of this genus by SPBiroB in Tramact. nf iht Lwm,
Soe, XI. 18 15, p. ia3--i6o,
Sp. Catopt chryBomdaUde; HdopB ehryaomdoidei, Paitzbb DeutaeKL Ins, Heft
57, Tab. I, ftc., Stubx Deutschl, Fauna, Int, xrr. Tab. 375, fig. b, B.
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INSECTA. 535
CoUm Hebbst.
Consult Stubm DetOschL Fauna, Ins. ziv. pp. 46 — 74.
8caj>htdmm Oliv. Antennae straight, moderate, thicker at the
apex, mostly provided with a club of five joints. Mandibles small,
homy, with border membranous internally, unidentate or bidentate.
Maxillae with two membranous lacinise. Maxillary palps quadri-
articulate, with last joint acuminate at the apex. Labial palps
very short. Body navicular, at both ends angustato-acuminate,
with head small, deflected. Elytra truncated at the apex. Feet
slender.
Sub-genera: Scaphidiwm Leach, Scaphium Kibbt, Amalocera
Erichson, Scaphtsoma Leach.
Sp. Seaphidium agaricifwm, SUpha offorieina li,, Fanzeb DeuUehl, Ins,
Heft 1, Tab. 3, &o.
Nittdula Fabr. Antennae straight, clavate, club mostly triarti-
culate. Labrum distinct. Mandibles partly exsert, emarginate or
bidentate at the apex. Maxillae mostly with single membranous
lacinia, the external lacinia obsolete. Palps short, filiform. Head
retracted within thorax almost as far as the eyes. Elytra often ab-
breviate. Tarsi with fourth joint very small, subindistinct.
Cercus LATa, Brachypterua Kugel., Ga/rpophUua Leach, NUidtila
ItATR., {Ipidia, EpurcBa, NUidula, Soronia, Amphotis, OmosUa
Erichs., Fria Kirby, Meligethes Kirby), Thalicra Erichs., Pooo-
dius Erichs., Cychrcmus Kugel. {Strongylua Herbst), Cyhocephalua
Erichs., CyUodes ejusd.
Compare Erichson, Gbbmab'b Zeiiseh, f. d, EnUm. iv. 1843, p. «i5,
&c. T. 1844, p. 438 &c. ; Eriohson's Nalurgemik. d. Int. DeutacU, lU.
pp. 135 — 210.
Sp. Nitidula hipuetulata Fabb., SUpha Inpugtulata L,, Panzeb, Deutschl, Im.
Heft 3, Tab. 10, SruBif DeuUchl, Fawna, Ins. XT. Tab. 398, &c.
Ip8, Fabr. Labrum membranous, concealed under the clypeus,
porrect between the mandibles. Maxillad with single lacinia. Tarsi
as in the preceding genus.
Bhizophagfia Herbst.
PeUis Geoffr., Thymalus Latr. Antennae with club triarticu-
late. Maxillae with two laciniae, the interior armed with a homy
hook. Body broad, depressed, with thorax and elytra marginate.
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586 CLASS YIII.
Trogosita Oliv., Fabr., Latr« Antennae with eleven joints,
moniliform, subclavate. Mandibles exsert, porrect, with apex
bidentate. Internal lacinia of maxillae obsolete. Palps filiform,
incrassated at the apex. First joint of tarsi small. Body elongate,
depressed.
Sp. TrogotUa maurUanica, Tenebrio maurUaniau L., Pahzkb DeuUcKl. Ins.
Heft 3, Tab. 4, S^dbm DeuUchL Fauna, Ins. 11. Tab. 48.
TenmockUa Erichs.
Nemoscma Latr.
Golydium Fabr. Antennae short, with club triarticulate. Max-
illary palps with last joint larger, subsecuriform. Tarsi quadri-
articulate. Body elongate, cylindrical.
NqU. — ^Here also ought to be placed, according to Eriohson, genera
Sa^rrctrium and CorHcua classed with the Tendtricnet (see aboye p. 496), and
Coxdui, placed amongst the Diaperiales, p. 494.
Rkysodea Illig.
Consult Gebmab PoMn, Ins, Europ. Fasc. vi. Tab. i, where the charac-
ters are set forth.
Passandra Dalm.
Prostomia Latr.
Sp. TrogosUa mandibularis Fabb., Stubx DeuisckL Fauna, Ins, n. Tab. 49,
Panzbb Deutschl, Ins. Heft 105, Tbb. 3.
Cucujits Fabr. Antennae filiform. Labrum small. Mandibles
exsert. Maxillas bilobed, with internal lacinia small, acuminate.
Palps short, filiform. Feet short, with first joint of tarsi mostly
very small. (Posterior tarsi in males with four joints.) Body
elongate, depressed.
Uleiota Latr. {Brontes Fabr. in part). Antenna of the length of
body, or longer than body.
Sp. Oucujus Jtaviceps Fabb., Cerambyx planatus L., Pahzxb Deutschl. Ins.
Heft 95, Tab. 4, Dumi^il, Cons. gin. a. L Ins. PI 17, ^. 7, &c.
Gryptophagus Herbst {Dacne Latr. in part). Antennse with
eleven joints, clavate. Mandibles short, strong. MaxillsB bilobed.
Palps short, incrassated towards the e2:tremity. All the tarsi in
both sexes pentameious, or posterior in males tetramerons.
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INSECTA. 537
Anierophagtu Kkoch, Emphylus Ebighb., Pcvrameeo9oma Oubtis,
Atomaria ElnffiT, £piHemu8 Wkstw., Ebichs.
Sp. Crypiophagw crenahu Hsrbst, DermetUt cellarii SooPOU, Stubm
DeuUchl. Jn$. ZYI. Tab. 513, fig. d.
Stlvarms Latk.
Mycetophagtia Fabr., Oliv. Antennae gradually thicker to-
wards the apex or clavate. Labrom distinct, homy, coyering man-
dibles. Mandibles with apex bidentate. Palps filiform, labial
short, maxillary longer, incrassate. Tarsi quadriarticulate, anterior
of male triarticiilate.
Sp. Mjfcdophagut piceut Fasbl., Myeetoph, variMlia QiLL., Pahzib Ihutiehl.
Ins, Heft I, Tab. 72, &c.
Add genera Tripht/Uue Meg., Typhcea Kiabt, and some others
here omitted.
Ptilium ScHUEPPEL, Trichopteryx KiRBY. Antennae straight,
with eleven joints, the three terminal joints incrassated. Tarsi
triarticnlate ; seta, ftimished with terminal pulvillus, placed be-
tween the claws. Wings narrow, lanceolate, ciliated at the margin.
6p. Ptilium fcudcularey Trichopt, grandicollis EuoHS., Stubx DmUtchl, In:
XVII. Tab. 313, fig. I, &c.
Small insects (scarcely more than \[*' long), living in ant-hills under
decaying leaves, the bark of trees, &c. with elytra often abbreviate, tmn-
cated. Compare Gillmeistkb, Stubm DeuUchl. Int. zvn. 1845, Ebiohboh
NtUursfeach. d. Ins, DeuUchl. ni. pp. 13 — 40.
Dermestes L. Antennae frontal, straight, short, clavate, with
club mostly triarticnlate. Mandibles short; maxillae with two
laciniae, mostly membranous. Palps short, incrassated. Tarsi
with five joints. Abdomen with five ventral segments, free. Feet
contractile under the body. Elytra entirely covering abdomen.
Sp. DermetUs lardariut L., Blankaabt Sehoubu^, Tab. zi. p. 96, figs. K,
L, u, DuiciBiL Cons. g&n. s. L Ins. PI. 6, fig. 10 ; z'" long, black, with a
grey, transverse band at the base of the shield-covers ; very common in
honses. Derm, muritms L., Panzkb Deutschl. Ins. Heft 40, Tab. 10, kc.
These insects in the larval and perfect states feed on all kinds of animal
substances : leather, bacon, &c.
I^ote. — The remaining sub-genera, composing the tribe of the
DermegUni, are furnished with a single frontal ocellus, which is
absent in DermaHes.
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538 CLASS YIU.
AUagemts Latb. {Jfegaioma ejiud), Megaiama Hebbst, Trogo-
derma Latb., Timias Stbph., AnihrenMU Gboffb. (ByrrkuM L.
ezclusiye of species), Trinode8 Meg.
Sp. AiUkrenMi WMueorum, JkrmaUt (and ^yrrfttif) mmteonm L., Pavssr
JhuUckL Int. Heft loo. Tab. i, ^tubm DetamJU. Im. n. Tab. 37. The
lanra is haiiy ; paiiicnlarij then are bandies of long hain at the posterior
extremity, with dart-shaped points Teiy elegant nnder the microecope.
Byrrhus anct. (species of Byrrhus L). Antennae incrassated
towards the apex, or clavate, with club distinctly articulate. Man-
dibles denticulate. Maxillae with two laciniae. Palps incrassated at
the apex, truncated. Head small, received in thorax. Feet con-
tractile, with tarsi replicate towards the tibiae, and tibiae upon the
femora. Tarsi with five joints. Body gibbous.
Sp. B^rrhm pHida'Ij., Panzbb DetdMchl. Int. Heft 4, Tab. 3, SruBM
DevBttehl. Ins. n. Tab. 33, Ac.
Cj/tUtu Ebichs. Palps with last joint acuminate. Antennae
clavate.
Sp. Byrrhnt variui Fabb., Pahzkb DentsehL Int. Heft 31, Tab. 3.
Lvmmchua Ziegl.
No9odendron Latb.
Agathidium Illig. Antennae with eleven joints, clavate. Man-
dibles strong, arcuate. Maxillae with two lobes. Palps filiform.
Head nutant. Abdomen with six ventral segments. Elytra en-
tirely covering abdomen. Feet somewhat slender, with tarsi of
posterior feet in males, sometimes also of middle or of all the feet in
females, quadriarticulate. Body contractile into a ball.
Compare on this genus and its division Ebichsoh Naturgesch, d. Int.
BtvUchl. m. pp. 87—134, SrruBM Deutschl. Int. 11. s. 53, &o. Small
ooleoptera living in fiingi and decaying trees.
Anisotoma Knogh, Illig.
Note. — ^A genus much resembling the preceding, bat with body not
contractile into a baU. Feet stronger, with tibie spinose. AH the tarsi
in some five-jointed, in others the posterior or middle foor-jointed, or
anterior four-jointed, middle and posterior three-jointed, the nnmbo* how-
ever not differing with the sex. Ebiohbok 1. 1. p. 44, whom see on the
division of this genus.
Pausua L. AntennsB large, very broad, with somewhat few
joints, sometimes only with two or three, the last veiy large.
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INSECTA. 539
Palps large, exsert. Elytra truncated. Tarsi short, with first joint
minute. Body oblong, depressed. Abdomen broader than thorax.
Small ooleopten, with almost the habit of Jffitpa or A lurmu, exotic, often
met with, as it seems, in ants' nests. Taksmvb first founded the genus in
A. Dahl Biffis Iruectorum 1775, AmomU. Academ. viii. p. 307. Compare
Ajzklius TrofMouA. of ike Linn, Soc. YoL iv. p. 343, &c. and Wkst-
wooD ibid. Vol. ZTI. pp. 607—684, PI. 33, and his Arean. Enlom. ii.
pp. I— 13, 37—40, 73—80, 361—190.
A. Head supplied posteriorly with a neck.
Cerobpterus Swed. Antennse with ten joints, dub with nine
joints.
Compare N. S. Swedkbus in Vdentk. Ahad. nya Handb. 1788, p. 303.
PerUapkUarthrua Westw. Antennie with seven or six joints,
club elongate, broad, with five joints.
Pau8SU8 L. Antennse with two or three joints, club broad, exar-
ticulate.
Sp. Pau$»u8 mieroeephahu L., Amoen, Acad. vin. Tab. vi. figs. 6 — 10;
— Paussut thoracictu DoNOV., Pauu. irigonicomu Latb., Oen, cnut, et
Ins. I. Tab. xi. fig. 8, &c.
B. Head immersed in thorax.
Hyhtorvs Dalm. Antenn» with two or three joints, club
exarticulate, lanceolate, incurved, acuminate. Head furnished with
two conical tubercles (ocelli )).
Sp. HyUAoT, hwepkaluM Gtllenh. in Sohoehh. B^, Int, i. 3, App, Tab. Ti.
Heterocerus Bosc. Antennae inserted under the lateral margin
of firons, with eleven joints, first two large, broad, two following
small, rest transverse, dilated, forming an elongated club. Head
received within thorax almost to the eyes. Mandibles porrect, den-
ticulate. Palps filiform. Body oblong, depressed, pubescent.
Tibia, especially anterior, spinose. Tarsi quadriarticulate.
Sp. ffderooena marginatuB Fabb., Dum^il Com. gin. 9. I. Ins. PL 7, fig. 7,
Pakzib DetOsefd. Ina. Heft 93, Tab. I3, ke. Small insects, the laiig;est
species of which is scarcely more than 2'" long, and which in all their states
fire in passages near the edge of the water. On the internal structure
compare LioN BuFOUB Ann. de$ Se. natur. ae S^rie I. Zoolog. p. 60 and
foUowing, with figures.
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540 CLASS yui.
Dryopa Oliv., Pamus Fabe. Antennae frontal, very short,
with ten or eleven joints, anriculate, with second joint large, cover-
ing the rest. Mandibles not exsert. Maxillae with two laciniae,
the internal not broader than the external. Tarsi with five joints,
the last joint and claws strong. Bodj pubescent and pilose.
Helichus Ebichson, Dryopa Leach, Poflrygrus EaiCHa, Pamua
Fabr., Erichs.
Sp. Dryops aurieulatua Oliy., Latb., Pamui prUifericornia Fabb., Panisb
DetOtcM, Ins. Heft 13, Tab. i, Cuv. JL Ant,, id. ill., Ins. PL 37, fig. 5 ;
1^'" long^ black-brown, feet brown, coYered with a silky glistering felt :
this insect lives at the bottom of fresh water, amongst the roots of water-
plants.
Potamophilvs Germar, Hydera Latr.
Elmia Latr. Antennae mostly with eleven joints, filiform or
scarcely incrassated towards the apex. Maxillae with two homy
laciniae, the external narrow, palpiform. Feet elongate, tarsi with
five joints.
Sub-genera: Limniua Muell., Erichs., Stenelmia Leon Dufoub,
Macronychu8 Mtjell., Latr. Antemm sexarticulate, with last
joint oval, forming capitulum.
Sp. Macron, quadrituberctdatua, Gbbmab Faun. Ins. Ewr. z. Tab. 4, Qu^rin
Icon., Ins, PL 30, fig. 4.
Georissua Latr. Antennas short, with nine joints, clavate,
with club globose, triarticulate. Head nutant. Body globose,
apterous. Elytra connate. Tarsi with four joints.
Fig. Panzeb DeiUschZ. Ins. Heft 6a, Tab. 5, Gebmab Faun. Ins. Eur.
Fasc. r5, Tab. 3, GuiaiN Icon., Ins. PI. 10, fig. 5 &c. Small insects,
scarcely ^'" long, living on the banks of rivers and streams.
Family LXI. Palpicomia, AntennsB short, with six to nine
joints, inserted in a deep sinus under the lateral margin of head,
terminated by a club perfoliate or solid. Palps filiform, unequal,
labial shorter, maxillary nearly of the length of antennae or some-
times longer than antennas. Body gibbous, mostly hemispherical
or oval. Mentum large, clypeiform.
Sphcerulium Fabr. Maxillary palps nearly of the length of
antennse, with four joints, the first small, the second incrassated,
obconical. MaxillsB with two laciniaB membranous or coriaceous.
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INSECTA. 541
First joint of tarsi equalling or surpassing the second in length.
Antennae with nine or eight joints, the first elongate, the last three
incrassated, forming a club. Body ovate or orbicular.
Sub-genera : Cercyon Leach {Gercy<m and Trichopoda Brull^),
Sphceridium Fabr. (in part), Cyclonotvm D£r. {CcdosUyma Brull^).
Sp. Sphceridium acarabcBOldes Fabb., DermeHes tcarabceoidea L., Yoet Chleopt.
I, Tab. $2, fig. I, Panzeb Deutschl. Int. Heft 6, Tab. i, Stubh DeuttcU.
Faun., Ins, Tab. xxi. &c. These insects liye on the land in dung. Other
spedee live under wet moss, or in muddy ground.
Hydrophilua Geoffr., Fabr. Maxillary palps with second
joint not incrassated. Maxillae with two coriaceous laciniee. First
joint of tarsi very short, connate with the second. Antennae with
eight or nine joints. Body ovate. Thorax broader posteriorly.
A. WHk anteMUB of nine joinU.
I. With sternum caainate.
Hydrophilus Fabr. (in part), Hydrophilua and Hydrchiua Leach.
Sp. HifdrophtluM pioeua Fabb., Dytiscus pieeue h., Dum^il Oons. gSn. s. I,
Ins. PL 6, fig. 9, Panzkb Deutschl. Ins. Heft no, Tab. 12 ; the largest
species of this genus 1} to 1} inches long, pitch-black, shining, the
antenna and feelers red-brown ; the thorax terminating in a sharp point
between the hind-feet This beetle lives in fresh water, and feeds especiaUj
as perfect insect, yet also as larva, on water-slugs, and swims very quickly.
The abdomen is covered beneath with a stratum of air, as though with
silver. This air (a provision for respiration) is renewed by the beetle, by
means of air-bubbles taken up on his antenns, which he brings from time
to time to the surface of the water (Nitzsoh in Rsil'b Arckiv f. d.
Physiol, z. p. 440). The female spins for her eggs an oblong, pod-shaped
nest of a ydlow-brown colour, terminating above in a little curved horn,
which floats on the water amongst the duckweed. In this the eggs lie in
regular order. The young larvse, some time after they have come forth
perforate this web, and cast themselves into the water. See Ltonet in his
translation of Lbsseb, ThkHogie des Ins. I. p. 159, fig. zvi. and Ouvrage
posthwne, pp. 133 — 151, PI. 13 ; compare also Miokb, Ann. du Mus.
d^Sist, not, XIV. pp. 441—459, PI. 28.^Bydrophilus scarabaoUies Fabb.,
{BydrMus) Pakzbb Deutschl, Ins. Heft 67, Tab. 11, &c.
3. With sterwum simple.
CyUidium Erighson.
B. Wilh antewncB of eight joints,
Berosvs, Laccobius, Limr^ebius Leaoh.
Helophcyrus Illig. {Elophorua Fabr.). Maxillae with two cori-
aceoiis lacinisB. Maxillary palps with second joint not incrassated.
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542 CLASS VIII.
First joint of tarsi veiy short, strictly connate with second, last
elongate. Antennae with seven to nine joints. Body oblong. Thorax
narrowed towards the base.
Sub-genera: HydrcBna Kuo., 0ehikebiu8 Leach, Eydrochui
Gesicab, Hdophonis Iluo.
Sp. Hdoph, ffrandis Illio., JSloph. aquaUctu Fabb., Dum^bil Cons. ffSn, s. I.
Int. PI. 6, ^. S, Sturm DeuUehX, Faun., In$. z. Tab. 119, fig. A &c.
Spercheus Fabr. Maxilla with external lacinia homy, palpi-
form, acuminate, incurved, ciliated internally, with internal lacinia
coriaceous, pilose at the apex. First joint of tarsi not less than
second, last elongate, equalling the first four in length. Antennae
short, thick, sex-articulate.
Sp. Sperckem emarginahu Fabb., Panzbb DetUmM. Ins. Heft 91, Tab. 4,
Stdbh L 1. IX. Tab. 114, Cuv. R. Ani, id, iU., Int. PI. 38, fig. 4; 3"
long, dull brown, very convex on the back ; head-shield excised in finmt :
this little beetle creeps upon the roots of water-plants.
Tbu genus forms the transition to the following family, and from the fonn
of the under-jaws belongs to the neighbourhood of Gyriaut.
Family LXII. HydrocarUharina or Dytiadna. Maxillas homy,
with external lacinia palpiform, articulate. Mandibles, as to the
greatest part, covered. Four posterior feet natatory, two posterior
remote from the rest. Body oval , thorax broad, transverse.
A. Abdomen composed of six segments.
OyrinuB Geoffr., L. AntennsB shorter than head, fusiform,
with nine joints, auriculate at the base. Eyes divided by the
lateral margin of head into two parts quite separate. Anterior
feet long, porrect ; four posterior feet short, broad. Elytra poste-
riorly rotundate, not covering the apex of abdomen.
Sub-genera: Orectochiltis Eschsch., Cybister Esghsch. (Oyretes
Brull^), Dineutes Mao L. {Cydous Esghsch., Dij.), Gyrinus^ Erir
hydrus Laporte. Comp. Aunouiir et Brull]£ Hist. not. des Ins, v.
pp. 234—242.
Sp. Oyrinut nataior L., Rosel Int. in. Tab. xzxi., Pakzkb DeuttckL Int.
Heft 3, Tab. 5, Ahbkns et Gkbmab Faun. Int. Bump. n. Thh. 5 ; the
male 1'", the female 5'" long, blue-black, shining ; the shields are stippled
in longitudinal rows. These whirling beeUet make their appearance early in
spring in fresh water, revolving in orcles on the sur&oe of the water with
much rapidity. On the least alarm they (tive suddenly below. Several other
European species are figured in Ahrens and Gkbmab Faun. Int. Eur.
Tab. n.
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INSECTA. 543
In OrtdocKUui (O^rirnu viUotus Fabb., Stdbx DeuUchl. Ins, x. Tab.
317) the outermost palpiform lobe of the under-jaws is wanting, and thus
they have only one feeler. 80 also in many large exotic Bpedea. Hence
thia character of the family is not constant^ but the agreement of habitut
does not permit these species to be placed elsewhere.
B. AbdoTnen eompoged 0/ seven segments^ the first three connate.
Dyttscus L. ^exclusive of the HydropMKy Dytwcus Geoffr.).
Antennae longer than head, filiform or setaceous, with eleven joints.
Eyes marginal. Head retracted within thorax as far as the
posterior margin of eyes. Posterior coxbb ample. Maxillse arcuate,
acuminate ; external maxillary palps with last joint oval, larger,
thicker or truncated.
Compare G. F. Ebichsoh Gtnera Dytkwrum Berolini, 1834 ; ejusd. DU
K&fer der Mark Brandenburg, I. p. 140, Ac
A« All the tarsi with five joints, anterior of males patellate.
Dyttscus Clairv., nob. (Sub-genera : Cyhister Curtis, AcUius
Leach, Hydaticus Leach, Dyttscus Leach, Ebichs.)
Sp. Dytitcui marginal^ L., Rosbkl Int. n. Tn», Aguat, CL i. Tab. i, Ltokbt
Omfr. podh. PL zi. figs. 1—8, Stubm DetOacM. Fauna, Int, vm. Tab. 185,
i86| D&riAN et Aub^ Jconogr. et BUt. not, dea Coleopt. d^ Europe, Tom. v.
PI. 5, figs. 3,4; 15'" long, feet and body red-brown, thorax and shield-
ooyers gleaming black, a yellow-brown border along all the sides of the
thorax and along the outer edge of shield-coyers ; the female {Dytieeue
aemietriaiut L.) has longitudinal furrows on the shield-covers more than half
their length. The females also of other species are distinguished by simi-
larly striated elytra, although in some of these females with smooth shield-
covers occur. The first three joints of the tarsi of the fore-feet form in the
male a round disc which is haired on the margin, and supplied with suckers
helow.—DjfHscue {ActUut) atdeaiue L,, Panzeb DeuUchl, Ins. Heft 31,
Tab. 9, 10, Stubx L 1. Tab. 189, &c.
The larvs of these water-beetles have a large oval head, armed with
strong curved jaws, three pairs of thin haired feet, and two appendages at
the hind part of the body subservient to respiration. They are very vora-
cious, and like the larvsB of libellse are incessantly on the hunt for other
water-insects. The nymphs lie concealed in the ground under the water,
Tlie perfect insects breathe on the surface of the water, by extending the
abdomen above it, and rainng the shield-covers a little, whilst the head
is kept under.
The intestinal canal does not undergo any remarkable change in the
metamorphosis. The four vessels that secrete urine are especially long,
the rectum has a lateral appendage, membranous and folded. Below the
rectum are the openings of vessels which secrete a milky fluid. Ramdohb
Ahhandl. Hb. d. VerdauungmBerJa, s. 79—83, Tab. n. figs. 1—5, L^K
DuFOUB, Ann. dee Sc. not. in. PI. 10, fig. 8.
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544 CLASS viii.
B. All the tarsi with five joints. Fint three joints of anterior tani in
males broad, transverse.
Colymbetea Clairv. Scutellum oonspicuoua. (Sub-genera : Coiym-
betes, ffyUtis Ebichs., Agahua Leach, Ebichs.)
Sp. Dyi. fuacut L., Panzeb Deuischl. Int, Heft 86, Tab. 5, GlaibviIiLE
Entom. helvet. n. PI. 2g, fig. B, &c.
Laccophilua Leach. Scutellum covered. Aifteniue tliiii, seta-
ceous. ^
Sp. Dyt, hyalinui De Geeb, LaccopkUiLS minutua Stubh, I>yt. obscwrus,
Pakzeb DeuUcM. Ins, Heft 26, Tab. 3, &o,
Noterua Clairv. Scutellum covered. Antennse incrassated in the
middle (especially in males).
Sp. Dyi, ercusicomtB Fabb., &c.
G. Anterior tarsi in both sexes subeqnal. (First and seoond pairs of
feet with tarsi of four joints, the penultimate joint obsolete or concealed.)
Hyphydrua (Ilug., Gyllenh. Sp. of Hydradvna Fabr.), Hydro-
poms Clairv.
Sp. Dytisc. palvstrii L., Dyt. Ut/wnOtu Pakzeb DeiUtdU. Ins. Heft 14, Tab. 4 ;
—Dyt, rufifrom Duftsohm., Stubm 1. 1. n. Tab. 407, fig. a, &c.
PeloUus ScHOENH., Erichs, (Spccies of Hydrcuilina Fabr.,
Hygrohia Lath.) Antennse filiform, short, with eleven joints.
Head sub-exsert, with eyes somewhat prominent. Labnim emar-
ginate. Thorax very short, much broader than head. Posterior
coxae narrow. Tarsi all with five joints, slender. Scutellum
distinct.
Sp. PelcbiuB ffermanni, ffydrachna Hermanni Fabb., Panzeb DeuUchl. Int.
Heft 10 1, Tab. i, Stubm 1. 1. vui, Tab. loo.
Haltplvs Latr. Antennse scarcely longer than head, filiform,
thin, with ten joints. Labrum entire, with margin ciliated. Pos-
terior coxfle narrow, covered by an orbicular plate. All the tarsi
with five joints. (External maxillary palps with last joint narrow,
subulate).
Sp. Saliplw r^ficoaia, Dyt, ruJicoUit, Db Gbeb Int, iv. Tab. 16^ fig. 9,
Panzbb DeuttOU, Int. Heft 14, ^. 10.
Cnemidotus Illig., Ebichs.
Note, — Differs from ffaliplua in the last joint of external maxillary palps
being greater, conioal. Sp. ffaliplut ccetut Gtllsnh., Panzkb DewttM.
Int, Heft 14, Tab. 7, D^jbak et AuBig, lixm^ygr, v. PL 3, fig. 2, ^
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INSECTA. 545
Family LXIII. Carahtcina {Adephaga terrestria, Carahici and
CtcindeletcB Latr.). Palps six; maxillae with double palp (ex-
ternal lacinia palpiform). Mandibles exsert. MaxillsB incurved at
the apex. Antennae thin, filiform. Body mostly oblong, with
thorax cordate, oval or quadrate. Abdomen composed of six seg-
ments, the three first connate.
A numerous family, in which D^tean in 1839 distinguished
nearly 2,800 species, and which with Linn.£US consisted of two
genera Ca/rabas and Cicindela numbering together, in the last
edition of the Si/stema Natv/rce, only 57 species.
Compare G. db Patkull Monographia ccurabwwn SuecicB, Upsalise, 1 790,
8yo; Glaibville Entomol, Belv^tique, voL 2, Zuric, 1806, Svo ; F. A.
BoNSLLi OhservtUUmB entamologiquet, 4to, 1809, {AeaaUm. des Sc.de Turin) ;
DiJEAN Specia gifUral des CoUoptirea d' Europe, Tom. i — iv. Paris, 1849—
183 1, Svo, &c
Phalanx I. Ligula porrect (often augmented by lateral ap-
pendages or paraglossse). Apex of maxilla (with few exceptions)
without hook distinct, articulate. Labial palps with three joints,
set upon labium by means of a peduncle elongate, immoveable.
Head not broader than thorax, mostly narrower. Many species
apterous. {Carahtcina in the stricter sense, Carahidce Leach,
Westw.)
Running-heedes, earthrbeetles. Most of the species live under-
ground, imder stones, under bark of trees. They run very swiftly.
The oesophagus is at its origin narrow, but it soon expands
hu'g^lyi ^ ^his crop succeeds the muscular stomach and a long
narrow stomach beset with villi, at the inferior extremity of which
the four vessels secreting urine are implanted. The small intestine
is short, the rectum oval and wide. Ramdohr, op. dt. pp. 82 — 84,
Tab. III. figa 7, 8, Tab. xxv. fig. 2, LioN Dupour Arm, des Sc. not,
n. PI. 20, 21.
The larvse have a longitudinal body, divided into twelve rings,
without counting the head. The first ring (prothorctx) is homy ;
the last has two conical, homy, or membranous appendages. The
head has strong jaws, six simple eyes on each side, and two antennsd
consisting of four joints. The six feet are of moderate length, with
two booklets at the extremity.
See fi^rnres of the larva of Carah, awroniteiu, Cwrab. hortentis, Carab.
depretam, and Cychnu rodraiut in O. Ukkb, Obierv. tniam. Tab. I. 11. and
VOL. I. 35
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546 CLASS YUI.
some others carefiilly collected and pnbHshed by WtsiwOOD, Jfodem
€fla$i\fication, i. pp. 64 — 71.
A, External nuunllary palpi mibulate at the apex, wUk pewiMmaU
jwid dbconieal, laet coniedL
BemMdium Latr. External maxillaiy and labial palps abraptly
subulate, with last joint very small, thin. Elytra entire. Anterior
tarsi of male with two first joints dilated. Anterior tibiae excised
at the inner margin. Thorax mostly cordate or subcordate.
Bembidium JUwipes Latb., Cieindda Jtavipee L., Panzxb Deutsckt. Im.
Heft ao, Tab. 2, Guv. R. Ant. Sd. iU,, In». PL 25, fig. 9 :—BenUnd. qvadn-
guttatum, Carab. quadriguttatus Fabb., Pakzsb Deut§ehl, Ine, Heft 40,
Tab. 5, Duicfo. Cone, gin, s. I, Ins, PI. i, fig. 6, kc.
Note. — Here belong sub-genera CiUenum Lbaoh, Blemtu Zaah,, 7VK%f-
put Mbobblb, Lopha, Ltjct, Bembidium, Notaphfu, and Taehys ejuad.
Compare D&riAK S^. gin. d. Col. v. pp. 31 — 194.
liochnophorua Dij.
Trechua Clairy. External maxillary palps with last joint
acuminate, conical, larger, of the same breadth at the base as the
penultimate joint. Anterior tarsi of male with two first joints
dilated, triangular. Anterior tibias emarginate internally.
Sp. Trechm rubens, Carabtte minutut Fabb., Panzbb J>eiUtchl. Int. Heft 73,
Tab. 6 (Oarab, tempeetivue), CLAiBvniiJi Entam. hdv. 11. Tab. n. fig. 6,
B, &c.
Anophihalmua SoHMnyr.
Sp. AnophthalfMU Sehmiddi Stubm D. Fauna, Ine. XV. Tikb. 303 ; 3'" long,
wingless, and without eyes.
B. External maailUxirypaipi not iubulate, ufith last joint a^
or truncated.
t Anterior tibia not emarginate on the internal nde.
Elaphrua Fabr. Labrum entire. External maxillary and labial
palps filiform, with last joint elongate, truncated. Anterior tibi»
with two terminal spines, the one inserted at the apex, the other
before the apex. Eyes globose, somewhat prominent. Body ob-
long. Anterior tarsi of male with first joints dilated.
Sp. Baphrui ripaHui, Cicindela riparia L., Paitzeb DeuUehl. Ine. Heft fto,
Tab. I, DuxtoL Cone. gin. s. I. Ine. PI. 4, fig. 6, &c.
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INBECTA. 647
Blethisa Bokelli (Blethisa and Fdophila D£r.).
NotiophUua Dumebil. Labrum rotundate. Anterior tarsi of
nude with £urst joints scarcely dilated
Sp. Elaphnu agucUicut Fabu,, Vtcindda aquoHea L., Pakzsb DeuUchl. Ins.
Heft ^o, Tfcb. 3.
Chncphron Lats., Scolytua Fabr« Labrum transverse, slightly
bisinnate. Anterior tarsi of male with two first joints dilated,
densely spongy beneath. Body subglobose, with thorax short,
transverse, covering scutellum. Feet elongate, thin.
8p. Omophron limbaiwm, JjATBl., DmiiaaL Oon». gin, s, I. Im. PI. i, fig. i,
Sturm D. Fauna, Int. vn. Tab. 184. (Genus allied to the Dytuci;
DssXAUBT has observed the larva ; see SiUBM Tab. dt.)
Nebria Latr.
ui^p{9U« BONISLU.
Leistus Froelich, Pogcmophorus Latr.
Sp. Leiihu tpinibarbis, Carah. apinibarhit F^BK., Paztzib DeuUchZ. In$,
Heft 89, Tab. a, Stubu D. Pawna, Ins. m. Tab. 70, &c.
Carabus L., (in part). Labrum bilobed, sinuate in the middle.
Labial and external maxillary palps elongate, exsert, with last
joint obconical or subsecuriform. Tibi» with two terminal spines.
Anterior tarsi of male with four first joints dilated, spongy beneath.
Ccdosoma Webeb, Fabb. Labrum deeply sinuate in the middle.
Mentum with lateral lobes exceeding the middle process. Thorax
short, transversely obcordate. Abdomen quadrate, with rounded
angles. Wings mostly distinct.
Sp. Caircbtma tycophanta L., Yoet Col, I. Tab. 37, fig. 3a, "DxntiBJL, Cons,
ffin. s. I. In$. PI. I, fig. 4, Stubm jD. Fauna, Int. m. Tab. 66, fig. a.
Ooe of our meet beautiful native insects, nearly one inch long ; head and
feet black, thorax violet-steel coloured, the shield-covers gold-green and
striated. Both in the state of larva and beetle it belongs to the useful
insects from devouring various kinds of caterpillars. See Bj^umttb Mim,
n. pp. 455—458, PI. 37, figs. 14—19, Ratzkb. Font-Int, i. pp. aa — 14,
46, 2y, Tab. i, fig. 1 1.
Carabus La.tr. Labrum broadly sinuate in the middla Mentum
with middle triangular process extending beyond the lateral lobes.
Thorax quadrate, with margins rounded, or obcordata Abdomen
oval. Wings either none or unfit for flight.
35—2
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548 CLASS vrii.
8p. Carahia emraitu L., Voir CoUopt. i. Tab. 36, fig. ^9, Pafier DevttehL
fns. Heft 81, Tab. 4, BruiiM D. Fauma, Int. m. Tab. 55 ; Carab. graMu-
laiui L. {Carab, eaneeOaJhu Fabb.), Pahzkb DentKld. Int, Heft 85,
Tab. I, Ac.
Procrugtea BoNELLi. Labmm trilobate. Labial and external
maxillaiy palps exsert, with last joint secnrifomu Mandibles
strong, denticulate. Mentom with middle process truncated, bifid.
Anterior tarsi of male with four first joints dilated, spongy beneath.
Eljtra connate, wings none.
Sp. ProenuUi eoriaeeus, (Jarab, eoriaeeus L., Panzxb DeuitekL /lu. Heft 81,
Tab. I, Smuf 2>. Fauna, Ina. m. Tab. 54.
Procerus Meqerle.
Tefflus Leach. (Is this its place ?)
Cychrus Fabr. Labrum deeply bifid. External maxillaiy
and labial palps porrect, with last joint broad, obtrigonal, ex-
cavated; internal maxillary palps dilated, cultriform. Mandibles
elongate, porrect, denticulate internally. Elytra connate.
a) Anterior tani of male with two or three first joints dilated.
Sub-genera : SphoBroderus Dij., Seapkinottis Latb., Dij.
Compare DitJMAJX Spec, gen. d. Col. n. pp. 14 — m.
b) AU the tarsi in both sexes simple.
Cj/chrua Lat&, D£r.
Sp. Cyeknu rotiraiut Fabb., Tenebrio rotbraltuB L., Dxtk^bil CcnM. gin. t. I.
I'M, PI. I, ^. i, Stubm 2>. Fawna, Int, m. Tab. 53.
Damaeter Kollail (Characters of Cychrus; but differs by labrum
transverse, only slightly sinuate, mandibles armed at the base with
only a single broad tooth). Body elongate. Elytra posteriorly
acuminate, dehiscent at the apex.
Sp. Damaster hUvptoldea Kollab, Awn, da Wiener Mutetan, 1, pp. 333, 334,
Tab. 31, fig. I. Habitat Japan, from whence also the Leyden Museum
has received specimens by the traveller V. Siebold.
Famborus Latr.
1 1 Anterior tibia emarginate on the internal aide,
• Ehftra nM truncated at the apex, rotundate.
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TNSECTA. 549
u) Abdomen continuous toUh thoraa:, not petiolate.
a) Tani of first pair in males toiih first two joints, or thrte dilaied,
subquadrate, forming a patella densely spongy beneath.
Badister ClAlRV.
Ltctmis Latr. External maxillary and labial palps with last
joint securiform. Mentum exsert in the middle, without tooth.
Sp. Licinus depressusy Lieinus cossyphoides, Sturm D. Fauna, Ins. in.
Tab. 74, fig. o.
Dtccdua BoNELU.
Bernima Latr.
Oodea Bonelli.
Epomis Bon., Latr.
DinodeB Bonelli, D£r.
Ghlceniua BoNELLi, GaUistus ejusd.
Loricera Latr.
Panagoeus Latr.
Sp. PanagcBus crux major, Carabus crux major It., SiUBM 2>. Fawna, Ins, ni.
Tab- 73i fig- «> A.
P) Tarsi of four anterior feet in males with three or four first joints
dilated, obeordate.
Harpahia Latr.
Sub-genera : Acupaipiu Latr, Stenolophua Zvsah,, Ophomu Ziegl.,
Ha^rpcUus Dij., Daptus Fibch., Acinoptis Zibol.
Sp. Barpalus r^fieomis, Carab, rt^ficomis Fabb., Pavzxb Deutsehl, Ins,
Heft 30, Tab. 3 ; a very common inflect in fields and gardens ; it hides
under stones.
7) Tarsi of first pair in males dilated, triangular or obeordate,
Zabrus Clairv.
Pogonvs Ziegl.
Tetragonoderus Dej.
Feronia Latr.
Sub-genera : Percua Bonelli, Molopa ejusd., Pterostichua ejusd.,
Abax ejusd.^ Cophoaua Ziegl., Gatadromua -^Mao L., Platyama
Bonelli, Ama/ra Bonelli, Dij., Pcecilua. Comp. Latr., R, AnL,
Sec ed, iv. p. 393, Dejean et Boisduyal IcoTuygraphie iil
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650 CLASS YIII.
Compare C. ZnofXHMAnr UA. die biaheriffe Qattmmg Amara, in Oian.
Fcnmtu, ZeiUekr, f. Zool, 1. 1, s. 1—40 ;— Scmdors in Kboteb's Tiddarift
I. pp. 39—65 And p. 99.
Genera: Myaa Ziegl., CepAahtes Bonelu, Golpodes Mac L.,
Bphodrvs Claibv., Bon., Ctenipw Latb., Ccdathua Bonelu,
Taphria BoNELLl.
b) Abdomen petioUxie. AnienncB ofien hroheft^ monUifarm.
Scaritea Fabb.
Add aeveral genera and sub-genera: Apotomiua HoFFifAir&,
DUomua Bonelu, Morio Latb., Dyschirius Bonelli, Clivina Latb,
CamptodarUus Dij., Oxystamua Latb., OxygjuUhus Djej., Scarite$
Fabb. in part, AearUhaaceUs Latb., Scapterui Di^., Panmackua
BONELLL
Sp. Secnilet artnariut Fabb., Tenebrio fimor L., Panzbb DeuUckl. Iiu, Heft
43, Tab. II, Claibyilli £fU. hdv, u. Tab. vin. fig. ▲, a.
Careniim BoNELLi.
8iag<ma Latb.
Encdadvs Bonelli.
Ozcena Oliv., ifortb Latb.
iVbte. — ^Add flub-genera BanUia and Meliiodera WnrwoOD, whom see
in GuiBiH MagoM. de Zool. 1835, PL 133. These ooleoptera fonn the
transition to the carabiei with tnmcated elytra, with which, aooording
to the opinion of some authors, they wonld be more ooirectly dassed.
* * Elytra tmneeUed at the apex or dUuae,
Truncatipenei Latb. ; a subdivision scarcely to be defined by oertun
limits, separate from the preceding by negatiye characiers; compare WxBT-
WOOD Modem Claeei/ication i. p. 75. Tarsi mostly not different in the
sexes. Claws of tarsi often denticiilate beneath.
Different species of the typical genus of this diTiaion, Brackinus
(Aptmua Bonelli), eject for their defence an acrid fluid from the
anus, which presently evaporates with an explosion and causes an
inflammation on the skin. Hence the name of Bombardter-beeile,
BoLANDKB gave the first account of this peculiarity in the TransactionB
of the Aoa<fSmy of Stockholm for 1 750. See, on the parts which secrete
the fluid, LlfiON DurouB, Ann, du Muefum, zvin. 1811, pp. 70 — 81, PL 5
(in Aptimu baUiata), and espedaUy Ami, dee So. natur. vm. pp. 6--I4.
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INSECTA. 651
a) External maacUHofry paipa tcith kut jakU ovcU, aeumMyUe^ or
robwndfUe,
a) WUk dano8 of tarti timple,
Odcxantka Payk., Fabr. Tarsi with fourth joint entire. Head
constricted posteriorly. Antennae moderate.
Sp. Odacamifha meUmura, Audabfu mdaMtrtu L., Pahzbb DmUsM. In$,
Heft lo. Tab. i, Stubm D. Fauna, Int. vn. Tab. 178.
Lqptatrachdut Latb. Sp. Odaoaniha dondlii Fabk.
CasnofUa Latb.
Mormolyce Haqekb. Antennsd elongate, almost of the length of
body, with first joint thick, clavate, second verj small, third and
fonrth very long, cylindrical. Head constricted posteriorly, elongate.
Elytra marginate, truncated posteriorly, with lobes of margin
romided, produced on each side beyond abdomen. Tarsi with
joints entire.
Sp. Mormolyce phyUodei Haobnb., GutiBJN leonogr., Ina. PL 6, £g. 7. An
insect of remarkable suse {ii inobes and more), from tbe island Java. See
J. J. HA.GXNBAOH MormoHyce, novum Ooleopterorum gen/va, NorimbergsBi,
1835, 8yo, c. Tab. color, (also transferred io Afm, du 8e, nai, Yl. pp.
500— 503, PL ai).
Cordistes Latr., Cahphwna Klug. Tarsi with four first joints
cordate. Antennae ahnost of the length of body.
Species from Soi;iih America, Odaeantha acuminakb SoHOEHH., OdacaiUha
hifaaeiata Fabb. &c.
/3) WUh clcnoi of tarti denticulaU heneaih.
DromivM Bonelli, D^. Mentum bilobed, without middle
tooth. Joints of tarsi entire.
Sp. Dnmitti guadrirnaculatut, Corah. guadrimaetdcUus L., ftc.
DemetrioB Bonelli, Dej. Mentum bilobed, with middle tooth
triangular, broad, small. Fourth joint of tarsi bilobed.
Sp. DetMtrioB dongaim D&r., Carabva ai/ricapiUui L., Stubx D, Fanna, Im.
▼u. Tab. 173, fig. a, A.
b) External maxiUary pcUpa with Ictet joint eylmdricaly or mUh
mcraeeated, tnmoatecL
a) With elawB of tarti denticulate beneath,
Lebia Latb. Mentum bilobed, no middle tooth. Last joint of
labial palps secoriform.
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552 CLASS viii.
a. With fourth joint of tarsi bilobed.
Sp. Lebia eyanocephalay Carah. cyanocephaltt* L., Panzbb Deutsckl. Ins.
Heft 75, Tab. 5.
h. With joints of tarsi entire. Cymindu Latr.
Add sub-genera : Flochioma DijEAN, GaUeida Dij.
Orthogonvua Dej.
Coptodera Dej.
Ctenodactyla Dej.
Agra Fabr. Last joint of labial palps very broad, securiform.
Mentum with middle tooth. Fourth joint of tarsi bilobed. Body
elongate. Head narrowed posteriorly. Thorax elongato-conical.
Species from South America. Compare KLua EnUm^ Monographienf pp. 9
— 42, Tab. I. n. and ni. figs, i, a.
fi) WUh clatDS of tarsi simple,
Brachtmia Weber, Fabr. Labrum sinuate in the middle,
broad, short. Mandibles exsert, with two basal teeth. Mentum
with middle tooth mostly none. Tarsi with joints entire. Thorax
obcordate, truncated. Elytra broader than thorax, truncated.
Sp. Erachinus crepitans Fabr., Carab. crepitans L., DuiciBiL Cons, pifn. s. I.
Ins. PL I, fig. 5, Sturm D. Fauna, Ins, vii. Tab. 176, &c.
Aptinus BcneiUi. Mentum furnished with middle tooth. Wings none.
£l3^tra truncated obliquely. Sp. Brack, mwtilatiu Fabr., Sturm D. Patma,
Ins. vn. Tab. 175 ; — Aptinus baUista Illiq., Germar Fowm, Ins. Europ.
Fasc. vm. Tab. 5.
Corsyra Steven.
Axinophorus Dej.
Eucheila Dej.
Cataacopus Kirby.
Anthia Weber, Fabr. Labrum porrect, rotundate. Mandibles
large, covered at the base by labrum. Mentum with middle tooth
indistinct. Tarsi with joints entire. Thorax obcordate, narrow
posteriorly. Eyes somewhat prominent. Wings none.
OraphiptertM Latb., D^j. Ligula in greatest part membranous.
Body short Abdomen broad, oval, almost orbicular.
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INSECTA. 653
Anthia Latr., D£j. Ligula homy. Body oblong. Abdomen
elongato-ovaL
Sp. Anthia decemfftOkOa, Carabut decemffuUahu L., Vosr Col. i. Tab. 38,
fig. 45» Cuv. R. Am., ed. ill, Int. PI. 18, fig. i ; habit. Cape of Good
Hope.
Helluo BONELLI.
c) External maasiXLary palps vnth kut joint obconical or sub-
secvriform, (Claws qf tarsi simple.)
Oalerita Fabr. Mandibles little exsert, short. Mentum with
middle tooth. Thorax elongate, obcordate.
Zuphivm Latb. {Zuphiwn P£j. and Folistickus Dej., Bonelll)
Joints of tarsi entire.
8p. Oalerita faaciolata Fabb., Stubm D. Fauna, Ins.Yii. Tab. 174 ;^Oalerita
oleni Fabb., Glaibv. Ent. hdv. u. PL xvii. fig. a, a, p. iii, CuY. R. Ani.,
id. UL, Ins. PI. 18, fig. 8.
Galerita Latr. Fourth joint of tarsi bifid- (Anterior tarsi of
male serrate internally, with joints produced into an auricula.)
Sp. Oalerita america/na Fabb., Cardb. americanua h., Ds Geeb /9ft. IV. Tab.
17, fig. 21, Latbeille Oen. Onut. el Int. Tab. y, fig. 2.
Drypta Fabr. Mandibles exsert, elongate, narrow, incurved at
the apex. Mentum with no middle tooth. Penultimate joint of
tarsi deeply bilobed. Thorax oblong.
Sp. I>rypta emarginata Fabb., Panzbb Deuttehl. Int. Heft 18, Tab. 15,
Glaibv. Bntom. hdv. n. Tab. xvni. p. 115, Stubm D. Pamia, Int. vu.
Tab. 179 ; in south of Europe ; lives under the bark of trees.
Phalanx II. Ligula inflected, concealed under mentum, homy,
with no paraglosssB. Maxillae mostly famished with a hook apical,
articulate. Labial palps with four joints, the basilar formed of
stipes free. Mandibles strong, dentate. Head large, with eyes
globose. Anterior tibiae with internal side entire. Tarsi long, slen-
der. {Gicmdelitm Latr., Cicindelidce Leach, Westw.)
This division consists in great part of the genus Gicrndda L.
Most of the species live in sandy places and run very rapidly.
Whence the name of Samd-runnjers^ Sand-heetles. The intestinal
canal nearly resembles that of the Garabi, but is shorter ; the
oesophagus is longer.
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554 CLASS YIII.
Ramdohb lib, ek. pp. 84 — 86, Tab. in. figs, i — 3, g, to, LioN DuiouB Arm,
d. Se. nai, m. PI. 10, fig. 2 ; both are figures of Oicindda camjteMrit,
The larvsB of Cidndda dig for themselyee pafisages undergTound,
and watch for their prey at the entrance of the hola They suspend
themselves in it by two curved booklets situated on the back at the
eighth ring of the body. The head and the first segment are broad
and strong, the last inversely cordate.
Compare J. A. Wbstwood, M6m. pow servir d rffiit. not. de la famUU det
(Hcmdditet. Atm. dea Se, nai. xxn. 183 1, pp. 399 — 317, PL 8 ; Eatkib.
Font-Im. I. Tab. t, fig. 13.
A. MenHim wUh no middle tooth,
CoUiuria Latb., CoUyris Fabb. Body narrow, elongate. Wings
distinct.
Sp. CM. UmgiooUii Fabb., Cuv. IL ArU,, id, OL, Im, PI. 17, fig. 5 ; Java.
IHcondyla Latb. Body narrow, elongate. Wings none.
Sp. J^ricondjfla altera, Oicindda aptera ToNBiB Luhd, Shrivter of Natwrh.
Sdthabei i. 1790, Tab. vi., Gufaiv Icanoffr,, Ina, PL 3, fig. a.
TTieKotea Latb. Body oblong. Internal maxillary palps very
short, nniarticnlate.
Sp. Therattei labiata, Oicindda tabida F., CuvncB R, Ani., id. iU,, Ina. PL
17, £g, 4; habit. Islands of the Padfic y^— 2%«pato» htuneraUa Mac L.,
habit. Java, ftc. — Mabkna of Oidndda.
R Mentum toith middle tooth,
a) MaxiUcB with no doM) apiccU cbrticulate,
Ctenostoma Klug.
Species from South America. Compare Kjajq, Sntom, Mcnogr, pp. 3 — 8,
Tab. ra. %g. 3.
Stenooera Bbulli^, PogomsUma Klug.
Sp. Stenocera degam Bbull^ Audoudt et 'BamiA, Hid. not, d. Ina, Tom. i.
PI. 3^ fig. 3 ; from Madagascar.
b) MctooiUcB supplied toith claw apical articulate.
Cicindela L. (in part). Anterior tarsi of male with three first
joints dilated.
* PaJpa egwU.
Sub-genera : (Xcindela Latb., Eupresopus Latb.^ Dij.
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INSECTA. 655
Sp. Oidndda hybrida h,, Pavzxb DeuUchl, Int, Heft 85, Tkb. 4, Voir
Cdeopi. I. Tab. 39, fig. t ; f" long, the onJ organs whitiBh-yeUow, elytra
and thorax copper-coloured ; on the elytra at the base and apex of each a
milk-white spot and a bent transyerse band of the same oolour in the
• middle. On a fine summer's day this insect is frequently seen on our sand-
hills. Oic. mariiima D^. differs from this species. lofmograjplUe et HUlL
neA, de» Col. SEwr, i. PL 3, fig. 3 }—€ficindela campettrii L., Pavzib L 1.
Tab. 3, Batzisbubo Pontt-Iiu. I. Tab. i, ^, 13, ftc.
* * Labial palps longer than manUary,
Sub-genera : OxycheUa Dij., Megaeephala Latb.
MarUicora Fabb. Tarsi not different in the sexes^ with joints
cjlindrical. (Mandibles ezsert, strong. Head large. Elytra
connate.)
Sp. MaiUioora fMudUoga Fabb.^ Oicifidda giffaiUea Thuitb., Duk^bil Com.
gH, f . I. Ina, Tt a, fig. 4 ; habitat Cape of Qood Hope.
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CLASS IX.
ARACHNIDS {ARACHNOIDEAy.
The Arachnids were counted by Linnjeus, as was stated above
(p. 248), to belong to tbe class of insects. They were first separated
from these as a distinct class by Lamarck (1801), who placed all
the wingless insects of Linn^us, which undergo no transformation
and breathe by air-slits, in his class of the Arachntda^, To this
then the myriapods, spring-tails, and parasites, would also belong,
which we have placed amongst the insects (pp. 288 — 303), and
Lamarck accordingly divides his Arackinula into those which
have antennas, and those which are without these parts. It is the
last alone, which we, following Latreille, refer to this class.
The class of arachnids, thus limited, is a veiy well defined and
natural division of the animal kingdom. The arachnids are arti-
culate animals with articulate feet or legs (see pp. 33, 248) and
without antennae. The head is with them joined to the thorax, so
^ For this daas we cite especially the following works :
Albih Spiders. — Elsazab Albir, Natural HitUyry of Spidert^ ilhatrated with 53
plaiet. London, 1736, 4to.
Clbbok Aran. — Gaboli Glkbok Aranei Sueeici ducriptionSmt etfgfirit ameit iUut-
trati. Stockholtniffi, 1757, 4to.
MuKLLEB ffydraehn.—ffydrachnoi qtuu in aqui» DanicBpalustribui dOexit, deaeripsU,
pingi et tabulia XL mteis incidi euravit O. F. Muelleb. LipsiiB^ 1781, 4to.
Hebm. Mhn. apt. — MSmoire apterologique par J. F. Hibmakn. Ayec 9 PI. enliun.
Strasbourg, 1804, foL
Walokbh. AramHd.--niMtoWt naiurd des Araniides par C. A. Walokbkabb.
Paris et Strasbourg, 1806. (5 parts in ismo, with col. fig. in the taste of Pahzbb
Deuitchl. Ins.)
SUNDKV. CompecL — C.J. Sundkvall Oompedua Arachnidum. Londini Gothomm,
1833, 8vo.
Hank u. Koch Araehn. — C. W. Hank Die Arachniden, gdrtu naeh der Natwr
abgebilda 1*. baehrieben. i, 1. 8vo. NUmbeig, 1831— 1835; C. J. Koch Die Arack-
mden (continuation of Hann'b work). Nttmberg, 1836— -1846. 3 — 11 vols.
Koch DeuttcMands Arachniden, in HBBBiOH-ScBiBFFBB's continuation of Pakueb's
Deutsehl. In$. (Aeaaij,
Koch Uebenichl.-^C. J. Koch Uebereieht der Arachmden-Systems. Nttmberg, 1837
— 1847 (4 parts with plates).
Walckxn. et GvRT.—ffistaire naJturdU dee Ineectea aptira. 4 vol. Paris, 1837 —
1848. (Suitee d BurrON ekez Robet ; the first three parts ; the third is by Gbbtaib.)
' SytUm^ dee AnimauK eoM verL p. 171.
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ARACHNIDS. 557
as to form a single piece, to which Latreille has given the name
of Cephahihorax : to this piece alone are the feet attached. Be-
hind the cephahihorax is connected with the abdomen, the second
principal piece of the body. A longitudinal arterial heart or dorsal
vessel is present, and in many a more or less developed vascular
system for the circulation of blood. Respiration is effected either
by means of air-tubes, as in insects, or of lungs in determinate
parts of the body. In all, however, there are lateral openings or
air-slits (see above, p. 260, 261), which conduct to the respiratory
organs. The sexes are distinct.
According to the theory of Savignt, no oral parts are present
in this class, which correspond to the upper and lower jaws of
insects. The parts, which in spiders and scorpions are usually
called upper jaws {mandibles) , are, according to Savigny, to be
compared with the second pair of auxiliary jaws, or feet changed
into jaws in the cray-fish and other ten-footed crustaceans. In
some arachnids these parts undergo such a change, that they
assume a flattened form and compose a sucker. The under-jaws
which succeed these, and which in the scorpions sustain large shear-
shaped feelers, are, according to Savignt, to be compared with the
third pair of auxiliary jaws, or feet changed into jaws of decapod
crustaceans. To these, in the arachnids, four pairs of feet succeed,
of which the first pair, according to the same writer, corresponds to
the second pair of untransformed feet of decapod crustaceans. The
untransformed first pair of feet of the decapod crustaceans, the so-
called chelcB or shears of cray-fish and crabs, would thus, like the
proper jaws (mandibles), be wanting in arachnids^. The abdomen
is never provided with feet.
' This view, however, is not altogether free from objection. Thus Latbsille oon-
sidere the first pair of jaws (the upper-jaws) of arachnids to be modified antennse.
[Ebiohson rejects this opinion of LATBSiiJiB; Bntomographien, erstes Heft. Berlin,
1840, s. 9 ; OwsN, on the contrary, on the ground of the origin of the nerves distributed
to these p^rts, defends it. Lectures on Comp. Anai. I. 1843, p. 2^3, ind edit. p. 448.]
Still more may it be doubted whether the first pair of feet of arachnids really corresponds
to the second pair of unaltered feet in decapod crustaceans ; this comparison may be
looked on as merely an arbitrary conception. Bather does the opinion deserve the
preference, that these parts correspond to the lateral parts of the under-lip. [This
opinion, I think, was first offered by W. Db Haan in an essay, of which the other
propositions appear to me to be less happy, entitied: Vergdijking tvMcken de tost-
haauw-en hcuxgingt-werhuigen der geUde diercn in Van Hall, Yrolik and Muldeb,
Bijdragen tot de natuurk. Wetcnsch. n. 1817, bl. 134, afterwards by Duqbs Ann. des
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558 CLASS IX.
The feet or legs of arachnids do not present exactly the same
tifpus as those of insects. The^ consist usually of seven joints, and
are not thinner towards the extremity, so that the tarsi differ less
from the other parts than in insects. K we suppose that the two
last joints belong to the tarsus, then the tibia consists of two joints
of which the first in some is the longest {e. g. Scorpio and Phrymuijy
in others the second. The long joint that precedes it is the Jkmur
which succeeds a short, inversely conical joint corresponding to the
trochanter in hexapod insects. The first joint, broader, commonly
inversely conical, by which the feet are attached to the cephalo-
thorax, corresponds to the hip (coxa) of insects. In Pkryfius^ the
tarsus of the fore legs consists of a great number of joints (thirty
and more), of extraordinary fineness, and in the remaining feet of
three joints. The last joint of the tarsus has usually two curved
booklets, and in some a membranous or vesicular cushion (j[ndviUua)
in addition at the under side. What is most distinctive in the com-
position of the feet in arachnids, is the division of the tibia into
two pieces. In PhryntAS, the tibia of the fore-feet, like the tarsus,
consists of numerous thin joints ,- those of the second and third pairs
of feet of the usual number, two ; those of the fourth pair of five, of
which the first exceeds the rest in length remarkably.
The ctphalothorax often presents a line or groove on each side
of the simple eyes placed in the middle, indicating an imperfect
separation, so that the head looks like a wedge or triangle, with the
point turned backwards, upon the thorax. The upper surfeoe of
Se, not 46 S^rie, i. Zoolog. p. 7, and by Eriohbon, EnUmogr, s. S.] In the scorpions
there are two homy triangular plates between the first pair of feet, which may be
regarded as metUum, a part wanting in the rest of the anachnids. An upper fip
{IdbrwaC^ is not present in these any more than in the rest. According to this view-
arachnids would have only three pairs of feet, like most insects. Yet Zoology must
neyertheless still continue to assert that arachnids have four pairs of feet, since, in
reality, four pairs of jointed appendages of the cephaUthoroiX do always act as feet
Even the jaws of hexapod insects are in a wider sense modified feet. Feet and jaws
are parts having the same fundamental form. This is proved by the histoiy^of develop-
ment. Both these parts are, according to the investigation of Bathkb in the Cray-fish,
of the same form, and the dissimilarity between feet and jaws, which is seen in perfect
cray-fishes, is merely the consequence of subsequent development {Untertwkungtn Hher
die BUdung u. Eniwkiodvmg dea Pluuhrtbws, Leipsig, 1819, fol. s. 67, 68). In young
and still imperfect individuals of Cfydopt quadricorwU, according to the figures of
JuBiKB, the upper jaws and the last pair of under jaws serve for swimming, and have
much resemblance to the feet whidi are developed only at a later period. Rathks loo.
cit. p. 73.
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ARACHNIDS. 559
the thorax consists of a lamina, in which sometimes, more or less
obviously, four sutures or grooves are seen, which proceed obliquely
from the feet to the center, and indicate the original composition of
the thorax of four pieces. On the under surface, between the cox»
of the feet, is a lamina, which may be considered as atemum^ formed
of the union of four pieces^ The abdomen of the Scorpions, of
Phrynus and Tel^honvs and (amongst the Arachnoidea tro/ckearia)
of Obisium is divided into rings; in others, as the spiders, it is
without rings, and has a much softer integument than the cephalo-
thorax. On the whole, the external covering of the Arachnids is
soft and very extensible ; the skin is homy and hard in Scorpio^
Phrynus, in some EpeircB {Epeira cancriformis)^ &c. Two layers
may usually be distinguished in the skin ; the external is firmer,
sometimes cellular, often provided with sinuous folds ; the internal
consists of a finely granular substance or of very delicate fibres, be-
neath which is a layer of pigment. In MygaJe, these pigment cells
are very apparent. The external membrane exhibits many concen-
tric rings with spots between them, which, under the microscope,
present a deceptive resemblance to the corpuscles and lamellsB of
bone.
The intestinal canal of the arachnids proceeds without tortu-
osities to the posterior extremity of the body. Not in all of them,
however, is the anus situated at the posterior extremity, but in the
Aeari more forward on the ventral surfiu>e^. In Phrytms and
Scorpio f the intestine is a narrow canal, nearly of the same width
throughout. In all the other Arachnids, the intestinal canal has
many protrusions or blind appendages of very different form ; and
in many Aca/rina these appendages are ramified, reminding us of
the form of the intestine in Planarice and DistomcUa. In the I^fo-
nogonida blind appendages proceed from the stomach, even into the
shear-shaped feelers and the feet. In the genus Phalangium the
intestinal canal forms a wide sac, which has in front, on each side,
five blind appendages that become wider towards the extremity, of
which the last pair is longer than the rest, and descends to the
termination of the intestinal canal. Between these appendages are
still twelve others smaller and vesicular, and at the hindmost part
^ See the figure of Jxodeg, Acartu amerieaiwM (or crmuiiui Kollab) in Tbbtiba-
iruSy ZeUtckr,far Phynol. iv. «. 1831.
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660 CLASS IX.
of the intestinal canal there are in addition four small blind appen-
dages on each side^ In the spiders {Aranea L.) the oesophagus is
narrow, homy, bent first downwards, then upwards, and temunates
in a part which proceeds backwards, and can be dilated by muscles
attached to it'. This part becomes at its extremity more membra-
nous, and passes into the intestinal canal, which sends two lateral
arms forwards that bend themselves round in the ccphalothorax
and approach each other or unite so as to form an annular stomach.
From this ring blind eversions proceed towards the feet, bend
downwards, and communicate with each other. From the same
point where the two lateral arms arise a middle canal exists, which
runs towards the abdomen and is gradually dilated in some degree.
The widened part receives on each side two large canals which
branch through the adipose body ; the intestinal canal then narrower
curves upwards and afterwards proceeds downwards, where, pre-
vious to its termination, an oval coecum is situated laterally.
Organs of different form for the secretion of saliva are observed
in most genera of this class, and probably are wanting in none of
the arachnids, although their opening into the intestinal canal has
not been detected in all. In the Scorpions, where Meckel* de-
nied altogether the existence of salivary organs, two longitudinal
sacs placed in the fore part of the abdonjinal cavity, their tortuous
canals running forwards in the ccphalothorax, probably serve for
the secretion of saliva*. From the secretion of saliva the poison
should be distinguished, with which different species kill their prey
previous to devouring it. In the spiders there is situated at the
base of each of the mandibles, or sometimes in the mandibles them-
selves, a vesicle, surrounded by a layer of flat spirally turned fila-
ments, from the anterior part of which a narrow duct arises, that
runs towards the point of the upper jaw. This poison has an effect
* Ramdohb Verdauungt toerJa. d. Ins, 8. 104, ao5, Tesviranus Verm. Schr, i.
«• ^9— 3'> Tab. III. figa. 16, 17, Tulk in Ann. of not. Bid, xn. 1843, PP- 446— 248.
PI. IV. fig. 17.
* Tbevibanus, in his work Uther den innem Ban der Araehniden, 8. 29, 30. Tab.
II. fig. 44, ha8 described the intestinal canal imperfectly. Bbandt Medk, Zool. ii.
has partially improved on him. We follow here especially Wasmann in Abkand-
lungen herautgegeben von den naturvisgenich. Verein, Hambuiig, 1846. I. 4to. s. 141 —
144.
' Syst, der vergl, Anaiom. iv. b. 145.
* J. MuKLLKB in Meokbl'8 Archivf. Anat, u. Physiol, i8a8. s. 52.
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ARACHNIDS. 561
often rapidly mortal upon those insects on which spiders feed, and
even large flies die quickly when they have been wounded on a
single foot alone by the bite of Clubtona cUrox^.
We have seen that in the spiders, firom the dilatation of the
intestinal canal situated in the abdomen, which might be taken for
the hindmost stomach, large canals proceed to the adipose body.
Similarly in the scorpions from the intestinal canal, at nearly equal
and very large distances from each other, there arise on each side
five transverse branches which subdivide into finer branches and
run through the granular adipose mass. Already, when treating of
the Class of Insects, we directed attention to the suggestion that
there perhaps the adipose body assumed the place of the liver
(p. 257). In most arachnids {Scorpio, Aranea L.) this part cer-
tainly, with still stronger claim, deserves to be thus considered*.
It consists of lobes, formed of small blind vesicles united in clusters,
and filled with cells. The ducts in question, which run towards
the intestinal canal or proceed from it as branching eversions, are
to be considered then as gall-ducts. In other arachnids {Phalan-
fftta, Acarinay Arctisca and Pycnog(mid<i) the glandular walls of the
blind intestinal appendages probably serve for the secretion of bile'.
In most arachnids there exist also thin tubes with blind ex-
tremities which correspond to the Malpighian vessels of insects (see
above, pp. 255, 256), and so are to be considered as subservient to
the urinary secretion. They differ, however, from the vessels of
insects alluded to, inasmuch as they usually divide into many
branches.
The organs for respiration and the circulation of blood are not
in these animals formed after one and the same type. When
respiration is performed by means of air-tubes, there is a dorsal
vessel, as in insects, a longitudinal heart, without branches ; vessels,
on the other hand, are found in those genera in which the respira-
tory organs are sacciform lungs, and are not spread throughout the
body as air-tubes. In PAalangimn, the heart is a dorsal vessel
without branches, which becomes narrower at both extremities,
and is divided by constrictions into three chambers, or dilata-
1 TBEYiBAirnB, Ueber d, inn. Bau d, Arachn, s. 31, 31, Tab. i, figs. 21, 22,
* J. F. MiOKEL JBeytrtige z. vergt. Anat, i, 2, 1809. a. 108. See also especially
Wasxahn 1. c. pp. 145 — 148, Tab. 13, figs. 10— aa.
> Y. SlEBOLD, Lehrb, der vergl. Anai. I. s. 519.
TOL. I. 36
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562 CLASS IX.
tions^. In many of the inferior arachnids neither heart nor yessels
have been met with.
In the spiders and scorpions, on the contrary, there are vessels
present for the circulation of the hlood* The heart has, indeed, still
the form of a longitudinal vessel, hnt other vessels arise from it,
whilst from the respiratory organs the returning arterialised blood
penetrates the heart through lateral, transverse fissures on the
upper or dorsal surface, which are provided with valves. The
course of these returning vessels, corresponding to the pulmonary
veins of the human body, is not yet perfectly understood. Pro-
bably the whole heart is surrounded by a sinus, into which the
arterial blood is poured, before it penetrates the above-mentioned
fissures. Vessels also from the heart proceed to the respiratory
organs, but these probably serve for their nutrition, and not for
respiration. The heart of arachnids is then, beyond doubt, an
arterial heart, like that of cruatacea and of moUuscaK
We have already said that the respiratory organs are in some
arachnids air-tubes, in others pulmonary sacs. In both cases the
air has access to the respiratory organs by air^slits {st^mata), aa in
insects ; but these stigmata are always in less number than in
most insects. In some arachnids no respiratory organs at all have
been discovered {I^cnogonumy the so-named tardigraday many
Acatx), In these also no stigmata are present. Sometimes, indeed,
parts have been taken for stigmata, which have an entirely different
signification ; I allude to two rows of spots on the back, which are
for the attachment of muscles, which, coimecting the dorsal and
ventral surfaces, contract the abdomen, and which we also meet
with again in lAmulus, amongst the Crustacea^. In the scorpions
^ TBKVimAKUB Fmik Sckr, i. s. 31, Tab. m. fig. 18, Tolk L 0. p. i49> ^^ ^-
fig. 17 a", ir, p.
* The most complete description of the heart and yessels of the sooipions has been
given by Niwpobt in PAfl. Transact, for 1843, Part i. pp. 486— 198, with beautiful
figures. From the dorsal vessel, that as an aorta springs from the heart at the fore part,
there arises, before the origin of the arteries for the last pair of foet, an arteiy on each
side, which passes beneath the oosophagus ; the two form a single vessel lying on the
nervous cord. It is this artery which was described by Tbxvibanus as a third nervous
cord, and by Mukllbb as a ligament. Under the nervous oord in the abdomen a
venous trunk is situated.
* This mistake ^jpia made even by the celebrated Tbmvirawjb in Chdifer {Verm.
Sehr. I. 8. 18, 19, Tab. 11. figs. 6, 7, a), who could not on that account find any air-
tubes there.
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ARACHKIDS. 663
the air-filits are obliquely transyerse fiBSures on the ventral plates
of the abdomen. The uppermoBt or anterior lip of these slits
coTors in some degree the lower, from which last a membranona
margin arises, which bears the respiratory organ situated in a
small eavily. This lung (or gill) consists here, as in spiders, in
Theiyfho/»x» and Phrynus^ of a number of double, very thin plates
lying upon each other. If now, as is asserted, the air on respira-
tion really penetrates this chamber so as to fill the spaces between
the duplicatures, then the name of lung would be justifiable ^ The
ordinary position of the stigmata is on the inferior surface of the
uppermost part of the abdomen. There also AuDOUiN discovered
four stigmata in the genus Chelifer that breathes by air-tubes'.
The Phalomgia that also breathe by air-tubes have only a single
pair of stigmata. In Ixodes Lyonet and Audouin found the two
stigmata famished with a plate, and upon it, besides a larger open-
ing, many other smaller ones with a stellate margin'. In spiders
also the stigma is not always a simple fissure, as in the scorpions,
but is sometimes closed by a plate perforated like a sieve ^. The
trachece of arachnids differ often firom those of insects by the
absence of the spiral thread. Usually also they are parcelled in
bundles, and not divided into branches. However, in Phalangtumy
a system of air-tubes is met with divided into branches and spread
throughout the body, and provided also with a spiral thread.
There are two wide principal stems which, nmning forwards in the
cephalothorax obliquely towards each other, divide into branches,
whilst a transverse branch on the inside, behind the thoracic gan-
glion, forms an arch by uniting with that of the opposite side. In
tiie abdomen, behind the stigmata, the lateral principal branches do
not continue their course, but three smaller branches alone firom the
principal stem penetrate backwards on each side*.
^ Not on account of respiring in air ; for all animals that livs in air have not lungs,
the land-orabs, for example, have gills. The respiratory organs of the ffoUAwria,
on the other hand, although these animals inhale water, are formed alter the tjfpug
of hmgs.
* Ann. dm 8e. nad, xxvn. 1834. p. 6a.
* Ltonet Reeh, PL 6, ^, 5, AuDOUiv, Ann: dt9 Se. nai, zxv. p. 419, and Todd'b
Oj^doptBdia i. p. 405.
* LTOim, 1. 1. PI. 10, £g, 10.
* TBBvniAinTa Verm, Sehr, i. s. 34, 33, Tab. iv. fig. 19, and especially Tulk 1. c.
pp. 5«7— 3«9. W. ▼• fig- 33.
36—2
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564 CLASS IX.
The male and female organs of propagation are, with the excep-
tion of the Tardigraday in all arachnids distinct, and not united in the
same individual. The ovaries are usually two sacs that are some^
times connected in the middle ; the oviduct is constantly an imme-
diate continuation of the ovary. In Phalangium the ovaiy consists
of a tortuous, circular tube connected with a wide sac, in which the
eggs await for a time their farther development. From this sac a
second oviduct arises, which running tortuously first backwards
then forwards, terminates in the ovipositor. This ovipositor con*
sists of homy transverse filaments, boimd mutually together by a
tough elastic membrane. It can be protruded by muscles through
the external sexual opening situated between the hind feet\ In
the scorpions, the internal female organs of propagation consist of
three tubes running longitudinally ; the middle tube is united with
the lateral by three transverse branches; the two lateral tubes,
moreover, unite with each other in an arch below. They extend
farther forwards than the middle tube and thus form two oviducts
as their immediate continuation, which open at the external sexaal
aperture under a valve between the base of the third and fourth
pair of feet. Here each of these oviducts, before it terminates,
forms a roundish expansion, which may be considered as a rec^pta-
The male organs of propagation consist of two blind saccules or
two tortuous and blind canals ; in others of two bundles or of a single
group of numerous blind saccules or tubules, as in Phalanffium. In
the scorpions two loops or tortuous strings of a narrow canal, with
two transverse canals, are foimd. The vcu deferens also receives the
openings of two blind sacs before reaching the external sexual aper-
ture, which is in the same situation as that of the female. Here is
1 TBXvnLAXVB ibid. s. 47— 49» T»b. v. fig. i8, Tab. vi. fig^ 3a, 35, Tdlk L 0.
pp. 318—344. PL V. figg. «6, 17, «8.
* Y. SuBOLD {LeM. der vergL Anat. i. a. 548) found in it SpermaUuoa in liriog
females. Cknnp. on the described Ovaria in Scorpio, Muillib in Miguel's ArMt /.
AneU. «. Phyriol. i8a8. i. 54, 55, Tab. n. ^, 16. In the dereloped state of the otary,
pear-shaped eversions proceed from these tubes of the impregnated female. Muu-i^
saw these in the form of elongate blind sacs, in the middle excavated like a bean. lo
this wider part the embryo of the scorpion is contained. I also haye met with the
ovary of Scorpio reiieulatui fit>m Java in the developed state. Scorpions are vivipvo^
of which mention was made already by the ancients ; Abistotslbs Hid. Ami^ ^'
cap. 15, Mluxvb de AninuU. not. Lib. ▼. cap. 40.
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ARACHNIDS. 565
found a small papilla, which may be considered as the rudiment of
Apents^ Also in some Acarina a short penis is present. In Pha'
lanfftuniy on the other hand, there is a long homy penis which can
be extended from the body at the same part as the ovipositor in
the female, and through which the ductus efaculatoriua extends;
at the upper extremity of the gland is a small booklet. In the
spiders the two efferent vessels, arising from the blind tubes that
secrete the seed, run towards the base of the abdomen, where they
open, without any copulative organ, between the gills. The organ
of copulation lies veiy distant from this opening. In the spiders,
namely, the last joint of the feeler is excavated like a spoon, with
various appendages in addition, different in different species, and a
homy, curved filament connected, which lies concealed in the exca-
vation, and can be extraded from it. With this club of the palpiy
the male, at the time of pairing, touches the sexual parts of the
female, having previously moistened it with a drop of seminal fluid
from the opening of the vasa deferentia. This is thus the copula-
tion itself, and by no means, as Treviranus supposed, a prelimi-
nary sport to excite the sexual passion : this is the copulation itself,
which Lister, De Geer, Ltonet, and others, described so frilly
and so interestingly*. The danger of being cruelly put to death by
the female spider causes the male to make his approaches to her
with anxious caution, and after the congress to betake himself
rapidly to flight.
In the development of the arachnids from the egg, afrer the
germinal vesicle has disappeared, there appear to occur, as in many
other animals, clefts and grooves in the yolk-mass. The ger-
minal membrane or disc grows slowly round the yolk, closing
finally on the dorsal surface. Before this closure is completed,
the commencement of the embryo appears on the ventral surface of
the yolk. Here are seen, at least in the embryo of the scorpions
according to the observations of Rathke, different thickenings
lying in pairs near each other in a row, which are the beginnings
of the segments of the body. From the mucous layer of the ger-
minal membrane arises the intestinal canal, which at first is not
1 See, for insUnce, Ltosst in bis tranBlation of LssssB Thiol, der Ins, I. p. 184,
and in his Jtechereh4!t, pp. 73 — 75, and Walokknabb Hist, not, de$ Araniides, ▼. 8,
respecting Theridion heniffnum. See also Owen Ledwret en eomp. Anal, of inveH,
Anim. p. 264, snd ed. p. 463.
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566 CLASS IX.
separated firom the jolk, afterwards lies between two lateral portionfl
of the jolk, which, by transverse iBdentations, change into ooecal
sacs placed in pairs. These sacs, in connexion with the intestinal
canal, are the rudiments of the biliaiy yessels and of the liyer. The
yolk changes into the liver, or the so-called adipose body. The
limbs arise as conical appendages placed under the ventral sur&oe,
with the extremities turned downwards towards each other. On
the dorsal surfsuse of the yolk is seen a streak running longitadi-
nallj as the rudiment of the heart, that is at first without veasek.
The nervous system, in its central parts, is formed at a still earlier
period, and the cerebral ganglion is in the beginning proportionally
much larger than it is afterwards ^
We have abeady said that the scorpions are viviparous. With
the egg-laying spiders, the egg, under the changes of development,
slowly loses its previous form, and almost assumes that of a spider,
indicating all the external parts of the inclosed animaL At length
the shell bursts on the thorax, and the spider, first with the head,
and afterwards with the thorax, comes to view ; then follows the
abdomen, to which however the egg-membrane, like a scale, con-
tinues attached for a time ; then come the feelers and feet*. The
young spider, through whose integument the granules of the yolk
may be clearly distinguished, is not yet in a state to weave a web
and catch its prey ; for the spinning organs are still concealed be-
neath the common integument. After the lapse of a week, or, in
some species, a longer time, during which the spider takes no food,
it casts its skin for the first time, and is, as it were, bom for the
second time. The young spiders now quit, on some mild day in
May or June, the web in which the mother had hidden her eggs;
they allow themselves to £all on the ground by a thread, and begin
at once to weave their nets, or in some other way, according to the
instinct of their kind, to watch for small insects corresponding to
their age and powers.
^ On the deyelopment of spiders see M. Hbbold De Oeneratitme Afxmearwm in Ovo,
Marburgi, 1834, fol. oum tab. seneis: Di WimoH OburvationM de Ara/Mnrum ex ovo
evoUUione, Halls, 1845 ; of soorpions, H. Ratbkb Zur MorphologU, Riga a. Leipog,
1837. 4to. pp. 17—34. PI. I. figs. 1—1 1 ; compare also the shorter description of these
last observations and of those of Hkbold in Bcbdaoh Die Phyeiologie ale Efft^nnnge-
vjieeeneehaftf ate Ausg. 11. 1837. s. 943 — 948.
* D« GXXB M4m, pour eervir d VHiet. dm Ine, vn. pp. 195, 196, PI. 18, figi.
11—14.
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ABACHNIDS. 567
Most arachnids feed on other animals, which they either swal*
low alive, or wUbse blood and fluids they suck. Usually, after
their escape firom the egg, they undergo no metamorphosis. They
cast, however, their skin more than once, and are commonly, after
the fourth or fifth moult, in a state for pairing. In most Aoarina,
the young animals are at first supplied with only three pairs of feet,
which gave occasion to some writers to adopt some six-footed
genera, which were afterwards rejected as unfounded. The Pyana-
gcnida and the genus Eydrachna present the most interesting
changes of form, of which the most remarkable particulars will be
noticed in the systematic arrangement of these animals.
The power of reproduction in arachnids is commonly, as in the
preceding class, considered to be small ^. In many, however, lost
feet can grow again. Thus Geoffsoy once saw a Phalangium, in
which one foot was less than the remaining seven', and which pro-
bably might have grown af a later period. At all events it is
established that, in spiders, lost feet are regenerated^ In animals,
whose growth is limited, i,e* which do not grow after they are
capable of propagating, I think the reproductive power, in this ftdl-
grown state, is small. In such a case are insects after their last
metamorphosis (see above, p. 276), but by no means spiders and
crustaceans.
Very various is the form of the nervous system in the arachnids.
In the greatest number there is a large ganglion in the thorax
{cqihaloihorax)^ formed, as it seems, by the coalition of different
other ganglia, firom which the nerves for the under-jaws KsApdlpi^
and for the four pairs of feet, radiate. At the posterior margin of
this ganglion arise, under acute angles, close to each other (as in
the Cauda equina of mammals), the nervous trunks for the abdomen.
The two middlemost are sometimes thicker, lie closer together, and
unite towards the extremity, before dividing, to form a ganglion
^ Mboksl'b 8ytt. der vergl, AnaJt, i. s. I3i.
^ Hitt. abrigU des Int. i. p. 629.
' Seo an observation of ihe celebrated Banks, intereeting also in other respects
recorded by Leaoh 2Van«. of the Linn. Soc. xi. 18 15, pp. 393, 394; see also Heine-
Kent's experiments and observations, Zool. JourTud, lY. 1829, pp. 184, 994, and those
of Lefbletisb and Audouin in Todd's Oydopcedia, I. pp. 214, 115. Spiders must,
however, lose the entire foot as fiu* as the coxa; if it be broken off lower the spider
dies, unless it succeed in breaking oi¥ the stump that is left. The new foot (at first
very short) makes its appearance at the next moult.
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568 CLAS8 1X«
{Tdtphonus), or lying side by side fonn an abdominal cord with-
out ganglia, aa in the spiders. A chain of ganglia, as in most
insects, is found in Tardiffraday in I\fcnaffonida^j and in Scorpio
alone. In the I\fcnog(mida, the four ganglia, forming the row, lie
close together, without connecting cords, just as SwAMiCEfiDAM has
figored the nenrous system in Pediculus (see above, p. 301). In
Scorpio^ on the contrary, they lie, seven in number^ at considerable
distances apart, and are connected by two strings'. Above, and
commonly close upon the thoracic ganglion, is placed a ganglion
which may be considered as cerebral ganglion; in veiy simply
organised arachnids it is seen as a simple commissure on the oeso-
phagus; in others it is oblong and formed of two small parts,
mostly conical or pearnshaped, lying side by side. From it arise
the nerves of the mandibles and of the eyes. Between this cerebral
ganglion and the thoracic ganglion, there is* always an opening,
mostly very narrow, for the passage of the oesophagus, which is
surrounded on each side by the nervous connexion of tbe two gan-
glia. That the first ganglion is smaller than the second, and not
broad, as in most insects, depends, without doubt, principally upon
the absence of the compound eyes, whose nerves, in insects, have
such a large development. Interesting also is the exceptional form
of the nervous system in Phalangium, where the nerves proceeding
from the thoracic ganglion form eight ganglia in their course, four on
each side, not behind one another in a row, but at different heights,
dispersed on each side of the body, and distant from each other^
Traces of a distinct nervous system for the intestines, of that
system which we indicated above, in insects, as answering to the
1 QOATBiVAasB Ann, det 8e, not. ai^e S^rie, iv. 1845. Zoologie, p. 77, PL I.
And n.
' Tbeyiiuvus Utif, d, inn. Ban der Arachn. s, 14 — 16« Tab. I. fig. 13, and espe-
cially Zeit$ehr. fOr Phytiol. it. 1831. 8. 89 — 97, Taf. Yi. and the elaborate fig. of
Nkwfobt Phil, Tram, 1843, Part I. PL xii. That the nervous system in PhrynvM
and Tdephonm is formed not after the type of the scorpions, but of the spiders, was
announced by me in the T^jdschr, von Nat, Ouch, en Phytiol. ix. 1849, bL 75, and x.
1843, bL 369, 370. In Telephonut, at least, it would be difficult to haye anticipated
this, and it is also in contradiction to the rules, already contradicted indeed by other
instances, which Stbaus Dubckhbim formerly laid down for the form of the nervous
system of articulate animals. Contid, gin, », VAnai, comp. det Ani, art, pp. 364, 365,
371.
• TBBViBAin7B Verm, Schr, i. s. 38, 39, Tab. iv. fig. 14, Tulk L 1. p. 3a5» ^1- ^•
fig. 31.
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ARACHNIDS. 569
nervua sympaiheticua (p. 278), are onlj imperfectly recognised.
There has been found, in spiders and scorpions, a nerve that runs '
towards the stomach, and arises by two thin roots from the posterior
margin of the cerebral ganglion^.
The organs of sense in arachnids are limited, as far as they are
known, to those of sight and feeling. A distinct organ of hearing
has not yet been discovered. The seat of taste, probably also of
smell, is to be sought for perhaps in the commencement of the intes-
tinal canal. For feeling the palpi^ in many at least, officiate ; in
others, where, as in Scorpio and Chelijery they are covered with a
hard skin and are shaped like shears, they serve for seizing prey.
The long feet also of spiders and of Phalangiay and especially the
thread-like feet of the first pair in Fhryntis, are peculiarly adapted
for organs of feeling. The eyes are constantly simple. Behind the
convex cornea is placed a veiy round lens, which is received by a
vitreous humour concave on the anterior surface. Behind this
humour lies the retina, surrounded by a coloured pigment of the
choroideay which at the fore part, between the lens and the vitreous
humour, forms a ring like an irts. For the presence or absence and
the number of simple eyes, we refer to the systematic arrangement
of the class. Often, as in Scorpio, Phrynua and TelyphonuSy two
simple eyes are placed close together in the middle, and on each
side two groups of smaller simple eyes towards the margin of the
aphahthorax. If this disposition be compared with that of insects,
then the middle eyes may be likened to their simple eyes, and the
two lateral groups to their two compound eyes.
The muscles of the arachnids are principally inserted in the
external integument. The cavity of the cephalothorax is especially
occupied by a mass of muscle, because the limbs, which are moved
by the muscles, are placed in a circle round about that part. In
spiders there is found here, for the attachment of the muscles, in
addition, an internal oblong plate, tendinous, hollow above, and
with the anterior margin excised. This part was already observed
by Lyonet?. It was also found by me in Phrynus, and in the
^ Brandt Med. ZoU. u. a. 90, Nxwpobt 1. L p. 161.
' JRechereket mr VAnai. &c. Ouvrage potthume, p. too, PL lO, fig. 26, Tbkyibakub
Ueb. d. inn. Bau der Araehn. b. 30, Tab. ii. fig. 13. Tbivibawus calls this part
inoorrectly a cartilage (JTnorpet).
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570 CLASS IX.
class of the Crustacea occurs largely developed in Limuhu. The
' name of inner sternum has been given to it ; it occupies in some
degree the place of the entothcrax in insects, .is not however a
homy part penetrating inwards from the integoment^ but a pro-
duction from the numerous interwoven tendons of the different
muscles which mutually meet there ^
Arachnids are met with in eveiy quarter of the globe. I am
acquainted with no frunilies or principal groups (larger genera in the
sense of Linnjeus) which, as is the case in other classes, belong
exclusively to the eastern or the western hemispheie. The larger
arachnids inhabit warm regions, especially the genera Scorpio^
Mygahj Telyphonus and Phrynus. By fsx the most of the ^-nimftlg
of this class live on land, others in fresh water, in the sea only very
few, just the reverse of what we remark in the crustaceans.
^ A similar part is the Hnea oJIha, for the abdominal muscles of mammals.
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SYSTEMATIC
ARRANGEMENT OF ARACHNIDS,
CLASS EL
AKACHNOIDEA.
Abticulate animals with articulate feet. Head and thorax
conjoined to form a single part. Feet eight, placed at the sides of
cephalothorax; abdominal feet none. Heart placed in the back,
resembling an elongate vessel, in manj giving off arteries. Bespi-
ration in some tracheal, in others pnlmonal; in some no distinct
organs of respiration. Sexes mostly distinct.
Section I. Tmetothoraca s. Apneuata. Cephalothorax divided
into fonr segments. Stigmata none. Organs of respiration none.
(Seat of respiration either in the external integument of body or in
the digestive canal.)
Order I. Polyganopoda.
Feet elongate, of the length of body or longer than body. First
segment of body tubular, exsert, perforated at the apex by the
mouth. Ocelli four in a tubercle behind the tube at the middle part
of the second segment. Abdomen small, conical.
Family I. R/cnogonida (characters of the order).
SecHtpiders. The genus Pycnogowam of BRUENiracH {PofygonaptAs
Pallas), with some other allied genera of later writers^ forms a
small group of marine animals, on the true place of which in the
natural arrangement opinions differ; for Milne Edwabds, and espe-
cially QuATBBFAOSS and Kboeter, refer them to the crustaceans.
That some of these animals live parasitically on whales and other
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572 CLASS IX.
marine creatures appears to be a IIlistake^ Thej are dull animaLi
living on the beach under stones, or on marine plants. They
undergo, according to the observationB of EIboeyeb, a remarkable
metamorphosis, and have at first onlj two pairs of short feet^ and a
short, thick, undivided body. Afterwards a third pair of feet, still
short, appears, and only in a succeeding state is the fourth pair added
to them.
That division of the body which lies in front of the first pair of
feet and bears the eyes above, is in some more, in others lees dis-
tinctly distinguished from the first ring of the thorax. To this
division are attached on the under surface two filiform parts, consist-
ing of 7 — 11 joints, which in some species occur only in the female
and serve for the attachment of the egg& In some species in front of
these are two pairs of other parts at the base of the conical sucker,
of which the first pair, shorter and broader, mostly forms a shear-
shaped nipper, the second is filiform, and consists of five, sometimes
of more joints, and is commonly denoted as a feeler (^pcUptts). It
is worthy of remark that the first-named, the nippers or mcmdibtUcB,
occur also in the earlier imperfect state of those species which, in
the perfect form, are not provided with them.
The hardness of the integument of these animals renders it less
probable that they breathe by the skin. Probably they have an
intestinal respiration, such as occurs also in other animals.
Compare M. T. Bruxnnioh ErUomologui, Haffnis, 1764, 8vo. p. 84, with
the expUmation of fig. 4 {Pycnogowum) of Tab. ; various figures are found
in MusLLBB Zool, danica Tab. 1 19, in Lkaoh Zoolog. Mitedlany, i. PL 13,
19, Savioitt MSm, i. l. Anl «. v. i. p. 54, PL v. fig. 3, Gu^Bur leonogr,,
Arachn. PL 4, &a
G. J0HN8TOK Mag. of Zocl. and Botany ^ I. 1837 ; A. Philiffi Nea^i-
tanitehe Pycnogoniden in Ebiohson's Archiv f, Natwrgetck. ix. 1843,
s. 175 — 183, Tkb. IX. figs. I — 3; H. Kbobteb NaturMst, IHduhnfl, m.
1840, pp. 999 — 306, Tab. m. and ibid, new series, i. 1844, pp. 90 — 139 ;
H. D. GrOODBiB On the tpeeific cmd generic ehara^ien of the araneifonn
CruttaceOf Ann. tfnat. ffiet, xiv. pp. i — 4, PL I; Ds Quatbbfaobs if<^.
ewr rorganieat. dee PyenogonideSf Ann. dee Be. not. 30 S^rie,Tom. TV. 1845,
Zool. pp. 69—83, PI I. II ; DuJABDiK in Oompi. rend. 1849. T. xxix. p. 18 ;
ZsiroKEB in Mubllbb's Archiv f Anat. u. Phye. 1853, s. 379.
A. Mandibles and palps (first pair of maxillae according to
Kroeyeb) none. Accessory feet (second pair of maxilla Kroet,)
in females alone.
^ Fabbioiub considered the parasitic crustacean, named by LiNNJius Oniecue CMi,
to be a species of this genus Pyenogonum.
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ARACHNOIDEA. 573
I\/cnoffonum Bbuenn. Body depressed. Feet not longer than
body, thick.
Sp. Pycnogowum UttorcUe, Phalangiwn halamaftim L., BaBTEB Nat, UiUp.
n. Tab. zn. fig. 5, A— D, Pallas Mite, Zool. Tab. xiv. figs, si— 43,
Gu^BiN Jconogr., Arachn. I^ 4, fig. i; this animal, with other marine
productions, is often cast upon our beach dead. The pair of feet that the
female poaseasee aboye the nude was aheady noted by Bastxb as a sexual
difference.
Ptoxichilua Latb. (in part), Edw. Bodj linear, feet slender,
very long.
Sp. Proaoichihu tpmomu Moirr., Cuyhb JZL Ani. 4d. iU., Araehn, PI. 11,
B. Two mandibles chelate, palps none.
Phoxichilidtum Edw., Kkoeyer {Orithyta Johnst.) Accessory
feet proper to females abne, without claws.
Sp. PhoadchUidiumfemoraium, Nymphon femorahim, J. Rathkb ShrivUr of,
fuOurh, Selikabd, v. i. Kiobenhavn, 1799, p. 301, Fl. v. fig. H.
FaUene Johnst., Keoetee. Accessory feet in both sexes ter-
minated by a claw.
Sp. PaBene tpini^, Pffcnogomuti apUUpeM 0th. Fabb., Panma OroaU. p. 454.
C. Two mandibles ; two palps. Accessory feet, oviferous in
both sexes.
Zstes Kboeteb. Bostrum very large, biarticnlate. Feet scarcely
longer than body. Mandibles filiform, short, not chelate.
Sp. ZeUi hitpiduB Kbobtxb.
Partbcea Philippi.
Nymphon Fabr. Bostrum uniarticulate, elongate. Mandibles
chelate. Feet very long (surpassing the length of animal three
times or more).
Sp. Nymphon grattipes, Pycnog. groitipes O. Fabb. (in part), GuiBiK
Iconogr,, Arachn. PL 4, fig. 3.
ATnmathea JjRAxm.
Sp. Amm. carolinamt Leaoh L 1. Tab. 13, Eneydop, mith. Cruit, et Ins,
PL 327, fig. 6.
D. Mandibles none, two palps.
Endeia Philippi^
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574 ciiASS IX.
Ordeb II. Colopoda.
Feet veiy short, truncated, conical, indistinctly triarticolate,
armed with four little claws or three, posterior placed at the
extremity of body. Abdomen not distinct from trunk. Two ocn-
liform points in most. Mouth with two styles exsertile, calcareous.
(Androgynous animals).
Family II. Arctiaca. (Characters of the order.)
Here belong some small (microsoopio) animals which live in
mosses, in canals and sluices amongst confervae, and in rain-water in
spouts. We alluded to them above (p. 206) in pasaing, and cited
them when treating of the wheel-animalcule, with which they agree
in the property of reviving after long apparent death.
Compare J. E. Eiohhobn WcMerihieref Dantag, '775 > "««« ^^i^^o^ ^"^ ^'
TUd: BeUr&ge zwr Naihurffetch. der Heintten W<umihiere, Beriin, 1781,
t. 74, Tab. vu. fig. s {der Watferbdr) ; Sfallanzaki, OputcuUt de Phytique
anim, et vSgSi. n. Gen^ye 1777, n. pp. Z49—^S^t ^^- ^' %«. 7, 8 {letardi^
grade) ; J. A. E. Gobsb in his translation of Bonnxt AhktmdL der InadB-
tol. Anhang, p. 367, Tab. 4, fig. 7 (aooording to his dtation in the Joomal :
Nattirfoncher, zx. a, 114.)
O. F. MuxLLXB in Fuisblt, ArMves tPAUomoL Tab. 36, p. 8a,
Aeante vneUtu.
Fb. Von Paula Sohbaitk, Fcmna Boiea, m. Bd., 1803, «. 178, Areti»-
eon, 8. 195, Aretiieon tardigradum.
G. A. S. ScfHULTZE, Maerobiotui HttfeUmdii, earn tab. lith. Berolini,
1834, 4to. ejnsd. EchMeeuM Mlarmanm, cum tab. Eth. Berolini, 1840, 4to.
DoTiai Uhmoire mr lee Tardigradee, Ann, dee So. not, 3e S^e, Tom.
XIV. Zoologie, 1840, pp. 169 — 361, PL la — 18, xvn. pp. 193 — 205, xvm.
pp. 1—35.
Echtntscua ScHULTZE, Emtdium Dotere.
Mtlnesium DoTi^E.
NoU. — DoTXBB devised the generic name in honour of MOiMi Edwabds.
Here belongs the animalcule described by Spallaniamx under the name of
Twrdigradiu, and perhaps ArcUeeon Sohbanxu.
Macrobiotua SCHULTZE.
Sp. Acamu tureelhu O. F. Muxllxb, Ac.
(The affinity with the Aeariwue already rightly perodved by the sagadtj
of the famous Muxllxb, the most distinguished by far of aU the inyestiga-
tors of mierosoopic animalcules previous to Ehbikhibo.)
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ABAGHNOIDEA. 575
N€U, — Gemw Afyeoifoma LxuoK., oontuning epedes that live panriti-
cally in Comotuto, allied, according to JoH. Muxllbb, to the ^retiict, ought
perhaps to be placed amongst the parasitic cnuAwoM, See below.
Section 11. Hohthoraca. Cephalothorax undiyided. .Organs
of respiration in most internal, distinct, with two or more stigmata
in abdomen, serving for the inspiration of air.
A. Trachea the sole instruments of respiration.
Obder in. Acarina,
Cephalothorax conjoined with abdomen not divided into seg-
ments. Mouth in many in the form of a rostellum.
Family III. Acarea. Head terminated anteriorly by labium
emarginate, or by a single bifid process. Palps connate with labium,
little distinct Mandibles chelate. Ocelli none distinct. Feet
mostly terminated by an adhesive vesicle or acetabulum and claws.
Compare on this family and the whole order A. Duois Seehereket awr
VOrdrt de$ AearieM, Ann, det 8e, not,, sec. S^rie, i. Zool, pp. i — 40, pp.
144—174; n. pp. 19 — 63; which author we shall principally follow here,
for want of observatiGns of our own ; also Dujasdik jut lei Aearieiu, in
Compiet fwd. T. xix. p. 11 58, and Ann. de$ Se, na$, 1845, seo. S^rie,
Tom. m. p. 5 ; Nicx>LKT ffid. not, dea Acwrieiu qui te trowfent a/ux tnvirona
de Paris, Archives du Mut, T. vn. 1855, pp. 381 — 48a.
Acarus Latr. Tyroglyphua (Latr. formerly), Gerv. Body
divided by a transverse furrow between the second and third pairs
of feet. Four feet approximate at the insertion.
Sp. Acmnu dometUcut Ds Gekb, Acamu mro L. (in part), Hsrino, Db
GUB, Mim, vn. PL 5, figs, x— xi, Ltohkt IUcherehe$, PI. 14, fig. 15,
Dux^iUL Ckms. gH. 9. I. In$. PI. 5a, figs. 4 — 7 (under the wrong name of
tareopte de la gdU ;) this small mite, liying on old cheese, has in the young
state, like many AcaridOy only six feet.
Myobia V. Hetden.
Sp. SareopUi fMUCidinm KooE, HxBBiOH-SoH.SFnB J>etaaehl, In$. Heft 138,
Tkb. 13.
ffypopus DuG^
See DUJABDIN, Mim, 9, dei Aeariem aan$ haucke doni on a fait U genre
BypopuM, et qwi mmt le premier dge dee Oamatet, in Ann, dee 8e, nai., 30
^rie, Zool, Tom. xn. 1849, PP* *45» ^5^*
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576 CLASS IX.
Sarocptes Latr. Body not fxiirowed between the second and
third pairs of feet. Third pair of feet remote by a large interval
from the four anterior feet.
Nate, — Here are to be referred genera Trichodctctyltu Dufoub and
Olycypkagua Hebing.
Comp. E. Hebino, Di$ KrdemUben der Thiere (1835), Nw. Att. Acad.
Leap. Carol, Nat. Cwrioa, VoL xvni. a, p. 573 — 624, Tab. 43 — 45.
Sp. Sorcoptes scabiei, Acarut siro teabiei L., Bononki OUertfotumet circa
vivenUa &c. Boms, 169 1, fig. 113 — Db Gbxb M^. vn. PL 5, figs. la, 13 ;
DuoBB Ann. det Sc. not, se S^rie, m. PI. xi. B; Gu&iN Iconogr., Arackn.
PL 5, fig. 12; Sohxidt's Jahrlmcher /. Med. 1853, Bd. 76. a. 304. This
species, which has been yery incorrectly confounded with the cheese-mite,
digs into the human skin^, and causes the itch. Other species oocor in
animals. Here belongs, for instance, Sarooptea eqtU (commonly regarded as
the Acarut exuleerant L.) HKRura L 1. Tab. 43, figs, i, 3, Dujabdim
Obterv. au microteope, PL 16.
See on this subject the interesting Medeelinffen van Dr D. H. Yov
Lkeuwen in the Nederl. Lancet 1846, and his treatise Over de Sckarft der
Dieren in A. Numan'b Vee^artten^kundig Magaeyn, vi. 1847, biz. $2 — 159,
met 2 pL HxssLura in ih^ MUnehener mediciniAen Zexbvng 185a, Bd. 76,
has described some new parasitic mites in pUca poUmica of the human
subject.
I^ote, — ^In this family is commonly enumerated an elongate aoi-
malcule found within a few years by Sihok in the sebaceous follicles
of the skin of the ^ce, which scarcely has its place here. In the
shorty conical feet^ it agrees with the Arctiaca a Colopoda,
Compare G. Simon, Uei>er eine in den kranien «. normalen Saar-taeke
det Mentchen lebende MUbe, Mukllxb*8 Archiv f. Anai. u, PkjftiU. 1849,
s. a 18— 337, Tab. XI.
J>emodex foUieulorum, Owen's Lecturet on the comp. Anatomy of the inveri,
Animait, pp. 351, 153, and ed. p. 444, 445. The same, or a nearly alHed
species, was found in the pustules of a mangy dog in numbers^ by Mr ToP-
FINO, OwxN, p. 445.
Dermaleichus KoCH.
Consult Udtertichi der Arachniden-Sjfttemt, Illtes Heft 3, 1843, PP*
133 — 136.
Family IV. Noiaspidea. Body defended by a homy, hard
covering. Mandibles chelate. Palps fusiformy quinquearticulate.
^ That this animalcule resides here, and not in the pustules, was already ezprenly
announced by Ntandxb, Linn. Amoeniiai. Acad. v. p. 95.
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ARACHNOIDEA. 577
Feet gressorial, with coxss scarcely distant, ungaiculate, not pro-
vided with a vesicle or acetabulum.
Ortbata Latr. {Notaapis Hermann).
Sub-genera: Hopophora Koch, Gal'wnna Y. Heyden, Gebv.
(JPdopa Koch, Oribatei and Zetes ejusd.), Damoeua Koch, {Bdba
Y. Heyden), Nothrua Koch.
OoDBult Oliyisb Eneyd, nUthad., InB, vm. pp. 55»~534. Moat of the
species liye in mosses on the trunks of trees ; in some the body is girdled
by » lamella produced on each side ; Aeanu marginatui De Gekb Mint.
vn. PL vm. fig. 6, NoUup, humercUit Hebm. M^m, Apt, PI. 4, fig. 5 ;»
— iVbtaip. alatui HsBM . PI. 4, fig. 6, kc.
Family Y. Ixodea. Body defended by a coriaceous, extensile
covering. Palps sheathing rostrum, with four joints ; rostrum por-
rect, truncated, composed of two lateral parts (mandibles?), denti-
culate at the apex, and a middle part (labium) covered with nume-
rous recurved barbs. Feet with last joint armed with a caruncle
or vesicle, and with two claws.
Ixodes Latr.
Sp. Ixodes ricinm, Acarue ricimu h,, Dji 6sib M^m. vil. PI. 6, figs. i~8,
Ltonkt JReeherch, PI. 6, figs. 1—8, Tick, LouveUe ; these annuals live m
woods, and attach themselves (the female) to different animals, especially
to dogs ; from sucking the blood the body swells in form of a pea. The
female is nearly 5'" long ; the male, much smaller, on copulating attaches
itself beneath the abdomen of the female by means of the sucker, at the
base of which the wua drferenHa open, to the vulva, situated at the fore-
part of the body between the fijrst and second pairs of feet. This singular
copulation was already figured by Di Gkeb ; see also P. W. J. Muellzb
in Gbbmab u. Zikokek Magae, der Entomol, ii. 181 7, s. 378 — 289.
When laying her eggs, the female, according to the observations of Fbiboh,
gives out a clear fluid from her mouth, for fixing to her body the eggs,
which the animal advances to her mouth ; hence the erroneous opinion of
Ghabbdeb that the eggs are laid through the mouth.
Ixodet americanuM, Aeanu mgua Db Gbbb Mim. vii. PI. 37, figs. 1 1 — 13.
Compare 6. R. Tbbvibakub, ZeiUchr^f, PhyHol. iv. 11. s. 185 — 191, Taf.
XT. XYi. (probably Ixodet crencUut Kollab). This and other allied species
known in America by the name of Piquet are very distressing, and some-
times dangerous to man and cattle.
By the presence or absence of eyes (which are wanting iu Ixodet ricinut),
and some cither characters, Ko€H has divided this numerous genus into
several others. See Ebiohsok'b ilrcAtv /. Naturgeteh. 1844, s. 217 — 239,
and UAenickt, 4tes Heft, 1847.
Family VI. Gamasea. Palps free, filiform. Mandibles che-
late, didactylons. Feet terminated by two claws and a caruncle or
vesicle. Ocelli none, or indistinct. (Animalcules mostly parasitic).
VOL. I. 87
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678 GLASS IX.
This fiunily unites some characters of the third family with othBtB
of the fifth. The larvae, or the imperfect young animals, have
only six feet. Here belongs the genus Ccvris of Latbetlus, which
contains mites that Uye on bats, probably young individuals of
Dermany88U8.
Uropoda Latr. Body depressed, with dorsal shield orbicular.
Peduncle deciduous, infimdibuliform at the posterior part, serving for
fixing the body.
Sp. Uropoda tfegetant, Mitte vSgitaUve Db Gkxb Mim, vn. PI. 7, figs. 15—19 ;
KooH in Hebbioh-Sohaffxb DeuUeJU, Im, Heft 188, Tab. 19 ; Gniani
loonoffr,, Araehn. PL v. fig. 10 ; parasitic on beetles. Gbbvaib mentions
some other species, Aptirea, m. p. 9ii.
Pteroptus Leon-Dufour, Celenpea Montagu, Sptntumta V.
Heyden. Body depressed. Palps with last joint longer, oval.
Feet thick, with joints short
8p. Pteropiut vapertUumis, Hebmaitn M6m, Apt6r. PI. i, fig. 14, Koch in
HBBBiOH-SoHiErFBB DevtaM. Ins. Heft 167, Tab. 2$, panusitic on Vaper-
Hlio nochda; the same or a nearly allied species occm« on Veap. aeroHnvi,
See on other species KooH 1. 1. Heft 137 and 188, who joins this and the
preceding genus to the Sarooptidea (our Aearea).
Argas Latr. Bostrum inferior, concealed under the margin of
body produced. First joint of palps longer than the rest. Feet
approximate at the insertion, biunguiculate, with caruncle none or
indistinct
8p. Arffoa r^/Uapua Latbbillb, Argua r^/Uxm Fabb., Bhynehoprion eolvmba
Hbbm . Mhn, Apt. PLVi fig"- 10, 1 1, Kooh in Hbbbioh-Sohaffbb Deutaeld.
Ina, Heft 189, Tab. i ; found in France, Italy, ftc. on pigeons. A species
occurring in Persia, lives in houses, and by its puncture occasions in man
oonvulsions, deliriiun, and sometimes, as is asserted, even death: Argua
peraieua Fisohbb, Gbbvaib Apiirea, m. pp. 199 — 331, PI. 35, fig. 6.
Holothyras Gerv.
Consult Gbbvaib ApUrea, m. pp. 933. Is this its place T
Dermanyssus Duqes. Last joint of palps very small ; labium
acute ; mandibles in males chelate, with outer finger very long, in
females ensiform. Body soft. Anterior feet elongate ; last joint of
feet furnished with canmcle bilobed and two claws.
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ARACUNOIDEA. 579
Sp. Dermanjftsus avium Duo b, Gbbv., {Sma/ndie dea peUig oueaux) Dum^bil
Con$. gin, 8. 1. Int. PL 53, fig. t, Ltonst ReehercK. PL 5, fig. t i, Duqbb
Ann, det Se, nai. 9e S^rie, ZooL Tom. n. PL 7, fig. i ; DugIbb and Kooh
unite with tl^B species the Acarut ChUina Di Gexb Mitn, vn. PL 6, figs.
13, 14, ajAAcaruM hirundinU HBRMAWif, PL i, fig. 13. This species lives
in bird-cages and hen-houses, and sucks the blood of the animals, especially
by night. Other species liye on different birds, on bats, &c.
Gamasua Latr. (with the addition of genera Biro and Mcucro^
cAeZe^ ejosd.). Mandibles chelate, denticulate. Labium trifid. Body
with dorsal shield coriaceous, mostly double. Anterior feet mostly
longer ; second pair of feet in some incrassated.
Sp. QavMuut ccleoptratorum, Acarua coleoptraiorum L., Robbkl Int. TV. Tab.
I, figs. 10—13, ^^ Geer Mim, Yii. PL 6, fig. 15, KooH in HSBBICH-
SOHAFFEB J)eut8chl, Ins. Heft 168, Tab. 19 ; the beeUe-mUe : the body has
an orange-yellow or blood-red colour; many insects, that creep under-
ground, or live in dung, especially beetles (OeolrupeB vemalis, tUreora/riuM)
have sometimes their entire body covered with hundreds of these mites.
This numerous genus contains moreover many species that live on the
ground, in mosses, underwood, and moist pastures. Also the mite which
was met with by Ltonet on the caterpillar of Cottiu ligniperda belongs
here. Beck, PL 6, figs. 11, 11.
Sub-genus : Ladapa, Zercon, Sejus Koch.
CSonsuli Uebenieht da Arachnidenayttemif 3teB Heft, 30 Abtheil. 1843.
Family Vll. Hydrarachnidta. Palps with last joint nnguicu-
late or spinose. Eyes two or four, distinct Feet with broad coxsb,
mostly ciliated, natatory, posterior surpassing the rest in length.
Aquatic animals.
Compare Hydraehna, quoM m aquia Dania pahulriiua ddeaat, deacripaU,
pingi et tabuUa xi. ceneia incwU euravU O. F. Muellxb, Lipsia, 1781, 4to.
The distinction of these water-spiders as a peculiar genus is to be ascribed
to O. P. MUKLLEB. L1NKJEU8 had not received these animals into his
Syatema naiurce; Fabbioiub in his Sntamologia ayatemaiiea, united the
species then known to him with his genus Tnmbidium, and only after-
wards, in his Syatema andiatorum, made room for a distinct genus for these
species under the name of Atax, p. 366. Besides the genus ffydrachna of
MusLLiB we a]so receive into this family a species of Acarua of LiNVAUS,
which is the typua of the genus Limnochwrea of Latbbillb.
lAmnochares Late. Palps scarcely longer than rostrum. Ros-
trum conical, truncated ; mandibles indistinct, with last joint subu-
late. Ocelli four. Legs pilose, four posterior remote.
37—2
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580 CLASS IX.
Sp. AfMnodUwvt holoierieta, Aeartu aqwUieut L. (exeL dtatione Fbibchu ei
SuLZSBi), Boau InM, m. Tab. 15, I>s Gsbb MSm.Yn. PI. 9, figs. 15 — ly,
Koch in HsBBiOH-SoH^rFBR DeuUM, In$, Heft 150, T»b. 44 ; a"' or 3"'
long^ elongate, very soft, during life vennilion-red : this litUe animal creeps
on the bottom of morasses and ditches, and upon watei^plants. The young
animals have six feet, and attach themselres by their sucker to OerrU
lacutt/rU (p. 443) Duoia Aim. det Se, not., sec. S^rie, L p. 161.
Eylais Latr. Palps with basal joints short, penultimate large,
last subulate, armed with spines. Bostrum short. Mandibles ter-
minated bj a moveable claw. Ocelli four, approximate in the mid-
dle of the anterior part of body. Feet long, slender ; two posterior
pilose, the rest ciliated.
Sp. EyhUa exlendeni, Hydrackna extenden*, Muxll., ffydr. Tab. 9, fig. 4,
KoGH L L Tab. ax, si (according to Koch Acana <tquaUeu$ L., which
however is doubtAil, and is in opposition to the citation of B<xbel horn
Luksmvb).
Hydrackna MuELL. (in part), Latb., Duq^s (and Diplodofdm
ejusd.). Penultimate joint of palps forming with the last a forceps.
Ocelli two, or four bigeminous, lateral, distant. Feet ciliated.
Sp. Efdraehna cmaUa Mukll., Hydryfkamtn cmeidm KooH, Dk Gkkb
M4m, yn. PL 9, figs. 11, la, Muxllbb Hydr, PI. 9, fig. i, KooK in
HXBBIOH-SOHJVFXB 2>. /tu. Heft 150, Tab. x6. The young animals of
this and other species with six feet and a sucker, as Musllib had already
remarked (|ni22m «b ovm auhtdi tex tanfmn pedihus etprobatdde tingidari
inulruetot), are described as distinct species of Aearida by AuDOunr, under
the generic name Achlyna (Jf^m. de la Soc. dHid, not. de Parit, i. 1843,
pp. 98 — 109). DuoBS has illustrated this subject by continuous observa-
tions, and has made known the entire life of these water-spiders in its
different periods, Ann, det 8c, fiol. le S^rie, I. pp. 165 — 171. From the
eggs, which have been laid in spring in the sterna of water-plants peiforated
for the purpose, little hexapod animals come to view, with a laige heart-
shaped sucker, which might be taken for a head, but that the eyes are
situated behind it, on the anterior margin of the back. After an interval
they attach themselves to different water-insects {Nqpa, JtetncUra, Dyii§cut),
and, whilst the abdomen is growing and extending itself into an elongated
sac, the feet and the sucker remain of the same size. In thu fonn the
water-spiders were observed by Swammkbdam on Nq^ (Bijbd der Nahmr.
PI. in. figs. lY. and v.), and described by him as the eggs of this animal,
although he had already expressed his doubts whether they were not rather
distinct animals, which had their increase by sucking the blood of the J^^pa
(biz. 930). In this period the animalcules are named Achlyna by AuDOUUr;
Duoia then calls them nymphs; within the skin the perfect animal is
formed, like a fly in the pu^ eoaretata (see above, p. 473). Bubmbibtbb
also, rimultaneously with Duoka, announced firom his observations that
Achlyna is a youtUul form of Hydroehma, Okbh'b /«'«, 1834, s. 138 — 143.
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ARACUNOIDEA. 581
Atax Fab£. (in part), Duo^ (and Arrenurua ejusd.), Palps
subulate, with last joint falcate, or unguiculate. Ocelli two, lateral,
distant. Body in the males of some species narrowed posteriorly
and produced into an appendage truncated or cylindrical {Arrenurua
DUGES).
Add sub-genera: NesoBa, Fionaj HygrobaUa^y HydrochoreuiUa,
AtracHdea, Acercua^ Marica Koch.
Comp. UAemekt da Arachnidengyttenu, mtes Heft, erat AbtheiL 1843,
and P. T. YAJf Bbnbdsv Mecherehet wr TAtax ypnUtphwa, JSydraehna
eonchartim, Mim, de VAcad, rcyaU de JBdgiqite, Tom. xxxv.
Family Vlll. BdeUea. Bostrum subulate, separated firom
body by stricture, often large, resembling a head. Body oblong,
mostly parted by a transverse groove or stricture between the
second and third pairs of feet. Palps inserted at the base of ros-
trum, lateral, divaricate, large. Mandibles terminated by an in-
curved daw or two small fingers. Ocelli mostly distinct, four.
Feet cursorial, attenuated towards the apex, terminated by two
small claws.
BdeHa Lath., Scirtta Herm. Palps filiform, geniculate or in-
curved. Feet subequal.
BdeUa Dugbs, Koch, Ammonia Koca, Seirua Dxrcns, Kooh,
Eupaltia Koch.
Gump. Vebenieht da Arachnidentsfitemi, mtes Heft, 3, pp. 73 — 80. — Sp.
BdeUa vulgarit Latb., Hsbicann M4m, Aptir. PL 3, fig. 9, Kooh, Hibbioh-
SoRJBVFXB D. Ins. Heft 167, tab. 8.
CheyUtua Latr. Palps thick, incurved, filiform. Feet slender,
anterior longer. Ocelli none or indistinct.
See figures in Koch, HEsaiCH-ScHiEFFEB, Hefi. 167, Tab. 20,
21, 22.
Family IX. Trombidtna. Palps chelate, with last two joints
opposite, the external (penultimate) representing an incurved
claw. Feet terminated by two small claws, cursorial.
Tromhidium Fabr. (exclusive of some species). Two mandi-
bles homy, unguiculate, included in labium. Body suboval or
^ In the neighbourhood of Bydrobata KooH prohMy Ponlaraehna PBiLim ought
to be placed, if it ie to be considered a distinct genus ; ayery sniall Bjfdraehina obienred
in the bay of Naples. AnnaU tf nal, ffitUtry, Ti. 1841, pp. 98—100.
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582 CLASS IX.
oblong, divided into two paxts ; two anterior pairs of feet placed at
the anterior part, remote from the two posterior. Ocelli two, auh-
pednnculate.
« Sp. Trombidium hdoserictwm Fabb., Acotm hcHouriceia L., Db Gbsb Mi1^,
Tii. PL 8, figs. 19— 18, Hbbmann Mim, ApUr, PL i. fig. a, andPLii.
fig. I, Tbbyibanub Verm. Schr. i. Tab. v. fig. a8; scarletried, felty, almost
quadrangular: this little animal may be often seen^ espedally in spring, on
the ground in gardens or in mosses on the stems of trees.
Trambidium tmdorium Fabb., Slabbbb Natuvrh VerUutig, Tab. n.,
Hbbmanh M^m. ApUrol. PL i. fig. i, I>uk6bil Cans. gH, f. 2. /im. PL 55,
fig. 4, from Guinea.
The young individuals of different species live pansiticaUy on insects.
These are the six-footed Acarida, of which Latbeillb formed the genus
il stoma {Gen, CrvM, et Ins. i. p. 163). On Phalangiwn also audi hesapod
parasites are met with (Genus Leptta Laxb. L 1. p. 161).
Stigmams Koch. With ocelli indistinct
Erytkrceus Latr. {Rhynchohphus Duo^S, KocH, Hrythreaa
DuGES, Koch).
Note. — Body undivided. Four posterior feet not very remote from
.. anterior. Other characters almost of Trombidium.
On some other divisions see Koch UeberHcht, m. 2, pp. 41 — 60. Here
also is to be referred genus Tetrcmchus DurouB {Ann. des Se. nai. zxv.
183a, pp. 276 — 185)1 ^ which division belong Trombidwrn idarium, T.
tUiarum, T. Socium Hebm., and other minute species of Acari, forming
fine close webs that invest and suffocate shrubs and trees.
Order IV. Phalangita.
Cephalothorax conjoined with abdomen. Abdomen ringed or
transversely folded. Palps filiform.
Family X. Phalangita, (Characters of the order those of the
single family). Mandibles didactylous, mostly exsert. Feet elon-
gate, terminated by a single claw.
Trogulus Latr. Cephalothorax produced anteriorly into a
clypeus, covering the organs of the mouth. Ocelli two at the base
of clypeus. Palps filiform, not imguiculate at the apex. Tarsi not
elongate, with three joints. Body elongate, depressed.
Sp. Trogylus nqpcrformie Latb., Oener. Onid. et Ina. Tab. vi. fig. i, GuiBiH
Iconogr., Ina, PL 4, fig. 6. To this genus belongs also PhakBugium trieari-
natum L. and Fabb., but is, according to Koch, a different species from
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ARACHNOIDEA. 583
the finrmer : see other species of this genus figured and desoribed in KooH'B
Araehn, T. Bd. pp. 138 — 149. Tbb. 176—179.
Cryptostemma Guia. With ocelli none or indistinct ; ^th tarsi
composed of four or five joints ; with mandibles exsert See GufsiN
Rifoue Z06L 1S38, p. 11, Oertaib ApUres^ m. pp. 130, 131, FL 47,
fig. 4.
Note. — Genus Ootculut LjioN Dufotjb Ann, det 8e. not. zxv. 1839, pp.
989 — 396, PL 9, figs. I — 3, is referred here by its author himself, by
GuiBlK and others ; but this scarcely seems to be its place.
Phalangium Latr. (species from genus Phalaryium L.). Ce-
phalothorax not produced anteriorlj. Mandibles exsert. Palps
unguiculate at the apex. Tarsi mostly with several (6 — 8, or very
numerous) joints. Ocelli two in the middle of cephalothorax,
seated on a common tubercle; two others accessory in many,
lateral, remote, situated more forward.
Gcnyl^ptea Kibbt. Palps spinose, dilated Coz» of last pair of
feet very broad Dorsal scutum homy, hard (Tarsi mostly with
not more than ten joints. Posterior feet longer than the rest),
Sp. ChnylepUs horridut Kibbt Transact. Linn. Soc. xn. PL 72, ^g. 6,
Centurie d'Int. Paris, 1834, PL 4, ^g. 8, Bnai^-^Oonyl. curvipet GuiBOr
Iconoffr., Araehn. PL 4, fig. 5, Chili, kc
Note. — ^All the species are exotic; in some the posterior legs are very
long, slender, and surpass the body three times or more: Mcutigopvt (genus
ineditum Musei L. B.) or Mitobaies Sundev. Contpect, Araehn. p. 54.
Pebtt and Koch have proposed several other genera, on which see KooH
Ueberticht, 2 Heft^ pp. 8 — 22. Grenera Cosmelus and Ditcotama PiBTT
form the transition between Q<mylept€» and Phala/ngivm.
PhcU<mgvum (species from genus Phakt/ngivm auctor). Posterior
coxffi scai'cely thicker than the rest. Feet slender, with tand having
numerous joints (10 — 15 or more) ; second and last pairs sub-
equal, longer than the others.
Sp. Phalangium opilio L., OpUio parietinu9 Hebbst, Kooh, Db Gebb Mhn.
vn. PI. 10, fig. I, Hahn Arachnid, n. PL 69; — Phal. comtOumlj., Oeraa-
Uma comtUum KooH, Db Gebb ib. fig. 19, Hahn Araehn. n. PL 70,
Hebmanb Mim. ApUr. PL 8, fig. 6 (Geoffbot and Latbeillb hold the
two for one species, and the last for the male of Opilio; Hebmakv, Tbb-
yiBAiTOB, Hahb and Koch consider them to be different species. These
animals {harvettspiders, hatterd-tpinnen hooiwagens, faueheura) run very
rigidly; they lurk in chinks of walls, live on the ground between stones,
on trunks of trees, Ac. The long thin legs, after their separation from the
body, present for a long time indications of remaining irritability. The
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584 CLASS IX.
anatomy of Phalamgiimn OpUio has been deacribed by 6. R. TsB^nLAVus
Verm. Schr, i. 1816, a. 30 — ^40, and A. Tulk Aim. and Magaz. of not
Huit, VoL xn. 1843, pp. 153—1^5, PP. «43— «53* PP. 3i»— 33i. PI- 3—5-
Ncte. — Several genera have been proposed by KooH for the azrangement
of the numerouB apeciea of Pkalanffiwui; aee Uebeniehl, 2te8 Hefl^ a. 43 —
SB.—Pkalangodet Tbllkampv ia diatingoiahed by the defect of ocelH.
Order V. Bfeudoscorpiones.
Cephalothorax conjoined with abdomen. Abdomen annulate.
Palps large, terminated hj a hand didactjloos (chela)«
Family XI. Pseudoscarpianes. (Characters of the order those
of the single family). Palps longer than feet, thicker. Feet mode-
rate, terminated by two claws. Habitus of little scorpions without
tails.
Chelifsr Geopfr., Latr. {Ohisium Illig.), species of PhaUm-
gium L. The single genus. (Mandibles didactylous. Ocelli two
or four, distant, lateral).
Chdi/er Leach. Cephalothorax parted in two by a transYerse
furrow. OceUufl single on each side.
Sp. CKdifer eancroJdes, Pkatangivm canenAda L., BosKL Int, m. Tab. 64 ;
the hooh'tcorpion ; brown red ; the ahear-shaped palps are twioe as long aa
the body ; the body without the palp ia scaroely i'" long. TioB little animal
livea in dark and moiat places in houses, between books, &c., and feeds on
mites and wood-lice.
Obiaium Leach. Cephalothorax undivided Ocelli on each ade
two.
Comp. Lbaoh Tran9(ui. of the Linn. Soe. XI. p. 391 ; LxAGH On Ae
Charaeten of Scorpionidea, with de$cripiiont of the Britiih Speeie8 qf C^ifer
and Obitium, Zocl, MitceU. ill. 18 17, PI. 4S— 53; Db Thus i;€Mr« d M.
AuDOunr iur qwlquu Arciehnidea, Ann. dm 8e. not. xxrn. 1834, pp. 61 —
78, PI. 1—3 ; KooH Arachnid, X. Bd. 3, 4 Heft (new genera ChAiomm
and PekruM),
Order VI. SoUfugiB.
Cephalothorax distinct from abdomen. Abdomen annulate.
Palps filiform, porrect, of the length of feet.
Family XII. OaUodea. (Characters of the order those of the
single &mily).
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AlUCHNOIDEA. 585
Galeodea Oliv., Latr. {Solpuga Lichtenst., Fabr.). Mandi-
bles ventricose, large, porrect, with chelae vertical, the lower finger
moveable. Palps with apex rounded, clawless, longer than the
first pair of feet. First pair of feet unarmed, remaining feet
armed with two claws at the apex. Ocelli two, placed in a
common tubercle at the middle of the thoracic shield, near the
anterior margin. Body villose ; legs and palps covered with long
scattered hairs. Posterior coxae famished at the inferior margin
with a row of appendages membranous, triangular, petiolate.
Sp. Oaleodet araneo^deSy Phalangium arcmeoUiei Pall. SpieU. iz. Tab. m.
figs. 7, 8, 9, DuM^iL ConM. gin. t. I Jn». PI. 55, fig. 3, Southern Buasu^
Greece, ice., and many oiher spedes, especiaDiy firom Africa. See Koca
in Ebiohbon'b Areh.f. Naiurgeseh. vni. 1S42, s. 350—356. Most of the
species are from the old world ; there are, however, American species also
enmnerated by KooH, to which may be added OaUodea Umbtda and O,
Oabat, Lucas in Gu^bin Mag, de Zool, 1834, Araekn. PI. 5, 1835, Araekn,
PL n. These animals appear to reside in warm sandy regions, and to come
frt>m their Inrking-places by night especially. On the dorsal sur&oe of the
upper-jaws some have a leaf -like appendage, whidi, according to Gbbvaib,
is a character of the male.
According to J. Muellxb, besides the two laiger eyes, there are in an
Egyptian species two smaller eyes on pedicles, and two lateral eyes {Ver^.
Phynol. da Oesiehtsinnes, s. 333) ; in the specimens investigated by me I
could not perceive these eyes.
The bite of Oaleodet is considered to be very venomous and dangerous;
Olivibb, who met with many of these animals in Persia and Arabia, was
not himself bitten, nor was any one of his companions, by them, and was
not able to discover any sure proof of the reality of the danger. Voyage
dans V Empire othoman, Paris, Tom. vi. 1807, p. 306. (Gomp. also the
figures there, PI. 4a, figs. 3—6, and in SoKNun Vogage en Cfrice, Paris,
1 80 1, PI. 3 ; the latter traveller found his specimen On the island of Qypms.
Seel. pp. 115 — 114.)
Sub-genera: Sdpuga, (?a20(x2^«, ile{2optMKooH (with jointed tani),iZAate,
Qluvia ejusd. (with tarsi not jointed).
B. Respiratory organs either lungs or trache», together with
longs. Cephalothorax distinct from abdomen.
Order VII. Pedipalpi.
Palps large, resembling feet, chelate at the apex. Abdomen
divided by segments. Pulmonary sacs without trachea ; eight or
four stigmata.
Family XIII. Phrynides. Abdomen separated from thorax by
a slight constriction. Stigmata two on each side at the base of
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586 CLASS IX.
abdomen. Mandibles monodactylous. Palps spinose. Ocelli eight,
situated at the anterior part of cephalothorax ; the two middle
approximate on a common tabercle ; three on each side marginal,
disposed in a triangle. Two anterior feet longer, with tarsi claw-
less, slender, multiarticulate.
Phrynus Oliv. (Species of Phalangium L., Tarantula Fabb.
in part). Palps supplied at the apex with a homy claw, of the
length of body or longer than body. First pair of feet slender,
very long, resembling antennse, with tibiee and tarsi multiarticulate.
Body depressed. Cephalothorax broad, semicircular, emarginate
posteriorly.
Sp. PhrynuB luncBttu, Phalangium htnatum Vall., Pkalangium ren^orme
L. (in part), Pallas SpicU. ZoU. ix. Tab. 3, fig. 5, Hxbbst Naiun^ der
ung^. Int. I. Berlin, 1797, Tab. m., Latb., Hid. not. des CruH. et de$
Ins. PL 61, fig. I ; — Phryntu rm\formu, Phalangmm rmiforme Pall, (not
L.), 8pie, Zool. Tab. dt. ^g. 3, DuiciRiL Chm. gSn. $, L Ins, PL 56,
fig. 1, &c.
Gomp. J. Van Deb Hoevkn, Sijdragen tot de kennis van het gedaeht
Phryrvus, Tijdsehr. voor not. Oesehied. en Physiol, ix. pp. 68 — 91. PL I. n.
TelyphontL8 Latr. (Species oi Phalangium L., Tarantula Fabb.
in part). Palps thick, terminated by hand didactylous, shorter
than body. First pair of feet with tarsi eight-jointed, and tibia
with two joints. Cephalothorax oblong, oval, not broader than
abdomen. Abdomen terminated by an articulate seta.
Sp. Telyphonus pro8Corpio Latb., Phalangium caudaium L., Spic Zool, ix.
Tab. 3, figs. I, 2, Gu^BiN Iconogr., Arach. PL 3, ^, 3 (is it the same
species 1) ; hah. in Java.
NoU, — The species of this, as well as of the preceding genns, are difficult
to distinguish. They are found in tropical regions of both hemispheres:
none European is known. On the TeUphoni oomp. Luoas in Gu&iiir
Magas. de Zool. 1835. Arachn. PL 8—10.
Family XIV. Scorpianes. Abdomen sessile. Stigmata four
on each side, in the second, third, fourth and fifth ventral scuta.
Mandibles didactylous. Palps longer than feet, terminated by
hand didactylous with external finger mobile. Six last segments of
abdomen abruptly narrowed, resembling a tail ; last segment vesi-
cular, terminated by an incurved sting. Feet increasing in length
firom the first towards the last pair, moderate, all biunguiculate at
the apex. Two pectinate appendages, with teeth different in number,
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ARACHNOIDEA. 587
at the inferior part of body behind the coxa of the fourth pair of
feet
Scorpio L. (Characters of the family). Ocelli two in the
middle of cephalothorax, approximate, larger. Lateral ocelli in
the anterior margin of cephalothorax, number various.
The soorpions live in wann regions of the temperate zone and in
tropical countries. In the last joint of the abdomen is a poison-
gland, which renders the wound of the scorpion dangerous, (see Bedi
De GenercU. Ineedor. Amstelod 1686, pp. 84 — 91 ; De Mauper-
Tuis Experiences auar lea Soorpions, Mem, de VAcad. de Pcvrisy
powr 1731, p. 223). Moreover these animals are distinguished by
two comb-shaped appendages at the base of the abdomen. These
appendages consist of a flat pedicle that runs fcransversely outwards
and becomes thinner towards its apex ; this on its inferior margin
bears parallel lancet-shaped leaflets (teeth), standing perpendicular
to its axis. The number of these teeth diflers in difierent species,
yet is not sufficiently determinate but subject to change, so that it
it is an uncertain character for the distinction of species.
Oomp. on the divifidons and the species of this numerous family, LiAOH
TrcaiB, of the Linn. Soc. xi, p. 391 ; Hbmpbioh and Ehbbnbbbo Symb,
fhytie., Everts, I. Ara>chnofdea, Berolini, folio 1828, cum tabulis 2; P.
Gkbvaib Remarquea sur la famiUe dea Scorpions, Archives du Museum,
Tom. lY. 1845, PP> ^01 — ^4^* ^^' ^^* ^^' ^^ ^® number of the eyes alone
no natural groups or sub-genera can be founded. The hoibUus, the greater
or less breadth of the six last abdominal rings, and the form of the forceps,
indicate better sab-divisions, or ought at least to be conjoined with the
character from the eyes.
Androctonua Ehbenr With twelve ocelli, five lateral on each
side. Centru/rus Ehsenb. With ten ocelli, four lateral on each
side. BtUhue Leach, Ehrenr With eight ocelli, three lateral on
each sida Scorpio Leach, Scorpiua Ehbenb. With six ocelli, two
lateral on each side^
Note. — Sub-genus JButhus, far from natural, ought to be distributed into
other sub-genera, after the example of Ehbekbkbo and KooH; comp.
Gbbtais L L, who admits three sub-genera Ischnwrus, .Sv^Aiwand TeU^onuM.
Sp. Scorpio ewropcBut, Scorpio Jlavicaudus, Di Gebb M6m. vn. Fl. 40, figs.
II — 13, BoBMErrat Elea^ Entom. Tab. 113, Guy. H. Anim. id. ill., Arachn.
PI. 19, fig. 2, hab. south of Europe and north of Africa; — Scorpio afet L.,
Koubl Ins. m. Tab. 65, Koch Arachniden, m. Tab. 79, &c.
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588 CLASS IX.
Order VUL Araneidea.
Palps subfilifomi, with last joint in males supplied with varioos
appendages, subservient to copulation. Abdomen coveied with
skin continuous, mostly soft, constricted at the base or joined to
the oephalothorax bj means of a petiole. Stigmata ncTer more
than four, mostly only two* Respiration in all pulmonaiy, in some
tracheal at the same time.
Family XV. Araneidea {Aranece Latr.) Characters of the
order also those of the single iamily.
(Mandibles monodactylous, with terminal claw perforate, for the
excretion of a poisonous liquid. Four or six papillae cylindrical or
conical at the inferior surface of abdomen, situated towards the
posterior part, perforated by very minute foramina for the passage
of a silky substance. Feet different in length, similar in form,
terminated by a double or triple claw.)
The spiders. Ail these animalg prepare from a silky substaiioe
certain filaments with which they cover their egg^ Many, fixnn
the same Bubetanoe, form alao webs and nets iu which they capture
their prey. At the hind part of the body four, or in most spedefl,
six spinarets are found, beset with fine tubules through which the
silky matter escapes. The secretion is effected in the form of an
adhesive fluid by means of glands, or tubes of very diff^ent form,
pear-shaped, glandular sacs united in groups, blind tubes convdated
and ramified. See Tbevirakus Ueber den innem Bau derAraehmcL
s. 41—44, Tab. iv, v. figs. 42 — 44 ; Verm. Schr. l & 11, 12, Tab. i.
fig. 4, H. Meckel Arch. /. AnaL u. Physiol. 1846, s. 50^-^6, Ta£
lu. figs. 38 — 45.
The long threads that cover the fields or float in the air, espedallj
in the fall of the year, {Gossamery Her/stdradeuy JUs de la Vierge,
Herbstgam, der fliegender Sommer) are connidered by some writen
to be products of the atmosphere, or exhalations firom plants ; the
chemical investigation of G. J. Mulder has proved that these
threads agree in composition with silk, and beyond doubt are the
work of spiders.
See NiUwi^'m Scheibundig Archirf, and a postscript by my hand, in
which I have cited some works on this subject. Latrbilli ascribes these
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ARACHNOIDEA. 589
threads to young spiders. lUgne Anim. i dd. lY. pp. 319, 410. Comp. J.
Blaokwall Catalogue pf Brituh Spiden, imdvding Remarki on their Strue^
iftre, Fwnetiona, JBeonomy and SyttemaUe ArrangemetU, Awn. nai, ffitt, md
Series, vn. — IL, 185 1 — 1853.
Phalanx I. Areneas tetrapneumones, Mygalides SuNDEY., Koch.
Spinarets four, two of them large, exsert, two others veiy short.
(Stigmata four, leading to four pulmonary sacs. Ocelli always
eight. Claw at the apex of mandible in the same direction as
the mandible, capable of inflection downwards, large, incurved).
Mygale Walck. Ocelli approximate, situated in the anterior
part of cephalothorax ( .': !.* ). Palps proceeding from the apex
of maxillsB. Labrum inserted under the base of maxillae, veiy
small, quadrate. Feet hirsute, subequal, first and fourth pairs
longer.
Sp. Mygale avievlaria Walok., Aranea avieuiaria L., Klksxann BeUrdge,
Tftb. zi. xn., Ds Gkbb Mim, yn. PL 38, fig. 8; a large spider of 8.
America ; the body i" 6'" and morei, the bind legs 1" $"*; it lires in a
tubular web narrowed behind, in chinks of bark of trees, between stones,
&c.; the female places the web in which she has laid her eggs close to her
nest. It has been asserted that these spiders are able to seize small birds
(humming-birds), and hence the name of this species ; there is, however, no
reason to think that the account is founded on any thing better than fable.
See on the web and mode of life of this animal Latbbillk Mim. du Mut.
vni. 183a, pp. 456 — 460, and W. 8. Mao Lkat TSrans. of the Zool, Sac.
I. 7, 1834, pp. 179 — 194. Other similarly large species are also found in
the eastern hemisphere, as Mygale fcueiata Walok. Hist. not. dea Aran.
IV. PL I ; Ceylon.
Cteniza Latb. Mandible fumished beneath, near the claw, with
a row of homy barbs.
Sp. Mygale eametOaria Lath., Walck., ffieL naJt. dee Aran. m. PL 10: —
Mygale fodnene Walok., Mygale Sauvageeii Latb. : these species of southern
Europe, which certain exotic species resemble, live in vertical tubular
cavities underground, closed by a circular coyer as by a door, and lined
internally with silken web. On the inside of ihe cover are small impres-
nons, to which the spider attaches itself by means of the booklet of the
jaws, holding iast to the walls of the tube with its feet, in order to keep the
door close when an attempt is made to open it; this shuts down by its own
weight. See Sauvaoxs M4m. de VAcad. dee 8c. de Parte, 1758, ffiel.
p. a6, Latbkills Mimoiree de la Soe. d^ffiet. not. de Parie, &c., An. vn.
4to. pp. 118 — 148, PL VI., AuDOUUi Ann. de la 8oc. entom. n. 1833, pp.
69—85. PL 4.
Oletera Walck., Atypm Late.
Eriodon Latr., Mismlena Walck.
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590 CLAJSS IX.
ArUhrchia Tellk. Ocelli none.
Livos in cavema. Ought it to be ooimted of tins fiGunilj %
Phalanx 11. Aranece dtpneumones. Spinarets six. (Pulmonary
sacs only two ; stigmata mostly two, sometimes four, the two
posterior leading to trachea, not to lungs. Ocelli mostly eight,
sometimes fewer. Claw of mandible capable of inflection trans-
versely or laterally to the inner margin of mandible).
Most of the spiders of this division have only two stigmata. In
some, however, four are observed, and on that account the genera
Dysdera and FUistcUa were placed by Latreille with MygdU
(amongst the Tetraprievmonea). But two of these stigmata do not
lead to pulmonary sacs, but to air-tubes, and were also obserred in
Argynoreta by Grube. For the arrangement, therefore, this cha-
racter of the number of stigmata is without weight, because it does
not indicate natural affinity. The number of lung-sacs, on the other
hand, appears to be in harmony with the characters derived from the
mandibles and from the number of spinarets.
A. TubitdcB. Spinarets cylindrical, parallel, collected into
a fiasciculus, directed backwards. Ocelli six or eight disposed
in two rows (in one genus only two). Fourth or first pair of legs
very long \ intermediate legs shorter.
Legs when at rest retracted uptoa/rds, vnth /emora not preesed
agamst the ground. Spiders toeaving vxhe, resemJbling tubes or fisk-
mg-netSf and hidden in chinks, comers, or under stones,
Nops Mac Leay. (The eyes excepted almost similar to Dys-
dera Latr.)
Sp. Nops Onanabaeooi Mao Lxat, Ann. of ntU, Hid, n. 1838, PL i, fig. i,
Cuba. The only species of tpider hitlwrto known with only two eyes.
ft Ocelli six,
Dysdera Latr. Ocelli almost contiguous, arranged in two rows ;
the fijrst row of two, the second of four ocelli. The first pair of
feet surpassing in length all the rest, the fourth pair the inter-
mediate.
Sp. Dyadera erytkrina Walok., Aranea rufipet Fab&., Latb. Oener, Cmtt,
H Int, Tab. v. fig. 3, UARsAraehn, Tkb. I. fig. 3 ; in south of Germany, &c
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ARACHNOIDEA. 591
Segestna Latb. Ocelli arranged in two rows ; the anterior row
of four, the posterior of two remote. Feet as in the preceding
genus.
Sp. Segestria aenoeulaia Waloksk., Aranea ienoculata L., Db Gxkb M^,
vn. PI. 15, fig. 5, Walckbn. AratUid. Tab. I. PI. 7, &c.
Scytodes Lata. Ocelli six, equal, disposed in pairs, forming a
triangle with the vertex forwards.
Sp. Scytodea ihoraeiea Latb. Oener. Crutt, a Ins, Tab. T. fig. 4, Walokxn.
AfXMM, I. PL 10.
Add genera Ariadne Say. and Uptwtea Walgk.
ttt Ocelli eight
Clotho Walcken.
Drassus Walcken.
Aaegena Sundev.
Clubwna Latr. Ocelli arranged in two lines transverse,
approximate. MaxillsB straight, subdilated outwards at the base,
rounded at the apex. Labium elongato-subquadrate, truncated or
emarginate at the apex.
Sp. Cfltihuma holoaericea Latb., Aranea hdUaericea L., Db Gbeb Mim, vn.
PI. 15, figa. 13 — 16, Walgk. ArantUid, iv. PL 3; moaBe-grey, satiny-
shining^ with an elongate pointed abdomen. The female lays her eggs
between one or two leaves spun together.
Anyph(ma Sundev.
Cimflo Blackw.
Argyroi(^ Latr. Ocelli eight, four middle forming a square
(.*::'.). Maxillss straight, rounded at the apex. Labium elon-
gato-trigonal.
Sp. Argynmtta aquaUca Latb., Aranea aquaUea "L., Glbbok Aran, PL 6,
Tab. 8, Db Gebb Mim, vn. PL 19, figs. 5—13, Hahn Araohn, Tab. 49,
fig. 118, HBBBiOH-SoHiBFFBB D. Int, Heft 134, l^b. 91, It; 6"' long; one
of onr largest native species; thorax red-brown, abdomen blackish, the
first pair of legs longer the rest. This spider lives in fresh water; as it
swims the abdomen and the th6raz have a silveiy aspect from the attached
stratum of air. This species spins a bell-shaped, water-proof web that is
filled with air, and open below; this it attaches to wat^r-plants by threads.
See the observations of Db Gbeb loc. oit. p. 303 — 313, those of Db Lignao,
ftc. (of which Walokbnabb gives an ample extract in his HiM. not. de$
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592 CTA8S TX.
ApUre$, n. pp. 380 andfoU.), and of E. Gbubb in Fmovsa^B NmieNaHam,
xixv. 1842, No. XXI, xxn. s. 311 — 348, s, 341—344.
Aranea Latr. (species from genus Aranea L.), Teffenaria
Walcken. Ocelli eight, arranged in two lines transverse, approxi-
mate. Maxillffi straight. Labium subquadrate. Feet of the fourth
and fifth pairs longer, subequal.
Sp. ArtMM domatiea L., Albiv Spiden, PL 18, fig. 87, Clkbok Anm.
PL 2, Tab. g, Walok. ApUre$, PL 16, fig. 2. They weave close, adheuTe,
nearly horizontal weba in the oornera of walla, in chamber-windowa, ftc., and
a tube doae by the net in a lurking-place, in which they wait for their prey.
The anatomy of thia q>eciea haa been treated of eapecially by Tbxvuubub^
in hia claaaical Monograph Ud>er den innem Bau der Araehmden,
Agelena Walckek. (and Ccdotea Blackw.)
Megamyrmakion Reuss^ Dydion Walcken. [Is this its placet]
Filistata Latr.
B. IniquitdoB Latb. Spinaiets conical, convergent. Ocelli
remote firom the anterior margin, not describing a segment of a
circle or a lune. Feet slender, drawn upwards when at rest; first
pair mostly the longest of all.
Spidere mahing neU, arranged irreguiarlf/y wUh threads intersect
ing one another in all direcHona,
Pholcua Walcken. Ocelli, two middle less, placed transversely,
three on each side larger, grouped in a triangle (.•* *.)• Feet
elongate, veiy slender; first pair longest of all, second longer than
fourth, third shortest of all. Maxillas incumbent on labium,
elongate, narrowed towards the apex.
Sp. Pholcui phalangiciidei Walok., Aran. v. PI. 10, Apt. PI. 8, figa. 2, 3 ;
pale yellowiah grey, the abdomen elongate. Thia apeoiea Krea in the anglea
of walla, kc. in hooaea.
Epiaimu Walcken.
la thia ita placet
Latrodectas Walcken. Ocelli subequal, four middle and two
lateral on each side placed on a tubercle, remote from each other.
First pair of feet longest of all, fourth longer than second and
third.
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ARACHNOIDEA. 593
Sp. Zairodeeiui malmignaiu»WALOK.,Aranea 13 guUata BoBSi, Fabb., Rossi
Fatin, etrutca PI. ix. fig. 10^ Walck. Aran. I. PI. 5 ; Uub species is
found in Italy and Gorsicay and named rognc malmigTiato ; it has thirteen
carmine-red spots on the abdomen. Its bite is poisonous, and causes strong
convulsions. Compare A. Raiksm Reeherchea awr It Tkeridion marmig-
natte de VoUenxk, ettwrUa ^Oa da aa morawra, Ann, dea 8c. not. le S^rie,
Tom. XI. 1839, ^<^ PP' ' — *7-
Note. — Latekillk joined this genus with the following, from which it
scarcely differs except in the arrangement of the eyes.
Theridum Wadcken* Ocelli snbequal, mostly small, four
middle disposed in form of a square, two lateral on both sides
approximate, sometimes contiguous (/t**.). Feet of first and
second pair longer than the rest.
Sub-genera: Eucha/ria KocH, Ero Koch, PaehygruUha Sundev.,
Steatoda Suot)EV., Dictyna Sundev. (ErgcUia Biackw.)
Sp. Theridion henign/um Walok., Walckxit. Araniid. v. PI. 8, KooH
Ara^n. m. Tab. 83, figs. 184, 185 ; a small grey spider much found
amongst grapes, and whose fine web protects them from other insects.
BolyphcmUs Koch. {Neriene Blaokw.)
Argu$ Walcken.
Erigone Savignt.
Micryphcmtea Koch. (J^alckeTiaera Blackw.)
C. OrbitelcB Latr. Spinarets conical, convergent. Ocelli near
the margin of the cephalothorax inflected forwards, not disposed in
a segment of a circle or a lune. MaxillsB straight, broader towards
the extremity. First and second pairs of feet longer than the rest,
the first pair longest of all.
Spiders tvecnnng nets orbieulate,/ormed of concerUric circular cmd
straight threcuis, radiaiimg from the common centre. Legs when at rest
drawn tipwards.
lAnyphia Latr. Ocelli four middle, placed in two rows ; the
posterior more remote; two lateral on each side approximate, placed
obliquely. Maxillae substraight, remote.
Compare Walckxnasb Hist, not, dea Apt, n. pp. 233 — 984. A genus
intermediate between this family and the preceding, to which it might per-
haps be more correctly consi^ed.
Mandncrdua Blaokw.
VOL. I. 38
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594 CLASS IX.
UhboruB Late.
Tetragnatha Walck. Ocelli subequal arranged in two rows
transverse, straight, almost parallel. MaxillsB elongate, divergent,
dilated on the outer side towards the apex. Feet elongate, slender.
Epeira Walck. Ocelli subequal, four middle arranged in a
square, two on each side more remote from the middle four, placed
obliquely. Maxillse broad, short.
Sub-genera: Singa Koch, ZiUa Koch, Miranda Koch, AUa
Koch, Argyopes Sav., Nephila Leach, MicrcUhena Sukbsv., Aero-
soma Pebtt, Gasteracantha Latb. {Plectatui Walck.)
Mithras KocH. With two external ocelli of the first row very
small, microsoopia
The net of the spidera of this genus is mostly placed in a Tertical plane,
sometimes obliquely. Only one species is known with which it lies horuraa-
tally. (JSpeira cucurhitana, Aranea cucurhitana L., Glbbok Aran, PL 2,
Tab. 4, Walok. ffitt. d. Aran, ii. 3.)
To this grenua belongs the Cross spider, Epeira diadema^ Aranea dia-
dema h,, Clxbck PL i, figs. 4, $, "RasRh Ins. TV, Tab. 35 to 40 ; Brandt
u. Batzsburo Med. Zool, 11. Tab. xiv, figs, i — ^4 ; with a broad, triangular,
toothed band of a darker colour on the dorsal surfiEuse of the abdomen, and
a triple cross of whitish-yellow spots upon the band ; the female has a lai^
oval abdomen. This species, everywhere known, is common in gardens,
especially in autumn.
Of the exotic species some have a resemblance to small crabs ; they have
a very hard homy sldn on the abdomen, which moreover is armed with
spines or long points ; Oasteracantha Latb. 8p. Epeira cancriformitf
Slabbbb Nat, Verlust, Tab. i, schulpspin, Walok. AranSid. ni. PL 4, Ac
D. Laterigradm Latk. Ocelli mostly arranged in a Innated
form or in a segment of a circle. Body depressed, with cephalo-
thorax suborhicular, small or moderate. Four anterior feet mostly
longer than the rest,
Spidera not weaving a wehy btU either drawing ovi some sparse
viscid strings, or constructing a house amongst leaves contorted at the
margins. Feet at rest extended, with femora pressed on the ground,
Selenops DuFOUR.
Sparassus Walck., Micrommata Latr. Ocelli arranged in two
rows, the anterior row narrower, convex (/. . ; ). Maxillae straight,
distant, parallel.
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AILiCHNOIDEA. 595
Sp. SpaimtuB imaraffdtUfu Walok., Aranea mnairagdimL Fabb., CIiBbok
Aran, PL 6, Tab. 4, and 7, Db Gbbb Mhn. yn. PL 18, fig. 6.
Olio8 Walck., Sarotes Sundev.
Philodromus Lat£.
Thomisus Walck. Ocelli subequal, arranged in two rows
convex anteriorly, the posterior row broader.
Xystictu Koch. External oceUuB of the anterior row large, the
rest small, subequaL
Compare KocalUAernchi de* Aracknidemyftema Entes Heft, 1837, 8.
IS, 16.
E. OttiffradcB Latr. Ocelli occupying a larger area, arranged
in form of a curvilinear triangle, or of a trapezium, or of an ellipse.
Cephalothorax lofty, ovate, narrower anteriorly.
ErrcUic spiders, not canstrvcHng webs or nets to capture prey,
pvo'suing their prey by running.
The femalee of many spedee sit upon the web of eggs to guard them, or
bear their eggs about with them in a round sac. They also protect their
newly hatched young for some time.
Ctenus Walck. Ocelli arranged in three rows 2, 4, 2, the last
two more remote f. ."..V
Most of the species American ; Comp. Walobbv. Aj^ i. pp. 363 — 370.
Dolomedes Latr. Ocelli unequal, arranged in three rows, in
the anterior row four, all or the two middle being smaller, in the
second and third rows two larger ; ocelli of the third row most
(.. .• \
.• •. )•
8p. IMomeda rnvrabUii Latb., Aranea dbtcwra Fabb., Clbbck Aran, PL 5,
Tab. to, Db Gbbb MSm. vn. PL 16, figs, i— S, Walck. AromHd, i. PL 9 ;
the female carries the sac of eggs with her under the thorax, holding them
&Bt with her jaws and feelers.
Sub-genus Oqfole Say., Sundet.
Lyoosa Latb. Ocelli unequal, arranged in three rows almost
equal, the first row having four smaller, the others two larger
(;■•)•
38—2
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596 GLASS IX.
Theae specieB live on the ground, and run very rapidly. The females
oany their egg-sac with them, which is attached by a few thxeads to the
abdomen. Sp. Lyeoaa aaecaia It., Fbisoh Ba^r. v. aUerl. Ins, vm. Tab.
2, Clbbok Aran, Suee, PI. 4, IVb. 8, fig. a, KooH in Hkbrioh-Sohafpsb
Deutschl. Ins, Heft 110, Tab. 8. This species is the most common in the
north of Europe. To this genus and probably to this species is to be
referred the account by Bonitbt of a contest between a spider and an ant-
lion, into whose hole it had fallen. Oontemp, de la not,, xnikne Parties
chap. 42, {(Eu/vrei compl. id., 8vo. Tom. ix. pp. 411, 413.)
To the genus Lycota some larger species also belong ; these are the cele-
brated TaaurdvlcB (Aranea Tarantula L.) of the south of Europe and Tariooa
other regions of the old world. They are of diflPerent species. The true
Tarantula of Italy and Spun (Lyeosa tarantvla Apulia Walck.) is figured,
amongst others, by Albik Spiden PL 38, Hahn Ara^nid, i. Tab. ^3,
Gu&iN Jconogr., AracKn, PI. \, ^, 6 \ according to Walokbbabb that
found by L^N Dcfocb in Spain is a different species, Ann, da Sc nal,
ae S^rie m. 1835, PI. 5, ^, 1 ; this spider, according to his observatioD,
lives in holes in the earth, of which the outlet is surrounded by a little
raised wall that is covered on the inside with web. That the bite of thiB
spider is said to be followed by a dancing madness, that can be cured by
music, is as well known as it is incredible.
Sphasus Walck., Oscyopes Latr. Ocelli unequal disposed in
four pairs, those of the first pair approximate, of the third pair most
remote of all l.*\) •
(The genus is allied to the next subdivision, and perhaps might
more correctly be placed there.)
Sp. Sphanu heUroptkalmua Walok., Oxyopet vatriegatw Latb., Qtn, Orud, ^
Ins, Tab. 4, ^, 9, Walck. Hitt, d:Aran. m. PI. 8, &c.
F. Salttffradce Latr. Ocelli unequal, arranged nearly in the
form of a square ; the anterior lateral placed towards the angles of
the anterior margin of cephalothorax, the two posterior remote from
each other by the whole breadth of cephalothorax, or opposite to
the former. First pair of feet with thighs mostly thick. Thorax
large, deep, oblong.
JErraUc spiders, not forming nets, puntUng their prey by running
cmd rushing upon it with a leap,
Myrmecium Latr. Ocelli arranged in three rows ; the anterior
of four ocelli, with two middle larger forming with the second row
a square; the two last distant, set on tubercles. Cephalothorax
divided by strictures, elongate. Abdomen small. Feet slender.
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ARACHNOIDEA.. 597
8p. Myrmedum rufwn Latb., AfmaL da 8e. not. ni. 1814, pp. 33 — 97, PI.
4, from Brazil. Other speoies, all from AmericA, are noticed by Waloksn.
Apt. u pp. 386—388.
Chersia Sav., Walck., {Paljnmanua Dup.)
Eresua Walck, Two middle ocelli of the first row forming
with the second row a square, inscribed in a larger square, formed
by the lateral ocelli of the first row and the two last ( ' •• j . Feet
thick.
Sp. Ere9fu HnnaberiwuM, Aranea moniliffera Yillkrb EiUom, Linn, Tab. xi.
fig. 8, Walok. AranHd, n. Pt 10, south of Europe, &c.
AUu8 Walck., Salticus Latil Ocelli arranged in form of a
square open behind or of a horse shoe, with two posterior remote,
opposite to the laterals of the first row ; two middle anterior ocelli
larger, the two of the second row smallest of all (*•' * •") •
Sp. AUut temieua Walok., Aranea sceniea L., Albin Spiders PL m. figs.
13, 14, Db Gexb MSm. vn. PI. 17, figs. 8—10, Panzkb Deuttehl. In»,
Heft 40, Tab. as (called Aran, cingulatce) &c.
This genus, with ^aeira the most numerous in species in the whole
division of spiders, presents, in the relative length of the legs, and in the
relative size of the different ocelli, some variations. See Walok. Apt. I.
PP« 483 — ^488 and SuinnvALL Comped. Arachnid, pp. 95 — 97.
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CLASS X.
CRUSTACEANS {GRUSTACEA)\
The class of crostaceans is the last of articulate animals with
articulate feet (see above, p« 248), all of which LiNNiEUS regarded
as insects. They breathe by means of gills, and there are no air-
passages or stigmata present as in insects. They are distingaished
from the preceding class by the presence of abdominal feet. The
myriapods alone agree with the crustaceans in this respect, and are
by some writers enumerated amongst them, with which, however,
on account of the presence of air-passages and air-tubes and of their
resemblance to insect-larvse, they ought not in our opinion to be
united.
The name of the class is derived from the nature of their
external covering ; this is in most cases hard, and contains a greater
or smaller quantity of carbonate of lime. In other instances, how-
ever, the integument is more leathery or homy. The tissue which
supplies a foundation for the hard calcareous shell, is still the same
chitine which forms the dermal skeleton in the entire division
of articulate animals with articulate feet (p. 284). The shell
consists of different layers, with a layer of pigment beneath or
equably penetrated by colouring matter. Between the rings
^ Besides the works of Latbeillr already cited under the insects, p. 147 (W*-
not. det Ortut. et da Ins,, and Otnera Oruttaceor. H In$eotor.) we wonld mention as
principal works: —
J. P. W. Hebbot, Vertuch einer NaturffetchichU der Krabbm und Krtbie, m. Bd.,
m. Ulum, Kwpfertaf, 4to. Berlin u. Strolsund, 178a — 1804.
SuoKOW^ Anatomischrphynologische UrUersuchungm der Insekten vnd Kriideniki^
Heidelberg, 18 18, 4ta
A. E. Dbsmasbst, C(wsidSrati4mi ginSraJUi iur la cUum dei OntitacSt, Tm
1825, 8vo. avec 56 pi. In this work (pp. 396 — ^420) there is a copious list of writings
on the class of the crustaceans.
MiLNB Edwabds, HUUnre natwreUe det enaUusis, ay. pL Paris, 1834— 1840, ni.
Vols. 8vo.
Th. Bbll, BiHory of Sritish Crtutacea, London, 1853.
Also the article Oruttaeea by the same author in Todd'8 CsfdopcBdia I. pp. 75^^
787 may be consulted. For the fossil species Bbogkiabt et Dbsmabbst, Sid. n^-
det Crtutaoii fotiiUi iout la rapporte zoologiquet et ffMogiqu/te. Paris, 1821, 4to.
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CRUSTACEANS. 699
(segments) and in the joints alone is the integcunent thinner and
more flexible.
Let ns first consider the external structtire of these animals
a little more closely. We have seen above that the body of insects,
the myriapods excepted, is divided into head, trunk and abdomen.
Such a division as this does not occur in the crustaceans. In many
the head is not distinct from the thorax, and its place is indicated
solely by the position of the eyes, the attachment of the antennas
and the presence of the mouth ; it is most completely connected
and, as it were, intimately fused with a large part of the body, in
which the principal viscera are contained, and which may be com-
pared not with the thorax alone of insects, but also with the an-
terior portion of their abdomen. To this part of the body another
succeeds, in which only the posterior portion of the intestinal canal
and of the nervous system is contained, and which is commonly
called the tail; thus it is for instance in the cray-fishes. In
other instances the hindmost portion is less obviously separated
from the trunk, and the entire body is parted into rings or segments.
In the Xiphisoura the divisions or segments are not recognisable on
the dorsal surface, for the body is parted into two shields alone, to
which a long and pointed appendage is attached posteriorly. In
others the cq>halothorax is more or less distinct from the rest of
the body, which is not obviously divided into segments but is
covered by a bivalve, membranaceo-homy shell [Cypria), In the
Cirrtpedia the partition into segments ceases ; the body ends with
a thin tail without appendage. They are surrounded by a mantle,
in which, as in the class of the mollusca, calcareous plates are formed
which resemble the shells of these animals.
To the anterior portion of the cephalothorax, or to the head
itself, whenever it is distinct, the antennsB, eyes and oral organs are
attached. When there are four antennae, as is the case with most,
they are placed either in the same plane, or the one pair is placed
above the other, so that according to their position they may be
distinguished as middle and external, or as upper and lower. The
external or lateral antennae are implanted close to the eyes, some-
times beneath the eyes. They consist commonly of three or four
larger and thicker joints at the base, and a filiform part, terminating
finely and composed of numerous joints. Sometimes the antennae
terminate in two or three filaments of this kind. In some the
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600 CLASS X.
antennaB are short, in others very long ; the external are commonly
the longest, and in the genus Paltnunis exceed twice the length of
the body. In some genera there are only two antennas present;
the genus Lirmdua has no antennae at all.
There are commonly two compound eyes, or two groups of
simple eyes. The simultaneous presence of two compound eyes
with simple eyes, so common in the winged insects, is observed in
only very few crustaceans. In most of these there are only two
compound eyes, which are often fixed upon a pedicle and moveable.
The mouth of crustaceans is situated on ^e inferior sur£Etoe of
the anterior portion of the body. The oral organs or jaws are
arranged in pairs and move sideways, as in manducating insects. In
some crustaceans, however, these parts acquire a modified form, and
compose a sucker for taking up fluids, on which these animals live.
The description of the jaws may well detain us some moments, and
requires a more particular survey according to the diflerent orders.
In the ten-footed crustaceans (lobsters and crabs) a transverse
upper lip is present ; beneath it lies a pair of upper jaws, which are
very hard, cut off straight at the extremity, and at the upper margin
provided with a short feeler of two or three joints. [Since the
upper jaws {mandtbuloe) in insects are without feelers, it has been
thought that these joints do not represent a, palpus, but a continua-
tion of the pedicle of the jaws : at all events this palpus differs from
that which occurs on the accessory under-jaws of crustaceans.]
A thin, membranous, bifid tongue, is situated at the base of the
upper jaws*. To it there succeed a first pair of under-jaws, which
are membranous, divided into lobes and beset with hairs at the
margin. Next follows a second pair of under-jaws, which are also
membranous and haired, and may be compared with the imder-lip
of insects which has been split into two parts. Then follow three
pairs of jaws, or rather of feet changed into jaws, to be compared
with the six feet of insects. At the outside of these accessory jaws
a palpus is attached, which, as external division of the foot, does
not correspond with the palpi of insects, and was named ^/2a^mim by
Savigny. This jlagrum consists of a flat, elongated part at the
basisj and a many-jointed filament running to a point at the end.
^ CuYiKB callB this part tiadhne mdchoir. Leg, d^Anat, camp, in. p. 304 ; Fabbi-
cins gave it the name of Id^um.
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CRUSTACEANS. 601
The innennost part of the first pair of modified feet or accessory
jaws is still fleshy and lobed, like the jaws that precede them ; but
the auxiliary jaws of the second and especially of the third pair,
indicate by their form that they correspond to feet, and bear small
gills at their base. In the short-tailed ten-footed crustaceans the
third pair has two very broad joints (the two joints of the femur^
according to Sayignt), so that it covers the oral organs on its under
surface.
The auxiliary jaws of the two last pairs, which, as we have
said, indicate by their form even in the decapod crustaceans most
manifestly their true nature of feet, remain in many crustaceans, as
in Oammarus and SquiUa^ unchanged feet. Hence these crustaceans
have fourteen unchanged feet, and not ten like lobsters and crabs.
The genus Apvs amongst the Eniomostroca possesses those oral
parts alone, which also occur in hexapod insects. To these succeed
numerous feet, of which the first pair terminates in filaments con-
sisting of many joints, but contributes nothing to mastication. In
Limulua there are six pairs of feet at the cephalothoraxy without
any jaws ; the broad basal pieces, armed with spines, of the ten
last feet surround the mouth and perform the office of jaws, whilst
the first pair of feet, scarcely a third of the length of the other feet,
is placed in firont of the mouth. It forms two false jaws that may
be compared with the mandtbulce of the ArachTwidea, whilst the
C4X)ca is membranous, and unites with that of the opposite side to
form a kind of upper-lip ^ If we compare the second pair of feet
with the so-named under-jaws of Arachnids , we shall observe the
greatest agreement between lAmulus and them. Behind the last
pair of feet also in Limulua there are even found two small
appendages, which may be compared with the pectinated organs of
the scorpions.
The body of crustaceans in the more restricted sense or the
trunk, the anterior portion of the abdomen, is almost always
divided beneath by transverse indentations into sections, but its
upper part in the decapod crustaceans is covered by a continuous
shield, named the shell [teata^ in French carapdce^. In the short
^ J. Van DIB HoKVBN, Beekerchea mm* Vffist, not, ei VAnat, da LimuUi, Leide,
1838, folio, p. II.
* In the shell DiaMABBflT has distinguished certain parts by particular terms, from
the position, relatiYe size and determinate form of which, the situation, magnitude
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CLASS X.
tailed crustaceans the pectoral shield is much deyeloped and coY^ts
the body beneath as the shell does above. The first sternal portion
is yery large and is situated in the mid line of the body. To it
succeed four smaller sternal pieces, which are divided in the middle
more or less obviously since they consist of the union of two lateral
pieces. To the first piece the first pair of feet is attached, that of
the so-called shears or claws ; to the four following the four follow-
ing pairs are united. To the basal piece of the feet the lateral
sternal pieces {episiemalia) are attached, which lie on the outside of
the middle pieces and fill up the truncated angles between them. The
most posterior portion of the abdomen in the decapod crustaceans is
always obviously divided into rings or segments, which are seven
in number, or may be fewer from the fusion of some of the rings.
The feet attached to these are short, commonly divided into two
filaments ; the hindmost ring bears no feet. These last abdominal
rings are usually called the tail. In the crabs they are veiy feebly
developed, and the fiat tail is curved round and lies with its
extremity turned forwards in a groove in the middle of the pectoral
shield.
The intestinal canal of crustaceans is short and straight, and
thus agrees with the nature of their food, which is animal. The
anus is situated at the extremity of the abdomen or of the tail. In
Limuliis, however, the long styliform tail is not perforate, and the
antM is situated on the inferior surface of the second shield, in
firont of the base of the tail. So is it also with the Cirripedia^
where the jointless tube, improperly named proboscis by some,
represents the tail, and where the intestinal canal opens at the base
of this tubular prolongation. In some lower crustaceans the intesti-
nal canal is nearly of the same width throughout its whole length,
or has in the middle or more forward an expanded part, which may
be considered as an imperfectly defined stomach^. In others an
obviously distinct stomach is present, which is commonly armed
with homy spines or with calcareous teeth. In Limulus the nar-
row oesophagus with longitudinal folds goes straight forwards and
and form of the internal organs that Ue beneath may be detennmed. See A.
Bboontabt and A. G. Dbbmabsst, But, not. dm OrfOt, fbaUn, pp. 73~-79> "^<1
Dbbmabbbt Cons, gin, «. I. Onut, pp. lo — 13.
^ This is the case, for inrtanoe, in AseUnt {OrUtcut aquoHeut L.), see Tbivibanub
Verm. 8ekr, i. ■. 73, Tab. xi. fig. 64, &c.
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CRUSTACEANS. 603
then cTuryes into the stomach, which is placed abnost perpendicu-
larly. The stomach is compressed laterally, has thick muscular
walls, and is armed internally with many oblong rows of homy
spines. The pylorus projects conically into the intestine, which
then proceeds horizontally backwards, lying in the first part of its
course close upon the oesophagus that runs in the opposite direction,
and lies below it^. In SquiUa the stomach is small, muscular,
triangular, armed in its posterior or pyloric portion with homy,
pointed spines. In the ten-footed crustaceans the oesophagus mounts
almost directly upwards. The stomach is capacious, and in its first
part entirely membranous. The uppermost and hindmost or pyloric
portion of the stomach is supported by hard parts, and therefore,
even in the empty state, remains expanded. To these hard parts
teeth are attached internally, by the assistance of which the food is
comminuted. The epithelium of the stomach, moreover, is covered
with numerous prolongations or colourless hairs, invisible to the
naked eye, whose points are directed backwards*. To this stomach
different muscles are attached, which arise on the inside of the
cephahikorax^ and which can also work upon the teeth described
above. These muscles are, beyond doubt, subject to the will of
the animal, and consequently we have here the rare example of an
organ of vegetative life that is moved by muscles of animal life.
In some ten-footed crustaceans one or more blind appendages to
the intestinal canal are observed, which probably are secreting
organs^. There are some species in which two such blind tubes
terminate in the intestine close to the inferior opening of the stomach,
whilst a single third tube is attached to the intestine lower down.
It is, however, this last unpaired tube which alone occurs in most.
Unless this tube be regarded as an organ for the secretion of
urine, no parts are known which correspond to the vasa urinaria of
1 Beeherchei mr VHid. not, H VAnat, de» Uinvles, p. 17, PL n. fig. i 0, figs. 1—4.
* At the hindmost part of the stomach, behind and under the aboYe-mentioned
teeth, these hairs may be distinguished even with the naked eje. Hie stomach of the
cray-fish has been often described and figured. We content ourselves with referring to
the latest investigations alone, those of F. OlSTXBLSN, in Muellbb'b Archiv, 1840,
8. 387— 44i» Taf. am.
> See Dttvebnot in the second edition of CuvniB Xep. d^Anat. eomp, v. pp. laS,
S39. In as far as they open dose to the pylorus these blind appendages may perhaps
be compared with the paneretu ; but ordinarily they are inserted further backwards.
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601 €XA» X.
in^ecU. In OmUcms and PcfnJbea, iMSOOidnig to TRETiSAsrs, tam
short rewels, irhi'rh open into the lectom dose to its extzemitj,
may perhaps correspond to the nrinaiy Tessels, or, as he supposes,
to the W/iwy, althoi^rh they diflfer from them gieady by their
shortness*. As little are salivaiy organs known hithoto, except
in the Cirripedia. The iirer, on' the contruy, is commonly much
developed here. In some lower crostsceans the intestinal canal is
sorrounded by a layer of small blind sacs (JoUiadi) <xr glandnles,
which may be regarded as a liver intimately connected with the
intestine. In the Cirripedia the intestine below the stomach is
surrounded by a liver formed of many blind sacs. In the On£9cide»
from two to six long blind liver-tabes are found, in most of the
genera four, often with dilations like a string of beads, which are
described by some writers as the adipose body of these animals.
In Bopyrus the intestinal canal, according to Bathke, receives
seven liver-tabes on each side, which lie behind each other in the
length, an arrangement which recalls that in the scorpions, whilst
besides an unpaired liver-mass, incised into three parts, lies in front
of the others on the stomach*. In Limtdus there are two veiy wide
gall-ducts on each side, at some distance behind the^^2t^ntf / they
receive the blind convoluted tubes, of which the large liver-mass of
this animal consists. In the decapod crustaceans only one gall-
duct is found on each side, terminating in the intestinal canal, be-
hind the lower orifice of the stomach. The liver is a doable and
symmetrical organ, as in most crustaceans, and each liver is divided
more or less distinctly into three lobes ; in each of these lobes runs
a tube, that terminates in the common gall-duct, and round about
the tubes blind sacs (JbUicuU) are set, which unite as fingers do.
These follicles consist of three membranes, of which the internal
and external present no special structure ; the external is more con-
sistent and more intimately connected with the middle membrane.
^ Verm. 8ch/r. i. s. 58, Taf. Yil. fig. 38, i. g.
' In SquiUa the liver oomiits of lateral blind aace divided into branches that
extend throughout the whole inteetinal oanal, a diepoeition which agrees with th»t in
BopyruMf and may also be compared with that in AphrodiUa (see above, p. 11 1).
Above the liver, on the dorMl surface, lie the taiet or ffoaria, which also extend longi-
tudinally, and oonslit of branched glandular lobes. The ovofria of SquiUa were de-
scribed by CuTIIB as liver. See Duvbbkot Ann, dea Se, ruU,, 2e Sdrie, Tom. vi.
1836. pp. «47— «5'.
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CRUSTACEANS. 605
The middle membrane is formed of a layer of nucleated cells and
granular tissue, and of cells with fat-globules*.
The blood-circulation, which was already more developed in
the highest arachnids, is here seen to be more and more perfected.
In all the heart is situated on the dorsal surface, and is arterial.
In the OniscideSy the Stomapods, and Inmulus the heart has the
form of an elongated dorsal vessel, from which on both sides
branches arise, that are distributed to the different paits. In the
Lophyropoda the heart is more oval, and situated in the anterior
part of the body on the dorsal surface. In the ten-footed crus-
taceans (crays and crabs) the heart is roundish or hexagonal, much
broader than the main trunks of the arteries, and is situated on the
back between the third and fourth pairs of feet. The arteries in
the crustaceans are very differently disposed ; in some there seem
to be only a few main trunks present, and the blood to flow else-
where merely in the interspaces of the organs in determinate direc-
tions, without being inclosed in vascular walls. In the ten-footed
crustaceans, where these vessels are best known, an artery arises
from the foremost part of the heart, in the middle, which is speci-
ally destined for the eyes; and, besides some smaller branches
which it gives off, divides close to the eyes into two branches.
Next to this artery, there is one on each side which is distributed to
the antennae and neighbouring parts. More behind there arise,
about the middle of the heart, on its under side, two arteries, one
on each side, which run to the liver ; lastly, at the posterior ex-
tremity of the heart, there arises a considerable single artery,
which appears to fill the office of a posterior aorta, and gives off
branches to the intestinal canal, to the generative organs, to the
muscles of the abdomen, &c. Audouin and Milne Edwards
name this vessel sternal artery {arth's stemale). It presently di-
vides into two main branches, of which one runs along the dorsal,
the other the abdominal surface.
The veins in crustaceans, even in the most perfect order of the
Decapods, are represented by cells or spaces between the organs of
the body, in which the blood is moved without proper walls.
1 See T. F. G. Schlbmm De ffepate et Bile Orustaceorum et MoUuscorum quo-
ruTidam, Berolinii 1844, 4to. pp. 13 — t6 ; H. Meckel in Mueller's ArcJUv, 1846,
8. 35—38.
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606 CLASS X.
These conyej it into siniises situated in the middle of the body
or along the feet, and from which it flows to the gills. From the
gills it returns to a sinus that surrounds the heart, and often in the
ten-footed crustaceans is described as ei pericardium. On the dorsal
surface of the heart are fissures, which during the diastole of the
organ afford an entrance to the arterial blood with which this sinus is
distended ^. The blood of crustaceans is whitish or purple, sometimes
red, as in Aptis. The velocity of the circulation is very different
in different families of this class. Cahus, in the cray-fish, observed
51 beats in a minute^, whilst in Daphnia 200 beats of the heart
in a minute were remarked by Jukine and Straus Durgkheim^
The respiratory organs in the crustaceans consist of gills, which
however are not met with in all, so that in some the skin itself
appears to be the only organ of respiration. Also it is probable
in a few that, at the same time that distinct gills are present, the
skin is still partly serviceable for respiration, as in the lateral parts
of the shield-like shell, in which the blood forms numerous currents,
in Apua (according to Zaddach) and in Argultia, in which last genus
VoGT regards these parts as the sole seat of the respiratory func-
tion. The gills have the form either of plates that, consisting of two
membranes, are properly flat sacs, or of filaments. In the Cirri-
pedia both forms are met with ; in AruUijh (and the other pediculated
genera, Lepadiced) there are two or more pairs of soft conical fila-
ments which, with the point turned upwards, are situated at the
base of the pedicle of the drri; whilst in the non-pediculated genera,
the sea-acorns (Balanidea), firinged plates at the inside of the mantle
are present. In many other genera, especially in the Isopoda, the
gills appear under the simple form of certain pairs of flattened sacs;
at the abdomen the feet are composed of two plates, of which the
^ Compare on the circulation of the cruBtaceans amongst others the ohservations
of AuDOUiN and Milwb Edwabds Ann, des Sc. not, xi. 1817, pp. «83— 314 and pp.
35»— 393 ; Milne Edwards Hid, not des CrvM. i. pp. 94—105 ; Luhd in Okkn's
Im 1829, 8. 1999, A. D. Kbohk {Atlacut jluvialUu) ibid., 1834, b. 518 — 519, T!af. XIL
Beautiful figures of the heart and vessels in Attaeus marinua, after the preparations of
HuvTBB, are to be found in the Cat4dog. of the Physiolog, Series of comp. anat, in the
Museum of ike BoyaL Coll, of Swgwns, Vol. 11. 1834. PL xv — xviii. pp. 136 — 140.
* G. G. Cabcb Von den amseren LAenthedingungen der weiss- und halAUUiffen
Thiere, 1824, 4to. s. 83. The number of beats was increased by the stimulus of wanntb,
but became fewer and irregular under the influence of galvanism.
' Mim. du Mushim v. pp. 412, 413.
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CRUSTACEANS. 607
external, of firmer stmctnre and commonly beset with hairs at the
margin, protects as a gill-cover the innermost soft and sacciform
plate, the proper gill. The normal number of gills is five pairs, but
in the land Oniscides and AaeJius there are only three pairs deve*
loped. In AaeUus and many marine OniscideSy the gill-covers
themselves contribute to respiration \ In most crustaceans indeed
these two chief forms of plates or filaments become modified through
greater development, and each gill consists not of a single plate or
of a single thread, but of a large number of plates or threads. Thus
in LimtduSy on the upper surface of the five last abdominal feet,
which have assumed the shape of flat semicircular discs, there are
found five pairs of gills, each of a hundred plates or more, whilst
the first pair of abdominal feet not bearing any gills at their base,
but the external sexual organs, covers all the succeeding feet on the
under-surface, after the manner of a gill-cover. In Squilla there are
five pairs of gills, in the form of numerous filaments placed pecti-
nately on a pedicle, which are attached to the base of the fin-like
posterior feet. In the ten-footed short-tailed crustaceans there are
commonly seven gills on each side, of which that in the middle is
the longest. They have a pyramidal form and are divided longi-
tudinally by a middle septum firom the base to the apex ; on this
septum numerous plates are set at right angles which make up the
pyramidal body of the gills, and consist of folds of the double
membrane of which the septum is composed; these plates thus
form sacs which may be inflated through the septum. In some long-
tailed decapods also the gills are leaf-shaped but in most they con-
sist of a multitude of fine cylindrical filaments which are attached
instead of plates to each side of the axis of the gill. The gills are
more numerous, sometimes even twenty-one on each side. In all the
decapod crustaceans the gills are situated imder the lateral parts of
the shell [carapace) in a proper cavity on each side, and are attached
to the basal piece of the five pairs of feet, or at the same time to the
hindmost foot-jaws also. The water penetrates to the respiratory
cavity by an opening on each side at the inferior margin ; in the
short-tailed this opening is situated in front of the basal piece of
1 TBBvniAznTS Verm. Sekr, i. s. 60—62, Tab. ix. figs. 50—52, b. 73—75, Tab.
zn. figs. 63 — 65. Compare also Dutxskot et Lbbebot7LLKT Eami cTtme MonographU
da organea de la retpiration de Vordre de$ OnutaeU uopodet, Ann. des Sc. not, le S^rie,
Tom. XV. Zoolog. pp. 177 — 240, PI. vi.
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608 CLASS X.
the first pair of feet ; in the long-tailed it is a fissure that extends
along the entire thorax on each side. The water is expelled, on
the other hand, along a fnrrow which opens forwards \>j the side of
the mouth. In this canal an oval lamina is situated, which is the
outermost lobe of the second pair of maxillsB, and produces by its
motion a current forwards to force the water out*.
In nearly all crustaceans the two sexes are distinct. In the
Ctrripedia, indeed, formerly counted amongst the Molluscs, the
male and female sexual organs are united in the same individual*:
[except in certain genera of the Lepadicea^ as Ibla and ScalpeHum
Leach, where there are not only, according to Darwin, males and
females distinct, but also the surprising fact of 'hermaphrodites
whose masculine efficiency is aided by one or more complemental
males',' often exceedingly minute, and rudimentary in structure,
which are permanently attached to difierent parts of the female.]
The ovary lies as an apparently granular mass in the pedicle oiAna-
tija, and under the microscope is seen to be composed of rounded
lobes ; a tube which traverses the pedicle and opens into the mantle
above by a fine aperture on the dorsal surface is the oviduct. In
Balanvs the ovary lies in the walls of the mantle. [According to
Darwin the female organs consist of true ovaria (salivary glands
Cuv.) seated on each side near the base of the labrum, of unbranched
ovarian ducts and of ovarian branching tubes and coeca. In the
Lepadicea the ovarian tubes branch out in all directions within the
peduncle, and ova are developed in the footstalks of their branches
as well as at their ends. Two unbranched tubes enter the body of
the Cirripede firom the peduncle and run into the two true ovaria
situated at the base of the labrum and resting on the upper edge of
^ See MiLNi Edwards Recherehes 9ur le nnScaniime de la respiration eha lea Oru»-
tad*, Ann, de» Sc. not. le S^rie, Tom. xi. Zoologie, pp. 119 — 143.
' A few jears ago Goodsib described Bmall cnurtaceanB, wiih five pairs of feet, as
the males of Balanus. Edinb. new PkHoe, Joum, xxy. 1843, p. 88, Ann, dea, Sc. not,
3ibme S^r. I. 1844, pp. 107 — 1x7. These supposed male individuals present however
little or no resemblance to the juvenile form of Girripedee, as we have been taught to
know them by Bcbicsibtbb.
> Dabwin LepadidcB, 185 1, p. iSa. Dabwin found the animals described by
Goodsib not to be males but females distended with ova. He considers them to be the
females of the unnamed genus belonging to the family of [oniens, described by Qoojy-
BIB, which live parasitically within the sac of the Balani. See Dabwin Mowogr. of
the Oirripedia, BalanidcK. London, 1853, p. 171.
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CRUSTACEANS. 609
the stomach. These tubular glands are generally diyided ^ the
end near the mouth into a few blunt branches. Their precise
connexion with the two main ducts has not been discovered. The
state of these two masses yaried much — ^but since, at times, they
contained aggregations of matter consisting of little balls closely
resembling, in general appearance and size, the ovigerms with their
germinal yesicles and spots, there could be no doubt that they are
ovaria. Darwin was unable to discover the orifice by which St.
Anqe supposes the ova to enter the sac. His observations lead
him to believe that the mode by which they enter the sac is quite
different: ** Immediately before one of the periods of exuviation,
the ova burst from the ovarian tubes in the peduncle and round the
sac, and, carried along the clrculatoiy channels, are collected beneath
the chitine-tunic of the sac, in the corium, at this period remarkably
spongy and full of cavities. The corium then forms, or rather, (as
Darwin believes,) resolves itself into a delicate membrane envelop-
ing each ovum and uniting them together in lamellss ; the corium,
having thus far retreated, then forms under the lamellae the chitine-
tunic of the sac, and the last-formed one is immediately moulted
with the other integuments of the body. The membranes harden,
the lamell® of ova become detached from the bottom of the sac,
and are attached to the ovigerous frsena.'' In the Balanotdea the
branching and inosculating ovarian cceca form a layer covering in
part the basis of the balanus, and in certain cases extending upwards
between the two layers of corium roimd the walls of the shelU.]
The testes lie on the sides of the body as small blind sacs that are
attached to branches that coalesce to form larger stems like veins.
On each side there arises from the union of these branches a wide
and tortuous canal {vaa deferem), which afterwards becoming nar-
rower advances towards that of the opposite side; thus these two
tubes lie close together at the base of the caudiform appendage of the
body, where they unite to form a ductus ejaculatoriiLSy which opens at
the apex of the appendage '• In some lower crustaceans the females
are much more numerous than the males, which are only imperfectly
1 Dabwin LepadicUe, pp. 56 — 60, and Monogr, of the Balamda, London, 1854,
p. 100.
* Waonee in MuELLiB*B Arekiv, 1834, s. 467 — 473, Tab. vin. figs. 8 — 11, 13;
Mabtin St. AiraB M4m. $ur Vorganitalion det Cirripidet, Parifl, 1835, 4to. pp.
11, PI. n. figs. 4—7.
VOL. I. 39
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610 CLASS X.
or noi at all known. In certain of them the males occur cmlj at
particular times of the jear, and the females, like the leaf-lioe, are
firuitful without copulation through several successire generations,
as in Daphnia aad, according to Jolt, in Artemia. In others the
males maj, on account of their minuteness, have eluded observation,
as in the parasitic genus Bopyrus, where they are surpassed in size
bj the female about four times. Besides the unequal size, the
full-grown female in this and in other parasitic genera (Siphonos-
tomes) often present a very deyiating form. The sexual difference
in the common form has not jet in the higher crustaceans been
sufficiently investigated in the different &milies. In the different
species of Limulus that difference is known pretty accurately. In
the long-tailed ten-footed crustaceans the males have larger daws
than the females, the last a broader abdomen than the first.
As a rule the internal and external sexual organs are double.
In Argulua there is only one ovary with one oviduct opening be-
tween the two hindmost natatory feet, whilst however the external
organs in the male are double ^ The ovaries are ordinarily two
blind sacs, as for instance in the Isopods, elongate blind tubes.
Only seldom are they divided into branches, as in ChandracantJua
according to Bathke, in Ajma^ and Limvlua. They pass into two
oviducts, which open each separately. In lAmvlus the two vulvae are
near the first pair of abdominal feet, close to the basis at the dorsal
surface. In the ten-footed crustaceans these two external openings
are situated on the cephalothorax, between the feet of the third
pair, or on the base of these feet themselves. Only seldom, as in
Bopyrua^ do the two oviducts coalesce to form a single tube that
opens into a single mdva. Sometimes there are yet accessory
female organs, for the secretion of a covering of viscid fluid, either
to cover the eggs, or to take up the yperma^ as the two large bursa
copulatrices in the short-tailed decapods'. The testes also have often
^ Here at the fint joint of the fourth pair of swimming-feet is seen a tuberob with
a hook directed backward. JuBim, Ann, dM Mui4um, vii. pp. 448, 449» ^^ ^^' ^
«, 3i flg. «i.
' Zaddaoh, De Apodis caneriformit Anatome et JEtidoria woluUonit, Boniue, 184 1,
4topp. 51, 5a, Tab. I, fig. 14.
• See Cabus Tabul, anat, comp, ilhutr, Faac. v. Tab. 3, fig. 7, /; oomp. MoHi
Edwards ffitt. not. dea Orutt. i. pp. 171 — 174 ; according to the obeervationa of tiiii
naturalist there is no room for doubt respecting the function of these parts (as po^et
eopulairieei).
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CRUSTACEANS. 611
the form of two blind sacs. Sometimes, as in the decapods, these
hare a more glandular aspect, and consist of an aggregate of many
follicles. In the long-tailed ten-footed crustaceans the two testes
form a single three-lobed body, though there are two vasade/erentta*
These efferent tubes open in these crustaceans, as also in the short-
tailed or crabs, at the base of the feet of the fifth pair. In these
animals the external male organs of copulation have homy accessory
organs attached to the first two abdominal rings which support the
penis in copulation, and which as to their function may be compared
with the little bone that occurs in this part in some mammals.
The eggs of crustaceans after they have been laid often continue
attached to different parts of the parent's body, and there advance
to further development. In the Oniscides, and many other crusta-
ceans, the development takes place in a brooding cavity at the
imder surface of the anterior part of the body, where it is covered
by a different number, commonly by five pairs, of ventral plates
lying on one another like roof-tiles *. In Daphnia the eggs remain
for some time in a cavity beneath the shell on the back; in Cyclops
they are carried about in two bunches by the female at the base of
the abdomen. The same is observed in many parasitic crustaceans.
In others they are attached to the feet by an adhesive fluid hardened
into threads; in the ten-footed crustaceans, to the feet of the abdo-
men or of the so-called tail. Other crustaceans divest themselves
of their eggs without bearing them about on their body, and attach
them to other objects. Thus the female oi Argulus foliaceus affixes
her numerous eggs (100 — 200) by means of a viscid covering to
stones'. Cypris also lays her eggs upon different bodies under
water, often in heaps of a hundred, which she covers with a green
thready matter*.
The egg of crustaceans, whilst still in the ovary, consists of the
yolk, the germ vesicle, and the vitelline membrane. The yolk is
slightly fluid, and consists of shapeless fat-particles and cells, with
only a small quantity of albuminous fluid. The germ-vesicle,
which again includes different small vesicles (germ-spots), disappears
after impregnation, as soon as the egg enters the oviduct. Here, on
the other hand, it receives an external covering, a chorion, formed
1 TBBmAinjB Verm, S^r. i. Tab. iz. figs. 51, 54.
• JuBWE, 1. 1. pp. 45a, 453. • Steaus, Mim. du, Mus. vn. p. 54-
39—2
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612 CLASS X.
from a coagulable fluid, which has been secreted by the oviduct
This covering is thicker than the vitelline membrane, and between
the two only a small space or none at all remains ; in the first case
a watery albuminous fluid occupies it. The production of the germ
is, as in the egg of many other animals, both vertebrate and inver-
tebrate, so also in that of many crustaceans, preceded by a parting
and cleaving of the yolk^ There arise first in the yolk a greater
or lesser number of membranous saccules, follicles which probably
take their origin from the albuminous fluid portion of the yolk,
and enclose a greater or lesser number of yolk-cells. The germ
first appears as a nebulous grey spot, which consists of cells that
have nuclei, and probably arise from modified yolk-cells. At first
the germ is small, afterwards the germinal membrane grows round
the whole of the yolk. It separates into two layers, which may
be compared to the serous lamina and the mtiams lamina in the
blastoderma of vertebrates. Just as in insects and arachnids, the
ventral parts of the walls of the body are the first that come into
being. The parting of the body into rings or segments begins on
the ventral surface. All the appendages (limbs, jaws, feet) greatly
resemble one another at first, and on the whole the anterior mem-
bers are formed first, the posterior last. Many crustaceans make
their appearance from the egg with fewer limbs than they afterwards
possess. But the development of crustaceans presents many dif-
ferences, in the different orders, of which the particular description
would demand too large a space. Such an uniformity of the plan
of development, as we observe in the classes of vertebrate animals,
seems in the lower classes of animals not to occur^.
^ EbdIi did not perceiye the deaving in the eggs of AttaeuB fnarintu, bat did in
those of Cancer numat. In this kst Rathkb also observed the phenomenon, as well
as in Oammanu JluviaHlU and O. Loctuta, Kobllikeb in ErgatUut and Oydops,
* As the oomparative history of development in general, so especially has that of
crustaceans, received its clearest iUustration from the miwearied and distanguished
investigations of H. Rathkb. We cite here his UrUenuchunffen Uber die Bildwig »■
JSrUwtcJxlung des Fluahrebset, mit 5 Eupfert. Leipzig. 1829, folio (from which ample
extracts are given with many figures in the Ann, dea Sc. natur, Tom. xz. 1830,
pp. 443 — 469); Ahhandlungen zur BUdufigt-und Entwickdungagesch, IL 1833, s. 69—
94, (in AuHlua ciqucUicut, Onitcits murarivs, 2>ajpA»ia, Lyncetu), and especially for the
numerous investigations in very different iamilies of crustaceans, his work, Zw Mor-
phologie, Rcisebemerhungen aus Taurien. Riga u. Leipzig, 1837, 4to. s. 35 — 15 '• ^
also the compressed review of this subject by Bathkb, in the second edition of Bub-
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CRUSTACEANS. 613
In yeij many crustaceans extremely remarkable metamorphoses
have be^ observed, which, however, are confined to the first period
of life ; long before they are full-grown, they have attained their
permanent form, and after that only undergo repeated moultings.
The yoimg of Cyclops, already observed by Leeuwenhoeck and
afterwards by De Geer, are so unlike in form to the parent animal
that O. F. Mueller formed distinct genera of them, Nauplius and
Amymone; flat, oval, without tail, with four or six feet very thick,
and radiating firom the body^ Young individuals of Ergasilus,
Achtheres, Lemceocera, present similar forms with four or six feet,
according to the observations of Nordmann^. Also the Ctrripedia
present themselves at first under such a form as this, with which,
however, that of the young of Aptta agrees still nearer, that has no
abdominal feet, and possesses two pairs of feet, of which the second
extends behind the body. [But the larves of cirripeds pass through
three different stages. In the first the larva, whilst yet in the sac
of the parent, is nearly globular, with lateral horns. Posteriorly, on
the sternal surface, are two other minute horns pointing backwards.
These horns are cases in which the antennae are being developed.
There are three pairs of feet close together in a longitudinal direc-
tion, some way apart in a transverse. The first pair has always a
single ramus or filament, not articulate in some genera, multi-
articulate in others. The abdomen, a little beyond the end of the
carapace, terminates in a slightly upturned tail. The mouth is
undergoing development on a prominence between the bases of the
feet. Towards the second stage, the most conspicuous character is
the great elongation of the posterior point of the carapace, whilst the
abdomen is prolonged, and has two very long spear-like processes
on its outside (Bcdanus Goodsir), or terminates in a single tapering
spinose projection {Lepas Thompson). In the second stage, the
body is prolonged behind the three pairs of limbs and ends in a
blunt point, in which, probably, the three posterior pairs of feet,
and the abdomen of the larva in the last stage, are developed. The
daoh's Phytidogit, ii. s. 448 — 376, and his CommentcUio de AnimeUium Chnutaceorvm
generatume, Begiomonti, 1844, 4to.
1 De Gkeb Mimoirtapour VHiat, des Ins, vii. pp. 489 — 491, PI. 30, figs. 6, 7, 8;
JUBINB ffist, dea Monocl.; Bahdohb BeUr, zur NcOurgeach. einiger deuiichm Monocu-
ItLiorten, s. 5, 6, &c.
* Mtkrogr. BeUrcige, u. Tab. u. fig, 7, Tab. iv. fig. 7, &c.
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614 CLASS X.
mouth is more forward, there are two simple eyes, and the smaller
pair of anteimae has disappeared. In the third stage, the. larva is
much compressed, nearly of the shape of a Gtfpris, and the thorax
and limbs hidden and enclosed by the carapace elongated back-
wards. The part of the head bearing the antennae is longer and
larger than the rest of the body. The antennae are large and con-
spicuous, consisting of three segments, of which the second (a
sucking disc) is much the largest, the third very small. The
antennae serve for walking, but their principal use is to attach the
larva, the attachment being at first voluntary, but soon becoming
permanent. There are now two large compound eyes close behind
the base of the antennae. The mouth, as in mature cirripeds, is
situated on a slight prominence in front of the thoracic limbs. It is
within the carapace, and still rudimentary. The thorax consists of
six segments with six pairs of feet, each with a pedicle bearing two
arms of two joints. The abdomen is small, but with three seg-
ments, of which the second is the longest, and bears two small
appendages between which the anus is situated. The bivalve shell
and compound eyes of the larva are first moulted: the antenna not
at all. The young cirriped is closely packed within the larva, and
there are two rudimentary eyes posterior to the cast-off eyes of the
larva. They are situated beneath the integument on the upper
part of the stomach*.] Scarcely less strange are the changes which
most of the decapod crustaceans undergo. The early states of short-
tailed crustaceans have been frequently recorded in systematic
works as distinct genera. Thus the genus Zoe Bosc, with large
eyes and a long bent beak and a recurved hook on the back, is
founded merely on individuals in the first period of life of Ccmcer
and Eji/as*. These animals have then a long tail, which only at
a later period is bent under towards the breast. But many long-
tailed crustaceans also imdergo changes of form, ex. gr. Pagurus.
In Astacua marimis the feet are at first provided with a jointed fila-
ment, which corresponds to the flagrum of the auxiliary jaws. Of
all the decapod crustaceans hitherto investigated ^«tocu« Jlufnatilts
1 Dabwin, Lepadidce, pp. 8 — 15.
* See Rathke BaUebemerhmffen aus Seandinavien, Neuetie Schriften der \
forwkenden OudUeh. zu Danzig, ni. 4te8 Heft, 1842, Tab. iv. The discovery of the
aingular change of forms in Decapods was first made by JoHH Thomfbon. Rathkk,
relying on his own inyestigations in AitaeuM Jluviatili$, at first doubted^ nftyi ^^^
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CBUSTACEAN8. 615
presents the smallest changes of form. The Amphipods, on the
other hand, come from the egg in a form similar to that of the
perfect animal. Many Isopods^ also, undergo only slight changes of
form ; many acquire another pair of feet in addition to those which
they had on leaving the egg. The yomig Limuliy according to
Milne Edwards, leave the egg without that ensiform appendage
or pointed tail which so strikingly distinguishes the full-grown
animal.
The crustaceans cast their shell several times. In yoimger indi-
viduals these moultings succeed each other at shorter periods, but
in full-grown animals, at least in the decapods, the hard calcareous
shell is cast off only once a year. In the River-cray the moulting
occurs towards the end of summer. The hard shell begins to
loosen itself from the body, which in the meantime is replaced
by a new covering situated beneath the former. The animal makes
many movements and contortions, until at last a fissure occurs on
the back between the abdomen and the large cephalothorax, from
which the animal with the fore part of its body and its feet makes
its appearance. At last the hinder part of the body divests itself
of its old covering. For these observations we are indebted to
Beaumub. In the short>tailed crabs the shell splits on each side
along the epimera. The cast-off shell presents penectly the form of
the living animaP.
As long as the new shell is still thin and flexible, the crays and
crabs are very sensitive. They then conceal themselves in holes,
imtil the new shell has attained sufficient hardness, for which a few
days only are requisite.
With the shell the inner coat or the epithelium of the stomach
is renewed in the crays and crabs. When a new internal tunic of
the stomach has been formed the old one is cast off and dissolved in
the cavity of the stomach. It has been supposed that the two
round calcareous plates which are situated on each side of the
contested the obeervations of Thokpson, but afterwards admitted, with that upright-
ness which always belongs to the inquirer after truth alone, his mistake respecting this
important discoyery, and that he had done the English naturalist a wrong ; Op. cit.
8. 46. See also the ob6er\'ations of Du Cans on the metamorphoses of Cancer mcauu,
Annals cf not. Hiti, ni. pp. 438—440, PL xi.
1 See B^UMUB Mim, de VAcad. des Sc, 17 18, p. 263 and foil., Ck>LLiN80ir Phil,
Trana. 1746 and 1751, MiLNX Edwaxixs Bist, nat, da OruH, i. pp. 53 — 57; oomp.
also Btmxb J0KI8 jiniffuU Kingdom, ad edit. 1855, pp. 434—436.
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616 CLASS X.
Btomach, and are named crab's-ejes {oeuli s. leqndes cancramm),
supplied the matter from which the new shell acquired its hardness.
These little stony substances are cast off with the old tunic of the
stomach, and being freed from their capsules come into the caTity
of the stomach, where they are broken up and partly diiBSolyed.
Thus it is possible that the calcareous matter, taken up into the
blood, may be useftil for the secretion of the hard shell (V. Baer).
The part, however, which the crab's-eyes take in the secretion
cannot be great, when we compare their weight with that of the
calcareous matter in the shell. During the time that the shell is
still increasing in hardness, no new crab's-eyes are produced ; but
only after the shell has attained its greatest hardness is calcareous
matter again secreted on the walls of the stomach, and new craVs-
eyes again appear. Thus the production of crab's-eyes would appear
to be a vicarious secretion ; a secretion of such constituents of the
blood as, if top abundant, would be injurious to the organism, like
the secretion of urine for instance, but with this difference, that the
calcareous matter is not set at liberty shortly after its secretion, but
remains accumulated for a long time in continuance ^
The power of restoration or reproduction is very great in this
class, so that even feet, amputated or broken off, are replaced bj
new ones.
We will now speak concisely concerning the organs of animal
life. The nervous system consists, as in articulate animals generally,
of a cerebral ganglion above or in front of the oesophagus and of a
ventral cord, which is formed of a greater or lesser number of ganglia
connected together by two filaments. Originally every nervous
ganglion consists here, just as we remarked formerly on the ringed
worms, of two lateral portions. In some crustaceans this separation
of the nervous ganglia persists even in the adult state, as in Cyamus,
Talitrus and Idotea; the lateral parts are merely connected by a
transverse commissure. In the Oniscidea the two nervous filaments
of the cord lie quite apart from each other, and the ganglia still
indicate evidently, by their compressed broad form, their original
^ Already in the first edition of this Handbook I offered eBsentially the same
opinion respecting the use of the secretion of crab*B-eyee, I. bl. 410. Gomp. on this
subject y . Babb U^ier die sogenannte Emeuerung de» Magma der Krdm il s. w. in
MuBLLEB*B Archvv, 1834, s. 510 — 533, and Dulk'b chemical investigations, ibid.
8- 5 ^3^5 ^7^ l>ut especially (EsTSBfiEN, Mublleb*8 Arckiv, 1840, #. 43i>-440.
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CRUSTACEANS. 617
condition of pairs. The number of the ganglia is, however, very
different, and varies from more than sixty in aptis to two in the
short-tailed ten-footed crustaceans. In cancer mcenas a cephalic
ganglion is situated above or in front of the oesophagus ; from it arise
the nerves of the eyes and of the antennae and neighbouring parts.
From the posterior part of this ganglion proceed the two nervous
filaments, which form the oesophageal ring, send in the middle
a nerve to the upper-jaws, and afterwards unite in the middle of
the thorax in a ganglion that is excavated in the centre. This
annular ganglion is almost eight times larger than the cephalic
ganglion in front of the oesophagus ; from its circumference nerves
arise that rxm to the feet and other parts, and, in addition, a single
nerve from the back part, that runs in the middle throughout the
tail and takes the place of the continuation of the ventral cord, but
has no ganglia \ In this crab, consequently, the ganglia of the
thorax and of the anterior part of the abdomen are represented by
a single ganglion, which, however, by the excavation in the middle
affords an indication of the lateral divisions. In the genus Majaf on
the other hand, the union has attained the highest degree, and the
large thoracic ganglion has lost the excavation in the middle.
Here then there are only two ganglia present, one for the head and
one for the thorax and rest of the body*. In the long-tailed ten-
footed crustaceans, on the contrary, there are different distinct
nervous ganglia in the ventral cord, twelve, for example, in the
1 CuyiSB Xep. cFAnat, amp. n. pp. 316, 317.
* See AUDOUIN and MiLNS Edwabds Reeherchea anat. awr le Sytt, nerveux dei
Cru9tacSs, Ann. dea Se. not. xiv. i8«8, pp. 76—102, PI. 1—6.
A fdUer descriptioii of the nenrouB system in different families of the Crustacea
would extend beyond the limits of this Handbook. I will here only add, that in
Linmlua the cephalic and thoracic ganglia coalesce to form a thick elongated nervous
ring, which behind the oesophagus is connected by three transverse commissures, and
from which backwards a nervous cord arises composed of two closely connected nervous
bundles, which in the ventral shield presents inconspicuous gangliform thickenings, and
afterwards divides into two lateral portions ; each of these strings then terminates in
an elongated semilunar ganglion, at the inferior extremity of which two nerves arise
that penetrate the caudiform appendage. On the part of the cephalothoracic ring lying
in front- of the oesophagus there are two conical swellings forwards, frt>m which the
two very long optic nerves arise for the two compound eyes situated above on the
dorsal shield. These nerves go first forwards and upwards, then bend outwards, and
afterwards go backwards to end at the inside of the eyes. Their length is more thax^
one quarter of that of the whole animal, the tail included.
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618 CLASS X.
lobster. In the hermit-crab (Poffurus) there are, besides the cephalic
ganglion^ only six ganglia, but the form of the nerrous system is
in other respects the same^*
The neryons system existing especially for the organic life,
which we have already noticed in insects and spiders, is not want-
ing in the crustaceans also, however it still in many families
remains for the most part or entirely unknown. It has been
especially investigated by Brandt in some Decapodsy Stomogpoda
and Oniscidea. In crays and in SquiUa it consists of a single
portion and two lateral portions. The siugle portion arises at the
posterior margin of the cerebral ganglion, and has no nervous
ganglion that lies in front of the brain as in insects ; the lateral
portions arise from the oesophageal band that connects the cerebral
ganglion with the succeeding ganglionic cord. Distinct ganglia are
not present at the origin of the lateral portions, but appear to
be fiised with the oesophageal band, which indicates a swelling at
the point of origin of these lateral nerves. The system of nerves
spreads itself over the stomach, presents here one or two ganglia,
and finally divides into two branches which axe distributed to the
liver. In PorceUio Brandt found no single portion of this system,
but only two small lateral swellings, situated behind the cerebral
ganglion and connected to it by two ihin filaments, which send fine
branches backwards to the stomach'.
In most crustaceans, common feeling on account of the hardness
of the covering cannot be otherwise than very small. The antennas,
which are often much developed and four in number, are wanting
only in few, and, like the whiskers in mammals, may serve for a fine
sense of touch even in the dark. In like manner, probably, the
jointed filaments of the cirrtpediay and the filaments on the foot-
jaws of the decapods and other appendages of the body, may serve
for touch in many species. A special organ for taste is not known,
but the seat of this sense appears to be in the commencement of the
oesophagus •.
^ SWAMMXBDAV Sijhd der Naiwur. bl. 20^—206, Tab. xi. fig. ix. ; oomp. abo
OwxK, CateUogue of the pKytiol, Seriet of the Miueum qf the OoU, of Surgeotu, it.
pp. t6, 17, who in other species of the same genus found only four gangUa beddes the
cerebral ganglion.
' Gomp. Bkandt in his Bemerhunffen Ober die EinffeweideHerven der EveriebraieH,
dted above (p. 479).
* According to MiLKX Edwards, who appeals to his obserrations made with
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CBUSTACEAN3. 619
In the riyer- and searcraTs Rosenthal first described «Dd
figured an organ as that of smell, which afterwards was also fonnd
in some other deoapoda^ hot is wanting in most species of this order,
as well as in the remaining crustaceans. At the base of the middle
or innermost antennas he found a triangular opening beset with
hairs, which is the entrance of a triangular cayitj lined with a soft
membrane, in which nervous branches are distributed^.
Just as little is known of the organ of hearing in most animals
of this class. In the ten-footed crustaceans it was discovered by
J. C. Fabrictus, and Minasi and Scarpa, and others, after him,
described and figured it in the river-cray (common cray-fish). At
the base of the outermost antennsB is a very hard wart-like ex*
crescence, at whose point is a round opening covered by a tense and
veiy elastic membrane. Behind this membrane is a vesicle filled
with fiuid, on the walls of which a nerve is distributed that arises
with the nerve of the external antennae from the cerebral ganglion**
The compound eyes in crustaceans are formed on the same plan
as in insects. Blainville found in them the vitreous humour, as
did JoH. Mueller after him in the eyes of insects ; and his descrip-
tion of the eyes of PaMnurus agrees with that which we have given
above of the compoimd eyes of insects*. In many crustaceans, for
instance in the crays, the facettes of the cornea are not hexangular,
as in insects, but quadrangular ; in some other ten-footed crustaceans,
AuDOunr, bat which haye not been made farther known, this sense woold seem to be
mnch developed in cmstaoeans. HUt. not des Onut, I. pp. iii, X13.
^ See BosnvTHAL in Rbil's Arehivf, d, Phytiol, x. 181 1, s. 433, 436, figs. 1—4.
Ck>mp. Trkvibanus BiologU, vi. s. 308, 309. This part is by Fabri considered to be
the auditory organ, PhUoi, TranBoeL 1843, p. 333 ; comp. however hereon Ekiohbon's
report in his Arehiv f, Naturg, 1844, s. 336, 337, who participates as little in that
opinion as I, for my part, am able. [Additional investigations by Leuokabt, Archiw
f, Naturgeaeh, 1853, i* b. 455, strongly oorroborate Fabbx'b conclunons.]
' A. Scarpa, AnoUomica dUquiskionei de Auditu H O^factu, 'Hcini, 1789, folio,
pp. 7, 3, Tab. IT. V. ; oomp. also £. H. Weber, de Av/re H Auditu hominit et aninudium,
Pan I. Lipei», 1820, 4to. pp. 8, 9, Tab. i. figs, t, 1. The membraneous tube situated
in the pedicle belongs to a larger sac placed behind it, Brandt Mfdk, ZoU. n. s. 64,
Tab. XT. fig. 13, a, a. In the short-tailed decapods this membrane covering the entrance
to the auditoiy sac is represented as a moveable calcareous plate. See on this Uttle
plate and its peculiar arrangement in Maia, Milne Edwards ffisL not, des Crust.
I. p. 194 ; on the auditory organs of Cfrustaeea, see T. H. Huxlet ZoUog. noUs and
oUervaiUms in Ann, of nai, Hisl. sec. S4r. VoL vn. 185 1, pp. 304 — 306, PL xrv.
* DuoROTAT Dl Blainvillb, De VorganiseOwn des Animaux, i. 1833, 8vo. pp.
433* 434.
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620 CLASS X.
in the Stomapods, &c., they are, howeyer, hexangular. In Limulus
thej are also hexangular, not very regular, and at the outside but
little raised. In some crustaceans the eyes are covered with an
undivided smooth cornea^ as in Apua; thej are compound ejes
with a cornea without facettes^. They make as it were the tran-
sition to the clustered eyes, placed in two groups at the side of the
head of the Iscpods, as Oniscus^ Idotea^ &c.
The passive organs of motion of crustaceans are the hard cover-
ings of the body and of the limbs, the dermal skeleton. There are
also frequently productions of this covering penetrating inwards^
homy or hard calcareous projections to which the muscles are
attached. In Limulus an elongated, rectangular tendinous plate,
somewhat excavated on the upper surface, is situated in the inside
of the cephalothorax, just as in spiders, in which, at the fore part,
are two cylindrical processes that pass iuto tendons. This part is
moved by many muscles, and, with its movements, those of the feet
are also connected. The legs of decapods have at every joint a
flexor and extensor muscle; those of the first pair or of the so-called
claws, in accordance with the greater strength of this pair of limbs,
are the most developed. The muscles of the tail in the crays
(lobsters, cray-fish, &c.) are divided into two layers; the layer
which is situated on the ventral surface, that of the flexor muscles,
is more composite and more powerfully developed than the layer
which lies towards the back*.
In art-instincts the crustaceans appear to stand below most
insects and arachnids. In them the vegetative or organic life is
more developed than the animal.
The geographic distribution of crustaceans has not been hitherto
sufficiently investigated, although Latheille, and after him espe-
^ See JOH. MnELLKBinMxoKEL'B Archiv, 1839, s. 54—59, ^'^^^ m. fig. 15; in Gam-
marut pulez, ibid. figs. 16, 17; in Branchipua, H. Bubmbibtkb in Mvellbr*8 Arekiv,
1833, B. 539— 534» »• 613, Tab. xra. figs, 1—4. The conical tranaparent bodies of the
separate divisions of the eye are either attached immediately to the cornea or have
lenses placed before them, which lie under the cornea. See on these pecnliaritieB
Mueller Handimeh der Phytiologie, 11. s. 309. The eyes of Limulus do not however
belong to the division in which Mubllbb places them. The organs of vision in Crus-
taceans are largely treated of in R. Waqneb Lehrb. der vergl. Anal,, 9te Auflage,
nter Theil, bearbeitet von Db H. Fbbt u. Db R. Leuoxabt, 1847, b. loi — 206.
■ The tail-muscles of the river-cray are described and figured by Cutibb, Zepoiu
d^Anai, camp, I. pp. 413—416, v. PL xiv.
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CRUSTACEANS. 621
ciallj Milne Edwards, have made copious contributions to it,
which however are confined principally to the order of the decapods.
With the different peculiarities that have an influence on the distri-
bution of species we are not yet satisfactorily acquainted. The
difference of salt-particles in the different seas comes here beyond
doubt into consideration, and not the mean temperature alone ; the
relative depth also, the nature of the coasts, &c. On the whole,
the general rule, that animal forms, the genera and species, are
more numerous in proportion as we advance from the poles to the
equator, has not such an unlimited prevalence for marine animals.
In size also and in brilliancy of colour the animals of the polar seas
often contest the palm with those of the tropics. Yet the short- tailed
ten-footed crustaceans, so rich in species in the seas of the warm
regions of our globe, are almost entirely wanting in the neighbour-
hood of the poles. On the other hand, the northern regions are
rich in forms of amphipods, which necessitate the adoption of
many distinct genera ; so that the colder seas may be regarded as
the proper home, the true father-land of these cnistcicea^.
^ H. Kbobtkb NatwrhiMoritk TicUakrifi, iv. 1842, pp. 141 — 166. On the geographio
distribntion of Onut<icea compare Latbbillb Mim, du Muaium, iii. 1817, p. 45; or
M^moiret nvr divert sujets de VSist. not. det IntecUt, de G^ographie ancienne, ftc.
PftriB, 18 19, 8yo. p. 174; and especially Mjlnb Edwasds, Ann. da Sc, ntUur. sec
S^e, Tom. x. pp. 119 — 174.
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SYSTEMATIC
ARRANGEMENT OF CRUSTACEA.
CLASS X.
CRUSTACEA-
ARTICULATE animals apterous, supplied with articulate feet
both thoracic and abdominal, breathing mostly bj bianchi», some-
times bj the skin, without stigmata. Heart very often distinct,
aortic, situated in the back.
Sexes in most distinct. Animals mostly aquatic.
Obdeb L PcscihpocUu
Jaws none. Feet of cephalothorax placed round the month,
with C0Z8B very spinose, performing the office of jaws. Abdominal
feet resembling semi-orbicular lamellae, bearing the branchiae. Two
shields, gibbous above, hollowed beneath, homy, hard, one covering
the cephalothorax, the other the abdomen.
Family L Xtphosurcu (Characters of the order.) Twelve
pairs of feet, six attached to the cephalothorax, six to the abdomen.
An appendage elongate, acuminate, carinate or gibbous above,
plane beneath, adhering by articulation to the posterior part of the
abdominal shield, covered by very hard skin.
Limulus MuELL. {Polyphemua Lam.).
This genus contains only few species, and is the only one in this
fiunily. The anterior shield is round forwards, and terminates on
each side behind in a point, which extends along the outside of the
second shield, which is of less breadth, about as fsff as its middle.
This second or abdominal shield has an irregular hexangnlar form,
and is armed on each side with seven sharp teeth and six moveable
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CRUSTACEA. 628
awl-shaped sj^nM between them. The feet of the Cephaloihorcuc
have a aihear-flhaped extremity j in the sixth pair the shear is snm<^.]l
and ahnoBt concealed by four lancet-foimed, homy laminn, that
are jwoTided on the outside with a projecting ridga Below on the
abdominal shield are six pairs of leaf-shaped natatory feet, of which
the first pair is the largest, and almost entirely covers the rest on
the inferior snr&ce ; the five following bear on their dorsal surface
the gills, which consist of many plates.
Above, on the shield of the Gephaiothorax, are situated two
kidney-shaped compound eyes, very remote from each other, and
more forward and near the middle, dose to each other, two very
small simple eyes of an oval form.
Oomp. Bakzaio OsBervaxiom nil Limuh polifemo in Opuscoli icientijici.
Bologna, u. 4to. 1818, pp. 175 — 285, and Van Dxb Hobvbk ReehereheM
tur VHUi, nalhi/r. et VAnatonUe det IAmnle», Avec 7 phnehei. "Lade, 1838,
folio.
a) Abdominal »hidd with Uut margmal iocA dongaie, with a^pex m the
vUddle.
Feet cf the ieeondpair alcne fMyMdaetylom in vujUes,
Sp. Limuliu Polyphemue Latb., Guianr leonogr,, OrutL PL 34, fig. i,
DxBiCABKST CtvmI, PI. 5 1, Yait DEB HoiVBK ReehtTch, PL VL (on the
easteam coart of America, espedaUy of N. America).
b) Ahdominal thidd wUh lad tooth eearcdy longer than the rett, hroad,
with tihairp point marginal,
* Feet of th^ eecond and Mrd pair fMnedaOy^^
Sp. LimvUie moluecanue, Pclyphemm gigae Lav., Buicph Awh. Sariteik,
Tab. xn., Yait dkb Hoevsn L L PL i. The Cancer pervereue or Balancae;
this Bpedes is fonnd on the shores of the idands of Sunda and Molucca ; its
Malay name is Mimie. The LimtUi live constantly in pairs, and are also
sold in pairs. In the months of July and August they are daily taken in
quantities near the roads of Batavia, and brought alive to nuurket. The
Malays eat the eggs with avidity, and the flesh also is agreeable to them
and to the Chinese. These animals can Hve more than a day out of water;
when laid on the back they are not able to right themselves. The tail is
a powerful instrument of defence.
Lim/ului langiepina V. J>. HOEVIN L L Tab. V. ; on the coast of Japan.
** AU the feet in loth aexet didaOgloue.
Sp. Limidut rotundicavda Latb., V. D. Hobvbn L L Tab. iv. figs. 1—3.
East Indies.
Note. — Limvhu vireeeene ItAXti, The species appears to me doubtful. A
single specimen mutilated and imperfect !» prsserved in the Paris Museum ;
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624 CLASS X.
I think it iR only a monstrous yariety of LimuluB molueeanus. Seveal
fossil spedes are known from the lithographic formation ; see my monograph.
Tab. 7.
Obder II. Ichihyophihira.
Mouth suctorial. Bostrum tubular, including two mandibles
setaceous, acuminate, formed of labium and labrum conjoined,
sometimes concealed, or a tubercle instead of haustellum. Anterior
feet supplied with hooks or acetabula for fixing the body. Animal-
cules, in the adult state, adhering parasitically to fishes, often
deformed, soft, with segments obsolete. Females supplied with
oviferous appendages (external ovaries).
Fi8h4ice, We prefer this name to that of ParasUoy introduoed
by WiEGHAinr, because in the class of insects an order of Fcmuidca
had been preyiouslj adopted The animals when young swim
freely about, by means of feet with long hairs^ and res^nble the
young animals of Cydopa.
Comp. on this order Blainyille MSfnoire wr lei Letnies, Jounudde
Phygique, de Chim., (Fffist, not. &c. Tom. 95, Paris, 1823, pp. 372 — 380,
and pp. 437^ — 447, and by the same the Article LemSe in Dictionnaire da
Scieneanat, ZZYi. 1823, pp. ti2 — 130. A. YoK Nobdmank Mikrogm-
phucKe BeiirUge, ates Heft. Berlin, 1832, 4to. H. Bubmbistsb Beackrei-
bunff einiger Schmarotzerkrdfte. Nov, Act, Acad. Ocu. Leop, xvn. 1835, PP-
269—336; H. Kbobtbb Om SnyUthrebicne, Natvrh, Tidukr, i. 1837, pp.
172 andfolL, pp. 252 and foil., pp. 470 and foil., pp. 605 — 628, n. 1838,
pp. 8—62, K>. 131—157.
Family II. Lemoeacea {PeneUina BuRM.) Body in adults not
articulate, cylindrical or sacciform. Articulate feet none.
Lemcea L. {Lemoeocera Blainv., V. NoRDM.). Body ventri-
cose, mostly incurved. Three or four appendages, branched at the
apex or bifid, around the head, for fixing the body.
Sp. Lenuxa branchicdis L., Sncyl mith,. Vert, PL 78, fig. 2, GuiBiK leonogr.,
Zooph, PI. 9, fig. I ; with three branched litUe horns on the head; the
ovisacs are two long tortnous strings ; this species fixes itself to the gills of
the cod-fish; comp. C. N. (Ozbhak) VUgezogU Verhanddinffin, n, 1757,
bL 282 and foil., PL 14, m. bL 232 and foil., PL 23.
There was much donbt formerly respecting the place of these gili-worma
in the natmul arrangement of the animal kingdom. CuvnsB placed them
amongst the intestinal worms (inUstmaux cavitairetf see 12. Ani. le Mit.
m. p. 255), as a distinct and anomalous family. The resemblance to some
other parasites, which had been refetred to the Crustacea, caused indeed a
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CRUSTACEA. 625
Buspicion that the LemcBa also might belong to this class (Debhabest
Com, gin. t . 2a CUuk des Crmt. p. 344), but it was only after the observa-
tions of N0BDMAJ7N on the young form of Lemcsocera eyprinaeta, L. etocina
BuRM. (Mikrogr. Beitr, n. Tab. vi. figs. 5, 6), that the matter oould be
regarded as definitively settled. The young animal has three pairs of feet
and a single eye in the middle at the anterior part of the body.
Pennella Oken^ nob. Head clavate. Two horns elongato-conical
at the base of head. Body straight, cylindrical, elongate, sub-
eqnally thick. Appendages small, disposed in pairs, resembling
fins, lanceolate, at the anterior part of body (rudiments of feet).
The posterior part of body as though pinnated, with several distinct
filaments. External ovaries round, very long.
Sp. PenneUafiota, PermatiUa JUoaa Gu., GuifiiH leonogr,, Zooph. PI.9, fig.
3; habit, in fishes of the Mediterranean.
Add genei'a LemeoneTna Milne Edwabds, and FemciUus Y.
NoBDiCANN, distinguished from PenneUa by defect of the posterior
. pinnated part especially. Crenus Lemeanema perhaps ought not
to be separated from the Lemcece; comp. the forms of Lemcea bran-
chialis which Kroeyer has delineated, i. PI. 3, fig. 10.
Spkyrion Cuv.
Family III. Lemceopoda. Antenna small. Feet uncinate,
two or more behind the rostrum ; natatory feet none.
A. Body affixed by two round arms, grown together from their
base throughout their whole length or conjoined at the apex alone,
terminated by an adhesive disc.
AnchoreUa Cuv. Body produced anteriorly into a cylindrical
part like a neck, rugose transversely. A short peduncle from the
two arms coalesced at the base of the neck, with adhesive disc at
the apex.
Sp. AnchoreUa uncinaUi, Lemcea vncinata Mubll., NoBDMAim 1.1. Tab.viii.
figs. 8 — 12, The characters of the genus apply only to the female, as
is common in the LemaxB, The male, which is much smaller, almost
spherical, has a conical rostmm, and two pairs of thick, short hooked feet
with one krge and two smaller booklets. SeeNoBDHANN 1. 1. Tab. x. fig. i.
[Pentastoma Run. ought to be brought here from the Entozoa,
according to Yan Beneden Mem. de VAcad. roy. de Brtixelles,
1849, and Ann. d. Sc. not. 3i$me S^. Zool. Tom. xi. pp. 313—348.
See also above, p. 189.]
Braokiella Cuv. Two arms extended in front of body, con-
joined at the apex alone, often very long.
VOL. 1. 40
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626 CLASS X.
A. Anterior part of body elongate, contracted, wemMmg a necL
G6iiera : Trachdiastes Nobdm., BrachieUa Nordh.
B. Anterior part of body oval.
1* Abdomen undivided.
Genera : Lemmopoda Blaiky., Bcuanistes Nobdh.
Sp. Biuanitte* ffuchonis Y. Koll^ Ann. des WUner Muae¥mt, i. Tab. i.
p. 86. {Tracheliastet steUifer KoLL. ibid. Tab. ix. figs, i — 8, seems to
belong to genua Lemceopoda.)
1 1 Abdomen divided by segments.
Achiherea Nobdh.
Sp. AchJOmt* perearum Y. Nordkakn L L Tab. TV.
B. Body not affixed hj two arms.
Cfhondracanthus De La Roche, Lemantoma Blainv. Upper
antennsB subulate, bi- or triarculate, inferior uncinate. Mouth in-
ferior, remote from the anterior part of body, bordered by two
hooks, scarcely prominent. Body with various lateral appendages
bifid or ramose. External ovaries two. Conical tubercles in many
in the mid line of the body, both dorsal and ventral.
Sp. Chondraeanthut La Roehei Guv. JR. Ani, PI. xv. fig. 3 (does not differ
perhaps from CJumdrac. gibboiua Kbostkb, i. PL a, fig. 4, pp. 953—257);
Clumdraeanthut Trigla Nordm. 1. 1. PI. 9, figs, i — 4 ; Lemaa aselUna L.
sec. KsosTER, L L n. pp. 135, 136; — ChondraeatUkia Lophii IUtejo,
BeUr. zur Fauna Norwegent. Tab. V. figs. 11 — 18, &c.
Clavella Oken, Kroeyer, {Habitus almost of Pe/itcuZw^NoRDM.)
Lemanthropua Blainv., Epachthes V. Nordm. Superior an-
tennae sexarticulate. Three pairs of feet behind the rostrum.
Cephalothorax separated from abdomen by stricture.
Sp. ZemofiMropiu ptipa Bubm. 1. 1. Tab. xxiv. figs. 7—- 1 1 ; LerMmik. para-
doxuBf EpachtKea paradoxm Kobdm., Burm. ibid. ftg. I4.
NoU. — ^To this family perhaps is to be referred genus Staurownia Will,
not yet 8u£Bciently known, androgynous in the opinion of the author, para-
sitic in AetinicB; see Ebiohbom*8 Archivf. Naturffetch. 1844, pp. 337—343-
Tab. z. figs. 1—9. Add genera Tueea and Seliua Kbokteb, 1. L I. pp. 476,
479. The last genus is referred by the author to the Srgatilina.
Myzostoma Leuckart. (A genus, whose place is uncertain,
perhaps to be counted amongst the parasitic crustaceans.)
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CRUSTACEA. 627
We h»ye already noiioed tbis genua (p. 575). It was regarded by Loyiir,
who has treated of its structure most exactly, as a form of transitioii
between the Trematodes and the ringed worms. I think that the short
jointed feet militate against this disposal of it. The surface beset with
yibratile dlia would seem to remove it from the crustaceans, with which
however it has the most affinity. The characters which Lov^ g^ves to
the genus are: Corpiu moUe, depressum, dUcoideum; proboscis retrtKtilu
mutica; anvs dUcretut art oppoaitut; hothria laleralia oppoaita ; pedes itrferi,
articulati, hamifert; genUalia mcucula duplicia mtUica, There are different
known species of these animalcules, living parasitically on CoTnattUa, only
one or two lines in size, of which one is distinguished by conical cirri, twenty
in number, at the margin of the disc of the body. See Lbdokabt Zoolo-
gische BruckstOclce, iii. 1842, pp. 5—11, Lot^n, Eriohson's Archiv /.
Naturgesch. 1842, s. 306 foU. or Ann. des 8c, not, ie S4ne, xym. Zool,
pp. -291 — 298.
Family IV. ErgasiUna. Cephalothorax large. Abdomen arti-
culate, made up of many segments. Two or four antennae. Mostly
four pairs of abdominal feet (natatory).
In these parasites the similarity of form with the genus Cyclops
is yery conspicuous^ so that even AunouiN and Milne Edwards at
first placed the genus Nioothoe, discovered by them, with the Mono-
culina,
A. Body elongate. Two filiform receptacles of eggs, at the
posterior extremity of body. Uncinate feet behind the rostrum.
Anthosoma Leach. Superior antennae sex-articulate, setaceous;
inferior antennae uncinate. Mouth produced into a rostrum. Ce-
phalothorax oval. Two foliaceous laminae at the back behind the
cephalothorax, and three pairs of laminae under the abdomen, in
place of natatory feet
Sp. Antho8(ma SmttkU JjujOB, CaJUgua crassus Ajbildoaabd, Skrivter of
Naturhistorie SdskaJbd, m. 3, 1794, p. 49, Tab. v. figs. 1—3, Desmabbst
Cons. gin. $. I. CfnuL PL 50, fig. 3 ; in the mouth and on the gills of sharks.
Nemesis Roux.
Fig. Gu^Biir Iconogr,, Crust. PI. 35, fig. 11.
Dtchelesthium Herm. Anterior antennae with seven joints,
setaceous, reflected backwards ; posterior antennae forcipate, resem-
bling two frontal chelae, with internal finger moveable. Cephalo-
thorax cordate, truncated anteriorly. Two mandibles denticidate
at apex, setaceous, between the folds of rostrum, and two palps
40—2
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628 CLASS X.
bifid, furnished at the longer apex with double seta. Five pairs of
feet, the two first uncinate, the third and fourth with apex bifid,
acideate, the fifth oval, simple. Two small oval appendages at the
end of the last segment.
Sp. JHcheUithiun Stwrionit Hebu., CaUgus ohlongut Abildoaabd, 1. L p.
$a, Tab. v. figs. 4— ii, Hermann Mhn. ofUrol, Tab. v. figs. *], 8, Rathkb
Nov. Ad, Acad. Ccu. Carol. Tom. iix. Part i. 1839, "• ^^5 — ^53> '^^
XYii. ; this elongated parasitic crustacean fixes itself by its two shear-shaped
antennas fast to the branchial arches of the stui^geon.
Lamproglena NoRDM. Mouth a truncated tubercle. Antennse
four subulate ; anterior larger with twelve rings, posterior not arti-
culate. Ocellus single. Two pairs of uncinate feet ; four pairs of
natatory feet, imperfect, very small.
Sp. LamprogUna jnUcheUa Kobdu., L 1. Tab. i. Milne Edwasds Hisi. det
Cfrud. PL 39, fig. 6.
B. Body oval or broad, depressed, posteriorly attenuated. Two
sacciform receptacles of eggs at the base of abdomen. Mouth a
truncated tubercle.
Nicotho^ AuD. and Edw. Cephalothorax in adults divided by
a stricture into an anterior cephalic portion, rotundate, small, and a
posterior portion very broad; sides of cephalothorax produced into
two wings lobed, inflected backwards. Ocelli two. Antennas two,
setaceous, small, ten-jointed. Two pairs of uncinate feet. Four
pairs of natatory feet.
Sp. Nicothoe AHaci AvJiOVis and Edwards, Ann. des Sc. not. dl 1826, pp.
345 — 359> ^^ 49> figB. I — 9 ; this little animal, which, with its two lateral
lobes and the two large ovisacs situated behind them, reminds us at first
sight of a four-winged insect, a little moth for instance, lives on the gills of
the lobster, and undergoes no such changes as most of the parasites and
even the monoeuli, but has merely a difiTerent form of the oephalothoraXy
and two fewer abdominal rings and feet. The male is not jiet known, as in
other genera of this family. Ck>mp. also Kbokteb 1. 1. ii. p. 146, Tab. ni.
^. 7, and especially Bathke Nov, Ad. Acad. Leof. Oar. xx. 1843, PP>
103 — To6,Tab.Y. figs, i — to, and Van Bbitsdbn ikf^m. twr U develcppemaU
et VorganiaaHon des Nicothoit. Mim. de VAcad. roy. de Belgtyue, Tom.
XXIV. av. une pi.
Bomolochua NoRDM. Antennae two, with basal joint large,
thick, denticulate above, the rest attenuate, setaceous. Feet bifid,
setiferous in four pairs.
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CRUSTACEA. 629
Sp. BomoloehuB parvuUu Y. Nobdil 1. 1. p. j^s;^Bamol. Bdonei BuBU.
L 1. Tab. XXIV. figs, i — 6.
Ergaaihia NoBDM. Antennas four ; anterior setaceous, sexarti-
culate, posterior quadriarticnlate, large, resembling arms, with last
joint incurved, acuminate. Mouth inferior, resembling a conical
tubercle in the middle nearly of cephalothorax. Ocellus single,
frontal. Four natatory feet divided into two oars. Abdomen
attenuated, terminated by double setiferous appendage.
Sp. ErgagUuM Sieholdu KOBDU. 1. 1. Tab. n. ; Erganlut gihbui NoBDM. ib.
Tab. m. figs. 1—6, &o. To this genus also belongs, as Kbobyeb justly
remarked, the LemoBa Lcwcureti described by Otto Fabbioiub, Skrivter of
naturh. SeWedbet, in. i, 1794, p. «i, Tab. m. figs, i — 5.
Family V. Caltgina. Body suboval, covered with a test
above. Cephalothoracic shield large, broader than the rest of
body, with anterior margin on each side mostly running out into a
transverse process or frontal lamina. Anterior antennse with two or
more, rarely with three joints, broad, short; posterior antennaB biar-
ticulate, uncinate, placed in front of the tubular mouth. Feet of
the second pair uncinate. Ovaries external, filiform. Animals
mostly not affixed, living parasitically upon the body of marine
fishes.
These crastaceaDS appear to me, more than the other families of
this order, to approach the Xiphosv/ra, a group that otherwise stands
much apart
A. Lamellose appendages on the back.
* Beceptacles of eggs contortuplicate, covered by shield.
LoBmargus Kroeyer. Anterior antennas triarticulate. Frontal
laminss none. Feet of the fourth, fifth and sixth pairs branchial,
with oars large, lamelliform and basal joints minute.
Sp, LasmarguB murieaiut Kboeteb 1. 1. I. pp. 487—501, Tab. v. figs. A— E,
Milne Edwabds Hist, d, Oruti. PL 39, fig. s; found on OrihagorUew
mola,
Cecraps Leach. Frontal margin of cephalothoracic shield ex-
cised in the middle, produced at the sides into a rounded lobe.
Anterior antennae Particulate. Feet of fourth, fifth and sixth pair
with basal joints large, lamellose, branchial and oars short, narrow.
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630 'CLASS X.
Sp. Ceeropi LatreUUi Dbbkasest Oiue. PI. 50, fig. i, Guiaiv Jeomogr.,
Onut. PI. 35, fig. 8, Cuv. jB. Ani, 6d. HI., OruH,, PI. 78, fig. 4; Uvoion
the Tunny, according to Kobdmank on the gills of OrthagorvKm aioIcl
* * Keoeptacles of eggs straight, ezsert, cylindiicaL
PhyUophora Edw.
Pandarvs Leach. Several dorsal shields. Two frontal lamina.
Sp. Pcmdartu Careharia Lkach, Bubmsistsr L L Tab. 15 ;— Paniorst
dentaiui MiLirs Edwards Bist. d. Crutt, PL 58, fig. 19, ftc.
Dmemura Latr, {Dinematura BuBM. in part). Two dorsal
shields. Two frontal laminae. Two pairs of palps. Rostrum
acuminate, produced. Feet of first pair terminated by two aculeate
cirri, of second pair by a strong incurved hook. Feet of third and
remaining pairs bifid; basal joints of last pair minute, with large
branchial oars.
Sp. Dinematuraferox Kbobter L 1. 11. pp. 40^45, Tab. i,fig. 5 ; — i>ifMiiMirt
aUUa, Panda/nu dUOtu Milkb Edwards J imi. dts Sc. neU, Tome 38, 18331
PI. 8.
The type of this genns was for Latrbillb the Calicut produetut of 0.
F. MnsLLKR (Entomottr, PI. 21, figs. 3, 4, copied bkEncycl mHk., OnataCt
Arachn. et Ins, PL a68, figs, i, a). On this species, known only imper-
fectly, compare Kroetsr L L pp. 45 — 47.
B. LamellsB incumbent on back none. Ocelli two (approximate
situated almost in the middle of cephalothorax, yet more towards
the anterior margin).
Caligu8 Muell. Cephalothorax running out anteriorly into
two transverse laminae, produced posteriorly by its outer margin
beyond the segment annexed to it, almost resembling a horse-shoe.
Furrows in the dorsal surface of cephalothorax forming the figure
H. Oviferous filaments exsert.
Add genera : Ewryphorua Nobdil, Nogagus Leach {Din&maiwra
BuBK. in part), Trebitia Kboeteb.
Sp. CdligvM htppoglossi Xroetkr, Binoculut pitcinus O. Tabbu, Kroxtsr I !•
Tab. Ti. fig. 3; on the holybut; this crustacean has been also figured (but
hind-foremost) by Bastbr Natuurk. Uitap, n. Tab. Vin. fig. rx. h, R—
Caligtu curtut Muell., Monoadus pitcinus L. in part, Cali^us M0eri
Leach, Desmar. Ortut. PL 50, fig. 4, on different species of Oadut.
NoU, — DvMmatura snracilisBvRU. is, in the opinion of the leamedKBOlTlB
the animal in an earlier state, not yet perfect. He thinks the same of
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CRUSTACEA. 631
genxm ChdHmui BuBK. contoining young CaUgi. His conjecture has been
confirmed by the observationB of Fb. Mueller, vid. Archivf, N<Uwrgt»ch,
1853, B. 91, Tab. IV. figs. 5, 6.
Family VI. Argulina. Shell membranons, scutifonn, placed
npon cephalothorax, extending posteriorly into two wings that
receive the body between them. Compound eyes, two. Antennae
four, concealed under cephalothorax, short; the anterior biarticu-
late, recurved, the posterior quadriarticulate. Rostrum acuminate,
situated in front of aperture of mouth. Six pairs of feet, the feet of
the first pair being changed in adults into two suctorial acetabula;
the feet of the second pair short, with basal joint aculeate, the last
eight feet cloven into two oars or pilose cirri. Tail short, bilobed,
behind the sixth pair of feet.
Argvlua MuELL. (Characters of the family).
Sp. Argulfu fcliaceM Jxtbuxe, MonoctUua foliaeeu$ L. Fauna Suee, 9044,
Hebm. Mim, apUrol, Tab. Yi. fig. 11, KooH in Panzer u. Hesbioh-
ScHJBFFSB Ins. DeuttcJU. Heft 186, Tab. 70, This little parasitic crusta-
cean lives in fresh water, and attaches itself to fishes (especially Chuteroiteui
punffitius) and to tadpoles. The suckers which had originally the form of
fore-feet, are only developed after the fourth moult. Compare on this
species the beautiful memoir of Jurine, the son, in the Ann, du Mu»6um
d'jffid. not, VII. 1806, pp. 431 — 459, PI. 16, from which the figures of
DE8MARB8T {Crutt. PI. 50) are borrowed, and C. Vogt BeUrdge swr Natwr-
getch, dor Schtpeiier-Ontatae. Nene Denhchr. der aUg, Schweizeritchen
CfeselUch. f, d. getammt, Naiiwrwmentch. Bd. vii. 1845, pp. 1—16^ Tab. I.
figs. I — XI,
Order III. Lophyropoda Latr. [Eatomostraca Edwards).
Mouth not produced into a rostrum, not suctorial; mandibles
suitable for manducation, maxillae lamellose. Feet never more
than ten, natatory, not foliaceous, often cloven into two oars,
furnished with set» or cilia. A single eye or two eyes, sessile.
Tufi-footedy or Oa/r-footed Grustacecma, Most of the known
species are from fresh-water, and all are of small size ; their motions
are rapid and frequently interrupted
Comp. on this order:
MuELL. Entom. (Entomotlritca 9eu Insecta tettaeea, qua in Aquit Dania
tt NorveguB repent, detcriptit d iconihuB iUustravit O. F. Mueller, Lipsus
et Haunioe, 1785, 4to.); — Jurike, Hitl. dt» Monocles qui setrouverU avx
eimrons de Qtniite, xSao, 4to. av. fig.
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632 CLA8S X.
Family VII. Copepoda Latr., Edw, {Cydopidm Bukai.).
Body oral, attenuated posteriorly, terminated by two appendages,
as if by a forked tail. Maxillas and foot-jaws various in num-
ber, lamellose, setigerous. Antennae two or (in many) four.
Natatory feet mostly eight, cloven into two oars.
This family is veiy closely allied to Caligtu, ErgctsUus, and some
other genera of the preceding order, from which it is distinguished
by the absence of the suctorial beak, and especially by the longer
antennm.
A. Eye single, frontal.
Cydopa Mueller [Monoculus L. in part).
Add sub-genera : Cydopaina, Arpactictu Edw. and I^oiodelphys
Allmann.
These Uttle aninudi »re to be met with almost eveiy where in water, even
in puddles, gnttera, and dsteniR of rain-water. The Cifcloptina tdpeBtriM of
G. YooT was found at a height of 8500 feet in water of the glacier of the
Aar ; Neue Denktchr. der allg. Sehtoeizer-CfeBdlsch, /. d, getamnU. Natur-
vntaench. vn. pp. 17 — 19.
The young (oomp. above, p. 611) have no tail, and a smaller number of
feet. The adult female differs from the male in having two oval sacs whidi
she bears about with her, and which are Med with eggs ; some species,
instead of two, have only a single sac attached to the middle of the body
beneath.
Sp. Cyclopt vulgaris Lbach, MtmoenUu quadricamu L., R(BSSL Im. m.
Suppl. Tab. 98, MuELL. EtUom. Tab. 18, Desmar. OrutL PI. 53, figs. 1—4;
very common in almost all fipesh water, so that of a certamty everyone has
often swallowed this little animal. They may be frozen in the water, and
after continuing motionless for days, be restored when the water is thawed,
after some hours, from their torpor to their former activity. They are
ordinarily thickly beset with Vorli^lot, The oopulaiion and impregnation
have been illustrated by Siebold's observations ^
B. Eyes two.
Qenera : FotUia Edw., Cetoehilua Roussel be Yauzemb, fferaUia
Philippi, Peltidium Philippi, Saphirina Thompson, Cananu9
Kboeter (not Leach).
Comp. Milne Edwabds Ann, det Sc. not. Tom. 13, pp. 295—297, PL
14;— R0UB8BL DS Vauz^e DeBcrtjOum de Cetochilut aut^ralu, Ann, da
1 BtUrHgt zur Nalurgetch, der vnrbellotm Thiere. Danzig, 1839, 4to. pp. 3^— 5©-
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CRUSTACEA. 633
Sc, not. 26 Q6ne, Tom. i. ZooloffU, pp. 333—338, PL 9, Philipfi in WiBO<
MANir*8 u. Erichson's Archiv, 1839, 1840, 1843, where Bome other genera
are proposed which are here omitted. Comp. Lubbock de novit genenhuB
et specieb, CcUanidarum, Ann. of not, ffitt. sec. Series, Vol. xi. 1853, pp.
*5 — *9> PP- ^o* — ^og, XII, pp. 115 — ia4.
Family VIII. Ostracoda [Cypridtna Edw.). Body com-
pressed, with segments indistinct, included in a bivalve shell, with
dorsal hinge. Anterior antennae setaceous; posterior geniculate,
pediform. Two pairs of maxillss; the second pair or foot-jaws with
a pectinate or flabelliform appendage at the base. Feet undivided,
acuminate towards the extremity, slender, four or six.
The two large jointed appendages, especially useful in swimming,
which we have here regarded as posterior antennse, are by many
writers, even by Erichson, considered to be the first pair of feet,
(placed in front of the mouth). On this supposition these little
animals have not two or three, but three or four pairs of feet.
A. Eye single.
Cyprts MuELL. (Species of Monoculus L.). Only two pairs of
feet besides the posterior pediform antennse ; feet of the last pair
recurved upwards, concealed beneath the shell. Abdomen termi-
nated by a bifurcated tail.
Comp. H. E. Stbaus Mimovre 9ur lea Cyp) it, Mim, du Mm, d^Higt, not,
vn. 1 8a I, pp. 33 — 61, PL I. and S. Fisoheh Abhandlung Uber das Otnut
Cyprii. M4m, det savantt Hrangeri der KaUed. AJcad, der WmeMch. Tom.
vn. St Petersbui^, 1851.
Sp. CyprUfuica Straus (Monoculua eonchaeetu L.?), Stbaus 1. l.i. figs, i —
16, GufiRHf I&mogr,, Orutt, PL 31, fig. 4, Dkbmab. CruH, PL 55, fig. i,
ftc. Tliese a^^p^^^ln live in fresh water.
Cythere MuELL., Oytherina Lam. Three pairs of feet besides
the posterior pediform antennae.
B. Eyes two.
Cypridtna Edw. [and Asterope Philippt].
Order IV. Cirripedia.
Crustaceans affixed in the adult period, included in a midtivalve
shell or coriaceous covering furnished with calcareous points, as
though rudiments of shell. Eyes in adults none. Six pairs of
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634 CLASS X.
feet, with short fleshy peduncle, and two cirri, mtdtiarticnlate,
homy. Mouth with mandibles and maxillae membranoso-homy.
Tail round, acuminate, reflected between the feet towards the ven-
tral surface of the animal.
Comp. PoLi Teatacea utriu$que SicUia, I. pp. i r — 39 ; CuviSB MSwuMra
iur lea Animaux det Anatifet et des Balanea {Lepaa L.) et aur lew anatamie;
Mim, du Mua. iFBiM. not, n. 1815;— 6. J. Mabtdt St. Ajtgb MHuirt
aur VOrganiaaition dea Cirripldea. Avee t planckea. Tarn, 1835, 4to. ; — ^H.
BuBMEiSTEB BeUrdffezur Oeachichle der Ranket^f&aaer. Mit 2 Kufferl. Berlin,
1834, 4to. — Catal, comp. AneU,, eoU, of Swrgeona, I. PL iv. pp. 255 — 160,
AnAtomy of PentaXaanUa vitrea and BaUmua tintinnabulum. See also the
article Cirrhopoda by J. Goldstrbam in ToDD*8 Oyelopcadia, i. 1836, pp.
683 — 694 : and especially DASwnr Monograph of ike Cirripedia {Lepadida),
London, 1851, and Monograph of ike Cirripedia {BalaaUdeB, VerrmeidtEf
kc,), London, 1854.
With LiKNiEUS these animals formed only a single genus (X^nu).
Lamasck was the first who made of them a distinct class, to which, on
account of the filiform arms, he gave the name of (Xrrhipedes {cvrrin
pedea^). Most writers place them amongst the molluscs, although
the resemblance to articulate animals was apparent to many, and
GirviEB shewed himself not averse to the opinion that they ought
perhaps to be arranged amongst these. The history of their develop-
ment, however, illustrated by J. V, Thompson* and Bcbmeisteb, can
leave no reasonable doubt that the cirripedes belong to the articulate
animals, and amongst them do not form a distinct class, but only an
order of the crustaceans. The place alone, which we allot them in
the series of the crustaceans, may perhaps admit of some doubt, but
we think that it ought to be preferred to a position at the end of
the crustaceans, which determines nothing respecting their true
affinity. According to us the Cirripedia have the same relation to
the Daphnidea and PhyUopoda as the Lemceaoea to the Cop^xxia.
Although the shells differ much in different species, and some of
these animals are pedunculated, others not, yet the cirripeds have
such an agreement in internal and external structure, that we have
eveiy reason to admire the sagacity of Lini7.£US who united them
all in a single genus.
The body of these animals is in the adult state inarticulate,
although on the dorsal sur&ce, between the different pairs of feet.
1 Philoaophie toologique, Paris, 18091 I. pp. 37 4, 315.
> Zoological Jleaearchea, Cork (1830), and Philoa, Tranaact, for 1835, pp. 355— 358#
PI. VI.
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CRUSTACEA. 635
a separation into segments is indicated. It tenninates backwards in
a thin tail, which earlier writers improperly called the proboscis*
Along the yentral surface there are six pairs of arms, each of which
bears on a short pedicle two long filaments turned inwards, that
consist of numerous joints and are beset with hairs. The ftniwiiyls
extend these filaments constantly during life, from the opening of
the shell and draw them into it again, by which they introduce
within the shell the water for respiration and with it their nutri-
ment
The nervous system consists of a row of six ganglia, or properly
pairs of ganglia, connected by two nervous cords. The first ganglion
is situated on the short oesophagus ; on each side a nervous string
descends — a ring being thus formed round the oesophagus — ^to the
second ganglion, which is situated between the first pair of feet. The
intestinal canal is straight and very short ; it runs along the dorsal
surface and terminates at the base of the tail Through this tail
runs the common efferent tube of the two wua de/erentia and opens
at its point. These animals are bisexual, with some exceptions (see
p. 608).
[The prehensile antennie of the larvae of Cirripeds in the last form
contain the ducts of the cement glands, which may be traced firom
within the discs of the antennae to the anterior or lower ends of the
incipient ovaria. The gland which secretes the cement appears to
be a part of an ovarian tube specially modified. If the base of at-
tachment of a Cirriped be carefully removed, the larval prehensile
antennae, from which the cement always escapes, will be found very
near to its extremity. In sessile Cirripeds a new cement gland is
formed at every period of growth. Dabwin Lepadidm, 28, 33 *.]
Cirripeds are found in the seas of every i*egion of the world ; they
attach themselves to rocks, to many marine animals as shell-fish,
turtles, whales, to different polyparies, to marine plants, to the hulls
of ships, to the floating wreck of vessels that have been lost, as frag-
ments of wood, bottles, &c.
^ Our countryman SlabbeBi as I lately diflcovered, was the first who observed and
figured the larvse of AnaHfa, without however recog^sing their true nature. See
Naiu^wh, VerluM, Tab. vm. fig. i, which almost entirely agrees with the figure of
Thompson PHUob, Trans. 1 1. fig. 5. Perhaps also PI. vi. fig. i. of Slabbxb is a tran-
sitory form of another species.
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636 CLASS X.
Family IX. Balancndea, Shell sessile, truncato-conical or
tubular, calcareons, open at the apex; calcareous valves at the
aperture. Animal conical, sometimes depressed. (Grenus BcUanus
Brug.).
Seor-AcomSj Sea-TiUips. The pieces of shell, that form the calca-
reous tube, shut upon each other with teetL In some the base is
closed by a calcareous plate, in which hollow tubes run fix>m the
middle to the margin, which communicate with other tubes in the
length of the shell. Poli Testae utriuaque SicUta, i. Tab. iv. figs.
9, 10. [Branching and inosculating ovarian tubes. Dabwin Bal<i^
nidof, pp. 100, 101.]
[To obviate the extreme confusion of the nomenclature, Darwin
proposes the following names for the external parts of Balanoids.
What is visible externally of sessile drripeds is composed of
shell and operculum, the operculum being generally seated a little
within the orifice of the shelL The shell consists of btuis or support
by which it is attached (membranous or shelly), and of compart-
ments (8 — 4) occasionally all calcified together. The compartment
as the end of the shell where the cirri are exserted is called carina,
that opposite to it rostru/m, those on the sides are the three lateral
compartments, that next the carina the carino-lcUeral, that next the
rostrum rostro-lcUeral, and the middle one simply the lateral com-
partmenL These three are rarely present together. Each com-
partment consists of a wall (paries) which always grows downwards
and forms the basal margin, and is furnished on the two sides with
alee or with radii, or with an ala on one side and a radius on the
other. The radii, not always developed, on the upper part over-
lap the alee, which usually extend about half-way down the
compartment. The caHna has always two alas. The carino-
lateral and lateral compartments have always an ala on the rostral
side and a radius on the carinal. The ro8tro4aleral (when present)
always radii on both sidea The rostrum normally has aloe on
both sides, but very often, when fused with the rostro-lateral com-
partments, it has radii on both sides. The i>peTculum consists of a
pair of scuta and a pair of terga, joined to the sheath of the shell by
the opercular membrane. On the internal surface of the scutuan
there is almost always a pit for the attachment of the Adductor scur
toruan muscle, and beneath the adductor ridge, often a pit for the
depressor lateralis muscle.
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CRUSTACEA. 637
A slit-like orifice between the opercular valves leads into the
sac in which the body is lodged. The valves are two on each side,
tergum and scuium. The tergum has three margins, the scutal, hcucU,
and carinaU ; on the basal margin a spur {ealcar) depends. The
sciUum has also three margins, the hasai, tergal, and ocdudent, so
called from opening and shutting against the opposite valve. Dabwin
BcUanid{jB, pp. 3 — 7.]
Coromtla Blainv. Operculum not articulated together, com-
posed of two or four calcareous pieces united by membrane.
Coronula Lam., Diadema Bakzani. Shell depressed, conical or
suborbicular, with very thick waJls excavated internally by radiating
cells.
Sp. Corowida diadema L., Blainv. Malaeol. PI. 86, fig. 4, Cuv. R. Ani. id,
iU., Mollvsq. PI. 139, fig. 2, on the Fin-fisb, Bakmemoptera, Balama
longimana.
TtUneinella Lah. Shell tubular or cylindrical, belted by trans-
verse ribs or rings. Operculum with four valves.
Sp. TubicmeUa hakenarum Lam. Ann, du Mut. z. Tab. 30, fig. i, Blainv.
Malaeol, Tab. 86, fig. 5, Gu^rih Iconogr,, MoUutq, PL 38, fig. 14 ; liyee
paraaitically in the akin of the whale of the soatheni hemiBphere {Balama
MiftUeehu anlarctica),
Balamia Bruo. (exclusive of some species), Lam. Shell conical,
composed of six laminss or valves. Operculum articulated, subver*
tical, consisting of four valves.
8p. BalanuM tulcatut Lak., Lepaa halaMts L., Bastbb Naiuurk. Uitap. i.
Tab. zii. figs. 7—10 ; Balanut miger Lak., Fncjfd. nUih. Ven, PI. 64,
fig. 4, Ac.
A casta Leach.
Creuna Leach.
Pyrgoma Savigny.
Note. — On this and some other genera, proposed by Ranzani^ and
others, compare Milnb Edwards in the new edition of Lamabck's Hid,
not, dee Anim, t. VerUbree, v. 1838, pp. 669—672.
* Observationi eu % Balani; Opu9coli scietUifice di Bologna i. 1817, pp. 195 — ^02,
pp. 269—176, n. 1818, pp. 63—93.
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638 CLA88 X.
Family X. Lepadicea (Blainy.) s. Anatiferce. Animal sus-
pended by a pedmicle cylindrical, flexible. Shell mostly com-
pressed or involucram coriaceous. (Genus Anattfo Britg.).
[These Cirripeds consist of a ctipUtUum, much flattened, and a
pedicle of vaiions lengtL The capitulum is generally formed of
five or more yalves, connected together by narrow or broad stripes
of membrane : sometimes the yalves are rudimental or absent, when
the whole consiBts of membrane. Of the valves the seuia are the
most persistent, then the terga, then the carina, the rostrum and
IcUera occur only in a few (Follicipes), The sctUa and terga are
always considerably larger than the other valves. Within the
capUukum is the sac which encloses the animal's body. The pednn-
de is usually flattened, sometimes quite cylindrical Its oorium is
very thick, and in those genera having numerous valves, scales are
found at its conuexion with the capitulum placed in whorls. The
peduncle is lined by three layers of muscles, running from its bottom
to the base of the capitulimi : and the cement ducts may be traced
upon them on each side until they expand into the two cement
glands, and which are connected with the ovarian tubea The
cement escapes through the larval prehensile antennse, which may
be always found quite close to the end of the peduncle if it be
carefully removed from the surface of attachment. Yid Dabwin
Lepadidw, pp. 28 — 37. See aJso above, pp 608, 609.]
Tetralaamis Cuv., Ibla Gray. Peduncle short, hirsute. Shell
with four valves, two dorsal long, narrow, and two ventral tri-
angular.
Sp. TOraUunUi hirgultu Cuv., Afiat\fa quadrivalvU CuvnsB MSa^ tur Us
AncUifei, fig. 14, GuiaiN Iconogr., MoU. PL 37, fig. 7.
Anai^a Lam. (Species from genus Anatifa Brug.). Peduncle
smooth, resembling intestine, elongate in some. Shell with five
valves; the single dorsal valve narrow, falciform.
Qoos&^nund. Tho four pieoes of shell in pairs are, as in the preceding
genus, to be compared with the opercular plates of the BaUud ; the single
piece of shell along the back of the animal earvna corresponds to the calca-
reous tube, and the stem is as it were an elongation of the basal piece of
the tube of the last-named animals.
Sp. AwjOAfa IcBcis Lau., Lepaa anati/era L., Blajnv. MdlacoL PL 86, fig. 3,
Guj^lN Iconogr, 1. 1. fig. i ; this species is often thrown on our shores,
especially in the winter months. In different districts of the north, a
fable is current that a species of goose {Anat hemicU^ has its origin from
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CRUSTACEA. 639
this cniBtaoean ; from this ridiculous popular fimoy the specific same gfHm-
mussel is derived.
Polltcipes Leach (and Scalpellum ejusd.), Polylepaa Blainv.
Peduncle rough, squamose. Shell compressed at the sides, with
valves subcontinuous, unequal, thirteen or more.
Sp. PoUicipes acalpdlum "Lam., Zep<u walpdluvii L., Gu^bin Iconogr, I. 1.
tg, 4 \—PoUxc. mitdla, Lepaa mUella L., Rumfh. Amb, BarUeitkam, Tab.
47^ ^. u, GuiBiN L 1. fig. 3, Ac.
Gymnolep(Z8 Blainv. Conchoderma Olfers. Mantle naked,
without valves, or with rudiments of valves, remote from each other.
Cvneraa Leach, Lah. Calcareous pieces five distinct, joined by
membrane, small, with one dorsal, two above the aperture of mantle,
two others below that apertura
Sp. Oineroi vUtata Leaob, Ltpas coriacea Poli Testae. I. Tab. Ti. fig. 30,
GuiaiN Iconogr, 1. 1. fig. 5 ; in the Mediterranean sea.
Otion Leach. Two lunated valves at the margin of the aperture
towards the peduncle. Body with two tubular ear^like appendages
pervious at the apex.
Sp. Otion Cktvieriif Lep<u aurUa L., Pou i 1. fig. ai, Gu^bin Iwnagr. 1. 1.
Alepas Rang. Mantle without any calcareous pieces, subpellucid,
continuous with peduncle.
Sp. Alepoi fatcicuUUa Lsssoir, Anatife jaune iant coqutUe Mabtin Sahtt-
Angb M&m, 8wr let Cirripidet, Tftb. I. Gui^lN Iconogr. 1. 1. fig. 8, Alep<u
squalicola Lov^, dfverngt of konigl, vet, AJcad, PMiardlingwr, 1844,
pp. 193, 193, Tab. III. (in this species, that lives parasitically on sharks,
the feet also are soft, and without bristles).
Note, — On the fossil species, which are chiefly found in chalk strata, of
Anatifa and PoUicipet comp. Stebnstbuf in Kbobteb's Tidekrift z. 1837,
pp. 358 — 366, n. 1839, PP' 39^ — ^4^5* *°*^ ^" *^® fossil Lepadida of Great
Britain, C. DABwnr's Monograph,, 1851, printed for the Pabeontographical
Society.
[From the investigations of Darwin, recorded in his two admir-
able memoirs so often cited, we learn that his subclass of Oir-
ripeds contains forms that dififer greatly from those of the Balor
noidea sxALepadicea and are much less perfect in their organisation.
He divides the Cinipeds into three orders — ^the Thoracica, Ahdomi-
ncUia, and Apodes, the limbs or cirri being thoracic in the first,
abdominal in the second, and entirely absent in the third. In the
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640 CLASS X.
Thoracica three families are included — ^the BcUamckB or sessile Qini-
peds, the VemuddcB, (the genus Clysia Leach, Creusia Lam., Ver-
ruca ScHUHACHEB, ScHUK. Essai cTun nouveau Syst^me des Habitor
lions des Vers testac^Sy 1817. CopenhaT.), remarkable for their
quite aBymmetrical shell, and the Lepadidce or pedunculated Cir-
ripeds. In the second order of AbdomincUia the seventh or last
cephalic segment is quite distinct, and bears radimentaiy org&Ds
answering to the first pair of foot-jaws in ordinary crustaceans,
of which organs, and of the segments supporting them, there is
no trace in the Thoracica; the seven succeeding thoracic seg-
ments are destitute of any appendages ; but the three segmentB
of the abdomen bear three pairs of cirri. In the metamorphosiB
all the first changes are merely indicated by changes of fonn
in the egg-like lai-va, without the development of distinct organs,
and the last or pupal condition, which is attained within the sac of
the parent, is very peculiar from the entire absence of rotatory
limbs. There is only one genus Cryptophiahis Dabwin, and one
known species Cryp, mintUvs Dabw., BcdanidcBy pi 566, Tab. 23,
24. The third order Apoda is the most peculiar of all, it resembles
the larva or maggot of a fly, the carapace is reduced to two separate
threads for attachment. The last cephalic, ihe seven thoracic, and
the three abdominal segments are all destitute of appendages. The
single known species, Proteolepas bivincta^ is parasitic within the sac
of Alepas comuta, Dabwin Lepadidof, p. 165^ Tab. 3^ %. 6. See
Dabwin Balanidce, pp. 20 — 22.]
Order V. Cladocera Latr.
Body compressed, included in a homy shell, divided into two
parts by a dorsal fold, without hinge. Head free, produced below,
terminating in a species of rostrum. Two small antennae at the
apex and two larger at the base of head, split into two or three
branches. Feet foliaceous, with four or five pairs. Abdomen ter-
minated posteriorly by two setaceous appendages.
Family XI. Daphnidea Straus. The characters of the order
are those of the single family. Single compound eye. Larger
antennae ramose, serving like oars for swimming. Ova in females
situated on tlie back in a space between the shell and the segments
of the thorax.
Polyphemus MuELL., Gephaloculvs Lam. Eye single, large,
occupying almost the whole head. Larger antennae bifid. Feet
eight, exsert from shell. Tail reflected, bisetose at the apex.
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CRUSTACEA. 641
Sp. Polyphemut ocuku Mubll., {Polyphemw pedicuku L. ? ) Ds GssR M^m.
M. I. Int. VII. PL 28, figs, g, 10, Muill. Eniom. Tab. xx. figs. 1—5,
Dbsmab. Onat. PI. 54, figs, i, 2, KooH, Hbbbioh-Sohjsfvsb DeuUchl.
Int. Heft 187, Tab. 2.
Evadne Loven.
Compare S. L. LoviN Ev<idne Nordmanni, tin biaher unbekanrUes Enio-
moUracon, in WiBauANK's Archiv f. Naiwrge$ch, 1838, pp. 143 — 166.
Daphnia MuELL. (and Lynceus ejusd.). Feet ten, included in
shell, the eight anterior foliaceous, ciliated. Larger antennas bifid,
with branches having from two to four joints.
Sp. Daphnia ptHex Latb., Monoculua pulex L., Swammbbdam Bibl, not.
Tab. 31, figs. I — lU, MuELL. Entomostr, Tab. xii. figs. 4 — 7, Dbsmabbbt
Cnut. PI. 54, figs. 3 — 5 (borrowed firom the figures of Stbaus) ; the
branched water-flea. This species is reddish, especially in spring, and fix>m
its abundance may give a red colour to the water, or cause an apparent
change of it into blood, of which Swammbbdam adduces a remarkable
example, bL 89, 90.
In the genus Lynceus Mdbll. there is a black spot in front of the eye ;
the antennae are usually very short. To these belongs Lynceus sphoericus,
&c. Ebwabds counts some of the species of Lynceus Muell. in the
genus Daphnia, Baibo adopts here different sub-genera, as MaerothriXf
Eurycerus, Chydorus Leach, CamptocercuSf Aeroperus, AUma, PleurogDus,
Peracantha {Ann. and Magaz. of Nat. Hist. Vol. xi. pp. 81 — 95, 1843),
Bosmina (ibid. Vol. xvn. 1846, pp. 410 — 413).
Compare on this genus H. £. Stbaus Mimoires svr les Daphnia, M4m.
du MusSum, v. 18*18, pp. 380 — 415, PL 19 ; vi. 1820, pp. 149 — 162 ;
also W. Baibb, Annals and Magaz. of Nat. Hist. i. 1838, pp. 245 — 256.
PL IX.
AccmthocercuB Schoedl.
Compare J. Ed. Sohoxdleb, Ueber Aeamihocercus rigidus, ein bisher unbe-
hanntes Entomostracon. Ebiohbon's Arch. f. Naturgesch. 1846, s. 301 —
374, Taf. XI. xn.
Genus LaUma Stbaus, with bifid antennse, differs from Daphnia,
Sp. Dixphnia settfera MuEiiL.
Oeder VI. Phylhpoda,
Feet lamellose, branchial, in eight pairs or more : in some, other
natatory feet besides inserted behind the former. Two compound
eyes.
Family XII. Branchiopoda, Body naked, without shell. Eleven
pairs of branchial feet. Two eyes petiolated and a frontal ocellus
sessile.
VOL. I. 41
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642 CLASS X.
A. E. Gbubs, Bemerkwngen liber die PkifUopoden, nebtt eme Uebenickl
ihrer QaUungen wnd Arten. Mit 4 Knpfertafdn. Berlin, 1853, 8w>.
LiEViN, Die Branchiopoden der Damiffer Gegend. Mit xi To/dn in
Stdndribck, Danzig, 1848, 4to.
Branchipus ScH-fiFFER (in part), Latb., {Branchicpoda Latr.
previously, Lam.)- Abdomen caudiform, long, composed of from
six to nine segments, bilobed at the apex, or terminated by two
pinnae. Antennee four, the superior filiform or setaceous, the
inferior incurved, resembling horns, directed downwards.
Sp. BranchipuB ttoffnalis, Cancer etagnaUs li., J. C. Scrjefwmb. Der fitck-
firmige Ki^enfues, m. einer color. Kvjpferpl, Begensburg, i754y 4^. Ejusd.
Elem. erUomol, Tab. 19, figs. 6, 7, Encyd. mSth. Cruetac. PL 336, figs. 4—16,
— Brarhchipua paludoaiu Latb., Chirocephalm diaphanut Pa^osr, Dss-
MAB. Crttat. PI. 56, figs. 2—5, Gu^BiN Iconogr., Oniat. PI. 33, fig. 3.
These animals resemble in some degree the larva of the common ephemera,
so that LiNNAUB himself was in doubt whether Cancer ttagnaUe was not
such a larva. They live in stagnant fresh water, and in pools formed by
summer-showers in roads. The young animals do not at first resemble the
old ones, have a short oval body, and besides the antennae, two pairs of
jointed appendages, of which the first pair consists of two long and strongly
developed rowing feet, curved, bent backward, and beset with many hain ;
these parts afterwards change into the posterior antenn» or horns of tlie
head ; the abdomen lengthens, and only after repeated moultings acqoixes
feet, which at first are less numerous because the posterior appear later than
the anterior.
Artemia Leach, ArtemMtia Lam.
Sp. Artemia aalina, Cancer eaUntit L., Baokxt Trana, of the Linn, Soeiety,
XI. Tab. ziY. figs. 8 — 10 ; N. Jolt JIutoire cftm petit enutad, Artemia
talina Lbaoh, &c. Montpelier, 1840, 4to. This little animal lives in fresh
water at Lymington^ Montpelier, &c ; it was named by the English author
Brineworm, and is almost ^ inch long. — Branchiput Mulhauaeni Fischsb,
Artemia ealina Rathke, Fauna der Krym^ M^m. dee Sav. Hrangen pre-
aenUt d VAcad, imp. dee Sc. de St. Peterebourg, m. 1837, vi. figs. 14— «i;
in a salt-water lake of the Crimea, and probably also in Siberia.
Eulimene Latr, Abdomen short, almost semi-globose.
Compare Latbih^lb in Cuv. R. Ani, ni. p. 168, le ^t. iv. p. 178.
Family XIII. Aspidephora. Body either covered by a clypei-
form or inclosed in a bivalve shell.
A. Eyes jpetiolate.
Nebalia Leach. Shell compressed, plicated at the back, descend-
ing by the sides of the body, covering head, thorax and part of
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CRUSTACEA. 643
abdomen ; a moveable appendage, resembling a rostrum at the
anterior part of the shell. Four antennas, terminated by a long
many-jointed seta, serving for swimming. Natatory feet bifid, in
diiFerent number behind four pairs of branchial feet, lamellose.
Sp. Nd>alia fferbttii, Cancer bipet O. Fabr., Fcwn, Groenl, fig. a, p. 946 ; —
NdKdia Qeofroy Edwards, Ann. des Sc. naiur, Tom. zm. 1828, pp. 297 —
300, PL TV. GuiRiN Iconogr., Crust. PL 39, fig. 2. Compare also
Edwards Ann. dea Sc. not, le B4ne, ni. pp. 300 — 311. To this genus
belongs also MonoetUus roHrattu Montagu {TransticL of the Linn. Soc. zi.
^8' 5)> NdtaUa Montagui Thompson.
B. Eyes sessile.
Limnadia Brongniart. Body inclosed in a bivalve, oval,
compressed shelL Antennae four, anterior short, undivided, pos-
terior and exterior large, with a cylindrical peduncle and two
branches setaceous, multiarticulate. Feet (eighteen to twenty-
seven pairs) lamellose, bifid, with a filament at the base recurrent
or ascending towards the back. Last segment of body terminated
by two diverging filaments.
Sp. Limnadia Hermanni Ad. Bbongniart, Mim, da Mm. vi. 1820, pp. 83
—92, Daphnia gigae Hebm., MSm. apUrol. Tab. v. figs. 4, 5, KocH in
Hekbich-Soh^ffbr Deuttchl. Ins. Heft 185, Tftb. 10; — Limn. nMuritiana
GufeiN, Magas. Zod. 1837, CL vn. PL 11, figs, i — 11, Jcofnogr.f Crust,
PI. 33, fig. «- .
Compare also J. Kbtnioki, Des Limnadies in the BtiUet. de la Soc. imp.
des Natv/raL. deMoscou, 11. 1830/ pp. 173 — iSa, PL vn. Lvmnadia tetracera.
This writer found males, and observed the copulation. Bbononiart
observed only females.
Estheria Rueppell.
Sp. Eiiheria dahalacensis BuxFP., Straus Durokhsiii, Museum Senchenher-
gian. n. 1837, pp. 117—128, Tab. vn.
Apu8 ScHJEFP., Limulus Muell., Lam. Shell produced firom
head over thorax and a great part of abdomen, clypeiform, thin,
deeply emarginate posteriorly. Eyes two, approximate, including
an ocelliform point. Two inferior antennae very short, styliform,
biarticulate. Mandibles strong, gibbous externally, with internal
margin straight, denticulate. Two pairs of maxillas. Sixty pairs
of feet; first pair incurved, filiform, with three very long set»
exsert beyond the shell, multiarticulate ; remaining feet lamellose.
Six last segments destitute of feet ; two setss at the last segment
long, multiarticulate.
41—2
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644 CLASS X.
Tbaae animalB, on a first impreaaioiiy seem to have some conformity with
the genus LimtUus, although they are much smaller. They diSer however
from these greatly, as well in their oral organs, which are formed on the
type of insects (compare Sayiont M€m, sur Ua Ani. s. v. i. pp. 63, 64,
PI. vn.) as in internal structure. They undergo, atf already stated in
p. 613, a metamorphosis resembling that of Branchipvs. They are found
in fresh water, canals, fish-ponds, and pools formed by abundant rain.
Compare 6. C. SoH^FrsB J>er Krebsartiffe kirfet^ftus, mit 7 Uhtm,
KupferUrfdn, Begensburg 1756. 4to, and E. G. Zaddach, De Apodis can-
criformu AntUome et Hirioria evoluHonia, acced, tabula 4 liikogr. Bonne,
184 T, 4to.
Sp. Apui eaneriformit Latb., Sohavfbb Monogr, cit,, Slem, eniom. Tab. 49,
figs. I. II. ; F. H. LosoHOB NcBtutfoncker, Tom. xix. 1783, Tah. m. ;
GuiBiN Jconogr,, Onut, PL 34, fig. 1.
Another smaller species has an elongate oval plate between the two
filaments of the tail, and forms the genus Lepidurua Lbach; it is the
Aput productu8 Latb., Mcnoculut aput L., Dbbmab. Cfrtui. PL 59, fig, a,
GuiBiK, n. fig. 3.
Appendix to the Phyllopoda.
Trildbitea or Palceades.
Fossil crustaceans. Shield cephalic, rounded anteriorly, mostly
supplied with two lateral eyes oval or reniform, often extending
posteriorly on each side into a homed point. Segments of thorax
iand abdomen transverse, different in number, mostly divided by
two longitudinal furrows into three convex lobes. The posterior
part of abdomen often undivided {pygidium). Body of some con-
tractile into a ball.
Very different opinions were entertained formerly respecting the
TrUcbUes, Because the transverse segments in these fossil indi-
viduals offered some resemblance to Chitony Latreillb was really of
opinion that they ought to be placed in the neighbourhood of this
genus. Mhn, du MuBewm^ vii. pp. 22 — 32, R^gne Ani. 2e ^t. iv.
pp. 202, 203, whilst he had previously considered them to be a
transitional form between the crtistctcea and the myriapocla, between
LimtUua and Glameris (jB. Ani, 1^ ^t. in. pp. 150, 151). That
these remains belong to the type of the articulates was already
apparent to Linnjeus, who collected the few petrifactions of this
division known to him under the name of ErUomolitkiis parcuioxus.
Wahlenbebg in 1818 {Aa, Societ, Beg. Scient Ups. VoL vul p. 1),
in a memoir in which new species were also described, illustrated
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CRUSTACEA. 645
and oonfinned this opinion of Likk.£US. He thonglit that tho
TrUobUes bad the closest affinity with Limulus, This opinion was
also adopted by some other writers, although to me the affinity with
Limtdua appears certainly not to be greater, or even not so great,
as with Branchipua or Aptis, The Trilobites were probably gigan-
tic PhyUopoda which peopled the seas of a former world. The re-
semblance to the Isopoda appears to me to be rather an analogy
than a real affinity. That the Trilobites were possessed of small
natatory feet which were soft, and so lost in the petrifying, is very
probable'. They are all confined to the oldest mountain-strata, in
which petrifactions occur, which preceded the coal-formation, and
are found especiaUy in the Silurian formation of Mubchison and
other modem geologists.
Compare on this division amongst others : A. Bbongniabt in the work
published by him and Dssmabest, Hist, natttr. dea OrustacSa fostHea, Paris,
1 8a a, 4to. pp. I — 65 ;— J. W. Dalman Ueber die PaUxaden oder die ioge-
nannten TrUdbiUn ; cmt dem, Schwedisehm von Fb. Engblhabt. Mit 6
Kupfert. Numberg 1818, 4to , — ^H. Bubmeistbb IHe organiiotion der Tri-
lobiien, aus ihren Ubtmdigtn Verwandlen enivncheU, Mit 6 Kupfertafdn,
Berlin, 1843, 4to ; — E. BSTBICH Udier eimge bdhmische TrUobiten. Berlin,
1845, ^^4^9 ^ Stticke, 4to. m. Kupfert. ; — ^Db Emmbioh Ueber die TrUo-
biten in y. Lbonhabd u. Bronn, Jakrbuch fUr MinercUogie 1845, s. 18 —
67 ; and especiaUy the admirable work of JOACHIK Babbande, with a great
number of beautiful plates, Sytthne SUvrien da Centre de la Bohime, ibre
partie ; Recherche$ paUontologiquei, YoL i. 185a. Prague et Paris, 4to.
Some Trilobites did not roll their body up. To these belong the
genera:
Trtnudeus MURGHISON {Cryptolithus), O^ry^wi Brokgn., Ceraums
Green, Argea Goldf., Brontes Goldf., Paradoxvdes Buongn.,
Olenus Dalm., Burm., Gcnocephalus Zenker, Ellipsocephalus Zen-
ker, Harpes Goldf.
Sp. OUnuB Tewini Dalm., EtUomolithtu paradoxus Lutn., Mm. Temnianum
fiohniffi, t753, ^ol. Tab. in. fig. t, pp. 98, 99, Dalm. 1. 1. Tab. vi. fig. 3.
Other Trilobites were able to roll themselves up, like Glomeris and
some Oniscidesy and to bring the shield of the tail to that of the
head. They had, as it seems, a harder shell, and commonly a
larger tail-shield. To these belong :
* It was principally because these feet are not known tbat Latbeillb arrived at
the singular opinion concerning the agreement with Chiton; he says of his own accord,
that under this point of view he must consider the eyes, which are observed in many
Trilobites, to be tubercles.
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646 CLASS X.
Galymene Bronqn., Homalonotua KoENiG, Asaphus Brongk.
(Add some other genera omitted here).
Sp. Calymene Blumenbaehii Bbonon., Torbubia Apporato para la ffui4>ria
natural Egpaflola, Madrid, 1754, fol. Tab. m. 6g. 4, p. 83 ; Blumbs-
BAOH Abb. naturhiatar. Qegenttdnde, No. 50, Dalm. 1. L Tab. 1. figs, a, 3,
€k-<f &c.
Note. — On these and some other genera^ more numerous than
seems to be necessary, consult the authors quoted ; on BaJUus Dalm.
see especially Beybich, i. s. 44. Other genera, EwrypUrua Dekay,
Cytheriria HisiNroER, seem scarcely to have their place here.
Order VII. Isopoda.
Head distinct from the segment bearing the first pair of feet.
Trunk divided into seven rings sustaining seven pairs of feet.
Feet of trunk all undivided. Tail with segments different in
number up to seven, supplied beneath with branchial foliaceons
feet. Mandibles and maxillse mostly distinct. AntennsB four, the
lateral at least setaceous. Eyes two, sessile, in most composed of
a cluster of ocelli, in others compound.
Family XIV. Epicarides or Bopyrina. Antennae very short,
imperfect, resembling tentacles. Feet short, incurved, hooked.
Parasitic animalcules ; females much larger than males, broad, with
eyes none or indistinct.
Bopyrus Latr. Five pairs of abdominal feet lamellose, con-
cealed under abdomen.
Compare H. Rathke, Dt Bopyro et Nereide commentatwnea antatcmico'
phyriologiea duo:. Oum tab, iii ceneU. Rig» et Dorpati, i737> 4to.
The species hitherto known of this and the following genus live parasiti-
cally on Palcemonea {Carides), in the grill-cavity or on the ventral surface of
these crustaceans (Bopyrus abdominalii Ebobtbr on ffippofyte). The sexual
difference is in these animals when adult very great. The male b elongatey
narrow, and keeps himself hidden between the gill-plates of the female.
The female is much larger, broader and asymmetrical. There are seven
pairs of short feet attached to the trunk, and five pairs of gill-leaves (abdo-
minal feet) on the abdomen, which is divided into six rings, of which the
hindmost is very small, and bears no gills. The young animals of each
sex are on the contrary of similar form, with four pairs of feet and long
posterior antennie (Rathkb, Zw MorpKologU, ReisAcmerhrngm ami
Taurien, s. 47 — 51).
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CRUSTACEA. 647
Sp. JBopyrtu tquiUarum Latb., Monocfdut Oranfforum Fabb., Latbeillb
Otn. CrvM, €t Ins. Tab. n. fig. 4, fern,, Gu^bin leanogr,, Cruat. PI. 29,
fig. I, Bathkb de Bopyro, Tab. i. This little animal (the male is at most
i^ lines long, the female 5 lines) liyes under the dorsal shield on the gills of
PaUanonea; on the Crangones, so nearly allied to them, and often living in
the same situation, this Bopjprui as it seems does not occur (Rathkb). The
fishermen think that these parasites are young soles {Plewronedei tdUa L.)
Phryxus Bathke. Brancliise exsert, bilobed, in four pairs.
« Sp. Phryxm ffippolytea Bathkb BeUrdgt zur Fauna Norwegens, Tab. u.
figs. I — 10, pp. 40 — 56. This little animal, not different from Bopyrtu
abdanUnalUKBOKYWL (Tidakrift, ni. 1840, pp. 102 — iii, 289 — 299, Tab.
I. n.) keeps to the ventral surface of HippolyU, The full-grown female has
on one side only a single foot which is attached to the first segment, whilst
on the other side, as is usual, seven feet are found. — Phryanu crangonit
Rathkb, ibid. Tab. i, figs. 13 — 15, Tab. n. figs. 11, 11.
lone Latr. Branchial feet in six pairs, exsert, filiform, arbor-
escent In female.
Sp. OnUciu ihoracieuM Momtaqu, Tratuae, of the Linn. Soc. Vol. iz. p. 103,
PL 3, figs. 3, 4, Dbsicab. Ortut. PI. 46, fig. 10, Gu^bin Iconogr., Cnat.
PI. 46, figs. I, 2 ; compare also Audouin and Milnb Edwabdb Ann. des
Se. not. Tom. iz., Edwabdb Hut. ncA. dea Ortut. PI. 33, figs. 14, 15. This
species lives on the gills of CaUioTuusa whterrtmea,
Kepon DxJVERNOY (Cepon), Feet unarmed, with last joint sub-
dilated. Abdomiual feet with pinnated branchiae.
Sp. Cfpon t^pttf, DUYSBNOT Ann. de$ 8c. natttr. 20 S^rie, Tom. zv. 184 1,
Zociog. pp. no — 123, PI. 4, B, figs, i— 11.
Family XV. Cymothoadea. Head small. Mandibles Aimished
with a large palp. Feet of trunk in seven pairs, either all or the
anterior terminated by a hook incnrved and moveable. Body oval,
depressed. Posterior part of abdomen or tail composed of from
three to six segments, with last segment {pygidium) large, scutiform.
Feet of caudal segments lamellose ; feet of last pair terminated by
two oars or oval styles, exsert on each side near the pygidium.
Comp. the article OyrrwthoadSea by W. E. Leach, Dictumnaire dea
Sciencea natur. zii. pp. 338 — 354.
Cymothoa Fabb. Antennae short, inserted under the exsert
frons. Eyes two, lateral, situated at the margin of head. Seven
pairs of feet ; feet terminated by a hook incurved and moveable,
bent and hidden under the trunk.
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648 (LASS X.
Urozeuctua {Owrozeuktea Edw.). Tail undivided, with segments
ooalesoed.
Cymatlioa nob. Tail composed of six segments, distinct, moveable.
Sub-genera: Cym/Ahoa Lkach, Livoneca Leaoh, Anilocra Leach, Edw.
(AnUocra and CanoUra Lbach), NerocUa Leach, £ow. Comp. W. £.
Leach, 1. 1.
These animals liye parasiticallj on different fishes. The young animals
have a larger head, the eyes more conspicuous^ a pair of feet less on Uie trunk,
and a long abdomen. See MiLNB Edwabss Ann, dea 8e. not,, sec. S^rie;,
Tom. Ill, ZooL PI. 14, fig. 3.
Sp. Cymothaa cutrum Leach {Onucui cairum L. in part), Dbbhaii. OmM.
PL 47, figs. 6, 7, Cuv. JL Ani. H. ill., Crust. PI. 65, fig. i ; in the Baltic ;
— OyvMthoa Banktii Leach, Milne Edwards Ann. de» Sc. nai. 1. L fign.
I, 2 (copied in Gu^iN Iconogr,, Crutt, PI. 29, fig. 7, under the name of
CyvMtk. trigonoeephjala), from the Cape of Good Hope, &c.
Alitrcpus Edw. Antennae not covered at the base, setaceous.
All the feet of trunk slender, terminated by a hook. Eyes two,
marginal.
Sp. Aliiroput typua MiLHE Edwards, HiiL not. det Onut. PI. 33, fig. i ;
from the bay of Bengal.
^ffa Leach. Antennae not covered at the base, anterior with
basal joints broad, plane and apex setaceous, multiarticulate.
Sub-genera : Fterdas Gu^n, Rocinda Leach ; with eyes lai^gey
contiguous, or confluent anteriorly.
Comp. GniRiN Magasin de Zool. 1S36, Crust. PI. 20.
Sp. JSga emarginaia Leach, Desmar., Crust. PI. 47, figs. 4, 5; — jBga
bicarinata Leach, Rathke Beitr. zur Fauna Norwegma, Tab. VL figs, i —
18, pp. «5— 35.
Oirolana Leach.
Ewrydice Leach, Edw. {Eurydice and species of Ndocira Leach).
Serolis Leach. Antennae four, long, not covered at the base,
with basal joints thicker, setaceous at the apex. Eyes two, large,
reniform, situated in a tubercle, remote from the margin of head.
Trunk broad, marked by a longitudinal furrow on each side.
Abdomen with only three or four segments distinct, the last very
large. Feet of the first pair (or in males, of first and second) thick,
short, with strong incurved hook.
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CRUSTACEA. 649
Sp. Serolu FabrieU Lbach, Cymothoa paradoxa Fabr., Bucklaixi) Mineral,
and Gfoloffy, London, 1836. PI. 45, figs. 6, 7 (after a drawing by Curtis),
Cuv. JL Ani, id, tU,, Crust, PI. 64, fig. 3, &c. This genus in the two
longitudinal furrows on the back of the trunk bears a strong resemblance
to the fossil Trilobites, but there is no proper affinity on this account.
Family XVI. SphcBromida, Body broad, oval, frequently con-
tractile into a ball. Head large, broad. Feet not fixing themselves
by hooks. Tail small, mostly composed of only two segments, the
first five having coalesced to form one. The last pair of caudal
feet with only a single moveable lamella extended near the outer
side of pygidium. Foot-jaws resembling palps.
ATlcinits Edw. First and second pairs of feet terminated by a
large moveable hook, subcheliform ; remaining feet gressorial,
slender, terminated by a straight claw. Body depressed, plane.
Antennae long, setaceous.
Sp. Ancin/uM depresttu Milnb Edwards, ffist, not. dea Orutt, PL 37, fig. 17.
Sphceroma Latr. Feet of trunk all gressorial, slender, termi-
nated by a short claw, often bifid. Body gibbous above, contrac-
tile into a ball {Onisci globatores).
Sp. Sphceroma aerratum Leach, Onitcva globator Pallas, SpicU, Zool. iz.
Tab. 4, fig. 18, Dbsmab., Crust, PL 47, fig. i ; from 2'" to morerthan 5"'
long, in the Atlantic ocean, the Mediterranean and the Black Sea.
Note. — Grenus Zuiara Lbach is scarcely distinct.
Add sub-genera : Cymodocea Leach, NcRsea Leach (and CUiccBa
ejusd.), Camnpecopea Leach, Cerceis Edw., Amphoraidea Edw., Cassi-
dina Edw.
Family XVII. Pramztdea. First two segments of trunk in-
distinct or joined with the head, destitute of feet. Five pairs of
feet of trunk. Antennee four, setaceous, unequal. Posterior part
of abdomen or tail with six segments, supplied with foliaceous
appendages at the apex.
Anceus Risso, Onaihia Leach. Head large, almost square,
armed anteriorly with two horns porrect, falcate (mandibles?).
Sp. Anceua fot^icularius Risso, Dxbmab., Crust. PI. 46, fig. 7 (not fig. 6),
GufiwN Iconogr., Crust. PI. 27, fig. 7; — Anceus maxillaris Lam., Cancer
maaeUlaris Montagu, Linn. Trans, vii. 1804, Tab. vi. fig. «, Dmmab.
Crust. PI. 46, fig. 6.
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650 CLASS X.
Prantza Leach. Head triangulax, narrower than thorax. Man-
dibles not exsert. Trunk in females consisting of only three
segments, the last three being confluent, and forming a single
oval scutum broader than the preceding segments.
Sp. Praniza ecendatc^, Onitcui marinus Slabbkb, Slabbsk Natuurk. Verftut,
Tab. IX. figs. I, 1 (copied in Eneyd. nUth., Int. et CrtuL PL 519, figs. 24,
35), Montagu Trans. Linn. xi. Tab. iv. fig. 2, &c.
Compare on this genus J. O. Wbstwood, Ann. det Se. natur. Tom. xxyh.
1831, pp. 316— 33«i I*L 6-
Family XVIII. Ontsctdes. Two middle antennae very short,
of only one or two joints. Last segment of tail short, with the
appendages of the last pair of caudal feet concealed or styliform.
Comp. J. F. "BsLAunn Conspectus Monographice Oruslaceorum Oniseodorum
Latbbillii, Bulletin de la 8oci4t£ imp. des Natural, de Moscou, Ti. 1833,
pp. 17T — 193, Tab. IV. figs. 6 — 41.
These Isopoda may be named Land-AssdUy for although some of
them keep to the searshore, they do not live in water. The most
live under stones, bark of trees, in chinks of walls, &c. They gnaw
various substances. At night especially they seek their food, which
consists principally of vegetable matters.
A. Oniscinea. Basal joint of the last pair of abdominal feet
short, not produced beyond the last segment of abdomen. Antennse
geniculate, with from six to nine joints (the five basal joints thicker,
the terminal seta with one or two, three or four joints).
T^los Latr. External antennas moderate, with nine joints, the
four terminal joints forming a short seta. Last pair of abdominal
feet lamellose, concealed under abdomen. Body contractile into a
ball.
Sp. Tylos Latreillii Audouin, T^los armadiUo Latb., Cloporte Savtgnt,
Deser. de VEgypU, Crust. PL 13, fig. r, Gu^iN Iconogr., Crust PL 31,
fig. 4, Edwards, Cov. R. Anim. H. iUtutr., Crust. PL 70, fig. a.
The first four pairs of abdominal feet have a broad quadraogular append-
age, in whicb longitudinal cavities with blind branchings are found. They
open at the inferior margin by a row of small foramina, and take up air
for respiration. The animal lives on the coasts of Egypt and Algiers.
Armadillo Latr. External antennse with seven joints. Styli-
form appendages of last pair of feet short, not prominent. Body
contractile into a ball.
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CRUSTACEA. 651
Sp. Armadillo variegatus Latb., ArfnadHlidium puitulatum Edwards,
YiLLXRS Entom. Linn, Tab. xi. fig. i6, Dbbmarkst Crrul. PL 49, figs.
6, 7, &c.
To this division belong the MiUepedce, more used in medicine formerly
than at present. Armadillo officinarum, Bbaitdt. u. RaTzkburo Mediz,
ZooL II. Tab. 43, figs. 8—10, Cuyier R. Ani. id. ill,, Crutt. PL 17,
fig. 4 ; from the south of Europe and Syria.
Sub-genera : ArmadiUidium Brandt, Diploexochus ejusd., Cvharia
ejusd, ArmadiUo ejusd.
Oniscus nob. (species from genus Ontscus L., Porcellio, Ontscus,
Philoscia Latr.). External antennae with from six to nine joints.
External appendage of last pair of abdominal feet styliform, exsert.
Deto Gv^XDS, External antennse with nine joints, the four ter-
minal joints small, forming a very short seta. Styliform appendages
at last segment of body elongate.
Sp. Deto tekynata Gu&in, Magcu, de Zool, 1836. Crust, PL 14.
Platyartkrus Brandt.
Trichoniscua Brandt.
NoU. — In these genera the external antennse are six-jointed.
Oniscus Latr. External antennsB with eight joints, inserted
under the anterior margin of head, which is somewhat prominent.
Body not contractile into a ball.
Sp. Oniacus asellut L., Oniscus mvrariiu CxTV., Dk Gxer Mim, p, s, A Vllist.
des Ins, vii. PL 35, fig. 3, Gboffr. Ins. n. PL xxii. fig. i, Brandt, u.
Batzeburo Medizin. ZooL ii. Tab. xii. fig. 7 ; the edlar-oniscus, mostly
7'" or 8'" long, ash-grey above, with yellow spots on the side. This little
ftTitmal is very common on walls in moist places.
FarceUio Latr. External antennae with seven joints. Body in
most not contractile into a ball.
Sp. Parcellio scaber Latr., Brandt u. Batzbburo Mediz, Zool. n. Tab. xii.
fig. 6, Gu6rin Iconogr., Crust, PL 31, fig. 7. In PorceUio and Armadillo
(not in Oniscus) the external plates or coyers of the two anterior abdominal
feet have a whitish body, which is hollow internally, and receives air from
a fissure by many openings. See Duyernot et Lkrbboullet Ann. des Se,
nal.,2e B6ne, Tom. XV. 184 1. Zool. pp. 197, 198; comp. V. Sirbold in
Mueller's Archiv, Jahre^>ericht, p. 141. Lereboullbt found a similar
structure in the external plate of the first five abdominal feet of two species
of PorceUio, which roll themselves up, whence he named one of these Pore.
armadUloides. Already Ds Geer had noticed such a species that rolls
itself up, Oniscus eonvexus, vii. p. 553, PL 35, fig. 11.
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652 CLASS X.
Philoscia Latr. External antennse with eight joints, naked at
the ba^ie. Segments of tail abruptly narrower behind the last
segment of trunk.
Sp. Onitctu mtucorum Guv.
B. Ligiea, Basal joint of last pair of abdominal feet slender,
elongate, exsert, terminated by two long styles. External antennse
terminated by a seta composed of several (twelve to twenty) joints,
long.
Ltgta Fabr.
Sp. Ligia oceanica Fabb., Basteb Naiuurk, Uitspann. n. Tab. xm. fig. 4,
Desmab. Oriut. PL 49, figs. 3, 4 ;— Ligia italica Fabr., Gu^tK leonogr.,
OrvM. PL 31, fig. 5, &c. These animals live od the sea-coast, but often
leave the water and climb on stones, piles, kc This genus forma the
tnmsition to the water-onisci.
Some s])ecies, in their antennse resembling Ligia, have at the last
abdominal feet a bifid basal piece, and at the extremity of these two
pedicles is a filiform appendage. They live under stones or under
moss, in moist meadows, in decaying wood, <fcc They form the
genus Ligidium Brandt, Zia Koch, in Hekrich-Schjsffer DeuUehL
Ina. Heft 180.
Sp. Ligidium Pertonii Brandt, Oniscus agilit Pbbsoon, Panzer DeuUM.
Ins. Heft 9, Tab. 14, ftc.
Family XIX. Asellota, Antennse four, distinct. Terminal
appendages of last pair of abdominal feet produced beyond the
body like a tail. Last segment of tail large, scutiform.
A. First pair of feet, either similar to the rest, or thickened at
the apex, and with hook single.
f Four antennae unequal, external much longer than internal.
* All the feet similar, tenninatcd by short double claw.
Oniscoda Latr. {Jantra Leach).
Jasridina Enw.
Sp. Janira Nordmanni Rathkk, Fauna der Krym, p. 388, Tab. VI. figs.
JcBra Leach.
Sp. JcBra nivalia Kboetxb, Cfrdnland'i Amfipoder, TVb. iv. fig. «i.
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CRUSTACEA. 653
* * Anterior feet short, thickened at the apex, furnished with hook in-
curved.
Munna Kroeyer. External antennae very long. Feet, except
the first pair, longer than body. Tail destitute of appendages.
8p. MuTvna BoeclcU Kboeteb, Tidnhrifi, ii. 1839, PP> 61 a— 616, with plate.
(The specimen, described by Ejlostsb, seems not to have been full grown).
AsdliM Geoffr., ScHiEFFER, Latr. External antennae elongate.
Feet shorter than body. Two abdominal appendages terminated by
two elongated filaments.
Sp. Asellua vtdgarU Latb., OniteuB aqualiciu L., Gbofvb. Jn$. n. PL 22,
fig. 2, Db Gsbb M^m, p, t. d Vffiat. dea Ins. vii. PI. 31, Desmab. CrtttL
PL 49, figs. I, 2, Tbevibands Verm, Schr, i. Tab. x. figs. 56, 57. This
species is very common in wet ditches, and is almost i" long. The little
animal passes the winter in the mud. It differs from the genus Onisctu, to
which LiNNJBUB united it, in the tail, which is covered above by only a single
shield-like plate, and further, in the greater development of the middle
antenns and in the feet, which increase in length from the head to the tail.
Limnoria Leach. Four antennae short, subequal. Body cylin-
drico-linear.
Sp. Limnoria terebroms Leach, Trans, of (he Linn. Soc. xi. pp. 370, 371 ; a
small animal (i — 2'") which perforates the piles and wood- work of marine
jetties, and in a short time may destroy them.
B. First pair of feet short, thick, terminated by a didactylons
chela.
Apseudes Leach. Tail composed of six segments, terminated
by two long filiform appendages of the last pair of feet.
Sp. Apseudes talpa Lbach, Cancer gammarus talpa Montagu, Trans, of the
Linn. Soc. ix. p. 98, PI. 4, fig. 6 (this figure is copied in the JEneyd. mSthod.,
Crust. H Ins. PL 336, fig. 16, Desmab. Crust. PL 46, fig. 9). Milne
Edwabdb has given a better figure, Cuv. R. Anim. id. iU., Crust. PL 61,
fig. I.
Tanaia Edwards.
Sp. Tcmais Cavdinii Milne Edwabdb, R4sumi d^Entom. pa/r MM. Audouin
H Edw. {Encycl. portative, 1819), i. p. 181, PL 29, fig, i, Edw. Hist. not.
des Crust. PL 31, fig. 6.
Comp. on this genua Kboeteb, Naturh. Tidsskr. IV. 1842, pp. 167—187.
Tab. II. figs. i—M.
Zeuxo Templeton.
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654 CLASS X.
Crossurus Rathke. (Not different perhaps from genus Tanais),
Note, — ^Add genus Rhoea Edw. {Ann. des Sc N(tL xni. 1828,
pp. 292—296, PL 13 a), unless it be better to refer it to tke
Amphipoda. Genera Eupheus and Olisca Risso, seem to be un-
certain, and require confirmation by more diligent observation.
Perhaps also will have to be placed here amongst the Isopoda, OniuMt
arenarius Slajbber Naiuurk, VeriuM, Tab. n. fig. 4, genus Ptaygocera
Latb., Cuv. M. Am. and ^ 1829, 17. p. 124.
Family XX. Idoteidea. Tail mostly composed of only three
distinct segments, with last segment large, scutiform. Abdominal
feet branchial; last pair of feet forming opercular laminsB covering
the branchiae beneath. No appendages exsert in the last segment
or at the sides of tail. Antennae four, very frequently unequal, the
middle approximate. Mandibles destitute of palps.
Anthura Leach. Four antennse, short, subulate. Body slen-
der, vermiform. Anterior feet incrassated, terminated by an in-
curved hook. Tail with two distinct segments.
Sp. Anthura gracilU Leaoh, Onisctu gracilia Moktaou, Desmab. OiuC. PL
46, fig. 13, MiLNB Edwards Hist. not. des Crtut. PI. 31, figs. 3^5.
Idatea Fabr. Middle antennse short, with four joints, the last
elongate, cylindrical; external antennee large, with first five joints
thicker, the rest more slender, running into a terminal seta. All
the feet of trunk terminated by an incurved claw; those of the first,
second and third pairs raptorial, incrassated at the apex and turned
forwards. Branchial feet of tail covered by the last pair of feet
changed into two valves articulated by means of a hinge to the
last segment.
Sp. Idotea entomon Fabb., Onitetu entamon L. (in part), Pallas SpiciL
ZooL IX. Tab. v. figs, i — 6, Ds Gebb MSm, p. s. d rffitt. de» Ins, yi. PI.
33, figs. I — 10, Bathkb Beitr. zwr Oesch. dtr 7%t«no6ft, i. Danzig, 1820^
Tab. iv. This animal was found in the Baltic Sea ; it attains a length of
\" 9"'. Its two small black eyes are situated at the side of the head in a
round pit, and are not, as is erroneously supposed, simple, but each of them
consists of a group of more than 50 small oval eyes placed dose together. —
Idoita Lichtensteinii Kbaubs, SOdqfrik. Cfrustaceen, Stuttgart^ 1843, Tab.
IV. fig. 4.
Idotea Unearis Latb., Stenosama lineare Lxaoh, Babteb Natuurk, UUap.
n. Tab. xin. fig. 1, Bbsmab. Crust, PI. 46, fig. 11; from the Baltic, about
i" long.
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CRUSTACEA. 655
To this division also belougs Onucua viridu of Slabbbb, Natuwh. Ver-
lust. PI. 12, figs. 4, 5, which, according to MiLNS Edwards, forms a
distinct genus.
Arcturvs Latr. Feet of first pair short, palpiform. Feet of
second, third and fourth pair ftimished with long hairs, with last
joint plane, elongate. External antennas of the length of body.
Sp. Archmfs Bafini Webtw., Idotea Bafini Sabine, Milne Edwards ^uf.
not, des Crust. PI. 31, fig. ij — Arctunu longicomit Westw., Gu^in
leonogr,, Crutt. PL 31, fig. 2; in this species the first four pairs of feet
stand very remote from the last three.
Order VIII. AmpMpoda,
Eyes sessile. Four antennae, the two middle inserted above the
lateral. Mandibles mostly furnished with palps. Two pairs of
maxillae and one pair of foot-jaws. Trunk mostly divided into
seven distinct segments, bearing seven pairs of feet not cloven into
two oars. Membranous vesicles for respiration adhering to the base
of some of the feet.
We unite here, after the idea of Kroeyer, the LcBmodipoda of
Latreille with the order of the Amphipods ; see Kroeyer Tid9-
shift, IV. 1843, pp. 490—495.
Section I. LoBmodipoda, Posterior part of abdomen or tail
short. First segment of trunk conjoined with head, an oblique
furrow alone on each side indicating the separation; feet of this
segment inserted forwards, beneath the head. Two or three pairs
of branchial vesicles in the anterior part of Irunk.
Lasmodipoda from Aai/uio« and hiwov^; these aminala were thus
named because they have two feet under the head, as though at the
throat. They do not swim, but creep on marine plants and animals
in search of food. The posterior part of the body is little developed.
In most the feet are wanting in those rings that carry gill-vesicles,
and conversely the gill-vesicles are wanting on those rings that have
feet.
Family XXI. Lcemodipoda, (Characters of the section).
Phalanx I. Cyamea. Body oval, mostly much depressed, with
transverse segments. Antennae with four joints, the last slender;
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656 CLASS X.
inferior anteim® small. Two pairs of cylindrical branchise at the
second and third segment of the trunk, in place of feet Feet
strong, terminated by an incurved hook, in five pairs. Mandibular
palps none. Females furnished with large laminae at the base of
branchiae, covering the eggs.
Cyamus Latr. (Species of Oniscus L.).
Sp. Cyamui eeti Latb.^ Oniscus eeti L., Pycfiogonwnceti Fabs., Pall. Spk.
ZoU. Fasc. IX. Tab. 14, Ds Gkeb Mhn. p. serv, d VHist, des Ins. vn. pp.
540 — 544, PI. 42, figs. 7, 8, Tbeviranus Verm, Schr. n. pp. 3 — 10, Tab. L
Thifl GruBtacean, whose length is about i" and more, lives on JBakma Umgi-
mana; Cyamus errcUictu RouBSEL DB Yauz., on the whale of the southem
hemisphere, is an allied species. Cyamtu gracilis Roues, db Yaui. is
smaUer and more elongate than the other species, and thus approaches to
the form of the next division.
Comp. on this genus, besides the anatomical notices of G. R. TBEYiBAinis^
RoussEL DE YauzAmb, Ann. des Sc, not., ae S^rie, Tom. i. 1834, Zool. pp.
439 — »65, PL 8, 9, and Kbobteb Natuurh. Tidsskr. nr. pp. 474—489.
Phalanx II. Gaprdlina. Body mostly elongate, the length of
the segments surpassing their breath. Feet slender. Branchi»
vesicular at the base of feet. Mandibles often furnished with
palp triarticulate. Superior antennsB setaceous at the apex,
multiarticulate.
Caprella Lam. Five pairs of feet, arranged in an interrupted
series, all terminated by a subcheliform hand. Two pairs of bran-
chial vesicles in the second and third segments, in place of feet.
Abdomen very minute, made up of a single or of two segments,
supplied with appendages or rudiments of feet.
Sub*genera: (7<ipreZZaand^^f»naKB0BTEB. Sp.Copfie^^tncarwLATB.,
Cancer linearis (L. X), Hbbbst Kraihen und Krebse, rr. p. 141, Tab. 36,
figs. 9, 10 \— Oniscus scdhpendru^desYAi^,, Spic, Zool. ix. Tab. 4, fig. 15,
Caprella lobata Latb., Gu^in Jconogr., Crust. PI. iS, fig. 1 ; to this or
to Capr. acviifrons Dbsh. the figure of Babteb, Naiuwrh. UOsp. r. PL rr.
fig. 1, seems also to belong.
Podaliriu8 Kroeyer.
Gercopa Kroeyer. Three pairs of branchial vesicles, the first
pair at the base of the second pair of feet, the rest in place of
feet. Abdomen distinct qidnquearticulate.
Sp. Cerrops ffolbodli Kroeter 1. 1. Tab. VL figs, i — 13.
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CRUSTACEA. 657
Leptomera Latr. {Proto Leach). Seven pairs of feet, arranged
in a continuous series. Fifth pair of feet shorter, furnished with a
small claw, slightly moveable. Three pairs of branchial vesicles at
the bases of the second, third and fourth pairs of feet. Abdomen
small, uniarticulate, with four appendages, elongate, biarticulate.
Sp. Leptomera pedaia, Ckmmarut pedatu$ Abildoaabd, Mubllbb Zool,
domic. Tab. loi, figs, i, i, GuiBiir leonogr,, Onut. PI. ^8, £g. 3. Aooording
to Kbostbb Leptomera vetUrieoea Dbshabkst, SquiUa vetUrieota Mubll.
Zool. dan. is the female of this species. To it belongs also Phthisica
marina Slabbbb Naiuurk. Verluet. PL x. figs, i, 1.
The genos Proto of Dbsmabxst {Crtut. p. 176) rests on imperfect observa-
tions, and appears not to differ from Naupredia Latb.
Section II. Genuine Amphipoda. Head distinct from the first
segment sustaining a pair of feet Segments of trunk mostly fur-
nished with a distinct lateral part {epimeron). Mandibles supplied
with palp. Tail made up of several segments, furnished with bifid
feet. Body mostly compressed, arcuate.
Gomp. MiLNB EowABDS Ann. des Sc. nat. zx. 1830, pp. 353 — 399, and
H. Kbobtbb Oroenland^t Amfipoder. Kjdbenhavn, 1838. 4to. (printed sepa-
rately firom vn. Th. der Kongd. Damhe Videntkabemes SeUhabt naturvid.
og mathem. Afhandlingerj p. 129, Ac.)
Family XXII. Hyperina or Uroptera. Foot-jaws smaU, not
covering the cibarious organs. Head large. Last abdominal feet
at the apex of tail mostly resembling a fiabelliform fin, foliaceous.
Oxycephahia Edw.
Pronoe GuiSrin.
Typhis Kisso, Edw.
Sp. TyphiiferuB Edw. Ann. dea Sc. not. 1. 1. PI. 1 1, fig. 8, Guisnr Iconogr.,
Crud, Fl. 17, fig. 8 ; in the Atlantic near the Canary Islands. This genus
and the preceding are remarkable for their long inferior antennn, which
are bent at three acute angles.
Phronima Latr. Antennse two short* Fifth pair of feet elon-
gate, terminated by a hand broad, didactylous. Tail elongate,
with five distinct segments, the sixth segment coalesced with fifth.
Sp. Phronima $edeniaria Latb. , Cancer 9edeniariu$ F0B8K., H1BB8T Erabben
und Krebte, n. p. 136, Tab. 16, fig. 8, Latb. Qmsr. CrueL et Int. Tab. u.
figs. 1, 3 ; this animal is found in the Mediterranean, where it lives in a
kind of gelatinous keg, open at both ends, probably from a dead Beroi.
VOL. I. 42
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658 CLASS X.
Primno Gu^RlN.
Themisto Gu^rin.
See GulRiN JI£Sm. delaSoc, tTffiit. ntU. de Parit, 19. pp. 37^3^i
PL 43 ; Kboxtbb Oroenl, Amjip. pp. 63—68, Tdh. TV, figs. 16, 17.
Phraatna Bisso, Dactylocera Latb.
Anchylomera Edw.
Hieraoonyx Orjsisxs,
Hyperia Late, Antenn® four. Feet slender, terminated by
an incuryed hook. Trunk broad, gibbous above; tail attenuated.
Zestrigonua Edw., Tyro Edw., Phorcus Edw., Hyperia Edw.,
Jf<5^oecfw Kbobteb.
8p. Byptrioi LatrmXHi Edw., Awn, <ki &. wot. zx. PL 11, figB. 1—7.
GuiBiN Iconogr,, Orutt. PL 95, fig. 5; ffidla Or&^ii Stbaub-Dubge*
Hxnc, MSm, du Mui. xnn. 1829, PL 4.
Note, — ^Genus Davra Edw., is distinguished by having only two
short anteniue, and by the second pair of feet being didactyloos.
MeMkvA ElaoETEB, furnished with four short subulate autenne,
differs from the other Hyperia by the first and second pairs of feet
being shorter and terminated by a didactylous chela. Gmenl.
Amfip, pp. 60—63, Tab. iv. fig. 15.
Family XXIII. Oammarina. Foot-jaws large, conjoined at
the base, forming an accessory inferior labium, covering the ciba-
rious organs; cloven anteriorly into two laminae, and on the outside
sustaining an articulate palp. Last caudal feet terminated by
appendages styliform, more rarely foliaceous, not resembling a fin.
Head congruous.
Gomp. H. Eboitxb Nye norduhe Slagter og Arter, kenJOrende til Famir
Um Oammarina, TSdnhnfi. iv. 1843, pp. 141 — 166.
A. Lateral parts of segments of trunk [epimerc^ moderate, not
covering the bases of feet. Abdominal feet six; the last terminated
by two small oVal laminae.
Vibilia Edw.
Corophium Latr. Two inferior antennae large, thick, pediform.
Sp. Cwrofhivm lonffieome. Cancer grostipei L., Oni$Gu$ wlutator Pill.,
MieeeU. Zocl. Tab. it. fig. 30, J^. Zod. ix. T»b. 4, fig. 9, Gbovot.
Zcophjflae, Tab. xvn. fig. 7, Gu^rik Iconoffr,, Onut. PL 27, ^. i.
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CRUSTACEA. 659
Add genera : Cerapua Say, AtyUia liKAca, Fodoeenu Leach, &c.
Gomp. MiLNB Edwabds, ffiH. NaL des Crust, m. pp. 6% — 70.
Genus Erickthanius Milnb Edwabbs, on the authority of Kboeyeb,
does not differ from Podacerus^ but rests on a sexual difference of
the males. Tidsshr. lY. 1842, pp. 163, 164.
B. Four first segments of tnmk with epimera large, covering
the bases of feet. Abdominal feet six; the last terminated by
styles, serving with the apex of the incurved tail for jumping.
Body depressed. Anterior either all or the inferior terminated
by a filament setaceous, multiarticnlate.
TaUirus Latr. Mandibles furnished with a rudiment only of
palp or destitate of palp. Superior antennas not longer than head,
shorter than the pedicle of inferior; inferior elongate*
Gtonera : TdUtrue and OrehegUa Leach.
Sp. TaHifUM ioUator E0W.» C(xncer loeutta L., Pall. Spie. Zool. ix. T»b. 4^
£g, 7, Dbsmab. Onut. PL 45, fig. 2, Cuv. JL AnL id. ill, Onut, PI. 59,
fig. 3. ThiB little animal, about )" in size, is veiy conunon on the Dutch
coast, where it is constantly progresBing in the sand by jumping.
Orchettia is distinguished by the sheer^shaped form of the first and
second pairs of feet; the second pair has a large oval joint at the apex, on
which the moveable curved daw is set; here belongs Orehedia Uttorta
JjitAOH, Basteb Natuurk. Uittp. n. Tab. 3, figs. 7^ 8, Dksicab. Crutt,
PI. 45, fig. 3.
Gamfnarus Latr. (Species from genus Cfammarus Fabr.).
Mandibles famished with palp. Superior antennee always longer
than the peduncle of inferior, sometimes longer than inferior
themselyes.
Sub-genera: Lysianassa Edw. {Anoni^ !^boeter), AUbrotua
Edw. (JJyaicmoMa previously), Phliaa Qcvisss^ Actmthonotus Owen,
Iseta Edw., Anisopua Templet., Amphitoe Leach, Gammarut Edw.,
Isehyrocerus ELboeteb, Leucothoe Leach {Lycesta Say.) and some
others proposed by Kroeyer 1. L
Sp. ChtMMuruM fuvialUU Edw., OammaruM ptdex KooB, B<bs>l /im. m.
Suppl Tab. 62 ; KocH and Gbbtais distinguish this species as Oammarut
Raudii from Chmmarut ptdex Fabb., Dbbxab. Onult, PI. 45, fig. 8, Qaimr
marus fotwrvm KooH in Hsrrioh-Sohafrb DtniUchl. Int. Heft 138,
Tab. I. L1KKAU8 has confounded not only these two species, but also
42—2
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660 CLASS X.
othen, M OrehaHa UUorta, under the niune of Ccmeerpidet, SjfA Mi, ed.
la, I. p. 1055 ».
OamnuMTUS ptdex Fabb., Gsby. is rerj oommon in ponds. The female
carries her eggs under the body »t the hase of the feet» until the young
emerge. B<B8IL ssw the animals nibbling »t roots, fraxtB, and other parii
of plants which he gave them. Usually, however, they live on animal
food, and according to Di Geer, do not spare eren the bodies of their
companions. They attain a length of about 1". The hinder end of tlie
body is always bent downwards. Compare also on this species J. C
Zevkbb J>e Crammari pulicis Fabb. ffiM. naiurali et »anffuim$ dratitu
commentaHo, Aecedit Tab. cm. Jens, 1833, 4to, and, on the diffuenoe of
the two species, Gbbvaib Ann. da 8e. ntU. le S&ie, iv. 1835, Zodit^
pp. 117, 138.
Order IX. Stamatcpoda.
Two compound ejes, placed on a biarticnlate moveable petiole.
Antennas four, placed nearly in the same horizontal row; internal
long, terminated bj two or three multiarticulate sets, extenud
mostly famished at the base with an oval ciliated lamina. Feet of
trunk mostly in seven or six pairs, more rarelj eight. Mouth
inferior, often situated between the bases of anterior feet. Branchiae
mostly adhering to the caudal feet, more rarely to the thoracic, never
covered. Tail mostly terminated posteriorly by a foliaceous pinna.
Shell thin, in many almost membranous.
Family XXTV. Unipekata, Body elongate, narrow. Cephalo-
thoracic shield elongate, often produced over the back of the ante-
rior segments of trunk. Anterior segment of cephalothoracic shield,
bearing the middle antenna and eyes, distinct. One pair of max-
illiform feet slender, elongate. Seven pairs of feet in trunk or
thorax; first pair large, with last joint mostly felcate, dentate in-
ternally; second, third and fourth pairs shorter, approximate to
mouth, terminated by an oval hand and moveable hook; three last
pairs slender, remote, mostly furnished with a styliform appendage.
Of caudal feet six pairs; the first five mostly bearing branchite of
^ From a passage in the Fanna Suecica p. 246 "habiieU ad Uttora marit vulgar
HmvMU* it would seem, that Liknaub probably by Cancer pulex had a species in viev
different from (romffiantf pulex or JiuviatUis, perhaps Oammartts loeutta Fabr. ; that
he also confounded OrchetHa with it appears, however, from his citation of Bastbs.
Henoe it is impossible to quote with justice Cancer pulex L. as a synonym of »
determinate species.
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CRUSTACEA. 681
many filaments set pectinatelj on a peduncular cirrus; last pair ter-
minated bj two laminsB oval, depressed, forming the pinna of tail.
SquiUa Fabb. (in part). Cephalothoracic shield tripartite from
two longitudinal furrows, not covering the last segment of trunk
at least, mostly not covering the three last segments. Branchias
distinct adhering to the first five pairs of abdominal feet.
Comp. Latbsilue Encyd, mith., BUt, not,, Int, Tom. x. pp. 467 — 475.
Caronia Latb. Six last feet of trunk with appendage oval, plane,
membranous.
Sp. CoronU ioolopendra Latb., GuiBiN Iconogr,, Onut» PL 14, £g, 1, &c.
SquiUa Latr. (and Oo7U>dactylu8 ejusd.). Six last feet of trunk
with appendage elongate, styliform.
Sp. SquiUa BcyUairw Fabb., Buhphiub AnA, BariteUkam, Tab. ni. fig. r ;—
SquiUa ehimgra Hkbbst, KrdNtm, Tab. 34, fig. 1, Latbeillb EneycL
mith,, Crutt. et Ina, PL 325, fig. a. These species, in which the tennioal
joint of the shears is almost or entirely toothless and swollen at the base,
form the genus Qonodactyhu Latb. In most species this joint is small,
canred and densely toothed within. As, for example, SquUla mantU Fabb.
(Cancer immH» L. in part), Dbsmab. Croat, PL 41, fig. 9, from the Medi-
terranean, SqviUa maculata Bumfh. Tab. dt. ^. b, finom the Mo-
luccas, Ac
SquiUerichthm Edw.
Erichtkus Latr., Smerdis Leach. Shell of cephalothorax not
sulcate, produced over the last three segments of trunk. Rudi-
ments alone of branchiae, adhering to the first pair of caudal feet, or
branchisB none distinct.
Sp. Brichlkm vUreui Latb., SqttUla vtirea Fabb., Smerdit vulgarit Lxaoh,
Dbbm. Cnut. PL 44, fig. 1, Ac.
Alima Leach.
CJomp. on this and the preceding genus, Lbaoh in Tuokbt's Nourratvfit
of on JBxpedition to explore the river Zoiire, London, 181 8, 4to, pp. 415,
416, and fig. PL xi.
Family XXV, BipeUcUa. Cephalothoracic shield membranous,
pellucid, produced over the segments of trunk. Middle antennas
terminated by two filaments. Six pairs of feet slender, elongate,
furnished with a cirrus multiarticulate, pilose.
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662 CLASS X.
FhyUosoma Leach. Body depressed, covered by two shidda,
the first oval, the second transverse, placed above the trunk. Eyes
set upon a long petiole.
Gomp. LXAOH in Tuokit's EtpedU, to the Zaire, pp. 416^ 417, and fig.
PL II., Journal de Physique, Tom. 86, 18 18, pp. 306, 307, Gii^Bnr Jf^
iur rOrgamitaiion dee PhyUoaomee ft Mcnographie de ce genre de CVatif.
Magasin de Zocl. 1833, CL vn. PI. 6 — 13, C. Gigknbaue UeberPhfimma
in SiEBOLD and Koilltksb Zeiitch. /. wieaeiMch. ZooL v. 1853, p. 351-
The latter author, from *the consideration of the circulationy which in lis
more minute details agrees with that of the decapods, thinks that PkyUo-
soma ought to be arranged with these.
a) Eaotemal antenna setaceous, longer than internal. Abdomen distinci
firom trunk, divided into segments, with terminal innna.
Sp. Phyllosoma commune Tuokkt 1. 1., ¥rith fig., Dxsmab. OrwsL PL 44t
fig. $,—PhyUosoma davicome Tuokbt ibid, with fig., Dbskab. ilnd. fig. 4*
b) EaOeimdl anOmnos lamellote, increased by a lobe at the baae, barter than
internal. Abdomen yery short or triangular, oontinuous with the shield
of trunk.
Sp. Phyllosoma laticome Tuokbt L L, Cancer eassideus J. Fobsteb, Natar-
fors^Ker xvii. pp. ao6 — 213, T^b. 5.
Most of the species come from the Indian and Afiican seas; yet one
species from the Mediterranean is known. These animals gaye occasiofD to
the name of the fiunily which does not apply to the following genus.
Amphvm Edw. Shell elongate, oblong, continaoas, covering
head and all the segments of trunk. Tail narrow, conical, com-
posed of seven segments, terminated by a flabelliform pinna. Ex-
ternal antennas long, famished with an oval scale at the base,
filiform.
Sp. Amphvm Iteynaudii Milkb Eowabds, Ann. de la Soe. entomoL L 1831,
pp. 336—340, PI. la ▲, ffist. not. des Crust. PL 18, fig. 8.
Family XXVI. Caridundea or Schtzopoda. Shell covering
head and trunk, descending to the sides over the bases of feet, com-
pressed. Abdomen long, made up of seven segments, the seventh
with two lamellsB of the last abdominal feet on each side composing
a flabelliform pinna. Feet of trunk slender, natatory.
This fiunily forms by its habiius the transition to tiie folloiring)
from which, however, it differs by the absence of shear-shaped feet
and by the gills not being included within the shell
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CRUSTACEA. 663
Leudjkr THOMPSON, Edw. Head narrow, produced into a pe-
duncle sustaining the antennsa and petiolate eyes, longer than the
thoracic shield. Abdomen slender, elongate, with segments oblong.
(Branchiae unknown.)
Compare Eowabdb HuA, not, dea Cnut. n. pp. 467 — ^469, FUte 36,
fig. TO. Habit, excepting the head, nearly that of Alima,
Thyaanopua {Thysanopoda Edw.). Shell anteriorly acuminate, or
supplied with a rostrum. Maxilliform feet none. Eight pairs of
feet of trunk; first seven slender, supplied with a cirrus or long
external oar. Branchiae ramose, fasciculate, adhering to the base
of thoracic feet, free. First fire pairs of abdominal feet bifid,
natatory.
Sp. ThjfMnoput trieutpidaiut MuiKB Edwards, Atm,dei8c. not, Tom. XUL
1830, pp. 451—460, PL 19, ffitt, not, de$ Crmt. PL a6, fig. i.
Cynthia Thompson.
Myais Latr. Shell with rostrum none or very short anteriorly,
deeply emarginate posteriorly. Antennae long, external, with a
basal lamellose appendage. Two pairs of maxilliform feet. Six
pairs of feet of trunk natatory, furnished with a long external
articulate cirrus. Branchiae none distinct.
Sp. Mysis JUaeuoia, Cancer Jiexitosus Muell.^ Zod. danie. Tab. 66, figs. 1—9,
Hbbbbt KttMen, Tab. 34, figs. 8, 9, Bathke Beitr. eur Fauna Norweffens,
pp. 18 — 20,--JtiynM oculata, Cancer oculahu O. F. Fabbioo Fauna GranU,
pp. 945, fig. I (copied in Hbbbst Tab. 34, figs. 5, 6), Ac. The size of the
species of this genus varies from 6'" — i" ; their synonymy is stUl very
nncertain and confused.
The thin dorsal shield or the shell probably senres for respiration, as in
Apu8, Compare on the stmcture of this genus H. Fbbt u. Lxugkabt,
JBeUrdffe eur Kenaitniee mrbeUoeer Thiere, Braunschweig, 1847, 4to, s.
no— 130.
Family of uncertain position. Oumaoea Kboeter.
Cuma Edw.
Compare Edwabdb Ann, dee Se, not. zm. 1848, pp. 194—- 296.
Edwabds, at a later period {ffiet. not, dee Cruei, m. p. 553) announced
the opinion that this genns has no founda*km» and that the little animal
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661 CLASS X.
obfenred by him might fbbUj be the brm of aome decapod. KBOmi
{Tidtihr. ni. 1841^ pp. 503—534) hss refuted this, hftTin|^ ibond other
species of this form, uid females with eggs in a sac beneath the abdamau
Cfuma is thus an adult animaL It has no eyes, almost the fbnn of a
long-tailed decapod, but fiye thoracic rings are not ooTered by the sheD.
The superior antenns are short, the inferior in the fiemale Ytry short and
rudimentary, in the male long. The feet haye no shears. These anioBsb
resemble the Caridina, but perhaps better still might they be rc&fred to
the stomapods. GrOODSiB and Kboxteb hftve adopted some new genera
in this small fiunily: Leueon Kboitxb, Bodolria Goodbib and Alatna
GooDsm, which last genos, howeyer, Kb(»tkb characteriaes dilfiratly.
See hereon Tid$8krift, iVy Maekke, n. 1846, pp. 133—111.
Obdeb X. Decapoda.
Two compoimd eyes, placed on a petiole moveable, biarticulate.
AntennsB four. Carapace large, covering head, thorax and anterior
abdomen. Branchiae adhering to the base of the posterior maxilli-
form feet and of the feet of trunk, pyramidal, covered by the sides
of the shield. Mostly three pairs of maxilliform feet, in a few only
two pairs. Feet of trunk not changed, almost always in five pairs,
in some six. Mandibles almost always furnished with a palp.
Most crustaceans of this order have a very hard covering, and to it^
with the exception of the Limtdiy the largest spe<aes belong : bat
there are many small ones alsa It is the most numerous and the
most important to man on account of the quantity of edible speoBB.
Section I. Macraura. Post-abdomen large, often longer than
cephalothorax. Penultimate segment with lateral appendages form-
ing with the last segment a terminal pinna of the tail, mostly
flabelliform.
Antennae long, the middle exsert, terminated by two or three
setaceous filaments. Vulvae situated in the first joint of the third
pair of feet.
The third pair of foot-jaws are elongate, whilst on the contrary
they are broad in the following division and cover the oral apparatus.
The caudal feet are mostly more developed than in the following
division, and serve for swimming. The shell is not so thick as in
the BrachyxMra and at the same time less hard
Family XXVII. Caridina. Middle antennae inserted mostly
above the lateral, more rarely in the same horizontal plane as lateral.
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CRUSTACEA. 665
Peduncle of lateral antennse entirely corcred by a large scale.
Body compressed, with post-abdomen incurred. Integuments thin,
almost membranous.
Acetea Edw. Maxilliform feet in two pairs. Only four pairs of
feet of trunk, the last two pairs being deficient; all filiform, long,
none didactylous.
Sp. Ac€te$ indicut Edwabdb, Ann. dea Se. noL zn. 1830, p. 350, PL xi.
fig. I.
Serffestes Edw. Maxilliform feet in two pairs. Feet of trunk
in six pairs, filiform, slender, last pair yeiy small, none didactylous.
Sp. 8erg. oOanHcua MiLNi Edwabds, Ann, da Se, ma. xix. PI. x. fig. i.
PencBua Fabr. Feet slender, furnished with a palpiform lamel-
lose appendage at the base; three anterior pairs armed with a small
didactylous chela. Lateral antennas yeiy long. Shell carinate.
Sp. Peiugus nUcaiui, Pakenwn mdeaiut Oliy., Edw. ffiti, not, dea OruH,
PI. 25, fig.i ;— Pemna tr%auicaiu9 Lxaoh, Desmab. Oru$t. PL 39, tg, 3 ^-~
Pencev* adifenu, Cancer aetifenu L., Seba l^eaaur, m. Tab. 17, fig. 9
(copied in Encyd. nUth., Ortut, PI. igj, ^g, 4) &c.
Add genera: Sicyonia Edw., Euphema Edw., Stenopua Latr.,
Ariiteus Duvkknot.
^pJiyra Eoux.
Pasiphasa Sationt. (In these and the following genus only two
anterior pairs of feet are didactylous.)
Oplophorus Edw.
Pal(jem(m Dald., Fabr. Middle antennae superior, with three
setflB. Shell carinate in the middle, keel produced anteriorly into
a recurved serrated rostrum. Feet of trunk in fire pairs, without
palp or lateral appendage ; two anterior pairs didactylous.
Sp. Pakmon aquiUa Fabr., Cancer aquUla L., Bastes Naiuurh. UUapann,
n. Tah. ra. fig. 5, Gu^bin leonogr,, CruaL PL 22, fig. i ; the prawn
(aleurhrab, creffette, aaUeocque) ftc. A fonil species, from the lithogmphio
stone^ Palamon apimpea, of which ^Bajbb (Oryelogr. Norica, 1758, Tab.
vm. Svppl. fig. 9) and Waloh and Ejtobb haye given figures, the last of
which is copied by Debhabbst CruaL foaaUea, Tab. xi. ig. 4, appears to
stand between Pakanon and SergeaUa and has long spines on the fore feet.
Add genera : PcmdaluB Leach, Lysmata Bisso.
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666 CLASS X.
Htppolyte Leach. Middle anteime siqierior, with two sete, the
external seta excavated beneath, external antennie with a lamel-
lose appendage elongate, obtoBe, and a long mnltiarticalate ^ta.
Shell carinate, supplied anteriorlj with a serrated frontal rostrom.
Third pair of maxilliform feet narrow, mostly furnished with a
pilose palp at the base. Fire pairs of feet, the two anterior didac-
lylous. Abdomen gibbous, with third segment large, fourth in-
flected beneath, and incurred.
Sp. HippclyU Sowerh^ Lsach, Dbsmab. Onut. PL 39, fig. i, Ac. Gompare
on this gwofl H. Kbostxk in KonffeL IkmtU VidenHr. Stbk. nofwr. eg
math. Afh, ix. 1849, pp. 109 — ^360.
Bhynehocmetea Edw.
GncOhophyllutn Latr.
Alpheua Fabr.
PonUmia Tjltil
Sp. PonUmia tjprrketM La9B., Alpheut piimopk^lax OtTO, Nov, Act. Acad,
Oce$, Leop, Carol, zn. 1818, Tab. xu. figs, i, 9.
Add genera : Athanaa Leach, NiJIca Kisso, Atya Leach and some
others, on which see Milzte Edwabds, ffisL ncU. des CrutL n.
pp. 347—366.
Orangon Fabr. Four antenna inserted almost in the same
plane, dilated at the base, terminated bj a double filament. Shell
depressed, with a yery short anterior process. Five pairs of feet,
first pair thicker than the rest, subdidactjlous, with internal finger
very short, immoveable.
Sp. Orangon vulgaria Fabb., Oaneer erangon It., Bobsbl Ins. m. Tth. 63,
Babteb Natuurk. UUtp. n. Tftb. m. figs, j — 4, Th. Bell BriL CnuL^
p. 156. The Shrimp, erepette ; attains a length of fdllj a" ; the second pur
of feet iB about aa long as the third ; this species is Yery oommon on our
shores, and weU known on account of the agreeable taste of its fiesh. In
some other species the second pair of feet is very short. Theij form the
genus Egeon Riaso, PoniophiUu Lsaoh.
Add genera : Sabinea Owen, Argia Kboeter.
Compare Kboitxb Tidsshr. iv. 1849, pp. 26^, 168.
^ A HiMory of BrkUK Onutaeea, by Th. Bbll. London, 1844 to 1848, 8vo. (only
Ti. parts have yet appeared ; this work contains yery beaiitiftil wood-cots).
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CBUSTACEA. 667
Family XXVIII. Astadna. Four antennse inserted in nearly
the same transverse line, unequal, the middle with two setae, shorter.
Anterior feet chelate. Shell mostly hard, calcareous. Branchisd
fasciculate, composed of cylindrical filaments.
Astcums Gronoy., Fabb. Lamellar appendage, dentiibrm or
hastate, mostly small, at the base of lateral antennae. Three first
pairs of feet didactylous, first pair large, thick. Caudal pinna with
lateral lamellae bipartite transversely.
Nephfrop8 Leach. Scale at the base of external antennse longer
than the peduncle. Eyes large, reniform. Frontal rostrum elongate,
denticulate on both aides.
Sp. Nephropi norwegieui Iaacb, Oaneer nonoegicui h,, DiSM. Onut, PL 37,
^. I, Gir^Biir Iconogr,, OnuL PI. 19, fig. i.
Astacus Lbaoh {Astacua and HoTnavrua Edw.).
a) Last segment of thorax mobile. Sqnama of external antenns
mobile, acuminate. A$tacu»,
AMaeua JkmaHlU Fabb., Chmeer oMaem L., BaexL Ins, m. Stippl,
Tab. 54—^1, Guiaiir Iconogr,, Onui, PL 19, fig. 2, HsBBiOH-SoHAFras
DeittecAZ. Int. Heft 186, Tab. 13 ; the cray-fith, rwer-eray; attains a length
of 3" to 4". Of this species, which is dispersed almost over the whole of
Europe, there are some Tarieties which KooH considers to be species.
HXBBIOH-SOHAFVES L L Heft 140, 186.
h) Last segment of thorax cohering with penultimate by an immoYe-
able connexion. Scale of external antenns small, dentiform. QouMMru*
OT ffomartu.
Sp. Attaew marinus, Cancer ffammanu L., Basteb Natwvrk. UiUp. n.
Tab. I. DiBXAB. Onut, PI. 41, ^. i, the Muter, eea-erap; on the coast of
Norway eq>eciaUy.
Compare on the different species of this genus Ebiohsov Archiv, 1S46,
pp. 86—103, 375—377.
Eryon Desxab. (Fossil genus ; see Diskab. BitL not. de» Onut.
foBiUet, p. 128.)
Sp. Bryan CvmerU Disk., Bajsbi Oryeiogr, nor. Tab. 8, Suppl, fig. i,
DiaXABBOT Onut. foes. PL 10, fig. 3, in the lithographio stone ; compare
Bbovn Lelh. ffeogn. pp. 473, 474.
Thalasaina Latb. Scale at the base of external antennae mostly
none, in a few veiy small. Fonr anterior feet, sometimes only
two, didactylous. Lateral lamellae of caudal pinna undivided.
Posterior part of abdomen or tail elongate.
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668 CLASS X.
Compare Lxaoh On the ckaraeUn nf <&« peniM Hifclaesinft of liATBXQU,
ZwA, Hue. in. 1817, pp. 17, 18.
t Aeeenofry hranckial offpmdaget Cj^ndrieal, diekoiomous, adkermg to
the abdominal feet.
CaUinidea Edw.
Callinisea Edw., Isea Gu^bin.
Compare Gu^BiN IktcripHon d'un noupeau gmre de enutaee wtacnmrc
Ann. de la 8oe. entom. de France, i. 1833, pp. 295 — 300.
1 1 EiBtemal Inxtnehia none,
a) Four anterior feet didactyUme,
ThaloBsina Latb. (in part), Leach, Axius Lkach, CMianasia
Leach.
Sp. Thalateina eeorpionidet, Cancer anomalut Hibbst, Lxaoh Zoeil. Mite.
Tab. 130, Desxab. CrvM, PI. 35, fig. i, Gu&iK leonogr., Oruet. PL 18,
fig. 4 ; from the coast of Chili ; — OalUanaeea eubterranea Lbach, Dxsxab.
Cfrtut. PI. 36, fig. 7, GuiRlN Iconogr., Cruet. PL 19, ^. 4, on aome parti
of the English coast and in the Mediterranean. According to Edwabds
CaUian. laUcauda Otto, Nov. Act. Acad. Ccee. Leop. Car. Tom. Jjr.
Tab. a I, fig. 3, is not specifically different from it.
b) Two anterior feet alone didadyloue.
Genera : Gebia Leach and Glaucothoe Edw. {Prophylax Latb. 9)
ilTo^— -Genus MegcUopa Leach, Megalops Edw., placed here by
Latbeille between GcUcUhea and the preceding genera, most be
suppressed ; from the observations of Thompson and Rathke it
belongs to the earlier forms of brachjurous Decapods. The same
conclusion is to be formed respecting Monol^oia Say.
Thoicfson Phil. Trane. 1835, PP* 358—361, PL ▼. ; Bathxx BeUr. c
Verffl. Anat. 1841, p. 46.
Oahihea Fabr. Middle antenna short, geniculate, terminated
at the apex by two setae ; external long, with elongate, multiarticu-
late seta, without a scale at the base. First pair of feet large,
chelate; fifth pair of feet slender, recurred. Shell ovate, depressed.
Galathea and Grimothea Leach.
Sp. Oalathea ttrigoea Fabb., Cancer etrigotut L., Dbsm. Cnut. PL 33, fig. '/
GuibiN leonogr., Cruet. PL 17, fig. 3.
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CRUSTACEA. 669
JEgha Leach.
Sp. JBgUa Icevit, Qolathea loms Latb., Encyel., ffiti, not, CruH, PI. 308,
fig. 9, DSSMAB. CruM, PL 33, fig. 1.
ParceUana Lam. Middle antenna very short, hidden beneath
frons, with double seta, external long, terminated by a mnltiar-
ticulate seta, and without scale at the base. First pair of feet didac-
tjlous, large, depressed, with chela large ; fifth pair of feet slender,
recurved. Shell depressed, suborbicular. Tail reflected under
thorax.
Sp. PorceUana longieomU Latb., Cancer UmgieomUTssvAST, £neyel. nUth,,
Ins. et Orrui, PI. 975, fig. 3, Bill Brit. Crual, PL T93 ; in the Baltic,
also on onr ooasts. This genus fonnfl the tranaition to the Decapoda
hraehywra. Lbaoh distinguishes this and some other species of PoreeUana
by the name PUidia, as a separate genus.
Family XXIX. Lorioaia {Locustce Latr.). Four antennas
inserted almost in the same transverse line, middle filiform, bisetose
at the apex, external without scale at the base. Feet monodacty-
lous, subequal. Sternum broad posteriorly. Shell hard.
Paltnurus Dald«, Fabr. External antennae setaceous, yery
long, thick at the base, aculeate. Shell muricate. Eyes large^
approximate.
To this genus belong different species of large crustaceans, which in-
habit rocky places in various seas. In the Mediterranean is found Pali*
nwrua vtdgaru Latb., Cancer homarfu L. (excL Synon.), Dbshab. Onui,
PL 31 ; the Greeks named this animal KopafiSs, the Romans locutta, from
which the French name langotute is derived. The flesh and also the eggs
of these crays are renowned as delicacies. Amongst the exotic species,
there are many of which the internal antenna have two long filaments, as
PaUrwriu gtdtahu Latb., Encyd.y Ins. et Crust. PL 315, from the West
Indies, Pal. japmicus Y. Siebold, Db Haan Fa/un. japon., Crust. Tab.
41, 4«, &c
ScyUanis Fabr. Lateral antennae without filament, peduncles
with joints foliaceous, plane, broad. Eyes mostly very remote.
Fifth pair of feet in females subchelate, with two small fingers.
Sp. ScyUarus arctus Fabb., Cancer arctus L. (ezd. Synon.), Sulzbb Gfeseh.
der Ins. Tab. 31, fig. Z, Cuv. jR. Ani., id. ill., Crust. PL 45, fig. i ; in the
Mediterranean. In ScyUarus oricntalis Fabb., the eyes stand still further
apart, entirely at the margin of the shell, which is very broad forwards
and becomes narrower backwards. Rumph. Amb. Raritk. Tab. n. fig. D,
Dbshab. Cnalt. PL 31, fig. i. This species forms the genus Thmus
Lbach. In some species from foreign seas the eyes are placed more in the
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670 CLASS X.
middle of the sheU, and thia is very broad, and indaed deeply on eacii ode.
They form the geniu IhacuM Lsaoh ; to it belonga ScjfUanu aMtanHaa
Fabu,, Bumph. L L fig. o.
Familj XXX. Anamura {Ariomala Latr.). Four antenns
inserted iiearlj in the same transverse row, or middle superior;
external antennae without scale at the base. Eyes placed on a
petiole, Tery often elongate. Two or firar posterior feet less by far
than the rest. Abdominal feet imperfect, small, deficient in some
of the segments. Lateral appendages of penultimate segment of
abdomen reflected at the sides, never composing a flabelliform
pinna.
Phalanx I. Pagurina {Poffuridce Bell). First pair of feet
didactylous, chelate. Appendages of penultimate segment of abdo-
men short, composed mostly of two parts, incurved, inserted on
a short peduncle, not foliaceous. Integuments of abdomen often
soft, membranous.
PaguTus Dald., Fabr. Abdomen contorted, membranous, sup-
plied with thin calcareous laminae scattered in the back. Foorth
and fifth pairs of feet very short, subequal.
Fagurtu Latr. Middle antennsB short, with two very short
terminal sets.
Sp. Pagwrut BemkardHi Fabb., Cancer BemKardug L., Bwamxesdax
Bifbd d«r Nat. Tab. xi. Dbbhab. OmT. PL 30, fig. 9, the hermit'cnb;
Uyet in different torbinated BheUa, when fiiU-grown in the shell of Bttc-
cinmm ufuletfttm.
iVbte.—Add genua CanetUvM Edw.
Compare Milnb Edwabdb Ob9ervalums tur let Pagwree, &c.. Awn. da
Sc, ma. ie S^e n. 1836, Zool. pp. 157—188, PL 13, 14 ; ejnad. i^
^[uetquee notsveUet eephes du genre Pagvire, Ann. dee Sc. mU. siteie 86ne x.
1848, Zool. pp. 59—64.
Goonolnta Latb. Middle antennae long, with peduncle elongate
and two unequal setSB, one of them long.
Sp. Cfamchita J>i4)gene9 Latb., EneycL m&h., Orud. PL 284, figs. 3, 3i
Edwabdb ffiel. nai. dee Onut. PL 11, figs. 11 — 13.
Birgus Leach. Abdomen broad, covered above by a tabulate
calcareous crust, membranous below. Fourth and fifth pairs of
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CRUSTACEA. 671
feet chelate, unequal, the last much smaller. Middle antennae
elongate, with two unequal setae at the apex.
Sp. Sirgui UUro, Pagwnu UOro Fabb., Dsbmab. Cfruit, PL 30, fig. 3.
Phalanx 11. Htppaidea. First pair of feet monodactjlous or
subchelate. Four or six following feet with last joint pinniform,
lamellose. Fifth pair of feet short, thin, incurved. Penultimate
segment of tail with two appendages on each side lamellose, oval,
ciliate. Shell ohlong.
Hippa Fabb. (exclusive of some species). External antennae
very long, terminated hy a multiarticulate ciliated seta. First pair
of feet with terminal joint lamelliform.
Sp. Hippa emeriia, Cancer emerihu h,, DiaCAB. Cfruti. PL 39, ^, ^, on the
coast of Bnzi].
Remipea Latb. Middle antennae bisetose at the apex, longer
than external. First pair of feet long, with last joint acuminate.
Sp. Bemipa tettudinariui Latb., Hippa adactyla Fabb., Cut. K AnLid, i.
PL la, fig. 2, Besxab. Onut, PL 29, fig. i, from the ooaat of New
Holland.
AUninea Dald., Fabr. (in part). Middle antennaa longer than
external, terminated hj a single seta, very long. Anterior feet sub-
chelate, with moveable finger incurved. Shell oblong, with anterior
margin broader, almost straight.
Sp. Albunea tymnuta Fabb., Bbsmab. Cnut, PL 39, fig. 5, Gu^bik leonogr.,
OniM, PL 15, fig. I, from the Indian Ocean.
Section II. Decapoda brachyura {KleiatagnathaYKBR.). Tail
or posterior part of abdomen small, reflected forwards, almost
always received in a furrow of sternum, not terminated by a pinna,
with appendages filiform, in females oviferous, not natatory. Third
pair of maxilliform feet broad, covering the cibarious organs. Vulvae
mostly situated in sternum, between the bases of third pair of feet.
Antenn» short ; middle received in a farrow under the anterior
margin of shell in most, terminated by a double subulate seta.
A. Two or four last feet inserted towards the back, not in the
same plane with anterior feet.
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672 CLASS X.
Familj XXXI. Xaicpoda Latb.
t VultXB at the base cf third pair qfjeet.
Banina JjAM. (Species of Albunea Fabr.). Anteiui» short.
Shell oblong, oval or obtrigonaL ChelsB compressed. Tail short,
extended.
Sp. JZohmm daUata Ljltb., Cancer rammwM L., Rumfh. AwA. Itarileiik.
Tab. vii. ^. T, ▼, GuiBiH Jamogr., CrutL PL 14, fig. 3, D» Haam
Ormd,, Fmm. Jap. Tah, xxxiT. zxxt. figs. 1—4 ; from tlie Uand of
Hamitiiii to Ji^Mn.
NcU. — ^Add genen: Natopui Di Haah {Noiopui donipet from wliich
SaninMm Edw. fcarody diffen), RaniUa £dw. and Lyreidut Di Haas.
Hamola Leach. External antennae terminated by a long seta,
middle not received in a sulcus of firons. Shell oblongo-quadrate,
spinose. Tail inflected beneath. Feet elongate ; fifth pair of feet
dorsal, subcheliform.
1^. Homola tphUfrom LiaOH, Dismab. Onui, PL 17, fig. i, from the Hedi-
ternuiean. Thu genus, like the preoeding, liM allumoe with Hippo.
Lithodea Lats. Shell obcordate, anteriorly rostrate, aculeate
and tuberculate. External antennas terminated by a long seta.
Tail inflected beneath. Second, third and fourth pairs of feet long,
spinose ; fifth pair of feet minute, almost hidden under the posterior
margin of shell, cylindrical, with apex didactylous, obtuse.
8p. JAAodei aretiea Lax., Cancer ffu^ (excL Syn.) PoNTOFpmAjr Narffa
Naiurl, Hid. n. p. 186 (with fig. »t p. 176), HnBST KnAhm, Tih. xr.,
Dbucab. CruM. PL 45. This species, living in the North Sea, is named
Trcldkrahbe^ DeviTt Crab. The feet measure foUy 4 feet, although the
shell 18 not laiger than 5 inohee.
Droma Dald., Fabr. Internal antennas received in sulcus of
frons. Shell orbicular, very convex. Feet short or moderate ; four
or more rarely {Dynomene Latr.) two last dorsal, subcheliform at
the apex.
Sp. Dr^mia Jtumpkii, Caneer DronUa h., Rumph. Amb. RarUeUk. Tab. zi.
fig. I ; — Drfmianodipet'LA.TK., Guiaor leonogr., Crud. PL 14, fig. 66, &c
lliese eruBtaoeans leize with their hind feet, which are placed on the back,
different objects met with in the lea, Alcffonidia, aponges, &c., and fix
them aa a covering on the shelL
tt Female orifices of generation in the middle of sternum,
between the bases of third pair offset.
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CRUSTACEA. 673
Dorippe Dald., Fabr. Shell depressed, anteriorly narrower,
truncated. Second and third pairs of feet very long,, with last
joint long, sutulate ; fourth and fifth pairs dorsal, short, sutcheli-
form. Tail inflected downwards.
Sp. Dorippe ktnata 'Boao, Cancer lanattu L., Desicab. Onut. PL 17, fig. 7 ;
from the Mediterranean.
B. Feet all in the same horizontal plane inserted at the inferior
surface of sternum. Tail always inflected downwards.
t Area of mouth (space bounded by the third pair of maxilli-
form feet) triangular, anteriorly narrower, often produced almost as
far as the anterior margin of shell. Third pair of maxilliform feet
slender or narrowed anteriorly, triangular.
Family XXXII. Oxystomata Edw. (in part).
Corysiea Latr. External antennsB setaceous, ciliated, very long.
Shell oblong, oval. First pair of feet cheliform, the rest with last
joint subulate, acute, elongate.
Sp. Ccryttei denkUtu, AUmnea deniata Fabb., Dxbmab. Cfntd. PI. 3, fig. 7,
Bell Brit. Orutt, p. 149, buries unclar the sand, sending out the antennn
alone above it; the males haye shears of twice the length of the body. This
species is found on the English and French coasts.
AteUcyclus Leach.
Add genus Hiia LSAOH and some others, on which see Milkx Edwabds
HkL not, dee Orutt. n. pp. 141— 151.
Leucosta Fabr., Latr. External antennae very small, internal
concealed in little pits. Eyes minute. Shell solid, mostly rotundato-
ovate* Branchial cavity closed below, no fissure at the bases of
maxilliform feet. Tail with four or three segments.
Several sub-genera proposed by Leach : Iphis, I^ureia, Peraephona,
Ixa, FhUyra, Ebcdia, Myra, Ilia, Add Oreophorus Bxnspp.
Note. — Here also Bellia Edw., a new genus, seems to be referrible. See
Ann. dee Se. not., si^me S^rie, Tom. ix. 1848, Zodog. p. 191.
Sp. Leueoeia crcmolarie Fabb., Dbsmab. Ortut. PI. 97, fig. 1.
HepcUus Latr.
Matuta Fabr. Antennae short, external minute. Two anterior
feet short, with chela concave internally, crested above ; remaining
VOL. I. 43
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674 CLASS X.
feet with last joint lamellose, lanceolate or oval. Shell snborbicolar,
with anterior margin dentate, with a strong spine on each side pro-
duced above the second pair of feet.
Sp. Matvia victor Fabb., Rumph. Amb. BarUeUk. T»b. TO. 8» Dismab.
Cnut, PL 7, fig. 5 ; in the Indimn Ooean, also in the Bed Sea.
Orithyia Dald., Fabr, Shell mnricate oval, truncated anteriorly.
Fifth pair of feet terminated by an oval lamina.
Sp. Orylkia fnomiUaini Fabb., Dbsmab. Cry/d, PI. 19, fig. i, GulEBni letmcgr.,
Onut, PL I, fig. 3 ; from the Indian Ocean and the coast of China.
MuTtia Leach.
PUUymera £dw.
Calappa Dald,, Fabr. External antennsB short, with basal
joint broad. Shell short, convex, broader posteriorly, covering
with its margin produced the contracted feet First pair of feet
with cheliB large, compressed, crested above, with inner concave
side excised at the outer margin of shell. Remaining eight feet
with last joint styliform,
Sp. Oalafpa granulata Fabb., Caneer granulaUu L. (exd. Synon.), Hibbst
Krahben i. Tab. 11, figs. 75, 76, Cuv. R, AtU., id. itt., Oruti, PL 38, fig. i,
from the Mediterranean i-^Cdlappa tubercuUOa Fabb., Bbsxab. Crud.
Fl. 10, fig. I ; Bed &e^ Indian Ocean, kc
iVbfo.— Add 8ub-genu8 Oafnara Db Haan. Sp. Calappa fonUcata,
Caneer calappa L.
tt Area of mouth quadrate, anteriorly remote from the margin
of shell. Maxilliform feet broad.
Family XXXIIl. Majacea De Haan, Oxyrhyncha Edw.
Space between the origin of middle antennae and the area of mouth
large, scarcely broader than long. Shell triangular or subovate,
anteriorly narrowed, mostly running into a frontal rostrum. Feet
mostly elongate, never natatory.
Inachtis Fabr. (in part). Feet slender, very long, elongate;
second pair very often longer than first. Third joint of maxilliform
feet obtrigonal or obovate, sustaining the fourth on its apex.
Leptopodia Leach. Rostrum very long. Peduncle of eyes
short.
I
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CRUSTACEA. 675
Sp. Inaeku9 tagittariut Fabb., Dbbmab. Crutt, PI. i6, fig. i, Gu^tn
lamogr., Orust, Fl. 1 1, fig. 4 ; finom the West Indies.
Nole.-—QeaauB PaOolm Lsaoh to be obliterated; comp. Di ^Ian
Fauna Japan,, Crrut. p. 89.
LaitreiUia Roux. (Is this its place ?)
Stenorhyrichua Lak.
Inaehu8 Leach. Peduncles of eyes capable of concealment back-
wards in a pit. Rostrum short. First pair of feet in males longer
than shell, in females short
Sp. Inachut Scorpio Fabb., Cancer Doraettensis Penn., Debmab. Crust,
PL 34, fig. I, Bell Brit. Cnut. p. 13, in the North Sea and the Atlantic
ooean. — Inaehut leplochirut Lsaoh, Bell L 1. p. 18.
Macrochira Db Ha an. Shell cordate, posteriorly broad, tuber-
culate and spinose, with rostrum bifid, and horns diverging. Chelse
cylindiical, in adult males surpassing in length the second feet.
Tail with seven segments in both sexes.
Sp. Inackus Kcempferi Db Haan, Faun, Japan,, Cnut, Tab. 35 — 48, a
very interesting crustacean which, according to the characters of the
oral organs belongs to Inacku8, but by its size and by the long fore-feet
or shears in the male differs from it. These may attain the length of
4 feet^ and the thickness of the thigh-bone of a man, as was already
figured by ELsMFFSB, Bctcknjv. van Japan, Amsterd. 1733, fol. p. 100,
PL XIV, A.
Campo8cia Latb., Oncinoptis De Haan, Ev/rypodiua Guj^sin,
AehcBus Leach, Microrhynckiu Belk Here also De Haan refers
genus ffalinvus Latr., resembling the Maja and FiacB.
Sp. ffalimua ariea Latb., GtJ^BiN Iconogr,, Crust, PL 9, fig. 1,
Egeria Latr. Feet slender, very long; second pair longer
than first. Third joint of maxilliform feet quadrate, receiving the
fourth joint in the upper margin at the internal angle.
* ShdU triangular, ttith rostrum two-homed,
Ev/rypodiu8 Guerin.
** SheU orbicular, vnik rostrum short, narrow, the apex entire or emar-
ginaJte.
Dodea Leach, Egeria Latr., Libinia Leach.
Sp. Egeria araehnddes Latb., Buhph. AmiK RarUeiik, Tab. vni. fig. 4 ;
East Indies.
Maja Lam. (exclusive of species). Feet moderate, first and
second pairs longer. Third joint of maxilliform feet of third pair
43-2
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676 CLASS X.
quadrate, receiving the fourth joint at the internal angle of the
superior margin. External antennse with seta pretty long, sub-
ulate, with elongated joints, first joint large, grown to the shell, with
the external margin shutting in the orbit beneath. Shell mostly
longitudinally triangular, or oval, spinose, with rostrum bicomute.
Abdomen in most composed of seven segments.
Piaa Leach (and Lissa ejusd). Add sub-genera : iTyow Leach,
Mydppa Leach, Leucippa Edw., Pericera Lata., Herhtiia Edw.
Sp. Hya$ araneut Lkaoh, Cancer araneHi L., Guy. R Ani., itL HL, CnuL
PI- 33* fig- «i Bill BrU. Cruat, p. 31.
Maja Lax. (exclusive of several species). Add sub-geneia:
AcarUhonyx Latb., Chorintu Leach, Mithra Leach and some
others. Comp. De Haak 1. L pp. 81 — 83.
Sp. Maja iquinado Latb., Cwneer iquinado Hkbbst, Dbsxab. OnuL PL 31,
Bill Cntd. p. 39 ; on the S. and W. coastB of England, in the Meditei^
ranean, &o.
Parthenope Fabr. First pair of feet very long, geniculate at
the chela, remaining feet moderate: (Third joint of maadlliform
feet as in Maja.) Basal joint of external antenn® not concrete
with shell, not closing the orbit. Shell mostly triangular, often
broader than long, tuberculate. Rostrum short.
P(Mihenope Fabb., Leach (and Larabiua Leach).
Sp. PartheMpe korrida, Caneer horridus li., Ruxph. AwUt. BariUiik. Tab.
n., Dbsmab. Cntae. PI. 40, fig. i ; from the Indian Ocean.
(Ethra Leach, Lam.
Cryptopodux, Edw,
NqU. — ^Theie sub-genera, acarcely distinct, diffsr from Partkmop* in tbe
feet, as in Oalappaf concealed under the sheik
Family XXXIV. Cancrina {AroucUa and Quadrtlatera Latb.,
Catometapa and Cyclomelapa Edw.). Space between the origin
of middle antennae and area of mouth short, transverse, much
broader than long. Shell never running anteriorly into a firontal
rostrum.
A. Fourth Joint of nuzxiUtJormJeei placed upon the middle ogpex
or the external angle of third joint.
Shell mostly svhquadrate or trapez^orm, with frona declminff or
perpendicularly injlected.
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CRUSTACEA. 677
Pinnoteres Latr. Shell orticulate. Peduncles of eyes shgrt.
Chelss short, thick.
Sp. PinnoUrei pwtm Lata, (and Pttm. mytilorum ejosd.), Ccmeer piswn L.,
Fabb., Babteb Natuurk. UUsp. n. Tab. lY. figs, i, 3, CuY.RArU., id. ill.,
OruH, PL 19, fig. I, Bkll Brii. OrutL p. lai; in muflsels. Another
species, PinnoUret vetenttn 'Bo^a, lives in Pinna and was supposed by the
ancients to warn this bivalye against impending danger (see CIOXBO de
Finib. mal, et hon. in. Cap. 39, the mid.).
The females are larger, have a broader shell and especially a much
broader tail than the maleSk
Sub-genera: Xanthatia, Pinniooa Whitb, Ann, of not. Eitt. xviu.
pp. 176, 177.
Orapsus Lam. Shell depressed, subquadrate, with firons broad
and lateral margins almost straight, i&jes placed at the lateral
angles of shell upon short thick peduncles. Middle antennas hori-
zontal, concealed under the teflected irons. First pair of feet short.
Sp. Qraptua pidua Latb., Cancer grapam L., Besicab. Cru$t. PI. t6, fig. i,
Cut. R, Ani.f id, ill., Cruat. Fl. aa 'y—Qrapavfi vaariegaUu Latb., Cane,
varieff. Fabb., GuisiN Jeonogr., Onui, PI. 6, fig. i.
Add genera: Sesarma Edw. {Pachyaoma De Haan), Tcuruna
Edw. {Trichopu8 De Haan), FaetLdograpsiu Edw. and some others,
which must be here omitted.
Oecardnvs Leach. Shell cordate, anteriorly broader, turgid,
posteriorly truncated. Peduncles of eyes short, received in rounded
furrows.
Sp. Qecwmnut ruricola. Cancer ruricola L., Dxsmab. OnuL PI. 17, fig. 2,
ftc ; from S. America.
This genus contains the so-called land-crabs, that live in woods. Some
betake themselves at a certain time of the year to the sea to lay their eggs,
and undertake the journey in large troops, as travellers relate, in a straight
line, finom which they cannot be turned by any opposition.
Flagusia Latb. Frons excised on both sides for middle an-
tennae denuded above, porrect. Shell broad, narrowed anteriorly.
First pair of feet short.
Sp. Plagutia clavimana, Dbbxab. Cnut. PL 14, fig. 2, 9^,
Ocypode Fabr. Shell quadrate or trapeziform, anteriorly
broader. Middle antennse short, with two very small seta? ; lateral
antennae small. Eyes placed on long peduncles. Chela in males
often very large on one side, far surpassing the chela of the other
side.
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678 CLASS X.
Gdatimus Latr. Pedcmcles of ejes slender, cylindricaly with
ejes termlnaL Shell broader than long, trapesiform.
Sp. OdoMimmi vocamt. Cancer toeamg L., Di Gkib M^m. p. s. a FHuL de$
In§. m. FL 36, fig. la, friHn the ocMsi of BnzQ. Comp. Limr. Anum.
Aead, ti. p. 414 ; — Gtiatin^ Mariomit DxsM. Cnut, V\. 13, fig. i, &c
Ocypode Latb. Peduncles of eyes acuminate, with a horn pro-
duced beyond the eye& Shell subquadrate.
Sp. Ocypode eeratopktkalma Fabr., Caneer enrtor L., Pallas SpicU. ZaoL
IX. p 83, Tab. T. figs. 7, 8, Dbskas. CruM. FL 12, fig. i ; from the Bed
Sea and the Indiaii Ocean.
Add genera Uca Leach, Carditoma Latb., MixerophthalmusIjATSLy
CleiHosiama De Haak, Hymenosama Leach, MydUris Latb., Doto
Ds Haak, dux CompL De Haak Faun, Japon^y CrusL p. 5,
ppi 24 — 30. Here also belongs genus ffalicarcinus Whitel
B. Fourth joint of nuuciUifyrm fiet placed on the inner angle of
third joint.
Shell mostly arcuate anteriorly, with margin convex, tnmcated
posteriorly.
Telphusa Latb, Shell depressed, smooth, cordate, broad. Ex-
ternal antennse very short, inserted near the peduncles of eyes.
Sp. TelphuM jluriatilis Latb., Cfrabe de rivih^ Olitier, VojfOffe done
V Empire Othamam, FL 30, itg. 1, Dksmab. OnuL PL 15, fig. 3 ; in fresh
water in the south of Italy, Greece, Egypt^ Ac
Here belong also the genera Boecia Edw. and Tridkodtutiflue Latb.
Eriphia Latb. Shell cordate or trapeziform. External antennae
exsert, remote at their insertion from the petioles of eyes.
Sp. JSriphia epinifrtnu Latb., Cancer epinifrone Hebbst, Fabb., Dbbmab.
Cnut. PL 14, fig. 1.
Add genera RuppeUia Edw., {Eudora De Haak), Trapezia Latb.
Cancer Fabr. (species from Cancer L., div, brachyurJ). Shell
broad, anteriorly arcuate, gibbous. Chelae thick. Eight remaining
feet with last joint styliform, not natatory. Third joint of maxilli-
form feet of third pair quadrate.
1^. Cancer poffurut, Platycarcinue pagurue Latb., Edw., Dbsmab. Omtt.
PI. 8, fig. I ; Bell Brit. Cruel, p. 59 ; the common eea-crah ; 6 inches
broad or broader, the shell reddish brown, granularly uneven and on each
side with nine incisures or grooves ; the shears smooth and, as in other
species of this division, black at the point ; this species is edible and
very tasty.
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CRUSTACEA. 679"
Compare on this genus Th. Bill OhaervaHons on the geMU CairciMU &c.
TranaacL oftheZooL Soc, I. 4, 1835, PP» 335— 343» PL 43—47-
Add genera : Oonoplax Leach, PUumrms Leach, Xamiho Leach,
and some others proposed by De Haan.
Poriunua Dald., Fabr. Shell depressed, posteriorly truncated,
mostly transverse, broader than long. Two last feet natatory, with
tarsus depressed, flat.
A. Two posterior fed alone ncBkUory.
dvtdnuB Leach. Tarsus of fifth pair of feet narrow, lanceolate.
Petioles of eyes short
Sp. Ca/rti'MU mcenae, Cancer motnaa h., Bastxb Naiuurk. UiUp, n. Tab. ii.,
BsLL Srit. Cnut, p. 76 ; the $hore-crab : forward between the eyes the
shell has three points, and on each side at the margin five triangular teeth.
This crab is very oonunon on our shores.
Portanus Leach. Tarsus of fifth pair of feet dilated, oval.
Petioles of eyes short.
Add sub-genus ThalaMUa Latb. and genus Lupa Leaogs in part.
Sp. Portunue puber Leaoh, Cancer puber IL,, Dksmab. Cruat, PL 5, ^. i,
Bill. Brit. Crud, p. 90.
Podophthalmus Lah. Tarsus of fifth pair of feet dilated, oval.
Peduncles of eyes cylindrical, very long, produced as feir as the
angles of shell, received in a canal in the margin of shelL Shell
short, transverse, depressed, on both sides at the anterior angle bi-
spinose.
Sp. Podophihalmtu epinoeue Lam., Latb., Portunue vigU Fabb. Suppl,
Enkm. ayet. p. 365, Desmab. CmM, PL 6, fig. i, GuiBiN Iconogr,, Cnut.
PL I, itg, 5 ; from the Indian Ocean.
B. Fovr poire of feet natatory, mth taraua foliaeeona.
PUUonychua Latb. (and Polyhms Leach, and species of genus
Lupa Leach, sub-genus Neplmvaa De Haan).
Sp. PolyUue Henslomi Leaoh, Deskab. Cnut. PI, 7, fig. i, Bell Brit.
Cruet, p. 116.
Lupa pdagica Leaoh, Catieer pdagicue L., Bumfh. Amb. Rariteitk.
Tab. vn. fig. b, Desmab. Cnut. PL 6, fig. 2, De Haan Cnut. Jap, PL
IX. X. ; this beautiful species with smooth shell running into a sharp point
on each side between the second and third pairs of feet, is found in the Bed
Sea, the whole Indian Ocean, and especially on the coast of Japan, where
it is very common, and a fayourite food of the natives.
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ON MOLLUSCS IN GENERAL'.
In what precedes we have treated of the greater part of those
animals that have no internal skeleton or ho vertebrse. There still
remain however for our consideration many that, in the system of
LinncBus, were referred to the class of worms, but which by Cuvier
were, first as a class, afterwards as a larger group constituting a
primary form {typus), collected together and separated from the rest
of the worms*. Since they do in reality form such a group, which
^ Compare on this diyiid<Mi of the Animal Kingdom^ amongst othen :
M. LiBTXB Exerdtatio anatomiea de CoeMeii numime terratribuM et Limaeibus.
Londini, 1694, Svo.
EjuwL jETut. nve Synopns meik4)d. Conehyliorum et Tabtdarum ctiuUomiearum edUio
aUera, Reeenxwi H indieihui aunt 6. Hudisford, Oxonii, 1770, folio.
M. AOAKBOK BiM. nafur. dn SinigaL Coquillagei, Avec 19 fL Paris, 1757, 4to.
J. X. Fou Taltaeea utriuMque SieilitB eorumfue kiMoria H anaUMne, TahuUt an,
illtutrata, PamuB, fol. Tom. I. 1791, Tom. n. 1795, Tom. m. Pan prima potthwna ;
edid, S. Dellb Chujx, Parme, JS26, Pan altera, aueton S. Dbllb Ghiajs, Panua,
1817 (this part remains unfinished).
CuviSB MSmoirti pour iervir d VHUbtire H d FAnatonie dea MoUutjuet, atcc
35 PI. Paris, 1817, 4to.
E. FoRBBS and 8. Hanlit Nat, ffut. of British MoUutca and ikeir SkdU,
London, 1848 — 50, 8vo.
Q. J0HK8TOK An Introd, to Coneht^ogy ; or ElemenU of the natural HitL 0/
MoUutcoue AnimcUt. London, 1850, 8to.
As manuak the following especially may be consulted :
Laxarck Biit. fiat, det Animaux »ant VerUbra, le idit, par Deshatbs et
MiLKE Edwabds, Tom. vi. 1835 ; H. Ducrotat db BLAnrviLLR Manud de Malaeo-
iogie et de Conchologie, i Vol. 8vd, Paris et Strasbourg, 1835—1827, ayec 107 Planchet,
and a. P. DxsHATBS Traiti elSmeniaire de Conchdogie, Paris, 1838 and folL 8to.
(hitherto only 9 numbers have appeared). We shall principally refer to these two
last works, besides Gu^iv's Iconographie, for the figures, and only occasionally quote
more extensive works of plates, as those of Mabtini and Chbmnitz, Kibkbr, ftc.
' See p. 33 and 208. To the works of Cuyibb referred to in the last of these
pages, there ought to have been added one that was printed separately from the
Decade philoeophique, a journal with which I am not further acquainted, and which
now lies before me : it is entitled, Mimoire tur la etrueiure interne et exteme et eur let
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ON MOLLUSCS IN GENERAL. 681
may be compared with that of the vertebrate animals, to which four
classes belong, it is not inexpedient, before we pass to the consi-
deration of the different classes which have here, with more or less
propriety, been adopted, to indicate the common characters which
distinguish the molluscs from the rest of the worms.
The body of these animals is covered by a skin, soft and con-
stantly moist, to which the muscles are attached, and in which or on
which a calcareous secretion is usually effected. The external in-
tegument, which indeed has also been named mantle (although
properly a free production on the dorsal surface ought alone to bear
that name), encloses as well the intestines as the nervous system
also. The central parts of this nervous system consist of ganglia,
which either form a ring round the oesophagus or lie more dis-
persedly, but not behind each other in a row on the abdominal sur-
face, as is the case in the insects. The molluscs ordinarily present a
much less similarity of the right and left half of the body than the
articulate animals already reviewed by us, or the vertebral animals
to be considered in the sequel. Many have no head distinct from
the rest of the body. Th« organs of sense are on the whole slightly
developed. In the most composite molluscs however, in the S^ice
and other Cephalopods, there are found not only two highly de-
veloped eyes, but also rudiments of auditory organs. The move-
ments are on the whole creeping and slow. Some, that live in
water, are immoveably attached to different objects. Many headless
bivalve molluscs have indeed a springing motion, yet this is in a far
less degree successive than in articulate animals.
The inferior degree of development of the organs of animal life
is the cause that many writers of the present day, as Linnjeus
formerly, still place the molluscs lower than the insects in the
arrangement of the animal kingdom.
More perfect than the organs of animal life are those of the vege-
tative, those for secretion, nutrition, and propagation. The respi-
ratory organs are usually gills. In most molluscs a heart is
present, which receives the arterial blood from the organs of respi-
ration, and distributes it by arterial tubes to the different parts of
affiniU$ des animaux auxqueli on a donvA le nom de Yen. From this memoir read in
1795 before the Soe, d'Hist, not. of Paris, it appears that even thus early the class of
the molluscs was distinguished by CuviER and defined as in his later works.
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682 ON MOLLUSCS
the bodj. Capillaries are wanting, and the veins are replaced hy
sinuses, which have no proper walls, bnt are excavated in the
different parts of the body. The blood of moUnscs is commonlj
white or whitish blue. Some moUnscs are bisexual, and require
mutual impregnation; in others the sexes are distinct. The mast
are oviparous. The eggs are surrounded by a thin shell {Chorion) y
between which and the yolk in some an albuminous fluid is inter-
posed, and which is sometimes homy, and but seldom covered with
calcareous incrustation. Often the eggs, when laid, are connected in
bunches, or adhere to each other by a gelatinous mass. The num-
ber of species that live on land is small in comparison of the much
greater quantity of species that live in fresh, and especially in salt
water.
Before we proceed to the division of the molluscs, we would
treat shortly of the shells which cover the body in most. The uni-
valve shells are called cochlecBy the bivalve, as those of mussels,
conchcB. There are also some molluscs that are covered by many
pieces of shell. Such a multivalve shell {testa multivalvis) has the
genus Chitorij where transverse calcareous plates, lying behind one
another in a row, cover the back. The bivalve shells are thicker
at the part where they are connected with each other. This part is
named thejxnnt {apex). At the point there are on the margin, within
the shell, usually projections and hollows, which mutually fit into
each other, and to which the name of hinge {cardo) is given. When
this margin is smooth, there is said to be no hinge {testa acardis).
In front of the point a slight depression is seen on the shell, which
is named the male depression {lunuUiy by Linnaeus anus) ; behind
the point is a chink, ordinarily smaller and more elongate, iii^Jlssure
{fissura^ in French Scussony with Linn^us vulva). On this the l^a-
ment is usually situated which, formed of elastic homy fibres, runs
transversely from one shell to the other. Where this ligament, as in
most of the bivalve molluscs, is attached to the outside of the shells,
it is obvious that they will be opened by its contraction. Yet
even where the ligament is placed internally the two shells are
separated from each other by its elasticity, because in this case the
fibres are forcibly compressed by the shell when closed. In those
bivalve molluscs which move freely, the opening of the shell is
turned downwards, the point upwards, and the ligament backwards.
LiNNiEUS, in his description, placed the bivalve shell with its point
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IN GENERAL. 683
downwards, and turned the ligament forwards. Blainville, on
the contrary, described it in the position which we have indicated ^
In the univalves {cochlew, testcB univalves, aubbivalves) the fol-
lowing parts may usually be distinguished. The extremity of the
spire is called the point (apex), which, when the animal is alive and
in motion, is turned upwards and backwards. Ordinarily the cavity
of the shell terminates in a point at this part, but it is sometimes
truncated horizontally {apex decollatus s. truncatus), which must not
be mistaken for an accidental firacture which always leaves an open-
ing. In some imivalve shells (as in the genus Patella), the cavity
between the point and the aperture is neither twisted to the right or
left, nor forwards or backwards. In most, on the other hand, this
cavity is turned. Usually all the turns run obliquely from above
downwards {turbinated shells, cochlea turbinata et iurrita); yet in
some the wreaths or turns run from left to right in a transverse
direction, whereby the last wreath includes the rest externally {testa
convoltUa s. involuta, the involute shell, as in the genera Conus and
Oliva), whilst others again are turned in one and the same vertical
plane, from behind forwards and from above downwards {cochlea
revoluta, as in the genus Nautilus).
The opening {apertura) of the shell is opposite to the point.
That part of the cavity, which is visible within the mouth, is called
the throat {Jaux). The mouth may be excised with a sinus ante-
riorly {apertura emarginata). Whenever it terminates forwards in
a channel {apertura canalifera), then the tube is called the tail
{cauda s. rostrum). At the mouth the external margin {labium
exterius s. labrum) is distinguished from the internal margin {labium
internum s. margo columeUaris). The spindle {columella) is that
part which runs in the middle of the shell directly from the point to
the aperture, and round which, as an axis, the wreaths are turned.
Such a calcareous axis is not present in all univalve shells; to see
this part distinctly, a shell sawn longitudinally must be provided.
At the internal margin of the aperture there is often on the ex-
tremity of the spindle, especially in the shells of younger molluscs, a
cavity which is called navel {umbilicus). Shells, in which this aper-
ture is wanting, are named imperforate {cochlear imperforatas).
^ That which with Linnaub is the right and the left shell still preserves this
name, since he turned the bivalve not only upside down, but also the fore part
backwards.
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684 ON MOLLUSCS
The internal cavity of the shell may extend uninterraptedly
through all the turns {cochlea monothalaviia s. uniloculars)^ or may
be divided by several partitions {cochUa polythalamicL). A cylindri-
cal tube traversing these partitions is named siphon (stpho). Such
a many-chambered shell is that of the genus Nautilus.
Cover {operculum) is the name given to a round calcareous or
sometimes homy lamina, which in many molluscs is attached to
the uppermost and hindmost part of the so-named foot, and which,
when the animal retracts itself within the shell, closes the aperture.
Sometimes a spiral line is seen on this part, as for instance in
the genus Turbo. Some writers name the shell, in this case, testa
suhhivalvis.
Finally, according to the direction of the wreaths, univalve
shells are distinguished as right- and left-turned. In most of them,
when placed on the aperture with the point backwards, the external
margin of this is on the right side, and the turns run from the point
to the aperture from the left side to the right {cochlea dextra). If
the converse takes place and the external margin of the aperture
lies on the left, then the turns run from the right to the left side
{cochlea sinistra s. contraria), and with this a displacement of the
internal organs is connected. Some species of molluscs have such a
left-handed shell as the rule, and in others (as in Helix) shells of
this sort occur as varieties or deviations, just as a displacement of
the internal organs, of the heart at the right-side, &c. has been also
observed occasionally in man*.
What has been stated may suflSce for the explanation of the
most common and necessary terms'. But for the general anatomy
and physiology it is desirable to contemplate shells in another
point of view, and to investigate the mode of their origin and
composition.
The formation of shells is effected by the external integument
^ On such left-tamed anail-shellB Chbicnitz has published Tarious remarks and
observations in the Journal Der Naturfoncher viiL s. 163 — 178, Xli. s. 76 — 84,
That from the left-handed variety of HeUx Pomatia right-handed young are again
produoedi was observed by him with perfect certainty ; ihid. xvn. s. i — 11.
' Comp. A. MuRRAT Fundamenla Testaceoloffiee, Upsalin, 1771, 4to, (also trans-
ferred to LiKK. Amomit. Acad. vni. pp. 107 — 150); Blaiitvills in Diet, det Sc not,
X. pp. 168 — 27$, article Cdnehologie (and the same writer in his Manual cited above) ;
Deshates, article CoquiUe, DicHonn, clastigvit dPBui, not Tome iv. 1813, pp. 431 —
449, &c.
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IN GENERAL. 685
or mantle of molluscs. B^aumub^ has illustrated the formation hj
his experiments. He found, on boring the shells of living snails, the
aperture to become closed again by a thin layer occupying the whole
of it, to which other layers were afterwards added. Thus there was
no calcareous matter secreted at the edge of the aperture by vessels
running in the shell, as in the reparation of bones where the forma-
tion of new osseous matter proceeds from the extremities of the
fractured bones. When, however, Reaumur supposes that the forma-
tion of shells is a mechanical transudation, that the secretion of lime
on the upper surface of the mollusc may be compared to calcareous
incrustations, which in certain waters and springs are formed round
bodies immersed in them, that the skin of the animal, like a sieve,
permits an adhesive fluid loaded with calcareous particles to escape,
and that this fluid by evaporation and rest loses its watery particles,
then his representation bears too forcibly the character of off-hand
ideas respecting living beings, and PoLi is so far justified in calling
the origin of shells an organic origin, and in rejecting this mechani-
cal explanation. It is clear that shells by the addition of new
laminae become thicker. In a shell several layers or calcareous
scales lie upon each other, which in bivalves originate from the
point; hence the shell is, at this part, of greatest thickness, and
becomes gradually thinner towards the margin. Thus every shell
consists, as it were, of many others, all of which becoming larger
and larger, lie under each other, whilst the innermost, the last
formed, extends beyond the others at the margin. In oyster-shells
and many other bivalves this may be clearly seen, and snail-shells
also indicate the same; in younger univalves the number of turns is
fewer; the larger shells of the same species present a greater num-
ber of wreaths than the smaller, without however those wreaths,
that were already present in the young ones, increasing in size. So
also the spines, tubercles, and other excrescences of imivalves are
at first short and obtuse, and become larger and more acute by the
addition of new layers. The increase however is not at all times
uniform, but in the cold of winter and the great drought of summer
^ Dela formation el de VdeeroiisemerU des eoquillet, MSm. de VAcad. royale de» Sc.
1709, Paris, 1733, pp. 364—400; Paris, 1741, pp. 303—311 ; comp. also Pou Teatae.
utriutque SicUice, Tom. I. (in the introduction} and HsuBiKOSB Syttem der Sialologie.
Eisenach, 1823, i. ates Heft, s. 236 — 242,
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686 ON MOLLUSCS
is suspended* Hence stronger stripes arise, which indicate sttcoe»-
sion in growth, and are the traces left of it.
As to chemical composition, besides some other less constant
constitnents and a small quantity of phosphate of lime, mdyalye
and bivalye shells consist principallj of carbonate of lime and an
animal substance, a membranous guhsiratum^ that remains after the
calcareous matter has been dissolved by acid, and is usuallj Teiy
nnall in quantity compared with the calcareous matter. FoLi,
when he exposed the membrane to the fire, saw it quickly take
flame, on which he perceived a smell like that of burning horn; a
spongy carbon remaiued.
This conducts us to a correct view of the nature of shells,
which belong to the same tissue as horn, hair, and in part also
scales, f.6. to homy tissue. In most animals the dermal skeleton
(such is the name given to the hard parts placed externally, to
which the muscles are attached,) is homy, whilst, on the other
hand, cartilage is almost always the foundation of an internal
skeleton, especially of a trae neural skeleton. The microscopic
stracture of bivalve and univalve shells has been chiefly illustrated
by the investigations of Carpenter. In some bivalves the entire
substance of the shell consists of layers of membrane, without
visible cells, in others such a membranous tissue forms the inside of
the shell, whilst on the outside columnar, often hexagonal cells are
visible under the microscope, which to the naked eye, or with the
use of a lens, look like fibres. They stand nearly at right angles
on the surface of the shell from within outwards, and are filled
with carbonate of lime. In each layer they are at that part onlj
which projects beyond the edge of the preceding layer; thus they
have been secreted by the edge pf the mantle, whilst the whole
mantle, in every new formation of a layer, produces a membrane
that covers the inner surface of the entire shell. The univalve
shells of the gasteropodous molluscs have only a small quantity of
organic substance; in many three layers of plates may be distin-
guished; the direction of these plates is different, and those of the
middle layer intersect those of the external and internal at right
angles. Every plate consists of a row of long columns, or pris-
matic cells, which axe arranged side by side^.
> See W. B. Carpenteb Annals of Nat, Hidory, VoL xn. 1843, PP.377— 39<'f
Pi. xni. XIV. Mid espedaUy his Uter, more general investigations, announced in
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IN GENERAL. 687
In many, especially bivalve shells, a homy, brown-yellow
oiitennost covering is found, which has been named epidermis or
re/o/Nii^j periostracum also (in French Drap marin). Sometimes it
is hairy or divided into scales, which however are more apparent at
the margin of the shell, having been worn off from the parts pre-
viously formed. This membrane has been regarded as a continua-
tion of the cuticle of the mantle, by which this is connected with
the margin of the shell. More correctly, perhaps, this covering may
be explained from a confluence of the intercellular matter, a resi-
duum of the formless homogeneous substance (the csyioblastemaYy in
which the cells filled with lime were formed. Let it be supposed
that at the outer margin of every layer this substance remains
without cells, and consequently hardens like horn. If these edges
should close upon each other, then a smooth epidermia will arise; if
they should remain more distant from each other, then a scaly, floc-
culent or hairy covering will be formed.
The colours, presented by bivalve and univalve shells, are de-
posited only in their outermost parts, the inner layers axe white.
This may be explained by the circumstance, that the colouring
matter is secreted especially by the edge of the mantle. But the
mantle grows with the animal, and thus each succeeding layer of
the shell is coloured at its outer margin alone, whilst its remaining
portion, secreted by the rest of the surface of the mantle, remains
white. In this way a series of coloured edges arises, which, closing
upon each other, form the outermost coloured surface of a shell.
There are however some univalves (the genus Cyprasa and some
OUwb) in which, when full grown, the colours are deposited not on
the surface alone, but also in a deeper layer, whilst at the same
time the superficial and the more deeply lying colours are different.
These molluscs are at first covered with a thin shell, of which the
colours must be ascribed to the edge of the mantle. As the animal
grows, lateral appendages of the mantle are developed, which throw
^emselves like wings over the shell, and secrete on their outer
BepwiM of ike Brititk Auociation for 1844 wid 1847. The chief puticulan of theae
inveetigatioiui may be also found in the Article SheU by the same writer in ToBD's
Cfjfdopaedia, TV. 1849, P- 55^^ ^*
1 Page 17, 18.
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688 ON MOLLUSCS
surface a calcaxeous layer as hard as ivory and of a different colour,
which covers the preceding layers*.
Many bivalve and univalve shells have on their inner snrface
lively and glancing colours, which must not be confounded with
the external colours. They are the play of colours from the refrac-
tion which is seen to arise on the transmission of white light
through a prism, and which causes the beautiful phenomenon of the
rain-bow. The mother of pearl therefore is not to be ascribed to
any special colouring matter, but to very fine streaks and folds of
the internal membrane of the shells '. It continues even after the
carbonate of lime has been removed by an acid, and Carpenter
observed that the play of colour disappeared when he extended the
membrane with needles so that the folds were obliterated*. These
changes of colour are particularly beautiful in the genus Hcdiotis^ a
species of which is used by the Japanese to ornament the lackered
furniture of their houses.
Whenever the fluid, which forms this innermost layer of bi-
valves, is efiused in the form of small drops, pearls arise, which
have frequently a very irregular shape. These may accordingly
exist in very different species of bivalves, also in turbinate uni-
valves; yet they arise usually in bivalves, and especially in Melea-
ffrina margaritifera and Unio margarittjerus. The former species
of conchifer is fished in the Persian gulph, the last is found in
fresh water in different parts of Europe, and also in our country.
According to Home, pearls have undeveloped eggs for a nucleus,
which have remained accidentally on the outside of the mantle
within the shell. Such may often be the case, but there is no
1 BBuauiiBB in the Jowmal SJSUU Nat. l. 179a, pp. 307—315, quoted by
SoHWEiooKB ffcmdb. der NcOwrgeMck. der dsdetUoten wtffeglkd, Thiere, b. 681. A dif-
ferent opinion of Bbuoui&e, that the animal of OjfproM leaves its sheU, and fonne a
new one, though adopted by Lakarck and Sohwzioobb, can no longer be defended ;
compare Dbbhateb in the new edition of Lamabox ffiM, nai, det Animmix «. Vert^-
brea, x. p. 485.
* This was first announced by Bbiwsteb Phil. Tram. 18 14. If a drop of sealing-
wax be allowed to fall on a mother of pearl shell, the surface of the wax, which has
been in contact with the shell, presents the same colours. It may be supposed that
this was an impression of the surface. But it appears, that in this experiment a thin
scale of the surface of the shell remains adhering to the wax,
3 AnncUt of Nat. Hid. xii. p. 383.
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IN GENERAL. 689
reason why the formation of pearls should not be ascribed also to
other irritants affecting the mantle. It has been observed at least
that injuries of the shells and wounds, caused by boring worms,
have had the production of pearls for a consequence; and the secret
of LiNN^us for favouring the production of pearls (in Unto)^ con-
sisted, most probably, in boring the shell in different places in con-
chifers, which were submitted to these experiments^.
The shells of molluscs, from their variety of colour and form,
constitute no small part of the ornament of natural-history collec-
tions. The knowledge of conchology is of the highest interest to
the Geologist, since the petrified and extinct species afford im-
portant characters for distinguishing the different strata. But,
more than this, the knowledge of the molluscs is of great value to
general physiology. To the celebrated Danish zoologist of the last
century, O. F. Mueller, the honour is principally due of having
raised this part of natural history from the fondness of collectors to
the scientific contemplation of naturalists; it was his impressive
exhortation that thenceforward attention should not be confined
solely to the hovse or the shell, but, above all, be extended
to an accurate investigation of its inhabitant*. Already had SwAM-
MERDAM in Holland, and Martin Lister in England, investi-
gated the internal structure of some molluscs. PoLi and Cuvier
made this subject a primary object of their numerous inquiries, and
thus, in the course of the last fifty years, and even in our own day,
through the labours of Delle Chiaje, Owen and others, a clear
and extensive view has. been obtained of a field of comparative
anatomy that previously was almost unknown. This was an inesti-
mable gain for a science which, if it is indeed to exercise an
important influence on physiology, must not, in any sense, be
limited to a few classes of animals, but must, in reality, be com-,
paring J and must compare generally.
* Compare Ghsmnitz Vom Ursprunge der PerUn, Naiu/ifortcher, xxv. Halle, 1791,
8. Ill — 130, and Bkokmann'b QeBchichie der Erfindungen, cited there. On the origin
of pearls from eggs of Conchifera see Phil. Trans. 1674, Vol. ix. pp. 11, 11, and
egpedally E. Homb in Phil, Transact, for the Tear 1816, Part 3, pp. 338—341. This
peculiarity had already been observed in 1673 ^7 ^ T)9Xie, H. Abnoldi, at Christiana
in Norway,
* See his Vermium terrettrium et JUtviatilium ffistoria, Hauni», 1774, 4to. Tom. ll.
Prsefat. p. i.
VOL. I. 44
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690 ON MOLLUSCS IN GENERAL.
Some mollnBCS have no head distinct from the bodj. Heie the
principal mass of the neryons system is not placed near to or aboTe
the mouth, but if nerrous ganglia be found in this situation, they
are inferior in development and size to other nervous ganglia;
no nerves for special organs of sense arise from the nervous mass
situated above the oesophagi. The mouth conducts immediately
into the oesophagus, and has no teeth or tongue. These are named
headless moUuecs {Aaphala). The other molluscs have a head more
or less clearly distinct. The head-bearing moUuscs {Cephalophara
or moUusca encephala) are more perfectly organised, and amongst
them alone are species found which live on land; most of the
species, however, reside in the water, like the acephalous molluscs.
These last may be arranged in two classes, the mihed and the
bivcdve acephalous molluscs.
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CLASS XI.
TUNICATES (TUNICATA)\
The Tunicates are acephalous molluscs without shells. The
external covering is perforated by two openings, in other respects
like a sac and entirely closed, of various thickness and hardness, in
some gelatinous, in others leathery. It consists of a tissue that is
sometimes very composite, includes cells, fibres, nuclei, crystals also
of carbAaate of lime, and of which the fundamental matter contains
no nitrogen, but only oxygen and hydrogen, and agrees in compo-
sition with the vegetable cell-membrane {cellulose)^ as was first
discovered by C. Schmidt in Ascidta mammilaria *.
The intestinal canal forms in most of the Salpm some convolu-
tions, which are united to form a clue which occupies a small part
alone of the cavity of the body, and to which Fokskal in his
descriptions gave the unmeaning name of Nucleus. In Salpapennata
8. cristaia Cuv., however, the intestinal canal runs straight from
the mouth to the opposite end, and has, close above and behind the
mouth, an expansion or stomach that terminates blindly and lies in
the opposite direction. The mouth, placed internally and bordered
by a tortuous band, is properly only the beginning of the oesopha-
gus. A furrow in the middle of the body runs from the opening
of the common integument, by which the water is introduced, to
this commencement of the oesophagus, and serves probably to con-
duct towards it the food that is conveyed with the water. [Above
this fturrow or semicanal, in the substance of the inner mantle is
an organ, called by Huxley endostyle^ a long tubular filament
with thick refracting walls. It is of various length, very short in
^ Compare on this class :
CuviKB Mimoire tv/r let ThaUdes et let Biphoret, Ann. du MutSum, iv. 1804, pp.
360 — 38a, PL 68 {Mimai/res awr lei MolUu^pies, No. 19) ; Mimoire tur let Aecidiet et
Uur AnaUmie, MSm, du Muaium, n. 181 5, pp. 10 — 39, PI. I. — in. {M6m, eur let
Mollutquee, No. 20.)
J. C. Sayigitt Mimoiret sur lee Anima/ux tans verUhret, n. premier fascicule, Paris,
1816, 8vo.
■ Zitr vergleiehenden Phytiologie der trirbeUaten Thiere, 1845, s. 62 — 65. Extensive
microscopic inyestigations respecting the structure of the external oorering in many
twniotOa, iUustrated by beautiful figures, have been published by Loxwiq and Koelli-
KXB, Ann, da 8e, natur, si^me S^rie, v. 1846, pp. 193—338.
44-2
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692 CLASS XI.
Doltolumy extending in some Salpw as far as the oesophageal aper-
ture. The tube is closed posteriorly, but, according to Leuckart,
communicates anteriorly with the longitudinal semicanal. Its walls
are composed of large nucleated cells, arranged perpendicular to its
cavity, and Leuckart suggests that it may perform a secreting
function. Huxley has seen it in SalpcBy Pyrasomaia and certsdn
BotryllidcBy as well as in Cynthia, and, as it seems to be figured
by Savigny and others, it may perhaps be concluded to be common
to the Tunicata ^]
In the AscidicB the intestinal canal usually forms two bends,
which lie toward each other, and have their convexity facing back-
wards. The intestinal canal commences at the bottom of the
branchial cavity, and becomes narrower posteriorly. The stomach
is sometimes merely indicated by a first expansion of the intestine,
which is not sharply defined; in other cases its form is elongate and
cylindrical, as in Didemnum and Botryllus^. The extremity of the
intestinal canal mounts higher than the commencement of the
oesophagus, in the direction of the second tubular opening of the
external integument of the body. The liver lies as a stratum of
glands on the walls of the stomach or the intestinal canal. Salivary
glands have as little been met with here as in the bivalve molluscs.
The blood-circulation of the tuniccUa presents the remarkable
phenomenon, that the direction in which the blood streams from
the heart is at intervals altogether reversed, so that the heart alter-
nately drives the blood to the branchiae, and may be called venous,
and alternately receives the blood from the branchiae and, as in the
rest of the invertebrates, is arterial. This peculiarity was first
discovered in 1821 by Van Hasselt in Salpa*, but was afterwards
observed by Lister, Milne Edwards, Van Beneden and others,
» [See HuxLBT Phil. Trana. 185 1, p. 588.]
* See Sayignt Mim. n. PI. zx. zxi. On the intestinal canal of Aseidia compare
CuviXB Aacidies, PI. i, fig. 5 ; CcOalogw of the Physiol. Series of Comp. Anat. of
the Mtiseum of the Boyal College of Surg. i. PI. 5, fig. i, Phallusia nigra Sav., and
Van Benxden Jtecherchee awr V Embryogenie, VAnatomie et la Phyeidlogie da Aacidiet
rimplee, Mim. de PAcad. royale de Belgiqne, Tome xx. 1846, PI. i. fig. 6.
* Alg. Kotut-en Letterbode iSai, I. bL 1x5, 116 (translated in Ann. dee Se. no/, ni.
pp. 78—81). Afterward Mbtxn in his Voyage obserred this motion of the blood in
two opposite directions in Salpa also; Act. Acad, Ccet. Leop. Carol, xvi. l p. 377.
With this in some degree may be compared, what was observed by J. Muellsb
(Meckel's Archiv, 1828, s. 32 — 29) in Nephdis respecting the inconstancy of direction
in the blood-current, and by E. H. Wkber (ibid. pp. 399, 400) in young leeches.
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TUNICATES. 693
in different species of simple and compound Asctdtce, so that it may
be safely concluded to be a common property of the animals of this
class. The heart has an elongate form and does not exhibit any
sudden expansion and contraction, but a sinuously progressive
constricting and widening, which is similar in form to the peri-
staltic motion of the intestinal canal. Besides larger vessels the
blood moves in interspaces which have no proper walls.
The respiratory organs are gills which are very different from
those of the bivalve molluscs. In the genus Salpa the gill is
extended as a longitudinal band running obliquely in the inner
cavity of the tubular body, with many transverse grooves closely
arranged side by side. In the Ascidice the respiratory organ forms
a sac at the bottom of which the entrance to the oesophagus is
situated. The very thin membrane, of which the sac consists,
presents transverse and longitudinal stripes that cross each other at
right angles like trellis-work, to which a net-work of blood-vessels
corresponds. The gills, as well of the SalpoB as of the AsddioBy
are beset with vibratile cilia ^.
Male and female organs are, at least in the Ascidtce, imited in
the same individual. Often they are intimately connected with each
other and are only to be distinguished by microscopic examination
of their content. Some Ascidice, as Ascidia ampuUaris V. Bened.,
have c^uite a double apparatus of sexual organs, which lies behind
in the intestinal sac on each side of the intestinal canal. In some
the oviduct is wanting, in others no vas deferens exists; a single,
long, tortuous canal, which is present in the compound Ascidice,
and is generally held to be the oviduct, ought, according to Milne
Edwards, to be regarded as the vas deferens. In the Ascidice,
besides the usual propagation by impregnated eggs, a multiplication
by buds has also been detected*.
^ Milne Edwabds and before him Lister admit openings like fissures between the
quadrangular meshes by which the water is expelled from the branchial sac (Milne
Edwabds Ohterv. s. I. Ate. comp. pp. 17 — 20). It is difficult to determine, in speci-
mens preserved in spirit, whether they are really openings or spaces closed by a trans-
parent membrane. [Huxley says that in Pyroaoma the respiratory cavity communi-
cates freely by means of apertures in the branchial network with the post-branchial or
anal cavity. PhU. Trans. 1851, p. 581.]
* Compare for some other peculiarities, the notice of the different genera in the
Bystematic arrangement, in which also the remarkable propagation of the Salpce is
noticed.
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694 CLASS XI.
The nervous ajstem presents a single ganglion, which in the
AscidicB is placed on the internal covering of the bodj, usually
regarded as the mantle, between the two tabes into which the body
is elongated. Besides other nerves which radiate &om this ganglion,
there appears to arise from it a nervous ring that surrounds the
tube by which the water penetrates (the oral and respiratoiy tube).
Other ganglia which are noticed in Ascidians bj some writers, are at
the least to be regarded as donbtful ^. In Salpa a ganglion (or a
nerve-mass formed of several ganglia united) lies close behind the
anterior, broader opening on that surface of the body which is directed
upwards •. Several nerves run radiately from this nerve-mass.
Vestiges' of organs of sense are not entirely absent. In the
Ascidice at the branchial aperture a circlet of small filiform feelers,
sometimes digitally incised or pennated, is found. In the ScUpa
there lies in front of the central mass of the nervous system an
organ of an elongated form, consisting of two laminsB with an
internal margin smooth and an external striated, which Eschricht
regards as an organ of feeling and compares with the four lamellsQ
which surround the mouth in bivalve molluscs'. Organs of sight
also have been observed in the Ascidice, There are found both
around the branchial aperture of the mantle and around the aper-
ture towards which the extremity of the intestinal canal proceeds,
a ring of eye-points, often of a yellow colour *. [In Salp^a a vesicle
^ ThvLB SoHALK, whose IHateri, de Aacidiarum Stmetura I un aoquainied with
from citations alone, records a nerve-ganglion in the posterior part of the body,
between the convolutions of the intestine ; Grant speaks even of three ganglia, (hi-
lines of Comp. Anai. i8ii, pp. 205, 106. Compare on the nervous system of the
AaciduB, Cuvikr Sur let Atcidies, p. 15, PI. n. fig. 1, 0, fig. 5, g, Saviqnt M6m. h
pp. 117, 118, PL X. fig. 2, I D, &c.
' This position of the central portion of the nervous system caused EscHSlCBT to
give to the surface the name of ventral side. See on the nervous system of Salpa
Metkn 1. 1. s. 395, and the fig., Eschricht Anat. physioL UndenSffdae over Salperne.
Kjobenhavn, 1840, 4to, printed from the Vidensk. SeUk. Naturvid. og math. Skr. vni.
pp. 12 — 14, Tab. I, fig. ^u, Tab. ii. fig. 8u, fig. 13; Milme Edwards in CxmaB.
Ani., ed, Ul., MoUusq, PL 120, fig. i 6, a, PL 121, fig. 2 a, h, fig. 2 h,
' [HUXLBT Anat. of Scdpa and PyrotomOy Ph. Trans. 185 1, p. 571^ calls this
'* Idnglichea organ** of Esohrioht " Usnguet." He supposes that it may subserve the
gustatory function. It floats freely in the respiratory cavity to which it is attadted by
its base.]
^ Grant OrUUnei, p. 361, says that in Aicidia {PhaUusia) iniatinedda eight red
eye-points lie round the branchial opening, and six around the efferent tube ; so abo,
according to the investigations of Will in various species of Cynthia Sav. and Ckte-
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TUNICATES. 696
attached to the lower snrfiEU^ of the nerve-mass and contaming
calcareous bodies would seem to represent the auditory capsule and
its otolithes ^.] Such an organ abo, but containing a concrement of
a more composite form, was discovered by Eschricht, similarly
situated in a simple Ascidian Chelyosoma Madeayanum, and noted
as the auditory organ * in 1841.
The muscular system in Ascidtce consists of a layer of fibres,
which cross each other on the inner covering of the body (the so-
called mantle), but ran principally in a longitudinal direction. In
the SaljHB muscular bands are seen remote from each other, which,
like a girdle not entirely closed at one part, surround the internal
cavity of the body, the respiratory space. These bands consist of
fibres that run parallel to each other in the direction of the bands,
and thus form a right angle with the longitudinal axis of the body.
On the position of these animals in the natural system there
can exist no doubt when they are referred to the common type of
the molluscs. They must be regarded as the lowest division of
these animals, that immediately close in upon the bivalves. The
compound Ascidians in propagating by gems remind us of lower
animals of the class of the Polyps. More especially still do the
Ascidians resemble the Bryozoa (comp. I. pp. 92, 93), but we should
prefer to tmite these last with the MoUusca^ rather than on account
of this affinity to separate the Tunicata firom the MoUusca.
Species of this class are met with in all seas. The Saljm are
numerous in the Mediterranean and in the ocean, especially between
30^ and 40* N. and S. Lat., in colder regions and in higher latitudes
they are of more rare occurrence although they extend to 60** N.
and S. Lat., and were once observed near the Norwegian coast even
Una. V. BEinEDBN in a spedes of Atcidia named by him Aaeidia vitrea, fomid eight eyes
at both openings; Rech. 9, 1, Atcid. Hmples, p. 6i, PL iv. fig. «. In Atcid. ampuUoidei
Y. Bbn. such eye-points cannot be seen, but when young and still moving freely the
animal has one or two black eye-points ; ibid. p. 40.
1 [Huxley L 1. p. 571, PL xvi. fig. a.]
* EscHBiCHT Anatomisk BederweUhe of Chelyo9oma Madeayam/um of Sowebbt,
Kjobenhavn, 1841, quoted by Siebold Ver^. Anat, i. p. a6o (note), who suspects
from the figures of Dellb Ghiajb and SAVioirr that the organ exists very generally in
this class. Milne Edwabdb (see the figures of Salpa referred to above in the illas-
trated edition of Guvieb) thinks that it must be regarded as an organ of vision.
That Mbtbn had observed this organ lying above the central nerve-mass, I collect
from his figure. Tab. xxvii. fig. 18.
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696 CLASS XI.
in 62° N. Lat.* They appear on the coasts, however, only raiely,
but keep by preference in the open sea, fax .from the continent
Ascidians, as well compound as simple, occnr in much higher lati-
tudes; Boltenia omfera was discoyered in Davis's Straits at 69*
N. L., Synoicum turgena in Phipps's voyage at the north coast of
Spitzbergen. Species of all the forms, nay, even of most of the
sub-genera, are found in the Mediterranean sea; the apparent pre-
ponderance in number of species in this sea, above all others, may
however be principally ascribed perhaps to this, that its productions
have been more perseveringly investigated by numerous observerB,
than those of the other seas.
^ In the autumn of 1839 in great numbers near the island Brtmamgmr; Sabs
Fauna littoraUi Nfyrvegia, Christiaoay 1846, folio, p. 63.
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SYSTEMATIC
ARRANGEMENT OF TUNICATES.
CLASS XI.
TUNIOATA.
Animals without distinct head, inclosed in an external covering
coriaceous or membranous, and furnished with two apertures. A
second tunic included in the former, perforated by two orifices cor-
responding to the apertures of the external covering. Cibarious
canal with mouth and anus internal, concealed. Bespiration bran-
chial, branchise situated internally in front of the internal mouth or
the commencement of the oesophagus, of different form in different
cases, never divided into four laminae. Heart tubular, propelling
the blood in a mutable direction.
Order I. Thaliacea,
Body prismatico-cylindrical, with two apertures opposite, sub-
terminal. Covering pellucid, thin, rigid. Animals swimming
freely, drawing in the water by one aperture, expelling it by
the other.
Family I, Salptna. (Characters of the order those also of the
single family.)
Salpa FoRSK., Ghel., Cuv., Lam., species of Hohthurta L.
( Thalia Brown, Blumenbach, Dagyaa Banks and Soland., Gmel.,
Home). Branchiae extended obliquely through the internal cavity
of body. Form in each species two-fold, the solitary progeny alter-
nating with the associated.
Comp. P. FobbkIl JkBcripUanea animalivm, Hauniffif i775> PP* '^^ —
117; quad. Icon. Rer. natur. ibid. 1776, Tab. 35, 36; Cuv. Mim, (▼.supra
p. 691) ; A. Ds Chamibso Ih AiMmalibu» qmbvMdain e CUum vemUtm.
PoBciewLvM I. Z>« SaJ/pa. Berolmi, 18 19, 4to; Mbtbit L i.; Esghbioer
1. 1. ; Kbohn ObHTvalumt tur lajf^tUraUan ei le dSvdoppetneiU det Biph/om,
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698 CULSS XI.
Arm.det Se. not,, zUme Sirie, yl 1846, Zoclogie, pp. no — 131. Huzliy
ObaervaJtioni upon the Anatomy and Phynoloffy of Salpa cmdPynmoma. PhiL
Trans. 185 1, Pt. n. p. 567.
The species of Salpcs ore verj generallj dispersed ; sometimeB
they are met with in incredible quantities together ; notwithstand-
ingy these animals a century ago were entirely unknown. The first
announcement of them was made by P. Bbowk in his CvoU and
Natural History of Jamaica^ published in 1756, under the name of
Thalia; afterwards followed the description of the species observed
in the Mediterranean and Bed Sea» under that of Salpa by the
Danish traveller Fobskal'. Bosc and Cuyier next pointed out
the resemblance of these animal species, described under two di£^
ent names, which ought to be referred to the self-same genus.
The body has an aperture at each extremity, but these apertures
have a different form. The one is broad and transverse, and presents
a valvular membrane, inasmuch as one of its margins is reflected
inwards. The valvular membrane prevents the efflux of the water,
which flows inwards by this opening and which is expelled by the
opposite one from the contraction of the body. This expulsion of the
water is the means by which the animal moves, so that the narrower
opening is tiumed backwards. Cuvieb thought that this opening
was the anterior, and that thus the animal moved backwards.
Since, however, the stream of water conducts also the food, and
since in the Aacidia the entrance to the oesophagus is situated
behind in the respiratory sac, the common opinion, that the broad
opening is the anterior, deserves to be preferred to this idea. The
intestinal canal is situated on the outside of the respiratory cavity,
in the space intervening between the external and internal covering,
but terminates by both its apertures in the respiratory cavity.
These animals, according to the testimony of Pjebon, Tilbbittb,
Meyen and others, are usually phosphorescent by night. The Salpa
are met with at one time singly, as distinct individuals, at another,
imited either in rings or in long chains, of which the arrangement
is various, yet similar in individuals of one and the same speciea
These are attached to each other by tubercles or prolongationa
Chakisso, from his observations on living animals, arrived at
the conclusion, that successively a generation of distinct Salpa
alternates with that of Salpa connected, and forming a chain.
Thus a metamorphosis occurs, which, however, does not take place
1 Bbuoui^b, who gftve in the JBneyd. method, an extract from the deacriptioDs of
FoRSK., changed the n«me Salpa into Biphora, which has found no genenl aooeptuce.
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TUNICATA. 699
in the same individual, but in two suooeeding generations. The
parent animal, for instance, always resembles her grand-daughter
and her sister, not her daughter, which, again, resembles the pre-
ceding generation. This conclusion seemed to most succeeding
observers so strange, that they thought it could not be adopted ;
Meyen supposed that these animals are always bom as single
individuals, and only become united subsequently. ISTevertheless,
EscHRiCHT discovered within single individuals embryos connected
together in a wreath, so that hereby Chamisso's conclusion was
established, that chains of connected Salpa were formed even
within the parent. Steenstbup brought this peculiarity in the
propagation of the ScUpce into xmison with other phenomena in the
animal kingdom, in his interesting work, Om FoHplantning og
Udvikling gjenem Generationsrackker, (see above, p. 70), and the
meritorious Norwegian observer Sabs, completely confirmed it
{Faun, lUtor. Norvegias). With it too the numerous observations
lately communicated by Krohn, are in accordance. The solitary
Salpw are sexless, and are propagated by internal germs or buds
connected in strings. Hence the associated form appears to be
the most perfect, that in which organs of propagation are developed,
(and organs of impregnation also, testes) ; each individual of the
associated Satpcs produces only a single young ona
Sp. Salpa maadma FoBSK. lean. Tab. xxxv. fig. ▲, Milne Edwards, Cuv.
R. Ani., id, iU., Moll, PI. 120, m, fig. i (here belongs, according to
Kbohn, ScUpa Forshalii Lesson and the solitary form Salpa africama
FoBSK.);— 5. runcinaia Chavisso 1. 1. fig. v. Salpa funformia Cuv. M6m,
8, I. Biphort8, fig. 10, Sabs 1. 1. Tab. vin. figs. 44, 45, Tab. ix. \^Salpa
pinnata Fobbk., S. crittata Cuv., Fobsk. Icon, Tab. cit. fig. b, Cuv. Mim,
figs. I — 3, IT, Chahisso L L fig. I (associated form arranged in a circle) &c.
Note, — The species of SalpCB are not yet sufficiently known, of many at
least, the synonymy is very difficult, full of doubt. Kbohn 1. L has endea-
voured to. tmravel the species from the Mediterranean. To the authors
already ctted, the French voyagers QuoT and Oaimard Voyage de rAttro-
lobe, Zoologie, Tom. iii. 1835, pp. 559 — 598, PI. 86—89, m« to be added.
Anchinia EscHSOH., Rathkb. Small Salpce aggregated in a single row
to a geUtinous filament. See Wieomann's Archiv, 1835, I. p. 85.
Doliolwm QuoY and Gaih. [Body cask-shaped. Branchise in two
bands with perpendicular bars, one on the dorsal {hcsmaC) the other
on the ventral {TiewraV) surface of the respiratory cavity, converging
and passing into each other posteriorly.]
DoUolum Otto differs from that bearing the same name : whether it be
a Beroi or a mntUated Salpa, I know not. Comp. Nov. Act. Acad, Lcop,
Car, XI. Tab. 41, fig. 4.
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700 CLASS XI.
[Sp. DoliUum dentieulaium QuOT Kod Gajm. The openings of the bnnuiul
cavity are short tabes, the anterior denticiilate, the posterior sonomidfldby
fine filaments. S. Pacific. If it be ever aggregated, it has not been met
with in this state. HuxuBT On DoUoium and Appendietdaria, Phil. IVou.
i85«* PP- 599» ^3]
[AppendicuUma Chamisso, Quot and Gaiil (Oikopteura Mebtkhs).
Body irregalarly OTate, with a long appendage attached to the
dorsal sur&ce at the posterior part, and making a large angle with
the axis of bodj. Opening into the respiratory cavity anterior,
outlets two short tubes ciliated where thej leave the cavity, open-
ing immediately on the dorsal surfiioe in front of the base of
appendage*. No proper branchias.
There has been much doubt respecting the pUoe and the nature of
Appendictdaria, Chamisso, who found his species in the N. Pacific^ con-
sidered it to be a Meduta allied to genus Otdum, Ebohsoholti, who
observed the same, or a nearly allied species, in the South Sea, refSened it
to the Heteropod MoHuscs, whilst Msbtbns, who voyaged in the nmc
seas, thought it was a Pteropod related to Clio. See Chamibso Nov, A<i.
Acad. Oca. Leopol, Car, x. Ft. 1, p. 361, Esohscholtz in Okih's Zm.
1895, s. 736, Mertens MSm. de VAead. imp. de St. POehbowg, 183T,
Tom. I. p. 905. Lbuokabt {Zoolog. Unienwk. n. 1854, pp. 80^ 81)
regards Appendieularia as a lanral state of some Ascidian, all of which haw
an appendage in the early condition, whilst with Huxlet {On Appeadkar
laria and Doliolum, Phil. Tram. 1851, p. 595) and Geqenbaubb (Soebold
and KoLUKBR'sZettoeA. /. tnHmach. Zod. B. vi. 1855, p. 406, &c.) it ii an
adult form.
Sp. Appendictdaria Jtabdlum Chavisbo, see Huxlet 1. cit., who found the
testis in the fully developed condition abounding with MpermaUma: the
ovary has not been ascertained ; coast of SootUnd, and common in the
Bristol Channel near Tenby. For a description of four Meditemnesn
species, with the anatomy, see Geoenbaueb loc cit.]
Anckinia EsoHSOH., Bathes. .Small Sa^pa aggregated in a singk row
by a gelatinous filament. See Wibomanit's Archiv, 1835, i. p. 58.
1 [These tubes were fint discovered by GBOEiniAUBR, but from the great tmspsr
rency, he did not detect their openings on the sur&oe of the animals, but supposed them
to oondiiH the fluid from the respiratoiy cavity, in order either to mix it directly with the
blood, or by expanding more largely to bring it in near contact with the blood through
their thin walls. See his paper, pp. 415, 416. Huxlbt, as well by direct obsemtion
as by feeding his animals on indigo, saw clearly the external apertures of the tubes in
front, and on each side of the anus. In one observation he found the current to set m
at these apertures, and out at the anterior opening of the respiratory sac See hit
paper on Appendictdaria JUibdlum, Quarierlif Journal qf Microi, Se. No. 15, 1856,
pp. 181— 191.]
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TUNICATA. 701
Order II. Tethyonidea.
Body sacciform, with two apertures mostly approximate. Cover-
ing coriaceous or gelatinous, mostly opaque. Branchial sac large,
reticulato-fenestrate, with rectangular areolae; the beginning of
oesophagus situated at the bottom of this sac. Animals mostly
affixed. Propagation oviparous and gemmiparous.
To this order some compound animals belong which were formerly
referred, for the most part, to the Alcyonidia (see above, p. 78).
The discovery of their tnie affinity is one of the most interesting
results of the accurate investigations of Savigny. Compare on this
order the above-eited works of this author, as well as those of Milns
Edwards, Van Beneden, kc
Besides the two divisions of Aacidite into simple and compound,
Milne Edwards has adopted a third, that of the social Ascidice,
which are distinguished by forming gems, without being grown
together like the compound Ascidiana. This gemmation has, how-
ever, been noticed in a species ordinarily simple^, and may perhaps
occur in all the animals of this division.
Young Aaddiatia, which do not originate in gemmation but
proceed from eggs, undergo an interesting metamorphosis. In the
early stage they move freely, and are provided with a long tail, as
was communicated by Milne Edwabds {Ann, dea Sc. ncU. xv. p. 10),
as early as 1828, and was afterwards more fully described by
Y. Beneden and others. They fix themselves by that extremity
which is opposite to the tail, which they then lose'. In compound
Aaddiana, according to the observations of Sabs on BotryJlua, such
a cercarisB-form larva may already enclose a group (eight) of united
Aacidiana, Thus, even before the Aacidia has become attached, by
the division of the gem the commencement is made of a colony
which is capable of further multiplication by the formation of
gems. Not, however, in all compound Aacidicma is this original
1 AocordiDg to BoHADBOH in Aacidia iniettinalis {PhaUusia inteBtinalit Say.). See
J. B. BoHADSOH De quibusdam Animalihui marinis. Dreads, 1761, 4to, pp. 132 —
135, Tab. X. fig. 5.
* As to the question, whicli is the anterior and which the posterior extremity of
tl^ese Cercariae-fonn larvae, consult R. Lsuckabt Ueber Morphologie und die Ver-
wmdiKhqftiverhaUniite der WirheUotm Thiere. Braunschweig, 1848, 8vo, s. 173, 174.
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CLASS XI.
to be detected, not st leasts aeooiding to Miuns Edwabds,
Familj IL Lucioe. Apertures of cctBcnal ooTering opposite,
termiiial. Branchial sac girdled anteriorlj hj a menibniioiis denti-
calate ring, open posteriorlj. Several animals aggregated to fam
a componnd bodj swimming fireelj, cylindrico-conical, hollow
internally.
PyroBoma Pl^SON.
This genus of compomid AscidioB was first discovered by Pebon and
his feUow-Yoyagers in the Atlantic ocean under the Tropics, when in
a dark night numerous specimens of it appeared to form a broad
band of light acroas the sea. From this phosphoric quality, the
name (^Fire-body) is derived. At first these compound animaJH were
supposed to be a single M^im^l^ and the single individuals of which
a PyroBoma is compounded, to be little tubercles on the sor&oe
of the animal See PiaoK Mem, 9wr le nauveau gtm/re Pyrowma^
Ann. du Musewn^ iv. pp. 437 — 446. For a more accurate know-
ledge of this remarkable genus we are indebted almost exduaivelj
to the investigfttions of SAViomr. The compound body is a much-
elongated cone, ordinarily six or seven inches in length, open at
one end and at the other closed and bluntly rounded oK The
little animals are placed perpendicularly to the axis of the cone, in
circles more or less irregular, whilst the posterior openings of their
body terminate in the cavity of the cona The gem, according
to the observations of Saviont, is already cloven into four animftls
even before they are bom. This is the conmiencement of the cylin-
der or cone, which may be imagined to be formed of a series of
circles or girdles of small Ascidias behind each other of increasing
size ; the thinner closed extremity of the compound body is thus
the first formed. Consult Saviont L dt. pp. 58, 206.
Sp. Pyro9oma aOanUeum Pl^aOK, 1. L PL fi, Vnyage omx Terru Auttrala,
^ 30, ^- I \—Pyromma giffonieum LisusuB, Satiovt Mim, n. PL 4,
fig. 7, PL «a, 23, Blaihv. Malac. PL 85, fig. 6, Cuv. R, Afd., 4d, iU,,
Moll. PL 133; in the Mediterranean : there is still % smaUer gpecieB in
the Bame sea in which the individual *Jiit«%ln are placed in regolar cirdes
round the cone ; Pyrot. degant LssuiUB.
Family III. Asctdice. Apertures of external covering not oppo-
site, mostly approximate. Branchial sac closed posteriorly. Animals
either single, or congregated into a common body, affixed.
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TUNICATA. 703
A. Compound Ascidians. Common body gelatinous, or gela-
tinoso-K^oriaceons, polTmorphous, often expanded, incrusting the
surface to which it adheres.
BotryUus G^ertner. Body of the animalcules not distinguished
into thorax and abdomen ; branchial aperture circular without rays.
Several clusters of animals, each of them mostly stellate.
Sp. BotryUut Sehloueri Say., Alqfonvum Sehloueri L., BctryOus tteUatus
Gmrtv., Pall. SpU. Zool. z. Tab. iv. figs. 1—5, Saviont 1. 1. Tab. xx.
fig. 5 ;—Boiryllu8 polycycltu Savignt 1. 1. Tab. iv. fig. 5, Tab. xxi. (re-
peated in Gu^iK Iconoffr,, MoUtuq. PL 35, fig. i). In this and other
species every group has a common central cavity, into which the posterior
aperture of each several animalcule opens. In other species the several
animalcules are not placed circularly round such a cavity, and the two
openings of the body are closer together. The common mass is penetrated
by canals. Here belong the species which SAyiGKT refers to his first divi-
sion of BotryUus; Milne Edwabds forms of them, with the addition of
some new species first described by him, a separate gemiB under the name
o{ £otryU<4de9.
Didemnum Sav. (and Euccdium ejusd.) Common body sessile,
incrusting, without central cavity. Animalcules divided into two
separate parts, with branchial sac separated like a thorax from the
abdomen. Anal orifice without rays. Many groups of animals
collected in a common body.
a. Branchial orifice rotundate, destitute of rays.
Eucodiwm Sav.
h. Branchial orifice with six equal rays or lobes.
DidmMwm Sav., Didenmum and Leptodinmn Milne Edw.
Sp. Didemnum candidum Sav. 1. 1. PI. nr. fig. 3, PI. xz. fig. z, Cuv. S,
AfU., MoU, PI. 129, fig. 3, &c.
Polydinvm Sav. (and Aplidium ejusd.) Common body sessile,
polymorphous, often incrusting. Many groups of animalcules col-
lected together in a common body. Animalcules divided into three
separate parts, with ovary and heart behind abdomen contained in
the last part. Branchial orifice sexradiate; anal orifice without rays.
Sp. Pclydimm conddUOum Sav. L 1. PI. iv. fig. 1, PL xvm. fig. i, Ou^Bnr
Iconogr.j MoU. PI. 35, fig. 4 ; on the coasts of the island Maoritius ; —
Apfidiwm ficm Sat., Alcycnium Jictu L., Ellis OoraUines, PI. xvn. fig.
b. B, Ac,
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704 CLASS XI.
NoU, — Here also ia to be referred sub-genuB AnMrimevtm Edw. (belter,
perhaps, Amarewnum from d/ia/>6(^, on account of the channelled canak
that permeate the oonmion body like doacce or sewers). Sp. ii«Mir. pnii-
ferum MiLKi Edw. ObtervcA, $. I. Ate. comp. PL I. fig. 3, Awior. onreMi
Milks Edw., Cut. JL Ani., id. iU., MoU, PI. 130, fig. i, Paraacidia Milhi
Edw., a new genus different from all its congeners by the octoradiate bian-
chial orifice: R. Ani, ibid. fig. 5.
Diatomus G^RTN., Diatoma Sav. (a name to be rejected as pre-
viously ascribed to a genus of Entozoa). Common body sessfle,
coriaceous, polymorphous. Several groups of animalcules. Ani-
malcules divided into three separate parts. Six radii in the branchial
and in the anal orifice.
^. JHdomut varioloiut Gartnkb, Pall. SpicU. Zoolog. Faac. x. Tab. iv.
fig. 7 a. A, &c.
SigiUina Sav.
Synoicum Phipps, Sav. Common body erect, cylindrical,
coriaceous, adhering by the base. Animalcules divided into three
parts, elongate, arranged in a circular group at the incrassated apex
of the common body. Branchial orifice with six equal rays, anal
aperture with unequal rays, three larger and three small, subihdis-
tinct.
Sp. Synoicum turgens Phipps, Voy<»ge towards the North Pole in 1773, Lon-
don, 1774, 4to, pp. 199, 900, PI. xin. fig. 3, Sat. MSm. 1. L PL m. fig. 3,
PL XV.
Diazona Sav. Common body gelatinous, sessile, orbicular,
excavated in the middle. Animalcules divided into three separate
parts, arranged in several concentric series. Six equal rays at the
orifice of the branchial and of the anal aperture.
Sp. DicoBona violacea Sat., Poiyelinum diatona Cut. JR. Ani., ^ i. Tab. XL
fig. 6, 4d. a. Tab. xiT. fig. 6, Sationt 1. L PL n. fig. 3, PL xn. The
entire system resembles a compound flower or an Actinia^ and attvns a
diameter of six inches ; the colour is a handsome Tiolet. This species has
been found in the Mediterranean on the Spanish coast.
B. Simple Ascidians, solitary or associated by gemmiferous
stolons. External integument sacciform, coriaceous or gelatinous,
with two apertures approximate. (Genus Asdiia L.)
The non-pedunculated sacciform spedes have been long known,
and were well described by Aristotle under the name of r^fivw.
De Aninu Hist. iv. Cap. 6. LnrNiEUS at first named this genus
Tethya, but confounded with it dififerent animals which alone now
keep this generic name. Baster on that account gave to this genus
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TUNICATA. 705
the name of Aacidivm (I^atuurk. UUspanningen, L bl. 97), and
LiNKiBUS, in the twelfth edition of his Syatema NaturcRy changed
this name, without any reason that I am aware of, into Ascidia.
These Aacidice or bag-pipea (zakpijpen), as Baster names them,
are always attached to other bodies, to rocks, shells, crabs, &c. Often
several individuals are united in a single group ; they never, how-
ever, form such a compound body as the preceding genera, which
are distinguished by the orifida analia being always turned towards
each other, and more or less really united, (Savigny, op, cit. p. 120),
whilst the external covering is common to all the individuals that
combine to form the group.
These animals ingurge water tiirough the branchial aperture, and
eject it chiefly by the same aperture in jets, which may serve as
a defensive means for chasing flnimalfl away that attack them.
CuviEB asserts that the expulsion of water can be performed
through the branchial aperture alone. Those writers, however, who
have observed them alive, are unanimous in their testimony that the
ejection of water is eflected through both apertures. Casus tells,
that in a large specimen of Aacidia microcoarnua he saw an opening
furnished with a membranous valve, which appeared to lead from
the branchial sac to the porua cmalis. Other writers, however, do
not speak of such an opening. On the supposition of Lister and
Milne Edwards that the branchial sac is perforated like a sieve
(see above, p. 693), the matter may be explained without difficulty.
Whether Aaddicma also ingurge water by the doacal aperture afler
a vacuum in the giU-sac has been caused by contraction, as Savigny
suspected {op, cit, p. 100), deserves further investigation.
Aaddicma live on small organic particles, which are brought with
the water into the respiratory sac and thence to the (esophagus that
opens at its bottom. Sometimes, indeed, small cnistaceans have
been found in the sac, but they would seem to have arrived there
fortuitously; for when they have been ingurged by an Aacidian they
are rather hurtful than beneficial, and in some cases even injure the
tissue of the gills.
Etbsnhardt has published observations from which it appears that the
body of Ascidians in a singular manner may change into a formleas mass,
on which other Ascidians attach themselves, and take root. Nov, Act,
Acad, 0(88, Leop. Ca/roL Vol. xi. 1813, pp. 249 — 372.
Gomp. also on these animals (besides the works of CuviER, SAViomr and
Van Benedsn already cited) Garub Beitrdge zur Anatomic und Phynologie
der Seetcheideny in Meckel's Arch, f, die Phynol, n. 18 16, s. 569—590
VOL. I. 45
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706 CLASS XI.
(the figoree rdating to this memoir were aim given, better executed, in
Pt X. of the Ncv, Act, Acad, Cm. Leap. Car, i8ax, p. 423) ;— W. SwMac
LiAT Anatomical ObiervoHona on the Tunicata, Linn, TVancxiY.iSis, pp.
527 — 555; — Philifpi Bhopakea cin neuea Cfenui der wnfacken AaadieM,
Muellrb's Arckiv, 1843, pp. 45 — 57 (which memoir containB some nokioeB
on different simple ABcidiMui found by the author near Naples).
A. External tunic ffelaiinoiu. Two orifices, either wUhovi
distinct rays, or furnished with more than /our rays.
Perophora WiEGM, Body compressed, sacciform, placed by
means of a short petiole on a creeping tabe, connecting seyeral
individuals. Branchial sac ample, produced to the posterior part of
body.
Sp. Perophora Luteri, a small animalcule living sodallj, fixed on Confena
donffoia by a creeping, cirrus-bearing pedide : found by Listkb on the
English coast, PhU, Trane,for 1834, Pt. a, pp. 378—382, PI. xi. In the
pedide, connecting the animal, a double blood-current was ofaeerved. The
name Perophora is given to this genus of Aicidia by WiBOiEAinr in hii
Meport on the Advancement of Zoology m 1834, Archiiv /. Nahurgeech, 1835,
I. s. 309.
Claveltna Sav. Branchial and cloacal pore without rays. Bran-
chial sac short, destitute of papillae, not plicated. Body oblong,
attenuate at the base, petiolate.
8p. Clavdina horealii Sav., Aeeidia davata Pall., SpieU. Zocl. x. Tab. i.
fig. 16, Sav. 1. L PI. i. f^. 3, PI. xi. fig. a. In the North and Polar seas ;
Clavel. lepadiformu Sav., Aeeidia Upadtformie O. F. Mdblleb, Zool.
Dan, Tab. 79, fig. 5, Hilki Edwa&ds Obeerv. FAee. Tab. l ^. i. Tab. 11.
fig. 1, ftc.
Sub-genus : BhopalcBa Philippl Bnmchial pore sexradiate, anal
octoradiate. Branchial sac shorty beset with papill», with transverse
8tri» subindistinct. Body elongate, davate, petiolate, with covering
pellucid.
Sp. Shop. neapoUtana Philippi 1. L Tab. iv.
PhaUusia Sav. Body sessile, mostly oblong. Branchial pore
mostly with eight or nine radii, cloacal pore sexradiate. Branchial
sac not plicated, ample, descending to the bottom of internal tonic
or reflected at the bottom.
Sp. PhaUusia mammiOata, Ateidia mammillata Oov. Mim, «. I, Aeddiet.
PI. m. J^. Ani,, td, ill., Moll. PL 115, fig. i ; in the Meditemmean. The
thick, white, gelatinous, semi-transparent external tunic presents under the
microscope large oval vesicular cells, without nuclei; this spedes atUuitf
a size of more than 4 inches, &c.
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TUNICATA. 707
B. External tunic corictceous, hard. BranchicU orifice cmd cmal
cloven irUofov/r rays. BrancJUal sac plicate,
Cynthia Sav. Body sessile.
Sp. CynUiia papillata Sav., Ascidia papUlota L., Bohadsoh de quihutd.
Anim. mar. p. 130. Tethyon eoriaeeum, Tab. z. fig. i, CuY. Mim. PI. ii.
figs. 1—3, Bang in Gu^rik Mag. de Z06L 1834, CI. v. PI. 43; Mediter. ;
— OyniKia microcogmuBf Aacidia microcotrMts Cuv., Ascidia Cuvierii Phi-
LiFFi, Guv. 1. L PI. I. figs. 1—6; the surface of the hard and wrinkled
covering is often beset with sertnlariffi, corals and conchifers ; this species
also is found in the Mediterranean Sea.
Sub-genus : Dendrodoa Mag L.
Sp. Jkndrodoa fflandaria Mao Liat I L Tab. 10, p. 547.
Boltenia Sav. Body oval, placed on a round petiole.
Sp. SoUmia fvM/ormis Sav., Vortiedla BoUeni L., Ascidia daivaUt Shaw
(not Pall.), J. E. Boltsn ^m^. a(2 C. A. LiNKi de novo qvodam Zoophytor.
ffenere, Amstelrod. 1771, 4to. c. Tab. color. ; — Boltenia or(f«ra Sav., Ascidia
pedunculata Shaw, Vortiedla ovifera L., Saviont 1. 1. PL l fig. i, B,
Blaikv. MoLacol, PI. 8, fig. 3, &c.
Subgenus : Cystingia Mac L.
Sp. OysHngia QriffitKsii Mao Lbat L L Tab. 19, pp. 540 — 545, near the
North Polar-circle in Foo^Channdf met with in Pabbt'b expedition.
Perhaps it belongs to sub-genus Boltenia legumen Lbssov, Oentur, tool.
^^ 5i %• I. pp. i49» 150.
0. Ext&mal tvmic hard, composed of separate scuteUa. Body
sessile.
Ghelyosoma SoWERBY. Body depressed, discoidal, adhering to
foreign bodies. Orifices conical, both closed by six triangular
valves.
Sp. Ohdyosoma Madeayan/um, Zoological Journal, 1819, xvii. p. 46, D.
F. Esohbioht AnaUynvisle Beshrivelse af Chdyos, Madeayamum, Vidensk.
Sdsh. naturvid. og riuUh. Shr. ix. 1843, pp. i — 16, Tab. I.
Note. — It may suffice here simply to name the genera Pedotethis
and Syphonotethis, of which the author, Gebvais, has given the
names alone {Ascidies, Diet. tmiv. cTHist. not. n. 1842, pp. 207,
208).
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CLASS XII.
CONCHIFERS {COFCHIFERAf.
With Lamarck we unite all the bivalve molluscs in a single
class, which also contains the Brtzch'opoda of CuviER, They are
acephalous molluscs, of which the mantle is always more or less,
and often entirely, cloven into two laminse, and which are covered
by a bivalve shell. Their respiratory apparatus is external, and
situated either between the mantle and the body in form of plates,
or in the substance of the mantle itself.
The intestinal canal, very various in length, is closely sur-
rounded by the other viscera. The oesophagus is short, or there
exists no oesophagus obviously distinct Ordinarily a stomach is
present, yet in Ligula the intestinal canal is almost of the same
width throughout; and in OrbicuJa also a stomachal expansion is
not apparent. Salivary glands are not present in the LameUi-
hranchiata ; in the Brachiopoda it is doubtful whether one of the
glandular masses, that surround the intestinal canal, is to be
regarded as a salivary gland'. Largely developed on the other
hand is the liver, which, as in the rest of the invertebrate animals,
receives arterial blood alone, and has no gall-bladder. Numerous
lobes, consisting either of blind sacs or of branching coecal tubes,
^ Most of the general works on this class treat also of the rest of the moUaacs, and
have been partly cited above. Here belongs especially the great work of Polt. Com-
pare besides, the article Ckmchifera of Dishatxs in Todd's CyclopcBdia of Anatomy and
Pkytiology, i. 1836, pp. 694 — 716, and B. Gabnsb, On the Anatomy of the UmdU-
hxmckiate Conchifera, Tran: af the Zool. Soe, ii. a, pp. 87 — loi, PL 18 — ao.
* In Ligula AnaHna CuviXB observed a difference of colour in these glandular
masses, which led him to consider the round white gland, situated in the middle, to be
salivaiy, the lateral, divided into many lobes, and yellow brown, to be a liver. Owes
could not perceive such difference of colour in Lingula Audd>ardii, and supposes that,
in the specimens examined by Cuyiib, it was to be ascribed to some accidental cause,
as bleaching by the spirit in which the animals were kept (Trans. Zool, Soc. I. p. 15 7)*
I must, however, observe that the specimen examined by me gave the same results as
CuviBfi has noticed, although I do not regard difference in colour as a certain proof of
difference in function. In the other genera of JBraehiopoda, whiph I have not examined,
TorebrtUula and Orbiada, Owkn found no glands except the liver.
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CONCHIFEES. 709
•
Bxurround the stomach and intestme, and pour the hile by more than
one opening into their cavity. In many LameUibranchtata there is
found, either in a blind appendage at the undermost part of the
stomach, or in the intestine itself below the stomach, a transpa-
rent elongated organ (the crystal style), on that extremity of which
that projects freely into the stomach, a small membranous cartilagi-
nous protuberance, divided into three or more irregular processes
or points, is seated ^ The use of this apparatus is not yet rightly
understood. Poli thinks the elasticity of the organ may press the
points of the protuberance towards and into the openings of the
gall-ducts, and thus moderate the influx of bile when not required ;
but such a regulator is unexampled in the animal kingdom. That
the style may effect the recoil of the foot, has been suggested by
Garner, without any accurate explanation of the mode in which
this is produced, whilst, at the same time, the free projection into
the stomach of the tricuspid protuberance remains unexplained.
We confess rather that we do not yet understand the organ,
because we can compare it with no other in the rest of the classes
of animals.
The circulation of the blood has in this class always a central
organ, a heart with a single ventricle, sometimes two hearts remote
from each other; for instance, in the Brachiopoda and in Area
amongst the LaTnellibranchicUa*. In this case, however, the two
hearts ftilfil the same oflSce on each side of the body, and are both
arterial like the single heart in the rest of the acephalous molluscs :
the blood, namely, flows from the gills to the heart, not from the
1 Poli names this protuberance aagitta trictapuj see on this snbject his celebrated
work Testae, utr. Sicil, i. IrUrod. p. 41, and the figures for example, from PhoUu
dactylus, Tab. vn. figs, g, 10, 1 1, from Tellvna planata, Tab. xiv. figs. 9, to, from
Cardium ruttumm, Tab. XYI. figs. 13, &c. The circumstance that the crystal style is
sometimes not to be found, and, as V. Siebold thinks, is developed and disappears
periodically (Lehrb. d. vergL Anai, i. s. 363, note 15), indicates an analogy with the
lapidet eancrorum (see above, p. 616) that promises, perhaps, to throw a clearer light
upon it.
* For the Brachiopoda compare Cuyibb Mimoire twr VAnimal de la Lingvla,
MSmoiret tw Ub Moll., and Owen, LeUre d M. Milne Edwabdb, Ann. d. Sc. Nat.,
3i^e S^rie, ZooL iii. 1845, PP; 3^5 — 3*Oi »l8o Huxlet Contributions to the Anatomy
of the Brachiopoda, in Proceedings of the Roy. Sac. Vol. vn. pp. 106 — 117, 1854, who
throws some doubts upon the office of the so-called hearts ; their number, also, he
states to be in some cases four. For Area see PoLi Test, utr. Sic. u. pp. 189, 183,
Tab. XXY. fig. 9.
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710 CLASS XII.
heart to the gills. When a single heart is present, it is usually
seated between the two membranous triangular auricles that have
their points turned towards the heart. From the heart arise the
arteries, which, however, do not pass from capillaries into yeins ;
the arterial blood flows, according to the interesting discoyeiy of
Milne Edwards S not through closed vessels, but in reticular inter-
spaces [lacuncB), which are emptied into larger venous sinuses, [or
rather the so-called close vessels expand, their attenuated proper
tunic being continued into these lacunae and sinuses'].
The respiratory organs are, in the Brachicpoda, situated on the
mantle, or the vascular mantle itself serves for respiration. In the
Lamelltbranchtata the gills lie as plates between the margins of
the mantle. Ordinarily two gills are present on each side. Each
gill consists of two plates, which are more remote &om each other
on the dorsal surface, and coalescent at their free outer margin.
Sometimes the triangular spaces which are thus formed in the gills
are capable of a great extension, and serve, as in AnodarUa, as tern*
porary repositories for the eggs, — brooding cavities. On each plate
numerous transverse stripes or projecting lines are seen, along
which the currents of blood pass in the gills. In Arca^ Peden^ and
8pondylu8, each of these projecting lines is changed into a free fila-
ment, and the gills thus consist here not of plates but of threads,
which, though separate, are still from their numbers placed close
together'. The gills in this condition resemble those of bony
fishes, whilst the laminated structure, which in the ZameZ2t%ran-
cJiiaia is that generally prevalent, occurs in these fishes (in Xiphias
gladiua) only as the exception. Another deviation from the ordi-
nary type of the gills is seen in some genera of LameUtbranckiaia
in the number of these organs, when on each side, instead of two,
only a single gill is present*. From the common opinion that the
^ Ann. da Se, Nat, 3i^me S^rie, in. 1845, pp. 300—304.
3 [Vid. Owen Introductixm to the Anatomy of Ter^fratula in DAVmsoK's Mono-
graph on Fossil TerebratulcBf published by the PalseoniographicAl Society, 1853, pp-
15, 16, and PI. III. fig. I. This structure of the spaces in which the blood flows was
first explained by Hunter, and exists equally in insects and crustaceans. Owxir L 1.
pp. 17, 18.]
> Meckel's System der vergl Anat. vi. 1833, P- ^- ^^ SoUnomya the gills also are
feather-shaped; seePHiLiFPi in WiEQMAVS*BArehivfarNaturge8ehickte, 1. 1835, ^ ^75-
* In Anatina and Pholadomya Soweebt the gill-plates on each side are so grown
together as to form a single gill ; Owen Lectures on the Oomp. Anat, of ths iwiert.
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COKCHIFEBS. 711
plates described are gills, Bojanus entirely recedes ; he thinks that
the only office of these organs is the custody of the eggs, whence
he does not name them gills, but brood-receptacles {BrutMUer).
Accordingly he has described an entirely different organ, not
noticed previously, as a respiratory organ*. According to his ob-
servations the venous blood, that flows back from the organs of
the body, is conducted to an elongated venous sinus situated at
the dorsal surface beneath the heart. Close to this sinus are
two spongy, dark-green or brown sacs, which are very rich in ves-
sels, and receive all the blood from the venous sinus. BojANUS
was of opinion that these organs serve for respiration, and are
lungs; from each sac arises a single vascular stem, which runs
along the two gills of its own side. From the parts which BojANUS
names lungs, come also some short vessels, which run immediately
to the heart, but the greater part of the blood that flows in them
goes to the two vascular stems of the gill-plates.
The investigations of Bojanus have made us much more accu-
rately acquainted with the circulation of the blood in LameUt-
branchtata, than what had previously been written upon it. The
mode, however, in which this anatomist explained what he observed,
is exposed to many objections. If his opinions be not adopted, it
is not clear what appellation ought to be assigned to the organs
described by Bojanus. At the present day it seems that these so-
named lungs are very commonly regarded as kidneys*. On this
supposition, however, it remains unexplained why they receive
all the venous blood of the body. This circumstance deserves
special consideration, whenever a conclusion is attempted con-
cerning the nature of these organs. Hence it Is that I offered,
now more than twenty years ago, the opinion that these so-
called lungs are venous sinuses, as much as the part that lies
between them, which Bojanus himself named sinus venostis; they
Animals, p. 383. AIbo in Lueina and Corbia Yalanoiekkes found only a single gill
on each side ; Campte$ rmdm, 9 Juin, 1845,
^ L. H. BojAKUS Sendachreiben d Mr le Chevalier G. De Cuvieb aber die Aihem- und
Krtida'uf'werkzeuge der zweiacluudigen Muscheln irubeaondere des Anodon cygneum.
Mit abbildungen, 4to. (Printed separately from Oeen'b Isis in 1820, Heft 7). PoLi
speaks in different parts of his work of this organ, under the name of visciis tettaceum,
^TnxviRAiras Zeitachr.f, Physiol, i. 1824, s. 53 ; Cajixjb Lehrbuch derZooUmie, 2te
Aufl. n. 1834, 8. 650; Y. 81EBOLD Lekrb. d, vergl. Anat, i. s. 281 — 284.
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712 CLASS XII.
are parts which correspond to the lateral venous sinnses of the
decapod crustaceans (see above, p. 605), and to the two latenl
venous hearts in the Cephalopoda dtbranchuzta^. Consequently the
two arterial stems of the gills convey venous blood, which, having
become arterial in the gills, is brought back to the heart by four
branchial veins (two on each side). The walls of these venous
sinuses may at the same time be regarded as secreting organs,
which correspond to the appendages of the veins in the Cqfhalopoda,
Each of these sinuses, moreover, is situated in a cavity or a space
with thin walls, to which the water has free access bj a fissure
which' opens close to the external opening of the genital organs.
Besides the circulation of the blood in vessels and in spaces
without proper walls, as already described, another system still of
canals or interspaces has been discovered, that is filled with
water in conchifers, as in many other invertebrate animals'. It is
probable that in the description of the circulating system of the
blood, a confusion with these canals has occasionally occurred'.
It wai^ generally supposed formerly, that all the conchifers
were of one sex, not so much bisexual, as indeed all female, there
being no other organs of propagation except ovaries\ But if
such were really the case, these animals ought not to be styled
female, but sexless. An organ for the preparation of germs covid
not, when the germ required no impregnation, be an ovaiy; the
germ which, without the influence of sperma is developed into a
new animal, ought rather to be named a detached bud than an
1 In a letter to my deceased friend NiTZSOH of Halle, Msckel's Arekivf. AnaLu,
Phydol. i8i8, 8. 503, and in the fint edition of this Handbook, n. bL 35. What
V. SiBBOLD advances as my opinion, and with which he professes to agree, that the
parts described by Bojanus correspond to the appendages of the veins in Oephalopodt,
is not mine, but indeed a later guess of Bojakus himself {Isu 1820), who in fact had
too much love of truth to conceal that his opinion respecting the respiration <^
conchifers was something of a paradox {eine in ironiscKer AnwancUung, etwat heck tuid
paradox atugetprochene Meinung),
* Dkllb Ghiajb Memcrie nUla ttoria e noUmia degli aninuUi 9ema voriAftj n.
p. 359, and foil. pp. 369, 270 ; Y. Basb in Fbokikp'b Noluen, i8a6.
* Ck)mpare on this point V. Siebold Lehrb, der vergl. AnaL i. s. 979 — 181. [^
Letdio Ud), Cyclat cornea Lam., Mueller's Ar^Mv, 1855, pp. 54 — 57, firom whoae
observations it would seem that the opinion of Delle Chiajb, that the fine pores and
canals of the water-system communicate with that of the blood, is perfectly conrect]
^ Even in the work of Debhayeb, TraiU de Oonehyliol,, begun a few yean1»ck,
this statement is found p. 384.
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CONCHIFERS. 713
egg*- We have already seen above in the case of the sea-nettles
and echinoderms, that the sexual glands {ovarian testes) might agree
entirely in position and external form, so that, without microscopic
investigation of their contents, it could not be determined whether
they were feminine or masculine, whether they served to prepare
the germ or to impregnate it. In the Brtzchtopada the ovary alone
is known hitherto, which lies behind the liver ; from the ovary the
eggs arrive at the lobes of the mantle, and cover the blood-vessels
that are distributed there^ In the LamellibranchtcUa Prevost
discovered in 1823 distinct sexes in Uh{o\ In the male sex, in
place of the ovary, a similarly formed part is found, filled with
white fluid, which swarms with seminal animalcules. Pkevost
saw neither from these nor from those that had ovaries any young
ones proceed when he kept them apart, but did when two were
placed in contiguity. These observations were confirmed some
years afterwards by others of the same kind, of Wagner, Milne
Edwards, and others, and especially by very exact investigations
of C. T. Von Siebold*. In the mean time, this interesting pecu-
. liarity is no new discovery of the last years ; and the observations
which have been alluded to tend to the confirmation rather of what
had already been observed by Leeuwenhoeck a century and a half
earlier^. The spermatozoa have a long thin tail and a conspicuously
distinct body, like a knob, which is very small and elongated.
The testes in the male individuals are situated at the same part as
the ovaries in the females ; they open at the same place, and appear
in the arrangement of their blind tubes to agree with the ovaries.
Now that it has been admitted that difference of sex exists in the
* Mubllkb's Physidoffie, n. p. 6i8.
* Cuyim could not trace the orgiuis of propagation in Lingula; OwXN describes
them very briefly in TerebraiiUa and OrhietUoy Trant, ZooL Soe, I. pp. 152 — 156, PL 21,
fig. 11^, i6r, PI. 33, fig. iiw, fig. T5f»; in these figures the eggs are represented
partly in the mantle ; there are eggs also lying on the mantle, which O. F. Mcellbb
described and figured as ovarium ramosum in Orhicula, Zool. Danic. Tab. 4, fig. 7.
[See, however, Owkn's further description of the geuerative organs in Terebratula, in
his Introduction cited above, p. 710 and Plate in. fig. i. He concludes that Ter,
Jlaveieent is diceceous.]
* De la g&iUnUion de la moule de» peirUres, MSm, de la Soc, de Phytique et d'B^itt,
Nat, de QeiUve, iii. i, pp. 11 1 and foil.
^ See these observations, illustrated by figures, in Mublleb*8 Archiv, 1837,
8. 381— 39«'
* A. Van Lbbuwenhosok Vijfde vervolg der Brieven, Delft, 1696, 4to, 95vte Mis-
sive, bk. 136—155 ; see especially bl*. 145.
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714 CLASS XII.
LcaneUibranchtataj perhaps hereafter external sexual difference
will be looked for. In AnodanUiy at least, the females may be
recognised by their more convex shells ^ However all LcmutJUr
branchtaia are not of different sex ; Peden, for instance, according
to Milne Edwards, is hermaphroditic'; also in CydaSy besides
the ovaries, two testes are present'.
The structure of the ovaries has, by the investigations of Polt,
become known in many genera of Lamellibranckiaia, The two
ovaries lie on each side of the intestinal canal and the liver, and
consist of tubes that divide into branches terminating blindly;
they have often a proper colour, ordinarily red or rose-red, by
which they are distinguished from the neighbouring parts. The
eggs pass from the ovary by an aperture situated on each side of
the foot or the abdomen, at the inside of the opening of the
vacuities in which the venous sinuses are contained. In the same
situation lie the apertures by which in the male conchifers the
sperma is evacuated. Afterwards the eggs proceed along the foot
into an opening between the foot and the inner gills, and arrive
at the canal of the inner gills, which conducts to the cloaca. From
here the eggs are carried to the lobes of the mantle, or they come
outward and are brought by the respiratory streams firom behind
into the canal of the external gills, and deposited between the
plates in the saccules of these gills, as in Unto and Anodonia*.
^ y. Subold op, cit. 8. 391 ; Kibtlanb appears to have remarked this difference
in Unio also.
* Ann.dcB Se, Nat. le Sine, ZTin. 1842, ZooloffU, pp. sat, 323, H. 10, fig. i>
* Waokkb found tpermatozoa in all the individuals of OycUu cordata which be
investigated ; Wisgmaitn'b Archiv, 1835, n. b. 218, Tab. ni. fig. 8. The presenoe of
two tettet and two ovaria in genus CtfcUu was observed by V. Sikbold, Hubllsb'b
Arekiv, 1837, s. 383, 384. [See on the genital organs of LamdUbraxkchiaUi, H. Leoazb-
DuTHiBBa Ann. de» Se. Nat,, ZoU. 4i^e Serie, Tom. n. pp. 155—348. He addi
Offrea, so much contested, to the list of the hermaphrodites. Peden, which ii vsaaltj
bisexual, has one spedes {Peden variut) unisexual, whilst Cardium, usually uniseznalt
has Oardium terratum and Card, hnigaium bisexual. In some hermaphrodites, the
sexual organs, though united in the body of the same animal, are quite distinct (Ptde^
&c.), in others they are quite confused, (Odrea).^
* The figure of PoLi Testae, u^utq. SicU. I. Tab. rx. fig. 18, gives a good ides of
these chambers formed by transverse septa, triangrular and much elongated, which have
their bases turned to the dorsal side of the gills. POLI and most observers have fouoa
the eggs in the external giUs alone, BojakT7B occasionally found some in the intenial
also. Thus the respiratory function is not prevented by these eggs ; but only parti*"/
interrupted, and since there are many conehifera in which the gills do not serve for tbe
reception of eggs, there is the less reason for refumng to these organs the fanetion of
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C0NCHIFER8. 715
Here the eggs remain for development. Since the young Anodonts
and Unios in the early period of life have a form differing from
the perfect animal, they have by some writers been regarded as
parasites that live in the gills^. It is almost needless to say that,
in the Lamdlibranchiaia, where the sexes are distinct, there is no
copulation. The eggs must leave the ovarium previously to im-
pregnation.
The nervous system was by Polt, whose investigations have
contributed so much to the knowledge of the structure of con-
chifers, described as aqui-vascular system, since the wide neurilema
admitted impletion with quicksilver. Mangili contested this
opinion. He observed four nervous ganglia in Anodonta^ two at
the side of the mouth, one, the largest of all, in the foot, and a
fourth under the posterior occludent muscle of the shell, from which
ganglion the nerves for the gills and the mantle arise. All these
ganglia are mutually connected by nerve-filaments'. This may be
regarded as the first exposition, in some measure accurate, of the
nervous system in this class. As the rule, three central nerve-
masses may be admitted in the Lamellibranchiatay each consisting
of two lateral ganglia, of which, however, the first two alone are
constantly distinct from each other. These lie near the mouth, and
are connected by a longer or shorter transverse arched filament, a
commissure which runs in front of the mouth. On each side there
arises from these ganglia a long nerve-cord that runs to the poste-
rior nerve-mass, the ganglia abdomtnalta, often connected in a single
ganglion, and thus forms a wide ring which may be compared to
the oesophageal ring in other invertebrate animals. The second
reflpiration, so generally attributed to them ; to say nothing of the male indiyiduals in
which the temporary function for the gills of brooding organs entirely falls to the
ground.
^ J. Bathkb (Shritfter of nat. SeUhahet, iv. i, 1797, pp. 166, 167, and after him
L. Jaoobson KongeL Damke Videntkab, SeUkaJbs NaturM, og McOhem, Afhandlinger,
m. 1828, pp. ^6S, 269), described the animal living in the gills as a distinct species
under the name of Glochidium. The observations of Y. Baeb (Mecksl's Archiv^
1830, 8. 331) and especially of Cabub (Neue UfUertuehungen Hb. d, ErUmckelungigtsck.
wucrer FlvMnm$ehel, Leipsig, 1832, 4to, printed separately from the Act, Acad, Cast,
Zeop. Cofid. zvi. i) have now cleared this matter up. The young animals have
a triangular bivalve shell, which gapes open, and has at the outer margins an
appendage or hook turned inwards and furnished with small spines. Gomp. also
QUATBBFAGEB, Ann, dcB Sc, NcU, sec. S^rie, v. 1836, Zool, pp. 321 — 336, PI. la.
' Nuove JUeherche zootomMchc sopra alcuni specie di Conckiglie hivcUvi, Milano,
1804 ; translated into German in Bsil's Archiv, ix. s. 113—220, Taf. xh.
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716 CLASS XII.
nerve-mass, the par pedalt^ also connected with the first ganglia by
two shorter strings, is, in those genera that have no foot, feebly
developed, or according to Garner, even absent. The nerve-
ganglia are osoallj distinguished bj a red or orange colonr^. The
nervous system of the Brachiopoda is not yet perfectly known, but
seems to be formed after a different typu8^.
Amongst the parts which may be regarded as the seats of senses,
the conical cirri are first to be noticed, which -are present sometimes
along the entire margin of the mantle of the LamellibrancAiata, or on
some parts of the mantle, at the opening of the siphon for example,
and which in the Brachiopoda are changed for long, stiflF, glistering
hairs. The mouth, moreover, in the Lamellibranchiata is surrounded
by two pairs of transversely striped organs of touch of considerable
size, triangular, elongated or oval laminse, which some* consider to
be accessory gills, from their external resemblance to these organs.
In the Brachicpoda there are two long arms, beset pectinately with
filaments like a finnge, situated at the side of the mouth, where
they are rolled up in a spiral form, and concealed within the shell.
Organs of vision have, of late years, been shewn by microscopic
investigation to be present in Pecten, Spondylus, and many other
genera amongst the Lamellibranchiata^ as green, red-yellow, or
brown tubers, shining brightly, and often set upon a small pedicle
at the margin of the mantle. In different species more than a hun-
dred such eyes have been counted*.
^ Compare R. Guutkb On the nervoiutyMem ofMoUutcouM AnimaU, Tnuu. of Linn,
Soe. XVII. 1835, PP- 485—488, PI. 14; Blahchahd Ol>$ervation» swrle SyaUme nenvuz
da MoUusqnea lamdlibranehei, Ann. des Se, not., 3i^me S^rie, in. 1845, PP* 3^' — 34Pf
PI. 12. Here may be found a copious historical review of this subject. The two small
nerve-ganglia that lie near the oval ganglia in Ottrea (Brandt and Katsebtjeq Median,
Zool. II. s. 340, 341), Blanchard regards as answering to the par pedale (here want-
ing according to Garner). Nervous branches that arise from the lateral ccmmiissunl
string of the first and hindmost pair, on which in some species {SoUUf Area) even gan-
gliform swellings are observed, correspond, so it appears, to the sympathetic nervons
system of articulate animals, especially to the lateral portions of it in the crays, which
in like manner arise from the collar round the neck (see above, p. 618). Those UaneQi-
branchiates, which are provided with a tubular prolongation of the mantle, have often
between the muscles, that retract the tube, small nerve-ganglia in addition.
■ CuviSR Mim, 9ur la LxnguU, p. 8, speaks very indecisively on this point; Owiif
describes a nerve-ganglion between the basal pieces of the two arms, and two others at
the side of the mouth. Tram, of Zool. Soe. I. p. 156. [On the nervous system oiTcrt-
hratula see Owbn Introduct. &c. pp. 11, 13 (cited above, p. 710).]
' PoLi spoke of such eyes tmaroffdino colore coruMsanteM, which are situated on the
larger cirri of the border of the mantle in f^ponflylus (ii. p. 107) and Pccten Jacobtgvf
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C0NCHIFER8. 717
As auditory organ, Sieeold regards a part met with by him in
CydaSy Anodonta, UntOy Mya, Gardium, Tdlina, &c., an organ
situated in front of and near to the nerve-mass of the foot. Here
namely on both sides he found a vesicle in which a flat, round,
transparent nucleus is in free motion. The nucleus is a concrement
comparable to the lapilli in the auditory sac of the bony fishes^.
Neither of this enigmatical organ nor of eyes has any vestige been
met with in the Brachiopoda.
The motions of these animals are very simple. Many Lamelli"
branchiata, and all the Brachiopods, are fixed to their places, and
are not able to move from them. Otlier conchifers have a springing
motion by means of the footy a name given to a production of the
abdomen which is muscular and very firm, possesses great con-
tractility and irritability, and may assume very difierent forms*.
By means of it bivalves move at the bottom of the water in which
they live.
The geographic distribution of conchifers deserves a more
special investigation than has hitherto been bestowed upon it.
All conchifers live in water, part of them in fresh water, but
the greatest part in the sea. Amongst the genera that live in
the sea are some of which species are met with in all parts of
the world, as genera Solen, Myay Anattnay Mactray Tellina, Lu'
ctruiy DonaXy VenuSy Cardiumy Arcay Pectunculiis, Mytilu3y PecteUy
Ostrea, It is however far from the fact that all these genera
are found in like manner in difierent seas ; of the genera VenibSy
Cardiumy Arcay Oatreay the species are much more numerous in
the Indian Sea and the South Pacific, than in seas of the northern
hemisphere. Olycimerts appears to be a northern form, of which
(ib. p. 153); they are particularly large and conspicuous in the species last named
(Tab. 17, figs. 5, 14); PoLi was not able to investigate their internal structure; and
the later writers on the molluscs neglected this peculiarity altogether. Only vdthin
the last few years has it been adequately illustrated by Gbuba (Muellbb's Archiv,
1840, s. 14, Taf. III. figs. 1, 3), Ebohk (ib. s. 381 — 386, Taf. XI. fig. 16) and Will
(Fbobiep's Neue Noiizen, xziz. Bd. January, 1844, No. 633, 623).
* C. Th. Von Siebold Ueher ein rdtkselhaftiges Organ einiger Birndven, Muellsb's
Archiv, 1838, s. 49—54 (transferred to Ann, des Sc. not., sec. S^rie, x. Zool. p. 311),
and WiEQMAiffN's Archiv, 1841, Ueb, d, Gehdrorgan der MdUushen, s. 148 and foil,
Ann, d. Sc. not., sec. S^rie, xix. p. 193, PI. 1 B, fig. i. [Also Lbydig Ueber Cyclai
cornea, Muelleb'b Archiv, 1855, pp. 51, 5a. PI. vi. fig. x8.]
' See Poli, i. Introd. p. 37.
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718 CLASS XIT.
genuB the only species hitherto known, Qlycimeris siliqua, is
found very abundantly on the great bank of Newfoundland. More
numerous, on the other hand, are the forms which occur excln-
sively in warm seas or in the southern hemisphere, or of which
one or only a few species are met with in the Medit^raaean
or the Red Sea. We mention here the genera GrcLaacjUella^ Tri-
dacna, Hippopua^ MaUeuSy Avicula, Meleagrina, Spondyltis (and
the genus Plicatula united with it), VuUeUay Placuna. The fre-
quent occurrence of well-preserved remains of this class in differ-
ent mountain-strata affords an assistance of the highest interest to
the investigations of geologists towards a true knowledge and
distinction of those strata. Of some genera the extinct species are
much more numerous than those now living ; some genera which
formerly peopled the sea have entirely disappeared in the present
period of the history of the earth. It is remarkable that of the
genera now living and which also count extinct species, whenever
these last are veiy numerous, by far the most of the living species are
now met with either exclusively, or principally, in the Indian ocean
and in the southern Pacific. Of the genus Trigonta, a single living
species alone is now known, and this is found in the South Sea at
New Holland, whilst numerous fossil species belong to it, especially
from the Jura- and chalk-formations. The genera SporidyluSy Lima,
Avtcula, Crassatella, Area and others, tend also to prove the same.
Regarding the fresh-water conchifers, we would call attention
to the great number of species of the genus Unioy which occur in
the western hemisphere, especially in North America.
The Brachiopoda in their geological and geographical distribu-
tion offer much that is peculiar. Of Orhicula and Terfhrcutula
species occur both in the north and in the south ; species are known
from the Indian ocean and from the Mediterranean, from the South
Sea on the west coast of America, and from the North Sea. At
the same time of these genera the species in the southern seas are
more numerous, and the few species of Lingula are all from the
southern hemisphere. Extinct species of TerebrcUula are uncom-
monly numerous, and occur in very old as well as in more recent
formations. Thus the Brachiopoda make one of the few fonnB
which are restricted to no limited period of the history of the
earth, and have survived its various catastrophes.
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SYSTEMATIC
ARRANGEMENT OF CONCHIFERA.
CLASS XII.
CONCHIFERA.
Animals without distinct head, covered on each side with
a lamina of mantle. Shells two, incumbent on mantle. Heart
aortic. Respiration branchial, with branchiae mostly lamellose,
running out near the margins of body on each side, covered by
mantle; in some the mantle itself discharging the oflSce of
branchiae. Some hermaphrodite, others with sex distinct: all
aquatic.
Order I. Palliohranchiata or Brachicpoda.
Body depressed, covered with mantle, bilobed, open. Branchiae
grown to the mantle or not distinct from mantle. Heart double,
arterial. Two long arms convoluted in form of a spiral, armed
with cirri or cilia. Mouth simple at the base of arms. Shell
bivalve, affixed, sessile or petiolate, adhering to mantle by several
oblique muscles.
Family I. Brachiopoda. Characters those of the order.
On this order and fiunily may be compared : —
CuvnsB Mhn. vwr V Animal de la LingtUa, Ann, du Mvs. l 1803,
pp. 69 — 80, PL V. reprinted in his M^movrea sfwr VHiat, et VAnat,
des Mollusq, No. 21.
Bboderip Desoriptiona of 807ne new species of Brachiopoda, Tra/ns.
Zool. Soc, L pp. Ul— U4.
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720 ciJLSS XII.
Owen On the Anatomy of the Brachtopoda; ib, pp. 145 — 164,
PL 22, 23 (both these memoirs are translated and reprinted in the
Ann. dea Sc Nat., sec. S^rie, in. 1835, pp 26 — 30 ; 52 — ^77).
R Owen Lettre d M. Milne Edwabds sitr VappareU de la Circur
hUion chez lea MoUuaqu^a de la classe des Brctchiopodes, Ann. des Sc
Nat., 3iSme S6rie, ni. 1845, ZooL pp 315—320, PL 4.
W. Kino Eemarke an certain Genera belonging to the Class FaUuh
branchiata, Annals of Natur. History, xviiL 1846, pp.26 — 42;
83—94.
Owen On the Anatomy o/ Terebratula, Introdttction to the British
Fossil Brachiopoda by Davidson. Printed for the Palseontographical
Soc 1854, pp 4—22.
The chief characteristic of this division consists in the respira-
tory organs. In LigtUa, on each lobe of the mantle on the inside,
two main trunks are seen that ran longitudinally and return the
blood to the heart, which is double, as we have seen above, at the
part where the two are most remote, and converge obliquely to the
free margin of the lobe of the mantle in the form of an inverted Y.
Lateral and parallel branches, that make a right angle with the
trunk, form an elegant pectinated design on the lanunse of the
mantle. It is these vascular divisions which compose the respiratory
organs ; the gills are, therefore, attached to the mantle, or rather the
mantle itself is the seat of respiration. From this disposition that
which presents itself in Orbicula and Terebratfda differs more in
appearance than in reality. The respiratory function is not limited
here, as in LigtUa, to a part of the mantle, but is distributed over
the whole of it, whilst two vascular stems on one lobe and four on
the other are formed from numerous branches, and bring back the
arterial blood to the double heart (Owen Transact. ZooL Soc i.
pp 147, 148, 154). The two hearts do not receive the blood
immediately, however, in these molluscs, from the stems of the
branchial veins, but these last pour it into a sintLS, from which it is
taken up by the free and wide opening of the two auricles of the
heart (Ann. des Sc NaL 3ieme sierie, in. pp 316 — 319).
A second character of this order is afforded by the two so-named
arms, which lie rolled up with their convex side outwards near the
mouth, and are margined with cirri or filaments. In TerebraJtida,
where these filaments are long and fine, these two arms thus acquire
the external appearance of gills, and have accordingly been so named
by earlier writers, ex. gr. Pallas MisceU. ZooL 1766, p. 182,
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CONCHIFEBA. 72t
GBUin>LEB Nat/wrfoTicher^ u. 1774, pp. 82, 83. The anns have
in their axis a tendinouB canal closed at the extremities, which,
according to Owen, is filled internally with a fluid. Muscular fibres
which surround the canal force, in his opinion, this fluid to the outer
extremity, and thus cause the arms to unfold.
In the third place the Brachiapoda are characterised by the
absence of an elastic ligament, which in the Lamellibranchiates
exists at the hinge of the shells and works antagonistically with the
adductor muscles. Thus the shell is opened here by the arms,
or also, in TerebrcUtdaf by the elasticity of a composite apparatus of
thin calcareous loops which are attached within, to the surfiace of
the lesser imperforate shell Hence arises a more complex dispo-
sition of the muscular system, which is formed of different oblique
bundles in place of the single or double adductor muscle found in,
the Lamellibranchiates.
I. Shell hingeless.
lAngula Brug., Lam. Shell subequivalve, flattened, oblong,
thin, gaping at both ends, aflSxed by a peduncle fleshy, cylindrical,
hollow internally.
Sp. Lingula anatina Lam., Blainy. PI. 51, fig. $, Gu^in Iconogr., MoU,
Fl. 36, fig. I ; from tbe East Indian Ocean. Fonnerly detached shellfl
alone of this animal were known ; such a single shell is figured by RuM-
PHius Amb, RariteUk, Tab. ZL. fig. L. LiKNiiUS named such an one
Patella vnguia. The first account of two shells connected together was
given by Gheukitz {NoAwrfoncker, xxii. 1787, pp. 13— 3a, Tab. m.).
Of late years a few other living species of this genus have been disoovered.
Compare Bbodkbip 1. 1. and on the anatomy of Linffula, besides Cuyibb
and OwKK, as cited above, also C. Yoot Neue DeiikichrifUn der dUg,
Schweizer QeadUchaftf, d, gea, Naturms$, vii. Nenchatel, 1845, ^"^ ^ ^^f*
Fossil spedes also of this genus are found, especially in the Silurian
formation, also in the mountain-Ume. In the Mutchd-kalk and hunter Sand^
atein Lingula tenuimma is found, Bbonn Leth. geogn. Tab. ZIL ^. 6 B.
Orbtcula Cuv., Lam. Shell inequivalve, orbicular. Affixed
valve plane, cloven in the disc ; superior valve conical.
Sp. Orbicula Norwegica Lam., Patella anomala Mukll., Zod. Danic. Tab.
v., Blaimt. Malacol. PI. 55, fig. 5 ; attached to rocks and shells in the
North Sea; — Orbic. lameUoea Bbodeb., Troate, Zool Soc. Tab. 23, fig. 2, on
the coast of Peru, &c.
Ncte. — Genus DUcina Lam. to be abolished, not being distinct from
Orhietda; see G. B. Sowbrbt Trans. Linn, Soe. xm. p. 473.
VOL. I. 46
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722 CLABS XII.
Crania Betzius, Bruo. Shell inequiTalye, suWbicolai.
Affixed valve plane, with four muscular impressions intemallj,
the two middle almost confluent to form one.
8p. Crania ringem HoimroH., Awmia tvMneda Pou, TOL «er. SiaL n.
Tab. 30, BLAJory. Malac, PL 59, fig. a ; Medhemneui. Tbii ^wciei ii
oomiiionly oonfoimded with Anomia cnmiolaint L.j which b fosnl, fike
many other species, firom the chalk-foxmation.
Compare A. J. Rnzius Crania oder Todteniopfi-MvMM, Sdirifta da
Berliner ChteUick, fuOwfortekender Freumde, Bd.11. 8.66, Sowikbt Jnnif.
Linn, 8oe, xm. pp. 469^-479, Tabw 16.
II. Shell furnished with hinge.
Calceola Lam. Fossil genus [of the Devonian period, Davidson
Introdtict. &c. p. 121].
Thecidea Defr.
Terebratula Lhwtd, Brug. (species of Anomia L.). Shell
inequivalve, subtrigonal. Apex of larger (ventral) valve perforated
by a round aperture transmitting a short affixed peduncle. Hioge
with two teeth. Internally two calcareous branches, slender,
arcuate, rising from the disc of the lesser (dorsal) valve, forming
the support of the arms.
[The ^(8re6rah<Zu2eBhave been subdivided, in the interest of Geology,
into genera and Bub-genera, chiefly according to the extent and form
of the calcareous appendage. In the true Terehrattda the appendage,
variable in length, is formed of two rihand-shaped lamellaB fixed to
the oniral base alone, and more or lees folded back upon itself
Tertir<Uulina D*Obbio. The crural processes united to fonn
a calcareous band behind the mouth, and the reflected loop always
in front of the moutL]
Sp. TerAraltMla (Ter^braiMnti) eap^i mrpenUt Luc., Anomia caput terpa^
L., BneyeL mUh., Ver§, PL 946, fig. 7, Bluky. MalacoL PL 52, fig. 6; in
the AtUntio ooean, the North Sea, the Soottiah ooast.
[Terebraiella DObbio. DifiTers from Terdfratula by its loopbeiog
doubly attached ; the lamells proceeding from the crural base, befors
attaining their greatest length, sending off a flat horizontal process
likewise affixed to a more or less elevated longitudinal septum.]
Sp. Terebratula (Terebraiella) dorwta Lah., Blaiky. MalucoL PL 51,^- h
from the Btraita of MageUan, &c.
[Megerlia King, Kra/usia Davidson, Morriaia Dav., Maga^
SowERBY, Bauehardia Dav., Agriopi Delongschamps, Megathyris
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CONCHIFERA. 723
D*ORa, RhyneonMa Fischeb (anatomy by Owen Zool. Trans,
YoL L pt 2). On all the genera of this division see especiallj
Davidsok on the dassiBcation of recent Brckchiopoday Arm. a/nd
Mag, of Nat, Hist, new ser. ix. 1851, pp. 361—377.]
Much more numerous than the recent speciei are the foasi], amongst
which we cite that alone, on account of its peculiarly symmetrical form,
which Fabio Ck)LONHA likened to two mussels joined together: Terebrahda
diMc&tdea IjAM,, TerOratula diphya Coh,, VOF BuoH Encyd, miih,, Vers,
PL 340, fig. 4, from the chalk-formation^ especially found in Italy.
Ciomp. L. Voir BuoH Udter Terdnrateln mit evnem, Veraueh tie su dauifi-
drtn und zu heackmben, vmI 3 lUho^. To^dn, Berlin, 1834, 4to. (Ahhandl.
der Ahad. der Witsensck. in Berlin, PhyHk. Klaate 1833).
Spirifer Sowerby, DeUhyris Daul, Trigonotreta Koenio, Bbonk.
Foramen of larger Talve triangular, marginal Supports of arms
rolled spirally.
Gomp. SowxBBT Linn. TramaCt, xn. p. 5T4, fig. 9; — L. Yov BnoH
Ueber Ddthyria oder Bpvrifer und Orihis, nUt 1 Utkogr. Taf, Berlin, 1837.
4to. (Abh. der Akad. in JBerlin, Physik, Kl. 1836).
The shells are usually m«ch extended in the breadth, as, Spirifer tpe*
eioius Bbokn, Leth, geogn. Tab. n. fig. 15.
All the species belong to the older, in great part to the transition-forma-
tions. In the Liat or lowest oolitic strata, the last species of this genus is
found, Spirifer WcUcoUii, which is thus met with no more amongst the
remains of living creatures.
y'- Produdua Sowerby, Strophomma Rafin., Beonn, Leptena
^ Dalm. Hinge linear, straight. Shell ineqtdvalve, alate, dmost
semiorbicular, with larger valve gibbous, imperforate.
'^^ An entirely extinct genus from the oldest periods of the history of our
r>> earth. Desha yes thinks that these oonchifera were not attached, but
^.., moved freely like most of the lamellibranchiates. Sp. Productus tuhtUiferua
^. Besh., Strephomena acvleaia Bbokit, Zethcea geogn. Tab. m. fig. i, &c.
^^^ Some writers distinguish Strophomena as a distinct genus from Produdua,
See on this and other genera which we here omit, W. Kino in Awn. and
« , Magaz, of Nat Hiit, Vol. x8, 1846, pp. 26 — 49 and pp. 83—94, and on
I p. Producttu L. Von Buoh in Ahhandl, der Akad, in Berlin, Phynk, Klaue,
1 84 1. The admirable Monograph of Davidbon, On the British Oolitic and
Liamc Braddopoda, with an introduction on the daesification, and a chap-
if' ter by Cabpentsb, On ike intimate Hrwtwre of the ihdU of Brachiopoda,
r;i> besides that of Owen, already cited on the Anatomy, were published for
^,^; the Pabeontographical Society of London in the years 1851, 1859, 1854,
'' ^ and are especially referred to in this division.
Order II. Laviellibranchiaia,
Body compressed, covered on both sides by a lamina of the
mantle. Branchise at the sides of the body placed under each lobe
^" 46—2
r-i'-
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724 . CLASS XII.
of mantle, mostly lamellose, two on each side, more rarely pectinate,
composed of thin* fimbriae or lacinias crowded together. Four lameUi-
form tentacles in by far the most, oval or triangular, placed in paiis
at the sides of mouth. Two lateral shells incumbent on mantle,
conjoined by an elastic ligament at the back.
On the internal structure of these lamellibranchiate oonchifen,
what was needful has been already said above. The beard-threads,
named byssua, still require a short notice. At the foot of the
oonchifers (see above, p. 717) there is often situated a bundle of haiis
or filaments to which the name of hyssua has been given ; sometimes
these filaments are united into a common mass ; they are attached
in a cavity at the base of the foot by a peculiar substance secreted
there (conjunctive matter). Prom the cavity a furrow nms to
nearly the extremity of the foot, and along this furrow is situated
the glandular tissue by which the bt/sstu is secreted. R£AU)n7B
maintained that these homy threads were spun just like the web of
caterpillars and the nets of spiders. With the assistance of the
muscular foot these threads are directed to determinate situations
and caused to adhere there firmly by their broad disciform extremi-
ties ; they grow again if out o£
Gomp. A. MuBLLKB de Byuo Acephalonm Aceed. Tab. n. Befrolini,
1836, 4to, and in Wieomann*s ArMv, 1837, s. i — 39, Tab. I. u. Varioui
opinions of writen on these filaments were noticed in the first edition of
this Handbook, 11. biz. 37.
Several conchifers, which are not attached by (y^^u^filaments,
grow fast by one of the two valves to rocks or different substances
fomid in the sea. That some of them move freely and get over the
ground with their foot has been already noticed (p. 717). The two
valves of the shell are separated from each other by the elastic
ligament situated at the hinge behind the point (p. 682). White
compact muscular fibres, which run transversely from one valve to
the other, close the shell on the other hand. These bundles leave
impressions on the interior of the shell, two on each valve, the one
before, the other behind, if, as in most of these animals, two adductor
muscles be present ; if, on the contrary, as in the oyster, there be
only one, then only a single muscular impression is to be seen on
each valve, situated nearly in the middle.
PoLi made many experiments with regard to the foroe of these mitfcles
in living conchifen. Thus he states, for instance, that the adductor fDn8<^^
in the animal of the Noah*8 shell {PechmcvZus) can oyercome the resisttfice
of 5olbfl.y and that more than 70 lbs. are required to tear them dtm^er.
The shell weighed only one pound, the animal its^f only 4 ounces.
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CONCHIFERA. 725
A. Mimomyartd. Muscular impression single, subcentral. In-
ternal ligament received in a cardinal pit, partially visible on the
outside in some, mostly included.
The single adductor muBcle here corresponds to the posterior adductor of
those conchifera that have two of them. All the ringle-muaded oonchifers
live in the sea.
Family II. Ostracea. Mantle open. Foot none or a small
rudiment of foot, not byssiferous. Shell irregular, lamellose.
Anomia Brug. (Species of genus Anomia L.). Shell inequi-
valve, thin, one valve flat, perforate or emarginate towards the
point, the other larger, more gibbous at the base. Animal {Echion
PoLi) with foot small, tentacles at mouth none, margin of mantle
cirriferous, adhering to marine bodies by a muscular cord perforat-
ing the shell and inserted into the calcareous cover.
The name Anomia was first employed by F. Columna {DePwrpu/ray
KomsB, 1616) and given to those oonchifera which are now named
TerdiToivla. LiNN^fiUS united with these some very different oon-
chifers also under the name of AnomMt,^ although what he says in the
description of the characters of this animal (Sys, N<U,y ed.12, l p. 1 150)
applies to jTere^ro^ti^ alone. BBnouiKBE(J^7u;^cZ.97i^^A., Vera, i. p. 70)
was the first who gave the name of Anomia to the present genus
and placed it in the neighbourhood of the oysters. From these,
however, it difiers by many essential characters, so that Deshayes
has placed it even in a separate family. There are properly three
muscles of which the impressions are seen on the convex valve, but
on the flat valve only a singular muscular impression appears. This
impression is that of the muscle which corresponds with the central
depressor of the oyster ; the two other muscles, which are attached
to the convex valve, go through the aperture of the flat valve and
fix themselves to the calcareous cover, which is often very hard,
whence it is named by several writers a little bone (ossiciUum). By
that cover the shell is attached to other conchifera or to rocks. See
Deshayes DicL Univ. d^Hiet Nat i. 1841, pp. 557 — 559, and the
figures of the three muscles in Pou Testae, uf/r, SidL Tab. 30, fig.
1, and in Cuv. E. Anl, id ill, Moll. PL 79.
Sp. Anomia Ephippium L., Listbb Conch. 104, Gheuk. Tab. 76, figs. 699,
693, Blainv. Malacol. PI. 59, fig. 31, Cuv. R. Ani., 4d, %U., Moll. PL 79;
Mediterranean, Atlantic; this spedeB i§ very flat, the shell is mostly of a
whitish colour ;—iitiont. eepa L., PoLl Tab. 30, figs, i, 8, is more convex,
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726 GLASS XII.
mauJlBt, roddish-puzpley and is foand in tlie aame seas. Tlie tpedes of
Anomia are difficult to chsrscterise, and sre not yet sufficiently dktuh
gnished. The fossil species occor especially in tertisiy farmatioiii.
PlcLCunaTiomia Bbode&
Comp. Proceedings of ike Zool, Soe. qf London, 183a, pp. ft8, 39.
Placuna Brug. (Species of Anomia L.}. Shell free, sabeqni-
Talve, flattened, thin, lamellose. Hinge interior, with two linear,
divaricate crests, converging in the apex in one valve, received in
two similar grooves in the other valve. Animal nnknown.
%>. Plaetma jptocewto, Anomia plaeenia L., Bujinr. Malae. n. 60, fig. 3,
GuiBiir Iconogr., MolL PL 37, fig. 7, Indian Sea ;— P^acfUM mUs Lax.,
CHiMif. Tikb. 79, fig. 714, HouTTUTir Nat. JHiM, L 15, PL 119, Ac. (under
the name of Ostrea qpMppium), &c.
Carolia Caktrains.
Comp. Bulletin de VAead. royale dee 8c.de BrtuuUee, v. X858, pp. in
— 113.
Ostrea L. (in part), BnUQ. Shell adhering, inequivalve, lamel-
lose, irregular, with left valve inferior, aflixed, and right superior,
plane, thinner. Hinge edentulous, a little cardinal pit receiving the
ligament Animal {Peloria PoLi) with body compressed, foot none,
margins of mantle thick, ftimished with a double row of short
fimbrise.
Sp. Oehnea ednlie L., Babtib NaiuwHL Uitap. X. pp. 71--81 ; Bladit.
Malaeol, PL 60, fig. i, Poli Teeiae. utr. SicU. n. Tab. 39, fig. i, Ciir. iZ.
Ant., id, UL, Moll. PL 73, fig. i ; Ojrter, (Eeter, ffvitre, Aneler; in the
North Sea» Mediterranean^ &c. ; on the sea-banka at the Helder and on
the TezeL This well-known speciee difien much in aiae and in the form of
the shell, as will appear at onoe from a comparison of the figures cited.
The oysters fished on the oyster-banks of Zealand become laige in a shorter
time than the English, and have flatter and less hard sheUs. Compare od
Oyster-banks, bemdes Babtib loe. cit. CABBomno. in Magaein deZod, par
Gu^iK, 1845.
Oetrea vifyiniana List., Gmsl., Listeb Oonek. Tab. too, ^. 341 '^'^'
aoi, fig. 35, JBncyd. miikod,, Vere. PL 79, figs. 1—5 ; frt>m the coast of
N. America, differs by its greatly elongated form, and attains the remark-
able length of 2 decimeters (nearly 8"). — ^By its maigin folded in a singolar
manner at acute angles and its yiolet-red colour, is distinguished Oitrai
erieta gaUi Chbmn., MytUue oriMa galU L., Rumfh. Amb. JRarUeUk. Tab.
47, fig. D, Blainv. Malac, PL 60, fig. 1, GuiBiN Iconogr., MdL PL 44»
fig. 7.
There are many fossil species of this genus both in the seoondaiy aod
tertiary formations. Comp. Lamabck Awn. dn liue. Tin. pp. 159^'^'
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CONCHIFERA. 727
ZIT. PL 90— 93, 93, figs. I, 9. In the nuyunt of St Peter is found Oitrea
larva of which the elongated crooked shell with folded margins has some
resemblanoe to the larra of an insect.
In some spedes the inferior shell, when the individuals are older, is
curved upwards at the point and very thick. Of these Lamabok formed
his genus Gryphaa. (Sp. Qryphcea anffuUOa Ljlm. from the coast of Peru.)
These forms in the fossil state especially abound in the Oolite and Lias
formations (as Qryphaa arcwUa Lam., Blaikv. Malae. PL 59, fig. 4,
Bbonn Lethaa, Tab. 19, figs. ia,h); hence the name (TrKpAtfo-Ztmestone,
in geological works. In Exogyra Sat, Sow. the point is not bent upwards,
but sideways. Species of this form occur especially in the chalk-formations.
Family III. Pectinidea. Mantle open with margin thick,
tentaculate, often ocelliferous. Foot small. Shell regular or sub-
regular, solid, often marked "by longitudinal ribs, mostly auriculate.
8p<mdyhi8 L. Shell inequiyalve, adherent, mostly auriculate,
Tude or muricate. Hinge with two strong teeth in each valve and
pit of ligament intermediate. Animal {ArgiLs PoLi) with body
compressed and margin of mantle cirrate and ocelliferous. Foot
small, disciform, supplied with a middle cylindrical filament sus-
taining an oval capitulum. Adductor muscle very large, central.
Sub-genUs : PliccUula Lam. Shell inauriculate, plicate ; inferior
valve without ertemal cardinal area.
1^ S^pandyhu pUeatut L., PUccEhda ramota Lam., Chimn. Conekyl, Tab.
4l> figB- 479f 4So;—PUeatula erittata Lam., BLAnrv. Malae, PL 62, fig. 2,
Gu^nr leonogr,, McU. PL 15, fig. 9, &c. There are also fossil species of
thisfonn.
Sub-genus : Spwidylus Lail Shell mostly auriculate. Inferior
valve produced beyond the hinge, area in superior trigonal, flat,
mostly divided by a median furrow.
8p. SpondyluB Cfcederopui L., Pou Tet^. vtr. SieU. n. Tab. ii, figs. 90, 11,
Tab. ai, Sncyd. nUth,, Vert, PL 190, fig. i; in the Mediterranean;—
Spcndyl, americanuB Lam., Guia. Iconogr,, Moll, PL 2$, fig. 8, &c.
Fossil spedes of Spondylug are found in the chalk-formation. To Sp&i^
df^, moreover, some fossil genera, whose characters are of no sufficient
Talue, are referred by Dbshatss, as Podopdt Lam., PackytcB Dunu,
JHanchora Sowbbbt. Comp. Ann, de$ Sc. naiwr, xv. iSaS, pp. 497 — 434,
and the second edition of Lamabok Jlitt. fuU, des AfU, i. vtrUhr, Tn. pp.
195—198.
Etnnttea Defr. Shell irregular, subovate or orbicular, auriculate,
aflSxed. Hinge edentulous, pit of ligament narrow, deeply cut
out.
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728 CLASS XII.
Comp. DiFBAFOi Diet. de$ Se. natur, XXIL i8«i, pp. 169, n^i
Debhatis Diet, dcM. d^HiM, ntU, Tin. pp. aoo, 101. This genue itudi
as in the middle between SpondyUu and Peeten. Besides the fosml ipeciet
upon which Defbance has founded it {Hinniles C<frU9ii, Blaihy. Mdae.
PL 61, fig. i), DB8HATB8 refers to it Peelen nnuonu Lax., Oj<rai<MMta
GXEL.
Pecien Brug. (Species of Ostrea L.). Shell free, inequiTalve,
mostly ribbed with longitudinal rays, auriculate, with apical margin
straight and apices contiguons. Hinge edentulous, with small
cardinal pit internal, receiving the ligament. Animal {ArgusTou)
orbicular, with mantle margined by cirri and ocelliferous tentacles,
the foot small, sometimes byssiferous.
Bp. Peeten Jacobonu Lav.. Otlrea Jacdbcea L., Pou Tett, ttfr. SieO. Tab. 97,
figs. I, 2 ; Blainy. Malaeol, PI. 60, fig. 4 ;—St Jamet* tkeU, Meditemneu ;
the shell is with us often used for stewing oysters, but the animal itself is,
when roasted, veiy agreeable aocording to Poll This species also is found
fossil in newer tertiary formations in Italy. — PecUn pleurtmeeUa JjM^l,
Otlrea pUuroneetes L., RuicPH. Amb, Jtariteitk. Tab. 45, figs. A, B, Bladtt.
Malae, PI. 60, fig. 5, Cuv. Jt, Ant., id, UL, MM. PL 75, fig. i. Areiy
numerous genus, of which many extinct species are found in the seoondaiy
fonnations.
iWum Brug. Shell free, inequivalve, elongato-trigonal. Hinge
edentulous. Inferior valve emarginate near the point. Animal
with mantle cirri and ocelliferous tentacles. Foot byssiferous, the
byssus passing through the excised valve.
8p. Pedum tpondyloXdeum Lam., Eneyd. mHh., Vert. PI. 17S, Bladit.
Malae. PI. 61, fig. 6, Cuv. R. Ani., id. iU., MoU. PI. 76, fig. 4, Deshatm
Conekyl. PL 48, figs. 4—6. In the Indian ocean and the South Pacific;
by QuoT and Gaimabd, who first described the animal of this shell, this
species was found yeiy numerous at the Island Vanikoro.
Lima Beug. Shell free, subequivalve, inequilateral, oblique,
frimished with small auriculas, subgaping at the side. Hinge
edentulous. Animal {Olaticua PoLi) with mantle margined with
long annulate cirri and without ocelliferous tentacles. Foot cylin-
drical, rugose, clavate, sulcated at the apex. (Animals swimming
hj contraction of the valves.)
Sp. Lima tquamota, Ottrea Lima L., Poli Te$t. utr, SiciL 11. Tab. 48, Bgt.
91—94, Blainy. Malac. PL 61, fig. 3, &c. The fossil genus Phgioitema
SowsBBT, Lam., contaios different species which belong to Podopti^ or
rather to Spondylut. The remaining, according to the remark of DssHATtfy
ought to be united with the genus Lima, for instance Plagioikma gigos^'
tewn froijA the Lias-formation.
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CONCUIFEIU. 729
Family IV. Malleacea. Mantle open, produced posteriorly.
Foot conical, Bulcate, mostly byssiferous. Ligament internal, often
divided and contained in several little pits. Shell lamellose, pearly
within, often deformed, undulato-plicate.
We transfer also to this group the genus ViUseUa, as was done by
Deshayes, who on the other hand removed Avicula from it, which
genus we with Lamarck leave in this family. The characters of the
group are not yet sufficiently determined, for the animals of these
shells are hitherto only imperfectly or not at all known. Some
genera exist only in the fossil state.
Inoceramus Sowerbt. Shell inequivalve, with points approxi-
mate. Hinge straight, incrassate, furnished with a row of foveolse
for the reception of the divided ligament. (Fossil genus.)
Many foseil species, especially from the Chalk-period, are contained in
thiB genus, whilst none from the tertiary formations are known. Debhatxs,
as also GoLDFUSS and Bronn, now unite the genera CatiUus and Myti-
loldes of Brokoniabt with this. Compare the article Inocerame of Dbs-
HATBS, Diet, univ, cTffist. not. vn. 1846, pp. 53, 54, and Bboitn Ldh,
ffeogn. pp. 689 — 696 (first edition).
Pachymya Sowebby ?
This genus Dsshateb united formerly with Catillui, and now with Ino-'
ceramiu; compare, however, Bbonn loc. cit. pp. 696, 697. There is only
one species known, also from the chalk-formation: Pachym. gig(U Sow.,
Bbonn leth. ffeogn. Tab. 31, figs. 9 a, b.
Pema Brug. (Species of Ostrea L.). Shell flattened, subequi-
valve, at the anterior side gaping, an apertnre emitting the byssus.
Hinge straight, linear, edentulous, with parallel furrows receiving
the divided ligament.
Sp. Pema itogtmum Lam., Ofitrea Uogonum L., Rukfh. Anib. Jtariteiik.
Tab. 47, fig. J, Cuv. Jt, Anim., id. ill., Moll. PI. 81, fig. i; Ind. Ocean;—
Pema femoralis Laic, and P, canina Lam. are, according to Dbbhates,
only varieties of this species. — Pema ephippium L., GufeiN Iconoffr.,
MM. Pi. 36, fig. 3, &c. Some fossil species also of this genus from the
secondary and tertiary formations are known.
OervilUa Defr.
A fossil genuB nearly allied to Pema, of which species from the oolite-
and chalk-period are known.
Grenatula Lam. Shell thin, flattened, lamellose, not emarginate
for the passage of byssus. Hinge edentulous, linear, crenate at the
margin, with semicircular pits receiving the ligament.
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730 CLASS xn.
Oomp. Laxabck Ann, du MtuHm, UL pp. 15 — 31, FL a.
Sp. Orenatvla avicularu Lam. 1. L PL 9, figs, i, a, Cuv. R, Ani., id, UL,
Moll, PI. 81, fig. 9 ; in the seas Bunounding the South American ooafte.
Vulsella Lam. Shell irregular, elongate, thin, stibeqmvalTC,
free. Hinge in both valves famished with a tubercle cut out
with a conical pit for receiving the ligament.
Sp. VuUdla Unguatula Lam., Mya vuUeUa L., RuMPH. Amh. SarHeiti. Tab.
46, Bg, A, Blauty. Malaeol. PL 62, fig. 5, from the Indian ocean, &c
Some spedes are met with amongst corals, others in sponges. One
spedes from the tertiary formation is known ( Vtdtdla deperdUa IiAJC, frtxn
the ealeaiirt grouier), and a few from the chalk-period.
MaUeua Lam. Shell subequivalve, rude, often undulate, de-
formed, elongate, flattened, auriculate, emarginate anteriorly for the
passage of bjssus. Straight, triangular, sinus between the two
valves; points small, divaricate. Hinge edentulous; cardinal
fossula oblongo-conical, receiving the ligament.
Sp. MaUeut vutgarii Lam., Otirea maUau L., RuMPH. Amb. BarUeUk. TA,
47, fig. n.; D'ABOBiryUiLX Coneh^iologie, 1757, 4to, PL 19, Sg. 1,
Blainy. Malae. PL 63, fig. 4; this shell, commonly caUed the PoUtk
Hammer, is found in the East-Indian Ocean. On account of its rarity in
collections, MaUeui albut Lam., Chemnitz Conehyl. Tab. 906, from the
South Sea» may be noticed.
Avtcula Bruq., Deshates. Shell subequivalve or inequivalve,
often squamose externally. Hinge edentulous or with one or two
small teeth. Area oblique, submarginal, sometimes subindistinct
for the internal ligament. Internal surface of valves pearly.
Animal compressed, with mantle open, the margin incrassate,
cirriferous. Foot conical, vermiform, with byssus thick, the fila-
ments sometimes coalesced.
Sub-genus : Avicula Lail Shell oval, oblique, with ears large*
A cardinal tooth in both valves beneath the points.
Sp. Avicula aemimgiUa Lam., MyHUu ffirundo L. in part, Blainv. Maiae.
Pi- <53, fig. S;—Avic. Tarmtina Lam., Poli Teatac, utr. Sk. Tab. 32, fig^
1 7 — 31, &c. Fossil species also from the Muachd-haOe and other fonnationB
are known.
Mdectgrina Lail Shell suborbicular, with ears not produced and
hinge edentulous.
Sp. Avicula margarUifera, MytUua marffarUiferut L., Bumps. Amb, Bari-
teiih Tab. 47, figs. F, a; D'Abqknt. PL oo, fig. a, Blaint. Malac PL
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CONCHIFEBA. 731
35 bis, fig. 7, Cuv. R. AnL, 4d. iU., Moll PL 85, fig. i; the pearl'theU,
The shell is sometimes four-cornered, with round margin, greenish-brown
striped with white spots, nacreous within, yellow or blueish white. This
oonchifer produces the finest pearls, and is found in the Persian gulf and
in the Indian ocean at Ceylon and Borneo, in the straits of Malacca^
ftc. The largest pearls are found in the deepest places, as in the Persian
gulf near the island Kharrak, where they Ue on yery deep banks. The
Persian pearls are bard, and consequently more esteemed than those of
Ceylon, which are often subject to desquamation.
B. Dvmyaria. Two muscular impressions, mostly remote,
sometimes approximate. Ligament of yalyes external in most.
Family V. Mytilacea. Mantle cloven anteriorly. Foot conical
or tongue-shaped, byssiferous. Two muscular impressions in each
valve remote. Anterior adductor muscle often very small.
Pirma L. Shell fragile, thin, often imbricato-squamose, elongate,
triangular, angustate towards the points. Hinge edentulous ; liga-
ment marginal. Animal {Ckimoera PoLi) with foot vermicular,
subulate, transversely rugose, delivering at its base the byssus
of long, dense, silken filaments. Anterior adductor muscle near the
apices, posterior larger, subcentral. A conical contractile appendage
at the posterior part of mantle {prdchea PoLi).
The shells are large, triangular, thin, and rery small at the point ; along
each shell there runs lengthwise a keel-shaped eminence sometimes feeble,
for each is formed of two sur&ces that meet at an obtuse angle. The
animal penetrates the sand with the point of the shell, and fastens itself
above it by the byssus to other objects. The silky byssus-threads of species
from the Mediterranean are used in Calabria and Sicily as guards for
gloves, stockings, purses, fto.
Sp. Pinna nchilU L., Piwna muHcata POLI, Te^tacea vJtr, Sic, Tab. 34, fig.
I, Blaiky. McUae. PI. 64, fig. i ; — Pinna rotundata L. (and P. fqtmmota
Ghbll., Lax.), Listeb Tab. 374, fig. 215, JBneyd, mHh., Ver$, PI. ooo,
fig. 9, Ac. Many fossil species of this genus from the secondary, and some
from the tertiary formations are known ; to the last belongs Pinna margari-
taeea Lax., Ann, d/it Mtu. ix. PI. 1 7, fig, 3, from the ealcaire grottier,
Mytihs L. (in part), Bruq. Shell elongate, not squamose,
rather smooth, equiyalye, with points placed forward, hinge edentu-
lous or with two teeth obsolete. Ligament dorsal, linear, received
in a narrow, elongate, marginal sinus. Two muscular impressions,
the anterior very small. Animal {CaUitnche PoLl) with foot
lingulate, canaliculate, byssiferous, the mantle concrete posteriorly
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732 CLASS XII.
to form a small anal tentaculiferous canal. Labial palps four,
elongate, lanceolate.
MytUvs Lam. Points terminal
Sp. Mytilut edulis L., Babteb Naiuurh, Uitsp. i. 117 — 117, Tab. ZL figi.
9—1 1, Ctjy. JH, Ani,, 4d. ill., MoUusq, PI. 89, tMuad. The shell is elongate
and, after the epidermis has been removed, purplish-blue. This species u
found in the North Sea and the Mediterranean. Mussels grow quickly,
and are already edible in the second year. Sometimes they are injurious,
and in the spring with us they are less pleasant to the taste ; as experience
in Zealand has taught, they are best from July to January. Some con-
stitutions cannot endure mussels, and this has in part contributed to give
these conchifers a bad name, as though they were sometimes poisonous.
The eggs are not taken up by the gills, as in AnodotUa and Uhio, but by
the mantle. Compare for the anatomy of the mussel A. Ds Hside Ana-
tome Mytili. Amstelod. 1683, 8vo, PoLi Teat, utr. Sicil. n. pp. 197 — 107,
Tab. 31, G. R. Tbbvibanub ZatocAr. /. Pkytiol, i. 1824, s. 41 — 47, Tab.
T. figs. 26 — 29. For the other species of this genus, see Lailabck Hid.
not, da Ani. », v. and Dbshates Encyd, nUth., Ven, n. pp. 557 and foIL
Modiola Lam. Points sublateraL
Sp. Modiola tvUpa Laic., Encyd, m&h,, Ver$. PI. 221, fig. x, from the golf
of Mexico, &c.
There are elongated species, that penetrate into stones, of which Cuvieb
forms the sub-genus LUhodomut, Sp. Mytiltu lithophagua L., Mod. litko-
phaga Lam., Blainy. Malac. PL 64, fig. 4.^
Tichogonia RosSM., Dreissena V. Beneden. Shell elongate,
trigonal, inflated posteriorly, flat or subconcave anteriorly. Points
terminal ; internal septum near the points. Mantle anteriorly con-
crete, excised for foot. Anal trachea distinct.
Sp. Tichogonia Chemnitzii Rosshaesslkr, Mylilui polymorphta J(unatilis
Pall., Mytilui Bagenii V. Baer, Deshateb Conchyl. PI. 37, figs. 9— ii>
&c. Pallas first discovered this species in the Wolga ; in Holland it was
first met with in the sea of Haarlem by H. W. Waabdbivbubo, ^0
described it under the name of Mytilua lineatu8 Lax., with which. However,
it does not agree (Ann. Acad. Lugd. Bat. 1826, 1827, CommaU. de Moll
indig.) ; it occurs also round Leyden in the Cingels and elsewhere. YaV
Bbhxdbn has given a further account of this animal m Ann. det Sc nai.,
^ In the columns, which are the remains of the temple of Serapis in the gulf oi
BaisB, an entire segment is seen, which is pejf orated by these animals, about 22 feet
above the surface of the sea; see PoLi Tett, utr. Sic. n. p. 216; recent geology has
borrowed a proof from this of the depression and upheaving of the gfround fitnn which
so many phsnomena receive their explanation. Compare the well-known work of
Ltkll PrincipUi of Geology.
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CONCUIFERA. 733
Bee. S^rie, m. 1835, pp. 193—2x3, PI. vni.; Cantrainx, who pUees this
geDHfl, on account of the m&ntle for the most part united, not in this family,
but in Bystomya, names it Mytilina. Ann, dea Sc. not., sec. S^rie, Yii. pp.
309 — 313. It agrees, however, very closely with MytUui, as well in the
shell as in the animal.
Family VI. Arcacea. Mantle cloven anteriorly, not produced
into a trachea. Foot large, in some serving for creeping, in others
furnished with byssus homy, lamellose, not divided into filaments.
Branchiae composed of loose filaments. Shell regular, with points
remote. Hinge with numerous teeth arranged in a row. Two
muscular impressions remote.
Area L. (in part). Hinge linear, straight. Shell transverse,
inequilateral, with points distant, a large area intervening. Animal
(Daphne PoLi) with foot compressed, cloven longitudinally, mostly
adhering by means of lamellose, homy byssus to marine bodies.
Sp. Area Noes L., Rumph. Amb. Rariteitk, Tab. 44, fig. p, Encycl. mith,,
Ver9. PI. 303, Blainv. MalacoL PI. 65, fig. 2 ; in the Mediterranean and
the Atlantic; the NoaKz Arc; the boat-like form occasioned the generic
name. — Area tortuosa L., Rumph. 1. 1. Tab. 47, fig. k, Encycl. mHh., Vera,
PI- 305^ fig. I, Blainv. Malae. PI. 65 bis, fig. i, from the Indian Ocean,
&c. A very numerous genus, of which also fossil species occur, some in
secondary, but the most in tertiary formations.
CucullcBa Lam. Shell ventricose. Hinge terminated at each end
by ribs exsert, parallel to the margin, vaiious in number.
Sp. Area eucullua GmEL., CucuUcea aurieulifera Lam., Blainv. Malae. PL
65, fig. 4 ; from the Indian Ocean. The fossil species of this small group
are the more numerous, since amongst the animal world now living, only
this single one is known.
PectunculiL8 Lam. (Species oi Area L.). Shell orbicular, thick,
shut. Hinge arcuate, with teeth oblique, the median obsolete.
Animal {Axinoea PoLl) with foot hatchet-shaped, serving for creep-
ing, very large, compressed, not byssiferous ; heart single.
Sp. PectuneuluB piloaus Lam., Area pUoaa L. (and glycimeris in part), Bo-
NANNi Beereat. Mentis H Oculi, Komse, 1684, fig. 80, PoLi Teat. vir. Sie,
Tab. 26, figs. 1, 3, Gu^iN Iconogr., Moll. PI. ^7, fig. 6, Dkshayes Con^
chyl. PI. 34, figs. 13, 24^;— Peat, fflycimeria Lam., Desh. 1. 1. figs. 21, 22,
1 The shells of this species are nsed by the Sicilians, acoording to the account of
FoLi, for making cameos.
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734 CLASS XII.
fto. Many extinet fossil species also of this genus oocor, whilst the newv
tertiBiy formatioBs oontun also the named living species in a fosnl sUk.
See on these fossil species Lamarck Ann. du Mum. ti. pp. 915—117, n.
Fl. 18, figs. 6--9, and Duhates Deter, det Chquittet fo$§. de$ enviftmi de
Paris.
Family VII. Trigoniacea, Mantle cloven anteriorly, trachea
none. Foot not byssiferous, serving for creeping. Shell equivalve,
inequilateral, hinge with several teeth, arranged in two rows
diverging from the point. Points contiguous, or slightly remote.
Two muscular impressions remote ; internal surface of valves often
pearly.
Nucula Lam. Shell transverse, with points contiguous. Hinge
linear, broken, interrupted by a pit for receiving the internal Uga-
ment, with numerous teeth.
Sp. NucvXa fnargaritaeea Lax., Area nueleut L., Blainv. Malae. FL 65,
Bg. s, DssHATKB Coneh^ FL ^ figs. 1 1— 13 ; in the North Sea and Medi-
terranean. This species also ooours fossil. The foot of theae oonchifen
resembles that of Peelunculut, This genus, which has filiform gills, might
be left in the preceding fiunily. Of late years many species haye heoome
known, especially through the English voyager Cvifxuia. They are aU
small oonchifen.
Trtgonia Bruq. Shell trigonal, sometimes suborbicular. Car-
dinal teeth oblong, grooved. Ligament external, marginal. Foot
elongate, geniculate, with the apical portion received in a groove
on the inferior surface of the basal portion.
6p. Trigonia peelinaia Lam., Tri^n. margaritaoeOf Ann. du Mut. IT. pp.
355—357, Fl. 67, ^, I, BLAlNvnJiB Malac. FL 70, fig. i; in the S.
Facifio. This is the only species now living which is known of this genus,
of which numerous fossil spedes occur, partly in the older seoondaiy
formations. Here belongs also the genus Myopkoria BftONV Zetk. geogn>
I. pp. 168 — 174, from the Mmehdrkalk, Especially, however, are the
species of Trtgonia numerous in the Jura- and Chalk-formations. Camp*
also AQA88I2 JEhides eritiquet tw let Mollu$que$, ire livraitan, NeufchaH
1840, 4to.
Family VIII. Naiadea. Mantle mostly cloven, sometimes
concrete posteriorly and produced into two short trachese. Foot
compressed, byssus none. Four oval buccal tentacles. Shell
transverse, inequilateral, pearly within. Two muscular impressions
distinct, remote. Ligament external.
This family contains fresh-water conchifers. LnmiEUB united the
species known to him partly with Myct, paiiily with MytUu$. They
might perhaps be all united in one large genus Unto,
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CONCHIFERA. 735
Phalanx I. Bhdl regular^ freSy with points approannuUe,
Unto Bruo. Animal {lAmnoea PoLi) with mantle cloven, con-
crete posteriorly by a commissure about the anus, surrounding the
anus with a trachea like an aperture. Posterior margin of each
lobe of mantle furnished with cirri or tentacles. Branchiae of each
side coalesced behind the foot. Shell thick, pearly within, with
points most frequently decorticate. Hinge mostly with two teeth
in each valve, the anterior thick, obliquely striated, the posterior
compressed like a lamella, elongate, sometimes obsolete. (Teeth
of left valve bipartite by a groove for receiving the teeth of the op-
posite valve.)
This genus is very numerona in species, especiaUj in North America.
The knowledge of these species is attended with great difficulties, since the
naturalists who have described them have dispersed their descriptions,
partly in Magasdnes, which are entirely unknown in Europe, and at the
same time have paid little regard to the contemporary or previous labours
of one another. We mention here only, as the most accessible, the papers
of Rafikesqus (Ann. des Sc. phynque* de MM. Bobt db St. Vinosnt et
Dbapisz, y. pp. 387 and foil.), J. Lea (Americ, PhUos, TrcmMoct,, new
Seriet, ni. Philadelphia, 18^8, pp. a59— 463; m. Pt. 2, 1830, pp. 403 —
457, IV. Pt. I, pp. 63— i«i), and the memoir of De F^ussao, in great part
bibliographical, in Gu^in's Magtuin de Zoologie, 1835, '^ which the
descriptions of Th. Sat and others are cited.
The form of the numerous shells of this class is very different. For some
of the forms distinct genera have been adopted. Thus Lamarck distin-
guishes the species with a triang^ular shell and transverse striie on the teeth
of the hinge, by the name of Ccutalia (Sp. Cattalia ambigua Lam., BLAiifT.,
Mtdacol, PL 67, fig. 4, from S. America). The species that have an oblique
shell, and at the posterior margin present a large, flat, triangular pro-
longation, he unites under the genus Hyria, (Sp. Unio tyrmalophorua O.
Fabb.\ Hyria avictUarU Lam., Mya aymuOophora Gbonov. Zoophylac,
Tab. xvni. fig. I, Unio corruff(Uu8 Blainv., Malacol. PI. 67, fig. i ; — Jfyria
corrugata Lam., Encycl. m6tk,, Ver$, PI. 247, fig. 2.) The other species
Lamabck leaves together under the genus Unio. But besides Ccutalia and
Hyria great varieties of form are found, of which sub-genera might be
made, as Unio orbictUatus Hildb. or Unio tvbrotundiu Kafin. of a round
form*, Unio arcarformis Lea, with a flat broad surface behind the points,
&c. In European species the shell is usually less different in form, elongate,
forwards round and obtuse, backwards pointed and prolonged.
1 O. Fabbioiub in Kongd. Damke Vidmahab, SeUkahs Naturvidenth. Afkand'
Unger, i. 1814, pp. 55— ^i*
* See, for example, Unio verrueo»a Yalbno., in Humboldt et Bonflakd BecueU
eTObierv. de Zool, d d'Anat. eomp. n. PI. 53, fig. 2.
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736 CLASS XII.
Sp. Unto pidorum Lax., Mya pietorum L., SruKM Fautk. Oerm. yl Hefk t.
Tab. a, Unto rottnUa PriiFPZB Jkttlteke Land- und Wanrrtchu. L Tab.¥.
Eg. 8 ; — Unto wuirgariHfenu Rets., Nilsov, Mya marffaritifera L., Unio
ginuala Lam. (and Unio eUmffota Lam.), Sohbobtes GtMckiekU der PUut-
eonchyl, 1779, 4to, Tab. lY. fig. i, Pfxiffkb L L Tab. v. ^. 11, ftc Tfai*
species, like the Mdeagrinaj produces fine pearls (see above, p. 688). — ^Ai
a very large species from N. America we note: Vnio undulaHu Sat,
Unio Peruviana Lam., Bneyd. mHk., Van. PL 148, ^. 7, VAUorcisssia
in Humboldt et Bonp. Obg. de Zool. H iTAnat. comp, 11. p. 239, PL 54,
fig. 3.
Anodonta Brug. The inhabitant of the shell {LimruBa Pou)
of the same form as the animal of the preceding genus. Shell
mostly thin, transverse, elongate posteriorly, rotmidato-troncate
anteriorly. Cardinal margin edentulous, linear.
Biver-muud. This genus differs from the preceding in the greater thm-
ness of the shell, and especially by the absence of hinge, but the animal is
entirely the same as in the genus Unio, with which too Dkshatss unites it
In the shell also Unio dtpretta V. Mukhlfibld (Pfkiffsb L L n. s. 32,
33, Tab. Tin. figs. 3, 4) forms the transition to Anodomta. Hie spedes of
Anodonta are much less numerous, and exhibit fewer varieties in the form
of the shell, than those of Uni4i,
Sp. Anodonta cygnea Lam., MytUus cygneut L., Blainv. Malae, PL 66, fig.
I, Pfeiffbb 1. L I. Tab. yl ^, 4, (on account of the coalesoenoe or
growing together of the shells on the dorsal side above the ligament, Lka
forms of this species and some others, the genus SympKynota; such a ooo-
nexion also occurs in certain species of Unio); Anod, anatina Lam., Myfi-
Itu anatinui L., Pfkiffsb Tab. vi. fig. a, &c.
Fossil species both of Unio and Anodonta occur in the coal-formation.
In the Oolite and Lias shells are found which were formeriy referred to
Unio, but now constitute the genus Cardinia of Aoassiz, Padkyodon
Stutchburt {Unio hybrida, Unio concinna Sowkbbt, &c.). See Stdtch*
BUBY Annals and Magaz. of Nat. Bitt. vin. 1841, pp. 481 — 485.
Iridtna Lam. Animal with lobes of mantle concrete poste-
riorly and furnished with two short tracheae. Shell transverse,
elongate, with hinge edentulous, linear, sometimes crenate.
Sp. Jridina exotica Lam., Le mutel Adanson, Hist. not. du SSnSgal, Co-
quillage», PL 17, fig. 21, Blainv. Malae. PL 66, fig. 3, in West Africa and
in the Nile ; the animal is described and figured by Debhates, MHh. de to
8oe. d'Hitt. not. de Paris, in. 18^7, pp. i— 16, PI. i;—Irid. mftmi DisH.,
, Anatina rubens Lam., Sncycl. mith.. Vers. PL aoi, fig. i.
Phalanx II. Shell affixed^ irregular.
Etheria Lam. (JEtherta). Mantle cloven, with margin cirri-
ferous. Four tentacles round the mouth oblong, rotundate. Liga-
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CONCHIFERA. 737
ment external, penetrating partly internally into a groove. Hinge
callous, edentulous,
Lamabok, who first made this genus known, thought that it was found
in the sea. It was placed in the neighbourhood of the Oysters, from which,
however, the shell differs by the two muscular impressions. Sinoe that
time these oonchifers were found in the Nile by the traveller Caillaud.
The animal was afterwards described and figured by Rang and QuoT. It
agrees, as Rako remarks, with that of Unio and Anodonta, so that in fiMst
the EthericB might be shortly characterised as irregular AnodorUa with one
of the two shells attached.
Comp. Lamabok Ann, dv Miisium, x. pp. 398 — 408, PL ag — 32 ; F^
BUSSAO Notice t, I. EthMt$ trouvies dana le Nil, Mim, de la Soc, d*ffitt.
not, de Paris, i. 1813, pp. 353 — 372 ,* Rang et Caillaud Mhn. wwrle genre
£thSrie et descr. de wn Animal, Nouv, Ann, du MtuSum, iih 1834, pp. 138
— 144, Qdot in Lesson lUtutr. de Zoologie, PL 58, 59 (1835).
Sp. Etheria Lamoflrckii F^usa., Eth, elUptica Lax. (and JSth, trigoniula
ejusd.). Arm, du Mus, x. PL 29, 30, fig. i, BLAimr. Malae. PL 70 bis, fig.
4, &c., Gu^BiH leonogr,, MoUuaq, PL a 6^ fig. 8.
All the known species live in fresh water in Africa, and would seem,
with Iridina, to represent the Uniones so numerous in America^ which
are not abundant in Africa.
Family IX. Carditacea. Mantle cloven. Foot compressed,
sulcate, not tyssiferous. Tentacles around the mouth divided into
several lobes. Shell regular, free, inequilateral, equivalve. Two
muscular impressions remote. Teeth of hinge mostly two, in some
a single tooth, the anterior being obsolete.
Cardita Brug. (exclusive of species), Deshates [Cardita and
Venericardta Lam.). Shell suborbicular, transverse or oblique,
often costate longitudinally.
An historical reyiew of the genus CardiUi, and of the modifications it
has undergone, was given by Dbshayeb Encyd, mith,, Vers. n. 1830, pp.
194 — 196. All the species now living are marine, mostly small or only of
moderate axe. The fossil species are numerous, especially in the calcaire
gromer, as Cardita imbrtcata, Venericardia imbricata Lam., Libtbb Biet.
Conchylior, Tab. 497, BLAnrv. Malacol. PL 68, fig. 3, Dbbhatbb Conchy
liol. PL 3r, fig. 6, &c.
Family X. Chamacea. Mantle excised with a middle aperture
for the foot, and furnished with two distinct foramina in the tentacu-
late margin (trachea of mantle and trachea of gills), elsewhere
VOL. I. 47
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788 CLASS XII.
closed. Foot round, geniculate, not byssiferons. Shell insular,
inequiyalTe, adhering. Two muscular impressions remote.
In all the preceding fiunilies of the Dvmya/ria the mantle is
entirely open at the anterior part. In the foUowiiig £eanilie8 the
marginR of the mantle are united anteriorly. Amongst these the
IH^Bent fiunily is distinguiahed by its irregnlar shell, just as the
^theriaoea are amongst the preceding &milies.
Chama L. (in part), Bruo. Shell irregular, mostly imbricate
or lamellose, with points incurved. Hinge with single oblique
tooth in each valve, received in a little pit of the other valve.
(Animal, Rilopua POLI, comp. characters of fBuooily.)
Sp. Chama Lazamu li., Ckama damcBcomu Lam., Bumph. Amb. Mariieitk.
Tab. 44, fig. 3, D'Akosnvills OonehyL PL to, fig. r, Chemst. Otmehr
lienhab. Tab. 51, figs. 507, 508 ; Eut Indiet ;— Chama maoeropkffUa GHun.
1. L Tab. 52, figs. 514, 515, Cuv. R Ani,, id, iU,, MoU, PL 95, fig. «, &c
All the species of this fiamily live in the sea.
Cleidothenu Sowerby.
Sp. Chama aCbida Lav., CUidoikerus chamoldes Sow., Dbshatks CcmckyUol,
PL 99, figs. 1—5 ; fi^>m the Soath Sea at New Holland.
Dieeraa Lak. Points large, divaricate, contorted into irregnlar
spires. Cardinal tooth very large, thick, incmrved.
A fossil genus, of whioh two or three species are known from the upper-
most oolite and chalk fonnations.
Sp. IHceras arietina Lam., Dksh. Conchf^icl. PL 28, figs. 4 — 6, Cur. R-
AfU,, id. m., MoU, PL 95, fig. I ; this species was first disooTored by
DSLUO in the chalk strata of Mont Sal^ Saussubb Voyaga dam laAlpa,
I. PL n. figs. 1—4 (M. Svo. Neuchatel, 1803, pp. 277—180). Comp. on
Dieerat Lamabok Ann, du Mua, vi. pp. 298 — 301, and Deshatss Did,
cUm, tTEitt. not, v. 1824, pp. 465—467.
Appendix to Gkamacea.
Family of uncertain position, Sudista or RudistcB, Fossils in
cretaceous strata*
Superior valve operculiform, smalL Inferior aflfixed, elongate,
often divided internally by septa.
The fossil conchifers designated by Lamabck under the unintel-
ligible name Rudistes, with which, however, as Deshates remarks^
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CONCHIFERA. 739
the genua ffippuaHies (placed by Laxabck amongst the Cephalopoda)
should be united {Ann, dea Sc not y. 1825, p. 205, xv. pp. 258—
266) seem, when the genera Crania and Calceola, belonging to the
Brachiopoda have been separated £rom them, to be allied to the
Chamctcea; they had probably, however, an entirely open mantle,
like the jEtheriacea, Oai/reacea &a (Deshates Encyd, meth,. Vers.
ni. p. 918). The Hippurttea, large, elongate, conical shells, resem-
bling in some degree, as to external form, the horns of mammals,
were first discovered at the foot of the Pyrenees by Picot La
Petbouse. Goldfuss, however, and D^Obbigkt also, is of opinion
that these EudisUe belong to the Brachicpoda and by no means
to the Lamellibranchiata {Ann. des Sc. mU., sea S6iie, xvui. 1842,
pp. 173 — 192), in which they now find many supportera
Genera: HippurUes Lail, Caprina D'Obbignt, Sphcerulites
Debmoul. (and Radiolites Lah.) Desh. {BirostrUes Lam. a fictitious
genus £rom the internal mould of SphcenUUes.)
Comp. Deshatxs in Lamabck Bitt. not, dea ArU. s. vert., ie ^ vn.
1836, pp. 178—397, PIOOT Db La Pbtbousb JM novU qiUbuad. Orihoeeror
Htum et OetraeUum apecieb. Korimb. 1781, foL; Dbshatbs Saaai eur lea
SphSrtUitta (BvUH. delaSoe. cTSiat. not. deBordeaux, 1. 1817); O. Bollakd
Du BoQUAV, J)«aeripHon dea eoquiUeafaaaiUa de lafamiiUe dea Budiseea, qui
ae irouvent d^na le terrain critaei da CorbUrea. Avec planches. Carcassone,
1841, 4to.
Family XI. Tridctcnocea* Mantle with lobes for the greatest
part concrete in the margin, furnished with a superior aperture for
the passage of foot, and two small inferior, the anterior anal, the
posterior respiratoiy. Four tentacles round the mouth elongate,
acuminate. Foot thick. Ligament external, narrow. Two adduc-
tor muscles approximate or confluent ; muscular impression single,
little distinct. Shell regular, equivalve, free.
CuviEB {Eigne Anim. IL 1817, p. 475) remarked that in the
conchifers of this &mily there is only a single adductor muscle.
Consequently Lamabck placed the genus Tridacna subsequently in
the division of the Monomya/ri4i, with which, however, it does not
agree. From the figures of the animal of Tridacna mtUica given
by QuOT and Gaimard in the narrative of the voyage of the Astro-
labe (copied in Guv. R Ani, ^cL iU., Moll. PL 96, fig. 3) it seems
that two muscles are present however, which lie close together, and,
although in other species they are confluent and form a single
muscle, this is no reason for separating Tridacna from those fiunilics
with which in other respects this genus has most agreement.
47-2
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740 CLASS xn.
Tridacna ' BnVQ. (Species of Chama L.). -Shell traMVcree,
inequilateral. Points approximate. Hinge with two teeth, placed
behind the points under the ligament ; the posterior in the right
valve bipartite, a groove receiving the posterior tooth of the left
valve.
Hippopus IiAiL, lunula closed Foot not bjasifeTOiis.
Sp. Tridaena Kippopu$ BLABrr., Chama kippoput L., ff^ppaput maevldM
Lam., Rumfh. AnJb, RarUekk. Tab. 43, fig. €, K.AIW. Malaatl. PL 68,
fig. a, GuiBiH leonogr., MoOutq. PL 19^ fig. 7; Indi&n Ooean.
Tridaena Lail Shell gaping at the lunukt, with aperture orate.
Foot byssiferoua
8p. Tridaena tquamoia Lam., Bttxph. Afiib. Raritettk. Tub. 43, fig. a, Auryr/.
mith., Ven. PL 136, fig. i ; Induuii Qoean 'y^Tridaena gigat, Cktma gigat L.
(in part), KUMPH. L L fig. B, Eme^. mie^., Ven, PL 335, % i. Thisspedes
alflo oooon in tbe Indian Ocean; it is Uie lar]^«8t copchifer known. Foi^
meriy it waa the custom to ornament gardens with them as basins of foan-
tains ; in ohnrehes these sheDs sometimes senre for holding the holy-water,
as two rery laige ones may be seen in St SiUpice at Paris; henoe the
derivation of the French name hiniUer for this species of shell. Lnm .sus
records a shell of this kind in the MMtmm Ludoviooe Ulrica, that weighs
53a pounds. — In Tridaena the opening of the Umvla in older mdividuak is
■mailer, so that the diffareikoe between Ilippopui and Tridaena is not very
Family XII. Cardiacea. Mantle with lobes concrete posteriorly,
open anteriorly for the passage of foot ; two very short trachefls dis-
tinct, foraminiform, with margin girdled by cirri. Foot large, fisdci-
form, not byssiferoos. Ligament external. Shell regular, equivalve,
free. Hinge with two teeth nnder the point, receiving each
other by mutual insertion, and one or two lateral. Two mnscular
impressions remote.
Cardium L. Shell subequilateral, shorter posteriorly, more
^bbous anteriorly, mostly furnished with longitudinal ribs. Four
teeth in each valve ; with two oblique under the points, receiving
each other crosswise by mutual insertion, and two lateral remote,
compressed. Animal {Cerastes PoLi) with foot falciform, very long,
serving for jumping.
Sp. Ckurdium eduU L., BiiAiiry. Malaeol. PI. 70 bis, ^. 3, Bastxb N^i^i^rL
Uiiap. n, bL 76—78, Tah. vin. figs. 1—4, Poli Teat, utr. Sic. i. Tab. xm,
figs. 13—15 ; on the shore in the Korth and Mediter. Seas; this biyalTe»
like some other species of this genus, is used for food; the common name
in Zealand is kokkaan.
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CQNCHIFERA. 741
Cardium i9ocardia L., Rumph. Arnib, Rmteitk, Tab. 48, fig. g, CHXicir.
Conchyl. Tab. 17, figs. 174 — 176, &c
Henhtcc^rdittm Klein^ Spenoler, Cut. Valves oompreased, a keel
dividing the anterior from the posterior plana
Sp. CarcUum henUe<urdium L., RuiCPH. Amib. RarUdtk, Tub. 44, fig. H,
Blautv. Malac, PL 70 bis, fig. 4 ; — Cardkim carditta L., Lax., Bumpb.
1. L Tub. 41, fig. E, fto.
Of the genua Cardiitm Spbnolsb has given a monogniph, J^kririer <tf
naiurhitt. Sdekahet, V. T. 1799, pp. i — 60, Tab. I.
The fossil species of Cardium are uncommonly namerous, (hey oocur
especiaUy in tertiary formations.
Isocardta Lam. Shell cordate, with points involnte spirally.
Two cardinal teeth under the points, compressed, entering; a single
lateral tooth posterior, elongate. Animal [Oloasua PoLi) with foot
compressed and tentacles of mouth elongate, acuminate.
Sp. Isocardia cor, Chama Cor L., Blainv. Malae. PL 69, ^. 9, Pou
Testae, utr. Sic. 11. Tab. xxni. figs, i, 9 (i. Tab. xv. figs. 34—36 animal) ;
Mediterranean, ftc. ; also fossiL Much more numerous are the extinct
species of this form, which fbr the most part occur in secondary fonnations.
Opts Defrance. Fossil genus. Is this its place ?
Comp. Dbshatss in Laicabok ffist, n. d. Ant, $, v. vi. pp. 519, 500.
Cyprtcoflrdia Lam. Shell inequilateral, elongate posteriorly,
oblique or transverse. Three teeth under the- points ; lateral tooth
single, posterior, compressed, acute.
Sp. Cypricardia obUmga Desh., Chama chlonga L., Cypric. gumaica Lax.,
Blainv. Malac. PL 65 bis, ^, 6, Dbbhatbb ConckyUol. PL 94, figs. 8, 9,
&c.; — ^Here belong also petrified species from the oolite and chalk
fonnations.
Family XIII. Vmeraxiea. Mantle with lobes fimbriated at
the margin, concrete posteriorly, open anteriorly for the passage of
large, compressed, acute foot ; two tracheae moderate, often coalesced
at the base. Ligament external, short. Shell equivalve, free,
regular, shut. Mostly three cardinal teeth, diverging obliquely
from the point; lateral tooth in some single, posterior, in most
none.
Aatarte SOWERBY, Crasshiu Lam. Cardinal teeth two, diver-
gent. Impression of mantle simple. Shell transversely rugose or
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742 CLASS XII.
striated, inequilateral, anteriorly truncato-rotundate, orbiculari-
trigonal or oblique, incrassated towards the points.
AtloTte danmonienais Sow., CroMtina dtmmotUetma Lam., Venui cnun-
teUa Blaikv. Malac. PL 75, fig. 7. The fossil spedes are numeroTis in
the later seoondaiy, and in the tertiary fonnations. C!omp. Ds La Joh-
KAIBB NoU nir le genre Aetarte, Mimoiru de la Soe. (THisL noL <U PtaVf
I. i8«3, pp. 127—131, PI. VI.
Venu8 L. (exclusive of species). Cardinal teeth in some three
in each valve, in others four in left valve. Lateral tooth none.
Impression of mantle below the posterior muscular impression
deeply sinuate. (Animal Callista Poll)
a) Cardinal teeth four in Ufi vahe, wih fourth eionffoie, eituated under
luntda, compreeeed, received in afaveola 0/ right valve,
Cytherea Lam.
Venus lutoria, Oyth. lutoria Lax., Venue chione L. (in part), Bukfh.
Amh. RarOeUh, Tab. xui. fig. o, Eneyel, mSth., Vere. PL 270, fig. i. The
Chinese and Japanese ooyer the inside of these shells with gold aodcolonr^
and use them in a certain game. — Venue Diane L., Ktjmph. Amb. BarileUk.
Tab. XLvra. fig. 4, J^icycL mith,, Vere. PI. 275, fig. i, Houttuyh Nat.
Hiet, I. i5e Stuk, PI. 1 1 7, fig. 6 ; East Ind. Seas. This species is described
by LiNNJEUB in detail and metaphorically, Syet. not., ed. 12, i. p. 11 99, &c.
b) Cardinal teeth three in each valve, approximate.
Vemis Lam.
Sp. Venue plicata Gm., Lam. ( Venue Dyeera L. in part), D'Abobsv. Con-
chyl. Tab. 11, fig. K, Encyd, mith., Vere, PI. 275, fig. 3; VenutpapiUo-
nacea Lam. (Venue rotundata L.), CuY. JL Ani., id. HI,, MoU, PL 104,
fig. 4, &C.1
Cyprina Lam. Shell obliquely cordate, thick. Hinge with
three unequal teeth, and lateral tooth remote, posterior, situated
under the fissure. Impression of mantle rotundate, entire. (Ani-
mal with trachess short, scarcely porrect beyond the margin of
shell.)
Sp. Cyprina ielandiea Lam., Venue idandiea L., Muell. ZooL. Danie. Tab.
28, Blainv. Malac, PI. 70 bis, fig. 5, Cuv. jB. Ani., id. iU,, MdH VI loi,
* The species belonging to the genus Venue (Venue and Cytherea Lam.) aw^
numerous, that they make up nearly ^ of all the known lamdUbramfhiata, so that this
genus much surpasses the genera Tellina, Cardium, Area, Oetrea eixid Peden w ndi
m species.
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CONCHIFEBA. 743
fig. 3. The Vmus isUmdiea of Bbooohi (and Ojfprina gigoi Lax.) a fossil
species irom the calcaire gromer and other tertiary strata, does not belong
to this genus, but to the preceding, Fimw Broechii Dsbh., Bbokh Ldh,
Tab. 38, fig. I.
Family XIV. Cychdea. Mantle concrete posteriorly, fur-
nished witt two tracheae. Ligament internal. Foot large, not
byssiferous. Shell equivalve, free, gibbouator ventricose. Hinge
with teeth under the point Yarious, sometimes obsolete, and
with lateral teeth gone. Two muscular impressions remote.
This small fieonily contains conchifers that live in rivers and
in fresh-water lakes, like the J^aidea, frt>m which it sufficiently
differs, although Iridina (see above, p. 736), on account of the
imited mantle and the two trachesa might be brought to it. The
present fiunily is nearly allied to the preceding, whose place they
represent in fresh water.
Olauconorm Gray. Cardinal teeth three, with middle one larger,
oflen bifid. Shell elongate, transverse, with points eroded.
Fresh-water bivalves, which in external appearance almost resemble
Anodonta or Unio, They are all exotic. It seems that some fossil fresh-
water shells, which preyiously had been referred to the genus Venw, find
their true place here.
Oalathea Brug., Lam., Potamophila Sow. Shell trigonal, sub-
equilateral. Teeth under the point two in right valve, with a
small triangular median pit receiving the third intermediate tooth
of left valve. Ligament prominent, turgid. Animal with two
trachesB, not concrete, and foot oblong, compressed.
Sp. Galaihea radiata Lau. ( Venus aubviridis and Venus hermaphrodiia Ghel.),
LiSTBB Canchyl Tab. 158, fig. 13, Encycl. nUlh., Vets, PI. ^50, fig. i,
Lahabck Ann, du Mua. v. p. 430, PL 28, Sakdsb Rang Notice swr la
CkUcUhSe, Ann, des Sc, ncA, Tom. xxv. 1832, pp. 152—164, PI. v. In the
rivers of W. Africa; the description of Bang first made the animal known,
the shell alone of this species having been previously known, which some
writers stated to be from Ceylon.
Cyrma Lam. Shell thick, ventricose, corticate, with points
eroded or decorticate. Hinge with three teeth in each valve.
(Animal with two tracheae separate from the base throughout.)
Sp. Cyrena Zeylimka Lam., Blainv. Malaccl, PI. 73, fig. 2 ; a larger variety
of this species occurs at Java, see A. MoussoN, JHeLand- wnd SOsstoasser'
MoUusken von Java, Zurich, 1849, ^^o* Taf. xm. All the species of this
genus are exotic.
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744 CLASS XII.
Gyclas Brug. (in part), Lam. Shell ovato-globose, thin, timiid
at the points. Hinge with teeth veiy small, sometimes sabnull.
Sp. Cycloi cornea Lam., TeUima cornea L., Blaiht. MaXaceH. PL 73, fig. if
Pfeiffeb Sjfilemat. Anordn. «. £eachr. deuUcker Land- vnd WattendM,
Oassel, 182 1, 4to, Tab. v. figs, i, a, ahell brown-yellowy tnospwent^ blue-
wbite within^ 4 or 5"' large; not rare in the canalB of Holland, l^e
species of this genns, which is mach allied to the preceding, are especially
found in Earope,\ut some occur also in other parts of the worid. — ^Pfeitfo
further distinguishes the genuis Pindium, by the abeenoe of long fraeiA^
which in the animal of these species are short and grown together. Sp.
CycUu obUqua Lax., OycU fontinalit Lax., C^. cUuoalis Lam.
Family XV. Lucinacea. Mantle open anteriorly, concrete
posteriorly, and furnished with branchial apertnre and single anal
trachea, or produced into two tracheas connate firom the base
throughout. Foot cylindrical, elongate, often vermiform. Liga-
ment external or subinternaJ concealed between the exsert margins
of fissure. Two muscular impressions remote, large, with anterior
elongate. Shell free ; hinge variable, in some edentulous, in others
with two teeth under the point, and one or two lateral teeth.
Cyrenotda Joannis, Cyrenella Deshayes. Shell ventricose,
corticate. Hinge with two small teeth divergent in right ralFC,
and one tooth bifid in left valve; lateral teeth none. Animal
with two trachesB concrete and four labial palps.
Sp. CyrenoUla Dupontia Joakkib, Gutianr Magasin de Zool, 1835, ^0^*^-
Pl. 64. Oomp. Deshatbs, ibid. No. 70 ; Senegal. This genus counts only
a few spedes, all exotic, from fresh water. According to psaHATES Cyrt-
ndla is a fresh-water Lucina, according to Da Joannis, a C^prena without
lateral teeth at the hinge of the shell. At all erents, this genus is inter-
mediate between this and the preceding Ssunily.
Lucina Brug. Shell orbiculate, transversely striated, subequi-
lateral, with points short, acuminate, incurved forwards. Hinge vari-
able. Animal {Loripes PoLi) with foot very long, vermiform, respira-
tory trachea foraminiform, anal conical, retractile by invagination.
Sp. Lucina lactea Lam., Tellina lactea L., Encycl. mSth., Ven. PI ^S6, fig-
I, PoLi Tulac, utr, Sio. I. Tab. XV. figs. 26 — 79; hab. Mediterranesn ;—
Lucina pentylvanica Lax., Venus pentylvanica L., D'Abqenv. Ccm^}/^
Tab. «i, fig. N, Encyd, nUth., Vers. PL 284, fig. i ; hab. in the ocean »t the
coast of K. Amer. ; Lucina divaricata Lax., Teilina divariaUa h., ^^^
1. 1. fig. 25, Encycl MSth., Vers. PL 385, fig. 4, Dkshatss Oonch^L PI- ^^'
figs. 9 — 1 1 ; hab. in Mediterranean, &c.
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CONCHIFERA. 745
Ungutma Daudin, Lah.
This genus is much allied to Lucina, with which D18HATK8 unites it.
Ungulina ohlonga Lau., Blainy. Mcdae, PI. 73, fig. 6 ; from the coast of
Senegal Since this animal penetrates stones and corals, the external form
of the shell is subject to much yariety. See on the animal, of which pre-
viously the shell alone was known, Duyxbkot Ann. de» Sc. not. sec. S6ne,
Tom. 18, pp. no— ia2, PI. v. B. Gomp. Deshates ibid. Tom. 19, pp.
5- II.
Corbis Cuv. Shell transverse, gibbous. Hinge with two teeth
under the point, and two lateral.
Sp. Corbis fimbriata Lam., VefMU fimbriata L., D'Aboeny. Conchyl. Tab.
21, fig. G, Encyd, mith., Vers, PL a86, fig. 3, Blainv. Malac, PL 71, fig.
4 ; Indian Sea. This shell is finely striated longitudinally between grooves
running parallel to the margin. Hence the French name la TricoUe or la
Oorbeille, of which last the generic name Corlns is a translation. There are
fossil species of this genus which are as finely or even more finely treliced.
Family XVI. Saxkavina (or Lithophagd). Lobes of mantle
as to the greater part concrete, with apertm-e narrow, anterior, emit-
ting the small foot. Tracheae two, coalesced at the base, tentacu-
late at the opening. Shell transverse, inequilateral, posteriorly
elongate, gaping. Ligament external. Two muscular impressions
remote. Impression of mantle deeply sinuate posteriorly. (Animals
living in stones, hence the form of shell often irregular.)
This small family is connected with the genus Veiiut on the one hand,
and with Oatlrochama on the other ; perhaps the genus Saacicava might be
placed in the last family. They are marine conchifers like all the following
families.
Petrtcola Lam. (and Venerupis ejusd.) Hinge with two teeth
in each valve, sometimes with three. Branchiae unequal, external
smaller.
Venerupis Lam. Tracheie separate at the apex alone. Hinge
with three teeth in one valve, sometimes in botL
Sp. Venerupis irus LaH., Donax Trus L., Encycl. nUth., Vers, PL a6a, fig.
4, PoLi Test, utr. Sic, 11. Tab. 19, figs. 25, 16 \ hab. Mediterranean ; —
Vener, perforans Lam., Venus perforans MoKtaou. (Phtlifpi has given a
figure of the Mollusc, see Wieomann Archivf. Natwrgesch, 1845, ^&^* "^^l-
figs. 15—18.)
Petricola Lam. Tracheae coalesced at the base, free at the
terminal part for nearly half their length. Hinge with two teeth
in each valve, or with only a single tooth in one of the valves.
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746 CLA8B XII.
Sp. Pehrieola roeedlaria Lam., Dbbh. Cfomek^iioL FL 19, %. 7, te. (ate
ftnimal of Petricola lee Pmuppi L L p. 188, Tab. TH. ligL 11— 14)-
Foflflil fpedes from the tertiary fonnatioiis aie known.
Saxtcava Fleuriau de Bellevue, Lam. {HicOella Dacik,
Lam., By89omya Cuv.). Shell with tooth obsolete, taberculifoim,
or edentulous. Foot very large, often bjssiferons.
Sp. Saxkava phdadU Lam., MglUut pkoiadu L., Mukllxk Zooi. Daaic
Tab. 87, figs. 1—3 (infer.), Bladit. MaUuxl. PL 80, fi^. 5; Saadeam
aretiea Philifpi, Solen minuhu L., Lam., J/jyo aretiea L., SutieBa mrttka
Lam., Ene^. vUtk., Ven. PL 934, f^. 4, Blaiht. Malae. PL 80, fig. 6
(named Momboldu ruffoi,), Cur. 12. AnL, MoU. PL no, filg. i, ftc.
Family XVII. Telltnacea. Mantle with lobes concrete poste-
riorly, margin tentaculiferons. Tracheae two, separate, oft^i very
long. Foot triangular, compressed. Shell transverse, equivalve,
inequilateral, with posterior part often shorter. Ligament external.
Two muscular impressions remote. Impression of mantle sinuate
posteriorly.
That margin of the shell which is ordinarily the longest, is here often the
shortest, the hindmost, namely, or that where the ligament is situated. It
was an error, therefore, in Lamabok to suppose that in JMmax the liga*
ment is displaced, and lies on the Umtdaj it lies, as in all bivalye sheUs,
behind the point.
Donax L. Shell transverse, posteriorly obtuse, flat. Cardi-
nal teeth two either in both valves or in left, a single tooth bipar-
tite in right ; lateral tooth single or two lateral teeth, snbremote,
sometimes obsolete. Ligament short. Animal {PeroncBa Poli)
with tracheae long, distinct, unequal, the inferior longer ; cirri at the
orifice of anal (saj)erior) trachea simple, at the orifice of branchial
trachea ramoso-digitate. Foot large, conspicuous.
Sp. DoTiax scortum L., D'Argxkvillb Conchyl. PL 9t, fig. l, Blaiht.
Afalac. PL 71, fig. i, Dsbhatbs ConchyL PL 14, figs, n— 15 {Donax
pttbescais L. Yariety); hab. Indian Ocean; — Donax truneulut L., Pou
Testae, ulriusq. Sicil, II. Tab. 19, figs. la— 30, Blainv. Malae, PL 71,
fig. ), &c.
(Cap$a Lam. Lateral teeth none.) Sp. Donax branlitntU Bladtv.,
Cap9a bratHieniii Lam., Bneyd. wUth., Ver$, PL «6i, fig. 10, Blaiitv.
Afalac. PL 71, fig. 3.
Psiunmobia Lam. (and Sanguinolaria ejusd.). Shell transverse,
Bubequilateral, subhiant at both ends. Hinge with two teeth,
more rarely with single tooth in each valve, lateral teeth none.
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^
CONCHIFERA, 747
Animal (Peroncea PoLi) with both tracheas extremely long, sub-
equal, and with margin of mantle as though serrate with acuminate,
conical cirri.
Capsa Bbug., Desh. (not Lail) Shell oonvex. Hinge narrow,
with two unequal teeth in each valve. Ligament thick, elongate.
Lunula indistinct.
Sp. Psammobia ruffosa Sowkbbt, Rixvb, SanguinotariarugoBaliAiK,, Venvi
d^/loraia L., Ghbmnitz Conekyl, Tab. 9, figs. 79 — 83, Deshatbb Oonehyl.
PI. 13, figs. I — 3 ; this species of sheU is met with in the Atliuitic, the Bed
Sea and the Indian Ocean, and is amongst the most generally dispersed
species. The animal is still unknown.
Sanguinolwria Lam., Desh. Shell flat Hinge with two unequal
teeth. Ligament elongate, flat.
Sp. Psammchia tangvinoUnia nob., SoUn aanffuinolentUB 6m., SangtUnolarta
rosea Lah., Ghshn. Conekyl. Tab. 7, fig. 56.
Paammoibia Lah., Desh. Shell flat Hinge with two teeth
small, diverging or with single tooth. Ligament elongate, gibbous,
protuberant
Sp. Pwmmdbia veapertina Lam., SoUn v$aperUnu» 6m., Blaiky. Malacol.
PI. 77, fig. 4 ; — Pscmmobia incamata Desh. {Tdlina ineamala L., Fauna
9ttee.)f Ptammob. feroenns Lam. ftc. These two species occur also fossil in
tertiary fonnations.
Tellina L. (exclusive of some species). Shell thin, mostly
transverse, inequilateral, anteriorly shorter, rotundate, posteriorly
plicate, with fold running obliquely from the margin towards the
points. Hinge narrow, with two teeth or single tooth in each
valve; in most two lateral teeth remote. Animal {Peroncea PoLi)
with foot large, lanceolate and margin of mantle cirrate with long
unequal trachese, hiding deep under the sand.
The species of this genus are numerous, especially in other parts of the
world. In most the shell is long and flat, as Tdlina radiata L., D'Aboef-
VILLE Conekyl. PL 22, fig. A, Encycl. mith., Vera. PL 289, fig. a, Blainv.
Malacol. PL 71^ fig. 4; sometimes ending behind with a thin elongated
part, as in Tellina ro$traia L., Tellina Spengleri Gm., Lam. Encyd.,
Vers. PL 287, fig. 5. — In others the shell is more round, as TeUina lingua
fdia L., Rumph. L L fig. o, Gu^n Iconogr., Moll, PL 31, fig. i. In the
North Sea is found Tellina fahula Gmel., Lam., Gbonoy. Zoopkylae. Tab.
18, fig. 9, TeUina tclidula Soland., Lam. &o.
Comp. on this genus Spenqler Skrivter of noiurk. Seltlabeif IT. 2.
Kjobenhavn, 1798, pp. 67 — lai^ Tab. 12.
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748 CLASS xir.
Tellinidea Laic. Posterior margin of shell without irregular
fold
Sp. TdUna Timoreruit Blainy., TeUinide$ Timariennt Lam., Bulehy. Maia-
col, PL 73, fig. 1, (DsBHATBS has rejected this sub-genus, caUing to mind
that some TdUntB by slow degrees conduct to this form.
Fragilia Desu. Shell subtrigonal, gibbous, anteriorly rotnn-
date, posteriorly longer. Hinge with two teeth diverging in each
valve.
Sp. TdlinafragUU !»,, Pdrieola oekroUuea Lam., Poli Tat, tttr. Sie. i. T^b.
XT. figs, aa, 34, Dbbh. Conchyliol. PI. 12, figs. 13 — 35. Hjbb. in Medi-
terranean.
Family XVIII. Maetracea. Mantle open anteriorly, concrete
posteriorly, produced into two tracheae, in some separate, in others
coalesced. Foot compressed, triangular. Shell equivalve, free.
Ligament internal, received in a pit ; in some an external ligament
as well as an internal. Two muscular impressions remote. Im-
pression of mantle marked posteriorly with a sinus, in some broad,
in others deep, narrower,
Amphidesma Lam. (in part), Recluz, Desh. Shell rotmidate,
suborbiculate, thin. Hinge with two teeth in each valve. Liga-
ment double; external short; internal in a pit of hinge oblique,
deep, narrow.
Sp. Amphideama variegata Lam., Encyd. mith.^ Ver$, PI. 391, fig. 3 ; Indian
Oceikn ]—Amph. $olida Gbat, Desh. ConchylioL PL 11, figs. 10 — 12,
South Sea, at the coast of Peru.
Syndosmya Recluz, Desh. Shell transverse, oval, thin, with
anterior side shorter. Hinge with two teeth in right valve, with
intermediate foveola receiving the single tooth of left valve. Two
lateral teeth. Ligament double ; external small, internal received
in a spoon-shaped process. Animal with two tracheae long,
separate.
Sp. Syndosmya dtba, Amphidama Boym Lam., Dksh. Conchy. PL 8 bis,
figs. 6—8, &c.
TrtgoneUa Da Costa, Desh. (Species of Mactra Gmel., sub-
genus of Mactra Cuv., spec, of Lutraria Lam.) Shell orbiculato-
trigonal, flat, subequilateral, with anterior side shorter, rotundate,
posterior subhiant. Hinge narrow, with two small teeth in right
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CONCHIFERA. 74&
valve, single tooth in left; a triangular pit receiving internal liga-
ment. External ligament small. Animal with tracheae long,
slender, separate, very nnequal.
8p. TriffoneUa plana Lovixr, Maclra piperata Gmkl., Lutraiia compreua
Lax., Blainv. Malae, PL 77, fig. 2; Babtbb Natwtrk. UiUp. n. Tab.
ym. figs. 5 — 7. This species keeps itself deep in the sand on sea-coasts,
and also occurs fossil in the Crag-formation.
Cumingia Sowerby.
8p. OunUngia UUinMei Bbsh. Oonchylidl, PL 8 bis, figs. 9 — 11 ; all the
species exotic.
Mesodeama Desh. Shell transverse with anterior side longer,
thick, shut. Ligament only internal, received in a narrow, deep
pit. Hinge with one or two teeth under the point, and two
lateral. Animal with two tracheae separate {Peroncea PoLi).
Sp. Meaodetma Qiuoyi Dbbh., Conchyl, PL 10, figs. 13, 14; — Metodesma
cornea Dksh., Amphidetma donaciUa Lah., Poli Testae, ulr. SicU. Tab.
ziz. figs. 8 — II {Mcietra cornea, habitus almost of Donax).
Onathodon Gray, Rangia Desmoulins.
Sp. QnaJ^hodon euneatut Gbat, Dbsh. Conchyl, PL 10, figs. 9—11. Comp.
Bang, Nouv. Ann, du Mue, ni. 1834, pp. 917 — 330, PL 11. This species
lires in the brackish water of the great lake Pontehartrain in Louisiana.
AnatineUa Sow.
Sp. AnatineUa SOtbaldii Sow., Mya CHBMir. Conchyl. Tab. 3, figs. 17, 18,
Dbbh. Conchyl, PL 8 bis, figs. 4, 5.
Mactra L. Shell transverse, inequilateral, subhiant at both
ends. An excavated process under the points in hinge, receiving
the internal ligament ; cardinal tooth bifid or forked in front of pit
for ligament. Animal {Calliata Poli) with trachea coalesced, foot
compressed.
Comp. L. Sfbholbb BederiveUe o>ver del Slaegt Mactra, Shrivter of
natwrhid, Selakabet, v. ). Kjobenhayn, 1802, pp. 93 — 128.
Madl/ra Lam. Shell trigonal, tumid at the points. Two lateral
compressed teeth in left valve, received between two lamella of
right valve.
Sp. JfcKrfra 9iMonm L., Poli TetA, ulr. Sic, i. Tab. 18, figs. 10— n, Blato v.
Malac, PL 73, fig. 5, Ouv. R. Ant,, id, iU,, MoU, PL 107, figs. 2, «a;
brownish grey, with light radiating stris, purplish within ; veiy common in
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750 CLAflB XIL
tke Kortk Sea ml MwBiemiMiw Mmetrm kdwmem Cbemx^
MmOrm gUmem SnsQUOL, Pou, L L figm. 1—3, Stsbgub L L Tih. m.
%. I ; m the XedhemnemB. TUa genus coanta TCfj mjoiy Bpeam fron
allaafl. Some IomB qpedei of h ako «c knpwa, whiA oceor in the ter-
tiaiy farnutkafc.
Lutraria Lax. (in part), Debh. Shell tnnsrerse, elongate pos^
teriorl J, flattened, lateral teeth nona
Sp. ifocfra lutrmria L., Luhyanm dlifiica Lax., Chdci. Cbudlfi. TUb. 24,
figs. X|Oy X|i, &C. Him ■ob-genna ■ znach leas mmMroiis tlun tbe pre-
ceding. The apedei died occvr alao fbaafl in tcrtijay formatlnna
Familj XIX. Myacea. Mantle sacciform, with lobes almost
entirelj conjoined at the margin, perforated anteriorly alone hj a
small aperture for the transmission of foot. Foot small. Tracheae
mostly coalesced, retractile. Shell mostly gaping. ligament
internal, inserted in a tooth or spoon-shaped process, somewhat
prominent at the hinge. Two mnscnlar impressions remote; im-
pression of mantle mostly excised posteriorly by a sinns.
Phalanx I. Osteodeamaoea. Cochleariform tooth in each valve,
receiving the ligament Mostly a loose calcareous particle {osaku"
lum Desh.) contained in ligament Two trachess often separate.
Cardaia Desh. (Is this its place?)
NdU. — ^Habitiia of Jmxardia, Spec Imeardia Memi-mleaia Lax. ud
Bomeothera; two an Ibaafl from the more reeentstnU. Comp. Dbbhatu
in Laxabck HiM, not, da Ani. «. v. 4dL 2, vi. pp. 448 — ^450, and GaiBi^
MagoM. de Zool. 1844, PI 99 — lot.
Myochama Stutchbuby. Shell affixed, irregular, ineqnivalve.
Animal with short separate trachea.
8p. Mfoekama mmnicidei Stutchburt, Dish. OonekyL PL 8 hla, figs, th
13, Ac
Thracia Leach, Desh. Shell free, inequivalve, transverse,
truncated posteriorly, hiant at both ends, with umbones tumid.
Ligament double; external smalL Animnl with two trachea
shprt, separate.
Sp. Thracia eomexa DssH., Blad^?. Malaeol, PL 76, fig. 6, Disa. 0»-
ckyUU, PL 9, figs, i--^; in the N. Sea» Mediter. Ac. There are fa^
apeoiea of this genus in the Jura- and ChAlk-fonnationB^ and in the tertiai7
stnia.
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CONCHIFERA. 751
Anatina JjAU. Shell free, subequivalve, transverse, thin, fra-
gile. Impression of mantle sinuate posteriorly. Animal with two
tracheae coalesced.
Sp. AnaUna tulrottrata Lam., Solm anatwiu "L,, Bumfh. Amh, RarUeUk.
Tab. 45, fig. o, Blainy. MaLac, PL 78, fig. 6, Desh. ConchylioL PI. 8,
figs. 20 — 33 ; in the Ind. Ocean, &c. Also of this genus fossil species
occur in different secondary and tertiary strata.
On the animal of this genus of oonchifers, oomp. H. Mittrb Mhn, ». I.
gewre Anatme, Gu^bin Magtu, de Zool, 1844, MoU, PI. 102 — 104.
Feriploma Schumacheb, Desh.
Osteodesma Desh.
Lyonsia Tubton, Desh.
Sp. Ancaina norvegiea Sowsrbt, Amphidesma eorimloidet Lam., Mya nor-
wegiea Ghbmk. Conchyl, Tab. 170, figs. 1667, 1668, Desh. Conchyl, PL
8, figs. 13 — 14.
Myadora Gbay.
Phalanx 11. Myacea. Ligament internal, fixed to a fovea of
each yalve or to a cochleariform process of one valve and a fovea of
the other. Trachesd coalesced into a fleshy thickened cylinder.
Pandora Brug. Shell inequilateral, thin, inequivalve, right
valve flat, left slightly convex. Hinge in flat valve guarded by an
exsert tooth, other valve emarginate to receive the tooth. Ligament
received in a pit behind the hinge on each side.
Sp. Paiiidora rattrata Lam., TeUina mcequivalvU L., Blainy. Mtdaedl, PL
78, fig. 5, Guy. B. Ant,, id. ill., MoU, PL no, fig. 3; Medit.; this species
occurs fossil in the orag-formation of England. There are also two other
fossil spedes from the tertiary formations, and some from the Indian Sea
known.
Corbula Bbug. Shell inequivalve, with right valve more con-
vex. Cardinal tooth in each valve conical with lateral pit. Liga-
ment inserted in tooth of left valve and in pit of right.
8p. Corbula nudeua Lam., Mya ificequtvahis Moktaou, Encyd, ffUth,, Vers.
PL 200, ^. 4, Dksh. Ocmehyl PL 8, figs. 7—9; in the N. Sea and Medi«
terianean. Within the last few years many species from all seas have
become known; the fossil species are even more numerous, and occur
some in secondary, but in great part in tertiary strata.
ErviUa Turton. Shell inequivalvft Other characters almost
those of Corbula,,
Oomp. Beoluz in GuiBiK Magas. de Zod, 1844, PL 95, 96.
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752 CLASS XII,
N<Ae. — Genus Neoera Gbat, related to the Corbtdce should, on uooimtof
the ossicle of the ligament, be removed to the first phalanx of this family.
My a L. (exclusive of several species). Shell subequiv^Ye,
transverse, hiant. Left valve with single cardinal tooth large, pro-
minent within towards the right valve, and received in the fovea of
this. Ligament inserted in tooth and cardinal fovea. Lnpression of
mantle emarginate with a deep sinus posteriorly.
Sp. Mya trwicaia L., Chexn. Conckyl, Tab. I. figa. i, i, Diae. Ora«4^.
PL 8, figs. I — 4 ; Mya armaria L., Bastkb Naiuurk, Uiisp. n. bL 71—76,
Tab. vii. figs. I — 3, Blainy. Malaeol. PL 77, fig. i; on the strand of the
N. Sea at Katwijk, Scheveningen, the Swedish coasts, &c. — ^l^ere are
some fossil species of Mya in the newer tertiary formations, in which alao
the two living species mentioned are met with.
Family XX. Pholadomyacea. Mantle sacciform, produced
posteriorly into two elongate conjoined tracheae, perforated ante-
riorly by a narrow aperture, emitting the small foot. Shell hiant
at both ends. Ligament external. Two muscular impressions
remote.
Pholadomya SOWERBY, Desh. Shell thin, transverse, longer
posteriorly, obtuse anteriorly. Hinge edentulous, inflected. Im-
pression of mantle excised posteriorly by a deep sinus.
Sp. Pholadomya Candida Sow.^ Desh. Conehyl, PI 4, figs. 4 — 6; a W.
Indian species still extremely rare in collections. The fossil species of this
genus, that occnr in almost aU geological formations, are numerous, moit
so in the Lias. To these also Dbshatjes refers the genera Lynanatta Mukn-
BTSB, AOABSiz, and Platymya AoASSiz. Comp. AOABSiz JBhtdet criL tur
les MoUMqv€tf 2elivrai*on, Keufchatel, 1841, Ac.
Ceromya Agass., Desh. (and Qreslyia Agass.).
Fossil species.
Olycimeria Lam., Cystodaria Daudin. Shell thick, trans-
verse, inequilateral, shorter posteriorly, rounded at both ends.
Hinge edentulous. Impression of mantle not excised posteriorly,
very distinct, with margin incrassate, undate.
Sp. Cflycimeris SUiqua Lam., Mya iUiqua SPEiraLXB, CmexF. OmekyL Tsb.
198, fig. 1934, BLAiir7. Malac, PL 80, fig. 3; comp. AuDOunr Mit^- 'w
P Animal de la GlycinUre, Ann, det Sc. nai. Tom. 38, 1833, pp. 33 '"3431
PL 14, 15. Hitherto the only known species of this genus; on the coasts
of Newfoundland.
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CONCHIFEBA. 753
Panopcea M^N. La Ge. Shell transverse, obtusely rotundate
at both ends, longer posteriorly. Hinge callous, with accessory
tooth in each valve. Impression of mantle excised posteriorly by a
sinus.
Sp. Panopcea Aldrovatidi M£nabd, Mya glycimerii Gmkl., Chxmn. Conchyl,
PL 3, fig. 35, Dksh. Conch. PI. 7, fig. i ; in the Medit. Sea. A few other
living species of this genus are known; they are large bivalves that in
some degree resemble Mya, The fossil species occur in the chalk-fonnation
and more numerously in the tertiary, to which Panopcea Faujaaii MiNASD
belongs, according to Deshates merely a variety of Pan, Aldrotfandi.
Comp. M^ABD DB LA Grote Ann, du ifia. ix. 1807, pp. 131 — 136*
PL IX. and VALENOiBinrBS Archives du Mui, i. 1839, PP- '— 33» ^^ '-~^'
Family XXI. Solenacea. Mantle sacciform, open anteriorly,
with foramen for transmitting foot which is large, mostly truncated
at the apex, or terminated by a broad disc. Tracheae very often
conjoined. Shell equivalve, transverse, elongate, gaping at both
ends. Ligament external. Muscular impressions remote.
Solen L. (exclusive of Anatina). Shell inequilateral, longer
posteriorly. Hinge with small -teeth, two in left valve, one in
right received between the opposite teeth, or with single tooth in
each valve. Ligament elongate, often protuberant.
Comp. L. Spekoleb Skrivter af Naiwrh. SeUkabd. Kjobenhayn, 1793,
m. 2, pp. 8t — 114.
Sclecu/rtua Blainv. (in part), Desh. Shell oval, rotundate at
both ends, subequilateraL Ligament large. Animal {HypogoBa
Pou) larger than shell, with foot large, lingulate. Trachea?
long, conjoined at base into a wide pipe.
Sp. Solen ttrigilatus L., PoLi Test, utr. Sic, I. Tab. 12, 13, Blainv. Malac.
^' 79« fig' 4; ui the MediterraDeaD. The fossil bivalve from the tertiary
formation, referred by Laicabok to Soiecurtus strigtdattu (figured Ann, du
Mus. xn. PI. 43, fig. 5) is, according to Deshates, a distinct species, SoU-
cwrius LamarcJni,
SoUfii Desh. Shell transverse, often very long, with hinge
sometimes almost terminal anterior. Ligament elongate. Animal
(ffj/pogcea Pou) with two short tracheae coalesced in most, in
some separate, with foot cylindrical, truncated.
Sazor-sheU, Manches de CouteaUj &c. The ancient names Solen, Aulos,
Donax (PLnnuB fftsL not. Lib. xxxn. cap. 1 1), already indicate the most
common form of these shells. They hide in perpendicular holes in the
sands of the beach, whilst the tubes are turned upwards. Some species are
edible, and are much esteemed by the Italians, as Solen siliqua, according
VOL. 1. 48
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754 CLASS XII.
to Pou. The species are numerous, and the fossil, of whidi some cont-
spond to forms still living, beloDg principally, though not exdosively, to
the tertiaxy formations.
Sp. Solen vagina L., Pou Tett. utr, Sie. I. Tab. X. figs. 5, 6, Blaikv.
Mataeol, PI. 79, fig. 1 ; in this species the hinge lies almost entirely at the
anterior end of the shell, and has only one tooth on each valTe. In other
species it is situated further backward, always however nearer to the
anterior than to the posterior extremity of the shell ; ex. gr. SoUn leyuwun
L. from the Medit., Solen radiahu L., RuMPH. Amh. BariteUk. Tah, 45,
fig. X, from the Indian Ocean, kc,
Solemya Lam. (more correctly Solenamya Menke, Phiuppi).
Shell inequilateral, shorter posteriorly, rotundate at both extremi-
ties, covered with a periostracon, produced beyond the margins of
shell. Hinge callous, edentulous. Animal with trachea ehort,
branchia on each side single, plumose, formed of lanceolate lamellas,
disjoined, disposed in two series. Foot large, cylindrical, termi-
nated by a disciform expansion, fimbriated at the margin.
Sp. SoUmmya mediterranean TeUina togala POLi, i. Tab. 15, ^, 10, Dssh.
Conchy, PI. 3, figs. 15—17. The animal is tgot^ by Dsllb Chiajb
Memorie nUla 9Uma t NoUm, d^i Amim. «. v. Tab. 62, figs. 10, 11, Phi-
LIFPI in WiBGifANH Arckiv flir Natwrgetek. 1835, Taf. nr. s. 171— 176,
and D£8HATB8 in Cuv. R. Ani,, id, UL, Moll. PI. 115. Comp. abo
Dbshatm Eneifd, mith., Ver$. m. pp. 956, g^f, —.Solen auglralie Lak.,
Blaif^. Maiac, PI. 79, fig. i.
Family XXII. Pholadacea. Mantle sacciform, open ante-
riorly, with a fissure emitting the truncated foot. Two fistulous
trachea, very long, conjoined in most, contained in a common
covering. Branchi® elongate, firee at the apex, produced within
the tracheflB. Shell at each end gaping, with no ligament, a
styliform or spoon-shaped process at the hinge in each valve
advancing into the cavity of shell.
Pholaa L. Shell inequilateral, elongate posteriorly, thin, equi-
valve. External accessory calcareous parts at the hinge. Animal
[HypogoBa Polt) with foot short, thick, its disciform apex flat or
excavated. Two long tracheae conjoined.
SUme^horern (0wXdt from ^wX^w, to lie hid in a hole) ; some of these
animals live at the bottom of the sea near the strand; others penetrate
very hard rocks, madrepores, corals, &c. — From the opening at the hinder
end of the sheU turned upwards proceed the two connected tubes into
which the mantle is prolonged. The mantle is thrown over the hinge, and
contains small supernumerary pieces of shell of various form in the difierent
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CONCHIFERA. 756
species. The points of the shell are covered with cftlcareons inequalities,
which are connected by transrerse projecting parallel lines at the inside
with the margin of the hinge.
Sp. PhoUu Daetifliu L., PoLi Test, utr. Sic, i. Testae, mulHv. pp. 40— 50,
Tab. vn. figs, i— 11, Tab. rm. ; in the Mediter. and N. Seas, edible, and,
as is said, of agreeable taste. Plinius speaks of the phosphorescence of
this animal in the dark, Sist. not. Lib. IX. cap. 61 ; — Photos crispaia L.,
Chxmv. Conehyl, Tab. loa, figs. 872 — 874, Pholas costata Lam. {Phol.
costatus L.), Blaint. Malae. PI. 79, fig. 6, ftc. Some fossU species are
known from the tertiaiy formations.
Teredtna Lam. Two valves furnished internally with cochleari-
form tooth, covered at the hinge with a shield, grown to a calca-
reous tube, elongate, conical, closed at one extremity, open at the
opposite. (Fossil species.)
Sp. Teredina penonata Lam., Ann. dm Mus. zn. PL 43, figs. 6, 7, Blautt.
Malae, PI. 81, fig. 5.
Teredo L. Mantle tubular, terminated by two trache» con-
crete at the base, open anteriorly for the passage of foot, short, toun-
cated. Shell equivalve, gaping on both ends, small, covering the
anterior part of the animal like a ring. Animal inhabiting a cylin-
drical tube covered with calcareous substance, and adhering to it
by two calcareous pinnae {palmulce) placed at the base of tracheae.
Pile-vjorm. These animals live in wood, which they perforate in all
directions. The cavity in which they reside is covered with a calcareous
incrustation^ but the true shell is bivalve and much smaller than the
mantle. How they penetrate and bore through the wood is not yet
sufficiently explained. They grow in the wood, and do not first enter
it as adult animals, for ^e external aperture, towards which the two
tubes {tracheei) are turned, is too narrow to allow the inclosed animal
to enter, however it may have been able to make itself a way at an earlier
period.
LiNKJEUS placed this genus incorrectly between Serpula and SabeUa
(amongst the ringed worms), and named the bivalve shell the jaws of the
animal. He united aU the individuals that had been described up to
his time under one species. Teredo navalis, which is a collective name.
The species even now are fg^ from being sufficiently distinguished. See
Spenoleb Shrivter of Naturk. Selskabd, n. i. Kjobenhavn, 1792, pp. 99
— 106, and QuATBEFAOBB M4m, sur le gen/re Taret, Ann. des Se, not.
^hme S^rie, Tom. zi. 1849, ^ooloffie, pp. 19—73, PI. i. n. The ana-
tomical investigations of the writer named last, are the latest and the most
complete : of the internal structure, to say nothing of older writers, notices
in modem times have also been given by DebhatEB, in the Eeploration
tcientifique de VAlg^rie (1846), and Fbbt und Leuokabt (BeMtge tur
48-2
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756 CLASS XII.
Ketudmm vtrMbfO* mere, Bramnehw. 1847, 4to, a. 46 —53). Hxy u«t
aeoocdiiig \o QcAlSSrACBB, of dktinct kx, but the made xttfTidnali ve
Biicfa nrtx than the female.
For the ^stioetioo of species moune is had siiMBgsi otiher chanetoi W
the form of the two nmll shoTcl-sh^ied eslcareoos pistes whid sie pboed
below on the msntle, si the base of the Uaebea, and are oommonlj named
palmmla. One qwdes, &mons lor the injmy it caused to the piks cf the
d jkes in Hollaad, in the beginning of the last ceniury €S|>eciaHy, migbt be
named Tendo SeOii {Teredo natniii L. ei anctor. in part^ Tardo iaterw
SnsGLKB). It ii figured in the woik of Sklutb and in BunonACH
Abk, natmrkukfrueker Geffeiutdmde, No. 89. The palmmla are fixed 00 a
short pedide, inverKly triangular, and terminating at the broad end od
each side in a points Piles that had been driven only six or seven weda
previoQslj were seen to be entirelj eaten throngfa by this wonn, sod robbed
of all their strength. In this waj the island of Waldieren was in 1730
threatened with destmciion. From time to time the same misrhii^f vm
discovered in other places, especially on the Zuidersee near MedembCk,
Lambertsliagen, Ac ; Wesi-Friesland was forced in conseqaenoe to vaatk
its dykes with large stones, whidi being brcm^t into the coontiyfnHn
abroad, occasioned a great expense ; according to Mabtdtxt, Caleekumnt
der Natmur, m. bL 53, f. 5,600,000 were appHed to the first repair of these
dykes. Since the middle of the Isstcentory the mischief has miich diminished,
so that hence may be explained thefiict, that even in the national mnseoms
only a few specimens of Teredo are preserved. This injnry to oar djkee
was the occasion of many writings, whidi however contributed fitUe to
the advancement of science. We soffioe oorselves with citing the large
work of 6. SlLLiUB, Bietor. not. TereditUe. Traj. ad Rhen. 4to. Foanl
Teredinet also are found in the tertiary formations and in the dialk staraia.
Septaria Lah.
Sp. Sepiaria aireneuria Lax., Serpula peiyUudomia L., Buxfh. Amh. Sari-
teiik. Tab. 41, figs. D, K. A long conical tube, with irregular constrictions
and corresponding septa projecting internally; at the thin end are two
calcareous pipes. The description of Bukphiub aheady indicates that the
enclosed animal has much agreement with Teredo. Ihis was confirmed by
the discovery of a similar species in the Mediterranean a few years ago by
IfATRiBON {Sepiaria mediterramea DxsH. Oondkyl. Pi. 3, figs. 9, 10). The
animal with its small shell is contained in the tube that sticks in the
sand. Periiaps this genus might be united with Teredo, as has been the
Indian species figured by Hohb under the name of Teredo gigamieuM (P^^-
Tranead. 1806).
Family XXIII. Tubicola. Mantle saccifonn, cloven ante-
riorly for the passage of a rudiment of foot ; two tracheae conjoined
into a fleshy pipe. Animal clavate anteriorly. Bivalve shell
gaping, without cochleariform process, with hinge edentulous
linear, contained in a calcareous tube, or concrete with the wall or
the tube.
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CONCHIFERA. 757
Oastrochcma Spenglee {Fiatulana Bkuq., Lam., and Oastro-
chcBna Lam.). Shell thin, equi valve, gaping at both ends, with
hinge linear edentulous, and ligament external elongate. Calca-
reous tube clavate, including the bivalve shell. Mantle with a
small aperture for the passage of a short foot, fiimished with a
small byssus. Two trache® coalesced, long, very contractile.
Sp. Gcutrochcena mumia Sfenol., FistuUma elava Lam., Sfbngl. Shritier
af Naturh, Selskab. II. i, PI. 2, fig. i, Blaiity. Malae. PI. 8i, fig. 3; on
the coast of Coromandel ; the tube is free, and gnuns of sand often adhere
to it. Other species, still, live enclosed in holes of madrepores and other
stony polyparies ; in these the calcareous tube was seen above the head,
and they were separated from the preceding as a distinct genus {Gcutro'
chama Lam.). [The name Ckcena substituted for Otutrochcma by A. J.
Bbtziub {Nova Tetlaceor. Genera, Diss. def. a L. Mubntbb Phillipbon,
Lunds, 1788, p. 19), although adopted by Sfsnoleb, has found no fiivour.]
Clavagella Lam. Sheath tubular, calcareous, free or enclosed
in other marine bodies, clavate, and often covered with spines, or
terminated by a cloven disc or calcareous branched tubes. One
valve fixed in the wall of tube, the other firee, contained in the
cavity of tube.
Most of the species of this geous are fossil, and occur in tertiary forma-
tions, as Clavoffdla echinala Lam., Ann. du Mut» xn. PI. 43, fig. 9, Clav,
haceiUaris DssH. Conckyl. PL I. figs. 4 — 10. A recent spedes is Clav.
aperia Sow., Cuv. J2. Ani., id. ill., MoU. PL 117, fig. 1, from the Mediter.,
also Clofvag. lata Bbod., of which the animal was described and the anatomy
given by Owen ; see W. J. Bbodbbip and B. Owbn in Tranaacb. of the
Zod. Soc. I. 3, 1835, pp. 161—^71, PL 35, figs. 1—8, PL 30, figs. 8—16.
AspergiUum Lam., FentciUtis Brug., Arytene Owen. Bivalve
shell incrusted in the wall of calcareous tube. Tube cylindrico-
clavate, open at the narrow part, at the opposite end closed by a
disc supplied with numerous tubular foramina and a central fissure.
Sp. AspergilluM Javanum Lam., SerptUa penit L., Ruhph. Amb. RarUeiik,
PL 41, fig. 7, BLAnnr. Malac. PL 81, fig. 2; from the Indian Sea;—
Aapergilhim vagmtferum Lam., Arytene vaginifera Lsuok., Bukfpell AU.
za der Reite im nSrd. Afrika, wirbeUote Thiertj 1828, Tab. xn. ; in the Bed
Sea. These animals live in the sand of level beaches and banks, and fix
themselTes in it with the broad end of the tube. The animal, which is
dub-shaped, became first known through Busfpell's TraveU,
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CLASS xm.
MOLLUSCS (MOLLUSC A).
Although the name Molluscs has a more general signification,
and is also so used by us in this work, yet we prefer employing it
in a more limited sense for a class of animals which, in the former
edition of this handbook, we named cephalophorons molluscs.
Names of classes ought in our judgment to be short and not to be
confounded with definitions or descriptions. Lamarck too has
already giren the name of molluscs in the same sense to this divi-
sion of the animal kingdom.
The animals of this class have a head more or less distinct
from the rest of the body. This head usually contains special
organs of sense for touch and vision, sometimes even for hearing.
Many of these animals have a shell, others are naked. Most of
them live in water, but some on land, which in the two classes
immediately preceding is never the case. However there are
amongst these animals very different degrees of perfection in the
organisation, yet we observe the same difference more or less in
other classes also of invertebrate animals, nay even in the last class
of vertebrates, that of fishes ^ Accordingly we must here trace the
principal differences in the arrangement of organs according to the
natural groups and families.
The oral cavity of molluscs forms a very muscular expansion,
at the base of which lies an organ, usually named tongue, which is
covered by little teeth or hooks placed in transverse rows. In some
this tongue is short and broad, in others ribbon-shaped and long*.
Above the tongue is a transverse homy plate with projecting lines
^ Here let it suffioe to point to Myxine, and especially Amphtoxfu; oomp. also what
we said above, pp. 34, 35.
* Compare on this eubject F. H. Tbobohbl Ueber die MuncUkeUe eMeimu^er
Schnecken, Wib6Makn*8 ArcKiv, 1836, Bd. I. a. ^5 7—179, Tkf. Dt. x., and especially
LoviN in Qfvenigt of Kongl, Vetentk-Akademiena Fl^rhandlingar, 1847, PP* i75-~'99»
who has described and figured these teeth in very many genera. That these hard parts
consist of silex was observed by Hanoook and Ehblston in SoUt, Ann. qfNai. Jfift-
XV. 1845, pp. 9, 10.
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MOLLUSCS. 769
{Helix f LinMx), or at the side of it two jaws are found. The
intestinal canal has varioos lengths, but is nsoallj tortuous and
longer than the body ; it is very long, for instance, in Chiton and
Patellay and makes many convolutions which are straitly enclosed by
the liver. The oesophagus is sometimes expanded at the lower end
like a crop, as in Limnceus, Planarbis, Nautilus and Octopus^ which
has longitudinal folds like those usually occurring in the oesopha-
gus. The stomach is in some, as in Helix, membranous, in others
muscular. In many this muscular stomach is armed internally
with hard parts, as in Scylloea, Bulla, Bidlcea, or the inner mem-
brane is homy, as in Tethya and the Cephalopoda, just as it is in
gallinaceous birds. There are often found pieces of Crustacea in this
stomach, whence it may be concluded that a hard covering of this
kind is serviceable for crushing hard food. In some the stomach
is compound, as in Onchidium and Aplysia, in which three (or
in the last named genus, according to Cuvier, even four) stomachs
are present. The first stomach is here membranous, the second
muscular, armed internally with larger cartilaginous teeth placed in
circles alternately, and with a single row of smaller teeth at the
upper edge ; the third stomach is armed on its inner surface with
hooks which are curved forwards; the fourth, finally, is an elongated
blind sac, which is surrounded by the liver. According to Owen,
this blind sac may be a secreting organ, a pancreas in the simplest
form, corresponding to the coecal appendages at the inferior orifice
of the stomach in fishes^. As such also he regards the so-called
second stomach in the Cephalopods, a membranous blind appendage
behind the" muscular stomach, which in Sepia and Octopus incloses
a spiral membranous valve, and in Nautilus is parted into many
chambers by thin transverse membranous valves". The termina-
tion of the rectum in molluscs is usually situated on the right
side of the body, always in the neighbourhood of the respiratory
organs.
Salivary glands, which are wanting in the preceding class,
appear very constantly to be present in this. Commonly there are
^ Lectures on the compar. Anai, of the invert. Anim. p. 300; sec. ed. p. 557.
* In some cephalopods there are foUiculi, larger ihan the liver itself, which are
attached to the gall-ducts; Huntbb r^rarded these as pancreas (Pkfsiol. Gatalogae I.
p. 329), in which opinion V. Suebold also participates. Lekrh. der vergL Anat.
». 393-
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79b ci^ias xiu.
twrsf <diaem.h^ in BMajr Ccphalopods mdaome Cfasten^oda, as
ia fcarirtlMi, tLtere are £?«r; m this cne tibe anterior pair is atnatei
T«fx c>»e t> lie mi'znih^
T^ lircr is acTiicL derel-iped. A gall-bladder is not preaent in this
clafls. asid i» secret:- o <4 bile is not effioded fitnn venous hai from
arterial U.-od. In A/ur the liTer is diiided into four Imge lobes.
In TaiaoiRa Ht lirer is doaUe, one on each side, w^hilst there are
tlrae of tlrm in Of^AHimm Uromii^ each with a distinct gaU-doct;
two of tLe.*e docta implant themselTes in the oesophagos, so that
tLe bile maj act opon the dijme as soon as it is in die stomach.
In manj CepLa!*>{Mds also the lirer is formed of aeveral lobes
distinct frc*m each other. The colour is yellow-brown or, as in
Aaif/i7tff , dark wine-coloured red. The finer stmctme of the liver
cfAisUts of nomer^os blind branched follicles which are ooveied
intemallj by a stratum dT bile-forming cells'.
The urinary secretion is not wanting in Molluscs, although it
has not jet been determined in all of them what organ effects it
Jacob^jS first obe«erTed that in the calcareous sac of Hdix and
LimaXy which CuviEH had named the mucus-secreting organ\ nric
acid is present, and that this part ought to be named the kidney of
Molluscs'. This organ is a sacciform cavity provided internally with
numerous folds or laming whidi is situated near the heart and has
an efferent duct that terminates close to the respiratory i^xertore.
In the Ctenobranchiates, as Murex and Bucdnumj the same is
described as the organ which secretes the purple fluid, and the dact
opens into the branchial cavity. In the cephalopoda the spongy
appendages of the venous stems are to be regarded as kidneys^
[In several families of this class it has been shewn that in
those molluscs which live in water, the urinary organ performs still
^ Comp. J. Fbavk J>e ffepate MoUuiconun, Berolini, 1844, 8yo, T. F. G. Schlbu
(•M ftbo^e, p. 605), H. O. LiKDiTKR NonnuUa de Hepaie H Bile everiArtOorum, Boo-
liiii, 1844, Sfo, H. Mboeel in Musllbr's Arehiv, 1846, a. 9 — 12.
* Organe dc la tntcoiiU, aee Ann, du Mus. vu. p. 165, PL 8, figt. i^ 3, 4* ^> *>
PI. 9, fig. 8, b b, fig. 10, c {Mimoire nir let MM, No. xi); M. Lister described the
Mme orgAn under the n^me of viteui Hnerewn, ExercUat, anat, de Ooddeit, Londini,
1694, Tab. I, fig. I, d.
* Journal de Physique, Tom. gi, p. 318, MiOKXL*8 Archiv, vi. i8ao, pp. 370, 37' >
Bidrag tU Bdddyrenet AnaUmie og Phytiol,, Kjdbenhayn, i8a8, pp. 34— «8.
^ See the fiill account of obeervations and opinions relating to thia subject in
v. BisnoLD Lehrh. der vergl. Anat, s. 391 — ^401.
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MOLLUSCS. 761
another fdnction, viz. that of introducing water for its admixture
with the blood. In the Pteropods, in which HuxLET first observed
it, Geqenbauer has shewn that this sac with thick spongy walls,
always seated within the mantle and in the neighbourhood of the
heart, communicates on the one hand with the respiratory cavity
by an aperture provided with circular and longitudinal muscular
fibres, and on the other with the pericardial sinus by a tubu-
lar opening, which has a sphincter muscle and cilia at its ex-
tremity*. In the Heteropods it has the same relations with the
branchial cavity and the pericardial sinus' ; except that in Firi-
loidea^ which has no respiratory organ, it opens externally on the
right side of the body near the vent'. In the Ctenobranchiata
Leydig has shewn in Paludina vivipara, that the triangular kid-
ney, situated near the heart, and receiving venous blood on its
passage to the gill, communicates with a wide space which is at
once its expanded duct and a sac receiving water from the gill-
cavity by a minute aperture. The water of this sac contains blood-
globules, so that a direct commtmication between the water and
the blood may be presumed to occur in the substance of the kidney^.
In the Oymnohranchiata, on the other hand, the kidney is de-
scribed by Hancock and Embleton as a branched, tubular,
spongy organ, inextricably connected with the liver on which it
lies, and opening by a duct near the vent, but presenting no other
aperture. It receives, according to these writers, blood partly fi:om
the aorta and partly from a pulsatory sac which delivers its con-
tents also to the liver. They regard this sac as a portal heart to
which the pericardial sinus serves as an auricle*; though by others
it is regarded as the kidney*.]
The heart is constantly arterial, that is, it receives the veins of
the respiratory organs and gives oflf the arteries of the body. In
^ [HuXLBT Morphology of Cephalous MdUuaea, PhU, Tram. 1853, P- 43> (^kc^ei^-
BAUEB Untemteh, Ac. pp. 10—13.
* HuzLST 1. L p. 61, Gbobnbaueb 1. L Atalanta, pp. iii, 113; Carmaria, pp.
148, 149, and ZeiUch,/, wiuentch., Zool. V. s. 115.
' HuxLXT 1. 1. p. 33, Lbuckabt UnUrswh. in. 1854, pp. 55—57.
* Lbtsio Zeitsch.f. Wisaentch,, Zool, n. pp. 175, 176. PI. xin. fig. 49.
* Hancx>ok and Embleton Anatomy of Dorit, Phil, Trans, 1851, pp. 116— a «8.
' Lbuokabt Zoolog, Unlertuch, in. 1854, p. 55.]
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762 CLASS XIII.
0ome {Hcdiatis, Crepidulay &c.) the rectuni, as in most of the li-
valves, passes through the heart Usually the veins of the lespi-
ratorj organs are collected in a membranons auricle (olrnim), to
whose broader part the broad part of the conical ventricle of the heut
is attached. In most of the Cephalopods three hearts are present;
namelj, two lateral hearts that drive the venous blood to the gills,
and a larger arterial heart placed in the middle, that receives the
veins of the gills. The blood of molluscs is whitish ; it is brown,
sometimes green or red in some Gymnobranchiatay where the oolonr
of the body often depends upon that of the bloods Blood-cor-
puscles are present in smaller quanti^ than in vertebrate animals;
they are round, in some degree flat and often filled with little
granules.
The venous part of the circulating system is always more or less
imperfect. Cuvibr had noticed in Aplysia as early as the beginning
of this century, that, in place of a vena cava and of branchial arteries,
spaces are present, which are perforated like meshes and communi-
cate freely with the cavity for the intestines*. This disposition
was, however, regarded as an exception. Within the last few
years it first became apparent, especially £rom the investigations of
Milne Edwards, that a large part of the circulating system in all
molluscs is formed by interspaces, which surround die different
organs of the body and are bounded by no special walls'. The
venous blood bathes the viscera, and is received into the cavity in
which these are contained, as into a sinus^ before being distributed
to the respiratory organs. The vessels which conduct the blood to
these organs (the branchial arteries), do not originate in capillaries,
but have oftien very conspicuous apertures ; in some the venous sys-
tem appears to be entirely absent, with the exception of the branchial
veins which, conveying arterial blood, run to the heart\ In some
1 E. FOBBEB in AnnaU of NcA. HitUiry, vi. 1841, p. 317.
" Ann, dtk Mus, n. pp. 499, 300, Mhn, #. I Moll, No. 9, PI. n. fig. h «• ^
fig"- h 3-
' [See note a, p. 7x0.]
^ See MiLKi Edwards Ann, de» Se, nai., ^\hme S^r. Tom. m. ZooL 1845* PP*
^89— 315* pp. 34J— 353» Tom. vin. Zool, 1847, pp. 37—76. In Octopus the vMoet*!
cavity, in Loligo the cavity surrounding the muscular bulb of the mouth, forma a venous
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MOLLUSCS. 763
molluscs the arterial system also is imperfect, especially in the
nudibranchiates*.
Some molluscs breathe atmospheric air by a cavity rich in
vessels ; others, and indeed the greatest number breathe, as fishes
do, the air diffused through water by means of gills. The form of
the gills is very various, as will be explained in the view of the
different families in the systematic arrangement. In those that have
a shell, this is a special protection for the respiratory apparatus,
and is usually situated, whenever it does not cover the entire animal,
upon these organs.
In the organs of propagation in molluscs great variety prevails.
Some are bisexual, others have distinct sexes.
In those molluscs in which the sexes are distinct, no external
difference is on the whole to be observed ; an extremely remarkable
exception to this is however formed by the genus Argonauta
amongst the Cephalopods, in which the male individuals have
hitherto been entirely unrecognised, as will be explained in the
sequel when we treat of this genus. In some there are no external
sexual or copulating organs present, and here, as in the bivalves, the
sexual difference can alone be recognised by microscopic investiga-
tion of the content of the secreting organs, which has been done
within the last ten years in Chiton^ Patella^ and Haliotis (the cych-
hranohiaia and aapidohranchiaUi)^ which were formerly supposed
to be all bisexual or rather female*. In others, on the contrary,
a copulation occurs, at least an approximation as in the Cephalo-
pods. Here there is a penis also, which in Buccinum is of
unusual size, in the Cephalopods short only. The testis or the
ovary is single and lies in the Gasteropoda dose to the liver. In
Nautilus the oviduct has a free opening on each side, and thus the
eggs do not come, as is elsewhere the general rule with invertebrate
animals, immediately by a closed canal from the ovary.
Amongst those molluscs which are hermaphrodite, to which,
besides many Gasteropoda, the Pteropoda belong, the arrangement
of the sexual apparatus has been best investigated in the genera
Helix and Limax, In these animals, backwards close to the liver
^ In HaUotia and PateUa Militb Edwabds found an arterial cayity in thfi head, in
which the large artery ends freely forwards.
' Compare what we said above on a similar opinion respecting bivalYee, p. 7i3*
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764 CLASS XIII.
a gland is situated, which consists of many long blind sacs that
unite digitally, whose branches or roots in consequence unite to
form a single efferent duct. This duct then runs along the inferior
extremity of another larger gland, elongate, and often ending like a
tongae ; this last was regarded by CUYIER as the testtSy and the
smaller compact gland as the ovaiy. An exactly opposite opinion
was advanced by Tkeviranus*, which however was contradicted
by Carus, who found eggs in the first-named gland*. With this
other observations again were in conflict which proved the presence
there of spermatozoa. The matter seemed inexplicable, until its
true bearing was illustrated by the microscopic investigations of
H. Meckel'. The gland which Cuvier names ovary, Tretira-
NUS testis^ is both one and the other ; it is a double, truly herma-
phrodite gland ; every foiliculus is double, and within the external,
which contains eggs, an internal is enclosed with seed and sperma-
iozoa in different states of development. The tongue-sbaped organ,
regarded by Cuvier as testis, is a gland secreting albumen, and
covering the eggs with its secretion. Hence this organ becomes
tumid subsequently to copulation, as on the other hand the true
sexual gland, according to CuviER the ovary, after the time of
copulation shrinks up\ The penis is a long filiform organ, capable
of eversion, which in copulation is received within a tube that ter-
minates at the end in a round vesicle ^
1 Ztitadkr. fm- Pkynctogie^ I. 1814, b. 1—30, Ueber die Zeugunffitheik wd dU
Fofiptansumg der MoUuaktn. So also PazvosT, who found ipermatoeoa in the compact
glMid. M^. delaSocde Pkfti^me H ^SimL nai. de Otnive, lY. v. tnd Anm^daSc^
mai. XXX. 1833, pp. 43~59» ^^ 5-
* Cabcb Beitrage twrgaunurn Kenntnim der OetehUckistheiU einigtr GtuUropodtn,
MmcLLKR's Arekir, 1835, a. 487—499, Taf. 11, figs, i, 1.
* MUBLL£B*8 Arckir, 1844, s. 484, Tkb. xiv. fig. 18.
* On this account especaaUy the opinion of Cttrb was contested by M. 6. YxBLOKX'
in his priae essay, Comwteniatic de Orgamie gemerationu m MoUmtcit ffotterop. jmeu94>-
nieis, which contains sereral good figures and accurate descriptions with an laatorial
review, very complete up to that time of the different previous observatiGiif «d<I
opinions. Amm. Acad. L, B, 1836, 1837. Comp. also A. Paabch dt OatieropoAm
nonnuUorum kenmapkrodiiieorum s^ttewuOe geniiaU e( wropoitieo, Berolini, 1843, 8to.
' This vesicle was aleady compared by Dbshatks and Prkvobt to thepodU cop*-
kUive of insects (see above, p. 366). The last writer established by actual obeervatioD
that the penis in Limmai k received at the time of copulation by the pedide of tl^B
vesicle. Moreover the puhnonaiy gasteropod molluscs present an anangemeot vbicb
is not easy to reoondle with the opinions of Stbnrbup, whose jn>tk we alhided to
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MOLLUSCS. 765
Most molluscs are oviparous; only few, as Paludina vivipara
and Claustlia verdricosay are viviparous. Development from the
egg begins with a cleaving or separating of the yolk, as has been
already noticed by us in other classes of animals. Presently the
yolk begins after a longer or shorter period to rotate round an
axis, which at first occurs more feebly, afterwards more rapidly and
regularly. With this rotatory motion a progressive one is connected
by which the yolk describes a circle. These motions are caused
by cilia. It is to be remarked, that, according to the observations
of Sars, Lov^n and others, young marine molluscs universally,
though they be afterwards naked, possess a shell which like that of
Nautilus is volute and covers the hind part of the body. Also
these molluscs undergo a remarkable change of form, since, before
tentacles appear, a large thin membrane, divided into two lobes,
surrounds the mouth. This part [velum) is surrounded by cilia, and
is the principal organ of motion. In some molluscs this organ is
found to be persistent, most conspicuously in Tethia^ where the fine
cilia surrounding the margin are replaced by ctrri^.
The power of restoration is in some species of this class very
great. In various Helices the reproduction of the head that has
been (partly) excised and of antennae has been observed by Spal-
LANZANi, Senebieb, and Bonnet*. That the entire head grows
again, as the experiments of Spallanzani appeared to indicate, may
however be doubted, since the anatomical investigation of such
snails as had been preserved by this observer in spirit afterwards
proved that by the excision the first or cerebral ganglion had not
been removed*.
The nervous system of molluscs presents various degrees of
development in the different orders and families. It may be re-
garded as the general type, that there is a central mass above and
(p. 135), and it ia not apparent from hiB figures why be confliden Tab. n. fig. z, to
represent a female, fig. 1 a male indiyiduai of HdixpomaHa,
^ For the sake of brevity it must suffice that we cite the investigations of C. VoOT,
9wr VembryoginiB des MoUugquea gatUropodea, Ann. dea Sc, not., 30 S^rie, Tom. vi.
1846, pp. 5 — 90, PL r — 4, where, besides, the earUer observations are noticed histo-
ricaDy.
* See of the last-mentioned (EumreB dCHitt, natur. et de PkUoiophie, Syo. xi. 1781,
pp. I — 61.
* ScHWSiooxB NcUwsfeteh. der SkdOU, unge^^. Thiere, s. 639, 630.
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766 CLASS XIII.
another below the oesophagus, each of which consists of two lateral
portions, that either remain distinct or are more or less fused
together to form an over- and an under-lying ganglion. From the
central nerve-mass situated above the oesophagus, or fix)m the two
lateral ganglia when these, as in Patella^ Haliotis^ BuUay &c, are
only united by a transverse commissure above the oesophagus, arise
the nerves of the eyes, of the feelers and of the mouth. On each
side this brain-like central organ is united with the central mass
lying under the oesophagus by one or more nervous filaments (in
Helix by three). The nervous ring, thus formed, surrounds the
oesophagus, and is in many short, because the central masses are
situated near the fore part of the body and in the neighbourhood
of each other ; only in some is this ring very long, as in the -Hete-
ropoda, because the second central mass is situated backwards. In
many nudibranchiates the central nerve-mass placed above the
oesophagus is much developed, and that which lies beneath it
feebly ; in the Ctenobranchiata also the first nerve-ganglion that lies
over the oesophagus is much larger than the second which lies
under it. The converse occurs in the Ptercpoda, where the upper-
most or cerebral nerve-center is ordinarily represented by a trans-
verse commissure alone above the oesophagus. From the central
nerve-mass, situated below the oesophagus, and consisting of a
greater or less number of difierent ganglia, arise the nerves for the
viscera, the respiratory and generative organs. In some several
ganglia or nerve-masses remote from each other are present, as in
Aplysia^ where a distinct ganglion is situated backwards at the origin
of the aorta. The nerve-ganglia are often coloured red, as in the
genus just referred to, in LimncBUs^ PlanorbiSy &c., by a substance
situated under the neurilema. In various molluscs a nervous sys-
tem corresponding to that of the visceral nerves of articulate ani-
mals has also been observed^, of which the anterior portion consists
of small ganglia that lie at the side of the oesophagus, and are con-
nected by nervous filaments with the cerebral ganglion, whilst back-
ward, firom one or two ganglia, branches are given off to the intes-
tinal canal, the liver, and the organs of propagation*.
1 Gomp. above ia the treating of InsectSy p. 378.
> The nervous system has been described and figured in different Gasteropocis, espe-
cially by GuviSB in his MSmoiretpour iervir d VBiti. et d VAnat, da MMaqvu; see
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MOLLUSCS. 767
The organs of sense are more developed, as a role, in this than
in the two preceding classes. The head is nsnallj Aimished (in
most Gasteropoda) on the upper surface with two feelers, on the
outer margin of which the eyes, sometimes on two little tubercles
or pedicles, are situated. When the eye-supporting pedicles are
developed independently, two pairs of feelers are present. In
some these feelers are hollow and can be everted like the finger of
a glove after they have been drawn in, as may be seen in the com-
mon snail. Besides these feelers situated on the head, different
productions of the mantle are probably the seat of a finer sense of
touch, as the ctrri round its margin in Patella and Haliotis. The
folded and indented fringe or circular lip in the Cephalopoda^ the
various feelers round the mouth in NatUiluSy may also be regarded
as organs of tact.
Taste cannot be very highly developed, since the tongue, as we
stated above, is homy. Of the organ of smell nothing certain is
known, although in NatUilus a part, first pointed out by Valen-
ciennes, situated close to the eyes, may probably be regarded as
such. That, however, many molluscs possess the sense of smell
seems to be demonstrated by observations; thus Swammebdam
found that vineyard snails, when he brought fresh food near them,
came out of their house and crept quickly towards it. According
to CuviER the entire mantle that covers these animals may be
analogous to the pituitary or mucous membrane of the nasal cavities,
and thus very appropriately the organ of smelP.
The auditory organ was known in the Cephalopods alone a few
years ago. In the cartilage of the head two small cavities are
found, which enclose a sacculus filled internally with a fluid, whilst
it is surrounded also by a fluid substance and attached to the larger
cavity, in which it is suspended by numerous fibrous threads. In
this sac, in most species, there lies a calcareous round or conical
little stone. The auditory nerve penetrates the saccule, and divides
into fine branches on its inner surface. There are neither apertures',
ako Garneb in Linn. Transact. Tom. xvii. cited above, p. 716. For the nervous
system of the Cephalopoda see below on this order.
^ liCfons SAncU. comp, u. p. 676. Blainvillb^ who regards the antennae of insects
as olfactory organs, ascribes similarly to the feelers of moUoscs the capacity of smeU.
Principes (TAnai, comp.
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768 CLASS XIII.
nor other external parts of the auditory organ present*. It is only
within the last ten years that a knowledge has been attained of the
auditory organs of Pteropoda and Gasteropods ; they are two round
saccules filled with fluid, each containing a small otolite or nume-
rous calcareous concrements, and situated on the central nerve-mass
beneath the OBSophagus, or in some behind the eyes, in which case
they receive a nervous branch firom the cerebral ganglion*.
In most molluscs two small eyes are present, in the Cephalo-
pods alone are they large. They never occur in greater number,
nor dispersed over other parts of the body, but are always situated
on or near the head; in some they are non-pediculate and are
placed at the base of the feeler, or stand on tubercles or sometiines
at the point of the feeler, as we lately stated. In Nautilus they
are seated on little pedicles at the side of the head, like hemispheres
flattened on the anterior surface. In the rest of the Cephalopoda
they lie in lateral excavations of the cartilaginous ring of the head,
protected in some degree by two long, thin, oval cartilagiiioiw
plates in front. In the Oaateropoda^ Swammerdam formerly mves-
tigated the eyes anatomically ; in our century this investigation has
been repeated and extended, especially by Stiebel, Mueller, and
Erohn. Under the skin, that in this situation is destitute o{ pig-
ment, lies the eye-ball, enclosed by a proper membrane {sclerotica)^
which is transparent in front, and sometimes very convex {cornea]-
The vascular choroid, coloured by a black pigment, is furnished
with an annular aperture (just as an iris is). A spherical lens and
a vitreous body occupy the interior of the eye-ball, whilst the optic
nerve expands within the choroid into a cup-shaped retinal
^ See Scarpa AnaA, disquUitumea de Auditu et O^adu, Ticini, 1789, fol. pp. 5) ^1
Tab. IV. figs. 7—9, GuviSB Mim. 9. I. MoUusq, No. i, p. 41, &c.
* See Etdouz and Soulztbt VIntiituL Journal ghUrcde, &c. Tom. VL 1838,
p. 376; Kbohk, Fbobisp^s neue NoHgen, 1840, 1841, s. 310—312 ; V. Sikbold, Wikg-
mann'8 Arehiv, 1841, and Ann, det Se. not. m fiWrie, Tom. 19, 1843, pp. 193— «"»
PL 1 B. These auditory yesicles appear very early in the development of the embryo»
but are at first simply filled with fluid, from which the otolite or the calcaz^ons concre-
ments are separated at a later period only and gradually by crystalization ; see Fut
in Ebiohbok's Arehiv, 1845, s. 117 — 222.
s Swammbbdam B^ der Natuure, i. bL 105, 106 ; Shbbbl in Msokkl'b Ar(^
V. 1819, B. ao6— axo, Tab. v. ; JoH. Mubllbb, Mbckbl'b Arekivfikr Anai, u, PAjsW-
1829, s. 308— 319, Tab. VI. figs. 4—8, Annal. dea Se. naL Tom. i«, 1831, pp. 7-19 J
Kbohx, Muelleb's Arehiv, 1837, s. 479—485, and ibid, 1839, «* 33^— 337» W- *•
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MOLLUSCS. 769
In the Cephalopods, the optic nerre is continued into a large
reniform ganglion, which is surrounded by a white fatty mass
divided into lobes, and is, with the eye-ball, surrounded by a
common case, a fibrous membrane, by which a sac is formed, that
is much larger than the eye-ball, and may be regarded as the cap-
sule {orbita) of the eye. In front this membrane is attached to the
common integument. There the skin becomes transparent, and
sometimes forms, by duplicatures or folds, two eye-lids as it were.
Behind this transparent membrane, perforated by a round opening,
the eye-ball is situated. A cornea is not present, and thus also
there is no anterior chamber, unless the space that intervenes be-
tween the transparent continuation of the common integument and
the eye-ball be so named. Within the eye-case lies a tunic of a
silvery lustre. The eye-ball itself has a cartilaginous external
membrane, which is perforated behind like a sieve by the filaments
of the optic nerve, and in front at the margin of the lens forms the
circle of the pupil. This covering may be regarded as sclerotica.
Within it lies the expansion of the optic nerve, the retina, which
also contains a purple-brown pigment-layer. The lens is large,
elongate, round, and at the posterior surface more convex than in
front^.
The muscles of molluscs are attached in general to the inner
surface of the skin. They do not exhibit the transverse stripes on
the primitive bundles which microscopic anatomy has detected in
the muscles of articulate animals'. Those Gastaropods that have a
figs. 6 — 8. Kbohn confirmed Swamhsbdam's statement of the independent existence
of lens and yitreous humour; whether an aqueous humour also is present, as Swui-
MEBDAM concludes, he leaves undetermined.
^ On the eye of Cephalopods compare amongst others CuYlEB M4m, 9. Ub MoU.
Ko. I, pp. 37— 4i> PL I. fig. 3, PI. n. fig. 5; D. W. S(emhbbino de Oculor, Seetione
horiaoatali. Oottingse, 181 8, fol. pp. 76 — 78, Tab.m.; Krows Nov, Ad, Acad. Oouar,
Leop. Carol, N. C. xvii. i, 1834, pp. 339—366, Taf« a6; DuG^ TraiU de PhyiiU,
foomp. Paris, 1838, 8vo, i. pp. 315 — 318. In i^au^ti«I am not able, any more than
OwiN, to discover a trace of the lens ; that this part should have escaped through the
small aperture existing in the middle of the flat anterior surface of the eye, appears to
me probable. Fresh specimens alone can afford a satisfactory solution.
* That they occur in Sagiita, is in my judgment a fiirther proof that this animal is
no mollusc. In Cephalopods also I cannot any more than in the muscular circular
belts of Salpa (p. 695) discover these transverse stripes, though they have been adopted
indeed in these animals, and also figured.
VOL. I. 49
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770 CLASS XIII,
wreathed shell are affixed to it by a muscle which is attached to
the uppermost turn near the columella, and descends along the toms
to attach itself to the head bj one portion, and to the hinder ex-
tremity of the foot by the other. This muscle, on its oontraction,
serves to draw the snail into its house.
Gasteropods adyance also by creeping on the yentral disc, or
foot, in which they fix alternately the fore part, then the hind
part, to the surface over which they are moving, whilst the disc is
contracted into sinuous transverse grooves that proceed from be-
fore backwards. Pteropods swim by means of the wing^like ap-
pendages that surround the mouth. The same purpose is served by
the flaps of skin on the body of most Cephalopods, which are
especially assisted when creeping on the bottom of the sea and in
other motions, as seizing their prey, &c. by the muscolar arms,
furnished with suckers, that surround the head.
Molluscs, as to their geographical distribution, are not suf-
ficiently ascertained, and it is difficult to form any general con-
clusions £rom comparison, since all coimtries, coasts, and seas, have
not been investigated with equal care. Molluscs are more nume-
rous than conchifers in great seas and on rocky coasts. In warmer
regions they surpass the bivalves greatly in multiplicity of species.
The greatest profusion of marine molluscs, as weU as to genera as
species, belongs, beyond doubt, to the great ocean between the
west coast of America, and the east coast of Asia and New Hol-
land. Tropical forms seem to extend more widely in the southern
than in the northern hemisphere. Amongst the genera that are
most numerous in species, we may here mention especially I\irpura,
MUra^ CyproM^ and Ganus; the species of the three last genera are
almost all from the great ocean, and the East Indian sea. It is
remarkable that the molluscs on the western and on the eastern
coast of South America differ to such a degree, that there is scarcely
a species that is common to both coasts. The genus Chiton, so
numerous in species in the great ocean, is represented by a conple
of species on the east coast of America. Species that dwell at con-
siderable depths are naturally further dispersed than those which
live more in the neighbourhood of coasts. Consequently it is not
sufficient to ascertain the longitude and latitude of the districts
within which certain species occur ; the depth also of the sea ought
to be ascertained in which they commonly live. With the depth
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MOLLUSCS. 771
the number of species decreases, just as that of land animals does
the greater the height is above the surface of the sea. Amongst
the Cephalopods that have a shell, and which occur sparingly in
the present creation, whilst in the previous world they were very
numerous and different in form, Naviilua fs confined entirely to the
Indian and South Seas. Of the molluscs that live on land, no
genus is more generally diffiised than Helix, of which some
hundreds of species are known^ and which occurs in all regions.
The species of molluscs are in a greater degree than those of
bivalves confined to limited regions, and thus, on the whole, differ
more in different countries^.
^ Interesting oooiributions to the knowledge of the geographical dintribution of
moUuBCB have been made m the last few years by the investigations of Sabs, Forbbs;
jyOiLBiavr, Philifpi and othen. Compare A. D*OBBiGinr Reeherchea aw les lots qui
jpriiidetU A la didribuiion giographifue de$ MoUuaques c6lier» fnariiu, Ann, de$ Se, not,
Ze 84ne, Tom. m. 1845, PP* '93 — '>^i ; £• Fobbbs On the iopogra/phietd distrtbvtion
of fMurvM Molluaea, Edinb, Phil. Jowm. New Ser. April, 1844, and Ann, des Se. not.
3e S^r. Tom. iv. pp. 1x7, 118, and JjOvts*B remarks on the geographical distribution
of Scandinavian Marine Molluscs, Cfvenigt of Kmgl. Vetentk, Ahademiens Fdrhand-
linear. 1846, pp. 252 — 274.
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774 CLASS XIII.
GlOSHBAUEB I. L 8. IIO.]
Aooording to CAKTBAnn it does not differ from the preceding genus, the
shell is cloYen at the sides as in HyaUa, L 1. p. 33.
Add genus Ewryhia Bako Ann. da Se. not. zn. 338.
Oreseis Rang. Shell subulate, furnished with an orbicnlai
opening anteriorly, not cloven at the sides. Mantle not producjed
at the sides.
Comp. Bavo Ann, de$ Se. not. xin. pp. 305 — 318.
NcU. — ^Here also or with the preceding genns is to be placed :
Sab-genns Chmeria Bavo. See Ann, de$ Se, noL xn. p. 311, PI 45 b,
figs. 1—8, y. BxvsDXir L L PL 4 b.
SpiraieUa Blainv., Limadna Cuv. Body anteriorly auriculate
on both sides with a triangular pinna, posteriorly convolute. Shell
fragile, discoidal, on one side deeply umbilicate, on the other widi a
spire slightly exsert.
Sp. SpiraieUa areUea nob., Li$naeina heUeinaUi Lam., Blaotv. Malae, PL
48 bis, ^. 5, y. BBKBDSir Mhn, mr la Litnaeina aretiea, Mim. de Vie.
TcyaU de £rux, Tom. XIT. 1841, PI. 5. This little animal is very numerous
in the North Polar Sea, and serves the whale for food ; see O. FABBiaus
Faun, Oranl, pp. 387 — 389, by which writer it is named ArffonatUa arc-
tiea. In Phifps'b Voyage towaids the N. Pole it is notioed under the name
of Clio kelicina, and it is probable that Cuyibb intended to name it Bdi-
cina, a name which we should prefer to that of Limacifui, which perhapi
originated merely in a mistake.
Cymbulia P^RON. Animal included in shell, except two large
pinnae, confluent [partially, with a middle lobe that represents the
foot. Gegenb.]. Two contractile tentacles in front of mouth.
Shell hyaline, elongate, cymbiform, truncated posteriorly.
Sp. Oymh, Peronii Cuv., PiBON Ann, da Mut, xv. PI. 3, pp. 10 — 11, GuiBnr
Iconogr,, MoU, PI. 4, fig. 3, Y. Bbkbdbn M4m, de VAcad, de Bmx. zn.
PI. X ; Mediter. Sea; according to CAirrBADni these animals often lose their
shell. [The shell is formed within the mantle, and normally is covered by
a thin lamella of the same. This last is easily torn and thus perfect speci-
mens are rare : hence also the esse with which the animal loses its shell
The shell has the consistence of soft cartilage and contains ehiiin, GioKV*
BAUBB L 1. pp. 41, 43.]
[Ttedemannia Delle Chiaje. Pinnae conjoined with the foot
to form an orbicular di%c. Mouth at the apex of a long proboscis,
very often bent backwards. Shell hyaline, gelatinous, slightly
excayated.
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MOLLUSCA. 775
8p. Tiedmanma neapoUkma Y. Beksd., proboBcis alender, pinxuD with
white and yellow spots at the nuuigin. Length 2 or 3". See Geoenbaub
1. 1. s. 54 — 68 for the anatomy and developmenty gen. char. p. 113.]
Famaj IL Clioidea. Naked Pteropods, with head distinct.
[The small foot on the ventral sturface never connate with pinnss.
Gegenb.]
Fneumoderman Cuv. Two small pinnae at the sides of neck
with an intermediate grooved conical appendage. Mouth with
numerous pediculated retractile suckers.
Sp. Pneumodermon Peronii Cuv., Ann, du Mus. TV, PI. 59 B, Blainy. MaUte,
PL 46, fig. 4 ; this animal was disoovered in the Atlantic Ocean by PfooN,
and appears to differ from that figured by QuoT and Gaimabd in the
Toyage of the AOroldbe (copied in Cuv. R, ani., id, HI., Mofhuq, PI. 17,
fig. 7), where the Backers stand like branches on two pedides. Pneumo-
dermon vMaoeum D'Obbiont has a greater resemblance to it, the anatomy
of which has been given by V. Bekeden with figures in Mueller's Arehw,
1838, pp. 296—304, PI. IX. X. There is stiU uncertainty respecting the
respiratoiy organs, and the name Pnbuicodbbmon rests perhaps on a
mistake. [Pn, eUiatum Gbobkb., see the anatomy, Ac. L 1. s. 74 — 100.
The folds of the integument^ here absent, which serve for respiration, are
described in Pn, vidaoeum and P. Mediterraneum, pp. 85, 86.]
Spongiobrcmchea D'Ob& (Is this genus distinct from the pre*
ceding))
D'Orbiont Voyage dans FAmir. mirid, (transferred to Okbk's Im,
1839, s. 498, Tab. I. iz. figs. i~9).
Clio L., Cltone Pall., Eschricht. Body oblong, acuminate
posteriorly. Head distinct from body by stricture, covered by a
bilobed hood, crowned by six conical appendages, and two retractile
conical tentacles. Two oval pinnae at the sides of neck. Two
black points (eyes) in the back of neck. Two genital apertures at
the right of the ventral side, the anterior in front of pinnae for
the exit of a very large penis, the posterior behind the pinnae
for the oviduct. Anus behind these orifices, more in the middle.
Sp. Clio lorealU Quel., Clicne lorealis Pall., Spk, Zool. x. Tab. i. figs.
18, 19, CxTViBB Ann, duMw, i. pp. 941 — 150, PI. 17, D. F. Esohbioht
Kangl Danehe Vdentk. SeUikaba Afhandl, Kjdbenhavn, 1838 (published
separately in German, AwUom, UtUerwchungen U, d, CI. boreal, Kopen-
hagen, 1838, 4to.) The six conical arms of the head are beset with spots,
microscopic suckers. This animal (Whale-bait), Clio Umoiema Phifps,
lives with Clio helicina principally in the neighbourhood of Greenland and
Spttzbergen, and forms the chief food of northern whales, although scarcely
i" long. CnvuEB regarded the fin-like appendages as respiratory organs, but
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776 CLASS xiii,
it WBB probably the reticalated mtueiiUr fibers that he took for Tcadi
in these dermal lamells [Clio Mediierranea Gbobvb., tentadet veiy Bkort*
no eye-points; CI. JUiv€9cen$ Gkobnb. G. thinks that the 3,000 sudLcn in
each tentacle described by Ebohrioht are merely forms of tpiikeUuM,]
Order II. Gasteropoda.
Molluscs with head distinct, in most tentaculate, the inferior
surface of abdomen flattened or grooved, or produced into a com-
pressed lamina. Some hermaphrodite, others distinct in sex ; many
aquatic, some terrestrial.
Gasteropoda, In most the heart lies on the left side of the body.
In those, howeyer, that have left-handed shells (see above, p 684)
the heart lies on the right side. In many the sexes are distinct;
others are bisexual, so that mutual impregnation of two or some-
times more individuals occurs.
The inferior surface of the belly forms an elongate flattened disc,
which is very muscular, as is commonly known in slugs and szuuls;
these animals, met with everywhere, give an idea of the typu^ of
this order. But in others, this ventral disc, usually named foot, is
compressed laterally, and serves for swimming. We separate these
molluscs from the rest as a distinct group.
Family III. A. Heteropoda. Foot compressed, resembling a
fin, furnished with a disc or suctorial acetabulum. Branchis pec-
tinate or pinnate. Sexes distinct.
Heteropoda, These molluscs all live in the sea, and usually swim
with the fin-shaped foot upwards, and the back downwards.
FoRSKAL, to whom we owe the first description of this family of
animals, gave them the name of Fterobranchea. The existence of
distinct sexes was discovered by Lavrillard and Milne Edwabds.
[The part named * Foot' is highly developed in this family, and in
some attains a high potentiality (Gegexb.). It does not exactly
correspond to the foot of Gasteropoda, but to one part of it alone.
Huxley {On the Morphology of the Cephaloua MoUuacOy FhS.
Trana. 1853, Pt. l pp. 29—65). A foot has four parts, the/>w-
podium, meaopodium, and metapodium, found in Heteropods, and a
fourth, the epipodmm, not found in them. The fin-shaped M 0^
heteropods is the propodiunu Besides the works just referred to,
comp. also, on this division, Leuckart Zoologiache Unterawhun^
Drittes Heft, Giessen, 1854, Der Bau der ffeteropoden, pp. 1—68.]
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MOLLUSCA, 777
Pterotrachea FoRSK., Firola Brug., PiSron. Body naked, elon-
gate, with integument almost pellucid. Two eyes. Tentacles (very
often) none. Head with a fleshy proboscis. Branchiae pinnate,
free, at the posterior part of back.
Sp. Pterotrachea coronata Fobbk., Icon. rer. not. Tab. 34, fig. ▲ (copied in
Encycl, mith., Vera, PI. 88, Firola, fig. i) ; in the Mediterranean. Lssbukb
has given a monograph of this genus in the Jowmal of the PhUos. Soc, of
Philadelphia, of which Blainvillb has made use in the article Firola,
Dictionn, dee Sc. nai, Tom. xvii. pp. 62 — 68. Usually the tentacula are
wanting. The species in which they are present were united by D'Obbignt
under the name of Cerophora. Here belongs Firola caudina Rang,
Gu6biw Magae. de Zool, 1833, CI. ▼. PI. 3, Gu^bin Iconogr,, MoU. PI. 11,
fig. 3 [For the Anat. and Hist, of Development, see Gxgbnb. 1. L pp.
153 — 185, and Huzlbt 1. 1. on the Anat, of Firoloides {Firola) L L pp.
30 — 36. The auditory vesicle was shewn by Lbtdig AnatonUsch, Bemar-
hung, iib. Carinaria, Firola wnd Amphicora Zeitechr. fUr tnssenech, Zool,
m. 1851, p. 3^8.]
According to D'Obbigkt the singular genus Sagitta (see above, p. 346)
should be placed here.
Cartnarta. Body elongate. Head with two long, subulate ten-
tacles, and two sessile eyes behind the base of tentacles. A small
shell at the posterior part of back, thin, pellucid, conical, com-
pressed, with apex turned backward.
These animals have much resemblance to the preceding genus, and
CuviEB at first thought that the Pterotrachecs described by FobskIl had
only accidentally lost their shell. Mdmairee, MoU. No. 18, p. 31. They
differ, however, by the constant presence of two tentacles in front of the
eyes and by small inequalities or tubercles on the skin.
Sp. Ca/rinaria cymbium {Argonauta Cymbium L.?) Ann. dee Sc, not, xvi.
PI. I, GuiBiN Iconogr., MoU, PI. 11, fig. i, PoLi Teetac. vir. Sic, in.
Tab. 44, figs. I — II ; in the Medit. Sea. Pou and Dellb Ghiajb gave an
anatomical description of this animal {Test. in. pp. 26 — 35, and Memorie
Sulla Storia e Notom. degli Anim. a. v.)^ which was afterwards completed
by MiLNB Edwabds as to several remarkable peculiarities, Ann. desSc. not,
sec. S<«r. Tom. 18, Zool. pp. 313—329, PL X. fig. 3» PL xi. The remark
of liVSTXMVS, that the' shell is not larger than a grain of sand, does not
apply to this species, which therefore has also been often confounded with
CaHna vitrea, a larger species from the Indian Ocean ; Mabtiki, Tab. 18,
fig. 163, Blainv. Malac, PI. 47, fig. 3 a,
Atlanta Lesueur. Head furnished with a proboscis, two cy-
lindrical tentacles, and two eyes. Foot compressed, acuminate
towards the end, with a suctorial disc in the margin. Thin oper-
culum, adhering to the ventral part below the foot. Shell very thin,
involute, carinate.
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778 CLASS XIII.
%>. AiUmia Permii Lbbubub, Cfofne ^ Amman Lamanoit, Vofogt d/t Li
Pbtbouss, pi. 63, figs. 1—3, BLAnry. Malac PL 48 \nB, fig. 9; AUmta,
Keraiudrtnii Lbsueub; this species fonns the genus LadoM CAaT&Aixi,
McUaeol. midUerr. Compare on these small animals tliat oocmr in &e
Mediterranean and in warm sones of the AUantic Ocean, Rako Mim. dt
la Soc. fFffitt. not. Tom. m. 1837, pp. 373—380, PI. 9, and D'0rbig5T
Voyage (see Oksn's In», 1839, pp. 511 — ^519).
B. Oasteropoda namudta.
Foot serving for creeping, flattened or grooved.
Section I. In some no distinct organs of respiration, in others
naked dorsal branchise, appendages or various productions of the
common integuments.
We part the common or normal Gasteropoda into three diyiaions, aoomd-
ing to their respiratory oi^gans, and unite, as was proposed b j the yoonger
Lbuckabt, the Qymnolmmehiata and Apneutta under the name of Jkrma-
tobranckiata of DuK^BiL ; see B. Lkuokabt Ueber Morfihologie der vkMI.
Thiere, 1848, s. 178.
Family FV. Dermatobranchiaia s. Oymnobranchtata, Cha-
racters of the section; Naked Gasteropods, hermaphrodite, marine,
often swimming on the back, with foot supine.
The I^udibranchiates {nudtbranehea) of CuviBR have been eepe-
ciallj investigated in the latest times. By Quatbefages, in many
an intestinal canal divided into branches was discovered, or rather a
branching appendage of the same, such as we have already noticed
in Distama, in some ringed worms, in jlcortno, in Phalangwm, kc.
These blind branches discharge at the same time the office of liver.
(Quatbefages gave to this arrangement the name of Fhlebenterismus
(see Ann. dea Sc not., 3idme S^rie, Tom. iv. p. 83), which has caofied
much misconception.) In some it would seem that no heart is
present j whilst there are such conflicting opinions on this point, ve
consider it inexpedient to found an arrangement upon it, which
might afterwards be subverted, and therefore cannot adopt the
&milies of Anangia and Angiophora (Y. Sisbold Lehrb. d. vargl
AnaL I. a 297). As little do we feel ourselves justified in sepa-
rating the Apneugta from the GynmobranchiatcL Those Gasteropods
alone can be regarded as Apneiuki, which, as Lisaoaoma, have no
external dermal appendages.
Compare Aldeb and Hakoook in dilforent paits of the Aw»ai* wd
Magaz. of Nat, ffia,, and their Monognqih of tke JBrUM Nudibran^i^
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MOLLUSCA. 779
MoUuica, wUhpUUei, fto., published by the Ray Society, London, 1845 —
1854, six parts, also Quatbifaqbs in the Annala des Se. not. since 1843.
A. BrtmehioB none,
Lisaosoma Koelliker and Quatbef.
Limapontia JoHNST., Chaltdts QuATBEF.
Actasonia Quatbbf.
Phyllirhoe Peron. Body naked, compressed, with two very
long, subulate tentacles. (Organs of respiration unknown.)
Comp. PIebon Ann, du Mua, TV, p. 65, PL n. figs, i — 3; QuoT et
Gaimabd Voyage de VAttrdabc, Zoologie, n, 1833, pp. 403—410, PL 18,
figs. 10—18 ; [H. MuBLLKB und G. Gxosnbauxb Ueber PhyUirkoe bucepha-
Itm in KoBLLiKXB u. Siebold's Zeittchr. f, wmmimcA. ZooL y. 1854, PP*
355 — 37 1» ^B^* Xi^- I^^OK placed this animal as well as all the Hetero-
pods amongst the Pteropoda, Lamabok amongst the ffeteropoda, Bang
amongst the Sdlpa, Etdouz and Soulktbt from the structure of the
animal first removed it to the NttdibranchieUa, The form of the nervous
system, the pouch-like appendages of the intestinal canal, and the disposition
of the genital organs, all refer it to this division, although from the com-
pressed form of the body the foot be not distinguishable from the rest of
the integument. In the md Edit, of his work Van dxb Hoeybn placed
gen. PhyUirhoe amongst the Heteropods. He now authorises its removal
to this place in the English Translat. of his Handbook. See the Anatomy
Ac. in the mem. of Muellbb and Gbqbkb.]
Cenia (previously Ictis) Aldeb and Hancock.
Note. — Comp. on those genera destitute of branchiee, the synonymy of
which is very obscure, QuATBEFAOBsifi^m. tier let Oatteropodes phlebenUrfy,
Ann. detSe. not., 30 S^rie, I. Zool. pp. 149 — 183, and J. Aldbb and Han-
cock on a proposed new order of Chuteropodous MoUtuca, Ann. of Natw.
HisUny, sec. Series, 1. 1848, pp. 401 — ^415, PL 19, 20. Genus Pelta Quatbbf.
if in reality it agrees with the Mollusc described by the English authors
A. and H. {Ann, of Not, Hitt, xviii. 1846, p. 289) cannot be united with
these dermaU^anchiata or abranchiaia,
AcUBon Oken, Elysia Eisso, Cantbaine. Body subcylindri-
cal, limaciform, bordered by a depressed expansion, acuminate
posteriorly. Two tentacles auriculate, subclavate, not retractile.
Two eyes behind the tentacles. Anus lateral, (sometimes middle,
posterior?) on right side.
Sp. Aetcdon viridis, Aplytia viridis Montagu, Linn. Trant, Tom. vn,, Quatbe-
FAOEB L L PL 3, lag. 1. Comp. Okbn Zool. i. 181 5, p. 307, and especially
G. F. AUJCAN on the Anatomy of Actceon, Annals of Nai. ffitt. YoL 16,
1845, pp. 145— 16«, PI.5— 7-
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780 CLASS XIII.
Plcicobrav,chu8 Y. Bjlss.
Comp. Van Habsxlt in BvUet, univ. Db FsBUsaAO, 1824, Oct. p. 14a
Scarcely diBiinct from the preceding genua, with which it Jb conjoined by
LoT&sr.
B. Brcmchia eoctemal <U the tides ofhacky nwnyerofu^ teniaadiform,
t Vent paeteriar, in the middle a/back,
Janus Verany, Antiopa Ald. and Hanc.
Comp. QuATRSFAQES Ann. des Sc. nai, 30 S^rie, xi. 1849, pp. 76-— 90,
PI. 3, 4. 8p. Janus Spinola, £olidia ctidaUk Dkllk Chiajs. See also
Ann, qf Nat, Hist, sec. Series, i. p. 190.
Proctanotus ( Venilia previously) Ald. and Hanc.
Comp. Aldsb and Hajbtcogk AnnaU of Nat. HiH. 13, 1844, pp. 161—
164 and p. 407, PL 4. Here also is to be referred as it seems g^enus Zefikf'
rina Quatbbf. [According to Aldbb and Hasoock the two ^jSx in thkt
four linear tentacles are seen from above in Zephyrina, whilst in ProdonotuM
the two larger are dorsal and the smaller arranged at the sidee of the Telum
which covers the head. Monograpk. No. y. Proeton."]
t+ Vent lateral, to the right.
Alderia Allman.
Comp. Allman Ann. of Nat. ffitt. ToL 17, 1846, pp. i — 5.
Stiliger Ehrenb.
Fterochtlus Alder and Hanc
.iSolidta Cuv. {^^lis or Eolis auctor.). Tentacles four.
Angles of foot lateral, the anterior mostly produced. Branchiss
placed at the sides of back, numerous, tentaculiform, emitting
stinging filaments at the perforate apex.
Sp. jBoUdia papulosa, Limax papillosut L., Sytt. not., Bastbb Natvmrk
Uitsp. I. Tab. z. fig. i, bl. 93, 94, Ann. of Nat. HUt. XY. PI. I. fig. h
Aldbb and Hanc. Monog. Pt. yi. Fam. 3, PL 9 ; this liUle animal attains
a length of 1 inches. Comp. Hancock and D. Emblbton on the Anatomy
of Eolis, Ann. of Nat. HUtory, XV. 1845, pp. i — 10, pp. 77 — 88, sec.
Series, I. 1848, pp. 88—105, ^^^^ u. Lbuckabt BeUr. z. KmnJtnim wfMk-
ser Thiere, 1847, "• 54 — <^4* '^^ ccBcal branches of the intestinal caoal
are extended in the Eolidoe into the branchial appendages. These app»-
dages have at the point a small vesicle, from whence filaments resembling
spermatozoa (nettle- threads, see above, p. 99) come to view. Ann. of Nat.
Hist, XV, PI. IV. PI. V. figs. 1— II.
In the neighbourhood of JBolidia some genera or sub-genera of IsUr
authors may be placed, all of which cannot be noticed here. BoiidvM
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MOLLUSCA. 781
QuATBEF. princip«Uy differs by the absence of the Uteral prolongations of
the foot, which Guvikb regarded as a third pair of tentacles in jEolidia,
See QuATBSFAOES Ann. de» Sc. not., ae S^rie, Tom. xix. Zoologie, pp.
971—313, PL II ; oomp. Aldeb and Hanoook AnndU of Nat, Hiti. xiY.
1844, pp. 125—129.
Tergipes Cuv. {Amphorina QuATREP., and Pstlocerus Menke).
Branchiae clavate or obovate, not numerous, placed in a double row.
Tentacles four or two.
Sp. Tergipes laemulatiis, Limax Tergipet Forse., Icon. rer. natur. Tab. XVT.
Hg. E ; Tergipet psilocerua nob. L. BoxxE, Verhandel, van het Zeeuwech
Oenootah, in. 1773, pp. 296—298, PI. (p. 318) fig. 3 ; on the sea-wiers of
the island of Walcheren. (This little animal certainly does not differ
notably from AmpTtorina Alberti Quatb. Ann. dee Sc, not,, 30 S^rie, i.
H. ni. fig. 5, which, however, is smaller.)
Comp. on Tergipes A. Y. Nobdmaitn Versttch einer Monographie von
Tergipes Sdwardsii, Mhn. de VAcad, de St. Petersbourg, sammls Grangers,
Tom. TV. 1845, (given in part in Ann, des Sc. not,, 30 S^rie, y. 1846,
pp. 109 — 160, PL I.) The generic name of Tergipes rests on an observa-
tion that the animal creeps on its back by means of the branchis, which is
very doubtful.
CaUwpoM D'Obbiokt.
Comp. D'Obbiont, Gu&ik Magca. de Zocl. 181 7, MdU. Pl. 108 ; Aldeb
and Hakoook Ann. of Nat. Hist. xii. p. 233, Milne Edwabds Ann, des
Sc. not. ae S^rie, xvin. 1843, p. 33, PI. x. fig. 1.
Hermcea LovfiN,
Comp. Qfversigt af Kongl. Vetensk. AJcad, Birhandl, 1844, p. 50.
CavoUna Brug.
C. BrcmchicR dorsal, cloven, peniciUate or wrboreacent
FlaheUina Cuv.
OUmcus FoRSTER, Body elongate, posteriorly subulate or fili-
form. Four short tentacles. Branchise digitate or palmate, lateral,
three on each side.
Sp. Qlaucus hexapterygius Cuv. Ann. du Mus. vi. PL 61, fig. 11, pp. 427 —
430 (Mim, swr les MoU, No. 6), R. Awi., id, HI., PI. 29, fig. 3 ; although
the figure in Blumeitbaoh Ahh. naturh, Oegenst. No. 48, has eight and not
six branchiffi, QuoT and Gaihabd are of opinion that this species denoted as
Cflaueus aUanOcus cannot be separated from the preceding, and that only a
single species of it is known ; comp. Vogage de V Astrolabe, n. pp. 279 — 283 ;
also G. Bekkett always saw only six branchiffi ; he observed that the animal
attacked and swallowed Porpitce that were placed in a glass with it. The
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782 CLASS XIII.
oolour is purpUBh or blue and ihe length neariy i) incheB. See Prooui.
Zod, Soe. 1836, pp. 113— 189. Theee uiiiiudi an found in the AUaotk
Ooean prindpaUy between the tropics.
ScyThea L. Body oconpressed, with foot farrowed longitudi-
nallj. Head frunished with twotentaclea. Back with two expan-
sions or membranous and flexible wings on each side. Branchis
penicillate, composed of filaments, scattered over back, especially
crowded in the wings.
Sp. SeyUfBa pdagica L., Guv. Ann, du Mum. yi. pp. 416, 417, PL 61, figs.
I — ^, MoUusq,, Mim, vi. ; Blaott. Malae, PL 46, fig. 5 ; this spedei
attaches itself by its furrowed foot to Fueui natant, and is found m the
Atlantic ocean ; ScyU. ghomfodentis observed by FoBaKAi* in the Bed Ses
appears to be the same species. Jkier. AninuU. p. 103^ ^., Tab. 39,
figs. C. c. Anatomy, Ald. and Haitc. Mcnogr. Fam. 2, Fl. 5.
Dot(f Ok£n\ Melibea (or Meltbasa) Bang. Body limaciform,
with foot narrowed posteriorly. Month with a circular velum.
Two filiform tentacles, defended by an ample cup-shaped sheath.
Branchias papillose, grouped in muricate clavae, disposed in a double
lateral row on the back. Apertures of generation and of rectum
at the right side.
Sp. Dato coronata, Dorii eoronaia Gkkl. (species of IWAmmo Cut.), Bojcmi
Verk. van hd Zeemnck Oenoottek, i. 1769, Tab. m. bL 314, eo^ or
erowmL Sec^ilug Johnston, Ann. of Nat, ffiit. i. p. 117, FL 3, ilgs.
5—8, D'OiiBioNT, QufoiN Moffoain de ZooL 1837, MoOmq. PL 105,
Anatomy in Ald. and Hang. Manogr, Fam. 3, PL 4.
Tritonta CuY. Body subtetragonal or compressed, elongate,
anteriorly rotundate, posteriorly acuminate. Apertures of genera-
tion and of rectum at the right side, with vent situated behind the
genital orifice. Two tentacles retractile into a sheath. Circular
velum in firont of mouth. Two lateral maxiUse, acute, with margin
denticulate. Branchise arborescent at the sides of back.
a) WUh arbore$eeiU brancMce remde, and theaih tf tentaelet with mar^
DtndrorwtuB Ald. and HANa
Sp. TriUmia ar&orsMSM Guv., Dofit aii%orueen» Gu., Cuv. Mothuq,, Mim, 6,
figs. 8—10 (probably not specifically different from Dorit eenina GmOm
ike amdered Sea-ilug Bommb, L L ni. bL 390, fig. i). Compare, on tb<
* Lekrh, der ZooUgie, i. 1815, s. 173.
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M0LLU8CA. 783
soond which thu little aaiinal prodnoee under water, GsAlv Atm, dm Se.
not, ym. pp. izi, 113, and S. Bbid Ann. of Nat. Sut. xvn. p. 389.
b) With branchke dupoaed in a continwnu Hiria; vnth cup of tentacUi
etUirt.
Trilonia Ald. and HANa
Sp. TrUonia ffombergii Cur. Ann,du Mua. i. pp. 418—498, PL 31, 33, MoU,,
Mim. 4 ; BiAiHV. PI. 46, figs. 6, &c.
Polycera Cuv. Body attenuate posteriorly with belly flat,
covered by a loose membrane. Dorsal branchias in little branches
or plumose, covered with membranous laminae or papillae. Veil of
head running out anteriorly into several cirri or tentaculiform
laciniae. Two clavate tentacles.
jEgirus Ijoy£s. Tentacles ya^inate, simple.
Sp. Pdycera puncHlueeru D'Obbiokt, Guiaiir Magarin de Zool, 1837,
MoUusq. p. 106.
Pdycera Loven. Tentacles naked, annulate with transverse
lamellsd.
Sp. Pchfcera comuta, DorU cormUa Abildqaaxd, Zocl. Jkmiea, Tab. 145,
figg. I — 3, Blainv. Malac. PL 46, fig. 10 ; aocordiDgtoLoviNl^om^tMui-
rilvmeata Mubll. Zool. Dan. Tab. 17, figs. ^--6, Tab. 138, figs. 5, 6, and
Pclyeera omaita D'Obb., Gubbin Mag. 1. L PL 107, ought to be brought
under the same Bpeciee. Comp. on this genus Aldkb Annals of Nat.
Hist. VI. 1841, pp. 337 — 34a, PL IX., Fbbt u. Lbuokabt BeUr. twr
Kenntniaa vfirhdl. Thiere, s. 66 — 70.
Teihya L. Body somewhat oblong, depressed, famished ante-
riorly with a broad funnel-shaped, fimbriated veil, behind the veil
contracted into a species of neck. Mouth proboscidious, retractile,
in the bottom of velum. Two conical tentacles at the base of
velum, margined by a broad cup-shaped sheath. Apertures of
generation and vent at the right side, in the anterior part of body.
Two rows of branchiae at the sides of back; cirrose pectinate
branchiae alternating with smaller bundles.
Sp. Tdhy leporvna L., Boitdslbt Piac. p. 536, tetiia leporit marmi Speciet, —
CuY. Ann. du Mu9. xn. pp. 359 — 170, PL 44, Mim. rar lea MoU. vn.,
Blaiky. MalaeU. PI. 46 bis, fig. 9 ; in the Mediterranean ; the Tdh, fmr
hria L., Bohadboh Dt qanhvad. Ainin. marin., Tab. v. figs, i, 7, appean
not to dif^ firom it. Dbllb Chiajx adopts two species, as does Linn^ous,
Memorie, m. p. 138, but his Teth. fimbria is not that of Ldtnaus, which is
distinguished by the absence of filaments at the margin of the vdwn.
Triopa JoHNST. (species of Trttonia Cuv., species of Euploca^
mu8 Phil.).
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784 CLAS xm.
Spi Tiiapm dmn^BT^ J>wm i imi^i m If wm , ZmL Ahl Ta^ 17. fiai. e-^
.Ami. < JKk. JImC. e. f^ 1x4^ IIS ^X"^' Ai^ XT
F^^mtcpitfias Leuck. PEaesMoom* CuT. from trpoei^ cmr .
Malia LeUi' k_ E^tp^rjmmiu PhiL-
C»ip. T.irmgy Br^aa A^imnl. fmrwmd. ZkaenfL ^ xf^ Tak. x.ff. l
ITur. IT, c^.«a^ pp. 98 — 90^ PL m. i^L u x.
/Acru L. CcT. Bj-It fiat or gibb«:'U3 &b^T^, wirh &bd>?tiKB £aL
cn-T^er^ hj & l<oe membrane, acd pILcat*>iiijr^iiiafie. Tent p«:«-
terl' T d- r«L in tie miJ line c^f bjilj. sanvjozided bj bnocoed .r
plinnefl bnnchix <iL«p:t?ed in & circle. Apertareg of gaiaxtijn 11
the rijh: «:ie- Tentacles £• or. two in^ri-T Tesdz^es of toI li
the pr V M'ii oa m:atn. tv j *nperi«>r letzactile within tiLbeSw in^i^-
Ute wirh tran^rer^e lame''-
m ^...li. JTml IT.,. 44; «iA fid.
Darit : wot tii« crrdi)3e «f Cutub
CmL, Ur^. CunL ttti p^ <rj — 52X, Tib. 3^ 17, awl G. Jumsatos
5i, Hajtoocx ana Emkcos «• Ac ^a^Mif «^ Avu^ MA. TV«m. iSfL
pp. JOX--jfi.
Spu DnriM Afy% L^ Bnganw 4^ \n\\mwi Amim^. mm. IUl t. %l 4, £>
DiUA Chultk Jfflii^ Ikr. jS, igiw i — % %atL, fap wd abore, j'^ Iok^.
«' fariMd: m the Mafits.; » fct ^wm of imHii ■» «ms m dbe
Kovtk Sa: />prir twitmi^m CcT^ Jossar., Alh. aid Hasbl Mmtogr,
Faa. L Fl. 3; i>!>rw jfaf^to Gmil., Bcmddi r<r*. «m i^ Zia—fci <^
■irtMl. m. p^ 51S, %. 4 idocB k fifcr frvB /Wit Ihmi^h 0»t., i>.
pl7iW JogCTT.r^
AoCeL — ^Add Mme ^Ruexm of modern wmen^ rSUerM lyOBHBST,
OMdUdkmi Blaist^ dmiodariM F<mdb.
Sectioo EL Organs of icspiiatkA biandiiae, inrfaiM in mantk,
or ooTcied br free margin of mantle.
Fanulj Y. ffypobrameiiaia, Naked gasteropoda, henna|^iio-
dite, marine. Branchial lamellje sitoated on eadi side in a bnow
between die mantk and foot, tnnsrene, in a single row.
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MOLLUSCA. 786
The heart lies in these mollusos in the middle of the bodj on the
dorsal surfiEtoe, and receives the blood from the gills placed on each
side. Milne Edwabds unites this fiunily with the preceding and
the next following into a common order under the name of Opia-
thobrcmchia. This family approaches, however, very near to the
preceding, and especially to the genera Doris and Tritania.
Phyllidia Cuv. Head with four tentacles, the two superior
retractile within a cavity. Anna in posterior and middle part of
back.
Comp. CnnXB Ann. du Mm. v. pp. 166—276, PL xvin. MoUmq., M4m.
"vm,
Sp. Phyllidia trilineata Cuv., Phyll, varicoia Lau., Cuv. 1. c. figs, i — 6,
Blainv. Malac. PI. 47, fig. z ; in the Indian and Red Beam, ms also PhyU
Udia putitUota Cuv. ibid. fig. 8, Lsuckabt in AUcu zu der Meite von
E. RuEPFSLL, Wvrbelloie Tkiert det roihen Meeres, p. 36, Tab. zi. fig. i.
Pleurophyllidia Meck., Diphyllidta Cuv., Armina Eafin. (on
the authority of Cantraine), Lingudla Blainv. Head with two
tentacles placed towards the back at the anterior margin of mantle.
Frontal veil with angle produced on each side. Vent in right side,
behind genital foramen. Mantle acuminate posteriorly.
Sp. Plewrophyllidia lineaia, DiphyU. linecOa Otto, Nov. Act, Acad. Ocu, Nat.
Cur. xvu.; Dblls Chiajb Memorie, i. p. ia8, Tab. 10, figs. 11 — so;
Mboksl Archiv /. d. Phytiol. vni. 1823, Taf. n. figs. 1—7, s. 190 — 207,
CAMTBAiins Malac mediterr. p. 63, PI. 2, fig. 4, in the Mediter. and aooord-
ing to Loy&r in the North Sea also. DiphyU. verruco$a Cantb. ibid. fig. 3.
Comp. ZHphyll. oceUata Dbsh., R. Ant., 4d. ill., Moll. PI. 31, fig. 2. (The
spedmen from the collection of Bbuohans, on which Cuvibb founded this
genns, now in the Rijkt Muttwn of Leyden, DiphyU. Brugmanni Cuv., is of
one colour, pale and with a smooth mantle.)
Family VI. Plearobrarichvxta, Marine gasteropods hermaphro-
dite, naked or furnished with a small internal shell, more rarely
external. Branchiae composed of divided laminae resembling leaves
pinnatifid or supradecomposed, on one side only, where are also the
vent and orifices of generation, mostly situated on the right side or
backwards on the back, more or less covered by the mantle.
This family, named by Cuvisr TeetihrcmcheBy which was trans-
lated by many writers FoTnatobranchia, by Menke Crypaibrcmchia,
has no greater claim to this name, or even less than many other
divisions of molluscs. We adopt therefore the name of PUwrchrcmr
chioUa instead of the sesquipedale of Blaik ville Monopleurobranchiata
VOL. I. 50
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786 CLABS XIII.
reoeived bj us in the first edition of this Handbook, n. U. 89. We
see that in this ire have been anticipated by €kuT and Rkkts.
Umbrella Lam. {Gagtrcplax BhAlSY.).
Heurobranchus CuT. Bodj with fiat abdomen, a fmrrow be-
tween the mantle and abdomen reoeiying the branchiae on tiie right
aide. Mantle covering the whole of back. Yelnm in front of
month acuminate on each side. Two tnbnlar tentacles fissnied by
an external {nnow. Vent behind branchia in right side Shell
dorsal, contained in mantle, homj or calcareous.
Compu OB this genos CumB Ann. du Jf«f. t. p. 366 and foIL PL Tm.
MoUm$q., Mim. vm.
Sp. Flemno^. Peronii CUY., 1. L 6g8. i, 7, JL Ami., id. Ql,, MalL FL 31,
fig. r; from the IndUn Sea; — Pleurobr. Fontalii LiUCK., Fobsk. Iftm.
ibr. natmral. Tab. iS, ^. a, Lepns 9unrinm$ Ruvrpbll, AUat L L TUi. 5..
fig. 3; in the Bed Sea;— PZoir. iahtdiMariia Cavtb., Plemr. Ponkali
Dkllb Chiajb, Memorie, PL 41, fig. 11; fixnn the Mediter. &c. Comp.
CABTMAnn Jfolof . nUd. pp. 87 — 90.
FleurobrancluBa Meck., Pleurdbranchidium Blaint. Four ten-
taculiform laciniae, not retractile. Yent aboTe branchia in the
right side. Shell none. Mantle shorter than bodj.
Spw PUmnlbfwikdkaM Medhdii Cur., Bbllb Chiajb Meautrie, FL 40^ fig. n.
Cabtbaibb L L PL m. ^. 3 ; from the Mediterr. Sea. Comp. & F.Liri
Dml de PiemrobramcktBa. Habe^ 1813, c. fig.
Aplysia L., GxEL. {Laplysia L., typogr. error, Lam.). Body
oblong, Umadform, mostlj margined by a broad Telnm reflected
over the back. Two contractile tentacles, conical, snlcated, in the
npper part of head ; two productions of the Telnm surrounding the
mouthy forming as it were a second pair of inferior tentacles. Eyes
sessile in firont of base of superior tentacles. Branchiae doisal,
covered by a production of mantle, with an operculum including a
flat membranoso-comeons or calcareous ahelL
Comp. Saitdbb Rabo HttL md, dm Ajdyna. Ptais, 1828, foL
Dolahdla Lail Operculum <^ branchiae towards the posterior
part of back, including a cakareous shell Body mostly tronoated
posteriorly with an orbicnlar declining area.
Sp. Ajiy&ia RrnmfkH Rabo, Dtdabdla Rumjpkn Cut., Buvpr. Awdt. Ran-
tekik, IUk X. fig. 5 (Tkb. XL. fig. B the shfiD); Cur. Am^. du Mv9, t. p-
437 and fcXL, PL 39, figs. 1—4, MoR., MHl 13, Bab« L 1. PL i.
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MOLLUSCA. 787
Aplysia a/actor. Body elongate, not truncated posteriorly, angus-
tate. Dorsal operculum of branchiie including a membranous homy
shell.
Comp. BoHADSOH De qmbu9d, animaUb. tMrima, pp. x — 53, Tkb. I. — nr.
CuYiSB Ann, du Mtu, n. pp. 287 — 314, PL ii. MoUuaq., M6m. iz.; Dsllx
Chiajb Memorie, i. pp. 15 — 76, Tab. n. — y.
These animalH, on a fint impresmon, resemble gigantic slugs {Limacea).
The ear-shaped tentacles have given occasion to stamp this genus with the
name of Seorhare (Lepua fMurimu). They live on /vci; and shed a purple
fluid at the margin of the mantle.
Sp. Apiyna depilana L., Bohadbch 1. 1 Tab. i.— iv. Rang L 1. PL 16,
GuviEB R. Am., id, HI,, Moll, PL 33 ; one of the largest species; of this
Mediterranean species, under the name of Lemcsa, Bohadbch gave L cit.
a detailed description and a masterly anatomical investigation. In that
and in other seas still different species of this genus are found. The eggs
are laid in long tortuous strings. The development of Aplysia has been
observed by Van BlirsDSN, Ann. des Sc, not, se S^rie, zv. pp. 133 — 12 8,
PL I.
Notarchu8 Cuv.
Bursatella Blainv.
Note. — ^Bang 1. L concludes that these genera form sections only of the
great genus Aplysia, Of Notarchtu the same has been noted by the cele-
brated voyagers QuOT and Gaiilabd, Voyage de VAUrolahe, Zoologie, u.
p. 31a.
Gastropteron Meck. Tentacles none. Margins of foot ex-
panded into ample wings. Shell none. Branchise pectinate,
almost entirely naked; a small duplicature of mantle, produced
beyond the body into a filiform lacinia.
Sp. Ckutropteron Meckdii J. F. J. KoBSSy Diss, de Pteropodum ordine et
novo ipdiM ffenere, HalsB, 181 3, 4to,fig8. 11 — 18, (Jlio Amati'DvLLE Chiaje,
Memorie, pp. 53 — 59, Tab. n. figs, i — 8; CAin'RAiNB L 1. PL 4, fig. 4; a
small mollusc from the Medit. Sea, of a rose-red colour ; the only species
hitherto known.
Acera Mueller (in part), Lam. {Daridium Meck., Lobaria
Blainv.). Tentacles none. Veil of head large, separated from
mantle by a transverse furrow. Vent, orifice of generation, bran-
chisB and heart placed at the right side; branchiae posterior. A
small shell included in the posterior part of mantle, flat, calcareous
at the incurved apex, elsewhere membranous, thin.
Comp. on this and the two following genera CuvnBB Ann, dv> Mue. xvi.
pp. I — 18, MoUueq., M4fn. x.
50—2
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788 CLASS xin.
Sp. Aeera eamoia CvY., Ami. du Mw, L 1. p. lo, PI. L figs. 15—19, MHk.
«. I. MoU, 1. L PL 3, figs. 15—19* J2. ^ni., id, m,, McU. PL 35, fig. 1. See
on another species from the Mediterranesiiy Cahtsaivr op. dt. pp. 73, 74-
BuUa L. (in part), Brug. Velum of head large, separated from
mantle hj a transverse furrow, in some produced Into four lacinis
or tentaculiform appendages. Vent, orifice of generation, bianchise
and heart placed at the right side. Foot shorter than body, with
lateral margin produced, undulato-plicate. Shell thin, convolute,
with aperture large.
BuUasa Lail {Philine Ascan., Lobciria Muell., Gxel., species of
Bidla L.). Shell very thin, concealod, not afiixed by muscles^ with
very wide aperture, and lip alone slightly involute.
Sp. BuUa aperia L., Qm. (m part), Lobaria quadrHoba Muxll., Gmkl.,
MusLUEB Zool. Danie, Tsb. 100, figs. 1—5, Guv. R, Ani,, id. UL, MoU.,
^ ZSf fig. I ; in the Mediterraneftn and North Sea.
BuUa Lail {Scaphander Montf., Altctda Ehbenb., dec) Shell
external affixed by muscles, involute, with spire little exsert or
depressed
Sp. Btdla amptdla L., Bumfh. Ami. RariteUk. Tab. 37, ^. q, BLAUrr.
Maiaeol. Fl. 45, fig. ii ; — the apex is somewhat projecting in JBmBa ApUMft
{BuOa AmpUuibre L.), Bncyd. mitk., Ven. PL 359, fig. 1, Blaihv. MoJm.
L L fig. 10, &c. There are also foeaU speciet of this gemu, almost sU
from the tertiary formationa.
Naie,'^Po8ierobraneh€Ba D'Ohmgnt. A genus related to the
AoercBy sheU none, but dififering from its oongenens by the branchis
placed on the left tide.
Sp. Potterobr. maevXata D'Obbiont, Voyage dam VAmSr. mirid. (reprinted in
Okin's Ins, 1839, pp. 516, 527; oomp. also Trobohil, Wivgk. Ar^v,
1838, n. p. 181).
The new genera Lobiger and Lophooenie Krohn, if indeed thej
belong to this fiunily, must as it seems be placed near Aplysia.
Gomp. KrokSj iw deva nowwaiut gmn$ de CkuUropcdet, Ann. det Se.
fMl. 30 S^e, yn. pp. 53--60, PI. n.
Family VH. Cydobranehiata. Grasteropods with branchi«
foliaceous or pyramidal, arranged in rows on each side under the
margin of mantle. Sexual organs without organs of copulation.
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MOLLUSCA. 789
Sexes distinct. Tongue long, lineax, mostly beset densely with
teeth and barbs. Shell resembling a shield, dorsal, not turbinated,
with aperture ample.
The position of the branohiBB brings this fiunily into the neigh-
bourhood of the ffypobranchiatiiy and thus Lamabck refers Fatdla
and Chiton with Phyllidia to the same family. In other respects,
however, there exists much difference amongst these animals, as at
once appears from the disposition of the sexual otgans, since in
this division the sexes are distinct Still, in a natural sequence of
the animal kingdom, the CyclobrcmchicUa ought to form as it were
the transition of the Ctenobrcmddata to the three preceding fiunilies^
the OpisthobranchicUa of Milne Edwabds.
That some indiyidualfl of Patdla are fiemale, othera male, was obsenred
by Gbat {Annals of Nat. Bitt. i. p. 48a), by Mnjrs Edwabds (Anndlei
det Se. not,, se S^e, xni. p. 376), by Pstkbs and BoBiir (Mukllxb's
Archiv, 1846, 8. 134) and by Waonbb (besides in Patella) ako in Chiton
{Annali of Nat. Hitt. VI. p. 70).
Chiton L. Shell multivalve, made up of (eight) testaceous
scales arranged in a longitudinal row, incumbent on back. Mantle
at the circumference not covered by shell, with margins hard,
coriaceous, often aculeate or squamose. Ventral disc elongate,
narrower than body. Eyes and tentacles none ; head crested by
a wavy veil.
With LiNK.£US there are three genera of Testacea muUivaZvia :
ChUony Lepaa and Phohs. The last genus belongs to the Conchifera
or BivcUvia ; Lepaa is, as we stated above, a family of the Crustaceck.
Thus there remaios the genus GhUon alone as a true multivalve
mollusc. That it has no affinity or true similarity with the Cirri-
pedia (Lepaa L.), to which Blainville united it under the name of
Malacoentoma, now requires no demonstration ; but many authors,
both of earlier and later periods, still maintain that it differs too
remarkably from the rest of the Gasteropods to allow it to remain
in the neighbourhood of Patella, and that it forms the transition to
the ringed worms (Milne Edwabds Ann, dea So, noLy 3e S^e, ix.
1848, p. 110). It appears to us that these writers attach too much
weight to tbe external resemblance of the pieces of shell to rings of
articulate ftnimiiliL In the internal structure there is, perhaps, with
the exception of the remarkable occurrence of two oviducts (or vaaa
deferentia), and two sexual apertures placed at the side, nothing to
be met with that can indicate a remote affinity with the ArticvJ<Ua,
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790 CLASS XIIX.
Gbmpare for the Anatomy of Ckiion, Pou Tuiae. utr. Sie. i. MuUwah.
pp. I — lo, Tab. ni.; Cuvieb MSm, pow tervir d VHui. nai. ei AT Anal, da
MoUuiq. No. i8, pp. 3i — 2S, PI. m. figs. 8—14, and A. Th. Middehvoeft
JBeKhreibung u. Anatomie n€uer CkUanen, Mhn. de TAcad. impfr, da Se.
de St. P^tenbowrg, 6e S^rie, Tom. vi. 1848, pp. 67—315, ay. 14 pL
The species of this genus are very numerous and difficult to distingiuih
otherwise than by accurate measurements of the comparative length and
breadth, the greater or lesser inequality &c. of the pieces of sheU, and bj
the nature of the maigins of the mantle which are not covered by the dor-
sal shields. In some species these margins of the mantle oover, as thoagfa
they came together by continued growth, the dorsal shields, so tiiat the
shell is concealed within them. Of these Middsiomrff forms the sob-
genus CryptochiUm (Sp. OryptochiUm Stdieri Meddekd. L L Taf. i. figs, i, 1,
CkUon amictdatui Sowerbt, Oonchol. lUtutratums, ChiUmes, fig. 80,
Reeve Conchologia tyttem, n. PI. 133, 133, fig. 80; from which Chitnii
aimieylal¥M Pall., according to Middend., does not differ). The remaining
species form the sub-genus PhanoehiUm Middend., in which the branchie
are sometimes situated backwards and the row of pieces of shell is inter-
rupted by the mantle {DUhackUan Middekd., ChUondtm Lail, Sp. ChiHh
ndlitt lasvU Lah., Blaiitv. Malae. PI. 87, fig. 5); or the pieces of sheU dose
upon one another and the brancluse are placed round about at the sides of the
mantle {HamachUon Middend.) To this last division belong most of the
species of the genus Ckiton of LamaBCK. On account of its slse we notice
ChiUm gigax Spenol., Ghemn. Cimckyl, Tab. 96, fig. 819, Encyd. mcth.,
Ven. PL i6t, fig. 3 (figured in the young state by F. Kbaubs SSdafrik,
MoUiuk. Tab. m. fig. 3).
Compare also on the species of the genus Chiton, Spekolxb Skrivter of
naturh. SdtkabH, IV. i, 1797, pp. 62—103, Tab. 6, and J. E. GvjLxAnwdi
of Nat, HUL XX. 1847, p. 131 and foil.
Patella L. (exclusive of several species). Shell univalve,
covering body entirely above, clypeate or retuso-<;onical. Branchia)
placed under the margin of mantle, lamellose, lateral, disposed in a
series surrounding the body. Head with two tentacles acuminate,
bearing the eyes externally at their base.
Limpets. A yery numerous genus of molluscs. The intestinal cual
which, as in the preceding genus, is veiy long and forms many convolutioDf
runs with the rectum upwards, so that the vent is placed under the head
on the right side, whilst in Ckiton it lies in the mid liue at the posterior
extremity of the body. The nervous system presents a band lying tran^
Tersely upon the oesophagus and swelling on each side into a ganglion,
irom which the nerves for the eyes, the feelers and the neighbouring parts
arise. Two nervous filaments descend on each side and form a transvene
ganglion, irom which nerves arise for the foot, the viscera and the drcalar
muscle that attaches the animal firmly to its shell. Compare Cuyibb M^n-
9ur let MoUuaq. No. 18, pp. 15—19, PI. n. figs. 8—19, and the fig. of
Patella algira DxflH. in Cuv. R. Ant., id. iU., MM. PL 66.
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MOLLUSCA. 791
ip. Patella tmlffota L., BLAimr. Malacol. PL 48, fig. i, PI. 49, fig. i ; PaieUa
granoLina L., LiSTBB Conchyl, Tab. 534, fig. 1$;— Patella wmpreua L.,
L18TBB, Tab. 541, fig. 35, Blainv. PI. 49, fig. a, &c.
Appendix to the CyclobranchicUa.
Nematobranchia Menke {Ctrrobranchiata BlAlNV.). (An ab-
noTinal, aberrant, family.)
Dentalium L. Shell tubular, elongato-conical, subarcuate, per-
vious at both extremities, with posterior aperture narrow, a fissure
often extending upon the shell. Animal conical, eyes none, bran-
chiae composed of many cirri arranged in two fasciculi.
The genus Dentalium, placed by Limr^us with PaUHla. and referred by
later writers to the ringed worms, belongs in reality, according to the
investigations of Sayignt and Dbshatbb, to the MoHueca, although it is
difficult to unite that genus with any of the other fiunilies. It forms a
small group between the Heteropoda and Aspidcbranchiata (Emarginvla),
The species are numerous ; many are extinct, and occur in various tertiary
and secondary formations.
Sp. Dentalium dephamitinum L., Mabtini Conchyl. Tab. i. fig. 4 a, Desh.
(see below) PL ivn. fig. .7 ;— Dctrf. ErUalie L., Debh. PI. xv. fig. 7, xvi.
fig. «, Gu^iN Iconogr., Annel. PI. 3, fig. i, Cuv. R, Ani., id, ill., Annel.
PI. 7, &c.
Comp. on this genus Debhatbs A natomie et Monographie du genre Den-
tate, Mim, de la Soc, d^ffiet, not, 11. Paris, 1825, pp. 321 — 378, PL 15—18.
Family VIII. AsptdohrancMata. Shell clypeiform, in most
not turbinate, with very laxge aperture, incumbent on the pectinate
branchiae. Heart with two auricles, perforated by the rectum as in
Acephula. Organs of sex distinct, some individuals male, others
female.
Lottia Gray {Patelloide QuOY and Gaim.). Branchia pectinate,
placed on the right side and forwards, often emerging beyond the
mantle. Shell conical, often thin.
Note. — A genus very similar to the Paielke, but different in the structure
of the branchiiB. Comp. QuoT et Gaihabd Voyage de rAetrolabe, Zoolo-
gie, m. 1834, pp. 349 — 366. Sp. Lottia tettudinaria, Patella testudinaria
Xi., Lax. ;— Patella virginea Mubll., LoUia pulcheUa Fobbes. Comp.
Aldbb Ann. of Nat. Hist, Ym. 1841, pp. 404 — 406.
Parmophonts Lam. Shell oblong, oval, depressed, clypeiform,
not turbinate, with margin entire, apex subincurved, not perforate.
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792 CLASS XIII.
Mantle with margin reflected, covering the circmnference of shell.
Tentacles thick, conical, bearing the eyes externally at the base.
Sp. Parmophorta auttralu BLAnrv., Malacol, PL 48, fig. 2, Ccvub JL Am.,
id, Ul., MoU. PL 65, &c.
Emarginula Lam. Shell scutellato-conical, anteriorly emaigi-
nate. Mantle famished with a single row of tentacles at the
margin of foot, incised anteriorly, under the emargioate part of
shell. Eyes set on tubercles at the base of tentacles.
Gomp. CuYiSB Mifa, awr lu MoUuaques, No. 18, pp. 14, 15, PL n. figs.
3-7.
Sp. Emarg, fitmura Lam., Paidlajumura L., Muxllsb Zool, Danic Thb. 14,
fig". 7—9-
Of^hifl genua, as of the preceding, fooail species are found. See La-
ICABOX Ann, du Mut, i. pp. 383, 384 ; vi. PL XLin. figs. 5, 6.
Fissurella Brug. Shell clypeiform, with vertex perforate.
Sides of foot margined by mantle, rugose, without tentacles.
Mantle with an oval aperture on the back, corresponding to the
perforation of the shell, leading to the branchial cavity. Anus
placed below the fissure of mantle. Eyes sessile, placed externally
at the base of tentacles.
GuTiEB ifoUutq. L L pp. 12 — 14, PL n. figs, i, 1.
Sp. FiuwreUa grcBoaf PateUa graca 1.., Blainv. Malaeol, PL 48, fig. 3, kc
This genus also is met with fossil ; see Laxabck Ann, du Mu». I. p. 312.
The fossil species are principally from the tertiary formationB, some firom
the Chalk-period.
Haliotis L. Shell depressed, auriform, with aperture very
ample, spire very short. Head with two very long tentacles, and
two oculiferous tubercles behind the tentacles. Branchial cavity
placed to the left, including two lamellose pectinate branchise.
Sub-genus StamcUia Helblikg, with addition of StomcUella Lajl
Shell imperforate, with spire prominent.
Sp. HaUotii itnpefforata Chxhn., Gh., J^ofnaHa phymUis Hxlbl., HSivs-
CHEN Hcsturfaneker, ZTiu. 1782, pp. 19, ao, Tab. n. fig. 18, Blahtt.
Malaeol. PL 49 biB, fig. 4.
Sub-genus : Haliotis Lah. Shell perforate by a row of foramina
near the left margin, pearly within.
Seoreara, Feelers pass through the holes of the shelL These
holes are closed up in the hinder part of the shell as the aninuJ
grows, and at the same time new ones are formed more forwards.
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MOLLUSCA. 793
In front, in the margin, an excision is seen in the direction of the
holes, which afterwards, during the growth, becomes changed into
a hola The ventral disc or foot is adorned with a double row of
feelers, tubercles, and fringes. The animal is attached to the shell
by a large oval muscle. The branchial cavity opens by a fissure of
the mantle on the right side, under the foremost holes of the shell.
Comp. CuviER Mollvaq, L L pp. 6—12, PL i. figs. 9 — 17.
Sp. ffaliotis parva L., ffal, eanaliculata Lam., Gu^in Iconogr., Moll, PL
^S> fig* I ; HaliotiM Mida L., Libtbb Conchyl. Tab. 613, fig. 5, Ac.
Many speciee of ffaHotis bear a great resemblance to each other, and are
difficult to disthigaish. Some fosail specimens of this genus are met with
in the tertiary formations.
Family IX. Aulobranchiata, Shell irregular, tubular, with
turns twisted spirally at the apex* Branchiae pectinate. Sexes
separate.
SiUqiuiria Brug. Shell thin, tubular, loosely spired ; aperture
rotundate, emarginate, with an incisure running with the spires to
a short distance from the apex. Mantle cloven in the same part ;
pectinate branchiae placed at the fissure of mantle.
Sp. SUiquaria anguina IjAM., SerptUa anguina L., Blaiky. Malacol., Prin-
cipet, PI. I. fig. II.
Magihis MoNTPORT.
Comp. £. RuEFPKLL USmoire swr U Ma^ui arUiquut Montf. ; Mimoiret
de la Soc. d^HUt, not, de Strcubaurg, Tom. i. Livr. a, 1833, ^^' ^g- Accord-
ing to BuEF. this animal belongs rather to the pectinibranchiate molluscs
than to this family ; it lives in the Red Sea inclosed in cavities of Mean-
drina,
Vermetm Adanson. Shell thin, tubular, loosely spired, with
spire adhering at the apex, elsewhere loose. Aperture orbicular,
with margins connected. Head with two tentacles, oculiferous exter-
nally at the base. Appendage of body replicate downwards, mostly
ftimished with an operculum. Branchiae arranged in a conical
series, at the left side of respiratory cavity.
Sp. Vermettit lumhricdlu, Serpula Iwmbricalts L., Blautt, Mtdacol. PI.
ZXUY. fig. I, Ac.
Family X. Ctenobranchiata. Shell spirate or conical. Bran-
chial cavity contained in the last turn of shell, open externally by
a large fissure between body and margin of mantle, containing
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794 CLASS XIII.
three branchiae, two or a single one, composed of nnmeioiis leaves,
arranged parallel like the teeth of a comb.
Very rarely in place of branchiae a vascnlar network in the walk
of respiratory cavity. Sexes separate, external organs of copula-
tion distinct.
Fectinibranchiaie$» In all two feelers and two eyes are present ;
these last are often pediculate. The mouth has the form of a pro-
boscis, of which the structure has been described at length by
CuviEB in the whelk {Buccinum). It consists of two tubes, which
push one into the other and are connected ^together by the upper
margin, so that when the innermost tube is tmroUed and elongated,
the outermost becomes shorter. This instrument is moved by many
muscles^. The hinder part of the body contains the liver and the
sexual organs. The sexes are distinct. In the male the penis is
situated on the right side, behind the head, and in some, as in the
whelk, is very larga It is folded round and concealed in the re^i-
ratory cavity, but is not retracted within the body, except in the
genus PcUtulinaf where it is protruded and retracted through an
aperture in the right tentacle, which had been observed already by
Lister, but was afterwards incorrectly denied by Drapaskauix
Through the penis runs a tortuous canal, which on copulation,
when that organ is erected, probably loses its tortuosities. Ac-
cording to Basteb and Blaikville, the shells indicate a difference
of sex, and those of the female are wider, particularly in the last
wreath*.
The females secrete a kind of common envelope for the eg^
which they deposit at the same time with the eggs. On our shores
roimd diunps of yellow vesicles may be frequently observed, which
resemble bunches of grapes, and are the masses of whelk's egg^ in
question*. According to Cuvibr, this envelope is secreted by a
1 Ann, du Mua, xi. 1808, MSmoiret t, I, Moll, No. 17, pp. 6, 7, figs. 8—10.
> See Basteb Natuurk, UiUp. i. bl. 39, 40; Blainv. Journal de Physique, xciv.
p. 91; Mbokbl's Arehiv /. d, Phytiol. vn. s. 571—573, 1822. Bastxr sayi th»t in
Buecinum the Bhell of the male ia somewhat smaller^ that it has a greater number of
wreaths, but which are thinner than thoee of the female.
* See figares of them in Basteb NiXtwirh, UiUp, i. Tab. V. figs. 2, 3 of Svceinum
undatum; Tab. vi. figs, i — 3, of a species of Murex, Gomp. ibid. bl. 38 — 45; mc
also Lund RecKerchet 9wr lea Envdopptt d^mufi dea GastSropodes peetinibranchet, Awt,
det Sc. nat, ae S^rie, i. Zoologie, pp. 84—111. By Abistoteles these masses of egga
are called /ueXiKtjf/Ku (translated /avo^mo); he did not, however, suppose that shdled
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MOLLUSCA. 795
lamellate or cellular organ situated in the respiratory cavity near
the gills, and of which the interspaces are filled by a slimy fluid.
This organ, however, occurs in the male, although less developed*.
Phalanx I. Capulmdea, Shell conical or orbicular, scarcely
turbinate. Aperture very large, without any sinus or canal.
Operculum in some none, in others subintemal.
Stffaretus Adans., Cryptostoma Blainv. Shell orbicular or
oblong, auriform, concealed imder the foot reflected towards the
back. Aperture ample, entire, with labium effuse, dilate, patent.
Cavity of branchisB furnished with a semicanal, in connexion with
an incisure of mantle, and containing two pectinate branchiae.
Head with two tentacles depressed, conical, furnished with a tuber-
cle at the base. (Eyes none ?)
Sp. Sigaretus haliotoideus JjAM., Hdiat haiiotoidea L., Ruhph. Afnh. Rari-
teitk. Tab. 40, fig. B, Adanbon CoquUl. PI. i, fig. 2, Ac. This genufl la
allied to Natica in the following division.
Coriocella Blainv. {Sigaretus Cuv., LameHaria Montagu,
LovjfiN. Shell with aperture ample, auriform, thin, homy, con-
cealed under mantle. Two eyes at the base of tentacles. Tongue
armed with barbs, very long, spirally convolute.
CJomp. Cuv. M6m, ». Z. MoUtiaqs No. 18, pp. a— 6, PL i. figs. 1—8.
Sp. Sigaretus Tonganua QuoT and Gaim., Attrolahe, Zool, n. p. 317, Cuv.
R, Ani., id. ill., MoU. PI. 49, fig. 2, &c.
Siphona/ria Sow.
Is this its place ? Comp. Blaikv. Diet, det 8c. not. Tom. 3a, p. ^67,
Malaecl. p. 475, Rebvb Conekol. syst. ii. pp. 19, 20, Michblin in Gu6iiN
Magaa. de Zool. 1831, 1831.
Calyptrcea Lam. (and Crepidula ejusd.). Shell with aperture
ample, orbicular or oblong, convex on the back. Branchial series
single, of filaments often very long. Tentacles triangular ; eyes at
the base of tentacles towards the outside.
CcUj/ptrcBa Lam. Shell conoid with base orbicular. A calcai*eous
lamina, conical or spiral, in the cavity of shell, and adhering to its
apex.
^yiiTTntlg proceeded from them, bat imagined, according to the notions then prevalent,
that these originated from slime and putrefaction. De Hist. Animal, v. c. 15 initio.
^ CuviEB Mim. 8vr lea MoUusq. 1. 1. p. $, and fig. 3/.
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796 CLASS XIII.
Sp. Calyptrixa equetiris Lam., PateUa equetlrit L., Buxfh. Amh, RarUaiL
Tab. 40, figs. F, Q; D'ABOBinr. PI. a, fig. k, Bladty. Maiaad. PL 49 fai%
fig. 1 ; habitat Indian Ocean. Comp. on this genus Dbbhatib Mimoin
awr la Calypirie, Ann. det 8e. naL m. 1824, pp. 535 — 344; BwmBir
DeaeripHom of tome new tpec. of Calyptreeidea, TnauaeL <iftke ZooL SocL
St 1835, pp.195— ao6, PI. 17— «9; OWBF On the Anatomy tfCai^ftraidte,
ibid. pp. 107 — an, PI. 30.
Sub-genera: Calypeopns Lbss., Zoo{., Coq. p. 399, lUuttr, de Zool. PL « ;
.6dAe(2af)Aitf OwXN (with calcareous lamina, affixed by its base). — Comp.
Rbevb Conch. 9yH, u. pp. 31, 3a.
CrepidtUa Lail Shell oyate or oblong, with apex mcaired, in-
clined towards margin. Aperture partly closed hy a horizontal
lamina.
Comp. CuTiSB MoUuaq., M4m. 18, pp. 10, a i, PI. m. figs. 3 — 7.
Sp. Orepidula poreellana Lax., PaUUa poredlcma L., Rumfh. Amb. Rari-
tekk. Tab. 40, fig. o; Blaihv. Mdlacol. PL 49 bis, fig. 3, kc.
Sub-genus Cfrq^ipatdia Lsss., ZooL, Coq., III. de Zool. PL 4a.
Capulus MoNTFORT, Pileopsis Lam. Shell obliquely conical,
with apex incurved backwards. Animal with long proboscis,
deeply chanelled above. Eyes set on tubercles or petioles at the
outside of tentacles. Neck furnished beneath with a folded velum.
Branchial series single, composed of narrow laminae.
Comp. CuYiiB 1. 1. pp. 19, ao, PL in. figs, i, a.
Sp. Capvlua hungartcui, Patella unga/rica L., D'Abgsht. PI. 4, fig. 3, Cur.
R. Ani., id, ill, MM. PI. 47, ^. 7, ko.
Hipponyx Defb. With lamina basal, calcareous.
Sp. Oa^pvUki cornucopia, PUeopHi comu eopioB Lam., Blaikt. MtUac. PL b^
fig. I, Bbonn Ldh. geogn. Tab. 40, fig. la; fossil in the ealcaire grottitr.
This sub-genus is related to Capulue as Lithedaphue is to CalyptrcM. Some
species still living are known, and some others fosnl, all from the tertiary
formations.
Phalanx II. Trochotdea. Shell spiral, with aperture entire,
not canaliferous ; animal mostly furnished with an operculum.
PileollAS COOKSON, SOWERBY.
Comp. Deshatxs Sneyd. nUth., Vert. m. p. 764. Fossil genus.
Navicella Lam., Septaria Feruss. Shell elliptical, or oblong,
convex above, concave below, with apex subspiral inflected as far as
the margin. Labium flattened, narrow, transverse. Operculiun
concealed, flat, irregular, at the upper part of foot. Animal with
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J
MOLLUSCA. 797
long tentacles not retractile, and eyes petiolate at the outside of
tentacles.
Sp. NaviceUa elUpHca Lam. {Patdla nerUoHdea L. X), Meubohen NcBtur-
foncher, xii. Tib. 5, fig. i, Encyd, mHh,, Vers, PL 456, fig. i, BLAimr.
Malcbcol, PL 36 bis, fig. i, PL 48, fig. 5, &c. ; fresh-water molluscs from
- the tropical regions.
Nerita L. (in part). Shell spiral, planato-ovate or semiglobose,
imperforate. Aperture semiorbicular, with columellar margin or
lahium dilate, transverse, truncate. Foot of animal broad, furnished
with operculum; eyes set on petioles or tubercles at the base of
tentacles, towards the outside.
* Flnyiatile species, with shell mostly thinner and lip edentulous. NeriHna
-Lam,
Sp. NerUaJlwviatilU L., Swajocsbdam Sibl. not. Tab. X. ^. «, PvEirrsB
Land- und Wasaendin, i. Tab. 4, figs. 57 — 39, &c.
* * Marine species, with shell thicker and lip often dentate. NerUa Lam.
Sp. NerUa Pdoronta L., Encyd. mith,, Vers, PL 454, fig. 9, Blainv. Malac,
PL 36 bis, fig. 6, &c. Species many, some fossil.
Narica Recluz (species of Sigaretua Lam.), Merry a Gray.
Shell subglobose, umbilicate, with short spire. Aperture ample,
semiorbicular. Operculum homy. Head proboscideous ; two tri-
angular tentacles bearing very small eyes externally at the base.
Single branchia, with triangular leaflets, contained in a large
respiratory cavity.
Sp. Nor, eanceUata Bxoluz, Sigaretui eanceUaiiu Lam., Bsoluz, Gu&iy
Magoi. de Zool. 1845, MoU, PL 119. Comp. on this and other species of
this genus, all of which live in the sea^ BaoLUZ op. dt.
Velutina Gray, Blainv.
• Sp. Vdutvna haliot&Sdea, BuUa vdutma MuxLL. Zod. Ikmic. Tab. loi, figs.
I — 4. ^o this species O. F. Mubllbb refers ffeUx halwMdea, which we
noted above at StgarOus,)
Nattca Lam. {NeritcB vmbilicaUB L.). Shell subglobose, umbi-
licate, with spire short and last wreath tumid, large. Aperture
semiorbicular, with lip oblique, edentulous, callous. Head of
animal with broad velum ; tentacles two ; eyes small, sessile at the
base of tentacles. Foot very ample.
Sp. NcAica mammilla Lam., NerUa mammilla L., Rumph. Amh, RariteUk,
Tab. i«, fig. P, Blainv. Malacd. PL 36 bis, fig. 5, &c. Species numerous.
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798 CLASS XIII,
Note, — G«nus Deshayeaia Raulin {Ouisis Magaa. 1844, M<^
111), fosaily intermediate between Neriia and N(U\ca^ still appean to
be more nearly related to the NatwuB,
Janthina Lam. Shell Tentncoae, thin, pellucid. Aperture
triangular. Columella straight, produced beyond the margin of
aperture. Animal with proboscis large, cylindrical, and two tenta-
cles deeply cloven. A yesicular or spumose organ adhering to
foot. Operculum none.
Comp. on thiB genuB, CuviSB ^nn. du Mu$. xi. pp. X3i — 130, MoB^t
Mhn. 15, pp. 1 — 10, figs. I — 8.
Sp. Janthina communis, ffdix Janthina L., RuKFH. Amb. Ra/riUUk, TA.
XX. fig. 1, Blajnv. Malacol. PI. 37 bis, fig. i ; in the Mediter. Sea and in
the Atl. Ocean, in wann climates. The foamy organ at the foot seireB for
attaching the covering of the eggs, and, according to "Rajxq, is afterwarde
cast off. Is this oi^gan peculiar to the females?
Scalarta Lam. Shell turrite, with ribs longitudinal, compressed,
interrupted. Aperture rotundate, with margins connected, margi-
nate, reflected. Animal with long retractile proboscis ; eyes sessile
at the base of tentacles. Operculum homy.
Sp. Scalaria pretio§a Lax., Twrbo $ealari» L., Ritkph. Amb, RarkeilL Tab.
49, fig. A, Gu^iN Iconogr,, Mod. PI. i a, fig. 11 ; the true winding ttair,
from the Indian Ocean ; — Seal, communis Lam., Turho claihrut L., Blact^-
Malae, PL 34, fig. a; in the AtL Ocean, the North 8ea» the Hediter.
There are ▼arions fossil species from the tertiary formations. Comp.
Lamabok Ann. du Mut. v. pp. 113—214, viii. PL 37, figs. 3—5.
Pyramidella Lam. Shell turrite. Aperture entire, scmioval,
with labrum acute. Columella with three transverse folds. Oper-
culum homy. Tentacles broad, long, sulcate ; eyes sessile at the
base of tentacles.
Sp. Pyramid, dolabrata Lam., Trochus dolahraiut L., Blaint. Malac. PL
II, fig. 4, Cuv. R. Ani., id. ill., Moll. PL 45, fig. 3 ;—jPyr. roi^rKWW
Gu^BiN, Magaa. de Zool. 1831, MoU. PL a, kc. (Species marine, exotic.)
Tomatella Lam. (in part). Shell convolute, oyato-cylindrical,
frequently transversely striated. Aperture entire, oblong, with
external margin acute. Columella plicate. Operculum homy.
Sp. TomaieUa fatciata Lam., Voluta tomoHlis L., Encyd, m&h.. Vert, FL
ASh fig- 3> Blainv. Malae. PL 38, fig. 5, Ac.
Trochtis L. Shell turbinate, with aperture sinuoso-quadrangu-
lar, depressed, obliquely truncated. Animal with mantle often
appendiculate on both sides.
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MOLLUSa^. 799
Solm/nwm Lail Shell orbicular, oonioo-depressed, umbilicate,
with umbilicus patulous, crenulate or dentate at the internal margins
of Tfrreaths. Columella nona Operculum homy.
Sp. Trochus perspeeUtms L., Solarium perspectivum. Lam., Libtbb Conch,
Tab. 636, fig. 24, BuMFH. Anib. JRariieitk, Tab. 37, fig. l, &c.
Euonvphalua Sow. (fossil genus).
BifironUa Desh., Schizoaioma Bbonn.
Rotella Lam. Shell orbicular^ shining, imperforate. Spire Very-
short, subcorneal Columella callous, convex.
Sp. Trochua vetHaHus L., Rotella UneokUa, &c.
Trochtis Lam. Shell conical, in some mnbilicate, in others
imperforate. Aperture transversely depressed, with margins dis-
joined above. Columella arcuate, somewhat prominent at the base.
Operculum mostly homy, more seldom calcareous.
Topt, Oomp. on the animal of this genus Cuvieb Ann. du Mus, XI.
pp. 184, 185, Moll, M6m. 16, pp. 15, 16, fig. 13.
Sp. Trochus solaria L., Chehn. Conchyl. Tab. 173, figs. 1700, 1701, Beeyb
Conch. Syal, n. PI. 114, figs, i, 1 ; — Troch. niloticua L., Bumfh. Amb,
Jiariieitk. Tab. 2j, fig. A, JEncyd. mith., Vera. PI. 444, fig. i, &c. Trochua
Cookii Chemk., Lam., Lesson, IU. de Zool. PI. 15, from which Lessok
forms the genus Cookia, belongs, according to Bebhates, to Jhirbo.
Trochua conchyliophorua Bobn., Gm., Cuv. R. Ani, €d. ill., Moll. PI. 41,
fig. 3. This species causes little stones and pieces of bivalves from the
bottom on which it creeps to adhere to its shell; these foreign bodies in
time grow to the shell which thus acquires a very irregular appearance.
Lamabck confounded with this species, from the West Indian Seas, a
fossil species from the tertiary formations, which presents the same pecu-
liarity, under the name of Trochus agglutinana. Gomp. on the fossil species
Lamaeoe Ann. du Mua. Vf. pp. 46—51, vii. PI. xv. figs. 5 — 7.
Monodonta Lam., Labia Oken.
Delphinula Lam. Shell umbilicate, subdiscoidal or conical,
with wreaths rough or angulate. Aperture entire, rotund, some-
times trigonal, with convex margins. Operculum homy or calca-
reous.
Sp. Ddphimda laciniata, Twrbo Ddphinua It., BuMFH. Amb. Rariteitk. Tab.
aq, fig. H, BLAnrvriLLB McUaeol, PI. 33, fig. 3, fto. This genus is scarcely
distinguishable from the preceding, but has a circular aperture separate
from the columella.
Turbo L. (in part), Lam. Shell conoid or subturrite, with wreaths
rotundate. Aperture entire, rotund, not deformed by the penultimate
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800 - cMss xiiT,
turn, with margins disjoined above. Colmnella arcoate, flat-
tened, not truncated at the base. Animal with slender acuminate
tentacles and pedunculate eyes at the outside of tentacles. Opercu-
lum mostly thick, calcareous.
The species of this genus are very numerous. Those which sluae nacreoni
and silvery within, with a mostly very thick operculum, are more espe-
cially named Tvf^, as Turbo chrysottomui L., Rukph. Amh. Rariiatk,
Tab. 19, fig. B, Turbo a/rgyrottomm L., Ohsmk. Oonchyl. Tab. 177, figs.
1758, 1759, ^^^- ^' '^**-' ^' *^'> ^^' PI- 4«. fig- If &c-
The species which have no nacreous splendour internally, and which
have constantly a homy operculum, are united imder the following sab-
genus.
LUtorina FiBUS&
Sp. Twrbo liUoreut L., Alikruih, Vigneau, OwgnOU; Swaxmkbd. BM. not.
I. pp. 180—185, Tib. IX. fig. 14, Babtib Natuur, UUtp, i. pp. 127, 128,
Tab. 14, ^, I ; this little snail, boiled with salt, is eaten in Zealand.
Phastanella Lam. Shell ovate or conical, solid, imperforate.
Columella smooth, rotundate. Aperture ovate, longitudinal. Animal
with two laciniate labia. Two long conical tentacles ; eyes seated
in tubercles at the base of tentacles. Body margined by a laciniate
membrane.
Comp. CuYDEB Ann, du Mm, xi. pp. 130—135, Mim^ 9, L Moll. No. 15,
pp. 10—15, figs. 9—13.
Sp. PkatianeUa hulimoidea Lam., Buccinum aiutrdU Gicbl., BLAnrv. Moda-
col, PI. 37, fig. 5, from the S. Pacific, at New ZeaUnd and New HoUand
Some species of this genus of Lamabok an referred by Dbshatbs to genus
Littorina, There are some fossil species of PheuianeUa from the tertiaiy
formations; Lamabok Ann. dn Mus, IT. pp. 195 — 397, vm. PI. 60, fig. i.
Turritella Lam. Shell elongato-conical, acuminate, turbinate.
Aperture circular, entire, with margins disjoined above, labnun
emarginate by a sinus.
Sp. Turritdla hiemsfulata Lam., Blainv. Mabtcol. PL a r, fig. 3; TiarriteUa
dupliccOa Lam., Turbo duplicahtt L., Rseye Conch, SyH, n. PL 234, fig. r,
ko. To this genus belong nmnerous fossil spedes.
Paludina Lam. Shell turbinate, with rotundate wreaths.
Aperture subrotund, somewhat oblong, angulate above, with mar-
gins connected, acute. Operculum homy, orbicular. Animal with
tentacles acuminate, eyes sessile at the base of tentacles, mantle
alate, appendiculate.
Sp. Pcdudina vivipara Lam., Hdix vitnpara L., Swammerdam Bibl. not. i.
pp. 169—180, Tab. IX. figs. 15—18; Cuv. Ann. du Mm, XT. p. 170, MoR,
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MOLLUSCA. 801
Mim. 15; Pfeiffeb Land- u. Wanerachn, i. Tab. iv. figs. 4«, 43; Stubm
DetUschl. Faun. vi. 2 ; Blainv. MalacoL. PI. 36, fig. 4 ; the shell is a duU-
green, with red-brown bands and very convex wreaths. The right tentacle
is perforated in the male individuals with an aperture which gives passage
to the penis. The gills consist of three series of filaments. This snul, com-
mon in fresh water, is viviparous, and the females are full of young in
spring. It is found in Holland in canals and other fresh water, in the
mud or hiding under stones, also creeping under water-plants, &c. [Con-
sult especially, in addition to the works cited above, LsTDio Ud>er Palu-
dina vivipara, ein BcUrag sur ndkem KenrUnist dietes Thiers in emJbryo-
logitcher, anatomischer und Kitklogueher jBeziehunff, in Sikbold u. Koel-
likeb'b Zeittch, /. vnssenck Zool. ii. 1850, pp. H5— 197, PI. xi. xn.
xni.]
Valvata Muell. Shell discoidal or conical, with cylindrical
wreaths. Aperture rotundate. Animal furnished with homy orbi-
cular operculum, with foot anteriorly truncate or emarginate, with
two slender tentacles, and with eyes sessile at the base of tentacles.
Sp. FaZrato pwCTfwiMF^usa., Lam., Cyclottoma obtusum Dbap., Pfkiffeb
Land- u. Wiutertchn, i. Tab. iv. fig. 33, Stubm Deutschl, Fauna, vi. Heft
4, Tab. a, Blainv. MalacoL PI. 34, fig. 4, in fresh water, canals, &c.
Cyclostoma Lam., Draparx. Shell conical, short, with cylin-
drical wreaths, the last ample, tumid. Aperture regular, rotund,
with margins connected orbiculately, reflected by age. Animal
with a thin orbicular operculum, two tentacles, and petiolate eyes
at the base of tentacles.
Sp. Cyclottoma elegant Drapabn., Nerita eUgant Muell., Blainv. Malacol.
PL 34, fig. 7, Pfbiffbb Land- u. Wattertchn. I. Tab. iv. figs. 30, 31,
Gu^iN Iconogr., MoU, PI. ii, fig. 12, &c. This genus also counts many
fossil species from the tertiary formations.
These animals differ from the rest of this family, inasmuch as they do
not live in water, but in moist places, and breathe atmospheric air. Also
they have a vascular net in place of gills, and on this account might
belong to the following family, if the whole form of the body and the
internal structure did not remove them from the Pneumonica and place
them in the neighbourhood of Turbo.
Helictna Lam.
Ampullaria Lam. Shell globose, ventricose, umbilicate, with
spire short, obtuse, sometimes discoidal. Aperture entire, oblong.
Animal furnished with operculum, with long tentacles, and petiolate
eyes at the base of tentacles. Respiratory cavity large, containing
a pectinate branchia composed of a row of lamellee, and a vascular
pulmonary cavity.
VOL. 1. 51
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802 CLASS XIII.
Sp. A iiyiJanq /wcMrta Lax., JTeliv ampuUaeea L., Bukph. Amh. BarUatt.
Tkb. 47, fig. ^ £nc!fcl. mStk., Van, PL 457, figs. 3, a, B> Ac. Fresh-water
molhiBcs of tropicil ooontricv, much Allied to Paludina and Cydottoma
See F. H. Tho6CHXL Anaiomie vom AmpuUaria wreeui vmd ueber die Gai-
tmmg LaniMtM MoSTT. in Ebzchboh's Arckiv /. NiUnrgekk. 1845, s. 197 —
916, Tkf. 8. To AmpviUana belongs also Planorbu Oomu Arietis Lam.
Planaxis Lam.
Sp. PtamaxU nUcaUt Lam., Blaiky. Malaeol. PL 16, fig. 14.
lii$9oa Fremixv. Shell mostly turrite, sometimes globose,
with oval aperture, labram incrassate, exsert anteriorly and sub-
arcnate near the columella, the apex acuminate. Opercnlnni
homy. Tentacles elongate. Eyes placed at the base of tenta-
cles. Foot oblong, anteriorly truncated, posteriorly acuminate.
TbJB genus ooosists of small marine animals, whose little shell has some
resemblance to Tmrbo or Scalaria, and of which the numerous species have
become known in the last few years alone; there are also many fossil
species, almost all from the tertiaiy formations. Compare Dbshatu in
Lamakck ffiat. mat. da Anu s. veri. w ^ Tin. pp. 461 — 485 ; J. Aldkb
Anm, of NaL Mut. XUL pp. 323 — 318; Lov^sr Ind, MoUuteor. dfvenigi,
1846, i^ 94 — 36.
Mehnia Lam. Shell torrite, with apex often decorticated,
eroded. Aperture entire, OYate or oblong, efinse at the base.
Columella smooth. Operculum homy, elongate. Animal elon-
gate, with foot small, and two elongate filiform tentacles.
JWle. — ^Gienus Pirena Lax., Pyrena Meztke, Gray is, according
to Dbshates, to be abolished Smooth and shining marine species
of Mdania form the subgenus E%d%ma Risso. The rest of the
species, aU fluviatile, form the sub-genera Mdania, and Mdanopsis
FEBU8&, LaJL
Sp. Mdania awutnda Lax., ffdix awutrula L., RuvPH. Amb. RariteUk. Tah.
33, fig. P, r; Blahty. Malaeol. PL 35, fig. 7;— ifefcmia RangU DssH.,
Mdam, tuberculota Rang, Gu^bik Mag. de ZooL 1832, McU. PI. 13 (a fig.
with the animal) ftc The species are especially numerous in the riverB and
fireeh waters of the Sanda Islands ; comp. Mousaoir MoUuthen von Java,
1849, "• ^5 — 79* ^^ European species, Melan. helvetica Michkldt, GviiB.
Mag. de Zoei. 1831, MoU. PL 37, forms the tgpue of the gen. Pgrguia Dl
Chbibtof. and Jan. ; Dsshates thinks this species ought not to be sepa-
rated from Metania; others refisr it to Paludina.
{Mdanopns. Columella truncated, disjoined firom labrum by a
sinus.)
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M0LLU8CA. 803
Sp. Meiania buceinoidea Oliv., JBuceinnm prceromim L., Olit., Voyoffe, PI.
17, fig. 8, RiKYX Conehyl, 8yd, n. Tab. 195, fig. t, kc, Comp. on the
species of this lub-genua F&ubbac Monogr, det etp^cci vivantea et fossilet
du ffenre MSlanopnde, MSm. de la Soc. d^ffitt. not de Paris, i. 1823, pp.
13a— 164, PL 7, 8.
Phalanx HI. Cerycoidea {Buceinoidea Cuv.). Cochlea spiral,
with aperture emarginate or canaliculate towards the columella, a
canal receiving the tube of mantle.
Cerithium Adans., Brug., Lam. Shell turrite. Aperture
oval, oblique, terminated by a short canal truncated or recurved.
Animal with two tentacles acuminate, furnished with an oculiferous
tubercle towards the base.
«6p. (krilAitm patuttre Lam., StvomJtms paluttrU L., Rukph. Amb. BofrUeUk,
Tkb. 30, fig. q; Blaikv. Malaeol, PL to, fig. 4; — Oeriih. tdtteopivm,
Troehu tdacopium L., Rumph. Amh, RarUeiik, Tab. a i , fig. 11, D'AEOmrv.
Conehyl. Tab. 1 1, fig. a, Blainv. Malae. PL 32 bis, fig. 1, &c. A very
numerous genus of which the species live in great part in the sea, partly
in brackish water or at the mouths of riyers. The fossil species which are
still more numerous (more than 300 according to Dbshatbs), are ahnoet
an limited to the tertiary formations. Amongst these Cerithium giganteum
deserves to be noticed, a species found in France near Orignon.
The sub-genus Potamides (Ann, du ifttf. xv. pp. 367 — 370) is usually
not aooepted by later writers.
FoBtigieUa Resve.
Sp. PatHg, carinata Rbeyb, Ann. of Nat. ffitt., second series, n. 1848, p. 66.
(MabUuB of Turritella, characters almost of CeriiMum,)
Pleurotoma Lam. Aperture produced into an elongated straight
canal. Shell turrite or fusiform. Columella smooth. Lip cloven
or emarginate bj a sinus. Two conical tentacles ; ejes at the base
of tentacles extemallj. Operculum homj.
Sp. Pleurotoma habylonia Lam., Mwrex hdbylonius L., RiTHFH. Am^. Rari-
teUJs. Tab. ^9, fig. L, Blainv. Malaeol. PI. 15, fig. 3, Cuv. R Ani., id. iU.,
MoU. PI. 60, fig. 1, &c. A numerous genus, of which also several fossil
species are known.
Fu8U8 Brug. (in part), Lam. Shell destitute of spines ; aper-
ture produced into a long straight canal. Columella smooth.
Labrum entire. Operculum homy.
Fu9U8 Lax. Shell fusiform, spire elongate, ventricoae in the
middla
51—2
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804 CLASS XIII.
Bp. Fuiut syraeutantu Lam., Mwrex tyracutanuB L., Poli Tettac, utr. Sic m.
Tab. 48, figs. II, 12, Blaiky. Malacol, PL 15, fig. j;—Fusut eoUmau
Lam., BncyoL m^., Ver$. PI. 427, fig. i, &c.
Pyrvia Laic (excL some species). Shell subpyriform, wiih short
spire.
Sp. Pyrvia mehnffena, Mvrex Mdongena L., Rumfh. Amb. Bariieitk. Tab.
xxiy. fig. 2, Blainy. Malae, PI. 17, fig. 3; — Pyr. pervena, Mvrex peroer-
tus L., Bncyd Ven. PI. 43$, fig. 4, Bekvx OondL Sytt. n. PI. 136, figs.
5, Ac.
Murex L. (in part), Brug. Shell spiral with transverse tuber-
cles or spines in rows. Aperture rotundate, produced into a
straight canal ; columella smooth. Operculum homy.
MtMrex Laic Shell with homy tubercles or spines disgpsed in
several longitudinal rows. Canal of aperture often very long.
Sp. Mwrex Brandarii It., Poli Teatae. utr. SicU. m. Tab. 49, fig. 8, Guianr
Iconoyr,, MoU. PL 19, fig. i, Leiblkin, HKUSiNaEB's Zeittchr, f. die oryan.
Phyeik. EiBenach, 1827, I. Heft t, pp. i — 33, Tab. I. Ann, deeSe. naL XTf.
pp. 176 — 306, PL 10, 11; Mediter. Sea; — Mwrex Tribvlua It., Mwrex
cramtpina Lam., Blainv. Malac. PL 17, fig. 2 ; — Murex ienuiapma Lam.
(Mur. Tribvlua var. L.), Bumph. Amb. Bariteiik. Tab. 26, fig. 3, Kuhxb
iS^. ff^nfral des Ooq. viv,, ca$Mliftre», m. PL 6, ^, 1, PL 7, fig. 1 ; —
Murex ffauttdlum It., Blainy. McdaeoL PL 19, fig. 5, Gu&ik leonogr.,
MoU. PL 19, fig. 2, &C. Of thifl genofl also there are various foBtal spedes.
RcmeUa Lam. Shell ovate or oblong, with ribs compressed, vari-
cose, opposite, remote at the middle part of the wreath.
Mwrex Rama L. (in part), RamtUa erumena Lam., Bumfh. Jm&. Jtorv
teilk. Tib. 14, fig. o, Eneyel. fn&h.. Vera. PL 411, fig. 3, &e.
Triton Montf., Lam. Shell ovate or oblong, with varices rare,
not forming longitudinal series.
Sp. Mwrex TrilUmia L., ^Vtton xariegatwn Lam., Bumph. AnA. Rariieiti, Tab.
38, fig. B and I, Eneyel. mith.. Vera. PL 421, fig. 2, Blainv. MalaeoL PL
iS« %• 3; from the Indian Ocean; — Triton nodiferum Lam., Poli Teai.
vtr. Sie, m. PL 49, fig. 9 ; from the Medit. Sea. To this last species
belongs the anatomical deecription of K. W. Etsikhabot, Mbcxzl's
Archivf. d. Phyaiol. vin. 1823, pp. 113 — 317, Tab. m.
Fasctolarta Lam. Aperture ending in a straight canal. Shell
oblong, scarcely tuberculate. Columella furnished with two or
three oblique folds.
Sp. Faaciolaria tulipa Lam., Murex Tulipa L., Bumph. Amb. Rariteitk. Tab.
49, fig. H, Blainy. Malac. PL 17, fig. 2.
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MOLLUSCA. 805
TurbtneUa Lam. (species of Valuta L.). Shell turbinate or
subAisiform, with aperture terminating in a canal. Columella with
three or more transverse compressed folds.
Sp. TwbineUa pifrum Lam., Volwta Pyrum L., Rumph. Amh, BarUeiik. Tab.
36, fig. 7, Gu^iN Iconogr., MoU. PI. 20, fig. g;—lhfrbm. craticuUUa
Lam., Mwrex craticuUUut L., Encyd, mUtk., Vers, PI. 419, fig. 3. With
the exception of this species, Lutnaub in the hut edition of the Sytt, n(U,
brings the species of this genus known to him to VoliUa. The species are
numerous and resemble in form either Fusua or Pvndd, from which they
differ by the transversely striped columella alone.
Cancellarta Lam. Shell oval or turrite, mostly reticulate with
transverse stripes and longitudinal rugee. Aperture ending in a
short canal. Columella with transverse folds; labrum iurrowed
internally. .
Sp. CanceUaria raiculcUa Lam., Valuta reticulata L., D'Aboskv. Conck, PL
17, fig. M, Encycl. mith., Vers. PI. 375, fig. 3, Blaikv. Malacol, PI. 22,
fig. I, &c. GifyiEB regarded this genus as a sub-division of Valuta;
Deshates is inclined to place it in the neighbourhood of Tomatdla, There
is no operculum present. Many fossil spedee from the tertiary formations
are known.
Struthiolarta Lam.
Sp. Strutkwlaria nodulosa Lam., Mwrex pes sbrutkiocameli Ghemv., Spen-
GLEE NcUurforscher, xvn. Tab. u. figs, a, b, Blauiv. MaJUtool, PI. 17,
fig. 10, ftc.
Rostellaria Lam. (in part), Chenopus Philippi, Aparrhais
Petiver, Gray.
Sp. Strombus Pes peUcani L., PoLi Test, utr, SicU. in. PI. 48, figs. 7—10,
Blainy. McUac, PI. 11, fig. 6, kc,
Stromhis L. Shell turrite, with labrum dilated into a very
ample wing excised towards the canal by a sinus. Canal straight
or inflected by the columella obliquely outwards. Operculum
homy. Tentacles bifid, the internal lacinia conical, the external
peduncle cylindrical, oculiferous at the apex.
* Canal subulate; sinus of lahrvm contiguous to the canal. Spec, of
Rostellaria Lam.
Sp. trombus Fusus L., JRostellaria rectirostris Lam., D'Abgbny. Conch, PI.
10, fig. D, Reeve Conch, Syst, n. Tab. 346, fig. 4; — Boslell, curvirostra
Lam., Encyd. mSth,, Vers. PL 411, fig. i, Blainy. Malacol. PI. 16,
fig. I.
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806 GLASS XIII.
* * Canal dongaU; iinrn of labrum not wntigwmt to oanal, Lakmm
produced into digitcUe laeinuB, Pterocera Lam.
Sp. Stromima Ckiragra L., Libtbb Omck. PI. 870, fig. 34;— Srom^itf Scot-
piua L., D'Akoenv. Conch. PI. 14, fig. b, Blainy. Malae, PL 15, figs. 3,
4, GuiBiN Iconogr., Moll, PL 21, fig. z, &o.
* * * Canal tkort, emarginate or trttneaied. Sintu of lahrtan diatimet
from eancU. Labrum in more adult theUs expanded into a teing simfle,
entire, Sironibue Lax.
Sp. Strombue gigae L., Listxb Conekgl. Tab. 863, ^. 18 ;—Strombue latietC-
mu8 L., L18TBB Tab. 861, fig. t8 (856, fig. la young) i—Slromhua ladigi-
nonu L., Cur, S, Ani., €d, iU,, Moll, PI. 61, fig. i, Ac
Cassis Lam., Cassidea Bkug. (spec, of Buccmum L.). Shell
yentricose or inflated, with spire short Aperture longitudinal,
terminating in a short canal. Columella transyerselj plicate or
rugose.
a) Canal incurved obli^uelg iotkel^ and upmarde or tMbtbraighi, Ca§-
eidaria "Lau,
Sp. Caatit eehinophora, Buccinum edUnopkomm L., BmcPB. Amb, RasriUitk.
Tib. 37, fig. I, Gu^BiN icmogr,^ MoU, PL 16, fig. 6, fto.— The genua
Oniecia SowBBBT it diatinguished by a abort csnal and a lamina, covered
with granular inequalities, extending fi!t>nL the oolumeUa over the last
wreath : CasHdaria Oniecua Lax., Strombua Oniacue 1,., Ac. See Rbevx
ConcK, Sgal, n, pp. 210 — 212, PL 353, 354.
b) Canal aacending, exeieed deeply towarda ike extremity. Cauia Lax.
Sp. Oaaaia eomuta Lax., Buccinum eomuHtm L., Libtbb Oonchyl. Tab.
1008; — Caaaia tuberoaa Lax., Buccinum tuberoaum L., Encgcl, mitk,, Vera.
PL 406, fig. r, PL 407, fig. 3, Blainy. MalacaiL PL 33, fig. i ^—CaaaU
glauea Lax., Buccinum giaucum L., Ruxph. Amb. Bariieiik. Tab. 35, figs.
A and 4, Cnv. R, Ani,, id. HI., Moll. PL 56, fig. i, &a
FoBsil species also of this genua are found.
Purpura Brug. Shell oval, often tuberculate, with spire short,
last wreath ample. Aperture terminating in a sinus oblique, sub-
canaliculate. Columella smooth, acuminate towards the apex.
Operculimi thin, homy.
DvcLOB {Ann, dea Se, nat, xxv. 1833, pp. 90—95, xxvi. pp. 103—113),
Dbshates {EncycL miih.. Vera, m. pp. 838 — 845) and Blaibyille {Nomf,
Ann, du Mua. I. pp. 189 — 363) have united under the name of Purpura,
besides Monoeeroa and Bicinula of Laxabck, also Concholqm Lax. ; they
regard these genera merely as divisions of a great genus, to which more
than 300 species now belong.
Bicinula Lam. Aperture oblong, narrowed by unequal teeth at
the columella and the internal surface of labrum. Shell often tu-
berculatoHspinose.
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MOLLUSCA. 807
Sp. Pwrpwra riciniMf Mv/rex Memui L., Riemula wrackwAdM Lam., Enc,
nUth., Vers, PL 395, fig. ^i—Jiicinula horrida Lam., Blainy. Malac. PL
22, fig. 2, &c.
Purpura Lah. Aperture dilated, with columella short
Sp. Purpura peniea Lam., Buceinum persieum L., BuMFH. Atnif, HariteUk.
Tab. 27, fig. b/ Blaihy. McUacol, PL 14, fig. 3. (Most of the spedea
belong to this division, of which fossil speoies also occur.)
{Monoceroe Lam. differs from the Purpura by a conical tooth at the in-
ternal mai-gin of labnim. Sp. Purpura monodon, Monoceros imbricattim
Lam., Buccinum monodon Gmel., Blaint. Malac, PL 17, fig. 3, &c.
Conchol^pas Lak. Vertex not spired, inclined obliquely towards
labium. Aperture longitudinal, oblique, very ample.
Sp. Purpura eonchoUpM nob., OimchoUpas peruvianui D'Arobitv. Conch.
PL a, fig. D, Blaimv. Malacol. PL 94, fig. i, Lessoit lUuMtr. de ZooL PL
27 (a figure with the animal). This singular shell, which at first sight
might be taken for a single conch of a bivalve, was received by Gmblin
into the thirteenth edition of the SyH. not, under the name of Patdla lepas,
Harpa Lam. Shell ovate, turgid, with longitudinal parallel
ribs. Aperture oval, terminating at the apex in an excised sinus.
Columella laevigato-gibbons, oblique. Spire short. Animal with
foot large, semicircular anteriorly, and operculum none. Tentacles
conical, incrassate at the base, oculiferous on the outside.
Sp. Harpa imptrialU Lam., Buccinum cosUUum L., D'Abgbnv. Conchyl.,
Append. PL iy fig. P, Chemn. Tab. 15a, fig. 1451; — Harpa veniricoea
Lam., Buccinum Harpa L. (in part), EncycL nUtk,, Vers. PL 404, fig. i,
Gui^iK Iconoffr., Moll. PL 18, fig. i. Comp. Rbtnaud Obdervations swr
V Animal de la Harpe, Mim. de la JSoc, d^Hisl. not, de Parte, v. 1834, pp.
33—40, PL 3-
Dolium Lam. Shell thin, ventricose, smooth, surrounded by
transverse ribs. Aperture longitudinal, terminated anteriorly by a
sinus. Columella oblique. Labrum transversely crenate. Spire
short. Foot large, without operculum.
Sp. Dolium perdix Lam., Buccinum Perdix L., D'Aboenv. Conchyl. Tab.
17, fig. A, Mabtini Tab. 117, fig. 1079, GufiRiif Iconogr., Moll. PL 17,
fig. 10, Cuv. JL Ant., id. ill., Moll. PL 54, fig. 2 ;—J)olium galea JjAM.,
Buccinum galea L., Mabtiki Tab. 1 16, fig. 10^, PoLi Test. utr. Sic. ni.
PL 47, figs. 3, 4 ; ib gigantic species from the Mediterranean Sea.
Buccinum L. (excl. many spec), Lam. (add genus Nassa Lam.).
Shell ovate or ovato-conical, gibbous. Aperture longitudinal,
emarginate by a sinus towards the apex. Operculum homy.
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808 CLASS XIII.
Sub-genera : Nassa, Buccinunh, Tritonium Muell. and Debhatis
(species of Bticdnwrn Lam.). Comp. Deshates in Lahabck Hist
not, dea Ani, a. vert, x. pp. 151, 152.
Sp. Buccinum undaiwn h., Encyd, mOh,, Yen. PL 399, fig. i, Blaxst.
Malae, PL ^2, fig. 4. (See an anatomicid description of this spedes bj
CuviBB, Ann. du Mtu. xi. pp. 447 — 457, Mim. s. I. MoUutq. No. 17.)
To Buccinum belongs also the genus Ebuma Lam. (with the excLouon of
Ehuma glabrata, Ancillaria glabrata Soweebt). Sp. Bttccinum tpiraimm
L., BUMFH. Amb. Bariteid. Tab. 49, fig. D, Encyd, mith.. Vers. PL 401,
fig. 4.
Terebra Brug. (species from genus Terehra Adans.), Subula
Schumacher, Blainy. Shell turrite, subulate. Aperture longi-
tudinal, much shorter than spire, excised at the apex by a sinus.
Columella oblique, plicate ; labrum with margin acute, thin. Oper-
culum homy.
Sp. Tertbra mactUata Lau., Biiccinum maculatum L., D'Abqbitv^illb Com-
ckyl. PL II, fig. A, Encyd, mith., Ven. PL 402, fig. i.
Conus L. Shell convolute, conical. Spire short, sometimes
acuminate, mostly flattened. Aperture narrow, longitudinal, eden-
tulous. Columella short. Animal with foot anteriorly truncated,
posteriorly attenuate, rotundate, with homy operculum ; head with
two tentacles subulate, bearing eyes on the outside towards the
apex.
Sp. Cmua imperialis li., RuuPH. Amb. RariicUk. Tab. 34, fig. H, Encydop.
mith., Vers. PL 319, figs, i, 2, Blainv. Malctc. PL 26, fig. 5;— Cwta
gloria maris Chemn., Conch. Tab. 143, figs. 1324, 1325, Enc. nUth., Vers.
PL 347, 6g. 7. A very numerous genus of more than 300 species (aooord-
ing tu Debhatbs, Lamabck Bist. not. d. Am. s. v., sec. id. xi. p. 7).
Beautiful figures of many species are found in the Encyd. mith.. Vers. PL
315 — 347. Almost all are from the tropical seas ; many fossil species from
the tertiary formations are known.
Voluta L. (exclusive of many species). Lam. Shell ovate, very
often ventricose, with apex obtuse. Aperture longitudinal, excised
towards the end by a sinus. Columella obliquely plicate. Ani-
mal without operculum.
Sp. Voluiu cethiopica Lau. {Voluta cethiopica L. in part), Encyd. mith..
Vers. PL 387, fig. i, Cuv. B. Ani., id. Ul., MoU, PL 52, fig. i, Ac.
Volvaria Lam. (in part), Sowebbt.
Species fossil. Comp. Rsbys Conchol, Syst, n. pp. 246 — 248.
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MOLLUSCA. 809
Mitra Lam. Shell mostly fusiform, with apex acute. Folds
of columella suboblique, transverse, decreasing forwards. Aper-
ture narrow, broader anteriorly, terminated by a deep oblique sinus
ascending upwards. Animal with tentacles oculiferous externally at
the sides or at base ; exsert proboscis surpassing the shell in length.
a) Wiih ipire dongaU,
Sp. MUra tpigcopalie Lam., VoltUa mitra L. (m part), Bumfh. Amh, Rari-
teith. Tab. ig, fig, K, D'Argbnvillb ConchyL PI. 9, fig. 0, Blajnv.
Malacol. PI. 2S bia, fig. i, Kienbb Coq. viv., Columellaires, MUre, PL i.
fig. I (fig. wiih the animal) ; — Mitra vulpeetda Lam., VoltOa ViUpecula "L,,
GuiRiN Iconogr., MM, PI. 17, fig. i.
b) With tpire shorter, {ffahitut nearly of Conus.) Conohdix Swains.,
SowEBBT, Imbricaria Schumachsb.
Sp. MUra dadylua Lam., Valuta Dactylut L., Blainy. MalacoL PI. 38 bis,
fig- 3> Gu&lN Iconogr., Moll. PI. 1 7, fig. 2.
Margtnella Lam. (with addit. of several spec, of Volvarta ejusd.).
Shell ovato-oblong, smooth, with short spire. Aperture scarcely
sinuate. Columella with folds transverse, oblique, subequal. La-
brum incrassated externally.
Sp. Marginella faha, VoltUa Faba L., Encycl. m^th., Vera. PI. 377, ^. 1,
Blainy. Malac. PI. 30, fig. 5, &c.
Terebellum Lam. Shell convolute, subcylindrical, with apex
acute. Aperture longitudinal, narrow, broader towards the end,
terminated by sinus. Columellar margin produced with apex
truncated beyond labrum. (Animal with tentacles filiform, oculife-
rous at the point.)
Sp. Terd)dlum tubuUUumt BuUa terebellum L., Humph. Anib. Mariteilk. Tab.
30, fig. B, Blainy. Malac. PI. 37, fig. i, Adams and L. Bseyb Mdlusca
of the Voyage of H.M.8. Samarang, 1848, PI. ix. fig. 6 (a fig. of the animal
with the shell) in the Ind. Ocean. Terebellum convohUum Blainy. 1. L
fig. a a foflsil species from the calcaire grostier.
Columbella Lam. Shell oval, spire short. Aperture emargi-
nate towards the end. Labrum gibbous internally, often denticu-
late, narrowing the aperture.
ColwnheiUa ruatica Lam., Valuta nutiea L., PoLi Teat. utr. Sic. iii. PI.
46, figs, s^—41 y—ColurnbeUa atrombiformia L., Blainy. Malac. PI. 30,
fig. 5. &c.
Ancillaria Lam. Shell oblong, smooth. Aperture longitudinal,
sinuate towards the end. Columella tumid, contorted. Spire short,
with sutures of wreaths not canaliculate.
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810 CLASS XIII.
Sp. AneiUaria cinnamonea Lam. {BtdlA eypraa L. I), &c. (A geons narcdy
distinct from Oliva^ of which perhaps it is to be regarded as a sub-generic
section.)
Oltva Brug., Lam. Shell smooth, subcylindrial, convolute,
polished, with spire short and sutures canaliculate. Aperture
longitudinal, terminating in a sinus. Columella obliquely striated.
Labrum somewhat thick, with margin inflected towards the aper-
ture. Animal with foot elongate, furnished anteriorly with a
triangular lobe, without operculum; tentacles incrassated at the
base, oculiferous on the outside, attenuated at the apex.
Sp. Oliva porphyria Lam., Voluta porphyria li,, D*ABOJiinriLLB Conck. PL
13, fig. K, Encycl. tn^lh., Vers. PL 361, fig. 4, Reeve Conch, SyH, n, PI.
a 7 3, fig. I ; — Oliva maura Lam., Voluta (Hiva L. (in part), Rumfh. Amb.
Rariteitk, Tab. 39, fig. a, Cuv. R, Anim,, €d. iU.,MoU, PL 51, fig. 3, ftc
The species are numerous, many much resembling each other, and on
account of the manifold varieUes which some present, difficult to distin-
guish. There are also some fossil species known, which occur in the ter-
tiary formations.
Ovula Brug. Shell involute, turgid, polished ; spire concealed^
Aperture longitudinal, narrow, produced at both ends beyond the
columella, with columellar margin edentulous. Labrum often in-
crassated, inflected, flat, transversely crenate.
Sp. Omda oviformia Lam., Bulla Ovum L., Rvmfh. Amb. Ba/nieUk, Tsb. 38;
fig. Q, Blainy. Malac. PL 31, fig. i ; white, the outer margin of the moath
toothed, just as in Cypraea, from which genus this species is in fact distin-
guished merely by the smooth, convex inner margin. The animals, more-
over, of the two genera are vexy similar. In other species the external
margin is smooth ; such is the case in Ovula vnha Lam., Bulla Volva L.,
Blainv., L L fig. 3, where it tenninates at both sides in a long tube.
Cyprcea L. Shell involute, polished, convex above, flattened
at the aperture. Aperture straightened, longitudinal, effiise at
both ends. Either labium transversely crenate. Spire very small,
concealed in adults. Animal with tentacles conical, the eyes
placed externally at the base of tentacles, foot broad, operculum
none ; mantle very ample, reflected upon the shell at the sides.
Porcelain ahells. A very numerous genus (about ^00 species are known),
which is defined very naturally. The shell of the young animal difiers
mudi firom that of the adult; it is thin, differently coloured and without
teeth at the aperture, the wreaths also being then visible (see above, p.
687, and compare the figure of Cyprcea exanthema in the young state,
Encycl. mith., Vera. PL 349, or in Blainy. Malac, PI. 30, fig. 1).
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MOLLUSCA. 811
Sp. (}yprmk ArgM L., Ruvph. Tab. 38, fig. d, Bneyd. mSth,, Vers, Fl. 350,
fig. I ;—OypraM varMaria Lam., City. R Ani,, id. iU., Moll. PI. 49, fig.
4 (a fig. of the aniinal with the shell transferred from QuoT et Gaimabo
Vay. de VAttrol.) i—OjfprcBa mtmda L., Rumfh. Amb. Bariteitk, Tab. 39,
^. 0, Encycl. tneth,, Vers. PI. 356, fig. 3, a species known by the name of
Kauri {Cowry), which serves for small money in some countries of Africa
and India.
Section III. Cavity for respiration furnished with a network
of vessels running on its walls, opening ektemally by an aperture
of mantle.
Family XI. Pneuman{ca{Coelopnoa ScHWEia.). Hermaphrodite
molluscs breathing air, most of them terrestrial, others aquatile,
ascending to the surface of the water for respiration. Some naked,
others testaceous.
This division of Mollnscs is treated of at length in the iUnstrated work of
Db F^ubsao, Hittcirt natur. des MoUutques terrutres ei Jhiviatiles, Paris,
1 8 19 and foU. Dbbhatjis, after the death of FfBUBSAO, began the con-
tinuation which was completed in 1851.
I. Operculum,
Ampullacera QuOT and Gaim.
Note. — Shell of AmpuHaria. Head bilobed, with rounded lobes in place
of tentacles. Here belong the species of A mpullaria (AmpuU. avdlana and
Amp. froffUis) of Lamabck. The genus seems to me to be of uncertain
position.
n. Operculum wme true.
Phalanx I. Eyes at the base of tentacles. Two tentacles
only. The most aquatic.
Ancylus Geoffr. Shell thin, obliquely conical, with apex
acute, inflected posteriorly. Aperture oval, with margins very
simple. CJreeping body entirely covered by shell.
Sp. AncyluB lacustris Muell., Patella lacustris 1,., PFEirFEB Schn. i. Pi. iv.
fig. 46 ; — AncyluB Jluviatilu, Patdla JUgnatUis GiCBLUi, Blaint. Afalaeol.
PL 48, fig. 6, Stubm DeuUchl. Fauna, vi. Heft 4, Tab. 9, Pfsiffeb 1. 1.
fig. 44.
Concerning the place of this genus in the natural arrangement opinions
are at yarianoe. Tbxvirakus thought, from the anatomical investigation
of Ancyl. fiuviatilit, that the genus might indeed have some affinity with
LymruEuSf but still does not belong to the pneuinmUca, because on the left
side, between the mantle and the border of the ventral disc, a gill is situated
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812 CLASS xnL
in fonn of a long Inf. Tsxvnujnjs, aooonfin^y, tumymita tkb aunl
with Pleurobranekut, but remarks at the ame time that in thie laat gm»
the gill liee on the right side, and hae many tnasrene folds, whikt that of
Aneylui preeenti an entirely smooth snz&ce. (Zoledbr. fwr Pkytiaiagkj Vf.
1, 1831, B. 194—197, Taf. xvn.). Tioscbsl icfets AmejfimM to the Bgpo-
Wanekiaia (see WixoMAinr's Ardkiv, 1838, n. s. ^5). Bat ainoe, however,
YooT has subsequently remaiked, that these animik appear to breathe air
alone, and that it has by no means been proved that the leaf4ike organ if
really to be regarded as a gill (Mukllke'b Ardkiv, 1841, s. 35 — 38), wcitil
think that Ancj^ui must remain amongst the fmemmomiea laqmatSia,
LtmruBus Cuv., LymruBa Lajc. Tentacles broad, short, com-
pressed, triangular, not retractile. Eyes placed between the bases
of tentacles. Shell thin, oblong, sometimes turrite; spire exsert
Aperture entire, longitudinal. Labrum acute, ascending towards
columella and forming an oblique fold.
Mud-tnaU. These animals have a thin shell; the last wreath is very
wide and round. On the right side of the animal is the respiratoiy sper-
ture, the walls of which can be elongated as a tube, and thos extend abo?e
the water. Such Lb the case when the animal respires at the saAce of the
water, with the foot or the ventral disc tumed upwards. The apertures of
the sexual organs lie far apart Under the right feeler is the aperture
which transmits the penis, under the respiratory aperture that of thefemak
organs. To this it is to be ascribed that in copukting one individasi ii
connected with two others, one of which impregnates it whilst the other in
impregnated by it. In this way, hanging together, th^ often form seveni
long chains. (Vok Basb has, however, observed self-impregnation also in
Limnetut awrieularu, MuBLUm's Arekiv, n. 1835, s. 114.) These snsik
live on water-pUnts.
Compare Swahmbsdam Bijbd der NaL i. U. 164—169, Tub. ix. ^. 4;—
CuviBR Ann. du Mus. vu. pp. 185—193, PI. x. figs. 1— 11, MoUusq., Min.
Ko. 14;— Stikbkl DUseri. inmtg. nttrnt Limtui stoffnalit Anatomen, Got-
tingn, 18 15, 4to, c. tabulis 1,
Sp. LimiMeuM ttagnaiit, ffdix dagnalis L. (703), Bladtv. Malae. PL 37i
fig. I ; Pfsiffsb Schn. I. Tab. iv. fig. 19, &c.
Sub-genu8 Amphipeplea Nilsson. Mantle inyolving dieU Shell
subglobose ; spire obtuse, scarcely prominent.
Sp. Limnceut glutinotut DBATAiur.
Physa Draparn. {Bulinu8 Adans.) Tentacles slender, seta-
ceous, long. Foot long, anteriorly rounded, posteriorly narrow, acute.
Margins of mantle sometimes fimbriate, reflected over shell. Shell
oval or oblong, often left-handed ; aperture longitudinal, produced
anteriorly, enlarged.
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HOLLUSCA. 813
Sp. Phyta fontinaZii Drap., BuUa fonttnalis L., Stubm Deutschl, Fauna,
VI. Heft 4, Tab. lo; — Phyia hypnorum Dbap., Bulla hypniorumli., Stubm
1. 1. Tab. II, Cuv. U. Ani,, id, iU., MoO, PI. i6, fig. 5.
Planarbis Brug. Tentacles setaceous, very long. Foot short.
Shell disciform, with all the wreaths conspicuous on both sides.
Aperture almost lunate, with margin not reflected. ^
These snails live with Limnaut in stagnant fresh water, and hare, how-
eirer remarkably the external form differs, almost the same internal structure
as the species of that genus. Compare Swammsboam B^bel der Nat. 1. bL
186 — 189, CuviEB Arm. du Mu9. vn. pp. 194 — 196, PI. x. figs. i« — 15,
MoUtuq., Mim, 14.
Sp. Planorhit comeus, Hdix cornea L., Blaikv. Malaeol. PL 37 bis, fig. 3,
Stubm L L Tab. 4, Gu^bim Iconogr., MoU, PI. 7, fig. 1, &;c.
Auricula Lam. (spec, from genus Voluta L.). Two conical
tentacles. Shell oval or oblongo-orate. Aperture longitudinal,
with labrum often reflected. Columella folded.
To this genus belong species that live on land, although elsewhere these
molluscs, like Hdix, are distinguished by four tentacles. Different genera
have been adopted here, as Carychium Mukll. and ScarabuM Montf.,
Scarabcea Cut. This bst genus is distinguished by a shell longitudinally
compressed into a margin on each side (carinated).
Compare Dbshatbs Diet, univers, cTffiat. not. 1. 1842, pp. 345 — 348,
the article Awicule; Tbosohxl in Wieqmank'b Archiv, 1838, 1, pp. 102 —
308, on Scardbiis, and Kkbvb on the same sub-genus, Ann. of Nat. Hut,
IX. pp. 218 — 221.
Sp. Avrieida mida, VohUa Aurit Midoi L., D'Abokky. Conchyl. PL 10,
fig. o, Encyd. mSth., Vers. PL 460, fig. 6, Gu^bik Iconogr., Mollusq. PL
7, fig. 7, QuoT et 6aih. Attrolabe, Zool. PL 14 (fig. with the animal, trans-
ferred in GuviXB R Ani., id, iU., MoU, PL 27, ftg, i) ; New Guinea, not
far from the shore ; Awricvlajttdas, VohUa AurU Judce L., Blainy. Malaeol.
PL 38, fig. I ; — Aurictda tcaraJbaa, Hdiz ScaaraboBUB L., Scarabus tmbrium
MoKTF., Chexk. Tab. 136, figs. 1249, 1250, Bejcve Ann, of Nat. Eitt. L L
PI. 4, fig. 1 1 ; lives in the E. Indies, in woods at the roots of trees, ko.
Phalanx II. Eyes at the point of tentacles. Tentacles most
frequently four ; two superior placed more backwards, larger ; in-
ferior short (sometimes obsolete). MoUusca pneumonica terrestria
aut amphibia.
A. Testacea (Helicina),
Pupa Drap., Lam. (spec, of Turbo L.). Animal with inferior
tentacles very short or none {Vertiffo Muell.). Shell elliptical or
cylindrical, with aperture semirotund, in most dentate.
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814 CLASS XllL
Sp. Pupa muaeorum Lail, Turbo muteorum L., PmmB SAn. i. Tib. 3,
figs. 17, jS;—Pupa mumia Laic, Blaihy. Malae. PL 38, fig. 5, fcc
Add genus Ckcmdrvi CuY.
ClauatUa Drap., Lam. (species of Turbo L.). Animal with
inferior tentacles veiy short Shell very often thin, fiisiform, with
apex sbmewhat obtuse. Aperture snboTate, with margins at each
side connate, reflected. (In the last vrreath in most a testaceous
lamella adhering to columella by a thin elastic pedicle^.)
This genus is much allied to Pupck, and may, as Deshatbs deddes, be
united with it. In most of the species the sheQ is left-haiided. Sndi ■
the case with aU the European species, which are small, aod are found os
trunks of trees, on the ground amongst faUen leaves and on stones. Sp.
ClaudLia rugo$a Dbap., HeUx penena Mubll., Pfstffbb Sckti. i. Tth.
HI, fig. 30. Here also belongs the genus Cylindrdla Ptkeffbii.
BuUmus Deshayes, nob. {Bulimua and Ackattna Lam., species
of genus Bulimus ScoPOLl, Brug.). Animal with four distinct
tentacles, and foot lanceolate posteriorly. Shell ovate, more rarelj
oblong, cylindrical, with last turn rather large, and apex of spiie
somewhat obtuse. Aperture longitudinal. Columella smooth.
AchctUna Lam. Columella truncated towards the end, not con-
tinuous with margin of aperture. Labrum always acute.
8p. Svlimut virginetu, BvBa virffiihea L., D'Abokkt. Ooneh^. PI. ii^ fig-
K, Blainv. Malae. PI. 38, fig. 2, Cuv. R. Ani., id. ill.,3foa. PL «5, fig. 4;
South America ; — BvJimut columna, ffdix columna Guel., EnqfcL miih.f
Ver$. PL 459» fig. 5, Blaihy. Malae. PL 40, fig. 3, kc.
JBviimits Lam. Columella not truncated, continuous with mai^
of aperture. Labrum often reflected, incrassated (spec, of Melix L).
Sp. BvlifMU lubriau Bbug. {ffelix iubcylindriea L. ?), Sturm DevttckL
Fauna, vi. Heft 4, l^b. 4, Pfeiffxb Sdm. i. Tab. 4, fig. 6. BuUmut
citrinui Bruo., Helix perversa h., D'AROXNvn.LB Conch. PL 9, fig. ^f
MouBSOK Tab. XX. fig. 5, fto.
Helix Brug. (Species from genus Helix L.). Four distinct
tentacles. Shell orbicular or depresso-conical. Aperture broader
than long, lunate, with penultimate wreath prominent intemallj.
Comp. Dbshatrs Did, cUusiq. SHUt, not, vm. 1815, pp. 74—^
Encyd, mStk., Ven. n. pp. 188—366, under Helix,
^ Compare on this part, to which the genus dautiHa owes its name, J. E. GbaI
in Annalt of Nat, Hitt, V. p. 143. It is no operculum, but rather a modification of tbe
tooth-like prominence at the aperture of the sheU occurring in many moUusoB.
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MOLLUSCA. 815
t Apertwn twmed upwardt, dcntaU on each nde,
Anostoma Fischer, Lah.
Sp. ffdix ringens L., D'Arosny. Conchyl, Fl. 38, figs. 13, 14, Blainv.
Malac, PI. 39, fig. 4; habit, in £. Indies^.
I I Aperiwre inclined downwards, tkell <it the periphery ecmnaie er wb-
earinate, moeUy depreend, Ldbrum often r^fUeied, eomeUmet denttUe,
CarocoUa Lam.
Sp. Helix Lapicida L., Pfeiffsb 8chn. i. Tab. n. fig. 26;^JIelix OaroeoUa
L., D'Abobnv. Conch, Tab. 8, fig. d, Gu6rik Iconogr,, MoU. PL 6, fig. i;
from the £. Indies, &c.
I I I Aperture inclined downwards. Shell rounded at the margin, moet
frequently eubglobose.
Ifelix Lajl
Sp. ffdix Pomatia L., Sturm Devtseld. Fauna, yi. Heft i, Pfeiffer Schn»
I. Tab. n. fig. 9, Cuv. R, Ani., Sd, ill., Moll. PI. 2 1. The Vineyard-Snail, le
grand Escargot. The name Pomatia, from Jlufia, cover {cochlea opercvlarie),
was given to this animal on account of the closure of the aperture of the
shell in winter that occurs in this and other species. In autumn the animal
retracts itself within the shell, and then a false operculum (epiphragma) is
secreted^ which is pushed off in spring when the snail again creeps out of
its house. During the winter these snails in our temperate climate take no
food, but lie asleep ; in Hke manner, in tropical regions the species of this
genus in the dry months fall into a state of insensibility.
Por the anatomical particulars we refer to Swammerdam JBijbd der Nat,
I. pp. 97 — 147; CuviBB Ann. du Mue. vn. 1806, pp. 140 — 184, MoUuaq.,
Mim. No. 1 1 ; and Wohnligh Dim. anat. de Hdice pomatia cum tab, am,
Wirceburgi, 18x3, 4to. In all the species of Hdix there is found in the
neighbourhood of the sexual organs a thick-walled pear-shaped purse, in
which a dart-shaped calcareous stile is contained ; this ox^gan is annually
renewed, just like the cover of the mouth of the shell.
Hdix advpersa MuELL., Sturm DeiUechl. Fauna, yi. Heft vi. Tab. 4 ;
this species nearly attains the size of the preceding; — Hdix nemordlie It.,
Sturm 1. 1. Heft 2, &c. On this and other native species, the figures of
Pfeiffer and Sturm may be compared. This genus is also remarkably
numerous in exotic species.
Sucdnea Drap. Animal larger than shell, not entirely covered
by it, with four tentacles short, especially the inferior. Shell ovate,
with spire short ; aperture ample, longitudinal, with labrum acute,
columella short.
^ Some fossil shells from the tertiary formations are known, which also have an
aperture turned upwards ; but this is round and not toothed. They form the genus
StrophotiMML DebhateS) and might be brought to Oyeloitoma (p. 801). See Dbbhates
M&m. »ur le StrophottOTne, Ann. dee Sc. not, xiii. 1826, pp. 481—287.
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816 CLASS XIII.
Sp. Suecinea amj^tbia Dbap., Hdixpuiru L., Bladty. Malae. PL 38, fig.
4, Stdbm DeutacM. Fauna, vi. Heft i, Pfeiffeb Sekn. i. Tab. l fig. 4,
Tab. m. figs. 36 — 38 ; very oommon in canals, on water-plants^ &c.
Vitrina Drap., Lam. {Heltcolimax F^RUSS.). Body elongate,
limaciform; mantle with an appendage reflected over the shell;
tentacles four, two inferior very short. Shell depressed, small,
very thin, imperforate, with spire short. Aperture large, rotundato-
lunate.
Gomp. D*AUDSBABD FisusSAO Nahuforweker, Vol. 29, pp. 336 — 141,
Tab. I.
Sp. Vitrina peOucida Dbap., ffdix peUueida MuxLL., Vitrina henfiina
Pfkiffxb, Stubm DeuUchi, Fauna, vi. Heft 3, Tab. 16, GufetiN letmogr.,
MoU. PI. 5, fig. 3; on water-plants, and on the ground amongst mo»ei
and fallen leaves.
B. Nuda. Shell none or concealed beneath the mantk
small. Limacina,
Ltmax L. Tentacles four.
Farmacetta Cut. Mantle placed upon the back, emarginate in
the middle of right margin, including posteriorly the shell spired
obsoletely. Orifice of respiration and vent below a fissure at the
right side of mantle, generative aperture between the tentacles of
the right side.
Sp. Parmacdla Olivierii Cuv., Ann. du Mva. v. PI. xxix. figs. 12—15;
MoUutq., Mim, xn. ; Blainv. MalaeoL PL XLI. fig. 3, GuiSBDr Iconoffr,,
MoUiuq, PL V. fig. 5, &c.
TeHaceUa Lam. Bespiratoiy cavity and vent above the posterior
extremity of body. Small mantle situated in the same place, in-
cluding a very small shell spired obsoletely at the apex. Aperture of
generative organs below«the larger tentacle of right side.
Sp. Tettacdla halioHdea Cuv., Ann. du Mut. v. p. 440, PL xxix. figs. 6—
1 1, MaUusq., M6m. ill., Blautv. Malacol, PL xu. fig. 2, GcAbht Ico-
nogr,, MoUu$q. Pi. v. fig. 4.
Vaginulua Lam.
Limax Lam. Body oblong, naked, creeping, convex on back, fur-
mshed anteriorly with a coriaceous, subrugose mantle, and below
with a longitudinal flat disc. Branchial cavity under the shield or
small mantle, in the anterior part of back. Respiratory orifice and
vent placed in the right side ; generative orifice beneath the ligl^t
tentadea
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BIOLLUSCA. 817
Ground-slug, PcUh-slug. The mantle is here small and in Bome contains
a calcareous grit, in others a small shell. At the hinder end of the body is
a small aperture, irom which adhesive raucous threads proceed. Here
the viscera are placed in the body and not in the mantle, as in the snails
(Helices), The internal structure however has much resemblance to that
of the Helices; but the calcareous dart and the purse that contains it are
here absent.
These animals live on herbs, fruits, &c., and dwell in moist places.
Comp. Guv. Ann. du Mus, vii., MoUusq., Mim. xi.
Sp. Limax einereus Muell., Limax maximus L., Blaiitv. Malacol. PI. xu.
fig. 5 ; Stdbm Dtulschl. Fauna, vi. Heft i ; — Limax ater L., Stubu 1. 1.
Heft I. ; Limax rufiu L., Stu&m 1. 1. Heft I., Blainv. 1. 1. fig. 6, &c.
OncMdium (Buchanan) Cuv., Peronia Blainv. Tentacles
four. Body oblong, creeping, marginate everywhere by thick
mantle. (Orifice for the passage of penis under the right tentacle ;
vent, respiratory foramen and aperture of female genitals beneath
the posterior extremity of mantle. Veil emarginate over the mouth,
or two auriform appendages at the sides of mouth.)
Half a century ago a slug-like animal was first discovered by Buchakan
in Bengal, on the leaves of Typha dephanHna, to which, on account of the
many small tubercles on the back, he gave the generic name cited above
(derived from 6yKos). See Linn, Trans, v. pp. 132 — 134. The animal dis-
covered by B. has not, to my knowledge, been described more particularly
by any later observer; the animals to which CuviJiB gave this generic name
Uve in the sea near the shore, and also come on land. According to
Ehbenbbbo {Symbol, physic., EverUbr. i.) there are seen behind on the back
arborescent appendages which contract when the animal is on dry land and
which may be regarded as gills. Thb animal may thus form the transition
to the Qymndbranchiata and have an affinity to Doris,
Sp. OncMdium Peronii Cuv., Ann. du Mus. v, pp. 37 — 51, PL vi.; Moll,,
MSm. No. 13, Peronia mawrUiana Blaikv. Malacol, PL 46, ^. 7.
Note. — Peronia ferruginea Less. III. de Zool. PL 19, is it an Onchidoraf
Order III. Cephalopoda.
Molluscs with distinct head, the organs of motion, tentacles or
arms, crowning the mouth. Body sacciform, open anteriorly.
Sexes distinct. Respiration branchial. Animals all marine, tes-
taceous or naked.
CephalopocU, These animals present clear vestiges of an internal
cartilaginous skeleton. In the head is found, in most, as in Sepia,
Loligo and Octopus, a cartilaginous ring, through the aperture of
which the oesophagus passes and of which the uppermost part covers
VOL. I. 52
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818 CLASS XIII.
the oerebnd gangjiim, the lowest oontains the aoditoiy oigajia, and
the latenJ puts, broad and excavated like a eup, receive the eyea
The orbits are goarded in front by two elongated cartilaginous plates,
which are attached by their base to the cartilaginous ring and else-
where are extended freely over those cavitiea In the S^na there is
an additional triangular cartilage, of which the point is attached to the
ring; the base tamed towards the armSy and to which the muscles
of the arms are affixed. In Sqna and Lcligo there are farther
found two catilaginoos plates on the back, and, on each side of the
body, an elongated flat cartilage which supports the lateral expan-
aioDB of the mantle or the fins'. In the NautUua there is only one
cartilage present in the head, which lies beneath the oesophagos, but
does not enclose this annularly, and is prolonged on each side to the
fimneL The eyes^ which in this genus are placed on pedicles at the
tide of the head, are not protected by this cartilage.
The head of these molluscs is round and broad and separated by
a constriction, as by a neck, from the body. Around the mouth an
placed in meet eight or ten arms; in Xauiilugf on the oontrazy,
several membranous sheath-like appendages^ perforated internal] j,
in which thin cylindrical tentacles, capable of extension and retzac-
tioUy are indoeed Where eight or ten arms are present^ eight of
ihem have constantly a somewhat conical form and stand in a circle
round the mouth On the upper sur&oe, that which feuyoR towards
the mouth, these ei^t arms are oovned with numerous suckers, bj
which they attach themselves to difforent bodiea (In the ten-armed
two are placed on the oatside beyond the circle of the other ei^t on
the ventral surfiM», and can by retortion be drawn entirely within
the body. Already had Ajustotui distinguished these two from the
other anna under the name of proboiddesS) The arms are hollow
internally, for the passage of the artery and nerve ; radiating mus-
cular fibres run from the middle outwards, and on the surfiAoe is a
layer of circular and longitudinal muscular fibres,* by which all the
various motions of retraction, contraction, flexure and convolution
become possible, which, on the seixure of prey, are executed by
these ofgana
Under the head is situated a ccmical organ, which is open at the
extremity, the frumel (injundibulum) formed by an extension of the
mantle and provided with muscular walla. In Haniilus it has two
^ See 0. A. QcBVvm in Msckkl's Arckiv /Or dU Pkytioiogie, iv. iBiS, a. 334"
338, Tab. rr. fig. I.
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M0LLU8CA. 819
margins, like a rdled lea^ free beneath, which are thrown one over
the other ; in the other Cephalopoda it opens at the anterior extremity
alon& Below, this funnel passes into the cavity within which the
giUs are situated, and into which the terminations of the sexual
oigans and of the intestinal canal open. The contraction of the
funnel forces out the water at its opening below the head. Before
the opening there is usually placed a valTular organ (not unlike an
epiglattia), by which the reflux of the water is {vevented.
The mouth lies in the middle between the arms, surrounded by a
circular fold of skin. Here are seen two large, curved, homy jaws,
which resemble in some degree the bill of a parrot, and of which the
lower overlaps the upper and smaller. Between the jaws is placed
a membranous tongue, which is soft at the point and beset with
papilke, but elsewhere is covered with hard, homy plates and
spinesL In the sac of the mantle most of the viscera (with exception
of the gills) are situated in a proper cavity, which is distinct from
that of the gills that opens externally through the funnel
The respiratory organs are two or four gills, which, pointed ooni-
cally, run obliquely upwards, and are situated, one or two on each
side, in the base of the sac, which terminates in the funnel They
are afiBxed here to a fleshy, projecting margin, and consist of mem-
branous leaflets, incised at the edge, each of which is affixed by a
separate stalk to the projectii^ margin. In those that have only
two gills, a venous heart is situated at the base of each of these ; in
the NcmtUu8y that has four gills, these two venous hearts are not
present. On the large venous stems which conduct the venous
blood towards the gills, spongy appendages, usually of a brown-
yellow colour, united in bundles, and sometimes subdivided blind
foUicuU^ which we noticed above (p. 760) as kidneys, are situated.
The cavities in which these veins lie, are washed by water that has a
free access to them through special apertures in the branchial cavity.
The sexual organs, in both sexes, are situated in the base or pos-
terior extremity of the visceral saa The ova/nvm is in all single,
although in many two oviducts are present, which then, however,
arise from a short common canal. This ovary consists of a sac with
thick waUs, on the inside of which, the eggs inclosed in proper cap-
sules, are affixed by pedicles. For the secretion of the envelop of
the eggs there is a glandular organ, which in Octopus surrounds each
oviduct, or the walls of the oviduct itself are thickened towards the
extremity by a glandular tissue which supplies the same secretion.
One or two glands composed of many laminee, which however are
52-2
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820 CLASS XIII.
wanting in OctopuB, are present in females, without any oonnexioa
with the oviducty and probably secrete an external cofvexmg, or an
adhesive substance, by which the egg? are attached to each other.
The testis in the nude agrees for the most part with the oTsry in
external form and in situation. It presents a sac in which a bundle
of glandular tubes that secrete the seed is found. The efferent duct
is narrow and very tortuous, and afterwards passes into a wider
canal with internal folds, into which also an elongated ^and
{prostata 9) situated laterally efiuses its secretion. This canal tenni-
nates in a muscular purse with thin walls, in which lie whitish cj-
lindrical bodies, about half an inch, and sometimes much more in
length, and which move when they are moistened long after the
death of the animal : which motion Nerdhax has fully described \
NsEOHAM compares these BinguUtr little machines^ as he names the
organs, with the Spermaioxoa of other animals, {rom which however ihej
differ in their siie and their composite stnicture. This is not exactly the
same in the various genera, yet it agrees in the chief point that in the
cylindrical body an elongated sac with thin walls and filled with a white
fluid (iperma) is situated, and extends to one of its extremities, whilst it
the other end there is a filament, turned spirally, and contained in a long
thin tube, which, after a pear-shaped expansion, is attached by a short
pedicle to the elongated sac at the other extremity. On contact with water
the cylindrical body is opened, probably by endosmose, and this occasioiH
the expulsion of the spiral organ which draws out with it the attached sac
Within the sac is contained a white sperma with cylindrical apermatotoa
terminating in a long fine filament. The Needkamian bodies are thus not
spermatozoa, but they contain tpermaUuoa; they are teed^nachimsg, cap$uke
§eminit or iperfnaiophore$, as Milkb Edwards has named them*.
The male Cephalopods hare a short, very thick, conical and per-
forate penis, which is situated at the side of the rectum at the same
part where the extremity of the oviduct lies in the females attached
1 NouvtUm dicouvertm faitet awe U Mieroteope, par T. Nskdham, tradttitet dc
VAnghU, Leide, 1747, 8vo, pp. 44—^7- London, 1745. Henoe, also, these oigans
aie named Needkamian bodies, although they had been preyiously obserred by
SWAMMBBDAX (Bijhd der Nat, Tab. Lll. figs. 6, 7). Some regard these litUe cylinders
as real animals ; see G. C. Cabus Needkamia eacpuUoria eepice ofieinalis, heeehreAen
und abgebiidH: iVbix Aei, Acad, Can. Leop, Carol, xxix. PI. i. 1839.
• Compare A. Kbohk, Fbobup's Neue Notiuii, xn. Bd. § 17— «3. October, 1839,
Philippi in Muillib's ArcAiv, 1839, § 310— 3"» Taf. xv. ; Pwaas ibid. § 98—100,
and MiLifB Kdwabds Aim. dee Sc, nctfur., «e S^rie, xvin. 184 1, pp. 3« 1—347, ^f^
many figures. In a well-preserved specimen of a male Nautilue I found the whole
frwrto Needhami {rwcula aeminalit) filled by a single cylindrical, long, and tortuooii
canal, within which was a fine spiral filament.
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MOLLUSCA* 821
in the gill-sac. Whether an introductioii of this organ into the
funnel of the female takes place, is not sufficiently cleared up.
[In certain genera of Cephalopods there is a very remarkable ac-
cessory organ of propagation in the males, which we now proceed to
notice. Amongst the doubtful genera of Entozoa (p. 188) ffectoco-
tylua is recorded. It was first noticed by Delle Chiaje in Argo-
natUa\ and by him referred to the genus Trichoc^fuUua by the
name of Trichocephaltia cLcetaibvlaria, Afterwards what was sup-
posed to be another species of the same genus was detected in Octo-
pvs granulastia Lam. by Laubillabd, and described by Cuyier*.
This presented more than 100 suckers, whereas that of Delle
Chiaje had only 35, whence it was named by Cuyieb Hectoootyle
from cKaroV. KoELLiKER*, from his examinations of the Hectocotylus
of Argonofuta, and of another discovered by Y^rant in Tremoetopua
violacetLSf declared it to be the male indiyidual of these animals
respectiyely, and not a parasite. This opinion was afterwards
shewn to be unfounded, for Yeslany* found in several specimens of
a Cephalopod, called by him Octopus Ca/rena (male Oct, grcmvlostit
Lam.), that the 3rd arm on the right side was longer and thicker
than the rest, and had a bladder at its extremity, whilst Philippi,
in a specimen examined by him, saw the same arm &11 off on being
touched, and found it to resemble exactly the Hectocotylua described
by CtTViEB.
At an early period the Hectoeotylua arm is developed in its future
position in a pediculated vesicle in which it is rolled up. The
vesicle alone contains chromatophores*. As the development pro-
ceeds, the containing vesicle or bladder shews a fissure on that side
which is towards the mouth of the Cephalopod fix)m which the hec-
toeotyhu emerges ; but since the latter is attached throughout a con-
siderable portion of its length to the inner surface of the otl^er side of
the bladder, this last is turned inside out when the hectocotylus has
escaped, and continues to be attached to it on its dorsal surfiice. The
structure of the hectocotylus is the same as that already described of
the other arms, except that in the central canal is an elongated mus-
cular pouch, closed at the near end and terminating in a fine tube,
and that there is a filament at the end of the hectocotyltis (which has
^ Memorie, n. p. 22^, PL xvi. fig. 1.
* Ann, det Sc, nal, xvm. 1819, pp. 147—156.
^ Ann, of Natural History, 1845.
* MoUtuqwi mediterranSen$, ihre Partie. G^nes, 1847 — 1^54» >• 34 '^^ ^^^> ^^ 4i-
* See p. 834.
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822 CLASS XIII.
been developed in its own proper vesicle within the larger bladder, and
of which the remains continue attached at the extremity of the hee-
tocotylus afler it has allowed the filament to escape). The filament is
tubular^ its cavity is continuous with that of the hedoootyhu^ and is
open at its extremity. The tube of the elongated pouch runs in the
central cavity of the heotocotylus and of its continuaticHi^ the fila-
menty and opens at the extremity of the latter externally. The
pouch or sac is found, when the hectocatyltu is fully developed, to
contain a spemuUophore, and it was the presence of ^^ermatozoa
and of an organ apparently for the secretion of them in the interic^-
of the hectocotylusy when found attached to the respiratory cavity of
the ArgonaiUa, that induced Koellikeb to suppose the ffectoeoiylus
to be the male of the ArgonatUa, H. Mueller found this Hedoco-
tylu8 in its state of imperfect development on that male, which is
much smaller than the female, and has no shells The hectoootylus
arm is the second of the left side in A, Argo, the third of the rig^t
side in Octopus granuhstis, {TremotAopus Cwrena Yer.) whilst the
hectoootylus only, and not the male of Tremoctopus violaeeus Dqj£
Ghiajb, has hitherto been discovered.
The sexual organs of the hectoootyliferous males are constructed
on exactly the same plan as those of the males o£ ordinary Cephalo-
pods. The vm de/erena opens at last into the respiratory sac, so
that the idea of a communication between it and the sac of the hec-
toootylus arm, entertained by some, is untenable. How then is the
spennatophore transferred to this sac ) Yogt' thinks that the spot-
ted pocket (the bladder in which the hectoeoiyhis was originally de-
veloped, and which is now turned inside out^ with a fissure
through which the arm has passed) serves as a receptacle for the
spermatophore, which has been transferred to it by the long filament
of t]|e hectoootylus embracing it ; at the same time he denies the ex-
istence of the muscular sac in the interior. Leuckakt', however^
has established the existence of this last, and has found a communi-
cation between the spotted bladder and the side of the muscular sac
attached to it, namely a canal by which they oommunicata It is not
1 See H. MuELLEB Ueber das Mdnnchen von Argonauta Argo und die Reetoeotylai,
SiEBOLD u. KoLLiKSB*8 ZeUtchr. fir wi$9ensck. Zool TV, 185a, pp. i — 35, PI. i. ibid,
pp. 34^—359-
* VfRANT et VooT Swr la not, de» HecHoeoiyUs, Ann. du Se. not. T. xvn. pp. 148,
191, PL 6—9, at p. 155.
* JjEUOHAXt Die HwtocGfyU€ von Oetoptu Oarentt, Zoel, ThUitrmtck, Hell m. 6ies-
sen, 1854, pp. 91—109, Tab. a, at p. 103.
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M0LLU8CA. 823
impossible that the spermatophore may be transferred in some casesy
in the vaj supposed bj Y ogt, but not always. For the spermatophore
has been found in the hectocatf/lu8y whilst the filament was yet in-
closed in its proper sac. In this case the other arms may have
effected ihe transfer, as Leuckart conjecturea After it has been
brought thus far, the propulsive force of the spiral portion of the
spermatophore must be supposed to come into play in order to begin
the passage through the tube of communication with the muscular
pouch of the heetocotylus, whose walls are also muscular. The fila-
ment at the end of the hectocotylui has been very generally supposed
to act as a peni& H. Muellbb found as many as six of them
within the capsule of the ovary in Argonawta Clio, that had been
broken off', and two others in one of the oviducts, so that no fewer
than eight males would seem to have fecundated this individual.
It is presumed that when the kectocUyluB arm has been cast ofi^
it may be successively reproduced.]
The eggs are laid in heaps, or are connected with each other in
cluster's by pedicles and an adhesive substance. In this respect
great variety prevails in the different genera. In Loligo, many eggs
are united in strings of a gelatinous substance; in Sqna they are
very large and surrounded by a homy dark envelop, which is pro-
duced into a pedicle at one extremity, and by which the eggs are
attached to each other, or to foreign bodies.
In the development of the embryo, when the egg has been laid,
grooves are formed in that part of the yolk where was previously
the germinal vesicle, and which do not extend over the whole
yolk. Here the embryo afterwards appears as a disciform germ, at
first flat ahd round, in which, at an early period, different inequali-
ties indicate the first existence of the mantle, of the eyes, and of the
funnel, formed at first of two separate halves. This germ-disc be-
gins to swell graduaUy in the center, extends itself constantly further
towai-ds the circumference, and finally surrounds the entire yolk.
In this way a part of the yolk-sac between the arms at the ventral
surface is included in the mantle, so that afterwards an internal as
well as an external yolk-sac is found, the two being connected by a
narrow pedicle*.
1 [Op. cU. p. 354.]
> Fonnerly it was supposed that the yolk-sac is connected ^nth the mouth by means
of this pedicle (Cavoliki); Guvhb presumed a connexion with the oesophagus j see
his memoir aur let ornfa de Seiehea, Nouv, Aim. du Mua. I. 1833, pp. 153 — 160, PI. 8.
The chief work that we now possess on the development of these animals and from
which we have borrowed what has been announced, is A. Kobllikxr BrUvfichelunffa-
geachicJUe der Cqahahpodm, Zurich, 1843, 4to.
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834 cuiss XIII.
The nervous system is kigelj developed in the Cephalopoda, and
in most of the genera the central masB for the head (the cerebral
mass) is endoeed in the cartilaginoiis ring, which we have already
noticed. Beneath the oesophagos a considerable nerve-mass is situ-
atedy which in Xauiilus is divided into two transverse bands, one in
front of the other. From the anterior portion of tlus central mass
(or from the first of the two half-rings, situated nnder the cesopha-
gns in XauiilHs) the nervons trunks for the arms arise, ot for the
tentacles surrounding the head. In OetopuSj Sepia, and Loligoj
these anterior portions form on eadi side a large, flat gan^<xi, from
the anterior margin of which the brachial nerves of that aide radiate
{gamglumM en paUe d'oie CuYiKa). From the posterior portion arise
at the side thick nervous stems for the mantle, which in Lcli^j
Sfpia, Ckiopus, Ac. terminate in two lai^ ganglia, from which the
nerves radiate at acute angles; in yautiltu these lateral gangiia
wUiiaia are wanting. In addition there arise from the middle of
the posterior margin of this portion two nervous trunks^ at first laid
dose together, from which the nerves for the viscera arise. 'From
the portion situated above the cesophagus nerves arise for the mus-
cular mass that surrounds the jaws, or these come from the gang^on
of the sympathetic system placed above the oesophagus (gangiian
pkaryngtum) as in Sfpia and Loiigo, From the lateral oommiasnres
of this portion, by which it is connected with the central mass situ-
ated beneath the oesophagus, the two remarkable optic n^ves arise,
which terminate in kidney-shaped ganglia; lower down arise the
two short auditory nerves which penetrate into the cartilaginous
ring of the head. The sympathetic nervous system has a ganglimi
(gangfioH labiale) seated under the oesophagus, and one on the sto-
mach formed by the union of two nervous stems.
On the organs of sense in this order, what we have stated above
(pp^ 7G6 — 7i>d) may be referred ta The skin of the Cephalopoda is
distinguished by a diange of colour exhibited during the life of the
animal, and depending upon a peculiar stratum of saccules fiUed
with pigment-granules {t^romaiophores of Saxoiotanni and Wag-
xer), which are connected by a delicate, elastic^ membranous tissua
These organs lie immediatdy under the cuticle, and by alternate
contraction of the vesides, containing red, blue, or yellow colouring
matter, a lively play of colour arises in definite situations, espedaUy
in Loiigo and Argonauia^.
> CompMv R. Wa65KB in OKKir*8 Im, 1835, pp. 159—161, Wuoxaks mid Ebich-
SOn's Arrkirf. Xfttmryetrk, 1841, s. 55 — 38.
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MOLLUSCA, 825
On this order, which many writers separate as a distinot dass from the
rest of the Molluscs, compare,
CuYiXB Mimovre ntr CephalopodeB et §ur leuranatomie, MoUuaq,,AfSm, i.
with 4 plates.
R. OwBN Cephalopoda in Todd*b Oydopcedia, i. pp. 517 — 563.
F^UBSAO et D'Orbiont Monographie des Cephalopodet erpptodibrancka.
Paris, 1834 — 1843. i81iynusoD8.
A. Tetrabranchtata.
Family XII. Nauttlacea. Branchidd four. Funnel cloven
beneath, with two lamellss covering each other obliquely. Tenta-
cles numerous, contractile, vaginate. Shell external revolute, spiral,
poljrthalamous, hard; with septa perforated in the middle; last
chamber ample, including the animal, with a membranous tube
(siphon) produced from the posterior part of the abdominal sac
through the foramina of the septa into the other chambers.
Nautilus L. (in part), Lam.
Sp. NauHlus PompHiut L., Rumph. Amb. JRariieitk. Tab. xvii. figs. ▲, 0,
Blainv. Malacol. PI. iv. fig. 8 ; Gu^bin Iconogr., MoUvsq. P. k fig. 7 ;
the pearly nautUut; the shell attains a size of more than J foot ; on the
inside it is whitish with a pearly lustre, on the outside milk-white, with
orange or red-brown stripes. This species, the only one of the seventeen
species recorded by Liknjkus that has remained in the genus Nautilus, is
found in the Indian Ocean, the Islands of the Moluccan Archipelago, &a
Nautilut wnUnUcatua Lam., in the shell of which the last wreath does not
cover that which precedes it, is in other respects comformable to the above.
Notwithstanding the shell is not altogether rare, the animal of the pearly
Nautilus was, until within the last few years, known only irom a very im-
perfect figure of RuiCPHius. Owen was the first who gave a good descrip-
tion and a careful anatomy of this interesting animal {Memoir on the pearly
Nautilus, with iUuttrationt of its external form and internal structure.
London, 183), 4to). Besides the characters already given above, it is dis-
tinguished from the rest of the Gephalopods now living by the absence of
the Ink-sac. The head is covered by a large membranous hood, which
represents exactly the circumference of the aperture of the shell, and arises
from the expansion and coalescence of the cases of the uppermost pair of
tentacles. The pediculated eye is partly covered by the over-hanging mai^n
of the hood. Beside the 19 or 90 larger external tentacles, the mouth is
surrounded by two pairs of flat, finger-shaped lappets, each of which encloses
19 — 16 smaller tentacles, but in other respects similar in structure to the
hunger. If a male specimen observed by me be not a monstrosity, then it
may be concluded that in these organs a sexual difference is presented ;
(compare Tijdsschrtfi voor de Wit-en Natuurk. Welenschappen, published by
the Koninld. Nederl. Instituul, i. 1848, biz. 67—73, ^1« i- ^^- '—3)» Trans.
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826 CLASS XIII.
</ ZoU. 8oe. lY. 1 85 1, pp. 76, tf. [From letten of Yak dkb HoBrnr,
July aad Sept. 1855, it appears that the above was no aboormity but the
usual oonfoimation of the males. " I have had the opportunity," he adds,
" of examining recently two other male specimens, and hope to have time
to publish this year or the following, some new observations on the struc-
ture .... The male differs from the female by the absence in the mantle of
the lamellated glandular apparatus (Owen's Mem, p. 43, PL Tin. fig. 10),
and by a different number of the digitations. (These are less nomenms in
the female.) A chief difference, however, is the presence of the great
conoid body at the left side (see my Con^nbuUong, p. 47, PL 7, figs. 10^ 11).
This singular body has at the extremity and on the outside a large disc
perforated by the orifices of numerous crypts. As the spermatophores
(Kbbdhak'8 mocAtMt) after their passage through the penis are iadoaed
in a bag formed of two coats, of a brown colour, and of neariy half an
inch in diameter, I believe that the glandular apparatus may secrete this
envelop, and that consequently it is a physiological analogon of the glan-
dular apparatus described by Owxv in the mantle of the female. The fold
connecting the labial processes at the inferior nde above the funnel is in
the male of a different conformation, (see CotUrib, pp. 16, 37). The fiae
folds of the actemal labial processes are totally wanting, and instead of the
so-called olfactory oi^gan at the commissure of the internal labial prooeasn,
there is the cushion-like part (with 8 — 1 1 digitations), (ConJtrQt, PL 8, ftg. 9).
As to the internal genital organs of the male^ a large gland (teiCu), whidi
in .bulk surpasses all the otiier organs of the body, except the liver. Is
situated ezactiy where the ovaiy is in the female ; another smaller, flat
gland, more at the fore part, seems to secrete the BpenMOftpkmu ; a bag
with an imperfect internal septum receives these spermotopAoree and brings
them to the conical penis, which is situated not exactly, but nearly, as is
the vulva (more in the mid-plane).**]
The genus NavUlus occurs also fossQ in secondary and tertiary formations.
It is the only genus still extant of a very numerous division of the Cepha-
lopods, which lived in the seas of a former world and of which the remains
are met with in mountun strata, espedaUy in the older seoondaiy fonna-
tions. Here belong the Amvumites.
Fossil genera related to Nautilus :
Clymenia Muenster.
Comp. Mim. mr lu Clyminea H le$ QoniaUtu du ealeairt de tramiUm
du Pichidgdtirgt par U Compte De Mukksteb, Ann, da Se. nai,, le S^e;, n.
1834, Zool. pp. 65—78, PL I.— L. Voif BuoH Ueber GaniaHien und Cly-
menien, in Sehienen, Phytikal. Abhandl, der BerL Akad, der Wi$$endi. 1838
(section of genus NautUut according to V. Buoh).
Lituitea Bretn, Mont.
Sp. Lituitet eonwlvani SohiiOTR., Brokit IMom geogn. Tab. 1. 1%, 3.
Campylites Desu., Oyrtocera QoLDFUSS.
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M0LLU8CA. 827
Orthoceratites Breyn. Shell elongato-conical, straight, poly-
thalamous, with septa perforated by siphon middle or subcentral.
(Straight, not spiral NatdiU.)
8p. Orthoceratites regularis SOHLOTH., Bbonn Loh, geogn. Tab. I. fig. lo, in
ThfyMckieferhdh {Werdock shale f)
Gen. Actinocerat Bbonn does not, according to Qubnstedt, differ finom
Orthoceratites.
Appendix to the Nautilacea. Family of Ammanitacea. Shell
poljthalamoiis, with siphon external, marginal. Septa at the margin
lobate, sinuate.
Comu Atnmonis, These petri&ctions are of various, some of very re-
markable, size. If the animal was placed, as we may conclude from analogy,
within the shell as Navtdus is, the siphon would lie on the ventral side
towards the margin of the funnel, and the expression Sipho dorsalis, used
by Paleontologists, is consequently incorrect. — Ammonitet are met with in
great numbers, especially in the transition and older secondary formations.
In the lowest chalk-strata they are still numerous, but afterwards decrease
rapidly and after the chalk-periods are met with no more on our earth.
Compare G. De Haax Spedmeih inaug. exhibens Monographiam Ammo-
niteorum et Ooniatiteorum, L. B. 1825, 8yo; L. Db Buoh Note tur les
Ammonites, Arm, des Sc, not. zvu. 1829, pp. 267 — 275 ; Sur la didribtUion
des Ammonites en famiUes, Ann. des Sc not. zvin. 1829, pp. 417 — 426;
Ueber Ammoniten, fiber ihre Sonderung in Famtlien, &c., PhysUc, Abhandl.
der Berliner Akad. d. Wissensch,, aus dem Jahre 1830; l/d>er Ceratiten,
ibid., a. d. J. 1848; F. A. Quenstei>t IHe Cqthalopoden, Nebst einem
AUas von 36 Tafeln. Tttbingen, 1849, 8vo.
Ammonites Breyn. Shell spiral, discoi'dal.
Sub-genera: GaniatUes De Haan, CercdUes ejusd. and some
others seem only to form sections, with species passing from one
section into another.
Sp. Ammonites nodosus Bbuo., Ceratites nodosus Db Haak, Bbokx Ldh,
geogn. Tab. xi. fig. 20; V. BuoH Ueb. Ceratiten, Tab. i. n. fig. i, from the
MuscheOcaUc, The species of the section of QoniatUes are found in the
transition formations.
Scaphitea PARKINSON.
Hamttes Sowerby. Shell incurved at the apex, produced into
a straight tube.
Sp. Hamites rotwndus Sowerby, Bbonn Leih, geogn. Tab. zzznT. fig. 9, from
the chalk-formations.
TurruUtes Montf., Lam., Twrriies Db Haan.
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828 CLASS XIII.
Baculites Lah. Shell straight, conical.
S^ Homlafet Fm^adi Lam., Faujab ds Sr Fokd Hiat. ttof. de la moiie de
Si Pierr€y PL 2t, figs, i, $. QuurariDT nnitee with this Baeulileg ameept
Lam. and BaeuiiU$ verfeftra/w Beoitk, Leik, geogn. Tkb. xxxm. fig. 6.
Th«M mkI BaeuiiUt neoeoMtfum D*Orb. are spedeB from the chalk-for
matioDB, BmnalUa mnutrimt QunremDT, Tkb. 21, fig. 15, ia hitherto the only
known speciee from the /iira-fornuition.
B. D!branchi<Ua,
In this division of Cephalopods, to which most of the species now
liTing belong, the funnel is dosed below. There is an organ for the
secretion of a black floid, which the aildmal can eject through, the liin-
nel, the ink-sac In the genos Octopus this ink-sac is endoeed within
the liver^ but in the rest is remote from it. The colouring matter of
some spedes is used as a paint, Sepia, Since some observations haTe
indicated the presence of an ink-sac in the fossil Belemnites^y it may
probably be conduded, with Owen, that these petrifiuitions are in-
ternal sheUs of Cephalopoda dibranehiataj which conduaion has^ in
&ct, been since established by the obeenration of petrified animals*.
Family XIII. Decacera s. Sepiacea, Arms ten, two longer
than the rest, round.
Beleninites Breyx, Lam. Shell fossil, conical, including inter-
nally a multilocular, shorter portion (alveolites).
Beifmniia or dart-tloikts ; thiu named on aooonnt of thdr dart-like form.
T^Mwe petrifibctionfl are found abundantly in the aeoondafy mountains of
the ooUte and chalk-periods. Compare, amongst others, Ducbotat DI
BXiAiimLLi Mfm. aur let BdemniieM, Paris, 1837, 4to; J. S. Miller
Obtrvaltuma on Bdemm,, Tnmsaet, of the OeoL Soe, of London, sec. Series,
n. 1839, pp. 43 — 67, PI. 6—9; Bronh Letk. geogn, pp. 403 — ^418, pp.
714—730.
Sptrula Lah. Animal with ten tentacles, two longer. Shell
placed in the posterior part of animal, thin, spiral, with wreaths
not contiguous. Partitions concave, perforated by siphon at the
inside.
1 A0A89U in V. LsoRHASD u. Bronn JVevei Jakrh. JUr Mineralogie, Cfeognome il
Oeol, 1835, 8. 163, BucKLAKD Oeologg and Mineralogy, London, 1836, 8to, {Bridge-
water TrtaUtes, vi.) p. 374, PI. 44', fig. 7, PL 44".
' Loiiuru on the comp. Anai. and Phgtiol. of invertebrate Animalt, p. 337, snd ed.
pp. 597—^3-
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MOLLUSCA. 829
Of thifl shell the animal is known from only imperfect figures, and some
specimens not entirely perfect. See E. J. Gray AnnaU of Nai, ffist. xv.
pp. 257 — 960, PI. zv. A mulilated specimen, investigated by Owen in
1848, has afforded, however, by the anatomical particulars of internal
structure the certainty that the animal resembles the Sepiacea or Deeactra,
There are two gills, two lateral or venous hearts, an ink-sac, a membranous
tubular siphon, kc, Sne B. Owen in Zocloffy of the Voycige of H. M, S.
Samarang, under the command of Oapt. Sir Edwabd Beeoheb, MoUvsca,
Part I. London, 1848, pp. 6 — 17, PL iv.
Sp. Spirtda Peronii Lah., Nautilus tpirula L. ; the shell has been several
times figm^, as in RuMFH. Amb. JlariteUk, Tab. 90, fig. i, D'Aboenv.
Conchyl, PL 5, fig. o, o, &c.
Sepia Lam. (species from genus Bepia L.). Body sacciform,
depressed, margined on each side by a narrow membrane or longi-
tudinal fin. A calcareous lamina elongate, dorsal, included in
mantle.
Seiche, DirOenfiach, Sea-caly Ink-fish, Cuttle-fish, The calcareous plate in
the back consists of many laminae. This part, known as os SepicB is used
by miniature-painters for polishing ivory plates and forms a component of
some tooth-powders.
Sp. Sepia officvndlis L., Bbakdt u. Ratzebubo Mediz, Zool. 11. Tab. 31,
fig. T ; Gu^iH Iconogr,, Moll, PL I. fig. 4, &c. Gabub has given coloured
figures from living animals, Nov. Act. Acad. Cos, L, C, Nat. Curios, xii.
1824, pp. 313— 3^^* Tab. 28—32.
Sepioteuthis Blainv,, Chondrosepia Leuck.
Loligo Lam. Body elongate, pinnate posteriorly on each side.
An elongate, narrow, homy, lamina at the back, covered by mantle.
Sp. Lcligo vuUfaris, Sepia loligo L., Blainv. Malacol. PL 3, fig. 2, Cabus 1. 1.
Tab. 29, fig. I, &c.
Ommcu^ephes D'Orr, OmmuUostrephea Lov^.
OnychoteiUhis Liohtenst. The longer or all the arms armed with
homy hooks.
Onychoteuthis Fabrieii Moelleb, Sepia loligo Fabb., Favma Grcenl. / —
LoUgo Banksii, LoUgo leptura, Loligo Smithii Leach in Tucket's Expedi-
Hon to the River Zaire.
Sqnola Leach.
Sp. Sepiola Jtondeletii, Sepia sepiola L., Blainvillb Malacol. PL n. fig. 3.
Comp. R. E. Gbant on the anatomy of Sepiola vulgaris Lbaoh, Ac.
Transact, of the Zool. Sor. i. 1833, pp. 77—86, PL 11.
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CLASS XIII.
Bessia Owen.
Loligapsia Lam., Perothis Eschscholtz, Bathke.
Laicarck hM described this geniu m huTiiig only ogiit ainiB ; the feog
tentaeula are often torn off. In common with Crtmehia this genus has no
membranous valye in the fonneL These animals are neariy transpsrat;
the eight arms are short ; the body is conical, posteriorly narrow and whh
two ronnd fins at the extremity. Compare Bathkx Peratkia a» nemei
QmvsderCepkalopoden, Mim. freaenUa d VAcad. impir.daScde Sfeten-
Ixmrg, n. 1835, PP- 149— '7^-
Cranchta Leach.
8p. Cfranchia 90ahra Leaoh in Tuokkt'b figwd. to the JUmt Zinre, p. 410.
The fins are situated quite at the end of the body and the dorsal sui&oe of
the mantle is grown fSiist to the head.
Family XIV. Octocera s. Octapoda. Arms eight, large, often
very long, surrounding the mouth in a circular row. Body sacci-
form, without fins. Funnel without valve.
Arffonauta L. Arms furnished with a double row of acetabula
(suckers), the two superior expanded into a membrane towards the
extremity. Shell thin, involute, external, unilocular, with spire
bicarinate.
This remarkable shelled animal excited long ago the admiratiosi of the
ancients ; see Puinus JTuC. Nat. ix. cap. 99. It was supposed that it made
use of its fin-shi^ed arms as a sail, and thus in still weather oould swim on
the surface of the sea; but the obeerTations of Rano (Gulbinr Maganm
de Zool. 1837^) have not confirmed this opinion; these arms lie expanded
along the outside of the shell and serve to fasten the animal in its sheD,
which is not attached to it by any muscles. Many naturalists (RAFonssQUi,
LSAOH and others) thought, that this animal like the homit^^aibb {Poffunu)
lived parasitically in a borrowed shell, and that the shell of Argtmaiita
belonged to some unknown species of mollusc. Apoordingly this genus of
Odopoda was named as a new genus Oeythoi, Although this animal can
readily move from its shell, just like some Pteropoda, this opinion is now,
however, sufficiently refuted. For the discrepant form of the male indivi-
duids which have no sheU see above, p. Sic — 815.
On the anatomy of Arffonauia compare Foli Teat. utr. Sicil. Tom. ni.
and Van Bbvsdxk Nouv. M4m. de VAeadSmie royaU detSe.de BruxeOee^
Tom. XI. 1833.
^ PoLI adopts this notion of the ancients, and he has even figfured the animal in
accordance with it (TVufac. tUr. Sicil. ni. Tab. 40), but says that he has never himself
seen the sails of the Ai*fronanla,
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MOLLUSCA. 831
Sp. Argonauia Argo L., Poli L 1. Tab. 40 — 43, Blainv. MdUbc, PL i. fig. i,
PI. I. bis; Rakq, GuiBiN Magaa. de Zool. 1837, MoU. PI. 86—88, from
the Medit. Sea; — Argonauia tvhercuUUa Lah., Rumph. Amb» JRariteitk,
Tab. i8y figs. I, 4, D'Aboeny. Conchyl, PI. 5, fig. o, from the Indian
Ooean, &c.
Nate. — Here would seem to be the place of BeUerophon Mont.
fossil, from the oldest strata.
Octopus Lam. Body naked. Two homy lamellse in the back,
inclosed in mantle. [Polypus of the ancients.)
Amongst these naked eight-armed Gephalopods there were some species
of which, as in the last, females alone were known, and of these it is that
the heetocotyltu of the male (now in some instances known) has been found
attached in the sac of the mantle or in the funnel, as Octopu* gramdoau$
Lam. and Tremoctopus violaceui Dxllb Chiaje. See above, p. 820.
Octopus auct With a double row of suckers in the arms.
Sp. Octopus vulgaris, Sepia Octopus L., Blaikv. Malacol. PI. 2, fig. i, Cuv.
JZ. Ant., id. iU., MoU. PL I.
Tremoctopus Dells Chiaje.
Sp. Tremoctopus violaceus Dellb Chiaje, Memorie, Tab. 70, 1830.
Eledone Leach {Ixclmvn Abistot.). With a single row of aceta-
bula in arms.
Sp. Octopus moschatus Lam., BLAimr. Malacol, PI. a, fig. a, Cuv. X. Ant,,
id. iUustr., MoU. PL 2, fig. i.
Oirroteuthis Eschbicht.
Sp. Oirroteuthis MuUeri, Ebchbioht Nov. Act. Acad. Ocbs. Carol. N. Cur.
zym. 2, 1838, pp. 637 — 634, Tab. 46 — ^48. A broad membrane connects
the arms almost as far as the extremity. This species is a native of the
coasts of Greenland.
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EXPLANATION OF PLATES.
Plate L Inpusories, figs. 1 — 6; Polyps, figs. 7 — 12.
Fig. 1. Vibrio lineola, p. 45, after Ehbenbebg InfusuyM-ihierchen^ TaU
V. ^. 4, (very higldy magnified, nat. size ^^ line).
Fig. 2. Atho^ dijluem, p. 46, after Ehrenbeeo, Tab. vm. fig. 12,
(magnified, nat. size ^ line).
Fig. 3. Siderdina calcttrap(ndeSf p. 49, magnified about 8 times, after a
specimen from Mont St Pierre.
Fig. 4. C}il(Miiidomona8 pulmscultM, p. 51, after Ehbenbebo, Tab. iil
fig. X. B, (highly magnified, nat size ^ line).
Fig. 5. Eerona mytUm, p. 55, after Ehrenbbrg, Tab. xu. fig. ix.
(magnified, nat. size J line).
Fig. 6. Varticella nebuli/era, p. 58, after Ehbenbebo, Tab. xxv. fig. 1,
(magnified, nat. size of the bell ^ line).
Fig. 7. Hydra griaea L., p. 73, from nature, nat. size ; a, a single polyp,
extended; 6, the like retracted; c, a polyp with two young buds
t,tt; the last still very incomplete.
Pig. 8. Clava parasitica, Hydra muUicomis, p. 75, aft»r Fobsk. Icon,
rer. natvflr. Tab. xxvi. ^g, 6, b.
Fig. 9. Goryne Qjebts,, Syncoryne ramosa Ehbekb. p. 75, highly magni-
fied, after LovfN, Wibom. Archiv, 1837, Tab. vl ^g. 19 6, b bell-
shaped excrescences with ^gs ; comp. p. 70.
Fig. 10. Campamdaria {gda4,ino8a\ p. 76, in the embryonal state,
magnified; b, in a more developed condition, in which it is
about to quit the ovary; after Yan Beneden M^m, de VAcad,
de BruoB, Tom. xvil PL n. figs. 9, 18.
Fig. 11. A, B, c, Tvhipora musica L., p. 82, aftier Fbeycinbt Voyage de
rUra/nie, ZooL PI. 88. a, development of a young polyp c; a,
empty tube, firee of its polyp ; by more developed polyp, opened
longitudinally d, e; b, upper extremity of a polyp with the 8
tentacles and the mouth in the middle ; c, longitudinal section
of a tube ; here the animal is seen, and below it the eggs within
the tube, attached to long filaments.
VOL. 1. 53
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834 EXPLANATION OF PLATES.
Plate II, Polyps.
Fig. 1. Anthdia glauca^ p. 77.
Fig. 2. Xenia umbellata, p. 77, both after Saviont DeaeripL de VEg^,
Polypes, PL i.
Fig. 3. CoraUium rubrumy p. 83, magnified, after Milke Edwabds, Cu-
TIER JR. Ani,, Zoophytes, PL 80, fig. 1. a, a, a, three polype, of
whioh the two lowest are retracted within the bark ; 6, 6, bark ;
e, stony axis.
Hg. 4. Mopsea, Isis eUmgaia, Espeb, p. 83, after a spedmen in the
National Museum of Nat Hist at Leyden.
Fig. 5. Caryophyllia ramea Lail, p. 86, after Milnb Edwabds, Cuyieb
R Ani., Zoopk,, PL 83, fig. 1.
Fig. 6. Actinia corioMa Cuv., p. 91, after Rapp Polypen und AcHnietif
Ta£ l 3, 4. a, in the oontracted state ; b, expanded.
Fig. 7. Halodactylus diaphanus Fabbe, p. 96. a, the gelatinous polj-
pary of nat size, after Yan Beneden Bryozoaires; b, a polyp
magnified 80 times, after Fabbe Phil Trans. 1837, PL 26,
fig. 7.
Plate IIL Polyps, fig. 1 ; Sea Nettles, figs. 2 — 8.
Fig. 1. PlymaUUa criHata Lam., p. 96, after Tbehblet Pdf^. PL x.
figs. 8, 9. Fig. A, natur. size; fig. b, three polyps magnified, of
which one is retracted within its cell, and another still young
and undeveloped.
Fig. 2. VdUUa seaphidia Pebon, p. 110, after PiaoK Voyage aux ierret
Ausir. PL xxz. fig. 6.
Yig. 3. Porpikt wmbella Eschsch., p. Ill, aft«r Pebon, PL xtxt. figs. 6
and 6 a; A, fixim the dorsal surfiuse, b, from below.
Fig. 4. Diphyes eampanulifera Eschsoh., p. 118, after Qvoy and Gai-
MABD Ann. des Sc. not. x. 1827, PL l figs. 1, 3. a, the animal
consisting of two pieces, nat size b.
Fig. 5. Cydippe pileus Eschsch., p. 121, after Milne Edwabdb in Cuyieb
B. Ani., Zooph., PL 56, % 2.
Fig. 6. Cassiopea borbonica Delle Ohiaje, p. 123, after the figure of
Dellb Ohl^ue reduced in Gui^bin's Iconographie.
Fig. 7. ^quorea violacea Milne Edwabds, p. 125, after the figure in tke
Ann. des. Sc noL, 2e S^e, Tom. xyl PL l fig. 1.
Fig. 8. Nettle-organs (pp. 99, 100), from Pelagia nocHluca (p 124),
after Wagnsb Icoti. Zootofn. Tab. xxxiii. figs. x. xl a, when
slightly magnified the round Yesides amongst the pigment-
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EXPLANATION OF PLATES. 835
granules are seen ; a, a, filaments, b, cell a more highly magni-
fiedy with the filament everted; c, filament lying in the cell
rolled in a spiral
Plate IY. £chinodebh&
All the figures of this plate are taken from specimens in the B^ka-
Museum of Nat History at Leyden.
Fig. 1. Ophidepis imbricata'NLxjmjj. et Tbosch., p. 147, 1^ the nat size,
seen from the under surfaoa To save space, the basal pieces
alone of four of the arms are delineated ; a, a, fissures near the
arms.
Fig. 2. Under sur&ce of a ray from a Japanese species of Astropecteriy
p, 148, nat. size. Two rows of terUaeula are seen here.
Fig. 3. Greasier hivlcus Muell. et Tbosch., p. 149, seen from above, of
nat. size; the left side is in great part removed, a, anus; 5,
Madrepore-plate (verruca dorst); compare p. 148.
Fig. 4. A ray of the same species opened a, Sand-canal or stone
canai; see p. 148.
Fig. 5. Under sur&ce of part of a ray of Asterias (Asteracanthum)
rubens, p. 149, nat size. Four rows of tentacles are seen;
see p. 149.
Fig. 6. Spaiangus purpureuSy p. 152, seen from above, reduced one half
There are four apertures of the genital organs (avcma and testes)
and four circumscribed amlmUicra,
Fig. 7. Oidaris imperialis Lam., p. 156, after a specimen from the Bed
Sea. The spines have been removed in order that the form of
the shell and the tubercles on which these spines are implanted
might be shewn. There are five sexual apertures round the
vent, on the same number of small pentangular plates of which
the hindmost is the largest : between them are the five eye-
plates. The am6««2aora here run quite through. Compare pp.150,
151.
Fig. 8. A spine of Cidariies glandifenis, or so-named pierre jtuia^ique,
fossil from the Jura formation.
Plate V. Fig. 1, Bchinodbrms ; figs. 2 — 7, Intestinal Wobms ;
figs. 8, 9, RonFEBa
Fig. 1. Sipunculus nudus, p. 160, after Gbube in Mueller's Archw,
Fig. 2. Ccenurus cerebralis, p. 182, after Bremseb Icones helminth. A
part of the common bladder, with two worms highly magnified.
Fig. 3. Gysticercus lanfficoUis, p. 182, magnified ; after the same, ibid,
53—2
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836 EXPLANATION OP PLATES,
Fig. 4. Tamia pliccOa, pp. 180, 181, the head and anteri(»- part of the
bodj nat size, after a specimen in the Rijks-Muaetun, Leyden.
Fig. 5. Echinorhynchua gigas, p. 184 ; A, nat. size, B, head of the same
species magnified. After Bremser Icones hdmijUhtan,
Fig. 6. Triatoma grantUaium, p. 186, magnified two diameters ; from a
specimen in the Bijka-Miueum, Lejden.
Fig. 7. Aacaris l^9U}pUra, p. 190, nat size, a, head, b, tail, both magni-
fied ; after Brehsbb Icariea helmirUhum.
Figs. 8 and 9, from Ehrenbbbg's Injuai4ma-thierchen. Fig. 8, Furcur
laria gracilia, p. 204 ; fig. 9, FkUodina megcdo^richa, p. 205 ;
both highly magnified.
Plate VI. Ringed Worms, Insects.
With the exception of fig. 3 0, all the figures are fix>m nature and of the
natural size unless the contrary be expressed.
Fig. 1. Gordiua aquaiiciia, p. 194.
Fig. 2. Planaria torva, p. 223, magnified two diam. b, head with tvo
eyes.
Fig. 3. Hirvdo fnedicincUia, p 226. b, ring^ fix>m the middle of the
body on the ventral sur&oe, magnified two diamet^^ c, the
anterior part of the head, from the inferior surfiuse with the
margin that surrounds the mouth. Three fissures are seen firom
which the jaws proceed. This magnified figure is copied fcom
that of Brandt u. Ratzeburg Mediz. Zool, n. Ta£ xxvul
fig. 10.
Fig. 4. A, Serpula bteomia Saviont, p. 236. b, the anterior part of the
inferior surface shewing the opercfUum.
Fig. 5, JTermelUiy p. 236. A new species fr:om the Cape of Good
Hope.
Fig. 6. Lj/caria, p. 241. The anterior part with proboada exserted.
A new species fit)m the Cape of Good Hope.
Fig. 7. PolyodmUea maxUloaa^ p. 245. a, the anterior portion with the
proboada exserted, b, the proboada on the inferior sur&oe with
the four jaws.
Fig. 8. Olomeria owtlia, Sphoeropcsua inaignia Brandt, p. 292. a, the
animal roUed together, b, the head and anterior rings from the
side, and c, fix)m before ; d, antenna ; o, first pair of feet ; h, two
rings frx)m the middle of the body with fo\ir pairs of feet
(d— H magnified).
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EXPLANATION OF PLATES. 837
Plate VIL Insects.
With the exception of figs. 4 and 5, all the figures are fix»m nature.
Fig. 1. LUkobma forficatus, p. 294. Natural size.
Fig. 2. Oral organs^ head and antennae of a large Scolopendra, slightly
magnified, a, head; b, antenna; a a, upper jaws; & h, under
jaws; c, under lip; d', first; d'\ second; rf'", third pair of feet.
Fig. 3. Oral organs of GryUotcUpa vulgaris, magnified, illustrating p. 249.
A, upper lip; B, upper jaws; c, under jaws; c'c', palps; (/V,
galea or helm of the lower jaws (internal palps); D, under
lip with its palps d d.
Figs. 4, 5. Head, spiral tongue and other oral organs of Noctua poce,
magnified ; after SAViaNY Memaires, i. PL in. ^g, 1. Compare
p. 250.
Fig. 4. A, basal piece of antenna, a a, compound eyes; bh,
upper jaws; </, under jaws (spiral tongue) ; rf' d\ attachment of
the palps of the under lip that have been removed.
Fig. 5, The under lip c, with its palps </ </, divested on the
left side of the hairs with which they are beset.
Fig. 6. Intestinal canal of GryUotaXpa vulgaris, natural size, to illustrate
pp.253 — 256; a, oesophagus; b, crop or proventriciUus; c, muscular
stomach, with two blind appendages near its inferior orifice ; d, e,
stomach, or duodenum, according to M. de Serbes; g, h, intes-
tinal canal, slightly unrolled, as in 6 ; ^ the short and numerous
urinary vessels, which terminate by a common efferent duct at g.
Fig. 7. The two secretory tubes of the web in Cossus ligniperda, natural
size, see p. 392.
Fig. 8. A Grylhtalpa vulgaris not yet full grown, shewing the air-slits
{stigmata) 1 — 10, see p. 261. This figure may serve at the same
time as an example of imperfect metamorphosis, see p. 273.
Fig. 9. A portion of a stem of the air-tubes with three branches from
a large Scolopendra, see p. 259.
Fig. 10. Internal sexual organs of male GryUotaXpa vulgaris, magnified
two diameters, to illustrate pp. 267 — 269 ; a a, testes; b b, efferent
vessels; c c, prostate (blind tubes, pp. 268, 269); </, such a tube un-
rolled; d, base o£ penis, covered on the upper surface with small
blind vascules like vUli (Cowprr's glands ?).
Fig. 11. Internal sexual organs of female of the same species; a a,
ovaries; bb, oviducts; c, blind sac (recepiaculum seminis), of
which the very fine duct c' terminates in the va^gina. See pp. 265,
266.
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838 EXPLAKATI09 OF PLATES.
Fig. 12. NenrooB system of GrydoUdpa ru^j^aHt magnified two <
see p. 276 ; 1, first nerre-gui^oii with the optie ncrres and
thoae of the anteimjSy bdiind it a rin^ thnni^ wkidi the
oesophagiis passes ; 2 — 10, the saooeeding nerre-gang^ik
Fig. 13. Head and firat piece of thorax {prcdtorax, eondd) c^GryOatalpa
vulgaris seen fi*om aboTe, sUghtlj magnified; A, A, oomponnd
eyes; a, a, simple ejes^ oeeUi or stemmaia. See p. 349.
Fig. 14. A small piece of the cornea of a compoond eye firam Spkmx
airapoi, with the hexangnlar /aceUe$^ magnified 200 diametos;
see p. 279.
Fig. 1^. A piece of the under surfiioe of an upper wing from Nym^fUtii
wrticcBf p. 411, magnified more than 40 diameters, and drawn by
means of incident light. The snr&ce divested of scales then
appears black ; the places for the attachment of the scales are
seen as white pointsi Comp. p. 390.
Fig. 16. Scales firom the upper surface of the upper wing of the sune
insect, magnified about 75 diameters, and drawn by means of
transmitted light under a microscope of Obebhaubeb, p. 390.
Plate VIIL Insects.
Fig. 1. Scales of MachiUs mariiima, p. 297, seen under the microso^
Fig. 2. LepUma ioccharina (p. 298) ; magnified and seen on the under
surface, after G. R TBEViRAinjs Verm, Schr. n. Tabi n. fig. I.
Fig. 3. TrichoiieeUi Equi magnified, as an examfde of Mallaphaga,
pp. 301, 302, after Gublt Magaz, f. d gesammt. Thierheilt ix
Tab. I. figs. 4 — 6. a, from aboye ; b, from below, attached to a
hair; c, the jaws; the absolute size is indicated by a line, as is
also done in figs. 2 and 4.
Fig. 4. Ptdex penek'onSf male, (p. 305).
Fig. 5, The pregnant female. Both this figure and the preceding after
DuMiuL C<m$. ghi. $, I Ins. PL 53, figs. 4, 5.
Fig. 6. Xenos Peckii, female, (p. 307), after Kirbt Linn. Transact xi.,
magnified ; a, a, the shield-coyers (elytra).
Fig. 7. Xenos Rossiij female, magnified, aft^r Y. Siebold.
Fig. 8. Bombylius discolor, p. 334.
Figs. 9, 10, Upper wings of ffymenoptera, fig. 9 of Apis meUi/ica, fig. 10
of Bombus; pp. 347, 348 ; m, e, outer margin {tnargo exterior);
Of apex ; 6, base ; m, p, posterior margin ; m, t, internal mazgin;
s, t, stigma; A^rculius; c, cubitus; c, r, ceUula radialis; c,Cy e,ey c,c,
esUuloi cubiiales; c, d, ceUula disc<ndaKs; e^ c,A^ oMvla hmtf-
ales; n, 6, nervi hrachidles.
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EXPLANATION OF PLATES. 839
Fig. 11. Vespa vulgana, p. 362.
Fig. 12. Mellmus arvensia^ p. 367.
Fig. 13. Trogtis kUorius, p. 381.
Fig. 14. Sirex gigas, female, p. 386.
Fig. 15. The larva of this insect, after Rcesel.
Fig. 16. The larva of Cimbex variabUisy p. 389, afber Rcesel.
Plate IX. Insects.
Fig. 1. Caterpillar of Cerura virnda, p. 403, after Rqbsbl.
Fig. 2. Sphinx convolvuti, p. 406, seen from below. At Oy the wing-
hooklet, retincKMLv/nhy is seen, p. 393.
Figs. 3, 4. Pupae of butterflies. Fig. 3, a pupa hanging free, as in the
Tetrapoda, p. 410 j flg. 4, a pupa feuitened by a transverse thread,
as in the ffexapoda, p. 409.
Fig. 5. SaJtyrus jEgeria, p. 410.
Fig. 6. Case of a larva of Phryganeay p. 414, opened to shew the smooth
internal surface.
Fig. 7. Larva of the Antrlion, MyrmeUonf(yrmiccvn%imj p. 419.
Fig. 8. A, larva; b, pupa of a lAbeUuUiy pp. 420, 421, after Ltonst : a,
mask-like under lip ; &, rudiments of the wings.
Figs. 9 — 11. Left upper wing oi LibeU<B, p. 421; fig. 9, jEahna grandis,
fig. 10, LiheUida depressa, fig. 11, jEahna/arcipaUi ; a, triangu-
lar wing-celL
Fig. 12. Agrion puella or furecUum Chabp., p. 422.
Fig. 13. Head of a Cicada as an example of the beak {rosirum) of the
ffomoptera (p. 430); A, seen from below; b, from the side; q,
in frt)nt; a, in ^g. A, the sucker; 5, 6, the insertion of the first
pair of feet; c, the antennie. In fig. c, the three filaments
which are enclosed in the sheath of the sucker, are represented
after the sheath has been removed.
Fig. 14. Anterior or upper wing of a Cicada; this wing is entirely mem-
branou& Comp. fig. 16.
Fig. 15. Fulgaraf p. 437; FtUgora macukUa Oltv. East Ind., nat size.
Fig. 16. Fore-wing of Bdostama indictim, (p. 441), to illustrate the
characters of the Hemiptera keieropteray p. 439.
Fig. 17. N^M, p. 441 ; iTepa rubra from Java^ much resembling ^epa
cinereay but larger.
Fig. 18. Redwwm peraofiatus, p. 444. It is the specimen to which I
referred in the note in p. 276, and which is distinguished by the
shortness of one of the feet, (the second on the left side).
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840 EXPLANATION OF PLATES.
Fig. 19. Head of a Reduvius from the side, as an example of the sacker
{rostrum frarUcde) in the ffeteroptera; see p. 439.
Plate X. Insects.
Fig. 1. Gryllua ccsrulescejis, p. 453.
Fig. 2. Phyllium aicci/oliumy p. 460, a female, the figure somewhat
diminished.
Fig. 3. Head of a species of EmpuaOy to exhibit the pectinate antenns,
the character of the males, p. 461.
Fig. 4. BUUta lappaniea, p. 462.
Fig. 5, Eumarj^UB marffinattu, p. 469.
Fig. 6. Adeamua Wallu^ii Hope, from Java, as an example of Ceramby-
eina, Lamioky p. 478.
Fig. 7. EvUimuB aeaptdosua Ohevrolat, as an example of th« Curai-
lumita. This species belongs to the same sub-genus as Cureulio
imperuUia, p. 487.
Fig. 8. LyUa vesicaiortay the Spanish flj, p. 500.
Fig. 9. MelolorUha vulgaris, p. 514.
Fig. 10. Under jaw of a Carabus magnified, to illustrate the primary
characteristic of the (hrabieinaf namelj the two palps, see
p. 545.
Plate XI. Arachnids and Cbustaceans.
Fig. 1. Oral organs and shears (first pair of unchanged feet) of PorUmus
momcbs, p. 679. See p. 600 ; a, upper jaws; a', palps; 5, first pair
of under jaws ; c, second pair of under jaws ; d\ first ; cT', second ;
d'"y tliird pair of foot-jaws; Jl, flagrumy or the palp attached to
these jaws; a, first pair of unchanged feet, the shears or claws.
ilg. 2. Oral organs and first pair of feet of Scorpio reticukOuSy natural
size. According to the theory of Savigny (p. 557), the upper
jaws p'* correspond to the second pair of foot-jaws of the
decapod crustaceans; the under jaws p'^' to the third pair of
these foot-jaws ; the first pair of feet, /S, correspond to the
second pair of feet of the decapodck Between these feet two
triangular plates are seen ; see the note^ p. 557.
Fig. 3. Nymphon grossipes (p. 573) in its first state, magnified (natural
size about \ line); after Kboeteb's Tidsskr. in. PL 3, fig. 26 ;
^g, 4; comp. p. 571.
•Fig. 4. Pycnogonum lUtorale, p. 573, after GuiiUN Ioonogr.y Araehn.
PL 4, fig. 1, nat siza
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EXPLANATION OF PLATES. 841
Fig. 5. Ixodes Eicimu^ p. 577, magnified after Gublt Magazin /, d
gesamnU. Thierheilk, ix. Tab. L fig. 18.
Fig. 6, Tdypkonua proacarpio Latb., p. 586, nat. siza Fig. 6 (, the
oephalothorax from the side, to shew the lateral eyes a.
Fig. 7. I^',p''\ oral organs of the same arachnid; p", upper jaws; />"',
under jaws ; magnified two diameters.
Fig. 8. Dorsal vessel or heart of TelyphonuSy with the adipose body,
magnified.
Fig. 9. Nervous system of Telyphonus^ magnified.
Plate XII. Arachnids and Chustaceans.
Fig. 1. An upper-jaw of Epeira diadema laid open, with the salivary
duct, magnified, from Treviranus Ueb. den innem Bau der
ArachnideUy Tab. IL fig. 21, see p. 560.
Fig. 2. Spinnarets of Arcmea tUrox (see p. 588), magnified, after
Treviranus op, ciL Tab. v. fig. 43; a, vent; 6,6, larger and
anterior; c, c, smaller posterior; d, d, lateral posterior papillae,
accessory feelers according to Treviranus {palpi accessorii).
Fig. 3. LimuLiia mohiccanua, p. 623, reduced. A female seen on the
under surface.
Fig. 4. Argvlua foliaceua, p. 631, on the under surfiice, laying its eggs ;
after Jurine Ann, du Mvs, vii. PL xxvl fig. 3.
Fig. 5. AnaH/a leevis, p. 638 ; one side of the shell is removed to exhibit
the animal in its natural position; nat size.
Fig. 6. Daphnia pulex, p. 641, a male, magnified after Straus MeTru du
Mu8. V. PL XLix. fig. 18.
Fig. 7. Idoiea entomon, p. 654, nat. size.
Figs. 8 — 10. Portunvs McsnaSj p. 679, nat size; ^, 8, from above,
fig. 9, under surface with the tail lying on it, fig. 10, the tail
thrown back.
Plate XIII. MoLLUSca
Fig. 1. GhUon Cummingii. Example of a multivalve shell, p. 682 ;
comp. p. 789.
Figs. 2—7. Examples of bivalves, p. 682.
Fig. 2. Terebralvlcu A species from New Guinea, see p. 722. a, ventral
shell with apex perforated ; B, dorsal shell from the inside with
the calcareous arc.
Fig. 3. Shell of Avicula {Meleagrina) margarUi/era, pp. 730, 731. On
the inside are seen some pearls attached, a, impression of
the adductor muscle.
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842 EXPLANATION OF PLATES.
Fig. 4. Area tortuoia L., p. 733.
Fig. 5. Shell of Trigonia peeUnata Lax., p. 73i. Fiom t^ insadt^ to
shew the hinge {cwrdo)»
Fig. 6. Vmu$ Dume L., Cytherea Diane Lajl, pi 742. The sheD is
placed in the position aooording to which Lm jeus hae described
it (see the note, p. 683), namelj, with the point. A, downwards;
the^tfttra, b, forwards ; the lunula, a, badcwarda.
Fig. 7. Venua {Cj/therea) erycina Lam. Right Talye from within, a, the
point; a, lunolar cavity; by fissnre; e, hinge; d, impression of
the anterior; «, of the posterior adductor mnsde; /, impression
of the mantle; g, g, inferior maigin of the shdL In this figme
the shell is placed in its natural position. On the left side the
fore end is placed, which the animal, when moviog, directs for-
wards.
Figs. 8 — 11. Examples of univalve shells or cochlecsy p. 683.
Fig. 8. Shell of Pwrpura ConcholepaSy p^ 807.
Fig. 9. Shell of PaUUa, p. 790 ; PateUa umbeUa Gm.
Fig. 10. Shell of Scalaria imperialiSy p. 798.
Fig. 11. Shell of BtUimus cUrinua B&uo., p. 814. Example of a Cochlea
einiatra, see p. 684.
Plate XIV. Molluscs.
Fig. 1. Shell of Oliva erythroHoma Lah., p. 810 ; example of an invo-
lute sheU, p. 683. a, the aperture ; a, external margin ; 6, in-
ternal margin.
Fig. 2. Shell of Nautilus PompUiua L, divided in the midplane, p. 825,
an example of a shell rolled upon itself, and afc the same time
many-chambered {cochlea revoluta, polythalanUa) pp. 683, 684;
A, A, the last chamber, or cavity, in which the animal lies; a, a, a,
the siphon which traverses the successive chambers.
Fig. 3. A cover (Operculum) of Turbo, from the inside ; see p. 684.
Fig. 4. Salpa (runcinata f) p. 699. Of the natural size, from a specimen
presented to me by Prof Y. Beneden. a, the animal seen fix>m
the surface which is oommonly turned upwards, according to
EsctfRicHT the ventral sur£ftoe. Here the muscular belts extend
throughout ; b, the animal seen frt>m the opposite side, where
the muscular belts are interrupted ; a, (in ^g, a) the fissured
aperture which the animal turns forwards; ft, the posterior
aperture ; c, the so-named nucleus, or the collection of the vis-
cera ; d, the iaUpa chain in process of development ; /, g, muscular
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EXPLANATION OP PLATES. 843
belts ; ky the gill ; Z, the heart ; r, dorsal folds {endottyle ?) ; t,
langaet ; b, (in b) the cerebral ganglion.
Fig. 5. Ascidia (fiynthia) Momus^ p. 707 ; after Savignt Mhnoirea, n.
PL Yi. fig. 1. A, respiratory aperture ; B, cloacal aperture ; c,c,e,
outer coyering, of which one half is removed in order to shew
the internal envelop d (the nuintle) ; e, muscular fibres on the
mantle.
Figs. 6, 7. Fig. 6. Lingtda anatina Lam., p. 721 ; after QuriESiAnn. du
Mils. I. fig. 6. The animal in its shell with the peduncle b.
Fig. 7. The animal removed from the shell, one of the laminn
of the mantle thrown upwards ; a, the cerebral ganglion, accord-
ing to Cuvieb; b, mouth; c,c, the two arms ; d,d,d,d, pectinated
gill-folds on the mantle.
Fig. 8. The animal of an Anodonta in the shell, of which the right valve
is removed, as an example of LamellibranchicUes, pp. 723, 724 ;
oomp. PL 15, fig. 1 ; a, shell (left valve from the inside) ; b, b,
the two laminae of the mantle j c, c, outermost gills ; d, e, inner
gills; g, the flat ventral disc projecting between the gills ; * ten-
tacles round the mouth, those of the right side alone are seen;
t mouth ; a, anterior adductor muscle ; b% V, the posterior
border of the mantle beset with fringe-like appendages.
Plate XV. Molluscs.
Fig. 1. A transverse section of ATwdcmta ancAina^ schematic after Neu-
WYLEB, Neu/6 DerJcschr. der allg. Schweizer, GeadUch. Jur die
geaaamnten N(Uv/nm8aen8chafien, Bd. vl 1842, Tab. 1 ; a, a, (, (,
and ^, as in fig. 8 of the preceding plate; c, external laminss of
the outer gills with the dorsal side attached to the mantle ; d,
internal lamina of the outer gill ; «, external ; f^ internal lamina
of the inner gill ; A, heart, traversed in the middle by the
rectum.
Fig. 2. Solen legvmen L., p. 754 ; after Poli Test, utr, SicU. l Tab. xl
fig. 15 ; A, the foot projecting at the anterior extremity of the
shell ; B, c, the two tubes (tracheae) coming into view from the
shell behind.
Fig. 3. Clio borealia Ghel., p. 775 ; after Eschkicht Anat, Untera,
Tab. L fig. 3 ; the animal is represented from the ventral sur&ce ;
a, a, fin-like appendages ; (, b, conical appendages ; c, c, tentacles
of the head ; d^ everted penis with a vesicle d' at the base.
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844 EXPLANATION OF PLATES.
Fig. 4. The oentnl neirooB system of Hdix Pamaiia L. ; after Bkasst
and Ratzebubg Mediz. ZooL n. Tab. 34, fig. 12; eompare p. 766 ;
1, ganglion sitimted above the OMophagas; 2, bdow it.
Fig. 5. Blood-circulation of Jlelix Fomatia L. ; after Cutier, as an
example of the Ptieumanica, p. 811 ; a, heart; c, auricle, both
lie in the pericardial sac, hud open; b, Vy artmee; e,e, Teaons
currents, from which the pulmonary veins in the ie^irat<»7
cavity arise, which pass into the trunk d that runs to the
auricle; f^f^ calcareous sac or kidney which surrounds the peri-
cardiuuL Compare pp. 760, 761.
Fig. 6. Limax cUer L., the anterior portion of the animal, after Brandt
u. Ratzbb. op. eii. Tab. 38, fig. 3, {Arion empirioorum brunneusy
A, respiratoiy aperture at the margin of the mantle. See
p. 817.
Fig. 7. Circulation of the blood in a cephalopod mollusc, LoUgo, afto-
CuviBB TM. elem. de VHui. not, des Animaux; a, vena cava,
which divides into two branches; b, b, lateral or venous hearts;
e, c, branchial veins; d, d, space where the gills are situated ; /,
the middle or arterial heart ; g, g, g, arteries. The direction of
the current is indicated by arrow& Compare pp. 762 and 819.
Fig. 8. A small specimen of Sepia officinalis L., p. 829; after Casus
Nov. Act. Acad. Ccbs. Natwr. Curios, xil; a, a, a, a, o, a, o, a, the
eight conical arms, the two long thin arms are seen at b, 6; c,
funnel; comp. pp. 818, 819.
Hg. 9. Detached arm {Hectocotylus) of a male individual of Argonauia
argo serving for copulation (magnified about three diameters) ;
after Koeluksb {Berickte der Zoot. Anstak zu WiinAurg^ 1849,
Tab. L fig. 19); comp. pp. 821—823.
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ALPHABETICAL INDEX
07 THE
LATIN GENERIC NAMES.
A.
Abyla. ii8.
Acsenitus. 381.
AcaDthia. 445.
Acanthomera. 337.
Acarufl. 575.
Acera. 787.
Acetes. 665.
Acheta. 456.
Achiaa. 321.
Acoetes. 245.
ActsoD. 779.
Actinia. 90.
Actinocrinites. 144.
Aotinodendron. 91.
Actinophrys. 51.
Actora. 319.
Adelinm. 493.
Adelostoma. 498.
Adeona. 93.
MgA. 648.
Mg\e&. 669.
JEolidisk, 780.
.^k>losoma. 229.
.^^norea. 125.
.^Esalus. 507.
i^hna. 422.
uEthalia. 436.
Agalma. 115.
Agahnopsia. 115.
Agaricia. 86.
Agathidiom. 538.
Aglaiflma. it 9.
Aglaura. 242.
Agra. 552.
Agrion. 422.
Agyrtes. 534.
AkiB. 498.
Albunea. 671.
Alcides. 484.
Alcyonidia. 78.
Alcyonium. 79.
Alderia. 780.
Aleochara. 531.
Aletirodes. 432.
Alima. 661.
Alitropus. 648.
AUecula. 492.
Alomya. 381.
AlpheuB. 666.
Alysia. 379.
Amerhinus. 484.
Ammonites. 827.
Ammothea. 79.
Amceba. 46.
Amphicoma. 511.
Amphidesma. 748.
AmphileptuB. 55.
Amphiuome. 243.
Amphion. 662.
Amphistoma. 185.
Ampbiitrite. 235.
Ampullacera. 811.
Ampulbbria. 801.
Amydetes. 524.
Anancbytes. 152.
Anaspis. 502.
Anatifa. 638.
Anatina. 751.
Anatinella. 749.
Anceus. 649.
Anchorella. 625.
Ancbvlomera. 658.
Ancillaria. 809.
AncinuR. 649.
AncuU. 784.
Ancylus. 811.
Ancyracantbus. 192.
An<b%na. 358.
Anekstes. 525.
Anguillula. 193.
Anguinaria. 95.
Anisoniera. 343.
Anisonyx. 511.
Anisotoma. 538.
Anobium. 520.
Anodonta. 736.
Anomala. 512.
Anomia. 725.
Anoplognatbus. 513.
Anotia. 437.
Anthelia. 77.
Anthia. 552.
Anthomyia. 321.
Autbopbora. 356.
Antbopbyllum. 86.
Antbosoma. 627.
Antbrax. 334.
Anthribus. 489.
Antbrobia. 589.
Antbura. 654.
Antipatbes. 84.
Aonis. 239.
Aphidius. 379.
Apbis. 432.
Apbodiua 519.
Apbrodita. 246.
Apiocrinites. 144.
Apis. 352.
Aplysia. 786.
Apolemia. 115.
Apseudes. 653.
Apus. 643.
Aradus. 445.
Aranea. 592.
Area. 733.
Arcella. 48.
Arcturus. 655.
Arenicoki. 237.
Argonauta. 830.
Argulus. 631.
Argus. 593.
Argyroneta. 591.
Aricia. 239.
Armadillo. 650.
Articerus. 532.
Asapbus. 646.
AscaUpbus. 420.
Ascaris. 190.
Aseffena. 591.
Asefius. 653.
Asilus. 331.
Asindulum. 342.
Aspergillum. 757.
Aspidisca. 55.
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846
ALPHABETICAL INDEX
AspisteB. ^AO.
ABtocufl. 607.
AsUrte. 741.
Aatasia. 51.
AflteriM. 147.
AMnsa. 86.
Atax. 581.
Atelecyolus. 673.
AthoiybU. 113.
Atlanta. 777.
Attelabus. 488.
Attus. 597.
AulacoB. 383.
AuletM. 488.
Auricula. 813.
Avicula. 730.
Ajunophorut. 559.
Bacttlites. 818.
Badister. ^49.
Balanufl. 037.
BanohuB. 38a.
BaridiuB. 484.
BaasuB. 381.
Bdella. 581.
B«leiimit6B. 8a8.
BeloBtoma. 44a
Bembex. 369.
Bembidium. 546.
BeriB. 337.
Bero«. III.
BiMo. 340.
BibliB. 41a
BiiguB. 670.
BittacuB. 415.
BlapB. 49jr.
Blatta. 461.
Boltenia. 707.
BombuB. 355.
BombyliuB. 333,
Bombyx. 403.
BomolochuB. 6a8.
Bonellia. 161.
BopyruB. 646.
BoreuB. 416.
BoBtrichuB. 530.
BothriooepbaluB. 179.
Botryllus. 703.
BotyB. 395.
Bk^ohiella. 615.
BraohinuB. 551.
BraohioDUB. 9or.
BrachyoeruB. 487.
Braohyopa. 318.
Braoon. 380.
Branobiobdella. 916.
BrancLipuB. 649.
BraBBolis. 410.
Braula. 313.
BrentuB. 488.
BrochuB. 489.
Buccinum. 807.
BulimuB. 814.
Bulla. 788.
BupreBtiB. 517.
Buraaria. 54.
Bunatella. 787.
ByrrhuB. 538.
Oalandra. 483.
Galappa. 674.
CalatbuB. 550.
Galoeola. 731.
GaliguB. 630.
Call^ira. 11 1.
Callioera. 336.
Callirhipis. 515.
Galobata. 319.
Calymene. 646.
CalypticuB. 431.
CalyptFBea. 795.
CamarotuB. 488.
Gampanularia. 76.
Campy lites. 826.
Campylomyza. 341.
CaDoellaria. 805.
Cancer. 678.
CanopuB. 448.
CapreUa. 656.
CapBUB. 446.
CapuluB. 796.
CarabuB. 547.
Cardilia. 75a
Cardita. 737.
Cardium. 740.
Carenum. 550.
Garinaria. 777.
GamuB. 324.
GaiyocriniteB. 144.
GaryophyllttUB. 178.
Garyopbyllia. 86.
GaBBida. 473.
GaBBiduloB. 154.
Gassiopea. 113.
GasaiB. 806.
GaBtnia. 406.
GataaoopuB. 552.
GatoM, 534.
Gavolina. 781.
Gebrio. 515.
Geddomyia. 341.
GeoropB. 619.
Cdlularia. 94.
GelyphuB. 320.
Genia. 779.
GentriB. 356.
Gephalotes. 55a
Gephea. 123.
GephuB. 386.
Gerarabyx. 479.
GenmioB. 360.
Geraphrcm. 574.
GercomonaB. 50.
Geria. 326.
Gerithium. 803.
Gerocoma. 501.
Geromya. 752.
Gerophytum. 527.
GeroplaiaB. 342.
Gerura. 402.
Cestum. 120.
Getonia. 509.
Ghstogaster. 229.
GhetonotuB. 202.
GhaetopieruB. 237.
GhalciB. 377.
Ghama. 738.
GhdracanUras. 191.
Ghelifer. 584.
Ghdyoeoma. 707.
ChenuBB. 435.
GhereiB. 597.
CheyletuB. 581.
Ghilodon. 54.
Ghionea. 343.
Ghirodota. 159.
GhironomuB. 344.
GhiroBoeliB. 490.
Chiton. 789.
Chlenius. 549.
Ghlamidodon. 55.
Ghlamidomonas. 51.
CholuB. 484.
GhondracanthuB. 626.
ChrysiB. 372.
GhryBomela. 471.
Chiysotoxum. 326.
Cicada. 438.
Gicindela. 554.
Cidaris. 156.
Gimbex. 389.
Gimex. 447.
GionuB. 484.
Girolana. 648.
GirratuluB. 238.
GiB. 520.
Cistela. 492.
Gladocora. 86.
GlauBilia. 814*
Clara. 74.
Clavagdla. 757.
darelina. 706.
Glavella. 626.
Ckviger. 532.
Gleodora. 773.
Glepeine. 225.
GleroB. 522.
Glidius. 533.
Glinocen, 329.
GKo. 775.
Glotho. 591.
Ghibiona. 591.
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OF THE LATIN OEJSTERIC NAMES.
847
Clymene. «33.
Clymenia. 8^6.
Clypeaster (Echinoderma^
to). 154. (Inteela), 468.
Gnodalon. 493.
Cobaz. 437.
GoocmeUa. 467.
C00CU8. 431.
GGelopa. 319.
GoBXiosia. 3)0.
Osnomyia. 336.
CoBDunis. 1 8a.
GoUetes. 359.
Golliaris. 554.
Colpoda. 54.
Golpodes. 550.
Columbella. 809.
Colorua. 101.
Colydimn. 536.
ComatulA. 145.
Gonops. 335.
Goniis. 808.
Gonvoluta. 331.
Gopris. 518.
Goptodera. 552.
Corbia. 745.
Gorbula. 751.
Gordisies. 551.
Gordyla. 34a.
Gordylura. 317.
GoreuB. 447.
Goriooella. 995.
Goronula^ 637.
Gorophium. 058.
Gonyra. 55a.
GofyDe. 75.
Goiymorpha. 76.
Gorysies. 673.
Go88onu8. 483.
GoasuB. 403.
GosByphus. 494.
Goxeiufl. 4^4.
Grabro. 307.
GrambuB. 394.
Granohia. 8^0.
Gntngon. 606.
Grania. 731.
Grenatula. 719.
GreBeis. 774.
Greusia. 637.
GriooeriB. 474.
GriBtateUa. 96.
GrypticuB. 407.
GryptocepbaloB. 471.
GiTptomonaB. 50.
GiyptopbagUB. 536.
GryptopB. 395.
GryptorfaynchuB. 484.
GiyptoB. 381.
GtenipuB. 550.
Gtenodaotyla. 551.
Gtenostoma. 554.
GtenuB. 595.
Gucujus. 536.
GucuIlanuB. 189.
Gulez. 345.
Guma. 663.
Gumingia. 749.
Gurculio. 486.
GyamuB. 656.
Gyamea. 134.
GyathocriniteB. 144.
GychruB. 548.
C^cUb. 744.
C^dooepbala. 516.
GyclopB. 633.
GycloBtoma. 801.
Gydippe. I3i.
Gylaa. 488.
Gymbnlia. 774.
C^moihoa. 647.
Gynipa. 385.
C^mthia {Crust,), 663.
(Tun.), 707.
Gyphomyia. 337.
(^bon. 534.
C^naa. 810.
(^pricardia. 741.
Gypridina. 633.
Carina. 743.
CypriB. 633.
Gyrena. 743.
Gyrenolda. 744.
C^ufl. 333.
GystioerciiB. 183.
Gythere. 633.
D.
Dacne. 470.
DanaiB. 411.
Dapbnia. 641.
DaBcUluB. 534.
Delpbax. 437.
Delphiniila. 799.
Demeirias. 551.
Dentalium. 791.
Derbe. 437.
DermaleichuB. 576.
DennanyBBus. 578.
DeimesteB. 537.
DeroBtoma. 333.
DesmooeruB. 477.
Diansea. 133.
Diaperis. 494.
Diapria. 375.
Diazona. 704.
DicbeleBtbium. 637.
Diooelus. 549.
Dicranopbora. 338.
Dictyoptera. 534.
Didemnum. 703.
Difflagia. 48.
DilopbuB. 340.
Dinemura. 630.
DinocbariB. 303.
DinopbyslB. 53.
Diopatra. 343.
Diopbrys. 55.
DiopBiB. 319.
DiphyeB. 118.
DiploBtomum. 185.
Diplozoon. 187.
Dircaea. 491.
DiBtoma. 184.
DiBtomuB. 704.
Dixa. 343.
DolicbopuB. 339.
Dolium. 807.
DolomedeB. 595.
Donacia. 475.
DoDaz. 746.
Dorippe. 673.
DoriB. 784.
DortbeBia. 433.
DoryluB. 366.
Doto. 783.
DrasBUB. 591.
DriloB. 533.
Dromia. 673.
DromiuB. 551.
Dxyinus. 374.
DryopB. 540.
DryopbtboruB. 483.
S'TPto. 553.
DysaBter. 153.
Dysdera. 590.
DyaideB. 520.
I>ytiBCii8. 543.
Eofainiacua. 574.
EcbinoooocuB. 181.
EcbinoDeua. 154.
Eobmorbyncbua. 184.
EobmoBpbceritea. 143.
EcbinuB. 155.
Edwardaia. 93.
Efferia. 675.
EUpbruB. 546.
Elater. 535.
ElencbuB. 308.
Eleutberia. 74.
Elraia. 540.
Emaiginula. 793.
Empia. 333.
Encauatea. 470.
Enoeladua. 550.
Encbelya. 54.
Encby^us. 330.
Encrinua. 144.
Encyrtua. 376.
Endeia. 573.
Endomycbua. 468.
Enoplua. 194.
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848
Enteroplea. 203.
Epeira. 594.
Ephemera. 433.
Ephesia. 240.
Ephippium. 338.
Ephyn. 665.
EpichariB. 356.
EpiBcapha. 470.
EpistyUs. 59.
EpitragoB. 493.
Epomifl. 549.
EreeoB. 597.
ErgaeiluB. 610.
ErichthiiB. 661.
Eriodon. 589.
Eriphia. 678.
ErodiuB. 498.
ErotyluB. 469.
Erasa. no.
Enrilia. 50.
Erycina^ 408.
ErythneuB. 581.
Eschara. 93.
Etheria. 736.
Euoera. 356.
Eucheila. 551.
EuchlaniB. 201.
EucnemiB. 527.
Eudozia. 119.
Euglena. 52.
Eurioeaa. 355-
Euumene. 642.
EulophuB. 376.
EumeaeB. 300.
EamorphuB. 469.
Eunice. 243.
EuploteB. 55.
Euprepia. 401.
Euryale. X46.
Eurychora. 498.
Eurytoma. 377.
Evadne. 641.
Eyagora. 79.
EraDia. 383.
Explanaria. 86.
Eytaifl. 580.
FaBciularia. 804.
Feronia. 549.
Fibularia. 154.
FiguluB. 507.
Filaria. 191.
FiliBtota. 59>-
FisBurella. 792.
FlabelliDa. 781.
FUta. 437.
Floscolaria. 199.
FluBtra. 94.
FoenuB. 383.
Forficula. 463.
ALPHABETICAL INDEX
Formica. 363.
FragUia. 748.
Fulgora. 437.
Fungia. 87.
Funiculina. 80.
Furcularia. 204.
FuBus. 803.
GaUthea (Crutl.). 668.
{Oonch,). 743.
Galba. 527.
GaleodeB. 585.
Galerita. 553.
Galerites. 153.
Galeruca. 473.
GalguluB. 441.
Gamasus. 570.
GammaruB. 059.
GastrochsBna. 757.
GastropteroD. 787.
Gastrus. 315.
GecarciDUB. 677.
GeophiluB. 295.
G^rissaB. 540.
Geotmpes. 517.
GerriB. 443.
Gervillia. 729.
Gibbium. 521.
GlaphyruB. 511.
Glaucoma. 54.
GUuconome. 743.
GlaucopiB. 405.
GlaucuB. 781.
GlomeriB. 292.
Glycera. 240.
Glycimeris. 752.
Gnathodon. 749.
Goniada. 239.
Gonium. 51.
GordiuB. 194.
Gorgoniat 83.
GrapsuB. 677.
Gromia. 48.
Gryllotalpa. 457.
GrylluB. 452.
Gymnolepaa. 639.
GymnoBoma. 323.
GyrinuB. 542.
GyropuB. 303.
HsBmoohariB. 228.
HalictophaguB. 308.
Haliolis. 792.
HalipluB. 544.
Halodactylus. 96.
Haltica. 472.
HamiteB. 827.
Harpa. 807.
HarpaluB. 549.
HebruB. 443.
Hegeter. 498.
HebeuB. 494.
Helcon. 380.
Helicina. 801.
HeliooniuB. 411.
Helix. 814.
Helluo. 553.
Hellwigia. 382.
HelophiluB. 327.
HelophoruB. 54 1.
HelopB. 493.
HemerobiuB. 418.
Hemioosmites. 143.
Hepatua. 673.
HepialuB. 4O4.
Hermsea. 781.
HermeUa. 236.
Hermetia. 337.
Herminia. 396.
Hesione. 241.
Hesperia. 40S.
HeteroceruB. 539.
HeterocheiluB. 189.
HeterotanuB 495.
HexodoD. 515.
HimantophoroB. 55.
Hinnites. 727.
Hippa. 671.
HippoboBca. 314.
Hippolyte. 666.
HippopodiuB. E16.
Hirudo. 226.
HiBpa. 473.
Hister. 533.
HoloptiluB. 445.
HolopuB. T43.
Holothuria. 158.
HolothyruB. 578.
Homola. 672.
Horia. 502.
Hughea. 89.
Hyalea. 773.
HyboB. 331.
HyboBoruB. 517.
HycleuB. 501.
Hydatina. 204.
Hydra. 73.
Hydrachna. 580.
Hydractinia. 75.
Hydrometra. 443.
HydrophiluB. 541.
HylflBUB. 359.
Hylotoma. 388.
Hyperia. 658.
HypopblAUB. 494.
Ichneiutaon. 381.
Ichthydium. 202.
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OF THE LATIN GENERIC NAMES.
849
Idalia. 784.
Idia. 32 T.
Idotea. 654.
InachuB. 674.
Inoceramus. 739.
lone. 647.
Ips. 535. ^
Iridina. 736.
leifl. 83.
Isocardia. %i.
lasus. 437.
Ixodes. 577.
J.
Jsra. 651.
Jantbina. 798.
Janus. 780.
Julus. 291.
L.
Lacinularia. 100.
Lacrymaria. 54.
Lsemargus. 629.
Lsena. 493.
Lagiia. 504.
Lamia. 477.
Lampyris. 523.
Lamproglena. 638
Languria. 470.
Larra. 368.
Lasiopiera. 342.
Latrodectos. 592.
Lauxania. 320.
Lebia. 551.
Lecanocephalus.
Leiatas. 547.
Lenticalites. 49.
Lepadella. 201.
Lepisma. 208.
Leptis. 330,
Leptocorisa. 447,
Leptomera. 657.
Leptoplana. 223.
Leptopus. 442.
Leptara. 476.
Lemflsa. 624.
Lernanthropus
Lethroa. 517.
Leadfer. 663.
Leucospis. 378.
Leucosia. 673.
Libellula. 421.
Libythea. 410.
Licinus. 549.
Ligia. 652.
Ligula. 178.
Lima. 728.
Limacodes. 402.
Limapontia. 779
192.
626.
Limaz. 816.
Limnadia. 643.
Limnsus. 812.
Limnias. 201.
Limnobia. 343.
Limnochares. 579.
Limnoria. 653.
Limulus. 622.
Lingula. 721.
Linyphia. 593.
Liorhynchus. 191.
Liosoma. 159,
Liotheum. 302.
Lispe. 321.
Liiisosoma. 779.
Lithobius. 294.
Lithodes. 672.
Lithophilus. 494.
Litfaosia. 401.
Lituites. 826.
Locuata. 455.
Loligo. 829.
LoligopsiB. 830.
Lophosia. 323.
Lophyrus. 388.
Lorioera. 549.
Lottia. 791.
Loxocera. 317.
Loxodes. 56.
Lucanns. 505.
Lucemaria. 92.
Lucina. 744.
Lumbricidus. 231.
Lumbricus. 231.
Lumbrineris. 242.
Lycosa. 595.
Lyooperdina. 468.
LycuB. 524.
Lyda. 387.
Lygaens. 446.
Lymexylon. 521.
Lysidice. 242.
Lystra. 437-
Lytta. 500.
M.
Machilis. 207.
Macrobiotus. 574.
Macrocera. 342.
Macropeza. 343.
Macroetoma. 222.
Mactra. 749.
Madrepora. 85.
Maeandrina. 86.
Ma^uB. 793.
Maja. 675.
Malleufl. 730.
Malthinus. 523.
Mammillifera. 89.
Mantioora. 555.
VOL. I.
Mantis. 461.
Marginella. 809.
Marsapites. 145.
Masaris. 360.
Mastigufl. 533.
Matuta. 673.
Megachile. 357.
Megasoelis. 474.
Megastelus. 381.
Megistocera. 343.
Melandrya. 491.
Melania. 802.
MelasiB. 527.
Me]icerta. 200.
Melocriniies. 144.
Meloe. 499.
Melolontha. 513.
Melophila. 313.
Melyris. 522.
Membracis. 436.
MermiB. 194.
Mesodesma. 749.
MesoBtoma. 222.
Metopius. 382.
Microdon. 326.
Microgaster. 380.
MicTopeza. 319.
Microstoma. 222.
Milesia. 327.
Miiiola. 50.
Millepora. 85.
Mihiesium. 574.
Minyas. 91.
Mitra. 809.
Molpadia. 160.
Moloris. 498.
Monas. 50.
MoooceliB. 223.
Monodonta. 799.
Mononyx. 441.
MonopnlebuB. 432.
Monostoma. 185.
Monticularia. 87,
Monnra. 202.
Mopsea. 83.
Mordella. 502.
Mormolyce. 551.
Morpho. 410.
Munna. 653.
Murex. 804.
Mursia. 674.
MuBca. 321.
Muldlla. 366.
Mya. 752.
Myadora. 751.
Myas. 550.
Mycetobia. 342.
Myoetochares. 492.
MyoetophaguB. 537.
Mycetophila. 342.
Mycoris. 444.
MycteroB. 490.
54
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Google
850
MydM. 330.
Mygale. 589.
MyUbrifl. 501.
Mymar. 373.
Myochama. 750.
Myopa. 345.
Mysifl. 663.
Mynnica. 364.
Myrmecium. 596.
Myrmecopfaila. 456.
Myrmeleon. 419.
Mytilus. 731.
MyzoBtoma. 626.
N.
Naifl. 319.
Narica. 797.
Naaaula. 54.
Natica. 797.
NaacoriB. 440.
Nautilus. 835.
Navicella. 796.
Nebalia. 64a.
Nebria. 547.
Necrophoros. 533.
Necydalis. 479.
Nemertes. 334.
Nemesis. 627.
Nemestrina. 334.
Nemoptera. 418.
Nemosoma. 536.
Nemotelus. 338.
Nepa. 441.
Nephelis. 125.
Nephtaea. 78.
Nephtfays. 240.
Nephrotoma. 344.
Nereis. 241.
Nerita. 797.
Nioothoe. 628.
Nilio. 494.
Nitidula. 535.
Noctua. 398.
Nodosaria. 48.
Nomada. 357.
Nops. 590.
NotarchuB. 787.
Notiphila. 320.
NoUnlonta. 402.
Notommata. 204.
Notonecta. 440.
NotoxuB. 503.
Nucleolites. 154.
Nucula. 734.
Nummulina. 49.
Nummulites. 49.
Nycteribia. 312.
Nympbalis. 411.
Nympbon. 573.
NysM>n. 368.
ALPHABETICAL INDEX
Obrium. 478.
Oceania. 125.
Octobotbrium. 187.
Octopus. 831.
Ocubna. 80.
Ocypode. 677.
Ocyptera, 323.
Odacaotba. 551.
Odontobius. 190.
CEdemera. 490.
(Enas. 500.
(Estnia. 315.
Ogygia. 645.
Olenus. 645.
Oletera. 589.
Olios. 595.
Oliva. 810.
OmalisuB. 524.
Omalium. 530.
Omopbron. 547.
Oncbidium. 817.
Oniscoda. 652.
Oniscus. 65 T.
Onupbis. 243.
Oodes. 549.
Opatrum. 496.
Opbion. 382.
(^biostoma. 189.
Opbiura. 147.
Opbrydium. 57.
Opbryoglena. 54.
Opis. 741.
Oplopborufl. 665.
Orbicula. 721.
OrbicuHna. 49.
Orcbestes. 484.
Oribata. 577.
Oritbyia. 674.
Omiukomyia. 313.
Orpbnus. 517.
Ortalis. 318.
Orsodacne. 476.
OrtbooeraUtes. 827.
OrtbogoniuB. 552.
Oiyssus. 386.
Oscinis. 320.
Ostrea. 726.
Otoglena. 203.
Ovula. 810.
Oxycepbalus. 657.
Oxyoera. 338.
Oxytelus. 530.
Oxyuris. 190.
Oaena. 550.
Osodicera. 343.
PachycDema. 511.
Paederus. 529.
Pagums. 670.
Pa]»moQ. iS^.
Palinurua. 669.
Pallene. 573.
Palmyra. 244.
Paludina. 800.
Palythoa. 89.
Pamborus. 548.
PaoagsBUS. 540.
Pandarus. 630.
Pandora. 751.
Pandorina. 51.
Panopiea. 753.
Panorpa. 415.
Panui^gus. 358
PapUio. 409.
Paragus. 326.
Panmeoium. 54*
Parandra. 480.
Paribcea. 573.
Parmopbonis. 791.
Pamopes. 372.
Partbenope. 676.
Passalus. 508.
Passandra. 536.
PateUa. 790.
Paussus. 538.
Pavonia. 86.
Pecten. 728.
Pectunculus. 733.
Pedicia. 344.
Pediculus, 300.
Pedinus. 497.
Pedum. 728.
Pelednus. 383.
Peliocera. 328.
Pelobius. 544.
Pelocopbora. 522.
Pelogonus. 441.
Peltis. 535.
Penaeus. 0(S5.
Pennaria. 76.
Pennatula. 8a
Pennella. 625.
Pentacrinus. 145.
Pentacta. 157.
Pentastoma. 188.
Pentatrematites. 143.
Pentbretria. 340.
Peridinium. 53.
Peripatus. 238.
Pex4a. 425.
Pema. 729.
Peropbora. 706.
Petricola. 745.
Pbalacras. 470.
Pbalaena. 396.
Pbalangium. 583.
Pbaleria. 495.
Pbailusia. 706.
Pbania. 323.
Pbasia. 323.
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OF THE LATIK GENERIC NAMES.
851
PhaaianeRa. 800.
Phasma. 458.
Phialma. 53.
PhilodromuB. 595.
PhilopteruB. 301.
Pholiulomja. 752.
Pholas. 754.
Pholcus. 592.
Phora. 317.
Phoxich^dinm. 573.
Phronima. 657.
Phronoa. 658.
Phiyganea. 414.
Phrynus. 586.
PhiTXUB. 647.
Phjllidia. 785.
Phyllirhoe. 779.
Pbjllium. 400.
Phyllodoce. 140.
Phyllophora. 630.
PhylloBoxna. 661.
Physa. 8 1 3.
Physalia. iii.
Physsophora. 113.
Pieris. 409.
PiestuB. 530.
Pileolua. 796.
Pixnelia. 498.
Pinna. 731.
Pinnoteres. 677.
PinophiliiB. 599.
Piopbila. 320.
Pipunculus. 329.
Pirnpla. 381.
Placana. 726.
Plagusia. 677.
Planaria. 223.
Planaxia. 80a.
Planorbia. 813.
Platycrinites. 144.
Platygaster. 374.
Platypeza. 329.
Pla^teryx. 398.
Platypus. 481.
Pla^^ura. 342.
Plecia. 340.
PleuTobranchna. 786.
PleurobranchuB. 786.
Pleurophyllidia. 785.
Pleurotoma^ 803.
PlocamopberuB. 784.
Ploiaria. 444.
Pliunatella. 96.
Pnemnodermon. 775.
Pneumora. 454.
PociUopora. 85.
Podura. 208.
Pogonua. 549.
PoUicipefi. 639.
Pollyxenus. 290.
Polyarthra. 202.
Polyoelin. 223.
Polycera. 783.
Polyclinum. 703.
Polydeamus. 290.
Polymera. 343.
Polynoe. 245.
Polyodontes. 245.
Polyommatus. 408.
Polyphemus. 640.
Polystoma. 186.
Polyzonium. 290.
Pompilus. 370.
Ponera. 365.
Pontobdella. 228.
Poroellana. 669.
Porphyrophora. 432.
Porpita. III.
Portunns. 679.
Poteriocrinitee. 144.
Potamopbiins. 540.
Praniza. 650.
Pray a. 119.
Priapulus. 161.
Primno. 658.
Prionos. 480.
Pristina. 229.
Procerus. 548.
Procirrus. 529.
Procrustes. 548.
Proctonotus. 780.
Proctotrupes. 375.
Productus. 723.
Pronoe. 657.
Prosoopia. 454.
Prostoma. 222.
Prostomis. 536.
Proteinus. 531.
Prozichilus. 573.
Psammobia. 746.
PsaniB. 326.
PselaphuB. 532.
Psilomyia. 318.
Psocus. 426.
Psolus. 158.
Psyche. 402.
Psychoda. 341.
Psylla. 435.
Pterochilus. 780.
Pterodina. 201.
Pteromalus. 376.
Pterophoi-UB. 393.
Pteroptus. 578.
Pterotrachea. 777.
Ptilinus. 521.
Ptilium. 537.
Ptiloccra. 337.
Ptilodactyla. 524.
Ptinus. 521.
Ptychoptera. 344,
Ptygura. 200.
Pulex. 303.
Pupa. 813.
Purpura. 806.
PycnogonuDL 573.
Pygaster. 153.
Pyramidella. 798.
I^igoma. 637.
I^ochroa. 503.
Pyrosoma. 702.
Ranina. 672.
Rapbidia. 4x6.
Rattulus. 202.
Keduyius. 444.
Rembus. 549.
Remipes. 671.
ReniUa. 81.
Retepora. 93.
Rhagium. 477.
Rhina. 483.
Rhingia. 328.
Rhinosimus. 490.
Rhinotia. 488.
Rbipicera. 525.
Rhipidia. 343.
Rhipiphorus. 501.
RhiiEophthiridium. 433.
Rhixophysa. 1x4.
Rhizostoxna. 123.
Rhodocrinites. 144.
Rhynchsenus. 485.
Rhyphus. 342.
Rbysodes. 536.
Rictularia. 189.
Riflsoa. 802.
Rophites. 358.
Rossia. 830.
Rostellaria. 805.
Rotifer. 205.
Rudista. 738.
Rutela. 512.
S.
Sabella. 235.
Ssenuris. 231.
Sagitta. 246.
Sagra. 475-
Salda. 442.
Salpa. 697.
Salpina. 202.
Sandaius. 525.
Sapyga. 370.
Sarcoptes. 570.
Saigus. 337.
Saropoda. 356.
Sarrotrium. 496.
Satumia. 403.
Satyrus. 410.
Saxicava. 746.
Scalaria. 798.
Scaphidium. 535.
Scaphites. 827.
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852
ALPHABETICAL INDEX
SoM^bntu. 516.
Scftrites. 550.
SoatophogA. 3x8.
ScatopM. 341.
Scaurus. 498.
ScenopinUB. 339.
Sciara. 343.
Soolia. 370.
Scolopeodra. 395.
Scoloplofl. 439.
Soolytua. 481.
Scorpio. 587.
Soraptia. 503.
Scutella. 155.
Soutellera. 448.
Scutigera. 194.
ScyclmiBDUB. 533.
ScyUna. 781.
ScyUanii. 669.
Soytodcfl. 591.
8eg«8tria. 591.
Selenops. 594.
Semblu. 417.
Sepia. 839.
Sepidium. 498.
Sepiola. 819.
Sepioteuthii. 839.
Sepsis. 319.
Septaiia. 756.
SergMtet. 665.
Serialaria. 95.
Seriatopora. 85.
Serioaria. 40a.
Sericomyia. 317.
Serolis. 648.
Serpula. 33d.
Serropalpua. 491.
Sortuiaria. 76.
StMiia. 405.
Siagona. 550.
Sideralina. 49.
Sigalion. 445.
Sigalphua. 3S0.
Sigaretus, 795.
SigiUina. 704.
SiUauaria. 793.
Silpha. 534.
Silvanua. 537.
Simulia. 341.
Stnod«ndron. 507.
SipboDaria. 795.
Sipbonophora. ^go,
Sipbonoetoma. 934.
Sipbonotus. 990.
Sipunculus. 1 00.
Sirex. 3S6.
Smyntburua. 999.
Solemya. 754.
Solen. 753.
SparaaioD. 374,
Sparanus. 594.
SpatanguB. 159.
SpercbeoB. 541.
Spbserocera. 320.
Spbfleridium. 540.
SpbaBroma. 649.
SpbaBroniteB. 143.
SphasuB. 596.
SpbeniscuB. 493.
Spbex. 369.
Spbinx. 405.
SpbodruB. 550.
Spio. 239.
Spiratella. 774.
Spiroptera. 190.
SpiroBtomum. 54.
Spirula. 828.
Spirolina. 49.
Spondylis. 480.
SpondyluB. 727.
Squilla. 661.
Squillericbtboa. 66 1.
Stapbylinufl. 528.
Statyra. 504.
Stenocera. 554.
StenoBtoma. 490.
StenotracheluB. 491.
Stentor. 56.
StenuB. 529.
StepbanoceroB. 200.
Stepbanomia. 114.
StepbanuruB. 190.
SteropeB. 503.
Stlliger. 780.
StomoxyB. 325.
Stratiomys. 338.
StrombuB. 805.
Strongylium. 492.
Stat>ngylosoma. 291.
Strongylostoma. 222.
StrongyluB. 189.
Strutblolaria. 805.
StylocbuB. 223.
StylopB. 307.
Suocinea. 815.
SyoocystiteB. 143.
Syllis. 241.
Sympodium. 79.
Syuagris. 360.
Synapba. 342.
Synapta. 159.
Syncbnta. 204.
Syndesus. 507.
Syudosmya. 748.
Synoicum. 704.
Synora. 51.
Syrpbus. 327.
SyrtiB. 445.
Systropba. 358.
TabanuB. 334.
Tkcbina. 323.
Tacbyponu. 531.
TsnJa. 180.
Tagenia. 498.
Taliiros. 659.
TaaaiB. 653.
Tapbria. 550.
Tarpa. 388.
TeffluB. 548.
TelephoruB. 523.
Tellina. 747.
Telpbusa. 678.
TelypbonuB. 586.
Tenora. 94.
Tenebrio. 496.
Tenthredo. 38S.
Tentyria. 498.
TerebeUa. 235.
Terebellum. 809.
Terebra. 808.
Terebratula. 722.
Teredina. 755.
Teredo. 755.
TergipeB. 781.
TermeB. 426.
Tetanooera. 317.
Tethys. 783.
TetraceliB. 223.
Tetragnatba. 594.
TetragonoderuB. 549.
TetraUsmis. 638.
Tetrarbyncbafl. 183.
Tetrix. 452.
Tettigometra. 437.
Tettigonia. 436.
Tbalassema. 161.
TbalaasiaxitbuB. 91.
TbalaBBina. 667.
Tbecidea. 722.
TbemJBto. 658.
Tberates. 554.
Tbereva. 330.
Tberidion. 593.
Tbomisus. 595.
Tbrada. 750.
Tbrips. 464.
TbroBcoB. 527.
Tbyone. 158.
Tbyreopbora. 319.
TbyriB. 405.
ThysaDopuB. 663.
Tbysanozoon. 222.
Tidiogonia. 732.
TietlemaDxiia. 774.
Tinea. 394.
TingiB. 445.
TintinnuB. 59.
Tipula. 344.
TmesistaTiua. 478.
Tomicus. 481.
Toracaniba. 377.
Tornatella. 798.
Tortrix. 395.
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OF THE LATIN GENEBIC NAMES.
853
Toxlcom. 496.
Trachynotus. 498.
TrachysceliB. 494.
Trechufl. 546.
Triarthra. aoa.
Tricelis. 1^3.
TrichiuH. 510.
Trichooera. 343.
Trichoda. 53.
Trichodecies. 309
TrichodiBCUB. 5a.
Tricbosoma. 191.
Trioocephalu8. 191.
Trioondyla. 554.
Trictenotoma. 508.
Tridacna. 740.
Trigonia. 734.
Trigonella. 748.
Trinucleus. 645.
Triopa. 783.
Triplatoma. 470.
Triplax. 470.
Tristoma. 186.
Tritonia. 78a.
Trochilia. 56.
TrochiM. 798.
Trogoflita. 536.
Trogulus. 582.
Trogus. 381.
Trombidium. 581.
Trox. 517.
Trypeta. 318.
Trypbon. 381.
Tubioolaria. aoo«
Tubifez. 330.
Tubipora. 8a.
Tubularia. 75.
Tubulipora. 94.
Turbinella. 805.
Turbo. 799.
Turritella. 800.
Tylos. 650-
Xypblocyba. 435.
T^pbloplana. 2 a a.
Typbifl. 657.
IJlidia. 319.
Uloborus. 594.
UloceruB. 488.
TJlopa. 430.
Umbelluiaria. 81.
Umbrella. 786.
Unio. 735.
Urania. 407.
Urceolaria. 57.
Urocentrum. 57.
Uropoda. 578.
Uvella. 50.
V.
Yaginicola. 57.
VaUieria. 95.
Yalvata. 801.
Vappo. 338.
Yelella. tio.
Velia. 443-
Veluima. 797.
Venus. 741.
Yeretillum. 81.
Yermetua. 793.
Yesicularia. 95.
Yespa. 361.
YibiUa. 658.
"Vibrio. 45.
Yirgtilaria. 80.
Yitrina. 816.
Yolucella. 328.
Yoluta. 808.
Yolvox. 51.
Yortex. 227.
Yortioella. 58.
Yulsella. 730.
Xenia- 77.
Xenos. 307.
Xipbydna. 386.
Xorides. 381.
Xya. 458.
Xyela. 387.
Xylocopa. 358.
XylopbaguB. 336.
Xysta. 323.
Z.
ZabruB. 549.
Zelus. 444.
Zetes. 573.
ZoantbuB. 89.
Zygsna. 405.
Zygoneura. 34a.
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LIST of WORKS in GENERAL LITERATURE
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Messrs. LONGMAN, BBOWN, GEEEN, LONGMANS, and EOBEETS,
89 PATBBNOSTEB BOW, LONDOV*
CLASSIFIED INDEX.
Acrtenltnrt and Rural
Affairs.
BayldOB on Valuing RenU, fte. - 4
Ceeir* Stud Farm - * " •
H(Mkyn«'> Talpa - - " " IJ*
Loudon'* Agrleultur* - - - ;i
Low'aElemenUorAcrienlturt - W
jjlortoa on Landed Propcrtf - 18
Art*. Mannfaetureflj and
AreUlteetnra.
4
4
Bourne on tbe Screw Propeller -
Brande's Dictionary of Sci«uc<,ao.
«« Oriranic ChemUtry -
OiCTreul on Colour - - -
Creey'eCiTUBngincerinx -
Fairbaim'a Inforxna. for Engineen
OwUt'e Encyclo. of Architecture -
Harford** Plate* from U. Aogelo -
Uumphrey*'* Parabt«$ IllumUiated
Jaae*oB'*Saercd A Legendary Art
" Commonplace- Book -
K5ni|r'* Pictorial Life of Luther -
Loudon'* Rural Architecture
MacI>ouffall'a Campaigns of Han-
" Theory of War
Moeeley** Engineering • - -
Piesee's ArtofPerftimery -
Richardson** Artof Horeemanship
Scoffiem on Projectiles, Ac. -
Serivenor on the Iron Trade •
Stark's Printing ....
Steam-Bngine,by the Artisan Clnb
Ure's Dictionary of Arts, Ae.
Btography.
^•s^'-tK^KaWMe-:
Bodenstedt and Wagner's Sehamyl
Briahnont'e Wellington
Bunscn's Hippobtus . - -
C*pgra«'s Henrie* . - -
Cockayne's Marshal Turenne -
Crosae^s (Andrew) Memories .
Forstn's De Foe and ChurchiU .
Green's Princesses of En«land -
Harford's Life of Michael Angelo -
Hayward's ( hesterileld and Selwyn
Holcroft's Memoir* - -
Latdner's Cabinet Cyclopedia -
Maunder's Blowaphlcaltreasury-
Memoir of the Duke of Wellington
MounUln's (Col.) ¥«»<>•« " "
Parry's (Admiral! Memoirs - -
Rogers's Life and Oeniua of Fullar
RnsseH's Memoirs of Moore -
•« (Dr.) MenofantJ -
SchlmmelPennlnck's rMrs.) Life -
8outhey*sLlfcofWedey - -
** Life and Correepondencc
Stephen's Ecclesiastical Biojjraphy
Strickland's Queens of England -
Sydney Smith's Memoirs -
S7mond*s (Admiral) Memoirs
l^ylor's Loyola . - - -
" Wesler - . - -
TJwins's Memoirs - ' '„ '
WaieTton*s Autobiography AEssays
Booka of Oenerai UtUitr.
Acton's Bread-Book
" Cookery - ". " "
Black's Treatise on Bre ving - -
Cabinet Oaiettecr - . . .
•• Lawyer - . . -
Gust's Invalid's Own Book -
G llbart's Logic for the MUUon -
Hints on Etiquette _ , - - -
How to Nuree Rick Children - -
Hudson's Executor's Guide -
" on Making Wills - -
Kesteren's Domeetic Medicine -
Lardner** Cabinet Cyclopwiia -
Loudon's Lady's Country Compa-
nion ----••
Maunder's TreasuTT of Knowledge
** Biographical Treasury
* Geographtcsl Treasury
MMundeT's Scientilc Treasury - 11
" Treasury of Hbtory - U
*< Natural History - . 14
jMh,!.,; p Arr.of Perliunery - - 17
I\.rL.i < 11,1 ide Stud ... 8
P^,Ti.4r^ KnfTlish Reading - - 18
Fin^rrp M,,]ical Guide - - - 18
Ti I >. to Latin Dictionary 18
Ui.Jj >. . .......
}Li, El ir^SiJM i Artof Horssmanshlp
II !'i,j I <■'» [..;iti a Dictionaries •
RnpfL'* Eriiilish Thcsauius - ^ -
Ft.>« iutz-j- Utbater . . - -
S;i,4.^r{, NVhj^l . . - . -
Tli''na,uui* I »terest Tables -
W i- 1 . * \j-j ■*■ I ^.omestic Economy
u' .-it > <.\, r h ^ I dren's Diseaaes -
V.M ; I . I k ■ ■^ r.i |»ular Tables
xfiL,Tj:.fB lU,kckstone . - -
Botany and Oardonln«.
Haasall's British Freshwater Alga 9
Hooker's BriUsh Flora - - - •
« Guide to Kew Gardens - 9
«< <« ** Kew Museum - 9
Lindley*8 Introduction to Botany p
" Theory of Horticulture - 13
Loudon's Hortus Britannicus - 13
** Amateur Gardener - IS
" Trees and Shrubs - - 12
** Gardening - . - 12
** PUnU - - - - IS
Pereira's MaterU Uedica - - 17
Rivers's Rose-Amateur's Guide - 19
Wilson's British Mosses - - 24
Ohronolory-
Blair's Chronological Tables - 4
Brewer's Historical Atlas . . 4
Bunsen's Ancient E«ypt - - 5
Calendars of English State Paper* 5
Haydn's Bnt^oiTs Index - - »
Jaquemefs Chronology - - 11
" Abridged Chronology - H
Nieolas's Chronology of History - U
Oommereo and MoreantUe
Affairs.
Gllbarfs TreatUe on Banking - B
Loriraer's Toung Master Mariner 12
Macleod's Bsnklng -_ - - 1*
M'Culloch'sCommerce&Natlgation 14
Murray on French Finance - - 16
Scrivenor on Iron Trade - - 1»
Thomson's I nterestTablrt - - 23
Tooke's History of Prices - - 28
Oritlelam, Hiaionr, and
Memolra.
Blair's Chron. and Hlstor.Tablei - «
Brewer's Historical Atlas - - - 4
Bunsen's Ancient Egypt " " *
Hippolytus
Calendars of Englbh State Papers
Chapman's GustaTusAdolphus -
Chronicles A Memorials of England
Conybcire and Howson's St. Paul
Connnllj's Sappers and Miners -
Croire's History of France -
Gleig's Css^iys . - - -
" Leipsic Campaign -
Gnmey's Historical Sketches
Hay ward's Essays - - - -
Herschels Essays and Addre»»#s -
Jeffrey's (Lord) Contributions -
Kemble'e Ang lo-Saaons
Lardner's Cabinet Cyclopiidia -
MaeauUy's Crit. and Hist. Essays
«« History of England -
" Speeches -
Mackintosh's MisceUaneous Works
«« Jlistory of England -
M'CulIoch'sGeographlcalDlctionary
Maunder's Treasury of HUtory -
Memoir of the Duke of Wellington
Merirale's History of Rome -
*' Roman Republic •
Mflner's Church History - -
Moore's (Thomas) Memoirs, Ac. -
Mure's Greek Literature
Normanby'a Tear of Rerolution • 17
Perry's Franks ... - 17
Raikes's Journal - - - .18
Ranke's Ferdinand A Maximilian 23
Riddle's Latin Lexicon -19
Rogers's Bss.%ys from Edinb. Reriewl9
Roget's Enclish Thesaurus > -19
Schmita's History of Greece * 19
Southey's Doctor - . . > SI
Stephen's Eecleeiastical Biography 21
" Lectures on French History 21
Sydney Smith's Works ... 20
« Select Works - 22
« Lecture* - - 21
** Memoirs - - 20
Taylor's Loyola .... 21
Wesley .... 21
Thlrlwall'sHistoryof Greece - 28
Thomas's Historical Notes . • 6
Townsend's State Trials > - 28
Turkey and Christendom > ■ - 22
Turner's Anglo-Saxons - • 28
** Middle Ages ... 28
** Sacred Hist, of the World 23
Uwins's Memoirs . - - . 23
V'ehse's Austrian Court- - - 28
Wade's England's Greatness • 21
Toaiig's Christ of History - . 24
Oeoirraphy and Atlasea.
Brewer's Historical Atlas - . 4
Butler's Geography and Atlases - 5
Cabinet Gasetteer .... 5
Cornwall: Ite Mines, Ac. . - 22
Darrieu's Morocco - - . 22
Hughes's Australian Colonies . 22
Johnston's General Gasetteer . II
M'Cnlloch's Geographical Dictionary 14
** Rnssta and Turkey . 22
Maunder's Treasury of Geography 1$
Ma) ne's Arctic Discorerics - - 22
Murray's Encrclo. of Geography • 16
Sharp's British Gasetteer - - 20
JuTonllo Booka.
Amy Herbert -
ClereHall - -
Earl's Daughter (The)
cc of Life
- 20
- 20
Experiencc^of Life ' - - - 20
Gertrude ..... 20
Hon-itt's Boy's Country Book . 10
" (Mary) Children's Year - 10
Ivor* ...... 20
Katharine Ashton ... 20
Laneton Parsonage - - . 20
Margaret Perciral - ... 20
Pycroll's Collegian's Guide <• - 18
Ursula ...... 20
MedielnOj Snrirenr, Ac.
Brodle's Psychological Inquiries . 4
Bull's Hinte to Mothsn- - . 5
" Management of Children - 5
Copland's Dictionary of Medicine • 6 |
Cust's luTslid's Own Book - - 7 I
Holland's Mental Physiolosy . » :
** Medical Notes sndReflect. 9 I
How to Nurse Sick Children • . 10
Kesteven's Domestic Medicine . 11
Pereira's Materia Medica - -17
Reece's Medical Guide - - - 18
Richardson's Cold- Water Cure - 18
Spencer's Psycholo«T - - - 21
West on Diseases of Infancy - • 24
MiaeeUanoous and General
Iiiteratare.
DACon's (Lord) Works ... 8
Carli^'s Lectures and Addrcssos 22
Defence of Srf<MWO/Fai«A - - 7
Eclipse of Faith - . - .7*
Fischer's Bacon and Realistic Phi-
losophy ..... 7
Greathed's Letters from Delhi - 8
Greyson's Select Correspondence - 8
Gumej's Erening Recreations - 8
HsssaQ's Adulterations Detected Ac* 9
Haydn's Book of Dignities - . 9
Holland's Mental Physioh)gy - 9
Hooker's Kew GuUes ... 9
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ALPHABETICAL CATALOGUE
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NEW WORKS and NEW EDITIONS
PUBLISHED BY
Messrs. LONGMAN, BEOWN, GEEEN, LONGMANS, and EOBEETS,
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Miss Acton's Modem Cookery for Private
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Amy. New Edition, with Supplement by
LuoyAikin; consisting of additional Selec-
tions from more recent Poets. 8vo. 18s.
AragoCF.)— BiographiesofDistmgnished
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Arnold. -lUrope, a Tragedy. By Matthew
Abipold. With a Preface and an Historical
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Lord Bacon's Works. A New Edition,
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Collected and edited by Bobbbt Lbslib
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Cambridge; Jambs Spedding, M.A. of
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Joanna Baillie's Dramatie and Poetical
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leave the reader notbinr
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tfaeaeofDr.Bartfa. Short of
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