Journal of the East Africa Natural History Society and National Museum

77

//X

Ji

JOURNAL

OF THE EAST AFRICA NATURAL HISTOR
SOCIETY AND NATIONAL MUSEUM

OCTOBER 1984

o

No. 181

i „

CHECKLIST OF THE SPECIES OF TERMITES (ISOPTERA)
RECORDED FROM EAST AFRICA

K. Wanyonyi*, J.P.E.C. Darlington + , and R.K.N. BagineT-

International Centre of Insect Physiology and Ecology,

P.O. Box 30772 , Nairobi, Kenya.

INTRODUCTION

This checklist is based on one originally prepared by Kisito Wanyonyi during a period of
training in Switzerland and Britain. It was apparently intended to form part of a thesis or hand-
book which he never completed. After his death in 1975 the list was used for reference by mem-
bers of the Termite Research Programme at I.C.I.P.E.. This programme finally closed down in
1982.

In 1983 the termite collection of the National Museums of Kenya was reorganised and
catalogued. It seemed an opportune moment to revise and update the checklist, and to publish it.
The revision was carried out by J.P.E.C. Darlington, formerly of I.C.I.P.E., and R.K.N. Bagine of
the National Museums of Kenya.

The list includes 1 77 species, of which two are recent introductions. Of the 32 species recorded
only from Uganda, most probably belong to the fauna of the West African savannas, or are relicts
of the Congo Forest which was formerly more extensive. The remaining 143 species comprise the
East African fauna proper which is continuous with that of Somalia, Ethiopia and Sudan to the
North, and of Malawi, Zimbabwe and Mozambique to the South. The actual distribution of
species is closely linked to vegetation zones (Bouillon 1970).

This list is not definitive but merely a statement of the status quo. The taxonomy and
distribution of East African termites are incompletely known, especially in the un-revised taxa,
and future work will certainly lead to changes in the list, both by the addition of newly-described
species and by the suppression of species names which are found to be synonymous.

The known distribution of each species in East Africa is coded in the list as follows: U =
Uganda, K = Kenya, T = Tanzania, mainland only, Z = Zanzibar Island (Tanzania).

SOURCES

In revising the list the following published sources have been used: Snyder (1949) for all
groups that have not subsequently been revised; Harris (1966a) for the genus Coptotermes; Sands
(1972) for the subfamily Apicotermitinae; Sands (1959) for the genus Amitermes ; Williams (1966)
for the genus Cubitermes\ Ruelle (1 970), 1977) for the genus Macrotermes\ Harris (1966b) for the
genus Ancistrotermes-, and Sands (1957) for the subfamily Nasutitermitinae, eastern .group only.
The “Catalogue ethiopien” of Bouillon and Mathot (1965, 1966) was also consulted. Additional
distribution records from Harris (1951) and Kemp (1955) have been included. The layout of the
checklist follows Snyder (1 949) except in the family Termitidae in which the revised arrangement
of Sands (1972) is used.

Distribution records derived from the published sources mentioned above are quoted in the list
without a subscript. Records derived solely from unpublished sources are indicated by a subscript,
as follows:

* Deceased

■f Present address: Section of Entomology, National Museums of Kenya, P.O. Box 40658,
Nairobi.

Page 2

No. 181

1. Wanyonyi’s original list dated 1972. This was compiled from two main sources:

a) Unpublished catalogue and notes of Dr. E. Ernst of the Schweizerisches Tropeninstitut,
Basle, Switzerland.

b) Records of the Termite Research Unit, Centre for Overseas Pest Research, London. This
included unpublished collection data of W.A. Sands and R.M.C. Williams; and a
catalogue, updated by R.W. Lamb, of the termite collection housed in the Entomology
Department, British Museum (Natural History).

2. A termite collection donated to the National Museums of Kenya by W. Wilkinson of the
East African Agriculture and Forestry Research Organisation, Muguga, Kenya.

3. Termites collected by J.P.E.C. Darlington, D.E. Pomeroy, K. arap Rotich, R.K.N. Bagine
and others, most of which were identified by S. Bacchus of the C.O.P.R. Termite Research
Unit, London. The material is housed partly in the National Museums of Kenya and partly in
the British Museum (Natural History).

4. An unpublished thesis by Buxton (1979) whose identifications were checked by W.A.
Sands and R.M.C. Williams and whose material is deposited at the British Museum (Natural
History) in London.

FAMILY KALOTERMITIDAE
Subfamily Kalotermitinae

Genus Epicalotermes Silvestri, 1918
E. aethiopicus Silvestri, 1918
E. kempae (Wilkinson, 1954)

E. mkuzii (Coaton, 1949)

Genus Neotermes Holmgren, 1911
N. aridus Wilkinson, 1959
N. cryptops (Sjbstedt, 1 900a)

N. firm us (Sjostedt, 1911a)

N. meruensis (Sjbstedt, 1 907a)

Genus Bijiditermes Krishna, 1961

B. angulatus (Wilkinson, 1959)

B. durbanensis Haviland, 1898
B. jeannelanus (Sjbstedt, 1915)

B. sibayensis (Coaton, 1 949)

B. sylvaticus (Wilkinson, 1959)

Genus Cryptotermes Banks, 1 906

C. dudleyi Banks, 1918 (introduced)
C. havilandi (Sjbstedt, 1 900b)

Genus Glyptotermes Froggatt, 1 896
G. ignotus Wilkinson, 1959
G. kawandae Wilkinson, 1954
G. parki Krishna & Emerson, 1962
G. parvulus (Sjbstedt, 1 907b)

G. reticulatus Wilkinson, 1954
G. uelensis Coaton, 1 955

Ki

K'2, 4 T

T

K

Ui T

K

K T

T

K 2 T

U K T

T i

U K

T

U K T

U

u

u

u

Genus Bicornitermes Krishna, 1961

B. bicornis Krishna, 1961 Ui

B. spinicollis (Wilkinson, 1958) T

ERRATA

page 4 line 30

”M. sp.” add "near notandus Silvestri 1914-15”
page 5 line 6

”genus Ceratotermes Silvestri 1914-15” delete and substitute
”genus Foraminitermes Holmgren 1912”

page 5 line 7

__”C. rhinoceros (Sjdstedt, 1905) U T” correct to read

f- "£• rhinoceros (Sjdstedt, 1905) U T”

These changes are made to conform with the synonymy established by
KRISHNA K., 1963. The African genus Foraminitermes Holmgren (Isoptera,
Termitidae, Termitinae) . American Museum Novi tates 2161: 1-23.

*'

FAMILY HODOTERMITIDAE

Subfamily Hodotermitinae

Genus Hodotermes Hagen, 1853

H. mossambicus (Hagen, 1853) U K3

FAMILY RHINOTERMITIDAE

Subfamily Coptotermitinae

Genus Coptotermes Wasmann, 1896
C. amanii (Sjostedt 1911b)

C. formosanus Shiraki, 1 909 (introduced) U K2

C. sjostedti Holmgren, 1911

Subfamily Rhinotermitinae

Genus Schedorhinotermes Silvestri, 1 909

S. lamanianus (Sjostedt, 1911c) U K

S. putorius (Sjostedt 1 896) U

FAMILY TERMITIDAE
Subfamily Apicotermitinae

Genus Apicotermes Holmgren, 1912
A. holmgreni Emerson, 1956

Genus Astalotermes Sands, 1972
A. amicus Sands, 1972

A. hapalus Sands 1972 K

Genus Astratotermes Sands, 1972
A. apoenetus Sands, 1972
A. mansuetus Sands, 1972

Genus Alyscotermes Sands, 1972

A. kilimandjaricus (Sjostedt, 1907a) U

A. trestus Sands, 1972

Genus Ateuchotermes Sands, 1972
A. pectinatus Sands, 1972
A. rastratus Sands, 1972

Genus Anenteotermes Sands, 1972
A. disluctans Sands, 1972
A. improelatans Sands, 1972

Genus Aderitotermes Sands, 1972
A. fossor Sands, 1972

Genus Adaiphrotermes Sands, 1972
A. choanensis (Fuller, 1925)

U
U

K

U K

K4

^7; * 7* 7> 7\

Page 4

No. 181

Subfamily Termitinae

Genus Amitermes Silvestri, 1901
A. acinacifer Sands, 1959

A. evuncifer Silvestri, 1912a U

A. hastatus (Haviland, 1898)

A. lonnbergianus (Sjostedt, 191 Id)

A. messinae Fuller, 1922

A. sciangallorum Ghidini, 1941

A. somaliensis Sjostedt, 1927

A. spinifer Silvestri, 1914-15 U

A. stephensoni Harris, 1957

A. truncatidens Sands, 1959

A. unidentatus (Wasmann, 1897) U

Genus Microcerotermes Silvestri, 1 90 1

M. brachygnathus Silvestri, 1914-15
M. edentatus Wasmann, 1911 U

M. masaiaticus Harris, 1954
M. nemoralis Harris, 1954
M. parvulus (Sjostedt, 1911c)

M. parvus (Haviland, 1898) U

M. zuluensis Holmgren, 1913a

Genus Crenetermes Silvestri, 1912a
C. albotarsalis (Sjostedt, 1897)

C. mixtus Williams, 1962

Genus Euchilotermes Silvestri, 1914-15
E. umbraticola (Williams, 1954)

Genus Op hiotermes Sjostedt, 1924a

O. ugandensis Sjostedt, 1925 U

Genus Megagnathotermes Silvestri 1914-15
M. sp.

Genus Cubitermes Wasmann, 1906
C. falcifer Williams, 1966
C. fulvus Williams, 1 966
C. glebae (Sjostedt, 1913)

C. inclitus Silvestri, 1912b U

C. latens Williams, 1 966
C. montanus Williams, 1966
C. muneris (Sjostedt, 1913)

C. oblectatus Harris, 1958

C. orthognathus (Emerson, 1928) U

C. pallidiceps (Sjostedt, 1913)

C. sankurensis Wasmann, 1911

C. speciosus Sjostedt, 1924a U

C. tenuiceps (Sjostedt, 1913)

C. testaceus Williams, 1966 U

C. ugandensis Fuller, 1923 U

C. umbratus Williams, 1 954
C. zavattarii Ghidini, 1937

Genus Lepidotermes Sjostedt, 1924b
L. goliathi (Williams, 1954)

L. planifacies (Williams, 1954)

K

K

T

K T

K T Z

Ka T

K

K T

K.

T

K T Z

T

T

K T

T
T

K4 T

T Z

T

K T

K T

K 3

T

T

K T

K T

T
T

K T

T

K T

T
T

T

K

K T

K T

K T

T

T

>

No. 181

Page 5

Genus

Genus

Genus

Genus

Genus

Genus

Genus

Genus

Genus

Genus

Genus

Genus

Noditermes Sjostedt, 1924a
N. wasambaricus Williams, 1 954 K

Basidentitermes Holmgren, 1912

B. amicus Harris, 1936

B aurivillii (Sjostedt, 1897) U

Ceratotermes Silvestri, 1914-15

C. rhinoceros (Sjostedt, 1 905) U

Angulitermes Sjostedt, 1924a
A. truncatus Sjostedt, 1926a U K

Promirotermes Silvestri, 1914-15
P. bellicosi (Wasmann, 1912) K4

P. gracilipes Schmitz, 1917 U K

P. massaicus (Sjostedt, 1 907a) K4

Pericapritermes Silvestri, 1914-15
P. dumicola Harris, 1 954
P. gloveri Harris, 1951

Subfamily Macrotermitinae

Acanthotermes Sjostedt, 1898
A. acanthothorax (Sjostedt, 1898)

Pseudacanthotermes Sjostedt, 1924a
P. grandiceps (Sjostedt, 1915)

P. militaris (Hagen, 1 858)

P. piceus (Sjostedt, 1911a)

P. spiniger (Sjostedt, 1 900a)

Synacathotermes Holmgren, 1910
S. zanzibarensis (Sjostedt, 1915)

Allodontermes Silvestri, 1914
A. morogorensis Harris, 1936

Macrotermes Holmgren, 1 909
M. bellicosus (Smeathman, 1781)

M. falciger (Gerstacker, 1891)

M. herus (Sjostedt, 1914a)

M. michaelseni (Sjostedt, 1914a)

M. subhyalinus (Rambur, 1842)

M. ukuzii (Fuller, 1922)

M. vitrialatus (Sjostedt, 1 899)

Odontotermes Holmgren, 1912
O. amanicus Sjostedt, 1925
O. amaniensis (Sjostedt, 1924b)

O. anceps (Sjostedt, 191 le)

O. a polio (Sjostedt, 1905)

O. aurora (Sjostedt, 1904)

O. badius (Haviland, 1 898)

O. boranicus (Ghidini, 1937)

O. caffrariae (Sjostedt, 1897)

O. culturarum Sjostedt, 1924c

U

K2

U K

U

U K2

K

U K

U K

K

K3

U K

K.i

U K

K

U Ka

T

T

T

T

T

T

T

T

T 1
T

T.

T Z

Page 6

No. 181

O. egregius Sjostedt, 1925
O. elgonensis (Sjostedt, 1925)

O. fallax Sjostedt, 1 924c
O. fulleri (Emerson, 1928)

O. kibarensis (Fuller, 1923)

0. latericius (Haviland, 1898)

O. mediocris (Sjostedt, 191 le)

0. microps (Sjostedt, 1899)

O. monodon (Gerstacker, 1891)

O. montanus Harris, 1 960
O. nolaensis Sjostedt, 1924c
O. palmquisti (Sjostedt, 1 907a)

O. patruus (Sjostedt, 1913)

O. ramulosus (Sjostedt, 1 904)

O. rectanguloides Sjostedt, 1924c
O. stercorivorus (Sjostedt, 1 907a)
O. tanganicus Sjostedt, 1924c
O. terricola (Sjostedt, 1 902a)

O. transvaalensis (Sjostedt, 1 902b
O. zambesiensis (Sjostedt, 1914b)

Genus A ncistrotermes Silvestri, 1914-15
A. cavithorax (Sjostedt, 1899)

A. equatorius (Harris, 1966b)

A. latinotus (Holmgren, 1912)

A. microdens (Harris, 1966b)

Genus Microtermes Wasmann, 1902
M. alluaudanus (Sjostedt, 1915)

M. kasaiensis (Sjostedt, 1913)

M. luteus (Harris, 1954)

M. magnocellus (Sjostedt, 1915)

M. redenianus (Sjostedt, 1904)

M. usambaricus Sjostedt, 1926b
M. vadschaggae (Sjostedt, 1907a)

K

K

K

U Ks

U Ks

U

K

U K

K

U K.3

K

Ui Ki

U K.

K.3

K

K

U

K.3,4

u

u

U K

U K

K

U. Ki
K2

K4

Subfamily Nasutitermitinae

Genus Nasutitermes Dudley, 1 890

N. arboreus (Smeathman, 1781) U

N. chapini Emerson, 1928 U

N. elegantulus (Sjostedt, 1911b) Ui

N. fulleri Emerson, 1928 U

N. incurvus (Sjostedt, 1924b) U

N. infuscatus (Sjostedt, 1902c) K

N. kempae Harris, 1954 K

N. latifrons (Sjostedt, 1896) U

N. maculiventris (Sjostedt, 1904) U

N. schoutedeni (Sjostedt, 1924b) U

N. torquatus (Sjostedt, 1 924c) U

Genus Rhadinotermes Sands, 1965a
R. coarctatus (Sjostedt, 1902b)

Genus Grallatotermes Holmgren, 1912

G. africanus Harris, 1954 K

Genus Trinervitermes Holmgren, 1912

T. auriterrae Sjostedt, 1926c U

T
T
T 2
T
T

T

T

T

T

T

T

T.

T

T

T

T

T

T

T

T

T

T

T

T

Z

Z.

N N

No. 181

Page 7

T. bettonianus (Sjostedt 1905)

U

K

T

T. carbonarius Sjostedt, 1926d

U

T. dispar (Sjostedt 1902d)

U

K

T

T. ebenerianus Sjostedt 1926e

u

T. geminatus (Wasmann, 1897)

u

T. gratiosus (Sjostedt 1924b)

u

K

T

T. lutzi (Emerson, 1928)

u

T. occidentalis (Sjostedt 1904)

u

T. oeconomus (Tragardh, 1904)

u

T. rapulum (Sjostedt 1904)

T

T. ruficeps (Holmgren, 1913a)

T

T. segelli (Sjostedt 1907a)

T

T. togoensis (Sjostedt 1899)

u

T. trinervius (Rambur, 1 842)

u

Genus Mimeutermes Silvestri, 1914-15

M. binghami Sands, 1968 K

M. majusculus Sands, 1965b T

Z

Z

ACKNOWLEDGEMENTS

The authors are grateful for the help of K.M. Shepherd of the Tropical Development and
Research Institute (formerly Centre for Overseas Pest Research), London, for help in tracing some
of the references.

REFERENCES

BANKS, N. 1906. Two new termites. Ent News 17: 336-337.

1918. The termites of Panama and British Guiana Bull. Amer. Mus. Nat. Histo. 38:659-667.

BOUILLON, A. 1970. Termites of the Ethiopian Region. Ch. 5 : 153-280 In: Biology of Termites. I. K.

Krishna & F.M. Weesner, eds. Academic Press.

BOUILLON, A. & G. MATHOT 1965. Quel est ce termite africain? Zooleo 1:1-1 14. Leopoldville.
BOUILLON, A. & G. MATHOT 1966. Quel est ce termite africain? Zooleo Suppl. 1:1-23. Kinshasa.
BUXTON, R.D. 1979. The role of termites in the ecology of Tsavo National Park, Kenya D. Phil, thesis.
University of Oxford.

COATON, W.G.H. 1949. Notes on some South African species of the families Hodotermitidae and Kaloter-
mitidae. J. Ent. Soc. Sth. Afr. 12:13-77.

1955. New Isoptera from Belgian Congo. (With redescriptions of some named species). J. Ent.

Soc. Sth. Afr. 18:109-136.

DUDLEY, P.H. 1890. Termites of the Isthmus of Panama Part II. Trans. N.Y. Acad. Sci. 9:157-180.
EMERSON, A.E. 1928. Termites of the Belgian Congo and the Cameroon. Bull. Am. Mus. Nat. Hist.
57:401-574.

1956. Ethospecies, ethotypes, taxonomy and evolution of Apicotermes and Allognathotermes

(Isoptera: Termitidae). Am. Mus. Novit. 177 LI -31.

FROGGATT, W.W. 1896. Australian Termitidae. Part I. Proc. Linn. Soc. New South Wales 21:510-552.
FULLER, C. 1922. The termites of South Africa. S. Afr. J. Nat. Hist. 3:70-131

1923. Two new termites from Uganda Ann. Mag. Nat. Hist. 11.-191-192.

1925. The termites of South Africa III. S. Afr. J. Nat. Hist 5:167-246.

GERST ACKER, A. 1891. Die Von Hern Dr. Fr. Stuhlmann in Ostafrika gesammelten Term i ten, Odonaten
und Neuropteren. Jb. Hamb. Wiss. Anst. 9:183-191.

GHIDINI, G.M. 1937. Missione del Prof. Edoardo Zavattari nel paese del BoranaNuove specie di Termitidi.

(Diagnose Preventive) Bol. Soc. Ent Ital. 69:141-143.

1941. Nouve specie di termitide. Bol. Soc. Ent Ital. 73:30-34.

HAGEN, H.A. 1 853. Hr. Peters bench tete liber die von ihm gesammelten und von Hm Dr. Hermann Hagen
bearbeiteten Neuropteren aus Mossambique. Bericht Konigl. Preuss. Akad.Wiss. Berlin. 479-481.

Page 8

No. 181

1858. Monographic der Termiten, Part 2. Linn Entomol. 12:1-342.

HARRIS, W.V. 1936. A list of the termites of Tanganyika Territory, with descriptions of two new species.
Bull. Ent. Res. 27:361-368.

1948. Termites of the Uganda Protectorate. Proc. R. Ent. Soc. Lond. (B) 17:73-83.

1951. Further records of East African termites. Proc. R. Ent. Soc. Lond. (B) 20:25-28.

1954. Further records of East African Termites. II. Proc. R. Ent. Soc. Lond. (B) 23:127-137.

1957. Isoptera. Report on the British Museum Expedition to South-West Arabia 1937-38

1(28): 42 1-433.

1958. Isoptera. Explor. Parc Natn. Upemba. I. Mission G.F. de Witte. 52:3-26

— — 1960. Further records of East African termites - III. Proc. R. Ent. Soc. Lond. (B), 29:17-21.

1 966a. On the genus Coptotermes in Africa (Isoptera: Rhinotermitidae). Proc. R. Ent. Soc. Lond.

(B) 35:161-171.

— — 1966b. The genus Ancistrotermes (Isoptera). Bull. Br. Mus. Nat. Hist. (Ent.) 18:1-20.

HAVILAND, G.D. 1898. Observations on termites with descriptions of new species. J. Linn. Soc. (Zool.)
26:358-442.

HOLMGREN, N. 1909. Termitenstudien. 1 Anatomische Untersuchungen. K. Svenska VetenskAkad. Handl.
44:1-25.

1910. Das System der Termiten. Zool. Anz. 35:284-286.

1911. Termitenstudien 2. Systematik der Termiten. Die Familien Mastotermitidae, Protermitidae

and Mesotermitidae. K. Svenska VetenskAkad. Handl. 46:1-88.

1912. Termitenstudien 3. Systematic der Termiten. Die Familie Mertatermitidae. K. Svenska

VetenskAkad. Handl. 48:1-66.

1913a. Termiten aus Natal und den Zululande. Gesammelt von Dr. Ivar Tragardh. Ent. Tidskr.

Arg. 34:321-366.

1913b. Termitenstudien 4. Versuch einer Systematise hen Monographic der Termiten der Orien-

talischen Region. K. Svenska VetenskAkad. Handl. 50:1-276.

KEMP P.B. 1955. The termites of North-eastern Tanganyika, their distribution and biology. Bull. Ent. Res.
46:1 13-135.

KRISHNA K. 1961. A generic revision and phylogenetic study of the family Kalotermitidae (Isoptera). Bull.
Am. Mus. Nat. Hist. 122:309-408.

& A.E. EMERSON 1962. New species of the Genus Glyptotermes (Froggatt) from the Papuan,

Oriental, Ethiopian and Neotropical Regions (Isoptera, Kalotermitidae). Am. Mus. Novit. 2089:65.
RAMBUR, J.P. 1842. Histoire naturelle des Insectes. Nevropteres. p534. Paris.

RUELLE, J.E. 1970. A revision of the termites of the genus Macrotermes from the Ethiopian region (Isop-
tera: Termitidae). Bull. Br. Mus. Nat. Hist. (Ent.) 9:365-444.

1977. Macrotermes michaelseni (Sjostedt) a new name for Macrotermes mossambicus (Hagen)

(Isoptera: Termitidae). J. Ent. Soc. Sth. Afr. 40:119.

SANDS,W.A. 1957. A revision of the East African Nasutitermitinae (Isoptera). Bull. Br. Mus. Nat. Hist.
(Ent.) 5:1-28.

1959. A revision of the termites of the genus Amitermes from the Ethiopian region, (Isoptera,

Termitidae, Amitermitinae). Bull. Br. Mus. Nat. Hist. (Ent.) 8:127-156.

1965a. A revision of the termite subfamily Nasutitermitinae (Isoptera, Termitidae) from the

Ethiopian region. Bull. Br. Mus. Nat. Hist.(Ent.)Suppl. 4:172pp.

1965b. A new species of Mimeutermes Silvestri and the imago of Nasutitermes diabolus (Sjostedt)

from Africa (Isoptera: Nasutitermitinae). Proc. R. Ent. Soc. Lond. (B) 34:132-136.

1968. New species and records of Nasutitermitinae (Isoptera:Termitidae) from Africa. Proc. R.

Ent. Soc. Lond. (B) 37:163-169.

1972. The soldierless termites of Africa (Isoptera: Termitidae). Bull. Br. Mus. Nat. Hist. (Ent.)

Suppl. 18:1-244.

SCHMITZ, H. 1917. Drei neue Termiten vom belgischen Kongo. Tijdschr. Ent. 60:225-231.

SHIRAKI, T. 1909. [On the Japanese termites.] Trans. Ent. Soc. Japan 2:229-242.

SILVESTRI, F. 1901. Nota preliminare sui Termitidi sud-americani. Boll. Mus. Zool. Anat. Comp. Torino

16:1-8.

1909. Isoptera. In: Die Fauna Sudwest-Austral. 2;279-314. W. Michaelsen & R. Hartmeyer, eds.

1912a. Termiti raccolte da L. Fea alia Guinea Portoghese e alia isole S. Thome, Annobon, Prin-
cipe e Fernando Poo. Ann. Mus. Civ. Stor. Nat. Genova 5:211-255.

1912b. Termitidi raccolte da S.A.R. la Duchessa d’Aosta nelle regione dei grandi laghi dell’ Africa

equatoriale. Ann. Mus. Zool. Univ. Napoli 3:1-5.

1914. Zoological Results of the Abor Expedition, 1911-12. 32. Termitidae. Rec. Indian Mus.

Calcutta 8:425-435.

No. 181

Page 9

1914-15. Contribuzione alia conoscenza dei Termitidae e Termitofili dell' Africa occidentale. 1.

Termitidi. Boll. Lab. Zool. Gen. Agr. R. Scuola Agric. Portici 9:1-146.

1918. Contribuzione alia conoscenza dei Termitidi e Termitofili dell' Africa occidentale. Boll. Lab.

Zool. Gen. Agr. R. Scuola Agric. Portici 12:287-346.

SJOSTEDT, Y. 1896. Termiten aus Kamerun. Ent. Tidskr. 17:297-298.

1897. Neue Termiten aus West-Africa (Vorlaufige Mitteilung) Ent. Tidskr. 18:123-126.

— 1898. Zwei neue Termiten aus der Westkuste Afrikas: Ent. Tidskr. 19:204-205.

1899. Neue afrikanische Termiten (Vorlaufige Mitteilung). Ent. Nachr. 25:34-39.

1900a. Vorlaufige Diagnosen einiger afrikanischen Termiten. Ent. Tidskr. 20:278.

1900b. Monographic der Termiten Afrikas. K. Svenska VetenskAkad. Handl. 34:1-236.

1902a. Termites novos ex Africa Reportatos. Ent.Tidskr. 23:40.

1902b. Eine neue termite aus kamerun. Ent. Tidskr. 23:252.

1902c. Neue afrikanische Termiten. Ent. Tidskr. 23:302-304.

1904. Monographic der Termiten Afrikas. K. Svenska VetenskAkad. Handl. 38:1-120.

— 1905. Ueber eine Termitensammlung aus Kongo und anderen Teilen von Africa. Ark. Zool. 2:1-

20.

1907a. Termitidae. Wiss. Ergebn. schwed. zool. Exped. Kilimandjaro - Meru, 1905-6. 3:1-24.

I 9C7b Ueber eine Termitensammlung aus Kongo. Ent. Tidskr. 28:233-250.

1911a. Neue Ost- and Westafrikanische Termiten. Ent. Tidskr. 32:173-188.

— 1911b. Neue afrikanische Termiten im Schwedischen Reichsmuseum. Ark. Zool. 7:1-18.

1911c. Zur Termitenfauna Kongos. Ent. Tidskr. 3 2:137-170.

191 Id. Ueber die von Herrn Professor Einar Lonneberg in Britisch Ostafrika eingesammelten

Termiten. Ark. Zool. 7:1-3.

191 le. Termitidae novae a Cl. Dom. Maurice de Rothschild ex Aethiopia reportatae. Ent. Tidskr.

32:171-172.

1913. Uber Termiten aus dem inneren Congo, Rhodesia, und Deutsch-Ostafrika. Rev. Zool. Afr.

2:354-391.

1914a. Isoptera. In. Beitr. Kennt. Land und Susswasserfauna Deutsch-Sudwest Afrikas 1:73-92.

W. Michaelsen, ed.

1914b. Termiten aus Zambesi, Rhodesia, Nyassa und Sud-Nigeria. Ark. Zool. 8:1-9.

1915. Insectes Pseudonevropteres. 1. Termitidae. Voyage Ch. Alluad et R. Jeannel en Afrique

Orientale 191 1-12:1-18.

1924a. Neue Arten und Gattungen afrikanischer Termiten. Rev. Zool. Afr. 12:253-257.

1924b. Neue Termiten im Kongo-Museum. Rev. Zool. Afr. 12:39-42.

1924c. Neue Afrikanischer Termiten. Rev. Zool. Afr. 12:490-497.

1925. Neue Termiten aus Afrika und Madagaskar. Konowia 4:53-55.

1926a. Termiten aus Accra. Ent. Tidskr. 47:238-246.

1926b. Revision der Termiten Afrikas. 3. Monographic. K. Svenska VetenskAkad. Handl. 3:1-419.

1926c. Neue Termiten aus der Goldkiiste. Ark. Zool. 18:1-5.

1 926d. Kongo-termiten aus dem Ituri- und Uelegebiet. Rev. Zool. . Afr. 14:140-164.

1926e. Wissenschaftliche Ergebnisse der mit Unterstiitzung der Akademia der Wissenschaften in

Wien aus der Erbschaft Treitl von F. Werner Unternommenen zoologischen Expedition nach dem
Anglo-Agyptischen Sudan (Kordofan) 1914. 21. Isoptera. Akad. Wiss. Wien Math. -Nat. 100:71-76.
1927. Termiten aus dem Somaliland. Rev. Zool. Afr. 15:97-104.

SMEATHMAN, H. 1781. Some account of the termites which are found in Africa and other hot climates.
Phil. Trans. R. Soc. 7 LI 39-1 92.

SNYDER, T.E. 1949. Catalog of the Termites (Isoptera) of the world. Smithson. Misc. Coll. 112:1-490.
TRAGARDH, I. 1904. Termiten aus dem Sudan. Results Swed. Zool. Exped. Egypt and White Nile, 1901.
12:1-47.

WASMANN, E. 1896. Neue Termitophilen und Termiten aus Indien. Viaggo di Leonardo Fea in Birmania e
Regioni Vicine. Ann. Mus. Civ. Stor. Nat. Genova 16:613-630.

1897. Termiten von Madagaskar und Ostafrika (Wiss. Ergeb. der Reisen in Madagaskar und

Ostafr. 1889-95 von Dr. A. Voeltzkow.) Abh. Senckenb. Naturforsch. Ges. 21:137-182.

1902. Termiten, Termitophilen und Myrmecophilen gesammelt auf Ceylon von Dr. W. Horn.

1899 mit andern ostindischen Material. Zool. Jahrb. Syst. 17:99-164.

1906. Beispiele rezenter Artenbildung bei Ameisen und Term itengasten. Biol. Zentralbl. 26:565-

580.

1911. Zur Kenntnis der Termiten und Termitengaste von Belgischen Kongo. Rev. Zool. Afr.

L91-117, 145-176.

1912. Neue Beitrage zur Kenntnis der Termitophilen und Myrmecophilen. Z. Wiss. Zool., Leip-
zig. 101:70-115.

Page 10

No. 181

WILKINSON, W. 1954. Three new species of Kalotermitidae (Isoptera) from East Africa. Proc. R. Ent. Soc.
Lond. (B) 23:75-82.

1958. Two new species of Kalotermes Hagen from West Africa. Proc. R. Ent. Soc. Lond. (B)

27:109-1 15.

1 959. Four new species of Kalotermitidae from East Africa (Isoptera). Proc. R. Ent. Soc. Lond.(B)

28:61-72.

WILLIAMS, R.M.C. 1954. New East African Termitinae (Isoptera: Termitidae). Proc. R. Ent. Soc. Lond. (B)
23:215-227.

1962. A correction concerning two East African Termitinae (Isoptera). Proc. R. Ent. Soc. Lond.

(B) 3L127-130.

— — 1966. The East African termites of the genus Cubitermes (Isoptera: Termitidae). Trans. R. Ent.

Soc. Lond. 118:73-118.

Received 5 January 1984
Editors: J.J. Hebrard, D. Widdowson

No. 181

Page 11

NOTICE TO CONTRIBUTORS

Papers on natural history subjects will be considered on the understanding that these are being offered
exclusively to the Society and have not been submitted, at the same time, to any other journal.

They should be addressed to the Editors, Journal of the East Africa Natural History Society and National
Museum, P.O. Box 44486, Nairobi, and be accompanied by a covering letter from the author responsible for
correspondence and decisions regarding the typescript.

Two copies of the contribution should be forwarded to the Society; a further copy should be kept by the
author for reference. The typescript should be on A4 size paper. Use one side only, double-spaced with margins
of at least 5 cm at the top and left-hand side of the pages to allow for editorial instruction to printers. The
pages must be numbered and all measurements are to be in metric units.

The title page should contain the title of the article followed by the name of the author(s) and institution(s)
where the research was carried out. A present address, if applicable, should be included as a footnote.

Abstract to be on the second page and contain not more than 150 words. State the purpose of the study,
procedures, main findings and conclusions. This should be intelligible in itself without reference to the paper.

Nomenclature In cases where there are well known and accepted English names, e.g. for birds and larger
mammals, these should be used throughout the text, accompanied by the scientific name on first mention.
Authorities should be given beside the generic and specific names, when quoted for the first time, forall groups,
with the exception of ornithological papers not dealing with taxonomic discussion.

Text This is usually, but not necessarily, divided into parts with the headings Introduction, Methods, Results
and Discussion, these being broken up with sub-headings where necessary. Main heads should be centred on
the page and typed in capital letters. Subheads should be ranged to the left margin and typed in capitals. Any
further headings should start with a capital letter followed by small letters on the left margin. No headings
should be underlined. Underlining is an instruction to the printer to use italic type and should be reserved
for all generic and species names, foreign words not adopted into English and for descriptions of bird. or
animal vocalisation.

Acknowledgements As acknowledgements may imply endorsement of the data and conclusions contained in
the authors work, it is advisable to obtain permission before quoting persons who have made contributions to
the study.

References Care should be taken to see that all references quoted in the text are given in full, in this section.
To avoid confusion, references to books and periodicals should not be abbreviated and neither should they be
underlined. For example:

(in text) Cave & Macdonald (1955) or (Cave & Macdonald 1955)

(in reference list) CAVE, F.O. & MACDONALD, J.D. 1955. Birds of the Sudan, Oliver & Boyd, London.

Tables Each table must be typed, double-spaced on a separate sheet and numbered. Leave good spaces
between columns and omit internal vertical and horizontal rules. Cite tables in the text in consecutive order.

Illustrations Submit two sets of figures which should be well, drawn on bristol board or heavy tracing paper
with a photographed or xeroxed copy. Lettering must be of first quality (e.g. lettraset) and consideration must
be given to size when reduced. Typewritten or freehand lettering will not be accepted unless put onto an overlay
which can be followed by the printers for typesetting. Each figure should have a label pasted on its back giving
the number of the figure, caption details and accreditation. Cite each figure in the text in consecutive order. A
separate caption sheet should be added to the typescript. Photographs are acceptable if they have good definition
and contrast and should be produced on glossy paper.

Proofs The author will be sent one set of galley proofs for correction of typographical errors. Authors may
be charged for any changes from the original typescript made by him at this stage, and which result in additional
printing expense. Plates or page proofs will not be sent.

Published material The author is entitled to 25 copies free of charge. Further copies can be supplied at cost
price, provided the order is placed when the typescript is submitted. If there is more than one author, 30 copies
will be sent to the corresponding author for distribution.

The society is supported entirely by membership subscriptions and may not, for financial reasons, be able
to accept all submissions of value.

Wherever possible an author should seek a grant, from his institution or other source, to help with costs of
publication.

Page 12

No 181

SMITHSONIAN INSTITUTION LIBRARIES

9088 01313 752

Published by The East Africa Natural History Society, Box 44486, Nairobi, Kenya. Phototypeset-
ting by Kul Graphics Ltd., P.O. Box 18095 and printed by Manhattan Displays, P.O. Box 30434,
Nairobi.