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SECOND  SERIES:   PULMONATA. 


MANUAL 


OF 


J 


STRUCTURAL  AND  SYSTEMATIC. 


WITH  ^LUSTRATIONS  OE  THE  SPECIES. 


BY  GEORGE  W,  TRYON,  JR. 

CONTINUED    BY 

HENRY  A.  PILSBRY. 

CONSERVATOR  OF  THE   CONCHOLOGICAL   SECTION    OF   THE    ACADEMY   OF 
NATURAL   SCIENCES   OF   PHILADELPHIA. 


Vol. 

HELICID^E,  Vol.  V. 


PHILADELPHIA  : 
Published  by  the  Conehologieal  Section; 

ACADEMY  OF  NATURAL  SCIENCES,  COR.  igTH  AND  RACE  STS. 

1891. 


MANUAL  OF  CONCHOLOGY. 


FAMILY  HELICIDsE,  Vol.  V. 


Genus  HELIX,  Linne,  (Concluded.) 


Sub-genus  PAPUINA  Martens. 

Geotrochus  BECK,  (in  part)  Index  p.  47,  1837,  not  o/v.  HASSELT, 
Algemeene  Konst-en-letterbode  voor  het  jaar  1823,  p.  233  ;  see  also, 
Bull,  des  Sci.  Naturelles  et  de  Geol.  iii,  Paris,  1824,  p.  83,  paragraph 
9  ;  and  also,  Ostas.  Zool.,  Landschn.  p.  245.— Geotrochus  of  authors 
generally.— Papuina  v.  MART.,  in  Albers,  Die  Heliceen,  2d.  Edit, 
p.  166,  1861.  Type  H.  lituus  Less. — Eugenia  ALBERS  Mss. — 
Insularia  TAPPARONE  CANEFRI,  Ann.  Mus.  Civ.  di  Storia  Nat.  di 
Geneva,  xix,  p.  115, 138  ;  1883,  type  H.  lituus— Pileolus  LESSON, 
Voy.  de  la  Coquille,  Zool.  ii,  p.  313,  first  species  H.  tu/etii  Less., 
H.  gaberti  also  included ;  (preoc).—  Cymotropis  v.  MART.,  Die  Hel. 
p.  169,  type  "  H.  vitrea  "=H.  antrorsa  Pfr.—Merope  ALBERS,  Die 
Heliceen,  2d  edit.  p.  158,  type  H.  fringilla  Pfr.  (name  preoc.  in 
Mactridce — Acavus  of  E.  A.  SMITH  and  TAPPARONE  CANEFRI,  not 
Acavus  MONTF. 

There  can  be  no  question  that  v.  Hasselt's  group  Geotrochus  was 
founded  on  species  of  Trochomorpha  or  Trochonanina.  "  Under  the 
generic  name  Geotrochus,  I  group  three  little  shells,  agreeing  with 
Trochus  in  form  of  the  shell,  but  with  Helix  in  the  animal."  The 
species  obtusus,  turriculatus  and  zonatus  are  enumerated  by  v.  Hasselt. 
The  absence  of  true  Papuina  in  Java,  and  the  abundance  of  Trocho- 
nanina at  the  localities  visited  by  Hasselt,  cause  Dr.  v.  Martens  to 
surmise  that  he  had  before  him  Helix  winteriana  and  conus  or  allied 
species.  Swainson  and  many  others  have  used  the  name  Geotrochus 
for  H.  pileus  and  allies,  and  it  is  an  unfortunate  circumstance  that 
the  change  is  necessary. 

Shell  turbinate  or  trochiform,  imperforate  or  umbilicate,  rounded 
at  the  periphery  or  carinated.  Surface  smooth  or  variously 
sculptured,  not  hirsute.  Aperture  oblique  ;  peristorne  expanded. 


4  HELIX-PAPUINA. 

The  species  are  illustrated  on  plates  1  to  17. 

No  exact  diagnosis  can  be  framed  for  a  group  in  which  such 
diverse  forms  occur  as  in  Papnina.  It  is  still,  however,  an  easy 
matter  to  recognize  a  species  as  belonging  here ;  for  with  all  its 
variety,  the  group  is  a  very  natural  one,  well  differentiated  from 
other  branches  of  the  Helix  stock.  The  central  point  for  the  sub- 
genus,  geographically,  is  New  Guinea ;  the  species  extending  eastward 
to  the  Louisiade,  Admiralty  and  Solomon  Islands,  with  a  few  in  the 
New  Hebrides,  and  northwestward  into  the  Moluccas.  The  distribu- 
tion is,  therefore,  practically  the  same  as  that  of  the  subgenus 
Chloritis  (-{-  Planispira,  etc.) 

In  a  previous  volume  the  writer  has  separated  the  South  American 
group  Oxyckona  from  Papuina.  It  remains  to  sunder  the  species  of 
southeastern  Asia  and  the  zoologically  dependent  islands,  w7hich  will 
be  discussed  after  the  true  Papuina-species.  The  latter  form  a 
moderately  homogeneous  group,  in  which  it  is  practically  impossible 
to  establish  "  sections  "  to  correspond  to  the  various  names  given  in 
the  above  synonymy.  It  seems  to  me  a  better  plan  to  form  the 
species  into  a  number  of  groups,  each  typified  by  its  more  prominent 
specific  type.  These  groups  are  of  somewhat  unequal  value,  and 
bear  various  degrees  of  affinity  to  each  other ;  but  they  undoubtedly 
form  a  basis  for  a  natural  system.  It  is  likely  that  anatomical  char- 
acters will  afford  a  good  clue  to  the  minor  groups. 

I.   True  PAPUINA  ;   including  species  of  the  Moluccas,  New  Guinea, 
Australia,  Admiralty,  Solomon,  Louisiade  Is.,  etc. 

(1.)  GROUP  OF  H.  BOIVINI. 

Imperforate,  turbinate,  the  periphery  rounded  or  bluntly  angled ; 
upper  lip  not  expanded  or  but  narrowly  expanded  ;  banded.  Sur- 
face smooth,  lightly  obliquely  striatulate,  sometimes  having  faint  spiral 
lines,  but  no  forward-descending  wrinkles. 

H.  CONGRUA  Pfeiffer.     PI.  9,  fig.  72. 

Imperforate,  trochiform,  rather  solid,  striatulate,  flesh-colored, 
variou.-ly  painted  with  brown  bands;  spire  subcon cave-conic,  apex 
ii  little  obtuse.  Whorls  6,  a  little  convex,  the  last  irregularly 
in  Hated,  slightly  descending,  distinctly  rib-striated  ;  base  nearly  fiat, 
more  shinin.L:.  Aperture  oblique,  subtetragonal-oval ;  peristome 
blackish-chestnut,  expanded,  encircled  by  a  streak  outside,  margins 


HELIX-PAPUINA.  O 

joined  by  a  blackish  callus  ;  the  right  margin  dilated,  columellar 
margin  wide,  complanate. 

Alt.  24,  greater  diam.  29,  lesser  25  mill.     (Pfr.) 

Admiralty  Is. 

H.  congrua  PFR.,  P.  Z.  S.  1858,  p.  21,  t.  40,  f.  7  ;  Mai.  Bl.  1859 
p.  14  ;  Monogr.  v.  p.  269. 

H.  CHANCEI  Cox.     PI.  14,  fig.  70. 

Shell  imperforate,  conoid,  faintly  striated  from  above  downwards, 
with  straight  striae,  and  longitudinally  striated  with  more  distinct  wavy 
striae,  white,  ornamented  on  the  center  of  the  whorls  with  a  broad 
brown  band,  and  around  the  base  with  a  broad  brown  zone  ;  spire 
pyramidal,  black  at  the  apex  ;  whorls  six,  the  last  rapidly  increasing 
in  size  and  inflated,  suddenly  deflected  in  front,  and  contracted 
behind  the  lip  ;  aperture  large,  irregularly  ovately  rounded,  margins 
approaching,  lip  slightly  thickened  and  reflected ;  collumellar 
margin  moderately  dilated  and  excavated,  and  darkened  by  a  black 
callus.  ( Cox.} 

Alt.  31,  greater  diam.  33,  lesser  27  mill. 

Ysabel,  Solomon  Is. 

H.  chancei  Cox,  P.  Z.  S.  1870,  p.  171,  t.  16,  f.  5. 

Resembles  closely  the  following  species  in  coloring,  but  is  very 
differently  proportioned. 

H.  AMPHIZONA  Pilsbry.     PI.  8,  figs.  52,  53,  54. 

Imperforate  ;  turbinate-conic,  the  spire  elevated,  apex  obtuse ; 
whorls  6J,  the  last  rounded  at  the  periphery,  descending  in  front. 
Surface  having  light  oblique  growth-lines,  and  very  close,  fine,  cren- 
ulated  spiral  striae.  Color  creamy-white  at  suture  and  periphery, 
the  base  and  a  wide  band  above  dark  brown  ;  earlier  whorls  purple- 
black. 

This  is  a  large  solid  shell ;  the  spire  is  conical,  whorls  somewhat 
convex,  the  earlier  three  or  four  of  a  dark-reddish  or  purple-black 
color.  The  last  whorl  has  no  trace  of  a  carina.  The  aperture  is 
very  oblique,  banded  inside ;  peristome  well  expanded,  reflexed,  the 
dark  bands  extending  upon  it  half-way  to  the  edge,  the  outer  portion 
being  white  ;  the  columella  and  parietal  wall  are  dark  brown. 
Columella  wide,  adnate,  flat,  its  inner  edge  somewhat  straightened, 
arcuate  and  deeply  entering  at  the  insertion. 

Alt.  34  greater  diam.  30,  lesser  23  mill. 

Alt.  32  greater  diam.  29,  lesser  23  mill. 

Solomon  Is. 


HELIX-PAPUINA. 


Like  H.  chancel  in  coloring,  but  so  different  in  proportions  that  I 
do  not  feel  justified  in  uniting  them. 

Differs  from  H.  pileus,  H.  lenta  and  H.  euchroes  in  being  im- 
perforate  at  all  stages  of  growth,  a  young  specimen  before  me  having 
the  slender  itxis  completely  solid.  The  specimens  before  me  are 
three  in  number.  They  were  presented  to  the  Academy  by  John 
H.  Campbell  Esq.,  of  Philadelphia.  Figures  52,  53  represent  a 
perfect  specimen.  The  original  of  fig.  54  is  somewhat  worn,  making 
the  upper  band  appear  more  broken  and  narrower  than  it  naturally 
is.  I  give  the  locality  with  much  doubt,  as  I  do  not  know  the 
history  of  the  specimens. 

H.  BOIVINI  Petit.     PI.  6,  figs.  13,  14,  15,  16,  19,  20. 

Imperforate,  turbinate,  smooth,  the  periphery  rounded ;  white, 
banded  with  blackish-brown  ;  lip  edged  with  bright  orange-red. 

The  spire  is  conical,  apex  obtuse  ;  whorls  6,  slightly  convex,  the 
last  rounded  at  the  periphery,  slightly  descending  in  front.  Surface 
Btriatulate,  smooth,  shining.  Color  white  or  cream,  conspicuously 
banded  with  blackish-brown  bands,  of  which  one  above  and  one 
below  the  periphery  are  most  constant  (there  being  also,  usually,  a 
band  a  short  distance  below  the  suture  and  one  or  several  on  the 
base).  Aperture  very  oblique,  banded  within  ;  peristome  narrowly 
expanded,  white,  edged  with  intense  orange.  Columella  sloping, 
straightened,  flattened,  dilated  and  closely  appressed  over  the 
umbilical  tract. 

Alt.  27,  greater  diam.  25,  lesser  22  mill. 

Alt.  20,  greater  diam.  21  mill. 

Bougainville  and,  Ysabel,  Solomon  Group. 

H.  boivini  PET.,  Kev.  Zool.  1841,  p.  184.— DESH.,  in  Fer.  Hist.  t. 
108C,  f.  7,  8.— REEVE,  f.  410.— PFR.,  Monogr.  i,  p.  230 ;  Conchyl. 
( 'ab.  p,  433,  t.  151,  f.  56. — v.  MARTENS,  Monatsber.  k.-preuss.  Akad. 
Wissensch.  zu  Berlin,  1877,  p.  276,  t.  2,  f.  11-13  (anatomy).— 
BRAZIER,  Journ.  de  Conchyl.  1880,  p.  306. — H.  subrepta  HOMBR.  & 
JACQ.,  Voy.  an  Pol  Sud,  Atlas,  t.  4,  f.  1-6. — H.  colorata  MOUSSON, 
teste  PFR.,  Monogr.  iii,  p.  174. 

Readily  recognized  by  the  brilliant  orange  edge  of  the  narrowly 
reflexed  lip,  and  the  black  bands.  There  are  often  some  reddish- 
brown  hands  also;  and  some  specimens  have  the  greater  part  of  the 
base  of  this  color. 


HELIX-PAPUINA.  7 

H.  AMBROSIA  Angas.     PI.  4,  figs.  75,  76,  77. 

Imperforate,  turbinate,  rounded  at  the  periphery,  having  a  broad- 
dark-brown  band  above  (often  split  into  two  bands)  which  is  macu- 
lated with  angular  patches  of  opaque  white  ;  base  either  unicolored 
deep  brown  or  having  a  brown  band  below  the  periphery ;  lips 
white. 

The  spire  is  conical,  apex  whitish,  obtuse;  whorls  5tr,  slightly 
convex,  the  last  a  little  descending  in  front.  Surface  smooth,  shin- 
ing, lightly  striatulate.  There  is  a  narrow  white  border  below  the 
suture,  a  wide  white  girdle  at  the  periphery,  and  usually  the  broad 
blackish-brown  band  on  the  upper  surface  is  split  into  two  by  a 
white  zone.  The  base  is  either  unicolored  blackish-brown,  or  is 
white  shading  into  brown  below  a  dark  band.  Aperture  oblique, 
subrhombic,  dark  within ;  the  face  and  reverse  of  the  narrowly  ex- 
panded lip  are  white.  The  columella  is  oblique,  wide,  subtruncate 
below. 

Alt.  24,  greater  diam.  20,  lesser  IS}  mill. 

Alt.  19,  greater  diam.  18,  lesser  16  mill. 

Galera  ("Sava}i  or  "  Russell?  s  Id"}  and  Ysabel  Is.,  Solomon 
Group. 

Geotrochus  ambrosia  ANGAS  P.  Z.  S.  1867,  p.  889,  t.  43,  f.  9,  10. — 
Helix  ambrosia  PFR.  Monogr.  vii,  p.  210. — DOHRN,  Conchyl.  Cab. 
p.|571,  t.  168,  f.  12, 13.— BRAZIER,  Journ.  de  Conchyl.  1880,  p.  302. 
—H.  ramsdeni  ANGAS,  P.  Z.  S.  1876,  p.  266,  t.  20,  f.  6,  7. 

The  pellucid  epidermis  forms  angular  spots  on  the  dark  bands, 
as  in  many  Philippine  Island  Cochlostylas. 

The  form  called  H.  ramsdeni  by  Angas  is  slightly  larger,  but  has 
no  constant  differences.  The  specimen  before  me  (typical  in  form 
and  color),  measures,  Alt.  25,  greater  diam.  24,  lesser  20  mill. 

On  PL  16,  fig.  19,  represents  Angas'  type  of  ramsdeni,  and 
fig.  16,  the  smallest  specimen  of  ambrosia  in  the  suite  of  sixteen 
specimens  before  me. 

H.  MALANTENSIS  Angas.     PI.  16,  figs.  17,  20,  21,  22. 

Imperforate,  elevated  turbinate,  the  spire  conical,  apex  obtuse ; 
whorls  5J,  slightly  convex,  the  earlier  3  unicolored  waxen-white, 
the  last  whorl  rounded  at  the  periphery,  a  trifle  descending  in  front. 
Surface  smooth,  shining,  lightly  striatulate.  Color  creamy-white, 
variously  zoned  with  wide  and  narrow  bands  of  dark  brown  and 
tawny. 


HELIX-PAPUINA. 

Aperture  oblique,  subrhombic,  banded  within;  lip  slightly 
expanded,  its  face  and  reverse  white;  columella  wide,  flat,  its  inner 
edge  slightly  arcuate,  scarcely  truncate  below. 

Alt.  24,  greater  diarn.  20 £,  lesser  18  mill. 

Alt.  22,  greater  diam.  25,  lesser  22  mill. 

Malanta  Is.,  Solomon  Group. 

H.  malantensis  ANGAS,  P.  Z.  S.  1876,  p.  488,  t.  47,  f.  1-3. 

Closely  allied  to  H.  ambrosia  var.  ramsdeni,  but  lacking  hydro- 
phanous  patches,  and  having  a  greater  number  of  bands.  It  differs 
from  H.  f/uadalcanarensis  in  being  somewhat  more  elevated,  and 
not  angled  at  the  periphery.  Figures  20,  21,  22  and  the  measure- 
ments last  given  above  are  from  Angas.  The  other  figure  repre- 
sents the  shell  before  me,  described  above. 

H.  PHILOMELA  Angas.     PI.  4,  figs.  66,  67. 

Shell  im perforate,  trochiform,  moderately  solid,  obliquely  obscurely 
striated  ;  whitish,  the  whorls  ornamented  in  the  middle  with  a  broad 
fascia  composed  of  numerous  purplish  chocolate  bands  with  a  similar 
fascia  below  the  periphery,  and  crossed  obliquely  with  close  narrow 
white  lines  corresponding  with  the  lines  of  growth ;  spire  conical, 
apex  obtuse,  purplish-black,  whorls  6,  moderately  convex,  sutures 
impressed,  last  whorl  rather  inflated  descending  in  front,  and  con- 
tracted behind  the  aperture,  base  somewhat  flattened;  aperture  sub- 
ovate,  very  oblique,  peristome  thickened  expanded  and  slightly 
reflected,  the  right  margin  a  very  little  sinuated  ;  columellar  margin 
dilated,  and  furnished  within  with  a  prominent  callus  terminating 
abruptly  within  the  aperture;  the  columella  and  the  interior  of  the 
aperture  dark  chocolate-brown,  the  inner  edge  of  the  lip  orange,  and 
the  reflected  portion  white,  immediately  behind  which  is  a  pigment- 
like  black  deposit  on  the  whorl.  (Angas.') 

Alt.  20,  greater  diam.  25,  lesser  20  mill. 

Ysabel  Island,  Solomon  Group. 

H.  philomela  ANGAS,  P.  Z.  S.  1872,  p.  610,  t.  42,  f.  2,  3. 

Differs  1'nnn  H.  gnadalcanarensis  in  the  form  of  the  columella. 

The  form  figured  by  Angas  is  said  by  Cox  to  be  the  rarest 
variety  of  this  species,  and  cannot  be  regarded  as  an  ordinary  rep- 
resentative. It  is  found  abundantly  at  the  Louisiade  and  Solomon 
Is. ;  and  the  prevailing  color  is  white,  with  a  dark  black  edge  behind 
th«-  reflexed  lip.  (See  Cox,  P.  Z.  S.  1873,  p.  566,  under  H.  re- 
f".  < 


HELIX-PAPUINA.  9 

H.  GUABALCANARENSIS  Cox.     Pl.v4,  figs.  80,  81  ;  pi.  16,  figs.  23,  24. 
Imperforate,  globose-turbinate,  the  spire  conical,    apex   obtCise, 


brownish-corneous;  whorls  5,  'somewhat  convex,  the  earlier  2  or 
uuicolored,  corneous  ;  the  last  whorl  distinctly  and  rather  acutely 
angled  in  front,  the  angle  becoming  indistinct  on  the  latter  part  of 
the  whorl  ;  toward  the  aperture  the  whorl  usually  descends  a  trifle. 
Surface  lightly  striatulate,  shining.  Color  pale  brown,  having  a 
wide  white  girdle  at  the  periphery,  and  (typically)  numerous  narrow 
brown  bands  above,  fewer  below.  Aperture  very  oblique,  large, 
more  or  less  distinctly  banded  inside  ;  lip  expanded,  white  ;  colu- 
mella  spreading  at  its  insertion. 

Alt.  22,  greater  diam.  23,  lesser  20  mill. 

Guadalcanar,  Solomon  Is. 

H.  guadalcanarentsis  Cox,  P.  Z.  S.  1871,  p.  645,  t.  52,  f.  8.  — 
DOHRN,  Contin.  Conchyl.  Cab.,  p.  586,  t.  172,  f.  12,  13. 

The  angular  periphery,  numerous  narrow  bands,  and  the  dark 
streak  just  behind  the  expansion  of  the  white  lip  are  characteristic 
marks  of  this  species.  H.  philomela  Angas  is  evidently  closely 
allied. 

Variety  (pi.  16,  figs.  23,  24).  A  color-variety  having  narrow 
bands  at  axis  and  suture  and  broad  bands  above  and  below  the 
peripheral  white  zone.  Form  typical. 

Alt.  22,  greater  diam.  21,  lesser  18  mill. 

H.  HARGREAVESI  Angas.     PI.  4,  figs.  72,  73,  74. 

*» 

Imperforate,  low-turbinate,  spire  conical,  apex  obtuse  white  ; 
whorls  5-54,  slightly  convex,  the  last  rounded  at  the  periphery, 
abruptly  descending  in  front,  but  not  descending  as  far  as  the  basal 
band.  Lightly  marked  by  growth-lines,  but  smooth  and  polished. 
White,  the  lips  and  columella  bordered  with  deep  chestnut-  brown, 
the  base  suffused  with  brown  around  the  columella  ;  and  usually 
there  is  in  addition  a  broad  brown  girdle  on  the  periphery,  a  nar- 
row7 brown  border  above  and  below  the  suture,  and  a  basal  stripe. 
There  are  some  sparsely  scattered,  small,  grayish  dots,  usually  few 
in  number  and  inconspicuous. 

The  aperture  is  very  oblique,  subrhombic,  white  and  showing  the 
band  inside.  The  upper  lip  is  not  at  all  expanded  ;  basal  lips 
slightly  expanded  ;  columellar  lip  suddenly  dilated,  spreading  in  a 


10  HELIX-PAPUINA. 

dark  callus  over  the  excavated  umbilical  region  ;  inner  edge  of  the 
columella  straightened,  thickened,  deeply  entering. 

Alt.  21,  greater  diam.  25,  lesser  2H  mill. 

Alt.  19,  greater  diam.  21,  lesser  18  mill. 

Bougainville,  Faro  and  Short  land  Is.,  Solomon  Group. 

H.  hargreavesi  ANG.,  P.  Z.  S.  1869,  p.  625,  t.  48,  f.  2.— PFR. 
Mon.  vii,  p.  310. — SMITH,  P.  Z.  S.  1885,  p.  591. — "  H.  Hargravesii 
Cox, "  PAETEL,  Catal.,  p.  90. 

The  broadly  brown-bordered  lip,  polished  surface  destitute  of  an- 
teriorly descending  wrinkles,  and  the  small  sparsely  scattered  bluish- 
translucent  dots,  are  the  more  prominent  characters  of  the  species. 
H.  gamelia  is  closely  allied,  but  it  lacks  the  dark  lip,  and  has  the 
upper  lip  more  arched  forward  ;  H.  hargreavesi  further  differs  from 
H.  gamelia  in  having  the  last  whorl  abruptly  and  considerably 
deflexed  in  front. 

H.  GAMELIA  Angas.     PL  3,  figs.  44,  45,  46,  47. 

Imperforate,  low-turbinate,  thin  ;  spire  conical,  apex  light,  obtuse; 
whorls  5-2,  slightly  convex,  the  last  either  rounded  or  subangular  at 
the  periphery,  not  descending  in  front;  surface  smooth,  lightly 
striatulate,  color  cream  white,  becoming  light  brown  on  the  latter 
half  of  the  body-whorl,  white  on  the  spire,  encircled  by  a  narrow 
dark  brown  band  bordering  the  suture  below,  a  wider  band  at  the 
periphery,  and  another  on  the  base  midway  between  axis  and  peri- 
phery;  all  visible  within  the  aperture.  Aperture  very  oblique,  sub- 
rhombic  ;  lip  white  or  light  brown,  the  columella  brown. 

The  lip  is  bordered  behind  by  a  light  and  then  a  dark  stfreak. 
The  upper  lip  is  slightly  expanded,  notably  arched  forward,  reced- 
ing toward  the  outer  angle  of  the  mouth.  The  columellar  lip  is 
spread  in  a  thin  brown  callus  over  the  impressed  umbilical  region. 
The  edge  of  the  columella  is  thickened  ;  it  is  not  truncated  at  the 
base.  Alt.  18-1  greater  diam.  21,  lesser  17$  mill.  % 

Stephen,  Ysabel,  Shortland  and  Treasury  Islands,  Solomon  Group. 

Geotrochus  gamelia  ANG.,  P.  Z.  S.  1867,  p.  888,  t.  43,  f.  1-3.— H. 
gamelia  PFR.,  Mon.  vii,  p.  313. — SMITH,  P.  Z.  S.  1885,  p.  591. 

Thinner,  less  elevated  than  II.  hargreavesi,  differently  colored,  and 
having  the  last  whorl  not  descending  to  the  aperture. 

H.  BRODIEI  Brazier.     PL  6,  fig.  9. 

Shell  imperforate,  flatly  conical,  thin,  shining,  obliquely  faintly 
and  irregularly  plicately  striated  ;  white,  with  a  chalk-white  band  at 


HELIX-PAPUINA.  11 

the  suture  and  periphery,  and  with  a  spiral  band  of  dark  brown  at 
the  base,  somewhat  diffused,  with  the  white  band  at  the  periphery ; 
keeled  ;  rather  blunt  at  the  apex  ;  whorls  5,  nearly  flat,  the  last  con- 
vex at  the  base  ;  aperture  oblique,  rhomboidally  ovate  ;  peristome 
dark  brown,  moderately  thickened,  margins  approximating,  the 
right  slightly  deflected,  the  columellar  margin  reflected  and  furnished 
with  a  straight  callus.  (Brazier.') 

Alt.  14,  greater  diam.  16,  lesser  13  mill. 

Choiseul  Island,  Solomon  Group. 

H.  brodiei  BRAZ.,  P.  Z.  S.  1872,  p.  20,  t.  4,  f.  6.— PFR.,  Mon.  vii, 
p.  301. 

Varieties  of  this  species  occur  in  which  the  dark  brown  band  is 
occasionally  found  above  the  chalk-white  one,  and  a  brown  band 
under  the  periphery.  (Braz.*) 

H.  DAMPIERI  Angas.     PL  5,  fig.  88  ;  pi.  16,  fig.  18. 

Shell  imperforate,  subglobose,  conoid,  light  brown  or  fawn  color, 
here  and  there  minutely  dotted  with  dark,  gray  specks  with  a  broad 
white  band  around  the  middle  of  the  penultimate  whorl,  and  two 
upon  the  last,  one  above  and  the  other  below  the  middle,  also  a  nar- 
row white  line  revolving  up  the  spire  beneath  the  suture,  and  a  dark 
brown  zone  surrounding  the  pale  or  yellowish  umbilical  region. 
Whorls  5,  a  little  convex  above,  somewhat  glossy,  obliquely  and 
very  finely  striated  by  the  lines  of  growth  ;  the  last  more  or  less  con- 
centrically striated  beneath,  shortly  descending  at  the  aperture, 
which  is  white  within.  Lip  a  little  thickened,  edged  with  reddish- 
brown,  only  slightly  expanded  on  the  right  side,  more  dilated  below, 
produced  into  a  thin,  transparent  callosity  over  the  umbilical  region, 
united  above  to  the  upper  extremity  of  the  peristome.  Columellar 
margin  oblique,  white  or  partly  tinged  with  reddish-brown,  thickened 
and  terminating  below  within  the  edge  of  the  lip.  Alt.  19,  greatest 
diam.  22,  smallest  19  mill.  (Smith.) 

Louisiade  Archipelago  (Angas) ;  Choiseul  Bay,  Bougainville 
Straits  (Guppy.) 

H.  dampieri  ANG.,  P.  Z.  S.  1869,  p.  47,  t.  2,  f.  6.— PFR.,  Mon., 
vii,  p.  310.— SMITH,  P.  Z.  S.  1885,  p.  592,  t.  36,  f.  5. 

The  specimens  collected  by  Guppy  and  described  by  Smith  are 
smaller  than  the  types ;  they  also  have  the  peristome  brown,  and 
the  basal  band  is  darker  (pi.  16,  fig.  18). 


12  HELIX-PAPUINA. 

H.  WALLERI  Brazier.     PI.  6,  figs.  11,  12. 

Shell  imperforate,  depressedly  trochiform,  rather  thin,  obliquely, 
irregularly  and  obscurely  striated  ;  yellowish-white,  with  two  dark 
chocolate  bands,  thickly  crossed  with  white  diaphanous  zigzag  mark- 
ings encircling  the  last  whorl,  the  uppermost  band  the  broadest ; 
spire  obtusely  conical,  apex  white ;  whorls  5,  flatly  convex,  sutures 
impressed,  last  whorl  descending  in  front,  base  a  little  tumid  ante- 
riorly ;  aperture  oblique,  subrhomboidal ;  peristome  very  slightly 
thickened  and  expanded,  and  but  little  reflexed,  the  right  margin 
moderately  sinuated ;  columellar  margin  somewhat  flattened  and 
dilated  with  a  straight  abrupt  callus ;  the  umbilical  region  and  the 
inner  edge  of  the  lip  with  the  reflected  portion  brown,  immediately 
behind  which  is  a  band  of  pigment-like  deposit  of  a  brilliant  orange 
color.  (Angas.) 

Alt.  13,  greater  diam.  22,  lesser  18  mill. 

Ysabel  Island,  Solomon  Group. 

H.  brenchleyi  ANGAS,  P.  Z.  S.  1878,  p.  861,  t.  54,  f.  7  (not  H. 
brenclileyi  Braz.). — H.  (Papuina)  walleri  BRAZIER,  Proc.  Linn.  Soc. 
N.  S.  Wales  viii,  p.  228,  1883. 

Has  the  orange  coloring  of  H.  boivini  behind  the  lip,  but  is  more 
depressed,  and  otherwise  different. 

H.  ALFREDI  COX.       PI.  6,  figS.  7,  8. 

Imperforate,  depressed-globose,  glabrous,  shining,  polished,  black- 
ish-brown, having  two  opaque-white  bands,  one  at  periphrey,  the 
other  below  the  suture.  Spire  wride-conic,  whorls  5,  slowly  widen- 
ing, nearly  flat,  the  last  shortly  deflexed  in  front.  Aperture  semi- 
oval  ;  peristome  white,  simple,  shortly  reflexed  ;  columella  dilated, 
appressed. 

Alt.  0.85,  greater  diam.  0.93,  lesser  0.82  inch.     (Cox.) 

Solomon  Is. 

H.  alfredi  Cox,  P.  Z.  S.  1871,  p.  323,  t.  34,  f.  1,  la.— PFR., 
Monogr.,  vii,  p.  311. — H.  alfredi  var.  trichroa  MARTENS,  Conchol. 
Mittheil.  i,  p.  5,  t.  2,  f.  8-10 ;  1880. 

A  polished  blackish-brown  shell,  with  strongly  contrasting  bands 
of  opaque-white  at  suture  and  circumference. 

Var.  TKICIIROA  Martens.     PI.  6,  figs.  4,  5,  6. 

Imperforate,  rotund-conoidal,  subtly  striatulate,  shining,  tawny, 
the  periphery  whitish,  bordered  above  and  below  with  black  ;  sutural 


HELIX-PAPUINA.  13 

zonule  white.  Spire  conic,  apex  submamillate ;  whorls  5,  a  little 
convex,  separated  by  moderately  impressed  sutures,  the  last  whorl 
rounded  at  the  periphery,  flattened  beneath.  Aperture  rhombic- 
semi-ovate,  very  oblique  ;  peristome  a  little  thickened,  white,  upper 
margin  straight,  arched  forward ;  the  lower  margin  shortly  ex- 
panded ;  columellar  margin  straightened,  very  oblique,  attenuated 
at  the  insertion. 

Alt.  20,  greater  diam.  28,  lesser  22  mill.     (Mart.~) 

Neiv  Ireland. 

H.  MACFARLANEI  Cox.      Unfigured. 

Shell  imperforate,  thin,  globosely  conical,  smooth,  white,  orna- 
mented with  one  broad  black  band  encircling  the  last  whorl  at  the 
middle,  and  a  broad  blackish-brown  zone  around  the  umbilical 
region,  also  with  dark  brown  or  black  interrupted  broad  zigzag  lines, 
running  interruptedly  across  the  last  three  whorls,  with  additional 
isolated,  irregularly  disposed,  round  black  spots,  principally  on  the 
body-whorl ;  apex  pale,  denuded  of  color ;  transversely  finely  striated 
above  and  below,  and  concentrically  striated  at  the  base  ;  whorls  6, 
rounded,  gradually  increasing  in  size,  last  slightly  deflected  in  front 
at  its  termination  ;  base  convex ;  aperture  obliquely  ovate-lunate ; 
peristome  slightly  thickened  and  everted,  margined  internally  with 
white,  and  externally  with  orange,  the  external  dark  band  on  the 
body-whorl  showing  through  ;  margins  approximating,  joined  by  a 
very  thin  callus  ;  columella  dilated  externally,  adnate  over  the  um- 
bilical region,  internal  margin  descending,  straight. 

Diam.  greatest  0*92,  least  O68  ;  height  0.72  of  an  inch.     (Cox.') 

Solomon  Islands. 

H.  (Geotrochus)  macfarlanei  Cox,  P.  Z.  S.  1873,  p.  567. — PFR., 
Monog.  vii,  p.  315. 

It  is  somewhat  allied  to  H.  coxiana  Angas. 

H.  COXIANA  Angas.     PI.  3,  figs.  36,  37. 

Shell  imperforate,  trochiform,  rather  thin,  obliquely  finely  striated, 
polished,  creamy-white,  the  lower  whorls  ornamented  with  irregular 
chocolate-brown  spots ;  spire  conical,  apex  rather  obtuse ;  whorls  6, 
flatly  convex,  the  last  scarcely  descending,  obtusely  angled  at  the 
periphery,  convex  at  the  base,  the  umbilical  region  pale  purplish- 
brown,  with  the  central  portion  chocolate-brown  ;  columella  sloping, 
flattened,  chocolate-brown  ;  aperture  very  oblique,  ovate  ;  peristome 


14  HELIX-PAPUINA. 

chocolate-brown,  the  margins  approximate,  slightly  thickened  and 
expanded,  the  right  flexuous  above. 

Alt.  22,  greater  diam.  22,  lesser,  19  mill.     (Angas.*) 

Ysabel  Island,  Solomon  Group. 

Geotrochus  coxianus  ANGAS,  P.  Z.  S.  1867,  p.  889,  t.  43,  f.  7,  8.- 
H.  coxiana  PFR.,  Mon.  vii,  p.  313. 

H.  FUCATA  Pfeiffer.     PI.  16,  figs.  9,  10. 

Shell  imperforate,  globosely-conical,  rather  solid,  shining,  porcel- 
laneous, under  the  lens-obliquely  striated,  also  but  more  faintly 
spirally  and  decussatedly  striated ;  white  with  three  black  spiral 
bands,  and  another  faint  reddish  one  above  each  of  the  central 
bands ;  spire  sharply  conical,  acute  and  reddish  at  the  apex,  suture 
smooth  ;  whorls  5,  nearly  flat,  last  convex  slightly  descending,  base 
rather  flat,  with  a  black  umbilical  mark ;  aperture  diagonal, 
Innately  oval ;  peristome  thin,  right  margin  rather  straight,  anteriorly 
produced  and  curved,  left  expanded  and  reflected,  columellar  margin 
tinged  with  pink.  (  Cox.} 

Alt.  14J,  greater  diam.  17,  lesser  15  mill. 

Wide  Bay  and  Marysborough,  Queensland,  Australia. 

H.  fucata  PFR.,  Zeitschr.  f.  Mai.  1853,  p.  56;  P.  Z.  S.  1853,  p. 
59;  Mou.Hel.  Viv.  iii,  p.  641.— Rv.  f.  1029.— Cox,  Monog.  p.  67, 
t.  2,  f.  8. 

H.  CONSCENDENS  Cox.     PI.  8,  figs.  50,  51. 

Shell  imperforate,  globosely  conical,  very  thin,  pellucid,  some- 
what glassy,  rather  shining,  under  the  lens  obsoletely  obliquely  stri- 
ated ;  whitish,  with  one  band  more  or  less  conspicuously  colored ; 
spire  conical,  colored  at  the  tip  ;  whorls  6,  slightly  convex,  last  very 
large,  slightly  deflected,  with  an  obsolete  keel ;  aperture  diagonal, 
lunately  ovate ;  peristome  thin,  outer  margin  somewhat  sinuated, 
columella  above  slightly  reflected. 

Diam.  greatest  0*60,  least  0'50,  height  0'55  of  an  inch.     (Cox.~) 

Lismore,  Upper  Richmond  River,  Australia. 

H.  conscendens  Cox,  P.  Z.  S.  1866,  p.  374 ;  Mon.  Austr.  L.  Sh. 
p.  67,  t.  2,  f.  6. 

(2.)  GROUP  OF  H.  META. 

Narr-oivly  obliquely  perforated;  elevated  conical,  the  periphery 
rounded  or  carinated  ;  last  whorl  not  descending  in  front.  Surface 
smooth,  lightly  obliquely  striatulate.  Distribution,  Solomon  Is. 


HELIX-PAPUINA.  15 

H.  XANTHOCHILA  Pfeiffer.     PI.  9,  figs.  67,  68,  69. 

Shell  obliquely,  narrowly  perforated,  elevated  trochiform,  smooth, 
alabastrine-white  ;  spire  conical,  apex  dark  ;  periphery  rounded ;  lip 
broadly  expanded,  of  a  yellow  color. 

The  shell  is  rather  thin,  but  strong,  having  slight  wrinkles  of 
growth,  but  no  other  sculpture  ;  shining,  white  ;  spire  conical ;  whorls 
6,  but  slightly  convex,  the  last  not  descending  in  front.  Columella 
widely  expanded,  its  inner  edge  nearly  straight  and  sub-vertical. 
Aperture  quite  oblique.  The  axis  is  very  narrowly  perforated 
throughout.  Young  shells  are  keeled. 

Alt.  46,  diam.  29  mill. 

Solomon  Is. 

H.  xanthochila  PFR.  P.  Z.  S.  1861,  p.  192  ;  Novit.  Conch,  p.  175,  t. 

47,  f.  5,  6  ;  Monogr.  v,  p.  334. — DOHRN  in  continuation  of  Kuster's 
Conchyl.  Cab.  p.  584,  t.  172,  f.  7,  8.— Cox,  P.  Z.  S.  1873,  p.  567,  t. 

48,  f.  7.     (variety.) 

A  beautiful  species,  easily  known  by  the  size,  form  and  the  yellow 
lip.  Part  of  the  specimens  before  me  are  marked  Bougainville  Island. 

Variety.  Lip  generally  pure-white,  and  converted  into  broad 
creuulated  frill.  Fig.  69. 

H.  BEATRIX  Angas.     PL  14,  figs.  59,  60,  61,  62,  63. 

Shell  narrowly  perforate,  ovately  conical,  rather  thin,  not  shining, 
finely  obliquely  striated,  variously  colored  ;  spire  conoidal :  whorls 
6*,  rounded,  narrowly  margined  at  the  sutures,  the  last  non-descend- 
ing, somewhat  inflated  and  obtusely  subangulated  at  the  periphery 
in  front,  moderately  convex  at  the  base  ;  aperture  subcircular ;  the 
outer  lip  very  slightly  expanded  and  reflected ;  the  columella 
descending  vertically  and  callously  reflected  over  the  perforation. 
(Angas.) 

Var.  a.  Rich  orange-yellow  throughout,  inner  lip  white,  apex 
sometimes  of  a  carneliau  color. 

Var.  b.  Bright  yellow,  apex  orange,  with  a  narrow  black  baud 
immediately  below  the  suture  of  the  last  whorl,  lip  black. 

Var.  c.     Pellucid- white,  upper  whorls  yellow  or  orange. 

Var.  d.  Upper  whorls  white,  last  whorl  light  brown,  with  a  nar- 
row pale  band  below  the  suture,  lip  white  stained  inside  with  purple. 

Var.  e.  Very  darM  purplish-chocolate  color,  reddish  toward  the 
apex. 


16  HELIX-PAPUINA. 

This  shell,  like  Helix  meta  Pfr.,  is  subject  to  remarkable  varia- 
tions in  color,  but  may  at  once  be  distinguished  by  its  smaller  size, 
the  convexity  of  the  whorls,  and  the  roundness  of  the  aperture,  in 
which  the  subflexuous  outer  lip  and  diagonal  form  of  H.  meta  are 
wanting.  The  general  form  of  the  shell  presents  somewhat  of  a 
Paludina-Yike  aspect.  The  outer  lip  is  much  less  expanded  and 
reflected  than  in  H.  meta,  and  the  texture  less  glossy  and  shining. 
(Angas.^) 

Alt.  26,  greater  diam.  20,  lesser  16  mill. 

Solomon  Archipelago  f 

H.  beatrix  ANG.,  P.  Z.  S.  1876,  p.  265.  t.  20,  f.  1-5. 

H.  CHOISEULENSIS  Brazier.     PI.  10,  fig.  90. 

Shell  perforate,  conical,  moderately  solid,  obliquely  very  finely 
striated,  reddish-brown,  ornamented  with  a  yellow  sutural  band,  and 
two  others  below  ;  spire  conical,  apex  acute ;  whorls  6,  moderately 
convex,  the  last  not  descending,  slightly  angled  at  the  periphery,  con- 
vex at  the  base ;  aperture  diagonal ;  peristome  rose  colored,  rather 
widely  expanded  and  reflexed  ;  columellar  margin  dilated  and 
reflected,  almost  covering  the  perforation.  (Brazier.) 

Alt.  16,  greater  diam.  16  mill. 

Choiseul  Id.,  Solomon  Group. 

H.  choiseulensis  BRAZ.,  P.  Z.  S.  1872,  p.  21,  t.  4,  f.  7. 

H.  SPLENDESCENS  Cox.     Unfigured. 

Shell  covered-perforate,  globose-conic,  thin,  pellucid,  smooth, 
polished,  chestnut  colored,  darker  at  the  carina ;  a  whitish  band  be- 
low the  suture,  and  beneath  two  wider  white  bands ;  spire  conic, 
apex  white;  whorls  6,  subplanate,  the  last  planate  above,  convex, 
very  obsoletely  carinated,  produced  forward  ;  aperture  very  oblique, 
rounded-elongate,  subsinuate  in  front ;  peristome  thin,  reflexed,  ex- 
panded at  the  straight  columella,  chestnut  colored,  narrowly  edged 
with  white.  Alt.  I'lO,  greater  diam.  0*95,  lesser  O'SO  inch.  (Cb.i\) 

Solomon  Islands. 

H.  splendescens  Cox,  P.  Z.  S.  1865,  p.  696 ;  Journ.  de  Conchyl. 
1866,  p.  48.— PFR.,  Monogr.  v,  p.  335. 

H.  mtENCHLEYi  Bra/ier.      U» figured. 

Shell  narrowly  perforate,  conical,  rather  thin,  very  finely 
obliquely  striated,  bright  straw-yellow  with  bright  reddish-chestnut 
band  on  the  periphery  running  spirally  above  the  suture,  below 


HELIX-PAPUINA.  17 

the  suture  broad  white  opaque  band ;  spire  conoid,  apex  rather 
obtuse  ;  whorls  6,  moderately  convex,  base  convex  ;  aperture  diagonal, 
triangularly  ovate,  peristome  white,  thickened  and  reflected  ;  mar- 
gins distant,  columellar  margin  broad  and  expanded  over  the  perfor- 
ation. (Braz.} 

Alt.  25,  greater  diam.  21,  lesser  16  mill. 

Maru  Sound  or  Caracoa  Harbor,  Guadalcanar  Island,  Solomon 
Archipelago. 

H.  Brenchleyi  BRAZ.,  Proc.  Linn.  Soc.  N.  S.  Wales,  i,  p.  3,  1875. 
— Pfr.  Monogr.  vii,  p.  582  (not  H.  brenchleyi  Angas,  P.  Z.  S.  1878, 
p.  861, =H.  walleri  Brazier.) 

H.  MENDANA  Angas.     PI.  8,  figs.  55,  56. 

Narrowly,  obliquely  umbilicated,  elevated  trochiform,  the  periph. 
ery  rounded.  Surface  striatulate,  nearly  smooth  ;  buff,  with  three 
chestnut  bands,  the  lower  one  occupying  the  greater  part  of  the 
base ;  suture  with  a  broad  white  or  buff  margin.  Lip  well  expanded, 
white. 

The  shell  is  thin,  not  so  strong  as  H.  meta.  The  upper  band  is 
often  faint ;  and  frequently  no  bands  are  seen  on  the  spire.  The 
apex  is  rather  acute  ;  the  last  whorl  has  no  trace  of  a  keel.  Aper- 
ture oblique,  rounded-subtriangular,  white  inside  and  showing  the 
three  dark  bands.  Lip  white,  well  reflexed  all  around. 

Alt.  28,  diam.  23  mill. 

Bougainville  Id.,  Shortland  Id.,  Stephen's  Id.  (and  Ysabel  Id.f), 
Solomon  Group. 

H.  mendana  ANCIAS,  P.  Z.  S.  1867,  p.  889,  t.  43,  f.  11,  12.— 
PFR.,  Monogr.  vii,  p.  384. — DOHRN  in  Contin.  of  Krister's  Conchyl. 
Cab.,  p.  571,  t.  168,  f.  14,  15.— SMITH,  P.  Z.  S.  1885,  p.  591. 

The  shell  is  less  solid,  and  the  upper  whorls  of  the  spire  rather 
slenderer  than  H.  meta  and  its  varieties.  The  color  pattern  is  very 
different,  also.  The  broad  basal  band  is  sometimes  replaced  by  a 
number  of  narrower  stripes. 

H.  META  Pfeiffer.     PI.  9,  figs.  59,  60,  61,  63,  70. 

Obliquely,  narrowly  perforated,  elevated  trochiform,  smooth, 
shining.  The  last  one  or  two  whorls  are  dark  olive,  brown  or  pur- 
plish-brown, or  clear  yellow,  becoming  blue  or  purple  on  the  spire  ; 
the  suture  has  a  conspicuous  snow-white  border ;  periphery  rounded 
or  bluntly  angular ;  face  and  back  of  the  lip  dark  brown. 


18  HELIX-PAPUIXA. 

Excessively  variable  in  coloring,  but  typically  as  described  above, 
(figs.  59, 60,  63),  the  dark  form  being  typical.  The  spire  is  conical, 
apex  obtuse;  whorls  5i  to  6,  their  convexity  slight,  the  last  usually 
obscurely  angular  at  the  periphery.  Aperture  oblique,  obtusely 
subtriangular ;  lip  not  widely  expanded,  typically  dark,  triangularly 
reflexed  at  the  umbilicus,  inner  edge  of  the  columella  concave. 

Alt.  25,  diarn.  20  mill. 

Alt.  24,  diam.  22  mill. 

Ysabel  and  Bougainville,  Solomon  Islands. 

H.  meta  PFR.,  P.  Z.  S.  1856,  p.  381,  t.  26,  f.  5  (notf.  4)  ;  Monogr. 
iv,  p.  257. — DOHRN  in  Contin.  of  Kiister's  Conchyl.  Cab.  p.  588,  t. 
173,  f.  1-3.— H.  deidamia  ANGAS,  P.  Z.  S.  1869,  p.  625,  t.  48,  f.  3.— 
BRAZIER,  Jouru.  de  Conchyl.  1880,  p.  311. — H.  acmella  PFR.,  P.  Z. 
S.  1860,  p.  135,  t.  50,  f.  4 ;  Monogr.  v,  p.  335. — DOHRN,  torn.  cit.  p. 
589, 1. 173,  f.  5,  6.— SMITH,  P.  Z.  S.  1885,  p.  590.— BRAZIER,  Journ. 
de  Conchyl.  1880,  p.  302. 

When  typical  this  form  may  be  known  by  the  dark  brown  or 
olive  body-whorl,  purple  spire  and  broad  white  sutural  border. 
There  are,  however,  many  variations  in  color  pattern.  One  of  the 
commonest  is  pictured  in  fig.  60.  The  color  is  clear  yellow  becom- 
ing purple  above ;  sutural  band  present ;  lip  dark.  Another  yellow 
form  lacks  the  purple  on  spire,  and  has  the  lip  white  ;  suture 
bordered. 

H.  deidamia  Angas  (PL  9,  fig.  70)  is  a  synonym  of  the  typical 
form. 

Var.  ACMELLA  Pfr.     PI.  9,  figs.  64,  65,  66,  62. 

Upper  portion  of  the  outer  lip  more  expanded  ;  color  clear-light- 
olive-yellow,  paler  on  the  spire ;  lip  white. 

The  alleged  differences  of  form  between  this  and  H.  meta  do  not 
hold  in  the  series  of  36  specimens  before  me,  in  which  some  shells 
with  the  widest  body-whorl,  have  the  typical  coloration  of  meta  (fig. 
63).  H.  acmella  has  been  collected  on  Bougainville,  Ysabel,  Faro  and 
Florida  Islands  of  the  Solomon  Group.  The  locality  "  Admiralty 
Islands"  originally  given  for  the  species,  is  incorrect. 

Smith  mentions  a  specimen  from  the  coral  limestone  region  of 
Florida  Island,  which  has  the  body-whorl  of  a  salmon  tint  and  the 
three  uppermost  volutions  bright  red.  The  lip  is  white.  A  speci- 
men before  me  lias  the  same  coloration.  This  form  corresponds 
with  fig.  4,  of  plate  36,  P.  Z.  S.  1856,  considered  by  Pfeiffer  as  a 
form  of  meta.  I  have  figured  it  on  PI.  17,  fig.  27. 


HELIX-PAPUINA.  19 

H.  PLAGIOSTOMA  Pfeiffer.     PL  10,  figs.  88,  89. 

Narrowly  perforated,  elevated  trochiform,  carinated  at  the  periph- 
ery ;  slightly  striated  obliquely  ;  covered  with  a  thin  yellowish-olive 
epidermis,  beneath  which  the  substance  of  the  shell  is  purplish-pink  ; 
the  lip  expanded  and  of  the  same  color. 

The  spire  is  conical ;  whorls  5J,  the  last  not  descending  in  front. 
The  upper  whorls  are  pale  purple-pink,  as  is  the  whole  shell  when 
denuded  of  the  thin  epidermis.  There  is  a  yellowish-white  border 
below  the  suture.  The  base  sometimes  shows  very  fine  concentric 
striae  under  a  lens  and  is  rather  flattened.  Aperture  subtriangular ; 
outer  lip  expanded,  thin,  flesh  colored  or  purplish ;  columella 
triangularly  reflexed,  its  inner  face  nearly  straight  and  vertical. 

Alt.  26-26J  diam.  21-22  mill. 

Solomon  Islands. 

H.plagiostoma  PFR.,  P.  Z.  8.  1856,  p.  381 ;  Mai.  Bl.  1856,  p.  238  ; 
Monogr.  iv,  p.  255. 

Allied  to  H.  adonis,  but  of  a  totally  different  tone  of  coloration, 
wider  body-whorl,  and  the  upper  part  of  the  outer  lip  is  more  ex- 
panded. The  locality  "  Admiralty  Is "  given  by  Pfeifier  is  an 
error. 

H.  GUPPYI  Smith.     PL  10,  fig.  95,  (enlarged). 

Shell  elevately  conical,  thin,  pale  yellow,  ornamented  with  con- 
spicuous nearly  black  or  black-brown  spiral  bands,  one  above  and 
one  below  the  sutures  of  the  upper  whorls,  and  three  upon  the  last, 
one  sutural,  the  second  peripheral,  and  the  third  basal ;  volutions  6, 
rather  slowly  enlarging,  a  little  convex,  sculptured  with  fine  oblique 
striae  of  growth,  not  glossy.  Three  first  whorls  livid  purplish,  the 
last  rather  sharply  angled  at  the  middle,  not  descending  in  front, 
having  the  basal  band  broad  around  the  almost  concealed  perfora- 
tion, and  obsolete  within  the  aperture.  The  latter  is  oblique,  some- 
what narrowed  and  pouting  in  front,  banded  within  with  three 
almost  black  and  two  white  bands,  the  central  one  of  the  former  be- 
ing squarely  truncate  at  the  end,  only  the  lower  corner  of  it  touch- 
ing the  margin  of  the  lip,  which  is  pale,  oblique,  receding,  a  little 
expanded  and  reflexed  in  front  at  the  columellar  margin,  the  upper 
end  of  which  is  spread  and  nearly  conceals  the  small  umbilicus. 

Height  22?  mill.,  greatest  diam.  19,  smallest  16.     (Smith.) 

Faro  Island,  Bougainville  Straits;  in  the  higher  parts  of  the 
island  up  to  the  summit,  1900  feet  above  the  sea. 


20  HELIX-PAPUINA. 

H.  guppyi  E.  A.  SMITH,  P.  Z.  S.  1885,  p.  591,  t.  36,  f.  4. 

This  species  is  remarkable  for  the  striking  contrast  of  its  color- 
bands  and  the  angular  character  of  the  last  whorl.  (Sin.) 

H.  MISER  Cox.      Unfigured. 

Shell  narrowly  perforate,  thin,  coniform,  obliquely  finely  striated  ; 
color  reddish  chestnut,  becoming  pale  yellowish-brown  towards  the 
apex ;  spire  elongated ;  whorls  7,  very  gradually  increasing  in  size, 
the  last  one  being  subcarinated  ;  base  rounded,  aperture  lunar  oval ; 
periphery  straight ;  margins  slightly  approximating,  and  united  by 
a  rather  thin  columellar  callus,  which  descends  perpendicularly,  and 
becomes  broadly  dilated,  concealing  the  umbilicus  ;  aperture  and 
interior  of  the  lip  pale,  the  lip  shortly  reflected  throughout. 

Diam.,  greatest  0'80,  least  0'68  ;  height  1*22  inch.     (Cox.) 

Solomon  Islands. 

H.  miser  Cox,  P.  Z.  S.  1873,  p.  146.— PFR.,  Monogr.  vii,  p.  383. 

H.  ADONIS  Angas.     PI.  10,  fig.  86. 

Narrowly  perforated,  elevated  trochiform,  the  periphery  carinated  ; 
lightly  obliquely  striated ;  of  a  light  orange-buff  tint,  the  lip  of  the 
same  color. 

The  shell  is  thin,  the  smoothness  of  the  surface  slightly  broken 
by  close  but  inconspicuous  growth  lines ;  spire  conical,  apex  sub- 
acute  ;  whorls  5J,  the  earlier  quite  convex,  the  last  two  less  so ;  last 
whorl  rather  acutely  keeled  at  the  periphery,  not  descending  in 
front ;  base  nearly  flat  and  microscopically  spirally  striated ;  aper- 
ture subtriangular,  oblique ;  outer  lip  thin,  scarcely  expanded  above, 
outer  and  basal  margins  expanded,  columellar  margin  triangularly 
reflexed  over  and  almost  covering  the  umbilicus.  Inner  edge  of  the 
columella  vertical,  straight.  Alt.  24,  diam.  18-19  mill. 

Bougainville  Island,  Solomon  Group. 

H.  adonis  ANGAS,  P.  Z.  S.  1869,  p.  624,  t.  48,  f.  4.— PFR.,  Monogr. 
Yii. — BRAZIER,  P.  Z.  S.  1872,  p.  805. — H.  metula  CROSSE,  Journal 
Conchyl.  1870,  p.  248. 

More  acutely  keeled  than  H.  meta  or  acmella,  not  so  smooth,  and 
having  the  lip  as  well  as  the  whole  shell,  of  a  fleshy-buff  color.  It 
differs  from  H.  plagiostoma  in  being  narrower,  lacking  a  subsutural 
white  band,  and  of  a  different  color. 


HELIX-PAPUINA.  21 

H.  BLANDA  Cox.      Unfigured. 

Shell  imperforate,  conical,  very  thin,  translucent,  obliquely  finely 
striated,  of  a  pale  brownish-yellow ;  spire  with  the  apex  acute  ; 
whorls  7,  gradually  increasing  in  size,  the  last  moderately  inflated 
and  sharply  carinated  at  the  periphery;  base  flattened;  suture  not 
margined ;  aperture  subrhomboidal,  margins  somewhat  approximat- 
ing, not  united  by  a  callus,  subangulate  at  the  periphery  of  the  last 
whorl ;  margins  simple,  slightly  everted,  columellar  margin  descend- 
ing vertically,  joining  the  basal  margin  at  an  obtuse  angle.  (Cox.) 

Alt.  '92,  greater  diam.  '76,  lesser  '64  inch. 

Solomon  Is. 

H.  blanda  Cox,  P.  Z.  S.  1873,  p.  147. 

This  species  is  closely  allied  to  H.  adonis  of  Angas,  but  after  com- 
paring it  with  a  number  of  specimens  of  that  species  I  feel  sure  that 
it  has  permanent  differences ;  it  is  a  much  thinner  and  lighter  shell, 
much  more  sharply  keeled,  and  much  less  coarsely  sculptured. 
(  Cox.) 

H.  MENDOZA  Brazier.     PI.  10,  fig.  87. 

Shell  narrowly  perforate,  conical,  rather  thin,  very  finely  obliquely 
striated,  light-straw  color  throughout ;  spire  conoid,  apex  rather 
obtuse;  whorls  6  to  61,  slightly  convex,  narrowly  channelled  at  the 
suture,  sharply  carinated  and  angled  at  the  periphery,  base  rather 
flat ;  aperture  oblique,  triangularly  ovate  ;  peristome  thin,  white, 
right  margin  a  little  expanded,  columellar  margin  reflexed,  and  ex- 
panded over  the  perforation.  (Brazier.) 

Alt.  16,  greater  diam.  13,  lesser  11  mill. 

Choiseul  Island,  Solomon  Group. 

H.  mendoza  BRAZIER,  P.  Z.  S.  1872,  p.  21,  t.  4,  f  8. 
Evidently  allied  to  H.  adonis,  but  more  acutely  keeled. 

H.  HERMIONE  Angas.     PI.  9,  figs.  73,  74. 

Narrowly,  obliquely  perforate,  thin,  elevated  trochoidal,  the  last 
whorl  carinated,  base  rather  flattened ;  surface  obliquely  striatulate, 
nearly  smooth.  Dark  purplish-brown,  with  a  buff  band  a  short  dis- 
tance below  the  carina;  and  generally  another  midway  between  carina 
and  suture.  Earlier  two  or  three  whorls  deep  blackish-purple. 

Spire  conical,  rather  slender ;  whorls  6,  convex,  the  last  not 
descending  in  front.  The  rather  flat  base  is  bounded  by  an  angle. 
Aperture  oblique,  rounded-subtriangular,  showing  the  bands  within. 


22  HELIX-PAPUINA. 

• 

Outer  lip  thin,  narrowly  expanded,  slightly  sinuous;  triangularly 
dilated  at  the  umbilicus,  nearly  closing  it ;  columella  vertical, 
slightly  arcuate.  Alt.  22,  diam.  18  mill. 

Bougainville  Island,  Solomon  Group. 

H.  hermione  ANGAS,  P.  Z.  S.  1869,  p.  625,  t.  48,  f.  5.— PFR., 
Monogr.  vii,  no.  2672.— BRAZIER,  P.  Z.  S.  1872,  p.  805.— IT.  bio- 
cheana  CROSSE,  Journ.  de  Conchyl.  1870,  p.  249. 

Smaller  and  thinner  than  any  form  of  H.  meta,  more  distinctly 
carinated,  differently  colored,  and  with  a  narrower  lip. 

H.  MIGRATORIA  Pfeiffer.     PI.  6,  figs.  99,  1,  2,  3 ;  pi.  17,  figs.  31,  32. 

Narrowly,  obliquely  perforated  ;  trochiform  periphery  obtusely 
angular ;  very  lightly  striated,  with  traces  of  spiral  lines  on  the  base  ; 
shining  flesh  colored,  often  varying  toward  white  or  brown ;  usually 
having  a  brown  band  below  the  periphery  and  another  on  the  upper 
surface,  but  often  bandless,  and  generally  showing  some  sparsely 
scattered  translucent-grayish  dots.  Lip  narrowly  expanded,  white. 
Suture  bordered  by  a  narrow  brown  line. 

The  shell  varies  greatly  in  form.  Spire  conical,  whorls  5,  convex, 
apex  obtuse ;  the  last  whorl  is  convex  above,  and  has  an  incon- 
spicuous angle  at  the  periphery.  The  base  is  convex.  The  whorl 
generally  descends  a  trifle  in  front.  Aperture  very  oblique,  flesh 
colored,  white  or  banded  within ;  outer  lip  markedly  sinuous,  nar- 
rowly reflexed,  dilating  at  the  columella,  almost  covering  the 
narrow  perforation. 

The  banding  is  excessively  variable. 

Alt.  16*,  diam.  17  mill. 

Alt.  22i,  diam.  18  mill. 

Alt.  20,    diam.  17  mill. 

Guadalcanar,  San  Christoval,  etc.,  Solomon  Is. 

H.  migratoriaPFR.  P.  Z.  S.  1855,  p.  108,  t.  32,  f.  3;  Monogr.  iv, 
p.  260— DOHRN,  Contin.  Conchyl.  Cab.  p.  587,  t.  172,  f.  16-19.— 
H.  leucophcea  Cox,  P.  Z.  S.  1872,  p.  20,  t.  4,  f.  5. 

This  species  has  a  pallid,  faded  aspect,  and  in  some  respects  it 
approaches  several  imperforate  species.  Ten  of  the  eleven  speci- 
mens before  me  show  the  grayish  dots  mentioned  above,  although 
there  are  very  few  on  some  of  the  specimens.  The  principal  color 
forms  are  as  follows : — 

a.  Straw-colored  without  band  or  dots. 

b.  Straw-colored,  sparsely  dotted. 


HELIX-PAPUINA.  23 

c.  Buffish-flesh-colored,  with  dots  and  two  faint  pink  bands,  one 
above,  one  below,  the  periphery. 

d.  The  same  ground  color,  with  dark-brown  bands. 

Figure  31,  of  PI.  17,  is  Pfeiffer's  type.  Fig.  1,  PI.  6,  is  the 
leucophcea  of  Cox. 

The  largest  specimen  I  have  seen  is  figured  on  PI.  17,  fig.  32. 

(3.)  GROUP  OF  H.  PILEUS. 

Narrowly,  obliquely  perforated ;  elevated-conical;  last  whorl  not 
descending ;  outer  lip  expanded  ;  surface  smooth,  obliquely  striatu- 
late.  Differs  from  the  group  of  H.  meta  only  in  distribution,  the 
present  group  being  confined  to  the  Moluccas  and  Papua. 

H.  EUCHROES  Pfeiffer.     PI.  8,  figs.  40,  41. 

Shell  half-covered  perforate,  conical,  rather  solid,  obliquely 
irregularly  striate,  and  very  minutely  granulate  under  a  lens ;  a  lit- 
tle shining,  reddish-brown,  ornamented  with  chestnut  bands  and  a 
wider  white  one  below  the  suture.  Spire  regularly  conical,  lilac- 
roseate  above,  apex  rather  acute.  Whorls  7,  moderately  convex, 
sensibly  increasing,  the  last  not  descending,  subrotund.  Aperture 
very  oblique,  subrhombic-lunar,  grayish-white  inside ;  peristome 
thin,  its  reverse  chestnut  colored,  margins  joined  by  a  thin  callus,  the 
right  margin  broadly  expanded  and  subreflexed,  basal  somewhat 
straightened,  reflexed,  forming  an  angle  with  the  columellar  margin, 
which  is  vertical,  slightly  arcuate,  triangularly  dilated,  half  cover- 
ing the  perforation.  Alt.  35,  diam.  36  mill.  (Pfr.') 

East  Indies  (Pfr.)  ;    Celebes  (Rve.). 

H.  euchroes  PFR.,  Mai.  Bl.  i,  1854,  p.  57  ;  Monogr.  iv,  p.  256 ; 
Novit.  Conch,  i.  p.  2,  t.  1,  f.  7,  8. — REEVE,  Conch.  Icon.  f.  1346. 

One  of  the  largest  species  of  Papuina.  The  locality  is  not  known. 
Only  one  specimen  has  been  placed  on  record,  the  figure  of  Reeve 
and  those  in  the  Novitates  being  evidently  drawn  from  the  same 
shell.  It  is  apparently  allied  to  H.  pileus,  but  is  larger,  broader,  and 
differently  colored.  Compare  also  H.  amphizona  (pi.  8,  figs.  52-54), 
a  species  with  solid,  not  perforated,  axis,  and  different  columella  and 
coloring,  but  of  similar  size. 

H.  LENTA  Pfeiffer.     PI.  9,  fig.  71. 

Rather  openly  perforate,  trochiform,  rather  thin,  very  lightly 
striate,  grayish-flesh-colored  or  reddish-corneous,  with  opaque  white 
bands;  spire  elongate-conic,  rather  acute;  whorls  7i,  nearly  flat, 


24  HELIX-PAPUINA. 

slowly  increasing,  the  last  slightly  descending  in  front,  subangulate, 
the  base  flat,  radiately  striated.  Aperture  very  oblique,  subtrigonal- 
oval,  concolored  within  ;  peristome  thin,  broadly  expanded  and  a 
little  reflexed,  violet-blackish  ;  right  margin  lightly  arched,  basal 
straightened,  the  columellar  vaultingly  reflexed. 

Alt.  29,  greater  diam.  29,  lesser  24  mill.     (Pfr.) 

Moluccas  f 

H.  lenta  PFR.,  Malak.  Bl.  1854,  p.  57  ;  Monogr.  iv,  p.  257.— H. 
pileus  var.  PFR.,  MOD.  Hel.  Viv.  i,  p.  324 ;  Kusters  Conch.  Cab.,  p. 
157,  t.  40,  f.  5. 

H.  PILEUS  Miiller.     PI.  8,  figs.  42,  43,  44,  45. 

Xarrowly,  obliquely  perforated,  turrited  trochiform,  thin  but 
strong.  Periphery  obtusely  subcarinated.  Base  rather  flat,  deep 
brown  ;  suture  margined  with  a  dark  line ;  upper  surface  of  whorls 
light,  with  or  without  dark  bands  or  lines,  lip  broadly  expanded, 
white. 

The  spire  is  high-conical,  base  rather  flat ;  surface  smooth,  lightly 
striatulate  ;  apex  obtuse,  the  earlier  two  whorls  dark  purple  or  red- 
dish ;  suture  shallow ;  whorls  6,  the  last  obtusely  angled  at  the 
periphery,  not  descending  in  front.  Aperture  very  oblique,  sub- 
triangular,  browD  or  banded  within.  The  entire  lip  is  broadly 
expanded,  white.  A  circular  tract  of  dark  brown  almost  covers  the 
base,  extending  nearly  to  the  periphery ;  above  this  the  surface  is 
cream  colored,  pinkish  or  yellowish,  with  or  without  brown  spiral 
bands  and  lines. 

Alt.  30,  diam.  22  mill. 

Alt,  32,  diam.  26  mill. 

Aru  Islands. 

H.  pileus  MULL.,  Hist.  Verm.  p.  80. — BORN,  Test.  Mus.  Cses.  t. 
16,  f.  11,  12.— CHEMNITZ,  Conch.  Cab.  ix,  f.  1046-1048=edit.  nov. 
t.  21,  f.  1-3.— FEB.,  Hist.  t.  63A,  f.  3-8.— LAM.,  edit.  Dh.,  viii,  p. 
64.— PFR.,  Monogr.  i,  p.  324.— CHENU,  Illust.  Conch,  t.  6,  f.  4-6.- 
REEVE,  Conch.  Icon.  f.  493. — v.  MARTENS,  Ostas.  Landschn.  p. 
323. — Helix  pileata,  bifasciata  and  Bulla  ambigua  of  GMEL.,  Syst. 
xiii,  pp.  3431,  3609. —  Geotrochm  pileus  BECK,  Ind.,  p.  47. — WAL- 
LACE, P.  Z.  S.,  1865,  p.  411. — H.  (Geotrochus)  pileus  TAP.  CAN., 
Ann.  Mus.  Civ.  Genov.  xix,  p.  137,  1883. 

Distinguished  by  its  high,  turrited  form.  The  pattern  of  band- 
ing is  very  variable.  Tapparone  describes  a  form  with  light  colored 
apex. 


HELIX-PAPUINA.  25 

H.  LACTEOLATA  Smith.       PL  1,  fig.  11. 

Shell  conical,  perforated,  encircled  with  zones  of  a  milk-white 
and  of  blackish-brown  blotched  with  white.  Whorls  5,  striated 
with  oblique  growth-lines,  the  last  whorl  rounded  at  the  periphery, 
slightly  convex  beneath,  a  little  descending  in  front.  Aperture 
very  oblique,  purplish-blackish  toward  the  lip,  blue  within  ;  peri- 
stome  white,  expanded  all  around  and  a  little  reflexed  ;  columellar 
margin  livid-brown  above,  broadly  dilated,  partly  closing  the  um- 
bilicus. Alt.  34,  greater  diam.  36,  lesser  28  mill.  (Smith.*) 

Foot  of  Owen  Stanley  Mts.,  British  New   Guinea. 

H.  (Geotroehus)  lacteolata  SMITH,  Ann.  Mag.  N.  5th.  Ser.  xix,  p. 
420,  t.  15,  f.  9,  1887. 

Bears  considerable  resemblance  to  H.  plurizonata  Ad.  &  Rv., 
from  Mindanao,  but  has  a  very  much  more  elevated  spire,  more 
raised  body-whorl,  and  the  volutions  increase  less  rapidly.  The 
colored  bands  are  eight  in  number  on  the  last  whorl.  Most  of  them 
are  almost  black,  and  generally  margined  with  brown,  and  more  or 
less  blotched  and  smeared  with  an  opaque  cream  color.  (Smith.*) 

H.  BLAINVILLEI  Le  Guillou.     PI.  3,  figs.  59,  60 ;  pi.  8,  figs.  57,  58  ; 
pi.  9,  figs.  75,  76. 

Shell  umbilicate,  conical,  solid,  irregularly  elevately  striate,  opaque, 
a  little  shining,  buffish-flesh-colored ;  spire  conic,  apex  obtuse ; 
suture  submargined ;  whorls  7,  convex,  the  last  subangular  at  the 
periphery,  encircled  by  a  reddish  line ;  not  descending  in  front ; 
flattened  somewhat  below,  umbilicus  very  narrow.  Aperture  little 
oblique,  subtetragonal ;  peristome  white,  upper  margin  very  strongly 
arched,  expanded,  basal  margin  substraighteued  ;  columella  lilac 
colored,  short,  vertical,  reflexed.  Alt.  22,  greater  diam.  22,  lesser 
19  mill.  (Pfr.,  under  H.  gdrtneriana.*) 

Aru  Is. 

H.  blainvillei'L'E  GUILL.,  Rev.  Zool.  1842,  p.  140. — PFR.,  Monogr. 
i,  p.  327. — TAPPARONE  CANEFRI,  Ann.  Mus.  Civ.  Genov.  vi,  p.  561, 
1874 ;  xix,  p.  129,  1883,  t.  3,  f.  4,  o.—H.  gdrtneriana  PFR.,  P.  Z.  S. 
1851,  p.  253  ;  Monogr.  iii,  p.  220;  Conchyl.  Cab.,  p.  402,  t.  145,  f. 
1,  2. — Rv.,  Conch.  Icon.,  f.  419. — FORBES,  Voy.  Rattlesnake,  p.  373, 
appendix. — WALLACE,  P.  Z.  S.  1865,  p.  411. — H.  zoce  PFR.,  Mai. 
Bl.  xii,  1865,  p.  121  ;  Monogr.  v,  p.  326  ;  Novit.  Conch.,  p.  274,  t. 
67,  f.  16,  17. — H.  pileolus  PFR.  (not  Fer.)  Symb.,  p.  75  ;  Mouogr.  i. 
p.  324. 


26  HELIX-PAPUINA. 

The  above  synonymy  was  first  worked  out  by  Tapparone  Canefri. 
The  prominent  characters  of  the  species  are  its  elevated  spire, 
angular  periphery  begirt  with  a  brown  line,  strong,  regular 
sculpture,  etc. 

Var.  a.  (pi.  3,  fig.  59).  Shell  more  convex,  umbilicus  more  open, 
periphery  less  angular,  destitute  of  a  brown  line.  (Tap.  Can.') 

Var.  b.  (pi.  3,  fig.  60).  Shell  less  elevated,  distinctly  carinated, 
whorls  less  convex. 

The  H.  gartneriana  of  Pfeiffer  (pi.  8,  figs.  57,  58)  was  reported 
from  Night  Island,  N.  Australia  by  Macgillivray,  but  doubtless 
erroneously,  mistaking  for  it  the  somewhat  similar  H.  poiretiana. 
See  also  TATE,  Trans.  &  Proc.  Roy.  Soc.  S.  Australia,  v,  p.  49,  50, 
1881-'82.  The  H.  ZOCK  Pfr.  is  figured  on  pi.  9,  figs.  75,  76.  It  seems 
to  be  completely  synonymous. 

H.  poiretiana,  which  also  has  a  basal  brown  line,  differs  from  this 
species  in  having  the  last  whorl  decidedly  descending  in  front. 

H.  CANOVARII  Tapparone  Canefri.     PI.  3,  fig.  56. 

Nearly  covered  umbilicate,  elevated  trochiform,  longitudinally 
very  closely  and  minutely  obliquely  striated  ;  carinated  ;  pale  horn 
colored,  but  when  the  epidermis  is  removed  it  is  pale  violaceous,, 
flesh-colored  toward  the  apex. 

Spire  high-conical,  apex  sub-acute.  Whorls  nearly  7,  a  little 
convex,  separated  by  a  distinct  suture,  margined  above  ;  last  whorl 
rather  acutelv  carinated  below  its  middle ;  base  little  convex,  hav- 

«/ 

ing  very  delicate  radiating  striae.  Aperture  ovate-subrhomboidal,. 
oblique ;  peristome  amethystine,  a  little  thickened,  all  around  sub- 
expanded  and  more  reflexed,  margins  not  approximating,  the 
outer  lip  sub-angular  in  the  middle,  basal  lip  nearly  straight,  colu- 
mella  little  oblique,  dilated,  almost  closing  the  umbilicus. 
Alt.  21,  greater  diam.  21,  lesser  19  mill.  (T.  Cf.) 

Banks  of  the  river  Fly,  New  Guinea* 

H.  canovarii  TAP.  CAN.  Ann.  Mus.  Civ.  Genov.  xix,  p.  137,  t.  3, 
f.  6,  1883. 

Allied  to  H.  blainvillei  and  H.  blanfordi.  Also  to  H.  plagiostoma 
of  the  Solomon  Is. 

H.  BLANFORDIANA  H.  Adams.     PI.  17,  fig.  33. 

Covered  perforate,  conoid,  rather  thin,  obliquely  striated,  and 
ornamented  with  spiral  minute  striae ;  shining,  corneous-white. 


HELTX-PAPUINA.  27 

Spire  elongated,  rather  acute  ;  suture  margined  ;  whorls  6,  a  little 
convex,  perceptibly  increasing,  the  last  not  descending  in  front, 
having  an  acute  carina  below  the  middle ;  base  but  little  convex. 
Aperture  oblique,  subrhomboidal-lunar  ;  peristome  white,  narrowly 
expanded,  the  upper  margin  acute,  basal  arcuate,  reflexed,  covering 
the  perforation.  Alt.  17,  greater  diam.  20,  lesser  16  mill.  (Ad.') 
Port  Dorey,  Andai,  Amberbaki,  New  Guinea;  Moluccas. 

Geotrochus  blanfordi  H.  AD.,  P.  Z.  S.  1865,  p.  415,  t.  21,  f.  1 
(preoc.). — WALLACE,  1.  c.,p.412. — H.  blanfordi  PFEIFFER,  Monogr. 
v,  p.  327,  501.— H.  blanfordiana  H.  AD.,  P.  Z.  S.  1869,  p.  275.— 
H.  blainfordiana  (sic)  TAPPARONE  CANEFRI,  Ann.  Mus.  Civ.  Genov. 
xix,  p.  133, 1883.— H.  poirieri  TAP.  CAN.,  Bull.  Soc.  Zool.  Fr.  1878, 
p.  270. — H.  t.urbinata  VAL.,  in  schedis  Mus.  Zool.  Parisiensis  teste 
Tapp.  Can.,  not  turbinata  of  Beck  or  Jan. 

The  variety  called  poirieri  by  Tapporone  Canefri  is  more  ele- 
vated, aperture  rounder,  base  less  dilated  than  the  type. 

(4.)    GROUP  OF  H.  POIRETIANA. 

Narrowly,  obliquely,  perforated ;  elevated-trochiform  :  the  last 
whorl  descending  in  front ;  the  superior  lip  not  expanded.  Surface 
obliquely  lightly  striatulate. 

Differs  from  the  groups  typified  by  H.  meta  and  H.  pileus  in  the 
descending  last  whorl,  and  simple,  not  expanded  outer  lip. 

H.  POIRETIANA  Pfeiffer.     PL  8,  figs.  46,  47,  48,  49. 

Narrowly,  obliquely  perforated,  elevated-conical,  the  periphery 
rounded  ;  last  whorl,  deeply  deflexed  in  front ;  color  white,  with  a 
narrow  brown  line  below  the  middle  of  the  body-whorl,  and  a  faint 
brown  spot  on  the  columella. 

The  spire  is  high-conic,  apex  obtuse  ;  whorls  7,  scarcely  convex, 
the  last  descending  in  front.  Surface  somewhat  shining,  obliquely 
striatulate ;  pure-white  above,  brown-tinged  beneath,  with  a  narrow 
band  (often  sub-obsolete)  just  under  the  periphery.  Aperture  trans- 
versely oblong  ;  upper  and  outer  lips,  not  expanded,  basal  lip  some- 
what expanded,  triangularly  dilated  at  the  umbilicus,  which  it 
nearly  conceals. 

Alt.  23  J,  greater  diam.  19,  lesser  16  mill. 

Night  Island,  N.  E.  Coast  Australia. 

H.  poiretiana  PFR.  P.  Z.  S.  1851,  p.  254;  Monogr.  iii,  p.  149; 
Conchyl.  Cab.  p.  372,  t.  140,  f.  5,  6— RVE.,  Conch.  Icon.  f.  418.— 


28  HELIX-PAPUINA. 

Cox,  Monogr.  Austr.  L.  Sh.  p.  66,  t.  2,  f.  1. — HEDLEY,  Proc.  Roy. 
Soc.  Queensl.  v,  p.  63. 

A  pyramidal  white  shell,  having  some  slight  resemblance  to 
H.  gartneriana,  but  differing  in  the  deflexion  of  the  whorl  at  the 
aperture,  non-expanded  outer  lip,  etc.  The  species  has,  in  fact,  no 
near  relatives. 

(5.)    GROUP  OF  H.  ANTTQUA. 

Globose-turbinate  species,  perforated,  the  spire  elevated,  last  whorl 
not  carinated,  not  descending  in  front ;  lip  broadly  expanded,  the 
concave  columellar  margin  having  a  small  tooth  in  the  middle. 

H.  ANTIQUA  Adams  &  Reeve.     PI.  14,  figs,  56,  57  ;  pi.  17,  fig.  36. 

Shell  covered-perforate,  globose-turbinate,  solid,  obliquely  striated, 
white.  Spire  turbinate,  apex  rather  obtuse.  Whorls  4J,  convex, 
rapidly  increasing,  the  last  rounded,  not  descending  in  front.  Aper- 
ture oblique,  truncate-oval ;  peristome  simple,  broadly  expanded 
and  reflexed,  the  columellar  margin  subcallus,  subdentate  inside, 
dilated-appressed  above  the  perforation.  (Pfr.*) 

Alt.  31,  greater  diam.  30  mill. 

Usang,  Borneo. 

H.  antiqua  AD.  &  Rv.,  Voy.  Samarang,  Zool.  p.  61,  t.  16,  f.  1. — 
RVE.,  Conch.  Icon.  f.  402. — PFR.,  Monogr.  iii,  p.  172;  Conchyl. 
Cab.  p.  400,  t.  144,  f.  14,  15. — v.  MART.,  Ostas.  Zool.  Landschn. 
p.  322. 

A  unicolored  white  shell,  with  rounded  periphery  and  toothed 
columella. 

Var.  XANTHOSOMA  Pilsbry.     PI.  17,  fig.  36. 

Obliquely  umbilicated,  turbinate ;  composed  of  4J  whorls,  the 
earlier  two  very  convex,  the  last  not  descending  in  front ;  suture 
impressed,  margined  below  by  an  impressed  line.  Surface  very 
delicately  obliquely  striatulate  ;  color  lemon-yellow,  paler  on  the 
last  whorl.  Aperture  rather  large,  oblique,  with  a  satin-like  luster 
inside  ;  lip  white,  very  broadly  expanded,  and  reflexed  ;  columellar 
margin  arcuate,  toothed  in  the  middle,  broadly  expanded  above  and 
around  the  oblique  umbilicus  which  rapidly  narrows  to  a  mere 
perforation. 

Alt.  24  J,  greater  diam.  22,  lesser  17*  mill. 

Aperture  (measured  outside  of  peristome),  oblique  alt.  19,  width 

171  mill. 

New  Guinea  f 


HELIX-PAPUINA.  29 

Agrees  with  H.  antiqua  in  form  and  proportional  size  of  aperture, 
but  differs  in  the  clear  lemon-yellow  color.  In  this  last  character  it 
resembles  the  form  described  by  Sowerby  as  H.  horderi,  but  that 
species  has  a  far  more  elevated  spire  and  greater  number  of  whorls. 
I  have  doubts  as  to  whether  this  form  and  the  next  are  actually 
from  New  Guinea,  as  alleged. 

H.  HORDERI  Sowerby.     PI.  14,  fig.  58. 

Narrowly  perforated,  elevated,  conical,  rather  solid,  obliquely 
very  finely  striated,  yellow.  Spire  elevated,  apex  acute.  Whorls 
6,  a  little  convex,  the  last  slightly  inflated,  obsoletely  angled  at  the 
periphery,  the  base  convex.  Aperture  wide,  oblique ;  peristome 
broadly  expanded  and  reflexed,  columellar  margin  provided  with  a 
small  tooth-like  tubercle  inside,  outwardly  dilated,  flattened. 

Alt.  35,  diam.  30  mill.     (Soivb.*) 

New  Guinea. 

H.  (Geotrochus)  horderi  G.  B.  SOWERBY,  JR.,  P.  Z.  S.  1889,  p. 
577,  t.  56,  f.  1. 

Differs  apparently  from  H,  antiqua  in  the  yellow  color,  greater 
number  of  whorls  and  higher  spire,  consequently  the  mouth  is 

shorter  in  proportion  to  the  total  length. 
* 

(6).  GROUP  OF  H.  PILEOLUS. 

Subperforate  or  imperforate ;  pyramidal,  acutely  carinated,  the 
base  flat.  Distribution,  Moluccas  and  New  Guinea. 

H.  PILEOLUS  Ferussac.     PI.  11,  figs.  99,  100,  1,  2,  3. 

Nearly  covered-perforate,  trochiform,  carinated,  rather  solid, 
striatulate,  the  base  minutely  decussate-gran ulose ;  opaque,  whitish 
or  brownish,  flamed  with  brown  ;  apex  acute,  liver-brown  ;  spire 
conical,  more  or  less  elevated ;  suture  superficial,  hair-margined. 
Whorls  5,  the  earlier  2  or  3  convex,  the  remainder  flat,  the  last  a 
little  convex  below,  descending  a  little  (rarely  not  at  all)  in  front. 
Aperture  very  oblique,  rhombic-triangular,  the  peristome  reflexed, 
white  or  pale  rose-brown  ;  at  the  outer  angle  it  is  curved  over  into  a 
beak,  and  at  the  columellar  insertion  it  is  dilated.  {Mart.") 

Batchian  and  Little  Tawalli  Islands,  Moluccas. 

H.  pileolus  FER.,  Hist.,  t.  63 A,  f.  1,  2. — v.  MART.,  Ostas.  Zool., 
Landschn.,  p.  231,  t.  17,  f.  8-10.— PFR.,  Monogr.  v,  p.  326,  501.- 
Geotrochus pileolns  WALLACE,  P.  Z.  S.  1865,  p.  412. 


30  HELIX-PAPUINA. 

The  following  forms  are  described  by  v.  Martens : 

Form  turrita  (figs.  2,  3).  White  or  pale  isabelline,  painted  with 
narrow  blackish-brown  lightning-streaks ;  last  whorl  distinctly 
descending.  Alt.  20,  diam.  20£  mill.  This  is  like  Ferussac's  fig.  2. 

Form  pyramidata  (figs.  99,  100).  Pale  brownish  ;  marbled  with 
broader  subdiaphanous  brown  lightning-streaks ;  base  nearly  uni- 
colored,  the  last  whorl  scarcely  descending.  Alt.  21,  diam.  24  mill. 
This  corresponds  to  Ferussac's  fig.  1. 

Form  convexa.  Subdepressed,  whitish,  marbled  with  wider 
lightning-streaks  of  pale  brown  ;  base  nearly  unicolored  ;  last  whorl 
abruptly  descending.  Alt.  14,  diam.  24  mill. 

Form  depressa  (fig.  1).  Whitish,  with  rather  narrow  brown 
markings,  the  base  unicolored. 

H.  EHYNCHOSTOMA  Pfeiffcr.     PL  11,  figs.  96,  97,  98. 

Narrowly  umbilicated,  trochiform,  acutely  carinated,  rather  thin, 
striatulate  and  under  a  lens  very  minutely  granulated ;  flesh-colored, 
with  two  reddish  bands,  one  above,  the  other  wider  and  on  the  base. 
Spire  conical,  somewhat  obtuse,  suture  carino-marginate;  whorls  6, 
nearly  flat,  slowly  widening,  the  last  scarcely  descending  in  front, 
base  nearly  flat.  Aperture  very  oblique,  subtriaugular ;  peristome 
thin,  margins  distant,  the  upper  slightly  expanded,  recurved  into  a 
beak  at  the  outer  angle  ;  basal  margin  narrowly  reflexed,  columellar 
dilated,  concealing  the  umbilicus.  Alt.  17,  greater  diam.  25,  lesser 
22  mill.  (P/r.) 

Batchian,  Moluccas. 

H.rhynchostomaPFR.,P.Z.  S.  1861,  p.  21,  t.  2,  f.  6 ;  Novit. 
Conch,  p.  166,  t.  45,  f.  9-11 ;  Monogr.  v,  p.  328,  501. 

Evidently  allied  to  H.  pileolus,  but  totally  different  in  coloration. 

H.  FERUSSACI  Lesson.     PI.  3,  fig.  51. 

Shell  imperforate,  trochiform,  deep-chestnut,  with  a  golden-yel- 
low line  at  the  carina,  and  ascending  the  spire  to  the  apex ;  whorls 
6,  completely  flat ;  the  apex  is  enlarged,  convex  ;  the  last  whorl  has 
a  salient  acute  carina.  Aperture  much  depressed,  triangular,  wider 
than  high,  very  acute  at  the  junction  of  the  upper  and  basal  lips  ; 
peristome  reflexed,  sinuous  at  the  angle. 

Port  Dorey,  New  Guinea,  on  trees. 

Helicogona  (helix)  Ferussacii  LESSON,  Voy.  de  la  Coquille,  Zool. 
ii,  p.  315,  t.  8,  f.  5. — Helix  Feruasati  PFR.  Monogr.  Hel.  Viv.  i,  p. 


HELIX-PAPUINA.  31 

231 — TAPPARONE  CANEFRI  Ann.  Mus.  Civ.  Genov.  xix.  p.  135,  t. 
3,  f.  7.     (Figure  copied  from  Lesson.) 

This  species  is  well  described  and  figured  by  Lesson  in  1830.  It 
has  not  been  seen  by  subsequent  writers.  The  deep-chestnut  shell  is 
begirt  with  a  yellow  peripheral  line.  See  below. 

Var.  EXSULTANS  Tapparone-Canefri.     PI.  3,  figs.  48,  49,  50,  58. 

Imperforate,  trochiform,  rather  solid,  obliquely  finely  striatulate, 
chestnut  or  cinnamon  colored,  with  a  carinal  yellowish  band  and  an 
infra-sutural  chestnut  band,  ascending  the  spire.  Spire  concave-conic, 
apex  obtuse ;  whorls  7,  the  upper  ones  convex,  the  rest  flat,  the  last 
acutely  carinated,  not  descending  in  front ;  base  but  little  convex, 
deep  chestnut.  Aperture  very  oblique,  depressed  subrhomboidal ; 
peristome  rather  thin,  the  upper  margin  expanded,  flexuous  at  the 
carina;  basal  margin  subdilated,  reflexed.  Alt.  20,  greater  diani. 
23,  lesser  19  mill.  (P/r.) 

Dorey  Hum,  Northern  New  Guinea. 

H.  ferussaci  PFR.,  Novit.  Conch.,  p.  212,  t.  56,  f.  7-9 ;  Mouogr. 
Hel.  Viv.  v,  p.  272. —  Geotrochus  ferussaci  WALLACE,  P.  Z.  S.  1865, 
p.  412. — H.  exsultans  TAP.  CAN.,  Ann.  Mus.  Civ.  Genov.  xix,  p.  135, 
t.  3,  f.  8  (fig.  copied  from  Pfr.)  ;  Ann.  Mus.  Civ.  Genov.  xxiv,  p. 
137,  1887. 

The  apex  is  submamillar,  not  acute.  It  may  be  a  species  distinct 
from  H.  ferussaci,  but  it  is  so  closely  allied  that  specimens  of  the 
latter  should  be  compared. 

H.  PYTHONISSA  Tapparone  Canefri.     PI.  3,  fig.  57. 

Shell  covered  perforate,  elevated  trochiform,  under  a  lens  seen  to  be 
minutely,  irregularly  obliquely  striated  and  having  obsolete  spiral 
stride ;  whitish-corneous ;  apex  rather  acute,  reddish-corneous. 
Whorls  7,  a  little  convex  at  the  sutures,  sub-concave  in  the  middle, 
separated  by  sharply  margined  sutures,  last  whorl  not  descending  in 
front,  acutely  compressedly  carinated  below  the  middle,  the  base 
nearly  flat.  Aperture  oblique,  triangular-lunate,  produced  into  a 
short  rostrum  at  the  keel ;  peristome  narrow,  thin,  upper  margin 
acute,  sinuous,  arched  forward,  basal  margin  arcuate-reflexed  ;  colu- 
mella  dilated  toward  its  insertion,  having  a  squarish  tooth  within. 

Alt.  22,  greater  diam.  25,  lesser  19  mill.     (T.  C.) 

Is.  of  Sorong,  near  New  Guinea. 

H.  pythonissa  T.  C.,  Ann.  Mus.  Civ.  Genov.  xix,  p.  136,  t.  3,  f.  9. 


32  HELIX-PAPUINA. 

Intermediate  between  H.  turris  and  H.  ferussaci.     It  differs  from 
«  •> 

the  latter  in  being  larger,  lighter-colored,  and  having  a  columellar 
tooth. 

H.  LEONARDI  Tapparone  Canefri.      Unfigured. 

Ovate-conoid,  perforate,  rather  solid,  pale  yellowish-green  or  red- 
dish, sometimes  subfasciate  on  the  base;  spire  conical,  apex  subacute. 
Whorls  5,  convex,  separated  by  rather  deep  sutures,  decussated  by 
spiral  and  longitudinal  subgranose  stride;  last  whorl  large,  rather 
tumid,  subcontabulate  above,  subangular  below,  the  base  convex. 
Aperture  very  oblique,  ovate-lunate  ;  fauces  citrinous  or  brownish- 
red  ;  peristome  white-edged,  broadly  retiexed,  margins  joined  by  a 
thin  callus,  the  upper  and  outer  regularly  curved,  basal  nearly 
straight;  columellar  margin  reflexed  above  the  perforation,  dilated 
at  the  insertion,  the  base  forming  a  more  or  less  developed  tooth. 
Alt.  34,  greater  diam.  30,  lesser  22  mill.  (Tapp.  Can.) 

Ansus,  Island  of  J obi,  Bay  of  Geelvink ;  Korido,  Is.  of  Misori, 
Geelvink  Bay,  New  Guinea. 

H.  leonardi  T.  C.,  Ann.  Mus.  Civ.  Genov.  1877,  p.  137. 

H.  TURRIS  H.  Adams.     PL  17,  fig.  28,  29. 

Shell  nearly  covered  perforate,  solid,  conical ;  sulcate-striate  ; 
buffish-tawny  ;  spire  elongated,  rather  acute ;  whorls  7,  a  little  con- 
vex at  the  sutures,  subconcave  in  the  middle,  the  last  not  descending 
in  front,  augulated,  the  base  nearly  flat ;  aperture  oblique, 
triangular ;  peristome  simple,  broadly  expanded,  white,  the  upper 
margin  sinuous,  arched  forward ;  basal  margin  reflexed,  with  oblique 
columella  triangularly  dilated,  reflexed  above  the  perforation  and 
subdentate  inside.  Alt.  37,  greater  diam.  36,  lesser  30  mill.  (Ad.~) 

Island  of  Waigheu. 

H.  turris  H.  AD.,  P.  Z.  S.  1865,  p.  415,  t.  21,  f.  4,  5.— PFE., 
Monogr.  v,  p.  325. —  GeotrocJms  turris  WALLACE,  1.  c.,  p.  411. 

H.  FERGUSONI  H.  Adams.     PL  3,  fig.  54. 

Shell  imperforate,  rather  thin,  conical,  obliquely  flexuously 
plicate-striate  ;  seen  under  a  lens  to  be  closely  spirally  striolate ; 
pale-tawny ;  spire  elevated  conical,  apex  rather  acute,  suture  hair- 
margined  ;  whorls  7,  nearly  flat,  the  last  not  descending,  acutely 
carinated  at  the  periphery,  the  base  nearly  flat.  Aperture  oblique, 
triangular-ovate,  the  peristome  broadly  expanded,  slightly  reflexed, 
white,  margins  joined  by  a  thin  callus  ;  right  margin  flexuous,  col- 


HELIX-PAPUINA. 

uniella  sloping,  nearly  straight,  triangularly  dilated,   forming    an 
angle  with  the  basal  lip. 

Alt.  32,  greater  diam.  18,  lesser  13  mill.     (Ad.) 

Island  of  New  Britian. 

Geotrochus  Fergusoni  AD.,  P.  Z.  S.  1872,  p.  614,  t.  42,  f.  14. 
Nearly  allied  to  H.  turris,  but  imperforate,  smaller,  thinner,  more 
acutely  keeled. 

H.  STEURSIANA  Shuttle  worth.      Unfigured. 

Shell  nearly-covered  perforate,  acutely  conical,  trochiform,  thin, 
subdiaphanous,  very  acutely  carinated ;  reddish-corneous,  covered 
with  very  fugacious  epidermis ;  delicately  striated,  and  having  very 
delicate  spiral  lines.  Spire  much  elevated,  apex  papillar;  whorls  8, 
flat  or  subconcave  in  the  middle,  the  last  whorl  flat  on  the  base ; 
suture  hair-like,  subimpressed,  margined  with  tawny.  Aperture 
very  oblique,  triangular- ax-shaped ;  peristome  rosy,  upper  margin 
expanded,  channelled  and  beaked  at  the  carina,  basal  margin 
reflexed,  callous,  thickened,  reflexed  at  the  columella,  covering  the 
perforation.  Columella  arcuate,  furnished  with  a  white  tooth-like 
callus  inside.  Alt.  30,  greater  diam.  40,  lesser  35  mill.  (Shutf) 

Amboyna. 

H.  steursiana  SHUTTLW.,  Bern.  Mittheil.  1852,  p.  200. — PFR., 
Monogr.  iii,  p.  179. 

(7).  GROUP  OF  H.  VITREA. 

Umbilicated ;  depressed,    acutely   carinated ;    aperture    angular. 

Moluccas  and  New  Guinea. 

H.  VITREA  Ferussac.     PI.  12  figs.  35,  36. 

Shell  very  narrowly  umbilicated,  depressed  pyramidal,  acutely 
keeled,  rather  thin,  closely  irregularly  striatulate,  minutely  gran- 
ulated, scarcely  shining,  isabelline-ivhitish,  with  a  very  narrow  per- 
ipheral band  ;  spire  conoidal,  apex  sub-globose ;  whorls  4i,  a  little 
convex,  the  suture  shallow  ;  last  whorl  slightly  and  nearly  equally 
convex  above  and  below,  the  keel  minutely  crenulated  ;  obtusely  an- 
gulated  around  the  umbilicus,  in  front  descending  a  little  below  the 
carina  ;  aperture  slightly  oblique,  trapezoidal-semiovate  ;  peristome 
expanded,  thin,  pale-roseate  or  white,  dilated  at  the  insertion  of  the 
columella.  Alt.  14?,  greater  diam.  27,  lesser  21  mill.  (3/te.) 

Islands  of  Moti,  Ternate  and   Gilolo,  Moluccas;  Port  Dorey  and 

Amberbaki,  New  Guinea. 
3 


34  HELIX-PAPUINA. 

H.  vitrea  FER.,  Hist.  t.  64,  f.  4  (bad). — PFR.,  Conchyl.  Cab.  p.  70, 
t.  76,  f.  18,  19,  (copy  of  Ferus.)  ;  Monogr.  i,  p.  326. — TAPPARONE- 
CANEFRI.  Ann.  Mus.  Civ.  Genov.  xix,  p.  116,  1883. — H.  albula 
LE  GUILL.  Kev.  Zool.  1842,  p.  139. — PFR,,  Monogr.  i,  p.  328- 
MARTENS  Ostas.  Exped.  Landschn.  p.  319,  t.  17,  f.  6. —  Geotrochus 
vitracea  BECK,  Index,  p.  48. 

H.  ARROWENSIS  Le  Guillou.     Unfigured. 

Shell  trochiform,  carinated,  subumbilicate,  white,  hyaline  ;  whorls 
5,  a  little  convex,  the  base  depressed  ;  substriate ;  last  whorl  having 
an  acute  carina,  convexly  depressed  below.  Spire  conical,  apex 
obtuse.  Aperture  subsemiovate,  margin  reflexed  ;  lip  broad,  nearly 
covering  the  umbilicus.  Alt.  12,  diam.  17  mill.  ((?.) 

Aru  Islands. 

H.  arrowensis  LE  GUILL.,  Rev.  Zool.  1842,  p.  139. — PFR.,  Monogr. 
i,  p.  327. — Papuina  arrowensis  WALLACE,  P.  Z.  S.  1865,  p.  411. 

Known  to  me  by  the  original  description  only. 

H.  LANCEOLATA  Pfeiffer.     PI.  11,  figs.  4,  5. 

Shell  half-covered  perforate,  pyramidal,  acutely  carinated,  obliquely 
striate  and  very  delicately  undulate-granulose  ;  above  pale  brownish, 
obsoletely  banded,  below  of  a  blackish  blood-red,  the  keel  pale. 
Spire  moderately  elevated,  apex  subglobose ;  whorls  5,  the  two 
earlier  convex,  the  rest  flat  above,  the  last  whorl  a  little  convex 
beneath,  descending  a  little  below  the  carina  in  front.  Aperture 
oblique,  triangular-trapezoidal ;  peristome  violet,  margin  at  the 
upper  insertion  slightly  expanded,  broadly  revolute  at  the  outer 
angle,  distinctly  reflexed  below,  and  dilated  at  the  columellar  inser- 
tion. (Mts.) 

Alt.  17  i,  greater  diam.  28,  lesser  22  mill. 

Alt.  17,  greater  diam.  26,  lesser  21 5-  mill. 

Halmaheira,  Ternate,  Tidore  and  Moti,  Moluccas;  Angate,  New 
Guinea. 

H.  lanceolata  PFR.,  P.  Z.  S.  1861,  p.  386,  t.  37,  f.  6  ;  Mouographia 
v,  p.  328,  501. — v.  MARTENS,  Ostas.  Exped.  Landschn.  p.  320,  t.  17, 
f.  7. — DOHRN  in  Conchyl.  Cab.  p,  567,  t.  168,  f.  1,  2. — TAPPARONE 
CANEFRI,  Ann.  Mus.  Civ.  Genov.  xix,  p.  117. 


HELIX-PAPUINA.  35 

H.  GRATA  Michelin.     P.  13,  figs.  50,  51. 

Narrowly,  obliquely  perforated,  pyramidal,  acutely  keeled  in  the 
middle  ;  yellow,  with  a  brown  band  above  the  keel  and  a  wider  one 
below  it. 

The  shell  is  solid,  with  oblique  delicate  growth  lines,  and  on  the 
base  microscopic  spiral  lines.  The  pyramidal  spire  is  composed  of  4| 
whorls,  the  earlier  two  shining,  convex.  Last  whorl  having  an 
acute,  compressed  white  peripheral  carina ;  base  convex  ;  whorl  not 
descending  below  the  keel  in  front.  Aperture  oblique,  sub-rhom- 
boidal.  Peristome  yellow,  broadly  expanded,  dilated  at  the  colu- 
mella  into  a  wide  vertical  plate.  Alt.  18,  diam.  23?  mill. 

Port  Dorey,  New  Guinea;  Island  of  Sorong,  Mansinam  and  Jobi. 

Carocolla  grata  MICH.,  in  Guerins'  Mag.  de  Zool.  1831,  t.  9.— 
Helix  grata  FER.  et  DESH.,  Hist,  i,  p.  335,  t.  63B,  f.  1,  2. — PFR. 
Monogr.  i,  p.  325 ;  Conch.  Cab.  p.  356,  t.  136,  f.  7,  8. — DOHRN  in 
Contin.  Conch.  Cab.  p.  595,  t.  174,  f.  12,  13.— TAP.-CAN.,  Ann. 
Mus.  Civ.  Genov.  xix,  p.  117,  t.  6,  f.  7 ;  t.  8,  f.  4,  10.  Anatomy. — 
Helix  aeuta  Q.  &  G.,  Voy.  Astrol.,  Zool.  ii,  p.  98,  t.  8,  f.  1-4  (not  of 
Lamarck). 

The  sulphur-yellow  and  brown  coloration  of  this  species  is 
peculiar.  It  is  variable  in  size  and  degree  of  depression. 

H.  PELECHYSTOMA  Tapparone  Canefri.     PI.  12,  figs.  28,  29,  30. 

Shell  narrowly  umbilicatecl,  depressed  trochiform,  most  acutely 
compressed-carinated,  irregularly  obliquely  striated,  under  a  lens 
seen  to  be  minutely  granulated  ;  corneous-whitish,  covered  with  a 
pale  and  deciduous  epidermis.  Spire  depressed-conoid,  rather  acute. 
Whorls  5  or  51,  nearly  flat,  separated  by  marginulated  sutures;  last 
whorl  deflexed  in  front,  the  base  convex,  radiately  striated.  Aperture 
oblique,  depressed,  ax-shaped ;  peristome  a  little  expanded,  beaked 
at  the  carina ;  upper  margin  nearly  straight,  moderately  reflexed, 
basal  margin  regularly  arcuate,  well  reflexed ;  columella  simple, 
dilated,  nearly  closing  the  narrow  umbilicus.  Alt.  11,  greater  diam. 
25*,  lesser  22  mill.  (Tapp.  Can.) 

Pulo  Faor,  New  Guinea. 

H.  pelechystoma  T.  C.,  Ann.  Mus.  Civ.  Genov.  xix,  p.  120,  t.  2,  f. 
21-23,  1883. 

Intermediate  between  H.  vitrea  and  H.  carinata.  It  is  more  de- 
pressed than  the  former,  the  aperture  more  oblique,  columella  nar- 


36  HELIX-PAPUINA. 

rower,  etc. ;  from  the  latter  species  it  is  separated  by  the  smaller 
size  and  simple,  not  tuberculose,  columella. 

H.  PENNANTIANA  Pfeiffer.     PI.  10,  figs.  81,  82. 

Narrowly  umbilicated,  orbiculate-conoid,  acutely  carinated,  thin, 
striatulate,  irregularly  and  lightly  malleated,  unicolored  fleshy,  the 
apex  reddish.  Whorls  5*,  a  little  convex,  the  last  slightly  descend- 
ing in  front,  base  nearly  flat.  Aperture  very  oblique,  angulate- 
lunar,  white  inside ;  peristome  roseate,  broadly  expanded,  the  basal 
margin  reflexed,  columella  short,  nearly  covering  the  umbilicus. 
Alt.  20,  greater  diam.  37,  lesser  30  mill.  (P/V.) 

Habitat  unknown. 

H.  pennantiana  PFR.,  P.  Z.  S.  1845,  p.  67  ;  Conchyl.  Cab.  p.  432, 
t.  151,  f.  1,  2  ;  Monogr.  i,  p.  .325— RVE.,  Conch.  Icon.  f.  123. 

H.  CARINATA  Hombron  &  Jacquinot.     PL  12,  figs.  31,  32,  33,  34. 

Narrowly  umbilicate,  depressed  trochiform,  irregularly  striated, 
tawny  ;  spire  conoid,  apex  acute ;  suture  light,  subcrenate.  Whorls 
5J,  moderately  convex,  rapidly  increasing,  the  last  very  acutely 
carinated,  the  keel  compressed  and  undulating.  Aperture  rhom- 
boidal ;  peristome  broadly  expanded  and  reflexed,  wavy-calloused 
inside;  columella  sloping,  obsoletely  bi-tuberculate.  Alt.  18,  diam. 
43  mill.  (Pfr.,  descr.  from  figures.) 

Neiv  Guinea. 

H.  carinata  HOMBR.  &  JACQ.,  Voy.  Pol  Sud,  Zool.  v,  t.  7,  f.  26- 
29. — ROSSEAU,  text  of  same,  p.  26. — PFR.,  Mon.  iv,  p.  254. 

H.  LEUCOTROPIS  Pfeiffer. 

Narrowly  umbilicated,  depressed,  rather  solid,  closely  striatulate, 
somewhat  shining,  flesh-colored;  spire  short  conoidal ;  suture  white- 
margined  ;  whorls  4J,  a  little  convex,  the  last  having  an  acute,  com- 
pressed white  carina,  slightly  descending  in  front,  base  more  con- 
vex. Aperture  oblique,  sub-rhombic-lunar,  peristome  thin,  margins 
converging,  the  upper  narrowly  expanded,  basal  reflexed,  dilated  at 
the  insertion.  Alt.  8,  greater  diam.  18,  lesser  15  mill.  (Pfr.) 

Am  Islands. 

H.  leucotropis  PFR.,  P.  Z.  S.  1861,  p.  22;  Monogr.  v.  p.  407.- 
Papuina  leucotropis  WALLACE  P.  Z.  S.  1865,  p.  411. 

An  unfigured  species,  collected  by  Wallace. 


HELIX-PAPUINA.  37 

H.  NODIFERA  Pfeiffer.     PL  13,  figs.  34,  35. 

Shell  umbilicate,  trochiform,  acutely  carinated,  rather  thin, 
striatulate,  diaphanous,  pale  buff.  Spire  conoid,  apex  rather  acute, 
suture  lightly  impressed ;  whorls  5,  slightly  convex,  the  last  not 
descending,  having  an  acute  white  keel  in  the  middle,  convex  above 
and  below.  Aperture  oblique,  subrhomboidal,  the  peristome  broadly 
expanded,  margins  subparallel,  columellar  margin  narrowly  vault- 
ingly  reflexed  above  the  narrow  umbilicus,  and  having  a  nodule  on 
its  inner  edge.  Alt.  18,  greater  diam.  30,  lesser  24  mill.  (Pfr.) 

Island  of  Batchian,  Moluccas. 

H.  nodifera  PFR.,  P.  Z.  S.  1861,  p.  21,  t.  2,  f.  4 ;  Novit.  Conch., 
p.  166,  t.  45,  f.  7, 8  ;  Monogr.  v,  p.  328. — Papuina  nodifera  WALLACE, 
P.  Z.  S.  1865,  p.  411. 

Distinguished  by  the  acute  keel,  nodiferous  columella  and  light 
color. 

(8.)  GROUP  OF  H.  LABIUM. 

Umbilicate ;  depressed-trochoidal.  not  acutely  keeled.  New 
Guinea. 

* 

H.  LITUUS  Lesson.     PL  10,  figs.  77,  78. 

Shell  narrowly  umbilicated,  orbicular-conoid,  apex  obtuse,  shin- 
ing, buffish-brown  or  reddish-chestnut,  encircled  by  a  red  or  black- 
ish zone ;  whorls  6,  a  little  convex,  the  last  rather  flattened  below. 
Aperture  transversely  ovate  ;  peristome  broadly  reflexed,  blackish, 
or  rosy  within  ;  basal  margin  straightened  somewhat,  forming  an 
angle  with  the  columellar  margin,  which  is  dilated,  nearly  closing 
the  umbilicus.  Alt.  25,  greater  diam.  37,  lesser  29  mill.  (Pfr.) 

Port  Dorey ;  Is.  Sorong ;  Is.  Jobi ,  Bay  of  Geelvink;  Andai. 
Amberbaki,  New  Guinea. 

H.  lituus  LESS.  Voy.  de  la  Coq.  Zool.,  p.  309. — FER.  &  DH.,  Hist. 
i,  p.  175,  t.  106,  f.  6-9.— PFR.,  Conchyl.  Cab.  p.  433,  t.  151,  f.  3,  4 ; 
Monogr.  i,  p.  326. — TAPP-CAN.,  Ann.  Mus.  Civ.  Genov.  xix,  p.  138. 
1883  ;  1.  c.  2d.  Ser.  iv,  p.  137,  1887. —  H.  ardouini  DESH.,  Encycl, 
Meth.  Vers.  ii,  p.  266. — H.papuensis  Q.  &  G.,  Voy.  dePAstrol,  Zool. 
ii,  p.  96,  t.  7,  f.  10-13. 

There  is  considerable  variation  in  this  species,  according  to  Tap- 
parone  Canefri.  The  shell  is  covered  with  a  thin,  pale  corneous 


38  HELIX-PAPUINA. 

epidermis,  which  is  readily  deciduous.     The  measurements  given  by 

Canefri  are  as  follows : — 

Alt.  25,  diam.  maj.  37,  min.  29  ;     Alt.  21,  diam.  maj.  35,  min.  27  ; 

Alt.  18,  diam.  maj.  26,  min.  21  mill. 

Var.  a.  Shell  whitish-gray,  peristome  white.     (Dh.) 
Var.  b.  Shell  whitish,  girdled  by  a  red  line.     (Z)A.) 
Var.  c.  Shell  reddish-chestnut,  with  a  broad  blackish-brown  per- 
ipheral band.     (D/i.) 

Var.  d.  Shell  large,  whorls  more    convex,  spire   more   elevated, 

apex  pallid,  concolored ;  no  peripheral  zone.     (Tapp.~) 

Var.  e.  Shells  smaller,  more  globose,  aperture  less  ovate,  umbilicus 

more  open,  columellar  lip  less  dilated,  less  covering  the  umbilicus ; 

no  peripheral  band.     (  Tapp.) 

H.  LABIUM  Ferussac.     PI.  12,  figs.  18,  19,  25,  26,  27. 

Narrowly  umbilicate,  depressed,  solid ;  spire  low-conic,  apex 
acute  ;  whorls  5,  the  last  obtusely  angular  at  the  periphery,  not 
descending  in  front ;  surface  shining,  the  growth-strife  fine,  light ; 
color  light  brown,  lighter  above,  the  periphery  girdled  with  brown  ; 
Aperture  transverse,  wide,  lip  white,  broadly  reflexed  all  around, 
basal  lip  straightened,  dilated  at  the  insertion,  partially  concealing 
the  deep,  narrow  umbilicus. 

Alt.  20,  greater  diam.  33,  lesser  25  2  mill. 

Port  Dorey,  Northern  New  Guinea;  Island  of  Sorong. 

H.  labium  FER.,  Hist.,  t.  64,  f.  6. — PFR.,  Mon.  Hel.  Viv.  v,  p. 
336.— DOHRN  in  Contin.  Conchyl.  Cab.  p.  570,  t.  168,  f.  10,  11.— 
TAPPARONE  CANEFRI,  Ann.  Mus.  Civ.  Genov.  xix,  p.  140,  1883. 

The  peristome  is  very  broadly  reflexed,  and  is  somewhat  angled 
at  the  outer  extremity.  Wallace  mentions  a  variety  having  no 
peripheral  girdle,  and  paler  in  color.  The  specimens  before  me  are 
larger  than  Ferussac's  figures,  and  somewhat  less  angular.  They 
correspond  well  with  Dohrn's  figures.  There  is  considerable  varia- 
tion in  the  degree  of  covering  of  the  umbilicus. 

This  species  has  a  more  convex  base  than  If.  lituus  and  its  other 
allies. 

H.  PSEUDOLABIUM  Pfeiflfer.     PI.  12,  figs.  20,  21. 

Narrowly  urnbilicated,  conoid,  obliquely  striatulate,  shining,  thin, 
pale  tawny,  with  a  red  girdle.  Spire  broad-conoidal,  apex  acute; 
whorls  6,  slightly  convex,  the  last  rather  depressed,  the  base,  nearly 
flat;  columella  short,  oblique.  Aperture  angular-lunar,  whitish  in- 


HELIX-PAPUINA.  39 

side,  with  a  pellucid  band  ;  peristome  thin,  very  broadly  expanded, 
reflexed,  white,  the  right  margin  angled,  columellar  margin  half 
covering  the  umbilicus.  Alt.  23,  greater  diam.  38,  lesser  31  mill. 


Mindanao,  Philippines  (/) 

H.  labium  PFR.  (not  Fer.)  Monogr.  Hel.  Viv.  i,  p.  325  ;  Conchyl. 
Cab.,  t.  55,  f.  3,  4.  —  H.  pseudolabium  PFR.,  Monogr.  v,  p.  336. 

Pfeiffer  gives  the  locality  Mindanao,  Philippines;  but  the  shell  is 
apparently  of  the  Papuan  type  and  probably  a  New  Guinea  species. 
It  differs  from  H.  labium  Fer.  in  the  more  conical  spire  and  flatter 
base. 

H.  MULTIZONA  Lesson.     PI.  10,  figs.  83,  84,  85. 

Shell  umbilicate,  depressed  turbinate,  rather  thin,  obliquely 
lightly  striated,  white,  encircled  with  rufo.us  zones  ;  spire  conoid, 
rather  acute  ;  whorls  5,  a  little  convex,  the  last  not  descending,  sub- 
depressed,  flattened  on  the  base  ;  columella  sloping,  obsoletely  nodose  ; 
aperture  oblique,  transversely  dilated,  lunar  ;  peristome  broadly  ex- 
panded, the  columellar  margin  vaultingly  reflexed,  nearly  covering 
the  narrow  umbilicus.  Alt.  20,  greater  diam.  36,  lesser  28  mill. 


Port  Dorey,  New  Guinea  ;  (and  Waigiou  Id. 

H.  multizona  LESS.,  Voy.  de  la  Coquille,  Zool.,  p.  308.  —  FER  & 
DESK.,  Hist,  i,  p.  174,  t.  106,  f.  1,  2,  3,  5.—  PFR.,  Conchyl.  Cab., 
p.  363,  t.  138,  f.  7,  8  ;  Monogr.  i,  p.  326.  —  TAPPARONE  CANEFRI, 
Ann.  Mus.  Civ.  Genov.  xix,  p.  140,  1883.  —  H.  multizonata  DESK,  in 
Lam.,  An.  s.  Vert,  viii,  p.  106.  —  H.  tenuiradiata  Q.  &  G.,  Voy.  de 
1'Astrol.  Zool.  ii,  p.  101,  t.  8,  f.  8-10.  —  H.  spectrum  RVE.,  Conch. 
Icon.,  f.  1424. 

The  synonymous  H.  spectrum  is  figured  on  pi.  6,  fig.  10. 

H.  TAUMANTIAS  Tapparone  Canefri.     PI.  5,  figs.  85,  86. 

Narrowly  umbilicated,  orbiculate-conoid,  obliquely  striated,  sub- 
pellucid,  pale  buff  or  red  iish-  tawny,  with  white  lines  above  and  a 
milk-white  zone  at  the  periphery  (bordered  with  black  or  brown)  ; 
base  tawny  ;  apex  brownish,  subacute.  Whorls  51  (nearly  6),  a  lit- 
tle convex,  having  impressed  obsolete  spiral  lines  ;  last  whorl  large, 
not  descending  in  front,  the  base  flattened.  Aperture  lunate-ellip- 
tical ;  peristome  white,  expanded  and  reflexed  throughout,  the  mar- 
gins not  approaching,  joined  by  a  pellucid,  very  thin  callus  ;  right 


40  HELIX-PAPUINA. 

margin  regularly  curved,  basal  margin  narrowed  somewhat,  and 
nearly  straight,  forming  an  angle  with  the  dilated  columellar  mar- 
gin, which  nearly  closes  the  umbilicus.  Alt.  22,  greater  diam.  33, 
lesser  26  mill.  (  T.  C.) 

Banks  of  the  Fly  River,  Southern  New  Guinea. 

H.  taumantias  T.  C.,  Ann.  Mus.  Civ.  Genov.  xix,  p.  141,  t.  3,  f. 
13,  14  (shell) ;  t.  vi,  f .  4  ;  t.  ix,  f.  16,  18  (anatomy),  1883. 

Resembles  H.  lituus  in  form,  and  H.  multizona  in  coloration. 
Form  a.  Shell  a  little  larger,  peristome  rosy-violet. 
Form  b.  Shell  small,  spire  more  raised,  peristome  rosy. 

H.  RIDIBUNDA  Tapparone  Canefri.     PI.  5,  figs.  82,  83. 

Narrowly  umbi Heated,  orbiculate-conoid,  obliquely  striated,  pel- 
lucid, buff,  variously  brown  banded  and  lineate,  and  having  a  brown 
peripheral  zone,  which  is  sometimes  bordered  below  with  white. 
Base  convex- depressed,  variously  banded  and  lineated  with  brown. 
Spire  conoid,  apex  sub-acute,  brownish  ;  whorls  5s,  a  little  convex, 
transversely  sub-striate  under  a  lens ;  last  whorl  large  not  descend- 
ing in  front.  Aperture  transversely  lunate,  sub-effuse  ;  peristome 
rather  narrow,  margins  expanded  and  reflexed,  not  approaching  ; 
right  margin  regularly  curved,  basal  straightened,  columellar 
margin  moderately  dilated  and  reflexed  above  the  umbilicus. 

Alt,  19,  greater  diam.  29,  lesser  23  mill.     (T.   C.) 

Banks  of  the  River  Fly,  New  Guinea. 

H.  ridibunda  T.  C.,  Ann.  Mus.  Civ.  Genov.  xix,  p.  142, t.  3,  f.  10, 
11  (shell)  ;  t.  vi,  f .  5  ;  t.  viii,  f.  17  (Anatomy)  1883. 

Closely  allied  to  H.  taumantias,  but  smaller,  the  spire  more  elevated, 
the  color  and  form  of  the  lip  different. 

H.  MEDITATA  Tapparone  Canefri.     PI.  5,  fig.  84. 

Narrowly  umbilicated,  globose-conoid,  rather  solid,  subcarinate 
at  the  periphery ;  whitish-buff,  variously  lineated  and  banded  with 
brown  ;  base  depressed  ;  spire  conoidal,  the  apex  subacute,  brownish. 
Whorls  5 1 ,  the  last  not  descending.  Aperture  lunar  rounded- 
ovate  ;  peristome  whitish,  narrowly  subreflexed  throughout,  the 
margins  not  approaching ;  right  margin  regularly  curved,  basal  sub- 
curved,  columellar  very  much  dilated  and  reflexed  above  the  um- 
bilicus. Alt.  15,  greater  diam.  21,  lesser  \1\  mill.  (  T.  C.) 

Katau,  Southern  Neiv  Guinea. 


HELIX-PAPUINA.  41 

H.  meditata  T.  C.,  Ann.  Mus.  Civ.  Genov.  1883,  xix,  p.  144,  t.  3, 
f.  5  (shell)  ;  t.  vi,  f.  6  (Anatomy). 

Close)}7  allied  to  If.  ridibunda,  H.  taumantias  and  If.  brazier  ce ; 
but  smaller  than  either,  more  distinctly  carinated,  and  with  a  very 
broadly  reflexed  columellar  lip. 

H.  AURORA  Pfeiffer.     PL  11,  figs.  10,  11,  2. 

Umbilicate,  turbinate-depressed,  rather  solid,  irregularly  striatu- 
late,  little  shining,  unicolored  brown ;  spire  short  conoidal,  apex 
minute,  rather  acute.  Whorls  4£,  a  little  convex,  the  last  carinated 
at  the  periphery,  scarcely  descending  in  front,  a  little  convex  below. 
Aperture  oblique,  wide,  depressed-lunar,  rosy  and  with  a  pearly  lus- 
ter inside ;  peristome  pale  rose  or  lilac-flesh  tinted,  the  margins  con- 
verging, upper  lip  horizontal,  expanded,  outwardly  subeffuse ;  basal 
lip  slightly  arcuate,  reflexed;  columellar  lip  dilated,  nearly  closing 
the  umbilicus.  Alt.  17,  greater  diam.  30,  lesser  24  mill.  (Pfr.) 

Waiyiou  Id.;  Is.  of  Sorony  and  Ramoi,  New  Guinea. 

H.  aurora  PFR.,  P.  Z.  S.  1862,  p.  271 ;  Novit.  Conch,  p.  207,  t. 
51,  f.  9-11 ;  Monogr.  v,  p.  341. — TAPPARONE  CANEFRI,  Ann.  Mus. 
Civ.  Genov.  xix,  p.  145. — Papuina  aurora  WALLACE,  P.  Z.  S.  1865, 
p.  411. 

From  the  localities  last  given,  Tapparone  Canefri  received  a  vari- 
ety which  is  larger,  without  a  peripheral  carina,  the  aperture  wide, 
peristome  dilated,  well  reflexed,  white ;  measuring,  alt.  20,  greater 
diam.  38,  lesser  30  mill. 

Specimens  from  Sorong  measure  :  alt.  16,  greater  diam.  28,  lesser 
22  mill. 

H.  .EROPE  Smith.      Unfiyured. 

Shell  rather  widely  umbilicated,  depressed  trochiform,  obtusely 
angulated  at  the  periphery,  pallid  or  whitish-brown,  encircled  with 
a  narrow  reddish  zone  about  the  middle.  Whorls  5,  a  little  con- 
vex, rather  slowly  widening,  obliquely  striated  with  delicate  growth- 
lines  ;  last  whorl  carinated,  the  carina  sensibly  disappearing  toward 
the  aperture,  equally  convex  above  and  below,  shortly  descending  in 
front,  about  the  middle  minutely  rugose.  Aperture  transverse, 
lunate,  whitish  or  pale  lilac  within ;  peristome  white,  dilated  and  re- 
flexed,  the  columellar  margin  widely  expanded. 

Alt.  15,  greater  diam.  23?,  lesser  19?  mill.     (Smith.') 

New  Guinea. 


42  HELIX-PAPUINA. 

H.  (Papuina)  cerope  E.  A.  SMITH,  Ann.  Mag.  N.  H.  (6th  ser.), 
vii,  p.  452,  May,  1891. 

This  species  somewhat  resembles  H.  aurora  Pfr.  in  form,  but  is  a 
trifle  higher  in  proportion  to  the  diameter.  It  also  differs  in  being 
a  smaller  form,  and  in  having  a  larger  umbilicus.  The  young  of 
this  species  is  evidently  sharply  keeled  at  the  middle,  but  in  the 
adult  shell  the  keel  gradually  disappears  toward  the  aperture. 
(Smith.*) 

H.  NOVOGUINEENSIS  Pfeiffer.     PI.  5,  figs.  89,  90,  91,  92. 

Imperforate  or  narrowly  perforate;  globose-conical,  solid,  the 
spire  conical,  composed  of  5i,  slightly  convex  whorls ;  apex  acute, 
pale ;  last  whorl  rounded  at  the  periphery,  slightly  descending  in 
front.  Surface  shining,  striatulate,  covered  with  a  thin  corneous 
epidermis,  white  under  the  epidermis,  variously  banded  with  chesnut, 
and  with  an  opaque-white  border  below  the  suture,  the  base  mostly 
chestnut.  Aperture  oblique,  wide  ;  lip  white,  broadly  expanded 
throughout,  reflexed  ;  columellar  margin  wide,  dilated  over  the  um- 
bilicus, usually  closing  it,  and  thickened  within  by  a  deposit  of  callus 
with  an  irregularly  nodulous  edge,  more  or  less  truncated  below. 

Alt.  26,  greater  diam.  41,  lesser  34  mill. 

Northern  New  Guinea ;  Island  of  Sorong. 

H.  novoguineensis  PFR.,  P.  Z.  S.  1862,  p.  270 ;  Novit.  Conch,  p. 
210,  t.  56,  f.  1,  2,  3,  4 ;  Mouogr.  v,  p.  276. — DOHRN,  Contin.  Conch. 
Cab.  p.  567,  t.  168,  f.  3,  4. — TAPPARONE  CANEFRI,  Ann.  Mus.  Civ. 
Genov.  xix  p.  146,  t.  6,  f.  8  ;  t.  9,  f.  3,  20  (Anatomy). — Papuina  novce- 
guineensis  WALLACE,  P.  Z.  S.  1865,  p.  411. — H.  triumplialisf 
TAPP.  CAN.  I  c.  vi,  1874,  p.  562 ;  ix,  1876,  p.  289.—?  H.  trium- 
phalis  REEVE,  Conch.  Icon.  f.  1421,  December,  1854. — H.  mysolen- 
sis  PFR.,  P.  Z.  S.  1862,  p.  270 ;  Novit.  Conch,  p.  206,  t.  54,  f.  7,  8  ; 
Monogr.  v,  p.  336.— TAPP.  CAN.  /.  c.  2d.  Ser.  iv,  p.  138,  1887.— 
Papuina  mysolensis  WALLACE,  P.  Z.  S.  1865,  p.  411. 

A  large,  globose-conic  shell,  variously  banded  with  chestnut,  and 
having  a  white  sutural  border.  The  following  seem  to  be  merely 
varieties ;  c/.  Dohrn,  I.  c. 

Var.  TRIUMPHALIS  Reeve.     PI.  17,  fig.  30. 

Imperforate ;  broadly  trifasciate  with  chestnut. 
Reeve's  name  has  priority  over  that  of  Pfeiffer,  and  I  would  adopt 
it  for  the  species  if  the  identity  were  established  by  an  actual  exami- 


HELIX-PAPUINA.  43 

nation  of  his  type.     This  examination  will  sooner  or  later  be  made 
by  someone,  and  the  nomenclature  revised  if  necessary. 

Var.  MYSOLENSIS  Pfr.     PI.  7,  figs.  29,  30. 

Smaller  and  more  elevated  than  the  type ;  sub-perforate  ;  having 
a  broad  basal  and  a  narrow  peripheral  band. 

Island  of  MysoL 

The  specimen  before  me  is  larger  than  Pfeiffer's  figures,  but  other- 
wise similar. 

H.  WAIGIOUENSIS    H.  Adams.     PI.  17,  figs.  34,  35. 

Subcovered  perforate,  depressed-turbinate,  rather  thin,  plicate- 
striate ;  pale  tawny,  banded  with  white  at  suture  and  periphery,  and 
ornamented  with  narrow  reddish  bands  ;  spire  conoidal,  rather  acute  ; 
whorls  5,  a  little  convex,  the  last  depressed-rounded,  descending  in 
front.  Aperture  very  oblique,  lunar-oval;  peristome white, broadly 
expanded  and  reflexed,  forming  an  angle  with  the  columellar  margin, 
which  is  dilated,  reflexed  above  the  perforation,  and  provided  inside 
with  an  elongated  tubercle.  .  Alt.  21,  diam.  maj.  40,  min.  32  mill. 
(H.  Ad.} 

Island  of  Waighiou. 

H.  waigiouensis  H.  AD.,  P.  Z.  S.  1865,  p.  415,  t.  21,  f.  6,  7. — PFR., 
Mon.  Hel.  Viv.  v,  p.  339,  502. — TAPPARONE  CANEFRI,  Ann.  Mus. 
Civ.  Genov.  xix,  p.  147,  1883. — Papuina  waigiouensis  WALLACE, 
P.  Z.  S.  1865,  p.  411. 

A  large  species,  evidently  closely  allied  to  H.  novoguineensis. 

H.  BRAZIER^E  Brazier.     PI.  5,  figs.  93,  94,  95,  96. 

Nearly  covered  perforate,  solid,  depressed  turbinate,  the  periphery 
rounded ;  spire  low-conic,  composed  of  4i  convex  whorls,  the  last 
descending  in  front,  rather  flattened  beneath.  Surface  roughened 
by  rude  shallow,  irregular  spiral  and  oblique  furrows  and  mallea- 
tions.  Color  opaque  white,  the  latter  two  whorls  marbled  and 
streaked  irregularly  with  dark  brown,  the  earlier  whorls  and  apex 
bluish  or  corneous-white.  Aperture  very  oblique,  transversely  oval, 
dark  brown  inside;  the  peristome  is  thin,  expanded,  white,  baso-col- 
umellar  margin  reflexed,  nearly  concealing  the  umbilicus.  Parietal 
wall  and  a  spot  at  the  insertion  of  the  columella  dark  brown. 

Alt.  19,  greater  diam.  25  i,  lesser  20-1  mill. 

Yule  Island,  Southern  Neiv  Guinea. 


44  HELIX-PAPUINA. 

H.  brazierce  BRAZIER,  Proc.  Linn.  Soc.  N.  S.  Wales,  i,  p.  107, 
126,  1876. — TAPPARONE  CANEFRI,  Ann.  Mus.  Civ.  Genov.  xix,  p. 
152,  t.  4,  f.  2  ;  t.  5,  f.  2  (shell)  ;  t.  vii,  f.  1  ;  t.  viii,  f.  7,  13  (anatomy). 

This  shell  is  curiously  mottled  with  dark  brown ;  figures  93-95 
are  drawn  from  a  specimen  received  from  Brazier. 

H.  TOMASINELLIANA  Tapparone  Canefri.     PI.  7,  figs.  27,  28. 

Covered  perforate,  globose-turbinate,  solid,  sharply  obliquely 
striate-crenulate  ;  tawny,  and  under  a  subpearly  epidermis  having 
a  number  of  brown-bluish  bands  and  minute  irregular  spots  ;  white 
at  the  suture.  Spire  conoid,  vertex  acute,  brown.  Whorls  4£,  con- 
vex and  separated  by  an  impressed  suture.  Last  whorl  large,  a  little 
descending  in  front,  subconstricted  in  the  middle  and  then  very 
obscurely  subangulated,  the  base  little  convex.  Aperture  ovate, 
very  oblique,  the  throat  brownish-pearly  ;  peristome  white,  margins 
converging,  joined  by  a  pellucid  brown  callus.  Lip  expanded  and 
reflexed  throughout,  subflexuous  above,  the  base  little  curved,  col- 
umellar  margin  subarcuate,  its  base  dentiform  and  truncate,  brown 
spotted  within,  and  expanded  at  the  insertion  above  the  umbilicus. 

Alt,  23,  greater  diam.  36,  lesser  29  mill.     (T.C.) 

On  the  River  Fly,  and  at  Katau,  Southern  New  Guinea. 

H.  tomasinelliana  T.  C.,  Ann.  Mus.  Civ.  Genov.  xix,  p.  148,  t.  4, 
f.  1,  t.  5,  f.  1  (shell)  ;  t.  7,  f.  3 ;  t.  8,  f.  6,  12  (Anatomy.) 
Allied  to  H.  brazierce,  but  larger,  flatter  beneath,  etc. 

H.  GESTROI  Tapparone  Canefri.     PI.  5,  figs.  97,  98. 

Covered  perforate,  globose-turbinate,  solid,  obliquely  rudely 
striated,  covered  with  a  buff  epidermis,  under  the  epidermis  white, 
sometimes  ornamented  with  very  obsolete  pale-gray  bands.  Spire 
conoidal,  apex  acute,  white.  Whorls  4£,  convex,  separated  by  im- 
pressed sutures ;  the  last  whorl  a  little  descending  in  front,  convex 
below  the  sutures,  then  depressed,  becoming  convex  and  sub-angu- 
lar at  the  periphery  the  base  a  little  convex.  Aperture  transversely 
ovate,  very  oblique,  throat  white  ;  peristome  white,  the  margins  con- 
verging and  joined  by  a  thin  pellucid  callus,  expanded  and  reflexed 
throughout ;  basal  margin  nearly  straight ;  columellar  margin  ex- 
panded at  the  insertion  and  nearly  covering  the  umbilicus,  its  base 
truncated,  tooth-like. 

Alt.  26,  greater  diam.  34,  lesser  29  mill.     (T.  C.) 

Banks  of  the  Fly  River,  Southern  New  Guinea. 


HELIX-PAPUINA.  45 

H.  gestroi  T.  C.,  Ann.  Mus.  Civ.  Genov.  xix,  p.  150,  t.  4,  f.  3  ;  t. 
5,  f.  3  (shell)  ;  t.  7,  f.  2,  and  t.  8,  f.  5,  14  (Anatomy). 

Differs  from  the  preceding  species  in  the  more  globose  form,  spire 
more  elevated,  base  more  convex,  aperture  less  elliptical ;  in  the 
pallid  coloration  and  white  interior  of  the  aperture ;  the  relative 
altitude  of  the  last  whorl ;  and  in  lacking  the  fine  strke  which  in  the 
other  species  intersect  the  elevated  stride  of  the  surface  and  render 
them  crenulated. 

H.  SICULA  Brazier.      Unfigured. 

Shell  narrowly  umbilicated,  depressly  globose,  rather  solid,  finely 
striated ;  pale  brown  ornamented  with  three  chestnut  bands,  two 
narrow  and  one  broad  contiguous  to  that  at  the  suture ;  spire 
conoid,  apex  dark  brown,  obtuse,  whorls  5,  slightly  convex,  rounded 
at  the  periphery,  convex  at  the  base,  ornamented  with  eight  narrow 
spiral  chestnut  lines ;  aperture  diagonal,  roundly  lunate,  peristome 
rose-pink,  slightly  expanded  and  reflected  columella  dilated  and 
reflected,  concealing  one-fourth  of  the  umbilicus.  Var.  With  chest- 
nut band  on  the  periphery,  and  running  spirally  around  the  suture 
to  the  apex.  (jBraz.)  Alt.  13,  greater  diam.  20,  lesser  16  mill. 

Banks  of  Katau  River,  New  Guinea. 

H.  (Geotrochus)  siculus  BRAZIER,  Proc.  Linn.  Soc.  N.  S.  Wales 
i,  p.  106,  126. — H.  (Papuina)  sicula  TAPP.  CAN.,  Ann.  Mus.  Civ. 
Genov.  xix,  p.  153,  1883 ;  1.  c.  2d  series,  iv,  p.  138,  1887. 

Two  specimens  only  were  found  ;  on  trees. 

H.  MACLAYANA  Brazier.      Unfigured. 

Shell  covered  with  a  thin  yellowish-brown  epidermis,  umbilicated, 
depressly  conoid,  rather  solid,  obliquely  plicately  striated,  every- 
where minutely  malleated,  encircled  with  numerous  spiral  broken 
purplish-chestnut  lines  and  whitish  opaque  patches  ;  spire  somewhat 
obtuse  ;  whorls  4J  moderately  convex,  the  two  apical  whorls  smooth, 
white,  the  last  large,  convex,  descending  in  front ;  suture  impressed, 
having  a  narrow  white  line;  below  the  suture  blotches  of  purplish- 
chestnut  ;  base  convex  having  four  spiral  purplish  lines  slightly 
broken  up  into  blotches ;  aperture  very  oblique,  large,  ovate,  interior 
purplish-brown ;  peristome  thickened,  white,  reflected,  margins 
approximating,  the  right  rather  thin  at  the  upper  part ;  columellar 
margin  thickened,  broadly  expanded  and  reflected  covering  half  of 
the  umbilicus. 


46  HELIX-PAPUINA. 

Only  one  specimen  of  this  species  was  found.  It  has  a  thick, 
solid,  white  reflected  lip  5  mill,  broad,  and  the  interior  of  an  intense 
purplish-chestnut. 

It  also  resembles,  in  its  markings,  Helix  Brazier  ce.  Braz.,  from 
Yule  Is.,  and  Helix  taumantias  Tapparone  Canefri,  from  the  Fly 
and  Katau  Rivers,  New  Guinea.  {Brazier, .) 

Alt.  24,  greater  diam.  39,  lesser  30  mill. 

Maclay  Coast,  North-eastern  New  Guinea. 

H.  maclayiana  BRAZ.,  Proc.  Linn.  Soc.  N.  S.  Wales,  x,  p.  841, 
1886. — H.  maclayana  TAPP.  CAN.,  Ann.  Mus.  Civ.  Genov.  2d.  Ser. 
iv,  p.  138,  1887. 

(9.)    GROUP  OF  H.  FLEXILABRIS. 

Imperforate/turbinate,  the  surface  corrugated  by  oblique,  forward- 
descending  wrinkles.  This  group  is  very  closely  allied  to  the  fol- 
lowing one. 

H.  VEXILLARIS  Pfeiffer.     PI.  14,  figs.  4,  65,  66,  67. 

Imperforate,  elevated,  trochoidal,  sculptured  with  rather  coarse 
forward-descending  wrinkles ;  opaque  white  above,  the  base  olive  or 
olive-brown. 

The  spire  is  conical,  straight-sided;  whorls  5J,  slightly  convex, 
the  last  bluntly  angular  at  the  periphery,  rather  flattened  beneath, 
not  descending  in  front.  The  surface  has  rather  separated  and 
coarse  obliquely  descending  wrinkles,  the  base  polished,  with  fine, 
subobsolete  spiral  striae.  Aperture  very  oblique,  white  within  ; 
peristome  white,  expanded,  reflexed,  the  outer  and  basal  lips  narrow, 
columella  white,  long,  sloping,  dilated  into  a  flat  plate  and  appressed 
to  the  base,  deeply  inserted. 

Alt.  22 £,  greater  diam.  20,  lesser  16  mill. 

Alt.  21,  greater  diam.  20,  lesser  17  mill. 

New  Georgia,  Solomon  Is. 

H.  vexillaris  PFR.,  P.  Z.  S.  1855,  p.  113 ;  Novit.  Conch,  i,  p.  109, 
t.  31,  f.  5,  6  ;  Mouogr.  iv,  p.  201. — DOHRN,  Contin.  of  Conchyl.  Cab. 
p.  593,  t.  174,  f.  5-7.— H.  phthisica  PFR.,  P.  Z.  S.  1856,  p.  383; 
Mai.  Bl.  1856,  p.  239  ;  Novit.  Conch,  i,  p.  110,  t.  31,  f.  7,  8 ;  Mon. 
Hel.  Viv.  iv.  p.  201. 

The  contrast  between  the  olive  base  and  white  or  bluish  upper 
surface,  and  the  coarse  wrinkling  are  the  more  striking  characters 
of  this  shell.  Pfeiffer  described  the  narrower  form  as  a  distinct  spe- 


HELIX-PAPUINA.  47 

cies,  but  there  is  not  the  slightest  ground  for  a  separation.  A  specimen 
before  me  has  an  indistinct  band  above,  and  the  col umella  and  pari- 
etal wall  are  of  a  blackish-chestnut  color  (pi.  16,  figs.  25,  26).  Pfeif- 
fer  and  Dohrn  give  Admiralty  Is.  as  the  locality  of  the  species,  but 
those  before  me  are  from  the  Solomons,  and  I  am  disposed  to  think 
the  other  locality  doubtful,  it  being  taken  from  the  Curning 
collection. 

H.  BOYERI  Fischer  &  Bernard!.     PL  13,  figs.  48,  49. 

Imperforate,  trochiform,  rather  solid,  obliquely  striate  and  decus- 
sated with  undulating,  forward-descending  striae.  Whitish  and 
tawny  clouded.  Spire  turbinate,  rather  obtuse,  whorls  5i,  convex, 
the  last  descending  in  front,  sub-carinated  and  banded  with  white 
below  the  middle ;  base  little  convex.  Aperture  very  oblique,  sub- 
rhombic,  white  within  ;  columella  entering,  compressed,  flexuously 
sloping,  surrounded  with  rosy ;  peristome  white,  right  margin  nar- 
rowly expanded,  basal  margin  reflexed,  forming  an  obtuse  angle 
with  the  columella. 

Alt.  22-23,  greater  diam.  27-29,  lesser  22-23i  mill.     (F.&B.) 

Louisiade  Islands. 

H.  boyeri  F.  &  B.,  Jouru.  de  Conchyl.  v,  p.  297,  t.  9,  f.  8,9;  Jan., 
1857.— PFR.,  Monogr.  iv,  p.  201  ;  v,  p.  270. 

H.  PH^OSTOMA  Martens.     PI.  7,  figs.  33,  34. 

Imperforate,  trochiform,  thin,  striatulate,  obsoletely  wrinkled- 
malleate,  a  little  shining  ;  isabelliue,  painted  with  zigzag  streaks  of 
blackish-brown,  often  interrupted ;  spire  conical,  elevated ;  whorls 
5,  a  little  convex,  the  earlier  2  unicolored,  diaphanous-corneous,  the 
third  carinated  at  the  suture,  the  last  whorl  rounded  at  the  periph- 
ery and  having  a  wide  white  band  there  ;  below  the  middle  flattened 
and  concentrically  striolate.  Aperture  subdiagonal,  elliptical,  cut  by 
the  adjacent  body-wall ;  peristome  broadly  expanded,  basal  and 
outer  lips  white,  the  columellar  lip  brown,  dilated,  appressed  ;  throat 
brownish.  Alt.  25,  greater  diam.  28,  lesser  20*  mill.  (Mart.) 

New  Hanover. 

H.  pkceostoma  v.  MARTENS,  Monatsber.  k.-p.  Akad.  der  Wissensch. 
zu  Berlin,  May,  1877,  p.  276,  1. 1,  f.  10,  11 ;  Novit.  Conch,  v,  t.  144, 
f.  13,  14. 

Allied  to  H.  gaberti  but  rounded,  not  angular,  at  the  periphery. 
Pfeiffer's  figures  of  H.  gaberti  agree  with  this  species  in  the  rounded 


48  HELIX-PAPUINA 

periphery,  but  they  show  no  light  zone  at  the  circumference.  As  far 
as  my  specimens  or  the  published  figures  show,  lambei,  gaberti  and 
phceostoma  have  always  a  light  peripheral  zone.  I  am  at  a  loss, 
therefore,  to  name  the  figures  of  PfeifFer,  which  I  have  copied  on 
plate  7,  figs.  36,  37. 

H.  GABERTI  Lesson.     PI.  7,  figs.  35,  38,  39. 

Imperforate,  elevated  trochiform,  the  spire  conical,  nearly 
straight-sided,  composed  of  6  whorls,  the  earlier  two  convex,  whit- 
ish, forming  a  papillar  or  sub-globose  nucleus,  the  remaining  whorls 
but  little  convex,  the  last  conspicuously  angled  at  the  periphery, 
slightly  descending  in  front.  Surface  having  light  growth-lines  and 
irregularly  developed,  fine,  forward-descending  wrinkles.  Color 
pale-yellowish  maculated  with  irregular,  interrupted  zigzags  of 
chestnut-brown,  often  forming  an  irregular  band  below  the  carina. 

Aperture  very  oblique,  streaked  and  spotted  with  dark-brown 
inside  ;  lip  broadly  expanded,  white.  Columella  deeply  entering. 

Alt.  25,  greater  diam.  23,  lesser  20    mill. 

Alt.  27,  greater  diam.  22,  lesser  18 5  mill. 

Alt.  22,  greater  diam.  22,  lesser  18    mill. 

New  Ireland ;  New  Guinea ;  Solomon  Is. 

H.  (Pileolus*)  yaberti  LESS.  Voy.  de  la  Coquille,  Zool.  ii,  p.  314. 
-PFR.,  Monogr.  i,  p.  231. — REEVE,  Conch.  Icon.  f.  100. — DESH. 
in  Fer.,  Hist.  p.  333,  t.  106,  f.  10-12.— v.  MART.,  Monatsb.  Preuss. 
Akad.  Wissensch.  zu  Berlin,  1877,  p.  276. — TAPP.  CAN.  Ann. 
Mus.  Civ.  Genov.  xix,  p.  138. — tGeotrochus  gaberti  WALLACE,  P. 
Z.  S.  1865,  p.  412.— IT.  trochus  Q.  &  G.,  Voy,  Astrol.  ii,  p.  110,  t. 
8,  f.  5-7. — H.  trochoides  DESH.,  in  Lamk.  viii,  p.  122  (not  trochoides 
Poir.) 

A  much  more  elevated  shell  than  H.  lambei  and  differing  from 
H.  phceostoma  in  being  distinctly  carinated.  Pfeiffer's  figures  in  the 
Conchylien  Cabinet  (copied  on  my  PI.  7  figs.  36,  37)  do  not  seem  to 
represent  gaberti.  They  may  perhaps  indicate  a  new  species  be- 
tween gaberti  and  phceostoma. 

My  figures  are  drawn  from  specimens  from  New  Britian  ;  I  do 
not  know  whether  the  last  two  localities  given  above  are  to  be  de- 
pended on,  but  consider  them  somewhat  doubtful. 

H.  LAMBEI  Pfeiffer.     PL  7,  figs.  21-26. 

Imperforate,  depressed,  wider  than  high,  the  spire  low-conic,  apex 
obtuse,  corneous  or  brown  ;  whorls  5s,  slightly  convex,  the  last  wide, 


HELIX-PAPUINA.  49 

descending  in  front ;  carinated  in  front,  becoming  obtusely  angular 
on  the  latter  part ;  base  rather  flattened.  Surface  sculptured  with 
fine,  close,  obliquely  descending  wrinkles.  Color  whitish,  buff  or 
rich  brown,  having  a  light  zone  at  the  periphery,  and  either  marked 
ivith  dark  brown,  oblique  zigzag  stripes  and  spots  above  and  below,  or 
having  a  band  above  and  one  below  the  periphery.  Aperture  very 
oblique,  transverse-oval,  white  and  showing  the  bands  within,  or  of 
a  uniform  reddish-brown  color,  peristome  broadly  expanded  through- 
out, reflexed,  white  or  rarely  reddish-brown. 
Alt.  22,  greater  diam.  31,  lesser  24  mill. 

New  Georgia,  Solomon  Is.;  (and  Admiralty  Is.  f) 

H.  lambei  PFR.,  Malak.  Bl.  1856,  p.  239  ;  Monogr.  iv,  p.  200.— 
DOHRN  in  Contin.  Conch yl.  Cab.  p.  593,  t.  174,  f.  1-4. — H.  lombei 
(error)  PFR.,  P.  Z.  S.  1856,  p.  382. 

Allied  to  H.  gaberti  and  H.  phceostoma,  but  much  broader  and 
more  depressed  than  either.  Scarcely  two  individuals  are  marked 
alike.  A  common  variation  is  shown  in  fig.  22,  in  which  there  are 
two  bands  but  no  zigzag  markings.  Fig.  25  represents  a  rarer  form, 
in  which  the  whole  shell  including  lip  is  reddish-brown,  with  darker 
flammules.  Of  the  large  suite  of  specimens  before  me  the  more 
recently  collected  are  from  the  Solomon  Is.  Two  old  trays  (one 
from  Cuming)  are  marked  A  dmirality  Is.,  but  this  locality,  although 
given  by  all  writers  above  referred  to,  requires  confirmation,  in  my 
opinion. 

H.  FLEXILABRIS  Pfeiffer.     PI.  4,  figs.  68,  69,  70,  71. 

Imperforate,  low-turbinate,  the  spire  conical,  last  whorl  rounded 
at  the  periphery,  descending  in  front  as  far  as  the  lower  band  ; 
sculptured  with  very  fine,  close,  forward-descending  wrinkles ;  color 
creamy-white,  with  a  chestnut-brown  band  above  the  periphery, 
ascending  the  spire,  and  another  below  the  periphery.  Aperture 
oblique,  lip  expanded,  white. 

The  spire  is  composed  of  6  rather  convex  whorls,  separated  by 
more  or  less  distinctly  impressed  sutures.  The  last  whorl  descends 
in  front  as  far  as,  or  just  below,  the  lower  brown  band.  The  bands 
are  continuous  and  have  smooth,  well-defined  edges ;  they  are 
prominently  seen  within  the  aperture.  The  lip  is  broadly  expanded, 
closely  appressed  in  the  vicinity  of  the  umbilicus,  which  is  covered 
by  a  thin  white  callus.  The  columella  has  a  thickened  edge,  and  a 


50  HELIX-PAPUINA. 

sort  of  obtuse  fold  near  its  insertion,  which  is  rather  deep  in  the  axis 

tii,'.  69.) 
Alt.  L'l,  -reatcr  diam.  23,  lesser  28  mill. 

Alt.  '-'»',.  -'-eater  diam.  27,  lesser  19'.  mill. 

Solomon  Is. 

H.  ilesi/abris  PFR.,  P.  Z.  S.  1856,  p.  382;  Malak.  Bl.  1856,  p. 
239  :  Mon.  Hel.  Viv.  iv,  p.  199.—  DOHRN  in  Contin.  Conchyl.  Cab., 
p.  587,  t.  172,  f.  14,  15. 

Tlu-  1.  M-ality  given  by  Pfeiffer,  Admiralty  Is.,  is  probably  erroneous. 
Three  trays  before  me  are  from  the  Solomons.  Figures  68,  69  are 
a  little  Hatter  than  Pfeiffer's  type.  Numerous  shells  before  me  agree 
in  characters;  others  are  more  elevated  (figs.  70,  71),  but  I  am  un- 
al.h  t<>  >c|>arate  them  specifically.  There  is  considerable  variation  in 
the  convexity  of  the  whorls,  but  it  is  probably  never  so  marked  asin 
H.  coniform  is,  which  is  from  another  island  group,  and  doubtless  dis- 
tinct specifically. 

There  is  a  translucent-bluish  patch  around  the  columella,  faint,  but 
\\ell-defined  against  the  opaque  cream  ground-color  of  the  shell. 
This  occurs  also  in  H.  gamelia.  All  of  the  specimens  seen  are  two- 
handed. 


II.  rn.viroKMis  Ferussac.     PI.  4,  figs.  78,  79;  pi.  17,  fig.  43. 

I  in  perforate,  orbiculate-conic,  smooth,  whitish-brown,  ornamented 
with  brown  bands  which  are  often  articulated.  Suture  deep  ;  whorls 
."»,  convex,  the  last  flattened  beneath,  descending  in  front.  Aperture 
very  ohliijiie,  large,  oval;  peristome  thin,  white,  broadly  expanded, 
the  margins  converging,  basal  lip  reflexed,  columellar  lip  dilated, 
Alt.  18,  greater  diam.  24,  lesser  18  mill.  (Pfr.) 

Port  Pralin,  New  Ireland. 


II.  II'  lii-nttijhC)  coniformis  FEE.,  Hist.  1.  108,  f.  1.  —  H.  coniformis 
1  >i>n.  in  Lam.,  An.  s.  V.  p.  103;  —  PFR.,  Monogr.  i,  p.  229  ;  Conchyl. 
Cab.  p.  4:;."),  t,  ir,l,  f.  9,  10.—  REEVE,  Conch.  Icon.  f.  101.—  H. 
hirhin'it'i  I  )i>n..  Kneycl.  Meth.  ii,  p.  265.  —  H.tuffetii  LESS.,  Voy.  de 
la  ('o(,nille,  p.  313,  t.  10,  f.  3. 

Tin-  whorls  are  more  convex  than  in  H.  flexilabris,  and  the  bands 
are  often  articulated.  H.  17,  fitf.  43,  is  one  of  Ferussac's  original 
illustrations. 


HELIX-PAPUINA.  51 

Var.  TUFFETII  Lesson.     PL  17,  fig.  42. 

Has  the  contour  of  typical  eoniformis,  but  the  color  is  a  uniform, 
intense  blue-black  ;  lip  bluish-white. 

"  Makakabat,"  New  Ireland. 
H.  SELLERSI  Cox.     PL  15,  fig.  75. 

Shell  imperforate,  trochiform,  brownish-yellow,  generally  or- 
namented with  two  very  dark-chestnut,  almost  black,  bands,  more 
or  less  wide,  one  above  and  one  below  the  sub-carinated  periphery 
of  the  last  whorl,  having  sometimes  between  them  an  opaque-white 
band ;  transversely  rather  regularly  striated  backwards,  decussated 
from  behind  forwards  by  rather  irregular  slightly  waved  propor- 
tionally broadly  separated  stride,  more  distinct  on  the  last  two  whorls 
than  at  the  apex  of  the  spire,  whorls  5,  rounded,  gradually  increas- 
ing in  size,  suture  impressed,  last  whorl  sharply  depressed  at  its  ter- 
mination and  flattened  from  the  periphery  to  its  insertion  ;  spire 
obtuse  ;  aperture  ovately  lunate ;  peristome  white  margined,  slightly 
everted  ;  columella  very  slightly  dilated  and  aduate. 

Diam.  greatest  0'78,  least  0'56  ;  height  0*60  of  an  inch.     (  Cox.~) 

Guadalcanar  Is.,  Solomon  Islands. 

H.  sellersi  Cox,  P.  Z.  S.  1871,  p.  646,  t.  52,  f.  9.—?  H.  sellersi 
DOHRN,  Conchyl.  Cab.  p.  585,  t.  172,  f.9-11. 

(10)       GROUP  OF  H.  BRUMERIENSIS. 

Imperforate,  turbinate  or  trochoidal,  the  last  whorl  descending  in 
front.  Surface  spirally  striate  or  having  forward-descending  wrinkles. 
Lip  more  or  less  sinuous  above,  often  vividly  colored.  Aus- 
tralia, New  Guinea,  Louisiade  Is. 

This  group  comprises  a  considerable  variety  of  forms,  but  they 
are  so  connected  by  intermediate  species  as  to  form  a  perfectly 
natural  assemblage. 

H.  CHAPMANI  Cox.     PL  1,  fig.  13. 

Shell  elevated-conic,  imperforate,  white  ;  the  lip  inwardly  purple- 
brown,  at  the  edge  deep  red. 

Whorls  5J,  a  little  convex,  obliquely  roughly  and  closely  striated, 
and  sculptured  with  flexuous  growth-lines.  Last  whorl  obtusely 
angulated  in  the  middle,  slightly  convex  beneath,  shortly  descend- 
ing in  front.  Aperture  oblique,  irregularly  subquadrangular,  white 
within ;  peristome  somewhat  thickened  and  expanded,  shortly 
reflexed,  the  outer  margin  obtusely  angulated  a  little  above  the 


52  HELIX-PArUINA. 

middle  :  <-.,  lamella  broadly  dilated,  appressed,  flattened,  lilac-colored 
at  its  inner  mar-in,  united  with  the  outer  lip  by  a  blackish-brown 
parietal  callus.  Alt.  34,  diam.  32  mill.  (Smith.') 

Rossel  Island,  Louisiade  Group. 

H  (Geotrochus)  chaj>m<tni  Cox.  Proc.  Linn.  Soc.  N.  S.  Wales,  iv, 
p.  115,  t.  16,  f.  2,  1879  ;  /.  c.  2d  ser.  ii,  p.  1063,  t.  2,  f.  10,  II.—  H. 
(Acavus)  cowliolnlri*  E.  A.  SMITH,  Ann.  Mag.  N.  H.  5th  series, 
six,  ,,.  419,  t.  15,  f.  4,  June,  1887. 

This  species  is  so  well  defined  by  the  remarkable,  contrast  of  the 
color  of  the  peristome  with  the  white  tint  of  the  rest  of  the  shell, 
that  it  will  be  readily  recognized.  The  edge  of  the  aperture  is  thick- 
nied  and  bright  coral-red,  and  this,  both  within  and  without,  is 
rather  deeply  bordered,  especially  within,  with  purple-brown. 

(Smith.') 

The  description  is  from  Smith.     The  identity  of  coraliolabris  with 

l  cannot  be  questioned. 


H.  BRUMERIENSIS  Forbes.     PL  1,  figs.  6,  7,  8. 

Imperforate,  globose-conic,  thin  but  solid,  opaque  white,  the  lip 
intensely  black,  Surface  lusterless,  with  rude,  irregular  lines  of 
•j-rowth,  obsoletely  granulated  by  spiral  lines.  Aperture  very 
oblique,  white  within  ;  upper  lip  roundly  expanded,  sinuated,  baso- 
columellar  lip  flat,  broad,  adnate  for  one-third  the  distance  from 
center  to  periphery. 

The  shell  is  globose  with  short  spire,'  consisting  of  4  rather  con- 
vex whorls.  Suture  impressed.  Last  whorl  not  angled  at  the 
periphery,  descending  in  front.  The  black  border  of  the  lip  is  very 
wide,  extending  .several  millimeters  back  of  the  expanded  portion. 

Alt.  25,  diam.  28  mill. 

J+i<1crii  \cw  (In'uiea,  on  the  shores  of  the   Gulf  of  Huon  ;  M'dl- 
j,»rf  II«rbor,  near  Amazon  Bay  ;  Brumer  Id.,  Louisiade  Group. 

If.  Itriiiii'-ri'-nxis  Foi;r,i>  Voy.  Kattlesnake,  Append,  p.  375,  t.  2, 
t'.  1.  —  Tri:.  Monogr.  iii,  p.  189.  —  REEVE,  Conch.  Icon.  f.  1448.- 
T.MM1.  CAN.  Ann.  Mns.  Civ.  Genov.  xix,  p.  156.  —  ANGAS,  P.  Z.  S. 
1S7G,  p.  48!).—  SMITH,  Ann.  Mag.  N.  H.  5th  ser.,  xix,  p.  419,  1887. 

A  white,  lu-terlrss  shell  with  deep  black  lip.  Although  this  spe- 
ei<  s  is  ])l;ieed  in  Acavux  by  Tapparone  Canefri  and  some  others,  it  is 
a  true  J'ujmiiin,  closely  allied  to  H.  diomedes,  zeno,  naso,  etc., 
mainly  in  the  rounded  instead  of  angular  periphery. 


HELIX-PA  PUIN  A.  53 

New  Guinea  specimens  are  larger  than  those  originally  described, 
and  sometimes  more  conically  elevated.  Smith's  measurements  are  : 
alt.  33,  diam.  38  mill.  This  is  larger  than  any  I  have  seen.  The 
aperture  is  sometimes  pale  rose  tinted  inside.  See  Smith,  /.  c. 

H.  RANGII  Lesson.     (H.  EXTRICANDA  Tapp.  Can.) 

Shell  suborbicular,  globulose,  ventricose,  imperforate,  convex,  the 
last  whorl  of  the  spire  much  swollen,  very  ventricose,  rounded  in 
every  part.  Lips  thick,  bordered,  the  left  margin  dilated,  covering 
the  umbilicus  with  a  smooth  plate.  The  mouth  is  rounded,  the  per- 
istome  level  and  convex.  The  earlier  whorls  of  the  spire  are  small, 
convex,  united  by  a  very  delicate  suture.  The  shell  is  entirely 
covered  with  excessively  fine  interrupted  stride,  which  viewed  under 
a  lens  are  seen  to  be  composed  of  regular,  elevated  aretes,  forming 
a  shagreened  surface. 

This  helix  is  thick,  robust,  with  a  diameter  of  18  lines  [33  mill.], 
alt.  13-14  lines  [26-28  mill.]  ;  of  a  clean  bright-chestnut  color,  the 
greater  portion  of  the  base  being  of  a  lighter  tint.  It  has  great 
resemblance  in  form  and  size  to  H.  melanotragus.  (Lesson.*) 

Port  Dorey,  New  Zealand. 

H.  rangii  LESS.,  Voy.  de  la  Coquille,  Zool.  ii,  p.  305  (not  of 
Desh. ;  not  rangiana  Fer.). — H.  extricanda  TAPP.  CAN.,  Ann.  Mus. 
Civ.  Genov.  xix,  p.  156,  1883. 

This  species  is  known  only  by  the  original  description  translated 
above.  The  color  of  the  lip  is  not  stated.  Later  collectors  in  New 
Guinea  have  not  found  a  shell  agreeing  with  it.  The  comparison 
with  H.  melanotragus  suggests  an  affinity  to  the  group  of  H.  bru- 
meriensis,  zeno,  etc. 

Accepting  the  synonymy  given  by  Dr.  Pfeiffer  as  probably 
founded  upon  some  knowledge  of  this  form  not  given  in  the  descrip- 
tion, I  placed  the  name  in  the  synonymy  of  Hadra  lessoni  (MANUAL 
VI,  p.  163),  but  not  without  expressing  a  doubt  as  to  the  pro- 
priety of  such  reference.  It  should  be  erased  from  that  place. 

H.  ZENO  Brazier.     PL  1,  figs.  1,  2,  3,  4. 

Imperforate,  depressed  trochiform,  the  spire  low-conic,  apex 
obtuse,  whorls  4s;  the  last  obsoletely  angular  at  the  periphery,  very 
deeply  deflexed  toward  the  aperture.  Surface  densely,  finely  and 
irregularly  granulated  by  the  decussation  of  spiral  and  oblique 
stride.  Color  white,  becoming  lilac  on  the  spire,  the  last  whorl  en- 


54  HKLIX-PAPUINA. 

circled  by  numerous  chestnut  bands,  which  are  sometimes  confluent 
above  and  below  the  white  peripheral  girdle,  giving  the  effect  of  a 
marly  nnicolored  chestnut  shell.  Aperture  very  oblique,  rosy  or 
lilac-white  inside:  peristome  expanded,  thickened,  black ;  columel- 
lar  mar-in  broad,  flattened,  black,  its  inner  edge  lilac  colored. 

Tlu  >liell  is  solid,  strong,  the  apical  whorl  dark.  The  last  whorl 
ha-  a  earina  at  its  origin,  which  becomes  rounded  on  the  latter  part 
of  the  volution.  Alt.  27,  diam.  34.  mill. 

//.///  Xnnnd;  Akmcj  Laloki  or  Goldie  River;  Mount  Astrolabe, 
Xn a ther H  New  Guinea. 

If.  zeno  BRAZIER,  Proc.  Linn.  Soc.  N.  S.  Wales,  i,  p.  107,  1876  ; 
/.  c.  ix,  ]>.  805,  1884. — H.  latiaxis  E.  A.  SMITH,  Ann.  Mag.  N.  H. 
5th.  Ser.,  xi,  p.  191 ;  /.  c.  xix,  p.  420,  t.  15,  f.  7,  1887. 

This  large  species  differs  from  H.  diomedes  in  being  less  acutely 
keeled  and  different  in  color.  The  specimens  before  me  show  that 
H.  (Acamis)  latiaxis  Smith  is  a  synonym.  They  vary  much  in  coloring, 
and  in  degree  of  carination.  Fig.  1  represents  Mr.  Smith's  type  ; 
the  other  figures  are  drawn  from  specimens  received  from  Mr. 
Brazier. 

II.  DIOMEDES  Brazier.     PL  1,  figs.  9,  10. 

Imperforate,  trochiform,  acutely  keeled,  solid,  opaque.  Surface 
cut  into  minute  granules  by  oblique  and  spiral  striae;  lusterless 
white,  with  irregular  radiating  patches  or  streaks  and  spiral  inter- 
rupted bands  of  purplish-pink.  Aperture  very  oblique,  pink  within, 
the  lip  intensely  black,  thickened  within,  upper  margin  expanded, 
basal  margin  flatly,  broadly  expanded,  columellar  margin  broad, 
flat,  adnate  to  the  base  for  about  half  the  distance  from  center  to 
periphery. 

The  shell  is  rather  thin  but  solid.      Spire  straightly  conical,  com- 
pox-d  of  4:j  whorls,  the  earlier  1£  rounded,  dark  purple,  the  follow- 
ing flattened  ;  last  whorl  acutely  keeled,  rather  deeply  descending 
in  front.     Aperture  oblong-triangular,  the  upper  lip  sinuous. 
Alt.  26,  diam.  :J2  mill. 

Constance  Island,  New  Guinea. 

II.  'Homedes  BRA/I  1:1;,  Proc.  Linn.  Soc.  N.  S.  Wales  ii,  p.  121, 
L878.  -TAPP.  CAN.,  Annali  Mns.  Civ.  Genov.  xix,  p.  122,  t.  3,  f. 
I-J,  1883, 

A  beautiful  specks,  intermediate  in  form  between  H.  brtuneriensis 
and  //.  zeno,  but  perfectly  distinct  from  either.  It  is  even  more 


HELIX-PAPUINA.  55 

closely    allied  to    H.    naso,   the   coloration    of  some  specimens   of 
the  two  species  being  identical. 

My  description  and  figures  are  from  a  specimen  received  from 
Mr.  Brazier. 

H.  MACGILLIVRAYI  Forbes.     PI.  3,  figs.  38,  39. 

Imperforate,  trochiform,  the  spire  straightly  conic,  periphery  car- 
inated,  base  rather  flattened  ;  of  an  opaque  fawn  or  fleshy-brown 
tint,  occellated  with  small  translucent  dark  dots,  irregularly  bestrewn  ; 
apex  purple-black  ;  lip  white,  sinuous. 

Surface  conspicuously  decussated  with  oblique  and  spiral  striae, 
the  latter  stronger  impressed.  Spire  straightly  conical,  whorls  5J, 
slightly  convex,  the  last  angulated  at  the  periphery,  rather  abruptly 
descending  in  front.  Aperture  very  oblique,  light  brown  and  dotted 
within  ;  upper  margin  of  peristome  bent  downward,  bisinuate  ;  basal 
margin  narrowly  reflexed,  adnate  toward  the  center ;  a  dark  brown 
spot  at  the  root  of  the  columella.  The  lip  is  bordered  outside  with 
creamy,  behind  which  there  is  a  blackish  stripe. 

Alt.  20,  greater  diam.  20*,  lesser  18  mill. 

Fitzroy  and  Frankland  Is.  off  N.  E.  Australia ;  and  on  the  main- 
land 28  miles  inland  from  Cardwell,  Rockingham  Bay  (alt.  3500  ft.) 

H.  macgillivrayi  FORBES,  Voy.  H.  M.  S.  Rattlesnake,  ii,  p.  337, 
t.  3,  f.  1 — REEVE,  Conch.  Icon.  f.  357. — Cox,  Monogr.  Austr.  L.  Sh. 
p.  62,  t.  2,  f.  12.— PFR.,  Monogr.  iii,  p.  168  ;  Conchyl.  Cab.  t.  140, 
f.  1,  2. — BRAZIER,  Proc.  Linn.  Soc.  N.  S.  Wales,  i,  p.  125  ;  Jour, 
of  Conchol.,  Leeds,  i,  p.  269,  1887. — HEDLEY,  Proc.  Roy.  Soc.  N.  S. 
Wales,  v,  p.  63.— TRYON,  1.  c.  p.  135,  1888. 

A  beautiful  species,  allied  to  H.  naso  and  some  other  Papuan  spe- 
cies in  form  of  the  lip,  but  totally  different  in  sculpture  and  color- 
ing. 

H.  BIDWILLI  Pfeiffer.     PI.  3,  figs.  52,  53. 

Imperforate,  elevated-trochiform,  the  spire  straightly  conic,  periph- 
ery keeled,  base  flat;  lusterless  whitish,  irregularly  spotted  with 
chestnut  above,  the  base  chestnut,  spotted  with  white;  peristome 
dark  chestnut. 

The  surface  is  decussated  by  very  fine  oblique  and  spiral  stride,  and 
besides  is  very  obsoletely  obliquely  wrinkled.  Whorls  51,  nearly 
flat,  the  last  angular,  a  little  descending  in  front,  flat  below,  con- 
tracted at  the  aperture.  Aperture  very  oblique,  somewhat  angled 
at  the  position  of  the  carina,  pinkish-brown  within  ;  peristome  edged 


BELIX-PAPUINA. 

will)  hlaekiMi-biown.  outer  margin  not  expanded  nor  thickened, 
ha<al  mar-in  narrowly  expanded,  columellar  margin  arcuate,  re- 
tlexe.l  ami  closely  appressed.  Parietal  wall  dark  brown. 

Ah.  I8i,  greatest  diani.  16.},  lesser  15  mill. 

Maryborough,  //^irirh.  Plmpamn  and  Burnett  River,  Queensland; 
Richmond  River,  A".  S.  Wales,  Australia.  On  trees. 

//.  britlirl/li  PI-MI.,  P.  Z.  S.  1853,  p.  49  ;  Monogr.  iii,  p.  644.— 
REEVE,  Conch.  Icon.,  f.  1034.— H.  bidwilli  Cox,  Monog.  Austr. 
Land  Sli.,  p.  63,  t.  2,  f.  3.— PFR.  Nomencl.  Hel.  Viv,  p.  193.— 
BEDLEY,  Proc.  K«»y.  Soc.  Queensl.  v,  p.  62,  and  p.  135,  1888. 

The  name  was  mis-spelled  by  Pfeifter  in  his  earlier  publications, 
hut  the  strict  rules  of  nomenclature  may  perhaps  be  "  more  honored 
in  the  breach  than  in  the  observance/1  in  this  case;  and  I  have 
therefore,  accepted  Cox's  emendation. 

H.  NASO  Martens.     PL  2,  figs.  32,  33,  34,  35. 

I  in  perforate,  depressed-trochiform  ;  spire  conic  or  concave-conic; 
la-t  whorl  conspicuously  carinated  in  the  middle,  abruptly  and 
deeply  descending  to  the  aperture.  Sculptured  with  inconspicuous 
th-lines  and  finely  obliquely  corrugated  in  a  direction  at  right 
to  the  lines  of  growth.  White  or  rosy-white,  having  spiral 
hands  <>r  irregular  radiating  patches  of  purplish-rose  color  above. 
Aperture  very  oblique,  rosy-white  inside,  the  parietal  wall  blackish  ; 
,  //x  iipjter  margin  forming  two  high  arches,  produced  down- 
them  in  an  acute  triangular  lobe ;  columellar  lip  broad, 
adherent  to  the  base. 

There  are  nearly  five  whorls,  the  earlier  one  or  two  pink  or  pur- 
ple with  a  sutural  white  band.  The  last  half  of  the  body-whorl  is 
t  narrowed,  the  keel  becoming  more  obtuse;  it  is  very 
M\ollcn  just  behind  the  aperture.  The  lip  is  broadly  black- 
liorderrd.  Alt.  21,  diani.  31  mill. 

T'lltiiri,  Axtr»/<ih(;  Bay ;  ^ft.  Astrolabe,  Southeast  New  Guinea. 

If,  //</xo  v.  MART.,  Jahrb.  D.  M.  Ges.  x,  p.  82,  1883. — H.  tappa- 
ronei  \\.  A.  SMITH,  Ann.  Mag.  N.  H.  (5th  ser.,)  xi,  p.  190,  March, 
1*^  TAPP.  CAN.,  Ann.  Mus.  Civ.  Genov.  xxiv,  p.  133. 

A  heautil'iil  >pecie>,  jieeiiliar  in  thebisinuate  black  lip.     It  repre- 
Qta  M  line  of  differentiation  intermediate  between  H.  diomedes  and 
//.  t>iiil<ii'inun.     The  specimens  before  me  (figured  on  plate  2)  ex- 
hibit .-..n-idenihle  divei>ity  in   color-pattern.     Although  published 


HELTX-PAPUINA.  57 

ill  the  same  year,  the  name  naso  has  about  a  month  priority  over 
tapparonei. 

H.  RHYNCHONELLA  Tapparone  Canefri.     Unfigured. 

Imperforate,  thin,  depressed-trochiform,  carinated,  obsoletely 
longitudinally  striated  and  obliquely  spirally  roughened  ;  shining, 
violaceous  above,  buff  beneath,  painted  all  over  with  brownish- 
purple  flames  and  spots  disposed  in  bands.  Spire  couoidal,  apex 
brown,  whorls  5  ?,  the  upper  ones  plane,  the  last  whorl  large,  some- 
what convex,  at  the  periphery  acutely  carinated,  at  the  carina  sub- 
excavated  ;  base  convex,  with  narrow  spiral  bands  of  brownish- 
purple.  Aperture  very  oblique,  produced  into  a  beak,  wider  than 
high,  the  throat  pale-rose ;  peristome  blackish-purple,  moderately 
reflexed,  the  upper  margin  deeply  sinuated  forward,  basal  margin 
sub-angulated  in  the  middle,  columellar  margin  sub-dilated. 

Alt.  18,  greater  diam.  32,  lesser  24  mill.  Aperture,  width  19, 
alt.  12  mill.  (Tapp.  Can.) 

Ansus,  Island  of  Jobi,  New  Guinea. 

H.  rhynehonella  TAPP.  CAN.,  Annali  Mus,  Civ.  Genov.  xxiv,  (2d. 
Ser.  iv),  p.  134,  1887. 

Has  much  resemblance  to  H.  tayloriana,  but  is  larger,  the  in- 
flection of  the  upper  lip  more  marked,  and  the  coloration  different. 

H.  HERO  Smith.      Unfigured. 

Shell  rimate,  depressed-trochiform,  rather  thin,  acutely  carinated 
in  the  middle  ;  whitish,  ornamented  above  with  oblique,  opaque 
buffish  subzigzag  lines,  below  with  a  few  concentric  brown  and 
opaque  buff  zones.  Whorls  4^,  the  upper  a  little  convex,  closely 
spirally  striate  and  having  oblique  lines  of  growth,  the  last  whorl 
less  convex,  similarly  striated,  but  minutely  rugose  forward,  shortly 
descending,  lightly  depressed  before  the  middle  of  the  lip,  at  the 
periphery  having  an  acute,  buff  carina,  beneath  radiately  and  con- 
centrically subrugosely  striated.  Aperture  very  oblique,  subrostrate, 
colored  the  same  inside  as  out ;  peristome  white  or  pale  lilac  above, 
upper  margin  lightly  reflexed,  depressed  in  the  middle,  lower  mar- 
gin widely  expanded,  white,  dilated  above  the  umbilicus,  nearly 
closing  the  rimation.  Alt.  16J,  greater  diam.  28,  lesser  22  mill. 
(Smith.) 

Neiv  Guinea. 

H.  (Papuina)  hero  SMITH,  Ann.  Mag.  N.  H.  6th  series,  vol.  vii, 
p.  45,  May,  1891. 


58  HELIX-PAPUINA. 

The  apical  whorls  of  this  very  pretty  species  and  the  region 
an»un»l  tin-  umbilicus  are  opalescent  white  and  devoid  of  the  oblique 
marking  and  bauds.  In  the  type,  the  upper  terminations  of  the 
brownish,  oblique,  irregular  stripes  on  the  body-whorl  are  almost  of 
a  reddish  tint,  and  form,  as  it  were,  a  series  of  spots  at  the  suture. 
A  variety  i>  .-i'a  uniform  opalescent  or  greenish-white  color,  varie- 
gated above  with  the  radiating,  somewhat  zigzag,  opaque,  deep 
civam-colored  markings,  which  on  the  lower  surface  assume  the  form 
of  /.on*-  and  irregular  spots  or  blotches.  In  form  this  species  bears- 
resemblance  to  H.  tayloriana  Ads.  &  Rve.  (Smith.') 


H.  IANTHE  Smith.     Unfigured. 

Shell  depressed-trochiform,  narrowly  rimate,  acutely  carinated  at 
the  periphery,  entirely  white,  sculptured  with  delicate  oblique  lines 
of  growth.  Whorls  4,  the  upper  a  little  convex,  the  last  slightly 
concave  above  and  below  the  carina,  scarcely  descending  in  front. 
Aperture  oblique,  subquadrate,  acuminate  at  the  carina  ;  peristome 
slightly  reflexed  above,  broadly  expanded  below,  the  columellar 
margin  widely  dilated,  nearly  closing  the  umbilicus.  Spire  short- 
conic,  obtuse  at  the  apex.  Alt.  13$,  greater  diam.  25i,  lesser  21 
mill.  (Smith.) 

New  Guinea. 

II.  (Papuind)  ianthe  E.  A.  SMITH,  Ann.  Mag.  N.  H.  1891,  p.  452. 

In  form  this  species  is  very  like  H.  hero,  but  more  obtuse  at  the 
apex,  and  the  aperture  is  not  quite  so  acuminate.  It  differs  also 
from  that  species  in  the  absence  of  color-markings  and  spiral  striae. 
The  last  whorl  also  is  a  trifle  narrower,  and  does  not  descend  in 
front.  (Smith.') 

II.  TAYLORIANA  Adams  and  Reeve.     PI.  17,  figs.  40,  41  ;  pi.  2,  figs. 
20,21,27. 

[mperforate,  trochiform,  thin,  the  spire  conic;  apex  mammillar, 
obtuse,  whorls  5,  the  last  acutely  keeled,  abruptly  deflexed  in  front. 
Surface  shining,  finely  obliquely  roughened  and  corrugated  by  ob- 
liquely .l.^ci-ndin^,  anastomosing  impressed  lines.  Cream  colored, 
bernmiii'j  HI  -h-t  iiited  on  the  spire,  and  yellower  toward  the  latter 
part  of  the  body  whorl,  more  or  less  closely  speckled  and  obliquely 
streaked  with  pinkish-brown  above,  the  base  having  a  band  of  the 
same  »-..h»r,  obsolete  in  front  of  the  mouth,  where  there  is  a  tract 
tinted  with  pink,  which  extends  inward  on  the  parietal  wall  ;  carinal 


HELIX-PAPUINA.  59 

band  white  or  cream  colored ;  the  whole  interior  of  the  mouth  being 
pink,  except  a  broad  white  band  at  the  position  of  the  carina ;  lip  deep 
purplish  or  brownish-black,  bisinuate  above,  prominently  beaked 
at  the  keel,  reflexed  below,  columellar  lip  sloping,  adnate  to  the 
base. 

Alt.  17,  diam.  26  mill,  (typical  form). 

Alt.  13,  diam.  20  mill.  (var.  yulensis}. 

D' Entrecasteaux  Is. ;  Southern  New  Guinea,  at  South  Cape ; 
Laloki  (or  Goldie]  River,  foot  of  ML  Astrolabe;  foot  of  Owen  Stan- 
ley Ranges,  and  Yule  Island. 

H.  tayloriana  ADAMS  &  REEVE,  Voy.  Samarang,  Moll.  p.  59,  t. 
15,  f.  2.— REEVE,  Conch.  Icon.  f.  524.--E.  A.  SMITH,  Ann.  Mag. 
N.  H.  5th  ser.,  xix,  p.  421, 1887. — H.  yulensis  BRAZIER,  Proc.  Linn. 
Soc.  N.  S.  Wales,  i,  p.  105,  1876.— TAPP.  CAN.,  Ann.  Mus.  Civ. 
Genov.  xix,  p.  123,  t.  3,  f .  2 ;  t.  vi,  f.  1  ;  t.  viii,  f.  11  (Anatomy).— 
Cox,  Proc.  Linn.  Soc.  N.  S.  Wales,  2nd  ser.  ii,  p.  1063  t.  2,  f.  5,  6  ; 
1888. 

This  species  is  closely  allied  to  H.  strabo,  but  apparently  con- 
stantly separated  by  the  creamy  peripheral  band  and  deep  purplish- 
black  peristome.  The  type  is  figured  on  pi.  17,  figs.  40,  41. 

The  variety  called  yulensis  by  Brazier  (pi.  2,  figs.  28,  29,  30,  31) 
is  smaller,  with  the  "  spout"  of  the  aperture  less  developed.  It  must 
however  be  regarded  as  a  variety  or  form  of  the  tayloriana. 

H.  ALBOCARINATA  Smith.     PL  1,  fig.  5. 

Short-conical,  imperforate,  thin,  semi-pellucid,  pale  corneous,  en- 
circled by  an  opaque-white  carina ;  whorls  5,  moderately  convex, 
obliquely  roughly  striate,  the  last  whorl  acutely  keeled  at  the  peri- 
phery, sculptured  below  with  rugose  concentric  strise,  ornamented 
with  few,  interrupted,  white  opaque  lines,  shortly  descending  in 
front,  margined  with  buff  or  milk-white  behind  the  lip.  Aperture 
oblique ;  peristome  white,  narrowly  expanded  and  reflexed,  outer 
margin  sinuous,  columellar  margin  thickened,  appressed  parietal 
callus  thin.  Alt.  17,  greater  diam.  24,  lesser  20  mill.  (Smith.} 

South  Cape,  Neiv  Guinea. 

H.  albocarinata  SM.,  Ann.  Mag.  N.  H.  5th.  Ser.  xix,  p.  423,  t. 
15,  f.  12,  1887. 

The  central  keel  is  rather  sharper  and  the  texture  more  transparent 
than  in  H.  tayloriana,  but  it  may  prove  to  be  an  albino  form  of  that 
species. 


(50  HI.LIX-PAPUINA. 

H.  STRABO  Brazier.     PI.  2,  figs.  22,  23  (typical);   figs.  17,  18,  19, 

21.  25,  •_>»;  (row.). 

Iniperforate,  low-trochiform,  spire  conical,  apex  obtuse;  whorls 
I  .  the  hist  one  acutely  carinated,  slightly  deflexed  in  front.  Sur- 
face shinini:,  obliquely,  finely  and  irregularly  corrugated,  as  in  H. 
t<m?»n''ina.  Color  creamy-white,  unicolored  or  with  spiral  bands, 
the  parietal  wall  and  base  in  front  of  the  aperture  flushed  with  pink. 
Ap'-rtmv  very  oblique  ;  lip  broadly  bordered  with  pink  (or  purple), 
bi>imiate  above,  produced  into  a  "spout'  or  beak  at  the  keel, 
expanded  below,  the  columellar  margin  adherent  to  the  base. 

Alt.  18,  diam.  .SO  mill. 

Southern  Xeiv  Guinea  at  Katau  River,  and  Maclachie  Point. 

H.  strabo  BRAZIER,  Proc.  Linn.  Soc.  N.  S.  Wales,  i,  p.  106,  1876. 

-//.  rotent'ibiata  E.  A.  SMITH,  Ann.  Mag.  N.  H.  xix,  p.  421,  t.  15, 
f.  2. — H.  katauensis  TAPF.  CAN.,  Annali  Mus.  Civ.  Genov.  xix,  p. 
126,  t.  3,  f.  1,  3  ;  t.  vi,  f.  3.  (Anatomy). 

Distinguished  from  the  closely  allied  H.  tayloriana  by  its  rosy  or 
lilac-purplish  lip,  absence  of  a  well-defined  peripheral  white  girdle, 
and  general  pattern  of  coloring.  The  apex  is  either  black,  corneous 
or  white.  The  following  color-varieties  have  but  slight  claims  to 
even  varietal  rank  ;  the  large  series  before  me  showing  great  varia- 
tion-  in  color-pattern,  as  well  as  typical  examples  of  the  several 
described  forms. 

The  typical  strabo  (pi.  2,  figs.  22,  23)  is  unicolored  creamy-white 
witli  rose-pink  lip.  Four  of  the  specimens  before  me  have  the  lip 
purple. 

Form  roseolabiata  Smith  (pi.  2,  figs.  17,  18).  Creamy-white  ; 
apical  whorls  blue-black;  periphery  girdled  with  a  sharply 

brown  band,  another  band  bordering  the  suture  ;  lip  rosy. 
I>laml,  D'Entrecasteaux  Group. 

l-'«.nn  kufiiuanxis  Tapp.  Can.  (pi.  1,  figs.  14,  15;  pi.  2,  figs.  24, 
I'.").  •_'•'•  .  Variously  obliquely  streaked  and  having  spiral  bands  of 
purplish-brown  ;  lip  pink. 

II.  RHOMBOSTOMA  I'f'eiifer.      PI.  16,  figs.  14,  15. 

[mperforate,  trodiiibnn,   rather  thin,  obliquely  striatulate,  sub- 
<!<•( -u-at. ..I  with  very  close  concentric  lines,  rather  shining,  whitish- 
tawny,  ornamental  with  a  number  of  chestnut  bands;  spire  short, 
ape\  acute.     NVhorU  5,  -rather  flat,  the  last  acutely  carinated, 
•1'iL'htly    convex     beneath,    little   descending   in    front;    peristome 


HELIX-PAPUINA.  61 

violaceous,  upper  margin  expanded,  impressed  above,  columellar 
margin,  straightened,  dilated,  flat,  appressed.  Alt.  15,  greater  diam. 
28,  lesser  23  mill.  (P/r.) 

Habitat  unknown. 

H.  rhombostoma  PFR.,  P.  Z.  S.  1845,  p.  72;  Monogr.  i,  p.  231. — 
RVE.,  Conch.  Icon.,  f.  1456. 

This  form  has  not  been  noticed  by  recent  writers.  It  is  apparently 
intermediate  between  H.  strabo  Braz.  and  H.  louisiadensis  Forbes, 
approaching  rather  near  to  the  var.  katauensis  of  the  former.  Should 
be  looked  for  in  Southern  New  Guinea  and  adjacent  islands. 

H.  GURGUSTII  Cox.     PL  1,  fig.  16 ;  PI.  14,  figs.  68,  69. 

Shell  imperforate,  broadly  conoid,  white,  diaphanous,  opaque, 
shining,  granular  on  the  surface,  whorls  five,  gradually  increasing 
in  size,  flat,  last  sharply  angled  at  the  periphery,  pinched  and 
everted  at  the  peristome,  reflexed  at  the  insertion  ;  base  flat ;  peri- 
stome  bright-pink,  Innately  elongated,  margins  approached,  everted 
and  beaked  at  the  center  ;  aperture  white  within. 

Diam.  greatest  I'lO  ;  least  0.84  ;  height  0*80  of  an  inch. 

This  fine  species  is  in  the  Hargravesian  Collection  in  the  Austra- 
lian Museum  ;  it  is  the  same  kind  of  shell  as  Helix  Louisiadensis  of 
MacGillivray,  but  is  a  larger  species,  easily  distinguished  from  that 
species  by  its  white  diaphanous  aspect,  light-pink  peristome  and  by 
the  absence  of  the  characteristic  oblique  fine  strise  on  the  surface  of 
the  whorls.  (Cox.*) 

Russell  Island,  Louisiade  Group. 

H.  (Geotrochus*)  gurgustii  Cox,  Proc.  Linn.  Soc.  N.  S.  Wales,  iv, 
p.  114,  t.  16,  f.  1,  1879 ;  I  c.  2d.  Ser.  ii,  p.  1063,  t.  2,  f.  3,  4. 

H.  LOUISIADENSIS  Forbes.     PI.  4,  figs.  61,  62,  63. 

Imperforate,  depressed-turbinate,  about  equally  convex  above  and 
below  the  periphery  ;  spire  low-conic,  apex  obtuse  ;  whorls  4-2-4 :|, 
slightly  convex,  the  last  whorl  somewhat  angulated  in  front,  becom- 
ing rounded,  abruptly  deflexed  to  the  aperture,  a  little  constricted 
behind  the  pink  lip. 

Surface  shining,  obliquely  striatulate,  and  covered  with  a  close  fine 
sculpture  of  fortvard-descending  wrinkles.  Ground-color,  corneous- 
white  or  pink,  having  an  opaque-white  peripheral  girdle,  above  ob- 
liquely streaked  in  a  very  irregularly  broken  pattern  with  light  brown ; 


62  HELIX-PAPUINA. 

streaked  and  indistinctly  spirally  banded  with  the  same  shade  below 
the  peripheral  zon.-.  Earlier  whorls  unicolored,  corneous  or  whitish. 
Aperture  oblique,  peristorue  pink,  its  face  thickened,  convex;  nar- 
rowly expanded,  reflexed,  sinuous. above;  columella  arcuate,  adnate, 
with  n<>  tooth  or  fold.  Alt.  18,  greater  diam.  21,  lesser  17*  mill. 

Sudest  Id.,  Louisiade  Is. 

II.  louisiadensis  FORBES,  Voy.  Rattlesnake,  appendix,  p.  376,  t, 
2,  f.  ^  I'ri:-.  Monogr.  iii,  p.  174. — REEVE,  Conch.  Icon.  f.  1449. — 
//.  millidntcB  Cox,  P.  Z.  S.  1871,  p.  323,  t.  34,  f.  2,  2a;  1.  c.  1873,  p. 
566.— PFR.  Monogr.  vii,  p.  313.— H.  Thomsoni  E.  A.  SMITH,  Ann. 
Ma-.  N-  H.  1889,  p.  202,  t.  13,  f.  12,  13. 

This  species  has  the  lip  like  H.  fringilla,  but  the  spire  is  conical 
and  the  surface  finely  wrinkled.  Slightly  differing  varieties  occur 
on  the  different  islands. 

Var.  MILLICENTJE  Cox.     PI.  4,  figs.  64,  65. 

Differs  from  the  type  in  having  the  lip  white,  and  more  produced 
forward  above. 

Var.  THOMSONI  Smith.     PI.  7,  figs.  31,  32. 

I m perforate,  subconic-globose,  rosy-purple,  spotted  and  variegated 
with  pale  buff,  closely  sculptured  with  lines  of  growth,  and  minutely 
obliquely  corrugated  ;  whorls  4J,  a  little  convex,  rapidly  widening, 
separated  by  simple  sutures,  the  last  whorl  at  first  carinated,  the 
carina  becoming  obsolete,  at  the  aperture  suddenly  deflexed,  con- 
st ri< -ted  behind  the  lip;  aperture  oblique,  elongated,  rosy -purple 
inside  ;  peristome  whitish,  expanded  and  reflexed,  upper  margin 
siniiated  forward  ;  columellar  margin  appressed,  obliquely  recti- 
lineal- within.  Alt.  18,  greater  diam.  27,  lesser  20  mill.  (Smith.) 
Var.  Shell  subdiapnanous,  varied  and  spotted  with  buff. 

St.  Aignan,  Louisiade  Group. 

H.  wnom.AKKiANA  Souvcrbie.     PI.  3,  figs.  42,  43. 

[inperforate,  semiglobose,  obliquely  striate,  very  finely  granulose- 
nialli-atf,  ln-rr  and  there  very  obsoletely  spirally  striate  ;  rather  thin, 
shining,  subtranslucid,  violaceous,  darker  at  the  apex,  paler  at  the 
umbilical  r« -j-iun,  having  four  whitish  bands,  the  three  upper  more 
01-  Irs-  interrupted  and  brown,  the  fourth  is  wider  and  continuous, 
at  tin-  periphery.  Circuin-mnbilical  area  paler,  having  three  narrow, 
ill-delim-d  whitish  band-.  Whorls  4,  convex,  the  suture  impressed, 
tin-  la-t  whorl  earinated,  mure  convex  beneath,  a  little  descending  in 


HELIX-PAPUINA.  63 

front,  subconstricted  behind  the  whitish-yellow  lip.  Aperture  con- 
colored  within,  oblique,  subrhornbic-lunar  ;  per  is  to  me  acute,  white, 
superior  margin  arched  forward,  subreflexed,  becoming  more 
reflexed  toward  the  columella,  which  suddenly  becomes  appressed  to 
the  base,  and  is  of  a  rosy  (slightly  violet)  color.  Alt.  14^,  greater 
diam.  19i,  lesser  15  mill.  (Souv.) 

WoodlarJc  Island,  Louisiade  Group. 

H.  woodlarkiana  Souv.,  Journ.  de  Conchyl.  1863,  pp.  76,  172,  t. 
5,  f.  2. — PFR.,  Monogr.  v,  p.  271. 

H.  GORENDUENSIS  Brazier.      Unfigured. 

Shell  imperforated,  trochus-shaped,  rather  thin,  obliquely,  finely 
striated  and  transversely  wrinkled,  flesh  tinted  or  cream  color ;  orna- 
mented with  pinkish  opaque  spots  and  dots ;  spire  rather  conoid  ; 
whorls  5,  convex,  the  last  more  convex,  the  first  three  apical  whorls 
dark  rose-pink,  the  fourth  a  little  lighter  in  color,  the  fifth  slightly 
keeled  in  front,  cream  colored,  and  marked  with  opaque  pinkish 
spots  and  dots,  base  convex,  sculptured  same  as  above ;  aperture 
triangular,  very  much  produced  and  contracted  in  front,  constricted 
behind  the  aperture,  interior  bright  pink,  peristome  blackish-purple, 
slightly  reflected  ;  the  right  margin  descending  in  front,  columellar 
margin  flattened  and  expanded,  tinged  with  brown,  margins  joined 
with  a  thin  pink  callus  entering  spirally  into  the  interior  of  the 
aperture. 

Two  specimens  of  this  very  pretty  species  were  found  by  Baron 
Miklouho-Maclay.  One  was  dead  and  weatherbeaten,  the  other  was 
found  with  the  animal  alive.  The  Baron  informs  me  that  the  ani- 
mal was  black,  and  that  the  back  of  the  neck  was  brown.  (Braz.} 

Alt.  15,  greater  diam.  25,  lesser  20  mill. 

Gorendu,  Maclay  Coast,  Neiv  Guinea. 

H.  ( Geotrochus)  gorenduensis  BRAZIER,  Proc.  Linn.  Soc.  N.  S. 
Wales,  x,  p.  841,  1886. 

H.  ROLLSIANA  Smith.     PI.  1,  fig.  12. 

Shell  imperforate  conoid-subglobose,  whitish  or  rubescent,  marked 
with  brown  at  the  suture,  variously  banded  with  brown  zones,  and 
having  a  streak  of  orange  behind  the  lip.  Whorls  5,  a  little  con- 
vex, sculptured  with  oblique  close  rough  striae  and  lines  of  growth; 
last  whorl  rounded  on  its  latter  part,  obtusely  carinated  in  front  of  the 
mouth,  always  zoned  with  white,  tinted  with  purple-rose  beneath,  in 


64  HELIX-PAPUINA. 

front    shortly  and    abruptly    descending.     Aperture   oblique,    sub- 
<iuadrute  :  prristome  white,  thin,  broadly  expanded  and  reflexed,  the 
upper  mar-in  simiated,  colurnellar  margin  dilated,  appressed. 
Alt.  •_'<>.  -n-ater  diam.  30-i,  lesser  2H  mill.     (Smith.') 

South  Cape,  British  New  Guinea. 

II.  (r'<ij>nin<D  KoUslaua  E.  A.  SMITH,  Ann.  Mag.  N.  H.  5th.  Ser. 
xix.  p.  423,  t.  15,  f.  3,  1887. 

II.  mMRiEi  Angas.     PI.  17,  figs.  37,  38. 

Shell  i  in  perforate,  conically  semi-globose,  rather  solid,  rugosely 
spirally  grooved,  and  obliquely  finely  striated,  opaque,  whitish  ; 
spire  conoidal,  obtuse  ;  whorls  4*,  rather  convex,  the  last  descending, 
subangulated  at  the  periphery,  somewhat  flattened  at  the  base,  and  a 
little  excavated  behind  the  aperture  ;  aperture  diagonal,  oval-oblong, 
pale  brown  within,  peristome  white,  margins  converging,  united  by 
a  thin  callus,  right  margin  sinuous,  rather  expanded  and  reflexed, 
columellar  margin  dilated  and  appressed.  (Adams  &  Angas.} 

Ah.  20,  greater  diam.  27,  lesser  21  mill. 

Shores  of  Huon  Gulf,  Eastern  New  Guinea. 

H.  comriei  ANGAS,  P.  Z.  S.  1876,  p.  489,  t.  47,  f.  4,  5.—  TAPP. 
(  'AN.,  Ann.  Mus.  Civ.  (lenov.  xix,  p.  155,  (transl.  of  desc.) 

(11.)    GROUP  OF  H.  EDDYSTONENSIS. 

U'Hit'ifi'-ate  ;  low-trochiform,  the  base  convex;  outer  lip  expanded, 
basal  lip  reflexed,  expanded  at  the  insertion  and  partly  concealing 
the  umbilicus;  surface  sculptured  with  fine  forward-  descending 
ir  r  inkles.  Columella  often  havin  a  callus  fold  or  obtuse  tooth. 


II.  iiDDvsToNKNsis  Reeve.     PI.  10,  figs.  79,  80. 

Umbilicate;  low-trochiform,  solid,  angulated  or  sub-carinate  at 
tin  periphery.  Surface  striatulate  and  closely  sculptured  with  fine 
forward-descending  wrinkles,  becoming  concentric  on  the  base. 
Spire  low-conic,  apex  obtuse  ;  earlier  whorls  ivhitish-corneous  ; 
whorl>f>,  somewhat  convex,  the  last  not  descending  in  front,  obtusely 
keeled  at  the  circumference.  Color  opaque  buff,  with  a  lighter  peri- 
pheral /one  having  a  brown  band  above  and  below  it,  and  often  a 
few  othi-r  obscure  bands  both  above  and  beneath. 

A  pei-tun-  oblique,  within  showing  the  white  and  dark  bands  and 
niewhat    -treaked    with    brown.     Peristome   white    (often    tinted 


\ 


HELIX-PAPUINA.  65 

where  the  bands  terminate),  outer  lip  broadly  expanded,  columellar 
lip  broad,  dilated,  about  half  closing  the  umbilicus. 
Alt.  17,  greater  diam.  23,  lesser  18*  mill. 

Eddy  stone  and  Simbo  Islands,  Solomon  Group. 

H.  eddystonensis  REEVE,  Conch.  Icon.  f.  1384 ;  1854. — DOHRN, 
in  Conchy  1.  Cab.  p.  568,  t.  168,  f.  5,  6. — H.  motacilla  SMITH,  P.  Z. 
S.  1885,  p.  591. 

Usually  has  two  rather  narrow  brown  bands,  bounding  a  peripheral 
whitish  zone.  There  is  an  ill-defined  opaque  whitish  band  below 
the  suture.  The  variations  include  forms  with  one  or  several  ad- 
ditional narrow  bands  above  and  below,  and  some  specimens  are 
bandless,  unicolored  straw-yellow. 

H.  GELATA  Cox.     PI.  10,  figs.  93,  94. 

Shell  conoidly  depressed,  rather  narrowly  umbilicated,  of  a  dark 
brown  color,  profusely  ornamented  and  zoned  with  opaque  white, 
apex  almost  black,  transversely  striated  from  left  to  right  with  very 
fine  strait-strise,  which  are  decussated  above  with  slightly  undulating 
coarser  stride  from  right  to  left,  at  the  base  these  undulating  striae 
become  longitudinal ;  whorls  5,  convex,  suture  deep  ;  base  convex  ; 
aperture  rotundately  lunar,  dark  chestnut  within  ;  lip  white,  margins 
somewhat  approximating,  joined  by  a  thin  callus ;  upper  margin 
broadly  expanded,  basal  reflected,  columellar  margin  triangularly 
dilated  and  reflexed,  half  concealing  the  umbilicus.  (Cox.) 

Diam.  greatest  0'87,  least  0*71,  height  0'68  of  an  inch. 

A  small  island  near  Eddystone  Id.,  Solomon  Group. 

H.  gelata  Cox.,  P.  Z.  S.  1873,  p.  149,  t.  16,  f.  5a,  5b.— BRAZIER, 
Proc.  Linn.  Soc.  N.  S.  Wales,  v,  p.  446,  (with  var.  maddocksi.) 

Closely  allied  to  and  perhaps  a  variety  of  H.  eddystonensis,  from 
which  the  dark  apical  whorls  and  more  variegated  color  separate  it. 
Cox  says : 

I  was  at  first  disposed  to  look  upon  it  as  a  variety  of  Helix  eddy- 
stonensis (Reeve),  but  the  sculpture  of  this  shell  appears  to  me  to  be 
always  coarser,  and  the  surface  invariably  more  or  less  covered  with 
a  thick  white,  opaque  enamel,  unlike  the  thin  epidermis  which  covers 
H.  eddystonensis.  Its  dark,  almost  black,  apex  and  white  zoned 
and  irregularly  ornamented  body  show  in  strong  contrast,  and 
resemble  more  some  of  the  Philippine  Island  species  than  any  of  those 
found  in  the  Solomon  Islands.  This  species  is  uniformly  of  a  smaller 
5 


66  HELIX-PAPUINA. 

size  than  H.  eddystonensis,  and  is  more  solid  and   more  raised  in 

the  spire.     (Cox.} 

Yar.  MADDOCKSI  Brazier.     PI.  16,  figs.  11,  12,  13. 

The  form  is  like  H.  eddystonensis,  from  which  this  variety  differs 
in  having  the  upper  whorls  red  or  dark  purple.  It  is  straw-yellow, 
unicolored,  or  with  one  or  two  bands  at  the  obscurely  angled  periph- 
ery. The  lip  is  broadly  expanded,  white.  Sculpture  as  in  H.  eddy- 
stonensis. 

Small  island  near  Eddystone  Id. 

Brazier  found  this  form  much  more  abundant  than  typical  gelata, 
which  inhabits  the  same  island.  He  enumerates  many  varieties  of 
striping  (see  below).  The  figures  on  my  plate  are  drawn  from 
authentic  specimens. 

Yar.  a.  Light-yellow,  ornamented  with  one  broad  chestnut-baud 
on  the  periphery  and  continuous  at  the  suture. 

Yar.  b.  Darker  yellow,  ornamented  on  the  base  with  two  fine 
lines  on  the  periphery,  two  wider  and  a  fine  one  below  the  suture. 

Yar.  c.  Nearly  white,  base  with  one  broad  band  nearly  black, 
periphery  and  upper  part  ornamented  with  three  broad  ba.nds. 

Yar.  d.  Light-brown,  base  ornamented  with  one  narrow  chest- 
nut line,  periphery  with  two  rather  broad,  white  lines  at  the  suture  ; 
below  a  light-brown  intermingled  with  a  few  dark-brown  opaque 
spots. 

Var.  e.  Bright  straw-yellow,  two  narrow  reddish  bands  on  the 
periphery,  apex  light-red. 

Var./.  Light-yellow,  nearly  white,  the  periphery  ornamented 
with  one  narrow  reddish  line,  obsolete  on  the  upper  whorls. 

Yar.  rj.  Light-yellow,  with  opaque  white  band  on  the  periphery 
witli  a  fine  reddish  line  above  and  below  it. 

Var.  h.  Dark-yellow,  ornamented  with  numerous  opaque  lines 
of  a  light-yellow  on  the  periphery  and  base,  upper  whorls  white, 
a]>»-\  dark-red. 

Yar.  L  Light-straw-yellow,  upper  whorls  white,  apex  dark-red. 
(Brazier.} 

11.  MOTACILLA  Pfeiffer.     PI.  11,  figs.  6,  7. 

Umbilicate,  solid,  opaque,  low-trochiform,  obtusely  carinated  at 
the  p.-riphcry  ;  spire  conoidal,  apex  obtuse;  surface  striatulate  and 
ol.liqm-ly  rorruiriited  by  very  fine  forward-descending  wrinkles. 
<  'I'M-  a  whitish-brown,  with  an  opaque  white  girdle  at  the  periphery, 


HELIX-PAPUINA.  67 

and  usually  under  the  suture.  Whorls  5,  convex,  separated  by 
impressed  sutures  ;  the  earlier  two  whorls  darker,  of  a  reddish  color  ; 
last  whorl  a  trifle  descending  in  front,  angular  at  the  periphery, 
convex  beneath. 

Aperture  oblique,  transversely  oval,  light  and  having  brown 
stains  within  ;  lip  expanded,  white,  dilated  at  the  columellar  inser- 
tion. Alt.  16 £,  greater  diam.  22,  lesser  18 £  mill. 

Eddy  stone  Id.,  Solomon  Group. 

H.  motacilla  PFR.,  Monogr.  iv,  p.  259  ;  Novit.  Conch.,  t.  31,  f. 
12,  13. 

Differs  from  H.  eddystonensis  in  lacking  brown  bands  and  in  hav- 
ing a  dark  apex.  The  lip  is  narrower  than  in  eddystonensis  or 
maddocksi,  and  the  spire  rather  more  conoidal.  All  of  these  species, 
eddystonensis,  gelata,  maddocksi,  motacilla  are  very  closely  allied,  and 
should  perhaps  be  considered  as  varietal  forms,  rather  than  fully 
differentiated  species. 

H.  ANTRORSA  Pfeiffer.     Unfigured. 

Shell  umbilicated,  subconic,  thin,  striate  and  decussated  with  for- 
ward-descending wrinkles ;  pellucid,  corneous-white ;  spire  conoid, 
rather  obtuse ;  suture  margined  ;  whorls  5,  a  little  convex,  rapidly 
increasing,  the  last  compressed  and  acutely  cariuated,  nearly  flat 
above,  deflexed  from  the  carina  in  front,  the  base  narrowly  inflated 
around  the  umbilicus-.  Aperture  very  oblique,  lanceolate-lunar; 
peristoine  thin,  margins  slightly  converging,  the  upper  lip  expanded, 
subrostrate  at  the  outer  angle,  basal  lip  strongly  arcuate,  reflexed, 
dilated  above  the  umbilicus. 

Alt.  12,  greater  diam.  23,  lesser  19 £  mill.     (Pfr.) 

Haman  Island,  Admiralty  Group. 

H.  (Cymotropis)  vitrea  v.  MART,  in  Die  Heliceen  2d.  edit.,  p.  169, 
notH.  vitrea  Fer.,  vide  v.  Mart.,  Ostas.  Zool.  Landschn.,p.  319. — H. 
antrorsa  PFR.,  P.  Z.  S.  1855,  p.  113 ;  Monogr.  iv,  p.  254. 

H.  SACHALENSIS  Pfeiffer.     Unfigured. 

Shell  narrowly  umbilicated,  conoid,  thin,  striate,  subdecussated 
with  forward-descending  striulse;  tawny-whitish,  with  a  white 
peripheral  band,  above  it  a  narrow  chestnut  band,  and  some  brown 
ones.  Spire  conoid ;  whorls  4£,  moderately  convex,  the  last  not 
descending,  carinated  in  the  middle.  Aperture  oblique,  rhombic- 


i;s  HKLIX-PAPUINA. 

lunar  ;  peristome  thin,  margins  subconniving,  the  upper  arcuate,  ex- 
}>an<U-<],  basal  reflexed,  dilated  and  vaulted  at  the  umbilicus. 
Alt.  \~2,  greater  diam.  19-1,  lesser  15*  mill.     (Pfr.) 

Saehala  Island,  Admiralty  Group. 

H.  sachalensis  PFR.,  P.  Z.  S.  1855,  p.  114;  Monogr.  iv,  p.  259  ; 
/.  c.  v,  p.  331. 

Yar.  b.  Shell  a  little  more  elevated,  distinctly  carinated,  having 
2  or  3  blackish-chestnut  bands. 

This  unfigured  species  is  perhaps  allied  to  H.  eddystonensis  Rv. 

H.  LEUCOTHOE  Pfeiffer.     PL  12,  figs.  22,  23,  24. 

Narrowly  umbilicated,  trochiform,  rather  thin,  striatulate,  and 
having  forward-descending  stride  above,  becoming  concentric  on  the 
base ;  whitish  ;  spire  turbinate,  apex  rather  obtuse ;  whorls  4£,  con- 
vex, the  last  slightly  descending  in  front,  acutely  carinated  at  the 
periphery,  having  a  single  band  below  the  carina ;  aperture  oblique, 
rhombic-lunar;  peristome  expanded,  upper  margin  thin,  the  basal 
margin  thickened  in  the  middle ;  columellar  margin  dilated, 
purplish.  Alt.  12£,  greater  diam.  2(H,  lesser  17  mill.  (Pfr.) 

New  Georgia,  Solomon  Is. 

H.  leucothoe  PFR.,  P.  Z.  S.  1861,  p.  192  ;  Novit.  Conch.,  p.  179, 
t.  48,  f.  7-9  ;  Monogr.  v,  p.  330. 

H.  CyERtiLESCENS  Angas.     PI.  10,  figs.  91,  92, 

Shell  subperforate,  conoidal,  thin,  obliquely  striatedjand  the  lower 
whorls  decussated  by  irregular  undulating  stria? ;  bluish-gray,  with 
darker  bands  of  the  same  color,  changing  into  pale  yellowish-olive 
above  and  at  the  base,  the  apical  whorls  being  dark  chestnut  and 
with  a  flesh  colored  band  immediately  below  the  suture,  and  a 
similar  one  at  the  periphery,  with  a  narrow  dark  chestnut  line  in  the 
middle  ;  spire  conical,  rather  obtuse  at  the  apex ;  whorls  4£,  slightly 
convex,  the  last  descending  in  front,  angled  at  the  periphery,  aper- 
ture diagonal,  sublunar ;  peristome  white,  margins  converging,  the 
right  scarcely  flexuous,  expanded,  the  basal  one  arcuate,  reflexed. 

Diam.  niaj.  9,  min.  8,  alt.  8->  lines.     (Angas.) 

Guadalcanar  Is.  Solomon  group. 

H.  ccerulescens  ANGAS,  P.  Z.  S.  1869,  p.  624,  t.  48,  f.  6.— PFR., 
Monogr.  vii,  p.,  381. 

Evidently  allied  to  H.  lienardiana. 


HELIX-PAPUINA.  69 

H.  PUDICA  Pfeiffer.     Unfigured. 

Narrowly  umbilicated,  trochiform,  rather  thin,  striatulate,  having 
close,  forward-descending  strise  above,  beneath  decussated  with  sub- 
concentric  striae ;  buffish-fleshy ;  spire  conic,  rubescent  above,  apex 
rather  obtuse ;  suture  light ;  whorls  5,  slightly  convex,  the  last  not 
descending,  subacutely  cariuated  in  the  middle,  more  convex  on  the 
base,  having  a  single  chestnut  band  at  the  carina ;  somewhat  tumid 
in  front.  Aperture  very  oblique,  rounded-subrhombic ;  peristome 
subroseous  calloused,  upper  margin  a  little  expanded,  basal  flexuous, 
reflexed,  dilated  above  the  umbilicus,  provided  with  a  nodule  within, 
at  the  insertion. 

Alt.  14-15,  greater  diam.  22,  lesser  19  mill.     (Pfr.) 

New  Britian. 

H.  pudica  PFR.,  Mai.  Bl.  vii,  1860,  p.  236 ;  Monogr.  v,  p.  330. 
An  unfigured  species,  evidently  allied  to  H.  eros,  lienardiana,  etc. 

H.  LEINARDIANA  Crosse.     PI.  11,  figs.  13,  14,  15,  16,  17,  18,  19. 

Half-covered  umbilicate,  depressed-trochiform,  solid  but  rather  thin, 
about  equally  convex  above  and  below  the  acute  peripheral  keel. 
Surface  striatulate  and  covered  with  a  close,  fine  sculpture  of  for- 
ward-descending wrinkles,  becoming  concentric  beneath.  Spire  low- 
conic,  apex  obtuse. 

Whorls  4J,  rather  convex,  the  earlier  red  or  purple  (rarely 
whitish),  the  remainder  white  or  buff,  with  a  brown  band  above, 
usually  wide  or  split  into  two  or  three,  and  a  broad  band  below  the 
white  or  buff  peripheral  zone ;  the  suture  is  bordered  below  with  white 
or  buff.  The  upper  surface  sometimes  has  a  faint  bluish  band,  and 
scattered  purplish  dots. 

Aperture  oblique,  vividly  banded  inside;  peristome  narrowly  ex- 
panded, pink  at  the  terminations  of  the  bands  and  at  the  root  of  the 
columella ;  basal  and  columellar  lips  reflexed,  the  latter  straightened, 
having  a  callous  fold  in  the  middle,  and  dilated  half  over  the  rather 
large  umbilicus.  Alt.  15,  greater  diam.  21,  lesser  17*  mill. 

New  Georgia,  Solomon  Is. 

H.  lienardiana  CR.,  Journ.  de  Conchyl.  1866,  p.  53,  t.  1,  f.  1. — 
PFR.,  Monogr.  v,  p.  330 ;  Nomencl.  Hel.  Viv.,  p.  197. 

The  expanded  columella  is  pink  ;  the  brown  bands  are  generally 
very  vividly  contrasted  against  the  opaque  white  or  creamy  ground- 
color, but  the  upper  one  is  sometimes  faint,  the  surface  then  usually 


7Q  HELIX-PAPUINA. 

having  scattered  purplish-translucent  dots.  It  is  larger  and  more 
s.,lid  than  H.  eros,  with  less  expressed  keel,  more  vivid  coloring,  and 
wider  umbilicus. 

H.  EROS  Angas.     PI.  15,  fig.  98,  99,  100,  101. 

Narrowly  perforated,  depressed  trochoidal,  about  equally  convex 
al  ,o ve  and  below  the  acute  peripheral  keel  Surface  obliquely  striat- 
ulate,  densely  sculptured  with  close,  fine  forward-descending  wrinkles, 
becoming  concentric  on  the  base.  Spire  low-conoidal,  whorls  4*, 
convex,  the  earlier  2  or  3  red,  the  remainder  opaque,  buff,  with 
scattered  grayish-translucent  dots,  and  a  brownish  band  above  and 
below  a  whitish  peripheral  zone  Last  whorl  convex  above  and 
below,  generally  shortly  deflexed  in  front. 

Aperture  oblique,  subrhombic,  more  or  less  streaked  and  dotted 
inside  on  a  white  ground.  Peristome  thin,  pink,  expanded,  the  basal 
and  columellar  margins  reflexed,  the  latter  straightened,  having  an 
obsolete  fold  of  callus  in  the  middle  and  dilated  at  the  insertion, 
nearly  closing  the  narrow  umbilicus. 

Alt.  14,  greater  diam.  18,  lesser  14*  mill. 

Shortland,  Stephens  and  Isabel  2s.,  Solomon  Group. 

Geotrochus  eros  ANGAS,  P.  Z.  S.  1867,  p.  888,  t.  43,  f.  4-6.— IT. 
eros  PFR.,  Mon.  vii,  p.  389. — BRAZIER,  Journ.  de  Conchyl.  1880,  p. 
313.— SMITH,  P.  Z.  S.  1885,  p.  592. 

This  species  resembles  H.  fringilla  in  the  red  or  pink  upper  whorls, 
and  the  callous  fold  on  the  columellar  lip  ;  it  differs  in  being  per- 
forated, smaller,  thinner,  and  sculptured  in  a  totally  different 
manner.  It  is,  in  fact,  very  closely  allied  to  H.  lienardiana,  but  is 
smaller,  with  narrower  umbilicus  and  far  less  vivid  coloring.  Lives 
upon  trees. 

II.  REDEMPTA  Cox.      PI.  3,  figS.  40,  41. 

Shell  obicularly  convex,  imperforate,  obliquely  finely  striated 
from  above  downwards  and  backwards  with  strait  stria?,  and  on  the 
last  two  whorls  from  behind  forwards  with  interrupted  malleated 
strire,  of  a  dull  opaque  white  color,  apex  pink,  variously  ornamented 
with  irregular  dark  chestnut  spots,  or  with  two  or  more  regular 
broad  bands;  spire  bluntly  coniform;  whorls  5,  slightly  convex,  last 
sharply  keeled  and  shortly  deflected  in  front,  base  flattened  ;  aperture 
quadrilateral,  beaked  at  the  periphery  of  the  last  whorl,  margins 
joined  by  a  thin  pink  callus;  peristome  pink,  narrowly  reflexed; 


HELIX-PAPUINA.  71 

columella  broadly  dilated,  blending  with  the  pink  callus  joining  the 
margins,  and  including  the  umbilicus.     (  Cox.) 
Alt.  0'72,  greater  diam.  1'09,  lesser  0'84  inch. 

Solomon  Is. 

H.  redempta  Cox,  P.  Z.  S.  1873,  p.  566,  t.  48,  f.  6,  6a.— PFR., 
Monogr.  vii. 

Differs  from  H.  eros  in  being  larger,  and  imperforate. 

H.  NIGROFASCIATA  Pfeiffer.     PI.  11,  figs.  8,  9. 

Narrowly  umbilicated,  globose  trochiform,  rather  solid,  obliquely 
regularly  striate,  closely  decussated  with  impressed  forward-descend- 
ing lines ;  buff  with  two  wide  black  bands,  bordered  sometimes  with 
brown,  spire  turbinate,  apex  rather  acute  ;  whorls  4,  convex,  the 
last  inflated,  slightly  descending  in  front,  lightly  spirally  striated 
beneath ;  aperture  oblique,  rounded-lunar,  white  and  black-banded 
within ;  peristome  white,  margins  converging,  the  outer  margin  ex- 
panded and  a  little  reflexed,  the  columellar  widely  reflexed,  dilated 
vaultingly  over  the  umbilicus. 

Alt.  13,  greater  diam.  19,  lesser  17  mill.     (Dohrn.') 

Admiralty  Is.  (Pfr.) 

H.  nigrofasciata  PFR.,  P.  Z.  S.  1863,  p.  524;  Monogr.  Hel.  Viv. 
v,  p. -337.— DOHRN,  Conchyl.  Cab.,  p.  591,  t.  173,  f.  9,  10. 

The  locality  may  not  be  correct.  It  has  the  appearance  of  a 
Solomon  Island  shell. 

H.  DONNAISABELL^E  AllgaS.      PI.  5,  fig.  87. 

Shell  perforate,  somewhat  depressedly  conical,  moderately  solid, 
very  finely  malleated  by  two  sets  of  oblique  stride  crossing  each  other 
at  right  angles  above  the  periphery,  whilst  below  it,  the  malleations 
become  more  irregular  and  flowing;  pale  yellowish-brown  ornamented 
with  a  narrow  white  band  below  the  sutures  and  one  or  two  dark 
brown  bands  on  each  whorl,  the  upper  one  being  the  narrowest ; 
spire  conical,  apex  rather  obtuse;  whorls  5£,  rather  convex,  the  last 
not  descending,  angled  at  the  periphery,  a  little  convex  at  base, 
which  has  a  single  broader  band  of  dark  brown  just  below  the 
periphery  ;  aperture  diagonal,  truncately  ovate,  peristome  white,  the 
upper  margin  narrow,  expanded,  the  lower  margin  arcuate  and  re- 
flexed,  almost  covering  the  umbilicus. 

Diam.  maj.  12,  min.  10J,  alt.  10  lines.     (Angas.) 

Eddy  stone  Is.,  Solomon  Group. 


72  HELIX-PAPUINA. 

H.  donna-isabellce  AXGAS,  P.  Z.  S.  1869,  p.  47,  t.  2,  f.  7.—  PFR., 
Monogr.  vii.  p.  380. 

(11)    GROUP  OF  H.  MOSELEYI. 

Iniperforate  ;  the  spire  low,  obtuse  ;  whorls  few  (about  4),  rapidly 
widening,  the  last  descending  in  front  ;  surface  obliquely  corrugated  ; 
lip  well-expanded. 

Allied  to  the  group  of  H.  motacilla,  but  imperforate  and  having 

fewer  whorls. 

H.  MOSELEYI  Smith.     PI.  6,  figs.  17,  18. 

Imperforate,  depressed,  the  spire  low,  apex  very  obtuse  ;  whorls 
3  -i  -4,  the  earlier  whorls  very  wide,  the  last  whorl  descending  in 
front,  rounded  at  the  periphery.  Surface  closely,  finely  sculptured 
with  forward-descending  wrinkles,  and  lightly  obliquely  striated. 
Earlier  1*  whorls  glossy,  corneous,  the  remainder  of  the  shell  opaque 
whitish,  except  a  corneous,  translucent  tract  around  the  umbilical 
region  ;  banded  with  from  one  to  five  translucent-brown  bands,  or 
bandless. 

Aperture  rounded-oval,  oblique;  peristome  white,  well  expanded 
all  around,  the  face  of  the  columella  flat. 

Alt.  14,  greater  diam.  21,  lesser  17*  mill.  (Specimen.) 
Alt,  11?,  greater  diam.  18,  lesser  15  mill.  (Specimen.) 
Alt.  15,  greater  diam.  23,  lesser  19  mill.  (Smith.) 

Wild  Island,  Admiralty  Group. 

H.  moseleyi  E.  A.  SMITH,  Proc.  Zool.  Soc.  Lond.  1884,  p.  263,  t. 
J,  f.  2,  2a. 


The  whorls  are  fewer  and  wider  than  in  any  other  Papuina. 
This,  in  combination  with  the  obliquely  corrugated  surface,  imper- 
forate axis  and  depressed  form,  render  it  easy  to  recognize. 

It  has  been  suggested  that  H.  moseleyi  belongs  to  Cristigibba,  but 
there  is  nothing  about  the  shell  to  warrant  such  reference. 

H.   NoV.l.CKORGIENSIS  Cox. 

This  species  was  included  in  Planispira  in  Vol.  VI  of  the  MANUAL, 
p.  L>(.M),  although  I  there  expressed  the  opinion  that  it  might  be  a 
I'<i/,'/i/i<i  (Geotrochns).  I  have  now  no  doubt  that  it  groups  here, 
th(.-  nearest  ally  known  of  the  preceding  species. 


HELIX-PAPUINA.  73 

(12)    GROUP  OF  H.  FRINGILLA. 

Iraperforate,  globose-trochoidal,  solid,  the  spire  rounded ;  surface 
striatulate  ;  lip  reflexed,  columella  bearing  an  obtuse  fold,  as  in  the 
group  of  H.  lienardiana. 

H.  FRINGILLA  Pfeiffer.     PL  16,  fig.  8  ;  vol.  VI,  pi.  49,  figs.  20,  21, 
22,  23,  24,  25. 

Imperforate,  solid,  depressed-globose,  the  spire  dome-shaped  or 
convexly  low-conical;  apex  obtuse;  whorls  4J,  slightly  convex,  the 
last  whorl  subcarinated  at  the  periphery,  abruptly  deflexed  in  front, 
strongly  constricted  behind  the  peristome,  which  is  narrowly  reflexed, 
white  or  bright  pink;  surface  smooth,  shining,  lightly  striatulate. 
Color  olive-yellow,  either  unicolored  or  having  brown  spiral  bands ; 
inner  whorls  white  or  pink.  Aperture  oblique,  subquadrate,  white 
within ;  columellar  margin  straight,  sloping,  having  a  more  or  less 
distinct  fold-like  callus,  or  blunt  tooth,  on  its  inner  margin,  near  the 
insertion. 

Alt.  18,  greater  diam.  24,  lesser  21  £  mill,  (large  specimen.) 
Alt.  16,  greater  diam.  21,  lesser  19  mill,  (average.) 

New  Georgia,  Solomon  Group ;  Admiralty  Is. 

H.  fringilla  PFR.,  P.  Z.  S.  1855,  p.  113;  Mon.  Hel.  Viv.  iv,  p. 
202.— DOHRN,  Conchyl.  Cab.,  p.  575,  t.  169,  f.  13-18.— Helicostyla 
fringilla  PFR.,  Mai.  Bl.  1855,  p.  145. — Merope  fringilla  ALB.,  Die 
Hel.  ii,  p.  158. — BRAZIER,  Journ.  de  Conchyl.  1880,  p.  319. —  Obba 
fringilla  FRAUENFELD. — Serpentulus  fringilla  TAPPARONE  CANE- 
FRI,  Viag.  Magent.,  p.  95. 

Differs  from  other  species  of  Papuina  in  the  con  vex  outlines  of  the 
spire,  the  constriction  behind  the  lip,  and  the  subdentate  columella. 
It  varies  much  in  color-pattern.  Some  specimens  are  pure  white, 
the  lip  either  white  or  rose-color. 

H.  BARNACLEI  Smith.      Unfigured. 

Shell  imperforate  in  the  adult  state,  (narrowly  umbilicated  when 
immature  and  acutely  angled  at  the  periphery)  conoidly  globose, 
with  a  very  shining  surface,  of  a  rich  fulvous  or  deep  fawn  color,  the 
upper  whorls  become  gradually  paler  than  the  last,  and  toward  the 
apex  are  semitransparent  horn  color  and  encircled  by  a  single  nar- 
row dark  brown  line  just  above  the  suture,  which  is  white;  this  line 
becomes  much  broader  upon  the  body  whorl,  which  has  a  second 
similar  band  beneath  the  more  or  less  white  periphery ;  this  is 


74  HELIX-PAPUIXA. 

obsoletely  keeled  or  angulated  near  the  upper  end  of  the  lip,  the 
keel  vanishing  altogether  on  the  last  half  of  the  whorl ;  whorls  4*,  a 
little  convex,  having  no  other  sculpture  than  the  arcuate  lines  of 
growth  (except  in  young  shells,  where  they  are  obsoletely  spirally 
striated),  the  last  somewhat  ventricose,  very  shortly  deflexed  at  the 
aperture ;  beneath  paler  or  horn  color  and  a  little  concave  in  the 
middle  or  umbilical  region  ;  the  aperture  is  very  oblique,  very  irreg- 
ularly triangular  lunate,  white  within  banded  with  black  and  brown 
(the  'latter  not  visible  exteriorly);  lip  not  much  thickened  on  the 
outer  and  basal  margins,  narrowly  expanded  and  reflected,  whitish, 
and  margined  on  the  outside  with  a  chestnut  band ;  the  columellar 
margin  white,  almost  straight,  thicker  than  elsewhere.  (Smith.) 
Alt.  15,  greater  diam.  24,  lesser  21  mill. 

Hawaii,  Sandwich  Islands. 

H.  (Merope  ?)  barnadei  E.  A.  SMITH,  Ann.  Mag.  K  H.  (4th  Ser.,) 
xx,  p.  242,  1877. 

This  very  pretty  species  closely  resembles  H.fringilla  of  Pfeiffer, 
which  inhabits  the  Admiralty  Islands.  Indeed  it  is  so  nearly  related 
to  it  that  were  I  not  certain  of  the  correctness  of  the  locality  whence 
it  was  obtained,  I  should  have  hesitated  to  describe  it  as  new. 

It  is,  however,  a  thinner,  lighter,  and  smoother  shell,  the  spire  is 
a  trifle  higher  and  less  obtuse,  the  whorls  a  little  more  convex,  more 
regular,  the  penultimate  proportionally  smaller,  and  the  last  larger 
and  not  showing  that  decided  constriction  behind  the  lip  which  is  so 
characteristic  of  Pfeiffer's  species.  The  lip,  too,  is  thinner  and  with- 
out the  least  trace  of  a  tooth-like  projection  on  the  columellar  edge. 
Specimens  offringilla  in  fine  condition  exhibit  distinct  spiral  stride 
on  the  entire  surface ;  such  sculpture  is  only  just  traceable  in  young 
examples  of  the  present  species. 

One  of  these  young  shells  has  a  particularly  beautiful  appearance, 
owing  to  the  broad  white  band  at  the  periphery,  which  is  margined 
on  both  sides  by  a  dark  brown  stripe  contrasting  prettily  with  the 
ground  color,  which  is  bright  fulvous  or  rich  faun. 

Three  specimens,  one  adult  and  two  immature,  have  been  presented 
to  the  British  Museum  bv  Mr.  H.  A.  Glanville  Barnacle,  whose 

».  * 

name  I  feel  much  pleasure  in  associating  with  this  species,  since  to 
him  is  owing  its  discovery. 

This  form  of  Helix,  being  so  different  from  any  other  hitherto 
found  in  the  Sandwich  Islands  led  me  to  think  that  there  must  be 
some  mistiikc  with  regard  to  the  alleged  habitat.  I  therefore,  wrote 


HELIX-PAPUINA.  75 

to  Mr.  Barnacle  upon  the  subject,  whose  reply  runs  as  follows: — "I 
am  perfectly  certain  that  the  specimens  I  have  were  grown  on  the 
Sandwich  Islands,  and  some  of  them  had  the  snails  in  them.  Whilst 
stationed  at  Kailua  I  used  at  times  to  go  shooting  at  some  brackish 
ponds  about  eight  miles  away  ;  and  whilst  going  quietly  amongst  the 
broad  reeds  to  shoot  a  duck  I  saw  one  of  the  shells  about  half  way 
up  a  reed.  I  took  it  and  found  two  or  three  more  that  day,  and 
now  and  then  others,  but  not  many.  Few  Europeans  go  to  these 
ponds,  as  it  is  impossible  to  find  them  without  a  guide  over  the  lava 
for  there  is  no  road.  Some  I  found  with  the  snail,  others  without ; 
those  with  the  animal  I  put  into  a  cigar  box  and  filled  it  up  with 
sea  sand  ;  and  so  the  animal  died  out,  and  then  I  washed  them." 

This  circumstantial  account  removes  all  doubt  respecting  the  true 
home  of  this  shell.  Notwithstanding  its  close  affinity  to  H.  fringilla, 
the  remoteness  of  the  habitats  of  the  two  species  and  the  extreme 
isolation  of  the  Sandwich  Islands  are  sufficient  reasons  in  themselves 
to  determine  the  specific  distinctness  of  these  shells.  (Smith.) 

(13)    GROUP  OF  H.  HELICINOIDES. 

Imperforate,  trochiform  species,  with  rhombic  aperture,  the  lip 
thickened  within,  not  expanded,  or  only  slightly  ;  the  columellar  lip 
not  expanded  or  reflexed. 

These  characters  define  one  of  the  most  distinct  groups  of  Papuina, 
— a  group  far  more  deserving  of  a  sectional  name  than  H.fringiUa, 
H.  antrorsa  or  other  of  the  species  representing  so-called  "  sections." 

I  have  examined  many  specimens  of  all  the  species  except  H. 
zelina,  H.  eyrene  and  H.  quirosi. 

The  species  inhabit  the  Admiralty  Is.,  Solomons  and  New 
Hebrides. 

In  all  other  groups  of  Papuina  the  columellar  lip  is  reflexed. 

H.  LABILLARDIEREI  Smith.     PL  15,  figs.  91,  92,  93. 

Imperforate,  trochiform,  spire  conical,  the  periphery  of  the  last  whorl 
acutely  keeled  in  front,  keel  more  or  less  obsolete  on  the  latter 
portion.  Surface  lusterless,  having  rather  coarse  and  irregular 
oblique  striae.  Color  whitish  unicolored,  or  with  a  broad  suffused 
brownish  band  below  the  middle  of  the  body-whorl,  or  entirely  light 
pinkish-brown  ;  the  apex  black.  Whorls  5 ;  base  very  convex,  deeply 
impressed  in  the  center. 


HELIX-PAPUINA. 

A  pi-mire  oblique;  lip  white,  very  broadly  and  flatly  expanded, 
thickened  within,  abruptly  narrowing  toward  the  columellar  termina- 
tion. Alt.  11,  greater  diani.  15  mill. 

Wild  Island,  Admiralty  Group. 

H.  labillirdierei  E.  A.  SMITH,  P.  Z.  S.  1884,  p.  264,  t.  22,  f.  3,  3a. 

Allied  to  H.  helicinoides,  but  readily  distinguished  by  the  broad 
lip,  constantly  black  apex,  and  more  globose  body-whorl,  upon 
which  the  keel  is  more  or  less  obsolete  on  the  latter  portion.  The 
convexity  of  the  base  around  the  umbilicus,  reminds  one  of  H. 
helicinoides. 

H.  HELICIXOIDES  Hombron  &  Jacquinot.     PI.  15,  figs.  72,  73,  74. 

Imperforate,  trochiform,  the  spire  conical,  periphery  acutely 
carinated.  Surface  lightly  striatulate,  and  densely,  most  minutely 
spirally  striated.  Pale  horn-colored,  somewhat  translucent ;  aperture 
subrhomboidal ;  peristome  thickened  within,  slightly  expanded,  its 
face  flattened;  slightly  angled  at  the  position  of  the  carina; 
columellar  margin  narrow,  having  an  obtuse  callous  fold  above, 
near  the  insertion. 

There  are  5?  whorls,  those  of  the  spire  nearly  rectilinear,  the  last 
whorl  slightly  convex  above  and  below.  The  base  is  deeply  impressed 
at  the  axis,  having  a  little  pit  at  the  place  of  the  umbilicus  ;  around 
the  depression  it  is  very  convex,  rendering  the  baso-columellar  lip 
strongly  arcuate.  Alt.  12,  greater  diam.  15,  lesser  14  mill. 

Solomon  Islands. 

H.  helicinoides  H.  &  J.  (PFR.,  Zeitschr.  f.  Mai.  1849,  p.  77)  Voy. 
Pol  Sud  Atlas,  t.  7,  f.  34-37.— PFR.,  Conchyl.  Cab.,  p.  294,  t.  125, 
f.  14,  1 5  ;  Monogr.  iii,  p.  104. — H.  cleryi  RECLUZ,  Journ.  de  Conchyl. 
1851,  p.  211,  t.  5,f.  10.— PFR.,  Monogr.  iii,  p.  179.— E.  A.  SMITH, 
I1.  Z.  S.  1885,  p.  592,  t.  36,  f.  6,  (with  varieties.) 

The  prominent  characters  of  typical  helicinoides  are  the  distinctly 
nurrowed,  sinuous  columella,  strongly  arcuate  baso-columellar  lip, 
and  flattened  peristome.  The  typical  helicinoides  is  of  a  uniform 
li;_rht  color.  Specimens  with  these  characters  strongly  developed 
form  Mr.  Smith's  var.  simboana. 

Vur.  CLERYI  Recluz.     PL  15,  figs.  81,  82,  83,  86-87. 

Rather  thinner  than  typical  helicinoides,  the  lip  generally  nar- 
rower, less  expanded,  columella  less  calloused,  basal  lip  less  arcuate  ; 


HELIX-PAPUINA.  77 

horny  or  whitish  with  a  broad  brown  band  above,  and  a  narrower 
one  just  below  the  periphery,  which  is  very  acute  and  white. 

Alt.  10,  greater  diam.  16,  lesser  14J  mill. 

Choiseul  Bay,  San  Christoval,  Shortland  and  Treasury  1$.,  Solomon 
Group. 

Typically  quite  distinct  from  helicinoides,  but  intermediate  forms 
occur.  Smith's  var.  septentrionalis  belongs  here. 

The  basal  band  is  wanting  in  one  suite  before  me  (pi.  15,  figs.  81- 
83.) 

The  form  described  as  var.  meridionalis  Smith  (pi.  16,  fig.  5)  is 
smaller,  pale  brown  above,  with  a  white,  thread-like  line  at  the 
suture  and  the  acutely  keeled  periphery,  paler  beneath,  especially  to- 
ward the  center,  and  have  the  aperture  particularly  acuminate  at 
the  termination  of  the  keel.  It  is  from  Santa  Anna  Island,  living 
on  young  cocoa-nut  palms,  growing  on  low  ground  bordering  the 
sea. 

H.  CINERACEA  Hombron  &  Jacquinot.     PI.  15,  figs.  88,  89 ;  pi.  16, 
figs.  6,  7. 

Imperforate,  globose-trochiform,  the  spire  conical,  periphery  of  the 
last  whorl  acutely  keeled.  Surface  striatulate,  most  minutely  and 
closely  spirally  striated.  Pale  brownish  horn-colored,  somewhat 
translucent,  paler  on  the  central  part  of  the  base.  Aperture  rounded- 
rhomboidal ;  peristome  white,  narrow,  very  little  expanded,  thick- 
ened within,  columellar  margin  straightened,  sloping,  slightly 
sinuous. 

There  are  5,  slightly  convex  whorls ;  the  apex  is  corneous-brown  ; 
the  last  whorl  is  convex  above,  very  convex  beneath,  far  more  so 
than  H.  helicinoides.  The  color  is  often  reddish-brown  with  a  light 
band  at  the  suture,  and  the  central  tract  of  the  base  light ;  the 
periphery  has  a  white  thread  also. 

Alt.  10,  greater  diam.  12  J,  lesser  11  mill. 

San  Christoval,  Ysabel  and  Stephens  Is.,  Solomons. 

H.  cineracea  HOMBR.  &  JACQ.,  Voy.  Pol  Sud  Atl.,  t.  7,  f.  30-33. 
— PFR.,  Monogr.  Hel.  Viv.  iv,  p.  203. — H.  cinerarea  Rouss.  in  text 
of  Voy.  Pol  Sud  p.  28. 

This  is  a  decidedly  more  globose  shell  than  H.  helicinoides.  The 
base  is  more  convex,  and  the  columella  less  sinuated  above. 


78  HELIX-PAPUINA. 

H.  ZELIXA  Cox.     PL  15,  fig.  90. 

Shell  imperforate,  pyramidally-conical,  transparent,  thin,  obliquely 
and  very  coarsely  striated,  straw  colored ;  whorls  7,  almost  flat,  the 
last  having  a  very  sharp  produced  keel ;  suture  broadly  margined 
above;  base  flat;  aperture  trapeziform ;  outer  margin  a  little  re- 
flected; peristome  thickened  and  white,  particularly  the  lower 
margin,  which  is  a  little  everted  and  reflexed,  columellar  margin, 

O         " 

rather  contracted  and  nodose.     (Cox.) 

Alt.  '60,  greater  diam.  '62,  lesser  '55  inch. 

Solomon  Is. 

H.  zelina  Cox,  P.  Z.  S.  1873,  p.  150,  t.  16,  f.  6. 

This  species  is  probably  more  closely  allied  to  H.  cineracea  H.  & 
J.  and  H.  subtecta  Pfr.  than  to  any  other.  It  is  a  much  more 
conical  shell  than  these,  and  its  sculpture  decidedly  coarser ;  the 
flattened-out  keel  of  the  periphery  is  also  characteristic  of  this 
species.  ( Cox.) 

H.  CYRENA  Crosse.     PI.  15,  figs.  79,  80. 

Shell  with  an  almost  closed  perforation,  subtrochiform,  quite  thin, 
rather  pellucid,  quite  shining,  very  finely  and  subobliquely  striat- 
ulate ;  opaque  milk-white,  with  a  rather  broad  translucent  zone 
above.  Spire  subconic,  rather  acute,  the  apex  rosy-violet ;  suture 
impressed;  whorls  5,  a  little  convex,  the  last  whorl  subobtusely  cari- 
nated,  not  descending,  somewhat  exceeding  the  spire  in  length ;  base 
convex,  subinflated,  more  shining,  whitish  ;  aperture  very  oblique, 
widely  lunate-rhombic;  peristome  milk-white,  margins  subparallel, 
the  columellar  margin  somewhat  thickened,  short ;  basal  margin  ex- 
panded ;  outer  margin  a  little  reflexed,  subangulate  in  the  middle. 

Alt.  11,  greater  diam.  17,  lesser  13?  mill.     (Crosse.) 

Habitat  unknown. 

H.  cyrene  CR.,  Journ.  de  Conchyl.  1869,  p.  183 ;  1870,  p.  102,  t. 
2  f  2 

—  ,    1  •  +j  • 

Allied  to  H.  eva  but  differing  in  the  slight  umbilical  perforation, 
obtuse  carina,  reflected  outer  border  of  the  peristome,  and  the  colora- 
tion. //.  helicinoides  differs  in  the  more  acute  keel  and  the  different 
suture  and  coloration.  (Cr.) 

H.  EVA  Pfeiffer.     PI.  15,  figs.  84,  85,  76,  77,  78;  pi.  16,  figs.  1,  2, 
3,4. 

Imperforate,  trochiform,  solid,  opaque;  the  spire  straightly  conic, 
periphery  acutely  carinated ;  surface  shining,  obliquely  striatulate, 


HELIX-PAPUINA.  79 

sometimes  showing  excessively  fine  spiral  lines  in  places  on  the  base. 
Opaque,  white  or  bluish,  creamy  or  pinkish  tinted,  unicolored  or 
having  a  brown  band  above,  often  showing  a  few  scattered  translu- 
cent dots.  Whorls  5J,  slightly  convex. 

Aperture  oblique,  subrhombic,  the  peristome  white  or  brown, 
somewhat  thickened  and  obtuse  but  not  expanded,  the  superior 
margin  straight,  basal  margin  forming  a  regular  curve  from  outer 
angle  to  insertion,  a  little  thickened  within  and  straightened  at  the 
columellar  slope. 

Alt.  9J,  greater  diam.  15,  lesser  13  mill. 

Alt.  9J,  greater  diam.  12*,  lesser  11  mill. 

New  Hebrides  (Island  of  Vate.) 

H.  em  PFR.,  P.  Z.  S.  1852,  p.  84 ;  Monogr.  iii,  p.  180 ;  Conchyl. 
Cab.,  p.  190,  t.  160,  f.  22,  23.— REEVE,  Conch.  Icon.,  f.  977.— 
BRAZIER,  Journ.  de  Conchyl.  1880,  p.  313. 

This  species  has  the  contour  of  H.  helicinoides  except  that  the  lip 
is  not  expanded,  the  columella  not  contorted,  the  base  flatter,  and 
consequently  the  basal  lip  is  much  less  strongly  arcuate.  It  further 
differs  in  being  more  solid  and  opaque.  These  same  characters 
separate  it  from  cleryi.  It  is  far  flatter  below  and  more  solid  and 
opaque  than  H.  cineracea. 

A  form  with  obtuse  periphery  also  occurs.     See  pi.  16,  figs.  1. 

H.  LAYARDI  Hartman.     PI.  15,  figs.  94,  95,  96,  97. 

Imperforate,  solid,  opaque,  trochiform,  the  spire  straightly  conical, 
periphery  acutely  carinated,  base  flattened.  The  surface  is  shining, 
lightly  striatulate.  Color  white,  more  or  less  suffused  or  streaked 
with  pink  in  places,  having  very  numerous  purplish  translucent  dots 
and  zigzag  marks  above,  fewer  below  ;  often  banded. 

There  are  6^  flat  whorls,  the  carina  of  the  last  acute,  having  a 
delicate  line  of  white  at  the  edge,  bordered  above  and  below  with 
dark  purplish-brown.  When  bands  are  present  there  is  a  faint  one 
in  the  middle  of  the  upper  surface,  and  a  broad  or  narrow  one  on 
the  base.  They  are  visible  within  the  aperture.  The  aperture  is 
rhomboidal,  very  oblique,  the  lip  not  expanded,  blunt,  white; 
columella  oblique,  straight,  often  brown. 

Alt.  14?,  greater  diam.  17,  lesser  14*  mill. 

Alt.  14J,  greater  diam.  15,  lesser  13  mill. 

Aura  Island,  New  Hebrides. 


Ml  HELIX-PAPUINA. 

Oxychowi  Injardi  HARTMAN,  Proc.  Acad.  Nat.  Sci.  of  Phila.  1889, 
p.  91,  t.  v,  f.  3. 

Allied  to  H.  eva  in  form  of  the  lip,  and  the  solid  opaque  texture. 
Differs    in    the    greater   elevation    of   the    spire,  larger    size    and 
coloration.     One   specimen  before  me   lacks  the   peripheral   dark 
girdle. 
H.  QUIROSI  Cox.     Unfigured. 

Shell  imperforate,  conoidly  depressed,  thin,  translucent,  rather 
coarsely  striated,  of  a  pale  brown  throughout;  spire  broadly  and 
flatly  conoid,  acute  at  the  apex ;  whorls  5i  to  6,  convexly  flattened, 
the  last  one  considerably  inflated,  and  sharply  keeled  at  the  periph- 
ery ;  base  rounded ;  aperture  sharply  lunar,  angled  at  the  periphery 
of  the  last  whorl  and  at  the  junction  of  the  lower  and  columellar 
margins ;  lip  slightly  thickened  and  expanded ;  columellar  margin 
straight,  thickened,  not  dilated  or  reflected. 

Diam.  greatest  0*80,  least  0'64  ;  height  0'40  of  an  inch.     (  Cox.) 

Solomon  Islands. 

H.  (Geotrochus)  quirosi  Cox,  P.  Z.  S.  1873,  p.  147. — PFR.,  Mon. 
vii,  p.  316. 

This  species  would  be  like  a  magnified  specimen  of  H.  cleryi 
Reel,  were  it  not  wholly  unornamented,  much  more  broadly  ex- 
panded and  less  conical.  (  Cox.) 

H.  PYXIS  Hinds.     PL  13,  figs.  42,  43. 

Small,  imperforate,  globose-turbinate,  the  periphery  acutely 
keeled  ;  white,  slightly  translucent.  Last  whorl  descending  in  front, 
swollen  and  then  contracted  behind  the  lip ;  columella  narrower  than 
basal  lip,  not  expanded  or  reflexed  except  immediately  at  the  inser- 
tion. 

The  surface  is  slightly  shining,  irregularly  obliquely  striate,  and 
has  irregular  spiral  impressed  lines  on  the  base.  The  spire  is 
conoidal,  apex  obtuse.  Whorls  4J,  convex,  the  last  swollen,  acutely 
keeled,  slowly  descending  in  front;  base  swollen,  then  contracted 
just  behind  the  lip.  Aperture  small,  very  oblique,  rounded,  not 
angular  at  the  position  of  the  keel,  which  becomes  evanescent  on  the 
latter  part  of  the  body-whorl.  Peristome  white,  right  margin 
slightly  expanded,  basal  margin  thickened  within;  columellar 
margin  narrow,  not  reflexed. 

Alt.  6,  greater  diarn.  7,  lesser  6}  mill. 

New  Ireland. 


HELIX.  81 

H.  pyxis  HINDS,  Voy.  Sulphur,  Moll,  in,  p.  55,  t.  19,  f.  10. — PFR., 
Monogr.  i,  p.  219  ;  iii,  p.  180  ;  Zeitschr.  f.  Mai.  1849,  p.  76  ;  Conchyl. 
Cab.,  p.  293,  t.  125,  f.  12,  13.— REEVE,  f.  793. 

This  is  a  very  pretty  little  shell,  the  smallest  of  the  H.  helicinoides 
group.  It  further  differs  from  the  others  in  its  white  color,  globose 
form,  convex  whorls,  and  rounded  mouth.  The  characters  of  the 
eolumella,  however,  show  unmistakably  that  pyxis  is  correctly 
grouped  here. 

H.  CRUCIBULUM  Pfeiffer.      Unfigured. 

Imperforate,  trochiform,  rather  solid,  covered  with  a  greenish- 
tawny  epidermis,  partly  removed ;  spire  regularly  conic,  rather 
acute;  suture  margined;  whorls  6*,  the  upper  a  little  convex, 
irregularly  striate,  the  last  smooth,  not  descending,  provided  with 
an  acute,  delicate  carina ;  base  more  convex,  shining,  impressed  in 
the  middle ;  aperture  oblique,  ax-shaped ;  peristome  straight,  mar- 
gins distant,  the  upper  acute,  labiate  with  white  within  behind  the 
edge,  basal  margin  white  callously  thickened. 

Alt.  10,  greater  diam.  13i,  lesser  12*  mill.     (P/r.) 

A  dmira  Ity  Is  lands. 

H.  crucibulum  PFR.,  Mai.  Bl.  iii,  p.  241 ;  Mon.  Hel.  Viv.  iv,  p. 
192. 

I  do  not  know  where  to  group  this  unfigured  species. 

*  ,  * 
* 

II.  Species  doubtfully  referred  to  Papuina,  inhabiting  southeastern 
Asia,  Andaman  Is.,  Java  and  Sumatra. 

The  species  included  here  agree  with  Papuina  in  general  form, 
but  differ  in  texture  and  usually  also  in  surface-sculpture  and  color- 
ing. In  shell  characters  they  approach  a  number  of  quite  diverse 
groups,  so  that  until  the  anatomy  is  investigated  we  will  remain 
ignorant  of  the  true  classification  of  some  of  them.  Subgeneric 
names  have  been  proposed  for  several  of  the  species. 

Under  the  circumstances  it  seems  best  to  group  them  in  this  place 
temporarily.  I  have  indicated,  however,  under  each  group  my  im- 
pressions as  to  its  affinities  and  correct  systematic  position. 

(1)    GROUP  OF  H.  PERAKENSIS. 

Thin,  straightly  conical,  carinated  species,  having  decided  affinities 

to  the  section  Satsuma,  a  portion  of  them  certainly  belonging  there. 
6 


82  HELIX. 

( '.nnpari.-  .>n-  should  also  be  made  with  TrocJiomorphoides  of  Nevill, 
type  H.  <icris  Bens.,  and  Ganesella  Blanf.,  type  H.  capitium  Bens. 
These  groups  having  been  monographed  by  Tryoii  in  Vol.  Ill  of  the 
MANUAL,  1887,  no  course  is  open  to  me  but  to  include  the  species 
here. 

Bulimus  sphceroconus  Pfr.,  and  some  other  species  should  have 
been  placed  in  Satsuma,  and  there  are  also  a  number  of  Philippine 
Island  species  likewise  belonging  to  that  characteristically  east 
Asiatic  section. 

H.  PERAKENSIS  Crosse.     PL  18,  figs.  44,  45. 

Shell  narrowly  umbilicated,  turrited-conic,  trochiform  ;  slightly 
subobliquely  striatulate ;  rather  thin,  subtranslucent,  whitish,  having 
a  narrow  chestnut  zone  immediately  above  the  suture ;  spire  rather 
elongated,  the  apex  rotund,  sutures  impressed.  Whorls  6,  nearly 
flat,  the  embryonic  earlier  2  smooth,  corneous ;  the  last  whorl  shorter 
than  the  spire  (last  whorl  and  spire  having  the  ratio  of  4  :  7)  ;  last 
whorl  acutely  carinated  at  the  periphery,  the  carina  brown,  the  base 
nearly  flat.  Aperture  subquadrate  ovate,  white  inside,  showing  by 
transparence  the  brown  line  at  the  carina.  Peristome  simple, 
shortly  reflexed,  white,  margins  separated,  the  columellar  margin 
short,  straight,  outwardly  dilated  and  covering  a  third  of  the  um- 
bilicus ;  basal  margin  subarcuate  ;  outer  lip  subangular  at  the  carina, 
narrowing  above  toward  the  insertion. 

Alt.  11,  greater  diarn.  10,  lesser  9  mill.     (Crse.) 

Perak. 

H.  (Getrochus')  perakensis  CR.,  Journ.  de  Conchyl.  1879,  p.  199, 
t.  8,  f.  4. 

Yar.  SUBPERAKENSIS  Pilsbry.     PI.  18,  figs.  46,  47. 

Shell  umbilicated,  turrited-conic,  thin.  Whorls  7-71,  the  earlier 
convex,  forming  an  obtuse  apex ;  last  whorl  carinated,  convex 
bem-ath,  slightly  descending  in  front.  Surface  finely  obliquely 
striatulate.  Aperture  oblique,  outer  lip  narrowly  expanded, 
colmiH'llar  lip  broadly  dilated. 

Alt.  1 1,  greater  diam.  10-],  lesser  9  mill.;  oblique  alt.  of  aperture 
5S,  width  7  mill. 

Tonquin. 

Differs  from  perakensis  in  the  blunter  carina,  more  convex  base, 
and  irivater  number  of  whorls.  The  specimens  being  dead  I  am  not 


HELIX.  83 

certain  of  the  coloring,  but  no  trace  of  a  brown  band  is  visible.      It 
is  probably  a  distinct  species. 

H.  PHONICA  Mabille.     Vol.  VI,  pi.  15,  figs.  74,  75. 

Nearly  covered  perforate,  trochiform,  thin  but  solid,  corneous, 
subpellucid,  shining,  subarcuately  and  irregularly  striated;  spire 
high-pyramidal,  the  apex  rather  obtuse,  concolored.  Whorls  8,  a 
little  convex,  regularly  increasing  separated  by  impressed  sutures  ; 
last  whorl  larger,  acutely  carinated,  sloping  above,  a  little  deflexed 
and  dilated  at  the  aperture.  Aperture  transversely  ovate,  lunate, 
angled  at  the  outer  lower  margin ;  peristome  scarcely  thickened, 
acute,  a  little  reflexed ;  outer  margin  a  little  curved,  slightly  in- 
flexed  ;  basal  margin  curved,  shortly  reflexed ;  columellar  margin 
dilated  in  a  white  triangular  plate,  nearly  covering  the  umbilicus. 

Alt.  12,  greater  diam.  11,  lesser  10  mill.     (Mabille?) 

Tonquin. 

H.  phonica  J.  MABILLE,  Moll.  Tonk.,  diagn.,  p.  3,  May  14, 1887  ; 
Bull.  Soc.  Mai.  Fr.  iv,  p.  94,  t.  2,  f.  8,  9,  1887. 

Evidently  allied  to  H.  perakensis,  but  larger,  more  acutely  keeled, 
and  with  narrower  umbilicus. 

H.  ROSTRELLA  Pfeiffer.     PL  18,  figs.  48,  49,  50. 

Shell  umbilicate,  globose-trochiform,  rather  thin,  irregularly  striat- 
ulate,  silky,  tawny-corneous.  Spire  conoid,  apex  rather  acute ; 
whorls  5J,  a  little  convex,  the  last  slightly  descending  in  front,  car- 
inated above  the  middle,  the  base  more  convex.  Umbilicus  nar- 
row. Aperture  oblique,  submargaritaceous  inside ;  peristome  white 
or  flesh  colored  the  margins  converging,  right  margin  expanded, 
subsinuate  above  the  carina,  then  shortly  beaked,  the  basal  margin  a 
little  reflexed  and  at  the  insertion  subdilated. 

Alt.  10,  greater  diam.  17,  lesser  15  mill.     (Pfr.) 

Laos  Mts.,  Cambodia. 

H.  rostrella  PFR.,  P.  Z.  S.  1862.  p.  270  ;  Monogr.  v,  p.  331 ;  Novit. 
Conch,  iii,  p.  379,  t.  88,  f.  1-3. 

Var.  Corneous-white,  with  a  chestnut  band  at  the  carina. 
Probably  a  Satsuma. 

H.  SCENOMA  Benson.     PI.  18,  figs.  53,  54. 

Shell  with  a  narrow  funnel-shaped  umbilicus,  subglobose-conoid, 
obliquely,  irregularly  striated,  decussated  by  very  minute  spiral 
stride,  whitish  under  a  corneous  epidermis,  sometimes  having  a  red- 


84  HELIX. 

dish-chestnut  hand  at  the  carina.  Spire  conoid,  apex  rather  obtuse, 
suture  impressed';  whorls  6,  a  little  convex,  the  last  slightly  descend- 
ing in  front,  at  the  periphery  obtusely  compressedly  carinated,  con- 
vex beneath  compressed  around  the  umbilicus  ;  aperture  oblique, 
^ibquadrate-lunate;  peristome  a  little  expanded,  the  columellar 
margin  a  little  reflexed. 

Alt.  11,  greater  diam.  16,  lesser  14  mill.     (Bens.} 

Moulmein. 

H.  scenoma  BENS.,  Ann.  Mag.  N.  H.  3d.  Ser.  xi,  May,  1863,  p. 
319.— PFR.,  Monogr.  v,  p.  331,  502.— HANL.  &  THEOB.,  Couch. 
Ind.,  t.  53,  f.  5,  and  var.  f.  3,  4. 

Seems  to  be  a  Satsuma. 

H.  BAXTAMENSIS  Smith.     PI.  18,  fig.  51. 

Shell  elevated  conic,  obtuse  at  the  apex,  narrowly  perforated, 
somewhat  thin,  grayish-white,  scarcely  shining;  whorls  7,  slowly 
increasing,  the  three  upper  ones  convex,  the  remainder  nearly  flat, 
all  over  minutely  granulate,  sculptured  with  very  oblique  delicate 
growth-stride,  at  the  scarcely  oblique  sutures  margined  with  a  slender 
cariua ;  last  whorl  acutely  cariuated  at  the  periphery,  nearly  flat 
beneath,  slightly  descending  in  front.  Aperture  oblique,  small, 
equaling  3  the  entire  length ;  peristome  whitish,  upper  margin 
oblique,  thin,  sinuous,  slightly  reflexed.  the  basal  margin  broadly 
expanded,  half-covering  the  umbilicus  above. 

Alt.  12,  greater  diam.  11,  lesser  10  mill.     (jSmith^) 

Bantam,  Java. 

H.  (  Geotrochus)  bantamensis  E.  A.  SMITH,  Ann.  de  la  Soc.  Roy. 
Malac.  de  Belgique,  xxii  (4th  ser.,  ii),  p.  217,  t.  9,  f.  11,  1887. 

Remarkable  for  its  elevated  form,  granulated  surface,  and  the 
acute  peripheral  carina,  which  continuing  upward  gives  the  suture  a 
carinated  appearance. 

H.  RUFOFILOSA  Bock.     PL  3,  fig.  55. 

Shell  shortly  conical,  narrowly  umbilicated,  thin,  semitrausparent, 
of  a  greyish  or  horn-color,  sometimes  with  a  thread-like  red  line 
bordering  the  sutures  and  encircling  the  last  whorl  at  the  periphery. 
Spire  with  rectilinear  outlines,  converging  at  an  angle  of  about  30°, 
rather  obtuse  at  the  apex.  Whorls  7,  very  obliquely  striated,  slowly 
increasing,  keeled  beneath  immediately  above  the  suture;  three  or 
four  uppermost  rather  more  convex  than  those  beneath,  which  are 


HELIX.  85 

but  slightly  so  ;  last  volution  acutely  carinate  at  the  middle,  and 
convexly  flattened  beneath.  Aperture  small,  oblique.  Peristome 
thin,  outer  margin  above  the  keel  scarcely  expanded,  beneath  it 
feebly  thickened  and  narrowly  reflexed,  in  the  umbilical  region 
more  expanded  and  partly  concealing  the  perforation. 

Length  from   10  to  11  mill.,  greatest  diameter  of  base  also  10- 

11  mill.     (Bock.) 

Paio,  Sumatra;  in  the  forests,  1500  feet  above  the  sea. 

H.  (Geotrochus)  rufo-filosa  BOCK,  P.  Z.  S.  1881,  p.  630,  t.  55,  f.  4. 
H.  NIAHENSIS  Godwin- Austin.  PL  18,  fig.  66. 

Shell  dextral,  pyramidal,  thin  texture,  base  flat ;  sculpture,  spiral 
lirse,  fine,  regular,  not  of  continuous  thickness,  crossed  by  lines  of 
growth  ;  color  white,  with  some  fine  marbling  in  a  greyer  color, 
within  the  aperture  porcellaneous  white ;  spire  high,  pyramidal, 
sides  slightly  concave  ;  apex  acuminate  ;  suture  fine,  linear ;  whorls 
7,  very  flat ;  aperture  ovate,  wide  and  ample,  oblique ;  peristome 
sharp,  coming  to  an  angulate  form  at  the  outer  keeled  margin,  ex- 
panded and  thickened  below  towards  the  columellar  side. 

Size:  maj.  diara.  18*8,  min.  16*0;  alt.  axis  12*3  mill.     (G.  Aust.) 

Niah  Hills. 

H.  (Geotrochus~)  niahensis  AUST.,  P.  Z.  S.  1891,  p.  44,  t.  2,  f.  3. 
Described  from  a  single  specimen. 

H.  TIG^ENSIS  Godwin-Austin.     PI.  18,  fig.  65. 

Shell  pyramidal,  rounded  below,  narrowly  umbilicated ;  sculpture, 
irregular  transverse  lines  of  growth,  indistinct  concentric  striation  on 
base ;  color  very  pale  ochre  throughout ;  spire  pyramidal,  sides  flat ; 
apex  blunt ;  suture  linear ;  whorls  6,  very  flat ;  aperture  semi- 
lunate  ;  peristome  reflected  upon  the  columellar  margin,  which  is 
very  oblique. 

Size:  maj.  diani.  9-3  ;  alt.  axis  8*0  mill.     (G.  Aust.) 

Tiga  Island,  Borneo. 

H.  (Geotrochus)  tigcensis  AUST.,  P.  Z.  S.  1891,  p.  44,  t.  2,  f.  5. 
Described  from  a  single  specimen. 

H.  SUBFLAVA  Godwin-Austin.     PI.  18,  fig.  52. 

Shell  pyramidal,  keeled,  rather  flat  on  the  base,  narrowly  per- 
forate ;  sculpture,  rough  pappillate  dottings  arranged  transversely ; 
color  ochre ;  spire  high,  sides  flat ;  apex  very  blunt ;  suture  linear ; 


HELIX. 

whorls  6,  sides  flatly  convex  ;  aperture  quadrate ;  peristome  very 
thin  ;  columellar  margin  much  reflected,  perpendicular. 
Size  :  maj.  diam.  7'0  ;  alt.  axis  6'5  mill.     (G.  Aust.*) 

Borneo. 

H.  (Geotrochu*)  subflava  AUST.,  P.  Z.  S.  1891,  p."  45,  t.  2,  f.  4. 
Has  the  appearance  of  Trochomorphoides. 

(2)  GROUP  OF  H.  WRAYI.     (Philidora.) 

Philidora  DE  MORGAN,  Bull,  de  la  Soc.  Zoologique  de  France, 
1885,  p.  384.  First  species,  P.  wrayi. 

The  name  Philidora  was  proposed  by  de  Morgan  as  a  generic  term 
to  replace  the  preoccupied  name  Philina  of  Albers.  If  it  were  really 
the  same  as  Philina,  the  name  would  become  a  synonym  of  Obbina 
Semper  ;  but  it  is  not  the  same.  In  the  absence  of  specimens  I  am 
unable  to  say  whether  the  group  belongs  in  the  neighborhood  of  the 
Chinese  Fruticicoloid  Helices,  such  as  Satsuma,  or  near  the  section 
Trachia.  De  Morgan  has  given  no  diagnosis  or  characters  of  his 
group. 

H.  WRAYI  de  Morgan.     PI.  18,  figs.  55,  56,  57,  58. 

Shell  dextral,  flattened,  bluish-white,  narrowly  umbilicated,  striate, 
strongly  carinated,  composed  of  5  whorls,  ornamented  with  ribs 
parallel  to  growth-lines ;  upper  surface  of  the  whorls  flattened,  lower 
surface  rounded ;  suture  linear,  very  distinct.  Aperture  oblique, 
elliptical  irregular,  encroached  upon  by  the  preceding  whorl ;  per- 
istome reflexed. 

Alt.  6,  diam.  15,  length  of  aperture  8  mill.     (Morgan.} 

Lahat,  Ipoh,  Valley  of  the  Kinta  River,  Perak. 

Philidora  Wrayi  MORGAN,  Bull.  Soc.  Zool.  Fr.  1885,  p.  384,  t.  6, 
f.  5. 

H.  HARDOUINI  de  Morgan.     PI.  18,  figs.  60,  61,  62,  63. 

Shell  dextral,  flattened,  translucid,  corneous,  widely  umbilicated, 
permitting  the  whorls  of  the  spire  to  be  seen  within  it ;  finely  striated 
and  very  strongly  carinated,  carina  extending  to  the  aperture  ;  com- 
posed of  5-6  whorls,  which  are  flattened  above,  rounded  beneath ; 
suture  a  little  distinct.  Aperture  oblique,  elliptical,  irregular, 
slightly  encroached  upon  by  the  preceding  whorl ;  peristome  reflexed. 

Alt.  8,  diam.  17  mill.  This  shell  is  of  a  corneous  brown  color,  the 
aperture  lighter,  (de  Morgan.} 


HELIX.  87 

Between  LaJiat  and  Ipoh,  valley  of  the  Kinta  river,  abundant  in 
moist  places  in  the  forest. 

Philidora  Hardouini  DE  MORG.,  Bull.  Soc.  Zool.  Fr.  1885,  p.  385, 
t.  5,  f.  10. 

• 

(3)  GROUP  OF  H.  ARFAKIENSIS.     (  Coliolus.) 

Coliolus  TAPPARONE  CANEFRI,  Ann.  Mus.  Civ.  Genov.,  2d.  ser.,  iv, 
p.  131,  1887. 

Shell  elevated-conical,  many-whorled ;  whorls  obliquely  costulate 
and  bristly ;  apex  obtuse,  mamillar ;  base  depressed ;  terminations 
of  the  peristome  connected  by  a  callus.  (T.  C.) 

This  section,  founded  upon  a  most  singular  shell,  is  placed  by 
Tapparone  Canefri  in  the  vicinity  of  the  section  Trochomorphoides 
of  Nevill. 

H.  ARFAKIENSIS  Tapparone  Canefri.     PI.  18,  fig.  59. 

Shell  trochiform,  narrowly  umbilicated,  corneous ;  spire  much 
elevated,  somewhat  constricted  above,  the  apex  mamillar.  Whorls 
11,  the  earlier  4  convex,  spirally  striate,  the  following  whorls  plano- 
convex, obliquely  sharply  costulate,  se.tigerous  at  middle  and  base, 
the  bristles  long,  nearly  covering  the  suture ;  last  whorl  acutely 
carinated  at  the  periphery,  the  base  nearly  flat,  radiately,  finely 
striated.  Aperture  dilated,  rhombic-quadrangular,  very  oblique, 
the  peristome  sinuous  above,  subexpanded,  reflexed  beneath,  in- 
curved and  in  the  middle  subaugulated,  the  margins  distant,  joined 
by  a  thick  callus. 

Alt.  11 |,  greater  diam.  8f,  lesser  7£  mill.     (T.  Cy.) 

Hatam,  Mt.  Arfak,  New  Guinea. 

H.  arfakiemis  T.  C.,  1.  c.,  p.  131,  t.  1,  f.  21. 

(4)    GROUP  OF  H.  TROCHALIA. 

Umbilicate ;  depressed,  strongly  carinated  ;  the  last  whorl  deflexed 
in  front ;  aperture  elliptical,  peristome  reflexed ;  surface  simply 
striate. 

The  following  species  has  been  placed  in  Geotrochus  by  Pfeiffer, 
in  Ampelitaby  Tryon,  and  in  Planispira  by  Nevill.  It  seems  to  me 
to  be  closely  allied  to  H.  gabata  Gld.,  merguiensis  Phil.,  etc.,  species 
belonging  very  close  to  Trachia,  and  probably  to  be  included  there- 
in rather  than  in  Plectotropis. 


HELIX. 


H.  TROCIIAUA  Benson.     PL  18,  figs.  67,  58. 

Shell  umbilicate,  depressed,  acutely  carinated  at  the  periphery, 
low  roiiiral  above,  convex  below;  chestnut  brown,  becoming  lighter 
toward  the  apex;  lightly,  obliquely  striated.  Whorls  5,  the  last 
descondinir  to  the  oblique,  transversely  oval  aperture;  lip  expanded, 
ivil.-xed,  pinkish-white,  its  ends  approaching,  the  columellar  termina- 
tion half-concealing  the  umbilicus. 

Tlu-  convexity  is  about  equal  above  and  below  the  pinched-out 
keel  ;  the  earlier  whorls  are  convex,  the  latter  two  flattened,  the  last 
whorl  being  excavated  just  above  and  below  the  keel,  and  rather 
abruptly  deflexed  in  front. 

Alt.  14,  greater  diara.  23,  lesser  19  mill.;  width  of  umbilicus  2 
mill.;  oblique  alt.  of  aperture  11,  width  15  mill.,  measurements  in- 

cluding peristome. 

Port  Blair,  Andaman  Is. 

H.  trochalia  BENS.,  Ann.  Mag.  N.  H.  3d.  Ser.,  vii,  p.  82,  Feb., 
1861.—  PFR.,  Monogr.  v,  p.  329.—  HANL.  &  THEOB.,  Conch.  Indica. 
t,  38,  f.  7.  —  STOLICZKA,  Proc.  Asiat.  Soc.  Beng.,  March,  1870,  p.  87. 
—  TRYON,  Amer.  Journ.  Conch,  vi,  p.  287.  —  Ampelita  (Helix') 
Bigsbyl  TRYOX,  Amer.  Journ.  Conch,  v,  p.  110,  t.  10,  f.  3. 

A  peculiar  shell,  of  doubtful  affinities,  but  apparently  more  nearly 
related  to  the  Indo-malayan  species  with  which  I  above  associate  it, 
than  to  Obba  or  Obbina. 

Unfigured,  undetermined  species. 

H.  TRITONIENSIS  Le  Guillou.  Shell  trochoidal,  sub-umbilicate, 
whitish-tawny,  substriate  ;  whorls  5-6,  a  little  convex,  the  last 
rounded,  convex  beneath  ;  spire  conic-depressed;  aperture  narrow; 
lip  .sub-reflexed  ;  umbilicus  small,  punctiform. 

Alt.  7-],  diam.  9  mill.     (Gidll  in  Revue  Zool.  1842,  p.  138.) 

Triton  Bay,  New  Guinea. 

H.  HYALIXA  Le  Guillou.  Shell  trochoidal,  carinated,  scarcely  um- 
bilicatcd,  hyaline,  subvitreus.  \Yhorls  6,  subconvex,  the  base  de- 
pressed ;  delicatelv  striated,  the  last  having  an  acute  carina  a  little 

*/  o 

rell«-\r  1  upward,  convex-depressed  beneath  ;  aperture  sublanceolate, 
acute,  anteriorly,  base  rounded,  subreflexed,  obliquely  straightened 
nl  the  umbilicus.  Alt.  10,  diam.  15  mill.  (Gui.ll.) 

Solomon  Islands. 


HELIX-ALBERSIA.  89 

H.  hyalina  LE  GUILL.,  Revue  Zoologique  1842,  p.  139. — H.  re- 
flexiuscula  PFR.,  Symbolse  ii,  p.  98,  Monogr.  i,  p.  327. 

PfeifFer  suggests  the  name  reflexiuscula  because  hyalina  was 
supposed  to  be  preoccupied.  The  species  has  not  been  recognized 
by  anyone  since  Le  Guillou. 

H.  TESTUDO  Pfeiffer.     PI.  18,  fig.  64. 

Imperforate,  trochiform,  rather  solid,  striate  and  decussated  by 
oblique  wrinkles  ;  pale  isabelline,  painted  with  spots  and  flames  of 
brown ;  spire  conoid,  rather  acute ;  suture  carino-marginate. 
Whorls  5,  a  little  convex,  regularly  increasing,  the  last  acutely 
carinated,  tumid  above,  slightly  deflexed  in  front ;  base  subplanate. 
Aperture  very  oblique,  rhombic-lunar  ;  peristome  whitish,  upper 
margin  expanded,  basal  reflexed,  columellar  margin  wide,  flat, 
adnate.  Alt.  17,  greater  diam.  35,  lesser  28  mill.  (Pfr.) 

Madagascar. 

H.  testudo  PFR.,  P.  Z.  S.  1858,  p.  21,  t,  40,  f.  6 ;  Mai.  Bl.  1859,  p. 
14;  Monogr.  v,  p.  271. 

If  this  is  really  from  Madagascar  I  would  refer  it  to  Ampelita. 
The  original  figure  is  copied  on  my  plate. 

Section  ALBERSIA  H.  Adams,  1865. 

Albersia  H.  AD.,  P.  Z.  S.  1865,  p.  410.  Type  H.  granulata  Q.  & 
G. — PFR.,  Nomencl.  Hel.  Yiv.,  p.  198,  1878. — v.  MARTENS,  Ostas. 
Zool.  Landschn.,  p.  329,  1867. — TAPPARONE  CANEFRI,  Ann.  Mus. 
Civ.  Genov.  xix,  p.  185, 1883. 

Shell  globose  ;  «aperture  but  little  oblique,  the  peristome  scarcely 
thickened,  narrowly  reflexed  ;  columellar  margin  steeply  ascending, 
narrowed  below.  Unicolored,  never  brilliantly  colored,  often 
hirsute. 

This  group  should  probably  be  regarded  as  a  section  of  the  genus 
or  subgenus  Chloritis.  This  view  of  its  position  is  that  taken  by  H. 
Adams  and  Tapparone  Canefri.  Von  Martens,  however,  expresses 
the  opinion  that  its  affinities  are  with  the  Philippine  Island  group 
Cochlostyla  (s.  lat.)  ;  and  he  is  followed  by  Pfeiffer,  who  places 
Albersia  at  the  end  of  the  Helices  between  Acavus  and  Axina. 

Albersia  is  herein  accepted  in  the  restricted  sense  as  it  is  under- 
stood by  Pfeiffer.  H.  najas,  H.  extensa  and  H.  obliquata  have  also 
been  included  by  some  authors,  but  they  are  not  separable  concho- 
logically  from  the  section  Corasia  of  Cochlostyla. 


(HI  HELIX-ALBERSIA. 

The  jaw  of  H.  pubicepa  is  arcuate  and  ribbed. 
H.  GRANULATA  Quoy  &  Gaimard.     PL  19,  figs.  1,  2. 

Shell  covered  perforate,  globose,  delicately  granulate,  golden  or 
rufous  encircled  with  a  brown  band.  Spire  short ;  whorls  5*,  nearly 
plane,  the  last  large,  inflated.  Aperture  ample,  lunate-elliptical ; 
peristome  little  reflexed,  white,  the  columellar  margin  emarginate, 
broadly  dilated,  wholly  closing  the  perforation. 

Alt.  33,  greater  diam.  50,  lesser  41  mill.     (Pfr.') 

Port  Doreij,  New  Guinea;  Is.  of  Waigheu. 

H.  granulata  Q.  &  G.,  Voy.  Astrol.  Zool.  ii,  p.  95,  t.  7,  f.  6-9.— 
FER.  &  DESH.,  Hist.,  p.  249,  t.  28 A,  f.  9-10.— RVE,  Conch.  Icon., 
t.  69,  f.  358.— PFR.,  Conchyl.  Cab.,  t.  137,  f.  13,  14;  Monogr.  i,  p. 
233;  iii,  p.  181.— H.  (Pomatia)  granulata  PFR.,  Versuch.  etc.,  p. 
•  ISS.—Albersia  granulata  WALLACE,  P.  Z.  S.  1865,  p.  410.— H. 
(Albersia)  granulata  PFR.,  Nomencl.  Hel.  Yiv.,  p.  198.— TAPPARONE 
CANEFRI,  Ann.  Mus.  Civ.  Genov.  xix,  p.  185. 

Variety.  Shell  dark  blackish-brown,  the  band  slightly  darker ; 
surface  shortly  hirsute ;  spire  a  little  higher ;  peristome  lilac,  broadly 
expanded  and  revolute. 

Alt.  33,  greater  diam.  50,  lesser  39.     (Pfr.) 

H.  PUBICEPA  Martens.     PI.  19,  figs.  9,  10,  11. 

Shell  subglobose,  thin,  striatulate,  downy,  the  hairs  standing  in 
quincunx ;  fleshy-brown,  with  a  reddish  peripheral  band,  pale, 
bordered  above  and  below.  Spire  convex ;  whorls  5,  the  last  in- 
flated, a  little  descending  in  front.  Aperture  moderately  oblique, 
lunate-rounded  ;  peristome  shortly  reflexed,  thin,  rufescent,  margins 
remote  from  one  another ;  columella  obliquely  descending,  thickened, 
flesh-color.  (Mts.) 

Alt.  24.  greater  diam.  36,  lesser  28  mill. 

Alt.  20 •>,  greater  diam.  29,  lesser  24  mill. 

Hiifniiiheira  and  Batjan,  Moluccas.  In  gardens  and  the  forest, 
not  common. 

Cochlostyla  pubicepa  MARTENS,  Monatsb.  Berl.  Akad.  1864,  p. 
26! ». — If.  (Albersia)  pubicepa  Mrs.,  Preuss.  Exped.  nach  Ostas., 
Landx-lin.,  p.  329,  t.  18,  f.  2. — PFR.,  Monogr.  v,  p.  281. — Helix 
tortistylis  Mousy.,  in  coll. 

Intermediate  between  the  coarsely  granulose  If.  granulata  and 
tli<-  smooth  H.  zonulata.  It  is  allied  to,  or  at  least  similar  to  H. 


HELIX-ALBERSIA.  91 

tennis  Pfr.,  the  habitat  of  which  is  unknown  ;  but  it  is  flatter,  the 
whorls  more  rapidly  increasing,  the  peristome  straight,  and  the  col- 
umella  differently  formed. 
The  jaw  is  arcuate,  ribbed. 

H.  ZONULATA.  Ferussac.     PL  19,  figs.  7,  8  (and  var.  f.  6.) 

Shell  imperforate,  depressed-globose,  thin ;  roughened  by  very 
minute  decussating  lines,  a  little  shining ;  of  a  tawny  flesh  color 
with  a  whitish  band  at  the  periphery,  and  a  narrow  rufous  band 
above  it.  Whorls  5,  a  little  convex,  the  last  ventricose ;  columella 
slender,  reddish,  calloused.  Aperture  rotund-lunar ;  peristome 
simple,  reflexed,  columellar  margin  dilated,  covering  the  place  of 
the  umbilicus,  its  junction  with  the  basal  margin  forming  a  sub- 
denticulate  angle. 

Alt.  15,  greater  diam.  26,  lesser  21  mill.     (Pfr.} 

Neiv  Guinea  at  Port  Dorey ;  Andai ;  Amberbaki;  Waigheu ; 
Wokan,  Wama  and  Giabu  Lengen,  Aru  Is. ;  Is.  of  Jobi,  in  the  bay 
of  Geelvink ;  Secaar,  S.-E.  New  Guinea. 

H.  listeri  FER.,  Prodr.,  p.  28,  no.  22,  (not  H.  listeri  Gray). — H. 
zonulata  FER.,  Prodr.,  p.  67  ;  Histoire,  t.  15,  f.  1,  2. — PFR.,  Conch. 
Cab.,  p.  310,  t.  53,  f.  3-5;  Monogr.  i,  p.  261. — MARTENS,  Ostas. 
ZooL,  p.  393. — Albersia  zonulata  WALLACE,  P.  Z.  S.  1865,  p.  410. — 
H.  (Albersia)  zonulata  TAPPARONE  CANEFRI,  Ann.  Mus.  Civ. 
Genov.  xix,  p.  185  ;  and  1.  c.  2d.  ser.,  iv,  p.  143, 1887. — H.  lemniscata 
LESSON,  Voy.  de  la  Coquille,  p.  306. — H.  reuluziana  LE  GUILL.  Rev. 
Zool.  1842,  p.  137.— PER.,  Monogr.  i,  334.  * 

Var.  RECLUZIANA  Le  Guill.     PI.  19,  fig.  6. 

Shell  larger.     Alt.  17,  greater  diam.  30,  lesser  24  mill. 

VAR.     Shell  smaller,  rather  solid,  peristome  without  a  tooth. 
Alt.  12,  greater  diam.  19,  lesser  16  mill.     (Tap.  Can.) 

Andai;  Amberbaki. 

H.  TENUIS  Pfeiffer.     Vol.  V,  pi.  6,  figs.  56,  57. 

Shell  imperforate,  subglobose,  thin,  brown  with  a  rufous  zone, 
marked  with  obsolete  impressed  concentric  lines,  covered  with  a  very 
thin  silky  epidermis;  spire  rather  obtuse ;  whorls  5,  rapidly  increas- 
ing, slightly  convex,  the  last  large,  inflated,  excavated  at  the  oblique, 
subtortuous  columella.  Aperture  lunate-oval ;  peristome  straight, 
simple,  subthickened  at  the  base. 

Alt.  20,  greater  diam.  32,  lesser  26  mill.     (Pfr.) 

Habitat  unknoivn. 


92  COOHLOSTYLA. 

H.  tennis  PFR.,  P.  Z.  S.  1845,  p.  41 ;  Monogr.  i,  p.  24  ;  Conchyl. 
Cab.,  t.  157,  f.  5,6.— REEVE,  Conch.  Icon.,  f.  759.— H.  (Enry  crater  a) 
tennis  PFR.,  Xomencl.  Hel.  Yiv.,  p.  169,  1878. 

A  species  known  only  by  Pfeiffer's  and  Reeve's  descriptions  and 
figures. 

Genus  COCHLOSTYLA  Ferussac,  1819. 

Cochlostyla  FERUSSAC,  Tab.  Syst.  des  Anim.  Moll.  Prodr.  gen.,  p. 
47,  1819. — PFR.,  Symbol*  i,  p.  21. — v.  MARTENS,  in  Albers,  Die 
Heliceen  2d.  edit.,  p.  173;  Preuss.  Exped.  Landmoll.,  p.  96. — 
SK.MPER,  Reis.  Archip.  Phil.,  p.  164. 

COCHLOSTYLA  includes  Cklorcea,  Corasia,  Axina,  Pfeifferia,  Calo- 
cocldia,  Helicostyla,  Orustia,  Cochlodryas,  Orthostylus,  Helicobulimus, 
Ptychostyliis,  Pkengus,  Eudoxus,  Hypselostyla,  Canistrum,  Prochilus, 
Chrysallis,  Phoenicobius. 

Shell  varying  from  depressed,  Helicoid,  to  elongated  and  Bulimi- 
form,  usually  imperforate  ;  rounded  at  the  circumference  or  keeled; 
the  lip  generally  reflexed  (rarely  simple  and  acute)  ;  usually  covered 
with  a  thin,  transparent  cuticle,  often  porous  in  places,  when  it  be- 
comes white  and  opaque,  producing  the  "  hydrophanous  ':  patterns 
with  which  most  of  the  species  are  ornamented. 

The  species  are  with  few  exceptions  confined  to  the  Philippine 
Archipelago.  They  are  mainly  arboreal  in  habit. 

The  species,  however  diverse  in  form  of  the  shell,  agree  in 
anatomy.  There  are  no  accessory  appendages  to  the  male  genital 
system  ;  the  female  system  has  a  single  mucus  gland,  globular  or 
oval  in  shape,  which  combines  with  the  sac  of  the  simple  stylet- 
formed  dart.  The  duct  of  the  receptaculum  seminis  is  long.  The 
jaw  has  strong  ribs. 

Diverse  as  the  shells  assembled  in  this  genus  at  first  appear,  it  is 
without  doubt  a  perfectly  natural  group.  The  extreme  forms  are  so 
closely  connected  by  intermediate  species  that  no  satisfactory  groups 
of  subgeneric  rank  can  be  defined. 

The  system  of  sections  or  groups  briefly  sketched  by  v.  Martens 
(Ostas.  Zool.,  p.  96),  and  improved  by  Semper  (Phil.  Archip.  Land- 
nioll.)  is  in  my  opinion  the  most  natural  yet  proposed.  I  have, 
therefore,  in  the  following  pages  followed  the  same  general  arrange- 
ment and  sequence  of  groups;  but  not  without  modifying  them 
materially  as  to  contents  and  limits,  and  adding  several  groups  to 
the  already  large  number.  In  the  system  herein  set  forth,  the  species 


COCHLOSTYLA-CHLOR^EA.  93 

are  divided  into  twenty  sections,  the  series  beginning  with  the  de- 
pressed, and  ending  with  the  elongated  species.  A  table  exhibiting 
the  characters  of  the  sections  will  conclude  the  account  of  the 
species. 

The  principal  special  authorities  consulted  on  the  Philippine 
Island  fauna  are : 

Dr.  C.  Semper,  Reisen  im  Archipelder  Philippinen,  vol.  iii,  Land- 
mollusken. 

We  are  indebted  to  Semper  for  a  vast  amount  of  information  on 
the  distribution  as  well  as  the  anatomy  of  Philippine  land  snails. 

Dr.  O.  F.  von  Mollendorff,  Die  Landschnecken  Fauna  der  Insul 
Cebu,  in  Bericht  liber  die  Senckeubergische  naturforschende  Gesell- 
schaft  ZLI  Frankfurt  a.  M.,  1890.  Also  numerous  papers  published 
in  Nachrichtsblatt  der  Deutschen  Malakozoologischen  Gesellschaft. 

A  most  excellent  paper  on  the  shells  of  the  island  Zebu. 

J.  G.  Hidalgo,  Obras  Malacologicas,  parte  I,  Estudios  preliminares 
sobre  la  Fauna  Malacologica  de  las  Mas  Filipinas,  in  Memorias  de 
la  Real  Academia  de  Ciencias  Exactas,  Fisicas  y  naturales  de 
Madrid,  xiv.  Also  papers  by  the  same  author  in  the  Journal  de 
Conchyliologie. 

Of  Sr.  Hidalgo's  Estudios,  etc.,  only  160  pages,  30  plates,  have  as 
yet  (Sept.,  1891)  been  published,  containing  of  the  genus  Coclilostyla 
only  Corasia,  Axina  and  part  of  Chlorcea.  This  magnificent  work 
will  when  completed,  leave  little  to  be  desired  as  far  as  the  illustra- 
tion of  the  shells  is  concerned.  The  text  is  of  equal  merit,  being 
characterized  by  a  broad  conservatism,  and  excellent  judgment  in 
the  treatment  of  species. 


Section  I.  CHLOR^A  Albers,  1850. 

Chlorcea  ALB.,  Die  Hel.,  p.  113. — v.  MART,  in  Albers,  2d.  edit., 
p.  169,  (type  H.  sirena  Beck). — SEMPER,  Reis.  Archip.  Phil.,  Land- 
moll.,  p.  226,  (in  part.) 

Imperforate,  varying  from  depressed-globose  to  lens-shaped, 
generally  solid  ;  whorls  4-5,  the  last  descending  or  not ;  lip  decidedly 
reflexed,  at  least  below.  Entire  surface  showing  under  a  lens 
excessively  fine,  close  spiral  lines;  lacking  hydrophanous  patches  on 
the  cuticle. 

Anatomy  as  in  Cochlostyla  generally,  except  that  the  simple 
mucus  gland  is  aciniform,  not  globular.  C.  sirena  has  a  short 
flagellum  on  the  penis. 


94  COCHLOSTYLA-CHLOR^EA. 

Heliciform  shells,  distinguished  at  once  from  the  closely  allied 
section  Corasia  by  the  excessively  close,  fine  spiral  striation  covering 
the  whole  surface  both  above  and  below,  except  the  smooth  apical 
embryonic  whorl.  These  snails  live  on  trees,  like  Corasia  and  most 
other  Cochlostylas. 

The  present  group  is  equivalent  to  Semper's  "  Gruppe  der  bunt- 
schaligen  Chlorceen"  His  "  Gruppe  der  einfarbigen  Chlorceen"  con- 
sists of  species  of  Dorcasia.  They  differ  from  the  true  Chlorcea 
species  in  having  a  compound  mucus  gland  ("  multifid  gland  "),  com- 
posed of  several  acinose  caeca,  whilst  that  gland  in  Chlorcea,  as  in 
all  Cochlostyla,  is  simple  ;  the  shells  also  are  notably  different. 

The  species  of  Chlorcea  fall  into  four  groups,  as  follows : 

A.  Latter  half  of  the  body-whorl  in  no  part  narrower  than  the 
adjacent  part  of  the  preceding  whorl,  viewed  from  above. 

1.  Solid,  keeled  or  rounded,  usually  banded.     Group  of  C.  fibula. 

2.  Thin,  acutely  keeled;   greenish,  often  banded. 

Group  of  C.  dry  ope. 

B.  Latter  half  of  the  body-whorl  narrower  in  some  part  than  the 
preceding  whorl,  viewed  from  above. 

3.  Last  ivhorl  not  acutely  keeled,  descending  in  front. 

Group  of  C.  paradoxa. 
2.  Last  whorl  acutely  keeled,  not  descending  in  front. 

Group  of  C.  thersites. 

(1)    GROUP  OF  C.  FIBULA. 

Solid,  varying  from  depressed-globose  to  lens-shaped,  carinated 
or  rounded  at  the  periphery,  the  last  whorl  circular  in  circumfer- 
ence, not  laterally  flattened  on  its  latter  half;  descending  in  front; 
lip  thickened  and  reflexed ;  usually  banded. 

These  shells  are  more  solid  than  in  the  closely  allied  group  of  C. 
dryope ;  they  do  not  have  the  last  whorl  compressed  at  the  periphery 
as  in  the  group  of  C.  paradoxa. 

C.  FIBULA  Broderip.     PI.  22,  figs.  21,  22,  23;  pi.  21,  figs.  31,  32, 

Oo. 

Imperforate,  depressed,  solid,  the  entire  circumference  acutely 
carinated  ;  white  or  pale  yellowish,  generally  three-banded ;  a  broad 
purplish-brown  band  below  and  another  above  the  periphery,  the 
latter  ascending  the  spire  in  company  with  a  narrow  dark  line  border- 
ing the  lower  edge  of  the  suture. 


COCHLOSTYLA-CHLOR.EA.  95 

The  shell  is  about  equally  ccmvex  above  and  below;  whorls  4-4J, 
the  last  abruptly  descending  in  front,  and  a  little  constricted  behind 
the  lip.  Surface  very  finely,  spirally  striated.  Aperture  very 
oblique  ;  peristome  white,  narrowly  expanded  above,  reflexed  below  ; 
columellar  margin  adnate.  Alt.  11,  diam.  20  mill. 

Luzon  ;  Lubang ;  Marinduque ;  Zebut 

Car ocolla  fibula  BROD.,  mss. — REEVE,  Conch.  Syst.  ii,  t.  167,  f.  5. 
— Helix  fibula  PHIL.,  Abbild.  i,  p.  12,  t.  1,  f.  8. — PFR.,  in  Conchyl. 
Cab.,  t.  69,  f.  5-7 ;  Monogr.  i,  p.  300.— REEVE,  Conch.  Icon.,  f.  137. 
— HIDALGO,  Obras  Malacol.  i,  p.  156,  t.  25,  f.  4,  5  (type),  6-8 
(vars.) 

This  is  a  larger  species  then  C.  hanleyi,  and  the  whorl  is  less  de- 
flexed  in  front.  Some  forms  of  C.  hugeli  approach  it,  but  they  are 
less  acutely  keeled  behind  the  lip,  and  differ  in  coloring.  Hidalgo 
describes  the  following  color-varieties  : 

1.  Whitish  or  whitish-yellow,  unifasciate  above  and  below. 

2.  No  band  beneath. 

3.  Smaller;  white,  not  banded.      From  Marinduque  (pi.  21,  fig. 
33.) 

4.  Chestnut  colored  above,  with  a  white  peripheral  band  continu- 
ing up  the  spire  above  the  suture;  base  4-zoned,  the  two  coliimellar 
zones  tawny,  outer  paler,  the  third  zone  chestnut,  the  peripheral  zone 
white  (pi.  21,  figs.  31,  32.) 

5.  Buff,  not  banded  below.     Island  of  Lubang. 

C.  HANLEYI  Pfeiffer.     PI.  22,  figs.  24,  25,  26,  27,  28. 

Imperforate,  depressed,  solid,  acutely  carinated;  light  greenish- 
yellow,  a  band  at  the  carina  and  a  border  below  the  suture  opaque 
white.  Last  whorl  very  deeply  deflexed  in  front,  the  aperture  nearly 
horizontal. 

The  shell  is  about  equally  convex  above  and  below;  surface 
microscopically  striate  spirally.  Whorls  4,  flat,  the  entire  circum- 
ference of  the  last  whorl  acutely  carinated.  Aperture  small,  oval, 
subhorizontal ;  the  peristome  white,  reflexed,  its  terminations 
approaching.  Alt.  9,  diam.  17  mill. 

Sinait,  Centro  del  Abra,  Monte  Vigan,  Ilocos  Sur ;  Bayabas 
province  of  Angat,  and  Santander,  province  of  Morong,  in  the  island 
of  Luzon,  Philippines. 

H.  hanleyi  PFR.,  P.  Z.  S.  1845,  p.  65  ;  Conchyl.  Cab.,  p.  31,  t.  69, 
f.  15,  16  (and  17,  18,  var.). — HIDALGO,  Obras  i,  p.  158. — H.  hugeli 
RVE.,  (part.)  f.  136e,  (and  136d,  var.) 


96  COCHLOSTYLA-CHLOBJEA. 

In  the  typical  C.  hanleyi  (pi.  22,  figs.  24-26)  the  whorl  is  very 
deeply  deflexetl  in  front,  the  suture  and  carina  are  bordered  below 
with  white.  There  are  no  other  bands  on  the  pale  greenish-yellow 
ground-color. 

( 'olor  var.  (pi.  22,  figs,  27,  28).  Blackish-purple  above,  banded 
with  white  at  suture  and  carina;  base  buff-white,  having  a  rufous 
band  below  the  carina.  Form  as  in  the  type. 

Var.  HI-GELI  Pfr.     PL  22,  figs.  15,  18,  19,  20. 

Slightly  larger  than  typical  hanleyi,  less  acutely  carinated,  the 
aperture  less  oblique  and  larger.  Color  greenish-buff  with  a  white 
girdle  at  periphery,  bordered  above  and  below  by  narrow  green 
bands.  Alt.  12,  diam.  18  mill. 

( 'arocolla  bifasciata  LEA,  Trans.  Amer.  Philos.  Soc.  (n.  ser.)  vii, 
1841,  p.  465,  t.  12,  f.  18,  (not  Helix  (Oxychona)  bifasciata  BURROW). 
-H.  hugeli  PFR.,  Symbolse  iii,  p.  31  ;  Monogr.  i,  p.  300;  Conchyl. 
Cab.,  p.  31,  t.  69,  f.  8-14.— FEB.,  Hist.,  t.  95,  f.  3  (central  fig.).— 
REKVK,  Icon.,  f.  136c. —  (7.  (Chlorcea)  hanleyi  var.  C.  HIDALGO, 
Obras  Malacol.  i,  p.  158,  t.  25,  f.  10. 

The  name  proposed  for  this  species  by  Lea  in  1841,  is  not  pre- 
occupied in  Cochlostyla,  and  it  should  probably  be  adopted. 

A  variation  of  this  form  (pi.  22,  fig.  15)  has  three  narrow  brown 
bands ;  one  below,  another  above  the  periphery,  and  the  third  closely 
bordering  the  suture  below.  The  basal  band  is  sometimes  lacking 
in  this  color-variety.  Pfeiffer's  figs.  11,  12,  of  pi.  69,  Conchyl.  Cab., 
represent  this  form. 

Hidalgo  believes  that  Pfeiffer  confused  two  forms  under  the  name 
lltlfjeli;  and  considers  the  figures  10  and  14  to  represent  the  true 
hi'ir/eli,  of  which  he  considers  Semper's  C.  antonii  a  synonym.  In 
this  view  I  am  unable  to  concur.  Pfeiffer's  fig.  10  is  reproduced  on 
my  pi.  22,  fig.  20. 

C.  I;I:N<;I:ETENSIS  Semper.     PL  21,  figs.  23,  24,  25. 

Im perforate,  lenticular,  thin,  acutely  carinated,  obliquely  striat- 
ulate,  whitish  ornamented  with  a  buff  band  above  the  carina  and  a 
chestnut  line  at  the  suture;  spire  little  elevated;  whorls  4,  rather 
flattened,  the  last  deflexed  and  constricted  in  front.  Aperture  large, 
aubhorizontal,  angular-elliptical;  peristome  simple,  margins  remote, 
the  upper  margin  slightly  expanded,  basal  margin  reflexed. 

Alt.  11,  greater  diam.  23,  lesser  18  mill.     (Semper.) 

Valley  of  Benguet,  Luzon,  Philippines. 


COCHLOSTYLA-CHLOR^EA.  97 

(Miloraia  benguetensis  SEMPER,  Eels.  Archip.  Phil.,  Landmoll.,  p. 
227,  t.  viii,  f.  11,  12  (shell  and  animal) ;  t.  xiv,  f.  5-8  (anatomy).— 
HIDALGO,  Obras  Malacol.  i,  p.  .158,  t.  25,  f.  1-3. 

The  color-mutations  recorded  by  Hidalgo  are  as  follows : 

1.  (Typical).     Whitish,  with  a  chestnut  sutural  line  and  a  buff 
band  above  the  periphery  (figs.  23,  24.) 

2.  Unicolored,  white. 

3.  Pale  straw-colored,  with  a  narrow  white  peripheral  band,  nar- 
rowly bordered  above  and  below  with  green,  (fig.  25.) 

"  The  second  and  third  varieties  have  much  similarity  to  C.  hanleyi 
and  its  variety  bifasciata  Lea,  but  they  have  the  aperture  more 
produced  transversely,  the  peristome  narrower,  and  the  last  whorl 
more  inflated,  around  the  basal  margin  of  the  peristome,  as  it  is  in 
typical  benguetensis,  of  which  I,  therefore,  consider  them  varieties." 
(Hidalgo.) 

C.  GEOTROCHUS  Mollendorff. 

Imperforate,  trochiform,  delicately  obliquely  striatulate  and 
decussated  with  close  spiral  lines ;  chestnut-brown  with  a  whitish 
band  at  the  carina  and  another  below  the  narrow  brown  subsutural 
zone  ;  spire  subregularly  conical,  apex  obtuse,  whitish.  Whorl  4J, 
nearly  flat,  the  last  acutely  white-carinated,  shortly  deflexed  in  front, 
swollen  toward  the  aperture  beneath,  having  a  pale  yellowish  band 
around  the  columella.  Aperture  very  oblique,  angulate-elliptical ; 
peristome  expanded,  albolabiate ;  columella  excavated,  oblique. 

Alt.  12,  greater  diam.  21*,  lesser  18  mill.     (Moll.) 

Mt.  Limutan,  Province  of  Morony,  Luzon. 

C/ilorcea  geotrochus  v.  MOLL.,  Nachrichtsbl.  D.  M.  Ges.  1888,  p. 
89. 

This  species  has  much  the  aspect  of  Papuina  cleryi.  It  is  closely 
allied  to  C.  hanleyi  and  C.  benguetensis. 

C.  ANTONII  Semper.     PI.  21,  figs.  34,  35. 

Imperforate,  globose,  shining,  spirally  very  minutely  striated,  pale 
green  or  straw  colored,  having  two  reddish  lines  above,  sometimes 
ornamented  with  a  brown  band  below;  spire  elevated;  suture 
smooth  ;  whorls  4-},  convex,  the  last  deflexed  in  front ;  columella 
short;  aperture  lunate-rounded;  peristome  simple,  subexpanded, 
margins  conniving. 

Alt.  13,  greater  diarn.  18,  lesser  15  mill.     (Semp.) 

Northivestern  Luzon. 
7 


Jl.s  COCHLOSTYLA-CHLOR^EA. 

Chlorcea  antonii  SEMPER,  Reis.  Phil,  Landmoll.  iii,  p.  228,  t.  10, 
f  loai  \),—H.  (Chlorcea)  hugeli  HIDALGO,  Obras  Malacol.  i,  p.  160. 

This  species  I  have  not  seen.  It  appears  to  be  more  globose,  with 
narrower  lip  and  less  oblique  aperture  than  C.  hugeli  Pfr. 

C.  AM.KXA  Pfeiffer.     PL  21,  figs.  21,  22. 

I  m  perforate,  depressed-globose,  solid  ;  white,  having  a  broad  purple- 
black  zone  above  the  periphery  and  ascending  the  spire.  Apex  and 
Up  white;  hist  whorl  abruptly  and  deeply  deflexed  in  front;  not 
cxrinated,  often  obscurely  angled  just  above  the  upper  insertion  of 
the  lip. 

Spire  convex,  apex  obtuse  ;  whorls  4£,  the  last  abruptly  descend- 
ing in  front,  slightly  constricted  behind  the  lip.  Surface  lightly 
obliquely  striatulate  and  microscopically,  spirally  striated.  Aperture 
very  oblique,  oval-truncate,  showing  the  band  within ;  peristorne 
narrowly  thickened  and  reflexed,  the  baso-columellar  margin  ad n ate, 
straightened,  thickened  within.  Alt.  12,  greater  diam.  16  mill. 

Luzon,  Philippines. 

H.  amcena  PFR.,  P.  Z.  S.  1845,  p.  65  ;  Conchyl.  Cab.,  t.  69,  f.  19, 
20  ;  Monogr.  i,  p.  270. — H.  hugeli  var.  REEVE,  Conch.  Icon.,  f.  136a. 

A  solid,  subglobose  species,  allied  to  C.  hugeli,  but  not  distinctly 
carinated  and  differently  colored.  It  is  much  smaller  than  C. 
paradoxa,  restricta  or  undina. 

C.  sfRENA  Beck.     PL  21,  figs.  J5-20. 

Imperforate,  globose-depressed,  thin  but  solid,  the  periphery 
rounded  except  just  in  front  of  the  aperture  where  it  is  carinated; 
sometimes  uniform  straw-yellow  with  white  apex,  but  usually  pale 
cream-white  or  pale  green  with  the  apical  whorl  pink,  the  body- 
whorl  encircled  with  one  or  two  green  or  greenish-purple  zones, 
visible  as  vivid  brown  bands  within  the  aperture,  the  upper  band 
aMvn.liiin-  the  spire  above  the  suture,  and  becoming  brown  there, 
and  the  peristome  dark  brown.  Spire  low,  convex;  suture  super- 
ficial ;  whorls  4,  the  last  descending  rather  abruptly  in  front.  Aper- 
ture oblique,  wide-lunar  ;  peristome  scarcely  expanded  above,  outer 
and  l.asil  margins  narrowly  expanded,  columellar  margin  adnate. 

Alt.  14,  diam.  2lM  mill. 
Ouimaras;   /'<///«//;  Z<i,ii/><><ui.f/<t)  Mindanao;  Zebu;  Philippines. 

II.  sirena  HKCK  (in  sched.  Cuming)  Pfr.,  Symbols  ii,  p.  39; 
<  Jonchyl.  ( lab.  ii,  p.  31:5,  t.  f>3,  f.  14-17  ;  Monogr.  i,  p.  263.— REEVE, 


COCHLOSTYLA-CHLOIkEA.  99 

Conch.  Icon.,  f.  96. — PHILIPPI,  Abbild.  i,  p.  78,  Helix,  t.  4,  f.  6. — 
HIDALGO,  Journ.  de  Conchyl.  1887,  p.  I21--Cfilorcea  sirena  v. 
MOLL.,  Landschn.  Fauna  Cebu,  p.  226  (var.  cebuana.") 

Subglobose,  with  the  whorl  less  deflexed  in  front  than  most  species 
of  Chlorcea.  It  is  very  variable  in  coloring. 

Typical  sirena  (figs.  16,  17,  18)  has  a  cream  colored  or  light 
greenish  ground-color,  with  a  single  band  above  the  periphery, 
which  is  brown  on  the  spire,  becoming  purplish,  green  or  blackish- 
green  on  the  body-whorl.  There  are  often  dots  of  translucent-gray- 
ish scattered  over  the  surface. 

Var.  cebuana  v.  Moll.  Smaller,  a  little  higher,  last  whorl  not  at 
all  angulated,  more  inflated  below ;  color  dull  green,  with  a  pale 
greenish  peripheral  girdle,  bordered  above  and  below  with  narrow, 
dark  green  bands. 

Island  of  Zebu. 

Var.  guimarasensis  Pils.  (fig.  15).  Larger  than  the  type,  dis- 
tinctly angular  in  front,  unicolored  yellow,  paler  toward  the  white 
apex.  Alt.  161,  diam.  26  mill. 

Island  of  Guimaras. 

C.  PELEWANA  Mousson.     PI.  22,  figs.  10,  11. 

Imperforate,  orbiculate-depressed,  striate,  sculptured  all  over  with 
undulating  oblique  and  subspiral  w r inkles  ;  shining,  white,  painted 
above  with  3  or  4  interrupted  blackish-brown  lines.  Spire  obtuse, 
depressed,  the  nucleus  roseate,  polished  ;  suture  simple,  subimpressed. 
Whorls  3i,  rapidly  increasing,  a  little  convex,  the  last  obtusely  car- 
inated,  abruptly  descending  at  the  aperture  and  somewhat  narrowed ; 
at  the  base  rather  convex.  Aperture  oblique  (inclined  at  60  degrees 
with  the  axis),  transverse  lunate-subcircular,  the  bauds  visible  inside ; 
peristome  acute,  expanded,  white,  sublabiate  inside,  margin  little 
approaching,  joined  by  a  pale,  thin  callus ;  right  margin  expanded, 
arcuate,  columellar  margin  very  oblique,  broadly  and  obtusely 
thickened,  flatly  expanded  above,  adnate. 

Alt.  13,  diam.  19  mill.     (Mouss.') 

Pelew  Is. 

H.  pelewana  Mouss.,  Journ.  de  Conchyl.  1869,  p.  58,  t.  4,  f.  2.- 
Chlorcea  pelewana  PSE.,  P.  Z.  S.  1871,  p.  474. 

The  fauna  of  the  Pelew  group  is  essentially  Polynesian,  the 
occurrence  of  this  and  a  few  other  western  forms  being  quite  excep- 


100  COCHLOSTYLA-CHLOR.EA. 

tional.     C.    pelewana    may    prove   to   belong   elsewhere    than    in 
( '/tlorcea. 

( '.  OMELINIANA  Pfeiffer.     PL  21,  figs.  29,  30. 

Im perforate,  globose-depressed,  rather  solid,  irregularly  rugosely 
malleated,  carinated,  shining,  pale  green,  girdled  with  white  at  the 
suture  and  below  the  carina;  spire  subelevated,  apex  obtuse,  whitish; 
whorls  4-j,  scarcely  convex,  gradually  increasing,  the  last  not  descend- 
infj;  base  a  little  flattened  ;  aperture  angular-lunar;  peristome  sub- 
thickened,  the  upper  margin  narrowly  expanded,  arched  forward; 
basal  margin  reflexed ;  columella  sloping,  dilated,  white  calloused. 

Alt.  13,  greater  diam.  23,  lesser  20  mill.     (Pfr.) 

Luzon,  Philippines. 

H.  gmelimana  PFR.,  P.  Z.  S.  1845,  p.  43;  Monogr.  i,  p.  300; 
Conchyl.  Cab.,  t.  150,  f.  10,  11. — REEVE,  Conch.  Icon.,  f.  138. 

A  species  known  only  by  the  description  of  Pfeiffer  and  the  figures 
of  Pfeiffer  and  Reeve.  I  have  italicized  the  more  important  portions 
of  the  diagnosis.  Reeve  remarks  :  A  delicate  and  very  characteristic 
species,  of  which  the  surface  is  irregularly  indented  throughout.  It 
is  of  a  peculiar  light  sea-green  color  encircled  by  a  narrow  opaque- 
white  band  of  epidermis  beneath  the  spiral  keel. 

(2)  GROUP  OF  C.  DRYOPE. 

Depressed,  rather  thin,  greenish  or  bluish,  usually  banded  ;  acutely 
carinated ;  last  whorl  strongly  deftexed  in  front ;  apex  and  columella 
usually  dark. 

These  shells  are  separated  from  the  C.  regince  group  of  section 
Corasia  by  the  excessively  fine  spiral  striation  of  the  surface.  The 
spiral  lines  are  much  coarser  or  more  separated  in  Corasia. 

C.  DRYOPE  Broderip.     PL  22,  figs.  1,  2,  3,  4,  5,  6,  7. 

I  in  perforate,  much  depressed,  acutely  carinated,  rather  thin ;  light 
bluish-green,  the  apical  ivhorl  purple-black,  and  usually  there  is  a  dark 
bund  above  the  carina  and  a  dark  columeUar  spot;  the  peristome 
either  white  or  brown.  Upper  and  lower  surface  about  equally  con- 
vex ;  under  a  lens  the  entire  surface  is  seen  to  be  very  finely  and 
very  closely,  spirally  striated.  Whorls  4] ,  flattened,  the  last  deeply 
in  front;  aperture  subhorizontal,  lanceolate-oval;  per- 


COCHLOSTYLA-CHLOR^A.  101 

istome  expanded  above,  reflexed  below,  the  baso-columellar  margin 
adnate,  arcuate. 

Alt.  12,  diam.  27  mill. 

Alt.  9,  diam.  19  mill. 

Province  of  Cagayan,  Luzon;  Burias ;  Tablets,  and  Sibuyan 
Philippine  Is. 

H.  dryope  BROD.,  P.  Z.  S.  1841,  p.  37. — REEVE,  Conch.  Syst.  ii, 
t.  168,  f.  10;  Conch.  Icon.,  f.  124.— PFR.,  Conchyl.  Cab.,  p.  315,  t. 
54,  f.  3?  4.— DESH.  in  Fer.,  Hist.,  t.  102,  f.  13,  14.— H.  prasina 
KOCH,  in  PHIL.,  Abbild.  i,  p.  151.  t.  5,  f.  1. — H.  (Chlorced)  dryope 
HIDALGO,  Obras  Malacol.  i,  p.  154. 

This  species  resembles  some  forms  of  C.  (Corasia)  regince,  but  the 
spire  is  raised,  the  apex  blackish,  and  the  last  whorl  deeply  deflexed 
in  front.  It  is  thinner  than  the  other  species  of  Chlorcea.  Semper 
did  not  find  the  species  on  the  island  Burias,  whence  it  was  reported 
by  Cuming. 

The  principal  color  varieties  represented  in  the  suite  before  me  are 
figured  on  my  plate. 

C.  CCERULEA  Mollendorff.     Unfigured. 

Imperforate,  depressed,  thin,  delicately  curved  striate,  decussated 
by  extremely  close  spiral  lines;  pellucid,  greenish-blue;  spire  de- 
pressed, apex  plane,  brown  ;  whorls  4,  flat,  rapidly  increasing,  the 
last  with  a  very  acute,  exserted  white  carina;  base  inflated,  gibbous 
before  the  aperture,  greenish,  at  thecolumellanodiferous.  Aperture 
oblique,  rounded-trapezoidal ;  peristome  yellow,  sublabiate,  expanded 
above,  expanded  a  little  at  the  keel,  appressed  and  whitish  toward 
the  columella;  columella  dilated,  sloping,  purple-brown  above,  then 
white,  armed  at  the  base  with  a  tooth-like  nodule. 

Alt.  11,  greater  diam.  23,  lesser  19  mill.     (Moll.) 

Confines  of  provinces  Manila,  Bulacan  and  Morong,  Luzon. 

Cochlostyla  (Corasia)  ccerulea  v.  MOLL.  Nachrichtsbl.  D.  M.  Ges. 
July-Aug.,  1888,  p.  97. 

This  species  recalls  C.  regince,  but  it  is  smaller,  the  last  whorl  not 
so  much  widened  ;  the  color  is  a  beautiful  sky-blue,  tinged  with 
bluish-green,  the  apex  brown  ;  the  lip  is  distinctly  but  slightly 
thickened,  and  of  a  beautiful  yellow  as  far  as  the  columella;  there 
is  a  purple-brown  spot  at  the  insertion  of  the  columella,  and  a  dis- 
tinct tooth  at  its  base  ;  finally,  there  is  a  nodule-like  thickening  on 
the  last  whorl  at  the  base  of  the  columella.  (v.  Moll.) 


COCHU  >ST  YLA-CHLOR^EA. 

(3)  GKOUP  OF  C.  PARADOX^. 

Bo<ly-wliorl  obtusely  or  not  obviously  keeled,  its  latter  half  some- 
what laterally  compressed  or  flattened  at  the  periphery  ;  descending 
tntr.ird  the  <ij>ertnre;  lip  broad. 

Closely  allied  to  the  group  of  C.  thersites,  but  with  the  last  whorl 
not  acutely  keeled,  less  distorted,  and  descending  in  front. 

1.  Basal  lip  with  a  central  strong  tooth-like  callus  within;  apical 
and  rnliunc/lar  spots  dark;  no  bands.  C.  parodoxa. 

2.  Basal  lip  broad  and  straightened,  without  a  central  tooth  ;  apex 
and  c<>/iimella  white;  body-whorl  2-banded.  C.  constricta. 

3.  Basal  lip  arcuate,  narrower ;  no  color  bands  or  spots. 

C.  undina. 

C.  PARADOX  A  Pfeiffer.     PL  22,  figs.  12,  13,  14. 

I m perforate,  globose-depressed,  solid,  not  obviously  carinated; 
\\v\\t  vellow  with  a  faint  tinge  of  green,  the  apex  blue-black,  a  large 
spot  at  the  insertion  of  the  columella  and  a  small  one  at  the  upper  end 
of  the  lip,  dark  brown;  having  a  callous  thickening  at  the  middle 
of  the  baso-columellar  lip. 

Spire  rather  elevated,  apex  obtuse;  whorls  5,  the  last  somewhat 
deflexed  in  front,  obtusely  carinated  just  above  the  upper  insertion 
of  the  lip,  having  a  broad  shallow  constriction  just  behind  the  lip. 
Surface  lightly  obliquely  striated,  and  showing  under  a  lens 
excessively  fine,  close  spiral  stria?.  Aperture  very  oblique ;  lip 
broadly  reflexed,  white;  baso-columellar  margin  aduate,  thickened 
within,  and  having  a  broad  tooth  or  callus  in  the  middle. 

Alt.  13.],  diam.  23  mill. 

Province  of  A  Ibay,  Luzon,  Philippines. 

II.  i>aradoxa  PFR.,  P.  Z.  S.  1845,  p.  39;  Monogr.  i,  p.  267; 
C..nehyl.  Cub.,  t.  71,  f.  1,  2.— REEVE,  Conch.  Icon.,  f.  1419.— 
I'liiui-i'i,  Abbild.  u.  Beschreib.  i,  p.  153,  t.  4,  f.  7. 

Distinguished  from  C.  constricta  by  the  absence  of  brown  bands, 
the  whorl  less  deflexed  toward  the  aperture,  and  the  more  prominent 
callus  or  wid«-  tooth  in  the  middle  of  the  baso-columellar  lip.  The 
lip  is  more  broadly  reflexed  than  in  C.  undina,  and  the  columella 
-traiirliter,  more  thickened  within. 

Form  immaculata  (pi.  LM,  figs.  27,  28).  Whitish,  without  dark 
Bpota  at  apex  and  eoluinella,  the  aperture  yellow  within. 


COCHLOSTYLA-CHLOR/F.A.  103 

C.  CONSTRICTA  Pfeiffer.     PI.  22,  figs.  16,  17. 

Imperforate,  globose-depressed,  solid  not  carinated  ;  yellow-white, 
the  apex  white,  body-whorl  encircled  by  two  chestnut-brown  zones,  one 
above,  the  other  below  the  rounded  periphery,  the  upper  one  ascend- 
ing the  spire ;  lip  white  or  yellow  within ;  baso-columellar  lip 
straightened,  thickened,  not  toothed  within. 

Spire  rather  elevated,  apex  obtuse ;  whorls  4£,  the  last  deeply  de- 
flexed  toward  the  aperture,  sometimes  a  little  angular  just  above  the 
upper  termination  of  the  lip,  broadly  constricted  behind  the  lip. 
Surface  obliquely  striatulate,  microscopically,  densely,  spirally, 
striated.  Aperture  very  oblique,  oval-triangular  ;  lip  very  broadly 
reflexed ;  baso-columellar  lip  adnate,  wide,  thickened  within,  but 
without  a  callus  or  tooth  in  the  middle. 

Alt.  14*,  diam,  22*,  mill. 

Alt.  12,  diam.  19  mill. 

Mindoro,  Philippines. 

H.  constricta  PFR.,  P.  Z.  S.  1845,  p.  39  ;  Conchyl.  Cab.,  t.  69,  f. 
21,  22  ;  Monogr.  i,  p.  267  (not  H.  constricta  Boub.). — H.  restricta 
PFR.,  Mai.  Bl.  1854,  p.  258;  Monogr.  iv,  p.  228.— IT.  stenopsis 
MOQ.-TAND.,  Rev.  et  Mag.  de  Zool.  1856,  p.  563. —  Chlorcea  constricta 
SEMPER,  Reis.  Phil.  Land  moll,  iii,  p.  228. 

Allied  to  C.  paradoxa,  but  having  the  last  whorl  more  deflexed  in 
front,  and  the  baso-columellar  lip  without  a  central  callous  or 
tooth. 

C.  UNDINA  Pfeiffer.     PI.  22,  figs.  8,  9. 

Imperforate,  globose-turbinate,  thin,  smooth,  shining,  whitish- 
hyaline;  spire  convex-conoid ;.  whorls  5,  a  little  convex,  the  last 
shorter  than  the  spire,  slightly  deflexed  in  front ;  periphery  obsoletely 
angulated,  base  rather  flattened.  Aperture  very  oblique,  truncate- 
elliptical  ;  peristome  thin,  margins  subconniving,  the  right  margin 
expanded,  columellar  lightly  arcuate,  flat,  appressed. 

Alt.  14*,  greater  diam.  23,  lesser  19  mill.     (Pfr.) 

Philippine  Is. 

H.  undina  PFR.,  P.  Z.  S.  1856,  p.  34 ;  Novit.  Conch,  i,  p.  77,  t. 
21,  f.  10-12 ;  Monogr.  iv,  p.  208. 

Columellar  lip  more  arcuate  than  in  the  two  preceding  species. 
The  shell  before  me  is  rather  solid,  and  the  lip  is  broader  than 
shown  in  Pfeiffer's  figures. 


]()4  ( '0( 'II  LOS  1Y  LA-TOR  ASIA. 

(4)    GROUP  OF  C.  THERSITES. 

i;..,lv- whorl  acutely  keeled  in  front,  laterally  compressed  and  dis- 

»  *" 

tnrtfl  "ii  its  latter  part,  not  descending  toward  the  aperture. 

C.  THERSI  riss  Broderip.     PI.  21,  figs.  11,  12,  13,  14. 

1  in pert< .rate,  much  depressed,  •  very  acutely  carinated,  solid; 
whitish-yellow  the  apex  white  or  purple-black,  having  usually  four 
dark-brown  hands.  Latter  third  or  half  of  the  body -whorl  compressed 
nt  tit'  i" 'i-ijihery,  narrowing  and  distorting  the  whorl.  Body-whorl 
not  descending  in  front. 

The  surface  is  finely  microscopically  striate  spirally;  whorls  4f, 
-carroly  convex,  the  last  acutely  keeled,  and  its  latter  portion  much 
compressed,  not  descending.  Aperture  very  oblique,  subtriangular  ; 
peristome  thickened,  white  or  yellowish,  broadly  expanded;  baso- 
columellar  margin  of  the  lip  white,  adnate  to  the  base  nearly  the 
whole  distance  from  axis  to  periphery. 

Alt.  15,  greater  diam.  32,  lesser  23  mill. 

Calapan,  Nan j an  and  Baco,  Mindoro,  Philippines. 

H.  (Carocolla)  thersites  BROD.,  P.  Z.  S.  1841,  p.  381. — PFR., 
Monogr.  i,  p.  299  ;  Conchyl.  Cab.,  p.  235,  t.  112,  f.  3,  4. —  Carocolla 
thersites  Rv.,  Conch.  Syst.  ii,  t.  168,  f.  9. — H.  thersites  Rv.,  Conch. 
Icon.,  f.  IL'I'I. — H.  (Chlorcea)  thersites  HIDALGO,  Obras  Malacol.  i, 
p.  15(J,  t.  i>3,  f.  5-9.  * 

rrhe  laterally  flattened  body-whorl,  not  descending  in  front,  amply 
distinguish  this  from  all  other  forms  of  Chlorcea. 

The  dark  apex  and  stitural  line  are  pretty  constant,  although  one 
specimen  before  me  has  the  apex  white.  The  other  bands  are  very 
variable,  some  or  all  of  them  being  absent  occasionally. 

Section  II.  CORASIA  Albers,  1850. 

'  \ra8ia  A i.i;.,  Die  Heliceen,  1850,  p.  111. — y.  MART,  in  Albers, 
I  >i"  Hel.  -Jd.  edit.,  p.  170.  (Type  H.  virgo}. — PFR.,  Nomencl.  Hel. 
Yiv.,  ]..  111'.). —  <',-ii*t>i11opsis  ANCEY,  Conchol.  Exchange,  ii,  p.  23, 
August,  1887,  (types  //.  Im  uteri,  H.  allisteri.} 

ImperfMnite  (rarely  perforate,)  depressed-globose,  often  carinated  ; 
thin,    Bubdiaphanoua ;    whorls    3-'-5,    rapidly    widening,   the    last 
icarcely    descending;  lip  >li-htly  expanded  or  narrowly   reflex ed. 
Surface   in. ,iv  or    l«-s  striated   spirally,  lucking  opaque  or   "hydro- 
plan,.,       '  1,,'irkinys.     A  natom y  as  in  Cochlostyla  generally. 


COCHLOSTYLA-CORASIA.  105 

Heliciform,  depressed  thin  Cochlostylas,  lacking  hydro phanous 
bands,  and  with  the  expansion  of  the  lip  narrow,  compose  the  section 
Corasia.  They  are  arboreal  in  habits. 

* 

Here  may  be  grouped  all  of  the  species  of  Cochlostyla  ranging 
outside  of  the  Philippine  Islands  ;  including  the  group  of  H.  hunteri, 
tricolor  and  their  allies,  of  the  Solomon  Islands,  a  portion  of  which 
have  hitherto  been  referred  erroneously  to  Geotrochus  by  PfeifFer 
and  other  authors. 

Like  all  of  the  sectional  groups  which  have  been  established  in 
Cochlostyla,  this  section  is  not  well  differentiated  from  several  others. 
Through  the  group  of  C.  regince  a  connection  with  the  section 
ChlorcBa  ( C.  dryope,  paradoxa,  etc.)  is  established  ;  and  again  such 
species  as  C.  intorta  lead  toward  the  section  Callicochlias.  In  a 
general  way,  however,  Chlorcea  differs  from  Corasia  in  its  deflexed 
body-whorl,  wide  lip,  and  the  frequent  presence  of  apical  and  col- 
umellar  dark  spots ;  and  Callicochlias  differs  in  its  more  globose 
form,  wider  lip  and  the  presence  of  "  hydrophanous"  cuticle. 

The  species  may  be  thrown  into  four  groups ; 

1.  Group  of  C.  lactiflua,  Solomon  Is. 

2.  Group  of  C.  extensa,  Moluccas  and  New  Guinea. 

3.  Group  of  C.  regince,  Philippines. 

4.  Group  of  C.  virgo,  Philippines. 


(1)    GROUP  OF  C.  LACTIFLUA. 

Thin  shells  of  glassy  texture,  light  colored,  usually  translucent- 
whitish,  sometimes  varied  with  brown.  Axis  perforate  or  closed ; 
form  varying  from  globose-turbinate  to  depressed,  the  periphery 
carinated  ;  apex  obtuse  ;  whorls  3 J-5.  Surface  generally  showing 
spiral  stride.  Lip  reflexed,  at  least  below.  Distribution,  Solomons 
and  some  adjacent  islands. 

The  shell  characters  given  above  define  a  natural  group,  the 
various  species  of  which  have  heretofore  been  distributed  by  authors 
in  Geolrochus  and  Corasia. 

Mr.  C.  F.  Ancey  has  proposed  as  a  genus  or  subgenus  the  name 
Crystallopsis  for  H.  hunteri  and  H.  a  Ulster  i ;  but  the  characters  he 
assigns  do  not  cover  the  group  as  here  understood. 

C.  HUNTERI  Cox.     PI.  13,  figs.  41,  44,  45. 

Umbilicus  open,  deep,  half-surrounded  by  the  broad  curved  col- 
umellar  lip  ;  globose-depressed,  carinated,  thin,  whitish-translucent. 


KM;  COCH  LOST  YLA-COE  ASIA . 

Surface  nearly  smooth,  lightly  striatulate,  and  showing  fine  im- 
pressed spiral  lines  under  a  lens.  Translucent-whitish,  with  a  nar- 
row opaque-white  band  at  the  subacute  peripheral  keel.  Spire  low  ; 
apex  obtuse;  whorls  4,  slightly  convex,  the  last  not  descending  in 
trout,  convex  below:  aperture  but  little  oblique,  large,  truncate- 
..val;  lip  thin,  expanded,  the  columella  arcuate,  subvertical,  very 
broadly  expanded  and  half  surrounding  the  umbilicus,  but  not 
covering  it,  Alt.  15?,  greater  diam.  21,  lesser  17*  mill. 

Guadalcanar,  Solomon  Islands. 

H.  hunteri  Cox,  P.  Z.  S.  1871,  p.  646,  t.  52,  f.  11.— PFR.,  Monogr. 
vii,  p.  377.— DOHRN  in  Conchyl.  Cab.,  p.  583,  t.  172,  f.  5,  6. 

Differs  from  the  other  species  of  Corasia  wrhich  are  white  and 
subtranslucent,  in  having  the  columellar  lip  produced  in  a  broad 
plate,  somewhat  as  in  H.  grata  Mich. 

( '.  AI.I.ISTERI  Cox.      Unfigured. 

Shell  with  a  deep  narrow  concealed  umbilicus,  semiglobose,  smooth, 
transversely  very  finely  striated  with  lines  of  growth;  the  whorls  of 
the  apex  also  longitudinally  finely  striated,  translucent;  pale  straw- 
colored,  ornamented  above  and  below  with  numerous  rather  narrow 
opaque  white  and  clear  bands  ;  whorls  5,  rapidly  increasing  in  size, 
the  last  much  inflated  and  obtusely  carinated,  not  depressed  in  front, 
convex  above  and  below ;  aperture  ovately  rounded,  subangulated 
at  the  periphery,  showing  the  opaque  and  clear  zones  within ;  per- 
istome  smooth  and  shining ;  lip  very  broadly  expanded  and  reflexed  ; 
margins  approximate,  upper  attached  to  the  carina  of  the  body-whorl, 
columellar  broadly  dilated,  overhanging  and  concealing  the  um- 
bilicus. Diam.,  greatest  0'98,  least  0'64 ;  height  0'60  of  an  inch. 
(  Cox.) 

Solomon  Islands  (Mus.  Cox.) 

This  beautiful  species  may  be  at  once  distinguished  by  the  very 
peculiar  alternate  opaque  white  and  clear  zones.  Amongst  a  large 
iiuiiiln-r  of  specimens  procured,  I  have  not  found  one  to  varv  in  this 
respect.  (Cox.) 

II.  n/listeri  Cox,  P.  Z.  S.  1873,  p.  564. 
II.  vii  i:  \n:\  I-Ynissac.     PI.  13,  figs.  52,  53. 

Shell  covered  perforate,  globose,  thin,  vitraceous,  carinated. 
\\  borla  1  ',-">.  :i  little  convex  ;  aperture  lunate-elliptical;  peristome 
>impl«-,  a  little  reflexed.  (Dh.) 

Australia  / 


COCHLOSTYLA-CORASIA.  107 

H.  vitracea  FEB.,  Prodr.  146;  Hist.,  t.  64,  f.  5. — DH.  in  text  of 
same,  i,  p.  340. — PFR.,  Mon.  Hel.  Viv.  i,  p.  233  (desc.  from  fig.) ; 
Conchyl.  Cab.,  t.  76,  f.  20,  21  (copied  from  Fer.).— Cox,  Mon. 
Austr.  L.  Sh.,  p.  109,  t.  11,  f.  20,  20a,  (copied  from  Fer.) 

Known  solely  by  the  original  figures  in  Ferussac's  Histoire,  which 
are  copied  on  my  plate. 

C.  SUBVITREA  Pfeiffer.     PI.  13,  figs.  54,  55. 

Shell  umbilicate,  trochiform,  rather  solid,  obliquely  distinctly 
striated,  decussated  with  close,  very  fine  spiral  strise,  satiny,  buffish- 
hyaline ;  spire  convex-conic,  rather  obtuse.  Whorls  6,  a  little  con- 
vex, slowly  increasing,  having  an  exserted  cord-like  carina ;  the 
last  whorl  slightly  deflexed  in  front,  more  convex  below  the  median 
carina,  a  little  turgid  in  front,  subcanaliculate  toward  the  narrow 
umbilicus.  Aperture  very  oblique,  subtriangular-lunar ;  peristome 
white,  shining,  the  margins  remote  ;  right  margin  expanded  a  little, 
columellar  margin  substraightened,  outwardly  reflexed,  subdentate 
within,  dilated  in  a  vaulting,  vitreous  plate  at  the  insertion. 

Alt.  15,  greater  diarn.  20*,  lesser  19  mill.     (P/r.) 

Moluccas. 

H.  subvitrea  PFR.,  P.  Z.  S.  1854,  p.  148 ;  Monogr.  iv,  p.  253 ; 
Novit.  Conch,  i,  p.  8,  t.  3,  f.  8,  9. — REEVE,  Conch.  Icon.,  t.  194,  f. 
1361. 

• 

The  locality  should  be  confirmed.  This  shell  has  much  the  aspect 
oflf.  (Satsuma)  japonica,  judging  from  the  figures. 

H.  CYMODOCE  Crosse.     PL  13,  figs.  39,  40. 

Imperforate,  turbinate-trochiform,  rather  thin,  subpellucid,  longitu- 
dinally minutely  striatulate;  dull  yellowish-white;  spire  subconic, 
apex  pale  violaceous,  suture  impressed  ;  whorls  5,  a  little  convex,  the 
last  slightly  descending,  ventricuse,  longer  than  the  spire,  obtusely 
carinated  below  the  middle,  having  a  pale  violaceous  zone  below  the 
carina,  but  little  conspicuous.  Base  moderately  convex,  spirally 
very  delicately  decussated.  Aperture  oblique,  lunate-subquadrate ; 
peristome  milk-white,  margins  distant,  columellar  margin  dilated, 
basal  and  outer  margins  narrowly  reflexed. 

Alt.  17,  greater  diam.  22,  lesser  18  mill.     (Crosse.) 

Habitat  unknown. 

H.  cymodoce  CR.,  Journ.  de  Conchyl.  1869,  p.  184;  1870,  p.  102, 
t.  2,  f.  3. 


COCHLOSTYLA-CORASIA. 

H.  LACTIFLUA  Pfeiffer.     PL  13,  fig.  36. 

Shell  imperforate,  globose-turbinate,  acutely  carinated  at  the 
periphery  ;  thin,  translucent-whitish,  having  a  narrow  opaque  milk- 
white  l)an(l  at  the  carina. 

Surface  smooth,  lightly  marked  by  oblique  growth-lines,  and  under 
a  h-n~  >eon  to  be  finely  spirally  striated,  these  strise  usually  obsolete 
on  t  he  last  whorl.  Apex  obtuse.  Whorls  4-4},  convex,  the  last  not 
descending  in  front,  or  but  slightly  descending.  Aperture  large, 
rounded-rhomboidal,  oblique;  outer  and  basal  margins  of  the  white 
peristome  evenly  reflexed,  becoming  wider  toward  the  base  of  the 
nearly  vertical  columella,  the  face  of  which  is  flattened  and  distinctly 
excavated,  the  columella  is  deeply  inserted  in  the  base,  and  the  axis 
is  narrowly  perforated  in  young  shells. 

Alt.  20,  greater  cliam.  23  mill. 

Alt.  17,  greater  diam.  22  mill. 

New  Georgia  and  Ysabel,  Solomon  Is. 

H.  lictlflaa  PFR.,  P.  Z.  S.  1861,  p.  190 ;  Novit.  Conch.,  p.  175,  t. 
47,  f.  7,  8 ;  Monogr.  v,  p.  270.— H.  isabellensis  Souv.,  Journ.  de 
Conchyl.  1863,  p.  74,  173,  t.  5,  f.  1.— PFR.,  Monogr.  v,  p.  270. 

A  tiirbinate,  glassy-white  shell,  having  a  white  line  at  the  acute 
carina.  The  face  of  the  columella  is  flattened  and  excavated  below. 
There  are  sometimes  inconspicuous  linear  whitish  streaks,  in  the 
direction  of  the  growth-lines. 

I  am  unable  to  separate  H.  isabellensis  (pi.  13,  fig.  37,  38)  from 
laetiflua.  Being  from  a  different  island  (  Ysabel}  it  may  prove  to  be 
a  variety,  but  the  characters  assigned  to  it  afford  no  distinctive 
marks  whatever,  so  far  as  I  can  see. 

H.  PURCHASI  Pfeiffer.     PI.  13,  figs.  46,  47. 

imperforate,  ovate-depressed,  very  thin,  very  closely  plicatulate, 
subdecussated  by  spiral  lines,  pellucid,  whitish  corneous.  Spire 
short-conoid,  obtuse;  suture  impressed,  margined.  Whorls  3*  to  4, 
rapidly  increasing,  convex,  the  last  large,  not  descending  in  front, 
encircled  by  a  milk-white,  acute,  compressed  peripheral  carina; 
beneath  inflated,  the  base  gibbous  ;  aperture  large,  oblique,  angulate- 
lunar;  peristome  thiii,  shortly  expanded,  basal  margin  a  little  re- 
fl''\rd,  columella  arcuate,  compressed,  slightly  calloused. 

Alt.  DI-14,  greater  diam.  22-23,  lesser  17-17}  mill.     (Dohrn.~) 

Admiralty  Islands. 


COCHLOSTYLA-CORASIA.  109 

H.  purchasi  PFR.,  P.  Z.  S.  1858,  p.  21,  t.  40,  f.  4;  Mon.  Hel.  Viv. 
v,  p.  271.— DOHRN  in  Conchyl.  Cab.,  p.  590,  t,  173,  f.  7,  8. 
A  thin,  clear  horn-colored  species. 

C.  ROSSTTERI  Angas.     PI.  21,  fig.  26. 

Shell  imperforate,  globosely  conical,  very  thin,  obliquely  irreg- 
ularly plicately  striated,  with  the  interstices  very  finely  transversely 
striated,  pale  straw-color  encircled  with  numerous  narrow  white 
diaphanous  bands,  the  one  nearest  the  sutures  the  broadest ;  spire 
obtusely  conoidal,  somewhat  flattened  at  the  apex  ;  whorls  four  con- 
vex, the  last  descending,  acutely  keeled  at  the  periphery,  rounded 
at  the  base ;  columellar  margin  arcuate,  narrowly  excavated,  sharp 
within;  aperture  oblique,  rhomboidal;  peristome  not  continuous, 
white,  with  a  purplish  rose-colored  spot  at  the  periphery,  and  another 
at  the  junction  of  the  right  margin  with  the  body-whorl.  Diam. 
maj.  10,  min.  8,  alt.  7!  lines.  (Angas.} 

Ysabel  Island,  Solomon  Group. 

H.  rossiteri  ANG.,  P.  Z.  S.  1869,  p.  46,  t.  2,  f.  5. 

This  species  belongs  to  the  same  group  as  H.  tricolor  Pfr.,  and  H. 
purchasi  Pfr. 

C.  WISEMANI  Brazier.      Un figured. 

Shell  imperforated,  globosely  depressed,  thin,  obliquely  finely 
striated,  shining,  diaphanous,  yellowish-white;  obtuse  at  the  apex; 
whorls  3^,  rather  flat,  rapidly  increasing,  the  last  very  large,  rounded 
at  the  periphery,  marked  with  faint  opaque  white  lines  running  into 
the  suture,  aperture  oblique,  ovately  rounded,  peristome  reflected, 
whitish-brown,  margins  joined  with  thin  callus,  the  right  curved 
down,  columellar  margin  rounded  and  broadly  expanded,  the  upper 
part  of  the  peristome  from  the  centre  takes  a  peculiar  bend  inwards. 
(BrazJ)  Alt.  14,  greater  diam.  28,  lesser  22  mill. 

Solomon  Archipelago. 

H.  (Corasia)  Wisemani  BRAZIER,  Proc.  Linn.  Soc.  N.  S.  Wales, 
i,  p.  8,  1876. 

Described  from  a  single  specimen,  in  the  collection  of  Mr.  Chas. 
Coxen,  of  Brisbane. 

C.  APHRODITE  Pfeiffer.     PL  23,  figs.  8,  9,  10,  11, 12. 

Shell  imperforate,  depressed,  thin,  acutely  keeled  at  the  periphery ; 
translucent-whitish  under  a  thin,  pale  straw-colored  epidermis. 
Surface  shining,  lightly  striatulate  and  having  very  obsolete  close 


COCHLOSTYLA-CORASIA. 

microscopic  spiral  lines,  scarcely  visible  on  the  last  whorl.  Spire 
v«-rv  Low,  apex  flat.  Whorls  3*,  the  last  scarcely  descending  in 
front,  and  encircled  by  an  acute,  compressed  white  keel. 

Aperture  oblique;  lip  white,  narrowly  reflexed  throughout, 
dilated  at  the  columellar  insertion  closing  the  axial  perforation. 

Alt.  17,  greater  diani.  35,  lesser  27?  mill. 

San  Christoval,  Solomon  Is. 

II.  ,,j,hrodite  PFR.,  P.  Z.  S.  1859,  p.  26,  t.  44,  f.  2  ;  Mai.  Bl.  1859, 
p.  L'7  :  Mouogr.  v,  p.  308,  499. 

A  depressed,  light,  ChrocoZws-shaped  shell,  having  much  the 
aspect  of  C.  virgo  of  the  Philippines.  The  thin  epidermis  is 
apparently  caducous  to  a  greater  or  less  degree.  The  whorl  is  con- 
stricted or  grooved  just  behind  the  expanded  upper  lip. 

A  variety  (figs.  11,  12)  is  smaller,  more  elevated,  the  last  whorl 
considerably  deflexed  at  the  aperture. 

Alt.  15i,  greater  diam.  27,  lesser  22*  mill. 

C.  ANADYOMENE  Adams  &  An  gas. 

Shell  orbicular,  depressed-conic,  imperforate,  white,  covered  with 
a  thin,  pellucid,  straw-colored  epidermis.  Whorls  4,  plane,  longitu- 
dinally very  delicately  striated;  the  last  whorl  dilated  in  front, 
constricted  at  the  lip,  at  the  periphery  angulated  and  ornamented 
with  a  conspicuous  filiform  carina.  Aperture  subquadrate,  callus 
of  the  inner  lip  covering  the  umbilical  region  ;  outer  lip  angulated 
in  the  middle,  margin  deeply  reflexed  and  thickened. 

Alt.  1'JA,  diam.  25  mill.     (Ad.  &  Aug.) 

Guadalcanar  and  Ugi,  Solomon  Is. 

H.  anadyomene  A.  &  A.,  P.  Z.  S.  1864,  p.  38.—  PFR.,  Monogr.  v, 
p.  '50s.  —  H.  (Corasia)  anadyomene  SMITH,  P.  Z.  S.  1885,  p.  591. 

Thi<  .species  has  not  been  figured.  It  differs  from  H.  aphroditein 
having  the  whorls  more  convex,  the  keel  and  peristome  not  opaque 
white,  and  the  aperture  less  produced  and  angular. 


<  '.  i-v<  'Hie  Anira^.     IM.  23,  figs.  13,  14. 

Shell  imperforate,  depressed,  thin,  obliquely  irregularly  plicately 
.-triatcd,  pellucid,  white,  covered  with  a  yellowish-brown  shining 
epidermic  spire  x-an-dy  elevated,  obtuse  at  the  apex;  whorls  3, 
flatly  mil  vex,  margined  at  the  sutures,  the  last  not  descending, 
prominently  keeled  at  the  periphery,  flattened  above  and  ventricose 
In-low  ;  eoluniellar  margin  a  rcuately  descending,  narrowly  excavated, 


COCHLOSTYLA-CORASIA.  Ill 

sharp  within  ;  aperture  large,  oblique,  subovate ;  peristome  not  con- 
tinuous, white,  slightly  reflected,  the  upper  margin  nearly  straight, 
not  flexuous.  (Any.) 

Alt.  14,  greater  26,  lesser  18  mill. 

New  Georgia,  Solomon  Is. 

H.  (Corasia) psyche  ANGAS,  P.  Z.  S.  1869,  p.  624,  t.  48,  f.  1. 

This  species,  which  I  have  received  from  Dr.  James  Cox  of 
Sydney,  New  South  Wales,  is  somewhat  intermediate  between  H. 
aphrodite  Pfr.  and  If.  purchasi  Pfr.  It  differs  from  the  former  in 
being  much  flattened  above  and  very  ventricose  below,  and  especi- 
ally in  the  form  and  amplitude  of  the  aperture,  and  from  the  latter 
in  texture,  form  of  spire,  color,  and  size.  (Ang.) 

C.  BALCOMBEI  Cox.       PI.  23,  fig.  15. 

Shell  imperforate,  depressed,  almost  flat  above,  convex  below,  very 
thin,  glossy,  pale  straw-colored,  coarsely  striated  above  and  below 
with  regular  oblique  transverse  stria?,  and  longitudinally  with 
strongly  marked  very  slightly  waved  striae,  giving  by  their  decussa- 
tion  a  granular  appearance  to  the  surface  of  the  shell ;  spire  scarcely 
raised,  suture  impressed;  whorls  4,  slightly  convex  above,  rapidly 
increasing  in  size ;  last  whorl  inflated  below,  and  very  prominently 
and  sharply  keeled,  the  keel  edged  with  an  opaque  white  line;  base 
rounded ;  aperture  large,  subtrapezoidal,  acutely  angled  at  the 
periphery  ;  peristome  slightly  thickened  and  everted,  upper  margin 
attached  to  the  carina  of  the  last  whorl ;  columella  dilated,  appressed, 
and  indented  over  the  umbilical  region.  (Cox.) 

Alt.  O48,  greater  diam.  1'09,  lesser  0'7S  inch. 

Solomon  Is. 

H.  (Corasia)  balcombei  Cox,  P.  Z.  S.  1873,  p.  565,  t.  48,  f.  4. 

C.  TRICOLOR  Pfeiffer.     PL  23,  figs.  1,  2,  3,  4,  5,  6. 

Shell  imperforate,  depressed,  thin  but  rather  solid,  acutely  keeled 
at  the  circumference;  translucent-whitish,  having  numerous  close 
spiral  lines  of  opaque  white  above  and  below,  the  keel  opaque  white 
bordered  with  lines  of  brown  and  articulated  with  brown  dots  ;  lip 
brown. 

Surface  lightly  obliquely  striatulate,  conspicuously  closely  and 
finely  spirally  striated.  Spire  low,  the  apex  flattened.  Whorls  4, 
the  last  wide,  a  little  deflexed  in  front ;  strongly  contracted  toward 
the  aperture  above  and  at  the  carina ;  base  more  or  less  swollen  or 
gibbous  around  the  columella.  Aperture  oblique,  triangular;  per- 


H  2  <  -OCHLOSTYLA-CORASIA. 

istome  edged  with  brown,  the  upper  margin  not  expanded,  basal 
mar-in  narrowly  expanded,  columellar  margin  wide,  flat,  brown, 
thickened  within  by  a  wide  callus  which  is  more  or  less  abruptly 
truncated  below. 

Alt.  14,  greater  diam.  30,  lesser  24*  mill. 

San  Christoval,  Ugi  and  Santa  Anna,  Solomon  Is. 

11.  tricolor  PFR.,  P.  Z.  S.  1849,  p.  129  ;  Conchyl.  Cab.,  t.  124,  f. 
'2-2-24:  Monogr.  iii,  p.  168;  I.  c.  iv,  p.  193.— KEEVE,  Conch.  Icon., 
f.  431.— E.  A.  SMITH,  P.  Z.  S.  1885,  p.  589,  (var.  picta.') 

This  is  a  variable  shell  both  in  form  and  coloring.  The  outline 
seen  from  above  or  below  is  often  more  oblong  than  the  figures 
show.  The  suture  is  white-margined,  sometimes  articulated  with 
brown.  The  circum-colurnellar  swelling  is  often  strongly  developed, 
often  obsolete. 

Color  var.  White  spiral  lines  inconspicuous ;  carina  white,  without 
brown  border  or  spots.  Smaller  than  the  type.  Diam.  25-27  mill. 

Santa  Anna;  San  Christoval. 

Color  var.  Similar  to  the  last,  but  the  white  peripheral  stripe 
bordered  with  brown,  not  articulated. 

San  Christoval. 

Cnlnr  I'm:  PICTA  Smith,  PL  18,  figs.  69-71. 

Undulating  reddish-brown  stripes  above  and  below,  distinctly 
visible  within  the  aperture. 

Northern  coast  of  San  Christoval. 

Var.  TRANSENNA  Pilsbry.      PL  23,  figs.  16,  17,  18,  and  fig.  between 

'1  and  5. 

Shell  im perforate,  depressed,  low-conic  above,  convex  beneath, 
a«-ut«'lv  keeled  at  the  periphery;  thin;  translucent-whitish,  having 
IIIIIIKTOU-  spiral  bands  and  lines  of  opaque- white ;  keel  white, 
bordered  above  and  below  with  irregularly  clouded  brown  stripes. 

Surface  having  liirht  growth-lines,  decussated  by  conspicuous  and 
<•!«,-<•  spiral  btriae.  Spire  conic,  apex  obtuse.  Whorls  4,  the  last  not 
•  I'-tli-M-d  in  front,  its  periphery  slightly  contracted  at  the  aperture; 
ba->'  convex,  not  gibbous  around  the  columella.  Aperture  oblique, 
subtriangular,  the  peristome  brown-edged,  its  upper  margin  not  ex- 
pamli-il  ;  basal  margin  expanded,  columella  dilated,  its  face  flattened 
and  bavin-  a  .-hallow  vertical  groove  or  excavation,  its  inner  edge 


COCHLOSTYLA-CORASIA.  113 

steeply  obliquely  descending,  straight,  forming  an  angle  at  its  junc- 
tion with  the  basal  margin. 

Alt.  14J,  greater  diam.  25 5,  lesser  20 J  mill. 

Solomon  Is. 

This  form  is  smaller  and  more  elevated  than  C.  tricolor,  the  col- 
umella  is  very  much  slenderer,  much  less  thickened,  and  mostly 
white.  The  structure  of  the  columella  is  so  different  that  I  am 
inclined  to  believe  the  shell  specifically  distinct.  The  young  have 
no  brown  markings  save  a  small  eolumellar  spot. 

(2)    GROUP  OF  C.  EXTENSA. 

Part  of  the  following  species  have  been  included  by  authors  in 
Albwsia ;  but  their  characters  seem  more  in  accord  with  Corasia  than 
with  the  other  group.  Distribution,  Moluccas  and  New  Guinea. 

C.  CONFORMIS  Ferussac.     PI.  19,  figs.  4,  5,  3. 

Shell  imperforate,  globose-depressed,  carinated,  thin,  obliquely 
closely  striated,  diaphanous,  whitish,  covered  with  a  pale  brownish 
epidermis ;  spire  little  elevated,  apex  very  obtuse  ;  suture  impressed  ; 
whorls  3J,  rapidly  increasing,  slightly  convex,  the  last  large,  not 
descending,  convex  beneath.  Columella  subvertical,  compressed, 
subexcavated ;  aperture  ample,  oblique,  lunar-subrotund,  having  a 
pearly  luster  inside;  peristome  simple,  shortly  reflexed. 

Alt.  20,  greater  diam.  34,  lesser  28  mill.     (Pfr.) 

New  Guinea. 

H.  conformis  FER.,  Prodr.,  p.  33  ;  Histoire,  t.  25a,  f.  10. — FER.  in 
QUOY  &  GAIMARD,  (Freycinet)  Voy.  autour  du  Monde,  1'Uranie  et 
la  Physicienne,  Zool.,  p.  467;  Atlas,  t.  67,  f.  4,  5. — WOOD,  Index 
Test.,  p.  22,  t.  7,  f.  37.— PFR.,  Conchyl.  Cab.,  p.  42,  t.  72,  f.  1-3 ; 
Monogr.  i,  p.  293  ;  iii,  p.  203  ;  vii,  p.  355  ;  Zeitschr.  f.  Mai.  1849,  p. 
76. — RVE.,  Couch.  Icon.,  f.  497. — MARTENS,  Ostas.  Landsch.,  p. 
393. —  Corasia  conformis  WALLACE,  P.  Z.  S.  1865,  p.  412. — H. 
(Corasia)  conformis  ALB.,  Die  Hel.,p.  HI.--TAPPARONE  CANEFRI, 
Ann.  Mus.  Civ.  Genov.  xix,  p.  158,  t.  8,  f.  1,  8,  15  (Anatomy.) 

C.  LEUCOPHTHALMA  Pfeiffer.     PI.  26,  figs.  16,  17. 

Imperforate,  subconoid-depressed,  carinated,  thin,  obliquely 
striated  and  under  a  lens  most  delicately  granulated  ;  spire  white, 
shortly  conoid-elevated,  apex  obtuse ;  suture  little  impressed,  indis- 
tinctly and  delicately  white-edged.  Whorls  nearly  4,  little  convex, 

rapidly  increasing,  the  last  not  descending,  subacutely  carinaled, 
8 


1  1  4  COCHLOSTYLA-CORASIA. 

(the  r:i ri mi  Incoming  more  obtuse)  ;  covered  with  a  thin,  tawny 
cuticle,  irregularly  streaked  and  four-banded  with  brown  ;  base  more 
convex  ;  columella  little  oblique,  compressed,  subexcavated  ;  aper- 
ture oblique,  wide  lunar,  the  right  side  obsoletely  angulated,  inside 
bluish-pearly,  shining ;  peristome  simple,  narrowly  reflexed,  upper 
margin  curving  forward,  basal  margin  forming  an  obtuse  angle  with 
the  columella. 

Alt.  21-22,  greater  diam.  42,  lesser  34  mill.     (Jjfr.) 

Celebes  f 

If.  leucophthalma  PFR.,  Mai.  Bl.  xvii,  1870,  p.  93  ;  Novit.  Conch, 
iv,  p.  10,  t.  Ill,  f.  8,  9 ;  Monogr.  vii,  p.  335. 

H.  EXTKXSA  Miiller.     PL  27,  figs.  9,  10,  11. 

Shell  imperforate,  globose-depressed,  rounded  at  the  periphery, 
thin  but  rather  solid;  white  under  a  fugacious  yellowish  epidermis; 
Hirface  shining,  very  lightly  striatulate,  having  faint,  almost  obsolete 
spiral  stride,  seen  only  under  a  strong  lens.  Spire  low,  apex  obtuse  ; 
whorls  4,  the  last  globose,  a  little  descending  in  front.  Aperture 
large,  oblique ;  peristome  narrowly  expanded  and  subreflexed 
throughout;  the  columella  steeply  sloping,  straight,  somewhat 
thickened,  forming  a  decided  and  conspicuous  angle  at  its  junction 
with  the  basal  margin. 

Alt.  21,  greater  diam.  27?,  lesser  22*  mill. 

Alt.  19,  greater  diam.  26,  lesser  21  mill. 

Island  of  Goram.     (Wallace.) 

If.  extensa  O.  FR.  MULL.,  Hist.  Verm.,  p.  60. — DESH.  in  FER., 
Hist.,  t.  96,  f.  5-7.— PFR.,  Monogr.  iii,  p.  193;  Conchyl.  Cab.,  p. 
193,  t.  1(50,  f.  6,  7. — MARTENS,  Ostas.  ZooL,  Landschn.,  p.  331. 

A  larger  shell  than  H.  najas,  more  flattened  above,  and  with  the 
columella  more  thickened,  straighter  and  longer,  forming  a  decided 
airj-lc  with  the  basal  margin. 

Tin-  axis  is  perforated  in  young  shells,  although  from  the  appear- 
auci-  of  adults  one  would  not  suspect  this  to  be  the  case. 

htrt  of  the  specimens  before  me  are  marked  "  New  Ireland,"  I 
know  not  on  whose  authority. 

C,  •  CELAXIS  I'ilsbry.     PL  26,  figs.  6,  7,  8. 

-li''ll  imperforate  but  having  a  hollow  axis,  globose-depressed,  thin, 

liu'ht,  covered  with  a  thin  cuticle  of  a  very  delicate  greenish-yellow 
tint,  becoming  white  on  the  spire;  the  suture  bordered  below  by  a 


COCHLOSTYLA-CORASIA.  115 

distinct  buff-white  margin  about  1  mill,  wide  on  the  last  whorl. 
Surface  nearly  lusterless,  delicately  obliquely  striate,  decussated  by 
obsolete  spiral  striae.  Spire  low,  apex  obtuse.  Whorls  4,  the  last 
slowly  descending  in  front,  rounded  at  the  periphery,  a  little  concave 
in  the  center  beneath.  Aperture  large,  oblique  ;  upper  margin  of 
the  peristome  slightly  expanded,  basal  subreflexed,  the  columellar 
margin  reflexed  and  closely  adnate,  closing  the  umbilicus  by  a  thin, 
convex,  dilated  plate.  Inner  edge  of  the  columella  concave,  not 
angled  where  it  passes  into  the  basal  margin. 

Alt.  20,  greater  diam.  23,  lesser  20  J  mill. 

Java.     (A.  D.  Brown  coll.) 

This  species  differs  from  its  allies  in  having  the  axis  perforated, 
the  perforation  closed  in  the  adult  by  a  thin  convex  plate  (fig.  8). 
It  is  described  from  the  collection  of  the  late  A.  D.  Brown,  now  in 
the  Academy  Museum. 

H.  NAJAS  Pfeiffer.     PI.  27,  fig.  13  (var.  fig.  12)  ;  pi.  28,  figs.  31,  32. 

Shell  depressed -globose,  thin,  regularly  lightly  sculptured  with 
close  striae;  yellowish-white;  spire  short,  obtuse;  whorls  4,  a  little 
convex,  rapidly  increasing,  the  last  inflated,  somewhat  descending 
in  front ;  aperture  oblique,  large,  subrotund ;  peristome  thin,  nar- 
rowly expanded  throughout,  the  upper  margin  arcuate,  columellar 
margin  subvertical,  lightly  thickened,  (r.  Mart.) 

Alt.  18  i,  greater  diam.  25,  lesser  19  mill. 

Alt.  22,  greater  diam.  31,  lesser  23  mill. 

Amberbaki,  New  Guinea;  Islands  of  Burn  and  Goram,  Moluccas. 

H.   najas   PFR.,  Symbolse  iii,  p.  71,  1846;  Monogr.  i,    p.  261; 
Conchyl.  Cab.,  t.  53,  f.  12,  13.— PHILIPPI,  Abbild.  etc.,  ii,  t.  9,  f.  5. 
— MARTENS,  Preuss.    Exped.    Laudschn.,  p.   330,   t.    18,   f.    4.- 
TAPPARONE  CANEFRI,  Ann.  Mus.  Civ.  di  Stor.  Nat.  di  Genov.  xix, 

p.  187,  1883. 

A  thin,  shining  species,  allied  to  H.  extensa,  H.  obliquata  and  H. 
physalis.  Numerous  specimens  are  before  me,  showing  considerable 
variation  in  contour  both  in  the  direction  of  H.  physalis  and  H. 
obliquata.  These  three  species  may  perhaps  prove  to  be  connected 
by  intermediate  forms. 

C.  PHYSALIS  Pfeiffer.     PI.  27,  figs.  2,  3. 

Imperforate,  globose-depressed,  thin,  striatulate  and  under  a  lens 
seen  to  be  minutely  punctulate  somewhat  shining,  translucent,  very 
pale  greenish-white.  Spire  very  shortly  elevated,  convex,  obtuse ; 


1  1  (I  COCHLOSTYLA-CORASIA. 


liirht,  impressedly-margined.  Whorls  4,  a  little  convex, 
rapidly  increasing,  the  last  depressed-globose,  visibly  a  little  descend- 
ing in  front;  columella  white,  straightened,  oblique,  compressed, 
excavated  above  ;  aperture  oblique,  lunate-subcircular,  with  a  pearly 
luster  inside;  peristome  thin,  margins  converging,  the  upper  margin 
curving  forward,  narrowly  expanded,  basal  margin  reflexed,  forming 
an  obsolete  angle  with  the  columella. 

O 

A  It,  16,  greater  diam.  27,  lesser  22  mill.     (Pfr,) 

Tular  \_Talaur  /]  Islands. 

H.pliysaUs  PFR.,  Mai.  Bl.  xviii,  1871,  p.  123;  Novit.  Conch,  iv, 
p.  73,  t.  121,  f.  10,  11  ;  Monogr.  vii,  no.  2316. 

Allied  to  H.  najas,  obliquata  and  extensa  but  much  more  depressed, 
the  last  whorl  less  inflated,  little  descending,  columella  much  slop- 
ing, forming  a  very  obsolete  angle  with  the  basal  margin. 


H.  OBLKJUATA  Deshayes.     PI.  27,  figs.  14,  15. 

Shell  subglobose,  pure  white,  delicately  striate  ;  spire  short,  obtuse  ; 
whorls  4,  nearly  flat,  the  last  large,  inflated,  obliquely  deflexed  to- 
ward the  aperture;  base  imperforate.  Aperture  ovate-lunar,  dilated 
in  the  middle,  coarctate  posteriorly  ;  peristome  margined,  narrowly 
reflexed,  base  covered  with  a  short  callus. 

Alt.  22,  diam.  27  mill.     (Desk.} 

Habitat  unknown. 

H.  obliquata  DH.  in  Fer.,  Hist,  i,  p.  219,  t.  28a,  f.  3,  4.—  PFR., 
Monogr.  iii,  p.  192. 

(3)    GROUP  OF  C.  REGINJE. 

Acutely  carinated  forms,  flattened  above,  of  a  more  or  less  pro- 
nounced green  tint;  lip  white;  surface  seen  under  a  lens  to  be  dis- 
tinctly spirally  striated,  often  obliquely  corrugated. 

C.  KI:<.INM;  Broderip.     PI.  25,  figs.  45,  46. 

Impi  i  •!•»  rate,  depressed,  piano-con  vex  above,  convex  below;  thin;  of 
a  bright  bluish-green  color,  the  apical  H  whorls  and  the  lip  white. 
Surface  delicately  obliquely  striatulate,  obsoletely,  very  finely  spirally 
.-triated.  Whorls  3-],  the  last  acutely  keeled,  slightly  concave  above 
and  below  the  keel,  not  descending  in  front,  tumid  beneath.  Aper- 
ture subtrian^uliir-lunar;  peristome  thin,  outer  margin  expanded, 
baso-columellar  margin  adnate,  slightly  thickened  within,  dilated 
ov<-r  the  umbilicus.  Alt.  11,  greater  diam.  28,  lesser  22  mill. 


COCHLOSTYLA-CORASIA.  117 

Is.  Catanduanes  and  Potillo,  off  E.  coast  of  Luzon  (Moll.)  ;  Baler 
Casiguran  and  Cordillera,  Ambubuk,  on  the  E.  coast  of  N.  Luzon 
(Semper);  Albay,  Luzon;  also  Sibonga,  Zebu?  (Cuming.) 

H.  regince  BROD.,  P.  Z.  S.  1841,  p.  36.— PFR.  in  Conchyl.  Cab.,  t. 
68,  f.  8-11 ;  Monogr.  i,  p.  299.— REEVE,  Conch.  Icon.,  f.  125.- 
Caracolla  regince  E.V.,  Conch.  Syst,  ii,  t.  167,  f.  2. —  Cochlostyla 
regince  MARTENS  in  Mai.  Bl.  xv,  p.  1 62, 1868. —  Cochlostyla  (  Corasia) 
regince  SEMPER,  Phil.  Archip.  Landmoll.  iii,  p.  168. — v.  MOLL.,  Ber. 
Senck.  Nat.  Ges.  1890,  p.  229. — Helix  (Chlorcea)  regince  HIDALGO, 
Obras  MalacoL,  p.  153,  t.  24,  f.  3,  4. — H.  smaragdina  GRAT.,  Act. 
Soc.  Bord.  xi,  p.  405,  t.  3,  f.  26,  27.  (Young.) 

The  bright  bluish-green  hue  of  this  species  distinguishes  it  for 
beauty,  even  in  this  genus  where  vividly  colored  shells  are  the  rule. 
The  typical  regince  has  the  apical  whorls  white,  and  the  periphery 
has  no  bands.  I  am  disposed  to  follow  Hidalgo  in  considering  the 
following  two  forms  geographic  varieties  rather  than  distinct  species. 

Var.  ALMS  v.  Moll.     PI.  25,  figs.  42,  43,  44. 

Spire  flatter,  the  apex  purple-brown ;  last  whorl  encircled  by 
a  brown  girdle,  and  descending  a  little  in  front, 

Alt.  11*,  greater  diam.  30  mill. 

Algeria  and  Boljoon,  Zebu;  Puerto  Galera,  Mindoro. 

H.  regince  var.  b,  PFR.,  Monogr.  i,  p.  299. — ?  FER.,  Hist.,  1. 102,  f. 
15,  16. —  Cochlostyla  almce  v.  MOLL.,  Landschu.  Fauna  Cebu,  p.  227, 
t.  8,  f.  5. — H.  ( Chlorcea)  regince  var.  almce  HIDALGO,  Obras 
Malacol.  i,  p.  153. 

This  variety  forms  a  transition  toward  certain  forms  of  C.  dryope. 

Var.  ELIZABETHS  O.  Semper.     PI.  25,  figs.  47,  48. 

Shell  larger,  the  carina  more  acute,  and  the  spire  flat. 
Alt.  14,  diam.  38  mill. 

Island  of  Calayan,  off  Northern  Luzon. 

Cochlostyla  (Corasia)  elizabethce  O.  SEMPER,  Journ.  de  Conchyl. 
1866,  p.  261,  t.  8,  f.  2. — C.  SEMPER,  Phil.  Archip.  Landmoll.  iii,  p. 
168. — H.  elizabethce  PFEIFFER,  Monogr.  v,  p.  307. — H.  (Chlorcea) 
regince  var.  elizabethce  HIDALGO,  Obras  Malacol.  i,  p.  154. 

C.  PAPYRACEA  Broderip.     PI.  24,  figs.  34,  35,  36. 

Imperforate,  convexly  flattened  above,  acutely  carinated  at  the 
periphery;  thin,  white,  under  a  very  light  greenish-buff  tinted  cuti- 
cle, having  a  small  brown  dot  at  the  apex.  Surface  delicately 


COCHLOSTYLA-CORASIA. 

striated,  decussated  by  spiral  striae,  some  of  which  are 
ratlu-r  prominent,  readily  seen  without  a  lens.  When  magnified 
the  surface  is  seen  to  be  finely,  obliquely  corrugated  in  a  direction 
at  riirht  un.des  to  the  lines  of  growth.  Whorls  4,  the  last  somewhat 
concave  immediately  above  and  below  the  keel,  not  descending  in 
front;  columella  surrounded  by  a  conspicuous  swelling  or  ridge. 
Lip  white,  narrowly  expanded,  columellar  margin  dilated,  arcuately 
>l..pincr.  Alt.  13-1,  diam.  26J,  lesser  21  mill. 

Mindoro,  Philippines. 

Carocolla  papyracea  BROD.,  P.  Z.  S.  1841,  p.  36.— Helix  papyra- 
cea  PFR.  Symb.  iii,  p.  70;  Conchyl.  Cab.,  t.  52,  f.  11,  12  ;  Monogr. 
i,  p.  218. — REEVE,  Conch.  Icon.,  f.  90. — H.  (Corasia)  papyracea 
HIDALGO,  Obras  Malacol. i,  p.  147,  t.  21,  f.  1,  2—Cochlostyla  (Cora- 
sia') papyracea  SEMPER  Phil.  Archip.  Landmoll.,  iii,  p.  168. — MOLL., 
Mai.  Bl.  (new  ser.)  x,  p.  160  ;  Landschn.  Fauna  Cebu,  p.  229  (var. 
dilatatd) — ?  H.  acutangula  BURROW,  Elem.  Conch.,  t.  26,  f.  1. 

This  is  like  C.  regime  in  form,  but  less  depressed  and  more 
swollen  around  the  columella ;  the  color  is  a  very  light  tint  of  buff, 
faintly  tinged  with  green. 

Var.  DILATATA  v.  Moll.     Larger  and  flatter  than  typical  papyracea, 
the  keel  more  produced,  last  whorl  more  widened.    Alt.  14,  greater 

diam.  31  mill. 

Zebu,  Philippines. 

('.  i-MTT.u'iNA  Deshayes.     PI.  25,  figs.  49,  50,  51. 

Im perforate,  orbiculate-depressed,  thin,  obliquely  closely  striate 
and  decussated  with  spiral  stria?  visible  under  a  lens,  subpellucid, 
shining;  greenish-buff  above  and  below;  the  middle  of  the  last 
whorl  vivid  blue ;  spire  convex,  obtuse ;  suture  slightly  impressed ; 
whorls  4-2,  slightly  convex,  the  last  more  convex,  subangular  at  the 
periphery,  not  descending  in  front.  Aperture  oblique,  subtrigonal- 
clliptiral ;  peristome  simple,  thin,  margins  remote,  the  right  margin 
expanded,  rolium-llar  margin  arcuately  sloping,  compressed,  dilated, 
adnatc,  whitish.  Alt.  15,  greater  diam.  25,  lesser  19  mill. 

Cordillera  of  Palanan,  Northern  Luzon. 

If.  i^lf/'iriitd  Dir.,  Journ.  de  Conchyl.  1861,  p.  350,  t.  16,  f.  3-5. 
I'M:.,  Monoirr.  v,  p.  285. —  Cochlostyla  (Corasia)  psittacina  SEMPER, 
Phil.  Ai-chip.  Landmoll.  iii,  p.  172  (with  var.) 

Allied  to  tin-  L^roup  of  C.  regince  by  coloring  and  general  form, 
luit  dim-ring  in  the  mure  globose  and  obtusely  keeled  body-whorl. 


COCHLOST  YLA-CORASIA.  119 

(4)    GROUP    OF    C.    VIRGO. 

Globose-depressed  shells,  varying  from  acutely  carinatecl  to  ob- 
tusely subangular  or  rounded ;  thin,  very  light  colored  ;  surface 
showing  more  or  less  obsolete  spiral  lines  on  the  earlier  whorls,  not 
distinct  on  the  last ;  not  microscopically  obliquely  wrinkled  or  cor- 
rugated. Whitish  (or  varying  toward  greenish  or  yellowish),  uni- 
colored  or  1,  2  or  3-banded. 

Includes  most  of  the  species  of  the  Philippine  Islands. 

C.  VIRGO  Broderip.     PL  24,  figs.  29,  30. 

Imperforate,  depressed,  low-convex  above,  convex  beneath,  acutely 
keeled  at  the  circumference  ;  thin,  entirely  white  (covered  with  a  thin 
yellowish  cuticle  when  fresh),  the  lip  narrowly  edged  with  dark 
brown.  Surface  shining,  smooth,  obliquely  striatulate,  showing 
under  a  strong  lens  almost  obsolete  traces  of  spiral  lines  on  the 
earlier  whorls.  Whorls  4.  Aperture  oblique  ;  lip  very  narrowly 
expanded,  browrn-edged  ;  columellar  lip  widened,  flat,  nearly  straight, 
white.  Alt.  18,  greater  diam.  32,  lesser  27  mill. 

Is.  of  Negros ;    Western  Zebu;  Zamboanga,  Mindanao. 

Carocolla  virgo  BROD.,  P.  Z.  S.  1841,  p.  44. — KEEVE,  Conch. 
Syst.  ii,  t.  168,  f.  13.— H.  virgo  PFR.  in  Conchyl.  Cab.,  t.  68,  f.  1,  2 ; 
Monogr.  i,  p.  217. — Rv.,  Conch.  Icon.,  f.  89. — DESK,  in  Fer.,  Hist, 
t.  101,  f.  13-15. — H.  (Corasia)  virgo  HIDALGO,  Obras  Malacal.  i, 
p.  147,  t.  22,  f.  1. —  Cochlostyla  (  Corasia)  virgo  SEMPER,  Phil.  Archip. 
Land  Moll.,  iii,  p.  166. — v.  MOLL.,  Landschn.  Fauna  Cebu,  p.  229. 

This  pure  white  shell  with  narrowrly  brown-bordered  outer  lip  is 
quite  distinct  from  other  earinated  species.  The  surface  is  almost 
free  from  spiral  sculpture.  The  spire  is  sometimes  much  depressed. 

C.  DEALBATA  Broderip.     PI.  21,  fig.  36. 

Shell  dull  white,  slightly  yellowish,  subdiaphanous,  subglobose, 
substriated  with  very  close  oblique  growth-lines ;  slightly  subum- 
bilicated  ;  aperture  auriculate,  large,  lip-edge  slightly  substriated. 

Alt.  21,  diam.  37*  mill.     (Brod.) 

Island  of  Siquijor,  Philippines. 

H.  (Carocolla)  dealbata  BROD.,  P.  Z.  S.  1841,  p.  45  (not  Helix 
dealbata  LOWE,  1830). — H.  Broderipi  REEVE,  Conch.  Icon.,  f.  88, 
(not  H.  broderipi  Pfr.) 


]  0|  i  (  ( )(  II  LOS TYLA-CORASIA. 

Hidalgo  has  already  (Obras  i,  p.  147)  raised  the  question  of  the 
identity  of  the  H.  broderipi  of  Pfr.  and  H.  broderipi  Keeve.  They 
seem  to  me  perfectly  distinct;  and  I  have,  therefore,  restored  the 
•  .riii-iiia!  name  given  by  Broderip  to  the  form  figured  in  the  Iconica, 
which  represents  Broderip's  species.  The  name  is  not  preoccupied 
in  the  genus  ( 'ochlostyla. 
C.  CASTA  Pfeiffer.  PL  25,  figs.  37,  38. 

Imperforate,  depressed,  about  equally  convex  above  and  below, 
car  mated,  striatulate  and  obsoletely  malleated,  shining,  white  under 
a  fim-aremis  pale  yellowish  epidermis;  suture  linear,  chalky-white; 
whorls  4,  nearly  fiat,  the  last  obsoletely  angled  at  the  suture  and 
below  tin-  carina  ;  eolumella  short,  sloping,  excavated ;  base  sub- 
tortuous  ;  aperture  subtrapezoidal ;  peristome  expanded,  white, 
basal  margin  slightly  arcuate,  forming  an  angle  with  the  eolumella. 

Alt.  '2:},  greater  diam.  47,  lesser  38  mill.     (P/r.) 

Philippines. 

H.  casta  PFR.,  P.  Z.  S.  1848,  p.  110  ;  Conchyl.  Cab.  t.  123,  f.  1, 
L> ;  Monogr.  iii,  p.  206. — REEVE,  Conch.  Icon.  f.  86. 

A  large  form,  not  yet  found  by  recent  collectors.  The  entire 
surface  of  the  shell  is  superficially  indented  or  malleated. 

C.  FUELLA  Broderip.     PI.  24,  figs.22,  23,  25 ;  pi.  25;  fig.  39. 

Imperforate,  globose-depressed,  carinated  at  the  periphery,  thin, 
white  under  a  very  thin,  deciduous  yellowish  cuticle,  having  a  brown 
band  at  the  peripheral  angle,  becoming  evanescent  on  the  earlier 
part  nf  the  body-whorl.  Surface  smooth,  very  delicately  striated, 
and  showing  under  a  lens  fine  spiral  lines  on  the  earlier  whorls. 
future  linear,  white.  Whorls  4,  nearly  flat,  the  last  carinated, 
descending  a  little  below  the  carina  at  the  aperture.  Aperture 
ol>li(|iie.  subtrigonal-lunar ;  peristome  thin,  narrowly  expanded; 
eolumella  wide,  fiat,  its  inner  edge  straight,  oblique,  forming  an 
with  the  basal  margin. 

Alt.  lir.,  greater  diam.  31,  lesser  26  mill. 

I. -<l<i nil  of  Camiguing ;    Lianga,   Rio  Salos,  Rio  Jibon,  Pasian, 
),  Rio  Af/tixnn,  Itlnnd  of  Mindanao,  Philippines. 

puella  BROD.,  P.  Z.  S.  1841,  p.  45. — H.  puella  PFR. 
Symb.  iii,  p.  74  ;  Conchyl.  Cab.  t.  68,  f.  3-5  ;  Monogr.  i,  p.  298.- 
REEVE,  Conch,  Icon.  f.  (;<;.-  //.  (Corasia)  puella  HIDALGO,  Obras 
Midarol.  i,  p.  148,  t.  21,  f.  4. —  Cochlottyla  ( Corona)  puella  SEMPER, 
K'-i-«-ii  in  Archip.  Phil.,  Landmoll.  iii,  p.  167. 


COCHLOSTYLA-CORASIA.  121 

Allied  to  C.  virgo,  but  the  carinaismore  obtuse  and  the  lip  white. 
Specimens  from  Mindanao  are  more  depressed,  and  sometimes  larger 
than  the  types,  measuring  over  40  mill.  diam.  according  to  Hidalgo. 
See  pi.  25,  fig.  39 

Var.  LAIS  Pfr.     PI.  25,  fig.  40. 

Similar  to  C.  puella  but  more  globose,  the  peripheral  angle  more 
obtuse ;  columella  more  arcuate  at  base,  not  forming  so  distinct  an 
angle  with  the  basal  margin  as  C.  puella.  Peripheral  band  more 
prominent. 

Min  danao,  Philippines. 

H.  Ms  PFR.,  P.  Z.  S.  1853,  p.  49  ;  Monogr.  iii,  p.  647.— KVE.  f. 
1016. — HIDALGO,  Obras  Malacol.  i.  p.  148,  t.  21,  f.  5. 

Figure  40  represents  the  type.  Hidalgo  believes  lais  to  be  a  vari- 
ety of  puella. 

Var.  SUBPUELLA  Pilsbry.     PI.  24,  figs.  26,  19,  20,  21,  24. 

Thinner  than  puella ;  multilineate  with  pale  yellow  on  a  white 
ground-color,  the  periphery  encircled  by  a  dark  chestnut  band 
which  continues  up  the  spire,  forming  a  narrow  border  above  the 
suture  to  the  apex. 

Basilan,  Philippines  (P.  Acad  coll.)  Tukan  Bessi  Id.  (Pfr.) 

H.  lais  var.  PFR.  in  Novit.  Conch,  iv,  p.  114,  t.  126,  f.  6,  7. 

Many  specimens  of  this  variety  are  before  me,  showing  a  remark- 
ably constant  color-pattern. 

C.  IROSINENSIS  Hidalgo.     PI.  28,  figs.  29,  30. 

Imperforate,  depressed,  about  equally  convex  above  and  below, 
the  young  carinated,  adults  angulated,  the  angle  gradually  dis- 
appearing ;  thin,  subpellucid,  little  shining,  smooth ;  (under  a  lens 
seem  to  be  decussated  with  very  subtle,  inconspicuous  spiral  and 
oblique  striae)  ;  unicolored  whitish  under  a  pale  buffish  cuticle ;  apex 
obtuse ;  suture  simple ;  whorls  4,  slightly  convex,  rapidly  widening, 
the  last  dilated  in  front,  not  descending,  somewhat  turgid  at  the  col- 
umella. Aperture  trapezoidal ;  peristome  thin  or  sublabiate,  the 
upper  margin  simple,  basal  margin  arcuate,  slightly  reflexed,  form- 
ing an  obtuse  angle  in  the  young  with  the  white,  flat,  sloping  col- 
umella. Alt.  13,  greater  diam.  24,  lesser  19  mill.  (Hid.) 

Irosin  (around  the  river),  Prov.  Albay,  Luzon,  Philippines. 

H.  irosinensis  HID.,  Journ.  de  Conchyl.  1887,  p.  119,  t.  4,  f.  6; 
Obras  Malacol.  i,  p.  26,  t.  25,  f.  9. 


12l_>  COCHLOSTYLA-CORASIA. 

C.  asRUGiKOSA  Pfeiffer.     PI.  26,  fig.  6. 

Imperforate,  globose-depressed,  rather  thin,  striatulate,  shining, 
irivenish- white ;  spire  convex,  vertex  small,  obtuse ;  whorls  4,  a  little 
convex,  the  last  not  descending,  inflated;  periphery  obsoletely 
angulated,  at  the  suture  banded  with  rust  color;  columella  sloping, 
sliirhtly  arcuate,  subexcavated,  white  ;  aperture  oblique,  wide-lunar  ; 
peristome  slightly  reflexed,  margins  subconniving,  right  margin 
sinuous,  basal  margin  arcuately  passing  into  the  columella. 

Alt.  18,  greater  diam.  31,  lesser  26  mill.     (Pfr.) 

Is.  of  Bohol  and  Panglao,  Philippines. 

H.  ceriiginosa  PFR.,  P.  Z.  S.  1854,  p.  56 ;  Monogr.  iv,  p.  215.— 
REEVE,  Conch.  Icon.,  f.  1265.—  Cochlostyla  (Corasia)  aruginosa 
SEMP.,  Reis.,  p.  169.— H.  (Corasia)  ceruginosa  HIDALGO,  Obras,  p. 
152,  t.  21,  f.  3. 

Closely  allied  to  C.  eydouxi,  cegrota,  etc. 

C.  FILARIS  Valenciennes.     PL  25,  figs.  52,  53,  54  ;   PL  26,  figs.  10, 

11,  12. 

Imperforate,  depressed-globose,  convex  above  and  below,  obtusely 
angulated  at  the  periphery  ;  white  (covered  with  a  thin  deciduous 
yellowish  cuticle)  the  lip  broadly  edged  with  purplish-brown,  the 
suture  narrowly  white-margined  ;  surface  shining,  obliquely  striated 
earlier  whorls  seen  to  be  obsoletely,  spirally  striated  under  a  lens. 
Whorls  4-4-].  Aperture  oblique,  wide;  peristome  narrowly  ex- 
panded, the  columella  subvertical,  flattened,  somewhat  excavated, 
its  inner  edge  straight  or  a  little  irregular,  brown  below,  becoming 
white  above.  Alt.  20,  greater  diam.  33,  lesser  27  mill. 

Marinduque ;  Zamboanga,  Mindanao. 

H.filiiris  (YAL.  in  Paris  Mus.,  teste  Chiming)  PFR.  P.  Z.  S.  1845, 
p.  38  ;  Monogr.  i,  p.  202. — HIDALGO,  Obras  Malacol.  i,  p.  152,  t.  23, 
f.  1-4.— H.  nympha  PFR.  P.  Z.  S.  1849,  p.  129  ;  Monogr.  iii,  p.  147. 

KKKVE,  Conch.  Icon.,  f.  85. — /  C.  broderipi  var.  fasciata  v.  MOLL., 
Landsrlm.  Fauna  (1ebu,  p.  230. 

The  periphery  is  far  less  acutely  angled  than  in  C.  virgo,  and  the 
dark  border  of  the  lip  is  broader.  Fig.  52  of  pi.  25  represents  the 
typical  form.  Hidalgo  arranges  the  varietal  forms  as  follows: 

Yar.  1.  (PI.  24,  fig.  31).  Periphery  of  the  last  whorl  rounded. 
This  is  the  //.  nympha  of  Pfeiffer  and  Reeve. 

Var.  2.  (  PL  2(1,  tigs.  10,  11,  12).  Apical  whorls  purplish-black  ; 
nit ure  usually  narrowly  margined  with  brown. 

*  */  O 


COCHLOSTYLA-CORASIA.  123 

Var.  3.  (PI.  25,  fig.  54).  Like  the  last,  but  the  peripheral  angle 
brown.  Hidalgo  considers  C.  broderipi  var.  fasciata  Moll,  the  same 
as  this  variety.  It  is  from  the  western  coast  of  Zebu. 

Var.  4.  (PI.  25,  fig.  53).  Shell  tawny  or  chestnut,  peripheral 
angle  more  obtuse,  dark  banded. 

C.  EYDOUXI  Hidalgo.     PI.  26,  figs.  8,  9. 

Imperforate,  depressed-globose,  thin ;  wrhite  with  a  faint  greenish 
tint,  and  slightly  diaphanous  ;  above  the  suture  there  is  a  narroiv  red- 
dish-brown band,  obsolete  on  the  last  half- whorl.  Surface  smooth, 
shining,  obliquely  striatulate.  Generally  no  spiral  strise  are  visible, 
even  on  the  inner  whorls.  Spire  low,  convex,  apex  plane  ;  whorls  4, 
the  last  very  obtusely  subangular  at  the  periphery,  not  descending 
in  front.  Aperture  oblique,  wide ;  lip  white,  narrowly  expanded  ; 
columella  flattened,  adnate. 

Alt.  21,  greater  diam.  31,  lesser  36  mill. 

Capul,  Philippines. 

H.  valenciennesii  PFR.  in  Conch yl.  Cab.,  p.  309,  t.  53,  f.  1,  2; 
Monogr.  i,  p.  262. — REEVE,  Conch.  Icon.,  f.  87. — DELESSERT, 
Recueil,  t.  38,  f.  8,  (not  H.  valenciennii  EYDOUX,  Mag.  de  Zool. 
1838,  plate  115,  f.  2  !) — H.  eydouxi  HIDALGO,  Journ.  de  Conchyl. 
1887,  p.  115 ;  Obras  Malacol.  i,  pp.  25,  152,  t.  22,  f.  4. 

Distinguished  by  the  narrow  brown  margin  above  the  suture  of 
the  whorls  of  the  spire,  the  entire  shell  otherwise  being  white. 

The  true  H.  valenciennii  of  Eydoux  is  no  doubt  a  cromyodes 
deprived  of  epidermis,  as  Hidalgo  states. 

C.  BRODERIPI  Pfeiffer.     PL  24,  figs.  27,  28,  32,  33. 

Imperforate,  depressed-globose,  more  or  less  obviously  angular  at 
the  periphery,  thin,  very  finely  striated,  subtranslucent,  somewhat 
shining,  whitish.  Spire  little  raised,  with  the  apex  flat  or  often  a 
trifle  impressed.  Whorls  4,  flattened,  rapidly  widening,  the  last 
not  descending  in  front,  rather  swollen  beneath.  Aperture  oblique, 
broadly  lunar ;  columella  very  little  curved,  slightly  sloping,  some- 
what excavated,  deeply  inserted  above  ;  peristome  simple,  narrowly 
expanded,  a  little  thickened  toward  the  columella,  and  reflexed. 

Alt.  20,  greater  diam.  29,  lesser  24  mill. 

Philippines. 

H.  broderipi  PFR.  in  Conchyl.  Cab.  ii,  p.  27,  t.  68,  f.  6,  7,  (not 
H.  broderipi  Kve.) 


1^4  COCHLOSTYLA-CORASIA. 

The  above  description,  mainly  taken  from  Pfeiffer,  corresponds 
well  with  the  specimens  before  me,  two  of  which  are  figured  on  my 

plate. 

The  surface  is  almost  devoid  of  spiral  stria? ;  the  shell  is  entirely 
white,  fresh  speciim-ns  having  a  very  delicate  greenish  tint,  scarcely 
perceptible.  It  is  smaller  and  less  solid  than  C.  cegrota,  and  the 
peripheral  angle  is  more  obvious  ;  to  C.  eydouxi  it  is  very  closely 
allied,  but  it  lacks  the  constant  sutural  brown  line  characteristic  of 
that  species. 

C.  .EiiROTA  Reeve.     PI.  26,  fig.  1. 

Imperforate,  depressed-globose,  smooth,  white  or  of  a  very  delicate 
greenish  tint;  surface  shining  very  delicately  striatulate,  showing 
slight  traces  of  subobsolete  spiral  lines  on  the  earlier  whorls.  Spire 
low,  convex,  apex  plane ;  whorls  4£,  the  last  very  obtusely  sub- 
angular  at  the  periphery ;  aperture  oblique,  wide ;  peristome  white, 
narrowly  reflexed,  the  columellar  margin  dilated  and  adnate ;  col- 
umella  nearly  vertical,  its  inner  edge  straight,  forming  a  slight  angle 
with  the  basal  lip  at  their  junction. 

Alt.  23s-,  greater  diam.  37,  lesser  29  mill. 

Mind 'or o  ;  Badajos,  Tablets;   Tuburan,  Zebu. 

H.  cegrotus  Rv.,  Conch.  Icon.,  f.  95.— H.  cegrota  PFR.,  Conchyl. 
Cab.,  p.  437,  t.  152,  f.  3 ;  Monogr.  iii,  p.  191.— H".  (Corasia)  cegrota 
HIDALGO,  Obras  Malacol.  i,  p.  152,  t.  21,  f.  8. — Cochlostyla 
(Corasia)  cegrota  v.  MOLL.,  Landschn.  Fauna  Cebu,  p.  230  (a 
variety.) 

«.     / 

The  yellowish-green  tint  of  this  shell  is  so  delicate  as  to  be  scarcely 
perceptible.  The  apex  and  lip  are  pure  white.  It  is  more  solid  and 
stronger  than  C.  eydouxi  or  C.  virgo  and  its  immediate  allies. 
The  angulation  of  the  periphery  is  scarcely  sufficiently  marked  to  be 
noticeable. 

C.  -AMI;OAX<;A  Hombron  &  Jacquinot.     PI.  26,  fig.  13,  14,  15. 

Imperforate,   subdepressed,   thin,    delicately    striate,    moderately 
sliming,  waxen-white  with  two  reddish-brown  zones,  one  at  periphery, 
the  other  at  the  suture  ;    the  suture  opaque,  white,  irregularly  sub- 
crenulated.     Whorls  4,  convex,  depressed  above,  the  last  moder- 
at'-ly    inflated;    columella    obliquely   descending,   flattened,    acute, 
white  ;  aperture  rounded-rhombic  ;  peristome  obtuse,  straight,  white. 
Alt.  20,  -renter  diam.  30,  lesser  24  mill.     (v.  Mart.) 
Islands  of  Zamboanga  ;    Zebu  ;    Magtan ;    Mindanao ;    Basilan  ; 
L'i in I^H  It/an  ;   Canaaraman ;  Balabac;  Philippines. 


COCHLOSTYLA-CORASIA.  125 

H.  samboanga  H.  &  J.  Voy.  Pol  Slid  v,  p.  15,  t.  5,  f.  18-20.— 
PFR.  Mon.  Hel.  Viv.  iv,  p.  215. —  Cochlostyla  zamboangce  v.  MART., 
Ostas.  Zool.  ii,  p.  89. — H.  (Coraaia)  zamboangce  HIDALGO,  Obras. 
Malacol.  p.  151,  t.  22,  f.  2,  3. —  Cochlostyla  (Corasia)  Magtanensis 
SEMPER,  Archip.  Phil.  Landmoll.  iii,  p.  170,  t.  10,  f.  11. — v.  MOLL. 
Landschn.  Fauna  Cebu,  p.  231,  with  var.  nana  and  globosa. 

More  depressed  than  C.  intorta,  and  with  less  expanded  peristome. 
The  following  are  considered  varieties  by  Hidalgo. 

The  C.  magtanensis  of  Semper,  (pi.  28,  figs.  27,  28)  is  a  small 
form  from  the  island  of  Magtan.  It  is  not  separable  from  the  typi- 
cal samboanga,  being  identical  in  coloring. 

Var.  intaminata  Gould,  (pi.  27,  fig.  1)  is  entirely  white,  very  thin, 
and  visibly  angulated  at  the  periphery.  It  is  from  the  island  of 
Mangsi,  straits  of  Balabac. 

C.  INTORTA  Sowbery.     PL  28,  figs.  16-24. 

Im perforate,  globose-depressed,  thin,  rounded  at  the  periphery; 
typically  light  straw-colored  with  peripheral,  basal  and  sutural  bands 
of  chestnut  brown ;  (see  below).  Surface  smooth,  having  light 
growth-lines,  and  on  the  earlier  whorls  spiral  striae  are  visible  under 
a  lens.  Apex  flattened,  polished,  white,  brown,  pink  or  purple. 
Suture  linear,  margined  by  a  white  line.  Whorls  4-4},  the  last 
very  little  descending.  Aperture  broad,  rounded-lunar,  oblique  ; 
entire  lip  narrowly  expanded,  varying  from  flesh-tinted  to  dark 
brown.  Columella  oblique,  nearly  straight  above,  arcuate  below, 
somewhat  flattened  and  excavated. 

Alt.  21,  greater  diam.  32  mill. 

Alt.  24,  greater  diam.  32  mill. 

Alt.  17,  greater  diam.  26  mill. 

Islands  of  Tablas,  Panay,  Guimaras,  Negros,  Zebu,  Siquijor, 
Bohol,  Capul  and  Mindanao,  Philippines. 

H.  intorta  SOWERBY,  P.  Z.  S.  1840,  p.  168.— KEEVE,  Conch.  Syst. 
ii,t.!64,f.  10;  Conch.  Icon.,  t.  20.— PFR.,  Symboke ii,  p.  31 ;  Conchyl. 
Cab.,  p.  311,  t.  53,  f.  8-11 ;  Monogr.  i,  p.  262.— DELESSERT,  Rec.,  t. 
38,  f.  7. —  Cochlostyla  (  Corasia)  intorta  SEMPER,  Reisen,  p.  170.- 
v.  MOLL.  Landschn.  Fauna  Cebu,  p.  231  (with  varieties).— IT. 
(Corasia)  intorta  HIDALGO,  Obras  Malacol  i,  p.  149,  t.  22,  f.  5-9. 

The  form  and  elevation  of  the  spire  vary  in  this  beautiful  species, 
as  the  measurements  and  figures  show.  Specimens  having  an  obtuse 
peripheral  carina  also  occur.  The  principal  color-forms  are  as 
follows : 


]:>»>  COCHLOSTYLA-CORASIA. 

1.  Ground  color  yellowish,  bands  three,  distinct,  not  spreading  on 
the  latter  part  of  the  whorl. 

'2.  Ground  color  yellowish,  no  bauds. 

3.  Ground  color  light-brown  below,  whitish  above,  bauds  three, 
dark-chestnut. 

4.  Chestnut  colored,  with  darker  bands ;  apex  purple. 

5.  Delicate  light-greenish,  with  a  subsutural   and    a  peripheral 
purple-brown  baud,  the  bands  spreading  over  the  entire  whorl  on  its 
latter  portion. 

6.  Delicate  greenish-white,  peripheral  and  sutural  bands  present, 

but  narrow. 

v.  Mollendorff  describes  the  following  varieties: 

1.  var.  tennis  v.  Moll.     Thin  shelled,  with  a  narrow  brown  sutural 
baud,  a  wider  brown  peripheral  band,  and  a  narrow  vertical  brown 
streak  behind  the  blackish-brown  lip,  to  which  the   spiral   bands 

extend. 

j\R.  Mangilao. 

2.  Var.  expansilabris  v.  Moll.     Spire  more  depressed  ;  shell  white, 
-Inning,  the  periphery  distinctly  angulated,  without  peripheral  band  ; 
sutural  band  evanescent ;  lip  more  expanded,  upper  margin  at  first 
horizontal,  then  well  curved. 

Mts.  Uling  and  Licos,  Zebu. 

This  is  the  Cochlostyla  (Corasia)  virgo  var.  subglobosaof  v.  Moll., 
Mai.  Bl.  (X.  F.)  x,  p.  161. 

3.  Yaf.  craxsa  v.  Moll.     Shell  smaller  than  the  type,  more  solid, 
>pire  more  depressed  ;  whorls  4  (uot4£)  ;  white  under  the  deciduous 
epidermis,  with  narrow  sutural  and  broad  peripheral  bauds,  widening 
and  confluent  toward  the  aperture;  peristome  thickened,  little  ex- 
panded, white;  aperture  brown  inside. 

Medellin,  Northen  Zebu. 

4.  Var.  slquijorica  v.  Moll.     See  pi.  28,  figs.  19,  20,  21,  22.     This 
i-  tin-  sum-  as  the  fifth  color-form  described  above. 

C.  LIMANSAUENSIS  Semper.     PL  28,  figs.  25,  26. 

Imperforate,  globose  or  globose-depressed,  papery,  very  diapha- 
nous, whitish  or  cinnamon  colored,  having  a  brown  band  at  the 
periphery;  suture  usually  manrined  with  rufous  in  the  earlier 
whorls;  whorls  4,  the  upper  depressed,  last  very  convex,  inflated 
beneath;  rolinm-lla  whitish,  nearly  straight;  aperture  nearly  tri- 
angular; peristome  simple,  rufous  within,  scarcely  inflexed. 


COCHLOSTYLA-PFEIFFERIA.  127 

Alt.  26-28,  greater  diam.  34-38,  lesser  27-30  mill.     (Semper.) 

Island  of  Limansau  a,  between  Leyte  and  Mindanao. 

C.  (Corasia)  limansauensis  SEMP.,  Reis.  Phil.  Arch.  Landmoll.  iii, 
p.  171,  t.  9,  f.  6.  —  If.  (Corasia)  limansauensis  HIDALGO,  Obras,  i,  p. 
149,  t.  21,  f.  7. 

Very  closely  allied  to  C.  intorta,  but  thin  as  paper,  lacking  com- 
pletely a  basal  brown  tract,  having  an  almost  vertical  columella, 
and  of  larger  size.  Like  all  other  Cochlostylas  it  lives  upon  trees 
near  the  beach,  the  leaves  of  which  it  twists  into  a  bag  for  the 
reception  of  its  eggs.  It  is  the  only  species  of  Cochlostyla  found  by 
Semper  on  the  island  Limansaua.  Hidalgo  doubts  its  distinctness 
from  C.  intorta. 


C.  BOURDILLONII  Theobald.     PI.  33,  fig.  55. 

Shell  conoid,  covered-perforate,  carinated,  thin,  polished,  whitish, 
translucent  ;  ornamented  under  the  deciduous,  pale  straw-colored 
cuticle  with  very  fine  flexuous  spiral  lines.  Whorls  4J,  rapidly 
increasing,  a  little  convex  toward  the  mamillate  apex,  but  the  last 
whorl  flat  above,  and  a  little  inflated  around  the  umbilicus.  Aper- 
ture large,  subquadrate;  margins  simple,  joined  by  a  very  thin 
callus.  Alt.  14'8,  gr.  diam.  25*4,  lesser  19'5  mill.  ;  apert.  alt.  16, 
width  14  mill.  (Theob.) 

Prov.  Travancore,  India. 

Corasia  bourdillonii  THEOB.,  Journ.  Asiat.  Soc.  Beng.  XLV,  (pt. 
ii)  p.  185,  t.  14,  f.  3  (1876). 

Judging  from  the  sharp  lip,  as  well  as  the  locality,  this  is  no 
Corasia.  It  may  prove  to  be  a  Nanina.  In  the  absence  of  certain 
information  it  may  as  well  remain  where  first  described  for  the 
present. 

Section  III.     PFEIFFERIA  Gray,  1853. 

Pfei/eria  GRAY,  P.  Z.  S.  1853,  p.  110.      Type  H.  micans  Pfr.- 
SEMPER,  Phil.  Archip.  p.  225. 

Shell  globose,  imperforate,  the  slender  axis  solid,  thin,  brittle, 
white,  with  few  whorls  ;  aperture  lunar,  lip  thin,  acute,  fragile  ; 
columella  slightly  thickened,  vertical,  deeply  inserted  in  the  base. 

Animal  like  Cochlostyla,  but  the  mantle  reflexed  over  the  acute 
edge  of  the  shell. 


1  J.x  COCHLOSTYLA-PFEIFFERIA. 

Q  MI-  una  Pfeiffer.     PI.  19,  figs.  12,  13,  14. 

Shell  impertbrate,  subglobular,  thin,  brittle;  translucent-whitish, 
with  an  opaque  white  margin  below  the  suture.  Surface  shining, 
obliquely  striatiilate,  obsoletely,  finely  spirally  striated.  Whorls  4, 
the  last  globose,  not  descending  in  front.  Aperture  a  little  oblique, 
lunar ;  peristome  1hin,  fragile  and  acute,  the  upper  margin  curving 
forward  ;  coltiraella  vertical,  slightly  sinuous,  deeply  inserted  in  the 
base.  Alt.  22,  greater  diara.  26,  lesser  22  mill. 

Province  of  Cagayan,  Northern  Luzon,  Philippines. 

H.  micans  PFR.,  P.  Z.  S.  1845,  p.  71 ;  Conchyl.  Cab.  p.  227,  t.  28, 
f.  3-5;  Monogr.  i,  p.  24. — REEVE,  Conch.  Icon.  f.  46a. — Pfeifferia 
micans  GRAY,  P.  Z.  S.  1853,  p.  110. —  Cochlostyla  micans  SEMPER, 
Reis.  Archip.  Phil.  Landmoll.  p.  225. 

The  thin  fragile  lip  of  this  species  separates  it  at  once  from  all 
other  forms  of  Cochlostyla. 

Semper  describes  the  animal  of  this  interesting  species  as  follows: 
The  border  of  the  mantle-edge  all  around,  in  the  contracted  speci- 
mens, covers  about  1-2  mm.  of  the  shell.  On  the  left  there  is  a 
-mall  mantle-lappet,  but  none  on  the  right. 

The  foot  is  broad,  flat,  without  mucus-gland  and  without  middle 
area  to  the  sole. 

The  color  of  the  mantle  in  the  region  of  the  lung  is  greenish  ;  the 
foot  and  mantle-margin  are  yellowish. 

The  right  tentacle  passes  with  its  retractor  muscle,  between  the 
branches  of  the  genitalia,  an  exception  to  the  rule  in  Cochlostyla. 
The  genitalia  are  just  as  in  Cochlostyla.  The  receptaculum  semi- 
nis  has  a  long  duct,  without  diverticulum.  The  dart-sac  is  globu- 
lar, thick ;  and  on  the  base  of  the  short  globule  (or  mucus)  gland 
there  is  a  small  sac,  apparently  a  gland.  The  vas  deferens  is  wide, 
and  parses  gradually  into  the  simple  penis,  which  lacks  all  acces- 
sory LI  lands  ;  the  retractor  penis  is  short  and  thick. 

The  jaw  has  numerous  (about  8)  pretty  wide  and  close  ribs,  pro- 
jecting on  the  cutting  edge. 

The  radula  is  entirely  typical  of  Cochlostyla;  the  central  tooth 
has  a  broad  cusp ;  there  are  fourteen  or  fifteen  side-teeth  or  true 
lateral- ;  al ready  the  eighteenth  tooth  being  distinctly  tricuspid.  In 
all  there  are  101  teeth  in  each  transverse  row. 


COCHLOSTYLA-LEYTIA-CALOCOCHLEA.  129 

Section  IV.     LEYTIA  Pilsbry. 

Tin  perforate,  globose,  thin  ;  whorls  few  (4),  the  last  angulated  at 
the  periphery.  Aperture  very  large,  lip  simple,  a  little  expanded 
below ;  columella  simple,  thin,  vertical,  deeply  inserted.  Surface 
spirally  striate,  having  hydrophanous,  cream-white  bands. 

This  section  differs  from  Calocochlea  in  its  thin,  bubble-like  shell, 
composed  of  few  whorls  ;  from  Corasia  in  the  possession  of  hydro- 
phanous cuticle,  and  the  globose  contour.  The  anatomy  has  not 
been  investigated. 

C.  FRAGILTS  Sowerby.     PI.  29,  figs.  5,  6. 

Imperforate,  globose,  thin  ;  light  blue-green  above  the  periphery, 
becoming  pinkish-white  on  the  spire,  the  last  whorl  having  numer- 
ous narrow  interrupted  white  bands;  below  the  periphery  dark 
green,  with  two  broad  straw-colored  zones  and  numerous  narrow, 
broken,  whitish  lines ;  lighter  around  the  center  of  the  base.  Sur- 
face finely  spirally  striated,  obliquely  striatulate.  AVhorls  4,  the 
apex  minute,  obtuse,  brown  ;  last  whorl  angulated,  not  descending 
in  front.  Aperture  large,  oblique  ;  peristome  white. 

Alt.  21,  greater  diam.  26,  lesser  22J  mill. 

Tanauan,  Island  of  Leyte,  Philippines. 

H.jragilis  &OWB.,  P.Z.  S.  1841,  p.  40. — H.  ( Corasia)  fragilis, 
HID.,  Obras.  i,  p.  149,  t.  24,  f.  8,  9. — H.  leytensis  PFR.,  Symbols  iii, 
p.  71  ;  Couchyl.  Cab.  t.  42,  f.  7,  8  ;  Monogr.  i,  p.  242.— RYE.,  Conch. 
Icon.  f.  55. — CROSSE,  Journ  de  Conchy],  xxiii,  p.  134,  t.  6,  f.  3.  1875. 

A  bubble-like  species,  rare  in  collections.  It  has  an  unfinished 
aspect,  even  when  quite  mature. 

Section  V.     CALOCOCHLEA  Hartmann,  1840. 

Calocochlea  HARTM.,  Erd  u.  Siisswasser  Gasteropoden  der  Schweiz, 
etc.,  p.  163.  Type  C.  pulcherrimaSowb.,  1.  c.,  pi.  54. —  C-dlicochlias 
AGASSIZ,  Nomencl.  Zool.,  1847. —  ALBERS,  Die  Hel.  p.  105,  et  al. 

Imperforate,  solid  shells,  varying  from  globose  to  depressed- 
globose  or  lens-shaped  and  carinated.  Generally  covered  with  hy- 
drophanous cuticle,  variously  patterned.  Anatomy  as  in  Cocldostyla 
generally. 

Under  Calocochlea  must  be  placed  Axina  and  a  number  of  other 
groups  which  stand  too  close  to  one  another  to  be  separated  as  pri- 
mary divisions  of  Cochlostyla,  although  they  have  a  certain  systematic 

value. 

9 


1 30  COCHLOSTYLA-CALOCOCHLEA. 

As  subsections  of  r.ihcochlea,  therefore,  we  may  range  the  follow- 
ing groups: 

Subsection  1.     CALOCOCHLEA  Hartm.,  1840. 

Subglobose  or  globose-depressed,  the  periphery  rounded.     Surface 
with  oblique  light  growth-lines,  and  on  the  whorls  of  the  spire,  fine 
spirals,  often  subobsolete ;   hydrophanous  cuticle  generally  present. 
Type,  ( '.  pukherrima. 
Subsection  2.     AXINA  Albers,  1850. 

Varying  from  globose-turbinate  to  carinated  and  lens-shaped ; 
solid,  dark-colored  ;  hydrophanous  cuticle  present.  Surface  totally 
lacking  spiral  stria1.  Type,  0.  zebuensis. 

Subsection  3.     TRACHYSTYLA  Pilsbry. 

Solid,  globose-depressed,  dark  colored,  with  a  thin  ashen  hydro- 
phanous cuticle  ;  surface  rough  and  lusterless.  Whorls  few.  Type, 
C.  crypt ica. 


Subsection  1.  CALOCOCHLEA  Hartm. 

Includes  Semper's  "  3  Gruppe,  Calliocochlias  Ag."  and  "4  Gruppe> 
Cochlottylu'  hypomelanice  Mts." 

(1.)    Group  of  C.  cromyodes. 

Globose  or  globose  depressed  shells,  thin  rather  than  solid,  yellow 
or  reddish,  not  conspicuously  marked  with  hydrophanous  cuticle, 
and  frequently  lacking  it  entirely. 

These  shells  connect  the  ('.  intorta  group  of  Corasia  with  Cah- 
cochle<i.  The  dividing  line  is  extremely  difficult  to  locate,  but  still 
there  is  some  difference,  as  these  shells  have  a  wider  lip,  and  gener- 
ally have  more  or  less  hydrophanous  cuticle,  —  characters  not  found 
in 


(  '.  .  ROMYODES  Pfeifier.     PI.  2!>,  figs.  13,  14,  15,  16. 

Imprrf'orate,  rather  thin,  depressed-globose.  Surface  microscop- 
ically spirally  striate,  having  light  growth-lines.  Color  varying 
from  greenish-yellow  to  reddish-brown,  the  upper  whorls  purple,  edged 
\\ith  darker  purple.  Hydrophanous  cuticle  either  absent  or  form- 
in-  hand-  at  suture  and  periphery,  with  narrower  intervening  bands 
or  line-. 


COCHLOSTYLA-CALOCOCHLEA.  131 

Whorls  4J  ;  suture  shallow  ;  apex  very  obtuse ;  last  whorl  but  lit- 
tle descending.  Aperture  oblique,  white  or  tinted  inside;  lip  re- 
flexed,  white,  edged  with  dark  brown;  coluraella  oblique. 

Alt.  27,  diam.  35  mill. 

Alt.  21,  diam.  28  mill. 

Province  of  Misamis,  northern  Mindanao. 

H.  cromyodes  PFR.,  P.  Z.  S.  1842,  p.  150;  Monogr.  i,  p.  261  ; 
Conchyl.  Cab.  t.  53,  f.  6,  7. — RVE.,  Conch.  Icon.  f.  1429. —  Cochlo- 
styla  cromyodes  SEMPER,  Reis.  p.  171. — H.  valenciennii  EYDOUX, 
Mag.  de  Zool.  1838,  t.  115,  f.  2,  a  dead  shell,  denuded  of  cuticle. 
(Not  H.  valenciennesii  Pfr. ;  see  Corasia  eydouxi  Hidalgo,  this  vol- 
ume p.  123.) 

Rather  thin,  and  generally  unicolored  except  for  the  darker  spire. 
Some  specimens  show  a  faint  peripheral  dark  band,  and  rarely 
a  dark  band  is  developed  around  the  columella.  Numerous  narrow 
hydrophanous  bands  are  seen  on  some  specimens. 

C.  DENTICULATA  Jay.     PI.  52,  figs.  17,  18,  19. 

Imperforate,  depressed-globose,  decidedly  solid.  Surface  smooth 
the  growth  lines  inconspicuous.  Color  a  light  brown  tint,  the  spire 
white,  having  a  dark  brown  margin  above  the  suture. 

Spire  very  low,  apical  whorl  plane.  Suture  superficial.  Whorls 
4,  the  last  wide,  slightly  descending  and  then  ascending  in  front. 
Aperture  oblique,  white  within  ;  lip  rather  thick,  reflexed,  edged 
with  dark  brown.  Columella  very  oblique,  wide,  white,  ending  be- 
low in  a  stout  tooth.  Alt.  25,  greater  diam.  34,  lesser  29  mill. 

Habitat  unknown. 

H.  denticulata  JAY,  Catalogue  (edit,  of  1839),  p.  114,  t.  1,  f.  21. 
-PrR.,  Monogr.  i,  311. 

The  above  description  and  the  figures  are  from  Jay's  type  speci- 
men, kindly  loaned  by  Prof.  R.  P.  Whitfield,  of  the  American 
Museum  of  Natural  History,  New  York  City. 

The  shell  is  decidedly  solid  ;  the  columella  is  wide,  its  face  gently 
convex,  its  base  strongly  toothed,  much  as  in  C.  curia  Sowerby. 
The  cuticle  is  wholly  worn  from  the  specimen,  but  under  the  very 
thin  parietal  callus  it  is  seen  to  be  of  the  yellow  color  of  that  of  C. 
cromyodes,  a  species  also  resembling  denticulata  very  closely  in  con- 
tour, in  the  suprasutural  margination  of  the  spire  whorls,  in  the 
appearance  of  the  surface,  and  the  dark-edged  lip.  From  C. 
cromyodes,  the  present  species  differs  only  in  being  more  solid,  in  the 


COCHLOSTYLA-CALOCOCHLEA. 

wider,  heavier  and  toothed  columella,  and  the  wider,  thicker  lip. 
These  characters  compel  me  to  regard  denticulata  as  a  distinct 
-]•  >cies,  even  though  but  one  specimen  of  it  is  known  to  me. 

C.  AKMAIKNSIS  Sowerby.     PI.  52,  figs.  20,  21. 

Imperforate,  depressed-globose,  shining,  white,  uuicolored  or 
brown-banded  ;  spire  depressed,  suture  linear  ;  whorls  4,  nearly  flat, 
the  last  inflated  beneath  ;  columella  thin,  nearly  straight,  sloping, 
profoundly  excavated.  Aperture  lunate-oval,  upper  margin  sub- 
depressed;  peristome  widely  expanded,  white,  the  margin  subthick- 
rned,  forming  a  distinct  angle  with  the  base.  (Pfr.) 

Alt.  26,  diam.  39  mill. 

Palanan  to  Camiguiii  de  Luzon,  E.  coast  of  N.  Luzon,  and  Amber- 
link,  Province  of  La  Isabella,  Luzon. 

H.  albaiensis  Sown.,  P.  Z.  S.,  1840,  p.  100.— PFR.,  Monogr.  i,  p. 
2IJO;  Conchyl.  Cab.  p.  p,  215,  t.  108,  f.  1,2.— REEVE,  Conch.  Icon, 
t.  13.  f.  48. —  Cochlostyla  albaiensis  SEMPER,  Reis.,  p.  176,  t.  8,  f.  7  ; 
t.  18,  f.  14  (anatomy). 

( J.  TUKANEXSIS  Pfeiffer.     PL  29,  figs.  7,  8,  9,  12. 

Imperforate,  subglobose,  thin  ;  surface  smooth,  the  spiral  and  in- 
cremental striae  almost  obsolete.  White  or  rosy  under  a  thin  yel- 
lowish or  reddish  cuticle,  sometimes  having  a  dark  band  at  periph- 
ery and  another  at  suture;  hydrophanous  cuticle  generally  absent, 
but  sometimes  forming  a  few  bands. 

Spire  moderately  elevated,  apex  very  obtuse ;  whorls  4s,  convex, 
the  last  not  perceptibly  descending  in  front.  Aperture  large,  pink 
or  white  within  ;  peristome  thin,  pink  or  white,  broadly  expanding ; 
columella  oblique,  flattened,  thin,  nearly  straight. 

Alt.  29,  diam.  31  mill. 

Tukan  Bessi  Id.  (Pfr.)  ;  Pular  Is.  (Horn}. 

II .  tnL-iiiiensis  PFR.,  Mai.  Bl.  xviii,  p.  122,  1871  ;  Xovit.  Conch, 
iv,  p.  72,  t.  121.  f.  5-<> ;  Monogr.  vii,  p.  324. 

A  more  elevated  and  thinner  shell  than  C.  cromyodes.  Part  of 
the  specimens  before  me  are  yellow  instead  of  reddish,  and  some  are 
very  small,  alt.  17,  diam.  23  mill. 

C.  OBTUSA  I'foih-or.      PI.  35,  figs.  23,  24. 

[mperf'orate,  subglobose,  thin,  subdecussated    with    growth-lines 
ami  impn-v-cd  obsolete  concentric  lines,  shining,  chestnut.     Spire 
,  obtuse.       Whorls  4,  convex,  the  last  inflated.     Colu- 


COCHLOSTYLA-CALOCOCHLEA.  133 

mella  oblique,  acute,  white.  Aperture  dilated,  subauriform,  with  a 
pearly  luster  inside  ;  peristome  simple,  expanded.  Alt.  26,  diam. 
35  mill.  (P/r.) 

Luzon,  Philippines.     (Cuming.) 

H.  obtusa  PFR.,  P.  Z.  S.  1845,  p.  38  ;  Monogr.  i,  p.  253  ;  Conchyl. 
Cab.  p.  437,  t.  152,  f.  1,  2.— REEVE,  Conch.  Icon.  f.  49. 

C.  AMICTA  Reeve.     PI.  34,  figs.  7,  8. 

Imperforate,  subturbinate,  semiglobose,  smooth,  white,  covered 
with  an  oily  huffish  epidermis  ;  spire  conoid,  a  little  obtuse;  whorls 
4i,  a  little  convex,  the  last  not  descending,  subdepressed,  about 
equaling  the  spire;  base  rather  flattened;  columella  subvertical, 
lightly  arcuate,  white.  Aperture  oblique,  subtrapezoidal-lunar, 
white  inside ;  peristome  white,  somewhat  thickened,  narrowly  ex- 
panded. Alt.  19,  greater  diam.  30,  lesser  26  mill.  (Pfr.) 

Philippines. 

H.  amicta  RVE.,  Conch.  Icon.  f.  52. — PER.,  Monogr.  iii,  p.  173  ; 
Conchyl.  Cab.  p.  442,  t.  152,  f.  17,  18. 

C.  DECORA  Adams  &  Reeve.     PI.  52.  figs.  23,  24. 

Imperforate,  conic-semi-globose,  solid,  nearly  smooth,  little  shin- 
ing, straw-colored,  irregularly  variegated  with  darker  streaks. 
Spire  conoid,  apex  obtuse,  pale.  Whorls  4,  little  convex,  the  last 
subcarinated,  not  descending  in  front :  base  smooth.  Aperture 
oblique,  obsoletely  quadrangular,  white  within;  peristome  simple, 
a  little  expanded,  columellar  margin  flat,  dilated.  Alt.  18,  greater 
diam.  26  mill.  (P/r.) 

Mindoro,  Philippines. 

H.  decora  AD.  &  Rv.,  Voy.  Samarang,  Zool.,  Moll.,  p.  62,  t.  16,  f. 
7.— PER.,  Monogr.  iii,  p.  178;  Conchyl.  Cab.  p.  401,  t.  144,  f.  18, 

19. — REEVE,  Conch.  Icon.  f.  586. 

• 

(2.)    Group  of  C.  pulcherrima. 

C.  PULCHERRIMA  Sowerby.     PL  33,  figs.  60-64. 

Imperforate,  globose,  somewhat  depressed,  generally  solid ; 
ground-color  uniform,  varying  from  buff  or  cream-color  to  a  dark 
chestnut-maroon  ;  sometimes  unicolored,  apparently  without  any 
hydrophanous  markings  whatever,  but  generally  encircled  by  nu- 
merous white  or  creamy  bands  which  are  closely  articulated  or 
broken  into  dots  and  spots. 


(  '<  >CHLOSTYLA-CALOCOCHLEA. 

The  surface  is  nearly  smooth.  Whorls  5,  the  last  one  scarcely 
or  slightly  deflexed  in  front.  Aperture  white  within,  rounded-lunar  ; 
peristome  reflexed  throughout,  white,  often  edged  with  flesh-color. 
( olumella  deeply  entering,  gently  arcuate. 

Alt.  •'>!,  diam.  35  mill,  (average  specimen). 

Alt,  31,  diam.  43  mill,  (large  thin  specimen  from  Luzon  ;  mouth 
deep  pink  within). 

Alt.  26,  diam.  31  mill,  (small  buff  specimen). 

H.  pulcherrima  SOWB.,  P.  Z.  S.  1840,  p.  90.— REEVE,  Conch. 
Syst.  ii,  t.  KU.  f.  12,  13;  Conch.  Icon.  f.  26.— PFR.,  Monogr.  i,  p. 
•2  ~>0  ;  Conchyl.  Cab.  t.  45,  f.  1-4. — DELESSERT,  Rec.  de  Coq.  t.  38, 
f.  11,  12, 15,  16. —  Galocochlea  pulcherrima  HARTM.,  Gastr.  Schweiz, 
i,  j).  163,  f.  54. —  Coehlostyla  pulcherrima  Semper,  Reis.,  p.  174, 1. 13, 
f.  9;t.  18,  f.  16  (anatomy). — HIDALGO,  Journ.  Conch.  1887,  p. 
126. — H.f estiva  MORCH.,  not  Donovan. 

The  ground-color  is  extremely  variable,  every  shade  between 
light  buff  or  light  olive-green  and  deep  maroon  being  represented. 
It  is  nearly  uniform  in  tint,  becoming  pink  on  the  earlier  whorls  in 
most  specimens.  In  about  one-third  of  the  specimens  before  me 
there  is  a  narrow  dark  band  bordering  the  columella,  generally  not 
very  distinct.  In  any  of  the  color-forms,  from  light  to  dark,  bands 
of  close  spots  of  hydrophanous  cuticle  are  frequently  developed. 
The  supraperipheral  hydrophanous  band  is  the  most  constant  and 
prominent. 

( '.  FKM-IVA  Donovan.     PI.  52,  fig.  25  ;  pi.  35,  figs.  11-16. 

Imperforate,  globose,  somewhat  depressed,  the  spire  generally 
rather  conoidal ;  solid;  ground-color  (beneath  the  hydrophanous 
cuticle)  dark  reddish-chestnut,  sometimes  yellowish-chestnut,  usu- 
ally with  a  light  peripheral  girdle.  Covered  with  a  cream-colored 
or  li!_rht  brown  cuticle,  with  a  white  zone  at  periphery. 

The  hydrophanous  cuticle  is  often  closely  streaked  obliquely,  and 
•nerally  has  some  or  many  .spiral  dark  lines.  The  apical  whorls 
arc  pink  or  purple-pink.  Whorls  43-5,  convex.  Aperture  wide- 
lunar,  pink  or  white  inside;  peristome  broadly  reflexed,  typically 
Imirht  pink,  but  often  white,  with  or  without  a  pink  edge.  Colu- 
mella pink  or  white,  broad,  irregularly  arcuate. 

Alt.  29,  diai. i.  35  mill. 

Alt.  38,  diam.  44  mill. 

Province  of  Cagayan,  Luzon,  Philippines* 


COCHLOSTYLA-CALOCOCHLEA.  135 

H.  /estiva  DONOVAN,  Naturalist's  Repository  iii,  pi.  103  (Feb., 
1825).— PER.,  Monogr.  iii,  p.  187  ;  Conchyl.  Cab.  t.  139,  f.  19,  '20. 
-RvE.,  Conch.  Icon.  f.  107. — H.  luzonica  SOWB.  (in  sched.  Cum- 
ing),  PFR.,  Monogr.  i,  p.  252  ;  Conchyl.  Cab.  t.  45,  f.  7,  9,  10, 11.— 
RVE.,  f.  41. — SEMPER,  Reis.,  t..  8,  f.  1. — H.  annceO.  SEMPER,  Jo  urn. 
de  Conch.  1862,  p.  146,  t.  5,  f.  8,  9. —  Cocklostyla  Jestiva  SEMP.,  Reis. 
p.  176. — HIDALGO,  Journ.  de  Conchyl.  1887,  p.  127. 

Fig.  25  of  pi.  52  is  the  typical  f  estiva.  On  pi.  35,  figs.  11, 
12  represent  the  C.  annce  Semper ;  the  figures  13-16  are  the  form 
luzonica  Sow. 

This  species  has  less  variegated  cuticle  than  C.  pulcherrima  and 
the  white  equatorial  zone  is  conspicuous.  The  lip  is  heavier  than  in 
pulcherrima ;  its  face  is  typically  pink,  but  many  specimens  have  a 
pure  white  lip  and  columella. 

C.  DUBIOSA  Pfeiffer.     PI.  32,  figs.  52,  54. 

Imperforate,  globose-depressed,  rather  thin  but  solid.  Ground- 
color under  the  hydrophanous  cuticle,  varying  from  yellow  to  brown, 
and  having  two  dark  chestnut  zones,  one  above,  the  other  below  the 
periphery.  Covered  with  a  cream-white  hydrophanous  cuticle, 
which  is  cut  into  a  great  number  of  very  fine  lines  by  fine  close  darker 
spiral  lines. 

The  surface  is  finely  spirally  striated.  Whorls  4A,  the  last  a  lit- 
tle descending  in  front.  Aperture  rounded,  white  and  showing  the 
zones  within ;  lip  broadly  reflex ed,  thin,  white ;  columella  nearly 
straight,  white,  forming  an  obtuse  angle  at  its  base  with  the  lower 
lip. 

Alt.  30,  diam.  31  mill. 

Mts.  of  Morong  district,  Luzon, 

H.  dubiosa  PER.,  Symbols  iii,  p,  71  ;  Conchyl.  Cab.  t.  46,  f.  5, 
6  ;  Mouogr.  i,  p.  252. — RVE.,  Conch.  Icon.  f.  22. — H.  decipiens  var. 
DELESSERT,  Req.  de  Coq.  t.  38,  f.  10. —  Cochlostyla  dubiosa  HIDALGO, 
Journ.  de  Conch.  1887,  p.  145. — H.  batanica  RVE.,  Couch.  Icon.  t. 
9,  f.  2. — H.  volubills  Rv.  Errata  to  Conch.  Icon. —  Cochlostyla 
batanica  HIDALGO,  Journ.  de  Conchyl.  1887,  p.  129. — H.  speeiosa 
JAY,  Catal.  (2nd  edit.),  plate  iii,  f.  9,  9  (1836). 

The  most  prominent  feature  of  this  species  is  its  thin,  broadly  ex- 
panded lip,  yellow-brown  ground-color  two-banded  with  chestnut, 
and  the  excessively  fine  and  numerous  lines  of  hydrophanous  cuti- 
cle of  the  exterior. 


1  :•!•;  COCHLOSTYLA-CALOCOCHLEA. 

Hidalgo  believes  the  H.  batanica  (+  volubilis)  of  Reeve  to  be  a 
distinct  species.  I  am  unable,  with  the  material. before  me,  to  make 
the  separation,  but  still  it  may  exist.  Reeve's  figure  of  batanica  is 
copied  on  my  plate  32.  fig.  53. 

I  have  examined  the  type  of  Helix  speciosa  Jay,  now  in  the 
American  Museum  of  Natural  History,  Central  Park,  New  York 
( 'ity.  It  is  a  typical  C.  diibiosa.  The  name  speciosa  has  priority 
nver  dubiosa,  but  as  Jay's  description  and  figure  have  been  demon- 
strated to  be  utterly  inadequate  by  the  failure  of  Pfeiffer  and  others 
to  identify  the  species  correctly,  its  rehabilitation  as  a  specific  name 
would  be  obviously  contrary  to  the  recognized  canons  of  nomencla- 
ture. 

C.  LALLOENSIS  Pfeiffer.     PL  51,  fig.  16  ;  pi.  26,  figs.  2,  3. 

Im perforate,  globose-depressed,  rounded  at  the  periphery,  thin, 
bluish- white  (denuded  of  cuticle)  with  two  dark  brown  bands,  one 
above,  the  other  below  the  periphery  ;  there  is  also  a  narrow  brown 
line  below  the  suture,  ascending  the  spire. 

Whorls  4-4.;  ;  apex  obtuse,  white;  last  whorl  subglobose,  a  little 
descending  in  front ;  aperture  rounded-lunar,  oblique,  showing  the 
bands  within  ;  peristome  white,  well  expanded  ;  columellar  margin 
widened,  sloping,  slightly  arcuate,  flattened.  Alt.  26,  diam.  33  mill. 

Lallo,  Province  of  Cagayan,  Luzon,  Philippines. 

H.  lalloensis  PFR.,  P.  Z.  S.  1855,  p.  Ill ;  Monogr.  iv,  p.  214. 

The  specimen  figured  on  pi.  26  is  immature.  Numerous  speci- 
mens are  before  me,  but  all  are  white,  as  if  denuded  of  cuticle.  It 
seems  to  be  allied  to  C.  dvbiosa,  differing  in  the  possession  of  a  sut- 
u ml  band  and  the  more  oblique  columella.  The  surface  is  very 
finely  roughened  and  spirally  substriated. 

(1.  AUGUSTA  Albers.     PI.  35,  figs.  17,  18. 

Imperforate,  depressed-globose,  rather  solid,  spirally  closely 
striated,  shining,  tawny,  ornamented  with  two  wide  blackish-chest- 
nut hands  with  a  white  space  of  the  same  width  between  them. 
Spire  convex,  obtuse ;  suture  white,  margined  with  chestnut ;  whorls 
4,  a  little  convex,  the  last  rounded,  slightly  descending  in  front. 
Ap.-nure  oblique,  lunate-round,  whitish  inside,  showing  pellucid 
bands;  peristome  white,  expanded;  columellar  margin  oblique, 
nearly  Mraightly  descending,  flat,  white,  subtruncate  at  base. 

Alt.  _'l .  greater  diam.  34,  lesser  28  mill.  (Pfr.  from  spec,  in  coll. 
Albers. 

Habitat  unknown. 


COCHLOSTYLA-CALOCOCHLEA.  137 

H.  augusta  ALB.,  Mai.  Bl.  i,  1854,  p.  214. — PFR.,  Mon.  iv,  p. 
214;  Novit.  Conch,  i,  p.  41,  t.  11,  f.  9,  10. 

The  locality  " New  Guinea?"  given  by  Albers  is  extremely 
doubtful. 

VAR.  On  pi.  32,  fig.  5],  I  have  figured  a  specimen  which  may 
be  a  variety  of  this  species,  but  I  am  in  doubt  of  it. 

C.  PRINCEPS  Keeve.     PI.  33,  figs.  58,  59. 

Imperforate,  oblong-turbinated,  solid,  rather  swollen,  very  finely 
rugulosely  striated  ;  yellowish,  purple  at  the  apex,  covered  with  a 
slight  hydrophanous  epidermis.  Aperture  lunar-oval ;  lip  thinly 
reflected,  columellar  margin  appressly  tubercled,  callous,  drawn  out. 

(Reeve.) 

Philippines  Is. 

H.  princeps  RVE.,  Conch.  Icon.,  f.  1422. — PFR.,  Monogr.  iv,  p. 
209. 

The  specimen  before  me  (fig.  59)  does  not  agree  in  all  respects 
with  Reeve's  description  and  his  figure  (fig.  58).  It  is  thinner, 
smoother,  not  so  distinctly  banded. 

C.  ERYTHROSPIRA  MollendorfF.     Unfig ured. 

Imperforate,  subglobose,  quite  thin,  above  bright  violet-red,  then 
chestnut  brown-black,  covered  with  a  wood-colored  hydrophanous 
cuticle,  banded  with  buff  at  the  periphery ;  spire  semiglobose,  apex 
obtuse,  whorls  41,  convex,  rather  rapidly  increasing;  suture 
margined,  almost  crenulated  ;  finely  plicate-striate,  the  last  whorl 
tumid,  base  around  the  columella  denuded,  shining,  blackish. 
Aperture  little  oblique,  excised  circular;  peristome  little  expanded, 
a  trifle  reflexed,  thin  for  this  genus,  blackish ;  columella  slightly 
curved,  white,  much  dilated  above.  (Moll.) 

Malunu,  Prov.  Isabela,  northern  Luzon. 

C.  (Callicochlias)  erythrospira  v.  MOLL.  Nachr.  D.  M.  Ges.  1890, 
p.  204. 

C.  GENERALIS  Pfeiffer.     PL  52,  fig.  22. 

Imperforate,  globose  turbinate,  striatulate  and  sculptured  with 
oblique  wrinkles  especially  beneath ;  not  shining  ;  white  under  a 
buff-greenish  cuticle.  Spire  convex-conoid,  rather  obtuse;  suture 
light,  sub  margined.  Whorls  4£,  a  little  convex,  rapidly  increasing, 
the  last  little  descending  in  front,  encircled  above  the  periphery  with 
a  rather  wide,  flat  ridge ;  base  little  convex.  Columella  slightly 


COCHLOSTYLA-CALOCOCHLEA. 

oblique,  compressed,  excavated,  white.     Aperture  diagonal,  obliquely 
truncate-oval,  milk-white  inside;  peristome thin,  margins  conniving, 
the  right  one  scarcely  expanded,  basal  thickened,  narrowly  reflexed, 
forming  an  obtuse  angle  with  the  columella. 
Alt.  40,  diam.  57  mill.     (Pfr.) 

Philippines. 

H.  generate  PFR.,  P.  Z.  S.  1854,  p.  123  ;  Novit.  Conch,  i,  p.  2, 
t.  1,  f.  6 ;  Monogr.  ii,  p.  206. — REEVE,  Conch.  Icon.,  f.  1349. 

C.  CIIRVSOCHILA  Sowerby.     PL  34,  figs.  9,  10. 

I  in  perforate,  globose-subdepressed,  solid;  surface  delicately 
striate-obliquely,  having  microscopic  spiral  lines  above,  and  some- 
times obscurely  malleated.  Bright  greenish-yellow,  the  earlier  whorls 
tinged  with  rufous  ;  suture  edged  with  a  white  line ;  no  hydrophanous 
cuticle  ;  lip  broadly  reflexed,  yellow. 

The  spire  is  somewhat  conoid,  apex  very  obtuse,  whorls  nearly  5, 
the  last  slightly  descending  and  then  ascending  in  front.  Aperture 
large,  white  inside  ;  lip  broadly  expanded,  yellow;  columella  thick, 
yellow,  expanding  in  a  concave  white  callus. 

Alt.  39,  diam.  43  mill. 

Calayan,  Philippines. 

H.  chrysocheila  SOWB.,  P.  Z.  S.  1841,  p.  3. — PFR.,  Monogr.  i,  p. 
248  ;  Conchyl.  Cab.,  p.  275,  t.  44,  f.  5,  6. — REEVE,  Conch.  Syst.  ii, 
p.  163,  f.  1  ;  Conch.  Icon.,  f.  25. —  Cochlostyla  chrysochila  SEMPER, 
Reis.,  p.  175. — HIDALGO,  Journal  de  Conchyl.  1887,  p.  125. 

A  large,  smooth  yellow  species,  altogether  lacking  hydrophanous 
cuticle,  and  with  a  yellow  lip. 

(3).     Group  of  C.  polillensis. 

<  .  POLILLENSIS  Pfeiffer.     PI.  32,  figs.  43-50. 

I  in  perforate,  globose-conoid,  somewhat  solid,  nearly  smooth; 
ground-color  varying  from  light-yellow  to  reddish-chestnut,  plain  or 
having  2  to  4  band-  :  covered  with  a  light  brown  or  creamy 
hydrophanous  cuticle,  which  generally  forms  a  finely  checkered 
pan. TII  without  any  conspicuous  bands.  Columella  having  a  strong 
{<>/<!  near  //*  iii.wrlion. 

Tin-  contour  varies  much.  Whorls  5-j,  the  last  scarcely  descend- 
in.-- in  front.  Aperture  oblique ;  lip  reflexed  and  somewhat  thick- 


COCHLOSTYLA-CALOCOCHLEA.  139 

ened,  white  or  flesh-colored  ;  columella  subvertical,  inserted  deeply 
in  the  base,  and  having  a  strong  lobe  or  fold  above. 
Alt.  44,  diam.  42  mill,  (average  specimen.) 

Island  of  Polillo  off  the  E.  coast  of  S.  Luzon. 

H.  polillensis  PFR.,  P.  Z.  S.  1861,  p.  190  ;  Novit.  Conch.,  p.  233, 
t.  60,  f.  1,  2;  Monogr.  v,  p.  281.— H.  portei  PFR.,  P.  Z.  S.  1861,  p. 
191 ;  Novit.  Conch.,  t.  60,  f.  7-11.— H.  ajax  PFR.,  P.  Z.  S.  1861,  p. 
191  ;  Novit.  Conch.,  t.  60,  f.  3.—H.  hector  PFR.,  P.  Z.  S.  1861, 
p.  191. 

This  is  a  large,  globose-conoid  species,  with  finely  checkered 
cuticle. 

Color-var.  polillensis  (typical ;  figs.  43,  44)  has  a  brownish-yellow 
ground-color,  bandless,  or  with  very  faint  sutural  and  columellar 
bands. 

Color-var.  portei  (figs.  47,  50)  is  very  light  greenish-yellow,  with 
a  distinct  band  at  suture  and  umbilicus. 

Color-var.  ajax  (figs.  45,  46, 48)  has  a  dark  reddish-brown  ground- 
color. 

Color-var.  hector  (fig.  49)  has  four  dark  bands. 

Transition  forms  are  not  uncommon.  The  large  series  before  me 
confirms  Pfeiffer's  judgment  in  uniting  the  several  races. 

C.  ANDROMACHE  Pfeiffer.     Unfigured. 

Imperforate,  subglobose,  rather  thin,  striatulate,  chestnut  colored, 
little  shining.  Spire  convex,  apex  minute.  Whorls  4-4o,  moderately 
convex,  the  upper  white,  penultimate  tawny,  the  last  globose,  not 
descending,  sometimes  encircled  by  light  spiral  line.  Columella 
white,  subvertical,  compressed.  Aperture  oblique,  lunate-rotund, 
inside  pearly ;  peristome  simple,  whitish,  everywhere  narrowly  ex- 
panded, margins  converging,  the  right  margin  flexuous. 

Alt.  28,  greater  diam.  37,  lesser  30  mill.     (PfV.) 

Polillo,  Philippines. 

H.  andromache  PFR.,  P.  Z.  S.  1861,  p.  191  ;  Monogr.  v,  p.  282. 
Probably  a  variety  of  C.  polillensis. 

C.  PERAFFINIS  Pilsbry.     PI.  54,  figs.  47,  48. 

Imperforate,  globose-conoid,  solid,  having  low  regular  wrinkles  of 
growth  and  on  the  spire  faint  spiral  strue.  Ground-color  brownish- 
yellow  with  inconspicuous  darker  oblique  streaks,  and  having  a  nar- 
row black-chestnut  band  at  the  suture  and  around  the  columel/a. 


140  COCHLOSTYLA-CALOCOCHLEA. 

Hydrophanous  cuticle   disposed  in  dose  narrow  bands  and  lines, 
closely  interrupted  by  the  wrinkles  of  growth.     Earlier  whorls  whitish. 

Spire  convex-subconoid,  obtuse ;  whorls  4f ,  the  last  slowly  and 
-lightly  descending.  Aperture  large,  oblique,  white  or  nearly  so  in- 
side ;  peristome  broadly  reflexed,  rather  thick,  its  face  and  reverse 
irhite.  Columella  wide,  heavy,  oblique,  white. 

Alt.  38,  greater  diam.  41,  lesser  33  mill. 

Alt.  31,  greater  diam.  32  mill. 

Philippines. 

This  species  is  allied  to  C.  polillensis  and  C.  codonensis.  It  is  a 
distinctly  solider  shell  than  the  first,  with  a  much  broader  lip  and 
heavier  columella.  It  is  less  elevated  than  C.  codonensis,  with  an 
entirely  different  color-pattern,  the  lip  not  black-edged,  the  columella 
not  roseate,  etc. 

C.  CODONENSIS  Hidalgo.     PI.  53,  fig.  37. 

Imperforate,  subglobose,  solid,  obsoletely  sculptured  with  wrinkles 
of  growth  ;  shining  and  tawny,  with  a  black  columellar  band,  under 
an  opaque  white  more  or  less  interrupted  or  broadly  streaked 
hydrophanous  cuticle. 

Spire  obtuse,  whitish  or  rosy ;  whorls  44  to  5,  a  little  convex, 
rather  rapidly  widening,  the  last  large,  very  convex,  obliquely  some- 
what descending  in  front ;  columella  long,  suboblique,  rather  flat, 
rose  colored,  the  callus  becoming  outwardly  bluish-white.  Aperture 
ample,  oblique,  lunate-oval ;  peristome  widely  expanded  and  re- 
flexed  ;  its  inner  part,  with  the  aperture,  whitish-blue,  its  outer  edge 
intense  black.  (Hid.) 

Alt.  42,  greater  diam.  42,  lesser  35  mill. 

Codon,  Island  of  Catanduanes. 

C.  codonensis  HID.,  Journ.  de  Conchyl.  1888,  p.  312 ;  I.  c.  1889, 
t.  14,  f.  4. 

This  species  is  allied  to  C.  polillensis  and  its  varieties,  but  the 
aperture  is  relatively  higher,  the  columella  roseate  and  straighter, 
the  peristome  more  reflexed,  and  black  on  the  outside  ;  the  epidermis 
has  a  different  aspect  and  the  last  whorl  is  more  oblique,  etc.  (Hid.) 

< '.  DECIPIKNS  Sowerby.     PI.  21),  figs.  10,  11. 

Imperforate,  rather  thin, globose-depressed  or  subglobose;  surface 
nearly  smooth,  the  growth-lines  light,  the  earlier  whorls  seen  to  be 
ally  siriate  under  a  lens.     Straw-yellow,  with  irregular  oblique 


COCHLO8TYLA-CALOCOCHLEA.  141 

streaks  of  white  hydrophanous  cuticle,  and  with  a  peripheral  (some- 
times also  a  basal)  interrupted  band  of  the  same,  these  hydrophanous 
markings  are  usually  partly  lost  in  adult  shells. 

Whorls  4*,  the  last  a  little  descending  in  front.  Aperture  large  ; 
lip  well  expanded  subreflexed,  thin,  white ;  columella  arcuate, 
somewhat  sinuous.  Alt.  33,  diam.  35  mill. 

Luzon  and  Capul,  Philippines  (Cuming.) 

H.  decipiens  SOWB.,  P.  Z.  S.  1840,  p.  96. — PFR.,  Monogr.  i,  p. 
251  ;  Couchyl.  Cab.,  p.  277,  t.  45,  f.  5,  6.— RVE.,  f.  23. 

A  thin  species,  having  oblique  ragged  streaks  and  a  narrow  band 
of  white  cuticle.  Our  specimens  are  more  globose  than  the  figures 
on  pi.  29. 

(4).     Group  of  C.  zonifera. 

C.  ZONIFERA  Sowerby.     PI.  35,  figs.  19,  20,  22. 

Imperforate,  solid,  depressed-globose.  Ground-color,  under  the 
hydrophanous  cuticle,  light  greenish-buff  with  (typically)  a  broad 
peripheral  girdle  of  dark  brotvn,  the  upper  whorls  nearly  white. 
Hydrophanous  cuticle  cream- white  forming  two  wide  uninterrupted 
bands,  one  above  the  other  below  the  peripheral  dark  zone ;  the  base 
below  the  lower  band  has  no  hydrophanous  cuticle ;  above  the  upper 
band  it  is  present  in  irregular  patches,  cut  by  a  group  of  narrow  spiral 
lines  which  revolve  about  the  middle  of  the  upper  surface. 

Whorls  4-4 2,  the  last  wide  and  slightly  descending  in  front. 
Aperture  quite  oblique,  white  (or  flesh-tinted)  inside ;  lip  white, 
roundly  reflexed ;  columella  oblique,  straight,  deeply  inserted. 

Alt.  30,  diam.  37  mill. 

Leyte ;  La  Laguna,  Luzon;  Loquilocon,  Samar ;  Visayes ;  Cat- 
anduanes,  E.  coast  of  Mindanao ;  Siargas,  in  the  Strait  of  Surigao ; 
Philippines. 

H.  zonifera  SOWB.,  in  sched.  Cuming. — PFR.,  Monogr.  i,  p.  251  ; 
Conchyl.  Cab.,  p.  280,  t.  48,  f.  3,  4.— REEVE,  Conch.  Icon.,  t.  9.  f. 
3. —  Cochlostyla  zonifera  MARTENS,  Ostas.  Landschn.,  p.  94,  with  var. 
purpurascens  and  modestior. —  Cochlostyla  zonifera  SEMPER,  Reis. 
Phil.,  p.  177,  t.  18,  f.  9  (anatomy). — HIDALGO,  Journ.  de  Conchyl. 
1887,  p.  129. 

A  globose-depressed  species  having  a  broad  zone  of  creamy  cuticle 
on  either  side  of  a  darker  peripheral  zone,  the  upper  surface  having 
irregular  patches  of  creamy  cuticle  cut  by  a  group  of  spiral  lines. 


142  <  OCHLOSTYLA-CALOCOCHLEA. 

Yur.  CIRCE  Pfr.      PI.  53,  figs.  28,  29. 

Ground-color  brown,  with  an  ill-defined  chocolate  band  at  the 
periphery,  the  whorls  of  the  spire  and  the  interior  of  the  aperture 
purple-tinged.  Otherwise  like  the  type. 

Is.  Samar  and  Leyte. 

II.  dree  PFK.,  Monogr.  iii,  p.  645.  —  RVE,  Conch.  Icon.,  f.  1025.  — 
Cochlostyla  zonifera  var.  purpurascens  MART.,  Ostas.  Landschn.,  p. 
95. 

Another  variety  is  represented  on  pi.  53,  figs.  '30,  31.  Two  broad 
hands  of  cream-white  cuticle  are  present  as  well  as  traces  of  a  sutural 
band,  but  there  is  a  wide  white  zone  above,  much  as  in  the  following 
variety,  which  is,  however,  a  much  more  globose  shell. 

Var.  FARALEUCA  Pilsbiy.     PL  53,  figs.  32,  33. 

Globose  ;  light-yellow  with  a  broad  w7hite  band  above.  Columella 
wide  and  strongly  curved  inward.  Alt.  29,  diam.  32J  mill. 

C.  CORONADOI  Hidalgo.     PL  34,  figs.  5,  6  ;  pi.  35,  fig.  21. 

Imperforate,  solid,  depressed,  the  spire  low-conoidal,  ground-color 
(under  the  hydrophanous  markings),  greenish-yellow,  becoming  brown 
on  the  base;  having  (normally)  two  narrow  dark  chestnut  bands, 
one  below,  the  other  above  the  periphery  and  ascending  the  spire, 
but  the  bands  arc  often  absent.  Hydrophanous  cuticle  torn  into 
HIII-  row,  oblique  shreds,  at  least  above,  not  forming  a  series  of  blotches 
below  the  xuture;  disposed  in  two  broad  zones,  one  below,  the  other 
above  a  wide  peripheral  zone  of  the  ground-color;  the  upper  zone 
bounded  above  by  a  narrow  dark  band. 

Whorls  4-j,  rather  flattened,  the  sutures  very  shallow,  becoming 
impressed  around  the  last  whorl,  which  descends  gradually  in  front. 
Aperture  quite  oblique,  white  inside;  lip  broadly  reflexed,  white; 
eolumrlla  deeply  entering,  more  or  less  arcuate,  its  face  deeply  ex- 
•  •avated.  Alt.  31,  diam.  44  mill. 

Leyte;   Catanduanes. 


<•<»-<>,  Ki<loi  1  1  1  1).,  Jon  ri)  .  dc  Conchyl.  1868,  t,  13,  f.  5; 
/.  c.  1887,  p.  1  .">•_>.—  //.  cailliaudi  var.,  I  Vis.,  Conchyl.  Cab.,  t.  42,  f. 

3,    I      nut    //.  cui/Haudi  I  Hi.) 

A  lar-e,  solid  sj.ccios,  allied  to  (\  zonifera,  but  larger,  with  differ- 
shaped    Columbia   and    differently    patterned    hydrophanous 
cuticle. 


COCHLOSTYLA-CALOCOCHLEA.  143 

Yar.  PULCHRA  Pilsbry.     PI.  54,  figs.  43,  44. 

I  m  perforate,  globose-depressed,  solid.  Yellowish -brown,  slightly 
streaked  obliquely  with  brown,  and  having  numerous  rather  narroiv 
bands  of  hydroplianous  cream-white  spots,  a  supra-peripheral  band 
sometimes  wider.  Whorls  of  the  spire  having  a  narrow  brown  *piral 
band. 

Spire  convex,  obtuse,  the  earlier  two  whorls  pink,  following  whorl 
white.  Whorls  5,  the  last  scarcely  descending.  Aperture  large, 
oblique,  white  inside  ;  lip  broadly  reflexed,  thick,  white  on  both  face 
and  reverse.  Columella  arcuate,  deeply  entering,  its  face  excavated. 

Alt.  37,  diam.  45  mill. 

Albay,  Philippines. 

A  more  elevated  shell  than  C.  coronadoi,  with  a  different  pattern 
of  cuticle.  A  part  of  the  hydroplianous  bands  are  doubtless  rubbed 
off  of  the  specimens  before  me. 

C.  NORRISII  Sowerby.     PI.  31,  figs.  41,  42. 

Imperforate,  globose-depressed,  solid,  obsoletely  malleated, 
glabrous,  tawny-citrinous,  having  opaque  white  zones  and  under  the 
suture  irregular  white  spots.  Spire  serniglobose.  Whorls  41,  con- 
vex, the  last  excavated  at  the  broad,  callous,  arcuately  prolonged 
columella.  Aperture  limate-subcircular,  white  inside  ;  peristome 
thick,  very  broadly  expanded. 

Alt.  28,  greater  diam.  42  mill.     (Pfr.) 

Luzon  and  Mindanao  (Pfr.)  ;  Leyte  and  Catanduanes  (Hidalgo), 
Philippines. 

H.  norrini  SOWB.  (in  sched.  Cuming). — PFR.,  P.  Z.  S.  1842,  p. 
85  ;  Monogr.  i,  p.  250  ;  Conchyl.  Cab.,  t.  46,  f.  1,  2. — REEVE,  Conch. 
Icon.,  f.  21. —  C.  norrisii  HID.,  Journ.  de  Conchyl.  1887,  p.  133. 

I  have  not  seen  this  species.  It  seems  very  closly  allied  to  C. 
zonifera,  but  the  lip  is  perhaps  broader  and  the  columella  more 
oblique.  Reeve's  figure  (fig.  42)  may  not  be  the  same. 

C.  DAMAHOYI  Pfeiffer.     PL  37,  figs.  40,  41,  42. 

Imperforate,  solid,  depressed-globose ;  having  superficial,  scarcely 
obvious  spiral  liruke  and  on  the  latter  part  of  the  last  whorl  lightly 
obliquely  malleated.  Ground-color  rich  brown,  with  two  broad 
darker  zones,  one  above,  the  other  below  a  wide  peripheral  zone  of 
greenish-buff.  Hydroplianous  cuticle  cream-white,  strongly  contrast- 
ing Avith  the  dark  back-ground,  and  forming  (over  this  ground-color) 


1 44  <  'OCHLOSTYLA-CALOCOCHLEA. 

a  wide  subsutural  and  a  wide  peripheral  zone  (one  or  two  narrow 
bands  between  them),   and  on  the  base  several  wide  and   narrow 

bands. 

Spire  low,  inconspicuous;  whorls  41,  little  convex,  the  last  but 

little  descending  in  front.     Aperture  large,  banded  inside,  perislome 
white,  broadly  reflexed ;    columella  oblique,  wide,  nearly  straight. 
Alt.  33-],  diam.  45  mill. 

Ca  lay  an,  Philippines. 

H.  damahoyi  PFR.,  P.  Z.  S.  1856,  p.  328;  Monogr.  iv,  p.  210; 
Novit.  Conch,  iv,  p.  8,  t.  Ill,  f.  1-4. —  Cochlostyla  damahoyi  SEMPER, 
Reis.,  p.  175,  t.  18,  f.  8  (anatomy.) 

Allied  to  C.  zonifera,  but  differs  markedly  in  the  arrangement  of 
colors  and  the  obscure  malleation.  The  two  blackish  bands  are 
sometimes  absent  (fig.  42.) 

C.  CAILLIAUDI   Deshayes.     PL  50,  figs.  2,  3  ;  pi.  52,  fig.  26,  27. 

I m perforate,  solid,  large,  globose-depressed-conoid:  Surface  finely 
and  obscurely  spirally  lirulate,  the  last  whorl  quite  obviously  sculpt- 
ured with  obliquely  forward-descending  malleations  forming  narrow 
ridges,  becoming  concentric  beneath.  Ground-color  clear  greenish- 
yellow,  having  a  dark  zone  below  and  another  above  the  periphery. 
Hydrophanous  white  cuticle  forming  numerous  narrow  encircling 
bands,  mostly  worn  off  in  front  of  the  aperture. 

Spire  rather  conoidal ;  whorls  5,  but  slightly  convex,  the  suture 
superficial;  last  whorl  slightly  descending  in  front.  Aperture  large, 
white  and  faintly  showing  the  bands  inside ;  peristome  broadly  re- 
flexed,  recurved,  white ;  columella  oblique,  nearly  straight. 

Alt.  43,  diam.  45  mill. 

Camaruan,  prov.  of  Camarines  Sur,  Luzon  (Hidalgo.) 

H.  cailliaudi  DH.  in  Guerin's  Mag.  de  Zool.  1839,  t.  5. — PFR., 
Monogr.  i,  p.  246.— REEVE,  Conch.  Icon.,  f.  ^.—  Cochlostyla 
t-tii/fi'muli  HIUALCJO,  Journal  de  Conchvliologie  1887,  p.  131. — 

•  ^5  i 

1 1 dix  ferruginea  LEA,  Trans.   Amer.  Philos.  Soc.,  2d  Ser.,   vii,  p. 
464,  t.  12,  f.  17  (1841.) 

A  large,  solid  shell,  decidedly  malleated,  light  yellow  with  two 
brown  bands,  the  whole  covered  with  numerous  narrow,  interrupted 
hydrophanous  bands.     The  typical  form  is  shown  in  fig.   3,  of  pi. 
50,   the  specimen   figured   being  nearly  denuded  of  hydrophanous. 
cuticle  in  front. 


COCHLOSTYLA-CALOCOCHLEA.  14.) 

The  shell  resembles  the  group  of  C.  latitans,  C.  cryptica,  etc.,  in 
sculpture  and  shape  of  the  mouth,  but  it  is  in  coloring  and  shape 
of  the  spire  more  allied  to  C.  coronadoi,  etc. 

Color-var.  ferruginea  Lea.  (pi.  52,  fig.  27).  Unicolored,  brown- 
ish. Lea  included  the  true  cailliaudi  also  in  his  diagnosis,  but  the 
name  may  be  retained  in  a  restricted  sense  for  the  pale  reddish- 
brown  form. 

Color-var.  (pi.  52,  fig.  26).  Shell  smaller,  with  accessory  bands 
at  suture  and  columella. 

C.  MICROSPIRA  Pfeiffer.     PI.  53,  figs.  34,  35,  36. 

Imperforate,  solid,  globose-depressed  ;  surface  smooth,  with 
delicate  growth-lines,  the  spiral  striation  almost  or  entirely  obsolete, 
the  last  whorl  sometimes  slightly  malleated.  Yellow,  with  three 
dark  chestnut  or  purple  bands,  one  at  the  narrowly  white-margined 
suture,  one  at  the  periphery,  and  a  third  surrounding  the  columella. 
No  traces  of  hydrophanous  cuticle. 

Spire  short,  obtuse  ;  sutures  superficial ;  whorls  4s,  the  last  a  little 
descending  in  front.  Aperture  large,  white  inside  and  often  show- 
ing the  bands;  lip  reflexed,  white,  frequently  narrowly  edged  with 
brown ;  columella  oblique,  white,  subtruncate  below.  Earlier 
whorls  white  or  reddish. 

Alt.  37,  diam.  42  mill. 

Alt.  35,  diam.  49  mill. 

Philippines. 

H.  microspira  PFR.,  P.  Z.  S.  1853,  p.  127  ;  Monogr.  iv,  p.  213.- 
REEVE,  Conch.  Icon.,  f.  1268. 

A  large  and  solid  trifasciate  species,  somewhat  allied  to  C.zonifera 
var.  circe,  but  decidedly  heavier,  and  of  a  different  tone  of  color. 
The  variations  in  banding  are  shown  by  the  figures,  selected  from  a 
large  suite  of  specimens. 

C.  HEMispir^ERiON  Pfeiffer.     PI.  54,  figs.  45,  4(5. 

Imperforate,  serniglobose,  solid,  obliquely  striated,  and  under  a 
lens  seen  to  be  very  closely  spirally  sculptured;  shining,  buff,  with 
chestnut-tawny  bands.  Spire  convex,  apex  obtuse.  Whorls  4^,  a 
little  convex,  rapidly  increasing,  the  last  scarcely  descending,  the 
periphery  obtusely  angled  and  sometimes  irregularly  obliquely 
malleated-plicate  ;  base  subplanulated.  Aperture  oblique,  lunate- 
oval,  whitish  within  ;  columella  somewhat  sloping,  wide,  basal  mar- 
10 


14»i  COCHLOSTYLA-CALOCOCHLEA. 

gin  narrowly  reflexed,  forming  an  obtuse  angle  with  the  columella. 
(P/K)     Alt.  27,  diam.  43  mill. 

Philippines. 

H.  hcmi*iih<irinn  PFR.,  Zeitschr.  f.  Mai.  1851,  p.  15;  Monogr, 
iii,  p.  J86;  Conchyl.  Cab.,  p.  407,  t.  14fi,  f.  1,  2. 

C.  SAMARENSIS  Semper.     PI.  34,  figs.  1,  2  ;  pi.  54,  figs.  41,  42. 

I  m  per  for  ate,  globose,  striatulate,  chestnut  or  tawny,  apex  very 
obtuse,  usually  white,  sometimes  ornamented  with  two  chestnut 
bands  and  an  obsolete  white  band  at  the  periphery.  Spire  globose  ; 
whorls 4,  rapidly  increasing,  the  last  very  large;  columella  oblique; 
excavated,  white.  Aperture  lunate-circular,  milky  or  brown  ;  per- 
istome  widely  reflexed,  black-edged.  ($.) 

Alt.  34,  greater  diam.  46,  lesser  35  mill. 

Pampan,  Samar ;  San  Fernando,  Sibuyan. 

C.  samarensis  C.  SEMPER,  Reis.,  p.  179,  t.  10,  f.  9  (shell)  ;  t.  13, 
f.  15;  t.  18,  f.  12  (anatomy). — HIDALGO,  Journ.  de  Conchyl.  1887, 
p.  136,  t.  (>,  f.  1,  2. 

The  typical  form  is  thin,  translucent,  sometimes  with,  sometimes 
without  the  narrow  chestnut  bands.  A  brownish  tract  surrounds 
the  white  columella,  and  the  reflexed  peristome  is  brownish-black. 
Upon  all  3  examples  there  is  a  wide  whitish  and  lacerated  band  at 
the  periphery,  formed  of  the  otherwise  inconspicuous  hydrophanous 
cuticle.  ($.) 

Hidalgo  describes  three  color-varieties: 

1.  Shell  chestnut,  unicolored  (pi.  54,  fig.  42.) 

2.  Shell  tawny,  columellar  area  and  suture  brov\7n. 

3.  Like  the  last,  but  periphery  with  a  chestnut  band  (pi.  54,  fig. 
41.) 

S.-mper  considered  the  following  form  a  variety,  but  Hidalgo  says 
that  it  does  not  belong  to  the  same  species. 

Var.  (pi.  34,  figs.  3,  4).  Shell  much  thicker  ;  peristome  bluish- 
white;  no  dark  columellar  area. 

C.  < -in  <  OMELOS  Sowerby.     PI.  55,  figs.  51-55. 

Impi-rforate,  globose,  rather  solid;  lightly  obliquely  striated. 
<  iruimd  colo i-  varying  from  dark  chestnut-brown  to  olive-yellow, 

terally  unicolored,  but  sometimes  banded  ;  suture  white-margined. 
Apical  whorls  pink  or  purple.  Hydrophanous  cuticle  thin,  evanes- 
cent, often  cut  by  spiral  dark  lines. 


COCHLOSTYLA-CALOCOCHLEA.  147 

Spire  low-conoid,  very  obtuse ;  whorls  4-2,  the  last  slightly 
descending.  Aperture  oblique,  rounded,  white  or  tinted  inside  ;  lip 
well  reflexed,  whitish  with  a  brown  edge. 

Alt.  33,  diam.  35  mill. 

Alt.  39,  diam.  39  mill. 

Tablas,  Sibuyan  and  Samar,  Philippines. 

H.  coccomelos  SOWB.,  P.  Z.  S.  1840,  p.  167. — RVE.,  Conch.  Syst. 
ii,  t.  164,  f.  9. —  Cochlostyla  coccomelos  HIDALGO,  Journ.  de  Conch. 
1887,  p.  134. — H.  speciosa  PFR.,  Monogr.  i,  p.  248;  Conchyl.  Cab., 
p.  311,  t.  46,  f.  7-9,  and  of  authors  generally,  not  H.  speciosa  Jay. 

A  very  variable  species.  The  following  varieties  are  enumerated 
by  Hidalgo : 

1.  Pale  buff,  the  spire  and  lip  white.     An  albino  form. 

2.  Olivaceous,  3  banded  with  chestnut. 

3.  Olivaceous,  columella  surrounded  by  a  chestnut  zone. 

4.  Olivaceous,  not  banded. 

5.  Greenish,  not  banded. 

6.  Tawny,  spire  paler. 

7.  Reddish,  3-banded  with  chestnut. 

8.  Blackish,  spire  paler. 

9.  Chestnut,  becoming  rosy  on  the  spire  (figs.  52,  5,3.) 

Dr.  Jay's  collection  is  now  in  the  American  Museum  of  Natural 
History,  Central  Park,  New  York  City.  I  am  indebted  to  Prof. 
R.  P.  AVhitfield  for  the  opportunity  of  examining  the  type  of 
H.  speciosa  Jay,  contained  therein.  It  is  a  C.  dubiosa  (q.  v.) 

C.  PONDEPvOSA  Pfeiffer.     PI.  48,  fig.  65. 

Imperforate,  globose-depressed,  tumid,  solid,  ponderous ;  rather 
smooth  (incremental  and  concentric  strise  very  close,  scarcely 
visible) ;  white,  the  base  covered  with  a  shining,  pale  corneous 
cuticle.  Whorls  4-j,  a  little  convex,  the  last  subangulated,  some- 
times obsoletely  marked  with  a  white  hydrophanous  band  below  the 
angle.  Columella  straight,  wide,  very  oblique.  Aperture  subauri- 
form,  white  inside  ;  peristome  broadly  reflexed. 

Alt.  27,  greater  diam.  46,  lesser  34  mill.     (Pfr.~) 

Prov.  Ilocou,  Luzon,  Philippines. 

H.  ponderosa  PFR.,  P.  Z.  S.  1845,  p.  38  ;  Monogr.  i,  p.  259 ;  Con- 
chyl. Cab.,  p.  215,  t.  108,  f.  3,  4. — REEVE,  Conch.  Icon.,  f.  47. 


1  4S  COCHLOSTYLA-CALOCOCHLEA. 

(5).     Group  of  C.  mindanaensis. 

(  '.  MINDANAENSIS  Sowerby.     PI.  47,  figs.  61,  62. 

Imprrforate,    solid,     depressed-globose-conical,    large.       Surface 


lv  obliquely  striated.  OjHtque  brownish-yellow,  marked  all  over 
with  *iint.?  <nid  zigzag  streaks  of  dark  reddish-brown,  having  a  band 
of  hydrophanous  whitish  cuticle  immediately  below  the  periphery. 

The  spire  is  low-conoidal,  apex  very  obtuse,  the  earlier  whorls 
flesh-colored,  with  a  dark  line  just  above  the  suture.  The  last  whorl 
H  vrrv  obtusely  subanirular  at  the  periphery.  Aperture  oblique, 
livid  in>ide;  lip  reflexed,  flesh-colored,  darker  toward  the  outer 
<  dge  ;  columella  flesh-colored,  oblique,  subtruncated  below. 

Alt.  44,  diam.  53  mill. 

Alt.  41  ,  diam.  45  mill. 

Is.  of  Mindanao  and  Camiguin,  Philippines. 

H.  mlndunai'-nsis  SOWB.  in  sched.  Cuming.  —  PFR.,  P.   Z.  S.  1842 
p.  85;  Monogr.  i,  p.  245  ;  Conchyl.  Cab.,   p.  268,   t.  41,  f.  5,   6.- 
REEVE,  Conch.  Icon.,  f.  32.  —  H.  mindana  RVE.,  Conch.  Syst.  ii,  t. 
1(54,  f.  8.  —  C/iromocochlea  mindorcnia  HARTM.,  Gast.  Schw.  i,  p.  139, 
t.  42.  —  Cochlostyla  mindanaensis  SEMPER,  Reis.,  p.  187. 

A  peculiarly  mottled  species,  very  distinct  from  other  known 
firms.  * 

(  '.  HARFORDII  Sowerhy.     PI.  55,  figs.  49,  50. 

I  MI  perforate,  depressed-globose-turbinate,  large,  solid  ;  obliquely 
lightly  striated.  Light  brown  (when  worn,  light  green),  blotched 
irregularly  with  dark  chestnut,  having  numerous  narrow  light  bands 
above  and,  a  broad  light  band,  covered  with  creamy  hydrophanous 
cuticle  just  below  the  periphery. 

Spire  low-conoidal,  apex  very  obtuse;  whorls  5,  the  last  a  trifle 
defending  in  front.  Aperture  oblique,  \yhite  within  ;  lip  expanded, 

'M'llr\cd,  white.     Columella  steeply  sloping,  white. 

All.  4!i,  diam.  57  mill,  (specimen.) 

Alt.  4X,  diam.  70  mill.  (Pfr.) 

Negros,  Philippines. 

If.  Itarfordii  BROD.,  P.  Z.  S.  1840,  p.  123.—  REEVE,  Conch.  Syst. 
ii,  t.  Hi5,  f.  15;  Condi.  Icon.,  f.  3d.—  PER.,  Monogr.  i,  p.  244; 
Conchyl.  Cab.,  p.  I'M,  t.  42,  f.  1,2. 

Allied  \<>  C.  miml'tiKii'ii.*;*,  but  larger  and  differently  colored. 


COC'HLOSTYL,  A-CALOCOCHLEA .  1  49 

(6).      Group  of  C.  melanochila. 

C.  PAN  Broderip.     PI.  3.3,  figs.  56,  56,  57. 

Imperforate,  globose-depressed-conoid,  solid,  smooth,  lusterless ; 
ground-color  chocolate-brown  below,  yellowish-brown  above  the  periph- 
ery ;  suture  narrowly  margined  with  dark  brown,  covered  with  a 
hydrophanous  creamy  cuticle  which  is  everywhere  conspicuously 
banded  with  rust-brown  above  and  chocolate  below. 

The  spire  is  rather  con oidal,  apex  obtuse;  whorls  5],  the  upper 
nearly  flat,  the  last  convex,  a  little  descending  in  front.  Aperture 
tinted  inside,  not  showing  bands ;  peristome  reflexed,  more  or  less 
conspicuously  dark  edged  ;  columella  oblique,  nearly  straight  or 
obsoletely  bisinuate. 

Alt.  43,  diarn.  42  mill,  (average  size.) 

BoJwl  and  Panglao,  Philippines. 

H.pan  BROD.,  P.  Z.  S.  1841,  p.  23.— REEVE,  Conch.  Syst,,  t.  165, 
f.  19 ;  Conch.  Icon.,  f.  24.— PER.,  Monogr.  i,  p.  245  ;  Conchyl.  Cab., 
p.  265,  t.  39,  f.  5-7.— DELESSERT,  Rec.  de  Coq.,  t.  38,  f.  9. 

The  ground-color  is  much  as  in  Hadra  bipartita.  The  hydropha- 
nous cuticle  has  bands  of  rust-brown  as  well  as  zones  of  the  chocolate 
ground-color.  It  is  a  beautiful  shell,  exhibiting  but  little  variation. 

C.  DEPRESSA  Semper.     PI.  50.  figs.  4,  5,  6,  7,  8. 

Imperforate,  solid,  depressed  ;  surface  having  light  growth-lines 
and  very  minute,  subobsolete  spiral  strise.  Ground-color  dark  brown, 
having  a  darker  band  at  the  periphery  and  under  the  suture. 
Hydrophauous  cuticle  forming  a  subsutural  series  of  blotches,  a 
peripheral  zone  and  numerous  narrow  bands  on  the  base,  of  cream- 
white,  the  rest  of  the  surface  closely  lineate  with  brownish-white 
cuticle. 

The  spire  is  low,  rounded  ;  sutures  superficial ;  apex  obtuse. 
Whorls  4i,  the  last  obtusely  subangular  at  the  periphery,  not  in  the 
least  descending  in  front.  Aperture  oblique,  leaden-bluish  inside; 
lip  narrowly  expanded  and  subreflexed,  dark  brown;  columella 
wide,  flattened,  subtruncate  below7. 

Alt,  28,  diam.  40  mill. 

Mindanao;  Surigao,  Philippines. 

H.  lignaria  PER.,  P.  Z.  S.  1846,  p.  Ill  ;  Monogr.  i,  p.  259 ;  Con- 
chyl. Cab.,  p.  424,  t.  149,  f.  15. — REEVE,  fig.  40.  (not  Bulimus 
lignarius  PER.,  also  a  Cochlostykt). —  Cochlostyla  depressa  SEMPER, 
Reis.,  p.  180. — HIDALGO,  Jo  urn.  de  Couchyl.  1887,  p.  137. 


1")0  COCHLOSTYLA-CALOCOCHLEA. 

This  species  differs  from  C.  damahoyi  which  it  somewhat  resem- 
bles in  pattern  of  coloring,  in  its  constantly  brown  lip  and  more  de- 
pressed contour. 

Var.  perdepressa  (figs.  6,  7,  8).  Much  more  depressed  above  than 
the  type;  upper  surface  (under  the  hydrophanous  patches)  green- 
ish-yellow, base  dark  brown.  Dark  bands  at  suture,  periphery  and 
and  columella.  Alt.  25,  diam.  42  mill. 

C.  MELANOCHILA  Valenciennes.     PI.  30,  figs.  17,  18,  19. 

Imperforate,  solid,  depressed-globose ;  growth-lines  delicate ; 
spiral  stria?  subobsolete.  Dark  chestnut-brown,  with  a  wide  yellowish 
bond  above,  and  a  narrow  one  on  the  base;  apex  purple  or  pink. 
Hydrophanous  cuticle  brownish,  usually  lost  by  erosion  for  the 
greater  part. 

Spire  low,  apex  very  obtuse;  whorls  5,  the  last  a  little  descending 
in  front.  Aperture  oblique,  wide,  white  inside  ;  lip  reflexed,  dark 
brown ;  columella  white,  nearly  straight. 

Alt.  31,  diam.  44  mill. 

Alt.  34,  diam.  36  mill. 

Calapan,  Mindoro,  Philippines. 

H.  melanocheila  VAL,  in  Paris  Mus. — GRATELOUP,  Actes  Soc. 
Linn.  Bord.,  xi,  p.  397,  t.  4,  f.  12.— PFR.,  Monogr.  i,  p.  258  ;  Con- 
chyl.  Cab.  p.  276,  t.  44,  f.  9,  10.— REEVE,  Conch.  Icon.  f.  SQ.—H. 
In-nunea  SOWB.,  P.  Z.  S.  1841,  p.  40  (not  of  Anton). —  Cochlostyla 
melanocheila  HIDALGO,  Journ.  de  Conchyl.  1887,  p.  125. 

A  large  black-lipped  species,  intermediate  between  C.  pan  and 
C.  roissyana. 

( 5.  CHLOROCHROA  Sowerby.     PI.  37,  figs.  38,  39. 

Imperforate,  subglobose,  rather  thin,  tawny  or  pale  brown, 
M  reaked  and  marbled  with  white ;  covered  with  an  ashen-brown 
cuticle.  Spire  semiglobose.  Whorls  4o,  a  little  convex,  the  last 
obsoletely  angular  at  the  periphery.  Columella  oblique,  arcuate, 
denticulate.  Aperture  rounded-lunar  ;  peristome  subthickened,  ex- 
panded, brown.  Alt.  34,  greater  diam.  50,  lesser  42  mill.  (Pfr.) 

Camiguin  de  Mindanao  (Semper)  ;  Bohol  (Cuming),  Philippines. 

If.  chlorochroa  Sow.,  P.  Z.  S.  1841,  p.  2. — PER.,  Monogr.  i,  p. 
246;  Conchyl.  Cab.  p.  '214,  t.  44,  f.  1,  2.— REEVE,  f.  27.—  Cochlo- 

/'i  ctilorochroa  SKMI-KR,  Reis.,  p.  187. 


COCHLOSTYLA-CALOCOCHLEA.  151 

C.  MATRUELIS  Sowerby.     PI.  48,  figs.  70,  71. 

Imperforate,  depressed-globose,  rather  thin.  Ground-color  brown 
with  three  blackish  bands:  one  below  the  suture,  one  at  the  periph- 
ery, the  third  (occasionally  absent)  bounding  a  light  yellow  coin- 
mellar  tract  ;  apex  purple.  Hydrophanous  cuticle  very  thin,  but 
forming  an  opaque  creamy  band  at  suture  and  immediately  below 
the  periphery. 

Spire  low,  apex  very  obtuse.  Whorls  slightly  exceeding  4,  the 
apex  slightly  sunken  ;  last  whorl  descending  a  little  in  front.  Aper- 
ture oblique,  livid  and  showing  a  band  inside  ;  lip  rather  narrowly 
expanded,  brown  ;  columella  very  wide,  brown,  truncated  and  sub- 
dentate  below,  the  tooth  white.  Alt.  25,  diam.  33  mill. 

Mindanao,  Philippines. 

H.  matruelis  SOWB.  P.  Z.  S.  1841,  p.  24.  —  PFR.,  Monogr.  i,  p. 
259  ;  Conchyl.  Cab.  p.  282,  t.  46,  f.  10,  11  .—REEVE,  Conch.  Icon.  f. 
75.  —  DELESSERT,  Rec.,  t.  38,  f.  4. 

Somewhat  allied  to  C.  melanocheila  but  smaller,  the  columella  of 
a  different  form,  and  the  arrangement  of  colors  diverse. 

C.  DIFFICILIS  Pfeiffer.     PL  48,  fig.  72. 

Imperforate,  globose-depressed,  thin,  very  minutely  striated,  shin- 
ing, pale  tawny,  at  the  white  suture  banded  with  chestnut.  Spire 
small,  semiglobose.  Whorls  4s,  a  little  convex,  the  last  inflated,  at 
the  periphery  and  base,  banded  with  chestnut  ;  not  descending  in 
front.  Aperture  oblique,  tetragonal-lunar,  white  inside;  peristome 
subthickened,  narrowly  reflexed,  the  columellar  margin  straightened, 
much  dilated,  flat,  white.  Alt.  26,  greater  diam.  42,  lesser  34  mill. 


Philippines. 

H.  difficilis  PFR.,  P.  Z.  S.  1853,  p.  127;  Monogr.  iv,  p.  213.— 
REEVE,  Conch.  Icon.  f.  1269. 

C.  ROISSYANA  Ferussac.     PI.  30,  figs.  23-28. 

Imperforate,  subglobular,  solid,  compact.  Surface  lusterless, 
smooth.  Color  either  yellow,  variously  banded  ivith  chocolate-brown, 
or  chocolate  with  a  light  band  above;  the  inner  whorls  either  white 
or  purplish-blue.  The  hydrophanous  cuticle  is  thin,  light  brown, 
and  nearly  uniform,  but  it  is  usually  almost  entirely  worn  off. 

The  form  is  globose  and  compact  ;  growth-lines  light  ;  spiral  stri- 
ation  nearly  obsolete.  Color  very  variable.  Whorls  5,  the  last 


1 .-,-_'  COCHLOSTYLA-CALOCOCHLEA. 

>liirhtly  descending  in  front.     Aperture  oblique,  whitish  inside;  lip 
,iurroii-/i!  e.r/i'inded,  blackish-brown,  T&rely  white;    columella  white, 
wide,  short,  nearly  straight,  steeply  sloping. 
Alt.  •'>!,   liam.  :>2  mill,  (average  specimen). 

Jfindoro,  Philippines. 

H.  roi**if<ui<t  Vi-n.,  Hist,  t,  104,f.  2-3.— SOWB.,  P.  Z.  S.  1840,  p. 
101.— PFR.,  Monogr.  i,  p.  249  ;  Conchyl.  Cab.  p.  283,  t.  47,  f.  1-5  ; 
Novit.  Conch,  iii,  p.  4!>7,  t.  107,  f.  10,  11.— REEVE,  Conch.  Icon.  f. 
~:\,—  ('nchlostii/<t  roi#«ii«na  SEMP.,  Reis.,  p.  185. — HIDALGO,  Obras. 
Malacol.  i,  t.  51,  f.  1-7. — H.  rossiana  GRAY,  Ann.  of  Phil.  (n.  s.), 
ix,  p.  412. 

A  compact,  subglobular  species,  with  narrow,  dark  lip.  The 
typical  form  as  figured  by  Ferussac  is  yellow  with  dark  bands. 
'  Color-var.  lutea  Pfr.  (pi.  30,  figs.  23,  24).  Light  yellowish,  with 
a  lighter  band  above. 

Color-var.  subatra  (pi.  30,  fig.  28).  Body-whorl  dark  chocolate- 
brown,  a  light  band  appearing  on  the  penultimate  whorl. 

C.  CNFUSCATA  Albers.     PI.  30,  figs.  20,  21. 

Imperforate,  conoid-subglobose,  solid,  obliquely  striated,  obsoletely 
decussated  with  spiral  stride ;  tawny ;  spire  convex-conoid,  whitish 
above;  suture  white-margined.  Whorls  5,  convex,  the  last  rotund, 
shortly  descending  in  front,  radiated  and  obsoletely  banded  with 
l»rown;  columellar  area  chestnut.  Columella  somewhat  sloping, 
wide,  white,  subexcavated.  Aperture  oblique,  lunate-rounded,  milk- 
white  inside;  peristome  thick,  white,  narrowly  reflexed,  the  right 
margin  arched  forward.  (Pfr.  from  spec,  in  coll.  Albers.) 

Alt.  24,  greater  diam.  31,  lesser  27  mill. 

Habitat  supposed  to  be  Philippines, 

II.  iiij'iiafufti  ALII.,  Mai.  Bl.  i,  1854,  p.  214.— PFR.,  Novit.  Conch. 
i,  p.  41  i it.  ll,f.7,  8;  Monogr.  iv,  p.  211. 

C.  M  \IMTKNSIS  Hidalgo.     PI.  55,  fig.  5(5. 

[raperforate,  subglobose,  rather  solid,  smooth,  chestnut  colored, 
under  an  ashen-brown  hydrophanous  cuticle,  transversely  darker, 

Unrated,  and  at  the  periphery  having  a  white  baud. 
moderate,  apex  obtuse,  pale.      Suture  simple;    whorls  5,  a 
little  convex,  the  last   whorl  very  rotund,  somewhat  descending  in 
front,  half  the  length  of  the  shell.     Aperture  rotund-truncate,  livid 
inside;  columella  oblique,  rather  wide,  depressed-concave,  brown, 


COCHLOSTYLA-CALOCOCHLEA.  153 

the  margin  whitish.  Peristome  simple;  nearly  semicircular,  margin 
somewhat  obtuse,  chestnut,  slightly  thickened  toward  the  columella. 
(Hid.)  Alt.  42,  greater  diam.  41,  lesser  39  mill. 

Mainit,  Inland  of  Mindanao. 

C.  mainifensis  HID.,  Journ.  de  Conchyl.  1888,  p.  310;  /.  c.  18*11, 
t.  14,  f.  2. 

Under  a  considerable  magnification  are  seen  stria?  of  growth  and 
transverse  stria?,  but  very  indistinct.  (Hid.') 

C.  LIGNICOLOR  Mollendorff.      Unfigured. 

Imperforate,  subdepressed-globose  or  globose,  subtly  striated, 
chestnut-brown,  covered  with  a  streaked  and  lineated  wood-colored 
hydrophauous  cuticle,  a  rather  wide  tawny-whitish  band  at  the 
periphery,  and  sometimes  another  around  the  columella  ;  spire  glo- 
bose-conoid, apex  flat.  Whorls  5,  convex,  the  last  tumid,  above  the 
aperture  denuded  and  shining.  Aperture  lunate-rotund  ;  peristome 
straight,  acute,  subthickened,  brown  ;  columella  wide,  rose- white, 
rather  straightened,  sloping.  (Moll.) 

Alt.  32,  greater  diam.  44,  lesser  38  mill. 
Alt.  37,  greater  diam.  44,  lesser  38  mill. 
Alt.  35,  greater  diam.  39,  lesser  33  mill. 

Mts.  on  L.  Mainit,  Mindanao. 

C.  (Callicochlias)  lignicolor  v.  MOLL.  Nachr.  D.  M.  Ges.  July- 
Aug.  1888,  p.  100. 

In  color  and  markings  it  approaches  C.  depressa  Semp.  and  we- 
'lanochila  Val.  while  it  is  more  globular  than  the  last,  and  in  contour 
resembles  most  C.  sphcerion  Sow7.  The  non-expanded,  weakly 
thickened  peristome  distinguishes  it  from  all  the  foregoing,  being 
found  only  in  C.  suprabqdia  Semp.  The  white  peripheral  band  is 
lacking  on  none  of  the  ten  examples  before  me,  whilst  the  circum- 
umbilical  is  present  upon  four  only.  The  height  of  the  spire  varies, 
as  the  above  dimensions  show7,  not  inconsiderably,  but  the  higher 
forms  seem  to  predominate.  (Moll.) 

Compare  C.  mainitensis  Hidalgo. 

C.  SOLIDA  Pfeiffer.     PI.  30,  fig.  22. 

Imperforate,  conoid-semiglobose,  thick,  striate,  tawny,  covered 
with  a  thin  brown  lusterless  cuticle;  spire  convex,  obtuse,  the  apex 
reddish  ;  whorls  5,  slightly  convex,  the  last  more  convex,  obsoletely 
angulated  in  middle,  slightly  descending  in  front.  Columella 


COCHLOSTYLA-CALOCOCHLEA. 

straightened,  sloping,  wide,  whitish.  Aperture  oblique,  subtetra- 
gonal-lunar,  white  inside;  peristome  somewhat  thickened,  a  trifle 
expanded,  brown-edged.  (Pfr.) 

Alt.  27,  greater  diam.  37,  lesser  33  mill. 

Naujan,  Mindoro. 

H.  solida  PFR.,  P.  Z.  S.  1851  ;  Monogr.  iii,  p.  187.— H.  sphwrion 
PFR.,  Conchyl.  Cab.  t.  42,  f.  5,  6  (not  description).— REEVE,  Conch. 
Icon.  f.  114. 

C.  SPH.ERION  Sowerby.     PL  31,  figs.  34,  35. 

Shell  globose-orbicular,  thin,  smooth,  unicolored,  scarcely  shining  ; 
spire  short,  little  elevated ;  whorls  4,  ventricose ;  sculptured  with 
delicate  growth-lines  only;  last  whorl  large,  much  larger  than  the 
others  ;  suture  a  distinct  white  line ;  aperture  rounded,  little  modi- 
fied by  the  last  whorl,  inside  whitish ;  outer  lip  brown,  a  little  re- 
flexed  in  front;  columellar  lip  white,  rather  widened,  deeply  in- 
serted. (Soivb.)  Alt.  1-3,  diam.  1-6  inch.  (Sowb.) 

Tanauan,  Island  of  Leyte,  Philippines. 

H.  sphwrion  Sows.,  P.  Z.  S.  1841,  p.  2.— PFR.,  Conchyl.  Cab.  t. 
150,  f.  5,  6  (not  /.  c.,  t.  42,  f.  5,  6). — H.  intincta  SHUTTLW.,  Bern. 
Mittheil.,  Aug.  1852,  p.  196. —  Coclilostyla  sphcerion  SEMPER,  Reis. 
p.  184. — HIDALGO,  Journ.  de  Conchyl.  1887,  p.  127. 

I  have  not  seen  the  typical  form  of  this  species.  Sowerby's  ori- 
ginal description  is  given  above.  As  varieties  the  following  forms 
have  been  described. 

Var.  IXTINCTA  Shuttlw.     PI.  31,  figs.  31,  32,  33. 

Buff  above,  chestnut-brown  below  the  periphery;  a  dark  streak 
behind  the  peristome ;  lip  but  slightly  expanded,  brown  ;  columella 
white,  wide,  slightly  concave,  flattened  and  somewhat  excavated. 
Alt.  30,  diam.  32-1  mill. 

This  is  the  only  form  of  C.  spJucrion  I  have  seen.  The  coloring 
and  scarcely  expanded  lip,  as  well  as  the  rather  thin  texture,  re- 
mind one  of  C.  dimera.  Like  that  shell,  it  shows  no  spiral  stride 
when  examined  under  a  lens.  This  is  the  H.  spkarion  of  Philippi 
(Abl)ild.  iii,  p.  54,  Helix  t.  10,  f.  2)  and  of  Reeve  (Conch.  Icon.  f. 
Ilia  . 

Y\i:.  Form  as  in  the  last,  but  unicolored  dark  brown  all  over; 
siit ure  with  a  light  line.  Reeve's  figure  lllb  represents  this  color- 
pattern. 


COCIILOSTYLA-CALOCOCHLEA.  155 

VAR.  Three-banded  with  brown,  one  band  at  the  white  suture, 
one  at  the  periphery,  the  third  around  the  columella  (Hidalgo.} 

VAR.  Shell  buff-tawny,  the  base  and  three  narrow  zones  brown. 
Leyte.  (Hidalgo.) 

VAR.  Much  larger  and  thicker  than  specimens  from  Leyte,  the 
lip  much  more  strongly  reflexed,  and  pure  white.  Alt.  39,  diam. 
47  mill. 

Island  of  Panaon,  off  S.  Leyte. 
Doubtfully  referred  to  sphcerion.     Described  by  Semper. 

VAR.  Another  form  resembling  v.  intincta  was  collected  by 
Semper  at  Higaquit,  Tandag,  S.  Juan  de  Bislig  and  Placer,  on  the 
east  coast  of  Mindanao. 

Var.  NANA  Semper. 

A  small  form  from  Maasin  and  Malitboc,  on  the  southern  coast 
of  Leyte.  There  are  5  principal  color-patterns,  as  follows:  (1) 
Unicolored  yellow,  no  trace  of  bands ;  (2)  Clear  yellow,  with  a 
brown  line  at  suture,  and  gradually  becoming  brown  toward  the 
peristome  and  beneath  ;  (3)  brownish-yellow,  beneath  dark  brown  ; 
(4)  last  whorl  unicolored  brown-black ;  (5)  black  with  a  light  yel- 
low band  above.  The  largest  specimen  measures:  alt.  25,  diam. 
34  mill.;  the  smallest  specimen,  alt.  21,  diam.  27  mill. 

C.  XANTHOBASIS  Pilsbry.     PL  54,  figs.  38,  39,  40. 

Subglobose,  thin  but  rather  strong,  obliquely  striatulate  and  some- 
times very  superficially  spirally  malleated.  White  above,  bright 
straw-yellow  beneath.  Suture  narrowly  bordered  below  with  yellow. 

Spire  convex,  less  obtuse  than  in  C.  intincta  Shuttlw.  Whorls  4s. 
the  earlier  nearly  flat,  with  linear  non- impressed  suture ;  last  whorl 
somewhat  descending  and  then  ascending  in  front.  Peristome 
entirely  pure  white  on  both  face  and  reverse,  broadly  expanded;  col- 
umella arcuate,  thin,  its  face  excavated  ;  outer  edge  of  the  columel- 
lar  callus  brown -tinted. 

Alt.  33,  greater  diam.  35,  lesser  29  mill. 

Alt.  28,  greater  diam.  34  mill. 

A  Ibay,  Ph  Hipp  ines. 

Allied  to  certain  forms  of  C.  sphcerion  in  color-pattern,  but  differ- 
ing in  the  broadly  expanded  lip,  etc. 

The  cuticle  is  extremely  thin  above,  and  is  readily  lost,  leaving 
that  part  of  the  surface  chalk-white.  When  present,  it  is  nearly 


COCHLOSTYLA-CALOCOCHLEA. 

wliite.     The  earlier  whorls  are  white.     The  base  is  of  a  glossy  bright 
yellow. 

C.  DIMERA  Jonas.     PL  30,  figs.  29,  30. 

Imperforate,  globose-elevated,  rather  thin  ;  surface  showing  faint 
-TO  \\th-lines  but  no  spiral  striae.  Buff  above,  chocolate-brown  below 
the  periphery,  the  suture  margined  above  and  below  with  brown  ; 
•ipicnl  whorl  dark  purple,  fading  into  pink  on  the  second  and  third 
whorls.  Under  the  thin  buff  cuticle  the  upper  surface  is  white. 

Spire  elevated,  apex  obtuse.  Whorls  5,  slightly  convex,  the  last 
but  slightly  descending.  Aperture  half-round,  bicolored  within  ; 
lip  very  slightly  expanded,  dark  brown;  columella  rather  wide, 
white. 

Alt.  31,  diam.  25*  mill. 

Alt,  254,  diara.  24  mill. 

Mindoro,  Philipp  ines. 

H.  dimera  JONAS,  Moll.  Beitr.,  p.  23,  t.  11,  f.  17.  —  PFR.,  Monogr. 
i,  p.  22-")  ;  Conchyl.  Cab.  t.  50,  f.  3,  4.  —  REEVE,  Conch.  Icon.  f.  61. 
-H.  Jonasi  var.   beta   PFR.,   P.   Z.   S.  1845,  p.  126  (teste  Pfr.).- 
Cochlostyla  dimera  SEMPER,  Reis.  p.  187.  —  HIDALGO,   Jouru.    de 
Conchyl.  1887,  p.  141  (with  var.). 

Hidalgo  describes  a  variety  with  whitish  shell,  the  apex  violaceous, 
i-olumella  surrounded  writh  tawny  zone. 


C.  <  i.'n-sKi  Hidalgo.     PI.  56,  figs.  57,  58. 

Imperforate,  subglobose-conoid,  the  apex  obtuse;  rather  solid, 
opaque,  not  shining,  scarcely  striatulate;  blackish  under  an  ashen- 
brown  epidermis,  which  is  transversely  lineated  with  brown. 
Whorls  4  to  41,  convex,  the  first  nude,  blue  black,  the  last  equaling 
one-half  the  total  length,  the  base  less  convex.  Aperture  rounded- 
subquadrangular,  blue  or  blue-brown  inside;  columella  oblique, 
subdilated,  white;  peristome  black,  slightly  reflexed. 

Alt.  26,  greater  diam.  28,  lesser  25  mill.     (Hid.') 

Agcalatao,  near  Badajos,  Tablas. 

<  .  Crossei  HID.,  Journ.  de  Conchyl.  1887,  p.  141,  t.  5,  f.  4. 

This  species  presents  analogies  with  C.  dimera,  glaucopthalma, 
/ihnfd  and  carbonaria.  It  is  distinguished  from  the  first  by  its  colora- 
tion,  its  cuticle,  and  its  more  ventricose  form  ;  from  the  second  by 
it<  ircin'ra!  form  and  cj)idermis  ;  from  the  third  by  its  coloration  and 


COCHLOSTYLA-CALOCOCHLEA.  157 

peristome  ;  finally  from  the  last  named  species  by  its  coloration,  col- 
umella, and  the  non-angular  periphery.     (Hid.} 

This  species,  with  the  two  following,  forms  a  passage  to  the 
globose  forms  of  the  section  Axina. 

C.  RETUSA  Pfeiffer.     PI.  56,  figs.  59,  60. 

Imperforate,  subglobose,  the  apex  retuse;  thin,  smooth,  the  base 
concentrically  striolate ;  chestnut-tawny,  broadly  flamed  and 
streaked  with  white ;  partly  covered  with  a  hydrophanous  white, 
deciduous  cuticle,  pallid  above,  chestnut-banded.  Spire  short. 
Whorls  4,  a  little  convex,  the  last  subangular,  scarcely  descending ; 
columella  entering,  white,  dentate-dilated  above.  Aperture  rounded- 
lunar,  pearly  within  ;  peristome  broadly  expanded,  a  little  reflexecl. 

Alt.  20,  greater  diam.  28,  lesser  24  mill.     (Pfr.) 

Samar,  Philippines. 

H.  retusa  PFR.,  P.  Z.  S.  1845,  p.  132;  Monogr.  i,  p.  255;  Con- 
chyl. Cab.,  t.  47,  f.  9,  10.— PHILIPPI,  Abbild.  ii,  p.  84,  t.  7,  f.  2.— 
REEVE,  f.  82. 

C.  LIBATA  Reeve.     PI.  56,  figs.  61,  62,  63. 

Imperforate,  subglobose,  solid,  nearly  smooth  ;  epidermis  brown, 
covered  with  darker  spiral  lines.  Spire  serniglobose,  apex  obtuse, 
nude,  brown.  AVhorls  4*,  a  little  convex,  the  last  rounded,  not 
much  wider  than  the  preceding  volution  ;  base  denuded,  buff;  col- 
umellar area  chestnut ;  columella  sub  vertical,  flat,  white.  Aperture 
oblique,  subtetragonal-lunar,  white  within  ;  peristome  brownish,  nar- 
rowly expanded,  right  margin  bowed. 

Alt.  24,  greater  diam.  29,  lesser  25  mill.     (Pfr.) 

Benguet  and  Mancayan,  N.-  W.  Luzon. 

H.  libata  RVE.,  Conch.  Icon.,  f.  35. — PFR.,  Monogr.  iii,  p.  189; 
Conchyl.  Cab.,  p.  438,  t.  152,  f.  7,  8. —  Cochlostyla  libata  SEMPER, 
Reis.,  p.  186. — HIDALGO,  Journ.  de  Conchyl.  1887,  p.  147. 

Hidalgo  notices  these  color-varieties : 

1.  Like  the  type,  but  lip  whitish,  and  the  blackish  columellar 
band  absent. 

2.  Buff,  ornamented  with  two  very  broad  chestnut  bands;  suture 
and  columellar  area  black,  apex  violaceous. 

3.  Tawny,  last  whorl  unifasciate  in  the  middle ;  suture  and  col- 

* ' 

lellararea  black,  apex  violaceous,  (fig.  61.) 


umellar 


l.-,X  COCHLOSTYLA-AXINA. 

Subsection  2.     AXINA  Albers,  1850. 

Axina  ALB.,  Die  Hel.,  p.  113.— v.  MART.,  Die  Hel.  1861,  p.  71 
type  H.  zebnensis). — PFEIFFER,  Nomencl.  Hel.  Viv.  p.  199,  1878. 
-SKMTER,  Land  moll.  Phil.  190.— v.  MOLL.,  Landschn.  Cebu.  p. 
'2-\4. —  Carocolla  of  some  early  authors. 

Im perforate  shells,  varying  from  globose  turbinate  to  flattened 
carinated  and  lens-shaped  ;  solid,  dark  colored]  more  or  leas  covered 
with  a  hydrophanous  cuticle ;  the  surface  smooth  or  at  least  totally 
licking  spim/  sculpture;  whorls  42-6  ;  lip  either  narrow  or  broadly 

reflexed. 

Anatomy  as  in  Cochlostyla  generally.  The  group  seems  to  replace 
Calocochlea,  on  the  islands  Siquijor  and  Zebu. 

Key  to  species  of  Axina. 

A.  Peripheral  keel  very  acute 

b.  Lip  broad,  flat,  angled  and  somewhat  guttered  at  the  keel, 
c.  Whorl  deflexed  at  aperture  ;  diam.  about  70  mill., 

garibaldiana. 
cc.  Whorl  not  deflexed  at  aperture;    diam.  about  40  mill., 

siquijorensis. 
bb.  Lip  narrow,  its  face  not  flattened, 

c.  Shell  about  equally  convex  above  and  below  the  keel, 

d.  Surface  smooth,  obliquely  streaked  above,       zebuensis. 
dd.  Surface  densely,  finely  lamellose,  not  variegated  above, 

striatissima. 

cc.  Shell  flattened  below,  elevated  above  the  keel      carbonaria. 
AA.  Peripheral  keel  rather  obtuse,  or  periphery  rounded 

b.  Lip  broadly  reflexed  ;  columella  distinctly  toothed  at  base, 
c.  Periphery  obtusely  keeled  ;  whorl  deflexed  at  aperture 

pfel/eri. 
bb.  Lip  rather  narrowly  reflexed 

c.  Alt.  exceeding  the  lesser  diameter  of  the  shell;  columella 
broad,  milk-white  ;  periphery  rounded  or  very  indistinctly 
angled. 

d.  Cuticle  obliquely  streaked  ;  alt.  24,  diam.  25  mill., 

bruguieriana. 

</'/.  Cuticle  scarcely  streaked  ;  alt.  27-29,  diam.  28-29  mill. 

montfortiana. 

cc.  Alt.  less  than  the  lesser  diam.  of  shell;    columella  pink  or 
brown  tinted. 


COCHLOSTYLA-AXINA.  159 

d.  Depressed  ;    alt.  about  one-half  the  diam. ;    periphery 
rounded,  phloiodes. 

dd.  Rather  elevated  ;   decidedly  angled  at  periphery,  flat- 
tened below,  carbonaria. 
ddd.  Not  very  obviously  angled  at  periphery, 

e.  Whorls  narrowly  revolving,  aperture  narrow  ;  base 

very  convex  magistra. 

ee.  Whorls  more  rapidly    increasing ;    aperture   wide, 

base  rather  flattened  moreleli. 


<C.  GARFBALDIANA  Dohrii  &  Semper.     PL  20,  figs.  1,  2. 

Lenticular,  solid,  sculptured  with  growth-stride;  chestnut,  spotted 
and  figured  all  over  with  straw-color,  marked  with  interrupted 
darker  concentric  lines ;  acutely  and  compressly  carinated,  carina 
subexserted  on  the  whorls  above ;  spire  moderately  elevated,  apex 
depressed ;  whorls  5,  scarcely  convex,  depressed  at  the  suture, 
the  last  whorl  descending  in  front,  more  convex  beneath  ;  aperture 
ax-shaped,  bluish  inside ;  peristome  thickened,  expanded,  reflexed, 
fleshy-violet,  the  right  margin  angular,  somewhat  channelled  at  the 
angle;  columellar  margin  dilated,  planate,  very  oblique. 

Alt.  27,  greater  diam.  72,  lesser  64  mill. 

Northern  Luzon. 

Cochlostyla  (Axinci)  garibaldiana  D.  &  S.,  Mai.  Bl.  ix,  1862,  p. 
206. — Helix  garibaldiana  PFR.,  Novit.  Conch,  p.  219,  t.  58,  f.  1,  2  ; 
Monogr.  v.  p.  806. 

G.  SIQUIJORENSIS  Broderip.     PL  20,  figs.  8,  9,  10. 

Imperforate,  solid,  depressed  and  very  acutely  keeled ;  convex 
above  and  below ;  reddish-brown,  the  earlier  whorls  purple,  dark 
brown  or  whitish  ;  body-whorl  not  descending  in  front ;  lip  broad, 
reflexed. 

The  surface  is  smooth,  often  slightly  malleated  above  the  acute, 
knife-like  keel,  and  usually  obliquely  malleated  below  it ;  growth- 
striseilight.  The  hydrophanous  epidermis  is  generally  entirely 
rubbed  oft',  but  often  persists  near  the  keel  and  behind  the  lip. 
AVhorls  5,  flattened,  the  last  not  descending  in  front.  Aperture 
oblique,  fleshy-white  inside ;  lip  broadly  reflexed  on  its  outer  and 


160  COCHLOSTYLA-AXINA. 

basal  mam-ins,  angled  at  the  carina,  brown  in  color.     Columella  not 
Toothed. 

Air.  Ill,  greater  diam.  39,  lesser  33  mill. 

Alt.  22,  greater  diam.  43,  lesser  37  mill. 

Sin  Juan,  etc.,  Island  oj  Siquijor,  Philippines. 

Carocolln  .<!</,! i/orensis  BROD.,  P.  Z.  8.  1841,  p.  38.— Helix  siqui- 
joren*i*  PFK,.  Monogr.  i,  p.  295 ;  Conch.  Cab.  p.  29,  t.  69,  f.  1,  2,  3, 
4._I>VE.,  Conch.  Icon.  f.  119. — HIDALGO,  Obras,  i,  p.  143,  t.  12,  f. 


2.  3. 


Keel  more  acute  and  lip  broader  than  in  the  allied  C.  zebuensis, 
which  further  differs  in  having  the  whorl  deflexed  in  front.  There 
is  some  variation  in  height  of  spire  and  in  color. 

Var.  pallens  v.  Moll.  Shell  smaller,  often  higher,  buff.  (v.  Moll., 
.lahr.  I).  M.  Ges.  1891,  p.  46.  Vide  Raster,  pi.  69,  f.  3,  4.) 

(  '.  s<  IIADEXBERGI  Molleudorff.      Unfigured. 

Im  perforate,  angulate-globose,  carinated,  solid,  bright  chestnut, 
the  hydrophanous  cuticle  yellowish-white,  arranged  in  a  series  of 
bands  of  spots  ;  spire  convex-conoid,  apex  obtuse,  paler.  Whorls 
5-j,  nearly  flat,  slowly  increasing,  separated  by  rather  im- 
pressed sutures,  transversely  striated,  microscopically  granulose, 
the  last  impressed  under  the  rather  thick  exserted  carina,  then  a 
little  convex,  in  front  suddenly  and  deeply  deflexed,  in  adults 
malleated.  Aperture  very  oblique,  ax-shaped;  peristome  rather 
broadly  expanded,  reflexed,  thickened,  white  lipped,  outwardly 
brown,  upper  margin  sinuous,  outer  margin  subcanaliculate  in  the 
middle;  columella  wide,  somewhat  straightened,  obliquely  descend- 
ing, forming  an  obtuse  angle  with  the  basal  margin,  surrounded  by 
a  wide  callus.  (Moll.} 

Alt.  39,  diam.  59  mill.;  aperture  width  with  callus,  39,  alt.  32 
mill. 

District  of  Quiangan,  province  Nueia  Vizcaya,  central  Luzon. 

C.  f  .l./-//m)  schadenberyi  v.  MOLL.,  Xachr.  D.  M.  Ges.  1890,  p. 
205. 


I  Mollendorff.      Unfigured. 

Imperforate,  depressed-globose,  angulate,  solid,  somewhat  shining, 
olive-buff,  brown  at  the  suture,  ornamented  with  bands  of  whitish 
hydrophanous  cuticle,  regularly  interrupted  at  the  suture.  Spire 
convex,  apex  obtuse,  nearly  planate.  Whorls  5,  slowly 


COCHLOSTYLA-AXINA.  161 

increasing,  a  little  convex,  separated  by  little  impressed  suture, 
lightly  striated,  decussated  by  almost  microscopic  spiral  lines, 
sculptured  with  obliquely  forward  descending  wrinkles ;  last  whorl 
nearly  flat  on  the  base,  malleate-rugose,  in  front  shortly  but  dis- 
tinctly deflexed.  Aperture  oblique,  rotund  pentagonal ;  peristbme 
broadly  expanded,  a  little  reflexed,  callous  labiate,  reddish-brown, 
the  upper  margin  sigmoidly  sinuate,  the  outer  chanelled  in  the 
middle ;  columella  short,  obliquely  descending,  outwardly  plicate, 
base  nodiformly  thickened.  (Moll.) 

Alt.  32,  diam.  48  mill.;  a  pert.,  width  with  callus  32,  alt.  25  mill. 

ML  Limutan,  (list.  Morong,  Luzon. 

C.  (Axina)  Kobelti  v.  MOLL.,  Nachr.  D.  M.  Ges.  1890,  p.  205. 

C.  ZEBUENSIS  Broderip.     PI.  20,  figs.  3,  4,  5,  6. 

Imperforate,  depressed,  solid,  acutely  carinated,  about  equally 
convex  above  and  below  the  carina.  Covered  with  a  buff  cuticle, 
becoming  reddish  on  the  body-whorl,  and  either  continuous  above, 
with  dark  spots  at  the  carina  (fig.  3),  or  conspicuously  streaked 
(figs.  5,  6)  ;  the  base  more  or  less  cut  by  spiral  dark  and  light  lines. 
Apical  whorls  purple  or  livid  reddish-brown.  Surface  smooth. 

The  surface  is  quite  smooth,  even  in  perfectly  fresh  examples. 
Whorls  4J,  nearly  flat,  the  last  descending  more  or  less  below  the 
carina  in  front.  Aperture  oblique,  bluish-white  inside;  peristome 
dark  brown,  obtuse,  narrowly  expanded  above,  angled  at  the  keel, 
narrowly  reflexed  beneath.  Columella  obliquely  sloping,  nearly 
straight,  more  or  less  obviously  toothed  below. 

Alt.  19,  greater  diam.  36,  lesser  30?  mill. 

Alt.  16,  greater  diam.  29,  lesser  25*  mill. 

Western  Zebu. 

Carocolla  zebuensis  BROD.  (in  part),  P.  Z.  S.  1841,  p.  46. — RVE., 
Conch.  Syst.  ii,  t.  168,  f.  14,  16. —  H.  zebuensis  PFR.,  Monogr.  i,  p. 
296 ;  Conchyl.  Cab.  t.  68,  f.  12,  13,  14,  15.— HIDALGO,  Obras,  i,  p. 
143,  t.  11,  f.  4,  5. —  Cochlostyla  (Axina)  zebuemis  SEMP.,  Landschn. 
Phil.  p.  191. — MLLDFF.,  Landschn.  Cebu,  p.  234. 

Broderip  and  authors  having  included  two  species  under  the  name 
zebuensis,  it  becomes  necessary  to  select  one  for  the  type.  The  pub- 
lished figures  all  represent  the  variegated  form  with  smooth  epider- 
mis, and  that  may,  therefore,  retain  the  name  zebuensis. 

The  range  of  variation  is  considerable  in  size  as  well  as  in  color- 
ing. Figs.  5,  6  of  plate  20  represents  the  largest  specimen  I  have 

seen. 

11 


162  COCHLOSTYLA-AXINA. 

C.  BTRIATISSIMA  PiUbry.     PL  57,  figs.  73,  74,  75  ;  pi.  20,  fig.  7. 

I  in  perforate,  depressed,  solid,  acutely  carinated,  about  equally 
convex  above  and  below  the  keel.  Covered  with  a  brown  cuticle 
which  is  finely  and  regularly  raised  into  narrow  lamellce  in  the 
direction  of  growth-lines,  the  intervals  between  the  lamellse  being 
microscopically  obliquely  wrinkled.  Unicolored  above,  except  that 
the  earlier  whorls  are  denuded  of  cuticle  and  dark  purplish  ;  the 
base  having  usually  numerous  darker  concentric  lines. 

The  surface  is  lusterless,  densely  lamellose,  the  cuticular  folds 
beinfc  close  and  acute.  The  upper  surface  has  neither  oblique 
streaks  nor  spiral  lines.  Whorls  4f,  nearly  flat,  the  last  slightly  or 
considerably  descending  in  front.  Aperture  oblique,  livid-whitish 
inside  ;  peristome  blackish-brown,  narrowly  expanded  above,  angled 
at  the  periphery,  narrowly  reflexed  below  ;  columella  sloping,  not 
toothed. 

Alt.  18,  greater  diam.  33,  lesser  30  mill. 

Zebu,  Philippines. 

H.  (Carocolla)  zebuensis,  var.  e.  tota  fusca,  BROD.,  P.  Z.  S.  1841, 
p.  46. 

This  form  differs  from  C.  zebuensis  in  being  unicolored  above,  and 
having  densely  lamellose  cuticle. 

C.  PFEIFFEKI  Semper.     PI.  44,  figs.  34,  35,  36,  37. 

Ira  perforate,  depressed  or  depressed-turbinate,  bluntly  angled  at 
the  periphery,  more  convex  above  than  below  the  angle ;  dark 
brown,  having  a  whitish-buff  hydrophanous  epidermis,  radiating 
from  the  suture  above,  and  cut  by  spiral  dark  lines,  forming  a  wide 
band  just  below  the  peripheral  angle,  and  cut  into  interrupted  nar- 
row concentric  bands  on  the  base. 

Surface  smooth,  lightly  striatulate.  Whorls  4f ,  somewhat  con- 
vex, the  last  descending  in  front.  Aperture  oblique,  livid-whitish 
within;  peristome  brown,  rather  broadly  expanded  and  reflexed  ; 
columella  sloping,  abruptly  truncated  or  toothed  below. 

Alt.  22,  greater  diam.  33,  lesser  28  mill. 

Alt.  26,  greater  diam.  31,  lesser  28  mill. 

Dalaguete,  western  Zebu,  Philippines. 

H.cuming  I'KIJ.,  P.  Z.  S.  1842,  p.  88  ;  Monogr.  i,  p.  264;  Conchyl. 
Cab.  t.  70,  f.  3,  4.— PHIL.,  Abbild.  i,  p.  69,  t.  3,  f.  4.— REEVE, 
Conch,  Iron.  f.  .11.-  -I!ij>Au;o,  Obras,  i,  pp.  144,  t.  11,  f.  6,  7.— (Not 
cumingi  Pfr.,  also  a  Cochlostyla^—Cochlostyla  (Axina) 


COCHLOSTYLA-AXINA.  163 

Pfeifferi  SEMPER,  Landmoll.  Phil.  p.  191. — v.  MOLL.,  Landschn. 
Cebu,  p.  236. 

Closely  allied  to  C.  magistra  Pfr.,  C.  gloynei  Sowb.,  but  less  glo- 
bose, with  more  oblique  aperture  and  wider  lip.  The  base  is  not 
excavated  arouni  the  axis  as  it  is  in  magistra,  and  the  whorls  of 
the  spire  do  not  revolve  nearly  so  closely. 

C.  PHLOIODES  Pfeiffer.     PI.  40,  figs.  8,  9. 

Imperforate,  depressed,  solid  ;  shining  blackish-red  under  the 
papery  epidermis  which  is  varied  with  darker  streaks  ;  spire  sub- 
planulate;  suture  profound ;  whorls  4£,  convex;  columella  nearly 
straight,  very  oblique,  subexcavated.  Aperture  wide-lunar,  livid 
inside,  shining ;  peristome  thickened,  reflexed,  brown.  (Pfr.) 

Alt.  17i,  greater  diam.  29,  lesser  25  mill. 

Zebu,  Philippines. 

H.  phloiodes  PFR.,  P.  Z.  S.  1842,  p.  151  ;  Conchyl.  Cab.  p.  427,  t, 
150,  f.  3,  4;  Monogr.  i,  p.  265.— RVE.,  Conch.  Icon.  f.  55.— 
HIDALGO,  Obras,  i,  p.  146. —  Cochlostyla  (Axina)  phloiodes  MOLL., 
Landschn.  Cebu,  p.  236. 

C.  CARBONARTA  Sowerby.     PI.  43,  figs.  30,  31,  32 ;  pi.  44,  fig.  33. 

Imperforate,  solid,  low-trochiform,  varying  from  subangular  to 
carinated  at  the  periphery,  decidedly  flattened  below  the  angle,  very 
convex  above  it.  Very  dark  brown,  covered  with  a  light  brown 

«/  o 

hydrophanous  cuticle,  usually  obliquely  streaked  dark  and  light 
above,  the  base  streaked  radially  with  blackish  and  light,  and  usu- 
ally spirally  lineolate. 

The  surface  is  lightly  wrinkled  by  growth  lines;  the  hydroph- 
anous streaks  are  broAvnish,  less  conspicuously  contrasting  with  the 
dark  streaks  than  in  most  species.  The  apical  whorls  are  dark 
purple  or  rosy.  The  lateral  outlines  of  the  spire  are  quite  convex. 
Whorls  5J,  slightly  convex,  the  last  not  descending  in  front. 
Aperture  oblique,  pinkish-plumbeous  within;  peristome  very  nar- 
rowly expanded,  deep  brown  ;  the  columella  nearly  vertical,  dilated, 
not  toothed,  pinkish-brown  or  whitish. 

Alt  26,  greater  diam.  29,  lesser  27  mill. 

Alt.  22,  greater  diam.  27-.  lesser  25 £  mill. 

Negros ;  Boljoon  and  Sambuan,  Zebu,  Philippines. 

H.  carbonaria  (SoAVB.,  in  sched.)  PFR.,  P.  Z.  S.  1842,  p.  86 ; 
Conchyl.  Cab.  p.  285,  t.  47,  f.  6-8 ;  Monogr.  i,  p.  296.— REEVE, 


COCHLOSTYLA-AXINA. 

Conch.  Icon.  f.  63— HIDALGO,  Obras,  i,  p.   145,  t.   12,  f.   6,   7.- 
Cochlostyla  (Axina)  carbonaria  SEMPER,  Reis.,  p.  191. — v.  MOLL., 
Landschn.  Cebu,  p.  237. 

Among  typical  examples,  at  Alegria  and  Malabuyoc,  are  found 
some  of  a  handsome  red  color,  with  fiery-red  peristome,  which  v. 
Mollendorff  proposes  to  call  deviation  rubens. 

('.  MAGISTRA  Pfeiffer.     PL  41,  figs.  17,  18,  19;   pi.  57,  fig.  76;   pi. 

21,  figs.  37,  38. 

Imperforate,  solid,  subglobose-depressed,  the  periphery  very  ob- 
tusely angled.  Blackish-brown  obliquely  streaked  and  spirally 
lineolate  with  buff-white  hydrophanous  cuticle  above,  having  a 
broad  white  band  just  below  the  periphery,  the  base  having  many 
concentric  whitish  lines  and  some  radiating  streaks. 

The  surface  is  smooth,  lightly  striatulate.  There  is  a  row  of  dark 
and  light  blotches  along  the  suture,  as  in  C.  pfeifferi.  Whorls  5s, 
convex,  closely  coiled,  the  last  a  very  little  descending  in  front. 
Aperture  whitish-leaden  within ;  lip  reflexed,  blackish-brown  ;  col- 
umella  white,  deeply  inserted,  its  edge  slightly  convex,  not  toothed 
at  base.  Alt.  22  J,  greater  diam.  29s,  lesser  26  mill. 

Medellin,  northern  Zebu;  Islands  of  Magtan  and  Timobo  near 
Leyte. 

ff.  magistra  PFR.,  Zeitschr.  f.  Mai.  1852,  p.  91  ;  Conchy  1.  Cab.  p. 
48f>,  t.  159,  f.  38-40;  Monogr.  iii,  p.  178. — HIDALGO,  Obras,  i,  p. 
145,  t.  xi,  f.  3. — H.  magister  REEVE,  Conch.  Icon.  f.  1018. —  Cocltlo- 
xtijld  (Axina)  magistra  v.  MOLL.,  Landschu.  Cebu,  p.  239  (with  var. 
ecarinata). —  Cochlontyla  (Axina)  gloynei  SOWB.,  Journ.  Linn.  Soc. 
Loud,  xx,  p.  398,  t.  25,  f.  16,  17  (1889).— H.  (Axina)  gloynei 
HIDALGO,  Obras,  i,  p.  144,  t.  11,  f.  1,  2. 

This  species  was  originally  described  from  specimens  denuded  of 
their  hydrophanous  cuticle.  The  original  figures,  copied  on  my  pi. 
41,  figs.  17-19,  represent  them  in  this  condition.  A  more  satisfac- 
tory illustration  is  pi.  57,  fig.  76. 

From  C.  pfeifferi  it  is  constantly  and  readily  distinguished  by  the 
more  narrowly  revolving  whorls  seen  from  above,  and  the  more 
Mvollcii  base,  which  is  far  more  excavated  at  the  axis  than  in  C. 
pfeifferi. 

The  following  varietal  forms  have  received  names : 


COCHLOSTYLA-AXINA.  165 

Var.  gloynei  Sowb.  (1889).     PL  21,  figs.  37,  38. 

More  globose,  elegantly  fenestrated  with  cream- white;  periphery 
very  obtusely  carinated.  Apical  whorls  rose. 

Var.  ecarinata  v.  Moll.  (1890). 

Nearly  6  little-convex  whorls,  the  last  not  angulated ;  cuticle  lat- 
ticed with  white,  apex  rose,  right  margin  of  the  aperture  not  in  the 
least  angulated,  peristome  less  expanded  and  reflexed. 

Alt.  22J,  diam.  32*  mill. 

Magtan ;    Tim  obo. 

C.  MORELETI  Pfeiffer.     PL  45,  figs.  45,  46. 

Imperforate,  conoid-globose,  rather  solid ;  brown-black,  cov- 
ered with  a  shining  buff-chestnut  cuticle,  variously  striped  and  zig- 
zagly  streaked:  Spire  sub-semi-globose,  apex  roseate,  rather  acute. 
Whorls  4*,  little  convex,  separated  by  somewhat  impressed  sutures, 
the  last  whorl  large,  peculiarly  flattened  at  the  periphery,  toward 
the  base  angulated,  beneath  somewhat  flattened.  Aperture  very 
oblique,  roundly-quadrangular,  milk-white  inside  ;  peristome  black, 
narrowly  reflexed  ;  columella  straighened,  suboblique,  flesh-colored, 
whitish  above,  (v.  Moll.} 

Alt.  28,  greater  diam.  37  mill.;  apert.  width  25*,  alt.  21  i  mill. 

Boljoon,  Zebu. 

A.  moreleti  PFR.,  Zeitschr.  f.  Mai.  1848,  p.  118  ;  Conchy  1.  Cab. 
p.  330,  t.  130,  f.  9,  10;  Monogr.  iii,  p.  177.—  Cocfilostyla  (Axina) 
moreleti  v.  MOLL.,  Landschn.  Cebu,  p.  238. 

The  only  published  figures  represent  a  shell  from  which  the  epi- 
dermis had  been  rubbed.  Mollendorff  has  recently  described  fresh 
examples  and  discovered  the  locality,  heretofore  unknown. 

O.  MONTFORTIANA  Pfeiffer.     PL  41,  figs.  14,  15,  16. 

Imperforate,  turbinate,  thick,  ponderous,  blackish-rufous,  covered 
with  a  hydrophanous  ashen-brown  cuticle ;  spire  conoid  ;  apex  violet- 
purple,  denuded,  shining,  obtuse ;  whorls  4*,  slightly  convex,  the 
last  angulated,  angle  evanescent  on  the  latter  part  of  the  whorl ; 
columella  sloping,  callous,  white,  subdentate  at  base;  aperture  sub- 
tetragonal,  shining  white  within ;  peristome  narrowly  reflexed,  bor- 
dered with  blackish-brown.  (Pfr.) 

Alt.  22,  greater  diam.  31,  lesser  27  mill. 

Philippine  Is. 


166  COCHLOSTYLA-TRACHYSTYLA. 

H.  montfortiana  PFR.,  P.  Z.  S.  1846,  p.  38 ;  MoDogr.  i,  p.  224.— 
REEVE,  Conch.  Icon.  f.  76. 

The  specimens  before  me,  one  of  which  is  figured  on  pi.  41,  are 
not  "  ponderosa,"  nor  is  the  aperture  "  intm  alba"  Otherwise  they 
correspond  with  Pfeiffer's  diagnosis,  translated  above.  They  differ 
from  C.  bruguieriana  in  being  larger,  somewhat  more  turbiuate  in 
contour,  the  hydrophanous  cuticle  not  streaked,  the  columella  sub- 
dentate  below,  and  the  apex  purple.  These  distinctions  may  prove 
to  be  variable  when  more  shells  are  examined. 

( '.  BRUGUIERIANA  Pfeiffer.     PI.  40,  figs.  6,  7  ;  pi.  57,  figs.  77,  78. 

Imperforate,  globose-turbinate,  rather  thin  but  solid,  rounded  at 
the  periphery.  Covered  with  a  hydrophanous  cuticle  light  brown 
in  color,  obliquely  streaked  with  cream-white;  the  apical  whorls 
pink,  with  a  darker  band  along  the  suture. 

Surface  smooth ;  cuticle  removed  around  the  columella  and  in 
front  of  the  mouth,  exposing  the  dark  brown  ground-color.  Whorls 
5,  somewhat  convex,  the  last  not  descending  in  front,  or  but  slightly, 
rounded  at  the  periphery,  or  more  rarely  very  obtusely  subangular,. 
convex  beneath.  Aperture  wide  semi-lunar,  purplish-pink  inside; 
lip  very  narrowly  expanded,  blackish-brown;  columella  dilated, 
deeply  inserted,  snow-white,  brown  at  the  base. 

Alt.  24,  greater  diam.  25,  lesser  22  mill. 

Philippines. 

H.  bruguieriana  PFR.,  P.  Z.  S.  1845,  p.  44;  Conch}7!.  Cab.  p. 
L".)9,  t.  44,  f.  3,  4 ;  Monogr.  i,  p.  224.— REEVE,  Conch.  Icon.  f.  38. 

More  elevated  than  the  other  forms  of  Axina,  and  usually  quite 
rounded  ac  the  periphery.  It  is  very  difficult  to  draw  the  line  be- 
tween Axina  bruguieriana  and  Calocochlea  roissyana,  etc.,  but  still, 
what  little  difference  there  is  seems  to  be  in  favor  of  placing  brugui- 
eriana in  the  present  group. 

The  preceding  species  is  doubtfully  distinct,  but  differs  in  being 
larger,  with  less  streaked  cuticle  and  purple  apex. 

Subsection  3.     TRACIIYSTYLA  Pilsbry. 

are  the  Cocklostyhe  cinerece  of  Semper.      The  shell  is  solid^ 
colored,  with  a  dull  ashen  hydrophanous  cuticle.      The  spire  is 
short,  whorls  few  and  rapidly  ividening,  columella  oblique,  straight ; 
dull  and  roughened. 


COCHLOSTYLA-TRACHYSTYLA.  167 

The  species  live  on  the  ground  under  leaves,  etc.,  contrary  to  the 
usual  arboreal  station  of  Cochlostyla. 

C.  CRYPTICA  Broderip.     PI.  50,  fig.  1  ;  pi.  51,  fig.  9. 

Imperforate,  capacious,  solid,  depressed-globose.  Surface  sculpt- 
ured with  low  spiral  cords,  and  close,  obliquely  forward-descending 
wrinkles.  Dark  brown,  covered  with  a  thin  brownish  hydropha- 
nous  cuticle,  which  is  worn  from  the  prominences  of  the  surface. 

Spire  short,  the  first  1?  whorls  planorboid  ;  suture  impressed. 
Whorls  4A,  convex,  rapidly  widening,  the  last  somewhat  descend- 
ing in  front.  Aperture  large,  oblique,  livid  inside;  peristome flesh- 
colored,  reflexed.  Columella  wide,  very  oblique,  curving  inward  in 
the  middle.  Alt.  46,  diam.  56  mill. 

Northern  Samar ;  Eastern  Mindanao,  Philippines. 

H.  cryptica  BROD.,  P.  Z.  S.  1841,  p.  22. — PFR.,  Monogr.  i,  p. 
258;  Conchyl.  Cab.  p.  272,  t.  43,  f.  1,  2.— REEVE,  Conch.  Icon.  f. 
7. —  Cochlostyla  cryptica  SEMPER,  1.  c.  p.  189. 

I  am  unable  to  separate  as  species  the  cryptica,  latitans  and  cretata 
of  Broderip,  and  have  therefore  considered  the  two  latter  geographic 
races. 

Yar.  LATTTANS  Sowerby.     PI.  51,  figs.  10,  11. 

Imperforate,  capacious,  solid,  depressed-globose  ;  surface  rudely 
roughened  by  low  spiral  cords,  and  subobsoletely  obliquely  malleated. 
Dark  brown,  more  or  less  covered  with  a  creamy-ashen  hydropha- 
nous  cuticle,  which  is  absent  for  a  distance  above  the  periphery,  and 
in  front  of  the  mouth. 

Spire  low,  very  obtuse;  whorls  4,  the  last  but  slightly  descending 
in  front.  Aperture  very  large,  oblique,  livid-pink  inside ;  lip  re- 
flexed,  flesh-colored  ;  columella  broad,  very  oblique,  nearly  straight, 
whitish.  Alt.  50,  diam.  60  mill. 

Forests  of  Sierra  Bullones,  on  the  north  coast  of  Bohol ;  Dapa, 
Siargao,  Pta  Malimono,  near  Surigao,  Mindanao,  Philippines. 

H.  latitans  BROD.,  P.  Z.  S.  1841,  p.  23. — REEVE,  Conch.  Syst.  ii, 
t.  165,  f.  18;  Conch.  Icon.,  f.  10.— PER.,  Monogr.  i,  p.  257  ;  Con- 
chyl. Cab.,  p.  273,  t.  43,  f.  3,  4.— DELESS.,  Rec.,  t.  38,  f.  3.— 
DH.  in  Fer.,  Hist.,  t.  108 A,  f.  1,  2. —  Cochlostyla  latitans  SEMPER, 
Reis.,  p.  188,  t.  18,  f.  15  (anatomy). — ?  H.  dionacea  DH.,  in  Fer., 
Hist.,  t.  28 A,  f.  11,  12,  texte,  p.  276.— PER.,  Monogr.  iii,  p.  191. 


168  COCHLOSTYLA-TRACHYSTYLA. 

Differs  from  C.  cryptica  in  the  absence  or  obsolescence  of  oblique 
wrinkles,  and  in  the  broad  band  just  above  the  periphery,  denuded 
of  hydrophanous  cuticle. 

Var.  CRETATA  Broderip.     PI.  51,  fig.  12. 

Shell  subglobose,  swollen  ;  whorls  3,  the  last  much  the  largest,  and 
begirt  with  blackish  bands  and  lines;  closely  striated  with  oblique 
growth-lines,  and  encircled  with  elevated  spiral  lines;  brownish- 
black,  with  a  chalky  cuticle  ;  aperture  silvery-bluish,  the  lip  having 
an  obscure  reddish  border.  (Brod.') 

Tananuan,  Is.  of  Leyte,  Philippines. 

H.  cretata  BROD.,  P.  Z.  S.  1841,  p.  23. — PFR.,  Monogr.  i,  p.  258. 
-REEVE,  Conch.  Icon.,  f.  13. 

Seems  to  differ  mainly  in  the  more  conspicuous  development  of 
the  hydrophanous  cuticle. 

C.  PANAENSIS  Semper.      Unfigured. 

Imperforate,  turbinate-globose,  solid,  purple-brown,  margined 
with  tawny  at  the  suture  ;  spire  elevated,  globose ;  whorls  4,  a  little 
convex,  encircled  by  evanescent  impressed  lines ;  the  last  whorl  not 
descending ;  suture  generally  impressed ;  columella  very  much  dilated, 
excavated,  rather  sloping,  whitish-rosy  ;  aperture  ample,  subtetrago- 
nal,  livid  ;  peristome  broadly  expanded  and  reflexed,  horny-livid, 
upper  margin  strongly  arched,  basal  margin  forming  an  angle  with 
the  columella.  ($.)  Alt.  50,  greater  diam.  65,  lesser  46  mill. 

Panaon,  near  Surigao. 

C.  panaensis  SEMP.,  Reis.,  p.  189. 

The  very  obliquely  placed  aperture,  with  the  upper  lip  strongly 
arched  forward,  the  high,  globular  form  of  the  shell,  the  quite  im- 
pressed, yellow-edged  suture,  and  the  comparative  smoothness  of 
the  shell,  with  very  weakly  inclined  columella,  cause  me  to  regard 
this  as  a  species.  (S.) 

C.  CINERACEA  Semper.     PI.  59,  fig.  3. 

I  )c|)t-cssed,  solid,  rough,  and  grooved  by  irregular  impressed  lines  ; 
ashen  ;  obtusely  carinated  at  the  periphery ;  spire  shortly  conoid ; 
whorls  41,  n  little  convex,  the  last  one  nearly  flat  above,  very  con- 
vex below,  not  descending  in  front.  Columella  white,  nearly 
straight.  Aperture  oblique,  subtetragonal,  livid  within  ;  peristome 


COCH  LOSTYLA-CHROMATOSPH^ER  A.  1 69 

narrowly  expanded,  fleshy-livid  ;  columellar  margin  forming  a  right 
angle  with  the  basal.     (&) 

Alt.  35-42,  greater  diam.  70,  lesser  55  mill. 

Lianga,  Mindanao. 

C.  cineracea  C.  SEMPER,  Reis.,  p.  190,  t.  9,  f.  1. 

Stands  between  C.  cryptica  and  C.  garibaldiana. 

C1.  DATAENSIS  Semper.     PL  51,  figs.  13,  14,  15. 

Imperforate,  solid,  depressed-globose  ;  surface  roughened  by  spiral 
sulci  below,  and  obliquely  descending  wrinkles  above.  Chestnut-brown 
unicolored,  or  having  a  yellow  band  at  the  periphery,  the  lower 
edge  of  which  is  not  sharply  defined. 

Spire  low-conoid,  apex  obtuse.  Whorls  4s,  the  last  slightly  de- 
scending in  front.  Aperture  wide,  fleshy-white  within  ;  lip  well  ex- 
panded, subreflexed,  flesh-colored  ;  columella  wide,  not  deeply  in- 
serted, oblique,  truncated  below.  Alt.  32,  diam.  37  mill. 

Foot  of  Mt.  Data,  near  the  mines  of  Mancdyan,  Western  Cordillera 
of  N.  Luzon,  at  5-6000  ft.  alt. 

C.  dataensis  O.  SEMPER,  Journ.  de  Conchy!,  xiv,  p.  152,  t.  5,  f.  1, 
1866.— C.  SEMPER,  Reis.,  p.  186,  t.  8,  f.  8,  t.  13,  f.  17  (anatomy).— 
ff.  dataensis  PER.,  Monogr.  v,  p.  283. 

The  rudely  sculptured  surface  is  very  characteristic,  and  shows 
this  species  to  be  closely  allied  to  the  group  of  C.  cryptica  rather 
than  to  the  "  Cochlostylce  hypomelanue,"  where  it  is  grouped  by 
Semper. 

Section  VI.     CHROMATOSPH.ERA  Pilsbry. 

Gnippe  der  Cochlostylce  globosce  SEMPER,  Reisen,  Landmoll.,  p. 
181. 

The  shell  is  imperforate,  heliciform,  depressed-globose  solid  and 
opaque,  richly  colored,  but  lacking  hydrophanous  cuticle;  suri'ace 
lacking  spiral  sculpture ;  last  whorl  scarcely  descending  at  the  aper- 
ture. Lip  blunt,  usually  very  narrowly  reflexed  throughout ;  col- 
umella subvertical,  deeply  inserted,  the  umbilico-columellar  area 
covered  with  a  concave  white  callus.  Type,  C.  aurata. 

Anatomy  as  in  typical  Cochlostyla. 

A  small  group  of  rare  shells,  confined  to  northern  Luzon.  It 
is  impossible  to  place  these  species  in  either  Corasia,  Calocochlea  or 
Pfeifferia; ,  they  form  an  independent  group  allied  to  these  sections 
and  to  Pachysphwra. 


1 70  COCHLOSTYLA-CHROMATOSPH^ERA. 

C.  AUBATA  Sowerby.     PL  56,  figs.  68,  69,  70 ;  pi.  26,  figs.  4,  5,  7. 

Imperforate,  depressed-globose,  solid  ;  sulphur-yellow,  with  a  wide 
orange  zone  below  the  suture  and  (generally)  a  dark  blue  line  above 
the  periphery ;  apical  whorl  purple,  the  next  outer  whorl  red. 

The  surface  is  smooth,  lusterless,  lightly  striatulate.  Spire  low, 
apex  plane  ;  whorls  44,  convex,  the  last  a  trifle  descending  in  front, 
constricted  behind  the  lip.  Aperture  slightly  oblique,  wide-lunar, 
white  inside,  showing  a  black-brown  supra-peripheral  band  wider 
than  that  of  the  outside.  Peristome  very  narrowly  reflexed  through- 
out, white ;  columella  arcuate,  sinuous  and  deeply  entering  above, 
spreading  in  a  wide,  concave  white  callus,  the  slightly  elevated  edge 
of  which  bounds  the  umbilical  area. 

Alt.  22'.,  greater  diam.  30,  lesser  24  mill. 

Northern  Luzon  and  Manilla,  Philippines. 

H.  aurata  SOWB.,  P.  Z.  S.  1840,  p.  100. — REEVE,  Couch.  Syst.  t. 
166,  f.  26;  Conch.  Icon.,  f.  46. — PFR.,  Monogr.  i,  p.  260;  Conchy!. 
Cab.,  p.  217,  t.  108,  f.  13,  14.— HIDALGO,  Obras,  t,  24,  f.  7. 

The  outer  lip  is  decidedly  curved  forward  above,  receding  in  a 
peculiar  manner  at  its  upper  insertion.  The  colors  of  this  shell  are 
peculiarly  pleasing. 

Var.  Golden  above,  banded  with  white  at  the  suture,  the  base 
white.  (Pfr.) 

C.  ERUBESCENS  Semper.     PI.  56,  figs.  64,  65. 

Imperforate,  depressed -globose,  thin,  shining,  lightly  striate  and 
irregularly  malleated,  golden,  ornamented  with  a  somewhat  inter- 
rupted blue  band  at  the  suture.  Spire  little  exserted,  very  obtuse, 
red  ;  whorls  4,  a  little  convex,  the  last  much  inflated,  not  descend- 
ing, very  obsoletely  angulated,  coarctate  at  the  margin.  Columella 
straight,  flat,  white.  Aperture  wide-lunar,  oblique ;  peristome  sim- 
ple, narrowly  reflexed,  the  upper  margin  curving  forward. 

Alt.  25,  greater  diam.  33,  lesser  26  mill.     (S.) 

Cabicungaii)  prov.  of  Cagayan,  Luzon. 

C.  erubescens  S.,  Reis.  Landmoll.  p.  182,  t.  9,  f.  8;  t.  10,  f.  12 
(  vur.) 

A  single  example  was  taken  by  my  servant  Antonio  at  the 
northern  point  of  Luzon.  It  stands  rather  near  to  C.  aurata,  but 
differs  in  the  greater  height,  want  of  a  wide  band  at  periphery,  the 
nuilleations  on  the  last  whorl,  and  the  contraction  of  the  whorl  be- 
hind the  lip.  OS'.) 


COCHLOSTYLA-HELICOSTYLA.  171 

Var.  LUTEOCINCTA  Semper.     PI.  56,  figs.  66,  67. 

Pure  white  in  color;  more  globular;  not  malleated.     (&) 

Casiguran,  N.  E.  Luzon. 

C.  LIVIDOCINCTA  Semper.     PI.  56,  figs.  71,  72. 

Imperforate,  globose,  thin,  ornamented  with  a  broad  livid  band, 
and  another  of  brown  at  the  suture ;  spire  scarcely  exserted,  apex 
obtuse,  bluish,  ornamented  with  a  white  cuticular  band.  Whorls  4- 
4t,  nearly  flat,  the  last  much  inflated,  not  descending;  coluinella 
subvertical,  excavated,  white.  Aperture  little  oblique,  lunar ;  per- 
istome  simple,  scarcely  thickened,  not  reflexed. 

Alt.  27,  greater  diam.  32,  lesser  26  mill.     (&) 

Casiguran,  Baler,  Mariquit,  island  of  Luzon,  very  rare. 

C.  liyido-dnctaS.,  Reis.  Phil.  Landmoll.,  p.  182,  t.  9,  f.  9. 

Of  this  remarkable  species  I  have  found  in  all,  only  two  adult 
and  two  young  specimens.  ($.) 

C.  PUDIBUNDA  Semper. 

Subperforate,  conic-globose,  thick,  white;  spire  exserted,  apex 
obtuse,  red  or  reddish -golden.  Whorls  5,  a  little  convex,  the  last 
inflated,  not  descending.  Columella  lightly  arcuate,  vertical,  white. 
Aperture  little  oblique,  rounded-lunar ;  peristome  slightly  thickened, 
not  reflexed.  Alt.  31,  greater  diam.  35,  lesser  28  mill. 

Minanga,  Ambubuk  and  Digollorin,  Luzon. 

C.  pudibunda  SEMPER,  1.  c.,  p.  183. 

Agrees  entirely  with  typical  Cochlostyla  in  anatomy.  This  species 
has  not  been  figured. 

Section  VII.     HELICOSTYLA  Ferussac,  1819. 

As  used  here,  this  section  comprises  a  number  of  minor  groups, 
too  closely  allied  to  be  given  separate  rank  as  primary  sections. 

The  shells  have  generally  a  more  elevated  contour  than  those  in- 
cluded in  Calocochlea,  and  usually  they  lack  hydrophauous  cuticle 
and  spiral  strise. 

It  must  freely  be  admitted  that  the  differences  between  this  group 
and  Calocochlea  are  not  definite  enough  to  permit  one  to  frame 
diagnoses  of  any  practical  value.  In  the  arrangement  of  the  spe- 
cies no  two  authors  agree  in  detail,  thi?  diversity  of  opinion  being 
the  natural  result  of  the  indiscriminate  inosculation  of  characters 
shown  by  the  shells  themselves. 


1  ;_>  ,,-HLOSTYLA-PACHYSPH/ERA. 

In    //.  Ocostyla,  as  iu    Calocochlea,  there  exist  a  number  of  minor 
lpe  deserving  individual  discussion.     They  are  as  follows:  § 


l.     PACHYSPMSRA  Pilsbry. 

Small,  fj/olxil'ir  or  globose-elevated,  solid,  brightly  colored  ;  lacking 
"X  cuticle  and  spiral  striae.     Type,  C.  sphceriea. 

Siil»rrti.»n  ±     OursTiA  Morch,  1852. 

Turl  -mate-globose,  thin,   brightly   colored;    partly   covered  with 
ns  cuticle  ;  no  spiral  striae.     Lip  narrowly  refiexed,  thin  ; 
a  r,Tfical,  not  folded,  deeply  entering.      Type,  C.  monticula. 

•">.     COCHLODRYAS  Martens,  1860. 

Globose-elevated;  colors  brilliant  green  or  brown;  no  hydropha- 
„<>".<  '-utirle;  suture  white-margined;  surface  shining,  having  faint 
>piral  >tri:r  above.  Type,  C.  pplychroa. 

Sul  section  4.     HELICOSTYLA  s.  str. 

'Jlobose,  globose-conic  or  oblong;    colors  not  vivid;    hydroph- 
cuticle  sometimes  present.     Type  C.  mirabilis. 


Subsection  1.     PACHYSPH.ERA  Pilsbry. 

der   Cochlostylce  sphaericce  MARTENS,  Ostas.    Landschu. 
l».  '.'7  -SKMI-KI:,  Reis.  Landmoll.  p.  197. 

A  .-mall  group  of  compact,  solid  little  shells,  confined  to  north- 
\vc-t«-ni  Lu/on. 

Key  to  species. 

a.   Fan-  <>f  lip  dark-edged;  shell  globose 

/-    .Hitural  band  white;  columellar  band  dark  brown;    colu- 
"1(>ll:i  white  sphaerica. 

!>/'.  Mitural  and  rolnm,.llar  bands  green;  columella  reddish 

balteata. 
aa.   Fari-  .if  lip  nut  dark-cd^cd  ;  shell  elevated 

I/ip  distil. rtly  rcflcxr.!  iloconensis. 

l>!>.   Lip  very  hluntly  refiexed  annullata. 

\  Sowerby.     PI.  :;:),  iigs.  82,  83,  84. 

solid,    -lobular;  ground-color    straw-yellow 
*ith  in-.-ular  <,l)li(,ue  yellowish-chestnut  streaks, 


COCHLOSTYLA-PACHYSPHvERA. 

having  a  narrow  white  band  below  the  suture  bordered  by  a  bright 
yellow  band,  a  supra-peripheral  chestnut  line,  and  a  dark  columellar 
area ;  the  lip  edged  with  dark  brown. 

The  surface  is  smooth,  sometimes  slightly  malleated.  Whorls  4J 
to  5,  the  last  deflexed  at  the  aperture  and  decidedly  constricted  be- 
hind the  lip.  The  aperture  is  half-round,  white  inside ;  lip  very 
strongly  thickened  within,  narrowly  reflexed,  white,  edged  with  deep 
brown  ;  columella  heavy,  white. 

Alt.  25,  diam.  23  £  mill. 

Alt.  20,  diam.  19  mill. 

Alt.  23!,  diam.  24  mill. 

Ylocos,  north-ivestern  Luzon. 

H.  splmrica  Sows.,  P.  Z.  S.  1841,  p.  26.— PFR.,  Monogr.  i,  p.  225  ; 
Conchyl.  Cab.,  t.  39,  f.  3,  4. — RVE.,  f.  104. —  Cochlostyla  splmrica 
SEMPER,  Phil.  Archip.,  Landmoll.,  p.  198. 

One  of  the  most  attractive  Helices.  Semper  says  that  the  brown 
line  at  the  periphery  is  wanting  in  one  specimen  among  19  examined. 
I  have  seen  many  more  than  this,  but  in  none  of  them  was  it  lack- 
ing. The  dark  oblique  streaks  are  rarely  so  prominent  as  in  fig. 
82. 

C.  BALTEATA  Sowerby.     PI.  39,  figs.  89,  90,  91,  92,  93. 

Imperforate,  solid,  globose  or  globose-turbinate ;  ground-color 
pale  yellow ;  suture  bordered  by  a  grass-green  band,  sometimes 
split  into  two ;  on  the  middle  of  the  upper  surface  there  is  a  wide 
band  of  olive  or  purplish,  becoming  reddish  on  the  earlier  whorls, 
this  band  often  bordered  below  by  a  supra-peripheral  line  or  zone  of 
darker  color ;  columellar  tract  grass-green.  Columella  red.  Apex 
pink,  rarely  white. 

The  surface  is  smooth.  Whorls  5,  the  last  somewhat  deflexed  in 
front,  slightly  constricted  behind  the  lip.  Aperture  half  round, 
oblique;  lip  somewhat  thickened  within,  very  narrowly  expanded, 
edged  with  purple-brown.  Columella  wide,  pinkish  or  purplish- 
brown. 

Alt.  22,  greater  diam.  20!  mill. 

Ait.  20,  greater  diam.  19  mill. 

Alt.  19,  greater  diam.  21  mill.  (pi.  39,  figs.  91,  92.) 

Provinces  of  South  and  North  Ylocos.  north-western  Luzon. 

H.  balteata  SOWB.,  P.  Z.  S.  1840,  p.  136.— PFR.,  Symb.  ii,  p.  23  ; 
Conchyl.  Cab.,  p.  302,  t,  51,  f.  9-12  ;  Monogr.  i,  p.  226.— RVE., 


]74  -TYLA-PACHYSPHJERA. 

.  ,nch,  Syst.  ii,  t.  166,  f.  25;  Conch.  Icon,  f.  11.— Co  chlo  sty  la 
,,;;;:.  Reis.  Landmoll.,  p.  197,  t.  18,  f.  24  (dentition.) 

Allied  to  C.  sphcerica,  hut  the  ground-color  is  paler,  the  banding 
ni:l;M  :  the  lip  is  narrower  and  the  columella  is  colored. 

Tl.  3  considerable  variation  in  contour,  figs.  91,  92  representing 
the  most  depressed  specimen  before  me.  This  example  has  several 
:i  bands  on  the  base.  The  coloring  of  this  species  is  not- 
ablv  varialile.  The  principal  patterns  are  figured  on  pi.  39. 

Tin-  apical  whorls  are  generally  pink  ;  but  when  they  are  white, 
tin  columella  also  is  light-colored. 

(  .  v  NMI.ATA  Sowerby.     PL  39,  figs.  85,  86,  87,  88. 

I  m  perforate,  solid,  globose-turbinate ;  ground-color  yellow,  usually 
havini:  a  broad  brown  or  purple-brown  zone  at  the  circumference,  a 
pink  line  or  a  yellow  band  at  the  suture,  and  a  streak  of  reddish- 
brown  back  of  the  lip  and  around  the  columella.  lAp  very  little 
i-i'ffi'.ri  tl,  ri-ri/  limit  «n<l  thick. 

The  surface  is  very  lightly  striatulate;  the  banding  is  very  vari- 
able, but  the  ground-color  is  always  yellow.  Apex  obtuse,  buff  or 
pink.  \Yhori-  5,  the  last  a  little  deflexed  in  front.  Aperture  white 
within,  half-round,  oblique;  lip  thickened,  flattened,  its  face  usually 
pink,  its  reverse  red. 

Alt.  25,  diani.  21  mill. 

Alt.  L'n.  <liam.  17  mill. 

I'mi'ince,*  »f  South  and  North  Yloco*,  Luzon,  Philippines. 

II.  annulata  SOWB.,  P.  Z.  S.  1840,  p.  135.— PFR.,  Mon.  i,  p.  227  ; 

'     Qchyl.  Cab.  p.  :;OL>,  t.  51,  f.  5-8.— REEVE,  Conch.  Icon.  f.  110.- 
('<,<•/</,, ..•/,//<!  ,mnit/<ifu  SK.MP.,  Reis.  Landmoll.  p.  197. 

Closely  allied  to  ('.  ilnmucuxis,  differing  mainly  in  the  much 
more  narrowly  rdlcxrd  and  much  more  obtuse  lip. 

Th<-    n-iial   I'onn   (li^.  .SM)   has  a  wide  peripheral  girdle,  a  sub- 

sutnral  line  of  pink  or  kind  of  yellow,  and  back  of  the  lip  there  is 

a  broad   red-brown  streak.       Some  examples  have  no  color-bands, 

ire  In-ill-  bordered  with  white,  the  lip  streaked  as  usual  (fig. 

Specimens  are  als..  found   having  no  color  bands  whatever, 

being  of  a  uniform  yellow  color ;    lip   white,   its  reverse  without  a 

<  MM-  such  is  belb re  me,  and  Semper  mentions  another. 

one  of  the  most,  abundant  snails  of  South  and    North 

^  !  ••• 


COCHLOSTYLA-PACHYSPH^RA.  175 

C.  ILOCONENSIS  Soweiby. '  PL  39,  figs.  97,  98,  99,  100. 

Imperforate,  solid,  high-globose,  the  altitude  decidedly  exceeding 
the  diameter.  Color  varying  from  light  yellow  to  dark  brown  or 
purplish-olive-brown,  the  lighter  forms  having  two  yellow  or  yellow- 
ish-brown bands,  or  one  yellow  and  one  pink  band  ;  the  darker 
forms  having  several  lighter  bands.  Lip  rather  broadly  or  very 
broadly  reflexed,  of  a  uniform  white  or  pinkish  tint. 

The  surface  is  lightly  obliquely  striatulate  ;  whorls  about  5,  the 
last  a  little  descending  and  then  slightly  ascending  at  the  aperture. 
The  aperture  is  oblique,  white  or  pink  inside.  The  lip  is  generally 
broadly  reflexed,  both  lip  and  columella  white  or  light  pink.  The 
columella  is  generally  encircled  by  a  pink  tract. 

Alt.  31,  diam.  23  mill. 

Alt.  20*,  diam.  18  mill. 

Alt.  24,  diam.  23  mill. 

Provinces  of  North  and  South  Ylocos,  Luzon,  Philippines. 

H.  iloconensis  SOWB.,  P.  Z.  S.  1840,  p.  116. — PFR.,  Syrnb.  ii,  p. 
30  ;  Conchyl.  Cab.,  p.  301,  t.  51,  f.  1-4  ;  Alonogr.  i,  p.  223.— REEVE, 
Conch.  Icon.,  f.  109. —  Cochlostyla  iloconensis  SEMP.,  Reis.  Phil.  Laud- 
moll,  p.  197,  t.  18,  f.  10  (dentition.) 

Closely  allied  to  C.  annulata,  but  appreciably  wider  in  the  spire, 
having  a  more  broadly  reflexed  lip  and  probably  never  having 
exactty  the  same  color-patterns  as  that  species.  On  many  specimens 
there  are  scattered  dots  of  a  pale  or  purplish  tint  (fig.  98.) 

All  of  the  specimens  I  have  seen,  over  30  in  number,  maybe 
referred  to  the  following  three  color-varieties : 

(1)  Var.  heterotwniata.     Shell  very  solid,    light  buff,   having  a 
supra-peripheral  yellow  band  and  a  subsutural  pink  band ;  a  streak 
back  of  the  broadly   reflexed  lip,    and    a  circum-columellar   patch 
pink.     Both  of  the  bands  are  sometimes  yellow.     The  apex  is  pink. 

(2)  Var.  xanthotceniata.     Shell  thinner  ;  lip  much  narrower,  thin  ; 
pale  yellow,  having  subsutural  and  peripheral  bands  of  yellow.    No 
streak  behind  the  lip,  no  columellar  patch.     The  apex  is  white. 

(3)  Var.  intensior  (fig.  100.)     Shell  solid,  lip  wide  ;  ground-color 
olive-brown  or  purplish-brown  ;  light  bands  at  periphery  and  suture, 
darker  bands  in  the  usual  positions,  sometimes  obsolete  on  the  body- 
whorl.     Apex  purple. 


1  7 1  i  COCHLOSTYLA-ORUSTIA. 

Subsection  2.     ORUSTIA  Morch,  1852. 

0rtw*ta  MORCH,  Catal.  Yoldi,  p.  15   (in  part;    first  species  H. 
nt'intlriif-i  Suw.). — SKMI-KR  (group  of  Cochlostyla),  Reis.  Phil.  Laud- 
.  p.  196. 

Shdl  imperforate,  turbinate-globose,  rather  thin,  green  under  a 
iilmnnus  yellow  cuticle,  the  base  green.  Lip  thin,  narrowly 
reflexed;  culumella  vertical,  deeply  entering;  the  umbilico-colu- 
iin-llar  tract  notably  concave,  covered  with  a  white  callus.  Surface 
wholly  free  from  spiral  sculpture. 

Anatomy  as  in  Cochlostyla  generally. 

The  entire  absence  of  spiral  sculpture  and  the  presence  of  hy- 
tlrophanous  cuticle  are  the  more  important  characters  of  the  sub- 
trochiform,  thin-lipped  species  to  which  the  name  Orustia  is  restricted 
by  recent  authorities. 

( '.  MONTICULA  Sowerby.     PI.  39,  figs.  94,  95,  96. 

Imperforate,  globose  turbinate,  rather  thin,  lightly  obliquely  stri- 
at u late.  Upper  surface  rich  yellow  and  cream-colored,  fading  into 
white  on  the  earlier  whorls  through  loss  of  the  cuticle ;  periphery 
girdled  by  a  dark  brown  band,  below  which  is  a  wider  yellow  band  ; 
ki-c  bright  green. 

There  is  no  trace  of  spiral  sculpture.  The  hydrophanous  cuticle 
is  rich  yellow,  sometimes  almost  orange,  on  the  last  whorl, 
In  •coming  lighter  above,  and  lost  by  erosion  on  the  earlier  whorls. 
The  dark  girdle  at  the  circumference  is  constant  throughout  the 
lar-v  x-rios  which  I  have  examined.  There  are  5  slightly  convex 
\\horU,  the  last  not  deflexed  in  front,  not  angled  at  the  periphery, 
although  the  base  is  rather  flattened.  The  apex  is  quite  obtuse. 
Aperture  oblique,  half-round,  white  within,  faintly  showing  the  brown 
band.  The  lip  is  very  narrowly  subreflexed,  white;  columellasub- 
v.-rtical,  (Icrply  inserted,  dilated  over  the  umbilico-columellar  tract 
in  a  white  concave  callus,  somewhat  as  in  C.  (Chromatosplmra) 
aurata. 

Alt.  20,  irrcater  diam.  1!).]  mill. 

Alt.  1-T,  Lfreater  diam.  10  mill. 

//.  monticula  Snwi-:i:i;y,  p.  /.  S.  1840,  p.  167.— PFR.,  Monogr.  i, 

p.  226;  Conchyl.  Cab.  t.  51,  f.  13-17.— KVE.,  f.  103.—  Cochlostyla 

i  monticula  SKMIMOI:,  /.  c.,  p.  197,  t.  8,  f .  6 ;   t.  13,  f .  1  ;    t. 

i'-  -'•>  '  iinatoiny;. 


COCHLOSTYLA-COCHLODRYAS.  177 

Easily  recognized  by  the  yellowish  upper  surface,  banded  with 
brown  at  the  circumference,  and  bright  green  base. 

Of  this  species  Semper  says:  I  found  this  species  exclusively  in 
the  wide  valley  of  the  Rio  Grande  de  Cagayan,  in  northern  Luzon  ; 
and  as  well  at  the  village  Aparri,  hard  by  the  valley's  mouth,  as  in 
the  upper  mountainous  course,  on  the  western  side  of  the  eastern 
Cordillera,  in  the  valleys  of  the  rivers  Ilagon  and  Catalangan,  at 
Satpat,  Ilagan  and  Minanga.  The  shell  varies  much  .  .  .  One 
example  from  Ilagan  before  me  is  entirely  green,  with  a  sulphur- 
yellow  band  at  the  periphery.  Another  from  Satpat  has  a  second 
broad  brown  band  upon  the  upper  surface,  and  the  periphery  is 
girdled  by  two  narrow  dark  yellow  bands,  strongly  contrasted  with 
the  clear  yellow  of  the  upper  surface.  The  colors  of  the  shell  fade 
very  readily.  I  suspect  that  Pfeiffer's  vat:  ft  having  a  white  shell, 
is  founded  upon  a  bleached  example. 

The  anatomy  is  as  in  typical  Gochlostyla. 

Subsection  3.     COCHLODRYAS  Martens,  1860. 

Cochlodryas  MTS.  in  Albers,  Die  Hel.  (2d  edit.),  p.  176,  (Type, 
C.  polychroa). — SEMPER,  Reis.  Landmoll.,  p.  198. — Poedlus  ALB. 
ms.,  preoc. 

Elevated,  vividly  colored  shells,  lacking  hydrophanous  cuticle, 
and  with  a  distinct  sutural  band  which  is  generally  white. 

The  Madagascar  species  H.  viridis  Dh.  has  been  included  here, 
but  its  place  is  in  the  vicinity  of  Ampelita  (q.  v.) 

C.  FLORIDA  Sowerby.     PI.  38,  figs.  70,  71,  72,  73. 

I m perforate,  solid,  globose-elongated  ;  ground-color  grass-green 
(sometimes  olive),  having  a  very  narrow  white  band  at  the  suture, 
and  frequently  some  brown  bands,  of  which  one  is  below  the  sutural 
white  line,  one  at  the  columella,  and  two  encircle  the  body-whorl 
near  the  middle.  Some  or  all  of  these  brown  bands  are  often  lack- 


ing. 


The  surface  has  light  growth-lines,  and  under  a  lens  faint  fine 
spiral  impressed  lines  are  seen  on  the  upper  whorls.  The  grass- 
green  or  olive  ground  color  fades  into  white  or  sometimes  pink  on 
the  earlier  whorls.  Whorls  nearly  6,  the  last  scarcely  deflexed  in 
front.  Aperture  white  within,  oblique;  lip  reflexed,  white;  col- 
umella subvertical,  slightly  arcuate. 

Alt.  37,  diam.  29  mill. 

Alt.  36,  diam.  26  mill. 

12  Mindoro,  Philippines. 


1  ;  v  COCHLOSTYLA-COCHLODRYAS. 

Jf.  florida  BOWB.,  P.  Z.  S.  1840,  p.  87.—  REEVE,  Conch.  Syst.  ii, 
t.  163,  f.  2;  Conch.  Icon,  f.  43.—  PFB.,  Mon.  i,  p.  222;  Conchyl. 

1...  p.  L'(.».I,  t.  50,  f.  1,  2—Bulimus  helicoides  PFR,  Zeitschr.  f. 
Mali'  1849,  p.  176;'  Conch.  Cab,  Bui,  t.  36,  f.  7,  8.—  RVE,  Conch. 

I-  'ii.,  t'.  72. 

\  brilliant  Lrreen  shell,  having  the  ground-color  less  streaked  and 
tin-  sutiinil  white  hand  narrower  than  C.  polychroa.  The  green  is 
sometimes  changed  to  olive. 

C.  i'"i.y<  IIROA  Sowerby.     PI.  38,  figs.  74,  75,  76,  77,  78. 

Impi-rforate,  globose  or  globose-elongated,  rather  thin  ;  conspicu- 
o//x/;/  i,l>/i<]uely  streaked  with  grass-green  and  whitish,  having  a  wide 
//•///'/,  (  rarely  red)  band  at  the  suture,  and  frequently  banded  with 
brown. 

Surface  smooth,  lightly  obliquely  striatulate,  the  earlier  whorls 
having  microscopic  spiral  strue  when  not  worn.  Whorls  nearly  6, 
convex  :  apex  obtuse.  Aperture  white  within,  oblique.  Peristome 
rather  narrowly  reflexed,  white;  columella  vertical,  nearly  straight, 
perceptibly  truncated  at  its  base. 

Alt.  43,  diam.  31  mill. 

Alt.  4:;,  diam.  34  mill. 

Alt.  •">•'),  diam.  29  mill,  (shortest  specimen). 

Alt.  39,  diam.  2!)  mill,  (most  elongated  specimen). 

Island  of  Temple,  near  Burias,  Philippines. 


H.  pnlychrnti  SOWB,  P.  Z.  S.  1840,  p.  87.  —  PFR,  Monogr.  i,  p. 
•_"_'  1  :  Conchy!.  Cah.  p.  295,  t.  50,  f.  13-15.—  RVE,  Couch.  Icon.  f. 
44.—  DII.  in  I-Y-r.  Hist.  t.  104  A,  1-4;  110  A;  f.  3-6.—  H.  galactites 
fir.  /,ictnr<it<i  (  IIIAT,  Actes  Bord.  xi,  p.  163.  —  ?  Helicostyla  smarag- 
•  In.--  r>  i  <  K,  Index  p.  37  (no  desc.).  —  Bidimus  virido-striatus  LEA, 
Trans.  Amer.  Philos.  Soc.  1841,  p.  455,  t.  11,  f.  2.  —  Cochlostyla 
1  •'•li/,l<lrij,i.-<)  polychroa  SEMPER,  Reis.  Landmoll.  j).  199. 

The  Hitural  hand  is  wider  than  that   of   C.  florida,  the  shell  is 
thinner,  and  the  columella  more  distinctly  truncated  at  its  base. 

The  color-forms  of  thU  beautiful  species  are  numerous,  the  princi- 
pal pattern.-  lieinu-  these  : 

<  ire,  ii  ami  white  streaked,  bandless  except  for  a  wide  sutural 
wliii«-  zsone  >  ii.Lr.  70). 

1'he  same,  luit.  sutural  /one  red,  a  brown  patch  around  the 
columella. 


COCHLOSTYLA-COCHLODRYAS.  179 

(3).  Green  and  white  streaked,  sutural  band  white,  having  two 
brown  zones  around  the  middle,  the  upper  ascending  the  spire,  the 
columellar  patch  dark.  There  is  often  an  additional  brown  line  or 
band  below  the  white  sutural  band  (figs.  77,  78). 

(4).  Green  largely  replaced  by  reddish  brown  (fig.  75),  or  by 
blackish-brown  (fig.  74).  Sometimes  greenish-black, 

C.  ORBITULA  Sowerby.     PL  58,  figs.  98,  99. 

Imperforate,  ovate-globose,  thick  and  solid,  smooth ;  light  straw- 
colored  with  bands  of  brown  and  green,  the  former  retained,  the 
latter  lost  when  the  epidermis  is  removed.  The  suture  has  an 
opaque  white  margin  below,  about  1*  millim.  wide  on  the  last  whorl. 

Spire  dome-shaped  ;  apex  obtuse  ;  sutures  superficial.  Whorls  5 
to  5J,  slightly  convex,  the  last  slightly  descending  in  front.  Aper- 
ture rather  small,  oblique  ;  lip  expanded,  reflexed,  white  ;  columella 
heavy,  slightly  arcuate,  rarely  dentate  below. 

Alt.  30,  diam.  26  mill. 

Alt.  26,  diam.  26  mill. 

Mindoi'o,  Philippines. 

H.  orbitula  SOWB.,  P.  Z.  S.  1840,  p.  103.— PFR.,  Monogr.  i,  p. 
224  ;  Conchyl.  Cab.,  p.  296,  t.  50,  f.  7,  8.— REEVE,  Conch.  Icon.,  f. 
60. — DH.  in  Fer.,  Hist,  i,  p.  311,  t.  108 A,  f.  7,  8. — H.  chlorogrammica 
VAL.,  in  Paris  Mus. 

The  shell  agrees  with  C.florida  and  C.  polychroa  in  possessing  a 
white  margin  below  the  suture,  but  it  is  much  more  compact  and 
solid  than  either  of  these  species.  It  is  a  more  solid  shell  than  C. 
tenera. 

The  coloring  is  very  variable.  There  are  generally  (?)  two  chest- 
nut-brown bands,  one  of  which  ascends  the  spire,  but  these  are  often 
lacking  on  the  body-whorl.  The  green  may  either  cover  most  of 
the  body-whorl,  or  be  restricted  to  a  narrow  band  or  two. 

C.  TENERA  Sowerby.     PI.  36,  figs.  31,  32. 

Imperforate,  globose-conic,  thin,  smooth  ;  white,  ornamented  with 
blackish  and  green  bands;  subhyaline;  spire  depressed-conoid. 
Whorls  4J,  a  little  convex ;  columella  almost  straight,  thin, 
excavated ;  aperture  lunate-rounded ;  peristome  simple,  little  ex- 
panded. Alt.  20,  greater  diam.  25,  lesser  21  £  mill.  (Pfr.) 

Mindoro  and  Tablas,  Philippines. 


1  xi  i  ,  ,  .CHLOSTYLA-COCHLODRYAS. 

//.  tenera  Sows.,  r.  /.  S.  1840,  p.  102.—  PFR.,  Conchyl.  Cab.,  p. 
293,  t.  4!».  1'.  12,  1:>:  M»>nogr.  i,  p.  228.  —  REEVE,  Conch.  Icon.,  f. 
62. 

A  IIIMIV  -lohose  shell  than    (.".  ignobilis,  with  shorter  spire,  more 
vrx  whorls  and  differently  formed  lip. 


(  '.  K.xonii.is  Snwerby.     PI.  33,  fig.  65. 

I  m  perforate,  rather  solid,  conical.  White  under  a  very  thin  light 
hulf  cuticle,  with  two  narrow  chestnut  bands,  one  above,  one  below 
i  In-  periphery,  and  a  dark  columellar  tract,  often  very  small. 

Surface  lightly  striatulate,  the  earlier  whorls  minutely,  obsoletely 
-pi  rally  striated. 

Spin-  distinctly  conical,  the  lateral  slopes  slightly  convex.  Sut- 
ures but  little  impressed.  Apex  obtuse,  but  more  acute  than  usual 
in  the  present  group.  Whorls  5i,  very  slightly  convex,  the  last 
unified  at  the  periphery  ;  base  rather  flattened.  Aperture  oblique, 
ovate  :  lip  expanded  and  reflexed,  white.  Columella  subvertical, 
-'•itivcly  arcuate. 

Alt.  :}:},  diam.  2-U  mill. 

Alt.  28,  diam.  25  mill. 

Romblon  and  Tablets,  Philippines. 

II.  ignobitis  Sown.,  P.  Z.  S.  1840,  p.  102.—  PFR.,  Monogr.  i,  p. 
228;  Conchyl.  Cab,  p.  292,  t.  49,  f.  11.—  REEVE,  Conch.  Icon.,  f. 
57. 

The  very  distinctly  conical  spire  of  this  species  is  a  notable  feature. 
Tin-  proportions  of  length  and  breadth  vary  considerable.  The 
l>;i!idiii'_r  is  usually  as  described  above,  but  it  varies  as  follows  : 

1.   Bifasciatc,  the  lower  band  wider  (fig.  65.) 

_.   rpper  hand  wanting. 

•'!.   Body-whorl  not  banded. 

C.  BOETTGERIANA  Mollendorff.      UnfigumL 

[mperforate,   globose-conic,   rather  solid,   transversely    striated, 

•iilptiin.l   with   minute  spiral  lines  and  forward-descending  fold- 

kvriiiUe<;  white,  covered  with  an  olive-buff  shining  deciduous 

iuticle,  and  eurii-cled  by  two  rather  wide  bands.      Spire  high  conic, 

:  i-:>tlM-r  obtuse.     Whorls  oi,  slightly  convex,  the  last  a  little 

sometimes  ornamented  with  a  third  band  around  the 

suture   broadly    margined,    white,    opaque,    crenulated. 

>hli<l'"-<    ample,     lunate-oval;     peristome    white-li}>ped, 


COCHLOSTYLA-HELICOSTYLA.  181 

widely  expanded  and  reflexed,  the  outer  margin  sinuous  ;  columella 
rather  straightened,  excavated  above,  dilated,  appressed.  (Moll.) 

Alt.  50,  greater  diam.  46^  lesser  36  mill.;  aperture  (with  perist.) 
36  mill,  long,  31  wide. 

High  mts.  on  the  boundary  between  the  provinces  Bulacan,  Manilla 
and  Morong. 

C.  (Helicostyla)  boettgeriana  v.  MOLL.,  Nachr.  D.  M.  Ges.  1888, 
p.  101. 

Belongs  to  the  group  of  C.  mirabilis  Fer.  The  acute  conic  spire, 
the  broad  dull  white  crenulated  suture,  the  strong,  almost  fold-like 
oblique  wrinkles,  separate  it  from  all  species  known  to  me.  (Moll.) 

Subsection  4.     HELICOSTYLA  Fer.,  1819. 

Helicostyla  FER.,  Prodr.  p.  46  (in  part). 

This  section  serves  to  contain  the  oblong  or  globose-conic  species 
remaining  after  the  segregation  of  the  various  allied  groups.  It  is 
consequently  rather  miscellaneous  as  to  contents  and  characters, 
but  perhaps  less  so  than  in  previous  works,  where  it  has  been  made 
still  more  comprehensive. 

(1.)    Group  of  C.  mirabilis. 

C.  MIRABILIS  Ferussac.     PL  57,  figs.  79-86  ;  pi.  31,  fig.  40. 

Imperforate,  rather  solid,  globose ;  ground-color  unicolored  yelloiv 
or  brownish-yellow,  without  bands,  or  variously  banded  with  brown 
or  black-brown,  the  bands  when  present  being  all  of  nearly  the  same 
color,  and  generally  4  in  number;  a  narrow  sutural  line,  two  wider 
zones  girdling  the  middle  part,  and  a  columellar  patch. 

Surface  nearly  smooth,  lightly  striatulate.  Bands  wide,  narrow 
or  none,  and  when  present  either  blackish-brown  or  reddish-brown 
in  color.  Whorls  nearly  5,  the  last  but  little  descending  in  front. 
Aperture  oblique,  white  inside  and  showing  the  bands  ;  lip  broadly 
reflexed,  white,  its  reverse  usually  snow-white  also;  columella  sub- 
vertical,  gently  arcuate,  very  deeply  inserted,  and  visibly  twisted 
near  the  insertion. 

Alt.  36,  diam.  34  mill,  (average  specimen.) 

Alt.  40,  diam.  29  mill,  (elongated  specimen.) 

Alt.  30,  diam.  32  mill,  (short  specimen.) 

Southeastern  and  middle  Luzon;  Islet  of  Alabat,  off  E.  coast 
Mauban  ;  Marinduque,  Philippines. 


COCHLOSTYLA-HELICOSTYLA. 

//  mirabUis  FKR.,  Prodr.  320  ;  Histoire,  t.  31,  f.  4-6.-SowB.,  P. 
z>g  L841,  p.  2.--PFR.,  Conchyl.  Cab.,  p.  288,  t.  48,  f.  5,  7,9,10; 
M,,i,,,_rr.  i,  p.  220.— RVE.,  Conch.  Icon,,  f.  45.— H.  fonnosa  WOOD, 
[ndex,  Buppl.,  t.  7,  f.  10.— H.  galactites  Lam.,  An.  s.  Vert,  iv,  p.  69. 
—  77.  pertimilis  DII.  in  Fer.  Hist,  p.  120,  t.  HOB,  f.  2.—Cochlo8tyla 
mirabilia  SEMPER,  Reis.,  p.  192.— HIDALGO,  Obras,  t.  49,  f.  1-7. 

A  globose  shell,  generally  a  little  higher  than  broad,  and 
excessively  variable  in  both  contour  and  pattern  of  coloring.  The 
surfu<v  i-  "ttni  somewhat  malleated.  The  ground-color  varies  from 
:1  v-rv  li-ht  buff  to  a  yellowish-brown.  It  is  allied  to  C.  fulgens, 
dirt'ei-ini:  mainly  in  the  pattern  of  coloring. 

Sliirht  traces  of  whitish  hydrophanous  cuticle  are  observable  near 
the  suture  in  some  specimens. 

Tin-  principal  varieties  of  banding  are  these: 

(1.)  Unicolored  yellow;  the  earlier  whorls  white  or  having  a 
l-m-plc  supra-sutural  band  (fig.85).  The  earlier  whorls  are  some- 
times pink. 

(2.)  Yellow  or  buff,  with  a  narrow  sutural  brown  line,  a  sub- 
peripheral  brown  band,  and  a  columellar  patch.  There  are  some- 
times numerous  yellow  lines  also  (fig.  79.) 

(3.)  Yellow  or  yellowish-brown,  with  4  wide  or  narrow  bands, 
'•Hi-  columellar,  another  sutural,  with  two  intermediate.  This  is  the 
c'Uiinionest  pattern  (figs.  81,  86.) 

I.     Yellow  with  reddish  bands,  Marinduque,  (fig.  80.) 
").)  Entire  shell  very  dark  (fig.  84.) 

Ti-o-it* if  ion  forms  to  var.fiilyens. 

I'nrm  fr!c]ti-on  (pi.  31,  fig.  38).  Having  4  wide  or  narrow  black- 
Mi-l>rown  hands,  and  three  yellowish-brown  bands  in  their  intervals. 
The  spe -'linen  figured  is  from  Albay,  Philippines.  The  figure  on 
pi.  ill  is  united  by  error  with  that  of  C.  tephrodes,  below  it. 

Another  deviation  has  the  three  brown  or  yellowish-brown  bands, 
but  no  blackish  hands  (pi.  ~>7,  fig.  87). 

A  very  lari_re  series  of  these  transition  forms  is  before  me. 

V\i;.  PDLGENS  Sowerby,     1M.  38,  figs.  79,  80,  81. 

Imperforate,   globose   or   globose-turbinate,   solid;    ground-color 

bove  the  periphery,  yellowish-brown  below  it,  having  normally 

blackish-brown  hands,  one  at  suture,  one  at  columella,  the  other 

encircling  the  body-whorl  at  equal  distances;  the  sutural  band 

'ii  absent,  and  sometimes  additional  bands  are  developed  on  base 


COCHLOSTYLA-HELICOSTYLA.  183 

and  at  periphery ;    and  there  is  usually  a  brown  border  below  the 
suttiral  and  above  the  supraperipheral  bands. 

Surface  smooth,  shining.  Whorls  5,  the  last  but  little  descending. 
Aperture  oblique,  white  within  ;  lip  thin,  narrowly  reflexed,  white ; 
columella  subvertical,  straight,  white. 

Alt.  32,  diam.  29  mill. 

Alt.  35,  diam.  28  mill. 

Mindoro,  Philippines. 

H.fulgens  SOWB.,  P.  Z.  S.  1841,  p.  3.— PFR.,  Monogr.  i,  p.  221  ; 
Conchyl.  Cab.  p.  287,  t.  48,  f.  3,  4.— KEEVE,  Conch.  Icon.  f.  31. 

This  is  typically  quite  distinct  in  coloring  from  C.  mirabilis,  but 
every  possible  stage  between  the  two  is  shown  in  the  series  before 
me.  That  it  is  idle  to  attempt  to  draw  specific  lines  in  such  a  case, 
I  am  convinced  by  the  examination  of  a  large  suite  of  shells. 

C.  TEPHRODES  Pfeiffer.     PI.  31,  fig.  39. 

Imperforate,  subglobose,  solid,  ponderous  ;  rufous  above,  nearly 
covered  by  wide  streaks  of  hydrophanous  ashen  cuticle,  the  base 
straw-colored,  banded  with  rufous.  Whorls  4i,  convex,  the  upper 
depressed,  rounded,  rufescent;  columella  wide,  white,  subarcuate. 
Aperture  lunate-subcircular,  white  inside;  peristome  simple,  re- 
flexed.  Alt.  30,  greater  diam.  37,  lesser  33  mill.  (-Pfr.) 

Prov.  Pangasinon,  Luzon,  Philippines. 

H.  tephrodes  PFR.,  P.  Z.  S.  1842,  p.  151  ;  Monogr.  i,  p.  254.— 
REEVE,  Conch.  Icon.,  f.  112. 

VAR.  Alternately  and  broadly  zoned  with  rufous  and  yellow 
under  a  streaked  cuticle.  (P/V.) 

This  variety  is  that  figured  by  Reeve  and  copied  on  pi.  31. 

C.  PLURIZONATA  Adams  &  Reeve.     PI.  59,  figs.  4,  5. 

Subglobose,  covered-perforate,  white,  clouded  with  pale  brown 
spots,  striate- roughened ;  encircled  by  several  fawn  and  purple 
bands.  Whorls  4,  convex,  the  last  rotund.  Aperture  lunate-trans- 
verse, brown  inside ;  peristome  strongly  reflexed,  white,  lower  mar- 
gin somewhat  calloused.  (A.  &  J?.) 

Mindanao,  Philippines. 

H.  plurizonata  A.  AD.  &  RVE.,  Voy.  Samarang,  Zool.,  Moll.,  p. 
62,  t.  16,  f.  9. — REEVE,  Conch.  Icon.,  f.  528. — PFR.,  Mon.  iii,  p. 
182. 


j  x  I  COCHLOSTYLA-HELICOSTYLA. 

V 

(2).     Group  of  C.  collodes. 

C,  .  OLLODES  Sowerby.     PI.  46,  fig.  56,  57. 

Iinperforate,  solid,  depressed -turbin ate,  the  base  distinctly  flat- 
tened. Surface  polished,  straw-yellow,  the  earlier  whorls  pink,  sut- 
UIV  bordered  below  by  a  reddish-brown  line.  Lip  expanded,  its 

tan-  and  reverse  black. 

The  form  is  somewhat  trochoidal,  base  obviously  flattened  ;  sur- 
face shining,  obliquely  lightly  striatulate.  The  color  is  a  bright 
>tra\\ -yellow.  Whorls  5J,  convex,  the  last  not  deflexed  in  front. 
Apt -rture  wide,  oblique,  white  inside;  columella  subvertical,  deeply 
entering,  arcuate  below. 

Alt.  27-1,  diam.  30  mill. 

Alt.  23,  diam.  26J  mill. 

Xear  Boljoon  and  Sambuan,  southern  Zebu,  Philippines. 

H.  collides  Sown.,  P.  Z.  S.  1840,  p.  102.— PFR.  Monogr.  i,  p. 
2:>3;  Conchy!.  Cab.  p.  216,  t.  108,  f.  7,  8.— RVE.,  Conch.  Icon.  f. 
Q4,—Cochlo8tyla  (Helicostylci)  collodes  \.  MOLL.,  Landschn.  Cebu, 
p.  2:59. 

Notable  for  its  bright  surface  entirely  free  from  hydrophanous 
cuticle,  its  flattened  base,  and  black  lip.  Mollendorff  gives  meas- 
urement-; of  larger  specimens  than  those  before  me:  alt.  30,  diam. 

364  mill. 

Variety.  Whorls  more  convex,  especially  beneath ;  no  brown 
Hitural  line;  lip  rather  more  broadly  expanded,  reddish-brown  in- 
>t»a«l  <>f  black. 

Alt.  27,  diam.  30  mill. 

Alt.  25,  diam.  28  mill. 

This  form  is  not  uncommon  ;  several  are  before  me. 

• 

\'»,-ii'hj.     Unicolored  blackish-chestnut. 

A  very  ran;  color-form,  recorded  by  Pfr.  and  v.  Moll. 

Tin  localities  given  by  Pfeiffer  and  others  are  very  doubtful.  V. 
MollemlorH1  found  it  in  Zebu,  and  upon  this  island  my  own  ex- 
ani|ih-  were  recently  collected  by  Mr.  E.  L.  Moseley. 

C.  [NDUSIATA  IT.-ifli-r.     1M.  46,  figs.  53,  54. 

[mperforate,  depressed-turbinate,  rather  solid,  nearly  smooth; 
flesh-colored,  covered  \\ith  a  tawny  epidermis  with  pallid  bands, 
havinu  a  Miiural  bum!  of  dark  chestnut,  another  at  the  periphery 
a  columellar  area  of  blackish.  Spire  short,  conoid-convex. 


COCHLOSTYLA-HELICOSTYLA.  185 

Whorls  4,  scarcely  convex,  regularly  increasing,  the  last  scarcely 
descending  in  front,  distinctly  carinated  at  the  periphery,  base 
more  convex.  Aperture  oblique,  subrhombic-lunar,  whitish  inside, 
with  translucent  bands.  Coluraella  oblique,  compressed,  subexca- 
vated  ;  peristome  blackish,  margins  subparallel,  the  upper  expanded, 
basal  shortly  reflexed.  (P/r.)  Alt.  18,  diam.  '30  mill. 

Tukan  Bessi  Island. 

H.  indusiata  PFR.,  Mai.  Bl.  1871,  p.  121  ;  Novit.  Conch,  iv,  p. 
71,  t.  121,  f.  3,  4;  Monogr.  vii.  p.  355. 

Specimens  of  this  species  received  through  the  kindness  of  MR. 
J.  H.  THOMSON  are  before  me.  The  surface  where  the  cuticle  is  re- 
moved, is  opaque-white,  faintly  tinted  with  pink.  The  cuticle  is 
straw-yellow,  \vith  a  narrow  sutural  brown  line,  extending  both 
above  and  below  the  actual  suture,  the  portion  above  the  suture  be- 
coming more  prominent  upward,  and  extending  to  the  apex.  At 
the  periphery  there  is  a  brown  line,  scarcely  visible  on  one  speci- 
men before  me,  the  same  specimen  lacking  the  columellar  dark 
tract,  which  is  quite  large  on  the  other  shell,  of  a  purple-brown 
color,  and  staining  the  greater  part  of  the  columella.  There  is  a 
very  thin  coat  of  hydrophanous  cuticle  above,  with  some  slightly 
more  distinct  creamy  spiral  lines.  Below  the  periphery  there  is  a 
wide  cream-white  zone,  and  the  base  shows  a  number  of  narrow 
bands  of  the  same  tint. 

C.  THOMSON:  Pfeiffer.     PJ.  29,  figs.  1,  2,  3,  4. 

Imperforate,  solid,  depressed  turbinate-semiglobose;  shining, 
lightly  striatulate  ;  bright  straw-yellow,  the  earlier  whorls  pale  pink- 
ish ;  suture  bordered  below  by  a  brown  line.  Lip  expanded,  its 
face  and  reverse  blackish-brown.  Whorl  not  deflexed  at  aperture. 

The  form  is  more  depressed  than  any  specimens  of  C.  collodes  I 
have  seen,  but,  as  in  that  species,  the  base  is  rather  flattened. 
Whorls  5,  convex,  the  last  very  obsoletely  angulated  in  front  of  the 
aperture.  Mouth  white  inside,  quite  oblique ;  lip  expanded,  deep 
brown.  Columella  arcuate,  deeply  entering. 

Alt.  22*,  diam.  30  mill. 

"  Ttikan  Bessi  Island  "  (J.  H.  Thomson.) 

H.  thomsoni  PFR.,  Mai.  Bl.  1871,  p.  120  ;  Novit.  Conch,  iv,  p.  70, 
t.  121,  f.  1,2. 


186  COC II  LOST  Y  L  A-H  ELICOST  Y  LA . 

This  shell  has  the  greatest  resemblance  to  C.  collodes  of  the  island 
/.I. u.  diilerini:  «»nly  in  being  a  little  more  depressed  than  any  spec- 
imens I  have  seen  of  that  species.  Under  the  yellow  cuticle  it  is 

white. 

I  il.i  n.it  know  the  location  of  the  island  Tukan  Bessi. 

41 

(3).      Group  of  C.  metaformis. 

C.  METAFORMIS  Ferussac.     PL  36,  figs.  25,  26,  27,  28,  29.  ] 

Imperforate,  solid,  oblong-ovate,  elevated,  smooth,  varying  from 
buff  to  chestnut-yellow,  unicolored  or  having  dark  chestnut  bands 
at  suture,  periphery  and  columella,  and  frequently  another  on  the 
base,  any  or  all  bands  often  absent,  but  the  columellar  band  most 
constant.  Generally  covered  with  a  hydrophanous  buff-white  cuticle 
worn  into  oblique  shreds. 

Spire  elevated,  the  apex  very  obtuse.  Whorls  6,  the  last  descend- 
ing in  front.  Aperture  oval,  oblique,  white  inside;  lip  white,  nar- 
row, slightly  expanded,  thickened  ;  columella  vertical,  white,  spread- 
ing in  a  shining  white  callus;  slightly  arcuate  above,  more  or  less 
strongly  toothed  below. 

Alt.  43,  diam.  31  mill,  (average  specimen.) 

Alt.  54,  diam.  33  mill,  (large  specimen.) 

Alt.  32,  diam.  25  mill,  (small  specimen.) 

Bohol ;  middle  Luzon.. 

II.  metaformis  FEE.,  Hist.  t.  108,  f.  2. — PER.  in  Conchyl.  Cab.  p. 
289,  t.  49,  f.  1-5;  Monogr.  i,  p.  222.— KEEVE,  f.  IQ.—  Cocldostyla  • 

metaformis  SKMP.,  Reis.  p.  192,  t.  13,  f.  16;  t.  18,  f.  21  (anatomy). 
HII>AL<;O,  Journ.  de  Conchyl.  1887,  p.  149. — Bulimus  metaformis 
<  M:  \v,  tt  <il. — Helix  ovularis  MKE.,  Catal.  Malsb.  p.  4. 

An  oblong  species,  allied  to  C.  rusticaa.nd  especially  C.  hydrophana, 
the  latter  being  scarcely  more  than  a  variety.  The  principal  varia- 
tions in  contour  and  banding  are  shown  on  my  plate. 

Then-  is  an  uncommon  form  having  a  chestnut  rather  than  yellow 
mid-color,  and  a  brown  lip.  A  large  specimen  of  this  is  shown 
in  li-.  25. 

I  am  inclined  to  consider  the  0.  rustlca  of  Hidalgo   (not   Mous- 
"ii  i  as  a  color  variety  of  this  species.     See  pi.  58,  fig.  89.     Several 

-  com •sj.onding  to  this  figure  are  before  me. 
A  form  very  closely  allied  to  the  true  C.  rustica  is  figured  on  pi. 

M). 

Another  n-, table  color-form  is  illustrated  on  pi.  58,  fig.  88. 


COCHLOSTYLA-HELICOSTYLA.  187 

C.  RUSTICA  Mousson.     PL  58,  figs.  91,  92. 

Imperforate,  oblong,  solid,  striate,  chestnut-colored,  covered  with 
a  gray  cuticle. 

Spire  elongate-conic,  summit  obtuse,  nude.  Whorls'  6,  slightly 
convex,  pallidly  unifasciate,  the  last  two-fifths  of  the  total  length, 
subinflated,  ornamented  with  a  black  and  a  buff  band  around  the 
umbilical  region.  Aperture  oblique,  lunate- circular,  milk-white 
within ;  peristome  narrowly  expanded,  subthickened ;  columella 
immersed,  white,  subcallous,  ^asi-unideutate.  (Mouss.^) 

Alt.  53,  greater  diam.  37,  lesser  32  mill. 

Eastern  Java. 

Bui.  rusticus  Mouss.,  Moll.  Java,  p.  114,  t.  22,  f.  1   (1849).- 
REEVE,  Conch.  Icon.  V,  t.  78,  f.  574. — cf.  HIDALGO,  Journ.de  Con- 
chyl.  1887,  p.  149. 

This  form  is  retained  separate  from  C.  metaformis  oaly  because  of 
its  different  habitat.  Certain  forms  of  metaformis  correspond  closely 
with  the  figures  and  description  of  rustica,  such  as  the  shell  figured 
as  rustica  by  Hidalgo  (pi.  58,  fig.  89).  Of  this  last  form  specimens 
are  before  me  from  Mt.  Calanan.  Another  shell,  even  closer  to 
rustica,  is  figured  on  pi.  58,  fig.  90.  Figure  92  is  copied  from 
Mousson's  original  illustration. 

C.  HYDROPHANA  Sowerby.     PI.  36,  figs.  35,  36,  37. 

Imperforate,  oblong-ovate,  elevated,  thin.  Smooth  ;  straw-yellow, 
with  dark  chestnut  bands  at  suture,  periphery  and  columella,  and 
often  a  narrow  band  between  the  last  two.  Rarely  all  bands  are 
lacking.  Hydrophanous  cuticle  forming  a  thin  film  of  oblique 
shreds,  scarcely  visible  except  on  the  bands. 

Spire  elevated,  summit  obtuse.  Whorls  5,  the  last  slightly  de- 
flexed  in  front.  Aperture  oval,  white  and  showing  the  band  inside ; 
lip  narrowly  expanded  and  reflexed,  white  ;  columella  subvertical, 
very  deeply  inserted,  having  an  obtuse  lobe  below  but  not  toothed. 

Alt.  33,  diam.  25  mill. 

Alt.  25,  diam.  22J  mill. 

Mindoro,  Philippines. 

H.  hydrophana  Sow.,  P.  Z.  S.  1840,  p.  88.— PFR.  in  Conchyl. 
Cab.  t.  49,  f.  8,  9 ;  Monogr.  i,  p.  222.— REEVE,  Conch.  Icon.  f.  69.— 
(?.  hydrophana  HIDALGO,  Journ.  de  Conchyl.  1887,  p.  148. 

Very  closely  allied  to  C.  metaformis,  but  smaller,  thinner,  with  a 
less  thickened,  non-dentate  columella. 


188  <  OCHLOSTYLA-HELICOSTYLA. 

C.  BUTLERI  Pfeiffer.     PL  37,  fig.  50,  51. 

Imperforate,  globose,  rather  thin  ;  apex  obtuse,  white  ;  spire  semi- 
irlobose,  suture  moderate.  Whorls  4£,  nearly  flat,  the  last  inflated, 
pale  buff,  ornamented  with  close  brown  interrupted  lines.  Colu- 
mella  nearly  straight,  wide,  deeply  entering.  Aperture  rounded- 
lunar  ;  peristome  subthickened,  forming  an  obtuse  angle  with  the 
columella.  (P/V.)  Alt.  28,  diam.  37  mill. 

Mountain  districts  of  Benguet  and  Mancayan,  and  S.  Nicolas  in  N. 
Ecija,  Luzon. 

H.  butleri  PFR.,  P.  Z.  S.  1842,  p.  87  ;  Monogr.  i,  p.  254;  Con- 
chyl.  Cab,  p.  284,  t.  47,  f.  11,  12.— PHILIPPI,  Abbild.,  etc.  i,  p.  80, 
t.  4,  f.  12.— RVE.,  Conch.  Icon.,  f.  37. —  Cochlostyla  butleri  SEMP., 
Reis,  p.  193,  t.  13,  f.  2,  3,  6  ;  t.  18,  f.  13  (Anatomy.) 

Semper  regards  this  as  probably  a  variety  of  C.  metaformis. 

< '.  FULIGINATA  Martens.     Unfigured. 

Shell  conic-globose,  lightly  striatulate,  smoky-brown,  hydroph- 
unous,  having  two  pale  yellow  zones,  one  just  above,  the  other 
below  the  suture,  the  last  visible  upon  the  last  whorl  only,  and 
.-mall,  irregular,  shining  brown-black  spots ;  spire  conical,  apex 
obtuse ;  whorls  5,  joined  by  a  superficial  suture,  the  last  rounded, 
somewhat  descending  in  front.  Aperture  oblique,  square-rounded, 
palate  cserulescent,  parietal  wall  brown-black ;  peristome  narrowly 
reflexed,  thickened,  bluish-white,  outwardly  rubescent ;  columellar 
margin  rather  oblique,  nearly  straightened,  (v.  Mart.) 

Alt.  25,  greater  diam.  26 £,  lesser  23  mill. ;  aperture,  oblique  alt. 
18,  width  including  peristome  16  mill. 

Luzon,  Philippines. 

<  'ofJilostylafuliginata  MART,  in  Mai.  Bl.  xxi,  p.  46,  1873. — v. 
M«M,L.,  Nachrbl.  D.  M.  Ges.  1890. — Bulimus  fuliginatus  PFR., 
Monogr.  viii,  p.  S. —  C.  fumigata  "Mart."  SEMPER,  Reis.  Phil. 
Landmoll.,  p.  198. 

In  -vnrral  form  it  reminds  one  of  C.  balteata  and  orbitula  Sowb., 
Imt  tin-  columellar  margin  is  more  oblique  than  in  the  first,  and 
longer  than  in  the  latter  of  these  two ;  the  spire  is  more  pointed 
abovi-  than  in  either  of  the  species  named,  so  that  this  snail  stands 
l>"t \vt-n  these  and  Cochlostyla  (7)  viridix.  The  coloring,  however, 
i-  UK m-  that  of  the  group  Orthostylus,  being  similar,  for  instance,  to 
that  of  C.  rnfnijustra.  (Mart.) 


COCHLOSTYLA-HELICOSTYLA.  189 

Var.  NIGROLABIATA  Moll. 

Less  elevated,  much  thinner;  peristome  scarcely  thickened,  little 
expanded,  black-brown  ;  columella  white,  distinctly  toothed.  Alt. 
22*,  diam.  23  mill.  (Moll.) 

C.  LANGUIDA  Pfeiffer.     PL  36,  fig.  30. 

Imperforate,  globose,  rather  thin,  regularly  and  closely  obliquely 
striated.  Apex  obtuse,  buff;  suture  marked  by  a  shining  brown 
line.  Whorls  4J,  a  little  convex,  the  last  rufous,  covered  with  an 
ashen  hydrophanous  cuticle,  lineolate  with  white ;  base  nude, 
shining.  Columella  vertical,  white- calloused  ;  aperture  suboval ; 
peristome  ?  (P/V.)  Alt.  30,  diam.  30  mill. 

Siquijo r,  Philippin es. 

H.  languida  PFR.,  P.  Z.  S.  1842,  p.  150  ;  Mon.  i,  p.  254. — REEVP:, 
Conch.  Icon.  f.  68. 

Described  and  figured  from  two  decidedly  immature  examples. 

C.  UNICA  Pfeiffer.     PL  36,  fig.  33. 

Imperforate,  globose,  rather  solid,  obliquely  striatulate,  whitish ; 
suture  profound;  whorls  5-],  inflated,  the  upper  nearly  flat ;  colu- 
mella profoundly  entering,  arcuate,  subcanaliculate.  Aperture  very 
oblique,  subcircular,  white  within ;  peristome  widely  expanded, 
inwardly  reddish,  margins  approximating,  basal  thickened.  Alt. 
19,  diam.  38  mill.  (P/V.) 

Zamboanga,  Mindanao,  Philippines  (Cuming). 

H.  unica  PFR.,  P.  Z.  8. 1842,  p.  151  ;  Monogr.  i,  p.  253. — REEVE, 
Conch.  Icon.  f.  74. 

C.  LACERA  Pfeiffer.     PL  59,  figs.  1,  2. 

Imperforate,  ovate-globose,  solid,  scarcely  striated,  brown,  varied 
with  a  brown-ashen  ragged  cuticle.  Spire  semiglobose,  obtuse ; 
suture  hair-margined  with  pale,  not  at  all  impressed  above,  becom- 
ing profound.  Whorls  4-]  the  upper  scarcely  convex,  the  last  two 
rounded,  the  last  whorl  somewhat  ascending  in  front.  Aperture 
oblique,  subtetragonal-oval,  livid  inside;  peristome  narrowly  ex- 
panded and  reflexed,  columellar  margin  somewhat  straightened, 
flat,  dilated,  forming  an  angle  with  the  basal  margin.  Alt.  30, 
greater  diam.  33,  lesser  27i  mill.  (P/V-) 

Philippines. 

•  H.laceraPFR.,P.  Z.   S.   1853,   p.   126;    Monogr.   iv,  p.   206.— 
REEVE,  Conch.  Icon.  f.  1266. 


]  «)()  COCHLOSTYLA-HELICOSTYLA. 

('.  KKHI-.KINI  Pfeiffer.     PL  37,  figs.  45,  46. 

Imperforate,  subglobose,  solid,  very  delicately  striatulate;  white 
mid. T  a  thin  greenish-yellow,  lusterless  epidermis,  encircled  by  sev- 
eral wide  dark  chestnut  bands.  Spire  conoid-semiglobose,  rather 
obtuse.  Whorls  4],  moderately  convex,  rapidly  increasing,  the  last 
rounded,  shortly  descending  in  front,  blackish  around  the  slightly 
sloping  wide,  white  subexcavated  columella.  Aperture  oblique, 
lunate-round,  whitish  inside;  peristome  white,  expanded-reflexed, 
s..me\vhat  thickened  within.  Alt.  20,  greater  diam.  27,  lesser  23 

mill.     (P/>.) 

Philippines. 

H.  rehbeini  PFR.,  P.  Z.  S.  1852;  Monogr.  iii,  p.  189;  Conchyl. 
Cal».  p.  480,  t.  159,  f.  1,  2.— REEVE,  Couch.  Icon.  f.  663. 

C.  PAPUENSIS  Hedley.     PI.  58,  fig.  97. 

Shell  globosely  conical,  imperforate,  solid  ;  color,  the  hydropha- 
QOUS  epidermis  when  wet  or  oiled  is  of  a  rich  chocolate  color;  when 
dry,  a  pale  yellowish-brown  crossed  by  numerous  narrow  spiral 
brown  lines;  deprived  of  the  epidermis  the  shell  is  a  pale  yellowr, 
with  a  brown  subsutural  band,  first  two  whorls  purple-blue,  interior 
of  shell  lustrous  pale  blue,  peristome  brown;  whorls  5J,  convex, 
regularly  increasing,  last  f  of  total  length;  sculpture,  entire  shell 
finely  obliquely  striated  and  encircled  by  microscopic  close  regular 
raided  lines;  apex  obtuse,  embryonic  whorls  H>  ;  suture  impressed, 
margined;  aperture  oblique,  ovate  lunate,  peristome  slightly  re- 
flected, a  thin  semi-transparent  callus  extends  over  the  axis  and 
curves  up  to  the  insertion  of  the  right  margin.  Alt.  55,  breadth  38 
mill.  (Hedley). 

The  Madcay  Museum  possesses  several  examples  of  this  species 
pin-chased  from  Mr.  Goldie,  who  collected  them  in  British  New 
Guinea.  Considerable  difference  in  size  and  form  exists  between 
these  specimens,  none  of  which  are  in  a  good  state  of  preservation. 
<  )ne  old  and  worn  shell  exhibits  on  the  inner  side  of  the  columella 
a  tubercle  10  mm.  long  and  2  mm.  wide.  Whether  this  be  a  dis- 
tinct species,  the  adult  form,  or  merely  a  variety  of  the  species 
d«-s  -rilted  above,  requires  further  material  to  decide.  (Hedley.) 

British  New  Guinea. 

<  ''„•/, in. s///A/  fxtpuensis  HEDLEY,  The  Land  Molluscan  Fauna  of 
r.iitish  New  ( Juinea,  in  Proc.  Linn.  Soc.  N.  S.  Wales,  2d  ser.,  vi,  p. 
96,  t.  12,  f.  :j()  (March  25,  1891). 


COCHLOSTYLA-HELICOSTYLA.  191 

The  contour  of  this  species  throws  it  into  the  group  of  C.  meta- 

formis.     In  the  case  of  this,  as  well  as  the  other  Australasian  species 

of  Cochlostyla,  it  is  highly  desirable  that  the  genitalia  be  examined. 

(4).     Group  of  C.  fenestrata. 

The  shell  in  this  group  is  notably  compact  and  solid,  showing  close, 
fine  spiral  strise  under  a  lens  ;  and  hydrophanous  cuticle  is  present. 
The  mouth  is  rather  small,  the  outer  lip  decidedly  thickened  within  with 
white  callus;  the  columella  is  pure  white,  and  decidedly  thick,  often 
truncated  at  base. 

The  limits  of  this  group  are  doubtful,  v.  Mollendorff  including  in 
it  C.  bruguieriana,  crossei,  roissyana,  fuliginata,  montana,  curta, 
fenestrata,  sollda,  dimera,  jonasi.  A  careful  study  of  the  whole 
matter  has  inclined  me  to  believe  that  bruguieriana  cannot  well  be 
sundered  from  carbonaria  and  other  species  of  Axina,  and  I  am  not 
certain  that  solida  and  crossei  should  not  also  be  included  there. 
C.  roissyana  and  dimera  had  better  be  left  in  the  Cochlostylce  hypo- 
melanice.  Mollendorff's  citation  of  C.  jonasi  in  this  connection  is 
evidently  due  to  Pfeiffer's  mistaken  reference  in  the  Monographic, 
to  the  figures  of  the  plate  in  Kiister,  the  figures  of  dimera  being 
given  in  error.  On  the  other  hand,  I  feel  impelled  to  place  in  the 
present  group  a  number  of  shells  heretofore  classed  under  Ortho- 
stylus,  such  as  C.  leucophcea  and  C.  mus;  but  until  the  entire  group 
can  be  studied  in  the  light  of  a  greater  mass  of  material  we  will  not 
be  able  to  speak  definitely  upon  these  obscure  group  relations. 

C.  MONTANA  Semper.     PI.  58,  figs.  93,  94. 

Imperforate,  turbinate-globose  orturbinate,  solid,  chestnut  colored 
covered  with  a  streaked  hydrophanous  cuticle,  ornamented  with  two 
white  bands,  one  at  periphery,  the  other  at  the  suture,  and  close 
whitish  lines  beneath.  Spire  conoidal,  apex  rubesceut,  obtuse. 
Whorls  5£,  a  little  convex,  the  last  convex  and  denuded  on  the  base. 
Columella  excavated,  dentate,  white.  Aperture  lunate-oval,  milk- 
white  within ;  peristome  narrowly  thickened,  edged  with  black  in- 
side. Alt.  29,  greater  diam.  25,  lesser  21  mill.  (&) 

Foot  of  Monte  Data,  Mancayan,  Luzon. 

C.  montana  SEMP.,  Reis.,  p.  194,  t.  9,  f.  4. 


1«IL)  COCHLOSTYLA-HELICOSTYLA. 

C.  FKNKSTRATA  Sowerby.     PI.  58,  figs.  95,  96. 

Imperforate,  globose,  solid;  smooth;  ground-color  (under  the 
hpdrophunons  cuticle)  dark  chestnut,  the  earlier  whorls  whitish  ; 
mrerfd  iriili  a  very  thin  hoary  cuticle  with  oblique  whiter  streaks,  and 
having  a  cream-ivhite  band  just  above  and  one  below  the  periphery. 
There  is  a  series  of  angular  dark  chestnut  spots  a  short  distance  be- 
low the  suture,  and  the  base  is  more  or  less  denuded  in  front  of  the 

aperture. 

Spire  elevated  ;  whorls  5,  slightly  convex,  the  last  deflexed  in 
front.  Aperture  rounded-truncate,  bluish-white  inside;  lip  thick- 
ened within,  expanded,  white  with  a  brown  edge.  Columella  steeply 
sloping,  inconspicuously  2-lobed,  its  face  flattened  and  shallowly 
grooved.  Alt.  20,  diam.  18^  mill. 

Cagayan  and  N.  Ecija,  Luzon. 

H.  fenestrata  SOWB.,  P.  Z.  S.  1840,  p.  137.— REEVE,  Conch.  Syst. 
ii,  t,  166,  f.  24 ;  Conch.  Icon.,  f.  50.— PER.,  Monogr.  i,  p.  227  ;  Con- 
chy 1.  Cab.,  t.  108,  f.  9,  10. —  Cochlostyla  fenestrata  SEMPER,  Reis. 
Phil.  Archip.  Landmoll.,  p.  198. 

A  small  species,  readily  known  by  the  peculiar  color-pattern.  It 
is  more  allied  to  C.  montana  Semper  than  to  any  other  species. 

( '.  CURTA  Sowerby.     PI.  37,  figs.  47,  48. 

Imperforate,  ovate-conoid,  solid  and  thick.  Ground-color  yellow 
(or  black-brown),  covered  with  a  hydrophanous  cuticle  which  is 
creamy  above,  brown  on  the  base,  and  is  deciduous  in  patches,  show- 
iii'_r  ;i  series  of  spots  of  the  ground-color  on  the  upper  surface  of  the 
whorl.  Surface  microscopically  spirally  striated. 

Spire  conoidal,  apex  very  obtuse.  Whorls  6,  almost  flat,  narrow, 
the  last  whorl  slightly  descending  in  front.  Aperture  small,  oblique, 
white  inside;  peristome  narrowly  reflexed,  strongly  thickened 
within,  white  or  brown  edged.  Columella  short,  oblique,  \vhite, 
very  .-trongly  thickened,  more  or  less  truncated  below. 

Alt.  29,  diam,  23  mill. 

Balanan,  prov.  La  Union,  northern  Luzon,  Philippines. 

II.  curia  Sown.,  P.  /.  S.  1841,  p.  20.— PER.,  Monogr.  i,  p.  223; 
<  bnchyl,  (  ah.  p.  429,  t.  150,  f.  8,  9.— REEVE,  Conch.  Icon.  f.  34.— 
t  'ochlostyla  curto  HID.,  J.  de  C.,  1887,  p.  140. — Bid.  dilatatus  REEVE, 
Conch,  [con.  f.  ('.!). 


COOHLOSTYLA-PTYCHOSTYLUS.  193 

I  have  not  seen  the  typical  CURTA  (pi.  37,  figs.  47,  48),  which  has 

a  light  buff  ground-color,  the  cuticle  being  pale  above,  brown  below. 

The  shell  called  B.  dilatatus  bv  Reeve  has  the  ground-color  almost 

*•  o 

black,  becoming  rufous  or  yellow  above,  the  lip  dark-edged. 

The  variety  ft  of  Hidalgo,  as  well  as  the  specimens  before  me, 
one  of  which  is  shown  in  fig.  15,  are  the  same  as  the  form  called  B. 
dilatatus  by  Reeve.  While  I  am  wholly  disposed  to  consider  these 
shells  the  same  as  Pfeiffer's  dilatatus,  it  should  be  mentioned  that 
Hidalgo  has  reached  an  opposite  conclusion,  from  a  study  of 
Pfeiffer's  figures.  The  suite  before  me  shows  very  great  variations 
in  the  degree  of  development  of  the  columellar  tooth,  it  being  in 
some  examples  scarcely  noticeably  truncated. 

Var.     PI.  60,  fig.  16. 

Larger,  more  elevated,  very  solid ;  black-brown  below,  rufous 
above.  Whorls  6i.  Columellar  tooth  very  strongly  developed. 
Alt.  40,  diam.  28J  mill. 

The  specimen  is  denuded  of  cuticle. 

Var.  DILATATUS  Pfr.     PI.  60,  figs.  13,  14. 

Chestnut-brown,  tawny  above,  the  last  whorl  having  a  white  band 
below  the  suture. 

B.  dilatatus  PFR.,  P.  Z.  S.  1846,  p.  42  ;  Conchyl.  Cab.  p.  110,  t.  34, 

f.  7,  8 ;  Monogr.  ii,  p.  8. 

I  am  wholly  inclined  to  consider  this  a  mere  variety  of  C.  curta, 
founded  upon  a  specimen  denuded  of  hydrophanous  cuticle,  except 
for  a  band  remaining  below  the  suture  on  the  last  whorl.  In  case 
my  supposition  is  correct,  the  name  dilatatus  may  be  retained  in  a 
varietal  sense  for  all  specimens  of  C.  curta  having  a  dark  ground- 
color, including  Reeve's  dilatatus. 

Section  VIII.     PTYCHOSTYLUS  Mollendorff,  1888. 

Ptychostylus  MLLDFF.,  Nachrichtsbl.  D.  M.  Ges.,  1888,  p.  74.- 
Stylodonta  (in  part)  of  authors. —  Columplica  (in  part)  HARTMANN, 
Gasterop.  Schweiz,  p.  187,  188  (H.  uniplicata  the  first  species;  see 
under  Stylodonta  unidentata,  this  work,    Vol.   VI,    p.    86). — Not 
Ptychostylis  Gabb,  a  group  of  Trochidce. 

Shell  globose-turbinate,  formed  of  many  closely  coiled  whorls,  the 
surface  having  hydrophanous  cuticle;  aperture  narrowly  lunar; 
columella  spirally  twisted,  having  a  strong  tooth-like  fold  at  the  base. 

Animal  externally  like  Cochlostyla. 
13 


194  COCHLOSTYLA-HELICOBULINUS. 

The  single  species  contained  in  this  group  has  an  aspect  very 
different  from  the  usual  types  of  Cochlostyla.  There  are  certain 
forms  of  C.  grandis  Pfr.,  however,  which  present  an  approach  to  the 
peculiar  features  of  C.  cepoides,  especially  in  the  short,  twisted  col- 
umella  and  the  numerous  whorls. 

C.  CEPOIDES  Lea.     PL  48,  fig.  69. 

Imperforate,  globose-turbinate,  the  spire  short-conoid.  Ground- 
color brown  above,  yellowish  beneath ;  hydrophanous  cuticle  form- 
ing a  series  of  creamy  blotches  under  the  suture,  and  a  band  below 
periphery. 

There  are  6*  narrow,  closely  revolving,  convex  whorls.  The 
base  is  very  convex.  Aperture  narrowly  lunar,  oblique ;  lip  some- 
what expanded,  thickened  within,  brown-edged;  columella  vertical, 
very  short,  having  a  strong  tooth-like  fold.  Alt.  40,  diam.  44  mill. 

Is.  of  Luban,  Philippines. 

H.  cepoides  LEA,  Trans.  Amer.  Philos.  Soc.  (n.  ser.),  vii,  1841,  p. 
462,  1. 12,  f.  14.— SOWB.,  P.  Z.  S.  1840,  p.  88.— Rv.,  Conch.  Syst.  ii, 
1. 164,  f.  14 ;  Conch.  Icon.  f.  39.— PFR.,  Monogr.  i,  p.  302  ;  Conchyl. 
Cab.  t.  41,  f.  3,  4. — H.  (Stylodonta)  cepoides  PFR.,  Nomencl.  Hel. 
Viv.  p.  170. — H.  dolium  SOWB.,  ined. —  Columpliea  do  Hum  HART- 
MANX,  Gast.  Schweiz,  p.  188,  t.  67,  f.  1,  2. — Nanina  (Stylodonta) 
cepoides  TRYON,  Manual  of  Conchology  (2d  ser.)  ii,  p.  26,  t.  5,  f. 
86,  87. —  Cochlostyla  (Ptycho  stylus)  cepoides  MLLDFF.,  Nachr.  D.  M. 
Ges.  1888,  p.  74. 

This  peculiar  shell  has  long  been  classed  with  Stylodonta  uni- 
dentata  of  the  Seychelles  Is.,  but  there  is  now  no  doubt  that  Semper 
and  Mollendorff  are  right  in  considering  it  a  Cochlostyla. 

Section  IX.     HELICOBULINUS  Broderip,  1840. 

Helicobulinus  BROD.,  P.  Z.  S.  1840,  p.  123,  type  H.  sarcinosa. — 
Helicobulimus  MOLLENDORFF,  Landschn.  Cebu,  p.   241. —  Chromo- 
•cochlea  HARTMANN,  Gast.  Schweiz,  p.  137, 1844,  type  C.  turbinoides. 
-  Chromatocochlias  AGASSIZ. 

Si  icll  capacious,  turbinate-globose,  solid,  variegated  with  green  or 
brown,  covered  with  a  variously  patterned  hydrophanous  cuticle. 
<  'ohimolhi  more  or  less  folded. 

This  section,  while  closely  allied  to  Orthostylus,  presents  affinities 
with  BO  many  groups  that  it  must  be  regarded  as  an  intermediate  or 
synthetic  type.  Through  C.  grandis  it  approaches  the  section 


COCHLOSTYLA-HELICOBULINUS.  195 

Ptychostylus ;  through  certain  depressed  forms  of  C.  turgens  it  ap- 
proaches C.  harfordii  and  other  allied  species  of  Calocochlea ;  and 
on  the  other  hand,  it  is  difficult  to  so  diagnose  Helicobulinus  that 
all  species  of  Orthostylus  will  be  excluded. 

The  several  species  of  Helicobulinus  are,  however,  more  nearly 
related  to  each  other  than  the  various  forms  are  to  any  of  these 
diverse  groups,  so  that  despite  several  atttempts  by  various  authors 
to  dismember  the  section,  it  is  likely  to  maintain  its  integrity 
permanently. 

C.  GRANDIS  Pfeiffer.     PL  59,  figs.  7,  8  ;  pi.  60,  figs.  9,  10,   11  ;    pi. 
48,  figs.  66,  67,  68. 

Imperforate,  large,  trochoidal  or  turbinate,  solid,  dark  reddish- 
chestnut,  covered  with  a  cream  colored  hydrophanous  cuticle  which 
is  cut  by  dark,  narrow  spiral  bands,  and  is  largely  lost  through 
erosion  on  the  earlier  whorls  and  the  apertural  part  of  the  base. 

The  spire  is  elevated,  conical ;  apex  rather  obtuse,  whitish.  Whorls 
5J-6J,  the  last  obtusely  angular  at  periphery,  somewhat  concave  or 
flattened  below  the  angle.  Aperture  oblique,  white  or  blue  inside ; 
penstome  narrowly  reflexed,  purple-brown  ;  columella  very  short, 
deeply  entering,  and  having  a  strong  fold  near  the  insertion. 

Alt.  50,  diam.  60  mill. 

Alt.  43,  diam.  60  mill. 

Northern  Luzon,  at  Baler,  Palanan,  Digollorin,  Casiguran  and 
Camiguin  de  Luzon. 

H.  grandis  PFR.,  P.  Z.  S.  1845,  p.  43. — EEEVE,  Conch.  Icon.  t.  7, 
f.  30. —  Cochlostyla  grandis  DOHRN,  Mai.  Bl.  ix,  p.  211,  1862. — 
SEMPER,  Reis.  Landm.  p.  199. — H.  collosea  PFR.,  Zeitschr.  f.  Mai. 
1852,  p.  153  ;  Conchyl.  Cab.  p.  422,  t.  149,  f.  1,  2.— IT.  carolus  DH., 
Journ.  de  Conchyl.  1861,  p.  348,  t.  16,  f.  1,  2. —  Cochlostyla  carolus 
HIDALGO,  Journ.  de  Conchyl.  1887,  p.  138. 

This  magnificent  species  may  be  known  at  once  by  its  very  short 
columella.  It  varies  greatly  in  contour,  the  height  of  the  spire  and 
the  number  of  whorls  being  unusually  mutable.  Semper  is  disposed 
to  include  "  Bulimus  "  lignarius  Pfr.  with  grandis,  but  the  material 
before  me,  of  both  species,  causes  me  to  believe  them  entirely 
distinct. 

C.  SARCINOSA  Ferussac.     PI.  47,  fig.  58. 

Imperforate,  capacious,  globose-turbiuate.  Green,  becoming  fawn- 
tinted  on  the  spire,  encircled  at  the  periphery  by  a  chestnut  band, 


1 !  1 1  i  COCHLOST  YLA-HELCIOBULIXUS. 

which  ascends  the  spire  above  the  sutures.  Having  a  band  of 
hydrophanous  cuticle  below  the  peripheral  girdle,  usually  almost 
entirely  lost. 

« 

Spire  conoid,  apex  very  obtuse.  Whorls  5-J,  the  first  nearly 
planate,  the  last  slightly  deflexed  in  front.  Aperture  large,  blue- 
white  within  ;  lip  narrowly  expanded,  purplish-pink  ;  columella  of 
the  same  color,  subvertical,  nearly  straight,  faintly  obliquely  trun- 
cated below.  Alt.  72,  diam.  64  mill. 

Northern  Negros;  Masbate ;    Visayas ;  Batayan,  Zebu. 

H.  sarcinosa  FER.,  Hist.  t.  109,  f.  1,  2.— BROD.,  P.  Z.  S.  1840,  p. 
121. — REEVE,  Conch.  Icon.  f.  5b ;  Conch.  Syst.  ii,  t.  163,  f.  7. — 
PFR.,  Monogr.  i,  p.  244;  Conchyl.  Cab.  t.  40,  figs.  1,  2  (not  3,  4). 
-Bulimus  sarcinosus  PFR.,  Symbolse  ii,  p.  50. —  Cochlostyla  (Ortho- 
sti/lus')  sarcinosa  SEMP.,  Reis.  p.  201. —  C.  (Helicobulimus)  sarcinosa 
MLLDFF.,  Landschn.  Cebu,  p.  241. —  C.  sarcinosa  HIDALGO,  Journ. 
de  Conchyl.  1887,  p.  138. 

The  typical  C.  sarcinosa  is  a  large,  green  shell,  with  a  brown 
peripheral  band.  It  is  so  intimately  connected  with  the  following 
that  I  am  unable  to  say  whether  some  of  the  specimens  before  me 
should  be  considered  turgens  or  sarcinosa,  and  am  therefore  com- 
pelled to  unite  the  species,  as  Pfeiffer  at  one  time  did. 

Var.  TURGENS  Desh.     PI.  46,  figs.  50,  51,  52. 

Typically  smaller  than  sarcinosa,  the  prevailing  color  brown 
instead  of  green.  The  hydrophanous  cuticle  is  disposed  in  numer- 
ous revolving  bands,  interrupted  irregularly  by  oblique  brown 
streaks.  Peripheral  brown  band  either  present  or  wanting. 

Alt.  55,  diam.  52  mill. 

Alt.  70,  diam.  55  mill. 

Alt.  53,  diam.  53  mill. 

Panay,  Guimaras  and  Luzon,  Philippines. 

H.  turgens  Dn.  in  For.  Hist.  p.  316,  t.  108  C,  f.  11,  13.— H.  sar- 
t-lnosa  var.  PFR.,  Conchyl.  Cab.  t.  40,  f.  3,  4. —  Cochlostyla  turgens 
SEMP.,  Jlois.  p.  201.— HIDALGO,  Journ.  de  Conch.  1887,  p.  138. 

Kxcessively  variable  in  contour. 

C.  n  I:HINOIDES  Broderip.     PI.  47,  fig.  63  ;  pi.  62,  fig.  29. 

[mperforate,   globose-turbinate ;    ground-color   green,   becoming 
on  the  spire;  having  silvery  hydrophanous  bands. 


COCHLOSTYLA-HELCIOBULINUS.  197 

Spire  conic,  obtuse,  reddish.  Apex  plan  ate;  whorls  5},  convex, 
the  last  slightly  descending  in  front.  Aperture  large,  blue  within  ; 
peristome  expanded,  reddish-brown. 

Alt.  54,  diam.  45  mill. 

Alt.  50,  diam.  70  mill. 

Luzon,  Leyte,  Catanduanes  and  Camarines,  Philippines. 

H.  turbinoides  BROD.,  P.  Z.  S.  1840,  p.  23. — PFR.,  Mon.  i,  p. 
243  ;  Conchyl.  Cab.  p.  264,  t.  39,  f.  1,  2.— RYE.,  Conch.  Syst.  ii,  t. 
166,  f.  21,  22;  C.  Ic.,  f.  6. —  Cochlostyla  turbinoides  HIDALGO, 
Journ.  de  Conchyl.  1887,  p.  139. —  Chromocochlea  turbinoides 
HARTM.,  Gast.  Schweiz,  p.  137,  t.  41. 

Differs  from  the  preceding  in  its  red  upper  whorls,  darker  lip 
and  columella,  etc. 

C.  CINERASCENS  Pfeiffer.     PI.  47,  fig.  60. 

* 

Imperforate,  globose-turbinate,  rather  thin;  straw  colored  with 
two  wide  bands  and  a  blackish  basal  area,  covered  with  a  thin 
hydrophanous  cuticle  ;  spire  short-turbinate,  apex  obtuse.  Whorls 
5J,  a  little  convex,  the  last  inflated  beneath.  Columella  subarcuate, 
flesh-brown.  Aperture  round-lunar,  white  within ;  peristome  nar- 
rowly expanded,  somewhat  thickened,  chestnut-edged. 

Alt.  37,  greater  diam.  41,  lesser  35  mill.     (Pfr.*) 

Masbate,  Philippines. 

H.  cinerascens  PFR.,  P.  Z.  S.  1845,  p.  64;  Monogr.  i,  p.  220; 
Conchyl.  Cab.  p.  427,  t.  150,  f.  1,  2. — REEVE,  Conch.  Icon.  no.  78 
(description),  figure  81,  not  fig.  78! 

I  have  not  seen  the  typical  C.  cinerascens,  but  from  a  study  of 
the  figures  and  descriptions  I  am  wholly  inclined  to  consider  H. 
turbo  Pfr.  a  mere  variety  of  it.  Reeve  has  transposed  the  figures  of 
these  two  forms  on  his  plate. 

Var.  TURBO  Pfr.     PL  47,  fig.  59. 

Shell  turbinate,  solid  :  fleshy-brown  under  a  straw-colored  cuticle, 
having  a  peripheral  band  and  a  small  columellar  patch  of  chestnut. 
Spire  conical,  apex  obtuse  ;  whorls  5,  the  last  a  mere  trifle  descend- 
ing in  front.  Aperture  tinted  within  ;  lip  well  expanded,  brown  ; 
columella  steeply  sloping,  straight,  white.  Alt.  36,  diam.  36  mill. 

H.  turbo  PFR.,  P.  Z.  S.  1845,  p.  64;  Monogr.  i,  p.  220;  Conchyl. 
Cab.  p.  286,  t.  48,  f.  1,  2. — REEVE,  Conch.  Icon,  no.  81  (descrip- 


198  COCHLOSTYLA-ORTHOSTYLUS. 

tion),  fig.  78,  not  fig.  81 !— DESH.  in  Fer.  Hist.  t.  HOB,  f.  1.— (7. 
turbo  HIDALGO,  Journ.  de  Conch.  1887,  p.  139. 

Several  specimens  before  me  exhibit  no  variation.     The  published    ^ 
figures  cited  above  are  also  very  similar. 

C.  BEMBICODES  Pfeiffer.     PL  31,  figs.  36,  37. 

Imperforate,  globose-turbinate,  solid,  striate ;  white  under  a  dull 
buffish  lusterless  epidermis,  with  a  chestnut  band  at  the  periphery. 
Spire  turbinate,  apex  obtuse.  Whorls  5,  convex,  the  last  rounded, 
little  descending  in  front.  Aperture  very  oblique,  lunate-rounded, 
white  inside  and  shining.  Columella  subvertical,  straightened, 
dilated,  flat,  white  ;  peristome  white,  upper  margin  expanded,  basal 
narrowly  reflexed,  forming  an  obtuse  angle  with  the  columella. 

Alt.  35,  diam.  44  mill.     (P/r.) 

Romblon,  Philippines. 

H.  bembicodes  PFR.,  Zeitschr.  f.  Mai.  1851,  p.  15 ;  Monogr.  iii,  p. 
171 ;  Conchyl.  Cab.  p.  408,  t.  146,  f.  6,  7.— Cochlostyla  bembicodes 
HIDALGO,  Journ.  de  Conch.  1887,  p.  140. 

This  species  seems  near  to  C.  turbo  in  form  and  coloring. 

Section  X.     ORTHOSTYLUS  Beck,  1837. 

Orthostylus  BECK  (in  part),  Index,  p.  49. — MARTENS  in  Alb. 
Die  Hel.  p.  177. — v.  MLLDFF.,  Landschn.  Cebu,  p.  242. — Pithohelix 
SWAINS.,  Malacol.  p.  166. — Pythohelix  SWAINS.,  /.  c.,  p.  332. 

The  shell  is  generally  large,  solid  ovate-conic,  covered  with  a 
variously  patterned  hydrophanous  cuticle.  Aperture  oblique, 
ovate ;  lip  reflexed  ;  columella  nearly  vertical,  more  or  less  obviously 
folded  below. 

Anatomy  typical  of  Cochlostyla. 

The  present  section  is  generally  considered  the  typical  group 
of  Cochlostyla.  It  is  allied  to  the  section  Helicobulinus,  which 
consists  of  more  inflated  shells,  and  to  the  section  Hypselostyla,  con- 
taining more  elongated  forms.  The  limits  of  Orthostylus  are  not 
easy  to  determine,  as  there  are  species  almost  or  entirely  interme- 
diate between  this  group  and  Helicostyla,  as  well  as  forms  connect- 
ing with  Hypselostyla. 


COCHLOSTYLA-ORTHOSTYLUS.  199 

(1)    Typical  species  of  Orthostylus. 

C.  BICOLORATA  Lea.     PI.  44,  figs.  40,  41. 

Imperforate,  pyramidal,  solid  ;  ground-color  chestnut  above,  green 
below  the  periphery.  Covered  with  a  brown  hydrophanous  cuticle 
which  is  streaked  obliquely  with  lighter,  and  is  varied  by  a  creamy 
band  near  the  periphery,  a  series  of  alternating  light  and  denuded 
dark  blotches  below  the  suture  followed  by  another  band,  and  frequently 
a  third  band  on  the  base. 

The  spire  is  notably  pyramidal  and  elevated,  apex  rather  obtuse. 
Whorls  6*.  Aperture  small,  oblique,  white  inside  ;  peristome  dark 
chestnut  colored,  broadly  expanded  and  reflexed.  Columella 
vertical,  obliquely  truncated  beneath,  brownish. 

Alt.  60,  diam.  38  mill. 

Alt.  63,  diam.  42  mill. 

Lucban,  Prov.  Tayabas,  Luzon ;  Boac  and  Gasan,  Marinduque ; 
A  labat,  Philippines. 

Bui.  bicoloratus  LEA,  Proc.  Amer.  Philos.  Soc.,  p.  173,  1840; 
Trans.  Amer.  Philos.  Soc.  vii,  p.  459,  t.  11,  f.  8, 1841. — EVE.,  Conch. 
Icon.,  f.  22. —  Cochlostyla  bicolorata  MLLDFF.  Nachr.  D.  M.  Ges. 
1889,  p.  W4.—Bul.  alberti  BROD.,  P.  Z.  S.  1841,  p.  35.— PFR., 
Monogr.  ii,  p.  3  ;  Conchyl.  Cab.  Bui.,  t.  50,  f.  3. —  Cochlostyla  alberti 
SEMP.,  Reisen,  p.  202. — HIDALGO,  Journ.  de  Conch.  1887,  p.  163  ; 
Obras,  t.  65,  f.  2,  3. — Bulinus  onyx  BROD.,  P.  Z.  S.  1841,  p.  34.- 
Bnlimus  onyx  PFR.,  Monogr.  ii,  p.  32. —  Cochlostyla  alberti  var.onyx 
HIDALGO,  Obras,  Malacol.  i,  t.  65,  f.  4,  5. 

The  ground-color  is  peculiar,  being  green  below,  chestnut-brown 
above  the  periphery.  When  quite  denuded,  the  green  color,  being 
epidermal,  disappears,  white  replacing  it.  There  is  sometimes  a 
broad  chestnut  band  beneath,  conspicuously  modifying  the  green 
color,  and  leaving  green  bands  only  at  periphery  and  columella. 

Var.  ONYX  Brod.     PI.  61,  fig.  22. 

More  elongated  and  with  the  last  whorl  notably  narrower. 
Alt.  60,  diam.  27  mill. ;  apert.,  alt.  29,  diam.  (inside)  1 6  mill. 

Luzon. 

C.  IMPERATOR  Pfeiffer.     PI.  45,  figs.  42,  43,  44. 

Imperforate,  pyramidal,  solid.  Ground-color  brown-black  on  the 
body- whorl,  light  brown  on  the  two  preceding  whorls,  the  earlier 


200  COCHLOSTYLA-ORTHOSTYLUS. 

whorls  purple.      Hydrophanous  cuticle  forming  conspicuous  cream- 
white  and  rusty-brown  obliquely  longitudinal  ragged  stripes. 

Spire  decidedly  pyramidal,  elevated,  rather  slender.  Whorls 
nearly  7,  the  last  indistinctly  angular  at  the  periphery,  at  times 
somewhat  malleated,  the  indentations  large  and  shallow.  Aperture 
quite  oblique,  blue-white  inside ;  peristome  black-brown,  well  ex- 
panded. Columella  vertical. 

Alt.  71,  diam.  43  mill. 

Alt.  45,  diam.  29  mill. 

Is.  Catanduanes ;  Prov.  Tayabas,  Luzon. 

B.  imperator  PFR..  P.  Z.  S.  1848,  p.  110;  Monogr.  ii,  p.  533  ; 
Conchyl.  Cab.  Bui.,  p.  158,  t.  46.  f.  1,  2. — KEEVE,  Conch.  Icon.  f. 
183. —  Cochlostyla  imperator  HID.,  Journ.  de  Conch.  1887,  p.  163; 
Obras  Malacol.  i,  t.  65,  f.  1. 

A  beautiful  species,  boldly  painted  with  cream-white  longitudinal 
stripes.  From  the  province  of  Tayabas,  Luzon,  we  have  a  variety 
scarcely  half  the  size  of  typical  examples  (fig.  44.) 

C.  PITHOGASTER  Ferussac.     PI.  40,  figs.  2,  3,  4,  5. 

Imperforate,  oblong-conic  or  ovate-conic,  solid.  Ground-color 
chocolate-black  on  the  body-whorl,  chestnut  on  the  next  earlier,  yellow- 
ish brown  on  the  next,  the  earlier  2  or  2^  whorls  dark  blue-purple. 
Hydrophanous  cuticle  uniformly  distributed  or  slightly  streaked 
obliquely,  of  a  drab  or  olive-slate  color,  varying  from  dark  to  rather 
light. 

Spire  high-conoidal,  apex  very  obtuse.  Whorls  6-6i,  the  last  a 
trifle  deflexed  in  front.  Aperture  oblique,  white  inside  ;  peristome 
reflexed,  inwardly  flesh-pink,  edged  with  black-brown ;  columella  irreg- 
ularly arcuate,  flesh-pink,  generally  not  perceptibly  truncated  at 
base. 

Alt.  61,  diam.  40  mill,  (typical  form.) 

Alt.  57,  diam.  47  mill,  (obese  specimen.) 

Alt.  50,  diam.  33  mill,  (small  specimen.) 

Camarines,  Luzon;  Alabat ;  Masbate  and  Marinduque,  Philip- 
pines. 

H.  pithogaster  FER.,  Hist.  t.  108,  f .  3 ;    t.  110,  f.  1-3. — Bulimus 

l>'ithogaster  PFR.,  Monogr.  ii,  p.  4;  Conchyl.  Cab.  p.  169,  t.  51,  f.  1. 

-/*'"/.  pythogaster  REEVE,  Conch.  Icon.  Bui.  t.  1,  f.  26. —  Cochlo- 

styin  pitfiogn.tti',1  SI;MI-I;K,  Reisen,  p.  201,  t.  13,  f.  12  (anatomy). — 

C.  pithogaster  IIn>\i.<;o,  Journ.  de  Conch.  1887,  p.    163;    Obras 


COCHLOSTYLA-ORTHOSTYLUS.  201 

Malacol,  t.  67,  f.  1-4. — Bui.  philippinensis  PFR.,  P.  Z.  S.   1846,  p. 
42 ;   Conchyl.  Cab.  p.  164,  t.  50,  f.  1.— REEVE,  Conch.  Icon.  f.  5- 
DH.  in  Fer.  Hist.  t.  110A,  f.  10,   11. —  Cochlostyla  philippinensis 
SEMP.,  Reisen,  p.  202. —  C.  pithogaster  var.  philippinensis  HIDALGO, 
Obras,  t.  67,  f.  5. 

The  typical  C.  pithogaster  is  an  elongated  shell,  covered  with  a 
uniform  dark  cuticle  (fig.  5).  It  is  impossible,  however,  to  draw  a 
line  between  the  elongated  forms  and  the  short,  obese  forms,  such  as 
that  shown  in  fig.  3.  The  intergradation  is  complete.  It  is  equally 
impracticable  to  separate  as  a  distinct  species,  the  form  called 
philippinensis,  as  all  stages  from  a  uniform  to  a  streaked  cuticle  are 
present  in  the  suite  before  me. 

Color-var.  philippinensis  Pfr.     PL  61,  fig.  24. 

Hydrophanous  cuticle  distinctly  obliquely  streaked. 

Color-var.  bipartita  Pils.     PI.  61,  fig.  23. 

Ground-color  dark  brown  below,  lighter  brown  or  yellowish  above 
the  periphery,  earlier  whorls  whitish  ;  hydrophanous  cuticle  not 
especially  streaked. 

Alt.  67,  diam.  43  mill. 

Alt.  59,  diam.  38  mill. 

C.  VILLARI  Hidalgo.     PL  40,  fig.  1. 

Imperforate,  ovate-conoid,  solid,  little  shining,  striatulate.  Buff, 
under  a  whitish-ashen  cuticle,  which  is  scarcely  streaked;  above  paler, 
apex  flesh-colored. 

Spire  conoid,  apex  obtuse ;  suture  seen  under  a  lens  to  be  minutely 
denticulated.  Whorls  6,  convex,  the  last  large,  about  £  the  total 
length,  descending  in  front,  the  base  slightly  depressed.  Aperture 
rounded-subquadrate,  white  inside ;  peristome  reflexed,  white, 
angularly  joining  the  white,  rather  wide  and  scarcely  oblique  col- 
umella.  (Hid.} 

Alt.  69,  diam.  50  mill. 

Boac,  Gasan  and  Saban,  Marinduque. 

C.  villari  HID.,  Journ.  de  Conchyl.  1887,  p.  166,  t.  6,  f.  3. 

Var.  Last  whorl  chestnut,  the  rest  tawny,  apex  flesh-colored, 
columella  rose-brown,  peristome  purple-black.  (Hid.) 

C.  DAPHNIS  Broderip.     PL  42,  figs.  20,  21,  22,  23. 

Imperforate,  ovate-conic,  solid.  Ground-color  either  (1)  light 
olive-buff  becoming  lighter  toward  the  white  apex,  or  (2)  chestnut- 


202  COCHLOSTYLA-ORTHOSTYLUS. 

I 

brown,  darker  below,  becoming  yellowish  above,  the  apex  flesh- 
tinted,  or  (3)  base  dark  chestnut-brown,  upper  surface  light  buff- 
brown,  or  (4)  body-whorl  black-brown,  the  spire  either  pale  or  be- 
coming purple  towrard  the  apex.  Hydrophanous  cuticle  cream-white, 
rarely  almost  uniform,  generally  conspicuously  streaked  obliquely  or 
cut  by  spiral  dark  bands,  or  both  cut  and  streaked. 

The  spire  is  very,  distinctly  conoidal  and  more  acute  than  usual. 
Whorls  63,  the  last'  somewhat  descending.  Aperture  quite  oblique, 
white  or  blue-white  inside ;  peristome  well  expanded,  brown,  edged 
with  black-brown.  Columella  vertical,  simple  and  straight,  flesh- 
tinted  or  purple-pink. 

Alt.  50,  diam.  36  mill. 

Alt.  63,  diam.  46  mill. 
Islands  Bohol,  Siquijor,  Magtan  and  southern  Zebu,  Philippines.. 

H.  daphnis  BROD.,  P.  Z.  S.  1840,  p.  180. — DELESSERT,  Kec.,  t. 
39,  f.  13,  14. — Bulimus  daphnis  PFR.,  Monogr.,  ii,  p.  4 ;  Conchyl. 
Cab.,  t.  51,  f.  4. — RVE.,  Conch.  Icon.,  f.  8. — DH.  in  Fer.,  Hist.,  t. 
151,  f.  5, 6. —  Cochlostyla  daphnis  SEMPER,  Reisen,  p.  205. — HIDALGO,. 
Journ.  de  Conch.  1887,  p.  166. — v.  MOLL.,  Landschn.  Cebu,  p.  243. 

This  species  is  excessively  variable  in  the  ground-color  as  well  as 
in  the  pattern  of  cuticle.  The  latter,  however,  has  no  zones  of 
greater  intensity  or  whiteness,  such  as  C.  lignaria  shows  below  the 
periphery.  The  spire  is  notably  conoidal,  the  narrower  specimens 
of  typical  daphnis  approaching  in  this  respect,  C.  imperator,  a  species 
quite  different  in  the  form  of  the  columella. 

C.  CUNCTATOR  Reeve.     Vol.  VIII,  pi.  12,  figs.  1,  2. 

Imperforate,  ovate- conic,  solid.  Ground-color  brownish  olive- 
green,  becoming  yellowish  on  the  spire,  the  earlier  whorls  flesh-col- 
ored or  purple.  Hydrophanous  cuticle  cream-colored,  becoming 
brownish  below,  disposed  in  very  close,  narrow,  oblique  stripes.  A 
fewT  narrow,  inconspicuous  darker  spirals  are  visible. 

Spire  conoidal,  obtuse.  Whorls  6.  Aperture  blue  within ;  peri- 
stome narrowly  expanded,  edged  with  black-brown.  Columella 
purple-pink,  straight  above,  gently  arcuate  below ;  not  in  the  least 
truncated  below. 

Alt.  52,  diam.  38  mill,  (average  specimen). 

Alt.  49,  diam.  39  mill,  (short  specimen). 

Alt.  60,  diam.  38  mill,  (elongated  specimen). 

Zebu,  Philippines. 


COCHLOSTYLA-ORTHOSTYLUS.  203 

Bul.  cunctator  REEVE,  Conch.  Icon.  f.  554,  1849. 

This  form  is  closely  allied  to  C.  'daplinis,  differing  in  being 
slightly  less  conoidal,  and' in  the  pattern  of  cuticle.  It  is  likely 
that  intermediate  forms  will  be  found.  Reeve's  figure  was  drawn 
from  an  immature  example. 

a  FAUNUS  Broderip.     PI.  61,  figs.  17,  18,  19. 

Imperforate,  ovate-conoid,  solid.  Ground-color  varying  from 
dark  chestnut  to  light  yellow-brown,  the  darker  specimens  becoming 
lighter  above  the  periphery  and  on  the  spire  ;  apical  whorls  purplish. 
Covered  with  a  light  brown  hydrophanous  cuticle,  which  is  either 
almost  uniform,  or  obliquely  streaked  having  few  or  many  darker 
spiral  lines. 

Spire  conoid,  apex  decidedly  obtuse.  Whorls  6,  the  last  slightly 
descending.  Aperture  quite  oblique ;  peristome  narrowly  reflexed, 
having  a  black-brown  edge.  Columella  vertical,  nearly  straight, 
flesh-tinted. 

Alt.  51,  diam.  34  mill,  (average  example.) 

Alt.  43,  diam.  29  mill,  (small  example.) 

Alt.  54,  diam.  31  mill,  (narrow  example.) 

Zebu ;   Camotes  Is. ;  Masbate,  Philippines* 

B.faunus  BROD.,  P.  Z.  S.  1840,  p.  180. — PFR.,  Mouogr.  ii,  p.  5 ; 
Conchyl.  Cab.  t.  51,  f.  7. — REEVE,  Conch.  Icon.  f.  13. — HIDALGO, 
Journ.  de  Conch.  1867,  p.  167  ;  Obras,  t.  62,  f.  4,  5. —  Cochlostyla 
daphnis  var.faunus  SEMPER,  Reisen,  p.  205. — MLLDFF.,  Landschn. 
Cebu,  p.  243. 

Less  variegated  than  0.  daphnis,  narrower  in  form,  the  spire  not 
so  conical  as  that  of  C.  daphnis,  and  not  so  acute. 

Both  Semper  and  Mollendorff  believe  faunus  to  be  a  variety  of 
daphnis,  but  the  differences  of  contour  are  so  constant  throughout 
the  large  suites  of  each  before  me,  that  I  am  still  inclined  to  con- 
sider them  distinct  species.  There  is  a  common  form  of  C.  daphnis 
having  the  cuticle  uniform,  not  variegated.  A  specimen  of  this 
from  Zebu  is  shown  on  pi.  42,  fig.  21. 

C.  TICAONICA  Broderip.     PI.  62,  figs.  25,  26,  27,  28. 

Imperforate,  ovate,  solid.  Ground-color  dark  chestnut,  becoming 
lighter  on  the  spire,  the  earlier  whorls  light  brown  or  purple-brown. 
Hydrophanous  cuticle  white  above,  tinted  with  brown  below,  dis- 
posed in  wide  or  narrow  oblique  streaks,  uninterrupted  or  cut  by 
dark  spiral  lines. 


204  COCHLOSTYLA-ORTHOSTYLUS. 

The  spire  is  obtusely  conoid  ;  apex  obtuse.  Whorls  5J,  the  last 
somewhat  descending  in  front.  Aperture  quite  oblique,  bluish- 
white  inside ;  lip  narrowly  reflexed,  dark  brown  or  edged  with  dark 
brown  ;  columella  subvertical,  deeply  inserted,  flesh-tinted. 

Alt.  58,  diam.  42  mill. 

Alt.  44,  diam.  34  mill. 

Alt.  56,  diam.  38  mill. 

Islands  of  Ticao,  Masbate,  northern  Zebu,  northern  Negros,  Philip- 
pines. 

H.  (Cochlostyla)  ticaonica  BROD.,  P.  Z.  S.  1840,  p.  155. — Bui. 
ticaonicus  PFR.,  Monogr.  ii,  p.  6 ;  Conchyl.  Cab.  p.  175,  t.  52,  f.  1- 
3. — REEVE,  Conch.  Icon.  f.  14. —  Cochlostyla  ticaonica  SEMPER, 
Reisen,  p.  202. — HIDALGO,  J.  de  C.  1887,  p.  167. — MLLDFF.,  Land- 
schn.  Cebu,  p.  242. — Bui.  subglobosus  LEA,  Trans.  Arner.  Philos. 
Soc.  vii,  p.  455,  t.  11,  f.  1  (young  shell). 

This  species  is  allied  to  C.  daphnis,  but  it  is  shorter,  with  lower, 
much  more  obtuse  spire. 

The  ground-color  is  sometimes  dark  below,  decidedly  lighter 
above  the  periphery.  The  prevailing  form  upon  the  island  Zebu 
(figs.  26,  27)  is  small,  with  conspicuous  longitudinal  white  stripes, 
but  few  spiral  lines  or  none. 

Color-form  lutea.     PI.  62,  fig.  25. 

With  the  typically  colored  examples  of  ticaonica,  live  others  hav- 
ing a  light  buff  ground-color,  overlaid  with  cuticle  forming  oblique 
white  streaks  above,  generally  rusty-brown  beneath.  The  lip  is 
white,  its  edge  often  flesh-pink.  Columella  purple-pink. 

C.  LIGNARIA  Pfeiffer.     PI.  62,  fig.  30  ;  pi.  41,  figs.  10,  11,  12,  13. 

Imperforate,  turbinate-conic,  solid.  Ground-color  uniform  dark 
chestnut,  becoming  lighter  and  then  yellow-chestnut  above,  the  apex 
whitish.  Hydrophanous  cuticle  wood-brown  streaked  with  cream, 
having  a  cream-white  zone  under  the  periphery,  and  few  (fig.  10)  or 
many  (fig.  12)  dark  spiral  lines. 

Spiral  conoidal,  apex  obtuse.  Whorls  5*,  the  last  somewhat  de- 
scending in  front,  obtusely  angular  at  the  periphery  in  front,  the 
degree  of  angulation  varying  from  distinct  to  almost  imperceptible. 
Aperture  quite  oblique,  nearly  white  inside;  peristome  expanded 


COCHLOSTYLA-ORTHOSTYLUS.  205 

and  reflexed,    dark    brown.      Columella   oblique,    indistinctly   or 
obviously  folded  below. 

Alt.  64,  diam.  58  mill,  (large  typical  form.) 

Alt.  51,  diam.  882  mill.  (var.  minor.) 

Alt.  60,  diam.  41*  mill,  (elongated  form.) 

Prov.  Cagayan,  Luzon. 

Bulimus  lignarius  PFR.,  P.  Z.  S.  1842,  p.  88  ;  Monogr.  ii,  p.  3  ; 
Conchy].  Cab.  Bui.  t.  19,  f.  1,  2. — KEEVE,  Conch.  Icon.  Bui.  t.  1,  f. 
3. — DH.  in  Fer.  Hist.  t.  151,  f.  8,  9. —  Coclilostyla  lignaria  SEMPER, 
Reisen,  p.  202. — HIDALGO,  Journ.  de  Conch.  1887,  p.  160. 

The  creamy  subperipheral  zone  is  a  constant  and  leliable  feature. 
The  peripheral  angulation  is  scarcely  noticeable  in  the  smaller  speci- 
mens, such  as  those  figured  on  pi.  41,  and  which  are  much  more 
numerous  in  collections  than  the  larger  form.  These  small  speci- 
mens are  about  the  size  of  C.  daphnis,  but  that  species  is  more  dis- 
tinctly conical,  more  acute  than  this,  and  it  lacks  the  subperipheral 
creamy  zone. 

Semper  has  stated  that  it  is  difficult  to  distinguish  the  more 
obviously  angular  forms  of  C.  lignaria  from  C.  grandis  Pfr.,  and  he 
has  therefore  united  the  two  ;  but  with  all  respect  for  the  opinion 
of  the  distinguished  German  zoologist,  we  must  still  consider  these 
species  perfectly  distinct.  It  is  hard  to  even  imagine  a  connection. 

C.  GILVA  Broderip.     PL  42,  figs.  24,  25,  26. 

Imperforate,  globose-ovate,  solid.  Ground-color  dark  chestnut, 
begirt  with  an  indistinct  darker  peripheral  band,  and  becoming 
lighter  above,  the  earlier  whorls  whitish  with  a  brown  subsutural 
margin.  Hydrophanous  cuticle  forming  a  uniform  cream-white  or 
light  brown  coat. 

Spire  bluntly  conoid,  the  apex  very  obtuse.  Whorls  5J,  the  last 
generally  deeply  descending  in  front.  Aperture  large,  blue-white 
witfiin  ;  lip  rather  broadly  reflexed,  white,  sometimes  brown-edged. 
Columella  nearly  vertical,  wide,  white. 

Alt.  45,  diam.  40  mill. 

Alt.  42,  diam.  37  mill. 

Philippines. 

B.  gilvus  SOWERBY,  (in  Coll.  Cuming) ;  PFR.,  P.  Z.  S.  1845,  p. 
45;  Conchyl.  Cab.,  p.  179,  t.  53,  f.  1. — REEVE,  Conch.  Icon.,  f.  7. 

Allied  to  C.  ticaonica,  but  more  globose,  with  shorter  spire,  more 

uniformly  distributed  cuticle,  whiter,  wider  lip,  and  having  an  in- 
distinct darker  peripheral  band. 


206  COCHLOSTYLA-ORTHOSTYLUS. 

C.  WOODIANA  Lea.     PI.  43,  figs.  28,  29. 

Imperforate,  oblong-conic,  solid.  Ground-color  dark  chestnut, 
becoming  light  yellowish-brown  above,  and  black  on  the  latter  part 
of  the  body-whorl ;  covered  with  cream-colored  hydrophanous  cuticle* 
which  is  cut  into  bands  by  a  number  of  dark,  denuded  broad  zones 
and  narrow  lines. 

Spire  conical,  apex  obtuse,  yellow-brown.  Whorls  5J,  the  last 
whorl  slowly  descending,  equal  to  the  spire  in  height.  Aperture 
oblique,  blue-white  inside ;  peristome  broadly,  roundly  reflexed, 
brown  or  fleshy-violet;  columella  vertical,  gently  arcuate,  flesh- 
colored,  forming  a  slightly  projecting  angle  where  it  joins  the  basal 
lip.  Alt.  70,  diam.  50  mill. 

Provinces  Tayabas  and  Laguna,  Luzon,  Philippines. 

Bulimus  woodianus  LEA,  Proc.  Amer.  Philos.  Soc.  i,  p.  173  (read 
Feb.  21,  1840;  Proc.  for  Jan.  and  Feb.,  1840,  distributed  March 
20,  1840)  ;  Trans.  Amer.  Philos.  Soc.  vii,  p.  457,  t.  11,  f.  5.— NOT 
B.  woodianus  PFR.  et  al. — H.  Reevii  BROD.,  P.  Z.  S.  1841,  p.  34.- 
Bulimus  Reevii  Rv.,  Conch.  Syst.  ii,  t.  172,  f.  1 ;  Conch.  Icon.,  f.  6. 
— B.  reevei  PFR.,  Monogr.  iii,  p.  295  ;  Conchyl.  Cab.,  Bui.,  t.  19,  f. 
5,  6. — DESH.  in  Fer.,  Hist.,  1. 116,  f.  1, 2. —  Cochlostyla  reevei  SEMPER, 
Reisen,  p.  203. — HIDALGO,  Journ.  de  Conchyl.  1887,  p.  160. —  C. 
woodiana  v.  MOLL.,  Nachr.  D.  M.  Ges.  1889,  p.  104. 

This  is  a  magnificent  species,  allied  to  C.  portei,  but  differing  in 
coloring.  It  is  not  especially  related  to  any  other  species. 

Lea's  heretofore  unquoted  description  in  the  Proc.  Amer.  Philos. 
Soc.  has  indisputable  priority  of  publication  over  Broderip's  descrip- 
tion in  P.  Z.  S.  under  the  name  Reevii.  The  change  of  name  is, 
therefore,  inevitable.  It  is  surprising  that  so  careful  an  author  as 
Pfeiffer  should  confound  Lea's  dark  brown  shell  with  the  totally 
diverse  white  species  which  has  always  been  called  woodianus. 

I  have  examined  Lea's  type  of  woodiana.  It  is  denuded 
of  the  creamy  hydrophanous  cuticle,  except  just  behind  the  ex- 
panded lip,  where  it  shows  the  characteristic  banding  of  the  species. 

C.  PORTEI  Pfeiffer.     PI.  44,  figs.  38,  39. 

Imperforate,  ovate-conic,  solid  ;  ground-color  green,  becoming  buff 
on  the  earlier  whorls  ;  covered  with  a  hydrophanous  cuticle  of  white, 
obliquely  streaked  with  tawny,  which  is  cut  by  a  couple  of  peripheral 
denuded  zones  and  a  number  of  lines. 


COCHLOSTYLA-ORTHOSTYLUS.  207 

Spire  conoid,  apex  obtuse ;  whorls  5*,  the  last  scarcely  descending, 
more  than  half  the  total  height  of  the  shell.  Aperture  oblique, 
large,  blue-white  inside ;  peristome  broadly  and  roundly  reflexed, 
dark  chestnut-brown.  Columella  vertical,  slightly  arcuate,  slightly 
salient  at  its  junction  with  the  basal  lip. 

Alt.  70,  diam.  54  mill. 

Alt.  60,  diam.  45  mill. 

Island  of  Polillo ;  District  of  Infanta,  Prov.  Laguna,  Luzon, 
Philippines. 

B.  portii  PFR.,  P.  Z.  S.  1861,  p.  193  (typog.  err.  for  portei)  ;  B. 
portei  PFR.  Novit.  Conch.,  p.  308,  t.  75,  f.  1-3  ;  Monogr.  vi,  p.  5. 

The  ground-color  is  green,  and  the  brown-lipped  aperture  is  larger 
than  in  C.  woodiana  Lea  (reevei  Brod.),  a  species  similar  in  pattern 
of  cuticle. 

C.  RUFOGASTER  Lesson.     PI.  43,  figs.  26,  27  ;  pi.  45,  fig.  47. 

Imperforate,  ovate-conoid,  solid.  Ground-color  chocolate  or  dark 
chestnut,  with  a  buff  band  at  the  periphery.  Hydrophanous  cuticle 
dull  brown,  varied  by  a  series  of  creamy,  oblique  dashes  below  the 
suture,  alternating  with  blotches  of  the  dark  ground-color  of  equal 
size,  and  having  a  light  creamy  zone  at  the  periphery,  and  another  at 
the  columella. 

Spire  high-conic,  apex  obtuse.  Whorls  6-6  J,  the  last  very  indis- 
tinctly angled  at  the  periphery  in  front.  Aperture  oblique,  white 
or  lilac-white  inside  ;  peristome  rather  narrowly,  roundly  reflexed, 
chocolate  colored ;  columella  subvertical,  somewhat  arcuate,  flesh- 
white  or  brown,  forming  a  rather  salient  angle  where  it  joins  the 
basal  lip. 

Alt.  76,  diam.  54  mill,  (typical.) 

Alt.  54,  diam.  43  mill. 

Alt.  45,  diam.  35  mill.  (v.  minor.) 

Tuan,  Prov.  of  Bulacan;  Porac,  prov.  Pampanga;  Los  Banos, 
Laguna ;  S.  Nicolas,  N.  Ecija,  and  Baler,  on  the  E.  coast,  Luzon. 

H.  rufogaster  LESS.,  Illustr.  de  Zool.,  t.  22,  1831. — Bulimus 
rufogaster  PFR.,  Monogr.  iii,  p.  6  ;  Conchyl.  Cab.,  t.  50,  f.  2. — REEVE, 
Conch.  Icon.,  f.  4.— DH.  in  Fe*r.,  Hist.,  t.  116,  f.  3,  6.—  Cochlostyla 
rufogaster  SEMP.,  Reisen,  p.  202. — HIDALGO,  Journ.  de  Conchyl. 
1887,  p.  160— Bui.  monozonus  PFR.,  P.  Z.  S.  1848,  p.  110 ;  Monogr. 
ii,  p.  533  ;  Conchyl.  Cab.,  t.  46,  f.  7,  8.— REEVE,  Conch.  Icon.,  f. 
195. 


208  COCHLOSTYLA-ORTHOSTYLUS. 

This  large  species  has  the  contour  of  C.  woodiana  Lea  (not  Pfr.),, 
but  is  differently  colored  and  the  lip  is  narrower.  It  is  even  more 
allied,  probably,  to  the  narrower,  more  pyramidal  C.  bicolorata  Lea. 

Pfeiffer's  B.  monozonus  (pi.  45,  fig.  47)  is  merely  a  rather  small 
example,  but  I  have  seen  specimens  still  smaller.  Localities  for 
these  small  specimens  are:  Santa  Ana,  prov.  Manilla;  Lucban 
prov.  Tayabas ;  San  Rafael,  prov.  Bulacan,  Luzon. 

C.  MACROSTOMA  Pieiffer.     PL  61,  fig.  20. 

Imperforate,  ovate,  rather  solid.  Rufous-black,  the  apex  paler  ; 
covered  with  a  brown  hydrophanous  cuticle,  encircled  by  black 
lines.  Whorls  6,  a  little  convex,  the  last  about  equal  to  the  spire 
in  length.  Columella  vertical,  dilated,  somewhat  thickened  in  the 
middle,  its  inner  part  dark  brown,  outer  lip  strongly  arcuate. 

Alt.  61,  diam.  39  mill.     (Pfr.) 

Mts.  near  Balanga,  prov.  Bataan,  Luzon,  Philippines* 

B.  macrostomus  PFR.,  P.  Z.  S.  1842,  p.  152  ;  Monogr.  ii,  p.  24  ; 
Conchyl.  Cab.,  t.  50,  f.  4,  5. — REEVE,  Conch.  Icon.,  f.  3. —  Cochlo- 
styla  macrostoma  HID.,  Journ.  de  Conch.  1887,  p.  161. 

C.  VIDALI  Hidalgo.     PI.  60,  fig.  12. 

Imperforate,  ovate,  solid,  obliquely  striated.  Under  a  thin, 
deciduous  hydrophanous  cuticle,  of  a  deep  purple-brown  color. 
Spire  conoid,  apex  obtuse,  concolored,  purple ;  suture  impressed, 
destitute  of  spots.  Whorls  6,  nearly  flat,  the  last  about  equaling 
the  spire,  encircled  with  a  little-conspicuous  pale  buff  band ;  base 
attenuated.  Aperture  augulate-ovate,  moderate,  inside  livid-white ; 
peristome  narrowly  reflexed,  edged  with  dark  purple,  margins  dis- 
tant, joined  by  a  thin,  wide  callus  ;  columellar  margin  deeply  intorted, 
quasi-truncate,  purple-white ;  basal  margin  narrowly  rounded,  form- 
ing an  angle  where  it  joins  the  columella,  outer  margin  becoming 
thin  toward  its  insertion. 

Alt.  67,  diam.  40  mill. ;  apert.  with  perist.,  37  mill,  long,  26 
wide.  (Hid.') 

Galiano,  district  of  Benguet,  Luzon. 

C.  vidali  HID.,  Journ.  de  Couch.  1887,  p.  162,  t.  7,  f.  2. 

This  species  is  truly  intermediate  between  C.  rufogaster  and  C. 
macrostoma.  It  differs  in  being  more  ovoid,  less  conoidal;  the  whorls 
more  swollen  ;  the  spire  is  more  obtuse  and  darker  than  the  rest  of 
the  shell,  not  lighter,  as  it  is  in  the  two  allied  species.  The  cuticle 


COCHLOSTYLA-ORTHOSTYLUS.  209 

lias  no  blotches  below  the  suture  ;  it  is  thinner  than  in  macro stoma, 
and  the  hydrophanous  band  is  not  accompanied  by  a  dark  line,  etc. 

C.  JUGLANS  Pfeiffer.     PL  59,  fig.  6. 

Imperforate,  elongate-globose,  the  apex  obtuse ;  rather  solid, 
obliquely  distinctly  striated,  unicolored  rufous.  Whorls  5,  convex, 
the  last  about  equal  to  the  spire.  Columella  wide,  white,  arcu- 
ately  curved  inward.  Aperture  large,  irregularly  suboval,  sinuated 
at  base  of  the  columella,  pale  livid  inside ;  peristome  scarcely 
reflexed,  margined  with  rufous. 

Alt.  53,  diam.  35  mill.     Aperture  28  mill,  long  inside.     (Pfr.) 

Mountains  of  northern-central  Luzon. 

Bul.juglans  PFR.  P.  Z.  S.  1842,  p.  89  ;  Monogr.  ii,  p.  7  ;  Conchy  1. 
Cab.,  p.  177,  t.  52,  f.  7. — REEVE,  Conch.  Icon.,  f.  12. —  Cochlostyla 
juglans  SEMPER,  Reisen,  p.  204. — MLLDFF,  Nachr.  D.  M.  Ges.  1888, 
p.  105. 

Fresh,  un rubbed  shells,  says  Mollendorff,  are  covered  with  a  thin 
gray-brown  cuticle,  which  often  shows  close  spiral  lines.  The  per- 
istome is  lipped  with  beautiful  purple-brown  ;  the  columella  is  white 
above. 

Var.  olivacea  Mlldff.     Smaller,  more  ventricose,  whorls  more  con- 
vex ;  olive-brown  or  olive-yellowish,  the  peristome  white.     (JMlldff^) 
Alt.  55,  diam.  43  mill. 
Alt.  50,  diam.  41  mill. 

Var.  roseolimbata  Mlldff.  More  ventricose,  olive-yellowish,  varied 
with  irregular  streaks;  peristome  more  thickened,  margin  pale 
purple  or  rose,  columella  more  spirally  twisted,  arcuate.  (J///o?jf.) 

Alt.  63,  diam.  49  mill. 

Alt.  59,  diam.  46  mill. 

Cuyapo,  prov.  Nueva  Ecija,  Luzon. 

C.  MARINDUQUENSIS  Hidalgo.     PL  45,  figs.  48,  49. 

Imperforate,  ovate-conic,  solid.  Under  an  evanescent  deciduous 
cuticle  the  surface  is  white,  unicolored  or  with  peripheral  and  colu- 
mellar  dark  bands,  or  having  the  middle  and  base  of  the  body- 
whorl  dark  brown.  Sometimes  there  is  a  narrow  sutural  band  ;  lip 
lilac-pink,  black  edged,  rarely  pure  white. 

Spire  conoidal ;  whorls  6,  convex,  especially  the  latter  two.  Last 
whorl  generally  obliquely  malleated  around  the  middle.  Aperture 
oblique,  white  inside ;  peristome  broadly  expanded  and  reflexed,  lilac 
14 


210  COCHLOSTYLA-ORTHOSTYLUS. 

pink  with   a  black-brown  edge ;  columella  very  deeply   inserted, 
lilac-pink,  expanding  in  a  blackish  callus. 

Alt,  64,  diara.  46  mill. 

Alt.  57,  diani.  42  mill. 

Boac,  Gasan  and  Saban,  Marinduque;  Romblon,  Philippines. 

C.  marinduquensis  HIDALGO,  Journ.  de  Conch.  1887,  p.  157. — 
Bulimus  ivoodianus  PFR.,  Conchyl.  Cab.,  p.  171,  t.  51,  f.  5. — REEVE, 
Conch.  Icon.,  f.  11.— DH.  in  Fer.,  Hist.,  t.  110A,  f.  7-9.— NOT 
Bulimus  ivoodianus  LEA,=  C.  reevei  Brod. 

The  peculiar  coloration  of  this  species  readily  distinguishes  it 
from  all  others,  although  it  is  quite  variable.  Some  specimens  are 
uniform  white ;  others  have  the  median  and  basal  portions  of  the 
body-whorl  blackish-brown  ;  but  the  most  frequent  form  is  that  re- 
presented by  fig.  48. 

This  species  has  long  been  known  under  the  name  "  C.  woodiana 
Lea,"  but  Lea's  species  is  a  totally  different  shell. 


INDEX  to  Subgenera  and  Sections 
contained  in  Vol.  VII, 


NOTE. — Aii  index  to  the  species  of  the  genus  Helix  will  accom- 
pany the  next  volume  of  the  MANUAL. 


Acavus  Srn.=Papuina, 
Albersia  H.  Ad., 
Axina  Alb., 

Callicochlias  Ag.  =  Caloco- 

chlea, 

Calocochlea  Hartm.,   .  129, 
Carocolla  Brod.— Axina 
Chlorsea  Alb., 
Chromatocochlias    Agas.= 

Helicobulinus  Brod., 
Chromatosphrera  Pils., 
Chromocochlea     Hartm.  = 

Helicobulinus  Brod., 
Cochlodryas  Mts.,        .  172, 
Cochlostyla  Fer., 
Coliolus  T.-C.,     . 
Columplica     Hartm.=Pty- 

chostylus, 
Corasia  Alb., 
Crystallopsis    Anc.=C  o  r  - 

asia, 
Cymotropis  Mts.=Papuina, 

Eugenia  Alb.— Papuina, 

Geotrochus  Beck=Papuina, 
Geotrochus     Hasselt=Tro- 
chomorpha,     .         0 


.      3 
.    89 
30,  158 

co- 
.  129 
29,  130 
.  158 
.    93 

Helicobulimus  Mlldff.—  He- 
licobulinus  Brod.,    .         .194 
Helicobulinus  Brod.,  .         .194 
Helicostyla  Fer.,    171,172,181 

Insularia  T.-C.—  Papuina,  .  3 
Ley  tia  Pils.,  .  .  .129 
JNIerope  Alb.—  Papuina,  .  3 

Orthostylus  Beck,  .  .  198 
Orustia  Morch.,  .  .  172,  176 

Paehysphsera  Pils.,      .         .172 
Papuina  Mts.,     ...       3 
Philidora  de  Morg.,     .         .     86 
Pfeifferia  Gray,  .         .         .127 
Pileolus  Less.^Papuina,     .       3 
Pithohelix  Swains.=Ortho- 
stylus  Beck,     .         .         .198 
Ptychostylus  Moll.,      .         .  193 
T^vtlioliplix"  ftwRina  —  Ortlio- 

.  194 

.  169 

.  194 

72,  177 
.     92 

.     87 

'ty- 

!  104 
>r- 
04,  105 
na,       3 

.       3 

na,       3 
'ro- 
.      3 

J.  V  Li-J.  \J  11  VxA  1  J.V  l-J  »  ClillJO*"  \^f  d.  Ul±\J 

stylus  Beck,     .         .         .198 

Satsuma,      .         .         .         .81 
Stylodonta   A  uct.—  Ptycho- 
stylus,     .         .         .         .193 

Trachystyla  Pils.,  .  130,  166 

(211) 


9  0  ,-i 


EXPLANATION  OF  PLATES. 

Vol.  VII. 


PLATE  1. 

FIGURE.  PAGE. 

1.  Papuina  zeno  Braz.  (latiaxis  Sm.).     Ann.  Mag.,       .  .     53 

2-4.  Papuina  zeno  Braz.     Specimens,           .         .         .  .53 

5.  Papuina  albocarinata  Sm.     Ann.  Mag.,    .         .         .  .59 

6.  Papuina  brumeriensis  Fbs.     Voy.  Rattlesn.,    .         .  .52 

7.  8.  Papuina  brumeriensis  Fbs.     Specimen,         .         .  .52 
9,  10.  Papuina  diomedes  Braz.     Specimens,          .         .  .54 

11.  Papuina  lacteolata  Sm.     Ann.  Mag.,        .         .         .  .25 

12.  Papuina  rollsiana  Sm.     Ann.  Mag.,          .         .         .  .63 

13.  Papuina  chapmani  Cox.  (coraliolabris  Sm.).     Ann.  Mag.,     51 

14.  15.  Papuina  strabo  var.  katauensis  T.-C.     Mus.  Geuov.,  .     60 

16.  Papuina  gurgustii  Cox.     P.  Z.  S.,     .         .         .         .  .     61 

PLATE  2. 

17,  18.  Papuina  strabo  var.  roseolabiata  Sm.     Ann.  Mag.,  .     60 
19.  Papuina  strabo  var.  (purple  lip).     Specimen,  .         .  .60 
20,21.  Papuiua  tayloriana  A.  <fe  R.     Ann.  Mag.,  .         .  .58 
22,  23.  Papuina  strabo  Braz.  typical.     Specimen,            .  .     60 
24-26.  Papuina  strabo  var.  katauensis  T.-C.     Specimen,  .     60 

27.  Papuina  tayloriana  A.  &  R.     Specimen,           .         .  .58 

28.  Papuina  tayloriana  var.  yulensis  Braz.     Mus.  Genov.,  .     59 
29-31.  Papuina  tayloriana  var.  yulensis  Braz.     Specimen,  .     59 
32-35.  Papuina  uaso  v.  Mart.     Specimens,    .         .         .  .56 

PLATE  3. 

36,37.  Papuina  coxiana  Aug.     P.  Z.  S.,         .         .         .  .13 

38,  39.  Papuina  macgillivrayi  Fbs.     Kiister,          .         .  .55 

40,  41.  Papuina  redempta  Cox.     P.  Z.  S.,       .         .         .  .     70 

42,  43.  Papuina  woodlarkiana  Sow.     J.  de.  C.,       .         .  .     62 

44-47.  Papuina  gamelia  Ang.     P.  Z.  S.  and  Specimen,  .     10 

48-50,  58.  Papuina  exsultans  Tap.-Can.     Novit,  Conch.,  .     31 

51.  Papuina  ferrussaci  Less.     Yoy.  Coq.,        .         .         .  .30 

52-53.  Papuina  bidwilli  Pfr.     Specimen,     , .         .         .  .55 

54.  Papuina  fergusoni  Ad.     P.  Z.  S.,               .         .         .  .     32 

55.  Satsuma  rufofilosa  Bock.     P.  Z.  S.,           .         .         .  .     84 

56.  Papuina  canovarii  T.-C.     Mus.  Genov.,    .         .         .  .26 

(212) 


EXPLANATION    OF   PLATES.  213 

FIGURE.  PAGE. 

57.  Papuina  pythonissa  T.-C.     Mus.  Genov.,  .         .         .31 

59,  60.  Papuina  blainvillei  Le  Guill.     Mus.  Genov.,       .         .     25 

PLATE  4. 

61-63.  Papuina  louisiadensis  Fbs.     Specimen,       .  .         .61 

64,  65.  Papuina  louisiadensis  var.  millicentse  Cox.  P.  Z.  S.,     62 

66,  67.  Papuina  philomela  Aug.       .     P.  Z.  S.,       .  .         .       8 

68,  69.  Papuina  flexilabris  Pfr.     Ku'ster,       .         .  .         .49 

70,  71.  Papuina  flexilabris  Pfr.     Specimens,           .  .         .49 

72-74.  Papuina  hargravesi  Cox.         .     Specimens,  .         .       9 
75-77.  Papuina  ambrosia  Aug.     Specimens,           ...       7 

78,  79.  Papuina  coniformis  Fer.     Ku'ster,       ,         .  .         .50 

80,  81.  Papuina  guadalcanarensis  Cox.     Specimens,  .         .       9 

PLATE  5. 

82,  83.  Papuina  ridibunda  T.-C.     Mus.  Genov.,     .         .         .40 

84.  Papuina  meditata  T.-C.     Mus.  Genov.,     .         .         .         .40 

85,  86.  Papuina  taumantias  T.-C.     Mus.  Genov.,    .         .         .39 

87.  Papuina  donnaisabella  Ang.     P.  Z.  S.,  .         .         .71 

88.  Papuina  dampieri  Ang.     P.  Z.  S.,    .         .         .         .         .11 

89.  90.  Papuina  novreguiueensis  Pfr.     Ku'ster,        .         .         .42 
91,  92.  Papuina  novseguineensis  Pfr.     Novit.,         .         .         .42 
93-95.  Papuina  braziene  Braz.     Specimen,    ....     43 

96.  Papuina  brazierre  Braz.     Mus.  Genov.,    .         .         .         .43 

97,  98.  Papuina  gestroi  T.-C.     Mus.  Genov.,          .         .         .44 

PLATE  6. 

99,  1.  Papuina  migratoria  Pfr.  Conch.  Cab.,  .  .  22,  23 
2,  3.  Papuina  migratoria  Pfr.  Conch.  Cab.,  .  .  22,  23 
4-6.  Papuina  alfredi  v.  trichroa  Mts.  C.  Mittheil.,  .  .12 

7,  8.  Papuina  alfredi  Cox.     P.  Z.  S., 12 

9.  Papuina  brodiei  Braz.     P.  Z.  S.,  .         .         .         .10 

10.  Papuina  spectrum  Rv.  (— multizona).     C.  Ic.,          .         .     39 

11,  12.  Papuina  walleri  Braz.  (brenchleyi  Ang.).     P.    Z.  S. 

18v  8,     .         .         .         .         .         .  .         . .       .         .12 

13,  14.  Papuina  boivini  Petit.  Specimen,  ....       6 

15,  16.  Papuina  boiviiii  Petit.  C.  Cab.,  ....       6 

17,  18.  Papuina  moseleyi  Sm.  P.  Z.  S.,  .         .         .         .     72 

19,  20.  Papuina  boiviui  Petit.  Specimen,  ....       6 

PLATE  7. 

21-26.  Papuina  lambei  Pfr.     C.  Cab.  and  Specimens,  .  .     48 

27,  28.  Papuina  tomasinelliana  T.-C.     Mus.  Genov.,  .  .     44 

29,  30.  Papuina  mysolensis  Pfr.     Novit.  Conch.,    .  .  .43 

31,  32.  Papuina.  thomsoni  Sm.     Ann.  Mag.  N.  H.,  .  .     o2 

33,  34.  Papuina  phseostyla  Mts.     Novit.  Conch.,    .  .  .47 


214  EXPLANATION    OF   PLATES. 

FIGURE.  PAGE. 

35,  38,  39.  Papuina  gaberti  Less.     Specimens,       .  .  .48 

36,37.  Papuina  gaberti  Lets,  var.?     C.  Cab.,         .  .  .48 

PLATE  8. 

40,  41.  Papuina  euchroes  Pfr.     Novit.  Conch.,       .  .  .23 

42-45.  Papuina  pileus  Mull.     Specimens,      .         .  .     24 

46,  47.  Papuina  poiretiana  Pfr.     Specimens,           .  .  .27 

48,  49.  Papuina  poiretiana  Pfr.     Conch.  Cab.,        .  .  .27 

50,  51.  Papuina  conscendens  Cox.     Mon.  Austr.  L.  Sh.,  .     14 
52-54.  Papuina  amphizona  Pils.     Specimens,         ...       5 

55,  56.  Papuina  mendana  Ang.         .     Conch.  Cab.,  .  .     17 

57,  58.  Papuina  gsertneriana  Pfr.     Couch.  Cab.,     .  .  .26 

PLATE  9. 

59-62.  Papuina  meta  Pfr.     Conch.  Cab.,        .         .  .  .17 

63.  Papuina  meta  Pfr.     Specimen,         .         .         .  .  .17 

64.  Papuina  meta  v.  acmella  Pfr.     Specimen,         .  .  .18 

65.  66.  Papuina  meta  v.  acmella  Pfr.     Conch.  Cab.,  .  .18 
67,  68.  Papuina  xanthochila  Pfr.     Conch  Cab.,     .  .  .15 

69.  Papuina  xanthochila  var.     P.  Z.  S.,          .         .  .  .15 

70.  Papuina  deidamia  Ang.  (—meta).     P.  Z.  S.,    .  .  .18 

71.  Papuina  lenta  Pfr.     Conch.  Cab.,     .         .         .  .  .23 

72.  Papuina  congrua  Pfr.     P.  Z.  S.,                .         .  .  .4 

73.  74.  Papuina  hermione  Ang.     P.  Z.  S.,               .  .  .     21 
75,  76.  Papuina  zose  Pfr.  (— blainvillei).     Novit.  Conch.,  25,  26 

PLATE  10. 

77,  78.  Papuina  lituus  Less.     Conch.  Cab.,     .         .  .  .37 

79,  80.  Papuina  eddystonensis  Rv.     Conch.  Cab.,  .  .     64 

81,  82.  Papuina  pennantiana  Pfr.     Conch.  Cab.,    .  .  .36 

83-85.  Papuina  multizona  Less.     Voy.  Astrol.,      .  .  .39 

86.  Papuina  adonis  Ang.     P.  Z.  S.,        .         .         .  .  .     20 

87.  Papuina  mendoza  Braz.     P.  Z.  S.,    .         .         .  .     '    .     21 

88.  89.  Papuina  plagiostoma  Pfr.     Specimens,        .  .  .19 

90.  Papuina  choisuelensis  Braz.     P.  Z.  S.,               .  .  .16 

91,  92.  Papuina  ccerulescens  Ang.     P.  Z.  S.,           ...     68 
93,  94.  Papuina  gelata  Cox.     P.  Z.  S.,    .         .         .  .  .     65 
95.  Papuina  guppyi  Smith  (enlarged).     P.  Z.  S.,    .  .  .19 

PLATE  11. 

96-98.  Papuina  rhynchostoma  Pfr.     Novit:  Conch.,  .  .     30 

99,  100,  1-3.  Papuina  pileolus  Fer.     Ostas.  Conch.,  .     29 

4,  5.  Papuina  lanceolata  Pfr.     Conch.  Cab.,          .  .  .34 

6,  7.  Papuina  niotacilla  Pfr.     Specimen,       .         .  .  .66 

8,  9.  Papuina  nigrofasciata  Pfr.     Conch.  Cab.,      .  .  .71 

10,  11,  2.  Papuina  aurora  Pfr.     Novit.  Conch.,      .  .  .41 


EXPLANATION    OF    PLATES.  215 

FIGURE.  PAGE. 

13-15.  Papuina  lienardiana  Crosse.     J.  de  C.,  69 

16-19.  Papuina  lienardiana  Crosse.     Specimens,  .  .     69 

PLATE  12. 

18,  19.  Papuina  labium  Fer.     Conch.  Cab.,    .         .  .  .38 

20,  21.  Papuina  pseudolabium  Pfr.     Conch.  Cab.,  .  .     38 

22-24.  Papuina  leucothoe  Pfr.     Novit.  Conch.,      .  .  .66 

25-27.  Papuina  labium  Fer.     Histoire,          .         .  .  .38 

28-30.  Papuina  pelechystoma  T.-C.     Mus.  Genov.,  .  .     35 

31-34.  Papuina  carinata  H.  &  J.     Astrol.  &  Zel.  .  .     36 

35,  36.  Papuina  vitrea  Fer.     Ostas.  Conch.,  .         .  .  .33 

PLATE  13. 

34,  35.  Papuina  nodifera  Pfr.     Novit.  Conch.,        .  .  .37 

36,  Papuina  lactiflua  Pfr.     Novit.  Couch.,      ....  108 

37,  38.  Papuina  isabellensis  (==lactiflua).     J.  de  C.,  .  .  108 
39,  40.  Papuina  cymodoce  Crosse.     J.  de  C.,           .  .  .107 

41.  Papuina  hunteri  Cox.     Specimen,    .         .         .  .  .105 

42,  43.  Papuiua  pyxis  Hinds.     Specimen,      .         .  .  .80 

44,  45.  Papuina  hunteri  Cox.     Conch.  Cab.,           .  .  .  105 
46,  47.  Papuina  purchasi  Pfr.     Conch.  Cab.,           .  .  .108 

45,  49.  Papuina  boyeri  C.  &  F.     J.  de  C.,      .         .  .  .47 
50,  51.  Papuina  grata  Mich.     Conch.  Cab.,    .         .  .  .35 
52,  53.  Papuina  vitracea  Fer.     Histoire,        ....  1.06 
54,  55.  Papuina  (?)  subvitracea  Pfr.     Novit.  Conch.,  .  .  107 

PLATE  14. 

56,  57.  Papuina  antiqua  A.  &  R.     Conch.  Cab.,     .  .  .28 

58.  Papuina  horderi  Sowb.     P.  Z.  S.,              .         .  .  .  .  29 

59-63.  Papuina  beatrix  Ang.     P.  Z.  S.,          .         .  .  .15 

64,  65-67.  Papuina  vexillaris  Pfr.     Conch.  Cab.,  .  .  .46 

68,  69.  Papuiua  gurgustii  Cox.     P.  L.  S.  N.  S.  W.,  .     61 

70.  Papuina  chancei  Cox.     P.  Z.  S.,                .         .  .  .5 

71,  72.  Cochlostyla  hinclei  Cox.     P.  L.  S.  N.  S.  W.  (see  next 

vol.). 

PLATE  15. 

72-74.  Papuiua  helicinoides  H.  &  J.     Specimen,  .  .  .76 

75.  Papuina  sellers!  Cox.     P.  Z.  S.,        .         .         .  .  .51 

76-78.  Papuina  eva  Pfr.     Specimens, 78 

79,  80.  Papuina  cyrene  Crosse.     J.  de  C.,       .         .  .  .78 

81-83.  Papuina  helicinoides  var.  cleryi  Reel.,         .  .  .76 

84,  85.  Papuina  eva  Pfr.     Conch.  Cab.,          .         .  .  .78 
86-87.  Papuina  helicinoides  var.  cleryi  Reel.     Specimen,       .     76 

88,  89.  Papuina  cineracea  H.  &  J.     Specimen,       .  .  .77 

90.  Papuina  zelina  Cox.     P.  Z.  S.,          .         .         .  .  .     78 


216  EXPLANATION    OF    PLATES. 

FIGURE.  PAGE. 

91.  Papuina  labillardieri  Sm.     Specimen,       .         .         .  .75 

92,  93.  Papuina  labillardieri  Sm.     P.  Z.  S.,             ...     75 
94-97.  Papuina  layardi  Hartm.     Specimens,          .         .  .79 
98,  99.  Papuina  eros  Aug.     Specimens,           .         .         .  .70 

100,  101.  Papuina  eros  Aug.     P.  Z.  S., 70 

PLATE  16. 

1-4,  Papuina  eva  Pfr.     Specimens,      .         .         .         .  .78 

5.  Papuina  heliciuoides  v.  meridionalis  Sm.     P.  Z.  S.,  .     77 

6,  7.  Papuina  cineracea  H.  &  J.     Specimen,         .         .  .77 

8.  Papuina  fringilla  Pfr.     Specimen,    .         .         .         .  .73 

9,  10.  Papuiua  fucata  Pfr.     Mon.  Aust.  L.  Sh.,    .       • .  .14 
11-1.3.  Papuina  gelata  v.  maddocksi  Braz.     Specimens,  .     66 
14,  15.  Papuina  rhombostoma  Pfr.     Couch.  Icon.,          .  .     60 

16.  Papuiua  ambrosia  Ang.     Specimen,          ....       7 

17.  Papuiua  malantensis  Aug.     Specimen,     ....       7 

18.  Papuina  dampieri  Sm.     P.  Z.  S.,               .         .         .  .11 

19.  Papuina  ramsdeni  Ang.  (—ambrosia).     P.  Z.  S.,  .7 
20-22.  Papuina  malantensis  Aug.     P.  Z.  S.,           .         .  .7 
23,  24.  Papuina  guadalcanarensis  var.     Specimen,         .  .       9 
25,  26.  Papuina  vexillaris  Pfr.     Specimen,    .         .         .  .47 

PLATE  17. 

27.  Papuina  meta  var.  bicolor  Pils.     P.  Z.  S.,         .         .  .18 

28,  29.  Papuina  tun-is  Ad.     P.  Z.  S.,      .  .32 

30.  Papuina  triumph alis  Rve.     C.  Icon.,        .         .         .  .42 

31.  Papuina  migratoria  Pfr.     P.  Z.  S.,            .         .         .  .     22 

32.  Papuina  migratoria  Pfr.     Specimen,         .         .         .  .22 

33.  Papuina  blanfordiana  Ad.     P.  Z.  S.,                 .         .  .     26 

34.  35.  Papuiua  waigionensis  Ad.     P.  Z.  S.,  .         .         .  .43 

36.  Papuina  xanthosoma  Pils.     Specimen,      .         .         .  .28 

37,  38.  Papuina  comriei  Aug.     P.  Z.  S.,         .         .         .  .     64 

39.  Papuina  antiqua  A.  &  R.     Voy.  Samarang.,    .         .  .28 

40,  41.  Papuiua  tayloriana  A.  &  R,     Voy.  Samarang.,  .     58 

42.  Papuina  coniformis  var.  tuifettii  Less.     Voy.  Coq.,  .     51 

43.  Papuina  coniformis  Fer.     Histoire,           .         .         .  .50 

PLATE  18. 

44.  45.  Satsuma  perakensis  Crosse.     J.  de  C.,  82 
46,  47.  Satsuma  subperakensis  Pils.     Specimen,     .         .  .82 
48-50.  Satsuma  rostrella  Pfr.     Novit.  Conch.,         .         .  .83 

51.  Satsuma  bantamensis  Sm.     Soc.  Mai.  Belg.,     .         .  .84 

52.  Satsuma  subflava  Aust.     P.  Z.  S.,     .         .         .         .  .     85 

53.  54.  Satsuma  (?)  scenoma  Bens.     Conch.  lud.,  .         .  .83 
55-58.  Satsuma  (?)  wrayi  de  Morg.     Bull.  Soc.  Zool.  Fr.,  .     86 
59.  Coliolus  arfakiensis  T.-C.     Mus.  Genov.,           .         .  .87 


EXPLANATION    OF    PLATES.  217 

FIGURE.  PAGE. 

60-63.  Satsuma  (?)  hardouini  de  Morg.     Soc.  Zool.  Fr.,  .     86 

64.  Helix  testudo  Pfr.     P.  Z.  S., 89 

65.  Satsuma  tigsensis  Anst.     P.  Z.  S..              .         .         .  .     85 

66.  Satsuma  niahensis  Aust.     P.  Z.  S.,  .         .         .         .  .     85 

67.  68.  Helix  trocbalia  Bens.     Specimen,       .         .         .  .88 
69-71.  Corasi a  tricolor  v.  picta  Sm.     P.  Z.  S.,        .        .  .  112 

PLATE  19. 

1,  2.  Albersia  granulata  Q.     Conch.  Cab.  and  Voy.  Astrol.,     90 

3.  Albersia  confbrmis  Fer.     Conch.  Icon.,    .         .         .  .113 

4,  5.  Albersia  conformis  Fer.     Histoire,       .         .  .113 

6.  Albersia  zouulata  Fer.   var.  recluziana  Guill.      Conch. 

Icon.,     ..........     91 

7,  8.  Albersia  zonulata  Fer.     Conch.  Cab.,  .         .         .  .91 
9-11.  Albersia  pubicepa  Mts.     Ostas.  Landschn.,          .  .     90 

12,  14.  Pfeifferia  micans  Pfr.     Conch.  Cab.,   .         .         .  .128 

13.  Pfeifferia  micans  Pfr.     Specimen,     .         .         .         .  .128 

PLATE  20. 

1,  2.  Axina  garibaldiana  D.  &  S.     Novit.  Conch.,        .  .159 

3-6.  Axina  zebuensis  Brod.     Couch.  Cab,  and  Specimen,  .  1 61 

7.  Axina  striatissima  Pils.     Specimen,          .         .         .  .162 

8.  Axina  siquijorensis  Brod.     Specimen,      .         .         .  .159 

9.  10.  Axina  siquijorensis  Brod.     Conch.  Cab.,     .         .  .159 

PLATE  21. 

11,  12,  14.  Chloraea  thersites  Brod.     Obras  Mai.,             .  .  104 

13.  Chlorsea  thersites  Brod.     Specimen,          .         .         .  .104 

15.  Chlorsea  sirena  var.  guimarasensis.     Specimen,         .  .     98 

16-18.  Chlorsea  sirena  Beck.     Specimen,       .         .         .  .98 

19,  20.  Chlorsea  sirena  Beck.     Couch.  Cab.,             .         .  .98 

21,  22.  Chlorsea  amoena  Pfr.     Conch.  Cab.,    .         .         .  .98 

23-25.  Chlorsea  benguetensis  Semp.     Phil.  Laudm.,       .  .     96 

26.  Corasia  rossiteri  Cox.     P.  Z.  S.,        .         .         .         .  .109 

27,28.  Chlorsea  paradoxa  v.  immaculata.     Couch.  Cab.,  .  102 

29,30.  Chlorsea  gmeliuiana  Pfr.     Couch.  Icon.,     .         .  .100 

31,  32,  33.  Chloraaa  fibula  var.                     .     J.  de  C.,    .  .     94 

34,  35.  Chlorsea  antonii  Semper.     Reiseu,       .         .         .  .97 

36.  Corasia  dealbata  Brod.     Conch.  Icon.,      .         .         .  .119 

37,  38.  Axina  magistra  var.  gloynei  Sowb.     Obras  Mai.,  .  165 

PLATE  22. 

1-7.  Chlorsea  dryc-pe  Brod.     Specimens,       .         .         .  .100 

8,  9.  Chlorsea  uudiua  Pfr.     I^ovit.  Couch.,    .         .         .  .103 

10,  11.  Chlorsea  pelewaua  Mouss.     J.  de  C.,            .         .  .99 

12-14.  Chlonea  paradoxa  Pfr.     Specimen,     .         .         .  .102 


218  EXPLANATION    OF    PLATES. 

FIGURE.  PAGE. 

15.  Chlorsea  hugeli  Pfr.     Specimen,        .         .         .         .         .1)6 

16,  17.  Chlorsea  restricta  Pir.  (=constricta  Pfr.).     Specimen,  103 
18-20.  Chloraa  hugeli  Pfr.     Conch.  Cab.,      .         .         .         .96 
21-23.  Chlortea  fibula  Brod.     Conch.  Cab.,    .         ...     94 
24-26.  Chlonea  hanleyi  Pfr.     Specimen.         .         .         .         .95 
27,  28.  Chlorsea  hanleyi  Pfr.  var.     Conch.  Cab.,     .         .         .95 

PLATE  23.  • 

1-3.  Corasia  tricolor  Pfr.     Specimen,   .  .  .  .  .111 

Fig.  below  fig.  2.  Corasia  transenna  Pils.  Specimen,  .  .112 

4-6.  Corasia  tricolor  Pfr.     Conch.  Cab.,  .  .  .  .111 

8-12.  Corasia  aphrodite  Pfr.     Specimens,  .  .  .  .109 

13,  14.  Corasia  psyche  Ang.     P.  Z.  S.,   .  .  .  .  .110 

15.  Corasia  balcomei  Cox,     P.  Z.  S.,       .  .  .  .  .111 

16-18.  Corasia  transenna  Pils.     Specimen,  .  .  .  .112 

PLATE  24. 

19-21.  Corasia  puella  Brod.  var.  subpuella  Pils.     Specimen, .  121 
22,  23.  Corasia  puella  Brod.     Conch.  Cab  ,    .         .         .         .120 

24.  Corasia  puella  var.  subpuella  Pils.     Specimen,         .         .  121 

25.  Corasia  puella  Brod.     Specimen,      .....  120 

26.  Corasia  puella  var.  subpuella  Pils.  (typical  of  var.).   Novit. 

Conch.,          .........  121 

27.  28.  Corasia  broderipii  Pfr.     Specimen,      .         .         .         .123 
29,  30.  Corasia  virgo  Brod.     Specimen,          .         .         .         .119 

31.  Corasia  nymph  a  Pfr.  (==filaris  Val.).     Conch.  Icon.,         .  122 

32,  33.  Corasia  broderipii  Pfr.     Specimen,      ....  123 
34-36.  Corasia  papyracea  Brod.     Specimen,  .         .         .         .117 

PLATE  25. 

37,  38.  Corasia  casta  Pfr.     Conch.  Cab.,         .  .  .  .120 

39.  Corasia  puella  Brod.  large  var.     Obras  Mai.,  .  .121 

40.  Corasia  puella  var.  lais  Pfr.     Obras  Mai.,  .  .  .  121 

41.  Corasia  lais  Pfr.  typical.     Conch.  Icon.,  .  .  121 
42-44.  Corasia  almse  Moll.     Landschu.  Cebu,  .  .  .117 
45,46.  Corasia  reginre  Brod.     Specimen,        .  .  .  .116 
47,  48.  Corasia  elizabethre  Semp.     J.  de  C.,    .  .  .  .117 
49-51.  Corasia  psittacina  Dh.     J.  de  C.,         .  .  .  .118 
52-54.  Corasia  tilaiis  Val.     Obras  Mai.,  .  122 

PLATE  26. 

1.  Corasia  segrota  Reeve.     Conch.  Cab.,        .                  .  .124 

2,  3.  Calocochlea  lallaoensis  Pfr.     Specimen,        .         .  .136 
4.  Corasia  seruginosa  Pfr.     Conch.  Icon.,      .         .         .  .122 
5-7.  Chromatosphsera  aurata  Sowb.     Conch,  and  C.  Cab.,  .  170 
8,  9.  Corasia  eydouxi  Hid.     Conch.  Cab.,     .         .         .  .123 


EXPLANATION    OF    PLATES.  219 

FIGURE.  PAGE. 

10-12.  Corasia  filaris  Val.     Specimen,  .....  122 
13-15.  Corasia  samboanga  H.  &  J.     Astro!.  &  Zelee,    .         .124 

16,  17.  Corasia  leucophthalma  Pfr.     Novit.  Conch.,       .         .  113 

PLATE  27. 

1,  Corasia  intaminata  Old.     U.  S.  Exp.,       .         .         .         .125 

2,  3.  Corasia  phy  sails  Pfr.     Novit.  Conch.,  .         .         .         .115 

4,  5.  Eudoxus  halichlora  Semp.     J.  de  C.  and  Specimen, 

6-8.  Corasia  coelaxis  Pils.     Specimen,           .  .  .  .114 

9,  10.  Corasia  extensa  Pfr.     Conch.  Cab.,     .  .  .  .114 

11.  Corasia  extensa  Pfr.     Specimen,       .         .  .  .  .114 

12.  Corasia  najas  Pfr.  var.     Specimen,  .         .  .  .  .115 

13.  Corasia  uajas  Pfr.  typical.     Ostas.  Conch.,  .  .  .  115 

14.  15.  Corasia  obliqnata  Dh.     Histoire,         .  .  .  .116 

PLATE  28. 

16,19,21.  Corasia  intorta  Sowb.     Specimens,         .         .         .  125 

17,  22.  Corasia  intorta  Sowb.     Conch.  Icon.,          .         .         .125 

18,  23.  Corasia  iutorta  Sowb.     Conch.  Cab.,  .         .         .  125 
20,  24.  Corasia  intorta  Sowb.     Obras  Mai.,    ....  125 
25,  26.  Corasia  limansauensis  Semp.     Reis.,  ....  126 

27,  28.  Corasia  magtanensis  Semp.     Reis.,      ....  125 
29,  30.  Corasia  irosinensis  Hid.     Obras  &  J.  de  C.,        .         .  121 
31,  32.  Corasia  najas  Pfr.     Abbild., 115 

PLATE  29. 

1,2.  Helicostyla  thomsoni  Pfr.     Novit.  Conch.,   .         .         .  185 

3,  4.  Helicostyla  thomsoni  Pfr.     Specimen,  .         ".         .  185 

5,  6.  Leytia  fragilis  Sowb.     J.  de  C.,     .         .         .         .         .129 
7-9.  Calocochlea  tukanensis  Pfr.     Novit.  Conch.,          .         .132 

10,  11.  Calocochlea  decipiens  Sowb.     Conch.  Cab.,         .         .  140 

12.  Calocochlea  tukanensis  Pfr.     Specimen,  ....  132 

13,  14.  Calocochlea  cromyodes  Pfr.     Conch.  Cab.,  .         .  130 

15.  16.  Calocochlea  cromyodes  Pfr.     Specimen,      .         .         .130 

PLATE  30. 

17, 19.  Calocochlea  melanochila  Val.     Specimens,          .  .  150 

18.  Calocochlea  melanochila  Val.     Conch.  Cab.,     .         .  .150 

20,21.  Calocochlea  infuscata  Alb.     Novit.  Conch.,         .  .  152 

22.  Calocochlea  solida  Pfr.     Conch.  Cab.,       ....  153 

23,  24.  Calocochlea  roissyana  var.  lutea.     Novit.  Conch.,  .  151 
25-27.  Calocochlea  roissyana  Fer.     Specimens,      .         .  .  151 

28,  Calocochlea  roissyana  var.  subatra.     Conch.  Cab.,    .  .152 

29,  30.  Calocochlea  dimera  Jonas.     Specimens,      .         .  .  156 


220  EXPLANATION  OF  PLATES. 

PLATE  31. 

FIGURE.  PAGE. 

31,  32.  Calocochlea  spliserion  var.  intincta  Shutt.     Abbild.,    .  154 

33.  Calococlilea  sphserion  var.  intincta  Shutt.     Specimen,       .  154 

34,  35.  Calocochlea  sphserion  Sow.,  typical.     Conch.  Cab.,      .  154 
36,  37.  Helicobulinus  bembicocles  Pfr.     Conch.  Cab.,     .         .  198 
38.  Helicostyla  mirabilis  var.  trichroa.     Specimen,         .         .182 
Fig.  below  fig.  38.  Helicostyla  tephrodes  Pfr.     Couch.  Icon.,  183 

40.  Helicostyla  mirabilis  Fer.     Specimen,      ....  181 

41.  Calocochlea  norrisii  Sowb.     Conch.  Cab.,          .         .         .  143 

42.  Calocochlea  norrisii  Sowb.     Conch.  Icou.,         .         .         .143 

PLATE  32. 

43-50.  Calocochlea  pollilensis  Pfr.  and  van*.     Novit.  Couch.,  138 

51.  Calocochlea  augusta  var.  (?)     Specimen,  .         .         .137 

52.  Calocochlea  dubiosa  Pfr.     Specimen,        .         .         .         .135 

53.  Calocochlea  batanica  Rv.  (=dubiosa).     Conch.  Icon.,      .  135 

54.  Calocochlea  batanica  Rv.     Conch.  Cab.,  .         .         .         .135 


00 


PLATE  66. 

55.  Corasia  (?)  bourdilloni  Theob.     J.  A.  S.  B 127 

56,56.  Calocochlea  pan  Brocl.     Specimens,    .         .  .  .149 

57.  Calocochlea  pan  Brod.     Conch.  Cab.,       .         .  .  .  149 

58.  Calocochlea  princeps  Rv.,  typical.     Conch.  Icon.,  .  .137 

59.  Calocochlea  princeps  Rv.  var.     Specimen,        .  .  .137 

60.  Calocochlea  pulcherriua  Sowb.     Conch.  Cab.,  .  .  .133 
61-64.  Calocochlea  pulcherrima  Sowb.     Specimens,  .  .133 
65.  Helicostyla  ignobilis  Sowb.     Conch.  Cab.,         .  .  .180 

PLATE  34. 

1,  2.  Calocochlea  samarensis  Semper.     Reiseu,      .  .  .  146 

3,  4.  Calocochlea  samarensis  var.  ?     Reisen,          .  .  .146 

5,  6.  Calocochlea  coronadoi  Hid.     J.  de  C.,          .  .  .142 
7,  8.  Calocochlea  amicta  Rv.     Conch.  Cab., 

9.  Calocochlea  chrysochila  Sowb.     Specimen,       .  .  138 

10.  Calocochlea  chrysochila  Sowb.     Couch.  Cab.,  .  .  .138 

PLATE  35. 

11,  12.  Calocochlea  festiva  var.  anna?  Semp.     J.  de  C.,  . 

13,  16.  Calocochlea  festiva  Don.     Specimens, 

14,  15.  Calocochlea  festiva  Don.     Conch.  Cab.,      .  .  134 
17,18.  Calocochlea  augusta  Alb.     Novit.  Conch.,  .         .         .  136 

19.  22.  Calocochlea  zonifera  Sowb.     Conch.  Cab.,  .  .  141 

20.  Calocochlea  zonifera  Sowb.     Conch.  Icon., 

21.  Calocochlea  coronadoi  Hid.     Specimen,  . 

22.  Calocochlea  zonifera  Sowb.     Conch.  Cab., 

23.  24.  Calocochlea  obtusa  Pfr.     Conch.  Cab.,        .         .         .132 


EXPLANATION  OF  PLATES.  221 
PLATE  36. 

FIGURE.  PAGE. 

25-27.  Helicostyla  metaformis  Fer.     Specimens,    .  .  .186 

28,  29.  Helicostyla  metaformis  Fer.     Couch.  Cab.,  .  .  186 

30.  Helicostyla  languida  Pfr.     Conch.  Icon.,          .  .  .  189 

31,32.  Helicostyla  tenera  Pfr.     Conch.  Cab.,          .  .  .  179 

33.  Helicostvla  unica  Rv.     Conch.  Icon.,        ....  189 

•/ 

34.  Helicostyla  nux  Semper.     Reisen.,  ..... 
35-37.  Helicostyla  hydrophana  Sow.     Specimens,  .  .187 

PLATE  37. 

38,  39.  Calocochlea  chlorochroa  Sowb.     Couch.  Cab.,  .  .150 

40.  Calocochlea  dalmahoyi  Pfr.     Specimen,    ....  143 

41,  42.  Calocochlea  dalmahoyi  Pfr.     Xovit.  Conch.,  .  .143 
43,44.  Eudoxus  jonasi  Pfr.     Conch.  Cab.,     .... 

45,  46.  Helicostyla  rehbeini  Pfr.     Conch.  Cab.,      .  .  .  190 

47,  48.  Helicostyla  curta  Pfr.     Conch.  Cab.,  .         .  .  .192 

49.  Helicostyla  senckendorffiaua  Pfr.     Conch.  Cab., 

t/ 

50,  51.  Helicostyla  butleri  Pfr.     Conch.  Cab.,        .  .  .188 

PLATE  38. 

70.  73.  Cochlodryas  florida  Sowb.     Conch.  Cab.,  .  .177 

71.  Cochlodryas  florida  Sowb.     Histoire,        ....  177 

72.  Cochlodryas  florida  Sowb.     Specimen,      .         .  .  .177 
74-78.  Cochlodryas  polychroa  Sowb.     Specimen,  .  .  .178 
79,  80.  Helicostyla  fulgeus  Sowb.     Specimen,         .  .  .182 

81.  Helicostyla  fulgens  Sowb.     Conch.  Cab.,           .  .  .  182 

PLATE  39. 

82,  83.  Pachysphrera  sphserica  Sowb.     Conch.  Cab.,  .  .172 

84.  Pachysphsera  sphserica  Sowb.     Specimen,         .  .  .  172 

85,  86.  Pachysphrera  annullata  Sowb.     Conch.  Cab.,  .  .174 
87,  88.  Pachysphsera  annullata  Sowb.     Conch.  Icon.,  .  .  174 
89,  90.  Pachysphsera  balteata  Sowb.     Conch.  Cab.,  .  .173 
91,  92,  93.  Eachysphsera  balteata  Sowb.     Specimens,  .  .  173 
94,  95.  Orustia  monticula  Sowb.     Specimens,         .  .  .176 

96,  Orustia  monticula  Sowb.     Conch.  Cab.,    ....  176 

97,  99.  Pachysphsera  iloconensis  Sowb.     Conch.  Cab.,  .  .  175 

98,  100.  Pachysphaera  iloconensis  Sowb.     Specimens,  .  .175 

PLATE  40. 

1.  Orthostylus  villari  Hid.     J.  de  C.,    .  .  .  201 

2.  Orthostylus  pithogaster  Fer.     Conch.  Icon.,      .  .  .  200 
3-5.  Orthostylus  pithogaster  Fer.     Specimens,     .  .  .  200 
6,  7.  Axina  bruguieriana  Pfr.     Conch.  Cab.,         .  .  .166 
8,  9.  Axina  phloides  Pfr.     Conch.  Cab.,       .         .  .  .163 


222  EXPLANATION  OF  PLATES. 

PLATE  41. 

FIGURE.  PAGE. 

10-12.  Orthostylus  lignaria  Pfr.     Specimens,  .  .  .  204 

13.  Orthostylus  lignaria  Pfr.     Conch.  Icon.,  .  .  .  204 

14-16.  Axina  montfortiana  Pfr.     Specimen,  .  .  .165 

17-19.  Axina  magistra  Pfr.     Conch.  Cab.,     .  .  .  .164 

PLATE  42. 

20-22.  Orthostylus  daphnis  Brod.     Specimens,  .  .  .  201 

23.  Orthostylus  daphnis  Brod.     Conch.  Icon.,  .  .  .  201 

24,  25.  Orthostylus  gilvus.Brod.     Conch.  Icon.,  .  .  .  205 

PLATE  43. 

26,  27.  Orthostylus  rufogaster  Less.     Specimens,  .  .  .  207 

28.  Orthostylus  woodiana  Lea.     Specimens,  ....  206 

29.  Orthostylus  woodiana  Lea  (Reevei  Brod.).     Conch.  Icon.,  206 
30-32.  Axina  carbonaria  Sowb.     Specimen,  .  .  .163 

PLATE  44. 

33.  Axina  carbonaria  Sowb.     Specimen,         .  .  .  .163 

34,35.  Axina  pfeifferi  Semper.     Specimen,    .  .  .  .162 

36,  37.  Axina  pfeifferi  Semper.     Conch.  Cab.,  .  .  .162 

38,  39.  Orthostylus  portei  Pfr.     Novit.  Conch.,  .  .  .206 

40.  Orthostylus  bicolorata  Lea.     Couch.  Icon.,  .  .199 

tf 

41.  Orthostylus  bicolorata  Lea.     Specimen,    .  .  .  .199 

PLATE  45. 

42-44.  Orthostylus  imperator  Pfr.     Specimens,  .  .  .  199 

45,46.  Axina  moreleti  Pfr.     Conch.  Cab.,      .  .  .  .165 

47.  Orthostylus  monozonu?  Pfr.     Conch.  Icon.,  .  .  .  208 

48.  Orthostylus  marinduqueusis  Hid.  (woodiana  Pfr.).  Conch. 

Icon.,     ......         ^  ...  209 

49.  Orthostylus  marinduquensis  Hid.     Specimen,  .  .  .  209 

PLATE  46. 

50,51.  Helicobulinus  turgens  Dh.     Conch.  Cab.,  .  .  .196 

52.  Helicobulinus  turgens  Dh.     Specimen,     .  .  .  .196 

53,54.  Helicostyla  indusiata  Pfr.     Novit.  Conch.,  .  .  184 

56,  57.  Helicostyla  collodes  Sowb.     Specimens,  .  .  .184 

PLATE  47. 

58.  Helicobulinus  sarciuosa  Fer.     Conch.  Cab.,  .  .  .195 

59.  Helicobulinus  turbo  Pfr.     Conch.  Icon.,  .  .  .  .137 

60.  Helicobulinus  ciuerasceus  Pfr.     Conch.  Icon.,  .  .197 

61.  Calocochlea  mindanaensis  Sowb.     Conch.  Cab.,  .  .148 


EXPLANATION    OF    PLATES.  223 

FIGURE.  PAGE. 

62.  Calocochlea  mindanaensis  Sowb.     Conch.  Syst.,        .  .  148 

63.  Helicobulinus  turbinoides  Dh.     Conch.  Syst.,            .  .196 

PLATE  48. 

65.  Calocochlea  ponderosa  Pfr.     Conch.  Icon.,        .         .  .147 

66,  67.  Helicobulinus  grandis  Pfr.  (carolus  Dh.).     J.  de  C.,  .  195 

68.  Helicobulinus  grandis  Pfr.     Specimen,     .         .         .  .195 

69.  Ptychostylus  cepoides  Lea.     Specimen,    .         .         .  .194 

70.  71.  Calocochlea  matruelis  Sowb.     Specimen,    .         .  .151 
72.  Calocochlea  difficilis  Pfr.     Conch.  Icon.,  ....  151 

PLATE  49. 

25-28.  Orthostylus  ticaonica  Brod.     Specimens,     .         .  .  203 

29.  Helicobulinus  turbinoides  Dh.     Specimen,       .         .  .196 

30.  Orthostylus  lignaria  Pfr.     Specimen,        ....  204 

PLATE  50. 

1.  Trachystyla  cryptica  Brod.     Specimen,    .         .         .  .167 

2.  Calocochlea  calliaudi  Dh.     Conch.  Icon.,          .         .  .144 

3.  Calocochlea  calliaudi  Dh.     Specimen,      .         .         .  .144 

4.  Calocochlea  depressa  Semp.     Conch.  Icon.,       .         .  .149 

5.  Calocochlea  depressa  Semp.     Couch.  Cab.,        .         .  .149 
6-8.  Calocochlea  depressa  Semp.  var.     Specimen,         .  .150 

PLATE  51. 

9.  Trachystyla  -cryptica Brod.     Conch.  Icon.,        .         .  .167 

10.  Trachystyla  latitans  Brod.     Conch.  Icon.,         .         .  .167 

11.  Trachystyla  latitans  Brod.     Specimen,     .         .         .  .167 

12.  Trachystyla  cretata  Brod.     Conch.  Icon.,          .         .  .168 

13.  Trachystyla  dataensis  Semp.     J.  de  C.,    .         .         .  .  169 

14.  15.  Trachystyla  dataensis  Semp.     Specimen,    .         .  .169 
16.  Calocochlea  lalloensis  Pfr.     Specimen,      ....  136 

PLATE  52. 

17-19.  Calocochlea  denticulata  Jay.     Type  specimen,    .  .  131 

20,21.  Calocochlea  albaiensis  Sowb.     Conch.  Cab.,         .  .  132 

22.  Calocochlea  generalis  Pfr.     Novit.  Conch.,       .         .  .137 

23,  24.  Calocochlea  decora  Ad.  &  Rv.     Voy.  Samarang.,  .  133 

25.  Calocochlea  festiva  Don.     Conch.  Cab.,    .         .         .  .134 

26.  Calocochlea  calliaudi  Dh.  var.     J.  de  C.,          .         .  .144 

27.  Calocochlea  calliaudi  Dh.  var.     Tr.  Am.  Phil.  Soc.,  .  144 

PLATE  53. 

28.  Calocochlea  circe  Pfr.     Specimen, 142 

29.  Calocochlea  circe  Pfr.     Conch.  Icon.,       .         .         .  .142 


224  EXPLANATION    OF    PLATES. 

FIGURE.  PAGE.. 

30,  31.  Calocochlea  zonifera  var.     Specimen,          .         .  .  142 

32,  33.  Calocochlea  zonifera  var.  paraleuca.     Specimen,  .  142 

34-36.  Calocochlea  microspira  Pfr.     Specimens,    .         .  .145 

37.  Calocochlea  codouensis  Hid.     J.  de  C.,     .         .         .  .  140 

PLATE  54. 

38-40.  Calocochlea  xanthobasis  Pils.     Specimen,  .         .  .  155 

41,  42.  CalococLlea  samareusis  Semp.  var.     J.  de  C.,      .  .  146 

43,  44.  Calocochlea  pulchra  Pils.     Specimen,          .         .  .  143 

45,  46.  Calocochlea  hemisphserion  Sowb.     Conch.  Cab.,  .  145 

47,  48.  Calocochlea  peraffinis  Pils.     Specimen,       .         .  .  139 

PLATE  55. 

49.  Calocochlea  harfordii  Brod.     Specimen,  .         .         .  .148 

50.  Calocochlea  harfordii  Brod.     Conch.  Syst,       .         .  .  148 

51,  53,  54.  Calocochlea  coccomelas  Brod.     Specimens,  .  .  146 

52,  55.  Calocochlea   coccomelas   Brod.     Conch.   Icon.    &    C. 

Syst.,     ..........  146 

56.  Calocochlea  mainitensis  Hid.     J.  de  C.,    ....  152 

PLATE  56. 

57,  58.  Calocochlea  crossei  Hid.     J.  de  C.,     .         .         .  .  156 
59,  60.  Calocochlea  retusa  Pfr.     Conch.  Cab.,         .         .  .  157 

61.  Calocochlea  libata  Rv.  var.     J.  de  C.,      ....  157 

62,  63.  Calocochlea  libata  Rv.     Conch.  Cab.,          .         .  .  157 
64,  65.  Chromatosphaera  erubesceus  Semp.     Reisen.,       .  .  170 
66,  67.  Chromatosphrera  erubescens  v.  luteocincta.     Reisen.,  .  171 
68,  69.  Chromatosphsera  aurata  Sow.     Specimen,            .  .170 

70.  Chroraatosphsera  aurata  Sow.     Obras  Mai.,      .         .  .170 

71,  72.  Chromatosphaera  lividocincta  Semp.     Reisen.,    .  .  171 

PLATE  57. 

73-75.  Axina  striatissima  Pils.     Specimen,    .         .         .  .162 

76.  Axina  magistra  Pfr.     Obras.  Mai.,  .         .         .         .  .164 

77,  78.  Axina  bruguieriana  Pfr.     Specimen,           .         .  .166 
79.  Helicostyla  mirabilis  Fer.     Conch.  Cab.,           .         .  .  181 
80-82.  Helicostyla  mirabilis  Fer.     Specimens,        .         .  .181 
83-86.  Helicostyla  mirabilis  Fer.     Obras  Mai.,      .         .  .181 

87.  Helicostyla  mirabilis  var.  near  trichroa.     Conch.  Cab.,     .  182 

PLATE  58. 

88.  Helicostyla  metaformis  Fer.  var.     Specimen,             .  .  186 

89.  Helicostyla  metaformis  var.     J.  de  C.,               .         .  .186 

90.  Helicostyla  metaformis  var.     Specimen,  .                  .  .  186 

91.  Helicostyla  rustica  Mouss.     Conch.  Icon.,         .         .  .187 


EXPLANATION    OF    PLATES.  225 

FIGURE.  PAGE. 

92.  Helicostyla  rustica  Mouss.     Moll.  Java,  ....   187 

93,  94.   Helicostyla  montana  Serap.     Reisen.,          .         .         .  191 

95.  Helicostyla  tenestrata  Sowb.     C.  Icon 192 

96.  Helicostyla  fenestrata  Sowb.     Specimen,  .         .         .192 

97.  Helicostyla  papuensis  Hedley.     P.  L.  S.  N.  S.  W.,  .  190 

98.  99.   Helicostyla  orbitula  Sowb.     Conch.  Icon.,  .         .  179 

PLATE  59. 

1,  2.  Helicostyla  lacera  Pfr.     Conch.  Icon.,  .         .         .189 

3.  Trachystyla  cineracea  Semp.     Reisen.,     .         .         .         .168 

4,  o.  Helicostyla  plurizonata  A.  &  R.     Samarang.,       .         .183 

6.  Orthostylus  juglans  Pfr.     Conch.  Icon.,    ....  209 

7,  8.  Helicobulinus  grandis  Pfr.     Typical.     Specimen,          .  195 

PLATE  60. 

9.   Helicobuliuus  grandis  Pfr.     Conchyl.  Cab.,      .         .         .195 
10,11.  Helicobulinus  grandis  Pfr.  var.     Specimen,        .         .  195 

12.  Orthostylus  vidali  Hid.     J.  de  C., 208 

13,14.  Helicostyla  dilatatus  Pfr.     Conch.  Cab.,     .         .         .  193 

15.  Helicostyla  curta  Sowb.     Specimen,          .         .         .         .193 

16.  Helicostyla  curta  var.  dentata.     Specimen,       .         .         .193 

PLATE  61. 

17.  Orthostylus  faunus  Brod.     Obras  Mai.,    ....  203 

18.  19.  Orthostylus  faunus  Brod.     Specimen,          .         .         .  203 

20.  Orthostylus  macrostoma  Pfr.     Conch.  Cab.,      .         .         .  208 

21.  Calocochlea  luengoi  Hid.     J.  de  C., 

22.  Orthostylus  onyx  Brod.     Obras  Mai.,       .         .         .         .199 

23.  Orthostylus  pithogaster  var.  bipartita.     Specimen,  .         .  201 

24.  Orthostylus  pithogaster  var.  philippinensis.      Obras  Mai.,  201 


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