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SMITHSONIAN INSTITUTION 


Contributions from the United States National Herbarium 
Volume 41: 1-255 


Catalogue of New World Grasses 
(Poaceae): II. Subfamily Chloridoideae 


by 
Paul M. Peterson 
Robert J. Soreng 
Gerrit Davidse 
Tarciso S. Filgueiras 
Fernando O. Zuloaga 
and 
Emmet J. Judziewicz 


Chief Editor: 
Robert J. Soreng 


Department of Systematic Biology - Botany 
National Museum of Natural History 
Washington, DC 
2001 


ABSTRACT 


Peterson, Paul M., Robert J. Soreng, Gerrit Davidse, Tarciso S. Filgueiras, Fernando O. Zuloaga, and 
Emmet J. Judziewicz. Chief Editor, Robert J. Soreng. Catalogue of New World Grasses: II. Subfamily 


Chloridoideae.-- Nomenclature, taxonomy, synonymy, original publications, type collections, secondary references using the 
accepted names, and distribution by country and Caribbean region within the New World are presented for taxa within the grass 
subfamily: Chloridoideae. Taxonomic synonymy is presented for all taxa at all ranks (other than ranks applying to cultivated 
taxa) known to occur natively or naturalized in the New World. In addition, data on introduced species are recorded. The work 
covers 3654 binomial and trinomial names, which are denoted as; 1 - accepted, 2 - tentatively accepted (bolded), synonyms 
(italicized), and status not yet determined. Accepted names are additionally marked as cultivated (45 species) or introduced (68 
species) to our region, Other works using the accepted names are cited. The number of accepted genera = 70 (62 native, 8 
introduced). The number of accepted or tentatively accepted species = 672 (604 native species); 148 taxa are further separated 
into subspecific rank. The status of 25 species is not yet determined. A classification of subfamily Chloridoideae is presented 
and a new combination, Muhlenbergia sericea, is made. 

KEY WORDS: Grasses, Classification, Nomenclature, Taxonomy, Poaceae, Gramineae, Chloridoideae. 


DATE OF PUBLICATION: December 2001 


Cover Design and Illustrations by Alice Tangerini: front Muhlenbergia jaime-hintonii P. M. Peterson & 
Valdés-Reyna (Valdés-Reyna & Carranza Pérez 2560); back Eragrostis pastoensis (Kunth) Trin. 
(Peterson et al. 8910 & 9268). 


Contributions from the United States National Herbarium (ISSN 0097-1618) Department of Systematic 
Biology - Botany, National Museum of Natural History, Smithsonian Institution, Washington, DC, 20560- 
0166, USA. 


POSTMASTER: Send address changes to Contributions US National Herbarium, Department of 
Systematic Biology - Botany, National Museum of Natural History, Smithsonian Institution, Washington, 
DC, 20560-0166, USA. 


The paper used in this publication meets the minimum requirements of the American National Standard 
for Permanence of Paper for Printed Library Materials Z39.48--1984. 


Contributions from the U.S. National Herbarium was first published in 1890 by The United States 
Department of Agriculture. From July 1 1902 forward it was published as a Bulletin of the United States 
National Museum. The series was discontinued after volume 38, 1974, and has been revived with volume 
39, as a venue for publishing longer taxonomic papers, checklists, floras, and monographs, produced by 
the staff and associates at the U.S. National Herbarium. It is externally peer reviewed, and published at 
irregular intervals. Subscription and other correspondence should be addressed to CUSNH, Department 
of Systematic Biology - Botany, National Museum of Natural History, Smithsonian — Institution, 
Washington, DC, 20560-0166, USA e-mail: CUSNH@nmnh.si.edu. The present issue is available for 
free while supplies last. 


CONTENTS 


INTRODUCTION 
MATERIALS AND METHODS 
GUIDE TO USING THE CATALOGUE OF NEW WORLD GRASSES 
Nomenclature 
Taxonomy 
Synonymy 
Levels of Acceptance and Status 
Distribution 
Abbreviations used for distribution by country or Caribbean Region 
Abbreviations used in subheadings 
Abbreviations used in NOTES 
Abbreviations used for Types 
Consulted Literature and Also Accepted By 
A CLASSIFICATION OF NEW WORLD POACEAE SUBFAMILY Chloridoideae 
ACKNOWLEDGMENTS 
CATALOGUE OF NEW WORLD GRASSES 


Owommwowo MwAIANwAAYNANADAAAAAADAAADA UN 


Aegopogon 

Allolepis 

Bealia 

Blepharidachne 

Blepharoneuron 

Bouteloua 20 
Calamovilfa 35 
Chaboissaea 38 
Chloris 39 
Cladoraphis 55 
Cottea 56 
Crypsis 56 
Ctenium 57 
Cynodon 59 
Dactyloctenium 64 
Dasyochloa 65 
Dinebra 67 
Distichlis 70 
Eleusine 73 
Enneapogon 78 
Enteropogon 78 
Eragrostis 81 
Erioneuron 115 
Eustachys 117 
Fingerhuthia 121 
Gouinia 122 
Gymnopogon 124 
Hilaria 128 
Jouvea 129 
Leptochloa 130 


Leptothrium 137 


Lepturidium 
Lycurus 
Microchloa 
Monanthochloe 
Muhlenbergia 
Munroa 
Neesiochloa 
Neobouteloua 
Neostapfia 
Neyraudia 
Orcuttia 
Pappophorum 
Pereilema 
Pleuraphis 
Redfieldia 
Reederochloa 
Rheochloa 
Saugetia 
Schaffnerella 
Schedonnardus 
Scleropogon 
Sohnsia 
Spartina 
Sporobolus 
Steirachne 
Swallenia 
Tetrachne 
Tragus 
Trichloris 
Trichoneura 
Tridens 
Triplasis 
Tripogon 
Triraphis 
Tuctoria 
Uniola 
Vaseyochloa 
Willkommia 
Zoysia 


BIBLIOGRAPHY 


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Catalogue of New World Grasses: II. 5 


Catalogue of New World Grasses (Poaceae): 
II. Subfamily Chloridoideae. 


Paul M. Peterson, Robert J. Soreng, Gerrit Davidse, Tarciso S. Filgueiras, 
Fernando O. Zuloaga, and Emmet J. Judziewicz.' 
Chief Editor, Robert J. Soreng 


INTRODUCTION 


The Catalogue of New World Grasses 
(CNWG) project is an in-progress effort led by 
the authors to present the current nomenclature, 
taxonomy, types, and distribution of grasses for 
the region. Contributions of treatments of genera 
were/are solicited from the agrostological 
community. For the present volume contributors 
were: Sonja Boechat (Eragrostis and Sporobolus), 
Yolanda Herrera-Arrieta (Bouteloua), Khidir Hilu 
(Eleusine), Neil Snow (Leptochloa), Jesus Valdes- 
Reyna (Blepharidachne, Dasyochloa, Erioneuron, 
Munroa, and Tridens), and Jeff Veldkamp 
(Sporobolus). Paul Peterson contributed all other 
genera. 


MATERIALS AND METHODS 


The data were recorded in and downloaded 
from TROPICOS (Fica and Beiser, 1995), the 
Missouri Botanical Garden’s (MBG) taxonomic 
database. TROPICOS is a PICK based database 
program developed by Bob Magill, Director of 
Research MBG. All data were entered into 
TROPICOS from terminals at the Missouri 
Botanical Garden, or by using Telnet access 
from remote locations (especially Washington, 
DC), or by downloading data from a PC version 
of TROPICOS. The complete and continuously 
updated data are accessible online through 


W°TROPICOS at: 
http://mobot.mobot.org/W3T/Search/nweg.html 


Names of taxa, their original publication 
citations, authorities, and type  protologues, 
primarily taken from the /ndex to Grass Species 
(Chase and Niles, 1962), were recorded in the 
Nomenclature and Types files of TROPICOS. 
Direct links were established between basionyms, 
combinations, new names, and  autonyms. 
Additional names and types were recorded from 
Kew Index supplements from 1950 to present 
(Jackson, 1893+), Gray Herbarium Index (Gray 
Herbarium Staff, 1903+), and other primary and 
secondary sources (see Bibliography). Type 
specimen data were augmented by the records 
from the Smithsonian Type Register. 

Synonymy and distribution according to 
authors of various floras, monographs, revisions, 
and checklists were recorded in the Synonymy 
and Distribution files which are linked to the 
Nomenclatural files. For CNWG, the contributors 
of each genus were provided these nomenclatural, 
taxonomic, and distributional data and asked to 
review the data, add missing names, and assign 
to each name a level of acceptance or indicate an 
accepted name for synonyms. In addition, the 
contributors were asked to accept, reject, or add 
to the distributions provided by the collective 
sources. 

Synonymy and distribution recognized by the 
CNWG contributors for each genus were then 
recorded in the synonymy and distribution files. 
Additional data were then linked to the names in 


| Paul M. Peterson, Smithsonian Institution, Washington, DC , USA. Robert J. Soreng, Smithsonian Institution, Washington, 
DC, USA. Gerrit Davidse, Missouri Botanical Garden, St. Louis, MO, USA. Tarciso S. Filgueiras, UPIS-Faculdades 
Integradas, Brasilia, Brazil. Fernando O. Zuloaga, Darwinion, San Isidro, Argentina. Emmet J. Judziewicz, University of 


Wisconsin, Stevens Point, WI, USA. 


6 Catalogue of New World Grasses: II. 


a separate Flora file, including the status of the 
name, the level of acceptance, the generic 
reference from synonymy and_ distribution, 
Special notes and_ references, and_ the 
suprageneric classification. 

A list, with formatting codes, of all names 

applicable to the four subfamilies (using our 
classification; see below) was generated from the 
database. Camera-ready copy was then 
produced through a macro written by Peter 
Jorgensen for Microsoft Word 97. 
Standards set forth for authors of plant names 
(Brummitt & Powell 1992), literature citations 
(Stafleu & Cowan 1983+; Stafleu & Mennega 
1992+; Lawrence et al. 1968; Birdson & Smith 
1991), and herbarium acronyms (Holmgren et al. 
1990), were followed. 


GUIDE TO USING THE 
CATALOGUE OF NEW WORLD 
GRASSES 


Nomenclature 

Taxa of all ranks (excluding taxa treated 
under the /nternational Code of Nomenclature 
for Cultivated Plants) within Poaceae were 
included. 


Taxonomy 

Accepted and tentatively accepted names are 
in bold face. Names of unknown or of uncertain 
application, and synonyms are in italics. 


Synonymy 

We attempted to include all names published 
for native New World taxa and tried to include 
the better known synonyms for introduced taxa. 


Levels of Acceptance and Status 

Each name not placed in synonymy was 
assigned a level of acceptance or confidence in 
the taxonomy. Names of all bolded taxa are 
validly published and legitimate, unless otherwise 
noted. 


| = Accepted. 
2 = Tentatively accepted. 


Status of names refers to their recognition as 
either accepted or tentatively accepted, 
synonyms, and as native or introduced. 
Synonymy may or may not be indicated for 
invalid names depending on the state of 
knowledge concerning them. They are presented 
here so that users will not have to re-verify them 
each time they turn up in the literature. We 
recommend they not be repeated again 
elsewhere. Names that we know nothing about 
are included and are designated as = Status not 
yet determined. 


Cultivated = Cultivated (native or naturalized) 
Introduced = Introduced to the New World. 


Distribution 

The geographical range covered is North 
America, Mesoamerica, and South America, 
from Greenland to Tierra del Fuego. As this is a 
large-scale project, floristically related islands 
are included with adjacent countries (e.g., 
Galapagos Islands are included with Ecuador, 
St.-Pierre et Miquelon are included with Canada, 
the Falkland or Malvinas islands are included 
with Argentina). The Caribbean island nations 
are treated collectively. 


Abbreviations for distributions by 
country or Caribbean Region 

Arge = Argentina (including Falkland Islands). 
Beli = Belize. 

Boli = Bolivia. 

Braz = Brazil. 

Cana = Canada. 

Cari = Caribbean (including all island nations). 
Chil = Chile (including Juan Fernandez Islands). 
Colo= Colombia 

CoRi = Costa Rica. 

Guya = Guyana. 

Ecua = Ecuador (including Galapagos Islands). 
EISa = El Salvador. 

FrGu = French Guiana. 

Guat = Guatamala. 

Hond = Honduras. 

Mexi = Mexico. 

Nica = Nicaragua. 

Pana = Panama. 

Para = Paraguay. 

Peru = Peru. 


Catalogue of New World Grasses: II. 7 


Suri = Surinam. 

USA = United States of America. 
Urug = Uruguay. 

Vene = Venezuela. 


Abbreviations used in subheadings 
ALSO ACCPTD. BY = Also Accepted by. 
CNSLTD. LIT. = Consulted literature. 
DST. = Distribution. 

E. NAME = Earlier name. 

LVL. ACCPTC. = Level of Acceptance. 
NHT = Next Higher Taxon (in rank). 

SPP. = Number of accepted species. 
SUBSP. = Number of accepted infraspecies. 
SYN. = Synonym(s). 

TRTD. BY = Treated by. 


Abbreviations and codes used in NOTES 


nom, nov. = nomen novum, new name or 
avowed substitute name 

nom. nud. = nomen nudum, naked name. 

nom. inval. = nomen invalidum, invalid name. 

nom. illeg. = nomen illegitimum, illegitimate 
name. 

nom, illeg. superfl. = nomen illegitimum et 
superfluum, superfluous illegitimate name. 

hom. illeg. = homonymum _ illegitimum, 
illegitimate name, a later homonym. 

isonym = a subsequent unnecessary publication 
of a combination already validly published. 

as syn. = in synonymo, an invalid name originally 
published as a synonym of another, earlier 
validly published name. 

Basionym: A name on which combinations or 
transfers of rank, if any, are based. 

Blocking Name: A name (most often an earlier 
homonym) preventing the adoption of 
another name. 

Replaced Name: A name (later homonym) which 
a nomen novum replaced. 

Correct Name: A nomenclaturally valid and 
legitimate name that could have been 
adopted for an illegitimate name. 


Abbreviations used for Types 

T-Protol. = type protologue (type specimen data 
cited in the original publication of a name). 

T-Specim. = type specimen (from specimen 
labels). 


Kinds of Types: ET = epitype, HT = holotype, 
IT = isotype, ILT = isolectotype, INT = 


isoneotype, IST =  isosyntype, LT = 
lectotype, NT = neotype, OM = original 
material, ST = syntype, T = = any 
uncharacterized type material, fragm. = 


fragment of another type (mainly used for 
very incomplete specimens taken off other 
original specimens). 

TRIN = C. B. Trinius herbarium LE, numbers 
corresponding to IDC microfiche specimen 
numbers (Soreng et al. 1995). 


Types are cited twice if entered into the 
database in two ways, once under T-Protol., and 
again under T-Specim. This is because the data 
sometimes differ between what is stated in the 
published protologue and what is stated on 
specimen labels. 


Consulted Literature and Also Accepted 


By 


Primary and secondary literature sources are 
cited as CNSLTD. LIT. under entries for genera. 
Other publications accepting the same taxonomy 
as the Catalogue of New World Grasses are 
listed under the heading: ALSO ACCPTD. BY. 


A CLASSIFICATION OF NEW 
WORLD POACEAE, SUBFAMILY: 
CHLORIDOIDEAE 


Poaceae subfamilies, tribes, subtribes, and 
genera are listed below for grasses occurring in 
the New World (“i”= genera with no native 
species in the New World; genera 
occurring in the New World principally in 
cultivation). Alignments reflect recent 
morphological and molecular — syntheses 
(Columbus et al. 1998; Duvall et al. 1994; Hilu 
and Alice 2000, 2001; Ortiz and Culham 2000; 
Peterson 2000; Peterson et al. 1995, 1997; Van 
den Borre and Watson 1997, 2000), and 
interpretations by the present authors. 


66 99 
Cc = 


subfam. Chloridoideae (SYN.: Eragrostoideae, 
Pappophoroideae) 

tribe Cynodonteae (SYN.: Chlorideae, 
Eragrostideae) 


8 Catalogue of New World Grasses: II. 


Incertae Sedis: Blepharidachne, Chloris, 
Cladoraphis i, Ctenium, Cynodon, Dinebra i, 
Enteropogon, Eragrostis, Eustachys, 
Gouinia, Gymnopogon, Hilaria, Leptochloa, 
Lepturidium, Microchloa, Neesiochloa, 
Neobouteloua, Neyraudia i, Pleuraphis, 
Rheochloa, Saugetia, Scleropogon, Sohnsia, 
Spartina, Steirachne, Trichloris, 
Trichoneura, Tridens, Triplasis, Tripogon, 
Vaseyochloa, Willkommia 

subtribe Boutelouinae 
Bouteloua 

subtribe Eleusininae 
Dactyloctenium i, Eleusine, Triraphis 

subtribe Muhlenbergiinae 
Aegopogon, Bealia, Blepharoneuron, 
Chaboissaea, Lycurus, Muhlenbergia, 
Pereilema, Redfieldia, Schaffnerella, 
Schedonnardus 

subtribe Monanthochloinae 


Allolepis, Distichlis, Jouvea, Monanthochloe, 


Reederochloa, Swallenia 
subtribe Munroinae 

Dasyochloa, Erioneuron, Munroa 
subtribe Sporobolinae 

Calamovilfa, Crypsis i, Sporobolus 
subtribe Uniolinae 

Fingerhuthia i, Tetrachne, Uniola 
subtribe Zoysiinae 

Leptothrium, Tragus i, Zoysia i 

tribe Orcuttieae 


Neostapfia, Orcuttia, Tuctoria 
tribe Pappophoreae 
Cottea, Enneapogon, Pappophorum 


ACKNOWLEDGEMENTS 


Many thanks are extended to the contributing 
authors Sonja Boechat, Yolanda Herrera-Arrieta, 
Khidir Hilu, Neil Snow, Jesus Valdes-Reyna, 
and Jeff Veldkamp; Chris Tuccinardi and Dawn 
Arculus (formerly) of US-type Register for 
providing the U.S. National Herbarium grass 
type records; Smithsonian Institution 
Fellowships and Grants, Seidell Foundation, and 
James Smithson Society for providing funding 
for aspects of the databasing; Missouri Botanical 
Garden Staff and especially Bob Magill, Director 
of Research MBG, for TROPICOS support, 
programming, and web site development and 
production, James Reveal for work on 
suprageneric names, and Peter Jorgensen for 
development of macros for downloading the files 
in publication format; Jeany Davidse, Victoria 
Batista, Donna Herendeen, Jill Marsden, Susan 
Pennington, Cirri Rohrbach, and volunteers John 
S. Skinner, and Virginia G. Laschober for data 
entry; Dan H. Nicolson for extensive help in 
nomenclatural matters; and K. N. Gandhi and 
John H. Wiersema for their careful reviews. 


Catalogue of New World Grasses: II. 9 


CATALOGUE OF NEW WORLD GRASSES 


Acamptoclados Nash, Fl. S.E. U.S. 139, 1327. 1903. TYPE: 
Acamptoclados sessilispicus (Buckley) Nash. 
= Eragrostis Wolf. 


Acamptoclados sessilispicus (Buckley) Nash, Fl. S.E. U.S. 
140. 1903. Basionym: Eragrostis sessilispica Buckley. 
= Eragrostis sessilispica Buckley, 


Achnatherum soboliferum (Muhl. ex Willd.) P. Beauv., Ess. 
Agrostogr. 20, 146, 148. 1812. Basionym: Agrostis 
sobolifera Muhl. ex Willd. 

= Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. 


Acroxis Steud., Nomencl. Bot. (ed. 2) 1: 22. 1840. 
NOTE: pro. syn. sub Muhlenbergia. 
= Muhlenbergia Schreb. 


Actinochloa Willd. ex Roem. & Schult., Syst. Veg. 2: 22, 
417. 1817. 
NOTE: nom. illeg. superfl. 
= Bouteloua Lag. 


Actinochloa barbata (Lag.) Roem. & Schult., Syst. Veg. 2: 
420. 1817. Basionym: Bouteloua barbata Lag. 
= Bouteloua barbata Lag. 


Actinochloa bromoides (Lag.) Roem. & Schult., Syst. Veg. 2: 
420. 1817. Basionym: Bouteloua bromoides Lag. 
= Bouteloua repens (Kunth) Scribn. 


Actinochloa gracilis (Kunth) Willd. ex Roem. & Schult., 
Syst. Veg. 2: 418. 1817. Basionym: Chondrosum 
gracile Kunth. 

= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Actinochloa hirsuta (Lag.) Roem. & Schult., Syst. Veg. 2: 
419. 1817. Basionym: Bouteloua hirsuta Lag. 
= Bouteloua hirsuta Lag. 


Actinochloa humilis (P. Beauv. ex Kunth) Willd. ex Roem. & 
Schult., Syst. Veg. 2: 417. 1817. Basionym: 
Chondrosum humile P. Beauv. ex Kunth. 

= Bouteloua simplex Lag. 


Actinochloa juncifolia (Lag.) Roem. & Schult., Syst. Veg 2: 
420. 1817. Basionym: Bouteloua juncifolia Lag. 
= Bouteloua juncifolia Lag. 


Actinochloa ovata (Lag.) Roem. & Schult., Syst. Veg. 2: 420. 
1817. Basionym: Bouteloua ovata Lag. 
= Bouteloua chondrosioides (Kunth) Benth. ex S. Watson. 


Actinochloa procumbens (P. Durand) Roem. & Schult., Syst. 
Veg. 2: 417. 1817. Basionym: Chloris procumbens P. 
Durand. 

= Bouteloua simplex Lag. 


Actinochloa prostrata (hort. ex Lag.) Roem. & Schult., Syst. 
Veg. 2: 419. 1817. Basionym: Bouteloua prostrata 
hort. ex Lag. 

= Bouteloua simplex Lag. 


Actinochloa scorpioides (Lag.) Roem. & Schult., Syst. Veg. 
2: 420. 1817. Basionym: Bouteloua scorpioides Lag. 
= Bouteloua scorpioides Lag. 


Actinochloa simplex (Lag.) Roem. & Schult., Syst. Veg. 2: 
418. 1817. Basionym: Bouteloua simplex Lag. 
= Bouteloua simplex Lag. 


Actinochloa tenuis (P. Beauv. ex Kunth) Willd. ex Roem. & 
Schult., Syst. Veg. 2: 418. 1817. Basionym: 
Chondrosum tenue P. Beauv. ex Kunth. 

= Bouteloua simplex Lag. 


Aegilops aromatica Walter, Fl. Carol. 249. 1788. T-Protol.: 
USA: South Carolina. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Aegilops saccharinum Walter, Fl. Carol. 249. 1788. T- 
Protol.: USA: South Carolina. 
= Dactyloctenium aegyptium (L.) Willd. 


Aegopogon Humb. & Bonpl. ex Willd., Sp. Pl. 4(2): 
899. 1805 [1806]. TYPE: Aegopogon cenchroides 
Humb. & Bonpl. ex Willd. 

TRTD. BY: Peterson, P. M. 

NHT: Muhlenbergiinae. NO. ACCPTD. SPP.: 4. 

CNSLTD. LIT.: Beetle, A. A., (1948). 

SYN.: Atherophora Steud.; Hymenothecium 
Schellingia Steud. 


Lag.; 


Aegopogon argentinus Mez, Repert. Spec. Nov. Regni Veg. 
17(8-12): 145. 1921. T-Protol.: Argentina: Prov. Salta: 
Sierra Nevada del Castillo, Lorentz & Hieronymus. T- 
Specim.: 1 of 1. Argentina: Salta, no date, P.G. 
Lorentz & G. Hieronymus 182 [HT: B; IT: US-75037 
(fragm. ex B)]. 
= Aegopogon bryophilus Ddll. 


Aegopogon breviglumis (Scribn.) Nash, N. Amer. FI. 17(2): 
139. 1912. Basionym: Aegopogon geminiflorus var. 
breviglumis Scribn. 

= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Aegopogon bryophilus Doll, Fl. Bras. 2(3): 239. 1880. T- 
Protol.: Brasilia: Habitat prope Rio de Janeiro, Glaziou 
11661. T-Specim.: 1 of 1. Brazil: Minas Gerais, 10 
Apr 1879, 4.F.M. Glaziou 11661] [IT: US-1280026]. 

DST.: Arge, Boli, Braz, Colo, Ecua, Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1948); Brako, L. & J. 
L. Zarucchi (1993); Doll, J. C. (1880); Foster, R. C. 
(1958); Hitchcock, A. S. (1927); Jorgensen, P. M. & 
S. Leén-Yanez (eds.) (1999); Macbride, J. F. (1936); 
Renvoize, S. A. (1998); Tovar, O. (1993); Zuloaga, 
F. O., E. G. Nicora, Z. E. Rugolo de Agrasar, O. 
Morrone, J. Pensiero & A. M. Cialdella (1994). 
Aegopogon argentinus Mez; Aegopogon fiebrigii 
Mez; Aegopogon geminiflorus var. muticus Pilg. 


SYN.: 


10 Catalogue of New World Grasses: II. 


Aegopogon cenchroides Humb. & Bonpl. ex Willd., Sp. PI. 
4(2): 899. 1806. T-Protol.: Habitat in Cumana. T- 
Specim.: | of 1. Venezuela: Sucre: Cumana, Humboldt 
& Bonpland s.n. [3002] (HT: B-W; IT: P, US-75957 
(fragm. ex P)]. 

DST.: Boli, Braz, Colo, CoRi, Ecua, ElSa, Guat, Guya, 

Hond, Mexi, Pana, Peru, USA, Vene. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1948); Beetle, A. A. 
(1983); Berendsohn, W. G. & A. E. Araniva de 
Gonzalez (1991); Brako, L. & J. L. Zarucchi (1993); 
Breedlove, D. E. (1986); Davidse, G., M. Sousa S. & 
A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Foster, R. C. (1958); Hitchcock, A. S. (1927): 
Jorgensen, P.M. & C. Ulloa U. (1994); Jorgensen, P. 
M. & S. Leén-Yanez (eds.) (1999); Judziewicz, E. J. 
(1990); Molina R., A. (1975); Pohl, R. W. (1980); 
Pohl, R. W. (1994); Renvoize, S. A. (1998); Shreve, 
F. & |. L. Wiggins (1964); Swallen, J. R. (1955); 
Tovar, O. (1993). 

Aegopogon breviglumis (Scribn.) Nash; Aegopogon 
cenchroides var. breviglumis (Scribn.) Beetle; 
Aegopogon cenchroides var. geminiflorus (Kunth) 
Griseb.; Aegopogon cenchroides var. multisetus E. 
Fourn.; Aegopogon cenchroides var. trisetus (Lag.) 
E. Fourn.; Aegopogon geminiflorus var. breviglumis 
Scribn.; | Aegopogon — guatemalensis —_Gand.; 
Aegopogon multisetus Steud.; Aegopogon pusillus P. 
Beauv.; Aegopogon quinquesetus (Lag.) Roem. & 
Schult., Aegopogon  setifer Nees; Aegopogon 
submuticus Rupr.; Aegopogon trisetus (Lag.) Roem. 
& Schult; Hymenothecium quinquesetum Lag.; 
Hymenothecium trisetum Lag. 


SYN.: 


Aegopogon cenchroides var. breviglumis (Scribn.) Beetle, 
Univ. Wyoming Publ. 8(2): 23. 1948. Basionym: 
Aegopogon geminiflorus var. breviglumis Scribn. 

= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Aegopogon cenchroides var. geminiflorus (Kunth) Griseb., 
Abh. Konigl. Ges. Wiss. Gottingen 24: 301. 1879. 
Basionym: Aegopogon geminiflorus Kunth. 

= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Aegopogon cenchroides var. multisetus E. Fourn., Mexic. Pl. 
2: 72. 1886. T-Specim.: | of 2. Mexico: Moran. in 
rupibus, 1840, H. Galeotti 5808 [HT: BR; IT: P, US- 
75958 (fragm. ex P)]. 2 of 2. Mexico: Toluca, 
Berlandier 1181 (ST: LE, P]. 

= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Aegopogon cenchroides vat. trisetus (Lag.) E. Fourn., Mexic. 
Pl. 2: 72. 1886. Basionym: Hymenothecium trisetum 
Lag. ‘ 
= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Aegopogon fiebrigii Mez, Repert. Spec. Nov. Regni Veg. 


17(8-12): 145. 1921. T-Protol.: Bolivia: Camacho, 


Fiebrig. T-Specim.: | of 1. Bolivia: Camacho, no date, 
K. Fiebrig 2865 |HT: B; IT:US-75308 (fragm. ex B)]. 
= Aegopogon bryophilus Doll. 


Aegopogon geminiflorus var. abortivus E. Fourn., Mexic. Pl. 
2: 71. 1886. T-Specim.: | of 5. Mexico: Veracruz: 
Orizaba, 1864, Weber & Thomas s.n. [ST: P; IST: US- 
75961 (fragm. ex P)]. 2 of 5. Mexico, no date, M 
Bourgeau 750 [IST: US-75303 (fragm.)]. 3 of 5. 


Mexico: Veracruz: Orizaba, Botteri 4] [ST: P]. 4 of 5. 
Mexico: Orizaba, M. Botteri & A.L. Sumicrast 84 [ST: 
P|. 5 of 5. Mexico, Schaffner 7 |ST: P}. 

= Aegopogon tenellus (DC.) Trin. 


Aegopogon geminiflorus var. breviglumis Scribn., Zoe 4(4): 
386. 1894. T-Protol.: Mexico: Saucito, 14 Oct 1893, 
Brandegee 69 [HT: US]. T-Specim.: | of 1. Mexico: 
Baja California: Saucito, 14 Oct 1893, 7.8. Brandegee 
s.n. [HT: US-75306]. 

= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Aegopogon geminiflorus var. muticus Pilg., Bot. Jahrb. Syst. 
27: 25. 1899. T-Protol.: Bolivia: crescit ad vicum 
Sorata, flor. coll. mens. Mai, 1892, Bang /307. T- 
Specim.: | of I. Bolivia: La Paz, May 1892,M. Bang 
1307 [IT: US-926725]. 

= Aegopogon bryophilus Déll. 


Aegopogon geminiflorus subvar. purpureus Griseb. ex E. 
Fourn., Mexic. Pl. 2: 71. 1886. T-Specim.: | of 2. 
Mexico: Tacubaya, Schaffner 168a (pl. ed. Hohen.) 
[ST]. 2 of 2. Panama, Seemann (Bot. Herald 225) [ST: 
K]. 

= Aegopogon tenellus (DC.) Trin. 


Aegopogon geminiflorus var. unisetus (Lag.) E. Fourn., 
Mexic. Pl. 2: 71. 1886. Basionym: Hymenothecium 
unisetum Lag. 

= Aegopogon tenellus (DC.) Trin. 


Aegopogon gracilis Vasey, Bull. Torrey Bot. Club 13(12): 
230. 1886. T-Protol.: Mexico: Southwestern 
Chihuahua, 1885, Dr. Ed. Palmer s.n. T-Specim.: | of 
1. Mexico: Chihuahua, Aug 1885-Nov 1885, E. 
Palmer 28 (HT: US-75305]. 

= Aegopogon tenellus (DC.) Trin. 


Aegopogon guatemalensis Gand., Bull. Soc. Bot. France 
66(7): 298. 1919 [1920]. T-Protol.: Habitat Guatemala, 
ad Alta Verapaz, Tuerckheim s.n. T-Specim.: | of 1. 
Guatemala: Alta Verapaz, 1901, H. von Tuerckheim 
s.n. [HT: LY; IT: US-75304 (fragm.)]. 
= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Aegopogon imperfectus Nash, N. Amer. Fl. 17(2): 138. 1912. 
T-Protol.: Collected on cool mossy ledges, Arroyo 
Aucho, Sierra Madre, Chihuahua, Mexico, 15 Oct 
1887, C.G. Pringle 1408 [HT: NY]. T-Specim.: | of 1. 
Mexico: Chihuahua, 15 Oct 1887,C.G. Pringle 1408 
[HT: NY; IT: US-740860]. 

= Aegopogon tenellus (DC.) Trin. 


Aegopogon multisetus Steud., Syn. Pl. Glumac. 1: 146. 1854. 
T-Specim.: | of 1. Venezuela, no date, NV. Funck & 
L.J. Schlim 827 [HT: P; IT: US-75960 (fragm. ex P)], 
Locality published as Mexico. 
= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Aegopogon pusillus P. Beauv., Ess. Agrostogr. 122, t. 22, f. 
4.1812. T-Protol.: Colombia. 
= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Aegopogon quinquesetus (Lag.) Roem. & Schult., Syst. Veg. 
1: 805. 1817. Basionym: = Hymenothecium 
quinquesetum Lag. 

= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Aegopogon rigidisetus Steud., Syn. Pl. Glumac. 1: 146. 1854. 
T-Protol.: USA: Texas, Drummond s.n. T-Specim.: | 


Catalogue of New World Grasses: II. 1] 


of 1. USA: Texas, 1856, Drummond 374 (HT: P; IT: 
US-879824 (fragm. ex P), US-1063200 (fragm. ex E)]. 
= Bouteloua rigidiseta (Steud.) Hitche. 


Aegopogon setifer Nees, Linnaea 19(6): 691. 1847. T- 
Specim.: | of 1. Mexico, no date, A. Aschenborn 132 
[HT: B; IT; US-75953 (fragm. ex B)]. 
= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Aegopogon solisii GA. Levin, Mem. San Diego Soc. Nat. 
Hist. 16: 61. 1989. T-Protol.: Mexico: Isla Socorro, E 
slope at 980 m near summit of Cerro Evermann, near 
18.46N 110.57W, 4 Apr 1981, Moran 29504 [HT: SD; 
IT: CHAPA, MO, US]. T-Specim.: | of 1. Mexico: 
Socorro I., 4 Apr 1981, R. Moran 29504 [IT: US- 
3170268]. 

DST.: Mexi. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Levin, G. A. & 
R. Moran (1989). 


Aegopogon submuticus Rupr., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1): 25. 1840. T-Specim.: | of 1. Peru, Haenke s.n. 
[T: PR]. 

= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Aegopogon tenellus (DC.) Trin., Gram. Unifl. Sesquifl. 164. 
1824. Basionym: Lamarckia tenella DC. 

DST.: CoRi, ElSa, Guat, Hond, Mexi, Pana, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1948); Berendsohn, 
W. G. & A. E. Araniva de Gonzalez (1991); 
Breedlove, D. E. (1986); Davidse, G., M. Sousa S. & 
A. O. Chater (editores generales) (1994); Gould, F. 
W. & R. Moran (1981); Hitchcock, A. S. (1951); 
Molina R., A. (1975); Pohl, R. W. (1980); Pohl, R. 
W. (1994); Swallen, J. R. (1955). 

Aegopogon geminiflorus subvar. purpureus Griseb. 
ex E. Fourn.; Aegopogon geminiflorus var. abortivus 
E. Fourn.; Aegopogon geminiflorus var. unisetus 
(Lag.) E. Fourn.; Aegopogon gracilis Vasey; 
Aegopogon imperfectus Nash; Aegopogon tenellus 
var. abortivus (E. Fourn.) Beetle; Aegopogon 
unisetus (Lag.) Roem. & Schult.; Chloris pedicellata 
Steud.; Chloris pedicellata Steud. ex E. Fourn.; 
Cynosurus tenellus Cav. ex DC.; Hymenothecium 
tenellum (Cav. ex DC.) Lag.; Hymenothecium 
unisetum Lag.; Lamarckia tenella DC.; Schellingia 
tenera Steud. 


SYN:: 


Aegopogon tenellus var. abortivus (E. Fourn.) Beetle, Univ. 
Wyoming Publ. 8(2): 19. 1948. Basionym:Aegopogon 
cenchroides var. abortivus E. Fourn. 

= Aegopogon tenellus (DC.) Trin. 


Aegopogon trisetus (Lag.) Roem. & Schult., Syst. Veg. 2: 
805. 1817. Basionym: Hymenothecium trisetum Lag. 
= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Aegopogon unisetus (Lag.) Roem. & Schult., Syst. Veg. 2: 
805. 1817. Basionym: Hymenothecium unisetum Lag. 
= Aegopogon tenellus (DC.) Trin. 


Aeluropodeae trib. Nevski ex Bor, Oesterr. Bot. Z. 112: 184. 


1965. TYPE: Aeluropus Trin. 


Aeluropodinae subtrib. Jacq.-Fél., Gram. Afr. Trop. 1: 94. 
1962. 
NOTE: nom. inval., without Latin description. 


Agropyron geminatum (Spreng.) Schult., Mant. 3(Add. 1): 
655. 1827. Basionym: Triticum geminatum Spreng. 
= Eleusine tristachya (Lam.) Lam. 


Agropyron peruvianum (Lam.) Roem. & Schult., Syst. Veg. 
2: 761. 1817. Basionym: Triticum peruvianum Lam. 
= Distichlis spicata (L.) Greene. 


Agrosticula Raddi, Fl. Bras. Enum. Pl. 33. 1823. TYPE: 
Agrosticula muralis Raddi. 
= Sporobolus R. Br. 


Agrosticula brasiliensis (Raddi) Herter, Revista Sudamer. 
Bot. 6: 145. 1939. Basionym: Aira brasiliensis Raddi. 
NOTE: cf. Gray Herb Card Cat. 
= Eragrostis airoides Nees, 


Agrosticula muralis Raddi, Agrostogr. Bras. 33t. 1, f. 2. 
1823. T-Protol.: Brasilia: invenitur in veteribus muris 
Rio-Janeiro, Raddi s.n. T-Specim.: | of 1. Brazil: Rio 
de Janeiro, no date, G. Raddi s.n. [HT: PI; IT: US- 
76254 (fragm. ex PI)]. 

= Sporobolus tenuissimus (Mart. ex Schrank) Kuntze. 


Agrostis airoides Torr., Ann. Lyceum Nat. Hist. New York 
1(1): 151-152. 1824. T-Protol.: USA: On the branches 
of the Arkansas, near the Rocky Mts., Dr. E. James. T- 
Specim.: | of 1. USA: Colorado, no date, E. James s.n. 
[HT: NY-327612; IT: US-76255 (fragm. & photostat 
ex hb. Torrey)]. 
= Sporobolus airoides (Torr.) Torr. 


Agrostis apetala Bosc ex Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(3-4): 287. 1841. 

NOTE: nom. inval., as syn. of Muhlenbergia diffusa 
Schreber. 
= Muhlenbergia schreberi J.F. Gmel. 


Agrostis arachnoides Poir., Encycl. 1: 249. 1810. T-Protol.: 
USA: Carolina, Bosc. T-Specim.: | of 1. USA: South 
Carolina, no date, M4 Bosc s.n. [T: Fl, US-76290a 
(fragm. & photostat ex FI)]. 

= Muhlenbergia expansa (Poir.) Trin. 


Agrostis atrovirens (Kunth) Roem. & Schult., Syst. Veg. 2: 
361. 1817. Basionym: Vilfa atrovirens Kunth. 
= Sporobolus atrovirens (Kunth) Kunth. 


Agrostis barbata Pers., Syn. Pl. 1: 75. 1805. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Agrostis littoralis Lam. 
= Sporobolus virginicus (L.) Kunth. 


Agrostis barbata var. senegalensis Pers., Syn. Pl. 1: 76. 1805. 
T-Specim.: | of 1. Senegal, Roussillon s.n. [HT: P- 
LAM]. 

= Sporobolus virginicus (L.) Kunth. 


Agrostis bermudiana Tussac ex Kunth, Enum. Pl. 1: 259. 
1833. T-Protol.: Bermuda. 
NOTE: nom. inval., as syn. of Cynodon dactylon (L.) Pers. 
= Cynodon dactylon (L.) Pers. 


Agrostis boeckeleri Seub. ex Steud., Syn. Pl. Glumac. 1: 218. 
1854. T-Protol.: USA: Texas. 


12 Catalogue of New World Grasses: II. 


NOTE: nom. inval., as syn. of Gymnopogon distichophyllus 
Steudel. 

= Gymnopogon ambiguus (Michx.) Britton, Sterns & 
Poggenb. 


Agrostis brevifolia Nutt., Gen. N. Amer. Pl. 1: 44. 1818. T- 
Protol.: USA: near Fort Mandan: Missouri Rv., in 
sterile naked plains and arid agillaceous soils. 

= Muhlenbergia cuspidata (Torr. ex Hook.) Rydb. 


Agrostis caespitosa Torr., Ann. Lyceum Nat. Hist. New York 

1(1): 152-153. 1824. T-Protol.: USA: On the prairies 
of the Missouri and Platt rivers. T-Specim.: | of 1. 
USA: Prairies of the Missouri and Plate, £. James s.n. 
[T: NY-327621 (possible)]. 

NOTE: nom. illeg. 

E. NAME: Agrostis caespitosa (L.) Salisb. 

CORRECT NAME: Agrostis peckii House. 

= Muhlenbergia torreyi (Kunth) Hitche. ex Bush. 


Agrostis clandestina Biehler, Pl. Nov. Herb. Spreng. 8. 1807. 
T-Protol.: USA: Pennsylvania: eastern, Muhlenberg. 
T-Specim.: | of 1. USA: Pennsylvania, no date, Ex 
Herb. Muhlenberg 115 (HT: PH; IT: US-1535795 
(fragm. ex PH)]. 

= Sporobolus clandestinus (Biehler) Hitchc. 


Agrostis compressa Poir., Encycl., Suppl. 1: 258. 1810. T- 
Protol.: Caroline, M. Bosc. T-Specim.: | of 1. USA: 
Carolina, Bose s.n. [HT: P?; IT: P]. 

NOTE: hom. illeg. 
E. NAME: Agrostis compressa Willd. 
= Sporobolus indicus (L.) R. Br. 


Agrostis compressa Torr., Cat. Pl. New York 91. 1819. T- 
Protol.: USA: New Jersey: in swamps, Goldy s.n. T- 
Specim.: | of 2. USA: New Jersey: pine barrens, Oct. 
1817, J. Goldie s.n. [ST: NY-327622, NY-327623; 
IST: US-76265 (fragm. and photostat ex NY hb. 
Torrey)]. 2 of 2. USA: New Jersey: Quaker Bridge, 
Sep 1829 [ST: NY-327624 (possible)]. 

NOTE: hom. illeg. 

E. NAME: Agrostis compressa Willd. 
CORRECT NAME: Agrostis torreyana Schult. 
= Muhlenbergia torreyana (Schult.) Hitche. 


Agrostis congener Schumach., Beskr. Guin. Pl. 46. 1827. T- 
Protol.: Guinea, Thonning s.n. [C]. 
NOTE: also in’ Kongel. Danske  Vidensk. 
Naturvidensk. Math. Ath. 3: 66 (1828). 
= Sporobolus virginicus (L.) Kunth. 


Agrostis cruciata L., Syst. Nat. (ed. 10) 2: 873. 1759. T- 
Specim.: | of 2. Jamaica, no date, Swartz hb. [IT: US- 
80823 (fragm. ex hb. Swartz)], Basis of Chloris 
cruciata. 2 of 2. Jamaica, P. Browne s.n. [LT: LINN], 
LT designated by Hitchcock, Contr. U.S. Natl. Herb. 
12: 120 (1908). 

NOTE: also nom. nud. in Pl. Jam. Pugill. 2: 7, description 
cited as L. in Syst. Nat. ed 10, 2: 873 as Andropogon. 
= Chloris cruciata (L.) Sw. 


Selsk. 


Agrostis cryptandra Torr., Ann. Lyceum Nat. Hist. New 
York I(1): 151. 1824. T-Protol.: USA: Texas or 
Oklahoma: Canadian Rv., James s.n. T-Specim.: | of 
1. USA: Texas or Oklahoma, Canadian River, no date, 
E. James s.n. [T: NY-128298; IT: US-76269 (fragm. & 
photostat ex NY)]. 

= Sporobolus cryptandrus (Torr.) A. Gray. 


Agrostis debilis (Kunth) Spreng., Syst. Veg. 1: 262. 1825. 
Basionym: Podosemum debile Kunth. 
NOTE: hom. illeg. 
E. NAME: Agrostis debilis Poir. 
= Muhlenbergia microsperma (DC.) Kunth. 


Agrostis diandra Retz., Observ. Bot. 5: 19. 1789 [1788]. T- 
Specim.: | of |. India, Koenig s.n. [HT: LD; IT: BM 
(photo BRI), C, K (photo, BRI), L, NSW]. 

= Sporobolus diandrus (Retz.) P. Beauv. 


Agrostis diffusa Muhl., Descr. Gram. 64. 1817. T-Specim.: 1 
of 1. [North America], no. /02 [HT: PH; IT: US- 
1535800 (ex PH)]. 

NOTE: hom. illeg. 

E. NAME: Agrostis diffusa Host. 

CORRECT NAME: Agrostis sylvatica Torr. 

= Muhlenbergia sylvatica (Torr.) Torr. ex A. Gray. 


Agrostis distichophylla Phil., Fl. Atacam. 54. 1860. T-Protol.: 
Chile: prope Tilopozo ad marginem paludis. T- 
Specim.: | of 1. Chile: prope Tilopozo ad marginem 
paludos illius ingentis salsae fere sicae, 23.20 lat., 
7000 ft [T]. 

NOTE: hom. illeg. 

E. NAME: Agrostis distichophylla Roem. & Schult. 

CORRECT NAME: Agrostis eremophila Speg. 

= Muhlenbergia asperifolia (Nees & Meyen ex Trin.) 
Parodi. 


Agrostis domingensis (Trin.) Schult., Mant. 3(Add. 1): 570. 
1827. Basionym: Vilfa domingensis Trin. 
= Sporobolus domingensis (Trin.) Kunth. 


Agrostis elongata Lam., Tabl. Encycl. 1: 162. 1791. T- 
Protol.: Ex Amer. Merid. 
NOTE: nom. illeg. superfl., based on same type as A. indica 
L. 
CORRECT NAME: Agrostis indica L. 


= Sporobolus indicus (L.) R. Br. 


Agrostis elongata var. flaccida Roth, Syst. Veg. 2: 368. 1817. 
T-Specim.: | of 2. India, Roth 80B [ST: BM (photo, 
BRI); IST: LE-TRIN-1699.01 (photo, L, BRD]. 2 of 2. 
India, R. Wight 30 [ST: L]. 

= Sporobolus diandrus (Retz.) P. Beauv. 


Agrostis eremophila Speg., Anales Mus. Nac. Hist. Nat. 
Buenos Aires 7: 190. 1902. 

REPLACED NAME: Agrostis distichophylla Phil. 

BLOCKING NAME: Agrostis distichophylla Roem. & 
Schult. 

= Muhlenbergia asperifolia (Nees & Meyen ex Trin.) 
Parodi. 


Agrostis filiformis J. Kénig ex Kunth, Enum. Pl. 1: 261. 
1833. 
NOTE: nom. inval., as syn. of Cynodon elongatus Trin. 
E. NAME: Agrostis filiformis Vill. 
= Cynodon dactylon (L.) Pers. 


Agrostis frondosa Poir., Encycl. 1: 252. 1810. T-Protol.: 
Germany. 
= Muhlenbergia frondosa (Poir.) Fernald. 


Agrostis glauca (Nees) Steud., Syn. Pl. Glumac. 1: 175. 
1854. Basionym: Podosemum glaucum Nees. 
NOTE: hom. illeg. 
E. NAME: Agrostis glauca Muhl. 
= Muhlenbergia glauca (Nees) B.D. Jacks. 


Catalogue of New World Grasses: II. 13 


Agrostis heterolepis (A. Gray) Alph. Wood, Class-book Bot. 
598. 1847. Basionym: Vilfa heterolepis A. Gray. 
= Sporobolus heterolepis (A. Gray) A. Gray. 


Agrostis implicata (Kunth) Spreng., Syst. Veg. 1: 262. 1825. 
Basionym: Podosemum implicatum Kunth. 
= Muhlenbergia implicata (Kunth) Trin. 


Agrostis indica L., Sp. Pl. 63. 1753. T-Protol.: Habitat in 
India. T-Specim.: | of 1. USA: Virginia, Clayton 460B 
[LT: LINN-84.36; ILT: SI (fragm.)], LT designated by 
Hubbard, Agron. Lus. 28 (1967). 
= Sporobolus indicus (L.) R. Br. 


Agrostis involuta Muhl., Cat. Pl. Amer. Sept. 11. 1813. 
NOTE: nom. nud. 
= Sporobolus compositus (Poir.) Merr. 


Agrostis involuta Muhl., Descr. Gram. 72. 1817. T-Protol.: 
USA: Pennsylvania & New York: Susquehanna Rv. T- 
Specim.: | of 1. USA: Pennsylvania, no date, H. C. 
Muhlenberg 114 (HT: PH; IT: US-1535794 (fragm. ex 
PH)]. 

NOTE: hom. illeg. 

E. NAME: Agrostis involuta Poir. 
CORRECT NAME: Agrostis longifolia Torr. 
= Sporobolus compositus (Poir.) Merr. 


Agrostis juncea Lam., Encycl. 1: 60. 1783. T-Specim.: | of 1. 
India, Sonnerat s.n. [HT: P; IT: US (fragm.)]. 
= Sporobolus virginicus (L.) Kunth. 


Agrostis juncea Michx., Fl. Bor.-Amer. 1: 52. 1803. T- 
Protol.: Michaux s.n. [HT: P]. T-Specim.: | of 1. USA: 
in aridis Carolinae, Michaux s.n. [HT: P]. 
NOTE: hom. illeg. 
E. NAME: Agrostis juncea Lam. 
CORRECT NAME: Heleochloa juncea P. Beauv. 
= Sporobolus junceus (P. Beauv.) Kunth. 


Agrostis lanata Kunth, Nov. Gen. Sp. 1: 136. 1815 [1816]. T- 
Protol.; Mexico: 1300 & 1460 hexap., Humboldt & 
Bonpland s.n. T-Specim.: 1 of 1. Mexico: Guanuato: 
Villalpando, Humboldt & Bonpland s.n. (HT: P}. 

= Muhlenbergia pubescens (Kunth) Hitchce. 


Agrostis lateriflora Michx., Fl. Bor.-Amer. 1: 53. 1803. T- 
Protol.: Hab. praesertim in praecipitibus saxosis 
fluminis Mississipi et ripariiis Illinoensibus, Michaux 
s.n. [HT: P]. T-Specim.: | of 1. USA: in ripis sabulosis 
inundatis fluminis Misisipi ripariis Illinoensisbus, A. 
Michaux s.n. [HT: P; IT: US-76274 (fragm. & photo 
ex P)}. 

= Muhlenbergia frondosa (Poir.) Fernald. 


Agrostis littoralis Lam., Tabl. Encycl. 1: 161. 1791. T- 
Protol.; America merid., Communic. a D. Richard. T- 
Specim.: 1 of 2. South America, no date, herb. J.B.P. 
Lamarck s.n. [ST: US-76275 (fragm. ex P)]. 2 of 2. 
Antilles: Virgin Islands: bayis Trium  insularum 
Danarum, May 1787, L.C.M. Richard s.n, (LT: P; ILT: 
P-LAM], LT designated by Baaijens & Veldkamp, 
Blumea 35: 446 (1991). 

= Sporobolus virginicus (L.) Kunth. 


Agrostis matrella L., Mant. Pl. 185. 1771. T-Protol.: Habitat 
in Malabariae arenosis. T-Specim.: | of 1. Sri Lanka ?, 
Koenig 56 [LT: LINN-84.11; ILT: L], LT designated 
by Goudswaard, Blumea 26: ? (1980). 

= Zoysia matrella (L.) Merr. 


Agrostis mexicana L., Mant. Pl. 31. 1767. T-Protol.: Habitat 
in America calidiore, D. Jacquin, erroneously credited 
to tropical America [see Man. U.S. Grasses]. T- 
Specim.: 1 of 1. Cultivated in Hortus Upsaliensis, 
Anon. [T: LINN; IT: US-76278 (fragm. ex LINN)], see 
Hitchcock, Contr. U.S. Natl. Herb. 12: 119 (1908). 
= Muhlenbergia mexicana (L.) Trin. 


Agrostis microsperma Lag., Gen. Sp. Pl. 2. 1816. T-Protol.: 
D. Sessé. T-Specim.: 1 of 1. Mexico: plants grown at 
the Botanical Garden of Madrid from seeds collected 
by Sessé in Mexico, Sessé [HT: MA?]. 

= Muhlenbergia microsperma (DC.) Kunth. 


Agrostis minutissima Steud., Syn. Pl. Glumac. 1: 171. 1854. 
T-Protol.: USA: New Mexico: 1847, Fendler 986. T- 
Specim.: 1 of 1. USA: New Mexico, 1847, 4. Fendler 
986 [IT: MO, NY-327637, US-825378, US-997292]. 

= Muhlenbergia minutissima (Steud.) Swallen. 


Agrostis mucronata (Kunth) Spreng., Syst. Veg. 1: 262. 
1825. Basionym: Podosemum mucronatum Kunth. 


= Muhlenbergia mucronata (Kunth) Trin. 


Agrostis peruviana (P. Beauv.) Spreng., Syst. Veg. 1: 262. 
1825. Basionym: Clomena peruviana P. Beauv. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Agrostis plumosa Ten., Fl. Napol. 1: LIX. 1811. T-Protol.: 
Naples, presumably. 
= Sporobolus pyramidatus (Lam.) Hitchc. 


Agrostis pubescens Kunth, Nov. Gen. Sp. 1: 136. 1815 
[1816]. T-Protol.: Mexico: 1300 et 1460 hexap., 
Humboldt & Bonpland s.n. T-Specim.: | of 1. Mexico: 
Guanuato: Villalpando: Santa Rosa et Cerro del 
Cubilete, Humboldt & Bonpland s.n. [HT: P; IT: LE- 
TRIN-1557.01 b (fragm. [ hrbr. no. 1746] & fig.), US- 
76283]. 

= Muhlenbergia pubescens (Kunth) Hitchc. 


Agrostis pungens Muhl., Cat. Pl. Amer. Sept. 11. 1813. 
NOTE: nom. nud. 
= Sporobolus virginicus (L.) Kunth. 


Agrostis pungens Muhl., Descr. Gram. 72. 1817. T-Protol.: 
USA: [Pennsylvania? ]. 
NOTE: hom. illeg. 
E. NAME: Agrostis pungens Schreb. 
= Sporobolus virginicus (L.) Kunth. 


Agrostis purpurascens Sw., Prodr. 25. 1788. T-Protol.: 
Jamaica. T-Specim.: | of 1. Jamaica, Swartz s.n. [HT: 
S; IT: BM]. 
= Sporobolus purpurascens (Sw.) Ham. 


Agrostis pyramidalis Rich. ex Steud., Syn. Pl. Glumac. 1: 
153. 1854. T-Protol.: West Indies. 
NOTE: nom. inval., as syn. of Vilfa richardii Steudel. 
= Sporobolus pyramidatus (Lam.) Hitchce. 


Agrostis pyramidata Lam., Tabl. Encycl. 1: 161. 1791. T- 
Protol.. Amer. merid., Richard s.n. [HT: P]. T- 
Specim.: | of 2. Martinique, Richard. s.n. [T: LE- 
TRIN-1645.01 (fragm. & illust.)]. 2 of 2. South 
America, Richard s.n. [HT: P-LA]. 

= Sporobolus pyramidatus (Lam.) Hitche. 


Agrostis racemosa Michx., Fl. Bor.-Amer. 1: 53. 1803. T- 
Protol.: USA: Illinois: in ripas sabulosis inundatis 


14 Catalogue of New World Grasses: II. 


fluminis Mississipi, Michaux s.n. [HT: P]. T-Specim.: 
1 of 1. USA: in ripis sabulosis inundatis fluminis 
Mississippi, Michaux s.n. [HT: P; IT: US-76287 
(fragm. & photo ex P)]. 

= Muhlenbergia racemosa (Michx.) Britton, Sterns & 
Poggenb. 


Agrostis ramulosa (Kunth) Roem. & Schult., Syst. Veg. 2: 
361. 1817. Basionym: Vilfa ramulosa Kunth. 
= Muhlenbergia ramulosa (Kunth) Swallen. 


Agrostis sericea (Michx.) Elliott, Sketch Bot. S. Carolina 1: 
135. 1816. Basionym: Stipa sericea Michx. 
= Muhlenbergia sericea (Michx.) P.M. Peterson. 


Agrostis serotina Torr., Fl. N. Middle United States 1: 88. 

1823. T-Protol.: USA: New Jersey. T-Specim.: | of 1. 
USA: New Jersey: Pine barrens, 1803 [?],Nuttal s.n. 
"Agrostis serotina" map (IT: GH, LE-TRIN-1739.05, 
US-76292 (fragm. & photostat ex G)]. 

NOTE: hom. illeg. 

E. NAME: Agrostis serotina (L.) L. 

= Muhlenbergia uniflora (Muhl.) Fernald. 


Agrostis setosa (Kunth) Spreng., Syst. Veg. 1: 262. 1825. 
Basionym: Podosemum setosum Kunth. 
= Muhlenbergia microsperma (DC.) Kunth, 


Agrostis sobolifera Muhl, ex Willd., Enum. Pl. 95. 1809. T- 
Protol.: USA: Pennsylvania: sylvis. T-Specim.: | of 1. 
USA: Pennsylvania, no date, Ex Herb. Muhlenberg 
/11 [HT: PH; IT; US-1535796 (fragm. ex PH)]. 
= Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. 


Agrostis sylvatica Torr., Fl. N. Middle United States 1: 87. 
1823. T-Protol.; USA: Mountians of New Jersey. 
NOTE: hom. illeg. 
E. NAME: Agrostis sylvatica Huds. 
REPLACED NAME: Agrostis diffusa Muhl. 
BLOCKING NAME: Agrostis diffusa Host. 
= Muhlenbergia sylvatica (Torr.) Torr. ex A. Gray. 


Agrostis tenacissima L. f., Suppl. Pl. 107. 1781. T-Protol.: 
Hort. Upsal.: "India orientali", [error for America] see 
Bor, Kew Bull. 1957: 233. 
= Muhlenbergia mexicana (L.) Trin. 


Agrostis tenuiflora Willd., Sp. Pl. 1: 364. 1797. T-Protol.: 
North America [America boreali]. T-Specim.: | of 1. 
cult. in hort Beroliense, no date, Anonymous s.n. [T: 
IT: US-76295 (fragm. ex LE-TRIN)]. 
= Muhlenbergia tenuiflora (Willd.) Britton, Sterns & 
Poggenb. 


Agrostis tenuissima Spreng., Syst. Veg. 1: 258. 1825. T- 
Protol.: Ind. Occ. Amer. Austr., coll. not idicated. 
= Sporobolus indicus (L.) R. Br. 


Agrostis thyrsoides Bosc ex Trin., Mem. Acad. Imp. Sci. 
Saint-Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. 
Nat. 6,4(1-2): 76. 1840. 
NOTE: nom. inval., as syn. of Vilfa fulvescens Trin. 
= Sporobolus junceus (P. Beauv.) Kunth. 


Agrostis torreyana Schult., Mant. 2: 203. 1824. 
REPLACED NAME: Agrostis compressa Torr. 
BLOCKING NAME: Agrostis compressa Willd. 
= Muhlenbergia torreyana (Schult.) Hitchce. 


Agrostis torreyi Kunth, Enum. Pl. 1: 226. 1833. 


NOTE: nom. nov., also Revist. Gram. 1: Suppl. 1834. p. 
XVII. 

REPLACED NAME: Agrostis caespitosa Torr. 

BLOCKING NAME: Agrostis caespitosa (L.) Salisb. 

= Muhlenbergia torreyi (Kunth) Hitchc. ex Bush. 


Agrostis tremula Willd., Sp. Pl. 1: 372. 1797. T-Protol.: Hab. 
in Indiae lucis. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Agrostis juncea Lam. 
= Sporobolus virginicus (L.) Kunth. 


Agrostis trichopodes Elliott, Sketch Bot. S. Carolina |: 135, t, 
8, f. 1. 1816. T-Protol.: USA. T-Specim.: | of 1. USA: 
South Carolina, Charleston Co., Charleston Neck, no 
date, s.n. [T: IT: NY-22645 (possible)]. 
= Muhlenbergia expansa (Poir.) Trin. 


Agrostis virginica L., Sp. Pl. 1: 63. 1753. T-Protol.: Habitat 
in Virginia. T-Specim.: | of I. USA: habitat in 
Virginia, J. Clayton 507 [LT: LINN-84.30; ILT: BM, 
US-76294 (fragm. ex BM)], LT designated by 
Hitchcock, Contr. U.S. Natl. Herb. 12: 119 (1908). 

= Sporobolus virginicus (L.) Kunth. 


Agrostomia aristata Cerv., Naturaleza (Mexico City) 1: 345. 
1870. T-Protol.: Mexico: San Angel, San Agustin de 
las Cuevas, in agris. T-Specim.: | of 1. Mexico: 
Distrito Federal: San Angel, San Agustin de las 
Cuevas, in agris [T: BR?, F?, FI?, MEXU?, NY?, 
OXF? ]. 

= Chloris rufescens Lag. 


Agrostomia barbata Cerv., Naturaleza (Mexico City) 1: 346. 
1870. T-Protol.: Mexico: Cuernavaca. 
= Chloris virgata Sw. 


Agrostomia mutica Cerv., Naturaleza (Mexico City) 1: 345. 
1870. T-Protol.: Mexico: San Angel, San Agustin de 
las Cuevas: in agris. 

= Chloris submutica Kunth. 


Aira brasiliensis Raddi, Agrostogr. Bras. 36. 1823. T-Protol.: 
Brasilia: invenitur in veteribus muris prope Rio- 
janeiro. T-Specim.: | of 1. Brazil: Rio de Janeiro, 
Raddi s.n. (HT: FI. 

= Eragrostis airoides Nees. 


Aira capillacea Lam., Tabl. Encycl. 1: 177. 1791. T-Protol.: 
USA: E. Carolinia, D. Fraser. T-Specim.: 1 of 1. 
USA: "Carolina", no date, D. Fraser s.n. [T: US-76301 
(fragm.)]. 
= Eragrostis capillaris (L.) Nees. 


Aira congener Schumach., Beskr. Guin. Pl. 46. 1827. T- 
Specim.: | of 1. Ghana, Thonning s.n. [HT: C]. 
NOTE: also in Dansk. Vid. Selsk. Skrivt. 3: 66 (1828). 
= Sporobolus virginicus (L.) Kunth. 


Aira panicea Willd. ex Steud., Nomencl. Bot. 1: 45. 1840. 
NOTE: nom, inval., as syn. of Leptochloa filiformis (Lam.) 
P. Beauv. 
= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Aira purpurea Walter, Fl. Carol. 78. 1788. T-Protol.: USA: S. 
Carolina. T-Specim.: | of 1. USA [T: BM(not found)]. 
= Triplasis purpurea (Walter) Chapm. 


Catalogue of New World Grasses: II. 15 


Aira sabulonum Labill., Sert. Austro-Caledon. 16, t. 21. 
1824. T-Specim.: 1 of 1. New Caldonia: Balade, 
Labillardiere s.n. [HT: FI]. 

= Sporobolus virginicus (L.) Kunth. 


Aira sabulonum var. uniflora Balansa, Bull. Soc. Bot. France 
19: 318. 1872. T-Specim.: | of 1. New Caldonia, 
Labillardiére s.n. (HT: FY. 

= Sporobolus virginicus (L.) Kunth. 


Aira tenuissima (Mart. ex Schrank) Spreng., Syst. Veg. 1: 
276. 1825. Basionym: Panicum tenuissimum Mart. ex 
Schrank. 

= Sporobolus tenuissimus (Mart. ex Schrank) Kuntze. 


Aira triglumis Steud., Syn. Pl. Glumac. 1: 223. 1854. T- 
Protol.: USA: Texas, Vincenti 62. 
= Sporobolus junceus (P. Beauv.) Kunth. 


Airopsis millegrana Griseb., Abh. Konigl. Ges. Wiss. 
Gottingen 19: 252. 1874. T-Protol.: Argentina: 
Tucuman in monte Cuesta de Berico, Lorentz. T- 
Specim.: | of 1. Argentina: Tucuman: in monte Cuesta 
de Berico, no date, P.G. Lorentz & G. Hieronymus 842 
[HT: ?; IT: BA-38791, US-76315 (fragm.)]. 

= Eragrostis airoides Nees. 


Alloiatheros Elliott ex Raf., Bull. Bot. (Geneve) 1: 221. 1830. 
NOTE: nom. superfl. pro Gymnopogon. 
= Gymnopogon P. Beauy. 


Alloiatheros ambiguus Elliott ex Kunth, Enum, Pl. 1: 284. 


1833. 
NOTE: nom. inval., as syn. of Gymnopogon racemosus P. 
Beauv. 
= Gymnopogon ambiguus (Michx.) Britton, Sterns & 
Poggenb. 
Alloiatheros ambiguus Elliott ex B.D. Jacks., Index Kew. 1: 
83. 1893. 
NOTE: nom. inval., as syn. of Gymnopogon racemosus P. 
Beauv. 
= Gymnopogon ambiguus (Michx.) Britton, Sterns & 
Poggenb. 


Alloiatheros aristatus Raf. ex B.D. Jacks., Index Kew. 83. 


1893. 

NOTE: nom. inval., as syn. of Gymnopogon racemosus P. 
Beauv. 

= Gymnopogon ambiguus (Michx.) Britton, Sterns & 
Poggenb. 


Alloiatheros lepturoides (Nutt.) Steud., Nomencl. Bot. 2: 55. 
1840. Basionym: Anthopogon lepturoides Nutt. 
NOTE: nom. inval., as syn. of Gymnopogon racemosus P. 
Beauv. 
= Gymnopogon ambiguus (Michx.) Britton, Sterns & 
Poggenb. 


Allolepis Soderstr. & H.F. Decker, Madrofio 18(2): 34. 
1965. TYPE: Allolepis texana (Vasey) Soderstr. & 
H.F. Decker. 
TRTD. BY: Peterson, P. M. 
NHT: Monanthochloinae. NO. ACCPTD. SPP.: 1. 
CNSLTD. LIT.: Soderstrom, T. R. & H. F. Decker, (1965). 


Allolepis texana (Vasey) Soderstr. & H.F. Decker, Madrofio 
18(2): 34. 1965. Basionym: Poa texana Vasey. 
DST.: Mexi, USA. 


LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Beetle, A. A. (1983); Correll, D. S. & M. C. 
Johnston (1970); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Soderstrom, T. 
R. & H. F. Decker (1965). 

Distichlis texana (Vasey) Scribn.; Poa texana Vasey; 
Sieglingia wrightii Vasey. 


SYN.: 


Alopecurus glomeratus (Willd.) Poir., Encycl. 5: 495. 1817. 
Basionym: Polypogon glomeratus Willd. 
= Muhlenbergia glomerata (Willd.) Trin. 


Ammophila brevipilis (Torr.) Benth. ex Vasey, Grass. U.S. 
29. 1883. Basionym: Arundo brevipilis Torr. 
= Calamovilfa brevipilis (Torr.) Scribn. 


Ammophila curtissii Vasey, Bull. Torrey Bot. Club 11: 7. 
1884. T-Protol.: USA: Indian River, Florida , 4.1. 
Curtiss. T-Specim.: 1 of 1. USA: Florida: Brevard Co., 
Jul 1879, A.H. Curtiss 3412 (HT: US-76420]. 

= Calamovilfa curtissii (Vasey) Scribn. 


Andropogon ambiguus Michx., Fl. Bor.-Amer. 1: 58. 1803. 
T-Protol.: Hab. in sabulosis Carolinae. T-Specim.: | of 
1. USA: in sabulosis Carolinae, Michaux s.n. [HT: P; 
IT: P]. 
NOTE: as Andropogon “ambiguum". 
= Gymnopogon ambiguus (Michx.) Britton, Sterns & 
Poggenb. 


Andropogon barbatus L., Syst. Nat. (ed. 10) 2: 1305. 1759. 
T-Specim.: 1 of 1. Jamaica, P. Browne s.n. [LT: 
LINN-1211.28, US-76445, ~-US-2830895], LT 
designated by Hitchcock, Contr. U.S. Natl. Herb. 
12(3): 126 (1908). 

NOTE: as Andropogon "barbatun'. 
= Chloris elata Desv. 


Andropogon barbatus L., Mant. Pl. 302. 1771. T-Protol.: 
Habitat in India orientali. T-Specim.: | of 1. India [LT: 
LINN-1211.21], LT designated (as holotype) by 
Judziewicz in Gérts-van Rijn, Fl. Guianas, Ser. A, 8: 
136 (1990). 

NOTE: hom. illeg.; as Andropogon "“barbatum", SEE 
NOTES and see Taxon 25: 177 (1976) and Rhodora 
94: 135 (1992). Fosberg (Taxon 25:176-178.1976) 
does not consider this to be a new publication of a 
later hom. as does Anderson (Brigham Young Univ. 
Sci. Bull., Biol. Ser. 19(2):34 1974. Fosberg believes 
that L. was giving a n emended description of his A. 
harbatus LL. 1759, and , therefore, that L. was not 
creating a later hom. in 1763. 

E,. NAME: Andropogon barbatus L. 

= Chloris barbata Sw. 


Andropogon capensis Houtt., Nat. Hist. 13: prelim. 6(name), 
§79 (descr. as A. muticus), t. 93, f. 3. 1782. T-Specim.: 
1 of 1. South Africa: Cape, Houttyn s.n. [T: G]. 
NOTE: as "capense". 
= Eustachys paspaloides subsp. paspaloides. 


Andropogon curtipendulum (Michx.) Spreng. ex Steud., 
Nomencl. Bot. 1: 90. 1840. Basionym: Chloris 
curtipendula Michx. 

NOTE: nom. inval., as syn. of Eutriana curtipendula Trin. 
= Bouteloua curtipendula (Michx.) Torr. 


16 Catalogue of New World Grasses: II. 


Andropogon polydactylon L., Sp. Pl. (ed. 2) 2: 1483. 1763. T- near Harper's Ferry, Virginia and in the pine barrens of 


Protol.: Habitat in Jamaica, Brown jam. 364 [2647]. 
NOTE: as "polydactylon"; nom. illeg. superfl.; same type as 
A. barbatus L. (1759) [non 1771]. 
= Chloris elata Desv. 


Andropogon pubescens Aiton, Hort. Kew. 3: 423. 1789. 
NOTE: nom. nov., Dryander not author, see ICBN 46.6 ex. 
21. 
REPLACED NAME: Chloris ciliata Sw. 
BLOCKING NAME: Andropogon ciliatum Thunb. 
= Chloris ciliata Sw. 


Anoplia Steud., Syn. Pl. Glumac. 1: 210. 1854. 
NOTE: nom. inval., as syn. of Leptochloa anoplia Nees. 
= Leptochloa P. Beauv. 


Anoplia mexicana Nees ex Steud., Syn. Pl. Glumac. 1: 210. 
1854. 
NOTE: nom. inval., as syn. of Leptochloa anoplia Nees. 
= Leptochloa anoplia Nees. 


Anthephora axilliflora Steud., Syn. Pl. Glumac. |: 111. 1854. 
T-Protol.: USA: Texas. T-Specim.: 1 of 1. USA: 
Texas, 1835, Drummond 359 [IT: US-A865418 
(fragm. ex herb. Delessert)]. 

= Bouteloua dactyloides (Nutt.) Columbus. 


Anthephora belangeri Steud., Syn. Pl. Glumac. 1: 111. 1854. 
T-Protol.: Mexico , Belanger 1428. T-Specim.: | of 1. 
Mexico / USA, no date, C.P. Belanger 1428 [HT: ?; 
IT: US-A865423 (fragm. ex hb Delessert)]. 

= Hilaria belangeri (Steud.) Nash. 


Anthipsimus Raf., J. Phys. Chim. Hist. Nat. Arts 89: 105. 
1819. TYPE: Anthipsimus gonopodus Raf. 
= Muhlenbergia Schreb. 


Anthipsimus gonopodus Rat., J. Phys. Chim. Hist. Nat. Arts 
89: 105. 1819. T-Protol.: USA: Collines seches de 
l'Ohio, Oct. 

= Muhlenbergia schreberi J.F. Gmel. 


Anthochloa colusana (Burtt Davy) Scribn., Bull. Div. 
Agrostol., U.S.D.A. 17: 221, f. 517. 1899. Basionym: 
Stapfia colusana Burtt Davy. 

= Neostapfia colusana (Burtt Davy) Burtt Davy. 


Anthopogon Nutt., Gen. N. Amer. Pl. 1: 81. 1818. 
NOTE: nom. illeg. superfl. 
= Gymnopogon P. Beauv. 


Anthopogon brevifolius Nutt. ex Trin., Gram. Unifl. Sesquifl. 
238. 1824. 

NOTE: nom. inval., as syn. of Gymnopogon brevifolius 
Trin. 
= Gymnopogon brevifolius Trin. 

Anthopogon filiforme Nutt., Trans. Amer. Philos. Soc., n.s., 
5: 152. 1835. T-Protol.: Arkansas Territory: in shrubby 
prairies near the banks of the Arkansas; I first detected 
this very distinct species in Sussex county, Delaware a 
few miles from Lewiston, Apr. 1818, Nuttall s.n. T- 
Specim.: | of 1. USA: Arkansas: near the banks of the 
Arkansas River, Nuttall s.n.[HT: BM; IT: PH]. 

= Gymnopogon brevifolius Trin. 


Anthopogon lepturoides Nutt., Gen. N. Amer. Pl. 1: 82. 1818. 
T-Protol.: USA: Virgina: On the banks of the Potomac, 


Carolina and Georgia, H7:BM; IT: PH. 
= Gymnopogon ambiguus (Michx.) Britton, Sterns & 
Poggenb. 


Antichloa Steud., Nomencl. Bot. (ed. 2) 1: 108. 1840. 
NOTE: nom. inval., as syn. of Actinochloa. 
= Bouteloua Lag. 


Antitragus Gaertn., Fruct. Sem. Pl. 2: 7. 1790. TYPE: Crypsis 
aculeata (L.) Aiton. 
NOTE: nom. superfl. pro Crypsis. 
= Crypsis Aiton. 


Antonella Caro, Dominguezia 2: 18, f. IE-I. 1981. TYPE: 
Antonella nicorae (Anton) Caro. 
= Tridens Roem. & Schult. 


Antonella nicorae (Anton) Caro, Dominguezia 2: 18, f. 1E-1. 
1981. Basionym: Tridens nicorae Anton. 
NOTE: without exact basionym page. 
= Tridens nicorae Anton. 


Apera tenuiflora (Willd.) P. Beauv., Ess. Agrostogr. 142, 
151. 1812. Basionym: Agrostis tenuiflora Willd. 

= Muhlenbergia tenuiflora (Willd.) Britton, Sterns & 
Poggenb. 


Aplocera Raf., Med. FI. 2: 193. 1830. 
NOTE: nom. superfl. pro Monocera Elliott. 
= Ctenium Panz. 


Aplocera maritima Raf., Med. Fl. 2: 193. 1830. T-Protol.: sin. 
loc. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Arcangelina Kuntze, Revis. Gen. Pl. 2: 759. 1891. 
NOTE: nom. superfl. pro Kralikia. 
= Tripogon Roem. & Schult. 


Aristida adscensionis var. americana (L.) Kuntze, Revis. 
Gen. Pl. 33: 340. 1898. Basionym: Aristida americana 
L. 
= Bouteloua americana (L.) Scribn. 


Aristida americana L.., Syst. Nat. (ed. 10) 879. 1759. T- 
Protol.: Jamaica. T-Specim.: | of 1. Jamaica, P. 
Browne s.n. [LT: LINN-98.2; IT: US-2947074 
(fragm.)], LT designated by Hitchcock, Contr. U.S. 
Natl. Herb. 12: 123 (1908), but specific sheet not 
indicated; specific sheet indicated by Davidse in 
Cafferty et al., Taxon 49(2): 247 (2000). 

= Bouteloua americana (L.) Scribn. 


Aristida antillarum Poir., Encycl. 1: 451. 1810. T-Specim.: 1 
of 2. Antilles, Richard [ST]. 2 of 2. Porto-Ricco, Ledru 
[ST]. 
= Bouteloua americana (L.) Scribn. 


Aristida chloris Salzm. ex Dll, Fl. Bras. 2(3): 83. 1878. T- 
Protol.: Brazil. 
NOTE: nom. nud. 
= Gymnopogon foliosus (Willd.) Nees. 


Aristida geminata Willd. ex Steud., Nomencl. Bot. 1: 131. 
1840. T-Protol.: West Indies: St. Thomas. 
NOTE: nom. inval., as syn. of Gymnopogon foliosus 
(Willd.) Nees. 
= Gymnopogon foliosus (Willd.) Nees. 


Catalogue of New World Grasses: II. 17 


Aristida secunda Rud. ex Roemer & Schultes, Syst. Veg. 2: 
711. 1817. 
NOTE: nom. inval., as syn. of Dinebra secunda (Pursh) 
Roemer & Schultes. 
= Bouteloua curtipendula (Michx.) Torr. 


Aristida subbiflora Steud., Syn. Pl. Glumac. 1: 138. 1854. T- 
Protol.: West Indies: Guadeloupe , Duchaissing. T- 
Specim.: 1 of 1. Guadeloupe, no date, EP. de F. 
Duchassaing s.n. [T: IT: US-2830955 (fragm.)]. 

= Bouteloua americana (L.) Scribn. 


Aristidium (Endl.) Lindl., Veg. Kingd. (ed. 2) 116. 1847. 
Basionym: Eutriana sect. Aristidium Endl. 
= Bouteloua Lag. 


Arundinaria paraguayensis Kuntze, Revis. Gen. Pl. 3(2): 
341. 1898. T-Protol.: Paraguay: Nordliches Paraguay: 
Puerto Esperanza, Kuntze 56 [HT: NY]. T-Specim.: | 
of |. Paraguay: norliches Paraguay: Puerto Esperanza, 
Sep 1892, C.E.O. Kuntze 56 [HT: NY; IT: US- 
2809010]. 

NOTE: nom. altern. Sieglingia paraguayensis Kuntze, same 
page. 
= Gouinia paraguayensis (Kuntze) Parodi. 


Arundo brevipilis Torr., Fl. N. Middle United States 1: 95. 
1823. T-Protol.: USA: sandy swamps in the pine- 
barrens of New Jersey, near Quaker Bridge. T- 
Specim.: | of 1. USA: New Jersey, 1820, Anonymous 
sn. [T: IT: US-A865368 (fragm. ex herb. Torrey), US- 
78850 (fragm. ex herb. Torrey)]. 

= Calamovilfa brevipilis (Torr.) Scribn. 


Arundo quitensis Spreng., Pl. Min. Cogn. Pug. 2: 6. 1815. T- 
Protol.; Habitat ad Quito, cites in syn.: "Agrostis 
quitensis Humb., Willd. in lit". 

= Muhlenbergia tenuifolia (Kunth) Kunth. 


Arundo quitensis (Kunth) Poir., Encycl. 4: 704. 1816. 
Basionym: Calamagrostis quitensis Kunth. 
NOTE: hom. illeg. 
E. NAME: Arundo quitensis Spreng. 
= Muhlenbergia tenuifolia (Kunth) Kunth. 


Arundo reynaudiana Kunth, Revis. Gramin. 2: 275, t. 49. 
1830. T-Specim.: | of 1. unknown, unknown s.n. [T: 
L]. 
= Neyraudia reynaudiana (Kunth) Keng ex Hitchc. 


Arundo tenella Spreng., Pl. Min. Cogn. Pug. 2: 6. 1815. T- 
Protol.: Habitat in nova Hispania, in syn. cites: 
"Agrostis tenella Willd. in lit.". T-Specim.: | of 1. 
Mexico, Humboldt & Bonpland s.n. (HT: B-W]. 

NOTE: hom. illeg. 
E. NAME: Arundo tenella Schrad. 
= Muhlenbergia tenella (Kunth) Trin. 


Arundo tenuifolia (Kunth) Poir., Encycl. 4: 704. 1816. 
Basionym: Calamagrostis tenuifolia Kunth. 
= Muhlenbergia tenuifolia (Kunth) Kunth. 


Arundo zollingeri Biise, Pl. Jungh. 3: 343. 1854. T-Specim.: 1 
of 2. Java, Zollinger 337 [ST: L]. 2 of 2. Java, 
Zollinger 380 [ST: L]. 

= Neyraudia reynaudiana (Kunth) Keng ex Hitche. 


Athernotus  longifolius (Hook.) Lunell, Amer. Mid. 
Naturalist 4: 218. 1915. Basionym: Calamagrostis 
longifolia Hook. 


= Calamovilfa longifolia (Hook.) Hack. ex Scribn. 


Atherophora Steud., Nomencl. Bot. (ed. 2) 1: 167. 1840. 
TYPE: Hymenothecium tenellum (Cav. ex DC.) Lag. 
NOTE: nom. nud. 
= Aegopogon Humb. & Bonpl. ex Willd. 


Atheropogon Muhl. ex Willd., Sp. Pl. 4(2): 937. 1806 
("1805"). TYPE: Atheropogon apludioides Muhl. ex 
Willd. 

= Bouteloua Lag. 


Atheropogon acuminatus E. Fourn., Mexic. PI. 2: 139. 1886. 
T-Specim.: | of 3. Mexico: Mirador, Nov 1941, F.M. 
Liebmann 583 [ST: US-734843]. 2 of 3. 3 of 3. 
Mexico: Potrero de Consquitla, Oct, Liehmann 584 
[ST]. 

= Bouteloua curtipendula var. caespitosa Gould & 
Kapadia. 


Atheropogon affinis (Hook. f.) E. Fourn., Mexic. PI. 2: 141. 
1886. Basionym: Eutriana affinisHook. f. 
NOTE: incidental mention in Benth. & Hook. f., Gen. PI. 3: 
1169 (1883). 
= Bouteloua curtipendula var. curtipendula. 


Atheropogon americanus (L.) E. Fourn., Mexic. Pl. 2: 139. 
1886. Basionym: Aristida americana L. 
= Bouteloua americana (L.) Scribn. 


Atheropogon americanus var.depauperata E. Fourn., Mexic. 
Pl. 2: 139. 1886. T-Specim.: | of 2. Karwinski s.n. 
[ST: LE]. 2 of 2. Cuba, Wright 739 [ST: GH, GOET, 
P, US]. 
= Bouteloua repens (Kunth) Scribn. 


Atheropogon antillarum (Poir.) Spreng., Syst. Veg. 1: 294. 
1825. Basionym: Aristida antillarum Poir. 
= Bouteloua americana (L.) Scribn. 


Atheropogon apludioides Muhl. ex Willd., Sp. Pl. 4: 937. 
1806. T-Protol.. Habitat in America boreali. T- 
Specim.: | of 1. North America, Coll. Ukn. s.n. [HT: ?: 
IT: US-1535780 (fragm.)]. 

= Bouteloua curtipendula var. curtipendula. 


Atheropogon aristidioides (Kunth) Roem. & Schult., Syst. 
Veg. 2: 415. 1817. Basionym: Dinebra aristidoides 
Kunth. 

= Bouteloua aristidoides (Kunth) Griseb. 


Atheropogon bromoides (Kunth) Roem. & Schult., Syst. Veg. 
2: 415. 1817. Basionym: Dinebra bromoides Kunth. 
= Bouteloua radicosa (E. Fourn.) Griffiths. 


Atheropogon chondrosiodes (Kunth) Roem. & Schult., Syst. 
Veg. 2: 416. 1817. Basionym: Dinebra chondrosioides 
Kunth. 

= Bouteloua chondrosioides (Kunth) Benth. ex S$. Watson. 


Atheropogon curtipendulus (Michx.) E. Fourn., Mexic. PI. 2: 
138. 1886. Basionym: Chloris curtipendula Michx. 
= Bouteloua curtipendula (Michx.) Torr. 


Atheropogon distichus (Kunth) Spreng., Syst. Veg. 1: 294. 
1825. Basionym: Polyodon distichum Kunth. 
= Bouteloua disticha (Kunth) Benth. 


Atheropogon domingensis Spreng., Syst. Veg. 1: 293. 1825. 
T-Protol.: West Indies: Hispaniola. 
= Bouteloua juncea (Desv. ex P. Beauv.) Hitche. 


18 Catalogue of New World Grasses: II. 


Atheropogon filiformis E. Fourn., Mexic. Pl. 2: 140. 1886. T- 
Specim.: | of 4. Mexico: Hacienda de la Naranja, 
Karwinski 99/b [LT: LE], LT designated by ?:; cited 
by McVaugh, Fl. Novo-Galic. 14: 91 (1983). 2 of 4. 
Mexico: Consoquitla, Aug, Liebmann 573 [ST: C]. 3 
of 4. Mexico, Liebmann 585 [ST: C]. 4 of 4. Mexico, 
Liebmann 568 |ST: C]. 

= Bouteloua repens (Kunth) Scribn. 


Atheropogon gracilis (Kunth) Spreng., Syst. Veg. 1: 293. 
1825 [1824]. Basionym: Chondrosum gracile Kunth. 
= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Atheropogon hirtus (Kunth) Spreng., Syst. Veg. 1: 293. 1825. 
Basionym: Chondrosum hirtum Kunth. 
= Bouteloua hirsuta var. hirsuta. 


Atheropogon humilis (P. Beauv. ex Kunth) Spreng., Syst. 
Veg. 1: 293. 1825. Basionym: Chondrosum humile P. 
Beauv. ex Kunth. 

= Bouteloua simplex Lag. 


Atheropogon juncifolius (Desv.) Spreng., Syst. Veg. 1: 294. 
1825. Basionym: Heterosteca juncifolia Desv. 
= Bouteloua repens (Kunth) Scribn. 


Atheropogon medius E. Fourn., Mexic. Pl. 2: 139. 1886. T- 
Specim.: | of 2. Mexico: Hacienda del Limon, 
Karwinsky [ST]. 2 of 2. Mexico: between T. Miguel 
and Sadani, Oct, Liebmann 581 [LT: US-2307995 
(fragm.)], LT designated by Swallen, N. Amer. FI. 17: 
633 (1939), 

= Bouteloua media (E. Fourn.) Gould & Kapadia. 


Atheropogon oligostachyus Nutt., Gen. N. Amer. Pl. 1: 78. 
1818. T-Specim.: | of 1. USA: plains of the upper 
Missouri, Nuftall s.n. [HT: BM; IT: PH]. 

= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Atheropogon papillosus Engelm., Amer. J. Sci. 46: 104, 
1843. T-Specim.: | of 1. USA: Illinois: Cass Co., 

1842, Geyer s.n. [HT: MO]. 

= Bouteloua hirsuta var. hirsuta. 


Atheropogon polymorphus E. Fourn., Mexic. Pl. 2: 141. 
1886. T-Protol.: Mexico: San Luis de Potosi, Virlet 
1393, T-Specim.: | of 1. Mexico: San Luis Potosi, 
Virlet 1395 [T: 1T: US-A865361 (fragm.)]. 

= Bouteloua polymorpha (E. Fourn.) Columbus. 


Atheropogon procumbens (P. Durand) J. Jacq., Ecl. Gram. 
Rar. 2: 16, t. 12. 1813. Basionym: Chloris procumbens 
P. Durand. 
= Bouteloua simplex Lag. 


Atheropogon racemosus (Lag.) Roem. & Schult., Syst. Veg. 
2: 414. 1817. Basionym: Bouteloua racemosa Lag. 
= Bouteloua curtipendula var. caespitosa Gould & 
Kapadia. 


Atheropogon radicosus E. Fourn., Mexic. Pl. 2: 140. 1886. T- 
Specim.: | of 1. Mexico: near Mexico City, 8 Jun 
1865-18 Jun 1866,M. Bourgeau 450 [LT: P; IST: BR, 
LE, US-81791, US-87200 (fragm. ex P), US-883896 
(fragm)], LT designated by ?; cited by McVaugh, FI. 
Novo-Galic. 14: 92 (1983). 

= Bouteloua radicosa (E. Fourn.) Griffiths. 


Atheropogon repens (Kunth) Roem. & Schult., Syst. Veg. 2: 
416. 1817. Basionym: Dinebra repens Kunth. 


= Bouteloua repens (Kunth) Scribn. 


Atheropogon scorpiodes (Lag.) Spreng., Syst. Veg. 1: 293. 
1825. Basionym: Bouteloua scorpioides Lag. 
= Bouteloua scorpioides Lag. 


Atheropogon stolonifer E. Fourn., Mexic. Pl. 2: 140. 1886. T- 
Protol.; Mexico: La Parada, Sierra de Oaxaca, 8500 ft, 
Apr, Liebmann 588. T-Specim.: | of 1. Mexico: La 
Parada, Sierra de Oaxaca, 1842, Liebmann 588 [T: IT: 
US-A865362 (fragm.)]. 
= Bouteloua reederorum Columbus. 


Atheropogon triaena Trin. ex Spreng., Syst. Veg. 1: 293. 
1825. 
REPLACED NAME: Triaena racemosa Kunth. 
BLOCKING NAME: Atheropogon racemosus (Lag.) Roem. 
& Schult. 
= Bouteloua triaena (Trin. ex Spreng.) Scribn. 


Atropis carinata Griseb., Abh. Konigl. Ges. Wiss. Gottingen 
24: 291. 1879. T-Protol.: Argentina: Jujuy [error for in 
salsis pr. el Volcan (Paraguay: Bal. pl. parag. 131. a.)], 
Balansa. 
= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Axonopus poiretii Roem. & Schult., Syst. Veg. 2: 318. 1817. 
REPLACED NAME: Agrostis compressa Poir. 
BLOCKING NAME: Agrostis compressa Willd. 
= Sporobolus junceus (P. Beauv.) Kunth. 


Baldomiria Herter, Revista Sudamer. Bot. 6(5-6): 145. 1940. 
TYPE: Baldomiria chloridiformis (Hack. ex Stuck.) 
Herter. 

= Leptochloa P. Beauv. 


Baldomiria chloridiformis (Hack. ex Stuck.) Herter, Revista 
Sudamer. Bot. 6(5-6): 145, f. 10. 1940. Basionym: 
Diplachne chloridiformis Hack. ex Stuck. 

= Leptochloa chloridiformis (Hack. ex Stuck.) Parodi. 


Bauchea E. Fourn., Mexic. Pl. 2: 87. 1886. TYPE: Bauchea 
karwinskyi E. Fourn. 
= Sporobolus R. Br. 


Bauchea karwinskyi E. Fourn., Mexic. Pl. 2: 87, 1886. T- 
Specim.: | of 2, Mexico, no date, WF. Karwinski von 
Karwin 1015 (IST: US-998324 (fragm.)]. 2. of 2. 
Mexico: Tanquecillas, Karw. /0/5b [ST]. 

NOTE: Fourn. ex Hemsl. Biol. Centr. Amer. Bot. 3: 550. 
1885, nom. nud. 
= Sporobolus wrightii Munro ex Scribn. 


Bealia Scribn., True Grasses 104, f. 45a. 
Bealia mexicana Scribn. 
TRTD. BY: Peterson, P. M. 
NHT: Muhlenbergiinae. NO. ACCPTD. SPP.: 1. 
CNSLTD. LIT.: Peterson, P. M., (1989); Peterson, P. M., 
M. R. Duvall & A. H. Christensen, (1993). 


1890. TYPE: 


Bealia mexicana Scribn., True Grasses 103, f. 45a. 1890. T- 
Protol.: Mexico: Chihuahua, Pringle 819. T-Specim.: | 
of 1. Mexico: Chihuahua: near Chihuahua, thin soil of 
dry porphyry mts., 7 Oct 1886,C.G. Pringle 819 [LT: 
US-995611; ILT: LE, MO, NME, NY, UC, US- 
967198, US-723648, US-820886, US-1645965, US- 
90613, VT, WIS], LT designated by Hitchcock, Contr, 
U.S. Natl. Herb. 17(3): 294 (1913); redesignated by 
Peterson (without sheet no.), Madrofio 36: 262 (1989). 


Catalogue of New World Grasses: II. 19 


NOTE: name definitely accepted by Scribner (although only 
provisionally by Hackel himself) and validated by 
illustr. (ICBN 44.1) and diagnosis. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Peterson, P. M. & Y. Herrera A. 
(1995): Peterson, P. M. (1989); Peterson, P. M. 
(2000). 

SYN.: Bealia mexicana Scribn. ex Beal; Epicampes 
mexicana (Scribn.) M.E. Jones; Muhlenbergia biloba 
Hitche. 


Bealia mexicana Scribn. ex Beal, Grass. N. Amer. 2: 267. 
1896. 
NOTE: isonym. 
E. NAME: Bealia mexicana Scribn. 
= Bealia mexicana Scribn. 


Bealia speciosa (Vasey) Beal, Grass. N. Amer. 2: 268. 1896. 
Basionym: Muhlenbergia speciosa Vasey. 
= Muhlenbergia speciosa Vasey. 


Bennetia Raf., Bull. Bot. (Geneve) 1: 220. 1830. TYPE: 
Agrostis juncea Michx.; hom. illeg. Heleochloa 
juncea P. Beauv. 

NOTE: later hom., non Gray 1821. 

= Sporobolus R. Br. 


Bennetia juncea Raf. ex B.D. Jacks., Index Kew. 1: 291. 
1893. 
NOTE: nom. inval., as syn. of Sporobolus junceus (Michx.) 
Kunth. 
= Sporobolus junceus (P. Beauv.) Kunth. 


Biatherium Desv., Mem. Soc. Agric. Angers 1: 172, t. 8, f. 2. 
1831. TYPE: Biatherium foliosa (Willd.) Desv. 
NOTE: also in Opusc. Sci. Phys. Nat. 72 [reprint] 1831- 
1833. 
= Gymnopogon P. Beauv. 


Biatherium foliosa (Willd.) Desv., Mem. Soc. Agric. Angers 
1: 176, t. 8 £ 2. 1831. Basionym: Chloris foliosa 
Willd. 

NOTE: also in Opusc. Sci. Phys. Nat. 72, t. 5, f. 2 [reprint] 
1831-1833. 
= Gymnopogon foliosus (Willd.) Nees. 


Blepharidachne Hack., Nat. Pflanzenfam. 2(2): 68. 
1887. 
REPLACED NAME: Eremochloe S. Watson. 
BLOCKING NAME: Eremochloa Biise. 
TRTD. BY: Valdés-Reyna, J. & P. M. Peterson. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 4. 
CNSLTD. LIT.: Hunziker, A. T. & A. M. Anton, (1979). 
SYN.: Eremochloe S. Watson. 


Blepharidachne benthamiana (Hack. ex Kuntze) Hitchc., 
Bull. Div. Agrostol., U.S.D.A. 772: 78. 1920. 
Basionym: Munroa benthamiana Hack. ex Kuntze. 

DST.: Arge. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 

SYN.: Munroa benthamiana Hack. ex Kuntze. 


Blepharidachne bigelovii (S. Watson) Hack., Monogr. Phan. 
6: 261. 1889. Basionym: Eremochloe bigelovii S. 
Watson. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Beetle, A. A. (1987); Correll, D. S. & M. C. 
Johnston (1970); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Hitchcock, A. S. 
(1951). 

SYN.: Eremochloe  bigelovii 
thurberi S. Watson. 


S. Watson; 


Eremochloe 


Blepharidachne hitchcockii Lahitte, Physis (Buenos Aires) 
15: 305. 1939. T-Protol.: Argentina: La Rioja: cerca 
del Pozo de Avila, 31 Mar 1901.4. Giacomelli [Hrb. 
Ministerio de Agricultura 1638] (HT: hrb. Ministerio 
de Agricultura]. T-Specim.: 1 of I. Argentina: La 
Rioja: cerca del Pozo de Avila, 31 Mar 1901, A. 
Giacomelli 1638 |HT: BAB; IT: BAF, US-3168447 
(fragm.)]. 

DST.: Arge. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 


Blepharidachne kingii (S. Watson) Hack., Monogr. Phan. 6: 
261. 1889. Basionym: Eremochloe kingii S. Watson. 

DST.: USA, 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Cronquist, A., 
A. H. Holmgren, N. H. Holmgren & J. L. Reveal 
(1977); Hickman, J. C. (1993); Hitchcock, A. S. 
(1951); Munz, P. A. & D. D. Keck (1959); Munz, P. 
A. (1974). 

SYN.: Eremochloe kingii S. Watson. 


Blepharoneuron Nash, Bull. Torrey Bot. Club 25: 
88. 1898. TYPE: Blepharoneuron tricholepis (Torr.) 
Nash. 

TRTD. BY: Peterson, P. M. 
NHT: Muhlenbergiinae. NO. ACCPTD. SPP: 2. 
CNSLTD. LIT.: Peterson, P. M. and C. R. Annable, (1990). 


Blepharoneuron shepherdii (Vasey) P.M. Peterson & 
Annable, Syst. Bot. 15(4): 519. 1990. Basionym: 
Sporobolus shepherdii Vasey. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Peterson, P. M. 
& C.R. Annable (1990); Peterson, P. M. (2000). 

Muhlenbergia  shepherdii (Vasey) —Swallen; 

Sporobolus shepherdii Vasey. 


SYN:: 


Blepharoneuron tricholepis (Torr.) Nash, Bull. Torrey Bot. 
Club 25(2): 88. 1898. Basionym: Vilfa_tricholepis 
Torr. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Beetle, A. A. (1987); Correll, D. S. & M. C. 
Johnston (1970); Cronquist, A., A. H. Holmgren, N. 
H. Holmgren & J. L. Reveal (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W. & R. Moran (1981): Hitchcock, A. S. 


20 Catalogue of New World Grasses: II. 


(1951); McVaugh, R. (1983); Peterson, P.M. & C. R. 
Annable (1990); Peterson, P. M. (2000). 

SYN.: Sporobolus tricholepis (Yorr.) J.M. Coult.; Vilfa 
tricholepis Torr. 


Boriskellera Terechoy, Del. Sem. Hort. Reg. Bot. Kujbyshev 
13. 1938. TYPE: Boriskellera arundinacea (L.) 
Terechov. 

= Eragrostis Wolf. 


Botelua Lag., Varied. Ci. 2(4): 134. 1805. 
NOTE: orth. var. 
= Bouteloua Lag. 


Botelua americana (L.) Scribn., Proc. Acad. Nat. Sci. 
Philadelphia 1891: 306. 1891. Basionym: Aristida 
americana L. 

NOTE: nom. inval. orth. var. 
CORRECT NAME: Bouteloua americana (L.) Scribn. 
= Bouteloua americana (L.) Scribn. 


Bouteloua Lag., Varied. Ci. 2(4,21): 134. 1805. TYPE: 
Bouteloua racemosa Lag. 
NOTE: orth. cons.; originally as "Botelua"; emend. Lag., 
Gen. Sp. Nov. 5 (1816). 

TRTD. BY: Peterson, P. M. & Y. Herrera-Arrieta. 

NHT: Boutelouinae. NO. ACCPTD. SPP.:54 SUBSP:16. 
CNSLTD. LIT.: Gould, F. W., (1979); Griffiths, D., (1912); 
Columbus, J. T., M. S. Kinney, R. Pant & M. E. 
Siqueiros Delgado, (1998). 

Actinochloa Willd. ex Roem. & Schult.; Antichloa 
Steud.; Aristidium (Endl.) Lindl.; Atheropogon Muhl. 
ex Willd.; Botelua Lag.; Bouteloua Hornem. ex P. 
Beauv.; Buchloe Engelm.; Buchlomimus Reeder, C. 
Reeder & Rzed.; Bulbilis Raf. ex Kuntze; Calanthera 
Hook.; Cathestecum J. Presl; Chondrosium Desv.; 
Chondrosum Desv.; Cyclostachya Reeder & C. 
Reeder; Erucaria Cerv.; Eutriana sect. Aristidium 
Endl.; Eutriana sect. Triplathera Endl.; Eutriana 
Trin.; Fourniera Scribn.; Griffithsochloa G.J. Pierce; 
Heterosteca Desv., Lasiostega Benth.; Nestlera 
Steud.; Opizia J. Presl; Pentarrhaphis Kunth; 
Pleiodon Rchb.; Polyadon Kunth; Polyschistis J. 
Presl; Pringleochloa Scribn.; Soderstromia C.V. 
Morton; Strombodurus Steud.; Triaena Kunth; 
Triathera Desv.; Triplathera (Endl.) Lindl. 


SYN.: 


Bouteloua Hornem. ex P. Beauv., Ess. Agrostogr. 40. 1812. 
NOTE: later hom., non Lagasca 1805. 
= Bouteloua Lag. 


Bouteloua acuminata (E. Fourn.) Griffiths, Contr. U.S. Natl. 
Herb. 14(3): 406, f. 55. 1912. Basionym: A theropogon 
acuminatus E, Fourn. 

NOTE: missapplied to Palmer 1655 which is B. reflexa 
Swallen. 

= Bouteloua curtipendula var. caespitosa Gould & 
Kapadia. 


Bouteloua alamosana Vasey, Contr. U.S. Natl. Herb. 1(4): 
115. 1891. T-Protol.: Mexico: on rocky ridges, 
Alamos, 10 - 30 Sep, E. Palmer 698. T-Specim.: | of 
1. Mexico: Sonora: Alamos, 16-30 Sep 1890, E. 
Palmer 698 |HT: US-81717; IT: NY, TAES, US- 
89471, US-883372]. 

DST.: CoRi, ElSa, Guat, Hond, Mexi, Nica. 
LVL. ACCPTC.: 1. 


ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Berendsohn, W. G. & A. E. Araniva de 
Gonzalez (1991); Breedlove, D. E. (1986); Davidse, 
G., M. Sousa S. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. (1980); Molina 
R., A. (1975); Pohl, R. W. (1980); Pohl, R. W. 
(1994); Reyes-Garcia, A. & M. Sousa S. (1997). 

SYN.: Bouteloua longiseta Gould. 


Bouteloua americana (L.) Scribn., Proc. Acad. Nat. Sci, 
Philadelphia 1891: 306. 1891. Basionym: Aristida 
americana L. 

DST.: Braz, Cari, Colo, CoRi, Ecua, ElSa, Guat, Guya, 
Hond, Mexi, Nica, Pana, USA, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Anonymous 
(1986); Beetle, A. A. (1987); Berendsohn, W. G. & 
A. E. Araniva de Gonzalez (1991); Breedlove, D. E. 
(1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J.  Valdes-Reyna (2000); 
Filgueiras, T. S. (1999); Gould, F. W (1979): Gould, 
F. W. (1980); Hitchcock, A. S. (1909); Hitchcock, A. 
S. (1936); Jorgensen, P. M. & S. Leon-Yanez (eds.) 
(1999); Judziewicz, E. J. (1990); Liogier, H. & L. 
Martorell (1982); Molina R., A. (1975); Pohl, R. W. 
(1980); Pohl, R. W. (1994). 

SYN.: Aristida adscensionis var. americana (L.) Kuntze; 
Aristida americana 1., Aristida antillarum Poir.; 
Aristida subbiflora Steud.; Atheropogon americanus 
(L.) E. Fourn.; Atheropogon  antillarum (Poir.) 
Spreng.; Botelua americana (L.) Scribn.; Bouteloua 
americana (L.) Hitche.; Bouteloua americana (L.) 
Pilg.; Bouteloua  elatior Griseb.; Bouteloua 
humboldtiana Griseb.; Bouteloua litigiosa Lag.; 
Bouteloua porphyrantha C. Wright; Chaetaria 
antillarum (Poir.) P. Beauv. ex Roem. & Schult.; 
Dinebra americana (L.) P. Beauv.; Eutriana 
antillarum (Poir.) Steud.; Heferostega americana (L.) 
Desv.; Triathera americana (L.) Desv. 


Bouteloua americana (L.) Hitche., Annual Rep. Missouri 
Bot. Gard. 1893: 148. 1893. Basionym: Aristida 
americana L. 

NOTE: isonym. 

E. NAME: Bouteloua americana (L.) Scribn. 
= Bouteloua americana (L.) Scribn. 


Bouteloua americana (L.) Pilg., Symb. Antill. 5: 288. 1907. 
Basionym: Aristida americana L. 
NOTE: isonym. 
E. NAME: Bouteloua americana (L.) Scribn. 
= Bouteloua americana (L.) Scribn. 


Bouteloua annua Swallen, J. Wash. Acad. Sci. 25(9): 414. 
1935. T-Protol.: Mexico: Baja California Sur: 4 mi E 
of San Ignacio, 6 Mar 1935, Shreve 7032 [HT: US- 
1611715]. T-Specim.: 1 of 1. Mexico: Baja California 
Sur: 4 mi E of San Ignacio, 6 Mar 1935, F. Shreve 
7032 (HT: US-1611715; IT: GH, US-1614953]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. & R. Moran 


Catalogue of New World Grasses: II. 21 


(1981); Gould, F. W. (1980); Shreve, F. & I. L. 
Wiggins (1964). 


Bouteloua arenosa Vasey, U.S.D.A. Div. Bot. Bull, 12(1): t. 
34. 1890. T-Protol.: Mexico: loose sandy soil, at 
Guaymas on the Gulf of California, Palmer 189. T- 
Specim.: 1 of 1. Mexico: Sonora: Guayamas, 1887, E. 
Palmer 189 {HT: US-81716]. 

NOTE: Vasey in S. Wats.; Proc. Amer. Acad. 24: 81. 1889, 
nom. nud. 
= Bouteloua barbata var. barbata. 


Bouteloua aristidoides (Kunth) Griseb., Fl. Brit. W. I. 537. 
1864. Basionym: Dinebra aristidoides Kunth. 

DST.: Arge, Boli, Braz, Cari, Ecua, Mexi, Para, Peru, USA. 

LVL. ACCPTC.:: |. 

ALSO ACCPTD. BY: Abrams, L. (1923); Anonymous 
(1986); Beetle, A. A. (1977); Beetle, A. A. (1987); 
Bor, N. L. (1960); Brako, L. & J. L. Zarucchi (1993); 
Correll, D. S. & M. C. Johnston (1970); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Foster, R. C. (1958); Gould, F. W. & R. 
Moran (1981); Gould, F. W. (1980); Hitchcock, A. S. 
(1927): Hitchcock, A. S. (1951); Jorgensen, P. M. & 
S. Leén-Yanez (eds.) (1999); Lindeman, J. C. & A. 
L. Stoffers (1963); McVaugh, R. (1983); Munz, P. A. 
& D. D. Keck (1959); Munz, P. A. (1974); Renvoize, 
S. A. (1998); Shreve, F. & I. L. Wiggins (1964); 
Tovar, O. (1993); Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 

SYN.: Atheropogon aristidioides (Kunth) Roem. & Schult.; 
Bouteloua ciliata Griseb.; Bouteloua gracilis Vasey; 
Dineba hirsuta J. Presl; Dinebra aristidoides Kunth; 
Dinebra_ hirsuta J, Presl,; Eutriana  aristidoides 
(Kunth) Trin.; Eutriana hirsuta (J. Presl) Kunth; 
Triathera aristidoides (Kunth) Nash. 


Bouteloua aristidoides (Kunth) Griseb. var. aristidoides. 
LVL. ACCPTC:: 1. 
ALSO ACCPTD. BY: Gould, F. W. (1980); Hickman, J. C. 
(1993). 


Bouteloua aristidoides var. arizonica M.E. Jones, Contr. W. 
Bot. 14: 13. 1912. T-Protol.: USA: Arizona: Tucson, 
Thornber 177 [HT: US]. T-Specim.: | of 1. USA: 
Arizona, no date, J.J. Thornber 177 [HT: ?; IT: US- 
3168446 (fragm.)]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. 
(1980); Hitchcock, A. S. (1951). 


Bouteloua aschenborniana Griseb. ex E. Fourn., Mexic. PI. 
2: 137. 1886. 
NOTE: nom. inval., as syn. 
aschenbornianum Nees. 
= Bouteloua hirsuta var. hirsuta. 


of Chondrosium 


Bouteloua barbata Lag., Varied. Ci. 2(4): 141. 1805. T- 
Specim.: | of 1. Philippinis Insulis [error for Mexico], 
Nee [HT: MA; IT: US-A865567 (fragm. ex MA & 
photo)]. 

DST.: Arge, Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Abrams, L. (1923); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 


Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Esparza Sandoval, S. & Y. Herrera 
Arrieta (1996); Espejo Serna, A., A. R. L6pez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. & 
R. Moran (1981); Gould, F. W. (1980); Great Plains 
Flora Association (1986); Hitchcock, A. S. (1951); 
McVaugh, R. (1983); Munz, P. A. & D. D. Keck 
(1959); Munz, P. A. (1974); Shreve, F. & I. L. 
Wiggins (1964). 

Actinochloa barbata (Lag.) Roem. & Schult.; 
Bouteloua barbata var. barbata;  Bouteloua 
micrantha Scribn. & Merr.; Bouteloua microstachya 
(E. Fourn.) L.H. Dewey; Bouteloua polystachya 
(Benth.) Torr; Bouteloua  polystachya — var. 
polystachya; Bouteloua pumila Buckley; 
Chondrosium polystachyum Benth.; Chondrosium 
subscorpioides Mill. Hal.; Chondrosum barbatum 
(Lag.) Clayton; Chondrosum exile FE. Fourn.; 
Chondrosum microstachyum E. Fourn.; Chondrosum 
polystachyum Benth.; Chondrosum subscorpioides 
Mill. Hal.; Eutriana barbata (Lag.) Kunth. 


SYN.: 


Bouteloua barbata var. arenosa (Vasey) Beetle, Phytologia 
54(1): 1. 1983. Basionym: Bouteloua arenosa Vasey. 
= Bouteloua barbata var. barbata. 


Bouteloua barbata Lag. var. barbata. 

DST.: Arge, Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1987), Esparza 
Sandoval, S. & Y. Herrera Arrieta (1996); Gould, F. 
W. & R. Moran (1981); Gould, F. W. (1980): 
Hickman, J. C. (1993). 

Actinochloa barbata (Lag.) Roem. & Schult.; 
Bouteloua arenosa Vasey; Bouteloua barbata var. 
arenosa (Vasey) Beetle; Bouteloua micrantha Scribn. 
& Merr.; Bouteloua microstachya (E. Fourn.) L.H. 
Dewey; Bouteloua polystachya (Benth.)  Torr.; 
Bouteloua pumila Buckley; = Chondrosium 
polystachyum Benth.; Chondrosium subscorpioides 
Mull. Hal.; Chondrosum exile E.  Fourn; 
Chondrosum microstachyum E. Fourn.; Chondrosum 
polystachyum Benth.; Chondrosum subscorpioides 
Mill. Hal.; Erucaria tetrastachya Cerv.; Eutriana 
barbata (Lag.) Kunth. 


SYN:: 


Bouteloua barbata var. major (Vasey) Govaerts, World 
Checklist of Seed Plants 2(1-2): 13. 1996. Basionym: 
Bouteloua polystachya var. major Vasey. 

NOTE: validated here. 
= Bouteloua barbata var. rothrockii (Vasey) Gould. 


Bouteloua barbata var. rothrockii (Vasey) Gould, Ann. 
Missouri Bot. Gard. 66(3): 403. 1979 [1980]. 
Basionym: Bouteloua rothrockii Vasey. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Esparza 
Sandoval, S. & Y. Herrera Arrieta (1996); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Gould, F. W. & R. Moran (1981); Gould, F. 
W. (1980). 

Bouteloua barbata var. major (Vasey) Govaerts; 
Bouteloua polystachya var.major Vasey, Bouteloua 
rothrockii Vasey. 


SYN.: 


22 Catalogue of New World Grasses: II. 


L 


Bouteloua barbata var. sonorae (Griffiths) Gould, Ann. 
Missouri Bot. Gard. 66: 403. 1979[1980]. Basionym: 
Bouteloua sonorae Griffiths. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Esparza Sandoval, S. & Y. Herrera 
Arrieta (1996); Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. 
(1980). 

SYN.: Bouteloua sonorae Griffiths. 


Bouteloua brachyanthera Phil., Verz. Antofagasta Pfl. 85. 
1891. T-Protol.: Chile: Tarapaca: pariter ad Calcalhuay 
lecta, F. Philippi s.n. T-Specim.: 1 of 1. Chile: 
Yarapaca, FL Philippi sn. [IT: US-A0865572 
(fragm.)]. 

= Bouteloua simplex Lag. 


Bouteloua bracteata (McVaugh) Columbus, Aliso 18(1): 63. 


1999. Basionym: Opizia bracteata McVaugh. 
DST.: Mexi. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000). 
SYN.: Opizia bracteata McVaugh. 


Bouteloua brasiliensis Ekman, Ark. Bot. 10(17): 29, t. 4, f. 4, 


t.6, f£. 15. 1911. T-Protol.: Brazil: Matto Grosso: locis 
glareosis ad marginnum silvulae, in fructicetis, 23 Apr 
1903, G.O. Malme 3106 [Exp. I Regn.] [HT: S; IT: 
US]. T-Specim.: | of 1. Brazil: Mato Grosso, 23 Apr 
1903, G.O.A. Malme 3106 [HT: S; IT: US-702282, 
US-2765856 (fragm.)]. 

= Bouteloua curtipendula (Michx.) Torr. 


Bouteloua brevifolia Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 93. 1862.  T-Protol.: USA: 
northwestern Texas. T-Specim.: | of 2. USA: Texas, 
May-Oct 1849, C.H. Wright 748 [LT: PH; ILT: US- 
81715], LT designated by Gould, Ann. Missouri Bot. 
Gard. 66: 397 (1979), bt the only the specific 
herbarium of a isolectotype designated. 2 of 2. USA: 
Texas, Fendler 950 [ST: PH, US]. 

= Bouteloua eriopoda (Torr.) Torr. 


Bouteloua breviseta Vasey, Contr. U.S. Natl. Herb. 1(2): 58. 
1890. T-Protol.: USA: Texas: Presidio Co., /C.G. 
Nealley 669]. T-Specim.: | of 1. USA: Texas: Presidio 
Co.: Screw Bean, 1889, Neally 669 [HT: US-80198 
(fragm. ex F)], Reeder & Reeder, Syst. Bot. 5: 318 
(1980) point out that Presidio of Vasey County 
actually is the modern Reeves Co, 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Gould, F. W. (1980): Hitchcock, A. S. 
(1951); Reeder, J. R. & C. G. Reeder (1980). 

SYN.: Chondrosum brevisetum (Vasey) Clayton. 


Bouteloua bromoides Lag., Gen. Sp. Pl. 5. 1816. T-Protol.: 
Habitat in Acapulco in N. Hispania. T-Specim.: | of 1. 
Mexico: Guerrero, Coll. Ukn. S\N. [HT: MA-181580; 
IT: US-A865568 (fragm.)]. 

= Bouteloua repens (Kunth) Scribn. 


Bouteloua bromoides Lag. var. bromoides. 


= Bouteloua repens (Kunth) Scribn. 


Bouteloua bromoides var. radicosa (E. Fourn.) Vasey ex L.H. 
Dewey, Contr. U.S. Natl, Herb. 2(3): 533. 1894. 
Basionym: Atheropogon radicosus E. Fourn. 

= Bouteloua radicosa (E. Fourn.) Griffiths. 


Bouteloua burkii Scribn. ex S. Watson, Proc. Amer, Acad. 
Arts 18: 179. 1883. T-Specim.: | of 3. Berlandier 167 
[ST: GH?]. 2 of 3. USA: Texas: Laredo, Berlandier 
1427 [ST: GH; IST: K, LE-TRIN-2130.01, P, US- 
A865684 (1 ex P; 2 ex W), US-204622, US-1084518 
(ex BM), US-883356, W]. 3 of 3. USA: West Texas, 
Upper Concho, sandy plains, Apr 1882,Curtiss [North 
American Plants] 3440 [=Reverchon] |ST: GH?; IST: 
US-820915, US-883346]. 

NOTE: as Bouteloua Burkei, to be treated as an 
orthographic variant as Burk [not Burke] is the 
correct spelling. 

= Bouteloua trifida Thurb. 


Bouteloua burkii Scribn., Bull. Torrey Bot. Club 11: 5, 1884. 
T-Specim.: | of 2. USA: western Texas and New 
Mexico, Berlandier 1427 [ST: US-883356, US- 
204622; IST: GH, K, LE-TRIN-2130.01, P, US- 
883356, US-1084518 (ex BM), US-a865684 (1 ex P; 2 
ex W), W]. 2 of 2. USA: Texas: Laredo, Mrs. Anna B. 
Nickles (Comm. Mr. Isaac Burk) [ST], Possibly 
destroyed when Scribner's herbarium burned. fide. 
Griffiths (1912). not found at US by Griffiths or by 
RJS 1999, 

NOTE: nom. superfl. illeg.; as Bowteloua Burkii. 
= Bouteloua trifida Thurb. 


Bouteloua chasei Swallen, Proc. Biol. Soc. Wash. 56: 81. 
1943. T-Protol.: Mexico: Nuevo Leén: Galeana, Mesa, 
5400 ft, 30 Jul 1939, Chase 7673 1/2 |HT: US- 
1763078]. T-Specim.: 1 of 1. Mexico: Nuevo Leon: 
Galeana Mun., 30 Jul 1939, V.H. Chase 7673-1/2 [HT: 
US-1763078]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. LOpez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. (1980). 

SYN.: Chondrosum chasei (Swallen) Clayton. 


Bouteloua chihuahuana (M.C. Johnst.) Columbus, Aliso 
14(3): 227. 1996. Basionym: Bouteloua rigidiseta var. 
chihuahuana M.C. Johnst. 

DST.: Mexi. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Columbus, J. T. (1996); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000). 

SYN.: Bouteloua rigidiseta var. chihuahuana M.C. Johnst. 


Bouteloua chondrosioides (Kunth) Benth. ex S. Watson, 
Proc. Amer. Acad. Arts 18: 179. 1883. Basionym: 
Dinebra chondrosioides Kunth. 

DST.: CoRi, ElSa, Guat, Hond, Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A, 
(1977); Beetle, A. A. (1987); Berendsohn, W. G. & 
A. E. Araniva de Gonzalez (1991); Breedlove, D. E. 
(1986); Correll, D. S. & M. C. Johnston (1970); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lopez- 


Catalogue of New World Grasses: II. 23 


SYN.: 


Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. 
(1980); Hitchcock, A. S. (1951); McClure, F. A. 


(1955); McVaugh, R. (1983); Molina R., A. (1975); 


Pohl, R. W. (1980); Pohl, R. W. (1994); Shreve, F. & 
I. L. Wiggins (1964). 

Actinochloa ovata (Lag.) Roem. & Schult.; 
Atheropogon chondrosiodes (Kunth) Roem. & 
Schult.; Bouteloua havardii Vasey ex S. Watson; 
Bouteloua ovata Lag., Chondrosum humboldtianum 
Kunth; Dinebra chondrosioides Kunth; Eutriana 
cristata Trin. 


Bouteloua ciliata Griseb., Abh. Konigl. Ges. Wiss. Gottingen 


24: 302. 1879. T-Protol.: Argentina: S. ad_ fl. 
Juramento, Lorentz & Hieronymus 352. T-Specim.: | 
of 1. Argentina: Salta: Juramento, P.G. Lorentz & G. 
Hieronymus 352 [HT: GOET; IT: US-A865569 
(fragm.)], CS\Ex griseb. herb. 


= Bouteloua aristidoides (Kunth) Griseb. 


Bouteloua curtipendula (Michx.) Torr., Not. Milit. Reconn. 


153. 1848. Basionym: Chloris curtipendula Michx. 


NOTE: prov. name, basionym not cited. 
= Bouteloua curtipendula (Michx.) Torr. 


Bouteloua curtipendula (Michx.) Torr., Explor. Red River 


Louisiana 300. 1853. Basionym: Chloris curtipendula 
Michx. 


NOTE: also Emory Not. Mil. Reconn. 154. 1848. 


DST. 


LVL. 


: Arge, Boli, Braz, Cana, Colo, Ecua, ElSa, Guat, 
Hond, Mexi, Nica, Peru, USA, Urug. 
ACCPTC.: 1; cultivated. 


ALSO ACCPTD. BY: Abrams, L. (1923); Allen, C. M., C. 


SYN:: 


S. Reid & C. H. Doffitt (1999); Anderson, D. E. 
(1974); Beetle, A. A. (1987); Berendsohn, W. G. & 
A. E. Araniva de Gonzalez (1991); Bor, N. L. (1960); 
Brako, L. & J. L. Zarucchi (1993); Burkart, A. 
(1969); Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Darke, R. (1999); Davidse, G., M. 
Sousa S. & A. O. Chater (editores generales) (1994); 
Fernald, M. (1950); Foster, R. C. (1958); Gleason, H. 
A. & A. Cronquist (1968); Gould, F. W. (1980); 
Great Plains Flora Association (1986); Hickman, J. 
C. (1993); Hitchcock, A. S. (1927); Hitchcock, A. S. 
(1951): Hitchcock, C. L., A. Cronquist, M. Ownbey 
& J. W. Thompson (1969); Jorgensen, P. M. & S. 
Leén-Yanez (eds.) (1999); McClure, F. A. (1955); 
McVaugh, R. (1983); Molina R., A. (1975); Munz, P. 
A. & D. D. Keck (1959); Munz, P. A. (1974); Pohl, 
R. W. (1994); Radford, A. E., H. E. Ahles & C. R. 
Bell (1968); Renvoize, S. A. (1998); Reyes-Garcia, 
A. & M. Sousa S. (1997); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 
Saravia, E. F. (1996); Scoggan, H. J. (1978 [1979]); 
Shreve, F. & I. L. Wiggins (1964); Small, J. K. 
(1933); Tovar, O. (1993); Voss, E. G. (1972); 
Wunderlin, R. P. (1998). 

Andropogon curtipendulum (Michx.) Spreng. ex 
Steud.; Aristida secunda Rud. ex Roemer & Schultes; 
Atheropogon  affinis (Hook. f.)  E.  Fourn., 
Atheropogon  apludioides Muhl. ex  Willd.; 
Atheropogon  curtipendulus  (Michx.) E. Fourn.; 
Atheropogon medius  E. Fourn.; Atheropogon 
racemosus (Lag.) Roem. & Schult.; Bouteloua 
brasiliensis Ekman; Bouteloua curtipendula (Michx.) 
Torr.; Bouteloua curtipendula var. aristosa A. Gray; 


Bouteloua curtipendula var.curtipendula; Bouteloua 
meliciformis Hornem. ex Roem. & Schult; 
Bouteloua melicoides P. Beauv.,; Bouteloua racemosa 
Lag.; Bouteloua racemosa var. aristosa (A. Gray) S. 
Watson & J.M. Coult.; Chloris curtipendula Michx.; 
Chloris secundus  (Pursh) Eaton; Cynodon 
curtipendula (Michx.) Raspail; Cynodon melicoides 
(P. Beauv.) Raspail; Cynosurus secundus Pursh; 
Dinebra curtipendula (Michx.) P. Beauv.; Dinebra 
melicoides P. Beauv.; Dinebra secunda (Pursh) 
Roem. & Schult.; Eutriana affinis Hook. f.; Eutriana 
curtipendula (Michx.) Trin.; Heterostega 
curtipendula Schwein. ex Hook. f. 


Bouteloua curtipendula var.aristosa A. Gray, Manual (ed. 2) 


553. 1856. 


REPLACED NAME: Eutriana affinis Hook. f. 
= Bouteloua curtipendula (Michx.) Torr. 


Bouteloua curtipendula var. caespitosa Gould & Kapadia, 


DST. 


LVL 


Brittonia 16(2): 203, f. 43. 1964. T-Protol.: USA: 
Arizona: Cochise Co.: 3 mi E of Bisbee at ca. 1725 m, 
Gould 10021 [HT: TAES; IT: ARIZ, DS, GH, NY, 
TEX, UC, US]. T-Specim.: | of 1. USA: Arizona: 
Cochise Co.: 3 mi E of Bisbee at ca. 1725 m, 10 Aug 
1962, F.W. Gould 10021 (HT: TAES; IT: ARIZ, DS, 
GH, NY, TEX, UC, US-2473990]. 

: Arge, Boli, Ecua, Guat, Mexi, Nica, Peru, USA, 

Urug, Vene. 


.ACCPTC:: 1. 


ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 


SYN: 


(1977); Beetle, A. A. (1987); Brako, L. & J. L. 
Zarucchi (1993); Burkart, A. (1969); Correll, D. S. & 
M. C. Johnston (1970); Cronquist, A. A. H. 
Holmgren, N. H. Holmgren & J. L. Reveal (1977); 
Davidse, G., M. Sousa S. & A. QO. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. & 
R. Moran (1981); Gould, F. W. (1969); Gould, F. W. 
(1980); McVaugh, R. (1983); Pohl, R. W. (1994); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

Atheropogon acuminatus E. Fourn.; Atheropogon 
racemosus (Lag.) Roem. & Schult.; Boufeloua 
acuminata (E. Fourn.) Griffiths; Bouteloua pendula 
hort. ex Lag.; Bouteloua racemosa Lag. 


Bouteloua curtipendula (Michx.) Torr. var. curtipendula. 


DST. 
-ACCPTC.: 1. 


LVL 


: Cana, Mexi, USA. 


ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 


SYN.: 


(1987); Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J, Valdes-Reyna (2000); Gleason, H. A. 
& A. Cronquist (1991); Gould, F. W. (1980); 
McVaugh, R. (1983). 

Andropogon curtipendulum (Michx.) Spreng. ex 
Steud.; Atheropogon affinis (Hook. f.) E. Fourn.; 
Atheropogon  apludioides Muhl. ex  Willd.; 
Atheropogon curtipendulus (Michx.) E. Fourn.; 
Bouteloua curtipendula var. aristosa A. Gray; 
Bouteloua racemosa var. aristosa (A. Gray) S. 
Watson & J.M. Coult.; Chloris curtipendula Michx.; 
Cynodon curtipendula (Michx.) Raspail; Cynosurus 
secundus Pursh; Dinebra curtipendula (Michx.) P. 


24 


Catalogue of New World Grasses: II. 


Beauv.; Dinebra secunda (Pursh) Roem. & Schult.; 
Eutriana_ affinis Hook. f.; Eutriana curtipendula 
(Michx.) Trin.; Heterostega curtipendula Schwein. 
ex Hook. f.; Melica curtipendula Michx. ex Steud. 


Bouteloua curtipendula var. tenuis Gould & Kapadia, 
Brittonia 16(2): 201, f. 42. 1964. T-Protol.: Mexico: 
Zacatecas: 10 mi NW of Sombrerete (68 mi SE of 
Durango), 30 Oct 1959, Gould 9000 [HT: TAES; IT: 
TEX, UC, US]. T-Specim.: 1 of 1. Mexico: Zacatecas, 
30 Oct 1959, FW. Gould 9000c [HT: TAES; IT: TEX, 
UC, US-2551354]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. (1980); 
McVaugh, R. (1983). 


Bouteloua dactyloides (Nutt.) Columbus, Aliso 18(1): 63. 
1999. Basionym: Ses/leria dactyloides Nutt. 

DST.: Cana, Mexi, USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000). 

Anthephora axilliflora Steud.; Bouteloua mutica 
Griseb. ex E. Fourn.; Buchloe dactyloides (Nutt.) 
Engelm.; Bulbilis dactyloides (Nutt.) Raf. ex Kuntze; 
Calanthera dactyloides (Nutt.) Kunth ex Hook.; 
Casiostega dactyloides (Nutt.) E. Fourn.; Casiostega 
hookeri Rupr. ex E. Fourn.; Lasiostega humilis Rupr. 
ex Munro; Melica mexicana Link ex E. Fourn.; 
Sesleria dactyloides Nutt. 


SYN:: 


Bouteloua dimorpha Columbus, Aliso 18(1): 63. 1999. 
REPLACED NAME: Opizia stolonifera J. Presl. 
BLOCKING NAME: Bouteloua stolonifera Scribn. 

DST.: Cari, Hond, Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000). 

SYN.: Opizia stolonifera J. Presl. 


Bouteloua distans Swallen, Contr. U.S. Natl. Herb. 29(9): 
401. 1950. T-Protol.: Mexico: ca. 170 km N of Oaxaca 
City: 13 Dee 1945, JA. Jenkins & E. Hernandez X- 
808 [HT: US-1961989]. T-Specim.: | of 1. Mexico: 
Oaxaca, 13 Dec 1945, £. Hernandez-Xolocotzi & JA. 
Jenkins X-808 (HT: US-1961989; IT: CHAPA, US]. 

DST.: Mext. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. (1980). 


Bouteloua disticha (Kunth) Benth., J. Linn. Soc., Bot. 19: 
105. 1881. Basionym: Polyodon distichum Kunth. 
DST.: Arge, Cari, Colo, CoRi, Ecua, ElSa, Guat, Hond, 

Mexi, Nica, Pana, Peru, Vene. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A, 
(1987); Berendsohn, W. G. & A. E. Araniva de 
Gonzalez (1991); Brako, L. & J. L. Zarucchi (1993); 
Catasus Guerra, L. (1997); Davidse, G., M. Sousa S. 
& A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Gould, F. W. (1980); Hitchcock, A. S. 
(1909); Hitchcock, A. S. (1927); Hitchcock, A. S. 


(1936); Jorgensen, P. M. & S. Leén-Yanez (eds.) 
(1999); Lawesson, J. E., H. Adsersen & P. Bentley 
(1987); Macbride, J. F. (1936); McClure, F. A. 
(1955); McVaugh, R. (1983); Molina R., A. (1975); 
Pohl, R. W. (1980); Pohl, R. W. (1994); Porter, D. 
M. (1983); Reeder, J. R. & C. G. Reeder (1971); 
Tovar, O. (1993); Wiggins, I. L. & D. M. Porter 
(1971). 

Atheropogon distichus (Kunth) Spreng.; Bouteloua 
pilosa (Hook. f.) Benth. ex S. Watson; Bouteloua 
piurensis Pilg.; Eutriana gracilis Hook. f., Eutriana 
mucronata F. Aresch.; Eutriana pilosa Hook. f.; 
Eutriana polyodon Trin.; Leptochloa hirta Nees ex 
Steud.; Polyodon distichum Kunth. 


SYN:: 


Bouteloua diversispicula Columbus, Aliso 18(1): 63. 1999. 
REPLACED NAME: Cathestecum brevifolium Swallen. 
BLOCKING NAME: Bouteloua brevifolia Buckley. 

DST.: ElSa, Guat, Hond, Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000). 

SYN.: Cathestecum brevifolium Swallen, Cathestecum 
brevifolium var. brevifolium. 


Bouteloua elata Reeder & C. Reeder, Brittonia 15(3): 215, f. 
1-2. 1963. T-Protol.: Mexico: Colima: growing in 
large clumps on a rocky slope, common, 425 m, ca. 13 
mi SW of Ciudad Colima, 19 Sep 1963,/. R. Reeder 
& C. G. Reeder 2365 (HT: RM; IT: GH, US]. T- 
Specim.: | of 1. Mexico: Colima: ca. 13 mi SW of 
Ciudad Colima, 19 Sep 1953, JR. Reeder & C.G. 
Reeder 2356 [HT: RM; IT: GH, US-2382028], 
Collection date 1963 in protologue. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Breedlove, D. E. (1986); Espejo Serna, A., A. 
R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W. (1980); McVaugh, R. (1983). 

SYN.: Chondrosum elatum (Reeder & C. Reeder) Clayton. 


Bouteloua elatior Griseb., Fl. Brit. W. 1. 537. 1864. T-Protol.: 
Habitat Antigua, at Grace bay , S._ Johns, 
Wullschlaegel s.n. T-Specim.: | of 2. Leeward Islands: 
Antigua and Barbuda: Antigua I., Wullschlaegel 660 
[ST: GOET; IST: US-A865570 (fragm.)]. 2 of 2. 
Leeward Islands: Antigua, Wullschlaegel 619 [ST: 
GOET; IST: US (fragm.)]. 

= Bouteloua americana (L.) Scribn. 


Bouteloua eludens Griffiths, Contr. U.S. Natl. Herb. 14(3): 
401, t. 78-80A. 1912. T-Protol.; USA: Arizona: on 
southern exposures upon the northern slope of the 
Santa Rita Mts., 17 Sep 1904, Griffiths 7269 [HT: 
US]. T-Specim.: | of 1. USA: Arizona: Santa Rita 
Mountains, 17 Sep 1904, D. Griffiths 7269 [HT: US- 
81714; IT: GH], CS. 

DST.: Mexi, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1987); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W. (1980); Hitchcock, A. S. (1951); 
Reeder, J. R. & C. G. Reeder (1990); Shreve, F. & I. 
L. Wiggins (1964). 


Catalogue of New World Grasses: II. 25 


Bouteloua erecta (Vasey & Hack.) Columbus, Aliso 18(1): 
63. 1999. Basionym: Cathestecum erectum Vasey & 
Hack. 

DST.: ElSa, Guat, Hond, Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000). 

SYN.: Cathestecum erectum Vasey & Hack. 


Bouteloua eriopoda (Torr.) Torr., Pacif. Railr. Rep. 4(5): 
155. 1857. Basionym: Chondrosum eriopodum Torr. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Anonymous (1986); Bailey, L. H. & 
E. Z. Bailey (1976); Beetle, A. A. (1977); Beetle, A. 
A. (1987); Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. & 
R. Moran (1981); Gould, F. W. (1980); Great Plains 
Flora Association (1986); Hickman, J. C. (1993); 
Hitchcock, A. S. (1951); Munz, P. A. & D. D. Keck 
(1959); Munz, P. A. (1974); Shreve, F. & I. L. 
Wiggins (1964). 

Bouteloua brevifolia Buckley; Chondrosium 
eriopodum Torr., Chondrosum eriopodum Torr. 


SYN:: 


Bouteloua eriopoda var. eriostachya Swallen, Proc. Biol. 
Soc, Wash. 56: 81. 1943. T-Protol.: Mexico: Coahuila: 


vicinity of Aguaje de Pajarito, canyon at west end of 


Sierra de la Fragua, 2-3 km N of Puerto Colorado, 
frequent on rocky flats, 1-3 Sep 1941,/. M. Johnston 
8718 [HT: US-154692; IT: GH, LL]. T-Specim.: | of 
1. Mexico: Coahuila: cerca de Aguaje de Pajarito, 1-3 
Sep 1941, 1M. Johnston 8718 (HT: US-154692; IT: 
GH, LL, US-2209363]. 

= Bouteloua eriostachya (Swallen) Reeder. 


Bouteloua eriostachya (Swallen) Reeder, Bull. Torrey Bot. 
Club 94: 7. 1967. Basionym: Bouteloua eriopoda var. 
eriostachya Swallen. 

DST.: Mexi. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. (1980). 

SYN.: Bouteloua eriopoda var. eriostachya Swallen; 
Chondrosum eriostachyum (Swallen) Clayton. 


Bouteloua filiformis (E. Fourn.) Griffiths, Contr. U.S. Natl. 
Herb. 14(3): 413, t. 82A, 83. 1912. Basionym: 
Atheropogon filiformis E. Fourn. 

= Bouteloua repens (Kunth) Scribn. 


Bouteloua foenea (Torr.) Torr. ex S. Watson, Cat. Pl. 18. 
1874. Basionym: Chondrosum foeneum Torr. 
= Bouteloua hirsuta var. hirsuta. 


Bouteloua gentryi Gould, Leafl. W. Bot. 5(12): 199. 1949. T- 
Specim.: | of 1. Mexico: Sinaloa: Imala, Gentry 5000 
{IT: ARIZ]. 
NOTE: as "Gentry1". 
= Bouteloua parryi var. gentryi (Gould) Gould. 


Bouteloua glandulosa (Cerv.) Swallen, N. Amer. Fl. 17(8): 
621. 1939. Basionym: Erucaria glandulosa Cerv. 
= Bouteloua hirsuta var. glandulosa (Cerv.) Gould. 


Bouteloua gracilis (Kunth) Lag. ex Steud., Nomencl. Bot. 
(ed. 2) 1: 219. 1840. Basionym: Chondrosum gracile 
Kunth. 

NOTE: comb. not made by Steud. 

CORRECT NAME: Bouwteloua gracilis (Kunth) Lag. ex 
Griffiths. 

= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Bouteloua gracilis Vasey, Rep. U.S. Geogr. Surv., Wheeler 
6: 287. 1878. T-Protol.: USA: Arizona: Riley's Well, 
1874, Rothrock 701 [HT: US]. T-Specim.: | of 1. 
USA: Arizona, Riley's Well, 1874, Wheeler's 
Expedition, Dr. J.R. Rothrock 701 (HT: US-879645]. 
NOTE: nom. rej. prop. 1479; cites "Hook." ex Vasey. 
= Bouteloua aristidoides (Kunth) Griseb. 


Bouteloua gracilis (Kunth) Lag. ex Griffiths, Contr. U.S. 
Natl. Herb. 14: 375. 1912. Basionym: Chondrosum 
gracile Kunth. 

NOTE: nom. cons. prop. 1479, by Gandhi et al., Taxon 
50(2): 573-175 (2001). 

E. NAME: Bouteloua gracilis Vasey. 

DST.: Arge, Cana, Mexi, USA. 

ALSO ACCPTD. BY: Darke, R. (1999); Hickman, J. C. 

(1993). 

Actinochloa gracilis (Kunth) Willd. ex Roem. & 

Schult.; Atheropogon gracilis (Kunth) Spreng.; 

Atheropogon oligostachyus Nutt.; Bouteloua gracilis 

(Kunth) Lag. ex Steud.; Bouteloua gracilis fo.pallida 

(Scribn. ex Beal) B. Boivin; Bouteloua gracilis var. 

gracilis, Bouteloua gracilis var. major (Vasey ex 

L.H. Dewey) Beetle; Bouteloua gracilis var. stricta 

(Vasey) Hitchc.; Bouteloua major Vasey; Bouteloua 

oligostachya (Nutt.) Torr. ex A. Gray; Bouteloua 

oligostachya var. major Vasey ex L.H. Dewey; 

Bouteloua oligostachya var. pallida Scribn. ex Beal; 

Bouteloua stricta’ Vasey; Chondrosium — gracile 

Kunth; Chondrosum gracile Kunth; Chondrosum 

gracile var. gracile, Chondrosum gracile var. 

polystachyum Nees; Chondrosum  oligostachyum 

(Nutt.) Torr.; Eutriana gracilis (Kunth) Trin.; 

Eutriana oligostachya (Nutt.) Kunth. 


SYN:: 


Bouteloua gracilis (Kunth) Lag. ex Steud. var. gracilis. 
= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Bouteloua gracilis var. major (Vasey ex L.H. Dewey) Beetle, 
Phytologia 52(1): 11. 1982. Basionym: Bouteloua 
oligostachya var. major Vasey ex L.H. Dewey. 

= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Bouteloua gracilis fo. pallida (Scribn. ex Beal) B. Boivin, 
Naturaliste Canad. 94(4): 521. 1967. Basionym: 
Bouteloua oligostachya var. pallida Scribn. ex Beal. 

= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Bouteloua gracilis var. stricta (Vasey) Hitche., J. Wash. 
Acad. Sci. 23(10): 454. 1933. Basionym: Bouteloua 
stricta Vasey. 

= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Bouteloua havardii Vasey ex S. Watson, Proc. Amer, Acad. 
Arts 18: 179. 1883. T-Protol.: USA: Limpio Mts., 
Western Texas, 1849, V. Havard s.n. (HT: US; IT: 
GH]. T-Specim.: | of 1. USA: Texas, 1881, V. Havard 
53 (HT: US-81723], CS. 

= Bouteloua chondrosioides (Kunth) Benth. ex S. Watson. 


26 Catalogue of New World Grasses: II. 


Bouteloua heterostega (Trin.) Griffiths, Contr. U.S. Natl. 
Herb, 14(3): 414, f. 59. 1912. Basionym: Eutriana 
heterostega Trin. 

= Bouteloua repens (Kunth) Scribn. 


Bouteloua hirsuta Lag., Varied. Ci. 2(4): 141. 1805. T- 
Protol.: [MA]. T-Specim.: | of 1. Cult. Madrid Bot. 
Gard. from seed collected in Mexico by L. Nee, 
Lagasca s.n, [LT: MA-245716; ILT: BM (possible ex 
herb. Roemer), M-7433 LE-TRIN-2125.6 (fragm. ex 
M and drawings)], LT designated by Griffiths, Contr. 
U.S. Natl. Herb. 14: 372 (1912); also cited by Gould, 
Ann. Missouri Bot. Gard. 66: 7??? (1980), 

DST.: Cana, Guat, Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Abrams, L. (1923); Allen, C. M., C. 
S. Reid & C. H. Doffitt (1999); Bailey, L. H. & E. Z. 
Bailey (1976); Beetle, A. A. (1977); Beetle, A. A. 
(1987); Breedlove, D. E. (1986); Correll, D. S. & M. 
C. Johnston (1970); Cowan, C. P. (1983); Cronquist, 
A., A. H. Holmgren, N. H. Holmgren & J. L. Reveal 
(1977); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Fernald, M. (1950); Gleason, 
H. A. & A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Gould, F. W. & R. Moran (1981); 
Gould, F. W. (1980); Great Plains Flora Association 
(1986); Hitchcock, A. S. (1951); Long, R. W. & O. 
Lakela (1971); McVaugh, R. (1983); Munz, P. A. & 
D. D. Keck (1959); Shreve, F. & I. L. Wiggins 
(1964); Small, J. K. (1933); Veldkamp, J. F. (2001); 
Wunderlin, R. P. (1998). 

Actinochloa_ hirsuta (Lag.) Roem. & Schult.; 
Atheropogon hirtus (Kunth) Spreng.; Atheropogon 
papillosus Engelm.; Bouteloua aschenborniana 
Griseb. ex E, Fourn.; Bouteloua foenea (Torr.) Torr. 
ex S. Watson; Bouteloua hirsuta var. minor Vasey; 
Bouteloua hirsuta var. palmeri Vasey ex Beal; 
Bouteloua hirta (Kunth) Scribn.; Bouteloua hirta 
hort. ex Lag.; Bouteloua hirta Scribn. ex Vasey; 
Bouteloua hirta var. minor Vasey ex L.H. Dewey; 
Bouteloua palmeri Vasey; Chondrosium hirsutum 
(Lag.) Sweet; Chondrosum aschenbornianum Nees; 
Chondrosum drummondii E. Fourn.; Chondrosum 
foeneum Torr.; Chondrosum hirsutum (Lag.) Sweet; 
Chondrosum hirtum Kunth; Chondrosum papillosum 
(Engelm.) Torr.; Eutriana hirta (Kunth) Trin. 


SYN.: 


Bouteloua hirsuta var. glandulosa (Cerv.) Gould, J. Arnold 
Arbor. 60(2): 320. 1979. Basionym: Erucaria 
glandulosa Cerv. 

DST.: Guat, Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W. & R. Moran (1981); Gould, F. W. 
(1980). 

Bouteloua glandulosa (Cerv.) Swallen,; Bouteloua 
hirticulmis Scribn.; Chondrosum hirsutum var. 
glandulosum (Cerv.) R.W. Pohl; Erucaria glandulosa 
Cerv.; Erucaria hirsuta Cerv. 


SYN.: 


Bouteloua hirsuta Lag. subsp. hirsuta. 
= Bouteloua hirsuta var. hirsuta. 


Bouteloua hirsuta Lag. subsp. hirsuta. 
ALSO ACCPTD. BY: Wipff, J. K. & S. D. Jones (1996). 


Bouteloua hirsuta Lag. var. hirsuta. 


DST.: Cana, Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1987); Gould, F. W. & 
R. Moran (1981); Gould, F. W. (1980). 

SYN.: Actinochloa hirsuta (Lag.) Roem. & Schult.; 
Atheropogon hirtus (Kunth) Spreng.; Atheropogon 
papillosus Engelm.; Bouteloua aschenborniana 
Griseb. ex E. Fourn.; Bouteloua foenea (Torr.) Torr. 
ex S. Watson; Bouteloua hirsuta subsp. hirsuta: 
Bouteloua hirsuta var. minor Vasey; Bouteloua 
hirsuta var. palmeri Vasey ex Beal; Bouteloua hirta 
(Kunth) Scribn.; Bouteloua hirta var. minor Vasey ex 
L.H. Dewey; Bouteloua pappilosa (Engelm.) Torr.: 
Chondrosium hirsutum (Lag.) Sweet; Chondrosium 
hirtum Kunth; Chondrosum aschenbornianum Nees; 
Chondrosum drummondii E. Fourn.; Chondrosum 

Joeneum Tort.; Chondrosum hirsutum (Lag.) Sweet; 

Chondrosum hirsutum var. hirsutum, Chondrosum 

hirtum Kunth; Chondrosum papillosum (Engelm.) 

Torr. 


Bouteloua hirsuta var. minor Vasey, U.S.D.A. Div. Bot. Bull. 
12(1): t. 39, f. 2. 1890. T-Protol.: [USA: Texas], 
[Reverchon 1153], drawn. 

NOTE: nom. seminud. 
= Bouteloua hirsuta Lag. 


Bouteloua hirsuta var. palmeri Vasey ex Beal, Grass. N. 
Amer. 2: 417. 1896. T-Protol.: cult.: seed collected in 
Mexico, Palmer s.n. T-Specim.: 1 of 1. Mexico: 
cultivated from seed collected in Mexico, 1886, 
Palmer s.n. {HT: US}. 

= Bouteloua hirsuta Lag. 


Bouteloua hirsuta subsp. pectinata (Feath.) Wipff & S.D. 
Jones, Sida 17(1): 109. 1996. Basionym: Bouteloua 
pectinata Feath. 

DST.: USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Wipff, J. K. & S. D. Jones (1996). 

SYN.: Bouteloua hirsuta var. pectinata (Feath.) Cory; 
Bouteloua hirta var. major (Vasey) Vasey ex L.H. 
Dewey; Bouteloua pectinata Feath.; Chondrosum 
pectinatum (Feath.) Clayton. 


Bouteloua hirsuta var. pectinata (Feath.) Cory, Rhodora 
38(455): 405. 1936. Basionym: Bouteloua pectinata 
Feath. 

= Bouteloua hirsuta subsp. pectinata (Feath.) Wipfft & 
S.D. Jones. 


Bouteloua hirta hort. ex Lag., Varied, Ci, 2(4): 141. 1805. 
NOTE: nom. inval., as syn. of Bouteloua hirsuta Lag. 
= Bouteloua hirsuta Lag. 


Bouteloua hirta Scribn. ex Vasey, Proc. Amer. Acad. Arts 

xxil: 531. 1887. 

NOTE: from Index 
illegitimate. 

= Bouteloua hirsuta Lag. 


Kewensis; apparently invalid or 


Bouteloua hirta (Kunth) Scribn., Contr. U.S. Natl. Herb. 2(3): 
531. 1894. Basionym: Chondrosum hirtum Kunth. 
= Bouteloua hirsuta var. hirsuta. 


Bouteloua hirta var. major (Vasey) Vasey ex L.H. Dewey, 
Contr. U.S. Natl. Herb. 2(3): 531. 1894. Basionym: 
Bouteloua hirsuta var. major Vasey. 


Catalogue of New World Grasses: II. 27 


= Bouteloua hirsuta subsp. pectinata (Feath.) Wipff & 
S.D. Jones. 


Bouteloua hirta var. minor Vasey ex L.H. Dewey, Contr. 
U.S. Natl. Herb. 2(3): 531. 1894. T-Protol.: USA: 
Central Texas. T-Specim.: | of 1. USA: Texas: central 
Texas [T]. 

= Bouteloua hirsuta Lag. 


Bouteloua hirticulmis Scribn., Circ. Div. Agrostol. U.S.D.A. 
30: 4. 1901. T-Protol.: Mexico: Sierra de San 
Francisquito Mts., Lower [Baja] California, 29 Sep 
1899, 2 syntypes. T-Specim.: 1 of 2. Mexico: Baja 
California Sur: Sierra de San Franciso, 29 Sep 1899, 
TS. Brandegee 11 [LT: IST: US-81722], LT 
designated by ?; cited (as type) by McVaugh, Fl. 
Novo-Galic. 14; 89 (1983). 2 of 2. Mexico: Lower 
California, Sierra de San Francisquito Mounttains, 
October 18, 1890, Brandegee 30 [ST: UC]. 

= Bouteloua hirsuta var. glandulosa (Cerv.) Gould. 


Bouteloua humboldtiana Griseb., Mem. Amer. Acad. Arts, 
n.s., 8: 532. 1862. 
REPLACED NAME: Heterosteca juncifolia Desv. 
BLOCKING NAME: Bouteloua juncifolia Lag. 
= Bouteloua repens (Kunth) Scribn. 


Bouteloua humilis (P. Beauv. ex Kunth) Hieron., Bol. Acad. 
Nac. Ci. 4: 495. 1882. Basionym: Chondrosum humile 
P. Beauv. ex Kunth. 
= Bouteloua simplex Lag. 


Bouteloua johnstonii Swallen, Proc. Biol. Soc. Wash. 56: 
79. 1943. T-Protol.: Mexico: Coahuila: S end of 
Canada Oscuro near Tanque La Luz: 27 Sep 1942, 
Johnston 8491 [HT: NA-154624; IT: GH]. T-Specim.: 
1 of 1. Mexico: Coahuila: S de Canada Oscuro, near 
Tanque La Luz, Johnston 849] [HT: NA; IT: GH, US- 
1817790]. 

DST.: Mexi. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. (1980). 


Bouteloua juncea (Desv. ex P. Beauv.) Hitche., Contr. U.S. 
Natl. Herb. 17(3): 343. 1913. Basionym: Triathera 
jJuncea Desvy. ex P. Beauv. 

DST.: Cari, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Catasus Guerra, L. (1997); Gould, F. W. 
(1980); Hitchcock, A. S. (1936); Liogier, H. & L. 
Martorell (1982). 

Atheropogon domingensis Spreng.; Eutriana juncea 
(Desv. ex P. Beauv.) Trin.; Eutriana ledebourii Trin.; 
Triaena juncea (Desv. ex P. Beauv.) Griffiths; 
Triathera juncea Desv. ex P. Beauv. 


SYN.: 


Bouteloua juncifolia Lag., Gen. Sp. Pl. 5. 1816. T-Protol.: 
Habit. in Imperio Mex. 
SYN.: Actinochloa juncifolia (Lag.) Roem. & Schult.; 
Eutriana lagascae Kunth. 
= Status not yet determined. 


Bouteloua karwinskii (E. Fourn.) Griffiths, Contr. U.S. Natl. 
Herb. 14(3); 394, f. 47, t. 76. 1912. Basionym: 
Chondrosum karwinskii E. Fourn. 

DST.: Mexi. 


LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. (1980). 

SYN.: Chondrosium karwinskii E. Fourn.; Chondrosum 
karwinskii E. Fourn. 


Bouteloua kayi Warnock, Field & Lab. 23(1): 15-16, f. s.n. 
1955. T-Protol.: USA: Texas: southern Brewster Co.: 
infrequent perennial in limestone crevices on lower 
portion of Maravillas Creek, Overton Ranch, elev. 
2200 ft, 4 Aug 1954, Ll. Kay s.n. [HT: Sul Ross State 
College; IT: SMU]. T-Specim.: | of 1. USA: Texas: 
Brewster Co.: near Rio Grande River, in limestone, 4 
Aug 1954, Kay K-/ [HT: SRSC: IT: SMU, TEX]. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Gould, F. W. (1980). 

SYN.: Chondrosum kayi (Warnock) Clayton. 


Bouteloua latifolia Swallen, N. Amer. Fl. 17(8): 631. 1939. 
T-Protol.: Mexico: on hills near Yautepec, Morelos, 
1200 m alt., 22 Oct 1902, Pringle 11217 (US-460593]. 
T-Specim.: | of 1. Mexico: Morelos, 22 Oct 1902, 
C.G. Pringle 11217 [HT: US-460593; IT: CM, GH, 
MO, TAES]. 

= Bouteloua media (E. Fourn.) Gould & Kapadia. 


Bouteloua litigiosa Lag., Gen. Sp. Pl. 5. 1816. T-Protol.: 
[West Indies]: "Hab. in Jamaica.", Swart. s.n. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: 4ristida americana L. 
= Bouteloua americana (L.) Scribn. 


Bouteloua longiseta Gould, Brittonia 21: 271. 1969. T- 
Protol.: Mexico: Chiapas: 9 km SW of Cintalapa, 8 
Oct 1968, Gould 12759 [HT: TAES; IT: GH, TEX]. T- 
Specim.: | of 1. Mexico: Chiapas, 8 Oct 1968, FLW. 
Gould 12759 [HT: TAES; IT: GH, TEX, US- 
2551471]. 
= Bouteloua alamosana Vasey. 


Bouteloua lophostachya Griseb., Abh. Konigl. Ges. Wiss. 
Gottingen 24: 302. 1879. T-Protol.: Argentina: in 
confinio Prov. Salta, syntypes?. T-Specim.: | of 1. 
Argentina: Tucuman, G. //ieronymus 97/1 [HT: 
GOET(fragm., US-A865560)], CS\Ex griseb. herb. 

= Neobouteloua lophostachya (Griseb.) Gould. 


Bouteloua major Vasey, Bull. Torrey Bot. Club 14: 9. 1887. 
T-Protol.: plant grown from seed collected in Mexico 
by Palmer, Palmer s.n., Name only, for a plant grown 
from seed collected in Mexico by Palmer. 

NOTE: nom. nud. 
= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Bouteloua media (E. Fourn.) Gould & Kapadia, Brittonia 
16(2): 196, f. 35. 1964. Basionym: Atheropogon 
medius E. Fourn. 

DST.: Arge, Braz, CoRi, Ecua, ElSa, Guat, Hond, Mexi, 
Nica, Pana, Para, Peru, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Brako, L. & J. L. Zarucchi (1993); 
Breedlove, D. E. (1986); Davidse, G., M. Sousa S. & 
A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 


28 Catalogue of New World Grasses: II. 


(2000); Gould, F. W. (1980); Jorgensen, P. M. & S. 
Leén-Yanez (eds.) (1999); McVaugh, R. (1983); 
Pohl, R. W. (1980); Pohl, R. W. (1994); Reyes- 
Garcia, A. & M. Sousa S. (1997); Zuloaga, F. O., E. 
G. Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

Atheropogon medius E. Fourn.; Bouteloua 
brasiliensis Ekman; Bouteloua latifolia Swallen; 
Bouteloua pringleiScribn. 


SYN.: 


Bouteloua megapotamica (Spreng.) Kuntze, Revis. Gen. PI. 
3(2):; 341. =: 1898. Basionym: = Pappophorum 
megapotamicum Spreng. 

NOTE: Index of Grass Species 3(3): 341. 

DST.: Arge, Boli, Braz, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Burkart, A. (1969); Fabris, H. A. 
(1970); Foster, R. C. (1958); Gould, F. W. (1980); 
Hitchcock, A. S. (1927); Nicora, E. G., M. E. De 
Paula, A. M. Faggi, M. del C. Mariano, A. M. Miante 
A., L. R. Parodi, C. A. Petetin, F. A. Roig & Z. 
Rugolo de Agrasar (1978); Renvoize, S. A. (1998); 
Rosengurtt, B., B. R. Arrillaga de Maffei & P. 
Izaguirre de Artucio (1970); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

Bouteloua multiseta (Nees) Griseb.; Bouteloua 
multiseta var. pallida Hack.; Eutriana multiseta 
Nees; Pappophorum eutrianoides ‘Trin. ex Nees; 
Pappophorum megapotamicum Spreng. 


SYN.: 


Bouteloua meliciformis Hornem. ex Roem. & Schult., Syst. 
Veg. 2: 414. 1817. 
NOTE: nom. inval., as syn. of Atheropogon apludioides 
Muhl. 
= Bouteloua curtipendula (Michx.) Torr. 


Bouteloua melicoides P. Beauv., Ess. Agrostogr. 40, 155, t. 9, 
f. 6. 1812. 

NOTE: based on "Boutel. melicoides Horn.", evidently an 
error for the invalidly published name: Bouteloua 
melicaeformis. 

= Bouteloua curtipendula (Michx.) Torr. 


Bouteloua mexicana (Scribn.) Columbus, Aliso 18(1): 63. 
1999, Basionym: Fourniera mexicana Scribn. 

DST.: ElSa, Guat, Mexi. 

LVL. ACCPTC.:: I. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000). 

SYN.: Fourniera mexicana Scribn.; Soderstromia mexicana 
(Scribn.) C.V. Morton. 


Bouteloua micrantha Scribn. & Merr., Circ. Div. Agrostol. 
U.S.D.A. 32: 8. 1901. T-Protol.: Collected at Fort 
Lowell, Arizona, Sep 1900, D. Griffiths 1556. Y- 
Specim.: | of 1. USA: Arizona: Pima Co.: Fort 
Lowell, Sep 1900, D. Griffiths 1556 [HT: US-81724]. 

= Bouteloua barbata Lag. 


Bouteloua microstachya (E. Fourn.) L.H. Dewey, Contr. U.S. 
Natl. Herb. 2(3): 531. 1894. Basionym: Chondrosum 
microstachyum E. Fourn. 

= Bouteloua barbata Lag. 


Bouteloua multifidum (Griffiths) Columbus, Aliso 18(1): 
63. 1999. Basionym: Cathestecum — multifidum 
Griffiths. 


DST.: Mexi. 

LVL. ACCPTC.: |. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000). 

SYN.: Cathestecum multifidum Griffiths; Griffithsochloa 
multifida (Griffiths) G.J. Pierce. 


Bouteloua multiseta (Nees) Griseb., Abh. Konigl. Ges. Wiss. 
Gottingen 19: 303. 1879. Basionym: Eutriana 
multiseta Nees. 

= Bouteloua megapotamica (Spreng.) Kuntze. 


Bouteloua multiseta var. pallida Hack., Anales Mus. Nac. 
Buenos Aires 13: 491. 1906. T-Protol.: Argentina: 
Altos del cementerio de San Geronimo de Cordoba, 
Stuckert hb. 5268 [T: Hb. Arg.]. T-Specim.: 1 of 1. 
Altos del cemetario. San Geronimo de Cordoba, 
Argentina, 1898, Stuckert herb arg 5268 [IT: US- 
72695 (fragm. ex W)]. 

= Bouteloua megapotamica (Spreng.) Kuntze. 


Bouteloua mutica Griseb. ex E. Fourn., Bull. Soc. Roy. Bot. 
Belgique 15: 471. 1876. T-Protol.: Mexico, Schaffner 
/34, |staminate plant]. Griseb. in sched (planta 
mascula). T-Specim.: | of 1. Mexico, JG. Schaffner 
134 [OM: US-3048117]. 
NOTE: nom. inval., as syn. of Buchloe dactyloides (Nutt.) 
Engelm. 
= Bouteloua dactyloides (Nutt.) Columbus. 


Bouteloua nana Griseb., Abh. Konig]. Ges. Wiss. Gottingen 
24: 303. 1879. T-Protol.: Argentina: Cérdoba und 
Santiago del Estero, pr. Gauzapampa, Hieronymus 
721. T-Specim.: 1 of 1. Argentina: Cordoba / Santiago 
del Estero: Guazapampa, G. Hieronymus 72] [HT: 
GOET; IT: US-A865561 (fragm. ex GOET)]. 

= Neobouteloua lophostachya (Griseb.) Gould. 


Bouteloua nervata Swallen, N. Amer. Fl. 17(8): 626. 1939. 
T-Protol.: Mexico: on rocky hill, Pachuca, Hidalgo, 
8000 ft alt., 6-7 Sep 1910, dry mesas and rocky hills, 
Hitchcock 6734 [HT: US-883931]. T-Specim.: | of 1. 
Mexico: Hidalgo, 6 Sep 1910-7 Sep 1910, 4.8. 
Hitchcock 6734 [HT: US-883931]. 

DST.: Mexi. 

LVL. ACCPTC.: |. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000). 

SYN.: Buchlomimus nervatus (Swallen) Reeder, C. Reeder 
& Rzed. 


Bouteloua oligostachya (Nutt.) Torr. ex A. Gray, Manual (ed. 
2) 553. 1856. Basionym: Atheropogon oligostachyus 
Nutt. 
= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Bouteloua oligostachya var. major Vasey ex L.H. Dewey, 
Contr. U.S. Natl. Herb. 2(3): 531. 1894. T-Protol.: 
USA: Texas to Arizona, //.G. Lemon 427], with name 
in Vasey script. Note by A.C. [cult. from Mex. seed 
from Dr. Palmer]. T-Specim.: 1 of 2. USA: Arizona, 
1882, .G. Lemmon 427 (HT: US-883201], CS. 2 of 2. 

NOTE: check Bouteloua major Vasey. 
= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Bouteloua oligostachya var. pallida Scribn. ex Beal, Grass. 
N. Amer. 2: 418. 1896. T-Protol.: Mexico, Pringle 
407, T-Specim.: 1 of 1. Mexico: Chihuahua, 23 Aug 
1885, C.G. Pringle 407 [HT: US-81708]. 


Catalogue of New World Grasses: II. 29 


= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Bouteloua oligostachya var. ramosa (Scribn. ex Vasey) 
Scribn. ex Beal, Grass. N. Amer. 2: 418. 1896. 
Basionym: Bouteloua ramosa Scribn. ex Vasey. 

= Bouteloua ramosa Scribn, ex Vasey. 


Bouteloua ovata Lag., Gen. Sp. Pl. 5. 1816. T-Protol.: Habit. 
in Imper. Mex. 
= Bouteloua chondrosioides (Kunth) Benth. ex S. Watson. 


Bouteloua palmeri Vasey, Bull. Torrey Bot. Club 14: 9. 
1887. T-Protol.: Mexico, Palmer s.n. 
NOTE: nom. nud.; later described as B. hirsuta var. palmeri 
Vasey ex Beal. 
= Bouteloua hirsuta Lag. 


Bouteloua pappilosa (Engelm.) Torr., Explor. Red River 
Louisiana 300. 1853. Basionym: Atheropogon 
papillosus Engelm. 

= Bouteloua hirsuta var. hirsuta. 


Bouteloua parryi (E. Fourn.) Griffiths, Contr. U.S. Natl. 
Herb. 14(3): 381, f. 37, t. 73. 1912. Basionym: 
Chondrosum parryi E, Fourn. 

DST.: Mexi, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Beetle, A. A. (1987); Gould, F. W. (1980); 
Hitchcock, A. S. (1951); Shreve, F. & I. L. Wiggins 
(1964); Viegi, L. & G. Renzoni. (1981). 

SYN.: Bouteloua vestita (S. Watson) Scribn. ex L.H. 
Dewey; Chondrosium parryi E. Fourn. 


Bouteloua parryi var. gentryi (Gould) Gould, Ann. 
Missouri Bot. Gard. 66: 408. 1979[1980]. Basionym: 
Bouteloua gentryi Gould. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1987); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W. (1980). 

SYN.: Bouteloua gentryiGould. 


Bouteloua parryi (E. Fourn.) Griffiths var. parryi. 

DST.: Mexi, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1987); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W. (1980). 

Bouteloua_ polystachya S. Watson; Bouteloua 
polystachya var. vestita S. Watson, Bouteloua vestita 
(S. Watson) Scribn. ex L.H. Dewey; Chondrosum 
parryi E. Fourn., 


SYN.: 


Bouteloua pectinata Feath., Bot. Gaz. 91: 103, f. 1-4. 1931. 

T-Protol.: USA: Wichita Mt. near Fort Sill, Oklahoma, 

17 Aug 1929, B. English 71 (HT: US-1445571]. T- 

Specim.: | of 1. USA: Oklahoma: Comanche Co.: near 

Fort Sill, 17 Aug. 1929, B. English 71 (HT: US- 
1445571]. 

= Bouteloua hirsuta subsp. pectinata (Feath.) Wipff & 

S.D. Jones. 


Bouteloua pedicellata Swallen, N. Amer. Fl. 17(8): 627. 
1939. T-Protol.: Mexico: rocky hill, Chalchicomula, 
Puebla, 9000 ft alt., 19 Aug 1910,4.8. Hitchcock 6302 
[HT: US-883950]. T-Specim.: | of 1. Mexico: Puebla, 


19 Aug 1910, A.S. Hitchcock 6302 [HT: US-883950; 
IT: US]. 
DST.: Mexi. 
LVL. ACCPTC:: 1. 
ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. L6pez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. (1980). 


Bouteloua pendula hort. ex Lag., Waried. Ci. 2(4): 141. 1805. 
T-Specim.: | of 1. Cultivated from seed collected in 
Mexico [T]. 

NOTE: nom. inval., as syn. of Bouteloua racemosa Lag. 
= Bouteloua curtipendula var. caespitosa Gould & 
Kapadia. 


Bouteloua pilosa (Hook. f.) Benth. ex S. Watson, Proc. 
Amer. Acad. Arts 18: 179. 1883. Basionym: Eutriana 
pilosa Hook. f. 

= Bouteloua disticha (Kunth) Benth. 


Bouteloua piurensis Pilg., Repert. Spec. Nov. Regni Veg. 
17(19-30): 447. 1921. T-Protol.: Peru: Prov. Piura: 
zwischen Morropon und Salitral, Mar 1912, 
Weberbauer 5961. T-Specim.: | of 1. Peru: Piura, Mar 
1912, A. Weberbauer 5967 [HT: B; IT: US-81712]. 

= Bouteloua disticha (Kunth) Benth. 


Bouteloua polymorpha (E. Fourn.) Columbus, Aliso 18(1): 
63. 1999. Basionym: Atheropogon polymorphus E. 
Fourn, 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000). 

SYN.: Atheropogon polymorphus E. Fourn.; Pentarrhaphis 
polymorpha (E. Fourn.) Griffiths. 


Bouteloua polystachya (Benth.) Torr., Pacif. Railr. Rep. 5(2): 
366, t. 10. 1857. Basionym: Chondrosum 
polystachyum Benth. 

NOTE: error for Bouteloua. 
= Bouteloua barbata Lag. 


Bouteloua polystachya §. Watson, Proc. Amer. Acad. Arts 
18: 177. 1883. T-Specim.: | of 1. Mexico: Sierra 
Madre, 40 mi S of Saltillo, Palmer 1357 (HT: GH]. 

= Bouteloua parryi var. parryi. 


Bouteloua polystachya var. major Vasey, Rep. U.S. Geogr. 
Surv., Wheeler 6: 287. 1878. T-Protol.: USA: Arizona: 
Sanoita Valley, 1874, Rothrock 691, 
[Sanoyta/Sonoyta]. T-Specim.: | of 2. USA: Arizona: 
Santa Cruz Co: Sanoita Vallet, 1874, /.7. Rothrock 
691 (HT: US-81711; IT: GH], CS. 2 of 2. 

= Bouteloua barbata var. rothrockii (Vasey) Gould. 


Bouteloua polystachya Gould var. polystachya. 
= Bouteloua barbata Lag. 


Bouteloua polystachya var. vestita 8. Watson, Proc. Amer. 
Acad. Arts 18: 177. 1883. T-Protol.: Mexico: Sierra 
Madre S. of Saltillo, 1880, Palmer 1357. T-Specim.: 1 
of 1. Mexico: Sierra Madre S of Saltillo, Palmer 1357 
{HT: GH]. 

= Bouteloua parryi var. parryi. 


Bouteloua porphyrantha C. Wright, Anales Acad. Ci. Med. 
Habana 8: 201. 1871. T-Protol.: Cuba, Wright 739 pro 
parte 734, 3816, [reprint Fl. Cubana 192. 1873]. T- 
Specim.: 1 of 2. Cuba: Oriente, Wright 739 [ST: 


30 Catalogue of New World Grasses: II. 


GOET; IST: G, GH, P, US-A865563 (fragm.)]. 2 of 2. 
Cuba, 1856, C. Wright 734 [ST: NY-70988; IST: US- 
2830957 (fragm.)]. 

= Bouteloua americana (L.) Scribn. 


Bouteloua pringlei Scribn., Circ. Div. Agrostol. U.S.D.A. 30: 
4. 1901. T-Protol.: Mexico: Mountains of Iguala, 1900, 
C.G. Pringle 8374 [HT: US]. T-Specim.: 1 of 1. 
Mexico: Guerrero: limestone ledges of mountains 
above Iguala, elev. 4000 ft, 24 Oct 1900,C.G. Pringle 
8374 [HT: US-382009; IT: CM, GH, LL, MO- 
2876580, NY, TAES]. 

= Bouteloua media (E. Fourn.) Gould & Kapadia. 


Bouteloua procumbens (P. Durand) Griffiths, Contr. U.S. 
Natl. Herb. 14(3): 364, f. 27. 1912. Basionym: Chloris 
procumbens P. Durand. 

NOTE: nom. superfl. as aplied; cites B. prostrata Lag. in 
syn., and that epithet has priority. 
= Bouteloua simplex Lag. 


Bouteloua prostrata hort. ex Lag., Varied. Ci. 2(4): 141. 
1805. T-Protol.: cult.[MA]. T-Specim.: | of 2. cult.: 
from seed from Mexico, Lagasca s.n. (HT: MA- 
230004 (photocopy, L); IT: M-7454, LE-TRIN-2126.3 
(upper left-hand corner)], Originally collected from 
Mexico by L. Nee, then cultivated in Madrid. 2 of 2. 
Ex Philippinis Insulis [error for Mexico], L. Nee [OM: 
MA; US-A865567 (fragm. ex MA & photo)]. 

NOTE: nom. seminud.; emend. Gen. & Sp. Nov. 5 (1816). 
= Bouteloua simplex Lag. 


Bouteloua pubescens Pilg., Verh. Bot. Vereins Prov. 
Brandenburg 51: 193. 1909. T-Protol.: Guatemala: 
dept. Huehuetenango: prope Malacatan, in arenosis, 
flor. Sept. E&. Seler 3234. T-Specim. 1 of 1. 
Guatemala: Huechuetenango: prope Malacatan, 13 Nov 
1896, E. Seler 3234 [IT: US-81707 (fragm.)]. 

NOTE: alternate pagination: Abh. p. 15. 

= Bouteloua repens (Kunth) Scribn. 


Bouteloua pumila Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 93. 1862.  T-Protol.: USA: 
northwestern Texas, Wright 754. T-Specim.: | of 1. 
USA: New Mexico and Texas, Oct 1849, C.H. Wright 
754 |LT: PH; ILT: GH, US-81706], LT designated (as 
type) by Griffiths, Contr. U.S. Natl. Herb. 14(3): 383 
(1912). 

= Bouteloua barbata Lag. 


Bouteloua purpurea Gould & Kapadia, Brittonia 16(2): 197, 
f. 40. 1964. T-Protol.: Mexico: México: 30 mi W of 
Tepeji del Rio, 10 Sep 1962, Gould 10211 [HT: 
TAES; IT: GH, NY, TEX, UC, US]. T-Specim.: 1 of 
1. Mexico: México: 30 mi W of Tepeji del Rio, 10 Sep 
1962, F.W. Gould 1/0211 |HY: TAES; IT: GH, NY, 
TEX, UC, US-2551353]. 

DST.: Mext. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. (1980); 
McVaugh, R. (1983). 


Bouteloua pusilla Vasey, Bull. Torrey Bot. Club 11: 6. 1884. 
T-Protol.: USA: New Mexico: Kingman: 1881, G.R. 
Vasey s.n. T-Specim.: | of 2. USA: New Mexico, Jun 
1881, GR. Vasey s.n. HT: US-81725], CS. 2 of 2. 


= Bouteloua simplex Lag. 


Bouteloua quiriegoensis Beetle, Phytologia 59(4): 287. 
1986. T-Protol.: Mexico: Sonora: Mun. de Quiriego: 
Rancho Los Varillerosy Ejido Nogales Ganaderos, 
rara, suelos areno-gravosos,, arcillo arenosos y franco- 
arenosos en el matorral arborescente, ocurriendo en 
orillas de arroyos temporales, R. Aguirre Murrieta & 
R. Cuadra Garcia s.n. [HT: COCA], acronym of 
holotype only indirectly, not explicitly, cited. T- 
Specim.: | of 1. Mexico: Sonora, Quiriego Mun., 25 
Oct 1985, R. Aguirre, Ro Cuandra s.n. (HT: COCA; 
IT: US-03206090], OS. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1986); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000). 


Bouteloua racemosa Lag., Varied. Ci. 2(4): 141. 1805. T- 
Protol.: [HT: MA], "Bot. pendula H.R.M." [=MA]. T- 
Specim.: 1 of 1. Mexico [HT: MA], Collected by L. 
Nee in Mexico, possibly then cultivated in Madrid. 

= Bouteloua curtipendula (Michx.) Torr. 


Bouteloua racemosa vat. aristosa (A. Gray) S. Watson & 
J.M. Coult., Manual 656. 1890. Basionym: Bouteloua 
curtipendula var. aristosa A. Gray. 

= Bouteloua curtipendula (Michx.) Torr. 


Bouteloua radicosa (E. Fourn.) Griffiths, Contr. U.S. Natl. 
Herb. 14: 411, t. 81. 1912. Basionym: Atheropogon 
radicosus E. Fourn. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Abrams, L. (1923); Beetle, A. A. 
(1977); Beetle, A. A. (1987); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Gould, F. 
W. (1980); Hitchcock, A. S. (1951); Munz, P. A. & 
D. D. Keck (1959); Shreve, F. & I. L. Wiggins 
(1964); Viegi, L. & G. Renzoni. (1981). 
Atheropogon bromoides (Kunth) Roem. & Schult.; 
Atheropogon  radicosus FE. Fourn.; Bouteloua 
bromoides Lag.; Bouteloua bromoides var. radicosa 
(E. Fourn.) Vasey ex L.H. Dewey; Dineba bromoides 
Kunth; Dinebra  bromoides Kunth; FEutriana 
bromoides (Kunth) Kunth; Eutriana bromoides 
(Kunth) Trin.; Nestlera festuciformis Willd. ex Steud. 


SYN.: 


Bouteloua rahmeri Phil., Verz. Antofagasta Pfl. 85. 1891. T- 
Protol.: Chile: Tarapaca: ad Calcalhuay reperta, F. 
Philippi s.n. T-Specim.: 1 of 2. Chile: Tarapaca, R.A. 
Philippi 268 [HT: SGO (photo, US); IT: US-A865562 
(fragm. ex SGO)}]. 2 of 2. 

NOTE: as Bouteloa "Rahmeri". 
= Bouteloua simplex Lag. 


Bouteloua ramosa Scribn. ex Vasey, U.S.D.A. Div. Bot. 
Bull. 12(1): t. 44. 1890. T-Protol.:G. C. Nealley 797. 
T-Specim.: | of 2. USA: Texas: Presidio Co.: Chenate 
Mts., SW Texas, Nealley 797 [NT: US-883244], NT 
designated by Beal, Grass. N. Amer. 2: 418 (1896). 2 
of 2. USA: Texas: Presidio Co.: Chenate Mts., 1888, 
G.C. Nealley 797 [LT: US-883244], LT designated by 
Reeder, Taxon 35; 153 (1986). See also Beal, Grasses 
N. Amer. (1896); Griffiths (1912); Gould (1979: 411). 

NOTE: also publ. as nom. nud. in Contr. U.S. Natl. Herb. 
(1890). 
DST.: Mexi, USA. 


Catalogue of New World Grasses: II. 3] 


LVL. ACCPTC:: |. 

ALSO ACCPTD. BY: Beetle, A. A. (1987); Correll, D. S. 
& M. C. Johnston (1970); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Reeder, 
J.R. & C. G. Reeder (1980). 

SYN.: Bouteloua oligostachya var. ramosa (Scribn. ex 
Vasey) Scribn. ex Beal. 


Bouteloua reederorum Columbus, Aliso 18(1): 63. 1999. 
REPLACED NAME: Atheropogon stolonifer E. Fourn. 
BLOCKING NAME: Bouteloua stolonifera Scribn. 

DST.: Mexi. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000). 

SYN.: Atheropogon — stolonifer E. Fourn.; Bouteloua 
stolonifera Scribn.; Cathestecum stoloniferum (E. 
Fourn.) Griffiths; Cyclostachya stolonifera (Scribn.) 
Reeder & C. Reeder; Pringleochloa stolonifera (E. 
Fourn.) Scribn. 


Bouteloua reflexa Swallen, N. Amer. Fl. 17: 632. 1939. T- 
Protol.: Mexico: on Mt. sides near Lodiego, Sonora, 9- 
15 Oct 1891, Palmer 1655 [HT: US-471374]. T- 
Specim.: | of 1. Mexico: Sonora: mountain sides near 
Lodeigo, 9-15 Oct 1891, &. Palmer 1655 [HT: US- 
471374; IT: ARIZ, GH, US]. 

DST.: Mexi. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. & R. Moran 
(1981); Gould, F. W. (1980). 


Bouteloua repens (Kunth) Scribn., Bull. Div. Agrostol., 
U.S.D.A. 4: 9. 1897. Basionym: Dinebra_ repens 
Kunth. 

NOTE: basionym not explicitly cited, only the basionym 
author cited; also by Scribn. & Merr. in U.S.D.A. 
Div. Agrostol. Bull. 24: 26 (1901). 

DST.: Beli, Cari, Colo, CoRi, ElSa, Guat, Hond, Mexi, 
Nica, Pana, USA, Vene. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Balick, M. J., 
M. H. Nee & D. E. Atha (2000); Beetle, A. A. 
(1977); Beetle, A. A. (1987); Berendsohn, W. G. & 
A. E. Araniva de Gonzalez (1991); Breedlove, D. E. 
(1986); Catasus Guerra, L. (1997); Correll, D. S. & 
M. C. Johnston (1970); Davidse, G., M. Sousa S. & 
A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Gould, F. W (1979); Gould, F. W. & R. 
Moran (1981); Gould, F. W. (1980); Liogier, H. & L. 
Martorell (1982); McClure, F. A. (1955); Molina R., 
A. (1975); Pohl, R. W. (1980); Pohl, R. W. (1994); 
Reyes-Garcia, A. & M. Sousa S. (1997); Viegi, L. & 
G. Renzoni. (1981). 

SYN.: Actinochloa bromoides (Lag.) Roem. & Schult.; 
Atheropogon americanus var.depauperata E. Fourn.; 
Atheropogon filiformis FE. Fourn.; Atheropogon 
juncifolius (Desv.) Spreng.; Atheropogon repens 
(Kunth) Roem. & Schult.; Bouteloua bromoides 
Lag.; Bouteloua  bromoides var. bromoides; 
Bouteloua filiformis (E. Fourn.) Griffiths; Bouteloua 
heterostega (Trin.) Griffiths; Bouteloua 
humboldtiana Griseb.; Bouteloua pubescens Pilg.; 
Dinebra juncifolia (Desv.) Steud.; Dinebra repens 


Kunth; Eutriana bromoides (Kunth) Trin.; Eutriana 
bromoides (Lag.) Trin.; Eutriana heterostega Trin.: 
Eutriana juncifolia (Desv.) Kunth; Eutriana repens 
(Kunth) Trin.;  Heterosteca  juncifolia  Desv.: 
Heterosteca rhadina Nash. 


Bouteloua rigidiseta (Steud.) Hitchce., J. Wash. Acad. Sci. 
23(10): 453. 1933. Basionym: Aegopogon rigidisetus 
Steud. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Allen, C. M., C. S. Reid & C. H. 
Doffitt (1999); Anonymous (1986); Beetle, A. A. 
(1977); Beetle, A. A. (1987): Correll, D. S. & M. C. 
Johnston (1970); Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. 
(1980); Great Plains Flora Association (1986); 
Hitchcock, A. S. (1951). 

SYN.: Aegopogon rigidisetus Steud.; Bouteloua texana S. 
Watson; Polyodon texanus (S. Watson) Nash. 


Bouteloua_ rigidiseta var. chihuahuana M.C.  Johnst., 
SouthW. Naturalist 27(1): 29. 1982. T-Specim.: 1 of 1. 
Mexico: Chihuahua, 24 Jul 1972, F. Chiang et al. 
8817 [HT: LL; IT: CAS, MEXU, NY]. 

= Bouteloua chihuahuana (M.C. Johnst.) Columbus. 


Bouteloua rothrockii Vasey, Contr. U.S. Natl. Herb. 1(8): 
268. 1893. T-Protol.. USA: Arizona: Cottonwood, 
1874, J.T. Rothrock 347 [HT: US]. T-Specim.: | of 1. 
USA: Arizona: Yavapai Co.: Cottonwood, 1874,/.7. 
Rothrock 347 [HT: US-81709; IT: GH]. 
= Bouteloua barbata var. rothrockii (Vasey) Gould. 


Bouteloua scabra (Kunth) Columbus, Aliso 18(1); 64, 1999, 
Basionym: Pentarrhaphis scabra Kunth. 
DST.: Beli, Colo, Guat, Hond, Mexi. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000). 
SYN.: Pentarrhaphis scabra Kunth. 


Bouteloua scorpioides Lag., Gen. Sp. Pl. 5. 1816. T-Protol.: 
Habitat in N. Hisp. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977): Beetle, A. A. 
(1987); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. (1980); Viegi, L. 
& G. Renzoni. (1981). 

SYN.: Actinochloa scorpioides (Lag.) Roem. & Schult.; 
Atheropogon scorpiodes (Lag.) Spreng.; 
Chondrosium scorpoides (Lag.) Kunth; Chondrosum 
scorpioides (Lag.) Kunth. 


Bouteloua simplex Lag., Varied. Ci. 2(4): 141. 1805. T- 
Specim.: | of 1. Peru, Lagasca s.n. [HT: MA; IT: M, 
LE-TRIN-2127.02]. 

DST.: Arge, Boli, Ecua, Mexi, Pana, Peru, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Beetle, A. A. (1987); Brako, L. & J. L. 
Zarucchi (1993); Correll, D. S. & M. C. Johnston 
(1970); Cronquist, A., A. H. Holmgren, N. H. 
Holmgren & J. L. Reveal (1977); Espejo Serna, A.. 
A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Foster, R. C. (1958); Gleason, H. A. & A. Cronquist 
(1968); Gould, F. W. & R. Moran (1981); Gould, F. 


32 Catalogue of New World Grasses: II. 


W. (1980); Hitchcock, A. S. (1927); Hitchcock, A. S. 
(1951); Macbride, J. F. (1936); Tovar, O. (1993); 
Viegi, L. & G. Renzoni. (1981). 

Actinochloa humilis (P. Beauv. ex Kunth) Willd. ex 
Roem. & Schult.; Actinochloa procumbens (P. 
Durand) Roem. & Schult.; Actinochloa prostrata 
(hort. ex Lag.) Roem. & Schult.; Actinochloa simplex 
(Lag.) Roem. & Schult.; Actinochloa tenuis (P. 
Beauv. ex Kunth) Willd. ex Roem. & Schult.; 
Atheropogon humilis (P. Beauv. ex Kunth) Spreng.; 
Atheropogon procumbens (P, Durand) J. Jacq.; 
Bouteloua brachyanthera Phil.; Bouteloua humilis 
(P. Beauv. ex Kunth) Hieron.; Bouteloua procumbens 
(P. Durand) Griffiths; Bouteloua prostrata hort. ex 
Lag.; Bouteloua pusilla Vasey; Bouteloua rahmeri 
Phil.; Bouteloua simplex var. actinochloides Henrard; 
Bouteloua simplex var. rahmeri (Phil.) Henrard; 
Bouteloua tenuis (P. Beauv. ex Kunth) Griseb.; 
Bouteloua tenuis var. humilis (P. Beauv. ex Kunth) 
Griseb.; Chloris filiformis Poir.; Chloris procumbens 
P. Durand; Chloris tenuis Poir.; Chondrosium humile 
Kunth; Chondrosium simplex (Lag.) — Kunth; 
Chondrosium  tenue P. Beauv. ex Kunth; 
Chondrosum humile P. Beauv.; Chondrosum humile 
P. Beauv. ex Kunth; Chondrosum procumbens (P. 
Durand) Desv.; Chondrosum procumbens (P. 
Durand) Desv. ex P. Beauv.; Chondrosum 
prostratum (hort. ex Lag.) Kunth; Chondrosum 
prostratum (hort. ex Lag.) Sweet; Chondrosum 
simplex (Lag.) Kunth; Chondrosum tenue P. Beauvy.; 
Chondrosum tenue P. Beauv. ex Kunth; Cynodon 
procumbens (P. Durand) Raspail; Erucaria lutescens 
Cerv.; Eutriana humilis (P. Beauv. ex Kunth) Trin.; 
Eutriana tenuis (P. Beauv. ex Kunth) Trin. 


SYN.: 


Bouteloua simplex var. actinochloides Henrard, Meded. 
Rijks-Herb. 40: 65, f. 1921. T-Protol.: Bolivien: dans 
une prairie marecageuse, avant darriver a Cochypata, 
3000m : 7 Nov 1857, Mandon pro parte [HT: L- 
908.86-226], "mixt. cum Bouteloua simplice. typ.". T- 
Specim.: | of 1. Bolivia: Cochypata, 7 Nov 1857,G. 
Mandon 908 (HT: L; IT: US-A865564 (fragm.)]. 

= Bouteloua simplex Lag. 


Bouteloua simplex var. rahmeri (Phil.) Henrard, Meded. 
Rijks-Herb. 40: 66. 1921. Basionym: Bouteloua 
rahmeri Phil. 

= Bouteloua simplex Lag. 


Bouteloua sonorae Griffiths, Contr. U.S. Natl. Herb. 14(3): 
389, f. 43. 1912. T-Protol.: Mexico: Yaqui River, 
1869, Palmer s.n. (HT: US]. T-Specim.: 1 of 2. 
Mexico: Sonora: Yaqui River, 1869, E. Palmer s.n. 
[HT: US-2808965], CS. 2 of 2. 

= Bouteloua barbata var. sonorae (Griffiths) Gould. 


Bouteloua stolonifera’ Scribn., Proc. Acad. Nat. Sci. 
Philadelphia 1891: 302. 1891. T-Protol.: Mexico: 
plains, Zacatecas, between San Luis Potosi and Aguas 
Calientes, 19 Aug 1890,C.G. Pringle 3174 (HT: US]. 
T-Specim.: | of 1. Mexico: Zacatecas: La Honda 
Station, 19 Aug 1890, C.G. Pringle 3174 [HT: US- 


72212; IT: MO-2959825, US-820985, US-883933, 


US-747802]. 
= Bouteloua reederorum Columbus. 


Bouteloua stricta Vasey, Bull. Torrey Bot. Club 15: 49. 1888. 


T-Protol.: USA: Texas: 1887, G.C. Nealley s.n. [HT: 


US]. T-Specim.: | of 1. USA: Texas, 1887, G.C. 
Nealley s.n. (HT: US-81710], CS. 
NOTE: nom. seminud.; U. S. Dept. Agr. Div. Bot. Bul. 
12(1): pl. 45. 1890. 
= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Bouteloua swallenii Columbus, Aliso 18(1): 63. 1999. 
REPLACED NAME: Pentarrhaphis annua Swallen. 
BLOCKING NAME: Bouteloua annua Swallen. 
DST.: Colo, CoRi, ElSa, Hond, Peru, Vene. 

LVL. ACCPTC.: 1. 
SYN.: Pentarrhaphis annua Swallen. 


Bouteloua tenuis (P. Beauv. ex Kunth) Griseb., Abh. Konigl. 
Ges. Wiss. Gottingen 19: 259. 1874. Basionym: 
Chondrosum tenue P. Beauv. ex Kunth. 

= Bouteloua simplex Lag. 


Bouteloua tenuis var. humilis (P. Beauv. ex Kunth) Griseb., 
Abh. Konigl. Ges. Wiss. Gottingen 24: 303. 1879. 
Basionym: Chondrosum humile P. Beauy. ex Kunth. 

= Bouteloua simplex Lag. 


Bouteloua texana S. Watson, Proc. Amer. Acad. Arts 18: 
196. 1883. T-Specim.: | of 6. USA: Texas: San 
Antonio de Bejar, Berlandier 1535 [LT: GH], LT 
designated by Griffiths, Contr. U.S. Natl. Herb. 14: 
404 (1912). 2 of 6. USA: Texas, no date, Drummond 
340 [IST: US-1063201 (fragm. ex E), US-1063202 
(fragm.)]. 3 of 6. USA: Texas, no date, 7) Drummond 
374 [ST: US-1063200 (fragm. ex E)]. 4 of 6. , 
Lindheimer 732 [ST]. 5 of 6. : Wright 752 [ST]. 6 of 6. 
: Hall 771 [ST]. 

= Bouteloua rigidiseta (Steud.) Hitche. 


Bouteloua triaena (Trin. ex Spreng.) Scribn., Proc. Acad. 
Nat. Sci. Philadelphia 1891: 307. 1891. Basionym: 
Atheropogon triaena Trin. ex Spreng. 

NOTE: as Bouteloua "Triaena". 

DST.: Guat, Mexi. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Breedlove, D. E. (1986); Davidse, G., M. 
Sousa S. & A. O. Chater (editores generales) (1994); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Gould, F. W. (1980); McClure, F. A. 
(1955); Pohl, R. W. (1994); Sousa S.. M. & E. F. 
Cabrera C. (1983); Viegi, L. & G. Renzoni. (1981). 

SYN.: Atheropogon triaena Trin. ex Spreng. Dinebra 
cristata J. Presl; Eutriana triaena Trin.: Triaena 
racemosa Kunth; Triathera gracilis E. Fourn.; 
Triathera racemosa (Kunth) Desv. 


Bouteloua trifida Thurb., Proc. Amer. Acad. Arts 18: 177. 
1883. T-Protol.: [Mexico]: Monclova, Coahuila, 1880, 
Palmer 1355. T-Specim.: | of 1. Mexico: Coahuila: 
Monclova, Aug 1880 [Feb-Oct given on specimens at 
GH and US, this crossed out on the GH label and Aug 
written in; the 2nd US sheet has Aug 14-17], £. 
Palmer 1355 (HT: GH; IT: NY, US-A865565, US- 
821003, US-883967, US-photo neg. 83761 sheet 
2474825]. 

DST.: Mexi, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Beetle, A. A. (1987); Correll, D. S. & M. C. 
Johnston (1970); Cronquist, A., A. H. Holmgren, N. 
H. Holmgren & J. L. Reveal (1977); Espejo Serna, 


Catalogue of New World Grasses: II. 33 


A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W. & R. Moran (1981); Gould, F. W. 
(1980); Hickman, J. C. (1993); Hitchcock, A. S. 
(1951); Munz, P. A. & D. D. Keck (1959); Munz, P. 
A. (1974); Shreve, F. & I. L. Wiggins (1964). 
Bouteloua burkii Scribn.,; Bouteloua burkii Scribn. ex 
S. Watson; Bouteloua trifida var. burkii (Scribn. ex 
S. Watson) Vasey ex L.H. Dewey; Bouteloua trinii 
(E. Fourn.) Griffiths; Chondrosum polystachyum 
Trin. ex E. Fourn.; Chondrosum trifidum (Thurb.) 
Clayton; Chondrosum trinii E. Fourn.; Chondrosum 
virletii E. Fourn.; Chondrosum virletii E. Fourn. ex 
Hemsl. 


SYN.: 


Bouteloua trifida var. burkii (Scribn. ex S. Watson) Vasey ex 
L.H. Dewey, Contr. U.S. Natl. Herb. 2(3): 532. 1894. 
Basionym: Bouteloua burkiiScribn. ex S. Watson. 

NOTE: cites B. burkii Scribn., (1884), which shares a 
syntype with B. burkei Scribn. ex S. Wats., (1883). 
= Bouteloua trifida Thurb. 


Bouteloua trinii (E. Fourn.) Griffiths, Contr. U.S. Natl. Herb. 
14(3): 387, f. 41, 42, t. 74. 1912. Basionym: 
Chondrosum triniiE. Fourn. 

= Bouteloua trifida Thurb. 


Bouteloua uniflora Vasey, Bot. Gaz. 16: 26. 1891. T-Protol.: 
USA: Texas: Crockett Co.: 1890, G.C. Nealley 222 
[HT: US]. T-Specim.: | of 1. USA: Texas: Crockett 
Co., 1890, G.C. Nealley 222 [HT: US-2307997; IT: 
GH]. 

DST.: Mexi, USA. 

LVL. ACCPTC.:: |. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1987); Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Gould, F. W. (1980); Hitchcock, 
A. S. (1951); Viegi, L. & G. Renzoni. (1981). 


Bouteloua uniflora var. coahuilensis Gould & Kapadia, 
Brittonia 16(2): 191, f. 27, 41. 1964. T-Protol.: 
Mexico: Coahuila: Rancho Los Angeles, 40 km SW of 
Saltillo, 4 Nov 1962, Gould 10300 [HT: TAES; IT: 
DS, GH, GH, TEX, UC, US]. T-Specim.: 1 of 1. 
Mexico: Coahuila: Rancho Los Angeles, 40 km SW of 
Saltillo, 4 Nov 1962, FW. Gould 10300 [HT: TAES; 
IT: DS, GH, TEX, UC, US-2551355]. 

DST.: Mexi. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. (1980); Viegi, L. 
& G. Renzoni. (1981). 


Bouteloua uniflora Vasey var. uniflora. 
DST.: Mexi, USA. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. (1980). 


Bouteloua vaneedenii Pilg., Symb. Antill. 6: 2. 1909. T- 
Protol.: [West Indies]: Leeward Isls.: Anguilla, Sep, 
Boldingh 3512B, fl. et fruct. T-Specim.: 1 of 1. St. 
Christopher-Nevi: Anguilla I., 1906, /. Boldingh 3512 
[HT: B; IT: US-2307996 (fragm. ex B)]. 

DST.: Cari, Vene. 
LVL. ACCPTC.:: 1. 


ALSO ACCPTD. BY: Catasus Guerra, L. (1997); Gould, F. 
W (1979); Gould. F. W. (1980); Hitchcock, A. S. 
(1936). 


Bouteloua varia (Swallen) Columbus, Aliso 18(1): 63. 1999. 
Basionym: Cathestecum varium Swallen. 
DST.: Mexi. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000). 
SYN.: Cathestecum varium Swallen. 


Bouteloua vestita (S. Watson) Scribn. ex L.H. Dewey, Contr. 
U.S. Natl Herb. 2(3): 531. 1894. Basionym: 
Bouteloua polystachya var. vestita S. Watson. 

= Bouteloua parryi var. parryi. 


Bouteloua warnockii Gould & Kapadia, SouthW. Naturalist 
7: 176. 1962. T-Protol.: USA: Texas: Culberson Co.: 2 
mi W of Kent, 18 Jul 1961, Gould & Kapadia 9533 
[HT: TAES; IT: MICH, MO, SMU, SRSC, TEX, UC, 
US]. T-Specim.: | of 1. USA: Texas: Culberson Co., 
18 Jul 1961, F.W. Gould & Z. Kapadia 9533 (HT: 
TAES; IT: MICH, MO, SMU, SRSC, TEX, UC, US- 
2551367]. 

DST.: Mexi, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Beetle, A. A. (1987); Correll, D. S. & M. C. 
Johnston (1970); Espejo Serna, A.. A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. 
(1980). 


Bouteloua williamsii Swallen, Ceiba 4(5):; 285. 1955. T- 
Protol.: Honduras: El Paraiso: near Las Mesas, 900 m, 
on open rocky hillsides, 2 Oct 1949, L.O. Williams 
16902 [HT: US-2044038]. T-Specim.: 1 of 1. 
Honduras: El Paraiso: open rocky hillsidesnear Las 
Mesas, alt. 900 m, ca. 14°N, 87°W, drainage of the Rio 
Yeguare, 2 Oct 1949, L.O. Williams 16902 [HT: US- 
2044038; IT: F, ISC, MO-1731263, UC, WIS]. 

NOTE: as Bouteloua "Williamsii". 

DST.: Guat, Hond, Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Breedlove, D. E. (1986); Davidse, G.. M. 
Sousa S. & A. O. Chater (editores generales) (1994); 
Espejo Serna, A., A. R. L6pez-Ferrari, and J. Valdes- 
Reyna (2000); Gould, F. W. (1980); Molina R., A. 
(1975); Pohl, R. W. (1994); Viegi, L. & G. Renzoni. 
(1981). 


Boutelouinae subtrib. Stapf, Fl. Trop. Afr. 9: 22. 
1917. TYPE: Bouteloua Lag. 

TRTD. BY: Soreng, R. J., G. Davidse, T. S. Filgueiras, E. J. 
Judziewicz, P. M. Peterson, and F. O. Zuloaga. 

NHT: Cynodonteae. 

CNSLTD. LIT.: Columbus, J. T., M. S. Kinney, R. Pant & 
M. E. Siqueiros Delgado, (1998); Columbus, J. T., 
(1999). 


Brachyelytrum pringlei Vasey ex Beal, Grass. N. Amer. 2: 
266. 1896. T-Protol.: Mexico, Pringle 1739, 2661. 
NOTE: nom. inval., as syn. of Muhlenbergia trinii Fourn. 
= Muhlenbergia diversiglumis Trin. 


34 Catalogue of New World Grasses: II. 


Briza barbata (Nees) Trin., Mem. Acad. Imp. Sci. St.- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 363. 1830. 
Basionym: Calotheca barbata Nees. 

= Neesiochloa barbata (Nees) Pilg. 


Briza caroliniana Lam., Encycl. 1: 465. 1785. T-Protol.: 
USA: Caroline & la Virginie. 


= Uniola paniculata L. 


Briza dura Desv., Mem. Soc. Agric. Angers 1: 207. 1831. T- 
Protol.: Habitat in America australi. T-Specim.: | of 1. 
Argentina: Buenos Aires?, Desvaux /2 |HT: P; IT: 
US-A865554]. 

NOTE: also in Opusc. Sci. Phys. Nat. 103 [reprint] 1831- 
1833. Presumably an Old World species of 
Eragrostis introduced, but the species 
undeterminable from the tiny type fragment at US. 

= Status not yet determined. 


Briza eragrostis L., Sp. Pl. 70. 1753. T-Protol.: Habitat in 
Europa australi ad agrorum versuras. T-Specim.: | of 
1. "Europa Australi", no date, J. Clayton 582 [T: BM; 
IT: US-3152245 (fragm. ex BM)]. 
CORRECT NAME: Eragrostis major Host. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Briza megastachya (Koeler) hort. ex Roem. & Schult., Syst. 
Veg. 2: 585. 1817. Basionym: Poa megastachya 
Koeler. 

NOTE: nom. inval., as syn. of Poa megastachya. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Briza megastachya (Koeler) Steud., Nomencl. Bot. (ed. 2) 1: 
225. 1840. Basionym: Poa megastachya Koeler. 
NOTE: nom. inval., as syn. of Poa megastachya Koeler. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Briza oblonga Moench, Methodus 185. 1794. 
REPLACED NAME: Briza eragrostis L. 
BLOCKING NAME: Poa eragrostis L. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Briza purpurascens Muhl., Descr. Gram. 154. 1817. T- 
Protol.: USA: habitat in Carolina. T-Specim.: | of 1. 
USA: no date, Coll. Ukn. s.n. (IT: US-1535783]. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Briza spicata (L.) Lam., Encycl. 1: 465. 1785. Basionym: 
Uniola spicata L. 
NOTE: hom. illeg. 
E. NAME: Briza spicata Burm. f. 
= Distichlis spicata (L.) Greene. 


Brizopyrum americanum (L.) Link, Hort. Berol. 1: 160. 1827. 
Basionym: Uniola spicata L. 
= Distichlis spicata (L.) Greene. 


Brizopyrum boreale J. Presl, Relig. Haenk. 1(4-5): 280. 1830. 
T-Protol.: Hab. in sinu Nootka, Haenke s.n. T- 
Specim.: | of 1. Canada: British Columbia: Vancouver 
Island: Nootka Sound, Haenke s.n. [HT: PR; IT; MO- 
1837457]. 

= Distichlis spicata var. borealis (J. Presl) Beetle. 


Brizopyrum condensatum (J. Presl) E. Fourn., Mexic. Pl. 2: 
121. 1886. Basionym: Megastachya condensata J. 
Presl. 

= Eragrostis condensata (J. Pres!) Steud. 


Brizopyrum cyperoides (Thunb.) Nees, Fl. Afr. Austral, Ill, 
374. 1841. 
= Cladoraphis cyperoides (Thunb.) S.M. Phillips. 


Brizopyrum obtusiflorum E. Fourn., Mexic. Pl. 2: 120. 1886. 
T-Protol.: Mexico: Veracruz: Orizaba: in ora 
occidentali, Emy s.n. (HT: P]. 

= Eragrostis obtusiflora (E. Fourn.) Scribn. 


Brizopyrum ovatum Nees ex Steud., Syn. Pl. Glumac. 1: 282. 
1854. T-Protol.: Peru, Cuming s.n. (HT: P?], or Chile?. 
= Distichlis spicata (L.) Greene. 


Brizopyrum pilosum J. Presl, Reliq. Haenk. 1(4-5): 280. 
1830. T-Protol.: Hab. ad Acapulco, Haenke s.n. T- 
Specim.: | of 1. Mexico: Guerrero: Acapulco, Haenke 
sn. (HT: PR; IT: MO-2112280, MO-5117060 
(drawing), US-865534 (fragm. ex PR)]. 

= Jouvea pilosa (J. Presl) Scribn. 


Brizopyrum prostratum (Kunth) E. Fourn., Bull. Soc. Roy. 
Bot. Belgique 15: 474. 1876. Basionym:Poa prostrata 
Kunth. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Brizopyrum prostratum (Kunth) Benth. ex E. Fourn., Mexic. 
Pl. 2: 120. 1886. Basionym: Poa prostrata Kunth. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Brizopyrum prostratum var. erectum E. Fourn., Mexic. Pl. 2: 
120. 1886. T-Specim.: | of 2. Mexico, Berlandier 376 
[ST: LE]. 2 of 2. Mexico: Prov. Puebla: secus rivos 
Tehuacan de las Granadas, Galleotti 5746 [ST]. 
= Distichlis spicata (L.) Greene. 


Brizopyrum prostratum var. humile E. Fourn., Mexic. Pl. 2: 
120. 1886. T-Specim.: | of 3. Mexico, Berlandier 923 
[ST: LE]. 2 of 3. Mexico, Bourgeau 230 [ST]. 3 of 3. 
Mexico, /Berlandier?] 376 [ST: P]. 
= Distichlis spicata (L.) Greene. 


Brizopyrum spicatum (L.) Hook. & Arn., Bot. Beechey Voy. 
403. 1840. Basionym: Uniola spicata L. 
= Distichlis spicata subsp. spicata. 


Brizopyrum spicatum var. strictum (Torr.) A. Gray ex S. 
Watson, Botany Fortieth Parallel 385. 1871. 
Basionym: Uniola stricta Torr. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Brizopyrum thalassicum (Kunth) Nees, Nov. Actorum Acad. 
Caes. Leop.-Carol. Nat. Cur. 1: 29. 1841. Basionym: 
Poa thalassica Kunth. 
NOTE: 161. 1843. 
= Distichlis spicata (L.) Greene. 


Brizopyrum uninervium (J. Presl) E. Fourn., Mexic. Pl. 2: 
121. 1886. Basionym: Megastachya uninervia J. Presl. 
= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Bromus spicatus Nees, Fl. Bras. Enum. Pl. 2: 471. 1829. T- 
Protol.: Brasil: habitat in campis, Campo mimoso 
dictis, provinciae Piauhianae, floret Maio et Junio, 
Martius s.n. [HT: M]. T-Specim.: | of |. [Brasilia]: 
Piauhia [IT: LE-TRIN-2313.01 (& fig.)]. 

= Tripogon spicatus (Nees) Ekman. 


Bromus virgatus J. Presl, Relig. Haenk. 1(4-5): 263. 1830. T- 
Protol.: Hab. in montanis Peruviae, in Mexico? 
[changed to Hab. ad monte-Rey Californiae," in the 
Addendum on p. 351], Haenke s.n., Probably from 


Catalogue of New World Grasses: II. 35 


Peru [gd, 1995]. T-Specim.: | of 1. Mexico: Guerrero: 
Acapulco, Haenke s.n. [HT: PR; IT: BM, MO-14696, 
MO-134997 (line drawing), US-A865483 (fragm.)], 
cited by Ortiz (1993). 

= Gouinia virgata (J. Presl) Scribn. 


Brousemichea Bal., J. Bot. (Morot) 4(8): 163. 1890. TYPE: 
Brousemichea seslerioides Bal. 
= Zoysia Willd. 


Buchloe Engelm., Trans. Acad. Sci. St. Louis 1: 432. 1859. 
TYPE: Buchloe dactyloides (Nutt.) Engelm. 
NOTE: nom. cons. 
= Bouteloua Lag. 


Buchloe dactyloides (Nutt.) Engelm., Trans. Acad. Sci. St. 
Louis 1: 432, t. 12, 14, f 1-17. 1859. Basionym: 
Sesleria dactyloides Nutt. 

= Bouteloua dactyloides (Nutt.) Columbus. 


Buchlomimus Reeder, C. Reeder & Rzed., Brittonia 17(1): 
29. 1965. TYPE: Buchlomimus nervatus (Swallen) 
Reeder, C. Reeder & Rzed. 

= Bouteloua Lag. 


Buchlomimus nervatus (Swallen) Reeder, C. Reeder & Rzed., 
Brittonia 17: 30. 1965. Basionym: Bouteloua nervata 
Swallen. 

= Bouteloua nervata Swallen. 


Bulbilis Raf. ex Kuntze, Revis. Gen. Pl. 2: 763. 1891. 
NOTE: nom. superfl. pro Buchloe. 
= Bouteloua Lag. 


Bulbilis dactyloides (Nutt.) Raf. ex Kuntze, Revis. Gen. PI. 2: 
763. 1891. Basionym: Sesleria dactyloides Nutt. 
= Bouteloua dactyloides (Nutt.) Columbus. 


Calamagrostis andina Nutt., J. Acad. Nat. Sci. Philadelphia, 
ser. 2, 1: 187. 1848. T-Protol.: USA: In Upper 
California on the Colorado of the West, Dr. W. 
Gambel. T-Specim.: | of 1. USA: California, Sept-Oct 
1894, W. Gambel [T]. 

= Muhlenbergia andina (Nutt.) Hitchce. 


Calamagrostis brevipilis (Torr.) L.C. Beck, Bot. North. 
Middle States 401. 1833. Basionym: Arundo brevipilis 
Torr. 
= Calamovilfa brevipilis (Torr.) Scribn. 


Calamagrostis compressa DOll, Fl. Bras. 2(3): 56. 1878. T- 
Protol.: Hort.: seminibus, Brazil: pr. Rio de Janeiro, 
Glaziou [Cult in horto bot Monacensi anno 1869]. 

= Muhlenbergia frondosa (Poir.) Fernald. 


Calamagrostis curtissii (Vasey) Vasey, Bot. Gaz. 15: 269. 
1890. Basionym: Ammophila curtissii Vasey. 
= Calamovilfa curtissii (Vasey) Scribn. 


Calamagrostis gigantea Nutt., Trans. Amer. Philos. Soc., 
n.s., 5: 143. 1837. T-Protol.: Arkansas Territory: Hab. 
on the sandy banks of Great Salt River of the 
Arkansas. T-Specim.: | of 1. USA: Oklahoma: Hab. 
on the sandy banks of Great Salt River of the 
Arkansas., Nuttall s.n. (HT: BM?; IT: PH, US-865774 
(fragm. ex BM), US-A865773 (fragm. ex PH)]. 

= Calamovilfa gigantea (Nutt.) Scribn. & Merr. 


Calamagrostis longifolia Hook., Fl. Bor.-Amer. 2: 241. 1840. 
T-Specim.: | of |. Canada: Saskatchawan, no date, 7. 


Drummond s.n. [HT: K; IT: US-3168581 (fragm. ex 
K)]. 
= Calamovilfa longifolia (Hook.) Hack. ex Scribn. 


Calamagrostis mexicana Kunth, Nov. Gen. Sp. 7: 488. 1825. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Calamagrostis quitensis Kunth. 
= Muhlenbergia tenuifolia (Kunth) Kunth. 


Calamagrostis pumila Hook. f., Trans. Linn. Soc. London 20: 
176. 1847. T-Protol.: Ecuador: Galapagos Isls.: 
Albermarle Isl., Macrae [LT]. T-Specim.: | of 1. 
Ecuador: Galapagos Islands, Macrae s.n. [LT: kK; 
ICGE, K], LT designated by Porter, Bot. J. Linn. Soc. 
81: 98 (1980). 

= Trichoneura lindleyana var. lindleyana. 


Calamagrostis quitensis Kunth, Nov. Gen. Sp. 1: 133. 1815 
[1816]. T-Protol.: Regni Mexicani prope Mescala, 
Sochipala et Valle Zopilote, inter 265 et 517 hexap.., 
Humboldt & Bonpland s.n. [HT: P]. T-Specim.: | of 1. 
Mexico: Guerrero: Sochipala et Valle Zopilote, 
Humboldt & Bonpland s.n. (HT: P-Bonpl.; IT: P]. 

= Muhlenbergia tenuifolia (Kunth) Kunth. 


Calamagrostis tenuifolia Kunth, Nov. Gen. Sp. 1: 134. 1815 
[1816]. T-Protol.: Crescit juxta urbem Mexico in 
cultis, alt. 1170 hexap., floret Aprili, Humboldt & 
Bonpland s.n. T-Specim.: | of 1. Mexico: Distrito 
Federal: Mexico City, Humboldt & Bonpland s.n. (HT: 
P-Bonpl.; IT: P]. 

= Muhlenbergia tenuifolia (Kunth) Kunth. 


Calamagrostis subg. Calamovilfa A. Gray, Manual 582. 
1848. TYPE: Calamagrostis brevipilis (Yorr.) L.C. 
Beck; LT designaated by Hitchcock, U.S.D.A. Bull. 
772: 126 (1920). 

= Calamovilfa (A. Gray) Hack. ex Scribn. & Southw. 


Calamochloa E. Fourn., Bull. Soc. Bot. France 24: 178. 1877. 
TYPE: Calamochloa filifolia E. Fourn. 
NOTE: hom. illeg. 
= Sohnsia Airy Shaw. 


Calamochloa filifolia E. Fourn., Bull. Soc. Bot. France 24: 
178. 1877. T-Protol.: Mexico: San Luis Potosi, Virlet 
d'Aoust 1461. T-Specim.: | of 1. Mexico: San Luis 
Potosi, no date, Virlet /46/ [T: P; US-A865757 
(fragm.)]. 

= Sohnsia filifolia (E. Fourn.) Airy Shaw. 


Calamovilfa (A. Gray) Hack. ex Scribn. & Southw., 


True Grasses 113. 1890. Basionym: Calamagrostis 
subg. Calamovilfa A. Gray. 
NOTE: This genus was definitely added by Scribner. 
TRTD. BY: Peterson, P. M. 
NHT: Sporobolinae. NO. ACCPTD. SPP.: 5 SUBSP.: 2. 
CNSLTD. LIT.: Thieret, J. W., (1966). 
SYN.: Calamagrostis subg. Calamovilfa A. Gray. 


Calamovilfa arcuata K.E. Rogers, Rhodora 72(789): 72-79, 
f. 1-2. 1970. T-Protol.: USA: Tennessee: Cumberland 
Co.: ca. 8 mi NNE of Crab Orchard, off of old gravel 
road from Crab Orchard to Rockwood, downstream 
about 600-800 ft from Antioch Bridge over Daddy's 
Creek, in a shrub-dominated community, 3 Oct 1968, 
Rogers, Sharp, Delgadillo & Meijer 42409 [HT: US- 
281796; IT: ISC, LAF, TAES, TENN, US]. T- 


36 Catalogue of New World Grasses: II. 


Specim.: | of |. USA: Tennessee: Cumberland Co.: ca. 
8 mi NNE of Crab Orchard, off of old grave | road 
from Crab Orchard to Rockwood, downstream about 
600-800 ft from Antioch Bridge over Daddy's Creek, 
in a shrub-dominated community, 3 Oct 1968, Rogers, 
Sharp, Delgadillo & Meijer 42409 (HT: US-281796; 
IT: ISC, LAF, TAES, TENN, US-74241]. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Rogers, K. E. 

(1970). 


Calamovilfa brevipilis (Torr.) Scribn., True Grasses 113. 
1890. Basionym: Arundo brevipilis Torr. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Fernald, M. 
(1950); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Godfrey, R. 
K. & J. W. Wooten (1979); Hitchcock, A. S. (1951); 
Radford, A. E., H. E. Ahles & C. R. Bell (1968); 
Rogers, K. E. (1970); Small, J. K. (1933); Thieret, J. 
W. (1966). 

Ammophila_ brevipilis (Torr.) Benth. ex Vasey; 
Arundo_ brevipilis Torr.; Calamagrostis brevipilis 
(Torr.) L.C. Beck; Calamovilfa brevipilis var. 
brevipilis, Calamovilfa brevipilis var. calvipes 
Fernald; Calamovilfa brevipilis var. heterolepis 
Fernald; Calamovilfa brevipilis var. typica Fernald. 


SYN:: 


Calamovilfa brevipilis (Torr.) Scribn. var. brevipilis. 
REPLACED NAME: Calamovilfa brevipilis var. typica 
Fernald. 
= Calamovilfa brevipilis (Torr.) Scribn. 


Calamovilfa brevipilis var. calvipes Fernald, Rhodora 
41(490): 501-502, pl. 573, f. 1-2, map 9. 1939. T- 
Protol.: Fernald & Long 8548 [HT: GH; IT: PH]. T- 
Specim.: | of 1. USA: Virginia: Greensville Co., 15 
1938, M.L. Fernald & B.H. Long 8548 (HT: GH; IT: 
US-3168573 (fragm. ex GH), US-1912828]. 

= Calamovilfa brevipilis (Torr.) Scribn. 


Calamovilfa_ brevipilis var. heterolepis Fernald, Rhodora 
41(490): 502, pl. 573, f. 4. 1939. T-Protol.: [USA]: 
North Carolina: Harnett Co.: edge of swamp between 
Coats and Erwin, 15 Jul 1935, Correll & Blomquist 
2539 [HT: GH]. T-Specim.: | of 1. USA: North 
Carolina: Harnett Co., 15 Jul 1935, D.S. Correll & 
HL. Blomquist 2539 (HT: GH; IT: US-2207854]. 

= Calamovilfa brevipilis (Torr.) Scribn. 


Calamovilfa brevipilis var. typica Fernald, Rhodora 41: 502. 
1939. 
NOTE: nom. inval., see ICBN Art 24.3. 
CORRECT NAME: Calamovilfa brevipilis var. brevipilis. 
= Calamovilfa brevipilis (Torr.) Scribn. 


Calamovilfa curtissii (Vasey) Scribn., Bull. Div. Agrostol., 
U.S.D.A. 17: 199, £. 495. 1899. Basionym:Ammophila 
curtissii Vasey. 

DST.: USA. 

LVL. ACCPTC.: |. 

ALSO ACCPTD. BY: Anonymous (1986); Godfrey, R. K. 
& J. W. Wooten (1979); Hitchcock, A. S. (1951); 
Johnson, A. E. & A. Blyth (1988); Rogers, K. E. 
(1970); Small, J. K. (1933); Thieret, J. W. (1966); 
Wunderlin, R. P. (1998). 


SYN.: Ammophila curtissii Vasey; Calamagrostis curtissii 
(Vasey) Vasey. 


Calamovilfa gigantea (Nutt.) Scribn. & Merr., Circ. Div. 
Agrostol. U.S.D.A. 35: 2. 1901.  Basionym: 
Calamagrostis gigantea Nutt. 

DST.: Cana, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Cronquist, A., A. H. 
Holmgren, N. H. Holmgren & J. L. Reveal (1977); 
Great Plains Flora Association (1986); Hitchcock, A. 
S. (1951); Rogers, K. E. (1970); Thieret, J. W. 
(1966). 

SYN.: Calamagrostis gigantea Nutt. 


Calamovilfa longifolia (Hook.) Hack. ex Scribn., True 
Grasses 113. 1890. Basionym: Calamagrostis 
longifolia Hook. 

DST.: Cana, USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Fernald, M. (1950); Gleason, H. A. 
& A. Cronquist (1968); Great Plains Flora 
Association (1986); Hitchcock, A. S. (1951); Moss, 
E. H. (1983); Rogers, K. E. (1970); Scoggan, H. J. 
(1978 [1979]): Thieret, J. W. (1966); Voss, E. G. 
(1972). 

Athernotus 

Calamagrostis 

Buckley. 


SYN:: (Hook.) 


Hook.;  Vilfa 


Lunell; 
rigida 


longifolius 
longifolia 


Calamovilfa longifolia | (Hook.) Hack. ex Scribn. var. 
longifolia. 

DST.: Cana. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Gleason, H. A. & A. Cronquist 
(1968); Gleason, H. A. & A. Cronquist (1991); 
Hitchcock, C. L., A. Cronquist, M. Ownbey & J. W. 
Thompson (1969); Scoggan, H. J. (1978 [1979]); 
Thieret, J. W. (1966). 


Calamovilfa longifolia var. magna Scribn. & Merr., Circ. 
Div. Agrostol. U.S.D.A. 35: 3. 1901. T-Protol.: 
Michigan: collected on the lake shore at themouth of 
the Kalamazoo river, 1894, WA. Taylor s.n. T- 
Specim.: | of 1. USA: Michigan: Allegan Co.: lake 
shore at the mouth of the Kalamazoo River, 1894, 
W.A. Taylor s.n. (HT: US-207912]. 

DST.: Cana, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Gleason, H. A. 
& A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Hitchcock, A. S. (1951); Scoggan, 
H. J. (1978 [1979]); Thieret, J. W. (1966); Voss, E. 
G. (1972). 


Calanthera Hook., Hooker's J. Bot. Kew Gard. Misc. 8: 18. 
1856. 
NOTE: nom. nud. 
= Bouteloua Lag. 


Calanthera dactyloides (Nutt.) Kunth ex Hook., Hooker's J. 
Bot. Kew Gard. Misc. 8: 18. 1856. Basionym: Ses/eria 
dactyloides Nutt. 

= Bouteloua dactyloides (Nutt.) Columbus. 


Calotheca barbata Nees, Fl. Bras. Enum. Pl. 2: 477. 1829. T- 
Protol.: Brasilia: habitat in graminosis irriguis, Campo 


Catalogue of New World Grasses: II. 37 


mimoso dictis, prov Piauhianae passim, Martius s.n. 

T-Specim.: | of 1. Brazil: Piauhy: in graminosis 

irriguis, no date, Martius s.n. [HT: M; IT: B, BAA-467 

(fragm. ex B), K, LE-TRIN-2543.01a (mis, Nees ab 

Es. 1830), MO-2112275, US-A865750 (fragm. ex M)]. 
= Neesiochloa barbata (Nees) Pilg. 


Calotheca cuspidata Spreng., Syst. Veg. 1: 348. 1825. 
REPLACED NAME: Poa cuspidata Roth. 
BLOCKING NAME: Poa cuspidata Nutt. 
= Eragrostis rufescens var. rufescens. 


Calotheca purpurascens (Muhl.) Spreng., Syst. Veg. 1: 348. 
1825. Basionym: Briza purpurascens Muhl. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Calotheca sabulosa Steud., Flora 12(2): 488. 1829. T-Protol.: 
South Africa: Cape of Good Hope, Ecklon 944. T- 
Specim.: | of 1. South Africa: Cape, Ecklon 944 (HT: 
Pi}. 

= Sporobolus virginicus (L.) Kunth. 


Calotheria Steud., Syn. Pl. Glumac. 1: 199. 1854. 
NOTE: insyn. sub Enneapogon elegans. 
= Enneapogon Desv. ex P. Beauv. 


Calycodon Nutt., Proc. Acad. Nat. Sci. Philadelphia 4: 23. 21 
Mar-4 Apr 1848. TYPE: Calycodon montanum Nutt. 
= Muhlenbergia Schreb. 


Calycodon montanum Nutt., J. Acad. Nat. Sci. Philadelphia, 
ser. 2, 1: 186. 1848. T-Protol.: Hab. in the Rocky Mts., 
near Santa Fe, Mexico. T-Specim.: 1 of 1. USA: New 
Mexico: near Santa Fe, W. Gambel s.n. [HT: BM]. 

= Muhlenbergia montana (Nutt.) Hitchc. 


Campuloa Desv., Mem. Soc. Agric. Angers 1: 167. 1813. 
NOTE: nom. illeg. superfl.; also in Opusc. Sci. Phys. Nat. 
63 [reprint] 1831-1833. 
= Ctenium Panz. 


Campuloa chapadensis Trin. ex DOll, Fl. Bras. 2(3): 74. 
1878. 
NOTE: nom. inval., as syn. of Ctenium chapadense. 
= Ctenium chapadense (Trin.) Ddll. 


Campuloa gracilis Desv., J. Bot. (Desvaux) 69. 1813. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Chloris monostachya Michx. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Campuloa monostachya (Michx.) Roem. & Schult., Syst. 
Veg. 2: 516. 1817. Basionym: Chloris monostachya 
Michx. 

= Ctenium aromaticum (Walter) Alph. Wood. 


Campulosus Desv., Nouv. Bull. Sci. Soc. Philom. Paris 2: 
189. 1810. TYPE: Campulosus gracilior Desv. 
NOTE: nom. rejic. 
= Ctenium Panz. 


Campulosus aromaticus Trin, ex Steud., Nomencl. Bot. (ed. 
2) 1: 272. 1840. T-Specim.: | of 1. South Carolina 
[OM: LE: TRIN-2068.01]. 
NOTE: nom. inval., as syn. of Campulosus monostachys. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Campulosus aromaticus (Walter) Scribn., Mem. Torrey Bot. 
Club 5(4): 45. 1894. Basionym: Aegilops aromatica 
Walter. 

= Ctenium aromaticum (Walter) Alph. Wood. 


Campulosus brachystachyus Nees, Fl. Bras. Enum. Pl. 2: 
417-418. 1829. T-Protol.: Brasilia habitat ignoto, 
Sellow s.n. [HT: B]. T-Specim.: | of 1. Brazil, no date, 
F. Sellow s.n. [HT: B; IT: US-2606902 (fragm. ex B)]. 

= Ctenium brachystachyum (Nees) Kunth. 


Campulosus chapadensis Trin., Sp. Gram. 3: t. 303. 1831. T- 
Protol.: Brazil: Figura ad specimen Brasilianum. T- 
Specim.: | of 2. Brazil [IT: K]. 2 of 2. Brazil [HT: LE- 
TRIN 2070.02]. 

= Ctenium chapadense (Trin.) Doll. 


Campulosus cirrosus Nees, Fl. Bras. Enum. Pl. 2: 416-417. 
1829. T-Protol.: Brasilia: habitat in campis elatioribus, 
3000 pedum alt., inter Villa Rica et Tejuco, prov. 
Minarum, Majo-Junio, Martius s.n. [HT: B]. T- 
Specim.: 1 of 1. Brazil: Minas Gerais: in campis 
elatioribus, 3000 pedum alt., inter Villa Rica et Tejuco, 
May-Jun, A.F.P. von Martius s.n. [IT:US: US- 
2606904 (fragm. ex B)}. 

= Ctenium cirrosum (Nees) Kunth. 


Campulosus floridanus Hitche., Amer. J. Bot. 2: 306. 1915. 
T-Specim.: | of 1. USA: Florida, 1875, 4.1. Curtiss 
s.n. [HT: US-726521]. 
= Ctenium floridanum (Hitchce.) Hitche. 


Campulosus gangitis (L.) Kuntze, Revis. Gen. Pl. 2: 764. 
1891. Basionym: Nardus gangitis L. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Campulosus gracilior Desv., Nouv. Bull. Sci. Soc. Philom. 
Paris 2: 189. 1810. 
NOTE: nom. illeg. et superfl. for Chloris monostachya 
Michx. 
REPLACED NAME: Chloris monostachya Michx. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Campulosus gracilis Bertol., Mem. Reale Accad. Sci. Ist. 
Bologna 2: 602, t. 43, f. a-c. 1850. T-Protol.: USA: 


Alabama. 
= Ctenium aromaticum (Walter) Alph. Wood. 
Campulosus  monostachyus (Michx.) P. Beauv., Ess. 


Agrostogr. 64, 157, 158, t, f. 1. 
Chloris monostachya Michx. 
= Ctenium aromaticum (Walter) Alph. Wood. 


1812. Basionym: 


Campulosus planifolius J. Presl, Relig. Haenk. 1(4-5): 287. 
1830. T-Protol.: Hab. in Mexico, Haenke s.n. T- 
Specim.: 1 of 1. Mexico, Haenke s.n. [HT: PR; IT: 
MO-1990361, US-2607350 (fragm.)]. 

= Ctenium planifolium (J. Pres!) Kunth. 


Campulosus plumosus Hitche., Contr. U.S. Natl. Herb. 17(3): 
330. 1913. T-Protol.: Mexico: Sinaloa: Cerro 
Colorado, vicinity of Culican, 5 Nov 1904, 7S. 
Brandegee [HT: US-555998]. T-Specim.: 1 of 1. 
Mexico: Sinaloa: Cerro Colorado, vicinity of Culican, 
5 Nov 1904, 7.8. Brandegee 7 [HT: US-555998]. 

= Ctenium plumosum (Hitchc.) Swallen. 


Capriola Adans., Fam. Pl. 2: 31, 532. 1763. TYPE:Capriola 
dactylon (L.) Kuntze. 
NOTE: nom. rejic. against Cynodon L.C. Rich. 1805. 
= Cynodon Rich. 


Capriola dactylon (L.) Kuntze, Revis. Gen. Pl. 2: 764. 1891. 
Basionym: Panicum dactylon L. 
= Cynodon dactylon (L.) Pers. 


38 Catalogue of New World Grasses: II. 


Capriola dactylon (L.) Hitche., Annual Rep. Missouri Bot. 
Gard. 1893: 147. 1893. Basionym: Panicum dactylon 
L. 
NOTE: isonym. 
E. NAME: Capriola dactylon (L.) Kuntze. 
= Cynodon dactylon (L.) Pers. 


Capriola dactylon var. maritima (Kunth) Hitche., U.S.D.A. 
Bull. (1915-23) 772: 179. 1920. Basionym: Cynodon 
maritimus Kunth. 

= Cynodon maritimus Kunth. 


Casiostega Galeotti, Bull. Acad. Roy. Sci. Bruxelles 9: 232. 
1842. 
NOTE: nom. nud, 


Casiostega dactyloides(Nutt.) E. Fourn., Bull. Soc. Roy. Bot. 
Belgique 15: 470. 1876. Basionym: Sesleria 
dactyloides Nutt. 

= Bouteloua dactyloides (Nutt.) Columbus. 


Casiostega hookeri Rupr. ex E. Fourn., Bull. Soc. Roy. Bot. 


Belgique 15: 471. 1876. 
NOTE: nom. inval., as syn. of Buchloe dactyloides (Nutt.) 
Engelm. 
= Bouteloua dactyloides (Nutt.) Columbus. 


Catabrosa micrantha Hochst. ex A. Rich., Tent. Fl. Abyss. 2: 
426. 1851. T-Protol.: Abyssinia, Pl. Schimp. Abyss., 
sect, 1, 401, 
NOTE: nom. inval., as syn. of Poa sporoboloides A. Rich. 
= Eragrostis japonica (Thunb.) Trin. 


Cathestecum J. Presl, Relig. Haenk. 1(4-5): 294, t. 42. 1830. 
TYPE: Cathestecum prostratum J. Pres|. 
= Bouteloua Lag. 


Cathestecum annuum Swallen, J. Wash. Acad. Sci. 27(12): 
497. 1937. T-Protol.: Mexico: Guerrero: Balsas, on 
open gravelly ground, 1500 ft, 9 Sep 1910, 4. S. 
Hitchcock 6776 [=Amer. Gr. Nat. Herb. 393] [HT: 
US-1720269]. T-Specim.: | of 1. Mexico: Guerrero: 
Balsas, 9 Sep 1910, A. S. Hitchcock 6776 [=Amer. Gr. 
Nat. Herb. no. 393] (HT: US-1720269; IT: MO- 
790204, MO-2959034, US-726629]. 

= Bouteloua griffithsii Columbus. 


Cathestecum brevifolium Swallen, J. Wash. Acad. Sci. 
27(12): 500. 1937. T-Protol.: Mexico: Jalisco: Tequila, 
on thin gravelly soil, 5 Oct 1893,C.G. Pringle 4559 
[HT: US-884043]. T-Specim.: | of 1. Mexico: Jalisco: 
Tequila, on thin gravelly soil, 5 Oct 1893, C.G. 
Pringle 4559 [HT: US-884043; IT: MO-2959035, US- 
821160, US-747968]. 

= Bouteloua diversispicula Columbus. 


Cathestecum brevifolium Swallen var. brevifolium. 
= Bouteloua diversispicula Columbus. 


Cathestecum erectum Vasey & Hack., Bull. Torrey Bot. Club 
11: 37, t. 45. 1884. T-Specim.: | of 1. USA: Texas: 
Bone Spring (Tornillo Cr.), between El Paso and 
Presidio, Aug 1883, V. Havard 2 [LT: US-884020; 
ILT: 884019], LT designated by ?. 
= Bouteloua erecta (Vasey & Hack.) Columbus. 


Cathestecum multifidum Griffiths, Contr. U.S. Natl. Herb. 


14(3): 360, f. 24. 1912. T-Protol.: Mexico: Guerrero: 
Iguala, 9 Sep 1909, Griffiths s.n. [HT: US]. T-Specim.: 


1 of 1. Mexico: Guerro: Iguala, 9 Sep 1909, D. 
Griffiths s.n. [HT: US-2489485,; IT: F, US-81732]. 
= Bouteloua multifidum (Griffiths) Columbus. 


Cathestecum prostratum J. Presl, Relig. Haenk. 1(4-5): 295, 
t. 42. 1830. T-Protol.: Hab. in Mexico, Haenke s.n. T- 
Specim.: | of 1. Mexico, no date, 7) Haenke s.n. [HT: 
PR; IT: LE-TRIN-2209.01, US-2489484 (fragm. ex 
PR)]. 

= Bouteloua griffithsii Columbus. 


Cathestecum stoloniferum (E. Fourn.) Griffiths, Contr. U.S. 
Natl. Herb. 14(3): 362, f. 26. 1912. Basionym: 
Atheropogon stolonifer E. Fourn. 

= Bouteloua reederorum Columbus. 


Cathestecum varium Swallen, J. Wash. Acad. Sci. 27(12): 
498. 1937. T-Protol.: Mexico: Puebla: Tehuacan, on 
dry soilunder mesquite, 9 Aug 1910.4. S. Hitchcock 
6072 [=Amer. Gr. Nat. Herb. 395] [HT: US-1720270]. 
T-Specim.: | of 1. Mexico: Puebla: Tehuacan, on dry 
soil under mesquite, 9 Aug 1910,A4.S8. Hitchcock 6072 
[=Amer. Gr. Nat. Herb, 395] (HT: US-1720270; IT: 
MO-790200, US-726631]. 

= Bouteloua varia (Swallen) Columbus. 


Cenchrus racemosus L., Sp. Pl. 2: 1049. 1753. T-Protol.: 
Habitat in Europa australioris maritimis. T-Specim.: | 
of 2. Herb, Burser 1: 5 [LT: UPS], LT designated by 
Scholz in Cafferty et al., Taxon 49(2): 249 (2000). 2 of 
2. (LT: LINN-1217.183 [1-3]], LT cited by Sherif & 
Siddiqui, Fl. Libya 145: 336 (1988)], but ineffective as 
this is not original material [see Scholz in Cafferty et 
al., Taxon 49(2): 149 (2000). 

= Tragus racemosus (L.) All. 


Centotheca malabarica (L.) Merr., Philipp. J. Sci. 1(Suppl.): 
385. 1906. Basionym: Poa malabarica L. 
NOTE: nom. rej., basionym rej. [ICBN art. 56.2]. 
= Leptochloa fusea (L.) Kunth. 


Ceytosis Munro, J. Linn. Soc. Bot. 6: 54. 1862. 
NOTE: nom. nud. 
= Crypsis Aiton. 


Chaboissaea EF. Fourn., Mexic. Pl. 2: 112. 
TYPE: Chaboissaea ligulata E. Fourn. 
TRTD. BY: Peterson, P. M. 
NHT: Muhlenbergiinae. NO. ACCPTD. SPP.: 4. 
CNSLTD. LIT.: Peterson, P. M. & Y. Herrera A., (1995); 
Peterson, P.M. & C. R. Annable , (1992). 


1886. 


Peterson & 
Basionym: 


Chaboissaea atacamensis (Parodi) P.M. 
Annable, Madrofio 39(1): 19. 1992. 
Muhlenbergia atacamensis Parodi. 

DST.: Arge, Boli. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Peterson, P. M. & C. R. Annable 
(1992); Peterson, P. M. & Y. Herrera A. (1995); 
Peterson, P. M. (2000); Zuloaga, F. O., E. G. Nicora, 
Z. E. Rugolo de Agrasar, O. Morrone, J. Pensiero & 
A. M. Cialdella (1994). 

Muhlenbergia atacamensis Parodi; Muhlenbergia 
atacamensis var. atacamensis, © Muhlenbergia 
atacamensis vat. brachyanthera Parodi. 


SYN:: 


Catalogue of New World Grasses: II. 39 


Chaboissaea decumbens (Swallen) Reeder & C. Reeder, 
Phytologia 65(2): 156. 1988. Basionym: Muhlenbergia 
decumbens Swallen. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Peterson, P. M. 
& C. R. Annable (1992); Peterson, P. M. & Y. 
Herrera A. (1995); Peterson, P. M. (2000). 

SYN.: Muhlenbergia decumbens Swallen. 


Chaboissaea ligulata E. Fourn., Mexic. Pl. 2: 112t. 1. 1886. 
T-Protol.: Mexico: San Luis de Potosi, Virlet. T- 
Specim.: | of 1. Mexico: San Luis Potosi, 1851, Virlet 
d'Aoust s.n. [HT: P; IT: US-A865709 (fragm. ex P)]. 

DST.: Mexi. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); McVaugh, R. (1983); 
Peterson, P. M. & C. R. Annable (1992); Peterson, P. 
M. & Y. Herrera A. (1995); Peterson, P. M. (2000). 

SYN.: Muhlenbergia ligulata (E. Fourn.) Scribn. & Merr. 


Chaboissaea subbiflora (Hitchc.) Reeder & C. Reeder, 
Phytologia 65(2): 156. 1988. Basionym:Muhlenbergia 
subbiflora Hitche. 

DST.: Mexi. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Peterson, P. M. 
& C. R. Annable (1992); Peterson, P. M. & Y. 
Herrera A. (1995): Peterson, P. M. (2000). 

SYN.: Muhlenbergia subbiflora Hitche. 


Chaetaria antillarum (Poir.) P. Beauv. ex Roem. & Schult., 
Syst. Veg. 2: 395. 1817. Basionym: Aristida 
antillarum Poir. 

= Bouteloua americana (L.) Scribn. 


Chascolytrum barbatum (Nees) Kunth, Enum. Pl. 1: 373. 
1833. Basionym: Calotheca barbata Nees. 
= Neesiochloa barbata (Nees) Pilg. 


Chauvinia Steud., Syn. Pl. Glumac. 1: 362. 20-21 Jul 1854 
("1855"). TYPE: Chauvinia chilensis Steud. 
NOTE: later hom., non Bory 1829. 
= Spartina Schreb. 


Chauvinia chilensis Steud., Syn. Pl. Glumac. 1: 362. 1854. T- 
Specim.: | of 1. Chile: Concepcion, no date, /.S.C.D. 
D'Urville s.n. [HT: P; IT: US-A865687 (fragm. ex P)]. 

= Spartina densiflora Brongn. 


Chlorideae fam. Bercht. & J. Presl, Prir. Rostlin 265. 1820. 
NOTE: nom. inval., Art. 33.5. 


Chlorideae trib. Trin., Gram. Unifl. Sesquifl. 117. 1824 [late 
Oct or | Nov]. 


Chloridinae subtrib. J. Presl, Relig. Haenk. 1(4-5): 286. 1830. 
TYPE: Chloris Sw. 
NOTE: as "Chlorideae". 


Chloridoideae subfam. Kunth ex Beilschm., Flora 
16(2): 52, 105. 1833. 
NOTE: ref. to Kunth sect. Chlorideae [apparently 1815], 
supplying description. 
TRTD. BY: Soreng, R. J., G. Davidse, T. S. Filgueiras, E. J. 
Judziewicz, P. M. Peterson, and F. O. Zuloaga. 


Chloridoideae subfam. Burmeist., Handb. Naturgesch. 205. 
1837. TYPE: Chloris Sw. 
NOTE: isonym, as "Chlorideae". 


Chloridopsis Hack., Nat. Pflanzenfam. 2(2): 59. 1887. 
NOTE: insyn. sub Trichloris blanchardiana. 
= Trichloris E. Fourn. ex Benth. 


Chloris Sw., Prodr. 1, 25. 1788. TYPE: Chloris cruciata 
(L.) Sw. 

TRTD. BY: Peterson, P. M. 

NHT: Cynodonteae. NO. ACCPTD. SPP.: 35 SUBSP.: 10. 

CNSLTD. LIT.: Anderson, D. E., (1974). 

SYN.: Actinochloris Steud.; Agrostomia Cerv.,; Apogon 
Steud.; Chloris subg. Pterochloris A. Camus; 
Chlorostis Raf.; Geopogon Steud., Heterolepis 
Ehrenb. ex Boiss.; Phacellaria Willd. ex Steud.; 
Pterochloris (A. Camus) A. Camus. 


Chloris abyssinica Hochst. ex A. Rich., Tent. Fl. Abyss. 2: 
406. 1851 [1850]. T-Specim.: 1 of 3. Ethiopia: Tigray, 
Adoua, 20 Sept 1837, Schimper, pl. Schimp. Abyss., 
sect. 1, 79 [ST: P; IST: K, US-3243722, WAG]. 2 of 3. 
Ethiopia (Djeladjeranne), Schimper, pl.  Schimp. 
Abyss., sect. Ill, 1800 (ST: P; IST: K, US-3243723]. 3 
of 3. Abyssinia (Adoua), Quartin Dillon s.n. [ST: P]. 

= Chloris gayana Kunth. 


Chloris accumbens Hack. ex Arechav., Anales Mus. Nac. 
Montevideo 5: 391. 1896. T-Protol.: Uruguay, Arech. 
s.n. [T: W], as Chloris berroi. 
NOTE: nom. inval., as syn. of Chloris berroi Arechav.; 
Index of Grass Species collat. 1: 391. 
= Chloris berroi Arechav. 


Chloris acuminata Trin., Sp. Gram. 3: t. 305, 1831. T-Protol.: 
Brazil: "Figura ad specimen Brasilianum.". T-Specim.: 
1 of 1. Brazil, G.H. von Langsdorff s.n. [HT: LE- 
TRIN-2075.01 (& fig.); IT: US-80848 (fragm. ex 
LE)]. 
= Eustachys distichophylla (Lag.) Nees. 


Chloris affinis Caro & E.A. Sanchez, Kurtziana 6: 224, f. 2. 
1971. T-Protol.: Argenina: Tucuman, R. Schreiter 
8367. 
DST.: Arge. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 


Chloris alba J. Presl, Relig. Haenk. 1(4-5): 289. 1830. T- 
Protol.: Mexico: Hab. in Mexico, Haenke s.n. T- 
Specim.: 1 of 1. Mexico, no date, 7; Haenke s.n. [T: 
(fragm., US-2830891)]. 

= Chloris virgata Sw. 


Chloris alba var. aristulata Torr., Pacif. Railr. Rep. 4: 155. 
1857. T-Protol.: USA: Texas: Banks of the Upper Rio 
Grande: Oct, [Emory Exped._] s.n. 

= Chloris virgata Sw. 


Chloris albertii Regel, Acta Horti Petrop. 7: 650. 1881. T- 
Protol.: Mongolia occidentali prope Takiansi, Rege/ 
s.n. T-Specim.: | of 1. China: near Takiansi: 1000'alt., 
24 Aug 1878, Regel A. s.n.[HT: LE]. 
= Chloris virgata Sw. 


40) Catalogue of New World Grasses: II. 


Chloris andropogonoides E. Fourn., Mexic. Pl. 2: 143. 
1886. T-Protol.: Mexico: San Luis de Potosi, Virlet 
1462. T-Specim.: | of 1. Mexico: San Luis Potosi, 
1851, Virlet 1462 (HT: P; IT: US-80845 (fragm.)]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Anonymous 
(1986); Beetle, A. A. (1977); Beetle, A. A. (1987); 
Correll, D. S. & M. C. Johnston (1970); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Hitchcock, A. S. (1951). 

SYN.: Chloris tenuispica Nash. 


Chloris angustiflora F. Aresch., Pl. Itin. Eugenia 3: 118. 
1910. T-Protol.: Ecuador: In insula Puna, Andersson, 
N.J. 23. T-Specim.: | of 1. 00/00/1852, 23, Andersson 
{HT: S]. 
= Enteropogon mollis (Nees) Clayton. 


Chloris anisopoda Scribn. ex B.L. Rob., Proc. Amer. Acad. 
Arts 38: 118. 1902. T-Protol.: Ecuador: Galapagos 
Islands: Charles Island., Baur 333. T-Specim.: 1 of 2. 
Galapagos Islands, Jun 1891, G. Baur 333 (ST: GH; 
IST: US-2830887 (fragm.)]. 2 of 2. Galapagos Islands, 
May 1899, RE. Snodgrass & E.G. Heller 459 (ST: 
GH; IST: US-2830885 (fragm.)]. 

= Enteropogon mollis (Nees) Clayton. 


Chloris anoplia (Nees) E. Fourn. ex Hemsl., Biol. Cent.- 
Amer., Bot. 3(19): 558. 1885. Basionym: Leptochloa 
anoplia Nees. 

= Leptochloa anoplia Nees. 


Chloris arenaria Hitchc. & Ekman, Man. Grasses W. Ind. 
131. 1936. T-Protol.: Cuba: Prov. Pinar del Rio: 
collected in pinelands on white sand, La Grifa, 14 Jun 
1920, Ekman 11253 (HT: US-1387449]. T-Specim.: | 
of 1, Cuba: Pinar del Rio: La Grifa, 14 Jun 1920, L. 
Ekman 11253 (HT: US-1387449; IT: F, K, NY- 
70982]. 

DST.: Cari. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Catasus 
Guerra, L. (1985); Catasus Guerra, L. (1997); 
Hitchcock, A. S. (1936). 

SYN.: Chloris eleusinoides var. vestita Greenm. ex Combs. 


Chloris argentina (Hack. ex Stuck.) Lillo & Parodi, Physis 
(Buenos Aires) 4: 180. 1918. Basionym: Chloris 
distichophylla var. argentina Hack. ex Stuck. 

= Eustachys retusa (Lag.) Kunth. 


Chloris argentinensis (Hack. ex Stuck.) Lillo & Parodi, 

Revista Fac. Agron. Veterin. (Buenos Aires) 2: 283. 
1919. Basionym: Chloris distichophylla var. argentina 
Hack. ex Stuck. 

NOTE: hom. illeg. 

E. NAME: Chloris argentina (Hack. ex Stuck.) Lillo & 

Parodi. 
= Eustachys retusa (Lag.) Kunth. 


Chloris aristata (Cerv.) Swallen, N. Amer. Fl. 17(8): 596. 
1939. Basionym: Agrostomia aristata Cerv. 
= Chloris rufescens Lag. 


Chloris aristida Salzm. ex Steud., Syn. Pl. Glumac. 1: 218. 
1854, 
NOTE: nom. inval., as syn. of Gymnopogon foliosus Nees. 
= Gymnopogon foliosus (Willd.) Nees. 


Chloris arundinacea Nees ex Steud., Syn. Pl. Glumac. 1: 
207. 1854. T-Protol.: Legit in Brasilia, Se/low s.n. T- 
Specim.: | of 1. Brazil, no date, F. Se/low s.n. [HT: B; 
IT: BAA-652 (fragm. ex B), US-2830894 (fragm.)]. 

= Chloris elata Desv. 


Chloris bahiensis Steud., Syn. Pl. Glumac. 1: 208. 1854. T- 
Protol.: Bahia[ST: Hb. Moricand]. T-Specim.: | of 1. 
Brazil: Bahia, Salzmann s.n. (Mordicand Herb. 2242) 
[ST: P; IT: US-2639951 (fragm. ex P), K, MO- 
1837389, BAA (fragm. ex MO)]. 

= Eustachys paspaloides subsp. caribaea (Spreng.) 
Nowack. 


Chloris bahiensis Nowack fo. bahiensis. 
= Eustachys caribaea (Spreng.) Herter. 


~ 


Chloris bahiensis fo. glabrescens Hack., Repert. Spec. Nov. 
Regni Veg. 8: 46. 1910. T-Protol.: Paraguay: Sep 
1909, Fiebrig 4575 [T: herb. Hassler]. T-Specim.: | of 
1. Paraguay: Nordl. Paraguay, 22-23 lat., entre Rio 
Apa y Aquidaban, Niederein camp, leiwaldinsel, Sep 
1909, K. Fiebrig 4575 [HT: W; IT: BAA, G, L, US- 
80855 (fragm.)]. 

= Eustachys paspaloides subsp. caribaea 

Nowack. 


(Spreng.) 


Chloris barbata Sw., Fl. Ind. Occid. 1: 200. 1797. 

NOTE: nom. nov., as comb., but the cited basionym is a 
hom. illeg. 

REPLACED NAME: Andropogon barbatus L. 

BLOCKING NAME: Andropogon barbatus L. 

DST.: Arge, Beli, Braz, Cari, Colo, CoRi, Ecua, FrGu, 
Guya, Hond, Mexi, Pana, Para, Peru, Suri, USA, 
Vene. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Bor, N. L. 
(1960); Brako, L. & J. L. Zarucchi (1993); Davidse, 
G., M. Sousa S. & S. Knapp (1998-); Doll, J. C. 
(1878); Fosberg, F. R. (1976); Jorgensen, P. M. & S. 
Leén-Yanez (eds.) (1999); Judziewicz, E. J. (1990); 
Kartesz, J. T. & K. N. Gandhi (1992); Lazarides, M. 
(1961); Lazarides, M. (1972); Phillips, S. (1995); 
Renvoize, S. A. (1984): Rutenberg, C. (1880-1889); 
Smith, L. B., D. C. Wasshausen & R. M. Klein 
(1981); Tovar, O. (1993); Wunderlin, R. P. (1998). 

SYN.: Andropogon barbatus L.; Chloris barbata_ var. 
divaricata Kuntze; Chloris inflata Link; Chloris 
longifolia Steud.; Chloris paraguaiensis Steud.; 
Chloris rufescens Steud. 


Chloris barbata (L.) Nash, Bull. Torrey Bot. Club 25: 443. 
1898. Basionym: Andropogon barbatus L. 
NOTE: hom. illeg. 
E. NAME: Chloris barbata Sw. 
= Chloris elata Desv. 


Chloris barbata var. decora (Nees ex Steud.) Benth., FI. 
Austral. 7: 613. 1878. Basionym: Chloris decora Nees 
ex Steud. 

= Chloris virgata Sw. 


Chloris barbata var. divaricata Kuntze, Revis. Gen. PI. 2: 
771. 1891. T-Specim.: 1 of 1. Indo-China: Annam: 
Turong, feb 1875, Kuntze 164] [HT: NY; IT: K]. 

= Chloris barbata Sw. 


Chloris barbata var. formosana Honda, Bot. Mag. (Tokyo) 
40: 437. 1926. T-Specim.: 1 of 2. Taiwan: Takao, 


Catalogue of New World Grasses: II. 4] 


1905, Nakahara 635 [ST]. 2 of 2. Bachu, 1919, 
Hayata s.n. [ST]. 
= Chloris formosana (Honda) Keng. 


Chloris berazainiae Cat. Guerra, Acta Bot. Cub. 25: 5. 1985. 
T-Protol.: Cuba: Provincia de Santiago de Cuba: 
Alcarraza, Sierra Maestra, 22 Jul 1974, 1. Catasus 553 
{[HT: HAC]. 

NOTE: as Chloris "berazainae". 

DST.: Cari. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Catasus Guerra, L. (1985); Catasus 
Guerra, L. (1997). 


Chloris berroi Arechav., Anales Mus. Nac. Montevideo 1: 
388, t. 44. 1896. T-Protol.: Uruguay: Vera, Dept. 
Soriano, en terrenos secos, aridos, A.B. Berro s.n. T- 
Specim.: | of 1. Uruguay: Soriano, Dec-Apr, M.B. 
Berro 6385 |IT: US-879335]. 

DST.: Arge, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Anonymous 
(1986); Burkart, A. (1969); Cabrera, A. L. (1970); 
Hitchcock, A. S. (1951); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

SYN.: Chloris accumbens Hack. ex Arechav. 


Chloris beyrichiana Kunth, Revis. Gramin. 1: 89, 289. 1829. 
T-Protol.: Brazil: Crescit in Brasiliae graminosis, 
prope Rio Janeiro. T-Specim.: 1 of 1. Brazil: 
Guanabara: prope Rio de Janeiro, no date, H.C. 
Beyrich s.n. [T: (fragm., US-2830902)]. 

= Chloris pycnothrix Trin. 


Chloris beyrichiana Steud., Nomencl. Bot. 1: 353. 1840. 
NOTE: nom. inval., as syn. of Chloris pycnothris Trin. 
1824. 
= Chloris pycnothrix Trin. 


Chloris biflora Regel, Annot. Bot. Ind. Sem. 28. 1863. 
NOTE: nom. inval., as syn. of Chloris radiata (L.) Sw. 
= Chloris radiata (L.) Sw. 


Chloris boliviensis Renvoize, Gram. Bolivia 247, f. 74. 
1998. T-Protol.: Bolivia: Beni: Ballivian, Espiritu, 
Renvoize 4704 (HT: LPB; IT: K]. 

DST.: Boli. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Renvoize, S. A. (1998). 


Chloris brachyathera (Hack. ex Parodi) Herter, Revista 
Sudamer. Bot. 6: 146. 1940. Basionym: Chloris ciliata 
var. brachyathera Hack. ex Parodi. 

NOTE: cf. Gray Herb. Card Cat. 
= Chloris ciliata Sw. 


Chloris — brachystachys Andersson, Naturw. — Reise 
Mossambique 556. 1864. 
REPLACED NAME: Chloris alba J. Presl. 


= Chloris virgata Sw. 


Chloris brandegei (Vasey) Swallen, Amer. J. Bot. 22: 41. 
1935. Basionym: Diplachne brandegei Vasey. 
= Enteropogon brandegei (Vasey) Clayton. 


Chloris breviaristata (Hack. ex Kiitz.) Herter, Revista 
Sudamer. Bot. 6: 146. 1940. Basionym: Chloris 
polydactyla var. breviaristata Hack. ex Kitz. 

NOTE: cfr. Gray Herb. Card Cat. 

NOTE: possibly same as C. ciliata SW. note in Anderson 
1974. 

= Chloris polydactyla var. breviaristata Hack. ex Kiitz. 


Chloris breviglumis C. Wright, Anales Acad. Ci. Med. 
Habana 8: 200. 1871. T-Protol.: Cuba: [Canuela]: 
"sobre piedras de las sabanas y en parages expuestos al 
sol y aridos, a la orilla de los rios.", Wright 1848/error 
for 1548] [HT: GH]. T-Specim.: 1 of 1. Cuba: 
Canuela, 1860-1864, C. Wright 1548 |HT: GH, US- 
80833 (fragm. ex GH & photo); IT: BM, G, NY- 
70893], Collection number cited erroneously as 1848 
in original publication. 

= Chloris cruciata (L.) Sw. 


Chloris brevipila Roseng. & Izag., Bol. Soc. Argent. Bot. 12: 
120, f. 2. 1968. T-Protol.: Uruguay: Tacuarembo: Ruta 
5 & Achar Tacuarembo, 12 Mar 1966, 8. Rosengurtt, 
del Puerto & Marchesi 10173 (HT: MVFA\]. 
= Eustachys brevipila (Roseng. & Izag.) Caro & E.A. 
Sanchez. 


Chloris brevispica Nash, Bull. Torrey Bot. Club 25: 438. 
1898. T-Protol.: USA: Texas: Nueces Co.: 21 Mar 
1894, A.A. Heller 1471. T-Specim.: 1 of 1. USA: 
Texas, Nueces Co., 21 Mar 1894, 4.4. Heller 1471] [T: 
US-747615, US-2474329, US-821282]. 
= Chloris x subdolichostachya Miill. Hal. 


Chloris calvescens Hack., Bull. Herb. Boissier, ser. 2, 4(3): 
279. 1904. T-Protol.: Paraguay: in arenosis regione 
cursus superioris fluminis Apa: Jan, Hassler 8369. T- 
Specim.: | of 1. Paraguay: In arenosis in regione 
cursus superioris fluminis Apa, Hassler 8369 [HT: G; 
IT: BAA, G, MO, US-879348 (fragm. ex G)]. 

= Eustachys calvescens (Hack.) Caro & E.A. Sanchez. 


Chloris canterae Arechay., Anales Mus. Nac. Montevideo |: 
385. 1896. T-Protol.: Uruguay: Abunda en los campos 
graminosos de Paysandu, en tierras bajas, frias, Sr. C. 
B. Cantera. T-Specim.: 1 of 1. Uruguay, Mar-Apr, J. 
Arechavaleta s.n. {[LT: MVM; ILT: BAA, USS- 
879336], LT designated by Rosengurtt & Izaguirre, 
Bol. Soc. Arg. Bot. 12: 124 (1968). 

NOTE: as "Chloris canterai", the ending correctable to ae. 

DST.: Arge, Braz, Para, USA, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 
(1987); Anonymous (1986); Beetle, A. A. (1987); 
Cabrera, A. L. (1970); Correll, D. S. & M. C. 
Johnston (1970); Hitchcock, A. S. (1951); 
Rosengurtt, B., B. R. Arrillaga de Maffei & P. 
Izaguirre de Artucio (1970); Smith, L. B., D. C. 
Wasshausen & R. M. Klein (1981). 

Chloris barbata Sw.; Chloris parodiana Roseng. & 
Izag.; Chloris polydactyla fo. pauciradiata Kurtz; 
Chloris polydactyla fo. stolonifera Parodi. 


SYN:: 


Chloris canterae Arechav. var. canterae. 
DST.: Arge, Braz, Para, USA, Urug. 
LVL. ACCPTC.:: I. 
ALSO ACCPTD. BY: Anderson, D. E. (1974). 
SYN.: Chloris parodiana Roseng. & Izag. 


42 Catalogue of New World Grasses: II. 


Chloris capensis var. bahiensis (Steud.) Parodi, Revista 
Argent. Agron. 20: 26. 1953. Basionym: Chloris 
bahiensis Steud. 

= Eustachys paspaloides 
Nowack. 


subsp. caribaea (Spreng.) 


Chloris capensis var. glabrescens (Hack.) Parodi, Revista 
Argent. Agron. 20: 20. 1953. Basionym: Chloris 
bahiensis fo. glabrescens Hack. 

= Eustachys paspaloides subsp. caribaea (Spreng.) 
Nowack. 


Chloris caribaea Spreng., Syst. Veg. 1: 295. 1825. T-Protol.: 
Ins. Carib., Bertero s.n., Locality perhaps in error, 
never found in the West Indies since the type 
collection, see Nowack, 1995. T-Specim.: 1 of 2. 
"West Indies", no date, Anonymous [IT: US-879338 
(fragm.)]. 2 of 2. Guadeloupe, Bertero s.n. [HT: B; IT: 
US (fragm. ex B)], (probabelmente erro por ser 
desconhecida nas Antilhas). 

= Eustachys paspaloides subsp. caribaea (Spreng.) 
Nowack. 


Chloris castilloniana Lillo & Parodi, Physis (Buenos Aires) 
4: 176, f. 1-2. 1918. T-Protol.: Argentina: Amaicha 
(valle Calchaqui), 1800 m, L. Castillon s.n. T-Specim.: 
| of |. Argentina: Tucuman, Dec 1914, L. Castillon 
3450 (IT: US-1256553], Nom. nud. in Lillo, Fl. de la 
Provinicia de Tucuman, Gramineas, 1916. 

DST.: Arge, Boll. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Burkart, A. 
(1969); Renvoize, S. A. (1998); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 


Chloris castilloniana Lillo & Parodi var. castilloniana. 
LVL. ACCPTC.: 1. 


Chloris castilloniana var. pubescens Caro & E.A. Sanchez, 
Kurtziana 6: 230. 1971. 
DST.: Arge. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 


Chloris caudata Trin. ex Bunge, Enum. Pl. China Bor. 70. 
1833. T-Protol.: Hab. prope Pekinum". T-Specim.: 1 
of 1. N China, 1831, Bunge A. Herb.Ledebour |IT: 
LE]. 

= Chloris virgata Sw. 


Chloris chloridea (J. Presl) Hitche., Proc. Biol. Soc. Wash. 
41: 162. 1928. Basionym: Dinebra chloridea J. Presl. 
= Enteropogon chlorideus (J. Pres!) Clayton. 


Chloris ciliata Sw., Prodr. 25. 1788. T-Protol.: West Indies: 
Jamaica, Swartz s.n. [HT: S]. T-Specim.: | of 1. 
Jamaica, O.P. Swartz s.n. [HT: S; IT: BAA (fragm.), 
US-80821 (fragm. ex S)]. 

DST.: Arge, Beli, Boli, Braz, Cari, Colo, Mexi, Nica, Para, 
USA, Urug, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Allem, A. C. 
& J. F. M. Valls (1987); Anderson, D. E. (1974); 
Anonymous (1986); Balick, M. J., M. H. Nee & D. E. 
Atha (2000); Beetle; A. A. (1977); Beetle, A. A. 
(1987); Breedlove, D. E. (1986); Burkart, A. (1969); 


Catasus Guerra, L. (1985); Correll, D. S. & M. C. 
Johnston (1970); Cowan, C. P. (1983); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994); Doll, J. C. (1878); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J.  Valdes-Reyna (2000); 
Filgueiras, T. S. (1999); Gleason, H. A. & A. 
Cronquist (1968); Gould, F. W (1979); Hitchcock, A. 
S. (1909); Hitchcock, A. S. (1936); Hitchcock, A. S. 
(1951); Lindeman, J. C. & A. L. Stoffers (1963); 
Liogier, H. & L. Martorell (1982); McClure, F. A. 
(1955); Pohl, R. W. (1994); Renvoize, S. A. (1998); 
Rosengurtt, B., B. R. Arrillaga de Maffei & P. 
Izaguirre de Artucio (1970); Sousa S., M. & E. F. 
Cabrera C. (1983). 

Andropogon pubescens Aiton; Chloris brachyathera 
(Hack. ex Parodi) Herter; Chloris ciliata var. texana 
Vasey; Chloris nashii A. Heller; Chloris propinqua 
Steud.; Chloris texana (Vasey) Nash; Cynodon 
ciliatus (Sw.) Raspail. 


SYN.: 


Chloris ciliata var. brachyathera Hack., Bol. Acad. Ci. 
(Cordoba) 16: 257. 1900. T-Protol.: Argentina. 
NOTE: nom. nud.; equals Chloris ciliata var. brachyathera 
Hack. ex Parodi. 
= Chloris ciliata var. ciliata. 


Chloris ciliata var. brachyathera Hack. ex Parodi, Revista 
Fac. Agron. Veterin. (Buenos Aires) 2: 271. 1919. T- 
Protol.: Argentina: Cordoba: San Roque, Sierra Chica, 
12 Jan 1881, G. Hieronymus. 

= Chloris ciliata var. ciliata. 


Chloris ciliata fo. breviseta Hack. 
DST.: Arge. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 


Chloris ciliata Sw. fo. ciliata. 
LVL. ACCPTC.: |. 


Chloris ciliata var. texana Vasey, U.S.D.A. Div. Bot. Bull. 
12(1): t. 30. 1890. T-Protol.: USA: Texas: near 
Brownsville, Nealley s.n. [HT: US]. T-Specim.: | of 1. 
USA: Texas: near Brownsville, 1889, G.C. Nealley 
s.n. [HT: US-879016]. 

= Chloris ciliata Sw. 


Chloris clandestina Scribn. & Merr., Bull. Div. Agrostol., 
U.S.D.A. 24: 25. 1901. 
REPLACED NAME: Gymnopogon longifolius E. Fourn. 
BLOCKING NAME: Chloris longifolia Steud. 
= Enteropogon chlorideus (J. Pres!) Clayton. 


Chloris compressa DC., Cat. Pl. Horti Monsp. 94. 1813. T- 
Protol.: [Cult.; at Montpellier from S$. America]. T- 
Specim.: | of 1. "South America", Sep 1828 and 20 
Aug 182, A.P. de Candolle 4 |T: (fragm., US-2830890, 
US-80851)], Cultivated at montpellier southern france. 

= Chloris virgata Sw. 


Chloris confertifolia Trin., Neue Entdeck. Pflanzenk. 2: 74. 
1821. T-Protol.: "Hab. in insula Sanctae Catharinae.". 
T-Specim.: | of 1. Brazil: Santa Catarina, G.H. von 
Langsdorff s.n. (HT: LE?; IT: US-879359 (fragm.)]. 

= Eustachys distichophylla (Lag.) Nees. 


Catalogue of New World Grasses: II. 43 


Chloris consanguinea Kunth, Revis. Gramin. 1: 89. 1829. T- 
Protol.: Brasilia. 
NOTE: nom. nud. 
= Chloris elata Desv. 


Chloris crinita Lag., Varied. Ci. 2(4): 143. 1805. T-Specim.: 
1 of |. Philippines: Ex Philippinis Insulis [from Nee 
letter] [error for Mexico], Nee [HT: MA; IT: US- 
80822 (fragm. ex MA & drawing by Parodi)]. 

= Trichloris crinita (Lag.) Parodi. 


Chloris crinita Salzm. ex DOll, Fl. Bras. 2(3): 64. 1878. 
NOTE: nom. inval., as syn. of Chloris radiata (L.) Sw. 
E. NAME: Chloris crinita Lag. 
= Chloris radiata (L.) Sw. 


Chloris cruciata (L.) Sw., Prodr. 25. 1788. Basionym: 
Agrostis cruciata L. 

DST.: Cari. 

LVL. ACCPTC.:: |. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Anderson, D. 

E. (1974); Catasus Guerra, L. (1985); Catasus 
Guerra, L. (1997); Hitchcock, A. S. (1909); 
Hitchcock, A. S. (1936). 
Agrostis cruciata L.; Chloris breviglumis C. Wright; 
Chloris humboldtiana Steud.;Cynodon cruciatus (L.) 
Raspail; Rhabdochloa cruciata (L.) P. Beauv.; Vilfa 
cruciata (L.) P. Beauv. 


SYN:: 


Chloris cryptostachya Steud. ex J.A. Schmidt, Beitr. Fl. Cap 
Verd. Ins. 148. 1852. T-Protol.: Ethiopia: It. nubic. 
Cordofan, Hochstetter 23 (HT: P; IT: K]. 

= Enteropogon prieurii (Kunth) Clayton. 


Chloris cubensis Hitche. & Ekman, Man. Grasses W. Ind. 
131. 1936. T-Protol.: West Indies: Cuba: Oriente: 
collected at Gamboa, 26 Aug 1922,E.L. Ekman 14960 
[HT: US-1387446]. T-Specim.: | of 1. Cuba: Oriente: 
Gamboa, 26 Aug 1922, E.L. Ekman 14960 [HT: US- 
1387446; IT: HAC], CS. 

= Chloris sagraeana subsp. cubensis (Hitchc. & Ekman) 
Cat. Guerra. 


Chloris cucullata Bisch., Ann. Sci. Nat., Bot., ser. 3, 19: 
357. 1853. T-Protol.: Habitat Mexico boreali. Semina 
in Prov. Tamaulipas prope Matamoros lecta absque 
nomine, misit Dr. Engelmann 1849, Anon. T-Specim.: 
1 of 1, Cultivated in Horto Botanico Heidelbergensis 
from seed collected near Matamoros, Tamaulipas, 
Mexico, and sent by Englemann in 1849, 1849,Anon. 
[T]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Anonymous 
(1986); Beetle. A. A. (1977); Beetle, A. A. (1987); 
Correll, D. S. & M. C. Johnston (1970); Davidse, G., 
M. Sousa S. & S. Knapp (1998-); Espejo Serna, A., 
A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Great Plains Flora Association (1986); Hitchcock, A. 
S. (1951); Wunderlin, R. P. (1998). 


Chloris curtipendula Michx., Fl. Bor.-Amer. 1: 59. 1803. T- 
Protol.: Hab. in aridis regionis [llinoensis ad Wabast et 
in rupibus ad prairie du rocher, 1795, Michaux s.n. T- 
Specim.: | of 1. USA: hauteurs du Missouri et Poste 
Vicenne, Michaux s.n. [HT: P; IT: US (fragm.)]. 
= Bouteloua curtipendula (Michx.) Torr. 


Chloris cynodon Trin., Gram. Unifl. Sesquifl. 229, 1824. 


NOTE: nom. illeg. superfl. 
CORRECT NAME: Panicum dactylon L. 
= Cynodon dactylon (L.) Pers. 


Chloris dandyana C.D. Adams, Phytologia 21: 408. 1971. 
REPLACED NAME: Andropogon barbatus L. 
BLOCKING NAME: Chloris barbata Sw. 
= Chloris elata Desy. 


Chloris decora Nees ex Steud., Syn. Pl. Glumac. 1: 205. 
1854. T-Protol.: Nepal, Royle hrbr. 87, see notes. T- 
Specim.: | of 1. Nepal, Royle 87 [IT: K]. 

= Chloris virgata Sw. 


Chloris digitaria Kunth, Nov. Gen. Sp. 1: 168. 1815 [1816]. 
T-Protol.: Peru: Crescit in temperatis inter pago 
Chontali et Colazey, 600 hexap., prov. Jaen de 
Eracamoros, Humb. & Bonpland s.n. T-Specim.: | of 
1. Peru: Cajamarca, Aug, &.W.H.A. von Humboldt, 
A.J.A, Bonpland s.n. |TV: US-2830889 (fragm.)]. 

= Leptochloa virgata (L.) P. Beauv. 


Chloris distichophylla Lag., Gen. Sp. Pl. 4. 1816. T-Protol.: 
Habitat in planitie Bonariensi et in Chilensi Regno. T- 
Specim.: | of 1. Argentina: Bonariensi planitie,L. Née 
s.n. [HT: MA-252893; IT: BA, US-80825 (fragm. ex 
MA)]. 

= Eustachys distichophylla (Lag.) Nees. 


Chloris distichophylla var. acuminata (Trin.) Hack., Bull. 
Herb. Boissier, ser. 2, 4: 279. 1903. Basionym: Chloris 
acuminata Trin. ; 

= Eustachys distichophylla (Lag.) Nees. 


Chloris distichophylla var.argentina Hack. ex Stuck., Anales 
Mus. Nac. Buenos Aires 11: 113. 1904. T-Protol.: 
Argentina, Stuckert 1/045 [T: Herb. Arg. 8783, 10188, 
11045]. T-Specim.: | of 1. Argentina: Cordoba: 
alrededores de la ciudad, 2 Jan 1902, 7./.V. Stuckert 
11045 |ST: BAA; IST: US-80850 (fragm.)]. 

= Eustachys distichophylla (Lag.) Nees. 


Chloris distichophylla var. genuina Hack., Anales Mus. Nac. 
Hist. Nat. Buenos Aires 11: 113. 1904. 
NOTE: nom. inval., see ICBN Art. 24.3. 
CORRECT NAME: Chloris — distichophylla _ var. 
distichophylla. 
= Eustachys distichophylla (Lag.) Nees. 


Chloris divaricata R. Br., Prodr. 186. 1810. T-Specim.: | of 
2. Australia: Queensland: Port Curtis Dist: Broad 
Sound, Sep 1802, Brown 6244 [ST: BM; IST: E, Kk, 
MEL]. 2 of 2. Australia: "Port I, Keppel Bay, 
Whorlwater Bay", R. Brown [LT: BM], LT designated 
by Anderson, Brigham Young Univ. Sci. Bull., Biol. 
Ser., 19(2): 85 (1974). 

DST.: USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Lazarides, 
M. (1972). 


Chloris divaricata R. Br. var. divaricata. 
DST.: USA. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Lazarides, M. (1972). 


Chloris divaricata var. minor J.M. Black, Trans. & Proc. 
Roy. Soc. South Australia 37: 124, t. 5. 1913. T- 
Specim.: | of 1. Australia: South Australia: 


44 Catalogue of New World Grasses: II. 


Oodnadatta, 15 Dec 1913, Staer s.n. [HT: AD; IT: K, 
MEL]. 
= Chloris pectinata Benth. 


Chloris divaricata var. muelleri Domin, Repert. Spec. Nov. 
Regni Veg. 9: 553. 1911. T-Specim.: | of 1. Australia: 
Western Australia: Ereman Prov: Sturt's Creek, Feb 
1856, Mueller s.n. (HT: K; IT: MEL]. 

= Chloris pectinata Benth. 


Chloris dubia Kunth, Nov. Gen. Sp. 1: 169. 1815 [1816]. T- 
Protol.: Mexico: crescit in calidis Provinciae Jaen de 
Bracamoros prope Tomependa, alt. 207 hex. Floret 
Augusto, Humboldt & Bonpland s.n. T-Specim.: | of 
1. Mexico: Distrito Federal, Aug, A.J.A. Bonpland 
4158 [HT: P; IT: BAA-6751 (fragm. ex P), US-865875 
(fragm. ex P)]. 

= Leptochloa dubia (Kunth) Nees. 


Chloris durandiana Schult., Mant. 2: 341. 1824. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Chloris gracilis P. Durand. 
= Chloris radiata (L.) Sw. 


Chloris dusenii Ekman, Ark. Bot. 10(11): 26, t. 4, f. 2, t. 6, f. 
14. 1911. T-Protol.: Brasil: Prov. Parana, Serrinha, in 
campo subhumido, 9 Dec 1908, P. Dusén 7335. T- 
Specim.: 1 of |. Brazil: Parana: Serrhina, in campo 
subhumido, 9 Dec 1908, P. Dusén 7335 [HT: S?; IT: 
BAA, L, NY, US-601103, US-80828 (fragm. ex S)]. 

= Eustachys ulignosa (Hack.) Herter. 


Chloris ekmanii Hitche., Man. Grasses W. Ind. 129. 1936. 
T-Protol.: Cuba: Matanzas: Jaguey Grande: collected 
in pastures at Babujal, 2 August 1923, E.L. Ekman 
16954 [HT: US-1387451]. T-Specim.: 1 of 1. Cuba: 
Matanzas, 2 Aug 1923, E.L. Ekman 16954 (HT: US- 
1387451]. 

DST.: Cari. 

LVL. ACCPTC.:: |. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Anderson, D. 
E. (1974); Catasus Guerra, L. (1985); Catasus 
Guerra, L. (1997); Hitchcock, A. S. (1936). 


Chloris elata Desv., Mem. Soc. Agric. Angers 1: 177. 1831. 
T-Protol.: Habitat in Brasilio. 
NOTE: also in Opusc. Sci. Phys. Nat. 73 [reprint] 1831- 
1833. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Kartesz, J. T. & K. N. Gandhi 
(1992); Wunderlin, R. P. (1998). 

Andropogon barbatus L.; Andropogon polydactylon 
L.; Chloris arundinacea Nees ex Steud.; Chloris 
barbata (L.) Nash; Chloris consanguinea Kunth; 
Chloris dandyana C.D. Adams; Chloris polydactyla 
(L.) Sw. 


SYN:: 


Chloris elegans Kunth, Nov. Gen. Sp. 1: 166, t. 49. 1815 
[1816]. T-Protol.: Crescit iisdem fere locis inter 
Mexico et Queretaro, floret} Aprili, Septembri, 
Humboldt & Bonpland 4194, T-Specim.: 1 of 1. 
Mexico: between Mexico City and Queretaro, 
Humboldt & Bonpland 4194 (HT: P; IT: US-80853 
(fragm.)]. 

= Chloris virgata Sw. 


Chloris eleusinoides Griseb., Fl. Brit. W. 1. 539. 1864. T- 
Specim.: | of 2. Jamaica, no date, H.R. Wullschlaegel 


g.n. [ST: US-80829 (fragm.)]. 2 of 2. Antigua, March 
gn. [ST]. 
= Chloris sagraeana A. Rich. 


Chloris eleusinoides var. vestita Greenm. ex Combs, Trans. 

Acad. Sci. St. Louis 7: 477, t. 39. 1897. T-Protol.: 
West Indies: Cuba: high grassy river banks of Rio 
Damuji at Rodas: 17 Sep 1895, R. Combs 631. T- 
Specim.: 1 of 1. Cuba: Las Villas, Cienfuegas Mun., 
17 Sep 1895, R. Combs 631 [T: (fragm., US-80830)]. 

CORRECT NAME: Chloris arenaria Hitche. & Ekman. 

= Chloris arenaria Hitchc. & Ekman. 


Chloris elongata Poir., Encycl., Suppl. 2: 236. 1811. T- 
Specim.: | of 1. Indonesia: Lesser Sunda Is., Timor I., 
no date, C. Gaudichaud-Beaupre s.n. [HT: Fl; IT: L, 
US-1912444 (fragm.)]. 
= Chloris truncata R. Br. 


Chloris equitans Trin., Sp. Gram, 3: t. 309, 1831, T-Protol.: 
Figura ad specimen Capense. T-Specim.: | of 1. [HT: 
LE-TRIN-2089.01]. 

= Eustachys paspaloides (Vahl) Lanza & Mattei. 


Chloris exilis Renvoize, Kew Bull. 37(2): 323, f. 2. 1982. T- 
Protol.: Brazil: Bahia: Senhor do Bonfim-Juazeiro, 26 
Feb 1974, Harley et al. 16363 [HT: CEPEC; IT: K]. 
DST.: Braz. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Filgueiras, T. S. (1999); Renvoize, S. 
A. (1982); Renvoize, S. A. (1984). 


Chloris fasciculata Schrad. ex Schult., Mant. 2: 339. 1824. T- 
Protol.: Brazil: "In Brasilia. Princeps  Sereniss, 
Maximil. Neowidens.". T-Specim.: 1 of 1. Brazil: 
Minas Gerais, 1817-1821, H.W. Schott 4858 [IT: US- 
80854]. 

= Eustachys distichophylla (Lag.) Nees. 


Chloris filiformis Poir., Encycl., Suppl. 2: 237. 1811. T- 
Protol.: [cult.; at Paris, the source unknown][T: P]. T- 
Specim.: | of 1. [IT: L]. 

NOTE: Poir replaced this with C. tenuis (Nov. 1817), but C. 
filiformis (Vahl) Poir was relaced in Jul 1817 by 
Leptochloa cynosuroides Roemer & Schult. 

= Bouteloua simplex Lag. 


Chloris floridana (Chapm.) Alph. Wood, Amer. Bot. FI. (ed. 
2) 407. 1871. Basionym: Eustachys floridana Chapm. 
= Eustachys floridana Chapm. 


Chloris foliosa Willd., Sp. Pl. 4: 924. 1806. T-Protol.: W.L.: 
Virgin Islands: "Habitat in insula St. Thomas 
Americes (v.s.), Ventenat s.n. T-Specim.: | of 1. 
Virgin Islands: St. Thomas, Ventenat s.n. [HT: B, US 
(fragm.)]. 

= Gymnopogon foliosus (Willd.) Nees. 


Chloris gabrielae Domin, Biblioth. Bot. 85: 368, f. 83. 1915. 
T-Specim.: | of |. Australia: Queensland: Burke Dist: 
am Flinders River,, Feb 1910, Domin s.n. [HT: PR]. 

NOTE: as Chloris "Gabrielae". 
= Chloris virgata Sw. 


Chloris gayana Kunth, Revis. Gramin. |: 89. 1829. 
NOTE: nom. nud. 
CORRECT NAME: Chloris gayana Kunth. 
= Chloris gayana Kunth. 


Catalogue of New World Grasses: II. 45 


Chloris gayana Kunth, Revis. Gramin. 2: 293, t. 58. 1830. T- 
Specim.: | of 2. Senegal, 4 Feb 1823, Doellinger 21 
[ST: K, P]. 2 of 2. Senegal, 4 Feb 1823, Herb. Gay 40 
[ST: K, P]. 
NOTE: as nom. nud. in Kunth, Rev. Gram. 1(6): 89. 1829. 
DST.: Arge, Boli, Braz, Cari, Chil, Colo, CoRi, Ecua, ElSa, 
Guat, Peru, Suri, USA, Urug, Vene. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Beetle, A. 
A. (1977); Beetle, A. A. (1987); Berendsohn, W. G. 
& A. E. Araniva de Gonzalez (1991); Bor, N. L. 
(1960); Breedlove, D. E. (1986); Burkart, A. (1969); 
Cabrera, A. L. (1970); Catasus Guerra, L. (1985); 
Catasus Guerra, L. (1997); Correll, D. S. & M. C. 
Johnston (1970); Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Edgar, E., H. E. 
Connor & J. E. Shand (1991); Espejo Serna, A., A. 
R. L6pez-Ferrari, and J. Valdes-Reyna (2000); 
Filgueiras, T. S. (1991); Filgueiras, T. S. (1999); 
Gibbs Russell, G. E, W. G. Welman, E. Reitief, K. L. 
Immelman, G. Germishuizen, B. J. Pienaar, M. van 
Wyk, and A. Nicholas (PRECIS) (1987); Gould, F. 
W. & R. Moran (1981); Hickman, J. C. (1993); 
Hitchcock, A. S. (1936); Hitchcock, A. S. (1951); 
Jorgensen, P. M. & S. Leon-Yanez (eds.) (1999); 
Judziewicz, E. J. (1990); Lazarides, M. (1972); 
Marticorena, C. & M. Quezada (1985); McClure, F. 
A. (1955); McVaugh, R. (1983); Molina R., A. 
(1975); Munz, P. A. & D. D. Keck (1959); Munz, P. 
A. (1974); Phillips, S. (1995); Pohl, R. W. (1980); 
Pohl, R. W. (1994); Renvoize, S. A. (1988); 
Renvoize, S. A. (1998); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 
Small, J. K. (1933); Smith, L. B., D. C. Wasshausen 
& R.M. Klein (1981); Tovar, O. (1993); Wunderlin, 
R. P. (1998); Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 

Chloris abyssinica Hochst. ex A. Rich.; Chloris 
gayana fo. genuina Maire & Weiller; Chloris gayana 
fo. oligostachys (Barratte & Murb.) Maire & Weiller; 
Chloris gayana Kunth; Chloris gayana_ subsp. 
oligostachys Barratte & Murb.; Chloris glabrata 
Andersson; Chloris multiradiata var.ragazzii Pirotta; 
Chloris repens Hochst.; Eustachys gayana Mundy. 


SYN:: 


Chloris gayana fo. genuina Maire & Weiller, Fl. Afrique N. 
2: 204. 1953. 
NOTE: nom. inval., see ICBN Art. 24.3. 
= Chloris gayana Kunth. 


Chloris gayana fo. oligostachys (Barratte & Murb.) Maire & 
Weiller, Fl. Afrique N. 2: 204. 1953. Basionym: 
Chloris gayana subsp. oligostachys Barratte & Murb. 

= Chloris gayana Kunth. 


Chloris gayana subsp. oligostachys Barratte & Murb., Acta 
Univ. Lund. 36: 8, t. 13. 1900. T-Protol.; Tunisia 
septentrionale, Entre Menzelbou-Zelfaet Soliman 
(Miss. bot. 1883). 

= Chloris gayana Kunth. 


Chloris glabrata Andersson, Naturw. Reise Mossambique 2: 
557. 1864. T-Protol.: Standort: auf feuchtem sandigen 
und moorigen Boden; Wiesengrund bei Tette, 17° S.B., 
120 Meilen im Innern des Landes, Kohlen- 
Sandsteinfrmation, Peters in 1842-1848. 


= Chloris gayana Kunth. 


Chloris glabrescens (Hack.) Roseng. & Izag., Bol. Soc. 
Argent. Bot. 12: 123. 1968. Basionym: Chloris 
bahiensis fo. glabrescens Hack. 

= Eustachys paspaloides subsp. caribaea (Spreng.) 
Nowack. 


Chloris glauca (Chapm.) Alph. Wood, Amer. Bot. FI. (ed. 2) 
407. 1871. Basionym: Eustachys glauca Chapm. 
= Eustachys glauca Chapm. 


Chloris glaucescens Steud., Syn. Pl. Glumac. 1: 206. 1854. 
T-Protol.: West Indies: Guadeloupe, Duschaissing s.n. 
[HT: P(fragm., US)]. T-Specim.: | of 1. Guadeloupe, 
1852, E.P. de F. Duchassaing 13 (HT: P (fragm., US- 
79823)]. 
= Chloris radiata (L.) Sw. 


Chloris gracilis P. Durand, Quib. Chlor. Spec. 10. 1808. T- 
Specim.: | of 1. Jamaica, no date, Anonymous 396 [T: 
(fragm., US-2830899)]. 

= Chloris radiata (L.) Sw. 


Chloris gracilis Kunth, Nov. Gen. Sp. 1: 168. 1815 [1816]. 
T-Protol.: Peru: Crescit in calidis Prov. Jaen de 
Bracamores pr. Tomependa, alt. 207 hex. 

NOTE: hom. illeg. 
E. NAME: Chloris gracilis P. Durand. 
= Leptochloa virgata (L.) P. Beauv. 


Chloris grandiflora Roseng. & Izag., Bol. Soc. Argent. Bot. 
12: 124, f. 3. 1968. T-Protol.: Uruguay: Florida: 
Estancia Rincén de Santa Elena, Picada de Castro, 
Arroyo Mansavillagra, Nov 1946, Gallinal & 
Rosengurtt PE-5790 [HT: MVFA]. 

= Chloris canterae var. grandiflora (Roseng. & Izag.) 
D.E. Anderson. 


Chloris guaranitica Parodi, Revista Argent. Agron. 28: 106, 
f. 2. 1961 [1962]. T-Protol.: Argentina: Corrientes: 
Dept. Mbrerucuya: Estancia St. Maria, 7-11-1956, 
Troels Myndel Pedersen 3742. T-Specim.: | of 1. 
Argentina: Corrientes: Mburucuya, Santa Maria, 7-11 
[II?]-1956, Troels Myndel Pedersen 3742 (IT: K, L]. 

= Chloris orthonoton Doll. 


Chloris halophila Parodi, Revista Argent. Agron. 12(1): 45. 
1945. T-Protol.: Argentina: Buenos Aires: Pergamino, 
Parodi 2769. T-Specim.: | of 1. Argentina: Buenos 
Aires, 20 Feb 1920, L.R. Parodi 2769 [IT: US- 
1256554]. 

DST.: Arge, Boli, Braz, Ecua, Para, Peru. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Brako, L. & 
J. L. Zarucchi (1993); Burkart, A. (1969); Cabrera, 
A. L. (1970); Foster, R. C. (1958); Jorgensen, P. M. 
& C. Ulloa U. (1994); Jorgensen, P. M. & S. Leén- 
Yanez (eds.) (1999); Renvoize, S. A. (1998); Tovar, 
0. (1993); Zuloaga, F. O., E. G. Nicora, Z. E. Rigolo 
de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 


Chloris halophila Parodi var. halophila. 
LVL. ACCPTC.: 1. 


Chloris halophila var. humilis Christa Miller, Feddes 
Repert. 94(9-10): 627, f. 4. 1983. T-Protol.: Peru: 
Cusco: Urubamba: Tal des Rio Urubamba unterhalb 


46 Catalogue of New World Grasses: II. 


Pisac, Hang, unter Stauchern, 28 May 1972, Mueller & 
Mueller 2634 [HT: LZ; IT: SMF}. 
DST.: Arge, Boli, Ecua, Peru. 
LVL. ACCPTC:: I. 
ALSO ACCPTD. BY: Muller, C. (1983); Zuloaga, F. O., E. 
G. Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994), 


Chloris humboldtiana Steud., Syn. Pl. Glumac. 1: 205. 1854. 
= Chloris cruciata (L.) Sw. 


Chloris humilis Kunth, Revis. Gramin. 1: 89. 1829. T-Protol.: 
Brasilia: Crescit prope Rio-Janeiro. T-Specim.: | of 2. 
Brazil: Guanabara, 4. Richard s.n. [T: (fragm., US- 
2830900)]. 2 of 2. 

= Chloris pycnothrix Trin. 


Chloris inermis Trin., Gram. Unifl. Sesquifl. 232. 1824. 
REPLACED NAME: Chloris polystachya Lag. 
= Chloris submutica Kunth. 


Chloris inflata Link, Enum. Hort. Berol. Alt. 1: 105, 1821. T- 
Protol.: Hab. in California [probably from Mex. acc. to 
A.S.H.] E. seminibus, D. Chamisso s.n. [HT: B], typ. 
cult., seed come from California, probably from 
Mexico. T-Specim.: | of 1. USA: California; Cult. en 
Berlin, no date, 1.4. von Chamisso s.n. [HT: B 
(fragm., US-2830897)]. 

= Chloris barbata Sw. 


Chloris intermedia A. Rich., Tent. Fl. Abyss. 2: 407. 1850. 
T-Protol.: Abyssinia: Chire, Dillon & Petit s.n. [HT: 
PI. 
= Chloris pycnothrix Trin. 


Chloris latisquamea Nash, Bull. Torrey Bot. Club 25: 439. 
1898. 
REPLACED NAME: Chloris verticillata var. intermedia 
Vasey. 
BLOCKING NAME: Chloris intermedia A. Rich. 
= Chloris * subdolichostachya Mill. Hal. 


Chloris leptantha Hitche. ex Urb., Symb. Antill. 7: 166. 
1912. T-Protol.: "Hab. in insulis orae Venezuelensi 
adjacentibus Bonaire: 19 Feb 1885, WFR. Suringar 
s.n. [HT: US690357]. T-Specim.: | of 1. Netherlands 
[Antilles]: Bonaire, Fontein, 19 Feb 1885, WFR, 
Suringar 19 [HT: L, US-690357], TS. 

= Enteropogon mollis (Nees) Clayton. 


Chloris leptostachya Hochst. ex A. Rich., Tent. Fl. Abyss. 2: 
407. 1851. T-Protol.: Abyssinia: ad agrorum margines 
et in incultis prope Adoam, Schimper, pl. Schimp. 
Abyss., sect. I], 951 [HT:P. IT: BM, G, K, P]. T- 
Specim.: | of 1. Ethiopia: Tigray, Adua, 29 Oct 1838, 
G.H.W. Schimper 951 (IV: WAG; IST: US-3243725]. 

= Chloris pycnothrix Trin. 


Chloris leptostachya var. intermedia (A. Rich.) T. Durand & 
Schinz, Consp. Fl. Afr. 5: 861. 1895. Basionym: 
Chloris intermedia A. Rich. 

= Chloris pycnothrix Trin. 


Chloris longifolia Steud., Syn. Pl. Glumac. 1: 205. 1854. T- 
Protol.: Cuming 696 [HT: P(fragm., US)]. T-Specim.: 
1 of 1. Phillipines: Luzon I.: Cavite Prov.: Corregidor 
Island, H. Cumming 696 [T: L; IT: US-80840 
(fragm.)]. 
= Chloris barbata Sw. 


Chloris longifolia (E. Fourn.) Vasey, Contr. U.S. Natl. Herb. 
1(8): 284, t. 19. 1893. Basionym: Gymnopogon 
longifolius E. Fourn. 

NOTE: hom. illeg. 
E. NAME: Chloris longifolia Steud. 
= Enteropogon chlorideus (J. Presl) Clayton. 


Chloris luetzelburgii Hitchc., Proc. Biol. Soc. Wash. 36: 197. 
1923. T-Protol.: Brazil: Parahyba do Norte: Serra do 
Borborema, Apr, Luetzelburg 1/245] |HT: US- 
1127373]. T-Specim.: | of 1. Brazil: Paraiba, Apr, P. 
von Luetzelburg 1245] [HT: US-1127373]. 

= Enteropogon mollis (Nees) Clayton. 


Chloris madagascariensis Steud., Syn. Pl. Glumac. |: 206. 
1854. T-Protol.: Le Jolis s.n. [HT: PJ], in 1848. T- 
Specim.: | of 1. Madagascar, 1849, 4.F. Le Jolis 7 
[HT: P (fragm., US-80841)]. 

= Chloris virgata Sw. 


Chloris maritimus (Kunth) Trin., Gram. Unifl. Sesquifl. 235. 
1824. Basionym: Cynodon maritimus Kunth. 
= Cynodon maritimus Kunth. 


Chloris meccana Hochst. ex Steud., Syn. Pl. Glumac. 1: 205. 
1854. T-Specim.: | of 1. 11 Feb 1836, Schimper 802 
{HT: G, US-1127159; IT: L]. 
= Chloris virgata Sw. 


Chloris megastachya Schrad. ex Schult., Mant. 2: 340. 1824. 
T-Protol.: Australia. 
= Chloris truncata R. Br. 


Chloris mendocina Phil., Anales Univ. Chile 36: 208. 1870. 
T-Specim.: | of 1. Argentina: Mendoza, no date, R.A. 
Philippi s.n. [T: P (fragm., US-80842)]. 

= Trichloris crinita (Lag.) Parodi. 


Chloris mollis (Nees) Swallen, N. Amer. Fl. 17: 596. 1939. 
Basionym: Gymnopogon mollis Nees. 
= Enteropogon mollis (Nees) Clayton. 


Chloris monostachya Michx., Fl. Bor.-Amer. 1: 59. 1803. T- 
Protol.: USA: "Hab. in sylvis Carolinae inferioris.", 
Michaux s.n. 

= Ctenium aromaticum (Walter) Alph. Wood. 


Chloris morales-coelloi Leon ex Britton, Bull. Torrey Bot. 
Club 53: 458. 1926. T-Protol.: Cuba: Oriente: 
Cajobabo, not far from mouth of the Jojo River, Ledn 
12320 [HT: US]. T-Specim.: | of 1. Cuba: Oriente: 
southern Baracoa region, not far from the mouth of the 
Joj6 River, Cajobabo, 17 Jul 1924-4 Aug 1924, Bro. 
Leon 12320 (IT: NY, US-1258978]. 

= Chloris sagraeana A. Rich. 


Chloris mucronata Michx., F1. Bor.-Amer. |: 59. 1803. T- 
Protol.: USA: "Hab. in cultis Carolinae.", Michaux s.n. 
T-Specim.: | of 1. USA: in cultis Carolinae, Michaux 
gn. [HT: PJ. 

= Dactyloctenium aegyptium (L.) Willd. 


Chloris multiradiata Hochst., Flora 38: 204. 1855. T-Protol.: 
Abyssinia [1853], Schimper, Hrbr. abyss. Buch. 486 
{[HT:STR; IT:K]. 

= Chloris virgata Sw. 
Chloris multiradiata var. ragazzii Pirotta, Annuario Reale Ist. 


Bot. Roma 6: 157. 1896. 
= Chloris gayana Kunth. 


Catalogue of New World Grasses: II. 47 


Chloris nashii A. Heller, Muhlenbergia 5: 120. 1909, 
NOTE: nom. illeg. superfl.; equals Chloris texana (Vasey) 
Nash. 
REPLACED NAME: Chloris ciliata var. texana Vasey. 
CORRECT NAME: Chloris texana (Vasey) Nash. 
= Chloris ciliata Sw. 


Chloris nealleyi Nash, Bull. Torrey Bot. Club 25: 435. 1898. 
T-Protol.: Texas, G.C. Nealley s.n. 
NOTE: nom. illeg. superfl.; as Chloris "Nealleyi". 
CORRECT NAME: Chloris texensis Nash. 
= Chloris texensis Nash. 


Chloris neglecta Nash, Bull. Torrey Bot. Club 22(10): 423- 
424. 1895. T-Protol.: USA: Coll. near Orange Bend, 
Lake Co., Florida, in low pine lands,G. V. Nash 2/49 
[HT: US]. T-Specim.: | of 1. USA: Florida, Lake Co., 
1 Jul 1895-10 Jul 1895, G.V. Nash 2149 (ST: US- 
252261]. 
= Eustachys neglecta (Nash) Nash. 


Chloris notocoma Hochst., Flora 38: 204. 1855. T-Protol.: 
Abyssinia [1853], Schimper, Hb. abyss. Hohenacker 
2125 {IT: G, P]. 
= Chloris virgata Sw. 


Chloris obtusifolia Desv., Mem, Soc. Agric. Angers 1: 177. 
1831. T-Protol.: Crescit in Brasilio. 
NOTE: hom. illeg.; also in Opusc. Sci. Phys. Nat. 73 
[reprint] 1831-1833. 
E, NAME: Chloris obtusifolia Trin. 
= Chloris pyenothrix Trin. 


Chloris orthonoton Doll, Fl. Bras. 2(3): 64. 1878. T-Protol.: 
Hab. prope Rio de Janeiro, Glaziou 36/0. T-Specim.: | 
of |. Brazil: Guanabara, 21 Apr 1869, 4.F.M. Glaziou 
3610 [IT: K, US-290153}. 

DST.: Arge, Braz. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 
(1987); Anderson, D. E. (1974); Doll, J. C. (1878); 
Filgueiras, T. S. (1991); Filgueiras, T. S. (1999); 
Molina R., A. (1975); Renvoize, S. A. (1984); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

SYN.: Chloris guaranitica Parodi. 


Chloris pallescens Regel, Annot. Bot. Ind. Sem. 28. 1863. 
NOTE: nom. inval., as syn. of Chloris radiata (L.) Sw. 
= Chloris radiata (L.) Sw. 


Chloris paniculata Scribn., Proc. Amer. Acad. Arts 38: 262. 
1902. T-Protol.; Costa Rica: Nuez Isl. (an islet near 
Cocos Isl.), abundant in bunches 3-5 dm in diameter, 
covering considerable areas on cliffs near the coast, 
Snodgrass & Heller 968. T-Specim.: | of 2. Galapagos 
Islands, Jul 1889, RE. Snodgrass & E.G. Heller 968 
[IT: US-80858], Endemic to Cocos Isl., not found on 
Galapagos, mislabeled collection?. 2 of 2. Costa Rica: 
Isla del Coco, R.E. Snodgrass & E.G. Heller 968 [HT: 
GH]. 

DST.: CoRi. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994); Pohl, R. W. (1994). 


Chloris paraguaiensis Steud., Syn. Pl. Glumac. 1: 204. 1854. 
T-Protol.: Paraguay, Rengger s.n. 
= Chloris barbata Sw. 


Chloris parodiana Roseng. & Izag., Bol. Soc. Argent. Bot. 
12: 128, f. 4. 1968. T-Protol.: Paraguay: Paraguari: 
costado de la via ferrea en cruce camino, gregaria, 7 
abril 1950, Rosengurtt B-5979 [HT: MFVA]. 

= Chloris canterae var. canterae. 


Chloris parvispicula Caro & E.A. Sanchez, Kurtziana 6: 
227, f. 3. 1971. T-Protol.: Argentina: La Rioja, 
Castellanos 1940. 

DST.: Arge. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 


Chloris paspaloides Hochst., Flora 38: 206. 1885. T-Protol.: 
Abyssinia [1853], Schimper, Hb. abyss. Buch. 400 
[error for 1400] [HT:STR}. 

= Eustachys paspaloides (Vahl) Lanza & Mattei. 


Chloris paytensis Steud., Syn. Pl. Glumac. 1: 207. 1854. T- 
Protol.: Peru: Payta, Urville s.n. 
= Eustachys distichophylla (Lag.) Nees. 


Chloris pectinata Benth. Fl. Austral. 7: 612. 1878. T- 
Specim.: | of 2. Australia: Northern Territory: 
Charlotte Waters, Giles s.n. [LT: K; ILT: MEL], LT 
designated by ?. 2 of 2. Australia: Queensland: Cook 
Distr: Cashmere, 20 Mar 1875, Armit 188 [ST: MEL]. 

DST.: USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Lazarides, 
M. (1972). 

SYN.: Chloris divaricata var. minor J.M. Black; Chloris 
divaricata var. muelleri Domin; Chloris pectinata 
var. fallax Domin; Chloris pectinata var. typica 
Domin. 


Chloris pectinata var. fallax Domin, Biblioth. Bot. 85: 366. 
1915. T-Specim.: 1 of 2. Australia: Queensland: Burke 
Dist: Hughenden, Feb 1910, Domin s.n. [LT: PR]. 2 of 
2. Australia; Queensland: Mitchell Dist: Winton, Mar 
1910, Domin s.n. [ST: PR]. 
= Chloris pectinata Benth. 


Chloris pectinata var. typica Domin, Biblioth. Bot. 85: 366. 
1915S. 
NOTE: nom. inval., see ICBN Art. 24.3. 
CORRECT NAME: Chloris pectinata var. pectinata. 
= Chloris pectinata Benth. 


Chloris pedicellata Steud., Flora 33: 232. 1850. T-Protol.: 
[Mexico][T: Herb. Delessert], In note following 
Schellingia tenera Steud. In Herb. Delessert. 

NOTE: nom. inval., as syn. of Schellingia tenera Steudel. 
CORRECT NAME: Schellingia tenera Steud. 
= Aegopogon tenellus (DC.) Trin. 


Chloris pedicellata Steud. ex E. Fourn., Mexic. Pl. 2: 71. 
1886. 

NOTE: nom. inval., as syn. of Aegopogon gemniflorus 
Kunth. 

= Aegopogon tenellus (DC.) Trin. 


Chloris pendula Salzm. ex Steud., Syn. Pl. Glumac. 1: 208. 
1854. T-Protol.: Brazil. 


48 Catalogue of New World Grasses: II. 


NOTE: nom. inval., as syn. of Chloris bahiensis Steudel. 
= Eustachys caribaea (Spreng.) Herter. 


Chloris penicillata Jan. ex Trin., Gram. Unifl. Sesquifl. 231. 
1824. 
NOTE: nom. inval., as syn. of Chloris compressa DC. 
E. NAME: Chloris penicillata (Vahl) Pers. 
= Chloris virgata Sw. 


Chloris penicillata Willd. ex Steud., Nomencl. Bot. (ed. 2) 1: 
353. 1840. 
NOTE: nom. inval., as syn. of Chloris elegans Kunth. 
E. NAME: Chloris penicillata (Vahl) Pers. 
= Chloris virgata Sw. 


Chloris petraea Sw., Prodr. 25. 1788. T-Protol.: Jamaica, 
Swartz s.n. [HT: S]. T-Specim.: | of 1. Jamaica, no 
date, 7. Drummond s.n. |T: S; IT: US-80832 (fragm.)]. 

REPLACED NAME: Chloris swartziana Dall. 
= Eustachys petraea (Sw.) Desv. 


Chloris petraea Thunb., Prodr. Pl. Cap. 1: 20. 1794. T- 
Specim.: | of 1. South Africa: Cape, Thunberg 23939 
{HT: UPS]. 
NOTE: hom. illeg. 
E. NAME: Chloris petraea Sw. 
= Eustachys paspaloides (Vahl) Lanza & Mattei. 


Chloris piperita Michx. ex Steud., Nomencl. Bot. (ed. 2) 1: 
353. 1840. 
NOTE: nom. inval., as syn. of Campulosus monostachyus P. 
Beauv. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Chloris pluriflora (E. Fourn.) Kuntze, Revis. Gen. Pl. 2: 771. 
1891. Basionym: Trichloris pluriflora E. Fourn. 
NOTE: error, actually published as Chloropsis pluriflora. 
= Trichloris pluriflora E. Fourn. 


Chloris pluriflora (E. Fourn.) Clayton, Kew Bull. 21: 102. 
1967. Basionym: Trichloris pluriflora E. Fourn. 
= Trichloris pluriflora E. Fourn. 


Chloris poiformis Kunth, Nov. Gen. Sp. 1: 169. 1815 [1816]. 
T-Specim.: | of 1. #L\WH.A. von Humboldt, AJA. 
Bonpland 96 1/2 [T: (fragm., US-2830888)]. 

= Leptochloa virgata (L.) P. Beauv. 


Chloris polydactyla (L.) Sw., Prodr. 26. 1788. Basionym: 
Andropogon polydactylon L. 
NOTE: comb. illeg., basionym is a nom. illeg. superfl., see 
Art. 58.1 Ex. 1. 
CORRECT NAME: Chloris barbata Sw. 
= Chloris elata Desv. 


Chloris polydactyla var. breviaristata Hack. ex Kiitz., Bol. 
Acad. Ci. (Cordoba) 16: 257. 1899. T-Protol.: 
Argentina: Tucuman, cercanias de la ciudad, 6-22 Dec 
1872, Lorentz & Hieronymus 157, Hack in litt. 

DST.: Arge. 
SYN.: Chloris breviaristata (Hack. ex Kitz.) Herter. 
= Status not yet determined. 


Chloris polydactyla subsp. multiradiata (Hochst.) Chiov., 
Annuario Reale Ist. Bot. Roma 8: 54. 1903. Basionym: 
Chloris multiradiata Hochst. 

= Chloris virgata Sw. 


Chloris polydactyla to. pauciradiata Kurtz, Bol. Acad. Ci. 
(Cordoba) 16: 257. 1900. T-Protol.: Argentina: 


"Cordoba: entre Arroyo de la Parra y Los Molles cerca 
de la, Leg. W. Bodenbender [Kurtz 445 7a]. 
= Chloris canterae Arechav. 


Chloris polydactyla fo. stolonifera Parodi, Revista Argent. 
Agron. 20: 24. 1953. T-Protol.: Cult. in Hort. Bot. Fac. 
Agr. et Veter., 1932, L.R. Parodi 10256 [T: cult. Hort. 
Bot. Fac. Agr. et Veter.]. T-Specim.: 1 of 1. Cultivated 
in Argentina at Buenos Aires, Mar 1932, 1.R. Parodi 
10256 (IT: US-1539641]. 
= Chloris canterae Arechav. 


Chloris polystachya Lag., Gen. Sp. Pl. 4. 1816. T-Protol.: 
Habitat in Nueva Hispania unde demina missit D. 
Sessé. 

= Chloris submutica Kunth. 


Chloris pratensis Lesible ex Steud., Syn. Pl. Glumac. 1: 207. 
1854. T-Protol.: Lesible s.n. [T: Hb. Lenormand.]. 
NOTE: nom. inval., as syn. of Chloris subbiflora Steudel. 
= Chloris submutica Kunth. 


Chloris prieurii Kunth, Revis. Gramin. 1: 89. 1829. T- 
Specim.: | of 1. Senegal: Ile St. Louis, Leprieur s.n. 
{HT: P; IT: kK]. 
NOTE: see also Rev. Gram. 2: 441 (1831). 
= Enteropogon prieurii (Kunth) Clayton. 


Chloris procumbens P. Durand, Quib. Chlor. Spec. 16, 22. 
1808. T-Specim.: 1 of 2. Cultivated in Montpelier 
from seed of plants grown in Madrid from seed said by 
Née to have come from the Philippines Islands, P. 
Durand s.n. |HT: MPU-1469 (photocopy, CIIDIR, L); 
IT: US-80856  (fragm.), MPU-1468 (photocopy, 
CIIDIR, L), US-80856 (fragm. ex MPU, & illustr. of 
2nd MPU specimen)], Cultivated at Madrid, but 
probably from Mexico or South America. The same 
Nee material was cultivated in Madrid and named as 
Bouteloua prostrata Lag. 2 of 2. 

= Bouteloua simplex Lag. 


Chloris propinqua Steud., Syn. Pl. Glumac. 1: 204. 1854. T- 
Protol.: West Indies: Guadeloupe, Duchaissing s.n. 
[HT: P(fragm., US)]. T-Specim.: | of 1. Guadeloupe, 
no date, E.P. de F. Duchassaing s.n. |HT: P (fragm., 
US-2830904)]. 
= Chloris ciliata Sw. 


Chloris pubescens Lag., Varied. Ci. 2(4): 143. 1805. T- 
Specim.: | of 1. Peru: ex Peruvia [HT: MA-185242; 
IT: G, US-2830893 (fragm. ex MA & photo)]. 
= Chloris virgata Sw. 


Chloris punctulata Hochst. ex Steud., Syn. Pl. Glumac. 1: 
205. 1854. T-Protol.: Nubia, Kotschy, Hochstetter 
herb. un. it. nubic. 23 (HT: P; IT: K]. T-Specim.: 1 of 
2. Egypt, 20 Sept 1839, 7. Kotschy 33350 [T: L]. 2 of 
2. Sudan: Locis Siccis prope pagum Cordofanum Abu- 
Gerad, 20 Sep 1839, K. G. T. Kotschy 23 {IT: US- 
3243709]. 

NOTE: nom. illeg. superfl. 

CORRECT NAME: Chloris cryptostachya Steud. ex J.A. 
Schmidt. 

= Enteropogon prieurii (Kunth) Clayton. 


Chloris pycnothrix Trin., Gram. Unifl. Sesquifl. 234. 1824. 
T-Protol.: Ins. Santa Catharinae, Chamisso s.n. (HY: 
LE]. 


Catalogue of New World Grasses: II. 49 


DST.: Arge, Boli, Braz, Cari, Colo, Ecua, Hond, Nica, Para, 
Peru, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Brako, L. & 

J. L. Zarucchi (1993); Davidse, G., M. Sousa S. & A. 

QO. Chater (editores generales) (1994); Filgueiras, T. 

S. (1991); Filgueiras, T. S. (1999); Gibbs Russell, G. 
E, W. G. Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987); Jorgensen, P. M. & S. 
Leén-Yanez (eds.) (1999); Lawesson, J. E., H. 
Adsersen & P. Bentley (1987); ORSTOM (1988); 
Phillips, S. (1995); Pohl, R. W. (1994); Porter, D. M. 
(1983); Renvoize, S. A. (1984); Renvoize, S. A. 
(1988); Smith, L. B., D. C. Wasshausen & R. M. 
Klein (1981); Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 

Chloris beyrichiana Kunth; Chloris beyrichiana 
Steud.; Chloris humilis Kunth; Chloris intermedia A. 
Rich.; Chloris leptostachya Hochst. ex A. Rich.; 
Chloris leptostachya var. intermedia (A. Rich.) T. 
Durand & Schinz; Chloris obtusifolia Desv.; Chloris 
radiata var. beyrichiana (Kunth) Hack. ex Stuck.; 
Chloris salzmannii Steud.; Gymnopogon 
beyrichianus (Kunth) Parodi; Gymnopogon haumanii 
Parodi; Gymnopogon radiatus var. beyrichiana 
(Kunth) Parodi. 


Chloris radiata (L.) Sw., Prodr. 26. 1788. Basionym: 
Agrostis radiata L. 
DST.: Beli, Boli, Braz, Cari, Chil, Colo, CoRi, Ecua, ElSa, 
FrGu, Guat, Guya, Hond, Mexi, Nica, Pana, Para, 
Peru, Suri, USA, Vene. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Anderson, D. 
E. (1974); Balick, M. J.. M. H. Nee & D. E. Atha 
(2000); Beetle, A. A. (1977); Beetle, A. A. (1987); 
Berendsohn, W. G. & A. E. Araniva de Gonzalez 
(1991); Brako, L. & J. L. Zarucchi (1993); Catasus 
Guerra, L. (1985); Catasus Guerra, L. (1997); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Dodson, C. H., A. H. Gentry & PF. 
M. Valverde (1985); DGll, J. C. (1878); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Foster, R. C. (1958); Gould, F. W (1979); Hitchcock, 
A. S. (1909); Hitchcock, A. S. (1927); Hitchcock, A. 
S. (1936); Hitchcock, A. S. (1951); Jorgensen, P. M. 
& S. Leén-Yanez (eds.) (1999); Judziewicz, E. J. 
(1990); Lawesson, J. E., H. Adsersen & P. Bentley 
(1987); Lindeman, J. C. & A. L. Stoffers (1963); 
Liogier, H. & L. Martorell (1982); Long, R. W. & O. 
Lakela (1971); Macbride, J. F. (1936); Marticorena, 
C. & M. Quezada (1985); McClure, F. A. (1955); 
McVaugh, R. (1983); Molina R., A. (1975); Pohl, R. 
W. (1980); Pohl, R. W. (1994); Porter, D. M. (1983); 
Reeder, J. R. & C. G. Reeder (1971); Renner, S. S., 
H. Balslev & L. B. Holm-Nielsen (1990); Renvoize, 
S. A. (1998); Tovar, O. (1993); Wiggins, I. L. & D. 
M. Porter (1971): Wunderlin, R. P. (1998). 

Agrostis radiata L., Chloris biflora Regel; Chloris 
crinita Salzm. ex D6OIl; Chloris durandiana Schult.; 
Chloris glaucescens Steud.; Chloris gracilis P. 
Durand; Chloris pallescens Regel; Chloris radiata 
var. /ongiaristata Christa Miiller; Chloris scoparius 
(Lam.) Desf.; Chloris scoparius Regel; Chloris 


SYN.: 


SYN.: 


tacnensis Steud. ex Lechler; Cynosurus scoparius 
Lam.; Gymnopogon radiatus (L.) Parodi. 


Chloris radiata var. beyrichiana (Kunth) Hack. ex Stuck., 
Anales Mus. Nac. Hist. Nat. Buenos Aires 6: 489. 
1906. Basionym: Chloris beyrichiana Kunth. 

NOTE: Index of Grass Species collat. 13: 489. 
= Chloris pycnothrix Trin. 


Chloris radiata var. longiaristata Christa Miiller, Feddes 
Repert. 94(9-10): 625, f. 2. 1983. T-Protol.: Ecuador: 
Galapagos-Inseln: Santiago, Fagerlind & Wibom 3434 
{HT: S]. 

= Chloris radiata (L.) Sw. 


Chloris repens Steud., Nomencl. Bot. (ed. 2) 1: 353. 1840. 
NOTE: nom. inval., as syn. of Eleusine indica (L.) Gaertner. 
= Eleusine indica (L.) Gaertn. 


Chloris repens Hochst., Flora 38: 204. 1855. T-Protol.: 
Ethiopia: Semien, Schimper, in hb. Buchinger 1110 
[HT:STR; IT:P]. 

= Chloris gayana Kunth. 


Chloris retusa Lag., Gen. Sp. Pl. 5. 1816. T-Protol.: Habitat 
in Bonariensi regno. T-Specim.: | of 2. Uruguay: del 
Cerro y Pefia Blanca, L. Nee s.n. [HT: MA; IT:BAA, 
MO, US-80834 (fragm. & photo)], The locality of the 
type has been confused. See Molina, Candolea 51: 246 
(1996) for a discussion. 2 of 2. Argentina: "Hab. in 
Bonariensi Regno.", no date, Nee s.n. [T: MA, F- 
029381]. 

= Eustachys retusa (Lag.) Kunth. 


Chloris rogeoni A. Chev., Rev. Int. Bot. Appl. Agric. Trop. 
14(150): 127. 1934. T-Specim.: | of 3. Dire,Chevalier 
43871 (ST: P]. 2 of 3. El Oualadjii, Chevalier 43937 
[ST: P]. 3 of 3. El Oualadjii, Chevalier 42357 (ST: P]. 

NOTE: as Chloris "Rogeoni"; nom. inval., without Latin 
description. 
= Chloris virgata Sw. 


Chloris rufescens Lag., Varied. Ci. 2(4): 143. 1805. T- 
Specim.: | of 1, Cultivated in the Montpellier 
Botanical Garden from material collected in Mexico 
[erroneously attributed to the Philippines], L. Née s.n. 
[NT: MA-252909; INT: US-80836 (fragm. ex MA & 
drawing by Parodi)], NT designated by Veldkamp, 
Taxon 50: 846. (2001); wrongly attributed to the 
Philippines by Lagasca according to Chase & Niles. 

DST.: CoRi, Guat, Hond, Mexi. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Espejo Serna, A., 
A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
McClure, F. A. (1955); McVaugh, R. (1983); Pohl, 
R. W. (1994); Reyes-Garcia, A. & M. Sousa S. 
(1997); Veldkamp, J. F. (2001). 

Agrostomia aristata Cerv.; Chloris aristata (Cerv.) 
Swallen. 


SYN.: 


Chloris rufescens Steud., Syn. Pl. Glumac. 1: 206. 1854. T- 
Protol.: Ins. Maurit., Urville s.n. 
NOTE: hom. illeg. 
E. NAME: Chloris rufescens Lag. 
= Chloris barbata Sw. 


Chloris rupestris (Ridl.) Hitche., Man. Grasses W. Ind. 126. 
1936. Basionym: Gymnopogon rupestre Ridl. 


50 Catalogue of New World Grasses: II. 


= Enteropogon mollis (Nees) Clayton. 


Chloris sagraeana A. Rich., Hist. Fis. Cuba, Bot. 11: 315. 
1850. ‘T-Protol.: [West Indies]: Cuba: "Crescit in 
ruderatis et locis arenosis circa Habanam.", Ramon de 
la Sagra s.n. [HT: P T-fragm.: US]. T-Specim.: 1 of 1. 
Cuba: La Habana, no date, R. de la Sagra A. 
RICHARD 2 |HT: P (fragm., US-80837)]. 

DST.: Cari. 

LVL. ACCPTC:: |. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Anderson, D. 
E. (1974); Anonymous (1986); Catasus Guerra, L. 
(1985); Catasus Guerra, L. (1997); Gould, F. W 
(1979); Hitchcock, A. S. (1936); Liogier, H. & L. 
Martorell (1982). 

SYN.: Chloris eleusinoides Griseb.; Chloris morales-coelloi 
Leon ex Britton. 


Chloris sagraeana subsp. cubensis (Hitchc. & Ekman) Cat. 
Guerra, Fontqueria 44: 145. 1996. Basionym: Chloris 
cubensis Hitche. & Ekman. 

DST.: Cari. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Catasus Guerra, L. (1985); Catasus 

Guerra, L. (1997), 

Chloris cubensis Hitche. & Ekman; Chloris 
sagraeana UNRANKED cubensis (Hitche. & 

Ekman) Cat. Guerra. 


SYN.: 


Chloris sagraeana UNRANKED cubensis (Hitche. & 
Ekman) Cat. Guerra, Acta Bot. Cub. 25: 6. 1985. 
Basionym: Chloris cubensis Hitchc. & Ekman. 

NOTE: nom. inval., rank not indicated. 
= Chloris sagraeana subsp. cubensis (Hitchc. & Ekman) 
Cat. Guerra. 


Chloris sagraeana A. Rich. subsp. sagraeana. 
LVL. ACCPTC:: 1. 


Chloris salzmannii Steud., Syn. Pl. Glumac. 1: 206. 1854. T- 
Protol.: Bahia, "C. crinita Salzm.". T-Specim.: | of 1. 
Brazil: Bahia: in argillosis aridis, P. Salzmann 664 
[HT: G; IT: K, US-1614946]. 

= Chloris pyecnothrix Trin. 


Chloris sclerantha Lindl., J. Exped. Trop. Australia 31. 1848. 
T-Specim.: | of 1. Australia: New South Wales: banks 
of Bogan River, 1846, Mitchell 116 (HT: CGE; IT: K]. 

= Chloris ventricosa R. Br. 


Chloris scoparius (Lam.) Desf., Tabl. Ecole Bot. 14. 1804. 
Basionym: Cynosurus scoparius Lam. 
NOTE: cites Cynosurus scoparius Link [error for Lam.?]. 
= Chloris radiata (L.) Sw. 


Chloris scoparius Regel, Annot. Bot. Ind. Sem. 28. 1863. 
NOTE: nom. inval., as syn. of Chloris radiata (L.) Sw. 
E. NAME: Chloris scoparius (Lam.) Dest. 
= Chloris radiata (L.) Sw. 


Chloris secundus (Pursh) Eaton, Man. Bot. (ed. 5) 173. 1829. 
Basionym: Cynosurus secundus Pursh. 
= Bouteloua curtipendula (Michx.) Torr. 


Chloris septentrionalis Mill. Hal., Bot. Zeitung (Berlin) 
19(46): 340. 1861. T-Protol.: USA: America 
septentrionalis, ubi ad Rio Brazos legit, Drummond 
sn. T-Specim.: | of 1. USA: Texas, no date, 7. 
Drummond s.n. (IT: MO-1837394, US-1063206 
(fragm.)]. 


= Eustachys petraea (Sw.) Desv. 


Chloris sesquiflora Burkart, Bol. Soc. Argent. Bot. 12: 287, 
f, 2, 1968. T-Protol.: Argentina: Entre Rios: Depto. 
Concepcién del Uruguay, in pratis subsalsis flum. 
Gualegguaychu, vencindades del Parque Unzue, 19 
Apr 1965, Burkart 25676 [HT: SI; IT: HSC]. T- 
Specim.: | of 1. Argentina: Entre Rios: Dept. 
Concepcién del Uruguay: bajos cerca del rio 
Gualeguaychu, vencdendades del Parque Unzué, 19 
Apr 1965, 4.E. Burkart et al. 25676 [HT: SI; IT; US- 
2959126]. 

DST.: Arge, Para. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Burkart, A. 
(1969); Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 


Chloris subbiflora Steud., Syn. Pl. Glumac. |: 206. 1854. T- 
Protol.: [USA]: America Septentrionalis, Lesible s.n. 
[T: CN T: US]. T-Specim.: | of 1. "North America", 
1823, H. Lesible s.n. [HT: P (fragm., US-2732913)]. 
= Chloris submutica Kunth. 


Chloris x subdolichostachya Mill. Hal. Bot. Zeitung 
(Berlin) 19(46): 341. 1861. T-Protol.: America 
septentrionalis, ubi forsan in Texas, 7. Drummond 
372. T-Specim.: 1 of 1. USA: Texas, no date, 7: 
Drummond 372 [HT: ?; IT: US-879091 (fragm.)]. 

NOTE: determined to be a hybrid by Anderson in Gould, 

Grasses of Texas, 1975, p. 327. 

DST.: Mexi, USA. 

LVL. ACCPTC:: L. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Beetle, A. A. (1987); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000): 
Hitchcock, A. S. (1951); Thomas, R. D. & C. M. 
Allen (1993). 

Chloris brevispica Nash; Chloris latisgquamea Nash; 
Chloris verticillata var. aristulata Torr. & A. Gray: 
Chloris verticillata var. intermedia Vasey. 


SYN.: 


Chloris submutica Kunth, Nov. Gen. Sp. 1: 167, t. 50. 1815 
[1816]. T-Protol.: Crescit in temperatis convallis 
Mexicanae propter littora lacus Tezcucensis, alt. 1150 
hexap, Humboldt & Bonpland s.n. T-Specim.: | of 1. 
Mexico: Tezcucensi, Feb, /.WH.A. von Humboldt, 
A.J.A. Bonpland 4182 (HT: P (fragm., US-2830903)]. 

DST.: Colo, Ecua, Guat, Mexi, USA, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Anonymous 
(1986); Beetle, A. A. (1977); Beetle, A. A. (1987); 
Breedlove, D. E. (1986); Davidse, G., M. Sousa S. & 
A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lépez-Ferrari, and J. Valdes-Reyna 
(2000); Hitchcock, A. S. (1951); Jorgensen, P.M. & 
S. Leén-Yanez (eds.) (1999); McVaugh, R. (1983); 
Pohl, R. W. (1994). 

Agrostomia mutica Cerv., Chloris inermis Trin.; 
Chloris polystachya Lag.; Chloris polystachya Lag.; 
Chloris pratensis Lesible ex  Steud.; Chloris 
subbiflora Steud.; Eustachys polystachya_ (Lag.) 
Kunth; Eustachys submutica (Kunth) Roem. & 
Schult.; Phacellaria panicea Willd. ex Steud. 


SYN.: 


Chloris subtriflora Steud., Syn. Pl. Glumac. |: 208. 1854. T- 
Protol.: Senegambia, LaPrieur s.n. 


Catalogue of New World Grasses: II. 5] 


= Enteropogon prieurii (Kunth) Clayton. 


Chloris suringarii Hitchc., Symb. Antill. 7: 167. 1912. T- 
Protol.: West Indies: Curacao ad Brakkeput: 14 Jan 
1885, WFR Suringar (HT: Krug and Urban 
Herb.(fragm., US)]. T-Specim.: 1 of 1. Netherlands 
Antilles, 14 Jan 1885, WFR. Suringar s.n. [HT: Krug 
and Urban Herb.; IT: L, US-690358 (fragm.)]. 

DST.: Cari. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Catasus 
Guerra, L. (1985); Catasus Guerra, L. (1997); 
Lindeman, J.C. & A. L. Stoffers (1963). 


Chloris swartziana DOll, Fl. Bras. 2(3): 68-69. 1878. 
NOTE: nom. illeg. superfl., based on Chloris petraea Sw., 
non Thunb. 
CORRECT NAME: Chloris petraea Sw. 
= Eustachys petraea (Sw.) Desv. 


Chloris swartzii Mill. Hal., Bot. Zeitung (Berlin) 19(46): 
341. 1861. 
NOTE: nom. illeg. superfl.; based on Chloris petraea Sw., 
non Thunb. unpbulished. 
CORRECT NAME: Chloris petraea Sw. 
= Eustachys petraea (Sw.) Desv. 


Chloris tacnensis Steud. ex Lechler, Berberid. Amer. Austr. 
56. 1857. T-Protol.: Peru, Lechler 1569. 
NOTE: nom. nud. 
= Chloris radiata (L.) Sw. 


Chloris tenuis Poir., Encycl., Suppl. 5: 614. 1817 [Nov]. 
NOTE: nom. superfl.; based on Chloris filiformis Poir. 
Encycl. Suppl. 2: 237, n. 8. 1811. Not P. 238. 

= Bouteloua simplex Lag. 


Chloris tenuispica Nash, Bull. Torrey Bot. Club 25: 436. 
1898, T-Protol.: USA: Texas: 1889,G.C. Nealley s.n. 
T-Specim.: | of 1. USA: Texas, 1889, G.C. Nealley 
s.n. [LT: US-80860], LT designated by ?. 

= Chloris andropogonoides E. Fourn. 


Chloris tetrastachys Hack. ex Hook. f., Fl. Brit. India 7: 291. 
1896, 
NOTE: nom. inval., as syn. of C. virgata Swartz. 
= Chloris virgata Sw. 


Chloris texana (Vasey) Nash, Bull. Torrey Bot. Club 25: 441. 


1898. Basionym: Chloris ciliata var.texana Vasey. 
= Chloris ciliata Sw. 


Chloris texensis Nash, Bull. Torrey Bot. Club 23: 151. 1896. 


T-Protol.: USA: Texas, Thurow & G.C. Nealley s.n. T- 
Specim.: | of 1. USA: Texas, 188-, G.C. Nealley s.n. 
[ST: US-80852]. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Correll, D. 

S. & M. C. Johnston (1970); Hitchcock, A. S. (1951). 
SYN.: Chloris nealleyi Nash. 


Chloris tibestica Quézel, Bull. Soc. Hist. Nat. Afrique N. 48: 
84. 1957. T-Protol.: Emi Koussi. 
= Chloris virgata Sw. 


Chloris trichodes Lag. ex Parodi, Revista Argent. Agron. 14: 


62. 1947. 
NOTE: nom. inval., as syn. of 7richloris crinita (Lag.) L. 
Parodi. 


= Trichloris crinita (Lag.) Parodi. 


Chloris truncata R. Br., Prodr. 186. 1810. T-Specim.: | of 1. 
Australia: New South Wales: Port Jackson, Brown 
6243 [HT: BM; IT: E, K]. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Edgar, E., 
H. E. Connor & J. E. Shand (1991); Gibbs Russell, 
G. E, W. G. Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987); Hitchcock, A. S. (1951); 
Lazarides, M. (1972). 

SYN.: Chloris elongata Poir.; Chloris megastachya Schrad. 
ex Schult.; Chloris truncata fo. abbreviata Thell. 


Chloris truncata fo. abbreviata Thell., Vierteljahrsschr. 
Naturf. Ges. Zurich 64: 706. 1919. T-Specim.: | of 1. 
Switzerland: Derendingen, Solothurn, 27 Sep 1917, 
Probst s.n. (HT: Z; IT: K]. 

= Chloris truncata R. Br. 


Chloris uliginosa Hack., Repert. Spec. Nov. Regni Veg. 7: 
320. 1909. T-Protol.: Uruguay: in locis uliginosis 
prope Cuarain, 1902, Berro s.n. [HT: W?}. T-Specim.: 
1 of 1. Uruguay: Cuareim, 23 Jan 1902,4.B. Berro 
2678 [T: BAA, US-80861 (fragm.)]. 

= Eustachys ulignosa (Hack.) Herter. 


Chloris ventricosa R. Br., Prodr. 186. 1810. T-Specim.: | of 
1. Australia; New South Wales: Richmond, Brown 
6242 [HT: BM; IT: K, E]. 
DST.: USA. 
LVL. ACCPTC.:: 1. 
ALSO ACCPTD. BY: Anderson, D. E. (1974); Hitchcock, 
A. S. (1951); Lazarides, M. (1972). 
SYN.: Chloris sclerantha Lindl.; Chloris ventricosa var. 
tenuis Benth. 


Chloris ventricosa var. tenuis Benth., Fl. Austral. 7: 613. 
1878. T-Specim.: | of 2. Australia: Queensland: Port 
Curtis Dist: Rockhampton, 15 Mar 1876, O'Shanesy 
1679 [LT: K; ILT: MEL]. 2 of 2. Australia: 
Queensland: Neerkool Creek, near Rockhampton, 
Bowman 22 |ST: MEL]. 

= Chloris ventricosa R. Br. 


Chloris verticillata Nutt., Trans. Amer. Philos. Soc., n.s., 5: 
150. 1835. T-Protol.: Arkansas Territory: Hab. on the 
sandy banks of the Arkansas, near Fort Smith, rare, 
Nuttall s.n. T-Specim.: | of 1. USA: Arkansas: near 
Fort Smith, no date, 7. Nuttall s.n. [HT: PH(fragm.., 
US-2830886): IT: K]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Beetle, A. 
A. (1977); Beetle, A. A. (1987); Correll, D. S. & M. 
C. Johnston (1970); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Fernald, M. 
(1950); Gleason, H. A. & A. Cronquist (1968): 
Gleason, H. A. & A. Cronquist (1991); Great Plains 
Flora Association (1986); Hickman, J. C. (1993); 
Hitchcock, A. S. (1951); Munz, P. A. & D. D. Keck 
(1959); Voss, E. G. (1972). 


Chloris verticillata var. aristulata Torr. & A. Gray, Pacif. 
Railr. Rep. 2: 176. 1855. T-Protol.: [USA]: "Type 
collected on the lower Rio Grande.", Dr. Gregg s.n. 


52 Catalogue of New World Grasses: II. 


= Chloris x subdolichostachya Mill. Hal. 


Chloris verticillata var. intermedia Vasey, Contr. U.S. Natl. 
Herb. 2(3); 528. 1894. T-Protol.: USA: Texas: 
Houston: sandy land, central Texas to Oklahoma, Hall 
773. T-Specim.: 1 of 1. USA: Texas: Harris Co., 20 
Apr 1872, E. Hall 773 [TYPE: US-879085]. 

CORRECT NAME: Chloris latisquamea Nash. 
= Chloris x subdolichostachya Mill. Hal. 


Chloris virgata Sw., Fl. Ind. Occid. 1: 203. 1797. T-Protol.: 
Provenit in Antigua, Swartz s.n. T-Specim.: | of 1. 
Antigua, Swartz s.n. [|HT: S; IT: BM]. 

DST.: Arge, Boli, Braz, Cari, Chil, Colo, Ecua, ElSa, Guat, 

Mexi, Nica, Pana, Para, Peru, USA, Vene. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Abrams, L. (1923); Anderson, D. E. 
(1974): Beetle, A. A. (1977); Beetle, A. A. (1987); 
Berendsohn, W. G. & A. E. Araniva de Gonzalez 
(1991): Bor, N. L. (1960); Brako, L. & J. L. Zarucchi 
(1993): Breedlove, D. E. (1986); Burkart, A. (1969); 
Cabrera, A. L. (1970); Catasus Guerra, L. (1985); 
Catasus Guerra, L. (1997): Correll, D. S. & M. C. 
Johnston (1970); Cronquist, A., A. H. Holmgren, N. 
H. Holmgren & J. L. Reveal (1977); Davidse, G., M. 
Sousa S. & A. O. Chater (editores generales) (1994); 
Doll, J. C. (1878); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Fernald, M. 
(1950); Filgueiras, T. S. (1999); Foster, R. C. (1958); 
Gibbs Russell, G. E, W. G. Welman, E. Reitief, K. L. 
Immelman, G. Germishuizen, B. J. Pienaar, M. van 
Wyk, and A. Nicholas (PRECIS) (1987); Gleason, H. 
A. & A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Gould, F. W (1979); Gould, F. W. 
& R. Moran (1981); Great Plains Flora Association 
(1986); Hickman, J. C. (1993); Hitchcock, A. S. 
(1927): Hitchcock, A. S. (1936); Hitchcock, A. S. 
(1951); Jorgensen, P. M. & S. Leén-Yanez (eds.) 
(1999); Killeen, T. J. (1990); Lawesson, J. E., H. 
Adsersen & P. Bentley (1987); Lazarides, M. (1972); 
Lindeman, J. C. & A. L. Stoffers (1963); 
Marticorena, C. & M. Quezada (1985); McClure, F. 
A. (1955); McVaugh, R. (1983); Molina R., A. 
(1975); Munz, P. A. & D. D. Keck (1959); Munz, P. 
A. (1974); Phillips, S. (1995); Pohl, R. W. (1994); 
Porter, D. M. (1983); Radford, A. E., H. E. Ahles & 
C. R. Bell (1968); Reeder, J. R. & C. G. Reeder 
(1971); Renvoize, S. A. (1984); Renvoize, S. A. 
(1998); Reyes-Garcia, A. & M. Sousa S. (1997): 
Shreve, F. & I. L. Wiggins (1964); Sousa S., M. & E. 
F. Cabrera C. (1983); Tovar, O. (1993); Veldkamp, J. 
F. (2001); Wiggins, |. L. & D. M. Porter (1971); 
Wunderlin, R. P. (1998); Zuloaga, F. O., E. G. 
Nicora, Z. E. Ragolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

Agrostomia barbata Cerv., Chloris alba J. Presl; 
Chloris alba var. aristulata Torr.; Chloris albertii 
Regel; Chloris barbata var. decora (Nees ex Steud.) 
Benth.; Chloris brachystachys Andersson; Chloris 
caudata Trin. ex Bunge, Chloris compressa DC.; 
Chloris decora Nees ex Steud.; Chloris elegans 
Kunth; Chloris — gabrielae =Domin; — Chloris 
madagascariensis Steud.; Chloris meccana Hochst. 
ex Steud.; Chloris multiradiata WHochst.; Chloris 
notocoma Hochst.; Chloris penicillata Jan. ex Trin.; 
Chloris penicillata Willd. ex Steud.; Chloris 
polydactyla subsp. multiradiata (Hochst.) Chiov.; 


SYN:: 


Chloris pubescens Lag.; Chloris rogeoni A. Chev.; 
Chloris tetrastachys Hack. ex Hook. f.; Chloris 
tibestica Quézel; Chloris virgata var. elegans 
(Kunth) Stapf; Lepeocercis serrata (Retz.) Trin.: 
Rabdochloa virgata (Sw.) P. Beauv.; Rhabdochloa 
virgata (Sw.) P. Beauv. 


Chloris virgata var. elegans (Kunth) Stapf, Fl. Cap. 7: 642. 
1900. Basionym: Chloris elegans Kunth. 
= Chloris virgata Sw. 


Chloroides Fisch. ex Regel, Index Sem. (St. Petersburg) 28. 
1863. 
NOTE: in syn. sub E. petraea. 
= Eustachys Desv. 


Chloropsis Hack. ex Kuntze, Revis. Gen. Pl. 2: 771. 1891, 
NOTE: nom. superfl. pro Trichloris. 
= Trichloris E. Fourn. ex Benth. 


Chloropsis blanchardiana Gay ex Scribn., Bull. Torrey Bot. 
Club 9: 146. 1882. 
NOTE: as "Chloridopsis"; nom. inval., as syn. of Trichloris 
blanchardianum Fourn. 
= Trichloris crinita (Lag.) Parodi. 


Chloropsis blanchardiana (E. Fourn. ex Scribn.) Kuntze, 
Revis. Gen. Pl. 2: 771. 1891. Basionym: Trichloris 
blanchardiana E. Fourn. ex Scribn. 

= Trichloris crinita (Lag.) Parodi. 


Chloropsis crinita (Lag.) Kuntze, Revis. Gen. Pl. 2: 771. 
1891. Basionym: Chloris crinita Lag. 
= Trichloris crinita (Lag.) Parodi. 


Chloropsis mendocina (Phil.) Kuntze, Revis. Gen. Pl. 3(2): 
348. 1898. Basionym: Chloris mendocina Phil. 
= Trichloris crinita (Lag.) Parodi. 


Chloropsis pluriflora (E. Fourn.) Kuntze, Revis. Gen. Pl. 2: 
771. 1891. Basionym: Trichloris pluriflora E. Fourn. 
= Trichloris pluriflora E. Fourn. 


Chondrosium Desv., J. Bot. Agric. 1: 68. 1813. 
NOTE: orth. var. 
= Bouteloua Lag. 


Chondrosium eriopodum Yorr., Not. Milit. Reconn. 154. 

1848. T-Protol.: USA: [New Mexico]: Del Norte and 
Gila rivers, /Bigelow] s.n. T-Specim.: | of 1. USA: 
New Mexico, 15 1846, /.' Bigelow et al. s.n. [HT: 
NY; IT: US-A865571 (fragm. ex NY)]. 

NOTE: orth, var. as Chondrosium. 

CORRECT NAME: Chondrosum eriopodum Torr. 

= Bouteloua eriopoda (Torr.) Torr. 


Chondrosium gracile Kunth, Nov. Gen. Sp. 1: 176. 1816. 
NOTE: orth. var. 
CORRECT NAME: Chondrosum gracile Kunth. 
= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Chondrosium hirsutum (Lag.) Sweet, Hort. Brit. 455. 1827. 
Basionym: Bouteloua hirsuta Lag. 

NOTE: orth. var. "Chondrosum"; Index of Grass Species, 
Man US. collat. 455, date 1826, presumably based 
on Actinochloa hirsuta Roem. & Schult. 

= Bouteloua hirsuta Lag. 


Chondrosium hirtum Kunth, Nov. Gen. Sp. 1: 176. 1815 
[1816]. T-Protol.: Mexico: Novae Hispaniae, inter 
Mexico et Gasave, 1800 hex., Humboldt & Bonpland 


Catalogue of New World Grasses: II. 53 


s.n. T-Specim.: | of 1. Mexico, no date, Anonymous 
s.n. [HT: B-W ; IT: US-A865681 (fragm. ex B-W)]. 
NOTE: orth. var. 
CORRECT NAME: Chondrosum hirtum Kunth. 
= Bouteloua hirsuta var. hirsuta. 


Chondrosium humile Kunth, Nov. Gen. Sp. 1: 175, t. 56. 
1815 [1816]. T-Specim.: | of 1. Ecuador: Llactacunga 
Quitensium, monte Guadalupens, Anonymous s.n. 
[HT: P; IT: US-A865683 (fragm. ex P-Bonpl.)]. 

NOTE: orth. var. 

CORRECT NAME: Chondrosum humile P. Beauv. ex 
Kunth. 

= Bouteloua simplex Lag. 


Chondrosium karwinskii E. Fourn., Mexic. Pl. 2: 137. 1886. 
T-Protol.: Mexico: Canon de las Minas et Victoria, 
Karwinskii 1479. 

NOTE: orth. var. Chondrosum. 
CORRECT NAME: Chondrosum karwinskii E. Fourn. 
= Bouteloua karwinskii(E. Fourn.) Griffiths. 


Chondrosium parryi E. Fourn., Mexic. Pl. 2: 150. 1886. T- 
Specim.: | of 1. Mexico: San Luis Potosi, 1878,C.C. 
Parry & E. Palmer 943-1/2 |IT: US-81783, US- 
883835], Publ. cites coll. no. 923-1/2 erroneously. 

NOTE: orth. var. 
CORRECT NAME: Chondrosum parryi E. Fourn. 
= Bouteloua parryi (E. Fourn.) Griffiths. 


Chondrosium polystachyum Benth., Bot. Voy. Sulphur 56. 
1844. T-Specim.: | of 1. Mexico: Baja California Sur: 
Magdalena Bay, Barclay s.n. [HT: K; IT: BM, US- 
A865566 (fragm. ex K)]. 

NOTE: “orth. var." "Chondrosum". 
CORRECT NAME: Chondrosum polystachyum Benth. 
= Bouteloua barbata Lag. 


Chondrosium scorpoides (Lag.) Kunth, Revis. Gramin. 1: 94. 
1829. Basionym: Bouteloua scorpioides Lag. 
NOTE: "orth. var." Chondrosum", 
CORRECT NAME: Chondrosum scorpioides (Lag.) Kunth. 
= Bouteloua scorpioides Lag. 


Chondrosium simplex (Lag.) Kunth, Revis. Gramin. 1: 94. 
1829. Basionym: Bouteloua simplexLag. 
NOTE: "orth. var." "Chondrosum". 
CORRECT NAME: Chondrosum simplex (Lag.) Kunth. 
= Bouteloua simplex Lag. 


Chondrosium  subscorpioides Mill. Hal., Bot. Zeitung 
(Berlin) 14: 347. 1856. T-Specim.: 1 of 1. USA: 
California and Texas, 18-, G.W. Barclay s.n. [HT: ?; 
IT: K, US-8883457 (fragm.)], CS\Baja california. 

NOTE: "orth. var." "Chondrosum". 
CORRECT NAME: Chondrosum subscorpioides Mill. Hal. 
= Bouteloua barbata Lag. 


Chondrosium tenue P. Beauv. ex Kunth, Nov. Gen. Sp. 1: 
176. 1816. 
NOTE: orth. var. Chondrosum. 
CORRECT NAME: Chondrosum tenue P. Beauv. ex Kunth. 
= Bouteloua simplex Lag. 


Chondrosum Desv., Nouv. Bull. Sci. Soc. Philom. Paris 2: 
188. 1810. TYPE: Chondrosum procumbens (P. 
Durand) Desv. 

NOTE: as "Chondrosum". 
= Bouteloua Lag. 


Chondrosum aschenbornianum Nees, Linnaea 19(6): 692. 
1847. T-Protol.: Mexico, Aschenb. exs. 331. T- 
Specim.: | of 2. Mexico: A. Aschenborn 331 [HT: B; 
IT: US-865680 (fragm.)], CS. 2 of 2. 

NOTE: as "Chondrosium" aschenbornianum. 
= Bouteloua hirsuta Lag. 


Chondrosum barbatum (Lag.) Clayton, Kew Bull. 37: 417. 
1982. Basionym: Bouteloua barbata Lag. 
= Bouteloua barbata Lag. 


Chondrosum brevisetum (Wasey) Clayton, Kew Bull. 37: 418. 
1982. Basionym: Bouteloua breviseta Vasey. 
= Bouteloua breviseta Vasey. 


Chondrosum chasei (Swallen) Clayton, Kew Bull. 37: 418. 
1982. Basionym: Bouteloua chasei Swallen. 
= Bouteloua chasei Swallen. 


Chondrosum drummondii E. Fourn., Mexic. Pl. 2: 137. 1886. 
T-Specim.: | of 2. USA: Texas, Drummond 323 [ST: 
P; IST: LE, US-865677 (fragm.), US-2182917 
(fragm.), US-1063203 (fragm. ex E)], CS. 2 of 2. 
Mexico: La Miquiguana, Karwinski s.n. [ST]. 
NOTE: as "Chondrosium" drummondii. 
= Bouteloua hirsuta Lag. 


Chondrosum elatum (Reeder & C. Reeder) Clayton, Kew 
Bull. 37: 418. 1982. Basionym: Bouteloua elata 
Reeder & C. Reeder. 

= Bouteloua elata Reeder & C. Reeder. 


Chondrosum eriopodum Tort., Not. Milit. Reconn. 154. 
1848. T-Specim.: | of 1. USA: New Mexico, Las 
Cruces, Wooton 458 [NT: US-A865571 (fragm.)], NT 
designated by Gould, Ann. Missouri Bot. Gard. 
66:397.1979. 

NOTE: as "Chondrosium" eriopodum. 

= Bouteloua eriopoda (Torr.) Torr. 


Chondrosum eriostachyum (Swallen) Clayton, Kew Bull. 37: 
418. 1982. Basionym: Bouteloua eriopoda_ var. 
eriostachya Swallen. 

= Bouteloua eriostachya (Swallen) Reeder. 


Chondrosum exile E. Fourn., Mexic. Pl. 2: 137. 1886. T- 
Protol.: Mexico, Berlandier 842. T-Specim.: | of 1. 
Mexico, JL. Berlandier 842 [HT: P; IT: US-865678 
(fragm. ex P), W]. 

NOTE: as "Chondrosium". 
= Bouteloua barbata Lag. 


Chondrosum foeneum Torr., Not. Milit. Reconn. 153, t. 12. 
1848. T-Protol.: USA: [New Mexico]: uplands 
bordering the valley of the Del Norte, /Emory exped.] 
s.n. T-Specim.: 1 of 3. USA: Rio Cabeza, Sep 1850, 
G.Thurber 27 [T: LL: NY-73937]. 2 of 3. USA, 
Collector unspecified on label [T: 1: NY-73938]. 3 of 
3. USA, 9 Oct 1846, Major W. H. Emory [T: 1: NY- 
73939]. 

NOTE: as "Chondrosium" foenum. 
= Bouteloua hirsuta Lag. 


Chondrosum gracile Kunth, Nov. Gen. Sp. 1: 176, t. 58. 1815 
[1816]. T-Protol.: Crescit in crepidinibus et devexis 
montis porphyritici) La Buffa de Guanaxuato 
Mexicanorum, alt. 1270 hexap., floret Septembri, 
Humboldt & Bonpland s.n. Y-Specim.: | of 1. Mexico: 
Guanajuato: La Bufa, Humboldt & Bonpland s.n. (HT: 
B-W-1628; IT: P, US-A865679 (fragm.)]. 


54 Catalogue of New World Grasses: II. 


NOTE: as "Chondrosium" gracile. 
= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Chondrosum gracile var. polystachyum Nees, Linnaea 19(6): 
692. 1847. T-Protol.: Mexico, Aschenborn exs. 153. T- 
Specim.: | of 1. Mexico, no date, A. Aschenborn 153 
[HT: ?: IT: US-883615 (fragm.)]. 

NOTE: as "Chondrosium" gracile var. polystachyum. 
= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Chondrosum hirsutum (Lag.) Sweet, Hort. Brit. 1: 455. 1826. 
Basionym: Bouteloua hirsuta Lag. 
NOTE: as "Chondrosium". 
= Bouteloua hirsuta Lag. 


Chondrosum hirsutum var. glandulosum (Cerv.) R.W. Pohl, 
Novon 2(2): 102. 1992. Basionym: Erucaria 
glandulosa Cerv. 

= Bouteloua hirsuta var. glandulosa (Cerv.) Gould. 


Chondrosum hirsutum Gould var. hirsutum. 
= Bouteloua hirsuta var. hirsuta. 


Chondrosum hirtum Kunth, Nov. Gen. Sp. 1: 176, t. 59. 1815 
[1816]. T-Protol.: Crescit in temperatis, apricis Novae 
Hispaniae, inter Mexico et Gasave, alt. 1180 hex., 
floret Majo, Humboldt & Bonpland s.n. T-Specim.: 1 
of |. Mexico: inter Mexico et Gasave, Humboldt & 
Bonpland s.n. [HT: P; TT: B-W-1626, M, LE-TRIN- 
2126.1, US-A865681 (fragm.)]. 

NOTE: as "Chondrosium" hirtum. 
= Bouteloua hirsuta Lag. 


Chondrosum humboldtianum Kunth, Revis. Gramin. 1: 93. 
1829, 
NOTE: nom. _ illeg. 
humboldtianum. 
CORRECT NAME: Dinebra chondrosioides Kunth. 
= Bouteloua chondrosioides (Kunth) Benth. ex S. Watson. 


superfl., as = "Chondrosium" 


Chondrosum humile P. Beauv., Ess. Agrostogr. 41, 158. 
1812. 
NOTE: nom. nud.; as "Chondrosium". 
CORRECT NAME: Chondrosum tenue P. Beauv. ex Kunth. 
= Bouteloua simplex Lag. 


Chondrosum humile P. Beauv. ex Kunth, Nov. Gen. Sp. 1: 
175, t. 56. 1815 [1816]. T-Protol.: Crescit in frigidis 
exsiccatis, argillosis propter Llactacunga Quitensium 
et in monte Guadelupensi, cui urbs Santae Fidei 
Bogotensium subjecta est, alt. 1480-1690 hezap., 
floret Junio, Julio, Humboldt & Bonpland s.n. T- 
Specim.: | of 1. Ecuador: Crescit in frigidis exsiccatis, 
argillosis propter Llactacunga Quitensium et in Monte 
Guadelupensi, Humboldt & Bonpland s.n. [HT: P; IT: 
US-A865683 (fragm.)]. 

NOTE: as "Chondrosium" humile. 
= Bouteloua simplex Lag. 


Chondrosum karwinskii E. Fourn., Mexic. Pl. 2: 137. 1886. 
T-Protol.: Mexico: Cafion de las Minas et Victoria, 
Karwinski 1479 (HT: P?]. T-Specim.: | of 1. Mexico: 
Cafion de las Minas et Victoria, Karwinsky 1479 [HT: 
P?: IT: LE, US-A865682 (fragm.)]. 

NOTE: as "Chondrosium" karwinskii Fourn. 
= Bouteloua karwinskii(E. Fourn.) Griffiths. 


Chondrosum kayi (Warnock) Clayton, Kew Bull. 37: 418. 
1982. Basionym: Bouteloua kayi Warnock. 
= Bouteloua kayi Warnock. 


Chondrosum microstachyum E. Fourn., Mexic. Pl. 2: 138. 
1886. T-Protol.: Mexico: Guadalupe, Bourgeau 667. 
T-Specim.: | of 1. Mexico: Guadelupe, 1865-1866,M. 
Bourgeau 667 [HT: P; IT: LE, MO, US-74908, US- 
87194 (fragm.)]. 

NOTE: as "Chondrosium" microstachyum. 
= Bouteloua barbata Lag. 


Chondrosum oligostachyum (Nutt.) Torr., Explor. Red River 
Louisiana 300. 1853. Basionym: Atheropogon 
oligostachyus Nutt. 

NOTE: as "Chondrosium". 
= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Chondrosum papillosum (Engelm.) Torr., Explor. Red River 
Louisiana 300. 1853. Basionym: Atheropogon 
papillosus Engelm. 

NOTE: as "Chodrosium" pappilosum. 
= Bouteloua hirsuta var. hirsuta. 


Chondrosum parryi E. Fourn., Mexic. Pl. 2: 150. 1886. T- 
Protol.: Mexico: circa San Luis de Potosi, Parry & 
Palmer 923 1/2 [error for 943 1/2]. T-Specim.: | of 1. 
Mexico: San Luis Potosi: circa San Luis Potosi, Parry 
& Palmer 943 1/2 (HT: BR?;, IT: US-81783, US- 
883835]. 

NOTE: as "Chondrosium" parryi. 
REPLACED NAME: Chondrosium parryi E. Fourn. 
= Bouteloua parryi var. parryi. 


Chondrosum pectinatum (Feath.) Clayton, Kew Bull. 37: 418. 
1982. Basionym: Bouteloua pectinata Feath. 
= Bouteloua hirsuta subsp. pectinata (Feath.) Wipff & 
S.D. Jones. 


Chondrosum polystachyum Benth., Bot. Voy. Sulphur 56. 
1844. T-Protol.: Bay of Magdalena, Barclay s.n. T- 
Specim.: | of 1. Mexico: Baja California Sur: Bay of 
Magdalena, G.W. Barclay s.n. [HT: K; IT: BM, US- 
865566 (fragm. ex K)]. 

NOTE: as "Chondrosium". 
= Bouteloua barbata Lag. 


Chondrosum polystachyum Trin. ex E. Fourn., Mexic. Pl. 2: 
136. 1886. T-Specim.: | of 1. Mexico: Lauredo: 
Laredo, /427 [T: LE-TRIN-2130.01]. 

NOTE: nom. inval., as syn. of Chondrosium trinii Fourn.; as 
"Chondrosium". 
= Bouteloua trifida Thurb. 


Chondrosum procumbens (P. Durand) Desv., Nouv. Bull. Sci. 
Soc. Philom. Paris 2: 188. 1810. Basionym: Chloris 
procumbens P. Durand. 

NOTE: as "Chondrosium". 
= Bouteloua simplex Lag. 


Chondrosum procumbens (P. Durand) Desy. ex P. Beauv., 
Ess. Agrostogr. 1: 41, 158, t. 9, f. 7. 1812. Basionym: 
Chloris procumbens P. Durand. 

NOTE: isonym; as "Chondrosium". 

E. NAME: Chondrosum procumbens (P. Durand) Desv. 

CORRECT NAME: Chondrosum procumbens (P. Durand) 
Desv. 

= Bouteloua simplex Lag. 


Chondrosum prostratum (hort. ex Lag.) Sweet, Hort. Brit. I: 
455. 1826. Basionym: Bouteloua prostrata hort. ex 
Lag. 
NOTE: as "Chondrosium", 


Catalogue of New World Grasses: II. 55 


= Bouteloua simplex Lag. 


Chondrosum prostratum (hort. ex Lag.) Kunth, Revis. 
Gramin. 1: 94. 1829. Basionym: Boufeloua prostrata 
hort. ex Lag. 

NOTE: as "Chondrosium"; comb. illeg. superfl. 

E. NAME: Chondrosum prostratum (hort. ex Lag.) Sweet. 

CORRECT NAME: Chondrosum prostratum (hort. ex Lag.) 
Sweet. 

= Bouteloua simplex Lag. 


Chondrosum scorpioides (Lag.) Kunth, Revis. Gramin. 1: 94. 
1829. Basionym: Bouteloua scorpioides Lag. 
NOTE: as "Chondrosium". 
= Bouteloua scorpioides Lag. 


Chondrosum simplex (Lag.) Kunth, Revis. Gramin. |: 94. 
1829. Basionym: Bouteloua simplexLag. 
NOTE: as "Chondrosium" simplex. 
= Bouteloua simplex Lag. 


Chondrosum subscorpioides Mill. Hal., Bot. Zeitung (Berlin) 
14(20): 347. 1856. T-Protol.: Patria California, 
Barclay s.n., Watson donavit. T-Specim.: | of 1. 
Mexico: Baja California, Barclay s.n. [HT: 9; IT: K, 
US-883457 (fragm.)]. 

NOTE: as "Chondrosium" subscorpioides. 
= Bouteloua barbata Lag. 


Chondrosum tenue P. Beauv., Ess. Agrostogr. 41, 158. 1812. 
NOTE: nom. nud.; as "Chondrosium". 
CORRECT NAME: Chondrosum tenue P. Beauv. ex Kunth. 
= Bouteloua simplex Lag. 


Chondrosum tenue P. Beauv. ex Kunth, Nov. Gen. Sp. 1: 
176, t. 57. 1816. T-Protol.: Crescit in alta planitie 
Novae Hispaniae prope Mexico et San Agustin de las 
Cuevas, alt. 1160 hexap., floret Septembri, Humboldt 
& Bonpland s.n. T-Specim.: | of 1. Mexico: Novae 
Hispaniae prope Mexico et San Agustin de las Cuevas, 
Humbolt & Bonpland s.n. (HT: P?]. 

NOTE: as "Chondrosium" tenue. 
= Bouteloua simplex Lag. 


Chondrosum trifidum (Thurb.) Clayton, Kew Bull. 37: 418. 
1982. Basionym: Bouteloua trifida Thurb. 
= Bouteloua trifida Thurb. 


Chondrosum trinii E. Fourn., Mexic. Pl. 2: 136. 1886. T- 
Protol.: [USA]: Texas: Laredo, in prov., Berlandier 
1427, "Ch. (?) polystachyum Trin. in sched. non 
Benth.". T-Specim.: 1 of 1. USA: Texas: Webb Co.: 
Laredo, JL. Berlandier 1427 (HT: P; IT: GH, K, LE- 
TRIN-2130.01, US-883356, US-A1186684 (fragm. 1 
ex P, 2 ex W), US-204622, US-108418 (ex BM)]. 

NOTE: cites "Ch. (?) polystachyum Trin. in sched., non. 
Benth.". 
= Bouteloua trifida Thurb. 


Chondrosum virletii E. Fourn. ex Hemsl., Biol. Cent.-Amer., 
Bot. 3: 564. 1885. 
NOTE: as "Chondrosium". 
CORRECT NAME: Chondrosum virletii E. Fourn. 
= Bouteloua trifida Thurb. 


Chondrosum virletii E. Fourn., Mexic. Pl. 2: 136. 1886. T- 
Protol.; Mexico: San Luis de Potosi, Virlet 1373. T- 
Specim.: | of 2. Mexico: San Luis Potosi, 1851, Virlet 
1373 [LT: P; IT: US-865685 (fragm.)], LT designated 


by Griffiths, Contr. U.S. Natl. Herb. 14: 423 (1912). 2 
of 2. 

NOTE: as "Chondrosium" virletii. 

= Bouteloua trifida Thurb. 


Cinna arachnoidea (Poir.) Kunth, Revis. Gramin. |: 67. 
1829. Basionym: Agrostis arachnoides Poir. 
= Muhlenbergia expansa (Poir.) Trin. 


Cinna lanata (Kunth) Kunth, Revis. Gramin. 1: 67. 1829. 
Basionym: Agrostis lanata Kunth. 
= Muhlenbergia pubescens (Kunth) Hitchce. 


Cinna lateriflora (Michx.) Kunth, Revis. Gramin. 1: 67. 
1829. Basionym: Agrostis lateriflora Michx. 
= Muhlenbergia frondosa (Poir.) Fernald. 


Cinna macroura (Kunth) Kunth, Revis. Gramin. 1: 67. 1829. 
Basionym: Crypsis macroura Kunth. 
= Muhlenbergia macroura (Kunth) Hitchc. 


Cinna mexicana (L.) P. Beauv., Ess. Agrostogr. 32, 148, 158. 
1812. Basionym: Agrostis mexicana L. 
= Muhlenbergia mexicana (L.) Trin. 


Cinna phleoides (Kunth) Kunth, Revis. Gramin. 1: 67. 1829. 
Basionym: Crypsis phleoides Kunth. 
= Muhlenbergia angustata (J. Pres!) Kunth. 


Cinna pubescens (Kunth) Kunth, Revis. Gramin. 1: 67. 1829. 
Basionym: Agrostis pubescens Kunth. 
= Muhlenbergia pubescens (Kunth) Hitchc. 


Cinna racemosa (Michx.) Kunth, Revis. Gramin. 1: 67. 1829. 
Basionym: Agrostis racemosa Michx. 
= Muhlenbergia racemosa (Michx.) Britton, Sterns & 
Poggenb. 


Cinna setifolia (J. Pres!) Kunth, Revis. Gramin. Suppl. xvi. 
1830. Basionym: Crypsis setifolia J. Presl. 
= Muhlenbergia macroura (Kunth) Hitche. 


Cinna sobolifera (Muhl. ex Willd.) Link, Enum. Hort. Berol. 
Alt. 1: 71. 1821. Basionym: Agrostis sobolifera Muhl. 
ex Willd. 

= Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. 


Cinna stricta (Kunth) Kunth, Revis. Gramin. 1: 67. 1829. 
Basionym: Crypsis stricta Kunth. 
= Muhlenbergia angustata (J. Pres!) Kunth. 


Cinna tenuiflora (Willd.) Link, Enum. Hort. Berol. Alt. 1: 71. 
1821. Basionym: Agrostis tenuiflora Willd. 
= Muhlenbergia tenuiflora (Willd.) Britton, Sterns & 
Poggenb. 


Cladoraphis Franch., Bull. Mens. Soc. Linn. Paris 1: 
673. 1887. TYPE: Cladoraphis duparquetii Franch. 
TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: I. 


Cladoraphis cyperoides (Thunb.) S.M. Phillips, Kew Bull. 
37(1): 159. 1982. Basionym: Poa cyperoides Thunb. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Gibbs Russell, G. E, W. G. Welman, 
E. Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987). 


56 Catalogue of New World Grasses: II. 


SYN.: Brizopyrum cyperoides (Thunb.) Nees; Eragrostis 
cyperoides (Thunb.) P. Beauv.; Poa cyperoides 
Thunb. 


Clomena P. Beauv., Ess. Agrostogr. 28. 1812. TYPE: 
Clomena peruviana P. Beauv. 
= Muhlenbergia Schreb. 


Clomena peruviana P. Beauv., Ess. Agrostogr. 28, t. 7, f. 10; 
t. 3, f 20. 1812. T-Protol.: Peru: "Pl. nouv. 
Communiquee.", M. Thibaut s.n. T-Specim.: 1 of 1. 
Peru, Thibaut s.n. (HT: PJ. 

= Muhlenbergia peruviana (P. Beauv.) Steud. 


Clomena peruviana P. Beauv. var. peruviana. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Clomena peruviana var. pulvinata Nees, Nova Acta Phys.- 

Med. Acad. Caes. Leop.-Carol. Nat. Cur. 19(Suppl. 1): 
12. 1841. T-Protol.: Peru: "planis summis  iugi 
Peruviani ad lacum Titicacam: Apr, Meyen s.n. T- 
Specim.: | of 1. Peru: Lake Titicaca, Meyen s.n. (HT: 
B; IT; US-3376134 (fragm. ex B)]. 

NOTE: see also p. 144 (1843). 

= Muhlenbergia peruviana (P. Beauv.) Steud. 


Colpodium compressum Trin., Neue Entdeck. Pflanzenk. 2: 
58. 1821. 
NOTE: nom. inval., as syn. of Vilfa compressa Trin. 
= Muhlenbergia torreyana (Schult.) Hitche. 


Colpodium junceum (P. Beauv.) Trin., Neue Entdeck. 
Pflanzenk. 2: 37. 1821. Basionym: Heleochloa juncea 
P. Beauv. 
= Sporobolus junceus (P. Beauv.) Kunth. 


Cottea Kunth, Revis. Gramin. 1: 84. 1829. TYPE: Cottea 
pappophoroides Kunth. 
TRTD. BY: Peterson, P. M. 
NHT: Pappophoreae. NO. ACCPTD. SPP.: I. 


Cottea pappophoroides Kunth, Revis. Gramin. 1: 84. 1829. 
T-Protol.: Peru, Anon. [?]. T-Specim.: 1 of 1. Peru, 
1820, Coll. Ukn. s.n. (IT: US-A865660 (fragm. ex B- 
W)]. 

DST.: Arge, Boli, Ecua, Mexi, Peru, USA. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Beetle, A. A. (1987); Brako, L. & J. L. 
Zarucchi (1993); Correll, D. S. & M. C. Johnston 
(1970); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Hitchcock, A. S. (1927); 
Hitchcock, A. S. (1951); Jorgensen, P. M. & S. Leén- 
Yanez (eds.) (1999); Macbride, J. F. (1936); 
McVaugh, R. (1983); Renvoize, S. A. (1998); 
Shreve, F. & I. L. Wiggins (1964); Tovar, O. (1993); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 


Cotteinae subtrib. Reeder, Madrofio 18: 25. 1965. 
Craspedorhachis texana (Hitchc.) Pilg., Bot. Jahrb. Syst. 74: 


27. 1947. Basionym: Willkommia texana Hitche. 
= Willkommia texana Hitchc. 


Craspedorhachis texana var. stolonifera (Parodi) Pilg., Bot. 
Jahrb. Syst. 74: 27. 1945. 
= Willkommia texana var. stolonifera Parodi. 


Craspedorhachis texana Parodi var.texana. 
= Willkommia texana var. texana. 
Crossotropis Stapf, Fl. Cap. 7: 317. 1898. TYPE: 
Crossotropis grandiglumis (Nees) Rendle. 
= Trichoneura Andersson. 


Crypsidinae subtrib. Maire & Weiler, Fl. Afrique N, 2: 89. 
1953. 
NOTE: nom. inval., without Latin description. 


Crypsinna E. Fourn., Mexic. Pl. 2: 90. 1886. TYPE: 
Crypsinna macroura (Kunth) E. Fourn. 


= Muhlenbergia Schreb. 


Crypsinna breviglumis M.E. Jones, Contr. W. Bot. 14: 8. 
1912. T-Protol.: Mexico: Chihuahua: Guayanopa 
Canyon, Sierra Madre Mts., Sep 1903, ME. Jones s.n. 
T-Specim.: | of 1. Mexico: Chihuahua, 1903, ACE. 
Jones s.n. (IT: US-3168555 (fragm.)]. 

= Muhlenbergia dubia E. Fourn. 


Crypsinna macroura (Kunth) E. Fourn., Mexic. Pl. 2: 90. 
1886. Basionym: Crypsis macroura Kunth. 
= Muhlenbergia macroura (Kunth) Hitche. 


Crypsinna rigens (Benth.) M.E. Jones, Contr. W. Bot. 14: 8. 
1912. Basionym: Epicampes rigens Benth. 
= Muhlenbergia rigens (Benth.) Hitchc. 


Crypsinna setifolia (J. Presl) E. Fourn., Mexic. Pl. 2: 90. 
1886. Basionym: Crypsis setifolia J. Presl. 
= Muhlenbergia macroura (Kunth) Hitche. 


Crypsinna setiglumis M.E. Jones, Contr. W. Bot. 14: 8. 1912. 
T-Protol.: Mexico: Chihuahua: Marsh Lake, Sierra 
Madre Mts. near Garcia in dry pine woods: Sep 1903, 
M.E. Jones s.n. 

= Muhlenbergia dubia E. Fourn. 


Crypsinna stricta (Kunth) E. Fourn., Mexic. Pl. 2: 90. 1886. 
Basionym: Crypsis stricta Kunth. 
= Muhlenbergia angustata (J. Presl) Kunth. 


Crypsis Aiton, Hortus Kew, 1: 48. 7 Aug-1 Oct 1789. 
TYPE: Crypsis aculeata (L.) Aiton. 
NOTE: nom. cons. 
TRTD. BY: Peterson, P. M. 
NHT: Sporobolinae. NO. ACCPTD. SPP.: 3. 
CNSLTD. LIT.: Hammel, B. E. & J. R. Reeder, (1979). 
SYN.: Antitragus Gaertn.,; Ceytosis Munro; Heleochloa 
Host ex Roem.; Pallasia Scop.; Pechea Lapeyr.; 
Raddia Mazziari; Torgesia Bornm. 


Crypsis alopecuroides (Piller & Mitterp.) Schrad., Fl. Germ. 
1: 167. 1806. Basionym: Phleum alopecuroides Piller 
& Mitterp. 

DST.: USA, 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Baldini, R. M. (1995); Cronquist, A., 
A. H. Holmgren, N. H. Holmgren & J. L. Reveal 
(1977); Hammel, B. E. & J. R. Reeder (1979); 
Hickman, J. C. (1993). 

Crypsis explicata (Link) F. Herm.; Crypsis 
phalaroides M. Bieb.; Heleochloa alopecuroides 
(Piller & Mitterp.) Host; Phalaris explicata Link; 
Phalaris  geniculata’ Sibth, & Sm. Phleum 
alopecuroides Piller & Mitterp. 


SYN:: 


Catalogue of New World Grasses: II. 57 


Crypsis compacta Steud., Syn. Pl. Glumac. 1: 151. 1855. T- 
Protol.: Senegal. 
= Crypsis vaginiflora (Forssk.) Opiz. 


Crypsis explicata (Link) F. Herm., Hercynia 1: 121. 1937. 
Basionym: Phalaris explicata Link. 
NOTE: in obs.; cf. Mansf. Fedde, Repert, 45: 230, 1938. 
= Crypsis alopecuroides (Piller & Mitterp.) Schrad. 


Crypsis juncea (P. Beauv.) Steud., Nomencl. Bot. 1: 242. 
1821. Basionym: Heleochloa juncea P. Beauv. 
= Sporobolus junceus (P. Beauv.) Kunth. 


Crypsis macroura Kunth, Nov. Gen. Sp. 1: 140. 1815 [1816]. 
T-Protol.: Crescit in apricis montanis regni Mexicani 
juxta montem Tolucae, alt. 1760 hexap. Floret 
Septembri, Humboldt & Bonpland s.n. T-Specim.: | of 
1. Mexico: México: near the Nevado de Toluca, Sep, 
F_W.H.A. von Humboldt & A.J.A. Bonpland s.n. (HT: 
P; IT: US-A865654 (fragm.)]. 

= Muhlenbergia macroura (Kunth) Hitchc. 


Crypsis niliacea Fig. & De Not., Mem. Acad, Roy. Sci. Turin 
14: 322. 1854. T-Protol.: Egypt: In insulis niloticis 
Algypti inferioris. 

= Crypsis vaginiflora (Forssk.) Opiz. 


Crypsis phalaroides M. Bieb., Fl. Taur.-Caucas. 1: 45. 1808. 
T-Protol.: Prouenit in desertis sabulosis Caucaso 
finitimis: etiam in sabulis ad Wolgam circa coloniam 
Sareptanam. 

= Crypsis alopecuroides (Piller & Mitterp.) Schrad. 


Crypsis phleoides Kunth, Nov. Gen. Sp. 1: 140. 1815 [1816]. 
T-Specim.: 1 of 1. Venezuela: Sucre, no date, 
F_W.H.A. von Humboldt & A.J.A. Bonpland s.n. (HT: 
P; IT: US-A865652 (fragm. ex P)]. 

= Muhlenbergia angustata (J. Presl) Kunth. 


Crypsis schoenoides (L.) Lam., Tabl. Encycl. 1: 166, t. 42, f. 
1.1791. Basionym: Phleum schoenoides L. 

DST.: Mexi, USA. 

LVL. ACCPTC.: |; introduced. 

ALSO ACCPTD. BY: Beetle, A. A. (1987); Bor, N. L. 
(1960); Cronquist, A., A. H. Holmgren, N. H. 
Holmgren & J. L. Reveal (1977); Espejo Serna, A., 
A. R. L6opez-Ferrari, and J. Valdes-Reyna (2000); 
Gleason, H. A. & A. Cronquist (1991); Gould, F. W. 
& R. Moran (1981); Hammel, B. E. & J. R. Reeder 
(1979); Hickman, J. C. (1993); Phillips, S. (1995). 

Heleochloa schoenoides (L.) Host; Phleum 
schoenoides L. 


Crypsis setifolia J. Pres|, Relig. Haenk. 1(4-5): 245. 1830. T- 
Protol.: Hab. in Mexico, Haenke s.n. T-Specim.: 1 of 
1. Mexico, 7. Haenke s.n. [IT: US-A865655 (fragm.)]. 

= Muhlenbergia macroura (Kunth) Hitchc. 


SYN.: 


Crypsis squarrosa Nutt., Gen. N. Amer. PI. 1: 49. 1818. T- 
Protol.: [USA]: South Dakota: On arid plains near the 
Grand Deour of the Missouri, Nutall s.n. 

= Munroa squarrosa (Nutt.) Torr. 


Crypsis stricta Kunth, Nov. Gen. Sp. 1: 140. 1815 [1816]. T- 
Protol.: Crescit in frigidis, praeruptis Andium inter 
Ipiales et Guachucal, alt. 1614 hexap., Humboldt & 
Bonpland s.n. T-Specim.: 1 of 1. Peru: Prov. de los 
Pastos, no date, Humboldt & Bonpland s.n. [IT: US- 
A865653 (fragm. ex B-W)]. 

= Muhlenbergia angustata (J. Presl) Kunth. 


Crypsis vaginiflora (Forssk.) Opiz, Naturalientausch 8: 83. 
1823. Basionym: Phalaris vaginiflora Forssk. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Baldini, R. M. (1995); Gould, F. W. 
& R. Moran (1981); Hammel, B. E. & J. R. Reeder 
(1979); Hickman, J. C. (1993); Phillips, S. (1995). 

SYN.: Crypsis compacta Steud.; Crypsis niliacea Fig. & De 
Not.; Heleochloa compacta (Steud.) T. Durand & 
Schinz; Phalaris vaginiflora Forssk. 


Crypsis virginica (L.) Nutt., Gen. N. Amer. Pl. 1: 49. 1818. 
Basionym: Agrostis virginica L. 
= Sporobolus virginicus (L.) Kunth. 


Cryptostachys Steud., Syn. Pl. Glumac. |: 181. 1854. TYPE: 
Cryptostachys vaginata Steud. 
= Sporobolus R. Br. 


Cryptostachys vaginata Steud., Flora 33: 229. 1850. T- 
Protol.: [USA]: "Am. septr." North America. 
NOTE: nom. nud. 
= Sporobolus vaginiflorus (Torr. ex A. Gray) Alph. Wood. 


Cryptostachys vaginata Steud., Syn. Pl. Glumac. 1: 181. 
1854. T-Protol.: [USA]: "Am. septr." North America. 
= Sporobolus vaginiflorus (Torr. ex A. Gray) Alph. Wood. 


Ctenium Panz., [deen Revis. Gras. 38, 61. 1813. TYPE: 
Ctenium carolinianum Panz. 
NOTE: nom. cons. 
TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 10. 
CNSLTD. LIT.: Longhi-Wagner, H. M., (1988). 
SYN.: Aplocera Raf.; Campuloa Desv.; Campulosus Desv.; 
Monathera Raf.; Monocera Elliott; Triatherus Raf. 


Ctenium americanum Spreng., Syst. Veg. 1: 274. 1825. T- 
Protol.: Amer. bor. T-Specim.: | of 1. USA: Carolina, 
no date, L.4.G. Bosc s.n. [IT: US-2606901 (fragm. ex 
B)]. 


= Ctenium aromaticum (Walter) Alph. Wood. 


Ctenium aromaticum (Walter) Alph. Wood, Class-book 


Bot. (ed. 1861) 806. 1861. Basionym: Aegilops 
aromatica Walter. 
DST.: USA. 


LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Anonymous (1986); Darke, R. 
(1999); Fernald, M. (1950); Gleason, H. A. & A. 
Cronquist (1968); Gleason, H. A. & A. Cronquist 
(1991); Godfrey, R. K. & J. W. Wooten (1979); 
Hitchcock, A. S. (1951); Radford, A. E., H. E. Ahles 
& C. R. Bell (1968); Swallen, J. R. (1939); 
Wunderlin, R. P. (1998). 

SYN.: Aegilops aromatica Walter; Aplocera maritima Raf., 
Campuloa gracilis Desv.; Campuloa monostachya 
(Michx.) Roem. & Schult.; Campulosus aromaticus 
(Walter) Scribn.; Campulosus aromaticus Trin. ex 
Steud.; Campulosus  gangitis (L.) Kuntze; 
Campulosus gracilior Desv.; Campulosus gracilis 
Bertol.; Campulosus | monostachyus (Michx.)  P. 
Beauv.; Chloris monostachya Michx.; Chloris 
piperita Michx. ex Steud.; Clenium americanum 
Spreng.; Cfenium aromaticum (Walter) Hitchc.; 
Ctenium carolinianum Panz.; Ctenium gangitum (L.) 
Druce; Cynodon monostachyos (P. Beauv.) Raspail; 


58 Catalogue of New World Grasses: II. 


Monerma gangitis (L.) Roem. & Schult.; Monocera 
aromatica (Walter) Elliott; Nardus gangitis L.; 
Nardus scorpioides Lam.;, Rottboellia scorpioides 
Poir. ex Steud.; Triatherus aromaticus Raf. 


Ctenium aromaticum (Walter) Hitche., Rhodora 8(95): 210. 
1906. Basionym: Aegilops aromatica Walter. 
NOTE: isonym. 
E. NAME: Ctenium aromaticum (Walter) Alph. Wood. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Ctenium brachystachyum (Nees) Kunth, Enum. Pl. 1: 275. 
1833. Basionym: Campulosus brachystachyus Nees. 
NOTE: also Revis. Gram. 3: Suppl. 1834 p. (as in 1835 ed.) 
XXIl. 
DST.: Braz, Para. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: D6ll, J. C. (1878); Filgueiras, T. S. 
(1991); Filgueiras, T. S. (1999); Longhi-Wagner, H. 
M. (1988); Renvoize, S. A. (1988). 
SYN.: Campulosus brachystachyus Nees. 


Ctenium brevispicatum J.G. Sm., Bot. Gaz. 21: 363. 1896. 
T-Protol.: Brazil: Campulosus brachystachyus Nees ab 
Esenb. 

NOTE: based on Campulosus brachystachyus sensu. Trin., 
Sp. Gram. 3: 302 (1836). 

DST.: Braz. 

LVL. ACCPTC:: |. 

ALSO ACCPTD. BY: Filgueiras, T. S. (1991); Filgueiras, 
T. S. (1999); Longhi-Wagner, H. M. (1988); Longhi- 
Wagner, H. M. (1990); Renvoize, S. A. (1988). 

SYN.: Crenium trinii Ekman. 


Ctenium carolinianum Panz., Denkschr. Bayer. Bot. Ges. 
Regensburg 4: 311, t. 13, f. 1-2. 1813 [1814]. T- 
Protol.: USA: Habitat in sylvis Carolinae inferiores. 

NOTE: nom. illeg. superfl. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Ctenium chapadense (Trin.) Dll, Fl. Bras. 2(3): 73. 1878. 
Basionym: Campulosus chapadensis Trin. 

DST.: Boli, Braz. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: DOIl, J. C. (1878); Filgueiras, T. S. 
(1991); Filgueiras, T. S. (1999); Longhi-Wagner, H. 
M. (1988); Longhi-Wagner, H. M. (1990); Renvoize, 
S. A. (1984). 

SYN.: Campuloa chapadensis Trin. ex Déll; Campulosus 
chapadensis Trin. 


Ctenium cirrosum (Nees) Kunth, Revis. Gramin. 2: 445, t. 
136. 1831. Basionym: Campulosus cirrosus Nees. 

DST.: Braz, Suri. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Doll, J. C. (1878); Filgueiras, T. S. 
(1991); Filgueiras, T. S. (1999); Judziewicz, E. J. 
(1990); Longhi-Wagner, H. M. (1990). 

SYN.: Campulosus cirrosus Nees. 


Ctenium concissum Swallen, N. Amer. Fl. 17(8): 601. 1939. 
T-Protol.: Panama: Coclé: collected at Picacho de Ola, 
alt. 350-600 m, Pittier 5065 |[HT: US-715193]. T- 
Specim.: | of 1. Panama: Coclé: Pichaché de Ola, 8 
Dec 1911, H. Pittier 5065 (HT: US-715193]. 
DST.: Pana. 
LVL. ACCPTC.: I. 


ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Pohl, R. W. & G. 
Davidse (1994); Swallen, J. R. (1939). 


Ctenium floridanum (Hitchc.) Hitchce., Proc. Biol. Soc. 


Wash. 41: 162. 1928. Basionym: Campulosus 
floridanus Hitche. 
DST.: USA. 


LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Hitchcock, A. S. (1951); Swallen, J. 
R. (1939); Wunderlin, R. P. (1998). 

SYN.: Campulosus floridanus Hitche. 


Ctenium gangitum (L.) Druce, Bot. Exch. Club Brit. Isles 
Rep. 3: 416. 1914. Basionym: Nardus gangitis L. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Crenium glandulosum Scribn. & J.G. Sm., Bot. Gaz. 21(6): 
362. 1896. T-Protol.: Mexico: Oaxaca: collected 
between Nietepec and Zanatepec, 15 Jul 1895, Nelson 
2814. T-Specim.: | of 1. Mexico: Oaxaca: between 
Nietepec and Zanatepec, 15 Jul 1895, Nelson 28/4 
{[HT: US-878837]. 

= Ctenium planifolium (J. Pres!) Kunth. 


Ctenium planifolium (J. Presl) Kunth, Enum. Pl. 1: 275, 
1833. Basionym: Campulosus planifolius J. Presl. 

DST.: Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
L6pez-Ferrari, and J. Valdes-Reyna (2000); Pohl, R. 
W. & G. Davidse (1994); Swallen, J. R. (1939). 

Campulosus  planifolius J. Presl; =Ctenium 
glandulosum Scribn. & J.G. Sm. 


SYN.: 


Ctenium plumosum (Hitche.) Swallen, N. Amer. Fl. 17(8): 
602. 1939. Basionym: Campulosus plumosus Hitche. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Beetle, A. A. 
(1987); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); McVaugh, R. (1983); Swallen, 
J. R. (1939). 

SYN.: Campulosus plumosus Hitche. 


Ctenium polystachyum Balansa, Bull. Soc. Bot. France 32: 
244. 1885. T-Specim.: | of 3. Paraguay, 18 1844, B. 
Balansa 4341 (PL. DU PARAG.,) |ST: US-2606905}]. 2 
of 3. Paraguay: 24 1876, B. Balansa 196 (Pl du 
Parag.) [ST: US-865648 (fragm. ex P)]. 3 of 3. 
Paraguay, Balansa 2962 [ST]. 

DST.: Braz, Para. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Longhi-Wagner, H. M. (1988); 
Renvoize, S. A. (1988). 


Ctenium trinii Ekman, Ark. Bot. 13(10): 45. 1913. T-Protol.: 
Brazil. 
NOTE: Based on Campulosus brachystachyus Nees sensu 
Trin., Sp. Gram. 3: 302 (1836). 
CORRECT NAME: Crenium brevispicatum J.G. Sm. 
= Ctenium brevispicatum J.G. Sm. 


Cyclostachya Reeder & C. Reeder, Bull. Torrey Bot. Club 
90: 195. 1963. TYPE: Cyclostachya — stolonifera 
(Scribn.) Reeder & C. Reeder. 

= Bouteloua Lag. 


Catalogue of New World Grasses: II. 59 


Cyclostachya stolonifera (Scribn.) Reeder & C. Reeder, Bull. 
Torrey Bot. Club 90: 196. 1963. Basionym:Bouteloua 
stolonifera Scribn. 

= Bouteloua reederorum Columbus. 


Cynodon Rich., Syn. Pl. 1: 85. 1805. TYPE: Cynodon 
dactylon (L.) Pers. 
NOTE: nom. cons. 
TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 17 SUBSP.: 13. 
CNSLTD. LIT.: Harlan, J. R., J. M. J. de Wet, W. W. 
Huffine & J. R. Deakin, (1970); Caro, J. A. & E. 
Sanchez, (1969). 
SYN.: Capriola Adans.: Dactilon Vill.; Fibichia Koeler. 


Cynodon abyssinicus (Jacq.) Raspail, Ann. Sci. Nat. (Paris) 5: 
302. 1825. Basionym: Poa abyssinica Jacq. 
= Eragrostis tef (Zucc.) Trotter. 


Cynodon aethiopicus Clayton & J.R. Harlan, Kew Bull. 24: 
187. 1970. T-Protol.: Ethiopia, de Wet OKLA9224 
(HT: K]. 

DST.: USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Clayton, W. D. & J. R. Harlan 
(1970); Gibbs Russell, G. E, W. G. Welman, E. 
Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987): Phillips, S. (1995). 


Cynodon affinis Caro & E.A. Sanchez, Kurtziana 5: 223, f. 
7. 1969. T-Protol.: Argentina: Camino entre General 
Madariaga y Conesa, 28 Jan 1960, Nicora 6976 [HT: 
BAA]. 

DST.: Arge, Braz, Chil, Urug. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 
(1987); Caro, J. A. & E. Sanchez (1969); Caro, J. A. 
& E. Sanchez (1970); Marticorena, C. & M. Quezada 
(1985); Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

SYN.: Cynodon dactylon var. affinis (Caro & E.A. Sanchez) 
Romero Zarco. 


Cynodon amabilis (L.) Raspail, Ann. Sci. Nat., Bot. 5: 302. 
1825. Basionym: Poa amabilis L. 
= Eragrostis amabilis (L.) Wight & Arn. ex Nees. 


Cynodon aristiglumis Caro & E.A. Sanchez, Kurtziana 5: 
236, f. 11. 1969. T-Protol.: Argentina: Rio Negro: 
Dpto. Avellaneda: Isla Choele-Choel, Brazo Sud del 
rio Negro-Sifon, 14 Feb 1948, Boelcke & Serrano 
3171 (HT: BAA]. 

= Cynodon dactylon (L.) Pers. 


Cynodon aristulatus Caro & E.A. Sanchez, Darwiniana 17: 
514, f. 2. 1972. T-Protol.: Peru: San Martin: Tingo 
Maria, 625-1100 m, near Huallaga River at Tingo 
Maria, 30 Oct 1949-19 Feb 1951, 1.4. Allard 22027 
[HT: US-2011844]. T-Specim.: 1 of 1. Peru: San 
Martin, 30 Oct 1949-19 Feb 1950, HA. Allard 22027 
[HT: US-2011844]. 

= Cynodon dactylon (L.) Pers. 


Cynodon carolinianus (P. Beauv.) Raspail, Ann. Sci. Nat., 
Bot. 5: 302. 1825. Basionym: Tricuspis caroliniana P. 
Beauv. 


= Tridens flavus (L.) Hitche. 


Cynodon ciliatus (Sw.) Raspail, Ann. Sci. Nat. (Paris) 5: 303. 
1825. Basionym: Chloris ciliata Sw. 
= Chloris ciliata Sw. 


Cynodon cruciatus (L.) Raspail, Ann. Sci. Nat. (Paris) 5: 303. 
1825. Basionym: Agrostis cruciata L. 
= Chloris cruciata (L.) Sw. 


Cynodon curtipendula (Michx.) Raspail, Ann. Sci. Nat. 
(Paris) 5: 303. 1825. Basionym: Chloris curtipendula 
Michx. 

= Bouteloua curtipendula (Michx.) Torr. 


Cynodon cynosuroides (L.) Raspail, Ann. Sci. Nat., Bot. 5: 
303. 1825. Basionym: Dactylis cynosuroides L. 
= Spartina cynosuroides (L.) Roth. 


Cynodon dactylon (L.) Pers., Syn. Pl 1: 85, 
Basionym: Panicum dactylon L. 

DST.: Arge, Beli, Boli, Braz, Cana, Cari, Chil, CoRi, Ecua, 
ElSa, FrGu, Guat, Guya, Hond, Mexi, Nica, Pana, 
Peru, Suri, USA, Urug, Vene. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Adams, C. D. (1972): Allem, A. C, 
& J. F. M. Valls (1987); Anderson, D. E. (1974); 
Balick, M. J.. M. H. Nee & D. E. Atha (2000); 
Beetle, A. A. (1977); Beetle, A. A. (1987); 
Berendsohn, W. G. & A. E. Araniva de Gonzalez 
(1991): Bor, N. L. (1960); Brako, L. & J. L. Zarucchi 
(1993); Breedlove, D. E. (1986); Burkart, A. (1969); 
Caro, J. A. & E. Sanchez (1969); Caro, J. A. & E. 
Sanchez (1970); Catasus Guerra, L. (1997); Clayton, 
W.D. & J. R. Harlan (1970); Correll, D. S. & M. C. 
Johnston (1970); Cowan, C. P. (1983); Cronquist, A., 
A. H. Holmgren, N. H. Holmgren & J. L. Reveal 
(1977); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Dodson, C. H. & A. H. 
Gentry (1978); Doll, J. C. (1878); Edgar, E., H. E. 
Connor & J. E. Shand (1991); Espejo Serna, A., A. 
R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Fernald, M. (1950); Filgueiras, T. S. (1991): 
Filgueiras, T. S. (1999): Foster, R. C. (1958); Gibbs 
Russell, G. E, W. G. Welman, E. Reitief, K. L. 
Immelman, G. Germishuizen, B. J. Pienaar, M. van 
Wyk, and A. Nicholas (PRECIS) (1987): Gleason, H. 
A. & A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Gould, F. W (1979): Gould, F. W. 
& R. Moran (1981); Great Plains Flora Association 
(1986); Hickman, J. C. (1993); Hitchcock, A. S. 
(1927); Hitchcock, A. S. (1936); Hitchcock, A. S. 
(1951); Hitchcock, C. L., A. Cronquist, M. Ownbey 
& J. W. Thompson (1969); Jorgensen, P. M. & C. 
Ulloa U. (1994); Jorgensen, P. M. & S. Leon-Yanez 
(eds.) (1999); Judziewicz, E. J. (1990); Killeen, T. J. 
(1990); Lawesson, J. E., H. Adsersen & P. Bentley 
(1987); Lazarides, M. (1972); Levin, G. A. & R. 
Moran (1989); Lindeman, J. C. & A. L. Stoffers 
(1963); Liogier, H. & L. Martorell (1982); Long, R. 
W. & O. Lakela (1971); Marticorena, C. & M. 
Quezada (1985); McClure, F. A. (1955); McVaugh, 
R. (1983); Molina R., A. (1975); Munz, P. A. & D. 
D. Keck (1959); Munz, P. A. (1974); Nicora, E. G., 
M. E. De Paula, A. M. Faggi, M. del C. Mariano, A. 
M. Miante A., L. R. Parodi, C. A. Petetin, F. A. Roig 
& Z. Rugolo de Agrasar (1978); ORSTOM (1988); 
Phillips, S. (1995); Pohl, R. W. (1980); Pohl, R. W. 


1805. 


60 Catalogue of New World Grasses: II. 


(1994); Porter, D. M. (1983); Radford, A. E., H. E. 
Ahles & C. R. Bell (1968); Reeder, J. R. & C. G. 
Reeder (1971); Renvoize, S. A. (1984); Renvoize, S. 
A. (1988); Renvoize, S. A. (1998); Rosengurtt, B., B. 
R. Arrillaga de Maffei & P. Izaguirre de Artucio 
(1970); Santos, A. M. P. V. dos & S. de Castro 
Boechat (1994); Scoggan, H. J. (1978 [1979]); 
Shreve, F. & I. L. Wiggins (1964); Skottsberg, C. 
(1922); Smith, L. B., D. C. Wasshausen & R. M. 
Klein (1981); Sousa S., M. & E. F. Cabrera C. 
(1983); Tovar, O. (1993); Veldkamp, J. F. (1973); 
Voss, E. G. (1972); Wiggins, I. L. & D. M. Porter 
(1971); Wunderlin, R. P. (1998). 

Agrostis bermudiana Tussac ex Kunth; Agrostis 
filiformis J. Konig ex Kunth; Capriola dactylon (L.) 
Hitche.; Capriola dactylon (L.) Kuntze; Chloris 
cynodon Trin.; Chloris paytensis Steud.; Cynodon 
aristiglumis Caro & E.A. Sanchez; Cynodon 
aristulatus Caro & E.A, Sanchez; Cynodon dactylon 
fo. vivipara Beetle; Cynodon dactylon subsp. 
glabratus (Steud.) Chev.; Cynodon dactylon var. 
elegans Rendle; Cynodon dactylon var. glabratus 
(Steud.) Chiov.; Cynodon erectus J. Presl; Cynodon 
glabratus Steud.; Cynodon occidentalis Willd. ex 
Steud.; Cynodon pascuus Nees; Cynodon polevansii 
Stent; Cynodon portoricensis Willd. ex Steud.; 
Cynodon tenuis Trin. ex Spreng.; Cynodon 
umbellatus (Lam.) Caro; Cynosurus dactylon (L.) 
Pers.; Cynosurus uniflorus Walter; Dactilon 
officinale Vill.; Digitaria dactylon (L.) Scop.; 
Digitaria glumaepatula (Steud.) Miq.; Digitaria 
littoralis Salisb.; Digitaria stolonifera  Schrad.; 
Fibichia dactylon (L.) Beck; Fibichia umbellata 
Koeler; Milium dactylon (L.) Moench; Panicum 
dactylon L.; Panicum glumaepatulum  Steud.; 
Paspalum dactylon (L.) Lam.; Paspalum umbellatum 
Lam.; Phleum dactylon Pall. ex Georgi. 


SYN.: 


Cynodon dactylon var. affinis (Caro & E.A. Sanchez) 
Romero Zarco, Lagascalia 14(1): 171. 1986. 
Basionym: Cynodon affinis Caro & E.A. Sanchez. 

= Cynodon affinis Caro & E.A. Sanchez. 


Cynodon dactylon var. aridus J.R. Harlan & de Wet, Crop 
Sci. (Madison) 9: 774. 1969. T-Protol.: South Africa: 
Buffelsvlei, Huffine 41-219 [HT: CEL]. 

DST.: USA. 
LVL. ACCPTC.: |; introduced. 


Cynodon dactylon var. biflorus Merino, Fl. Galicia 3: 310. 
1909. T-Protol.: (Herbar. nim.1.938).La variedad muy 
rara vive en nuestra costa entre La Guardia y la 
parroqui de Oya, y tambien en la Toja, Pontevedra. 

DST.: Arge, Braz. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Burkart, A. (1969); Caro, J. A. & E. 
Sanchez (1969); Caro, J. A. & E. Sanchez (1970); 
Santos, A. M. P. V. dos & S. de Castro Boechat 
(1994). 


Cynodon dactylon (L.) Pers. var. dactylon, -. 

DST.: Arge, Braz, Cana, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Burkart, A. (1969); Caro, J. A. & E. 
Sanchez (1969); Caro, J. A. & E. Sanchez (1970); 
Santos, A. M. P. V. dos & S. de Castro Boechat 
(1994). 


Cynodon dactylon var. elegans Rendle, Cat. Afr. Pl. 2(1): 
221. 1899. T-Specim.: | of 1. Angola, Welwitsch 2290 
{IT: K]. 
= Cynodon dactylon (L.) Pers. 


Cynodon dactylon subsp. glabratus (Steud.) Chev., Rev. Int. 
Bot. Appl. Agric. Trop. 27(297-298): 282. 1947. 
Basionym: Cynodon glabratus Steud. 

= Cynodon dactylon (L.) Pers. 


Cynodon dactylon var. glabratus (Steud.) Chiov., Nuovo 
Giorn. Bot. Ital. 26: 82. 1919. Basionym: Cynodon 
glabratus Steud. 

= Cynodon dactylon (L.) Pers. 


Cynodon dactylon to. glabrescens (Beck.) Sod, Acta Bot. 
Acad. Sci. Hung. 17(1-2): 123. 1971 [1972]. 
NOTE: nom. inval., basionym not indicated. 
= Cynodon hirsutissimus (Litard. & Maire) Caro & E.A. 
Sanchez. 


Cynodon dactylon subvar. hirsutissima Litard. & Maire, 
Mem. Soc. Sci. Nat. Maroc 4(1): 25. 1924. T-Protol.: 
Morocco, Casablanca, Litardiere. 

= Cynodon hirsutissimus (Litard. & Maire) Caro & E.A. 
Sanchez. 


Cynodon dactylon var. hirsutissimus (Litard. & Maire) Maire, 
Bull. Soc. Hist. Nat. Afrique N. 30: 368. 1939, 
Basionym: Cynodon dactylon subvar. hirsutissima 
Litard. & Maire. 

= Cynodon hirsutissimus (Litard. & Maire) Caro & E.A. 
Sanchez. 


Cynodon dactylon var. longiglumis Caro & E.A. Sanchez, 
Kurtziana 5: 210, f. 2G. 1969. T-Protol.: Argentina: 
Tucuman: Dpto. Capital: prope urbem, Mar 1918, 
Schreiter s.n. (=Herb Osten 12150) (HT: BAA}. 

DST.: Arge, Para, Urug. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Caro, J. A. & E. Sanchez (1969); 
Caro, J. A. & E. Sanchez (1970). 


Cynodon dactylon var. maritimus (Kunth) Hack., Ark. Bot. 
8(8): 40. 1909. 
= Cynodon maritimus Kunth. 


Cynodon dactylon var. pilosus Caro & E.A. Sanchez, 
Kurtziana 5: 210. 1969. T-Protol.: Argentina: 
Cordoba: Dpto. Capital: Ciudad de Cordoba, Barrio 
San Martin, orilla del rio Primero, 17 Apr. 1952,Caro 
333 [HT: BAA; IT: JC]. 

DST.: Arge, Braz, Para. 

LVL. ACCPTC.:: I. 

ALSO ACCPTD. BY: Caro, J. A. & E. Sanchez (1969); 
Caro, J. A. & E. Sanchez (1970); Santos, A. M. P. V. 
dos & S. de Castro Boechat (1994). 


Cynodon dactylon var. sarmentosus Parodi, Revista Argent. 
Agron. 23: 185. 1956. T-Specim.: | of 1. Argentina, 
Luna s.n. [TY: K]. 
NOTE: as Cynodon dactylon var. "sarmentosum". 
CORRECT NAME: Cynodon parodii Caro & E.A. 
Sanchez. 
= Cynodon nlemfuensis Vanderyst. 


Cynodon dactylon to. vivipara Beetle, Phytologia 48(2): 189. 
1981. 
= Cynodon dactylon (L.) Pers. 


Catalogue of New World Grasses: II. 61] 


Cynodon diffusus (Willd.) Raspail, Ann. Sci. Nat., Bot., ser. 
5, 5: 303. 1825. Basionym: Muhlenbergia diffusa 
Willd. 

= Muhlenbergia schreberi J.F. Gmel. 


Cynodon distichloides Caro & E.A. Sanchez, Kurtziana 5: 
254, f. 1. 1969. T-Protol.: Uruguay: Piriapolis, verano 
1924, Bordale s.n. [HT: BAA]. 
DST.: Urug. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Sanchez, E. & J. A. Caro (1969). 


Cynodon domingense (Jacq.) Raspail, Ann. Sci. Nat., Bot. 5: 
302. 1825. Basionym: Cynosurus domingensis Jacq. 
= Leptochloa virgata (L.) P. Beauv. 


Cynodon erectus J. Presl, Reliq. Haenk. 1(4-5): 290. 1830. T- 
Protol.: Hab. in Mexico, et in Peruviae montanis 
huanaccensibus, Haenke s.n. T-Specim.: | of 1. 
Mexico. Peruano montano, Haenke s.n. [HT: PR- 
495767; IT: PRC], HT indicates both Peru and 
Mexico; IT indicates Mexico only. Protologue 
indicates both Mexico and Peru. Caro & Sanchez, 
Darwiniana 17: 522 (1972) consider the species to 
occur in Peru, not Mexico. 

= Cynodon dactylon (L.) Pers. 


Cynodon fascicularis (Lam.) Raspail, Ann. Sci. Nat., Bot. 5: 
303. 1825. Basionym: Festuca fascicularis Lam. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Cynodon glabratus Steud., Syn. Pl. Glumac. 1: 212. 1854. T- 
Specim.: | of 3. Ethiopia, Schimper 3/1] (ST: P]. 2 of 
3. , Schimper 311 (IST: K]. 3 of 3. , Schimper 431 [ST: 
P 


= Cynodon dactylon (L.) Pers. 


Cynodon grandispiculus Caro & E.A. Sanchez, Kurtziana 5: 
258, f. 2. 1969. T-Protol.: Uruguay: Montevideo, 
Felippone 4555 (HT: SI. 

= Cynodon maritimus var. grandispiculus Caro & E.A. 
Sanchez. 


Cynodon hirsutissimus (Litard. & Maire) Caro & E.A. 
Sanchez, Kurtziana 5: 220. 1969. Basionym: Cynodon 
dactylon subyar. hirsutissima Litard. & Maire. 

DST.: Arge, Braz, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Caro, J. A. & E. Sanchez (1969); 
Caro, J. A. & E. Sanchez (1970). 

SYN.: Cynodon dactylon fo. glabrescens (Beck.) Soo; 
Cynodon dactylon subvar. hirsutissima Litard. & 
Maire; Cynodon dactylon var. hirsutissimus (Litard. 
& Maire) Maire. 


Cynodon hirsutus Stent, Bothalia 2: 277, 286. 1927. T- 
Protol.: Transvaal, Burtt-Davy, several syntypes cited. 

DST.: Arge, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Burkart, A. (1969); Caro, J. A. & E. 
Sanchez (1969); Caro, J. A. & E. Sanchez (1970); 
Gibbs Russell, G. E, W. G. Welman, E. Reitief, K. L. 
Immelman, G. Germishuizen, B. J. Pienaar, M. van 
Wyk, and A. Nicholas (PRECIS) (1987). 

Cynodon incompletus var. hirsutus (Stent) de Wet. & 
J.R. Harlan. 


SYN.: 


Cynodon hirsutus Stent var. hirsutus. 
DST.: Arge, Urug. 


LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Caro, J. A. & E. Sanchez (1969); 
Caro, J. A. & E. Sanchez (1970). 


Cynodon hirsutus var. sesquiflorus Parodi, Revista Argent. 
Agron. 17(3): 216, f. 14. 1950. T-Protol.: Argentina: 
Buenos Aires: Partido Tornquist: Sierra de la Ventana, 
9 Jan 1950, Petetin 514 (HT: BAA; IT: SI]. 

DST.: Arge. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Caro, J. A. & E. Sanchez (1969); 
Caro, J. A. & E. Sanchez (1970); Zuloaga, F. O., E. 
G. Nicora, Z. E. Ragolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 


Cynodon incompletus var. hirsutus (Stent) de Wet & J.R. 
Harlan, Crop Sci. (Madison) 10: 291. 1970. Basionym: 
Cynodon hirsutus Stent. 

= Cynodon hirsutus Stent. 


Cynodon indicus (L.) Raspail, Ann. Sci. Nat., Bot. 5: 303. 
1825. Basionym: Cynosurus indicus L. 
= Eleusine indica (L.) Gaertn. 


Cynodon laeviglumis Caro & E.A. Sanchez, Kurtziana 5: 
217, f. 4. 1969. T-Protol.: Argentina: La Pampa: 
Depto. Leventue: Victoria, Carro Quemado, Estancia 
las Vertientes, Feb 1960, Legaspi s.n. [HT: BAA- 
2237]. 

DST.: Arge. 
LVL. ACCPTC.:: 1. 
ALSO ACCPTD. BY: Caro, J. A. & E. Sanchez (1969). 


Cynodon x magennisii Hurcombe, J. S. African Bot. 13: 
109. 1947. T-Protol.: Experimental plots — at 
Frankenwald, the Botanical Research Station of the 
University of Witwatersrand, known to gardeners as 
‘Magennis'. 

NOTE: pro. hyb. [Cynodon dactylon (L.) Pers. x C. 
transvaalensis Burtt Davy]. 

DST.: USA. 

LVL. ACCPTC.: 1; introduced. 


Cynodon maritimus Kunth, Nov. Gen. Sp. 1: 170. 1815 
[1816]. T-Protol.: Peru: Crescit locis planis apricis, 
arenosis regni Peruviani, inter Lima, Santa et Truxillo, 
Humboldt & Bonpland s.n. (HT: P]. 

NOTE: as Cynodon "maritimum". 

DST.: Arge, Braz, Chil, Para, Peru, Urug. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 
(1987); Caro, J. A. & E. Sanchez (1969); Caro, J. A. 
& E. Sanchez (1970); Santos, A. M. P. V. dos & S. 
de Castro Boechat (1994); Smith, L. B., D. C. 
Wasshausen & R. M. Klein (1981); Tovar, O. (1993). 

SYN.: Capriola dactylon var. maritima (Kunth) Hitchce.; 
Chloris maritimus (Kunth) Trin.; Cynodon dactylon 
var. maritimus (Kunth) Hack. 


Cynodon maritimus var. breviglumis Caro & E.A. Sanchez, 
Kurtziana 5: 234, f. 9A. 1969. T-Protol.: Argentina: 
Buenos Aires: Partido Monte: San Miquel del Monte, 
16 Feb 1938, Cabrera 4314 [HT: LP]. 

DST.: Arge. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Caro, J. A. & E. Sanchez (1969); 
Caro, J. A. & E. Sanchez (1970). 


62 Catalogue of New World Grasses: II. 


Cynodon maritimus var. grandispiculus Caro & E.A. 
Sanchez, Kurtziana 5: 234, f. 2A-C,E-F. 1969. T- 
Protol.: Argentina: Prov. Tucuman: Dpto. Capital: Rio 
Sali, 31 Jan 1924, Venturi 2737, T-Specim.: | of 1. 
Argentina: Tucuman: Capital Dept., 31 Jan 1924, S. 
Venturi 2737 {IT: BAA, BAB, BA, LP, LIL, US- 
1547122]. 

DST.: Arge, Braz, Chil, Para, Urug. 
LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Caro, J. A. & E. Sanchez (1969); 
Caro, J. A. & E. Sanchez (1970); Santos, A. M. P. V. 
dos & S. de Castro Boechat (1994); Smith, L. B., D. 
C. Wasshausen & R. M. Klein (1981). 

SYN.: Cynodon grandispiculus Caro & E.A. Sanchez. 


Cynodon maritimus Kunth var. maritimus. 

DST.: Arge, Boli, Chil, Para, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Caro, J. A. & E. Sanchez (1969); 
Caro, J. A. & E. Sanchez (1970); Santos, A. M. P. V. 
dos & S. de Castro Boechat (1994); Smith, L. B., D. 
C. Wasshausen & R. M. Klein (1981). 

SYN.: Cynodon dactylon var. maritimus (Kunth) Hack. 


Cynodon maritimus var. vaginiflorus Caro, Dominguezia 
6: 20. 1983. T-Protol.: USA: Pennsylvania: EE. UU:: 
Below Navy Yd., Philadelphia, 8 Nov 1865, / Burk 
s.n. (HT: PH]. 

DST.: Arge, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Caro, J. A. (1983); FNA GRAY- 
CARDS (); Hartman, R. L. & B. E. Nelson (1998); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994), 


Cynodon melicoides (P. Beauv.) Raspail, Ann. Sci. Nat., Bot. 
5: 303. 1825. Basionym: Bouteloua melicoides P. 
Beauv. 


= Bouteloua curtipendula (Michx.) Torr. 


Cynodon monostachyos (P. Beauv.) Raspail, Ann. Sci. Nat., 
Bot. 5: 303. 1825. Basionym: Campulosus 
monostachyus P. Beauv. 

NOTE: basionym erroneously attributed to Desv. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Cynodon mucronatus Caro & E.A. Sanchez, Kurtziana 5: 
239, f. 12. 1969, T-Protol.: Argentina: Misiones: Dpto. 
Montecarlo: Colonia Caraguatay, 19 Dec. 1943, Porta 
85 [HT: BAA]. 

DST.: Arge, Braz. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Caro, J. A. & E. Sanchez (1969); 
Santos, A. M. P. V. dos & S. de Castro Boechat 
(1994); Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 


Cynodon nitidus Caro & E.A. Sanchez, Darwiniana 17: 510, 

f. 1. 1972. T-Protol.: Chile: Dpto. Arica: Azapa, Aug 
1925, Wedermann 704 (HT: SI, IT: GH, LIL, NY, US- 
1498176], Chile: Prov. Tacna, dep. Arica, Azapa. 

DST.: Chil. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Caro, J. A. & E. Sanchez (1972); 

Marticorena, C. & M. Quezada (1985). 


Cynodon nlemfuensis Vanderyst, Bull. Agric. Congo Belge 
13: 342. 1922. T-Specim.: | of 1. Zaire, Vanderyst 
6095 [ST: BR]. 
NOTE: Cynodon lumfensis, nom. prov., in Bull. Agric. 
Congo Belge 11: 121 (1920). 
DST.: Arge, Beli, Boli, Braz, Cari, CoRi, ElSa, Hond, Mexi, 
USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Balick, M. J., M. H. Nee & D. E. 
Atha (2000); Beetle, A. A. (1987); Breedlove, D. E. 
(1986); Catasus Guerra, L. (1997); Clayton, W. D. & 
J. R. Harlan (1970); Davidse, G., M. Sousa S. & A. 
O. Chater (editores generales) (1994); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Filgueiras, T. S. (1991); Gibbs Russell, G. E, W. G. 
Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987); Killeen, T. J. (1990); 
Liogier, H. & L. Martorell (1982); McVaugh, R. 
(1983); Phillips, S. (1995); Pohl, R. W. (1980); Pohl, 
R. W. (1994); Renvoize, S. A. (1998); Santos, A. M. 
P. V. dos & S. de Castro Boechat (1994); Zuloaga, F. 
O., E. G. Nicora, Z. E. Rugolo de Agrasar, O. 
Morrone, J. Pensiero & A. M. Cialdella (1994). 
Cynodon dactylon var. sarmentosus Parodi; Cynodon 
parodii Caro & E.A. Sanchez. 


SYN.: 


Cynodon nlemfuensis Vanderyst var. nlemfuensis. 
DST.: Arge, Braz. 
LVL. ACCPTC.: 1; introduced. 
ALSO ACCPTD. BY: Clayton, W. D. & J. R. Harlan 
(1970); Phillips, S. (1995); Santos, A.M. P. V. dos & 
S. de Castro Boechat (1994). 
SYN.: Cynodon dactylon var. sarmentosus Parodi. 


Cynodon nlemfuensis var. robustus Clayton & J.R. Harlan, 
Kew Bull. 24: 189. 1970. T-Protol.: Kenya, Trapnell 
2110 [HT: K]. 
DST.: USA. 
LVL. ACCPTC.: |; introduced. 
ALSO ACCPTD. BY: Clayton, W. D. & J. R. 
(1970); Phillips, S. (1995). 


Harlan 


Cynodon occidentalis Willd. ex Steud., Nomencl. Bot. (ed. 2) 
1: 463. 1840. 
NOTE: nom. inval., as syn. of Cynodon dactylon (L.) Pers. 
= Cynodon dactylon (L.) Pers. 


Cynodon parodii Caro & E.A. Sanchez, Kurtziana 5: 193. 
1970. 
REPLACED NAME: Cynodon dactylon var. sarmentosus 
Parodi. 
= Cynodon nlemfuensis Vanderyst. 


Cynodon pascuus Nees, Fl. Bras. Enum. Pl. 2: 425. 1829, T- 
Specim.: | of 1. Brazil: Piauhy: Cachian et alibii, 
Martius s.n. (HAT: M; IT: US-878403]. 

= Cynodon dactylon (L.) Pers. 


Cynodon pedicellatus Caro, Dominguezia 6: 9, f. 2. 1983. 
T-Protol.: Uruguay: Depto. Montevideo: Sayago, 26 
Nov. 1959, Praderi 579 [HT: LIL-458958],. 
DST.: Urug. 
LVL. ACCPTC.: |. 
ALSO ACCPTD. BY: Caro, J. A. (1983). 


Catalogue of New World Grasses: II. 63 


Cynodon plectostachyus (K. Schum.) Pilg., Bot. Jahrb. Syst. 
40: 82. 1907. Basionym: Leptochloa plectostachya K. 
Schum. 

DST.: Arge, Cari, Mexi, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Beetle, A. A. (1987); Bor, N. L. (1960); 
Burkart, A. (1969); Caro, J. A. & E. Sanchez (1969); 
Catasus Guerra, L. (1997); Clayton, W. D. & J. R. 
Harlan (1970); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gibbs Russell, 
G. E, W. G. Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987); Molina R., A. (1975); 
Munz, P. A. & D. D. Keck (1959); Nicora, E. G. 
(1995); Phillips, S. (1995); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

Cynodon plectostachyus var. ruspolianus (Chiov.) 
Chiov.; Cynodon ruspolianus Chiov.; Leptochloa 
plectostachya K. Schum. 


SYN.: 


Cynodon plectostachyus var. ruspolianus (Chiov.) Chiov., 
Webbia 8: 111. 1951. Basionym: Cynodon ruspolianus 
Chiov. 

= Cynodon plectostachyus (K. Schum.) Pilg. 


Cynodon portoricensis Willd. ex Steud., Nomencl. Bot. (ed. 
2) 1: 463. 1940. 
NOTE: nom. inval., as syn. of Cynodon dactylon (L.) Pers. 
= Cynodon dactylon (L.) Pers. 


Cynodon procumbens (P. Durand) Raspail, Ann. Sci. Nat. 
(Paris) 5: 303. 1825. Basionym: Chloris procumbens 
P. Durand. 
= Bouteloua simplex Lag. 


Cynodon radiatus Roth, Syst. Veg. 2: 411. 1817. T-Protol.: 
In India orient., Heyne s.n. T-Specim.: | of 1. 1814, B. 
Heyne s.n.[T: L]. 

LVL. ACCPTC.:: 1. 


Cynodon ruspolianus Chiov., Annuario Reale Ist. Bot. Roma 
7: 70, t. 7. 1897. T-Specim.: | of 1. Ethiopia, Riva 248 
[HT: FI]. 
= Cynodon plectostachyus (K. Schum.) Pilg. 


Cynodon scabrifolius Caro, Dominguezia 6: 13, f. 3. 1983. 
T-Protol.: Brasil: Distrito Federal: arrededores de 
Brasilia, Apr 1968, N. Lima 232 [HT: US-2578849A]. 
T-Specim.: | of |. Brazil: Federal District, May 1968, 
N. Lima 232 (HT: US-2578849]. 

DST.: Braz. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Caro, J. A. (1983). 


Cynodon tenuis Trin. ex Spreng., Neue Entdeck. Pflanzenk. 
2: 63. 1821. T-Protol.: Hab. in America boreal. 
= Cynodon dactylon (L.) Pers. 


Cynodon transvaalensis Burtt Davy, Bull. Misc. Inform. 
Kew 1921: 281. 1921. T-Specim.: 1 of 1. South Africa, 
Dec 1919, .B. Davy 18156 [T: US-1295750 (fragm.)]. 

DST.: Ecua, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Anonymous (1986); Gibbs Russell, 
G. E, W. G. Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987); Hickman, J. C. (1993); 


Hitchcock, A. S. (1951); Jorgensen, P. M. & S. Leén- 
Yanez (eds.) (1999); Munz, P. A. & D. D. Keck 
(1959); Munz, P. A. (1974); Phillips, S. (1995). 


Cynodon umbellatus (Lam.) Caro, Dominguezia 6, f. 4: 16. 
1983. Basionym: Paspalum umbellatum Lam. 
= Cynodon dactylon (L.) Pers. 


Cynodon virgatus (L.) Raspail, Ann. Sci. Nat., Bot. 5; 302. 
1825. Basionym: Cynosurus virgatus L. 
= Leptochloa virgata (L.) P. Beauv. 


Cynodonteae trib. Dumort., Observ. Gramin. Belg. 
83, 90, 140. Jul-Sep 1824. TYPE: Cynodon Rich. 

NOTE: as "Cynodoneae". 

TRTD. BY: Soreng, R. J., G. Davidse, T. S. Filgueiras, E. J. 
Judziewicz, P. M. Peterson, and F. O. Zuloaga. 

NHT: Chloridoideae. 

CNSLTD. LIT.: Peterson, P. M. & Y. Herrera A., (1995); 
Peterson, P. M., R. D. Webster & J. Valdés-Reyna, 
(1997); Phillips, S. M., (1982); Hilu, K. W. & K. 
Wright, (1982); Swallen, J. R., (1939): Lazarides, M., 
(1972). 


Cynosurus aegyptius L., Sp. Pl. 72. 1753. T-Protol.: Habitat 
in Africa, Asia, America. T-Specim.: | of 4. Egypt, 
Anon. [ST: LINN-91.12]. 2 of 4. Asia [ST]. 3 of 4. 
America [ST]. 4 of 4. : van Royen s.n. [LT: L]. 

= Dactyloctenium aegyptium (L.) Willd. 


Cynosurus carolinianus Willd. ex Steud., Nomencl. Bot. (ed. 
2) 1: 465. 1840. 
NOTE: nom. nud. 
= Dactyloctenium aegyptium (L.) Willd. 


Cynosurus coracanus L., Syst. Nat. (ed. 10) 2: 875. 1759. T- 
Specim.: | of 1. Plukenet, Phytographia t. 91, f. 5 
(1691) [T]. 

NOTE: as Cynosurus "coracan”. 
= Eleusine coracana (L.) Gaertn. 


Cynosurus dactylon (L.) Pers., Syn. Pl. 1: 85. 
Basionym: Panicum dactylon L. 
= Cynodon dactylon (L.) Pers. 


Cynosurus distachyos Rottler ex Steud., Nomencl. Bot. (ed. 
2) 465. 1840. 
NOTE: as syn of Dactyloctenium aegyptiacum. 
= Dactyloctenium aegyptium (L.) Willd. 


Cynosurus domingensis Jacq., Misc. Austriac. 2: 363. 1781. 
T-Specim.: | of 1. Dominican Republic, Jacquin s.n. 
[HT]. 

= Leptochloa virgata (L.) P. Beauv. 


Cynosurus indicus L., Sp. Pl. 1: 72-73. 1753. T-Protol.: In 
Indiis. T-Specim.: 1 of 3. Burman, Thes. Zeylan. t. 47, 
f 1 (1737) [LT], LT designated by Phillips in Cafferty 
et al., Taxon 49(2): 249 (2000). 2 of 3. Sri Lanka: 
Central Province: Matale District: 5 mi S of Matale on 
Kandt Road, 1970, Clayton 5330 [ET: K; IET: PDA, 
US], ET designated by Phillips in Cafferty et al., 
Taxon 49(2): 249 (2000). 3 of 3. .van Royen s.n. [OM: 
L]. 

= Eleusine indica (L.) Gaertn. 


Cynosurus paspaloides Vahl, Symb. Bot. 2(27): 21. 1791. T- 
Specim.: 1 of 1. South Africa: Cape, Biillow s.n. [HT: 
C-Herb. Vahl]. 


64 Catalogue of New World Grasses: II. 


= Eustachys paspaloides (Vahl) Lanza & Mattei. 


Cynosurus retroflexus Vahl, Symb. Bot. 2: 20. 1791. T- 
Protol.: India, LaMark s.n. [HT: C]. 
= Dinebra retroflexa (Vahl) Panz. 


Cynosurus scoparius Lam., Encycl. 2: 188. 1786. T-Protol.: 
St. Dominique[HT: P]. T-Specim.: 1 of 1. Dominican 
Republic: Hispaniola I., no date, Crosnier s.n. [HT: P- 
LA; IT: US-2830901 (fragm. ex P-LA)]. 

= Chloris radiata (L.) Sw. 


Cynosurus secundus Pursh, Fl. Amer. Sept. 728. 1814. T- 
Protol.: USA: In upper Lousiana, Bradbury s.n. 
= Bouteloua curtipendula (Michx.) Torr. 


Cynosurus tenellus Cav. ex DC., Cat. Pl. Horti Monsp. 120. 
1813. 
NOTE: nom. inval., as syn. of Lamarckia tenella DC. 
= Aegopogon tenellus (DC.) Trin. 


Cynosurus tristachyos Lam., Encycl. 2: 188. 1786. T- 
Specim.: | of 1. Uruguay: Montevideo, Commerson 
sn. [T]. 

= Eleusine tristachya (Lam.) Lam. 


Cynosurus uniflorus Walter, Fl. Carol. 82. 1788. 
= Cynodon dactylon (L.) Pers. 


Cynosurus virgatus L., Syst. Nat. (ed. 10) 876. 1759. T- 
Specim.: | of 1. Jamaica, P. Browne s.n. (LT: LINN- 
91.18], LT designated by Hitchcock, Contr. U.S. Natl. 
Herb. 12: 122 (1908). 

= Leptochloa virgata (L.) P. Beauv. 


Czerniaevia rupestris Turez. ex Kom., Fl. URSS 2: 297. 
1934. 
NOTE: nom. inval., as syn. Pappophorum boreale Griseb. 
CORRECT NAME: Pappophorum boreale Griseb. 
= Enneapogon desvauxii P. Beauv. 


Dactilon Vill., Hist. Pl. Dauphine 2: 69. 1787. TYPE: 
Dactilon officinale Vill.; nom. illeg. superfl. Panicum 
dactylon L. 

NOTE: nom. rejic. against Cynodon. 
= Cynodon Rich. 


Dactilon officinale Vill., Hist. Pl. Dauphine 2: 69. 1787. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Panicum dactylon L. 
= Cynodon dactylon (L.) Pers. 


Dactylis cynosuroides L., Sp. Pl. 1: 71. 1753. T-Specim.: | of 
1. USA: Virginia, D. Clayton 577 [LT: BM; ILT: US- 
A865636 (fragm.)], LT designated by Hitchcock, 
Contr. U.S. Natl. Herb. 12: 121 (1908). 

= Spartina cynosuroides (L.) Roth. 


Dactylis fasciculata C. Sm., Tabl. Encycl. 1: 180. 1791. T- 
Specim.: | of 1. America, D. Richard s.n. (IT: US- 
A865637 (fragm. ex P)]. 

= Spartina alterniflora Loisel. 


Dactylis maritima Curtis, Enum. Brit. Grasses 46. 1787. 

NOTE: see notes. Notes from Index of Grass Species: 
Based on Dactylis cynosuroides as given in Hudson, 
Fl. Angl. ed 2. 43. 1778, the name misapplied to 
English species S. Stricta, the "cynosuroides" 
(misapplied) of Loefl. Portugal, the basis of 
"Lusitania" reference of Linnaeus. 

= Spartina maritima (Curtis) Fernald. 


Dactylis maritima Walter, Fl. Carol. 77. 1788. T-Protol.: 
South Carolina. 
NOTE: nom. illeg. hom.; not Curtis, 1787. 
E. NAME: Dactylis maritima Curtis. 
= Spartina alterniflora Loisel. 


Dactylis paspaloides Willd., Hort. Berol. 111. 1809. 
NOTE: nom. illeg. superfl. 
= Dinebra retroflexa var. retroflexa. 


Dactylis patens Aiton, Hort. Kew. |: 104. 1789. T-Protol.: 
North America. 
= Spartina patens (Aiton) Muhl. 


Dactyloctenium Willd, Enum. PI. 2: 1029. 1809. 


TYPE: Dactyloctenium aegyptium (L.) Willd.; LT 
designated by Hitchcock, U.S.D.A. Bull. (1915-23) 
772: 175 (1920). 

TRTD. BY: Peterson, P. M. 

NHT: Eleusininae. NO. ACCPTD. SPP.: 5. 

CNSLTD. LIT.: Fisher, B. S. & H. G. Schweickerdt, 
(1941). 


Dactyloctenium aegyptiacum Willd., Enum. PI. 2: 1029-1030. 
1809. 
NOTE: orth. var., see D. aegyptium. 
CORRECT NAME: Dactyloctenium aegyptium (L.) Willd. 
= Dactyloctenium aegyptium (L.) Willd. 


Dactyloctenium aegyptium (L.) Willd, Enum. Pl. 2: 1029. 
1809. Basionym: Cynosurus aegyptius L. 
NOTE: as Dactyloctenium “aegyptiacum", correctable to 
“aegyptium" as this a orth. var. of aegytium,and 
Willd. cites Cynosurus aegyptius as the basionym. 
DST.: Arge, Beli, Boli, Braz, Cari, Colo, CoRi, Ecua, ElSa, 
FrGu, Guat, Guya, Hond, Mexi, Nica, Pana, Para, 
Peru, Suri, USA, Urug, Vene. 
LVL. ACCPTC.: 1; introduced. 
ALSO ACCPTD. BY: Adams, C. D. (1972); Allem, A. C. 
& J. F.M. Valls (1987); Balick, M. J., M. H. Nee & 
D. E. Atha (2000); Beetle, A. A. (1977); Beetle, A. 
A. (1987); Berendsohn, W. G. & A. E. Araniva de 
Gonzalez (1991); Catasus Guerra, L. (1997); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Filgueiras, T. S. 
(1999); Gould, F. W (1979); Gould, F. W. & R. 
Moran (1981); Hickman, J. C. (1993); Hitchcock, A. 
S. (1909); Jorgensen, P. M. & S. Leén-Yanez (eds.) 
(1999); Judziewicz, E. J. (1990); Killeen, T. J. 
(1990); Lawesson, J. E., H. Adsersen & P. Bentley 
(1987); McClure, F. A. (1955); Phillips, S. (1995); 
Pohl, R. W. (1994); Porter, D. M. (1983); Pringle, J. 
S. (1985); Renvoize, S. A. (1984); Renvoize, S. A. 
(1998); Smith, L. B., D. C. Wasshausen & R. M. 
Klein (1981); Tovar, O. (1993); Wiggins, I. L. & D. 
M. Porter (1971); Wunderlin, R. P. (1998); Zuloaga, 
F. O., E. G. Nicora, Z. E. Rugolo de Agrasar, O. 
Morrone, J. Pensiero & A. M. Cialdella (1994). 
Aegilops saccharinum Walter, Chloris mucronata 
Michx.; Cynosurus  aegyptius L.;  Cynosurus 
carolinianus Willd. ex Steud.; Cynosurus distachyos 
Rottler ex Steud.; Dactyloctenium aegyptiacum 
Willd.; Dactyloctenium aegyptium (L.) P. Beauv.; 
Dactyloctenium aegyptium (L.) Richt.; 
Dactyloctenium aegyptium to. viviparum Beetle; 
Dactyloctenium aegyptius var. mucronatum (Michx.) 


SYN:: 


Catalogue of New World Grasses: II. 65 


Lanza & Mattei; Dactyloctenium distachyum Trin., 
Dactyloctenium meridionale Ham.; Dactyloctenium 
mucronatum  (Michx.)  Willd.; © Dactyloctenium 
mucronatum var. erectum E. Fourn.; Eleusine 
aegyptiaca (L.) Desf.; Eleusine aegyptia (L.) Desf.; 
Eleusine aegyptia (L.) Pers.; Eleusine aegyptia (L.) 
Roberty; Eleusine aegyptia (L.) Roxb.; Eleusine 
aegyptia Raf.; Eleusine cruciata Elliott; Eleusine 
cruciata Lam.; Eleusine egyptia Raf.; Eleusine 
mucronata (Michx.) Hornem.; Eleusine mucronata 
Stokes; Eleusine pectinata Moench. 


Dactyloctenium aegyptium (L.) P. Beauv., Ess. Agrostogr. 
15, 72, 159, t. 15, f. 2. 1812. Basionym: Cynosurus 
aegyptius L. 

NOTE: isonym. 
E. NAME: Dactyloctenium aegyptium (L.) Willd. 
= Dactyloctenium aegyptium (L.) Willd. 


Dactyloctenium aegyptium (L.) Richt., Pl. Eur. 1: 68. 1870. 
Basionym: Cynosurus aegyptius L. 
NOTE: hom. illeg. 
E. NAME: Dactyloctenium aegyptium (L.) Willd. 
= Dactyloctenium aegyptium (L.) Willd. 


Dactyloctenium aegyptium var. aristatum (Link) A. Chev., 
Rev. Int. Bot. Appl. Agric. Trop. 14(150): 130. 1934. 
Basionym: Dactyloctenium aristatum Link. 

= Dactyloctenium aristatum Link. 


Dactyloctenium aegyptium fo. viviparum Beetle, Phytologia 
48(2): 190. 1981. T-Protol.: Mexico. T-Specim.: | of 
1. Mexico; Sonora: Hermosillo, edge of irrigation 
ditch, 19 Oct 1980, B. Bracamonte 31 [HT: COCA]. 
= Dactyloctenium aegyptium (L.) Willd. 


Dactyloctenium aegyptius var. mucronatum (Michx.) Lanza 
& Mattei, Boll. Reale Orto Bot. Giardino Colon. 
Palermo 9: 58. 1910. Basionym: Chloris mucronata 
Michx. 

= Dactyloctenium aegyptium (L.) Willd. 


Dactyloctenium aristatum Link, Hort. Berol. 1: 59. 1827. T- 
Protol.: Hab. in Aegypto, Eherenb. s.n. 

DST.: Mexi. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Beetle, A. A. (1987); Bor, N. L. 
(1960); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Phillips, S. (1995). 

SYN.: Dactyloctenium aegyptium var. aristatum (Link) A. 
Chev. 


Dactyloctenium distachyum Trin., Fund. Agrost. 140. 1820. 

NOTE: nom. nud., based on unpublished Cynosurus 
distachyos Rottl., later published as C. distachyos 
Rottl. ex Steud. which equals D. aegyptium (L.) 
Willd. 

= Dactyloctenium aegyptium (L.) Willd. 


Dactyloctenium geminatum Hack., Bull. Herb. Boissier 
7(1): 26. 1899. T-Protol.: Delagoa Bay, in arenosis, 
Junod s.n, T-Specim.: | of 1. Mozambique, 1890, 
Junod 231 [IT: US-81794]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Beetle, A. A. (1987); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Gibbs Russell, G. E, W. G. Welman, E. Reitief, K. L. 
Immelman, G. Germishuizen, B. J. Pienaar, M. wan 


Wyk, and A. Nicholas (PRECIS) (1987); Phillips, S. 
(1995). 


Dactyloctenium glaucophyllum Courbai, Ann. Sci. Nat., Bot., 
ser. 4, 18: 133. 1862. T-Protol.: In locis  siccis, 
sabulosis. L'ile de Dissee (Mer Rouge). 

= Dactyloctenium scindicum Boiss. 


Dactyloctenium meridionale Ham., Prodr. Pl. Ind. Occid. 6. 
1825. T-Protol.: Habitat vulgo in pascuis Insularum 
Carribearum, et passim in America calidiore. 

NOTE: nom. illeg. superfl. 
CORRECT NAME: Cynosurus aegyptius L. 
= Dactyloctenium aegyptium (L.) Willd. 


Dactyloctenium mucronatum (Michx.) Willd., Enum. PI. 
1029. 1809. Basionym: Chloris mucronata Michx. 
= Dactyloctenium aegyptium (L.) Willd. 


Dactyloctenium mucronatum var. erectum E. Fourn., Mexic. 
Pl. 2: 144. 1886. T-Specim.: | of 3. Mexico: Veracruz, 
20 1865, Gouin 68 [ST: US-A865633 (fragm. ex P)]. 2 
of 3. Mexico: Colipa, Karwinsky 989b [ST]. 3 of 3. 
Mexico: Hacienda el Coxo, Karwinsky 989 [ST]. 
= Dactyloctenium aegyptium (L.) Willd. 


(R. Br.) P. Beauv., Ess. 
1812. Basionym: Eleusine 


Dactyloctenium radulans 
Agrostogr. 72, 160. 
radulans R. Br. 

DST.: USA. 
LVL. ACCPTC.: 1; introduced. 


Dactyloctenium scindicum Boiss., Diagn. PI. Orient., ser. 2, 


4: 131. 1859. T-Protol.: Hab. in ditione Scinde, 
Griffith 637. 
DST.: USA. 


LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Beetle, A. A. (1987); Bor, N. L. 
(1960); Phillips, S. (1995). 

SYN.: Dactyloctenium glaucophyllum Courbai; Eleusine 
glaucophylla (Courbai) Munro ex Benth.; Eleusine 
scindica (Boiss.) Duthie. 


Dactylogramma Link, Hort. Berol. 2: 248. 1833. TYPE: 
Dactylogramma cinnoides Link. 
= Muhlenbergia Schreb. 


Dactylogramma cinnoides Link, Hort. Berol. 2: 248. 1833. T- 
Protol.; Cult. in Berlin: Hab. in America septentrionali 
occidentali, seed from Richardson. 

= Muhlenbergia glomerata (Willd.) Trin. 


Dasyochloa Willd. ex Rydb., Agric. Exp. Sta. Agric. 
Coll. Colorado Bull, 100: 18, 37. 1906. TYPE: 
Dasyochloa pulchella (Kunth) Willd. ex Rydb. 

NOTE: Alt. title, Fl. Colorado. 

TRTD. BY: Valdés-Reyna, J. 

NHT: Munroinae. NO. ACCPTD. SPP.: 1. 

CNSLTD. LIT.: Valdés-Reyna, J. & S. L. Hatch, (1995); 
Caro, J. A., (1981). 

SYN.: Dasyochloa Willd. ex Steud. 


Dasyochloa argentina (Kuntze) Caro, Dominguezia 2: 4. 
1981. Basionym: Sieglingia argentina Kuntze. 
= Erioneuron pilosum var. longearistatum (Kurtz) Anton. 


Dasyochloa argentina (Kuntze) Caro var. argentina. 
= Erioneuron pilosum var. longearistatum (Kurtz) Anton. 


66 Catalogue of New World Grasses: II. 


Dasyochloa argentina var. aristiglumis Caro, Dominguezia 
2: 4, f. 1. 1981. T-Protol.: Argentina: Cordoba: Dpto. 
Punilla: Capilla del Monte, 1 Mar 1941, Nicora s.n. 
[HT: SI-18502]. T-Specim.: | of 1. Argentina: Prov. 
Cordoba, Capilla del Monte, | Mar 1941, £. Nicora 
s.n. (HT: SI-18502]. 

= Erioneuron pilosum var. longearistatum (Kurtz) Anton. 


Dasyochloa argentina var. parodiana (E.A. Sanchez) Caro, 
Dominguezia 2: 7, f. 2. 1981. Basionym: Erioneuron 
pilosum var. parodianum E.A. Sanchez. 

= Erioneuron pilosum var. longearistatum (Kurtz) Anton. 


Dasyochloa avenacea (Kunth) Willd. ex Steud., Nomencl. 
Bot. (ed. 2) 1: 484. 1840. Basionym: Triodia avenacea 
Kunth. 

= Erioneuron avenaceum (Kunth) Tateoka. 


Dasyochloa kurtziana (Parodi) Caro, Dominguezia 2: 7, f. 3. 
1981. Basionym: Tridens avenaceus var. kurtziana 
Parodi. 

= Erioneuron avenaceum 
Anton. 


var. kurtzianum (Parodi) 


Dasyochloa longiglumis (Parodi) Caro, Dominguezia 2: 8, f. 
4. 1981. Basionym: Tridens  avenaceus — var. 
longiglumis Parodi. 

= Erioneuron avenaceum var. longiglume (Parodi) Anton, 


Dasyochloa longiglumis var.cabrerae Caro, Dominguezia 2: 
13. 1981. T-Protol.: Argentina: La Rioja: Dpto. 
Famatima: Quebrada de la Aguadita, Cabrera 18096, 
pro parte |HT: LP], type mixed with Dasyochloa 
longiglumis var. longiglumis. 

= Erioneuron avenaceum var. longiglume (Parodi) Anton. 


Dasyochloa longiglumis (Parodi) Caro var. longiglumis. 
= Erioneuron avenaceum var. longiglume (Parodi) Anton. 


Dasyochloa pulchella Willd. ex Steud., Nomencl. Bot. (ed. 2) 
1: 484. 1840. 
NOTE: nom. inval., genus not yet validly published; based 
on Triodia pulchella Kunth. 
= Dasyochloa pulchella (Kunth) Willd. ex Rydb. 


Dasyochloa pulchella (Kunth) Willd. ex Rydb., Agric. Exp. 
Sta. Agric. Coll. Colorado Bull. 100: 18, 37. 1906. 
Basionym: Triodia pulchella Kunth. 

NOTE: Alt. title, Fl. Colorado. 

DST.: Mexi, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Valdés-Reyna, 
J. & S. L. Hatch (1997). 

SYN.: Dasyochloa pulchella Willd. ex Steud.; Erioneuron 
pulchellum (Kunth) Tateoka; Koeleria pulchella 
(Kunth) Spreng.; Sieglingia pulchella (Kunth) 
Kuntze; Sieglingia pulchella var. parviflora Vasey ex 
Beal; Trichodiclida_ prolifera Cerv.;  Tricuspis 
pulchella (Kunth) Torr.; Tridens pulchellus (Kunth) 
Hitche.; Triodia pulchella Kunth; Uralepis pulchella 
(Kunth) Kunth. 


Dasyochloa pygmaea (Hack.) Caro, Dominguezia 2: 8. 1981. 


Basionym: Triodia avenacea var. pygmaea Hack. 
= Erioneuron avenaceum var. pygmaeum (Hack.) Anton. 


Davyella Hack., Oesterr. Bot. Z. 49: 134. 1899. 
NOTE: nom. superfl. pro Neostapfia. 
= Neostapfia Burtt Davy. 


Davyella colusana (Burtt Davy) Hack., Oesterr. Bot. Z. 49: 
134. May 1899. Basionym: Stapfia colusana Burtt 
Davy. 

= Neostapfia colusana (Burtt Davy) Burtt Davy. 


Diachroa Nutt., Trans. Amer. Philos. Soc., n.s., 5: 147. 1835. 
TYPE: Diachroa procumbens (Muhl.) Nutt. 
= Leptochloa P. Beauv. 


Diachroa procumbens (Muhl.) Nutt., Trans. Amer. Philos. 
Soc., ms. 5: 147. 1837. Basionym: Festuca 
procumbens Muhl. 

= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Diachyrium Griseb., Abh. Konigl. Ges. Wiss. Gottingen 19: 
257. 1874. TYPE: Diachyrium arundinaceum Griseb. 
= Sporobolus R. Br. 


Diachyrium arundinaceum Griseb., Abh. Konigl. Ges. Wiss. 
Gottingen 19: 257, t. 2, f. 8. 1874. T-Protol.: 
Argentina: Catamarca: "in salsis Laguna Blanca, alt. 
10,000'.", [Lorentz] s.n. T-Specim.: | of 1. Argentina: 
Catamarca: Laguna Blanca, Lorentz 440 [IT: US- 
A865784 (fragm.)]. 

= Sporobolus rigens (Trin.) Desv. 


Diachyrium rigens (Trin.) Mez, Repert. Spec. Nov. Regni 
Veg. 17(19-30): 300. 1921. Basionym: Vilfa rigens 
Trin. 

= Sporobolus rigens (Trin.) Desv. 


Diacisperma Kuntze, Lex. Gen. Phan. 169. 1903. 
NOTE: nom. illeg. superfl. 
= Leptochloa P. Beauv. 


Diandrochloa De Winter, Bothalia 7: 387. 1960. TYPE: 
Diandrochloa namaquensis (Nees ex Schrad.) De 
Winter. 

= Eragrostis Wolf. 


Diandrochloa diplachnoides (Steud.) A.N. Henry, Bull. Bot. 
Surv. India 9: 290. 1968. Basionym: Eragrostis 
diplachnoides Steud. 

= Eragrostis japonica (Thunb.) Trin. 


Diandrochloa japonica (Yhunb.) A.N. Henry, Bull. Bot. 
Surv. India 9: 290. 1968. Basionym: Poa japonica 
Thunb. 

= Eragrostis japonica (Thunb.) Trin. 


Diandrochloa namaquensis (Nees ex Schrad.) De Winter, 
Bothalia 7: 388. 1960. Basionym: Eragrostis 
namaquensis Nees ex Schrad. 

= Eragrostis japonica (Thunb.) Trin. 


Digitaria dactylon (L..) Scop., Fl. Carniol. (ed. 2) 1: 52. 1772. 
Basionym: Panicum dactylon L. 
= Cynodon dactylon (L.) Pers. 


Digitaria glumaepatula (Steud.) Miq., Fl. Ned. Ind. 3: 439. 
1857. Basionym: Panicum glumaepatulum Steud. 
= Cynodon dactylon (L.) Pers. 


Digitaria littoralis Salisb., Prodr. Stirp. Chap. Allerton 19. 
1796. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Panicum dactylon L. 
= Cynodon dactylon (L.) Pers. 


Digitaria malabarica (L.) Roem. & Schult. Syst. Veg. 2: 
474. 1817. Basionym: Poa malabarica L. 


Catalogue of New World Grasses: II. 67 


NOTE: nom. rej., basionym rej. [ICBN art. 56.2]; see note 
Index of Grass Species page 583. 
= Leptochloa fusea (L.) Kunth. 


Digitaria stolonifera Schrad., Fl. Germ. 1: 165, pl. 3, f. 9. 
1806. 
NOTE: nom. illeg. superfl. 
E. NAME: Digitaria dactylon (L.) Scop. 
CORRECT NAME: Panicum dactylon L. 
= Cynodon dactylon (L.) Pers. 


Dilepyrum Michx., Fl. Bor.-Amer. 1: 40. 1803. TYPE: 
Dilepyrum minutiflorum Michx. 


= Muhlenbergia Schreb. 


Dilepyrum diffusum P. Beauv., Ess. Agrostogr. 160. 1812. 
NOTE: nom. nud.; not accepted by the author, referred to 
Muhlenbergia, probably an error for Muhlenbergia 
diffusa Willd. 
= Muhlenbergia schreberi J.F. Gmel. 


Dilepyrum minutiflorum Michx., Fl. Bor.-Amer. 1: 40. 1803. 
T-Protol.: USA: "Hab. in apricis, pratensibus regionum 
Kentucky et Illinoensium.". T-Specim.: 1 of 1. USA: 
In apricis, pratis Kentucky, Illinoensium pabulum 
parestantissimum, Michaux s.n. [HT: P]. 

= Muhlenbergia schreberi J.F. Gmel. 


Dineba bromoides Kunth, Nov. Gen. Sp. 1: 172, t. 51. 1815 
[1816]. T-Protol.: Mexico: Crescit in  temperatis 
scopulosis aridis inter Guanaxuato et Cubilete 
Mexicanorum", Humboldt & Bonpland s.n. 

NOTE: orth. var. 
CORRECT NAME: Dinebra bromoides Kunth. 
= Bouteloua radicosa (E. Fourn.) Griffiths. 


Dineba hirsuta J. Presl, Relig. Haenk. 1(4-5): 292. 1830. T- 
Protol.: Peru, Haenke s.n. [HT: PR]. 
NOTE: orth. var. 
CORRECT NAME: Dinebra hirsuta J. Pres]. 
= Bouteloua aristidoides (Kunth) Griseb. 


Dinebra Jacg., Fragm. Bot. 77. 1809. TYPE: Dinebra 
arabica Jacq. 
TRTD. BY: Peterson, P.M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 1 SUBSP.: 1. 
CNSLTD. LIT.: Phillips, S. M., (1973). 


Dinebra aegyptiaca Delile, Descr. Egypte, Hist. Nat. 26, t. 
11, f. 3. 1813. T-Specim.: | of 1. Egypt, Damiette, 
Delile s.n. (HT: MPU}. 

NOTE: as "Dinaeba", nom. illeg. superfl. 
= Dinebra retroflexa var. retroflexa. 


Dinebra americana (L.) P. Beauv., Ess. Agrostogr. 98, 160, t. 
16, f. 1-3. 1812. Basionym: Aristida americana L. 
NOTE: as "Dineba" americana. 
= Bouteloua americana (L.) Scribn. 


Dinebra arabica Jacq., Fragm. Bot. 77, 98, t. 121, f. 1. 1809. 
T-Specim.: | of 1. cultivated at the Vienna Botanic 
Garden [HT: W]. 

= Dinebra retroflexa var. retroflexa. 


Dinebra aristidoides Kunth, Nov. Gen. Sp. 1: 171. 1815 
[1816]. T-Protol.: Crescit in asperis frigidis convallis 
Tolucensis, alt. 1320 hexap. (regno Mexicano), floret 
Septembri, Humboldt & Bonpland s.n. T-Specim.: | of 


1. Mexico: near Nevada Toluca, Humboldt & 
Bonpland 67 (HT: P; IT: US (fragm.)]. 


= Bouteloua aristidoides (Kunth) Griseb. 


Dinebra brevifolia Steud., Syn. Pl. Glumac. 1: 299. 1854. T- 
Specim.: | of 1. Ethiopia, near Schire Dschogarti, 22 
Nov 1839, Schimper 527 [HT: P; IT: K]. 
= Dinebra retroflexa var. retroflexa. 


Dinebra bromoides Kunth, Nov. Gen. Sp. 1: 172, t. 51. 1815 
[1816]. T-Specim.: 1 of 1. Mexico: Guanajuato, 
Humboldt & Bonpland s.n. [HT: P; IT: US-865871 
(fragm. ex P, photo)]. 

NOTE: non Bouteloua bromoides Lag. (1816). 
= Bouteloua radicosa (E. Fourn.) Griffiths. 


Dinebra chloridea J. Presl, Relig. Haenk. 1(4-5): 291. 1830. 
T-Protol.: Hab. in Mexico, Haenke s.n, T-Specim.: | 
of 1. Mexico, Haenke s.n. [HT: PR; IT; MO]. 

NOTE: as "Dineba" chloridea. 
= Enteropogon chlorideus (J. Pres!) Clayton. 


Dinebra chondrosioides Kunth, Nov. Gen. Sp. 1: 173, t. 53. 
1815 [1816]. T-Specim.: 1 of 1. Mexico: Michoacan: 
Valladolid, Humboldt & Bonpland s.n. (HT: P; IT: LE, 
US (fragm.)]. 

= Bouteloua chondrosioides (Kunth) Benth. ex S. Watson. 


Dinebra cristata J. Presl, Relig. Haenk. 1(4-5): 293. 1830. T- 
Specim.: | of 1. Mexico, Haenke s.n. [HT: PR; IT: 
US-2307999 (fragm.), US photo], fragm. probably 
from PR. 

NOTE: as "Dineba" cristata. 
= Bouteloua triaena (Trin. ex Spreng.) Scribn. 


Dinebra curtipendula (Michx.) P. Beauv., Ess. Agrostogr. 98, 
158, 160. 1812. Basionym: Chloris curtipendula 
Michx. 

NOTE: as "Dineba" curtipendula. 
= Bouteloua curtipendula (Michx.) Torr. 


Dinebra hirsuta J. Presl, Relig. Haenk. 1(4-5): 292. 1830. T- 
Protol.: Hab. in Peruviae, Haenke s.n. T-Specim.: | of 
1. Peru, Haenke s.n. {HT: PR; IT: US (fragm.)]. 
NOTE: as "Dineba" hirsuta. 
= Bouteloua aristidoides (Kunth) Griseb. 


Dinebra juncifolia (Desv.) Steud., Nomencl. Bot. (ed. 2) 1: 
510. 1840. Basionym: Heterosteca juncifolia Desv. 
NOTE: as "Dineba" juncifolia. 
= Bouteloua repens (Kunth) Scribn. 


Dinebra melicoides P. Beauv., Ess. Agrostogr. 160. 1812. 
NOTE: nom. nud.; as "Dineba" melicoides. 
= Bouteloua curtipendula (Michx.) Torr. 


Dinebra paspaloides (Willd.) P. Beauv., Ess. Agrostogr. 98, 
160. 1812. Basionym: Dactylis paspaloides Willd. 
NOTE: as "Dinaeba" paspaloides. 
= Dinebra retroflexa var. retroflexa. 


Dinebra repens Kunth, Nov. Gen. Sp. 1: 172, t. 52. 1815 
[1816]. T-Protol.: Crecit ad litora Ocwani Pacifici 
prope Acapulco, Humboldt & Bonpland s.n._ Y- 
Specim.: 1 of 1. Mexico: Guerrero: Acapulco, 
Humboldt & Bonpland s.n. [LT: P], LT designated 
by?; cited by McVaugh, Fl. Novo-Galic. 14: 91 
(1983). 

= Bouteloua repens (Kunth) Scribn. 


68 Catalogue of New World Grasses: II. 


Dinebra retroflexa (Vahl) Panz., Ideen Revis. Gras. 59-60, 


20, t. Hl, f. 1, 2 a-f. 1814. Basionym: Cynosurus 
retroflexus Vahl. 
NOTE: also, Denkschr. Kaiserl. Akad. Wiss., Math.- 


Naturwiss. Kl. 4: 309-310, 270 pl. 12 (1814). 

DST.: USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Bor, N. L. (1960); Phillips, S. 
(1995); Phillips, S. M. (1973). 

SYN.: Cynosurus retroflexus Vahl; Dinebra arabica Jacq.; 
Eleusine calycina Roxb.; Eleusine calycina Roxb.; 
Leptochloa arabica (Jacq.) Kunth. 


Dinebra retroflexa var. brevifolia (Steud.) T. Durand & 
Schinz, Consp. Fl. Afr. 5: 865. 1894. Basionym: 
Dinebra brevifolia Steud. 

= Dinebra retroflexa var. retroflexa. 


Dinebra retroflexa var. retroflexa. 

DST.: USA. 

LVL. ACCPTC.:: 1; introduced. 

ALSO ACCPTD. BY: Phillips, S. (1995); Phillips, S. M. 
(1973). 

Cynosurus retroflexus Wahl; Dactylis paspaloides 
Willd.; Dinebra aegyptiaca Delile; Dinebra arabica 
Jacq.; Dinebra — brevifolia Steud.; | Dinebra 
paspaloides (Willd.) P. Beauv.; Dinebra retroflexa 
var. brevifolia (Steud.) T. Durand & Schinz; Eleusine 
calycina Roxb.; Leptochloa arabica (Jacq.) Kunth; 
Leptochloa calycina (Roxb.) Kunth. 


SYN.: 


Dinebra secunda (Pursh) Roem. & Schult., Syst. Veg. 2: 711. 
1817. Basionym: Cynosurus secundus Pursh. 
NOTE: as "Dineba" secunda. 
= Bouteloua curtipendula (Michx.) Torr. 


Diplachne P. Beauv., Ess. Agrostogr. 80. 1812. TYPE: 
Diplachne fascicularis(Lam.) P. Beauv. 
= Leptochloa P. Beauv. 


Diplachne acuminata Nash, Man. Fl. N. States 128. 1901. T- 
Specim.: 1 of 1. USA: Kansas, Pottawatomie Co. 
Manhattan, 19 Jul 1892, C.H. Thompson s.n. [HT: NY- 
19505; IT: US-865882 (fragm.)]. 

= Leptochloa fusea subsp. fascicularis (Lam.) N. Snow. 


Diplachne barbata Hack., Oesterr. Bot. Z. 52: 240. 1902. T- 
Protol.: Brasilia: Pernambuco: Boa Viagem, in 
arenosis, Schenck 43/0 [HT: W]. T-Specim.: 1 of 1. 
Brazil: Pernambuco: Boa Viagem, in arenosis, /.H.R. 
Schenck 4310 (HT: W; IT: US-78813 (fragm.)]. 

= Gouinia barbata (Hack.) Swallen. 


Diplachne brandegei Vasey, Proc. Calif. Acad. Sci., ser, 2, 2: 
213. 1889, T-Specim.: | of 1. Mexico: Baja California 
Sur: Magdalena Island, 18 Jan 1889, 7.8. Brandegee 
/1 {IT: US-78812]. 
= Enteropogon brandegei (Vasey) Clayton. 


Diplachne carinata (Griseb.) Hack., Bol. Acad. Ci. 
(Cordoba) 16: 253. 1900. Basionym: Atropis carinata 
Griseb. 

= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Diplachne cearensis Ekman, Ark. Bot. 10(17): 32, t. 5, f. 3, t. 
6, f. 18. 1911. T-Protol.: Brazil: Ceara: in_ silvis 
caatinga dictis viam entre Crato et Barbalha, 23 Apr 
1810, Loefgren 672 [HT:S]. T-Specim.: | of 1. Brazil: 


Ceara: Ent. Crato & Barbalha, A.Leofgren 672 [T: IT: 
US-865879 (fragm.)]. 
= Gouinia cearensis (Ekman) Swallen. 


Diplachne chloridiformis Hack. ex Stuck., Anales Mus. Nac. 
Hist. Nat. Buenos Aires 13: 498. 1906. T-Protol.: 
Argentina: Cordoba: Rio Seco, Dep. Rio Seco, Feb 
1897, remitido por D. Napoleon Saravia, Stuckert 
Herb. Arg. 2329a. T-Specim.: | of 2. Argentina: 
Cérdoba: Rio Seco, Feb 1897, Stuckert 2329a [HT: W; 
IT: BAA-907 (fragm.)]. 2 of 2. Argentina: Cordoba: 
Rio Seco, 1897, TV. Stuckert 2329 [LT: ILT: US- 
865878 (fragm.)], Spec. annotated by N. Snow as a 
fragm. of a Lectotype. 

= Leptochloa chloridiformis (Hack. ex Stuck.) Parodi. 


Diplachne domingensis (Jacq.) Chapm., Fl. South. U.S. (ed. 
3) 609. 1897, Basionym: Cynosurus domingensis Jacq. 
= Leptochloa virgata (L.) P. Beauv. 


Diplachne dubia (Kunth) Scribn., Bull. Torrey Bot. Club 
10(1): 30. 1883. Basionym: Chloris dubia Kunth. 
= Leptochloa dubia (Kunth) Nees. 


Diplachne dubia var. aristata Vasey, Proc. Calif. Acad. Sci., 
ser. 2, 2: 213. 1889. 
NOTE: nom. nud. 
= Leptochloa dubia (Kunth) Nees. 


Diplachne dubia (Kunth) Scribn. var. dubia. 
= Leptochloa dubia (Kunth) Nees. 


Diplachne dubia var. humboldtiana Kuntze, Revis. Gen. Pl. 
3(2): 349. 1898. 
= Leptochloa dubia (Kunth) Nees. 


Diplachne dubia var. kurtziana Kuntze, Revis. Gen. Pl. 3(3): 
349. 1898. T-Specim.: | of 1. Argentina: Cordoba, 
EB. Kurtz 6647 [T: US-86574}. 
= Leptochloa dubia (Kunth) Nees. 


Diplachne dubia var. pringleana Kuntze, Revis. Gen. PI. 
3(2): 349. 1898. T-Specim.: 1 of 1. Mexico: 
Chihuahua: hills and plains near Chihuahua, Aug 
1885, C.G. Pringle 422 [LT: US-899043; ILT: GH, 
NY-19498, NY-19499, NY-19500, P, RAS], LT 
designated by Snow. 

= Leptochloa dubia (Kunth) Nees. 


Diplachne fascicularis (Lam.) P. Beauv., Ess. Agrostogr. 81, 
160, t. 16, f. 9. 1812. Basionym: Festuca fascicularis 
Lam. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Diplachne fusca (L.) P. Beauv. ex Roem. & Schult., Syst. 
Veg. 2: 615. 1817. Basionym: Festuca fusca L. 
NOTE: Ess. Agrost. 80, 163. 1812. 
= Leptochloa fusca (L.) Kunth. 


Diplachne fusca (L.) P. Beauv. ex Stapf, Fl. Cap. 7: 591. 
1900. Basionym: Festuca fusca L. 
NOTE: nom. illeg. hom. 
E. NAME: Diplachne fusca (L.) P. Beauv. ex Roem, & 
Schult. 
= Leptochloa fusea (L.) Kunth. 


Diplachne fusca (L.) P. Beauv. ex Stuck., Anales Mus. Nac. 
Buenos Aires 11: 128. 1904. Basionym: Festuca fusca 
L. 
NOTE: nom. illeg. hom. 


Catalogue of New World Grasses: II. 69 


E. NAME: Diplachne fusca (L.) P. Beauv. ex Roem. & 
Schult. 
= Leptochloa fusca (L.) Kunth. 


Diplachne guatemalensis Hack., Oesterr. Bot. Z. 52: 275. 
1902. T-Protol.: Guatemala, Friedrichstahl 1748 (HT: 
W]. T-Specim.: 1 of 1. Guatemala, E.R. von 
Friedrichsthal 1748 [T: US-78815]. 
= Gouinia latifolia var. guatemalensis (Hack.) J.J. Ortiz. 


Diplachne halei Nash, New York Bot. Gard. 1: 292, 1889. T- 
Specim.: | of 2. USA: Louisiana, 188-, J. Hale s.n. 
[HT: NY-19504]. 2 of 2, USA: Louisiana, 188-, J. 
Hale s.n. (ST: US-78814]. 
= Leptochloa panicoides (J. Presl) Hitchc. 


Diplachne imbricata (Thurb.) Scribn., Bull. Torrey Bot. Club 
10(1):; 30. 1883. Basionym: Leptochloa_ imbricata 
Thurb. 

= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Diplachne latifolia (Griseb.) Hack., Oesterr. Bot. Z. 52: 274. 
1902. Basionym: Tricuspis latifolia Griseb. 
= Gouinia latifolia (Griseb.) Vasey. 


Diplachne malabarica (L.) Merr., Bull. Torrey Bot. Club 60: 
635. 1933. Basionym: Poa malabarica L. 
NOTE: nom. rej., basionym rej. [ICBN art. 56.2]. 
= Leptochloa fusca (L.) Kunth. 


Diplachne maritima E.P. Bicknell, Bull. Torrey Bot. Club 
35(4): 195. 1908. 
REPLACED NAME: Festuca procumbens Muhl. 
BLOCKING NAME: Diplachne procumbens Arechav. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Diplachne mendocina (Phil.) Kurtz, Bol. Acad. Ci. (Cordoba) 
15: 521. 1897. Basionym: /pnum mendocinum Phil. 
= Leptochloa dubia (Kunth) Nees. 


Diplachne mexicana (Scribn.) Hack., Oesterr. Bot. Z. 52: 
275. 1902. Basionym: Leptochloa mexicana Scribn. 
= Gouinia mexicana (Scribn.) Vasey. 


Diplachne monticola (Chase) McNeill, Brittonia 31(3): 401. 
1979. Basionym: Leptochloa monticola Chase. 
= Gouinia latifolia (Griseb.) Vasey. 


Diplachne panicoides (J. Presl) McNeill, Brittonia 31(3): 
402. 1979. Basionym: Megastachya panicoides J. 
Presl. 

= Leptochloa panicoides (J. Pres!) Hitche. 


Diplachne patens (J. Presl) E. Desv., Fl. Chil. 6: 371. 1854. 
Basionym: Schismus patens J. Presl. 
= Leptochloa dubia (Kunth) Nees. 


Diplachne patens E. Fourn. ex Hemsl., Biol. Cent.-Amer., 
Bot. 3: 570. 1885. 
NOTE: nom. nud. 
CORRECT NAME: Diplachne patens E. Fourn. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Diplachne patens E. Fourn., Mexic. PI. 2: 148. 1886. T- 
Protol.: Mexico: Vera Cruz, Gouin 93 [HT: P]. T- 
Specim.: 1 of 1. Mexico: Vera Cruz, 28 Jul 1865, 
Gouin 93 [HT: P; US-2875408b (fragm. ex P)]. 

NOTE: hom. illeg. 
E. NAME: Diplachne patens (J. Presl) E. Desv. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Diplachne peacockii Maiden & Betche, Agric. Gaz. New 
South Wales 15: 925, with plate. 1904. T-Specim.: | 
of 1. Australia: New South Wales: Coolabah, 4 Dec 
1904, Maiden & Boorman s.n. [LT: NSW-126631; IT: 
BM, BRI, K, W], LT designated (as holotype) by 
Lazarides, Brunonia 3: 263 (1980). 

= Leptochloa decipiens subsp. peacockii (Maiden & 
Betche) N. Snow. 


Diplachne pringlei Vasey ex Beal, Grass. N. Amer. 2: 436. 
1896. 
NOTE: nom. inval., as syn. of /eptochloa pringlei Beal. 
= Leptochloa dubia (Kunth) Nees. 


Diplachne procumbens Arechav., Anales Mus. Nac. 
Montevideo 1: 414. 1894. T-Specim.: 1 of 1. 
Argentina: Buenos Aires, Feb 1892, Spegazzini s.n. 
[HT: MNM; IT: BAA-914]. 

= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Diplachne procumbens (Muhl.) Nash, Man. FI. N. States 128. 
1901. Basionym: Festuca procumbens Muhl. 
NOTE: hom. illeg. 
E. NAME: Diplachne procumbens Arechav. 
CORRECT NAME: Diplachne maritima E.P. Bicknell. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Diplachne reptatrix(L.) Druce, Bot. Arr. Brit. Pl. (ed. 2) 129. 
1928. Basionym: Festuca reptatrix L. 
= Leptochloa fusea (L.) Kunth. 


Diplachne reverchonii Vasey, Bull. Torrey Bot. Club 13(7): 
118. 1886. T-Protol.. USA: Texas: Llano Co.: 
collected on granitic rocks, J. Reverchon 1613. T- 
Specim.: | of 1. USA: Texas: Llano Co: on granite 
rocks, May 1885, J. Reverchon 1613 [IT: ST: US- 
78819; IT: K, LL, MO-1837490, MO-2108836]. 

NOTE: as Diplachne "Reverchoni". 
= Tripogon spicatus (Nees) Ekman. 


Diplachne rigida Vasey, U.S.D.A. Div. Bot. Bull, 12(2): t. 
44. 1891. T-Specim.: | of 1. USA: Texas, J/ 
Reverchon 30 [HT: US-90883 1]. 

= Eragrostis sessilispica Buckley. 


Diplachne scabra (Nees) Nicora, Hickenia 2(19): 91. 1993. 
Basionym: Leptochloa scabra Nees. 
= Leptochloa scabra Nees. 


Diplachne simplex D6ll, Fl. Bras. 2(3): 97. 1878. 
NOTE: nom. illeg. superfl. 
= Tripogon spicatus (Nees) Ekman. 


Diplachne spicata (Nees) Doll, Fl. Bras. 2(3): 160. 1878. 
Basionym: Bromus spicatus Nees. 
= Tripogon spicatus (Nees) Ekman. 


Diplachne tarapacana Phil., Verz. Antofagasta Pfl. 88. 1891. 
T-Protol.: Chile: habitat prope oppidum Tarapaca, F- 
Philippi s.n. 

= Leptochloa fusea subsp. uninervia (J. Presl) N. Snow. 


Diplachne tracyi Vasey, Bull. Torrey Bot. Club 15: 40. 1888. 
T-Specim.: 1 of 2. USA: Nevada: Washoe Co., 1887, 
S.M. Tracy 216 [IT: US-78818]. 2 of 2. USA: Nevada, 
Reno, 17 Jul 1887, 8. M. Tracy [IT: NY-19502, NY- 
19503]. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


70 Catalogue of New World Grasses: II. 


Diplachne uninervia (J. Presl) Parodi, Revista Centro Estud. 


Agron, 18: 147. 
uninervia J. Presl. 
= Leptochloa fusea subsp. uninervia (J. Pres!) N. Snow. 


1925. Basionym: Megastachya 


Diplachne uninervia fo. abbreviata Parodi, Revista Fac. 
Agron. Univ. Nac. La Plata 6: 36. 1927. T-Protol.: 
Argentina: Buenos Aires: Pergamino, Basualdo, 13 
Nov 1925, L.R. Parodi 6703 [HT:BAA]. 

= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Diplachne uninervia vat. procumbens (Arechav.) Parodi, 
Revista Fac. Agron. Veterin. (Buenos Aires) 6: 37. 
1927. Basionym: Diplachne procumbens Arechav. 
= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Diplachne uninervia (J. Presl) Parodi fo. wninervia. 
= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Diplachne uninervia (J. Presl) Parodi var. uninervia. 
= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Diplachne virgata (J. Presl) Hack., Oesterr. Bot. Z. 52: 276. 
1902. Basionym: Bromus virgatus J. Presl. 
= Gouinia virgata (J. Presl) Scribn. 


Diplachne viscida Scribn., Bull. Torrey Bot. Club 10(1): 30, 
1883. T-Protol.: USA: Arizona: Santa Cruz Valley, 
near Tuscon, 28 Jun 1881, C.G. Pringle s.n. T- 
Specim.: | of 1. USA: Arizona, Pima Co. Near 
Tucson, C. G. Pringle [T: IT:NY-19501 (Possible)]. 
= Leptochloa viscida (Scribn.) Beal. 


Diplocea Raf., Amer. J. Sci. 1: 252. 1818. 
= Triplasis P. Beauv. 


Diplocea barbata Raf., Amer. J. Sci. 1: 252. 1818. T-Protol.: 
Raf. s.n. 
NOTE: nom. prov. 
= Triplasis purpurea (Walter) Chapm. 


Disakisperma Steud., Syn. Pl. Glumac. 1: 287. 12-13 Apr 
1854 ("1855"). TYPE: Disakisperma mexicana Steud. 
= Leptochloa P. Beauv. 


Disakisperma mexicana Steud., Syn. Pl. Glumac. 1: 287. 
1854. T-Specim.: | of 1. Mexico, 1845, E.G. von 
Steudel 112 OR 738 [IT: US-865873 (fragm.)]. 

= Leptochloa dubia (Kunth) Nees. 


Distichlis Raf., J. Phys. Chim. Hist. Nat. Arts 89: 104, 
1819. TYPE: Distichlis maritima Raf. 
TRTD. BY: Peterson, P. M. 
NHT: Monanthochloinae. NO. ACCPTD. SPP.: 6, SUBSP.: 
12. 
CNSLTD. LIT.: Beetle, A. A., (1943); Beetle, A. A., 
(1955). 
SYN.: Trisiola Raf. 


Distichlis araucana Phil., Anales Mus. Nac. Santiago de 
Chile 94: 162. 1896. T-Specim.: 1 of 2. Chile: 
Araucania ad Nacimiento: in arenosa ripa fluminis 
Vergara, Jan 1877 [ST]. 2 of 2. Chile: in arenis praedii 
Renaico, 1871 [ST]. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis australis (Speg.) Villamil, Kurtziana 5: 388. 1969. 
Basionym: Monanthochloe australis Speg. 
DST.: Arge. 


LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Nicora, E. G., M. E. De Paula, A. M. 
Faggi, M. del C. Mariano, A. M. Miante A., L. R. 
Parodi, C. A. Petetin, F. A. Roig & Z. Ruigolo de 
Agrasar (1978). 

SYN.: Monanthochloe australis Speg. 


Distichlis condensata (J. Presl) Hemsl., Biol. Cent.-Amer., 
Bot. 3(20): 578. 1885. Basionym: Megastachya 
condensata J. Presl. 

= Eragrostis condensata (J. Presl) Steud. 


Distichlis dentata Rydb., Bull. Torrey Bot. Club 36: 536. 
1909. T-Protol.: USA: Washington: 1893, Sandberg & 
Leiberg 463 (HT: NY], Pistillate plant. T-Specim.: | 
of 1. USA: Washington, 14 Jul 1893,/.H. Sandberg & 
J.B. Leiberg 463 (HT: NY; IT: US-2947337]. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis deserticola Phil., Anales Univ. Chile 36: 209. 
1870. T-Protol.: Chile: viaje al desierto de Atacama, 
Philippi 407. 

= Distichlis spicata var. mendocina (Phil.) Hack. 


Distichlis distichophylla (Labill.) Fassett, Rhodora 27: 71. 
1925. Basionym: Uniola distichophylla Labill. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis hirsuta Phil., Anales Univ. Chile 36: 209. 1870. T- 
Protol.: Chile: Santiago. 
= Distichlis spicata var. mendocina (Phil.) Hack. 


Distichlis hirta Phil., Anales Univ. Chile 48: 570. 1873. T- 
Protol.: Chile: en los arenales de Llico i Vichuqen, 
Luis Landbeck s.n. 
= Distichlis spicata var. mendocina (Phil.) Hack. 


Distichlis humilis Phil., Verz. Antofagasta Pfl. 86. 1891. T- 
Protol.: Chile: Tarapaca: prope Antofagasta lecta, F. 
Philippi s.n. 

DST.: Arge, Boli, Chil. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1955); Foster, R. C. 
(1958); Hitchcock, A. S. (1927); Macbride, J. F. 
(1936); Marticorena, C. & M. Quezada (1985); 
Renvoize, S. A. (1998); Tovar, O. (1993). 

SYN.: Distichlis misera Phil.; Distichlis spicata var. humilis 
(Phil.) Kuntze. 


Distichlis laxiflora Hack., Anales Mus. Nac. Buenos Aires 
21: 141. 1911. T-Protol.: Argentina: Buenos Aires: 
Chapal-Malal, La Barrancosa, F. C. Sur, Stuckert 
[Hrb. arg. 17468] [HT: Stuckert Hrb.]. T-Specim.: | 
of 1. Argentina: Buenos Aires, Stuckert 17468 [HT]. 

DST.: Arge. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Nicora, E. G. (1970); Nicora, E. G., 
M. E. De Paula, A. M. Faggi, M. del C. Mariano, A. 
M. Miante A., L. R. Parodi, C. A. Petetin, F. A. Roig 
& Z. Rugolo de Agrasar (1978). 

SYN.: Distichlis scoparia fo. laxiflora (Hack.) Beetle. 


Distichlis marginata Phil., Verz. Antofagasta Pfl. 86. 1891. 
T-Protol.: Chile: Tarapaca: ad Breas lecta est, F. 
Philippi s.n., [Philippi Tarapaca] from title but Breas is 
in Antofagasta. 

= Distichlis spicata var. mendocina (Phil.) Hack. 


Catalogue of New World Grasses: II. 71 


Distichlis  marginata var. stricta (Torr.) Thurb., Bot. 
California 2: 306. 1880. Basionym: Uniola_ stricta 
Torr. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis maritima Raf., J. Phys. Chim. Hist. Nat. Arts 89: 

104. 1819. T-Protol.: Atlantic coast of North America. 

NOTE: based on Uniola spicata L. as noted by A.S. Hitchce. 
= Distichlis spicata subsp. spicata. 


Distichlis maritima var. stricta (Torr.) Thurb., Bot. California 
2: 306. 1880. Basionym: Uniola stricta Torr. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis mendocina Phil., Anales Univ. Chile 36: 209. 1870. 


T-Protol.: Mendoza. T-Specim.: | of 1. Philippi s.n. 
(HT: SGO-045729]. 
= Distichlis spicata var. mendocina (Phil.) Hack. 


Distichlis misera Phil., Verz. Antofagasta Pfl. 86. 1891. T- 
Protol.: Chile: Tarapaca: locis dictis Vegas del Diablo 
reperta, F. Philippi s.n. T-Specim.: | of 1. Chile: 
Vegas del Diablo prope Antofagasta, Jan 1885, 
Philippi s.n. (HT: SGO]. 

= Distichlis humilis Phil. 


Distichlis multinervosa (Vasey) Piper, Proc. Biol. Soc. Wash. 
18: 147. 1905. Basionym: Melica multinervosa Vasey. 
= Vaseyochloa multinervosa (Vasey) Hitchce. 


Distichlis nodosa Raf., J. Phys. Chim. Hist. Nat. Arts 89: 
104. 1819. 
REPLACED NAME: Festuca distichophylla Michx. 
BLOCKING NAME: Distichlis distichophylla (Labill.) 
Fassett. 
= Distichlis spicata (L.) Greene. 


Distichlis palmeri (Vasey) Fassett ex I.M. Johnst., Proc. 
Calif. Acad. Sci., ser. 4, 12: 984. 1924. Basionym: 
Uniola palmeri Vasey. 

DST.: Mexi. 

LVL. ACCPTC:: |. 

ALSO ACCPTD. BY: Beetle, A. A. (1987); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W. & R. Moran (1981); Shreve, F. & I. L. 
Wiggins (1964). 

SYN.: Uniola palmeri Vasey. 


Distichlis prostrata (Kunth) E. Desv., Fl. Chil. 6: 398. 1854. 
Basionym: Poa prostrata Kunth. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis scoparia (Nees ex Kunth) Arechay., Anales Mus. 
Nac. Montevideo 1: 457, t. 58. 1897. Basionym: Poa 
scoparia Nees ex Kunth. 

DST.: Arge, Chil, Urug. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1955); Burkart, A. 
(1969); Marticorena, C. & M. Quezada (1985); 
Nicora, E. G. (1970); Nicora, E. G., M. E. De Paula, 
A. M. Faggi, M. del C. Mariano, A. M. Miante A., L. 
R. Parodi, C. A. Petetin, F. A. Roig & Z. Rugolo de 
Agrasar (1978); Rosengurtt, B., B. R. Arrillaga de 
Maffei & P. Izaguirre de Artucio (1970). 

Poa scoparia Nees ex Kunth; Uniola scoparia (Nees 
ex Kunth) Steud. 


SYN:: 


Distichlis scoparia fo. erinacea Beetle, Revista Argent. 
Agron. 22(2): 90. 1955. T-Protol.: Argentina: Chubut: 


Manantial, 19 May 1903[HT: SI (herb. Hicken no. 
9645, ex herb. Minist. Agric.)]. 
= Distichlis scoparia var. erinacea (Beetle) Nicora. 


Distichlis scoparia var. erinacea (Beetle) Nicora, FI. 
Patagonica 3: 498. 1978. Basionym: Distichlis 
scoparia fo. erinacea Beetle. 

DST.: Arge. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Nicora, E. G., M. E. De Paula, A. M. 
Faggi, M. del C. Mariano, A. M. Miante A., L. R. 
Parodi, C. A. Petetin, F. A. Roig & Z. Rugolo de 
Agrasar (1978). 

SYN.: Distichlis scoparia fo. erinacea Beetle. 


Distichlis scoparia fo. laxiflora (Hack.) Beetle, Revista 
Argent. Agron. 22(2): 90. 1955. Basionym: Distichlis 
laxiflora Hack. 

= Distichlis laxiflora Hack. 


Distichlis scoparia (Nees ex Kunth) Arechavy. fo. scoparia. 
= Distichlis scoparia var. scoparia. 


Distichlis scoparia (Nees ex Kunth) Arechav. var. scoparia. 

DST.: Arge, Chil, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Nicora, E. G., M. E. De Paula, A. M. 
Faggi, M. del C. Mariano, A. M. Miante A., L. R. 
Parodi, C. A. Petetin, F. A. Roig & Z. Rugolo de 
Agrasar (1978). 

SYN.: Distichlis scoparia fo. scoparia. 


Distichlis spicata (L.) Greene, Bull. Calif. Acad. Sci. 2: 415. 
1887. Basionym: Uniola spicata L. 

DST.: Arge, Beli, Boli, Cana, Cari, Chil, Ecua, Guat, Mexi, 
Peru, USA, Urug. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Abrams, L. (1923); Balick, M. J., M. 
H. Nee & D. E. Atha (2000); Beetle, A. A. (1943); 
Beetle, A. A. (1987); Brako, L. & J. L. Zarucchi 
(1993); Burkart, A. (1969); Catasus Guerra, L. 
(1997); Correll, D. S. & M. C. Johnston (1970); 
Cowan, C. P. (1983); Cronquist, A., A. H. Holmgren, 
N. H. Holmgren & J. L. Reveal (1977); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994); Edgar, E., H. E. Connor & J. E. Shand 
(1991); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Fernald, M. (1950); Foster, R. 
C. (1958); Galan-Mera, A. (1989); Gleason, H. A. & 
A. Cronquist (1968); Godfrey, R. K. & J. W. Wooten 
(1979); Gould, F. W. & R. Moran (1981); Great 
Plains Flora Association (1986); Hickman, J. C. 
(1993); Hitchcock, A. S. (1909); Hitchcock, A. S. 
(1927); Hitchcock, A. S. (1936); Hitchcock, A. S. 
(1951); Jorgensen, P. M. & C. Ulloa U. (1994); 
Jorgensen, P. M. & S. Le6n-Yanez (eds.) (1999); 
Lawesson, J. E., H. Adsersen & P. Bentley (1987); 
Long, R. W. & O. Lakela (1971); Macbride, J. F. 
(1936); McClure, F. A. (1955); McVaugh, R. (1983); 
Nicora, E. G. (1970); Nicora, E. G., M. E. De Paula, 
A. M. Faggi, M. del C. Mariano, A. M. Miante A., L. 
R. Parodi, C. A. Petetin, F. A. Roig & Z. Rugolo de 
Agrasar (1978); Pohl, R. W. (1994); Radford, A. E., 
H. E. Ahles & C. R. Bell (1968); Renvoize, S. A. 
(1998); Rosengurtt, B., B. R. Arrillaga de Maffei & 
P. Izaguirre de Artucio (1970); Scoggan, H. J. (1978 
[1979]); Seberg, O. & G. Petersen (1998); Small, J. 


72 Catalogue of New World Grasses: II. 


K. (1933); Tovar, O. (1993); Wunderlin, R. P. 
(1998). 

Agropyron peruvianum (Lam.) Roem. & Schult.; 
Briza spicata (L.) Lam.; Brizopyrum americanum 
(L.) Link; Brizopyrum ovatum Nees ex Steud.; 
Brizopyrum  prostratum var. erectum E. Fourn.; 
Brizopyrum  prostratum var. humile E.  Fourn.; 
Brizopyrum thalassicum (Kunth) Nees; Distichlis 
nodosa Raf.; Distichlis spicata var. dentata (Rydb.) 
C.L. Hitche.; Festuca multiflora Walter; Festuca 
triticea Lam. ex Kunth; Festuca triticoides Lam.; 
Megastachya thalassica (Kunth) Roem. & Schult.; 
Poa michauxii Kunth; Poa thalassica Kunth; 
Triticum peruvianum Lam.; Uniola spicata L.; 
Uniola thalassica (Kunth) Trin. 


SYN:: 


Distichlis spicata var. andina Beetle, Revista Argent. Agron. 
22(2): 93. 1955. T-Protol.: Bolivia: Guaqui, 6 Feb 
1937, L.R. Parodi 10082 (HT: BAA]. 
DST.: Arge, Boli, Ecua, Peru. 
LVL. ACCPTC:: 2. 
ALSO ACCPTD. BY: Beetle, A. A. (1955). 


Distichlis spicata var. borealis (J. Pres!) Beetle, Bull. Torrey 
Bot. Club 70(6): 643, f. 1, 8. 1943. Basionym: 
Brizopyrum boreale J. Presl. 

DST.: Cana, USA. 

LVL. ACCPTC.: 2. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1943); Hitchcock, C. L., A. Cronquist, M. Ownbey 
& J. W. Thompson (1969). 

SYN.: Brizopyrum boreale J. Presl; Poa borealis (J. Presl) 
Kunth. 


Distichlis spicata var. distichophylla (Michx.) Beetle, 
Rhodora 47(557): 148. 1945. Basionym: Festuca 
distichophylla Michx. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis spicata var. divaricata Beetle, Bull. Torrey Bot. 
Club 70(6): 647, f. 10. 1943. T-Protol.: USA: 
California: Riverside Co.: Salton, 1902,/.B. Davy s.n. 
{HT: UC]. 

DST.: Mexi, USA. 

LVL. ACCPTC.:: 2. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1943); Beetle, A. A. (1977); Beetle, A. A. (1987); 
Espejo Serna, A., A. R. L6pez-Ferrari, and J. Valdes- 
Reyna (2000); Munz, P. A. & D. D. Keck (1959), 


Distichlis spicata fo. hirta (Phil.) Kuntze, Revis. Gen. PI. 
3(3): 350. 1898. Basionym: Distichlis hirta Phil, 
= Distichlis spicata var. mendocina (Phil.) Hack. 


Distichlis spicata var. humilis (Phil.) Kuntze, Revis. Gen. PI. 
3(3): 350. 1898. Basionym: Distichlis humilis Phil. 
= Distichlis humilis Phil. 


Distichlis spicata var. laxa Vasey ex Beal, Grass. N. Amer. 2: 
519. 1896. T-Protol.: USA: Utah: Lake Park, 1887, 
Tracy s.n. T-Specim.: | of 1. USA: Utah: Lake Park, 
1887, Tracy s.n. [HT]. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis spicata var. marginata (Phil.) Kuntze, Revis. Gen. 
Pl. 3(2); 350. 1898. Basionym: Distichlis marginata 
Phil. 
= Distichlis spicata var. mendocina (Phil.) Hack. 


Distichlis spicata var. mendocina (Phil.) Hack., Anales 
Mus. Nac. Buenos Aires 6: 513. 1906. Basionym: 
Distichlis mendocina Phil. 

DST.: Arge, Boli. 

LVL. ACCPTC.: 2. 

ALSO ACCPTD. BY: Beetle, A. A. (1955); Burkart, A. 

(1969). 

Distichlis deserticola Phil.; Distichlis hirsuta Phil.; 

Distichlis hirta Phil.; Distichlis marginata Phil.; 

Distichlis mendocina Phil.; Distichlis spicata fo. 

hirta (Phil.) Kuntze; Distichlis spicata var.marginata 

(Phil.) Kuntze; Distichlis spicata var. thalassica 

(Kunth) Kuntze; Distichlis thalassica (Kunth) E. 

Desv.; Distichlis thalassica var. mendocina (Phil.) 

Kurtz; Distichlis thalassica var. pectinata Griseb.; 

Distichlis thalassica var. thalassica. 


SYN:: 


Distichlis spicata var. mexicana Beetle, Bull. Torrey Bot. 
Club 70: 648, f. 5. 1943. T-Protol.: Mexico: Durango: 
Durango, 1896, FE. Palmer 388 [ST: UC], (pistillate). 
T-Specim.: | of 2. Mexico: Durango, Jul 1896, E. 
Palmer 388 [ST: US-913771, US-748683]. 2 of 2. 
Mexico: Durango, E. Palmer /82 [ST]. 

DST.: Mexi. 

LVL. ACCPTC:: 2. 

ALSO ACCPTD. BY: Beetle, A. A. (1943); Beetle, A. A. 
(1977); Beetle, A. A. (1987): Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000). 


Distichlis spicata var. nana Beetle, Bull. Torrey Bot. Club 
70: 647, f. 3, 9. 1943. T-Protol.: USA: California: 
Tulare Co.: Whitaker Forest, 28 Jun 1928, P.B. 
Kennedy s.n. (HT: AHUC]. 

DST.: USA. 

LVL. ACCPTC.:: 2. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1943); Hitchcock, A. S. (1951); Munz, P. A. & D. 
D. Keck (1959). 


Distichlis spicata (L.) Greene subsp. spicata. 
DST.: Boli, Cana, Cari, Chil, Mexi, USA, Urug. 
LVL. ACCPTC:: 2. 
SYN.: Brizopyrum spicatum (L.) Hook. & Arn.; Distichlis 
maritima Raf. 


Distichlis spicata (L.) Greene var. spicata. 

DST.: Cana, Mexi, USA. 

LVL. ACCPTC.: 2. 

ALSO ACCPTD. BY: Beetle, A. A. (1943): Beetle, A. A. 
(1977); Beetle, A. A. (1987); Correll, D. S. & M. C. 
Johnston (1970); Gleason, H. A. & A. Cronquist 
(1991); Marticorena, C. & M. Quezada (1985). 


Distichlis spicata var. stolonifera Beetle, Bull. Torrey Bot. 
Club 70(6): 644, f. 4, 12. 1943. T-Protol.: USA: 
California: Humboldt Co.: near Ferndale, pistillate, 30 
Jul 1899, 1B. Davy & W.C. Blasdale 6202 [HT: UC]. 
T-Specim.: | of 1. USA: California: Humboldt Co.: 
near Ferndale, 30 Jul 1899, J.B. Davy & W.C. Blasdale 
6202 [HT: US; IT: US-913745]. 

DST.: Mexi, USA. 

LVL. ACCPTC:: 2. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1943); Beetle, A. A. (1977); Beetle, A. A. (1987); 
Espejo Serna, A., A. R. L6pez-Ferrari, and J. Valdes- 
Reyna (2000); Munz, P. A. & D. D. Keck (1959). 


Catalogue of New World Grasses: II. 73 


Distichlis spicata subsp. stricta (Torr.) Thorne, Aliso 9(2): 
195. 1978. Basionym: Uniola stricta Torr. 

DST.: Arge, Boli, Cana, Chil, Ecua, Mexi, Peru, USA, 
Urug. 

LVL. ACCPTC.:: 2. 

SYN.: Brizopyrum prostratum (Kunth) Benth. ex E. Fourn.; 
Brizopyrum  prostratum (Kunth) EE. Fourn.; 
Brizopyrum spicatum var. strictum (Torr.) A. Gray ex 
S. Watson; Distichlis araucana Phil.; Distichlis 
dentata Rydb.; Distichlis distichophylla (Labill.) 
Fassett; Distichlis marginata var. stricta (Torr.) 
Thurb.; Distichlis maritima var. stricta (Torr.) 
Thurb,; Distichlis nodosa Raf.; Distichlis prostrata 
(Kunth) E.  Desv.; Distichlis spicata var. 
distichophylla (Michx.) Beetle; Distichlis spicata var. 
laxa Vasey ex Beal; Distichlis spicata var. stricta 
(Torr.) Beetle; Distichlis spicata var. stricta (Tort.) 
Scribn.; Distichlis stricta (Torr.) Rydb.; Distichlis 
stricta var. dentata (Rydb.) C.L. Hitche.; Distichlis 
stricta var. laxa (Vasey ex Beal) Fawc. & West ex 


Munz; Distichlis stricta var. stricta; Distichlis 
tenuifolia Phil.; Distichlis viridis Phil.; Festuca 
distichophylla (Labill.) Hook. f.; Festuca 
distichophylla | Michx.; Megastachya _ prostrata 


(Kunth) Roem. & Schult.; Poa distichophylla 
(Labill.) R. Br.; Poa michauxii Kunth; Poa paradoxa 
Roem. & Schult.; Poa prostrata Kunth; Uniola 
distichophylla (Michx.) Roem. & Schult.; Uniola 
distichophylla Labill.; Uniola flexuosa Buckley; 
Uniola multiflora Nutt.; Uniola prostrata (Kunth) 
Trin.; Uniola stricta Torr. 


Distichlis spicata var. stricta (Torr.) Scribn., Mem. Torrey 
Bot. Club 5(4): 51. 1894. Basionym: Uniola stricta 
Torr. 
NOTE: varietal status is indicated in the introduction (p. 7) 
of this paper. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis spicata var. stricta (Torr.) Beetle, Bull. Torrey Bot. 
Club 70(6): 645, f. 6. 1943. Basionym: Uniola stricta 
Torr. 
NOTE: nom. illeg. 
E. NAME: Distichlis spicata var. stricta (Torr.) Scribn. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis spicata var. thalassica (Kunth) Kuntze, Revis. Gen. 
PI. 3(3): 350. 1898. Basionym: Poa thalassica Kunth. 
= Distichlis spicata var. mendocina (Phil.) Hack. 


Distichlis stricta (Torr.) Rydb., Bull. Torrey Bot. Club 
32(11): 602. 1905. Basionym: Uniola stricta Torr. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis stricta var. dentata (Rydb.) C.L. Hitche., Vasc. PI. 
Pacific NW 1: 553. 1969. Basionym: Distichlis 
dentata Rydb. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis stricta var. laxa (Vasey ex Beal) Fawc. & West ex 
Munz, Man. S. Calif. Bot. 52, 597. 1935. Basionym: 
Distichlis spicata var. laxa Vasey ex Beal. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis stricta (Torr.) Rydb. var. stricta. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis tenuifolia Phil., Anales Univ. Chile 94: 162. 1896. 
T-Protol.: Chile: in arena litorea insulae Quiriquinae et 
prope Talcahuano crescit, Philippi s.n. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Distichlis texana (Vasey) Scribn., Circ. Div. Agrostol. 
U.S.D.A. 16: 2. 1899. Basionym: Poa texana Vasey. 
= Allolepis texana (Vasey) Soderstr. & H.F. Decker. 


Distichlis thalassica (Kunth) E. Desv., Fl. Chil. 6: 397. 1854. 
Basionym: Poa thalassica Kunth. 
= Distichlis spicata var. mendocina (Phil.) Hack. 


Distichlis thalassica var. mendocina (Phil.) Kurtz, Bol. Acad. 
Ci. (Cordoba) 15: 521. 1897. Basionym: Distichlis 
mendocina Phil. 

= Distichlis spicata var. mendocina (Phil.) Hack. 


Distichlis thalassica var. pectinata Griseb., Abh. Konigl. 
Ges. Wiss. Gottingen 24: 291. 1879. T-Protol.: 
[Lorentz] s.n., the following 3 specimens may all refer 
to the same collection. T-Specim.: | of 3. Cordoba, 
[Lorentz] s.n. [T]. 2 of 3. Chile: Santiago del Estero, 
[Lorentz] s.n. [T]. 3 of 3. Pampa pr. Laguna de Pocho 
[T]. 

= Distichlis spicata var. mendocina (Phil.) Hack. 


Distichlis thalassica (Kunth) E. Desv. var. thalassica. 
= Distichlis spicata var. mendocina (Phil.) Hack. 


Distichlis viridis Phil., Anales Univ. Chile 94: 163. 1896. T- 
Protol.: Chile: prope Monte Amargo inter Caldera et 
Copiapo lecta fuit, Philippi s.n. T-Specim.: | of 1. 
Chile: Atacama, Coll. Ukn. 2/2 [IT: US-1763463], CS. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Doellochloa Kuntze, Revis. Gen. Pl. 2: 773. 1891. TYPE: 
Doellochloa fastigiata (Nees) Kuntze. 
NOTE: nom. illeg. superfl. 
= Gymnopogon P. Beauv. 


Doellochloa fastigiata (Nees) Kuntze, Revis. Gen. Pl. 2: 773. 
1891. Basionym: Gymnopogon fastigiatus Nees. 
= Gymnopogon fastigiatus Nees. 


Echinanthus Cerv., Naturaleza (Mexico City) 1: 351. 1870. 
= Tragus Haller. 


Ectosperma Swallen, J. Wash. Acad. Sci. 40(1): 19. 1950. 
TYPE: Ectosperma alexandrae Swallen. 
NOTE: hom. illeg. 
= Swallenia Soderstr. & H.F. Decker. 


Ectosperma alexandrae Swallen, J. Wash. Acad. Sci. 40(1): 
19, f. 1. 1950. T-Protol.: USA: California: Inyo Co.: 
Eureka Valley, on lower slopes of big sand hill, 3050 
ft, 24 May 1949, A. M. Alexander & L. Kellogg 5655 
[HT: US-1963637]. T-Specim.: 1 of 1. USA: 
California: Inyo Co.: Eureka Valley, 24 May 1949,M. 
Alexander & L. Kellogg 5655 (HT: US-1963637]. 

= Swallenia Soderstr. & H.F. Decker. 


Eleusine Gaertn., Fruct. Sem. Pl. 1: 7. 1788. TYPE: 
Eleusine coracana (L.) Gaertn.; LT designated by 
Nash in Britton & Brown, Ill. Fl. N. U.S. (ed. 2) 1: 228 
(1913). 
TRTD. BY: Peterson, P. M. & K. W. Hilu. 
NHT: Eleusininae. NO. ACCPTD. SPP.: 4 SUBSP.: 2. 


74 Catalogue of New World Grasses: II. 


CNSLTD. LIT.: Phillips, S. M., (1972); Werth, C. R., K. 
Hilu, C. A. Langer & W. V. Baird, (1993). 


Eleusine aegyptia (L.) Desf. Fl. Atlant. I: 85. 1798. 
Basionym: Cynosurus aegyptius L. 
NOTE: as "aegyptiaca", a correctable error. 


= Dactyloctenium aegyptium (L.) Willd. 


Eleusine aegyptia (L.) Pers., Syn. Pl. 1: 87. 1805. Basionym: 
Cynosurus aegyptius L. 
NOTE: isonym. 
E. NAME: Eleusine aegyptia (L.) Desf. 
= Dactyloctenium aegyptium (L.) Willd. 


Eleusine aegyptia Raf., Chlor. Aetn. 7. 1815. 
NOTE: without citation of basionym, presumably same as 
Eleusine aegyptia Desf. 
E. NAME: Eleusine aegyptia (L.) Desf. 
= Dactyloctenium aegyptium (L.) Willd. 


Eleusine aegyptia (L.) Roxb., Fl. Ind., ed. 1820 1: 345. 1820. 
Basionym: Cynosurus aegyptius L. 
NOTE: isonym, as "aegyptica", a correctable error. 
E. NAME: Eleusine aegyptia (L.) Desf. 
= Dactyloctenium aegyptium (L.) Willd. 


Eleusine aegyptia (L.) Roberty, Petite Fl. l'Quest-Afr. 386. 
1954. Basionym: Cynosurus aegyptius L. 
NOTE: isonym, also Roberty in Bull. Inst. France. Afr. 
Noire, Ser.A., 18: 50, 1955. 
E. NAME: Eleusine aegyptia (L.) Desf. 
= Dactyloctenium aegyptium (L.) Willd. 


Eleusine aegyptiaca (L.) Desf., Fl. Atlant. 1: 85. 1798. 
Basionym: Cynosurus aegyptius L. 
NOTE: orth var., see E. aegyptia. 
= Dactyloctenium aegyptium (L.) Willd. 


Eleusine africana Kenn.-O'Byrne, Kew Bull. 12: 65. 1957. T- 
Protol.: South Africa: Cape Province: Kimberly 
District: Warrenton-on-Waal, Mar 1952, Wilman HK1. 
T-Specim.: | of 1. South Africa: Cape Province, 
Kimberley Dist.: Warrenton-on-Waal, Mar 1950, M. 
Wilman H.K.1 (HT: K; IT: US-2589173]. 

= Eleusine coracana subsp. africana (Kenn.-O'Byrne) Hilu 
& de Wet. 


Eleusine barbata Desy., Mem. Soc. Agric. Angers 1: 208. 
1831. T-Protol.: Crescit in Brasilio. T-Specim.: | of 1. 
Brazil, Coll. Ukn. s.n. [T: US-88709]. 

NOTE: also in Opusc. Sci. Phys. Nat. 104 [reprint] 1831- 
1833. 
= Leptochloa barbata (Desv.) Parodi ex Nicora. 


Eleusine calycina Roxb., Hort. Bengal. 8. 1814. 
NOTE: nom. nud. 
= Dinebra retroflexa (Vahl) Panz. 


Eleusine calycina Roxb., Fl. Ind., ed. 1820 1: 347. 1820. T- 
Protol.: On dry pasture ground, but generally amongst 
bushes. 

= Dinebra retroflexa (Vahl) Panz. 


Eleusine cerealis Salisb., Icon. Stirp. Rar. 19. 1796. 
NOTE: nom. illeg. superfl. 
= Eleusine coracana (L.) Gaertn. 


Eleusine cerealis Ehrenb. ex Sweet, Hort. Brit. (ed. 2) 571. 
1830. 
NOTE: hom. illeg. 


E. NAME: Eleusine cerealis Salisb. 

= Eleusine coracana (L.) Gaertn. 
Eleusine coracana (L.) Gaertn., Fruct. Sem. Pl. 1: 8, t. 1, f. 

11. 1788. Basionym: Cynosurus coracanus L. 

DST.: USA. 

LVL. ACCPTC.: 1; cultivated, introduced. 
ALSO ACCPTD. BY: Bor, N. L. (1960); Hitchcock, A. S. 
(1951); ORSTOM (1988); Phillips, S. (1995); 
Phillips, S. M. (1972). 

Cynosurus coracanus L.; Eleusine cerealis Ehrenb. 
ex Sweet; Eleusine cerealis Salisb.; Eleusine 
coracana var. stricta (Roxb.) Nees; Eleusine 
coracana var. tocussa (Fresen.) Franch.; Eleusine 
indica subsp. coracana (L.) Lye: Eleusine indica var. 
coracana (L.) Fiori; Eleusine indica var. stricta 
(Roxb.) Chiov.; Eleusine luco Welw.; Eleusine pilosa 
Gilli; Eleusine sphaerosperma Stokes; Eleusine 
stricta Roxb.; Eleusine tocussa Fresen. 


SYN:: 


Eleusine coracana subsp. africana (Kenn.-O'Byrne) Hilu & 
de Wet, Econ. Bot. 30: 202. 1976. Basionym:Eleusine 
africana Kenn.-O'Byrne. 

NOTE: nom. inval., basionym not cited. 
= Eleusine coracana subsp. africana (Kenn.-O'Byrne) Hilu 
& de Wet. 


Eleusine coracana subsp. africana (Kenn.-O'Byrne) Hilu & 
de Wet, Phytologia 78(5): 411. 1994. Basionym: 
Eleusine africana Kenn.-O'Byrne. 

DST.: USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Gibbs Russell, G. E, W. G. Welman, 
E. Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987). 

Eleusine africana Kenn.-O'Byrne; Eleusine coracana 

subsp. africana (Kenn.-O'Byrne) Hilu & de Wet: 

Eleusine indica subsp. africana (Kenn.-O'Byrne) 

S.M. Phillips. 


SYN.: 


Eleusine coracana (L.) Gaertn. subsp. coracana. 
LVL. ACCPTC.: 1. 


Eleusine coracana var. stricta (Roxb.) Nees, Fl. Afr. Austral. 
Ill. 1: 251. 1841. Basionym: Eleusine stricta Roxb. 
= Eleusine coracana (L.) Gaertn. 


Eleusine coracana vat. tocussa (Fresen.) Franch., Bull. Soc. 
Hist. Nat. Autun 8: 377. 1893. Basionym: Eleusine 
tocussa Fresen. 

= Eleusine coracana (L.) Gaertn. 


Eleusine cruciata Lam., Tabl. Encycl. 1: 203, t. 48, f. 2. 
1791. T-Protol.: ex Indiis utrisque. 
NOTE: nom. illeg. superfl. for Cynosurus aegyptius. 
CORRECT NAME: Eleusine aegyptia (L.) Desf. 
= Dactyloctenium aegyptium (L.) Willd. 


Eleusine cruciata Elliott, Sketch Bot. S. Carolina 1: 176. 
1816. T-Protol.: Grows in cultivated ground, common. 
NOTE: hom. illeg. 
E. NAME: Eleusine cruciata Lam. 
= Dactyloctenium aegyptium (L.) Willd. 


Eleusine digitata (R. Br.) Spreng., Syst. Veg. 4: Cur. Post. 
36. 1827. Basionym: Poa digitata R. Br. 
= Leptochloa digitata (R. Br.) Domin. 


Catalogue of New World Grasses: II. 75 


Eleusine digitata Willd. ex Steud., Nomencl. Bot. (ed. 2) 549. 
1840. 
NOTE: nom. illeg., as syn. of Leptochloa digitaria (Kunth) 
Nees. 
= Leptochloa virgata (L.) P. Beauv. 


Eleusine distachya Nees, F1. Bras. Enum. PI. 440. 1829. 
NOTE: nom. inval., as syn. of Eleusine indica, Steud. Nom. 
Bot. ed. 2 1: 549. 1840. 
= Eleusine indica (L.) Gaertn. 


Eleusine distans Moench, Methodus 210. 1794. 
NOTE: nom. illeg. superfl. 
= Eleusine indica (L.) Gaertn. 


Eleusine domingensis (Jacq.) Pers., Syn. Pl. 1: 87. 1805. 
Basionym: Cynosurus domingensis Jacq. 
= Leptochloa virgata (L.) P. Beauv. 


Eleusine domingensis Sieber ex Schult., Mant. 2: 323. 1824. 
NOTE: nom. inval., as syn. of Eleusine indica (L.) Gaertn., 
hom. illeg. 
E. NAME: Eleusine domingensis (Jacq.) Pers. 
= Eleusine indica (L.) Gaertn. 


Eleusine egyptia Raf., Precis Decouy. Somiol. 45. 1814. 
NOTE: Presumably same as Eleusine aegyptia Desf. 
E. NAME: Eleusine aegyptia (L.) Desf. 
= Dactyloctenium aegyptium (L.) Willd. 


Eleusine elongata Willd. ex Steud., Nomencl. Bot. (ed. 2) 1: 


549. 1840. 
NOTE: nom. inval., as syn. of Leptochloa filiformis (Lam.) 
P. Beauv. 
= Leptochloa panicea subsp. mucronata (Michx.) 
Nowack. 


Eleusine filiformis Pers., Syn. Pl. 1: 87. 1805. 
REPLACED NAME: Festuca filiformis Lam. 
BLOCKING NAME: Festuca filiformis Pourret. 
= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Eleusine glabra Schumach., Beskr. Guin. Pl, 53-54. 1827. T- 
Protol.; Guinea. T-Specim.: | of 1. Ghana, Thonning 
392 |HT: C: IT: kK]. 
NOTE: also in’ Kongel. Danske  Vidensk. 
Naturvidensk. Math. Afh. 3: 73-74 (1828). 
= Eleusine indica (L.) Gaertn. 


Selsk. 


Eleusine glaucophylla (Courbai) Munro ex Benth., J. Linn. 
Soc., Bot. 19: 107. 1881. Basionym: Dactyloctenium 
glaucophyllum Courbai. 

= Dactyloctenium scindicum Boiss. 


Eleusine gonantha Schrank, Syll. Pl. Nov. 1: 191. 1824. T- 
Protol.: Brazil: Habitat in Brasilia, unde D. de Martius 
semina misit. 

= Eleusine indica (L.) Gaertn. 


Eleusine gouini E. Fourn., Mexic. Pl. 2: 146. 1886. T-Protol.: 
Mexico: Veracruz, Gouin 66. T-Specim.: 1 of 2. 
Mexico: Veracruz, Gouin 66 [HT]. 2 of 2. 

= Eleusine indica (L.) Gaertn. 


Eleusine gracilis Salisb., Prodr. Stirp. Chap. Allerton 19. 
1796. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Cynosurus indicus L. 
= Eleusine indica (L.) Gaertn. 


Eleusine gracilis (Kunth) Spreng., Syst. Veg. 1: 350. 1825. 
Basionym: Chloris gracilis Kunth. 
NOTE: nom. illeg. 
= Leptochloa virgata (L.) P. Beauv. 


Eleusine inaequalis E. Fourn. ex Hemsl., Biol. Cent.-Amer., 
Bot. 3: 565. 1885. 
NOTE: nom. inval., as syn. Eleusine indica. 
= Eleusine indica (L.) Gaertn. 


Eleusine inaequalis E. Fourn., Mexic. Pl. 2: 145. 1886. T- 
Specim.: | of 7. Mexico: Vera Cruz, Gouin 67 [ST]. 2 
of 7. Orizaba, Bott. 125 [ST: herb. Van Heurck]. 3 of 
7. Orizaba, Bott. 126 {ST: herb. Van Heurck]. 4 of 7. 
Orizaba, Bott. 1/258 [ST: herb. Van Heurck]. 5 of 7. 
Izhuatlancillo, Bourg. 2378 part. [ST]. 6 of 7. 
Chapultepec, Schaffn. 34 [ST]. 7 of 7. San. Luis de 
Potosi, Virl. s.n. [ST]. 

= Eleusine indica (L.) Gaertn. 


Eleusine indica (L.) Gaertn., Fruct. Sem. Pl. 1: 8. 1788. 
Basionym: Cynosurus indicus L. 

DST.: Arge, Beli, Boli, Braz, Cana, Cari, Chil, Colo, CoRi, 
Ecua, ElSa, FrGu, Guat, Guya, Hond, Mexi, Nica, 
Pana, Para, Peru, Suri, USA, Urug, Vene. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Abrams, L. (1923); Adams, C. D. 
(1972); Allem, A. C. & J. F.M. Valls (1987); Balick, 
M. J., M. H. Nee & D. E. Atha (2000); Beetle, A. A. 
(1977); Berendsohn, W. G. & A. E. Araniva de 
Gonzalez (1991); Bor, N. L. (1960); Brako, L. & J. 
L. Zarucchi (1993); Breedlove, D. E. (1986); Burkart, 
A. (1969); Cabrera, A. L. (1970); Catasus Guerra, L. 
(1997); Correll, D. S. & M. C. Johnston (1970); 
Cowan, C. P. (1983); Cronquist, A., A. H. Holmgren, 
N. H. Holmgren & J. L. Reveal (1977); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994); Dodson, C. H. & A. H. Gentry (1978); 
Dodson, C. H., A. H. Gentry & F. M. Valverde 
(1985); Doll, J. C. (1878); Edgar, E., H. E. Connor & 
J. E. Shand (1991); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Fernald, M. 
(1950); Filgueiras, T. S. (1991); Foster, R. C. (1958); 
Gleason, H. A. & A. Cronquist (1968); Gleason, H. 
A. & A. Cronquist (1991); Gould, F. W (1979); 
Gould, F. W. & R. Moran (1981); Great Plains Flora 
Association (1986); Hickman, J. C. (1993); 
Hitchcock, A. S. (1909); Hitchcock, A. S. (1927); 
Hitchcock, A. S. (1936); Hitchcock, A. S. (1951); 
Hitchcock, C. L., A. Cronquist, M. Ownbey & J. W. 
Thompson (1969); Jorgensen, P. M. & S. Leon- 
Yanez (eds.) (1999); Judziewicz, E. J. (1990); 
Killeen, T. J. (1990); Lawesson, J. E., H. Adsersen & 
P. Bentley (1987); Lindeman, J. C. & A. L. Stoffers 
(1963); Liogier, H. & L. Martorell (1982); Long, R. 
W. & O. Lakela (1971); Marticorena, C. & M. 
Quezada (1985); McClure, F. A. (1955); McVaugh, 
R. (1983); Molina R., A. (1975); Munz, P. A. & D. 
D. Keck (1959); Munz, P. A. (1974); ORSTOM 
(1988); Phillips, S. (1995); Phillips, S. M. (1972); 
Pohl, R. W. (1980); Pohl, R. W. (1994); Porter, D. 
M. (1983); Radford, A. E., H. E. Ahles & C. R. Bell 
(1968); Reeder, J. R. & C. G. Reeder (1971); Renner, 
S. S., H. Balsley & L. B. Holm-Nielsen (1990); 
Renvoize, S. A. (1984); Renvoize, S. A. (1988); 
Renvoize, S. A. (1998); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 


76 Catalogue of New World Grasses: II. 


Rutenberg, C. (1880-1889); Scoggan, H. J. (1978 
[1979]); Shreve, F. & I. L. Wiggins (1964); 
Skottsberg, C. (1922); Small, J. K. (1933); Smith, L. 
B., D. C. Wasshausen & R. M. Klein (1981); Sousa 
S.,M. & E. F. Cabrera C. (1983); Tovar, O. (1993); 
Voss, E. G. (1972); Wiggins, I. L. & D. M. Porter 
(1971); Wunderlin, R. P. (1998). 

SYN.: Chloris repens Steud.; Cynodon indicus (L.) Raspail; 
Cynosurus indicus L., Eleusine distachya Nees; 
Eleusine distans Moench, Eleusine domingensis 
Sieber ex Schult.; Eleusine glabra Schumach.; 
Eleusine gonantha Schrank; Eleusine gouini E. 
Fourn.; Eleusine gracilis Salisb.; Eleusine inaequalis 
E. Fourn.; Eleusine inaequalis E. Fourn. ex Hemsl.; 
Eleusine indica var. major E. Fourn.; Eleusine indica 
var. sandaensis Vanderyst; Eleusine rigidifolia E. 
Fourn.; Eleusine rigidifolia E. Fourn. ex Hemsl.; 
Eleusine scabra E. Fourn.; Eleusine scabra_ E. 
Fourn.; Eleusine scabra E. Fourn. ex Hemsl.; 
Eleusine textilis Welw. 


Eleusine indica subsp. africana (Kenn.-O'Byrne) S.M. 
Phillips, Kew Bull. 27(2): 259. 1972. Basionym: 
Eleusine africana Kenn.-O'Byrne. 

= Eleusine coracana subsp. africana (Kenn.-O'Byrne) Hilu 
& de Wet. 


Eleusine indica var. brachystachya Trin., Sp. Gram. |: t. 72. 
1828. 
REPLACED NAME: Triticum geminatum Spreng. 
= Eleusine tristachya (Lam.) Lam. 


Eleusine indica var. condensata D6ll, Fl. Bras. 2(3): 86. 
1878. T-Protol.: Brazil and Uruguay: Brazil: In 
Brasilia meridonali et ad Montevideo, Se//o. 

= Eleusine tristachya (Lam.) Lam. 


Eleusine indica subsp. coracana (L.) Lye, Lidia 4: 150. 1999, 
Basionym: Cynosurus coracanus L. 
= Eleusine coracana (L.) Gaertn. 


Eleusine indica var. coracana (L..) Fiori, Nuov. FI. Italia 1: 
114. 1923. Basionym: Cynosurus coracanus L. 
= Eleusine coracana (L.) Gaertn. 


Eleusine indica var. major E. Fourn., Mexic. Pl. 2: 145. 1886. 
T-Specim.: | of 5. Jicaltepec, Karw. & Liebm. 223 
[ST]. 2 of 5. in itinere a Colipa ad Jicaltepec, Karw. 
sn. [ST]. 3 of 5. Paso de Dona Juana, Liebm. 222 
[ST]. 4 of 5. Mundo neuvo, Karw. 955 [ST]. 5 of 5. 
Morro, Rancho nuevo, Lieb. 227 [ST]. 
= Eleusine indica (L.) Gaertn. 


Eleusine indica var. sandaensis WVanderyst, Bull. Agric. 
Congo Belge 11: 122. 1920. 
NOTE: nom. prov. based on Vanderyst 5459. 
= Eleusine indica (L.) Gaertn. 


Eleusine indica var. stricta (Roxb.) Chiov., Nuovo Giorn. 
Bot. Ital. 26: 83. 1919. Basionym: Eleusine stricta 
Roxb. 

= Eleusine coracana (L.) Gaertn. 


Eleusine indica var. tristachya (Lam.) Fiori, Nuov. FI. Italia 
1: 114. 1923. Basionym: Cynosurus tristachyos Lam. 
= Eleusine tristachya (Lam.) Lam. 


Eleusine leucosperma Bernh. ex Steud., Nomencl. Bot. (ed. 
2) 1: 549. 1840. 
NOTE: nom. inval., as syn. of Eleusine oligostachya. 


= Eleusine tristachya (Lam.) Lam. 


Eleusine luco Welw., Apont. 591. 1859[1859]. 
NOTE: nom. nud. 
= Eleusine coracana (L.) Gaertn. 


Eleusine mucronata Michx., Fl. Bor.-Amer. |: 65. 1803. T- 
Protol.: Hab. in cultis Hlinoensibus, Michaux s.n. T- 
Specim.: | of 1. USA: Illinois, Michaux s.n. [HT: P- 
MICH]. 

= Leptochloa  panicea 
Nowack. 


subsp. mucronata (Michx.) 


Eleusine mucronata Stokes, Bot. Mat. Med. 1: 150. 1812. T- 
Protol.: Species gathered by Broughton in Jamaica, 
Broughton s.n. 

NOTE: hom. illeg. 
E. NAME: Eleusine mucronata Michx. 
= Dactyloctenium aegyptium (L.) Willd. 


Eleusine mucronata (Michx.) Hornem., Hort. Bot. Hafn. 116. 
1819. Basionym: Chloris mucronata Michx, 
NOTE: hom. illeg., same sp. as Stokes 1812, but different 
type. 
E. NAME: Eleusine mucronata Michx. 
= Dactyloctenium aegyptium (L.) Willd. 


Eleusine multiflora Hochst. ex A. Rich., Tent. Fl. Abyss. 2: 
412. 1851 [1850]. T-Protol.: Crescit in pratis et locis 
graminosis circa Adowa nec non in provincia Chiré, 2 
syntypes. T-Specim.: | of 2. Ethiopia: in graminosis et 
campis Poae Abyssinicae prope Adoam, 20 Sep 1837, 
W. Schimper, pl. Schimp. Abyss., sect I, 110 [LT: P; 
IST: K, MO, US-3298790], LT designated by ?; cited 
(as type) by McVaugh, Fl. Novo-Galic. 14: 157 
(1983). 2 of 2. Abyssinia: circa Adoua, nec. non in 
provincia Chire, Quartin Dillon s.n. [ST]. 

DST.: Mexi. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Gibbs 
Russell, G. E, W. G. Welman, E. Reitief, K. L. 
Immelman, G. Germishuizen, B. J. Pienaar, M. van 
Wyk, and A. Nicholas (PRECIS) (1987); McVaugh, 
R. (1983); Phillips, S. (1995); Phillips, S. M. (1972); 
Pohl, R. W. (1994). 

SYN.: Eragrostis kwaiensis Peter. 


Eleusine oligostachya Link, Hort. Berol. 1: 60. 1827. T- 
Protol.: Hab. Hort. 
= Eleusine tristachya (Lam.) Lam. 


Eleusine pectinata Moench, Methodus 68. 1802. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Cynosurus aegyptius L. 
= Dactyloctenium aegyptium (L.) Willd. 


Eleusine pilosa Gilli, Ann. Naturhist. Mus. Wien 69: 50. 
1965[1966]. T-Specim.: | of 1. Tanzania: NW shore of 
Lake Nyassa near Mwaya, Gilli 78 [HT: W]. 
= Eleusine coracana (L.) Gaertn. 


Eleusine procera Spreng. ex Steud., Nomencl. Bot. (ed. 2) 1: 
549. 1840. 
NOTE: nom. illeg., as syn. of Uniola racemiflora Trin. 
= Uniola virgata (Poir.) Griseb. 


Catalogue of New World Grasses: II. 77 


Eleusine rigidifolia E. Fourn. ex Hemsl., Biol. Cent.-Amer., 
Bot. 3: 565. 1885. 
NOTE: nom. nud. 
= Eleusine indica (L.) Gaertn. 


Eleusine rigidifolia E. Fourn., Mexic. Pl. 2: 146. 1886. T- 
Specim.: | of 2. Mexico: In arenis prope Vera Cruz, 
Gouin s.n. [ST]. 2 of 2. Oajaca, Ghiesbreght s.n. [ST]. 

= Eleusine indica (L.) Gaertn. 


Eleusine scabra E. Fourn., Bull. Soc. Bot. France 27: 296. 
1880. 
NOTE: nom. nud. 
= Eleusine indica (L.) Gaertn. 


Eleusine scabra E. Fourn. ex Hemsl., Biol. Cent.-Amer., Bot. 
3: 565. 1885. T-Specim.: 1 of 1. Mexico, 20 Sep 1865- 
1866, M. Bourgeau 1030 (OM: US-884368]. 
NOTE: nom. nud. 
= Eleusine indica (L.) Gaertn. 


Eleusine scabra E. Fourn., Mexic. Pl. 2: 145. 1886. T- 
Specim.: | of 9. Mexico: Secus agros prope Mejico, 
Bourg. 1030 [ST: US: 884368]. 2 of 9. Secus agros 
Mejico, Bourg. 2378 part. [ST]. 3 of 9. San Luis de 
Potosi, Virl. 1435 [ST]. 4 of 9. Orizaba, Bilimek 454 
[ST: in herb. Coss.]. 5 of 9. Orizaba, Thomas s.n. [ST: 
in herb. Buchinger]. 6 of 9. Orizaba, Bourg. 2743 
[ST]. 7 of 9. Orizaba, Bourg 2634 [ST]. 8 of 9. 
Orizaba, F. Mell. 1392 [ST]. 9 of 9. Vera Cruz, Bouin 
67 [ST]. 

= Eleusine indica (L.) Gaertn. 


Eleusine scindica (Boiss.) Duthie, Fodder Grasses North. 
India 58. 1888. Basionym: Dactyloctenium scindicum 
Boiss. 

= Dactyloctenium scindicum Boiss. 


Eleusine sparsa Muhl., Descr. Gram. 135. 1817. T-Protol.: 
USA: Habitat in Carolina et Georgia. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Eleusine filiformis Pers. 
= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Eleusine sphaerosperma Stokes, Bot. Mat. Med. 1: 149. 
1812. T-Protol.: Cult: Specimens gathered in Paris 
garden. 

NOTE: nom. illeg. superfl. 

= Eleusine coracana (L.) Gaertn. 


Eleusine stricta Roxb., Fl. Ind., ed. 1820 1: 344. 1820. T- 
Protol.: In the Rajamundry Circar. 
= Eleusine coracana (L.) Gaertn. 


Eleusine stricta Willd. ex Steud., Nomencl. Bot. (ed. 2) 1: 
549. 1840. 
NOTE: nom. inval., as syn. of Leptochloa filiformis (Lam.) 
P. Beauv. 
E. NAME: Eleusine stricta Roxb. 
= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Eleusine textilis Welw., J. Travel Nat. Hist. 1: 31. 1868. 
NOTE: nom. prov. based on Welwitsch 1104 (BM). 
= Eleusine indica (L.) Gaertn. 


Eleusine tocussa Fresen., Mus. Senckenberg. 2: 141. 1837. T- 
Specim.: | of 2. Ethiopia: Dagussa, prope Adoam. 
Cult. at Frankfurt from Abyssinian seed, 15 Nov 1837, 
W.Schimper 347 [POSSIBLE TYPE: FR, US- 


3298794]. 2 of 2. Ethiopia: Celitur, W.Schimper 1969 
[POSSIBLE TYPE: US-3298784]. 
= Eleusine coracana (L.) Gaertn. 


Eleusine tristachya (Lam.) Lam., Tabl. Encycl. 1: 203. 1791 
[1792]. Basionym: Cynosurus tristachyos Lam. 

DST.: Arge, Boli, Braz, Chil, Para, USA, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Burkart, A. (1969); Cabrera, A. L. 
(1970); Edgar, E., H. E. Connor & J. E. Shand 
(1991); Gibbs Russell, G. E, W. G. Welman, E. 
Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987); Gleason, H. A. & A. Cronquist (1968); 
Hickman, J. C. (1993); Hitchcock, A. S. (1951); 
Marticorena, C. & M. Quezada (1985); Munz, P. A. 
(1968); Nicora, E. G., M. E. De Paula, A. M. Faggi, 
M. del C. Mariano, A. M. Miante A., L. R. Parodi, C. 
A. Petetin, F. A. Roig & Z. Rugolo de Agrasar 
(1978); Phillips, S. (1995); Renvoize, S. A. (1988); 
Renvoize, S. A. (1998); Rosengurtt, B.. B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 
Small, J. K. (1933); Smith, L. B., D. C. Wasshausen 
& R. M. Klein (1981); Trinajstic, I. & N. Jasprica 
(1998). 

Agropyron geminatum (Spreng.) Schult.; Cynosurus 
tristachyos Lam.; Eleusine indica var. brachystachya 
Trin.; Eleusine indica var. condensata D6ll; Eleusine 
indica var. tristachya (Lam.) Fiori; Eleusine 
leucosperma Bernh. ex Steud.; Eleusine oligostachya 
Link; Eleusine tristachya (Lam.) Kunth; Eleusine 
tristachya fo. latifolia Hack., Eleusine tristachyos 
(Lam.) Lam.; Triticum geminatum Spreng. 


SYN: 


Eleusine tristachya (Lam.) Kunth, Revis. Gramin. 1: 92. 
1829. Basionym: Cynosurus tristachyos Lam. 
NOTE: isonym. 
CORRECT NAME: Eleusine tristachya (Lam.) Lam. 
= Eleusine tristachya (Lam.) Lam. 


Eleusine tristachya fo. latifolia Hack., Bull. Herb. Boissier, 
ser. 2, 4(3): 280. 1904. T-Protol.: Paraguay: in campis 
in regione cursus superioris fluminis Apa, Feb, 
Hassler 8455. 

= Eleusine tristachya (Lam.) Lam. 


Eleusine tristachyos (Lam.) Lam., Tabl. Encycl. 1: 203. 1791. 
Basionym: Cynosurus tristachyos Lam. 
NOTE: orth. var. 
= Eleusine tristachya (Lam.) Lam. 


Eleusine unioloides Willd. ex Steud., Nomencl. Bot. (ed. 2) 
1: 549. 1840. 
NOTE: nom. inval., as syn. of Leptochloa virgata Pers. 
= Leptochloa virgata (L.) P. Beauv. 


Eleusine virgata (L.) Pers., Syn. Pl. 1: 87. 1805. Basionym: 
Cynosurus virgatus L.. 
= Leptochloa virgata (L.) P. Beauv. 


Eleusininae subtrib. Dumort., Anal. Fam. PI. 63. 1829. 
TYPE: Eleusine Gaertn. 
NOTE: as "Eleusineae". 
TRTD. BY: Soreng, R. J., G. Davidse, T. S. Filgueiras, E. J. 
Judziewicz, P. M. Peterson, and F. O. Zuloaga. 
NHT: Cynodonteae. 


78 Catalogue of New World Grasses: II. 


Enneapogon Desy. ex P. Beauy., Ess. Agrostogr. 81, 
161. 1812. TYPE: Enneapogon desvauxii P. Beauv. ex 
Desv.; LT designated by Hitchcock, U.S. Dept. Agric. 
Bull. 772: 83 (1920). 

TRTD. BY: Peterson, P. M. 

NHT: Pappophoreae. NO. ACCPTD. SPP.: 2. 

CNSLTD. LIT.: Renvoize, S. A. , (1968); Kakudidi, E. K, 
Z., M. Lazarides & J. A. Carnahan, (1988). 

SYN.: Calotheria Steud. 


Enneapogon brachystachyus (Jaub. & Spach) Stapf, Fl. Cap. 
7: 654. 1900. Basionym: = Pappophorum 
brachystachyum Jaub. & Spach. 

= Enneapogon desvauxii P. Beauv. 


Enneapogon desvauxii P. Beauv., Ess. Agrostogr. 82, 161, t. 
16, f. 11. 1812. T-Protol.: Manilia [error for Mendoza, 
Argentina], type not designated. T-Specim.: | of 1. P. 
Beauv., Ess. Agrost. t. 16, f 11 (1812) [LT], LT 
designated (as type) by Ali Chaudahry, Grass. Saudi 
Arabia 221 (1989). 

NOTE: validly published here because an illustration with 
analysis is included by P. Beauv. 

DST.: Arge, Boli, Chil, Mexi, Peru, USA. 

LVL. ACCPTC.:: 1; introduced. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Brako, L. & J. L. Zarucchi (1993); Correll, 
D.S. & M. C. Johnston (1970); Cronquist, A., A. H. 
Holmgren, N. H. Holmgren & J. L. Reveal (1977); 
Gibbs Russell, G. E, W. G. Welman, E. Reitief, K. L. 
Immelman, G. Germishuizen, B. J. Pienaar, M. van 
Wyk, and A. Nicholas (PRECIS) (1987); Gould, F. 
W. & R. Moran (1981); Hickman, J. C. (1993); 
Hitchcock, A. S. (1951); Marticorena, C. & M. 


Quezada (1985); McVaugh, R. (1983); Munz, P. A. 


& D. D. Keck (1959); Munz, P. A. (1974); Phillips, 
S. (1995); Renvoize, S. A. (1998): Shreve, F. & I. L. 
Wiggins (1964); Tovar, 0. (1986); Tovar, O. (1993); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O, Morrone, J. Pensiero & A. M. Cialdella 
(1994), 

Czerniaevia rupestris Turcz. ex Kom.; Enneapogon 
brachystachyus (Jaub, & Spach) Stapf; Enneapogon 
desvauxii P. Beauv. ex Desv.; Enneapogon 
lophotrichus Chiov, ex Chiarugi; Enneapogon 
phleioides Roem. & Schult.; Enneapogon wrightii (S. 
Watson) C.E. Hubb.; Enneapogon wrightii (S. 
Watson) Roshev.; Pappophorum boreale C.C. Gmel. 
ex B. Fedtsch.; Pappophorum boreale Griseb.; 
Pappophorum  brachystachyum Jaub. & Spach; 
Pappophorum — brachystachyum 
Pappophorum fasciculatum Chiov.; Pappophorum 
figarianum Fig. & De Not; Pappophorum 
Jaminianum Coss. & T. Durand &  Boiss.; 
Pappophorum mexicanum Griseb. ex E. Fourn.; 
Pappophorum mexicanum Griseb. ex  Hemsl.; 
Pappophorum ~~ nanum — Steud.; = Pappophorum 
phleoides Turez.; Pappophorum pusillum (Rendle) 
K.  Schum.; Pappophorum vincentianum  J.A. 
Schmidt; Pappophorum  wrightii §S. Watson; 
Raspailia rupestris Turcz. ex Kom. 


SYN.: 


Enneapogon desvauxii P. Beauv. ex Desv., Mem. Soc. Agric. 
Angers 1: 202. 1831. T-Protol.: Argentina: Mendoza, 


Nee s.n. T-Specim.: 1 of 1. Manilia [error for 
Argentina: Mendoza, Neé [HT: MA; IT: BM (ex 


Jaub. & Spach; 


MA)], see Burbidge, Proc. Linn. Soc. London 57 
(1940-41). 
NOTE: hom. illeg.; also in Opusc. Sci. Phys. Nat. 98 
[reprint] 1831-1833. 
= Enneapogon desvauxii P. Beauv. 


Enneapogon lophotrichus Chiov. ex Chiarugi, Webbia 8: 

119. 1951. T-Specim.: 1 of 3. Fortino de Dande, | 
Aug, Chiovenda? 860 [ST]. 2 of 3. Fortino di Dande, | 
Aug, Chiovenda? 898 [ST]. 3 of 3. Elolo, 6 Aug, /73 
[ST]. 

NOTE: nom. nud. 

REPLACED NAME: Pappophorum brachystachyum Jaub. 

& Spach. 
= Enneapogon desvauxii P. Beauv. 


Enneapogon mollis Lehm., Nov. Stirp. Pug. 3: 40. 1831. T- 
Protol.: Ecuador: Habitat in Promontorio Bonar Spei, 
in terrae tractu, qui vocatur Beaufort (v. s.). T- 
Specim.: 1 of 1. South Africa, Eck/on s.n. [T]. 
DST.: USA. 
LVL. ACCPTC.: 1; introduced. 
SYN.: Pappophorum molle (Lehm.) Kunth, 


Enneapogon phleioides Roem. & Schult., Syst. Veg. 2: 616. 
1817. T-Protol.: Amer. Meridion[T: Herb. Zeae]. 
= Enneapogon desvauxii P. Beauv. 


Enneapogon wrightii (S. Watson) C.E. Hubb., Hooker's Icon. 
Pl. 34t: 3337. 1937. Basionym: Pappophorum wrightii 
S. Watson. 
NOTE: in adnot. 
= Enneapogon desvauxii P. Beauv. 


Enneapogon wrightii (S. Watson) Roshev., Zlaki SSSR 418. 
1937. Basionym: Pappophorum wrightii S. Watson. 
= Enneapogon desvauxii P. Beauv, 


Enteropogon Nees, Intr. Nat. Syst. Bot. 448. 1836. 
TYPE: Enteropogon melicoides(Rottler) Nees. 
TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 4. 
SYN.: Macrostachya A. Rich. 


Enteropogon brandegei (Vasey) Clayton, Kew Bull. 37: 
419. 1982. Basionym: Diplachne brandegei Vasey. 

DST.: Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY; Clayton, W. D. (1982); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000). 

SYN.: Chloris brandegei (Vasey) Swallen,; Diplachne 
brandegei Vasey; Gouinia  brandegei (Vasey) 
Hitche.; Leptochloa brandegei (Vasey) Hitche. 


Enteropogon chlorideus (J. Presl) Clayton, Kew Bull. 37: 
419. 1982, Basionym: Dinebra chloridea J, Pres). 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Clayton, W. D. (1982): Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Pohl, R. W. & G. Davidse 
(1994). 

Chloris chloridea (J. Presl) Hitche.; Chloris 
clandestina Scribn. & Merr.; Chloris longifolia (E. 
Fourn.) Vasey; Dinebra chloridea J. Presl; Eutriana 
chloridea (J. Presl) Kunth; Gymnopogon longifolius 
E. Fourn.; Gymnopogon virletii E. Fourn. 


SYN.: 


Catalogue of New World Grasses: II. 79 


Enteropogon fasciculatus Clayton & Renvoize, Kew Bull., 
Addit. Ser. 13: 239. 1986, 
NOTE: nom. inval., without description and without 
basionym reference. 
= Saugetia fasciculata Hitchc. & Chase. 


Enteropogon mollis (Nees) Clayton, Kew Bull. 37: 419. 
1982. Basionym: Gymnopogon mollis Nees. 
DST.: Braz, Cari, Colo, CoRi, Ecua, Guat, Hond, Nica, 
Peru, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Brako, L. & J. L. Zarucchi (1993); 
Catasus Guerra, L. (1997); Clayton, W. D. (1982); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Jorgensen, P. M. & S. Leén-Yanez 
(eds.) (1999); Pohl, R. W. & G. Davidse (1994); 
Renvoize, S. A. (1984); Tovar, O. (1993). 

Chloris angustiflora F. Aresch.; Chloris leptantha 
Hitche. ex Urb.; Chloris luetzelburgii Hitchc.; 
Chloris mollis (Nees) Swallen; Chloris rupestris 
(Ridl.) Hitche.; © Gymnopogon mollis Nees; 
Gymnopogon rupestre Ridl. 


SYN:: 


Enteropogon prieurii (Kunth) Clayton, Kew Bull. 37: 419. 
1982. Basionym: Chloris prieuriiKunth. 

DST.: USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Clayton, W. D. (1982); Gibbs 
Russell, G. E, W. G. Welman, E. Reitief, K. L. 
Immelman, G. Germishuizen, B. J. Pienaar, M. van 
Wyk, and A. Nicholas (PRECIS) (1987); Phillips, S. 
(1995). 

Chloris cryptostachya Steud. ex J.A. Schmidt; 
Chloris prieurii Kunth; Chloris punctulata Hochst. 
ex Steud.; Chloris subtriflora Steud. 


Epicampes J. Presl, Relig. Haenk. 1(4-5): 235. 1830. TYPE: 
Epicampes stricta J. Presl. 
= Muhlenbergia Schreb. 


SYN:: 


Epicampes anomala Scribn. ex Beal, Grass. N. Amer. 2: 311. 
1896. T-Protol.: Mexico: Chihuahua, 11 Oct 1887, 
Pringle 1423. T-Specim.: 1 of 1. Mexico: Chihuahua: 
Arroyo Aucho, Sierra Madre, 11 Oct 1887, C.G. 
Pringle 1423 (HT: MSC; IT: US-746682, US-155141, 
US-822729]. 

= Muhlenbergia longiligula Hitche. 


Epicampes argentea (Vasey) M.E. Jones, Contr. W. Bot. 14: 
7. 1912. Basionym: Muhlenbergia argentea Vasey. 
= Muhlenbergia argentea Vasey. 


Epicampes arundinacea (Griseb.) Hack. ex Stuck., Anales 
Mus. Nac. Buenos Aires 11: 107. 1904. Basionym: 
Diachyrium arundinaceum Griseb. 

NOTE: basionym erroneously attributed to Trin. 
= Sporobolus rigens (Trin.) Desv. 


Epicampes berlandieri E. Fourn. ex Hemsl., Biol. Cent.- 
Amer., Bot. 3: 548. 1885. T-Protol.: Mexico, Berl. 
670. 
NOTE: nom. nud. 
= Muhlenbergia robusta (E. Fourn.) Hitchce. 


Epicampes berlandieri E. Fourn., Mexic. Pl. 2: 89. 1886. T- 
Specim.: | of 5. Mexico, 1839, JL. Berlandier 670 
[ST: US-1127013]. 2 of 5. Mexico: Orizaba, F. Mill. 
2130 [ST: P]. 3 of 5. Mexico: Orizaba, Bort. 102 [ST: 
P]. 4 of 5. Mexico: Orizaba, Schaffner 169 [ST: "herb. 


Frang."]. 5 of 5. Mexico: pr. Tacubaya, Jun, Schaffner 
315 (ST: "herb. Franq."]. 
= Muhlenbergia robusta (E. Fourn.) Hitchc. 


Epicampes bourgaei E. Fourn. ex Hemsl., Biol. Cent.-Amer., 
Bot. 3: 548. 1885. 
NOTE: nom. nud. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Epicampes bourgeaei E. Fourn., Mexic. Pl. 2: 88. 1886. T- 
Protol.: Mexico: Escamela pr. Orizaba, Aug, Bourgeau 
2973. T-Specim.: | of 1. Mexico: Veracruz: Escamala: 
Refrou D'Orizaba, 26 Aug 1866, E.Bourgeau 2973 
[IT: US-A0865984 (fragm.)]. 
NOTE: as Epicampes "bourgaei". 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Epicampes bourgeaei (E. Fourn.) M.E. Jones, Contr. W. Bot. 
14: 7. 1912. Basionym: Muhlenbergia bourgeaei E. 
Fourn. 

NOTE: hom. illeg. 
E. NAME: Epicampes bourgeaei (E. Fourn.) M.E. Jones. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Epicampes bourgeaei E. Fourn. var. bourgeaei. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Epicampes bourgeaei var. grandis (Vasey) M.E. Jones, 
Contr. W. Bot. 14: 7. 1912. Basionym: Muhlenbergia 
grandis Vasey. 

= Muhlenbergia grandis Vasey. 


Epicampes bourgeaei var. mutica E. Fourn., Mexic. Pl. 2: 88. 
1886. T-Specim.: 1 of 3. Mexico: Mirador, Oct-Nov, 
Liebmann 676 {ST: IST: MO-1837818, MO-1837819]. 
2 of 3. Mexico: Mirador, Oct-Nov, Liebmann 678 
[ST]. 3 of 3. Mexico: Tlaltengo, Oct, Liebmann 677 
(ST: IST: MO-1837820]. 

CORRECT NAME: Epicampes bourgeaei E. Fourn. 
= Muhlenbergia gigantea (E. Fourn.) Hitchc. 


Epicampes buchingeri E. Fourn. ex Hemsl., Biol. Cent.- 
Amer., Bot. 3: 548. 1885. 
NOTE: nom. nud. 
= Epicampes buchingeri E. Fourn. 


Epicampes buchingeri E. Fourn., Mexic. Pl. 2: 88. 1886. T- 
Protol.: Mexico: Orizaba, Thomas s.n. [HT: Herb. 
Buchinger], (Type not found by Chase & Niles, Index 
Grass Sp.). 

NOTE: Apparently an annual Muhlenbergia. 
SYN.: Epicampes buchingeri E. Fourn. ex Hemsl. 
= Status not yet determined. 


Epicampes clomena M.E. Jones, Contr. W. Bot. 14: 7. 1912. 
NOTE: nom. illeg.; based on Muhlenbergia clomema 
Kunth, a superfl. name for M. peruviana (P.Beauv.) 
Steud. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Epicampes coerulea Griseb., Abh. Konigl. Ges. Wiss. 
Gottingen 19: 256. 1874. T-Protol.: Argentina: 
Tucuman, ad vias pr. Anfama, S. de Aconquija in M. 
Cuesta de Juntas. T-Specim.: | of 2. Argentina: 
Tucuman, Lorentz 85 [ST: IST: US]. 2 of 2. Tucuman, 
Lorentz 86 [ST: IST: US]. 

= Muhlenbergia angustata (J. Pres!) Kunth. 


Epicampes coerulea var. submutica Hack. ex Stuck., Anales 
Mus. Nac. Buenos Aires 13: 471. 1906. T-Protol.: 


80 Catalogue of New World Grasses: II. 


Argentina: Tucuman: Cuesta de Malamala, Dep. Tafi, 
Stuckert herb. Arg. 14900 ex Lillo 3402. 
= Muhlenbergia angustata (J. Presl) Kunth. 


Epicampes crassiculmis Piper, Proc. Biol. Soc. Wash. 18: 
144. 1905. T-Protol.: Mexico: Sonora: in alkalai soil at 
Alamos, mouth of Yaqui River, Apr 1890, £. Palmer 
414 (HT: US]. T-Specim.: | of 1. Mexico: Sonora, Apr 
1890, E. Palmer 414 (HT: US-81870]. 

= Sporobolus wrightii Munro ex Scribn. 


Epicampes distichophylla Vasey, Descr. Cat. Grass. U.S. 45. 
1885. 
NOTE: nom. nud.; cites "Muhlenbergia, Munro" which may 
be M. distichophylla (Presl) Kunth, but misapl. by 
Vasey. 
= Muhlenbergia emersleyi Vasey. 


Epicampes distichophylla var. mutica Scribn., Grass. N. 
Amer. 2: 308. 1896. T-Protol.: USA: Arizona, Toumey 
740, (Mexican specimens cited do not agree with the 
type] Accd. to Chase & Niles. 
= Muhlenbergia longiligula Hitchc. 


Epicampes ehrenbergii Mez, Repert. Spec. Nov. Regni Veg. 
17: 212. 1921. T-Protol.: Mexico: Cuesta de Pinolco, 
Ehrenberg [1156]. T-Specim.: 1 of 1. Mexico: Cuesta 
de Pinolco, C. Ehrenberg 1156 [IT: US-A0865980 
(fragm.)]. 

= Muhlenbergia gigantea (E. Fourn.) Hitche. 


Epicampes emersleyi (Vasey) Hitche., U.S.D.A. Bull. (1915- 
23) 772: 144-145. 1920. Basionym: Muhlenbergia 
emersleyi Vasey. 

= Muhlenbergia emersleyi Vasey. 


Epicampes expansa E. Fourn. ex Hemsl., Biol. Cent.-Amer., 
Bot. 3: 548. 1885. 
NOTE: nom. nud. 
= Muhlenbergia gigantea (E. Fourn.) Hitche. 


Epicampes expansa E. Fourn., Mexic. Pl. 2: 88. 1886. T- 
Protol.: Mexico: Orizaba, Bort. 104 part. [HT: P]. T- 
Specim.: | of 1. Mexico: Orizaba, Bofteri & 
Sumichrast 104 {IT: US-865979 (fragm.)]. 

= Muhlenbergia gigantea (E. Fourn.) Hitche. 


Epicampes gigantea E. Fourn. ex Hemsl., Biol. Cent.-Amer., 
Bot. 3: 548. 1885. 
NOTE: nom. nud. 
= Muhlenbergia gigantea (E. Fourn.) Hitche. 


Epicampes gigantea E. Fourn., Mexic. Pl. 2: 88. 1886. T- 
Protol.: Mexico: Rio Blanco pr. Orizaba, Sep, 
Bourgeau 3137, T-Specim.: | of 1. Mexico: Orizaba: 
Rio Blanco, 30 Sep 1886, E.Bourgeau 3137 [HT: P; 
IT: US-865978 (fragm.), US-865977 (fragm.)]. 

= Muhlenbergia gigantea (E. Fourn.) Hitchc. 


Epicampes gracilis Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(3-4): 271. 1841. T-Protol.: Mexico [part now USA, 
Texas]. T-Specim.: | of 1. [USA]: Texas oriental, Dec 
1870, s.c. s.n. PHT: LE-TRIN-1556.01]. 

= Muhlenbergia lindheimeri Hitche. 


Epicampes grandis (Vasey) Beal, Grass. N. Amer. 2: 309, 
1896. Basionym: Muhlenbergia grandis Vasey. 
= Muhlenbergia grandis Vasey. 


Epicampes kunthiana Griseb., Abh. Konigl. Ges. Wiss. 
Gottingen 19: 256. 1874. 
NOTE: nom. nov., Based on Cinna stricta Kunth, this based 
on "Crypsis stricta Humb. & Kunth". 
REPLACED NAME: Crypsis stricta Kunth. 
BLOCKING NAME: Epicampes stricta J. Presl. 
= Muhlenbergia angustata (J. Pres!) Kunth. 


Epicampes lanata (Kunth) J. Presl ex Kunth, Enum. Pl. 1: 
209. 1833. Basionym: Agrostis lanata Kunth. 
= Muhlenbergia pubescens (Kunth) Hitchce. 


Epicampes laxiuscula E, Fourn. ex Hemsl., Biol. Cent.- 
Amer., Bot. 3: 549. 1885. 
NOTE: nom. nud. 
= Muhlenbergia gigantea (E. Fourn.) Hitchce. 


Epicampes laxiuscula E. Fourn., Mexic. Pl. 2: 88. 1886. T- 
Specim.: | of 2. Mexico: Orizaba, Botteri 1/55 [ST: P: 
IST: US-865975 (fragm.)]. 2 of 2. Mexico: Orizaba, 
Weber s.n. (ST: hb. Buchinger]. 
= Muhlenbergia gigantea (E. Fourn.) Hitche. 


Epicampes leptoura Piper, Proc. Biol. Soc. Wash. 18: 143. 
1905. T-Protol.: Mexico: Chihuahua: near Colonia 
Garcia, in the Sierra Madre Mts., 7000 fit, 21 Sep 1899, 
C.H. Townsend & C.M. Barber 341 [HT: US]. T- 
Specim.: | of 1. Mexico: Chihuahua: collected in the 
Sierra Madres near Colonia Garcia, 2134m, 21 Sep 
1899, CHT. Townsend & C.M. Barber 341] [HT: US- 
347144; IT: MO-1837827, US-347144, US-998992]. 

= Muhlenbergia rigens (Benth.) Hitchce. 


Epicampes ligulata Scribn. ex Vasey, Contr. U.S. Natl. Herb, 
3(1): 58. 1892. T-Specim.: | of 2. USA: Arizona, 23 
Jul 1892, JW. Toumey s.n. [ST: US-A865974 
(fragm.)]. 2 of 2. USA: Arizona, 26 Jul 1884, C.G. 
Pringle s.n. [LT: US-746686; ILT: F, GH, US-87018], 
LT designated by Hitchcock, Amer. J. Bot. 21: 136 
(1934). 

= Muhlenbergia longiligula Hitchce. 


Epicampes macrotis Piper, Proc. Biol. Soc. Wash. 18: 144, 
1905. T-Protol.: Mexico: Zacatecas: Sierra Madre 
Mts., 7 Aug 1897, JN. Rose 3528 [HT: US]. T- 
Specim.: | of 1. Mexico: Zacatecas: Sierra Madre 
Montains, ca. 40 km W of San Juan Capistrano, 7 Aug 
1897, J.N. Rose 3528 [HT: US-302505]. 

= Muhlenbergia robusta (E. Fourn.) Hitche. 


Epicampes macroura (Kunth) Benth., J. Linn. Soc., Bot. 19: 
87. 1881. Basionym: Crypsis macroura Kunth. 
= Muhlenbergia macroura (Kunth) Hitche. 


Epicampes mexicana (Scribn.) M.E. Jones, Contr. W. Bot. 
14: 7. 1912. Basionym: Bealia mexicana Scribn. 
= Bealia mexicana Scribn. ex Beal. 


Epicampes minutiflora Mez, Repert. Spec. Nov. Regni Veg. 
17: 212. 1921. T-Protol.: Mexico: Michoacan bei El 
Canizal, Langlassé 750. T-Specim.: 1 of 1. Mexico: 
Michoacan: near El Canizal, 15 Jan 1899, E. 
Langlassé 750 (IT: US-386160]. 

= Muhlenbergia robusta (E. Fourn.) Hitche. 


Epicampes mutica Rupr., Bull. Acad. Roy. Sci. Bruxelles 
9(2): 230. 1842. 
NOTE: nom. nud. 
= Muhlenbergia mutica (Rupr. ex E. Fourn.) Hitche. 


Catalogue of New World Grasses: II. 8] 


Epicampes mutica Rupr. ex E. Fourn., Mexic. Pl. 2: 87. 1886. 
T-Specim.: | of 4. Mexico: secus sylvulas in practis 
calidis Mirador, Zacuapan, Cantaranas, 1000-3500", 
Galeotti 5797 [LT: P], LT designated by Soderstrom, 
Contr. U.S. Natl Herb. 34: 141 (1967). 2 of 4. 
Mexico: Orizaba, Bott. 1/04 in part [ST]. 3 of 4. 
Mexico: Orizaba, Bott. ///] [ST: herb. Coss. & Franq.]. 
4 of 4. Mexico: Orizaba, Thomas [ST: herb. 
Buchinger]. 

NOTE: cites "E. mutica Rupr." nom. nud. 
= Muhlenbergia mutica (Rupr. ex E. Fourn.) Hitche. 


Epicampes phleoides (Kunth) Griseb., Abh. Konig]. Ges. 
Wiss. Gottingen 19: 256. 1874. Basionym: Crypsis 
phleoides Kunth. 

= Muhlenbergia angustata (J. Presl) Kunth. 


Epicampes pubescens (Kunth) J. Presl ex Kunth, Enum. PI. 1: 
209. 1833. Basionym: Agrostis pubescens Kunth. 
= Muhlenbergia pubescens (Kunth) Hitchce. 


Epicampes pubescens (Kunth) M.E. Jones, Contr. W. Bot. 
14: 7. 1912. Basionym: Agrostis pubescens Kunth. 
= Muhlenbergia pubescens (Kunth) Hitchce. 


Epicampes rigens Benth., J. Linn. Soc., Bot. 19: 88. 1881. T- 
Protol.: USA: California, H.N. Bolander 6124. T- 
Specim.: | of 1. USA: California: Sonoma, 1860-1867, 
H.N. Bolander 6124 \YT: US-323974, US-323975]. 

NOTE: cites Cinna macroura Thurb., as misaplication of C. 
macroura (Kunth) Kunth. 
= Muhlenbergia rigens (Benth.) Hitchc. 


Epicampes rigens (Trin.) Phil., Anales Mus. Nac. Santiago de 
Chile 8: 82. 1891. Basionym: Vilfa rigens Trin. 
NOTE: hom. illeg. 
E. NAME: Epicampes rigens Benth. 
= Sporobolus rigens (Trin.) Desv. 


Epicampes robusta E. Fourn. ex Hemsl., Biol. Cent.-Amer., 
Bot. 3: 550. 1885. 
NOTE: nom. nud. 
= Muhlenbergia robusta (E. Fourn.) Hitche. 


Epicampes robusta E. Fourn., Mexic. Pl. 2: 89. 1886. T- 
Specim.: 1 of 3. Mexico: Distrito Federal: Santa Fe, 2 
Oct 18?, M. Bourgeau 1153 (LT: P; ILT: US-999036, 
US-999031  (fragm.), US-90734 (fragm.)], LT 
designated by Hitchcock, N. Amer. Fl. 17(6): 462 
(1935). 2 of 3. Mexico: supra Tacubaya, E.Bourgeau 
683 [ST: P; IST: US-999030 (fragm.)]. 3 of 3. Mexico: 
Cuernavaca, Hahn 420 [ST: P]. 

= Muhlenbergia robusta (E. Fourn.) Hitchce. 


Epicampes speciosa (Vasey) M.E. Jones, Contr. W. Bot. 14: 
7.1912. Basionym: Muhlenbergia speciosa Vasey. 
= Muhlenbergia speciosa Vasey. 


Epicampes stricta J. Presl, Relig. Haenk. 1(4-5): 235, t. 39. 
1830. T-Protol.: Mexico: Hab. in Mexico, Haenke s.n. 
T-Specim.: | of 1. Mexico, 7. Haenke (HT: PR; IT: 
LE-TRIN-1558.01 (fragm. & — fig.), US-865970 
(fragm.)]. 

NOTE: as Epicampes "strictus”. 
= Muhlenbergia robusta (E. Fourn.) Hitchce. 


Epicampes stricta vat. distichophylla (J. Presl) M.E. Jones, 
Contr. W. Bot. 14: 6. 1912. Basionym: Podosemum 
distichophyllum J. Presl. 

= Muhlenbergia distichophylla (J. Presl) Kunth. 


Epicampes stricta var. mutica (Scribn.) M.E. Jones, Contr. 
W. Bot. 14: 6. 1912. Basionym: Epicampes 
distichophylla var. mutica Scribn. 

= Muhlenbergia longiligula Hitchc. 


Epicampes subpatens Hitche., U.S.D.A. Bull. (1915-23) 772: 
144. 1920. T-Protol.: USA: New Mexico: collected 
near Queen, in a rocky ravine, Guadelupe Mountains, 
7000 ft, 5 Sep 1915, Hitchcock 13541 [HT: US- 
905799]. T-Specim.: | of 1. USA: New Mexico: 
Ranger Station, Queen, Guadeloupe Mountains, 3 Sep 
1915-6 Sep 1915, A.S. Hitchcock 13541 [Amer. Gr. 
Herb. 1382] (HT: US-905799; IT: LE, US-3215629, 
US-3278409]. 

= Muhlenbergia emersleyi Vasey. 


Epicampes virletii E. Fourn., Mexic. Pl. 2: 88. 1886. T- 
Protol.: Mexico: San Luis de Potosi, Virlet d'Aoust 
1415. T-Specim.: | of 1. Mexico: San Luis de Potosi, 
Virlet d'aoust 1415 (HT: P; IT: US-73225]. 

= Muhlenbergia virletii (E. Fourn.) Soderstr. 


Eragrostideae trib. Stapf, Fl. Cap. 7: 316. 1898. TYPE: 
Eragrostis Wolf. 


Eragrostidinae subtrib. J. Presl, Relig. Haenk. 1(4-5): 273. 
1830. TYPE: Eragrostis Wolf. 
NOTE: as "Eragrostideae". 


Eragrostis Wolf, Gen. Pl. 23. 1776. TYPE: Eragrostis 
minor Host; LT designated by Ross, Acta Bot. Neer. 
15: 157 (1966). 
TRTD. BY: Peterson, P. M., and S. C. Boechat. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 110 SUBSP.: 25. 
CNSLTD. LIT.: Harvey, L. H., (1948); Nicora, E. G., 
(1998); Boechat, S. C. & H. M. Longhi-Wagner, 
(2000); Van den Borre, A. & L. Watson, (1994). 
Acamptoclados Nash; Boriskellera  Terechov; 
Diandrochloa De Winter; Erochloe Raf.; Erosion 
Lunell; Exagrostis Steud.; Macroblepharus Phil.; 
Neeragrostis Bush; Poa UNRANKED Psilantha K. 
Koch; Psilantha (K. Koch) Tzvelev; Roshevitzia 
Tzvelev; Stiburus Stapf, Thellungia Stapf; Triphlebia 
Stapf, Vilfagrostis DOll. 


SYN.: 


Eragrostis abessinica (Jacq.) Link, Hort. Berol. 1: 192. 1827. 
Basionym: Poa abyssinica Jacq. 
NOTE: orth. var. "abyssinica". 
CORRECT NAME: Eragrostis abyssinica (Jacq.) Link. 
= Eragrostis tef (Zucc.) Trotter. 


Eragrostis abyssinica (Jacq.) Link, Hort. Berol. 1: 192. 1827. 
Basionym: Poa abyssinica Jacq. 
NOTE: as Eragrostis "abessinica", 
= Eragrostis tef (Zucc.) Trotter, 


Eragrostis acicularis Trin, Mem, Acad. Imp. Sci. St- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 406. 1830. T- 
Protol.: V. subsp. Brasilia. (in arenosis ad ripas fluv. 
Jacare Pr. Parana), Riedel s.n. T-Specim.: 1 of 1. 
Brazil: in arenos. ad rip. fluv. Rio Jacare pr. Parauna, 
Dec 24, Riedel 1140 [HT: LE-TRIN-2299.01; IT: K, 
LE, P, SI, US-2891442, US-2765424 (fragm.)]. 

= Eragrostis rufescens var. rufescens. 


Eragrostis acuminata Doll, Fl. Bras. 2(3): 153. 1878. T- 
Protol.: Brazil: in prov. Piauhy, Gardner 2338 [IT: kK]. 
= Eragrostis maypurensis (Kunth) Steud. 


82 Catalogue of New World Grasses: II. 


Eragrostis acuta Hitche., Proc. Biol. Soc. Wash. 41: 159, 
1928. T-Protol.: Collected at Punta Rassa, Florida, Jul 
1990, A.S. Hitchcock 263 |[HT: US-731236]. T- 
Specim.: | of 1. USA: Florida: Punta Rassa, Jul-Aug 
1900, A.S. Hitchcock 263 |HT: US-731236; IT: US- 
1503824]. 
= Eragrostis elliottii S. Watson. 


Eragrostis acutiflora (Kunth) Nees, Fl. Bras. Enum. Pl. 2: 
501-502. 1829. Basionym: Poa acutiflora Kunth. 
DST.: Beli, Boli, Braz, Cari, Colo, CoRi, Ecua, ElSa, FrGu, 

Guat, Guya, Hond, Mexi, Nica, Pana, Suri, Vene. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Balick, M. J., M. H. Nee & D. E. 
Atha (2000); Beetle, A. A. (1977); Boechat, S.C. & 
H. M. Longhi-Wagner (2000); Brako, L. & J. L. 
Zarucchi (1993); Breedlove, D. E. (1986); Cowan, C. 
P. (1983): Davidse, G. (1994): Davidse, G., M. Sousa 
S. & A. O. Chater (editores generales) (1994); Doll, 
J. C. (1878); Espejo Serna, A., A. R. Lopez-Ferrari, 
and J. Valdes-Reyna (2000); Gould, F. W (1979); 
Harvey, L. H. (1984); Hitchcock, A. S. (1936); 
Jorgensen, P. M. & S. Leén-Yanez (eds.) (1999); 
Judziewicz, E. J. (1990); Macbride, J. F. (1936); 
McClure, F. A. (1955); McVaugh, R. (1983); Molina 
R., A. (1975); Nelson, C. (1978); Pohl, R. W. (1980); 
Renner, S. S., H. Balsley & L. B. Holm-Nielsen 
(1990); Renvoize, S. A. (1998); Reyes-Garcia, A. & 
M. Sousa S. (1997). 

Eragrostis acutiflora var. humilior J. Presl; 
Eragrostis flavescens J, Presl; © Megastachya 
acutiflora (Kunth) Roem. & Schult.; Poa acutiflora 
Kunth. 


SYN.:: 


Eragrostis acutiflora var. humilior J. Presi, Relig. Haenk. 
1(4-5): 277. 1830. T-Protol.: Hab. in Panama, Haenke 
SN. 
= Eragrostis acutiflora (Kunth) Nees. 


Eragrostis acutiglumis Parodi, Notas Mus. La Plata, Bot. 
2(Bot. Il): 7, f. 3. 1937. T-Protol.: Argentina: 
Corrientes: Mercedes, L.R. Parodi 6168. T-Specim.: | 
of 1. Argentina: Corrientes: Mercedes, 5 Feb 1925, 
L.R. Parodi 6168 (HT: BAA; IT: L, US-1256543]. 

DST.: Arge, Braz, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Burkart, A. (1969); Nicora, E. G. 
(1998); Rosengurtt, B., B. R. Arrillaga de Maffei & 
P. Izaguirre de Artucio (1970). 


Eragrostis affinis Salzm. ex Steud., Syn. Pl. Glumac. 1: 277. 
1854. T-Protol.: Bahia, Sa/zmann s.n., cites: Salzm. 
(Hrbr. Bahia sub: Poa.). T-Specim.: | of 1. Brazil: 
Bahia: in maritimis, 1838, 7) Salzmann s.n. (HT: LE- 
TRIN-2572.01; IT: K, LE-TRIN-2572.02, MO, US- 
2767409, US-911819, US-1614948]. 

NOTE: cites Eragrostis affinis Salzm., herb. Bahia sub: 
Poa, nom. nud. 
= Eragrostis rufescens var. rufescens. 


Eragrostis airoides Nees, Fl. Bras. Enum. Pl. 2: 509-510. 
1829. 
NOTE: nom. nov., Nees elected to rename Aira brasiliensis 
and recombine Poa brasiliensis, both Raddii names 
of 1823, 
REPLACED NAME: Aira brasiliensis Raddi. 
BLOCKING NAME: Eragrostis brasiliensis (Raddi) Nees. 
DST.: Arge, Boli, Braz, Cari, Colo, Para, USA, Urug, Vene. 


LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Boechat, $. C. & H. M. Longhi- 
Wagner (2000); Burkart, A. (1969); Cabrera, A. L. 
(1970); Doll, J. C. (1878); Judziewiez, E. J. (1990); 
Killeen, T. J. (1990); Longhi-Wagner, H. M. & I. I. 
Boldrini (1988); Nicora, E. G. (1998); Nicora, E. G., 
M. E. De Paula, A. M. Faggi, M. del C. Mariano, A. 
M. Miante A., L. R. Parodi, C. A. Petetin, F. A. Roig 
& Z. Rugolo de Agrasar (1978); Renvoize, S. A. 
(1988); Renvoize, S. A. (1998); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 
Smith, L. B., D. C. Wasshausen & R. M. Klein 
(1981). 

Agrosticula brasiliensis (Raddi) Herter; Aira 
brasiliensis Raddi; Airopsis millegrana  Griseb.; 
Eragrostis airoides var. pluriflora Doéll, Eragrostis 
brasiliensis (Raddi) Nees; Eragrostis triflora Ekman; 
Poa airoides (Nees) Kunth; Poa brasiliensis Raddi; 
Sporobolus brasiliensis (Raddi) Hack.; Sporobolus 
ramosissimus Kunth. 


SYN:: 


Eragrostis airoides var. pluriflora Doll, Fl. Bras. 2(3): 137. 
1878. T-Protol.: In Brasilia meridionali lecta, Se//o s.n. 
= Eragrostis airoides Nees. 


Eragrostis alba J. Presl, Relig. Haenk. 1(4-5): 279. 1830. T- 
Protol.; Monte-rey, Californiae, Haenke s.n., according 
to A.S.Hitchcock in Index of Grass Species probably 
collected in Peru, not California. T-Specim.: | of 1. 
Peru: no date, 7 Haenke s.n. [HT: PR; IT: MO- 
2111118, US-2942409 (fragm. ex P)], US specimen a 
possible type. 

= Eragrostis mexicana (Hornem.) Link. 


Eragrostis amabilis (L.) Wight & Arn. ex Nees, Bot. 
Beechey Voy. 251. 1838. Basionym: Poa amabilis L. 
NOTE: in Hooker and Arnott. 
DST.: Beli, Boli, Braz, Cari, CoRi, Ecua, ElSa, FrGu, Guat, 
Guya, Hond, Mexi, Nica, Pana, Para, Peru, Suri, 
USA, Vene. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Balick, M. J.. M. H. Nee & D. E. 
Atha (2000); Boechat, S.C. & H. M. Longhi-Wagner 
(2000); Brako, L. & J. L. Zarucchi (1993); Correll, 
D.S. & M. C. Johnston (1970); Davidse, G. (1994); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Hitchcock, A. S. 
(1927): Hitchcock, A. S. (1936); Hitchcock, A. S. 
(1951); Jorgensen, P. M. & S. Leon-Yanez (eds.) 
(1999); Judziewicz, E. J. (1990); Lindeman, J. C. & 
A. L. Stoffers (1963); McClure, F. A. (1955); Molina 
R., A. (1975); Radford, A. E., H. E. Ahles & C. R. 
Bell (1968); Renvoize, S. A. (1998): Small, J. K. 
(1933); Wunderlin, R. P. (1998), 

Cynodon amabilis (L.) Raspail; Eragrostis amabilis 
var. plumosa (Retz.) E.G. Camus & A. Camus; 
Eragrostis ciliaris var. patens Chapm. ex Beal; 
Eragrostis plumosa (Retz.) Link; Eragrostis tenella 
(L.) P. Beauv. ex Roem. & Schult.; Eragrostis tenella 
var. plumosa (Retz.) Stapf; Erochloe amabilis Raf. ex 
B.D. Jacks.; Megastachya amabilis (L.) P. Beauv.: 
Poa amabilis L.; Poa plumosa Retz.; Poa tenella L. 


SYN:: 


Eragrostis amabilis var. plumosa (Retz.) E.G. Camus & A. 
Camus, FI. Indo-Chine 7: 557. 1923. Basionym: Poa 
plumosa Retz. 


Catalogue of New World Grasses: II. 83 


= Eragrostis amabilis (L.) Wight & Arn. ex Nees. 


Eragrostis amoena J. Presl, Relig. Haenk. 1(4-5): 275. 1830. 
T-Protol.: Hab. in Mexico, Haenke s.n. T-Specim.: | 
of 1. Mexico, no date, 7) Haenke s.n. {HT: PR; IT: US- 
2850746 (fragm.)]. 

= Eragrostis maypurensis (Kunth) Steud. 


Eragrostis ancashensis P.M. Peterson, Refulio & Tovar, 
Sida 19(1): 66, f. 1. 2000. T-Protol.: Peru: Dptm. 
Ancash: Prov. Recuay: Cordillera Blanca, ca. 20 km E 
of Raquia on Rt. 02-014, on rd to Huaraz, 
10°08'55.8"S, 77°19'48.8"W, 3000 m, 20 Mar 1997, 
P.M. Peterson & N. Refulio Rodriguez 13793 [USM; 
IT: K, MO, NY, RSA, TAES, UC, US, WIS]. T- 
Specim.: | of 1. Peru: Dptm. Ancash: Prov. Recuay: 
Cordillera Blanca, ca. 20 km E of Raquia on Rt. 02- 
014, on road to Huaraz, 10°08'S5.8"S, 77°19'48.8"W, 
3000 m, along roadside with shrubby Asteraceae, 
Lupinus, Vicia, and Commelina, 20 Mar 1997, P.M. 
Peterson & N. Refulio Rodriguez 13793 [HT: USM; 
IT: K, MO-5156401, NY, RSA, TAES, UC, US, WIS]. 

DST.: Peru. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Peterson, P. M., N. Refulio R. & O. 
Tovar (2000). 


Eragrostis andicola R.E. Fr., Nova Acta Regiae Soc. Sci. 
Upsal., ser. 4, 1(1): 180. 1905. T-Protol.: Argentina: 
Jujuy: Moreno in montibus saxosis apricis siccis, 26 
Nov, 15 Dee 1901, Fries 924a, 924. T-Specim.: | of 2. 
Argentina: Prov. Jujuy: Moreno in montibus saxosis 
apreicis siccis, 3500-3700 m, 26 Nov 1901,Fries 924a 
[ST: BAA-998, CORD]. 2 of 2. Prov. Jujuy: Moreno 
in montibus saxosis apricis siccis, 15 Dec 1901, Fries 
924 [ST: BAA, CORD, US-2891444 (fragm.)]. 

DST.: Arge. 
LVL. ACCPTC:: I. 
ALSO ACCPTD. BY: Nicora, E. G. (1998). 


Eragrostis andicola Pilg., Bot. Jahrb. Syst. 37: 377. 1906. T- 
Protol.: Peru: Ancash: inter Samanco et Caraz, infra 
Hacienda Cajabamba, in formatione aperta, praecipue 
gramina intermixtis fruticibus gerente, 3000-3500 m, 
May 1903, Weberbauer 3114 (HT: B]. T-Specim.: | of 
1. Peru: Ancash: Hacienda Cajabamba, entre Samanco 
y Caraz, 27 May 1903, 4. Weberbauer 3114 (HT: B; 
IT: BAA-999 (fragm. ex B), US-2766205 (fragm.)]. 

NOTE: hom. illeg. 

E. NAME: Eragrostis andicola R.E. Fr. 
CORRECT NAME: Eragrostis pilgeri Fedde. 
= Eragrostis pilgeri Fedde. 


Eragrostis andicola fo. humilior Pilg., Bot. Jahrb. Syst. 37: 
377. 1906. T-Protol.: Peru: Ancash: inter Samanco et 
Caraz, infra Hacienda Cajabamba, in formatione 
aperta, 3700 m, May 1903, Weberbauer 3036. T- 
Specim.: | of 1. Peru: Ancash: Hacienda Cajabamba, 
entre Caraz y Samanco, 3700m, Weberbauer 3036 [IT: 
MOL]. 

= Eragrostis pilgeri Fedde. 


Eragrostis andicola var. robustior Pilg., Bot. Jahrb. Syst. 37: 
377. 1906. T-Protol.: Peru: Ancahs: Cajatambo: infra 
Ocros, in formatione aperta, fruticibus et plantis 
herbaceis mixta, 3000-3200 m, Weberbauer 2746. T- 
Specim.: | of 1. Peru: Ancash: Cajatambo Prov., no 
date, A. Weberbauer 2746 [HT: B; IT: US-2850747]. 


= Eragrostis pilgeri Fedde. 


Eragrostis annulata Chiov., Annuario Reale Ist. Bot. Roma 
8: 63. 1903. T-Protol.: Erythreae. T-Specim.: | of 2. 
Samhar: Isola Scek Said, 3 Mar 1892, Terracciano & 
Pappi 2730 [ST]. 2 of 2. Mista conErag. aspera Nees 
neil, isola Scek Said, 3 Mar 1892, Terracciano & 
Pappi 2732 [ST]. 

= Eragrostis cylindriflora Hochst. 


Eragrostis apiculata Doll, Fl. Bras. 2(3): 145. 1878. T- 
Protol.: Brasilia: Habitat ad Caldas prov. Minarum, 
Regnell II. 1411, T-Specim.: | of 1. Brazil: Minas 
Gerais, 25 Jun 1868, A.F. Regnell III, 141] [{IT: US- 
911759, US-2891446 (fragm.)]. 

DST.: Braz. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Dall, J. C. (1878). 


Eragrostis argentina Jedwabn., Bot. Arch. 5(3-4): 193. 1924. 
T-Protol.: Argentina: in Rio Colorado valle, ubi 
primum saxa granitica in lucem evadunt, et Rio Negro. 
super insulam Choele-Choel, Lorentz & Niederlein s.n. 
T-Specim.: | of 2. Argentina: Rio Colorado valle, ubi 
primum saxa granitica in lucem evadunt, Lorentz s.n. 
[ST: BAA-1001, CORD, SI, US-2850748 (fragm.)]. 2 
of 2. Argentina: Rio Negro, super insulum Choele- 
Choel, Niederlein s.n. [ST]. 

= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis arida Hitche., J. Wash. Acad. Sci. 23(10): 449. 
1933. T-Protol.: USA: Texas: Del Rio, 14 Sep 1915, 
A.S. Hitchcock 13650 [HT: US-905937]. T-Specim.: 1 
of 1. USA: Texas: Val Verde Co.: Del Rio, 14 Sep 
1915, A.S. Hitchcock 13650 [HT: US-905937]. 
= Eragrostis pectinacea var. miserrima (E. 
Reeder. 


Fourn.) 


Eragrostis articulata (Schrank) Nees, Fl. Bras. Enum. PI. 2: 
502-503. 1829. Basionym: Poa articulata Schrank. 
DST.: Arge, Boli, Braz, Para. 
LVL. ACCPTC:: 1. 
ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 
(1987); Boechat, S. C. & H. M. Longhi-Wagner 
(2000); Filgueiras, T. S. (1991); Filgueiras, T. S. 
(1999): Foster, R. C. (1958); Hitchcock, A. S. (1927); 
Killeen, T. J. (1990); Longhi-Wagner, H. M. (1990): 
Nicora, E. G. (1998); Renvoize, S. A. (1984); 
Renvoize, S. A. (1998); Smith, L. B., D. C. 
Wasshausen & R. M. Klein (1981). 
Eragrostis articulata var. eglandulosa Nicora; 
Eragrostis articulata. var. glabrescens Henrard; 
Eragrostis articulata var. pauciflora Doll; Eragrostis 
brasiliensis var. delta Nees; Eragrostis glareosa 
Trin.; Eragrostis neesii var. laxa  Jedwabn.: 
Eragrostis pilosissima Link; Eragrostis villosa 
Salzm. ex Steud.; Poa articulata Schrank. 


SYN.: 


Eragrostis articulata var. eglandulosa Nicora, Revista 
Argent. Agron. 7(4): 272, 259, f. 4C, 5B. 1940. T- 
Protol.: Paraguay: Guaira, Villarrica, P. Jérgensen 
3554 [HT: SI; IT: LP]. T-Specim.: | of 1. Paraguay: 
Guaira: Villa Rica: comtin campo, P. Jorgensen 3554 
[HT: BAA; IT: BA, LP, SI, US-1936682, US- 
1447249, US-1517041). 

= Eragrostis articulata (Schrank) Nees. 


84 Catalogue of New World Grasses: II. 


Eragrostis articulata var. glabrescens Henrard, Meded. 
Rijks-Herb. 40: 69. 1921. T-Protol.: Bolivia, Herzog 
/309a, T-Specim.: | of 1. Bolivia, no date, 7. Herzog 
1309A [IT: US-2767389, US-2850750]. 

= Eragrostis articulata (Schrank) Nees. 


Eragrostis articulata var. pauciflora Doll, Fl. Bras. 2(3): 142. 
1878. T-Protol.: sin. loc., of 13 specimens of the 
species cited none applying to the — var. 
weredistinguished. 

= Eragrostis articulata (Schrank) Nees. 


Eragrostis arundinacea Jedwabn., Bot. Arch. 5(3-4): 192. 
1924, T-Protol.: "Am. sept., Texas", anon. s.n. [HT: 
B]. 

= Tridens flavus (L.) Hitche. 


Eragrostis atherstonei Stapf, Fl. Cap. 7: 607. 1900. T-Protol.: 
Kalahari Region: Transvaal; near Lydenburg, 
Atherstone s.n. 

= Eragrostis trichophora Coss. & Durieu. 


Eragrostis atrovirens (Desf.) Trin. ex Steud., Nomencl. Bot. 
(ed. 2) 1: 562. 1840. Basionym: Poa atrovirens Desf. 

DST.: Beli, Boli, Cari, Guat, Mexi, Nica, Peru, USA, Vene. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Anonymous (1986); Balick, M. J., 
M. H. Nee & D. E. Atha (2000); Bor, N. L. (1960); 
Brako, L. & J. L. Zarucchi (1993); Breedlove, D. E. 
(1986); Catasus Guerra, L. (1997); Cowan, C. P. 
(1983); Davidse, G. (1994); Davidse, G., M. Sousa S. 
& A. QO. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Foster, R. C. (1958); Godfrey, R. K. & J. W. 
Wooten (1979); Koch, S. D. (1978); Nicora, E. G. 
(1998); Phillips, S. (1995); Tovar, O. (1993); 
Wunderlin, R. P. (1998). 

Eragrostis biformis (Kunth) Benth.; Eragrostis 
bromoides Jedwabn.;, Eragrostis chariis (Schult.) 
Hitche.; Eragrostis elegantula (Kunth) Nees ex 
Steud.; Poa atrovirens Desf.; Poa biformis Kunth; 
Poa chariis Schult. 


SYN.: 


Eragrostis attenuata Hitche., Contr. U.S. Natl. Herb. 24(8): 
340. 1927. T-Protol.: Peru: Arequipa: collected on 
sandy slopes of hills about 3 km back of Mollendo, 17 
November 1923, 4.8. Hitchcock 22424 [HT: US- 
1164909]. T-Specim.: | of |. Peru: Arequipa: on sandy 
slopes of hills ca. 3 km back of Mollendo, 17 Nov 
1923, AS. Hitchcock 22424 [HT: US-1164909; IT: 
BAA-1007, US-1 164361]. 

DST.: Chil, Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Hitchcock, A. S. (1927); Macbride, 
J. F. (1936); Marticorena, C. & M. Quezada (1985): 
Nicora, E. G. (1994); Nicora, E. G. (1998); Tovar, O. 
(1993). 

SYN.: Sporobolus scaber Phil. 


Eragrostis aturensis (Kunth) Trin. ex Steud., Nomencl. Bot. 
(ed. 2) 1: 562. 1840. Basionym: Poa aturensis Kunth. 
= Eragrostis japonica (Thunb.) Trin. 


Eragrostis bahamensis Hitchc., Annual Rep. Missouri Bot. 
Gard. 1893: 149, t. 14. 1893. T-Protol.: Bahamas: 
Inagua, growing on rocky shore, winter 1890-1891, 
Hitchcock s.n. T-Specim.: | of 1. Bahamas: Inagua, 3 


Dec 1890, A.S. Hitchcock s.n. [HT: MO-47402; IT: 
NY-70972, US-748406]. 

DST.: Cari. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Hitchcock, A. S. (1936). 


Eragrostis bahiensis Schrad. ex Schult., Mant. 2: 318. 1824. 
T-Protol.: Brasilia, Princeps Serenniss. Maximilian 
Neowid. s.n. T-Specim.: | of 1. Brazil, Maximilian 
Neowidensis [HT: LE]. 

DST.: Arge, Boli, Braz, Colo, Ecua, FrGu, Guat, Hond, 
Mexi, Para, Peru, USA, Urug, Vene. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 

(1987); Anonymous (1986); Beetle, A. A. (1977); 

Boechat, S$. C. & H. M. Longhi-Wagner (2000); 

Brako, L. & J. L. Zarucchi (1993): Breedlove, D. E. 

(1986); Burkart, A. (1969); Cabrera, A. L. (1970); 

Cowan, C. P. (1983); Davidse, G. (1994); Davidse, 

G., M. Sousa $. & A. O. Chater (editores generales) 

(1994); Doll, J. C. (1878); Espejo Serna, A., A. R. 

Lopez-Ferrari, and J. Valdes-Reyna (2000); 

Filgueiras, T. S. (1999); Foster, R. C. (1958); 

Godfrey, R. K. & J. W. Wooten (1979); Hitchcock, 

A. S. (1927); Hitchcock, A. S. (1951); Jorgensen, P. 

M. & S. Le6n-Yanez (eds.) (1999); Killeen, T. J. 

(1990); Koch, S. D. (1978); Long, R. W. & O. Lakela 

(1971); Longhi-Wagner, H. M. (1990); McClure, F. 

A. (1955); Nicora, E. G. (1998); Renner, S. S., H. 

Balslev & L. B. Holm-Nielsen (1990); Renvoize, S. 

A. (1984); Renvoize, S. A. (1988); Renvoize, S. A. 

(1998); Rosengurtt, B., B. R. Arrillaga de Maffei & 

P. Izaguirre de Artucio (1970); Small, J. K. (1933); 

Smith, L. B., D. C. Wasshausen & R. M. Klein 

(1981); Wunderlin, R. P. (1998). 

Eragrostis bahiensis fo. riparia Burkart,; Eragrostis 

bahiensis var. bahiensis, Eragrostis bahiensis var. 

contracta Doll; Eragrostis bahiensis var. laxiuscula 

Doll; = Eragrostis — blepharophylla — Jedwabn.; 

Eragrostis elatior Stapf; Eragrostis expansa Link; 

Eragrostis firma Trin., Eragrostis macra Jedwabn.; 

Eragrostis microstachya (Link) Link; Eragrostis 

pilosa var. bahiensis (Schrad. ex Schult.) Kuntze; 

Eragrostis psammodes Trin.; Eragrostis psammodes 

var, microstachya (Link) Doll; Poa expansa (Link) 

Kunth; Poa microstachya Link; Poa psammodes 

(Trin.) Kunth, 


SYN.: 


Eragrostis bahiensis Schrad. ex Schult. var. bahiensis. 
= Eragrostis bahiensis Schrad. ex Schult. 


Eragrostis bahiensis var. boliviensis Henrard, Meded. Rijks- 
Herb. 40: 68. 1921. T-Protol.: Bolivien: Prov. 
Larecaja, in viciniis Sorata, colle Ullontiji, undique, in 
aridis, 2900 m,Feb 1858, Mandon 1/332 [HT: L- 
908,88-303; IT: L-908,88-342]. T-Specim.: | of 1. 
Bolivia: La Paz: Larecaja: Sorata, 2900 m, Feb 1858, 
G. Mandon 1332 |HT: L-908,88-303; IT: K, L-908,88- 
342, P, US-3161127, US-256470, US-911775 
(fragm.)]. 

= Eragrostis lurida J. Presl. 


Eragrostis bahiensis var. contracta Doll, Fl. Bras. 2(3): 151. 
1878. T-Specim.: | of 3. Brazil: Rio de Janeiro, 
Weddell 175 |LT: BJ], LT designated by Scholz, 
Willdenowia 22: 268 (1992). 2 of 3. , Aug 1874,4. 


Catalogue of New World Grasses: II. 85 


Glaziou 6952 [ST: K]. 3 of 3. : Mar 1872,A4. Glaziou 
492 [ST: K]. 
= Eragrostis bahiensis Schrad. ex Schult. 


Eragrostis bahiensis var. laxiuscula DOll, Fl. Bras. 2(3): 151. 
1878. T-Specim.: | of 1. Brasilia meridionalis, F. 
Sellow s.n. [LT: B], LT designated by Scholz, 
Willdenowia 22: 268 (1992). 

= Eragrostis bahiensis Schrad. ex Schult. 


Eragrostis bahiensis fo. riparia Burkart, Fl. Il. Entre Rios 
6(2): 189. 1969. T-Specim.: 1 of 2. Argentina: Entre 
Rios: Concepcién del Uruguay, Isla Almirén Chico, 
Burkhart & Crespo 22874 [ST: BAA, SI]. 2 of 2. 
Argentina: Entre Rios: Orillas humedas, Burkhart 
22871] (ST: BAA, SI]. 
= Eragrostis bahiensis Schrad. ex Schult. 


Eragrostis barbata Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
4,2(1): 76. 1836. T-Protol.: V. subsp. Brasilia, Riedel 
s.n. T-Specim.: | of 4. Brazil, Jun 1822, s.c. s.n. [IT: 
LE]. 2 of 4. Brasilia, s.d., Riedel s.n. [IT: K, LE]. 3 of 
4. Bahia, 1831, Riedel [HT: LE-TRIN-2309.01]. 4 of 
4. Brazil, no date, L. Riedel s.n. [IT: US-911782, US- 
2891449 (fragm.)]. 

= Steirachne barbata (Trin.) Renvoize. 


Eragrostis barbiglumis Jedwabn., Bot. Arch. 5(3-4): 192. 
1924. T-Specim.: | of 3. Brazil: Rio de Janeiro: loco 
non indicato, A.f.M. Glaziou 15615 [ST: B; IST: C, 
US-2891450 (fragm.)]. 2 of 3. Brazil: Minas Gerais: 
ad Caldas, 4.F. Regnell s.n. [ST: ?; IST: US- 
A0865985 (fragm.)]. 3 of 3. Argentina: Misiones: Rio 
Alto Parana, 3 May 1883, W. Niederlein s.n. [ST: ?; 
IST: BAA-10I1 (fragm. ex B), US-A0865986 
(fragm.)]. 

= Eragrostis seminuda Trin. 


Eragrostis barrelieri Daveau, J. Bot. (Morot) 8: 289. 1894. 
T-Protol.: Plantae per Galliam, Hispaniam et Italiam 
observatae, Tab. 44, f. 2, type not indicated in the 
protologue. T-Specim.: | of 4. Egypt, Ascherson s.n. 
[ST: K]. 2 of 4. Algeria, Balansa 734 [ST: K]. 3 of 4. 
Italy: Sicily, herb. Todaro s.n. [ST: K]. 4 of 4. South 
Europe, Barrelier [ST]. 

DST.: Arge, Braz, Cari, Ecua, Mexi, USA, Urug. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Anonymous 
(1986); Boechat, S. C. & H. M. Longhi-Wagner 
(2000); Boechat, S. C. & P. M. Peterson (1995); Bor, 
N. L. (1960); Burkart, A. (1969); Cabrera, A. L. 
(1970); Cope, T. A. (1992); Correll, D. S. & M. C. 
Johnston (1970); Espejo Serna, A., A. R. L6pez- 
Ferrari, and J. Valdes-Reyna (2000); Gibbs Russell, 
G. E, W. G. Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987); Great Plains Flora 
Association (1986); Hickman, J. C. (1993); 
Hitchcock, A. S. (1936); Hitchcock, A. S. (1951); 
Jorgensen, P. M. & S. Leén-Yanez (eds.) (1999); 
Koch, S. D. (1978); Liogier, H. & L. Martorell 
(1982); Munz, P. A. & D. D. Keck (1959); Munz, P. 
A. (1974); Nicora, E. G. (1998); Phillips, S. (1995); 
Rosengurtt, B., B. R. Arrillaga de Maffei & P. 
Izaguirre de Artucio (1970); Wunderlin, R. P. (1998). 


SYN.: Eragrostis poaeoides var. barrelieri (Daveau) Fiori; 
Eragrostis vulgaris subsp. barrelieri (Daveau) 
R.C.V. Douin. 


Eragrostis berteroniana (Schult.) Steud., Nomencl. Bot. 
(ed. 2) 1: 562. 1840. Basionym: Megastachya 
berteroniana Schult. 

DST.: Cari. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Catasus Guerra, L. (1997); Catasus 
Guerra, L. J. (1980); Hitchcock, A. S. (1936), 

SYN.: Megastachya berteroniana Schult.; Poa berteroniana 
(Schult.) Kunth. 


Eragrostis beyrichii J.G. Sm., Annual Rep. Missouri Bot. 
Gard. 6: 117, t. 56. 1895. T-Protol.; USA: Arkansas: 
1834, Beyrich s.n. [HT: MO-Bernhardi Herb.]. 
= Eragrostis secundiflora J. Presl. 


Eragrostis biformis (Kunth) Benth., Niger Fl. 568. 1849. 
Basionym: Poa biformis Kunth. 
= Eragrostis atrovirens (Desf.) Trin. ex Steud. 


Eragrostis bleeseri Pilg., Notizbl. Bot. Gart. Berlin-Dahlem 
10(91): 112. 1927. T-Protol.: Nord-Australien: Port 
Darwin, offner Wald, May 1926, F.A.K. Bleeser 659. 

= Eragrostis cumingii Steud. 


Eragrostis blepharophylla Jedwabn., Bot. Arch. 5(3-4): 197. 
1924. T-Protol.: Brasilia: loco non indicato, Se//ow 
3688. T-Specim.: 1 of 1. Brazil, F.Se//lo 3688 [IT: US- 
2891454 (fragm.)]. 

= Eragrostis bahiensis Schrad. ex Schult. 


Eragrostis boliviensis Jedwabn., Bot. Arch. 5(3-4): 205. 
1824. T-Specim.: | of 2. Bolivia: ad Soratam prope 
San Pedro, G. Mandon /330 [ST: BAA-1014 (fragm. 
ex B), K, US-2767395 (fragm.)]. 2 of 2. Bolivia: ad 
Camataqui, Fiebrig 3266 [ST: B]. 
= Eragrostis pastoensis (Kunth) Trin. 


Eragrostis botterii E. Fourn., Mexic. Pl. 2: 115. 1886. T- 
Specim.: 1 of 2. Mexico: Orizaba, Bott. 107 [OM: P]. 
2 of 2. Mexico: S. Lorenzo, Dec, Liebmann 530 [OM: 
Pi. 
NOTE: nom. inval., as syn. of £. poaeoides Beauv. var. B. 
= Eragrostis soratensis Jedwabn. 


Eragrostis brachypodon Hack., Bot. Centralbl. 120: 548. 
1912. T-Protol.: Bolivia: La Paz, Buchtien s.n. 
= Eragrostis soratensis Jedwabn. 


Eragrostis brasiliana Nees, Fl. Bras. Enum. Pl. 2: 510-511. 
1829. T-Protol.: Brazil, based on Poa brasiliensis 
Spreng. (1825), non. Raddi (1825). 

NOTE: nom. inval., not accepted by the author, who states 
in the "Addenda et emandanda" p. 510. Nro. 14. pro 
Eragrostis brasiliana lege E. elegans."; Accd. to 
Chase & Niles, Index Grass Sp. equals £. elegans 
Nees; not £. brasiliensis (Raddi) Nees, Fl. Bras. 
Enum. Pl. 2: 497, Fl. Bras. Enum. PI. 2: 608. 

CORRECT NAME: Eragrostis elegans Nees. 

= Eragrostis japonica (Thunb.) Trin. 


Eragrostis brasiliensis (Raddi) Nees, Fl. Bras. Enum. PI. 2: 
497-499, 1829. Basionym: Poa brasiliensis Raddi. 
= Eragrostis airoides Nees. 


Eragrostis brasiliensis var. delta Nees, Fl. Bras. Enum. PI. 2: 
497. 1829. 


86 Catalogue of New World Grasses: II. 


NOTE: nom. inval. 
= Eragrostis articulata (Schrank) Nees. 


Eragrostis brevipedicellata A. Gray, Proc. Acad. Nat. Sci. 
Philadelphia 14: 336. 1862. 
NOTE: nom. inval., as syn. of Eragrostis curtipedicellata 
Buckley. 
= Eragrostis curtipedicellata Buckley. 


Eragrostis brizoides Schult., Mant. 2: 319. 1824. 
REPLACED NAME: Poa tenella Nutt. 
BLOCKING NAME: Poa tenella L. 
= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis bromoides Jedwabn., Bot. Arch. 5(3-4): 190. 
1924. T-Protol.: Africa tropica, Sudan centr., in 
paludibus flum. Bailli, Chevalier 8588 [HT: ?]. 

NOTE: hom. illeg. 
E. NAME: Eragrostis bromoides (Vahl) Steud. 
= Eragrostis atrovirens (Desf.) Trin. ex Steud. 


Eragrostis buchtienii Hack., Repert. Spec. Nov. Regni Veg. 
6: 157. 1908. T-Protol.: Bolivia: Stid-Yungas, 
Simpayas bei Yanacachi, Dr. O. Buchtien 428 (in 
part). T-Specim.: | of 1. Bolivia, no date, O. Buchtien 
428 (HT: W; IT: US-77388],. 

= Eragrostis pastoensis (Kunth) Trin. 


Eragrostis campestris Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
4,2(1): 72. 1836. T-Protol.: V. subsp. Am. bor., two 
speciemns in hb. Trinius. T-Specim.: | of 2. USA: 
"Jackson v. 4" [North Carolina?, Georgia?, Florida?] 
[ST: LE-TRIN-2316.01]. 2 of 2. USA: Georgia: in 
campis arenosis, Beyrich mis. 1834 [ST: LE-TRIN- 
2316.02], packet Poa nitida (Ell.) Spreng] herb, 91. 

= Eragrostis refracta (Muhl. ex Elliott) Scribn. 


Eragrostis campestris var. refracta Chapm., Fl. South. U.S. 
(ed. 3) 617. 1897. T-Protol.: "Dry soil, middle 
Florida.". 

NOTE: cites Poa refracta Muhl. (nom. nud). 
= Eragrostis refracta (Muhl. ex Elliott) Scribn. 


Eragrostis capillacea Jedwabn., Bot. Arch. 5(3-4): 196. 
1924. ‘T-Protol.: Americae sept. Civit. Unitae, 
Nebraska centr., ad Plumer Ford, Dismal River, 
Thomas Co., Rydberg /832, syntype specimen at B 
presumably destroyed. T-Specim.: | of 1. USA: 
Nebraska: Thomas Co.: on sand hills, near Plumer 
Ford, Dismal River, 23 Aug 1893, P.A. Rydberg 1832 
[LT: NY-380756; ILT: US-207974], LT designated by 
Witherspoon, A Numerical Tax. Stud. Eragrostis 
intermedia Complex, Ph.D. thesis, 252 (1975), but not 
effectively published here because it is an unpublished 
thesis. 

= Eragrostis trichodes (Nutt.) Alph. Wood. 


Eragrostis capillaris (L.) Nees, Fl. Bras. Enum. Pl. 2: 505. 
1829. Basionym: Poa capillaris L. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Fernald, 
M. (1950); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Great Plains 
Flora Association (1986); Hitchcock, A. S. (1951); 


Radford, A. E., H. E. Ahles & C. R. Bell (1968); 
Small, J. K. (1933); Voss, E. G. (1972). 

Aira capillacea Lam.;, Eragrostis capillaris (L.) 
Steud.; Eragrostis pilosa var. capillaris (L.) Kuntze; 
Eragrostis tenuis Steud.; Poa capillaris L.; Poa 
tenuis Elliott. 


SYN:: 


Eragrostis capillaris (L.) Steud., Syn. Pl. Glumac. 1: 273. 
1854. Basionym: Poa capillaris L. 
NOTE: hom. illeg. 
E. NAME: Eragrostis capillaris (L.) Nees. 
= Eragrostis capillaris (L.) Nees. 


Eragrostis capillaris var. frankii (C.A. Mey. ex Steud.) 
Farw., Rep. Michigan Acad. Sci. 17: 182. 1916. 
Basionym: Eragrostis frankiiC.A. Mey. ex Steud. 

= Eragrostis frankiiC.A. Mey. ex Steud. 


Eragrostis capitata (Nutt.) Nash, Man. Fl. N. States 1042. 
1901. Basionym: Poa capitata Nutt. 
= Eragrostis reptans (Michx.) Nees. 


Eragrostis carazensis Pilg., Bot. Jahrb. Syst. 56(Beibl. 123): 
27. 1920. T-Protol.: Peru: Ancachs: Caraz, offene 
Formation bei 2200-2500 m, May 1903, Weberbauer 
2999, T-Specim.: 1 of 1. Peru: Ancash: Craz, alt. 
2200-2500 m, 12 May 1903, 4. Weberbauer 2999 
[HT: B; IT: BAA-1018 (fragm. ex B), US-2891460]. 

= Eragrostis pilgeri Fedde. 


Eragrostis caroliniana (Biehler) Scribn., Mem. Torrey Bot. 
Club 5(4): 49. 1894. Basionym: Poa caroliniana 
Biehler. 

= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis cataclasta Nicora, FI. Il. Entre Rios 2: 189. 1969. 
T-Protol.: Argentina: Entre Rios: Holt, R. Martinez 
Crovetto 8819 [HT: BAB]. 

DST.: Arge, Braz, Para, Urug. 

LVL. ACCPTC.:: I. 

ALSO ACCPTD. BY: Burkart, A. (1969); Cabrera, A. L. 
(1970); Nicora, E. G. (1998); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970). 


Eragrostis caudata E. Fourn., Mexic. Pl. 2: 115. 1886. T- 
Protol.: Mexico, 2 syntypes cited. T-Specim.: 1 of 2. 
Mexico: Consoquitla, Aug 1841, FM. Liebmann 520 
[ST: C, US-207543 (fragm.)]. 2 of 2. Mexico: near 
Matamoras, Sept 1851, J.-L. Berlandier 2345 [ST: MO- 
129490, US-911405 (fragm.), US-77390 (fragm.)]. 

NOTE: hom. illeg. 
E. NAME: Eragrostis caudata Nees ex Steud. 
= Eragrostis palmeri S. Watson. 


Eragrostis chariis (Schult.) Hitche., Lingnan Sci. J. 7: 193. 
1929 [1931]. Basionym: Poa chariis Schult. 
= Eragrostis atrovirens (Desf.) Trin. ex Steud. 


Eragrostis chilensis (Moris) Nees, Nova Acta Acad. Caes. 
Leop.-Carol. German. Nat. Cur. 19: Suppl. 1: 32, 
1841; 164. 1843. Basionym: Poa chilensis Moris. 

= Eragrostis mexicana (Hornem.) Link. 


Eragrostis chiquitaniensis Killeen, Ann. Missouri Bot. 
Gard. 77(1): 153, f. 3. 1990. T-Protol.: Bolivia: Santa 
Cruz: Estancia San Ignacio, 25 km N of San José de 
Chiquitos, Prva. Chiquitos, 17°35'S 60°45'W, 320 m, | 
Feb 1986, 7. Killeen 1728 |HT: ISC; IT: CTES, F, 
LPB, MO, NY, SI, TEX, US]. T-Specim.: | of 1. 
Bolivia: Santa Cruz: Estancia San Ignacio, 25 km N of 


Catalogue of New World Grasses: II. 87 


San José de Chiquitos, Prva. Chiquitos, 17°35'S 
60°45'W, 320 m, 1 Feb 1986, 7. Killeen 1728 [HT: 
ISC; IT: CTES, F, GA, LPB, MO, NY, SI, TEX, US- 
3181404]. 

DST.: Boli, Para. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Killeen, T. J. (1990); Nicora, E. G. 

(1998): Renvoize, S. A. (1998). 


Eragrostis chloromelas Steud., Syn. Pl. Glumac, 1: 271. 
1854. T-Protol.: Pr. b. sp. T-Specim.: | of 1. South 
Africa: Cape Province: Schiloh, Drege [IT: K]. 

NOTE: based on "E. atrovirens sensu Nees, Agrostogr. Cap. 
400, non Desf.". 

NOTE: sometimes regarded as a separate species. 

= Eragrostis curvula (Schrad.) Nees. 


Eragrostis cilianensis (All.) Link ex Vignolo, Malpighia 18: 
386. 1904. Basionym: Poa cilianensis All. 

NOTE: nom. inval., combination not accepted by author; 
See Taxon 35: 697. 1986. Cited by fna 1494 as: "E. 
cilianensis (All.) Mosher" Simon (1983) notes that 
when Vignolo-Lutati made the new combination E. 
cilianensis, mistakingly attributing it to Link, he did 
not not accept the combination, which was thus not 
validly published. The first valid use was by Janchen 
(1907). 

CORRECT NAME: Eragrostis cilianensis (All.) Vignolo ex 
Janch. 

= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis cilianensis (All.) Vignolo, Malpighia 18: 386. 
1904. Basionym: Poa cilianensis All. 
NOTE: nom. inval., combination not accepted by author; 
see Taxon 35: 697 (1986). 
CORRECT NAME: Eragrostis cilianensis (All.) Vignolo ex 
Janch. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis cilianensis (All.) Vignolo ex Janch., Mitt. 
Naturwiss. Vereins Univ. Wien, n.s., 5: 110. 1907. 
Basionym: Poa cilianensis All. 

NOTE: for authorship see: Taxon 35: 697 (1986). 

DST.: Arge, Boli, Braz, Cana, Cari, CoRi, Ecua, ElSa, Guat, 
Hond, Mexi, Nica, Para, Peru, USA, Urug, Vene. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Berendsohn, W. G. & A. E. Araniva 
de Gonzalez (1991); Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Brako, L. & J. L. Zarucchi (1993); 
Burkart, A. (1969); Cabrera, A. L. (1970); Cope, T. 
A. (1992); Davidse, G. (1994); Davidse, G., M. 
Sousa S. & A. O. Chater (editores generales) (1994); 
Edgar, E., H. E. Connor & J. E. Shand (1991); Espejo 
Serna, A., A. R. L6pez-Ferrari, and J. Valdes-Reyna 
(2000); Filgueiras, T. S. (1991); Gleason, H. A. & A. 
Cronquist (1991); Gould, F. W (1979); Gould, F. W. 
& R. Moran (1981); Hickman, J. C. (1993); 
Hitchcock, A. S. (1936); Jorgensen, P. M. & S. Leén- 
Yanez (eds.) (1999); Kerguelen, M. (1993); 
McVaugh, R. (1983); Nicora, E. G., M. E. De Paula, 
A. M. Faggi, M. del C. Mariano, A. M. Miante A., L. 
R. Parodi, C. A. Petetin, F. A. Roig & Z. Rugolo de 
Agrasar (1978); Phillips, S. (1995); Pohl, R. W. 
(1980); Reeder, J. R. & C. G. Reeder (1971); 
Renvoize, S. A. (1998); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1968); 
Rosengurtt, B., B. R. Arrillaga de Maffei & P. 


Izaguirre de Artucio (1970); Shreve, F. & I. L. 
Wiggins (1964); Tovar, O. (1993); Wunderlin, R. P. 
(1998). 

SYN.: Briza eragrostis L.; Briza megastachya (Koeler) 
hort. ex Roem. & Schult.; Briza megastachya 
(Koeler) Steud.; Briza oblonga Moench; Briza 
purpurascens Muhl.; Calotheca purpurascens 
(Muhl.) Spreng.; Eragrostis argentina Jedwabn.; 
Eragrostis cilianensis (All.) Link ex Vignolo; 
Eragrostis cilianensis (All.) Mosher; Eragrostis 
cilianensis (All.) Vignolo; Eragrostis cilianensis 
subsp. megastachya (Koeler) Maire & Weiller; 
Eragrostis eragrostis (L.) Blatt. & McCann; 
Eragrostis eragrostis (L.) MacMill.; Eragrostis 
eragrostis var. megastachya (Koeler) — Farw.; 
Eragrostis major Host; Eragrostis major Host; 
Eragrostis megastachya (Koeler) Link; Eragrostis 
megastachya fo. nana Lorentz & Niederl.; Eragrostis 
megastachya var. cilianensis (All.) Asch. & Graebn.; 
Eragrostis minor var. megastachya (Koeler) Burtt 
Davy; Eragrostis multiflora (Forssk.) Asch.; 
Eragrostis multiflora var. cilianensis (All.) Maire; 
Eragrostis multiflora var. glandulifera Chiov.; 
Eragrostis multiflora var. insularis A. Terracc.; 
Eragrostis multiflora var. subbiloba  Chiov.; 
Eragrostis pappiana var. insularis A. Terrace. ex 
Chiov.; Eragrostis pappii Gand.; Eragrostis 
poaeoides var. megastachya (Koeler) A. Gray; 
Eragrostis virletii E. Fourn.; Eragrostis vulgaris 
subsp. megastachya (Koeler) Douin; Eragrostis 
vulgaris var. megastachya (Koeler) Coss. & Germ.; 
Erosion ciliare Lunell; Megastachya eragrostis (L.) 
P. Beauv. ex Roem. & Schult.; Megastachya obtusa 
(Nutt.) Schult.; Megastachya purpurascens (Muhl.) 
Schult.; Poa cilianensis All.; Poa nuttallii Spreng.; 
Poa obtusa Nutt.; Poa philadelphica W.P.C. Barton. 


Eragrostis cilianensis (All.) Mosher, Univ. Illinois Agric. 


Exp. Sta. Bull. 205: 381. 
cilianensis All. 
NOTE: isonym; See Taxon 35: 697 (1986). 
E. NAME: Eragrostis cilianensis (All.) Vignolo ex Janch. 
CORRECT NAME: Eragrostis cilianensis (All.) Vignolo ex 
Janch. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


1918. Basionym: Poa 


Eragrostis cilianensis subsp. megastachya (Koeler) Maire & 


Weiller, Fl. Afrique N. 2: 175. 1953. Basionym: Poa 
megastachya Koeler. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis ciliaris (L.) R. Br. var. ciliaris. 

DST.: Arge, Beli, Boli, Braz, Cari, Colo, CoRi, Ecua, ElSa, 
FrGu, Guat, Guya, Hond, Mexi, Nica, Pana, Para, 
Peru, Suri, USA, Vene. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Balick, M. J.. M. H. Nee & D. E. 
Atha (2000); Bor, N. L. (1960); Davidse, G. (1994); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Lindeman, J. C. 
& A. L. Stoffers (1963). 

SYN.: Eragrostis ciliaris vat. comta (Link) Schrad. 


Eragrostis ciliaris vat. comta (Link) Schrad., Linnaea 12(4): 
451. 1838. Basionym: Eragrostis comta Link. 
= Eragrostis ciliaris var. ciliaris. 


88 Catalogue of New World Grasses: II. 


Eragrostis ciliaris var. laxa Kuntze, Revis. Gen. Pl. 2: 774. 
1891. T-Specim.: | of 2. Barbados, Kuntze s.n. [ST: 
NY-70975, MO (microfiche ex NY)]. 2 of 2. Virgin 
Islands: St. Thomas Isl., St. Thomas, Kuntze 735 [ST: 
NY-70974]. 

DST.: Cari, Mexi, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Davidse, G. (1994); Davidse, G., M. 
Sousa S. & A. O. Chater (editores generales) (1994); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Hitchcock, A. S. (1936). 


Eragrostis ciliaris var. patens Chapm. ex Beal, Grass. N. 
Amer. 2: 479. 1896. T-Protol.: USA: Georgia, Curtiss 
3493, T-Specim.: | of 1. USA: Georgia: Wayne Co., 
Sep 1880-Oct 1880, 4.H. Curtiss 3493 [IT: NY; US- 
748296, US-821866, US-909340]. 
= Eragrostis amabilis (L.) Wight & Arn. ex Nees. 


Eragrostis cognata Steud., Syn. Pl. Glumac. 1: 273. 1854. T- 
Protol.: Ohio Am. Spt., 1837, 4b. Frank s.n., "P. 
capillaris? hrbr. Frank un. it. 1837; Prioris [E. unionis] 
vix varietas". T-Specim.: | of 1. USA: Ohio, 1837, 
Hrbr. Frank s.n, (HT: P; IT: US-2851265 (fragm.)]. 

= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis collocarpa kK. Schum., Pflanzenw. Ost-Afrikas C: 
114. 1895. T-Specim.: | of 3. Usagara - Usambara, 
Holst 54 [ST]. 2 of 3. Usagara - Usambara, Holst 3228 
{ST]. 3 of 3. Usagara - Usambara, Holst 8923 [ST]. 
= Eragrostis tenuifolia (A. Rich.) Hochst. ex Steud. 


Eragrostis compacta Salzm. ex Steud., Syn. Pl. Glumac. 1: 
275. 1854. T-Protol.: Brazil. in sabulosis maritimus 
subhumidis prope Bahia, P. Salzman, "E. inconstans 
Nees var.". T-Specim.: | of 1. Brazil: Bahia, P. 
Salzmann s.n. [HT: P; IT: K, MO, US-911749, US- 
911748]. 

= Eragrostis secundiflora J. Presl. 


Eragrostis condensata (J. Presl) Steud., Syn. Pl. Glumac. 1: 
278. 1854. Basionym: Megastachya condensata J, 
Presl. 

DST.: Ecua. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Jorgensen, P. M. & C. Ulloa U. 
(1994); Jorgensen, P. M. & S. Leén-Yanez (eds.) 
(1999); Valencia, R., N. Pitman, S. Leén-Yanez & P. 
M. Jorgensen (2000). 

Brizopyrum condensatum (J. Presl) E. Fourn.; 
Distichlis condensata (J. Presl) Hemsl.; Eragrostis 
densissima Hack.; Eragrostis tenax (Kunth) Steud.; 
Megastachya condensata J. Presl; Megastachya tenax 
(Kunth) Roem. & Schult.; Poa condensata (J. Presl) 
Kunth; Poa tenax Kunth. 


SYN.: 


Eragrostis conferta (Elliott) Trin., Mem. Acad. Imp. Sci. St.- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 409. 1830. 
Basionym: Poa conferta Elliott. 

= Eragrostis japonica (Thunb.) Trin. 


Eragrostis contracta Pilg., Bot. Jahrb. Syst. 37: 376. 1906. T- 
Specim.: | of 3. Peru: Depart. Ancachs, in provincia 
Cajatambo, ad Chiquian, in formatione aperta, plantas 
herbaceas (maxime Gramineas) intermixtis fruticibus 
gerente, 3000-3300 m, Apr 1903,4. Weberbauer 2841 
[ST: B, BAA-1020 (fragm. ex B)]. 2 of 3. Peru: prope 
Hacienda Cajabamba inter Samanco et Caraz, in 


formatione aperta. 3700 m, A. Weberbauer 3035 (ST: 
US-2767408 (fragm.)]. 3 of 3. Peru: prope Ocros, in 
formatione aperta, 3200-3400 m,A. Weberbauer 2663 
[ST: US-1061508 (fragm.)]. 

= Eragrostis pastoensis (Kunth) Trin. 


Eragrostis contrerasii R.W. Pohl, lowa State J. Res. 51(3): 
323, f. 1. 1977. T-Protol.: Guatemala: Peten: in 
clearing, bordering Rio Usumacinta, ca. 9 km W of 
Lacondon, 8 Mar 1962, E. Contreras 3484 [HT: F; IT: 
LL, TEX]. T-Specim.: | of 1. Guatemala, 8 March 
1962, Contreras 3484 |HT: F; IT: US-2486077, US- 
2454156, F-05351 1]. 

DST.: Beli, Guat, Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Balick, M. J., M. H. Nee & D. E. 
Atha (2000); Davidse, G. (1994); Davidse, G., M. 
Sousa S. & A. O. Chater (editores generales) (1994). 

SYN.: Neeragrostis contrerasii (R.W. Pohl) P.M. Peterson. 


Eragrostis contristata Meyen, Reise Erde 1: 484. 1834. 
NOTE: nom. nud. 
CORRECT NAME: Eragrostis contristata Nees & Meyen. 
= Eragrostis lurida J. Presl. 


Eragrostis cordobensis Jedwabn., Bot. Arch. 5(3-4): 208. 
1924. T-Specim.: 1 of 3. Argentina: Cordoba: prope 
urbem, 1881, Galander 54b (ST: US-2767407]. 2 of 3. 
Argentina: Salta: Rio Juramento, Lorentz & 
Hieronymus 356 [ST]. 3 of 3. Argentina: Santa Fé: 
Canada de Gomez, Galander 25 [ST]. 

= Eragrostis mexicana (Hornem.) Link. 


Eragrostis cubensis Hitchc., Contr. U.S. Natl. Herb. 12(6): 
243. 1909. T-Protol.: Cuba: Isle of Pines, Curtiss 420 
[HT: US-522037]. T-Specim.: | of 1. Cuba: Pinar del 
Rio: near Nueva Gerona, Isla de Pinos, 20 Mar 1904, 
A.H. Curtiss 420 (HT: US-522037; IT: MO-2067266, 
NY-70976]. 

DST.: Cari. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Catasus 
Guerra, L. (1997); Hitchcock, A. S. (1909); 
Hitchcock, A. S. (1936). 


Eragrostis cumingii Steud., Syn. Pl. Glumac. 1: 266. 1854. 
T-Specim.: | of 2. Philippines: Luzon: Laguna, 1841, 
Cuming 672 [ST: L, LE]. 2 of 2. Philippines: Luzon, 
Cuming 1104 (ST: L]. 

DST.: USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Anonymous (1986); Bor, N. L. 
(1960); Koch, S. D. (1978); Lazarides, M. (1961); 
Wunderlin, R. P. (1998). 

SYN.: Eragrostis bleeseri Pilg.; Eragrostis distans Hack.; 
Eragrostis simplex Scribn. 


Eragrostis curtipedicellata Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 97. 1862. T-Protol.: USA: Northern 
Texas, June. T-Specim.: | of |. USA: northern Texas, 
Buckley s.n. [LT: PH], LT designated by Hitchcock, 
Man. Grass. U.S. 849 (1935), but without citing a 
specific sheet in a specific herbarium. 

DST.: Mexi, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. L6pez-Ferrari, and J. Valdes- 


Catalogue of New World Grasses: II. 89 


Reyna (2000); Great Plains Flora Association (1986); 
Hitchcock, A. S. (1951). 

SYN.: Eragrostis brevipedicellata A. Gray; Eragrostis 
viscosa Scribn. 


Eragrostis curvula (Schrad.) Nees, Fl. Afr. Austral. Ill. 397. 
1841. Basionym: Poa curvula Schrad. 
DST.: Arge, Boli, Braz, Chil, Colo, CoRi, USA, Urug, 
Vene. 

LVL. ACCPTC.: 1; cultivated, introduced. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Boechat, S. C. 
& H. M. Longhi-Wagner (2000); Bor, N. L. (1960); 
Brako, L. & J. L. Zarucchi (1993); Burkart, A. 
(1969); Cabrera, A. L. (1970); Correll, D. S. & M. C. 
Johnston (1970); Cronquist, A., A. H. Holmgren, N. 
H. Holmgren & J. L. Reveal (1977); Darke, R. 
(1999); Davidse, G. (1994); Davidse, G., M. Sousa S. 
& A. O. Chater (editores generales) (1994); Edgar, 
E., H. E. Connor & J. E. Shand (1991); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Filgueiras, T. S. (1991); Gibbs Russell, G. E, W. G. 
Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987); Gleason, H. A. & A. 
Cronquist (1991); Great Plains Flora Association 
(1986); Hitchcock, A. S. (1951); Kerguelen, M. 
(1993); Marticorena, C. & M. Quezada (1985); 
Molina R., A. (1975); Munz, P. A. & D. D. Keck 
(1959); Munz, P. A. (1974); Phillips, S. (1995); Pohl, 
R. W. (1980); Radford, A. E., H. E. Ahles & C. R. 
Bell (1968); Renvoize, S. A. (1988); Renvoize, S. A. 
(1998); Rosengurtt, B., B. R. Arrillaga de Maffei & 
P. Izaguirre de Artucio (1970); Smith, L. B., D. C. 
Wasshausen & R. M. Klein (1981); Wunderlin, R. P. 
(1998). 

Eragrostis chloromelas Steud.; Eragrostis curvula 
var. atrata (Schweinf.) Cufod.; Eragrostis robusta 
Stent; Eragrostis thunbergiana var.atrata Schweinf., 
Poa curvula Schrad. 


SYN.: 


Eragrostis curvula vat. atrata (Schweinf.) Cufod., Bull. Jard. 
Bot. Belg. 38: 1251. 1968. Basionym: Eragrostis 
thunbergiana var. atrata Schweinf. 

= Eragrostis curvula (Schrad.) Nees. 


Eragrostis curvula (Schrad.) Nees var. curvula. 
DST.: Braz. 
LVL. ACCPTC.: 1; introduced. 
ALSO ACCPTD. BY: Hickman, J. C. (1993); Smith, L. B., 
D.C. Wasshausen & R. M. Klein (1981). 


Eragrostis curvula var. valida Stapf, Fl. Cap. 7: 600. 1900. 
T-Protol.: Near Durban, Plant, 57. T-Specim.: | of 1. 
Africa do Sul, Natal, Berea., Wood 5940 [T]. 

DST.: Braz. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Smith, L. B., D. C. Wasshausen & R. 
M. Klein (1981). 


Eragrostis cuspidata (Schult.) Link, Hort. Berol. 2: 293. 
1833. Basionym: Megastachya cuspidata Schult. 
= Eragrostis rufescens var. rufescens. 


Eragrostis cylindriflora Hochst., Flora 38: 324. 1855. T- 
Protol.; Abyssinia, Semien [1852], Schimper in Herb. 
Buchinger 772 {HT: STR], in pl. abyss. anno 1852 a 
Schimpero lectis. 

DST.: USA. 


LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Gibbs Russell, G. E, W. G. Welman, 
E. Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987); Phillips, S. (1995). 

Eragrostis annulata Chiov.; Eragrostis cylindriflora 
var. gymnorhachis Schweinf.; Eragrostis horizontalis 
Peter; Eragrostis multipilosa Hochst. ex Mattei. 


SYN.: 


Eragrostis cylindriflora var. gymnorhachis Schweint., Bull. 
Herb. Boissier 2(App. 2): 40. 1894. T-Specim.: | of 6. 
Aethiopian colonia Eritrea, Schweinfurth 1151 [ST]. 2 
of 6. Aethiopian, Colonia Eritrea, Schweinfurth 1102 
[ST]. 3 of 6. Aethiopien, Colonial — Eritrea, 
Schweinfurth 1289 [ST]. 4 of 6. Aethiopien, Colonia 
Eritrea, Schweinfurth 1489 [ST]. 5 of 6. Aethiopien, 
Colonia Eritrea, Schweinfurth 1623 [ST]. 6 of 6. 
Aethiopien, Colonia Eritrea, Schweinfurth 9] [ST]. 

= Eragrostis cylindriflora Hochst. 


Eragrostis cyperoides (Thunb.) P. Beauv., Ess. Agrostogr. 
71, 162, 174. 1812. Basionym: Poa cyperoides Thunb. 
= Cladoraphis cyperoides (Thunb.) S.M. Phillips. 


Eragrostis damiensiana (Bonnet) Thell., Repert. Spec. Nov. 
Regni Veg. 24: 323. 1928. Basionym: Eragrostis 
pilosa var. damiensiana Bonnet. 

= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis damiensiana var. condensata (Hack.) Thell., 
Repert. Spec. Nov. Regni Veg. 24: 328. 1928. 
Basionym: Eragrostis pilosa var. condensata Hack. 

= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis damiensiana var. laxior Thell., Repert. Spec. Nov. 
Regni Veg. 24: 327. 1928. 
NOTE: cites Poa (Eragrostis) purshii. var. Fisch. Mey. Lall. 
Accd. to Chase & Niles, Index Grass Sp. 
= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis decipiens (R. Br.) Steud., Syn. Pl. Glumac. 1: 279. 
1854. Basionym: Poa decipiens R. Br. 
= Leptochloa decipiens (R. Br.) Stapf ex Maiden. 


Eragrostis delicatula Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
2(1): 73. 1836. T-Protol.: Brazil: "V. subsp. Bras. in 
cultis prope Rio Jan.". T-Specim.: | of 1. Brazil: In 
cultis prope Rio de Janeiro, May-June 1823,L. Riedel 
[HT: LE-TRIN-2330.01; IT; LE, LE, US-2891464 
(fragm.)]. 

= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis densissima Hack., Oesterr, Bot. Z. 52: 304. 1902. 
T-Protol.: Ecuador: in regione temperata et subandina 
prope Tumbaco etc., Sodiro s.n. T-Specim.: | of 1. 
Ecuador: in reg. temperata et Subandina Prope 
Tumbaco, 1886, S.J.Sodiro [LT: US-2891465], LT 
designated by Hitchcock, Contr. U.S. Natl. Herb. 
24(8): 340, no herbarium indicated. 
= Eragrostis condensata (J. Presl) Steud. 


Eragrostis deserticola Phil., Fl. Atacam. 55. 1860. T-Protol.: 
Chile: circa Hueso parado 25° 26' lat m. et 1200 p. s. 
m. inveni. T-Specim.: | of 2. Chile: circa Hueso 
Parado, 25°26'S, 1200 m [HT: SGO-63452; IT: BAA 
(fragm.)]. 2 of 2. Chile, R.A. Philippi 658 or 608 [IT: 
US-556537 (fragm.)]. 
= Eragrostis peruviana (Jacq.) Trin. 


90 Catalogue of New World Grasses: II. 


Eragrostis diffusa Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 97. 1862. T-Protol.: USA: Northern 
Texas, Buckley s.n. T-Specim.: | of 1. USA: Texas, 
S.B. Buckley [LT: PH; IT: US-91621 (fragm.)], LT 
designated by Hitchcock, Man. Grass. U.S. 849 
(1935), but without citing a specific sheet in a specific 
herbarium. 

= Eragrostis pectinacea var. pectinacea. 


Eragrostis diffusa var. arida (Hitche.) Beetle, Phytologia 
37(4): 317-407. 1977. Basionym: Eragrostis arida 
Hitche. 

= Eragrostis pectinacea var. miserrima (E. Fourn.) 

Reeder. 


Eragrostis diffusa Buckley var. diffusa. 
= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis diplachnoides Steud., Syn. Pl. Glumac. 1: 268. 
1854. T-Specim.: | of 1. Sudan, Kotschy 346 [HT: ?; 
IT: K, L], L cites coll. as B.P.G. Hochreutiner. 
= Eragrostis japonica (Thunb.) Trin. 


Eragrostis distans Hack., Philippine Bur. Govt. Lab. Bull. 
35: 81. 1906. T-Protol.: Kias, Benguet, Luzon, Jun 
1904, Elmer 6608. 

= Eragrostis cumingii Steud. 


Eragrostis diversiflora Vasey, Contr. U.S. Natl. Herb. 1(8): 
285. 1893. T-Protol.: Mexico: Manzanillo, 2-18 Mar 
1891, E. Palmer 1335. T-Specim.: | of 1. Mexico: 
Colima: Manzanillo, 2-18 Mar 1891, £. Palmer 1335 
{HT: US-1749577; IT: 821924, US-910000]. 
= Eragrostis prolifera (Sw.) Steud. 


Eragrostis domingensis (Pers.) Steud., Syn. Pl. Glumac. 1: 
278. 1854. Basionym: Poa domingensis Pers. 
= Eragrostis prolifera (Sw.) Steud. 


Eragrostis echinochloidea Stapf, Fl. Cap. 7: 627. 1900. T- 
Protol.: Kalahari Region: Orange Free State, between 
Kimberly anf Bloemfontein, Buchanan 284. 

DST.: USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Gibbs Russell, G. E, W. G. Welman, 
E. Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987). 


Eragrostis ekmanii Hitche., Man. Grasses W. Ind. 43, f. 22. 
1936. T-Protol.: West Indies: Cuba: Los Indios, Isla de 
Pinos: collected in sandy pinelands: 8 Nov 1920.,E.L. 
Ekman 12143 (HT: US-1387320]. T-Specim.: 1 of 1. 
Cuba: Isla de la Juventud, Isla de Pinos, Los Indios, 8 
Nov 1920, E. L. Ekman 12/43 [HT: US-1387320; IT: 
NY-70973 [7 Nov]]. 

= Eragrostis elongata (Willd.) J. Jacq. 


Eragrostis elatior Stapf, Fl. Cap. 7: 617. 1900. T-Specim.: 1 
of 3. Caput bonae spei, 1839, Zeyher s.n. [IST: LE]. 2 
of 3. Coast Region: Cape Div.: Cape Flats, near Doorn 
Hoogte, 1839, Zeyher s.n. [ST]. 3 of 3. Coast Region: 
Paarl Div.: by the Berg River near Paarl, Drége s.n. 
[ST]. 

= Eragrostis bahiensis Schrad. ex Schult. 


Eragrostis elatior Hack., Repert. Spec. Nov. Regni Veg. 7: 
374. 1909. T-Protol.: Paraguay: Sierra de Amambay: 
ad margines silvarum, Apr 1908, Rojas (Hassler 
10380). T-Specim.: 1 of |. Paraguay, ad margines 


silvarum: Sierra de Amambay, Apr 1908, 7. Rojas 
10380 [IT: BAA, BAF, P, US-911888, US-77387 
(fragm.), US-77392 (fragm.)]. 

NOTE: hom. illeg. 

E. NAME: Eragrostis elatior Stapf. 

CORRECT NAME: Eragrostis hackelii Hassl. 

= Eragrostis hackelii Hass. 


Eragrostis elegans Nees, Fl. Bras. Enum. Pl. 2: 510, 608. 
1829. 
REPLACED NAME: Poa brasiliensis Spreng. 
BLOCKING NAME: Eragrostis brasiliensis (Raddi) Nees. 
= Eragrostis japonica (Thunb.) Trin. 


Eragrostis elegans var. laxiflora Arechav., Anales Mus. Nac. 
Montevideo 1: 444. 1896. ‘T-Protol.: Uruguay: 
Western border of Uruguay: in front of Salto. 

= Eragrostis japonica (Thunb.) Trin. 


Eragrostis elegantula Nees, Anal. Bot. Ind. 4: 39. 1851. T- 
Protol.: India, Wight 1781. 
= Eragrostis gangetica (Roxb.) Steud. 


Eragrostis elegantula (Kunth) Nees ex Steud., Syn. PI. 
Glumac. 1: 266. 1854. Basionym: Poa elegantula 
Kunth. 

NOTE: hom. illeg. 

E. NAME: Eragrostis elegantula Nees. 

CORRECT NAME: Eragrostis chariis (Schult.) Hitche. 

= Eragrostis atrovirens (Desf.) Trin. ex Steud. 


Eragrostis elliottii S. Watson, Proc. Amer. Acad. Arts 25: 
140. 1890. 

REPLACED NAME: Poa nitida Elliott. 

BLOCKING NAME: Eragrostis nitida Link. 

DST.: Beli, Cari, Hond, Mexi, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Anonymous 
(1986); Balick, M. J.. M. H. Nee & D. E. Atha 
(2000); Beetle, A. A. (1977); Breedlove, D. E. 
(1986); Catasus Guerra, L. (1997); Correll, D. S. & 
M. C. Johnston (1970); Cowan, C. P. (1983); 
Davidse, G. (1994); Davidse, G., M. Sousa S. & A. 
O. Chater (editores generales) (1994); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000): 
Godfrey, R. K. & J. W. Wooten (1979); Hitchcock, 
A. S. (1909); Hitchcock, A. S. (1936); Hitchcock, A. 
S. (1951); Koch, S. D. (1978); Liogier, H. & L. 
Martorell (1982); Long, R. W. & O. Lakela (1971); 
McClure, F. A. (1955); Molina R., A. (1975); 
Radford, A. E., H. E. Ahles & C. R. Bell (1968); 
Sousa S., M. & E. F. Cabrera C. (1983); Wunderlin, 
R. P. (1998). 

Eragrostis acuta Hitche.; Eragrostis macrapoda 
Pilg.; Eragrostis nitida (Elliott) Chapm.; Poa nitida 
Elliott. 


SYN.: 


Eragrostis elongata (Willd.) J. Jacq., Ecl. Gram. Rar. 3. 
1813. Basionym: Poa elongata Willd. 

DST.: Chil, CoRi, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Anonymous (1986); Brako, L. & J. 
L. Zarucchi (1993); Davidse, G. (1994); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994): Koch, S. D. (1978); Marticorena, C. & M. 
Quezada (1985); Skottsberg, C. (1922); Wunderlin, 
R. P. (1998). 


Catalogue of New World Grasses: II. 9] 


SYN.: Eragrostis ekmanii Hitche.; Eragrostis stenoclada J. 
Presl; Poa elongata Willd. 


Eragrostis eragrostis (L.) P. Beauv., Ess. Agrostogr. 71, 174, 
t. 14, f. 11. 1812. Basionym: Poa eragrostis L. 
NOTE: nom. inval., tautonym. 
CORRECT NAME: Eragrostis poaeoides P. Beauv. ex 
Roem. & Schult. 
= Eragrostis minor Host. 


Eragrostis eragrostis (L.) H. Karst., Deut. Fl. 389. 1880. 
Basionym: Poa eragrostis L. 
NOTE: nom. inval., tautonym. 
E. NAME: Eragrostis eragrostis (L.) P. Beauv. 
= Eragrostis minor Host. 


Eragrostis eragrostis (L.) MacMill., Metasp. Minnesota 
Valley 75. 1892. Basionym: Briza eragrostis L. 
NOTE: nom. inval., tautonym. 
E. NAME: Eragrostis eragrostis (L.) P. Beauv. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis eragrostis (L.) Blatt. & McCann, J. Bombay Nat. 
Hist. Soc. 33: 492. 1929. Basionym: Briza eragrostis 
L. 
NOTE: nom. inval., tautonym. 
E. NAME: Eragrostis poaeoides P. Beauv. ex Roem. & 
Schult. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis eragrostis var. megastachya (Koeler) Farw., Rep. 
Michigan Acad. Sci. 17: 182. 1916. Basionym: Poa 
megastachya Koeler. 

NOTE: nom. inval., because the species is invalid. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis eragrostis var. microstachya (Coss. & Germ.) 
Farw., Amer. Midl. Naturalist 10: 306. 1927. 
Basionym: Eragrostis vulgaris var. microstachya 
Coss. & Germ. 

NOTE: nom. inval., because the species is invalid. 
= Eragrostis minor Host. 


Eragrostis erosa Scribn. ex Beal, Grass. N. Amer. 2: 483. 
1896. T-Protol.: Mexico: Chihuahua, Pringle 415, T- 
Specim.: | of 1. Mexico: Chihuahua: Santa Eulalia 
Mountains, Oct 1885, C.G. Pringle 415 [HT: MSC; 
IT: MO-3728015, US-821925, US-1749578]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Hitchcock, A. S. (1951). 


Eragrostis erythrogona Nees ex Steud., Syn. Pl. Glumac. 1: 
273. 1854. T-Protol.; USA: Missouri: St. Louis, 
Drummond s.n. T-Specim.: | of 1. USA: Missouri: St. 
Louis, 7. Drummond [IT: LE-TRIN-2338.01, US- 
909490 (fragm.)], herb. Nees. 

= Eragrostis frankiiC.A. Mey. ex Steud. 


Eragrostis eucampta Pilg., Bot. Jahrb, Syst. 70: 346, f. 1939. 
T-Protol.: Brazil: S40 Paulo "E. solida Nees", Sellow 
s.n. (HT: B-W-5533]. T-Specim.: 1 of 2. Brazil: Sao 
Paulo, F. Sellow s.n. [ST]. 2 of 2. : Sellow s.n. [ST: B- 
W-5533]. 
= Eragrostis solida Nees. 


Eragrostis excelsa Griseb., Cat. Pl. Cub. 227. 1866. T- 
Protol.: Cuba: Cuba occ., C. Wright 3425. T-Specim.: 
| of 1. Cuba, 1860-1864,C. Wright 3425 [HT: GOET; 
IT: BM, MO-2067269, NY, US-911718 (fragm.), US- 
911719 (fragm.), US-2767393 (fragm.)]. 

DST.: Beli, Cari, Mexi. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Balick, M. J.. M. H. Nee & D. E. 
Atha (2000); Catasus Guerra, L. (1997); Davidse, G. 
(1994); Davidse, G.. M. Sousa S. & A, O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J.  Valdes-Reyna (2000); 
Hitchcock, A. S. (1909); Hitchcock, A. S. (1936): 
Sousa S., M. & E. F. Cabrera C. (1983). 

SYN.: Eragrostis glumacea E. Fourn. 


Eragrostis expansa Link, Hort. Berol. 1: 190. 1827. T- 
Protol.: Habitat in Montevideo, Se//ow s.n. T-Specim.: 
1 of 1. Uruguay: Montevideo, F. Se/low [IT: US- 
2850751 (fragm.)]. 
= Eragrostis bahiensis Schrad. ex Schult. 


Eragrostis filiformis Link, Hort. Berol. 1: 191. 1827. T- 
Protol.: Hab. in America boreali. T-Specim.: | of 1. 
America Boreali, Ex Herb. Elliot {IT: US-91380 
(fragm.)]. 

= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis firma Trin. Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
4,2(1): 74. 1836. T-Protol.: V. subsp. Brasilia. T- 
Specim.: 1 of 1. St. Paulo Brasil., inter plantas a [HT: 
LE-TRIN-2342.01; IT: US-2891470 (fragm.)]. 

= Eragrostis bahiensis Schrad. ex Schult. 


Eragrostis flaccida Lindm., Kongl. Svenska Vetenskapsakad. 
Handl. 34(6): 17, t. 9A. 1900. T-Specim.: | of 3. 
Brazil: Rio Grande do Sul: Porto Alegre, in regione 
camporum, in dumetis paulum umbrosis, solo duro 
graminoso, 7 Nov 1899, Exp. I. Regnell, A 613 |ILT: 
LIL-19370, P, U, US-703148], LT designated by ?. 2 
of 3. Paraguay, Colonia Risso, in ripa calcarea 
graminosa fluvii Paraguay, loco aprico, 20 Oct 1893, 
Exp. I. Regnell., A 2301 [ST: S, IST: U]. 3 of 3. 
Paraguay: Asuncidén, dans les lieux sablonneux 
humides, mens Febr. florens et fructifera, Balansa 241 
[ST: S]. 

= Eragrostis lugens Nees. 


Eragrostis flavescens J. Presl, Relig. Haenk. 1(4-5): 278. 
1830. T-Protol.: Hab. in Mexico, Haenke s.n. T- 
Specim.: | of 1. Mexico, 7}Haenke [IT: LE-TRIN- 
2344.01, US-2891471 (fragm.)]. 

= Eragrostis acutiflora (Kunth) Nees. 


Eragrostis floribunda Schrad. ex Schult., Mant. 2: 319. 1824. 
T-Protol.: Brazil: In Brasilia Princeps, Sereniss, 
Maximil, Neowidense. 

NOTE: Poa floribunda Schrad. M. S., cited, Accd. to Chase 
& Niles, Index Grass Sp. 
= Eragrostis rufescens Schrad. ex Schult. 


Eragrostis floridana Hitche., Amer. J. Bot. 2: 308. 1915. T- 
Protol.: USA: Florida: in dry pine woods near Tampa: 
March, 4.H. Curtiss 3494* [HT: US-726520]. T- 
Specim.: | of 1. USA: Florida: Hillsborough Co.: near 
Tampa, Mar, Curtiss 3494 [HT: US-726520; IT: BR, 


92 Catalogue of New World Grasses: II. 


F, ISC, LE, M, MO, NY, PH, TAES, TENN, US- 
748356, US-823092]. 
= Eragrostis polytricha Nees. 


Eragrostis fragilis Swallen, Fieldiana, Bot. 28(1): 18. 1951. 
T-Protol.: Venezuela: Bolivar: Gran Sabana, between 
Kun and Uaduara-paru, in valley of Rio Kukenan, S of 
Mount Roraima, 1065-1220 m, | Oct 1944, JA. 
Steyermark 59062 [HT: US-1911657; IT: F, VEN]. T- 
Specim.: | of 1. Venezuela: Bolivar, | Oct 1944,/.4. 
Steyermark 59062 [HT: US-1911657; IT: F, VEN]. 

= Eragrostis polytricha Nees. 


Eragrostis frankii C.A. Mey. ex Steud., Syn. Pl. Glumac. 1: 
273. 1854. T-Protol..: USA: Ohio, Am. septr., Hrbr. 
un. it. 1835 [HT: LE], distributed as Poa tenella. T- 
Specim.: | of |. USA: Ohio, Frank, in C.A. Meyer (Ind. 
sem. H. Petrop. sub nr. 2454) (IT: BM; US-2891472 
(tragm.)]. 

DST.: Cana, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Anonymous (1986); Fernald, M. 
(1950); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Godfrey, R. 
K. & J. W. Wooten (1979); Great Plains Flora 
Association (1986); Hitchcock, A. S. (1951); 
Kerguelen, M. (1993); Radford, A. E., H. E. Ahles & 
C. R. Bell (1968); Schwegman, John E. (1991); 
Scoggan, H. J. (1978 [1979]); Small, J. K. (1933); 
Voss, E. G. (1972). 

Eragrostis capillaris var. frankii (C.A. Mey. ex 
Steud.) Farw.; Eragrostis erythrogona Nees ex 
Steud.; Eragrostis frankii var. brevipes Fassett; 
Eragrostis frankii var. frankii, Poa micrantha 
Schult.; Poa parviflora Nutt. 


SYN.: 


Eragrostis frankii var. brevipes Fassett, Rhodora 34(401): 95. 
1932. T-Protol.: USA: Wisconsin: mud-flats, 
Mississippi River bottoms, Glenhaven, 9 Sep 1930, 
Fassett 12899 |HT: WIS]. T-Specim.: 1 of 1. USA: 
Wisconsin: Grant co., 9 Sep 1930,N.C. Fassett 12899 
{IT: MO, US-1535776]. 

= Eragrostis frankiiC.A. Mey. ex Steud. 


Eragrostis frankii C.A. Mey. ex Steud. var. frankii. 
= Eragrostis frankiiC.A. Mey. ex Steud. 


Eragrostis gangetica (Roxb.) Steud., Syn. Pl. Glumac. 1: 
266. 1854. Basionym: Poa gangetica Roxb. 

DST.: Beli, USA, Vene. 

ALSO ACCPTD. BY: Balick, M. J., M. H. Nee & D. E. 
Atha (2000); Boechat, S.C. & P. M. Peterson (1995); 
Bor, N. L. (1960); Davidse, G. (1994); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994); Gibbs Russell, G. E, W. G. Welman, E. 
Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987); Koch, S. D. (1978); ORSTOM (1988); 
Phillips, S. (1995); Wunderlin, R. P. (1998). 

Eragrostis elegantula (Kunth) Nees ex Steud.; 
Eragrostis elegantula Nees; Eragrostis luzoniensis 
Steud.; Eragrostis ovina Hochst. ex A. Rich.; 
Eragrostis stenoclada J. Presl; Eragrostis stenophylla 
Hochst. ex Miq.; Poa gangetica Roxb.; Poa ovina A. 


SYN:: 


Rich. 
Eragrostis geyeri Steud., Syn. Pl. Glumac. 1: 272. 1854. T- 
Protol.: USA: Illinois, Geyer hrbr. as "Poa 


pectinacea". T-Specim.: | of 1. USA: Illinois [IT: US- 
2891474 (fragm.)]. 
= Eragrostis spectabilis (Pursh) Steud. 


Eragrostis gigantea Trin., Mem. Acad. Imp. Sci. St.- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 403. 1830. T- 
Protol.: V. sp. Domingo. T-Specim.: | of 2. St. 
Domingo [HT ?: LE-TRIN-2346.01], Rudolphi herb.: 
Poitcan Paris 3026: Poa giganthea mihi. 2 of 2. 
Dominican Republic, 1802 [IT: US-2891475 (fragm.)]. 

= Eragrostis prolifera (Sw.) Steud. 


Eragrostis glandulosa L.H. Harv., Bull. Torrey Bot. Club 
81(5): 406. 1954. T-Protol.: Mexico: Guerrero: 
Temisco, ¥. Mexia 8774 [HT: US-1724746]. T- 
Specim.: | of 1. Mexico: Guerrero: Sierra Madre del 
Sur, north of Rio Balsas, Temisco, Barranca de la 
Suriana, streambank, alt. 312 m, 9 Nov 1937, ¥. Mexia 
8774 [HT: US-1724746, IT: MO-1269264, US- 
1834505]. 

DST.: Mexi. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Breedlove, D. E. (1986); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); McClure, F. A. (1955). 


Eragrostis glareosa Trin., Mem. Acad. Imp. Sci. St- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 406. 1830. T- 
Protol.: V. subsp. Brasilia., in glareosis, Barra de 
Jiquitiba. T-Specim.: | of |. Brazil: in glareosis, Barra 
do Jiquitiba, Nov 24, 847 [HT: LE-TRIN-2349.01; IT: 
BAA-1038 (fragm.), K, LE, US-2891476 (fragm.)]. 

= Eragrostis articulata (Schrank) Nees. 


Eragrostis gloeodes Ekman, Ark. Bot. 10(17): 38, t. 4, 6. 

1911. T-Protol.: Brazil: Mato Grosso: Porto Murtinho, 
2 Jan 1903 ("2.1.03"),G.O. Malme 2784 Exp. IT Regn. 
T-Specim.: | of 1. Brazil: Matto Grosso: Porto 
Murtinho, O.Malme 2784 (IT: US-3274383 (fragm.)]. 

DST.: Braz. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 

Wagner (2000). 


Eragrostis glomerata (Walter) L.H. Dewey, Contr. U.S. Natl. 
Herb. 2(3): 543. 1894. Basionym: Poa glomerata 
Walter. 

= Eragrostis japonica (Thunb.) Trin. 


Eragrostis glutinosa (Sw.) Trin., Mem. Acad. Imp. Sci. St.- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 397. 1830. 
Basionym: Poa glutinosa Sw. 

DST.: Cari, Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Beetle, A. A. 
(1977); Catasus Guerra, L. (1997); Hitchcock, A. S. 
(1909); Hitchcock, A. S. (1936); Liogier, H. & L. 
Martorell (1982). 

SYN.: Eragrostis sudans Griseb.; Poa glutinosa Sw. 


Eragrostis gracilis Schrad., Linnaea 12(4): 451. 1838. T- 
Protol.: Chile. T-Specim.: 1 of 1. 1835, Herb. 
Schrader [HT: LE], P. chilensis Hort. Paris. 

NOTE: cites Poa chilensis Hort. Par. (nom. inval., as syn. of 
E. gracilis Schrad., in Steudel Nom. Bot. 2 2:359, 
1841) non Poa chilensis Moris. 

= Eragrostis pilosa (L.) P. Beauv. 


Catalogue of New World Grasses: II. 93 


Eragrostis grandiflora J.G. Sm. & Bush, Annual Rep. 
Missouri Bot. Gard. 6: 117, t. 55. 1895. T-Protol.: 
USA: Oklahoma: Creek Co.: Sapulpa, Indian 
Territory: | Oct 1894, BF. Bush 808. T-Specim.: 1 of 
1. USA: Sapulpa, Indian Territory, 1 Oct 1894, 
BF. Bush 808 [IT: US-91622 (fragm.)]. 

= Eragrostis trichodes (Nutt.) Alph. Wood. 


Eragrostis guatemalensis Witherspoon, Ann. Missouri Bot. 
Gard. 64: 324. 1977. T-Protol.: Guatemala, Johnston 
930 (HT: F]. 
DST.: Braz, FrGu, Guat, Guya, Mexi, Vene. 
LVL. ACCPTC:: 1. 
ALSO ACCPTD. BY: Davidse, G. (1994); Davidse, G., M. 
Sousa S. & A. O. Chater (editores generales) (1994); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Witherspoon, J. T. (1978). 


Eragrostis guianensis Hitche., Contr. U.S. Natl. Herb. 22(6): 
454, f. 78. 1922. T-Protol.: British Guiana: collected at 
Kwaimatta, Rupununi River, southern British Guiana, 
G.S. Jenman 5970 (HT: US-1039337]. T-Specim.: | of 
1. Guyana: Rupununi River, Kwaimatta, Oct. 1888, 
G.S. Jenman 5970 [HT: US-1039337; IT: K, W, US- 
1126593 (fragm.)]. 

DST.: Braz, FrGu, Guya, Vene. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Judziewicz, E. J. (1990). 


Eragrostis hackelii Hassl., Repert. Spec. Nov. Regni Veg. 8: 
47. 1910. 
REPLACED NAME: Eragrostis elatior Hack. 
BLOCKING NAME: Eragrostis elatior Stapf. 
DST.: Para. 
LVL. ACCPTC.: I. 
SYN.: Eragrostis elatior Hack. 


Eragrostis haenkei J. Presl, Relig. Haenk. 1(4-5): 279. 1832. 
T-Protol.: Mexico: Hab. in Mexico, Haenke s.n. T- 
Specim.: | of 1. Mexico, 7:Haenke [IT: US-2942412 
(fragm.)]. 

= Eragrostis prolifera (Sw.) Steud. 


Eragrostis hapalantha Trin., Mem. Acad. Imp. Sci. St.- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 409. 1830. T- 
Protol.: V. subsp. Ind. or. Manil. Bras. (ad flum. 
Parana in arenosis), Riedel s.n. T-Specim.: | of 5. 
India orientali [ST: LE; NY], a. = Poa nutans Koenig 
Ex India orientali m.; b. = Poa interrupta ex India 
orientali. 2 of 5. Ex India orientali, 1820, Hornemann 
[ST: LE], a: P. diandra R.; b. = Poa interrupta Lam. & 
Roth. non Br. Trinius scripsit ; c. = Poa diandra hr. 
Berol. Otto. 3 of 5. , 428 [ST: LE]. 4 of 5. In arenosis 
ripae rivi Parana, Aug. 1826,s.c. s.n. [IT: K, LE]. 5 of 
5. Brazil: Parana riv. in arenosis [HT: LE-TRIN- 
2351.03]. 

= Eragrostis japonica (Thunb.) Trin. 


Eragrostis henrardii Jansen, Blumea, Suppl. 3: 42. 1946. T- 
Protol.: South-West-Africa: Grassteppe at Okahandja, 
1912, K. Dinter 2572 {HT: L]. T-Specim.: 1 of 1. Mar 
1912, MK. Dinter 2572 (HT: L]. 
= Eragrostis trichophora Coss. & Durieu. 


Eragrostis hirsuta (Michx.) Nees, Fl. Bras. Enum. PI. 2: 
508. 1829. Basionym: Poa hirsuta Michx. 
DST.: Mexi, USA. 


LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Cowan, C. P. (1983); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Fernald, M. 
(1950); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Hitchcock, A. 
S. (1951); McClure, F. A. (1955); Radford, A. E., H. 
E. Ahles & C. R. Bell (1968); Small, J. K. (1933); 
Wunderlin, R. P. (1998). 

Eragrostis hirsuta var. hirsuta; Eragrostis hirsuta 
var. laevivaginata Fernald; Eragrostis sporoboloides 
J.G. Sm. & Bush; Poa hirsuta Michx. 


SYN.: 


Eragrostis hirsuta var. laevivaginata Fernald, Rhodora 
41(490): 500-501. 1939. T-Protol.: USA: Virginia: 
Southhampton Co.: from wooded alluvial bottomland 
of Meherrin River, near Haley's Bridge, Fernald & 
Long 9273 [HT: GH; IT: PH]. 

= Eragrostis hirsuta (Michx.) Nees. 


Eragrostis hirta E. Fourn., Mexic. Pl. 2: 115. 1886. T- 
Protol.: Mexico: San Luis de Potosi, 1851, Virlet 1390. 
T-Specim.: | of 1. Mexico: San Luis Potosi, Aug 
1851, Virlet 1390 [HT: P; IT: US-77382 (fragm.)]. 

DST.: Beli, Guat, Hond, Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Balick, M. J., M. H. Nee & D. E. 
Atha (2000); Beetle, A. A. (1977); Breedlove, D. E. 
(1986); Davidse, G. (1994); Davidse, G., M. Sousa S. 
& A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); McClure, F. A. (1955); Molina R., A. (1975). 
Eragrostis intermedia var. praetermissa (L.H. Harv.) 
Witherspoon; Eragrostis praetermissa L.H. Harv. 


SYN:: 


Eragrostis hirta E. Fourn. var. hirta. 
DST.: Beli, Guat, Hond, Mexi. 
LVL. ACCPTC.: 1. 


Eragrostis hirta var. longiramea (Swallen) Witherspoon, 
Ann. Missouri Bot. Gard. 64: 328. 1977. Basionym: 
Eragrostis longiramea Swallen. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. L6pez- 
Ferrari, and J. Valdes-Reyna (2000); Witherspoon, J. 
T. (1978). 

SYN.: Eragrostis longiramea Swallen. 


Eragrostis hondurensis R.W. Pohl, lowa State J. Res. 54(3): 
319, f. 1. 1980. T-Protol.: Honduras: El Paraiso: 10.5 
km SE of Yuscaran, elev. 550 m, abandoned field near 
a small stream, 31 Dec 1977, Pohl & Gabel 13442 
[HT: ISC-329297; IT: MO]. 

DST.: Hond, Mexi, Nica. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Breedlove, D. E. (1986); Davidse, G. 
(1994); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000). 


Eragrostis horizontalis Peter, Repert. Spec. Nov. Regni Veg. 
Beih. 40(1 Lief. 2): 107, t. 68, f. 2 [in 40 (1, Lief. 4), 
1936]. 1930. T-Protol.: D. O.Afr., Ugogo: Garten in 
Dodoma, Peter 46834. 

= Eragrostis cylindriflora Hochst. 


94 Catalogue of New World Grasses: II. 


Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb., 
Prelim. Cat. 69. 1888. Basionym: Poa hypnoides Lam. 
DST.: Arge, Beli, Boli, Braz, Cana, Cari, CoRi, Ecua, ElSa, 
FrGu, Guat, Guya, Hond, Mexi, Nica, Pana, Para, 
Peru, Suri, USA, Urug, Vene. 
LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Abrams, L. (1923); Adams, C. D. 
(1972); Allem, A.C. & J. F.M. Valls (1987); Balick, 
M. J.. M. H. Nee & D. E. Atha (2000); Beetle, A. A. 
(1977); Berendsohn, W. G. & A. E. Araniva de 
Gonzalez (1991); Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Brako, L. & J. L. Zarucchi (1993); 
Breedlove, D. E. (1986); Burkart, A. (1969); Cabrera, 
A. L. (1970); Catasus Guerra, L. (1997); Correll, D. 
S. & M. C. Johnston (1970); Cowan, C. P. (1983); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Davidse, G. (1994); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000): Fernald, M. (1950); Filgueiras, 
T. S. (1991); Foster, R. C. (1958); Gleason, H. A. & 
A. Cronquist (1968); Gleason, H. A. & A. Cronquist 
(1991); Godtrey, R. K. & J. W. Wooten (1979); 
Gould, F. W (1979): Gould, F. W. & R. Moran 
(1981); Great Plains Flora Association (1986); 
Hickman, J. C. (1993); Hitchcock, A. S. (1909); 
Hitchcock, A. S. (1927); Hitchcock, A. S. (1936); 
Hitchcock, A. S. (1951); Hitcheock, C. L., A. 
Cronquist, M. Ownbey & J. W. Thompson (1969); 
Jorgensen, P. M. & S. Leon-Yanez (eds.) (1999); 
Judziewicz, E. J. (1990); Liogier, H. & L. Martorell 
(1982); Long, R. W. & O. Lakela (1971); McClure, 
F. A. (1955); McVaugh, R. (1983); Molina R., A. 
(1975); Munz, P. A. & D. D. Keck (1959); Munz, P. 
A. (1974); Pohl, R. W. (1980); Radford, A. E., H. E. 
Ahles & C. R. Bell (1968); Renvoize, S. A. (1984); 
Renvoize, S. A. (1988); Renvoize, S. A. (1998); 
Rosengurtt, B., B. R. Arrillaga de Maffei & P. 
Izaguirre de Artucio (1970); Schwegman, John E. 
(1991); Scoggan, H. J. (1978 [1979]); Small, J. K. 
(1933); Smith, L. B., D. C. Wasshausen & R. M. 
Klein (1981); Tovar, O. (1993); Voss, E. G. (1972); 

Wunderlin, R. P. (1998). 
Eragrostis reptans var, contracta Doll; Eragrostis 
reptans var. laxior Doll; Eragrostis reptans var. 
pygmaea Dll; Erosion hypnoides (Lam.) Lunell: 
Megastachya_ breviflora EE. Fourn.; Megastachya 
corymbifera EE. Fourn.; Megastachya gouinii E. 
Fourn.; Megastachya gouinti E. Fourn. ex Hemsl.; 


SYN: 


Megastachya hypnoides” (Lam.) P.  Beauv.; 
Neeragrostis hypnoides (Lam.) Bush; Poa hypnoides 
Lam. 


Eragrostis incana Maury, J. Bot. (Desvaux) 3: 163, f. 6. 
1889. T-Specim.: | of 2. Venezuela: Boca del Pao, J 
Chaffanjon 289 |ST]. 2 of 2. Venezuela: savane 
humide, Mapire, J. Chaffanjon 296 [ST]. 

= Status not yet determined. 


Eragrostis inconstans Nees, Fl. Bras. Enum, Pl. 2: 495. 1829. 
T-Protol.: Brazil: In lapidosis, ad saxa, prope Fanado, 
in minis Novis provinciae Minarum, Mart. s.n. 
NOTE: nom. incorrect; included older name Eragrostis 
rufescens in var. alpha. 
CORRECT NAME: Eragrostis rufescens Schrad. ex Schult. 
= Eragrostis rufescens Schrad. ex Schult. 


Eragrostis inconstans var. rufescens (Schrad. ex Schult.) 
Nees, Fl. Bras. Enum. Pl. 2: 495. 1829. Basionym: 
Eragrostis rufescens Schrad. ex Schult. 

NOTE: nom. incorrect, E. rufescens Schrad has_ priority 
over E. inconstans as the name for the sepcies. 
= Eragrostis rufescens Schrad. ex Schult. 


Eragrostis intermedia Hitchc., J. Wash. Acad. Sci. 23(10): 
450. 1933. T-Protol.: USA: Texas: San Antonio, in 
moist place in gravel pit, 3 Jul 1910, 4.8. Hitchcock 
5491 [HT: US-1535749]. T-Specim.: | of 1. USA: 
Texas: Bexar Co.: San Antonio, 3 Jul 1910, 4.8. 
Hitchcock 5491 [HT: US-1535749; IT: US-908993, 
US-1535750]. 

DST.: Colo, CoRi, Guat, Mexi, USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Davidse, G. (1994); Davidse, G., M. Sousa S. 
& A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Fernald, M. (1950); Gleason, H. A. & A, 
Cronquist (1968); Gleason, H. A. & A. Cronquist 
(1991); Gould, F. W. & R. Moran (1981); Great 
Plains Flora Association (1986); Hitchcock, A. S. 
(1951); McClure, F. A. (1955); McVaugh, R. (1983); 
Shreve, F. & I. L. Wiggins (1964); Wunderlin, R. P. 
(1998). 

SYN.: Eragrostis lugens var. major Vasey ex L.H. Dewey. 


Eragrostis intermedia var. appressa Witherspoon, Ann. 
Missouri Bot, Gard. 64: 325. 1977 [1978]. T-Protol.: 
Mexico, Harvey & Witherspoon 9344 [HT: US; IT: 
ENCB, MO, MONTU, NY, RM, TAES, W]. 

DST.: Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Witherspoon, J. 
T. (1978). 


Eragrostis intermedia Hitche. var. intermedia. 
DST.: CoRi, Guat, Mexi, USA. 
LVL. ACCPTC.: 1. 


Eragrostis intermedia var. oreophila) (L.H. Harv.) 
Witherspoon, Ann. Missouri Bot. Gard. 64: 326. 1977 
[1978]. Basionym: Eragrostis oreophila L.H. Harv. 

DST.: Mexi. 

LVL. ACCPTC.: |. 

ALSO ACCPTD. BY: Espejo Serna, A.. A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Witherspoon, J. 
T. (1978). 

SYN.: Eragrostis oreophila L.H. Harv. 


Eragrostis intermedia var. praetermissa (L.H. Harv.) 
Witherspoon, Ann. Missouri Bot. Gard, 64; 327, 1977 
[1978]. Basionym: Eragrostis praetermissa L.H. Harv. 

= Eragrostis hirta E. Fourn. 


Eragrostis interrupta (Lam.) DOll, Fl. Bras. 2(3): 157, 1878. 
Basionym: Poa interrupta Lam. 
NOTE: hom. illeg. 
E. NAME: Eragrostis interrupta P. Beauv. 
= Eragrostis japonica (Thunb.) Trin. 


Eragrostis interrupta (Nutt.) Trel., Rep. U.S. Geol. Geogr. 
Surv. Territ. 4: 237. 1891. Basionym: Poa interrupta 
Nutt. 
NOTE: hom. illeg. 


Catalogue of New World Grasses: II. 95 


E. NAME: Eragrostis interrupta P. Beauv. 
= Eragrostis secundiflora subsp. oxylepis (Torr.) S.D. 
Koch. 


Eragrostis interrupta var. laxiflora Doll, Fl. Bras. 2(3): 158. 
1878. T-Protol.: Martio ad littora fluminus Amazonum 
reperta — Etiam in India orientali, in Mosul, Syria, 
Nubia Abyssinia et in Africa australi obvia. 

= Eragrostis japonica (Thunb.) Trin. 


Eragrostis interrupta vat. parviflora Doll, Fl. Bras. 2(3): 158. 
1878. T-Protol.: Brazil: Prov. Goias "Goyaz", Burchell 
SM. 
= Eragrostis japonica (Thunb.) Trin. 


Eragrostis japonica (Thunb.) Trin., Mem. Acad. Imp. Sci. 
St.-Petersbourg, Ser. 6, Sci. Math. 1(4): 405. 1830. 
Basionym: Poa japonica Thunb. 

NOTE: Boechat considers Eragrostis glomerata Walter to 

be dist. 

DST.: Arge, Boli, Braz, Cari, CoRi, Ecua, ElSa, Guat, 

Guya, Hond, Mexi, Para, Peru, USA, Urug, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Baaijens, G. J. & J. F. Veldkamp 
(1991); Bor, N. L. (1960); Catasus Guerra, L. (1997); 
Cope, T. A. (1992); Jorgensen, P. M. & S. Leén- 
Yanez (eds.) (1999); Judziewicz, E. J. (1990); 
Killeen, T. J. (1990); Phillips, S. (1995); Renvoize, S. 
A. (1984); Renvoize, S. A. (1988); Renvoize, S. A. 
(1998). 

Catabrosa micrantha Hochst. ex A. Rich.; 
Diandrochloa diplachnoides (Steud.) A.N. Henry; 
Diandrochloa japonica’ (Thunb.) A.N. Henry; 
Diandrochloa namaquensis (Nees ex Schrad.) De 
Winter; Eragrostis aturensis (Kunth) Trin. ex Steud.; 
Eragrostis brasiliana Nees; Eragrostis conferta 
(Elliott) Trin.; Eragrostis diplachnoides Steud.; 
Eragrostis elegans Nees; Eragrostis elegans var. 
laxiflora Arechav.; Eragrostis glomerata (Walter) 
L.H. Dewey; Eragrostis hapalantha Trin.; Eragrostis 
interrupta (Lam.) DOll; Eragrostis interrupta var. 
laxiflora DO\l, Eragrostis interrupta var. parviflora 
D6ll; Eragrostis namaquensis Nees ex Schrad.; 
Eragrostis namaquensis var. diplachnoides (Steud.) 
Clayton; Eragrostis pallida Vasey; Glyceria 
micrantha Steud.; Megastachya aturensis (Kunth) 
Roem. & Schult.; Megastachya brasiliensis Schult. 
ex B.D. Jacks.; Megastachya glomerata (Walter) 
Schult.; Poa aturensis Kunth; Poa brasiliensis 
Spreng.; Poa glomerata Walter; Poa interrupta 
Lam.; Poa japonica Thunb.; Poa sporoboloides A. 
Rich.; Roshevitzia diplachnoides (Steud.) Tzvelev; 
Roshevitzia glomerata (Walter) Tzvelev; Roshevitzia 
Japonica (Thunb.) Tzvelev; Sporobolus confertiflorus 
A. Rich.; Sporobolus verticillatus Nees; Vilfa 
confertiflora (A. Rich.) Steud. 


SYN:: 


Eragrostis kuschelii Skottsb., Ark. Bot., n.s., 4(15): 485, t. 
21-2, 22. 1962 [1963]. 

NOTE: nom. nov., incidental mention in Ark. Bot., n.s., 
4(12): 417 (1961 [1963]) by Kuschel without citation 
of basionym publication. 

REPLACED NAME: _ Eragrostis 
macranthera Skottsb. 

DST.: Chil. 

LVL. ACCPTC.: 1. 


peruviana — var. 


ALSO ACCPTD. BY: Marticorena, C. & M. Quezada 
(1985); Nicora, E. G. (1994). 
SYN.: Eragrostis peruviana var. macranthera Skottsb. 


Eragrostis kwaiensis Peter, Repert. Spec. Nov. Regni Veg. 
Beih. 40(1) Anhang: 99. 1930. T-Specim.: 1 of 1. 
Tanzania: W Usambaras, Kwai, Apr 1911, Eichinger 
3333 [HT: EA; IT: K]. 

= Eleusine multiflora Hochst. ex A. Rich. 


Eragrostis lasseri Luces, Bol. Soc. Venez. Ci. Nat. 15: 6, f. 


3. 1953. T-Protol.: Venezuela: en  formaciones 
xerofilas ca. Maracaibo, 12 Jan 1948, 7. Lasser 2660 
[HT: VEN]. 

DST.: Vene. 


LVL. ACCPTC:: I. 


Eragrostis lehmanniana Nees, Fl. Afr. Austral. Ill, 402. 
1841. T-Protol.: Ad rupes nudas prope Slengerfontein 
in collibus ad Nieuwe Hantom, alt. 4000-5000 ft, Feb, 
Drége s.n. T-Specim.: | of 1. Cap.{ut] b.[onae] sp.[ei]. 
Drége IT: LE]. 

DST.: Arge, Braz, Mexi, USA, Vene. 

LVL. ACCPTC.: 1; cultivated, introduced. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Boechat, S$. C. & H. M. Longhi-Wagner 
(2000); Boechat, 8. C. & P. M. Peterson (1995); Bor, 
N. L. (1960); Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Hickman, J. C. (1993); Hitchcock, A. 
S. (1951). 


Eragrostis lehmannii Pilg., Bot. Jahrb. Syst. 27: 32. 1899. T- 
Protol.: Ecuador: crescit in montanis praeruptis, massa 
lava formatis, circa Banos ad flumen Tunguragua, alt. 
s. m. 1800-2500 m, Lehmann 5283, 5283 is error for 
5823, Accd. to Chase & Niles, Index Grass Sp. T- 
Specim.: | of 1. Ecuador: Circa Banos ad flumen 
Tunguragua, alt. s.m. 1800-2500 m, F.C.Lehmann 
5283 (error for 5823) (IT: US-2767406 (fragm.)]. 

= Eragrostis pastoensis (Kunth) Trin. 


Eragrostis leptantha Trin., Mem. Acad. Imp. Sci. St.- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 405. 1830. T- 
Protol.: V. subsp. Brasilia, in siccis arenosis. T- 
Specim.: 1 of 1. Brazil: in siccis arenosesum pr. Moji 
{[Maji, Majr?, Sao Paulo], 23 Apr, /Riedel s.n.] G.H. 
von Langsdorff s.n. (HT: LE-TRIN-2361.01; IT: BAA- 
1043 (fragm.), K, LE, LE, LE, MO-2111141, P, US- 
2941514 (fragm.)]. 

= Eragrostis mexicana (Hornem.) Link. 


Eragrostis leucosticta Nees ex DOoll, Fl. Bras. 2(3): 144. 
1878. T-Specim.: | of 3. Brazil: in altis auriferis prope 
Itambe de Matte Dentro aliisque ejus regionalis locis, 
Apr-May, K.F.P. Martius 509] (ST: BAA-4601, LE- 
TRIN-2362.01, IST: US-82041]. 2 of 3. Caldas prov. 
Minarum, Mar, Regnell Ill, 14/3 [ST: P]. 3 of 3. 
Brazil, Schott 4850 [ST]. 

DST.: Braz. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Dll, J. C. (1878); Filgueiras, T. S. 
(1999); Renvoize, S. A. (1988). 


Eragrostis limbata E. Fourn., Bull. Soc. Bot. France 27: 295. 
1880. 
NOTE: nom. nud. 


96 Catalogue of New World Grasses: II. 


= Eragrostis mexicana subsp. mexicana. 


Eragrostis limbata E. Fourn., Mexic. Pl. 2: 116. 1886. T- 
Specim.: 1 of 5. Mexico: in Scopuloris Regnii 
Mexicana, 1070 mt, 1833, 4.J.4. Bonpland 4573 (LT: 
US-2941517 (fragm.)], LT designated by ?; cited by 
McVaugh, FI. Novo-Galic. 14: 168 (1983). 2 of 5. 
Mexico: Veracruz: 1070 mt, 20 Jul 1865, EP. 
Fournier 266 [ST: US-2941518 (fragm.)]. 3 of 5. 
Mexico: Vallee Mexico, J.L. Berlandier 765 [ST: US- 
2850913]. 4 of 5. Mexico: Tacubaya, W. Schaffner 72 
[ST: US-2850914 (fragm.)]. 5 of 5. Mexico: Cerro de 
los Banos, Valle de Mexico, 7 July 1865,£. Bourgeau 
437 (ST: US-911297 (fragm.)]. 

NOTE: cites Poa eragrostis Willd. msc. in herb. Mus. 
Parr.,. 
= Eragrostis mexicana subsp. mexicana. 


Eragrostis limbata var. major E. Fourn., Mexic. Pl. 2: 116. 
1886. T-Specim.: | of 3. Mexico, J.L.Berlandier 935 
[ST: P; IST: US-2850911 (fragm.)]. 2 of 3. Mexico: 
Region de Orizaba, 12 Jul 1866, E. Bourgeau 2635 
[ST: US-1614955]. 3 of 3. Mexico: San Luis de Potosi, 
M. Virlet D'Aoust 1386 [ST: P; IST: US-2850912 
(fragm.), US-2941516 (fragm.)]. 
NOTE: E. tephrosanthos Steud. in part, in: Biol. Cent.- 
Amer. Bot. 3: 573 (1883). 
= Eragrostis mexicana (Hornem.) Link. 


Eragrostis lindeniana Steud., Syn. Pl. Glumac. 1; 278. 1854. 
T-Protol.: Venezuela "N. Granada": Carabobo, Linden 
1554. T-Specim.: 1 of 1. Venezuela: Carabobo, 
J.L.Linden 1554 (IT: US-2941520 (fragm.)]. 
= Eragrostis maypurensis (Kunth) Steud. 


Eragrostis lindmanii Hack., Kongl. Svenska Vetensk. Acad. 
Handl, 34(6): 19, t. LOA. 1900. T-Protol.: Hab. in 
Brasilia austr., Rio Grande do Sul, Porto Alegre, in 
pascuis siccioribus (campo dictis), mens Sept. florens 
et fructificans, Exp. / Regnell., A, 267 (HT: S]. T- 
Specim.: | of 1. Brazil: Rio Grande do Sul, Regnell A, 
267 (HT: S; IT: P (large plant only), US-703 146]. 

NOTE: as Eragrostis "Lindmani”. 
= Eragrostis neesii var. lindmanii (Hack.) Ekman. 


Eragrostis linkii (Kunth) Steud., Syn. Pl. Glumac. 1: 273. 
1854. Basionym: Poa linkii Kunth. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Eragrostis filiformis Link. 
= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis longeradiata Steud., Syn. Pl. Glumac. |: 272. 
1854. T-Protol.: USA: Carolina, M.A. Curtis s.n. 
= Eragrostis refracta (Muhl. ex Elliott) Scribn. 


Eragrostis longipila Hack., Anales Mus. Nac. Hist. Nat. 
Buenos Aires 21: 132. 1911. T-Protol.: Argentina: 
Rioja: Dep. San Martin: Estancia La Diana, 3 Mar 
1907, Stuckert 17090. T-Specim.: 1 of 1. Argentina: 
La Rioja, 3 Mar 1907, 7..V. Stuckert 17090 (STUCK. 
HERB. NO.) |IT: BAA (fragm.), CORD, SI, US- 
911821, US-2941521]. 

= Eragrostis orthoclada Hack. 


Eragrostis longiramea Swallen, J. Wash. Acad. Sci. 21(17): 
437. 1931. T-Protol.: Mexico: Tamaulipas: on Pico del 
Diablo, vicinity of Marmolejo, Sierra de San Carlos, 
12 Aug 1930, 1. H. Bartlett 10910 [HT: US-1501524]. 
T-Specim.: | of 1. Mexico: Tamaulipas, 12 Aug 1930, 


H.H. Bartlett 10910 [HT: US-1501524; IT: GH, 
MICH, US-1611156, US-3160925]. 

= Eragrostis hirta’ var. longiramea 
Witherspoon. 


(Swallen) 


Eragrostis lugens Nees, Fl. Bras. Enum. Pl. 2: 505-506. 
1829. T-Protol.: Habitat ad Monte-video et in 
confinibus regni Paraguayani, Se//ow s.n. [HT: B]. T- 
Specim.: | of 1. Uruguay: Montevideo, no date, F. 
Sellow s.n. {[LT: US-732957; ILT: B, BM (possible) 
BAA-2932], LT designated by Witherspoon, Ph. D. 
thesis (1975) [not effectivly published]; sheet label 
cites location as Brazil. 

DST.: Arge, Boli, Braz, Chil, Ecua, ElSa, Guat, Mexi, Para, 
Peru, USA, Urug, Vene. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 

(1987); Anonymous (1986); Beetle, A. A. (1977); 

Berendsohn, W. G. & A. E. Araniva de Gonzalez 

(1991); Boechat, S. C. & H. M. Longhi-Wagner 

(2000); Breedlove, D. E. (1986); Burkart, A. (1969); 

Cabrera, A. L. (1970); Correll, D. S. & M. C. 

Johnston (1970); Davidse, G. (1994); Davidse, G., 

M. Sousa S. & A. O. Chater (editores generales) 

(1994): Doll, J. C. (1878); Espejo Serna, A., A. R. 

Lépez-Ferrari, and J.  Valdes-Reyna (2000); 

Filgueiras, T. S. (1999); Foster, R. C. (1958); 

Hitchcock, A. S. (1927); Hitchcock, A. S. (1951); 

Jorgensen, P. M. & S. Leén-Yanez (eds.) (1999); 

Killeen, T. J. (1990); Macbride, J. F. (1936); 

McClure, F. A. (1955); Nicora, E. G., M. E. De 

Paula, A. M. Faggi, M. del C. Mariano, A. M. Miante 

A., L. R. Parodi, C. A. Petetin, F. A. Roig & Z. 

Rugolo de Agrasar (1978); Radford, A. E., H. E. 

Ahles & C. R. Bell (1968); Renvoize, S. A. (1984); 

Renvoize, S. A. (1988); Renvoize, S. A. (1998); 

Rosengurtt, B., B. R. Arrillaga de Maffei & P. 

Izaguirre de Artucio (1970); Small, J. K. (1933); 

Smith, L. B., D. C. Wasshausen & R. M. Klein 

(1981); Tovar, O. (1993); Wunderlin, R. P. (1998). 

Eragrostis flaccida Lindm.; Eragrostis lugens fo. 

pallida Hack.; Eragrostis lugens subsp. flaccida 

(Lindm.) Hack.; Eragrostis lugens subsp. lugens; 

Eragrostis lugens var. glabrata Dé6ll, Eragrostis 

lugens var. glabrescens DOll; Eragrostis pilosa var. 

lugens (Nees) Griseb.; Eragrostis pilosa var. lugens 

Kuntze; Poa /ugens (Nees) Kunth. 


SYN.: 


Eragrostis lugens subsp. flaccida (Lindm.) Hack., Anales 
Mus. Nac. Buenos Aires 21: 134. 191]. Basionym: 
Eragrostis flaccida Lindm. 

= Eragrostis lugens Nees. 


Eragrostis lugens var. glabrata Doll, Fl. Bras. 2(3): 141. 
1878. T-Protol.: Brazil, Regne// s.n. T-Specim.: | of 1. 
Brazil: Minas Gerais, 2 Nov 1861, 4.F. Regnell Ex. 
Herb. Regn. Ser. 3 No. 1404 (ILT: US-911911], LT 
designated by ?. 

= Eragrostis lugens Nees. 


Eragrostis lugens var. glabrescens DOll, Fl. Bras. 2(3): 140. 
1878. T-Protol.: Brasilia: ad Curvello et ad Lagoa 
Santa ejusd. provinciae, Warming s.n., also cites 
Regnell and Weddell collections. T-Specim.: | of 1. 
Brazil, 20 Mar 1868, 4.F. Regnell 1405 |ILT: US- 
911912, US-911847 (fragm.)], LT designated by ?. 

= Eragrostis lugens Nees. 


Catalogue of New World Grasses: II. 97 


Eragrostis lugens var. major Vasey ex L.H. Dewey, Contr. 
U.S. Natl. Herb. 2(3): 542. 1894. T-Protol.: USA: Low 
prairies. 

= Eragrostis intermedia Hitchce. 


Eragrostis lugens fo. pallida Hack., Anales Mus. Nac. 
Buenos Aires 21: 134. 1911. T-Protol.: Argentina: 
Carnerillo, F.C. And., Dep. Rio IV. C., Stuckert 20053. 
T-Specim.: | of |. Argentina: Rio Negro, Apr 1909, 
T.JV. Stuckert 20053 (IT: US-2941522 (fragm.)]. 

= Eragrostis lugens Nees. 


Eragrostis lugens var. villosa Déll, Fl. Bras. 2(3): 140. 1878. 
T-Protol.: Brazil, no collection specified for this var. 
= Eragrostis polytricha Nees. 


Eragrostis lurida J. Presl, Relig. Haenk. 1(4-5): 276. 1830. 
T-Protol.: Hab. in montanes Peruviae, Haenke s.n. T- 
Specim.: | of 1. Peru, 7) Haenke s.n. [HT: PR; IT: 
MO-2111146, US-2941523 (fragm.)]. 

DST.: Arge, Boli, Ecua, Peru. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Foster, R. C. (1958); Hitchcock, A. 
S. (1927); Jorgensen, P. M. & C. Ulloa U. (1994); 
Jorgensen, P. M. & S. Leén-Yanez (eds.) (1999); 
Macbride, J. F. (1936); Renvoize, S. A. (1998); 
Tovar, O. (1993). 

Eragrostis bahiensis var. boliviensis Henrard; 
Eragrostis contracta Pilg.; Eragrostis contristata 
Meyen; Poa lurida (J. Pres!) Kunth. 


SYN.: 


Eragrostis luzoniensis Steud., Syn. Pl. Glumac. 1: 266. 1854. 
T-Protol.: Ins. Luzon, Cuming. 1416. T-Specim.: | of 
1. Philippines: Luzon, H. Cuming 1416 [T: L]. 
= Eragrostis gangetica (Roxb.) Steud. 


Eragrostis macra Jedwabn., Bot. Arch. 5(3-4): 200. 1924. T- 
Specim.: | of 6. Brazil: without further locality 
information, 3 Nov 1887, 4.F.M. Glaziou 16624 [ST: 
C, US-289924, US-1280048, US-1280047]. 2 of 6. 
Brazil: loco non indicato, Glaziou 16623 [ST: C]. 3 of 
6. Brazil: Santa Catarina: ad Itajahy, Schenck 1093 
[ST]. 4 of 6. Paraguay: loco non indicato, Fiebrig s.n. 
[ST]. 5 of 6. Paraguay: in Sierra Amambay, Hassler 
9973 [ST: B, BAA-1035 (fragm. ex B), K, LIL- 
45973]. 6 of 6. Paraguay: in Sierra Amambay, Hassler 
11638 [ST]. 

= Eragrostis bahiensis Schrad. ex Schult. 


Eragrostis macropoda Pilg., Symb. Antill. 4: 106. 1903. T- 
Protol.: Porto Rico: Bayamon in litoralibus ad Catafio, 
Sintensis 1233. T-Specim.: 1 of 1. Puerto Rico: 
Batamon, in litoralibus ad Catafio, 27 Mar 1885, 
P.ELE. Sintenis 1233 [HT [ST?]: B; IT: NY-70977, 
US-821975, US-2941525 (fragm.)]. 

= Eragrostis elliottii S. Watson. 


Eragrostis macrothyrsa Hack., Repert. Spec. Nov. Regni 
Veg. 8: 47. 1910. T-Protol.: Paraguay septen. a. 1909, 
Fiebrig 5166 [HT: Hb. Hassler]. T-Specim.: | of 1. 
Paraguay: Paraguay septentrionale, Sep 1909, K. 
Fiebrig 5166 [IT: BAA-1047, K, US-1099649, US- 
2764900]. 

DST.: Boli, Braz, Para. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Boechat, S. C. & P. M. Peterson 
(1995); Killeen, T. J. (1990); Renvoize, S. A. (1998). 


Eragrostis magna Hitchc., Contr. U.S. Natl. Herb. 24(8): 
341. 1927. T-Protol.: Peru: collected on steep rocky 
grasslands at Huacachi Estacion near Muna, alt. 2000 
m, 20 May to | June 1923, /.F. Macbride 4069 [HT: 
F-535136]. T-Specim.: | of 1. Peru: near Muna, 20 
May-! Jun 1923, /.F. MacBride 4069 [HT: F-535136; 
IT: US-1256382]. 

DST.: Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Hitchcock, A. S. (1927); Macbride, 
J. F. (1936); Tovar, O. (1993). 


Eragrostis major Host, Icon. Descr. Gram. Austriac. 4: 14, t. 
24. 1809. 
NOTE: nom. nov., cites Briza eragrostis L. 
REPLACED NAME: Briza eragrostis L. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis major Host, Fl. Austriac. 1: 135. 1827. 
NOTE: isonym. 
E. NAME: Eragrostis major Host. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis mattogrossensis Pilg., Bot. Jahrb. Syst. 30(1): 140. 
1902. T-Protol.: Brasil: Mattogrosso:  zerstreut 
zwischenSandsteinblocken am trocknen Uferrand des 
Quellflusses des Ronuro, May 1899,H. Meyer 558. T- 
Specim.: | of 1. Brazil: Mato Grosso: Ronuro, May 
1899, H. Meyer 588 [IT: US-2767405 (fragm.)]. 

= Eragrostis solida Nees. 


Eragrostis mattogrossensis f0. glabrescens Pilg., Bot. Jahrb. 
Syst. 30(1): 141. 1902. T-Specim.: 1 of 2. Brazil: 
Mattogrosso: zerstreut an sandigen, offener Stellen im 
Serrado bei Cuyaba, Feb 1899, H. Meyer 223 [ST: 
BAA-1049 (fragm. ex B)]. 2 of 2. Brazil: Mattogrosso: 
auf offener Sandflache an der Bandeira bei Cuyaba, 
Mar 1899, H. Meyer 336 [ST]. 

= Eragrostis solida Nees. 


Eragrostis maxima E. Fourn., Mexic. Pl. 2: 114. 1886. T- 

Specim.: 1 of 3. Mexico: S. Jean d'Ulloa, Gouin 9] 
[ST: US-A2767419]. 2 of 3. Mexico: St. Agustin, Oct 
1842, F.M. Liebmann 482 [ST: US-207541]. 3 of 3. 
Saint-Dominique, Poiteau s.n. [ST: Herb. Juss. ]. 

NOTE: hom. illeg.; also E. maxima Fourn. in Helmsley, 

Biol. Centr. Amer. Bot. 3: 574 (1885). 
E. NAME: Eragrostis maxima Baker. 
= Eragrostis prolifera (Sw.) Steud. 


Eragrostis maypurensis (Kunth) Steud., Syn. Pl. Glumac. 1: 
276. 1854. Basionym: Poa maypurensis Kunth. 

DST.: Beli, Boli, Braz, Colo, CoRi, ElSa, FrGu, Guat, 
Guya, Hond, Mexi, Nica, Pana, Peru, Suri, USA, 
Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Balick, M. J.. M. H. Nee & D. E. 
Atha (2000); Beetle, A. A. (1977); Berendsohn, W. 
G. & A. E. Araniva de Gonzalez (1991); Boechat, S. 
C. & H. M. Longhi-Wagner (2000); Brako, L. & J. L. 
Zarucchi (1993); Breedlove, D. E. (1986); Cowan, C. 
P. (1983); Davidse, G. (1994); Davidse, G., M. Sousa 
S. & A. O. Chater (editores generales) (1994); DOIl, 
J. C. (1878); Espejo Serna, A., A. R. Lopez-Ferrari, 
and J. Valdes-Reyna (2000); Filgueiras, T. S. (1991); 
Filgueiras, T. S. (1999); Foster, R. C. (1958); 
Hitchcock, A. S. (1927); Judziewicz, E. J. (1990); 
Killeen, T. J. (1990); McClure, F. A. (1955); 


98 Catalogue of New World Grasses: II. 


McVaugh, R. (1983); Molina R., A. (1975); Nelson, 
C. (1978); Pohl, R. W. (1980); Renvoize, S. A. 
(1984); Reyes-Garcia, A. & M. Sousa S. (1997); 
Tovar, O. (1993). 

Eragrostis acuminata Doll, Eragrostis amoena J. 
Presl; Eragrostis lindeniana Steud.; Eragrostis 
maypurensis (Kunth) Déll; Eragrostis maypurensis 
var. densiuscula Pilg.; Eragrostis maypurensis vat. 
meratiana (Kunth) Pilg.; Eragrostis meratiana 
(Kunth) Steud.; Eragrostis panamensis J. Presl; 
Eragrostis vahlii (Roem. & Schult.) Nees; Eragrostis 
vahlii var. polyantha Doll; Megastachya amoena E. 
Fourn. ex Hemsl.; Megastachya maypurensis (Kunth) 
Roem. & Schult.; Megastachya panamensis (J. Presl) 
E. Fourn.; Megastachya panamensis E. Fourn, ex 
Hemsl.;  Megastachya swainsonii Raddi; Poa 
maypurensis Kunth; Poa racemosa Wahl; Poa 
squarrosa Salzm. ex Steud.; Poa vahlii Roem. & 
Schult. 


SYN:: 


Eragrostis maypurensis (Kunth) Doll, Fl. Bras. Enum. PI. 
2(3): 153. 1878. Basionym: Poa maypurensis Kunth. 
NOTE: isonym. 
E. NAME: Eragrostis maypurensis (Kunth) Steud. 
= Eragrostis maypurensis (Kunth) Steud. 


Eragrostis maypurensis var. densiuscula Pilg., Bot. Jahrb. 
Syst. 70: 349. 1939. T-Protol.: Brazil: Sao Paule: rio 
Clare, auf Campo, Jun 1888, Lofgren 576. T-Specim.: 
| of 1. Brazil: So Paulo, 5 Jun 1888,4. Loefgren 576 
(IT: US-911855). 

= Eragrostis maypurensis (Kunth) Steud. 


Eragrostis maypurensis var. meratiana (Kunth) Pilg., Bot. 
Jahrb. Syst. 70: 348. 1939. Basionym: Poa meratiana 
Kunth. 

= Eragrostis maypurensis (Kunth) Steud. 


Eragrostis megapotamica (Spreng.) Schult., Mant. 3(add. 1): 
617. 1827. Basionym: Poa megapotamica Spreng. 
= Eragrostis purpurascens (Spreng.) Schult. 


Eragrostis megastachya (Koeler) Link, Hort. Berol. 1: 187. 
1827. Basionym: Poa megastachya Koeler. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis megastachya var. cilianensis (All.) Asch. & 
Graebn., Syn. Mitteleur. Fl. 2: 371. 1900. Basionym: 
Poa cilianensis All. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis megastachya fo. nana Lorentz & Niederl., Exped. 
Rio Negro, Bot. 272. 1881. T-Protol.: Argentina. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis mendozina (Phil.) Jedwabn., Bot. Arch. 5(3-4): 
192. 1924. Basionym: Jpnum mendocinum Phil. 
= Leptochloa dubia (Kunth) Nees. 


Eragrostis meratiana (Kunth) Steud., Syn. Pl. Glumac. 1: 
276. 1854. Basionym: Poa meratiana Kunth. 
= Eragrostis maypurensis (Kunth) Steud. 


Eragrostis mexicana (Hornem.) Link, Hort. Berol. 1: 190. 
1827. Basionym: Poa mexicana Hornem. 
DST.: Arge, Boli, Braz, Cana, Chil, CoRi, Ecua, ElSa, Guat, 
Hond, Mexi, Nica, Pana, Peru, USA, Urug, Vene. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Abrams, L. (1923); Anonymous 
(1986); Beetle, A. A. (1977); Berendsohn, W. G. & 


A. E. Araniva de Gonzalez (1991); Brako, L. & J. L. 
Zarucchi (1993); Breedlove, D. E. (1986); Correll, D. 
S. & M. C. Johnston (1970); Davidse, G. (1994); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Dll, J. C. (1878); Edgar, E., H. E. 
Connor & J. E. Shand (1991); Espejo Serna, A., A. 
R. L6pez-Ferrari, and J. Valdes-Reyna (2000); 
Fernald, M. (1950); Foster, R. C. (1958); Gleason, H. 
A. & A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Gould, F. W. & R. Moran (1981); 
Hitchcock, A. S. (1927); Hitchcock, A. S. (1951): 
Jorgensen, P. M. & S. Leén-Yanez (eds.) (1999); 
Kerguelen, M. (1993); Koch, S. D. (1978); 
Lawesson, J. E., H. Adsersen & P. Bentley (1987); 
Marticorena, C. & M. Quezada (1985); McClure, F. 
A. (1955); MeVaugh, R. (1983); Molina R., A. 
(1975); Munz, P. A. & D. D. Keck (1959); Munz, P. 
A. (1974); Nicora, E. G., M. E. De Paula, A. M. 
Faggi, M. del C. Mariano, A. M. Miante A., L. R. 
Parodi, C. A. Petetin, F. A. Roig & Z. Rugolo de 
Agrasar (1978); Pohl, R. W. (1980); Porter, D. M. 
(1983); Reeder, J. R. & C. G. Reeder (1971); Shreve, 
F. & I. L. Wiggins (1964); Tovar, O. (1993); 
Wiggins, I. L. & D. M. Porter (1971); Wunderlin, R. 
P. (1998). 

SYN.: Eragrostis alba J. Presl; Eragrostis chilensis (Moris) 
Nees; Eragrostis cordobensis Jedwabn.; Eragrostis 
leptantha Trin.; Eragrostis limbata E. Fourn.; 
Eragrostis limbata var. major E. Fourn.; Eragrostis 
mexicana var. pygmaea DO6ll; Eragrostis pauciflora 
Trin. ex E. Fourn.; Eragrostis pauciflora Trin. ex E. 
Fourn. ex Hemsl.; Eragrostis purshii var. major E. 
Fourn.; Eragrostis rahmeri Phil.; Poa chilensis 
Moris; Poa mexicana Hornem. 


Eragrostis mexicana (Hornem.) Link subsp. mexicana. 

DST.: Arge, Boli, Braz, Cana, Chil, CoRi, Ecua, ElSa, Guat, 
Hond, Mexi, Nica, Pana, Peru, USA, Urug, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Brako, L. & J. L. Zarucchi (1993); 
Davidse, G. (1994); Davidse, G., M. Sousa S, & A, 
O. Chater (editores generales) (1994); Hickman, J. C. 
(1993); Sanchez Vega, I. & S. D. Koch (1988). 

SYN.: Eragrostis limbata E. Fourn.; Eragrostis limbata E. 
Fourn.; Eragrostis mexicana var. mexicana; 
Eragrostis neomexicana Vasey ex L.H. Dewey. 


Eragrostis mexicana var. pilosior DOll, Fl. Bras. 2(3): 143. 
1878. T-Protol.: Uruguay: Prope Montevideo, Sel/o 
SM. 
ALSO ACCPTD. BY: Déll, J. C. (1878). 
= Status not yet determined. 


Eragrostis mexicana var. pygmaea Doll, Fl. Bras. 2(3): 143. 
1878. T-Protol.: Uruguay: Prope Montevideo, Se/lo 
S.A. 
= Eragrostis mexicana (Hornem.) Link. 


Eragrostis mexicana subsp. virescens (J. Presl) S.D. Koch 
& Sanchez Vega, Phytologia 58: 380. 1985. 
Basionym: Eragrostis virescens J. Presl. 

DST.: Arge, Boli, Braz, Cana, Chil, Ecua. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Boechat, S.C. 
& H. M. Longhi-Wagner (2000); Brako, L. & J. L. 


Catalogue of New World Grasses: II. 99 


Zarucchi (1993); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Hickman, J. C. 
(1993); Sanchez Vega, I. & S. D. Koch (1988). 

Eragrostis cordobensis Jedwabn.; Eragrostis 
delicatula’ Trin.; Eragrostis leptantha — Trin.; 
Eragrostis orcuttiana Vasey; Eragrostis pilosa var. 
delicatula (Trin.) Hack.; Eragrostis rahmeri Phil.; 
Eragrostis virescens J. Presl; Poa virescens (J. Presl) 
Kunth. 


SYN.: 


Eragrostis microstachya (Link) Link, Hort. Berol, 2: 294. 
1933. Basionym: Poa microstachya Link. 
= Eragrostis bahiensis Schrad. ex Schult. 


Eragrostis minor Host, Icon. Descr. Gram. Austriac. 4: 15. 
1809, 

REPLACED NAME: Poa eragrostis L. 

BLOCKING NAME: Eragrostis eragrostis (L.) P. Beauv. 

DST.: Arge, Braz, Cana, Mexi, USA, Urug, Vene. 

LVL. ACCPTC.: |; introduced. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Boechat, S. C. 
& H. M. Longhi-Wagner (2000); Burkart, A. (1969); 
Cabrera, A. L. (1970); Cope, T. A. (1992); Edgar, E., 
H. E. Connor & J. E. Shand (1991); Gibbs Russell, 
G. E, W. G. Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987); Gleason, H. A. & A. 
Cronquist (1991); Great Plains Flora Association 
(1986); Hickman, J. C. (1993); Kerguelen, M. 
(1993); Koch, S. D. (1978); Phillips, S. (1995); 
Wunderlin, R. P. (1998). 

Eragrostis eragrostis (L.) H. Karst.; Eragrostis 
eragrostis (L.) P. Beauv.; Eragrostis eragrostis var. 
microstachya (Coss. & Germ.) Farw.; Eragrostis 
multiflora var. pappiana Chiov.; Eragrostis pappiana 
(Chiov.) Chiov.; Eragrostis pilosa var. minor (Host) 
Kuntze; Eragrostis poaeoides P. Beauv. ex Roem. & 
Schult.; Eragrostis poaeoides var. laxiflora Ddll; 
Eragrostis poiformis Link; Eragrostis vulgaris Coss. 
& Germ.; Eragrostis vulgaris J. Presl ex Steud.; 
Eragrostis vulgaris subsp. poaeoides (P. Beauv. ex 
Roem. & Schult.) R.C.V. Douin; Eragrostis vulgaris 
var. microstachya Coss. & Germ.; Poa eragrostis L.; 
Poa poaeiodes P. Beauv. 


SYN.: 


Eragrostis minor var. megastachya (Koeler) Burtt Davy, FI. 
W. Calif. 60. 1901. Basionym: Poa megastachya 
Koeler. 

= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis minor var. suaveolans (A.K. Becker ex Claus) 
Schmalh., Izv. Imp. Bot. Sada Petra Velikago 
14(Suppl. 2): 68. 1915. Basionym: Eragrostis 
suaveolens A.K. Becker ex Claus. 

= Eragrostis suaveolens A.K. Becker ex Claus. 


Eragrostis mokensis Pilg., Bot. Jahrb. Syst. 51(3-4): 419. 
1914. T-Protol.: Fernando Po: Grasland (Bergweiden) 
von Moka im Siidosten der Insel, 1200-1800 m, Nov 
1911, Mildbraed 7/02. T-Specim.: 1 of |. Equatorial 
Guinea: Fernando Po, no date, J. Mildbraed 7102 (HT: 
B; IT: US-2941533]. 

DST.: Braz, Vene. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Boechat, S. C. & P. M. Peterson 
(1995). 

SYN.: Eragrostis moritzii Jedwabn. 


Eragrostis montufari (Kunth) Steud., Nomencl. Bot. (ed. 2) 
1: 563. 1840. Basionym: Poa montufari Kunth. 
NOTE: see also Eragrostis montufari (Kunth) Steud., Syn. 
Pl. Glum. 1: 275 (1854), isonym. 
= Eragrostis pastoensis (Kunth) Trin. 


Eragrostis moritzii Jedwabn., Bot. Arch. 4: 328. 1923. T- 
Protol.: Venezuela, Merida, in der Alpinen Region, 
Moritz 1570. T-Specim.: 1 of 1. Venezuela: Merida, 
J.W.K. Moritz 1570 (IT: US-2941535 (fragm.)]. 

= Eragrostis mokensis Pilg. 


Eragrostis multicaulis Steud., Syn. Pl. Glumac. 1: 426. 1854. 
T-Protol.: Japonia. 
= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis multiflora (Forssk.) Asch., Fl. Brandenburg 1: 
841. 1864. Basionym: Poa multiflora Forssk. 
NOTE: hom. illeg. 
E. NAME: Eragrostis multiflora Trin. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis multiflora var. cilianensis (All.) Maire, Bull. Soc. 
Hist. Nat. Afrique N. 30: 369. 1939, Basionym: Poa 
cilianensis All. 

= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis multiflora var. glandulifera Chiov., Annuario 
Reale Ist. Bot. Roma 8: 370. 1908. T-Protol.: Assaorta: 
lungo il torrente Haddas,? 26//. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis multiflora var. insularis A. Terracc., Annuario 
Reale Ist. Bot. Roma 8: 65. 1903. T-Protol.: 
Arcipelago Dahalak: Isola Dahalak Cumbeibak Ras 
Antalo 11-12 Apr 1892, Terracciano 778 [HT: 
Romani]. 

= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis multiflora var. pappiana Chiov., Annuario Reale 
Ist. Bot. Roma 8: 65. 1903. T-Protol.: Samhar: Saati, 8 
Mar 1892, Terracciano & Pappi 2868 |HT: P]. 
= Eragrostis minor Host. 


Eragrostis multiflora var. subbiloba Chiov., Annuario Reale 
Ist. Bot. Roma 8: 370. 1908. T-Specim.: 1 of 12. 
Dembelas: Dintorni di Ferfer, Pappi 6/56 [ST]. 2 of 
12. Dembelas: lunge il Mai Makatar, Pappi 6057 [ST]. 
3 of 12. Dembelas: Mai Giahan, Pappi 6625 [ST]. 4 of 
12. Beni Amer: Monte Damba, Pappi 5964 [ST]. 5 of 
12. Beni Amer: Carajai, Pappi 6/86 [ST]. 6 of 12. 
Beni Amer: Carajai, Pappi 6346 [ST]. 7 of 12. Beni 
Amer: Carajai, Pappi 6374 [ST]. 8 of 12. Beni Amer: 
Carajai, Pappi 6641 [ST]. 9 of 12. Beni Amer: lungo il 
flume Scetel presso Dalul, Pappi 6453 [ST]. 10 of 12. 
Saraé: lungo il flume Mareb presso Debarroa, Pappi 
412 [ST]. 11 of 12. Medriod Tesfa: Mumat Mahader, 
Pappi 6675 {ST}. 12 of 12. Medriod Tesfa: Mumat 
Mahader, Pappi 7248 [ST]. 

= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis multipes 8. Moore, Trans. Linn. Soc. London 4: 
511. 1895. T-Protol.: Brazil: Hab. In cacumine 
montium Serra de Chapada prope Santa Anna de 
Chapada, Matto Grosso Exped, Spencer Le M. Moore, 
Botanist 13] (HT: BM]. 

= Eragrostis rufescens var. rufescens. 


Eragrostis multipilosa Hochst. ex Mattei, Boll. Reale Orto 
Bot. Giardino Colon. Palermo 7: 29-36. 1909. T- 


100 


Catalogue of New World Grasses: II. 


Protol.: Ethiopia, Schimper 117.= Herb. Buchinger 
772, PL ex Agow 2279 [IT:K]. 
= Eragrostis cylindriflora Hochst. 


Eragrostis namaquensis Nees ex Schrad., Linnaea 12(4): 
452. 1838. T-Protol.: Cap. b. sp., Drége s.n. T- 
Specim.: | of 1. South Africa: Namaqualand, Drége 
2569 |HT: B; IT: K.P, PRE (fragm. ex B)]. 

= Eragrostis japonica (Thunb.) Trin. 


Eragrostis namaquensis var. diplachnoides (Steud.) Clayton, 
Kew Bull. 25(2): 251. 1971. Basionym: Eragrostis 
diplachnoides Steud. 

= Eragrostis japonica (Thunb.) Trin. 


Eragrostis neesii Trin., Mem. Acad. Imp. Sci. St.- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 405. 1830. T- 
Protol.: V. subsp. Brasilia. T-Specim.: 1 of 1. Brazil: 
Brasilia meridionalis, /\Se/low [HT: LE-TRIN- 
2369.02; IT: US-2767412 (fragm.)]. 

DST.: Arge, Boli, Braz, Para, Urug. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Boechat, S$. C. & H. M. Longhi- 
Wagner (2000); Burkart, A. (1969); Doll, J.C. 
(1878); Killeen, T. J. (1990); Renvoize, S. A. (1988); 
Renvoize, S. A. (1998); Rosengurtt, B.. B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970). 

SYN.: Eragrostis villosa Salzm. ex Steud.; Poa neesii 
(Trin.) Kunth. 


Eragrostis neesii var. expansiflora Nicora, Revista Argent. 
Agron. 7: 258, 265, f. 3. 1940. T-Protol.: Argentina: 
Santa Fé: costa, en praderas virgenes, 3 Jul 1938, A. 
Burkart 9061 [HT: SI]. T-Specim.: | of 1. Argentina: 
Santa Fé: costa, en pradereas virgenes con Acacia 
cavenia, Trifolium polymorphum, Polygala linoides, 
Ornithopus micranthus y Dichondra repens, 3 Jul 
1938, A. Burkhart 9061 (HT: SI; IT: BAA]. 

DST.: Arge, Para. 
LVL. ACCPTC:: 1. 
ALSO ACCPTD. BY: Burkart, A. (1969). 


Eragrostis neesii var. laxa Jedwabn., Bot. Arch. 5(3-4): 206. 
1924. T-Specim.: | of 6. Paraguay: inter Rio Apa et 
Rio Aquidaban, 1908-1909, K. Fiebrig 5106 [ST: G, 
US-1645643]. 2 of 6. Brazil: loco non indicato, 
Glaziou 20/43 [ST]. 3 of 6. Brazil: loco non indicato, 
F. Sellow s.n. [ST]. 4 of 6. Brazil: Rio Grande do Sul: 
ad Neu-Wiirttemberg, Bornmiiller 401 [ST]. 5 of 6. 
Brazil: Minas Gerais: loco non indicato, Mendonga 
335 [ST]. 6 of 6. Brazil: Sao Paulo: Jacarahy, 
Mendonga 684 |ST]. 

= Eragrostis articulata (Schrank) Nees. 


Eragrostis neesii var. lindmanii (Hack.) Ekman, Ark. Bot. 
13(10): 51. 1913. Basionym: Eragrostis lindmanii 
Hack. 

DST.: Arge, Boli, Braz, Para, Urug. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Burkart, A. (1969); Cabrera, A. L. 
(1970); Rosengurtt, B., B. R. Arrillaga de Maffei & 
P. Izaguirre de Artucio (1970). 

SYN.: Eragrostis lindmanii Hack. 


Eragrostis neesii Trin. var. neesii. 
DST.: Arge, Boli, Braz, Para, Urug. 
LVL. ACCPTC.: 1. 


ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Burkart, A. (1969); Longhi-Wagner, 
H. M. & I. I. Boldrini (1988); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970). 


Eragrostis neomexicana Vasey ex L.H. Dewey, Contr. U.S. 
Natl. Herb. 2(3): 542. 1894. T-Protol.: USA: Mountain 
sides, western Texas to southern California. T- 
Specim.: | of 2. USA: New Mexico: Dona Ana Co., 
1881, GR. Vasey s.n. (IT: BAA-1061, US-822049, 
US-909912, US-1761631]. 2 of 2. USA: New Mexico, 
Vasey s.n. [LT: US], LT designated by Koch & 
Sanchez, Phytologia 58: 379 (1985). 
= Eragrostis mexicana subsp. mexicana. 


Eragrostis nigricans (Kunth) Steud., Nomencl. Bot. (ed. 2) 
1: 563. 1840. Basionym: Poa nigricans Kunth. 

DST.: Arge, Boli, Chil, Ecua, Peru. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Brako, L. & J. L. Zarucchi (1993); 
Foster, R. C. (1958); Hitchcock, A. S. (1927); 
Jorgensen, P. M. & S. Ledn-Yanez (eds.) (1999); 
Macbride, J. F. (1936); Marticorena, C. & M. 
Quezada (1985); Renvoize, S. A. (1998); Tovar, O. 
(1993). 

SYN.: Eragrostis nigricans var. tristis (Jedwabn.) Pilg.; 
Eragrostis subatra Jedwabn., Eragrostis  tristis 
Jedwabn.; Megastachya nigricans (Kunth) Roem. & 
Schult.; Poa atrovirens Willd. ex Spreng.; Poa 
nigricans Kunth. 


Eragrostis nigricans (Kunth) Steud. var. nigricans. 
DST.: Boli, Chil, Ecua, Peru. 
LVL. ACCPTC.: I. 


Eragrostis nigricans var. punensis Nicora, Boissiera 54: 65, 
f. 16. 1998. T-Protol.: Argentina: Prov. Jujuy: Dpto. 
Tumbaya: subida de Purmamarca al Abra de Pives, 
3800 m, 24 Apr 1975, A4.L. Cabrera et al. 26382 (HT: 
SI]. 

DST.: Arge, Chil. 
LVL. ACCPTC.: 1. 


Eragrostis nigricans var. tristis (Jedwabn.) Pilg., Notizbl. 
Bot. Gart. Berlin-Dahlem 11: 778. 1933. Basionym: 
Eragrostis tristis Jedwabn. 

= Eragrostis nigricans (Kunth) Steud. 


Eragrostis nitida (Elliott) Chapm., Fl. South. U.S. 564. 1860. 
Basionym: Poa nitida Elliott. 
NOTE: hom. illeg. 
E. NAME: Eragrostis nitida Link. 
= Eragrostis elliottii S. Watson. 


Eragrostis nuttalliana Steud., Nomencl. Bot. (ed. 2) 1: 563. 
1840. 
NOTE: nom. nov., nom. illeg. superfl. 
REPLACED NAME: Poa tenella Nutt. 
BLOCKING NAME: Poa tenella L. 
CORRECT NAME: Eragrostis brizoides Schult. 
= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis obtusiflora (E. Fourn.) Scribn., Bull. Div. 
Agrostol., U.S.D.A. 8: 10, t. 5. 1897. Basionym: 
Brizopyrum obtusiflorum E. Fourn. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 


Catalogue of New World Grasses: II. 


101 


ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Hitchcock, A. S. (1951); McVaugh, R. (1983). 

SYN.: Brizopyrum obtusiflorum E. Fourn. 


Eragrostis olmedoi (Kunth) Steud., Nomencl. Bot. (ed. 2) 1: 
564. 1840. Basionym: Poa olmedoi Kunth. 
= Eragrostis pastoensis (Kunth) Trin. 


Eragrostis orcuttiana Vasey, Contr. U.S. Natl. Herb. 1(8): 
269. 1893. T-Protol.: USA: California: San Diego, 
1885, CR. Orcutt 1313 [HT: US]. T-Specim.: 1 of 1. 
USA: California: Chollus Valley, San Diego, Aug 
1885, Orcutt 13/3 [HT: US-1761633; IT: MO- 
2646277]. 

= Eragrostis mexicana subsp. virescens (J. Presl) S.D. 
Koch & Sanchez Vega. 


Eragrostis oreophila L.H. Harv., Bull. Torrey Bot. Club 
81(5): 407. 1954. T-Protol.: Mexico: Hidalgo: in 
declivitate saxosa ad altitudinem 1500 m, prope Jacala, 
29 Jun 1939, V.H. Chase 7223 1/2 [HT: US-1763115; 
IT: GH, TEX]. T-Specim.: | of 1. Mexico: Hidalgo: 
Mun. Jacala: mountain ravine, alt. 4500 ft, 29 Jun 
1939, V.H. Chase 7223 1/2 [HT: US-1763115; IT: GH, 
MO-1179378, TEX]. 

= Eragrostis intermedia var. oreophila (L.H. Harv.) 
Witherspoon. 


Eragrostis orthoclada Hack., Bull. Herb. Boissier, ser. 2, 
4(3): 281. 1904. T-Protol.: Paraguay: in arenosis salsis 
in regione cursus superioris fluminis Apa., Jan, 
Hassler s.n. T-Specim.: | of 2. Paraguay: in arenosis 
salsis in regione cursus superiores fluminis Apa, Jan, 
Hassler 8347 [ST: BAA-1063 (fragm.)]. 2 of 2. 
Paraguay: in arenosis salsis in regione cursus 
superioris fluminis, Apa, Jan, Hassler 8460 [ST: BAA- 
1066 (fragm.), K, P]. 

DST.: Arge, Boli, Braz, Para. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Boechat, S. C. & P. M. Peterson 
(1995); Burkart, A. (1969); Killeen, T. J. (1990); 
Renvoize, S. A. (1998). 

SYN.: Eragrostis longipila Hack.; Eragrostis villamontana 
Jedwabn. 


Eragrostis ovina Hochst. ex A. Rich., Tent. Fl. Abyss. 2: 428. 
1851. T-Protol.: Abyssinia, Schimper, pl. Schimp. 
Abyss., sect. Ill, 1831] [K]. T-Specim.: 1 of 1. In 
planitie montana Schire, 10 Oct. 1840, Schimper 1831 
[OM: K, LE]. 

NOTE: nom, inval., as syn. of Poa ovina A. Rich. 
= Eragrostis gangetica (Roxb.) Steud. 


Eragrostis oxylepis (Torr.) Torr., Pacif. Railr. Rep. 4: 156. 
1857. Basionym: Poa oxylepis Torr. 

= Eragrostis secundiflora subsp. oxylepis (Torr.) S.D. 
Koch. 


Eragrostis oxylepis var. beyrichii (J.G. Sm.) Shinners, Field 
& Lab. 20(1): 34. 1952. Basionym: Eragrostis 
beyrichii J.G. Sm. 

= Eragrostis secundiflora subsp. oxylepis (Torr.) S.D. 
Koch. 


Eragrostis pallida Vasey, Contr. U.S. Natl. Herb. 1(8): 285. 
1893. T-Protol.: Mexico: Colima, collected in a ditch, 
9 Jan-6 Feb 1981, Edward Palmer 1268 (HT: US]. T- 


Specim.: | of 1. Mexico: Colima, Colima, 9 Jan-6 Feb 
1891, E. Palmer 1268 [HT: US-1749579; IT: NY- 
277651, NY-277652; US-911251]. 

= Eragrostis japonica (Thunb.) Trin. 


Eragrostis palmeri S. Watson, Proc. Amer. Acad. Arts 18: 
182. 1883. T-Protol.: Mexico: Coahuila: Juarez, on the 
Sabinas River, 1879-1880, Edward Palmer 1368. T- 
Specim.: | of 1. Mexico: Coahuila, Feb 1880-Oct 
1880, E. Palmer 1368 (JT: US-1761635, US-821984]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Correll, D. S. 
& M. C. Johnston (1970); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J.  Valdes-Reyna (2000); 
Hitchcock, A. S. (1951). 

SYN.: Eragrostis caudata E. Fourn. 


Eragrostis panamensis J. Presl, Relig. Haenk. 1(4-5): 277. 
1830. T-Protol.: Panama: Hab. in Panama, Haenke s.n. 
T-Specim.: 1 of 1. Panama, 7: Haenke s.n. [IT: US- 
2850745 (fragm.)]. 

= Eragrostis maypurensis (Kunth) Steud. 


Eragrostis paniciformis (A. Braun) Steud., Syn. Pl. Glumac. 
1: 268. 1854. Basionym: Poa paniciformis A. Braun. 

DST.: Braz. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Boechat, S.C. & 
Wagner (2000); Boechat, S. C. & 
(1995); Phillips, S. (1995). 

SYN.: Poa paniciformis A. Braun, 


Longhi- 


H. 
P. Peterson 


M. 
M. 


Eragrostis panicoides (J. Presl) Steud., Syn. Pl. Glumac. 1: 
278. 1854. Basionym: Megastachya panicoides J. 
Presl. 

= Leptochloa panicoides (J. Presl) Hitche. 


Eragrostis pappiana (Chiov.) Chiov., Annuario Reale Ist. 
Bot. Roma 8(3): 371. 1908. Basionym: Eragrostis 
multiflora var. pappiana Chiov. 

= Eragrostis minor Host. 


Eragrostis pappiana var. insularis A. Terrace. ex Chiov., 
Annuario Reale Ist. Bot. Roma none given. none 
given. T-Specim.: | of 2. Hamasen: Ghinda, Feb 
1907,? 770 [ST]. 2 of 2. Dembesan: Braucaga, 29 May 
1907,? 770 [ST]. 

NOTE: in Pirotta accd. to Chase & Niles, Index Grass Sp. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis pappii Gand., Bull. Soc. Bot. France 66(7): 299- 
300. 1919 [1920]. T-Protol.: Habitat Africa orient.: 
Erythraea ad Beni Amer., Pappi 5964. T-Specim.: | of 
1. Ethiopia: Eritrea, no date, 4. Pappi 5964 [HT: LY; 
IT: US-1445661]. 

NOTE: as Eragrostis "Pappii". 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis parviglumis Hochst. ex Steud., Syn. Pl. Glumac. 
1: 267. 1854. T-Protol.: Mont. Nilagiri, in Hohnkr. 
Hrbr. Ind. 936. T-Specim.: 1 of 1. India orientalis: 
Tamil Nandu, coimbatore, in montibus Nilagiri, 1851, 
Metz 936 [IT: L, LE, LE, US-1127146], 
Ed.R.F.Hohenaker.1851 [Herb.]. 
= Eragrostis tenuifolia (A. Rich.) Hochst. ex Steud. 


Eragrostis parvula Steud., Syn, P!. Glumac. |: 277. 1854. T- 
Protol.: Mexico. T-Specim.: 1 of 2. Mexico, Parry 172 


102 


Catalogue of New World Grasses: II. 


[LT: CN], LT designated (as holotype) by Koch, Ill. 


Biol. Monogr. 48: 44 (1974). 2 of 2. Mexico: M. 
Parreyss, 1845, Parry 127 [IT: US-79704 (fragm.)]. 


NOTE: Accd. to Chase & Niles, Index Grass Sp.,£. purshii 


var. parvula, based on this, but do not list they var. in 
their Index. 


CORRECT NAME: Eragrostis purshii var. miserrima E. 


Fourn. 


= Eragrostis pectinacea var. miserrima (E. Fourn.) 


Reeder. 


Eragrostis pastoensis (Kunth) Trin., Mem. Acad. Imp. Sci. 


DST. 
LVL. 


Saint-Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. 


Nat. 4,2(1): 71. 1836. Basionym: Poa pastoensis 


Kunth. 
: Arge, Boli, Braz, Colo, Ecua, Peru, Vene. 
ACCPTC:: 1. 


ALSO ACCPTD. BY: Boechat, S. C. & P. M. Peterson 


SYN.: 


(1995); Brako, L. & J. L. Zarucchi (1993); 
Hitchcock, A. S. (1927); Jorgensen, P. M. & S. Leon- 
Yanez (eds.) (1999); Macbride, J. F. (1936); 
Renvoize, S. A. (1998); Tovar, O. (1993). 

Eragrostis  boliviensis  Jedwabn.,; — Eragrostis 
buchtienii WHack.; Eragrostis  contracta  Pilg.; 
Eragrostis lehmannii Pilg., Eragrostis montufari 
(Kunth) Steud.; Eragrostis olmedoi (Kunth) Steud.; 
Eragrostis setifolia (Benth.) Steud.; Eragrostis 
virescens var. trachyphylla Hack.; Eragrostis 
virescens var. trichophylla (Benth.) Hack. ex Stuck.; 
Megastachya montufarii (Kunth) Roem. & Schult.; 
Megastachya olmedoi (Kunth) Roem. & Schult.; 
Megastachya pastoensis (Kunth) Roem. & Schult; 
Poa montufari Kunth; Poa olmedoi Kunth; Poa 
pastoensis Kunth; Poa setifolia Benth. 


Eragrostis patula (Kunth) Steud., Nomencl. Bot. (ed. 2) 1: 


DST. 


564. 1840. Basionym: Poa patula Kunth. 
: Ecua. 


ALSO ACCPTD. BY: Brako, L. & J. L. Zarucchi (1993); 


SYN 


Hitchcock, A. S. (1927); Macbride, J. F. (1936); 
Renvoize, S. A. (1998). 


| Megastachya patula (Kunth) Roem. & Schult.; Poa 


patula Kunth. 


= Status not yet determined. 


Eragrostis pauciflora Trin. ex E. Fourn. ex Hemsl., Biol. 


Cent.-Amer., Bot. 3: 575. 1885, 


NOTE: nom. nud. 
CORRECT NAME: Eragrostis pauciflora Trin. ex E. 


Fourn. 


= Eragrostis mexicana (Hornem.) Link. 


Eragrostis pauciflora Trin. ex E. Fourn., Mexic. Pl. 2: 117. 


1886. T-Specim.: | of 4. Mexico: Hacienda del Sefior 
Posado pr. Orizaba, Schaffner pl. ed Hohen. 51 [ST]. 2 
of 4. Mexico: Veracruz: Hacienda de Sefior Posado pr. 
Orizaba, Thomas s.n. [ST: herb. Buchinger]. 3 of 4. 
Mexico: San Luis de Potosi, Virlet /383 [ST]. 4 of 4. 
Mexico: circa Mejico, Berlandier 843 [ST]. 


NOTE: cites "£. pauciflora Trin. in sched., E. nigricans as 


misapl. by Griseb. in sched., non. Kunth. 


= Eragrostis mexicana (Hornem.) Link. 


Eragrostis pectimacea (Michx.) Nees, Fl. Afr. Austral. Il. 


NOTE: Nees 


406. 1841. Basionym: Poa pectinacea Michx. 
clearly accepts Eragrostis pectinaceae 
"Michx.", provides a diagnosis that corresponds to 


DST. 


LVL. 


Poa pectinaceae Michx., and the combination is 
effectively made by indirect citation. 

: Arge, Braz, Cana, Cari, CoRi, Ecua, ElSa, FrGu, 
Guat, Guya, Hond, Mexi, Pana, Para, Peru, Suri, 
USA, Urug, Vene. 

ACCPTC.: 1. 


ALSO ACCPTD. BY: Beetle, A. A. (1977); Berendsohn, 


SYN.: 


W.G. & A. E. Araniva de Gonzalez (1991); Boechat, 
S.C. & H. M. Longhi-Wagner (2000); Brako, L. & J. 
L. Zarucchi (1993); Cabrera, A. L. (1970); Correll, 
D.S. & M. C. Johnston (1970); Cronquist, A., A. H. 
Holmgren, N. H. Holmgren & J. L. Reveal (1977); 
Davidse, G. (1994); Davidse, G., M. Sousa S. & A. 
QO. Chater (editores generales) (1994); Espejo Serna, 
A., A. R. L6pez-Ferrari, and J. Valdes-Reyna (2000); 
Fernald, M. (1950); Filgueiras, T. S. (1991); Gleason, 
H. A. & A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Godfrey, R. K. & J. W. Wooten 
(1979); Gould, F. W. & R. Moran (1981); Great 
Plains Flora Association (1986); Hitchcock, A. S. 
(1951); Hitchcock, C. L., A. Cronquist, M. Ownbey 
& J. W. Thompson (1969); Jorgensen, P. M. & S. 
Leén-Yanez (eds.) (1999); Kerguelen, M. (1993); 
Levin, G. A. & R. Moran (1989); McVaugh, R. 
(1983); Munz, P. A. & D. D. Keck (1959); Munz, P. 
A. (1974); Pohl, R. W. (1980); Renner, S. S., H. 
Balslev & L. B. Holm-Nielsen (1990); Reyes-Garcia, 
A. & M. Sousa S. (1997); Scoggan, H. J. (1978 
[1979]); Small, J. K. (1933); Voss, E. G. (1972); 
Wunderlin, R. P. (1998). 

Eragrostis brizoides Schult.; Eragrostis caroliniana 
(Biehler) Scribn.; Eragrostis cognata  Steud.; 
Eragrostis delicatula Trin.; Eragrostis diffusa var. 
diffusa; Eragrostis nuttalliana Steud., Eragrostis 
pectinacea (Michx.) Steud.; Eragrostis 
pennsylvanica Scheele; Eragrostis pilosa var. 
caroliniana (Biehler) Farw.; Eragrostis purshii 
Schrad.; Eragrostis purshii Schrad. ex A. Gray; 
Eragrostis purshii var. delicatula Munro ex Scribn.; 
Eragrostis purshii var. pauciflora  E. Fourn.; 
Eragrostis spectabilis var. sparsihirsuta  Farw.; 
Eragrostis tracyi Hitche.; Eragrostis unionis Steud.; 
Poa diandra hort. ex Schrad.; Poa eragrostis Elliot; 
Poa nuttallii Kunth; Poa pectinacea Michx.,; Poa 
tenella Nutt. 


Eragrostis pectinacea (Michx.) Steud., Syn. Pl. Glumac. 1: 


272. 1854. Basionym: Poa pectinacea Michx. 


NOTE: isonym. 
E. NAME: Eragrostis pectinacea (Michx.) Nees. 
= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis pectinacea var. miserrima (E. Fourn,) Reeder, 


LVL 


Phytologia 60(2): 154. 1986. Basionym: Eragrostis 


purshii var. miserrima E. Fourn. 
DST. 


: Arge, Braz, Cana, Cari, CoRi, ElSa, Guat, Hond, 
Mexi, Nica, Pana, Peru, USA, Urug. 


. ACCPTC.: 1; introduced. 


ALSO ACCPTD. BY: Brako, L. & J. L. Zarucchi (1993); 


SYN.: 


Davidse, G. (1994); Davidse, G., M. Sousa S. & A. 
O. Chater (editores generales) (1994); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Hickman, J. C. (1993); Wunderlin, R. P. (1998). 

Eragrostis arida Hitche.; Eragrostis diffusa var. 
arida (Hitche.) Beetle; Eragrostis parvula Steud.; 
Eragrostis pilosa var. delicatula (Trin.) Hack.; 
Eragrostis purshii var. miserrima’ E. Fourn.; 


Catalogue of New World Grasses: II. 


103 


Eragrostis tephrosanthos Schult.; Poa tephrosanthos 
Spreng. ex Schult. 


Eragrostis pectinacea (Michx.) Nees var. pectinacea. 

DST.: Arge, Braz, Cana, Cari, CoRi, Ecua, ElSa, FrGu, 
Guat, Guya, Hond, Mexi, Nica, Pana, Para, Peru, 
Suri, USA, Urug, Vene. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Brako, L. & J. L. Zarucchi (1993); 
Davidse, G. (1994); Davidse, G., M. Sousa S. & A. 
O. Chater (editores generales) (1994); Hickman, J. C. 
(1993); Wunderlin, R. P. (1998). 

SYN.: Eragrostis diffusa Buckley; Eragrostis purshii var. 
diffusa (Buckley) Vasey. 


Eragrostis pectinacea var. refracta (Muhl. ex Elliott) 
Chapm., Fl. South. U.S. 564. 1860. Basionym: Poa 
refracta Muhl. ex Elliott. 

= Eragrostis refracta (Muhl. ex Elliott) Scribn. 


Eragrostis pectinacea var. spectabilis (Pursh) A. Gray, 
Manual (ed. 2) 565. 1856. Basionym: Poa spectabilis 
Pursh. 

= Eragrostis spectabilis (Pursh) Steud. 


Eragrostis pectinacea var. tracyi (Hitchc.) P.M. Peterson, 
Sida 17(1): 106. 1996. Basionym: Eragrostis tracyi 
Hitche. 

DST.: USA. 
LVL. ACCPTC:: 1. 
SYN.: Eragrostis tracyi Hitche. 


Eragrostis pennsylvanica Scheele, Flora 27: 58. 1844. T- 
Protol.: Amer. Bor. 
NOTE: IK gives p. 56. 
= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis peregrina Wiegand, Rhodora 19: 95. 1917. 
REPLACED NAME: Eragrostis pilosa var. condensata 
Hack. 
BLOCKING NAME: Eragrostis condensata (J. Presl) 
Steud. 
= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis perennis Doll, Fl. Bras. 2(3): 144. 1878. T- 
Protol.: Brasilia: habitat inter Rio Grande et Tejuco 
Prov. Minarum, Burchell 5689. T-Specim.: 1 of 1. 
Brazil: Minas Gerais: Inter Rio Grande et Tejuco, WJ. 
Burchell 5689 [HT: ?; IT: K, US-79708 (fragm.)]. 

DST.: Arge, Boli, Braz, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Doll, J. C. (1878); Filgueiras, T. S. 
(1999); Killeen, T. J. (1990); Longhi-Wagner, H. M. 
(1990); Renvoize, S. A. (1988); Renvoize, S. A. 
(1998): Rosengurtt, B., B. R. Arrillaga de Maffei & 
P. Izaguirre de Artucio (1970); Smith, L. B., D. C. 
Wasshausen & R. M. Klein (1981). 


Eragrostis perplexa L.H. Harv., Bull. Torrey Bot. Club 
81(5): 409. 1954. T-Protol.: USA: South Dakota: 
Mellette Co.: in alkaline soil, 30 Aug 1935, WL. 
Tolstead s.n. (HT: US-1645027]. T-Specim.: 1 of 1. 
USA: South Dakota: Mellette Co., 30 Aug 1935,W.L. 
Tolstead s.n. (HT: US-1645027; IT: US-1649186]. 

= Eragrostis pilosa var. perplexa (L.H. Harv.) S.D. Koch. 


Eragrostis peruviana (Jacq.) Trin., Mem. Acad. Imp. Sci. 
St.-Petersbourg, Ser. 6, Sci. Math. 1(4): 396. 1830. 
Basionym: Poa peruviana Jacq. 

DST.: Chil, Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Brako, L. & J. L. Zarucchi (1993): 
Hitchcock, A. S. (1927); Macbride, J. F. (1936); 
Marticorena, C. & M. Quezada (1985): Nicora, E. G. 
(1994): Tovar, O. (1993). 

Eragrostis deserticola Phil.,; Eragrostis peruviana 
var. brachythyrsa Pilg.; Koeleria multiflora Regel & 
Herter; Poa peruviana Jacq. 


SYN:: 


Eragrostis peruviana var. brachythyrsa Pilg., Bot. Jahrb. 
Syst. 37: 375. 1906. T-Specim.: | of 2. Peru: 
Mollendo, in formatione "loma", 200-600 m, Oct 
1902, A. Weberbauer 1462 [LT: B; ILT: US-2851259 
(fragm.)], LT designated (as holotypus) by Nicora, 
Gayana Bot. 51(1): 2 (1994). 2 of 2. Peru: Tacna, 
gramen principale a collibus apertis, arenosis, 800- 
1000 m, 1891, Woitschach s.n. [ST]. 

= Eragrostis peruviana (Jacq.) Trin. 


Eragrostis peruviana var. macranthera Skottsb., Kongl. 
Gotheborgska Wetensk. Samhallets Handl., Wetensk. 
Afd. 5(6): 81, f. 43, 44 b-i. 1937. T-Protol.: Chile: II 
Region: San Ambrosio, Johow form 3, 5-6 Oct. 
1896[HT: UPS]. 
CORRECT NAME: Eragrostis kuschelti Skottsb. 
= Eragrostis kuschelii Skottsb. 


Eragrostis petrensis Renvoize & Longhi-Wagner, Kew 
Bull. 41(1): 71. 1986. T-Protol.: Brazil: Bahia: Rio de 
Contas, 13 Dec 1984, Longhi-Wagner et al. CFCR 
6752 [HT: SPF]. 

DST.: Braz. 

LVL. ACCPTC:.: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Filgueiras, T. S. (1999); Renvoize, S. 
A. & H. M. Longhi-Wagner (1986). 


Eragrostis pilgeri Fedde, Just's Bot. Jahresber. 3: 18, t. 34. 
1908. 

REPLACED NAME: Eragrostis andicola Pilg. 

BLOCKING NAME: Eragrostis andicola R_E. Fr. 

DST.: Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Tovar, O. (1993). 

SYN.: Eragrostis andicola fo. humilior Pilg.; Eragrostis 
andicola Pilg.; Eragrostis andicola var. robustior 
Pilg.; Eragrostis carazensis Pilg.; Eragrostis 
pilgeriana Hitche. 


Eragrostis pilgeriana Hitche., Contr. U.S. Natl. Herb. 24(8): 
342. 1927. 

NOTE: nom. nov., nom. illeg. superfl. and hom. 
E. NAME: Eragrostis pilgeriana Dinter ex Pilg. 
REPLACED NAME: Eragrostis andicola Pilg. 
BLOCKING NAME: Eragrostis andicola R.E. Fr. 
CORRECT NAME: Eragrostis pilgeri Fedde. 
= Eragrostis pilgeri Fedde. 


Eragrostis pilifera Scheele, Linnaea 22(3): 344. 1849. T- 
Protol.. USA: Texas: auf trockner, hochliegender 
Prairie bei Neubraunfels, Lindheimer s.n. T-Specim.: | 
of 1. USA: Texas: Neubraunfels, 1847, F. Lindheimer 
736 [LT: US-909190; ILT: US-1871150], LT 
designated by ?. 


104 


Catalogue of New World Grasses: II. 


= Eragrostis trichodes (Nutt.) Alph. Wood. 


Eragrostis pilosa (L.) P. Beauv., Ess. Agrostogr. 71, 162, 
175. 1812. Basionym: Poa pilosa L. 
DST.: Arge, Boli, Braz, Cana, Cari, Chil, CoRi, Ecua, ElSa, 
FrGu, Guya, Hond, Mexi, Nica, Para, Peru, USA, 
Urug, Vene. 
LVL. ACCPTC.: 1; introduced. 
ALSO ACCPTD. BY: Adams, C. D. (1972); Allem, A. C. 
& J. F. M. Valls (1987); Beetle, A. A. (1977); 
Berendsohn, W. G. & A. E. Araniva de Gonzalez 
(1991); Boechat, S. C. & H. M. Longhi-Wagner 
(2000); Bor, N. L. (1960); Brako, L. & J. L. Zarucchi 
(1993); Burkart, A. (1969); Cabrera, A. L. (1970); 
Correll, D. S. & M. C. Johnston (1970); Davidse, G. 
(1994); Davidse, G., M. Sousa S. & A. QO. Chater 
(editores generales) (1994); Doll, J. C. (1878); Edgar, 
E., H. E. Connor & J. E. Shand (1991); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Fernald, M. (1950); Filgueiras, T. S. (1991); Foster, 
R. C. (1958); Gibbs Russell, G. E, W. G. Welman, E. 
Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Gould, F. W 
(1979); Gould, F. W. & R. Moran (1981); Great 
Plains Flora Association (1986); Hitchcock, A. S. 
(1927); Hitchcock, A. S. (1936); Hitchcock, A. S. 
(1951); Hitchcock, C. L., A. Cronquist, M. Ownbey 
& J. W. Thompson (1969); Jorgensen, P. M. & S. 
Leon-Yanez (eds.) (1999): Judziewicz, E. J. (1990); 
Kerguelen, M. (1993); Killeen, T. J. (1990); 
Lawesson, J. E., H. Adsersen & P. Bentley (1987); 
Lindeman, J. C. & A. L. Stoffers (1963); Liogier, H. 
& L. Martorell (1982); McVaugh, R. (1983); Munz, 
P. A. & D. D. Keck (1959); Munz, P. A. (1974); 
Nicora, E. G., M. E. De Paula, A. M. Faggi, M. del 
C. Mariano, A. M. Miante A., L. R. Parodi, C. A. 
Petetin, F. A. Roig & Z. Rugolo de Agrasar (1978); 
Phillips, S. (1995); Pohl, R. W. (1980); Porter, D. M. 
(1983); Radford, A. E., H. E. Ahles & C. R. Bell 
(1968); Renvoize, S. A. (1984); Renvoize, S. A. 
(1988); Renvoize, S. A. (1998); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 
Schwegman, John E. (1991); Shreve, F. & I. L. 
Wiggins (1964); Small, J. K. (1933); Smith, L. B., D. 
C. Wasshausen & R. M. Klein (1981); Tovar, O. 
(1993); Voss, E. G. (1972); Wiggins, I. L. & D. M. 
Porter (1971); Wunderlin, R. P. (1998). 
Eragrostis damiensiana (Bonnet) Thell.; Eragrostis 
damiensiana var. condensata’ (Hack.)  Thell.; 
Eragrostis damiensiana var. laxior Thell.; Eragrostis 
filiformis Link; Eragrostis — gracilis Schrad.; 
Eragrostis linkii (Kunth) Steud.; Eragrostis 
multicaulis Steud.; Eragrostis peregrina Wiegand; 
Eragrostis pilosa fo. multicaulis (Steud.) 1.C. Chung; 
Eragrostis pilosa subsp. damiensiana (Bonnet) 
Thell.; Eragrostis pilosa var. condensata Hack.; 
Eragrostis pilosa var. condensata Hack.; Eragrostis 
pilosa var. damiensiana Bonnet; Eragrostis 
verticillata (Cav.) P. Beauv.; Eragrostis verticillata 
(Cav.) Roem. & Schult.; Poa eragrostis Walter; Poa 
linkii Kunth; Poa pilosa L.; Poa tenella Willd.? (L. 
misappl. by) Pursh; Poa verticillata Cav. 


SYN.: 


Eragrostis pilosa subsp. abyssinica (Jacq.) Asch. & Graebn., 
Syn. Mitteleur. Fl. 2: 374. 1900. Basionym: Poa 
abyssinica Jacq. 

= Eragrostis tef (Zucc.) Trotter. 


Eragrostis pilosa var. bahiensis (Schrad. ex Schult.) Kuntze, 
Revis. Gen. Pl. 3(2,2): 353. 1898. Basionym: 
Eragrostis bahiensis Schrad. ex Schult. 

= Eragrostis bahiensis Schrad. ex Schult. 


Eragrostis pilosa var. capillaris (L.) Kuntze, Revis. Gen. PI. 
3: 353. 1898. Basionym: Poa capillaris L. 
= Eragrostis capillaris (L.) Nees. 


Eragrostis pilosa var. caroliniana (Biehler) Farw.; Rep. 
Michigan Acad. Sci. 17: 182. 1916. Basionym: Poa 
caroliniana Biehler. 

= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis pilosa var. condensata Hack., Allg. Bot. Z. Syst. 

7: 13. 1901. T-Protol.: Karlsruhe, Kneucker Gram. 
Exs, 115, T-Specim.: 1 of |. Germany: Karlsruhe: 
Baden-Wurttemberg, elev. 117 m, 17 Jul 1900, 1A. 
Exs Kneucker 115 (IT: BAA-1072, SI, US-81897, US- 
79709 (fragm.), W, WAG]. 

CORRECT NAME: Eragrostis peregrina Wiegand. 

= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis pilosa var. condensata Hack., Vierteljahrsschr. 
Naturf. Ges. Zurich 52: 439. 1908. 
NOTE: Thellung provides quadrinomial taxonomy here, as 
E. pilosa subsp. damiensiana var. condensata. 
E. NAME: Eragrostis pilosa var. condensata Hack. 
= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis pilosa subsp. damiensiana (Bonnet) Thell., 
Vierteljahrsschr. Naturf. Ges. Zurich 52: 438. 1908. 
Basionym: Eragrostis pilosa var. damiensiana Bonnet. 

= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis pilosa var. damiensiana Bonnet, Naturaliste 3: 
412. 1881. T-Protol.: France: Paris: dans la cour du 
ministere de la Guerre, Af, Damiens s.n. 

NOTE: No. 52. 412. 1881., Accd. to Chase & Niles, Index 
Grass Sp. 
= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis pilosa var. delicatula (Trin.) Hack., Anales Mus. 
Nac. Buenos Aires 11: 133. 1904. Basionym: 
Eragrostis delicatula Trin. 

= Eragrostis pectinacea var. miserrima (E. 
Reeder. 


Fourn.) 


Eragrostis pilosa var. lugens (Nees) Griseb., Abh. Konig]. 
Ges. Wiss. Gottingen 24: 290. 1879. Basionym: 
Eragrostis lugens Nees. 

= Eragrostis lugens Nees. 


Eragrostis pilosa var. lugens Kuntze, Revis. Gen. Pl. 3(3): 
352, 353. 1898. T-Protol.: Argentina: Patagonia, 
Moreno & Tonini s.n. 

NOTE: hom. illeg. 
E. NAME: Eragrostis pilosa var. lugens (Nees) Griseb. 
= Eragrostis lugens Nees. 


Eragrostis pilosa var. minor (Host) Kuntze, Revis. Gen. PI. 
3(2): 353. 1898. Basionym: Eragrostis minor Host. 
= Eragrostis minor Host. 


Catalogue of New World Grasses: II. 


105 


Eragrostis pilosa fo. multicaulis (Steud.) I.C. Chung, J. 
Wash. Acad. Sci. 45(7): 215. 1955. Basionym: 
Eragrostis multicaulis Steud. 

= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis pilosa var. perplexa (L.H. Harv.) S.D. Koch, 
Illinois Biol. Monogr. 48: 28. 1974. Basionym: 
Eragrostis perplexa L.H. Harv. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Great Plains Flora Association 
(1986). 

SYN.: Eragrostis perplexa L.H. Harv. 


Eragrostis pilosa (L.) P. Beauv. var. pilosa. 
DST.: USA. 
LVL. ACCPTC.: 1; introduced. 
ALSO ACCPTD. BY: Great Plains Flora Association 
(1986); Hickman, J. C. (1993). 


Eragrostis pilosa var. tef (Zucc.) Fiori, Nuov. Fl. Italia 1: 
123. 1923. Basionym: Poa tef Zucc. 
= Eragrostis tef (Zucc.) Trotter. 


Eragrostis pilosa var. versicolor Kuntze, Revis. Gen. PI. 
3(2): 353. 1898. T-Protol.: Bolivia: Rio Tapacari, 
Kuntze s.n. 

= Status not yet determined. 


Eragrostis pilosissima Link, Hort. Berol. 1: 189. 1827. T- 
Protol.: Germany: Berlin. 
= Eragrostis articulata (Schrank) Nees. 


Eragrostis plana Nees, FI. Afr. Austral. Ill, 390. 1841. T- 
Protol.: Ad flumen prope Kachu et Zandplaat alt. 1500' 
et inter Geckau et Basche alt. 1500-2000', Drége s.n. 
T-Specim.: | of 3. South Africa: Cap. b. sp., Drége 
[ST: LE (hrbr. Fischer)]. 2 of 3. South Africa: ad 
flumen prope Kachu et Zandplast, Drége s.n. [ST]. 3 
of 3. South Africa: inter Geckau et Basche, Drége s.n. 
[ST]. 

DST.: Braz, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Boechat, §. C. & H. M. Longhi- 
Wagner (2000); Bor, N. L. (1960); Edgar, E., H. E. 
Connor & J. E. Shand (1991); Filgueiras, T. S. 
(1991); Gibbs Russell, G. E, W. G. Welman, E. 
Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987); Koch, S. D. (1978). 


Eragrostis plumbea Scribn. ex Beal, Grass. N. Amer. 2: 
484. 1896. T-Protol.: Mexico: Jalisco: Guadalajara. T- 
Specim.: 1 of 2. Mexico: Jalisco, 1 Oct 1889, C.G. 
Pringle 2311 [LT: MSC?; ILT: TEX, US-822001, US- 
1761636], LT designated by McVaugh, Fl. Novo-Gal. 
14: 484 (1983), without specifically citing the 
lectotype, just one of the isolectotypes. 2 of 2. Mexico: 
Jalisco: Guadalajara, Jul-Oct 1880, E. Palmer 240 [ST: 
MSC, MO-820709]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
MceVaugh, R. (1983). 


Eragrostis plumosa (Retz.) Link, Hort. Berol. 1: 192. 1827. 
Basionym: Poa plumosa Retz. 
= Eragrostis amabilis (L.) Wight & Arn. ex Nees. 


Eragrostis plurinodis Swallen ex Luces, J. Wash. Acad. Sci. 
32(6): 158, f. 2. 1942. T-Protol.: Venezuela: Apure: in 
fields around Cunaviche: 13 Feb 1941,C.E. Chardon 
249 (HT: VEN]. T-Specim.: | of 1. Venezuela: Apure: 
Cerca de Curnaviche, 13 Feb 1941,C.E. Chardon 249 
{HT: VEN; IT: US-1817505 (fragm. ex VEN)]. 

DST.: Braz, Guya, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Judziewicz, E. J. (1990). 


‘Eragrostis poaeoides P. Beauv. ex Roem. & Schult., Syst. 


Veg. 2: 574. 1817. 
NOTE: nom. illeg. superfl. 
REPLACED NAME: Eragrostis poaeoides P. Beauv. 
CORRECT NAME: Eragrostis minor Host. 
= Eragrostis minor Host. 


Eragrostis poaeoides var. barrelieri (Daveau) Fiori, FI. Italia 
1: 82. 1908. Basionym: Eragrostis barrelieri Daveau. 
= Eragrostis barrelieri Daveau. 


Eragrostis poaeoides vat. laxiflora Dill, Fl. Bras. 2(3): 149. 
1878. T-Protol.: Brazil: loco accuratius non indicato, 
Widgren, Freyreiss & Forstrom  [HT: — hrbr. 
Holmiensis]. 

= Eragrostis minor Host. 


Eragrostis poaeoides var. megastachya (Koeler) A. Gray, 
Manual (ed. 2) 563. 1856. Basionym: Poa 
me gastachya Koeler. 

= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis poiformis Link, Hort. Berol. 1: 188. 1827. 
NOTE: nom. nov., nom. illeg. superfl. 
REPLACED NAME: Poa eragrostis L. 
BLOCKING NAME: Eragrostis eragrostis (L.) P. Beauv. 
CORRECT NAME: Eragrostis minor Host. 
= Eragrostis minor Host. 


Eragrostis polytricha Nees, Fl. Bras. Enum. PI. 2: 507-508. 
1829. T-Protol.: Habitat in Brasili australi, Se//ow s.n. 
[HT: B]. T-Specim.: | of 1. Brazil, F. F. Sellow s.n. 
[1882] |HT: B; IT: BAA-2668 (ex B), US-77386 
(fragm.)]. 
DST.: Arge, Beli, Boli, Braz, Chil, Colo, Guat, Guya, Hond, 
Mexi, Nica, Para, USA, Urug, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Balick, M. J.. M. H. Nee & D. E. 
Atha (2000); Boechat, S. C. & H. M. Longhi-Wagner 
(2000); Burkart, A. (1969); Cabrera, A. L. (1970); 
Davidse, G. (1994); Davidse, G., M. Sousa S. & A. 
O. Chater (editores generales) (1994); Dll, J. C. 
(1878); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Filgueiras, T. S. (1991); 
Filgueiras, T. S. (1999); Foster, R. C. (1958); 
Hitchcock, A. S. (1927); Judziewicz, E. J. (1990); 
Killeen, T. J. (1990); Longhi-Wagner, H. M. (1990); 
Marticorena, C. & M. Quezada (1985); Renvoize, S. 
A. (1984); Renvoize, S. A. (1988); Renvoize, S. A. 
(1998); Rosengurtt, B., B. R. Arrillaga de Maffei & 
P. Izaguirre de Artucio (1970). 

Eragrostis floridana Hitche.; Eragrostis fragilis 
Swallen; Eragrostis lugens var. villosa  DOll; 
Eragrostis polytricha var. glabrior Doll; Eragrostis 
polytricha var. hirsutior Doll, Eragrostis purpusii 
Jedwabn.; Eragrostis trichocolea var. floridana 


SYN.: 


106 


Catalogue of New World Grasses: II. 


(Hitche.) Witherspoon; Poa conceptionis Steud.; Poa 
polytricha (Nees) Kunth. 


Eragrostis polytricha var. glabrior Doll, Fl. Bras. 2(3): 140. 
1878. T-Protol.: Eragrostis polytricha var. alpha, Nees 
ab Esenb. Agr. bras. T-Specim.: 1 of 1. Brazil: in 
campis —- prope Villa Rica aliisque locis provinciae 
Minarum [IT: US]. 

= Eragrostis polytricha Nees. 


Eragrostis polytricha var. hirsutior D6ll, Fl. Bras. 2(3): 140. 
1878. T-Protol.: Eragrostis polytricha var. beta Nees 
ab Esenb. Agr. bras. T-Specim.: 1 of 1. Brazil: in 
campis limosis altis prope Villa Rica aliisque locis 
provinciae Minarum, Martius s.n. [IT: US]. 

= Eragrostis polytricha Nees. 


Eragrostis praetermissa L.H. Harv., Bull. Torrey Bot. Club 
81(5): 408. 1954. T-Protol.: Guatemala: Baja Verapaz: 
prope Santa Rosa, in rupibus, Jul 1887,von Turckheim 
1292 [HT: US-821939]. T-Specim.: | of 1. Guatemala: 
Baja Verapaz, Jul 1887, H. von Tuerckheim 1292 [HT: 
US-821939]. 

= Eragrostis hirta E. Fourn. 


Eragrostis pringlei Mattei, Contr. Biol. Veg. 4: 241. 1909. 
REPLACED NAME: Eragrostis pusilla Scribn. ex Beal. 
BLOCKING NAME: Eragrostis pusilla Hack. 

DST.: Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983). 

SYN.: Eragrostis pusilla Scribn. ex Beal; Eragrostis 
scribneriana Hitche. 


Eragrostis prolifera (Sw.) Steud., Syn. Pl. Glumac. 1: 278. 
1854. Basionym: Poa prolifera Sw. 
DST.: Beli, Braz, Cari, Colo, CoRi, Ecua, ElSa, Guat, 
Hond, Mexi, Nica, Pana, Peru, Vene. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Balick, M. J., M. H. Nee & D. E. 
Atha (2000); Beetle, A. A. (1977); Berendsohn, W. 
G. & A. E. Araniva de Gonzalez (1991); Boechat, S. 
C. & H. M. Longhi-Wagner (2000); Brako, L. & J. L. 
Zarucchi (1993); Breedlove, D. E. (1986); Catasus 
Guerra, L. (1997); Davidse, G. (1994); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994); Doll, J. C. (1878); Gould, F. W (1979); 
Hitchcock, A. S. (1936); Jorgensen, P. M. & S. Leon- 
Yanez (eds.) (1999); Lawesson, J. E., H. Adsersen & 
P. Bentley (1987); McVaugh, R. (1983); Nelson, C. 
(1979): Pohl, R. W. (1980); Renvoize, S. A. (1984); 
Sousa S., M. & E. F. Cabrera C. (1983); Wunderlin, 
R. P. (1998). 


SYN.: Eragrostis — diversiflora Vasey; — Eragrostis 
domingensis (Pers.) Steud.; Eragrostis excelsa 
Griseb.; Eragrostis gigantea Trin.; Eragrostis 


haenkei J. Presl; Eragrostis maxima E. Fourn.; 
Eragrostis salzmannii Steud.; Poa domingensis Pers.; 
Poa prolifera Sw. 


Eragrostis psammodes Trin., Mem. Acad. Imp. Sci. St.- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 400. 1830. T- 
Protol.: V. subsp. Brasilia (in ripa rivi Coxim, das 
Velhas). T-Specim.: | of 7. In sabulosis ripam rio das 
Velhas Barra ab Jiquitiba, Nov. 1824, Riedel s.n. [ST: 
K, LE]. 2 of 7. Paraguay: i ripa Taquary, Nov 1826 


{ST: LE]. 3 of 7. Brasilia, #. Se/low s.n. [ST: LE]. 4 of 
7. In ripa rivi Coxine, Nov 1826 [ST: LE]. 5 of 7. , 
Nov 1824, 846 [IT: LE]. 6 of 7. [Brasil]: in arenosis 
ripis rivi das Velhas, Nov 24, s.c. 846 [HT: LE-TRIN- 
2386.04], b="in ripa rivi Coxine. Cuyaba". 7 of 7. 
Brazil: in arenosis ripas rivi das Velhas, GH. von 
Langsdorff [error for Riedel] 846 \IT: NY, US-79712 
(fragm.)]. 
= Eragrostis bahiensis Schrad. ex Schult. 


Eragrostis psammodes var. microstachya (Link) DOll, FI. 
Bras. 2(3): 153. 1878. T-Protol.: Uruguay: prope 
Montevideo, Sello?. 

= Eragrostis bahiensis Schrad. ex Schult. 


Eragrostis purpurascens (Spreng.) Schult., Mant. 2: 317. 
1824. Basionym: Poa purpurascens Spreng. 

DST.: Braz, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Boechat, S.C. & H. M. Longhi- 
Wagner (2000); Doll, J. C. (1878); Gould, F. W 
(1979); Hitchcock, A. S. (1936); Longhi-Wagner, H. 
M. & I. I. Boldrini (1988); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970). 

SYN.: Eragrostis megapotamica (Spreng.) Schult.; Poa 
purpurascens Spreng. 


Eragrostis purpusii Jedwabn., Bot. Arch. 5(3-4): 201. 1924. 
T-Protol.: Mexico: Cerro de Gavilan, Purpus 4084. T- 
Specim.: | of 1. Mexico: Puebla: Cerro del Gavilan, 
C.A. Purpus 4084 [LT: UC; IT: US-77385 (fragm.)], 
LT designated by Witherspoon, Ann. Missouri Bot. 
Gard. 64:328(1977). 

NOTE: as Eragrostis "Purpusii". 
= Eragrostis polytricha Nees. 


Eragrostis purshii Schrad., Linnaea 12(4): 451. 1838. T- 
Protol.: China: Guangxi: Hepu, Wenli, in open 
wilderness, 400 m, 6 April 1956, Hepu Exped. 1648 
(HT: ?; IT: IBSC]. 

NOTE: nom. nud.; cites Poa diandra hort., non. R.Br., 

name later validated by A. Gray. 
= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis purshii Schrad, ex A, Gray, Manual 564, 1856. T- 
Protol.: Sand New Jersey to Virginia, and southwardy 
or sterile open ground. 

NOTE: cites E. purshii (Bernh.?) Schrad. [nom. nud.], Poa 
tenella sensu Pursh, P. caroliniana Spreng. [actually 
published first by Biehler], P. petinacea of authors, 
not of Michx. 

CORRECT NAME: Eragrostis 
Scribn. 

= Eragrostis pectinacea (Michx.) Nees. 


caroliniana  (Biehler) 


Eragrostis purshii var. delicatula Munro ex Scribn., Bull. 
Torrey Bot. Club 10(1): 30. 1883. T-Protol.: USA: 
Arizona: Santa Cruz Valley, near Tucson, Pringle s.n. 

NOTE: nom. nud. 
= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis purshii var. diffusa (Buckley) Vasey, Contr. U.S. 
Natl. Herb. 1(2): 59. 1890. Basionym: Eragrostis 
diffusa Buckley. 

= Eragrostis pectinacea var. pectinacea. 


Eragrostis purshii var. major E. Fourn., Mexic. Pl. 2: 116. 
1886. T-Specim.: | of 4. Mexico, Bourgeau 445 [ST]. 
= E. mexicana. 2 of 4. Mexico, Bourgeau 1031] [ST]. 3 


Catalogue of New World Grasses: II. 


107 


of 4. Nicaragua, P. Levy s.n. [ST]. 4 of 4. Mexico, /.L. 
Berlandier 794 |ST: US-2850915 (fragm.)]. 

NOTE: cites "E. poaeoides Griseb. in sched. part.". 

= Eragrostis mexicana (Hornem.) Link. 


Eragrostis purshii var. miserrima E. Fourn., Mexic. Pl. 2: 
116. 1886. 
REPLACED NAME: Eragrostis parvula Steud. 
= Eragrostis pectinacea var. miserrima (E. Fourn.) 
Reeder. 


Eragrostis purshii var. pauciflora E. Fourn., Mexic. Pl. 2: 
116. 1886. T-Specim.: 1 of 6. Mexico: Tampico, 
Berlandier 43 [OM]. 2 of 6. Mexico: Tampico, 
Berlandier 147 [OM]. 3 of 6. Mexico: Tampico, 
Berlandier 621 {OM: L]. 4 of 6. Mexico: Tampico, 
Berlandier 755 |OM]. 5 of 6. Mexico: Tampico, 
Berlandier 844 in part [OM]. 6 of 6. Mexico: 
Tampico, Berlandier 92] [OM]. 

NOTE: nom. nud. 

= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis pusilla Scribn. ex Beal, Grass. N. Amer. 2: 481, 
Nov 1896. T-Protol.: Mexico: Jalisco, Pringle 2327. 
T-Specim.: | of 1. Mexico: Jalisco: dry rocky hills 
near Guadalajara, 28 Oct 1889, C.G. Pringle 2327 
[HT: MSC; IT: LL, MO-2524105, US-1761637, US- 
748411, US-822031). 

NOTE: hom. illeg.; as Eragrostis "pusillus”. 

E. NAME: Eragrostis pusilla Hack. 

CORRECT NAME: Eragrostis scribneriana Hitche. 
= Eragrostis pringlei Mattei. 


Eragrostis pycnantha (Phil.) Parodi ex Nicora, Gayana, Bot. 
51(1): 4. 1994. Basionym: Poa pycnantha Phil. 
DST.: Chil. 
LVL. ACCPTC:: L. 
ALSO ACCPTD. BY: Nicora, E. G. (1994). 
SYN.: Poa pycnantha Phil. 


Eragrostis quitensis Yrin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
4,2(1): 76. 1836. T-Protol.: V. subsp. Brasilia. T- 
Specim.: | of 1. [HT: LE-TRIN-2388.01]. 

NOTE: nom. inval., as syn. of F. caldasii Kunth; based on 
Festuca quitensis Willd. in Nees Agrost. Bras. 1829. 
p 473, as syn. 

= Steirachne barbata (Trin.) Renvoize. 


Eragrostis rahmeri Phil., Verz. Antofagasta Pfl. 88. 1891. T- 
Protol.: Chile: in Quebrada de Guavifia provinciae 
Tarapaca lecta, /. Philippi s.n. T-Specim.: 1 of 1. 
Chile: Tarapaca, 13 Mar 1885, R.A. Philippi 359 (HT: 
SGO-37293;, IT: BAA-1080 (fragm.), CORD, SGO- 
62670, US-556538]. 

NOTE: as Eragrostis "Rahmeri". 
= Eragrostis mexicana (Hornem.) Link. 


Eragrostis refracta (Muhl. ex Elliott) Scribn., Mem. Torrey 
Bot. Club 5(4): 49. 1894. Basionym: Poa refracta 
Muhl. ex Elliott. 

DST.: Suri, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Correll, D. S. & M. C. Johnston 
(1970); Fernald, M. (1950); Gleason, H. A. & A. 
Cronquist (1968); Gleason, H. A. & A. Cronquist 
(1991); Godfrey, R. K. & J. W. Wooten (1979); 
Hitchcock, A. S. (1951); Judziewicz, E. J. (1990); 


Long, R. W. & O. Lakela (1971); Radford, A. E., H. 
E. Ahles & C. R. Bell (1968); Small, J. K. (1933). 

Eragrostis campestris Trin.; Eragrostis campestris 
var. refracta Chapm.; Eragrostis longeradiata 
Steud.; Eragrostis pectinacea var. refracta (Muhl. ex 
Elliott) Chapm.; Eragrostis virginica (Zuccagni) 
Steud.; Poa reflexa Elliott ex Scribn. & Merr.; Poa 
refracta Muhl. ex Elliott; Poa virginica Zuccagni. 


SYN:: 


Eragrostis reptans (Michx.) Nees, Fl. Bras. Enum. Pl. 2: 
514. 1829. Basionym: Poa reptans Michx. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Correll, D. S. 
& M.C. Johnston (1970); Cowan, C. P. (1983); Déll, 
J. C. (1878); Fernald, M. (1950); Foster, R. C. 
(1958); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Godfrey, R. 
K. & J. W. Wooten (1979); Gould, F. W. & R. Moran 
(1981); Great Plains Flora Association (1986); 
Hitchcock, A. S. (1951). 

Eragrostis capitata (Nutt.) Nash; Eragrostis reptans 
(Michx.) J. Presl; Eragrostis squarrosa E. Fourn.; 
Eragrostis weigeltiana Bush; Megastachya 
Jasciculata E. Fourn.; Megastachya fasciculata E. 
Fourn.; Megastachya reptans (Michx.) P. Beauv.; 
Neeragrostis reptans (Michx.) Nicora; Neeragrostis 
weigeltiana Bush; Poa capitata Nutt.; Poa dioica 
Michx. ex Poir.; Poa reptans Michx.; Poa 
weigeltiana Rchb. ex Trin. 


SYN.: 


Eragrostis reptans (Michx.) J. Presl, Relig. Haenk. 1(4-5): 
275. 1830. Basionym: Poa reptans Michx. 
NOTE: nom. illeg. superfl. et hom. 
E. NAME: Eragrostis reptans (Michx.) Nees. 
= Eragrostis reptans (Michx.) Nees. 


Eragrostis reptans vat. contracta Doll, Fl. Bras. 2(3): 148. 
1878. T-Protol.: In palude exsiccata ad S. Cruz (Jund) 
et ad vias prov. Minarum. T-Specim.: | of 2. Brazil: 
Santa Cruz, Lund s.n. [ST]. 2 of 2. Brazil: Minas 
Gerais, Warming s.n. [ST]. 

= Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb. 


Eragrostis reptans var. laxior Doll, Fl. Bras. 2(3): 148. 1878. 
T-Protol.: America boreali obvia, in Brasilia adhuo 
non lecta. 

= Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb. 


Eragrostis reptans var. pygmaea Doll, Fl. Bras. 2(3): 149. 
1878. T-Protol.: Brazil: prope Bahia, Blanchet 2960 

(IT: OXF]. 
= Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb. 


Eragrostis retinens Hack. & Arechav., Anales Mus. Nac. 
Montevideo 1: 445-446, t. 56. 1896. T-Protol.: 
Uruguay: comun en campos graminosos, prefiere los 
terrenos arenosos. T-Specim.: | of 1. [T: BAA-1083]. 

DST.: Arge, Braz, Urug. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Burkart, A. (1969); Cabrera, A. L. 
(1970); Rosengurtt, B., B. R. Arrillaga de Maffei & 
P. Izaguirre de Artucio (1970). 

SYN.: Eragrostis retinens var. pergaminensis Parodi. 


Eragrostis retinens var. pergaminensis Parodi, Notas Mus. 
La Plata, Bot. 2(11): 6. 1937. T-Protol.: Argentina: 


108 


Catalogue of New World Grasses: II. 


Pergamino, Prov. Buenos Aires, 2 Feb 1923, L.R. 
Parodi 4943. T-Specim.: 1 of 1. Argentina: Buenos 
Aires: Pergamino, hab. a orillas del ferrocarril Central 
Argentino cerca de la Estaci6n Mauel Ocampo, 2 Feb 
1923, L.R. Parodi 4943 [HT: BAA; IT: SI, US- 
1721254). 

= Eragrostis retinens Hack. & Arechav. 


Eragrostis retinorrhoea Steud., Syn. Pl. Glumac. 1: 268. 
1854. T-Protol.: Abyssinia. T-Specim.: 1 of 1. 
Ethiopia: Gapdiam, 21 Sep 1838, GHW. Schimper 
824 [ST: L, WAG]. 

= Eragrostis viscosa (Retz.) Trin. 


Eragrostis rigida (Vasey) Scribn., Proc. Acad. Nat. Sci. 
Philadelphia 1891: 304. 1891. Basionym: Diplachne 
rigida Vasey. 

= Eragrostis sessilispica Buckley. 


Eragrostis riobrancensis Judz. & P.M. Peterson, Brittonia 
42: 47, f. 1. 1990. T-Protol.: Guyana: Rupununi Distr.: 
3-11 km S of Sand Creek on road to Dadanawa, E of 
Rupununi River, ca. 02.03°N 59.30°W, elev. 110 m, 20 
Jun 1989, grassland with scattered small trees,, 
Peterson & Gopaul 7545 |HT: US; IT: BRG]. T- 
Specim.: | of 1. Guyana: Rupununi, 20 Jun 1989,P.M. 
Peterson & D. Gopaul 7545 (HT: US-3055106; IT: 
BRG]. 
DST.: Braz, Guya. 
LVL. ACCPTC:: I. 
ALSO ACCPTD. BY: Boechat, $. C. & H. M. Longhi- 
Wagner (2000); Judziewicz, E. J. & P. M. Peterson 
(1990); Judziewicz, E. J. (1990). 


Eragrostis robusta Stent, Bothalia 2: 288. 1927. T-Protol.: 
Maritzburg, Natal Angus 2, Govt. Herb. 13862. 
REPLACED NAME: Eragrostis valida Stent. 
BLOCKING NAME: Eragrostis valida Pilg. 
= Eragrostis curvula (Schrad.) Nees. 


Eragrostis rojasii Hack., Repert. Spec. Nov. Regni Veg. 7: 
373. 1909. T-Protol.: Paraguay: Sierra de Amambay, 
in campis siccis Punta Pora: Dec 1908, Rojas hbr. 
Hassler 9858. T-Specim.: | of 1. Paraguay: Sierra de 
Amambay, Dec 1908, Rojas hbr. Hassler 9858 |IT: 
BAA-1085 (fragm.), BAF, G, US-899326, US-79718]. 

NOTE: as Eragrostis "Rojasii". 

DST.: Arge, Braz, Para. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000). 


Eragrostis rufescens Schrad. ex Schult., Mant. 2: 319. 1824. 
T-Protol.: In Brasilia. Princeps Sereniss, Maximilian 
Neowidens s.n. [HT: LE]. T-Specim.: | of 1. Brazil, 
Maximilian Neowidens s.n. (HT: LE; IT: BAA-1087]. 

DST.: Beli, Boli, Braz, CoRi, Guat, Hond, Mexi, Nica. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 
(1987); Boechat, S. C. & H. M. Longhi-Wagner 
(2000); Davidse, G. (1994); Davidse, G., M. Sousa S. 
& A. O. Chater (editores generales) (1994); Doll, J. 
C. (1878); Filgueiras, T. S. (1991); Filgueiras, T. S. 
(1999): Killeen, T. J. (1990); Longhi-Wagner, H. M. 
(1990); Renvoize, S. A. (1984); Renvoize, S. A. 
(1988); Renvoize, S. A. (1998). 

Eragrostis floribunda Schrad. ex Schult.; Eragrostis 
inconstans Nees; Eragrostis inconstans — var. 


SYN:: 


rufescens (Schrad. ex Schult.) Nees; Eragrostis 
rufescens Schrad.; Eragrostis vahlii var. coarctata 
Doll; Poa rufescens Schrad. ex Schult. 


Eragrostis rufescens Schrad., Linnaea 12(4): 450. 1838. 
NOTE: isonym. 
E. NAME: Eragrostis rufescens Schrad. ex Schult. 
= Eragrostis rufescens Schrad. ex Schult. 


Eragrostis rufescens var. interrupta Doll, Fl. Bras. 2(3): 152. 
1878. 
ALSO ACCPTD. BY: DOIl, J.C. (1878). 
= Status not yet determined. 


Eragrostis rufescens var. laurentiana Benoist, Bull. Mus. 
Hist. Nat. (Paris) 17: 69. 1945. T-Specim.: 1 of 1. 
French Guiana: St. Laurent du Maroni, Herb. Lemee 
sn. (HT: P?]. 

= Eragrostis scaligera Salzm. ex Steud. 


Eragrostis rufescens var. mesoamericana Davidse, Novon 
2(2): 101. 1992. T-Protol.: Honduras: Morazan: 2.5 
km N of Zambrano on road to San Francisco de 
Soroguara, pine forest along small stream, on dry tuff 
outcrops, elev. 1370 m, | Jul 1970,R.W. Pohl & G. 
Davidse 12125 [HT: MO; IT: ISC]. 

DST.: Beli, CoRi, Guat, Hond, Mexi, Nica. 

LVL. ACCPTC:.: I. 

ALSO ACCPTD. BY: Balick, M. J., M. H. Nee & D. E. 
Atha (2000); Davidse, G. & R. W. Pohl (1992); 
Davidse, G. (1994); Davidse, G., M. Sousa S. & A. 
QO. Chater (editores generales) (1994); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000). 


Eragrostis rufescens Davidse var. rufescens. 

DST.: Boli, Braz, Para, Vene. 

LVL. ACCPTC:.: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000). 

Calotheca cuspidata Spreng.; Eragrostis acicularis 
Trin.; Eragrostis affinis Salzm. ex Steud. Eragrostis 
cuspidata (Schult.) Link; Eragrostis multipes 5S. 
Moore; Eragrostis polyneura Jedwabn.; 
Megastachya cuspidata Schult.; Poa cuspidata Roth. 


SYN:: 


Eragrostis rufescens var. spiciramea Dé6ll, Fl. Bras. 2(3): 
152. 1878. T-Specim.: | of 1. Brazil, Anonymous [IT: 
FL, K]. 
= Eragrostis solida Nees. 


Eragrostis salzmannii Steud., Syn. Pl. Glumac. 1: 277. 1854. 
T-Protol.: Maritimis Bahiae; Poa prolifera Salzm. 
herb. [herb. name] vix Swartz.), Salzmann s.n. T- 
Specim.: | of 1. Brazil: Bahia, no date, P. Sa/zmann 
s.n. (IT: K, US-911901, US-2767414]. 

= Eragrostis prolifera (Sw.) Steud. 


Eragrostis scaligera Salzm. ex Steud., Syn. Pl. Glumac. 1: 
277. 1854. T-Protol.: Bahia: (herb. Bah. sub: Poa). T- 
Specim.: | of 1. Brazil: Bahia, no date, P. Salzmann 
s.n. (HT: P; IT: US-911752]. 

NOTE: cites Poa scaligera Salzm. ex Steud., nom. nud. 

DST.: Braz, FrGu, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Judziewicz, E. J. (1990); Koch, S. D. 

- (1978); Renvoize, S. A. (1984); Wunderlin, R. P. 

(1998). 

SYN.: Eragrostis rufescens vat. laurentiana Benoist. 


Catalogue of New World Grasses: II. 


109 


Eragrostis sclerophylla Trin., Mem. Acad. Imp. Sci. St.- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 412. 1830. T- 
Protol.: Brasel: ad rivulum Jequitinhonha. T-Specim.: 
1 of 2. Brazil: ad rivul. Jequitinhonha, Dec. 1824, L. 
Riedel 1275 [HT: LE-TRIN; IT: LE, LE, LE, US- 
2765426 (fragm.), US-79719 (fragm.)]. 2 of 2. Brazil: 
in ripa rivi Jiquitiba et Jequitinhonha, Dec. 1824,s.c. 
1275 {IT: LE]. 

DST.: Braz. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Doll, J. C. (1878); Filgueiras, T. S. 
(1999). 

SYN.: Poa sclerophylla (Trin.) Kunth. 


Eragrostis scribneriana Hitche., Contr. U.S. Natl. Herb. 
17(3): 361. 1913. 
REPLACED NAME: Eragrostis pusilla Scribn. ex Beal. 
BLOCKING NAME: Eragrostis pusilla Hack. 
= Eragrostis pringlei Mattei. 


Eragrostis secundiflora J. Presl, Relig. Haenk. 1(4-5): 276. 
1830. T-Protol.. Hab. in Mexico, Haenke s.n. T- 
Specim.: | of 1. Mexico, 7: Haenke s.n. [HT: PR; IT: 
LE, MO-123764, US-79720 (fragm. ex PR)]. 

DST.: Boli, Braz, Colo, Guya, Mexi, Peru, USA, Vene. 

LVL. ACCPTC.:: I. 

ALSO ACCPTD. BY: Abrams, L. (1923); Beetle, A. A. 
(1977); Boechat, S. C. & H. M. Longhi-Wagner 
(2000); Breedlove, D. E. (1986); Cowan, C. P. 
(1983); Davidse, G. (1994); Davidse, G., M. Sousa S. 
& A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Filgueiras, T. S. (1999); Great Plains Flora 
Association (1986); Judziewicz, E. J. (1990); Killeen, 
T. J. (1990); Renvoize, S. A. (1984); Renvoize, S. A. 
(1998): Small, J. K. (1933); Sousa S., M. & E. F. 
Cabrera C. (1983); Wunderlin, R. P. (1998). 
Eragrostis beyrichii J.G. Sm.; Eragrostis compacta 
Salzm. ex Steud.; Eragrostis oxylepis (Torr.) Torr.; 
Eragrostis secundiflora var. capitata (E. Fourn.) 
Beetle; Eragrostis vahlii var. subfasciculata DOll, 
Eragrostis yucatana L.H. Harv.; Megastachya 
oxylepis var. capitata E. Fourn.; Poa secundiflora (J. 
Presl) Kunth. 


SYN: 


Eragrostis secundiflora var. capitata (E. Fourn.) Beetle, 
Phytologia 54(1): 1. 1983. Basionym: Megastachya 
oxylepis var, capitata E. Fourn. 

= Eragrostis secundiflora J. Pres. 


Eragrostis secundiflora subsp. oxylepis (Torr.) S.D. Koch, 
Rhodora 80(823): 397. 1978. Basionym:Poa oxylepis 
Torr. 

DST.: Mexi, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Great Plains Flora Association 
(1986); Koch, S. D. (1978). 

Eragrostis interrupta (Nutt.) Trel.; Eragrostis 
oxylepis (Torr.) Torr.; Eragrostis oxylepis var. 
beyrichii (J.G. Sm.) Shinners; — Eragrostis 
secundiflora var. capitata (E. Fourn.) Beetle; 
Eragrostis veraecrucis Rupr.; Megastachya oxylepis 
(Torr.) E. Fourn.; Megastachya oxylepis E. Fourn. ex 
Hemsl.; Poa oxylepis Torr. 


SYN:: 


Eragrostis secundiflora J. Pres! subsp. secundiflora. 
DST.: Boli, Braz, Guya, Mexi, Peru, Vene. 


LVL. ACCPTC.:: 1. 
ALSO ACCPTD. BY: Koch, S. D. (1978). 


Eragrostis seminuda Trin, Mem. Acad. Imp. Sci. St.- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 406. 1830. T- 
Protol.: V. subsp. Brasilia (Sto. Paulo), Riede/ 1641. T- 
Specim.: | of 3. Pr. Castro, March 1826,s.c. 250 [CT: 
LE]. 2 of 3. Brazil: S. Paolo: in arenosis pr. Parauna ad 
rivuli Jacare, Dec. 24, //4/ [HT: LE; IT: K]. 3 of 3. 
Brazil: St. Paulo in arenosis Pr. Parana ad Rivul Jacau, 
G.H. von Langsdorff s.n. (IT: US-77384 (fragm.)]. 

DST.: Arge, Braz, Para. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Dll, J. C. (1878); Filgueiras, T. S. 
(1999); Longhi-Wagner, H. M. (1990); Renvoize, S. 
A. (1984); Renvoize, S. A. (1988). 

SYN.: Eragrostis barbiglumis Jedwabn.; Eragrostis 
seminuda var. glabrata Doll; Eragrostis seminuda 
var. hirta Doll; Eragrostis seminuda var. pilosissima 
D6ll; Poa seminuda (Trin.) Kunth. 


Eragrostis seminuda var. glabrata Doll, Fl. Bras. 2(3): 143. 
1878. T-Specim.: | of 2. Brazil: Prov. S. Paulo, in 
campsi siccis graminosique vulgatissima, prope Castro 
et alibi, Riedel sn. [ST]. 2 of 2. Brazil: Prov. 
Minarum: in campis prope Lagoa Santa, Warming s.n. 
[ST]. 

NOTE: placed here with doubt in E. seminuda. 
= Eragrostis seminuda Trin. 


Eragrostis seminuda var. hirta Doll, Fl. Bras. 2(3): 143. 
1878. T-Specim.: | of 1. [T], perhaps LE TRIN- 
2398.01 in part [rjs 2001]. 

NOTE: erroneously attributed to Trin. by Doell. 

= Eragrostis seminuda Trin. 


Eragrostis seminuda var. pilosissima Doll, Fl. Bras. 2(3): 
143. 1878. T-Specim.: | of 1. [T], see notes under var. 
glabrata. 

= Eragrostis seminuda Trin. 


Eragrostis sessilispica Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 97. 1862. T-Protol.: USA: Texas: near 
Austin. T-Specim.: | of 1. USA: Texas: near Austin, 
Buckley s.n. {LT: PH], LT designated by Hitchcock, 
Man. Grass. U.S. 852 (1935), but without citing a 
specific sheet in a specific herbarium. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Correll, D. S. 
& M. C. Johnston (1970); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Great 
Plains Flora Association (1986); Harvey, L. H. 
(1984); Hitchcock, A. S. (1951). 

Acamptoclados  sessilispicus (Buckley) Nash; 
Diplachne rigida Vasey; Eragrostis rigida (Vasey) 
Scribn.; Leptochloa rigida Munro ex Vasey; 
Leptochloa rigida Munro ex Vasey; Rhabdochloa 
rigida (Munro ex Vasey) Kuntze. 


SYN.: 


Eragrostis setifolia (Benth.) Steud., Syn. Pl. Glumac. 1: 274. 
1854. Basionym: Poa setifolia Benth. 
NOTE: hom. illeg. 
E. NAME: Eragrostis setifolia Nees. 
= Eragrostis pastoensis (Kunth) Trin. 


110 


Catalogue of New World Grasses: II. 


Eragrostis silveana Swallen, Amer. J. Bot. 19(5): 438, f. 3. 
1932. T-Protol.: USA: Texas: collected along railroad 
near Taft, | Oct 1931, W.A. Silveus 360 [HT: US- 
1501596]. T-Specim.: | of 1. USA: Texas: Bexar Co., 
1-31 Oct 1931, WA. Silveus 360 [HT: US-1501596; 
IT: US-1501597, US-1501598, US-1501599, US- 
1501600]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Correll, D. S. 
& M. C. Johnston (1970); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J.  Valdes-Reyna (2000); 
Hitchcock, A. S. (1951). 


Eragrostis simplex Scribn., Bull. Div. Agrostol., U.S.D.A. 7: 
(ed. 3)250, f. 244. 1900. T-Protol.: USA: Florida, 
Curtiss 6073. T-Specim.: 1 of 1. USA: Florida: 
Istachatta, dooryard, 26 Aug 1897,4.H. Curtiss 6073 
[LT: US-314438; ILT: US-7481235], LT designated 
by Koch, Rhodora 80: 395 (1978). 

NOTE: as £. brownii Nees (?) 7: (ed. 1) 262, f. 244 (1987). 
= Eragrostis cumingii Steud. 


Eragrostis simpliciflora (J. Presl) Steud., Syn. Pl. Glumac. 
1: 278. 1854. Basionym: Megastachya simpliciflora J. 
Presl. 

DST.: CoRi, ElSa, Guat, Hond, Mexi, Nica, Pana. 

LVL. ACCPTC:.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Berendsohn, 
W. G. & A. E. Araniva de Gonzalez (1991); 
Breedlove, D. E. (1986); Davidse, G. (1994); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J, Valdes-Reyna (2000); McClure, F. A. 
(1955); Molina R., A. (1975); Pohl, R. W. (1980). 
Megastachya simpliciflora J. Pres|; Poa simpliciflora 
(J. Presl) Kunth. 


SYN.: 


Eragrostis solida Nees, Fl. Bras. Enum. Pl. 2: 501. 1829. T- 
Protol.: Brasilia in altis pascuis inter Gaspar Soares et 
Tapanhoacanga provinciae Minarum, Martius s.n. 
{HT: B]. T-Specim.: | of 2. Brasilia, FL Se/low [ST?: 
LE (ex B)]. 2 of 2. Brazil: Minas Gerais: in altis 
Pascuis inter Gasper Soares et Tapanhoacanga, f° K.P. 
von Martius s.n. [HT: B; IT: BAA-1094 (fragm. ex B), 
M, US-2767404 (fragm.)]. 

DST.: Arge, Boli, Braz, Para. 
LVL. ACCPTC.: |. 
ALSO ACCPTD. BY: Boechat, S$. C. & H. M. Longhi- 

Wagner (2000); Filgueiras, T. S. (1991); Filgueiras, 

T. S. (1999); Foster, R. C. (1958); Killeen, T. J. 

(1990); Longhi-Wagner, H. M. (1990); Renvoize, S. 

A. (1984); Renvoize, S. A. (1998). 

Eragrostis eucampta Pilg.; Eragrostis 
mattogrossensis to. glabrescens Pilg.; Eragrostis 
mattogrossensis Pilg., Eragrostis rufescens var. 
spiciramea Doll; ~~ Eragrostis — solida var. 
mattogrossensis (Pilg.) Pilg.; Poa solida (Nees) 
Kunth. 


SYN.: 


Eragrostis solida var. mattogrossensis (Pilg.) Pilg., Bot. 
Jahrb. Syst. 70(3): 346. 1939. Basionym: Eragrostis 
mattogrossensis Pilg. 

= Eragrostis solida Nees. 


Eragrostis soratensis Jedwabn., Bot. Arch. 5(3-4): 213. 
1924. T-Specim.: | of 2. Bolivia: La Paz: vicinis 
Sorata, colle Ticacirca, Feb 1858, G. Mandon 1331 


[ILT: US-1126604], LT designated by Hitchcock, 
Contr. U.S. Natl. Herb. 24(8): 343 (1927). 2 of 2. 
Bolivia: la Paz prope Huancapampa, Pflanz 366 [ST]. 
DST.: Boli, Peru. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Renvoize, S. A. (1998); Saravia, E. 
F. (1996); Tovar, O. (1986); Tovar, O. (1993). 
Eragrostis botterii E.  Fourn.; — Eragrostis 
brachypodon Hack. 


SYN:: 


Eragrostis spectabilis (Pursh) Steud., Nomencl. Bot. (ed. 2) 
1: 564. 1840. Basionym: Poa spectabilis Pursh. 

DST.: Cana, Mexi, USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Correll, D. S. 
& M. C. Johnston (1970); Darke, R. (1999); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Fernald, M. (1950); Gleason, H. A. & A, 
Cronquist (1968); Gleason, H. A. & A. Cronquist 
(1991); Godfrey, R. K. & J. W. Wooten (1979); 
Great Plains Flora Association (1986); Hitchcock, A. 
S. (1951); Long, R. W. & O. Lakela (1971); Radford, 
A. E., H. E. Ahles & C. R. Bell (1968); Scoggan, H. 
J. (1978 [1979]); Small, J. K. (1933); Wunderlin, R. 
P. (1998). 

Eragrostis geyeri Steud.; Eragrostis pectinacea var. 
spectabilis (Pursh) A. Gray; Eragrostis spectabilis 
var. sparsihirsuta Farw., Eragrostis spectabilis var. 
spectabilis, Eragrostis velutina Schrad.,; Erochloe 
spectabilis (Pursh) Raf. ex B.D. Jacks.; Megastachya 
spectabilis (Pursh) Roem. & Schult.; Poa amabilis 
Walter; Poa hirsuta var. spectabilis (Pursh) Torr.; 
Poa pectinacea Geyer ex Steud.; Poa spectabilis 
Pursh; Poa villosa Beyr. ex Schrad. 


SYN.: 


Eragrostis spectabilis var. sparsihirsuta Farw., Amer. Midl. 
Naturalist 10: 306, 1927, 
NOTE: cites E. pectinacea sensu Nees, Steud., A. Gray, non 
Poa pectinacea Michx. 
= Eragrostis spectabilis (Pursh) Steud. 


Eragrostis spectabilis (Pursh) Steud. var. spectabilis. 
= Eragrostis spectabilis (Pursh) Steud. 


Eragrostis spicata Vasey, Bot. Gaz. 16(5): 146. 1891. T- 
Protol.: Mexico: collected at San José del Cabo, Lower 
California, 1890, 7.8. Brandegee s.n. T-Specim.: | of 
1. Mexico: Baja California Sur: San José del Cabo, 12 
Sep 1890, 7.8. Brandegee 10 [HT: US-1761638]. 

DST.: Arge, Mexi, Para, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Burkart, A. 
(1969); Correll, D. S. & M. C. Johnston (1970): 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Gould, F. W..& R. Moran (1981); 
Hitchcock, A. S. (1951). 

SYN.: Sporobolus tenuispica Hack. 


Eragrostis sporoboloides J.G. Sm. & Bush, Annual Rep. 
Missouri Bot. Gard. 6: 116, t. 54. 1895. T-Protol.: 
USA: Sapula, Indian Territory: Jul 1894, BF. Bush 
SM. 

= Eragrostis hirsuta (Michx.) Nees. 


Eragrostis stenoclada J, Presl, Relig. Haenk. 1(4-5): 278. 
1830. T-Protol.: Habitat in montanis Peruviae [changes 
to "Hab. ad Monte-Rey, Californiae," in the 
Addendum on p. 351], Haenke s.n. [HT: PR], probably 


Catalogue of New World Grasses: II. 


111 


Philippines, not Peru as cited in protologue; see notes 
for more data on the type. T-Specim.: 1 of 2. 
Philippines: Luzon I., 7. Haenke s.n. (IT: US-79721 
(fragm.)]. 2 of 2. Peru, 7. Haenke s.n. (IT: US-79722 
(fragm.)]. 

= Eragrostis gangetica (Roxb.) Steud. 


Eragrostis stenophylla Hochst. ex Miq., Anal. Bot. Ind. 2: 27. 
1851. T-Protol.: prope urbem Mangalor, Metz 664, 
"Hochst. mss.". T-Specim.: | of 1. India: Mangalore 
[ST: L]. 

NOTE: also Verh. Nederl. Inst. II. 4: 39 (1851). 
= Eragrostis gangetica (Roxb.) Steud. 


Eragrostis subatra Jedwabn., Bot. Arch. 5(3-4): 202. 1924. 
T-Protol.: Bolivia: prope La Paz, Rusby 80, The 
citation of Rusby as the collector is an error for Bang, 
fide Nicora, Boissiera 54: 63 (1998). T-Specim.: | of 
1. Bolivia: prope La Paz, 1889, M. Bang 80 [IT: K, 
US-822065], The collector is erroneously cited as 
Rusby in the protologue. 

= Eragrostis nigricans (Kunth) Steud. 


Eragrostis sudans Griseb., Cat. Pl. Cub. 227. 1866. T-Protol.: 
Cuba or., C. Wright 3422. T-Specim.: 1 of 1. Cuba, 
1860-1864, C. Wright 3422 [HT: ?; IT: MO-2067270, 
US-911645, US-79723 (fragm.)]. 

= Eragrostis glutinosa (Sw.) Trin. 


Eragrostis superba Peyr., Sitzungsber. Kaiserl. Akad. Wiss., 
Math.-Naturwiss. Cl. 38: 584. 1860. T-Protol.: In locis 
umbrosis prope Benguela ac Loanda, Wawra 244. T- 
Specim.: | of 1. Angola, Wawra 244 [HT: W]. 

DST.: Arge, Boli, Guya, Hond, Mexi, USA, Urug. 

LVL. ACCPTC.: 1; cultivated, introduced. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Bor, N. L. 
(1960); Cope, T. A. (1992); Davidse, G. (1994); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gibbs Russell, 
G. E, W. G. Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987); Judziewicz, E. J. (1990); 
Molina R., A. (1975); Phillips, S. (1995). 


Eragrostis swallenii Hitche., J. Wash. Acad. Sci. 23(10): 
451. 1933. T-Protol.: USA: Collected on sandy prairie 
at Riviera, Texas, 8 Jun 1931,/.R. Swallen 1847 (HT: 
US-1535332]. T-Specim.: | of 1. USA: Texas, 8 Jun 
1931, AR. Swallen 1847 (HT: US-1535332; IT: US- 
3160855, US-2852654]. 

DST.: Mexi, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Correll, D. S. 
& M. C. Johnston (1970); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Hitchcock, A. S. (1951). 


Eragrostis tef (Zucc.) Trotter, Boll. Soc. Bot. Ital. 1918: 62. 
1918. Basionym: Poa tef Zucc. 
DST.: Braz, USA. 
LVL. ACCPTC.: 1; cultivated, introduced. 
ALSO ACCPTD. BY: Anonymous (1986); Boechat, S. C. 
& H. M. Longhi-Wagner (2000); Bor, N. L. (1960); 
Cope, T. A. (1992); Gibbs Russell, G. E, W. G. 
Welman, E. Reitief, K. LL. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 


Nicholas (PRECIS) (1987); Hitchcock, A. S. (1951); 
Phillips, S. (1995). 

Cynodon abyssinicus (Jacq.) Raspail; Eragrostis 
abessinica (Jacq.) Link; Eragrostis abyssinica (Jacq.) 
Link; Eragrostis pilosa subsp. abyssinica (Jacq.) 
Asch. & Graebn.; Eragrostis pilosa var. tef (Zucc.) 
Fiori; Poa abyssinica Jacq.; Poa cerealis Salisb.; Poa 
tef Zucc. 


SYN:: 


Eragrostis tenax (Kunth) Steud., Nomencl. Bot. (ed. 2) 1: 
564. 1840. Basionym: Poa tenax Kunth. 
= Eragrostis condensata (J. Presl) Steud. 


Eragrostis tenella (L.) P. Beauv. ex Roem. & Schult., Syst. 
Veg. 2: 576. 1817. Basionym: Poa tenella L. 
NOTE: see Notes under Eragrostis amabilis for the use of 
this name. 
= Eragrostis amabilis (L.) Wight & Arn. ex Nees. 


Eragrostis tenella var. plumosa (Retz.) Stapf, Fl. Brit. India 
7: 315. 1896. Basionym: Poa plumosa Retz. 
= Eragrostis amabilis (L.) Wight & Arn. ex Nees. 


Eragrostis tenella var. viscosa (Retz.) Stapf, Fl. Brit. India 
7(22): 315-316. 1896. Basionym: Poa viscosa Retz. 
= Eragrostis viscosa (Retz.) Trin. 


Eragrostis tenuifolia (A. Rich.) Hochst. ex Steud., Syn. PI. 
Glumac. 1: 268. 1854. Basionym: Poa tenuifolia A. 
Rich. 

DST.: Arge, Boli, Braz, Chil, Colo, CoRi, Ecua, ElSa, Guat, 

Hond, Mexi, Pana, Peru, Vene. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Bor, N. L. (1960); Brako, L. & J. L. 
Zarucchi (1993); Breedlove, D. E. (1986); Davidse, 
G. (1994); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J.  Valdes-Reyna (2000); 
Filgueiras, T. S. (1991); Gibbs Russell, G. E, W. G. 
Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987); Jorgensen, P. M. & C. 
Ulloa U. (1994); Jorgensen, P. M. & S. Leon-Yanez 
(eds.) (1999); Killeen, T. J. (1990); Nelson, C. 
(1979); Phillips, S. (1995); Pohl, R. W. (1980): 
Tovar, O. (1993). 

Eragrostis collocarpa &. Schum.; Eragrostis 
parviglumis Hochst. ex Steud.; Poa tenuifolia A. 
Rich. 


SYN:: 


Eragrostis tenuis Steud., Syn. Pl. Glumac. 1: 273. 1854. 
NOTE: nom. nov., as acomb., but based on hom. illeg. 
REPLACED NAME: Poa tenuis Elliott. 

BLOCKING NAME: Poa tenuis Clairv. 
= Eragrostis capillaris (L.) Nees. 


Eragrostis tenuis A. Gray, Manual (ed. 6) 661. 1890. T- 
Protol.: USA: Ohio to Illinois, Kansas and southward, 
sandy soil. 

NOTE: hom. illeg. 
E. NAME: Eragrostis tenuis Steud. 
= Eragrostis trichodes (Nutt.) Alph. Wood. 


Eragrostis tenuis var. texensis Vasey, Contr. U.S. Natl. Herb. 
1(2): 59. 1890. T-Protol.: USA: Texas: In the region of 
the rio Grande, in Texas, from Brazos Santiago to El 
Paso Co., Reverchon & Nealley s.n. T-Specim.: | of 1. 


112 


Catalogue of New World Grasses: II. 


USA: Texas, 1884, G.C. Nealley & J. Reverchon 848 
[HT: US-1761634]. 
= Eragrostis trichodes (Nutt.) Alph. Wood. 


Eragrostis tephrosanthos Schult., Mant. 2: 316. 1824. T- 
Protol.: Martinica. T-Specim.: 1 of 1. Martinique, no 
date, F.W. Sieber 33 (Fl. Martin no.) (HT: M; IT: kK, 
MO-201170, P, US-1127056, US-1127055]. 

= Eragrostis pectinacea var. miserrima (E. Fourn.) 
Reeder. 


Eragrostis terecaulis Renvoize, Gram. Bolivia 324. 1998. T- 
Protol.: Bolivia: La Paz: Aroma, Patacamaya, Lara & 
Parker 3le (HT: MO]. 

DST.: Boli. 
LVL, ACCPTC.: 1. 
ALSO ACCPTD. BY: Renvoize, S. A. (1998). 


Eragrostis thunbergiana var. atrata Schweinf., Bull. Herb. 
Boissier, ser. 2, App. 2: 101. 1894. T-Protol.: Eritrea: 
Kohaito, Schwinfurth 75. T-Specim.: | of 1. [HT: ?; 
IT:K]. 

= Eragrostis curvula (Schrad.) Nees. 


Eragrostis tracyi Hitche., Amer. J. Bot. 21(3): 130, f. 1. 
1934. T-Protol.: USA: Florida: collected on Sanibel 
Island, 19 May 1901, S.M. Tracy 7/68 [HT: US- 
441983]. T-Specim.: | of 1. USA: Florida: Lee Co.: 
Sanibel Island, 19 May 1901, S8.M. Tracy 7168 [HT: 
US-441983; IT: US-909970]. 

= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis tremula Hochst. ex Steud., Syn. Pl. Glumac. 1: 
269. 1854. T-Protol.: Sendung von Pflanzen aus 
Nubien und Kordofan, Kotschy, Hrbr. un. it. 6. T- 
Specim.: | of |. Nubia: in arenosis prope pagum Abu- 
Gerad, 18 Sept. 1839, Kotschy 6 [IT: LE]. 

DST.: Braz. 

LVL. ACCPTC:.: 1; introduced. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (2000); Boechat, S. C. & P. M. Peterson 
(1995); Bor, N. L. (1960); ORSTOM_ (1988); 
Phillips, S. (1995), 


Eragrostis triangularis Henrard, Repert. Spec. Nov. Regni 
Veg. 23(18-25): 301. 1927. T-Protol.: Paraguay: 
Coaguazu, in campis, 24 Mar 1876, B. Balansa 236 
[HT: L-908.88-310]. T-Specim.: | of 1. Paraguay: 
Caaguazu: in campis, 24 Mar 1876, B. Balansa 236 
[HT: L-908.88-310; IT: K, P, US-77378 (fragm.), US- 
91397 (fragm.)]. 

DST.: Para. 
LVL. ACCPTC:: 1. 


Eragrostis trichocolea Hack. & Arechav., Anales Mus. Nac. 
Montevideo 1: 444. 1896. T-Protol.: En terrenos 
arenosos de la costa del Rio de la Plata. T-Specim.: 1 
of J. Uruguay [T: BAA-3933 (fragm.), MVFA 
(fragm.)]. 

NOTE: also, Las Gramim. Urug, 384 (1896). 

DST.: Arge, Braz, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Boechat, S. C. & H. M. Longhi-Wagner 
(2000); Breedlove, D. E. (1986); Burkart, A. (1969); 
Hitchcock, A. S. (1951); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 
Small, J. K. (1933); Wunderlin, R. P. (1998). 


Eragrostis trichocolea var. floridana (Hitchc.) Witherspoon, 
Ann. Missouri Bot. Gard. 64: 328. 1977. Basionym: 
Eragrostis floridana Hitche. 

= Eragrostis polytricha Nees. 


Eragrostis trichodes (Nutt.) Alph. Wood, Class-book Bot. 
(ed. 1861) 796. 1861. Basionym: Poa trichodes Nutt. 

DST.: USA. 

LVL. ACCPTC.:: 1; cultivated. 

ALSO ACCPTD. BY: Anonymous (1986); Bor, N. L. 
(1960); Correll, D. S. & M. C. Johnston (1970); 
Darke, R. (1999); Fernald, M. (1950); Gleason, H. A. 
& A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Great Plains Flora Association 
(1986); Hitchcock, A. S. (1951). 

Eragrostis capillacea  Jedwabn.; Eragrostis 
grandiflora J.G. Sm. & Bush; Eragrostis pilifera 
Scheele; Eragrostis tenuis A. Gray; Eragrostis tenuis 
var. fexensis Vasey; Eragrostis trichodes var. pilifera 
(Scheele) Fernald; Eragrostis trichodes — var. 
trichodes, Poa trichodes Nutt. 


SYN: 


Eragrostis trichodes var. pilifera (Scheele) Fernald, Rhodora 
40(477): 331. 1938. Basionym: Eragrostis pilifera 
Scheele. 

= Eragrostis trichodes (Nutt.) Alph. Wood. 


Eragrostis trichodes (Nutt.) Alph. Wood var. trichodes. 
= Eragrostis trichodes (Nutt.) Alph. Wood. 


Eragrostis trichophora Coss. & Durieu, Bull. Soc. Bot. 
France 2: 311. 1855. 

NOTE: nom. nov., see also Expl. Sci. Alger. 2: 150 (1855); 
based on E. geniculata Durieu (sub. Poa) in Expl. 
Sci. Alg. t. 40, f. 4, non E. geniculata Nees, Accd. to 
Chase & Niles, Index Grass Sp. 

REPLACED NAME: Eragrostis geniculata (Dur.) Dur. 

BLOCKING NAME: Eragrostis geniculata Nees & 
Meyen. 

DST.: USA. 

LVL. ACCPTC.: 1; cultivated, introduced. 

ALSO ACCPTD. BY: Anonymous (1986); Gibbs Russell, 
G. E, W. G. Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987); Phillips, S. (1995). 

SYN.: Eragrostis atherstonei Stapf; Eragrostis henrardii 
Jansen. 


Eragrostis tricuspis Trin. Mem. Acad. Imp. Sci. St.- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 414. 1830. T- 
Protol.: USA: Carolinas, Beauv. s.n. 

NOTE: nom. illeg.; base on Tricuspis caroliniana Beauv., 
but no blocking name present in 1830. 
= Tridens flavus (L.) Hitche. 


Eragrostis triflora Ekman, Ark. Bot. 11(4): 42, t. 4, f. 1. 
1912. T-Protol.: Bonpland, locis siccis, saxosis, apricis 
(campinas, incol.) trans "Martires chico" amnem, 
20.1.08, Ekman 7/4 [HT: S]. T-Specim.: 1 of 1. 
Argentina: Misiones, 20 Jan 1908, E.L. Ekman 7/4 
[HT: S; IT: BAA-1100 (fragm.), CORD, SI, US- 
602662, US-77381 (fragm.)]. 

= Eragrostis airoides Nees. 


Eragrostis tristis Jedwabn., Bot. Arch. 5(3-4): 205. 1924. T- 
Specim.: | of 3. Bolivia: ad Huancapamba, 15 Feb 
1910, Pflanz 359 [ST: BAA-L101 (fragm. ex B)]. 2 of 
3. Bolivia: Quechisla, K. Bender 19 [ST: ?; IST: US- 


Catalogue of New World Grasses: II. 


113 


2767396 (fragm.)]. 3 of 3. Peru: prope Tacna, Meyen 
s.n. [ST]. 
= Eragrostis nigricans (Kunth) Steud. 


Eragrostis uninervia (J. Presl) Steud., Syn. Pl. Glumac. 1: 
278. 1854. Basionym: Megastachya uninervia J. Presl. 
= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Eragrostis unioloides (Retz.) Nees ex Steud., Syn. PI. 
Glumac. |: 264. 1854. Basionym: Poa unioloides 
Retz. 

DST.: Cari, Ecua, FrGu, Guya. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Bor, N. L. (1960); Breedlove, D. E. 
(1986); Davidse, G. (1994); Davidse, G., M. Sousa S. 
& A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Godfrey, R. K. & J. W. Wooten (1979); 
Hitchcock, A. S. (1951); Jorgensen, P. M. & S. Leén- 
Yanez (eds.) (1999); Judziewicz, E. J. (1990); Small, 
J. K. (1933); Wunderlin, R. P. (1998). 

SYN.: Poa unioloides Retz. 


Eragrostis unionis Steud., Syn. P|. Glumac. 1: 273. 1854. T- 
Protol.: USA: Ohio: Miami, "Poa capillaris Hrbr. 
Frank. un. it 1835 (non1837) Miami, Am. sptr." Ohio, 
Accd. to Chase & Niles, Index Grass Sp. T-Specim.: | 
of 1. USA: Ohio: in ditione Miami, 1835, hrbr. Frank 
[IT: herb. Caen; IT: US-2851266 (fragm.)]. 

= Eragrostis pectinacea (Michx.) Nees. 


Eragrostis urbaniana Hitche., Symb. Antill. 7: 167. 1912. 
T-Protol.: West Indies: Netherlands Antilles: Ins. 
Bonaire, 25 Feb 1885, WFR. Suringar s.n. (HT: US- 
690359]. T-Specim.: 1 of 1. Netherlands Antilles: 
Bonaire, 25 Feb 1885, W.F.R. Suringar s.n. [HT: US- 
690359]. 

DST.: Cari. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Hitchcock, A. S. 
(1936); Lindeman, J. C. & A. L. Stoffers (1963); 
Liogier, H. & L. Martorell (1982). 


Eragrostis vahlii (Roem. & Schult.) Nees, Fl. Bras. Enum. PI. 
2: 499-500. 1829. Basionym: Poa vahlii Roem. & 
Schult. 

NOTE: var. alfa Nees. 
= Eragrostis maypurensis (Kunth) Steud. 


Eragrostis vahlii var. coarctata Doll, Fl. Bras. 2(3): 155. 
1878. T-Protol.: Brazil, 2 syntypes cited. T-Specim.: 1 
of 2. Brazil: ad Congo Rubim, no date, J.B.E. Pohl 
3247 [ST: IST: US-1126606, W]. 2 of 2. Brazil: Bahia: 
prope Ilheos in sabulosis ad ostia fluvii Rio de 
Caxoeira, Martius s.n. [ST: MJ. 

NOTE: pro parte. 
= Eragrostis rufescens Schrad. ex Schult. 


Eragrostis vahlii var. polyantha Doll, Fl. Bras. 2(3): 155. 
1878. T-Protol.: [Brazil], no specimens mentioned for 
this var. 

= Eragrostis maypurensis (Kunth) Steud. 


Eragrostis vahlii var. sejuncta DOll, Fl. Bras. 2(3): 155. 1878. 
T-Protol.: [Brazil], no specimens mentioned for this 
var. 

ALSO ACCPTD. BY: Dall, J. C. (1878). 
= Status not yet determined. 


Eragrostis vahlii var. subfasciculata DOll, Fl. Bras. 2(3): 155. 
1878. T-Protol.: [Brazil], no specimens mentioned for 
this var. 

= Eragrostis secundiflora J. Presl. 


Eragrostis vallsiana Boechat & Longhi-Wagner, lheringia, 
Bot. 51(2): 179, f. 1A-C, 2-3A-D. 1998. T-Protol.: 
Brasilia: Mato Grosso do Sul: Corumba, 19 Nov 1977, 
Allem & Vieria 1335 [HT: CEN; IT; MO, US]. 

DST.: Braz. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Boechat, S. C. 
Wagner (1998); Boechat, S. C. 
Wagner (2000). 


& H. M. Longhi- 
& H. M. Longhi- 


Eragrostis velutina Schrad., Linnaea 12(4): 451. 1838. T- 
Protol.: USA: Carolina, Beyrich. 
NOTE: cites "Poa villoa Beyr.—-Carolina" (pub. as syn.). 
= Eragrostis spectabilis (Pursh) Steud. 


Eragrostis veraecrucis Rupr., Bull. Acad. Roy. Sci. Bruxelles 
9(2): 235. 1842. 
NOTE: nom. nud.; cited again by Fourn. 1886 as syn. of 
Megastachya oxylepis captitata Fourn. 
= Eragrostis secundiflora subsp. oxylepis (Torr.) S.D. 
Koch. 


Eragrostis vernix Boechat & Longhi-Wagner, Iheringia, 
Bot. 51(2): 181, f. ID-E, 2-3E-H. 1998. T-Protol.: 


Brasilia: Pernambuco: Garanhuns, 18 Aug 1969, 
Tenorio 69-808 (HT: IPA]. 
DST.: Braz. 


LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (1998); Boechat, S. C. & H. M. Longhi- 
Wagner (2000). 


Eragrostis verticillata (Cav.) P. Beauv., Ess. Agrostogr. 71, 
[162 queried], 176. 1812. Basionym: Poa verticillata 
Cav. 
NOTE: nom. nud.????. 
= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis verticillata (Cav.) Roem. & Schult., Syst. Veg. 2: 
575. 1817. Basionym: Pea verticillata Cav. 
NOTE: hom. illeg.???. 
= Eragrostis pilosa (L.) P. Beauv. 


Eragrostis villamontana Jedwabn., Bot. Arch. 5(3-4): 197. 
1924. T-Protol.: Bolivia: prov. Tarija, ad Villamontes, 
Pflanz 632. T-Specim.: 1 of 1. Bolivia: Tarija: 
Villamontes, K. Pflanz 632 [HT: B; IT: BAA-1105 
(fragm. ex B), US-77380 (fragm.)]. 

NOTE: as Eragrostis "Villamontana". 
= Eragrostis orthoclada Hack. 


Eragrostis villosa Salzm. ex Steud., Syn. Pl. Glumac. 1: 276. 

1854. T-Protol.: Brasil, Sa/zmann s.n. T-Specim.: | of 
1. Brazil: Bahia, P. Salzmann s.n. (Herb. Fischer) 
[HT: P; IT: K, LE (2 sheets), BAA-1106 (fragm.), US- 
79727 (fragm.)]. 

NOTE: hom. illeg. 

E. NAME: Eragrostis villosa Trin. 

= Eragrostis articulata (Schrank) Nees. 


Eragrostis virescens J. Presl, Relig. Haenk. 1(4-5): 276. 
1830. T-Protol.: Hab. in Cordilleris chilensibus, 
Haenke s.n. T-Specim.: | of 1. Chile, 7. Haenke s.n. 


114 


Catalogue of New World Grasses: I. 


[HT: PR; IT: B, BAA-1100 (fragm. ex B), LE-TRIN- 
2413.01, US-2942410 (fragm.)]. 
= Eragrostis mexicana subsp. virescens (J. Presl) S.D. 
Koch & Sanchez Vega. 


Eragrostis virescens var. trachyphylla Hack., Anales Mus. 


Nac. Buenos Aires 13: 505. 1906. T-Protol.: 
Argentina: Prov. Tucuman, Lillo hb. 3192 [hb. 
Stuckert 14886]. T-Specim.: | of 1. Argentina: 


Tucuman, Stuckert herb. arg. 14866 ex Lillo Herb. 
arg. 3192 [IT: US-2942411 (fragm.)]. 
= Eragrostis pastoensis (Kunth) Trin. 


Eragrostis virginica (Zuccagni) Steud., Syn. Pl. Glumac, 1: 
273. 1854. Basionym: Poa virginica Zuccagni. 
= Eragrostis refracta (Muhl. ex Elliott) Scribn. 


Eragrostis virletii E. Fourn., Mexic. Pl. 2: 116. 1886. T- 
Protol.: Mexico: San Luis de Potosi, Virlet 1391. T- 
Specim.: | of 1. Mexico: San Luis Potosi, 1851, A¢ 
Virlet D'Aoust 1391 [I1T: US-2850749 (fragm.)]. 

= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis viscosa (Retz.) Trin., Mem. Acad. Imp. Sci. St.- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 397. 1830. 
Basionym: Poa viscosa Retz. 

DST.: CoRi, Ecua, ElSa, Guat, Hond, Mexi, Nica, Pana. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Berendsohn, W. G. & A. E. Araniva de 
Gonzalez (1991); Bor, N. L. (1960); Breedlove, D. E. 
(1986); Davidse, G. (1994); Davidse, G., M. Sousa S. 
& A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Gibbs Russell, G. E, W. G. Welman, E. 
Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987); Gould, F. W. & R. Moran (1981); Hitchcock, 
A. S. (1927); Jorgensen, P. M. & S. Leén-Yanez 
(eds.) (1999); McClure, F. A. (1955); McVaugh, R. 
(1983); Molina R., A. (1975); Phillips, S. (1995); 
Pohl, R. W. (1980); Shreve, F. & IL L. Wiggins 
(1964). 

Eragrostis retinorrhoea Steud.; Eragrostis tenella 
var. viscosa (Retz.) Stapf; Eragrostis viscosa var. 
pilosissima (Hochst. ex A. Rich.) Hochst.; Eragrostis 
warmingti Hack., Poa viscosa Retz.; Poa viscosa var. 
pilosissima Hochst. ex A. Rich. 


SYN.: 


Eragrostis viscosa Scribn., Bull. Div. Agrostol., U.S.D.A. 11: 
51, t. 7. 1898. T-Specim.: | of 2. USA: Texas: 
Midland, 2 Aug 1897, 1G. Smith s.n. [ST: US- 
1768944]. 2 of 2. USA: Texas: Laredo, Mrs. Anna B. 
Nickels s.n. [ST: US]. 

NOTE: hom. illeg. 
E. NAME: Eragrostis viscosa (Retz.) Trin. 
= Eragrostis curtipedicellata Buckley. 


Eragrostis viscosa var. pilosissima (Hochst. ex A. Rich.) 
Hochst., Flora 38: 329. 1855, Basionym: Poa viscosa 
var. pilosissima Hochst. ex A. Rich. 

= Eragrostis viscosa (Retz.) Trin. 


Eragrostis vulgaris J. Pres| ex Steud., Nomencl. Bot. (ed. 2) 
1: 564. 1840. 
NOTE: nom. inval., as syn. of Eragrostis poaeformis Link. 
= Eragrostis minor Host. 


Eragrostis vulgaris Coss. & Germ., Fl. Env. Paris 2: 641. 
1845, 

NOTE: nom. nov., nom. illeg. superfl.; equals Eragrostis 
minor Host (E. vulgaris Presl ex Steud. Nom. Bot. 
2(1): 564 (1840), as syn. of E. poaeformis Link) 
[Index of Grass Species]. 

REPLACED NAME: Poa eragrostis L. 

BLOCKING NAME: Eragrostis eragrostis (L.) P. Beauv. 

CORRECT NAME: Eragrostis minor Host. 

= Eragrostis minor Host. 


Eragrostis vulgaris subsp. barrelieri (Daveau) R.C.V. Douin, 
Fl. Ill, France 12: 32. 1927-1932. Basionym: 
Eragrostis barrelieri Daveau. 

= Eragrostis barrelieri Daveau. 


Eragrostis vulgaris subsp. megastachya (Koeler) Douin, FI. 
Ill. France 12: 32. 1934. Basionym: Poa megastachya 
Koeler. 

= Eragrostis cilianensis (All.) Vignolo ex Janch, 


Eragrostis vulgaris var. megastachya (Koeler) Coss. & 
Germ., Fl. Env. Paris 2: 641. 1845. Basionym: Poa 
megastachya Koeler. 

= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Eragrostis vulgaris var. microstachya Coss. & Germ., FI. 
Env. Paris 2: 641. 1845, 
NOTE: nom. nov., nom. illeg.  superfl.; Eragrostis 
poaeoides P. Beauv. and Poa eragrostis L. cited. 
REPLACED NAME: Poa eragrostis L. 
BLOCKING NAME: Eragrostis eragrostis (L.) P. Beauv. 
= Eragrostis minor Host. 


Eragrostis vulgaris subsp. poaeoides (P. Beauv. ex Roem. & 
Schult.) R-C.V. Douin, Fl. Ill. France 12: 32. 1927- 
1932. Basionym: Eragrostis poaeoides P. Beauv. ex 
Roem. & Schult. 
= Eragrostis minor Host. 


Eragrostis warmingii Hack., Oesterr. Bot. Z. 52: 305. 1902. 
T-Protol.: Venezuela: in campis prope Las Trincheras, 
Warming s.n. T-Specim.: | of 1. Venezuela: Las 
Trincheras, 1891-1892, .E.B. Warmingii 315 (IT: US- 
79729]. 

NOTE: as Eragrostis "Warmingii" 
= Eragrostis viscosa (Retz.) Trin. 


Eragrostis weberbaueri Pilg., Bot. Jahrb. Syst. 37: 375. 
1906. T-Protol.: Peru: Ancachs: inter Samanco et 
Caraz infra Pampa Romas, in formatione aperta, 
praecipue gramina gerente intermixtis fruticibus et 
Cactaceis, 2300 m, May 1903, Weberbauer 3189 [HT: 
B]. T-Specim.: | of 1. Peru: Ancash: Pampa Roman 
entre Samanco y Caraz, 2300 m, 29 May 1903, 4. 
Weberbauer 3189 [HT: B; IT: BAA-1100 (fragm. ex 
B), US-2767411]. 

NOTE: as Eragrostis "Weberbaueri". 

DST.: Chil, Peru. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Hitchcock, A. S. (1927); Macbride, 
J. F. (1936); Nicora, E. G. (1994); Tovar, O. (1993). 


Eragrostis weigeltiana Bush, Trans. Acad. Sci. St. Louis 13: 
180. 1903. T-Protol.: Surinam, 1827, Weigelt. 
NOTE: nom. altern., cites Poa weigeltiana Rchb. ex Trin., 
but that a nom. inval. (see also Neeragrostis w.). 
= Eragrostis reptans (Michx.) Nees. 


Catalogue of New World Grasses: II. 


115 


Eragrostis yucatana L.H. Harv., Bull. Torrey Bot. Club 
81(5): 406. 1954. T-Protol.: Mexico: Yucatan: in locis 
aridis secus viam prope Progreso, J.R. Swallen 2933 
[HT: US-1537194]. T-Specim.: 1 of 1. Mexico: 
Yucatan: near Progreso, 11 Aug 1932-15 Aug 1932, 
JR. Swallen 2933 (HT: US-1537194]. 
= Eragrostis secundiflora J. Presl. 


Eremochloe S. Watson, Botany Fortieth Parallel 5: 382. 
1871. TYPE: Eremochloe kingii S. Watson. 
NOTE: hom. illeg. 


Eremochloe bigelovii S. Watson, Botany Fortieth Parallel 
382, t. 40, f. 1-9. 1871. T-Protol.: USA: Texas: near El 
Paso, Frontera, Wright 2028. T-Specim.: | of 3. USA: 
Texas: Grassy rocky hills near Frontera, in small 
compact bushes, 4 May 1852, C. Wright 2028 (ST: 
GH; IST:K, US-81934 (fragm.)], field number 277. 2 
of 3. USA: Texas: Mex. Boundary Surv. chiefly in the 
valley of the Rio Grande, below Dofiana, rocky hills 
Frontera, 4 May 1852 (or 4 July 1852), Parry, 
Bigelow, Wright & Schott s.n. [ST: GH; IST:K]. 3 of 
3. USA: Texas: Frontera, Mex. Boundary Surv., 4 May 
1852, Bigelow s.n. [ST: GH; IST: US-81934 (fragm. 
ex GH)]. 

= Blepharidachne bigelovii(S. Watson) Hack. 


Eremochloe kingiiS. Watson, Botany Fortieth Parallel 382, t. 
40, f. 10-16. 1871. T-Protol.: USA: Nevada: Trinity 
Mountains, Watson s.n. T-Specim.: | of 1. USA: 
Nevada: Trinity Mts., May 1868, Watson 1300 [HT: 
GH; IT: US-81943 (fragm.)]. 

= Blepharidachne kingii (S. Watson) Hack. 


Eremochloe thurberi S. Watson, Botany Fortieth Parallel pl 
40, f. 1-9. 1871. T-Protol.: USA: Texas. 
= Blepharidachne bigelovii(S. Watson) Hack. 


Erioneuron Nash, FI. S.E. US. 143, 1327. 1903. 


TYPE: Erioneuron pilosum (Buckley) Nash. 
TRTD. BY: Valdés-Reyna, J. & P. M. Peterson. 
NHT: Munroinae. NO. ACCPTD. SPP.: 3 SUBSP.: 7. 
CNSLTD. LIT.: Valdés-Reyna, J. & S. L. Hatch, (1997); 
Valdés-Reyna, J. & S. L. Hatch, (1995). 


Erioneuron avenaceum (Kunth) Tateoka, Amer. J. Bot. 
48(7): 572. 1961. Basionym: Triodia avenacea Kunth. 

DST.: Arge, Boli, Mexi, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); McVaugh, R. 
(1983); Renvoize, S. A. (1998); Valdés-Reyna, J. & 
S. L. Hatch (1997). 

Dasyochloa avenacea (Kunth) Willd. ex Steud.; 
Erioneuron avenaceum var. grandiflorum (Vasey) 
Gould; Erioneuron grandiflorum (Vasey) Tateoka; 
Erioneuron nealleyi var. grandiflorum (Vasey) 
Beetle; Koeleria avenacea (Kunth)  Spreng.; 
Sieglingia avenacea (Kunth) Kuntze; Sieglingia 
avenacea var. grandiflora (Vasey) L.H. Dewey; 
Sieglingia grandiflora (Vasey) Beal, Sieglingia 
nealleyi (Vasey) L.H. Dewey; Tricuspis avenaceus 
(Kunth) Thurb. ex A. Gray; Tridens avenaceus 
(Kunth) Hitche.; Tridens grandiflora (Vasey) 
Wooton & Standl.; Triodia avenacea Kunth; Triodia 
avenacea var. breviaristata Kurtz,  Triodia 


SYN: 


grandiflora Vasey; Uralepis avenacea (Kunth) 
Kunth; Uralepis avenacea var. viridiflora E. Fourn. 


Erioneuron avenaceum (Kunth) Tateoka var. avenaceum. 
DST.: Mexi, USA. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Beetle, A. A. (1977). 
SYN.: Tridens avenaceus var. avenaceus; Triodia avenacea 
var. avenaced. 


Erioneuron avenaceum var. cabrerae (Caro) E.A. Sanchez, 
Lilloa 36: 115. 1983. Basionym: Dasyochloa 
longiglumis var. cabrerae Caro. 

= Erioneuron avenaceum var. longiglume (Parodi) Anton. 


Erioneuron avenaceum var. grandiflorum (Vasey) Gould, 
Brittonia 26(1): 60. 1974. Basionym: Triodia 
grandiflora Vasey. 

= Erioneuron avenaceum (Kunth) Tateoka. 


Erioneuron avenaceum var. kurtzianum (Parodi) Anton, 
Kurtziana 10: 62. 1977. Basionym: 7ridens avenaceus 
var. kurtziana Parodi. 

DST.: Arge. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anton, A. M. (1977); Valdés-Reyna, 
J. & S.L. Hatch (1997), 

SYN.: | Dasyochloa  kurtziana_ (Parodi) 
avenaceus vat. kurtziana Parodi. 


Caro; Tridens 


Erioneuron avenaceum var. longiaristatum (Kurtz) Beetle, 
Phytologia 49(1): 39. 1981. Basionym: Triodia 
avenacea Var. longearistata Kurtz. 

NOTE: as Erioneuron avenaceum "longearistatum". 
= Erioneuron pilosum var. longearistatum (Kurtz) Anton. 


Erioneuron avenaceum var. longiglume (Parodi) Anton, 
Kurtziana 10: 65. 1977. Basionym: Tridens avenaceus 
var. /ongiglumis Parodi. 

DST.: Arge. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Anton, A. M. (1977); Valdés-Reyna., 

J. & 8. L. Hatch (1997). 

Dasyochloa longiglumis (Parodi) Caro; Dasyochloa 

longiglumis var. cabrerae Caro; Dasyochloa 

longiglumis var. longiglumis, Erioneuron avenaceum 
var. cabrerae (Caro) E.A. Sanchez, Tridens 
avenaceus var. longiglumis Parodi. 


SYN.: 


Erioneuron avenaceum var. nealleyi (Vasey) Gould, Brittonia 
26(1): 60. 1974. Basionym: Triodia nealleyi Vasey. 
= Erioneuron nealleyi (Vasey) Tateoka. 


Erioneuron avenaceum var. pygmaeum (Hack.) Anton, 
Kurtziana 10: 63. 1977. Basionym: Triodia avenacea 
var. pygmaea Hack. 

DST.: Arge, Boli. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anton, A. M. (1977); Valdés-Reyna, 
J. & S. L. Hatch (1997). 

SYN.: Dasyochloa pygmaea (Hack.) Caro; Tridens 
avenaceus var. pygmaea (Hack.) Parodi; Triodia 
avenacea var. pygmaea Hack. 


Erioneuron grandiflorum (Vasey) Tateoka, Amer. J. Bot. 48: 
572. 1961. Basionym: Triodia grandiflora Vasey. 
= Erioneuron avenaceum (Kunth) Tateoka. 


Erioneuron nealleyi (Vasey) Tateoka, Amer. J. Bot. 48(7): 
572. 1961. Basionym: 7riodia nealleyi Vasey. 


116 


Catalogue of New World Grasses: II. 


DST.: Mexi, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Valdés-Reyna, 
J. &S.L. Hatch (1997). 

Erioneuron avenaceum vat. nealleyi (Vasey) Gould; 
Sieglingia nealleyi (Vasey) L.H. Dewey; Tricuspis 
nealleyi (Vasey) A. Heller; Tridens nealleyi (Vasey) 
Wooton & Standl.; 7riodia nealleyi Vasey; Triodia 
nealleyi Vasey. 


SYN:: 


Erioneuron nealleyi var. grandiflorum (Vasey) Beetle, 
Phytologia 27(6): 442. 1974. Basionym: Triodia 
grandiflora Vasey. 

= Erioneuron avenaceum (Kunth) Tateoka. 


Erioneuron pilosum (Buckley) Nash, Fl. S.E. U.S. 144, 
1327. 1903. Basionym: Uralepis pilosa Buckley. 
DST.: Arge, Mexi, USA. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Hickman, J. C. 
(1993); McVaugh, R. (1983); Munz, P. A. (1974); 
Valdés-Reyna, J. & S. L. Hatch (1997). 
Sieglingia acuminata (Benth. ex Vasey) Kuntze; 
Sieglingia pilosa (Buckley) Nash; Tricuspis 
acuminata Munro ex A. Gray; Tricuspis pilosa 
(Buckley) A. Heller; 7ridens pilosus (Buckley) 
Hitche.; 7riodia acuminata Benth. ex Vasey; Triodia 
pilosa (Buckley) Merr.; Uralepis pilosa Buckley. 


SYN:: 


Erioneuron pilosum var. argentinum (Kuntze) Nicora, 
Darwiniana 18: 104. 1973. Basionym: Sieglingia 
argentina Kuntze. 

= Erioneuron pilosum var. longearistatum (Kurtz) Anton. 


Erioneuron pilosum var. aristiglumis (Caro) E.A. Sanchez, 
Lilloa 36: 135. 1983. Basionym: Dasyochloa 
argentina var. aristiglumis Caro. 

= Erioneuron pilosum var. longearistatum (Kurtz) Anton. 


Erioneuron pilosum var. longearistatum (Kurtz) Anton, 
Kurtziana 10: 58. 1977. Basionym: Triodia avenacea 
var. /ongearistata Kurtz. 

DST.: Arge. 

LVL. ACCPTC.:: |. 

ALSO ACCPTD. BY: Anton, A. M. (1977); Nicora, E. G., 
M. E. De Paula, A. M. Faggi, M. del C. Mariano, A. 
M. Miante A., L. R. Parodi, C. A. Petetin, F. A. Roig 
& Z. Rugolo de Agrasar (1978); Valdés-Reyna, J. & 
S. L. Hatch (1997). 

Dasyochloa argentina (Kuntze) Caro; Dasyochloa 
argentina var. argentina; Dasyochloa argentina var. 
aristiglumis Caro; Dasyochloa argentina var. 
parodiana (E.A. Sanchez) Caro; Erioneuron 
avenaceum var. longiaristatum (Kurtz) Beetle; 
Erioneuron pilosum var. argentinum (Kuntze) 
Nicora; Erioneuron pilosum var. aristighumis (Caro) 
E.A. Sanchez; Erioneuron pilosum var. parodianum 
E.A. Sanchez; Sieglingia argentina Kuntze; Tridens 
pilosus var. argentina (Kuntze) Parodi; Triodia 
argentina (Kuntze) K. Schum.; 7riodia avenacea var. 
longearistata Kurtz. 


SYN:: 


Erioneuron pilosum var. mendocinum (Parodi) Nicora, 
Darwiniana 18: 104. 1973. Basionym: 7ridens pilosus 
var. mendocina Parodi. 

DST.: Arge. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anton, A. M. (1977); Valdés-Reyna, 
J. & S.L. Hatch (1997). 

SYN.: Tridens pilosus var. mendocina Parodi. 


Erioneuron pilosum var. parodianum E.A. Sanchez, 

Darwiniana 22(1-3): 171. 1979. T-Protol.: Argentina: 

La Pampa: Dpto. Lihuel-Calel: Sierra Lihuel-Calel, 21 

Dec 1947, A. Krapovickas 3633 [HT: BAB; IT: SI]. T- 

Specim.: 1 of 1. Argentina: La Pampa: Dpto. Lihuel- 

Calel: Sierra Lihuel-Calel, 21 Dec 1947, A. 
Krapovickas 3633 [HT: BAB; IT: CORD, SI]. 

= Erioneuron pilosum var. longearistatum (Kurtz) Anton. 


Erioneuron pilosum (Buckley) Nash var. pilosum. 
DST.: Mexi, USA. 
LVL. ACCPTC.:: 1. 


Erioneuron pulchellum (Kunth) Tateoka, Amer. J. Bot. 48(7): 
572. 1961. Basionym: Triodia pulchella Kunth, 
= Dasyochloa pulchella (Kunth) Willd. ex Rydb. 


Erochloe Raf., Neogenyton 4. 1825. ; type not designated. 
= Eragrostis Wolf. 


Erochloe amabilis Raf. ex B.D. Jacks., Index Kew. 1: 886. 
1893. 
NOTE: nom. inval., as syn. of Eragrostis amabilis (L.) 
Wight & Arn. ex Nees. 
= Eragrostis amabilis (L.) Wight & Arn. ex Nees. 


Erosion Lunell, Amer. Midl. Naturalist 4: 221. 1915, 
NOTE: nom. illeg. superfl. 
= Eragrostis Wolf. 


Erosion ciliare Lunell, Amer. Midl. Naturalist 4: 221. 1915. 
NOTE: basis erroneously attributed to Eragrostis ciliaris 
(All.) Link, rather than E. cilianensis (All.) Link. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Erosion hypnoides (Lam.) Lunell, Amer. Midl. Naturalist 4: 
221. 1915. Basionym: Poa hypnoides Lam. 
= Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb. 


Erucaria Cerv., Naturaleza (Mexico City) 1: 347. 1870. 
TYPE: Erucaria glandulosa Cerv. 
NOTE: hom. illeg. 
= Bouteloua Lag. 


Erucaria glandulosa Cerv., Naturaleza (Mexico City) |: 347. 
1870. T-Protol.: Mexico: in collibus de Guadalupe, et 
de Moctezuma prope Mexicum, floret Septembri, 
Anon, T-Specim.: 1 of 1. Mexico: Guadelupe et 
Moctezuma, lV. Cervantes s.n. [ST: possibly in BR, F, 
FI, MEXU, NY, OXF]. 

= Bouteloua hirsuta var. glandulosa (Cerv.) Gould. 


Erucaria hirsuta Cerv., Naturaleza (Mexico City) 1: 348. 
1870. T-Protol.: Mexico: In collibus de Guadalupe, et 
de Moctezuma prope Mexicum. 

= Bouteloua hirsuta var. glandulosa (Cery.) Gould. 


Erucaria lutescens Cerv., Naturaleza (Mexico City) 1: 349. 
1870. T-Specim.: | of 1. Mexico [T]. 
= Bouteloua simplex Lag. 


Catalogue of New World Grasses: II. 


117 


Erucaria tetrastachya Cery., Naturaleza (Mexico City) 1: 
349. 1870. T-Protol.: Mexico: In collibus de 
Guadalupe, et de Moctezuma prope Mexicum. 

= Bouteloua barbata var. barbata. 


Eufournia Reeder, Brittonia 19: 244. Jul-Sep 1967. 
NOTE: nom. nov., nom. superfl. pro Sohnsia. 
BLOCKING NAME: Calamochloe Rchb. 
= Sohnsia Airy Shaw. 


Eufournia filifolia (E. Fourn.) Reeder, Brittonia 19: 244. 
1967. Basionym: Calamochloa filifolia E. Fourn. 
= Sohnsia filifolia (E. Fourn.) Airy Shaw. 


Eustachys Desv., Nouv. Bull. Sci. Soc. Philom. Paris 2: 
188. 1810. TYPE: Eustachys petraea (Sw.) Desv. 
TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 12 SUBSP.: 2. 
CNSLTD. LIT.: Molina, A. M., (1996). 
SYN.: Chloroides Fisch. ex Regel; Langsdorffia Regel; 
Schultesia Spreng. 


Eustachys argentina (Hack. ex Stuck.) Herter, Estud. Bot. 
Reg. Uruguay 1: 86, f. 341. 1941. Basionym: Chloris 
distichophylla var. argentina Hack. ex Stuck. 

= Eustachys retusa (Lag.) Kunth. 


Eustachys argentina (Hack. ex Stuck.) Herter, Revista 
Sudamer. Bot. 7(6-8): 196. 1943. Basionym: Chloris 
distichophylla var. argentina Hack. ex Stuck. 

NOTE: hom. illeg. 

E. NAME: Eustachys argentinensis (Hack. ex Stuck.) 
Herter. 

= Eustachys retusa (Lag.) Kunth. 


Eustachys argentinensis (Hack. ex Stuck.) Herter, Revista 
Sudamer. Bot. 6(5-6): 146. 1940. Basionym: Chloris 
distichophylla var. argentina Hack. ex Stuck. 

= Eustachys retusa (Lag.) Kunth. 


Eustachys argentinensis (Lillo & L. Parodi) Herter, Revista 
Sudamer. Bot. 6: 146. 1940. 
NOTE: erroneous citation, cf. Gray Herb. Card Cat. 
= Eustachys retusa (Lag.) Kunth, 


Eustachys bahiensis (Steud.) Herter, Estud. Bot. Reg. 
Uruguay 14(1): 85, f. 339. 1941 [1942]. Basionym: 
Chloris bahiensis Steud. 

NOTE: basionym not explicitly cited, only the basionym 
author; cf. Gray Herb. Card Cat.; Herter, Rev. 
Sudamer. Bot. 7: 196 (1943). 

= Eustachys paspaloides subsp. caribaea (Spreng.) 
Nowack. 


Eustachys bahiensis (Steud.) Herter var. bahiensis. 
= Eustachys paspaloides subsp. caribaea (Spreng.) 
Nowack. 


Eustachys bahiensis var. robusta A.M. Molina, Candollea 
51(1): 260, f. 7A-I, 8D. 1996. T-Protol.: Brasil: 
Parana:4: Mun. Lapa: Engl. Bey, do campo, 17 Jun 
1951, G. Hatschbach 2294 [HT: BAA; IT: MBM]. 

NOTE: Presumably belongs in E. paspaloides, but comb. 
not available. 

DST.: Braz. 

LVL. ACCPTC:: 2. 

ALSO ACCPTD. BY: Molina, A. M. (1996). 


Eustachys brevipila (Roseng. & Izag.) Caro & E.A. 
Sanchez, Kurtziana 6: 124. 1971. Basionym: Chloris 
brevipila Roseng. & Izag. 

DST.: Arge, Braz, Urug. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Molina, A. M. (1996). 
SYN.: Chloris brevipila Roseng. & Izag. 


Eustachys calvescens (Hack.) Caro & E.A. Sanchez, 
Kurtziana 6: 128. 1971. Basionym: Chloris calvescens 
Hack. 

DST.: Arge, Braz, Para. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Molina, A. M. (1996); Renvoize, S. 
A. (1988). 

SYN.: Chloris calvescens Hack. 


Eustachys capensis (Houtt.) Chiov., Webbia 8: 115. 1951. 
Basionym: Andropogon capensis Houtt. 
= Eustachys paspaloides subsp. paspaloides. 


Eustachys caribaea (Spreng.) Herter, Revista Sudamer. Bot. 
6: 147. 1940. Basionym: Chloris caribaea Spreng. 
NOTE: cf. Gray Herb. Card Cat, "caribaeus", Accd. to 
Chase & Niles, Index Grass Sp. 
= Eustachys paspaloides subsp. caribaea (Spreng.) 
Nowack. 


Eustachys confertifolia (Trin.) Kunth, Revis. Gramin. 1: 88. 
1829. Basionym: Chloris confertifolia Trin. 
= Eustachys distichophylla (Lag.) Nees. 


Eustachys distichophylla (Lag.) Nees, Fl. Bras. Enum. PI. 2: 
418. 1829. Basionym: Chloris distichophylla Lag. 

DST.: Arge, Boli, Braz, Chil, Para, Peru, USA, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Brako, L. & 
J. L. Zarucchi (1993); Filgueiras, T. S. (1999); Jones, 
S. D. & J. K. Wipff (1992); Killeen, T. J. (1990); 
Marticorena, C. & M. Quezada (1985); Molina, A. 
M. (1996); Renvoize, S. A. (1984); Renvoize, S. A. 
(1988); Renvoize, S. A. (1998); Tovar, O. (1993); 
Wunderlin, R. P. (1998). 

Chloris acuminata Trin.; Chloris confertifolia Trin.; 
Chloris distichophylla Lag.; Chloris distichophylla 
var. acuminata (Trin.) Hack.; Chloris distichophylla 
var. argentina Hack. ex  Stuck.; Chloris 
distichophylla var. genuina Hack.; ~~ Chloris 
dolichostachya Lag.; Chloris fasciculata Schrad. ex 


SYN.: 


Schult.; Chloris paytensis Steud.; Eustachys 
confertifolia (Trin.) Kunth; Paspalum superbum 
Spreng. 


Eustachys floridana Chapm., Fl. South. U.S. 557. 1860. T- 
Protol.: USA: Dry Pine barrens, middle Florida. 

DST.: USA. 

LVL. ACCPTC.: |. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Anonymous 
(1986); Jones, S. D. & J. K. Wipff (1992): 
Wunderlin, R. P. (1998). 

SYN.: Chloris floridana (Chapm.) Alph. Wood. 


Eustachys gayana Mundy, Rhodesian Agric. J. 14: 142. 1922. 
NOTE: presumably based on Chloris gayana but basionym 
not cited. 
= Chloris gayana Kunth. 


118 Catalogue of New World Grasses: II. 


Eustachys glabrescens (Hack.) Caro & E.A. Sanchez, 
Kurtziana 6: 120. 1971. Basionym: Chloris bahiensis 
fo. glabrescens Hack. 

= Eustachys paspaloides subsp. caribaea (Spreng.) 
Nowack. 


Eustachys glauca Chapm., Fl. South. U.S. 557. 1860. T- 
Protol.: Brackish marshes, West Florida, Chapman s.n. 
[Aug-Sep]. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Anonymous 
(1986); Jones, S. D. & J. K. Wipff (1992); Molina, A. 
M. (1996); Wunderlin, R. P. (1998). 

SYN.: Chloris glauca (Chapm.) Alph. Wood. 


Eustachys neglecta (Nash) Nash, Bull. Torrey Bot. Club 25: 
450. 1898. Basionym: Chloris neglecta Nash. 

DST.: USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Jones, S. D. & J. 
K. Wipff (1992); ref. missing (1998); Wunderlin, R. 
P. (1998). 

SYN.: Chloris neglecta Nash. 


Eustachys paranensis A.M. Molina, Candollea 51(1): 234, f. 
3A-J, 8A. 1996. T-Protol.: Brasil: Parana: Ponta 
Grossa, rare on campo, Fazenda de Criacao, 4 Feb 
1946, JR. Swallen 8459 [HT: US-1960543; IT: PEL]. 

DST.: Braz. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Molina, A. M. (1996), 


Eustachys paspaloides (Vahl) Lanza & Mattei, Boll. Reale 
Orto Bot. Giardino Colon. Palermo 9: 56. 1910. 
Basionym: Cynosurus paspaloides Vahl. 

DST.: Arge, Braz, Cari, Para. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Gibbs 
Russell, G. E, W. G. Welman, E. Reitief, K. L. 
Immelman, G. Germishuizen, B. J. Pienaar, M. van 
Wyk, and A. Nicholas (PRECIS) (1987); Phillips, S. 
(1995). 

SYN.: Chloris equitans Trin.; Chloris paspaloides Hochst.; 
Chloris petraea Thunb.; Cynosurus  paspaloides 
Vahl. 


Eustachys paspaloides subsp. caribaea (Spreng.) Nowack, 
Bull. Mus. Natl. Hist. Nat., B, Adansonia Ser. 4, 17(1- 
2): 57. 1995. Basionym: Chloris caribaea Spreng. 

NOTE: Type trom Caribbean, Guadaloupe, may be in error, 
as the taxon has not been found there since. 

DST.: Arge, Braz, Para. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Nowack, R. (1995), 

SYN.: Chloris bahiensis fo. glabrescens Hack.; Chloris 
bahiensis Steud.; Chloris capensis var. bahiensis 
(Steud.) Parodi; Chloris capensis var. glabrescens 
(Hack.) Parodi; Chloris caribaea Spreng.; Chloris 
glabrescens (Hack.) Roseng. & Izag.; Eustachys 
bahiensis (Steud.) Herter; Eustachys bahiensis var. 
bahiensis, Eustachys caribaea (Spreng.) Herter; 
Eustachys glabrescens (Hack.) Caro & E.A. Sanchez. 


Eustachys petraea (Sw.) Desv., Nouv. Bull. Sci. Soc. 
Philom. Paris 2: 189. 1810. Basionym: Chloris petraea 
Sw. 


DST.: Arge, Beli, Boli, Braz, Cari, CoRi, Hond, Mexi, Nica, 
Pana, USA, Urug. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Anonymous 
(1986); Balick, M. J.. M. H. Nee & D. E. Atha 
(2000); Beetle, A. A. (1977); Catasus Guerra, L. 
(1997); Cowan, C. P. (1983); Davidse, G.. M. Sousa 
S. & A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Fosberg, F. R. & M.-H. Sachet (1982); 
Gould, F. W (1979); Jones, S. D. & J. K. Wipff 
(1992); Liogier, H. & L. Martorell (1982); Molina, 
A. M. (1996); Pohl, R. W. (1980); Pohl, R. W. 
(1994); Renvoize, S. A. (1998); Sousa S.. M. & E. F. 
Cabrera C. (1983); Wunderlin, R. P. (1998). 

SYN.: Chloris petraea Sw.; Chloris septentrionalis Mill. 
Hal.; Chloris swartziana Doll; Chloris swartzii Mill. 
Hal.; Schultesia petraea (Sw.) Spreng. 


Eustachys polystachya (Lag.) Kunth, Revis. Gramin. |: 88. 
1829. Basionym: Chloris polystachya Lag. 
= Chloris submutica Kunth. 


Eustachys retusa (Lag.) Kunth, Revis. Gramin. 1: 88. 1829. 
Basionym: Chloris retusa Lag. 

DST.: Arge, Boli, Braz, Para, USA, Urug. 

LVL. ACCPTC.:: I. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Anonymous 
(1986); Jones, S. D. & J. K. Wipff (1992); Molina, A. 
M. (1996); Nicora, E. G., M. E. De Paula, A. M. 
Faggi, M. del C. Mariano, A. M. Miante A., L. R. 
Parodi, C. A. Petetin, F. A. Roig & Z. Rugolo de 
Agrasar (1978); Renvoize, S. A. (1988); Wunderlin, 
R. P. (1998). 

SYN.: Chloris argentina (Hack. ex Stuck.) Lillo & Parodi; 
Chloris argentinensis (Hack. ex Stuck.) Lillo & 
Parodi; Chloris distichophylla var. argentina Hack. 
ex Stuck.; Chloris retusa Lag.; Eustachys argentina 
(Hack. ex Stuck.) Herter; Eustachys argentina (Hack. 
ex Stuck.) Herter; Eustachys argentinensis (Hack. ex 
Stuck.) Herter; Eustachys argentinensis (Lillo & L. 
Parodi) Herter. 


Eustachys submutica (Kunth) Roem. & Schult., Syst. Veg. 2: 
614. 1817. Basionym: Chloris submutica Kunth. 
= Chloris submutica Kunth. 


Eustachys swalleniana A.M. Molina, Candollea 51(1): 232, 
f. 2A-H, 8A. 1996. T-Protol.: Brasil: Rio Grande do 
Sul: Pelotas, Instituto Agronémico do Sul, 6 Jun 1945, 
J. R. Swallen 7287 (HT: PEL}. 
DST.: Braz, Urug. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Molina, A. M. (1996). 


Eustachys ulignosa (Hack.) Herter, Revista Sudamer. Bot. 
6(5-6): 147. 1940. Basionym: Chloris uliginosa Hack. 

NOTE: spelled "uliginosus", Accd. to Chase & Niles, Index 
Grass Sp. 

DST.: Arge, Boli, Braz, Para, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Filgueiras, 
T. S. (1999); Molina, A. M. (1996): Renvoize, S. A. 
(1988). 

SYN.: Chloris dusenii Ekman; Chloris uliginosa Hack. 


Eutriana Trin., Fund. Agrost. 161. 1820. TYPE: Eutriana 
curtipendula (Michx.) Trin. 


Catalogue of New World Grasses: II. 


119 


= Bouteloua Lag. 


Eutriana affinis Hook. f., Trans. Linn. Soc. London 20: 174. 
1847. T-Specim.: | of 2. North America: Missouri: St. 
Louis, Schweinitz s.n. [ST: K]. 2 of 2. USA: Texas, 
Drummond s.n. |ST: K; IST: E?, US-1063204 
(fragm.)]. 

= Bouteloua curtipendula (Michx.) Torr. 


Eutriana antillarum (Poir.) Steud., Syn. Pl. Glumac. 1: 217. 
1854. Basionym: Aristida antillarum Poir. 
= Bouteloua americana (L.) Scribn. 


Eutriana aristidoides (Kunth) Trin., Gram. Unifl. Sesquifl. 
242. 1824. Basionym: Dinebra aristidoides Kunth. 
= Bouteloua aristidoides (Kunth) Griseb. 


Eutriana barbata (Lag.) Kunth, Revis. Gramin. 1: 96. 1829. 
Basionym: Bouteloua barbata Lag. 
= Bouteloua barbata Lag. 


Eutriana bromoides (Kunth) Trin., Fund. Agrost. 161, 162. 
1820. Basionym: Dinebra bromoides Kunth. 
= Bouteloua radicosa (E. Fourn.) Griffiths. 


Eutriana bromoides (Lag.) Trin., Gram. Unifl. Sesquifl. 241. 
1824. Basionym: Bouteloua bromoides Lag. 
NOTE: hom. illeg. 
E. NAME: Eutriana bromoides (Kunth) Trin. 
= Bouteloua repens (Kunth) Scribn. 


Eutriana bromoides (Kunth) Kunth, Revis. Gramin. 1: 95. 
1829. Basionym: Dinebra bromoides Kunth. 
E. NAME: Eutriana bromoides (Kunth) Trin. 
= Bouteloua radicosa (E. Fourn.) Griffiths. 


Eutriana chloridea (J. Presl) Kunth, Enum. PI. 1: 280. 1833. 
Basionym: Dinebra chloridea J. Presl. 
NOTE: also Revis. Gram. 3: Suppl. 1834 p. (as in 1835 ed.) 
XXIHI. 
= Enteropogon chlorideus (J. Presl) Clayton. 


Eutriana cristata Trin., Gram. Unifl. Sesquifl. 241. 1824. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Atheropogon chondrosiodes (Kunth) 
Roem. & Schult. 
= Bouteloua chondrosioides (Kunth) Benth. ex S. Watson. 


Eutriana curtipendula (Michx.) Trin., Fund. Agrost. 161. 
1820. Basionym: Chloris curtipendula Michx. 
= Bouteloua curtipendula (Michx.) Torr. 


Eutriana gracilis (Kunth) Trin., Gram. Unifl. Sesquifl. 240. 
1824. Basionym: Chondrosum gracile Kunth. 
= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Eutriana gracilis Hook. f., Trans. Linn. Soc. London 20: 175. 


1847. T-Protol.: Argentina: Tucman, Tweedie s.n. T- 
Specim.: 1 of 1. Argentina: Tucuman, Tweedie s.n. 
[HT: K; IT: US (fragm.)]. 

NOTE: hom. illeg. 

E. NAME: Eutriana gracilis (Kunth) Trin. 

= Bouteloua disticha (Kunth) Benth. 


Eutriana heterostega Trin., Gram. Unifl. Sesquifl. 242. 1824. 
REPLACED NAME: Heterosteca juncifolia Desv. 
BLOCKING NAME: Bouteloua juncifolia Lag. 
= Bouteloua repens (Kunth) Scribn. 


Eutriana hirsuta (J. Presl) Kunth, Enum, Pl. 1: 280. 1833. 


Basionym: Dinebra hirsuta J. Presl. 


NOTE: also Revis. Gram. 3: Suppl. 1834 p. (as in 1835 ed.) 
XXIII. 
= Bouteloua aristidoides (Kunth) Griseb. 


Eutriana hirta (Kunth) Trin., Gram. Unifl. Sesquifl. 240. 
1824. Basionym: Chondrosum hirtum Kunth. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Bouteloua hirsuta Lag. 
= Bouteloua hirsuta Lag. 


Eutriana humilis (P. Beauv. ex Kunth) Trin., Gram. Unifl. 
Sesquifl. 239. 1824. Basionym: Chondrosum humile P. 
Beauv. ex Kunth. 

= Bouteloua simplex Lag. 


Eutriana juncea (Desv. ex P. Beauv.) Trin., Gram. Unifl. 
Sesquifl. 238. 1824. Basionym: Triathera juncea 
Desv. ex P. Beauv. 

= Bouteloua juncea (Desy. ex P. Beauv.) Hitche. 


Eutriana juncifolia (Desv.) Kunth, Revis. Gramin. 1: 95. 
1829. Basionym: Heterosteca juncifolia Desv. 
= Bouteloua repens (Kunth) Scribn. 


Eutriana lagascae Kunth, Revis. Gramin, 1: 95. 1829. 
NOTE: nom. nov., Type not found, unidentifiable, Accd. to 
Chase & Niles, Index Grass Sp. 
REPLACED NAME: Bouteloua juncifolia Lag. 
BLOCKING NAME: Eutriana juncifolia (Desv.) Kunth. 
= Bouteloua juncifolia Lag. 


Eutriana ledebourii Trin., Gram. Unifl. Sesquifl. 238. 1824. 
T-Protol.: Santo Domingo, Ledebourii s.n. [HT: Herb. 
Rudolphii]. 

= Bouteloua juncea (Desv. ex P. Beauv.) Hitche. 


Eutriana mucronata F. Aresch., Pl. Itin. Eugenia 3: 118. 
1910. T-Protol.: Ecuador: In insula Puna, Andersson 
sn. T-Specim.: | of 1. 00 xxx 1852, 23, Andersson 
[HT: S]. 

= Bouteloua disticha (Kunth) Benth. 


Eutriana multiseta Nees, Fl. Bras. Enum. Pl. 2: 413. 1829. T- 
Protol.: Ad Monte Video, Sellow s.n. [HT: B]. T- 
Specim.: | of 1. Uruguay: Montevideo, F. Sedlo s.n. 
[HT: B; IT: LE, MO-2108848, US-72694 (fragm. ex 
B, Willd. hb.)]. 

= Bouteloua megapotamica (Spreng.) Kuntze. 


Eutriana oligostachya (Nutt.) Kunth, Revis. Gramin. 1: 96. 
1829. Basionym: Atheropogon oligostachyus Nutt. 
= Bouteloua gracilis (Kunth) Lag. ex Griffiths. 


Eutriana pilosa Hook. f., Trans. Linn. Soc. London 20: 173. 
1847. T-Specim.: | of 2. Galapagos Islands: Abelmarle 
Island, Macrae s.n. [LT: ?; IST: US (fragm.)], LT 
designated by Porter, Bot. J. Linn. Soc. 81: 97 (1980). 
2 of 2. Ecuador: Galapago Islands: Albemarle Island, 
Macrae s.n. [LT: CGE]. 

= Bouteloua disticha (Kunth) Benth. 


Eutriana polyodon Trin., Gram. Unifl. Sesquifl. 242. 1824. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Polyodon distichum Kunth. 
= Bouteloua disticha (Kunth) Benth. 


Eutriana repens (Kunth) Trin., Gram. Unifl. Sesquifl. 241. 
1824. Basionym: Dinebra repens Kunth. 
= Bouteloua repens (Kunth) Scribn. 


120 


Catalogue of New World Grasses: II. 


Eutriana tenuis (P. Beauv. ex Kunth) Trin., Gram. Unifl. 
Sesquifl. 240. 1824. Basionym: Chondrosum tenue P. 
Beauv. ex Kunth. 

= Bouteloua simplex Lag. 


Eutriana triaena Trin., Gram. Unifl. Sesquifl. 239. 1824. T- 
Specim.: | of 1. Mexico, Humboldt & Bonpland s.n. 
[HT: P]. 
NOTE: nom. illeg.; based on Triaena racemosa Kunth 
[legit.]. 
CORRECT NAME: 4theropogon triaena Trin. ex Spreng. 
= Bouteloua triaena (Trin. ex Spreng.) Scribn. 


Eutriana sect. Aristidium Endl., Gen. Pl. 94. 1836. TYPE: 
Eutriana aristidoides (Kunth) Trin. 
= Bouteloua Lag. 


Eutriana sect. Triplathera Endl., Gen. Pl. 94. 1836. TYPE: 
Eutriana multiseta Nees. 
= Bouteloua Lag. 


Exagrostis Steud., Nomencl. Bot. (ed. 2) 1: 622. 1840. 
NOTE: nom. nud. 
= Eragrostis Wolf. 


Festuca aquatica Bosc ex Roem. & Schult., Syst. Veg. 2: 
615. 1817. 
NOTE: nom. inval., as syn. of Diplachne fascicularis P. 
Beauv. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Festuca brevifolia Muhl., Cat. Pl. Amer. Sept. 13. 1813. T- 
Specim.: | of 1. USA: Delaware, Georgia and New 
York, no date, Coll. Unkn. [OM: US-1535782]. 

NOTE: nom. nud. 
CORRECT NAME: Festuca brevifolia Muhl. 
= Triplasis purpurea (Walter) Chapm. 


Festuca brevifolia Muhl., Deser. Gram. 167. 1817. T-Protol.: 
USA: Habiata in Delaware, Georgia et N. Eboraco. T- 
Specim.: | of 1. no date, Coll, Unkn. [ST: IST: US- 
1535782]. 

= Triplasis purpurea (Walter) Chapm. 


Festuca caroliniana Steud., Syn. P|. Glumac. 1: 312. 1854. 
T-Protol.: USA: Carolina, Bose s.n. T-Specim.: | of 1. 
USA: Carolina, Bosc s.n. [HT: herb. Caen]. 
= Tridens carolinianus (Steud.) Henrard. 


Festuca clandestina Muhl., Cat. Pl. Amer. Sept. 13. 1813. 
NOTE: nom. nud. 
CORRECT NAME: Festuca clandestina Muhl. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Festuca clandestina Muhl., Descr. Gram. 162. 1817. T- 
Protol.: USA: New York: N. Eboraco. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Festuca distichophylla Michx., Fl. Bor.-Amer. 1: 67. 1803. 
T-Protol.: USA: Hab. in maritimis Carolinae. T- 
Specim.: | of 1. USA: in maritimis Carolinae, 
Michaux s.n. [HT: P, US-2851254 Photo ex P]. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Festuca distichophylla (Labill.) Hook. f., Fl. Tasman. 2: 127. 
1858. Basionym: Uniola distichophylla Labill. 
NOTE: hom. illeg. 
E. NAME: Festuca distichophylla Michx. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Festuca domingensis (Jacq.) Lam., Tabl. Encycl. 1: 189. 
1791. Basionym: Cynosurus domingensis Jacq. 
= Leptochloa virgata (L.) P. Beauv. 


Festuca fascicularis Lam., Tabl. Encycl. 1: 189. 1791. T- 
Protol.: Ex Amer. merid., D. Richard. s.n. T-Specim.: 
| of 1. South America, D. Richard s.n. (HT: P; IT: US- 
2875408, US-2875408a (fragm. ex P)]. 

= Leptochloa fusea subsp. fascicularis (Lam.) N. Snow. 


Festuca filiformis Lam., Tabl. Encycl. 1: 191. 1791. T- 
Protol.: Ex. Amer. Merid.[Richard s.n.]. T-Specim.: | 
of 1. South America, L. Richard s.n. [HT: P-LA]. 

NOTE: hom. illeg. 
E. NAME: Festuca filiformis Pourret. 
= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Festuca flava (L.) F. Muell., Select Pl. 87. 1876. Basionym: 
Poa flava L. 
= Tridens flavus (L.) Hitche. 


Festuca fournieriana Hemsl., Biol. Cent.-Amer., Bot. 3(20): 
581. 1885. T-Protol.: Mexico. 
NOTE: nom. nud., the intended replaced name, Gouinia 
polygama Fourn., not published until 1886. 
= Gouinia virgata (J. Presl) Scribn. 


Festuca fusca L., Syst. Nat. (ed. 10) 2: 876. 1759. T-Specim.: 
1 of 1. Palestine, F. Hasselquist s.n. [LT: LINN- 
92.21], LT designated by Phillips, Fl. Trop. E. Afr., 
Gram (2): 281 (1974). 

= Leptochloa fusca (L.) Kunth. 


Festuca laxiflora A. Rich., Hist. Fis. Cuba, Bot. 11: 318. 
1850. T-Protol.: Cuba: Crescit in pratis non procul ab 
urbe Havanae, Ramon de la Sagra s.n. T-Specim.: | of 
1. Cuba: Habana, de la Sagra s.n. [HT: P; IT: BM]. 

= Gouinia virgata (J. Presl) Scribn. 


Festuca macrostachya Torr. & A. Gray, Pacif. Railr. Rep. 
2(4): 177. 1855. T-Protol.: USA: Texas: Pecos, 
Staminate specimen. T-Specim.: | of 1. [USA]: On the 
Pecos viseo, Capt. Pope [OM: NY (US photo)]. 

NOTE: nom. nud. 
= Scleropogon brevifolius Phil. 


Festuca multiflora Walter, Fl. Carol. 81. 1788. T-Protol.: 
USA: S. Carolina. 
= Distichlis spicata (L.) Greene. 


Festuca obtusiflora Willd. ex Spreng., Syst. Veg. 1: 356. 
1825. T-Protol.: Mexico. 
= Leptochloa dubia (Kunth) Nees. 


Festuca pilosa Nees, Fl. Bras. Enum. P|. 473. 1829. 
NOTE: hom. illeg. 
E. NAME: Festuca pilosa Hall. f. 
= Steirachne diandra Ekman. 


Festuca polystachya Michx., Fl. Bor.-Amer. 1: 66. 1803. T- 
Protol.: USA: Illinoensibus: in arvis. T-Specim.: 1 of 
1. USA: Illinois, Michaux s.n. [HT: P]. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Festuca procumbens Muhl., Descr. Gram. 160. 1817. T- 
Protol.: USA: Carolina. T-Specim.: | of 1. USA: 
Carolinas, Muhlenberg 235 (HT: PH]. 

REPLACED NAME: Festuca procumbens Muhl. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Catalogue of New World Grasses: II. 


121 


Festuca prostrata Muhl. ex Scribn. & Merr., Circ. Div. 


Agrostol. U.S.D.A. 27: 5. 1900. T-Protol.: USA: 
Carolina. 

NOTE: nom. inval., as syn. of Festuca prostrata Muhl. 

= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Festuca purpurea Schreb. ex Steud., Nomencl. Bot. (ed. 2) 1: 


632. 1840. 
NOTE: nom. inval., as syn. of Uralepis cuprea Kunth. 
= Tridens flavus (L.) Hitche. 


Festuca purpurea (Walter) F. Muell., Select Pl. 88. 1876. 
Basionym: Aira purpurea Walter. 
= Triplasis purpurea (Walter) Chapm. 


Festuca quadridens Poir., Encycl. 2: 640. 1812. T-Protol.: 


USA: Cett plante croit a la Caroline, M. Bosc. 
= Tridens flavus (L.) Hitche. 


Festuca quadridens Bosc ex Trin., Mem. Acad. Imp. Sci. 
Saint-Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. 
Nat. 4,2(1): 76. 1836. 
NOTE: nom. inval., as syn. of Eragrostis tricuspis Trin. 
= Tridens flavus (L.) Hitche. 


Festuca quitensis Willd. ex Nees, Fl. Bras. Enum. PI. 473. 
1829. T-Protol.: Brazil. 
NOTE: nom. inval., as syn. of F. pilosa Nees. 
E. NAME: Festuca quitensis Spreng. 
= Steirachne diandra Ekman. 


Festuca reptatrix L., Sp. Pl. (ed. 2) 1: 108. 1762. T-Specim.: 
1 of 1. [LT: LINN-92.20 (left-hand specimen)], LT 
designated by Snow in Cafferty et al., Taxon 49(2): 
250 (2000). 

NOTE: as nom. nud. in Fl. Palaest. 13 (1756). 
= Leptochloa fusca (L.) Kunth. 


Festuca texana Steud., Syn. Pl. Glumac. 1: 310. 1854. T- 
Protol.: USA: Texas, Drummond 387. T-Specim.: | of 
1. USA: Texas: San Felipe, Drummond 387 [HT: P; 
IT: K, US-1445735]. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Festuca thouini Steud., Syn. Pl. Glumac. 1: 311. 1854. T- 
Protol.: West Indies: Ins. Antillae in Domingo, see 
note Index of Grass Species vol. 2. p. 181. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Festuca triticea Lam. ex Kunth, Enum. PI. 1: 325. 1833. 
NOTE: nom. inval., as syn. of Poa michauxil Kunth; 
probably error for Festuca triticoides Lam. 
= Distichlis spicata (L.) Greene. 


Festuca triticoides Lam., Tabl. Encycl. 1: 191. 1791. T- 
Protol.: USA: E. Carolina, Fraser. 


= Distichlis spicata (L.) Greene. 


Festuca virgata (L.) Lam., Tabl. Encycl. 1: 189. 1791. 
Basionym: Cynosurus virgatus L. 
= Leptochloa virgata (L.) P. Beauv. 


Fibichia Koeler, Descr. Gram. 308. 1802. TYPE: Fibichia 
umbellata Koeler; nom. illeg. superfl. Panicum 
dactylon L. 

= Cynodon Rich. 


Fibichia dactylon (L.) Beck, Wiss. Mitt. Bosnien & 
Herzegovina 9: 436. 1904. Basionym: Panicum 
dactylon L. 

= Cynodon dactylon (L.) Pers. 


Fibichia umbellata Koeler, Descr. Gram. 302. 1802. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Panicum dactylon L. 
= Cynodon dactylon (L.) Pers. 


Fingerhuthia Nees, Sem. Hort. Bot. Hamburg. 1834: 
7. 1834. TYPE: Fingerhuthia africana Nees ex Lehm. 
TRTD. BY: Peterson, P. M. 
NHT: Uniolinae. NO. ACCPTD. SPP.: 1. 
SYN.: Lasiotrichos Lehm. 


Fingerhuthia afghanica Boiss., Fl. Orient. 5: 569. 1884. 
REPLACED NAME: Fingerhuthia africana Lehman ex 
Aitch. 
BLOCKING NAME: Fingerhuthia africana Nees ex Lehm. 
= Fingerhuthia africana Nees ex Lehm. 


Fingerhuthia africana Nees ex Lehm., Sem. Hort. Bot. 
Hamburg. 1834: 7. 1834. T-Protol.: Habitat in Africa 
australi in vicinia fluminis Guaritz, Ecklon s.n. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Bor, N. L. (1960); Gibbs Russell, G. 
E, W. G. Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987). 

SYN.: Fingerhuthia afghanica Boiss. 


Fourniera Scribn., Bull. Div. Agrostol., U.S.D.A. 4: 7. 1897. 
TYPE: Fourniera mexicana Scribn. 
NOTE: hom, illeg. 
= Bouteloua Lag. 


Fourniera mexicana Scribn., Bull. Div. Agrostol., U.S.D.A. 
4: 8. 1897. T-Specim.: | of 2. Mexico: Acapulco, 
growing in loose gravelly soil, Oct 1894, Palmer 43 
[LT: US-81932, US-81933, US-494824, US-740915, 
US-740916, US-822265; ILT: BM, MO], LT 
designted by Pohl & Davidse, Fl. Mesoamericana 6: 
297 (1994). 2 of 2. Mexico: Acapulco, growing in 
loose, gravelly soil, Oct 1894, Palmer 41 [ST: MO, 
US]. 

= Bouteloua mexicana (Scribn.) Columbus. 


Glyceria brevifolia (Muhl.) Schult., Mant. 2: 387. 1824. 
Basionym: Festuca brevifolia Muhl. 
NOTE: as G. ? brevifloia. 
= Triplasis purpurea (Walter) Chapm. 


Glyceria micrantha Steud., Syn. Pl. Glumac. 1: 287. 1854. T- 
Specim.: | of 1. Ethiopia: Tigray, Scholoda, 27 Oct 
1837, G.H.W. Schimper 406 (IT: WAG]. 
NOTE: nom. illeg. superfl., cites Poa sporoboloides A. Rich 
in syn. 
= Eragrostis japonica (Thunb.) Trin. 


Glyceria stricta Buckley, Proc. Acad. Nat. Sci. Philadelphia 
14: 95-96. 1862. T-Protol.: USA: middle Texas, 
diseased. T-Specim.: | of 1. USA: Texas, Buckley s.n. 
[LT: PH; IT: US (fragm. & photostat ex PH)], LT 
designated by Hitchcock, Man. Grass. U.S. 958 
(1935), but without citing a specific sheet in a specific 
herbarium. 

NOTE: hom. illeg. 
E. NAME: Giveeria stricta Hook. f. 
= Sporobolus compositus (Poir.) Merr. 


Gossweilerochloa Renvoize, Kew Bull. 33(3): 525. 1979. 


122 


Catalogue of New World Grasses: II. 


= Tridens Roem. & Schult. 


Gouinia E. Fourn. ex Benth. & Hook. f., Gen. Pl. 3: 
1178. 1883. TYPE: Gouinia polygama E. Fourn.; LT 
designated by Swallen, Amer. J. Bot. 22: 32 (1935). 

TRTD. BY: Peterson, P. M. 

NHT: Cynodonteae. NO. ACCPTD. SPP.: 10 SUBSP.: 6. 
CNSLTD. LIT.: Swallen, J. R., (1935); Ortiz, J. J., (1993). 
SYN.: Hackelia Vasey ex Beal; Pogochloa S. Moore. 


Gouinia barbata (Hack.) Swallen, Amer. J. Bot. 22: 37. 

1935. Basionym: Diplachne barbata Hack. 

DST.: Braz. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Nicora, E. G. (1995); Swallen, J. R. 
(1935). 

SYN.: Diplachne barbata Hack.; Schenckochloa barbata 
(Hack.) J.J. Ortiz. 


Gouinia brandegei (Vasey) Hitche., U.S.D.A. Bur. PI. 
Industr. Bull. 33: 21. 1903. Basionym: Diplachne 
brandegei Vasey. 

= Enteropogon brandegei (Vasey) Clayton. 


Gouinia brasiliensis (S. Moore) Swallen, Amer. J. Bot. 22: 
36. 1935. Basionym: Pogochloa brasiliensis S. Moore. 

DST.: Arge, Boli, Braz, Para. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Filgueiras, T. S. (1999); Ortiz, J. J. 
(1993); Renvoize, S. A. (1998); Swallen, J. R. 
(1935); Tirpe, A. M. (1975); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rigolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

SYN.: Pogochloa brasiliensis S. Moore. 


Gouinia cearensis (Ekman) Swallen, Amer. J. Bot. 22: 39. 
1935. Basionym: Diplachne cearensis Ekman. 
DST.: Braz. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Ortiz, J. J. (1993); Swallen, J. R. 
(1935). 
SYN.: Diplachne cearensis Ekman. 


Gouinia ekmanii Alain, Contr. Ocas. Mus. Hist. Nat. Colegio 
"De La Salle" no, 12: 5, f. 6. 1953. T-Protol.: [West 
Indies]: Cuba. 

DST.: Cari. 
= Status not yet determined. 


Gouinia gracilis Ekman ex Swallen, Amer. J. Bot. 22: 33, f. 
1. 1935. T-Protol.: Cuba: Oriente: in thickets on Cerro 
del Fraile at Holguinn, 17 Nov 1922,£. L. Ekman s.n. 
[HT: US-1161268]. T-Specim.: | of 2. Cuba: Oriente, 
Cerro del Fraile at Holguin, thickets, 17 Nov 1922, 
ELL. Ekman s.n. [Amer. Grass Natl. Herb. 1020] (HT: 
US-1161268; IT: CHAPA, GH, L, MO-1709940, MO- 
1702531, NY-71114, SI, UB]. 2 of 2. Cuba: Holguin, 
Cerro del Fraile, Province Oriente, 17 Nov 1922,F. L. 
Ekman 3235 {IT?: NY-71115 (possible)]. 
DST.: Cari. 
LVL. ACCPTC.:: 1. 
ALSO ACCPTD. BY: Catasus Guerra, L. (1997); 
Hitchcock, A. S. (1936); Ortiz, J. J. (1993); Swallen, 
J. R. (1935). 


Gouinia guatemalensis (Hack.) Swallen, Publ. Carnegie Inst. 
Wash. 436: 334. 1934. Basionym: Diplachne 
guatemalensis Hack, 


= Gouinia latifolia var. guatemalensis (Hack.) J.J. Ortiz. 


Gouinia guatemalensis (Hack.) Swallen, Amer. J. Bot. 22: 
40, 21. 1935. Basionym: Diplachne guatemalensis 
Hack. 

NOTE: isonym. 
E. NAME: Gouinia guatemalensis (Hack.) Swallen. 
= Gouinia latifolia var. guatemalensis (Hack.) J.J. Ortiz. 


Gouinia isabelensis J.J, Ortiz, Acta Bot. Mex. 23: 18. 1993. 
T-Protol.: Mexico: Nayarit: Isla Isabel, 27 Jul 1925, 
Solis 1 [HT: MEXU; IT: US-1268804]. 
DST.: Mexi. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Ortiz, J. J. 
(1993). 


Gouinia latifolia (Griseb.) Vasey, Contr. U.S. Natl. Herb. 
1(9): 365. 1895. Basionym: Tricuspis latifolia Griseb. 

DST.: Arge, Boli, Braz, Guat, Hond, Mexi, Nica, Para, 
Peru, Vene. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Brako, L. & J. L. Zarucchi (1993); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Filgueiras, T. S. (1999): Foster, R. 
C. (1958); Hitchcock, A. S. (1927); Killeen, T. J. 
(1990); Nicora, E. G. (1995); Ortiz, J. J. (1993); 
Renvoize, S. A. (1998); Swallen, J. R. (1935); Tovar, 
O. (1993); Tirpe, A. M. (1975); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

SYN.: Diplachne latifolia (Griseb.) Hack.; Diplachne 
monticola (Chase) McNeill; Sieglingia latifolia 
(Griseb.) Kuntze; Tricuspis latifolia Griseb. 


Gouinia latifolia var. guatemalensis (Hack.) J.J. Ortiz, Acta 
Bot. Mex. 23: 21. 1993. Basionym: Diplachne 
guatemalensis Hack. 

DST.: Guat, Hond, Mexi, Nica. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Ortiz, J. J. 


(1993). 

SYN.:  Diplachne — guatemalensis — Hack.; | Gouinia 
guatemalensis (Hack.) Swallen; Gouinia 
guatemalensis (Hack.) Swallen; Gouinia ramosa 
Swallen. 


Gouinia latifolia (Griseb.) Vasey var. latifolia. 
DST.: Arge, Boli, Braz, Para, Peru, Vene. 
LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Ortiz, J. J. (1993). 


Gouinia longiramea Swallen, Publ. Carnegie Inst. Wash. 
436: 334, f. 2. 1934. T-Protol.; Mexico: Yucatan: in 
brush around clearing at Chichen Itza, 7-13 Jul 1932, 
JR. Swallen 2385 [HT: US-1538059; IT: US]. T- 
Specim.: | of 1. Mexico: Yucatan: in brush around 
clearing at Chichen Itza, 7 Jul 1932-13 Jul 1932.7. 
Swallen 2385 [HT: US-1538059]. 

= Gouinia virgata (J. Pres!) Scribn. 


Gouinia mexicana (Scribn.) Vasey, Contr. U.S. Natl. Herb. 
1(9): 365. 1895. Basionym: Leptochloa mexicana 
Scribn. 

DST.: Mexi. 
LVL. ACCPTC.: 1. 


Catalogue of New World Grasses: II. 


123 


ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Ortiz, J. J. (1993); Swallen, J. R. (1935). 

SYN.: Diplachne mexicana (Scribn.) Hack.; Leptochloa 
mexicana Scribn. 


Gouinia papillosa Swallen, Publ. Carnegie Inst. Wash. 436: 
334, f. 3. 1934. T-Protol.: Mexico: Quintana Roo: in 
brush along trail in forest, Tancah, 4-5 Aug 1932,/.R. 
Swallen 2829 [HT: US-1538062]. T-Specim.: 1 of 1. 
Mexico: Yucatan: in brush along trail in forest, 
Tancah,, 4 Aug 1932-5 Aug 1932,/R. Swallen 2829 
[HT: US-1538062]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Davidse, G., 
M. Sousa S. & A. QO. Chater (editores generales) 
(1994): Espejo Serna, A., A. R. L6pez-Ferrari, and J. 
Valdes-Reyna (2000); Ortiz, J. J. (1993); Pohl, R. W. 
& G. Davidse (1994): Sousa S., M. & E. F. Cabrera 
C. (1983); Swallen, J. R. (1935). 


Gouinia paraguayensis (Kuntze) Parodi, Revista Mus. La 
Plata 34: 192, 176. 1934. Basionym: Arundinaria 
paraguayensis Kuntze. 

DST.: Arge, Boli, Cari, Guat, Hond, Para, Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Ortiz, J. J. (1993); Renvoize, S. A. 
(1998); Swallen, J. R. (1935); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

SYN.: Arundinaria paraguayensis Kuntze; Sieglingia 


paraguayensis Kuntze; Triodia paraguayensis 
(Kuntze) Hack. 

Gouinia paraguayensis (Kuntze) Parodi var. 
paraguayensis. 


DST.: Arge, Boli, Cari, Guat, Hond, Para, Peru. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Ortiz, J. J. (1993); Tiirpe, A. M. 
(1975). 


Gouinia paraguayensis var. tortuosa (Swallen) Tipe, 
Lilloa 34(6): 77, f. 2B. 1975. Basionym: Gouinia 
tortuosa Swallen. 

DST.: Arge, Boli. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Ortiz, J. J. (1993); Turpe, A. M. 

(1975). 
SYN.: Gouinia tortuosa Swallen. 


Gouinia polygama E. Fourn. ex Hemsl., Biol. Cent.-Amer., 
Bot. 3: 581. 1885. 
NOTE: nom. inval., as syn. of Festuca fournieriana 
Hemsley. 
= Gouinia virgata (J. Presl) Scribn. 


Gouinia polygama E. Fourn., Mexic. Pl. 2: 103. 1886. T- 
Specim.: | of 6. Mexico: Veracruz: pr. Veracruz, in 
sylvis umbrosis, 23 Aug 1865, Gouin 76 [LT: P], LT 
designated by Ortiz, Acta Bot. Mex. 23: 28 (1993). 2 
of 6. Mexico: Oaxaca: San Agustin, Oct, Liebmann 
505 [ST: MO-134695, US]. 3 of 6. Mexico: absque 
loco, Karwinsky 1000 [ST: US]. 4 of 6. Mexico: San 
Agustin, Oct, Liebmann 504 [ST: US]. 5 of 6. Mexico: 
Acapulco secus rivum pr. Pointe Griffon, Thiebaut 
1042 [ST]. 6 of 6. Florida, Karwinsky 44 [ST]. 

= Gouinia virgata (J. Presl) Scribn. 


Gouinia polygama var. major E. Fourn., Mexic. Pl. 2: 103. 
1886. T-Protol.: Mexico: Veracruz, Gouin 77. T- 
Specim.: 1 of 1. Mexico: Veracruz: 6 Sep 1865,Gouin 
77 (HT: P]. 

= Gouinia virgata (J. Presl) Scribn. 


Gouinia ramosa Swallen, Publ. Carnegie Inst. Wash. 436: 
333, f. 1. 1934. T-Protol.: Mexico: Yucatan: in brush 
at Chichen Itza, 7-12 Jul 1932, Swallen 2436 [HT: US- 
1538060]. T-Specim.: | of 2. Mexico: Yucatan: in 
brush at Chichen Itza, 7 Jul 1932-13 Jul 1932, /7R 
Swallen 2436 [IT: US-1538085]. 2 of 2. Mexico: 
Yucatan: in brush at Chichen Itza, 7 Jul 1932-13 Jul 
1932, LR. Swallen 2436 |HT: US-1538060]. 

= Gouinia latifolia var. guatemalensis (Hack.) J.J. Ortiz. 


Gouinia tortuosa Swallen, Amer. J. Bot. 22: 41. 1935. T- 
Protol.: Argentina: La Rioja: Patquia, Mar 1933, 
Lahitte & Castro 13 [HT: US-1538642]. T-Specim.: | 
of 1. Argentina: La Rioja, Mar 1933, Lahitte, Castro 
13 (REP. ARG. HERB. NAC. NO.) [HT: US-1538642; 
IT: BAA, BAB-47612]. 

= Gouinia paraguayensis var. tortuosa (Swallen) Tiirpe. 


Gouinia virgata (J. Presl) Scribn., Bull. Div. Agrostol., 
U.S.D.A. 4: 10. 1897. Basionym: Bromus virgatus J. 
Presl. 
DST.: Boli, Braz, Cari, Colo, CoRi, Ecua, ElSa, Guat, 
Hond, Mexi, Nica, Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Berendsohn, 
W.G. & A. E. Araniva de Gonzalez (1991); Brako, 
L. & J. L. Zarucchi (1993); Breedlove, D. E. (1986); 
Catasus Guerra, L. (1997); Davidse, G., M. Sousa S. 
& A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lépez-Ferrari, and J. Valdes-Reyna 
(2000); Foster, R. C. (1958); Hitchcock, A. S. (1927); 
Hitchcock, A. S. (1936); Jorgensen, P. M. & S. Leén- 
Yanez (eds.) (1999); Killeen, T. J. (1990); Macbride, 
J. F. (1936); McVaugh, R. (1983); Ortiz, J. J. (1993): 
Pohl, R. W. & G. Davidse (1994); Pohl, R. W. 
(1980); Swallen, J. R. (1935); Tovar, O. (1993). 
Bromus virgatus J. Presl; Diplachne virgata (J. Presl) 
Hack.; Festuca fournieriana Hemsl.; Festuca 
laxiflora A. Rich.; Gouinia longiramea Swallen; 
Gouinia polygama E. Fourn.; Gouinia polygama E. 
Fourn, ex Hemsl.; Gouinia polygama var. major E. 
Fourn.; Hackelia angustifolia Vasey ex Beal; 
Leptochloa palmeri Vasey ex Scribn.; Leptochloa 
polygama (E. Fourn.) Beal. 


SYN.: 


Gouinia virgata var. robusta J.J. Ortiz, Acta Bot. Mex. 23: 
30. 1993. T-Protol.: Mexico: Guerrero: near 
Tlalixtaquilla, 10 Dec 1894, Nelson 2255 [HT: US]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Ortiz, J. J. 
(1993). 


Gouinia virgata (J. Presl) Scribn. var. virgata. 
DST.: Boli, Braz, Cari, Colo, CoRi, Ecua, ElSa, Guat, 
Hond, Mexi, Nica, Peru. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Ortiz, J. J. (1993). 


Graphephorum flexuosum Thurb. ex A. Gray, Proc. Acad. 
Nat. Sci. Philadelphia 1863: 78. 1864. T-Specim.: | of 


124 


Catalogue of New World Grasses: II. 


1. USA: Nebraska, 1862, E. Hall & J.P. Harbour 635 
[IT: US-82055]. 
= Redfieldia flexuosa (Thurb. ex A. Gray) Vasey. 


Griffithsochloa G.J, Pierce, Bull. Torrey Bot. Club 105(2): 
134. 1978. TYPE: Griffithsochloa multifida (Griffiths) 
G.J. Pierce. 
= Bouteloua Lag. 


Griffithsochloa multifida (Griffiths) G.J. Pierce, Bull. Torrey 
Bot. Club 105(2): 134. 1978. Basionym:Cathestecum 
multifidum Griffiths. 

= Bouteloua multifidum (Griffiths) Columbus. 


Gymnopogon P. Beauy., Ess. Agrostogr. 41, 164. 
1812. T-Protol.: F19-20443. TYPE: Gymnopogon 
ambiguus (Michx.) Britton, Sterns & Poggenb. 

TRTD. BY: Peterson, P. M. 

NHT: Cynodonteae. NO. ACCPTD. SPP.: 12 SUBSP.: 4. 

CNSLTD. LIT.: Smith Jr., J. P., (1971). 

SYN.: Alloiatheros Elliott ex Raf.; Anthopogon Nutt.; 
Biatherium Desv.; Doellochloa Kuntze; Monochaete 
DOIl; Sciadonardus Steud. 


Sterns & 
Basionym: 


Britton, 
1888. 


Gymnopogon ambiguus (Michx.) 
Poggenb., Prelim. Cat. 69. 
Andropogon ambiguus Michx. 

DST.: Cari, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Fernald, M. (1950); Gleason, 
H. A. & A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Great Plains Flora Association 
(1986); Hitchcock, A. S. (1936); Hitchcock, A. S. 
(1951); Radford, A. E., H. E. Ahles & C. R. Bell 
(1968); Small, J. K. (1933); Smith Jr., J. P. (1971); 
Wunderlin, R. P. (1998). 

Agrostis boeckeleri Seub. ex Steud.; Alloiatheros 
ambiguus Elliott ex B.D. Jacks.; Alloiatheros 
ambiguus Elliott ex Kunth; Alloiatheros aristatus 
Raf. ex B.D. Jacks.; Alloiatheros lepturoides (Nutt.) 
Steud.; Andropogon ambiguus Michx.; Anthapogon 


SYN.: 


lepturoides Nutt.; Gymnopogon — distichophyllus 
Steud.; Gymnopogon  racemosus P. Beauv.; 
Gymnopogon — scoparius  Trin.; Sciadonardus 
distichophyllus Steud.; Stipa expansa Willd. ex 
Steud. 


Gymnopogon aristiglumis Hitche., Proc. Biol. Soc. Wash. 
40: 82. 1927. T-Protol.: El Salvador: Rosario, Jan 
1924, Calderén 1924 [HT: US-1169956]. T-Specim.: 
1 of 1. El Salvador: Rosario, Jan 1924, 8. Calderén 
1924 (HT: US-1169956]. 

DST.: Boli, ElSa. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Berendsohn, W. G. & A. E. Araniva 
de Gonzalez (1991); Davidse, G., M. Sousa S. & A. 
QO. Chater (editores generales) (1994); Pohl, R. W. 
(1994); Smith Jr., J. P. (1971). 


Gymnopogon beyrichianus (Kunth) Parodi, Revista Centro 
Estud. Agron. 18: 148, f. 55. 1925. Basionym:Chloris 
beyrichiana Kunth. 

= Chloris pyenothrix Trin. 


Gymnopogon biflorus Pilg., Bot. Jahrb. Syst. 30(1): 139. 
1901. T-Protol.: Brasil: Mattogrossso: auf trocknem, 


steinigen Boden im Oberen Cuyabathal, Apr. 1899,/7. 
Meyer 489 (HT: B; IT: BAA (fragm. ex B)]. 
= Gymnopogon spicatus (Spreng.) Kuntze. 


Gymnopogon biflorus Pilg. var. biflorus. 
= Gymnopogon spicatus (Spreng.) Kuntze. 


Gymnopogon biflorus var. ciliatilemma Burkart, Bol. Soc. 
Argent. Bot. 12: 290, f. 3. 1968. T-Protol.: Argentina: 
Prov. Entre Rios: Dept. Gualeguaychu, predelta, 
Ibicuy, 4. Burkart 3598 (HT: SI]. T-Specim.: 1 of 2. 
Argentina: Entre Rios: Dpto. Gualeguaychu: Ibicuy, 
Burkart 3598 |HT]. 2 of 2. Argentina: Prov. Entre 
Rios, Ibicuy., 4 Dec 1930, A. Burkart [IT]. 

= Gymnopogon legrandii Roseng., B.R. Arill. & Izag. 


Gymnopogon brevifolius Trin., Gram. Unifl. Sesquifl. 238. 
1824. T-Protol.: USA: Delaware, Bernhardi s.n. [HT: 
LE]. T-Specim.: | of 1. USA: Delaware, Bernhardi 
[HT: LE-TRIN-2118.01 (& fig.)]. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Fernald, M. (1950); Gleason, 
H. A. & A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Hitchcock, A. S. (1951); Radford, 
A. E., H. E. Ahles & C. R. Bell (1968); Small, J. K. 
(1933); Smith Jr., J. P. (1971); Wunderlin, R. P. 
(1998). 

Anthopogon brevifolius Nutt. ex Trin.; Anthopogon 
filiforme  Nutt.; | Gymnopogon  racemosus — vat. 
filiformis Chapm.; Gymnopogon racemosus var. 
racemosus. 


SYN.: 


Gymnopogon brevisetus (Hack.) J.P. Smith, lowa State J. Sci. 
45(3): 344. 1971. Basionym: Gymnopogon spicatus 
var. brevisetus Hack. 

= Gymnopogon grandiflorus Roseng., B.R. Arill. & Izag. 


Gymnopogon burchellii (Munro ex Déll) Ekman, Ark. Bot. 
11(4): 35, t. 1, f. 4. 1912. Basionym: Leptochloa 
burchellii Munro ex DOll. 

DST.: Arge, Boli, Braz, Para, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Burkart, A. (1969); Caro, J. A. 
(1982); Filgueiras, T. S. (1991); Filgueiras, T. S. 
(1999): Nicora, E. G. (1995); Renvoize, S. A. (1988); 
Renvoize, S. A. (1998); Rosengurtt, B. B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 
Smith Jr., J. P. (1971); Zuloaga, F. O., E. G. Nicora, 
Z. E. Riugolo de Agrasar, O. Morrone, J. Pensiero & 
A. M. Cialdella (1994). 

Gymnopogon muticus Hack.; Leptochloa burchellii 
Munro ex DOIl. 


SYN.: 


Gymnopogon chapmanianus Hitchc., Amer. J. Bot. 2: 306. 
1915. T-Protol.: USA: Florida: Seminole Co.: 
Sandford, sandy meadow, A. Chase 4/35 [HT: US- 
733684]. T-Specim.: | of 1. USA: Florida: Seminole 
Co., Sandford, 25 Sep 1907, A. Chase 4135 [HT: US- 
733684). 

DST.: USA. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Hitchcock, A. S. 
(1951); Small, J. K. (1933); Smith Jr., J. P. (1971); 
Wunderlin, R. P. (1998). 

SYN.: Gymnopogon floridanus Swallen. 


Catalogue of New World Grasses: II. 


125 


Gymnopogon distichophyllus Steud., Syn. Pl. Glumac. 1: 218. 
1854. T-Specim.: | of 2. USA: Texas, Seubert herb. 
no. 128 [ST], as Agrostis boeckeleri [nom. inval.]. 2 of 
2. USA: Louisiana, Hartmann 57, Louisaina herb. 
[ST]. 

= Gymnopogon ambiguus (Michx.) Britton, Sterns & 
Poggenb. 


Gymnopogon doellii Boechat & Valls, Bradea 5(28): 314, f. 
1. 1990. T-Protol.: Brazil: Distrito Federal: Area de 
Protecdo Ambiental da Bacia do Rio Sado Bartolomeu, 
cerca de 5km S do local onde sera construida a 
barragem do futero lago, em leito seco de um cérego 
temporario, 6 Jun 1989, 7S. Filgueiras & S.P. Arauo 
1803 [HT: CEN. IT: IBGE]. 

NOTE: new taxon name, description, and type, for G. 
rigidus D6ll. hom. illeg. 

DST.: Braz. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Filgueiras, T. S. (1991); Filgueiras, 
T.S. (1999). 

SYN.: Gymnopogon rigidus Doll. 


Gymnopogon fastigiatus Nees, Fl. Bras. Enum. Pl. 2: 430. 
1829. T-Specim.: 1 of 1. Brazil: Sado Paulo, 
Anonymous s.n. (HT: B; IT: LE-TRIN-2119.01 (fragm. 
ex B)]. 

DST.: Boli, Braz, Colo, CoRi, Pana, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Filgueiras, T. S. 
(1999): Killeen, T. J. (1990); Pohl, R. W. (1980); 
Pohl, R. W. (1994); Renvoize, S. A. (1998); Smith 
Jr., J. P. (1971). 

SYN.: Doellochloa fastigiata (Nees) Kuntze; Monochaete 
Jastigiata (Nees) DOll. 


Gymnopogon fastigiatus Nees subsp. fastigiatus. 
DST.: Boli, Braz, Colo, CoRi, Pana, Vene. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Pohl, R. W. 
(1980); Pohl, R. W. (1994); Smith Jr., J. P. (1971). 


Gymnopogon fastigiatus subsp. jubiflorus (Hitchc.) J.P. 
Smith, Iowa State J. Sci. 45: 361, 1971. Basionym: 
Gymnopogon jubiflorus Hitche. 

DST.: Boli. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Pohl, R. W. 
(1994); Smith Jr., J. P. (1971). 

SYN.: Gymnopogon jubiflorus Hitche. 


Gymnopogon filiformis Griseb., Fl. Brit. W. I. 538. 1864. T- 
Protol.: Trinidad, Crueger s.n. T-Specim.: 1 of 1. 
Trinidad, no date, Crueger s.n. [HT: GOET; IT: B, 
US-822349]. 

= Gymnopogon spicatus (Spreng.) Kuntze. 


Gymnopogon floridanus Swallen, N. Amer. FI. 17(8): 607. 
1939. T-Protol.: USA: Florida: Clay Co.: collected on 
open, sandy prairie, Penny Farms, 15 Nov 1938, 
Swallen 5596 [HT: US-1723536]. T-Specim.: | of 1. 
USA: Florida: Clay Co., Penny Farms, 15 Nov 1938, 
JR. Swallen 5596 (HT: US-1723536; IT: L]. 
= Gymnopogon chapmanianus Hitchce. 


Gymnopogon foliosus (Willd.) Nees, Fl. Bras. Enum. PI. 2: 
426. 1829. Basionym: Chloris foliosa Willd. 

DST.: Braz, Cari, FrGu, Guya, Peru, Suri, USA, Vene. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 
(1987); Anonymous (1986); Brako, L. & J. L. 
Zarucchi (1993); Doll, J. C. (1878); Filgueiras, T. S. 
(1991); Filgueiras, T. S. (1999); Hitchcock, A. S. 
(1936); Judziewicz, E. J. (1990); Liogier, H. & L. 
Martorell (1982); Longhi-Wagner, H. M. (1990); 
Macbride, J. F. (1936); Renvoize, S. A. (1984); 
Smith Jr., J. P. (1971); Tovar, O. (1993). 

Aristida chloris Salzm. ex DO6ll; Aristida geminata 
Willd. ex Steud.; Biatherium foliosa (Willd.) Desv.; 
Chloris aristida Salzm. ex Steud.; Chloris foliosa 
Willd.; Gymnopogon pullulans Doll. 


SYN.:: 


Gymnopogon glaber Caro, Dominguezia 5: 17, f. 5. 1982. 


T-Protol.: Argentina: Salta: Dpto. Anta: Cerro 
Maldonado, Jun 1934, Ragonese /13/ [HT: BA]. 
DST.: Arge. 


LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Caro, J. A. (1982); Zuloaga, F. O., E. 
G. Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 


Gymnopogon grandiflorus Roseng., B.R. Arill. & Izag., 
Bol. Fac. Agron. Univ. Montevideo 103: 20, f. 4. 
1968. T-Protol.: Uruguay: Depto Maldonado: arroyo 
Solis, proximo a la barra, 8 Apr 1955, Rosengurtt B- 
6347 [HT: MVFA]. T-Specim.: 1 of 1. Uruguay: 
Maldonado: Arroyo Solis, proxima a la barra, 8 Apr 
1955, B. Rosengurtt B-6347 (HT: MVFA; IT: BAA, 
US-2946505]. 

DST.: Arge, Braz, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Burkart, A. (1969); Caro, J. A. 
(1982); Rosengurtt, B., B. R. Arrillaga de Maffei & 
P. Izaguirre de Artucio (1970); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rigolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

SYN.: Gymnopogon  brevisetus (Hack.) J.P. Smith; 
Gymnopogon spicatus var. brevisetus Hack. 


Gymnopogon haumanii Parodi, Physis (Buenos Aires) 4: 183. 
1918. T-Specim.: | of 2. Paraguay, Sapucay, Nov 
1894, Dresse [ST]. 2 of 2. [Argentina]: Trinidad, 
Asuncion, M.S. Bertoni 2493 [ST]. 
NOTE: cites Chloris radiata sensu. Hack., non SW, in 
Berttoni, Anal, Ci. Paraguay (ser. I1) 2: (1918). 
= Chloris pycnothrix Trin. 


Gymnopogon jubiflorus Hitche., Contr. U.S. Natl. Herb. 
24(8) 412. 1927. T-Protol.: Bolivia: collected at 
Guanay, northern Bolivia, alt. 600 m, May 1886,/7. 1. 
Rusby 215 [HT: US-879400]. T-Specim.: | of 1. 
Bolivia: Guanay, May 1886, H.-H. Rusby 215 [HT: US- 
879400]. 

= Gymnopogon fastigiatus subsp. jubiflorus (Hitchc.) J.P. 
Smith. 


Gymnopogon laevis Nees, Fl. Bras. Enum. Pl. 2: 428. 1829. 
T-Protol.: Habitat ad Monte Video, Se//ow s.n. [HT: 
B]. T-Specim.: | of 1. Brazil, Sedlow s.n. [12088 at L] 

[HT: B; IT: L, US (fragm. ex B)]. 

= Gymnopogon spicatus fo. spicatus. 


Gymnopogon laevis var. laevis. 


126 


Catalogue of New World Grasses: II. 


= Gymnopogon spicatus fo. spicatus. 


Gymnopogon laevis vat. pluriflorus Doll, Fl. Bras. 2(3): 81. 
1878. T-Protol.: Brasil: prope Ypanema, Riedel s.n. 
= Gymnopogon spicatus fo. spicatus. 


Gymnopogon legrandii Roseng., B.R. Arill. & Izag., Bol. 
Fac. Agron. Univ. Montevideo 103: 22, f. 5. 1968. T- 
Protol.: Uruguay: Canelones: Carrasco, Aeropuerto, en 
arenas, 4 Mar 1949, Rosengurtt B-529] |HT: MVFA; 
IT; BAA, US]. T-Specim.: | of 1. Uruguay: 
Canelones: Carrasco, Aeropuerto, en arenas, 4 Mar 
1949, B. Rosengurtt B-5291 (HT: MVFA,; IT: BAA, 
SI, US]. 

DST.: Arge, Braz, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Burkart, A. (1969); Caro, J. A. 
(1982); Rosengurtt, B., B. R. Arrillaga de Maffei & 
P. Izaguirre de Artucio (1970); Zuloaga, F. O., E. G. 
Nicora, Z. E. Ruigolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

Gymnopogon biflorus var. ciliatilemma Burkart; 
Gymnopogon swalleniiJ.P. Smith. 


SYN: 


Gymnopogon longifolius E. Fourn., Mexic. Pl. 2: 144. 1886. 
T-Protol.; Mexico: Veracruz: Oct, Gouin 52, T- 
Specim.: | of 1. Mexico: Veracruz, 1867, Gouin 52 
[HT: P; IT: US-s.n. (fragm. ex P)]. 

= Enteropogon chlorideus (J. Pres!) Clayton. 


Gymnopogon mollis Nees, Fl. Bras. Enum. PI. 2: 427. 1829. 
T-Specim.: | of 3. Brazil: Pernambuco, Martius [ST: 
M?]. 2 of 3. Brazil: Piauhi: Martins Iter Poros, K.F. P. 
von Martius s.n. [ST: M?; IST: US-3386738 (fragm.)]. 
3 of 3. Brazil: Bahia, Martius s.n. [ST: M?]. 
= Enteropogon mollis (Nees) Clayton. 


Gymnopogon muticus Hack., Anales Mus. Nac. Buenos Aires 
21: 117. 1911. 
NOTE: nom. nud. 
= Gymnopogon burchellii (Munro ex Déll) Ekman. 


Gymnopogon pullulans Doll, Fl. Bras. 2(3): 82, pl. 23, f. 1. 
1878. T-Specim.: | of |. Brazil, Burchell 5398 [LT: k; 
ILT: B, GH, P], ILT designated by Smith, lowa St. J. 
Sci. 45: 366. (1974). 
= Gymnopogon foliosus (Willd.) Nees. 


Gymnopogon racemosus P. Beauv., Ess. Agrostogr. 41, 164, 
t.9, f.3. 1812. 

NOTE: nom. nov., nom. illeg. superfl. for Andropogon 
ambiguus Michx. 

REPLACED NAME: Andropogon ambiguus Michx. 

CORRECT NAME: Andropogon ambiguus Michx. 

= Gymnopogon ambiguus (Michx.) Britton, Sterns & 
Poggenb. 


Gymnopogon racemosus Var. filiformis Chapm., Fl. S.E. U.S. 
556. 1860. 
= Gymnopogon brevifolius Trin. 


Gymnopogon racemosus Trin. var. racemosus. 
= Gymnopogon brevifolius Trin. 


Gymnopogon radiatus (L.) Parodi, Physis (Buenos Aires) 4: 
180. 1918. Basionym: Agrostis radiata L. 
= Chloris radiata (L.) Sw. 


Gymnopogon radiatus var. beyrichiana (Kunth) Parodi, 


Physis (Buenos Aires) 4: 173. 
Chloris beyrichiana Kunth. 
= Chloris pycnothrix Trin. 


1918. Basionym: 


Gymnopogon rigidus Doll, Fl. Bras. 2(3): 80. 1878. T-Protol.: 
Brazil: Habitat ad Lagoa Santa prov. Minarum, 
Warming s.n. T-Specim.: | of 1. Brazil: Minas Gerais: 
in campis ad Lagoa Santa, Warming s.n. [HT: B]. 

NOTE: hom. illeg. 

E. NAME: Gymnopogon rigidus Thwaites. 

CORRECT NAME: Gymnopogon doellii Boechat & Valls. 
= Gymnopogon doellii Boechat & Valls. 


Gymnopogon rupestre Ridl., J. Linn. Soc., Bot. 27: 73. 1890. 
T-Protol.: Brazil: Fernando do Noronha, cliffs on 
Portugese Bay, 14 Sep 1887, Ridley & Ramage s.n. T- 
Specim.: | of 1. Brazil: Fernando do Noronha, cliffs 
on Portugese Bay, 14 Sep 1887, H.N. Ridley et al. 165 
{HT: BM; IT: US-821287]. 

= Enteropogon mollis (Nees) Clayton. 


Gymnopogon scoparius Trin., Gram. Unifl. Sesquifl. 237. 
1824. T-Protol.: USA: V. sp. e New Jersey[HT: LE]. 
T-Specim.: | of 1. Nov. Coesar., 1835,Gray [HT: LE- 
TRIN-2123.01 (& fig.)]. 

= Gymnopogon ambiguus (Michx.) Britton, Sterns & 
Poggenb. 


Gymnopogon spicatus (Spreng.) Kuntze, Revis. Gen. PI. 
3(2): 354. 1898. Basionym: Polypogon  spicatus 
Spreng. 

DST.: Arge, Beli, Boli, Braz, Cari, Colo, Guat, Guya, Hond, 

Mexi, Para, Urug, Vene. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 
(1987); Balick, M. J.. M. H. Nee & D. E. Atha 
(2000); Beetle, A. A. (1977); Burkart, A. (1969); 
Caro, J. A. (1982); Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Espejo Serna, A., 
A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Filgueiras, T. S. (1991); Filgueiras, T. S. (1999); 
Foster, R. C. (1958); Hitchcock, A. S. (1927); 
Hitchcock, A. S. (1936); Judziewicz, E. J. (1990): 
Killeen, T. J. (1990); McClure, F. A. (1955); Molina 
R., A. (1975); Pohl, R. W. (1994); Renvoize, S. A. 
(1984); Renvoize, S. A. (1988); Renvoize, S. A. 
(1998); Smith Jr., J. P. (1971); Smith, L. B., D.C. 
Wasshausen & R. M. Klein (1981); Zuloaga, F. O., 
E. G. Nicora, Z. E. Rugolo de Agrasar, O. Morrone, 
J. Pensiero & A. M. Cialdella (1994). 

Gymnopogon biflorus Pilg., Gymnopogon biflorus 
var. biflorus; Gymnopogon _filiformis  Griseb.; 
Muhlenbergia spicata (Spreng.) Kunth; Polypogon 
spicatus Spreng.; Trichochloa spicata (Spreng.) 
Schult. 


SYN:: 


Gymnopogon spicatus var. brevisetus Hack., Anales Mus. 
Nac. Buenos Aires 21: 117. 1911. T-Protol.: [HT: G]. 
NOTE: ser 3, 14. 
= Gymnopogon grandiflorus Roseng., B.R. Arill. & Izag. 


Gymnopogon  spicatus fo. longiaristatus (Kuntze) J.P. 
Smith, lowa State J. Sci. 45(3): 371. 1971. Basionym: 
Gymnopogon spicatus var. longiaristatus Kuntze. 

DST.: Boli, Braz, Colo, Hond, Vene. 
LVL. ACCPTC:: I. 


Catalogue of New World Grasses: II. 


127 


ALSO ACCPTD. BY: Foster, R. C. (1958); Smith Jr., J. P. 
(1971). 
SYN.: Gymnopogon spicatus var. longiaristatus Kuntze. 


Gymnopogon spicatus var. longiaristatus Kuntze, Revis. 
Gen. Pl. 354. 1891. T-Protol.: Bolivia: Ost-Velasco, 
200 m. T-Specim.: | of 1. [LT: NY], LT designated by 
Smith, Iowa St. J. Sci. 45: 372. 1971. 
= Gymnopogon spicatus fo. longiaristatus (Kuntze) J.P. 
Smith. 


Gymnopogon spicatus var. pluriflorus (D6ll) Stuck., Anales 
Mus. Nac. Buenos Aires 21: 117. 1911. Basionym: 
Gymnopogon laevis var. pluriflorus DOll. 

NOTE: comb. illeg. 
= Gymnopogon spicatus fo. spicatus. 


Gymnopogon  spicatus. var. pluriflorus (D6ll) Parodi 
Basionym: Gymnopogon laevis var. pluriflorus Doll. 
NOTE: comb. superfl. 
= Gymnopogon spicatus fo. spicatus. 


Gymnopogon spicatus (Spreng.) Kuntze fo. spicatus. 

DST.: Arge, Beli, Boli, Braz, Cari, Colo, Guat, Guya, Hond, 
Mexi, Para, Urug, Vene. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Smith Jr., J. P. (1971). 

SYN.: Gymnopogon biflorus Pilg.; Gymnopogon filiformis 
Griseb.; Gymnopogon laevis Nees; Gymnopogon 
laevis var. laevis; Gymnopogon laevis var. pluriflorus 
Doll; Gymnopogon spicatus var. pluriflorus (D6ll) 
Parodi; Gymnopogon spicatus var. pluriflorus (D6ll) 
Stuck.; Gymnopogon spicatus vat. spicatus. 


Gymnopogon spicatus (Spreng.) Kuntze var. spicatus. 
= Gymnopogon spicatus fo. spicatus. 


Gymnopogon swallenii J.P. Smith, lowa State J. Sci. 45(3): 
373. 1971. T-Protol.: Brazil: Rio Grande do Sul: Rio 
Grande, Swallen 9235 [HT: US-1960896]. T-Specim.: 
1 of 1. Brazil: Rio Grande do Sul, 3 May 1946,./R. 
Swallen 9235 [HT: US-1960896; IT: US-3135306]. 

= Gymnopogon legrandii Roseng., BR. Arill. & Izag. 


Gymnopogon virletii E. Fourn., Mexic. Pl. 2: 144. 1886. T- 
Protol.: Mexico: San Luis de Potosi, Virlett 1441 [HT: 
P}. 
= Enteropogon chlorideus (J. Pres!) Clayton. 


Hackelia Vasey ex Beal, Grass. N. Amer. 2: 438. 1896. 
NOTE: nom. inval., as syn. of Gouinia polygama, non Opiz 
1838. 
= Gouinia E. Fourn. ex Benth. & Hook. f. 


Hackelia angustifolia Vasey ex Beal, Grass. N. Amer. 2: 438. 
1896. 
NOTE: nom. inval., as syn. of Gouinia polygama Fourn. 
= Gouinia virgata (J. Presl) Scribn. 


Halochloa Griseb., Abh. Konigl. Ges. Wiss. Gottingen 24: 
285. 1879. 
NOTE: later hom., non Kuetzing 1843. 
= Monanthochloe Engelm. 


Halochloa acerosa Griseb., Symb. Fl. Argent. 285. 1879. T- 
Specim.: | of 1. Lorentz & Hieronymis 488 [IT: US 
(fragm. ex Griseb herb.)]. 

= Monanthochloe acerosa (Griseb.) Speg. 


Heleochloa Host ex Roem., Collecteana 233. 1809. TYPE: 
Heleochloa alopecuroides (Pill. & Mitt.) Roem. 
= Crypsis Aiton. 


Heleochloa alopecuroides (Piller & Mitterp.) Host, Icon. 
Descr. Gram. Austriac. 1: 23, t. 29. 1801. Basionym: 
Phleum alopecuroides Piller & Mitterp. 

= Crypsis alopecuroides (Piller & Mitterp.) Schrad. 


Heleochloa compacta (Steud.) T. Durand & Schinz, Consp. 
Fl. Afr. 5: 814. 1894. Basionym: Crypsis compacta 
Steud. 

= Crypsis vaginiflora (Forssk.) Opiz. 


Heleochloa juncea P. Beauv., Ess. Agrostogr. 24, 147, 164, t. 
7, £.2. 1812. 
REPLACED NAME: Agrostis juncea Michx. 
BLOCKING NAME: Agrostis juncea Lam. 
= Sporobolus junceus (P. Beauy.) Kunth. 


Heleochloa schoenoides (L.) Host, Icon. Descr. Gram. 
Austriac. I: 23, t. 30. 1801. Basionym: Phleum 
schoenoides L. 

= Crypsis schoenoides (L.) Lam. 


Hemigymnia malabarica (L.) Henrard, Meded. Rijks-Herb. 
61: 12. 1930. Basionym: Poa malabarica L. 
NOTE: nom. rej., basionym rej. [ICBN art. 56.2]. 
= Leptochloa fusca (L.) Kunth. 


Hemimunroa (Parodi) Parodi, Notas Mus. La Plata, Bot. 2: 4. 
1937. Basionym: Munroa subg. Hemimunroa Parodi. 
= Munroa Torr. 


Hemimunroa andina (Phil.) Parodi, Notas Mus. La Plata, Bot. 
2: 6. 1937. Basionym: Munroa andina Phil. 
= Munroa andina Phil. 


Hemimunroa andina (Phil.) Parodi var. andina. 
= Munroa andina Phil. 


Hemimunroa andina var. breviseta (Hack.) Parodi, Notas 
Mus. La Plata, Bot. 11: 6. 1937. Basionym: Munroa 
andina var. breviseta Hack. 

= Munroa andina Phil. 


Heterosteca Desv., Nouv. Bull. Sci. Soc. Philom. Paris 2: 
188. 1810. TYPE: Heterosteca juncifolia Desv. 
= Bouteloua Lag. 


Heterosteca juncifolia Desv., Nouv. Bull. Sci. Soc. Philom. 
Paris 2: 188. 1810. T-Protol.: Habitat in Antillis. T- 
Specim.: | of 3. Cuba: La Habana, 4./.A. Bonpland 
132B [HT: P; IT: US-2830954(fragm.)]. 2 of 3. 
Antilles, Coll. Ukn. s.n. [T: P(frgam., US-2830952)]. 3 
of 3. Cuba, no date, C. Wright 739 [T: L:NY-70990, 
NY-70991]. 

= Bouteloua repens (Kunth) Scribn. 


Heterosteca rhadina Nash, Bull. Torrey Bot. Club 30(7): 
386. 1903. T-Protol.: Puerto Rico: collected on the 
Adjuntas road, near Ponce, Heller 6057 [HT: NY]. T- 
Specim.: | of 1. Puerto Rico: on the Adjuntas road, 3 
miles from Ponce, 18 Nov 1902, 4.4. Heller 6057 
{HT: NY-71113; IT: US-426292]. 

= Bouteloua repens (Kunth) Scribn. 


Heterostega americana (L.) Desv., J. Bot. Agric. 1: 68. 1813. 
Basionym: Aristida americana L. 
= Bouteloua americana (L.) Scribn. 


128 


Catalogue of New World Grasses: II. 


Heterostega curtipendula Schwein. ex Hook. f., Trans. Linn. 
Soc. London 20: 175. 1847. 
NOTE: as "Heterostegon" curtipendula; as syn. of Eutriana 
affinis Hook. f. 
= Bouteloua curtipendula (Michx.) Torr. 


Hexarrhena J. Presl, Relig. Haenk. 1(4-5): 326. 1830. TYPE: 
Hexarrhena cenchroides J. Pres}. 
= Hilaria Kunth. 


Hexarrhena cenchroides J. Presl, Relig. Haenk. 1(4-5): 326, 
t. 14. 1830. T-Protol.: Hab. in Mexico, Haenke s.n., 
Possibly the same as Hilaria cenchroides Kunth type. 
T-Specim.: 1 of 1. Mexico, 7’Haenke [IT: US- 
2906093 (fragm.)]. 

= Hilaria cenchroides Kunth. 


Hilaria Kunth, Nov. Gen. Sp. 1: ed. qu. 116. 1815 
[1816]. TYPE: Hilaria cenchroides Kunth. 
TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 7, SUBSP.: 2. 
CNSLTD. LIT.: Sohns, E. R., (1956). 
SYN.:  Hexarrhena J. Presl; Schleropelta 
Symbasiandra Steud, 


Buckley; 


Hilaria annua Reeder & C. Reeder, Madrofio 35: 7. 1988. T- 


Protol.: Mexico: Colima: 10 km S of Ciudad Colima, 
300 m, abundant along roadside with other rank 
weeds, 300 m, 24 Sep 1974, Reeder & Reeder 6333 
[HT: ARIZ; IT: MEXU, MICH, RM, US]. T-Specim.: 
| of 1. Mexico: Colima, 24 Sep 1974, /.R. Reeder & 
C.G. Reeder 6333 [HT: ARIZ; IT: MEXU, MICH, 
RM, US-3206089]. 
DST.: Mexi. 
LVL. ACCPTC.:: 1. 
ALSO ACCPTD. BY: Espejo Serna, A., A. R. Léopez- 
Ferrari, and J. Valdes-Reyna (2000); Reeder, J. R. & 
C. G. Reeder (1988). 


Hilaria belangeri (Steud.) Nash, N. Amer. Fl. 17(2): 135. 
1912. Basionym: Anthephora belangeri Steud. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Hitchcock, A. S. (1951); McVaugh, R. 
(1983); Munz, P. A. (1968); Munz, P. A. (1974); 
Shreve, F. & I. L. Wiggins (1964); Sohns, E. R. 
(1956). 

Anthephora belangeri Steud.; Hilaria cenchroides 
var. texana Vasey; Hilaria texana (Vasey) Nash; 
Schleropelta stolonifera Buckley. 


SYN.: 


Hilaria belangeri (Steud.) Nash var. belangeri. 
DST.: Mexi, USA. 
LVL. ACCPTC.: L. 


Hilaria belangeri UNRANKED longifolia (Vasey) Hitchc., 
Proc. Biol. Soc. Wash. 41: 162. 1928. Basionym: 
Hilaria cenchroides var. longifolia Vasey. 

= Hilaria belangeri var. longifolia (Vasey) Hitchc. 


Hilaria belangeri var. longifolia (Vasey) Hitche., Man. 
Grasses U.S. 200: 870. 1935. Basionym: Hilaria 
cenchroides var. longifolia Vasey. 

NOTE: first ranked combination. 

DST.: Mexi, USA. 


LVL. ACCPTC.:: 1. 
SYN.: Hilaria belangeri UNRANKED longifolia (Vasey) 
Hitche. 


Hilaria cenchroides Kunth, Nov. Gen. Sp. 1: 117, t. 37. 
1815 [1816]. T-Protol.: Crescit in planitie montana 
regni Mexicani, inter Zelaya et Guanaxuato, locis 
subfrigidis, alt. 980 hexap., Humboldt & Bonpland s.n. 
T-Specim.: | of 1. Mexico: inter Zelaya et 
Guanaxuato, Humboldt & Bonpland s.n. [HT: P; IT: 
LE-TRIN-0379.01 (fragm.), US-81367 (fragm.)]. 

DST.: Guat, Mexi, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Breedlove, D. E. (1986); Davidse, G. (1994); 
Davidse, G., M. Sousa 8. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. & 
R. Moran (1981); McClure, F. A. (1955); McVaugh, 
R. (1983); Shreve, F. & I. L. Wiggins (1964); Sohns, 
E.R. (1956). 

SYN.: Hexarrhena cenchroides J. Presl. 


Hilaria cenchroides var. ciliata Scribn., Proc. Acad. Nat. Sci. 
Philadelphia 1891: 293. 1891. T-Protol.: Mexico: near 
Guadalajara, 17 May 1890, Pringle 3/28 [HT: US]. T- 
Specim.: | of 1. Mexico: Jalisco: valley of the Rio 
Grande de Santiago at Atequiza, 17 May 1898,Pringle 
3/28 (HT: US-70845; IT: CM, NY], despite the 
difference in citation in the protologue, based on the 
same collection.—gd. 

NOTE: as "ciliatus". 
= Hilaria ciliata (Scribn.) Nash. 


Hilaria cenchroides var. longifolia Vasey, Proc. Amer. Acad. 
Arts 24: 80. 1889. T-Specim.: | of 1. Mexico: Sonora: 
Guayamas, 1887, £. Palmer 347 (HT: US-926525; IT: 
US-81938]. 

= Hilaria belangeri 
Hitche. 


UNRANKED longifolia (Vasey) 


Hilaria cenchroides var. texana Vasey, Contr. U.S. Natl. 
Herb. 1(2): 53. 1890. T-Specim.: 1 of 1. USA: Texas: 
Duval Co., 1890, G.C. Nealley 600 [HT: US-81368}. 

= Hilaria belangeri (Steud.) Nash. 


Hilaria ciliata (Scribn.) Nash, N. Amer. Fl. 17(2): 136. 1912. 
Basionym: Hilaria cenchroides var. ciliata Scribn. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W. & R. Moran (1981); McVaugh, R. 
(1983); Sohns, E. R. (1956). 

SYN.: Hilaria cenchroides var. ciliata Scribn.; Hilaria 
ciliata (Scribn.) Sohns. 


Hilaria ciliata (Scribn.) Sohns, J. Wash. Acad. Sci. 46: 317. 
1956. Basionym: Hilaria cenchroides var. ciliata 
Scribn. 

NOTE: isonym. 
E. NAME: Hilaria ciliata (Scribn.) Nash. 
= Hilaria ciliata (Scribn.) Nash. 


Hilaria hintonii Sohns, J. Wash. Acad. Sci. 46(10): 319, f. 
30-34. 1956. T-Protol.: Mexico: Temascaltepec, 
Lluvianos, lIlano. 9/8/1933, Hinton 4502 [HT: US- 
1840874]. T-Specim.: | of 1. Mexico: Temascaltepec, 


Catalogue of New World Grasses: II. 


129 


9 Aug 1933 or 8 Sep 1933,G.B. Hinton 4502 (HT: 
US-1840874]. 
DST.: Mexi. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Sohns, E. R. (1956). 


Hilaria jamesii (Yorr.) Benth., J. Linn. Soc., Bot. 19: 62. 
1881. Basionym: Pleuraphis jamesii Torr. 
= Pleuraphis jamesii Torr. 


Hilaria mutica (Buckley) Benth., J. Linn. Soc., Bot. 19: 62. 
1881. Basionym: Pleuraphis mutica Buckley. 
= Pleuraphis mutica Buckley. 


Hilaria rigida (Thurb.) Benth. ex Scribn., Bull. Torrey Bot. 
Club 9: 86. 1882. Basionym: Pleuraphis rigida Thurb. 
= Pleuraphis rigida Thurb. 


Hilaria semplei Sohns, J. Wash. Acad. Sci. 46(10): 319, f. 
36-50. 1956. T-Protol.: Mexico: Michoacan: Llanos de 
Antufiez, ca. 12 mi E of Apatzingan, alt. 1000 ft, Nov 
1955, dense heavy stands on very heavy clay soil, 
dominant grass over many areas, A. 7. Semple s.n. 
[HT: US-2183565]. T-Specim.: | of 1. Mexico: 
Michoacan: Llanos de Antufiez, ca. 12 mi E of 
Apatzingan, elev. 1000 ft, Nov 1955,4.T- Semple s.n. 
[HT: US-2183565; IT:US-2378480]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Sohns, E. R. (1956). 


Hilaria sericea Benth., J. Linn. Soc., Bot. 19; 62. 1881. 
NOTE: nom. nud. 
= Pleuraphis jamesii Torr. 


Hilaria swallenii Cory, Wrightia 1: 215. 1948. T-Specim.: | 
of 1. USA: Texas: Jeff Davis Co.: Musquiz Canyon, 
28 Jul 1938, O.R. Sperry 7778 [TYPE: US-1839917]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Hitchcock, A. S. (1951); Sohns, E. R. 
(1956). 


Hilaria texana (Vasey) Nash, Fl. South. U.S. 68. 1903. 
Basionym: Hilaria cenchroidesvar. texana Vasey. 
= Hilaria belangeri (Steud.) Nash. 


Hubbardochloinae subtrib. Auquier, Bull. Jard. Bot. Belg. 
50: 246. 1980. 


Hymenothecium Lag., Gen. Sp. Pl. 4. 1816. TYPE: 
Hymenothecium tenellum (Cav. ex DC.) Lag. 
= Aegopogon Humb. & Bonpl. ex Willd. 


Hymenothecium quinquesetum Lag., Gen. Sp. Pl. 4. 1816. T- 
Protol.: Habitat in Mex. Iperio, Ludovicus Nee 
{HT:MA]. 

= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Hymenothecium tenellum (Cav. ex DC.) Lag., Gen. Sp. PI. 4. 
1816. Basionym: Cynosurus tenellus Cav. ex DC. 
= Aegopogon tenellus (DC.) Trin. 


Hymenothecium trisetum Lag., Gen. Sp. Pl. 4. 1816. T- 
Protol.: Habit. in Mexico Imperio[HT: MA], cites 
Cynosurus gracilis Cav. H.R.M. L. t. 5, f. 2. 

= Aegopogon cenchroides Humb. & Bonpl. ex Willd. 


Hymenothecium unisetum Lag., Gen. Sp. Pl. 4. 1816. T- 
Protol.: Cult. from seed: H. in Imp. Mexicano, missit 
D. Sessé (HT: MA]. 
= Aegopogon tenellus (DC.) Trin. 


Ipnum Phil., Anales Univ. Chile 36: 211. 1870. TYPE:/pnum 
mendocinum Phil. 
= Leptochloa P. Beauv. 


Ipnum mendocinum Phil., Anales Univ. Chile 36: 211. 1870. 
T-Protol.: Mendoza. T-Specim.: 1 of 1. Argentina: 
Mendoza [HT: SGO; IT: BAA-1452]. 

= Leptochloa dubia (Kunth) Nees. 


Jouvea E. Fourn., Bull. Soc. Roy. Bot. Belgique 15(3): 
475. 1876. TYPE: Jouvea straminea E. Fourn. 
TRTD. BY: Peterson, P. M. 
NHT: Monanthochloinae. NO. ACCPTD. SPP.: 2. 
CNSLTD. LIT.: Weatherwax, P., (1939). 
SYN.: Rhachidospermum Vasey. 


Jouvea pilosa (J. Presl) Scribn., Bull. Torrey Bot. Club 23: 
143. 1896. Basionym: Brizopyrum pilosum J. Presl. 

DST.: CoRi, ElSa, Guat, Hond, Mexi, Nica. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Berendsohn, 
W. G. & A. E. Araniva de Gonzalez (1991): 
Breedlove, D. E. (1986); Davidse, G., M. Sousa S. & 
A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. L6pez-Ferrari, and J. Valdes-Reyna 
(2000); Gould, F. W. & R. Moran (1981); Levin, G. 
A. & R. Moran (1989); McClure, F. A. (1955); 
McVaugh, R. (1983); Molina R., A. (1975); Pohl, R. 
W. & G. Davidse (1994); Pohl, R. W. (1980); 
Shreve, F. & I. L. Wiggins (1964). 

Brizopyrum pilosum J. Presl; Poa preslii Kunth; 
Rhachidospermum mexicanum Vasey. 


SYN.: 


Jouvea straminea E. Fourn., Bull. Soc. Roy. Bot. Belgique 
15(3): 475-476. 1876. T-Protol.: Mexico: in arena 
littorali maris Pacifici prope San Augustin, decembri 
florifera, Liebmann s.n. [HT: BR]. T-Specim.: | of 1. 
Mexico: Oaxaca: prope San Augustin, Oct 1842, 
Liebermann 738 [HT: BR; IT: US-91915 (fragm. ex 
herb, musei botanici Hauniensis)]. 

DST.: CoRi, Ecua, ElSa, Guat, Hond, Mexi, Nica, Pana. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Berendsohn, 
W.G. & A.E. Araniva de Gonzalez (1991); Davidse, 
G., M. Sousa S. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Jorgensen, P. M. & S. Leon- 
Yanez (eds.) (1999); McClure, F. A. (1955); 
McVaugh, R. (1983); Pohl, R. W. & G. Davidse 
(1994); Pohl, R. W. (1980). 


Jouveae trib. Pilg., Nat. Pflanzenfam. (ed. 2) 14d: 168. 1956. 
TYPE: Jouvea E. Fourn. 


Koeleria avenacea (Kunth) Spreng., Syst. Veg. 1: 332. 1825. 
Basionym: Triodia avenacea Kunth. 
= Erioneuron avenaceum (Kunth) Tateoka. 


130 


Catalogue of New World Grasses: II. 


Koeleria multiflora Regel & Herter, Index Sem. (St. 
Petersburg) 23. 1858. T-Protol.: Cult.: Hort. Petrop.: 
seed from Chile, M4. Philippe [T: LE]. 

= Eragrostis peruviana (Jacq.) Trin. 


Koeleria pulchella (Kunth) Spreng., Syst. Veg. 1: 332. 1825. 


Basionym: Triodia pulchella Kunth. 
= Dasyochloa pulchella (Kunth) Willd. ex Rydb. 


Kralikia Coss. & Durieu, Bull. Soc. Bot. France 14: 89. 1867. 
= Tripogon Roem. & Schult. 


Kralikiella Batt. & Trab., Fl. Alg. Monocot. 245. 1895. 
NOTE: nom. superfl. pro Kralikia. 
= Tripogon Roem. & Schult. 


Lamarckia tenella DC., Cat. Pl. Horti Monsp. 120. 1813. T- 
Protol.: Cavanillas ex hort. madr., patriae ignoro. T- 
Specim.: | of 1. Cultivated at Montpelier, Anon. [HT: 
MPU?; IT: US-75926 (fragm.)]. 

= Aegopogon tenellus (DC.) Trin. 


Langsdorffia Regel, Index Sem. (St. Petersburg) 26. 1863. 
NOTE: later hom., non Mart. 1818, insyn. sub  E. 
distichophylla. 
= Eustachys Desv. 


Lappagininae subtrib. Endl., Fl. Poson. 100. 1830. TYPE: 
Lappago Schreb.; nom. illeg. superfl. Tragus Haller. 
NOTE: nom. illeg., genus illeg. 


Lappago Schreb., Gen. Fil. (Ann. Cryptog. Phytopathol. 5:) 
55. 1789. 
NOTE: nom. illeg. superfl. 
= Tragus Haller. 


Lappago aliena Spreng., Neue Entdeck. Pflanzenk. 3: 15. 
1822. T-Protol.: Hab. in Brasilia, Zeyher. 
= Tragus berteronianus Schult. 


Lappago berteroniana Schult. ex Steud., Syn. Pl. Glumac. 1: 
112. 1854. 
NOTE: nom. inval., as syn. 
= Tragus berteronianus Schult. 


Lappago occidentalis Nees ex Hook., Fl. Brit. India 7: 97. 
1896. 
NOTE: nom. inval., as syn. of Tragus racemosus Scop. 
= Tragus berteronianus Schult. 


Lappago racemosa (L.) Honck., Syn. Pl. Germ. 1: 440. 1792. 
Basionym: Cenchrus racemosus L. 
= Tragus racemosus (L.) All. 


Lasiostega Benth., Pl. Hartw. 347. 1857. 
NOTE: nom. nud. 
= Bouteloua Lag. 


Lasiostega humilis Rupr. ex Munro, Pl. Hartw. 347. 1857. 
= Bouteloua dactyloides (Nutt.) Columbus. 


Lasiotrichos Lehm., Del. Sem. Hort. Hamburg. 1834: 7. 
1834. 
NOTE: in syn. sub Fingerhuthia. 
= Fingerhuthia Nees. 


Latipes Kunth, Revis. Gramin. 1: 53. 1829. TYPE: Latipes 
senegalensis Kunth. 
= Leptothrium Kunth. 


Lepeocercis serrata (Retz.) Trin., Fund. Agrost. 203, t. 18. 
1820. Basionym: Andropogon serratum Retz. 


= Chloris virgata Sw. 


Lepiureae trib. Dumort., Observ. Gramin. Belg. 83, 140. 
1824. 
NOTE: nom. inval., orth. var. Leptureae. 


Leptochloa P. Beauv., Ess. Agrostogr. 71. 1812. 
TYPE: Leptochloa virgata (L.) P. Beauv.; LT 
designated by Niles & Chase, Contr. U.S. Natl. 24(6): 
180 (1925); the earlier designation by Nash, Ill. FI. 
North. U.S. Canada, ed. 2, 229 (1919) is ineffective 
according to ING Art. 10.5 *Ex. 6. 

TRTD. BY: Peterson, P.M. & N. Snow. 

NHT: Cynodonteae. NO. ACCPTD SPP.: 17 SUBSP.: 9. 

CNSLTD. LIT.: Lazarides, M., (1980); Nicora, E. G., 
(1995). 

SYN.: Anoplia Steud.; Baldomiria Herter; Diachroa Nutt.; 
Diacisperma Kuntze; Diplachne P. Beauv.: 
Disakisperma Steud.; Ipnum Phil.; Leptostachys G. 
Mey.; Oxydenia Nutt.; Rabdochloa P. Beauv. 


Leptochloa acuminata (Nash) Mohlenbr., Ill. Fl. Iinois 293. 
1973. Basionym: Diplachne acuminata Nash. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Leptochloa albemarlensis B.L. Rob. & Greenm., Amer. J. 
Sci. 50: 145, 1895. T-Protol.: Galapagos Islands: S. 
Albemarle Isl., Jul 1891, Baur 347. T-Specim.: | of 1, 
Ecuador: Galapagos Islands: Isla Isabela, Southern 
Portion, (as Albemarle 1.), Jul 1891,G. Baur 34] [IT: 
US-91025, US-91027 (fragm.)]. 

= Trichoneura lindleyana var. albemarlensis (B.L. Rob. 
& Greenm.) Reeder & C. Reeder. 


Leptochloa aquatica Scribn. & Merr., Bull. Div. Agrostol., 
U.S.D.A. 24: 26. 1901. T-Protol.: Mexico: Morelos: in 
shallow water near Cuernavaca, alt. 1700 m, 22 Aug 
1897, C.G. Pringle 6664 [HT: US]. T-Specim.: | of 1. 
Mexico: Morelos: in shallow water near Cuernavaca, 
5200 ft, 22 Aug 1897, C.G. Pringle 6664 [HT: US- 
316891; IT: B, CM, GH, L, M, MCU, MEXU, MO- 
2972338, NY-19495, P, TAES]. 

DST.: Mexi. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983). 


Leptochloa arabica (Jacq.) Kunth, Revis. Gramin, 1: 91, 
1829. Basionym: Dinebra arabica Jacq. 
= Dinebra retroflexa (Vahl) Panz. 


Leptochloa attenuata (Nutt.) Steud., Syn. Pl. Glumac. |: 209. 

1854. Basionym: Oxydenia attenuata Nutt. 

= Leptochloa panicea’ subsp. mucronata 
Nowack. 


(Michx.) 


Leptochloa barbata Desy. ex Killeen, Ann. Missouri Bot. 
Gard. 77(1): 164. 1990. 
NOTE: nom. inval., as syn. of Leptochloa virgata (L.) P. 
Beauv. 
= Leptochloa virgata (L.) P. Beauv. 


Leptochloa barbata (Desv.) Parodi ex Nicora, Hickenia 
2(19): 91. 1993. Basionym: Eleusine barbata Desv. 
DST.: Arge, Boli, Braz, Para. 
LVL. ACCPTC:: 1. 
ALSO ACCPTD. BY: Nicora, E. G. (1995). 


Catalogue of New World Grasses: II. 


131 


SYN.: Eleusine barbata Desv. 


Leptochloa brachiata Steud., Syn. Pl. Glumac. 1: 209. 1854. 
T-Protol.: Guadeloupe, Duchaissing s.n. [HT: P], as 
spelled by Steudel. T-Specim.: 1 of |. Guadeloupe, 
Duchassaing s.n. {HT: P (fragm., US)], as spelled in 
Authors of Plant Names (1992). 

= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Leptochloa brandegei (Vasey) Hitche., U.S.D.A. Bur. PI. 
Industr. Bull. 33: 21. 1903. Basionym: Diplachne 
brandegei Vasey. 

= Enteropogon brandegei (Vasey) Clayton. 


Leptochloa burchellii Munro ex DO6ll, Fl. Bras. 2(3): 93. 
1878. T-Protol.: Habitat in prov. S. Paulo, Burchell 
4462. T-Specim.: | of 1. Brazil: S40 Paulo, Burchell 
4462 {HT: K; IT: BAA (fragm.), BR, US (fragm. ex 
K), US (fragm. ex BR)]. 

= Gymnopogon burchellii (Munro ex Déll) Ekman. 


Leptochloa calycina (Roxb.) Kunth, Revis. Gramin. 1: 91. 
1829. Basionym: Eleusine calycina Roxb. 
= Dinebra retroflexa var. retroflexa. 


Leptochloa carinata Hack. 
NOTE: entry error for Diplachne carinata (Griseb.) Hack. 
= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Leptochloa chloridiformis (Hack. ex Stuck.) Parodi, Physis 
(Buenos Aires) 4: 184, f. 5-6. 1918. Basionym: 
Diplachne chloridiformis Hack. ex Stuck. 

NOTE: typographical error as Letopchloa. 

DST.: Arge, Para, USA, Urug. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Burkart, A. 
(1969); Cabrera, A. L. (1970); Correll, D. S. & M. C. 
Johnston (1970); Hitchcock, A. S. (1951); Nicora, E. 
G. (1995); Zuloaga, F. O., E. G. Nicora, Z. E. Ragolo 
de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 

Baldomiria chloridiformis (Hack. ex Stuck.) Herter; 
Diplachne chloridiformis Hack. ex Stuck. 


SYN.: 


Leptochloa decipiens (R. Br.) Stapf ex Maiden, Agric. Gaz. 
New South Wales 20: 307. 1909. Basionym: Poa 
decipiens R. Br. 

DST.: USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Blake, S. T. (1972); Lazarides, M. 
(1980). 

SYN.: Eragrostis decipiens (R. Br.) Steud.; Leptochloa 
decipiens (R. Br.) Druce; Poa decipiens R. Br. 


Leptochloa decipiens (R. Br.) Druce, Rep. Bot. Exch. Club 
1916: 632. 1917. Basionym: Poa decipiens R. Br. 
NOTE: iso. et hom. 
E. NAME: Leptochloa decipiens (R. Br.) Stapf ex Maiden. 
= Leptochloa decipiens (R. Br.) Stapf ex Maiden. 


Leptochloa decipiens subsp. peacockii (Maiden & Betche) 
N. Snow, Novon 8(1): 78. 1998. Basionym: Diplachne 
peacockii Maiden & Betche. 

DST.: USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Snow, N. & B. K. Simon (1999); 
Snow, N. (1998). 

SYN.: Diplachne peacockii Maiden & Betche; Leptochloa 
peacockii (Maiden & Betche) Domin. 


Leptochloa digitaria (Kunth) Nees, Fl. Bras. Enum. Pl. 2: 
433. 1829. Basionym: Chloris digitaria Kunth. 
= Leptochloa virgata (L.) P. Beauv. 


Leptochloa digitata (R. Br.) Domin, Biblioth. Bot. 85: 379. 

1915. Basionym: Poa digitata R. Br. 

DST.; USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Blake, S. T. (1972); Lazarides, M. 
(1980); Snow, N. & B. K. Simon (1999), 

SYN.: Eleusine digitata (R. Br.) Spreng.; Leptochloa 
subdigitata Trin. ex Steud.; Poa digitata R. Br. 


Leptochloa digitatiformis Beetle, Phytologia 52(1): 14. 1982. 
T-Protol.: Mexico: Sonora: Ejido km 47, matorral 
subinerme, 19 Aug 1981, Bernal & Cuadra s.n., 
herbarium not indicated by the author. 

= Leptochloa dubia (Kunth) Nees. 


Leptochloa divaricatissima S.T. Blake, Contr. Queensland 
Herb. 14: 8, f. 5. 1972. T-Protol.: Australia: 
Queensland: Darling Downs Distr.: Goondiwindi, 
Blake 10517 (HT: BRI; IT: K, NSW]. T-Specim.: 1 of 
1. Australia: Queensland: Goondiwindi, 8.7. Blake 
/0517 (HT: BRI-131665-6; IT: CANB, K, MO, NSW, 
PRE]. 

DST.: USA, 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Lazarides, M. (1980); Snow, N. & B. 
K. Simon (1999). 


Leptochloa domingensis (Jacq.) Trin., Fund. Agrost. 133. 
1820. Basionym: Cynosurus domingensis Jacq. 
= Leptochloa virgata (L.) P. Beauv. 


Leptochloa dubia (Kunth) Nees, Syll. Pl. Nov. 1: 4. 1824. 
Basionym: Chloris dubia Kunth. 
DST.: Arge, Boli, Cari, Chil, Colo, Ecua, Hond, Mexi, Peru, 
USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Anderson, D. E. (1974); Anonymous 
(1986); Beetle, A. A. (1977); Breedlove, D. E. 
(1986); Correll, D. S. & M. C. Johnston (1970); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Foster, R. C. 
(1958); Gould, F. W. & R. Moran (1981); Hitchcock, 
A. S. (1927); Hitchcock, A. S. (1951); Jorgensen, P. 
M. & S. Leon-Yanez (eds.) (1999); Long, R. W. & 
O. Lakela (1971); McVaugh, R. (1983); Pohl, R. W. 
& G. Davidse (1994); Renvoize, S. A. (1998); 
Shreve, F. & I. L. Wiggins (1964); Small, J. K. 
(1933); Snow, N. & B. K. Simon (1999); Tovar, O. 
(1993); Wunderlin, R. P. (1998). 

Chloris dubia Kunth; Diplachne dubia (Kunth) 
Scribn.; Diplachne dubia var. aristata Vasey; 
Diplachne dubia var. dubia; Diplachne dubia var. 
humboldtiana Kuntze; Diplachne dubia — var. 
kurtziana Kuntze; Diplachne dubia var. pringleana 
Kuntze; Diplachne mendocina (Phil.) Kurtz: 
Diplachne patens (J. Presl) E. Desv.; Diplachne 
pringlei Vasey ex Beal; Disakisperma mexicana 
Steud.; Eragrostis mendozina (Phil.) Jedwabn.; 
Festuca obtusiflora Willd. ex Spreng.; Jpnum 
mendocinum Phil.; Leptochloa digitatiformis Beetle; 
Leptochloa dubia subsp. dubia; Leptochloa dubia 
subsp. patens (J. Presl) Covas & Steibel: Leptochloa 
dubia var. humboldtiana (Kuntze) Beetle; Leptochloa 


SYN.: 


132 


Catalogue of New World Grasses: II. 


dubia var. pringleana (Kuntze) Scribn. & Merr.; 
Leptochloa obtusiflora Trin. ex Steud.; Leptochloa 
patens (J. Presl) Kunth; Leptochloa pringlei Vasey ex 
Beal; Leptostachys dubia (Kunth) G. Mey.; Molinia 
retusa Griseb. ex E. Fourn.; Rabdochloa dubia 
(Kunth) Kuntze ex Stuck.; Schismus patens J. Presl; 
Sieglingia dubia (Kunth) Kuntze ex Stuck.; Uralepis 
brevicuspidata Buckley. 


Leptochloa dubia Kuntze subsp. dubia. 
= Leptochloa dubia (Kunth) Nees. 


Leptochloa dubia var. humboldtiana (Kuntze) Beetle, 
Phytologia 54(1): 4. 1983. Basionym: Diplachne dubia 
var. humboldtiana Kuntze. 

= Leptochloa dubia (Kunth) Nees. 


Leptochloa dubia subsp. patens (J. Presl) Covas & Steibel, 
Apuntes Fl. Pampa 61: 243. 1981. Basionym: 
Schismus patens J. Presl. 

= Leptochloa dubia (Kunth) Nees. 


Leptochloa dubia var. pringleana (Kuntze) Scribn. & Merr., 
Bull. Div. Agrostol, U.S.D.A. 24: 27. 1901. 
Basionym: Diplachne dubia var. pringleana Kuntze. 

= Leptochloa dubia (Kunth) Nees. 


Leptochloa fascicularis (Lam.) A. Gray, Manual 588. 1848. 
Basionym: Festuca fascicularis Lam. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Leptochloa fascicularis var. acuminata (Nash) Gleason, 
Phytologia 4(1): 21. 1952. Basionym: Diplachne 
acuminata Nash. 

= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Leptochloa fascicularis(Lam.) A. Gray var. fascicularis. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Leptochloa fascicularis var. 
Gleason, Phytologia 4(1): 21. 
Diplachne maritima E.P. Bicknell. 

= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow, 


maritima (E.P. Bicknell) 
1952. Basionym: 


Leptochloa filiformis (Pers.) P. Beauy., Ess. Agrostogr. 71, 
163, 166. 1812. Basionym: Eleusine filiformis Pers. 
= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Leptochloa filiformis fo. attenuata (Nutt.) F.C. Gates, Annot. 
List Pl. Kansas 391: 130. 1940. Basionym: Oxydenia 
attenuata Nutt. 

NOTE: basionym not explicitly cited, basionym author cited 
only as Stuedel; from Lepfochloa attenuata (Nutt.) 
Steudel. 

= Leptochloa  panicea 
Nowack. 


subsp. mucronata (Michx.) 


Leptochloa filiformis var. attenuata (Nutt.) Steyerm. & 
Kucera, Rhodora 63(745): 26. 1961. Basionym: 
Oxydenia attenuata Nutt. 

NOTE: also isonym, Grass. Missouri 138 (1961). 

= Leptochloa panicea subsp. mucronata (Michx.) 
Nowack. 


Leptochloa filiformis var. humilior J. Pres|, Relig. Haenk. 1: 
288. 1830. 
= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Leptochloa filiformis var. pulchella’ (Scribn.) Beetle, 
Phytologia 49(1): 40. 1981. Basionym: Leptochloa 
mucronata var. pulchella Scribn. 


= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Leptochloa filiformis var. remota Doll, Fl. Bras. 2(3): 93. 
1878. T-Protol.: "in Brasilia adhuc non observata.": 
[up to present time not observed in Brazil], Gray Herb 
cards give as [var.]. 
= Status not yet determined. 


Leptochloa floribunda DO6ll, Fl. Bras. 2(3): 89, t. 26. 1878. T- 
Protol.: Brasilia habitat ad ripas fluminis Amazonam 
inter Manaos et Santarem, Spruce s.n. T-Specim.: | of 
1. Brazil: Amazonas: inter Manaus et Santaren, R. 
Spruce 908 [HT: M; IT: BAA-1508 (fragm. ex M), 
GH, US-88700 (fragm. ex M)]. 

= Leptochloa panicoides (J. Pres!) Hitche. 


Leptochloa fusca (L.) Kunth, Revis. Gramin. 1: 91. 1829. 
Basionym: Festuca fusca L. 

DST.: Arge, Beli, Boli, Braz, Cari, Chil, CoRi, Ecua, Hond, 
Meni, Nica, Para, Peru, USA, Urug. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Phillips, S. (1995). 

SYN.: Centotheca malabarica (L.) Merr.; Digitaria 
malabarica (L.) Roem. & Schult.; Diplachne fusca 
(L.) P. Beauv. ex Roem. & Schult.; Diplachne fusca 
(L.) P. Beauv. ex Stapf; Diplachne fusca (L.) P. 
Beauv. ex Stuck.; Diplachne malabarica (L.) Merr.; 
Diplachne reptatrix (L.) Druce; Festuca fusca L.; 
Festuca reptatrix L.; Hemigymnia malabarica (L.) 
Henrard; Leptochloa malabarica (L.) Veldkamp; 
Ottochloa malabarica’ (L.) Dandy; Panicum 
malabaricum (L.) Merr.; Poa malabarica L.; 
Uralepis fusca (L.) Steud. 


Leptochloa fusca subsp. fascicularis (Lam.) N. Snow, 
Novon &(1): 78. 1998. Basionym: Festuca fascicularis 
Lam. 

DST.: Arge, Beli, Boli, Braz, Cana, Cari, Chil, CoRi, Ecua, 
Hond, Mexi, Nica, Para, Peru, USA, Urug. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Snow, N. (1998). 

SYN.: Cynodon fascicularis (Lam.) Raspail; Diachroa 
procumbens (Muhl.) Nutt.; Diplachne acuminata 
Nash; Diplachne fascicularis (Lam.) P. Beauv.; 
Diplachne maritima E.P. Bicknell; Diplachne patens 
E. Fourn.; Diplachne patens E. Fourn. ex Hemsl.; 
Diplachne procumbens (Muhl.) Nash; Diplachne 
tracyi Vasey, Festuca aquatica Bose ex Roem. & 
Schult.; Festuca  clandestina Muhl.; Festuca 
clandestina| Muhl.; Festuca fascicularis  Lam.; 
Festuca polystachya Michx.,; Festuca procumbens 
Muhl.; Festuca procumbens Muhl.; Festuca 
prostrata Muhl. ex Scribn. & Merr.; Festuca texana 
Steud.; Festuca thouini  Steud.; ~—Leptochloa 
acuminata (Nash) Mohlenbr.; Leptochloa 
Jascicularis (Lam.) A. Gray; Leptochloa fascicularis 
var. acuminata (Nash) Gleason;  Leptochloa 
fascicularis var. fascicularis, Leptochloa fascicularis 
var. maritima (E.P. Bicknell) Gleason; Leptochloa 
polystachya (Michx.) Kunth; Leptochloa  tracyi 
(Vasey) Beal; Tridens veralensis Cat. Guerra; 
Tridens virens Nees; Uralepis virens (Nees) Kunth; 
Uralepsis composita Buckley. 


Leptochloa fusca (L.) Kunth subsp. fusca. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Snow, N. & B. K. Simon (1999). 


Catalogue of New World Grasses: II. 


133 


Leptochloa fusca subsp. uninervia (J. Presl) N. Snow, 
Novon 8(1): 79. 1998. Basionym: Megastachya 
uninervia J. Presl. 

DST.: Arge, Beli, Boli, Braz, Cari, Chil, Colo, CoRi, Ecua, 
Hond, Mexi, Nica, Para, Peru, USA, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Snow, N. & B. K. Simon (1999); 

Snow, N. (1998). 

Atropis carinata Griseb.; Brizopyrum uninervium (J. 

Presl) E. Fourn.; Diplachne carinata (Griseb.) Hack.; 

Diplachne imbricata (Thurb.) Scribn.; Diplachne 

procumbens Arechav., Diplachne tarapacana Phil.; 

Diplachne uninervia (J. Presl) Parodi; Diplachne 

uninervia fo. abbreviata Parodi; Diplachne uninervia 

fo. uninervia; Diplachne uninervia var. procumbens 

(Arechav.) Parodi; Diplachne — uninervia vat. 

uninervia, Eragrostis uninervia (J. Presl) Steud.; 

Leptochloa_ carinata Hack.; Leptochloa imbricata 

Thurb.; Leptochloa uninervia (J. Presl) Hitchc. & 

Chase; Leptochloa virletii E. Fourn.; Leptochloa 

virletii E. Fourn. ex Hemsl.; Megastachya uninervia 

J. Presl; Poa uninervia (J. Presl) Kunth; Puccinellia 

carinata (Griseb.) Ponert; Rabdochloa imbricata 

(Thurb.) Kuntze; Tridens duartei Cat. Guerra, 

Tridens verticillatus Meyen; Uralepis anderssonii F. 

Aresch.; Uralepis verticillata (Nees & Meyen) 

Steud.; Uralepsis verticillata (Nees & Meyen) Steud. 


SYN.: 


Leptochloa gracilis Nees, Syll. Pl. Nov. 1: 4. 1824. 
REPLACED NAME: Chloris gracilis Kunth. 
BLOCKING NAME: Chloris gracilis P. Durand. 
= Leptochloa virgata (L.) P. Beauv. 


Leptochloa halei (Nash) Scribn, & Merr., Bull. Div. 
Agrostol., U.S.D.A. 24: 27. 1901. Basionym: 
Diplachne halei Nash. 

= Leptochloa panicoides (J. Pres!) Hitche. 


Leptochloa hirta Nees ex Steud., Syn. Pl. Glumac. 1: 209. 
1854. T-Protol.: [Ecuador]: Galapagos Islands: Ins. 
Albermarle, Gallipogorum. T-Specim.: 1 of 1. 
Galapagos Islands: Isla Isabela: (as Albermarle 1.), 
1853, NJ Anderson [T: IT:US-91034 (fragm.)]. 

= Bouteloua disticha (Kunth) Benth. 


Leptochloa hookeri (Andersson) Andersson, Freg. Eugenies 
Resa, Bot. 2: 51. 1861. Basionym: Trichoneura 
hookeri Andersson. 

NOTE: nom. illeg. 
CORRECT NAME: Calamagrostis pumila Hook. f. 
= Trichoneura lindleyana var. lindleyana. 


Leptochloa humilis Nees, Syn. P|. Glumac. 1: 210. 1854. T- 
Protol.: Mexico[T: Herb. Lindley]. 
= Status not yet determined. 


Leptochloa imbricata Thurb., Bot. California 2: 293. 1880. T- 
Specim.: 1 of 2. USA: California: San Diego Co., 
Larkin's Station, S part of Co., 1875, E. Palmer 404 
[ST: NY-19497; IST: US-88701 (fragm.)]. 2 of 2. 
USA: Arizona: Fort Yuma, Major Thomas s.n. [ST]. 

= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Leptochloa langloisii Vasey, Bull. Torrey Bot. Club 12: 7. 
1885. T-Protol.: [USA]: Louisiana, A.B. Langlois s.n. 
T-Specim.: 1 of 1. USA: Louisiana, 1883, 4. B. 
Langlois s.n. [HT: US-78811, IT: NY-19496 [1 Sep 
1884]]. 


= Leptochloa scabra Nees. 


Leptochloa liebmannii E. Fourn. ex Hemsl., Biol. Cent.- 
Amer., Bot. 3: 565. 1885. 
NOTE: nom. nud. 
CORRECT NAME: Leptochloa liebmannii E. Fourn. 
= Leptochloa scabra Nees. 


Leptochloa liebmannii E. Fourn., Mexic. Pl. 2: 147. 1886. T- 
Specim.: | of 2. Mexico: Antigua, Feb, F.M. Liebmann 
248 (ST: US-899151 (fragm.)]. 2 of 2. Absque loco, 
Liebm. 244 [ST]. 
= Leptochloa scabra Nees. 


Leptochloa lindleyana Kunth, Enum. PI. 1: 525. 1833. T- 
Protol.: {Ecuador}: Insula  Albermarle, una ex 
Gallapogos [Galapagos]. T-Specim.: 1 of 2. Ecuador: 
Galapagos Islands: Albemare Island, Macrae s.n. [HT: 
B; IT: CGE, K]. 2 of 2. Galapagos Islands, 1825, 
J. Macrae [T: IT:US-91035 (fragm.)]. 

= Trichoneura lindleyana (Kunth) Ekman. 


Leptochloa longa Griseb., Fl. Brit. W. 1. 538. 1864. T- 
Protol.; Trinidad, Crueger s.n. T-Specim.: | of 2. 
Trinidad: without locality, Crueger [Bot. Gard. Herb. 
3359] [HT: K; IT: US-899223]. 2 of 2. Trinidad and 
Tobago: Trinidad I., A.H.R.Grisebach 3359 [IT: US- 
822634]. 

DST.: Cari, Pana. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Hitchcock, A. S. 
(1936); Pohl, R. W. & G. Davidse (1994). 

SYN.: Rhabdochloa longa (Griseb.) Kuntze. 


Leptochloa malabarica (L.) Veldkamp, Blumea 19(1): 64. 
1971. Basionym: Poa malabarica L. 
NOTE: nom. rej., basionym rej. [ICBN art. 56.2]. 
= Leptochloa fusca (L.) Kunth. 


Leptochloa mexicana Scribn., Proc. Acad. Nat. Sci. 
Philadelphia 1891: 302. 1891. T-Protol.: Mexico: [San 
Luis Potosi]: about dry ledges, Tamasopo Cafion, 28 
Sep 1890, Pringle 3252. T-Specim.: | of 1. Mexico: 
San Luis Potosi: limestone ledges, Tamasopo Canyon, 
28 Sep 1890, C.G. Pringle 3252 [HT: NY; IT: MEXU, 
MO-2369804, MO-2369805, MO-3727962, NY, US- 
481015, US-78809, US-899311]. 
NOTE: as Leptochloa "Mexicana". 
= Gouinia mexicana (Scribn.) Vasey. 


Leptochloa monticola Chase, J. Wash. Acad. Sci. 17: 73, f. 

2. 1927. T-Protol.: Haiti: on summit of Pio de Brouet, 
in the vicinity of Purcy, 1300 m, 13 Jun 1920, E.C. 
Leonard 4751 [HT: US-1077272]. T-Specim.: 1 of 1. 
Haiti: Ouest: vicinity of Furcy, 13 Jun 1920, E.C. 
Leonard 475] [HT: US-1077272; IT: NY-19492, US- 
1298292]. 

DST.: Cari. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Catasus 

Hitchcock, A. S. (1936). 


Guerra, L. (1985); 


Leptochloa mucronata (Michx.) Kunth, Revis. Gramin. 1: 91. 
1829. Basionym: Eleusine mucronata Michx. 
= Leptochloa panicea subsp. mucronata (Michx.) 
Nowack. 


Leptochloa mucronata (Michx.) Kunth var. mucronata. 


134 


Catalogue of New World Grasses: II. 


= Leptochloa subsp. mucronata (Michx.) 


Nowack. 


panicea 


Leptochloa mucronata var. multiflora Eggers, Vidensk. 
Meddel. Dansk Naturhist. Foren. Kjobenhavn 8: 151. 
1876. 
NOTE: nom. nud. 
= Leptochloa 
Nowack. 


panicea subsp. mucronata (Michx.) 


Leptochloa mucronata var. pulchella Scribn., Bull. Torrey 
Bot. Club 9: 147. 1882. T-Protol.: USA: Arizona: 
Santa Cruz Valley, near Tucson, July, Pringle s.n. T- 
Specim.: | of 1, USA: Arizona: Santa Cruz Valley, Jul 
1881, C.G. Pringle s.n. |HT: US-78810]. 

= Leptochloa’ panicea’ subsp. mucronata (Michx.) 
Nowack. 


Leptochloa mutica Steud., Syn. Pl. Glumac. 1: 208. 1854. T- 
Protol.: Surinam, Kappler Hrbr. 1553, "L. virgata 
Hochst.". T-Specim.: | of 1. Surinam, Feb 1884-Apr 
1844, A. Kappler 1553 (HT: P; IT: US-3266093 
(fragm.)]. 

= Leptochloa virgata (L.) P. Beauv. 


Leptochloa nealleyi Vasey, Bull. Torrey Bot. Club 12: 7. 
1885. T-Protol.: USA: Texas, G.C. Nealley s.n. T- 
Specim.: | of 1. USA: Texas, 1884, G.C. Nealley s.n. 
[HT: US-78808, IT: MCU, NY-798 14]. 

DST.: Cari, Mexi, USA. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 

(1977); Catasus Guerra, L. (1985); Catasus Guerra, 

L. (1997); Correll, D. S. & M. C. Johnston (1970); 

Davidse, G., M. Sousa S. & A. O. Chater (editores 

generales) (1994); Espejo Serna, A., A. R. Lopez- 

Ferrari, and J, Valdes-Reyna (2000); Godfrey, R. K. 

& J. W. Wooten (1979); Hitchcock, A. S. (1936); 

Hitchcock, A. S. (1951); Liogier, H. & L. Martorell 

(1982); Pohl, R. W. & G. Davidse (1994), 

Leptochloa stricta E. Fourn.; Leptochloa stricta E. 

Fourn. ex Hemsl. 


SYN.:: 


Leptochloa obtusiflora Trin, ex Steud., Nomencl. Bot. (ed. 2) 
2: 30. 1841. 
NOTE: nom. inval., as syn. of Leptochloa dubia (Kunth) 
Nees. 
= Leptochloa dubia (Kunth) Nees. 


Leptochloa palmeri Vasey ex Scribn., Bull. Div. Agrostol., 
U.S.D.A. 4: 10. 1897. T-Specim.: 1 of 1. Mexico: 
Colima, | Dee 1890-31 Dec 1890, E. Palmer 1087 
[OM: US-8993 18]. 

NOTE: nom. inval., as synonym of Gouinia virgata (J. S. 
Presl) Scribner. 
= Gouinia virgata (J. Presl) Scribn. 


Leptochloa panicea (Retz.) Ohwi, Bot. Mag. (Tokyo) 
55(655): 311. 1941. Basionym: Poa panicea Retz. 
DST.: Arge, Beli, Boli, Braz, Cari, CoRi, Ecua, ElSa, Guat, 

Hond, Mexi, Nica, Pana, Peru, USA, Vene. 

ALSO ACCPTD. BY: Bor, N. L. (1960); Fosberg, F. R. & 
M.-H. Sachet (1982); Gibbs Russell, G. E, W. G. 
Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987); Lazarides, M. (1980); 
Lindeman, J.C. & A. L. Stoffers (1963). 

SYN.: Poa panicea Retz. 


Leptochloa panicea subsp. brachiata (Steud.) N. Snow, 
Novon 8&(1): 79. 1998. Basionym: Leptochloa 
brachiata Steud. 

DST.: Arge, Beli, Boli, Braz, Cari, Colo, CoRi, Ecua, EISa, 
Guat, Hond, Mexi, Nica, Pana, Para, Peru, USA, 
Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Snow, N. & B. K. Simon (1999); 

Snow, N. (1998). 

Aira panicea Willd. Ex Steud.; Eleusine filiformis 

Pers.; Eleusine sparsa Muhl.; Eleusine stricta Willd. 

ex Steud.; Festuca filiformis Lam., Leptochloa 

brachiata Steud.; Leptochloa filiformis (Pers.) P. 

Beauv.; Leptochloa filiformis var. humilior J. Presl: 

Leptochloa filiformis var. pulchella (Scribn.) Beetle; 

Leptochloa paniculata’ E. Fourn.; Leptochloa 

pellucidula  Steud.; Leptochloa pilosa Scribn.; 

Leptostachys filiformis (Lam.) G. Mey.,; Oxydenia 

filiformis Nutt. ex B.D. Jacks. 


SYN:: 


Leptochloa panicea subsp. mucronata Nowack, Rheedea 
4(2): 88. 1994, 

NOTE: nom. inval., without 
basionym not cited. 

= Leptochloa panicea 
Nowack. 


Latin description, and 


subsp. mucronata (Michx.) 


Leptochloa panicea subsp. mucronata (Michx.) Nowack, 
Rheedea S(1): 93. 1995. Basionym: Eleusine 
mucronata Michx. 

DST.: USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Snow, N. (1998). 

SYN.: Eleusine elongata Willd. ex Steud.; Eleusine 
mucronata Michx.; Leptochloa attenuata (Nutt.) 
Steud.; Leptochloa filiformis fo. attenuata (Nutt.) 
F.C. Gates; Leptochloa filiformis var. attenuata 
(Nutt.) Steyerm. & Kuéera, Leptochloa mucronata 
(Michx.) Kunth; Leptochloa mucronata — yar. 
mucronata, Leptochloa mucronata var. multiflora 
Eggers; Leptochloa mucronata var.pulchella Scribn.; 
Leptochloa panicea subsp. mucronata Nowack: 
Rabdochloa mucronata (Michx.) P. Beauv. 


Leptochloa panicea (Retz.) Ohwi subsp. panicea. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Snow, N. & B. K. Simon (1999). 


Leptochloa panicoides (J. Presl) Hitche., Amer. J. Bot. 
21(3): 137. 1934. Basionym: Megastachya panicoides 
J. Presl. 

DST.: Braz, Cari, Colo, CoRi, Guat, Hond, Mexi, Peru, 
USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Correll, D. S. 
& M. C. Johnston (1970); Cowan, C. P. (1983); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Gleason, H. A. 
& A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Godfrey, R. K. & J. W. Wooten 
(1979); Gould, F. W. & R. Moran (1981): Hitchcock, 
A. S. (1951); Liogier, H. & L. Martorell (1982); 
McClure, F. A. (1955); McVaugh, R. (1983); Molina 
R., A. (1975); Pohl, R. W. & G. Davidse (1994): 
Tovar, O. (1993); Wunderlin, R. P. (1998). 


Catalogue of New World Grasses: II. 


135 


SYN.: Diplachne halei Nash; Diplachne panicoides (J. 
Presl) McNeill; Eragrostis panicoides (J. Presl) 
Steud.; Leptochloa floribunda Déll; Leptochloa halei 
(Nash) Scribn. & Merr.; Megastachya panicoides J. 
Presl; Poa panicoides (J. Pres!) Kunth. 


Leptochloa paniculata E. Fourn., Bull. Soc. Bot. France 27: 
296. 1880. T-Protol.: Nicaragua: absque loco, M. P. 
Levy 1079 [HT: P]. T-Specim.: 1 of 1. Nicaragua: 
Granada: absque loco, Fall 1869, MP. Levy 1079 [IT: 
US-91036 (fragm.)]. 
= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Leptochloa patens (J. Presl) Kunth, Enum. Pl. 1: 271. 1833. 
Basionym: Schismus patens J. Presl. 
NOTE: also Revis. Gram. 3: Suppl. 1834 p. (as in 1835 ed.) 
XXII. 
= Leptochloa dubia (Kunth) Nees. 


Leptochloa peacockii (Maiden & Betche) Domin, Biblioth. 
Bot. 85: 379. 1915. Basionym: Diplachne peacockii 
Maiden & Betche. 

= Leptochloa decipiens subsp. peacockii (Maiden & 
Betche) N. Snow. 


Leptochloa pellucidula Steud., Syn. Pl. Glumac. 1: 209. 
1854. T-Protol.: Panama, Duchaissing s.n. [HT: P]. T- 
Specim.: | of 1. Panama, Duchassaing s.n. [HT: P]. 

= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Leptochloa perennis Hack., Informe Anual Estac. Centr. 
Agron. Cuba |: 411. 1906. T-Protol.: Cuba: Prov. 
Santa Clara prope La Magdalena, Cayamas, C.F. 
Baker 4617. T-Specim.: 1 of 1. Cuba: Las Villas 
[Santa Clara], no date, C.F. Baker 4617 (HT: W; IT: 
US-88698 (fragm.)]. 

= Leptochloa virgata (L.) P. Beauv. 


Leptochloa pilosa Scribn., Cire. Div. Agrostol. U.S.D.A. 32: 
9. 1901. T-Protol.: USA: Texas: Travis Co.: in sandy 
soil, Dappan, Sep 1891,/.E. Bodin 294, T-Specim.: 1 
of 1. USA: Texas: Travis Co.: Dappan, in sandy soil, 
Sep 1891, AE. Bodin 294 [HT: US-78807]. 
= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Leptochloa plectostachya K. Schum., Pflanzenw. Ost-Afrikas 
112. 1895, T-Specim.: 1 of 1. Tanzania, Volkens 477 
[HT: B]. 
= Cynodon plectostachyus (K. Schum.) Pilg. 


Leptochloa polygama (E. Fourn.) Beal, Grass. N. Amer. 2: 
437. 1896. Basionym: Gouinia polygama E. Fourn. 
= Gouinia virgata (J. Presl) Scribn. 


Leptochloa polystachya (Michx.) Kunth, Revis. Gramin. 1: 
91. 1829, Basionym: Festuca polystachya Michx. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Leptochloa pringlei Vasey ex Beal, Grass. N. Amer. 2: 436. 
1896. T-Protol.: USA: Arizona, 1884, Pringle s.n. T- 
Specim.: 1 of 1. USA: Arizona: Pima Co.: Sierra 
Tuscon, 27 Apr 1884, C.G. Pringle 13 [IT: US- 
78806], 5. 

= Leptochloa dubia (Kunth) Nees. 


Leptochloa procera Nees, Syll. Pl. Nov. 1: 2. 1824. T- 
Protol.: Brasilia. T-Specim.: | of 1. Brazil, M.A.P. 
Wied-Neuwied 244 (HT: B; IT: US-3266091 (fragm.)]. 

DST.: Arge, Boli, Braz, Cari, Para, Vene. 
LVL. ACCPTC.: 1. 


ALSO ACCPTD. BY: Dll, J. C. (1878); Nicora, E. G. 
(1995): Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 


(1994). 

Leptochloa procera var. major Brongn., Voy. Monde 
Phan(8): 48. 1831. 

= Status not yet determined. 

Leptochloa procera vat. minor Brongn., Voy. Monde 


Phan(8): 48. 1831. T-Protol.: Peru: environs de Lima. 
= Status not yet determined. 


Leptochloa rigida Munro ex Vasey, Grass. U.S. 35. 1883. 
NOTE: nom. inval. as syn. of Diplachne rigida Vasey. 
= Eragrostis sessilispica Buckley. 


Leptochloa rigida Munro ex Vasey, U.S.D.A. Div. Bot. Bull. 
12(2): t. 44. 1891. 
= Eragrostis sessilispica Buckley. 


Leptochloa scabra Nees, Fl. Bras. Enum. Pl. 2: 435. 1829. 
T-Protol.: Brasilia: Provinciae Paraensis: in  ripa 
fluminum Amazonum, Tagipurie et Tocantins, Martius 
s.n. T-Specim.: 1 of 1. Brazil: Para, K.F.P. von 
Martius s.n. [HT: M; IT: BAA-1514 (fragm. ex M), 
US-88699 (fragm. ex M)]. 

NOTE: alternate title Agrost. Bras. 435. 

DST.: Arge, Boli, Braz, Cari, Colo, CoRi, Ecua, ElSa, 

FrGu, Guat, Guya, Hond, Mexi, Nica, Pana, Para, 
Peru, Suri, USA, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Berendsohn, 
W. G. & A. E. Araniva de Gonzalez (1991); Brako, 
L. & J. L. Zarucchi (1993); Breedlove, D. E. (1986); 
Cowan, C. P. (1983); Davidse, G., M. Sousa S. & A. 
O. Chater (editores generales) (1994); Doll, J. C. 
(1878); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Hitchcock, A. S. (1927); 
Hitchcock, A. S. (1936); Hitchcock, A. S. (1951): 
Jorgensen, P. M. & S. Leon-Yanez (eds.) (1999); 
Judziewicz, E. J. (1990); Killeen, T. J. (1990); 
Liogier, H. & L. Martorell (1982); Macbride, J. F. 
(1936); McClure, F. A. (1955); McVaugh, R. (1983); 
Molina R., A. (1975); Pohl, R. W. & G. Davidse 
(1994); Pohl, R. W. (1980); Renner, S. S., H. Balslev 
& L. B. Holm-Nielsen (1990); Renvoize, S. A. 
(1998); Snow, N. & M. G. Lelong (1991). 

Diplachne scabra (Nees) Nicora; Leptochloa 
langloisii Vasey; Leptochloa liebmannii E. Fourn.; 
Leptochloa liebmannii E. Fourn. ex Hemsl. 


SYN:: 


Leptochloa spicata (Nees) Scribn., Proc. Acad. Nat. Sci. 
Philadelphia 1891: 304. 1891. Basionym: Bromus 
spicatus Nees. 

NOTE: basionym not cited directly. 

= Tripogon spicatus (Nees) Ekman. 


Leptochloa stricta E. Fourn. ex Hemsl., Biol. Cent.-Amer., 
Bot. 3: 566. 1885. 
NOTE: nom. nud. 
E. NAME: Leptochioa stricta Trin. ex Steud. 
= Leptochloa nealleyi Vasey. 


Leptochloa stricta E. Fourn., Mexic. Pl. 2: 147. 1886. T- 
Protol.; Mexico: Veracruz, Gouin 73 [HT:P]. T- 
Specim.: 1 of 1. Mexico: Veracruz, 1867,Gouin 73 [T: 
IT: US-91037 (fragm.)]. 

= Leptochloa nealleyi Vasey. 


136 


Catalogue of New World Grasses: II. 


Leptochloa subdigitata Trin. ex Steud., Nomencl. Bot. (ed. 2) 
2(2): 30. 1841. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Leptochloa digitata (R. Br.) Domin. 
= Leptochloa digitata (R. Br.) Domin. 


Leptochloa tracyi (Vasey) Beal, Grass. N. Amer. 2: 436. 
1896. Basionym: Diplachne tracyi Vasey. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Leptochloa uninervia (J. Presl) Hitche. & Chase, Contr. U.S. 
Natl. Herb. 18(7): 383. 1917. Basionym: Megastachya 
uninervia J. Presl. 

= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Leptochloa villosa Ekman, Ark. Bot. 10(17): 31, t. 3, f. 3, t. 
6, f. 16. 1911. T-Protol.: Brazil: Matto Grosso: 
Corumba, 11 Apr 1903, G.O [Malme 3075 [Exp. II 
Regn.] (HT: S]. T-Specim.: 1 of 1. Brazil: Mato 
Grosso: Corumba, 11 Apr 1903, G.O. Malme 3075 
[HT: S; IT: US-78805 (fragm.)], Explor. I regn. 

= Leptochloa virgata (L.) P. Beauv. 


Leptochloa virgata (L.) P. Beauv., Ess. Agrostogr. 71, 161, 
166, t. 15, f. 1. 1812. Basionym: Cynosurus virgatus L. 
DST.: Arge, Beli, Boli, Braz, Cari, Colo, CoRi, Ecua, ElSa, 
FrGu, Guat, Guya, Hond, Mexi, Nica, Pana, Para, 

Peru, Suri, USA, Urug, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Allem, A. C. 
& J. F. M. Valls (1987); Anonymous (1986): Balick, 
M. J., M. H. Nee & D. E. Atha (2000); Beetle, A. A. 
(1977); Berendsohn, W. G. & A. E. Araniva de 
Gonzalez (1991); Brako, L. & J. L. Zarucchi (1993); 
Breedlove, D. E. (1986); Burkart, A. (1969); Cabrera, 
A. L. (1970); Catasus Guerra, L. (1997); Correll, D. 
S. & M. C. Johnston (1970); Cowan, C. P. (1983); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Doll, J. C. (1878); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Foster, R. C. (1958); Gould, F. W (1979); Hitchcock, 
A. S. (1909); Hitchcock, A. S. (1927); Hitchcock, A. 
S. (1936); Hitchcock, A. S. (1951); Jorgensen, P. M. 
& S. Leén-Yanez (eds.) (1999); Judziewicz, E. J. 
(1990); Killeen, T. J. (1990); Lawesson, J. E., H. 
Adsersen & P. Bentley (1987); Lazarides, M. (1980); 
Lindeman, J.C. & A. L. Stoffers (1963); Liogier, H. 
& L. Martorell (1982); Long, R. W. & O. Lakela 
(1971); Macbride, J. F. (1936); McClure, F. A. 
(1955); McVaugh, R. (1983); Molina R., A. (1975); 
Pohl, R. W. & G. Davidse (1994); Pohl, R. W. 
(1980); Porter, D. M. (1983); Reeder, J. R. & C. G. 
Reeder (1971); Renner, S. S., H. Balslev & L. B. 
Holm-Nielsen (1990); Renvoize, S. A. (1984); 
Renvoize, S. A. (1988); Renvoize, S. A. (1998); 
Reyes-Garcia, A. & M. Sousa S. (1997); Small, J. K. 
(1933); Smith, L. B., D. C. Wasshausen & R. M. 
Klein (1981); Sousa S.. M. & E. F. Cabrera C. 
(1983); Tovar, O. (1993); Wiggins, I. L. & D. M. 
Porter (1971); Wunderlin, R. P. (1998); Zuloaga, F. 
O., E. G. Nicora, Z. E. Rugolo de Agrasar, O. 
Morrone, J. Pensiero & A. M. Cialdella (1994). 

Chloris digitaria Kunth, Chloris gracilis Kunth; 
Chloris poiformis Kunth; Cynodon domingense 
(Jacq.) Raspail; Cynodon virgatus (L.) Raspail; 
Cynosurus domingensis Jacq.; Cynosurus virgatus L.; 
Diplachne domingensis (Jacq.) Chapm.; Eleusine 


SYN.: 


digitata Willd. ex Steud.; Eleusine domingensis 
(Jacq.) Pers.; Eleusine gracilis (Kunth) Spreng.; 
Eleusine unioloides Willd. ex Steud.; Eleusine 
virgata (L.) Pers.; Festuca domingensis (Jacq.) Lam.,; 
Festuca virgata (L.) Lam.; Leptochloa barbata Desv. 
ex Killeen; Leptochloa digitaria (Kunth) Nees; 
Leptochloa domingensis (Jacq.) Trin.; Leptochloa 
gracilis Nees; Leptochloa mutica Steud.; Leptochloa 
perennis Hack.; Leptochloa villosa Ekman; 
Leptochloa virgata’ var. aristata E.  Fourn.; 
Leptochloa virgata var. domingensis (Jacq.) Link ex 
Griseb.; Leptochloa virgata var. gracilis (Nees) Link; 


Leptochloa virgata var. intermedia §E. Fourn.; 
Leptochloa virgata var. mutica (Steud.) Déll; 
Leptostachys domingensis  (Jacq.) G.  Mey.; 


Leptostachys virgata (L.) G. Mey.; Oxydenia virgata 
Nutt. ex B.D. Jacks.; Rabdochloa domingensis 
(Jacq.) P. Beauv. 


Leptochloa virgata var. aristata E. Fourn., Mexic. Pl. 2: 146. 
1886. T-Specim.: | of 7. [Mexico]: Tampico [ST: ST: 
herb. Uzac]. 2 of 7. [Mexico]: Colipa, Karw. 996 d 
part. [ST]. 3 of 7. [Mexico]: Colipa ad Jicaltepec, 
Karw. 996 b [ST]. 4 of 7. [Mexico]: Jicaltepec, Apr, 
Liebm. 252 [ST]. 5 of 7. [Mexico]: Huitamalco, Jun, 
Liebm. 245 [ST]. 6 of 7. [Mexico]: San Luis de Potosi, 
Virl. 1450 [ST]. 7 of 7. Jamaica, Galapagos, Peru, 
Dombey s.n. {ST}. 

= Leptochloa virgata (L.) P. Beauv. 


Leptochloa virgata var. domingensis (Jacq.) Link ex Griseb., 
Fl. Brit. W. [. 538. 1864. Basionym: Cynosurus 
domingensis Jacq. 

= Leptochloa virgata (L.) P. Beauv. 


Leptochloa virgata var. gracilis (Nees) Link, Fl. Brit. W. I. 
538. 1864. Basionym: Leptochloa gracilis Nees. 
= Leptochloa virgata (L.) P. Beauv. 


Leptochloa virgata var. intermedia E. Fourn., Mexic. Pl. 2: 
146. 1886. T-Protol.: [Mexico]. T-Specim.: 1 of 3. 
Mexico, Jul 1841, F.AZ Liebmann s.n. [ST: US- 
3266092 (fragm.)]. 2 of 3. Mexico: Paso de Dona 
Juana, Liebmann 243 [ST]. 3 of 3. Mexico: Colipa, 
Liebmann 251 |ST]. 

NOTE: cites "Cynosurus domingensis Sw. Obs. 40". 

REPLACED NAME: Leptochloa virgata var. intermedia E. 
Fourn. 

= Leptochloa virgata (L.) P. Beauv. 


Leptochloa virgata var. mutica (Steud.) Doll, Fl. Bras. 2(3): 
91. 1878. Basionym: Leptochloa mutica Steud. 
= Leptochloa virgata (L.) P. Beauv. 


Leptochloa virgata fo. nana Lindm., Kongl. Svenska 
Vetenskapsakad. Handl. 14(6): 33. 1900. T-Protol.: 
Rues de l'Assomption, Balansa /2/. T-Specim.: | of 1. 
Paraguay: Rues de l'Assomption, Balansa /2/ [HT: 
°?(fragm., MVFA)}. 
= Leptochloa virgata var. virgata. 


Leptochloa virgata var. puberula Hack., Repert. Spec. Nov. 
Regni Veg. 6(21-26): 345. 1909. T-Protol.: Paraguay: 
Gran Chaco, in sylvis ad flumen Pilcomayo, flor mens 
Maj., 7. Rojas 94 [HT: G]. T-Specim.: 1 of 1. 
Paraguay: Gran Chaco, in sylvis ad flumen Pilcomayo, 
May, 7. Rojas 94 {HT: G; IT: BAA-1516, US-88708 
(fragm.)]. 


Catalogue of New World Grasses: II. 


137 


DST.: Arge, Boli, Para, Vene. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Burkart, A. (1969); Nicora, E. G. 
(1995); Zuloaga, F. O., E. G. Nicora, Z. E. Rigolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

SYN.: Leptochloa virgata var. puberula Hack. 


Leptochloa’ virgata var. puberula Hack., Trab. Mus. 
Farmacol. 21: 34. 1909, T-Protol.: Montes en las 
Orillas del rio, May, 7: Rojas 94. 

NOTE: nom. nud. 
= Leptochloa virgata var. puberula Hack. 


Leptochloa virgata (L.) P. Beauv. var. virgata. 
DST.: Arge, Beli, Boli, Braz, Cari, Colo, CoRi, Ecua, ElSa, 
FrGu, Guat, Guya, Hond, Mexi, Nica, Pana, Para, 
Peru, Suri, USA, Urug, Vene. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Nicora, E. G. (1995). 
SYN.: Leptochloa virgata fo. nana Lindm. 


Leptochloa virletii E. Fourn. ex Hemsl., Biol. Cent.-Amer., 
Bot. 3: 566. 1885. 
NOTE: nom. nud. 
CORRECT NAME: Leptochioa virletii E. Fourn. 
= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Leptochloa virletii E. Fourn., Mexic. Pl. 2: 147. 1886. T- 
Protol.: Mexico: San Luis de Potosi, Virlet 1404 
[HT:P]. T-Specim.: | of 1. Mexico: San Luis Potosi, 
1851, M.D'aoust Virlet 1404 (IT: US-91039 (fragm.)]. 

= Leptochloa fusea subsp. uninervia (J. Presl) N. Snow. 


Leptochloa viscida (Scribn.) Beal, Grass. N. Amer. 2: 434. 
1896. Basionym: Diplachne viscida Scribn. 

DST.: Mexi, USA. 

LVL. ACCPTC.:: |. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. & R. Moran 
(1981); Hickman, J. C. (1993); Hitchcock, A. S. 
(1951); Munz, P. A. (1968); Shreve, F. & I. L. 
Wiggins (1964). 

SYN.: Diplachne viscida Scribn. 


Leptochloa subg. Diplachne (P. Beauv.) A. Gray, Manual 
588. 1848. Basionym: Diplachne P. Beauv. 


Leptochloa P. Beauv. subg. Leptochloa, NONE LISTED. 


Leptochloopsis H.O. Yates, SouthW. Naturalist 11(3): 382. 
1966. TYPE: Leptochloopsis virgata (Poir.) H.O. 
Yates. 

= Uniola L. 


Leptochloopsis condensata (Hitche.) H.O. Yates, SouthW. 
Naturalist 11(3); 389. 1966. Basionym: Uniola 
condensata Hitche. 

= Uniola condensata Hitchc. 


Leptochloopsis virgata (Poir.) H.O. Yates, SouthW. 
Naturalist 11(3); 384. 1966. Basionym: Poa virgata 
Poir. 


= Uniola virgata (Poir.) Griseb. 


Leptochloris Kuntze, Revis. Gen, Pl. 2: 771. 1891. 
NOTE: insyn. sub Trichloris crinita. 
= Trichloris E. Fourn. ex Benth. 


Leptochloris crinita (Lag.) Munro ex Kuntze, Revis. Gen. PI. 
2: 771. 1891. Basionym: Chloris crinita Lag. 
= Trichloris crinita (Lag.) Parodi. 


Leptochloris greggii Munro ex Merr., U.S.D.A. Div. Bot. 
Circ. 32: 7. 1901. 
NOTE: nom. inval., as syn. of Chloropsis mendocina (R.A. 
Phil.) Kuntze. 
= Trichloris crinita (Lag.) Parodi. 


Leptostachys G. Mey., Prim. Fl. Esseq. 73. 1818. TYPE: 
Leptostachys virgata (L.) G. Mey. 
NOTE: nom. illeg. superfl. 
= Leptochloa P. Beauv. 


Leptostachys domingensis (Jacq.) G. Mey., Prim. Fl. Esseq. 
74. 1818. Basionym: Cynosurus domingensis Jacq. 
= Leptochloa virgata (L.) P. Beauv. 


Leptostachys dubia (Kunth) G. Mey., Prim. FI. Esseq. 74. 
1818. Basionym: Chloris dubia Kunth. 
= Leptochloa dubia (Kunth) Nees. 


Leptostachys filiformis (Lam.) G. Mey., Prim. Fl. Esseq. 74. 
1818. Basionym: Festuca filiformis Lam. 
= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Leptostachys virgata (L.) G. Mey., Prim. Fl. Esseq. 74. 1818. 
Basionym: Cynosurus virgatus L. 
NOTE: basionym erroneously attributed to Willd. 
= Leptochloa virgata (L.) P. Beauv. 


Leptothrium Kunth, Revis. Gramin. 1: 156. 1829. 
TYPE: Leptothrium rigidum Kunth. 
TRTD. BY: Peterson, P. M. 
NHT: Zoysiinae. NO. ACCPTD. SPP.: 1. 
SYN.: Latipes Kunth. 


Leptothrium rigidum Kunth, Revis. Gramin. 156, 1829. T- 
Protol.: America calidor. T-Specim.: 1 of 1. South 
America: habitat Orinoco [IT: US (fragm. ex B- 
WILLD.); US (fragm. ex P-HBK)]. 

NOTE: based on unpublished Zoysia rigida Willd. 

DST.: Cari, Colo, Guya, Peru, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Hitchcock, A. 
S. (1936); Raimondo, F. M. (2001); Tovar, O. 
(1993). 


Lepturidium Hitchc. & Ekman, Man. Grasses W. Ind. 
111. 1936. TYPE: Lepturidium insulare Hitche. & 
Ekman. 

TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 1. 


Lepturidium insulare Hitchc. & Ekman, Man. Grasses W. 
Ind. 111, f. 71. 1936. T-Protol.: Cuba: Isla de Pinos: 
Santa Barbara, collected in white sand, saline flats, 3 
Nov. 1920, E.L. Ekman 12060 (HT: US-1296125]. T- 
Specim.: | of 2. Cuba: Isla de Pinos, 11 Mar 1920, 
E.L. Ekman 12060 [T: US-1161850]. 2 of 2. Cuba: La 
Habana: Isla de Pinos, 3 Nov 1920, E.L. Ekman 12060 
[HT: US-1296125]. 

DST.: Cari. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Catasus 
Hitchcock, A. S. (1936). 


Guerra, L. (1997); 


138 


Catalogue of New World Grasses: II. 


Lepturinae subtrib. Benth., J. Linn. Soc., Bot. 19: 31. 1881. 
NOTE: as "Leptureae". 


Lepturus paniculatus Nutt., Gen. N. Amer. Pl. 1: 81. 1818. T- 
Protol.: [USA]: North Dakota: on dry saline plains, 
near Fort Mandan, on the Missouri. 

= Schedonnardus paniculatus (Nutt.) Trel. 


Lepyroxis E. Fourn., Mexic. Pl. 2: 92. 1886. 
NOTE: nom. inval., pro syn. sub Polypogon canadensis. 
= Muhlenbergia Schreb. 


Lepyroxis canadensis P. Beauy. ex B.D. Jacks., Index Kew. 
1: 244. 1906, 
NOTE: nom. inval., as syn. of Agrostis mexicana L. 
= Muhlenbergia mexicana (L.) Trin. 


Lesourdia E. Fourn., Bull, Soc. Bot. France 27: 102. 1880. 
= Scleropogon Phil. 


Lesourdia karwinskyana E. Fourn., Bull. Soc. Bot. France 27: 
102, t. 4, f. 12. 1880. T-Specim.: | of 2.Karwinski 992 
[LT], LT designated by ?. 2 of 2. : Parry et Palmer 
925 [ST]. 
= Scleropogon brevifolius Phil. 


Lesourdia multiflora E. Fourn., Bull. Soc. Bot. France 27: 
102, t. 3-4. 1880. T-Protol.: Mexico: Tampico[HT: 
herb. Uzac]. 

= Scleropogon brevifolius Phil. 


Limnetis Rich. in Persoon, Syn. Pl. 1: 72. 1805. 
NOTE: nom. superfl. pro Spartina. 
= Spartina Schreb. 


Limnetis cynosuroides (L.) Rich., Syn. Pl. 1: 72. 1805. 
Basionym: Dactylis cynosuroides L. 
= Spartina cynosuroides (L.) Roth. 


Limnetis glabra Nutt., Gen. N. Amer. Pl. 1: 38. 1818. 
NOTE: nom. nud. 
= Spartina alterniflora Loisel. 


Limnetis glabra (Muhl. ex Elliott) Eaton & J. Wright, Man. 
Bot. (ed. 8) 301. 1840. Basionym: Spartina glabra 
Muhl. ex Elliott. 

= Spartina alterniflora Loisel. 


Limnetis polystachia (Michx.) Rich., Syn. Pl. 1: 72. 1805. 
Basionym: Trachynotia polystachya Michx. 
= Spartina cynosuroides (L.) Roth. 


Lycurinae subtrib. Pilg., Nat. Pflanzenfam. (ed. 2) 14d: 167, 
50. 1956. 


Lycurus Kunth, Nov. Gen. Sp. 1: 141. 1815 [1816]. 
TYPE: Lycurus phleoides Kunth, LT designated by 
Hitchcock, U.S.D.A. Bull. 772: 139 (1920). 

TRTD. BY: Peterson, P. M. 

NHT: Muhlenbergiinae. NO. ACCPTD. SPP.: 3. 

CNSLTD. LIT.: Reeder, C. G., (1985); Peterson, P. M. & 
O. Morrone, (1997); Sanchez, E. & Z. E. Rigolo de 
Agrasar, (1986). 

SYN.: Pleopogon Nutt. 


Lycurus alopecurus Griseb., Abh. Konigl. Ges. Wiss. 
Gottingen 19: 207. 1874. T-Protol.: Argentina: 
Catamarca: in alpinis convallis Granadillas pr. Belen, 
Lorentz s.n. T-Specim.: | of 1. Argentina: Prov. 
Catamarca: cerca de Belén, en el altivalle de las 


Granadillas, Feb 1872, P. G. Lorentz s.n., [Hb. Kurtz 
548) [HT: ?; IT: BA, BAA, SI, US-996080 (fragm. ex 
herb. Griseb)]. 

= Lycurus setosus (Nutt.) C. Reeder. 


Lycurus phalaroides Kunth, Nov. Gen. Sp. 1: 142. 1815 
[1816]. T-Protol.: Crescit in = montanis —regni 
Mechoacanensis juxta Valladolid, Alberca de Palangeo 
et Patzcuaro, alt. 950-1060 hexap., floret Septembri, 
Humboldt & Bonpland s.n. T-Specim.: | of 1. Mexico: 
Michoacan: near Valladolid, Alberca de Palangeo and 
Patzcuaro, Sept, Humboldt & Bonpland s.n. [HT: P: 
IT: B-W-1630, BM, BAA-1530, US-91988 (fragm. ex 
P-Bonpl.), US-610837 (fragm. ex LE-TRIN)]. 

DST.: Arge, Boli, Colo, Ecua, Guat, Mexi, Peru, USA, 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Brako, L. & J. L. Zarucchi (1993); 
Breedlove, D. E. (1986); Davidse, G. (1994): 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A.. A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Foster, R. C. 
(1958); Jorgensen, P. M. & S. Leén-Yanez (eds.) 
(1999); McVaugh, R. (1983); Peterson, P. M. (2000); 
Reeder, C. G. (1985); Renvoize, S. A. (1998); 
Sanchez, E. & Z. E. Rugolo de Agrasar (1986); 
Tovar, O. (1993); Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 

Lycurus phleoides var. brevifolius Scribn. ex Beal; 
Muhlenbergia lycuroides Vasey ex Beal. 


SYN.: 


Lycurus phleoides Kunth, Nov. Gen. Sp. |: 142, t. 45. 1815 
[1816]. T-Protol.: Crescit in temperatis Mexici, inter 
Guanaxuato et Temascatio et in radicibus aridissimi 
montis La Buffa, alt. 1030 hexap., floret Septembri, 
Humboldt & Bonpland s.n. T-Specim.: | of 1. Mexico: 
inter Guanaxuato et Temascatio et in  radicibus 
aridissimi montis La Buffa, alt. 1030 hexap., Sept, 
Humboldt & Bonpland s.n. (HT: P; IT: BAA (fragm.), 
US-610840 (fragm. ex P-Bonpl.), US-610841 
(fragm.)]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Cronquist, A., A. H. 
Holmgren, N. H. Holmgren & J. L. Reveal (1977); 
Espejo Serna, A., A. R. L6pez-Ferrari, and J. Valdes- 
Reyna (2000); Foster, R. C. (1958); Gould, F. W. & 
R. Moran (1981); Great Plains Flora Association 
(1986); Hitchcock, A. S. (1927); Hitchcock, A. S. 
(1951); Macbride, J. F. (1936); McVaugh, R. (1983); 
Peterson, P. M. (2000); Reeder, C. G. (1985); 
Renvoize, S. A. (1998); Sanchez, E. & Z. E. Ragolo 
de Agrasar (1986). 

SYN.: Lycurus phleoides var. phleoides. 


Lycurus phleoides var. brevifolius Scribn. ex Beal, Grass. N. 
Amer. 2: 271. 1896. T-Specim.: 1 of 3. Mexico: 
Jalisco: plains of Guadalajara, 23 Oct 1889, C.G. 
Pringle 2470 [LT: MSC; ILT: BAA, GH, MEXU, 
MO-2972926, NY, P, US-996049], LT designated (as 
holotype) by C. Reeder, Phytologia 57(4): 288 (1985). 
2 of 3. Mexico, Botteri 680 [ST: US]. 3 of 3. Mexico, 
Palmer 489 [ST: US]. 

= Lycurus phalaroides Kunth. 


Catalogue of New World Grasses: II. 


139 


Lycurus phleoides var. glaucifolius Beal, Grass. N. Amer. 2: 
271. 1896. T-Specim.: | of 3. Mexico: Chihuahua: 
rocky hills near Chihuahua, 28 May 1885, C.G. 
Pringle 426 |LT: MSC; ILT: MO-3701765, NY, P, SI, 
US-996079], LT designated (as type) by Hitchcock, 
Contr. U.S. Natl. Herb. 17(3): 305 (1913), without 
explicitly citing a specific sheet in a_ specific 
herbarium; fully lectotypified by Reeder, Phytologia 
57(4): 287 (1985), who cited the MSC specimen as 
"holotype". 2 of 3. USA: Texas, Havard s.n. [ST]. 3 of 
3. USA: Texas, Nealley s.n. [ST]. 

= Lycurus setosus (Nutt.) C. Reeder. 


Lycurus phleoides Kunth var. phleoides. 
= Lycurus phleoides Kunth. 


Lycurus schaffneri (E. Fourn.) Mez, Repert. Spec. Nov. 
Regni Veg. 17: 212. 1921. Basionym: Muhlenbergia 
schaffneri E. Fourn. 

= Muhlenbergia depauperata Scribn. 


Lycurus setosus (Nutt.) C. Reeder, Phytologia 57: 287. 
1985. Basionym: Pleopogon setosum Nutt. 

DST.: Arge, Boli, Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Peterson, P. M. 
(2000); Reeder, C. G. (1985); Sanchez, E. & Z. E. 
Rugolo de Agrasar (1986); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

Lycurus alopecurus Griseb.; Lycurus phleoides var. 
glaucifolius Beal; Pleopogon setosum Nutt. 


SYN.: 


Macroblepharus Phil., Linnaea 29(1): 100. 1858. TYPE: 
Macroblepharus contractus Phil. 
= Eragrostis Wolf. 


Macrostachya A. Rich., Tent. Fl. Abyss. 2: 408. 1851. 
NOTE: nom. nud. 
= Enteropogon Nees. 


Matrella Pers., Syn. Pl. 1: 73. 1805. TYPE: Matrella juncea 
Pers. 
= Zoysia Willd. 


Matrella juncea Pers., Syn. Pl. 1: 73. 1805. 
REPLACED NAME: Agrostis matrella L. 
= Zoysia matrella (L.) Merr. 


Megastachya acutiflora (Kunth) Roem, & Schult., Syst. Veg. 
2: 588. 1817. Basionym: Poa acutiflora Kunth. 
= Eragrostis acutiflora (Kunth) Nees. 


Megastachya amabilis (L.) P. Beauy., Ess. Agrostogr. 74, 
167, 173. 1812. Basionym: Poa amabilis L. 
= Eragrostis amabilis (L.) Wight & Arn. ex Nees. 


Megastachya amoena E. Fourn. ex Hemsl., Biol. Cent.- 
Amer., Bot. 3: 572. 1885. 
NOTE: nom. inval., as syn. of Eragrostis amoena J. Presl. 
= Eragrostis maypurensis (Kunth) Steud. 


Megastachya aturensis (Kunth) Roem. & Schult., Syst. Veg. 
2: 587. 1817. Basionym: Poa aturensis Kunth, 
= Eragrostis japonica (Thunb.) Trin. 


Megastachya berteroniana Schult., Mant. 2: 330. 1824. T- 
Protol.: W.I.: Santo Domingo: In Domingo, D. Bertero 
sn. T-Specim.: | of 1. W.1.: Dominican Republic: 


Santo Domingo, D. Bertero [IT: IT: US-79730 
(fragm.)]. 


= Eragrostis berteroniana (Schult.) Steud. 


Megastachya brasiliensis Schult. ex B.D. Jacks., Index Kew.. 
NOTE: error in IK?, said to be published by Schult., Mant. 
2: 620 (1824), but no such name in the Mantissa. 
= Eragrostis japonica (Thunb.) Trin. 


Megastachya breviflora E. Fourn., Mexic. Pl. 2: 119. 1886. 
T-Protol.: Mexico: Veracruz, Gouin 84 (pro parte) 
{HT: P]. T-Specim.: 1 of 1. Mexico: Veracruz, Gouin 
84 (pro parte) [HT: P; IT: US-2821450 (fragm.)]. 
= Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb. 


Megastachya condensata J. Pres|, Relig. Haenk. 1(4-5): 284. 
1830. T-Protol.: Hab. in Mexico [error for Ecuador], 
Haenke s.n. T-Specim.: | of 1. Ecuador, Haenke s.n. 
[HT: PR; IT: US-78689, W]. 

= Eragrostis condensata (J. Presl) Steud. 


Megastachya corymbifera E. Fourn., Mexic. Pl. 2: 119. 1886. 
T-Protol.: Mexico: Laguna de S. Nicolas, Berlandier 
1949. T-Specim.: | of 1. Mexico: Laguna de S. 
Nicolas, JL. Berlandier 1949 [IT: IT: US-2821452 
(fragm.)]. 

= Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb. 


Megastachya cuspidata Schult., Mant. 2: 329. 1824. 
REPLACED NAME: Poa cuspidata Roth. 
BLOCKING NAME: Poa cuspidata Nutt. 
= Eragrostis rufescens var. rufescens. 


Megastachya eragrostis (L.) P. Beauv. ex Roem. & Schult., 
Syst. Veg. 2: 575, in obs.; 584. 1817. Basionym:Briza 
eragrostis L. 

= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Megastachya fasciculata E. Fourn., Mexic. Pl]. 2: 120. 1886. 
T-Protol.: Mexico: Matamoros, Berl. 2325 [T:US]. T- 
Specim.: | of 1. Mexico: Matamoros, /.L. Berlandier 
2325 [T: IT: US-2821451 (fragm.)]. 

= Eragrostis reptans (Michx.) Nees. 


Megastachya glomerata (Walter) Schult., Mant. 2: 327. 1824. 
Basionym: Poa glomerata Walter. 
= Eragrostis japonica (Thunb.) Trin. 


Megastachya gouinii E. Fourn. ex Hemsl., Biol. Cent.-Amer., 
Bot. 3: 576. 1885. 
NOTE: nom. nud. 
= Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb. 


Megastachya gouinii E. Fourn., Mexic. Pl. 2: 119. 1886. T- 
Protol.: Mexico: circa Veracruz, Gouin 85 [T:US]. T- 
Specim.: | of 1. Mexico: circa Veracruz, 1865, Gouin 
85 [IT: IT: US-2850741 (fragm.)]. 

= Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb. 


Megastachya hypnoides (Lam.) P. Beauv., Ess. Agrostogr. 
74, 167, 175. 1812. Basionym: Poa hypnoides Lam. 
= Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb. 


Megastachya leersioides C. Presl, Cyper. Gramin. Sicul. 44. 
1820. T-Protol.: Sicily: Hab. in arenosis maritimis 
Panormi. 

= Eragrostis eragrostis subvar. leersioides (C. Pres!) Farw. 


Megastachya maypurensis (Kunth) Roem. & Schult., Syst. 
Veg. 2: 588. 1817. Basionym: Poa maypurensis 
Kunth. 


140 


Catalogue of New World Grasses: II. 


= Eragrostis maypurensis (Kunth) Steud. 


Megastachya montufarii (Kunth) Roem. & Schult., Syst. 
Veg. 2: 586. 1817. Basionym: Poa montufari Kunth. 
= Eragrostis pastoensis (Kunth) Trin. 


Megastachya nigricans (Kunth) Roem. & Schult., Syst. Veg. 
2: 586. 1817. Basionym: Poa nigricans Kunth. 
= Eragrostis nigricans (Kunth) Steud. 


Megastachya obtusa (Nutt.) Schult., Mant. 2: 326. 1824. 


Basionym: Poa obtusa Nutt. 
NOTE: nom. illeg. superfl.; should be based on Poa 
pennsylvanica Nutt. 
CORRECT NAME: Poa pennsylvanica Nutt. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Megastachya olmedoi (Kunth) Roem. & Schult., Syst. Veg. 
2: 586. 1817. Basionym: Poa olmedoi Kunth. 
= Eragrostis pastoensis (Kunth) Trin. 


Megastachya oxylepis E. Fourn. ex Hemsl., Biol. Cent.- 
Amer., Bot. 3: 574. 1885. 

NOTE: nom. inval.; as comb. in syn. based on Poa oxylepis 
Torrey; as syn. of Eragrostis oxylepis (Torrey) 
Torrey. 

= Eragrostis secundiflora subsp. oxylepis (Torr.) S.D. 
Koch. 


Megastachya oxylepis (Torr.) E. Fourn., Mexic. Pl. 2: 118. 
1886. Basionym: Poa oxylepis Torr. 
= Eragrostis secundiflora subsp. oxylepis (Torr.) S.D. 
Koch. 


Megastachya oxylepis var. capitata E. Fourn., Mexic. Pl. 2: 
118. 1886. T-Protol.: Mexico: Veracruz. T-Specim.: 1 
of 2. Mexico: Veracruz, 22 Jul 1865,Gouin 87 [ST: P; 
IST: US-2850754 (fragm.)]. 2 of 2. Mexico: Veracruz, 
Galeotti 5705 [ST]. 
= Eragrostis secundiflora J. Presl. 


Megastachya panamensis E. Fourn. ex Hemsl., Biol. Cent.- 
Amer., Bot. 3: 574. 1885. 
NOTE: nom. inval., as syn. of Eragrostis maypurensis 
Steud. 
= Eragrostis maypurensis (Kunth) Steud. 


Megastachya panamensis (J. Presl) E. Fourn., Mexic. Pl. 2: 
118. 1886. Basionym: Eragrostis panamensis J. Presl. 
= Eragrostis maypurensis (Kunth) Steud. 


Megastachya panicoides J. Presl, Relig. Haenk. 1(4-5): 283. 
1830. T-Protol.: Habitat in Mexico ad Acapulco, 
verosimiliter in lutosis, //aenke s.n. T-Specim.: | of 1. 
Mexico: Guerrero: Acapulco, 7) Haenke s.n. [HT: PR; 
IT; LE-TRIN-2121.02, MO-2109566, MO-2109567 
(line drawing), US-78688 (fragm.)]. 

= Leptochloa panicoides (J. Pres!) Hitche. 


Megastachya pastoensis (Kunth) Roem. & Schult., Syst. Veg. 
2: 587. 1817. Basionym: Poa pastoensis Kunth. 
= Eragrostis pastoensis (Kunth) Trin. 


Megastachya patula (Kunth) Roem. & Schult., Syst. Veg. 2: 
585. 1817. Basionym: Poa patula Kunth. 
= Eragrostis patula (Kunth) Steud. 


Megastachya prostrata (Kunth) Roem. & Schult., Syst. Veg. 
2: 590. 1817. Basionym: Poa prostrata Kunth. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Megastachya purpurascens (Muhl.) Schult., Mant. 2: 326. 
1824. Basionym: Briza purpurascens Muhl. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Megastachya reptans (Michx.) P. Beauy., Ess. Agrostogr. 74, 
167, 175. 1812. Basionym: Poa reptans Michx. 
= Eragrostis reptans (Michx.) Nees. 


Megastachya simpliciflora J. Presl, Relig. Haenk. 1(4-5): 
283. 1830. T-Protol.: Hab. in Mexico, Haenke s.n. T- 
Specim.: | of 1. Mexico, 7. Haenke s.n. [HT: P; IT: 
US-80149 (fragm.)]. 

= Eragrostis simpliciflora (J. Presl) Steud. 


Megastachya spectabilis (Pursh) Roem. & Schult., Syst. Veg. 
2: 589. 1817. Basionym: Poa spectabilis Pursh. 
= Eragrostis spectabilis (Pursh) Steud. 


Megastachya swainsonii Raddi, Agrostogr. Bras. 52. 1823. 
T-Protol.: Brasilia: Species rarissima ex itinere 
Pernambucano ad Urbem Rio Janeiro, D. Swainson 
sn. T-Specim.: | of 1. Brazil: Pernambucano: ad 
Urbem Rio Janeiro, D. Swainson [HT: Pl; IT: US- 
2850752 (fragm.)]. 

= Eragrostis maypurensis (Kunth) Steud. 


Megastachya tenax (Kunth) Roem. & Schult., Syst. Veg. 2: 
587. 1817. Basionym: Poa tenax Kunth. 
= Eragrostis condensata (J. Presl) Steud. 


Megastachya thalassica (Kunth) Roem. & Schult., Syst. Veg. 
2: 590. 1817. Basionym: Poa thalassica Kunth. 
= Distichlis spicata (L.) Greene. 


Megastachya uninervia J. Presl, Relig. Haenk. 1(4-5): 283. 
1830. T-Protol.: Hab. in Mexico, Haenke s.n. T- 
Specim.: | of 1. Mexico, Haenke s.n. [LT: PR; ILT: 
LE-TRIN-2424.01, US-78697 (fragm.), WJ], LT 
designated by Snow, Novon 8: 79 (1998). 

= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Melica anomala Scribn. ex Beal, Grass. N. Amer. 2: 311. 
1896. 
NOTE: nom. inval., as syn. of Epicampes anomala Scribner 
ex Beal. 
= Muhlenbergia longiligula Hitchc. 


Melica curtipendula Michx. ex Steud., Nomencl. Bot. (ed. 2) 
1: 91, 519. 1821. 
NOTE: nom. inval., as syn. of Atheropogon apludioides 
Muhl. 
= Bouteloua curtipendula var. curtipendula, 


Melica x curtisetosa (Scribn.) R.W. Pohl, Amer. Midl. 
Naturalist 82: 528. 1969. Basionym: Muhlenbergia 
schreberi subsp. curtisetosa Scribn. 

= Muhlenbergia x curtisetosa (Scribn.) Bush (pro sp.). 


Melica mexicana Link ex E. Fourn., Bull. Soc. Roy. Bot. 
Belgique 15: 471. 1876. 
NOTE: nom. inval., as syn. of Buchloe dactyloides (Nutt.) 
Engelm. 
= Bouteloua dactyloides (Nutt.) Columbus. 


Melica multinervosa Vasey, Bot. Gaz. 16(8): 235. 1891. T- 
Protol.: USA: Texas: collected at Brazos Santiago, 
1891, G.C. Nealley s.n. T-Specim.: | of 1. USA: 
Texas: Brazos Santiago, G.C. Nealley s.n. (HT: ?; IT: 
US-81963]. 

= Vaseyochloa multinervosa (Vasey) Hitche. 


Catalogue of New World Grasses: II. 


141 


Merisachne Steud., Syn. Pl. Glumac. I: 117. 1854 ("1855"). 
TYPE: Merisachne drummondii Steud. 
= Triplasis P. Beauv. 


Merisachne drummondii Steud., Syn. Pl. Glumac. 1: 117. 
1854. T-Protol.:; Texas, Drummond herb, 330. 
= Triplasis purpurea (Walter) Chapm. 


Micrairoideae subfam. Pilg., Nat. Pflanzenfam. (ed. 2) 14d: 
167, 1. 1956. TYPE: Micraira F. Muell. 


Microchloa Rk. Br.. Prodr. 208. 1810. TYPE: 


Microchloa setacea (Roxb.) R. Br.; nom. illeg. superfl. 
Microchloa indica (L. f.) P. Beauv. 
TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 2 SUBSP.: 2. 
CNSLTD. LIT.: Launert, E., (1966). 
SYN.: Micropogon Pfeiff.; Rendlia Chiov. 


Microchloa abyssinica Hochst., Flora 24(Intell.): 19. 1841. 
NOTE: nom. nud. 
= Microchloa kunthii Desv. 


Microchloa abyssinica Hochst. ex A. Rich., Tent. Fl. Abyss. 
2: 404. 1851. T-Specim.: 1 of 2. Ethiopia: in montibus 
et collibus ad rupes nudas prope Adoam, 28 Sep 1837, 
Schimper, pl. Schimp. Abyss., sect I, 321 (ST: ? BM, P. 
IST: K, US-878303]. 2 of 2. Abyssinia: In montosis 
provincie Chire, Quartin Dillon s.n. [ST]. 
= Microchloa kunthii Desv. 


Microchloa elongata R. Br., Numer. List 3807. 1831. 
NOTE: nom. nud. 
= Microchloa kunthii Desv. 


Microchloa indica (L. f.) P. Beauv., Ess. Agrostogr. 13, t. 
20, f. 8. 1812. Basionym: Nardus indica L. f. 

DST.: Arge, Boli, Braz, Chil, Para, Urug. 

LVL. ACCPTC:.: I. 

ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 
(1987); Bor, N. L. (1960); Brako, L. & J. L. Zarucchi 
(1993); Filgueiras, T. S. (1991); Filgueiras, T. S. 
(1999); Judziewicz, E. J. (1990); Killeen, T. J. 
(1990); Launert, E. (1966); Lazarides, M. (1972); 
Phillips, S. (1995); Renvoize, S. A. (1984); 
Renvoize, S. A. (1998); Rosengurtt, B. B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 
Smith, L. B., D. C. Wasshausen & R. M. Klein 
(1981); Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

Microchloa indica (L. f.) Hack.; Microchloa indica 
(L. f.) Kuntze; Microchloa indica var. gracilis 
Rendle; Microchloa setacea (Roxb.) R. Br.; Nardus 
indica L. f.; Rottboellia setacea Roxb. 


SYN.: 


Microchloa indica (L. f.) Kuntze, Revis. Gen. PI. 3(2): 356. 
1898. Basionym: Nardus indica L. f. 
NOTE: isonym. 
E. NAME: Microchloa indica (L. f.) P. Beauv. 
CORRECT NAME: Microchloa indica (L. f.) P. Beauv. 
= Microchloa indica (L. f.) P. Beauv. 


Microchloa indica (L. f.) Hack., Repert. Spec. Nov. Regni 
Veg. 7(152-156): 373. 1909. Basionym:Nardus indica 
Lf. 
NOTE: hom. illeg. 
CORRECT NAME: Microchloa indica (L. f.) P. Beauv. 


= Microchloa indica (L. f.) P. Beauv. 


Microchloa indica var. gracilis Rendle, Cat. Afr. Pl. 2(1): 
219. 1899. T-Specim.: 1 of 1. Angola: Pungo 
Andongo, Feb 1857, Welwitsch 2811 (HT: BM]. 

= Microchloa indica (L. f.) P. Beauv. 


Microchloa indica P. Beauv. var. indica, -. 
DST.: Boli, Braz, Chil, Para, Urug. 
LVL. ACCPTC.: 1. 


Microchloa indica var. kunthii (Desv.) B.S. Sun & Z.H. Hu, 
FI. Reipubl. Popularis Sin. 10(1): 88, t. 27: 6-7. 1990. 
Basionym: Microchloa kunthii Desv. 

= Microchloa kunthii Desv. 


Microchloa indica var. tenuis Hack. & Stuck., Anales Mus. 
Nac. Hist. Nat. Buenos Aires 21: 113. 1911. T-Protol.: 
Argentina: Prov. Tucuman: Cafiada Alegre: Dep. 
Burruyacu, Stuckert 21,598. 

DST.: Arge. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 


Microchloa kunthii Desv., Mem. Soc. Agric. Angers 1: 179. 
1831. T-Protol.: Habitat in America aequinoxialia. T- 
Specim.: 1 of 1. Mexico: Hidalgo: Omitlan et Llano de 
las Tinaxas, Humboldt & Bonpland s.n. [HT: P]. 

NOTE: also in Opuse. Sci. Phys. Nat. 75 [reprint] 1831- 
1833. 

DST.: Arge, Boli, Braz, Chil, Ecua, Guat, Hond, Mexi, 
Nica, Para, Peru, USA, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Bor, N. L. 
(1960); Breedlove, D. E. (1986); Davidse, G. (1994); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Filgueiras, T. S. 
(1991); Gibbs Russell, G. E, W. G. Welman, E. 
Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987); Gould, F. W. & R. Moran (1981); Hitchcock, 
A. S. (1951); Jorgensen, P. M. & S. Leén-Yanez 
(eds.) (1999); Launert, E. (1966); McClure, F. A. 
(1955); McVaugh, R. (1983); Molina R., A. (1975); 
Phillips, S$. (1995); Renvoize, S. A. (1988); 
Renvoize, S. A. (1998); Shreve, F. & I. L. Wiggins 
(1964); Tovar, O. (1993). 

SYN. Microchloa abyssinica Hochst.; Microchloa 
abyssinica Hochst. ex A. Rich.; Microchloa elongata 
R. Br.; Microchloa indica var. kunthii (Desv.) B.S. 
Sun & Z.H. Hu; Paspalum tenuissimum M.E. Jones. 


Microchloa setacea (Roxb.) R. Br., Prodr. 208. 1810. 
Basionym: Roftboellia setacea Roxb. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Microchloa indica (L. f.) P. Beauv. 
= Microchloa indica (L. f.) P. Beauv. 


Micropogon Pfeiff., Nomencl. Bot. 2: 310. 1847. 
NOTE: nom. superfl. pro Microchloa. 
= Microchloa R. Br. 


Milium dactylon (L.) Moench, Methodus 67. 1802. 
Basionym: Panicum dactylon L. 
= Cynodon dactylon (L.) Pers. 


142 


Catalogue of New World Grasses: II. 


Milium microspermum Lag., Gen. Sp. Pl. 2. 1816. T-Protol.: 
Habitat in Nova Hispania. Semina communicavit 
perill. D. Sesse. T-Specim.: | of 1. Mexico, D. Sessé 
[HT: MA; IT: US-91019 (fragm.)]. 

= Muhlenbergia minutissima (Steud.) Swallen. 


Molinia retusa Griseb. ex E. Fourn., Mexic. Pl. 2: 147. 1886. 
NOTE: nom. inval., as syn. of Leptochloa dubia (Kunth) 
Nees. 
= Leptochloa dubia (Kunth) Nees. 


Monanthochloe Engelm. Trans. Acad. Sci. St. 


Louis 1: 436. 1859. TYPE: Monanthochloe littoralis 
Engelm. 
TRTD. BY: Peterson, P. M. 
NHT: Monanthochloinae. NO. ACCPTD. SPP.: 2. 
CNSLTD. LIT.: Villamil, C. B., (1969). 
SYN.: Halochloa Griseb.; Solenophyllum Baill. 


Monanthochloe acerosa (Griseb.) Speg., Anales Mus. Nac. 
Buenos Aires 7: 194. 1879. Basionym: Halochloa 
acerosa Griseb. 

DST.: Arge. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Villamil, C. B. (1969); Zuloaga, F. 
O., E. G. Nicora, Z. E. Rugolo de Agrasar, O. 
Morrone, J. Pensiero & A. M. Cialdella (1994). 

SYN.: Halochloa acerosa Griseb. 


Monanthochloe australis Speg., Anales Mus. Nac. Hist. Nat. 
Buenos Aires 7: 196. 1902. T-Protol.: [Argentina]: 
Hab. in aridis subsalsis inter San Julia et Rio Deseado, 
1899, C. Ameghino s.n. T-Specim.: | of 1. Argentina: 
Santa Cruz: entre San Julian y Rio Deseado, 1899, 
Ameghino s.n. (HT: CORD]. 
= Distichlis australis (Speg.) Villamil. 


Monanthochloe littoralis Engelm., Trans. Acad. Sci. St. 
Louis 1: 437, pl. 13-14. 1859. T-Specim.: | of 6. USA: 
Texas (Matamoras Region), no date, JL. Berlandier 
3227 |ST: MO-1837526, MO-1837527; IST: US- 
899643]. 2 of 6. USA: Texas: Galveston Island, 
covering large tracts of moist, sandy, saline soil, May 
1843, Lindheimer 63 [LT: MO-1837530; ILT: MO- 
1837531, MO-1837528], LT designated by Davidse & 
Pohl, Fl. Mesoamer. 6: 258 (1994). 3 of 6. USA: 
Texas: Galveston Island, 1842, Lindheimer s.n. [ST: 
MO-1837529]. 4 of 6. USA: Texas: Galveston Island, 
Sep 1841, Lindheimer s.n. [ST: MO-1837532]. 5 of 6. 
USA: Texas, Drummond s.n. [ST: BM]. 6 of 6. USA: 
Florida: Key West, Blodgett s.n. [ST]. 

DST.: Cari, Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Abrams, L. (1923); Beetle, A. A. 
(1977); Breedlove, D. E. (1986); Catasus Guerra, L. 
(1997); Correll, D. S. & M. C. Johnston (1970); 
Davidse, G. & R. W. Pohl (1994); Davidse, G., M. 
Sousa S. & A. O. Chater (editores generales) (1994); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Godfrey, R. K. & J. W. Wooten 
(1979); Gould, F. W. & R. Moran (1981); Hickman, 
J. C. (1993); Hitchcock, A. S. (1936); Hitchcock, A. 
S. (1951); Long, R. W. & O. Lakela (1971); Munz, P. 
A. & D. D. Keck (1959); Munz, P. A. (1974); ref. 
missing (1994); Shreve, F. & I. L. Wiggins (1964): 
Small, J. K. (1933); Villamil, C. B. (1969); 
Wunderlin, R. P. (1998). 


Monanthochloinae subtrib. Pilg. ex Potztal, 
Willdenowia 5(3): 472. 1969. TYPE: Monanthochloe 
Engelm. 

TRTD. BY: Soreng, R. J., G. Davidse, T. S. Filgueiras, E. J. 
Judziewicz, P. M. Peterson, and F. O. Zuloaga. 
NHT: Cynodonteae. 


Monathera Raf., Amer. Monthly Mag. & Crit. Rev. 4: 190. 
1819. 
NOTE: nom. superfl. pro Monocera Elliott. 
= Ctenium Panz. 


Monerma gangitis (L.) Roem. & Schult., Syst. Veg. 2: 800. 
1817. Basionym: Nardus gangitis L. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Monermeae trib. C.E. Hubb., Brit. Flow. Pl. 332. 1948. 
NOTE: nom. inval., without Latin description. 


Monerminae subtrib. Janch., Phyton (Horn) 5: 60. 1953, 
NOTE: nom. nud. 


Monocera Elliott, Sketch Bot. S. Carolina 1: 176. 1816, 
TYPE: Monocera aromatica (Walter) Elliott. 
= Ctenium Panz. 


Monocera aromatica (Walter) Elliott, Sketch Bot. S. Carolina 
1: 176. 1816. Basionym: Aegilops aromatica Walter. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Monochaete Doll, Fl. Bras. 2(3): 78. 1875. TYPE: 
Monochaete fastigiata (Nees) D6ll. 


= Gymnopogon P. Beauv. 


Monochaete fastigiata (Nees) D6ll, Fl. Bras. 2(3): 79. 1878. 
Basionym: Gymnopogon fastigiatus Nees. 
= Gymnopogon fastigiatus Nees. 


Muehlenbergia annua (Vasey) Swallen, Contr. U.S. Natl. 
Herb. 29(4) 204. 1947. Basionym: Sporobolus annuus 
Vasey. 

NOTE: orth. var. 
CORRECT NAME: Muhlenbergia annua (Vasey) Swallen. 
= Muhlenbergia annua (Vasey) Swallen. 


Muehlenbergia lanata (Kunth) Hitche., N. Amer. Fl. 17; 459. 
1935. Basionym: Agrostis lanata Kunth. 
NOTE: orth. var. 
CORRECT NAME: Muhlenbergia lanata (Kunth) Hitche. 
= Muhlenbergia pubescens (Kunth) Hitchce. 


Muehlenbergia macroura (Kunth) Hitche., N. Amer, Fl. 17: 
468. 1935. Basionym: Crypsis macroura Kunth. 
NOTE: orth. var. 
CORRECT NAME: Muhlenbergia macroura (Kunth) 
Hitche. 
= Muhlenbergia macroura (Kunth) Hitche. 


Muehlenbergia pubescens (Kunth) Hitche., N. Amer. Fl. 17: 
460. 1935, Basionym: Agrostis pubescens Kunth. 
NOTE: orth. var. 
CORRECT NAME: 
Hitche. 
= Muhlenbergia pubescens (Kunth) Hitche. 


Muhlenbergia pubescens (Kunth) 


Muehlenbergia ramulosa (Kunth) Swallen, Contr. U.S, Natl. 
Herb. 29(4) 205. 1947. Basionym: Vilfa ramulosa 
Kunth. 

NOTE: orth. var. 


Catalogue of New World Grasses: II. 


143 


CORRECT NAME: 
Swallen. 
= Muhlenbergia ramulosa (Kunth) Swallen. 


Muhlenbergia ramulosa (Kunth) 


Muehlenbergia robusta (E. Fourn.) Hitche., N. Amer. Fl. 17: 
462. 1935. Basionym: Epicampes robusta E. Fourn. 
NOTE: orth. var. 
CORRECT NAME: Muhlenbergia robusta (E. Fourn.) 
Hitche. 
= Muhlenbergia robusta (E. Fourn.) Hitche. 


Muehlenbergia torreyana (Schult.) Hitche., Amer. J. Bot. 21: 
136. 1934. Basionym: Agrostis torreyana Schult. 
NOTE: orth. var. 
CORRECT NAME: Muhlenbergia torreyana_ (Schult.) 
Hitche. 
= Muhlenbergia torreyana (Schult.) Hitche. 


Muehlenbergia utilis (Yorr.) Hitche., J. Wash. Acad. Sci. 23: 
453. 1933. Basionym: Vilfa utilis Torr. 
NOTE: orth. var. 
CORRECT NAME: Muhlenbergia utilis (Torr.) Hitche. 
= Muhlenbergia utilis (Torr.) Hitchc. 


Muhlenbergia Schreb., Gen. Pl. 44. 1789. TYPE: 
Muhlenbergia schreberi J.F. Gmel.; LT designated by 
Nash in Britton & Brown, III. Fl. N. U.S. (ed. 2) 1: 184 
(1913). 
TRTD. BY: Peterson, P. M. 
NHT: Muhlenbergiinae. NO. ACCPTD SPP.: 147, SUBSP.: 
6. 

CNSLTD. LIT.: Hitchcock, A. S., (1935); Soderstrom, T. 
R., (1967); Peterson, P. M. & C. R. Annable, (1991); 
Herrera-Arrieta, Y., (1998); Peterson, P. M., (2000); 
Peterson, P. M. and Y. Herrera-Arrieta, (2001). 
Acroxis Steud., Anthipsimus Raf.; Calycodon Nutt.; 
Clomena P.  Beauv.; Crypsinna E.  Fourn.; 
Dactylogramma Link; Dilepyrum Michx.; Epicampes 
J. Presl; Lepyroxis E. Fourn.; Podosaemum Kunth; 
Podosemum Desv.;, Sericrostis Raf.; Tosagris P. 
Beauv.; Trichochloa DC.;, Trichochloa P. Beauv.; 
Vaseya Thurb. 


SYN.: 


Muhlenbergia abata 1.M. Johnst., J. Arnold Arbor. 24(4): 
387-388. 1943. T-Protol.: USA: New Mexico: valley 
of the Rio Grande 10-100mi above El Paso, Wright 
1/982 [HT: GH]. T-Specim.: 1 of I. USA: New 
Mexico: valley of the Rio Grande, 1851-1852, C. 
Wright 1983 [HT: GH; IT: US-556896]. 

= Muhlenbergia repens (J. Presl) Hitchc. 


Muhlenbergia acuminata Vasey, Bot. Gaz. 11(12): 337-338. 
1886. T-Protol.: USA: New Mexico, C. Wright 1993. 
T-Specim.: | of 1. USA: New Mexico, 1851-1852,C. 
Wright 1993 [IT: US-81605, US-995177]. 

= Muhlenbergia dubia E. Fourn. 


Muhlenbergia acutifolia E. Fourn. ex Hemsl., Biol. Cent.- 
Amer., Bot. 3: 538. 1885. 
NOTE: nom. nud. 
= Muhlenbergia spiciformis Trin. 


Muhlenbergia acutifolia E. Fourn., Mexic. Pl. 2: 86. 1886. T- 
Protol.: Mexico: Orizaba, Nov, Bourg. 3327. T- 
Specim.: 1 of 1. Mexico: Veracruz: Orizaba, 8 Nov 
1866, M. Bourgeau 3327 (HT: P; IT: MO-2974301, 
US-87235 (fragm.), US-2561240]. 

= Muhlenbergia spiciformis Trin. 


Muhlenbergia adspersa Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(3-4): 291. 1841. T-Protol.: Cult. "Cynosurus 
tenellus ht. Madrit. (in herb. Mertensii). Lima?". T- 
Specim.: | of 1. Lima? [HT: LE; IT: LE-TRIN- 
1486.01 (fragm. ex LE hb. Mertensii), US-87236 
(fragm. ex LE hb. Mertens)], "Cynosurus tenellus ex 
Hh. Madr. in herb. Mertensii". 

NOTE: preprint p. 45 (1841, later). 
= Muhlenbergia ciliata (Kunth) Trin. 


Muhlenbergia affinis Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(3-4): 301. 1841. T-Protol.: Mexico: Toluca, 
Berlandier 1083. T-Specim.: | of 1. Mexico: México: 
Toluca, J.L. Berlandier 1083 (IT: LE-TRIN-1485.01 
(fragm.), US-87237 (fragm.)]. 

= Muhlenbergia rigida (Kunth) Kunth. 


Muhlenbergia aguascalientensis Y. Herrera & De la Cerda- 
Lemus, Novon 5(3): 278, f. 1. 1995. T-Protol.: Mxico: 
Aguascalientes: Mun. de San José de Gracia: NW of 
Aguascalientes, 12 km NW from La Congoja, Playa 
Mariquiotas, Monte Grande de la Sierra Fria, 2850 m, 
22.25.7°N 102.37.25°W, 8 Nov 1986,G. Garcia 2801 
{[HT:HUAA; IT:CHDIR, ENCB, IEB]. T-Specim.: | of 
1. Mexico: Aguascalientes: San Jose de Gracia Mun.: 
Playa Mariquitas, Monte Grande de Sierra Fria: 2850 
mt, 8 Nov 1986, G.Garcia 280] [HT: HUAA; IT: 
CIIDIR, ENCB, IEB, US-3335737]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Herrera-Arrieta, 
Y. & M. de la Cerda-Lemus (1995); Herrera-Arrieta, 
Y. (1998); Peterson, P. M. (2000). 


Muhlenbergia alamosae Vasey, Bot. Gaz. 16(5): 146. 1891. 
T-Protol.: Mexico: collected at Alamosa in Sonora in 
1890, Palmer 407 [HT: US-81604]. T-Specim.: | of 1. 
Mexico: Sonora: Sierra de Almos, 25 Mar-8 Apr 1890, 
E. Palmer 407 [HT: US-81604; IT: US-822802]. 

DST.: Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000): 
Gould, F. W. & R. Moran (1981); McVaugh, R. 
(1983): Peterson, P. M. (2000). 


Muhlenbergia alta Hitchc., N. Amer. FI. 17(6): 461. 1935. T- 
Protol.: Mexico: hills east of Zaptolan, Jalisco, alt., 
1500 m, 25 Sep 1910, 4.8. Hitchcock 7180 [HT: US- 
998980]. T-Specim.: | of 1. Mexico: Jalisco: hills east 
of Zapotlan, 25 Sep 1910, A.S. Hitchcock 7180 (HT: 
US-998980]. 
= Muhlenbergia gigantea (E. Fourn.) Hitchc. 


Muhlenbergia ambigua Torr., Rep. Miss. Rv. 164. 1843. T- 
Protol.; USA: Stony banks of Okaman lake, Sioux 
country, 20 Sep, C. Geyer s.n. T-Specim.: 1 of 1. 
USA: Okaman Lake, Sioux Country, 20 Sep 1838, 
C.A, Geyer s.n. (IT: US-952523]. 

= Muhlenbergia mexicana var. filiformis (Torr.) Scribn. 


Muhlenbergia ambigua Torr. var. ambigua. 
= Muhlenbergia mexicana var. filiformis (Torr.) Scribn. 


144 


Catalogue of New World Grasses: II. 


Muhlenbergia ambigua var. filiformis (Torr.) Farw., Rep. 
Michigan Acad. Sci. 20: 168. 1919. Basionym: 
Agrostis lateriflora var. filiformis Torr. 

NOTE: basionym erroneously attributed to Muhl. 
= Muhlenbergia mexicana var. filiformis (Torr.) Scribn. 


Muhlenbergia andina (Nutt.) Hitche., U.S.D.A. Bull. (1915- 
23) 772: 145. 1920. Basionym: Calamagrostis andina 
Nutt. 

DST.: USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Abrams, L. (1923); Correll, D. S. & 
M. C. Johnston (1970); Cronquist, A., A. H. 
Holmgren, N. H. Holmgren & J. L. Reveal (1977); 
Hickman, J. C. (1993); Hitchcock, A. S. (1951); 
Hitchcock, C. L., A. Cronquist, M. Ownbey & J. W. 
Thompson (1969); Munz, P. A. & D. D. Keck 
(1959); Munz, P. A. (1974); Peterson, P. M. (2000). 

SYN.: Calamagrostis andina Nutt.; Muhlenbergia comata 
(Thurb.) Thurb. ex Benth.; Vaseya comata Thurb. 


Muhlenbergia angustata (J. Pres!) Kunth, Enum. Pl. 1: 202. 
1833. Basionym: Podosemum angustatum J. Presl. 
NOTE: also Revis. Gram. 3: Suppl. 1834 p. (as in 1835 ed.) 
XVI. 

DST.: Arge, Boli, Colo, Ecua, Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Brako, L. & J. L. Zarucchi (1993); 
Foster, R. C. (1958); Hitchcock, A. S. (1927); 
Jorgensen, P. M. & C. Ulloa U. (1994); Jorgensen, P. 
M. & S. Leén-Yanez (eds.) (1999); Macbride, J. F. 
(1936); Peterson, P. M. (2000); Renvoize, S. A. 
(1998); Tovar, O. (1993); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

Cinna phleoides (Kunth) Kunth: Cina’ stricta 
(Kunth) Kunth; Crypsinna stricta (Kunth) E. Fourn.; 
Crypsis phleoides Kunth; Crypsis stricta Kunth; 
Epicampes coerulea Griseb.; Epicampes coerulea 
var. submutica Hack. ex  Stuck.; Epicampes 
kunthiana Griseb.; Epicampes phleoides (Kunth) 
Griseb.; Muhlenbergia coerulea (Griseb.) Mez; 
Podosemum angustatum J. Presl. 


SYN.: 


Muhlenbergia angustifolia Swallen, N. Amer. Fl. 17(6): 457. 
1935. T-Protol.: Mexico: collected on rocky hills near 
Guadalajara, 11 Nov 1889, Pringle 2346 [HT: US- 
822822]. T-Specim.: | of 1. Mexico: Jalisco: rocky 
hills, near Guadalajara, 11 Nov 1889, C.G. Pringle 
2346 [HT: US-822882; IT: LE, MO-1837815, US- 
995828, US-3274342 (fragm.)]. 

= Muhlenbergia distichophylla (J. Presl) Kunth. 


Muhlenbergia annua (Vasey) Swallen, Contr. U.S. Natl. 
Herb. 29(4): 204. 1947. Basionym: Sporobolus annuus 
Vasey. 

DST.: Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Peterson, P.M. & C. R. Annable (1991); Peterson, P. 
M. (2000). 

SYN.: Muehlenbergia annua (Vasey) Swallen; Sporobolus 
annuus Vasey. 


Muhlenbergia anomala E. Fourn., Mexic. Pl. 2: 82. 1886. T- 
Protol.: Absque loco, Liehmann 687. T-Specim.: 1 of 


1. Mexico, no date, F.M. Liebmann 687 |IT: US-87238 
(fragm.)]. 

NOTE: as Muhlenbergia "anomalis". 

= Muhlenbergia quadridentata (Kunth) Trin. 


Muhlenbergia appressa C.O. Goodd., J. Wash. Acad. Sci. 
31: 504. 1941. T-Protol.: USA: Arizona: Pinal or Gila 
Co.: Devils Canyon, 28 Mar 1926, Harrison & 
Kearney 14933 [HT: US-1296986]. T-Specim.: 1 of 1. 
USA: Arizona: Pinal or Gila Co.: Devil's Canyon, 28 
Mar 1926, Harrison & Kearney 1493 [HT: US- 
1296986]. 

DST.: Mexi, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W. & R. Moran (1981); Hickman, J. C. 
(1993); Hitchcock, A. S. (1951); Peterson, P. M. & 
C. R. Annable (1991); Peterson, P. M. (2000); 
Shreve, F. & I. L. Wiggins (1964). 


Muhlenbergia arachnoidea Trin. ex Kunth, Enum. Pl. 1: 207. 
1833. T-Protol.: USA: South Carolina. 
NOTE: nom. inval., as syn. of Cinna arachnoidea Kunth. 
= Muhlenbergia expansa (Poir.) Trin. 


Muhlenbergia arenacea (Buckley) Hitchc., Proc. Biol. Soc. 
Wash. 41: 161. 1928. Basionym: Sporobolus 
arenaceus Buckley. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Correll, D. S. 
& M. C. Johnston (1970); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Hitchcock, A. S. (1951); Peterson, P. M. (2000). 


SYN.: Sporobolus  arenaceus Buckley; Sporobolus 
asperifolius var. brevifolius Vasey; Sporobolus 
asperifolius var. brevifolius Vasey; Sporobolus 


auriculatus Vasey. 


Muhlenbergia arenicola Buckley, Proc. Acad, Nat. Sci. 
Philadelphia 14: 91. 1862. T-Protol.: USA: arid places 
in western Texas. T-Specim.: | of 3. USA: Texas: arid 
places in western Texas, no date, C. Wright 735 [LT: 
PH; ILT: US-87239 (fragm.)], LT designated by 
Hitchcock, Man. Grass. U.S. 886 (1935), but without 
citing a specific sheet in a specific herbarium. 2 of 3. 
USA: Texas, Fendler 968 [ST: PH]. 3 of 3. USA: 
Texas, Fendler 969 [ST: PH]. 

DST.: Arge, Mexi, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Correll, D. S. 
& M. C. Johnston (1970); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Great 
Plains Flora Association (1986); Hitchcock, A. S. 
(1951); Peterson, P. M. & J. J. Ortiz-Diaz (1998); 
Peterson, P. M. (2000). 

SYN.: Podosemum arenicola (Buckley) Bush. 


Muhlenbergia argentea Vasey, Bull. Torrey Bot. Club 
13(12): 232. 1886. T-Protol.: Mexico: S.W. 
Chihuahua, 1885, Palmer 160 [HT: US; IT: US]. T- 
Specim.: | of 1. Mexico: Chihuahua, Aug 1885-Nov 
1885, E. Palmer 160 [HT: US-995727; IT: US- 
995726], Annotated by Reeder. 

DST.: Mexi. 
LVL. ACCPTC.: I. 


Catalogue of New World Grasses: II. 


145 


ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Herrera-Arrieta, Y. (1998); Peterson, P. M. (2000). 

SYN.: Epicampes argentea (Vasey) M_E. Jones. 


Muhlenbergia aristulata Rydb., Bull. Torrey Bot. Club 
32(11): 600. 1905. 
REPLACED NAME: Sporobolus aristatus Rydb. 
BLOCKING NAME: Muhlenbergia aristata Pers. 
= Muhlenbergia filiformis (Thurb. ex S. Watson) Rydb. 


Muhlenbergia arizonica Scribn., Bull. Torrey Bot. Club 
15(1): 8, t. A, f. 1-6. 1888. T-Specim.: | of 2. Mexico: 
State Chihuahua, 16 Sep 1885,C.G. Pringle 402 |ST: 
LE, US]. 2 of 2. USA: Arizona: mesas near the 
Mexican boundary, 11 Sep 1884, C.G. Pringle s.n. 
[LT: US-81603], LT designated by Hichc. in Mex. 
Grasses (1913). 

DST.: Mexi, USA. 

LVL. ACCPTC.:: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W. & R. Moran (1981); Hitchcock, A. S. 
(1951); Peterson, P. M. (2000); Shreve, F. & I. L. 
Wiggins (1964). 


Muhlenbergia arsenei Hitchc., Proc. Biol. Soc. Wash. 41: 
161. 1928. T-Protol.. USA: New Mexico: Sandoval 
Co.: Sulfer Springs, 2400 m, 17 Aug 1926,G. Arséne 
& A. Benedict 16405 (HT: US-1298444]. T-Specim.: | 
of 1. USA: New Mexico: Sandoval Co.: Sulphur 
Springs, 17 Aug 1926, Bro. Arsene 16405 (HT: US- 
1298444; IT: US-81602]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Cronquist, A., 
A. H. Holmgren, N. H. Holmgren & J. L. Reveal 
(1977); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. & R. Moran 
(1981); Hickman, J. C. (1993); Hitchcock, A. S. 
(1951); Munz, P. A. & D. D. Keck (1959); Munz, P. 
A. (1974); Peterson, P. M. (2000). 


Muhlenbergia articulata Scribn., Proc. Acad. Nat. Sci. 
Philadelphia 1891: 298, f. 1. 1891. T-Protol.: Mexico: 
San Luis Potpsi: Cardenas, calcareous banks, 7 Oct 
1890, Pringle 3477. T-Specim.: | of 1. Mexico: San 
Luis Potosi: Cardenas, 7 Oct 1890, C.G. Pringle 3477 
{IT: US-74629 1]. 
DST.: USA. 
LVL. ACCPTC:.: I. 
ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Peterson, P. M. (2000). 


Muhlenbergia aspera Trin. ex Kunth, Enum. PI. 1: 210. 1833. 
NOTE: nom. inval., as syn. of Sporobolus asper (Michx.) 
Kunth. 
= Sporobolus asper (P. Beauv.) Kunth. 


Muhlenbergia aspericaulis Nees ex Trin., Mem. Acad. Imp. 
Sci. Saint-Petersbourg, Ser. 6, Sci. Math., Seconde Pt. 
Sci. Nat. 6,4(1-2): 103. 1840. 
NOTE: nom. inval., as syn. of Vilfa richardsonis Trin. 
= Muhlenbergia richardsonis (Trin.) Rydb. 


Muhlenbergia asperifolia (Nees & Meyen ex Trin.) Parodi, 
Revista Fac. Agron. Veterin. (Buenos Aires) 6: 117, f. 


1. 1928. Basionym: Vilfa asperifolia Nees & Meyen ex 

Trin. 
DST.: Arge, Boli, Cana, Chil, Mexi, USA. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Beetle, A. A. (1977); Cabrera, A. L. 
(1970); Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Fernald, M. (1950); Gleason, H. A. 
& A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Gould, F. W. & R. Moran (1981); 
Great Plains Flora Association (1986); Hickman, J. 
C. (1993); Hitchcock, A. S. (1951); Hitchcock, C. L., 
A. Cronquist, M. Ownbey & J. W. Thompson (1969): 
Marticorena, C. & M. Quezada (1985); Moss, E. H. 
(1983); Munz, P. A. & D. D. Keck (1959); Munz, P. 
A. (1974); Nicora, E. G., M. E. De Paula, A. M. 
Faggi, M. del C. Mariano, A. M. Miante A., L. R. 
Parodi, C. A. Petetin, F. A. Roig & Z. Rugolo de 
Agrasar (1978); Peterson, P. M. (2000); Renvoize, S. 
A. (1998); Scoggan, H. J. (1978 [1979]); Zuloaga, F. 
O., E. G. Nicora, Z. E. Rugolo de Agrasar, O. 
Morrone, J. Pensiero & A. M. Cialdella (1994). 
Agrostis distichophylla Phil.; Agrostis eremophila 
Speg.; Muhlenbergia asperifolia (Nees & Meyen ex 
Trin.) Matthei; Sporobolus asperifolius (Nees & 
Meyen ex Trin.) Nees; Sporobolus asperifolius var. 


SYN.: 


major Vasey; Sporobolus  copiapinus — Phil.; 
Sporobolus deserticolus Phil.; — Sporobolus 
distichophyllus Phil.; Sporobolus —sarmentosus 


Griseb.; Vilfa asperifolia Meyen,; Vilfa asperifolia 
Nees & Meyen ex Trin. 


Muhlenbergia asperifolia (Nees & Meyen ex Trin.) Matthei, 
Gayana, Bot. 41(1-2): 55. 1984. Basionym: Vilfa 
asperifolia Nees & Meyen ex Trin. 

NOTE: nom. superfl. illeg. 

E. NAME: Muhlenbergia asperifolia (Nees & Meyen ex 
Trin.) Parodi. 

= Muhlenbergia asperifolia (Nees & Meyen ex Trin.) 
Parodi. 


Muhlenbergia atacamensis Parodi, Revista Argent. Agron. 
15: 248. 1948. T-Protol.: Argentina: Prov. Jujuy: La 
Quiaca, en cienagas, 3450 m, 15 Feb 1931, Parodi 
9656. T-Specim.: | of 1. Argentina: Jujuy: La Quiaca, 
3442 m, 15 Feb 1931, L.R. Parodi 9656 [HT: BAA; 
IT: BAA, SI, US-1502428]. 

= Chaboissaea atacamensis (Parodi) P.M. Peterson & 


Annable. 

Muhlenbergia atacamensis P.M. Peterson & Annable var. 
atacamensis. 

= Chaboissaea atacamensis (Parodi) P.M. Peterson & 
Annable. 

Muhlenbergia atacamensis var. brachyanthera_ Parodi, 
Revista Argent. Agron. 15: 250. 1948. T-Protol.: 


Argentina: Jujuy: Depto. de Cochinoca: Puesto del 
Marques: a 3500 m: 30 Jan 1943, Cabrera 7785. T- 
Specim.: 1 of 1. Argentina: Jujuy: Depto. Cochinoca: 
Puesto del Marques, 30 Jan 1943, Cabrera 7785 {HT: 
BAA]. 
= Chaboissaea atacamensis (Parodi) P.M. Peterson & 
Annable. 


Muhlenbergia attenuata Swallen, Ann. Missouri Bot. Gard. 
30(2): 138. 1943. T-Protol.: Panama: Chiriqui: El 
Boquete, foothills, 1000-1300 m, Hitchcock 8174 [HT: 


146 


Catalogue of New World Grasses: II. 


US-995843]. T-Specim.: | of 1. Panama: Chiriqui, 28 
Sep-7 Oct 1911, A.S. Hitchcock 8174 |HT: US- 
995843}. 

= Muhlenbergia lehmanniana Henrard. 


Muhlenbergia aurea Swallen, Contr. U.S. Natl. Herb. 29(9): 
411. 1950. T-Protol.: Guatemala: Dept. — of 
Quezaltenango: in thickets at base of vertical slopes 
along railroad, Finca Pirineos, lower S facing slopes of 
Volcan Santa Maria, between Santa Maria de Jesus 
and Calahuache, alt. 1300-1500 m, 31 Dec 1939./4. 
Steyermark 33175 (HT: F-1057948]. T-Specim.: | of 
1. Guatemala: Quezaltenango, 31 Dec 1939, JA. 
Steyermark 33175 (HT: F-1057948; IT: US-2236470]. 

DST.: Guat. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); McClure, F. A. 
(1955); Peterson, P. M. (2000); Reeder, C. G. (1994); 
Soderstrom, T. R. (1967). 


Muhlenbergia berlandieri Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(3-4): 299. 1841. T-Protol.: Mexico: in montibus, 
Berlandier s.n. T-Specim.: | of 3. Mexico: Distrito 
Federal: mountains near México, Aug 1827, 
Berlandier 676, 684 [ST: LE-TRIN-1487.01 (incl. 
fig.)], both nos. appear on the same TRIN sheet, there 
is one specimen and a fig. 2 of 3. Mexico, 26 Aug 
1827, AL. Berlandier 676 |IST: US-2557457, US- 
87241 (fragm.)]. 3 of 3. Mexico: cerca Mexico in 
montibus, Berlandier 684 [IST: MO-2974285]. 

= Muhlenbergia rigida (Kunth) Kunth. 


Muhlenbergia beyrichianum Kunth, Enum. PI. 1: 200. 1833. 
T-Protol.: Brazil: "Brasilia", Anon. T-Specim.: | of 1. 
Brasil: Province de Saint-Paul, Voyage d'Auguste de 
Saint-Hilaire de 1816 a 1821, Catal B' No 1028 [T: P]. 

= Pereilema beyrichianum (Kunth) Hitche. 


Muhlenbergia biloba Hitche., Contr. U.S. Natl. Herb. 17(3): 
294. 1913. 
REPLACED NAME: Bealia mexicana Scribn. 
BLOCKING NAME: Muhlenbergia mexicana (L.) Trin. 
= Bealia mexicana Scribn. 


Muhlenbergia botteri E. Fourn., Mexic. Pl. 2: 85. 1886. T- 
Specim.: | of 3. Mexico: Orizaba, Sep, Bott. 87 [ST: 
P; IST: US-087242 (fragm. ex P)]. 2 of 3. Mexico: 
Orizaba, Sep, Ff. Meull. 2006 [ST]. 3 of 3. Mexico: 
absque loco, Karw. /0// [ST]. 
= Muhlenbergia schreberi J.P. Gmel. 


Muhlenbergia bourgeaei E. Fourn. ex Hemsl., Biol. Cent.- 
Amer., Bot. 3: 539. 1885. 
NOTE: nom. nud. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Muhlenbergia bourgeaei E. Fourn., Mexic. Pl, 2; 86. 1886. 
T-Specim.: | of 3. Mexico: Montezuma pr. Cuantepec 
in Valle Mexicensi, 11 Oct 1866, Bourgeau 1155 
IIT: US-1645092, US-995481]. 2 of 3. Mexico: 
Gualzimalpan in valle Mexicensi, Jul, Schaffner 2 [ST: 
P}. 3 of 3. Mexico: Valle de Mexico, Desierto Viejo, 3 
Nov 1865, M. Bourgeau 1309 |LT: P; IT; US-87243 
(fragm. ex P)], LT designated by Peterson, Sys. Bot. 
Monogr. 31: 73 (1991). 

= Muhlenbergia peruviana (P. Beauv.) Steud. 


Muhlenbergia brachyphylla Bush, Amer. Midl. Naturalist 6: 

4]. 1919. T-Protol.: USA: Missouri: Webb City, 20 
Sep 1909, Palmer 2734 [HT: MO-757387]. T- 
Specim.: | of 1. USA: Missouri, Webb City, 20 Sep 
1909, Palmer 2734 [HT: MO-757387; IT: US (fragm. 
ex MO)]. 

NOTE: hom. illeg. 

E. NAME: Muhlenbergia brachyphylla (Nees) B.D. Jacks. 

CORRECT NAME: Muhlenbergia bushii R.W. Pohl. 

= Muhlenbergia bushii R.W. Pohl. 


Muhlenbergia brachyphylla_ fo. aristata EJ, Palmer & 
Steyerm., Brittonia 10(3): 110-111. 1958. T-Protol.: 
USA: Missouri, | 3/4 mi se of Spenceburg, Pike Co., 
Stillhouse Cave, 7 Oct 1956, /. A. Steyermark 8292] 
[HT: F; IT: US]. 

= Muhlenbergia frondosa (Poir,) Fernald. 


Muhlenbergia brandegei C. Reeder, Madrofio 13; 248, f. 1. 
1956. T-Protol.: Mexico: Baja California: Santa 
Margarita Isl., 6 Mar 1889, Brandegee 1/6. T-Specim.: 
1 of 1. Mexico: Baja California Sur: Santa Margarita 
Island, 6 Mar 1889, 7.8. Brandegee 16 [HT: UC; IT: 
US-995612]. 

DST.: Mexi. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W. & R. Moran (1981); Peterson, P.M. & 
C.R. Annable (1991); Peterson, P. M. (2000). 


Muhlenbergia breviaristata (Hack.) Parodi, Physis (Buenos 
Aires) 9: 219. 1928. Basionym: Muhlenbergia 
phragmitoides var. breviaristata Hack. 

DST.: Arge. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Peterson, P. M. (2000); Zuloaga, F. 
O., E. G. Nicora, Z. E. Rugolo de Agrasar, O. 
Morrone, J. Pensiero & A. M. Cialdella (1994). 

Muhlenbergia phragmitoides var. breviaristata 
Hack. 


SYN: 


Muhlenbergia breviculmis Swallen, Contr. U.S. Natl. Herb. 
29(9): 408. 1950. T-Protol.: Guatemala: Dept. 
Huehuetenango; top of Cerro Chemalito, sierra de los 
Cuchumatanes, 3.5 mi W of Santa Eulalia; 3100-3150 
m; 2 Aug 1942, JA. Steyermark 49905 [HT: US- 
1935054]. T-Specim.. 1 of 1. Guatemala: 
Huehuetenango: Cerro Chemalito, Sierra de los 
Cuchumatanes, 3.5 mi W of Santa Eulalia, 3100-3150 
m, 2 Aug 1942, Steyermark 49905 [HT: US-1935054; 
IT: F, US-2208654]. 

DST.: Guat. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); McClure, F. A. 
(1955): Reeder, C. G. (1994). 


Muhlenbergia brevifolia Scribn. ex Beal, Grass. N. Amer. 
2: 254, 255. 1896. T-Protol.: Mexico: Jalisco, Pringle 
4736. T-Specim.: 1 of 1. Mexico: Jalisco: near 
Guadalajara, 6 May 1893, C.G. Pringle 4736 [HT: 
MSC; IT: US-250853]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Peterson, P. M. (2000). 


Catalogue of New World Grasses: II. 


147 


Muhlenbergia brevifolia (Nutt.) M.E. Jones, Contr. W. Bot. 
14: 12. 1912. Basionym: Agrostis brevifolia Nutt. 
NOTE: hom. illeg. 
E. NAME: Muhlenbergia brevifolia Scribn. ex Beal. 
= Muhlenbergia cuspidata (Torr. ex Hook.) Rydb. 


Muhlenbergia brevifolia var. richardsonis (Trin.) M.E. Jones, 
Contr. W. Bot. 14: 12. 1912. Basionym: Vilfa 
richardsonis Trin. 

= Muhlenbergia richardsonis (Trin.) Rydb. 


Muhlenbergia breviligula Hitchc., N. Amer. Fl. 17(6): 458. 
1935. T-Protol.: Guatemala: Guatemala City, collected 
on clay hill, alt. 1500 m, 2 Dee 1911, Hitchcock 9063 
[HT: US-995888]. T-Specim.: 1 of 1. Guatemala, 2 
Dec 1911, A.S. Hitchcock 9063 [HT: US-995888]. 

DST.: Guat, Hond, Mexi, Nica. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); McClure, 
F. A. (1955); Molina R., A. (1975); Peterson, P. M. 
(2000); Reeder, C. G. (1994); Soderstrom, T. R. 
(1967). 


Muhlenbergia brevis C.O. Goodd., J. Wash. Acad. Sci. 31: 
505. 1941. T-Protol.: USA: New Mexico: Socorro 
County: Mogollon Mountains, on Mogollon Creek, 8 
Sep 1903, Metcalfe 671 [HT: US-495644]. T-Specim.: 
| of |. USA: New Mexico: Socorro Co.: Mogollon 
Mts., 8000 ft, 8 Sep 1903,O0.B. Metcalfe 671 [HT: US- 
495644; IT: GH, K. MO, NMC, NY, P]. 

DST.: Mexi, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Correll, D. S. 
& M. C. Johnston (1970); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J.  Valdes-Reyna (2000); 
Hitchcock, A. S. (1951); Peterson, P, M. & C. R. 
Annable (1991); Peterson, P. M. (2000). 

SYN.: Muhlenbergia schaffneri var. longiseta Scribn. 


Muhlenbergia breviseta Griseb. ex E. Fourn., Mexic. Pl. 2: 
83. 1886. T-Protol.: Mexico: Orizaba, 5 syntypes cited. 
T-Specim.: | of 5. Mexico: Veracruz: Orizaba, 1853, 
F. Miiller 1453 [LT: P; ILT: US-81601], LT 
designated by Hitchcock, N. Am. FI. 17(6): 464 
(1935), but herbarium not specified; cited by 
McVaugh, FI. Novo-Gal. 14: 233 (1983). 2 of 5. 
Mexico: Orizaba, fF. Miiller 2003 [ST: IST: US- 
81601]. 3 of 5. Mexico: Orizaba, F. Miller 2093 [ST: 
IST: US-81601]. 4 of 5. Mexico: Orizaba, Schaffner 
I/1 (ST: P]. 5 of 5. Mexico: Orizaba, Schaffner 157 
[ST: P], "in pl. ed. Hohen.". 

DST.: Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Peterson, P. M. (2000). 


Muhlenbergia brevivaginata Swallen, Bol. Soc. Bot. 
Mexico 23: 30, f. 3. 1958 [1959]. T-Protol.: Mexico: 
Jalisco: 6 mi NW of Cuautla, 1850 m, 21 Oct 1952, in 
grassy openings in open forest of Quercus 
macrophylla, on shallow soil over gently sloping 
rocks, road to Mascota, McVaugh 13671 |HT: US- 
2118495]. T-Specim.: | of 1. Mexico: Jalisco, 21 Oct 
1952, R. McVaugh 13671 (HT: US-2118495]. 


DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); McVaugh, R. 
(1983); Peterson, P. M. (2000), 


Muhlenbergia buckleyana Scribn., Contr. U.S. Natl. Herb. 
1(2): 56. 1890. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Muhlenbergia texana Buckley. 
= Muhlenbergia texana Buckley. 


Muhlenbergia bushii R.W. Pohl, Amer. Midl. Naturalist 82: 
534. 1969. 

REPLACED NAME: Muhlenbergia brachyphylla Bush. 

BLOCKING NAME: Muhlenbergia brachyphylla (Nees) 
B.D. Jacks. 

DST.: USA. 

LVL. ACCPTC.: L. 

ALSO ACCPTD. BY: Gleason, H. A. & A. Cronquist 
(1991); Great Plains Flora Association (1986); 
Peterson, P. M. (2000); Schwegman, John E. (1991). 

SYN.: Muhlenbergia brachyphylla Bush. 


Muhlenbergia caespitosa Chapm., Bot. Gaz. 3: 18. 1878. T- 
Protol.: USA: Florida: Dry pine barrens, near 
Apalachicola, /Chapman/. 

= Muhlenbergia expansa (Poir.) Trin. 


Muhlenbergia calamagrostidea Kunth, Revis. Gramin. |: 63. 
1829. 
NOTE: nom. nov. for Calamagrostis quitensis Kunth, but 
illeg.; see M. quitensis (Kunth) Hitche. 
CORRECT NAME: Calamagrostis quitensis Kunth. 
= Muhlenbergia tenuifolia (Kunth) Kunth. 


Muhlenbergia calcicola Swallen, Contr. U.S. Natl. Herb. 
29(9): 407. 1950. T-Protol.: Guatemala: Dept. 
Huehuetenango: open limestone Juniperus forest, 
along road in region of Chemal, alt. 3300 m, Sierra de 
los Chuchumatanes, 31 Dec 1940, P.C. Standley 
81703 [HT: US-1910686]. T-Specim.: | of 1. 
Guatemala: Huehuetenango: Chemal, Sierra de los 
Chuchumatanes, 3300 m, 31 Dec 1940, Standley 
81703 [HT: US-1910686; IT: US-2236500]. 

= Muhlenbergia ligularis (Hack.) Hitche. 


Muhlenbergia californica Vasey, Bull. Torrey Bot. Club 13: 
53. 1886. 

REPLACED NAME: 
brevifolia Vasey. 

DST.: USA. 

LVL. ACCPTC.: |. 

ALSO ACCPTD. BY: Abrams, L. (1923); Hickman, J. C. 
(1993); Hitchcock, A. S. (1951); Munz, P. A. & D. 
D. Keck (1959); Munz, P. A. (1974); Peterson, P. M. 


Muhlenbergia glomerata — var. 


(2000). 

SYN.: Muhlenbergia californica (Vasey) Abrams; 
Muhlenbergia glomerata var. brevifolia Vasey; 
Muhlenbergia — parishii Nasey; = Muhlenbergia 


racemosa var. brevifolia (Vasey) Beal; Muhlenbergia 
sylvatica var. californica Vasey. 


Muhlenbergia californica (Vasey) Abrams, Fl. Los Angeles 
32. 1904. Basionym: Muhlenbergia sylvatica var. 
californica Vasey. 

NOTE: hom. illeg. 
E. NAME: Muhlenbergia californica Vasey. 


148 


Catalogue of New World Grasses: II. 


= Muhlenbergia californica Vasey. 


Muhlenbergia calycina (Trin.) Trin., Gram. Unifl. Sesquifl. 
193, 296, t. 5, f Ll. 1824. Basionym: Trichochloa 
calycina Trin. 

NOTE: as "Miihlenbergia". 
= Muhlenbergia glomerata (Willd.) Trin. 


Muhlenbergia capillaris (Lam.) Trin., Gram. Unifl. 


Sesquifl. 191-192, 296, t. 5, f. 15. 1824. Basionym: 


Stipa capillaris Lam. 
NOTE: as "Miihlenbergia”. 
DST.: Cari, Guat, Mexi, USA. 
LVL. ACCPTC.: 1; cultivated. 
ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Breedlove, D. E. (1986); Catasus Guerra, L. 
(1997); Correll, D. S. & M. C. Johnston (1970); 
Darke, R. (1999); Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Espejo Serna, A., 
A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Fernald, M. (1950); Gleason, H. A. & A. Cronquist 
(1968); Godfrey, R. K. & J. W. Wooten (1979); 
Great Plains Flora Association (1986); Hitchcock, A. 
S. (1909); Hitchcock, A. S. (1936); Hitchcock, A. S. 
(1951); Liogier, H. & L. Martorell (1982); Long, R. 
W. & O. Lakela (1971); McClure, F. A. (1955); 
Peterson, P. M. (2000); Radford, A. E., H. E. Ahles 
& C. R. Bell (1968); Reeder, C. G. (1994); Small, J. 
kK. (1933); Sousa S., M. & E. F. Cabrera C. (1983); 
Wunderlin, R. P. (1998). 
Muhlenbergia polypogon (DC.) Kunth; Podosemum 
agrostideum P. Beauy.; Podosemum capillare (Lam.) 
Desv.; Stipa capillaris Lam.; Stipa diffusa Walter; 
Tosagris  agrostidea P.  Beauv.;  Trichochloa 
capillaris (Lam.) DC.; Trichochloa polypogon DC. 


SYN.: 


Muhlenbergia capillaris var. filipes (M.A. Curtis) Chapm. ex 
Beal, Grass. N. Amer. 2: 256. 1896. Basionym: 
Muhlenbergia filipes M.A. Curtis. 

= Muhlenbergia sericea (Michx.) P.M. Peterson. 


Muhlenbergia capillaris var. trichopodes (Elliott) Vasey, 
Contr. U.S. Natl. Herb. 3(1): 66. 1892. Basionym: 
Agrostis trichopodes Elliott. 

= Muhlenbergia expansa (Poir.) Trin. 


Muhlenbergia capillipes (M.E. Jones) P.M. Peterson & 
Annable, Syst. Bot. Monogr. 31: 27. 1991. Basionym: 
Sporobolus capillipesM.E. Jones. 

DST.: Mexi. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Peterson, P. M. 
& C. R. Annable (1991); Peterson, P. M. (2000). 

Sporobolus capillipes M.E. Jones; Sporobolus 
capillipes var. delicatulus M.E. Jones. 


SYN.: 


Muhlenbergia carinata Mez, Repert. Spec. Nov. Regni Veg. 
17; 213. 1921. T-Protol.: Mexico: Edo. Jalisco: bei 
Quadalajara, Pringle 2350. 

NOTE: nom. illeg. superfl. 
CORRECT NAME: Muhlenbergia scoparia Vasey. 
= Muhlenbergia scoparia Vasey. 


Muhlenbergia caxamarcensis Legaard & Sanchez Vega, 
Nordic J. Bot. 10: 437. 1990. T-Protol.: Peru: Prov. 
Cajamarca: Micuypampa, 62 km from Cajamarca 
towards Celendin, 3600 m as.l., 26 Mar 1988, 


Renvoize & Legaard 4962 |HT: PCUN; IT: AAU, K, 
MO, US]. T-Specim.: | of 1. Peru: Prov. Cajamarca: 
Micuypampa, 62 km from Cajamarca towards 
Celendin, growing in mud flat at bottom of limestone 
quarry, 3600 m, 26 Mar 1988, S.A. Renvoize & S. 
Legaard 4962 [HT: PCUN; IT: AAU, K, MO- 
3712393, US-3185350]. 

DST.: Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Legaard, S. & I. Sanchez V. (1990); 

Peterson, P. M. (2000); Tovar, O. (1993). 


Muhlenbergia ciliata (Kunth) Trin., Gram. Unifl. Sesquifl. 
193, t. 5, f. 16. 1824. Basionym: Podosemum ciliatum 
Kunth. 

DST.: Arge, Boli, Braz, Colo, Ecua, ElSa, Guat, Hond, 

Mexi, Nica, Pana. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Berendsohn, W. G. & A. E. Araniva 
de Gonzalez (1991); Brako, L. & J. L. Zarucchi 
(1993); Breedlove, D. E. (1986); Davidse, G., M. 
Sousa S. & A. O. Chater (editores generales) (1994); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Gould, F. W. & R. Moran (1981); 
Hitchcock, A. S. (1927); Jorgensen, P. M. & C. Ulloa 
U. (1994); Jorgensen, P. M. & S. Leén-Yanez (eds.) 
(1999); Macbride, J. F. (1936); McClure, F. A. 
(1955); McVaugh, R. (1983); Peterson, P.M. & C. R. 
Annable (1991); Peterson, P. M. (2000); Reeder, C. 
G. (1994); Renvoize, S. A. (1998); Zuloaga, F. O., E. 
G. Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

Muhlenbergia adspersa Trin.; Muhlenbergia ciliata 
(Kunth) Kunth; Podosemum brachyphyllum Nees; 
Podosemum ciliatum Kunth; Polypogon 
brachyphyllus (Nees) E. Fourn. ex  Hemsl.; 
Polypogon ciliatus (Kunth) Spreng.; Trichochloa 
ciliata (Kunth) Roem. & Schult. 


SYN.: 


Muhlenbergia ciliata (Kunth) Kunth, Revis. Gramin. 1: 63. 
1829. Basionym: Podosemum ciliatum Kunth. 
NOTE: isonym. 
E. NAME: Muhlenbergia ciliata (Kunth) Trin. 
= Muhlenbergia ciliata (Kunth) Trin. 


Muhlenbergia circinata Kuntze, Revis. Gen. Pl. 3(3): 356. 
1898. T-Protol.: Argentina: 1500 m nahe dem Paso 
Cruz 34 degree, O. Kuntze s.n. 
NOTE: as "Muehlenbergia" circinata. 
= Status not yet determined. 


Muhlenbergia clandestina (Biehler) Trin., Gram. Unifl. 
Sesquifl. 190-191, 296, t. 5, f. 9. 1824. Basionym: 
Agrostis clandestina Biehler. 

NOTE: as “Miithlenbergia". 

= Sporobolus clandestinus (Biehler) Hitchc. 


Muhlenbergia cleefii Lagaard, Caldasia 17(82-85): 409- 
411, f. 1. 1995. T-Protol.: Colombia: Boyaca: Sierra 
Nevada del Cocuy, Alto Valle Lagunillas, alt. 4060 m, 
26 Jun 1972, paramo pantanoso al sur de Ja Laguna 
Cuadrada, cauce relleno con Muhlenbergia fastigiata 
predominante, graminea pequefia muy abundante, 
A.M. Cleef & P.A.F. Florschiitz 5578 [HT: COL]. T- 
Specim.: | of 1. Colombia: Boyaca: Sierra Nevada del 
Cocuy, Alto Valle Lagunillas, paramo pantanosos al 
“Sur de la Laguna Cuadrada, 4060 m, 26 Sep 1972, 


Catalogue of New World Grasses: II. 


149 


A.Cleff & P.A.Florschutz 5578 (HT: COL; IT: U, US- 
2785756]. 
NOTE: as "cleefi". 
DST.: Colo. 
LVL. ACCPTC:: 1. 
ALSO ACCPTD. BY: Legaard, S. (1995); Peterson, P. M. 
(2000). 


Muhlenbergia clomena Kunth, Revis. Gramin. 1: 64. 1829. 
NOTE: nom. nov., nom. illeg. superfl.; based on Clomena 
peruviana P. Beauv. 
CORRECT NAME: Muhlenbergia peruviana (P. Beauv.) 
Steud. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Muhlenbergia coerulea (Griseb.) Mez, Repert. Spec. Nov. 
Regni Veg. 17: 213. 1921. Basionym: Epicampes 
coerulea Griseb. 

= Muhlenbergia angustata (J. Presl) Kunth. 


Muhlenbergia coloradensis Mez, Repert. Spec. Nov. Regni 
Veg. 17: 213. 1921. T-Protol.: USA: Colorado: Chiann 
Canyon, Jones. T-Specim.: | of 1. USA: Colorado, 5 
Oct 1878, ME. Jones 806 [IT: US-994951]. 

= Muhlenbergia wrightii Vasey ex J.M. Coult. 


Muhlenbergia comata (Thurb.) Thurb. ex Benth., J. Linn. 
Soc., Bot. 19: 83. 1881. Basionym: Vaseya comata 
Thurb. 

= Muhlenbergia andina (Nutt.) Hitchce. 


Muhlenbergia commutata (Scribn.) Bush, Amer. Midl. 
Naturalist 6: 61. 1919. Basionym: Muhlenbergia 
mexicana subsp. commutata Scribn. 

= Muhlenbergia frondosa fo. commutata (Scribn.) 
Fernald. 


Muhlenbergia composita Trin. ex Kunth, Enum. Pl. 1: 229. 
1833. 
NOTE: nom. inval., as syn. of Agrostis composita Poiret. 
= Sporobolus compositus (Poir.) Merr. 


Muhlenbergia confusa (E. Fourn.) Swallen, Contr. U.S. Natl. 
Herb. 29(4) 207. 1947. Basionym: Vilfa confusa E. 
Fourn. 

= Muhlenbergia minutissima (Steud.) Swallen. 


Muhlenbergia crispiseta Hitchc., N. Amer. Fl. 17(6): 440. 
1935. T-Protol.: Mexico: Chihuahua: Sierra Madre, 
collected on thin soil of porphyry ledges, 23 Oct 1887, 
Pringle 1411 (HT: US-995491]. T-Specim.: 1 of 1. 
Mexico: Chihuahua, 23 Oct 1887,C.G. Pringle 1411 
[HT: US-995491; IT: RSA, VT]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Peterson, P. M. & C. R. Annable (1991); Peterson, P. 
M. (2000). 


Muhlenbergia cualensis Y. Herrera & P.M. Peterson, Novon 
2(2): 114, f. 1. 1992. T-Protol.: Mexico: Jalisco: SSE 
of Puerto Vallarta, 14 km from the opening of the 
Zimapan Mine, 1 km W of Providencia boundary El 
Tuito-E] Caule, 20.1SN 105.15W, 31 Jan 1983, R. 
Guzman 6090 [HT: COCA; IT: IBUG, US]. T- 
Specim.: | of 1. Mexico: Jalisco, 31 Jan 1983, R. 
Guzman M. 6090 [HT: COCA; IT; IBUG, US- 
3239031]. 


DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Herrera A., Y. 
& P. M. Peterson (1992); Herrera-Arrieta, Y. (1998); 
Peterson, P. M. (2000). 


Muhlenbergia curtifolia Scribn., Bull. Torrey Bot. Club 38: 
328. 1911. T-Protol.: USA: Utah: collected between 
Kanab and Carmel, 1894, ME. Jones 6047/. T- 
Specim.: | of 1. USA: Utah, collected between Kanab 
and Carmel, 6000 ft, 13 Sep 1894, ME. Jones 6047J 
[HT: US-278700]. 

DST.: USA. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Anonymous (1986); Hitchcock, A. S. 

(1951); Peterson, P. M. (2000). 

Muhlenbergia curtifolia subsp. — curtifolia; 

Muhlenbergia curtifolia subsp. griffithsii Scribn. 


SYN.: 


Muhlenbergia curtifolia Scribn. subsp. curtifolia. 
= Muhlenbergia curtifolia Scribn. 


Muhlenbergia curtifolia subsp. griffithsii Scribn., Bull. 
Torrey Bot. Club 38: 328. 1911. T-Protol.: USA: 
Arizona: in Du Chelly Canyon: 1903, D. Griffiths 
5837. T-Specim.: 1 of 1. USA: Arizona: Apache Co., 
Canyon De Chelly, 16-17 Sep 1903,D. Griffiths 5837 
[HT: US-556088; IT: US-995190]. 

= Muhlenbergia curtifolia Scribn. 


Muhlenbergia curtisetosa (Scribn.) Bush, Amer. Mid. 
Naturalist 6: 35. 1919. Basionym: Muhlenbergia 
schreberi subsp. curtisetosa Scribn. 

= Muhlenbergia x curtisetosa (Scribn.) Bush (pro sp.). 


Muhlenbergia x curtisetosa (Scribn.) Bush (pro sp.), Amer. 
Mid]. Naturalist 6: 35. 1919. Basionym: Muhlenbergia 
schreberi subsp. curtisetosa Scribn. 

NOTE: hybrid status first recognized by Pohl, Amer. 
Midland Nat. 82: 528 (1969) [AZ schreberi x 
unknown]. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Peterson, P. M. (2000). 

SYN... Melica x curtisetosa (Scribn.) R.W. Pohl; 
Muhlenbergia curtisetosa (Scribn.) Bush; 
Muhlenbergia schreberi subsp. curtisetosa Scribn.; 
Muhlenbergia schreberi var. curtisetosa (Scribn.) 
Steyerm. & C.L. Kucera. 


Muhlenbergia curvula Swallen, Contr. U.S. Natl. Herb. 
29(9): 410. 1950. T-Protol.: Mexico: in dry oak- 
wooded slopes ca. 12 mi from Guanajuato on road to 
Santa Rosa, Guanajuato, 30 Sep 1946, H.E. Moore, Jr. 
1353 [HT: US-1963089]. T-Specim.: | of 1. Mexico: 
Guanajuato, 30 Sep 1946, H.E. Moore 1353 [HT: US- 
1963089]. 

= Muhlenbergia virescens (Kunth) Trin. 


Muhlenbergia cuspidata (Torr. ex Hook.) Rydb., Bull. 
Torrey Bot. Club 32(11): 599. 1905. Basionym: Vilfa 
cuspidata Torr. ex Hook. 

DST.: Cana, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Fernald, M. (1950); Gleason, H. A. 
& A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Great Plains Flora Association 


150 


Catalogue of New World Grasses: II. 


(1986); Hitchcock, A. S. (1935); Hitchcock, A. S. 
(1951); Hitchcock, C. L., A. Cronquist, M. Ownbey 
& J. W. Thompson (1969); Moss, E. H. (1983); 
Peterson, P. M. (2000); Scoggan, H. J. (1978 [1979}]); 
Voss, E. G. (1972), 

Agrostis brevifolia Nutt. Muhlenbergia brevifolia 
(Nutt.) M.E. Jones; Sporobolus cuspidatus (Torr. ex 
Hook.) Alph. Wood; Sporobolus gracilis (Trin.) 
Merr.; Vilfa cuspidata Torr. ex Hook.; Vilfa gracilis 
Trin. 


SYN:: 


Muhlenbergia debilis (Kunth) Trin., Gram. Unifl. Sesquifl. 
193, t. 5, f. 18. 1824. Basionym: Podosemum de bile 
Kunth. 

NOTE: basionym p. 193, comb. in tab. 
= Muhlenbergia microsperma (DC.) Kunth. 


Muhlenbergia debilis (Kunth) Kunth, Revis. Gramin. 1: 63. 
1829. Basionym: Podosemum debile Kunth. 
= Muhlenbergia microsperma (DC.) Kunth. 


Muhlenbergia decumbens Swallen, Bol. Soc. Bot. Mexico 23: 
30-32, f. 4. 1958 [1959]. T-Protol.: Mexico: 
Chihuahua: on orilla carretera, suelo migajon arenoso 
gris, km 33, Cuauhtemoc - V. Guerrero, 2200 m, 27 
Oct 1954, E. Hernandez & C. Tapia N359 [HT: US- 
2237023]. T-Specim.: | of 1. Mexico: Chihuahua: 
orilla carretera, suelo migajon arenoso gris, km 33, 
Cuahtemoc-V. Guerrero, 2200 m,, 27 Oct 1954, E. 
Hernandez X. & Tania J. N-359 [HT: US-2237023}. 

= Chaboissaea decumbens (Swallen) Reeder & C. Reeder. 


Muhlenbergia densiflora Scribn. & Merr., Bull. Div. 
Agrostol., U.S.D.A. 24: 18, f. 4. 1901. T-Protol.: 
Mexico: on lava beds, Serrania de Ajusco, Federal 
district, alt. 3000 m, 13 Aug 1897, Pringle 6675 (HT: 
US]. T-Specim.: 1 of 1. Mexico: Distrito Federal: 
Serrania de Ajusco, on lava beds, elev. 10000 ft, 13 
Aug 1897, C.G. Pringle 6675 [HT: US-316900; IT: 
LE, US-822867]. 

= Muhlenbergia dubia E. Fourn. 


Muhlenbergia depauperata Scribn., Bot. Gaz. 9: 187. 1884. 
T-Protol.: USA: Arizona, Pringle. T-Specim.: | of 2. 
USA: Arizona, 13 Sep 1884, C.G. Pringle s.n. [ST: 
US-746200]. 2 of 2. USA: Arizona: Cochise Co. or 
Santa Cruz Co.: Mustang Mts., 13 Sep 1884, Pringle 
s.n. [LT: US-994221; ILT: K, MO, MSC, NY, P, US, 
VT], LT designated (as type) by McVaugh, Fl. Novo- 
Galiciana 14: 236 (1983), but without indicating a 
specific sheet, specific sheet indicated by Peterson, 
Syst. Bot. Monogr. X (1991). 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Hitchcock, A. S. (1951); McVaugh, R. 
(1983), Peterson, P. M. & C. R. Annable (1991); 
Peterson, P. M. (2000). 

Lycurus schaffneri (E. Fourn.) Mez; Muhlenbergia 
schaffneri E. Fourn.; Muhlenbergia schaffneri E. 
Fourn. ex Hemsl.; Muhlenbergia schaffneri var. 
schaffneri. 


SYN.: 


Muhlenbergia diehlii M.E. Jones, Contr. W. Bot. 14: 13. 
1912. ‘T-Protol.: Mexico: Chihuahua: Guayanopa 
canyon, Sierra Madre Mts., /AZE. Jones] 7316. T- 


Specim.: | of 1. Mexico: Chihuahua: Guaryanopa 

Canyon, Sierra Madre Mts., 23 Sep 1903, ME. Jones 

7316 {HT: POM; IT: MO, US-1447451, US-591131]. 
= Muhlenbergia schmitzii Hack. 


Muhlenbergia diffusa Muhl., Trans. Amer. Philos. Soc. 3: 
160. 1793. 
NOTE: nom. nud. 
= Muhlenbergia schreberi J.F. Gmel. 


Muhlenbergia diffusa Willd., Sp. Pl. 1: 320. 1797. T-Protol.: 
Habitat in America boreali. 
= Muhlenbergia schreberi J.F. Gmel. 


Muhlenbergia diffusa Schreb., Beschr. Gras. 2: t. 51. 1810 
[1810]. 

NOTE: nom. illeg. superfl., et hom., descr. by Schreber, 
Gen. Pl. 1: 44 (1791) in Muhlenbergia, but without a 
species name. 

E. NAME: Muhlenbergia diffusa Willd. 

CORRECT NAME: Muhlenbergia schreberi J.F. Gmel. 

= Muhlenbergia schreberi J.F. Gmel. 


Muhlenbergia diffusa Farw., Rep. Michigan Acad. Sci. 20: 
168. 1919. 
NOTE: hom. illeg.; as comb. but basionym Agrostis diffusa 
Muhl. a hom. illeg. 
E. NAME: Muhlenbergia diffusa Willd. 
= Muhlenbergia sylvatica (Torr.) Torr. ex A. Gray. 


Muhlenbergia diffusa var. attenuata (Scribn.) Farw., Pap. 
Michigan Acad. Sci. 23: 125. 1938. Basionym: 
Muhlenbergia umbrosa subsp. attenuata Scribn. 

= Muhlenbergia frondosa fo. frondosa. 


Muhlenbergia distans Swallen, N. Amer. Fl. 17(6): 461. 
1935. T-Protol.: Mexico: Oaxaca: Las Sedas, collected 
on dry hills, alt. 1800 m, 1S Sep 1894, Pringle 5575 
[HT: US-746297]. T-Specim.: | of 1. Mexico: Oaxaca: 
dry hills, Las Sedas, alt. 6000 ft, 15 Sep 1894, C.G. 
Pringle 5575 |HT: US-746297; IT: MO-1837832, 
TAES, US-134322, US-305676]. 

DST.: Guat, Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. L6pez-Ferrari, and J, 
Valdes-Reyna (2000); McClure, F. A. (1955); 
McVaugh, R. (1983); Peterson, P. M. (2000): Reeder, 
C. G. (1994); Soderstrom, T. R. (1967). 


Muhlenbergia distichophylla (J. Presl) Kunth, Enum. Pl, 1: 
202. 1833. Basionym: Podosemum distichophyllum J. 
Pres}. 

NOTE: not Revis. Gram. suppl. published in 1834, not 
1830, as commonly cited. 

DST.: Guat, Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Peterson, P. M. (2000); Reeder, 
C. G. (1994): Soderstrom, T. R. (1967). 


SYN.: Epicampes stricta var. distichophylla (J. Presl) M.E. 
Jones;  Muhlenbergia ~~ angustifolia Swallen; 
Muhlenbergia distichophylla B.D. Jacks.; 


Podosemum distichophyllum J. Presl. 


Catalogue of New World Grasses: II. 


151 


Muhlenbergia distichophylla B.D. Jacks., Index Kew. 269. 
1894. T-Protol.: USA. 
NOTE: nom. inval., as syn. Epicampes distichophylla 
Vasey. 
E. NAME: Muhlenbergia distichophylla (J. Presl) Kunth. 
= Muhlenbergia distichophylla (J. Presl) Kunth. 


Muhlenbergia distichophylla var. mutica Scribn. ex Urbina, 
Cat. Pl. Mexic. 393. 1897. T-Protol.: Mexico, Pringle 
2360, Pringle 5577. 
NOTE: nom. nud. 
= Muhlenbergia versicolor Swallen. 


Muhlenbergia diversiglumis Trin., Mem. Acad. Imp. Sci. 
Saint-Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. 
Nat. 6,4(3-4): 298. 1841. T-Protol.: Mexico: Porto 
Pedro, Karwinsky s.n. T-Specim.: | of 1. Mexico: 
Pedro, Karwinsky s.n. [HT: LE-TRIN-1497.01; IT: 
US-84831 (fragm. ex LE-TRIN-1497.02)]. 
NOTE: reprint p. 52. 
DST.: Colo, CoRi, Ecua, ElSa, Guat, Hond, Mexi, Pana, 
Peru, Vene. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Beetle, A. A. (1977); Berendsohn, 
W. G. & A. E. Araniva de Gonzalez (1991); 
Breedlove, D. E. (1986); Davidse, G., M. Sousa S. & 
A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Hitchcock, A. S. (1927); Jorgensen, P.M. & 
C. Ulloa U. (1994); Jorgensen, P. M. & S. Leon- 
Yanez (eds.) (1999); Macbride, J. F. (1936); 
McClure, F. A. (1955); McVaugh, R. (1983); Molina 
R., A. (1975); Peterson, P. M. & C. R. Annable 
(1991); Peterson, P. M. (2000); Pohl, R. W. (1980); 
Reeder, C. G. (1994): Tovar, O. (1993). 
Brachyelytrum  pringlei Vasey ex Beal; 
Muhlenbergia triniiE. Fourn.,; Muhlenbergia trinit E. 
Fourn. ex Hemsl. 


SYN.: 


Muhlenbergia dubia E. Fourn., Biol. Cent.-Amer., Bot. 
3(29): 540. 1885. T-Protol.: Mexico: South Mexico, 
Chinantla, Ligebmann 688. T-Specim.: | of 1. Mexico: 
Oaxaca: Chinantla, May 1841, F.M. Liebmann 688 
{IT: US-91018 (fragm.)]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1; cultivated, 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977), Correll, D. S. & M. C. Johnston (1970); 
Darke, R. (1999); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Hitchcock, A. S. 
(1951); McVaugh, R. (1983); Peterson, P. M. (2000). 

Crypsinna breviglumis M.E. Jones; Crypsinna 
setiglumis M.E. Jones; Muhlenbergia acuminata 
Vasey; Muhlenbergia densiflora Scribn. & Merr.; 
Muhlenbergia dubia E. Fourn.; Muhlenbergia firma 
Beal; Sporobolus inflatus Vasey & L.H. Dewey ex 
Beal; Sporobolus ligulatus Vasey & L.H. Dewey. 


SYN.: 


Muhlenbergia dubia E. Fourn., Mexic. PI. 2: 82. 1886. T- 
Protol.: Mexico: South Mexico, Chinantla, Liebmann 
[688]. T-Specim.: | of 1. Mexico: Chinantla, May 
1841, F&M. Liebmann 688 [T: IT: US-91018 (fragm.)]. 

NOTE: isonym. 
E. NAME: Muhlenbergia dubia E. Fourn. 
= Muhlenbergia dubia E. Fourn. 


Muhlenbergia dubioides C.O. Goodd., J. Wash. Acad. Sci. 
30: 20. 1940. T-Protol.: USA: Arizona: in Box 


Canyon, 25 Sep 1936, W.A. Silveus 3490 [HT: SCS 
Hb. in Tucson]. T-Specim.: | of 1. USA: Arizona: 25 
Sep 1938, W.A. Silveus 3490 [HT: USDA-SCS herb. in 
Tucson; IT: LL, TEX, US-1761611, US-1723903]. 

= Muhlenbergia palmeri Vasey. 


Muhlenbergia dumosa Scribn. ex Vasey, Contr. U.S. Natl. 
Herb. 3(1): 71. 1892. T-Specim.: 1 of 3. USA: 
Arizona: Santa Catalina Mountains, 1884, C.G. 
Pringle s.n. {LT: US-994324], LT designated by ?; 
cited by McVaugh, FI. Novo-Gal. 14: 239 (1983). 2 of 
3. USA: Arizona, Pringle & Lemmon [ST]. 3 of 3. 
USA: Southern California, Orcurt s.n. [ST]. 

DST.: Mexi, USA. 

LVL. ACCPTC.:: 1; cultivated. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Darke, R. (1999); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Gould, F. 
W. & R. Moran (1981); Hitchcock, A. S. (1951); 
McVaugh, R. (1983); Peterson, P. M. (2000); Shreve, 
F.& I. L. Wiggins (1964). 

SYN.: Muhlenbergia dumosa var. minor Scribn. ex Beal. 


Muhlenbergia dumosa var. minor Scribn. ex Beal, Grass. N. 
Amer. 2: 261. 1896. T-Protol.: Mexico, Pringle 2355. 
T-Specim.: | of 1. Mexico: Jalisco: cool wet ledges, 
Sierra de San Estéban near Guadalajara, 7 Nov 1889, 
C.G. Pringle 2355 [HT: MSC; IT: MO-2974115, US- 
822888, US-746096, US-81625]. 

= Muhlenbergia dumosa Scribn. ex Vasey. 


Muhlenbergia durangensis Y. Herrera, Phytologia 63(6): 
457. 1987. T-Protol.: Mexico: Durango: Mun. de Los 
Charcos: Mezquital, 32 km de Charcos por el camino a 
La Guajolota, alt. 2040 m, bosque de pino-encino, 15 
Mar 1985, M. Gonzalez et al. 1659 (HT: CIIDIR]. 

DST.: Mexi. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Herrera Arrieta, 
Y. (1987); Herrera-Arrieta, Y. (1998); Peterson, P. 
M. (2000). 


Muhlenbergia elata Vasey, Contr. U.S. Natl. Herb. 1(8): 282. 
1893. T-Protol.: Mexico: near Guadalajara, 1886, E. 
Palmer 770 [HT: US]. T-Specim.: | of 1. Mexico: 
Jalisco: Guadalajara, Jul-Oct 1886, Edward Palmer 
770 [HT: US-81624; IT: LE, MO-2974305, MO- 
2974306]. 

= Muhlenbergia stricta (J. Presl) Kunth. 


Muhlenbergia elegans (Kunth) Trin., Gram. Unifl. Sesquifl. 
194, t. 5, f. 19. 1824. Basionym: Podosemum elegans 
Kunth. 

NOTE: basionym p. 194, comb. in tab. 

CORRECT NAME: Muhlenbergia elegans var. subviridis 
Kuntze. 

= Muhlenbergia rigida (Kunth) Kunth. 


Muhlenbergia elegans var. atroviolacea Kuntze, Revis. Gen. 
Pl. 3(3): 357. 1898. T-Protol.: Bolivia, Kuntze, 
syntypes. T-Specim.: | of 2. Bolivia: Tunarigebirge: 
3400 m, Kuntze [ST]. 2 of 2. Bolivia: Cochabamba 
3000 m, Kunize [ST]. 

= Muhlenbergia rigida (Kunth) Kunth. 


152 Catalogue of New World Grasses: II. 


Muhlenbergia elegans var. subviridis Kuntze, Revis. Gen. PI. Muhlenbergia eriophylla Swallen, Contr. U.S. Natl. Herb. 


3(3): 357. 1898. T-Protol.: Bolivia: Tunarigebirge: 
1600 m, Kuntze. 
REPLACED NAME: Muhlenbergia elegans (Kunth) Trin. 
= Muhlenbergia rigida (Kunth) Kunth. 


Muhlenbergia elongata Scribn. ex Beal, Grass. N. Amer. 2: 

251. 1896. T-Protol.: Mexico, Pringle s.n. T-Specim.: 
1 of 2. Mexico: Chihuahua: rocky ledges near 
Chihuahua, Aug 1885, C.G. Pringle 398 [LT: US- 
746292; ILT: LE (2 sheets), MO-2974118, TEX, US- 
78502, US-822891, US-1645960, US-746293, US- 
2959891], LT designated by A.S. Hitchcock in Contr. 
U.S. Natl. Herb. 17: 302 (1913), without explicitly 
citing a specific herbarium. 2 of 2. Mexico, C.G. 
Pringle 3477 [ST]. 

DST.: Mex1. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Peterson, P. M. 

(2000). 
SYN.: Muhlenbergia xerophila C.O. Goodd. 


Muhlenbergia eludens C. Reeder, J. Wash. Acad. Sci. 39: 
365. 1949. T-Protol.: Mexico: Minaca, in bed or rocky 
run, about 2100 m, 13 Oct 1910, Hitchcock 7768 [HT: 
US; IT: ARIZ, YU]. T-Specim.: 1 of 1. Mexico: 
Chihuahua: Minaca, 13 Oct 1910,4.8. Hitchcock 7768 
[HT: US-995423; IT: ARIZ, YU]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Hitchcock, A. S. (1951); Peterson, P. 
M. & C. R. Annable (1991); Peterson, P. M. (2000). 


Muhlenbergia emersleyi Vasey, Contr. U.S. Natl. Herb. 
3(1): 66. 1892. T-Protol.: USA: Southern Arizona, 
JD. Emersley (HT: US]. T-Specim.: 1 of 1. USA: 
Arizona, 1890, J.D. Emersley 46 [HT: US-73223]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Darke, R. (1999); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. & 
R. Moran (1981); Hitchcock, A. S. (1951); McClure, 
F. A. (1955); McVaugh, R. (1983); Molina R., A. 
(1975); Peterson, P. M. (2000); Shreve, F. & I. L. 
Wiggins (1964); Soderstrom, T. R. (1967). 


SYN.: Epicampes  distichophylla | Vasey; Epicampes 
emersleyi (Vasey) Hitche.; Epicampes subpatens 
Hitche.;  Muhlenbergia gooddingii — Soderstr.; 


Muhlenbergia vaseyana Scribn. 


Muhlenbergia enervis (Scribn. ex Beal) Hitche., Contr. U.S. 
Natl Herb. 17(3): 302. 1913. — Basionym: 
Muhlenbergia gracilis var. enervisScribn. ex Beal. 

= Muhlenbergia montana (Nutt.) Hitche. 


Muhlenbergia erecta J. Presl, Relig. Haenk. 1(4-5): 231. 
1830. T-Protol.: Mexico: "Hab. in Mexico", Haenke 
s.n. T-Specim.: | of |. Mexico, Haenke s.n. [HT: PR]. 
NOTE: hom. illeg. 
E. NAME: Muhlenbergia erecta Schreb. 
= Muhlenbergia implicata (Kunth) Trin. 


29(9): 409. 1950. T-Protol.: Mexico: Mexico: in oak 
woods at Luvianos, Temascaltepec, alt. 1100 m, 12 
May 1933, G.B. Hinton 5307 [HT:US-1840917]. T- 
Specim.: | of 1. Mexico: México: Temascaltepec 
Dist., Luvianos P.G., 1100 m, 12 May 1933, GB. 
Hinton 5307 (HT: US-1840917; IT: US-1867750]. 
DST.: Mexi. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Herrera-Arrieta, Y. (1998); Peterson, P. M. (2000). 


Muhlenbergia exilis E. Fourn., Mexic. Pl. 2: 84. 1886. T- 
Specim.: | of 8. Mexico: secus rivos pr. Jalapa, 3000 
fi, et pr. Mirador 6500 ft, Nov-Dec, Galiotti 5691 [ST: 
P; IST: US-87246 (fragm.)|. 2 of 8. Absque loco, 
Bonpland |ST: P]. 3 of 8. Absque loco, Liebmann 641 
[ST: P]. 4 of 8. Mexico: Orizaba, Bott, 1207 [ST: herb. 
Van Heurck]. 5 of 8. Mexico: Barranca pr. 
Cuernavaca, Nov, Bourg. 1298 [ST: P]. 6 of 8. 
Mexico: Cordova, Dec, Bourg. 1665 [ST: P]. 7 of 8. 
Mexico: Consoquitla, Liebmann 642 [ST: P]. 8 of 8. 
Mexico: Consoquitla, Liebmann 643 [ST: P]. 

NOTE: based on Muhlenbergia tenella Trin. as misapl. by 
Rupr. in Galiotti. 
= Muhlenbergia tenella (Kunth) Trin. 


Muhlenbergia expansa (Poir.) Trin., Gram. Unifl. Sesquifl. 
193. 1824. Basionym: Stipa expansa Poir. 

NOTE: comb. not made here. 

DST.; USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Espejo Serna, 
-A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 

Fernald, M. (1950); Godfrey, R. K. & J. W. Wooten 

(1979); Hitchcock, A. S. (1951); Peterson, P. M. 

(2000); Radford, A. E., H. E. Ahles & C. R. Bell 

(1968); Small, J. K. (1933). 

Agrostis arachnoides Poir.; Agrostis trichopodes 

Elliott; © Cinna = arachnoidea (Poir.) — Kunth; 

Muhlenbergia  arachnoidea Trin. ex Kunth; 

Muhlenbergia caespitosa Chapm.; Muhlenbergia 

capillaris var. trichopodes (Elliott) Vasey; 

Muhlenbergia  trichopodes (Elliott) © Chapm.; 

Podosemum purpureum (P. Beauv.) P. Beauv.; 

Podosemum trichopodes (Elliott) Bush; Trichochloa 

expansa (Poir.) DC.; Trichochloa purpurea P. 

Beauv.; Vilfa arachnoidea (Poir.) P. Beauv. 


SYN.: 


Muhlenbergia fasciculata Trin., Gram. Unifl. Sesquifl. 192, 
296, t. 5, f. 21. 1824. T-Protol.: "boreali-Amer. sub 
nom. Agrostis debilis". T-Specim.: 1 of 1. North 
America, Anon. [HT: LE?]. 

NOTE: "boreali-Amer. sub nom. Agrostis debilis (nec sane 
Poir)"; as "Miihlenbergia". 
= Muhlenbergia microsperma (DC.) Kunth. 


Muhlenbergia fastigiata (J. Pres!) Henrard, Meded. Rijks- 
Herb. 40: 59. 1921. Basionym: Sporobolus fastigiatus 
J. Pres. 
NOTE: as "Muehlenbergia" fastigiata. 
DST.: Arge, Boli, Chil, Ecua, Peru. 
LVL. ACCPTC:: 1. 
ALSO ACCPTD. BY: Brako, L. & J. L. Zarucchi (1993); 
Foster, R. C. (1958); Hitchcock, A. S. (1927); 
Macbride, J. F. (1936); Marticorena, C. & M. 


Catalogue of New World Grasses: II. 


153 


Quezada (1985); Matthei, O. (1984); Morden, C. W. 

& S. L. Hatch (1996); Peterson, P. M. (2000): 

Renvoize, S. A. (1998); Tovar, O. (1993); Zuloaga, 

F. O., E. G. Nicora, Z. E. Rigolo de Agrasar, O. 

Morrone, J. Pensiero & A. M. Cialdella (1994). 
SYN.: Sporobolus fastigiatus J. Pres}. 


Muhlenbergia filiculmis Vasey, Contr. U.S. Natl. Herb. 
1(8): 267. 1893. T-Protol.: USA: Colorado: El Paso 
Co.: sandy soil, Ute Pass, Green Mountains, 8500 ft 
alt., 2 Aug 1892, C.S. Sheldon 321 [HT: US]. T- 
Specim.: | of 1. USA: Colorado, El Paso Co., 2 Aug 
1892, C.S. Sheldon 321 (HT: US-81618]. 

DST.: USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Cronquist, A., 
A. H. Holmgren, N. H. Holmgren & J. L. Reveal 
(1977); Herrera-Arrieta, Y, (1998); Hitchcock, A. S. 
(1951); Peterson, P. M. (2000). 


Muhlenbergia filiculmis M.E. Jones, Contr. W. Bot. 14: 12. 
1912. T-Protol.: USA: New Mexico: Plaza Larga, 
Bigelow 778, Whipple Exped. 

NOTE: hom. illeg. superfl.; based on "Vilfa_filiculmis 
Thurber" nom. inval. as syn. Sporobolus; basis of S. 
Thurberi Scribn. which is the basionym of M. 
thurberi (Scribn.) Rydb. 

E. NAME: Muhlenbergia filiculmis Vasey. 

CORRECT NAME: Muhlenbergia thurberi (Scribn.) Rydb. 

= Muhlenbergia thurberi (Scribn.) Rydb. 


Muhlenbergia filiformis (Thurb. ex S. Watson) Rydb., Bull. 
Torrey Bot. Club 32(11): 600. 1905. Basionym: Vilfa 
depauperata var. filiformis Thurb. ex S. Watson. 

DST.: Cana, Mexi, USA. 
LVL. ACCPTC:: 1. 
ALSO ACCPTD. BY: Abrams, L. (1923); Cronquist, A., A. 

H. Holmgren, N. H. Holmgren & J. L. Reveal (1977); 

Gould, F. W. & R. Moran (1981); Great Plains Flora 

Association (1986); Hickman, J. C. (1993); 

Hitchcock, A. S. (1951); Hitchcock, C. L., A. 

Cronquist, M. Ownbey & J. W. Thompson (1969); 

Munz, P. A. & D. D. Keck (1959); Munz, P. A. 

(1974); Peterson, P. M. & C. R. Annable (1991); 

Peterson, P. M. (2000); Scoggan, H. J. (1978 [1979]). 

Muhlenbergia aristulata| Rydb.; Muhlenbergia 
filiformis var. fortis E.H. Kelso; Muhlenbergia 
idahoensis H. St. John; Muhlenbergia simplex 

(Scribn.) Rydb.; Sporobolus  aristatus Rydb.; 

Sporobolus depauperatus var. filiformis Beal; 

Sporobolus filiformis (Thurb. ex S. Watson) Rydb.; 

Sporobolus gracillimus (Thurb. ex S. Watson) Vasey, 

Sporobolus simplex Scribn.; Sporobolus simplex var. 

thermale Merr., Vilfa depauperata var. filiformis 

Thurb. ex S. Watson; Vilfa gracillima Thurb. ex S. 

Watson. 


SYN.: 


Muhlenbergia filiformis var. fortis E.H. Kelso, Rhodora 
38(452): 298. 1936. 
REPLACED NAME: Sporobolus simplex Scribn. 
= Muhlenbergia filiformis (Thurb, ex S. Watson) Rydb. 


Muhlenbergia filipes M.A. Curtis, Amer. J. Sci. Arts 44: 83. 
1843. T-Protol.: USA: "Habitat, sandy soil of the sea 
islands of North Carolina; also East Florida.". T- 
Specim.: | of 1. USA: North Carolina: Sea islands, M. 
A. Curtis sn. {HT: GH?; IT: NY, US (fragm. & 
photostat ex NY)]. 


= Muhlenbergia sericea (Michx.) P.M. Peterson. 


Muhlenbergia firma Beal, Grass. N. Amer. 2: 243. 1896. T- 
Protol.: Mexico: Oaxaca, Pringle 49/4. T-Specim.: | 
of 1. Mexico: state of Oaxaca: near M. Palmeri, 
summit ledges, Sierra de San Felipe, 10500 ft, 18 Sep 
1894, C.G. Pringle 4914 [HT: MSC; IT: LE, MO- 
2974120, US-746252, US-822866, US-25084 1]. 

= Muhlenbergia dubia E. Fourn. 


Muhlenbergia flabellata Mez, Repert. Spec. Nov. Regni 
Veg. 17: 213. 1921. T-Protol.: Costa Rica: Cerro de 
Bueno Vista, Pittier 3372. T-Specim.: | of 1. Costa 
Rica: San José: Cerro de Bueno Vista, 3000 m, 19 Jan 
1891, H. Pittier 3372 (HT: B; IT: LE, US-577110]. 

DST.: CoRi, Pana. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Herrera-Arrieta, 
Y. (1998); Peterson, P. M. (2000); Pohl, R. W. 
(1980); Reeder, C. G. (1994). 


Muhlenbergia flavida Vasey, Contr. U.S. Natl. Herb. 1(8): 
282. 1893. T-Protol.: Mexico: Rio Blanco, 1886, £. 
Palmer 645 (HT: US]. T-Specim.: | of 1. Mexico: 
Jalisco: Rio Blanco, Jun-Oct 1886, E. Palmer 645 
[HT: US-81623; IT: K, LE, MO-2974124, NY, P, US- 
822896, US-995427]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Peterson, P. M. & C. R. 
Annable (1991); Peterson, P. M. (2000). 


Muhlenbergia flavida UNRANKED strictior (Scribn. ex 
Beal) Hitche., Contr. U.S. Natl. Herb. 17(3): 290. 
1913. Basionym: Muhlenbergia strictior Scribn. ex 
Beal. 

NOTE: trinomial. 

= Muhlenbergia strictior Scribn. ex Beal. 


Muhlenbergia flaviseta Scribn., U.S.D.A. Div. Bot. Bull. 8: 
Il, t. 7. 1897. T-Protol.: Mexico: found at Dos 
Cajetes, 30 mi nearly west from Durango, on the road 
to Otinapa, alt. 8500 ft, among pines and oaks, 23 Oct 
1896 1896, E. Palmer 834. T-Specim.: | of 1. Mexico: 
Durango: collected at the city of Durango and vicinity, 
Dos Cajetes, Apr-Nov 1896, E. Palmer 834 [HT: US- 
81622; IT: LE, MO-2974125, US-476182, US- 
746184]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Herrera-Arrieta, Y. (1998); Peterson, P. M. (2000). 


Muhlenbergia flexuosa Hitchc., Contr. U.S. Natl. Herb. 

24(8): 388. 1927. T-Protol.; Peru: at summit of rocky 
crest, Huacachi Estacion near Muna, alt. 2000 m, 20 
May-1 Jun 1923, J. F. Macbride 3874 [HT: F- 
534937]. T-Specim.: | of 1. Peru: Huacachi: Muna, 20 
May-1 Jun 1923, J. Francis Machride 3874 [HT: F- 
534937; IT: LE, US-1256339, F-050164]. 

DST.: Peru. 

LVL. ACCPTC.: 1. 


154 


Catalogue of New World Grasses: II. 


ALSO ACCPTD. BY: Hitchcock, A. S. (1927); Macbride, 
J. F. (1936); Peterson, P. M. (2000); Tovar, O. 
(1993). 


Muhlenbergia foliosa (hort. ex Roem. & Schult.) Trin., 
Gram. Unifl. Sesquifl. 190, 297. 1824. Basionym: 
Agrostis foliosa hort. ex Roem. & Schult. 

NOTE: as "Mihlenbergia". 
= Muhlenbergia mexicana var. filiformis (Torr.) Scribn. 


Muhlenbergia foliosa fo. ambigua (Torr.) Wiegand, Rhodora 
26(301): 1. 1924. Basionym: Muhlenbergia ambigua 
Torr. 
= Muhlenbergia mexicana var. filiformis (Torr.) Scribn. 


Muhlenbergia foliosa subsp. ambigua (Torr.) Scribn., 
Rhodora 9(98): 20. 1907. Basionym: Muhlenbergia 
ambigua Torr. 

= Muhlenbergia mexicana var. filiformis (Torr.) Scribn. 


Muhlenbergia foliosa subsp. setiglumis (S. Watson) Scribn., 
Rhodora 9(98): 20. 1907. Basionym: Muhlenbergia 
sylvatica var. setiglumis S. Watson. 

= Muhlenbergia mexicana var. filiformis (Torr.) Scribn. 


Muhlenbergia fournieriana Hitche., J. Wash. Acad. Sci. 
23(10): 453. 1933. 
REPLACED NAME: Epicampes berlandieri E. Fourn. 
BLOCKING NAME: Muhlenbergia berlandieri Trin. 
= Muhlenbergia robusta (E. Fourn.) Hitche. 


Muhlenbergia fragilis Swallen, Contr. U.S. Natl. Herb. 
29(4): 206. 1947. T-Protol.: USA: Texas: Brewster 
Co.: Sunny Glen, west of Alpine, 29 Sep 1935, B.H. 
Warnock 235. T-Specim.: 1 of 1. USA: Texas: 
Brewster Co.: Sunny Glen, W of Alpine, 29 Sep 1935, 
B.H. Warnock 235 {HT: US-1829290; IT: GH]. 

DST.: Mexi, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Breedlove, D. E. (1986); Correll, D. S. & M. 
C. Johnston (1970); Davidse, G., M. Sousa S. & A. 
O. Chater (editores generales) (1994); Gould, F. W. 
& R. Moran (1981); Hickman, J. C. (1993); 
Hitchcock, A. S. (1951); McVaugh, R. (1983); 
Peterson, P. M. & C. R. Annable (1991); Peterson, P. 
M. (2000); Reeder, C. G. (1994): Shreve, F. & I. L. 
Wiggins (1964). 


Muhlenbergia frondosa (Poir.) Fernald, Rhodora 45(534): 
235, pl. 750. 1943. Basionym: Agrostis frondosa Poir. 

DST.: Cana, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Fernald, M. (1950); Gleason, 
H. A. & A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Godfrey, R. K. & J. W. Wooten 
(1979); Great Plains Flora Association (1986); 
Hitchcock, A. S. (1951); Peterson, P. M. (2000); 
Radford, A. E., H. E. Ahles & C. R. Bell (1968); 
Schwegman, John E. (1991); Scoggan, H. J. (1978 
[1979]). 

SYN.: Agrostis frondosa Poir.; Agrostis lateriflora Michx.; 
Calamagrostis compressa DO6l\l; Cinna lateriflora 
(Michx.) Kunth; Muhlenbergia brachyphylla fo. 
aristata EJ, Palmer & Steyerm.; Muhlenbergia 
lateriflora Trin. ex Kunth; Vilfa lateriflora (Michx.) 
P. Beauv. 


Muhlenbergia frondosa fo. commutata (Scribn.) Fernald, 
Rhodora = 45(534): 235. 1943. Basionym: 
Muhlenbergia mexicana subsp. commutata Scribn. 

DST.: USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Fernald, M. (1950); Hitchcock, A. S. 
(1951); Scoggan, H. J. (1978 [1979]); Voss, E. G. 
(1972). 

SYN.: 9 Muhlenbergia =commutata (Scribn.) — Bush; 
Muhlenbergia mexicana fo. commutata (Scribn.) 
Wiegand; Muhlenbergia mexicana subsp. commutata 
Scribn.; Muhlenbergia mexicana var. commutata 
(Scribn.) Farw. 


Muhlenbergia frondosa (Poir.) Fernald fo. frondosa. 

DST.: Cana, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Fernald, M. (1950); Scoggan, H. J. 
(1978 [1979]). 

SYN.: Muhlenbergia diffusa var. attenuata (Scribn.) Farw.; 
Muhlenbergia sylvatica to. attenuata (Scribn.) E.J. 
Palmer & Steyerm.; Muhlenbergia umbrosa_ fo. 
attenuata (Scribn.) Deam; Muhlenbergia umbrosa 
subsp. attenuata Scribn. 


Muhlenbergia gigantea (E. Fourn.) Hitchc.. N. Amer. FI. 
17(6): 460, 1935. Basionym: Epicampes gigantea E. 
Fourn. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J.  Valdes-Reyna (2000); 
McVaugh, R. (1983); Peterson, P. M. (2000); Reeder, 
C. G. (1994); Soderstrom, T. R. (1967). 

SYN.: Epicampes bourgeaei var. mutica E. Fourn.; 
Epicampes ehrenbergii Mez; Epicampes expansa E. 
Fourn.; Epicampes expansa FE. Fourn. ex Hemsl.; 
Epicampes gigantea E. Fourn.; Epicampes gigantea 
E. Fourn. ex Hemsl.; Epicampes laxiuscula E. 
Fourn.; Epicampes laxiuscula E. Fourn. ex Hemsl.; 
Muhlenbergia alta Hitche.; Muhlenbergia magna 
Hitche. 


Muhlenbergia glabrata (Kunth) Trin., Gram. Unifl. Sesquifl. 
194, t. 5, f. 20. 1824. Basionym: Podosemum 
glabratum Kunth, 

NOTE: basionym p. 194, comb. in tab, 
= Muhlenbergia rigida (Kunth) Kunth. 


Muhlenbergia glabriflora Scribn., Rhodora 9(98): 22-23. 
1907. T-Protol.: USA: Texas, /. Reverchon 5 [HT: US- 
81621]. 

NOTE: as "glabrifloris". 

DST.: Mexi, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Fernald, M. (1950); Gleason, 
H. A. & A, Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Hitchcock, A. S. (1951); Peterson, 
P.M. (2000). 


Muhlenbergia glauca (Nees) B.D. Jacks., Index Kew. 2: 
269. 1895. Basionym: Podosemum glaucum Nees. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Peterson, P. M. (2000). 


Catalogue of New World Grasses: II. 


155 


SYN.: Agrostis glauca (Nees) Steud.; Muhlenbergia glauca 
(Nees) Mez; Muhlenbergia huachucana Vasey; 
Muhlenbergia lemmonii  Scribn.; = Podosemum 
glaucum Nees. 


Muhlenbergia glauca (Nees) Mez, Repert. Spec. Nov. Regni 
Veg. 17: 214. 1921. Basionym: Podosemum glaucum 
Nees. 
NOTE: isonym. 
E. NAME: Muhlenbergia glauca (Nees) B.D. Jacks. 
= Muhlenbergia glauca (Nees) B.D. Jacks. 


Muhlenbergia glomerata (Willd.) Trin., Gram. Unifl. 
Sesquifl. 191, 297, t. 5, f. 10. 1824. Basionym: 
Polypogon glomeratus Willd. 

NOTE: as "Miihlenbergia". 

DST.; Cana, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Cody, W. J. (1996); Cronquist, A., 
A. H. Holmgren, N. H. Holmgren & J. L. Reveal 
(1977); Fernald, M. (1950); Gleason, H. A. & A. 
Cronquist (1991); Godfrey, R. K. & J. W. Wooten 
(1979); Great Plains Flora Association (1986); 
Hitchcock, A. S. (1951); Hultén, E. (1968); Moss, E. 
H. (1983); Peterson, P. M. (2000); Radford, A. E., H. 
E. Ahles & C. R. Bell (1968); Scoggan, H. J. (1978 
[1979]): Voss, E. G. (1972); Welsh, S. L. (1974). 

Alopecurus — glomeratus — (Willd.) Poir.; 
Dactylogramma  cinnoides Link; Muhlenbergia 
calycina (Trin.) Trin.; Muhlenbergia glomerata var. 
cinnoides (Link) FJ.  Herm.; Muhlenbergia 
glomerata var. glomerata; Muhlenbergia racemosa 
subsp. violacea Scribn.; Muhlenbergia racemosa vat. 
cinnoides (Link) B. Boivin; Muhlenbergia setosa 

(Biehler) Trin. ex B.D. Jacks.; Muhlenbergia setosa 

var. cinnoides (Link) Fernald; Podosemum 

glomeratum (Willd.) Link; Polypogon glomeratus 

Willd.; Polypogon setosus Biehler; Trichochloa 

calycina Trin., Trichochloa glomerata (Willd.) Trin. 


SYN:: 


Muhlenbergia glomerata var. brevifolia Vasey, Bot. Gaz. 7: 
92. 1882. T-Protol.: USA: Collected in S._ E. 
California, Parish. T-Specim.: | of 1. USA: California, 
San Bernardino Co., 15 Aug 1882,S.B. Parish, W F. 
Parish 1028 (HT: US-81620]. 

CORRECT NAME: Muhlenbergia californica Vasey. 
= Muhlenbergia californica Vasey. 


Muhlenbergia glomerata var. cinnoides (Link) F.J. Herm., 
Rhodora 48: 64. 1946. Basionym: Dactylogramma 
cinnoides Link. 

= Muhlenbergia glomerata (Willd) Trin. 


Muhlenbergia glomerata (Willd.) Trin. var. glomerata. 
= Muhlenbergia glomerata (Willd.) Trin. 


Muhlenbergia glomerata var. ramosa Vasey, Descr. Cat. 
Grass. U.S. 40. 1885. T-Protol.: USA, syntypes. T- 
Specim.: 1 of 1. USA: Illinois / Colorado / Montana, 
no date, G.R. Vasey s.n. [ST: US-994606]. 

= Muhlenbergia racemosa (Michx.) Britton, Sterns & 
Poggenb. 


Muhlenbergia gooddingii Soderstr., Contr. U.S. Natl. Herb. 
34(4): 115. 1967. T-Protol.: USA: Pima Co.: collected 
in canyon north of Moristo Canyon, Baboquivari 
Mountains, alt. 4000 ft, 19 Nov 1945,L.N. Goodding 
462-45 [HT: YU; IT: ARIZ, NY]. T-Specim.: | of 1. 


USA: Arizona: Pima Co.: Baboquivari Mountains, 
canyon north of Moristo Canyon, steep north-facing 
slopes, elev. 4000 ft, 19 Nov 1945, L.N. Goodding 
462-45 (HT: YU; IT: ARIZ, MO-5073874, NY, US- 
2550348). 

= Muhlenbergia emersleyi Vasey. 


Muhlenbergia gracilis (Kunth) Trin., Gram. Unifl. Sesquifl. 
193, t. Sa, f. 6. 1824. Basionym: Podosemum gracile 
Kunth. 

NOTE: basionym p. 193, comb. in tab. 
= Muhlenbergia quadridentata (Kunth) Trin. 


Muhlenbergia gracilis var. breviaristata Vasey, Cat. Pl. 54. 
1874. T-Protol.: USA: Colorado: Twin Lakes: 1873, 
Wolf 1090. T-Specim.: | of 1. USA: Colorado, 1873, 
J. Wolfe 1090 [IT: US-81619], Also in Rothrock, J. T., 
Catalogue 284. 1878. 1098 given as collection number. 
CORRECT NAME: Muhlenbergia subalpina Vasey. 
= Muhlenbergia montana (Nutt.) Hitchc. 


Muhlenbergia gracilis var.enervis Scribn. ex Beal, Grass. N. 
Amer. 2: 242. 1896. T-Protol.: Mexico: Chihuahua, 
Pringle 1413. T-Specim.: | of 2. Mexico: Chihuahua, 
7 Oct 1887, C.G. Pringle 1413 |HT: US-995814]. 2 of 
2. Mexico: Chihuahua: Sierra Madre, dry ledges, 
Pringle 1413 [T]. 

= Muhlenbergia montana (Nutt.) Hitchce. 


Muhlenbergia gracilis var. major Vasey, Rep. U.S. Geogr. 
Surv., Wheeler 6: 284. 1878. T-Protol.: USA: Arizona, 
744 [Rothrock Exped.]. T-Specim.: | of 1. USA: 
Arizona, Sep 1874, 7. Rothrock 744 [IT: US- 
995140], Also in Rothrock, J. T. Catalogue 284. 1878. 

= Muhlenbergia montana (Nutt.) Hitche. 


Muhlenbergia gracillima Torr., Pacif. Railr. Rep. 4: 155. 
1857. T-Protol.: USA: Texas: "Llano Estacado, and 
near the Antelope hills of the Canadian River, Sep, 2 
syntypes. T-Specim.: | of 2. USA: Texas: Llano 
Estacado, 1853-1854,..M. Bigelow s.n. [ST: NY; IST: 
US (fragm. ex NY)]. 2 of 2. USA: Texas: Antelope 
Hills, 1853-1854, /.M. Bigelow s.n. [ST: NY; IST: US 
(fragm. ex NY )]. 

= Muhlenbergia torreyi (Kunth) Hitche. ex Bush. 


Muhlenbergia grandis Vasey, Contr. U.S. Natl. Herb. 1(8): 
283. 1893. T-Specim.: | of 3. Mexico: Jalisco: Rio 
Blanco, Jun-Oct 1886, £. Palmer 5/5 [LT: US-73222; 
ILT: LE, MO-1837830, US-822914], LT designated 
by Hitchcock, Contr. U.S. Natl. Herb. 17(3): 301 
(1913), but without specifying a specific sheet; also by 
Soderstrom, Contr. U.S. Natl. Herb. 34: 161 (1967). 
who designated a specific sheet (as type). 2 of 3. 
Mexico; Jalisco, 1888, C.G. Pringle 1760 |ST: US}. 3 
of 3. Mexico: Jalisco: Rio Blanco, 1886, E. Palmer 
515A (IST: US-999059, US-822915]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Peterson, P. M. (2000): 
Soderstrom, T. R. (1967). 

SYN.: Epicampes bourgeaei var. grandis (Vasey) M.E. 
Jones; Epicampes grandis (Vasey) Beal. 


Muhlenbergia gypsophila Reeder & C. Reeder, Madrofo 
18: 186. 1966. T-Specim.: | of 1. Mexico: Nuevo 


156 


Catalogue of New World Grasses: II. 


Leon, 30 Oct 1964, /.R. Reeder et al. 3963 [IT: US- 
2524092]. 
DST.: Mexi. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Peterson, P. M. (2000). 


Muhlenbergia herzogiana Henrard, Meded. Rijks-Herb. 40: 
58. 1921. T-Protol.: Bolivia; Zwerggras auf 
Schieferschutt am  Ufer eines  Glazialtiimpels 
(Cordillera Santa Benito), 4000 m, Jun 1911, 7 
Herzog 2226 {HT: L]. T-Specim.: 1 of 1. Bolivia: 
Cordillera de Santa Bonita, Jun 1911, 7. Herzog 2226 
{HT: L; IT: US-87248 (fragm. ex L), US-1161342]. 

NOTE: as "Muehlenbergia". 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Muhlenbergia hintonii Swallen, Contr. U.S. Natl. Herb. 
29(9): 409. 1950. T-Protol.: Mexico: México: in 
meadow, Crucero, Temascaltepec, alt. 2880 m, 30 Dec 
1932, GB. Hinton 3059 [HT: US-1840838}. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Peterson, P. M. (2000). 


Muhlenbergia holwayorum Hitche., Contr. U.S. Natl. Herb. 
24(8): 389. 1927. T-Protol.: Bolivia: collected at 
Sorata, 16 Apr 1920, £. WD. Holway & M.M. Holway 
530 [HT: US-1108445]. T-Specim.: 1 of 1. Bolivia: 
Sorata, 16 Apr 1920, E.W.D. Holway et al. 530 (HT: 
US-1108445)]. 

= Muhlenbergia rigida (Kunth) Kunth. 


Muhlenbergia huachucana Vasey, Contr. U.S. Natl. Herb. 
3(1): 69. 1892, T-Protol.: USA: Arizona: Huachuca 
Mts., .G. Lemmon. T-Specim.: | of 1. USA: Arizona, 
Jul 1882, 4G. Lemmon 2915 (HT: US-81617]. 
= Muhlenbergia glauca (Nees) B.D. Jacks. 


Muhlenbergia idahoensis H. St. John, Fl. S. E. Washington 
50. 1937. T-Protol.: USA: Idaho: meadow along creek, 
4500 ft., Zaza, Craig Mts. Nez Perce Co.: 9 Oct 1927, 
St. John 9085 (HT: WSU]. T-Specim.: 1 of 1. USA: 
Idaho: Nez Perce Co.: Zaza, Craig Mts., 1372 m, 9 Oct 
1927, St. John 9085 |HT: WSU; IT: US-3412355 
(fragm.)]. 

= Muhlenbergia filiformis (Thurb. ex S. Watson) Rydb. 


Gram. Unifl. 
1824. Basionym: 


Muhlenbergia implicata (Kunth) Trin., 
Sesquifl. 193: t. Sa, f. 26. 
Podosemum implicatum Kunth. 

NOTE: basionym p. 193, comb. in tab. 

DST.: Colo, CoRi, ElSa, Guat, Hond, Mexi, Nica, Pana, 

Vene. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Breedlove, D. E. (1986); Davidse, 
G., M. Sousa S. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); McClure, F. A. (1955); 
McVaugh, R. (1983); Peterson, P. M. & C. R. 
Annable (1991); Peterson, P. M. (2000); Reeder, C. 
G. (1994), 

Agrostis implicata (Kunth) Spreng.; Muhlenbergia 
erecta J. Presl; Muhlenbergia implicata (Kunth) 


SYN:: 


Kunth; Podosemum implicatum Kunth; Trichochloa 
implicata (Kunth) Roem. & Schult. 


Muhlenbergia implicata (Kunth) Kunth, Revis. Gramin. 1: 
63. 1829. Basionym: Podosemum implicatum Kunth. 
NOTE: isonym. 
E. NAME: Muhlenbergia implicata (Kunth) Trin. 
= Muhlenbergia implicata (Kunth) Trin. 


Muhlenbergia inaequalis Soderstr., Contr. U.S. Natl. Herb. 
34(4): 142. 1967. T-Protol.: Venezuela: Trujillo: 
collected in Quebrada de Duri, 11 Nov 1939, A. 
Mueller 973 (HT: US-2380325; IT: VEN]. T-Specim.: 
1 of 1. Venezuela: Trujillo, 11 Nov 1939, 4.8. Miiller 
973 [HT: US-2380325; IT: MO-fragm., VEN]. 

DST.: Colo, Vene. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Peterson, P. M. (2000); Soderstrom, 
T. R. (1967). 


Muhlenbergia involuta Swallen. 
= Muhlenbergia x involuta Swallen. 


Muhlenbergia x involuta Swallen (pro sp.), Amer. J. Bot. 
19(5): 436, f. 2. 1932. T-Protol.: USA: Texas: 
collected in draws in hills, 20 mi. northeast of San 
Antonio, 10 Oct 1931, WA. Silveus 358 [HT: US- 
1501595]. T-Specim.: | of 1. USA: Texas: Bexar Co., 
10 Oct 1931, W.A. Silveus 358 [HT: HT: US-1501595; 
IT: US-1501594, US-3215628, US-32784 10]. 

NOTE: possibly hybridogenic [Af lindheimeri x M. 
reverchonii| according to Correll & Johnston, p. 232 
Man. Vasc. PI. Texas (1970). 

DST.: USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Darke, R. (1999); Hitchcock, 
A. S. (1951); Peterson, P. M. (2000); Soderstrom, T. 
R. (1967). 

SYN.: Muhlenbergia involuta Swallen. 


Muhlenbergia iridifolia Soderstr., Contr. U.S. Natl. Herb. 
34(4): 145. 1967. T-Protol.: Mexico: Jalisco: steep 
mountains 11-12 mi S of Talpa de Allende, in the 
headwaters of a west branch of Rio de Talpa, barranca 
above a rapid clear stream, in dense forest of Quercus, 
Carpinus, Distylium, Magnolia, Podocarpus, with pine 
forest on the ridges above, abundant in the pine forest, 
in clumps, inflorescence purple, alt. 1200-1700 m, 23- 
25 Nov 1960, McVaugh 2/313 [HT: MICH; IT: US]. 
T-Specim.: | of 1. Mexico: Jalisco, 25 Nov 1960, R. 
McVaugh 2/313 [HT: MICH; IT: US-2432724, US- 
2960226]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Peterson, P. M. (2000); 
Soderstrom, T. R. (1967). 


Muhlenbergia jaime-hintonii P.M. Peterson & Valdés- 
Reyna, Sida 18(3): 686, f. 1. 1999. T-Protol.: Mexico: 
Nuevo Leén: Municipio General Zaragoza: La Joya, 
Cuesta Blanca, approximately 15 km of Aramberri on 
road towards Zaragoza 23°59" 37.1"N-99°47'38.7"W, 
1345m, 31 Oct 1998, J. Valdés-Reyna & M.A. 
Carranza Pérez 2560 |HT: ANSM; IT: US]. T- 
Specim.: 1 of |. Mexico: Nuevo Leén: La Joya, Cuesta 


Catalogue of New World Grasses: II. 


157 


Blanca, approximately 15 km de Arramberri Rumba a 
Zaragoza, 31 Oct 1998, JValdés-Reyna & 
M.A.Carranza Pérez 2560 (IT: US-3377556, US- 
3377557]. 
DST.: Mexi. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Peterson, P. M. 
& J. Valdés-Reyna (1999). 


Muhlenbergia jaliscana Swallen, Bol. Soc. Bot. Mexico 23: 
32-35, f. 5. 1958 [1959]. T-Protol.: Mexico: Jalisco: 
Sierra de Manantlan, 15-20 mi SE of Aytlan, on the 
bajada S and W of the divide between Aserradero San 
Miguel Uno and Durazno, 1700 m, 7 Nov 1952, on the 
steep west-facing slope in pine forest, 7 Nov 1952, 
McVaugh 13991 (HT: US-2118498]. T-Specim.: 1 of 
1. Mexico: Jalisco: Sierra de Manantlan, 15-20 mi SE 
of Aytlan, on the bajada S and W of the divide 
between Aserradero San Miguel Uno and Durazno, alt. 
1700 m, 7 Nov 1952, R. McVaugh 13991 (HT: US- 
2118498]. 

DST.: Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J, Valdes-Reyna (2000); McVaugh, R. 
(1983); Peterson, P. M. (2000). 


Muhlenbergia japonica Steud., Syn. Pl. Glumac. 1: 422. 
1854. T-Specim.: | of 1. Japan, P.F. von Siebold s.n. 
{HT: L]. 
NOTE: known only from cultivation. 
DST.: USA. 
LVL. ACCPTC.: 1; cultivated, introduced. 
ALSO ACCPTD. BY: Peterson, P. M. (2000). 


Muhlenbergia jonesii (Vasey) Hitche., Fl. Calif. 1: 111. 
1912. Basionym: Sporobolus jonesii Vasey. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Abrams, L. (1923); Herrera-Arrieta, 
Y. (1998); Hickman, J. C. (1993); Hitchcock, A. S. 
(1951); Munz, P. A. & D. D. Keck (1959); Peterson, 
P. M. (2000). 

SYN.: Sporobolus jonesii Vasey. 


Muhlenbergia lanata (Kunth) Hitche., N. Amer. Fl. 17: 459. 
1935. Basionym: Agrostis lanata Kunth. 
= Muhlenbergia pubescens (Kunth) Hitche. 


Muhlenbergia lateriflora Trin. ex Kunth, Enum. Pl. 1: 207. 
1833. 
NOTE: nom. inval., as syn. of Cinna lateriflora Kunth. 
= Muhlenbergia frondosa (Poir.) Fernald. 


Muhlenbergia laxa Hitchc., N. Amer. FI. 17(6): 445. 1935. 
T-Protol.: Mexico: Orizaba: in moist places: October, 
Botteri 1259 [HT: US-822840]. T-Specim.: 1 of 1. 
Mexico: Orizaba, M. Botteri 1259 [HT: US-822840]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. L6pez- 
Ferrari, and J. Valdes-Reyna (2000); Peterson, P. M. 
(2000). 


Muhlenbergia laxiflora Scribn., Zoe 4(4): 389. 1894. T- 
Protol.: Mexico: Baja California Sur: La Chuparosa: 
17 Oct 1893, Brandegee 74. T-Specim.: 1 of 2. 


Mexico: La Chuparosa, 17 Oct 1893, Brandegee 74 
[LT: UC; ILT: US (fragm. ex UC)], LT designated by 
A.S. Hitchcock, Contr. U.S. Natl Herb. 17: 298 
(1913). A Pringle collection is also cited in the 
Scribner's protologue, without the two collections 
being clearly distinguished as types, except in the 
context of the title of the article. 2 of 2. Mexico: 
Chihuahua: Cool slopes of Sierra Madre, 2286 to 2591 
m, 23 Sept 1887,C.G. Pringle 1412 [IST: US-822919, 
US-91919 (fragm. ex UC)]. 
= Muhlenbergia mucronata (Kunth) Trin. 


Muhlenbergia lehmanniana Henrard, Meded. Rijks-Herb. 
40: 59. 1921. T-Protol.: Colombia: prob. prope 
Popayan, Lehmann 1267 (HT: L]. T-Specim.: | of 1. 
Colombia: Popayan?, Lehmann 1267 |HT: L-908.350- 
519; IT: US-72979 (fragm. ex L), US-72977 (fragm. 
ex L), US-72978 (fragm. ex K)]. 

NOTE: as "Muehlenbergia Lehmanniana". 

DST.: Vene. 

LVL. ACCPTC:.: 1. 

ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Jorgensen, P. M. 
& S. Leén-Yanez (eds.) (1999); Peterson, P. M. 
(2000); Pohl, R. W. (1980); Reeder, C. G. (1994); 
Soderstrom, T. R. (1967). 

SYN.: Muhlenbergia attenuata Swallen, Muhlenbergia 
multinodis Asp. 


Muhlenbergia lemmoniiScribn., Contr. U.S, Natl. Herb. 1(2): 
56. 1890. T-Specim.: | of 2. USA: Arizona, Lemon 
and wife 2918 [LT: US-746111; ILT: US-994769, US- 
994778, US-746113], Lectotypified by A. S. 
Hitchcock, North American Flora 17(pt. 6): 449. 1935. 
2 of 2. USA: Texas: Runnels Co.: Ballinger, G.C. 
Nealley [ST]. 

= Muhlenbergia glauca (Nees) B.D. Jacks. 


Muhlenbergia leptoura (Piper) Hitche., N. Amer. Fl. 17(6): 
468. 1935. Basionym: Epicampes leptoura Piper. 
= Muhlenbergia rigens (Benth.) Hitchc. 


Muhlenbergia ligularis (Hack.) Hitche., Contr. U.S. Natl. 
Herb. 24(8): 388. 1927. Basionym: Sporobolus 
ligularis Hack. 

DST.: Arge, Boli, Colo, Ecua, Guat, Peru, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Brako, L. & J. L. Zarucchi (1993); 
Foster, R. C. (1958); Hitchcock, A. S. (1927); 
Jorgensen, P. M. & C. Ulloa U. (1994); Jorgensen, P. 
M. & S. Leon-Yanez (eds.) (1999); Macbride, J. F. 
(1936); Peterson, P. M. & C. R. Annable (1991); 
Peterson, P. M. (2000); Renvoize, S. A. (1998); 
Tovar, O. (1993); Young, K. R. & B. Leon (1990); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

Muhlenbergia breviculmis Swallen; Muhlenbergia 
calcicola  Swallen; Muhlenbergia minuscula HU. 
Scholz; Sporobolus ligularis Hack.,; Sporobolus 
ligularis Hack. ex Sodiro. 


SYN:: 


Muhlenbergia ligulata (E. Fourn.) Scribn. & Merr., Bull. Div. 
Agrostol., U.S.D.A. 24: 19. 1901. Basionym: 
Chaboissaea ligulata E. Fourn. 

= Chaboissaea ligulata E. Fourn. 


158 


Catalogue of New World Grasses: II. 


Muhlenbergia lindheimeri Hitche., J. Wash. Acad. Sci. 
24(7): 291. 1934. T-Protol.: USA: Texas, 1847, F 
Lindheimer 725 [HT: US-998949]. T-Specim.: 1 of 1. 
USA: Texas, 1847, FL Lindheimer 725 [HT: US- 
998949; IT: F, GH, MO, UC, US-998947]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Darke, R. (1999); Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Hitchcock, A. S. 
(1951); Peterson, P. M. (2000); Soderstrom, T. R. 
(1967). 

SYN.: Epicampes gracilis Trin. 


Muhlenbergia longifolia Vasey, Contr. U.S. Natl. Herb. 1(8): 
283. 1893. T-Protol.: Mexico: Rio Blanco, 1886, E. 
Palmer 523 (HT: US]. T-Specim.: 1 of 1. Mexico: 
Jalisco: Rio Blanco, 1886, E. Palmer 523 [HT: US- 
995747; IT: MO-2947307]. 

= Muhlenbergia stricta (J. Pres!) Kunth. 


Muhlenbergia longiglumis Vasey, Contr. U.S. Natl. Herb. 
1(8): 283. 1893. T-Specim.: | of 2. Mexico: Jalisco: 
Guadalajara, Jul-Oct 1886, E. Palmer 766 |LT: US- 
2383429; ILT: LE, MO-1837829, US-822922, US- 
995812], LT designated by Hitchcock, Contr. U.S. 
Natl. Herb. 17(3): 301 (1913), but without specifying a 
sheet; also by Soderstrom, Contr. U.S. Natl. Herb. 
34(4): 166 (1967), who did specify a specific sheet. 2 
of 2. Mexico: Jalisco: Guadalajara, 1889,C.G. Pringle 
2365 [ST: MO-1837828, US]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Peterson, P. M. (2000); 
Soderstrom, T. R. (1967). 


Muhlenbergia longiligula Hitchc., Amer. J. Bot. 21(3): 136. 
1934. 
REPLACED NAME: 
Vasey. 
BLOCKING NAME: Muhlenbergia ligulata (E. Fourn.) 
Scribn. & Merr. 

DST.: Mexi, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Hitchcock, A. S. (1951); 
Peterson, P. M. (2000); Soderstrom, T. R. (1967). 

Epicampes anomala Scribn. ex Beal; Epicampes 
distichophylla. var. mutica Scribn.; Epicampes 
ligulata Scribn. ex Vasey; Epicampes stricta var. 
mutica (Scribn.) M.E. Jones; Melica anomala Scribn. 
ex Beal. 


Epicampes ligulata’ Scribn. ex 


SYN:: 


Muhlenbergia longiseta Benth., Pl. Hartw. 28. 1840. T- 
Protol.: Mexico: Ad fossas, Aguas Calientes, Hartweg 
248. T-Specim.: | of |. Mexico: Aguascalientes: ad 
fossas, Aguas Calientes, /artweg 248 [HT: K; IT: K, 
NY. P, US-91927 (fragm. ex W)]. 
= Muhlenbergia tenuifolia (Kunth) Kunth. 


Muhlenbergia lucida Swallen, J. Wash. Acad. Sci. 26(5): 
208. 1936. T-Protol.: Mexico: Chihuahua: on ledges of 
igneous rock, Barranca Island, Sierra Gazachic, 2300- 
2500 m, 35 km SW of Minaca, 16-17 Sep 1934,F. W. 


Pennell 18955 |HT: US-1614380]. T-Specim.: | of 1. 
Mexico: Chihuahua, 16 Sep 1934-17 Sep 1934,F.W. 
Pennell 18955 [HT: US-1614380; IT: US-1614381]. 

DST.: Mexi. 

LVL. ACCPTC.:: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Peterson, P. M. & Y. Herrera A. (1995); Peterson, P. 
M. (2000). 


Muhlenbergia lycuroides Vasey ex Beal, Grass. N. Amer, 2: 
239. 1896. T-Protol.: Mexico, Palmer 489. T-Specim.: 
| of 1. Mexico: Jalisco: Guadalajara, Jul-Oct 1886, E. 
Palmer 489 (HT: MSC; IT: LE, MEXU, MO-2972929, 
NY, P, US-822925, US-81642]. 
NOTE: as Muhlenbergia "lycurioides", as nom. nud. in 
Vasey 1887. 
= Lycurus phalaroides Kunth. 


Muhlenbergia macrotis (Piper) Hitche., N. Amer. Fl. 17(6): 
463. 1935. Basionym: Epicampes macrotis Piper. 
= Muhlenbergia robusta (E. Fourn.) Hitche. 


Muhlenbergia macroura (Kunth) Hitche., N. Amer. FI. 
17(6): 468. 1935. Basionym: Crypsis macroura Kunth. 

DST.: Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); McClure, 
F. A. (1955); MeVaugh, R. (1983); Peterson, P. M. 
(2000); Reeder, C. G. (1994). 

Cinna macroura (Kunth) Kunth; Cinna setifolia (J. 
Presl) Kunth; Crypsinna macroura (Kunth) E. 
Fourn.; Crypsinna_ setifolia (J. Pres!) E. Fourn.; 
Crypsis macroura Kunth; Crypsis setifolia J. Presl; 
Epicampes macroura (Kunth) Benth.; Muehlenbergia 
macroura (Kunth) Hitche.; Phlewm macrourum 
(Kunth) Willd. ex Steud. 


SYN:: 


Muhlenbergia madrensis M.E. Jones, Contr. W. Bot. 14: 12. 
1912. T-Protol.: Mexico: Chihuahua: In meadows, 
Round Valley near Garcia, Sierra Madre Mts., Sept. 
21, 1903, [ALE. Jones] 7315. T-Specim.: | of 1. 
Mexico: Chihuahua, 17 [21] Sep 1903. Coll. Ukn. 
(7315 M.E. JONES) |HT: POM; IT: US-00591130; US 
(fragm. ex POM). 

= Muhlenbergia strictior Scribn. ex Beal. 


Muhlenbergia magna Hitche., N. Amer. Fl. 17(6): 460. 1935. 
T-Protol.: Mexico: under cliffs of barranca near 
Guadalajara, 3 Nov 1890, Pringle 3335 [HT: US- 
825277]. T-Specim.: | of 1. Mexico: Jalisco: under 
cool cliffs of barranca near Guadalajara, 3 Nov 1890, 
C.G. Pringle 3335 [HT: US-825277, IT: MO- 
1837821, US-998977]. 

= Muhlenbergia gigantea (E. Fourn.) Hitche. 


Muhlenbergia majalcensis P.M. Peterson, Syst. Bot. 14(3): 
316-318, f. 1. 1989. T-Protol.: Mexico: Chihuahua: 
Cumbres de Majalca, 20 Sep 1986, Peterson & 
Annable 4519 [HT: WS; IT: ARIZ, ENCB, GH, 
MEXU, MICH, MO, NMC, NY, RSA, TAES, UC, 
UNLV, US,]. T-Specim.: | of 1. Mexico: Chihuahua: 
Sierra Madre Occidental, 21.1 mi W of Hwy. 45, 0.4 
mi E of Cumbres de Majalca, Parque Nacional, elev. 
2200 m, barranca with Pinyon pines andJuniperus, 20 


Catalogue of New World Grasses: II. 


159 


Sep 1986, P.M. Peterson & C.R. Annable 4519 [HT: 
WS; IT: ARIZ, ENCB, GH, HSC, MEXU, MICH, 
MO-3643064, NMC, NY, RSA, TAES, UC, US- 
3127034, WIS]. 
DST.: Mexi. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Peterson, P. M. 
& C. R. Annable (1991); Peterson, P. M. (1989); 
Peterson, P. M. (2000). 


Muhlenbergia marshii \.M. Johnst., J. Arnold Arbor. 24(4): 
392. 1943. T-Protol.: Mexico: Coahuila: Sierra del 
Carmen, 8 Sep 1936, E. G. Marsh Jr. 746 [HT: GH]. 
T-Specim.: | of 1. Mexico: Coahuila, 8 Sep 1936,£.G. 
Marsh 746 [HT: GH; IT: LL, US-1647094]. 

= Muhlenbergia rigens (Benth.) Hitche. 


Muhlenbergia matudae Sohns, J. Wash. Acad. Sci. 46(12): 
382, f. 32-38. 1956. T-Protol.: Mexico: collected en 
madera humeda, orilla de bosque mixto de pinos y 
oyamel, Lago de Zempoala, Edo. de Morelos, alt. 3000 
m, 7 Oct 1951, £. Matuda 25601 [HT: US-2079186]. 
T-Specim.: | of 1. Mexico: Morelos, 7 Oct 1951, E. 
Matuda 25601 [HT: US-2079 186]. 

= Muhlenbergia orophila Swallen. 


Muhlenbergia maxima Legaard & Sanchez Vega, Nordic J. 
Bot. 10: 439. 1990, T-Protol.: Peru: Prov. Cajamarca: 
Choten between Paso El Gavilan and San Juan at km 
153 on the road to the coast, 2900 m a.s.l., 29 May 
1984, Sanchez Vega & Ruiz Vigo 3561 [HT: CPUN; 
IT: AAU, K, US, USM]. 

DST.: Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Laegaard, S. & I. Sanchez V. (1990); 
Peterson, P. M. (2000); Tovar, O. (1993). 


Muhlenbergia metcalfei M.E. Jones, Contr. W. Bot. 14: 12. 
1912. T-Protol.: USA: New Mexico: Grant Co.: Santa 
Rita Mt.: on rocky slopes, Metcalf 1485. 
NOTE: as "Muhlenbergia metcalfi', named for O.B. 
Metcalfe. 
= Muhlenbergia rigida (Kunth) Kunth. 


Muhlenbergia mexicana (L.) Trin., Gram. Unifl. Sesquifl. 
189, 190, 297, t. 5, f. 8. 1824. Basionym: Agrostis 
mexicana L. 

NOTE: as "Miihlenbergia". 

DST.: Cana, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Bor, N. L. (1960); Cody, W. J. 

(1996); Cronquist, A., A. H. Holmgren, N. H. 

Holmgren & J. L. Reveal (1977); Fernald, M. (1950); 

Gleason, H. A. & A. Cronquist (1968); Gleason, H. 

A. & A. Cronquist (1991); Great Plains Flora 

Association (1986); Hickman, J. C. (1993); 

Hitchcock, A. S. (1951); Hitchcock, C. L., A. 

Cronquist, M. Ownbey & J. W. Thompson (1969); 

Munz, P. A. & D. D. Keck (1959); Peterson, P. M. 

(2000); Radford, A. E., H. E. Ahles & C. R. Bell 

(1968); Scoggan, H. J. (1978 [1979]); Small, J. K. 

(1933); Welsh, S. L. (1974). 

Agrostis mexicana \..; Agrostis tenacissima L. f.; 

Cinna mexicana (L.) P.  Beauv.; Lepyroxis 

canadensis P. Beauv. ex B.D. Jacks.; Muhlenbergia 

mexicana fo. mexicana, Muhlenbergia mexicana var. 
purpurea Alph. Wood; Muhlenbergia polystachya 


SYN.: 


Mack. & Bush; Podosaemum mexicanum (L.) Link; 
Podosemum mexicanum (L.) Link; Polypogon 
canadensis E. Fourn.; Trichochloa mexicana (L.) 
Trin.; Vilfa mexicana (L.) P. Beauv. 


Muhlenbergia mexicana fo. ambigua (Torr.) Fernald, 
Rhodora = 45(534): 236. 1943. Basionym: 
Muhlenbergia ambigua Torr. 

= Muhlenbergia mexicana var. filiformis (Torr.) Scribn. 


Muhlenbergia mexicana fo. commutata (Scribn.) Wiegand, 
Rhodora 26(301): 1. 1924. Basionym: Muhlenbergia 
mexicana subsp. commutata Scribn. 

= Muhlenbergia frondosa fo. commutata (Scribn.) 
Fernald. 


Muhlenbergia mexicana subsp. commutata Scribn., Rhodora 
9(98): 18-19. 1907. T-Specim.: | of 1. USA: Maine: 
Aroostook Co., Fort Fairfield, 9 Sep 1896, ML. 
Fernald 522 [LT: US-994763], LT designated by 
Hitchcock, Manual U.S. Grasses, p. 890 (1935). 

= Muhlenbergia frondosa fo. commutata (Scribn.) 
Fernald. 


Muhlenbergia mexicana var. commutata (Scribn.) Farw., 
Rep. Michigan Acad. Sci. 17: 181. 1916. Basionym: 
Muhlenbergia mexicana subsp. commutata Scribn. 

= Muhlenbergia frondosa fo. commutata (Scribn.) 
Fernald. 


Muhlenbergia mexicana var. filiformis Vasey, Grass. U.S. 23. 
1883. 
NOTE: nom. nud., no basionym cited. 
= Muhlenbergia mexicana var. filiformis (Torr.) Scribn. 


Muhlenbergia mexicana var. filiformis (Torr.) Scribn., 
Mem. Torrey Bot. Club 5: 36. 1894. Basionym: 
Agrostis lateriflora var. filiformis Torr. 

DST.: Cana, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Peterson, P. M. (2000). 

SYN.:  Muhlenbergia ambigua Yorr., Muhlenbergia 
ambigua var. ambigua,; Muhlenbergia ambigua var. 
filiformis (Yorr.) Farw.; Muhlenbergia foliosa (hort. 
ex Roem. & Schult.) Trin.; Muhlenbergia foliosa fo. 
ambigua (Torr.) Wiegand; Muhlenbergia foliosa 
subsp. ambigua (Torr.) Scribn.; Muhlenbergia foliosa 
subsp. setiglumis (S. Watson) Scribn.; Muhlenbergia 
mexicana fo. ambigua (Yorr.) Fernald; Muhlenbergia 
mexicana fo. setiglumis (S. Watson) Fernald; 
Muhlenbergia mexicana var. filiformis Vasey; 
Muhlenbergia setiglumis (S. Watson) A. Nelson & 
J.F. Macbr.; Muhlenbergia sylvatica var. setiglhumis 
S. Watson; Podosemum foliosum (hort. ex Roem. & 
Schult.) Link. 


Muhlenbergia mexicana (L.) Trin. fo. mexicana. 
= Muhlenbergia mexicana (L.) Trin. 


Muhlenbergia mexicana (L.) Trin. var. mexicana. 
DST.: Cana, USA. 
LVL. ACCPTC:: 1. 


Muhlenbergia mexicana var. purpurea Alph. Wood, Amer. 
Bot. Fl. pt. 2: 386. 1871. T-Specim.: | of 1. USA: 
Illinois, 1880, J. Wolfs.n. (IT: US-952457]. 

= Muhlenbergia mexicana (L.) Trin. 


160 


Catalogue of New World Grasses: II. 


Muhlenbergia mexicana fo. setiglumis (S. Watson) Fernald, 
Rhodora = 45(534): 236. 1943. Basionym: 
Muhlenbergia sylvatica var. setiglumis S. Watson. 

= Muhlenbergia mexicana var. filiformis (Torr.) Scribn. 


Muhlenbergia meziana Hitche., N. Amer. Fl. 17(6): 461. 
1935. 
NOTE: nom. nov., as Muhlenbergia "Meziana". 
REPLACED NAME: Epicampes minutiflora Mez. 
BLOCKING NAME: Muhlenbergia minutiflora (Michx.) 
Hitche. 
= Muhlenbergia robusta (E. Fourn.) Hitche. 


Muhlenbergia michisensis Y. Herrera & P.M. Peterson, 
Novon 2(2): 117, f. 2. 1992. T-Protol.: Mexico: 
Durango: Mun. de Suchil: Las Escobas, San Juan de 
Michis, 23.24°N 104.08°W, 17 Sep 1989, Herrera A. 
& Acevedo 986 [HT: CIIDIR; IT; MTMG, US]. T- 
Specim.: | of 1. Mexico: Durango: Suchil Mun.: Las 
Escobas, San Juan de Michis, Bosque de Quercus- 
Pinus, 2420 m, 18 Sep 1989, ¥.Herrera & S.Acevedo 
986 [HT: CUDIR; IT; MTMG, US-3239026], dates 
verified in pub. and on US specimen, 17 & 18, 
respectively. 

DST.: Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Herrera A., Y. 
& P.M. Peterson (1992); Herrera-Arrieta, Y. (1998): 
Peterson, P. M. (2000). 


Gram. Unifl. 
Trichochloa 


Muhlenbergia microsperma (DC.)_ Trin., 
Sesquifl. 193. 1824. — Basionym: 
microsperma DC. 

NOTE: nom. inval., Trinius did not explicitly make the 
combination in either the text or the figure list. 
= Muhlenbergia microsperma (DC.) Kunth. 


Muhlenbergia microsperma (DC.) Kunth, Revis. Gramin. 
1: 64. 1829. Basionym: Trichochloa microsperma DC. 

DST.: Boli, Colo, Ecua, Guat, Mexi, Peru, USA, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Abrams, L. (1923); Beetle, A. A. 
(1977); Brako, L. & J. L. Zarucchi (1993); Davidse, 
G., M. Sousa S. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. & R. Moran 
(1981); Hickman, J. C. (1993); Hitchcock, A. S. 
(1927); Hitchcock, A. S. (1951); Jorgensen, P. M. & 
C. Ulloa U. (1994); Jorgensen, P. M. & S. Leén- 
Yanez (eds.) (1999); Lawesson, J. E., H. Adsersen & 
P. Bentley (1987); McClure, F. A. (1955); McVaugh, 
R. (1983); Munz, P. A. & D. D. Keck (1959); Munz, 
P. A. (1974); Peterson, P. M. & C. R. Annable 
(1991); Peterson, P. M. (2000); Porter, D. M. (1983); 
Reeder, C. G. (1994); Reeder, J. R. & C. G. Reeder 
(1971); Renvoize, S. A. (1998); Shreve, F. & I. L. 
Wiggins (1964); Tovar, O. (1993); Wiggins, I. L. & 
D. M. Porter (1971). 

SYN.: Agrostis debilis (Kunth) Spreng.; Agrostis 
microsperma Lag.;, Agrostis setosa (Kunth) Spreng.; 
Muhlenbergia debilis (Kunth) Kunth; Muhlenbergia 
debilis (Kunth) Trin.; Muhlenbergia fasciculata 
Trin.; Muhlenbergia microsperma (DC.)_ Trin.; 
Muhlenbergia purpurea Nutt.; Muhlenbergia 
ramosissina Vasey, Muhlenbergia setosa (Kunth) 
Kunth; Muhlenbergia  setosa (Kunth) © Trin.; 


Podosemum debile Kunth; Podosemum setosum 
Kunth; 7richochloa debilis (Kunth) Roem. & Schult.; 
Trichochloa microsperma DC.; Trichochloa setosa 
(Kunth) Roem. & Schult. 


Muhlenbergia minuscula H. Scholz, Willdenowia 14: 393. 
1984.  T-Protol.: Bolivia: Canton — Ulla-Ulla: 
Apolobamba Cordillera, Pampa von Ulla-Ulla, 4450 
m, Menhofer 1974 (HT: B]. 

= Muhlenbergia ligularis (Hack.) Hitchc. 


Muhlenbergia minutiflora (Michx.) Hitche., Trans. Kansas 
Acad. Sci. 14: 140. 1896. Basionym: Dilepyrum 
minutiflorum Michx. 

= Muhlenbergia schreberi J.F. Gmel. 


Muhlenbergia minutissima (Steud.) Swallen, Contr. U.S. 
Natl. Herb. 29(4) 207. 1947. Basionym: Agrostis 
minutissima Steud. 

DST.: Mexi, USA, 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977): Correll, D. S. 
& M. C. Johnston (1970); Cronquist, A., A. H. 
Holmgren, N. H. Holmgren & J. L. Reveal (1977); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Gould, F. W. & R. Moran (1981): 
Hickman, J. C. (1993); Hitchcock, A. S. (1951); 
Hitchcock, C. L., A. Cronquist, M. Ownbey & J. W. 
Thompson (1969); McVaugh, R. (1983); Munz, P. A. 
& D. D. Keck (1959); Munz, P. A. (1974); Peterson, 
P.M. & C. R. Annable (1991); Peterson, P. M. 
(2000). 

SYN.: Agrostis minutissima Steud.; Milium microspermum 
Lag.; Muhlenbergia confusa (E. Fourn.) Swallen; 
Panicum — microspermum  (Lag.)  E. — Fourn.; 
Sporobolus confusus (E. Fourn.) Vasey; Sporobolus 
microspermus (Lag.) Hitche.; Sporobolus 
minutissimus (Steud.) Hitche.; Vilfa confusa_ E. 
Fourn. 


Muhlenbergia montana (Nutt.) Hitche., U.S.D.A. Bull. 
(1915-23) 772: 145, 147. 1920. Basionym:Calycodon 
montanum Nutt. 

DST.: Guat, Mexi, USA. 

LVL. ACCPTC.:: I. 

ALSO ACCPTD. BY: Abrams, L. (1923); Beetle, A. A. 
(1977); Breedlove, D. E. (1986); Correll, D. S. & M. 
C. Johnston (1970); Cronquist, A., A. H. Holmgren, 
N. H. Holmgren & J. L. Reveal (1977); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Herrera-Arrieta, Y. (1998); Hickman, J. C. (1993); 
Hitchcock, A. S. (1951); Hitchcock, C. L., A. 
Cronquist, M. Ownbey & J. W. Thompson (1969); 
McClure, F. A. (1955); McVaugh, R. (1983); Munz, 
P. A. & D. D. Keck (1959); Peterson, P. M. (2000). 

SYN.: Calycodon montanum Nutt.; Muhlenbergia enervis 
(Scribn. ex Beal) Hitche.; Muhlenbergia gracilis vat. 
breviaristata| Vasey; Muhlenbergia gracilis var. 
enervis Scribn. ex Beal; Muhlenbergia gracilis var. 
major Nasey; Muhlenbergia  subalpina Vasey; 
Muhlenbergia trifida Hack. 


Muhlenbergia monticola Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 91. 1862. T-Protol.: USA: Texas: 
northwestern Texas. T-Specim.: | of 1. USA: Texas: 
side of hills in the Pass of the Limpia, 24 May 1849, 
C. Wright 731 (LT: PH; ILT: MO, US-81641], LT 
designated by Dorr & Peterson, Sida 15: 591 (1993). 


Catalogue of New World Grasses: II. 


161 


= Muhlenbergia tenuifolia (Kunth) Kunth. 


Muhlenbergia mucronata (Kunth) Trin, Gram. Unifl. 
Sesquifl. 194, t. 5, f. 23. 1824. Basionym: Podosemum 
mucronatum Kunth. 

NOTE: basionym p. 194, comb. in tab. 

LVL. ACCPTC.: |. 

ALSO ACCPTD. BY: Peterson, P. M. (2000). 

SYN.: Agrostis mucronata (Kunth) Spreng.; Muhlenbergia 
laxiflora Scribn.,; Podosemum mucronatum Kunth; 
Trichochloa mucronata (Kunth) Roem. & Schult. 


Muhlenbergia multinodis Aspl., Bot. Not. 1939: 796, f. 1. 
1939. T-Protol.: Colombia: Popayan: Hab. in Novae 
Granatae ad pagum El Tambo loco graminoso, 1700 
m, 22 Jun 1938, Sneidern 1323. 

= Muhlenbergia lehmanniana Henrard. 


Muhlenbergia mundula 1.M. Johnst., J. Arnold Arbor. 24(4): 
392-393. 1943. T-Protol.: Mexico: Chihuahua: by 
streams near Chihuahua, 13 Oct 1885, Pringle 417 
[HT: GH]. T-Specim.: 1 of 1. Mexico: Chihuahua: by 
streams near Chihuahua, 13 Oct 1885, C.G. Pringle 
417 (HT: GH; IT: MO-1837822, US-999023]. 

= Muhlenbergia rigens (Benth.) Hitche. 


Muhlenbergia mutica (Rupr. ex E. Fourn.) Hitche., N. 
Amer. FI. 17: 459. 1935. Basionym: Epicampes mutica 
Rupr. ex E. Fourn. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Peterson, 
P. M. (2000); Reeder, C. G. (1994); Soderstrom, T. 
R. (1967). 

Epicampes mutica Rupr.; Epicampes mutica Rupr. ex 
E. Fourn. 


SYN: 


Muhlenbergia nana Benth., Pl. Hartw. 262. 1846. T-Protol.: 
Ecuador; In radicibus montis ignivomi Cotopaxi, 
Hartweg s.n. T-Specim. 1 of 1. Ecuador: Mt. 
Cotopaxi, 1843, Hartweg 1458 (HT: K; IT: kK, LE, P, 
US-91916 (fragm. ex P, LE)]. 

= Muhlenbergia peruviana (P. Beauv.) Steud. 


Muhlenbergia nardifolia Griseb., Abh. Konigl. Ges, Wiss. 
Gottingen 24: 294. 1879. T-Protol.: Argentina: 
Cordoba and Santiago del Estero: Cordoba, Pampa pr. 
Laguna de Pocho. 

= Muhlenbergia torreyi (Kunth) Hitchce. ex Bush. 


Muhlenbergia nebulosa Scribn. ex Beal, Grass, N. Amer. 2: 
247. 1896. T-Protol.: Mexico: wet places, hills near 
Guadalajara, 1889, Pringle 2366. T-Specim.: | of 1. 
Mexico: Jalisco: wet places, hills near Guadalajara, 5 
Nov 1889, C.G. Pringle 2366 [HT: MSC; IT: GH, K, 
LE, LL, MICH, MO-2974337, NY, RSA, SI, TEX, 
US-995539, US-746248, US-746269, US-822939, US- 
1645933, VT]. 

= Muhlenbergia tenuissima (J. Presl) Kunth. 


Muhlenbergia neomexicana Wasey, Bot. Gaz. 11(12): 337. 
1886. T-Protol.: USA: New Mexico: "Rocky hills and 
mountain sides", /G.R. Vasey] s.n. T-Specim.: | of 1. 
USA: New Mexico, 1881, G.R. Vasey s.n. (HT: US- 
81640]. 

NOTE: as "Neo-Mexicana". 


= Muhlenbergia pauciflora Buckley. 


Muhlenbergia nigra Hitchc., N. Amer. Fl. 17(6): 468. 1935. 
T-Protol.; Mexico: Mexico: Nevado de Toluca, 
collected on cool slopes under pines, 3300 m, 2 Sep 
1892, Pringle 4211 [HT: US-746689]. T-Specim.: | of 
1. Mexico: Mexico: Nevada de Toluca, cool slopes 
under pines, 11000 ft, 2 Sep 1892,C.G. Pringle 4211] 
[HT: US-746689; IT: F, MO-2974170, MSC, US- 
821929]. 

DST.: CoRi, Guat, Mexi. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994): Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); McClure, F. A. (1955): 
Peterson, P. M. (2000); Pohl, R. W. (1980); Reeder, 
C. G. (1994). 


Muhlenbergia orophila Swallen, Contr. U.S. Natl. Herb. 
29(9): 408. 1950. T-Protol.: Guatemala: Dept. 
Huehuetenango: Cerro Chemal: in alpine meadow, 
summit of Sierra de los Cuchumantanes, alt. 3700- 
3800 m; 8 Aug 1942.,/.4. Steyermark 50309 (HT: F- 
1202399]. T-Specim.: 1 of 1. Guatemala: 
Huehuetenango, 8 Aug 1942, /.4. Steyermark 50309 
[HT: F-1202399; IT: US-132785, US-132784, US- 
1935066, US-2208672]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Espejo Serna, A., 
A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
McClure, F. A. (1955); Peterson, P. M. (2000); 
Reeder, C. G. (1994). 

SYN.:; Muhlenbergia matudae Sohns. 


Muhlenbergia palmeri Vasey, Bull. Torrey Bot. Club 
13(12): 231. 1886. T-Protol.: Mexico: in S.W. 
Chihuahua: 1885, Dr. E. Palmer s.n. T-Specim.: | of 
1. Mexico: Chihuahua, Aug-Nov 1885,£. Palmer 16 
[HT: US-995818; IT: US-995817]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: |; cultivated. 

ALSO ACCPTD. BY: Beetle, A. A. (1977), Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Peterson, P. M. (2000). 

SYN.: Muhlenbergia dubioides C.O. Goodd. 


Muhlenbergia palmirensis Grignon & Lagaard, Nordic J. 
Bot. 9(1): 47, f. 1-2. 1989. T-Protol.: Ecuador: 
Chimborazo: Panamerican highway between 
Riobamba and Alausi, N of Palmira, 3200 m, Legaard 
52721 [HT: AAU; IT: K, MO, NY, QCA, QCNE, US]. 
T-Specim.: | of 2. 2 of 2. Ecuador: Chimborazo: 3200 
m, Legaard 52721 (HT: AAU; IT: K, MO, NY, QCA, 
QCNE, US]. 

DST.: Ecua, 

LVL. ACCPTC:.: I. 

ALSO ACCPTD. BY: Grignon, I. & S. Legaard (1989); 
Jorgensen, P. M. & C. Ulloa U. (1994); Jorgensen, P. 
M. & S. Leén-Yanez (eds.) (1999); Peterson, P. M. 
(2000); Valencia, R., N. Pitman, S. Le6n-Yanez & P. 
M. Jorgensen (2000). 


Muhlenbergia palustris Scribn., Bull. Div. Agrostol., 
US.D.A. 11: 47. 1898. T-Protol.: USA: D.C.: 
Brightwood, swampy grounds, Sep 1896, E.S. Steele 


162 


Catalogue of New World Grasses: II. 


s.n. T-Specim.: | of 1. USA: District of Columbia: 
Brightwood, Sep 1896, E.S. Steele s.n. [HT: US- 
995038; IT: US-995040 (possible)]. 

= Muhlenbergia schreberi J.F. Gmel. 


Muhlenbergia parishii Vasey, Bull. Torrey Bot. Club 13: 53. 
1886. 
REPLACED = NAME: 
californica Vasey. 
BLOCKING NAME: Muhlenbergia californica Vasey. 
= Muhlenbergia californica Vasey. 


Muhlenbergia sylvatica — var. 


Muhlenbergia parviglumis Vasey, Contr. U.S. Natl. Herb. 
3(1): 71. 1892. T-Protol.: USA: Texas, G.C. Nealley 
sn. T-Specim.: 1 of 1. USA: Texas, 1887, G.C. 
Nealley s.n. [HT: US-81638; IT: US-994967 (Nealley 
2, possible IT)]. 

= Muhlenbergia spiciformis Trin. 


Muhlenbergia pauciflora Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 91. 1862. T-Protol.: Hillsides, western 
Texas. T-Specim.: | of 1. USA: Texas, no date, C. 
Wright 732 |LT: PH; ILT: US-81637], LT designated 
by Hitchcock, Man. Grass. U.S. 891 (1935), but 
without citing a specific sheet in a specific herbarium. 

DST.: Mexi, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. & 
R. Moran (1981); Hickman, J. C. (1993); Hitchcock, 
A. S. (1951); Peterson, P. M. (2000). 

Muhlenbergia neomexicana Vasey; Muhlenbergia 
pringlei (Scribn.) Scribn.; Muhlenbergia sylvatica 
var. pringlei Scribn. 


SYN.: 


Muhlenbergia pectinata C.O. Goodd., J. Wash. Acad. Sci. 
31: 505, 1941. T-Protol.: Mexico: Jalisco: moist ledges 
ot the barranca near Guadalajara, | Nov. 1888, Pringle 


1745 (HT: US-995478; IT: GH, NY]. T-Specim.: | of 


1. Mexico: Jalisco: near Guadalajara, 1 Nov 1888, 
C.G. Pringle 1745 [HT: US-995478; IT: GH, NY]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Hitchcock, A. S. (1951); 
Peterson, P.M. & C. R. Annable (1991); Peterson, P. 
M. (2000). 


Muhlenbergia peruviana (P. Beauv.) Steud., Nomencl. Bot. 
(ed. 2) 1: 41. 1840. Basionym: Clomena peruviana P. 
Beauv. 

DST.: Arge, Boli, Chil, Ecua, Guat, Mexi, Peru, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Brako, L. & J. L. Zarucchi (1993); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Foster, R. C. (1958); Hitchcock, A. S. 
(1927); Jorgensen, P, M. & C. Ulloa U. (1994): 
Jorgensen, P. M. & S. Leén-Yanez (eds.) (1999); 
Macbride, J. F. (1936); Marticorena, C. & M. 
Quezada (1985); Matthei, O. (1984); Peterson, P. M. 
& C. R. Annable (1991); Peterson, P. M. (2000); 
Renvoize, S. A. (1998); Saravia, E. F. (1996); Tovar, 
O. (1993); Zuloaga, F. O., E. G. Nicora, Z. E. Rigolo 


de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 

Agrostis peruviana (P. Beauv.) Spreng., Clomena 
peruviana P. Beauv.; Clomena_ peruviana var. 
peruviana, Clomena peruviana var. pulvinata Nees; 
Epicampes bourgaei E. Fourn. ex Hemsl.; Epicampes 
bourgeaei (E. Fourn.) M.E. Jones; Epicampes 
bourgeaei E, Fourn.; Epicampes bourgeaei var. 
bourgeaei; Epicampes clomena ME. — Jones: 
Muhlenbergia bourgeaei E, Fourn.; Muhlenbergia 
bourgeaei_ E. Fourn. ex Hemsl.; Muhlenbergia 
clomena Kunth; Muhlenbergia herzogiana Henrard: 
Muhlenbergia nana Benth.; Muhlenbergia peruviana 
fo. peruviana; Muhlenbergia peruviana fo.versicolor 
Kuntze, Muhlenbergia peruviana fo. viridis Kuntze; 
Muhlenbergia peruviana var. breviaristata Hack, ex 
Buchtien; Muhlenbergia peruviana var. elatior 
Kuntze; Muhlenbergia peruviana var. peruviana; 
Muhlenbergia peruviana var. pulvinata (Nees) Nees 
& Mey. ex Kuntze; Muhlenbergia peruviana var. 
subcaespitosa Kuntze; Muhlenbergia pulcherrima 
Scribn. ex Beal; Muhlenbergia pusilla Steud.; 
Sporobolus complanatus Scribn. ex Beal. 


SYN.: 


Muhlenbergia  peruviana var. breviaristata| Hack. ex 
Buchtien, Contr. Fl. Bolivia 1: 72. 1920. T-Protol.: 
Bolivia: La Paz, Buchtien [865] [US]. 

NOTE: nom. nud. 

= Muhlenbergia peruviana (P. Beauv.) Steud. 


Muhlenbergia peruviana var.elatior Kuntze, Revis. Gen. PI. 
3(2): 357. 1898. T-Protol.: [Bolivia: Tunari]. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Muhlenbergia peruviana Steud. fo. peruviana. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Muhlenbergia peruviana Steud. var. peruviana. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Muhlenbergia peruviana var. pulvinata (Nees) Nees & Mey. 
ex Kuntze, Revis. Gen. Pl. 3(3): 357. 1898. Basionym: 
Clomena peruviana var. pulvinata Nees. 

NOTE: Presumably based on Clomena peruviana var. 
pulvinata. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Muhlenbergia peruviana var. subcaespitosa Kuntze, Revis. 
Gen. Pl. 3(3): 357. 1898. T-Protol.: No loc. cited. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Muhlenbergia peruviana fo. versicolor Kuntze, Revis. Gen. 
PI. 3(3): 357. 1898. T-Specim.: 1 of 2. Bolivia: 
Tunarigebirge, 3000 m, May 1892, Kuntze s.n. [LT: 
NY; IT: US (fragm. ex NY)], LT designated by 
Peterson & Annable, Syst. Bot. Monogr. 31: 73. 1991. 
2 of 2. Argentina: Salta [ST]. 

= Muhlenbergia peruviana (P. Beauv.) Steud. 


Muhlenbergia peruviana fo. viridis Kuntze, Revis. Gen. PI. 
3(3): 357. 1898. T-Protol.: Bolivia: 4000 m,/Kuntze/] 
SM. 
= Muhlenbergia peruviana (P, Beauv.) Steud. 


Muhlenbergia phragmitoides Griseb., Abh. Konigl. Ges. 
Wiss. Gottingen 19: 255. 1874. T-Protol.: Argentina: 
Tucuman, Cuesta de Anfama: in sylvis montanis reg. 
Aliso, [Lorentz s.n.j. T-Specim.: | of 1. Argentina, 


Catalogue of New World Grasses: II. 


163 


Lorentz 79 [HT: GOET; IT: US-91911 (fragm. ex 
GOET)]. 
= Muhlenbergia rigida (Kunth) Kunth. 


Muhlenbergia phragmitoides var. breviaristata  Hack., 
Annuaire Conserv. Jard. Bot. Geneve 17: 291, 1914. 
T-Protol.: Argentina: Tucuman: Lara, 3200 m, 28 Mar 
1912, Lilla 11322 [Stuckert hb. 22470]. T-Specim.: | 
of 1. Argentina: Tucuman, Lara, a 3200 m. alt., Lillo 
1/322 (HT: W; IT: US-3412353 (fragm. ex W)]. 

= Muhlenbergia breviaristata (Hack.) Parodi. 


Muhlenbergia phragmitoides Griseb. var. phragmitoides. 
= Muhlenbergia rigida (Kunth) Kunth. 


Muhlenbergia pilosa P.M. Peterson, Wipff & S.D. Jones, 
Madrofio 39(2): 151. 1992. T-Protol.: Mexico: 
Mexico: Mun. Tejupilco: 17 km NE of Tejupilco on 
road to Temascaltepec, 18.59°N 10.04°W, 30 Oct 
1982, SD. Koch & P.A. Fryxell 82256 [HT:CHAPA; 
IT:CHAPA, IEB. MEXU, MICH, TAES, US, XAL]. 
T-Specim.: | of 1. Mexico: Mexico: Tejupilco Mun.: 
17 km NE de Tupilco, por la carr. a Temascaltepec, 
1,530 mt, 30 Oct 1992, SD. Koch & P.A, Fryxell 
82256 [HT: HT:CHAPA,; IT:CHAPA, IEB, MEXU, 
MICH, TAES, US-323217, XAL]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Peterson, P. M. 
(2000); Peterson, P. M., J. K. Wipff & S. D. Jones 
(1992). 


Muhlenbergia plumbea (Trin.) Hitchce., Contr. U.S. Natl. 
Herb. 17(3): 296. 1913. Basionym: Vilfa plumbea 
Trin. 

DST.: Guat, Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Morden, C. W. & S. L. Hatch 
(1996); Peterson, P. M. (2000); Reeder, C. G. (1994). 

SYN.: Sporobolus plumbeus (Trin.) Hemsl.; Sporobolus 
poaeoides Hack.; Vilfa plumbea Trin. 


Muhlenbergia polycaulis Scribn., Bull. Torrey Bot. Club 38: 
327. 1911. T-Protol.: Mexico: Chihuahua: 1887, 
Pringle 1414 [HT: US]. T-Specim.: | of 1. Mexico: 
Chihuahua: Sierra Madre, 30 Sep 1887,C.G. Pringle 
1414 [HT: US-81636; IT: US-822943, US-155173, 
US-995733]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977), Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Hitchcock, A. S. (1951); McVaugh, R. 
(1983); Peterson, P. M. (2000). 


Muhlenbergia polypogon (DC.) Kunth, Revis. Gramin. 1: 64. 
1829. Basionym: Trichochloa polypogon DC. 
= Muhlenbergia capillaris (Lam.) Trin. 


Muhlenbergia polypogonoides Hack., Ann. K.K. Naturhist. 
Hofmus. 17; 255. 1902. T-Protol.: Mexico, Schmitz 
862 [HT: W]. T-Specim.: 1 of 1. Mexico, Schmitz 862 
{HT: W; IT: US-3412356 (fragm. ex W)]. 


NOTE: as "Muehlenbergia" polygonoides. 
= Muhlenbergia setarioides E. Fourn. 


Muhlenbergia polystachya Mack. & Bush, Trans. Acad. Sci. 
St. Louis 12(7): 79-80, pl. 12. 1902. T-Protol.: USA: 
Open rocky woods E of Sibley, A. K. Mackenzie 637 
{HT: Herb. K. K. Mackenzie; IT: MO]. 

= Muhlenbergia mexicana (L.) Trin. 


Muhlenbergia porteri Scribn. ex Beal, Grass. N. Amer. 2: 
259. 1896. 

REPLACED NAME: Muhlenbergia texana Thurb. ex 
Porter & J.M. Coult. 

BLOCKING NAME: Muhlenbergia texana Buckley. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Abrams, L. (1923); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. & 
R. Moran (1981); Hickman, J. C. (1993); Hitchcock, 
A. S. (1951); Munz, P. A. & D. D. Keck (1959); 
Munz, P. A. (1974); Peterson, P. M. (2000); Shreve, 
F. & I. L. Wiggins (1964). 

SYN.: Muhlenbergia texana Thurb. ex Porter & J.M. Coult.; 
Podosemum porteri (Scribn. ex Beal) Bush. 


Muhlenbergia presliana Hitche., N. Amer. Fl. 17(6): 462. 
1935. 
NOTE: nom. nov., as Muhlenbrgis "Presliana". 
REPLACED NAME: Epicampes stricta J. Presl. 
BLOCKING NAME: Muhlenbergia stricta (J. Presl) Kunth. 
= Muhlenbergia robusta (E. Fourn.) Hitche. 


Muhlenbergia pringlei (Scribn.) Scribn., Contr. U.S. Natl. 
Herb. 3(1): 71. 1892. Basionym: Muhlenbergia 
sylvatica var. pringlei Scribn. 

= Muhlenbergia pauciflora Buckley. 


Muhlenbergia pubescens (Kunth) Hitche.. N. Amer. FI. 
17(6):; 460. 1935. Basionym: Agrostis pubescens 
Kunth. 

DST.: Mexi. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Peterson, P. M. (2000); 
Soderstrom, T. R. (1967). 

Agrostis lanata Kunth; Agrostis pubescens Kunth; 
Cinna lanata (Kunth) Kunth; Cinna pubescens 
(Kunth) Kunth; Epicampes lanata (Kunth) J. Presl ex 
Kunth; Epicampes pubescens (Kunth) J. Presl ex 
Kunth; Epicampes pubescens (Kunth) M.E. Jones; 
Muehlenbergia lanata (Kunth) Hitche.; 
Muehlenbergia pubescens (Kunth) — Hitche.; 
Muhlenbergia lanata (Kunth) Hitchce. 


SYN.: 


Muhlenbergia pubigluma Swallen, Proc. Biol. Soc. Wash. 
56: 78. 1943. T-Protol.: Mexico: Coahuila: Mun. de 
Cuatro Cienegas: Sierra de la Madera, Canon del 
Agua, common in shrub zones of lower canyon, 10 
Sep 1939, C. H. Muller 3264 [NA]. T-Specim.: | of 1. 
Mexico: Coahuila, 10 Sep 1939, C.H. Muller 3264 
[HT: NA-145092; IT:US-2209360, US-1585527]. 

DST.: Mexi. 
LVL. ACCPTC.: I. 


164 


Catalogue of New World Grasses: II. 


ALSO ACCPTD. BY: Beetle, A. A. (1977); Peterson, P. M. 
(2000); Soderstrom, T. R. (1967). 


Muhlenbergia pulcherrima Scribn. ex Beal, Grass. N. Amer. 
2: 240. 1896. T-Protol.: Mexico, Pringle 1416. T- 


Specim.; | of 1. Mexico: Chihuahua: dry ledges of 


porphyry, Sierra Madre, 30 Sep 1887, C.G. Pringle 
1416 (HT: MSC; IT: MO-3727978, NY, US-995494 
VT]. 

= Muhlenbergia peruviana (P. Beauv.) Steud. 


> 


Muhlenbergia pungens Thurb. ex A. Gray, Proc. Acad. Nat. 
Sci. Philadelphia 1863: 78. 1864. T-Protol.: USA: 
Colorado: on and near the Rocky Mts. lat. 39°-41°, 
1862, Hall & Harbour 632. T-Specim.: | of 1. USA: 
American Plains Flora, Lat. 41°, 1862, £. Hall & JP. 
Harbour 632 |HT: GH; IT: US-3376135 (fragm. ex 
GH)]. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Cronquist, A., A. H. 
Holmgren, N. H. Holmgren & J. L. Reveal (1977); 
Great Plains Flora Association (1986); Hitchcock, A. 
S. (1951); Peterson, P. M. (2000). 

SYN.: Podosemum pungens (Thurb. ex A. Gray) Bush. 


Muhlenbergia purpurea Nutt., J. Acad. Nat. Sci. 
Philadelphia, ser. 2, 1: 186. 1848. T-Protol.: USA: 
California: Santa Barbara, Upper California: and Santa 
Catalina Island, Gambel s.n. T-Specim.: | of 1. USA: 
California: Santa Barbara Co.: Santa Barbara and 
Santa Catalina Island, Gambel s.n. [HT: K]. 

= Muhlenbergia microsperma (DC.) Kunth. 


Muhlenbergia purpusii Mez, Repert. Spec. Nov. Regni 
Veg. 17(13-18): 214. 1921. T-Protol.: Mexico: San 
Luis Potosi: Minas de San Rafael, Purpus s.n. T- 
Specim.: | of 1. Mexico: San Luis Potosi: Minas de 
San Rafael, 1911, CLA. Purpus 5011] [HT: B; IT: MO- 
2974180, US-463679, US-72636 (fragm. ex B)]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 

Peterson, P. M. (2000). 


Muhlenbergia pusilla Steud., Syn. Pl. Glumac. 1: 177. 1854. 
T-Protol.: Mexico: Tolucca, Hrbr. Berlandier 1141. T- 
Specim.: | of 1. Mexico: México: Valley of Toluca, 
Oct. 1827, JL. Berlandier 1141] (HT: P; IT: K, MO- 
2974185, P, US-1084517, US-2561239, US-91910 
(fragm. ex P)]. 

= Muhlenbergia peruviana (P. Beauv.) Steud. 


Muhlenbergia quadridentata (Kunth) Trin., Gram. Unifl. 
Sesquifl. 194, t. Sb, f. 14. 1824. Basionym: 
Podosemum quadridentatum Kunth. 

NOTE: basionym p. 194, comb. in tab. 

DST.: Guat, Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Herrera- 
Arrieta, Y. (1998); McClure, F. A. (1955); McVaugh, 
R. (1983); Peterson, P. M. (2000); Reeder, C. G. 
(1994). 


SYN.: Muhlenbergia anomala E. Fourn.; Muhlenbergia 
gracilis (Kunth) Trin.; Muhlenbergia quadridentata 
(Kunth) Kunth; Muhlenbergia virescens subsp. 
quadridentata (Kunth) Y. Herrera; Podosemum 
gracile Kunth; Podosemum quadridentatum Kunth; 
Trichochloa gracilis (Kunth) Roem. & Schult; 
Trichochloa quadridentata (Kunth) Roem. & Schult. 


Muhlenbergia quadridentata (Kunth) Kunth, Revis. Gramin. 
1: 64. 1829. Basionym: Podosemum quadridentatum 
Kunth. 

NOTE: isonym. 
E. NAME: Muhlenbergia quadridentata (Kunth) Trin. 
= Muhlenbergia quadridentata (Kunth) Trin. 


Muhlenbergia quitensis (Kunth) Hitche., Contr. U.S. Natl. 
Herb. 17(3): 292. 1913. Basionym: Calamagrostis 
quitensis Kunth. 

= Muhlenbergia tenuifolia (Kunth) Kunth. 


Muhlenbergia racemosa (Michx.) Britton, Sterns & 
Poggenb., Prelim. Cat. 67. 1888. Basionym: Agrostis 
racemosa Michx,. 

DST.: Cana, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Abrams, L. (1923); Correll, D. S. & 
M. C. Johnston (1970); Cronquist, A.. A. H. 
Holmgren, N. H. Holmgren & J. L. Reveal (1977); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Fernald, M. (1950); Gleason, H. A. & 
A. Cronquist (1968); Gleason, H. A. & A. Cronquist 
(1991); Great Plains Flora Association (1986); 
Hitchcock, A. S. (1951); Hitchcock, C. L., A. 
Cronquist, M. Ownbey & J. W. Thompson (1969); 
Moss, E. H. (1983); Munz, P. A. & D. D. Keck 
(1959); Peterson, P. M. (2000); Scoggan, H. J. (1978 
[1979]); Small, J. K. (1933); Voss, E. G. (1972). 

SYN.: Agrostis racemosa Michx.; Cinna racemosa (Michx.) 
Kunth; Muhlenbergia glomerata var. ramosa Vasey; 
Muhlenbergia racemosa var, ramosa (Wasey) Vasey 
ex Beal; Polypogon racemosus (Michx.) Nutt.; Vilfa 
racemosa (Michx.) P. Beauv. 


Muhlenbergia racemosa var. brevifolia (Vasey) Beal, Grass. 
N. Amer. 2: 253. 1896. Basionym: Muhlenbergia 
glomerata var. brevifolia Vasey. 

= Muhlenbergia californica Vasey. 


Muhlenbergia racemosa var. cinnoides (Link) B. Boivin, 
Naturaliste Canad. 94: 526. 1967. Basionym: 
Dactylogramma cinnoides Link. 

= Muhlenbergia glomerata (Willd.) Trin. 


Muhlenbergia racemosa vat. ramosa (Vasey) Vasey ex Beal, 
Grass. N. Amer. 2: 253. 1896.  Basionym: 
Muhlenbergia glomerata var. ramosa Vasey. 

= Muhlenbergia racemosa (Michx.) Britton, Sterns & 
Poggenb. 


Muhlenbergia racemosa subsp. violacea Scribn., Rhodora 
9(98): 22. 1907. T-Protol.. USA: New York, North 
Hannibal: 1883, OLE. Pearce s.n. 

= Muhlenbergia glomerata (Willd.) Trin. 


Muhlenbergia ramosissima Vasey, Bull. Torrey Bot. Club 
13(12): 231. 1886. T-Protol.: Mexico: Chihuahua: in 
SW Chihuahua, 1885, Dr. FE. Palmer [158]. T- 
Specim.: | of 1. Mexico: Chihuahua: SW Chihuahua, 
Aug-Nov 1885, E. Palmer 158 [LT: NY; ILT: LE, 


Catalogue of New World Grasses: II. 


165 


MO-2974152, P, US-995580], LT designated by 
Hitchcock, N. Amer. Fl. 27: 441 (1935) but without 
indicating the specific specimen; also by Peterson, 
Syst. Bot. Monogr. 31: 61 (1991) where the specific 
specimen is designated. 

= Muhlenbergia microsperma (DC.) Kunth. 


Muhlenbergia ramulosa (Kunth) Swallen, Contr, U.S. Natl. 
Herb. 29(4): 205. 1947. Basionym: Vilfa ramulosa 
Kunth. 

NOTE: page 204 in Index of Grass Species. 

DST.: Arge, CoRi, Guat, Hond, Mexi, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); McClure, F. A. (1955); 
McVaugh, R. (1983); Molina R., A. (1975); Peterson, 
P. M. (2000); Pohl, R. W. (1980); Reeder, C. G. 
(1994); Zuloaga, F. O., E. G. Nicora, Z. E. Ragolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 


(1994). 
SYN.: Agrostis ramulosa (Kunth) Roem. & Schult.; 
Muehlenbergia ramulosa (Kunth) — Swallen; 


Muhlenbergia wolfii (Vasey) Rydb.,; Sporobolus 
racemosus Vasey; Sporobolus ramulosus (Kunth) 
Kunth; Sporobolus wolfii Vasey; Vilfa ramulosa 
Kunth. 


Muhlenbergia reederorum Soderstr., Contr. U.S. Natl. 
Herb. 34(4):; 123. 1967. T-Protol.: Mexico: Durango: 
Sierra Madre Occidental, W of Ciudad Durango, 7 mi 
E of Navios, elev. 7700 ft, 26 Sep 1963,/. R. Reeder 
& C. G. Reeder 3834 [HT: YU; IT: F, K, MEXU, 
MICH, NY, P, UC, US]. T-Specim.: | of 1. Mexico: 
Durango, 26 Sep 1963, /.R. Reeder & C.G. Reeder 
3834 [HT: HT: YU; IT: F, K, MEXU, MICH, NY, P, 
UC, US-2443512]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 

A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000), 
Peterson, P. M. (2000); Soderstrom, T. R. (1967). 


Muhlenbergia repens (J. Presl) Hitche., Fl. Calif. 1: 111. 
1912. Basionym: Sporobolus repens J. Presl. 

DST.: Mexi, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Abrams, L. (1923); Beetle, A. A. 

(1977); Correll, D. S. & M. C. Johnston (1970); 

Espejo Serna, A., A. R. L6pez-Ferrari, and J. Valdes- 

Reyna (2000); Gould, F. W. & R. Moran (1981); 

Hitchcock, A. S. (1951); McVaugh, R. (1983); 

Morden, C. W. & S. L. Hatch (1996); Peterson, P. M. 

(2000). 

Muhlenbergia abata \.M. Johnst.,; Muhlenbergia 

subtilis. Nees; Sporobolus repens J. Presl, Vilfa 

repens (J. Presl) Trin. 


SYN.: 


Muhlenbergia reverchonii Vasey & Scribn., Contr. US. 
Natl. Herb. 3(1): 66. 1892. T-Protol.: USA: Texas, J. 
Reverchon 73 [HT: US-81635]. T-Specim.: 1 of 1. J. 
Reverchon 73 (HT: US-81635]. 

DST.: USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Darke, R. (1999); Espejo 


Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Hitchcock, A. S. (1951); Peterson, P. M. 
(2000). 

SYN.: Podosemum reverchonii (Vasey & Scribn.) Bush. 


Muhlenbergia richardsonis (Trin.) Rydb., Bull. Torrey Bot. 
Club 32(11): 600. 1905. Basionym: Vilfa richardsonis 
Trin. 

NOTE: as "Richardsonis". 

DST.: Cana, Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Cody, W. J. 
(1996); Cronquist, A., A. H. Holmgren, N. H. 
Holmgren & J. L. Reveal (1977); Espejo Serna, A., 
A. R. L6opez-Ferrari, and J. Valdes-Reyna (2000); 
Fernald, M. (1950); Gleason, H. A. & A. Cronquist 
(1968); Gleason, H. A. & A. Cronquist (1991); 
Gould, F. W. & R. Moran (1981); Great Plains Flora 
Association (1986); Hickman, J. C. (1993); 
Hitchcock, A. S. (1951); Hitchcock, C. L., A. 
Cronquist, M. Ownbey & J. W. Thompson (1969); 
Hultén, E. (1968); Morden, C. W. & S. L. Hatch 
(1996); Moss, E. H. (1983); Munz, P. A. & D. D. 
Keck (1959); Munz, P. A. (1974); Peterson, P. M. 
(2000); Porsild, A. E. & W. Cody (1980); Scoggan, 
H. J. (1978 [1979]); Voss, E. G. (1972); Welsh, S. L. 
(1974). 

Muhlenbergia aspericaulis Nees ex Trin; 
Muhlenbergia brevifolia var. richardsonis (Trin.) 
ME. Jones; Muhlenbergia squarrosa (Trin.) Rydb.; 
Sporobolus  aspericaulis — Scribn., — Sporobolus 
depauperatus (Torr. ex Hook.) Scribn.; Sporobolus 
richardsonis (Trin.) Merr.; Vilfa depauperata Torr. 
ex Hook.; Vilfa richardsonis Trin.; Vilfa squarrosa 
Trin. 


SYN.: 


Muhlenbergia rigens (Benth.) Hitchce., J. Wash. Acad. Sci. 
23(10): 453. 1933. Basionym: Epicampes rigens 
Benth. 

E. NAME: Muhlenbergia rigens (Benth.) Hitche. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Darke, R. (1999); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. & 
R. Moran (1981); Hickman, J. C. (1993); Hitchcock, 
A. S. (1951); McVaugh, R. (1983); Munz, P. A. & D. 
D. Keck (1959); Munz, P. A. (1974); Peterson, P.M. 
(2000); Shreve, F. & I. L. Wiggins (1964). 


SYN.: Crypsinna rigens (Benth.) M.E. Jones; Epicampes 
leptoura Piper; Epicampes _ rigens Benth.; 
Muhlenbergia leptoura (Piper) Hitchce.; 


Muhlenbergia marshii 1.M. Johnst., Muhlenbergia 
mundula 1.M. Johnst. 


Muhlenbergia rigida (Kunth) Trin., Gram. Unifl. Sesquifl. 
194. 1824. Basionym: Podosemum rigidum Kunth. 
NOTE: nom. inval., Trinius did not explicitly make the 
combination in either the text or the figure list. 
= Muhlenbergia rigida (Kunth) Kunth. 


Muhlenbergia rigida (Kunth) Kunth, Revis. Gramin. 1: 63. 
1829. Basionym: Podosemum rigidum Kunth. 
DST.: Arge, Boli, Colo, CoRi, Ecua, Mexi, Pana, Peru, 
USA, Vene. 
LVL. ACCPTC.: 1; cultivated. 


166 Catalogue of New World Grasses: II. 


ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Brako, L. & J. L. Zarucchi (1993); 
Breedlove, D. E. (1986); Correll, D. S. & M. C, 
Johnston (1970); Darke, R. (1999); Davidse, G., M. 
Sousa S. & A. O, Chater (editores generales) (1994); 
Franquemont, C., T. Plowman, E. Franquemont, S. R. 
King, C. Niezgoda, W. Davis & C. R. Sperling 
(1990); Gould, F. W. & R. Moran (1981); Hitchcock, 
A. S. (1927); Hitchcock, A. S. (1951); Jorgensen, P. 
M. & S. Leén-Yanez (eds.) (1999); Macbride, J. F. 
(1936); McVaugh, R. (1983); Reeder, C. G. (1994); 
Renvoize, S. A. (1998); Saravia, E. F. (1996); Tovar, 
O. (1993); Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo 
de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 

SYN.:  Muhlenbergia  affinis Trin; ~~ Muhlenbergia 
berlandieri Trin.; Muhlenbergia elegans (Kunth) 
Trin.; Muhlenbergia elegans var. atroviolacea 
Kuntze; Muhlenbergia elegans var. subviridis 
Kuntze; Muhlenbergia glabrata (Kunth) Trin.; 
Muhlenbergia holwayorum Hitche.; Muhlenbergia 
metcalfei M.E. Jones; Muhlenbergia phragmitoides 
Griseb.; Muhlenbergia — phragmitoides var. 
phragmitoides; Muhlenbergia rigida (Kunth) Trin.; 
Podosemum affine (Trin.) Bush; Podosemum elegans 
Kunth; Podosemum glabratum Kunth; Podosemum 
rigidum Kunth; Trichochloa elegans (Kunth) Roem. 
& Schult.; 7richochloa rigida (Kunth) Roem. & 
Schult. 


Muhlenbergia robusta (E. Fourn.) Hitchc., N. Amer. FI. 
17(6): 462. 1935. Basionym: Epicampes robusta E. 
Fourn. 

DST.: Guat, Hond, Mexi, Nica. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994): Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Peterson, P. M. (2000): Reeder, 
C. G. (1994); Soderstrom, T. R. (1967). 

SYN.: Epicampes berlandieri FE. Fourn.; Epicampes 
berlandieri E, Fourn. ex Hemsl.; Epicampes macrotis 
Piper; Epicampes minutiflora Mez; Epicampes 
robusta E, Fourn.; Epicampes robusta E. Fourn. ex 
Hemsl.; Epicampes stricta J. Presl, Muehlenbergia 
robusta (EF. Fourn.) — Hitche.; Muhlenbergia 
fournieriana Hitche.; Muhlenbergia macrotis (Piper) 
Hitche.; Muhlenbergia meziana Hitche.; 
Muhlenbergia presliana Hitche. 


Muhlenbergia scabra (Kunth) Trin. & Rupr., Sp. Gram. 
Stipac. 183. 1842. Basionym: Streptachne scabra 
Kunth, 

= Muhlenbergia watsoniana Hitchc. 


Muhlenbergia scabra 8. Watson, Proc. Amer. Acad. Arts 18: 
174. 1883. T-Protol.: Mexico: at San Luis Potosi, 
Schaffner 1067. T-Specim.: | of 1. Mexico: San Luis 
Potosi: San Luis Potosi, Schaffner 1067 HT: GH; IT: 
US-397161, 822842]. 

NOTE: hom. illeg. 
E. NAME: Muhlenbergia scabra (Kunth) Trin. & Rupr. 
= Muhlenbergia watsoniana Hitchce. 


Muhlenbergia schaffneri E. Fourn. ex Hemsl., Biol. Cent.- 
Amer., Bot. 3: 542. 1885, 


NOTE: nom. nud. 
= Muhlenbergia depauperata Scribn. 


Muhlenbergia schaffneri E. Fourn., Mexic. Pl. 2: 85. 1886. T- 
Specim.: | of 3. Mexico: Tacubaya, W. Schaffner s.n. 
[LT: P; ILT: LE (2 sheets), MO, P, US-3412353, 
2312354 (fragm. ex P)], LT designated by Hitchcock, 
Contr. U.S. Natl Herb. 17(3):294. 1913. 2 of 3. 
Mirador, Schaffner 142 [ST: P: IST: US-3412354 
(fragm. ex P)]. 3 of 3. Mexico: Circa urbem Mexico in 
collibus pr. Tacubaya, Sept., Schaffner 50 |ST: P; IST: 
US-3412352]. 

= Muhlenbergia depauperata Scribn. 


Muhlenbergia schaffneri var. longiseta Scribn., Proc. Acad. 
Nat. Sci. Philadelphia 1891: 297. 1891. T-Protol.: 
Mexico: foothills of the Rio Hondo, 10 mi W of the 
City of Mexico, growing on thin soil of ledges, 25 Aug 
1890, C.G. Pringle 3444. T-Specim.: | of 1. Mexico: 
foothills of the Rio Hondo, 25 Aug 1890,Pringle 3444 
[HT: VT]. 

= Muhlenbergia brevis C.O, Goodd. 


Muhlenbergia schaffneri C.O. Goodd. var. schaffneri. 
= Muhlenbergia depauperata Scribn. 


Muhlenbergia schmitzii Hack, Ann. K.K. Naturhist. 
Hofmus. 17: 255. 1902. T-Protol.: Mexico, Schmitz 
380 [HT: W]. T-Specim.: | of 1. Mexico, no date, A. 
Schmitz 380 (HT: W; IT: US-84832]. 

NOTE: as "Muehlenbergia" schmitzii. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Peterson, P. M. & C. R. Annable (1991); Peterson, P. 
M. (2000). 

SYN.: Muhlenbergia diehlii ME. Jones. 


Muhlenbergia schreberi J.F. Gmel., Syst. Nat. 2: 171. 1791. 
T-Protol.: — [USA: presumably Pennsylvania), 
[Muhlenberg]. 

NOTE: description given in Schreb. Gen. Plant. Linn. I. p. 
44. n. 103. 1789. However, no species is mentioned. 

DST.: Arge, Braz, Mexi, USA, Urug. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Burkart, A. (1969): Cabrera, A. L. (1970): 
Correll, D. S. & M. C. Johnston (1970); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Fernald, M. (1950); Gleason, H. A. & A. 
Cronquist (1968); Gleason, H. A. & A. Cronquist 
(1991); Godfrey, R. K. & J. W. Wooten (1979); 
Great Plains Flora Association (1986); Hickman, J. 
C. (1993); Hitchcock, A. S. (1951); Munz, P. A. 
(1968); Peterson, P. M. (2000): Radford, A. E., H. E. 
Ahles & C. R. Bell (1968); Rosengurtt. B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970): 
Schwegman, John E. (1991); Scoggan, H. J. (1978 
[1979]); Small, J. K. (1933); Smith, L. B., D. C. 
Wasshausen & R. M. Klein (1982); Voss, E. G. 
(1972), Wunderlin, R. P. (1998); Zuloaga, F. O., E. 
G. Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

SYN.: Agrostis apetala Bosc ex Trin., Anthipsimus 
gonopodus Rat.; Cynodon diffusus (Willd.) Raspail; 
Dilepyrum — diffusum PP. Beauv.; — Dilepyrum 
minutiflorum Michx.; Muhlenbergia  botteri E. 


Catalogue of New World Grasses: II. 


167 


Fourn.; Muhlenbergia diffusa Muhl.; Muhlenbergia 
diffusa Schreb., Muhlenbergia diffusa Willd.; 
Muhlenbergia — minutiflora — (Michx.) — Hitche.; 
Muhlenbergia palustris Scribn.; Muhlenbergia 
schreberi subsp. palustris (Scribn.) — Scribn.; 
Muhlenbergia — schreberi — subsp. _schreberi, 
Muhlenbergia schreberi var. palustris (Scribn.) 
Scribn. ex B.L. Rob. 


Muhlenbergia schreberi subsp. curtisetosa Scribn., Rhodora 
9(98): 17. 1907. T-Protol.: USA: Illinois, 1881,/. Wolf 
s.n. T-Specim.: | of 1. USA: Illinois, 1881,/. Wolfs.n. 
{IT: US-952419]. 
= Muhlenbergia x curtisetosa (Scribn.) Bush (pro sp.). 


Muhlenbergia schreberi var. curtisetosa (Scribn.) Steyerm. & 
C.L. Kucera, Rhodora 63(745): 25. 1961. Basionym: 
Muhlenbergia schreberi subsp. curtisetosa Scribn. 

NOTE: as "Schreberi". 
= Muhlenbergia x curtisetosa (Scribn.) Bush (pro sp.). 


Muhlenbergia schreberi subsp. palustris (Scribn.) Scribn., 
Rhodora 9(98): 17. 1907. Basionym: Muhlenbergia 
palustris Scribn. 

NOTE: subsp. is implied by 1904 code. 
= Muhlenbergia schreberi J.F. Gmel. 


Muhlenbergia schreberi var. palustris (Scribn.) Scribn. ex 
B.L. Rob., Rhodora 10(112): 65. 1908. Basionym: 
Muhlenbergia palustris Scribn. 

= Muhlenbergia schreberi J.F. Gmel. 


Muhlenbergia schreberi J.F. Gmel. subsp. schreberi. 
= Muhlenbergia schreberi J.F. Gmel. 


Muhlenbergia scoparia Vasey, Contr. U.S. Natl. Herb. 1(8): 
283. 1893. T-Specim.: | of 2. Mexico: Jalisco: rocky 
hills near Guadalajara, 10 Nov. 1893, C.G. Pringle 
2350 [LT: US-999069; ILT: LE, MO-1837823, TEX, 
US-91909, US-822950], LT designated by Hitchcock, 
N. Amer. FI. 17(6): 457 (1935), but without explicitly 
citing a specific sheet in a specific herbarium; fully 
lectotypified by Sodertrom, Contr. U.S. Natl. Herb. 
34(4): 144 (1967), who designated the US specimen as 
lectotype. 2 of 2. Mexico: Chihuahua, Batopilas, 1885, 
Dr, E. Palmer (ST: US-999070]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Peterson, P. M. (2000); 
Soderstrom, T. R. (1967). 

SYN.: Muhlenbergia carinata Mez. 


Muhlenbergia seatonii Scribn., Proc. Amer. Acad. Arts 28: 
122. 1893. T-Protol.: Mexico: Veracruz: hills near 
Esperanza, 8000 ft, August 1891, Seaton 320. T- 
Specim.: 1 of 1. Mexico: Veracruz: hills near 
Esperanza, 8000 fi, 14 Aug 1891,H.E. Seaton 320 {IT: 
US-995669]. 

NOTE: as Muhlenbergia "Seatoni". 

DST.: Mext. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Peterson, P. M. (2000). 


Muhlenbergia sericea (Michx.) P.M. Peterson, comb. nov. 
Basionym: Stipa sericea Michx., Fl. Bor.-Amer. 1: 54. 
1803. 

DST.: USA. 

SYN.: Agrostis sericea (Michx.) Elliott; Muhlenbergia 
capillaris var. filipes (M.A. Curtis) Chapm. ex Beal; 
Muhlenbergia filipes M.A. Curtis; Podosemum filipes 
(M.A. Curtis) Bush; Polypogon sericeus (Michx.) 
Spreng.; Stipa sericea Michx. 


Muhlenbergia setarioides E. Fourn., Mexic. Pl. 2: 84. 1886. 
T-Protol.: Mexico, 7 syntypes cited. T-Specim.: | of 6. 
Mexico: Valle de Orizaba, W. Schaffner 16] [ST: P: 
IST: LE, MO-2940108 (fragm.)], W. Schaffner pl. 
Mexic. 2 of 6. Mexico, Thomas s.n. [ST: herb. 
Buchinger; IST: MO]. 3 of 6. Mexico, Mueller 2022 
[ST: IST: US (fragm. ex. LE)]. 4 of 6. Mexico, Botteri 
& Sum. 1202 [ST]. 5 of 6. Mexico, Botteri & Sum. 
1279 [ST]. 6 of 6. Mexico, Bourgeau 3362 [ST: P; 
IST: MO-2974294, US (fragm. ex P), US (fragm. ex 
LE)]. 

DST.: CoRi, ElSa, Guat, Hond, Mexi, Pana. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Berendsohn, 
W. G. & A. E. Araniva de Gonzalez (1991); 
Breedlove, D. E. (1986); Davidse, G., M. Sousa S. & 
A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); McClure, F. A. (1955); Molina R., A. (1975); 
Peterson, P. M. (2000); Pohl, R. W. (1980); Reeder, 
C. G. (1994). 

SYN.: Muhlenbergia polypogonoides Hack.,; Muhlenbergia 
sylvatica var. setarioides (E. Fourn.) Beal. 


Muhlenbergia setifolia Vasey, Bot. Gaz. 7: 92. 1882. T- 
Protol.: USA: Texas: on Guadalupe Mts. of Western 
Texas, V. Havard s.n. T-Specim.: | of 1. USA: Texas, 
1881, V. Havard s.n. [HT: US-81634]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Hitchcock, A. S. (1951); Peterson, P. 
M. (2000). 


Muhlenbergia setiglumis (S. Watson) A. Nelson & J.F. 
Macbr., Bot. Gaz. 61(1): 30. 1916. Basionym: 
Muhlenbergia sylvatica var. setiglumis S. Watson. 

= Muhlenbergia mexicana var. filiformis (Torr.) Scribn. 


Muhlenbergia setosa (Kunth) Trin., Gram. Unifl. Sesquifl. 
193. t. 5, f. 22. 1824. Basionym: Podosemum setosum 
Kunth. 

NOTE: basionym p. 193, comb. in tab. 
= Muhlenbergia microsperma (DC.) Kunth. 


Muhlenbergia setosa (Kunth) Kunth, Revis. Gramin. 1: 63. 
1829. Basionym: Podosemum setosum Kunth. 
NOTE: isonym. 
E. NAME: Muhlenbergia setosa (Kunth) Trin. 
= Muhlenbergia microsperma (DC.) Kunth, 


Muhlenbergia setosa (Biehler) Trin. ex B.D. Jacks.. Index 
Kew. 2: 269. 1894. Basionym: Polypogon setosus 
Biehler. 

NOTE: hom, illeg. 
E. NAME: Muhlenbergia setosa (Kunth) Trin. 


168 


Catalogue of New World Grasses: II. 


= Muhlenbergia glomerata (Willd.) Trin. 


Muhlenbergia setosa var. cinnoides (Link) Fernald, Rhodora 
45(534): 238, pl. 757, map 2. 1943. Basionym: 
Dactylogramma cinnoides Link. 

= Muhlenbergia glomerata (Willd.) Trin. 


Muhlenbergia shepherdii (Vasey) Swallen, Contr. U.S. Natl. 
Herb. 29(4): 204. 1947. Basionym: Sporobolus 
shepherdii Vasey. 

= Blepharoneuron shepherdii (Vasey) P.M. Peterson & 
Annable. 


Muhlenbergia simplex (Scribn.) Rydb., Bull. Torrey Bot. 
Club 32(11): 600. 1905. Basionym: Sporobolus 
simplex Scribn. 

NOTE: hom. illeg. 
E. NAME: Muhlenbergia simplex (Spreng.) Kunth. 
= Muhlenbergia filiformis (Thurb. ex S. Watson) Rydb. 


Muhlenbergia sinuosa Swallen, Contr. U.S. Natl. Herb. 
29(4): 204. 1947. T-Protol.: USA: New Mexico: Grant 
Co.: on the W side of San Luis Mts., 26 Sep 1893, E.A. 
Mearns 2457 (HT: US-234817]. T-Specim.: | of 1. 
USA: New Mexico: Grant Co.: W side of the San Luis 
Mts., 26 Sep 1893, E. A. Mearns 2457 [HT: US- 
234817; IT: K, US-1910761]. 

DST.: Mexi, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Hitchcock, A. S. (1951); Peterson, P. M. & C. R. 
Annable (1991); Peterson, P. M. (2000). 

SYN.: Sporobolus confusus var. aberrans M.E. Jones. 


Muhlenbergia sobolifera (Muhl. ex Willd.) Trin., Gram. 
Unifl. Sesquifl. 189, 297, t. 5, f. 4. 1824. Basionym: 
Agrostis sobolifera Muhl, ex Willd. 

NOTE: as "Miihlenbergia". 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Fernald, M. (1950); Gleason, 
H. A. & A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Great Plains Flora Association 
(1986); Hitchcock, A. S. (1951); Peterson, P. M. 
(2000); Radford, A. E., H. E. Ahles & C. R. Bell 
(1968); Small, J. K. (1933). 

SYN.: Achnatherum soboliferum (Muhl. ex Willd.) P. 
Beauv.; Agrostis sobolifera Muhl. ex Willd.; Cinna 
sobolifera (Muhl. ex Willd.) Link; Muhlenbergia 
sobolifera fo. setigera (Scribn.) Deam; Muhlenbergia 
sobolifera fo. sobolifera; Muhlenbergia sobolifera 
subsp. setigera Scribn.; Muhlenbergia sobolifera 
subsp. sobolifera,; Muhlenbergia sobolifera var. 
setigera (Scribn.) Hitche, & Chase; Muhlenbergia 
sobolifera var. sobolifera, Podosemum soboliferum 
(Muhl. ex Willd.) Link; Trichochloa  sobolifera 
(Muhl. ex Willd.) Trin. 


Muhlenbergia sobolifera fo.setigera (Scribn.) Deam, Indiana 
Dept. Cons., Publ. 82: 163. 1929. Basionym: 
Muhlenbergia sobolifera subsp. setigera Scribn. 

NOTE: basionym not explicitly cited, only the basionym 
author cited. 
= Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. 


Muhlenbergia sobolifera subsp. setigera Scribn., Rhodora 
9(98): 18. 1907. T-Specim.: | of 1. USA: Texas, 1879, 
J. Reverchon 70 (HT: US-49863]. 
= Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. 


Muhlenbergia sobolifera var. setigera (Scribn.) Hitchc. & 
Chase, Man. Grasses U.S. (ed. 2) 906. 1950. 
Basionym: Muhlenbergia sobolifera subsp. setigera 
Scribn. 

= Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. 

Muhlenbergia sobolifera (Muhl. Trin. fo. 

sobolifera. 

= Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. 


ex Willd.) 


Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. subsp. 
sobolifera. 
= Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. 


Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. var. 
sobolifera. 
= Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. 


Muhlenbergia speciosa Vasey, Bull. Torrey Bot. Club 
13(12): 231. 1886. T-Protol.: Mexico: Chihuahua: in 
S.W. Chihuahua, 1885, Palmer 30 [HT: US]. T- 
Specim.: | of 1. Mexico: Chihuahua: S.W. Chihuahua, 
Aug-Nov 1885, Edward Palmer 30 [HT: US-999062; 
IT: LE, US-999063, US-822953]. 

DST.: Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Peterson, P. M. (1989); 
Peterson, P. M. (2000); Soderstrom, T. R. (1967). 

SYN.: Bealia speciosa (Vasey) Beal; Epicampes speciosa 
(Vasey) M.E. Jones. 


Muhlenbergia spicata (Spreng.) Kunth, Revis. Gramin. |: 64. 
1829. Basionym: Polypogon spicatus Spreng. 
= Gymnopogon spicatus (Spreng.) Kuntze. 


Muhlenbergia spiciformis Trin. Mem. Acad. Imp. Sci. 
Saint-Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. 
Nat. 6,4(3-4): 288. 1841. T-Protol.; Mexico: "Southern 
Mexico", Karwinsky s.n. T-Specim.: 1 of 1. Mexico, 
Karwinsky s.n. [T: LE]. 

NOTE: reprint p. 42. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Peterson, 
P.M. (2000); Reeder, C. G. (1994). 

Muhlenbergia acutifolia E. Fourn.; Muhlenbergia 
acutifolia E. Fourn, ex Hemsl.; Muhlenbergia 
parviglumis Vasey. 


SYN.: 


Muhlenbergia sprengelii Trin., Gram. Unifl. Sesquifl. 189, 
297, t. 5, f. 5. 1824. 
NOTE: nom. nov., as "Miihlenbergia" "Sprengelii". 
REPLACED NAME: 4rundo tenella Spreng. 
BLOCKING NAME: Arundo tenella Schrad. 
= Muhlenbergia tenella (Kunth) Trin. 


Muhlenbergia squarrosa (Trin.) Rydb., Bull. Torrey Bot. 
Club 36: 531. 1909. Basionym: Vilfa squarrosa Trin. 
= Muhlenbergia richardsonis (Trin.) Rydb, 


Catalogue of New World Grasses: II. 


169 


Muhlenbergia straminea Hitchc., Contr. U.S. Natl. Herb. 
17(3): 302. 1913. T-Protol.: Mexico: Chihuahua: in the 
western Sierra Madre at Tecovichu, in pine and oak 
woods, 2200 m alt., 10 Apr 1906, R. Endlich 1210 
[HT: US-691888]. T-Specim.: 1 of 1. Mexico: 
Chihuahua, 10 Apr 1906, R. Endlich 1210 [HT: US- 
691888]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Peterson, P. M. (2000); Reeder, C. 
G. & J. R. Reeder (1995). 


Muhlenbergia stricta (J. Pres!) Kunth, Enum. Pl. 1: 202. 
1833. Basionym: Podosemum strictum J. Pres. 

NOTE: also Revis. Gram. 3: Suppl. 1834 p. (as in 1835 ed.) 
XVI. 

DST.: Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Peterson, P. M. (2000). 

SYN.: Muhlenbergia elata Vasey; Muhlenbergia longifolia 
Vasey; Podosemum strictum J. Presl. 


Muhlenbergia strictior Scribn. ex Beal, Grass. N. Amer. 2: 
263. 1896. T-Protol.: Mexico, Pringle 1/418. T- 
Specim.: | of |. Mexico: Chihuahua, 17 Sep 1887, 
C.G. Pringle 1418 [LT: US-86380; ILT: K, MEXU, 
MSC, NY, US-822895, VT], LT designated by P.M. 
Peterson in Syst. Bot. Monographs, (1991). 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Peterson, P. M. & C. R. Annable (1991); Peterson, P. 
M. (2000). 

SYN.: Muhlenbergia flavida UNRANKED strictior (Scribn. 
ex Beal) Hitche.; Muhlenbergia madrensis MLE. 
Jones. 


Muhlenbergia subalpina Vasey, Descr. Cat. Grass. U.S. 40. 
1885, 
REPLACED NAME: 
breviaristata Vasey. 
= Muhlenbergia montana (Nutt.) Hitche. 


Muhlenbergia — gracilis var. 


Muhlenbergia subaristata Swallen, J. Wash. Acad. Sci. 
26(5): 208. 1936. T-Protol.: Mexico: Durango: along 
river, above Arroyo de Agua, El Salto, Sierra Madre 
Occidental, 2600-2650 m, 1 Sep 1934, F. W. Pennell 
18572 [HT: US-1614379]. T-Specim.: | of 1. Mexico: 
Durango, | Sep 1934, F.W. Pennell 18572 [HT: US- 
1614379]. 

DST.: Mex1. 

LVL. ACCPTC:.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Peterson, P. M. (2000). 


Muhlenbergia subbiflora Hitche., N. Amer. Fl. 17(6): 437. 
1935. T-Protol.: Mexico: Durango: collected on rich 
moist soil in garden, city of Durango, Nov 1896, 
Palmer 948 [HT: US-995434]. T-Specim.: 1 of 1. 
Mexico: Durango: City of Durango and vicinity, Apr- 
Nov 1896, E. Palmer 948 [HT: US-995434; IT: L, 
MEXU, MO-2974308 US-3155932, US-995433, US- 
746245]. 

= Chaboissaea subbiflora (Hitchc.) Reeder & C. Reeder. 


Muhlenbergia subtilis Nees, Linnaea 19(6): 689. 1847. T- 
Protol.: Mexico, Aschenb. exs. 206. T-Specim.: | of 1. 
Mexico, Aschenborn 206 [HT: B; IT: US (fragm. ex 
B)]. 

= Muhlenbergia repens (J. Pres) Hitchce. 


Muhlenbergia sylvatica (Torr.) Torr. ex A. Gray, N. Amer. 
Gram. |: 13. 1834. Basionym: Agrostis sylvatica Torr. 

DST.: Cana, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Correll, D. S. & M. C. Johnston 
(1970); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Godfrey, R. 
K. & J. W. Wooten (1979); Great Plains Flora 
Association (1986); Hitchcock, A. S. (1951); 
Peterson, P. M. (2000); Radford, A. E., H. E. Ahles 
& C. R. Bell (1968); Scoggan, H. J. (1978 [1979]); 
Voss, E. G. (1972). 

Agrostis diffusa Muhl.; Agrostis sylvatica Torr.; 
Muhlenbergia diffusa Farw.; Muhlenbergia sylvatica 
fo. sylvatica; Muhlenbergia sylvatica var. gracilis 
Scribn.; Muhlenbergia sylvatica var.robusta Fernald; 
Muhlenbergia sylvatica var. sylvatica, Muhlenbergia 
umbrosa Scribn. 


SYN.: 


Muhlenbergia sylvatica fo. attenuata (Scribn.) E.J. Palmer & 
Steyerm., Ann. Missouri Bot. Gard. 22: 467. 1935. 
Basionym: Muhlenbergia umbrosa subsp. attenuata 
Scribn. 

= Muhlenbergia frondosa fo. frondosa. 


Muhlenbergia sylvatica var. californica Vasey, Bot. Gaz. 7: 
93. 1882. T-Specim.: | of 1. USA: California: San 
Bernardino Co., Aug 1882,S.B. Parish & W.F. Parish 
1076 (HT: US-81639; IT: LE]. 

CORRECT NAME: Muhlenbergia parishii Vasey. 
= Muhlenbergia californica Vasey. 


Muhlenbergia sylvatica var. flexuosa Vasey, Rep. US. 
Geogr. Surv., Wheeler 6: 284. 1878. T-Protol.: USA. 
T-Specim.: 1 of 2. USA: New Mexico, 1874, C. 
Wright 731 [ST: USZ-81641], Also a lectotype of M. 
monticola Buckley. 2 of 2. USA: Arizona: Camp 
Crittenden, 1874, Rothrock 681 [LT: US-995253], LT 
designated by Hitchcock, N. Amer. FI. 17: 474 (1935). 

= Muhlenbergia tenuifolia (Kunth) Kunth. 


Muhlenbergia sylvatica var. gracilis Scribn., Trans. Kansas 
Acad. Sci. 9: 116. 1885. T-Protol.: USA: Kansas: 
Topeka, E.A. Popenoe. 

= Muhlenbergia sylvatica (Torr.) Torr. ex A. Gray. 


Muhlenbergia sylvatica var, pringlei Scribn., Bull. Torrey 
Bot. Club 9: 89, 1882. T-Protol.: USA: New Mexico: 
Dry cliffs, Santa Rita Mts., 7000 ft, July, Pringle 480. 
T-Specim.: | of |. USA: Arizona: Santa Rita Mts., 14 
Jul 1881, C.G. Pringle 480 (HT: US-746158]. 
= Muhlenbergia pauciflora Buckley. 


Muhlenbergia sylvatica var. robusta Fernald, Rhodora 
45(534): 326. 1943. T-Protol.: USA: Maine: Sydney, 
18 Aug 1916, Fernald & Long 12597 (HT: GH]. T- 
Specim.: 1 of 1. USA: Maine, Kennebec Co., Valley of 
Kennebec River, dry open clay river terraces, Sydney., 
18 Aug 1916, Fernald & Long 12597 |HT: GH; IT: 
US-3412350 (fragm. ex GH)]. 

= Muhlenbergia sylvatica (Torr.) Torr. ex A. Gray. 


170 


Catalogue of New World Grasses: II. 


Muhlenbergia sylvatica var. setarioides (E. Fourn.) Beal, 
Grass. N. Amer. 2: 249. 1896.  Basionym: 
Muhlenbergia setarioides E. Fourn. 

= Muhlenbergia setarioides E. Fourn. 


Muhlenbergia sylvatica var. setiglumis S. Watson, Botany 
Fortieth Parallel 378. 1871. T-Protol.: USA: Nevada: 
Near warm springs in Humboldt Pass, 6000 ft, Sep, 
[Watson] 1288. T-Specim.: | of 1. USA: Nevada, Sep 
1886, S. Watson 1288 (IT: US-952519]. 
= Muhlenbergia mexicana var. filiformis (Torr.) Scribn. 


Muhlenbergia sylvatica (Torr.) Torr. ex A. Gray fo. sylvatica. 
= Muhlenbergia sylvatica (Torr.) Torr. ex A. Gray. 


Muhlenbergia sylvatica (Yorr.) Torr. ex A. Gray var. 
sylvatica. 
= Muhlenbergia sylvatica (Torr.) Torr. ex A. Gray. 


Muhlenbergia sylvatica var. vulpina A. Wood, Amer. Bot. FI. 
pt. 2: 86. 1871. T-Protol.: USA: New York, HB. Lord. 
NOTE: Index of Grass Species has p. 386. 
= Status not yet determined. 


Muhlenbergia tenella (Kunth) Trin., Gram. Unifl. Sesquifl. 
192, 297, t. 5, f. 17. 1824. Basionym: Podosemum 
tenellum Kunth. 

NOTE: as "Miihlenbergia". 

DST.: Colo, CoRi, ElSa, Guat, Hond, Mexi, Nica, Pana. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Berendsohn, 
W. G. & A. E. Araniva de Gonzalez (1991); 
Breedlove, D. E. (1986); Davidse, G., M. Sousa S. & 
A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 


(2000); McClure, F. A. (1955); McVaugh, R. (1983); 


Molina R., A. (1975); Peterson, P. M. & C. R. 
Annable (1991); Peterson, P. M. (2000); Pohl, R. W. 
(1980); Reeder, C. G. (1994); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rigolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

SYN.: Arundo tenella Spreng.; Muhlenbergia exilis E. 
Fourn.; Muhlenbergia sprengelii Trin.; Podosemum 
tenellum Kunth; Polypogon — gracilis Spreng.; 
Polypogon tenellus (Kunth) Spreng.; Trichochloa 
tenella (Kunth) Roem. & Schult. 


Muhlenbergia tenuiflora (Willd.) Britton, Sterns & 
Poggenb., Prelim. Cat. 67. 1888. Basionym: Agrostis 
tenuiflora Willd. 

DST.: Cana, USA. 

LVL. ACCPTC.:: I. 

ALSO ACCPTD. BY: Anonymous (1986); Fernald, M. 
(1950); Gleason, H. A. & A. Cronquist (1968): 
Hitchcock, A. S. (1951); Peterson, P. M. (2000); 
Radford, A. E., H. E. Ahles & C. R. Bell (1968); 
Scoggan, H. J. (1978 [1979]); Small, J. K. (1933); 
Voss, E. G. (1972). 

SYN.: Agrostis tenuiflora Willd.; Apera tenuiflora (Willd.) 
P. Beauv.; Cinna tenuiflora (Willd.) — Link; 
Muhlenbergia tenuiflora subsp. tenuiflora; 
Muhlenbergia tenuiflora subsp. variabilis Scribn.: 
Muhlenbergia tenuiflora var. tenuiflora; 
Muhlenbergia tenuiflora var. variabilis (Scribn.) 
R.W. Pohl; Muhlenbergia  willdenowii — Trin.; 
Podosemum tenuiflorum (Willd.) Link; Trichochloa 
longiseta Trin.; Trichochloa tenuiflora (Willd.) 
Sweet. 


Muhlenbergia tenuiflora (Willd.) Britton, Sterns & Poggenb. 
subsp. tenuiflora. 
= Muhlenbergia tenuiflora (Willd.) Britton, Sterns & 
Poggenb. 


Muhlenbergia tenuiflora (Willd.) Britton, Sterns & Poggenb. 
» var. tenuiflora. 
= Muhlenbergia tenuiflora (Willd.) Britton, Sterns & 
Poggenb. 


Muhlenbergia tenuiflora subsp. variabilis Scribn., Rhodora 
9(98): 18. 1907. T-Protol.. USA: North Carolina: 
Rutherford: Chimney Mt., Biltmore Herb. 654A [HT: 
US]. T-Specim.: | of 1. USA: North Carolina: 
Rutherford Co., no date, Ex Herb. Biltmore 654A (HT: 
US-10020)]. 

= Muhlenbergia tenuiflora (Willd.) Britton, Sterns & 
Poggenb. 


Muhlenbergia tenuiflora var. variabilis (Scribn.) R.W. Pohl, 
Amer. Midl. Naturalist 82(2): 538. 1969. Basionym: 
Muhlenbergia tenuiflora subsp. variabilis Scribn. 

= Muhlenbergia tenuiflora (Willd.) Britton, Sterns & 
Poggenb. 


Muhlenbergia tenuifolia (Kunth) Trin., Gram. Unifl. Sesquifl. 
193, 277. 1824. Basionym: Calamagrostis tenuifolia 
Kunth. 

NOTE: nom. inval., Trinius did not explicitly make the 
combination in either the text or the figure list. 
= Muhlenbergia tenuifolia (Kunth) Kunth. 


Muhlenbergia tenuifolia (Kunth) Kunth, Revis. Gramin. 
1(4): 63. 1829. Basionym: Calamagrostis tenuifolia 
Kunth. 

DST.: Arge, Boli, Mexi, Peru, USA, Vene. 

LVL. ACCPTC:.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Brako, L. & J. 
L. Zarucchi (1993); Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); McVaugh, R. 
(1983); Peterson, P.M. & C. R. Annable (1991); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 


(1994). 

SYN.: Arundo quitensis (Kunth) Poir.; Arundo quitensis 
Spreng.;  Arundo  tenuifolia (Kunth)  Poir.; 
Calamagrostis mexicana Kunth; Calamagrostis 
quitensis Kunth; Calamagrostis tenuifolia Kunth; 
Muhlenbergia calamagrostidea Kunth, 
Muhlenbergia longiseta Benth., Muhlenbergia 


monticola Buckley; Muhlenbergia quitensis (Kunth) 
Hitche.; Muhlenbergia sylvatica var. flexuosa Vasey; 
Muhlenbergia tenuifolia (Kunth) Trin.; 
Muhlenbergia trichodes — Steud.; ~~ Podosemum 
tenuifolium (Kunth) Nees ex Nees & Schauer. 


Muhlenbergia tenuissima (J. Presl) Kunth, Enum. Pl. 1: 
198. 1833. Basionym: Podosemum tenuissimum J. 
Presl. 

NOTE: also Revis. Gram. 3: Suppl. 1834 p. (as in 1835 ed.) 
XVI. 

DST.: Boli, Braz, CoRi, Hond, Mexi, Nica, Pana, Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Brako, L. & J. 
L. Zarucchi (1993); Breedlove. D. E. (1986); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Foster, R. C. 


Catalogue of New World Grasses: II. 


171 


(1958); Hitchcock, A. S. (1927); Macbride, J. F. 
(1936); McVaugh, R. (1983); Molina R., A. (1975); 
Peterson, P. M. & C. R. Annable (1991); Peterson, P. 
M. (2000): Pohl, R. W. (1980); Reeder, C. G. (1994); 
Tovar, O. (1993); Vargas C., C. (1943). 
Muhlenbergia nebulosa Scribn. ex Beal; Podosemum 
tenuissimum J. Presl. 


SYN: 


Muhlenbergia texana Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 91. 1862. T-Protol.: Northern Texas, 
May. T-Specim.: | of 2. USA: Texas: W Texas to El 
Paso, May-Oct 1849, Wright 739 [LT: US-994219; 
ILT: MO], LT designated by Peterson & Annable, 
Syst. Bot. Monogr. 31: 93 (1991). This 
lectotypification now superseded by next screen. 2 of 
2. USA: northern Texas: [Agrostis barbatis Buck.], 
May, S.B. Buckley s.n. [LT: PH], LT designated by 
Dorr & Peterson, Sida 15: 590 (1993). 

DST.: Mexi, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W. & R. Moran (1981); Hitchcock, A. S. 
(1951); Peterson, P. M. & C. R. Annable (1991); 
Peterson, P. M. (2000); Shreve, F. & I. L. Wiggins 
(1964). 

Muhlenbergia buckleyana_ Scribn.,; Podosemum 
texanum (Buckley) Bush. 


SYN:: 


Muhlenbergia texana Thurb. ex Porter & J.M. Coult., Syn. 
Fl. Colorado 144. 1874. T-Protol.: Texas, 3 syntypes. 
T-Specim.: | of 3. USA: Texas: Rio San Pedro, 5 Nov 
1850, JM. Bigelow s.n. [ST]. 2 of 3. USA: Texas: 
Presidio del Norte, Jul 1852, C.C. Parry [ST]. 3 of 3. 
USA: Texas: Western Texas to El Paso, May-Oct 
1849, C. Wright 734 [ST]. 

NOTE: hom. illeg. 

E. NAME: Muhlenbergia texana Buckley. 

CORRECT NAME: Muhlenbergia porteri Scribn. ex Beal. 
= Muhlenbergia porteri Scribn. ex Beal. 


Muhlenbergia thurberi (Scribn.) Rydb., Bull. Torrey Bot. 
Club 32(11): 601. 1905. Basionym: Sporobolus 
thurberi Scribn. 

NOTE: as "Thurberi". 

DST.: USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Cronquist, A., 
A. H. Holmgren, N. H. Holmgren & J. L. Reveal 
(1977); Hitchcock, A. S. (1951); Peterson, P. M. 
(2000). 

SYN.: Muhlenbergia filiculmis M.E. Jones; Sporobolus 
filiculmis Vasey, Sporobolus filiculmis Vasey ex 
Beal; Sporobolus thurberi Scribn.; Vilfa_filiculmis 
Thurb. ex Scribn.; Vilfa filiculmis Thurb. ex Vasey. 


Muhlenbergia torreyana (Schult.) Hitche., Amer. J. Bot. 
21(3): 136. 1934. Basionym: Agrostis torreyana 
Schult. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Fernald, M. 
(1950); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Hitchcock, A. 
S. (1951); Peterson, P. M. (2000). 

Agrostis compressa Yort., Agrostis torreyana 
Schult.; Colpodium compressum Trin.; 


SYN.: 


Muehlenbergia torrevana— (Schult.) —_ Hitche.; 
Sporobolus compressus (Torr.) Kunth; Sporobolus 
torreyanus (Schult.) Nash; Vilfa compressa Trin. ex 
Spreng. 


Muhlenbergia torreyi (Kunth) Hitchc. ex Bush, Amer. Mid. 
Naturalist 6: 84. 1919. Basionym: Agrostis torreyi 
Kunth. 

DST.: Arge, Mexi, USA. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 

(1977); Correll, D. S. & M. C. Johnston (1970); 

Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 

Reyna (2000); Great Plains Flora Association (1986); 

Hitchcock, A. S. (1951); Peterson, P. M. & J. J. 

Ortiz-Diaz (1998); Peterson, P. M. (2000); Zuloaga, 

F. O., E. G. Nicora, Z. E. Rigolo de Agrasar, O. 

Morrone, J. Pensiero & A. M. Cialdella (1994). 

Agrostis caespitosa Torr.; Agrostis torreyi Kunth; 

Muhlenbergia gracillima  Torr.; Muhlenbergia 

nardifolia Griseb.; Podosemum gracillimum (Torr.) 

Bush. 


SYN.: 


Muhlenbergia trichodes Steud., Syn. Pl. Glumac. 1: 177. 
1854. T-Protol.: South America: "Am. austr.", T. 
"Guadalupe, Duchaessing" in Steud. script. but 
published locality is "amer. Austr." not W. 

= Muhlenbergia tenuifolia (Kunth) Kunth. 


Muhlenbergia trichopodes (Elliott) Chapm., Fl. South. U.S. 
553. 1860. Basionym: Agrostis trichopodes Elliott. 
= Muhlenbergia expansa (Poir.) Trin. 


Muhlenbergia trifida Hack., Repert. Spec. Nov. Regni Veg. 
8: 518. 1910. T-Protol.: Mexico: Michoacan, Arséne 
3217. T-Specim.: | of 1. Mexico: Michoacan: vicinity 
of Morelia, Quinceo, 11 Nov 1909, Bro. Arséne 3217 
[HT: W; IT: BM, MO-843315, US, US-86637 
(fragm.)]. 

= Muhlenbergia montana (Nutt.) Hitchce. 


Muhlenbergia trinii E. Fourn. ex Hemsl., Biol. Cent.-Amer., 
Bot. 3(19): 543. 1885. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Muhlenbergia diversiglumis Trin. 
= Muhlenbergia diversiglumis Trin. 


Muhlenbergia triniiE. Fourn., Mexic. Pl. 2: 84. 1886. 
NOTE: nom. nov., nom. illeg. superfl.; 4 diversiglumis 
Trin. cited. 
CORRECT NAME: Muhlenbergia diversiglumis Trin. 
= Muhlenbergia diversiglumis Trin. 


Muhlenbergia umbrosa Scribn., Rhodora 9(98): 20. 1907. 
REPLACED NAME: Agrostis sylvatica Yorr. 
BLOCKING NAME: Agrostis sylvatica Host. 
= Muhlenbergia sylvatica (Torr.) Torr. ex A. Gray. 


Muhlenbergia umbrosa fo. attenuata (Scribn.) Deam, Indiana 
Dept. Cons., Publ. 82: 171. 1929. Basionym: 
Muhlenbergia umbrosa subsp. attenuata Scribn. 

NOTE: basionym not explicitly cited, only the basionym 
author cited. 
= Muhlenbergia frondosa fo. frondosa. 


Muhlenbergia umbrosa subsp. attenuata Scribn., Rhodora 
9(98): 21. 1907. T-Protol.: USA: South Dakota: 
Aurora Co., E.N. Wilcox 25. T-Specim.: 1 of 1. USA: 


172 


Catalogue of New World Grasses: II. 


South Dakota: Aurora Co., E.N. Wilcox 25 [HT: US- 
10019]. 
= Muhlenbergia frondosa fo. frondosa. 


Muhlenbergia uniflora (Muhl.) Fernald, Rhodora 29(337): 
10. 1927. Basionym: Poa uniflora Muhl. 

DST.: Cana, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Fernald, M. (1950); Gleason, H. A. 
& A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Hitchcock, A. S. (1951); Peterson, 
P. M. (2000); Scoggan, H. J. (1978 [1979]); Voss, E. 
G. (1972). 

Agrostis serotina Torr.; Muhlenbergia uniflora var. 
terrae-novae Fernald; Muhlenbergia uniflora var. 
uniflora; Poa modesta Tuck., Poa stricta’ var. 
uniflora Muhl. ex Scribn. & Merr.; Poa uniflora 
Muhl.; Poa uniflora Muhl.; Sporobolus serotinus A. 
Gray; Sporobolus uniflorus (Muhl.) Scribn. & Merr.; 
Vilfa serotina Torr. ex A. Gray; Vilfa serotina Trin.; 
Vilfa tenera Trin. 


SYN.: 


Muhlenbergia uniflora var. terrae-novae Fernald, Rhodora 
29(337): 11-12. 1927. T-Protol.: Canada: 
Newfounland: moss and silicious rocks along rill, 
slope of South Hill, St. John's, 14 Aug 1924, Fernald, 
Long & Dunbar 26244 |HT: GH]. 

= Muhlenbergia uniflora (Muhl.) Fernald. 


Muhlenbergia uniflora (Muhl.) Fernald var. uniflora. 
= Muhlenbergia uniflora (Muhl.) Fernald. 


Muhlenbergia utilis (Torr.) Hitche., J. Wash. Acad. Sci. 
23(10): 453. 1933. Basionym: Vilfa utilis Torr. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Correll, D. S. & M. C. Johnston (1970); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Hickman, J. C. 
(1993); Hitchcock, A. S. (1951); McVaugh, R. 
(1983); Morden, C. W. & S. L. Hatch (1996); Munz, 
P. A. & D. D. Keck (1959); Munz, P. A. (1974); 
Peterson, P. M. (2000); Reeder, C. G. (1994). 

Muehlenbergia utilis (Yorr.) Hitche.; Sporobolus 
sacatilla Griseb. ex E. Fourn.; Sporobolus sacatilla 
Griseb. ex E. Fourn.; Sporobolus utilis (Torr.) 
Scribn.; Vilfa sacatilla E. Fourn.; Vilfa utilis Torr. 


SYN.: 


Muhlenbergia vaginata Swallen, Contr. U.S. Natl. Herb. 
29(9): 406. 1950. T-Protol.: Guatemala: Dept. of San 
Marcos: in spring-fed meadow along quebrada, 4 km 
from San Sebastian, NW of San Marcos, 15 Feb 1940, 
JA, Steyermark 35598 [HT: F-1046643]. T-Specim.: | 
of 1. Guatemala: San Marcos: road between San 
Sebastian and San Marcos, 2700-3800 m, 15 Feb 
1940, JA. Steyermark 33589 [HT: F-1046643; IT: US- 
2240531, US-2236472]. 

DST.: Guat, Mexi. 

LVL, ACCPTC:: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); McClure, 
F. A. (1955); MeVaugh, R. (1983); Peterson, P. M. & 
C. R. Annable (1991); Peterson, P. M. (2000); 
Reeder, C. G. (1994). 


Muhlenbergia vaseyana Scribn., Annual Rep. Missouri Bot. 
Gard. 10: 52. 1899. T-Protol.: USA: Arizona, Rothrock 
283. T-Specim.: | of 1. USA: Arizona, Jul 1874.7. 
Rothrock 282 (IT: US-81633]. 

NOTE: cited Epicampes distichophylla Vasey nom. nud., 
same as M. distichophylla Munro ex Vasey ex B. D. 
Jackson, nom. inval. 

= Muhlenbergia emersleyi Vasey. 


Muhlenbergia venezuelae Luces, Bol. Soc. Venez. Ci. Nat. 
15: 14. 1953. T-Protol.: Venezuela: Edo Merida:en 
Paramo de Misinta, Muchuchies: 27 Nov 1943, Z. 
Luces 281 (HT: [VEN] Hrbr. del Inst. Botanico del 
Ministerio de Agricultura y Cria]. 

DST.: Vene. 
LVL. ACCPTC.: 2. 
ALSO ACCPTD. BY: Peterson, P. M. (2000). 


Muhlenbergia versicolor Swallen, Contr. U.S. Natl. Herb. 
29(9): 412. 1950. T-Protol.: Mexico: Oaxaca: 170 km 
N of Oaxaca, 13 Dec 1945, FE. Hernandez X. & JA. 
Jenkins X-810 [HT: US-1961991]. T-Specim.: | of 1. 
Mexico: Oaxaca, 13 Dec 1945, FE. Hernandez- 
Xolocotzi & JA, Jenkins X-810 [HT: US-1961991], 

DST.: Guat, Hond, Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G.. M. Sousa S. & A. O. Chater 
(editores generales) (1994); Peterson, P. M. (2000); 
Reeder, C. G. (1994); Soderstrom, T. R. (1967). 

SYN.: Muhlenbergia distichophylla var. mutica Scribn. ex 
Urbina. 


Muhlenbergia villiflora Hitchc., N. Amer. Fl. 17(6): 470. 
1935, 

REPLACED NAME: Vilfa pubescens E. Fourn. 

BLOCKING NAME: Muhlenbergia pubescens (Kunth) 
Hitche. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Morden, C. W. & S. L. Hatch (1996); Morden, C. W. 
(1995): Peterson, P. M. (2000). 

SYN.: Vilfa pubescens E. Fourn, 


Muhlenbergia villiflora Hitchc. var. villiflora. 
DST.: Mexi. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Morden, C. W. & S. L. Hatch 
(1996); Morden, C. W. (1995). 


Muhlenbergia villiflora var. villosa (Swallen) Morden, 
Phytologia 79(1): 29. 1995. Basionym: Muhlenbergia 
villosa Swallen. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Morden, C. W. & S. L. Hatch 
(1996); Morden, C. W. (1995); Peterson, P. M. 
(2000). 

SYN.: Muhlenbergia villosa Swallen. 


Muhlenbergia villosa Swallen, J. Wash. Acad. Sci. 31(8): 
350, f. 2. 1941. T-Protol.: USA: Texas: 15 mi S of 
Stanton, 11 Jul 1928, B.C. Tharp 5048 [HT: US- 
1386596; IT: TEX]. T-Specim.: 1 of 1. USA: Texas, 
11 Jul 1928, B.C. Tharp 5048 [HT: US-1386596; IT: 
GH, MO, TEX]. 


Catalogue of New World Grasses: II. 


173 


= Muhlenbergia villiflora var. villosa (Swallen) Morden. 


Muhlenbergia virescens (Kunth) Trin., Gram. Unifl. 
Sesquifl. 193, t. 5a, f. 7. 1824. Basionym: Podosemum 
virescens Kunth. 

NOTE: basionym p. 193, comb. in tab. 

DST.: Mexi. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Herrera-Arrieta, 
Y. (1998); Peterson, P. M. (2000); Reeder, C. G. & J. 
R. Reeder (1995). 

Muhlenbergia curvula Swallen; Muhlenbergia 
virescens (Kunth) Kunth; Muhlenbergia virescens 
subsp. virescens, Podosemum virescens Kunth; 
Trichochloa virescens (Kunth) Roem. & Schult.. 


SYN: 


Muhlenbergia virescens (Kunth) Kunth, Revis. Gramin. 1: 
64. 1829. Basionym: Podosemum virescens Kunth. 
NOTE: isonym. 
E. NAME: Muhlenbergia virescens (Kunth) Trin. 
= Muhlenbergia virescens (Kunth) Trin. 


Muhlenbergia virescens subsp. quadridentata (Kunth) Y. 
Herrera, Amer. J. Bot. 81(8): 1043. 1994. Basionym: 
Podosemum quadridentatum Kunth. 

= Muhlenbergia quadridentata (Kunth) Trin. 


Muhlenbergia virescens (Kunth) Trin. subsp. virescens. 
= Muhlenbergia virescens (Kunth) Trin. 


Muhlenbergia virletii (E. Fourn.) Soderstr., Contr. US. 
Natl. Herb. 34(4): 157. 1967. Basionym: Epicampes 
virletii E. Fourn. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Peterson, P. M. 
(2000); Soderstrom, T. R. (1967). 

SYN.: Epicampes virletii E. Fourn. 


Muhlenbergia watsoniana Hitche., N. Amer. Fl. 17(6): 471. 
1935. 

NOTE: nom. nov., as Muhlenbergia "Watsoniana". 

REPLACED NAME: Muhlenbergia scabra S. Watson. 

BLOCKING NAME: Muhlenbergia scabra (Kunth) Trin. 
& Rupr. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Herrera-Arrieta, Y. (1998); McVaugh, R. (1983); 
Peterson, P. M. (2000). 

SYN. Muhlenbergia scabra (Kunth) Trin. & Rupr.; 
Muhlenbergia scabra S$. Watson. 


Muhlenbergia willdenowii Trin., Gram. Unifl. Sesquifl. 188, 
189, 297, t. 5, f. 3. 1824. 
NOTE: nom. illeg. superfl.; as 
"Willdenowii". 
CORRECT NAME: Agrostis tenuiflora Willd. 
= Muhlenbergia tenuiflora (Willd.) Britton, Sterns & 
Poggenb. 


"Miihlenbergia" 


Muhlenbergia wolfii (Vasey) Rydb., Bull. Torrey Bot. Club 
32(11): 600. 1905. Basionym: Sporobolus wolfii 
Vasey. 

NOTE: as "Wolfi". 
= Muhlenbergia ramulosa (Kunth) Swallen. 


Muhlenbergia wrightii Vasey ex J.M. Coult., Man. Bot. 
Rocky Mt. 409. 1885. T-Protol.: USA, /Wright 1986/. 
T-Specim.: | of 1. USA: New Mexico, 1851-1852,C. 
Wright 1986 [HT: US-556872]. 

DST.: Mexi, USA. 
LVL. ACCPTC.: 1; cultivated. 
ALSO ACCPTD. BY: Beetle, A. A. (1977); Cronquist, A., 

A. H. Holmgren, N. H. Holmgren & J. L. Reveal 

(1977); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 

Valdes-Reyna (2000); Gould, F. W. & R. Moran 

(1981); Hitchcock, A. S. (1951); Peterson, P. M. 

(2000). 

Muhlenbergia coloradensis Mez; Muhlenbergia 
wrightii var. annulata Vasey. 


SYN.: 


Muhlenbergia wrightii var. annulata Vasey, Descr. Cat. 
Grass. U.S. 41. 1885. T-Protol.: [USA: Arizona], 
[Lemmon 3179]. 

NOTE: nom. nud. 

= Muhlenbergia wrightii Vasey ex J.M. Coult. 


Muhlenbergia xanthodas Soderstr., Contr. U.S. Natl. Herb. 
34(4): 173. 1967. T-Protol.: Mexico: Chiapas: 
collected on rock on Mt. Ovando, 2300 m, 14-18 Nov 
1939, E. Matuda 4003 [HT: US-1817864; IT: F, 
MICH]. T-Specim.: | of 1. Mexico: Chiapas, 14 Nov 
1939-18 Nov 1939, E. Matuda 4003 [HT: US- 
1817864; IT: F, MICH, US-2075810}. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Peterson, 
P. M. (2000); Reeder, C. G. (1994); Soderstrom, T. 
R. (1967). 


Muhlenbergia xerophila C.O. Goodd., J. Wash. Acad. Sci, 
30: 19. 1940. T-Protol.: USA: Arizona: in Sycamore 
Canyon, 22 Sep 1937,L.N. Goodding M262 |HT: SCS 
Hrb. at Tucson, Ariz.; IT: RM, US]. T-Specim.: | of I. 
USA: Arizona: Santa Cruz Co.: Sycamore Canyon, 22 
Sep 1937, L.N. Goodding M262 |HT: USDA-SCS hrb. 
Tucson; IT: MO-5062477, RM, US-1761610]. 

= Muhlenbergia elongata Scribn. ex Beal. 


Muhlenbergia sect. Epicampes (J. Presl) Soderstr., Contr. 


U.S. Natl Herb. 34(4): 112. 1967. Basionym: 
Epicampes J. Presl. 

LVL. ACCPTC.: 1. 

Muhlenbergia Schreb. subg. Muhlenbergia, NONE 
LISTED. 

LVL. ACCPTC.: I. 

Muhlenbergia sect. Podosemum (Desv.) Pilg., Nat. 
Pflanzenfam. (ed. 2) 14d: 71. 1956. Basionym: 


Podosemum Desv. 
LVL. ACCPTC.: |. 


Muhlenbergia subg. Trichochloa A. Gray, Manual 546. 
1848. 

REPLACED NAME: Trichochloa DC. 

BLOCKING NAME: Trichochloa P. Beauv. 


LVL. ACCPTC.: I. 


174 


Catalogue of New World Grasses: II. 


Muhlenbergiinae subtrib. Pilg., Nat. Pflanzenfam. 
(ed. 2) 14d: 168, 67. 1956. TYPE: Muhlenbergia 
Schreb. 

TRTD. BY: Soreng, R. J., G. Davidse, T. S. Filgueiras, E. J. 
Judziewicz, P. M. Peterson, and F. O. Zuloaga. 

NHT: Cynodonteae. 

CNSLTD. LIT.: Duvall, M. R., P. M. Peterson & A. H. 
Christensen, (1994); Peterson, P. M., (2000). 


Munroa Torr., Pacif. Railr. Rep. 4: 158. 1857 Aug-Sep. 
TYPE: Munroa squarrosa (Nutt.) Torr, 
NOTE: misspelled as "Monro", but named after W. Munro. 
TRTD. BY: Valdés-Reyna, J. & P. M. Peterson. 
NHT: Munroinae. NO. ACCPTD. SPP.: 5. 
CNSLTD. LIT.: Anton, A. M. & A. T. Hunziker, (1978), 
SYN.: Hemimunroa (Parodi) Parodi. 


Munroa andina Phil., Verz. Antofagasta Pfl. 8: 90, 1891. T- 
Protol.: Chile: Tarapaca: ad Paroma 3800 m.s.m. lecta, 
F. Philippi s.n. T-Specim.: | of 1. Chile: Tarapaca: ad 
Paroma 3800 m.s.m. lecta, F. Philippi s.n. (IT: US- 
556619 (fragm. ex herb. Phillipi)). 

DST.: Arge, Boli, Chil. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Foster, R. C. (1958); Marticorena, C. 
& M. Quezada (1985); Renvoize, S. A. (1998): 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

Hemimunroa andina (Phil.) Parodi; Hemimunroa 
andina (Phil.) Parodi var. andina; Hemimunroa 
andina var. breviseta (Hack.) Parodi; Munroa andina 
var. andina, Munroa andina var. breviseta Hack.; 
Munroa multiflora Phil. 


SYN:: 


Munroa andina Phil. var. andina. 
= Munroa andina Phil. 


Munroa andina var. breviseta Hack., Annuaire Conserv. Jard. 
Bot. Geneve 17: 294. 1914. T-Protol.: Argentina: 
Jujuy: Estacion 3 Cruces, 9 Apr 1909, Lillo 9645 
[Stuckert hb. 22443], F.C.A Bol. T-Specim.: 1 of 1. 
Argentina: Jujuy: Estacion 3 Cruces, 9 Apr 1909,Lillo 
9645 [IT: US (fragm. ex herb. W)]. 

= Munroa andina Phil. 


Munroa argentina Griseb., Abh. Konigl. Ges. Wiss. 
Gottingen 24: 300. 1879. T-Protol.: Argentina. T- 
Specim.: | of 1. Argentina: Cordoba and Santiago del 
Estero, Laguna de Pocho, Minas. Ct.: pr. Atajo [T]. 

DST.: Arge, Boli, Chil. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Foster, R. C. (1958); Hitchcock, A. 
S. (1927); Matthei, O., C. Marticorena, R. Rodriguez, 
M. Kalin Arroyo, M. Mufioz, F. A. Squeo & G. 
Arancio (1997); Renvoize, S. A. (1998); Zuloaga, F. 
O., E. G. Nicora, Z. E. Ragolo de Agrasar, O. 
Morrone, J. Pensiero & A. M. Cialdella (1994). 


Munroa benthamiana Hack. ex Kuntze, Revis. Gen. Pl. 3(3): 
357. 1898. T-Protol.: Argentina: Cordoba: Dept. 
Minas, en los montes ralos al E de la laguna de Pocho, 
21 Feb 1876, Hieronymus 435. T-Specim.: | of 1. 
Argentina; Cordoba: Dept. Minas, 21 Feb 1876, 
Hieronymus 435 (HT: K; IT: CORD, US (fragm.)]. 


NOTE: Publ. as nom. nud. in Rev. Mus. La Plata 5: 303. 
1893, 
= Blepharidachne benthamiana (Hack. ex Kuntze) Hitchce. 


Munroa decumbens Phil., Verz. Antofagasta Pf]. 90. 1891. 
T-Protol.: Chile: Tarapaca: prope Calcalhuay lecta, F. 
Philippi s.n. 

DST.: Arge, Boli, Chil, Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Brako, L. & J. L. Zarucchi (1993); 
Foster, R. C. (1958); Hitchcock, A. S. (1927): 
Macbride, J. F. (1936); Marticorena, C. & M., 
Quezada (1985); Renvoize, S. A. (1998): Tovar, O. 
(1993); Zuloaga, F. O., E. G. Nicora, Z. E. Rigolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 


Munroa mendocina Phil., Anales Univ. Chile 36: 210. 1870. 
T-Protol.: /Philippi] s.n. T-Specim.: | of |. Phillippi 
s.n. (IT: US-1939962 (fragm. ex SGO)}. 
DST.: Arge. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 


Munroa multiflora Phil., Verz. Antofagasta Pfl, 90. 1891. T- 
Protol.: Chile: Tarapaca: pariter ad Calcalhuay lecta, 
F. Philippi s.n. 
= Munroa andina Phil. 


Munroa squarrosa (Nutt.) Torr., Pacif. Railr. Rep. 4(5): 
158. 1857. Basionym: Crypsis squarrosa Nutt. 

DST.: Cana, Mexi, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Correll, D. S. 
& M. C. Johnston (1970); Cronquist, A.. A. H. 
Holmgren, N. H. Holmgren & J. L. Reveal (1977); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Fernald, M. (1950); Great Plains Flora 
Association (1986); Hickman, J.C. (1993); 
Hitchcock, A. S. (1951); Hitchcock, C. L., A. 
Cronquist, M. Ownbey & J. W. Thompson (1969); 
Moss, E. H. (1983); Scoggan, H. J. (1978 [1979]). 
Crypsis squarrosa Nutt., Munroa squarrosa var. 
floccuosa Vasey ex Beal; Munroa squarrosa var. 
squarrosa. 


SYN.: 


Munroa squarrosa var. floccuosa Vasey ex Beal, Grass. N. 
Amer. 2: 456. 1896. T-Protol.: USA: Arizona [Peach 
Springs], Jones s.n. T-Specim.: | of 1. USA: Arizona, 
7 Aug 1884, ME. Jones 4088 [IT: US-899721]. 

= Munroa squarrosa (Nutt.) Torr. 


Munroa squarrosa (Nutt.) Torr. var. squarrosa. 
= Munroa squarrosa (Nutt.) Torr. 


Munroinae subtrib. Parodi ex P.M. Peterson, Sida 
16(3): 541. 1995, TYPE: Munroa Torr. 
TRID. BY: Soreng, R. J., G. Davidse, T. S. Filgueiras, E. J. 
Judziewicz, P. M. Peterson, and F. O. Zuloaga. 
NHT: Cynodonteae. 
CNSLTD. LIT.: Peterson, P. M.. R. D. Webster & J. 
Valdés-Reyna, (1995). 


Nardus gangitis L., Sp. Pl. 53. 1753. 
NOTE: nom. rej. prop. 1390, Taxon 48(1): 155-156 (1999), 
reccomended Taxon 49(4): 802 (2000). 


Catalogue of New World Grasses: II. 


175 


= Ctenium aromaticum (Walter) Alph. Wood. 


Nardus indica L. f., Suppl. Pl. 105. 1782. T-Specim.: | of 2. 
India: Habitat in Tranquebaria, Koenig s.n. [ST: 
LINN-73.8]. 2 of 2. India: "Habitat in Tranquebaria", 
Koenig s.n. [T: LINN-73.8]. 

= Microchloa indica (L. f.) P. Beauv. 


Nardus scorpioides Lam., Tabl. Encycl. 1: 152. 1791. T- 
Specim.: | of 1. America, no date, P. de Beauvois s.n. 
(IT: US-A865647 (fragm. ex P-LAM)]. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Nazia Adans., Fam. Pl. 2: 31, 582. 1763. TYPE: Nazia 
racemosa (L.) Kuntze. 
NOTE: nom. rejic. 
= Tragus Haller. 


Nazia aliena (Spreng.) Scribn., Bull. Div. Agrostol., 
U.S.D.A. 28, f. 324. 1899. Basionym: Lappago aliena 
Spreng. 

= Tragus berteronianus Schult. 


Nazia occidentalis (Nees) Scribn., Zoe 4(4): 386. 1894. 
Basionym: 7ragus occidentalis Nees. 
= Tragus berteronianus Schult. 


Nazia racemosa (L.) Kuntze, Revis. Gen. Pl. 2: 780. 1891. 
Basionym: Cenchrus racemosus L. 
= Tragus racemosus (L.) All. 


Nazia racemosa var. aliena (Spreng.) Scribn. & J.G. Sm., 
Bull. Div. Agrostol., U.S.D.A. 4: 12. 1897. Basionym: 
Lappago aliena Spreng. 

= Tragus berteronianus Schult. 


Nazieae trib. Hitchc., U.S.D.A. Bull. (1915-23) 772: 15. 
1920. 
NOTE: nom. illeg.. generic name nom. rej. 


Neeragrostis Bush, Trans. Acad. Sci. St. Louis 13(7): 178. 
1903. TYPE: Neeragrostis weigeltiana Bush. 
= Eragrostis Wolf. 


P.M. Peterson, 
1997, Basionym: 


Neeragrostis contrerasii (R.W. Pohl) 
Smithsonian Contr. Bot. 87: 35. 
Eragrostis contrerasii R.W. Pohl. 

= Eragrostis contrerasii R.W. Pohl. 


Neeragrostis hypnoides (Lam.) Bush, Trans. Acad. Sci. St. 
Louis 13: 180. 1903. Basionym: Poa hypnoides Lam. 
= Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb. 


Neeragrostis reptans (Michx.) Nicora, Revista Argent. 
Agron. 29: 5. 1963. Basionym: Poa reptans Michx, 
= Eragrostis reptans (Michx.) Nees. 


Neeragrostis weigeltiana Bush, Trans. Acad. Sci. St. Louis 
13(7): 178. 1903. T-Protol.: Surinam, 1827, Weigelt. 
NOTE: nom. altern., cites Poa weigeltiana Rchb. ex Trin., 
nom. inval. (see also Eragrostis w.). 
= Eragrostis reptans (Michx.) Nees. 


Neesiochloa Pilg., Repert. Spec. Nov. Regni Veg. 48: 
119. 1940. TYPE: Neesiochloa barbata (Nees) Pilg. 
TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: I. 


Neesiochloa barbata (Nees) Pilg., Repert. Spec. Nov. Regni 
Veg. 48: 119. 1940. Basionym: Calotheca barbata 
Nees. 

DST.: Braz. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Pilger, R. (1940); Renvoize, S. A. 
(1984). 

SYN.: Briza barbata (Nees) Trin.; Calotheca barbata Nees; 
Chascolytrum barbatum (Nees) Kunth. 


Neobouteloua Gould, Bol. Soc. Argent. Bot. 12: 106. 
1968. TYPE: Neobouteloua lophostachya (Griseb.) 
Gould. 

TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 1. 


Neobouteloua lophostachya (Griseb.) Gould, Bol. Soc. 
Argent. Bot. 12: 108, f. 1-2. 1968. Basionym: 
Bouteloua lophostachya Griseb. 

DST.: Arge, Chil. 

LVL. ACCPTC:.: I. 

ALSO ACCPTD. BY: Gould, F. W. (1968); Marticorena, C. 
& M. Quezada (1985); Zuloaga, F. O., E. G. Nicora, 
Z. E. Rigolo de Agrasar, O. Morrone, J. Pensiero & 
A. M. Cialdella (1994). 

SYN.: Bouteloua lophostachya Griseb.; Bouteloua nana 
Griseb. 


Neostapfia Burtt Davy, Erythea 7: 43. 1899. 
REPLACED NAME: Stapfia Burtt Davy. 
TRTD. BY: Peterson, P. M. 

NHT: Orcuttieae. NO. ACCPTD. SPP.: 1. 
CNSLTD. LIT.: Crampton, B., (1959). 
SYN.: Davyella Hack.; Stapfia Burtt Davy. 


Neostapfia colusana (Burtt Davy) Burtt Davy, Erythea 7: 43. 
1899. Basionym: Stapfia colusana Burtt Davy. 

DST.: USA. 

ALSO ACCPTD. BY: Hickman, J. C. (1993); Hitchcock, A. 
S. (1951); Munz, P. A. & D. D. Keck (1959); Reeder, 
J.R. (1982). 

Anthochloa colusana (Burtt Davy) Scribn.; Davyella 
colusana (Burtt Davy) Hack.; Stapfia colusana Burtt 
Davy. 


Nestlera Steud., Nomencl. Bot. (ed. 2) 2: 192. 1841. 
NOTE: nom nud., non Sprengel 1818. 
= Bouteloua Lag. 


SYN.: 


Nestlera festuciformis Willd. ex Steud., Nomencl. Bot. (ed. 2) 
2: 192. 1841. 
NOTE: nom. inval., as syn. of Eutriana bromoides Trin. 
= Bouteloua radicosa (E. Fourn.) Griffiths. 


Nevroctola Raf., Neogenyton 4. 1825. 
NOTE: nom. superfl. pro Uniola. 
= Uniola L. 


Nevroctola maritima Raf. ex B.D. Jacks., Index Kew. 2: 311. 
1894. 
NOTE: nom. inval., as syn. of Uniola paniculata L. 
= Uniola paniculata L. 


Nevroctola paniculata Raf. ex B.D. Jacks., Index Kew. 2: 
311. 1894. 
NOTE: nom. inval., as syn. of Uniola paniculata L. 
= Uniola paniculata L. 


176 


Catalogue of New World Grasses: II. 


Neyraudia Hook. f, Fl. Brit. India 7: 305. 1896. TYPE: 
Neyraudia madagascariensis (Kunth) Hook. f. 
TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 1. 
CNSLTD. LIT.: Conert, H. J., (1959). 


Neyraudia arundinacea var. zollingeri (Biise) Henrard, 
Blumea 3: 439. 1940. Basionym: Arundo zollingeri 
Biise. 

= Neyraudia reynaudiana (Kunth) Keng ex Hitchc. 


Neyraudia madagascariensis var. zollingeri (Biise) Hook. f., 
Fl. Brit. India 7: 305. 1897. Basionym: Arundo 
zollingeri Biise. 

= Neyraudia reynaudiana (Kunth) Keng ex Hitchc. 


Neyraudia reynaudiana (Kunth) Keng ex Hitche., Amer. J. 
Bot. 21(3): 131. 1934. Basionym:Arundo reynaudiana 
Kunth. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Bor, N. L. 
(1960); Conert, H. J. (1959); Espejo Serna, A., A. R. 
L6pez-Ferrari, and J, Valdes-Reyna (2000); Godfrey, 
R. K. & J. W. Wooten (1979); Hitchcock, A. S. 
(1951); Long, R. W. & O. Lakela (1971); Wunderlin, 
R. P. (1998), 

SYN.: Arundo reynaudiana Kunth; Arundo zollingeri Buse; 
Neyraudia arundinacea’ var. zollingeri  (Biise) 
Henrard; Neyraudia madagascariensis var.zollingeri 
(Biise) Hook. f.; Phragmites zollingeri (Biise) Steud. 


Opizia J. Presl, Relig. Haenk. 1(4-5): 293, t. 41, f. 1-11. 1830. 
TYPE: Opizia stolonifera J. Pres|. 
= Bouteloua Lag. 


Opizia bracteata McVaugh, Fl. Novo-Galiciana 14: 269-270, 
f. 16a-e. 1983. T-Protol.: Mexico: Michoacan: wooded 
hills, roadsides, grassy openings in tropical deciduous 
forest, along the road from Apatzingan to Aguililla, ca. 
16 mi S of the steel bridge over the Rio Tepalcatepec, 
elv. 1450 m, flowering, 14 Sep 1958,McVaugh 17818 
{HT: MICH]. 

= Bouteloua bracteata (McVaugh) Columbus. 


Opizia stolonifera J. Presl, Relig. Haenk. 1(4-5): 293, t. 41, f. 
I-11. 1830. T-Protol.: Hab. ad Acapulco, Haenke s.n. 
T-Specim.: 1 of 1. Mexico: Guerrero: Acapulco, 
Haenke s.n. (HT: PR; IT: US (fragm.)]. 

= Bouteloua dimorpha Columbus. 


Orcuttia Vasey, Bull. Torrey Bot. Club 13: 219. 1886. 
. TYPE: Orcuttia californica Vasey. 
TRTD. BY: Peterson, P. M. 
NHT: Orcuttieae. NO. ACCPTD. SPP.: 5. 


Orcuttia californica Vasey, Bull. Torrey Bot. Club 13: 219, 

pl. 60. 1886. T-Protol.: Mexico: Baja California: near 

San Quentin Bay, C.R. Orcutt s.n. T-Specim.: | of 1. 

Mexico: Baja California Norte: near San Quintin Bay, 

21 Apr 1886, C_R. Orcutt 1439 [HT: US-81384; IT: 

ARIZ, DS, MEXU, MO-1837553, UC]. 

DST.: Mexi, USA. 
LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Abrams, L. (1923); Beetle, A. A. 

(1977); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 

Valdes-Reyna (2000); Gould, F. W. & R. Moran 


(1981); Hickman, J. C. (1993); Hitchcock, A. S. 
(1951); Munz, P. A. (1974); Reeder, J. R. (1982). 
SYN.: Orcuttia californica var. californica. 


Orcuttia californica Vasey var.californica. 
= Orcuttia californica Vasey. 


Orcuttia californica var. inaequalis (Hoover) Hoover, Bull. 
Torrey Bot. Club 68(3): 154-155. 1941. Basionym: 
Orcuttia inaequalis Hoover. 

= Orcuttia inaequalis Hoover. 


Orcuttia californica var. viscida Hoover, Bull. Torrey Bot. 
Club 68(3): 155. 1941. T-Protol.: USA: California: 
Sacramento Co.: 7 mi S of Folsom, Hoover 3709 (HT: 
UC]. T-Specim.; 1 of 1. USA: California: Sacramento 
Co.: S of Folsom, 30 Jul 1938, Hoover 3709 [HT: UC; 
IT: DS, US-1789362]. 

= Orcuttia viscida (Hoover) Reeder. 


Orcuttia fragilis Swallen, J. Wash. Acad. Sci. 34(9): 308. 
1944. T-Protol.: Mexico: southern Baja California, on 
playa, sandy clay, at Llano Dirai, Magdalena Plain, 
within the limits of the Sonoran Desert, H. S. Gentry 
4192 [HT: US-1865489]. T-Specim.: | of 1. Mexico: 
Baja California Sur: Magdalena Plain, Llano Dirai, 
sandy clay, an abundant forage grass over the great 
flood plain following rain stoage; reported excellent 
for cattle, 7 Jan 1939, H.S. Gentry 4192 [HT: US- 
1865489; IT: ARIZ, DS, UC, US-2209357], Reeder, 
Madrofio 30: 308 (1981), reports the actual locality to 
be: Llanos de Hiray, a few miles south of El Refugio. 

= Tuctoria fragilis (Swallen) Reeder. 


Orcuttia greenei Vasey, Bot. Gaz. 16(5):; 146. 1891. T- 
Protol.: USA: California: collected on moist plains of 
the upper Sacramenton ear Chico, Jun 1890, ELL. 
Greene s.n. T-Specim.: | of 1. USA: California: Butte 
Co.: near Chico, Jun 1890, Greene s.n. [HT: US- 
81385]. 

NOTE: as Orcuttia "Greenei". 
= Tuctoria greenei (Vasey) Reeder. 


Orcuttia inaequalis Hoover, Madrofio 3: 229. 1936. T- 
Protol.: USA: California: Satnislaus Co.: Montpellier, 
28 May 1935, Hoover 582 [HT: UC-534128]. T- 
Specim.: 1 of 1. USA: California: Stanislaus Co.: 
Montpellier, 28 May 1935, R.F. Hoover 582 [HT: UC; 
IT: US-1645282]. 
DST.: USA. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Hickman, J. C. (1993); Reeder, J. R. 
(1982). 
Oreuttia californica var. 
Hoover. 


SYN.: inaequalis (Hoover) 


Orcuttia mucronata Crampton, Madrofio 15: 107, f. 2-3. 
1959. T-Protol.: USA: California: Solano Co.: 12 mi S$ 
of Dixon, | Aug 1958, Crampton 5057 [HT: AHUC]. 
T-Specim.: | of 1. USA: California: Solano Co.: 12 mi 
S of Dixon, | Aug 1958, B. Crampton 5077 [HT: 
AHUC; IT: CAS, DS, JEPS, UC, US-2241392]. 

= Tuctoria mucronata (Crampton) Reeder. 


Orcuttia pilosa Hoover, Bull. Torrey Bot. Club 68(3): 155- 
156. 1941. T-Protol.: USA: California: Stanislaus Co.: 
12 mi E of Waterford, Hoover 3624 [HT: UC]. T- 
Specim.: | of 1. USA: California: Stanislaus Co.: E of 


Catalogue of New World Grasses: II. 


177 


Waterford, 10 Jul 1938, Hoover 3624 [HT: UC; IT: 
DS, US-1789361, US-1815889]. 
DST.: USA, 
LVL. ACCPTC:.: I. 
ALSO ACCPTD. BY: Hickman, J. C. (1993); Hitchcock, A. 
S. (1951); Munz, P. A. & D. D. Keck (1959); Reeder, 
J.R. (1982). 


Orcuttia tenuis Hitchc., Amer. J. Bot. 21(3): 131. 1934. T- 
Protol.: USA: California: Shasta Co., collected in open 
sandy oil, Goose Valley, 29 Jun 1912, 4. Eastwood 
1013 = Amer. Grass. Natl. Herb. 686 [HT: US- 
734402]. T-Specim.: 1 of 1. USA: California: Shasta 
Co.: Goose Valley, 29 Jun 1912, 4. Eastwood 1013 
[HT: US-734402; IT: CAS, MO-848913, RM, UC]. = 
Amer. Grass Nat. Herb. 686. 

DST.: USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Hickman, J. C. (1993); Hitchcock, A. 
S. (1951); Munz, P. A. & D. D. Keck (1959); Reeder, 
J.R. (1982). 


Orcuttia viscida (Hoover) Reeder, Phytologia 47(3): 221. 
1980. Basionym: Orcuttia californica var. viscida 
Hoover. 

DST.: USA. 

LVL. ACCPTC:: L. 

ALSO ACCPTD. BY: Hickman, J. C. (1993); Reeder, J. R. 
(1982). 

SYN.: Orcuttia californica var. viscida Hoover. 


Orcuttieae trib. Reeder, Madrofio 18: 20. 1965. TYPE: 
Orcuttia Vasey. 
TRTD. BY: Soreng, R. J., G. Davidse, T. S. Filgueiras, E. J. 
Judziewicz, P. M. Peterson, and F. O. Zuloaga. 
NHT: Chloridoideae. 
CNSLTD. LIT.: Roalson, E. H. & J. T. Columbus, (1999); 
Reeder, J. R., (1982); Reeder, J. R., (1965). 


Osterdamia Neck. ex Kuntze, Revis. Gen. Pl. 2: 781. 1891. 
TYPE: Osterdamia matrella (L.) Kuntze. 
NOTE: nom. illeg. superfl. pro Zoysia. 
= “Zoysia Willd. 


Osterdamia japonica (Steud.) Hitche., U.S.D.A. Bull. (1915- 
23) 772: 166. 1920. Basionym: Zoysia japonica Steud. 
= Zoysia japonica Steud. 


Osterdamia matrella (L.) Kuntze, Revis. Gen. Pl. 2: 781. 
1891. Basionym: Agrostis matrella L. 
= Zoysia matrella (L.) Merr. 


Osterdamia tenuifolia (Willd. ex Thiele) Kuntze, Revis. Gen. 
Pl. 2: 781. 1891. Basionym: Zoysia tenuifolia Willd. 
ex Thiele. 

= Zoysia tenuifolia Willd. ex Thiele. 


Osterdamia zoysia Honda, Bot. Mag. (Tokyo) 36: 113. 1922. 
NOTE: nom. illeg. superfl. for Agrostis matrella L. 
= Zoysia matrella (L.) Merr. 


Osterdamia zoysia var. tenuifolia (Willd. ex Thiele) Honda, 
Bot. Mag. (Tokyo) 36: 113. 1922. 
= Zoysia matrella (L.) Merr. 


Osterdamia zoysia var. typica Honda, Bot. Mag. (Tokyo) 36: 
113. 1922. 
NOTE: nom. inval., see ICBN Art. 24.3. 


= Zoysia matrella (L.) Merr. 


Ottochloa malabarica (L.) Dandy, J. Bot. 69(2): 55. 1931. 
Basionym: Poa malabarica L. 
NOTE: nom. rej., basionym rej. [ICBN art. 56.2]. 
= Leptochloa fusca (L.) Kunth. 


Oxydenia Nutt., Gen. N. Amer. Pl. 1: 76. 1818. TYPE: 
Eleusine filiformis Pers., LT designated by Pfeiffer, 
Nomencl. Bot. 2: 554 (1873). 

= Leptochloa P. Beauv. 


Oxydenia filiformis Nutt. ex B.D. Jacks., Index Kew. 2: 392. 
1894. 
NOTE: nom. inval., as syn. of Leptochloa filiformis (Lam.) 
P. Beauv. 
= Leptochloa panicea subsp. brachiata (Steud.) N. Snow. 


Oxydenia virgata Nutt. ex B.D. Jacks., Index Kew. 2: 392. 
1894. 
NOTE: nom. inval., as syn. of Leptochloa virgata (L.) P. 
Beauv. 
= Leptochloa virgata (L.) P. Beauv. 


Pallasia Scop., Intr. Hist. Nat. 72. 1777. ; not designated. 
NOTE: hom. illeg. 
= Crypsis Aiton. 


Panicularia brevifolia (Muhl.) Porter, Bull. Torrey Bot. Club 
20: 205. 1893. Basionym: Festuca brevifolia Muhl. 
= Triplasis purpurea (Walter) Chapm. 


Panicum dactylon L., Sp. Pl. 1: 58. 1753. T-Protol.: In europa 
australi. T-Specim.: | of 1. Portugal: Baseado no Pinax 
de Bauhin, Anon. [LT: LINN-80.35], LT designated by 
Clayton & Harlan, Kew Bull. 24: 185-189 (1970). 

= Cynodon dactylon (L.) Pers. 


Panicum festucoides Poir., Encycl., Suppl. 4: 283. 1816. T- 
Protol.: "Cette plante croit dans les Indes orientales.", 
Vs. in herb. Desv. 
= Tridens flavus (L.) Hitchce. 


Panicum glumaepatulum Steud., Syn. Pl. Glumac. 1: 41. 
1853. T-Protol.: Ins. Philipp., Herb. Cuming 550. T- 
Specim.: | of 1. Philippines: Luzon, Cuming 550 [HT: 
P: IT: L, MO]. 

= Cynodon dactylon (L.) Pers. 


Panicum malabaricum (L.) Merr., Philipp. J. Sci. 4: 248. 
1909. Basionym: Poa malabarica L. 
NOTE: nom. rej., basionym rej. [ICBN art. 56.2]. 
= Leptochloa fusca (L.) Kunth. 


Panicum microspermum (Lag.) E. Fourn., Mexic. Pl. 2: 492. 
1886. Basionym: Miliwm microspermum Lag. 
= Muhlenbergia minutissima (Steud.) Swallen. 


Panicum tenuiflorum Schrank, Syll. Pl. Nov. 1: 192. 1824. T- 
Protol.: "Patria: Habitat in Brasilia inter Tejuco et 
Bahiam, ubi D. de Martius semina collegit.". T- 
Specim.: 1 of 2. [see next record]. 2 of 2. cultivated in 
Regensburg from seeds collected in Brazil, Martius 
s.n. [HT: M]. 

NOTE: hom. illeg. 
E. NAME: Panicum tenuiflorum R. Br. 
= Sporobolus tenuissimus (Mart. ex Schrank) Kuntze. 


Panicum tenuissimum Mart. ex Schrank, Denkschr. Konigl.- 
Baier. Bot. Ges. Regensburg 2: 26. 1822. T-Specim.: 1 
of 1. Cultivated in Munich fron seeds collected in 


178 


Catalogue of New World Grasses: II. 


Brazil, Martius s.n. [HT: M], merotypic with Vilfa 
minutiflora Trin. 
NOTE: type grown from same seed source as Vilfa 
minutiflora Trin, 
= Sporobolus tenuissimus (Mart. ex Schrank) Kuntze. 


Pappophoreae trib. Kunth, Revis. Gramin. 1: 82. 
1829. TYPE: Pappophorum Schreb. 
TRTD. BY: Soreng, R. J., G. Davidse, T. S. Filgueiras, E. J. 
Judziewicz, P. M. Peterson, and F. O. Zuloaga. 
NHT: Chloridoideae. 
CNSLTD. LIT.: Reeder, J. R., (1965). 


Pappophorinae subtrib. Dumort., Anal. Fam. Pl. 63. 1829. 


TYPE: Pappophorum Schreb. 
NOTE: as "Pappophoreae". 


Pappophoroideae subfam. Burmeist., Handb. Naturgesch. 


205. 1837. TYPE: Pappophorum Schreb. 
NOTE: as "Pappophorae". 


Pappophorum Schreb., Gen. Pl. 2: 787. 1791, 
TYPE: Pappophorum alopecuroideum Vahl. 
TRTD. BY: Peterson, P. M. 
NHT: Pappophoreae. NO. ACCPTD. SPP.: 8. 
CNSLTD. LIT.: Reeder, J. R. & L. J. Toolin, (1989). 
SYN.: Pappophorum sect. Polyraphis Trin.; Polyraphis 
(Trin.) Lindl. 


Pappophorum alopecuroideum Vahl, Symb. Bot. 3: 10, t. 51. 


1794. T-Protol.: Ad fodinas Insulae Spanish Town 
Americae, von Rohr s.n., may refer to Spanish Town, 
Jamaica, but we have no specimen from that islland. 
T-Specim.: | of 2. Jamaica: Spanish Town, von Rohr 
sn. [HT]. 2 of 2. Jamaica: Spanish Town, von Rohr 
sn. (HT: MA?, BAA(fragm.)]. 

= Pappophorum pappiferum (Lam.) Kuntze. 


Pappophorum alopecuroideum fo. coerulescens  Stuck.. 
Anales Mus. Nac. Buenos Aires 11: 122. 1904. T- 
Protol.: Argentina, Stuckert [Hrb. arg. no 9167a] (HT: 
Stucker Hrb.]. 

DST.: Arge. 
= Status not yet determined. 


Pappophorum alopecuroideum var. glabrum Doll, Fl. Bras. 
2(3): 59. 1878. T-Protol.: Brazil, Gray cards give as: 
[var. ]. 

= Pappophorum pappiferum (Lam.) Kuntze. 


Pappophorum alopecuroideum var. pilosiusculum Doll, FI. 
Bras, 2(3): 59. 1878. T-Protol.: Brazil: in sabulosis 
maritimis prope Bahia, Salzmann 713 [IT: K], Gray 
cards give as: [var.]. 

= Pappophorum pappiferum (Lam.) Kuntze. 


Pappophorum apertum Munro ex Scribn., Bull. Torrey Bot. 
Club 9: 148. 1882. T-Protol.; USA: Arizona: mesas 
near Camp Lowell, Pringle s.n. [HT: US]. T-Specim.: 
1 of 1. USA: Arizona, 1881, C.G. Pringle s.n. {IT: 
MO, US-899491]. 

= Pappophorum vaginatum Buckley. 


Pappophorum apertum var. vaginatum (Buckley) Scribn. ex 
L.H. Dewey, Contr. U.S. Natl. Herb. 2(3); 535. 1894. 
Basionym: Pappophorum vaginatum Buckley. 

NOTE: based on Pappophorum vaginatum Buckl. 


= Pappophorum vaginatum Buckley. 


Pappophorum bicolor E. Fourn. ex Hemsl., Biol. Cent.- 
Amer., Bot. 3: 568. 1885. 
NOTE: nom. nud. 
CORRECT NAME: Pappophorum bicolor E. Fourn. 
= Pappophorum bicolor E. Fourn. 


Pappophorum bicolor E. Fourn., Mexic. Pl. 2: 133. 1886. T- 
Protol.: Mexico: Cerro de las Minas et Toluca, 
Karwinsky 1483 [IT: US-899499]. T-Specim.: | of 1. 
Mexico, Karwinsky 1483 |IT: US-899499 (fragm. ex 
LE)]. 

DST.: Mexi, USA. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Hitchcock, A. S. (1951); Reeder, J. R. 
& L. J. Toolin (1989). 

SYN.: Pappophorum bicolor E. Fourn. ex Hemsl. 


Ross. 4: 404. 1852. T- 
baikalensis rupibus as 


Pappophorum boreale Griseb., FI. 
Protol.: Hab. in Sibirae 
Selengam, 7urcz. 

= Enneapogon desvauxii P. Beauv. 


Pappophorum boreale C.C. Gmel. ex B. Fedtsch., Izv. Imp. 
Bot. Sada Petra Velikago 14(Suppl. 2): 67. 1915. 
NOTE: without description, nom. nud. or hom. illeg., cited 
Czernaevia rupestris Turcez., P. phleoides Turcz., and 
Raspalia rupestris Turcz. 

E. NAME: Pappophorum boreale Griseb. 

= Enneapogon desvauxii P. Beauv. 


Pappophorum brachystachyum Jaub. & Spach, Ann. Sci. 
Nat., Bot., ser. 3, 14: 365. 1850. T-Protol.: Arabia 
Felix, Taifa, Botta s.n. T-Specim.: 1 of 1. Arabia Felix, 
Taifa, Botta [HT: P]. 

= Enneapogon desvauxii P. Beauv. 


Pappophorum brachystachyum Jaub. & Spach, Ill. Pl. Orient. 
4: 34, t. 324. 1850-1853 [1851]. T-Protol.: Arabia 
Felix, Taifa, Botta s.n. [HT: P]. 

NOTE: isonym. 
= Enneapogon desvauxii P. Beauv. 


Pappophorum caespitosum R.E. Fr., Nova Acta Regiae 
Soc. Sci. Upsal. 1: 177, t. 9, f. 5-6. 1905. T-Protol.: 
Argentina: Prov. Jujuy: Huancar ad Salinas grandes in 
fissuris rupium, 20 Nov 1901, Fries 804. T-Specim.: | 
of |. Argentina: Jujuy, 20 Nov 1901. RE. Fries 804 
[1T: US-1162325]. 

DST.: Arge, Boli, Para. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Nicora, E. G., M. E. De Paula, A. M. 
Paggi, M. del C. Mariano, A. M. Miante A., L. R. 
Parodi, C. A. Petetin, F. A. Roig & Z. Rugolo de 
Agrasar (1978); Reeder, J. R. & L. J. Toolin (1989); 
Renvoize, S. A. (1998): Zuloaga, F. O., E. G. Nicora, 
Z. E. Rugolo de Agrasar, O. Morrone, J. Pensiero & 
A.M. Cialdella (1994). 


Pappophorum elongatum Spreng., Syst. Veg. 4(2): 34. 1827. 
T-Protol.: Uruguay: Montevideo, Sed/o s.n. 
= Pappophorum pappiferum (Lam.) Kuntze. 


Pappophorum eutrianoides Trin. ex Nees, Fl. Bras. Enum. PI. 
2: 414. 1829. 
NOTE: nom. inval., as syn. of Eutriana multiseta Nees. 


Catalogue of New World Grasses: II. 


179 


= Bouteloua megapotamica (Spreng.) Kuntze. 


Pappophorum fasciculatum Chioyv., Pl. Nov. Minus Not. 
Aethiopia 31. 1928. T-Specim.: 1 of 1. Somali 
Republic, Puccioni & Stefanini 506 (ST: FI]. 

= Enneapogon desvauxii P. Beauv. 


Pappophorum figarianum Fig. & De Not., Mem. Reale 
Accad. Sci. Torino, ser. 2, 12: 254. 1852. T-Specim.: 1 
of 1. Sudan, Figari s.n.[T]. 
= Enneapogon desvauxii P. Beauv. 


Pappophorum hassleri Hack., Bull. Herb. Boissier, ser. 2, 
4(3): 280. 1904. T-Protol.: Paraguay: in arenosis prope 
flumen Capibary, sep, Hassler 4456. T-Specim.: | of 
|. Paraguay: fl. Capibary, Hassler 4456 [IT: US-76118 
(fragm. ex Delessert)]. 

NOTE: as Pappophorum "Hassleri". 

DST.: Para. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Reeder, J. R. & L. J. Toolin (1989). 


Pappophorum jaminianum Coss. & T. Durand & Boiss., FI. 
Orient. 5: 558. 1884. 
NOTE: nom. inval., as syn. 
= Enneapogon desvauxii P. Beauv. 


Pappophorum krapovickasii Roseng., Comun. Bot. Mus. 
Hist. Nat. Montevideo 4(58): 1, text f. 1975. T-Protol.: 
Paraguay: Chaco: Mariscal Estigarribia, 22.00S 
60.42W, en claros de monte, 24 Jan 1950, Rosengurtt 
B-5816 [HT: MVFA\. 

DST.: Arge, Boli, Braz, Para. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 
(1987); Killeen, T. J. (1990); Reeder, J. R. & L. J. 
Toolin (1989); Renvoize, S. A. (1998); Zuloaga, F. 
O., E. G. Nicora, Z. E. Rugolo de Agrasar, O. 
Morrone, J. Pensiero & A. M. Cialdella (1994). 


Pappophorum laguroideum Schrad. ex Schult., Mant. 2: 342. 
1824. T-Protol.: In India occidentali. T-Specim.: 1 of 
|. {[IT: B, BAA-2007 (fragm. ex B)]. 
= Pappophorum pappiferum (Lam.) Kuntze. 


Pappophorum macrospermum Roseng., B.R. Arill. & Izag., 
Gram. Urug. 397, f. 178. 1970. T-Protol.: Uruguay: 
Rivera: ruta 30 entre Bajada de Pena y A. Rubio 
Chico, 7 Feb 1966, Del Puerto & Marchesi 6080 [HT: 
MFVA\]. 

= Pappophorum philippianum Parodi. 


Pappophorum macrostachyum Schrad. ex Schult., Mant. 2: 
342. 1824. T-Protol.: In Brasilia, Princeps Sereniss. 
Maximilian Neovidens s.n. T-Specim.: | of 1. Brazil: 
Barra de Vereda, Maximilian s.n. [HT: B, IT: BAA- 
2008 (fragm. ex B), K, US-76120 (fragm. ex B)]. 

= Pappophorum pappiferum (Lam.) Kuntze. 


Pappophorum megapotamicum Spreng., Syst. Veg. 4(2): 34. 
1827. T-Protol.: Brasilia: rio Grande, Sello s.n. T- 
Specim.: | of |. Brasil: Rio Grande, Se/lo s.n. [HT: 
B?; US-72694 (fragm. ex B, herb. Sprengel)]. 

= Bouteloua megapotamica (Spreng.) Kuntze. 


Pappophorum mexicanum Griseb. ex E. Fourn., Mexic. Pl. 2: 
133. 1886. T-Specim.: 1 of 2. Mexico: Distrito 
Federal: Guadalupe, Aug, Bourgeau s.n. [ST]. 2 of 2. 
Mexico: Distrito Federal: in valle Mexicensi, 
Schaffner 184 (ST: US-76123 (fragm. ex W)]. 


= Enneapogon desvauxii P. Beauv, 


Pappophorum molle (Lehm.) Kunth, Enum. Pl. 1: 255. 1833. 
Basionym: Enneapogon mollis Lehm. 
= Enneapogon mollis Lehm. 


Pappophorum mucronulatum Nees, FI. Bras. Enum. Pl. 2: 
412. 1829. T-Protol.: Habitat in capis interioribus 
provinciae Bahiensis, et campis Mimoso — dictis, 
provinciae Piauhiensis, Martius s.n.. syntypes. T- 
Specim.: | of 2. Brazil, 1834, Martius s.n. (IST: LE- 
TRIN-2199.01, M]. 2 of 2. Brazil: Bahia, Martius s.n. 
[ST: B; IST: BAA-2012 (fragm. ex B), US-76117 
(fragm. ex M)]. 

DST.: Braz, Colo, Hond, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Burkart, A. (1969); Cabrera, A. L. 
(1970): Correll, D. S. & M. C. Johnston (1970); 
Davidse, G. (1994); Davidse, G., M. Sousa S. & A. 
QO. Chater (editores generales) (1994); Dll, J. C. 
(1878); Filgueiras, T. S. (1999); Foster, R. C. (1958): 
Hitchcock, A. S. (1927); Hitchcock, A. S. (1951): 
Reeder, J. R. & L. J. Toolin (1989); Renvoize, S. A. 
(1984): Shreve, F. & I. L. Wiggins (1964); Tovar, O. 
(1993). 

Pappophorum pappiferum var.mucronulatum (Nees) 
Kuntze. 


SYN.: 


Pappophorum mucronulatum fo.elatior Stuck., Anales Mus. 
Nac. Buenos Aires 11: 123. 1904. T-Protol.: 
Argentina: el los immediasciones de Cordoba: 27 Jan 
1903 [first cited Index of Grass Species], Stuckert 
[herb. arg. 12682], syntypes. 

= Pappophorum vaginatum Buckley. 


Pappophorum mucronulatum fo. intermedia Hack., Anales 
Mus. Nac. Hist. Nat. Buenos Aires 21: 123. 1911. T- 
Protol.: Argentina: Rioja: La Diana, 12 Mar 1907, 
Stuckert 17259. 

= Pappophorum philippianum Parodi. 


Pappophorum mucronulatum var. subsimplex Hack. ex 
Stuck., Anales Mus. Nac. Hist. Nat. Buenos Aires 13: 
495. 1906. T-Protol.: Argentina: Cordoba, Stuckert 
herb. arg. 3853. ; 
= Pappophorum philippianum Parodi. 


Pappophorum mucronulatum var. vaginatum (Phil.) Hack., 
Anales Mus. Nac. Hist. Nat. Buenos Aires 21: 123. 
1911. Basionym: Pappophorum vaginatum Phil. 
= Pappophorum philippianum Parodi. 


Pappophorum nanum Steud., Syn. Pl. Glumac. 1: 200. 1854. 
T-Protol.: Africa: Cape Verde Isls., Steudel cites "P. 
vincentianum J.A. Schmidt.Ins. St. Vincent.". 

NOTE: nom. illeg. superfl., 

CORRECT NAME: Pappophorum 
Schmidt. 

= Enneapogon desvauxii P. Beauv. 


vincentianum — J.A. 


Pappophorum pappiferum (Lam.) Kuntze, Revis. Gen. PI. 
3(3): 365. 1898. Basionym: Saccharum pappiferum 
Lam. 

DST.: Arge, Boli, Braz, Cari, Ecua, Guya, Mexi, Para, Peru, 

USA, Vene. 

LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 
(1987); Anonymous (1986); Beetle, A. A. (1977); 
Brako, L. & J. L. Zarucchi (1993); Burkart, A. 


180 


Catalogue of New World Grasses: II. 


(1969); Catasus Guerra, L. (1997); Espejo Serna, A., 
A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Gould, F. W (1979); Hitchcock, A. S. (1936); 
Jorgensen, P. M. & C. Ulloa U. (1994); Jorgensen, P. 
M. & S. Leén-Yanez (eds.) (1999); Judziewicz, E. J. 
(1990); Killeen, T. J. (1990); Lindeman, J.C. & A.L. 
Stoffers (1963); Liogier, H. & L. Martorell (1982); 
Reeder, J. R. & L. J. Toolin (1989); Renvoize, S. A. 
(1984); Renvoize, S. A. (1998); Tovar, O. (1993); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

Pappophorum alopecuroideum Vahl; Pappophorum 
alopecuroideum var. glabrum Doll; Pappophorum 
alopecuroideum var. pilosiusculum Doll; 
Pappophorum elongatum Spreng.; Pappophorum 
laguroideum Schrad. ex Schult.; Pappophorum 
macrostachyum Schrad. ex Schult.; Pappophorum 
pappiferum var. alopecurodes (Vahl) Kuntze; 
Pappophorum pappiferum var.laguroideum (Schrad. 
ex Schult.) Kuntze; Pappophorum  polystachyum 
Kunth; = Pappophorum — saccharoides — Griseb.; 
Saccharum pappiferum Lam. 


SYN.: 


Pappophorum pappiferum var. alopecurodes (Vahl) Kuntze, 
Revis. Gen. Pl. 3(3): 365. 1898.  Basionym: 
Pappophorum alopecuroideum Vahl. 

= Pappophorum pappiferum (Lam.) Kuntze. 


Pappophorum pappiferum var. laguroideum (Schrad. ex 
Schult.) Kuntze, Revis. Gen. Pl. 3(3): 365. 1898. 
Basionym: Pappophorum laguroideum Schrad. ex 
Schult. 

NOTE: as Pappophorum laguroideum. 

= Pappophorum pappiferum (Lam.) Kuntze. 


Pappophorum pappiferum var.mucronulatum (Nees) Kuntze, 
Revis. Gen. Pl. 3(3): 365. 1898. Basionym: 
Pappophorum mucronulatum Nees. 

= Pappophorum mucronulatum Nees. 


Pappophorum philippianum Parodi, Notas Mus. La Plata, 
Bot. 8(40): 79. 1943. 

REPLACED NAME: Pappophorum vaginatum Phil. 

BLOCKING NAME: Pappophorum vaginatum Buckley. 

DST.: Arge, Boli, Chil, Mexi, Urug. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 

Ferrari, and J, Valdes-Reyna (2000); Filgueiras, T. S. 
(1999); Nicora, E. G., M. E. De Paula, A. M. Faggi, 
M. del C. Mariano, A. M. Miante A., L. R. Parodi, C. 
A. Petetin, F. A. Roig & Z. Rugolo de Agrasar 
(1978); Reeder, J. R. & L. J. Toolin (1988); Reeder, 
JR. & L. J. Toolin (1989); Renvoize, S. A. (1998); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 
Pappophorum macrospermum Roseng., B.R. Arill. & 
Izag.; Pappophorum mucronulatum fo. intermedia 
Hack.; Pappophorum mucronulatum var. subsimplex 
Hack. ex Stuck.; Pappophorum mucronulatum var. 
vaginatum (Phil.) Hack.; Pappophorum vaginatum 
Phil. 


SYN.: 


Pappophorum phleoides Turcz., Bull. Soc. Imp. Naturalistes 
Moscou 29(1): 30. 1856. 
NOTE: nom. inval., as syn. of Pappophorum boreale 
Griseb. 


= Enneapogon desvauxii P. Beauv. 


Pappophorum polystachyum Kunth, Revis. Gramin. 2: 435, t. 
131. 1831. T-Protol.: Uruguay: Crescit prope 
Montevideo. 

= Pappophorum pappiferum (Lam.) Kuntze. 


Pappophorum pusillum (Rendle) K. Schum., Just's Bot. 
Jahresber. 27: 458. 1901. Basionym: Enneapogon 
pusillus Rendle. 

= Enneapogon desvauxii P. Beauv. 


Pappophorum saccharoides Griseb., Abh. Konig]. Ges. Wiss. 
Gottingen 24: 301. 1879. T-Protol.: Paraguay: C.: in 
rupibus pr. S. Maria, Balansa 288a. T-Specim.: | of 2. 
Paraguay, Balansa 288a |ST]. 2 of 2. Argentina: 
Cordoba: an einem Felsen zwischen Cosquin und 
Santa Maria, 27 Jan 1876,G. Hieronymus 281 |ST: B; 
IST: BAA-2017 (fragm. ex B), US-76116 (fragm. ex 
Griseb. herb.)]. 

= Pappophorum pappiferum (Lam.) Kuntze. 


Pappophorum subbulbosum Arechav., Anales Mus. Nac. 
Montevideo 1: 405, t. 48. 1896. T-Protol.: Uruguay: 
Montevideo, Punta Brava, terrenos arenosis, Paso de 
los Toros, campos secos, aridos, Mercedes campos 
arensoso de las orillas del Rio Negro. T-Specim.: | of 
1. Uruguay: Montevideo, no date,J/. Arechavaleta s.n. 
[IT: US-899538]. 

= Pappophorum vaginatum Buckley. 


Pappophorum vaginatum Buckley, Prelim. Rep. Surv. 
Texas App. 1. 1866. T-Protol.: USA: Texas, western 
Texas. T-Specim.: 1 of 1. USA: Texas, May 1849-Oct 
1849, C. Wright 803 (IT: US-76124]. 

DST.: Arge, Mexi, Para, USA, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. & R. Moran 
(1981); Reeder, J. R. & L. J. Toolin (1989); Zuloaga, 
F. O., E. G. Nicora, Z. E. Rugolo de Agrasar, O. 
Morrone, J. Pensiero & A. M. Cialdella (1994). 

Pappophorum  apertum Munro” ex — Scribn.; 

Pappophorum apertum var. vaginatum (Buckley) 

Scribn. ex L.H. Dewey; Pappophorum mucronulatum 

fo. elatior Stuck.; Pappophorum  subbulbosum 

Arechavy. 


SYN:: 


Pappophorum vaginatum Phil., Anales Univ. Chile 36: 206. 

1870. T-Protol.: Argentina: Mendoza. T-Specim.: | of 
1. Mendoza, Anon. [HT: SGO; IT: B, BAA-2025 
(fragm. ex B), US-1939964 (fragm. ex SGO)]. 

NOTE: hom. illeg. 

E. NAME: Pappophorum vaginatum Buckley. 

CORRECT NAME: Pappophorum philippianum Parodi. 

= Pappophorum philippianum Parodi. 


Pappophorum vincentianum J.A. Schmidt, Beitr. Fl. Cap 
Verd. Ins. 144. 1852. T-Specim.: | of |. Cape Verde 
Islands, Bolles s.n. {IT: K]. 

= Enneapogon desvauxii P. Beauv. 


Pappophorum wrightii S. Watson, Proc. Amer. Acad. Arts 
18: 178. 1883. T-Specim.: | of 3. Mexico: Coahuila: 
Parras, 1880, E. Palmer 136! [ST: GH]. 2 of 3. USA: 
Texas: Devils River, 1851-1852,C. Wright 2029 [ST: 
US-76126]. 3 of 3. USA: Texas: Llano Estacado, in 
deep ravines, C. Wright 751 [ST]. 


Catalogue of New World Grasses: II. 181 


= Enneapogon desvauxii P. Beauv. 


Pappophorum sect. Polyraphis Trin., Mem. Acad. Imp. Sci. 
St.-Petersbourg, Ser. 6, Sci. Math. 1(1): 91. 1830. 
TYPE: Pappophorum alopecuroideum Vahl. 

= Pappophorum Schreb. 


Paspalum cynosuroides (L.) Brot., Fl. Lusit. 1: 83. 1804. 
Basionym: Dactylis cynosuroides L. 
= Spartina cynosuroides (L.) Roth. 


Paspalum dactylon (L.) Lam., Tabl. Encycl. 1: 176. 1791. 
Basionym: Panicum dactylon L. 
= Cynodon dactylon (L.) Pers. 


Paspalum superbum Spreng., Syst. Veg. 1: 248. 1825. T- 
Protol.: Brasil. 
= Eustachys distichophylla (Lag.) Nees. 


Paspalum tenuissimum M.E. Jones, Contr. W. Bot. 18: 24. 
1935. T-Protol.: On the prairie at the Laguna Mts., 
Lower California, lower temperate life zone, 6500 ft, 
and along the down trail, 22 Sep 1930, ME. Jones 
27584. T-Specim.: | of 1. Mexico: Baja California, 24 
Sep 1930, MLE. Jones 27584 [HT: POM, IT: US- 
1614906]. 

= Microchloa kunthii Desv. 


Paspalum umbellatum Lam., Tabl. Encycl. |: 177. 1791. T- 
Protol.: Insula Franciae, Commerson s.n. T-Specim.: | 
of 1. Mauritius, no date, P. Commerson s.n. (HT: P; 

IT: US-2856012 (fragm.)]. 

= Cynodon dactylon (L.) Pers. 


Pechea Lapeyr., Suppl. Pl. Pyr. 8. 1818. 
NOTE: in syn. sub Crypsis schoenoides (L.) Lam. 
= Crypsis Aiton. 


Pentarrhaphis Kunth, Nov. Gen. Sp. 1: 177. 1815 [1816]. 
TYPE: Pentarrhaphis scabra Kunth. 


Pentarrhaphis annua Swallen, Ceiba 4(5): 286. 1955. T- 
Protol.; Honduras: El Paraiso: road to Yuscaran, El 
Limonal, on bare rocks, 18 Oct 1951, / R. Swallen 
10861 {[HT: US-2076680]. T-Specim.: 1 of 1. 
Honduras: El Paraiso, 18 Oct 1951, JR. Swallen 
10861 (HT: US-2076680]. 

= Bouteloua swallenii Columbus. 


Pentarrhaphis polymorpha (E. Fourn.) Griffiths, Contr. U.S. 
Natl. Herb. 14(3): 357, f. 21. 1912. Basionym: 
Atheropogon polymorphus E. Fourn. 

= Bouteloua polymorpha (E. Fourn.) Columbus. 


Pentarrhaphis scabra Kunth, Nov. Gen. Sp. 1: 178, t 60. 
1815 [1816]. T-Protol.: Crescit in planatie montana 
Mexicana prope Tula et Queretaro, alt. 1000 hex., 
Humboldt & Bonpland s.n. [HT: P]. 

= Bouteloua scabra (Kunth) Columbus. 


Pereilema J. Presi, Relig. Haenk. 1(4-5): 233. 1830. 
TYPE: Pereilema crinitum J. Pres. 
TRTD. BY: Peterson, P. M. 
NHT: Muhlenbergiinae. NO. ACCPTD. SPP.: 4. 


Pereilema beyrichianum (Kunth) Hitche., Contr. U.S. Natl. 
Herb. 24(8) 385. 1927. Basionym: Muhlenbergia 
beyrichianum Kunth. 

DST.: Braz, Colo, Ecua, Guat, Mexi, Peru, Vene. 


LVL. ACCPTC:: |. 
ALSO ACCPTD. BY: Beetle, A. A. (1977); Brako, L. & J. 
L. Zarucchi (1993); Breedlove, D. E. (1986); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. L6opez- 
Ferrari, and J. Valdes-Reyna (2000); Filgueiras, T. S. 
(1991); Hitchcock, A. S. (1927); Jorgensen, P. M. & 
S. Leén-Yanez (eds.) (1999); McClure, F. A. (1955); 
Miller, C., G. K. Miller & P. Gutte (1981); Peterson, 
P. M. (2000); Pohl, R. W. (1980); Tovar, O. (1986): 
Tovar, O. (1993). 
Muhlenbergia beyrichianum 
brasilianum Trin. 


SYN:: Kunth; Pereilema 


Pereilema brasilianum Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(3-4): 382. 1841. T-Protol.: Brazil. T-Specim.: | of 
2. Brazil: in nemoribus Serr. d'os Orgos, Beyrich. s.n. 
(ST: IST: US-0090604 (fragm. ex LE)], OM: LE- 
TRIN 1520.01, 1520.02. 2 of 2. Brazil: in locis cultis 
prope Summodorio, Riedel. s.n. [ST]. 

= Pereilema beyrichianum (Kunth) Hitchce. 


Pereilema ciliatum E. Fourn., Mexic. PI. 2: 93. 1886. T- 
Specim.: | of 6. Mexico: Orizaba, Nov, Bourg. 3328 
[ST: P; IST: GH, K, L, US-996083 (fragm. ex LE)]. 2 
of 6. Mexico: Orizaba, Nov, fF. Mull. 2073 [ST: L, 
IST: K, NY, US-996084 (fragm. in Herb. Col. 
College), US-996083 (fragm.)]. 3 of 6. Mexico: Valle 
de Orizaba, Nov, Schaffn. 155 [ST: P; Herb. Franq. [P 
& PC]]. 4 of 6. Mexico: Cerro del Borreyo, Nov, Bott. 
87 [ST: P]. 5 of 6. Mexico: Orizaba, Nov, Bott. 40 
[ST: P; herb. Frang. [P & PC]; IST: US-0090799 
(fragm. ex Pitt & Dur.), US-996083 (fragm. ex P)]. 6 
of 6. Mexico: Cordova, Jul, Schaffn. 274 (ST: P; herb. 
Frang. [P & PC]; IST: US-090800 (fragm. ex Pitt & 
Dur)]. 

DST.: ElSa, Guat, Mexi. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Berendsohn, 
W. G. & A. E. Araniva de Gonzalez (1991); 
Breedlove, D. E. (1986); Davidse, G., M. Sousa S. & 
A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); hMcVaugh, R. (1983); Peterson, P. M. (2000); 
Pohl, R. W. (1994). 


Pereilema crinitum J. Presl, Relig. Haenk. 1(4-5): 233, t. 37, 
f. a-f. 1830. T-Protol.; Hab. in Panama, Haenke s.n. T- 
Specim.: | of 1. Panama, Haenke s.n. [HT: PR- 
198058; IT: PR-849, LE-TRIN-1519.01 (& fig.), MO- 
3047510, US-0090603 (fragm.)]. 

DST.: Braz, Colo, CoRi, Ecua, ElSa, Guat, Hond, Mexi, 
Pana, Vene. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Berendsohn, 
W. G. & A. E. Araniva de Gonzalez (1991); 
Breedlove, D. E. (1986); Davidse, G., M. Sousa S. & 
A. O. Chater (editores generales) (1994); Espejo 
Serna, A., A. R. Lopez-Ferrari, and J. Valdes-Reyna 
(2000); Gould, F. W. & R. Moran (1981); Hitchcock, 
A. S. (1927); Jorgensen, P. M. & S. Leon-Yanez 
(eds.) (1999); McClure, F. A. (1955); McVaugh, R. 
(1983); Molina R., A. (1975); Peterson, P. M. (2000); 
Pohl, R. W. (1980); Pohl, R. W. (1994); Reyes- 
Garcia, A. & M. Sousa S. (1997); Shreve, F. & 1. L. 
Wiggins (1964). 


182 


Catalogue of New World Grasses: II. 


cirratum E. Fourn.; 
crinitum, Pereilema 


Pereilema crinitum var. 
Pereilema  crinitum — var. 
setarioides E. Fourn. 


SYN.: 


Pereilema crinitum var. cirratum E. Fourn., Mexic. Pl. 2: 93. 
1886. T-Specim.: | of Il. Mexico: Veracruz: 
Escamella, Orizaba, 24-Oct.-1865-66, M4 Bourgeau 
3272 [ST: P; IST: K, L, LE, GH, US-996097, US- 
996091 (fragm. ex LE)], ex Mus. Paris. 2 of 11. 
[America]: Mexico: prope Cordova, W. Schaffner 275 
[ST: LE; IST: US-0090793 (fragm. ex Pitt.& Dur.)], 
W. Schaffner pl. Mexic. 3 of 11. Mexico: Orizaba, 
October, F. Mill. 2068 [ST: IST: K, NY, US-996088 
(fragm. ex LE), US-1127006 (fragm. ex W)]. 4 of 11, 
Mexico: Orizaba, October, fF. Mill. 2072 [ST: IST: 
US-996087 (fragm. ex LE & fragm. ex W)]. 5 of 11. 
Mexico: Orizaba, October, Thomas & Webber s.n. in 
herb. Buchinger {ST]|. 6 of 11. Mexico: Orizaba, 
October, Bott. 8/ [ST: P, IST: US-0090794 (fragm.)]. 
7 of 11. Mexico: Orizaba, October, Bott, & Sum, 33 
[ST: P; IST: US-0090796 (fragm. ex herb. Pitt & 
Dur)]. 8 of 11. Mexico: Orizaba, October, Bott. & 
Sum. 87 [ST]. 9 of Ll. Mexico: Orizaba, October, Bott, 
& Sum. 1272 [ST]. 10 of Ll. Mexico: Orizaba, 
October, Bott. & Sum. 1482 [ST]. 11 of 11. Mexico: 
Mirador, Hacienda del Sefior Posado, Schaffn. 142 pl. 
ed. Hohen. [ST: IST: US-0090797 (fragm. ex Herb. 
Boissier)]. 

= Pereilema crinitum J. Presl. 


Pereilema crinitum J. Pres| var. crinitum. 
= Pereilema crinitum J. Pres. 


Pereilema diandrum R.W. Pohl, Novon 2(2): 102. 1992. T- 
Protol.: Costa Rica: Heredia: Puente Mulas, S of San 
Antonio, canyon of Rio Virilla, 850 m, 28 Nov 1968, 
RW. Pohl & G. Davidse 11482 (HT: ISC; IT: MO 
(fragm. ex ISC), US]. 

DST.: CoRi. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Davidse, G. & R. W. Pohl (1992); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994): Peterson, P. M. (2000); Pohl, R. 
W. (1994). 


Pereilema setarioides E. Fourn., Mexic. Pl. 2: 93. 1886. T- 
Protol.: Mexico: Orizaba, Thomas s.n. [T: herb. 
Buchinger]. T-Specim.: | of 1. [IT: P, K, US-996083 
(fragm. ex P)]. 

= Pereilema crinitum J. Presl. 


Perotideae trib. C.E. Hubb., Grass. Burma, Ceylon, India & 
Pakistan 686. 1960. TYPE: Perotis W. Aiton. 


Phacellaria panicea Willd. ex Steud., Nomencl. Bot. (ed. 2) 
2: 313. 1841. 
NOTE: nom. inval., as syn. of Chloris submutica Kunth, 
also, Nom, Bot. |: 353 (nom. nud.) (1840). 
= Chloris submutica Kunth. 


Phalaris explicata Link, J. Bot. (Schrader) 1: 312, 1799, T- 
Protol.: Portugal: Ufern des Tajo und Douro. 
= Crypsis alopecuroides (Piller & Mitterp.) Schrad. 


Phalaris geniculata Sibth. & Sm., Fl. Graec. Prod. 1: 38. 
1806. T-Protol.: Greece: In aquis stagnantibus montis 
Olypi Bithyni. 
= Crypsis alopecuroides (Piller & Mitterp.) Schrad. 


Phalaris muricata Forssk., Fl. Aegypt.-Arab. 202. 1775. T- 
Protol.: Ad littora Maris Marmorae. 
= Tragus racemosus (L.) All. 


Phalaris vaginiflora Forssk., Fl. Aegypt.-Arab. 18. 1775. T- 
Protol.: Egypt: Alexandriae. 
= Crypsis vaginiflora (Forssk.) Opiz. 


Phleum alopecuroides Piller & Mitterp., Iter Poseg. Sclavon. 
147, t. 16. 1783. T-Protol.: Hungary: Iter per 
Poseganum Slavoniae provinciam. 

= Crypsis alopecuroides (Piller & Mitterp.) Schrad. 


Phleum dactylon Pall. ex Georgi, Beschr. Russ. Reich. 684. 
1800. 
NOTE: nom. nud., cites Pallas, Ind. Pl. Taur., may trace to 
Panicum dactylon L. 
= Cynodon dactylon (L.) Pers. 


Phleum macrourum (Kunth) Willd. ex Steud., Nomencl. Bot. 
(ed. 2) 1: 365. 1840. Basionym: Crypsis macroura 
Kunth. 

NOTE: as new, but actually a comb. 
= Muhlenbergia macroura (Kunth) Hitche. 


Phleum schoenoides L., Sp. P\. 1: 60. 1753. T-Specim.: | of 
|. Spain, Loefling s.n. [T: LINN]. 
NOTE: as "Schoenoides". 
= Crypsis schoenoides (L.) Lam. 


Phragmites zollingeri (Biise) Steud., Syn. Pl. Glumac. 1: 196. 
1855. Basionym: Arundo zollingeri Buse. 
= Neyraudia reynaudiana (Kunth) Keng ex Hitche. 


Plagiolytrum Nees, Proc. Linn. Soc. Lond. 1: 95. 1841. 
NOTE: also Ann. Mag. Nat. Hist. 7: 221 (May 1841). 
= Tripogon Roem. & Schult. 


Pleiodon Rchb., Nom. Gen. PI. 38. 1841. 
NOTE: nom. illeg. superfl. 
= Bouteloua Lag. 


Pleopogon Nutt., Proc. Acad. Nat. Sci. Philadelphia 4: 25, 21 
Mar-4 Apr 1848. TYPE: Pleopogon setosum Nutt. 
= Lycurus Kunth. 


Pleopogon setosum Nutt., Proc. Acad. Nat. Sci. Philadelphia 
4: 25. 1848. T-Specim.: | of 1. USA: New Mexico: 
mountains near Santa Fe, 1841 or 1842, William 
Gambel s.n. [HT: K; IT: US-610839 (fragm. ex K))]. 

NOTE: also in J. Acad. Nat. Sci. Philadelphia (ser. 2) 189 
(1848, Aug). 
= Lycurus setosus (Nutt.) C. Reeder. 


Pleuraphis Torr., Ann. Lyceum Nat. Hist. New York 1: 
148. 1824. TYPE: Pleuraphis jamesii Torr. 
TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 3. 


Pleuraphis jamesii Torr., Ann. Lyceum Nat. Hist. New York 
1(1): 148-150, t. 10. 1824. T-Protol.: James s.n. 
DST.: USA. 
LVL. ACCPTC.: 1; cultivated. 
ALSO ACCPTD. BY: Hickman, J. C. (1993). 
SYN.: Hilaria jamesii (Yorr.) Benth.; Hilaria sericea 
Benth.; Pleuraphis sericea Nutt. ex Benth. 


Pleuraphis mutica Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 95. 1862. T-Protol.: Northern Texas. 


Catalogue of New World Grasses: II. 


183 


T-Specim.: | of 2. USA: New Mexico, Wright 760 
[ST: PH]. 2 of 2. USA: New Mexico, Wright 2108 
[ST: PH]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 

Ferrari, and J. Valdes-Reyna (2000). 
SYN.: Hilaria mutica (Buckley) Benth. 


Pleuraphis rigida Thurb., Bot. California 2: 293-294. 1880. 
T-Specim.: | of 1. USA: California: Fort Mojave, 24 
Feb 1861, .G. Cooper 2230 [HT: US-323096]. 

DST.: Mexi, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Hickman, J. C. 
(1993). 

SYN.: Hilaria rigida (Thurb.) Benth. ex Scribn. 


Pleuraphis sericea Nutt. ex Benth., J. Linn. Soc., Bot. 19: 62. 
1881. 
NOTE: nom. inval., as syn. of Hilaria sericea Benth. 
= Pleuraphis jamesii Torr. 


Poa abyssinica Jacq., Misc. Austriac. 2: 364. 1781. T-Protol.: 
Abyssinia. 
NOTE: again Jacq. Icon. 1(2): pl. 17. 1782. 
= Eragrostis tef (Zucc.) Trotter. 


Poa acutiflora Kunth, Nov. Gen, Sp. 1: 161. 1815 [1816]. T- 
Protol.. Crescit in humidis, inundatis fluminis 
Magdalenae juxta Mompox and Isla del las Bruxas 
(Regno Novogranatensi), floret Aprili, Humboldt & 
Bonpland s.n. T-Specim.: | of 1. Colombia: on the Rio 
Magdalena, Humboldt & Bonpland 1603 (HT: P; IT: 
Pi. 

= Eragrostis acutiflora (Kunth) Nees. 


Poa airoides (Nees) Kunth, Enum. Pl. 1: 360. 1833. 
Basionym: Eragrostis airoides Nees. 
NOTE: hom. illeg.; also in Rev. Gram 1: suppl. XXVIII 
(1834). 
E. NAME: Poa airoides Koeler. 
= Eragrostis airoides Nees. 


Poa amabilis L., Sp. Pl. 68. 1753. T-Protol.: Habitat in India. 
T-Specim.: | of 3. India, Plukenet, Alm. Bot., t. 300, f 
2 (1696) [ST]. 2 of 3. Sri Lanka, Herb. Hermann 2: 
59, no. 46 {[LT: BM], LT designated by Veldkamp in 
Cafferty et al., Taxon 49(2): 254 (2000). 3 of 3. India, 
Plukenet 1: 187 (ST: BM-SL}. 
= Eragrostis amabilis (L.) Wight & Arn. ex Nees. 


Poa amabilis Walter, Fl. Carol. 80. 1788. T-Protol.: USA: 
South Carolina. T-Specim.: | of 1. Clayton 580 [T: IT: 
US-2891488 (fragm.)]. 

NOTE: hom. illeg. 

E. NAME: Poa amabilis L. 

CORRECT NAME: Poa spectabilis Pursh. 
= Eragrostis spectabilis (Pursh) Steud. 


Poa ambigua Elliott, Sketch Bot. S. Carolina 1: 165. 1816. T- 
Protol.: USA: Found in the mountains of Carolina, by 
Dr. Macbride and in the lower country of Georgia, by 
Dr. Baldwin. T-Specim.: | of 1. USA: In mont. 
Carolinae, Autumn, Macbride s.n. [ST: CHAS, IST: 
US-556771 (fragm. ex herb. Elliott, CHAS), US 
(photo ex CHAS)]. 

= Tridens ambiguus (Elliott) Schult. 


Poa articulata Schrank, Syll. Pl. Nov. 1: 194. 1824. T- 
Protol.: Patria: habitat in Brasilia. 
= Eragrostis articulata (Schrank) Nees. 


Poa arundinacea Poir., Encycl. 4: 329. 1816. 
NOTE: nom. nov., hom. illeg. 
E. NAME: Poa arundinacea Moench. 
REPLACED NAME: Poa sesleroides Michx. 
BLOCKING NAME: Poa sesleroides All. 
= Tridens flavus (L.) Hitche. 


Poa atrovirens Desf., Fl. Atlant. 1: 73, t. 14. 1798. T-Protol.: 
Algeria: Habitat in arvis incultis prope La Calle. T- 
Specim.: | of 1. Algeria: e Barbaria, Desfontaines 160 
(HT: FI; IT: LE, P]. 

= Eragrostis atrovirens (Desf.) Trin. ex Steud. 


Poa atrovirens Willd. ex Spreng., Syst. Veg. 1: 340. 1825. T- 
Protol.: Willd. herb. [no loc. given, Index of Grass 
Species|[T: US]. 

NOTE: hom. illeg. et nom. inval., as syn. of P. nigricans 
Humb. [Kunth]. 

E. NAME: Poa atrovirens Desf. 

= Eragrostis nigricans (Kunth) Steud. 


Poa aturensis Kunth, Nov. Gen. Sp. 1: 161. 1815 [1816]. T- 
Protol.: Crescit in calidis sylvarum Orinocosium prope 
Cataractas Aturensium, floret Aprili, Humboldt & 
Bonpland s.n. T-Specim.: 1 of 1. Venezuela: 
Amazonas: prope Cataractas Aturensium, Apr, 
Humboldt & Bonpland s.n. [HT: P; IT: BAA-2465 
(fragm.), SI]. 

= Eragrostis japonica (Thunb.) Trin. 


Poa berteroniana (Schult.) Kunth, Revis. Gramin. 1: 112. 
1829. Basionym: Megastachya berteroniana Schult. 
= Eragrostis berteroniana (Schult.) Steud. 


Poa biformis Kunth, Revis. Gramin. 2: 471, t. 149. 1831. T- 
Protol.: Africa: Senegambia, au Senegal. 
= Eragrostis atrovirens (Desf.) Trin. ex Steud. 


Poa borealis (J. Presl) Kunth, Enum. Pl. 1: 326. 1833. 
Basionym: Brizopyrum boreale J. Pres. 
NOTE: also Revis. Gram. 3: Suppl. 1834 p. (as in 1835 ed.) 
XXVIII. 
= Distichlis spicata var. borealis (J. Pres!) Beetle. 


Poa brasiliensis Raddi, Agrostogr. Bras. 51, 1823. T-Protol.: 
Brasilia: Invenitur in sepibus prope Rio de Janeiro, 
Raddi s.n., [type not found in Florence nor Pisa in 
1922, A.C.]. 

= Eragrostis airoides Nees. 


Poa brasiliensis Spreng., Syst. Veg. 1: 342. 1825. T-Protol.: 
Brazil: (P. elegans B. herb. Willd. ). T-Specim.: | of 1. 
Hoffmannsegg [T: B-Willd. (no. 1921. fo. 1 et 1939)]. 

NOTE: hom. illeg. 

E, NAME: Poa brasiliensis Raddi. 
CORRECT NAME: Fragrostis elegans Nees. 
= Eragrostis japonica (Thunb.) Trin. 


Poa caerulescens Michx. ex P. Beauv., Ess. Agrostogr. 77. 
1812. T-Protol.: USA: Virginia. 
NOTE: nom. nud. 
= Tridens flavus (L.) Hitche. 


Poa capillaris L., Sp. Pl. 68. 1753. T-Protol.: USA: virginaia. 
Canada. T-Specim.: | of 1. North America, Kalm s.n. 


184 


Catalogue of New World Grasses: II. 


{[LT: LINN], LT designated by Hitchcock, Contr. U.S. 
Natl. Herb. 12: 121 (1908). 
= Eragrostis capillaris (L.) Nees. 


Poa cerealis Salisb., Prodr. Stirp. Chap. Allerton 20. 1796. T- 
Protol.: Abyssinia. 
NOTE: Poa abyssinica Jacq. cited; equals Eragrostis tef 
(Zucc.) Trotter. 
= Eragrostis tef (Zucc.) Trotter. 


Poa chariis Schult., Mant. 2: 314. 1824. 
REPLACED NAME: Poa elegans Roxb. 
BLOCKING NAME: Poa elegans Poir. 
= Eragrostis atrovirens (Desf.) Trin. ex Steud. 


Poa chilensis Moris, Ann. Storia Nat. 4: 60. 1830. T-Protol.: 
Nata est in hort. bot. Taurinensi ex seminibus e Chile, 
Bertero s.n. T-Specim.: | of 1. Chile: E semina Chili 
de Taurio, C. G. Bertero s.n. {IT: IT: LE, US-911928 
(fragm. ex LE-TRIN)]. 

NOTE: Ann. Storia Nat. Bologna?. 
= Eragrostis mexicana (Hornem.) Link. 


Poa cilianensis All., Fl. Pedem. 2: 246. 1785. T-Protol.: In 
agris patrio Ciliani, Bellardi s.n. T-Specim.: | of 2. 
Italy, Bellardi s.n. [ST: TO-8242]. 2 of 2. Italy, Balbi 
sn. [ST: TO]. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Poa conceptionis Steud., Syn. Pl. Glumac. 1: 259. 1854. T- 
Protol.: Chile: Concepcion: 1844, Chauvin 187. T- 
Specim.: | of 1. Chile: Concepcion, 1844, Fu 
Chauvin 187 [d'Urville herb.] HT: P; IT: US- 
2891500 (fragm. ex P & fig. from STEUD-384)]. 

= Eragrostis polytricha Nees. 


Poa condensata (J. Presl) Kunth, Enum. Pl. 1: 344. 1833. 
Basionym: Megastachya condensata J. Presl. 
NOTE: also Revis. Gram. 3: Suppl. XXVIII (1834) (as in 
1835 ed.). 
= Eragrostis condensata (J, Presl) Steud. 


Poa curvula Schrad., Gott. Gel. Anz. 3: 2073. 1821. T- 
Specim.: | of 1. South Africa: Cape Province: Cape of 
Good Hope, Hess s.n. [T: ?]. 
= Eragrostis curvula (Schrad.) Nees. 


Poa decipiens R. Br., Prodr. 181. 1810. T-Protol.: Australia: 
Queensland: Kepple Bay: Aug 1802, R. Brown 6271. 
T-Specim.: | of 1. Australia: Queensland: Port Curtis 
Distr.: Keppel Bay, Brown 627] (HT: BM; IT: E, K]. 

= Leptochloa decipiens (R. Br.) Stapf ex Maiden. 


Poa diandra hort. ex Schrad., Linnaea 12(4): 451. 1838. 
NOTE: nom. inval., as syn. of Eragrostis purshii Schrader. 
E. NAME: Poa diandra R. Br. 
= Eragrostis pectinacea (Michx.) Nees. 


Poa digitata R. Br., Prodr. 182. 1810. T-Protol.: Australia: 
Queensland: Upper Head [Charon Point]: Broad 
Sound: 26 Sep 1802,R. Brown 6238 [error for 6266]. 
T-Specim.: | of 1. Australia: Queensland: Port Curtis 
Distr.: Broad Sound, Brown 6266 [HT: BM; IT: BM, 
E, K]. 

= Leptochloa digitata (R. Br.) Domin. 

Poa distichophylla (Labill.) R. Br., Prodr. 182. 

Basionym: Uniola distichophylla Labill. 


NOTE: hom. illeg. 
E. NAME: Poa distichophylla Gaudin. 


1810. 


= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Poa domingensis Pers., Syn, Pl. 1: 88. 1805. T-Protol.: West 
Indies: Dominican Republic: Santo Domingo: in 
arenosis maritimis ad St. Domingo. (Poiteau). 

= Eragrostis prolifera (Sw.) Steud. 


Poa elongata Willd., Enum. Pl. 1: 108. 1809. T-Protol.: India 
oriental. 
= Eragrostis elongata (Willd.) J. Jacq. 


Poa eragrostis L., Sp. Pl. 1: 68. 1753. T-Protol.: Habitat in 
Italia supra muros, D. Baeck s.n. T-Specim.: | of 1. 
Italy, Baeck s.n. [HT: LINN]. 
NOTE: as "Poa Eragrostis". 
CORRECT NAME: Eragrostis minor Host. 
= Eragrostis minor Host. 


Poa eragrostis Walter, Fl. Carol. 80. 1788. T-Protol.: USA: 
South Carolina. 
NOTE: hom. illeg. 
E. NAME: Poa eragrostis L. 
= Eragrostis pilosa (L.) P. Beauv. 


Poa eragrostis Elliott, Sketch Bot. S. Carolina I: 161. 1816. 
T-Protol.: USA: S. Carolina & Georgia: Grows in 
cultivated land. 

NOTE: hom. illeg. 
E. NAME: Poa eragrostis L. 
= Eragrostis pectinacea (Michx.) Nees. 


Poa expansa (Link) Kunth, Revis. Gramin. 1: 113. 1829. 
Basionym: Eragrostis expansa Link. 
NOTE: hom. illeg. 
E. NAME: Poa expansa J.F. Gmel. 
= Eragrostis bahiensis Schrad. ex Schult. 


Poa flava L., Sp. Pl. 68. 1753. T-Protol.: USA: Gron. Virg. 
13. Habitat in Virginia, Gronov?. T-Specim.: 1 of 1. 
USA: Virginia, Clayton 273 [HT: BM], see Hitchcock, 
Contr. U.S. Natl. Herb. 12: 120 (1908). 
= Tridens flavus (L.) Hitche. 


Poa gangetica Roxb., Fl. Ind., ed. 1820 1: 341. 1820. T- 
Protol.: India: native of the banks of the Ganges, but 
scarce. T-Specim.: | of 1. India: native of the banks of 
the Ganges, but scarce, Roxburgh s.n. [HT: K; IT: 
BM]. 

= Eragrostis gangetica (Roxb.) Steud. 


Poa glomerata Walter, Fl. Carol. 80. 1788. T-Protol.: USA: 
South Carolina, Walter s.n. T-Specim.: 1 of 1. USA: 
Carolinas, Walter s.n. [HT: BM (specimen lost)]. 

REPLACED NAME: Poa walteri Kunth. 
= Eragrostis japonica (Thunb.) Trin. 


Poa glutinosa Sw., Prodr. 26. 1788. ‘T-Protol.: Jamaica, 
Swartz s.n. [HT: S]. T-Specim.: | of 1. West Indies, 18 
Sep 1803 [IT: LE-TRIN-2629.01 a (fragm.)}. 
= Eragrostis glutinosa (Sw.) Trin. 


Poa hirsuta Michx., Fl. Bor.-Amer. 1: 68. 1803. T-Protol.: 
USA: Carolina inferiore, Michaux s.n. T-Specim.: | of 
1. USA: Carolina, Michaux s.n. [HT: P; IT: US-77389 
(fragm.)]. 
= Eragrostis hirsuta (Michx.) Nees. 


Poa hirsuta var. spectabilis (Pursh) Torr., Fl. N. Middle 
United States 1(1): 114. 1823. Basionym: Poa 
spectabilis Pursh. 


Catalogue of New World Grasses: II. 


185 


= Eragrostis spectabilis (Pursh) Steud. 


Poa hypnoides Lam., Tabl. Encycl. 1: 185. 1791. T-Protol.: 
Ex America merid., D. Richard s.n. T-Specim.: | of 1. 
Tropical America, D. Richard s.n. [HT: P-LA]. 

= Eragrostis hypnoides (Lam.) Britton, Sterns & Poggenb. 


Poa interrupta Lam., Tabl. Encycl. 1: 185. 1791. T-Protol.: 
India, Sonnerat s.n. T-Specim.: | of 1. India, Sonnerat 
g.n. (HT: P; IT: BAA-2585]. 
NOTE: Koen. ex Roxb., Fl. Ind. ed Carey & Wall., 1: 336, 
1820; Fl. Ind. ed Carey 1: 335, 1832. 
= Eragrostis japonica (Thunb.) Trin. 


Poa japonica Thunb., Fl. Jap. 51. 1784, T-Protol.: Crescit in 
Kosido et juxta Nagasaki. T-Specim.: 1 of 1. Japan, 
Thunberg s.n. [HT: UPS]. 

= Eragrostis japonica (Thunb.) Trin. 


Poa linkii Kunth, Revis. Gramin, 1: 113. 1829. 
REPLACED NAME: Eragrostis filiformis Link. 
BLOCKING NAME: Poa filiformis Thunb. 
= Eragrostis pilosa (L.) P. Beauv. 


Poa lugens (Nees) Kunth, Enum. PI. 1: 331. 1833. Basionym: 
Eragrostis lugens Nees. 
NOTE: also Revis. Gram. 3: Suppl. 1834 p. (as in 1835 ed.) 
XXVIII. 
= Eragrostis lugens Nees. 


Poa lurida (J. Presl) Kunth, Enum. Pl. 1: 342. 
Basionym: Eragrostis lurida J. Presl. 
NOTE: also Revis. Gram. 3: Suppl. XXVIII (1834) (as in 
1835 ed.). 
= Eragrostis lurida J. Presl. 


1833. 


Poa malabarica L., Sp. Pl. 69. 1753. T-Specim.: | of 1. 
Rheede, Hort. Mal. 12, t. 45 (1703) [LT], LT 
designated by Merrill, Bull. Torrey Bot. Club 60: 633- 
635 (1933); cited by Nowack, Rheedea 4(2): 84 
(1994). 

NOTE: nom. utiq. rej. [see Brummitt, Taxon 49(2): 266 
(2000)]. 
= Leptochloa fusca (L.) Kunth. 


Poa maypurensis Kunth, Nov. Gen. Sp. 1: 161. 1815 [1816]. 
T-Protol.: Crescit in opacatis sylvarum Orinocensium 
inter Cataractam Sancti Josephi Maypurensium et 
confluentem Sipapum, floret Aprili, Humboldt & 
Bonpland s.n. T-Specim.: 1 of 1. Venezuela: 
Amazonas: on the Rio Orinoco, Apr, Humboldt & 
Bonpland s.n. (HT: P; IT: BAA-1053 (fragm.), P]. 

= Eragrostis maypurensis (Kunth) Steud. 


Poa mexicana Hornem., Hort. Bot. Hafn. 2: 953. 1815. T- 
Protol.: Hab. in Mexico? Garden specimen from 
Mexican seed. T-Specim.: | of 2. Cultivated from seed 
collected in Mexico, Sessé s.n. [ST: MA]. 2 of 2. 
Brazil, Sessé & M. Lacasta {IST?: US-2891498 
(fragm.)]. 

= Eragrostis mexicana (Hornem.) Link. 


Poa michauxii Kunth, Revis. Gramin. 1: 111. 1829. 
REPLACED NAME: Festuca distichophylla Michx. 
BLOCKING NAME: Poa distichophylla Gaudin. 
= Distichlis spicata (L.) Greene. 


Poa micrantha Schult., Mant. 2: 305. 1824. 
REPLACED NAME: Poa parviflora Nutt. 
BLOCKING NAME: Poa parviflora R. Br. 


= Eragrostis frankii C.A. Mey. ex Steud. 


Poa microstachya Link, Hort. Berol. 1: 185. 1827. T-Protol.: 
Habitat in Montevideo. T-Specim.: 1 of 1. Uruguay: 
Montevideo, Anonymous |[HT: B; IT: BAA-989 
(fragm. ex B)]. 

= Eragrostis bahiensis Schrad. ex Schult. 


Poa modesta Tuck., Amer. J. Sci. 45(1): 45. 1843. T-Protol.: 
USA: Cambridge, wet margins of Fresh Pond brook. 
= Muhlenbergia uniflora (Muhl.) Fernald. 


Poa montufari Kunth, Nov. Gen. Sp. 1: 159. 1815 [1816]. T- 
Protol.: Crescit in alta planitie Chillensi prope la 
Hacienda de Don Carlos Montufar: item juxta Puembo 
et San Antonio de Lulumbabmba, inter 1200 et 1400 
hexap. (Regno Quitensi), floret Aprili, Majo, 
Humboldt & Bonpland s.n. (HT: P|. T-Specim.: 1 of 1. 
Ecuador: between Puembo and San Antonio de 
Lulumbamba, Humboldt & Bonpland s.n. (HT: P- 
Bonpl; IT: BAA-1056, US-2891497 (fragm.)]. 

= Eragrostis pastoensis (Kunth) Trin. 


Poa neesii (Trin.) Kunth, Enum. PI. 1: 339, 1833. Basionym: 
Eragrostis neesii Trin. 
= Eragrostis neesii Trin. 


Poa nigricans Kunth, Nov. Gen. Sp. 1: 159. 1815 [1816]. T- 
Protol.: Crescit cum Poa patula Kunth prope 
Conocoto, Chillo et Sangolqui, 1360 hexap., floret 
Aprili, Majo, Humboldt & Bonpland s.n. T-Specim.: | 
of 1. Ecuador: Chillo and Sangolqui, F. WH. Humboldt 
& AJA. Bonpland 229] (HT: P-Bonpl; IT: BAA-2281 
(fragm.), LE-TRIN-2371.01, US-2891495 (fragm. ex 
LE-TRIN, fragm. ex P)]. 

= Eragrostis nigricans (Kunth) Steud. 


Poa nitida Elliott, Sketch Bot. S. Carolina 1: 162. 1816. T- 
Protol.: USA: South Carolina: Common on Paris 
Island, grows on cultivated land, Elliott s.n. [HT: 
CHARL]. T-Specim.: 1 of 1. America boreal., E//iot 
com. Herb. Ledeb. : Herb. Zeyheri [1T: LE]. 

NOTE: hom. illeg. 

E. NAME: Poa nitida Lam. 

CORRECT NAME: Fragrostis elliottii S. Watson. 
= Eragrostis elliottii S. Watson. 


Poa nuttallii Spreng., Syst. Veg. 1: 344. 1825. 
NOTE: nom. nov., nom. illeg. superfl. 
REPLACED NAME: Poa obtusa Nutt. 
BLOCKING NAME: Poa obtusa Muhl. 
CORRECT NAME: Poa pennsylvanica Nutt. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Poa nuttallii Kunth, Revis. Gramin. 1: 116. 1829. 
NOTE: nom. nov., hom. illeg. 
E. NAME: Poa nuttallii Spreng. 
REPLACED NAME: Poa tenella Nutt. 
BLOCKING NAME: Poa tenella L. 
= Eragrostis pectinacea (Michx.) Nees. 


Poa obtusa Nutt., Gen. N. Amer. Pl. 1: 67. 1818. T-Protol.: 
USA: Pennsylvania?: Collected in the neighborhood of 
Philadelphia, W. Barton. T-Specim.: 1 of 1. USA: New 
Jersey, no date, Coll. Ukn. s.n. (IT: US-1535786]. 

NOTE: hom. illeg. 

E. NAME: Poa obtusa Muhl. 

CORRECT NAME: Poa pennsylvanica Nutt. 

= Eragrostis cilianensis (All.) Vignolo ex Janch. 


186 


Catalogue of New World Grasses: II. 


Poa olmedoi Kunth, Nov. Gen. Sp. 1: 159. 1815 [1816]. T- 
Protol.: Peru: Crescit in montibus Peruanis et Loxae ad 
vias, juxta domum botanici hispani Don Vicente 
Olmedo, cui cura caedendarum  Cinchonarum 
Uritusingensium demandata est, alt. 1100 hexap., 
floret Julio, Humboldt & Bonpland s.n. [HT: P]. 

NOTE: as Poa "olmedi". 
= Eragrostis pastoensis (Kunth) Trin. 


Poa ovina A. Rich., Tent. Fl. Abyss. 2: 428. 1851. T- 
Specim.: | of 2. Ethiopia: in campis montosis 
provinciae Chire, Schimper, pl. Schimp. Abyss., sect. 
HI, 1831 (ST: P; IST: K]. 2 of 2. Ethiopia: in campis 
montosis provinciae Chire, Quartin Dillon s.n. (ST: P]. 

= Eragrostis gangetica (Roxb.) Steud. 


Poa oxylepis Torr., Explor. Red River Louisiana 301, t. 19. 
1853. 
REPLACED NAME: Poa interrupta Nutt. 
BLOCKING NAME: Poa interrupta Lam. 
= Eragrostis secundiflora subsp. oxylepis (Torr.) S.D. 
Koch. 


Poa panicea Retz., Observ. Bot. 3: 11. 1783. T-Specim.: 1 of 
1. China, Bladh s.n. |HT: LD]. 
= Leptochloa panicea (Retz.) Ohwi. 


Poa paniciformis A. Braun, Flora 24: 274. 1841. T-Protol.: 
Cult. in) garden in Carlsruhe, Germany [from 
Abyssinia]. 

= Eragrostis paniciformis (A. Braun) Steud. 


Poa panicoides (J. Presl) Kunth, Enum. Pl. 1: 343. 1833. 
Basionym: Megastachya panicoides)J. Presl. 
NOTE: also Revis. Gram. 3: Suppl. 1834 p. (as in 1835 ed.) 
XXVIII. 
= Leptochloa panicoides (J. Presl) Hitche. 


Poa paradoxa Roem. & Schult., Syst. Veg. 2: 569. 1817. 
REPLACED NAME: Uniola distichophylla Labill. 
BLOCKING NAME: Poa distichophylla Gaudin. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Poa parviflora Nutt., Gen. N. Amer. Pl. 1: 67, 1818. T- 
Protol.: USA: Ohio. 
NOTE: hom. illeg. 
E. NAME: Poa parviflora R. Br. 
CORRECT NAME: Poa micrantha Schult. 
= Eragrostis frankiiC.A. Mey. ex Steud. 


Poa pastoensis Kunth, Nov. Gen. Sp. 1: 160. 1815 [1816]. T- 
Protol.: Colombia: Colombia: Crescit’ in regni 
Novogranatensis apricis, tempratis, prope urbem Pasto, 
in radicibus montis Arandensis: 1340 hexap.: Dec., 
Humboldt & Bonpland 2149. T-Specim.: 1 of 1. 
Colombia: Pasto Mt. Arand, Bonpland 2/49 [HT: P- 
HBK; IT: US-2767397 (fragm. ex P-HBK)]. 

= Eragrostis pastoensis (Kunth) Trin. 


Poa patula Kunth, Nov. Gen. Sp. 1: 158. 1815 [1816]. T- 
Protol.: Crescit in apricis, termperatis regni Quitensis 
juxta pagum Conocoto: 1360 hex., Humboldt & 
Bonpland s.n. T-Specim.: | of 1. Ecuador, no date, 
Humboldt & Bonpland s.n. [1V: US-2766206 (fragm.)]. 

= Eragrostis patula (Kunth) Steud. 


Poa pectinacea Michx., Fl. Bor.-Amer. 1: 69. 1803. T- 
Protol.: Habitat in arvis Ilinoensibus, Michaux s.n. T- 
Specim.: | of 1. USA: Illinois, Michaux s.n. [HT: P- 
MICH; IT: US-2851264 (fragm.)]. 


= Eragrostis pectinacea (Michx.) Nees. 


Poa pectinacea Geyer ex Steud., Syn. Pl. Glumac. 1: 272. 
1854. T-Protol.: USA: Illinois. 
NOTE: hom. illeg. et nom. inval., as syn. of Eragrostis 
geyeri Steudel. 
E. NAME: Poa pectinacea Michx. 
= Eragrostis spectabilis (Pursh) Steud. 


Poa peruviana Jacq., Collectanea 1: 107. 1786 [1787]. T- 
Specim.: | of 1. Peru, NV. von Jacquin [HT: ?; IT: 
US-2851260 (fragm.)]. 

= Eragrostis peruviana (Jacq.) Trin. 


Poa philadelphica W.P.C. Barton, Comp. Fl. Philadelph. 1: 
62. 1818. 
NOTE: nom. nov., nom. illeg. superfl. 
REPLACED NAME: Poa obtusa Nutt. 
BLOCKING NAME: Poa obtusa Muhl. 
CORRECT NAME: Poa pennsylvanica Nutt. 
= Eragrostis cilianensis (All.) Vignolo ex Janch. 


Poa pilosa L., Sp. Pl. 1: 68. 1753. T-Protol.: Habitat in Italia. 
T-Specim.: | of 3. Scheuzer, Agrostographia 193, t. 4, 
Sf 3 (1719) [LT], LT designated by Du Puy, Telford & 
Edgar, Fl. Australia 50: 472 (1993). 2 of 3. Italy: an 
Wegen zwischen den Reisfeldern von Pldenico unweit 
Vercelli in Oberitalien, 9-10 Aug 1902, 4. Kneucker, 
Gram. Exsicc. XII, 344 [ET: B], ET designated by 
Scholz in Cafferty et al., Taxon 49(2): 256 (2000). 3 of 
3. Italy, Tiumfetti s.n. [LT], LT designated by Clayton, 
Fl. Trop. East Africa, Gramineae 3: 214 (1974), but as 
this is not original material, the lectotypification is 
ineffective [fide Cafferty et al. Taxon 49(2): 256 
(2000)]. 

= Eragrostis pilosa (L.) P. Beauv. 


Poa plumosa Retz., Observ. Bot. 4: 20. 1786. T-Protol.: E. 
Tranquebaria, Adnig s.n. 
= Eragrostis amabilis (L.) Wight & Arn. ex Nees. 


Poa polytricha (Nees) Kunth, Enum. Pl. 1: 331. 1833. 
Basionym: Eragrostis polytricha Nees. 
NOTE: also Revis. Gram. 3: Suppl. XXVIII (1834) (as in 
1835 ed.). 


= Eragrostis polytricha Nees. 


Poa preslii Kunth, Enum. Pl. 1: 326. 1833. 
NOTE: nom. nov., also Revis. Gram. 3: Suppl. 1834 p. (as 
in 1835 ed.) XXVIHIL. 
REPLACED NAME: Brizopyrum pilosum J. Presl. 
BLOCKING NAME: Poa pilosa L. 
= Jouvea pilosa (J. Presl) Scribn. 


Poa prolifera Sw., Prodr. 27. 1788. T-Protol.: Insulae 
Caribbaeae: Santa Lucia, Guadeloupe. T-Specim.: | of 
1. Guadeloupe, Swartz s.n. (HT: S: IT:K]. 
= Eragrostis prolifera (Sw.) Steud. 


Poa prostrata Kunth, Nov. Gen. Sp. 1: 157. 1815 [1816]. T- 
Protol.: Venezuela: Crescit copiose in valle temperata, 
amoena Caripensi, alt. 420 hex., (Prov. Novae- 
Andalusiae), floret Septembri, Humboldt & Bonpland 
sn. (HT: P]. T-Specim.: 1 of 1. Venezuela: Pratis 
Carpio, F. WH. Humboldt & A.J.A. Bonpland 4161 [T: 
IT: US-2891494 (fragm.)}. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Poa psammodes (Trin.) Kunth, Enum, Pl. 1: 328. 1833. 


Basionym: Eragrostis psammodes Trin. 


Catalogue of New World Grasses: II. 


187 


= Eragrostis bahiensis Schrad. ex Schult. 


Poa purpurascens Spreng., Novi Provent. 33 no. 73. 1819. T- 
Protol.: Uruguay. 
= Eragrostis purpurascens (Spreng.) Schult. 


Poa pycnantha Phil., Anales Univ. Chile 94: 165. 1896. T- 
Protol.: Chile: prope Copiapo, Sep 1885, F. Philippi 
sn. T-Specim.: | of 1. Chile: prope Copiapo, Sep 
1885, F. Philippi s.n. (HT: SGO; IT: B, BAA-1079 
(fragm. ex B), W]. 

= Eragrostis pycnantha (Phil.) Parodi ex Nicora. 


Poa quinquefida Pursh, Fl. Amer. Sept. 1: 81. 1814. 
REPLACED NAME: Poa sesleroides Michx. 
BLOCKING NAME: Poa sesleroides All. 
= Tridens flavus (L.) Hitche. 


Poa racemosa Vahl, Eclog. Amer. 1: 7. 1796. T-Protol.: 
Habitat in America meridionali, von Rohr 95 [HT: C]. 
NOTE: hom. illeg. 
E. NAME: Poa racemosa Thunb. 
CORRECT NAME: Poa vahliiRoem. & Schult. 
= Eragrostis maypurensis (Kunth) Steud. 


Poa refracta Muhl. ex Elliott, Sketch Bot. S. Carolina 1; 162. 
1816. T-Protol.: USA: South Carolina: in damp soils, 
in pine barrens, and common along roads. T-Specim.: 
1 of 1. USA: South Carolina, S. Elliott s.n. (HT: 
CHARL; IT: PH, US-2767400 (fragm.)]. 

= Eragrostis refracta (Muhl. ex Elliott) Scribn. 


Poa rufescens Schrad. ex Schult., Mant. 2: 319. 1824. T- 
Protol.: In Brasilia, Maxim. Neowidens s.n. T-Specim.: 
1 of 2. 1819, herb. Schrader [OM: LE]. 2 of 2. , 
Schrader Herb. Ledebour: Hfortus] Dorp. 21 [OM: 
LE]. 
NOTE: nom. inval., as syn. of Eragrostis? rufescens Schrad 
MS. 
= Eragrostis rufescens Schrad. ex Schult. 


Poa sclerophylla (Trin.) Kunth, Enum. Pl. 1; 340. 1833. 
Basionym: Eragrostis sclerophylla Trin. 
= Eragrostis sclerophylla Trin. 


Poa scoparia Nees ex Kunth, Revis. Gramin. 2: 535, t. 182. 
1829 [1832]. T-Protol.: Crescit in Brasilia meridionali. 
T-Specim.: | of |. Brasilia meridionalis, Se//ow s.n. 
[IT: LE, LE-TRIN-2501.01 a, US-91379 (fragm.)]. 

NOTE: cites Uniola scoparia Nees in herb. 
= Distichlis scoparia (Nees ex Kunth) Arechav. 


Poa secundiflora (J. Presl) Kunth, Enum. Pl. 1: 342. 1833. 
Basionym: Eragrostis secundiflora J. Presl. 
NOTE: also Revis. Gram. 3: Suppl. XXVIII (1834) (as in 
1835 ed.). 
= Eragrostis secundiflora J. Presl. 


Poa seminuda (Trin.) Kunth, Enum. Pl. 1: 340. 1833. 
Basionym: Eragrostis seminuda Trin. 


= Eragrostis seminuda Trin. 


Poa sesleroides Michx., Fl. Bor.-Amer. |: 68-69. 1803. T- 
Protol.: USA: in regione Illinoensi et in montosis 
Carolinae, Michaux. T-Specim.: 1 of 1. USA: 
Carolina, Michaux s.n. [HT: P]. 

NOTE: hom. illeg. 

E. NAME: Poa sesleroides All. 

CORRECT NAME: Poa quinquefida Pursh. 
= Tridens flavus (L.) Hitche. 


Poa setifolia Benth., Pl. Hartw, 262. 1846. T-Protol.: 
Ecuador: in arvis prope Quito[HT: K]. T-Specim.: 1 of 
1. Ecuador: Quito, Hartweg 1/452 [HT: K; IT: US- 
2826937 (fragm. ex K)]. 
= Eragrostis pastoensis (Kunth) Trin. 


Poa simpliciflora (J. Presl) Kunth, Enum, Pl. 1: 343. 1833. 
Basionym: Megastachya simpliciflora J. Pres. 
NOTE: also Revis. Gram. 3: Suppl. XXVIII (1834) (as in 
1835 ed.). 
= Eragrostis simpliciflora (J. Presl) Steud. 


Poa solida (Nees) Kunth, Enum. Pl. 1: 336. 1833. Basionym: 
Eragrostis solida Nees. 
NOTE: also Revis. Gram. 3: Suppl. XXVIII (1834) (as in 
1835 ed.). 
= Eragrostis solida Nees. 


Poa spectabilis Pursh, Fl. Amer. Sept. 1: 81. 1814 [1813]. 
REPLACED NAME: Poa amabilis Walter. 
BLOCKING NAME: Poa amabilis L. 
= Eragrostis spectabilis (Pursh) Steud. 


Poa sporoboloides A. Rich., Tent. Fl. Abyss. 2: 426. 1850. T- 
Specim.: | of 3. Ethiopia: juxta Adiabo, non procul a 
convalle fluvii Mareb, Quartin Dillon et Ant. Petit 
[ST: P]. 2 of 3. Ethiopia: planitie montosa prope 
Kouaieta, in prov. Chiré, Quartin Dillon et Ant. Petit 
[ST: P]. 3 of 3. , Schimper, pl. Abyss., sect. 1, no. 401 
[ST]. 

= Eragrostis japonica (Thunb.) Trin. 


Poa squarrosa Salzm. ex Steud., Syn. Pl. Glumac. 1: 274. 
1854. T-Protol.: Bahiense. 
NOTE: nom. inval., as syn. of Eragrostis reptans Michx. 
E. NAME: Poa squarrosa Licht. ex Roem. & Schult. 
= Eragrostis maypurensis (Kunth) Steud. 


Poa stricta var. uniflora Muhl. ex Scribn. & Merr., Bull. Div. 
Agrostol., U.S.D.A. 27: 5. 1900. 
NOTE: nom. inval., as syn. of Sporebolous uniflorus Muhl. 
= Muhlenbergia uniflora (Muhl.) Fernald. 


Poa tef Zucc., Diss. Ditef s.p. 1775. T-Specim.: | of 1, Cult. 
at Florence from seed collected in Ethiopia, seed coll. 
by Bruce [HT: FI]. 

= Eragrostis tef (Zucc.) Trotter. 


Poa tenax Kunth, Prodr. 1: 160. 1816. T-Protol.: Ecuador: 
Crescit in apricis regni Quitensis, prope Chillo et Rio 
Guangopolo, alt. 1350 hexcap., floret Aprili, Majo, 
Humboldt & Bonpland s.n. (HT: P], Kunth 7: pl. 692. 
1825. 

= Eragrostis condensata (J. Presl) Steud. 


Poa tenella L., Sp. Pl. 1: 69. 1753. T-Protol.: Habitat in India. 
T-Specim.: | of 1. India, Anonymous [HT: LINN- 
87.33]. 

= Eragrostis amabilis (L.) Wight & Arn. ex Nees. 


Poa tenella Willd.? (L. misappl. by) Pursh, Fl. Amer. Sept. 1: 
80. 1814. T-Protol.: USA: In sandy fields New Jersy to 
Carolina in herb. Lambert sub. nominae P. pilosae. 

NOTE: misapl. [Index of Grass Species], hom. illeg.” again 
by Ell. Bot. S.C. & Ga 1: 160 (1816). 

E. NAME: Poa tenella L. 

= Eragrostis pilosa (L.) P. Beauv. 


Poa tenella Nutt., Gen. N. Amer. Pl. 1: 67. 1818. T-Protol.: 
[North America]. 


188 


Catalogue of New World Grasses: II. 


NOTE: hom. illeg. 

E. NAME: Poa tenella L. 

CORRECT NAME: Eragrostis brizoides Schult. 
= Eragrostis pectinacea (Michx.) Nees. 


Poa tenuifolia A. Rich., Tent. Fl. Abyss. 2: 425. 1851 [1850]. 
T-Protol.: Abyssinia. T-Specim.: | of 2. Ethiopia: in 
locis incultis Vallium prope Adoam, Jan 1841,G.H.W. 
Schimper 92 (ST: K, L, W, US-1127147 (fragm.)]. 2 
of 2. Ethiopia: crescit in montosis Chiré, Oct, Quartin 
Dillon s.n. (IST: P]. 

= Eragrostis tenuifolia (A. Rich.) Hochst. ex Steud. 


Poa tenuis Elliott, Sketch Bot. S. Carolina 1: 156. 1816. T- 
Protol.: US: South Carolina: Greenville, Moulins s.n. 
NOTE: hom. illeg. 
E. NAME: Poa tenuis Clairy. 
= Eragrostis capillaris (L.) Nees. 


Poa tephrosanthos Spreng. ex Schult., Mant. 2: 316. 1824. 

NOTE: nom. inval., as syn. of Eragostis tephrosanthos 
Schultes. 

= Eragrostis pectinacea var. miserrima (E. Fourn.) 
Reeder. 


Poa texana Vasey, Contr. U.S. Natl. Herb. 1(2): 60. 1890, T- 
Protol.: USA: Texas: from Brazos Santiago to El Paso 
Co. in the region of the Rio Grande, Nealley s.n. T- 
Specim.: | of 1. USA: Texas: El Paso Co., 1887,G.C. 
Nealley 1078 (HT: US-81565]. 
= Allolepis texana (Vasey) Soderstr. & H.F. Decker. 


Poa thalassica Kunth, Nov. Gen. Sp. 1: 157. 1815 [1816]. T- 
Protol.: Peru: Crescit in littore Oceani Pacifici prope 
Callao de Lima, Guamang et Urbem_ Santa 
Peruvianorum, floret Octobri, Humboldt & Bonpland 
sn. [3719] [HT: P|. T-Specim.: 1 of 1. Peru: Lima: 
Prope ad Litus Oceani, fF. WH. Humboldt Y A.J.A. 
Bonpland 3719 [T: IT: US-2891486 (fragm.)]. 

= Distichlis spicata (L.) Greene. 


Poa trichodes Nutt., Trans, Amer. Philos. Soc., n.s., 5: 146. 
1835. T-Protol.: Arkansas Territory: in brushy prairies 
and open alluvial lands, Nuttall s.n. T-Specim.: | of 1. 
USA: Arkansas: bushy prairies and open alluvial 
lands, 7. Nuttall s.n. [HT: PH; IT: US-77391 (fragm.)]. 
= Eragrostis trichodes (Nutt.) Alph. Wood. 


Poa uniflora Muhl., Cat. Pl. Amer. Sept. 11. 1813. 
NOTE: nom. nud. 
CORRECT NAME: Poa uniflora Muhl. 
= Muhlenbergia uniflora (Muhl.) Fernald. 


Poa uniflora Muhl., Descr. Gram. 151. 1817. T-Protol.: USA: 
Nova Anglia et plata simillima in Nova Caesarea. T- 
Specim.: | of 1. USA, Muhlenberg 223 [HT: PH; IT: 
US-1535784 (fragm. ex PH)], US specimen notes PH 
slips 146 & 413. 

= Muhlenbergia uniflora (Muhl.) Fernald. 


Poa uninervia (J. Presl) Kunth, Enum. Pl. 1: 344. 1833. 
Basionym: Megastachya uninervia J. Presl. 
NOTE: also Revis. Gram. 3: Suppl. 1834 p. (as in 1835 ed.) 
344, 
= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Poa unioloides Retz., Observ. Bot. 5: 19. 1788. T-Protol.: 
India. T-Specim.: 1 of |. India, 1776, Koenig s.n. [ST: 
BM (2 sheets), LE]. 
= Eragrostis unioloides (Retz.) Nees ex Steud. 


Poa vahlii Roem. & Schult., Syst. Veg. 2: 563. 1817. 
NOTE: pro. parte. = Eragrostis maypurensis. 
REPLACED NAME: Poa racemosa Vahl. 
BLOCKING NAME: Poa racemosa Thunb. 
= Eragrostis maypurensis (Kunth) Steud. 


Poa verticillata Cav., Icon. 1(3): 63, pl. 93. 1791. T-Protol.: 
Habitat in arenosis humidis ad fluminis Manzanares. 
= Eragrostis pilosa (L.) P. Beauv. 


Poa villosa Beyr. ex Schrad., Linnaea 12(4): 451, 1838. 
NOTE: nom. inval., as syn. of Eragrostis velutina Schrader. 
= Eragrostis spectabilis (Pursh) Steud. 


Poa virescens (J. Presl) Kunth, Enum. Pl. 1: 329. 1833. 
Basionym: Eragrostis virescens J. Pres. 
= Eragrostis mexicana subsp. virescens (J. Presl) S.D. 
Koch & Sanchez Vega. 


Poa virgata Poir., Encycl. 5: 78. 1804. T-Protol.: West 
Indies: Saint-Dominique[HT: P-LA]. 
= Uniola virgata (Poir.) Griseb. 


Poa virginica Zuccagni, Cent. Observ. Bot. 1: [p. 12]no. 25. 
1806. T-Protol.: USA: [Virginia]. 
NOTE: Preprint of J.J. Roemer, Collectanea p. 124 (1806). 
= Eragrostis refracta (Muhl. ex Elliott) Scribn. 


Poa viscosa Retz., Observ. Bot. 4: 20. 1786. T-Specim.: 1 of 
|. India, Koenig s.n. [HT: LD; IT: BM]. 
= Eragrostis viscosa (Retz.) Trin. 


Poa viscosa var. pilosissima Hochst. ex A. Rich., Tent. FI. 
Abyss. 2: 424. 1850. T-Protol.: Cresit in rupestribus 
siccis prope Guendepta, Sep, Schimper, pl. Abyss., 
sect. ll, no. 824. 

= Eragrostis viscosa (Retz.) Trin. 


Poa UNRANKED Psilantha K. Koch, Linnaea 21(4): 405. 
1848. TYPE: Poa collina (Trin.) K. Koch. 
= Eragrostis Wolf. 


Podosaemum Kunth, Mem. Mus. Hist. Nat. 2: 72. 1815. 
NOTE: orth. var. Podosemum. 
= Muhlenbergia Schreb. 


Podosaemum mexicanum (L.) Link, Hort. Berol. 1; 84. 1827. 
Basionym: Agrostis mexicana L. 
NOTE: orth. var. 
= Muhlenbergia mexicana (L.) Trin. 


Podosemum Desv., Nouv. Bull. Sci. Soc. Philom. Paris 2: 
188. 1810. TYPE: Podosemum capillare (Lam.) Desv. 
= Muhlenbergia Schreb. 


Podosemum affine (Trin.) Bush, Amer. Midl. Naturalist 7(2): 
40. 1921. Basionym: Muhlenbergia affinis Trin, 
= Muhlenbergia rigida (Kunth) Kunth. 


Podosemum agrostideum P. Beauv., Ess. Agrostogr. 176, 
179. 1812. 
NOTE: nom. nud., equals Tosagris agrostideum P. Beauv, 
= Muhlenbergia capillaris (Lam.) Trin. 


Podosemum angustatum J. Presl, Reliq. Haenk. 1(4-5): 229. 
1830. T-Protol.: Habitat in Peruvia, Haenke s.n. T- 
Specim.: 1 of 1. Peru, Haenke s.n. [HT: PR]. 

NOTE: as "Podosaemum” angustatum. 
= Muhlenbergia angustata (J. Presl) Kunth. 


Catalogue of New World Grasses: II. 


189 


Podosemum arenicola (Buckley) Bush, Amer. Midl. 
Naturalist 7(2): 40. 1921. Basionym: Muhlenbergia 
arenicola Buckley. 

NOTE: as Podosemum "arenicolum". 
= Muhlenbergia arenicola Buckley. 


Podosemum brachyphyllum Nees, Linnaea 19(6): 690. 1847. 
T-Protol.: Mexico, Aschenb. pl. exs. 114. T-Specim.: | 
of 1. Mexico, Aschenborn 114 [T: ?]. 

NOTE: as "Podosaemum". 
= Muhlenbergia ciliata (Kunth) Trin. 


Podosemum capillare (Lam.) Desv., Nouv. Bull. Sci. Soc. 
Philom. Paris 2: 188. 1810. Basionym: Stipa capillaris 
Lam. 
= Muhlenbergia capillaris (Lam.) Trin. 


Podosemum ciliatum Kunth, Nov. Gen. Sp. 1: 128. 1815 
[1816]. T-Protol.: Crescit in radicibus montis ignivomi 
Mexicani, Jorullo, alt. 490 hexap., floret Septembri, 
Humboldt & Bonpland s.n. T-Specim.: | of 1. Mexico: 
Michoacan: Volcan de Jorullo, Humboldt & Bonpland 
sn. [HT: P-Bonpl.; IT: BM, P-Bonpl., US-91918 
(fragm.)]. 

NOTE: as "Podosaemum" ciliatum. 
= Muhlenbergia ciliata (Kunth) Trin. 


Podosemum debile Kunth, Nov. Gen. Sp. 1: 128. 1815 
[1816]. T-Protol.: Ecuador: Crescit in umbrosis calidis 
vallis Guallabamba, alt. 1030 hexap., (Regno 
Quitensi), floret Januario, Humboldt & Bonpland s.n. 
T-Specim.: 1 of 1. Ecuador: Proy. Pichincha: Quito, 
Humboldt & Bonpland s.n. |HT: P-Bonpl.; IT: B-W, P, 
US-91924 (fragm. ex P-Bonpl.)]. 

NOTE: as "Podosaemum" debile. 
= Muhlenbergia microsperma (DC.) Kunth. 


Podosemum distichophyllum J. Presl, Relig. Haenk. 1(4-5): 
231. 1830. T-Protol.: Habitat in Mexico, Haenke s.n. 
T-Specim.: | of 1. Mexico, 7) Haenke s.n. (HT: PR 
(photo, US); IT: MO-1837831, US-90711 (fragm. ex 
PR, fragm. ex W)]. 

NOTE: as "Podosaemum" distichophyllum. 
= Muhlenbergia distichophylla (J. Presl) Kunth. 


Podosemum elegans Kunth, Nov. Gen. Sp. 1: 130. 1815 
[1816]. T-Protol.: Ecuador: "Crescit regione frigida 
Amdium Quitensium inter Calpi et radices montis 
Chimborazo, Paramo de las Puntas et Pomallacta: alt. 
1600 hexap.". 

NOTE: as "Podosaemum" elegans. 
= Muhlenbergia rigida (Kunth) Kunth. 


Podosemum filipes (M.A. Curtis) Bush, Amer. Midl. 
Naturalist 7(2): 29. 1921. Basionym: Muhlenbergia 
filipes M.A. Curtis. 

= Muhlenbergia sericea (Michx.) P.M. Peterson. 


Podosemum foliosum (hort. ex Roem. & Schult.) Link, Hort. 
Berol. 1: 83. 1827. Basionym: Agrostis foliosa hort. ex 
Roem. & Schult. 

NOTE: as "Podosaemum". 
= Muhlenbergia mexicana var. filiformis (Torr.) Scribn. 


Podosemum glabratum Kunth, Nov. Gen. Sp. 1: 130. 1815 
[1816].  T-Protol.: Crescit cum  Podosaemum 
mucronatum juxta Santa Rosa de la Sierra et Cafiada 
de Acabuca, floret Septembri, Humboldt & Bonpland 
s.n. T-Specim.: | of 1. Mexico: S. Rosa de la Sierra & 


Cafiada de Acabuca, n.d. [HT: P-HBK; US-91921 
(fragm. ex P-HBK Herb.)]. 

NOTE: as "Podosaemum" glabratum. 

= Muhlenbergia rigida (Kunth) Kunth. 


Podosemum glaucum Nees, Linnaea 19(6): 689. 1847. T- 
Protol.: Mexico, Aschenborn exs. 335. 
NOTE: as "Podosaemum". 
= Muhlenbergia glauca (Nees) B.D. Jacks. 


Podosemum glomeratum (Willd.) Link, Hort. Berol. 1: 84. 
1827. Basionym: Polypogon glomeratus Willd. 
NOTE: as "Podosaemum". 
= Muhlenbergia glomerata (Willd.) Trin. 


Podosemum gracile Kunth, Nov. Gen. Sp. 1: 131. 1815 
[1816]. T-Protol.: Crescit in aridis exustis montis 
Mexicani, Volcan de Jorullo: alt. 580 hexap, Humboldt 
& Bonpland s.n. T-Specim.: 1 of 1. Mexico: 
Michoacan: Volcan de Jorullo, Humboldt & Bonpland 
s.n. [HT: P; IT: LE-TRIN-1501.02, US-86636 (fragm. 
ex P-HBK Herb)]. 

NOTE: as "Podosaemum" gracile. 
= Muhlenbergia quadridentata (Kunth) Trin. 


Podosemum — gracillimum (Torr.) Bush, Amer. Midl. 
Naturalist 7(2): 33. 1921. Basionym: Muhlenbergia 
gracillima Torr. 

= Muhlenbergia torreyi (Kunth) Hitchc. ex Bush. 


Podosemum implicatum Kunth, Nov. Gen. Sp. 1: 127. 1815 
[1816]. T-Protol.: Crescit in humidis, uliginosis 
Andium Mexicanorum prope lacum Cuiseo et Puerto 
de Andaracus, alt. 940 hexap., floret Septembri, 
Humboldt & Bonpland s.n. T-Specim.: | of 1. Mexico: 
Michoacan: near Lake Cuiseo and Puerto de 
Andaracuas, Humboldt & Bonpland s.n. (HT: B-W; 
IT: P]. 

NOTE: as "Podosaemum" implicatum. 
= Muhlenbergia implicata (Kunth) Trin. 


Podosemum mexicanum (L.) Link, Hort. Berol. 1: 84. 1827. 
Basionym: Agrostis mexicana L. 
NOTE: as "Podosaemum"; "Muhlenbergia mexicana L." 
cited, error for Agrostis mexicana L. 
= Muhlenbergia mexicana (L.) Trin. 


Podosemum mucronatum Kunth, Nov. Gen. Sp. 1: 129. 1815 
[1816]. T-Protol.: Crescit in montanis prope Cerro de 
Serena, S. Rosa et Los loares, alt. 1270-1360 hexap., 
(Prov. Mex. Guanaxuatensi), floret Septembri, 
Humboldt & Bonpland s.n. (HT: P]. T-Specim.: | of 1. 
Mexico: Guanajuato: in montanis prope Santa Rosa, 
Humboldt & Bonpland s.n. {[HT: P; IT: US-91925 
(fragm. ex P-HBK Herb & photo)]. 

NOTE: as "Podosaemum" mucronatum. 
= Muhlenbergia mucronata (Kunth) Trin. 


Podosemum porteri (Scribn. ex Beal) Bush, Amer. Mid]. 
Naturalist 7(2): 36. 1921. Basionym: Muhlenbergia 
porteri Scribn. ex Beal. 

= Muhlenbergia porteri Scribn. ex Beal. 


Podosemum pungens (Thurb. ex A. Gray) Bush, Amer. Midl. 
Naturalist 7(2): 32. 1921. Basionym: Muhlenbergia 
pungens Thurb. ex A. Gray. 

= Muhlenbergia pungens Thurb. ex A. Gray. 


190 


Catalogue of New World Grasses: II. 


P. Beauv., Ess. 
1812. Basionym: 


Podosemum purpureum (P. Beauv.) 
Agrostogr. 176, 179, t. 8, f. 2. 
Trichochloa purpurea P. Beauv. 

= Muhlenbergia expansa (Poir.) Trin. 


Podosemum quadridentatum Kunth, Nov. Gen. Sp. 1: 130. 
1815 [1816]. T-Protol.: Crescit in siccis apricis regni 
Mexicani prope Toluca, Puente de Tepare et Playas de 
Jorullo, inter 400 et 1300 hexap. Floret Septembri, 
Humboldt & Bonpland s.n. T-Specim.: 1 of 1. Mexico: 
México: near Toluca, no date, Humboldt & AJA. 
Bonpland s.n. [LT: P; IT: US-2557456, US-86634 
(fragm. ex P), US-86635], LT designated by ?; cited 
by McVaugh, Fl. Novo-Gal. 14: 253 (1983). 

NOTE: as "Podosaemum" quadridentatum. 
= Muhlenbergia quadridentata (Kunth) Trin. 


Podosemum reverchonii (Vasey & Scribn.) Bush, Amer. 
Midl. Naturalist 7(2): 38. 1921.  Basionym: 
Muhlenbergia reverchonti Vasey & Scribn. 

= Muhlenbergia reverchonii Vasey & Scribn. 


Podosemum rigidum Kunth, Nov. Gen. Sp. 1: 129. 1815 
[1816]. T-Protol.: Crescit in subfrigidis regni Mexicani 
juxta Guanaxuato, Temascatio et Lo de Sierra, alt. 
900-1070 hexap., floret Septembri, Humboldt & 
Bonpland s.n. T-Specim.: | of |. Mexico: Guanajuato: 
near Guanajuato, Humboldt & Bonpland s.n. [HT: P; 
IT: US-91920 (fragm. ex P)]. 

NOTE: as "Podosaemum" rigidum. 
= Muhlenbergia rigida (Kunth) Kunth. 


Podosemum setosum Kunth, Nov. Gen. Sp. 1: 129. 1815 
[1816]. T-Protol.: Mexico: Crescit in alta planitie 
Mexicana inter Gueguetoque et Tula, alt. 1100 hexap., 
floret Augusto, Humboldt & Bonpland s.n. {HT: P]. T- 
Specim.: | of 1. Mexico: Gueguetoque & Tula, 
Humboldt & Bonpland 4174 |HT: P-HBK; IT: B-W, 
US-91917 (fragm. ex P-HBK)]. 

NOTE: as "Podosaemum" setosum. 

= Muhlenbergia microsperma (DC.) Kunth. 


Podosemum soboliferum (Muhl. ex Willd.) Link, Hort. Berol. 
1: 83. 1827. Basionym: Agrostis sobolifera Muhl. ex 
Willd. 

NOTE: as "Podosaemum". 
= Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. 


Podosemum strictum J. Presl, Relig. Haenk. 1(4-5): 230. 
1830. T-Protol.: Hab. ?, /Taenke s.n., Country of origin 
not indicated in the protologue [gd, 1995). 
NOTE: as "Podosaemum" strictum. 
= Muhlenbergia stricta (J. Pres!) Kunth. 


Podosemum tenellum Kunth, Nov. Gen. Sp. 1: 128. 1815 
[1816]. T-Protol.: Crescit in scopulosis — frigidis 
Andium Mexicanorum, inter Rio Frio et Baranca 
Honda, alt. 1220 hexap., (Prov. Xalapensi), floret 
Februario, /fumboldt & Bonpland s.n. T-Specim.: | of 
|. Mexico: Veracruz: inter Rio Frio et BArranca 
Honda, Humboldt & Bonpland s.n. |HT: P-Bonpl.; IT: 
P, US-91922 (fragm. ex P)]. 

NOTE: as "Podosaemum" tenellum. 
= Muhlenbergia tenella (Kunth) Trin. 


Podosemum tenuiflorum (Willd.) Link, Hort. Berol. 1: 82. 
1827. Basionym: Agrostis tenuiflora Willd. 
NOTE: as "Podosaemum". 


= Muhlenbergia tenuiflora (Willd.) Britton, Sterns & 
Poggenb. 


Podosemum tenuifolium (Kunth) Nees ex Nees & Schauer, 
Linnaea 19(6): 690. 1847. Basionym: Calamagrostis 
tenuifolia Kunth. 

NOTE: as "Podosaemum". 
= Muhlenbergia tenuifolia (Kunth) Kunth. 


Podosemum tenuissimum J. Presl, Relig. Haenk. 1(4-5): 230. 
1830. T-Protol.: Hab. in Panama et in Mexico, Haenke 
s.n. T-Specim.: | of |. Panama, Haenke s.n. [HT: PR; 
IT: MO-2974335, MO-2974336 (line drawing of MO 
isotype), US (fragm.)]. 
NOTE: as "Podosaemum" tenuissimum. 
= Muhlenbergia tenuissima (J. Pres!) Kunth. 


Podosemum texanum (Buckley) Bush, Amer. Midl. Naturalist 


7(2): 41. 1921. Basionym: Muhlenbergia texana 
Buckley. 

= Muhlenbergia texana Buckley. 

Podosemum trichopodes (Elliott) Bush, Amer. Midl. 
Naturalist 7; 30. 1921. Basionym: Agrostis trichopodes 
Elliott. 


NOTE: as "Podosaemum". 
= Muhlenbergia expansa (Poir.) Trin. 


Podosemum virescens Kunth, Nov. Gen. Sp. 1: 132. 1815 
[1816]. T-Protol.: Crescit locis asperis, excelsis regni 
Mexicani prope Santa Rosa de la Sierra et Puerto de 
Varientos, alt. 1350 hexap. [ca. 2400 m], floret Sep, 
Humboldt & Bonpland s.n. T-Specim.: | of 1. Mexico: 
Guanajuato: near Santa Rosa de la Sierra, no date, 
Humboldt & Bonpland s.n. [HT: P; IT: US-86633 
(fragm. ex P-HBK Herb)]. 

NOTE: as "Podosaemum" virescens. 
= Muhlenbergia virescens (Kunth) Trin. 


Podosemum virginicum (L.) Link, Hort. Berol. 1: 85. 1827. 
Basionym: Agrostis virginica L. 
NOTE: as "Podosaemum". 
= Sporobolus virginicus (L.) Kunth. 


Pogochloa S$. Moore, Trans. Linn. Soc. London, Bot. 4(3): 
509. 1895. TYPE: Pogochloa brasiliensis §. Moore. 
= Gouinia E. Fourn. ex Benth. & Hook. f. 


Pogochloa brasiliensis S. Moore, Trans. Linn. Soc. London, 
Bot. 4(3): 509, t. 37, f. 9-23. 1895. T-Protol.: Brazil: 
Matto Grosso: reperi ad Coimbra mens. Feb. 
florentem, S.L.M. Moore 1080, T-Specim.: | of 1. 
Brazil: reperi ad Coimbra, Feb 1891, Moore 1080 [HT: 
BM]. 

= Gouinia brasiliensis (S. Moore) Swallen. 


Polyodon Kunth, Nov. Gen. Sp. 1: 174. 1815 [1816]. TYPE: 
Polyodon distichum Kunth. 
= Bouteloua Lag. 


Polyodon distichum Kunth, Nov. Gen. Sp. 1: 175, t. 55. 1815 
[1816]. T-Protol.: Crescit in montosis, altis Regni 
Quitensis, Humboldt & Bonpland s.n. T-Specim.: | of 
I. Ecuador: Prov. Pichincha: Quito, Humboldt & 
Bonpland s.n. (HT: P]. 

= Bouteloua disticha (Kunth) Benth. 


Polyodon texanus (S. Watson) Nash, Fl. S.E. U.S. 138, 1327. 
1903. Basionym: Bouteloua texana S. Watson. 
= Bouteloua rigidiseta (Steud.) Hitche. 


Catalogue of New World Grasses: II. 


19] 


Polypogon brachyphyllus (Nees) E. Fourn. ex Hemsl., Biol. 
Cent.-Amer., Bot. 3(19): 552. 1885. Basionym: 
Podosemum brachyphyllum Nees. 

= Muhlenbergia ciliata (Kunth) Trin. 


Polypogon canadensis E. Fourn., Mexic. Pl. 2: 92. 1886. 
NOTE: nom. nov., nom. illeg. superfl.; based on Agrostis 
mexicana, the name changed for location excluded. 
= Muhlenbergia mexicana (L.) Trin. 


Polypogon ciliatus (Kunth) Spreng., Syst. Veg. 1: 243. 1825. 
Basionym: Podosemum ciliatum Kunth. 
= Muhlenbergia ciliata (Kunth) Trin. 


Polypogon glomeratus Willd., Enum. Pl. 87. 1809. T-Protol.: 
USA: Pennsylvania ["North America"]. T-Specim.: | 
of 1. USA: Habitat in Pensylvania, /67/ [HT: B 
(Willd. hb); IT: US-1535797 (fragm. ex Willd. hb) 
[67]]. 
= Muhlenbergia glomerata (Willd.) Trin. 


Polypogon gracilis Spreng., Syst. Veg. 5: 558 (index). 1828. 
REPLACED NAME: Polypogon tenellus (Kunth) Spreng. 
BLOCKING NAME: Polypogon tenellus R. Br. 
= Muhlenbergia tenella (Kunth) Trin. 


Polypogon racemosus (Michx.) Nutt., Gen. N. Amer. PI. 1: 
51. 1818. Basionym: Agrostis racemosa Michx. 
= Muhlenbergia racemosa (Michx.) Britton, Sterns & 
Poggenb. 


Polypogon sericeus (Michx.) Spreng., Syst. Veg. 1: 243. 
1825. Basionym: Stipa sericea Michx. 
= Muhlenbergia sericea (Michx.) P.M. Peterson. 


Polypogon setosus Biehler, Pl. Nov. Herb. Spreng. 7. 1807. 
T-Protol.: USA: E Pennsylvania, Muhlenberg s.n. T- 
Specim.: | of 1. Muhlenberg no. 107 [T: IT: PH, US- 
1535797 (fragm. ex PH-"107")]. 

= Muhlenbergia glomerata (Willd.) Trin. 


Polypogon spicatus Spreng., Syst. Veg. 1: 243. 1824. T- 
Protol.: Brasil, Se/iow s.n. T-Specim.: | of 1. Brazil, F. 
Sellow s.n. (HT: B; IT: BAA (fragm.), US (fragm. ex 
B)]. 

= Gymnopogon spicatus (Spreng.) Kuntze. 


Polypogon tenellus (Kunth) Spreng., Syst. Veg. 1: 243. 1825. 
Basionym: Podosemum tenellum Kunth. 
NOTE: hom. illeg. 
E. NAME: Polypogon tenellus R. Br. 
CORRECT NAME: Pedosemum tenellum Kunth, 
= Muhlenbergia tenella (Kunth) Trin. 


Polyraphis (Trin.) Lindl., Veg. Kingd. 115. 1847. Basionym: 
Pappophorum sect. Polyraphis Trin. 
NOTE: nom. superfl. 
= Pappophorum Schreb. 


Polyschistis J. Presl, Relig. Haenk. 1(4-5): 294. 1830. TYPE: 
Polyschistis paupercula J. Presl. 
= Bouteloua Lag. 


Pommereulleae trib. Bor, Grass. Burma, Ceylon, India & 
Pakistan 686. 1960. TYPE: Pommereulla L. f. 


Ponceletia L. M. A. A. Du Petit-Thouars, Esquisse Fl. Tristan 
D'Acugna 36. 1811. TYPE: Ponceletia arundinacea 
Thouars. 

NOTE: later hom., non R. Br. 1810. 
= Spartina Schreb. 


Pringleochloa Scribn., Bot. Gaz. 21: 137. 1896. TYPE: 
Pringleochloa stolonifera (E. Fourn.) Scribn. 
= Bouteloua Lag. 


Pringleochloa stolonifera (E. Fourn.) Scribn., Bot. Gaz. 21: 
138. 1896. Basionym: Atheropogon  stolonifer E. 
Fourn. 

= Bouteloua reederorum Columbus. 


Psammophila Schult., Syst. Veg. | Mant. 1: 231. 1822. 
REPLACED NAME: Ponceletia L. M. A. A. Du Petit- 
Thouars. 
BLOCKING NAME: Ponceletia R. Br. 
= Spartina Schreb. 


Psilantha (K. Koch) Tzvelev, Bot. Zhurn. (Moscow & 
Leningrad) 53: 311. 1968. Basionym: Poa 
UNRANKED Psilantha K. Koch. 

= Eragrostis Wolf. 


Puccinellia carinata (Griseb.) Ponert, Feddes Repert. 84(9- 
10): 739. 1974. Basionym: Atropis carinata Griseb. 
= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Rabdochloa P. Beauv., Ess. Agrostogr. 84, 176. 1812. TYPE: 
Rabdochloa domingensis (Jacq.) P. Beauv.; LT 
designated by Niles & Chase, Contr. U.S. Natl. Herb. 
24: 187 (1925). 

= Leptochloa P. Beauv. 


Rabdochloa domingensis (Jacq.) P. Beauy., Ess. Agrostogr. 
84, 176, t. 17, f. 3. 1812. Basionym: Cynosurus 
domingensis Jacq. 

= Leptochloa virgata (L.) P. Beauv. 


Rabdochloa dubia (Kunth) Kuntze ex Stuck., Anales Mus. 
Nac. Buenos Aires 11: 121. 1904. Basionym: Chloris 
dubia Kunth. 

= Leptochloa dubia (Kunth) Nees. 


Rabdochloa imbricata (Thurb.) Kuntze, Revis. Gen. Pl. 2: 
788. 1891. Basionym: Leptoch/oa imbricata Thurb. 
= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Rabdochloa mucronata (Michx.) P. Beauy., Ess. Agrostogr. 
84, 158, 176. 1812. Basionym: Chloris mucronata 
Michx. 

= Leptochloa 
Nowack. 


panicea subsp. mucronata (Michx.) 


Rabdochloa spicata (Nees) Kuntze ex Stuck., Anales Mus. 
Nac. Buenos Aires 11: 121. 1904. Basionym: Bromus 
spicatus Nees. 

= Tripogon spicatus (Nees) Ekman. 


Rabdochloa virgata (Sw.) P. Beauv., Ess. Agrostogr. 84, 158. 
1812. Basionym: Chloris virgata Sw. 
= Chloris virgata Sw. 


Raddia Mazziari, lonios Antologia 2: 448. 1834. TYPE: 
Raddia aculeata Mazziari. 
NOTE: later hom., non Bertol. 1819. 
BLOCKING NAME: Raddia Bertol. 
= Crypsis Aiton. 


Raspailia rupestris Turcz. ex Kom., Fl. URSS 2: 297. 1934. 
NOTE: nom. inval., as syn. Pappophorum boreale Griseb. 
= Enneapogon desyauxii P. Beauy. 


192 


Catalogue of New World Grasses: II. 


Redfieldia Vasey, Bull. Torrey Bot. Club 14: 133. 
1887. TYPE: Redfieldia flexuosa (Thurb. ex A. Gray) 
Vasey. 

TRTD. BY: Peterson, P. M. 
NHT: Muhlenbergiinae. NO. ACCPTD. SPP.: 1. 
CNSLTD. LIT.: Reeder, J. R., (1976). 


Redfieldia flexuosa (Thurb. ex A. Gray) Vasey, Bull. Torrey 
Bot. Club 14: 133. 1887. Basionym: Graphephorum 
flexuosum Thurb. ex A. Gray. 

DST.: USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Cronquist, A., A. H. 
Holmgren, N. H. Holmgren & J. L. Reveal (1977); 
Fernald, M. (1950); Great Plains Flora Association 
(1986); Hitchcock, A. S. (1951); Reeder, J. R. 
(1976). 

SYN.: Graphephorum flexuosum Thurb. ex A. Gray. 


Reederochloa Soderstr. & HF. Decker, Brittonia 
16(3): 334. 1964. TYPE: Reederochloa eludens 
Soderstr. & H.F. Decker. 

TRTD. BY: Peterson, P. M. 
NHT: Monanthochloinae. NO. ACCPTD. SPP.: 1. 
CNSLTD. LIT.: Soderstrom, T. R. & H. F. Decker, (1964). 


Reederochloa eludens Soderstr. & H.F. Decker, Brittonia 
16(3): 335, f. 1-10. 1964. T-Protol.: Mexico: Durango: 
1 mi S of Durango, alkali flats, common in dry heavily 
trodden path, 6200 ft, 7.R. Soderstrom 798 [HT: US- 
2378386; IT: K, MEXU, YU]. T-Specim.: 1 of 1. 
Mexico: Durango: | mi S of Durango, alkali flats, 
common in dry heavily trodden path, 6200 ft, 29 Sep 
1959, T.R. Soderstrom 798 [HT: US-2378386; IT: K, 
MEXU, YU]. 
DST.: Mexi. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Soderstrom, T. R. & H. F. Decker (1964). 


Rendlia Chiov., Ann. Bot. (Rome) 13: 53. 1914. TYPE: 
Rendlia obtusifolia Chiov. 
= Microchloa R. Br. 


Rhabdochloa cruciata (L.) P. Beauv., Ess. Agrostogr. 84, 
158, 176. 1812. Basionym: Agrostis cruciata L. 
= Chloris cruciata (L.) Sw. 


Rhabdochloa longa (Griseb.) Kuntze, Revis. Gen. Pl. 2: 788. 
1891. Basionym: Leptochloa longa Griseb. 
= Leptochloa longa Griseb. 


Rhabdochloa rigida (Munro ex Vasey) Kuntze, Revis. Gen. 
Pl. 2: 788. 1891. Basionym: Leptochloa rigida Munro 
ex Vasey. 

= Eragrostis sessilispica Buckley. 


Rhabdochloa virgata (Sw.) P. Beauv., Ess. Agrostogr. 84, 
158, 176. 1812. Basionym: Chloris virgata Sw. 
= Chloris virgata Sw. 


Rhachidospermum mexicanum Vasey, Bot. Gaz. 15(5): 106, 
110, t. 12. 1890. T-Protol.: Mexico: Lower California: 
La Paz, FE. Palmer s.n. T-Specim.: | of 1. Mexico: 
Baja California Sur: La Paz, 1890, £. Palmer 124 [IT: 


US-1015504, 
1015503}. 
= Jouvea pilosa (J. Presl) Scribn. 


US-1015505, US-1015498, — ~US- 


Rheochloa Filg., P.M. Peterson & Y. Herrera, Syst. 
Bot. 24(2): 123-127, f. 1-3. 1999. TYPE: Rheochloa 
scabriflora Filg., P.M. Peterson & Y. Herrera. 

TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: I. 


Rheochloa seabriflora Filg., P.M. Peterson & Y. Herrera, 
Syst. Bot. 24(2): 123-127, f. 1-3. 1999. T-Protol.: 
Brazil: Goias: Municipio de Mineiros: Parque 
Nacional de Emas, 18°17'5S5"S, 52°43'79"W, Campo 
Umido (humid grassland), erva decumbendente 
(decumbent grass), colmos flexuosas (culms flexuous), 
apoiandose em outras plantas (supported by other 
plants), inflorescéncia) avermelhada — (reddish 
inflorescence), rara (rare), apenas duas plantas vistas 
no meio da vegetacdo (only two plants seen in the 
vegetation),1 Apr 1994, Filgueiras 2854 [HT: IBGE; 
IT: MO (fragm.), US]. T-Specim.: 1 of 1. Brazil: 
Goias: Municipio de Mineiros: Parque Nacional de 
Emas, 18°17'55"S, 52°43'79"W, Campo Umido (humid 
grassland), erva decumbendente (decumbent grass), 
colmos flexuosas (culms flexuous), apoiandose em 
outras plantas (supported by other _ plants), 
inflorescéncia avermelhada (reddish inflorescence), 
rara (rare), apenas duas plantas vistas no meio da 
vegetacdo (only two plants seen in the vegetation), | 
Apr 1994, 1 Apr 1994, Filgueiras 2854 [HT: IBGE; 
IT: MO-04575749 (fragm., photocopy), US]. 

DST.: Braz. 

LVL. ACCPTC.: 4. 

ALSO ACCPTD. BY: Filgueiras, T. S., P. M. Peterson & 
Y. Herrera-Arrieta (1999), 


Rhombolytrum albescens (Vasey) Nash, Man. Fl. N. States 
129. 1901. Basionym: Triodia albescens Vasey. 
= Tridens albescens (Vasey) Wooton & Stand. 


Roshevitzia Tzvelev, Bot. Zhurn. (Moscow & Leningrad) 53: 
311. 1968. TYPE: Roshevitzia diarrhena (Schult. & 
Schult. f.) Tzvelev. 

= Eragrostis Wolf. 


Roshevitzia diplachnoides (Steud.) Tzvelev, Novosti Sist. 
Vyss. Rast. 7: 50. 1970 [1971]. Basionym: Eragrostis 
diplachnoides Steud. 

= Eragrostis japonica (Thunb.) Trin. 


Roshevitzia glomerata (Walter) Tzvelev, Novosti Sist. Vyss. 
Rast. 7: 50. 1970 [1971]. Basionym: Poa glomerata 
Walter. 

= Eragrostis japonica (Thunb.) Trin. 


Roshevitzia japonica (Thunb.) Tzvelev, Novosti Sist. Vyss. 
Rast. 7: 50. 1970[1971]. Basionym: Poa japonica 
Thunb. 

= Eragrostis japonica (Thunb.) Trin. 


Rottboellia paniculata (Nutt.) Spreng., Syst. Veg. 1: 300. 
1825. Basionym: Lepturus paniculatus Nutt. 
= Schedonnardus paniculatus (Nutt.) Trel. 


Rottboellia scorpioides Poir. ex Steud., Nomencl. Bot. (ed. 2) 
2: 474. 1841. 


Catalogue of New World Grasses: II. 


193 


NOTE: nom. inval., as syn. of Ctenium americanum 
Sprengel. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Rottboellia setacea Roxb., Pl. Coromandel 2: 17, t. 132. 
1798. T-Specim.: 1 of |. India: coast of Coromandel, 
Roxburgh s.n. {IT: K]. 

NOTE: nom. illeg. superfl., cited Nardus indica L.f. as syn. 
CORRECT NAME: Microchloa indica (L. f.) P. Beauv. 
= Microchloa indica (L. f.) P. Beauv. 


Saccharum pappiferum Lam., Tabl. Encycl. 1: 155. 1791. T- 
Protol.: ex Amer. merid., D. Richard s.n. T-Specim.: | 
of 2. South America, Richard s.n. [HT: P-LAM, IT: 
BAA(fragm.), US-76121 (fragm. ex P-LAM)]. 2 of 2. 

= Pappophorum pappiferum (Lam.) Kuntze. 


Saugetia Hitche. & Chase, Contr. U.S. Natl. Herb. 
18(7): 378. 1917 Aug 18. TYPE: Saugetia fasciculata 
Hitche. & Chase. 

TRTD. BY: Peterson, P.M. NO. ACCPTD. SPP.: 2. 


Saugetia fasciculata Hitchc. & Chase, Contr. U.S. Natl. 
Herb. 18(7): 378. 1917. T-Protol.: Cuba: collected in 
small wood, savanna San Julian, south of Guane, 
Province of Pinar del Rio, 28 Dec 1916,Leon 690. T- 
Specim.: | of 1. Cuba: Pinar del Rio: in small wood, 
savanna San Julian, south of Guane, 28 Dec 1916,Bro. 
Leon & M. Roca 6901 |HT: US-950204; IT: NY- 
71123). 

DST.: Cari. 

LVL. ACCPTC.:: I. 

ALSO ACCPTD. BY: Catasus 
Hitchcock, A. S. (1936). 

SYN.: Enteropogon fasciculatus Clayton & Renvoize. 


Guerra, L. (1997); 


Saugetia pleiostachya Hitchc. & Ekman, Man. Grasses W. 
Ind. 123, 1936. T-Protol.: Cuba: Prov. Pinar del Rio: 
collected on jurassic limestone, top of Cerro de 
Mendoza, Mendoza, 26 June 1920,E.L. Ekman 11483 
[HT: US-1296127]. T-Specim.: 1 of 1. Cuba: Pinar del 
Rio: Mendoza Mun., 26 Jun 1920,E.L. Ekman 11483 
(HT: US-1296127; IT: NY-71124]. 

DST.: Cari. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Catasus 
Hitchcock, A. S. (1936). 


Schaffnera Benth., Hooker's Icon. Pl. 14: 59, t. 1378. 1882. 
TYPE: Schaffnera gracilis Benth. 
NOTE: later near hom., non Schaffneria Moore 1857. 
= Schaffmerella Nash. 


Guerra, L. (1997); 


Schaffnera gracilis Benth., Hooker's Icon. Pl. 14: 59 pl. 1378. 
1882. T-Specim.: | of 2. Mexico: San Luis Potosi, 
1879, LG. Schaffner 1070 [ST: US-397116], CS. 2 of 
2. Mexico: San Louis Potosi, 1876,/.G. Schaffner 134 
[IST: US-825687]. 
= Schaffnerella gracilis (Benth.) Nash. 


Schaffnerella Nash, N. Amer. Fl. 17(2): 141. 1912. 
REPLACED NAME: Schaffnera Benth. 
BLOCKING NAME: Schaffneria Fée. 
TRTD. BY: Peterson, P. M. 
NHT: Muhlenbergiinae. NO. ACCPTD. SPP.: 1. 
SYN.: Schaffnera Benth. 


Schaffnerella gracilis (Benth.) Nash, N. Amer. FI. 17(2): 
141. 1912. Basionym: Schaffnera gracilis Benth. 
DST.: Mexi. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000). 
SYN.: Schaffnera gracilis Benth. 


Schedonnardus Steud., Syn. Pl. Glumac. 1: 146. 
1854 ("1855"). TYPE: Schedonnardus texanus Steud. 
TRTD. BY: Peterson, P. M. 
NHT: Muhlenbergiinae. NO. ACCPTD. SPP.: 1. 
SYN.: Spirochloe Lunell. 


Schedonnardus paniculatus (Nutt.) Trel., Annual Rep. 
Geol. Surv. Arkansas 1888(4): 236. 1891. Basionym: 
Lepturus paniculatus Nutt. 

DST.: Arge, Cana, Mexi, USA. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Burkart, A. (1969); Cabrera, A. L. 
(1970); Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Fernald, M. (1950); Gleason, H. A. 
& A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Great Plains Flora Association 
(1986); Hitchcock, A. S. (1951); Hitchcock, C. L., A. 
Cronquist, M. Ownbey & J. W. Thompson (1969); 
Moss, E. H. (1983); Scoggan, H. J. (1978 [1979]); 
Zuloaga, F. O., E. G. Nicora, Z. E. Ruigolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

Lepturus paniculatus Nutt.; Rottboellia paniculata 
(Nutt.) Spreng.; Schedonnardus texanus Steud.; 
Spartina dissitiflora Steud.; Spirochloe paniculata 
(Nutt.) Lunell. 


SYN.: 


Schedonnardus texanus Steud., Syn. Pl. Glumac. |: 146. 
1854. 
= Schedonnardus paniculatus (Nutt.) Trel. 


Schellingia Steud., Flora 33: 231. 1850. TYPE: Schellingia 
tenera Steud. 
= Aegopogon Humb. & Bonpl. ex Willd. 


Schellingia tenera Steud., Flora 33: 232. 1850. T-Protol.: 
Mexico, Galeotti herb. no. 5750. 
= Aegopogon tenellus (DC.) Trin. 


Schenckochloa barbata (Hack.) J.J. Ortiz, Candollea 46(2): 
243. 1991. Basionym: Diplachne barbata Hack. 
= Gouinia barbata (Hack.) Swallen. 


Schismus patens J. Presl, Relig. Haenk. 1(4-5): 269. 1830. T- 
Protol.: Habitat in Cordilleris chilensibus, Haenke s.n. 
T-Specim.: 1 of 1. Chile: 7) Haenke s.n. [T: US- 
A78816], CS. 

= Leptochloa dubia (Kunth) Nees. 


Schleropelta Buckley, Prelim. Rep. Surv. Texas App. I. 
1866. TYPE: Schleropelta stolonifera Buckley. 
= Hilaria Kunth. 


Schleropelta stolonifera Buckley, Prelim. Rep. Surv. Texas 
App. |. 1866. 
= Hilaria belangeri (Steud.) Nash. 


Schultesia Spreng., Pl. Min. Cogn. Pug. 2: 17. 1815. 
NOTE: nom. rejic. et nom. superfl. pro Eustachys. 


194 


Catalogue of New World Grasses: II. 


BLOCKING NAME: Schultesia Mart. 
= Eustachys Desv. 


Schultesia petraea (Sw.) Spreng., Pl. Min. Cogn. Pug. 2: 17. 
1815. Basionym: Chloris petraea Sw. 
= Eustachys petraea (Sw.) Desv. 


Sciadonardus Steud., Flora 33: 229. 1850. 
NOTE: nom. nud. 
= Gymnopogon P. Beauv. 


Sciadonardus distichophyllus Steud., Flora 33: 229. 1850. 
NOTE: nom. inval., as syn. of Gymnopogon distichophyllus 
Steudel. 
= Gymnopogon ambiguus (Michx.) Britton, Sterns & 
Poggenb. 


Scleropogon Phil., Anales Univ. Chile 36: 205. 1870. 
TYPE: Scleropogon brevifolius Phil. 

TRTD. BY: Peterson, P. M. 

NHT: Cynodonteae. NO. ACCPTD. SPP.: 1. 

CNSLTD. LIT.: Anton, A. M., H. E. Connor & M. E. 
Astegiano, (1998); Peterson, P. M. & J. T. Columbus, 
(1997). 

SYN.: Lesourdia E. Fourn. 


Scleropogon brevifolius Phil., Anales Univ. Chile 36: 206. 
1870. T-Protol.: Mendoza, Chile, is this the same as 
Mendoza Argentina today?. 

DST.: Arge, Chil, Mexi, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Hickman, J. C. (1993); Hitchcock, 
A. S. (1951); Marticorena, C. & M. Quezada (1985); 
McVaugh, R. (1983); Shreve, F. & I. L. Wiggins 
(1964); Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

Festuca macrostachya Torr. & A. Gray; Lesourdia 
karwinskyana E. Fourn.; Lesourdia multiflora E. 
Fourn.; Scleropogon karwinskyanus (E.  Fourn.) 
Benth. ex S. Watson; Scleropogon longisetus Beetle. 


SYN.: 


Scleropogon karwinskyanus (E. Fourn.) Benth. ex S. Watson, 
Proc. Amer. Acad. Arts 18: 181. 1883. Basionym: 
Lesourdia karwinskyana E. Fourn. 

= Scleropogon brevifolius Phil. 


Scleropogon longisetus Beetle, Phytologia 49(1): 42. 1981. 


T-Protol.: Mexico: Coahuila[HT: YU]. T-Specim.: | 
of 1. Mexico: Coahuila: 28 mi S of Saltillo, along road 
to Concepcion del Oro, J. Reeder & C. Reeder 3626 
[HT: YU]. 

= Scleropogon brevifolius Phil. 


Sericrostis Raf., Neogenyton 4. 1825. TYPE: Stipa sericea 
Michx.; LT designated by Pfeiffer, Nom. Bot. 2: 1142 
(1874). 

= Muhlenbergia Schreb. 


Sesleria dactyloides Nutt., Gen. N. Amer. Pl. 1: 65. 1818. T- 
Specim.: | of 1. USA: plains of the Missouri River, 
Nuttall s.n. {T]. 

= Bouteloua dactyloides (Nutt.) Columbus. 


Sieglingia acuminata (Benth. ex Vasey) Kuntze, Revis. Gen. 
Pl. 2: 789. 1891. Basionym: Triodia acuminata Benth. 
ex Vasey. 

= Erioneuron pilosum (Buckley) Nash. 


Sieglingia albescens (Vasey) Kuntze, Revis. Gen. Pl. 2: 789. 
1891. Basionym: Triodia albescens Vasey. 
= Tridens albescens (Vasey) Wooton & Stand. 


Sieglingia ambigua (Elliott) Kuntze, Revis. Gen. Pl. 2: 789. 
1891. Basionym: Poa ambigua Elliott. 
= Tridens ambiguus (Elliott) Schult. 


Sieglingia americana (P. Beauv.) Beal, Grass. N. Amer. 2: 
466. 1896. Basionym: Triplasis americana P. Beauv. 
= Triplasis americana P. Beauv. 


Sieglingia argentina Kuntze, Revis. Gen. Pl. 3(2): 367. 1898. 
T-Specim.: 1 of 1. Argentina: Rio Negro: Rio 
Colorado, Lorentz s.n. [HT: LP; IT: CORD]. 

= Erioneuron pilosum var. longearistatum (Kurtz) Anton. 


Sieglingia avenacea (Kunth) Kuntze, Revis. Gen. Pl. 2: 789. 
1891. Basionym: Triodia avenacea Kunth. 
= Erioneuron avenaceum (Kunth) Tateoka. 


Sieglingia avenacea var. grandiflora (Vasey) L.H. Dewey, 
Contr. U.S. Natl. Herb. 2(3): 538. 1894. Basionym: 
Triodia grandiflora Vasey. 

= Erioneuron avenaceum (Kunth) Tateoka. 


Sieglingia buckleyana Vasey ex L.H. Dewey, Contr. U.S. 
Natl. Herb. 2(3): 540. 1894. T-Specim.: | of 1. USA: 
southern Texas, 1883, Buckley s.n. [IT: US-81808]. 

= Tridens buckleyanus (Vasey ex L.H. Dewey) Nash. 


Sieglingia chapmanii Small, Bull. Torrey Bot. Club 22(8): 
365. 1895. T-Specim.: | of 1. Florida, Chapman [T]. 
NOTE: as "Chapmani". 
= Tridens flavus var. chapmanii (Small) Shinners. 


Sieglingia congesta L.H. Dewey, Contr. U.S. Natl. Herb. 
2(3): 538. 1894. T-Specim.: 1 of 1. USA: Texas, 
Nueces Co., May 1891, G.C. Nealley 24 [IT: US- 
81809]. 

= Tridens congestus (L.H. Dewey) Nash. 


Sieglingia cuprea (Jacq.) Millsp., Bull. West Virginia Agric. 
Exp. Sta. 24(2): 471. 1892. Basionym: Triodia cuprea 
Jacq. 
= Tridens flavus (L.) Hitchc. 


Sieglingia dubia (Kunth) Kuntze ex Stuck., Anales Mus. Nac. 
Buenos Aires 11: 128. 1904. Basionym: Chloris dubia 
Kunth. 

= Leptochloa dubia (Kunth) Nees. 


Sieglingia elongata (Buckley) Nash, Ill. Fl. N. U.S. 3: 504. 
1898. Basionym: Uralepis elongata Buckley. 
= Tridens muticus var. elongatus (Buckley) Shinners. 


Sieglingia eragrostoides (Vasey & Scribn.) L.H. Dewey, 
Contr. U.S. Natl. Herb, 2(3): 539. 1894. Basionym: 
Triodia eragrostoides Vasey & Scribn. 

= Tridens eragrostoides (Vasey & Scribn.) Nash. 


Sieglingia eragrostoides var. scabra Vasey ex Beal, Grass. 
N. Amer. 2: 465. 1896. T-Specim.: | of 1. USA: 
Texas, Aug 1892, G.C. Nealley 96 [IT: US-908296], 
CS. 

= Tridens eragrostoides (Vasey & Scribn.) Nash. 


Catalogue of New World Grasses: II. 


195 


Sieglingia flava (L.) Kuntze, Revis. Gen. Pl. 2: 789. 1891. 
Basionym: Poa flava L. 
= Tridens flavus (L.) Hitche. 


Sieglingia grandiflora (Vasey) Beal, Grass. N. Amer. 2: 471. 
1896. Basionym: Triodia grandiflora Vasey. 
= Erioneuron avenaceum (Kunth) Tateoka. 


Sieglingia latifolia (Griseb.) Kuntze, Revis. Gen. Pl. 2: 789. 
1891. Basionym: Tricuspis latifolia Griseb. 
= Gouinia latifolia (Griseb.) Vasey. 


Sieglingia mutica (Torr.) Kuntze, Revis. Gen. Pl. 2: 789, 
1891. Basionym: Tricuspis mutica Torr. 
= Tridens muticus (Torr.) Nash. 


Sieglingia nealleyi (Vasey) L.H. Dewey, Contr. U.S. Natl. 
Herb. 2(3): 538. 1894. Basionym: Triodia nealleyi 
Vasey. 

= Erioneuron nealleyi (Vasey) Tateoka. 


Sieglingia paraguayensis Kuntze, Revis. Gen. Pl. 3(2): 341. 
1898. 
NOTE: nom. altern. Arundinaria paraguayensis Kuntze, 
same page. 
= Gouinia paraguayensis (Kuntze) Parodi. 


Sieglingia pilosa (Buckley) Nash, Ill. Fl. N. U.S. 3: 504. 
1898. Basionym: Uralepis pilosa Buckley. 
= Erioneuron pilosum (Buckley) Nash. 


Sieglingia pulchella (Kunth) Kuntze, Revis. Gen. Pl. 2: 789. 
1891. Basionym: Triodia pulchella Kunth. 
= Dasyochloa pulchella (Kunth) Willd. ex Rydb. 


Sieglingia pulchella var. parviflora Vasey ex Beal, Grass. N. 
Amer. 2: 468. 1896. T-Protol.: Western Texas. T- 
Specim.: | of 1. USA: California: southern California, 
Orcutt s.n. (HT: US-908213 (fragm., TAES)]. 

= Dasyochloa pulchella (Kunth) Willd. ex Rydb. 


Sieglingia purpurea (Walter) Kuntze, Revis. Gen. Pl. 2: 789. 
1891. Basionym: Aira purpurea Walter. 
= Triplasis purpurea (Walter) Chapm. 


Sieglingia schaffneri (S. Watson) Kuntze, Revis. Gen. Pl. 2: 
789. 1891. Basionym: Triodia schaffneri S. Watson. 
= Tripogon spicatus (Nees) Ekman. 


Sieglingia sesleroides (Michx.) Scribn., Mem. Torrey Bot. 
Club 5(4): 48. 1894. Basionym: Poa sesleroides 
Michx. 

= Tridens flavus (L.) Hitche. 


Sieglingia sesleroides var. intermedia Vasey ex L.H. Dewey, 
Contr. U.S. Natl. Herb. 2(3); 539. 1894. T-Specim.: 1 
of 1. USA: Texas: Indian Territory, Aug 1891, C.S. 
Sheldon 273 {IT: US-1611435], Portion from a sheet 
sent to Gray Herb. 
= Tridens flavus (L.) Hitche. 


Sieglingia spicata (Nees) Kuntze ex Stuck., Anales Mus. 
Nac. Buenos Aires 11: 128. 1904. Basionym: Bromus 
spicatus Nees. 

= Tripogon spicatus (Nees) Ekman. 


Sieglingia stricta (Nutt.) Kuntze, Revis. Gen. Pl. 2: 789. 
1891. Basionym: Windsoria stricta Nutt. 
= Tridens strictus (Nutt.) Nash. 


Sieglingia texana (S. Watson) Kuntze, Revis. Gen. Pl. 2: 789. 
1891. Basionym: Triodia texana S. Watson. 
= Tridens texanus (S. Watson) Nash. 


Sieglingia wrightii Vasey, Contr. U.S. Natl. Herb. 1(8): 269. 
1893. T-Specim.: | of 3. USA: Texas: Valley of the 
limpio, 1851-1852,C. Wright 2038 [ST: US-82080]. 2 
of 3. USA: Texas: Presidio Co., 1891, G.C. Nealley 
s.n. [ST: US]. 3 of 3. USA: Texas: Presidio Co., 1892, 
G.C. Nealley s.n. (ST: US]. 

= Allolepis texana (Vasey) Soderstr. & H.F. Decker. 


Soderstromia C.V. Morton, Leafl. W. Bot. 10: 327. 1966. 
REPLACED NAME: Fourniera Scribn. 
BLOCKING NAME: Fourniera J. Bommer ex E. Fourn. 
= Bouteloua Lag. 


Soderstromia mexicana (Scribn.) C.V. Morton, Leafl. W. 
Bot. 10(16): 327. 1966. Basionym: Fourniera 
mexicana Scribn. 

= Bouteloua mexicana (Scribn.) Columbus. 


Sohnsia Airy Shaw, Kew Bull. 18; 272. 1965. 
BLOCKING NAME: Calamochloe Rchb. 
TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 1. 
SYN.: Calamochloa E. Fourn.; Eufournia Reeder. 


Sohnsia filifolia (E. Fourn.) Airy Shaw, Kew Bull. 18: 272. 
1965. Basionym: Calamochiloa filifolia E. Fourn. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000). 

SYN.: Calamochloa /filifolia E. Fourn.; Eufournia filifolia 
(E. Fourn.) Reeder. 


Solenachne Steud., Syn. Pl. Glumac. 1: 12. 10-12 Dec 1853 
("1855"). TYPE: Solenachne phalaroides Steud. 
= Spartina Schreb. 


Solenachne ciliata (Brongn.) Herter, Revista Sudamer. Bot. 
6(5-6): 147. 1940. Basionym: Spartina ciliata Brongn. 
= Spartina ciliata Brongn. 


Solenachne phalaroides Steud., Syn. Pl. Glumac. 1: 12. 1853. 
T-Protol.: Brazil: Ins. St. Catharinae Brasil, Urville. 
= Spartina ciliata Brongn. 


Solenophyllum Baill., Hist. Pl. 12: 235. 1894. 
NOTE: in syn. sub Monanthochloe. 
= Monanthochloe Engelm. 


Spartina Schreb., Gen. Pl. 43. 1789. TYPE: Spartina 
cynosuroides (L.) Roth. 

TRTD. BY: Peterson, P. M. 

NHT: Cynodonteae. NO. ACCPTD. SPP.: 15. 

CNSLTD. LIT.: Mobberley, D. G., (1956). 

SYN.: Chauvinia Steud.; Limnetis Rich. in Persoon; 
Ponceletia LL. M. A. A. Du Petit-Thouars; 
Psammophila Schult.; Solenachne Steud.; 
Trachynotia Michx.; Tristania Poir. 


Spartina alterniflora Loisel., Fl. Gall. 719. 1807. T-Protol.: 
France: Bayonne. T-Specim.: | of 1. Sul da Franca 
{T]. 

DST.: Arge, Braz, Cana, Cari, FrGu, Guya, Suri, USA, 
Urug. 


196 


Catalogue of New World Grasses: II. 


LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Cabrera, A. L. (1970); Correll, D. S. 
& M. C. Johnston (1970); Edgar, E., H. E. Connor & 
J. E. Shand (1991); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Fernald, M. 
(1950); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Godfrey, R. 
K. & J. W. Wooten (1979); Gould, F. W (1979); 
Hickman, J. C. (1993); Hitchcock, A. S. (1951); 
Hitchcock, C. L., A. Cronquist, M. Ownbey & J. W. 


Thompson (1969); Judziewicz, E. J. (1990); Long, R. 


W. & O. Lakela (1971); Mobberley, D. G. (1956); 
Nicora, E. G., M. E. De Paula, A. M. Faggi, M. del 
C. Mariano, A. M. Miante A., L. R. Parodi, C. A. 
Petetin, F. A. Roig & Z. Rugolo de Agrasar (1978); 
Radford, A. E., H. E. Ahles & C. R. Bell (1968); 
Renvoize, S. A. (1984); Renvoize, S. A. (1988); 
Scoggan, H. J. (1978 [1979]); Small, J. K. (1933); 
Smith, L. B., D. C. Wasshausen & R. M. Klein 
(1981); Wunderlin, R. P. (1998); Zuloaga, F. O., E. 
G. Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

Dactylis fasciculata C. Sm.; Dactylis maritima 
Walter; Limnetis glabra (Muhl. ex Elliott) Eaton & J. 
Wright; Limnetis glabra Nutt.; Spartina alterniflora 
var. glabra (Muhl. ex Elliott) Fernald; Spartina 
alterniflora var. pilosa (Merr.) Fernald; Spartina 
brasiliensis Raddi; Spartina glabra Muhl.; Spartina 
glabra Muhl. ex Elliott; Spartina glabra var. 
alterniflora (Loisel.) Merr.; Spartina glabra var. 
pilosa Merr.; Spartina laevigata Bosc ex Link; 
Spartina maritima subvar. pilosa (Merr.) St.-Yves; 
Spartina maritima subvar. radii St.-Yves; Spartina 
maritima var. alterniflora (Loisel.) St.-Yves; 
Spartina maritima vat. brasiliensis (Raddi) St.-Yves; 
Spartina maritima var. glabra (Muhl. ex Elliott) St.- 
Yves; Spartina merrillii A. Chev., Spartina stricta 
var. alterniflora (Loisel.) A. Gray; Spartina stricta 
var. glabra (Muhl. ex Elliott) A. Gray; Spartina 
stricta var. maritima (Walter) Scribn.; Trachynotia 
alterniflora (Loisel.) DC.; Trachynotia alterniflora 
Steud. 


SYN.: 


Spartina alterniflora var. glabra (Muhl. ex Elliott) Fernald, 
Rhodora 18: 178. 1916. Basionym: Spartina glabra 
Muhl. ex Elliott. 

= Spartina alterniflora Loisel. 


Spartina alterniflora var. pilosa (Merr.) Fernald, Rhodora 18: 
179. 1916. Basionym: Spartina glabra var. pilosa 
Merr. 
= Spartina alterniflora Loisel. 


Spartina anglica C.E. Hubb., Grasses 359. 1968. 
NOTE: nom. inval., without Latin description. 
= Spartina anglica C.E. Hubb. 


Spartina anglica C.E. Hubb., Bot. J. Linn. Soc. 76(4): 364. 
1978. ‘T-Protol.: England: W. Sussex: Bosham, 
fringing shore on mud-flats and muddy shingle, 
extending into water of Chishester Harbour, forming 
extensive masses, 17 Aug 1868, CE. Hubbard 
D1I7868A [HT: K]. T-Specim.: | of 1. Great Britain: 
England: Sussex, 17 Aug 1968,C.E. Hubbard 17868A 
(HT: K; IT: US-2907471]. 

DST.: USA. 
LVL. ACCPTC.: 1; introduced. 


ALSO ACCPTD. BY: Edgar, E., H. E. Connor & J. E,. 
Shand (1991). 

SYN.: Spartina anglica C.E. Hubb.; Spartina townsendii 
var. anglica (C.E. Hubb.) Lambinon & Maquet. 


Spartina argentinensis Parodi, Revista Fac. Agron. Veterin. 
(Buenos Aires) 2: 258. 1919. 
= Spartina spartinae (Trin.) Merr. ex Hitche. 


Spartina bakeri Merr., U.S.D.A. Bur. Pl. Industr. Bull. 9: 
14. 1902. T-Specim.: | of 1. USA: Florida, 19 Apr 
1898, C.H. Baker 14 (IT: US-81737]. 
DST.: USA. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Anonymous (1986); Godfrey, R. kK. 
& J. W. Wooten (1979); Hitchcock, A. S. (1951); 
Long, R. W. & O. Lakela (1971); Small, J. K. (1933); 
Wunderlin, R. P. (1998). 
SYN.: Spartina juncea var. bakeri(Merr.) St.-Y ves. 


Spartina brasiliensis Raddi, Agrostogr. Bras. 21. 1823. T- 
Protol.: Brasilia: in inundatis prope Rio-Janeiro, Raddi 
s.n. T-Specim.: | of 1. Brazil: Rio de Janeiro: Rio de 
Janeiro, Raddi s.n. [HT: PI (2 sheets); IT: US-3376186 
(fragm. ex Pisa)]. 
= Spartina alterniflora Loisel. 


Spartina caespitosa A.A. Eaton, Bull. Torrey Bot. Club 25: 
338. 1898. T-Protol.: USA: New Hampshire: 
Seabrook, 4.4. Eaton s.n. T-Specim.: | of lL. USA: 
New Pachughan Co.: Hampshire: Hampton Falls, 26 
Aug 1896, 4.A. Eaton s.n. [LT: NEBC; ILT: US- 
1763051], LT designated by McDonnell & Crow, 
Rhodora 81: 125 (1979). 

CORRECT NAME: Spartina x caespitosa A.A. Eaton (pro 
sp.). 
= Spartina x caespitosa A.A. Eaton (pro sp.). 


Spartina x caespitosa A.A. Eaton (pro sp.), Bull. Torrey 
Bot. Club 25: 338. 1898. 

NOTE: [S. patens x S. pectinata|, hyb. stat. given by 
Fernald, Gray's Man. Bot. ed 8., 180 (1950). 

DST.: USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Fernald, M. 
(1950). 

SYN.: Spartina caespitosa A.A. Eaton; Spartina patens var. 
caespitosa (A.A. Eaton) Hitche. 


Spartina capensis Nees ex Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 111. 1840. T-Protol.: South Africa: Cbsp. 
[Cape Good Hope]. T-Specim.: | of 1. [HT: LE-TRIN- 
2055.01]. 

= Spartina maritima (Curtis) Fernald. 


Spartina ciliata Brongn., Voy. Monde 2(2): 15, t. 2. 1829. T- 
Protol.: Brazil: recueillie dans l'ile de Santa Catarina. 
T-Specim.: | of |. Brazil: Ilha de Santa Catarina, 
D'Urville 1304 [IT: P; IT: US-s.n. (fragm. ex P), US- 
878777 (fragm. ex P)]. 

DST.: Arge, Braz, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Cabrera, A. L. (1970); Mobberley, 
D. G. (1956); Renvoize, S. A. (1988); Rosengurtt, B., 
B. R. Arrillaga de Maffei & P. Izaguirre de Artucio 
(1970); Smith, L. B., D. C. Wasshausen & R. M. 
Klein (1981). 


Catalogue of New World Grasses: II. 


197 


SYN.: Solenachne ciliata (Brongn.) Herter; Solenachne 
phalaroides Steud.; Spartina coarctata Trin. 


Spartina coarctata Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 110. 1840. T-Protol.: Montivideo[HT: LE (ex 
B)], "perperum s.n. Spart. brasiliensis Radd.". T- 
Specim.: | of 1. [HT: LE-TRIN-2056.01]. 

= Spartina ciliata Brongn. 


Spartina cynosuroides (L.) Roth, Catal. Bot. 3: 10. 1806. 
Basionym: Dactylis cynosuroides L. 

DST.: Cari, Mexi, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Chase, A. (1929); Correll, D. S. & M. C. 
Johnston (1970); Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Fernald, M. 
(1950); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Godfrey, R. 
K. & J. W. Wooten (1979); Hitchcock, A. S. (1936); 
Hitchcock, A. S. (1951); Radford, A. E., H. E. Ahles 
& C. R. Bell (1968); Small, J. K. (1933); Wunderlin, 
R. P. (1998). 

Cynodon cynosuroides (L.) Raspail; Dactylis 
cynosuroides L.; Limnetis cynosuroides (L.) Rich.; 
Limnetis polystachia (Michx.) Rich.; Paspalum 
cynosuroides (L.) Brot.; Spartina cynosuroides x 
gracilis St.-Yves; Spartina  cynosuroides var. 
polystachya (Michx.) Beal; Spartina polystachya 
(Michx.) P. Beauv.; Trachynotia cynosuroides (L.) 
Michx.; Trachynotia polystachya Michx. 


SYN.: 


Spartina cynosuroides var.aureo-marginata W. Irving, Gard. 
Chron. 38(987): 372. 1905. T-Protol.: Received at 
Kew from the New York Botanical Garden in 1904. 
= Spartina pectinata Link. 


Spartina cynosuroides fo. major St.-Yves, Candollea 5: 61, 
62. 1932. T-Protol.: no type desiginated, 
NOTE: Published as S. cynosuroides var. Michauxiana fo. 
major. 
= Spartina pectinata Link. 


Spartina cynosuroides var. michauxiana (Hitche.) St.-Yves, 
Candollea 5: 58, t. 3, f. a-7. 1932. Basionym: Spartina 
michauxiana Hitche. 

= Spartina pectinata Link. 


Spartina cynosuroides var. polystachya (Michx.) Beal, Grass. 
N. Amer. 2: 398. 1896. Basionym: Trachynotia 
polystachya Michx. 

= Spartina cynosuroides (L.) Roth. 


Spartina cynosuroides x gracilisSt.-Y ves, Candollea 5: 66, t. 
4, f. b10. 1932. 
= Spartina cynosuroides (L.) Roth. 


Spartina densiflora Brongn., Voy. Monde 2(2): 14. 1829. T- 
Protol.: Chile: Concepcion, D'Urville s.n. T-Specim.: 1 
of 1. D’'Urville s.n. (IT: US-610991 (fragm. ex P), US- 
92018 (fragm. ex P & photo)]. 

DST.: Arge, Braz, Chil, Urug. 

LVL. ACCPTC.: |. 

ALSO ACCPTD. BY: Burkart, A. (1969); Cabrera, A. L. 
(1970); Hickman, J. C. (1993); Marticorena, C. & M. 
Quezada (1985); Nicora, E. G., M. E. De Paula, A. 
M. Faggi, M. del C. Mariano, A. M. Miante A., L. R. 
Parodi, C. A. Petetin, F. A. Roig & Z. Rugolo de 


Agrasar (1978): Renvoize, S. A. (1988); Smith, L. B., 
D. C. Wasshausen & R. M. Klein (1981); Zuloaga, F. 
O., E. G. Nicora, Z. E. Rugolo de Agrasar, O. 
Morrone, J. Pensiero & A. M. Cialdella (1994). 

Chauvinia chilensis Steud.; Spartina densiflora 
subvar. pauper St.-Yves; Spartina juncea_ var. 
laxiflora St.-Y ves; Spartina juncea var.montevidense 
(Arechav.) St.-Yves; Spartina —montevidensis 
Arechav.; Spartina patagonica Speg. 


SYN.: 


Spartina densiflora fo. acuta St.-Yves, Candollea 5: 76, 81. 
1932. T-Specim.: 1 of 1. USA: California: Humboldt 
Co., salt marsh at Eureka, 21 July 1924, A.A. Heller 
13781 (IT: US-1287859], specimen drawn by Chase in 
Manual as S. foliosa. 
= Spartina foliosa Trin. 


Spartina densiflora subvar. argentinensis (Parodi) St.-Yves, 
Candollea 5: 26, 79, t. 7, f. b-17. 1932. Basionym: 
Spartina argentinensis Parodi. 

NOTE: var. junciformis St. Yves. 
= Spartina spartinae (Trin.) Merr. ex Hitche. 


Spartina densiflora var.junciformis (Engelm. & A. Gray) St.- 
Yves, Candollea 5: 26, 77, t. 7, f. 16. 1932. Basionym: 
Spartina junciformis Engelm. & A. Gray. 

= Spartina spartinae (Trin.) Merr. ex Hitchc. 


Spartina densiflora var. obtusa Hack., Repert. Spec. Nov. 
Regni Veg. 6(21-26): 345. 1909. T-Protol.: Paraguay: 
Gran Chaco: in arenosis Loma Clavel, flor mens. Nov, 
Hassler 2621. T-Specim.: | of 1. Paraguay: Gran 
Chaco: Loma Clavel, 1903, Hassler 262/ [IT: US- 
92017(fragm. ex Vienna)]. 
= Spartina spartinae (Trin.) Merr. ex Hitchc. 


Spartina densiflora subvar. pauper St.-Yves, Candollea 5: 77. 
1932. 
NOTE: nom. inval.; as syn. of S. patagonica Speg. 
= Spartina densiflora Brongn. 


Spartina dissitiflora Steud., Syn. Pl, Glumac. 1: 215. 1854. 
T-Protol.: Bahia, Sa/zmann s.n. T-Specim.: | of 1. 
Brazil: Bahia: in aguis subsaltis, Sa/zmann s.n. (HT: 
P?: IT: MO-2108466]. 

NOTE: nom. illeg. superfl. 
CORRECT NAME: Rottboellia paniculata (Nutt.) Spreng. 
= Schedonnardus paniculatus (Nutt.) Trel. 


Spartina foliosa Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 114. T-Specim.: | of 1. California, 1831, D. 
Peters s.n. (IT: US-610993(fragm. ex LE), US-92016 
(fragm.)]. 

DST.: Mexi, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Abrams, L. (1923); Beetle, A. A. 
(1977); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. & R. Moran 
(1981); Hickman, J. C. (1993); Hitchcock, A. S. 
(1951); Mobberley, D. G. (1956); Munz, P. A. & D. 
D. Keck (1959); Munz, P. A. (1974); Shreve, F. & I. 
L. Wiggins (1964). 

Spartina densiflora fo. acuta St.-Yves, Spartina 
leiantha Benth. 


SYN.: 


Spartina glabra Muhl., Cat. Pl. Amer. Sept. 8. 1813. 
NOTE: nom. nud. 
= Spartina alterniflora Loisel. 


198 


Catalogue of New World Grasses: II. 


Spartina glabra Muhl. ex Elliott, Sketch Bot. S. Carolina 1: 
95, t. 4, f 2. 1816. T-Protol.: Grows in places 
overflowed with salt water at every tide, forming 
exclusively the 'salt water marsh’, 

= Spartina alterniflora Loisel. 


Spartina glabra var. alterniflora (Loisel.) Merr., U.S.D.A. 
Bur. Pl. Industr. Bull. 9: 9. 1902. Basionym: Spartina 
alterniflora Loisel. 

= Spartina alterniflora Loisel. 


Spartina glabra var. pilosa Merr., U.S.D.A. Bur. Pl. Industr. 
Bull. 9: 9. 1902. T-Specim.: | of 1. USA: New Jersey, 
Aug 1895, FLL. Scribner s.n. [IT: US-81736]. 
= Spartina alterniflora Loisel. 


Spartina gouini E. Fourn., Mexic. Pl. 2: 135. 1886. T-Protol.: 
Mexico: Veracruz, Gouin 72. T-Specim.: | of 1. 
Mexico: Veracruz, 1867, Gouin 72 {[IT: US- 
92015(fragm. ex P)]. 

= Spartina spartinae (Trin.) Merr. ex Hitchce. 


Spartina gracilis Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 110. 1840. T-Protol.: Amer. bor., Hooker 
s.n., “Hooker s.n. Sp. [artina] cynosuroides". T- 
Specim.: | of 1. Hooker s.n. [IT: LE-TRIN-2058.01, 
US-92014(fragm. ex LE)]. 

DST.: Cana, Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Abrams, L. (1923); Cronquist, A., A. 
H. Holmgren, N. H. Holmgren & J. L. Reveal (1977); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Gleason, H. A. & A. Cronquist 
(1991); Great Plains Flora Association (1986); 
Hickman, J. C. (1993); Hitchcock, A. S. (1951); 
Hitchcock, C. L., A. Cronquist, M. Ownbey & J. W. 
Thompson (1969); Moss, E. H. (1983); Munz, P. A. 
& D. D. Keck (1959); Munz, P. A. (1974); Porsild, 
A. E. & W. Cody (1980); Scoggan, H. J. (1978 
[1979}). 


Spartina intermedia Bosc ex St.-Y ves, Candollea 5: 50. 1932. 
NOTE: nom. inval., as syn. of Spartina maritima. 
= Spartina maritima (Curtis) Fernald, 


Spartina juncea (Michx.) Willd, Enum. Pl, 81. 1809, 
Basionym: Trachynotia juncea Michx. 


= Spartina patens (Aiton) Muhl. 


Spartina Juncea subvar. americana St.-Yves, Candollea 35: 
27, 84, t. 8, f. b-20. 1932. T-Protol.: no type cited. 
NOTE: Published as S. juncea var. juncea  subvar. 
americana, 
= Spartina patens (Aiton) Muhl. 


Spartina juncea var. bakeri (Merr.) St.-Yves, Candollea 5: 
27, 91, t. 9, f. c. 1932. Basionym: Spartina bakeri 
Merr. 

= Spartina bakeri Merr. 


Spartina juncea var. laxiflora St.-Yves, Candollea 5: 88. 
1932. T-Protol.: Dusen 23. 
= Spartina densiflora Brongn. 


Spartina juncea var. montevidense (Arechav.) St.-Yves, 
Candollea 5: 89. 1932. Basionym: Spartina 
montevidensis Arechav. 

= Spartina densiflora Brongn. 


Spartina juncea var. patens (Aiton) St.-Yves, Candollea 5: 
27, 86. 1932. Basionym: Dactylis patens Aiton. 
= Spartina patens (Aiton) Muhl. 


Spartina junciformis Engelm. & A. Gray, Boston J. Nat. Hist. 
5: 238. 1845. T-Protol.: Saline prairies near the coast, 
May. T-Specim.: | of 1. USA: Texas: saline prairies 
near the coast, Lindheimer 207 (HT: GH; IT: BM]. 
= Spartina spartinae (Trin.) Merr. ex Hitche. 


Spartina laevigata Bosc ex Link, Jahrb. Gewachsk. 1(3): 92. 
1820. T-Protol.: Nord-Amerika, Bosc s.n. [B-W]. T- 
Specim.: | of 1. North America, Link s.n. [IT: US- 
878539 (fragm. ex LE)]. 

= Spartina alterniflora Loisel. 


Spartina leiantha Benth., Bot. Voy. Sulphur 56. 1844. T- 
Protol.: Mexico: Baja California Sur: Bay of 
Magdalena[BM]. T-Specim.: | of |. Mexico: Baja 
California Sur: Bay of Magdalena [IT: US-92013 
(fragm. ex BM)]. 

= Spartina foliosa Trin. 


Spartina longispica Hauman & Parodi ex St-Yves, 
Candollea 5: 27, 92. 1932. T-Specim.: 1 of 3. 
Argentina, Parodi 5014 [ST: P; US-92012 (fragm. ex 
BAA)]. 2 of 3. Argentina, Parodi 9567 [ST: P]. 3 of 3. 
Argentina: Buenas Aires: Pipinas, 24 Mar 1923, L. 
Parodi 5030 [LT: US(fragm. ex SI)], LT selected by 
Parodi in letter to A. Chase & published in Index to 
Grass Species vol. 3: 336. 1962. 

NOTE: originally described as hybrid [Spartina brasiliensis 
(alterniflora) x S. montevidensis (densiflora)| but 
then recognized as species by Parodi in Revista 
Sudam. Bot. Montevideo 4: 177-178 (1937). 

DST.: Arge, Urug. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Cabrera, A. L. (1970); Nicora, E. G., 
M. E. De Paula, A. M. Faggi, M. del C. Mariano, A. 
M. Miante A., L. R. Parodi, C. A. Petetin, F. A. Roig 
& Z. Rigolo de Agrasar (1978); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 


Spartina maritima (Curtis) Fernald, Rhodora 18: 180. 1916. 
Basionym: Dactylis maritima Curtis. 

DST.: USA. 

LVL. ACCPTC.: |; introduced. 

ALSO ACCPTD. BY: Gibbs Russell, G. E, W. G. Welman, 
E. Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987). 

SYN.: Dactylis maritima Curtis; Spartina capensis Nees ex 
Trin.; Spartina intermedia Bosc ex St.-Yves; 
Spartina stricta (Curtis) Roth. 


Spartina maritima var. alterniflora  (Loisel.) St.-Yves, 
Candollea 5: 25, 53, t. 2, f. a-4. 1932. Basionym: 
Spartina alterniflora Loisel. 

= Spartina alterniflora Loisel. 


Spartina maritima var. brasiliensis (Raddi) St.-Yves, 
Candollea 5: 56, t. 2, f. 56. 1932. Basionym: Spartina 
brasiliensis Raddi. 

= Spartina alterniflora Loisel. 


Catalogue of New World Grasses: II. 


199 


Spartina maritima var. glabra (Muhl. ex Elliott) St.-Yves, 
Candollea 5: 24, 49, t. 1, f. b-2. 1932. Basionym: 
Spartina glabra Muhl. ex Elliott. 

= Spartina alterniflora Loisel. 


Spartina maritima subvar. pilosa (Merr.) St.-Yves, Candollea 
5: 51, t. 1, f. c-3. 1932. Basionym: Spartina glabra 
var. pilosa Merr. 

= Spartina alterniflora Loisel. 


Spartina maritima subvar. radii St.-Yves, Candollea 5: 56, t. 
2, f. 0-5. 1932. 
NOTE: subsp. glabra (Muhl.) St. Yves var. brasiliensis St. 
Yves. 
= Spartina alterniflora Loisel. 


Spartina merrillii A. Chev., Bull. Soc. Bot. France 80: 787, t. 
8, f. 3. 1933. 
NOTE: as Spartina x “Merrillii"; [Spartina glabra x S. 
polystachya]. 
= Spartina alterniflora Loisel. 


Spartina michauxiana Hitche., Contr. U.S. Natl. Herb, 12(3): 
153. 1908. T-Protol.: USA: Illinoensis. Hateurs des 
terres, Michaux s.n. [HT: P]. 

= Spartina pectinata Link. 


Spartina michauxiana var. suttiei Farw., Rep. Michigan 
Acad. Sci. 21: 352. 1920. T-Protol.: USA: Michigan: 
Orchard Lake, Farwell 1487a. 

= Spartina pectinata Link. 


Spartina michauxiana var. tenuior Farw., Rep. Michigan 
Acad. Sci. 21: 352. 1920. T-Protol.: USA: Michigan: 
River Rouge, Farwell 5/38. 
= Spartina pectinata Link. 


Spartina montevidensis Arechav., Anales Mus. Nac. 
Montevideo |: 378, t. 43. 1896. T-Protol.: Vive en las 
orillas del rio de la Plata, en parajes cenagosos 
ordinariamente. 

= Spartina densiflora Brongn. 


Spartina multiflora Vasey ex Beal, Grass. N. Amer. 2: 400. 


1896. 
NOTE: nom. inval., as syn. of Spartina junciformis Engelm. 
& A. Gray. 


= Spartina spartinae (Trin.) Merr. ex Hitche. 


Spartina patagonica Speg., Revista Fac. Agron. Univ. Nac. 
La Plata 3: 579. 1897. T-Protol.: Hab. Abunde in 
uliginosis inundatis martimis Isla de los Leones ad 
ostia Rio Santa Cruz, Feb. 1882, Spegazzini s.n. 

= Spartina densiflora Brongn. 


Spartina patens (Aiton) Muhl., Descr. Gram. 55. 1817. 
Basionym: Dactylis patens Aiton. 

DST.: Cana, Cari, Mexi, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Anonymous 
(1986); Beetle, A. A. (1977); Catasus Guerra, L. 
(1997); Correll, D. S. & M. C. Johnston (1970); 
Fernald, M. (1950); Gleason, H. A. & A. Cronquist 
(1968); Godfrey, R. K. & J. W. Wooten (1979); 
Gould, F. W (1979); Hickman, J. C. (1993); 
Hitchcock, A. S. (1936); Hitchcock, A. S. (1951); 
Liogier, H. & L. Martorell (1982); Long, R. W. & O. 
Lakela (1971); Munz, P. A. (1968); Radford, A. E., 
H. E. Ahles & C. R. Bell (1968); Scoggan, H. J. 


(1978 [1979]); Small, J. K. (1933); Voss, E. G. 
(1972); Wunderlin, R. P. (1998). 

Dactylis patens Aiton; Spartina juncea (Michx.) 
Willd.; Spartina juncea subvar. americana St.-Y ves; 
Spartina juncea var. patens (Aiton) St.-Yves; 
Spartina patens var. juncea (Michx.)  Hitchc.; 
Spartina patens var. monogyna (M.A. Curtis) 
Fernald; Spartina platensis Mez ex St.-Yves; 
Spartina pumila Roth; Trachynotia juncea Michx. 


SYN.: 


Spartina patens var. caespitosa (A.A. Eaton) Hitchc., 
Rhodora 8(95): 210. 1906. Basionym: Spartina 
caespitosa A.A, Eaton. 

= Spartina x caespitosa A.A. Eaton (pro sp.). 


Spartina patens var.juncea (Michx.) Hitchc., Rhodora 8(95): 
210. 1906. Basionym: 7rachynotia juncea Michx. 
= Spartina patens (Aiton) Muhl. 


Spartina patens var. monogyna (M.A. Curtis) Fernald, 
Rhodora 49(580): 114. 1947. Basionym: Limnetis 
Juncea var. monogyna M.A. Curtis. 

= Spartina patens (Aiton) Muhl. 


Spartina pectinata Link, Jahrb. Gewachsk. 1(3): 92-93. 
1820. T-Protol.: Aus Nordamerika, Bosc s.n. [B-W]. 
T-Specim.: | of 1. USA, Bose s.n. [HT: B; IT: US- 
92011 (fragm. ex B)]. 

DST.: Cana, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Correll, D. S. & M. C. Johnston 
(1970); Cronquist, A., A. H. Holmgren, N. H. 
Holmgren & J. L. Reveal (1977); Darke, R. (1999); 
Fernald, M. (1950); Gleason, H. A. & A. Cronquist 
(1968); Gleason, H. A. & A. Cronquist (1991); 
Godfrey, R. K. & J. W. Wooten (1979); Great Plains 
Flora Association (1986); Hitchcock, A. S. (1951); 
Hitchcock, C. L., A. Cronquist, M. Ownbey & J. W. 
Thompson (1969); Moss, E. H. (1983); Porsild, A. E. 
& W. Cody (1980); Radford, A. E., H. E. Ahles & C. 
R. Bell (1968); Schwegman, John E. (1991); 
Scoggan, H. J. (1978 [1979]). 

Spartina cynosuroides fo. major St.-Yves; Spartina 
cynosuroides var. aureo-marginata W. Irving; 
Spartina cynosuroides var. michauxiana (Hitchc.) 
St.-Yves; Spartina michauxiana Hitche.; Spartina 
michauxiana var. ——suttiei ~=Farw.; ~~ Spartina 
michauxiana var. tenuior Farw., Spartina pectinata 
fo. pectinata, Spartina pectinata fo. variegata Vict.; 
Spartina pectinata var. suttiei(Farw.) Fernald. 


SYN:: 


Spartina pectinata Link fo. pectinata. 
= Spartina pectinata Link. 


Spartina pectinata var. suttiei (Farw.) Fernald, Rhodora 35: 
260. 1933. Basionym: Spartina michauxiana vat. 
suttiei Farw. 

= Spartina pectinata Link. 


Spartina pectinata fo. variegata Vict., Nat. Canada 71: 209. 
1944. T-Protol.: Canada: Que: Ile Parrot sur le lac St. 
Louis, 12 Jul 1941, Victorin & Rolland 56603 [HT: 
MT]. 

= Spartina pectinata Link. 


Spartina pittieri Hack., Oesterr. Bot. Z. 52: 237. 1902. T- 
Specim.: 1 of 2. Costa Rica: Atlantic, pr. Limon, 
Pittier 4209 [ST: US-3412973 (fragm. ex W)]. 2 of 2. 
Guatemala, Friedrichstahl s.n. [ST]. 


200 


Catalogue of New World Grasses: II. 


NOTE: aS Spartina "Pittieri" 
= Spartina spartinae (Trin.) Merr. ex Hitche. 


Spartina platensis Mez ex St.-Yves, Candollea 5: 86. 1932. 
NOTE: nom. nud. 
= Spartina patens (Aiton) Muhl. 


Spartina polystachya (Michx.) P. Beauv., Ess. Agrostogr. 25, 
178, 179. 1812. Basionym: Trachynotia polystachya 
Michx. 

= Spartina cynosuroides (L.) Roth. 


Spartina pumila Roth, Catal. Bot. 3: 10. 1806. T-Protol.: 
Habitat circa Neuyorck. 
= Spartina patens (Aiton) Muhl. 


Spartina spartinae (Trin.) Merr., U.S.D.A. Bur. Pl. Industr. 
Bull. 9: 11. 1902. Basionym: Vilfa spartinae Trin. 
NOTE: nom. inval., not accepted and in syn. of Spartina 
Junciformis Engelm. & A. Gray. 
CORRECT NAME: Spartina spartinae (Trin.) Merr. ex 
Hitche. 
= Spartina spartinae (Trin.) Merr. ex Hitchce. 


Spartina spartinae (Trin.) Merr. ex Hitche., Contr. U.S. 
Natl. Herb. 17(3): 329. 1913. Basionym: Vilfa 
spartinae Trin. 

REPLACED NAME: Spartina spartinae (Trin.) Merr. 

DST.: Arge, Beli, Cari, CoRi, Hond, Mexi, Pana, Para, 

USA, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Balick, M. J., 
M. H. Nee & D. E. Atha (2000); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Cowan, C. P. (1983); Davidse, G., M. Sousa S. & A. 
O. Chater (editores generales) (1994); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Godfrey, R. K. & J. W. Wooten (1979); Hitchcock, 
A. S. (1951); Long, R. W. & O. Lakela (1971); 
Mobberley, D. G. (1956); Pohl, R. W. (1980); Pohl, 
R. W. (1994): Small, J. K. (1933); Sousa S., M. & E. 
F. Cabrera C. (1983); Swallen, J. R. (1955); 
Wunderlin, R. P. (1998). 

Spartina argentinensis Parodi; Spartina densiflora 
subvar. argentinensis (Parodi) St.-Yves; Spartina 
densiflora var. junciformis (Engelm. & A. Gray) St.- 
Yves; Spartina densiflora var. obtusa  Hack.; 
Spartina gouini E. Fourn.; Spartina junciformis 
Engelm. & A. Gray; Spartina multiflora Vasey ex 
Beal; Spartina pittieri Hack.; Spartina spartinae 
(Trin.) Merr.; Vilfa spartinae Trin. 


SYN.: 


Spartina stricta (Curtis) Roth, Neue Beytr. Bot. 101. 1802. 
Basionym: Dactylis maritima Curtis. 
= Spartina maritima (Curtis) Fernald. 


Spartina stricta var. alterniflora (Loisel.) A. Gray, Manual 
(ed. 2) 552. 1856. Basionym: Spartina alterniflora 
Loisel. 

= Spartina alterniflora Loisel. 


Spartina stricta var. glabra (Muhl. ex Elliott) A. Gray, 
Manual (ed. 2) 552. 1856. Basionym: Spartina glabra 
Muhl. ex Elliott. 

= Spartina alterniflora Loisel. 


Spartina stricta var, maritima (Walter) Scribn., Mem. Torrey 
Bot. Club 5: 45. 1894. Basionym: Dactylis maritima 
Walter. 


NOTE: isonym. 

E. NAME: Spartina stricta. var. maritima (Walter) Britton, 
Sterns & Poggenb. 

= Spartina alterniflora Loisel. 


Spartina x townsendii H. Groves & J. Groves, Rep. Bot. 
Exch. Club 37, 1881. 
DST.: USA. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Anonymous (1986); Edgar, E., H. E. 
Connor & J. E. Shand (1991). 


Spartina townsendii var. anglica (C.E. Hubb.) Lambinon & 
Maquet, Nouv. Fl. Belgique, Luxembourg, N. France 
(ed. 3) 923. 1983. Basionym: Spartina anglica C.E. 
Hubb. 
= Spartina anglica C.E. Hubb. 


Spartineae trib. Trin., Gram. Unifl. Sesquifl. 115. 1824 [late 
Oct-1 Nov]. TYPE: Lygeuwm Loefl. ex L. 
NOTE: nom. inval., Art. 19.1, based on Lygeum sparteum 
not Spartina Schreb., as "Spartea". 


Spartininae subtrib. Maire & Weiler, Fl. Afrique N. 2: 211. 
1953. 
NOTE: nom. inval., without Latin description. 


Spermachiton Llanos, Fragm. Pl. Filip. 25. 1851. TYPE: 
Spermachiton involutum Llanos. 
= Sporobolus R. Br. 


Spermachiton involutum Llanos, Fragm. Pl. Filip. 25. 1851. 
T-Protol.: Philippines: Calumpit, Boguion road, Feb. 
T-Specim.: | of 1. Philippines: Luzon I. Manila, Dec 
1914, ED. Merrill sp. Blancoan. 845 (NEOTYPE: 
US-904537]. 

= Sporobolus diandrus (Retz.) P. Beauv. 


Spirochloe Lunell, Amer. Midl. Naturalist 4: 220. 1915. 
NOTE: nom superfl. pro Schedonnardus. 


Spirochloe paniculata (Nutt.) Lunell, Amer. Midl. Naturalist 
4: 220. 1915. Basionym: Lepturus paniculatus Nutt. 
= Schedonnardus paniculatus (Nutt.) Trel. 


Sporoboleae trib. Stapf, Fl. Cap. 7: 315. 1898. 


Sporobolinae subtrib. Benth. J. Linn. Soc., Bot. 19: 
30. 1881. TYPE: Sporobolus R. Br. 
TRTD. BY: Soreng, R. J., G. Davidse, T. S. Filgueiras, E. J. 
Judziewicz, P. M. Peterson, and F. O. Zuloaga. 
NHT: Cynodonteae. 


Sporobolus Rk. Br, Prodr. 169. 1810. TYPE: 
Sporobolus indicus (L.) R. Br.; LT designated by 
L.K.G. Pfeiffer, Nom. Bot. 2:1274 (1874), also by 
Nash, III. Fl. N. US. (ed. 2) 1: 194 (1913). 

TRTD. BY: Peterson, P. M., S. C. Boechat, & J. F. 

Veldkamp. 

NHT: Sporobolinae. NO. ACCPTD SPP.: 77 SUBSP.: 13. 

CNSLTD. LIT.: Stent, S. M., (1927); Riggins, R., (1977); 
Simon, B. K. & S. W. L. Jacobs, (1999). 

SYN.: Agrosticula Raddi; Bauchea E. Fourn.; Bennetia 
Raf.; Cryptostachys Steud., Diachyrium Griseb.; 
Spermachiton Llanos; Triachyrum Hochst. ex A. 
Braun. 


Catalogue of New World Grasses: II. 


201 


Sporobolus acuminatus (Trin.) Hack., Repert. Spec. Nov. 
Regni Veg. 9: 373. 1909. Basionym: Vilfa acuminata 
Trin. 

DST.: Arge, Braz, Para. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 

Sporobolus acuminatus (Trin.) Kuhlm.; Vilfa 
acuminata Trin. 


SYN.: 


Sporobolus acuminatus (Trin.) Kuhlm., Comm. Lin. Telegr., 
Bot. 67(Bot. 11): 92. 1922. Basionym:Vilfa acuminata 
Trin. 
NOTE: isonym. 
E. NAME: Sporobolus acuminatus (Trin.) Hack. 
= Sporobolus acuminatus (Trin.) Hack. 


Sporobolus adustus (Trin.) Roseng., B.R. Arill. & Izag., 
Gram. Urug. 270, f. 112. 1970. Basionym: Vilfa adusta 
Trin. 
DST.: Arge, Braz, Para, Urug. 
LVL. ACCPTC.:: 1. 
ALSO ACCPTD. BY: Rosengurtt, B., B. R. Arrillaga de 
Maffei & P. Izaguirre de Artucio (1970). 


Sporobolus aeneus (Trin.) Kunth, Enum. Pl. 1: 213. 1833. 
Basionym: Vilfa aenea Trin. 

DST.: Arge, Boli, Braz, Para, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Burkart, A. (1969); Filgueiras, T. S. 
(1991); Filgueiras, T. S. (1999); Foster, R. C. (1958); 
Hitchcock, A. S. (1927); Longhi-Wagner, H. M. 
(1990); Renvoize, S. A. (1984); Renvoize, S. A. 
(1988); Renvoize, S. A. (1998); Smith, L. B., D. C. 
Wasshausen & R. M. Klein (1981); Zuloaga, F. O., 
E. G. Nicora, Z. E. Rugolo de Agrasar, O. Morrone, 
J. Pensiero & A. M. Cialdella (1994). 

Sporobolus aeneus var. barbicollis Hack.; 
Sporobolus aeneus fo. parviflora Parodi; Sporobolus 
aeneus var. subbulbosus  (Arechav.) — Parodi; 
Sporobolus aeneus var. typica Parodi; Sporobolus 
eximius (Nees ex Trin.) Ekman; Sporobolus 
sporobolus (Spreng.) Kuntze; Sporobolus sprengelii 
Kunth; Sporobolus subbulbosus Arechav.;, Vilfa 
adusta Trin.; Vilfa aenea Trin.; Vilfa aenea var. 
angustifolia Doll, Vilfa aenea var. latifolia Déll; 
Vilfa eximia Nees ex Trin. 


SYN:: 


Sporobolus aeneus var. barbicollis Hack., Bull. Herb. 
Boissier, ser. 2, 4(3): 278. 1904. T-Protol.: Paraguay: 
in campis pr. Tobaty, Sep, Hassler 6278. T-Specim.: | 
of |. [HT: W; IT: US (ex W)]. 

= Sporobolus aeneus (Trin.) Kunth. 


Sporobolus aeneus fo. parviflora Parodi, Revista Fac. Agron. 
Veterin. (Buenos Aires) 6(2): 161. 1928. 
= Sporobolus aeneus (Trin.) Kunth. 


Sporobolus aeneus var. subbulbosus (Arechav.) Parodi, 
Revista Fac. Agron. Veterin. (Buenos Aires) 6(2): 161, 
f. 16. 1928. Basionym: Sporobolus subbulbosus 
Arechav. 
= Sporobolus aeneus (Trin.) Kunth. 


Sporobolus aeneus var. typica Parodi, Revista Fac. Agron. 
Veterin. (Buenos Aires) 6(2): 160, 1. 11,1, f. 16A-B. 
1928. 


NOTE: nom. inval., see ICBN Art. 24.3. 
= Sporobolus aeneus (Trin.) Kunth. 


Sporobolus affinis Kunth, Revis. Gramin. 1: 68. 1829. T- 
Protol.: West Indies: Antillae. 
NOTE: hom. illeg. superfl.; based on Agrostis pyramidata 
Lam. 
= Sporobolus pyramidatus (Lam.) Hitche. 


Sporobolus airoides (Torr.) Torr., Pacif. Railr. Rep. Parke, 
Bot. 7: 21. 1856. Basionym: Agrostis airoides Torr. 

DST.: Cana, Mexi, USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Abrams, L. (1923); Bor, N. L. 
(1960); Correll, D. S. & M. C. Johnston (1970); 
Darke, R. (1999); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Fernald, M. 
(1950); Gleason, H. A. & A. Cronquist (1968); 
Gould, F. W. & R. Moran (1981); Great Plains Flora 
Association (1986); Hickman, J. C. (1993); 
Hitchcock, A. S. (1951); Hitchcock, C. L., A. 
Cronquist, M. Ownbey & J. W. Thompson (1969); 
McVaugh, R. (1983); Shreve, F. & I. L. Wiggins 
(1964). 

Agrostis airoides Torr.; Sporobolus diffusissimus 
Buckley; Sporobolus tharpii Hitche.; Vilfa airoides 
(Torr.) Trin. ex Steud. 


SYN.: 


Sporobolus airoides (Torr.) Torr. subsp.. airoides. 
LVL. ACCPTC.: 1. 


Sporobolus airoides (Torr.) Torr. var. airoides. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Cronquist, A., 
A. H. Holmgren, N. H. Holmgren & J. L. Reveal 
(1977); Gleason, H. A. & A. Cronquist (1991); 
Gould, F. W. & R. Moran (1981); Munz, P. A. & D. 
D. Keck (1959). 

SYN.: Sporobolus schaffneri Mez. 


Sporobolus airoides var. minor (Vasey) Beetle, Phytologia 
54(1): 5. 1983. Basionym: Sporobolus altissimus var. 
minor Vasey. 

= Sporobolus wrightii Munro ex Scribn. 


Sporobolus airoides subsp. regis (1.M. Johnst.) Wipff & 
S.D. Jones, Sida 16(1): 164. 1994. Basionym: 
Sporobolus regis 1.M. Johnst. 

DST.: Mexi. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000). 

SYN.: Sporobolus regis 1.M. Johnst. 


Sporobolus airoides var. wrightii (Munro ex Scribn.) Gould, 
Madrofio 10: 94. 1949. Basionym: Sporobolus wrightii 
Munro ex Scribn. 

= Sporobolus wrightii Munro ex Scribn. 


Sporobolus altissimus Vasey, Proc. Calif. Acad. Sci., ser. 2, 
2: 212. 1889. T-Protol.: USA: California: San Diego, 
Dr. E. Palmer s.n. (HT: US]. T-Specim.: | of 1. USA: 
California: San Diego Co., 1888, E. Palmer s.n. \IT: 
US-82015]. 
= Sporobolus wrightii Munro ex Scribn. 


Sporobolus altissimus var. minor Vasey, Proc. Calif. Acad. 
Sci., ser. 2, 2: 213. 1889. T-Protol.: Mexico: Baja 


202 


Catalogue of New World Grasses: II. 


California: San Enrique, Brandegee [HT: US]. T- 
Specim.: | of 1. Mexico: Baja California: San Enrique, 
4 May 1889, 7.S. Brandegee s.n. [IT: US-998328]. 

= Sporobolus wrightii Munro ex Scribn. 


Sporobolus angustus Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 88-89. 1862. T-Protol.: USA: Texas: 
Buchanan County: Jun., Buckley s.n. T-Specim.: | of 
1. USA: Texas: Buchanan, Jun, $B. Buckley s.n. [HT: 
PH; IT: US-611005 (neg. & fragm. ex PH)], 
Apparently there is no Buchanan Co. in Texas. 

= Sporobolus indicus (L.) R. Br. 


Sporobolus annuus Vasey, Bull. Torrey Bot. Club 14: 9. 


1887.  T-Protol.: Mexico: Chihuahua, — 1885, 
Dr.E. Palmer 4a (HT: US]. T-Specim.: 1 of 1. Mexico: 
Chihuahua: Noragachi, 1885, Palmer 4a [HT: US- 
82016; IT: K, NY-19971 [Palmer 4]]. 

= Muhlenbergia annua (Vasey) Swallen. 


Sporobolus apiculatus Boechat & Longhi-Wagner, Bradea 
6(20): 173-177, f. 1. 1993. T-Protol.: Brasil: Distrito 
Federal: Brasilia: 22 Oct 1990, Brochado & Filgueiras 
90 [HT: ICN; IT: IBGE]. 

DST.: Braz. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Boechat, S. C., and H. M. Longhi- 
Wagner (1993). 


Sporobolus arenaceus Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 89. 1862. T-Protol.: USA: Texas: Hill 
sides, Western Texas. T-Specim.: | of 1. USA: New 
Mexico / Texas, May 1849-Oct 1849,C.H. Wright 737 
[LT: GH; ILT: US-997371], LT designated by 
Hitchcock, Man. Grass. U.S. 886 (1935), but without 
citing a specific sheet in a specific herbarium. 

= Muhlenbergia arenacea (Buckley) Hitchc. 


Sporobolus argutus (Nees) Kunth, Enum. Pl. 1: 215. 1833. 
Basionym: Vilfa arguta Nees. 
NOTE: also Revis. Gram. 3: Suppl. 1834 p. (as in 1835 ed.) 
XVI. 
= Sporobolus pyramidatus (Lam.) Hitche. 


Sporobolus argutus fo. purpurascens Hack., Anales Mus. 
Nac. Buenos Aires 21: 90. 1911. T-Protol.: Argentina: 
Chaco: Colonia Benites, /Stuckert] 15967. T-Specim.: 
1 of |. Argentina: Chaco: Colonia Benites, Stuckert 
15967 [ST]. 

= Sporobolus pyramidatus (Lam.) Hitche. 


Sporobolus argutus var. tuberculatus (Hack.) Hack., Anales 
Mus. Nac. Buenos Aires 21: 90. 1911. Basionym: 
Sporobolus tuberculatus Hack. 

= Sporobolus pyramidatus (Lam.) Hitche. 


Sporobolus aristatus Rydb., Bull. Torrey Bot. Club 28(5): 
266. 1901. T-Protol.: USA: Wyoming: Sheridan co.: 
Big Horn Mts., F. Tweedy 2196 [HT: NY]. T-Specim.: 
1 of 1. USA: Wyoming: Sheridon Co.: Big Horn Mts., 
2438 m, Aug 1899, Tweedy 2196 [HT: NY; IT: US- 
84833 (fragm. ex NY)}. 
= Muhlenbergia filiformis (Thurb. ex S. Watson) Rydb. 


Sporobolus arizonicus Thurb. ex Vasey, Bull. Torrey Bot. 
Club 15: 8. 1888. 
NOTE: nom. inval., as syn. of Sporobolus interruptus 
Vasey. 
= Sporobolus interruptus Vasey. 


Sporobolus arkansanus Nutt. ex Vasey, Contr. U.S. Natl. 
Herb. 3(1): 61. 1892. 
NOTE: nom. inval., as syn. of Sporobolus argutus Kunth, 
= Sporobolus pyramidatus (Lam.) Hitchce. 


Sporobolus arundinaceus (Griseb.) Kuntze, Revis. Gen. PI. 
3(2): 369. 1898. Basionym: Diachyrium arundinaceum 
Griseb. 

NOTE: hom. illeg. 
E. NAME: Sporobolus arundinaceus (Trin.) Kunth. 
= Sporobolus rigens (Trin.) Desv. 


Sporobolus asper (P. Beauv.) Kunth, Revis. Gramin. 1: 68. 
1829. Basionym: Vilfa aspera P. Beauv. 
= Sporobolus compositus (Poir.) Merr. 


Sporobolus asper (P. Beauv.) Kunth var. asper. 
= Sporobolus compositus (Poir.) Merr. 


Sporobolus asper var. canovirens (Nash) Shinners, Rhodora 
56(662): 30. 1954. Basionym: Sporobolus canovirens 
Nash. 

= Sporobolus clandestinus (Biehler) Hitchce. 


Sporobolus asper var. clandestinus (Biehler) Shinners, 
Rhodora 56(662): 30. 1954. Basionym: Agrostis 
clandestina Biehler. 

= Sporobolus clandestinus (Biehler) Hitchc. 


Sporobolus asper var.drummondii (Trin.) Vasey, Contr. U.S. 


~ Natl Herb. 3(1): 60. 1892. Basionym: Vilfa 
drummondii Trin. 
= Sporobolus compositus var. drummondii (Trin.) 


Kartesz & Gandhi. 


Sporobolus asper var. hookeri (Vrin.) Vasey, Descr. Cat. 
Grass. U.S. 43. 1885. Basionym: Vilfa hookeri Trin. 
= Sporobolus compositus (Poir.) Merr. 


Sporobolus asper var. macer (Trin.) Shinners, Rhodora 
56(662): 29. 1954. Basionym: Vilfa macra Trin. 

= Sporobolus compositus var. macer (Trin.) Kartesz & 
Gandhi. 


Sporobolus asper UNRANKED pilosus (Vasey) Hitchc., 
Proc. Biol. Soc. Wash. 41: 161. 1928. Basionym: 
Sporobolus pilosus Vasey. 

= Sporobolus compositus var. 
Kartesz & Gandhi. 


drummondii (Trin.) 


Sporobolus asper var. pilosus (Vasey) Hitche., N. Amer. FI. 


17(7): 488. 1928. Basionym: Sporobolus pilosus 
Vasey. 
= Sporobolus compositus var. drummondii (Trin.) 


Kartesz & Gandhi. 


Sporobolus aspericaulis Scribn., Bot. Gaz. 21: 15. 1896. T- 
Specim.: | of 1. North America, Richardson s.n. [T: 
BM, US (fragm. ex BM)]. 
NOTE: nom. illeg. superfl., Muhlenbergia aspericaulis 
Nees ex Trin., nom. inval., cited. 
CORRECT NAME: Sporobolus richardsonis (Trin.) Merr. 
= Muhlenbergia richardsonis (Trin.) Rydb. 


Sporobolus asperifolius (Nees & Meyen ex Trin.) Nees, Nov. 
Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19(Suppl. 
1): 9. 1841. Basionym: Vilfa asperifolia Nees & 
Meyen ex Trin. 
NOTE: also p. 141. 1843. 


Catalogue of New World Grasses: II. 


203 


= Muhlenbergia asperifolia (Nees & Meyen ex Trin.) 
Parodi. 


Sporobolus asperifolius var. brevifolius Vasey, Contr. U.S. 
Natl. Herb. 1(2): 56. 1890. 
NOTE: nom. nud. 
= Muhlenbergia arenacea (Buckley) Hitchc. 


Sporobolus asperifolius var. brevifolius Vasey, Contr. U.S. 
Natl. Herb. 3(1): 64. 1892. 
NOTE: nom. inval., as syn. of Sporobolus auriculatus 
Vasey. 
= Muhlenbergia arenacea (Buckley) Hitche. 


Sporobolus asperifolius var. major Vasey, Contr. U.S. Natl. 
Herb. 3(1): 64. 1892. T-Protol.: [USA: Texas: Presidio 
Co.: Marfa, 1883], (Harvard 10]. T-Specim.: 1 of 1. 
USA: Texas, Presidio Co., Oct 1883, V. Havard 10 
[IT: US-556866]. 
= Muhlenbergia asperifolia (Nees & Meyen ex Trin.) 
Parodi. 


Sporobolus atrovirens (Kunth) Kunth, Revis. Gramin. 1: 68. 
1829. Basionym: Vilfa atrovirens Kunth. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W. & R. Moran 
(1981); McVaugh, R. (1983); Pohl, R. W., C. G. 
Reeder & G. Davidse (1994); Sousa S.. M. & E. F. 
Cabrera C. (1983). 

SYN.: Agrostis atrovirens (Kunth) Roem. & Schult. 


Sporobolus attenuatus Nash, Fl. S.E. U.S. 123. 1903. T- 
Protol.: USA: Mississipi: Oktibbeha Co.: Starkville, 
TH. Kearney 83 |HT: NY; IT: US]. T-Specim.: | of 1. 
USA: Mississippi: Oktibbeha Co., 30 Sep 1896, 7-H. 
Kearney 83 [IT: US-746652, US-556867]. 
= Sporobolus compositus var. drummondii 
Kartesz & Gandhi. 


(Trin.) 


Sporobolus auriculatus Vasey, Contr. U.S. Natl. Herb. 3(1): 
64. 1892. T-Protol.: USA: Texas: Pena, Nealley s.n. T- 
Specim.: | of 1. USA: Texas, Duval Co., 1890, G.C. 
Nealley 492 [IT: US-997369]. 

= Muhlenbergia arenacea (Buckley) Hitchc. 


Sporobolus bahamensis Hack., Oesterr. Bot. Z. 52(2): 56. 
1902. T-Protol.: West Indies: Bahamas:  Insula 
Bahamenses: Acklins Island, in fruticetis gregaria, 
Eggers 3905 [IT: US-3159457 (fragm. ex W)]. T- 
Specim.: | of 1. [HT: W; IT: US-3159457 (fragm. ex 
W)). 

DST.: Cari. 
LVL. ACCPTC.: I. 
ALSO ACCPTD. BY: Hitchcock, A. S. (1936). 


Sporobolus benthamii var. robustus Domin, Biblioth. Bot. 85: 
348. 1915. T-Protol.: Australia: Nord-Australian: Gulf 
of Carpentaria, Port Darwin, F. Schultz 212. T- 
Specim.: | of 1. Australia: Northern Territory: Port 
Darwin, Schultz 2/2 [LT: K], LT designated by 
Baaijens & Veldkamp, Blumea 35: 446 (1991). 
= Sporobolus virginicus (L.) Kunth. 


Sporobolus berteroanus (Trin.) Hitchc. & Chase, Contr. U.S. 
Natl. Herb. 18(7): 370. 1917. Basionym: Vilfa 
berteroana Trin. 

= Sporobolus indicus (L.) R. Br. 


Sporobolus bogotensis Swallen & Garcia-Barr., Caldasia 
2(8): 301, f. B. 1943. T-Protol.: Colombia: 
Cundinamarca: Macizo de Bogota, Sierra de Siete 
Picos, 2900-3100 m, 28 Jun 1939, J. Cuatrecasas 5680 
(HT: US; IT: COL]. T-Specim.: 1 of 1. Colombia: 
Cundinamarca, 28 Jun 1939, J. Cuatrecasas 5680 [HT: 
US-1795919; IT: COL]. 

DST.: Colo, Ecua. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Jorgensen, P. M. & S. Leén-Yanez 
(eds.) (1999); Swallen, J. R. & H. Garcia-Barriga 
(1943). 


Sporobolus brasiliensis (Raddi) Hack., Bull. Herb. Boissier, 
ser. 2, 4(3): 278. 1904. Basionym: Aira brasiliensis 
Raddi. 

= Eragrostis airoides Nees. 


Sporobolus buckleyi Vasey, Bull. Torrey Bot. Club 10: 128. 
1883. T-Protol.: USA: Texas, S.B. Buckley s.n. T- 
Specim.: 1 of 1. USA: Texas, 1883, D.B. Buckley s.n. 
[IT: US-556873]. 

DST.: Beli, Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Balick, M. J., 
M. H. Nee & D. E. Atha (2000); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Davidse, G.. M. Sousa S. & A. O. Chater (editores 
generales) (1994); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Hitchcock, A. S. 
(1951); Pohl, R. W., C. G. Reeder & G. Davidse 
(1994); Sousa S., M. & E. F. Cabrera C. (1983); 
Swallen, J. R. (1955). 


Sporobolus camporum Swallen, Sellowia 7: 10. 1956. T- 
Protol.: Brazil: Santa Catarina: Sao Joaquim, on 
campo, alt. 1200 m, 21-29 Jan 1950,P. R. Reitz 3431 
[HT: US-2042484]. T-Specim.: | of 1. Brazil: Santa 
Catarina, 21-29 Jan 1950, R. Reitz 343] |HT: US- 
2042484]. 

DST.: Braz, Urug. 
LVL. ACCPTC:.: 1. 
ALSO ACCPTD. BY: Renvoize, S. A. (1988). 


Sporobolus canovirens Nash, Man. Fl. N. States 1042. 1901. 
T-Protol.: USA: "In dry sandy soil, Tenn. to Mo., 
Kans., Miss. and Tex.". T-Specim.: 1 of 1. USA: 
Kansas: St. George, 3 Sep 1890, Kellerman s.n. (HT: 
NY; IT: US-611006(neg. & fragm. ex NY)]. 

= Sporobolus clandestinus (Biehler) Hitchc. 


Sporobolus capensis Kunth, Enum. Pl. 1: 212. 1833. 
NOTE: nom. illeg. superfl. 
= Sporobolus africanus (Poir.) Robyns & Tournay. 


Sporobolus capillaris Vasey, Contr. U.S. Natl. Herb. 1(8): 
283. 1893. T-Protol.: Mexico[LT]. T-Specim.: | of 2. 
Mexico: Rio Blanco, 1886, £. Palmer 5/2 [LT: LT: 
US-82013; ILT: NY-19973], LT designated by 
Hitchcock, Contr. U.S. Natl. Herb. 17(3): 311 (1913). 
2 of 2. Mexico: Jalisco: dry ledges, barranca near near 
Guadalajara, 15 Sep 1891, C.G. Pringle 3853 [ST: 
MO, US]. 


204 


Catalogue of New World Grasses: II. 


NOTE: hom. illeg. 

E. NAME: Sporobolus capillarisMiq. 
CORRECT NAME: Sporobolus trichodes Hitchc. 
= Sporobolus trichodes Hitchc. 


Sporobolus capillipes M.E. Jones, Contr. W. Bot. 14: 9. 
1912. T-Protol.: Mexico: Chihuahua: San Diego 
Canon, Sierra Madre Mts. T-Specim.: | of 1. Mexico: 
Chihuahua: San Diego Canyon (W of Colonia Juarez), 
Sierra Madre Mountains, 16 Sep 1903,Jones s.n. [HT: 
POM; IT: US-90605 (fragm.)]. 

= Muhlenbergia capillipes (M.E. Jones) P.M. Peterson & 
Annable. 


Sporobolus capillipes var. delicatulus M.E. Jones, Contr. W. 
Bot. 14: 10. 1912. T-Protol.: Mexico: Chihuahua: 
Soldier Cafion: Sierra Madre Mts.: Sep 1903, /ME. 
Jones] s.n. T-Specim.: | of 1. Mexico: Chihuahua: 
Soldier Canyon (Arroyo Soldado), Sierra Madre Mts., 
16 Sep 1903, Jones s.n. [HT: POM; IT: US-90606 
(fragm.)]. 

= Muhlenbergia capillipes (M.E. Jones) P.M. Peterson & 
Annable. 


Sporobolus ciliatus J. Presl, Relig. Haenk. 1(4-5): 242. 1830. 


T-Protol.: Panama: "Hab. in Panama.", Haenke s.n. 
= Sporobolus piliferus (Trin.) Kunth. 


Sporobolus ciliatus Munro ex Hook. f., Fl. Brit. India 7: 251. 


1896. T-Protol.: Munro in Herb. Ind. Or. Hf & T. 
NOTE: nom. inval., as syn. of Sporobolus piliferus Kunth. 
= Sporobolus piliferus (Trin.) Kunth. 


Sporobolus ciliatus var. subfasciculatus (D6ll) Hack., 
Ergebn. Bot. Exp. Sudbras. 1: 17. 1906. Basionym: 
Vilfa ciliata var. subfasciculata Doll. 
NOTE: also in Denkschr. Kaiserl. Akad. Wiss., Math.- 
Naturwiss. KI. 79: 73 (1908). 
SYN.: Vilfa ciliata var. subfasciculata DOI. 
= Status not yet determined. 


Sporobolus clandestinus (Biehler) Hitche., Contr. U.S. Natl. 


Herb. 12(3): 150. 1908. Basionym: Agrostis 
clandestina Biehler. 
DST.: USA. 


LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Anonymous (1986); Fernald, M. 
(1950); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Hitchcock, A. 
S. (1951); Radford, A. E., H. E. Ahles & C. R. Bell 
(1968); Small, J. K. (1933). 

Agrostis clandestina Biehler; Muhlenbergia 
clandestina (Biehler) Trin.; Sporobolus asper var. 
canovirens (Nash) Shinners; Sporobolus asper var. 


SYN.: 


clandestinus (Biehler)  Shinners; Sporobolus 
canovirens Nash; Sporobolus clandestinus — var. 
canovirens (Nash) Steyerm. & C.L. Kucera; 


Sporobolus compositus var. clandestinus (Biehler) 
Wipff & S.D. Jones; Vilfa clandestina (Biehler) Nees 
ex Steud. 


Sporobolus clandestinus var.canovirens (Nash) Steyerm. & 
C.L. Kucera, Rhodora 63(745): 25. 1961. Basionym: 
Sporobolus canovirens Nash. 

= Sporobolus clandestinus (Biehler) Hitchc. 


Sporobolus coahuilensis Valdés-Reyna, Phytologia 41(2): 
81. 1978. T-Protol.: Mexico: Coahuila, Hendrickson 
15363 [LL]. 


DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000). 


Sporobolus complanatus Scribn. ex Beal, Grass. N. Amer, 2: 
310. 1896. 
NOTE: nom. inval., as syn. of Epicampes bourgeaei E. 
Fourn. 
= Muhlenbergia peruviana (P. Beauv.) Steud. 


Sporobolus compositus (Poir.) Merr., Circ. Div. Agrostol. 
U.S.D.A. 35: 6. 1901. Basionym: Agrostis composita 
Poir. 

DST.: Cana, Mexi, USA. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Wunderlin, R. P. (1998). 

SYN.: Agrostis involuta Muhl., Glyceria stricta Buckley; 
Muhlenbergia composita Trin. ex Kunth; Sporobolus 
asper (P. Beauv.) Kunth; Sporobolus asper var. 
asper, Sporobolus asper var. hookeri (Trin.) Vasey; 
Sporobolus  longifolius (Torr.) Alph. Wood; 
Sporobolus pilosus Vasey; Vilfa aspera P. Beauv.; 
Vilfa composita (Poir.) P. Beauv. 


Sporobolus compositus var. clandestinus (Biehler) Wipff & 
S.D. Jones, Phytologia 78(4): 244. 1995. Basionym: 
Agrostis clandestina Biehler. 

= Sporobolus clandestinus (Biehler) Hitchc. 


Sporobolus compositus (Poir.) Merr. var. compositus. 
LVL. ACCPTC.:: I. 


Sporobolus compositus var. drummondii (Trin.) Kartesz & 
Gandhi, Phytologia 78(1): 11. 1995. Basionym: Vilfa 
drummondii Trin. 

LVL. ACCPTC.:: 1. 

SYN.: Sporobolus asper UNRANKED pilosus (Vasey) 
Hitche.; Sporobolus asper var. drummondii (Trin.) 
Vasey; Sporobolus asper var. pilosus (Vasey) 
Hitche.; Sporobolus attenuatus Nash; Sporobolus 
drummondii (Trin.) Vasey. 


Sporobolus compositus var. macer (Trin.) Kartesz & 
Gandhi, Phytologia 78(1): 11. 1995. Basionym: Vilfa 
macra Trin. 

LVL. ACCPTC.: 1. 

SYN.: Sporobolus asper var. macer (Trin.) Shinners; 
Sporobolus macer (Trin.) Hitche.; Sporobolus 
macrus Hitchc. 


Sporobolus compressus (Torr.) Kunth, Enum, Pl. 1: 217. 
1833. Basionym: Agrostis compressa Torr. 
NOTE: comb. illeg. based on Agrostis compressa Torr. 
hom. illeg., equals Muhlenbergia torreyana (Schult.) 
Hitche. 
= Muhlenbergia torreyana (Schult.) Hitche. 


Sporobolus confusus (E. Fourn.) Vasey, Bull. Torrey Bot. 
Club 15: 293. 1888. Basionym: Vilfa confusa E. Fourn. 
= Muhlenbergia minutissima (Steud.) Swallen. 


Sporobolus confusus var. aberrans M.E. Jones, Contr. W. 
Bot. 14: 10. 1912. T-Specim.: | of 3. USA: Arizona: 
Bowie Co., 21 Aug 1884, ME. Jones s.n. [ST: US- 
825374]. 2 of 3. USA: Arizona: Bowie, Jones s.n. 
{LT], LT designated by Peterson & Annable, Syst. 
Bot. Monogr. 31: 83. 1991. 3 of 3. Mexico: 
Chihuahua: Colonia Juarez [ST]. 


Catalogue of New World Grasses: II. 


205 


= Muhlenbergia sinuosa Swallen. 


Sporobolus contractus Hitchce., Amer. J. Bot. 2: 303. 1915. 

REPLACED NAME: Sporobolus cryptandrus — var. 

strictus Scribn. 

BLOCKING NAME: Sporobolus strictus Franch. 

DST.: Mexi, USA. 

LVL. ACCPTC.: |. 

ALSO ACCPTD. BY: Abrams, L. (1923); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gleason, H. A. 
& A. Cronquist (1968); Gleason, H. A. & A. 
Cronquist (1991); Gould, F. W. & R. Moran (1981); 
Hickman, J. C. (1993); Hitchcock, A. S. (1951); 
Munz, P. A. & D. D. Keck (1959); Munz, P. A. 
(1974); Shreve, F. & I. L. Wiggins (1964). 

Sporobolus cryptandrus var. strictus 
Sporobolus strictus (Scribn.) Merr. 


SYN:: Scribn.; 


Sporobolus copiapinus Phil., Anales Univ. Chile 94: 6. 1896. 
T-Protol.: Chile: Atacama: prope Copiapo lecta. T- 
Specim.: | of 1. Chile: prope Copiapo lecta [T: IT: 
US]. 

= Muhlenbergia asperifolia (Nees & Meyen ex Trin.) 
Parodi. 


Sporobolus creber De Nardi, Contr. New South Wales Natl. 
Herb. 4(7): 406, t. 1, 1A. 1973. T-Specim.: 1 of 1. 
Australia: New South Wales: Warrumbungle Range, c. 
30 mi E of Gulargambone, 20 May 1971, McDonald 
s.n. [HT: NSW-121997; IT: BRI, K, L, NSW, US- 
2884167, US-2892244]. 

DST.: USA. 
LVL. ACCPTC.: 1; introduced. 
ALSO ACCPTD. BY: Simon, B. K. & S. W. L. Jacobs 

(1999). 

Sporobolus 

Veldkamp. 


SYN:: indicus. var. creber (De Nardi) 


Sporobolus crucensis Renvoize, Gram. Bolivia 334. 1998. 
T-Protol.: Bolivia: Santa Cruz: Chavez, Concepcion 
hacia Lomerio, Seide/ 3097 [HT: LPB]. 

DST.: Boli. 
LVL. ACCPTC:.: 1. 
ALSO ACCPTD. BY: Renvoize, S. A. (1998). 


Sporobolus cryptandrus (Torr.) A. Gray, Manual 576. 
1848. Basionym: Agrostis cryptandra Torr. 

DST.: Arge, Cana, Mexi, USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Abrams, L. (1923); Beetle, A. A. 
(1977): Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Fernald, M. 
(1950); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Gould, F. W. 
& R. Moran (1981); Great Plains Flora Association 
(1986); Hickman, J. C. (1993); Hitchcock, A. S. 
(1951); Hitchcock, C. L., A. Cronquist, M. Ownbey 
& J. W. Thompson (1969); Moss, E. H. (1983); 
Munz, P. A. & D. D. Keck (1959); Munz, P. A. 
(1974); Nicora, E. G., M. E. De Paula, A. M. Faggi, 
M. del C. Mariano, A. M. Miante A., L. R. Parodi, C. 
A. Petetin, F. A. Roig & Z. Rugolo de Agrasar 
(1978); Scoggan, H. J. (1978 [1979]); Shreve, F. & I. 


L. Wiggins (1964); Small, J. K. (1933); Torres, M. A. 
(1970); Voss, E. G. (1972); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

Agrostis cryptandra Torr.,; Sporobolus cryptandrus 
subsp. fuscicolus (Hook.) E.K. Jones & Fassett; 
Sporobolus cryptandrus var. involutus Farw.; 
Sporobolus cryptandrus var. occidentalis E.K. Jones 
& Fassett; Sporobolus cryptandrus var. typicus 
(Torr.) E.K. Jones & Fassett; Sporobolus cryptandrus 
var. vaginatus Lunell; Sporobolus subinclusus fo. 
panicula expansa Stuck.; Sporobolus subinclusus 
Phil.; Sporobolus subinclusus var. expansus Parodi; 
Vilfa cryptandra (Torr.) Trin.; Vilfa tenacissima vat. 
fuscicola Hook.; Vilfa triniana Steud. 


SYN: 


Sporobolus cryptandrus var. flexuosus (Thurb. ex Vasey) 
Thurb., Bot. California 2: 269. 1880. Basionym: Vilfa 
cryptandra var. flexuosa Thurb. ex Vasey. 

= Sporobolus flexuosus (Thurb. ex Vasey) Rydb. 


Sporobolus cryptandrus subsp.fuscicolus (Hook.) E.K. Jones 
& Fassett, Rhodora 52: 126. 1950. Basionym: Vilfa 
tenacissima vat. fuscicola Hook. 

= Sporobolus cryptandrus (Torr.) A. Gray. 


Sporobolus cryptandrus var. giganteus (Nash) E.K. Jones, 
Contr. W. Bot. 14: 11. 1912. Basionym: Sporobolus 
giganteus Nash. 

= Sporobolus giganteus Nash. 


Sporobolus cryptandrus var. involutus Farw., Michigan 
Acad. Sci. Rep. 22: 179. 1921. T-Protol.: USA: 
Michigan: Rochester: 4 Sep 1919, [Farwell] 5393. 

= Sporobolus cryptandrus (Torr.) A. Gray. 


Sporobolus cryptandrus var, occidentalis E.K. Jones & 
Fassett, Rhodora 52: 125. 1950. T-Protol.: USA: 
Oregon: Baker Co.: Bars of Snake River, Ballard's 
Landing ca 3 mi NE of Homestead: 8 Jul 1899, Cusick 
2222 [HT: US-362493]. T-Specim.: 1 of 1. 8 Jul 1899, 
W.C. Cusick 2222 [HT: US-362493]. 

= Sporobolus cryptandrus (Torr.) A. Gray. 


Sporobolus cryptandrus var. robustus Vasey, Contr. U.S. 
Natl. Herb. 1(2): 56. 1890. T-Protol.: USA: Texas, 
G.C. Nealley [746]. T-Specim.: 1 of 1. USA: Texas: 
Presidio Co., 1887, G.C. Nealley 746 [IT: US- 
556883]. 

= Sporobolus giganteus Nash. 


Sporobolus cryptandrus var. strictus Scribn., Bull. Torrey 
Bot. Club 9: 103. 1882. T-Protol.: USA: Arizona: 
banks of the Rillita, near Camp Lowell, Pringle s.n. T- 
Specim.: | of 1. USA: Arizona, 15 Jun 1881, C.G. 
Pringle s.n. {(IT: US-825284]. 

CORRECT NAME: Sporobolus contractus Hitche. 
= Sporobolus contractus Hitche. 


Sporobolus cryptandrus var. typicus (Torr.) E.K. Jones & 
Fassett, Rhodora 52: 125. 1950. Basionym: Agrostis 
cryptandra Torr. 

NOTE: nom. inval., see ICBN Art. 24.3. 
= Sporobolus cryptandrus (Torr.) A. Gray. 


Sporobolus cryptandrus var. vaginatus Lunell, Amer. Midl. 
Naturalist 2: 123. 1911. T-Protol.: USA: North 
Dakota: Benson Co.: on bare, gravelly hillsides at 
Pleasant Lake: 28 Jul 1911, /. Lunell 93. 


206 


Catalogue of New World Grasses: II. 


= Sporobolus cryptandrus (Torr.) A. Gray. 


Sporobolus cubensis Hitchce., Contr. U.S. Natl. Herb. 12(6): 
237. 1909. T-Protol.: Cuba: Isle of Pines, A.H. Curtiss 
392 {HT: US-522010]. T-Specim.: 1 of 1. Cuba: La 
Habana, 6 Mar 1904, 4.H. Curtiss 392 (HT: US- 
522010]. 

DST.: Beli, Boli, Braz, Cari, Colo, CoRi, Ecua, FrGu, Guya, 
Hond, Mexi, Nica, Pana, Peru, Suri, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Balick, M. J., 
M. H. Nee & D. E. Atha (2000); Beetle, A. A. 
(1977); Catasus Guerra, L. (1997); Cowan, C. P. 
(1983); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J.  Valdes-Reyna (2000); 
Filgueiras, T. S. (1991); Filgueiras, T. S. (1999): 
Hitchcock, A. S. (1909); Hitchcock, A. S. (1936); 
Judziewicz, E. J. (1990); Killeen, T. J. (1990); 
Liogier, H. & L. Martorell (1982); Pohl, R. W. 
(1980); Pohl, R. W., C. G. Reeder & G. Davidse 
(1994); Renvoize, S. A. (1998); Swallen, J. R. 
(1955). 


Sporobolus curtissii Small ex Kearney, Bull. Div. Agrostol., 
U.S.D.A. 1: 24. 1895. T-Protol.: Florida: Jacksonville, 
in pine barrens, growing in open ground along 
railways, 4.1. Curtiss, cites "Sporobolus floridanus 
curtissii Vasey, in herb." and 2 paratypes. T-Specim.: 
| of 3. USA: Florida: Swamps, 1883, 4.1. Curtiss s.n. 
[HT: US-556876; IT: MSC], Sporobolus floridanus 
var. Curtissii in Vasey's hand. 2 of 3. USA: Florida: 
Moist pine barrens near Jacksonville, 13 Nov 1894, 
A.H, Curtiss 5181 [PT: US-221904, US-221903]. 3 of 
3. USA: Florida, 21 Aug 1893,4.H. Curtiss 4053 [PT: 
US-221755]. 

NOTE: diagonsis valid here. 

LVL. ACCPTC.: I. 

SYN.: Sporobolus curtissii Small ex Scribn.; Sporobolus 
loridanus var. curtissii (Small ex Kearney) Vasey ex 
Beal; Sporobolus floridanus var. curtissii Vasey; 
Sporobolus floridanus var. curtissii Vasey ex 
Kearney. 


Sporobolus curtissii Small ex Scribn., Bull. Div. Agrostol., 
U.S.D.A. 7: 142, f. 124. 1897. Basionym: <>. 
NOTE: isonym, said to be based on S. floridanus var. 
curtissli Vasey ex Beal. 
= Sporobolus curtissii Small ex Kearney. 


Sporobolus cuspidatus (Torr. ex Hook.) Alph. Wood, Amer. 
Bot. Fl. 2: 385. 1871. Basionym: Vilfa cuspidata Torr, 
ex Hook. 

NOTE: as S. cuspidata. 
= Muhlenbergia cuspidata (Torr. ex Hook.) Rydb. 


Sporobolus depauperatus (Torr. ex Hook.) Scribn., Bull. 
Torrey Bot. Club 9: 103. 1882. Basionym: Vilfa 
depauperata Torr. ex Hook. 

= Muhlenbergia richardsonis (Trin.) Rydb. 


Sporobolus depauperatus var. filiformis Beal, Grass. N. 
Amer. 2: 296. 1896. T-Specim.: 1 of 2. USA: Utah, 4 
Aug 1879, MLE. Jones 1179 [ST: US-87211]. 2 of 2. 
USA: Montana (probably Wyoming): Yellowstone 
National Park,, 23 Aug 1888, Williams s.n. [LT: MSC; 
ILT: US (fragm.)], LT designated by Hitchcock, N. 
Amer.Fl. 17: xxx (1935). 


= Muhlenbergia filiformis (Thurb. ex S. Watson) Rydb. 


Sporobolus deserticolus Phil., Verz. Antofagasta Pfl. 82. 
189]. T-Protol.: Chile: Tarapaca: ad Aguas calientes 
lectus, FL Philippi s.n. T-Specim.: | of 1. Chile: 
Tarapaca: ad Aguas calientes, 25 Jan 1885, R.A. 
Philippi s.n. (HT: SGO; IT: US-1763471 (fragm. ex 
SGO)]. 

= Muhlenbergia asperifolia (Nees & Meyen ex Trin.) 
Parodi. 


Sporobolus diander (Retz.) P. Beauv., Ess. Agrostogr. 26, 
147, 178. 1812. Basionym: Agrostis diandra Retz. 
NOTE: orth. var. inval. 
CORRECT NAME: Sporobolus diandrus (Retz.) P. Beauv. 
= Sporobolus diandrus (Retz.) P. Beauv. 


Sporobolus diandrus (Retz.) P. Beauv., Ess. Agrostogr. 26, 
147, 178. 1812. Basionym: Agrostis diandra Retz. 
NOTE: the more commonly used orth. is Sporobolus 
diander but Beauvois' use is correct - see Baaijens & 
Veldkamp, Blumea 35: 433 (1991). 

DST.: Beli, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Balick, M. J.. M. H. Nee & D. E. 
Atha (2000); Bor, N. L. (1960); Davidse, G., M. 
Sousa S. & A. O. Chater (editores generales) (1994); 
Pohl, R. W., C. G. Reeder & G. Davidse (1994); 
Simon, B. K. & S. W. L. Jacobs (1999), 

Agrostis diandra Retz.; Agrostis elongata var. 
flaccida Roth; Spermachiton involutum Llanos: 
Sporobolus diander (Retz.) P. Beauv.; Sporobolus 
indicus var. diandrus (Retz.) Jovet & Guédés: 
Sporobolus indicus var. flaccidus (Roth) Veldkamp; 
Vilfa diandra (Retz.) Trin.; Vilfa erosa Trin.; Vilfa 
retzii Steud. 


SYN.: 


Sporobolus diandrus var. major Biise, Pl. Jungh. 3: 343. 
1854. T-Protol.: Colombia: Habit. insulam Javae prope 
Tjibogo, Juhnhuhn s.n. [HT: LJ]. T-Specim.: 1 of 1. 
Java: Cibogo, June, Junghuhn s.n. (HT: L-903,342- 
96]. 

= Sporobolus fertilis (Steud.) Clayton. 


Sporobolus diffusissimus Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 90. 1862. T-Protol.: USA: Texas: 
Western Texas, /Wright 726]. T-Specim.: | of 1. USA; 
Texas, 1849, C. Wright 726 [IT: US-3198038 
(fragm.)]. 

= Sporobolus airoides (Torr.) Torr. 


Sporobolus distichivaginatus R.W. Pohl, Novon 2(4): 325, 
f. 2. 1992. T-Protol.: Guatemala: Petén: Dolores, en 
orillando Rio Machaquilla, lado sur este de caserio, 
fruto amarillo, en foresta alta, 18 Feb 1971, R. Tun 
Ortiz 1610 [HT: ISC; IT: F, US-27991 20]. 

DST.: Guat. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Davidse, G. & R. W. Pohl (1992); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Pohl, R. W., C. G. Reeder & G. 
Davidse (1994). 


Sporobolus distichophyllus Phil., Anales Univ. Chile 94: 7. 
1896. 
NOTE: nom. nov., comb. illeg., Agrostis distichophylla 
R.A. Phil. hom. illeg. 
REPLACED NAME: Agrostis distichophylla Phil. 


Catalogue of New World Grasses: II. 


207 


BLOCKING NAME: Agrostis distichophylla Roem. & 
Schult. 

= Muhlenbergia asperifolia (Nees & Meyen ex Trin.) 
Parodi. 


Sporobolus domingensis (Trin.) Kunth, Enum. Pl. 1: 214. 
1833. Basionym: Vilfa domingensis Trin. 
NOTE: also wrongly cited as Revis. Gramin. |: suppl. 17. 


1830, 
DST.: Cari, Mexi, USA, Vene. 
LVL. ACCPTC.:: 1. 


ALSO ACCPTD. BY: Adams, C. D. (1972); Anonymous 

(1986); Beetle, A. A. (1977); Correll, D. S. & M. C. 
Johnston (1970); Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Espejo Serna, A., 
A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Godfrey, R. K. & J. W. Wooten (1979); Hitchcock, 
A. S. (1936); Hitchcock, A. S. (1951); Liogier, H. & 
L. Martorell (1982); Long, R. W. & O. Lakela 
(1971); Pohl, R. W., C. G. Reeder & G. Davidse 
(1994); Small, J. K. (1933); Wiggins, I. L. & D. M. 
Porter (1971); Wunderlin, R. P. (1998). 

Agrostis domingensis (Trin.) Schult.; Sporobolus 
inordinatus Mez;, Sporobolus pyramidatus subsp. 
domingensis (Trin.) Cat. Guerra; Vilfa domingensis 
Trin. 


SYN.: 


Sporobolus drummondii (Trin.) Wasey, Descr. Cat. Grass. 
U.S. 44. 1885. Basionym: Vilfa drummondii Trin. 
= Sporobolus compositus var. drummondii (Trin.) 
Kartesz & Gandhi. 


Sporobolus effusus Desv., Mem. Soc. Agric. Angers |: 164. 
1831. T-Protol.: Crescit in Brasilio. 
NOTE: also in Opusc. Sci. Phys. Nat. 60 [reprint] 1831- 
1833. 
= Status not yet determined. 


Sporobolus ejuncidus Nash, Man. Fl. N. States 106. 1901. 
NOTE: nom. illeg. superfl. 
=§. junceus Kunt. 
= Sporobolus junceus (P. Beauy.) Kunth. 


Sporobolus eminens J. Presl, Relig. Haenk. 1(4-5): 242. 1830. 
T-Protol.: Hab. ad Acapulco, Haenke s.n. 
= Sporobolus wrightii Munro ex Scribn. 


Sporobolus erectus Hitche., Contr. U.S. Natl. Herb. 17(3): 
313. 1913. T-Protol.: Mexico: Veracruz: Jalapa, 
collected in a clay cut, 1400 m, 2 Sep 1910, Hitchcock 
6616 [HT: US-691231]. T-Specim.: 1 of 1. Mexico: 
Veracruz: Jalapa, 2 Sep 1910, A.S. Hitchcock 6616 
[=Amer. Gr. Natl. Herb. 1285] (HT: US-691231, IT: 
MO-1649620]. 

DST.: Hond, Mexi. 

LVL. ACCPTC:.: I. 

ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Espejo Serna, A., 
A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Pohl, R. W., C. G. Reeder & G. Davidse (1994). 


Sporobolus exilis (Trin.) Balansa, J. Bot. (Morot) 4: 164. 
1890. Basionym: Vilfa exilis Trin. 
NOTE: basionym implied by citing Trin. 
= Sporobolus indicus (L.) R. Br. 
Sporobolus exilis Renvoize, Kew Bull. 42(4): 923. 1987. T- 


Protol.: Brazil: Parana: Mun. Palmeira, Dombrowski 
6520 [HT: PKDC; IT: K]. 


NOTE: nom. illeg. hom. 

E. NAME: Sporobolus exilis (Trin.) Balansa. 
CORRECT NAME: Sporobolus linearifolius Nicora. 
= Sporobolus linearifolius Nicora. 


Sporobolus eximius (Nees ex Trin.) Ekman, Ark. Bot. 
13(10): 41. 1913. Basionym: Vilfa eximia Nees ex 
Trin. 

DST.: Arge, Braz. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Filgueiras, T. S. (1999); Foster, R. C. 
(1958); Hitchcock, A. S. (1927); Renvoize, S. A. 
(1988); Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

SYN.: Vilfa eximia Nees ex Trin. 


Sporobolus expansus Scribn., Zoe 4(4): 390. 1894. T-Protol.: 
Mexico: Lower California: Pescadero: 23 Sep 1893, 
T.S. Brandegee 16. T-Specim.: 1 of 1. Mexico: Baja 
California: Pescadero, 23 Sep 1893,7.S. Brandegee 16 
[T: IT: US-3351770 (fragm.), NY-19972]. 
= Sporobolus wrightii Munro ex Scribn. 


Sporobolus fastigiatus J. Presl, Relig. Haenk. 1(4-5): 241. 
1830. T-Protol.: Peruvia, Haenke s.n. T-Specim.: | of 
1. Peru, Haenke s.n. [HT: PR; IT: PRC, US-3048470 
(fragm. ex PR)]. 
= Muhlenbergia fastigiata (J. Presl) Henrard. 


Sporobolus fertilis (Steud.) Clayton, Kew Bull. 19: 291. 
1965. Basionym: Agrostis fertilis Steud. 
NOTE: known only from cultivation. 
DST.: USA. 
LVL. ACCPTC.: 1; cultivated, introduced. 
ALSO ACCPTD. BY: Simon, B. K. & S. W. L. Jacobs 
(1999). 
SYN.: Sporobolus diandrus var. major Buse; Sporobolus 
indicus var. fertilis (Steud.) Jovet & Guédes; 
Sporobolus indicus var. major (Biise) Baaijens. 


Sporobolus filiculmis Vasey, Descr. Cat. Grass. U.S. 44. 
1885. 
NOTE: nom. nud.; validly published by Beal 1896. 
= Muhlenbergia thurberi (Scribn.) Rydb. 


Sporobolus filiculmis L.H. Dewey, Contr. U.S. Natl. Herb. 
2(3): 519. 1894. 
REPLACED NAME: Sporobolus minor Vasey ex A. Gray. 
BLOCKING NAME: Sporobolus minor Trin. ex Kunth. 
= Sporobolus vaginiflorus (Torr. ex A. Gray) Alph. Wood. 


Sporobolus filiculmis Vasey ex Beal, Grass. N. Amer. 2: 288. 
1896. T-Protol.: USA: New Mexico: Whipple Exped. 
[Plaza Larga], /Bigelow 778]. 

NOTE: hom. illeg. 

E. NAME: Sporobolus filiculmis L.H. Dewey. 
CORRECT NAME: Sporobolus thurberi Scribn. 
= Muhlenbergia thurberi (Scribn.) Rydb. 


Sporobolus filiformis (Thurb. ex S. Watson) Rydb., Contr. 
U.S. Natl. Herb. 3(3): 189. 1895. Basionym: Vilfa 
depauperata var. filiformis Thurb. ex S. Watson. 

= Muhlenbergia filiformis (Thurb. ex S. Watson) Rydb. 


Sporobolus fimbriatus (Trin.) Nees, Fl. Afr. Austral. Ill. 
156. 1841. Basionym: Vilfa fimbriata Trin. 
DST.: USA. 
LVL. ACCPTC.: 1; introduced. 


208 


Catalogue of New World Grasses: II. 


ALSO ACCPTD. BY: Bor, N. L. (1960); Phillips, S. 
(1995). 


Sporobolus flexuosus (Thurb. ex Vasey) Rydb., Bull. Torrey 
Bot. Club 32(11): 601. 1905. Basionym: Vilfa 
cryptandra var. flexuosa Thurb. ex Vasey. 

DST.: Mexi, USA. 

LVL. ACCPTC:: |. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. & 
R. Moran (1981); Hickman, J. C. (1993); Hitchcock, 
A. S. (1951); Munz, P. A. & D. D. Keck (1959); 
Munz, P. A. (1974). 

Sporobolus cryptandrus var. flexuosus (Thurb. ex 
Vasey) Thurb.; Vilfa cryptandra var. flexuosa Thurb. 
ex Vasey. 


SYN:: 


Sporobolus floridanus Chapm., Fl. South. U.S. 550. 1860. 
T-Protol.: USA: Florida: Middle and West Florida, 
[Chapman] s.n. T-Specim.: | of 1. USA: Florida: 
Franklin Co.: Apalachiola, Chapman s.n. [LT: NY- 
127474; ILT: NY-127472, NY-127471, NY-127473, 
US-998263], LT designated by Weakley & Peterson, 
Sida 18(1): 256 (1998). 

NOTE: as Sporobolus "Floridanus". 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Godfrey, R. K. 
& J. W. Wooten (1979); Hitchcock, A. S. (1951); 
Radford, A. E., H. E. Ahles & C. R. Bell (1968); 
Small, J. K. (1933); Weakley, A. S. & P.M. Peterson 
(1998); Wunderlin, R. P. (1998). 


Sporobolus floridanus var.curtissii Vasey, Descr. Cat. Grass. 
U.S. 44. 1885. 
NOTE: nom. nud. 
= Sporobolus curtissii Small ex Kearney. 


Sporobolus floridanus var. curtissii Vasey ex Kearney, Bull, 
Div. Agrostol., U.S.D.A. 1: 24. 1895. 
NOTE: nom. inval., as syn. of S. curtissi. 
= Sporobolus curtissii Small ex Kearney. 


Sporobolus floridanus var. curtissii (Small ex Kearney) 
Vasey ex Beal, Grass. N. Amer. 2: 290. 1896. 
Basionym: Sporobolus curtissii Small ex Kearney. 

NOTE: as new, but same type as for S. curtissii Small ex 
Kearney. 
= Sporobolus curtissii Small ex Kearney. 


Sporobolus funckii (Steud.) F.M. Knuth, Repert. Spec. Nov. 
Regni Veg. 43 (Beih.): 123. 1926. Basionym: Vilfa 
Junckii Steud. 

SYN.: Vilfa funckii Steud. 
= Status not yet determined. 


Sporobolus giganteus Nash, Bull. Torrey Bot. Club 25(2): 
88-89. 1898. T-Protol.: USA: New Mexico: Dona Ana 
Co., E. O. Wooton 394 [HT: NY]. T-Specim.: 1 of 1. 
USA: New Mexico: Dona Ana Co., 26 Aug 1897, £.0. 
Wooton 394 [HT: NY-128297; IT: US-330618]. 
DST.: Mexi, USA. 
LVL. ACCPTC:: L. 
ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 


Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Great Plains 
Flora Association (1986); Hitchcock, A. S. (1951). 
Sporobolus cryptandrus var. giganteus (Nash) E.K. 
Jones; Sporobolus cryptandrus var. robustus Vasey. 


SYN.: 


Sporobolus gracilis (Trin.) Merr., Rhodora 4(39): 48. 1902. 
Basionym: Vilfa gracilis Trin. 
= Muhlenbergia cuspidata (Torr. ex Hook.) Rydb. 


Sporobolus gracillimus (Thurb. ex S. Watson) Vasey, Descr. 
Cat. Grass. U.S. 44. 1885. Basionym: Vilfa gracillima 
Thurb. ex S. Watson. 

= Muhlenbergia filiformis (Thurb. ex S. Watson) Rydb. 


Sporobolus heterolepis (A. Gray) A. Gray, Manual 576. 
1848. Basionym: Vilfa heterolepis A. Gray. 

DST.: Cana, USA. 

LVL. ACCPTC.: |; cultivated. 

ALSO ACCPTD. BY: Correll, D. S. & M. C. Johnston 
(1970); Darke, R. (1999); Fernald, M. (1950); 
Gleason, H. A. & A. Cronquist (1968); Gleason, H. 
A. & A. Cronquist (1991); Great Plains Flora 
Association (1986); Hitchcock, A. S. (1951); 
Radford, A. E., H. E. Ahles & C. R. Bell (1968); 
Scoggan, H. J. (1978 [1979]); Voss, E. G. (1972). 
Agrostis heterolepis (A. Gray) Alph. Wood; Vilfa 
heterolepis A. Gray. 


SYN.: 


Sporobolus hians Van Schaack, Ann. Missouri Bot. Gard, 
37(3): 397. 1950. T-Protol.: Brazil: Minas Gerais: 
Municipio Ituintaba: S. Terezinha, 20 Dec 1949, grass 
from resfriado, 4. Macedo 2045 [HT: MO]. T- 
Specim.: | of 1. Brazil: Minas Gerais: Ituiutaba Mun., 
20 Dec 1949, A. Macedo 2045 [HT: MO-2097004; IT: 
US-2010752]. 

DST.: Braz. 
LVL. ACCPTC:: I. 
ALSO ACCPTD. BY: Filgueiras, T. S. (1999). 


Sporobolus hintonii Hartley, Bull. Misc. Inform. Kew 
1941(1): 23. 1941. T-Protol.: Mexico: Temascaltepec 
District: Vigas, 1,080 m, 16 Nov 1932, George B. 
Hinton 2610 [HT: K]. T-Specim.: 1 of 1. Mexico: 
Mexico, Temascaltepec, Vigas, 16 Nov 1932, G. B. 
Hinton 2610 |HT: K; IT: NY-277646, US-1867701, 
US-2240742]. 

NOTE: as Sporobolus "Hintoni". 

DST.: Mexi. 

LVL. ACCPTC:.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. L6épez- 
Ferrari, and J. Valdes-Reyna (2000). 


Sporobolus humifusus (Kunth) Kunth, Revis. Gramin. |: 67. 
1829. Basionym: Vilfa humifusa Kunth, 
DST.: Vene. 
LVL. ACCPTC.: 1. 


Sporobolus indicus (L.) R. Br., Prodr. 170. 1810. Basionym: 
Agrostis indica L. 
DST.: Arge, Beli, Boli, Braz, Cari, Chil, Colo, CoRi, Ecua, 
ElSa, Guat, Hond, Mexi, Nica, Pana, Para, Peru. 
USA, Urug, Vene. 
LVL. ACCPTC.:: 1. 
ALSO ACCPTD. BY: Adams, C. D. (1972); Anonymous 
(1986); Baaijens, G. J. & J. F. Veldkamp (1991); 
Balick, M. J., M. H. Nee & D. E. Atha (2000); 


Catalogue of New World Grasses: II. 


209 


Beetle, A. A. (1977); Berendsohn, W. G. & A. E. 
Araniva de Gonzalez (1991); Brako, L. & J. L. 
Zarucchi (1993); Breedlove, D. E. (1986); Burkart, 
A. (1969); Catasus Guerra, L. (1997); Correll, D. S. 
& M. C. Johnston (1970); Cowan, C. P. (1983); 
Davidse, G., M. Sousa S. & A. O. Chater (editores 
generales) (1994); Filgueiras, T. S. (1991); 
Filgueiras, T. S. (1999); Foster, R. C. (1958); 
Gleason, H. A. & A. Cronquist (1991); Godfrey, R. 
kK. & J. W. Wooten (1979); Gould, F. W (1979); 
Hickman, J. C. (1993); Hitchcock, A. S. (1909); 
Hitchcock, A. S. (1927); Hitchcock, A. S. (1936); 
Hitchcock, A. S. (1951); Jorgensen, P. M. & C. Ulloa 
U. (1994); Jorgensen, P. M. & S. Leén-Yanez (eds.) 
(1999): Lawesson, J. E., H. Adsersen & P. Bentley 
(1987); Liogier, H. & L. Martorell (1982); Long, R. 
W. & O. Lakela (1971); Longhi-Wagner, H. M. 
(1990); Marticorena, C. & M. Quezada (1985); 
McVaugh, R. (1983); Nicora, E. G., M. E. De Paula, 
A. M. Faggi, M. del C. Mariano, A. M. Miante A., L. 
R. Parodi, C. A. Petetin, F. A. Roig & Z. Rugolo de 
Agrasar (1978); Pohl, R. W. (1980); Pohl, R. W., C. 
G. Reeder & G. Davidse (1994); Porter, D. M. 
(1983); Reeder, J. R. & C. G. Reeder (1971); 
Renvoize, S. A. (1984); Renvoize, S. A. (1988); 
Renvoize, S. A. (1998); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 
Saravia, E. F. (1996); Simon, B. K. & S. W. L. 
Jacobs (1999); Skottsberg, C. (1922); Smith, L. B., 
D. C. Wasshausen & R. M. Klein (1981); Swallen, J. 
R. (1955); Torres, M. A. (1970); Tovar, O. (1993); 
Wiggins, I. L. & D. M. Porter (1971); Wunderlin, R. 
P. (1998): Zuloaga, F. O., E. G. Nicora, Z. E. Ragolo 
de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 

Agrostis compressa Poir.; Agrostis elongata Lam.; 
Agrostis indica L.; Agrostis tenuissima Spreng.; 
Sporobolus — angustus Buckley; — Sporobolus 
berteroanus (Trin.) Hitchce. & Chase; Sporobolus 
exilis (Trin.) Balansa; Sporobolus indicus fo. 
africanoides Jovet & Guédés; Sporobolus indicus 
var. africanus (Poir.) Jovet & Guédes; Sporobolus 
indicus fo. indicus; Sporobolus indicus subsp. 
indicus, Sporobolus indicus var. exilis (Trin.)_ T. 
Koyama; Sporobolus indicus var. — indicus, 
Sporobolus indicus fo. microspiculus Jovet & 
Guédés; Sporobolus lamarckii Desy. ex Ham.,; 
Sporobolus littoralis var. elongatus (P. Beauv.) T. 
Durand & Schinz; Vilfa angusta Buckley; Vilfa 
berteroana Trin.; Vilfa elongata P. Beauv.; Vilfa 
exilis Trin.; Vilfa familiaris Steud. ex Lechler, Vilfa 
indica (L.) Trin. ex Steud.; Vilfa rupestris Trin.; Vilfa 
tenacissima (L. f.) Kunth; Vilfa tenacissima var. 
exilis (Trin.) E. Fourn. 


SYN: 


Sporobolus indicus fo. africanoides Jovet & Guédes, Taxon 
22: 163. 1973. T-Protol.: Barba de bode, Rio de 
Janiero, 4 Feb 1874, Glaziou 6940 [HT: P]. 

NOTE: as Sporobolus indicus var. indicus fo. africanoides 
in Bull. Cent. Etudes Rech. Sci. 7: 60 (1968), nom. 
inval. 

= Sporobolus indicus (L.) R. Br. 


Sporobolus indicus var. africanus (Poir.) Jovet & Guédes, 
Taxon 22: 163. 1973. Basionym: Agrostis africana 
Poir. 


NOTE: nom. illeg. Also in Bull. Cent. Etudes Rech. Sci. 7: 
(1968), nom. inval. 

CORRECT NAME: Sporobolus indicus var. capensis Engl. 

= Sporobolus indicus (L.) R. Br. 


Sporobolus indicus var. andinus Renvoize, Kew Bull. 
49(3): 543. 1994. T-Protol.: Bolivia:  Ingavi, 
Huacullani, Beck 328 [HT: LPB; IT: K]. 

DST.: Boli, Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Renvoize, S. A. (1994); Renvoize, S. 
A. (1998). 


Sporobolus indicus var. creber (De Nardi) Veldkamp, 
Blumea 35(2): 432. 1991. Basionym: Sporobolus 
creber De Nardi. 

= Sporobolus creber De Nardi. 


Sporobolus indicus var. diandrus (Retz.) Jovet & Guédés, 
Taxon 22: 163. 1973. Basionym: Agrostis diandra 
Retz. 
NOTE: as Sporobolus indicus var. "diander"; Also in Bull. 
Cent. Etudes Rech. Sci. 7: (1968), nom. inval. 
= Sporobolus diandrus (Retz.) P. Beauv. 


Sporobolus indicus var. exilis (Trin.) 1. Koyama, J. Jap. Bot. 
37: 235. 1962. Basionym: Vilfa exilis Trin. 
= Sporobolus indicus (L.) R. Br. 


Sporobolus indicus var. fertilis (Steud.) Jovet & Guédeés, 
Taxon 22: 163. 1973. Basionym: Agrostis fertilis 
Steud. 

NOTE: Also in Bull. Cent. Etudes Rech. Sci. 7: (1968), 
nom. inval. 
= Sporobolus fertilis (Steud.) Clayton. 


Sporobolus indicus var. flaccidus (Roth) Veldkamp, Blumea 
35(2): 433. 1991. Basionym: Agrostis elongata var. 
flaccida Roth. 

= Sporobolus diandrus (Retz.) P. Beauv. 


Sporobolus indicus (L.) R. Br. var. indicus. 

DST.: Arge, Beli, Boli, Braz, Cari, Chil, Colo, CoRi, Ecua, 
ElSa, Guat, Hond, Mexi, Nica, Pana, Para, Peru, 
USA, Urug, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Renvoize, S. A. (1998); Smith, L. B., 
D. C. Wasshausen & R. M. Klein (1981); Wunderlin, 
R. P. (1998). 


Sporobolus indicus var. major (Bisse) Baaijens, Blumea 
35(2): 437. 1991. Basionym: Sporobolus diandrus var. 
major Biise. 

= Sporobolus fertilis (Steud.) Clayton. 


Sporobolus indicus fo.microspiculus Jovet & Guédés, Taxon 
22: 163. 1973. T-Protol.: Antilles, 1868, Husnot Pl 
Antilles 56 (HT: P]. 

NOTE: as Sporobolus indicus var. africanus fo. 
microspiculus in Bull. Cent. Etudes Rech. Sci. 7: 
(1968), nom. inval. 

= Sporobolus indicus (L.) R. Br. 


Sporobolus indicus fo. pyramidalis (P. Beauv.) Peter, Repert. 
Spec. Nov. Regni Veg. Beih. 40: 280, 291. 1931. 
Basionym: Sporobolus pyramidalis P. Beauv. 

= Sporobolus jacquemontii Kunth. 


210 


Catalogue of New World Grasses: II. 


Sporobolus indicus var. pyramidalis (P. Beauv.) Veldkamp, 
Blumea 35(2): 439. 1991. Basionym: Sporobolus 
pyramidalis P. Beauv. 

= Sporobolus jacquemontii Kunth. 


Sporobolus inflatus Vasey & L.H. Dewey ex Beal, Grass. N. 
Amer, 2: 289. 1896. 

NOTE: nom. illeg. superfl., as inflata; error for Sporobolus 
ligulatus Vasey & Dewey-fide Hitchcock & Chase, 
Man. Grass. U. S. 900 (1950). 

CORRECT NAME: Sporobolus ligulatus Vasey & L.H. 
Dewey. 

= Muhlenbergia dubia E. Fourn. 


Sporobolus inordinatus Mez, Repert. Spec. Nov. Regni Veg. 
17(19-30): 294. 1921. T-Protol.: Cuba, Ramon de la 
Sagra s.n. 

= Sporobolus domingensis (Trin.) Kunth. 


Sporobolus interruptus Vasey, Bull. Torrey Bot. Club 
15(1): 8-9, t. B, f. 1-5. 1888. T-Specim.: 1 of 2. USA: 
Arizona: San Francisco Forest, 9 Aug 1883, HH. 
Rusby 885 [15] (ST: US-556885, US-746621], Rusby 
885 cited as "15" in the protologue, but probably 
identical number as sent to Vasey. 2 of 2. USA: 
Arizona, 1866, Coues & Palmer 66 [ST: US]. 

DST.: USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Hitchcock, A. S. (1951). 
SYN.: Sporobolus arizonicus Thurb. ex Vasey. 


Sporobolus jacquemontii Kunth, Revis. Gramin. 2: 427, t. 
127. 1831. T-Protol.: Crecsit in insula St. Domingo, 
1827, Jacquemont s.n. T-Specim.: | of 1. Santo 
Domingo, 1827, V. Jacquemont [HT: P; IT: LE-TRIN- 
1712.01]. 
DST.: Beli, Boli, Braz, Cari, Colo, CoRi, Ecua, FrGu, Guat, 
Guya, Hond, Mexi, Nica, Pana, Peru, Suri, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Allem, A. C. 
& J. F.M. Valls (1987); Balick, M. J.. M. H. Nee & 
D. E. Atha (2000); Breedlove, D. E. (1986); Davidse, 
G., M. Sousa $. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Filgueiras, T. S. (1999); 
Godfrey, R. K. & J. W. Wooten (1979); Gould, F. W 
(1979), Judziewicz, E. J. (1990); Killeen, T. J. 
(1990); Liogier, H. & L. Martorell (1982); Pohl, R. 
W. (1980); Pohl, R. W., C. G. Reeder & G. Davidse 
(1994); Renvoize, S. A. (1988); Simon, B. K. & S. 
W. L. Jacobs (1999); Sousa S., M. & E. F. Cabrera C. 
(1983); Tovar, O. (1986); Tovar, O. (1993). 
Sporobolus indicus fo. pyramidalis (P. Beauv.) Peter; 
Sporobolus indicus var. pyramidalis (P. Beauv.) 
Veldkamp; Sporobolus pyramidalis P. Beauv.; 
Sporobolus pyramidalis var. jacquemontii (Kunth) 
Jovet & Guédes; Vilfa jacquemontii (Kunth) Trin. 


SYN.: 


Sporobolus jonesii Vasey, Bot. Gaz. 6: 297. 1881. T-Protol.: 
USA: California: collected at Soda Srings, ME. Jones 
303. T-Specim.: | of 1. USA: California, 1881, ME. 
Jones 303 {IT: US-556886]. 
= Muhlenbergia jonesii (Vasey) Hitche. 


Sporobolus junceus (P. Beauv.) Kunth, Revis. Gramin. 1: 
68. 1829. Basionym: Heleochloa juncea P. Beauv. 
DST.: Mexi, USA. 
LVL. ACCPTC.:: 1. 


ALSO ACCPTD. BY: Baaijens, G. J. & J. F. Veldkamp 
(1991); Beetle, A. A. (1977); Correll, D. S. & M. C. 
Johnston (1970); Fernald, M. (1950); Gleason, H. A. 
& A. Cronquist (1968); Hitchcock, A. S. (1951); 
Long, R. W. & O. Lakela (1971); Radford, A. E., H. 
E. Ahles & C. R. Bell (1968); Veldkamp, J. F. 
(1990); Wunderlin, R. P. (1998). 

Agrostis juncea Michx.; Agrostis thyrsoides Bosc ex 
Trin.; Aira triglumis Steud.; Axonopus poiretii Roem. 
& Schult.; Bennetia juncea Rat. ex B.D. Jacks.; 
Colpodium junceum (P. Beauv.) Trin.; Crypsis 
juncea (P. Beauv.) Steud.; Heleochloa juncea P. 
Beauv.; Sporobolus ejuncidus Nash; Sporobolus 
gracilis (Trin.) Merr.; Sporobolus funceus (Michx.) 
Kunth; Sporobolus poiretii (Roem. & Schult.) 
Hitche.; Vilfa fulvescens Trin.; Vilfa gracilis Trin.; 
Vilfa juncea (P. Beauv.) Trin.; Vilfa schiedeana 
Trin.; Vilfa subsetacea Trin.; Vilfa vinzenti Steud. 


SYN.: 


Sporobolus junceus (Michx.) Kunth Basionym: Agrostis 
Juncea Michx. 
NOTE: correctable error for S. junceus (P.Beauv.) Kunth. 
CORRECT NAME: Sporobolus junceus (P. Beauv.) Kunth. 
= Sporobolus junceus (P. Beauv.) Kunth. 


Sporobolus lamarckii Desv. ex Ham., Prodr. P|. Ind. Occid. 
4. 1825. 
NOTE: nom. nov., nom. illeg. superfl.; as Sporobolus 
"Lamarckii". 
REPLACED NAME: Agrostis elongata Lam. 
BLOCKING NAME: Sporobolus elongatus R. Br. 
CORRECT NAME: Agrostis indica L. 
= Sporobolus indicus (L.) R. Br. 


Sporobolus lasiophyllus Pilg., Bot. Jahrb. Syst. 37: 504. 
1906. T-Specim.: | of 2. Peru: Junin: Tarma: in 
montibus ad orientem a Palca versus, in campis aridis, 
graminibus  numerosis sed parvis, _ fruticibus 
intermixtis, 2700-3000 m, 12 Feb 1903, Weberbauer 
2467 (ST: B; IST: BAA-2904 (fragm. ex B), US- 
82017]. 2 of 2. Peru: Junin: in montibus ad oppidum 
Tarma, in eadem formatione, 3600 m, Feb 1903, 
Weberbauer 238] [ST: B]. 

DST.: Colo, Ecua, Peru, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Hitchcock, A. S. (1927); Jorgensen, 
P.M. & C. Ulloa U. (1994); Macbride, J. F. (1936); 
Tovar, O. (1993). 


Sporobolus ligularis Hack. ex Sodiro, Anales Univ. Centr. 
Ecuador 1889: 7. 1889. 
NOTE: nom. nud. 
= Muhlenbergia ligularis (Hack.) Hitche. 


Sporobolus ligularis Hack., Oesterr. Bot. Z. 52(2): 57. 1902. 
T-Protol.: Ecuador: Pichincha: Pifo, "in locis sterilibus 
elevatis altoplanitiei", 4. Sodiro [23] ]. T-Specim.: 1 of 
1. Ecuador: Pichincha, 23 Jan 1899, Sodiro s.n. (HT: 
W; IT: US-32743 13 (fragm.)]. 

= Muhlenbergia ligularis (Hack.) Hitche. 


Sporobolus ligulatus Vasey & L.H. Dewey, Contr. U.S. Natl. 
Herb. 1(8): 268. 1893. T-Protol.: USA: Texas: Presidio 
Co.: Limpia Canon, Sep 1892,G.C. Nealley 127 [HT: 
US]. T-Specim.: | of 1. USA: Texas: Presidio Co.: 
Limpia Canon, Sep 1892,G.C. Nealley 127 [HT: US- 
82018; IT: US-556887]. 
= Muhlenbergia dubia E. Fourn. 


Catalogue of New World Grasses: II. 


211 


Sporobolus linearifolius Nicora, Hickenia 2(19): 90. 1993. 
REPLACED NAME: Sporobolus exilis Renvoize. 
BLOCKING NAME: Sporobolus exilis (Trin.) Balansa. 
DST.: Braz. 

LVL. ACCPTC:: I. 
ALSO ACCPTD. BY: ref. missing (1994). 
SYN.: Sporobolus exilis Renvoize. 


Sporobolus littoralis (Lam.) Kunth, Revis. Gramin. 1: 68. 


1829. Basionym: Agrostis littoralis Lam. 
= Sporobolus virginicus (L.) Kunth. 


Sporobolus littoralis var. elongatus (P. Beauv.) T. Durand & 
Schinz, Consp. Fl. Afr. 5: 821. 1894, Basionym: Vilfa 
elongata P. Beauv. 

= Sporobolus indicus (L.) R. Br. 


Sporobolus longifolius (Torr.) Alph. Wood, Class-book Bot. 


(ed. 1861) 1861: 775. 
longifolia Torr. 
= Sporobolus compositus (Poir.) Merr. 


1861. Basionym: Agrostis 


Sporobolus macer (Trin.) Hitche., Amer. J. Bot. 2: 203. 1915. 
Basionym: Vilfa macra Trin. 
NOTE: as "macrus". 
= Sporobolus compositus var. macer (Trin.) Kartesz & 
Gandhi. 


Sporobolus macrospermus Scribn. ex Beal, Grass. N. 
Amer. 2: 302. 1896. T-Protol.: Mexico: Jalisco, C.G. 
Pringle 2447. T-Specim.: | of 1. Mexico: Jalisco, 29 
Oct 1889, C.G. Pringle 2447 [HT: MSC; IT: CM, L, 
LL, MO, NY-19975, NY-19977, NY-19976, US- 
81955, US-746695, US-746696, US-825364]. 

DST.: Guat, Hond, Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983); Pohl, R. W., C. G. Reeder & G. 
Davidse (1994): Swallen, J. R. (1955). 


Sporobolus macrus Hitche. 
NOTE: orth. var. S. macer (Trin.) Hitche. 
CORRECT NAME: Sporobolus macer (Trin.) Hitche. 
= Sporobolus compositus var. macer (Trin.) Kartesz & 
Gandhi. 


Sporobolus mangaloricus Hochst. ex Migq., Anal. Bot. Ind. 2: 


36. 1851. T-Specim.: 1 of 1. India: Tamil Nadu, 
Canara, near Mangalore, 1847, Hochstetter 137 [HT: 
U: IT: BM, kK, L, TUB]. 

= Sporobolus tenuissimus (Mart. ex Schrank) Kuntze. 


Sporobolus matrella Nees, Fl. Afr. Austral. Ill. 152. 1841. 
NOTE: nom. illeg. superfl.; as comb. based on Vilfa 
matrella Nees, itself a nom. illeg. superfl. for Vilfa 
intermedia. 
CORRECT NAME: Vilfa intermedia Trin. 
= Sporobolus virginicus (L.) Kunth. 


Sporobolus maximus Hauman, Physis (Buenos Aires) 5: 52. 
1921. T-Protol.: Argentina: Salta: Cafayate: abondant 
en terrains un peu sales, sur les rives inondables du 
Rio San Carlos, pres de El Corralito: Nov 1919, 
Hauman s.n. 

DST.: Arge. 
LVL. ACCPTC:: 1. 


ALSO ACCPTD. BY: Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 


Sporobolus mendocinus E. Méndez, Bol. Soc. Argent. Bot. 
30(1-2): 13-15, f. 1. 1994. T-Protol.: Argentina: 
Mendoza: Dept. San Rafael: en niveles de terrazas o 
pseudoterrazas del arroyo Blanco a 1820 m, 8 Jan 
1985, formando comunidades densas en contacto con 
la comunidad de Cortaderia rudiuscula, en suelos 
salinos, ricos en materia organica, £. Méndez 4269 
[HT: MERL; IT: SI]. 

DST.: Arge. 
LVL. ACCPTC.: 1. 


Sporobolus metallicolus Longhi-Wagner & Boechat, Acta 
Bot. Brasil. 7(2): 150, f. 1. 1993. T-Protol.: Brazil: 
Minas Gerais: ca 10 km W de Barao de Cocais, Serra 
do Caraca, 22 Jan 1971, /rwin, H.S., R.M. Harley & E. 
Onishi 28813 [HT: MBM]. T-Specim.: 1 of 1. Brazil: 
Minas Gerais, 22 Jan 1971, Jrwin, H.S., R.M. Harley & 
E. Onishi 28813 (IT: US-2869348]. 

DST.: Braz. 
LVL. ACCPTC.: 1. 


Sporobolus microspermus (Lag.) Hitche., J. Wash. Acad. Sci. 
23(10): 453. 1933. Basionym: Milium microspermum 
Lag. 
= Muhlenbergia minutissima (Steud.) Swallen. 


Sporobolus minarum Boechat & Longhi-Wagner, Iheringia, 
Bot. 44: 34, f. 1-2. 1994. T-Protol.: Brasil: Minas 
Gerais: Lavras, 11 Mar 1981, HF. Leitdo Filho et al. 
1197 [HT: ESAL]. 

DST.: Braz. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (1994). 


Sporobolus minor Trin, ex Kunth, Enum. PI. 1: 212. 1833. 
REPLACED NAME: Vilfa rupestris Trin. 
BLOCKING NAME: Sporobolus rupestris Kunth. 
DST.: Arge, Boli, Braz, Chil, Para. 
LVL. ACCPTC:: I. 
ALSO ACCPTD. BY: Renvoize, S. A. (1988); Renvoize, S. 
A. (1998). 
SYN.: Vilfa rupestris Trin. 


Sporobolus minor Vasey ex A. Gray, Manual 646. 1890. T- 
Specim.: 1 of 1. USA: North Carolina, 1888, £.1. 
Boyton s.n. (ST: US-997198]. 

NOTE: hom. illeg. 

E. NAME: Sporobolus minor Trin. ex Kunth. 

CORRECT NAME: Sporobolus filiculmis L.H. Dewey. 

= Sporobolus vaginiflorus (Torr. ex A. Gray) Alph. Wood. 


Sporobolus minutiflorus (Trin.) Link, Hort. Berol. 1: 88. 
1827. Basionym: Vilfa minutiflora Trin. 
= Sporobolus tenuissimus (Mart. ex Schrank) Kuntze. 


Sporobolus minutissimus (Steud.) Hitche., Proc. Biol. Soc. 
Wash. 41: 161. 1928. Basionym: Agrostis minutissima 
Steud. 

= Muhlenbergia minutissima (Steud.) Swallen. 


Sporobolus mirabilis Pilg., Bot. Jahrb. Syst. 56(Beibl. 123): 
27. 1920. T-Protol.: Peru: Cajamarca: Prov. Jaen: 
zwischen dem Tale des Shumba, eines rechten 
Nebenflusses des Chinchipe und der Ortschaft Jaen, 


212 


Catalogue of New World Grasses: II. 


grassteppe mit Staucheren, 700-800 m, sehr Haufig, 
stellenweise reine Bestande bildend, Apr 1912, 
Weberbauer 6186. T-Specim.: 1 of 1. Peru: 
Cajamarca: zwischen Shumba und Jaén, 25 Apr 1912, 
A. Weberbauer 6186 [HT: B; IT: BAA-2910 (fragm. 
ex B), US-81956]. 

DST.: Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Hitchcock, A. S. (1927); Macbride, 

J. F. (1936); Tovar, O. (1993). 


Sporobolus monandrus Roseng., B.R. Arill. & Izag., Bol. 
Fac. Agron. Univ. Montevideo 103: 12, f. 3. 1968. T- 
Protol.: Uruguay: Dep. Artigas: camino a Tres Cruces, 
aprox. Km 6-8, en quebrada pedregosa, 10 Jun 1952, 
Rosengurtt B-6200 (HT: MVFA]. T-Specim.: | of I. 
Uruguay: Dep. Artigas: camino a Tres Cruces, aprox. 
Km 6-8, en quebrada pedregosa, 10 Jun 1952, B. 
Rosengurtt B-6200 [HT: MVEA; IT: US-2946506]. 

DST.: Arge, Boli, Braz, Para, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Burkart, A. (1969); Killeen, T. J. 
(1990); Renvoize, S. A. (1998); Rosengurtt, B., B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 
Smith, L. B., D. C. Wasshausen & R. M. Klein 
(1981); Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 


Sporobolus muelleri(E. Fourn.) Hitche., N. Amer. Fl. 17(7): 
490. 1937. Basionym: Vilfa muelleri E. Fourn. 
NOTE: as Sporobolus "Miilleri" 
= Sporobolus purpurascens (Sw.) Ham. 


Sporobolus multinodis Hack., Repert. Spec. Nov. Regni 

Veg. 7: 316. 1909. T-Protol.: Brasilien: Rio Grande do 
Sul: Municipio Rio Pardo: Fasenda Soledad, 70 
m.s.m., Dec 1905, C. Juergens s.n. [HT: W?]. T- 
Specim.: | of |. Brazil: Rio Grande do Sul: Rio Pardo 
Mun., Dec 1905, C. Jurgens G/53 [IT: US-1127181, 
US-81957]. 

DST.: Braz, Urug. 

LVL. ACCPTC.: 1, 

ALSO ACCPTD. BY: Rosengurtt, B., B. R. Arrillaga de 

Maffei & P. Izaguirre de Artucio (1970). 
SYN.: Sporobolus multinodis var. exasperatus Hack. 


Sporobolus multinodis var. exasperatus Hack., Repert. Spec. 
Nov. Regni Veg. 7: 317. 1909. T-Protol.: Brazil: Rio 
Grande do Sul, Campo de St. Angelo, Municipio de St. 
Angelo dos Missioes, alt. 400 m.s.m., Feb 1906, C. 
Juergens G259. T-Specim.: | of 1. Brazil: Rio Grande 
do Sul, Campo de St. Angelo Mun., Feb 1906, C. 
Jurgens G259 (IT: US-1127180, US-81958]. 

= Sporobolus multinodis Hack. 


Sporobolus multiramosus Longhi-Wagner & Boechat, Acta 
Bot. Brasil. 7(2): 155, f. 1. 1993. T-Protol.: Brazil: 
Para Maraba, 19 Apr 1985, Rosa & Silva 4680 [HT: 
SP]. 
DST.: Braz. 
LVL. ACCPTC.: 1. 


Sporobolus muralis (Raddi) Hitche. & Chase, Contr. U.S. 
Natl. Herb. 18(7): 368. 1917. Basionym: Agrosticula 
muralis Raddi. 

= Sporobolus tenuissimus (Mart. ex Schrank) Kuntze. 


Sporobolus nealleyi Vasey, Bull. Torrey Bot. Club 15: 49. 
1888. T-Specim.: | of 1. USA: Texas: Brazos Co., 
1887, G.C. Nealley 752 [OM: US-556888]. 
NOTE: nom. nud. 
= Sporobolus nealleyi Vasey. 


Sporobolus nealleyi Vasey, Contr. U.S. Natl. Herb. 1(2): 57. 
1890. T-Protol.: USA: Texas: Brazos Santiago, G.C. 
Nealley 752 [HT: NY]. T-Specim.: | of 1. USA: 
Texas: Brazos Co., 1887, G.C. Nealleyi 752 [IT: US- 
556888 (fragm.)]. 

DST.: Mexi, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000). 

SYN.: Sporobolus nealleyi Vasey. 


Sporobolus neglectus Nash, Bull. Torrey Bot. Club 22(11): 
464. 1895. T-Protol.: Britton s.n. (HT: NY]. T- 
Specim.: | of 1. USA: New Jersey, Sussex O.; 
Woodruff Gap, 17 Sep 1887, N.L. Britton {IT: US- 
91125(fragm.)]. 

DST.: Cana, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Abrams, L. (1923); Fernald, M. 
(1950); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Great Plains 
Flora Association (1986); Hitchcock, A. S. (1951); 
Hitchcock, C. L., A. Cronquist, M. Ownbey & J. W. 
Thompson (1969); Moss, E. H. (1983); Scoggan, H. 
J. (1978 [1979]); Small, J. K. (1933); Voss, E. G. 
(1972). 

Sporobolus vaginiflorus var. neglectus (Nash) 
Scribn.; Sporobolus vaginiflorus var. neglectus 
(Nash) Shinners. 


SYN:: 


Sporobolus neglectus var. ozarkanus (Fernald) Steyerm. & 
C.L. Kucera, Rhodora 63(745): 25. 1961. Basionym: 
Sporobolus ozarkanus Fernald. 

= Sporobolus vaginiflorus var. ozarkanus (Fernald) 
Shinners. 


Sporobolus nudiramus Boechat &  Longhi-Wagner, 
Iheringia, Bot. 44: 36, f. 3-6. 1994. T-Protol.: Brasil: 
Parana: Ponta Grossa, 15 Mar 1976, G. Davidse & 
D'Arcy 11354 [HT: SP; IT: MO], This is an erroneous 
citation for Davidse, Ramamoorthy & Vital 11354. 
D'Arcy did not collect with me in Brazil—G. Davidse, 
2 Mar 1996. T-Specim.: | of |. Brazil: Parana: ca. 25 
km W of Ponta Grossa, just west of the intersection of 
Highway BR-373 and Rio Tibagi, elev. 740° m, 
Trachypogon dominated grassland on deep, sany loam, 
15S Mar 1976, G. Davidse, T.P. Ramamoorthy & D.M. 
Vital 11354 (HT: SP; IT: MO-3113919], Erroneously 
cited as Davidse & D'Arcy 11354 in the protologue. 

DST.: Braz. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (1994). 


Sporobolus ovatus Beal, Grass. N. Amer. 2: 300. 1896. 
NOTE: nom. nov., nom. illeg. superfl. = S. filiculmis L.H. 
Dewey. 
REPLACED NAME: Sporobolus minor Vasey ex A. Gray. 
BLOCKING NAME: Sporobolus minor Trin. ex Kunth. 
= Sporobolus vaginiflorus (Torr. ex A. Gray) Alph. Wood. 


Catalogue of New World Grasses: II. 


213 


Sporobolus ozarkanus Fernald, Rhodora 35(411): 109-110. 


1933. T-Protol.: USA: Missouri: barrens, Web CitY, 8 
Sep 1910, E. J. Palmer 3133 [HT: GH; IT: US]. T- 
Specim.: | of 1. USA: Missouri, 8 Sep 1910, E./ 
Palmer 3133 (HT: GH; IT: US-671791, US-671790]. 

= Sporobolus vaginiflorus var. ozarkanus (Fernald) 
Shinners. 


Sporobolus palmeri Scribn., Bull. Div. Agrostol., U.S.D.A. 
11: 48, t. 5. 1898. T-Protol.: Mexico: in alkali bottoms, 
growing in rather dense bunches, near the city of 
Durango, June 1896, E. Palmer 180. T-Specim.: | of 
1. Mexico: Durango, Jun 1896, E. Palmer 180 [IT: 
MO, NY-19974, US-998310]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000). 


Sporobolus patens Swallen, J. Wash, Acad. Sci. 31(8): 352, f. 
5. 1941. T-Protol.: USA: Arizona: Wilcox: 26 Sep 
1938, W. A. Silveus 3504 [HT: US-1723881; IT: LL]. 
T-Specim.: 1 of 1. USA: Arizona, 26 Sep 1938, W.A. 
Silveus 3504 [HT: US-1723881; IT: US-3278411, US- 
F3215630]. 
= Sporobolus pyramidatus (Lam.) Hitchce. 


Sporobolus paucifolius Boechat & Longhi-Wagner, 
Iheringia, Bot. 44: 37, f. 7-9. 1994. T-Protol.: Brasil: 
Goids: Alto Paraiso-Teresina, 10 Oct 1979, E.P. 
Heringer et al. 2382 (HT: IBGE]. 

DST.: Braz. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (1994). 


Sporobolus phleoides Hack., Museo Nac. Buenos Aires 13: 
468. 1906. T-Protol.: Argentina: Cordoba: Rio Seco, 
Stuckert 2323 [HT: Herb. Arg.]. T-Specim.: 1 of 1. 
Argentina: Cordoba, 1897, 7..V. Stuckert 2323 [IT: 
US-72698 (fragm.)]. 

DST.: Arge. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 


Sporobolus piliferus (Trin.) Kunth, Enum. PI. 1: 211. 1833. 
Basionym: Vilfa pilifera Trin. 

NOTE: Veldkamp considers Sporobolus ciliatus J. Presl to 

be distinct. 

DST.: Braz, CoRi, FrGu, Guya, Nica, Pana, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Baaijens, G. J. & J. F. Veldkamp 
(1991); Bor, N. L. (1960); Davidse, G., M. Sousa S. 
& A. O. Chater (editores generales) (1994); Phillips, 
S. (1995); Pohl, R. W., C. G. Reeder & G. Davidse 
(1994); Renvoize, S. A. (1988). 

Sporobolus ciliatus J. Presl; Sporobolus ciliatus 
Munro ex Hook. f.; Sporobolus piliferus (Trin.) 
Kuhlm.; Sporobolus praecox A. Chev.; Sporobolus 
rupestris Kunth; Sporobolus stachyanthus A. Rich.; 
Sporobolus stachydanthus A. Rich.; Sporobolus 
villosus (Rchb. ex Spreng.) Kunth; 7riachyrum 
nilagiricum Hochst. ex  Steud.; Triachyrum 
stachydanthum (A. Rich.) Asch.; Vilfa pilifera Trin.; 
Vilfa rupestris (Kunth) Trin.; Vilfa stachydantha (A. 
Rich.) Steud. 


SYN.: 


Sporobolus piliferus (Trin.) Kuhlm., Comm. Lin. Telegr., 
Bot. 67(Bot. 11): 92. 1922. Basionym: Vilfa pilifera 
Trin. 
NOTE: isonym. 
E. NAME: Sporobolus piliferus (Trin.) Kunth. 
= Sporobolus piliferus (Trin.) Kunth. 


Sporobolus pilosus Vasey, Bot. Gaz. 16: 26. 1891. T-Protol.: 
USA: Kansas, B.B. Smyth s.n. T-Specim.: | of 1. USA: 
Kansas: Coolidge, 1890, B.B. Smyth 217 [HT: US- 
556890; IT: US-746585, US-75038]. 

= Sporobolus compositus (Poir.) Merr. 


Sporobolus pinetorum Weakley & P.M. Peterson, Sida 
18(1): 258, f. la-e, 2. 1998. T-Protol.: USA: North 
Carolina: Cumberland Co.: Fort Bragg Army Base, 
Macridge Danger Zone, ecotonal seepages and moist 
swales in upland Pinus palustris savannas, 31 Aug 
1993, Weakley s.n. [HT: US, IT: NCU]. T-Specim.: | 
of 1. USA: North Carolina: Cumberland Co.: Fort 
Bragg Army Base, Macridge Zone, 31 Aug 1993,5.N. 
Weakley [HT: US-3295350]. 

DST.: USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Weakley, A. S. & P. M. Peterson 
(1998). 


Sporobolus platensis Parodi, Revista Fac. Agron. Veterin. 
(Buenos Aires) 6(2): 133, f. 7. 1928. T-Protol.: 
Argentina: Buenos Aires: San Vicente, 14 Apr 1927, 
L.R. Parodi 8120 [HT: BAA]. T-Specim.: 1 of 1. 
Argentina: Buenos Aires, 14 Apr 1927, L.R. Parodi 
8120 |[HT: BAA; IT: US-1388644]. 

DST.: Arge, Braz, Urug. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Burkart, A. (1969); Rosengurtt, B., 
B. R. Arrillaga de Maffei & P. Izaguirre de Artucio 
(1970); Torres, M. A. (1970); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 


Sporobolus plumbeus (Trin.) Hemsl., Biol. Cent.-Amer., Bot. 
3(19): 546-547. 1885. Basionym: Vilfa plumbea Trin. 
= Muhlenbergia plumbea (Trin.) Hitche. 


Sporobolus poaevides Hack., Repert. Spec. Nov. Regni Veg. 
10(243-247): 167. 1911. T-Protol.: Mexico: in civitate 
Puebla: ad Rancho Posada, alt. 2194 m, Aug 1910,Fr. 
Nicolas 5423. T-Specim.: | of 1. Mexico: Puebla, Aug 
1910, F.G. Nicolas 5423 [HT: IT: US-87219]. 

= Muhlenbergia plumbea (Trin.) Hitchce. 


Sporobolus poiretii (Roem. & Schult.) Hitche., Bartonia 14: 
32. 1932. Basionym: Axonopus poiretii Roem. & 
Schult. 

= Sporobolus junceus (P. Beauv.) Kunth. 


Sporobolus potosiensis Wipff & S.D. Jones, Sida 16(1): 
165, 167, f. 1-4. 1994. T-Protol.: Mexico: San Luis 
Potosi, 9 km al E de Rio Verde, sobre la carretera a 
Rayon, alt. 1000 m, terrenos aluviales planos con 
vegetacion de zacatal haldfilo con Prosopis, graminea 
dominate, 16 Sep 1967, J. Rzedowski 24790 [HT: 
MICH. IT: LL, MICH, TAES, WIS]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000). 


214 


Catalogue of New World Grasses: II. 


Sporobolus pseudairoides Parodi, Revista Fac. Agron. 
Veterin. (Buenos Aires) 6(2): 144, f. 11. 1928. T- 
Protol.: Argentina: Misiones: Santa Ana, Feb 1922, 
L.R. Parodi 4316. T-Specim.. | of 1. Argentina: 
Misiones, Feb 1922, 1.R. Parodi 4316 [HT: BAA; IT: 
US-1448002]. 

DST.: Arge, Braz, Para. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Filgueiras, T. S. (1991); Filgueiras, 
T. S. (1999); Longhi-Wagner, H. M. (1990); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 


Sporobolus pulvinatus Swallen, J. Wash. Acad. Sci. 31(8): 
351, f. 4. 1941. T-Protol.: USA: Arizona: Apache co.: 
Adamana, 6-15 Aug 1903, D. Griffiths 5107 [HT: US- 
997877]. T-Specim.: | of 1. USA: Arizona: Apache 
Co., 6-15 Aug 1903, D. Griffiths 3107 [HT: US- 
997877]. 

= Sporobolus pyramidatus (Lam.) Hitche. 


Sporobolus purpurascens (Sw.) Ham., Prodr. Pl. Ind. 

Occid. 5. 1825. Basionym: Agrostis purpurascens Sw. 

DST.: Beli, Braz, Cari, Colo, CoRi, Ecua, ElSa, Guat, 
Hond, Mexi, Nica, Pana, Peru, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Beetle, A. A. 
(1977), Brako, L. & J. L. Zarucchi (1993); Catasus 
Guerra, L. (1997); Correll, D. S. & M. C. Johnston 
(1970), Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Gould, F. 
W (1979): Hitchcock, A. S. (1909); Hitchcock, A. S. 
(1927); Hitchcock, A. S. (1936); Hitchcock, A. S. 
(1951); Jorgensen, P. M. & C. Ulloa U. (1994); 
Jorgensen, P. M. & S. Leén-Yanez (eds.) (1999); 
Levin, G. A. & R. Moran (1989); Macbride, J. F. 
(1936); Pohl, R. W. (1980); Pohl, R. W., C. G. 
Reeder & G. Davidse (1994): Swallen, J. R. (1955); 
Tovar, O. (1993). 

SYN.: Agrostis purpurascens Sw.; Sporobolus muelleri (E. 
Fourn.) Hitche.; Sporobolus purpurascens (Sw.) 
Kuhlm.; Vilfa’ grisebachiana E. Fourn.; Vilfa 
liebmanni E. Fourn.; Vilfa muelleri E. Fourn.; Vilfa 
purpurascens (Sw.) P. Beauv. 


Sporobolus purpurascens (Sw.) Kuhlm., Comm. Lin. Telegr., 


Bot. 67(Bot. 11): 92. 
purpurascens Sw. 
NOTE: isonym. 
E. NAME: Sporobolus purpurascens (Sw.) Ham. 
= Sporobolus purpurascens (Sw.) Ham. 


1922. Basionym: Agrostis 


Sporobolus pyramidalis P. Beauv., Fl. Oware 2: 36, t. 80. 
1816. T-Specim.: | of 1. Nigeria, Palisot de Beauvois 
sn. [HT: G (photo, BRI, kK); IT: P?]. 
= Sporobolus jacquemontii Kunth. 


Sporobolus pyramidalis var. jacquemontii (Kunth) Jovet & 
Guédés, Taxon 22: 163. 1973. Basionym: Sporobolus 
Jacquemontii Kunth. 

NOTE: Also in Bull. Cent. Etudes Rech. Sci. 7: (1968), 
nom. inval. 
= Sporobolus jacquemontii Kunth. 


Sporobolus pyramidatus (Lam.) Hitche., Man. Grasses W. 
Ind. 84. 1936. Basionym: Agrostis pyramidata Lam. 


DST.: Arge, Boli, Braz, Cari, Chil, Colo, CoRi, Ecua, ElSa, 
FrGu, Guat, Hond, Mexi, Nica, Pana, Para, Peru, 
Suri, USA, Urug, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Allem, A. C. 
& J. F. M. Valls (1987); Beetle, A. A. (1977); 
Berendsohn, W. G. & A. E. Araniva de Gonzalez 
(1991); Brako, L. & J. L. Zarucchi (1993); 
Breedlove, D. E. (1986); Burkart, A. (1969); Catasus 
Guerra, L. (1997); Correll, D. S. & M. C. Johnston 
(1970); Davidse, G., M. Sousa S. & A. QO. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000): Fernald, 
M. (1950); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Gould, F. W 
(1979); Gould, F. W. & R. Moran (1981); Great 
Plains Flora Association (1986); Hitchcock, A. S. 
(1936); Hitchcock, A. S. (1951); Jorgensen, P. M. & 
C. Ulloa U. (1994); Jorgensen, P. M. & S. Leén- 
Yanez (eds.) (1999); Judziewicz, E. J. (1990); 
Killeen, T. J. (1990); Lawesson, J. E., H. Adsersen & 
P. Bentley (1987); Lindeman, J. C. & A. L. Stoffers 
(1963); Liogier, H. & L. Martorell (1982); Long, R. 
W. & O. Lakela (1971); McVaugh, R. (1983); Pohl, 
R. W. (1980); Pohl, R. W., C. G. Reeder & G., 
Davidse (1994); Porter, D. M. (1983); Reeder, J. R. 
& C. G. Reeder (1971); Renvoize, S. A. (1984); 
Renvoize, S. A. (1998); Rosengurtt, B.. B. R. 
Arrillaga de Maffei & P. Izaguirre de Artucio (1970); 
Shreve, F. & I. L. Wiggins (1964); Smith, L. B., D. 
C. Wasshausen & R. M. Klein (1981); Sousa S., M. 
& E. F. Cabrera C. (1983); Swallen, J. R. (1955); 
Torres, M. A. (1970); Tovar, O. (1993); Wiggins, I. 
L. & D. M. Porter (1971); Wunderlin, R. P. (1998); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

SYN.: Agrostis plumosa Ten.; Agrostis pyramidalis Rich. 
ex Steud.; Agrostis pyramidata Lam.; Sporobolus 
affinis Kunth; Sporobolus argutus (Nees) Kunth; 
Sporobolus argutus fo. purpurascens Hack.; 
Sporobolus argutus var. tuberculatus (Hack.) Hack.; 
Sporobolus arkansanus Nutt. ex Vasey; Sporobolus 
patens Swallen; Sporobolus pulvinatus Swallen; 
Sporobolus sabeanus Buckley ex Vasey;Sporobolus 
tuberculatus Hack.; Vilfa agrostoidea Buckley; Vilfa 
arguta Nees; Vilfa arkansana Trin.; Vilfa richardii 
Steud.; Vilfa sabeana Buckley. 


Sporobolus pyramidatus subsp. domingensis (Trin.) Cat. 
Guerra, Fontqueria 44: 143. 1996. Basionym: Vilfa 
domingensis Trin. 

= Sporobolus domingensis (Trin.) Kunth. 


Sporobolus racemosus Vasey, Bull. Torrey Bot. Club 14: 9. 
1887. T-Protol.: Mexico: Chihuahua: near Noragachie, 
E. Palmer s.n. Y-Specim.: | of 1. Mexico: Chihuahua: 
near Noragachie, 1885, E. Palmer 4B{IT: US-81959]. 
= Muhlenbergia ramulosa (Kunth) Swallen. 


Sporobolus ramosissimus Kunth, Revis. Gramin. 2: 269, t. 
46. 1830. T-Specim.: | of 1. Brazil: St. Paulo [ILT: L]. 
= Eragrostis airoides Nees. 


Sporobolus ramulosus (Kunth) Kunth, Revis. Gramin. 1: 68. 
1829. Basionym: Vilfa ramulosa Kunth. 
= Muhlenbergia ramulosa (Kunth) Swallen. 


Catalogue of New World Grasses: II. 


215 


Sporobolus recurvatus Boechat & Longhi-Wagner, Bradea 
6(20): 177, f. 2. 1993. T-Protol.: Brasil: Mato Grosso: 
Bonito-Rondonopolis, 8 Apr 1930,Chase 11887 [HT: 
SP; IT: US]. T-Specim.: | of 1. Brazil: Matto Gross: 
between Bonito & Rondonopolis, 8 Apr 1930, A. 
Chase 11887 (IT: IT: US-1500497]. 

DST.: Braz. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Boechat, S. C., and H. M. Longhi- 
Wagner (1993). 


Sporobolus reflexus Boechat & Longhi-Wagner, Iheringia, 
Bot. 44: 38, f. 10-11. 1994. T-Protol.: Brasil: Distrito 
Federal: Brasilia, 10 Oct 1990, A.L. Brochado & T. 
Filgueiras 68 [HT: IBGE; IT: ICN]. 

DST.: Braz. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Boechat, S. C. & H. M. Longhi- 
Wagner (1994). 


Sporobolus regis 1.M. Johnst., J. Arnold Arbor. 24: 393. 
1943. T-Protol.: Mexico: Coahuila: Salt flat 4 km. SE 
of Laguna del Rey, abundant, 18 Sep 1942, Stewart 
2653 [HT: GH]. T-Specim.: | of 1. Mexico: Coahuila: 
4 km SE of Laguna Del Rey, 18 Sep 1942, R.M. 
Stewart 2653 (HT: GH; IT: US-90729 (fragm.)]. 

= Sporobolus airoides subsp. regis ([.M. Johnst.) Wipff & 
S.D. Jones. 


Sporobolus repens J. Presi, Relig. Haenk. 1(4-5): 241. 1830. 
T-Protol.: Hab. in Mexico, Haenke s.n. T-Specim.: 1 
of 1. Mexico, Haenke s.n. [HT: PR; IT: LE-TRIN- 
1732.01 (fragm. & fig.), MO, US (fragm.)]. 

= Muhlenbergia repens (J. Presl) Hitche. 


Sporobolus richardsonis (Trin.) Merr., Rhodora 4(39): 46. 
1902. Basionym: Vilfa richardsonis Trin. 
= Muhlenbergia richardsonis (Trin.) Rydb. 


Sporobolus rigens (Trin.) Desv., Fl. Chil. 6: 295. 1854. 
Basionym: Vilfa rigens Trin. 

DST.: Arge, Chil. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Marticorena, C. & M. Quezada 
(1985): Nicora, E. G., M. E. De Paula, A. M. Faggi, 
M. del C. Mariano, A. M. Miante A., L. R. Parodi, C. 
A. Petetin, F. A. Roig & Z. Rugolo de Agrasar 
(1978); Torres, M. A. (1970); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

Diachyrium arundinaceum Griseb.; Diachyrium 
rigens (Trin.) Mez; Epicampes arundinacea (Griseb.) 
Hack. ex Stuck.; Epicampes rigens (Trin.) Phil.; 
Sporobolus arundinaceus (Griseb.) Kuntze; Vilfa 
rigens Trin. 


SYN.: 


Sporobolus rigens fo. atacamensis Parodi, Revista Fac. 
Agron. Veterin. (Buenos Aires) 6(2): 133. 1916. T- 
Protol.: Argentina: Los Andes, Susquis, 5 Mar 1927, 
Castellanos s.n. [HT: BAA-27/766]. 

= Sporobolus rigens var. atacamensis (Parodi) Asteg. 


Sporobolus rigens var. atacamensis (Parodi) Asteg., 
Kurtziana 24: 189. 1995. Basionym: Sporobolus rigens 
fo. atacamensis Parodi. 

DST.: Arge. 
LVL. ACCPTC.:: 1. 
SYN.: Sporobolus rigens fo. atacamensis Parodi. 


Sporobolus rigens var. expansa E. Méndez, Bol. Soc. 
Argent. Bot. 30(1-2): 15-17, f. 2. 1994. T-Protol.: 
Argentina: Mendoza: Depto. La Paz: entre Villa La 
Paz y La Dormida (Santa Rosa), muy abundante, 
fijadora de médanos, 17 Sep 1944, 4.R. Leal Herb. 
FA. Roig 9508 [HT: MERL]. 

DST.: Arge. 

LVL. ACCPTC.: 1. 


Sporobolus rigens (Trin.) Desv. var. rigens. 
LVL. ACCPTC.: 1. 


Sporobolus rupestris Kunth, Revis. Gramin. 1: 267, t. 45. 
1830. T-Protol.: Brazil: Crescit in rimis madidis 
rupium prope novum Friburgum Brasiliae[IT: LE]. T- 
Specim.: | of 1. in humidis petrosis pr. Friburgo 
Brasil., 1836, a cl. Beyrich |. dr. ill. a Roemer [\T: LE- 
TRIN 1737.01 (fig.)]. 

= Sporobolus piliferus (Trin.) Kunth. 


Sporobolus sabeanus Buckley ex Vasey, Contr. U.S. Natl. 
Herb. 3(1): 61. 1892. 
NOTE: nom. inval., as syn. of Sporobolus argutus (Nees) 
Kunth. 
= Sporobolus pyramidatus (Lam.) Hitche. 


Sporobolus sacatilla Griseb. ex E. Fourn., Mexic. Pl. 2: 101. 
1886. 
NOTE: nom. inval., as syn. of Vilfa sacatilla Griseb. ex 
Fourn. 
= Muhlenbergia utilis (Torr.) Hitche. 


Sporobolus sarmentosus Griseb., Abh. Konigl. Ges. Wiss. 
Gottingen 24: 295. 1879. T-Protol.: Argentina: 
Cordoba: Santiago del Estero: Pampa pr. Laguna de 
Pocho. T-Specim.: 1 of 1. Argentina: Pampa pr. 
Laguna de Pocho [T: IT: US]. 

= Muhlenbergia asperifolia (Nees & Meyen ex Trin.) 
Parodi. 


Sporobolus scaber Phil., Fl. Atacam. 54. 1860. T-Protol.: 
Chile: "prope Cachimal de la Costa, m.c., 1900 p.s.m. 
primum inveni, deinde ad Hueso Parado et alia loca 
herbosa litoralis partis deserti vidi, k. Philippi 118. T- 
Specim.: | of 1. Chile: Cachinal, R.A. Philippi 118 
[HT: SGO-45543; IT: BAA-2928 (fragm.), US- 
2891484 (fragm.)]. 

= Eragrostis attenuata Hitchc. 


Sporobolus schaffneri Mez, Repert. Spec. Nov. Regni Veg. 
17(19-30):; 295, 1921. T-Protol.: Mexico: San Luis 
Potosi, Schaffner [HT: US]. T-Specim.: 1 of 1. 
Mexico: San Luis Potosi, no date, W. Schaffner s.n. 
(IT: US-87214(fragm.)]. 

= Sporobolus airoides var. airoides. 


Sporobolus serotinus A. Gray, Manual 577. 1848. 
REPLACED NAME: Agrostis serotina Torr. 
BLOCKING NAME: Agrostis serotina (L.) L. 
= Muhlenbergia uniflora (Muhl.) Fernald. 


Sporobolus shepherdii Vasey, Bull. Torrey Bot. Club 14: 8. 
1887. T-Protol.: Mexico: Chihuahua,£. Palmer 9 (LT: 
US; IL: GH]. T-Specim.: 1 of 1. Mexico: Chihuahua: 
Hacienda San Miguel, near Batopilas, Aug-Nov 1885, 
E. Palmer 9 {LT: LT: US-825391; ILT: US-81961], 
LT designated by Peterson & Annable, Syst. Bot. 15: 
519. 1990, 


216 


Catalogue of New World Grasses: II. 


= Blepharoneuron shepherdii (Vasey) P.M. Peterson & 
Annable. 


Sporobolus silveanus Swallen, J. Wash. Acad. Sci. 31(8): 
350, f. 3. 1941. T-Protol.: USA: Texas: in open woods 
ca. 10 mi NE of Orange, 30 Sep 1940, W. A. Silveus 
6441 [HT: US-98476(NA); IT: TEX]. T-Specim.: | of 
1. USA: Texas: Orange Co: 10 mi NE of Orange on 
US 90, 30 Sep 1940, WA. Silveus 6441 [HT: US- 
2209343; IT: TEX, US-1867557, US-1817963]. 

DST.: USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Correll, D. S. & M. C. Johnston 
(1970); Hitchcock, A. S. (1951); Weakley, A. S. & P. 
M. Peterson (1998). 


Sporobolus simplex Scribn., Bull. Div. Agrostol., U.S.D.A. 
11: 48-49, f. 6. 1898. T-Protol.: USA: Colorado: 
Georgetown, 19 Aug 1895, P.A. Rydberg 2411. T- 
Specim.: | of 1. USA: Colorado: Clear Creek Co.: 
Georgetown, 19 Aug 1895, P.A. Rydberg 2411 [LT: 
NY; ILT: NY, US-556893]. 

= Muhlenbergia filiformis (Thurb. ex S. Watson) Rydb. 


Sporobolus simplex var. thermale Merr., Rhodora 4(39): 48. 
1902. T-Protol.: USA: Montana: Lolo Hot Springs, 17 
Sep 1898, D. Griffiths 302A [HT: US; IT: NY]. T- 
Specim.: | of 1. USA: Montana: Lolo Hot Springs, 17 

Sep 1898, Griffiths 302a [HT: US-556894; IT: NY]. 

= Muhlenbergia filiformis (Thurb. ex S. Watson) Rydb. 


Sporobolus spiciformis Swallen, Proc. Biol. Soc. Wash. 56: 
78. 1943. T-Protol.: Mexico: Coahuila: Puerto del 
Norte, Cuatro Cienegas, 1460 m, 18 Jul 1939, L. H. 
Harvey 1225 [HT: US-1763006]. T-Specim.: 1 of 1. 
Mexico: Coahuila, 18 Jul 1939, L.H. Harvey 1225 
{HT: US-1763006]. 

DST.: Mexi. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000). 


Sporobolus splendens Swallen, Bol. Soc. Bot. Mexico 23: 
35-37, f. 6. 1958 [1959]. T-Protol.: Mexico: Oaxaca: 
in saline soil near Tehuantepec, 7 Jun 1950, J. 7. 
Baldwin 14365 (HT: US-2014792]. T-Specim.: 1 of 1. 
Mexico: Oaxaca, 7 Jun 1950,/.7. Baldwin 14365 (HT: 
US-2014792]. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Breedlove, D. E. (1986); Davidse, 
G., M. Sousa S. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); McVaugh, R. (1983); Pohl, R. 
W., C. G. Reeder & G. Davidse (1994). 


Sporobolus sporobolus (Spreng.) Kuntze, Revis. Gen. PI. 
3(3): 369. 1898. Basionym: Agrostis sporobolus 
Spreng. 

NOTE: nom. illeg.; tautonym. 

= Sporobolus aeneus (Trin.) Kunth. 


Sporobolus sprengelii Kunth, Revis. Gramin. 1: 68. 1829. 
REPLACED NAME: Agrostis sporobolus Spreng. 
= Sporobolus aeneus (Trin.) Kunth. 


Sporobolus stachyanthus A. Rich., Tent. Fl. Abyss. 2: 394. 


1851. 
NOTE: error for S. stachydanthus A. Rich. 


CORRECT NAME: Sporobolus stachydanthus A. Rich. 
= Sporobolus piliferus (Trin.) Kunth. 


Sporobolus stachydanthus A. Rich., Tent. Fl. Abyss. 2: 394. 
1851. T-Protol.: Abyssinia: Crecit in montosis planitiei 
prov. Chiré. T-Specim.: | of 1. Ethiopia: Prov. Chire, 
Oct, Quartin-Dillon s.n. (HT: P; IT: B, K]. 

= Sporobolus piliferus (Trin.) Kunth. 


Sporobolus strictus (Scribn.) Merr., Circ. Div. Agrostol. 
U.S.D.A. 32: 6. 1901. Basionym: Sporobolus 
cryptandrus var. strictus Scribn. 

NOTE: hom. illeg. 
E. NAME: Sporobolus strictus Franch. 
= Sporobolus contractus Hitchc. 


Sporobolus subbulbosus Arechav., Anales Mus. Nac. 
Montevideo 1: 343, t. 35. 1896. T-Protol.: Uruguay: 
"en terrenos elevados, pedregosos, colinas de las 
sierras de las Minas, campos de Mrecedes, estancia del 
Sr. D.M.B. Berro.". 
NOTE: Index of Grass Species 1895. 
= Sporobolus aeneus (Trin.) Kunth. 


Sporobolus subinclusus Phil., Anales Univ. Chile 36: 207. 
1870. T-Protol.: Argentina: Mendoza. T-Specim.: | of 
1. Argentina: Mendoza, no date, R.A. Philippi s.n. {IT: 
US-81962]. 
= Sporobolus cryptandrus (Torr.) A. Gray. 


Sporobolus subinclusus var. expansus Parodi, Revista Fac. 
Agron. Veterin. (Buenos Aires) 6(2): 149, f. 12F. 
1928. T-Protol.: Argentina: Cordoba: dep. Rio IV, 
Carnerillo, Stuckert 20060 [HT: BAA]. 

NOTE: based on Sporobolus subinclusus var. pinicula 
expansa Stuckert, nom. inval. 
= Sporobolus cryptandrus (Torr.) A. Gray. 


Sporobolus subinclusus fo. panicula expansa Stuck., Anales 
Mus. Nac. Hist. Nat. Buenos Aires 21: 93. 1911. 
NOTE: nom. inval., phrase name. 
= Sporobolus cryptandrus (Torr.) A. Gray. 


Sporobolus sundaicus Ohwi, Bull. Tokyo Sci. Mus. 18: 13. 
1947. T-Protol.: Java: Soerabaja, van Slooten 2044 
[HT: BO]. T-Specim.: 1 of 1. Java: Surabaya, near 
Grissee, 29 Sep 1928, Van Slooten (HT: BO; IT: K]. 
= Sporobolus virginicus (L.) Kunth. 


Sporobolus temomairemensis Judz. & P.M. Peterson, Syst. 
Bot. 14(4): 525-528, f. 1. 1989. T-Protol.: Brazil: 
Amapa: Mt. Temomairem, ca. 2°00-20'N, 54°25-35'W, 
elev. ca. 400-450 m, rocky slopes, 26 Jul 1939, 
Geyskes 9 [HT: US; IT: U]. 

DST.: Braz, Suri. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Judziewicz, E. J. & P. M. Peterson 
(1989); Judziewicz, E. J. (1990). 


Sporobolus tenuispica Hack., Repert. Spec. Nov. Regni Veg. 
6(21-26): 344. 1909. T-Protol.: Paraguay: Gran Chaco, 
in campis altis in regione cursus inferiores fluminis 
Pilcomayo, Rojas 258 [HT: Herb. Hassler]. T-Specim..: 
1 of 1. Paraguay: Gran Chaco, in campis altis in 
regione cursus inferiores fluminis Pilcomayo, Jun, 7 
Rojas 258 [HT: W; IT: US-2891485 (fragm.)]. 

= Eragrostis spicata Vasey. 


Catalogue of New World Grasses: II. 


217 


Sporobolus tenuissimus (Mart. ex Schrank) Kuntze, Revis. 
Gen. Pl. 3(3): 369. 1898. Basionym: Panicum 
tenuissimum Mart. ex Schrank. 

DST.: Beli, Braz, Cari, CoRi, Ecua, ElSa, FrGu, Guya, 
Hond, Mexi, Pana, Para, Peru, Suri, USA, Vene. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Baaijens, G. J. 
& J. F. Veldkamp (1991); Balick, M. J., M. H. Nee & 
D. E. Atha (2000); Berendsohn, W. G. & A. E. 
Araniva de Gonzalez (1991); Bor, N. L. (1960); 
Brako, L. & J. L. Zarucchi (1993); Catasus Guerra, 
L. (1997); Catasus Guerra, L. J. (1980); Davidse, G., 
M. Sousa S. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Gould, F. W_ (1979); 
Hitchcock, A. S. (1936); Jorgensen, P. M. & S. Leon- 
Yanez (eds.) (1999); Judziewicz, E. J. (1990); 
Liogier, H. & L. Martorell (1982); Miller, C., G. K. 
Miiller & P. Gutte (1981); Phillips, S. (1995); Pohl, 
R. W. (1983); Pohl, R. W., C. G. Reeder & G, 
Davidse (1994); Renvoize, S. A. (1984); Tovar, O. 
(1993); Wunderlin, R. P. (1998). 

SYN.: Agrosticula muralis Raddi; Aira tenuissima (Matt. ex 
Schrank) Spreng.; Panicum tenuiflorum Schrank; 
Panicum tenuissimum Mart. ex Schrank; Sporobolus 
mangaloricus Hochst. ex Miq.; Sporobolus 
minutiflorus (Trin.) Link; Sporobolus —muralis 
(Raddi) Hitchc. & Chase; Sporobolus tenuissimus 
(Mart. ex Schrank) Hack.; Vilfa mangalorica Hochst. 
ex Steud.; Vilfa minutiflora Trin. 


Sporobolus tenuissimus (Mart. ex Schrank) Hack., Bull. 
Herb. Boissier, ser. 2, 4(3): 278. 1904. Basionym: 
Panicum tenuissimum Mart. ex Schrank. 

NOTE: isonym. 

E. NAME: Sporobolus tenuissimus (Mart. ex Schrank) 
Kuntze. 

= Sporobolus tenuissimus (Mart. ex Schrank) Kuntze. 


Sporobolus teretifolius R.M. Harper, Bull. Torrey Bot. Club 
33: 229-231. 1906. T-Protol.: USA: Georgia: Colquitt 
Co.: near Moultrie, 20 Sep 1902, R.M. Harper 1642. 
T-Specim.: 1 of 1. USA: Georgia: Colquitt Co.: moist 
pine barrens near Moultrie, 20 Sep 1902,R.M. Harper 
1642 [HT: NY; IT: GA, US-431954]. 
DST.: USA. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Hitchcock, A. S. (1951); Radford, A. 
E., H. E. Ahles & C. R. Bell (1968); Small, J. K. 
(1933); Weakley, A. S. & P. M. Peterson (1998). 


Sporobolus texanus Vasey, Contr. U.S. Natl, Herb. 1(2): 57. 
1890. T-Protol.: USA: Texas: Presidio Co.: Screw 
Bean, G.C. Nealley s.n. T-Specim.: | of 1. USA: 
Texas: Presidio Co.: Screw Bean, 1889, G.C. Nealley 
755 [HT: US-556895]. 

DST.: USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Correll, D. S. & M. C. Johnston 
(1970); Great Plains Flora Association (1986): 
Hitchcock, A. S. (1951). 


Sporobolus tharpii Hitche., Proc. Biol. Soc. Wash. 41: 161. 
1928. T-Protol.: USA: Texas: collected on Padre 
Island, 4 Sep 1927, B.C. Tharp 4772 [HT: US- 
1299827]. T-Specim.: | of 1. USA: Texas, 4 Sep 1927, 
B.C. Tharp 4772 (HT: US-1299827]. 


= Sporobolus airoides (Torr.) Torr. 


Sporobolus thurberi Scribn., Bull. Div. Agrostol., U.S.D.A. 
11: 48, f. 5. 1898. 
REPLACED NAME: Sporobolus filiculmis Vasey ex Beal. 
BLOCKING NAME: Sporobolus filiculmis L.H. Dewey. 
= Muhlenbergia thurberi (Scribn.) Rydb. 


Sporobolus torreyanus (Schult.) Nash, Manual 107. 1901. 
Basionym: Agrostis torreyana Schult. 
= Muhlenbergia torreyana (Schult.) Hitche. 


Sporobolus tremulus (Willd.) Kunth, Revis. Gramin. |: 67. 
1829. Basionym: Agrostis tremula Willd. 
NOTE: nom. illeg. superfl. 
= Sporobolus virginicus (L.) Kunth. 


Sporobolus trichodes Hitchc., Contr. U.S. Natl. Herb. 17(3): 
311. 1913. 

REPLACED NAME: Sporobolus capillaris Vasey. 

BLOCKING NAME: Sporobolus capillaris Miq. 

DST.: Mexi. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
McVaugh, R. (1983). 

SYN.: Sporobolus capillaris Vasey. 


Sporobolus tricholepis (Torr.) J.M. Coult., Man. Bot. Rocky 
Mt. 411. 1885. Basionym: Vilfa tricholepis Torr. 
= Blepharoneuron tricholepis (Torr.) Nash. 


Sporobolus tuberculatus Hack., Anales Mus. Nac. Buenos 
Aires 13: 470, t. 13. 1906. T-Protol.: Argentina: Salta: 
Rosario de la Frontera, Lillo 3908 [HT: Stuckert herb. 
arg. 15397]. T-Specim.: | of 1. Argentina: Salta: 
barrancas de los alrededores de Rosario de la Frontera, 
1905, M. Lillo 3908 (IT: US-87217 (fragm.)]. 

= Sporobolus pyramidatus (Lam.) Hitche. 


Sporobolus uniflorus (Muhl.) Scribn. & Merr., U.S. D. A. 
Circ. Div. of Agrostol. 27: 5. 1900. Basionym: Poa 
uniflora Muhl. 

= Muhlenbergia uniflora (Muhl.) Fernald. 


Sporobolus utilis (Torr.) Scribn., Bull. Div. Agrostol., 
U.S.D.A. 17: 171, f. 467. 1899. Basionym: Vilfa utilis 
Torr. 
= Muhlenbergia utilis (Torr.) Hitchc. 


Sporobolus vaginatus (Steud.) Scribn., Bot. Gaz. 21: 15. 
1896. Basionym: Cryptostachys vaginata Steud. 
= Sporobolus vaginiflorus (Torr. ex A. Gray) Alph. Wood. 


Sporobolus vaginiflorus (Torr. ex A. Gray) Alph. Wood, 
Class-book Bot. (ed. 1861) 775. 1861. Basionym:Vilfa 
vaginiflora Torr. ex A. Gray. 

DST.: Cana, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Correll, D. S. & M. C. Johnston 
(1970); Fernald, M. (1950); Gleason, H. A. & A. 
Cronquist (1968); Gleason, H. A. & A. Cronquist 
(1991); Great Plains Flora Association (1986); 
Hickman, J. C. (1993); Hitchcock, A. S. (1951); 
Munz, P. A. (1968); Radford, A. E., H. E. Ahles & Cc. 
R. Bell (1968); Scoggan, H. J. (1978 [1979]); Small, 
J. K. (1933); Wunderlin, R. P. (1998). 

Cryptostachys vaginata Steud., Cryptostachys 
vaginata Steud.; Sporobolus filiculmis L.H. Dewey, 
Sporobolus minor Vasey ex A. Gray; Sporobolus 


SYN.: 


218 


Catalogue of New World Grasses: II. 


ovatus Beal; Sporobolus vaginatus (Steud.) Scribn.; 
Sporobolus vaginiflorus var. inaequalis Fernald: 
Sporobolus vaginiflorus var. minor Scribn. ex 
Chapm.; Vilfa riehlii Steud.; Vilfa vaginiflora Torr. 
ex A. Gray. 


Sporobolus vaginiflorus var.inaequalis Fernald, Rhodora 35: 
109. 1933. T-Protol.: USA: New Hampshire: Concord: 

9 Sep 1901, FW. Batchelder |T?: GH]. 
= Sporobolus vaginiflorus (Torr. ex A. Gray) Alph. Wood. 


Sporobolus vaginiflorus var. minor Scribn. ex Chapm., FI. 
South. U.S. (ed. 3) 598. 1897. 
= Sporobolus vaginiflorus (Torr. ex A. Gray) Alph. Wood. 


Sporobolus vaginiflorus var. neglectus (Nash) Scribn., Bull. 
Div. Agrostol., U.S.D.A. 17: 170. 1901. Basionym: 
Sporobolus neglectus Nash. 

= Sporobolus neglectus Nash. 


Sporobolus vaginiflorus var. neglectus (Nash) Shinners, 
Rhodora 56(662): 29. 1954. Basionym: Sporobolus 
neglectus Nash. 

NOTE: isonym. 

E. NAME: Sporobolus vaginiflorus var. neglectus (Nash) 
Scribn. 

= Sporobolus neglectus Nash. 


Sporobolus vaginiflorus var. ozarkanus (Fernald) Shinners, 
Rhodora 56(662): 29. 1954. Basionym: Sporobolus 
ozarkanus Fernald. 

DST.: Cana, USA. 

LVL. ACCPTC:: 1. 

SYN.: Sporobolus neglectus var. ozarkanus (Fernald) 
Steyerm. & C.L. Kucera; Sporobolus ozarkanus 
Fernald. 


Sporobolus vaginiflorus (Torr. ex A. Gray) Alph. Wood var. 
vaginiflorus. ; 
DST.: Cana, USA. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Fernald, M. (1950); Scoggan, H. J. 
(1978 [1979}). 


Sporobolus villosus (Rchb. ex Spreng.) Kunth, Revis. 


Gramin. 1: 68. 1829. Basionym: Agrostis villosa Rchb. 
ex Spreng. 
= Sporobolus piliferus (Trin.) Kunth. 


Sporobolus virginicus (L.) Kunth, Revis. Gramin, 1: 67. 
1829. Basionym: Agrostis virginica L. 

DST.: Beli, Braz, Cari, Chil, CoRi, Ecua, ElSa, FrGu, Guat, 
Guya, Hond, Mexi, Nica, Pana, Peru, Suri, USA, 
Vene. 

LVL. ACCPTC.: |. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Baaijens, G. J. 
& J. F. Veldkamp (1991); Balick, M. J.,M. H. Nee & 
D. E. Atha (2000); Beetle, A. A. (1977); Bor, N. L. 
(1960); Brako, L. & J. L. Zarucchi (1993); 
Breedlove, D. E. (1986); Catasus Guerra, L. (1997); 
Correll, D. S. & M. C. Johnston (1970); Cowan, C. P. 
(1983); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Fernald, 
M. (1950); Gibbs Russell, G. E, W. G. Welman, E. 
Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987), Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Godfrey, R. 


K. & J. W. Wooten (1979); Gould, F. W (1979); 
Gould, F. W. & R. Moran (1981); Hitchcock, A. S. 
(1909); Hitchcock, A. S. (1927); Hitchcock, A. S. 
(1936); Hitchcock, A. S. (1951); Jorgensen, P. M. & 
S. Leon-Yanez (eds.) (1999); Judziewicz, E. J. 
(1990); Lawesson, J. E., H. Adsersen & P. Bentley 
(1987); Lindeman, J. C. & A. L. Stoffers (1963); 
Liogier, H. & L. Martorell (1982); Long, R. W. & O. 
Lakela (1971); Macbride, J. F. (1936); Marticorena, 
C. & M. Quezada (1985); McVaugh, R. (1983); 
ORSTOM (1988); Pohl, R. W. (1980): Pohl, R. W., 
C. G. Reeder & G. Davidse (1994); Porter, D. M. 
(1983); Radford, A. E., H. E. Ahles & C. R. Bell 
(1968); Reeder, J. R. & C. G. Reeder (1971); 
Renvoize, S. A. (1984); Renvoize, S. A. (1988); 
Shreve, F. & I. L. Wiggins (1964); Simon, B. K. & S. 
W. L. Jacobs (1999); Small, J. K. (1933); Smith, L. 
B., D. C. Wasshausen & R. M. Klein (1981); Sousa 
S., M. & E. F. Cabrera C. (1983); Swallen, J. R. 
(1955); Tovar, O. (1993); Wiggins, LL. & D. M. 
Porter (1971); Wunderlin, R. P. (1998). 

Agrostis barbata Pers.; Agrostis barbata var. 
senegalensis Pers.; Agrostis congener Schumach.; 
Agrostis juncea Lam.; Agrostis littoralis Lam.; 
Agrostis pungens Muhl.; Agrostis pungens Muhl.; 
Agrostis tremula Willd.; Agrostis virginica L.; Aira 
congener Schumach.; Aira sabulonum Labill.; Aira 
sabulonum var. uniflora Balansa; Calotheca sabulosa 
Steud.; Crypsis virginica (L.) Nutt.; Eragrostis 
sabulosa (Steud.) Schweick.; Podosemum virginicum 
(L.) Link; Sporobolus benthamii var. robustus 
Domin; Sporobolus littoralis (Lam.) Kunth; 
Sporobolus littoralis var. elongatus (P. Beauv.) T. 
Durand & Schinz; Sporobolus matrella Nees; 
Sporobolus pungens Kunth; Sporobolus sundaicus 
Ohwi; Sporobolus tremulus  (Willd.) Kunth; 
Sporobolus virginicus (L.) Brongn.; Sporobolus 
virginicus subsp. littoralis (Lam.) Borhidi & O. 
Mufiz; Sporobolus virginicus var. littoralis (Lam.) 
Hitche.; Sporobolus virginicus var. minor F.M. 
Bailey; Sporobolus virginicus var.minor P.M. Bailey 
ex B.K. Simon; Sporobolus virginicus var. pallidus 
Benth.; Sporobolus virginicus var. virginicus; Vilfa 
barbata P. Beauv.; Vilfa intermedia Trin; Vilfa 
littoralis (Lam.) P. Beauv.; Vilfa luxurians Steud. ex 
Lechler; Vilfa matrella Nees: Vilfa tremula Trin.; 
Vilfa virginica (L.) P. Beauv, 


SYN.: 


Sporobolus virginicus (L.) Brongn., Voy. Monde 2: 17. 1829. 
Basionym: Agrostis virginica L. 
NOTE: isonym., 
E. NAME: Sporobolus virginicus (L.) Kunth. 
= Sporobolus virginicus (L.) Kunth. 


Sporobolus virginicus subsp. littoralis (Lam.) Borhidi & O. 
Mufliz, Bot. Kozlem. 58(3): 175. 1971. Basionym: 
Agrostis littoralis Lam. 

NOTE: as Sporobolus virginicus var. "litoralis". 
= Sporobolus virginicus (L.) Kunth. 


Sporobolus virginicus var. littoralis (Lam.) Hitche., N. Amer. 
Fl. 17(7): 486. 1937. Basionym: Agrostis littoralis 
Lam. 
= Sporobolus virginicus (L.) Kunth. 


Catalogue of New World Grasses: II. 


219 


Sporobolus virginicus var. minor F.M. Bailey, Bot. Bull. 
Dept. Agric., Queensland 13: 15. 1896. T-Specim.: 1 
of 1. Australia: Queensland, Bailey s.n. [IT: K]. 

NOTE: nom. nud. accd. to Simon, Austrobailyea 2(1): 22. 
1984; validly published according to Baaijens & 
Veldkamp, Blumea 35: 446 (1991). 

= Sporobolus virginicus (L.) Kunth. 


Sporobolus virginicus var.minor F.M. Bailey ex B.K. Simon, 

Austrobaileya 2(1): 22. 1984. T-Protol.: Australia: 
New South Wales: Central Coast: Cape Solander, 
Botany Bay, Mar. |, 1961, EF. Constable s.n. NSW- 
55195 [HT: BRI; IT: NSW]. 

NOTE: illeg. hom. 

E. NAME: Sporobolus virginicus var. minor F.M. Bailey. 

= Sporobolus virginicus (L.) Kunth. 


Sporobolus virginicus var. pallidus Benth., Fl. Austral. 7: 
621. 1878. T-Specim.: | of 1. Australia: Queensland: 
Brisbane River, Bailey s.n. [LT: K], LT designated by 
Baaijens & Veldkamp, Blumea 35: 446 (1991). 

NOTE: as Sporobolus virginicus var. "pallida". 
= Sporobolus virginicus (L.) Kunth. 


Sporobolus viscidus Sohns, J. Wash. Acad. Sci. 46(12): 385, 
f. 56-61. 1956. T-Protol.; Mexico: México: collected 
en matorral seco, 550 m de altitud, La Junta, cerca de 
Tingambato, 20 Dec 1953, £. Matuda 29813 (HT: US- 
2181966]. T-Specim.: | of 1. Mexico: México: La 
Junta, 20 Dec 1953, E. Matuda 29813 [HT: US- 
2181966; IT: US-2206400]. 

DST.: Mexi. 
LVL. ACCPTC:: I. 
ALSO ACCPTD. BY: Sohns, E. R. (1956). 


Sporobolus wolfii Vasey, Bull. Torrey Bot. Club 10: 52. 
1883. T-Protol.: USA: Colorado: Twin Lakes, 1873, 
John Wolf s.n. 
= Muhlenbergia ramulosa (Kunth) Swallen. 


Sporobolus wrightii Munro ex Scribn., Bull. Torrey Bot. 
Club 9: 103. 1882. T-Protol.: USA: Arizona: Near 
Pantano, Jun 1881, Pringle 190. T-Specim.: 1 of 1. 
USA: Arizona, 28 Jun 1881, C.G. Pringle 190 [IT: 
US-825415]. 

DST.: Mexi, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Bor, N. L. (1960); Correll, D. S. & 
M. C. Johnston (1970); Hitchcock, A. S. (1951); 
Shreve, F. & J. L. Wiggins (1964). 

Bauchea karwinskyi EE. Fourn.; &Epicampes 
crassiculmis Piper; Sporobolus airoides var. minor 
(Vasey) Beetle; Sporobolus airoides var. wrightii 
(Munro ex Scribn.) Gould; Sporobolus altissimus var. 


SYN:: 


minor WNasey; Sporobolus altissimus Vasey; 
Sporobolus eminens J. Presl; Sporobolus expansus 
Scribn. 


Sporobolus sect. Agrosticula (Raddi) Veldkamp, Blumea 
35(2): 399. 1991. Basionym: Agrosticula Raddi. 
LVL. ACCPTC.: 1. 


Sporobolus sect. Fimbriatae Veldkamp, Blumea 35(2): 399. 
1991. TYPE: Sporobolus fimbriatus (Trin.) Nees. 
LVL. ACCPTC.: 1. 


Sporobolus Nees sect. Sporobolus. 
LVL. ACCPTC:: 1. 


Sporobolus sect. Triachyrum (Hochst. ex A. Braun) 
Veldkamp, Blumea 35(2): 398. 1991. Basionym: 
Triachyrum Hochst. ex A. Braun. 

LVL. ACCPTC:: I. 


Sporobolus sect. Virginicae Veldkamp, Blumea 35(2): 398. 
1991. TYPE: Sporobolus virginicus (L.) Kunth. 
LVL. ACCPTC.: I. 


Stapfia Burtt Davy, Erythea 6: 109. 1898. 
NOTE: hom. illeg. 
= Neostapfia Burtt Davy. 


Stapfia colusana Burtt Davy, Erythea 6: 110, t. 3. 1898. T- 
Specim.: | of 1. USA: California: Colusa Co.: near 
Princeton, 28 May 1898, /. Burtt Davy s.n. [HT: UC; 
IT: K, US-2947514]. 

= Neostapfia colusana (Burtt Davy) Burtt Davy. 


Steirachne Ekman, Ark. Bot. 10(17): 35, t. 5, f. 1, t. 6, 
f. 17, text f. 1-2. 1911. TYPE: Steirachne diandra 
Ekman. 

TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP: 2. 


Steirachne barbata (Trin.) Renvoize, Kew Bull. 39(1): 184. 
1984. Basionym: Eragrostis barbata Trin. 
DST.: Braz, Guya, Vene. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Filgueiras, T. S. (1999); Judziewicz, 
E. J. (1990); Renvoize, S. A. (1984). 
SYN.: Eragrostis barbata Trin., Eragrostis quitensis Trin. 


Steirachne diandra Ekman, Ark. Bot. 10(17): 36, t. 5, f. 1, t. 
6, f. 17, text f. 1-2. 1911. T-Specim.: 1 of 2. Brazil, 9 
Apr 1910, Loefgren 488 [ST: S]. 2 of 2. Brazil, 9 Apr 
1910, Loefgren 489 [ST: S]. 
DST.: Braz, Vene. 
SYN.: Festuca pilosa Nees; Festuca quitensis Willd. ex 
Nees; Steirachne pilosa Ekman. 


Steirachne pilosa Ekman, Ark. Bot. 10(17): 38. 1911. 
NOTE: in obs., nom. inval., as syn. 8. diandra. 
= Steirachne diandra Ekman. 


Stiburus Stapf, Fl. Cap. 7(4): 696-697. 1900. 
REPLACED NAME: Triphlebia Stapf. 
BLOCKING NAME: Triphlebia Baker. 
= Eragrostis Wolf. 


Stipa capillaris Lam., Tabl. Encycl. 1: 158. 1791. T-Protol.: 
E. Carolina, D. Fraser. T-Specim.: 1 of 2. USA: E. 
Carolina, D. Fraser sn. [HT: P-LAM; IT: US- 
A866136 (fragm. ex P-LAM)]. 2 of 2. 

= Muhlenbergia capillaris (Lam.) Trin. 


Stipa diffusa Walter, Fl. Carol. 78. 1788. T-Protol.: USA: 
South Carolina. 
= Muhlenbergia capillaris (Lam.) Trin. 


Stipa expansa Willd. ex Steud., Nomencl. Bot. (ed. 2) 2: 643. 


1841. 

NOTE: nom. inval., as syn. of Gymnopogon racemosus P. 
Beauv. 

= Gymnopogon ambiguus (Michx.) Britton, Sterns & 
Poggenb. 


Stipa sericea Michx., Fl. Bor.-Amer. 1: 54. 1803. T-Protol.: 
USA: Hab. in Carolina, Georgia maritima, Michaux 


220 


Catalogue of New World Grasses: II. 


s.n. [HT: P]. T-Specim.: | of 1. USA: South Carolina, 
no date, 4. Michaux s.n. [HT: P-MICHX; IT: US- 
866059 (fragm.)]. 

REPLACED NAME: Stipa cericea Michx. ex Raf. 

= Muhlenbergia sericea (Michx.) P.M. Peterson. 


Strombodurus Steud., Nomencl. Bot. (ed. 2) 2: 299. 1841. 
NOTE: nom. inval., as syn. of Pentarrhaphis scabra. 
= Bouteloua Lag. 


Swallenia Soderstr. & H.F. Decker, Madrofio 17(3): 88. 
1963. 
TRTD. BY: Peterson, P. M. 
NHT: Monanthochloinae. NO. ACCPTD. SPP.: 1. 
SYN.: Ectosperma  alexandrae Swallen; Ectosperma 
Swallen; Swallenia alexandrae (Swallen) Soderstr. & 
H.F. Decker. 


Swallenia alexandrae (Swallen) Soderstr. & H.F. Decker, 
Madrofio 17(3): 88. 1963. Basionym: Ectosperma 
alexandrae Swallen. 

DST.: USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Hickman, J. C. 
(1993); Munz, P. A. (1974); Soderstrom, T. R. & H. 
F. Decker (1963). 


Symbasiandra Steud., Nomencl. Bot. (ed. 2) 1: 767. 1840. 
NOTE: nom. inval., in syn. sub Hilaria cenchroides Kunth. 
= Hilaria Kunth. 


Tetrachne Nees, Fl. Afr. Austral. Ill. 375. 13-19 Jun 
1841. TYPE: Tetrachne dregei Nees. 
TRTD. BY: Peterson, P. M. 
NHT: Uniolinae. NO. ACCPTD. SPP.: 1. 


Tetrachne dregei Nees, Fl. Afr. Austral. Ill. 376. 1841. T- 
Protol.: In planitie  graminosa ad flumen 
Klipplaatsrivier alt. 3500' inter gramina, inter 
Compasberg et Rhinosterbergen montes alt. 4500 - 
5000", ad Wonderheuvel alt. 4500', ad zwartkey 
fluvium alt. 4000', ad radices montium Witbergen alt. 
5000', Decembri, [anuario.,, Drege s.n. 

NOTE: cites Poa glomerata Thunb., Prodr. 22 [non Walt. ]. 

NOTE: Cultivated and tested in Villa Mercedes, San Louis 
Argentina. 

DST.: Arge. 

LVL. ACCPTC.: 1; cultivated, introduced. 

ALSO ACCPTD. BY: Gibbs Russell, G. E, W. G. Welman, 
E. Reitief, K. L. Immelman, G. Germishuizen, B. J. 
Pienaar, M. van Wyk, and A. Nicholas (PRECIS) 
(1987). 


Thellungia Stapf, Bull. Misc. Inform. Kew 1920(3): 97, f. 1- 
11. 1920. TYPE: Thellungia advena Stapf. 
= Eragrostis Wolf. 


Torgesia Bornm., Mitth. Thuring. Bot. Vereins 30: 83. 1913. 
TYPE: Torgesia minuartioides Bornm. 
= Crypsis Aiton. 


Tosagris P. Beauv., Ess. Agrostogr. 29. 1812. TYPE: 
Tosagris agrostidea P. Beauv. 
NOTE: nom. prov. 


= Muhlenbergia Schreb. 


Tosagris agrostidea P. Beauv., Ess. Agrostogr. 29, t. 8, f. 3. 
1812. T-Protol.: USA: "Etats-Unis d'Amerique". 
= Muhlenbergia capillaris (Lam.) Trin. 


Trachynotia Michx., Fl. Bor.-Amer. 1: 63. 1803. 
NOTE: nom. superfl pro Spartina. 
= Spartina Schreb. 


Trachynotia alterniflora (Loisel.) DC., Fl. Franc. 5: 279. 
1815. Basionym: Spartina alterniflora Loisel. 
= Spartina alterniflora Loisel. 


Trachynotia alterniflora Steud., Nomencl. Bot. (ed. 2) 2: 695. 
1841. 
NOTE: orth. error for Spartina alterniflora Steudel. 
= Spartina alterniflora Loisel. 


Trachynotia cynosuroides (L.) Michx., Fl. Bor.-Amer. 1: 64. 
1803. Basionym: Dactylis cynosuroides L. 
= Spartina cynosuroides (L.) Roth. 


Trachynotia juncea Michx., Fl. Bor.-Amer. 1: 64. 1803. T- 
Specim.: | of 1. USA: bords des creeks sales Basse 
Caroline, Michaux s.n. [HT: P]. 

= Spartina patens (Aiton) Muhl. 


Trachynotia polystachya Michx., Fl. Bor.-Amer. 1: 64. 1803. 
T-Specim.: | of 1. USA: Basse Carolina, Michaux s.n. 
[HT: P]. 
= Spartina cynosuroides (L.) Roth. 


Trageae trib. Hitche., Contr. U.S. Natl. Herb. 24: 559. 1927. 
TYPE: Tragus Haller. 


Tragus Haller, Hist. Stirp. Helv. 2: 203. 1768. TYPE: 
Tragus racemosus (L.) All. 
NOTE: nom. cons. 
TRTD. BY: Peterson, P. M. 
NHT: Zoysiinae. NO. ACCPTD. SPP.: 4. 
CNSLTD. LIT.: Anton, A. M., (1981). 
SYN.: Echinanthus Cerv.; Lappago Schreb.; Nazia Adans. 


Tragus alienus (Spreng.) Schult., Mant. 2: 205. 1824. 
Basionym: Lappago aliena Spreng. 


= Tragus berteronianus Schult. 


Tragus australianus S.T. Blake, Univ. Queensland Pap., 
Dept. Bot. 1(18): 12. 1941. T-Protol.: Queensland— 
Maranoa District.Noondoo, near Dirranbandi, on sand 
in Callitris forest, also on heavier soils, ca. 550 ft., 
Blake 10660. 

DST.: Arge, Boli, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Renvoize, S. A. (1998); Zuloaga, F. 
O., E. G. Nicora, Z. E. Rugolo de Agrasar, O. 
Morrone, J. Pensiero & A. M. Cialdella (1994), 


Tragus berteronianus Schult., Mant. 2: 205. 1824. T- 
Specim.: | of 1. Dominican Republic: Santo Domingo, 
Bertero s.n. (HT: B; IT: M, US-92008 (fragm. ex B)}. 

DST.: Arge, Boli, Braz, Cari, Ecua, Hond, Mexi, Peru, 
USA, Vene. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Adams, C. D. (1972); Beetle, A. A. 
(1977); Bor, N. L. (1960); Catasus Guerra, L. (1997); 
Correll, D. S. & M. C. Johnston (1970); Davidse, G. 
(1994): Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Gibbs 


Catalogue of New World Grasses: II. 


221 


Russell, G. E, W. G. Welman, E. Reitief, K. L. 
Immelman, G. Germishuizen, B. J. Pienaar, M. van 
Wyk, and A. Nicholas (PRECIS) (1987); Gleason, H. 
A. & A. Cronquist (1968); Gould, F. W (1979); 
Gould, F. W. & R. Moran (1981); Hitchcock, A. S. 
(1936); Hitchcock, A. S. (1951); Jorgensen, P. M. & 
C. Ulloa U. (1994); Jorgensen, P. M. & S. Leon- 
Yanez (eds.) (1999); Lindeman, J. C. & A. L. 
Stoffers (1963); Liogier, H. & L. Martorell (1982); 
McVaugh, R. (1983); Phillips, S. (1995); Renvoize, 
S. A. (1984); Renvoize, S. A. (1998); Shreve, F. & I. 
L. Wiggins (1964); Tovar, O. (1993); Zuloaga, F. O., 
E. G. Nicora, Z. E. Rugolo de Agrasar, O. Morrone, 
J. Pensiero & A. M. Cialdella (1994). 

Lappago aliena Spreng.,; Lappago berteroniana 
Schult. ex Steud.; Lappago occidentalis Nees ex 
Hook.; Nazia aliena (Spreng.) Scribn.; Nazia 
occidentalis (Nees) Scribn.; Nazia racemosa var. 
aliena (Spreng.) Scribn. & J.G. Sm.; Tragus alienus 
(Spreng.) Schult.; Tragus occidentalis Nees; Tragus 
racemosus fo. erectus D6ll; Tragus racemosus var. 
berteronianus (Schult.) Hack.; Tragus racemosus 
var. brevispicula Doll. 


SYN:: 


Tragus echinatus (L.) Cav., Elench. Pl. 38. 1803. 
NOTE: cf. Rothm., Fedde, Repert. 49: 53, 1940. 
= Tragus racemosus (L.) All. 


Tragus heptaneuron Clayton, Kew Bull. 27(1): 151. 1972. 
T-Protol.: Kenya: Samburu, 13 Nov 1952, Bogdan 
3619 (HT: K]. 

DST.: USA. 
LVL. ACCPTC.: 1; introduced. 


Tragus muricatus Moench, Methodus 53. 1794. Basionym: 
<>. 
NOTE: nom. illeg. superfl., based on Cenchurs racemosus 
L. 
CORRECT NAME: Tragus racemosus (L.) All. 
= Tragus racemosus (L.) All. 


Tragus occidentalis Nees, Fl. Bras. Enum, Pl. 2: 286. 1829. 
T-Specim.: | of 1. Brazil: Bahia, Martius s.n. IT: K, 
US (fragm. ex Munich)]. 
= Tragus berteronianus Schult. 


Tragus racemosus (L.) All., Fl. Pedem. 2: 241. 1785. 
Basionym: Cenchrus racemosus L. 

DST.: Arge, Cari, Peru, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Baldini, R. M. (1995); Brako, L. & J. 
L. Zarucchi (1993); Catasus Guerra, L. (1997); 
Catasus Guerra, L. J. (1980); Correll, D. S. & M. C. 
Johnston (1970); Fabris, H. A. (1970); Gibbs Russell, 
G. E, W. G. Welman, E. Reitief, K. L. Immelman, G. 
Germishuizen, B. J. Pienaar, M. van Wyk, and A. 
Nicholas (PRECIS) (1987); Gleason, H. A. & A. 
Cronquist (1968); Hitchcock, A. S. (1951); Phillips, 
S. (1995); Tovar, O. (1993); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

Cenchrus racemosus L., Lappago racemosa (L.) 
Honck.; Nazia racemosa (L.) Kuntze; Phalaris 
muricata Forssk.; Tragus echinatus (L.) Cav.; Tragus 
muricatus Moench; Tragus racemosus fo. divaricatus 
D6Il; Tragus racemosus var. longispicula DOll. 


SYN.: 


Tragus racemosus var. berteronianus (Schult.) Hack., 
Oesterr. Bot. Z. 51(6): 195. 1901. Basionym: Tragus 
berteronianus Schult. 

= Tragus berteronianus Schult. 


Tragus racemosus vat. brevispicula Doll, Fl. Bras. 2(2): 123. 
1877. T-Specim.: 1 of |. Brazil, Martius s.n. [IST: K]. 
NOTE: third name in the quadrinomials fo. erectus and fo. 
expansus. 
= Tragus berteronianus Schult. 


Tragus racemosus fo. divaricatus DOll, Fl. Bras. 2(2): 122. 
1877. T-Protol.: regionis mediterraneis. 
= Tragus racemosus (L.) All. 


Tragus racemosus fo. erectus Doll, Fl. Bras. 2(2): 123. 1877. 
T-Protol.: Brazil: Prov. Piauhyensi, in aridis apricis 
Prov. Bahiensis, Gardner 2340; Martius s.n., 2 
syntypes listed. 

= Tragus berteronianus Schult. 


Tragus racemosus vat. longispicula DOll, Fl. Bras. 2(2): 122. 
1877. 
NOTE: third name of the quadrinomial fo. divaricatus and 
fo. erectus. 
= Tragus racemosus (L.) All. 


Triachyrum Hochst. ex A. Braun, Flora 24: 712. 1841. 
TYPE: Triachyrum adoense Hochst. ex Braun, A. 
= Sporobolus R. Br. 


Triachyrum nilagiricum Hochst. ex Steud., Syn. Pl. Glumac. 
1: 176. 1854. T-Specim.: 1 of 1. India: Orissa: 
Nilgirris, 1851, R.F. Hohenacker 93] [HT: P; IT: B, 
BM, K, L, US-1127182, US-2947339 (fragm.), W]. 

= Sporobolus piliferus (Trin.) Kunth. 


Triachyrum stachydanthum (A. Rich.) Asch., Beitr. FI. 
Aethiop. 302, 311. 1867. Basionym: Sporobolus 
stachydanthus A. Rich. 

NOTE: nom. inval., Art. 34. 1.a. 

= Sporobolus piliferus (Trin.) Kunth. 


Triaena Kunth, Nov. Gen. Sp. 1: ed qu. 178. 1815 [1816]. 
TYPE: Triaena racemosa Kunth. 
= Bouteloua Lag. 


Triaena juncea (Desv. ex P. Beauv.) Griffiths, Contr. U.S. 
Natl. Herb. 14(3): 354. 1912. Basionym: Triathera 
juncea Desv. ex P. Beauv. 

= Bouteloua juncea (Desv. ex P. Beauv.) Hitche. 


Triaena racemosa Kunth, Nov. Gen. Sp. 1: 179, t. 61. 1815 
[1816]. T-Protol.: Inter Guanaxuato Mexicanorum et 
Villalpando, Humboldt & Bonpland s.n. T-Specim.: 1 
of 1. MexicO; Guanajuato: between Guanajuato and 
Villalpando, Humboldt & Bonpland s.n. (HT: P]. 

= Bouteloua triaena (Trin. ex Spreng.) Scribn. 


Triathera Desv., Nouv. Bull. Sci. Soc. Philom. Paris 2: 188. 
1810. TYPE: 7riathera americana (L.) Desv. 
= Bouteloua Lag. 


Triathera americana (L.) Desv., Nouv. Bull. Sci. Soc. 
Philom. Paris 2: 188. 1810. Basionym: Aristida 
americana L. 

= Bouteloua americana (L.) Scribn. 


Triathera aristidoides (Kunth) Nash, Fl. S.E. U.S. 137. 1903. 
Basionym: Dinebra aristidoides Kunth. 
= Bouteloua aristidoides (Kunth) Griseb. 


222 


Catalogue of New World Grasses: II. 


Triathera gracilis E. Fourn., Mexic. Pl. 2: 141. 1886. T- 
Specim.: | of 3. Mexico: Cuernavaca, Berl /0/6 [ST]. 
2 of 3. Mexico: Chapulco, Nov, Liebmann 578 [ST]. 3 
of 3. Mexico: San Lorenzo pr. Tehuacan, Nov., 
Liebmann 582 [ST]. 
= Bouteloua triaena (Trin. ex Spreng.) Scribn. 


Triathera juncea Desy. ex P. Beauv., Ess. Agrostogr. 40, t. 8, 
f. 4. 1812. T-Protol.: Hispanola[Known only from 
illustration possibly at G]. 

= Bouteloua juncea (Desv. ex P. Beauv.) Hitche. 


Triathera racemosa (Kunth) Desv., Mem. Soc. Agric. Angers 
1: 176. 1831. Basionym: Triaena racemosa Kunth. 
NOTE: also in Opusc. Sci. Phys. Nat. 72 [reprint] 1831- 
1833. 
= Bouteloua triaena (Trin. ex Spreng.) Scribn. 


Triatherus Raf., Amer. Monthly Mag. & Crit. Rev. 3: 99. 
1818. 

NOTE: non Triathera Desvaux 1810, nom. superfl. pro 
Monocera. 

= Ctenium Panz. 


Triatherus aromaticus Raf., Amer. Monthly Mag. & Crit. 
Rev. 3: 99. 1818. T-Protol.: USA: South Carolina. 
NOTE: based on Monocera aromatica Elliot. 
= Ctenium aromaticum (Walter) Alph. Wood. 


Trichloris &. Fourn. ex Benth., J. Linn, Soc., Bot. 19: 
102. 1881. TYPE: Trichloris pluriflora E. Fourn.; LT 
designated by Hitchcock, U.S. Dept. Agric. Bull. 772: 
190 (1920). 

TRID. BY: Peterson, P. M. 

NHT: Cynodonteae. NO. ACCPTD. SPP.: 2 SUBSP.: 2. 

SYN.: Chloridopsis Hack., Chloropsis Hack. ex Kuntze; 
Leptochloris Kuntze. 


Trichloris blanchardiana E. Fourn. ex Scribn., Bull. Torrey 
Bot. Club 9: 146. 1882. T-Protol.: USA: Arizona: 
Santa Cruz Valley, near Tuscon, 19 May, Pringle [HT: 
US]. T-Specim.: | of 1. USA: Arizona: Santa Cruz 
Valley near Tuscon, 19 May 1881,C.G. Pringle 1392] 
[IT: US-879469; IT: MO-1837535]. 

= Trichloris crinita (Lag.) Parodi. 


Trichloris crinita (Lag.) Parodi, Revista Argent. Agron. 
14(1): 63. 1947. Basionym: Chloris crinita Lag. 

DST.: Arge, Boli, Braz, Chil, Mexi, Para, Peru, USA, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Bor, N. L. (1960); Brako, L. & J. L. 
Zarucchi (1993); Burkart, A. (1969); Cabrera, A. L. 
(1970); Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. L6pez-Ferrari, and J. Valdes- 
Reyna (2000); Hitchcock, A. S. (1951); Marticorena, 
C. & M. Quezada (1985); Nicora, E. G., M. E. De 
Paula, A. M. Faggi, M. del C. Mariano, A. M. Miante 
A., L. R. Parodi, C. A. Petetin, F. A. Roig & Z. 
Rugolo de Agrasar (1978); Renvoize, S. A. (1998); 
Rosengurtt, B., B. R. Arrillaga de Maffei & P. 
Izaguirre de Artucio (1970); Shreve, F. & I. L. 
Wiggins (1964); Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 


Cialdella (1994). 
SYN.: Chloris crinita Lag.; Chloris mendocina Phil.; 
Chloris trichodes Lag. ex Parodi; Chloropsis 


blanchardiana (E. Fourn. ex Scribn.) Kuntze; 


Chloropsis — blanchardiana Gay ex _— Scribn.; 
Chloropsis crinita (Lag.) Kuntze; Chloropsis 
mendocina (Phil.) Kuntze; Leptochloris crinita (Lag.) 
Munro ex Kuntze; Leptochloris greggii Munro ex 
Merr.; Trichloris blanchardiana E. Fourn. ex Scribn.; 
Trichloris crinita var. triflora Parodi; Trichloris 
crinita var. typica Parodi; Trichloris fasciculata E. 
Fourn.; Trichloris mendocina (Phil.) — Kurtz; 
Trichloris mendocina (Phil.) Kurtz ex Seckt; 
Trichloris mendocina fo. blanchardiana (E. Fourn. 
ex Scribn.) Kurtz; 7richloris verticillata E. Fourn. ex 
Vasey; Trichloris verticillata E. Fourn. ex Vasey. 


Trichloris crinita var. triflora Parodi, Revista Argent. Agron. 
14: 63. 1947. T-Protol.: Argentina: San Luis, Laguna 
Bebedero, Jan 1925, leg. R. Guifiazui [HT: BAA-7255]. 

= Trichloris crinita (Lag.) Parodi. 


Trichloris crinita var. typica Parodi, Revista Argent. Agron. 
14: 63. 1947. 
NOTE: nom. inval., see ICBN Art 24.3. 
= Trichloris crinita (Lag.) Parodi. 


Trichloris fasciculata E. Fourn., Mexic. Pl. 2: 142. 1886, T- 
Protol.: Mexico: San Luis Potosi, Virlett 1440 [HT: P]. 
NOTE: Fourn. ex Benth., nom. nud, J. Linn. Soc. Bot., 
London 19: 103. 1881. 
= Trichloris crinita (Lag.) Parodi. 


Trichloris latifolia Vasey, Dept. Agric. Special Rep. 63: 32. 
1883. 
NOTE: nom. nud. 
= Trichloris pluriflora E. Fourn. 


Trichloris lilloi Parodi, Physis (Buenos Aires) 9: 256, f. A-E. 
1928. T-Protol.: Jorgensen 1396. T-Specim.: 1 of 1. 
Argentina, Jorgensen 1396 [HT: ?: IT: BAF]. 

= Trichloris pluriflora E. Fourn. 


Trichloris mendocina (Phil.) Kurtz, Mem. Fac. Ci. Exact. Fis. 
Nat. Univ. Nac. Cordoba 1896: 37. 1897. Basionym: 
Chloris mendocina Phil. 

= Trichloris crinita (Lag.) Parodi. 


Trichloris mendocina (Phil.) Kurtz ex Seckt, Revista Univ. 
Nac. Cordoba 16-17: 156. 1929-30. Basionym:Chloris 
mendocina Phil. 

NOTE: isonym. 
= Trichloris crinita (Lag.) Parodi. 


Trichloris mendocina fo. blanchardiana (E. Fourn. ex 
Scribn.) Kurtz, Bol. Acad. Ci. (Cordoba) 16: 270. 
1900. Basionym: Trichloris blanchardiana E. Fourn. 
ex Scribn. 

= Trichloris crinita (Lag.) Parodi. 


Trichloris pluriflora E. Fourn., Mexic. Pl. 2: 142. 1886. T- 
Specim.: | of 2, USA: Texas: inter Laredo et Bejar, 
Berlandier 1430 [LT: P], LT designated by Hitchcock, 
Contr. U.S. Natl. Herb. 17: 335 (1917). 2 of 2. Mexico, 
Karwinsky s.n. (ST: LE]. 

DST.: Arge, Boli, Cari, Ecua, Guat, Mexi, Para, Peru, USA, 
Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Bor, N. L. (1960); Brako, L. & J. L. 
Zarucchi (1993); Burkart, A. (1969); Cabrera, A. L. 
(1970); Caro, J. A. (1981); Catasus Guerra, L. 
(1997); Correll, D. S. & M. C. Johnston (1970); 
Davidse, G. (1994); Davidse, G., M. Sousa S. & A. 


Catalogue of New World Grasses: II. 


223 


O. Chater (editores generales) (1994); Espejo Serna, 
A., A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Foster, R. C. (1958); Hitchcock, A. S. (1927); 
Hitchcock, A. S. (1936); Hitchcock, A. S. (1951); 
Jorgensen, P. M. & S. Ledn-Yanez (eds.) (1999); 
Macbride, J. F. (1936); Renvoize, S. A. (1998); 
Tovar, O. (1993); Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 

SYN.: Chloris pluriflora (E. Fourn.) Clayton; Chloris 
pluriflora (E. Fourn.) Kuntze; Chloropsis pluriflora 
(E. Fourn.) Kuntze; Trichloris latifolia Vasey; 
Trichloris lilloi Parodi. 


Trichloris pluriflora fo. macra Hack., Anales Mus. Nac. 
Buenos Aires 11: 116. 1904. T-Protol.: Argentina, 
Stuckert: Herb, arg. no. 12,921. 

DST.: Arge. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 


Trichloris pluriflora Hack. fo. pluriflora. 
LVL. ACCPTC.: I. 


Trichloris verticillata E. Fourn. ex Vasey, Descr. Cat. Grass. 
U.S. 61. 1885. 
NOTE: nom. nud. 
= Trichloris crinita (Lag.) Parodi. 


Trichloris verticillata E. Fourn. ex Vasey, U.S.D.A. Div. Bot. 
Bull. 12(2): t. 25. 1891, 
= Trichloris crinita (Lag.) Parodi. 


Trichochloa P. Beauv., Ess. Agrostogr. 29. 1812. TYPE: 
Trichochloa purpurea P. Beauv. 
= Muhlenbergia Schreb. 


Trichochloa DC., Cat. Pl. Horti Monsp. 151. 1813. 
NOTE: nom. illeg. superfl. 
= Muhlenbergia Schreb. 


Trichochloa calycina Trin., Fund. Agrost. 117. 1820. 
REPLACED NAME: Agrostis setosa (Kunth) Spreng. 
BLOCKING NAME: Agrostis setosa Muhl. 
= Muhlenbergia glomerata (Willd.) Trin. 


Trichochloa capillaris (Lam.) DC., Cat. Pl. Horti Monsp. 
152. 1813. Basionym: Stipa capillaris Lam. 
= Muhlenbergia capillaris (Lam.) Trin. 


Trichochloa ciliata (Kunth) Roem. & Schult., Syst. Veg. 2: 
386. 1817. Basionym: Podosemum ciliatum Kunth. 
= Muhlenbergia ciliata (Kunth) Trin. 


Trichochloa debilis (Kunth) Roem. & Schult., Syst. Veg. 2: 
385. 1817. Basionym: Podosemum debile Kunth. 
= Muhlenbergia microsperma (DC.) Kunth. 


Trichochloa elegans (Kunth) Roem. & Schult., Syst. Veg. 2: 
387. 1817. Basionym: Podosemum elegans Kunth. 
= Muhlenbergia rigida (Kunth) Kunth. 


Trichochloa expansa (Poir.) DC., Cat. Pl. Horti Monsp. 151. 


1813. Basionym: Stipa expansa Poir. 
= Muhlenbergia expansa (Poir.) Trin. 


Trichochloa glomerata (Willd.) Trin., Fund. Agrost. 117. 


1820. Basionym: Polypogon glomeratus Willd. 
= Muhlenbergia glomerata (Willd.) Trin. 


Trichochloa gracilis (Kunth) Roem. & Schult., Syst. Veg. 2: 
389. 1817. Basionym: Podosemum gracile Kunth. 
= Muhlenbergia quadridentata (Kunth) Trin. 


Trichochloa implicata (Kunth) Roem. & Schult., Syst. Veg. 
2: 385. 1817. Basionym: Podosemum implicatum 
Kunth. 

= Muhlenbergia implicata (Kunth) Trin. 


Trichochloa longiseta Trin., Fund. Agrost. 117. 1820. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Agrostis tenuiflora Willd. 
= Muhlenbergia tenuiflora (Willd.) Britton, Sterns & 
Poggenb. 


Trichochloa mexicana (L.) Trin., Fund. Agrost. 117. 1820. 
Basionym: Agrostis mexicana L. 
= Muhlenbergia mexicana (L.) Trin. 


Trichochloa microsperma DC., Cat. Pl. Horti Monsp. 151. 
1813. T-Specim.: | of 1. Mexico: cult. at Montpellier, 
from seeds collected in Mexico and disributed by the 
Botanical Garden of Madrid, Sésse & Mocifio s.n. 
[HT: MPU; IT: P, US (fragm. ex P)]. 

= Muhlenbergia microsperma (DC.) Kunth. 


Trichochloa mucronata (Kunth) Roem. & Schult., Syst. Veg. 
2: 387. 1817. Basionym: Podosemum mucronatum 
Kunth. 

= Muhlenbergia mucronata (Kunth) Trin. 


Trichochloa polypogon DC., Cat. Pl. Horti Monsp. 152. 
1813. T-Protol.: USA: Carolina, fraser s.n. 
= Muhlenbergia capillaris (Lam.) Trin. 


Trichochloa purpurea P. Beauv., Ess. Agrostogr. 29, t. 8, f. 2. 
1812. T-Protol.: USA: "Etats-Unis d'Amerique". 
= Muhlenbergia expansa (Poir.) Trin. 


Trichochloa quadridentata (Kunth) Roem. & Schult., Syst. 
Veg. 2: 388. 1817. Basionym: Podosemum 
quadridentatum Kunth. 

= Muhlenbergia quadridentata (Kunth) Trin. 


Trichochloa rigida (Kunth) Roem. & Schult., Syst. Veg. 2: 
386. 1817. Basionym: Podosemum rigidum Kunth. 
= Muhlenbergia rigida (Kunth) Kunth. 


Trichochloa setosa (Kunth) Roem. & Schult., Syst. Veg. 2: 
386. 1817. Basionym: Podosemum setosum Kunth. 
= Muhlenbergia microsperma (DC.) Kunth. 


Trichochloa sobolifera (Muhl. ex Willd.) Trin., Fund. Agrost. 
117. 1820. Basionym: Agrostis sobolifera Muhl. ex 
Willd. 

= Muhlenbergia sobolifera (Muhl. ex Willd.) Trin. 


Trichochloa spicata (Spreng.) Schult., Mant. 3: 577. 1824. 
Basionym: Polypogon spicatus Spreng. 
= Gymnopogon spicatus (Spreng.) Kuntze. 


Trichochloa tenella (Kunth) Roem. & Schult., Syst. Veg. 2: 
385. 1817. Basionym: Podosemum tenellum Kunth. 
= Muhlenbergia tenella (Kunth) Trin. 


Trichochloa tenuiflora (Willd.) Sweet, Hort. Brit. 443. 1826. 
Basionym: Agrostis tenuiflora Willd. 
= Muhlenbergia tenuiflora (Willd.) Britton, Sterns & 
Poggenb. 


224 


Catalogue of New World Grasses: II. 


Trichochloa virescens (Kunth) Roem. & Schult., Syst. Veg. 
2: 389. 1817. Basionym: Podosemum virescens Kunth. 
= Muhlenbergia virescens (Kunth) Trin. 


Trichodiclida prolifera Cerv., Naturaleza (Mexico City) 1: 
346. 1870. T-Specim.: 1 of 1. Mexico: in collibus de 
Guadelupe, et de Moctezuma prope Mexicum, 
Cervantes s.n. [T]. 

= Dasyochloa pulchella (Kunth) Willd. ex Rydb. 


Trichoneura Andersson, Kongl. Svenska 
Vetenskapsakad. Handl. 1853: 148. 1855. TYPE: 
Trichoneura hookeri Andersson; nom. illeg. superfl.; 
based on Calamagrostis pumila Hook.f., a legitimate 
nameCalamagrostis pumila Hook. f. 

TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 3 SUBSP.: 2. 
SYN.: Crossotropis Stapf. 


Trichoneura elegans Swallen, Amer. J. Bot. 19(5): 439, f. 4. 
1932. T-Protol.: USA: Texas: collected in sandy land 
south of Devine, 26 Sep 1931, W.A. Silveus 343 [HT: 
US-1501591]. T-Specim.: | of 1. USA: Texas, 26 Sep 
1931, W.A. Silveus 343 [HT: US-1501591; IT: US- 
1501592, US-1536745]. 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J.  Valdes-Reyna (2000); 
Hitchcock, A. $8. (1951). 


Trichoneura hookeri Andersson, Kongl. Vetensk. Acad. 
Handl. 1853: 148. 1855. T-Protol.: | Ecuador: 
Galapagos Islands, Albemarle Island. T-Specim.: | of 
1. Eduador: Galapagos Island, Albermarle Island, V./. 
Anderson [T: NY-19491]. 

NOTE: nom. illeg. superfl.; based on Calamagrostis pumila 
Hook.f., a legitimate name. 

CORRECT NAME: Calamagrostis pumila Hook. f. 

= Trichoneura lindleyana var. lindleyana. 


Trichoneura lindleyana (Kunth) Ekman, Ark. Bot. 11(9): 9. 
1912. Basionym: Leptochloa lindleyana Kunth. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Jorgensen, P. M. & S. Leén-Yanez 
(eds.) (1999); Lawesson, J. E., H. Adsersen & P. 
Bentley (1987); Porter, D. M. (1983); Valencia, R., 
N. Pitman, S. Leén-Yanez & P. M. Jorgensen (2000); 
Wiggins, I. L. & D. M. Porter (1971). 


Trichoneura lindleyana var. albemarlensis (B.L. Rob. & 
Greenm.) Reeder & C. Reeder, Madrofio 20: 253. 
1970. Basionym: Leptochloa albemarlensis B.L. Rob. 
& Greenm. 

DST.: Ecua. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Lawesson, J. E., H. Adsersen & P. 
Bentley (1987); Porter, D. M. (1983); Reeder, J. R. & 
C. G. Reeder (1971); Wiggins, I. L. & D. M. Porter 
(1971). 

SYN.: Leptochloa albemarlensis B.L. Rob, & Greenm. 


Trichoneura lindleyana (Kunth) Ekman var. lindleyana. 
DST.: Ecua. 
LVL. ACCPTC:: 1. 


ALSO ACCPTD. BY: Lawesson, J. E., H. Adsersen & P. 
Bentley (1987); Porter, D. M. (1983); Reeder, J. R. & 
C. G. Reeder (1971); Wiggins, I. L. & D. M. Porter 


(1971). 
SYN.: Calamagrostis pumila Hook. f.; Leptochloa hookeri 
(Andersson) Andersson; Leptochloa —lindleyana 


Kunth; Trichoneura hookeri Andersson. 


Trichoneura weberbaueri Pilg., Bot. Jahrb. Syst. 50(Beibl. 
111): 1. 1913. T-Protol.: Peru: ad viam ferream Limo- 
Oroya, in montibus septentrionem a statione Chosica 
versus, in declivibus saxosis aridis ad 1400 m, Apr 
1910, Weberbauer 5345 (HT: B]. T-Specim.: 1 of 1. 
Peru: Lima: Lima Province, Apr 1910, 4. Weberbauer 
5343 [HT: B; IT: US-2947334]. 

DST.: Chil, Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Macbride, J. F. (1936); Marticorena, 
C. & M. Quezada (1985); Pilger, R. (1913); Tovar, 
0. (1993). 


Tricuspis P. Beauv., Ess. Agrostogr. 77, f. 15/10. 1812. 
NOTE: hom. illeg. 
= Tridens Roem. & Schult. 


Tricuspis acuminata Munro ex A. Gray, Proc. Acad. Nat. Sci. 
Philadelphia 14: 335. 1862. T-Protol.: Western Texas, 
Wright 781 [HT: GH; IT: US]. T-Specim.: | of 1. C. 
Wright 78] (HT: HT: GH; IT: US-908404]. 

NOTE: nom. nud. 
= Erioneuron pilosum (Buckley) Nash. 


Tricuspis albescens Munro ex A, Gray, Proc. Acad. Nat. Sci. 
Philadelphia 14: 335. 1863. 
NOTE: nom. nud. 
= Tridens congestus (L.H. Dewey) Nash. 


Tricuspis ambigua (Elliott) Chapm., Fl. South. U.S. 559, 
1860. Basionym: Poa ambigua Elliott. 
= Tridens ambiguus (Elliott) Schult. 


Tricuspis avenaceus (Kunth) Thurb. ex A. Gray, Proc. Acad. 
Nat. Sci. Philadelphia 14: 335. 1863. Basionym: 
Triodia avenacea Kunth. 

NOTE: as Tricuspis avenacea. 
= Erioneuron avenaceum (Kunth) Tateoka. 


Tricuspis caroliniana P. Beauv., Ess. Agrostogr. 179, t. 3, f. 
29, t. 15, f. 10. 1812. T-Protol.: South Carolina. 
= Tridens flavus (L.) Hitche. 


Tricuspis congesta (L.H. Dewey) A. Heller, Cat. N. Amer. PI. 
(ed. 2) 28. 1900. Basionym: Sieglingia congesta L.H. 
Dewey. 

= Tridens congestus (L.H. Dewey) Nash. 


Tricuspis cornuta (Elliott) A. Gray, Manual 590. 1848. 
Basionym: Uralepis cornuta Elliott. 
= Triplasis americana P. Beauv. 


Tricuspis elongata (Buckley) A. Heller, Cat. N. Amer. PI. 


(ed. 2) 28. 1900. Basionym: Uralepis elongata 
Buckley. 
NOTE: basionym erroneously cited as "Triodia" 


elongataBuckley. 
= Tridens muticus var. elongatus (Buckley) Shinners. 


Tricuspis flava (L.) F.Y. Hubb., Rhodora 14(164): 186. 1912. 
Basionym: Poa flava L. 
= Tridens flavus (L.) Hitche. 


Catalogue of New World Grasses: II. 


225 


Tricuspis langloisii Nash, Bull. New York Bot. Gard. 1: 293. 
1899. T-Specim.: 1 of 1. USA: Louisana: Slidell: In 
pinewood clearings, 5 Oct 1891, A.B. Langlois s.n. 
[IT: US (fragm. ex NY)]. 

= Tridens ambiguus (Elliott) Schult. 


Tricuspis latifolia Griseb., Abh. Konigl. Ges. Wiss. 
Gottingen 19: 259, 1874. T-Specim.: | of 2. Argentina: 
Cordoba: pr. Ascochinga, Feb 1873, Lorentz & 
Hieronymus 256 |LT: GOET(fragm., US)], LT 
designated by Swallen, Amer. J. Bot. 22: 39 (1935). 2 
of 2. Argentina, Lorentz & Hieronymus 256 [LT: 
Grisebach Herb. ]. 

= Gouinia latifolia (Griseb.) Vasey. 


Tricuspis mutica Torr., Pacif. Railr. Rep. 4(5): 156. 1857. T- 
Specim.: | of 1. USA: New Mexico: Fort Smith to the 
Rio Grande, Laguna Colorado, 1853-1854, JM. 
Bigelow s.n. (IT: US (fragm. ex Torry herb., NY)]. 
= Tridens muticus (Torr.) Nash. 


Tricuspis nealleyi (Vasey) A. Heller, Cat. N. Amer. PI. (ed. 
2) 28. 1900. Basionym: Triodia nealleyi Vasey. 
= Erioneuron nealleyi (Vasey) Tateoka. 


Tricuspis novae-boracensis P. Beauv., Ess. Agrostogr. 77, 
179. 1812. 
NOTE: nom. illeg. 
= Tridens flavus (L.) Hitchc. 


Tricuspis pilosa (Buckley) A. Heller, Cat. N. Amer. PI. (ed. 
2) 2: 28. 1900. Basionym: Uralepis pilosa Buckley. 
= EFrioneuron pilosum (Buckley) Nash. 


Tricuspis pulchella (Kunth) Torr., Pacif. Railr. Rep. 4: 156. 
1857. Basionym: Triodia pulchella Kunth. 
NOTE: basionym erroneously cited as 
pulchella Kunth. 
= Dasyochloa pulchella (Kunth) Willd. ex Rydb. 


"Trichodia" 


Tricuspis purpurea (Walter) A. Gray, Manual 589. 1848. 
Basionym: Aira purpurea Walter. 
= Triplasis purpurea (Walter) Chapm. 


Tricuspis quinquefida P. Beauv. ex Don, Hort. Brit. 31. 1830. 
NOTE: based on Poa caerulescens Michx., nom. nud. 
= Tridens flavus (L.) Hitche. 


Tricuspis sesleroides Torr., Fl. N. Amer. 118. 1823. 
NOTE: nom. illeg. superfl., based on Poa seslerioides 
Michx. 
CORRECT NAME: Poa quinquefida Pursh. 
= Tridens flavus (L.) Hitche. 


Tricuspis sesleroides var. flexuosa Alph. Wood, Amer. Bot. 
Fl. 2: 398. 1871. T-Protol.: Pa. 
= Tridens flavus (L.) Hitche. 


Tricuspis sesleroides var. pallida T. Holm, Proc. Biol. Soc. 
Wash. 14: 19. 1901. T-Protol.: With the type near 
Marshall Hall, Ho/m s.n. (HT: private herb. T. Holm]. 
T-Specim.: | of 1. USA: Maryland: field and roadsides 
at Marshall Hall, 28 Sept 1898, Holm s.n. [IT: US- 
908342]. 

= Tridens flavus (L.) Hitche. 


Tricuspis simplex Griseb., Mem. Amer. Acad. Arts, n.s., 8: 
532. 1862. T-Protol.: Cuba: in rupidus aridis, Wright 
1551. T-Specim.: | of 1. CUBA, 1860,C. Wright 1551 
[IT: MO-2108840, MO-2108841, US-3048158]. 


= Tripogon spicatus (Nees) Ekman. 


Tricuspis stricta (Nutt.) Alph. Wood, Class-book Bot. (ed. 
1861) 792. 1861. Basionym: Windsoria stricta Nutt. 
= Tridens strictus (Nutt.) Nash. 


Tridens Roem. & Schult. Syst. Veg. 2: 34, 599. 1817. 

REPLACED NAME: Tricuspis P. Beauv. 

BLOCKING NAME: Tricuspis Pers. 

TRTD. BY: Valdés-Reyna, J., & P. M. Peterson. 

NHT: Cynodonteae. NO. ACCPTD. SPP.: 15, SUBSP.: 6. 

CNSLTD. LIT.: Tateoka, T., (1961); Sanchez, E., (1979). 

SYN.: Antonella Caro; Gossweilerochloa Renvoize; 
Rhombolytrum Link; Tricuspis P. Beauy.; Windsoria 
Nutt. 


Tridens albescens (Vasey) Wooton & Standl., New Mexico 
Agric. Exp. Sta. Bull. 81: 129. 1912. Basionym: 
Triodia albescens Vasey. 

DST.: Mexi, USA. 

LVL. ACCPTC:: I. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Great Plains Flora Association (1986); 
Hitchcock, A. S. (1951). 

Rhombolytrum albescens (Vasey) Nash; Sieglingia 
albescens (Vasey) Kuntze; Tridens  albescens 
(Vasey) N.T. Burb.; 7riodia albescens Vasey. 


SYN:: 


Tridens albescens (Vasey) N.T. Burb., Blumea 3: 84. 1946. 
Basionym: Triodia albescens Vasey. 
NOTE: isonym; in obs. 
= Tridens albescens (Vasey) Wooton & Standl. 


Tridens ambiguus (Elliott) Schult., Mant. 2: 333. 1824. 
Basionym: Poa ambigua Elliott. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Godfrey, R. K. & J. W. 
Wooten (1979); Hitchcock, A. S. (1951); Radford, A. 
E., H. E. Ahles & C. R. Bell (1968); Wunderlin, R. P. 


(1998). 
SYN.: Poa ambigua Elliott; Sieglingia ambigua (Elliott) 
Kuntze; Tricuspis ambigua (Elliott) Chapm.;: 


Tricuspis langloisii Nash; Tridens langoisii (Nash) 
Nash; Triodia ambigua (Elliott) Benth. ex Vasey; 
Triodia elliottii Bush; Triodia langoisii (Nash) Bush, 
Uralepis ambigua (Elliott) Kunth; Windsoria 
ambigua (Elliott) Nutt. 


Tridens avenaceus (Kunth) Hitche., Contr. U.S. Natl. Herb. 
17(3): 357. 1913. Basionym: 7riodia avenacea Kunth. 
= Erioneuron avenaceum (Kunth) Tateoka. 


Tridens avenaceus (Kunth) Hitche. var. avenaceus. 
= Erioneuron avenaceum var. avenaceum. 


Tridens avenaceus var. kurtziana Parodi, Revista Argent. 
Agron. 4(4): 256. 1937. T-Protol.: Argentina: San 
Juan: precordillera entre Barreales, tontal y Retamito 
(Maradona), 1740 m, 28 Nov 1897, F. Kurtz 9850 
[HT: Herb. Fac. Med. B. Aires]. T-Specim.: | of 1. 
Argentina: San Juan: Dpto. Zonda: Precordillera entre 
Barreales, Tontal y Retamito, entre Agua Pinta y 
Pedernal, cerca de Maradona, 1740 m, 28 Feb 1897, 
Kurtz 9850 [HT: BAF; IT: CORD, LP, US-908624]. 


226 


Catalogue of New World Grasses: II. 


NOTE: as Tridens "avenacea" var. "Kurtziana". 
= Erioneuron avenaceum var. kurtzianum 
Anton. 


(Parodi) 


Tridens avenaceus var. longiglumis Parodi, Revista Argent. 
Agron. 4(4): 255, f. 4. 1937. T-Protol.: Argentina: 
Catamarca: Andalgala, Mar 1916, Jorgensen 1718 
[BAF]. T-Specim.: | of |. Argentina: Catamarca: 
Depto. Andalgala: Las Minas, Mar 1916, Jorgensen 
1718 [HT: BAF; IT: BAF, GH, LIL, MO, SI, US- 
917965). 

NOTE: as Tridens "avenacea" var. longiglumis. 

= Erioneuron avenaceum var. longiglume (Parodi) Anton. 


Tridens avenaceus var. pygmaea (Hack.) Parodi, Revista 
Argent. Agron. 4(4): 253, f. 3. 1937. Basionym: 
Triodia avenacea var. pygmaea Hack. 

NOTE: as Tridens "avenacea" var. pygmaea. 
= Erioneuron avenaceum var. pygmaeum (Hack.) Anton. 


Tridens brasiliensis (Nees ex Steud.) Parodi, Revista 
Argent. Agron. 4(4): 245. 1937. Basionym: Uralepis 
brasiliensis Nees ex Steud. 

NOTE: treated by Parodi as 7ridens brasiliensis Nees, but 
to be treated as a new combintion since Parodi was 
the first to use and accept this combination and at the 
same same time made a clear and direct reference to 
the basionym by including it in his synonymy. 

DST.: Arge, Braz, Para, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 

(1987); Anton, A. M. (1977); Burkart, A, (1969); 

Cabrera, A. L. (1970); Doll, J. C. (1878); Filgueiras, 

T. S. (1991); Rosengurtt, B., B. R. Arrillaga de 

Maffei & P. Izaguirre de Artucio (1970); Zuloaga, F. 

O., E. G. Nicora, Z. E. Raigolo de Agrasar, O. 

Morrone, J. Pensiero & A. M. Cialdella (1994). 

Tridens brasiliensis Nees ex Steud. ex Jacks.; 

Triodia brasiliensis (Nees ex Steud.) Lindm.; Triodia 

figueirae Arechav.; Uralepis brasiliensis Nees ex 

Steud. 


SYN.: 


Tridens brasiliensis Nees ex Steud. ex Jacks., Index Kew. 2: 
ITLL. 1895, 
NOTE: nom. inval., as syn. of Uralepsis brasiliensis Nees; 
Nees unpublished. 
= Tridens brasiliensis (Nees ex Steud.) Parodi. 


Tridens buckleyanus (Vasey ex L.H. Dewey) Nash, FI. S.E. 
U.S. 143. 1903. Basionym: Sieglingia buckleyana 
Vasey ex L.H. Dewey. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Hitchcock, A. S. (1951). 

SYN.: Sieglingia buckleyana Vasey ex L.H. Dewey; 
Triodia buckleyana (Vasey ex L.H. Dewey) Vasey ex 
Hitche. 


Tridens carolinianus (Steud.) Henrard, Blumea 3(3): 424. 
1940. Basionym: Festuca caroliniana Steud. 

DST.: USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Hitchcock, A. S. 
(1951); Radford, A. E., H. E. Ahles & C. R. Bell 
(1968); Wunderlin, R. P. (1998). 

SYN.: Festuca caroliniana Steud.; Tridens drummondii 
(Scribn. & Kearney) Nash ex Small; Triodia 


caroliniana (Steud.) Chase; 7riodia drummondii 
Scribn. & Kearney. 

Tridens chapmanii (Small) Chase, Man. Grasses U.S. (ed. 2) 
213: 971. 1951. Basionym: Sieglingia chapmanii 
Small. 

= Tridens flavus var. chapmanii (Small) Shinners. 


Tridens congestus (L.H. Dewey) Nash, Fl. S.E. U.S. 143. 
1903. Basionym: Sieglingia congesta L.H. Dewey. 

DST.: USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Hitchcock, A. S. (1951). 

SYN.: Sieglingia congesta L.H. Dewey; Tricuspis albescens 
Munro ex A. Gray; Tricuspis congesta (L.H. Dewey) 
A. Heller; Triodia congesta (L.H. Dewey) Bush. 


Tridens drummondii (Scribn. & Kearney) Nash ex Small, FI. 
S.E. U.S. 143. 1903. Basionym: Triodia drummondii 
Scribn. & Kearney. 

= Tridens carolinianus (Steud.) Henrard. 


Tridens duartei Cat. Guerra, Acta Bot. Cub. 4: 3. 1980. T- 
Protol.: Cuba: Prov. de Camaguey: Santa Lucia, Fundo 
del Jucaro; Apr. 1973, Catasus 116 [HT: HAC]. 

= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Tridens elongatus (Buckley) Nash, Fl. S.E. U.S. 143. 1903. 
Basionym: Uralepis elongata Buckley. 
= Tridens muticus var. elongatus (Buckley) Shinners. 


Tridens eragrostoides (Vasey & Scribn.) Nash, Fl. S.E. U.S. 
142. 1903. Basionym: Triodia eragrostoides Vasey & 
Scribn., 

DST.: Cari, Mexi, USA, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Catasus Guerra, L. (1997); Correll, D. S. & 
M. C. Johnston (1970); Davidse, G. (1994): Davidse, 
G., M. Sousa S. & A. O. Chater (editores generales) 
(1994); Espejo Serna, A., A. R. Lopez-Ferrari, and J. 
Valdes-Reyna (2000); Hitchcock, A. S. (1951); Long, 
R. W. & O. Lakela (1971); Wunderlin, R. P. (1998). 

Sieglingia eragrostoides (Vasey & Scribn.) L.H. 
Dewey; Sieglingia eragrostoides var. scabra Vasey 
ex Beal; Triodia eragrostoides var.scabra (Vasey ex 
Beal) Bush; 7riodia eragrostoides Vasey & Scribn. 


SYN: 


Tridens flaccidus (D6Il) Parodi, Revista Argent. Agron. 
4(4): 249. 1937. Basionym: Uralepis flaccida Doll. 
DST.: Braz, Guya, Vene. 
LVL. ACCPTC:: 1. 
ALSO ACCPTD. BY: Judziewicz, E. J. (1990). 
SYN.: Triodia flaccida (DO6ll) Hitche.; Uralepis flaccida 
Doll. 


Tridens flavus (L.) Hitche., Rhodora 8(95): 210. 1906. 
Basionym: Poa flava L. 

NOTE: as "flava". 

DST.: Mexi, USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Darke, R. (1999); Espejo Serna, A.. A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Gleason, H. A. 
& A. Cronquist (1991); Great Plains Flora 
Association (1986); Hitchcock, A. S. (1951); Long, 
R. W. & O. Lakela (1971); Peterson, P. M., and S.C. 


Catalogue of New World Grasses: II. 


227 


Boechat (2001); Radford, A. E., H. E. Ahles & C. R. 
Bell (1968); Voss, E. G. (1972); Wunderlin, R. P. 


(1998). 
SYN.:  Cynodon carolinianus (P. Beauv.) Raspail; 
Eragrostis arundinacea  Jedwabn.; Eragrostis 


tricuspis Trin.; Festuca flava (L.) F. Muell.; Festuca 
purpurea Schreb. ex Steud.; Festuca quadridens 
Poir.; Panicum festucoides Poir.; Poa arundinacea 
Poir.; Poa caerulescens Michx. ex P. Beauv.; Poa 
flava L.; Poa quinquefida Pursh; Poa sesleroides 
Michx.; Sieglingia cuprea (Jacq.) Millsp.; Sieglingia 
flava (L.) Kuntze; Sieglingia sesleroides (Michx.) 
Scribn.; Sieglingia sesleroides var. intermedia Vasey 
ex L.H. Dewey; Tricuspis caroliniana P. Beauv.; 
Tricuspis flava (L.) F.T. Hubb.; Tricuspis novae- 
boracensis P. Beauv.; Tricuspis quinquefida P. 
Beauv. ex Don; Tricuspis sesleroides Torr.; Tricuspis 
sesleroides var. flexuosa Alph. Wood; Tricuspis 
sesleroides var. pallida T. Holm; Tridens flavus fo. 
cuprea (Jacq.) Fosberg; Tridens flavus var. aristatus 
(Scribn. & C.R. Ball) Shinners; Tridens quinquefidus 
(Pursh) Roem. & Schult.; Tridens sesleroides Nash; 
Triodia caerulescens (Michx.) Desv.; Triodia cuprea 
Jacq.; Triodia festucoides (Poir.) Desv.; Triodia flava 
(L.) Hitche.; Triodia flava (L.) Smyth; Triodia flava 
fo. cuprea (Jacq.) Fosberg; Triodia flava fo. flava; 
Triodia flava var. aristata (Scribn. & C.R. Ball) 
Fernald & Griscom; Triodia novaeboracensis (P. 
Beauv.) Desv.; Triodia sesleroides var. aristata 
Scribn. & C.R. Ball; Triodia triticoides C.A. 
Gardner; Uralepis cuprea (Jacq.) Kunth; Uralepis 
tricuspis (Trin.) Steud.; Windsoria poiformis Nutt.; 
Windsoria sesleroides (Michx.) Eaton. 


Tridens flavus var. aristatus (Scribn. & C.R. Ball) Shinners, 
Rhodora 56(662): 27. 1954. Basionym: Triodia 
sesleroides var. aristata Scribn. & C.R. Ball. 

= Tridens flavus (L.) Hitche. 


Tridens flavus var. chapmanii (Small) Shinners, Rhodora 
56(662): 27. 1954. Basionym: Sieglingia chapmanii 
Small. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Gleason, H. A. & A. 
Cronquist (1991); Radford, A. E., H. E. Ahles & C. 
R. Bell (1968); Wunderlin, R. P. (1998). 

Sieglingia chapmanii Small; Tridens chapmanii 
(Small) Chase; Triodia chapmanii (Small) Bush, 
Triodia flava var. chapmanii (Small) Fernald & 
Griscom. 


SYN:: 


Tridens flavus fo. cuprea (Jacq.) Fosberg, Castanea 20: 58. 
1957. Basionym: Triodia cuprea Jacq. 
= Tridens flavus (L.) Hitche. 


Tridens flavus (L.) Hitche. var. flavus. 
DST.: Mexi, USA. 
LVL. ACCPTC.: L. 
ALSO ACCPTD. BY: Gleason, H. A. & A. Cronquist 
(1991); Radford, A. E., H. E. Ahles & C. R. Bell 
(1968); Wunderlin, R. P. (1998). 


Tridens grandiflora (Vasey) Wooton & Standl., New Mexico 
Agric. Exp. Sta. Bull. 81: 129. 1912. Basionym: 
Triodia grandiflora Vasey. 

= Erioneuron avenaceum (Kunth) Tateoka. 


Tridens hackelii (Arechav.) Parodi, Revista Argent. Agron. 
4(4): 247, f. 1. 1937. Basionym: Triodia hackelii 
Arechav. 

NOTE: as Tridens "Hackeli". 

DST.: Arge, Braz, Para, Urug. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anton, A. M. (1977); Burkart, A. 
(1969); Rosengurtt, B., B. R. Arrillaga de Maffei & 
P. Izaguirre de Artucio (1970); Zuloaga, F. O., E. G. 
Nicora, Z. E. Rugolo de Agrasar, O. Morrone, J. 
Pensiero & A. M. Cialdella (1994). 

SYN.: Triodia hackelii Arechav. 


Tridens langoisii (Nash) Nash, Fl. S.E. U.S. 142. 1903. 
Basionym: Tricuspis langloisii Nash. 
= Tridens ambiguus (Elliott) Schult. 


Tridens muticus (Torr.) Nash, Fl. S.E. U.S. 143. 1903. 
Basionym: Tricuspis mutica Torr. 

DST.: Mexi, USA. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Cronquist, A., A. H. Holmgren, N. H. Holmgren & J. 
L. Reveal (1977); Espejo Serna, A., A. R. Lopez- 
Ferrari, and J. Valdes-Reyna (2000); Gould, F. W. & 
R. Moran (1981); Great Plains Flora Association 
(1986); Hickman, J. C. (1993); Hitchcock, A. S. 
(1951); Munz, P. A. & D. D. Keck (1959); Munz, P. 
A. (1974): Shreve, F. & I. L. Wiggins (1964). 


SYN.: Sieglingia mutica (Torr.) Kuntze; Tricuspis mutica 
Torr.; Triodia mutica (Torr.) Scribn.; Uralepis 
mutica E. Fourn. ex Hemsl.; Uralepis pilosa 
Buckley. 


Tridens muticus fo. effusus I.M. Johnst., SouthW. Naturalist 
14: 258. 1969. T-Specim.: | of 1. USA: Texas, 19 Aug 
1952, B.H. Warnock 10857 [HT: LL]. 
DST.: USA. 
LVL. ACCPTC.: 1. 


Tridens muticus var. elongatus (Buckley) Shinners, 
Rhodora 56(662): 28. 1954. Basionym: Uralepis 
elongata Buckley. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Great Plains 
Flora Association (1986). 

SYN.: Sieglingia elongata (Buckley) Nash; Tricuspis 
elongata (Buckley) A. Heller; Tridens elongatus 
(Buckley) Nash; 7riodia elongata (Buckley) Scribn.; 
Uralepis elongata Buckley. 


Tridens muticus (Torr.) Nash fo. muticus. 
= Tridens muticus var. muticus. 


Tridens muticus (Torr.) Nash fo. muticus. 
LVL. ACCPTC.: |. 


Tridens muticus (Torr.) Nash var. muticus. 
DST.: USA. 
LVL. ACCPTC.: 1. 
SYN.: Tridens muticus fo. muticus. 


Tridens nealleyi (Vasey) Wooton & Standl., New Mexico 
Agric. Exp. Sta. Bull. 81: 129. 1912. Basionym: 
Triodia nealleyi Vasey. 

= Erioneuron nealleyi (Vasey) Tateoka. 


228 


Catalogue of New World Grasses: II. 


Tridens nicorae Anton, Kurtziana 10: 51, f. 1. 1977. T- 
Protol.: Argentina: San Luis: Dpto. Pringles: Campo 
E! Totoral, 3 km al N de Los Tapiales, 1500 m, 29 Mar 
1972, rara en pastizal serrano, suelo pedrogoso, 
Anderson 2428 [HT: CORD]. 
DST.: Arge. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Anton, A. M. (1977); Zuloaga, F. O., 
E. G. Nicora, Z. E. Rugolo de Agrasar, O. Morrone, 
J. Pensiero & A. M. Cialdella (1994). 
SYN.: Antonella nicorae (Anton) Caro. 


Tridens x oklahomensis (Feath.) Feath., U.S.D.A. Bur. PI. 
Industr. Misc. Publ. 213: 973. 1951. Basionym: 
Triodia oklahomensis Feath., 

NOTE: [Tridens flavus (L.) Hitche. x T. strictus (Nutt.) 
Nash]. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Hitchcock, A. S. 
(1951). 


Tridens pilosus (Buckley) Hitche., Contr, U.S, Natl. Herb. 
17(3): 357. 1913. Basionym: Uralepis pilosa Buckley. 
= Erioneuron pilosum (Buckley) Nash. 


Tridens pilosus var. argentina (Kuntze) Parodi, Revista 
Argent. Agron. 4(4): 251, f 2C-D. 1937. Basionym: 
Sieglingia argentina Kuntze. 

NOTE: as Tridens "pilosa" var. argentina. 
= Erioneuron pilosum var. longearistatum (Kurtz) Anton. 


Tridens pilosus var. mendocina Parodi, Revista Argent. 
Agron. 4(4): 251, f. 2A-B. 1937. T-Protol.: Argentina: 
Mendoza: Las Heras, Potrerillos, estancia El Salto, to, 
1700 m.,s.m. Nov 1933, 4. Ragonese 94. 

NOTE: as Tridens "pilosa" var. mendocina. 
= Erioneuron pilosum var. mendocinum (Parodi) Nicora. 


Tridens pulchellus (Kunth) Hitche., Fl. Calif. 1: 141. 1912. 
Basionym: Triodia pulchella Kunth. 
= Dasyochloa pulchella (Kunth) Willd. ex Rydb. 


Tridens quinquefidus (Pursh) Roem. & Schult., Syst. Veg. 2: 
599. 1817. Basionym: Poa quinquefida Pursh. 
= Tridens flavus (L.) Hitchce. 


Tridens sesleroides Nash, Fl. S.E. U.S. 142. 1903. 
REPLACED NAME: Poa sesleroides Michx. 
BLOCKING NAME: Poa sesleroides All. 
= Tridens flavus (L.) Hitchc. 


Tridens strictus (Nutt.) Nash, Fl. S.E. U.S. 143. 1903. 
Basionym: Windsoria stricta Nutt. 

DST.: USA. 

LVL. ACCPTC.:: |. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Godfrey, R. K. & J. W. 
Wooten (1979); Great Plains Flora Association 
(1986); Hitchcock, A. S. (1951); Radford, A. E., H. 
E. Ahles & C. R. Bell (1968); Wunderlin, R. P. 
(1998). 

Sieglingia stricta (Nutt.) Kuntze; Tricuspis stricta 
(Nutt.) Alph. Wood; Triodia stricta (Nutt.) Benth. ex 
Vasey; Uralepis densiflora Buckley; Windsoria 
stricta Nutt. 


Tridens texanus (S. Watson) Nash, FI. S.E. U.S. 142. 1903. 
Basionym: Triodia texana S$. Watson. 


SYN.: 


DST.: Mexi, USA. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Correll, D. S. & M. C. Johnston (1970); 
Espejo Serna, A., A. R. Lopez-Ferrari, and J. Valdes- 
Reyna (2000); Hitchcock, A. S. (1951). 

SYN.: Sieglingia texana (S. Watson) Kuntze; Triodia 
texana S. Watson. 


Tridens veralensis Cat. Guerra, Acta Bot. Cub. 4: 4. 1980. T- 
Protol.: Cuba: Prov. de Pinar del Rio: Guanahacabibes, 
El Veral; 20 Apr. 1974, Catasus 298 [HT: HAC]. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Tridens verticillatus Meyen, Reise Erde 1: 408, 1834. T- 
Protol.: Chile: prov. Copiapo: Nantoco, /Meyen] s.n. 
NOTE: nom. nud. 
= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Tridens virens Nees, Fl. Bras. Enum. Pl. 2: 476. 1829. T- 
Protol.: Brazil: Habitat in graminosis ad fluvim S. 
Francisci in provincia Bahiensi et Permambucana, ad 
Joazeiro et alibi.". 

= Leptochloa fusea subsp. fascicularis (Lam.) N. Snow. 


Triodia acuminata Benth. ex Vasey, U.S.D.A. Div. Bot. Bull. 
12(2): 32. 1891. 
NOTE: cites Tricuspis acuminata Munro in herb, 
= Erioneuron pilosum (Buckley) Nash. 


Triodia albescens Vasey, U.S.D.A. Div. Bot. Bull. 12(2): t. 
33. 1891. T-Specim.: | of 1. USA: Texas, 1872, E. 
Hall 782 [IT: US-908505]. 
= Tridens albescens (Vasey) Wooton & Stand. 


Triodia ambigua (Elliott) Benth. ex Vasey, Dept. Agric. 
Special Rep. 63: 35. 1883. Basionym: Poa ambigua 
Elliott. 

NOTE: hom. illeg. 
E. NAME: Triodia ambigua R. Br. 
= Tridens ambiguus (Elliott) Schult. 


Triodia argentina (Kuntze) K. Schum., Just's Bot. Jahresber. 
26(1): 331. 1900. Basionym: Sieglingia argentina 
Kuntze. 

= Erioneuron pilosum var. longearistatum (Kurtz) Anton. 


Triodia avenacea Kunth, Nov. Gen. Sp. 1: 156, t. 48. 1815 
[1816]. T-Protol.: In covalle Mexicana inter montem 
Chapultepec et Penol de los Banos, alt. 1190 hexap, 
Humboldt & Bonpland s.n. T-Specim.: 1 of 1. Mexico: 
Distrito Federal: Chapultepec, Humboldt & Bonpland 
gn. [ST: P]. 

= Erioneuron avenaceum (Kunth) Tateoka. 


Triodia avenacea Tateoka var. avenacea. 
= Erioneuron avenaceum var. avenaceum. 


Triodia avenacea var. breviaristata Kurtz, Revista Mus. La 
Plata 5: 301. 1893. T-Protol.: Valle de Mexico, 
Schaffner 158. T-Specim.: | of 1. Mexico: Valle de 
Mexico, Schaffner 158 [HT: US-825506]. 

= Erioneuron avenaceum (Kunth) Tateoka. 


Triodia avenacea var. longearistata Kurtz, Revista Mus. La 
Plata 5: 299. 1893. T-Specim.: 1 of 1. Argentina: 
colinas secas cerca de la Estancia La Era, al SO de San 
Francisco del Chanar, Kurtz 6729 [LT: CORD], LT 
designated by Anton, Kurtziana 10: 58 (1977). 

= Erioneuron pilosum var. longearistatum (Kurtz) Anton. 


Catalogue of New World Grasses: II. 


229 


Triodia avenacea var. pygmaea Hack., Nova Acta Regiae 
Soc. Sci. Upsal. 1(1): 179. 1905. T-Specim.: | of 1. 
[HT: W; IT: CORD]. 
= Erioneuron avenaceum var. pygmaeum (Hack.) Anton. 


Triodia brasiliensis (Nees ex Steud.) Lindm., Kong]. Svenska 
Vetenskapsakad. Handl. 34(6): 35. 1900. Basionym: 
Uralepsis brasiliensis Nees ex Steud. 

= Tridens brasiliensis (Nees ex Steud.) Parodi. 


Triodia buckleyana (Vasey ex L.H. Dewey) Vasey ex 
Hitche., J. Wash. Acad. Sci. 23(10): 452. 1933. 
Basionym: Sieglingia buckleyana Vasey ex L.H. 
Dewey. 

= Tridens buckleyanus (Vasey ex L.H. Dewey) Nash. 


Triodia caerulescens (Michx.) Desv., Mem. Soc. Agric. 
Angers 1: 203. 1831. Basionym: Poa_sesleroides 
Michx. 

NOTE: also in Opusc. Sci. Phys. Nat. 99 [reprint] 1831- 

1833. 
= Tridens flavus (L.) Hitche. 


Triodia caroliniana (Steud.) Chase, Amer. J. Bot. 24(1): 34. 
1937. Basionym: Festuca caroliniana Steud. 
= Tridens carolinianus (Steud.) Henrard. 


Triodia chapmanii (Small) Bush, Trans. Acad. Sci. St. Louis 
12(6): 74. 1902. Basionym: Sieglingia chapmanii 
Small. 

NOTE: as Triodia "Chapmani". 
= Tridens flavus var. chapmanii (Small) Shinners. 


Triodia congesta (L.H. Dewey) Bush, Trans. Acad. Sci. St. 
Louis 12(6): 67. 1902. Basionym: Sieglingia congesta 
L.H. Dewey. 
= Tridens congestus (L.H. Dewey) Nash. 


Triodia cuprea Jacq., Ecl. Gram. Rar. 2: 21. 1812, T-Protol.: 
Vaterland. unbekandt. kam 1803 durch Samen aus 
England. 

= Tridens flavus (L.) Hitche. 


Triodia drummondii Scribn. & Kearney, Bull. Div. Agrostol., 
U.S.D.A. 4: 37. 1897. T-Specim.: 1 of 4. USA: 
Florida: Duval Co., 1833, 7. Drwmmond s.n. [ST: US- 
81810]. 2 of 4. USA: South Carolina, Ravenel s.n. [ST: 
US]. 3 of 4. USA: Mississippi: Biloxi, growing in dry 
soil in low pine barrens, 1896, Kearney 324 [ST]. 4 of 
4. USA: Georgia, Anon. [ST: US]. 

= Tridens carolinianus (Steud.) Henrard. 


Triodia elliottii Bush, Trans. Acad. Sci. St. Louis 12(6): 73. 
1902. 
NOTE: nom. illeg. superfl. — as "Elliottii". 
CORRECT NAME: Poa ambigua Elliott. 
= Tridens ambiguus (Elliott) Schult. 


Triodia elongata (Buckley) Scribn., Bull. Div. Agrostol., 
U.S.D.A. 17(ed.2): 210, f. 506. 1901. Basionym: 
Uralepis elongata Buckley. 

= Tridens muticus var. elongatus (Buckley) Shinners. 


Triodia eragrostoides Vasey & Scribn., Contr. U.S. Natl. 
Herb. 1(2): 58. 1890. T-Specim.: 1 of 2. USA: Texas, 
1889, G.C. Nealley 822 [IT: US-81811]. 2 of 2. USA: 
Texas, Nealley s.n. [HT: US]. 
= Tridens eragrostoides (Vasey & Scribn.) Nash. 


Triodia eragrostoides var. scabra (Vasey ex Beal) Bush, 
Trans. Acad. Sci. St. Louis 12(6): 71. 1902. Basionym: 
Sieglingia eragrostoidesvar. scabra Vasey ex Beal. 

= Tridens eragrostoides (Vasey & Scribn.) Nash. 


Triodia festucoides (Poir.) Desv., Mem. Soc. Agric. Angers 
1: 202. 1831. Basionym: Panicum festucoides Poir. 
NOTE: also in Opusc. Sci. Phys. Nat. 98 [reprint] 1831- 
1833. 
= Tridens flavus (L.) Hitche. 


Triodia figueirae Arechay., Anales Mus. Nac. Montevideo 1: 
408, t. 49. 1896. T-Specim.: 1 of 2. Uruguay: en 
campos de Concordia villa occidental del Uruguay, 
Figueira s.n. [ST: MVM]. 2 of 2. Uruguay: Mercedes, 
1895, D.M.B. Berro s.n.[ST: MVM]. 

NOTE: as Triodia "Figueirae" 
= Tridens brasiliensis (Nees ex Steud.) Parodi. 


Triodia flaccida (Doll) Hitche., Contr. U.S. Natl. Herb. 22(6): 
457. 1922. Basionym: Uralepis flaccida DOll. 
= Tridens flaccidus (Doll) Parodi. 


Triodia flava (L.) Smyth, Trans. Kansas Acad. Sci. 25: 95. 
1913. Basionym: Poa flava L. 
= Tridens flavus (L.) Hitche. 


Triodia flava (L.) Hitche., U.S.D.A. Bull. (1915-23) 772: 76, 
f. 35. 1920. Basionym: Poa flava L. 
NOTE: isonym. 
E. NAME: Triodia flava (L.) Smyth. 
= Tridens flavus (L.) Hitchce. 


Triodia flava var. aristata (Scribn. & C.R. Ball) Fernald & 
Griscom, Rhodora 37(436): 134. 1935. Basionym: 
Triodia sesleroides var. aristata Scribn. & C.R. Ball. 

= Tridens flavus (L.) Hitche. 


Triodia flava var. chapmanii (Small) Fernald & Griscom, 
Rhodora 37(436): 133. 1935. Basionym: Sieglingia 
chapmanii Small. 

NOTE: as "Chapmani". 
= Tridens flavus var. chapmanii (Small) Shinners. 


Triodia flava fo. cuprea (Jacq.) Fosberg, Castanea 11: 67. 
1946. Basionym: Triedia cuprea Jacq. 
= Tridens flavus (L.) Hitche. 


Triodia flava (L.) Smyth fo. flava. 
= Tridens flavus (L.) Hitchce. 


Triodia grandiflora Vasey, Contr. U.S. Natl. Herb. 1(2): 59. 
1890. T-Specim.: | of 1. USA: Texas, Presidio Co.: 
Chenate Mountains, no date, G.C. Nealley 823 [IT: 
US-81812]. 

= Erioneuron avenaceum (Kunth) Tateoka. 


Triodia hackelii Arechay., Anales Mus. Nac. Montevideo 1: 
410. 1896. T-Protol.: "Recogida por el Sr. Berro, D. 
Mariano B., en campos graminosos de Vera. T- 
Specim.: 1 of 1. Uruguay: Depart. Vera: Mercedes, 
MB. Berro s.n. [HT: MVM; IT:US-908629]. 

NOTE: as Triodia "Hackeli". 
= Tridens hackelii (Arechav.) Parodi. 


Triodia langoisii (Nash) Bush, Trans. Acad. Sci. St. Louis 
12(6): 72. 1902. Basionym: Tricuspis langloisii Nash. 
NOTE: as "Langloisii". 
= Tridens ambiguus (Elliott) Schult. 


230 


Catalogue of New World Grasses: II. 


Triodia multinervosa (Vasey) Hitchc., Proc. Biol. Soc. Wash. 
41: 159. 1928. Basionym: Melica multinervosa Vasey. 
= Vaseyochloa multinervosa (Vasey) Hitchce. 


Triodia mutica (Torr.) Scribn., Bull. Torrey Bot. Club 10(1): 
30. 1883 [Mar]. Basionym: Tricuspis mutica Torr. 
NOTE: isonym by Benth. in Aug 1883. 
= Tridens muticus (Torr.) Nash. 


Triodia nealleyi Vasey, Bull. Torrey Bot. Club 15: 49. 1890. 
T-Specim.: | of 1. USA: Texas, 1887,G.C. Nealley 77 
{IT: US-908406], CS. 
NOTE: nom. nud. 
= Erioneuron nealleyi (Vasey) Tateoka. 


Triodia nealleyi Vasey, U.S.D.A. Div. Bot. Bull. 12(2): t. 36. 
1891. T-Specim.: | of 1. USA: Texas: Pesidio Co.: 
Cibolo Canyon, Chenate Mountains, 1887,Neally s.n. 
[I'T: US-908406]. 

= Erioneuron nealleyi (Vasey) Tateoka. 


Triodia novaeboracensis (P,. Beauv.) Desv., Mem. Soc. 
Agric. Angers 1: 203. 1831. Basionym: Tricuspis 
novae-boracensis P. Beauv. 

NOTE: also in Opusc. Sci. Phys. Nat. 99 [reprint] 1831- 

1833. 
= Tridens flavus (L.) Hitche. 


Triodia paraguayensis (Kuntze) Hack., Anales Mus, Nac. 
Buenos Aires 11: 127. 1904. Basionym: Arundinaria 
paraguayensis Kuntze. 

= Gouinia paraguayensis (Kuntze) Parodi. 


Triodia_ pilosa (Buckley) Merr., Circ. Div. Agrostol. 
U.S.D.A. 32: 9. 1901. Basionym: Uralepis pilosa 
Buckley. 

= Erioneuron pilosum (Buckley) Nash. 


Triodia pulchella Kunth, Nov. Gen. Sp. 1: 155, t. 47. 1815 
[1816]. T-Protol.: In subfrigidis, siccis, apricis regnii 
Mexicani inter Guanaxuati, Mina de Belgrado et 
Cubilete, Humboldt & Bonpland s.n. T-Specim.: | of 
2. Mexico: Guanajuato, Humboldt & Bonpland s.n. 
[HT: P; IT: B-W-2046]. 2 of 2. Mexico, ex hb. 
Humboldt [T: 1T: US-91469 (fragm.)]. 

= Dasyochloa pulchella (Kunth) Willd. ex Rydb. 


Triodia purpurea (Walter) Smyth, Trans. Kansas Acad. Sci. 
25: 95. 1913. Basionym: Aira purpurea Walter. 
= Triplasis purpurea (Walter) Chapm. 


Triodia schaffneri S. Watson, Proc. Amer. Acad. Arts 18: 
181. 1883. T-Protol.: Mexico, Schaffner 1077. T- 
Specim.: | of 1. Mexico: San Louis Potosi, August 
1877, Schaffner 1077 [1T: US (fragm. ex GH), K]. 

= Tripogon spicatus (Nees) Ekman. 


Triodia sesleroides var. aristata Scribn. & C.R. Ball, Bull. 
Div. Agrostol., U.S.D.A. 24: 45. 1901. T-Protol.: 
USA: Florida: Orange Co.: collected at Clarcona, Oct? 
1899, Marie Meislahn 90 [HT: US]. T-Specim.: 1 of 1. 
USA: Florida: Orange Co.: Clarcona, 16 Oct. 1899, 
MarieMeislahn 90 [HT: US-1538641]. 
NOTE: as Triodia "seslerioides" var. aristata. 
= Tridens flavus (L.) Hitche. 


Triodia stricta (Nutt.) Benth. ex Vasey, Dept. Agric. Special 
Rep. 63: 35. 1883. Basionym: Windsoria stricta Nutt. 
= Tridens strictus (Nutt.) Nash. 


Triodia texana S. Watson, Proc. Amer. Acad. Arts 18: 180. 
1883. T-Specim.: | of 1. USA: Texas, 1851-1852.C. 
Wright 2055 [IT: US-908320]. 

= Tridens texanus (S. Watson) Nash. 


Triodia triticoides C.A. Gardner, Fl. W. Australia |: 78. 
1952. T-Protol.: Australia: Western Australia: 
Kimberley Research Station, Ord River, 28 Jan 1949, 
E.C.B. Langfield 32 |HT: PERTH, CANB]. 

= Tridens flavus (L.) Hitche. 


Triphlebia Stapf, Fl. Cap. 7: 318. 1898. TYPE: Triphlebia 
alopecuroides (Hack.) Stapf. 
NOTE: hom. illeg.; see also Hook. Ic. Pl. 27: 1612 (1899); 
as syn. of Stiburus Stapf in Fl. Cap. 7(4): 696 (1900). 
= Eragrostis Wolf. 


Triplasis Pp. Beauv., Ess. Agrostogr. 81. 1812. TYPE: 
Triplasis americana P. Beauv. 
TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.:; 2, SUBSP.: 2. 
SYN.: Diplocea Raf.; Merisachne Steud.: Uralepis Nutt. 


Triplasis americana P. Beauv., Ess. Agrostogr. 81, t. 16, f. 


10. 1812. T-Protol.: [USA]: "des Etats-Unis 
d'Amerige.", M. Delille s.n.. communiquee par M. 
Delille. 

DST.: USA. 


LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Anonymous (1986); Hitchcock, A. S. 
(1951); Radford, A. E., H. E. Ahles & C. R. Bell 
(1968); Small, J. K. (1933); Wunderlin, R. P. (1998). 

SYN.: Sieglingia americana (P. Beauv.) Beal; Tricuspis 
cornuta (Elliott) A. Gray; Triplasis cornuta Benth. ex 
W.D. Jacks.; Uralepis cornuta Elliott. 


Triplasis caribensis (R.W. Pohl) Beetle, Phytologia 54(1): 5. 
1983. Basionym: Triplasis purpurea var. caribensis 
R.W. Pohl. 
= Triplasis purpurea (Walter) Chapm. 


Triplasis cornuta Benth. ex W.D. Jacks., Index Kew. 2: 1121. 
1895, 
NOTE: nom. inval., as syn. of Triplasis americana P. 
Beuv.; based on Uralepsis cornuta Ell. 
= Triplasis americana P. Beauv. 


Triplasis floridana Gand., Bull. Soc. Bot. France 66(7): 303. 
1919 [1920]. T-Specim.: | of 2. USA: Florida: Lee 
Co.: Punta Rassa, 1900, 4.8. Hitchcock 533 |ST: LY, 
US-387544]. 2 of 2. USA: Mississippi: Deer Island, 
Tracy 4561 |ST: LY]. 

= Triplasis purpurea (Walter) Chapm. 


Triplasis glabra Gand., Bull. Soc. Bot. France 66(7): 303. 
1919 [1920]. T-Protol.: [USA]: America borealis. T- 
Specim.: | of 2. USA: Florida: Sneed's Island, 1899, 
Tracy 6461 [ST: LY, US-1109617]. 2 of 2. USA: 
Rhode Islad: Providence, MacDougal s.n. [ST: LY]. 

= Triplasis purpurea (Walter) Chapm. 


Triplasis intermedia Nash, Bull. Torrey Bot. Club 25: 564. 
1898. T-Protol.: [USA]: Florida: Point near Tampa, 20 
Aug 1895, G.V. Nash 2426. T-Specim.: | of 1. USA: 
Florida: Hillsborough Co., 20 Aug 1895,G./ Nash 
2426 (IT: US-252278]. 
= Triplasis purpurea (Walter) Chapm. 


Catalogue of New World Grasses: II. 


231 


Triplasis purpurea (Walter) Chapm., Fl. South. U.S. 560. 
1860. Basionym: Aira purpurea Walter. 

DST.: Cana, CoRi, Hond, USA. 

LVL. ACCPTC.: I. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Davidse, G., M. Sousa S. & 
A. O. Chater (editores generales) (1994); Fernald, M. 
(1950); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Great Plains 
Flora Association (1986); Hitchcock, A. S. (1951); 
Long, R. W. & O. Lakela (1971); Pohl, R. W. (1980); 
Pohl, R. W. (1994): Radford, A. E., H. E. Ahles & C. 
R. Bell (1968); Scoggan, H. J. (1978 [1979]); Small, 
J. K. (1933); Voss, E. G. (1972); Wunderlin, R. P. 
(1998). 

Aira purpurea Walter; Diplocea barbata Raf.; 
Festuca brevifolia Muhl., Festuca brevifolia Muhl.; 
Festuca purpurea (Walter) F. Muell.; Glyceria 
brevifolia (Muhl.) Schult.; Merisachne drummondii 
Steud.; Panicularia  brevifolia | (Muhl.) Porter; 
Sieglingia purpurea (Walter) Kuntze; Tricuspis 
purpurea (Walter) A. Gray; Triodia purpurea 
(Walter) Smyth; 7riplasis caribensis (R.W. Pohl) 
Beetle; Triplasis floridana Gand.; Triplasis glabra 
Gand.; Triplasis intermedia Nash; Triplasis 
sparsiflora Chapm.; Uralepis  aristulata Nutt; 
Uralepis purpurea (Walter) Nutt. 


SYN:: 


Triplasis purpurea var. caribensis R.W. Pohl, Iowa State J. 
Res. 47(1): 76, f. 3. 1972. T-Protol.: Costa Rica: 
Limon: sandy, open beach, between Limén Airport 
and the Rio Banano along Caribbean Coast, 2 m, 
scattered but forming an extensive stand, 16 Sep 1969, 
Pohl & Davidse 11080 [HT: ISC; IT: CR, F, US]. T- 
Specim.: | of 1. Costa Rica: Lim6én: between Limon 
Airport and the Rio Banano, 2 m,9.56N 83.00W, 16 
Sep 1968, R.W. Pohl & G. Davidse 11080 (HT: ISC, 
IT: CR-50763, F, US-2644177]. 

DST.: CoRi, Hond. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Espejo Serna, A., 
A. R. Lépez-Ferrari, and J. Valdes-Reyna (2000); 
Pohl, R. W. (1980); Pohl, R. W. (1994). 

SYN.: Triplasis caribensis (R.W. Pohl) Beetle. 


Triplasis purpurea (Walter) Chapm. var. purpurea. 
DST.: Cana, USA. 
LVL. ACCPTC:: 1. 


Triplasis setacea Griseb., Abh. Konigl. Ges. Wiss. Gottingen 
24: 304. 1879. T-Protol.: Argentina. T-Specim.: | of 2. 
Argentina: Catamarca, 22 Feb 1873, P.G. Lorentz 337 
(ST: US-294571]. 2 of 2. Argentina: Salta: La Merced 
S., ad fl. Juramento [ST]. 

= Tripogon spicatus (Nees) Ekman. 


Triplasis sparsiflora Chapm., Bot. Gaz. 3: 19. 1878. T- 
Protol.; [USA]: Florida: Punta Rassa, S. Florida, 
Chapman s.n., [specimen affected by fungus]. T- 
Specim.: 1 of 1. USA: Florida, Oct 1875, A.W. 
Chapman s.n. [IT: US-980578]. 

= Triplasis purpurea (Walter) Chapm. 


Triplathera (Endl.) Lindl., Veg. Kingd. (ed. 2) 116. 1846. 
Basionym: Eutriana sect. Triplathera Endl. 
= Bouteloua Lag. 


Tripogon Roem. & Schult., Syst. Veg. 2: 34, 600. 1817. 
TYPE: Tripogon bromoides Roth ex Roem. & Schult. 
TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 2. 
CNSLTD. LIT.: Phillips, S. M. & E. Launert, (1971). 
SYN.: Arcangelina Kuntze; Kralikia Coss. & Durieu; 
Kralikiella Batt. & Trab.; Plagiolvtrum Nees. 


Tripogon ekmanii Nicora & Rugolo, Candollea 46(2): 533. 
1991. T-Protol.: Paraguay: In regione fluminis Alto 
Parana, 1909-1910., K. Fiebrig 53905 [HT: SI; IT: 
BAA, G, LIL]. 

DST.: Arge. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Nicora, E. G. & Z. E. Rugolo de 
Agrasar (1991); Zuloaga, F. O., E. G. Nicora, Z. E. 
Rugolo de Agrasar, O. Morrone, J. Pensiero & A. M. 
Cialdella (1994). 


Tripogon spicatus (Nees) Ekman, Ark. Bot. 11(4): 36. 1912. 
Basionym: Bromus spicatus Nees. 

DST.: Arge, Boli, Braz, Cari, Chil, Ecua, Guat, Hond, Mexi, 
Nica, Para, Peru, USA, Urug, Vene. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Allem, A. C. & J. F. M. Valls 
(1987); Anonymous (1986); Beetle, A. A. (1977); 
Brako, L. & J. L. Zarucchi (1993); Burkart, A. 
(1969): Cabrera, A. L. (1970); Correll, D. S. & M. C. 
Johnston (1970); Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Espejo Serna, A., 
A. R. Lopez-Ferrari, and J. Valdes-Reyna (2000); 
Foster, R. C. (1958); Hitchcock, A. S. (1927); 
Hitchcock, A. S. (1936); Hitchcock, A. S. (1951): 
Jorgensen, P. M. & S. Le6n-Yanez (eds.) (1999): 
Judziewicz, E. J. (1990); Killeen, T. J. (1990); 
Macbride, J. F. (1936); Marticorena, C. & M. 
Quezada (1985); McVaugh, R. (1983); Nicora, E. G. 
(1995); Pohl, R. W. (1994); Renvoize, S. A. (1984); 
Renvoize, S. A. (1988); Renvoize, S. A. (1998); 
Rosengurtt, B., B. R. Arrillaga de Maffei & P. 
Izaguirre de Artucio (1970); Tovar, O. (1993); 
Zuloaga, F. O., E. G. Nicora, Z. E. Rugolo de 
Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella 
(1994). 

SYN.: Bromus spicatus Nees; Diplachne reverchonii Vasey; 
Diplachne simplex Doll; Diplachne spicata (Nees) 
Doll; Leptochloa spicata (Nees) Scribn.; Rabdochloa 
spicata. (Nees) Kuntze ex Stuck.; Sieglingia 
schaffneri (S. Watson) Kuntze; Sieglingia spicata 
(Nees) Kuntze ex Stuck.; Tricuspis simplex Griseb.; 
Triodia schaffneri S. Watson; Triplasis setacea 
Griseb. 


Triraphis R. Br., Prodr. 185. 1810. TYPE: Triraphis 
pungens R. Br.; LT designated by Maire, Fl. Afrique 
Nord 2: 191 (1953). 
TRTD. BY: Peterson, P. M. 
NHT: Eleusininae. NO. ACCPTD. SPP.: 2. 


Triraphis devia Filg. & Zuloaga, Novon 9(1): 36, f. 1. 1999. 
T-Protol.: Brazil: Goias: Chapada dos Veadeiros, 
municipio de Alto Paraiso, ca. 17 km da cidade, em 
diregdo a Colinas de Goias, 14.09.30S 47.39.54W, ca. 
1200 m, ereto, cerca 60 cm de altura, campo limpo, 23 
Nov 1994, 7.S. Filgueiras & D. Alvarenga 3168 [HT: 
IBGE; IT: F, ICN, K, MEXU, MO, SI, SP, US]. T- 


232 


Catalogue of New World Grasses: II. 


Specim.: | of 1. Brazil: Goias, Alto Paraiso Mun., 
Chapado dos Veadeiros, ca.l17 km de cidade em 
diregdéo a Colinas de Goias, 23 Nov 1994, TS. 
Filgueiras & D. Alvarenga 3168 (IT: US-3382566]. 

DST.: Braz. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Filgueiras, T. S. & F. O. Zuloaga 

(1999). 


Triraphis mollis R. Br., Prodr. 185. 1810. T-Protol.: Littora 
Novae Hollandiae intra tropicum. T-Specim.: | of 1.2. 
Br. 6248 (HT: K]. 
DST.: USA. 
LVL. ACCPTC.: 1; introduced. 
ALSO ACCPTD. BY: Anonymous (1986). 


Trisiola Raf., Fl. Ludov. 144. 1817. 
NOTE: nom. superfl. pro Uniola. 
= Uniola L. 


Trisiola paniculata (L.) Raf., Fl Ludov. 144, 1817. 
Basionym: Uniola paniculata L. 


= Uniola paniculata L. 


Tristania Poir., Encycl. 4: 526. 1816. 
NOTE: nom. prov. pro Ponceletia. 
= Spartina Schreb. 


Triticum geminatum Spreng., Syst. Veg. 1: 326. 1825, T- 
Protol.: Uruguay: Monte Video, Sellow. 
= Eleusine tristachya (Lam.) Lam. 


Triunila Raf., Neogenyton 4. 1825. 
NOTE: nom. prov. pro Uniola. 
= Uniola L. 


Tuctoria Reeder, Amer. J. Bot. 69: 1090. 1982. TYPE: 


Tuctoria fragilis (Swallen) Reeder. 
TRTD. BY: Peterson, P. M. 
NHT: Orcuttieae. NO. ACCPTD. SPP.: 3. 
CNSLTD. LIT.: Reeder, J. R., (1982). 


Tuctoria fragilis (Swallen) Reeder, Amer. J. Bot. 69(7): 
1090. 1982. Basionym: Orcuttia fragilis Swallen. 

DST.: Mexi. 

LVL. ACCPTC:: 1. 

ALSO ACCPTD. BY: Espejo Serna, A., A. R. Lépez- 
Ferrari, and J. Valdes-Reyna (2000); Reeder, J. R. 
(1982). 

SYN.: Orcuttia fragilis Swallen. 


Tuctoria greenei (Vasey) Reeder, Amer. J. Bot. 69(7): 1091, 
1982. Basionym: Orcuttia greenei Vasey. 
DST.: USA. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Hickman, J. C. (1993); Reeder, J. R. 
(1982). 
SYN.: Orcuttia greenei Vasey. 


Tuctoria mucronata (Crampton) Reeder, Amer. J. Bot. 
69(7): 1091. 1982. Basionym: Orcuttia mucronata 
Crampton. 

DST.: USA. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Hickman, J. C. (1993); Reeder, J. R. 
(1982). 

SYN.: Oreuttia mucronata Crampton. 


Uniola L., Sp. Pl. 1: 71. 1753. TYPE: Uniola paniculata 

L.; LT designated by Nash in Britton & Brown, Ill. Fl. 
N. U.S. (ed. 2) 1: 248 (1913). 

TRTD. BY: Peterson, P. M. 

NHT: Uniolinae. NO. ACCPTD. SPP.: 5. 

CNSLTD. LIT.: Yates, H. O., (1966); Yates, H. O., (1966). 

SYN.: Leptochloopsis H.O. Yates; Nevroctola Raf.; Trisiola 

Raf.; Triunila Raf. 


Uniola condensata Hitche., Contr. U.S. Natl. Herb. 24(8): 
345. 1927. T-Protol.: Ecuador: Prov. of Chimborazo: 
collected on dry grassy slopes at Huigra, alt. 1200 m, 4 
July 1923, A.S. Hitchcock 20333 |HT: US-1164907]. 
T-Specim.: 1 of 1. Ecuador: Chimborazo: on dry 
grassy slopes at Huigra, alt. 1200 m, 4 Jul 1923,4.8. 
Hitchcock 20333 [HT: US-1164907; IT: F, GH, NY]. 

DST.: Ecua. 

LVL. ACCPTC.:: 1. 

ALSO ACCPTD. BY: Hitchcock, A. S. (1927); Jorgensen, 
P.M. & S. Leon-Yanez (eds.) (1999); Lagaard, S. & 
I. Sanchez V. (1990); Valencia, R., N. Pitman, S. 
Leon-Yanez & P. M. Jorgensen (2000). 

SYN.: Leptochloopsis condensata (Hitchc.) H.O. Yates. 


Uniola distichophylla Labill., Nov. Holl. Pl. 1; 21, t. 24. 
1804. T-Protol.: Habitat in capite Van-Diemen, 
= Distichlis spicata subsp. stricta (Torr.) Thorne, 


Uniola distichophylla (Michx.) Roem. & Schult., Syst. Veg. 
2: 596. 1817. Basionym: Festuca distichophylla 
Michx,. 

NOTE: nom. illeg. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Uniola flexuosa Buckley, Proc. Acad. Nat. Sci. Philadelphia 
14: 99. 1862. T-Protol.: USA: Texas: On the Brazos at 
Fort Belknap, Humboldt & Bonpland s.n. T-Specim.: | 
of 1. USA: Texas: on the Brazos at Fort Belknap, 
Buckley s.n. [LT: PH], LT designated by Hitchcock, 
Man. Grass. U.S. 838 (1935), but without citing a 
specific sheet in a specific herbarium. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Uniola floridana Gand., Bull. Soc. Bot. France 66(7):; 304. 
1919 [1920]. T-Protol.: Florida: in insula Sta Rosa, 
Tracy 4545, T-Specim.: | of 1. USA: Florida: Santa 
Rosa Island, 29 Aug 1899, S.M. Tracy 4545 [HT: LY; 
IT: US-383700]. 

= Uniola paniculata L. 


Uniola heterochroa Gand., Bull. Soc. Bot. France 66(7): 304. 
1919 [1920]. T-Protol.: Florida: ad Punta Rassa Lee, 
Hitchcock 335. T-Specim.: | of 1. USA: Florida: Lee 
Co.: ad Punta Rassa Lee, no date, 4.8. Hitchcock 535 
[HT: LY; IT: US-387546]. 

= Uniola paniculata L. 


Uniola macrostachys Gand., Bull. Soc. Bot. France 66(7): 
304. 1919 [1920]. T-Protol.: America sept., Louisiana 
in insula Breton, SM. Tracy & Lloyd 462. T-Specim.: 
1 of 1. USA: Louisiana, 17 Sep 1900,S.M4 Tracy & 
Lloyd 462 [HT: LY; IT: US-383665]. 

= Uniola paniculata L. 


Uniola maritima Michx., Fl. Bor.-Amer. 1: 71. 1803. T- 
Protol.: USA: "Hab. in littoribus arenosis, a Virginia 
ad Ludovisianam.", Michaux s.n. T-Specim.: | of 1. 


Catalogue of New World Grasses: II. 


233 


USA: Carolina, sur la bord de la mer, Michaux s.n. 
[HT: P]. 

NOTE: nom. inval., U. paniculata L. cited. 

= Uniola paniculata L. 


Uniola multiflora Nutt., Trans. Amer. Philos. Soc., n.s., 5: 
148. 1837. T-Protol.: Arkansas Territory: Hab. on the 
sand beaches of the Arkansas, above the garrison, 
Nutall s.n. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Uniola palmeri Vasey, Gard. & Forest 2: 401, f. 124. 1889. 
T-Protol.: Mexico: Sonora: Colorado Rv., Horseshoe 
Bend, 35 mi. S of Lerdo, 12-15 mi from its mouth, Apr 
1889, Palmer s.n. T-Specim.: | of 1. Mexico: Sonora: 
Colorado River, Horseshoe Bend, 35 mi S of Lerdo, 
12-15 mi from its mouth, Apr 1889, £. Palmer s.n. 
{IT: IBUG-87337, US-913540, US-913539, US- 
81764, US-81763]. 

= Distichlis palmeri (Vasey) Fassett ex I.M. Johnst. 


Uniola paniculata L., Sp. Pl. 1: 71. 1753. T-Protol.: USA: 
"In Carolina.". T-Specim.: 1 of 1. USA, Anon. [LT: 
LINN-98.1], LT designated by Hitchcock, Contr. U.S. 
Natl. Herb. 12: 121 (1908); also by Yates, Southw. 
Nat. 11: 377 (1966). 

DST.: Cari, Mexi, USA. 

LVL. ACCPTC.: 1; cultivated. 

ALSO ACCPTD. BY: Anonymous (1986); Beetle, A. A. 
(1977); Catasus Guerra, L. (1997); Correll, D. S. & 
M. C. Johnston (1970); Cowan, C. P. (1983); Darke, 
R. (1999); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Fernald, 
M. (1950); Gleason, H. A. & A. Cronquist (1968); 
Gleason, H. A. & A. Cronquist (1991); Hitchcock, A. 
S. (1909); Hitchcock, A. S. (1936); Hitchcock, A. S. 
(1951); Long, R. W. & O. Lakela (1971); Pohl, R. W. 
& G. Davidse (1994); Radford, A. E., H. E. Ahles & 
C. R. Bell (1968); Small, J. K. (1933); Wunderlin, R. 
P. (1998); Yates, H. O. (1966). 

SYN.: Briza caroliniana Lam.; Nevroctola maritima Raf. ex 
B.D. Jacks.; Nevroctola paniculata Raf. ex B.D. 
Jacks.; Trisiola paniculata (L.) Raf.; Uniola 
floridana Gand.; Uniola heterochroa Gand.; Uniola 
macrostachys Gand.; Uniola maritima Michx. 


Uniola peruviana Legaard & Sanchez Vega, Nordic J. Bot. 
10(4): 439. 1990. T-Protol.: Peru: Dpto. Amazonas: at 
bofer between Prov. Bongar and luya, following the 
road Pedro Ruiz to Bagua, 700 m a.s.1., 23 Jun 1981, 
Sanchez Vega 2642 [HT: CPUN; IT: AAU, K, USM]. 

DST.: Peru. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Laegaard, S. & I. Sanchez V. (1990); 
Tovar, O. (1993). 


Uniola pittieri Hack., Oesterr. Bot. Z. 52: 309. 1902. T- 
Protol.: Costa Rica: in ora maritima Pacifica ad Baie 
de Salinas, Pittier, 2 syntypes. T-Specim.: | of 2. 
Costa Rica: in ora maritima Pacifica ad Baie de 
Salinas, no date, H. Pittier 2688 [LT: W; ILT: BM, 
US-825669], LT designated by Yates, Southw. 
Naturalist 11: 380 (1966). 2 of 2. Costa Rica, Pittier 
6826 [ST]. 
NOTE: as Uniloa "Pittieri". 
DST.: Colo, CoRi, Ecua, ElSa, Guat, Hond, Mexi, Nica, 
Pana, Peru. 


LVL. ACCPTC.: |. 

ALSO ACCPTD. BY: Beetle, A. A. (1977); Breedlove, D. 
E. (1986); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J. Valdes-Reyna (2000); Gould, F. 
W. & R. Moran (1981); Hitchcock, A. S. (1927); 
Jorgensen, P. M. & S. Leoén-Yanez (eds.) (1999); 
Lawesson, J. E., H. Adsersen & P. Bentley (1987); 
McVaugh, R. (1983); Pohl, R. W. & G. Davidse 
(1994); Pohl, R. W. (1980); Porter, D. M. (1983); 
Reeder, J. R. & C. G. Reeder (1971); Swallen, J. R. 
(1955); Tovar, O. (1993); Wiggins, I. L. & D. M. 
Porter (1971); Yates, H. O. (1966). 


Uniola prostrata (Kunth) Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
4,.2(1): 55. 1836. Basionym: Poa prostrata Kunth. 

= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Uniola racemiflora Trin. Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
4,2(1): 55. 1836. T-Protol.: "V. sp. Doming."(Mis. b. 
Sprengel s.n. Eleusines proceras sibi.), Sprengel s.n. T- 
Specim.: | of 1. [HT: LE-TRIN-2500.01 (fragm.)]. 

= Uniola virgata (Poir.) Griseb. 


Uniola scoparia (Nees ex Kunth) Steud., Syn. Pl. Glumac. 1: 
281. 1854. Basionym: Poa scoparia Nees ex Kunth. 
= Distichlis scoparia (Nees ex Kunth) Arechav. 


Uniola sparta Trin., Linnaea 10(3): 307. 1836. T-Protol.: 
Cuba: in rupibus aridis, Poeppig s.n. T-Specim.: | of 
1. Cuba [HT?: LE-TRIN-2502.01; IT: US (fragm. ex 
LE-TRIN)], mis. Poppig, 1834: pistillum. 

= Uniola virgata (Poir.) Griseb. 


Uniola spicata L., Sp. Pl. 71. 1753. T-Protol.: In America 
borealis maritimis. T-Specim.: | of |. USA, Kalm s.n. 
[LT: LINN-89.4], LT designated by Hitchcock, Contr. 
U.S. Natl. Herb. 12: 121 (1908). 
= Distichlis spicata (L.) Greene. 


Uniola stricta Torr., Ann. Lyceum Nat. Hist. New York 1(1): 
155-156. 1824. T-Protol.: USA: Oklahoma: shores of 
the Canadian River, Marcy exped. s.n. T-Specim.: | of 
1. /misit Hooker] T. 1/13 [T?: LE-TRIN-2505.02a 
(fragm.)]. 

NOTE: as "Uniola stricta"?. 
= Distichlis spicata subsp. stricta (Torr.) Thorne. 


Uniola thalassica (Kunth) Trin., Mem. Acad. Imp. Sci. St.- 
Petersbourg, Ser. 6, Sci. Math. 1(4): 359. 1830. 
Basionym: Poa thalassica Kunth. 

= Distichlis spicata (L.) Greene. 


Uniola virgata (Poir.) Griseb., Fl. Brit. W. I. 531. 1864. 
Basionym: Poa virgata Poir. 

DST.: Cari. 

LVL. ACCPTC.: 1. 

ALSO ACCPTD. BY: Catasus Guerra, L. (1997); 

Hitchcock, A. S. (1909); Hitchcock, A. S. (1936); 

Liogier, H. & L. Martorell (1982). 

Eleusine procera Spreng. ex Steud.; Leptochloopsis 

virgata (Poir.) H.O. Yates; Poa virgata Poir., Uniola 

racemiflora Trin.; Uniola sparta Trin. 


SYN:: 


Uniolinae subtrib. Clayton, Kew Bull. 37: 417. 1982. 
TYPE: Uniola L. 


234 


Catalogue of New World Grasses: II. 


TRTD. BY: Soreng, R. J., G. Davidse, T. S. Filgueiras, E. J. 
Judziewicz, P. M. Peterson, and F. O. Zuloaga. 
NHT: Cynodonteae. 


Uralepis Nutt., Gen. N. Amer. Pl. 1: 62. 1818. TYPE: 
Uralepsis purpurea (Walter) Nutt. 

NOTE: as "Uralepsis". The derivation accompanying the 
generic name indicates that the spelling Uralepsis 
was a typographical error (fide Hitchcock, U.S.D.A. 
Bull. 772: 76 (1920). 

= Triplasis P. Beauv. 


Uralepis ambigua (Elliott) Kunth, Revis. Gramin. 1: 108. 
1829. Basionym: Poa ambigua Elliott. 
= Tridens ambiguus (Elliott) Schult. 


Uralepis anderssonii F. Aresch., PI. Itin. Eugenia 119. 1910. 
T-Protol.: Ecuador: circa Guajaquil et in insula Puna, 
NJ. Andersson s.n. 
= Leptochloa fusca subsp. uninervia (J. Presl) N. Snow. 


Uralepis aristulata Nutt., Gen. N. Amer. Pl. 1: 63. 1818. T- 
Protol.. USA: Delaware: near Wilmington, Dr. 
Baldwin s.n. 

NOTE: as "Uralepsis". 
= Triplasis purpurea (Walter) Chapm. 


Uralepis avenacea (Kunth) Kunth, Revis. Gramin. 1: 108. 
1829. Basionym: Triodia avenacea Kunth. 
= Erioneuron avenaceum (Kunth) Tateoka. 


Uralepis avenacea vat. viridiflora E. Fourn., Mexic. Pl. 2: 
110. 1886. T-Protol.: Mexico: San Louis Potosi, Virlet 
1379. 
NOTE: nom. nud. 
= Erioneuron avenaceum (Kunth) Tateoka. 


Uralepis brasiliensis Nees ex Steud., Syn. Pl. Glumac. 1: 
248. 1854. T-Protol.: Brasil. 
= Tridens brasiliensis (Nees ex Steud.) Parodi. 


Uralepis brevicuspidata Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 93-94. 1862. T-Protol.: Northern 
Texas, May. T-Specim.: | of 1. USA: northern Texas, 
Wright 767 [LT: PH; US-2786821 (photo)], LT 
designated by Hitchcock, Man. Grass. U.S. 877 
(1935), but without citing a specific sheet in a specific 
herbarium, 

= Leptochloa dubia (Kunth) Nees. 


Uralepis cornuta Elliott, Sketch Bot. S. Carolina 1: 580. 
1821. T-Protol.: sine loc. 
= Triplasis americana P. Beauv. 


Uralepis cuprea (Jacq.) Kunth, Revis. Gramin. 1: 108. 1829, 
Basionym: Triodia cuprea Jacq. 
= Tridens flavus (L.) Hitche. 


Uralepis densiflora Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 94, 1862. T-Protol.: Middle Texas. T- 
Specim.: | of 1, USA: Texas, Buckley s.n. (IST: US 
(fragm. ex PH)]. 

= Tridens strictus (Nutt.) Nash. 


Uralepis elongata Buckley, Proc. Acad. Nat. Sci. 
Philadelphia 14: 89. 1862. T-Protol.: Northern Texas, 
May. T-Specim.: | of 1. USA: Texas, Wright 2054 
{IST: US (fragm. ex PH)]. 

= Tridens muticus var. elongatus (Buckley) Shinners. 


Uralepis flaccida Déll, Fl. Bras. 2(3): 95, pl. 27. 1878. T- 
Protol.: Brazil: Goias: ad Porto Imperial, Burchell 
8477 (HT: KR?; IT: K, P, US (fragm. ex FB)], Fragm. 
from FB, herb. disc., HT may be at KR. T-Specim.: | 
of 1. Brazil: Prov. Goyas: Porto Imperial, Burchell [IT: 
US (fragm. ex FB)], fragm. from Freiburg (FB), herb. 
disc., may be at KR. 

= Tridens flaccidus (D6ll) Parodi. 


Uralepis fusca (L.) Steud., Syn. Pl. Glumac. 1: 247. 1854. 
Basionym: Festuca fusca L. 
= Leptochloa fusea (L.) Kunth. 


Uralepis mutica E. Fourn. ex Hemsl., Biol. Cent.-Amer., Bot. 
3: 569. 1885. 
NOTE: nom. inval., as syn. of Triodia mutica Benth. 
= Tridens muticus (Torr.) Nash. 


Uralepis pilosa Buckley, Proc. Acad. Nat. Sci. Philadelphia 
14: 94. 1862. T-Protol.: Middle Texas, Name from 
page 94. T-Specim.: | of 2. USA: Texas, Buckley s.n. 
[LT: PH; ILT: P-DU?], LT designated by Hitchcock, 
Man. Grass. U.S. 971 (1935), but without citing a 
specific sheet in a specific herbarium. 2 of 2. USA: 
western Texas, Wright 78] [LT: PH; ILT: US 
(fragm.)], LT designated by Valdés-Reyna & Hatch, 
Sida 17: 657 (1997). 

NOTE: This name was published again by Buckley on the 
next page in this publication, with a different 
description, and a different type locality.—gd/Jan 
1995, 

= Erioneuron pilosum (Buckley) Nash. 


Uralepis pilosa Buckley, Proc. Acad. Nat. Sci. Philadelphia 
14: 95. 1862. T-Protol.: Northern Texas, type is 
actually from Western Texas. T-Specim.: | of 3. 
Western Texas, Wright 779 (IST: US (fragm. ex PH)]. 
2 of 3. USA: Texas, Wright 760 [ST]. 3 of 3. USA: 
New Mexico, Wright 2046 [ST: PH]. 

NOTE: This name was previously published by Buckley in 
this same publication on p. 94 with a different 
description and a different type locality. —gd. 

= Tridens muticus (Torr.) Nash. 


Uralepis pulchella (Kunth) Kunth, Revis. Gramin. 1; 108. 
1829. Basionym: Triodia pulchella Kunth. 
= Dasyochloa pulchella (Kunth) Willd. ex Rydb. 


Uralepis purpurea (Walter) Nutt., Gen. N. Amer. Pl. 1: 62. 
1818. Basionym: Aira purpurea Walter. 
NOTE: as "Uralepsis". 
= Triplasis purpurea (Walter) Chapm. 


Uralepis tricuspis (Trin.) Steud., Nomencl. Bot. (ed. 2) 1: 
564. 1840. Basionym: Eragrostis tricuspis Trin. 
= Tridens flavus (L.) Hitchce. 


Uralepis verticillata (Nees & Meyen) Steud., Syn. Pl. 
Glumac. 1: 248. 1854. Basionym: Diplachne 
verticillata Nees & Meyen. 

= Leptochloa fusea subsp. uninervia (J. Pres!) N. Snow. 


Uralepis virens (Nees) Kunth, Enum. Pl. 1: 319. 1833. 
Basionym: 7ridens virens Nees. 
NOTE: also Revis. Gram. 3: Suppl. 1834 p. (as in 1835 ed.) 
XXVII. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Uralepsis composita Buckley, Proc. Acad. Nat. Sci. 


Philadelphia 14: 94-95. 1862. 


Catalogue of New World Grasses: II. 


235 


NOTE: orth. var. 
CORRECT NAME: Uralepis composita Buckley. 
= Leptochloa fusca subsp. fascicularis (Lam.) N. Snow. 


Uralepsis verticillata (Nees & Meyen) Steud., Syn. PI. 
Glumac. 1: 248. 1854. Basionym: Diplachne 
verticillata Nees & Meyen. 

NOTE: orth. var. 
= Leptochloa fusea subsp. uninervia (J. Presl) N. Snow. 


Vaseya Thurb., Proc. Acad. Nat. Sci. Philadelphia 15: 79. 
1863. TYPE: Vaseya comata Thurb. 
= Muhlenbergia Schreb. 


Vaseya comata Thurb., Proc. Acad. Nat. Sci. Philadelphia 
1863: 79. 1863. T-Protol.: USA: a native of the plains 
of Nebraska [Wyoming], Hall & Harbour 685. T- 
Specim.: | of |. USA: Wyoming, 1862,F. Hall & JP. 
Harbour 685 |IT: US-907389}. 

= Muhlenbergia andina (Nutt.) Hitchc. 


Vaseyochloa hitche., J. Wash. Acad. Sci. 23(10): 452. 
1933. TYPE: 
Hitche. 

TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: 1. 


Vaseyochloa_ multinervosa (Vasey) 


Vaseyochloa multinervosa (Vasey) Hitchc., J. Wash. Acad. 
Sci. 23(10): 452. 1933. Basionym: Melica 
multinervosa Vasey. 

DST.: USA. 

LVL. ACCPTC.:: |. 

ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Hitchcock, A. S. (1951). 

SYN.: Distichlis multinervosa (Vasey) Piper; Melica 
multinervosa Vasey; Triodia multinervosa (Vasey) 
Hitche. 


Vilfa adusta Trin., Mem. Acad. Imp. Sci. Saint-Petersbourg, 
Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 6,4(1-2): 80. 
1840. T-Protol.: Brasilia: in campis desertorum prope 
Rio de S. Francisco, Pohl s.n. T-Specim.: | of 1. 
Brazil: Trichod, AB.E. Pohl 326 [HT: LE-TRIN- 
1673.01; IT: US-1126778]. 

= Sporobolus aeneus (Trin.) Kunth. 


Vilfa aenea Trin., Gram. Panic. 23. 1826. T-Protol.: Brazil: " 
specimen Brasiliense.", Langsdorff s.n. T-Specim.: 1 
of |. Brazil, G.H. von Langsdorff s.n. [T: LE-TRIN- 
1647.01?]. 

= Sporobolus aeneus (Trin.) Kunth. 


Vilfa aenea var. angustifolia Doll, Fl. Bras. 2(3): 34. 1878. T- 
Protol.: Brazil: ad Rio S. Francisco, Pohl 326, 
apparently the same type as Vilfa adutsa Trin. 

= Sporobolus aeneus (Trin.) Kunth. 


Vilfa aenea var. latifolia DOll, Fl. Bras. 2(3): 34. 1878. T- 
Protol.: Brazil. 
= Sporobolus aeneus (Trin.) Kunth. 


Vilfa agrostoidea Buckley, Proc. Acad. Nat. Sci. Philadelphia 
14: 88. 1862. T-Protol.: USA: Texas: Llano County. 
= Sporobolus pyramidatus (Lam.) Hitche. 


Vilfa airoides (Torr.) Trin. ex Steud., Nomencl. Bot. (ed. 2) 
2: 766. 1841. Basionym: Agrostis airoides Torr. 
= Sporobolus airoides (Torr.) Torr. 


Vilfa angusta Buckley, Proc. Acad. Nat. Sci. Philadelphia 14: 
88. 1863. 
NOTE: nom. illeg., published as Sporobolus (Vilfa) 
angusta, within a treatment of Vilfa. 
CORRECT NAME: Sporobolus angustus Buckley. 
= Sporobolus indicus (L.) R. Br. 


Vilfa arachnoidea (Poir.) P. Beauv., Ess. Agrostogr. 147, 
181. 1812. Basionym: Agrostis arachnoides Poir. 
= Muhlenbergia expansa (Poir.) Trin. 


Vilfa arguta Nees, Fl. Bras. Enum. Pl. 2: 395, 1829. T- 
Protol.: Brazil. T-Specim.: 1 of 1. Brasilia, #. Se//ow 
[ST: LE-TRIN-1676.01 (& fig.)]. 
= var. Beta ad Montevideo, Uruguay. 
CORRECT NAME: Vilfa humifusa var. major E. Fourn. 
= Sporobolus pyramidatus (Lam.) Hitchc. 


Vilfa arkansana Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 64. 1840. T-Protol.: USA: Arkansas 
[territory], Beyrich s.n. T-Specim.: 1 of 1. [HT: LE- 
TRIN-1677.01; IT: US (fragm. ex LE-TRIN)]. 

= Sporobolus pyramidatus (Lam.) Hitchc. 


Vilfa aspera P. Beauv., Ess. Agrostogr. 16, 147, 181. 1812. 
REPLACED NAME: Agrostis aspera Michx. 
BLOCKING NAME: Agrostis aspera Weber. 
= Sporobolus compositus (Poir.) Merr. 


Vilfa asperifolia Nees & Meyen ex Trin., Mem. Acad. Imp. 
Sci. Saint-Petersbourg, Ser. 6, Sci. Math., Seconde Pt. 
Sci. Nat. 6,4(1-2): 95. 1840. T-Protol.: Chile: 
Atacama: Rio Mayno et Copiap6o, /Meyen] s.n. T- 
Specim.: | of 2. Chile: Rio Mayno, 1000' alt, Mar 
1831, Meyen s.n. [ST: B, BAA-3464, LE-TRIN- 
1679.01 b]. 2 of 2. Chile: Copiap6, Meyen s.n. [ST: 
LE-TRIN-1679.01 a]. 

= Muhlenbergia asperifolia (Nees & Meyen ex Trin.) 
Parodi. 


Vilfa asperifolia Meyen, Nov. Actorum Acad. Caes. Leop.- 
Carol. German. Nat. Cur. Sup. 1: 9. 1841. 
NOTE: isonym; also Reis. Erd. 1:408. 1834, nom. nud. 
E. NAME: Vilfa asperifolia Nees & Meyen ex Trin. 
= Muhlenbergia asperifolia (Nees & Meyen ex Trin.) 
Parodi. 


Vilfa barbata P. Beauv., Ess. Agrostogr. 16, 147, 181. 1812. 
NOTE: nom. illeg. superfl.; based on Agrostis barbata Pers. 
= Agrostis littoralis Lam. 
= Sporobolus virginicus (L.) Kunth. 


Vilfa berteroana Trin, Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 100. 1840. T-Protol.: Santo Domingo, 
Bertero s.n. T-Specim.: 1 of 1. In Santo Domingo lect., 
Bertero sn. [HT: LE-TRIN-1682.01 (with drawing 
and fragm.); IT: B, MO-2095245]. 

= Sporobolus indicus (L.) R. Br. 


Vilfa ciliata var. subfasciculata Doll, Fl. Bras. 2(3): 39. 1878. 
= Sporobolus ciliatus var. subfasciculatus (D6ll) Hack. 


Vilfa clandestina (Biehler) Nees ex Steud., Nomencl. Bot. 
(ed. 2) 2: 767. 1841. Basionym: Agrostis clandestina 
Biehler. 

= Sporobolus clandestinus (Biehler) Hitche. 


236 


Catalogue of New World Grasses: II. 


Vilfa composita (Poir.) P. Beauv., Ess. Agrostogr. 16, 147, 
181. 1812. Basionym: Agrostis composita Poir. 
= Sporobolus compositus (Poir.) Merr. 


Vilfa compressa Trin. ex Spreng., Neue Entdeck. Pflanzenk. 
2: 58. 1821. T-Protol.: North America: America 
boreali. 

NOTE: hom. illeg. 
E. NAME: Vilfa compressa P. Beauv. 
= Muhlenbergia torreyana (Schult.) Hitchc. 


Vilfa confertiflora (A. Rich.) Steud., Syn. Pl. Glumac. 1:. \ 
[1854]. Basionym: Sporobolus confertiflorus A. Rich. 
= Eragrostis japonica (Thunb.) Trin. 


Vilfa confusa E. Fourn., Mexic. Pl. 2: 101. 1886. T-Specim.: 
1 of 7. Mexico: Jalicingo, C..W. Schiede & Deppe 
9/3 [ST: US-998282 (fragm. ex P), US-3376139(split 
from 998282)]. 2 of 7. Mexico: Orizaba, Botteri | 17 
[ST: P; IST: US (fragm.)]. 3 of 7. Mexico: Orizaba, 
Schaffner 93 [ST: P; IST: USA (fragm. ex P)]. 4 of 7. 
Mexico: Orizaba, Schaffner 125 [ST: P]. 5 of 7. 
Mexico: Nevado de Toluca, Sep, Hahn s.n. [ST: P]. 6 
of 7. USA, Hall & Harbour 643 [ST: P]. 7 of 7. 
Mexico: Jorullo, in devexis arenosis montis ignivomi, 
Bonpland in part {ST}. 

NOTE: cites S. ramulosus Griseb. in sched. part.; 
ramulosa Schlect. in Linn. VI, 37, Kunth hb, part. 

= Muhlenbergia minutissima (Steud.) Swallen. 


Vilfa cruciata (L.) P. Beauv., Ess. Agrostogr. 16, 181. 1812. 
Basionym: Agrostis cruciata L. 
= Chloris cruciata (L.) Sw. 


Vilfa cryptandra (TYorr.) Trin., Mem, Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 69. 1840. Basionym: Agrostis cryptandra 
Torr. 

= Sporobolus cryptandrus (Torr.) A. Gray. 


Vilfa cuspidata Torr. ex Hook., Fl. Bor.-Amer. 2: 238. 1840. 
T-Protol.: Canada: banks of the Saskatchawan near the 
Rocky Mts., Drummond s.n. T-Specim.: | of 1. [HT: 
K; IT: NY]. 

= Muhlenbergia cuspidata (Torr. ex Hook.) Rydb. 


Vilfa depauperata Torr. ex Hook., Fl. Bor.-Amer. 2: 257, t. 
236. 1840. T-Protol.: Northwest America, barren 
sandy parts of the Columbia from Menzies' island 
[now Haydens's Isl.] upwards, Douglas s.n. T-Specim.: 
1 of 1. USA, Douglas s.n. (HT: K; IT: GH, NY- 
431744, US (fragm. ex K)]. 

= Muhlenbergia richardsonis (Trin.) Rydb. 


Vilfa depauperata var. filiformis Thurb. ex S. Watson, 
Botany Fortieth Parallel 376. 1871. T-Specim.: | of 3. 
USA: Utah: Vinta Mts., Aug 1869, S. Watson 1281 
[ST: US-83989]. 2 of 3. USA: California: Yosemite 
Valley, 1935, H.N. Bolander 6091 [LT: NY; ILT: GH, 
MO, NY, US-556881, US-344165], LT designated by 
Hitchcock, N. Amer. Fl. 17: xxx. 1935. 3 of 3. USA: 
California: Donner Lk, Torrey 565 [ST]. 

= Muhlenbergia filiformis (Thurb. ex S. Watson) Rydb. 


Vilfa diandra (Retz.) Trin., Gram. Unifl. Sesquifl. 154. 1824. 
Basionym: Agrostis diandra Retz. 
= Sporobolus diandrus (Retz.) P. Beauv. 


Vilfa domingensis Trin., Neue Entdeck. Pflanzenk. 2: 59. 
1821. T-Protol.: Hab. in Domingo. T-Specim.: | of 2. 


In S. Domingo, Cl. Poiteau [HT: LE-TRIN-1694.01 
(& fig.); IT: US-91124]. 2 of 2. Santo Domingo, Anon. 
[T: LE}. 

= Sporobolus domingensis (Trin.) Kunth. 


Vilfa elongata P. Beauv., Ess. Agrostogr. 16, 147, 181. 1812. 
NOTE: nom. nov., nom. illeg. superfl. 
REPLACED NAME: Agrostis elongata Lam. 
CORRECT NAME: Agrostis indica L. 
= Sporobolus indicus (L.) R. Br. 


Vilfa erosa Trin., Mem. Acad. Imp. Sci. Saint-Petersbourg, 
Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 6,4(1-2): 86. 
1840. T-Specim.: | of 1. Sri Lanka, Koenig s.n. [LT: 
LE-TRIN-1698.01 (photo, L, BRI); IT: BM (photo, 
BRI], LT designated by Baaijens & Veldkamp, 
Blume 35: 434 (1991). 

= Sporobolus diandrus (Retz.) P. Beauy. 


Vilfa exilis Trin., Mem. Acad. Imp. Sci. Saint-Petersbourg, 
Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 6,4(1-2): 89. 
1840. T-Protol.: Mexico: Veracruz: Jalapa, Schiede 
s.n. T-Specim.: | of |. Mexico: Jalapa, Aug 28,C./W. 
Schiede s.n. (HT: LE-TRIN-1699.03 (& fig.)]. 

= Sporobolus indicus (L.) R. Br. 


Vilfa eximia Nees ex Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 77. 1840. T-Protol.: Brasilia: Sto Paolo, 
Sellow s.n. T-Specim.: | of 2. Brasilia: St. Paulo, Mar 
1826, Riedel 266 [OM: LE-TRIN-1700.02 (2 sheets)], 
Trinius (Vilfa elatior Nees ). 2 of 2. Brazil: Sao Paulo, 
Sello s.n. [HT: LE-TRIN-1700.01; IT: B, BAA-3469 
(fragm.)]. 

= Sporobolus aeneus (Trin.) Kunth. 


Vilfa familiaris Steud. ex Lechler, Berberid. Amer. Austr. 56. 
1857. T-Protol.: Peru, Lechler 1728. 
NOTE: nom. nud. 
= Sporobolus indicus (L.) R. Br. 


Vilfa filiculmis Thurb. ex Vasey, Descr. Cat. Grass. U.S. 44. 
1885. T-Protol.: USA: New Mexico: Plaza Larga, 
Bigelow 778. 

NOTE: nom. inval., as syn. of Sporobolus filiculmis Vasey. 
= Muhlenbergia thurberi (Scribn.) Rydb. 


Vilfa_filiculmis Thurb. ex Scribn., Bull. Div. Agrostol., 
U.S.D.A. 11: 48. 1898. 
NOTE: nom. inval., as syn. of Sporobolus thurberi Scribner. 
E. NAME: Vilfa filiculmis Thurb. ex Vasey. 
= Muhlenbergia thurberi (Scribn.) Rydb. 


Vilfa fulvescens Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 76. 1840, T-Protol.: America borealis?, Bosc, 
in Willd. herb. 1750,"Agrostis thyrsoides" herb. name. 
T-Specim.: | of 1. [HT: B-Willd.-1750; IT: LE-TRIN- 
1703.01 (frag. ex B-Willd.-1750)], may be the same 
type coll. as Agrostis compressa Poir. (hom. illeg.) = 
Axonopus poiretii Roem. & Schult. (1817) [fide 
Veldkamp, Taxon 39: 327-328 (1990)]. 

= Sporobolus junceus (P. Beauv.) Kunth. 


Vilfa funckii Steud., Syn. Pl. Glumac. 1: 159. 1854. T-Protol.: 
Venezuela: Caripe, hb. Funck 229. 
= Sporobolus funckii (Steud.) F.M. Knuth. 


Vilfa gracilis Trin., Mem. Acad. Imp. Sci. Saint-Petersbourg, 
Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 6,4(1-2): 74. 


Catalogue of New World Grasses: II. 


237 


1840. T-Protol.: USA: Carolina. T-Specim.: 1 of 1. 
Carolina, Zimmermann misit 1836 (HT: LE-TRIN- 
1705.01 (& fig.)]. 

NOTE: hom. illeg., lecta Trinius, p. 133 (1840). 

E. NAME: Vilfa gracilis Trin. 

CORRECT NAME: Vilfa subsetacea Trin. 

= Sporobolus junceus (P. Beauv.) Kunth. 


Vilfa gracilis Trin., Mem. Acad. Imp. Sci. Saint-Petersbourg, 
Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 6,4(1-2): 104. 
1840. T-Protol.: North America: Amer. boreal "?", 
Hooker [misit]. T-Specim.: 1 of 1. [HT: LE-TRIN- 
1704.01]. 
NOTE: non. Vilfa gracilis Trin. p. 74 (1940). 
= Muhlenbergia cuspidata (Torr. ex Hook.) Rydb. 


Vilfa gracillima Thurb. ex S. Watson, Bot. California 2: 268. 
1880. T-Specim.: | of 2. USA: California, 1866, H.N. 
Bolander 6090 [ST: US-556882]. 2 of 2. USA: 
California: High Sierra, 1864, Brewer 2827 [LT: MO; 
ILT: NY], LT designated by Peterson & Annable, 
Syst. Bot. Monogr. 31: 44 (1991). 

= Muhlenbergia filiformis (Thurb. ex S. Watson) Rydb. 


Vilfa grisebachiana E. Fourn., Mexic. Pl. 2: 98. 1886. T- 
Specim.: | of 2. Cuba, Wright 3427a [ST]. 2 of 2. 
Mexico: Orizaba, Bourgeau 32 [ST]. 

NOTE: cites "Sporobolus purpurascens Griseb. part.". 

= Sporobolus purpurascens (Sw.) Ham. 


Vilfa heterolepis A. Gray, Ann. Lyceum Nat. Hist. New York 
3: 233. 1835. T-Specim.: | of 3. USA: New York, 
Watertown, Dr. Crawe [HT: NY], right hand plant on 
sheet of 3 collections. 2 of 3. USA: Connecticut, New 
Haven, J.D. Dana [PT: NY]. 3 of 3. Canada: Quebec, 
Montreal, Dr. A. Holms [PT: NY]. 

= Sporobolus heterolepis (A. Gray) A. Gray. 


Vilfa indica (L.) Trin. ex Steud., Nomencl. Bot. (ed. 2) 2: 
767. 1841. Basionym: Agrostis indica L. 
= Sporobolus indicus (L.) R. Br. 


Vilfa intermedia Trin., Gram. Unifl. Sesquifl. 156. 1824. T- 
Protol.: West Indies: Ind. occ. [error, corrected in 1840 
to Mauritan. Sieber]. T-Specim.: | of 1. Mauritius, 
Sieber 11-38 [HT: LE-TRIN-1710.01; IT: K, L]. 

= Sporobolus virginicus (L.) Kunth. 


Vilfa jacquemontii (Kunth) Trin., Mem. Acad. Imp. Sci. 
Saint-Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. 
Nat. 6,4(1-2): 92. 1840. Basionym: Sporobolus 
jacquemontii Kunth. 

= Sporobolus jacquemontii Kunth. 


‘ilfa juncea (P. Beauv.) Trin., Gram. Unifl. Sesquifl. 157, 
1824. Basionym: Heleochloa juncea P. Beauv. 
= Sporobolus junceus (P. Beauv.) Kunth. 


Vilfa lateriflora (Michx.) P. Beauv., Ess. Agrostogr. 16, 147, 
181. 1812. Basionym: Agrostis lateriflora Michx. 
NOTE: appears erroneously as "laterifolia" on p. 16 and 147 

but correctly on p. 181. 
= Muhlenbergia frondosa (Poir.) Fernald. 


Vilfa liebmanni E. Fourn., Mexic. PI. 2: 100. 1886. T-Protol.: 
Mexico: "Absque loco.", Liebmann 693. 
= Sporobolus purpurascens (Sw.) Ham. 


‘ilfa littoralis (Lam.) P. Beauv., Ess. Agrostogr. 16, 147, 
181. 1812. Basionym: Agrostis littoralis Lam. 


= Sporobolus virginicus (L.) Kunth. 


Vilfa luxurians Steud. ex Lechler, Berberid. Amer. Austr. 56. 
1857. T-Protol.: Peru, Lechler 1544. 
NOTE: nom. nud. 
= Sporobolus virginicus (L.) Kunth. 


Vilfa mangalorica Hochst. ex Steud., Syn. Pl. Glumac. 1: 
159. 1854. Basionym: <>. 
= Sporobolus tenuissimus (Mart. ex Schrank) Kuntze. 


Vilfa matrella Nees, Fl. Bras. Enum. Pl. 2: 400. 1841. T- 
Protol.: Herb. Maurit., Sieber 38. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Vilfa intermedia Trin. 
= Sporobolus virginicus (L.) Kunth. 


Vilfa mexicana (L.) P. Beauv., Ess. Agrostogr. 16, 148, 181. 
1812. Basionym: Agrostis mexicana L. 
= Muhlenbergia mexicana (L.) Trin. 


Vilfa minutiflora Trin., Gram. Unifl. Sesquifl. 158. 1824. T- 
Protol.: cult. T-Specim.: | of |. cultivated in Leningrad 
from seeds collected in Brazil, Martius s.n. (HT: LE- 
TRIN-1718.01; IT: MJ], merotypic with Sporobolus 
tenuissimus and S. minutiflorum. 

NOTE: type grown from same seed source as Panicum 
tenuissium Mart. ex Schrank (1822). 

CORRECT NAME: Sporobolus tenuissimus (Mart. ex 
Schrank) Kuntze. 

= Sporobolus tenuissimus (Mart. ex Schrank) Kuntze. 


Vilfa muelleri E. Fourn., Mexic. Pl. 2: 98. 1886. T-Specim.: 1 
of 2. Mexico: Orizaba, Mueller 2/17 [LT: US], LT 
designated by Hitchcock, N. Amer. FI. 17: 490 (1937). 
2 of 2. Mexico: Mirador, Liebmann 689 [ST]. 

= Sporobolus purpurascens (Sw.) Ham. 


Vilfa pilifera Trin., Gram. Unifl. Sesquifl. 157. 1824. T- 
Protol.: V. sp. Nepal. T-Specim.: 1 of 2. Nepal, 
Wallich 3774 {LT: LE; ILT: BM, Kk, US], LT 
designated by Baaijens and Veldkamp, Blumea 35: 
441 (1991). 2 of 2. Nepal, a cl. Rudge il. com. a 
Lindley [HT?: LE-TRIN-1723.01 parte Nepal]. 

= Sporobolus piliferus (Trin.) Kunth. 


Vilfa plumbea Trin., Mem. Acad. Imp. Sci.  Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 98. 1840. T-Protol.: Mexico: Min. del Monte, 
Schlechtendal s.n. T-Specim.: | of 1. Mexico: Mineral 
del monte, Schlechtendal ex Schiede s.n. (HT: LE- 
TRIN-1724.01; IT; US-557435 (fragm. ex LE)], fig. 

= Muhlenbergia plumbea (Trin.) Hitche. 


Vilfa pubescens E. Fourn., Mexic. Pl. 2: 102. 1886. T-Protol.: 
Mexico: inter Michiguana et Tanquecillos, Canyon de 
las Minas et Victoria, Karwinsky 1012. T-Specim.: 1 
of 1. [HT: P; IT: US (fragm. ex P)]. 
= Muhlenbergia villiflora Hitchc. 


Vilfa purpurascens (Sw.) P. Beauv., Ess. Agrostogr. 16, 182. 
1812. Basionym: Agrostis purpurascens Sw. 
= Sporobolus purpurascens (Sw.) Ham. 


Vilfa racemosa (Michx.) P. Beauv., Ess. Agrostogr. 16, 148, 
182. 1812. Basionym: Agrostis racemosa Michx. 
= Muhlenbergia racemosa (Michx.) Britton, Sterns & 
Poggenb. 


238 


Catalogue of New World Grasses: II. 


Vilfa ramulosa Kunth, Nov. Gen. Sp. 1: 137. 1815 [1816]. T- 
Protol.: Crescit in devexis arenosis montis ignivomi 
Mexicani, Jorullo, alt. 460 hexap. Floret Septembri., 
Humboldt & Bonpland s.n. T-Specim.: | of 1. Mexico: 
Jorullo, Humboldt & Bonpland s.n. [LT: P-Bonpl. (the 
middle plant on the sheet); ILT: B-W, BM, LE-TRIN- 
1730.01, US-91131 (fragm.)], LT designated by 
Peterson & Annable, Syst. Bot. Monogr. 31: 77 
(1991). 

= Muhlenbergia ramulosa (Kunth) Swallen. 


Vilfa repens (J. Presl) Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 102. 1840. Basionym: Sporobolus repens J. 
Presl. 

= Muhlenbergia repens (J. Presl) Hitchce. 


Vilfa retzii Steud., Nomencl. Bot. (ed. 2) 1: 40. 1840. 
NOTE: nom. illeg. superfl. 
CORRECT NAME: Agrostis diandra Retz. 
= Sporobolus diandrus (Retz.) P. Beauv. 


Vilfa richardsonis Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 103. 1840. T-Protol.: Amer. — boreal, 
Richardson s.n. T-Specim.: | of 1. North America, 
Richardson s.n. [HT: LE; IT: US (fragm. ex LE)]. 

= Muhlenbergia richardsonis (Trin.) Rydb. 


Vilfa riehlii Steud., Syn. Pl. Glumac. 1: 154. 1854. T-Protol.: 
Am. sptr. T-Specim.: | of 1. USA: Missouri: St. Louis, 
bord des chemins, no date, . Rieh/ s.n. [HT: P; US- 
A865650 (fragm. ex P)]. 

= Sporobolus vaginiflorus (Torr. ex A. Gray) Alph. Wood. 


Vilfa rigens Trin., Sp. Gram, 3: t. 250. 1836 [1829-1830]. T- 
Protol.: Chile: [Mendoza], /Gillies/. T-Specim.: | of 1. 
[HT: LE-TRIN-1734.02]. 
= Sporobolus rigens (Trin.) Desv. 


Vilfa rigida Buckley, Proc. Acad. Nat. Sci. Philadelphia 14: 
89. 1862. T-Protol.: USA: Oregon?, Anymous_ s.n. 
[HT: PH], "in the herbarium of the academy without 
label". T-Specim.: 1 of 1. USA: Oregon: in the 
herbarium of the academy without label, Nuttall s.n. 
[HT: PH], According to Manual of the Grasses of the 
Unites States: 844 (1950): “the locality probably 
erroneous". 

= Calamovilfa longifolia (Hook.) Hack. ex Scribn. 


Vilfa rupestris Trin., Gram. Panic. 22. 1826. T-Protol.: V. sp. 
Brasilia. T-Specim.: | of |. In rupium fissus_ pr. 
Itamaratim Brasiliae, mensu Febr., Langsdorff (HT: 
LE-TRIN-1746.01 (fig.)]. 

= Sporobolus indicus (L.) R. Br. 


Vilfa rupestris (Kunth) Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 66. 1840. Basionym: Sporobolus rupestris 
Kunth. 

NOTE: hom. illeg. 

E. NAME: Vilfa rupestris Trin. 
= Sporobolus piliferus (Trin.) Kunth. 


Vilfa sabeana Buckley, Proc. Acad. Nat. Sci. Philadelphia 14: 
90. 1862. T-Protol.: USA: Texas: San Saba Co. T- 
Specim.: | of 1. USA: Texas: San Saba Co, Buckley 
sn. [LT: PH], LT designated by Hitchcock, Man. 


Grass. U.S. 957 (1935), but without citing a specific 
sheet in a specific herbarium. 

NOTE: as Vilfa “Sabeana". 

= Sporobolus pyramidatus (Lam.) Hitche. 


Vilfa sacatilla E. Fourn., Mexic. Pl. 2: 101. 1886. T-Specim.: 
1 of 3. Mexico: Chapultepec, Schaffner 165 [ST: W; 
IST: US (fragm. ex W)]. 2 of 3. Mexico: San Luis de 
Potosi, Virlet 1455 [ST: P; IST: US (fragm. ex P)]. 3 
of 3. USA: Texas, Wright s.n. [ST: P; IST: US (fragm. 
ex P)]. 

NOTE: nom. illeg. superfl.; cites Sporobolus sacatilla 

Griseb. in Sched. Agrostis brevifolia Nutt. 
= Muhlenbergia utilis (Torr.) Hitche. 


Vilfa schiedeana Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 73. 1840. T-Protol.: USA: Arkansas, 
"Schiede” [error for Beyrich according to Ruprecht]. 
T-Specim.: 1 of 1. [HT: LE-TRIN-1738.01], Type 
specimen annotated by Ruprecht “Beyrich non 
Schiede" [fide Index of Grass Species]. 

= Sporobolus junceus (P. Beauv.) Kunth. 


Vilfa serotina Trin., Sp. Gram. 3: t. 251. 1829-30. T-Protol.: 
USA: America boreal, Nuttall, Agrostis serotina Nutt. 
ined. 

= Muhlenbergia uniflora (Muhl.) Fernald. 


Vilfa serotina Torr. ex A. Gray, N. Amer. Gram. 1: 2. 1834. 
NOTE: hom. illeg.; same species as Vilfa serotina Trin., but 
different type, Gray's name proposed as a new name 
for Agrostis serotina Torr. 
E. NAME: Vilfa serotina Trin. 
CORRECT NAME: Sporobolus serotinus A. Gray. 
= Muhlenbergia uniflora (Muhl.) Fernald. 


Vilfa spartinae Trin, Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 82. 1840. T-Protol.: USA: Texas, misit 
Hooker. T-Specim.: | of 1. USA: Texas, Hooker s.n. 
[HT: LE-TRIN-1742.01]. 

= Spartina spartinae (Trin.) Merr. ex Hitche. 


Vilfa squarrosa Trin, Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 
6,4(1-2): 100. 1840. T-Protol.: USA: Columbia River: 
Ins. Menzies [now Hayden's Island], /Douglas/ misit 
Hooker. T-Specim.: 1 of 1. USA: Washington: 
Menzies Island, Columbia River, Douglas s.n. [HT: 
LE; IT: US (fragm. ex LE)]. 

NOTE: nom. illeg. superfl.; November 1840, same as Vilfa 
depauperata Torr. Jul 1840. 

CORRECT NAME: Vilfa depauperata Torr. ex Hook. 

= Muhlenbergia richardsonis (Trin.) Rydb. 


Vilfa stachydantha (A. Rich.) Steud., Syn. Pl. Glumac. 1: 
156. 1854. Basionym: Sporobolus stachydanthus A. 
Rich. 

= Sporobolus piliferus (Trin.) Kunth. 


Vilfa subsetacea Trin., Mem. Acad. Imp. Sci. Saint- 
Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci, Nat. 
6,4(1-2): 133. 1840. 

NOTE: nom. nov., in nota. 

REPLACED NAME: Vilfa gracilis Trin. 
BLOCKING NAME: Vilfa gracilis Trin. 
= Sporobolus junceus (P. Beauv.) Kunth. 


Catalogue of New World Grasses: II. 


239 


Vilfa tenacissima (L. f.) Kunth, Nov. Gen. Sp. 1: 138. 1815 
[1816]. Basionym: Agrostis tenacissima L. f. 
= Sporobolus indicus (L.) R. Br. 


Vilfa tenacissima var. exilis (Trin.) E. Fourn., Mexic. PI. 2: 
99. 1886. Basionym: Vilfa exilis Trin. 
= Sporobolus indicus (L.) R. Br. 


Vilfa tenacissima var. fuscicola Hook., Fl. Bor.-Amer. 2: 239. 
1839. T-Protol.. USA: Columbia River: Menzies Isl. 
[now Haydens Isl.], Douglas. 
= Sporobolus cryptandrus (Torr.) A. Gray. 


Vilfa tenera Trin., Mem. Acad. Imp. Sci. Saint-Petersbourg, 
Ser. 6, Sci. Math., Seconde Pt. Sci. Nat. 6,4(1-2): 87. 
1840. T-Protol.: USA: Boston, Boott, in hb. Kunth. 

= Muhlenbergia uniflora (Muhl.) Fernald. 


Vilfa tremula Trin., Gram. Unifl. Sesquifl. 155. 1824. 
NOTE: nom. illeg. superfl., cites Agrostis tremula Willd., 
nom. illeg. superfl. 
CORRECT NAME: Agrostis juncea Lam. 
= Sporobolus virginicus (L.) Kunth. 


Vilfa tricholepis Torr., Pacif. Railr. Rep. 4(5): 155. 1857. T- 
Protol.; USA: New Mexico: Sandia Mts., Bigelow s.n. 
T-Specim.: | of 1. USA: New Mexico: Sandia 
Mountains, 10 Oct 1853, Bigelow s.n. [LT: NY; ILT: 
MO}, LT designated by ?. 

= Blepharoneuron tricholepis (Torr.) Nash. 


Vilfa triniana Steud., Syn. Pl. Glumac. 1: 156. 1854. T- 
Protol.: Canada: British Columbia: "Columbia a 
Canadensi". 

= Sporobolus eryptandrus (Torr.) A. Gray. 


Vilfa utilis Torr., Pacif. Railr. Rep. 5(2): 365. 1858. T-Protol.: 
USA: California: between the Tejon Pass and the Lost 
Hills. T-Specim.: 1 of 1. USA: California: Lost 
Mountain Spring, trip from Tejon to the Lost Hills, in 
stony places, WP. Blake s.n. [HT: NY; IT: GH, MO, 
US (fragm. ex NY)]. 
= Muhlenbergia utilis (Torr.) Hitche. 


Vilfa vaginiflora Torr. ex A. Gray, N. Amer. Gram. 1: no. 3. 
1834. T-Protol.: USA: New Jersey. 
NOTE: same by Trinius in 1840. 
= Sporobolus vaginiflorus (Torr. ex A. Gray) Alph. Wood. 


Vilfa vinzenti Steud., Syn. Pl. Glumac. 1: 155. 1854. T- 
Protol.: USA: Texas: "Agrostis nova species", Vinzent 
hb. 62 [HT: Vinzent hb.]. T-Specim.: | of 1. [HT: P?}. 
= Sporobolus junceus (P. Beauv.) Kunth. 


Vilfa virginica (L.) P. Beauv., Ess. Agrostogr. 16, 149, 182. 
1812. Basionym: Agrostis virginica L. 
= Sporobolus virginicus (L.) Kunth. 


Vilfagrostis Déll, Fl. Bras. 2(3): 137. 1878. 
NOTE: nom. inval., as syn. of Eragrostis Wolf. 
= Eragrostis Wolf. 


Willbleibia Herter, Revista Sudamer. Bot. 10: 132. 1953. 
TYPE: Willbleibia texana (Hitchc.) Herter. 
NOTE: nom. illeg. superfl., Willkommia Hack. thought to be 
an illeg. hom., but not. 
= Willkommia Hack. 


Willbleibia stolonifera (Parodi) Herter, Revista Sudamer. 
Bot. 9: 85. 1952. Basionym: Willkommia texana var. 
stolonifera Parodi. 


= Willkommia texana var. stolonifera Parodi. 


Willbleibia texana (Hitche.) Herter, Revista Sudamer. Bot. 9: 
132. 1952. Basionym: Willkommia texana Hitche. 
= Willkommia texana Hitche. 


Willkommia Hack., Verh. Bot. Vereins Prov. 
Brandenburg 30: 145. 1888. TYPE: Willkommia 
sarmentosa Hack. 

TRTD. BY: Peterson, P. M. 
NHT: Cynodonteae. NO. ACCPTD. SPP.: | SUBSP.: 2. 
SYN.: Willbleibia Herter. 


Willkommia texana Hitchc., Bot. Gaz. 35: 283, f. 1-2. 1903. 
T-Protol.: USA: Texas: Ennis and Beeville,./.G. Smith 
s.n. [HT: US]. T-Specim.: | of 1. USA: Texas: Ennis, 
21 Jul 1897, 4G. Smith s.n. [HT: US-81733; IT: US- 
879624]. 
DST.: Arge, USA. 
LVL. ACCPTC.: 1. 
ALSO ACCPTD. BY: Anonymous (1986); Correll, D. S. & 
M. C. Johnston (1970); Hitchcock, A. S. (1951). 
SYN.: Craspedorhachis texana (Hitche.) Pilg.; Willbleibia 
texana (Hitchc.) Herter. 


Willkommia texana var. stolonifera Parodi, Physis (Buenos 
Aires) 8: 78. 1925. T-Protol.: Argentina: Buenos 
Aires: Pipinas (F.C.S.), Habita tierras calcareas, 25 
Mar 1923, L. Parodi 4975 [HT: SI; IT: US-1024458]. 
T-Specim.: | of 1. Argentina: Buenos Aires: Pipinas, 
25 Mar 1923, L. Parodi 4975 [HT: SI; IT: US- 
1024458]. 

DST.: Arge. 

LVL. ACCPTC:: |. 

ALSO ACCPTD. BY: Cabrera, A. L. (1970). 

SYN.: Craspedorhachis texana var. stolonifera (Parodi) 
Pilg.; Willbleibia stolonifera (Parodi) Herter. 


Willkommia texana Hitchce. var. texana. 
DST.: USA. 
LVL. ACCPTC.: 1. 
SYN.: Craspedorhachis texana var. texana. 


Windsoria Nutt., Gen. N. Amer. PI. 1: 70. 1818. 
= Tridens Roem. & Schult. 


Windsoria ambigua (Elliott) Nutt., Gen. N. Amer. Pl. 1: 70. 
1818. Basionym: Poa ambigua Elliott. 
= Tridens ambiguus (Elliott) Schult. 


Windsoria poiformis Nutt., Gen. N. Amer. Pl. 1: 70. 1818. 
NOTE: nom. nov., nom. illeg. superfl. 
REPLACED NAME: Poa sesleroides Michx. 
BLOCKING NAME: Poa sesleroides All. 
CORRECT NAME: Poa quinquefida Pursh, 
= Tridens flavus (L.) Hitchce. 


Windsoria sesleroides (Michx.) Eaton, Man. Bot. (ed. 5) 447. 
1829. Basionym: Poa sesleroidesMichx. 
= Tridens flavus (L.) Hitche. 


Windsoria stricta Nutt., Trans. Amer. Philos. Soc., n.s., 5: 
147. 1835. T-Protol.: Arkansas Territory: Hab. in 
prairies, near the town of Arkansas, syntypes. T- 
Specim.: 1 of 2. USA: Prairies, near the town of 
Arkansas., Nuttall s.n.? [IST: US (fragm. ex BM)]. 2 
of 2. , Wright 778 [ST: PH]. 

= Tridens strictus (Nutt.) Nash. 


240 


Catalogue of New World Grasses: II. 


Zoysia Willd., Ges. Naturf. Freunde Berlin Neue 
Schriften 3: 440. 1801. TYPE: Zoysia pungens Willd. 
NOTE: nom. cons. 
TRTD. BY: Peterson, P. M. 
NHT: Zoysiinae. NO. ACCPTD. SPP.: 3 SUBSP.: 2. 
SYN.: Brousemichea Bal.; Matrella Pers.; Osterdamia 
Neck. ex Kuntze. 


Zoysia aristata Mill. Hal., Bot. Zeitung (Berlin) 13(16): 272. 


1855. T-Specim.: | of 2. Insula Java: in arenosis 
maritimis prope Gradjakam_ prov. Banjuwangie, 
Zollinger 2890 [ST: B, destroyed; IST: BM, P]. 2 of 2. 
Java, Weltevreden, Hasskarl s.n. (ST: B, destoyed]. 

= Zoysia matrella (L.) Merr. 


Zoysia griffithiana Mill. Hal., Bot. Zeitung (Berlin) 13(16): 
273. 1855. T-Protol.: Serampore Indiae orientalis, 
Griffith s.n. T-Specim.: | of 1. Serampore, Griffith s.n. 
[HT: B, destroyed; IT: K]. 

NOTE: as Zoysia "Griffithiana”. 
= Zoysia matrella (L.) Merr. 


Zoysia japonica Steud., Syn. Pl. Glumac. 1: 414. 1854. 

DST.: Braz, USA. 

LVL. ACCPTC.: 1; cultivated, introduced. 

ALSO ACCPTD. BY: Bor, N. L. (1960); Filgueiras, T. S. 
(1991); Gleason, H. A. & A. Cronquist (1991); 
Hitchcock, A. S. (1951); Wunderlin, R. P. (1998). 

SYN.: Osterdamia japonica (Steud.) Hitche.; Zoysia 
pungens var. japonica (Steud.) Hack. 


Zoysia malaccensis Gand., Bull. Soc. Bot. France 66(7): 303. 
1919 [1920]. T-Protol.: Asia merid., Singapore, Ridley 
s.n. T-Specim.: | of 1. Singapore, Ridley s.n. [HT: LY: 
IT: 1445654]. 
= Zoysia matrella (L.) Merr. 


Zoysia matrella (L.) Merr., Philipp. J. Sci. 7(4): 230. 1912. 
Basionym: Agrostis matrella L. 

NOTE: as "Zoisia". 

DST.: Braz, Cari, CoRi, ElSa, Hond, Mexi, Nica, USA. 

LVL. ACCPTC.: |; cultivated, introduced. 

ALSO ACCPTD. BY: Bor, N. L. (1960); Catasus Guerra, L. 
(1997); Davidse, G., M. Sousa S. & A. O. Chater 
(editores generales) (1994); Espejo Serna, A., A. R. 
Lopez-Ferrari, and J.  Valdes-Reyna (2000); 
Goudswaard, P. C. (1980); Hitchcock, A. S. (1936); 
Hitchcock, A. S. (1951); McVaugh, R. (1983); Pohl, 
R. W. & G. Davidse (1994); Renvoize, S. A. (1988); 
Smith, L. B., D. C. Wasshausen & R. M. Klein 
(1981); Wunderlin, R. P. (1998). 

Agrostis matrella L.; Matrella juncea  Pers.; 
Osterdamia matrella (.) Kuntze; Osterdamia zoysia 
Honda; Osterdamia zoysia var. tenuifolia (Willd. ex 
Thiele) Honda; Osterdamia zoysia var. typica Honda; 
Zoysia aristata Mill. Hal.; Zoysia griffithiana Mill. 
Hal.; Zoysia malaccensis Gand.; Zoysia matrella var. 
typica Sasaki; Zoysia pungens Willd. 


SYN: 


Zoysia matrella (L.) Merr. var. matrella. 

DST.: Braz, Cari, CoRi, Nica, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994): Goudswaard, P. 
C. (1980); Liogier, H. & L. Martorell (1982); Pohl, 
R. W. & G. Davidse (1994); Smith, L. B., D. C. 
Wasshausen & R. M. Klein (1981); Wunderlin, R. P. 
(1998). 


Zoysia matrella var. pacifica Goudswaard, Blumea 26: 172, 
174, Map 1. 1980. T-Protol.: Ryukyu Islands: 
Yonakuni Island, Hatusima 24127 (HT: L;1T: FU. 

DST.: CoRi, ElSa, Hond, Nica. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Davidse, G., M. Sousa S. & A. O. 
Chater (editores generales) (1994); Goudswaard, P. 
C. (1980); Pohl, R. W. & G. Davidse (1994), 

SYN.: Zoysia pacifica (Goudswaard) M. Hotta & Kuroki; 
Zoysia tenuifolia Willd. ex Trin. 


Zoysia matrella subsp. tenuifolia (Willd. ex Thiele) T. 
Koyama, Grass. Jap. Neighb. Reg. 534. 1987. 
Basionym: Zoysia tenuifolia Willd. ex Thiele. 

= Zoysia tenuifolia Willd. ex Thiele. 


Zoysia matrella var. tenuifolia (Willd. ex Thiele) Sasaki, List 
PI. Formosa 80. 1928. Basionym: Zoysia tenuifolia 
Willd. ex Thiele. 

= Zoysia tenuifolia Willd. ex Thiele. 


Zoysia matrella var. typica Sasaki, List Pl. Formosa 80. 1928. 
NOTE: nom. inval., see ICBN Art. 24.3. 
= Zoysia matrella (L.) Merr. 


Zoysia pacifica (Goudswaard) M. Hotta & Kuroki, Acta 
Phytotax. Geobot. 45(1): 71-72, f. IA. 1994, 
Basionym: Zoysia matrella var. pacifica Goudswaard. 

= Zoysia matrella var. pacifica Goudswaard. 


Zoysia pungens Willd., Ges. Naturf. Freunde Berlin Neue 
Schriften 3: 441. 1801. T-Protol.: Klein s.n. [B-W]. T- 
Specim.: | of 1. India Orientali [T: IT; US-91043 
(fragm.)]. 

= Zoysia matrella (L.) Merr. 


Zoysia pungens var. Japonica (Steud.) Hack., Bull. Herb. 
Boissier 7(9): 642. 1899, Basionym: Zoysia japonica 
Steud. 

= Zoysia japonica Steud. 


Zoysia pungens var. tenuifolia (Willd. ex Thiele) T. Durand 
& Schinz, Consp. Fl. Afr. 5: 734. 1894. Basionym: 
Zoysia tenuifolia Willd. ex Thiele. 

= Zoysia tenuifolia Willd. ex Thiele. 


Zoysia tenuifolia Willd. ex Thiele, Linnaea 9(3): 309. 1835. 
T-Protol.: Sieber Maur. Exsicc. Il 40 [LT: B 
(Destroyed); ILT: BM, L], LT cited by P.C. 
Goudswaard in Blumea 26: 171 (1980). T-Specim.: | 
of 1. Sieber Maurit. II, no. 40 [ILT: MO]. 

DST.: Ecua, USA. 

LVL. ACCPTC.: 1; introduced. 

ALSO ACCPTD. BY: Dodson, C. H., A. H. Gentry & F. M. 
Valverde (1985); Jorgensen, P. M. & S. Leon-Yanez 
(eds.) (1999); Pohl, R. W. (1980); Tovar, O. (1993). 

SYN.: Osterdamia tenuifolia (Willd. ex Thiele) Kuntze; 
Zoysia matrella subsp. tenuifolia (Willd. ex Thiele) 
T. Koyama; Zoysia matrella var. tenuifolia (Willd. ex 
Thiele) Sasaki; Zoysia pungens var. tenuifolia (Willd. 
ex Thiele) T. Durand & Schinz. 


Zoysia tenuifolia Willd. ex Trin., Mem. Acad. Imp. Sci. 
Saint-Petersbourg, Ser. 6, Sci. Math., Seconde Pt. Sci. 
Nat. 4,2(1): 96. 1836. T-Protol.: Mascarene Islands. T- 
Specim.: 1 of 1. [IT: LE-TRIN 0397.01]. 
NOTE: hom. illeg. 
E. NAME: Zoysia tenuifolia Willd. ex Thiele. 
= Zoysia matrella var. pacifica Goudswaard. 


Catalogue of New World Grasses: II. 241 


Zoysieae trib. Benth., J. Linn. Soc., Bot. 19: 29. 1881. NHT: Cynodonteae. 
CNSLTD. LIT.: Clayton, W. D. & F. R. Richardson, 
(1973). 


Zoysiinae subtrib. Benth., Fl. Austral. 7: 453. 1878. 
TRTD. BY: Soreng, R. J., G. Davidse, T. S. Filgueiras, E. J. 
Judziewicz, P. M. Peterson, and F. O. Zuloaga. 


242 


Catalogue of New World Grasses: II. 


BIBLIOGRAPHY 


Abrams, L. 1923. Ferns to Birthworts. ///ustrated 
Flora of the Pacific States. 1: 1-557. Stanford 
University Press, Stanford. 


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