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OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


VOLUME 1. SECTION D 


Edited by 
FRANCIS HEMMING, c.M.G., C.B.E. 


Secretary to the International Commission on Zoological Nomenclature 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956—1957 


(All rights reserved) 


» INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
DIRECTIONS PUBLISHED IN THE PRESENT SECTION OF 
VOLUME 1 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JorRDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President : Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) 
(12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary: Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(Arranged in order of precedence by reference to date of election or of most recent re-election, 
as prescribed by the International Congress of Zoology) 


Professor H. Boscuma (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. is (Universitetets Zoologiske Museum, Copenhagen, Denmark) (27th 

y 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (A7th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEy (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEwsKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (Sth July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitat zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMoND (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) (12th August 
1953) (President) 

Professor Harold E. Voxes (University of Tulane, Department of Geology, New Orleans, 
Louisiana, U.S.A.) (12th August 1953 

Professor Béla HANKO (Mezégazdasadgi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y., 
U.S.A.) (12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. HoLtuuts (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (15th October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum v Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KUHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

Professor F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 


Professor Ernst MAyrR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954 

Professor Enrico ToRTONESE (Museo di Storia Naturale ‘‘ G. Doria’’, Genova, Italy) 
(16th December 1954) 


IV 


INTERNATIONAL COMMISSION ON ZOOLOGICAL 
NOMENCLATURE 


(continued) 


C. The Staff of the Secretariat of the Commission 


Honorary Secretary: Myr. Francis Hemming, C.M.G., C.B.E. 


Honorary Personal Assistant to the Secretary: Mrs. M. F. W. 
Hemming 


Honorary Archivist : Mr. Francis J. Griffin, A.L.A. 


Consulting Classical Adviser: Professor The Rev. L. W. 
Grensted, M.A., D.D. 


** Official Lists” Section : Miss D. N. Noakes, B.Sc. 
** Régles’”’ Section: Mrs. A. F. Wilson, M.A. 


Mrs. J. H. Newman 
Miss C. W. Kirton 
Mrs. I. Saltman 
Mrs. J. A. White 


Indexer : Miss Mary Cosh, M.A. 


Secretariat : 


INTERNATIONAL TRUST FOR ZOOLOGICAL 
NOMENCLATURE 


Chairman : The Right Hon. Walter Elliot, C.H., M.C., F.R.S., 
M.P. 


Managing Director and Secretary: Mr. Francis Hemming, 
CMG... CBE, 


Publications Officer : Mrs. C. Rosner 
Trust Duties Officer : Mrs. J. H. Newman 


ADDRESSES OF THE COMMISSION AND THE TRUST 


Secretariat of the Commission: 28 Park Village East, Regent’s 
Park, London, N.W.1. 


Offices of the Trust : 41 Queen’s Gate. London, S.W.7. 


FOREWORD 


The present Section contains a further instalment of Directions 
rendered by the International Commission on Zoological Nomen- 
clature embodying decisions taken in connection with the review 
of the entries made on the Official List of Generic Names in Zoology 
in the period up to the end of 1936 undertaken in pursuance of 
a General Directive issued to it by the Thirteenth International 
Congress of Zoology, Paris, 1948, as extended by a further such 
Directive issued by the Fourteenth International Congress of 
Zoology, Copenhagen, 1953. The immediately preceding 
Section (Section C) of the present volume contained twenty-three 
(23) instalments of the prescribed review—all in the form of 
Directions. In addition, prior to the commencement of the 
above Section, thirty-four (34) instalments had been published 
in other volumes of the Opinions and Declarations Series. Of 
these, thirty-two (32) had been rendered as Opinions! and two (2) 
as Directions*. Accordingly, at the time of the commencement 
of the present Section the total number of instalments already 
published amounted to fifty-seven (57). Thus, when account is 
taken of the eighteen (18) instalments contained in the present 
Section, the total number of instalments of the prescribed review 
so far published is seen to amount to seventy-five (75). 


2. The present volume (Section D of Volume 1) was published 
in 19 Parts. It contains 484 pages (T.P.—XIV, 1—470). It is 


— 


The thirty-two Opinions here referred to are Opinions 136, 149, 158, 180, 192, 
Poe 196. 201: 200,225, 226, 238, 239, 240, 242, 244.259" 260, 261, 262,°271, 
212s 21427821279, 283, 299, 312, 320; 333, 334, 356: 


The two Directions concerned are Directions | and 4, 


nw 


VI 


thus of substantially the same size as the preceding section 
(Section C) of the present volume. 


3. Eighteen Directions are included in the present instalment. 
Of these, three (Directions 40, 42, 52) are concerned exclusively 
with the determination of the gender to be attributed to generic 
names placed on the Official List of Generic Names in Zoology 
up to the end of 1936 ; three (Directions 37, 44, 56) contain Rulings 
on various miscellaneous matters relating to the names of genera 
of particular Classes (Crustacea (Order Decapoda); Aves ; 
Pisces ; Amphibia ; Reptilia) on which decisions were needed 
as a preliminary to the publication of the foregoing Official List 
in book-form ; three Directions (Directions 36, 43, 57) contain 
Rulings placing on the Official List of Specific Names in Zoology 
the specific names of the type species of genera in the Classes 
referred to above, the names of which were placed on the Official 
List of Generic Names in Zoology in the period in question. 
The remaining nine Directions are concerned with individual 
cases, seven with particular names and two (Directions 38, 48) 
with the status of particular books. Jn two of these nine cases 
(Directions 49, 50) the International Commission found it necessary 
to make use of its Plenary Powers. The seven Directions dealing 
with individual names (Directions 39, 45, 47, 49, 50, 51, 55) 
contain Rulings which validate, correct or complete entries made 
on the Official List of Generic Names in Zoology in the period up 
to the end of 1936. 


4. The cases dealing with individual names in the present 
Section, including those relating to the determination of the 
gender of generic names placed on the Official List of Generic 
Names in Zoology in the period up to the end of 1936 amount to 
sixteen ; of these, six contain Rulings relating to the names of 
taxa belonging to more than one Class of the Animal Kingdom, 
including one (Direction 42) which relates to the names of taxa 
belonging to no less than twenty-two Classes. The total number 
of cases involved in Rulings given in the Directions comprised in 
the present Section is thus fifty. The distribution by Classes 
of these cases is shown in the following table :— 


Vil 


TABLE 1 


Distribution of cases by Classes in the Animal Kingdom 


Number of 
Name of Class applications 


Rhizopoda 
Ciliata 
Mastigophora 
Sporozoa 
Neosporidia 
Hydrozoa 
Anthozoa 
Cestoda 
Trematoda 
Nematoda 
Nematomorpha 
Hirudinea 
Chaetopoda 
Crustacea 
Diplopoda 
Trilobita 
Insecta 
Merostomata 
Arachnida 
Gastropoda 
Scaphopoda 
Pelecypoda 
Cephalopoda 
Brachiopoda | 
Asteroidea | 
Holothuroidea 
Crinoidea 
Urochorda 
Cyclostomata 
Pisces 
Amphibia 
Reptilia 

Aves 


Total 


OO WW GO RD me ee ND OO ee RRR RR RE RE Ee 


On 
i=) 


Vill 


5. The three Directions relating exclusively to the gender 
attributable to generic names placed on the Official List in the 
period up to the end of 1936 to which reference has been made in 
paragraph 3 above contain determinations of gender in respect 
of 145 generic names. These Directions conclude the review 
undertaken by the Commission from this point of view of the 
entries made on the Official List of Generic Names in Zoology 
in the period referred to above, save for a small number of cases 
where special problems are involved. The total number of generic 
names placed on the Official List in the period up to the end of 
1936 amounted to 563. Gender determinations for 394 of these 
generic names were made in Section C of the present volume, and 
accordingly the determinations made in the present volume 
brings the total number of determinations made up to a total of 
539. As regards the remaining 24 names, gender determinations 
have also been made for six names, the number of determinations 
still to be made being thus reduced to 18. The distribution by 
Classes of the genera for the names of which gender determinations 
are made in the Directions included in the present Section is shown 
in the following table :— 


IX 
TABLE 2 


Distribution by Classes in the Animal Kingdom of the genera 
for the names of which gender determinations have 
been made in the present Section of Volume 1 


Number of generic 
Name of Class names for which genders 
have been determined 


Rhizopoda 2 
Ciliata 
Mastigophora 
Sporozoa 
Neosporidia 
Hydrozoa 
Anthozoa 
Cestoda 
Trematoda 
Nematoda 
Nematomorpha 
Hirudinea 


Chaetopoda 


— 


— 
ONWNHORK AOR NK KN 


Crustacea 
Diplopoda 
Trilobita 
Insecta 
Merostomata 
Arachnida 
Gastropoda 
Scaphopoda 
Pelecypoda 
Cephalopoda 
Brachiopoda 
Asteroidea 
Holothuroidea 
Crinoidea 
Urochorda 
Amphibia 
Reptilia 
Aves 

Total 145 


No 


rm KO 
= ee CNR KNOW OK ORR NRE 


xX 


6. The Directions comprised in the present Section contain 
529 additions to the Official Lists and Official Indexes. Of the 
entries so made 522 are in respect of the names of taxa of various 
categories and two are of the titles of zoological works. Particulars 
of the additions so made are given in the following table :— 


TABLE 3 
Additions to the ‘‘ Official Lists ’’ and ‘‘ Official Indexes ”’ 
respectively 
Category Official Lists Official Indexes 
Specific Names 322 22 
Generic Names 8 171 
Family-Group Names ] 3 
Titles of Works 1 1 
Totals 332 197 


7. The subject index to the present Section of Volume 1 is of 
the same scope as, and follows the style of the corresponding 
index for the preceding Section (Section C) of this volume. For 
this index the Commission is indebted to Mrs. J. H. Newman. 


FRANCIS HEMMING 


Secretary to the International Commission 
on Zoological Nomenclature 


28 Park Village East, 
Regent’s Park, 
LONDON, N.W.1. 


31st May 1957. 


TABLE OF CONTENTS 


DIRECTION 36 Addition to the Official List of Specific 
Names in Zoology (a) of the specific names of the type 
species of one hundred and seventy genera of the 
Order Decapoda (Class Crustacea), the names of which 
were placed on the Official List of Generic Names in 
Zoology in the period up to the end of 1936 and (b) of 
the specific names currently treated as senior sub- 
jective synonyms of the names of the type species of 
nine other such genera .. 


DIRECTION 37 Completion of the entries relating to 
the names of certain genera of the Order Decapoda 
(Class Crustacea) placed on the Official List of Generic 
Names in Zoology in the period up to the end of 1936. . 


DIRECTION 38 Rejection for nomenclatorial purposes 
of the anonymously published work entitled Orni- 
thologia britannica issued in 1771 and believed to have 
been written by Marmaduke Tunstall and matters 
incidental thereto (cancellation of Opinion 38) 


DIRECTION 39 Substitution of Gallinago Brisson, 1760, 
for Gallinago Koch, 1816 (Class Aves) on the Official 
List of Generic Names in Zoology (correction of an 
error in the Ruling given in Opinion 67) 


DIRECTION 40 Determination of the gender to be 
attributed to the names of fifty-five genera of the 
Phylum Arthropoda, exclusive of genera of Decapod 
Crustacea, placed on the Official List of Generic 
Names in Zoology in the period up to the end of 1936 


DIRECTION 42 Determination of the gender to be 
attributed to the names of seventy-eight genera of 
various Classes of Invertebrates and of six genera of 
the Class Urochorda placed on the Official List of 
Generic Names in Zoology in the period up to the end of 
1936 


XI 


Page 


47 


83 


oF 


139 


155 


XII 


DIRECTION 43 Addition to the Official List of Specific 
Names in Zoology (a) of the specific names of the type 
species of seventy-three genera in the Class Aves, 
the names of which were placed on the Official List 
of Generic Names in Zoology in the period up to the 
end of 1936 and (b) of twelve specific names which 
are currently regarded as senior subjective synonyms 
of such names 


DIRECTION 44 Correction of certain entries relating to 
the names of genera in the Class Aves made on the 
Official List of Generic Names in Zoology in the period 
up to the close of 1936 and addition to the Official 
Indexes of Rejected and Invalid Names in Zoology of 
certain names given to genera and species of the same 
Class he a 


DIRECTION 45 Substitution of Pinnotheres_ Bosc, 
[1801—1802] for Pinnotheres Latreille, [1802—1803] 
(Class Crustacea, Order Decapoda) on the Official 
List of Generic Names in Zoology (correction of an 
error in the Ruling given in Opinion 85) 


DIRECTION 47 Substitution on the Official List of 
Generic Names in Zoology of a revised entry relating 
to the generic name Pandalus Leach, 1815 (Class 
Crustacea, Order Decapoda) (Revision of a Ruling 
given in Opinion 104) mf 


DIRECTION 48 Determination of the authorship to. be 
attributed to the work published anonymously in the 
year 1798 under the title Museum Boltenianum 
(Direction supplementary to Opinion 96) 


DIRECTION 49 Validation under the Plenary Powers 
of the generic name [phis Leach, 1817 (Class Crustacea, 
Order Decapoda) (correction of an error in the Ruling 
given in Opinion 73) 


Page 


171 


211 


pe 


243 


255 


265 


DIRECTION 50 Validation under the Plenary Powers of 
the generic name Homalaspis Milne Edwards (A.), 
1863 (Class Crustacea, Order REN sean 
of an error in Opinion 85) Be tas 


DIRECTION 51 Revision of the entry on the Official 
List of Generic Names in Zoology relating to the name 
Homarus Weber, 1795 (Class Crustacea, Order Deca- 
poda) (revision of a Ruling given in Opinion 104) 


DIRECTION 52 Determination of the gender to be 
attributed to the generic names Cryptobranchus 
Leuckart, 1821 (Class Amphibia) and Eremias 
Wiegmann, 1834 (Class Reptilia) placed on the 
Official List of Generic Names in ae a the 
Ruling given in Opinion 92 ; 


DIRECTION 55 Insertion in the Official List of Generic 
Names in Zoology of an entry relating to the generic 
name Grus Brisson, 1760 (Class Aves) (correction of 
an error in the Ruling given in Opinion 103) .. 


DIRECTION 56 Completion and in certain cases correc- 
tion of entries relating to the names of genera belonging 
to the Classes Pisces, Amphibia and Reptilia made 
in the Official List of Generic Names in 2 i in 
the period up to the end of 1936 a 


DIRECTION 57 Addition to the Official List of Specific 
Names in Zoology (a) of the specific names of forty- 
seven species belonging to the Classes Cyclostomata, 
Pisces, Amphibia and Reptilia, each of which is the 
type species of a genus, the name of which was placed 
on the Official List of Generic Names in Zoology in 
the period up to the end of 1936 and (b) of the specific 
name of one species of the Class Amphibia which is 
currently treated as senior subjective synonym of the 
name of such a species .. 


XIU 


Page 


281 


295 


309 


319 


Sif) 


365 


XIV 


Page 
MINUTE, dated 3lst December 1956, by the Secretary 
containing directions regarding the generic names 
Teuthys Linck, 1790 (Class Pisces) and Teuthis 
Schneider, 1784 (Class Cephalopoda) .. ae +1) 391 
Corrigendan yt 5 Ne a a: aE: ‘aly ans 
Subject Index .. 7! a as a Ps so 395 
Particulars of dates of publication of the several Parts in 
which the present volume was published ne .. 469 


Instructions to Binders ri ry ae = cr 470 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, c.M.G., C.B.E. 


_ Secretary to the Commission 


VOLUME 1. SECTION D. Part D.1. Pp. 1—46 


DIRECTION 36 


Addition to the Official List of Specific Names in Zoology 
(a) of the specific names of the type species of one hundred 
and seventy genera of the Order Decapoda (Class 
Crustacea), the names of which were placed on the 
Official List of Generic Names in Zoology in the period 
up to the end of 1936 and (b) of the specific names 
currently treated as senior subjective synonyms of the 
names of the type-species_of nine other such genera. 


LONDON : 
Printed by Order-of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956 


Price Thirty Shillings 
(All rights reserved) 


Issued \st September, 1956 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 36 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JorDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President: Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., 
U.S.A.) (12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(Arranged in order of precedence by reference to date of election_or of most recent 
re-election, as prescribed by the International Congress of Zoology) 


Professor H. Boscuma (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. Henning Lemcue (Universitetets Zoologiske Museum, Copenhagen, Denmark) 
(27th July 1948) 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RiLey (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a. M., Germany) (5th July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitat zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMoND (University of Toronto, Toronto, Canada) (12th August 1953) 

eee a: oe BRADLEY (Cornell University, Ithaca, N. Y., U.S.A.) (12th August 1953) 
President 

Professor Harold E. Voxes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) 
(12th August 1953) 

Professor Béla HANKO (Mezégazdasdgi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N. Y.,.0 38:22) 
(12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. Hottuuts (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (15th October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, Berkeley, 
California, U.S.A.) (29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum v Praze, Prague, Czechoslovakia) (30th 
October 1954) : 

Professor Dr. Wilhelm KUHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

Professor F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 


Professor Ernst MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico TORTONESE (Museo di Storia Naturale, “ G. Doria,’ Genova, Italy) 
(16th December 1954) 


DIRECTION 36 


ADDITION TO THE “OFFICIAL LIST OF SPECIFIC 
NAMES IN ZOOLOGY ” (a) OF THE SPECIFIC NAMES 
OF THE TYPE SPECIES OF ONE HUNDRED AND 
SEVENTY GENERA OF THE ORDER DECAPODA 
(CLASS CRUSTACEA), THE NAMES OF WHICH 
WERE PLACED ON THE “OFFICIAL LIST 
OF GENERIC NAMES IN ZOOLOGY” IN 
THE PERIOD UP TO THE END OF 1936 
AND (b) OF THE SPECIFIC NAMES 
CURRENTLY TREATED AS SENIOR 
SUBJECTIVE SYNONYMS OF THE 
NAMES OF THE TYPE SPECIES OF 
NINE OTHER SUCH GENERA 


RULING :—(1) The under-mentioned specific names of 
species of the Order Decapoda (Class Crustacea), each 
of which is the type species of a genus, the name of which 
was placed on the Official List of Generic Names in 
Zoology in the period up to the end of 1936 by the Rulings 
given in the Opinions cited below, are hereby placed on the 
Official List of Specific Names in Zoology with the Names 
Numbers severally specified against the names in 
question :— 


(1) gayi Milne Edwards (H.) & Lucas, [1844], as 
published in the combination Acanthocyclus 
gayi (specific name of type species of Acantho- 
cyclus Milne Edwards (H.) & Lucas, [1844]) 
(Opinion 73) (Name No. 765) ; 


(2) erosa Miers, [1877], as published in the combina- 
tion Actaeomorpha erosa (specific name of type 
species of Actaeomorpha Miers, [1877]) (Opinion 
73) (Name No. 766) ; 


(3) tomentosus Dana, 1852, as published in the com- 
bination Actumnus tomentosus (specific name of 
type species of Actumnus Dana, 1851) (Opinion 
73) (Name No. 767) ; 


SEP 9 ° 1956 


OPINIONS AND DECLARATIONS 


(4) erinaceus Fabricius, 1787, as published in the com- 
bination Cancer erinaceus (specific name of type 
species of Arcania Leach, 1817) (Opinion 73) 
(Name No. 768) ; 


(5) sexdentatus Paulson, 1875, as published in the 
combination Archias sexdentatus (specific name 
of type species of Archias Paulson, 1875) (Opinion 
73) (Name No. 769) ; 


(6) cribrarius Lamarck, 1818, as published in the 
combination Portunus cribrarius (specific name of 
type species of Arenaeus Dana, 1851) (Opinion 
73) (Name No. 770) ; 


(7) integerrimus Lamarck, 1818, as published in the 
combination Cancer integerrimus (specific name 
of type species of Atergatis de Haan, [1833]) 
(Opinion 73) (Name No. 771); | 


(8) signatus Adams & White, [1849], as published in 
the combination Carpilius signatus (specific name 
of type species of Atergatopsis Milne Edwards 
(A.), 1862) (Opinion 73) (Name No. 772) ; 


(9) armata Milne Edwards (A.), 1869, as published 
in the combination Banareia armata (specific 
name of type species of Banareia Milne Edwards 
(A.), 1869) (Opinion 73) (Name No. 773) ; 


(10) picta Milne Edwards (H.), 1848, as published in the 
combination Bellia picta (specific name of type 
species of Bellia Milne Edwards (H.), 1848) 
(Opinion 73) (Name No. 774) ; 


(11) hemingi Alcock & Anderson, 1899, as published in 
the combination Benthochascon hemingi (specific 
name of type species of Benthochascon Alcock 
& Anderson, 1899) (Opinion 73) (Name No. 
TID) 3 


(12) rouxii Guérin-Ménéville, 1832, as published in the 
combination Caphyra rouxii (specific name of 
type species of Caphyra Guérin-Ménéville, 1832) 
(Opinion 73) (Name No. 776) ; 


DIRECTION 36 5 


(13) maculatus Linnaeus, 1758, as published in the 
combination Cancer maculatus (specific name of 
type species of Carpilius Desmarest, 1823) 
(Opinion 73) (Name No. 777) ; 


(14) tristis Dana, 1852, as published in the combination 
Carpilodes tristis (specific name of type species 
of Carpilodes Dana, 1851) (Opinion 73) (Name 
No. 778) ; 


(15) papulosus Stimpson, [1871], as published in the 
combination Carpoporus papulosus (specific name 
of type species of Carpoporus Stimpson, [1871]) 
(Opinion 73) (Name No. 779) ; 


(16) tenuipes Dana, 1852, as published in the combina- 
tion Carupa tenuipes (specific name of type 
species of Carupa Dana, 1851) (Opinion 73) 
(Name No. 780) ; 


(17) melanochirus Milne Edwards (A.), 1873, as pub- 
lished in the combination Chlorodopsis melano- 
chirus (specific name of type species of Chloro- 
dopsis Milne Edwards (A.), 1873) (Opinion 73) 
(Name No. 781) ; 


(18) tridentatus Milne Edwards (A.), 1879, as published 
in the combination Coenophthalmus tridentatus 
(specific name of type species of Coenophthalmus 
Milne Edwards (A.), 1879) (Opinion 73) (Name 
No. 782) ; 


(19) chilensis Milne Edwards (H.) & Lucas, [1844], as 
published in the combination Corystoides chilen- 
sis (specific name of type species of Corystoides 
Milne Edwards (H.), & Lucas [1844]) (Opinion 
73) (Name No. 783) ; 


(20) pentagonus Stimpson, 1858, as published in the 
combination Cryptocnemus pentagonus (specific 
name of type species of Cryptocnemus Stimpson, 
1858) (Opinion 73) (Name No. 784) ; 


OPINIONS AND DECLARATIONS 


(21) ornatus Dana, 1852, as published in the combina- 
tion Cyclodius ornatus (specific name of type 
species of Cyclodius Dana, 1851) (Opinion 73) 
(Name No. 785) ; 


(22) andreossyi Audouin, [1826], as published in the 
combination Pilumnus andreossyi (specific name 
of type species of Cymo de Hann, [1833]) 
(Opinion 73) (Name No. 786) ; 


(23) eremita Nobili, 1906, as published in the com- 
bination Dacryopilumnus eremita (specific name 
of type species of Dacryopilumnus Nobili, 1906) 
(Opinion 73) (Name No. 787) ; 


(24) imitatrix Hilgendorf, [1869], as published in the 
combination Deckenia imitatrix (specific name 
of type species of Deckenia Hilgendorf, [1869]) 
(Opinion 73) (Name No. 788) ; 


(25) hispida Eydoux & Souleyet, 1842, as published in 
the combination Domecia hispida (specific name 
of type species of Domecia Eydoux & Souleyet, 
1842) (Opinion 73) (Name No. 789) ; 


(26) tuberosus Pennant, 1777, as published in the 
combination Cancer tuberosus (specific name of 
type species of Ebalia Leach, [1817]) (Opinion 73) 
(Name No. 790) ; 


(27) cubensis Stimpson, 1860, as published in the 
combination Epilobocera cubensis (specific name 
of type species of Epilobocera Stimpson, 1860) 
(Opinion 73) (Name No. 791) ; 


(28) cessacii Milne Edwards (A.), 1878, as published in 
the combination Epimelus cessacii (specific name 
of type species of Epimelus Milne Edwards (A.), 
1878) (Opinion 73) (Name No. 792) ; 


(29) isenbeckii Brandt, 1848, as published in the com- 
bination Platycorystes (Podacanthus) isenbeckii 
(specific name of type species of Erimacrus 
Benedict, 1892) (Opinion 73) (Name No. 793) ; 


DIRECTION 36 7 


(30) spinosus Rathbun, 1894, as published in the com- 
bination Erimetopus spinosus (specific name of 
type species of Erimetopus Rathbun, 1894) 
(Opinion 73) (Name No. 794) ; 


(31) dovii Stimpson, 1860, as published in the com- 
bination Euphylax dovii (specific name of type 
species of Euphylax Stimpson, 1860) (Opinion 
wt tName No, 795): 


(32) jacquemontii Milne Edwards (H.), 1844, as published 
in the combination Gecarcinucus jacquemontii 
(specific name of type species of Gecarcinucus 
Milne Edwards (H.), 1844) (Opinion 73) (Name 
No. 796) ; 


(33) amica Smith (S.I.), 1869, as published in the com- 
bination Hepatella amica (specific name of type 
species of Hepatella Smith (S.I.), 1869) (Opinion 
73) (Name No. 797) ; 


(34) fallax Henderson, 1893, as published in the com- 
bination Ebalia fallax (specific name of type 
species of Heterolithadia Alcock, 1896) (Opinion 
73) (Name No. 798) ; 


(35) vesiculosa Alcock, 1896, as published in the com- 
bination Heteronucia vesiculosus (specific name 
of type species of Heteronucia Alcock, 1896) 
(Opinion 73) (Name No. 799) ; 


(36) rotundifrons Milne Edwards (A.), 1867, as pub- 
lished in the combination Heterozius rotundifrons 
(specific name of type species of Heterozius 
Milne Edwards (A.), 1867) (Opinion 73) (Name 
No. 800) ; 


(37) agilis Milne Edwards (A.), 1872, as published in the 
combination Hydrothelphusa agilis (specific name 
of type species of Hydrothelphusa Milne Edwards 
(A.), 1872) (Opinion 73) (Name No. 801) ; 


(38) subglobosa Stimpson, [1871], as published in the 
combination /liacantha subglobosa (specific name 
of type species of Jliacantha Stimpson, [1871)]) 
(Opinion 73) (Name No. 802) ; 


OPINIONS AND DECLARATIONS 


(39) spongiosus Adams & White, [1849], as published 
in the combination Iphiculus spongiosus (specific 
name of type species of Iphiculus Adams & White 
[1849]) (Opinion 73) (Name No. 803) ; 


(40) cylindricus Fabricius, 1777, as published in the 
combination Cancer cylindricus (specific name of 
type species of Jxa Leach, 1815) (Opinion 73) 
(Name No. 804) ; 


(41) jurinei Saussure, 1853, as published in the com- 
bination Guaia (Ilia) jurinei (specific name of 
type species of Leucosilia Bell, 1855) (Opinion 73) 
(Name No. 805) ; 


(42) polybioides Adams & White, [1849], as published 
in the combination Lissocarcinus polybioides 
(specific name of type species of Lissocarcinus 
Adams & White, [1849]) (Opinion 73) (Name 
No. 806) ; 


(43) cumingii Bell, 1855, as published in the combina- 
tion Lithadia cumingii (specific name of type 
species of Lithadia Bell, 1855) (Opinion 73) 
(Name No. 807) ; 


(44) rotundatus Adams & White, [1849], as published 
in the combination Lupocyclus rotundatus (speci- 
fic name of type species of Lupocyclus Adams & 
White, [1849]) (Opinion 73) (Name No. 808) ; 


(45) lambriformis Milne Edwards (A.), 1873, as published 
in the combination Merocryptus lambriformis 
(specific name of type species of Merocryptus 
Milne Edwards (A.), 1873) (Opinion 73) (Name 
No. 809) ; 


(46) eudactylus Bell, 1855, as published in the combina- 
tion Myrodes eudactylus (specific name of type 
species of Myrodes Bell, 1855) (Opinion 73)(Name 
No. 810) ; 


(47) speciosa Dana, 1852, as published in the combina- 
tion Nucia speciosa (specific name of type species 
of Nucia Dana, 1852) (Opinion 73) (Name No. 
811); 


DIRECTION 36 9 


(48) hardwickii Leach, 1817, as published in the com- 
bination Nursia hardwickii (specific name of 
type species of Nursia Leach, 1817) (Opinion 73) 
(Name 812) ; 


(49) dentata Bell, 1855, as published in the combination 
Nursilia dentata (specific name of type species of 
Nursilia Bell, 1855) (Opinion 73) (Name No. 
813); 


(50) lamelligera Stimpson, 1858, as published in the 
combination Onychomorpha lamelligera (specific 
name of type species of Onychomorpha Stimpson, 
1858) (Opinion 73) (Name No. 814) ; 


(51) horridus Riippell, 1830, as published in the com- 
bination Oreophorus horridus (specific name of 
type species of Oreophorus Riippell, 1830) 
(Opinion 73) (Name No. 815) ; 


(52) tuberosa Stimpson, [1871], as published in the 
combination Osachila tuberosa (specific name of 
type species of Osachila Stimpson, [1871]) 
(Opinion 73) (Name No. 816) ; 


(53) milneedwardsii Miers, 1886, as published in the 
combination Paracyclois milne-edwardsii (speci- 
fic name of type species of Paracyclois Miers, 
1886) (Opinion 73) (Name No. 817) ; 


(54) tridentata Milne Edwards (H.), 1853, as published 
in the combination Parathelphusa tridentata 
(specific name of type species of Parathelphusa 
Milne Edwards (H.), 1853) (Opinion 73) (Name 
No. 818) ; 


(55) orientalis Miers, 1886, as published in the com- 
| bination Lupocyclus (Parathranites) orientalis 
(specific name of type species of Parathranites 
Miers, 1886) (Opinion 73) (Name No. 819) ; 


(56) alcocki Wood-Mason, 1891, as published in the 
combination Parilia alcocki (specific name of 
type species of Parilia Wood-Mason, 1891) 
(Opinion 73) (Name No. 820) ; 


10 OPINIONS AND DECLARATIONS 


(57) coronata Alcock & Anderson, 1894, as published in 
the combination Randallia coronata (specific 
name of type species of Pariphiculus Alcock, 
1896) (Opinion 73) (Name No. 821) ; 


(58) crassipes Bell, 1855, as published in the combina- 
tion Phlyxia crassipes (specific name of type 
species of Phlyxia Bell, 1855) (Opinion 73) (Name 
No. 822) ; 


(59) denticulatus Montagu, 1808, as published in the 
combination Cancer denticulatus (specific name 
of type species of Pirimela Leach, [1816]) 
(Opinion 73) (Name No. 823) ; 


(60) gaudichaudii Milne Edwards (H.), 1837, as pub- 
lished in the combination Platymera gaudi- 
chaudii (specific name of.type species of Platy- 
mera Milne Edwards (H.), 1837) (Opinion 73) 
(Name No. 824) ; 


(61) henslowii Leach, [1820], as published in the com- 
bination Polybius henslowii (specific name of 
type species of Polybius Leach, [1820]) (Opinion 
73) (Name No. 825) ; 


(62) armatus Milne Edwards (H.), 1853, as published 
in the combination Potamocarcinus armatus 
(specific name of type species of Potamocarcinus 
Milne Edwards (H.), 1853) (Opinion 73) (Name 
No. 826) ; 


(63) perlata Milne Edwards (H.), 1837, as published in 
the combination Thelpheusa [sic] perlata (specific 
name of type species of Potamonautes Macleay, 
1838) (Opinion 73) (Name No. 827) ; 


(64) tridentata Miers, 1879, as published in the com- 
bination Pseudophilyra tridentata (specific name 
of type species of Pseudophilyra Miers, 1879) 
(Opinion 73) (Name No. 828) ; 


(65) americana Saussure, 1857, as published in the 
combination Pseudothelphusa americana (specific 
name of type species of Pseudothelphusa Saus- 
sure, 1857) (Opinion 73) (Name No. 829) ; 


DIRECTION 36 11 


(66) ornata Randall, 1840, as published in the com- 
bination J/ia ornata (specific name of type species 
of Randallia Stimpson, 1857) (Opinion 73) (Name 
No. 830) ; 


(67) nodosus Bell, 1855, as published in the combination 
Oreophorus nodosus (specific name of type species 
of Speloeophorus Milne Edwards (A.), 1865) 
(Opinion 73) (Name No. 831); 


(68) bedoti Zehnter, 1894, as published in the combina- 
tion Sphaerocarcinus bedoti (specific name of 
type species of Sphaerocarcinus Zehnter, 1894) 
(Opinion 73) (Name No. 832) ; 


(69) serratus Forskal, 1775, as published in the com- 
bination Cancer serratus (specific name of type 
species of Scy/la de Haan, [1833]) (Opinion 73) 
(Name No. 833) ; 


(70) admete Herbst, [1803], as published in the combina- 
tion Cancer admete (specific name of type species 
of Thalamita Latreille, 1829) (Opinion 73) (Name 
No. 834) ; 


(71) quadridens Milne Edwards (A.), 1869, as published 
in the combination Thalamitoides quadridens 
(specific name of type species of Thalamitoides 
Milne Edwards (A.), 1869) (Opinion 73) (Name 
ING. S35) ; 


(72) danae Milne Edwards (A.), 1869, as published in 
the combination Goniosoma danae (specific name 
of type species of Thalamonyx Milne Edwards 
(A.), 1873) (Opinion 73) (Name No. 836) ; 


(73) muriger Adams & White, [1849], as published in 
the combination Jlos muriger (specific name of 
type species of J/os Adams & White, [1849]) 
(Opinion 73) (Name No. 837) ; 


(74) corallinus Faxon, 1893, as published in the com- 
bination Trachycarcinus corallinus (specific name 
of type species of Trachycarcinus Faxon, 1893) 
(Opinion 73) (Name No. 838) ; 


12 OPINIONS AND DECLARATIONS 


(75) fluviatilis Latreille, [1828], as published in the 
combination Trichodactylus fluviatilis (specific 
name of type species of Trichodactylus Latreille, 
[1828]) (Opinion 73) (Name No. 8339) ; 


(76) nobile Milne Edwards (A.), 1880, as published in 
the combination Trichopeltarion nobile (specific 
name of type species of Trichopeltarion Milne 
Edwards (A.), 1880) (Opinion 73) (Name No. 
840) ; 


(77) serrata White, 1847, as published in the combina- 
tion Valdivia serrata (specific name of type species 
of Valdivia White, 1847) (Opinion 73) (Name No. 
841) ; 


(78) parvula Stimpson, 1858, as published in the com- 
bination Acmaeopleura parvula (specific name 
of type species of Acmaeopleura Stimpson, 1858) 
(Opinion 85) (Name No. 842) ; 


(79) inaequipes Stimpson, 1858, as published in the 
combination Asthenognathus inaequipes (specific 
name of type species of Asthenognathus Stimp- 
son, 1858) (Opinion 85) (Name No. 843) ; 


(80) typhlus Milne Edwards (A.), 1880, as published in 
the combination Bathyplax typhlus (specific 
name of type species of Bathyplax Milne 
Edwards (A.), 1880) (Opinion 85) (Name No. 

); | 

(81) sexdentatum Stimpson, 1858, as published in the 
combination Camptandrium sexdentatum (specific 
name of type species of Camptandrium Stimpson, 
1858) (Opinion 85) (Name No. 845) ; 


(82) coppingeri Miers, 1884, as published in the com- 
bination Camptoplax coppingeri (specific name 
of type species of Camptoplax Miers, 1884) 
(Opinion 85) (Name No. 846) ; 


(83) nitidus Milne Edwards (A.), 1870, as published in 
the combination Catoptrus nitidus (specific name 
of type species of Catoptrus Milne Edwards (A.), 
1870) (Opinion 85) (Name No. 847) ; 


DIRECTION 36 13 


(84) ciliatus Stimpson, 1858, as published in the com- 
bination Ceratoplax ciliatus (specific name of 
type species of Ceratoplax Stimpson, 1858) 
(Opinion 85) (Name No. 848) ; 


(85) typicus Rathbun, 1898, as published in the com- 
bination Chasmocarcinus typicus (specific name 
of type species of Chasmocarcinus Rathbun, 
1898) (Opinion 85) (Name No. 849) ; 


(86) balansae Milne Edwards (A.), 1873, as published 
in the combination Clistocoeloma balansae (speci- 
fic name of type species of Clistocoeloma Milne 
Edwards (A.), 1873) (Opinion 85) (Name No. 
850) ; 


(87) angulatus Dana, 1851, as published in the combina- 
tion Cyrtograpsus angulatus (specific name of 
type species of Cyrtograpsus Dana, 1851) ( Opin- 
ion 85) (Name No. 851) ; 


(88) nitidus Smith (S.I.), 1870, as published in the 
combination Dissodactylus nitidus (specific name 
of type species of Dissodactylus Smith (S.I.), 
1870) (Opinion 85) (Name No. 852) ; 


(89) carinipes de Man, 1889, as published in the com- 
bination Durckheimia carinipes (specific name 
of type species of Durckheimia de Man, 1889) 
(Opinion 85) (Name No. 853) ; 


(90) liguricus Milne Edwards (H.), 1853, as es ieee 
in the combination Euchirograpsus liguricus 
(specific name of type species of Euchirograpsus 
Milne Edwards (H.), 1853) (Opinion 85) (Name 
No. 854) ; 


(91) crenatus de Haan, [1835], as published in the com- 
bination Cancer (Eucrate) crenatus (specific name 
of type species of Eucrate de Haan, [1835)) 
(Opinion 85) (Name No. 855) ; 


14 OPINIONS AND DECLARATIONS 


(92) agassizii Milne Edwards (A.), 1880, as published 
in the combination Eucratodes agassizii (specific 
name of type species of Eucratodes Milne 
Edwards (A.), 1880) (Opinion 85) (Name No. 
856) ; 


(93) crassimanus Dana, 1851, as published in the 
combination Eucrate crassimanus (specific name 
of type species of Eucratopsis Smith (S.I.), 1869) 
(Opinion 85) (Name No. 857) ; 


(94) deplanatus Cano, 1889, as published in the com- 
bination Euryetisus deplanatus (specific name of 
type species of Euryetisus Cano, 1889) (Opinion 
85) (Name No. 858) ; 


(95) nitidus Stimpson, 1859, as published in the com- 
bination Euryplax nitidus (specific name of type 
species of Euryplax Stimpson, 1859) (Name No. 
859) ; 


(96) limosa Say, 1818, as published in the combination 
Cancer limosa (specific name of type species of 
Eurytium Stimpson, 1859) (Opinion 85) (Name 
No. 860) ; 


(97) subquadrata Dana, 1851, as published in the com- 
bination Fabia subquadrata (specific name of 
type species of Fabia Dana, 1851) (Opinion 85) 
(Name No. 861) ; 


(98) bispinosus Herbst, [1783], as published in the 
combination Cancer bispinosus (specific name of 
type species of Galene de Haan, [1833]) (Opinion 
85) (Name No. 862) ; 


(99) tridens Kreyer, 1837, as published in the combina- 
tion Geryon tridens (specific name of type species 
of Geryon Kroyer, 1837) (Opinion 85) (Name 
No. 863) ; 


(100) impressus Smith (S.I.), 1870, as published in the 
combination Glyptograpsus impressus (specific 
name of type species of Glyptograpsus Smith 
(S.I.), 1870) (Opinion 85) (Name No. 864) ; 


DIRECTION 36 15 


(101) pugnax Smith (S.I.), 1870, as published in the 
combination Glyptoplax pugnax (specific name 
of type species of Glyptoplax Smith (S.I.), 1870) 
(Opinion 85) (Name No. 865) ; 


(102) bicornis Gray (J.E.), 1831, as published in the 
combination Gomeza bicornis (specific name of 
type species of Gomeza Gray (J.E.), 1831) 
(Opinion 85) (Name No. 866) ; 


(103) fragifer de Haan, [1835], as published in the com- 
bination Cancer (Halimede) fragifer (specific 
name of type species of Halimede de Haan, 
[1835]) (Opinion 85) (Name No. 867) ; 


(104) tridens de Haan, [1835], as published in the com- 
bination Ocypode (Helice) tridens (specific name 
of type species of Helice de Haan, [1835)) 
(Opinion 85) (Name 868) ; 


(105) lugubris Alcock, 1899, as published in the com- 
bination Hephthopelta lugubris (specific name of 
type species of Hephthopelta Alcock, 1899) 
(Opinion 85) (Name No. 869) ; 


(106) sexpes Fabricius, 1798, as published in the com- 
bination Cancer sexpes (specific name of type 
species of Hexapus de Haan, [1833]) (Opinion 
85) (Name No. 870) ; 


(107) haematocheir de Haan, [1835], as published in the 
combination Grapsus (Pachysoma) haematocheir 
(specific name of type species of Holometopus 
Milne Edwards (H.), 1853) (Opinion 85) (Name 
No. 871) ; 


(108) trapeziformis Nauck, 1880, as published in the 
combination Holothuriophilus trapeziformis 
(specific name of type species of Holothurio- 
Bi). Nauck, 1880) (Opinion 85) (Name No. 
Si2y% 


16 OPINIONS AND DECLARATIONS 


(109) rodgersii Stimpson, 1858, as published in the com- 
bination Lachnopodus rodgersii (specific name 
of type species of Lachnopodus Stimpson, 1858) 
(Opinion 85) (Name No. 873) ; 


(110) exaratus Milne Edwards (H.), 1834, as published 
in the combination Chlorodius exaratus (specific 
name of type species of Leptodius Milne Edwards 
(A.), 1863) (Opinion 85) (Name No. 874) ; 


(111) rubromaculatus de Haan, [1833], as published in 
the combination Cancer (Liagore) rubromacu- 
latus (specific name of type species of Liagore 
de Haan, [1833]) (Opinion 85) (Name No. 875) ; 


(112) nitidus Milne Edwards (A.), 1867, as published in 
the combination Libystes nitidus (specific name 
of type species of Libystes Milne Edwards (A.), 
1867) (Opinion 85) (Name No. 876) ; 


(113) cinctimanus White, [1849], as published in the com- 
bination Carpilius cinctimanus (specific name of 
type species of Liomera Dana, 1851) (Opinion 85) 
(Name No. 877) ; 


(114) leeanus Rathbun, 1898, as published in the com- 
bination Lipaesthesius leeanus (specific name of 
type species of Lipaesthesius Rathbun, 1898) 
(Opinion 85) (Name No. 878) ; 


(115) bispinosa Kinahan, 1856, as published in the com- 
bination Litocheira bispinosa (specific name of 
type species of Litocheira Kinahan, 1856) (Opin- 
ion 85) (Name No. 879) ; 


(116) bella Stimpson, 1860, as published in the combina- 
tion Xantho bella (specific name of type species 
of Lophopanopeus Rathbun, 1898) (Opinion 85) 
(Name No. 880) ; 


(117) dilatipes Adams & White, [1849], as published in 
the combination Pilumnus dilatipes (specific 
name of type species of Lophopilumnus Miers, 
1886) (Opinion 85) (Name No. 881) ; 


DIRECTION 36 i? 


(118) tesselatus Latreille, 1812, as published in the 
combination Grapsus tesselatus (specific name 
of type species of Lybia Milne Edwards (H.), 
1834) (Opinion 85) (Name No. 882) ; 


(119) thalamita Stimpson, [1871], as published in the 
combination Melybia thalamita (specific name 
of type species of Melybia Stimpson, [1871)]) 
(Opinion 85) (Name No. 883) ; 


(120) rousseauxi Milne Edwards (H.), 1853, as published 
in the combination Metasesarma rousseauxi 
(specific name of type species of Metasesarma 
Milne Edwards (H.), 1853) (Opinion 85) (Name 
No. 884) ; 


(121) truncatus Stimpson, [1860], as published in the 
combination Metopocarcinus truncatus (specific 
name of type species of Metopocarcinus Stimp- 
son, [1860]) (Opinion 85) (Name No. 885) ; 


(122) sculptipes Stimpson, [1871], as published in the 
combination Micropanope sculptipes (specific 
name of type species of Micropanope Stimpson, 
[1871] (Opinion 85) (Name No. 886) ; 


(123) nitidus Milne Edwards (A.), 1873, as published in 
the combination Notonyx nitidus (specific name 
of type species of Notonyx Milne Edwards (A.), 
1873) (Opinion 85) (Name No. 887) ; 


(124) granulatus Rathbun, 1893, as published in the 
combination Oediplax granulatus (specific name 
of type species of Oediplax Rathbun, 1893) 
(Opinion 85) (Name No. 888) ; 


(125) macgillivrayi White, 1852, as published in the com- 
bination Ommatocarcinus macgillivrayi (specific 
name of type species of Ommatocarcinus White, 
1852) (Opinion 85) (Name No. 889) ; 


(126) transversus Rathbun, 1893, as published in the 
combination Opisthopus transversus (specific 
name of type species of Opisthopus Rathbun, 
1893) (Opinion 85) (Name No. 890) ; 


18 OPINIONS AND DECLARATIONS 


(127) microps Alcock & Anderson, 1894, as published 
in the combination Xanthodes microps (specific 
name of type species of Orphnoxanthus Alcock, 
1898) (Opinion 85) (Name No. 891) ; 


(128) depressa Stimpson, [1871], as published in the 
combination Panoplax depressa (specific name 
of type species of Panoplax Stimpson, [1871]) 
(Opinion 85) (Name No. 892) ; 


(129) neapolitana Kossmann, 1878, as published in the 
combination Paragalene neapolitana (specific 
name of type species of Paragalene Kossmann, 
1878) (Opinion 85) (Name No. 893) ; 


(130) ewagora de Man, 1895, as published in the com- 
bination Parapanope euagora (specific name of 
type species of Parapanope de Man, 1895) 
(Opinion 85) (Name No. 894) ; 


(131) roseus Nobili, 1906, as published in the combination 
Parapleurophrycoides roseus (specific name of 
type species of Parapleurophrycoides Nobili, 
1906) (Opinion 85) (Name No. 895) ; 


(132) planissimus Herbst, [1804], as published in the 
combination Cancer planissimus (specific name 
of type species of Percnon Gistl, 1848) (Opinion 
85) (Name No. 896) ; 


(133) excelsus Heller, 1862, as published in the com- 
bination Perigrapsus excelsus (specific name of 
type species of Perigrapsus Heller, 1862) (Opinion 
85) (Name No. 897) ; 


(134) perlatus Poeppig, 1836, as published in the com- 
bination Hepatus perlatus (specific name of type 
species of Pilumnoides Milne Edwards (H.) & 
Lucas, [1844]) (Opinion 85) (Name No. 898) ; 


(135) hirtellus Linnaeus, 1761, as published in the com- 
bination Cancer hirtellus (specific name of type 
species of Pilumnus Leach, 1815) (Opinion 85) 
(Name No. 899) ; 


DIRECTION 36 19 


(136) hirtipes Heller, [1865], as published in the combina- 
tion Pinnaxodes hirtipes (specific name of type 
species of Pinnaxodes Heller, [1865]) (Opinion 85) 
(Name No. 900) ; 


(137) cylindricum Say, 1818, as published in the com- 
bination Pinnotheres cylindricum (specific name 
of type species of Pinnixa White, 1846) (Opinion 
85) (Name No. 901) ; 


(138) /aevigata Milne Edwards (H.) & Lucas, [1844], as 
published in the combination Pinnotherelia 
laevigata (specific name of type species of 
Pinnotherelia Milne Edwards (H.), & Lucas, 
[1844]) (Opinion 85) (Name No. 902) ; 


(139) spectabilis de Man, 1896, as published in the 
combination Platychirograpsus spectabilis (speci- 
fic name of type species of Platychirograpsus 
de Man, 1896) (Opinion 85) (Name No. 903) ; 


(140) gracilipes Alcock, 1894, as published in the com- 
bination Platypilumnus gracilipes (specific name 
of type species of Platypilumnus Alcock, 1894) 
(Opinion 85) (Name No. 904) ; 


(141) orbignyi Milne Edwards (H.) & Lucas, [1844], as 
published in the combination Xantho orbignyi 
(specific name of type species of Platyxanthus 
Milne Edwards (A.), 1863) (Opinion 85) (Name 
No. 905) ; 


(142) cupulifer Latreille, 1825, as published in the com- 
bination Pilumnus cupulifer (specific name of 
type species of Polydectus Milne Edwards (H.), 
1837) (Opinion 85) (Name No. 906); 


(143) spinicarpus Milne Edwards (H.), 1852, as published 
in the combination Prionoplax spinicarpus (speci- 
fic name of type species of Prionoplax 
Milne Edwards (H.), 1852) (Opinion 85) (Name 
No. 907) ; 


20 OPINIONS AND DECLARATIONS 


(144) gigas Lamarck, 1818, as published in the combina- 
tion Cancer gigas (specific name of type species 
of Pseudocarcinus Milne Edwards (H.), 1834) 
(Opinion 85) (Name No. 908) ; 


(145) carinata Ortmann, 1894, as published in the com- 
bination Pseudopinnixa carinata (specific name of 
type species of Pseudopinnixa Ortmann, 1894) 
(Opinion 85) (Name No. 909) ; 


(146) gquadridentata Latreille, 1825, as published in the 
combination Melia quadridentata (specific name 
of type species of Pseudorhombila Milne Ed- 
wards (H.), 1837) (Opinion 85) (Name No. 910) ; 


(147) stridulans Wood-Mason, 1892, as published in the 
combination Psopheticus stridulans (specific name 
of type species of Psopheticus Wood-Mason, 
1892) (Opinion 85) (Name No. 911) ; 


(148) glaber Stimpson, 1858, as published in the com- 
bination Ptychognathus glaber (specific name of 
type species of Ptychognathus Stimpson, 1858) 
(Opinion 85) (Name No. 912) ; 


(149) granulosus Milne Edwards (A.), 1879, as published 
in the combination Pyxidognathus granulosus 
(specific name of type Pyxidognathus Milne 
Edwards (A.), 1879) (Opinion 85) (Name No. 
913); 


(150) harrisii Gould, 1841, as published in the combina- 
tion Pilumnus harrisii (specific name of type 
species of Rhithropanopeus Rathbun, 1898) 
(Opinion 85) (Name No. 914) ; 


(151) gracilipes Stimpson, 1858, as published in the 
combination Rhizopa gracilipes (specific name of 
type species of Rhizopa Stimpson, 1858) (Opinion 
85) (Name No. 915) ; 

(152) convexus Milne Edwards (A.), 1867, as published 
in the combination Ruppellioides convexus (speci- 
fic name of type species of Ruppellioides Milne 
es ((A.), 1867) (Opinion 85) (Name No. 
916); 


DIRECTION 36 21 


(153) crassum Dana, 1851, as published in the combina- 
tion Samartium crassum (specific name of type 
species of Samartium Dana, 1851) (Opinion 85) 
(Name No. 917) ; 


(154) spinosipes Stimpson, 1858, as published in the 
combination Scalopidia spinosipes (specific name 
of type species of Scalopidia Stimpson, 1858) 
(Opinion 85) (Name No. 918) ; 


(155) granulatus Rathbun, 1893, as published in the 
combination Scleroplax granulatus (specific name 
of type species of Scleroplax Rathbun, 1893) 
(Opinion 85) (Name No. 919) ; 


(156) carolinensis Stimpson, 1859, as published in the 
combination Speocarcinus carolinensis (specific 
name of type species of Speocarcinus Stimpson, 
1859) (Opinion 85) (Name No. 920) ; 


(157) nitidus Stimpson, 1858, as published in the com- 
bination Sphaerozius nitidus (specific name of 
type species of Sphaerozius Stimpson, 1858) 
(Opinion 85) (Name No. 921) ; 


ao bidentatus Milne Edwards (A.), 1880, as published 
in the combination Yanthodes bidentatus (specific 
name of type species of Tetraxanthus Rathbun, 
1898) (Opinion 85) (Name No. 922) ; 


(159) scabripes Rathbun, 1898, as published in the 
combination Tetrias scabripes (specific name of 
type species of Tetrias Rathbun, 1898) (Opinion 
85) (Name No. 923) ; 


(160) anomalipes Miers, 1881, as published in the com- 
bination Thaumastoplax anomalipes (specific 
name of type species of Thaumastoplax Miers, 
1881) (Opinion 85) (Name No. 924) ; 


(161) gracilipes White, 1847, as published in the com- 
bination Utica gracilipes (specific name of type 
species of Utica White, 1847) (Opinion 85) 
(Name No. 925) ; 


22 OPINIONS AND DECLARATIONS 


(162) litteratus Fabricius, 1798, as published in the 
combination Cancer litteratus (specific name of 
type species of Varuna Milne Edwards (H.), 1830) 
(Opinion 85) (Name No. 926) ; 


(163) murigera White, 1846, as published in the com- 
bination Xanthasia murigera (specific name of 
type species of Xanthasia White, 1846) (Opinion 
85) (Name No. 927) ; 


(164) sternberghii Stimpson, 1859, as published in the 
combination Xanthodius sternberghii (specific 
name of type species of Xanthodius Stimpson, 
1859) (Opinion 85) (Name No. 928) ; 


(165) moebii Richters, 1880, as published in the com- 
bination X enophthalmodes moebii (specific name 


of type species of Xenophthalmodes Richters, 
~ 1880) (Opinion 85) (Name No. 929) ; 


(166) pinnotheroides White, 1846, as published in the com- 
bination Xenophthalmus pinnotheroides (specific 
name of type species of Xenophthalmus White, 
1846) (Opinion 85) (Name No. 930) ; 


(167) aeneus Linnaeus, 1758, as published in the com- 
bination Cancer aeneus (specific name of type 
species of Zosimus Leach, 1823) oe 85) 
(Name No. 931) ; 


(168) carinipes Heller, 1861, as published in the com- 
bination Zozymodes carinipes (specific name of 
type species of Zozymodes Heller, 1861) (Opinion 
85) (Name No. 932) ; 


(169) pagurus Linnaeus, 1758, as published in the com- 
bination Cancer pagurus (specific name of type 
species of Cancer Linnaeus, 1758) (Opinion 104) 
(Name No. 933) ; 


(170) norvegicus Linnaeus, 1758, as published in the 
combination Cancer norvegicus (specific name of 
type species of Nephrops [Leach], [1814]) (Opinion 
104) (Name No. 934). 


DIRECTION 36 23 


(2) The under-mentioned specific names, each of which 
is currently regarded as a senior subjective synonym of 
the specific name of the type species of a genus, the name 
of which was placed on the Official List of Generic Names 
in Zoology in the Opinion cited against the name in 
question, are hereby placed on the Official List of Specific 
Names in Zoology with the Name Numbers severally 
specified below :— 


(1) savignyi (correction of savignii) Milne Edwards (H.), 
1834, as published in the combination of Cancer 
savignii (Opinion 73) (Name No. 935) ; 


(2) superba Costa, 1838, as published in the combination 
Portunus superba (Opinion 73) (Name No. 936) ; 


(3) punctatus Linnaeus, 1767, as published in the com- 
bination Cancer punctatus (Opinion 73) (Name 
No:2937); 


(4) vigil Weber, 1795, as published in the combination 
Portunus vigil (Opinion 73) (Name No. 938) ; 


(5) cheiragonus Tilesius, 1815, as published in the com- 
bination Cancer cheiragonus (Opinion 73) (Name 
No. 939) : 


(6) frontalis Milne Edwards (H.), 1834, as published in 
the combination Ozius frontalis (Opinion 85) 
(Name No. 940) ; 


(7) rhomboides Linnaeus, 1758, as published in the 
combination Cancer rhomboides (Opinion 85) 
(Name No. 941) ; 


(8) barbiger Poeppig, 1836, as published in the com- 
bination Gecarcinus barbiger (Opinion 85) (Name 
No. 942) ; 


(9) minutus Linnaeus, 1758, as published in the com- 
bination Cancer minutus (Opinion 85) (Name No. 
943). 


(3) The under-mentioned specific name is hereby 
placed on the Official Index of Rejected and Invalid 


24 OPINIONS AND DECLARATIONS 


Specific Names in Zoology with the Name No. 335 :— 
savignii Milne Edwards (H.), 1834, as published in the 
combination Cancer savignii (an Invalid Original Spelling 
for savignyi). 


(4) The dates attributed to the under-mentioned specific 
names in the Opinions specified below are hereby corrected 
as follows :— 


(1) rouxii Guérin-Ménéville, as published in the com- 
bination Caphyra rouxii (Opinion 73) : date to be 
corrected from 1830 to 1832; 


(2) amica Smith (S.L.), as published in the combination 
Hepatella amica (Opinion 73) : date to be corrected 
from 1870 to 1869 ; 


(3) signatus Adams & White, as published in the 
combination Carpilius signatus (Opinion 73) : 
date to be corrected from 1848 to [1849] ; 


(4) spongiosus Adams & White, as published in the 
combination Jphiculus spongiosus (Opinion 73) : 
date to be corrected from 1848 to [1849] ; 


(5) polybioides Adams & White, as published in the 
combination Lissocarcinus polybioides (Opinion 
73) : date to be corrected from 1848 to [1849] ; 


(6) rotundatus Adams & White, as published in the 
combination Lupocyclus rotundatus (Opinion 73) : 
date to be corrected from 1848 to [1849] ; 


(7) muriger Adams & White, as published in the com- 
bination Tlos muriger (Opinion 73): date to be 
corrected from 1848 to [1849] ; 


(8) fluviatilis Latreille, as published in the combination 
Trichodactylus fluviatilis (Opinion 73) : date to be 
corrected from 1825 to [1828] ; 


(9) cinctimanus Adams & White, as published in the 
combination Carpilius cinctimanus (Opinion 85) : 
date to be corrected from 1848 to [1849] ; 


DIRECTION 36 25 


(10) Jaevigata Milne Edwards (H.) & Lucas, as pub- 
lished in the combination Pinnotherelia laevigata 
(Opinion 85) : date to be corrected from 1843 to 
[1844] ; 


(11) orbignyi Milne Edwards (H.) & Lucas, as published 
in the combination Xantho orbignyi (Opinion 85) : 
date to be corrected from 1843 to [1844] ; 


(12) tesselatus Latreille, as published in the combina- 
tion Graspus tesselatus (Opinion 85): date to be 
corrected from 1818 to 1812. 


I—THE SUBJECT MATTER OF THE PRESENT 
** DIRECTION ” 


The present Direction has been prepared under a General 
Directive issued to the International Commission on Zoological 
Nomenclature by the Thirteenth International Congress of 
Zoology, Paris, 1948, that the specific names of the species 
which are the type species of genera, the names of which had 
been placed on the Official List of Generic Names in Zoology 
prior to the opening of that Congress should, if the oldest 
available names for the species concerned, be placed on the 
Official List of Specific Names in Zoology and that in other cases 
there should be placed on that List whatever is currently 
considered to be the oldest available names for the species 
concerned. The present, which is the second of this series of 
Directions, is concerned with the names of the type species of 
genera of the Order Decapoda (Class Crustacea), the names of 
which were placed on the Official List of Generic Names in 
Zoology in the pre-Lisbon (1935) period. The total number of 
‘such generic names is one hundred and ninety. At the time of the 
submission of the proposals relating to the names with which 
the present Direction is concerned, decisions on three names had 
been taken on separate Directions, proposals on three others 
had been published in the Bulletin of Zoological Nomenclature 
and were awaiting decision by the Commission, while in two 
other cases the material so far collected was not considered 


26 OPINIONS AND DECLARATIONS 


sufficient for framing proposals for the consideration of the 
Commission. The number of names on which proposals were 
submitted to the Commission amounted therefore to one hundred 
and eighty-two. In the case of three of these names additional 
information received during the Prescribed Voting Period led 
to the withdrawal of the proposals submitted in order to permit 
of the further examination of the issues so raised (paragraph 4 
below). Accordingly, the present Direction contains decisions on 
one hundred and seventy-nine of the names involved. The 
proposals which formed the basis of the decisions taken by the 
Commission in the present case were submitted by the Secretary 
in the following paper on 19th September 1955 :— 


Proposed addition to the ‘‘ Official List of Specific Names in Zoology ”’ 
of the specific names of the type species of one hundred and seventy- 
three genera of the Order Decapoda (Class Crustacea), the 
names of which were placed on the ‘* Official List of Generic 
Names in Zoology ”’ in the period up to the end of 1936 
and of nine specific names which are currently 
treated as senior subjective synonyms of the 
names of type species of such genera 


By FRANCIS HEMMING, C.M.G., C.B.E., 
(Secretary to the International Commission on Zoological Nomenclature) 


The purpose of the present paper is to lay recommendations before 
the International Commission to enable it to comply, so far as concerns 
names of species of the Order Decapoda (Class Crustacea), with a 
General Directive issued to it by the Thirteenth International Congress 
of Zoology, Paris, 1948. This Directive stated that the specific names 
of the type species of genera, the names of which had been placed on 
the Official List of Generic Names in Zoology in the period up to the 
end of 1936, should be placed on the Official List of Specific Names in 
Zoology, save where any given such name is not the oldest available 
name for the taxon concerned, in which case the oldest available name, 
whatever it may be, is to be placed on that Official List. 


2. It will be recalled that, in a paper (Z.N.(S.) 906) which I sub- 
mitted to the Commission on 11th February last, with Voting Paper 
V.P.(O.M.)(55)6, I dealt, so far as the Crustacea Decapoda are con- 
cerned, with the first of the two principle stages (namely the determina- 
tion of the gender of each name) involved in completing the entries 
on the Official List of Generic Names in Zoology made in the pre- 
Lisbon (1935) period, as a preliminary to the publication of the volume 
containing the first instalment of the Official Lists. In that paper 
I reported that in the period under review the names of one hundred 
and ninety genera of Decapoda had been placed on the Official List. 


DIRECTION 36 oP 


3. In compiling the present proposals, I have, as on previous 
similar occasions, consulted such catalogues, checklists and mono- 
graphs as are available. I have also consulted Dr. Isobel Gordon 
(British Museum (Natural History), London), who has rendered the most 
valuable assistance in this matter. In both of the two principal 
Opinions concerned (Opinions 73 and 85) a note was added in every 
case where the applicant (the late Miss Mary Rathbun) was of the 
opinion that the name of the type species of any given genus was a 
junior subjective synonym of some other name. Dr. Gordon points 
out that Miss Rathbun was the leading authority of her day in this 
group and that it is likely that her synonymisations, which were most 
carefully made, have been generally followed by later workers. It is 
accordingly believed that the 173 specific names of type species shown 
in Annexe | to the present paper are the oldest available names for the 
species concerned. Similarly, it is believed that in the case of the nine 
genera specified in Column (2) of Annexe 2, the name of the type 
species is a junior subjective synonym of the name specified in Column 
(1) in that Annexe. 


4. The eight generic names as regards the names of the type species 
of which no proposals are now submitted are the following :—(1) All 
outstanding questions have been settled in regard to the following 
three generic names, which have formed the subject of individual 
Directions by the Commission :—Astacus (Direction 12); Penaeus 
(Direction 15) ; Favus (Direction 25). (2) In the following three cases 
action under the Plenary Powers is required and proposals have been 
published in the Bulletin of Zoological Nomenclature!:—Iphis (Bull. 
11 : 79—81); Homalaspis (Bull. 11 : 83—85) ; Homarus (Bull. 11 : 
114—116). (3) The two remaining names are Daira and Chasmagna- 
thus. As regards each of these proposals for action under the Plenary 
Powers will, it is proposed, be submitted at an early date, in the first 
case for the purpose of validating the generic name Daira, in the second 
case, for the purpose of validating the specific name of the type species 
of Chasmagnathus. 


5. In twelve cases investigations undertaken in this Office have 
shown that the date given for a particular name at the time when it 
was placed on the Official List was incorrect. In some cases (e.g. 
Adams & White, The Voyage of H.M.S. Samarang) Miss Rathbun 
accepted the date printed on the Title Page as the date of publication, 
but later investigations have shown that publication did not take place 
untila later year. In some of the cases of this kind the date of a generic 
name already placed on the Official List is affected as well as the 
name of the type species of such a genus. Proposals for the necessary 
corrections, so far as specific names are concerned, are submitted in 
Annexe 3 to the present paper. 


1 Decisions have since been taken by the International Commission on each of 
these cases and the decisions have been embodied in Directions which will be 
published in the present Section of Volume 1. The Directions concerned are : 
(i) Direction 49 Uphis); (i) Direction 50 (Homalaspis) ; (iil) Direction 51 
(Homarus). 


28 OPINIONS AND DECLARATIONS 


6. In all cases where a specific name consists of a noun in the 
genitive singular based upon the modern patronymic of a man, to which 
was added a double “i” in place of a single ““i’’, the form with a 
double “i” has been retained in accordance with the decision on that 
subject taken by the Copenhagen (1953) Congress.? In one case, a 
name based upon a modern patronymic, (Savigny), was incorrectly 
formed by the substitution of the letter ‘“‘i’’ for the letter “‘ y’’, the 
name being published in the form “‘ savignii”’. In this case, in accord- 
ance with another of the decisions taken by the Copenhagen Congress? 
the form of the name has been corrected to savignyi. 


7. In two cases (Ebalia ; Portumnus) the author of a generic name 
introduced a new name for the type species, at the same time specifying 
the previously published name for which his new specific name was 
a substitute. In each of the cases concerned, the new (substitute) 
name was unnecessary and is invalid as being a junior objective 
synonym of the name which it was intended to replace. In both these 
cases the older (valid) specific name and not the objectively identical 
junior synonym is treated as the name of the type species of the genus 
concerned.* | 


8. In the case of the name Podophthalmus Lamarck, 1801, the name of 
the type species (Podophtalmus [sic] spinosus Lamarck, 1801) is 
currently treated as a junior subjective synonym of Portunus vigil 
Fabricius, 1798 (Suppl. Ent. syst. : 363). However, the latter name is 
a junior homonym of Portunus vigil Weber, 1795 (Nomencl. Ent. syst. 
Fabric. : 93). So many of the names in Weber’s ill-starred little book 
are nomina nuda that as a first step I consulted it to see what Weber 
had actually said. For the benefit of anyone who has never had 
occasion to look at this book, I should explain that Weber did not 
give individual diagnoses either for the genera or for the species 
included by him in it. What he did commonly do, however, was to 
divide up into groups the species placed by him in any given genus 
and to provide a few words (in Latin) to differentiate from one another 
the groups so established. It is only on the rare occasions when 
Weber placed a single species in a group that the words given as a 
definition for that group become also a definition for the species placed 
in it. The present is an example of this unusual situation, for in the 
first of the groups which he established within the genus Portunus, 
Weber placed only one species, Portunus vigil. In consequence this 
name is not a nomen nudum, but the indication given is so extremely 
brief, being simply “‘ thor. utr. bid.’’, that it might not be sufficient 
to permit of the identification of Weber’s species. In that case the 
name Portunus vigil Weber, though an available name, would be a 
nomen dubium. Weber’s book was, as its title indicates, a preview of 


2 See 1953, Copenhagen Decisions zool. Nomencl. : 54, Decision 91. 
3 See 1953, Copenhagen Decisions zool. Nomencl. : 51—52, Decision 86(1)(b). 


4 This procedure has since been formalised by the Ruling given in Declaration 
21 (1956, Ops. Decls. int. Comm. zool. Nomencl. 12 : i—viii). 


DIRECTION 36 29 


what Fabricius at that time intended to write in his Supplementum 
Entomologiae systematicae. Where, as happened in the present case, 
Weber cited a specific name which he knew was a manuscript name 
destined by Fabricius first to see the light of day in his Supplementum, he 
added the letter “‘ S ’ after the specific name concerned. The presence 
of this notation after the name vigil in Weber’s book opened the 
possiblity that in addition to being a senior primary homonym of 
Portunus vigil Fabricius, 1798 (the year in which that author’s Supple- 
mentum was published), the name Portunus vigil Weber might be also 
a senior objective synonym of P. vigil Fabricius. At this point I sought 
the views of two leading specialists, namely, Dr. Fenner A. Chace, Jr. 
(U.S. National Museum, Washington, D.C., U.S.A.) and Dr. L. B. 
Holthuis (Rijksmuseum van Natuurlijke Historie, Leiden, The Nether- 
lands). Each of these specialists agreed that the name Portunus vigil 
Weber and Portunus vigil Fabricius were applicable to the same species, 
Dr. Chace pointing out that there are very few Portunid crabs with two 
lateral teeth on each side of the carapace and Dr. Holthuis drawing 
attention to the significance to be attached to the addition by Weber 
of the letter ““S’’ after the name vigi/. Both explained that the name 
Portunus vigil Weber appeared to have been overlooked in the literature 
and that this binomen was in consequence currently attributed to 
Fabricius, 1798. Both were agreed that no confusion or difficulty 
need be apprehended if in future this name were to be attributed to 
Weber instead of to Fabricius. Since the change involved is one of 
authorship and date only and does not involve a change of name, 
I do not anticipate that anyone would consider it necessary for the 
Commission to use its Plenary Powers in order to suppress Weber’s 
name for the purpose of validating the same name as used by Fabricius 
for the same species three years later. I accordingly share the view 
expressed by the foregoing specialists that this is a case where the 
normal priority rule can properly be allowed to operate. In the 
proposals submitted in the present paper I accordingly recommend 
(Annexe 2) that the name vigil Weber, 1795, as published in the 
combination Portunus vigil, should, as the oldest name subjectively 
available for the type species of the genus Podophthalmus Lamarck, 
1801, be placed on the Official List of Specific Names in Zoology. 


9. The recommendations which are now submitted for the considera- 
tion of the Commission are as follows :— 


(a) that the one-hundred and seventy-three specific names specified 
in Annexe 1, each of which is the specific name of the type 
species of a genus of Crustacea Decapoda, the name of which 
was placed on the Official List of Generic Names in Zoology 
in the period up to the end of 1936, should now be placed on 
the Official List of Specific Names in Zoology (paragraph 3); 


_(b) that the nine specific names specified in Annexe 2, each of which 
is a senior subjective synonym of the specific name of the type 
species of a genus of Crustacea Decapoda, the name of which 


30 OPINIONS AND DECLARATIONS 


was placed on the Official List of Generic Names in Zoology 
in the period up to the end of 1936, should now be placed on 
the Official List of Specific Names in Zoology (paragraph 3) ; 


(c) that in the case of the twelve names specified in Annexe 3, for 
each of which an incorrect date of publication was cited in the 
Opinion concerned, the correct date, as shown in Column 
(3) of the said Annexe, be entered on the Official List of Specific 
Names in Zoology (paragraph 5) ; 


ANNEXE 1 


Specific names of type species of genera of Decapod Crustacea now 
proposed for addition to the ‘* Official List of Specific 
Names in Zoology ”’ 


Specific name Combination in which Name of genus of 
proposed to be placed name specified in which species 
on the “‘Official List — Col. 1 was originally specified in Col. 1 
of Specific Names in published is the type species 
Zoology ”’ 
(1) (2) (3) 
Opinion 73 
gayi Milne Edwards & Acanthocyclus gayi Acanthocyclus Milne 
Lucas, [1844] Edwards & Lucas, 
[1844] 
erosa Miers, [1877] Actaeomorpha erosa Actaeomorpha Miers, 
[1877] 
tomentosus Dana, 1852 Actumnus tomentosus Actumnus Dana, 1851 
erinaceus Fabricius, Cancer erinaceus Arcania Leach, 1817 
1787 


sexdentatus Paulson, Archias sexdentatus Archias Paulson, 1875 
1875 


cribrarius Lamarck,  Portunus cribrarius Arenaeus Dana, 1851 
1818 

integerrimus Lamarck, Cancer integerrimus Atergatis de Haan, 
1818 [1833] 

signatus Adams & Carpilius signatus Atergatopsis Milne 
White, [1849] Edwards, 1862 

armata Milne Banareia armata Banareia Milne 


Edwards, 1869 Edwards, 1869 


Specific name 
proposed to be placed 
on the “Official List 
of Specific Names in 

Zoology ” 


(1) 
picta Milne Edwards, 
1848 
hemingi Alcock & 
Anderson, 1899 
rouxii Guérin- 
Ménéville, 1832 


maculatus Linnaeus, 
1758 


tristis Dana, 1852 

papulosus Stimpson, 
[1871] 

tenuipes Dana, 1852 

melanochirus Milne 
Edwards, 1873 


tridentatus Milne 
Edwards, 1879 


chilensis Milne 
Edwards & Lucas, 
[1844] 


pentagonus Stimpson, 
1 


ornatus Dana, 1852 

andreossyi Audouin, 
[1826] 

ermita Nobili, 1906 


imitatrix Hilgendorf, 
[1869] 

hispida Eydoux & 
Souleyet, 1842 


tuberosus Pennant 
1777 


cubensis Stimpson 


9 


> 


cessacii Milne 
Edwards, 1878 


DIRECTION 36 


Combination in which 
name specified in 
Col. 1 was originally 
published 


(2) 
Bellia picta 


Benthochascon 
hemingi 
Caphyra rouxii 


Cancer maculatus 


Carpilodes tristis 
Car poporus 
papulosus 
Carupa tenuipes 
Chlorodopsis 
melanochirus 


Coenophthalmus 
tridentatus 


Corystoides chilensis 


Cryptocnemus 
pentagonus 


Cyclodius ornatus 


Pilumnus andreossyi 

Dacryopilumnus 
eremita 

Deckenia imitatrix 

Domecia hispida 

Cancer tuberosus 

Epilobocera 


cubensis 
Epimelus cessacii 


31 


Name of genus of 
which species 
specified in Col. 1 
is the type species 


(3) 
Bellia Milne Edwards, 
1848 


Benthochascon Alcock 
& Anderson, 1899 


Caphyra Guérin- 
Meénéville, 1832 


Carpilius Desmarest, 
1823 


Carpilodes Dana, 1851 

Carpoporus Stimpson, 
[1871] 

Carupa Dana, 1851 

Chlorodopsis Milne 
Edwards, 1873 


Coenophthalmus 
Milne Edwards, 1879 


Corystoides Milne 
Edwards & Lucas 
[1844] 


Cryptocnemus 
Stimpson, 1858 


Cyclodius Dana, 1851 
Cymo de Haan, [1833] 


Dacryopilumnus 
Nobili, 1906 


Deckenia Hilgendorf, 
[1869] 


Domecia Eydoux & 
Souleyet, 1842 


Ebalia Leach, [1817] 


Epilobocera Stimpson, 
1860 


Epimelus Milne 
Edwards, 1878 


3D OPINIONS AND DECLARATIONS 


Specific name 
proposed to be placed 
on the ‘Official List 
of Specific Names in 

Zoology ”’ 


(1) 
isenbeckii Brandt, 
1848 


spinosus Rathbun, 
1894 


dovii Stimpson, 1860 


jaquemontii Milne 
Edwards, 1844 

amica Smith, 1869 

fallax Henderson, 
1893 


vesiculosa Alcock, 
1896 


rotundifrons Milne 
Edwards, 1867 

agilis Milne Edwards, 
1872 


sublobosa Stimpson, 
[1871] 


spongiosus Adams & 


White, [1849] 


cylindricus Fabricius, 
1777 


jurinei Saussure, 1853 


polybioides Adams & 
White, [1849] 


cumingii Bell, 1855 


rotundatus Adams & 
White, [1849] 


lambriformis Milne 
Edwards, 1873 


eudactylus Bell, 1855 
speciosa Dana, 1852 
hardwickii Leach, 1817 
dentata Bell, 1855 


Combination in which 


name specified in 


Col. 1 was originally 


published 


(2) 
Platycorystes 
(Podacanthus) 
isenbeckii 


Erimetopus 
Spinosus 


Euphylax dovii 


Gecarcinucus 
Jaquemontii 

Hepatella amica 

Ebalia fallax 


Heteronucia 
vesiculosa 


Heterozius 
rotundifrons 


Hydrothelphusa 
agilis 

Tliacantha 
subglobosa 


Iphiculus 
Spongtosus 
Cancer cylindricus 


Guaia (Ilia) jurinei 
Lissocarcinus 
polybioides 
Lithadia cumingii 
Lupocyclus 
rotundatus 


Merocryptus 
lambriformis 


Myrodes_ eudactylus 
Nucia speciosa 
Nursia_ hardwickii 
Nursilia dentata 


Name of genus of 
which species 
specified in Col. 1 
is the type species 


(3) 
Erimacrus Benedict, 
1892 


Erimetopus Rathbun, 
1894 


Euphylax Stimpson, 
1860 


Gecarcinucus Milne 
Edwards, 1844 
Hepatella Smith, 1869 


Heterolithadia Alcock, 
1896 


Heteronucia Alcock, 
1896 


Heterozius Milne 
Edwards, 1867 


Hydrothelphusa Milne 
Edwards, 1872 


Tliacantha Stimpson, 
[1871] 


Iphiculus Adams & 
White, [1849] 


Ixa Leach, 1815 


Leucosilia Bell, 1855 


Lissocarcinus Adams 
& White [1849] 


Lithadia Bell, 1855 


Lupocyclus Adams & 
White [1849] 


Merocryptus Milne 
Edwards, 1873 


Myrodes Bell, 1855 
Nucia Dana, 1852 
Nursia Leach, 1817 
Nursilia Bell, 1855 


Specific name 
proposed to be placed 
on the “Official List 
of Specific Names in 

Zoology” 


(1) 


lamelligera Stimpson, 
horridus Riippell, 1830 


tuberosa Stimpson, 
[1871] 


milne-edwardsii 
Miers, 1886 


tridentata Milne 
Edwards, 1853 


orientalis Miers, 1886 


alcocki Wood-Mason, 
1891 — 


coronata Alcock & — 
Anderson, 1894 


crassipes Bell, 1855 


denticulatus Montagu, 
1808 


gaudichaudii Milne 
Edwards, 1837 


henslowii Leach, [1820] 
latipes Pennant, 1777 


armatus Milne 
Edwards, 1853 


perlata Milne 
Edwards, 1837 


tridentata Miers, 1879 

americana Saussure, 
1857 

ornata Randall, 1840 


nodosus Bell, 1855 


DIRECTION 36 


Combination in which 
name specified in 
Col. 1 was originally 
published 


(2) 
Onychomorpha 
lamelligera 


Oreophorus 
horridus 


Osachila tuberosa 


Paracyclois 
milne-edwardsii 


Parathelphusa 
tridentata 


Lupocyclus 
(Parathranites) 
orientalis 


Parilia alcocki 
Randallia coronata 


Phlyxia crassipes 
Cancer denticulatus 


Platymera 
gaudichaudii 


Polybius henslowii 
Cancer latipes 


Potamocarcinus 
armatus 


Thelpheusa [sic] 
perlata 


Pseudophilyra 
tridentata 


Pseudothelphusa 
americana 


Ilia ornata 


Oreophorus nodosus 


33 


Name of genus of 
which species 
specified in Col. | 
is the type species 


(3) 
Onychomorpha 
Stimpson, 1858 


Oreophorus Riippell, 
1830 


Osachila Stimpson, 
[1871] 


Paracyclois Miers, 
1886 


Parathelphusa Milne 
Edwards, 1853 


Parathranites Miers, 
1886 


Parilia Wood-Mason, 
1891 


Pariphiculus Alcock, 
1896 


Phlyxia Bell, 1855 
Pirimela Leach, [1816] 


Platymera Milne 
Edwards, 1837 


Polybius Leach, [1820] 


Portumnus [Leach], 
[1814] 


Potamocarcinus Milne 
Edwards, 1853 


Potamonautes 
Macleay, 1838 


Pseudophilyra Miers, 
1879 


Pseudothelphusa 
Saussure, 1857 


Randallia Stimpson, 


Speloeophorus Milne 
Edwards, 1865 


34 


Specific name 
proposed to be placed 
on the “Official List 
of Specific Names in 

Zoology” 


(1) 
bedoti Zehnter, 1894 


serratus Forskal, 1775 
admete Herbst, [1803] 


quadridens Milne 
Edwards, 1869 


danae Milne Edwards, 
1869 


muriger Adams & 
White, [1849] 


corallinus Faxon, 1893 


fluviatilis Latreille, 
[1828] 


nobile Milne Edwards, 
1880 


serrata White, 1847 


parvula Stimpson, 1858 
inaequipes Stimpson, 


typhlus Milne 
Edwards, 1880 


sexdentatum Stimpson, 
1858 


coppingeri Miers, 1884 

nitidus Milne 
Edwards, 1870 

ciliatus Stimpson, 1858 


typicus Rathbun, 1898 


Combination in which 
name specified in 
Col. 1 was originally 
published 


(2) 
Sphaerocarcinus 
bedoti 
Cancer serratus 
Cancer admete 


Thalamitoides 
quadridens 


Goniosoma danae 
Tlos muriger 


Trachycarcinus 
corallinus 


Trichodactylus 
fluviatilis 

Trichopeltarion 
nobile 

Valdivia serrata 


Opinion 85 


Acmaeopleura 
parvula 


Asthenognathus 
inaequipes 
Bathyplax typhlus 


Camptandrium 
sexdentatum 


Camptoplax 
coppingeri 
Catoptrus nitidus 


Ceratoplax ciliatus 


Chasmocarcinus 
typicus 


OPINIONS AND DECLARATIONS 


Name of genus of 
which species 
specified in Col. | 
is the type species 


(3) 
Sphaerocarcinus 
Zehnter, 1894 
Scylla de Haan, [1833] 
Thalamita Latreille, 
1829 


Thalamitoides Milne 
Edwards, 1869 


Thalamonyx Milne 
Edwards, 1873 


Tlos Adams & White, 
[1849] 


Trachycarcinus Faxon, 


Trichodactylus 
Latreille, [1828] 


Trichopeltarion Milne 
Edwards, 1880 


Valdivia White, 1847 


Acmaeopleura 
Stimpson, 1858 


Asthenognathus 
Stimpson, 1858 


Bathyplax Milne 
Edwards, 1880 


Camptandrium 
Stimpson, 1858 


Camptoplax Miers, 
1884 


Catroptrus Milne 
Edwards, 1870 


Ceratoplax Stimpson, 
1858 


Chasmocarcinus 
Rathbun, 1898 


Specific name 
proposed to be placed 
on the “Official List 
of Specific Names in 

Zoology” 


(1) 
balansae Milne 
Edwards, 1873 


angulatus Dana, 1851 
nitidus Smith, 1870 
carinipes de Man, 1889 


liguricus Milne 
Edwards, 1853 


crenatus de Haan, 
[1835] 

agassizii Milne 
Edwards, 1880 


crassimanus Dana, 
1851 


deplanatus Cano, 1889 
nitidus Stimpson, 1859 


limosa Say, 1818 


subquadrata Dana, 
1851 


bispinosus Herbst, 
[1783] 


tridens Kreyer, 1837 
impressus Smith, 1870 
pugnax Smith, 1870 
bicornis Gray, 1831 
fragifer de Haan, 


[1835] 


tridens de Haan, 
[1835] 


DIRECTION 36 


Combination in which 
name specified in 
Col. 1 was originally 
published 


(2) 
Clistocoeloma 
balansae 


Cyrtograpsus 
angulatus 
Dissodactylus nitidus 


Durckheimia carinipes 


Euchirograpsus 
liguricus 


Cancer (Eucrate) 
crenatus 


Eucratodes 
agassiZil 
Eucrate crassimanus 


Euryetisus deplanatus 
Euryplax nitidus 


Cancer limosa 


Fabia subquadrata 


Cancer bispinosus 
Geryon tridens 


Glyptograpsus 
impressus 


Glyptoplax pugnax 
Gomeza_ bicornis 


Cancer (Halimede) 
fragifer 

Ocypode (Helice) 
tridens 


35 


Name of genus of 
which species 
specified in Col. 1 
is the type species 


(3) 
Clistocoeloma Milne 
Edwards, 1873 


Cryrtograpsus Dana, 
1851 


Dissodactylus Smith, 


Durckheimia de Man, 
1889 


Euchirograpsus Milne 
Edwards, 1853 


Eucrate de Haan, 
[1835] 


Eucratodes Milne 
Edwards, 1880 


Eucratopsis Smith, 
1869 


Euryetisus Cano, 1889 
Euryplax Stimpson, 
1859 


Eurytium Stimpson, 
1859 

Fabia Dana, 1851 

Galene de Haan, 
[1833] 

Geryon Kregyer, 1837 

Glyptograpsus Smith, 
1870 

Glyptoplax Smith, 
1870 

Gomeza Gray, 1831 

Halimede de Haan, 


[1835] 
Helice de Haan, [1833] 


36 OPINIONS AND DECLARATIONS 


Specific name 
proposed to be placed 
on the “Official List 
of Specific Names in 

Zoology ” 


(1) 
lugubris Alcock, 1899 


Combination in which 


name specified in 


Col. 1 was originally 


published 


(2) 
Hephthopelta 
lugubris 


sexpes Fabricius, 1798 Cancer sexpes 


haematocheir de Haan, Grapsus (Pachysoma) 


[1835] 
trapeziformis Nauck, 


rodgersii Stimpson, 
1858 


exaratus Milne 
Edwards, 1834 


rubromaculatus de 
Haan, [1833] 


nitidus Milne 
Edwards, 1867 


cinctimanus White, 
[1849] 


leeanus Rathbun, 1898 


bispinosa Kinahan, 
1856 


bella Stimpson, 1860 


dilatipes Adams & 
White, [1849] 


tesselatus Latreille, 
1812 


thalamita Stimpson, 
[1871] 


rousseauxi Milne 
Edwards, 1853 


truncatus Stimpson, 
1860 


sculptipes Stimpson, 
[1871] 


haematocheir 


Holothuriophilus 
trapeziformis 


Lachnopodus 
rodgersii 


Chlorodius exaratus 


Cancer (Liagore) 
rubromaculatus 


Libystes nitidus 


Car pilius 
cinctimanus 


Lipaesthesius 
leeanus 


Litocheira bispinosa 


Xantho bella 


Pilumnus dilatipes 
Grapsus tesselatus 
Melybia thalamita 


Metasesarma 
rousseauxi 


Metopocarcinus 
truncatus 

Micropanope 
sculptipes 


Name of genus of 
which species 
specified in Col. 1 
is the type species 


(3) 
Hephthopelta Alcock, 
1899 


Hexapus de Haan, 
[1833] 


Holometopus Milne 
Edwards, 1853 


Holothuriophilus 
Nauck, 1880 


Lachnopodus 
Stimpson, 1858 


Leptodius Milne 
Edwards, 1863 


Liagore de Haan, 
[1833] 


Libystes Milne 
Edwards, 1867 


Liomera Dana, 1851 


Lipaesthesius Rathbun, 
1898 


Litocheira Kinahan, 
1856 


Lophopanopeus 
Rathbun, 1898 


Lophopilumnus Miers, 


Lybia Milne Edwards, 
1834 

Melybia Stimpson, 
[1871] 


Metasesarma Milne 
Edwards, 1853 


Metopocarcinus 
Stimpson, [1860] 

Micropanope 
Stimpson, [1871] 


Specific name 
proposed to be placed 
on the “Official List 
of Specific Names in 

Zoology” 


(1) 
nitidus Milne Edwards, 
1873 


granulatus Rathbun, 


macgillivrayi White, 
1852 


transversus Rathbun, 
1893 


microps Alcock & 
Anderson, 1894 


depressa Stimpson, 
[1871] 


neapolitana Kossmann, 
1878 


euagora de Man, 1895 


roseus Nobili, 1906 


planissimus Herbst, 
[1804] 


excelsus Heller, 1862 


perlatus Poeppig, 1836 


hirtellus Linnaeus, 
1761 


hirtipes Heller, [1865] 
cylindricum Say, 1818 


laevigata Milne 
Edwards & Lucas 
[1844] 


pisum Linnaeus, 1767 


DIRECTION 36 


Combination in which 
name specified in 
Col. 1 was originally 
published 


(2) 


Notonyx nitidus 
Oediplax granulatus 
Ommatocarcinus 


macgillivrayi 
Opisthopus 

transversus 
Xanthodes microps 


Panoplax depressa 
Paragalene 
neapolitana 
Parapanope euagora 
Parapleurophrycoides 


roseus 
Cancer planissimus 


Perigrapsus excelsus 


Hepatus perlatus 


Cancer hirtellus 
Pinnaxodes hirtipes 
Pinnotheres 
cylindricum 
Pinnotherelia 


laevigata 


Cancer pisum 


37 


Name of genus of 
which species 
specified in Col. | 
is the type species 


(3) 
Notonyx Milne 
Edwards, 1873 


Oediplax Rathbun, 
1893 


Ommatocarcinus 
White, 1852 


Opisthopus Rathbun, 
1893 


Orphnoxanthus 
Alcock, 1898 

Panoplax Stimpson, 
[1871] 


Paragalene Kossman, 
1878 


Parapanope de Man, 
1895 

Parapleurophrycoides 
Nobili, 1906 


Percnon Gistl, 1848 


Perigrapsus Heller, 
1862 


Pilumnoides Milne 
Edwards & Lucas, 
[1844] 


Pilumnus Leach, 1815 


Pinnaxodes Heller, 
[1865] 


Pinnixa White, 1846 


Pinnotherelia Milne 
Edwards & Lucas 
[1844] 


Pinnotheres Latreille, 
[1802—1803] 


38 


Specific name 
proposed to be placed 
on the “Official List 
of Specific Names in 

Zoology” 


(1) 


spectabilis de Man, 
gracilipes Alcock, 1894 


orbignyi Milne 
Edwards & Lucas 
[1844] 


cupulifer Latreille, 
1825 


spinicarpus Milne 
Edwards, 1852 


gigas Lamarck, 1818 


carinata Ortmann, 
1894 


quadridentata 
Latreille, 1825 


stridulans Wood- 
Mason, 1892 


glaber Stimpson, 1858 


granulosus Milne 
Edwards, 1879 


harrisii Gould, 1841 


gracilipes Stimpson, 
1858 


convexus Milne 
Edwards, 1867 


crassum Dana, 1851 
spinosipes Stimpson, 


granulatus Rathbun, 
1893 


carolinensis Stimpson, 
1859 


Combination in which 


name specified in 
Col. 1 was originally 
published 


(2) 
Platychirograpsus 
spectabilis 


Platypilumnus 
gracilipes 
Xantho orbignyi 


Pilumnus cupulifer 


Prionoplax 
Spinicarpus 
Cancer gigas 


Pseudopinnixa 
carinata 


Melia 
quadridentata 


Psopheticus 
stridulans 


Ptychognathus glaber 


Pyxidognathus 
granulosus 


Pilumnus harrisii 
Rhizopa gracilipes 


Ruppellioides 
convexus 


Samartium crassum 

Scalopidia 
Spinosipes 

Scleroplax 
granulatus 


Speocarcinus 
carolinensis 


OPINIONS AND DECLARATIONS 


Name of genus of 
which species 
specified in Col. 1 
is the type species 


(3) 
Platychirograpsus de 
Man, 1896 


Platypilumnus Alcock, 
1894 


Platyxanthus Milne 
Edwards, 1863 


Polydectus Milne 
Edwards, 1837 


Prionoplax Milne 
Edwards, 1852 


Pseudocarcinus Milne 
Edwards, 1834 


Pseudopinnixa 
Ortmann, 1894 


Pseudorhombila 
Milne Edwards, 
1837 


Psopheticus Wood- 
Mason, 1892 


Ptychognathus 
Stimpson, 1858 


Pyxidognathus Milne 
Edwards, 1879 


Rhithropanopeus 
Rathbun, 1898 


Rhizopa Stimpson, 
1858 


Ruppellioides Milne 
Edwards, 1867 


Samartium Dana, 1851 
Scalopidia Stimpson, 
1858 


Scleroplax Rathbun, 
1893 


Speocarcinus 
Stimpson, 1859 


Specific name 
proposed to be placed 
on the ‘Official List 
of Specific Names in 

Zoology” 


(1) 
nitidus Stimpson, 1858 


bidentatus Milne 
Edwards, 1880 


scabripes Rathbun, 
1898 


anomalipes Miers, 1881 


gracilipes White, 1847 


litteratus Fabricius, 
1798 


murigera White, 1846 
sternberghii Stimpson, 


moebii Richters, 1880 


pinnotheroides White, 
1846 


aeneus Linnaeus, 1758 
carinipes Heller, 1861 


pagurus Linnaeus, 
1758 


norvegicus Linnaeus, 
1758 


annulicornis Leach, 
1815 


DIRECTION 36 


Combination in which 
name specified in 
Col. 1 was originally 
published 


(2) 


Sphaerozius nitidus 
Xanthodes bidentatus 
Tetrias scabripes 


Thaumastoplax 
anomalipes 


Utica gracilipes 
Cancer litteratus 


Xanthasia murigera 


Xanthodius 
sternberghii 


Xenophthalmodes 
moebii 


Xenophthalmus 
pinnotheroides 


Cancer aeneus 
Zozymodes carinipes 


Opinion 104 
Cancer pagurus 


Cancer norvegicus 


Pandalus 
annulicornis 


39 


Name of genus of 
which species 
specified in Col. | 
is the type species 


(3) 
Sphaerozius 
Stimpson, 1858 


Tetraxanthus Rathbun, 
1898 


Tetrias Rathbun, 1898 
Thaumastoplax Miers, 


Utica White, 1847 


Varuna Milne 
Edwards, 1830 


Xanthasia White, 1846 


Xanthodius Stimpson, 
1859 


Xenophthalmodes 
Richters, 1880 


Xenophthalmus White, 
1846 


Zosimus Leach, 1823 
Zozymodes Heller, 
1861 


Cancer Linnaeus, 1758 


Nephrops {Leach}, 
[1814] 


Pandalus Leach, 1815 


40 OPINIONS AND DECLARATIONS 


ANNEXE 2 


Specific names of nine nominal species which are subjectively identified 
with nominal species of later date which are the type species of 
genera, the names of which were placed on the ‘‘ Official 
List of Generic Names in Zoology ”’ in the period up to 
the end of 1936 


Specific name proposed to be 
placed on the <“ Official List 
of Specific Names in Zoology””. 


(1) 
savignyi (correction of savignii) 
Milne Edwards, 1834, as pub- 
lished in the combination 
Cancer savignii. 


punctatus Linnaeus, 1767, as 


published in the combination 
Cancer punctatus. 


vigil Weber, 1795, as published 
in the combination Portunus 
vigil. 


cheiragonus Tilesius, 1815, as 
published in the combination 
Cancer cheiragonus. 


Remarks 


(not for inclusion in the entry to 


be made in the Official List) 
(2) 


The name savignyi Milne Edwards, 


1834, is a senior subjective syno- 
nym of granulatus de Haan, 
[1835], as published in the 
combination Cancer (Actaea) 
granulatus, the specific name 
of the type species of Actaea 
de Haan, [1833]. 


The name punctatus Linnaeus, 


1767, is a senior subjective 
synonym of /atrei/li Leach, 1817, 
as published in the combination 
Persephona lIatreilli, the specific 
name of the type species of 
Persephona Leach, 1817. 


The name vigil Weber, 1795, is a 


senior subjective synonym of 
spinosus Lamarck, 1801, as pub- 
lished in the combination 
Podophthalmus [sic] spinosus, the 
specific name of the type species 
of Podophthalmus Lamarck, 
1801. 


The name cheiragonus Tilesius, 


1815, is a senior subjective 
synonym of serratus White, 
1846, as published in the com- 
bination Telmessus serratus, the 
specific name of the type species 
of Telmessus White, 1846. 


DIRECTION 36 4| 


Specific name proposed to be 
placed on the “ Official List 
of Specific Names in Zoology’’. 


(1) 


frontalis Milne Edwards, 1834, 
as published in the combina- 
tion Ozius frontalis. 


rhomboides Linnaeus, 1758, as 
published in the combination 
Cancer rhomboides. 


barbiger Poeppig, 1836, as pub- 
lished in the combination 
Gecarcinus barbiger. 


minutus Linnaeus, 1758, as pub- 
lished in the combination 
Cancer minutus. 


superba Costa, 1838, as pub- 
lished in the combination 
Portunus superba. 


Remarks 
(not for inclusion in the entry to 
be made in the Official List) 


(2) 


The name frontalis Milne Edwards, 
1834, is a senior subjective 
synonym of kotschii Heller, 1861, 
as published in the combination 
Epixanthus kotschii, the specific 
name of the type species of 
Epixanthus Heller, 1861. 


The name rhomboides Linnaeus, 
1758, is a senior subjective 
synonym of bispinosa Lamarck, 
1801, as published in the com- 
bination Ocypoda [sic] bispinosa, 
the specific name of the type 
species of Goneplax [Leach], 
(1814). 


The name barbiger Poeppig, 1836, 
is a senior subjective synonym of 
hirtipes Milne Edwards & Lucas, 
[1844], as published in the com- 
bination Paraxanthus hirtipes, 
the specific name of the type 
species of Paraxanthus Milne 
Edwards & Lucas, [1844]. 


The name minutus Linnaeus, 1758, 
is a senior subjective synonym 
of clypeatus Bowditch, 1825, as 
published in the combination 
Planes clypeatus, the specific 
name of the type species of 
Planes Bowditch, 1825. 


The name superba Costa, 1838, is 
a senior subjective synonym of 
longispina Stimpson, [1871], as 
published in the combination 
Bathynectes longispina, the 
specific name of the type species 
of Bathynectes Stimpson, [1871]. 


42 OPINIONS AND DECLARATIONS 


ANNEXE 3 


Correction of the date given for certain specific names in 
‘** Opinions ’’ 73 and 85 


Specific Name Date cited in Corrected 
concerned the *‘ Opinion”? date 
(1) (2) (3) 
Opinion 73 


rouxii Guérin-Ménéville, as pub- 
lished in the combination Caphyra 


rouxii 1830 1832 
amica Smith, ‘as published in the 
combination Hepatella amica .. 1870 1869 


signatus Adams & White, as pub- 

lished in the combination C Has 

signatus.. 1848 [1849] 
spongiosus Adams & White, as pub- 

lished in the combination [phiculus . 

spongiosus . . ap a oS 1848 [1849] 
polybioides Adams & White, as 

published in the combination 

Lissocarcinus polybioides 1848 [1849] 
rotundatus Adams & White, as 

published in_ the combination 

Lupocyclus rotundatus.. 1848 [1849] 
muriger Adams & White, as pub- 

lished in the combination T/os 

muriger 1848 [1849] 
fluviatilus Latreille, as s published i in 

the combination Trichodactylus 

fluviatilis .. efi ne Be 1825 [1828] 


Opinion 85 


cinctimanus Adams & White, as 

published in the combination 

Carpilius cinctimanus _.. 1848 [1849] 
laevigata Milne Edwards & Lucas, 

as published in the combination 

Pinnotherelia laevigata .. 1843 [1844] 
orbignyi Milne Edwards & Lucas, 

as published in the combination 


Xantho orbignyi .. 1843 [1844] 
tesselatus Latreille, as pubtghea in 
the combination Gr apsus tesselatus 1818 1812 


2. Registration of the present application : Upon the receipt 
of Mr. Hemming’s paper, the question of placing on the Official 


DIRECTION 36 43 


List of Specific Names in Zoology the specific names of species 
of the Order Decapoda specified therein was allotted the 
Registered Number Z.N.(S.) 985. 


IlL—THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


3. Issue of Voting Paper V.P.(O.M.)(55)27 : On 19th September 
1955 a Voting Paper (V.P.(O.M.)(55)27) was issued in which 
each Member of the Commission was asked (1) to state whether 
he agreed “that, in conformity with the General Directives 
issued to the International Commission on Zoological Nomen- 
clature by the Thirteenth International Congress of Zoology, 
Paris, 1948, regarding the placing on the appropriate Official 
Lists and Official Indexes of names dealt with in, or otherwise 
involved in, Opinions rendered prior to 1948, the names referred 
to in Sections (a) and (b) of paragraph 9 of the paper bearing 
the Registered Number Z.N.(S.) 985 submitted by the Secretary 
simultaneously with the present Voting Paper [i.e. in the 
paragraph numbered as above in the paper reproduced in the 
first paragraph of the present Direction] should be placed on the 
Official List of Specific Names in Zoology as there recommended 
and that the other action recommended in the said paragraph be 
similarly approved ” and (2), if he did not so agree as regards any 
given item, to indicate that item. 


4. Withdrawal of the proposals in regard to three of the specific 
names submitted with Voting Paper V.P.(O.M.)(55)27 in order to 
permit of a further study of the issues involved in regard thereto : 
On 7th October 1955 Mr. Hemming, as Secretary, placed on the 
Commission’s File Z.N.(S.) 985 a Minute withdrawing from 
the scope of Voting Paper V.P.(O.M.)(55)27 the proposals 
submitted in regard to three of the specific names dealt with therein, 
in order to permit of a re-examination of the issues involved in 
the light of additional information which had just been received. 
In the same Minute Mr. Hemming drew attention also to the 
inadvertent omission from the proposals submitted with the 
foregoing Voting Paper of a recommendation for the addition to 


44 OPINIONS AND DECLARATIONS 


the Official Index of Rejected and Invalid Specific Names in 
Zoology of an Invalid Original Spelling of one of the specific 
names recommended therein for addition to the Official List 
of Specific Names in Zoology. Mr. Hemming’s Minute was in the 
following terms :— 


Withdrawal of proposals relating to three specific names from the 
scope of Voting Paper V.P.(O.M.)(55)27 


MINUTE by FRANCIS HEMMING, C.M.G., C.B.E., 
(Secretary to the International Commission on Zoological Nomenclature) 


In the case of three specific names on which proposals were submitted 
with Voting Paper V.P.(O.M.)(55)27 additional information now 
furnished by Dr. L. B. Holthuis calls for further study of the action 
required in these cases before decisions are taken thereon by the Com- 
mission. The names concerned are the specific names of the type 
species of the following genera: (1) Portumnus [Leach], [1814]; 
(2) Pinnotheres Latreille, [1802—1803] ; (3) Pandalus Leach, 1815. 


2. Accordingly, as Secretary to the Commission, I hereby withdraw 
from the scope of the foregoing Voting Paper the proposals submitted 
therein in regard to the specific names referred to above. I further 
direct that the issues which have now come to light in regard to these 
names be studied on the under-mentioned Registered Files and that 
proposals regarding these names be submitted to the Commission as 
soon as possible :—(a) Z.N.(S.) 1028 (Pandalus)?; (b) Z.N.(S.) 1029 
(Pinnotheres)® ; (c) Z.N.(S.) 1031 (Portumnus).’ 


3. I take this opportunity to draw attention to the fact that through 
some inadvertence the proposal submitted on the foregoing Voting 
Paper for the addition to the Official List of Specific Names in Zoology 
of the specific name savignyi Milne Edwards (H.), as published in the 
combination Cancer savignii, was not accompanied by a corresponding 
recommendation for the addition of the Invalid Original Spelling 
savignii to the Official List of Rejected and Invalid Specific Names in 
Zoology. Having regard to the fact that in a case such as this the 
addition of the above name to the Official Index is obligatory under the 
General Directives issued to the Commission by the International 
Congresses of Zoology, I, as Secretary, hereby direct that a proposal 
in the foregoing sense be ‘deemed to have been included in the Voting 
Paper referred to above. 


5 A decision has since been taken on this case and has been embodied in Direction 
47, which will be published in the present Section (Section D) of Volume 1. 

6 A decision has since been taken on this case and has been embodied in Direction 
45, which will be published in the present Section of Volume 1. 

* An application for the settlement of this case by the use of the Commission’s 
Plenary Powers has since been published (Holthuis, 1956, Bull. zool. Nomencl. 
12 : 99—105 ; Hemming, 1956, ibid, 12 : 105—106). 


DIRECTION 36 45 


5. The Prescribed Voting Period: As the foregoing Voting 
Paper was issued under the One-Month Rule, the Prescribed 
Voting Period closed on 19th October 1955. 


6. Particulars of the Voting on Voting Paper V.P.(O.M.)(55)27 : 
At the close of the Prescribed Voting Period, the state of the 
voting on Voting Paper V.P.(O.M.)(55)27 was as follows :— 


(a) Affirmative Votes had been given by the following twenty-four 
(24) Commissioners (arranged in the order in which Votes 
were received) : 


Boschma ; Holthuis; Stoll; Mayr; Riley ; Sylvester- 
Bradley ; Vokes ; Hering ; Jaczewski ; Prantl ; Lemche ; 
Mertens ; Bonnet ; do Amaral ; Hanko ; Esaki; Key ; 
Dymond; Cabrera; Hemming; Miller®; Bradley ; 
(J.C.) ; Kiihnelt ; Tortonese ; 


(b) Negative Votes : 
None ; 


(c) On Leave of Absence, one (1) : 
Bodenheimer ; 


(d) Voting Papers not returned : 
None. 


7. Declaration of Result of Vote: On Ist November 1955, 
Mr. Hemming, Secretary to the International Commission, acting 
as Returning Officer for the Vote taken on Voting Paper 
V.P.(O.M.)(55)27, signed a Certificate that the Votes cast were 
as set out in paragraph 6 above and declaring that the proposal 
submitted in the foregoing Voting Paper had been duly adopted 
and that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


8. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 22nd December 1955, Mr. Hemming prepared the Ruling 
given in the present Direction and at the same time signed a 


8 Commissioner Miller exercised in this case the right conferred by the Thirteenth 
International Congress of Zoology in Paris, 1948, under which a Commissioner 
may, if he so desires, signify his willingness to support the view or the majority 
view, of other members of the Commission (1950, Bull. zool. Nomencl. 4 : 50—S51). 


46 OPINIONS AND DECLARATIONS © 


Certificate that the terms of that Ruling were in complete accord 
with those of the proposal approved by the International 
Commission in its Vote on Voting Paper V.P.(O.M.)(55)27, subject 
(a) to the withdrawal therefrom of the proposals relating to the 
three specific names specified in the Minute executed by the 
Secretary on 7th October 1955, and (b) to the inclusion in the 
Official Index of Rejected and Invalid Specific Names in Zoology 
of a specific name specified in the same Minute which had been 
rejected by the Commission as being objectively invalid in its 
Vote on the above Voting Paper. The text of the Minute 
executed by the Secretary on 7th October 1955 has been repro- 
duced in paragraph 4 of the present Direction. 


9. Original references : The original references for the specific 
names placed on the Official List and Official Index by the Ruling 
given in the present Direction have already been noted in con- 
nection with the Official List of Generic Names in Zoology, the 
nominal species bearing the names concerned being either 
(i) the type species of genera, the names of which have been 
inscribed on that List or (ii) or nominal species which are 
currently subjectively identified as representing the same taxa 
as those represented by nominal species which are the type 
species of such genera. 


10. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in 
dealing with the present case, and the present Direction is 
accordingly hereby rendered in the name of the said International 
Commission by the under-signed Francis Hemming, Secretary to 
the International Commission on Zoological Nomenclature, in 
virtue of all and every the powers conferred upon him in that 
behalf. 


11. The present Direction shall be known as Direction Thirty- 
Six (36) of the International Commission on Zoological 
Nomenclature. 


Done in London, this Twenty-Second day of December, 
Nineteen Hundred and Fifty-Five. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


eS ee ee 
Printed in England by MretcaLtFe & Cooper LimiTeED, 10-24 Scrutton St., London EC 2 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, C.M.G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.2. Pp. 47—82 


DIRECTION 37 


Completion of the entries relating to the names of certain 
genera of the Order Decapoda (Class Crustacea) placed 
on the Official List of Generic Names in Zoology in the 
period up to the end of 1936 


he, 


— 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956 


Price Twenty-four Shillings 
(All rights reserved) 


Issued \st September, 1956 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 37 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JORDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President: Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., 
U.S.A.) (12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary ;: Mr. Francis HEMMING (London, England) (27th July 1948) 


_B. The Members of the Commission 


(Arranged in order of precedence by reference to date of election or of most recent 
re-election, as prescribed by the International Congress of Zoology) 


Professor H. BoscHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. Henning LeMcHE (Universitetets Zoologiske Museum, Copenhagen, Denmark) 
(27th July 1948) 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEy (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZESWKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (5th July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitat zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyYMOND (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N. Y., U.S.A.) (12th August 1953) 
(President) 

Professor Harold E. Voxes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) 
(12th August 1953) 

Professor Béla HANKO (Mezégazdasdgi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y., U.S.A.) 
(12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. Hottuurs (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (15th October 1954) 

Dr. Alden H. MiILLer (Museum of Vertebrate Zoology, University of California, California, 
U.S.A.) (29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum v Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KiiNHELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

Professor F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 


Professor Ernst MAyR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico TORTENESE (Museo di Storia Naturale “‘G. Doria’’, Genova, Italy) 
(16th December 1954) 


DIRECTION 37 


COMPLETION OF THE ENTRIES RELATING TO THE 
NAMES OF CERTAIN GENERA OF THE ORDER 
DECAPODA (CLASS CRUSTACEA) PLACED ON THE 
“OFFICIAL LIST OF GENERIC NAMES IN 
ZOOLOGY”? IN THE PERIOD UP TO THE 
END OF 1936 


RULING :—(1) The following particulars relating to the 
_ determination under Article 30 of the Régles of the type 
species of the under-mentioned genera of the Order 
Decapoda (Class Crustacea) the names of which were 
placed on the Official List of Generic Names in Zoology 
by the Ruling given in Opinion 85, are hereby inserted 
in the said Official List :— 


(a) Type species by original designation (Rule (a)) : 

(1) Chasmocarcinus Rathbun, 1898 ; 

(2) Eurytium Stimpson, 1859 ; 

(3) Geryon Kroyer, 1837; 

(4) Glyptograpsus Smith (S.I.), 1870 ; 

(5) Goneplax [Leach], [1814] ; 

(6) Lophopanopeus Rathbun, 1898 ; 

(7) Lophopilumnus Miers, 1886 ; 

(8) Pseudorhombila Milne Edwards (H.), 1837 ; 
(b) Type species by original aoe (Rule (a)) 

through the formula “n.g., n.sp.’ 

(1) Durckheimia de Man, 1889 ; 

(2) Euryetisus Cano, 1889 ; 

(3) Holothuriophilus Nauck, 1880 ; 


ISEP © © 1956 


OPINIONS AND DECLARATIONS 


(4) Pinnaxodes Heller, [1865] ; 

(5) Platychirograpsus de Man, 1896 ; 
(6) Platypilumnus Alcock, 1894 ; 

(7) Thaumastoplax Miers, 1881 ; 

(8) Xenophthalmodes Richters, 1880 ; 


(c) Type species by monotypy (Rule (c)) : 
(1) Acmaeopleura Stimpson, 1858 ; 
(2) Asthenognathus Stimpson, 1858 ; 
(3) Bathyplax Milne Edwards (A.), 1880 ; 
(4) Camptandrium Stimpson, 1858 ; 
(5) Camptoplax Miers, 1884 ; 
(6) Catoptrus Milne Edwards (A.),-. 1870 ; 
(7) Ceratoplax Stimpson, 1858 ; 
(8) Chasmagnathus de Haan, [1833] ; 
(9) Clistocoeloma Milne Edwards (A.), 1873 ; 
(10) Cyrtograpsus Dana, 1851 ; 
(11) Dissodactylus Smith (S.I.), 1870 ; 
(12) Epixanthus Heller, 1861 ; 
(13) Euchirograpsus Milne Edwards (H.), 1853 ; 
(14) Eucrate de Haan, [1835] ; 
(15) Eucratodes Milne Edwards (A.), 1880 ; 
(16) Eucratopsis Smith (S.1.), 1869 ; 
(17) Euryplax Stimpson, 1859 ; 
(18) Fabia Dana, 1851 ; 
(19) Galene de Haan, [1833] ; 
(20) Glyptoplax Smith (S.I.), 1870 ; 
(21) Gomeza Gray (J.E.), 1831 ; 


DIRECTION 37 51 


(22) Halimede de Haan, [1835] ; 

(23) Helice de Haan, [1833] ; 

(24) Hephthopelta Alcock, 1899 ; 

(25) Hexapus de Haan, [1833] ; 

(26) Holometopus Milne Edwards (H.), 1853 ; 
(27) Lachnopodus Stimpson, 1858 ; 

(28) Leptodius Milne Edwards (A.), 1863 ; 
(29) Liagore de Haan, [1833] ; 

(30) Libystes Milne Edwards (A.), 1867 ; 

(31) Liomera Dana, 1851 ; 

(32) Lipaesthesius Rathbun, 1898 ; 

(33) Litocheira Kinahan, 1856 ; 

(34) Lybia Milne Edwards (H.), 1834 ; 

(35) Melybia Stimpson, [1871] ; 

(36) Metasesarma Milne Edwards (H.), 1853 ; 
(37) Metopocarcinus Stimpson, [1860] ; 

(38) Micropanope Stimpson, [1871] ; 

(39) Notonyx Milne Edwards (A.), 1873 ; 
(40) Oediplax Rathbun, 1893 ; 

(41) Ommatocarcinus White, 1852 ; 

(42) Opisthopus Rathbun, 1893 ; 

(43) Orphnoxanthus Alcock, 1898 ; 

(44) Panoplax Stimpson, [1871] ; 

(45) Paragalene Kossmann, 1878 ; 

(46) Parapanope de Man, 1895 ; 

(47) Parapleurophrycoides Nobili, 1906 ; 

(48) saan Ne Milne Edwards (H.) & Lucas, 


3 


OPINIONS AND DECLARATIONS 


(49) Perigrapsus Heller, 1862 ; 


(50) Pilumnoides Milne Edwards (H.) & Lucas, 
[1844] ; 


(51) Pilumnus Leach, 1815 ; 
(52) Pinnixa White, 1846 ; 


(53) Pinnotherelia Milne Edwards (H.) & Lucas, 
[1844] ; 


(54) Planes Bowditch, 1825 ; 

(55) Platyxanthus Milne Edwards (A.), 1863 ; 
(56) Polydectus Milne Edwards (H.), 1837 ; 
(57) Prionoplax Milne Edwards (H.), 1852 ; 
(58) Pseudopinnixa Ortmann, 1894 ; 

(59) Psopheticus Wood-Mason, 1892 ; 

(60) Ptychognathus Stimpson, 1858 ; 

(61) Pyxidognathus Milne Edwards (A.), 1879 ; 
(62) Rhithropanopeus Rathbun, 1898 ; 

(63) Rhizopa Stimpson, 1858 ; 

(64) Ruppellioides Milne Edwards (A.), 1867 ; 
(65) Samartium Dana, 1851 ; 

(66) Scalopidia Stimpson, 1858 ; 

(67) Scleroplax Rathbun, 1893 ; 

(68) Speocarcinus Stimpson, 1859 ; 

(69) Sphaerozius Stimpson, 1858 ; 

(70) Tetraxanthus Rathbun, 1898 ; 

(71) Tetrias Rathbun, 1898 ; 

(72) Utica White, 1847 ; 

(73) Varuna Milne Edwards (H.), 1830 ; 

(74) Xanthasia White, 1846 ; 


DIRECTION 37 53 


(75) Xanthodius Stimpson, 1859 ; 
(76) Xenophthalmus White, 1846 ; 
(77) Zosimus Leach, 1823 ; 

(78) Zozymodes Heller, 1861 ; 


(d) Type species by subsequent selection (Rule (g)) : 
(1) aS Gistl, 1848 : by selection by Rathbun 
l ° 


oe b 


(2) Pseudocarcinus Milne Edwards (H.), 1834 : 
by selection by Miers (1886). 


(2) The type species of the under-mentioned genera were 
selected under Rule (g)in Article 30 by Miss Mary Rathbun 
in the application which formed the basis of, and which 
was reproduced in, Opinion 73 published in 1922 (Smith- 
son. misc. Coll. 73 (No. 1)) and it is hereby directed that 
a note to the above effect be entered in the Official List 
of Generic Names in Zoology, the page references for the 
type selections so made being as specified below :— 

(a) Actaea de Haan, [1833] (: 26) ; 

(b) Actumnus Dana, 1851 (: 26) ; 

(c) Atergatis de Haan, [1833] (: 26) ; 

(d) Chlorodopsis Milne Edwards (A.), 1873 (: 26) ; 

(e) Cyclodius Dana, 1851 (: 26) ; 

(f) Ebalia Leach, [1817] (: 26) ; 

(g) Lliacantha Stimpson, [1871] (: 27) ; 

(h) Pariphiculus Alcock, 1896 (: 27) ; 

(1) Persephona Leach, 1817 (: 28) ; 

(j) Phlyxia Bell, 1855 (: 28) ; 

(k) Pseudophilyra Miers, 1879 (: 28) ; 

(1) Speloeophorus Milne Edwards (A.), 1865 (: 28) ; 


54 OPINIONS AND DECLARATIONS 


(m) Thalamitoides Milne Edwards (A.), 1869 (: 28) ; 
(n) Thalamonyx Milne Edwards (A.), 1873 (: 28). 


(3) It is hereby directed that the corrections specified 
below be made in the entries on the Official List of Generic 
Names in Zoology relating to the under-mentioned generic 
names made by the Rulings given in the Opinions severally 
cited below :— 


(a) Bathynectes Stimpson (Opinion 73): The date for 
this name is to be cited as [1871] and not as 1871. 


(b) Bathynectes Stimpson, [1871] (Opinion 73): The 
incorrect statement that Bathynectes longispina 
Stimpson, [1871], became the type species of the 
above genus by original designation is to be deleted 
and to be replaced by the entry that this species 
became the type species by monotypy. 


(c) Caphyra Guérin-Ménéville (Opinion 73): The in- 
correct entries relating to the date and place of 
publication of the above generic name and its 
type species are to be deleted and the following 
revised entries are to be substituted therefor :-— 


(a) Caphyra Guérin-Ménéville, 1832, Ann. Sci. 
nat., Paris 25(99) : 285 ; 


(b) Caphyra rouxii Guérin-Meénéville, 1832, Ann. 
Sci. nat., Paris 25(99) : 287. 


(d) Hepatella Smith and Hepatella amica Smith (Opin- 
ion 73): The incorrect entry “1870” relating 
to the date of publication of the above names is 
to be deleted and the date “ July 1869 ” (printed 
on the Part (Part 5) of volume 3 of the American 
Naturalist in which these names were published) 
is to be substituted therefor. 


(e) Five generic names by Bell (T.) published in 1855 
(Opinion 73) : The under-mentioned generic names 


DIRECTION 37 55 


by Bell (T.) were first published in 1855 not in 
volume 21 of the Transactions of the Linnean 
Society of London (the reference cited in Opinion 
73) but in volume 2 of the Proceedings of the 
above Society, and the entries relating to these 
names are to be amended accordingly, the pages 
on which these names appeared being as specified 
below :— 


(a) Leucosilia Bell, 1855 (: 431) ; 
(b) Lithadia Bell, 1855 (: 434) ; 
(c) Myrodes Bell, 1855 (: 432) ; 
(d) Nursilia Bell, 1855 (: 434) ; 
(e) Phlyxia Bell, 1855 (: 433). 


(f) Speloeophorus Milne Edwards (A.), 1865 (Opinion 
73): For reasons similar to those specified in 
relation to the generic names enumerated in (e) 
above the incorrect entry (Trans. linn. Soc. Lond. 
21) given for the name Oreophorus nodosus 
Bell (T.), 1855 (name of type species of Speloeo- 
phorus Milne Edwards (A.), 1865) is to be deleted 
and the following revised entry to be inserted 
in its place :—Oreophorus nodosus Bell (T.), 1855, 
Proc. linn. Soc. Lond. 2 : 434. 


(g) Correction of date attributed to certain names 
published in Part 2 of the Crustacea Section of 
Arthur White's “‘ The Voyage of H.M.S. Sama- 
rang’? (Opinion 73): The date “1848” in- 
correctly attributed to the under-mentioned names 
published in Part 2 of the Crustacea Section of 
White (A.), The Voyage of H.M.S. Samarang 
(which commenced at page 33) is to be deleted 
and the date “1849” is to be inserted in its 
place :— 


(a) Carpilius signatus Adams & White, [1849] 
(: 37) (type species of Atergatopsis Milne 
Edwards (A.), 1862) ; 


OPINIONS AND DECLARATIONS 


(6) Iphiculus Adams & White, [1849] (: 57) ; 


(c) Iphiculus spongiosus Adams & White, [1849] 
(: 57) (type species of Iphiculus Adams & 
White, [1849]) ; 


(d) Lissocarcinus Adams & White, [1849] (: 45) ; 


(e) Lissocarcinus polybioides Adams & White, 
[1849] (: 46) (type species of Lissocarcinus 
Adams & White, [1849]) ; 


(f) Lupocyclus Adams & White, [1849) (: 46) ; 


(g) Lupocyclus rotundatus Adams & White, [1849] 
(: 47) (type species of Lupocyclus Adams & 
White, [1849]) ; 


(h) Tlos Adams & White, [1849] (: 57) ; 


(i) Tlos muriger Adams & White, [1849] (: 58) 
(type species of ZJ/os Adams & White, 
[1849]). 


(h) Trichodactylus Latreille (Opinion 73) : The incorrect 
entry of the date of publication of the above 
name as “ 1825” is to be deleted and the date 
** 1828 ” (the date of publication of Part 2 of the 
Crustacea Section of the Ency. méth., in which 
this name first appeared) is to be inserted in its 
place. 


(i) Cymo de Haan, [1833] (Opinion 73) : The incorrect 
entry of the date of publication of the name 
Pilumnus andreossyi Audouin (the type species 
of Cymo,de, Haan,. (1833), as - le2>, 5 18 10.4be 
deleted and the date ‘‘ [1826] ”’ is to be inserted 
in its place. 


(j) Scylla de Haan, [1833] (Opinion 73) : The incorrect 
entry that Cancer serratus Forskal, 1775, is the 
type species of Scylla de Haan, [1833], by mono- 
typy is to be deleted and is to be replaced by an 
entry that the above species is the type species 


DIRECTION 37 a7 


of the foregoing genus by selection by Rathbun 
(1922, Smithson. misc. Coll. 73 (No. 1) : 28). 


(k) Correction of date attributed to certain names 
published in the Crustacea Section of d’ Orbigny’s 
‘“* Voyage dans Il’ Amerique Méridionale ”’ (Opinion 
85): The date “‘ 1843’ incorrectly attributed to 
the under-mentioned names published in the 
Crustacea Section of d’Orbigny’s Voyage dans 
l’ Amerique Méridionale is to be deleted and the 
date ‘‘ [1844] ” inserted in its place :-— 


(a) Paraxanthus Lucas, [1844] ; 
(b) Pilumnoides Lucas, [1844] ; 
(c) Pinnotherelia Lucas, [1844] ; 


(1) Nephrops [Leach] (Opinion 104): The incorrect 
entry of the date of publication of the above name 
as “1815” is to be deleted and the date “1814” 
(in which year the foregoing name was published 
in an anonymous paper of Leach’s in Part 2 
of volume 7 of Brewster’s Edinburgh Encyclo- 
paedia) is to be inserted in its place, the revised 
entry to be made being as follows :—Nephrops 
[Leach], [1814], in Brewster’s Edinburgh Ency. 
7(2) : 398—400. 


(4) The under-mentioned generic names are hereby 
placed on the Official Index of Rejected and Invalid Generic 
Names in Zoology with the Name Numbers severally 
specified below :— 


(i) Acanthocyclus Dybowski, 1873 (a junior homo- 
nym of Acanthocyclus Milne Edwards (H.) & 
Lucas, [1844]) (Name No. 461) ; 


(11) Acanthopus Oken, 1816 (a junior homonym of 
Acanthopus Klug, 1807) (Name No. 462) ; 


(iii) Acanthopus Dahl, 1823 (a junior homonym of 
Acanthopus Klug, 1807) (Name No. 463) ; 


58 OPINIONS AND DECLARATIONS 


(iv) Acanthopus Latreille, 1829 (a junior homonym 
of Acanthopus Klug, 1807) (Name No. 464) ; 


(v) Acanthopus de Haan, [1835] (a junior homonym 
of Acanthopus Klug, 1807) (Name No. 465) ; 


(vi) Acanthopus Muenster, 1839 (a junior homonym 
of Acanthopus Klug, 1807) (Name No. 466) ; 


(vii) Acanthopus Giebel, 1872 (a junior homonym of 
Acanthopus Klug, 1807) (Name No. 467) ; 


(viii) Acanthopus Vernet, 1877 (a junior homonym of 
Acanthopus Klug, 1807) (Name No. 468) ; 


(ix) Asthenognathus Bocourt, 1884 (a junior homo- 
nym of Asthenognathus Stimpson, 1858 (Name 
No. 469) ; 


(x) Asthenognathus Broom, 1915 (a junior homonym 
of Asthenognathus Stimpson, 1858) (Name No. 
470) ; 


(xi) Bathynectes Guenther, 1878 (a junior homonym 
of Bathynectes Stimpson, [1871] (Name No. 
471) ; 


(xil) Bathynectes Brinkman, 1913 (a junior homonym 
of Bathynectes Stimpson, [1871] (Name No. 
472) ; 


(xiii) Bellia Bate, 1851 (a junior homonym of Bellia 
Milne Edwards (H.), 1848) (Name No. 473) ; 


(xiv) Bellia Gray (J.E.), 1869 (a junior homonym of 
Bellia Milne Edwards (H.), 1848) (Name No. 
474) ; 


(xv) Bellia Tutt, 1902 (a junior homonym of Bellia 
Milne Edwards (H.), 1848) (Name No. 475) ; 


(xvi) Caphyra Barrande, 1846 (a junior homonym of 
Caphyra Guérin-Ménéville, 1832 (Name No. 
476) ; 


(xvii) Chlorodopsis Milne Edwards (A.), 1863 (a nomen 
nudum) (Name No. 477) ; 


DIRECTION 37 59 


(xviii) Cyrtograpsus Carruthers, 1867 (a junior homo- 
nym of Cyrtograpsus Dana, 1851) (Name No. 
8 . 


9 


(xix) Eucrate Rafinesque, 1815 (a nomen nudum) 
(Name No. 479) ; 


(xx) Euryplax Klapalek, 1909 (a junior homonym of 
Euryplax Stimpson, 1859) (Name No. 480) ; 


(xxl) Gecarcinicus Dana, 1852 (an Erroneous Subse- 
quent Spelling for Gecarcinucus Milne 
Edwards (H.), 1844) (Name No. 481) ; 


(xxil) Goneplat [Leach], [1814] (an Invalid Original 
Spelling for Goneplax [Leach], [1814]) (Name 
No. 482) ; 


(xxiil) Gonoplax Leach, 1815 (an Invalid Emendation 
of Goneplax [Leach], [1814]) (Name No. 483) ; 


(xxiv) Halimede Rathke, 1843 (a junior homonym of 
Halimede de Haan, [1835]) (Name No. 484) ; 


(xxv) Halimede Oberthuer & Houlbert, 1922 (a junior 
homonym of Halimede de Haan, [1835)]) 
(Name No. 485) ; 


(xxvil) Helice Chambers, 1873 (a junior homonym of 
Helice de Haan, [1833]) (Name No. 486) ; 


(xxvil) Holometopus Angelin, 1854 (a junior homonym 
of Holometopus Milne Edwards (H.), 1853) 
(Name No. 487) ; 


(xxviii) Homalaspis Brunner von Wattenwyl, 1895 (a 
junior homonym of MHomalaspis Milne 
Edwards (A.), 1863) (Name No. 488) ; 


(xxix) Homalaspis Kaier, 1932 (a junior homonym of 
Homalaspis Milne Edwards (A.), 1863) (Name 
No. 489) ; 


(xxx) Homarus Broun, 1881 (a junior homonym of 
Homarus Weber, 1795) (Name No. 490) ; 


60 OPINIONS AND DECLARATIONS 


(xxx) Iphiculus White, 1847 (a nomen nudum) (Name 
No. 491); 


(xxxil) Iphis Koch (C.L.), 1835 (a junior homonym of 
Iphis Leach, 1817) (Name No. 492) ; 


(xxxill) Jphis Laporte, 1836 (a junior homonym of Iphis 
Leach, 1817) (Name No. 493) ; 


(xxxiv) Liagora Gistl, 1848 (an Erroneous Subsequent 
Spelling of Liagore de Haan, [1833]) (Name 
No. 494) ; 

(xxxv) Lissocarcinus Adams & White, 1847 (a nomen 
nudum) (Name No. 495) ; 


(xxxvi) Lupocyclus White, 1847 (a nomen nudum) (Name 
No. 496) ; 


(xxxvil) Persephona Leach, 1852 (a junior homonym of 
Persephona Leach, 1817) (Name No. 497) ; 


(xxxvill) Pilumnus Rafinesque, 1815 (a junior homonym 
of Pilumnus Leach, 1815) (Name No. 498) ; 


(xxxix) Pilumnus Dejean, 1821 (a junior homonym of 
Pilumnus Leach, 1815) (Name No. 499) ; 


(xl) Pilumnus Koch (C.L.), 1837 (a junior homonym 
of Pilumnus Leach, 1815) (Name No. 500) ; 


(xli) Pilumnus Gistl, 1848 (a junior homonym of 
Pilumnus Leach, 1815) (Name No. 501) ; 


(xlii) Pilumnus Bonaparte, 1854 (a junior homonym 
of Pilumnus Leach, 1815) (Name No. 502) ; 


(xliii) Planes Saussure, 1862 (a junior homonym of 
Planes Bowditch, 1825) (Name No. 503) ; 


(xliv) Planes Rondani, 1864 (a junior homonym of 
Planes Bowditch, 1825) (Name No. 504) ; 


(xlv) Podacanthus Brandt, 1848 (a junior homonym of 
Podacanthus Gray (J.E.), 1833) (Name No. 
505 


b) 


DIRECTION 37 61 


(xlvi) Podophtalmus Lamarck, 1801 (an Invalid Original 
Spelling for Podophthalmus Lamarck, 1801) 
(Name No. 506) ; 
(xlvii) Polydectus Rafinesque, 1815 (a nomen nudum) 
(Name No. 507) ; 
(xlviii) Pseudopinnixa Holmes, 1895 (a junior homonym 
of Pseudopinnixa Ortmann, 1894) (Name No. 
508) ; 
(xlix) Ptychognathus Owen, 1860 (a junior homonym 
of Ptychognathus Stimpson, 1858) (Name No. 
509) ; 
(1) Telmessus Stal, 1866 (a junior homonym of 
Telmessus White, 1846) (Name No. 510) ; 


(li) Tlos White, 1847 (a nomen nudum) (Name No. 

SEB ee 

(lit) Trichodactylus Dufour, 1839 (a junior homonym 
of Trichodactylus Latreille, 1828) (Name No. 
512), 

(li1) Utica Hewitson, [1865] (a junior homonym of 
Utica White, 1847) (Name No. 513) ; 

(liv) Valdivia Ragonot, 1888 (a junior homonym of 
Valdivia White, 1847) (Name No. 514) ; 


(lv) Valdivia Shannon, 1927 (a junior homonym of 
Valdivia White, 1847) (Name No. 515) ; 


(Ilvi) Zosimus Schaufuss, 1886 (a junior homonym of 
Zosimus Leach, 1825) (Name No. 516) ; 


(lvii) Zozymus Milne Edwards (H.), 1834 (an Erro- 
neous Subsequent Spelling for Zosimus Leach, 
1825) (Name No. 517). 


(5) The under-mentioned specific names are hereby 
placed on the Official Index of Rejected and Invalid Specific 
Names in Zoology with the Name Numbers severally 
specified below :— 


(i) granulatus de Haan, [1835], as published in the 
combination Cancer (Actaea) granulatus (a junior 


62 OPINIONS AND DECLARATIONS 


homonym of granulatus Linnaeus, 1758, as 
published in the combination Cancer granulatus) 
(Name No. 336) ; 


(ii) pennanti Leach, [1817], as published in the com- 
bination Ebalia pennanti (a junior objective 
synonym of tuberosus Pennant, 1777, as pub- 
lished in the combination Cancer tuberosus) 
(Name No. 337) ; 


(111) tresselata Milne Edwards (H.), 1834, as published 
in the combination Lybia tresselata (an Errone- 
ous Subsequent Spelling for tesselatus Latreille, 
1812, as published in the combination Grapsus 
tesselatus) (Name No. 338) ; 


(iv) jurinii Bell, 1855, as published in the combination 
Leucosilia jurinii (an Erroneous Subsequent 
Spelling for jurinei Saussure, 1853, as published 
in the combination Guaia (Ilia) jurinei) (Name 
No. 339) ; 


(v) vigil Fabricius, 1798, as published in the com- 
bination Portunus vigil (a junior objective syno- 
nym of vigil Weber, 1795, as published in the 
combination Portunus vigil) (Name No. 340). 


I. THE SUBJECT MATTER OF THE PRESENT 
“DIRECTION. 


The present Direction is the third of a series containing decisions 
by the International Commission on Zoological Nomenclature 
in regard to names of genera of the Order Decapoda (Class 
Crustacea) placed on the Official List of Generic Names in Zoology 
in the period up to the end of 1936 taken by the International 
Commission under General Directives issued to it by the Thirteenth 
International Congress of Zoology, Paris, 1948, with the object of 
completing the entries made on the Official List in the foregoing 
period. The first of the Directions already rendered (Direction 11) 
contained decisions by the Commission as to the gender to be 
attributed to the generic names in question, while the second 


DIRECTION 37 —663 


of these Directions (Direction 36) was concerned with the placing 
on the Official List of Specific Names in Zoology of the specific 
names of the type species of the foregoing genera in all cases where 
those names were the oldest available for the species concerned 
and, in other cases, of the specific names subjectively considered 
to be the oldest available for the species in question. The present 
_ Direction is concerned with various miscellaneous matters on which 
decisions by the Commission are needed in order to enable it to 
comply fully with the General Directives referred to above. 
Apart from a few names which have been reserved for separate 
consideration, the present Direction, coupled with its two pre- 
decessors, completes the action required in the case of the names of 
genera of the Order Decapoda placed on the Official List in the 
period under review. The proposals which formed the basis 
of the decisions embodied in the present Direction were submitted 
by the Secretary in the following paper on 19th September 
1955 :— 


Names of genera of the Order Decapoda in the Class Crustacea placed 
on the ‘‘ Official List of Generic Names in Zoology ”’ in the period 
up to the end of 1936 : proposed determination of certain out- 
standing matters additional to those dealt with in the 
proposals submitted with in Voting Papers 
V.P.(O.M.)(55)6 and V.P.(O.M.)(55)27 


By FRANCIS HEMMING, C.M.G., C.B.E., 


(Secretary to the International Commission on Zoological Nomenclature) 


The present paper contains a final instalment of proposals relating 
to the names of genera of the Order Decapoda in the Class Crustacea 
placed on the Official List of Generic Names in Zoology by Rulings 
given in Opinions rendered in the period up to the end of 1936. The 
two earlier instalments were concerned respectively with the determin- 
ation of the gender to be attributed to the generic names concerned 
(proposals submitted with Voting Paper V.P.(O.M.)(55)6) and with the 
addition to the Official List of Specific Names in Zoology of the specific 
names of the type species of the genera in question (proposals sub- 
mitted with Voting Paper V.P.(O.M.)(55)27). The present paper is 
concerned with various other matters on which decisions by the 
Commission are required before this large block of entries on the 
Official List of Generic Names in Zoology can be regarded as having 


64 OPINIONS AND DECLARATIONS 


been brought up to the standard required for inclusion in the forth- 
coming volume containing particulars of all names so far placed on 
the Official List. 


2. The first point which calls for note is that, whereas in Opinions 
73 and 104 some information (though often of a meagre character) was 
given on the question of how the species there cited as the type species 
of the genera concerned acquired that status under Article 30 of the 
Régles, no information of any sort bearing on this subject was provided 
in respect of the ninety-eight genera, the names of which were placed 
on the Official List of Generic Names in Zoology by the Ruling given 
in Opinion 85. In the case of these names it has been necessary to 
examine the book or serial in which each was published in order to 
ascertain whether the species cited in Opinion 85 was one of the included 
species and whether that species was there either designated or indicated 
as the type species. In those cases where none of the included species 
was so designated or indicated, it was necessary, in conjunction with 
interested specialists, to examine the later literature for the purpose of 
ascertaining by whom, when and where one of the originally included 
species was first selected to be the type species of the genus concerned. 
The results of these laborious investigations are set out in Annexe 1 
to the present paper. 


3. It is necessary next to note that in the case of fourteen names 
placed on the Official List of Generic Names in Zoology by the Ruling 
given in Opinion 73, the only note given as to the manner in which the 
type species of the genera concerned had been determined under 
Article 30 was the insertion of the formula “‘ tpd ”’, i.e. type by present 
designation. Since these type selections have sometimes been attri- 
buted to the Commission, it should be noted that the portion of the 
Opinion in which this formula is used consists of the application sub- 
mitted to the Commission by Miss Mary Rathbun, the applicant, 
to whom therefore these type selections are attributable. The names 
concerned are the following :—Actaea ; Actumnus ; Atergatis ; Chloro- 
dopsis ; Cyclodius ; Ebalia; Iliacantha ; Pariphiculus ; Persephona ; 
Phlyxia ; Pseudophilyra ; Speloeophorus ; Thalamitoides ; Thalamonyx. 


4. In a number of cases wrong dates or erroneous attributions for 
generic names were given in the Opinions in which the names concerned 
were placed upon the Official List. The necessary corrections are set 
out in Annexe 2 to the present paper. In the same Annexe particulars 
are given also regarding the dates of publication of certain books 
which were issued in Parts and which therefore present certain diffi- 
culties in dating. 


5. Annexe 3 gives particulars of certain objectively invalid names 
which are involved in one way or another with the names of genera 


DIRECTION 37 65 


placed on the Official List in the period up to the end of 1936 and 
which under the General Directives issued to the Commission by the 
International Congress of Zoology should now be placed on the 
appropriate Official Index of Rejected and Invalid Names. This 
Annexe is divided into two Sections : Section A which contains a list 
of invalid generic names, Section B, which contains a list of invalid 
specific names. 


6. Although the number of generic names of the Order Decapoda 
placed on the Official List in the period up to the end of 1936 is large, 
the genera concerned are in many cases closely allied to one another. 
In consequence, the number of family-group names involved is 
relatively small. These names are now being investigated and pro- 
posals in regard to them will be submitted in due course. 


7. The proposals now submitted are that, in accordance with the 
Directives issued by the International Congress of Zoology, the 
Commission should complete the Rulings in relation to the names of 
genera in the Order Decapoda (Class Crustacea) given in Opinions 
rendered in the period up to the end of 1936 by :— 


(a) entering on the Official List of Generic Names in Zoology the 
particulars specified in Annexe 1 regarding the manner in 
which type species for the genera there enumerated were 
determined under Article 30 of the Régles ; 


(b) taking note that the type species for the genera specified in 
paragraph 3 above were selected under Rule (g) in Article 
30 by Miss Mary Rathbun in her application to the Commission 
published in Opinion 73 ; 


(c) entering on the foregoing Official List the corrections, specified 
in Annexe 2, of wrong dates and erroneous attributions given 
in the Opinions in which generic names of Decapoda were 
placed on that List ; 


(d) placing on the Official Index of Rejected-and Invalid Generic 
Names in Zoology the generic names specified in Section A 
of Annexe 3 and on the corresponding Official Index of specific 
names the specific names specified in Section B of that Annexe. 


8. The proposals now submitted complete the action required for 
the completion and, where necessary, the correction, of entries relating 
to the names of genera of Decapoda placed on the Official List of 
Generic Names in Zoology in the period up to the end of 1936, except 
as regards two names which are being further investigated and on which 
proposals will be submitted to the Commission as soon as possible. 


66 OPINIONS AND DECLARATIONS 


ANNEXE 1 


Particulars as to the determination, under Article 30 of the ‘‘ Régles ”’, 
of the type species of certain genera in the Order Decapoda (Class 
Crustacea), the names of which were placed on the ‘‘ Ofiicial 
List of Generic Names in Zoology ”’ in the period up to 
the end of 1936 


Name of Genus 


(1) 


Acmaeopleura Stimpson, 1858 
Asthenognathus Stimpson, 1858 


Bathyplax Milne Edwards, 1880 


Camptandrium Stimpson, 1858 
Camptoplax Miers, 1884 
Catoptrus Milne Edwards, 1870 
Ceratoplax Stimpson, 1858 
Chasmagnathus de Haan, [1833] 
Chasmocarcinus Rathbun, 1898 
Clistocoeloma Milne Edwards, - 
1873 
Cyrtograpsus Dana, 1851 


Dissodactylus Smith, 1870 
Durckheimia de Man, 1889 


Epixanthus Heller, 1861 

Euchirograpsus Milne Edwards, 
1853 

Eucrate de Haan, [1835] 

Eucratodes Milne Edwards, 1880 

Eucratopsis Smith, 1869 

Euryetisus Cano, 1889 


Euryplax Stimpson, 1859 
Eurytium Stimpson, 1859 
Fabia Dana, 1851 

Galene de Haan, [1833] 
Geryon Kr@yer, 1837 
Glyptograpsus Smith, 1870 
Glyptoplax Smith, 1870 
Gomeza Gray, 1831 
Goneplax [Leach], [1814] 


Entry relating to the determination 
of the type species of the genus 
specified in Col. | now proposed 

to be inserted in the “Official List 
of Generic Names in Zoology ”’ 


(2) 


by monotypy 
by monotypy 


by monotypy 


by monotypy 
by monotypy 
by monotypy 
by monotypy 
by monotypy 
by original designation 
by monotypy 


by monotypy 


by monotypy 
by original designation through 
the formula “‘n.g., n.sp.”’ 


by monotypy 
by monotypy 


by monotypy 

by monotypy 

by monotypy 

by original designation through 
the formula “‘n.g., n.sp.”’ 

by monotypy 

by original designation 


by monotypy 


by monotypy 
by original designation 
by original designation 
by monotypy 
by monotypy 
by original designation 


DIRECTION 37 67 


Entry relating to the determination 
of the type species of the genus 
specified in Col. 1 now proposed 


Name of Genus to be inserted in the ‘‘ Official List 
of Generic Names in Zoology ”’ 
(1) (2) 

Halimede de Haan, [1835] by monotypy 

Helice de Haan, [1833] by monotypy 

Hephthopelta Alcock, 1899 by monotypy 

Hexapus de Haan, [1833] by monotypy 

Holometopus Milne Edwards, 1853 by monotypy 

Holothuriophilus Nauck, 1880 by original designation through 

the formula “n.g., n.sp.” 

Lachnopodus Stimpson, 1858 by monotypy 

Leptodius Milne Edwards, 1863 by monotypy 

Liagore de Haan, [1833] by monotypy 

Libystes Milne Edwards, 1867 by monotypy 

Liomera Dana, 1851 by monotypy 

Lipaesthesius Rathbun, 1898 by monotypy 

Litocheira Kinahan, 1856 by monotypy 

Lophopanopeus Rathbun, 1898 by original designation 
Lophopilumnus Miers, 1886 by original designation 

Lybia Milne Edwards, 1834 by monotypy 

Melybia Stimpson, [1871] by monotypy 


Metasesarma Milne Edwards, 1853 by monotypy 
Metopocarcinus Stimpson, [1860] by monotypy 


Micropanope Stimpson, [1871] by monotypy 
Notonyx Milne Edwards, 1873 by monotypy 
Oediplax Rathbun, 1893 by monotypy 
Ommatocarcinus White, 1852 by monotypy 
Opisthopus Rathbun, 1893 by monotypy 
Orphnoxanthus Alcock, 1898 by monotypy 
Panoplax Stimpson, [1871] by monotypy 
Paragalene Kossmann, 1878 by monotypy 
Parapanope de Man, 1895 by monotypy 


Parapleurophrycoides Nobili, 1906 by monotypy 
Paraxanthus Milne Edwards & by monotypy 
Lucas, [1844] 


Percnon Gistl, 1848 by subsequent selection by Rath- 
bun, 1925, Smiths. misc. Publ. 
TIT 6 

Perigrapsus Heller, 1862 by monotypy 


Pilumnoides Milne Edwards & by monotypy 
Lucas, [1844] 
Pilumnus Leach, 1815 by monotypy 
Pinnaxodes Heller, [1865] by original designation through 
the formula “‘n.g., n.sp.”’ 


68 


Name of Genus 


(1) 


Pinnixa White, 1846 

Pinnotherelia Milne Edwards & 
Lucas, [1844] 

Pinnotheres Latreille, [1802—1803] 

Planes Bowditch, 1825 

Platychirograpsus de Man, 1896 


Platypilumnus Alcock, 1894 


Platyxanthus Milne Edwards, 1863 

Polydectus Milne Edwards, 1837 

Prionoplax Milne Edwards, 1852 

Pseudocarcinus Milne Edwards, 
1834 


Pseudopinnixa Ortmann, 1894 

Pseudorhombila Milne Edwards, 
1837 

Psopheticus Wood-Mason, 1892 

Ptychognathus Stimpson, 1858 

Pyxidognathus Milne Edwards, 
1879 


Rhithropanopeus Rathbun, 1898 
Rhizopa Stimpson, 1858 
Ruppellioides Milne Edwards, 
1867 
Samartium Dana, 1851 
Scalopidia Stimpson, 1858 
Scleroplax Rathbun, 1893 
Speocarcinus Stimpson, 1859 
Sphaerozius Stimpson, 1858 
Tetraxanthus Rathbun, 1898 
Tetrias Rathbun, 1898 
Thaumastoplax Miers, 1881 


Utica White, 1847 
Varuna Milne Edwards, 1830 


Xanthasia White, 1846 
Xanthodius Stimpson, 1859 
Xenophthalmodes Richters, 1880 


OPINIONS AND DECLARATIONS 


Entry relating to the determination 
of the type species of the genus 
specified in Col. 1 now proposed 

to be inserted in the “‘ Official List 
of Generic Names in Zoology ” 


(2) 


by monotypy 
by monotypy 


by monotypy 

by monotypy 

by original designation through 
the formula “‘ n.g., n.sp.”’ 

by original designation through 
the formula “‘n.g., n.sp.”’ 

by monotypy 

by monotypy 

by monotypy 

by subsequent selection by Miers, 
1886, Rep. Voy. Challenger 
(Zool.) 17(no. 2) : 141, nota 1. 

by monotypy 

by original designation 


by monotypy 
by monotypy 
by monotypy 


by monotypy 
by monotypy 
by monotypy 


by monotypy 
by monotypy 
by monotypy 
by monotypy 
by monotypy 


by monotypy 

by monotypy 

by original designation through 
the formula “‘n.g., n.sp.”’ 


by monotypy 
by monotypy 


by monotypy 

by monotypy 

by original designation through 
the formula “‘ n.g., n.sp.”’ 


DIRECTION 37 69 


Entry relating to the determination 
of the type species of the genus 
specified in Col. 1 now proposed 


Name of Genus to be inserted in the “‘ Official List 
of Generic Names in Zoology ”’ 
(1) (2) 
Xenophthalmus White, 1846 by monotypy 
Zosimus Leach, 1823 by monotypy 
Zozymodes Heller, 1861 by monotypy 
ANNEXE 2 


Proposals for the correction of certain erroneous dates and attributions 
cited for the names of genera of the Order Decapoda (Class 
Crustacea) when placed on the ‘‘ Official List of Generic 
Names in Zoology ”’ by Rulings given in ‘‘ Opinions ”’ 73, 

85 and 104 or for the names of the type species of such 

genera 


(a) Corrections of entries in ‘‘ Opinion ’’ 73 
(1) ‘* Bathynectes ’’ Stimpson 


Two questions arise in connection with this generic name: first, 
the date on which it was published ; second, the manner in which the 
type species was determined. These matters are discussed separately 
below. 


The name Bathynectes Stimpson was published in Part 2 of vol. 2 of 
the Bull. Mus. comp. Zool. of Harvard. This name (and other new 
names occurring in the same paper) are treated as having been published 
in 1870 in some catalogues and in others as having been published in 
1871. In order to clear up this matter, I appealed to my colleague the 
late Dr. James L. Peters, who, as a member of the staff of the Museum 
of Comparative Zoology, was in a specially good position for investi- 
gating the dates of publication of parts of the Bulletin of that Museum. 
On Ist June 1944, Dr. Peters replied as follows :— 


The early numbers of the M.C.Z. Bulletin were issued without 
covers and no printed date of publication. The number in question 
bears the date and inscription at the end of the article, ‘“‘ Chicago 
Academy of Science, December 1, 1870”. This apparently is 
when the manuscript was completed and should not be accepted 
as the publication date. The accession catalogue in the Museum 
Library shows that a bound copy of No. 2, Vol. 2, was entered 
on April 12, 1871. Inquiring of the librarian of the Boston 
Society of Natural History reveals that a copy in paper covers, 


70 OPINIONS AND DECLARATIONS 


as issued, was received there on February 17, 1871. On the other 
hand, the price list of Museum publications claims December 
1870 as the date. This evidence seems to show that the actual 
date of publication was probably about the middle of Feb. 1871 
and the fact that our Museum catalogue doesn’t show an entry 
until April 12th is accounted for by the fact that the issue was 
not entered until it had been returned from the bindery. 


Further check-up with the Museum price-list and the date of 
receipt of the unbound parts at the Boston Society of Natural 
History shows that, in each case, the numbers of Vol. 2 were 
received by that institution a month to six weeks after the date 
claimed in the Museum price-list. 


In view of the information supplied by Dr. Peters and having 
regard to the fact that, as he shows, Stimpson’s paper was still in 
manuscript at the Chicago Academy as late as Ist December 1870, it 
must certainly be concluded that publication did not take place until 
some date in 1871. 


In the application published in Opinion 73 Miss Rathbun stated 
that the type species of Bathynectes Stimpson was “‘ tod. B. superba = 
Portunus superba Costa, 1838? = B. longispina Stimpson,, 1871’. 
This statement is incorrect in two respects. First Stimpson made no 
mention of Portunus superba Costa, which was therefore not an 
originally-included species and is accordingly ineligible for selection 
as the type species of Bathynectes. Second, contrary to the statement 
given by the abbreviated entry “‘ tod.’ (= type by original designation) 
Stimpson did not designate a type species for the genus Bathynectes. 
All that he did was to include in it two new nominal species, namely 
Bathynectes longispina (: 146) and Bathynectes brevispina (: 147) and to 
add as regards the latter, that he was doubtful whether it was 
specifically distinct from Jongispina. This latter statement, however, 
made the nominal species Bathynectes brevispina a species inquirenda 
from Stimpson’s standpoint. Accordingly, under the second subsection 
of Rule (e) in Article 30, the nominal species Bathynectes brevispina 
becomes ineligible for selection as the type species of Bathynectes. 
As a result, that genus is left with only one eligible included species, 
namely, Bathynectes longispina Stimpson, which thus becomes the 


type species of this genus, not by original designation as stated in- 


Opinion 73, but by monotypy. 


(2) ‘* Caphyra’’ Guérin-Ménéville 

Both the place and date of publication given for this generic name 
in Opinion 73, as also for the name of its type species, are incorrect. 
The entry given in this Opinion reads as follows :—‘‘ Caphyra Guérin, 
1830, 26, mt. C. rouxii Guérin, 1830.’’ Reference to the list of papers 
given earlier (: 24) in Opinion 73 shows that the expression ‘* Guérin, 


DIRECTION 37 A 


1830’ was intended to be interpreted as meaning ‘‘ Guérin, 1830, 
Voy. Coquille, Zool., v.2, Crust.” Reference to the Voy. Coquille 
shows that these names were published in vol. 2 (No. 2) (Part 1) of 
the Zoology Section (Caphyra on page 26 and C. rouxii on page 28). 
The point to be noted here is that a detailed analysis of the evidence 
relating to the dates of publication of the Voy. Coquille made by 
Sherborn in 1905 (Ann. Mag. nat. Hist. (7) 17 : 335) shows that the 
Crustacea portion was issued as Livraison 28 (which contained 60 
sheets, i.e. the whole of Vol. 2, No. 2) and that this Livraison was not 
published until 1838. Thus if Caphyra had first been published in the 
Voy. Coquille, it would be necessary to attribute it to the year “‘ 1838 ” 
and not to the year “* 1830” (the year cited in Opinion 73). But in 
fact that generic name and the name of the type species of the genus 
so named were first published in a paper in the Ann. Sci. nat. of Paris 
which appeared in 1832. The correct references for the two names in 
question are thus :—Caphyra Guérin-Ménéville, 1832, Ann. Sci. nat., 
Paris 25(99) : 285; Caphyra rouxii Guérin-Ménéville, 1832, ibid. 
25(99) : 287. 


(3) ‘* Hepatella ’’ Smith 


The name Hepatella Smith and the name (Hepatella amica Smith) of 
the type species of the genus so named were published in volume 3 of 
the American Naturalist. These names were assigned the date “‘ 1870” 
in Opinion 73. Inspection of this volume shows that Part 5 (the 
part in which the foregoing names appeared) bears the date “‘ July 
1869’. It also bears the following legend :— 


** Entered according to Act of Congress in the year 1869, by 
the Peabody Academy of Science, in the clerk’s office of the District 
Court of the District of Massachusetts.” 


On the other hand, the title page of the volume as a whole bears the 
date “* 1870’ and it was no doubt from this source that this date was 
cited in Opinion 73. There are no grounds for rejecting the evidence 
cited above and it is reasonable therefore to conclude that this volume 
appeared partly in 1869 and partly in 1870 and that the Part containing 
the names here under consideration was published in the earlier of these 
two years. 


(4) Relative dates of publication in 1855 of two papers by Bell (T.), 
each containing identical names as new names. 


A number of names dealt with in Opinion 73 were first published in 
1855 in a paper by Bell (T.), the place of publication being cited in 
that Opinion as volume 21 of the Transactions of the Linnean Society 
of London. It appears from the Minutes of the Society that the paper 
was read at a meeting held under Bell’s Presidency on 5th June 1855. 
It was published later in the same year in the Transactions, but previous 
to this an abstract of it containing new names with the requisite 
indications appeared in the issue of the Proceedings of the Society, 


Fp OPINIONS AND DECLARATIONS 


published on the 19th October 1855. The names concerned should 
therefore be attributed to the Proceedings and not to the Transactions. 


The names to which the foregoing correction applies and the number 
of the page in vol. 2 of the Proceedings in which those names first 
appeared are as follows :—Leucosilia (: 431) ; Lithadia (: 434) ; Myrodes 
(: 432) ; Nursilia (: 434) ; Phlyxia (: 433) ; the name Oreophorus nodosus 
(: 434) (specific name of type species of Speloeophorus). 


(5) Date of publication of the ‘* Crustacea ’’ Section of Arthur 
White’s ‘‘ The Voyage of H.M.S. Samarang ’’. 


A number of names first published in the Crustacea Section of Adams 
(A.), The Voyage of H.M.S. Samarang were dealt with in Opinion 73 
where they were all treated as having been published in 1848. Sherborn, 
however, has shown (1922, Index Anim., Sectio secund. (1) : cxi) that 
the Crustacea Section of this book was published in two parts, the first, 
containing pp. 1—32, in July 1848 ; the second, containing pp. 33—66, 
in April 1849. Re-examination of the dates assigned in Opinion 73 
to names published in this work shows that the following names must be 
dated ‘“‘ 1849” instead of ‘* 1848’ :—(i) Carpilius signatus (: 37) 
(type species of Atergatopsis) ; (11) Iphiculus (: 57) and its type species 
Iphiculus spongiosus (: 57) ; (iii) Lissocarcinus (: 45) and its type species 
Lissocarcinus polybioides (: 46); (iv) Lupocyclus (: 46) and its type 
species Lupocyclus rotundatus (: 47); Tlos (: 57) and its type species 
Tlos muriger (: 58). 


(6) ‘*‘ Trichodactylus ’’ Latreille 


This name was published in Part 2 of volume 10 (Crustacea) of 
the Ency. méth., and in Opinion 73 the date “‘ 1825’ was attributed 
to it. This is, however, incorrect, being the date of publication of the 
first part of the volume. Part 2 was published in 1828. 


(7) ‘*Cymo”’ de Haan, [1833] 


In Opinion 73 the date “ 1825”’ was attributed to Pilumnus andreossyi 
Audouin, the name of the type species of the genus Cymo. This 
matter, however, was investigated by Sherborn (1923, Index Anim., 
Sectio secund. : 305) who ascertained that the above name should 
be treated as having been published in 1826. 


(8) ‘*Scylla’’? de Haan, [1833] 


This genus was stated by Miss Rathbun in Opinion 73 to be mono- 
typical with Cancer serratus Forskal, 1775, as type species. Miss 
Rathbun was aware that de Haan had placed a second nominal species 
in this genus, namely, Portunus tranquebaricus Fabricius, 1798, for she 
noted this fact in the application published in the Opinion, and added 
that these species ‘‘are synonymous’. Although she was of the opinion 
that only one valid taxon had been placed by de Haan in this genus, 


DIRECTION 37 73 


it was erroneous from the nomenclatorial point of view to state that 
the genus was monotypical. She herself in the application referred 
to above definitely selected C. serratus Forskal to be the type species 
of Scylla de Haan, and this appears to be the first, and therefore the 
valid, type selection for this genus. 


b) Corrections of entries in ‘* Opinion ”’ 85 
p 


(9) Date of publication of the Crustacea Section of d’Orbigny’s 
** Voyage dans l’Amerique Meridionale ”’’ 


Both in Opinion 73 and in Opinion 85 there are names which were 
first published in the Crustacea Section of d’Orbigny’s Voyage dans 
l’ Amerique Meridionale. There is no consistency as to the dates assigned 
to those names in those Opinions. The dates given in Opinion 73 are 
correct but those given in Opinion 85 for the following names were 
incorrectly given as “1843” instead of “‘ 1844” :—Paraxanthus ; 
Pilumnoides ; Pinnotherelia. 


(10) ‘‘ Pinnotheres ’’ Latreille 


The name Pinnotheres Latreille was published in volume 3 of the 
Hist. nat. gen. partic. Crust. Ins. in Sonnini’s Buffon. In Opinion 85 
the date given for this name was “‘ 1801—2 ”’ and it was stated that the 
volume concerned was published in An X of the French Revolutionary 
Calendar. This ran from September 1801 to September 1802. While 
*“An X”’ is the date given on the title page of the volume, it has now 
been established that it cannot have been published before some date 
in “An XI”’, i.e. in the period September 1802 to September 1803 
(see Hemming, 1946, Bull. zool. Nomencl. 1 : 205, footnote 7). 


(c) Corrections of entries in ‘*‘ Opinion ’’ 104 
Pp 


(11) ‘‘Nephrops ”’ [Leach] 


Two points call for notice in connection with the name Nephrops. 
First, it must be noted that it is sometimes cited in the literature as 
having been first published in 1816 in the Supplementary Volume to 
the 4th—6th Editions of the Encyclopedia Britannica. In fact, however, 
it was first published in an anonymous article by Leach in Brewster’s 
Edinburgh Encyclopedia (7(2) : 398—400). Second, it should be noted 
that Part 2 of vol. 7 of Brewster’s Edinb. Ency. was published in 1814 
and not in 1815, the year attributed to it in Opinion 104. (For a note 
on the date of publication of this portion of the Edinb. Ency. see 
Rathbun, 1897, Proc. biol. Soc. Wash. 11 : 154, nota.). 


(12) ‘*Pandalus’’ Leach, 1815 


The name Pandalus was published by Leach twice in the year 1815. 
The first occasion was in the text to Plate 40(2) of his Malac. podophth. 
Brit. The second occasion was in the Trans. linn. Soc. Lond. (11(2) : 
346). By some oversight it was the latter instead of the former of 
these references which was cited in Opinion 104 as the original reference 
for this name. 


74 


10. 


OPINIONS AND DECLARATIONS 


ANNEXE 3 
Proposed addition of certain objectively invalid names to 
the ‘* Official Indexes of Rejected and Invalid Names ”’ 


Section A:—Proposed addition of fifty-one generic 
names to the “ Official Index of Rejected and 
Invalid Generic Names in Zoology ”’ 


. Acanthocyclus Dybowski, 1873, Arch. Naturk. Liv-, Ehst-u. Kuhl. 


Dorpat (1)5 : 333 (a junior homonym of Acanthocyclus Milne 
Edwards (H.) & Lucas, [1844], placed on the Official List in 
Opinion 73) 


. Acanthopus de Haan, [1835], in Siebold, Fauna japon. (Crust.) : 29 


(a junior homonym of Acanthopus Klug, 1807, Mag. f. Insektenk. 
(Illiger) 6 : 199, 226) (replaced by Percnon Gistl, 1848, placed on 
the Official List in Opinion 85) 


. The under-mentioned junior homonyms of Asthenognathus 


Stimpson, 1858, placed on the Official List in Opinion 85 :— 


(a) Asthenognathus Bocourt, 1884, Bull. Sci. Soc. Philomat. 
Paris (7)8 : 149 

(b) Asthenognathus Broom, 1915, Bull. Amer. Mus. nat. Hist 
25 ‘125 


The under-mentioned junior homonyms of Bathynectes Stimpson, 
[1871], placed on the Official List in Opinion 73 :— 


(a) Bathynectes Guenther, 1878, Ann. Mag. nat. Hist. (5)2 : 20 
(b) Bathynectes Brinkman, 1913, Bergens Mus. Aarb. 1912 (9) : 1 


The under-mentioned junior homonyms of Bellia Milne Edwards 
(H.), 1848 placed on the Official List in Opinion 73 :— 


(a) Bellia Bate, 1851, Ann. Mag. nat. Hist. (2)7 : 318 
(b) Bellia Gray (J.E.), 1869, Proc. zool. Soc. Lond. 1869 : 197 
(c) Bellia Tutt, 1902, Nat. Hist. Brit. Lep. 3 : 386 


. Caphyra Barrande, 1846, Not. prélim. Syst. Silur. Trilob. Bohéme : 


32 (a junior homonym of Caphyra Guérin-Ménéville, 1832, 
placed on the Official List in Opinion 73 


Chlorodopsis Milne Edwards, 1863, Ann. Sci. nat., Paris, Zool. 
(4)20 : 292 (a nomen nudum) 


Cyrtograpsus Carruthers, 1867, Intell. Observ. 11 : 369 (a junior 
homonym of Cyrtograpsus Dana, 1851, placed on the Official 
List in Opinion 85) 


Eucrate Rafinesque, 1915, Analyse Nature : 136 (a nomen nudum) 


Euryplax Klapalek, 1909, Wien. ent. Ztg. 28 :255 (a junior 
homonym of Euryplax Stimpson, 1859, placed on the Official 
List in Opinion 85) 


11. 


12. 


, 


14. 


3. 


16. 


¥2 


18. 


19. 


20. 


DIRECTION 37 To 


Gecarcinicus Dana, 1852, U.S. Explor. Exped. 13 (Crust.) : 375, 
pl. 1 (an Erroneous Subsequent Spelling for Gecarcinucus 
Milne Edwards (H.), 1844, placed on the Official List in 
Opinion 73) 


Goneplat [Leach], [1814], Brewster’s Edinb. Ency. 7 : 393 (an 
Invalid Original Spelling for Goneplax [Leach], [1814], placed 
on the Official List in Opinion 85) 


Gonoplax Leach, 1815, Trans. linn. Soc. Lond, 11(2) : 323 (an 
Invalid Emendation of Goneplax [Leach], [1814], placed on the 
Official List in Opinion 85) 


The under-mentioned junior homonyms of Halimede de Haan, 
[1835], placed on the Official List in Opinion 85 :— 


(a) Halimede Rathke, 1843, Nova Acta Acad. Caes. Leop. Carol. 
20(1) : 166 


(b) Halimede Oberthuer & Houlbert, 1922, C.R. Acad. Sci. 
Paris 174 : 190 


Helice Chambers, 1873, Canad. Ent. 5 : 187 (a junior homonym 
of Helice de Haan, [1833], placed on the Official List in Opinion 
85) | 


Holometopus Angelin, 1854, Pal. Scand. (ed. 1) fasc. 2 : 58 (a 
junior homonym of Holometopus Milne Edwards (H.), 1853, 
placed on the Official List in Opinion 85) 


The under-mentioned junior homonyms of Homalaspis Milne 
Edwards (A.), 1863, placed on the Official List in Opinion 85 :— 


(a) Homalaspis Brunner von Wattenwyl, 1895, Monogr. Pseudo- 
phylliden 9 : 217 


(b) Homalaspis Kaier, 1932, Skr. Svalbard Ishavet, Oslo, 52 : 14 
Homarus Broun, 1881, Man. N.Z. Coleopt. (fasc. 2) : 740 (a junior 


homonym of Homarus Weber, 1795, placed on the Official 
List in Opinion 104) 


Iphiculus White, 1847, List. Crust. Coll. B.M. : 51 (a nomen nudum) 
The under-mentioned junior homonyms of Jphis Leach, 1817, 
placed on the Official List in Opinion 73 :— 


(a) Iphis Koch (C.L.), 1835, Deutschl. Crust. Myriap. Arach. 
(Heft 2) : tab. 6 


(b) Iphis Laporte, 1836, Revue Entom. 4(1) : 7 


76 


a 


2. 


De 


24. 


2D) 


26. 


27. 


28. 


2. 


30. 


an: 


OPINIONS AND DECLARATIONS 


Liagora Gistl, 1848, Nat. Thierr. : ix (an Erroneous Subsequent 
Spelling for Liagore de Haan, [1833], placed on the Official 
List in Opinion 85) 


Lissocarcinus Adams & White, 1847, in White’s List. Crust. coll. 
B.M. : 126 (a nomen nudum) 


Lupocyclus White, 1847, List Crust. Coll. B.M. : 127 (a nomen 
nudum) 


Persephona Leach, 1852, Syn. Moll. Gt. Brit. : 146, 189 (a junior 
homonym of Persephona Leach, 1817, placed on the Official List 
in Opinion 73) 


The under-mentioned junior homonyms of Pilumnus Leach, 1815, 
placed on the Official List in Opinion 85 :— 


(a) Pilumnus Rafinesque, 1815, Analyse Nature : 122 

(b) Pilumnus Dejean, 1821, Cat. Coll. Coléopt. Dejean [1821 ed.] 
3365 1d-18237 Cola Lepiad-2156 

(c) Pilumnus Koch (C.L.), 1837, Uebers Arachnidensyst. 1 : 38 

(d) Pilumnus Gistl, 1848, Nat. Thierr. : xi (nom. nov. pro Alphus 
Dejean, 1833) 

(e) Pilumnus Bonaparte, 1854, Ateneo Ital. 2 : 123 


The under-mentioned junior homonyms of Planes Bowditch, 1825, 
placed on the Official List in Opinion 85 :— 
(a) Planes Saussure, 1862, Rev. Mag. Zool. (2)14 : 228 
(b) Planes Rondani, 1864, Arch. Zool. anat. Fisiol. 3 (fasc. 1) : 9 


Podacanthus Brandt, 1848, Bull. Acad. Sci. St. Petersb. 7(12/13) : 
180 (a junior homonym of Podacanthus Gray, 1833) (replaced 
by Erimacrus Benedict, 1892, placed on the Official List in 
Opinion 73) 


Podophtalmus Lamarck, 1801, Syst. Anim. : 152 (an Invalid Ori- 
ginal Spelling for Podophthalmus Lamarck, 1801, placed on the 
Official List in Opinion 73) 


Polydectus Rafinesque, 1815, Analyse Nature : 142 (a nomen 
nudum) 


Pseudopinnixa Holmes, 1895, Proc. Cal. Acad. Sci. (2)4 : 565 (a 
junior homonym of Pseudopinnixa Ortmann, 1894, placed on the 
Official List in Opinion 85) 


Ptychognathus Owen, 1860, Quart. J. geol. Soc. Lond. 16 : 49 (a 
junior homonym of Ptychognathus Stimpson, 1858, placed on the 
Official List in Opinion 85) 


a2. 


33. 
34. 


32. 


36. 


aT, 


38. 


DIRECTION 37 Th 


Telmessus Stal, 1866, Hemipt. Afric. 4 : 211 (a junior homonym of 
Telmessus White, 1846, placed on the Official List in Opinion 73) 


Tlos White, 1847, List Crust. Coll. B.M. : 129 (a nomen nudum) 


Trichodactylus Dufour, 1839, Ann. Sci. nat., Paris (Zool.) 11 : 276 
(a junior homonym of Trichodactylus Latreille, 1828, placed on 
the Official List in Opinion 73) 


Utica Hewitson, [1865], J//. diurn. Lep. Lycaenidae 1 : 56 (a 
junior homonym of Utica White, 1847, placed on the Official 
List in Opinion 85) 


The under-mentioned junior homonyms of Valdivia White, 1847, 
placed on the Official List in Opinion 73 :— 
(a) Valdivia Ragonot, 1888, Nouv. gen. esp. Phycit. Gall. : 27 
(b) Valdivia Shannon, 1927, Proc. U.S. nat. Mus. 70(art. 9) : 31 


Zosimus Schaufuss, 1886, Tijdschr. Ent.29 : 293 (ajunior homonym 
of Zosimus Leach, 1825, placed on the Official List in Opinion 85) 


Zozymus Milne Edwards (H.), 1834, Roret’s Suite a Buffon 
(Crust.) 1 : 383 (an Erroneous Subsequent Spelling for Zosimus 
Leach, 1825, placed on the Official List in Opinion 85) 


Section B :—Proposed addition of six specific names to the 
** Official Index of Rejected and Invalid Specific Names 
in Zoology ”” 


. granulatus de Haan, [1835], as published in the combination 


Cancer (Actaea) granulatus (in Siebold, Fauna japon. (Crust.) 
> 4, 18) (a junior homonym of granulatus Linnaeus, 1758, as 
published in the combination Cancer granulatus) 


. ponnanti Leach, [1817], as published in the combination Ebalia 


pennanti (Malac. podophth. Brit. (14) : text to pl. 25) (a junior 
objective synonym of tuberosus Pennant, 1777, as published 
in the combination Cancer tuberosus). 


. tresselata Milne Edwards, 1834, as published in the combination 


Lybia tresselata (Roret’s Suite 4 Buffon, Hist. nat. Crust. 1 : 431) 
(an Erroneous Subsequent Spelling for tesselatus Latreille, 1812, 
as published in the combination Grapsus tesselatus) 


. variegatus [Leach], [1814], as published in the combination 


Portumnus variegatus (Brewster’s Edinb. Ency. 7(2) : 391) (a 
junior objective synonym of /atipes Pennant, 1777, as published 
in the combination Cancer latipes) 


78 OPINIONS AND DECLARATIONS 


5. jurinii Bell, 1855 (as published in the combination Leucosilia 
jurinii Proc. linn. Soc. Lond. 2 : 431; id., 1855, Trans. linn. Soc. 
Lond. 21 : 295) (an Erroneous Subsequent Spelling for jurinei 
Saussure, 1853, as published in the combination Guaia (llia) 
jurinei) 

6. vigil Fabricius, 1798, as published in the combination Portunus 


vigil (Suppl. Ent. syst. : 363) (a junior objective synonym of vigil 
Weber, 1795, as published in the combination Portunus vigil) 


2. Registration of the present application : Upon the receipt 
of Mr. Hemming’s paper the question of making the additions to 
the Official Lists and Official Indexes recommended therein was 
allotted the Registered Number Z.N.(S.) 987. 


Il. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


3. Issue of Voting Paper V.P.(O.M.)(55)28 : On 19th September 
1955 a Voting Paper (V.P.(O.M.)(55)28) was issued in which each 
Member of the Commission was asked (1) to state whether he 
agreed “‘ that, in accordance with the General Directives issued to 
the International Commission on Zoological Nomenclature by 
the Thirteenth International Congress of Zoology, Paris, 1948, 
regarding the placing on the appropriate Official Lists and Official 
Indexes of names dealt with in, or otherwise involved in, Opinions 
rendered prior to 1948, and other matters on which action is 
required in order to complete, to clarify, or to correct Rulings 
given in Opinions rendered in the foregoing period, the action 
recommended in paragraph 7 of the paper numbered Z.N.(S.) 987, 
regarding names in the Order Decapoda (Class Crustacea), 
submitted by the Secretary simultaneously with the present 
Voting Paper [i.e. in the paragraph numbered as above in the 
paper reproduced in the first paragraph of the present Direction] 
be now taken as there recommended ”’, and (2), if he did not so 
agree as regards any given item, to indicate that item. 


) 


DIRECTION 37 79 


4. Withdrawal of the proposals in regard to three specific names 
submitted with Voting Paper V.P.(O.M.)(55)28 : Consequential 
upon a decision to withdraw from the scope of V.P.(O.M.)(55)27 
the proposals submitted therewith in relation to the names 
Pandalus Leach 1815, Pinnotheres Latreille [1802—1803] and 
Portumnus Leach, 1814, in order to permit of the consideration 
of certain new information received in regard to those names, 
the Secretary on 7th October 1955 executed the following Minute 
withdrawing from the scope of Voting Paper V.P.(O.M.)(55)28 
certain supplementary proposals relating to the foregoing names 
for the purpose of enabling the Commission to deal simultaneously 
with all outstanding problems in regard to these names when it 
should have before it revised proposals prepared in the light of 
the additional information referred to above :— 


Withdrawal of proposals relating to three specific names from the 
scope of Voting Paper V.P.(O.M.)(55)28 


MINUTE by FRANCIS HEMMING, C.M.G., C.B.E., 


(Secretary to the International Commission on Zoological Nomenclature) 


In the light of additional information now furnished by Dr. L. B. 
Holthuis I have this day by a Minute numbered Z.N.(S.) 985 withdrawn 
from the scope of Voting Paper V.P.(O.M.)(55)27 certain proposals 
submitted therewith in relation to the names Pandalus Leach, 1815, 
Pinnotheres Latreille, [1802—1803] and Portumnus Leach, 1814, in 
order to permit of the further study of the issues involved before 
decisions were taken by the Commission thereon. In completion of 
the action taken in the foregoing Minute I now, as Secretary, withdraw 
from the scope of Voting Paper V.P.(O.M.)(55)28 certain other pro- 
posals regarding the foregoing names for the purpose of enabling all 
aspects of the questions outstanding in regard to the above names to be 
taken into consideration by the Commission simultaneously. 


5. The Prescribed Voting Period : As Voting Paper V.P.(O.M.) 
(55)28 was issued under the One-Month Rule, the Prescribed 
Voting Period therefor closed on 19th October 1955. 


6. Addition to the ‘‘ Official Index of Rejected and Invalid 
Generic Names in Zoology ’”’ of six additional junior homonyms 


80 OPINIONS AND DECLARATIONS _ 


of the name ‘‘Acanthopus ”’ Klug, 1807 : On Ist November 1955, 
Mr. Hemming, as Secretary, placed the following Minute on the 
Commission’s File Z.N.(S.) 987 :— 


Addition to the ‘* Official Index of Rejected and Invalid Generic Names 
in Zoology ’’ of certain junior homonyms of ‘‘Acanthopus ”’ Klug, 
1807, additional to that specified in Annexe 3 to the paper 

submitted with Voting Paper V.P.(O.M.)(55)28 


MINUTE by FRANCIS HEMMING, C.M.G., C.B.E., 


(Secretary to the International Commission on Zoological Nomenclature) 


In returning his copy of Voting Paper V.P.(O.M.)(55)28, 
Mr. Sylvester-Bradley drew attention to the fact that in Annexe 3 to 
the paper numbered Z.N.(S.) 987 submitted with the foregoing Voting 
Paper! one only of a number of junior homonyms of Acanthopus 
Klug, 1807, had been put forward for addition to the Official Index of 
Rejected and Invalid Generic Names in Zoology and expressed the view 
that all of these ought now to be placed on the above Official Index. 


2. Having regard to the Directive issued to the Commission by the 
Thirteenth International Congress of Zoology, Paris, 1948, that all 
objectively invalid names dealt with in, or otherwise involved in con- 
nection with, cases submitted to the Commission for decision be placed 
on the appropriate Official Index, 1 now, as Secretary, hereby direct 
that the following names each of which is a junior homonym of 
Acanthopus Klug, 1807, be placed on the Official Index of Rejected and 
Invalid Generic Names in Zoology and that the said additions to the 
Official Index be included in the Ruling in the Direction embodying 
the decisions of the Commission in its vote on Voting Paper 
V.P.(O.M.)(55)28 :— 


(a) Acanthopus Oken, 1816, Lehrbuch Naturgesch. 3(2) : ui, 122 
(b) Acanthopus Dahl, 1823, Col. u. Lepid. : 45 


(c) Acanthopus Latreille, 1829, in Cuvier (G.L.C.F.D.), Régn. anim. 
(ed: 2) 5 2138 


(d) Acanthopus Muenster, 1839, Beitr. Petref. (A) : 94 
(e) Acanthopus Giebel, 1872, Thesaur. Orn. 1 : 260 


(f) Acanthopus Vernet, 1877, Arch. Sci. phys. nat. Genéve (n.s.) 
60 : 334 


ou ee eee 


1 Kor the text of the paper here referred to see the first paragraph of the present 
Direction. 


DIRECTION 37 8! 


7. Particulars of the Voting on Voting Paper V.P.(O.M.)(55)28 : 
At the close of the Prescribed Voting Period, the state of the 
voting on Voting Paper V.P.(O.M.)(55)28 was as follows :— 


(a) Affirmative Votes had been given by the following twenty- 
four (24) Commissioners (arranged in the order in which 
Votes were received) : 


Boschma; Holthuis; Stoll; Mayr; Riley; Vokes ; 
Hering ; Jaczewski ; Prantl ; Lemche ; Mertens ; Bonnet ; 
do Amaral ; Hank6o ; Esaki ; Key ; Dymond ; Cabrera ; 
Hemming ; Miller? : Bradley (J.C.) ; Kiihnelt ; Sylvester- 
Bradley ; Tortonese ; 


(b) Negative Votes : 


None ; 


(c) On Leave of Absence, one (1) : 


Bodenheimer ; 


(d) Voting Papers not returned : 


None. 


8. Declaration of Result of Vote: On Ist November 1955, 
Mr. Hemming, Secretary to the International Commission, acting 
as Returning Officer for the Vote taken on Voting Paper 
V.P.(O.M.)(55)28, signed a Certificate that the Votes cast were as 
set out in paragraph 7 above and declaring that the proposal 
submitted in the foregoing Voting Paper had been duly adopted 
and that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


9. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 29th December 1955, Mr. Hemming prepared the Ruling 
given in the present Direction and at the same time signed a 
Certificate that the terms of that Ruling were in complete accord 


* Commissioner Miller exercised in this case the right conferred by the Thirteenth 
International Congress of Zoology in Paris, 1948, under which a Commissioner 
may, if he so desires, signify his willingness to support the view or the majority 
view. of other members of the Commission (1950, Bull. zool. Nomencl. 4: 50—S1). 


82 OPINIONS AND DECLARATIONS 


with those of the proposal approved by the International Com- 
mission in its Vote on Voting Paper V.P.(O.M.)(55)28, subject 
(a) to the withdrawal therefrom of the proposals relating to the 
two generic names specified in the Minute executed by the Secretary 
on 7th October 1955, and (b) to the addition to the Official Index 
of Rejected and Invalid Generic Names in Zoology of the junior 
homonyms specified in the Secretary’s Minute of Ist November 
1955. The Minutes referred to above have been reproduced in 
paragraph 4 and paragraph 6 respectively of the present Direction. 


10. Original References : The original references for the generic 
names specified in Rulings (1) and (2) of the present Direction 
have already been noted in connection with the Official List of 
Generic Names in Zoology. The references for the names specified 
in Ruling (3) have been similarly noted except in so far as they 
have been supplemented by additional information given in 
Annexe 2 to the application reproduced in the first paragraph 
of the present Direction. The original references for the names 
specified in Rulings (4) and (5) are given in Annexe 3 to the paper 
referred to above, with the exception of those for six junior 
homonyms of Acanthopus Klug, 1807, the references for which are 
given in the Minute executed by the Secretary on Ist November 1955, 
the text of which is given in paragraph 6 of the present Direction. 


11. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in dealing 
with the present case, and the present Direction is accordingly 
hereby rendered in the name of the said International Commission 
by the under-signed Francis Hemming, Secretary to the Inter- 
national Commission on Zoological Nomenclature, in virtue of 
all and every the powers conferred upon him in that behalf. 


12. The present Direction shall be known as Direction Thirty- 
Seven (37) of the International Commission on Zoological 
Nomenclature. 


Done in London, this Twenty-Ninth day of December, Nineteen 
Hundred and Fifty-Five. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


Printed in England by Mretcatre & Cooper LiMiTED, 10-24 Scrutton St., London EC 2 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, c.M.G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.3. Pp. 83—94 


DIRECTION 38 


Rejection for nomenclatorial purposes of the anonymously 

published work entitled Ornithologia britannica issued 

in 1771 and believed to have been written by Marmaduke 

Tunstall and matters incidental thereto (cancellation of 
Opinion 38) 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956 


Price Eight Shillings 
(All rights reserved) 


Issued 1st September, 1956 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 38 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JorDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President: Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., 
U.S.A.) (12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


Arranged in order of precedence by reference to date of election or of most recent 
re-election, as prescribed by the International Congress of Zoology 


Professor H. BoscHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CaBRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. Henning Lemcue (Universitetets Zoologiske Museum, Copenhagen, Denmark) 
(27th July 1948) 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEY (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a. M., Germany) (5th July 1950 

Professor Erich Martin HERING) (Zoologisches Museum der Humboldt-Universitat zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMOND (University of Toronto, Toronto, Canada) (12th August 1953) 

ee 4; ics BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) (12th August 1953) 

resident 

Professor Harold E. Voxes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) 
(12th August 1953) E 

Professor Béla HANKO (Mezdgazdasdgi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. SToxt (Rockefeller Institute for Medical Research, New York, N.Y., U.S.A.) 
(12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. Hottuuts (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (15th October 1954) ; 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, Berkeley, 
California, U.S.A.) (29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum v Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KiiHNFLT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

ak Hy F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 
1954 

Professor Ernst MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 

Massachusetts, U.S.A.) (4th December 1954 

Professor Enrico TORTONESE (Museo di Storia Naturale “‘ G. Doria’, Genova, Italy) 

(16th December 1954) 


DIRECTION 38 


REJECTION FOR NOMENCLATORIAL PURPOSES OF 
THE ANONYMOUSLY PUBLISHED WORK ENTITLED 
**ORNITHOLOGIA BRITANNICA”? [ISSUED IN 1771 
AND BELIEVED TO HAVE BEEN WRITTEN BY 
MARMADUKE TUNSTALL AND MATTERS 
INCIDENTAL THERETO (CANCELLATION OF 
** OPINION ”’ 38) 


RULING :—(i) It is hereby ruled that in the anonymous 
work published in 1771 under the title Ornithologia 
britannica the author, who is reputed to have been 
Marmaduke Tunstall, did not apply the principles of 
binominal nomenclature, that in consequence no new 
name published in the foregoing work acquired the 
status of availability in virtue of being so published and 
therefore that the acceptance of that work for nomen- 
clatorial purposes signified in Opinion 38 was incorrect. 


(2) In view of (1) the Ruling given in Opinion 38 1s 
hereby cancelled for all except historical purposes. 


(3) The title of the work specified in (1) above is hereby 
placed on the Official Index of Rejected and Invalid Works 
in Zoological Nomenclature with the Title No. 48. 


(4) Attention is drawn to the action taken by the 
Commission by the Ruling given in Opinion 404 under 
which the generic name Pyrrhocorax [Tunstall], 1771, 
the only new generic name published in the Ornitho- 
logia britannica, was validated under the Plenary Powers 
and placed on the Official List of Generic Names in 
Zoology. 


SEP 9 9 1958 


86 OPINIONS AND DECLARATIONS 


(5) The under-mentioned generic names are hereby 
placed on the Official Index of Rejected and Invalid 
Generic Names in Zoology with the Name Numbers— 
severally specified below :— 


(a) Ampelis Nozeman & Vosmaer, 1758 (a name 
published in a work rejected for nomenclatorial 
purposes!) (Name No. 518) ; 


(b) Columbus [Tunstall], 1771 (an Erroneous Subse- 
quent Spelling for Columba Linnaeus, 1758) 
(Name No. 519) ; 


(c) Columbus Linnaeus, 1758 (an Invalid Original 
Spelling for Colymbus Linnaeus, 1758) (Name 
No. 520). 


(6) The under-mentioned generic name is _ hereby 
placed on the Official List of Generic Names in Zoology 
with the Name No. 1015 :—Columba Linnaeus, 1758 
(gender : feminine) (type species, by selection by Vigors 
(1825) : Columba oenas Linnaeus, 1758). 


(7) The under-mentioned specific name is hereby placed 
on the Official List of Specific Names in Zoology with the 
Name No. 944 :—oenas Linnaeus, 1758, as published in 
the combination Columba oenas (specific name of type 
species of Columba Linnaeus, 1758). 


I. THE SUBJECT MATTER OF THE PRESENT 
DIRECTION * 


The present Direction contains a revised Ruling regarding the 
status for nomenclatorial purposes of the anonymously published 
work entitled Ornithologia britannica commonly attributed to 
Marmaduke Tunstall which was published in 1771. It will be 


' For the rejection for nomenclatorial purposes of Nozeman & Vosmaer, 1758, 
Geslachten der Vogelen, see Opinion 241 (1954, Ops. Decls. int. Comm. zool. 
Nomencl. 4 : 13—21). 


DIRECTION 38 87 


recalled that, when on Ist September 1955 the Commission was 
invited on Voting Paper V.P.(O.M.)(55)22 to codify the Rulings 
in regard to the status of individual books and papers given by 
it in Opinions rendered up to the end of 1936, it was explained 
that further investigations were in progress in regard to the fore- 
going work and that in consequence it was not proposed on that 
occasion to submit any proposal for the codification of the Ruling 
given in Opinion 38, the Opinion in which in 1912 the Com- 
mission had previously dealt with the present subject?. For 
various reasons the issue of the Voting Paper referred to above 
did not take place at as early date as had originally been expected 
with the result that the above Voting Paper was issued concurrently 
with the Voting Paper covering the Secretary’s promised Supple- 
mentary Report on the Ornithologia britannica of 1771. That 
Report, which was also submitted on Ist September 1955, was as 
follows :— 


Proposed cancellation of ‘‘ Opinion ”’ 38 and matters 
incidental thereto 


By FRANCIS HEMMING, C.M.G., C.B.E., 


(Secretary to the International Commission on Zoological Nomenclature) 


Under decisions taken by the Fourteenth International Congress 
of Zoology, Copenhagen, 1953 (Copenhagen Decisions zool. Nomencl. 
24, Decision 24 ; ibid. : 23, Decision 23), an Official List of Works 
Approved as Available for Zoological Nomenclature and an Official 
Index of Rejected and Invalid Works in Zoological Nomenclature were 
. established and the Commission was instructed to place on this new 
Official List and Official Index the titles of works which it had, in 
the one case, accepted and, in the other case, rejected for nomen- 
clatorial purposes. The Commission has already complied with the 
above Directive as regards all Opinions rendered since 1936. Proposals 
are now being submitted?, in connection with Voting Paper V.P.(O.M.) 
(55)22, to enable it to do so in respect of all Opinions rendered before the 
above date, except in the present case which it was considered could 
be more conveniently dealt with separately, and four others, which for 
various reasons are also dealt with separately. 


2 For the decision taken by the International Commission on Voting Paper 
V.P.(O.M.)(55)22 see Direction 32 (1956, Ops. Decls. int. Comm. zool. Nomencl. 
1(C) : 307—328. 


88 OPINIONS AND DECLARATIONS 


2. Opinion 38, with which the present paper is concerned, was 
published in 1912 (Smithson. Publ. 2060 : 89—90) under the title 
‘On the Status of the Latin Names in Tunstall, 1771’. The work 
dealt with is a very slim large folio consisting of six unnumbered 
pages (T.P. and Verso blank, [1]—[4]). Its full title is as follows :— 
‘** Ornithologia Britannica: seu Avium omnium Britannicarum tam 
Terrestrium quam Aguaticarum Catalogus, Sermone Latino, Anglico, 
& Gallico redditus : cui subjicitur Appendix, Aves alienigenas, in Angliam 
rara advenientes, complectens. The Title Page states that this work 
was “* Printed for the Author by J. Dixwell in St. Martin’s Lane” 
and bears the date “‘ 1771”. The work is anonymous but is attributed 
to Marmaduke Tunstall (1743—1790). 


3. This book consists entirely of a tabular presentation of the names 
for the British birds known to the author. The entry in these tables 
for each species contains four columns devoted to (i) the Latin name of 
the genus, (ii) a Latin name for the species, (iii) an English vernacular 
name, (iv) a French vernacular name. The work has no pretention to 
originality, being, as 1s clearly shown by the following footnote, no 
more than a compilation from earlier works :—‘‘ Nomina Latina vel 
ex Linnaeo vel ex ultima editione Zoologiae Britannicae, Gallica vero 
.ornithologia Brissonii plerumque decerpta sunt ’’. No bibliographical 
references are given for any of the 53 generic names cited and very few 
such references are supplied for the names used for species, there being 
references for only 14 of the Latin specific names and for one of 
the English names cited. Thus for 226 out of the 241 “species ” 
recognised no bibliographical references are supplied. For fourteen 
of the species cited, polyverbal Latin specific names are employed. 


4. It was recognised by the Commission that in this book Tunstall 
did not apply the principles of binominal nomenclature, it being stated 
in the Section of Opinion 38 headed “ Discussion ”’ that “‘ the nomen- 
clature is binary”. That the Commission felt able to accept names 
in Tunstall’s Ornithologia as possessing the status of availability was 
due to the fact that at that time it was guiding itself by the interpretation . 
of Proviso (b) to Article 25 which it had given in 1910 in Opinion 20. 
In that Opinion it had ruled that the generic names but not the specific 
names, in the Zoophylacium Gronovianum of Gronovius were available, 
because, although that author had not applied the principles of 
binominal nomenclature, he had used a “ binary’ system of nomen- 
clature, that is, he had realised that the scientific name for an animal 
must be so constructed as to recognise the twofold concept of genus 
and species. In Paris in 1948 the International Congress of Zoology 
rejected as defective the Ruling given in Opinion 20 and underlined its 
decision in this matter by substituting the expression “‘ nomenclature 
binominale’’ for the expression “‘ nomenclature binaire’’ wherever 
it occurred in the Régles. Following this decision the Commission 
during its Paris Session dealt with a number of its older Opinions in 
which the so-called “‘ binary’’ system of nomenclature had been 


DIRECTION 38 89 


recognised as conferring availability upon names published in the works 
concerned. Tunstall’s Ornithologia had not then been re-examined 
from the foregoing point of view, and accordingly the disposal of 
generic names published in that work was left over for later considera- 
tion. 


5. It is necessary therefore to examine the generic names used by 
Tunstall, to see which of them require to be validated, now that, the 
Ornithologia, contrary to the statement in Opinion 38, is found to be 
unavailable for nomenclatorial purposes. It is here that a paradoxical 
situation is disclosed, for of the 46 generic names cited by Tunstall, 
41 were first published by Linnaeus in 1758, 3 by Brisson in 1760, 
and one by Linnaeus in 1766. Thus one new generic name only was 
published in this work. The names concerned are :— 


¢1) Names published by Linnaeus, 1758, forty-one (41) : 

Palco : Strix ; Lanius.; Corvus ; Picus ;.Jynx ; Cuculus ; Sitta ; 
Alcedo ; Certhia; Tetrao; Otis; Columbus ; (ex err. pro 
Columba) ; Turdus ; Sturnus ; Alauda ; Hirundo ; Motacilla ; 
Loxia; Fringilla; Emberiza; Parus; Ardea; Scolopax ; 
Tringa ; Haematopus ; Charadrius ; Rallus ; Fulica ; Colymbus ; 
Recurvirostra ; Alca; Mergus ; Larus ; Sterna; Procellaria ; 
Anas ; Pelecanus ; Coracias ; Upupa ; Platalea ; 


(2) Names published by Brisson, 1760, three (3) : 
Gallinula ; Phaleropus ; Merganser ; 


(3) Name published by Linnaeus, 1766, one ( a 
Ampelis ; 


(4) Name first published in Tunstall’s *‘ Ornithologia’’, one (1) : 
Pyrrhocorax. 


6. We see therefore that the only purpose to be served by validating 
Tunstall’s Ornithologia would be to provide a valid basis for the 
name Pyrrhocorax as the generic name for the Chough. That action 
is not now needed even for this purpose since in 1954 the Commission 
by its vote on Voting Paper V.P.(54)69 used its Plenary Powers to 
validate the name Pyrrhocorax [Tunstall] when dealing with Application 
Z.N.(S.) 492 (1952, Bull. zool. Nomencl. 9 : 53—61)?. 


7. Both the name Pyrrhocorax and the name Ampelis were published 
in 1758 in Nozeman & Vosmaer’s Ges/l. Vogel. : 1, 15 (Pyrrhocorax) 
2, 25 (Ampelis) and, as Tunstall’s Ornithologia makes no pretension to 
being an original work, it is likely that Tunstall obtained these two 
names, if not direct from Nozeman & Vosmaer, at least from some book 


3 The decision taken by the Commission on Application Z.N.(S.)492 (Pyrrho- 
corax) has since been embodied in Opinion 404 (1956, Ops. Decls. int. Comm, 
zool. Nomencl. 13 : 87—106). 


90 OPINIONS AND DECLARATIONS 


containing a reference to these names. Nozeman & Vosmaer were not 
binominal authors and the work referred to above has for this reason 
been rejected by the Commission in Opinion 241 (1954, Ops. Decls. int. 
Comm. zool. Nomencl. 5 : 13—22). Since the two names referred to 
above found their way into Neave’s Nomenclator, they are liable to 
mislead the unwary and it is desirable therefore that they should both 
now be placed on the Official Index of Rejected and Invalid Generic 
Names in Zoology. 


8. It will be observed that in paragraph 5(1) above the name Columbus, 
as used by Tunstall,is treated as being a variant, accidental or intentional, 
of Columba Linnaeus, 1758. That this interpretation is correct is 
shown by the fact that the three species cited by Tunstall under 
Columbus (Stock-Dove ; Ring-Dove ; Turtle-Dove) were all included 
by Linnaeus in his genus Columba. It is desirable that the name 
Columbus |Tunstall] be now disposed of by being placed on the Official 
Index as an Erroneous Subsequent Spelling. At the same time the 
generic name Columba Linnaeus, 1758 (Syst. Nat. (ed. 10) 1 : 162) 
(gender : feminine) (type species, by selection by Vigors, 1825 (Trans. 
linn. Soc. Lond. 14 : 481) : Columba oenas Linnaeus, 1758, Syst. Nat. 
(ed. 10) 1 : 162) should be placed on the Official List of Generic Names 
in Zoology, and the name oenas Linnaeus, 1758, as published in the 
foregoing combination, should be placed on the Official List of Specific 
Names in Zoology. Finally, it would be well to take note that there is 
an even earlier invalid usage of the name Columbus, namely, that by 
Linnaeus himself in 1758 in his preliminary section entitled “‘ Classis 
II. Aves. Caracteres Avium ”’ (: 84) where he used this spelling for the 
genus numbered “ 68”? which on page 134 he dealt with fully under 
the name Colymbus. This Invalid Original Spelling should now be 
placed on the Official Index. (No action is required in connection 
with the Valid Original Spelling Colymbus Linnaeus, 1758, for that name 
has already formed the subject of a decision by the Commission in its 
vote on Voting Paper V.P.(54)62)?. 


9. The specific names cited by Tunstall have not been examined in 
detail, for even under the erroneous Ruling given in Opinion 20 
(relating to the Zoophylacium of Gronovius), on which the Ruling 
given in Opinion 38 was based, the generic names only, of “* binary but 
not binominal ”’ authors, were acceptable. An inspection of Tunstall 
shows, it may be noted, that the great majority, if not all, the specific 
names which he cited in binominal combinations, as contrasted with 
his polyverbal specific names, were derived from Linnaeus. If any 
specific name were to be found to be currently accepted as from 


4 The decision in regard to the generic name Co/ymbus Linnaeus, 1758, taken by 
the International Commission in its vote on Voting Paper V.P.(54)62 has since 
been embodied in Opinion 401 (1956, Ops. Decls. int. Comm. zool. Nomencl. 
13 : 1—64). 


DIRECTION 38 ee | | 


Tunstall’s Ornithologia, it would be necessary to treat that name as 
constituting a separate case, for that usage would not have been 
justified under the Ruling given in Opinion 38, even if that Ruling 
had been correct which, as shown in paragraph 4 above, it is now seen 
not to have been. 


10. In the light of the particulars furnished above, I recommend 
the International Commission :— 


(1) to take note :— 


(a) that in the anonymous work published in 1771 under the 
title Ornithologia britannica, the author, who is believed 
to have been Marmaduke Tunstall, did not apply the 
principles of binominal nomenclature and therefore that 
the names published in the above work do not satisfy the 
requirements of Proviso (b) to Article 25 and accordingly 
that the acceptance of the above work for nomenclatorial 
purposes by the Ruling given in Opinion 38 was incorrect ; 


(b) that in the above work the only new generic name 
introduced was the name Pyrrhocorax which has already 
been validated under the Plenary Powers by the vote 
taken by the Commission on Voting Paper V.P.(54)69, 
when dealing with Application Z.N.(S.) 492 which was 
expressly concerned with that name’®; 


(2) for the reasons specified in (1) above, to cancel Opinion 38 for 
all except historical purposes ; 


(3) to place the under-mentioned title on the Official Index of 
Rejected and Invalid Works in Zoological Nomenclature :— 
[Tunstall (M.)], 1771, Ornithologia britannica ; 


(4) to place the under-mentioned generic names on the Official 
Index of Rejected and Invalid Generic Names in Zoology :— 


(a) Pyrrhocorax Nozeman & Vosmaer, 1758 (a name published 
in a work rejected for nomenclatorial purposes) ; 


(b) Ampelis Nozeman & Vosmaer, 1758 (a name published in 
a work rejected for nomenclatorial purposes) ; 


(c) Columbus [Tunstall], 1771 (an Erroneous Subsequent 
Spelling for Columba Linnaeus, 1758) ; 


(d) Columbus Linnaeus, 1758 (an Invalid Original Spelling 
for Colymbus Linnaeus, 1758) ; 


(5) to place the under-mentioned generic name on the Official List 
of Generic Names in Zoology :—Columba Linnaeus, 1758 


> See footnote 3. 


92 OPINIONS AND DECLARATIONS 


(gender: feminine) (type species, by selection by Vigors 
(1825) : Columba oenas Linnaeus, 1758) ; 


(6) to place the under-mentioned specific name on the Official 
List of Specific Names in Zoology :—oenas Linnaeus, 1758, 
as published in the combination Columba oenas (specific name 
of type species of Columba Linnaeus, 1758). 


2. Registration of the present application : Upon the receipt of 
Mr. Hemming’s Report, the question whether the International 
Commission should give a revised Ruling on the status of 
Tunstall’s Ornithologia britannica of 1771 was allotted the 
Registered Number Z.N.(S.) 944. 


II. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


3. Issue of Voting Paper V.P.(O.M.)(55)23 : On Ist September 
1955, a Voting Paper (V.P.(O.M.)(55)23) was issued in which 
each Member of the Commission was asked (1) to state whether 
he agreed that, ‘“‘ in conformity with the General Directive relating 
to the recording on the various Official Lists and Official Indexes 
of decisions in regard to particular names and particular books 
issued to the International Commission by the Thirteenth Inter- 
national Congress of Zoology, Paris, 1948, and with the General 
Directive supplementary thereto issued to the Commission by 
the Fourteenth International Congress of Zoology, Copenhagen, 
1953, the entries on Official Lists and Official Indexes specified 
in paragraph 10 of the paper in regard to Tunstall’s Ornithologia 
britannica of 1771 numbered Z.N.(S.) 944 submitted by the 
Secretary simultaneously with the present Voting Paper [i.e. in 
the paragraph numbered as above in the application reproduced 
in the first paragraph of the present Direction], together with the 
other recommendations specified in that paragraph, be approved ”’, 
and (2), if he did not so agree as regards any given item, to indicate 
the item concerned. 


4. The Prescribed Voting Period : As the foregoing Voting 
Paper was issued under the One-Month Rule, the Prescribed 
Voting Period closed on Ist October 1955. 


DIRECTION 38 93 


5. Particulars of the Voting on Voting Paper V.P.(O.M.)(55)23 : 
At the close of the Prescribed Voting Period, the state of the 
voting on Voting Paper V.P.(O.M.)(55)23 was as follows :— 


(a) Affirmative Votes had been given by the following twenty- 
five (25) Commissioners (arranged in the order in which 
Votes were received) : 


Bodenheimer ; Holthuis; Lemche; Hering; Riley ; 
Mayr ; Vokes; Miller ; Prantl; Key ; Esaki; Bradley 
(J.C.) ; Cabrera ; Stoll ; do Amaral ; Boschma ; Kiihnelt ; 
Hemming; Dymond; Sylvester-Bradley ; Jaczewski ; 
Tortonese ; Mertens ; Bonnet ; Hanko ; 


(b) Negative Votes : 


None ; 


(c) Voting Papers not returned : 


None. 


6. Declaration of Result of Vote: On 4th October 1955, 
Mr. Hemming, Secretary to the International Commission, 
acting as Returning Officer for the Vote taken on Voting Paper 
V.P.(O.M.)(55)23, signed a Certificate that the Votes cast were 
as set out in paragraph 5 above and declaring that the proposal 
submitted in the foregoing Voting Paper had been duly adopted 
and that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


7. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 9th January 1956, Mr. Hemming prepared the Ruling given 
in the present Direction and at the same time signed a Certificate 
that the terms of that Ruling were in complete accord with those 
of the proposal approved by the International Commission in 
its Vote on Voting Paper V.P.(O.M.)(55)23, subject to the exclusion 
from the said Ruling of the addition of the name Pyrrhocorax 
Nozeman & Vosmaer, 1758, to the Official Index of Rejected and 
Invalid Generic Names in Zoology in view of the fact that in the 
meantime the said action had been taken by the Ruling given 
in Opinion 4048. 


® See Footnote 3 of the present Direction. 


94 OPINIONS AND DECLARATIONS 


8. Original References : The following are the original refer- 
ences for the names placed on. Official Lists and Official Indexes 
by the Ruling given in the present Direction :— 


Ampelis Nozeman & Vosmaer, 1758, Geslacht. Vogel. (Dutch 
translation of Moehring, 1752, Avium Genera) : 1, 2, 25 

Columba Linnaeus, 1758, Syst. Nat. (ed. 10) 1 : 162 

Columbus Linnaeus, 1758, Syst. Nat. (ed. 10) 1 : 84 

Columbus [Tunstall], 1771, Ornith. brit. : [2] 

oenas, Columba, Linnaeus, 1758, Syst. Nat. (ed. 10) 1 : 162 


9. The following is the reference for the selection of a type 
species for Columba Linnaeus, 1758, specified in the Ruling given 
in the present Direction :—Vigors, 1825, Trans. linn. Soc. Lond. 
14 : 481. , 


10. Family-group-name position : The generic name Columba 
Linnaeus, 1758, placed on the Official List of Generic Names in 
Zoology by the Ruling given in the present Direction is the type 
genus of the well-known family COLUMBIDAE. It has not so far 
been possible to determine where this family-group name was first 
published. This matter is at present being investigated on a File 
numbered Z.N.(G.) 133, which has been specially opened for this 
purpose. 


11. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in dealing 
with the present case, and the present Direction 1s accordingly 
hereby rendered in the name of the said International Com- 
mission by the under-signed Francis Hemming, Secretary to the 
International Commission on Zoological Nomenclature, in virtue 
of all and every the powers conferred upon him in that behalf. - 


12. The present Direction shall be known as Direction Thirty- 
Eight (38) of the International Commission on Zoological 
Nomenclature. 


Done in London, this Ninth day of January, Nineteen Hundred 
and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


——S 
Printed in England by MercaLFe & Cooper Limitep, 10-24 Scrutton St. London EC 2 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, c.M.G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.4. Pp. 95—138, I pl. 


DIRECTION 39 


Substitution of Gallinago Brisson, 1760, for Gallinago 

Koch, 1816 (Class Aves) on the Official List of Generic 

Names in Zoology (correction of an error in the Ruling 
given in Opinion 67) 


EMNTHSON 
OCT 16 1955 


LONDON: 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956 


Price Twenty-Nine Shillings 
(All rights reserved) 


Issued 17th September, 1956 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 39 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JoRDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President : Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) 
(12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953). 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(arranged in order of precedence by reference to date of election or of most recent re-election 
as prescribed by the International Congress of Zoology) 


Professor H. BoscHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. care pct (Universitetets Zoologiske Museum, Copenhagen, Denmark) (27th 
uly 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEy (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (5th July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitat zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMoNnD (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) (12th August 
1953) (President) ; 

Professor Harold E. Voxes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) (2th 
August 1953) 

Professor Béla HANK6 (Mezégazdasdgi Muzeum, Budapest, Hungary) (i2th August 1953) 

Dr. Norman R. StToLt (Rockefeller Institute for Medical Research, New York, N.Y., 
U.S.A.) (12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. Hoituuts (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 

Canberra, A.C.T., Australia) (15th October 1954 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum V Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KUHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

. ea S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 
1954 

Professor Ernest MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico TorTcwEsE (Museo do Storia Naturale “‘ G. Doria’’, Genova, Italy) 
(16th December 1954) 


eS 


DIRECTION 39 


SUBSTITUTION OF ‘*‘GALLINAGO ” BRISSON, 1760, 
FOR ‘*“ GALLINAGO ”? KOCH, 1816 (CLASS AVES) ON 
THE ‘* OFFICIAL LIST OF GENERIC NAMES IN 
ZOOLOGY ” (CORRECTION OF AN ERROR IN 
THE RULING GIVEN IN ‘“* OPINION ” 67) 


RULING :—(1) The following entry is to be inserted 
in the Official List of Generic Names in Zoology in place 
of the name Gallinago Koch, 1816, made by the Ruling 
given in Opinion 67 :— 


Gallinago Brisson, 1760 (gender : feminine) (type 
species, by absolute tautonymy : Scolopax gallinago 
Linnaeus, 1758) (Name No. 58). 


(2) The under-mentioned specific name is hereby 
placed on the Official List of Specific Names in Zoology 
with the Name No. 945: gallinago Linnaeus, 1758, as 
published in the combination Scolopax gallinago (specific 
name of type species of Gallinago Brisson, 1760). 


(3) The under-mentioned generic names are hereby 
placed on the Official Index of Rejected and Invalid 
Generic Names in Zoology with the Names Numbers 
521 to 523 respectively :— 


(a) Gallinago Koch, 1816 (a junior homonym of, and 
a junior objective synonym of, Gallinago Brisson, 
1760) ; 


(b) Capella Frenzel, 1801 (a name published in 
synonymy without an independent description 
and rejected by its author) ; 


(c) Capella Keyserling & Blasius, 1840 (a junior 
objective synonym of Rupicapra Blainville, 1816). 


art 171 1956 


98 OPINIONS AND DECLARATIONS 


(4) The under-mentioned specific name is hereby 
placed on the Official Index of Rejected and Invalid 
Specific Names in Zoology with the Name Number 341 : 
media Koch, 1816, as published in the combination 
Gallinago media (a junior objective synonym of gallinago 
Linnaeus, 1758, as published in the combination Scolopax 
gallinago). 


I—THE SUBJECT MATTER OF THE PRESENT 
“DIRECTION |: 


On 23rd August 1952 Mr. Francis Hemming, as Secretary, 
submitted the following paper to the Commission in which he 
drew attention to an erroneous entry regarding the generic 
name Gallinago Koch, 1816 (Class Aves) made in the Official 
List of Generic Names in Zoology by the Ruling given in Opinion 
67 published in 1916 :— 


Proposed use of the Plenary Powers to suppress the name ‘* Gallinago ”’ 
Brisson, 1760 (Class Aves), and proposed substitution of ‘‘ Capella ”’ 
Frenzel, 1801, for ‘‘ Gallinago ’’ Koch, 1816, on the ‘‘ Official 
List of Generic Names in Zoology ”’ (proposed correction 
of an Erroneous Entry in ‘‘ Opinion ”’ 67) 


By FRANCIS HEMMING, C.M.G., C.B.E. 
(Secretary to the International Commission on Zoological Nomenclature) 
The subject matter of the present application came to notice in the 


course of the checking of the entries on the Official List of Generic 
Names in Zoology in connection with the projected publication of the 


Official List in book form and is concerned with the generic name » 


Gallinago Koch, 1816 (Syst. baier. Zool. 1 : 312) which was placed 
on the Official List in Opinion 67 published in 1916 (Smithson. Publ. 
2409 : 180). 


DIRECTION 39 99 


2. The generic name Gallinago Koch, 1816, was stated in Opinion 67 
to have as its type species, Scolopax gallinago Linnaeus, 1758 (Syst. 
Nat. (ed. 10) 1 : 147) by absolute tautonymy, through the ‘* media of 
Koch ” (i.e. Gallinago media Koch, 1816, loc. cit. 1 : 314). This name 
was therefore a generic name applied to the Common Snipe. 


3. When checking this part of Opinion 67, I observed that Hartert, 
after using the generic name Gallinago Koch for the Snipe in the main 
portion of his work relating to the genus concerned (Hartert, 1916, 
Vogel paldarkt. Fauna (2) : 1655), had later in the same work (1921, 
ibid. (3) : 2213) published a correction pointing out that Gallinago, 
Koch was a junior synonym of Capella Frenzel, 1801 (Beschr. Végel 
Wittenberg : 58), the type species of which was the nominal species 
Capella coelestis Frenzel, 1801 (ibid. : 58), a nominal species which 
represented the same taxonomic species as did the nominal species 
Scolopax gallinago Linnaeus. I noted also that my colleague Com- 
missioner James L. Peters (1934, Check List Birds World 2 : 274) 
accepted the name Capella Frenzel, sinking Gallinago Koch as a 
synonym. 


4. As it was clearly not possible in these circumstances to leave the 
name Gallinago Koch on the Official List without prior re-submission 
to the International Commission, I wrote a letter (on 14th October 
1945) to Dr. Peters asking for his views as to the action which it was 
desirable should be taken. Dr. Peters in his reply (of 6th December 
1945) wrote: ‘‘ After Capella Frenzel was shown to be an earlier 
name than Gallinago Koch and of equal applicability, it was immediately 
adopted and is now in current use for the different species of Snipe. 
For this reason I believe the Gallinago should be expunged from the 
Official List and Capella substituted in its place. No useful purpose 
would be served by reinstating Gallinago Koch under suspension of the 
rules and suppressing Capella’”’ 


5. Quite recently I was led, for the reasons which I have explained 
in Application Z.N.(S.) 701 (relating to the generic names Bubo, 
Coturnix, Egretta, and Oriolus)! to examine carefully M. J. Brisson’s 
Ornithologie published in 1760, for I had already discovered that one 
name (Egretta Forster, 1817) that was already on the Official List 
of Generic Names in Zoology was an invalid junior homonym of a 
totally overlooked name published in Brisson’s Ornithologie. This 
search brought to light the existence of the name Gallinago Brisson, 
1760 (Ornithologie 5 : 298—310), which thus greatly antedates, 
the name Capella Frenzel, 1801. The name Gallinago was used by 
Brisson as the name for a subdivision of the genus Scolopax ; he 
placed in it all except the first of the five species which he referred to 
Scolopax. All the species described in the Ornithologie are described 


1 The decision regarding these names has since been taken by the Commission 
and incorporated in Direction 21 (Ops. Decls. int. Comm. Zool. Nomencl. 1, 
Section C : 161—178). 


100 OPINIONS AND DECLARATIONS 


initially under a French name, followed by a Latin diagnosis, at the 
end of which is given, in different type, the Latin name accepted by 
Brisson for the species in question, this in turn being followed, in 
the case of previously described species, by a detailed synonymy. The 
first of the species placed by Brisson in his Gallinago was cited under the 
French name “‘ La Beccassine ’’, the scientific name at the end of the 
Latin diagnosis being given simply as “‘ Gallinago’’ (not because 
Brisson was a mononominalist, but because he customarily cited in 
this way the names of species when the “‘ species ”’ portion of the name 
consisted of the same word (i.e. was a single word) tautonymous with 
the generic name). In the synonymy of this species Brisson quoted 
the diagnosis given by Linnaeus in 1758 for his Scolopax gallinago, 
finishing this quotation as follows: ‘“* Gallinago. Linn. Syst.Nat. ed. 
10 Gen. 77 sp. 11” (Scolopax being the 77th genus of birds in the 10th 
edition of the Syst. Nat. and Scolopax gallinago being the eleventh of 
the species referred by Linnaeus to this genus). Thus, we see clearly 
that this species is Scolopax gallinago Linnaeus, 1758 (: 147) and that, 
as its trivial name is tautonymous with the generic name selected by 
Brisson, it is the type species of Gallinago Brisson by absolute 
tautonymy. Accordingly, the name Gallinago Brisson, 1760, is not 
only a senior homonym of Gallinago Koch, 1816, but in addition is a 
senior synonym of Koch’s generic name, Brisson’s and Koch’s nominal 
genera each having the same nominal species as its type species. The 
position so established created a new situation and one under which the 
entry on the Official List relating to Gallinago Koch was not only 
(as previously) subjectively defective (because of the subjective identi- 
fication of the nominal species which are respectively the type species 
of Capella Frenzel, 1801, and of Gallinago Koch, 1816), but also 
objectively incorrect (through Gallinago Koch, being both an objective 
junior homonym, and an objective junior synonym, of Gallinago 
Brisson, 1760). Faced with this situation, it seemed to me that, since 
(as Dr. Peters had explained) the name Capella Frenzel had by now 
completely replaced the name Gallinago Koch, it would be confusing 
if now that transition had to be reversed, the name Capella Frenzel 
being displaced by Gallinago Brisson. 


6. At this stage I consulted Colonel Richard Meinertzhagen, 
Chairman of the Standing Committee on Ornithological Nomenclature 
who in his reply (dated 12th September 1952) wrote as follows: “* If, 
as at first appeared, the question to be considered in the case of the 
name Gallinago had been whether the name Gallinago Koch, 1816, 
should be replaced on the Official List by its senior subjective synonym 
Capella Frenzel, 1801, I should have been strongly in favour of that 
course, for now that Gallinago Koch has been completely replaced by 
the name Capella Frenzel, I should have thought it most unfortunate 
if, through Gallinago Koch being already on the Official List, it had 
been necessary to abandon current practice by reverting to the use of 
the name Gallinago Koch. Now that it appears that the oldest generic 
name for the Common Snipe is Gallinago Brisson, 1760, I hold the 


DIRECTION 39 101 


same view for the same reason. I accordingly consider that the best 
course will be for the International Commission to suppress Brisson’s 
Gallinago, to remove Koch’s Gallinago from the Official List and to 
insert in its place the name Capella Frenzel. The trivial name gallinago 
Linnaeus, 1758, as published in the combination Scolopax gallinago, 
should, I agree, now be placed on the Official List of Specific Trivial 
Names. Naturally, however, the trivial name coelestis Frenzel, 1801 
(the trivial name of the type species of Capella Frenzel) ought not to 
be placed on that Official List, for, although nomenclatorially it is an 
available name, it is only a junior synonym of gallinago Linnaeus 
and therefore can never be needed ”’. 


\ 


7. A settlement of this case is urgently required, for at present the 
problem presented by the name Gallinago Koch represents one of the 
obstacles which is holding up the publication of the Official List of 
Generic Names in Zoology in book form. In view of the advice received 
in this case—as set out in paragraphs 4 and 6 above—I recommend 
that the International Commission on Zoological Nomenclature 
should :— | 


(1) use its Plenary Powers to suppress the generic name Gallinago 
Brisson, 1760, for the purposes of the Law of Priority but not 
for those of the Law of Homonymy ; 


(2) delete the name Gallinago Koch, 1816, from the Official List of 
Generic Names in Zoology, at the same time correcting Opinion 
67 to the extent necessary ; 


(3) substitute on the foregoing Official List the name Capella Frenzel, 
1801 (type species, by monotypy: Capella coelestis Frenzel, 
1801) for the name proposed, under (2) above, to be removed 
therefrom ; 


(4) place the trivial name gallinago Linnaeus, 1758, as published in 
the combination Scolopax gallinago, on the Official List of 
Specific Trivial Names in Zoology ; 


(5S) place the under-mentioned names on the Official Index of 
Rejected and Invalid Generic Names in Zoology :— 


(a) Gallinago Brisson, 1760, as proposed, under (1) above, to 
be suppressed under the Plenary Powers ; 


(b) Gallinago Koch, 1816 (junior homonym of Gallinago 
Brisson, 1760) ; 


(c) Capella Keyserling & Blasius, 1840, Wirbelth. Europas 
1:9 (junior homonym of Capella Frenzel, 1801). 


102 OPINIONS AND DECLARATIONS 


Il—THE SUBSEQUENT HISTORY OF THE PRESENT 
CASE 


2. Registration of the present application: When in 1951 
Mr. Hemming deposited with the Office of the Commission 
certain preliminary notes regarding the generic name Gallinago 
Koch, the question of the rectification of the entry on the Official 
List of Generic Names in Zoology in regard to the foregoing name 
was allotted the Registered Number Z.N.(S.) 575. 


3. Publication of the present application : The present applica- 
tion was sent to the printer on 24th August 1952 and was 
published in Triple-Part 1/3 of volume 9 of the Bulletin of 
Zoological Nomenclature on 15th October of the same year 
(Hemming, 1952, Bull. zool. Nomencl. 9 : 93—95). 


4. Issue of Public Notices: Under the’ revised procedure 
prescribed by the Thirteenth International Congress of Zoology, 
Paris, 1948 (1950, Bull. zool. Nomencl. 4 : 51—56) Public Notice 
of the possible use by the International Commission on Zoological 
Nomenclature of its Plenary Powers in the present case was given 
on 15th October 1952 (a) in Triple-Part 1/3 of volume 9 of the 
Bulletin of Zoological Nomenclature (the Part in which the 
application by the Standing Committee on Ornithological 
Nomenclature was published), (b) to the other prescribed serial 
publications and (c) to certain general zoological serials. In 
addition, such Notice was given also to fourteen serial publications 
or institutions specially concerned with ornithology. The names 
of the serials and institutions in question have been given in the 
Opinion (Opinion 401)? relating to the generic name Colymbus 
Linnaeus, the Opinion in which the Commission dealt with the 
first of the series of applications submitted by the Standing 
Committee on Ornithological Nomenclature. 


5. Comments of a general character : The issue of the Public 
Notices specified above elicited thirty-seven comments of a 
general character relating to the cases of ornithological nomen- 


2 Opinion 401 has been published as Part 1 of volume 13 of the Opinions and 
Declarations Series. 


DIRECTION 39 103 


clature published in Triple-Part 1/3 of volume 9 of the Bulletin 
of Zoological Nomenclature. These comments came from 
ornithological institutions, groups of ornithologists and individual 
specialists. Of these comments, the authors of thirty-six gave 
general support to the applications published in the foregoing 
Triple-Part. The single remaining comment was from a specialist 
who was opposed to any use of the Commission’s Plenary Powers. 
The communications so received have been reproduced in the 
Appendix annexed to Opinion 401 (Colymbus),*® those supporting 
the applications referred to above being given in Part 1, the single 
comment in opposition to the above applications being given in 
Part 2. 


6. Comments directly related to the present application: Six 
communications directly concerned with the present application 
were received in the Office of the Commission. Of these com- 
munications one was signed by six specialists. Seven of the 
specialists who commented on this case supported the action 
recommended, while four were opposed to that action. The 
communications so received are reproduced in the following 
paragraphs. 


7. Support received from Colonel Richard Meinertzhagen 
(London) : On 5th December 1952 Colonel Richard Meinertz- 
hagen (London) addressed a letter to the Commission commenting 
upon a number of applications relating to the names of birds 
then recently published in Triple-Part 1/3 of volume 9 of the 
Bulletin of Zoological Nomenclature. The portion of Colonel 
Meinertzhagen’s letter relating to the present case was as follows : 
“I wish to register my whole-hearted agreement with Recom- 
mendations Nos... . 174 [Gallinago]”’. 


8. Support received from Dr. Josselyn Van Tyne and five other 
United States ornithologists : On 18th March 1953 Dr. Josselyn 
Van Tyne (Ann Arbor, Michigan, U.S.A.) and the five under- 


3 See Footnote 2 above. 

4 The number here cited is that which was allotted to the present case in the 
list of papers published in Triple-Part 1/3 of volume 9 of the Bulletin of Zoo- 
logical Nomenclature which was printed on the back wrapper of that Part (cover). 


104 OPINIONS AND DECLARATIONS 


mentioned specialists addressed a letter to the Commission in 
support of the present application:—(a) Robert W. Storer 
(Museum of Zoology, University of Michigan, Ann Arbor, U.S.A.) ; 
(b) Andrew J. Berger (Ann Arbor, Michigan, U.S.A.) ; (c) Olin 
Sewall Pettingill, Jr. (Northfield, Minnesota, U.S.A.) ; (d) Frank A. 
Pitelka (Berkeley, California, U.S.A.); (e) John Davis (Los 
Angeles, California, U.S.A.). The letter so received was as 
follows :— 


We, the under-signed, wish to express our strong approval of 
proposal No. 12° (concerning Capella Frenzel) published on page 3 
of vol. 9 of the Bulletin of Zoological Nomenclature. We hope that 
your Commission will take favorable action on it. 


9. Objection received from Dr. Ernst Mayr (then of the American 
Museum of Natural History, New York) : On 8th January 1953 
Dr. Ernst Mayr (then of the American Museum of Natural History, 
New York, and now of the Museum of Comparative Zoology, 
Cambridge, Massachusetts, U.S.A.) addressed a letter to the 
Office of the Commission commenting upon certain of the 
applications relating to the names of birds published in Triple- 
Part 1/3 of volume 9 of the Bulletin of Zoological Nomenclature 
and as regards the present application raising objection to the 
action proposed. The following is the portion of Dr. Mayr’s 
letter relating to the present case :— 


The present point concerns the generic name Gallinago (Z.N.(S.) 575). 
It seems to me that in the statement contained on page 93 [of volume 9 
of the Bulletin of Zoological Nomenclature] there are several 
inaccuracies. To begin with, as pointed out by you, the name Gallinago, 
with Brisson as author, has priority over Capella, and Peters’ reasons 
for supporting Capella are therefore invalid. Equally erroneous 
is the assertion that Capella is now the name in universal use. It is 
true that Dr. Peters had overlooked that the name Gallinago was on the 
Official List when he published volume 2 of his checklist. I am guilty 
of the same negligence when publishing my list of New Guinea birds 
(1941 : 32). However, several correspondents brought this error to 
my attention and in the Birds of the Southwest Pacific and Birds of 


> The number here cited is that which was allotted to the present case in the 
Public Notice of cases involving the possible use of the Commission’s Plenary 
Powers given at the beginning of Triple-Part 1/3 of volume 9 of the Bulletin 
of Zoological Nomenclature. 


DIRECTION 39 105 


the Philippines, 1 have used the name Gallinago, as have other authors 
working in the Australian, Papuan, Malayan and India region. For 
instance it is the name used in Delacour’s Birds of Malaysia and his 
recent publications on Indo-Chinese birds. 


10. Objection received from Dr. John T. Zimmer (The American 
Museum of Natural History, New York) : On 9th January 1953 
Dr. John T. Zimmer (American Museum of Natural History, 
New York) addressed a letter to the Office of the Commission 
commenting upon certain of the applications relating to the 
names of birds which had been published in Triple-Part 1/3 
of volume 9 of the Bulletin of Zoological Nomenclature and in 
the present case raising objection to the action proposed. After 
stressing the importance of maintaining the principle of 
permanency for entries made in the Official List of Generic Names 
in Zoology, Dr. Zimmer proceeded as follows :— 


The principle of permanency is completely upset, however, by the 
proposal to remove Gallinago from the List and replace it by Capella. 
It would be less objectionable to alter the authority for, and reference 
to, Gallinago and credit Brisson with the name, although I am not 
convinced that this is not also an infraction of the authority of the 
List, but to supplant it by Capella destroys all faith in the Official List 
as a permanent standard. 


11. Objection received from M. Noél Mayaud (Faculté des 
Sciences de Paris) : Under cover of a letter dated 17th March 
1953 M. Noél Mayaud (Faculté des Sciences de Paris) furnished 
the following comment amounting to an objection to the action 
proposed in the present case: ‘“‘ Gallinago Brisson, 1760, est un 
nom d’espéce, qui n’invalide pas Capella Frenzel® ”’. 


12. Objection received from Dr. D. L. Serventy (Common- 
wealth Scientific and Industrial Research Organisation, Wildlite 
Survey Section, Perth, Western Australia) : On 22nd April 1953 
Dr. D. L. Serventy (Commonwealth Scientific and Industrial 


6 For a full account of the usage of the generic name Gallinago by Brisson in 
1760 see paragraph 5 of the application reproduced in the first paragraph of the 
present Direction. 


106 OPINIONS AND DECLARATIONS 


Research Organisation, Wildlife Survey Section, Perth, Western 
Australia) addressed the following letter to the Office of the 
Commission objecting to the action proposed in the present 
case :— 


I was interested in your article in The Emu, 53(1), 1953 : 71,7 con- 
cerning certain applications relating to the names of birds submitted 
to the International Commission. 


I would like to offer some comments on the case cited above. It 
would be a very retrograde step in my opinion to remove any name 
from the Official List of Generic Names merely because, after the 
acceptance of such a List, a prior name is subsequently discovered. 
Such a List is surely prepared and authorised to preserve names 
in perpetuity and, to quote the late Joseph Grinnell (Condor, xli, 
1939 : 118) to place them “‘ permanently beyond the reach of name- 
shifters; 7’, 


It must be confessed that in the Gallinago case, the majority of 
ornithologists followed Gregory Mathews in defying the Commission 
in this matter and used Capella. The Royal Australasian Ornitholo- 
gists Union was an exception and Australian publications have almost 
invariably continued to use Gallinago. It is pleasing to find that some 
overseas ornithologists, after having used Capella, are now reverting to 
Gallinago. 1 may quote Ernst Mayr (Birds of the Southwest Pacific, 
1945, p. 44), Jean Delacour and Ernst Mayr (Birds of the Philippines, 
1946, p. 72) and Jean Delacour (Birds of Malaysia, 1947, p. 87). 


If the Commission gives way on this point an unfortunate precedent 
will have been established and one which will undermine one’s faith 
in the security of any other name on the Official List. 


13. Supplementary Report submitted by the Secretary in October 
1955: In May 1954 there was issued to the Members of the 
Commission a Voting Paper (V.P.(54)78) in which particulars 
were given of the comments received in the present case and a 
decision invited on the action recommended in the application 
submitted in 1952. From comments received from certain of the 
Members of the Commission during the Prescribed Voting 


_ 


?. The article here referred to is the Public Notice regarding the possible use 
by the International Commission on Zoological Nomenclature of its Plenary 
Powers in the present case referred to in paragraph 4 of the present Direction 
which was issued to, and published in, The Emu and other ornithological 
serial publications. 


DIRECTION 39 107 


Period on the foregoing Voting Paper it was evident that, having 
regard to the fact that this was not a case where the only action 
possible was a straight choice between two opposing alternatives, 
the Commissioners concerned found it difficult to decide how to 
record their vote. Accordingly, at the close of the Prescribed 
Voting Period the Secretary took the view that the best course 
would be to resubmit this case to the Commission for a fresh 
vote, the Revised Voting Paper so to be issued to be prepared in 
such a way that instead of being asked to vote affirmatively or 
negatively on a single proposal Commissioners would have before 
them alternative proposals of an affirmative character from which 
they could choose whichever seemed to them best calculated to 
serve the general interest. In pursuance of this decision Mr. 
Hemming on 4th August 1954 executed a Minute of Direction 
withdrawing the proposals submitted on the Voting Paper 
referred to above. In the ensuing period, Mr. Hemming carried 
out, as opportunity offered, a survey of the extent to which 
during the years 1916 to 1954 the names Gallinago and Capella 
respectively had been used as the generic name for the Common 
Snipe. On 27th October 1955 Mr. Hemming submitted to the 
Commission the following Supplementary Report in which he 
summarised the history of the present case, gave particulars of 
the information which he had collected regarding the relative 
usage of the foregoing generic names during the last forty years 
and submitted alternative proposals for arriving at a decision 
on the action to be taken in this matter :— 


Request for a Ruling on the question whether the generic name 
‘** Gallinago ’’ or the name ‘‘ Capella ”’ shall be the generic name 
correctly applicable to the Common Snipe (Class Aves) 
(proposed review of an erroneous Ruling given in 
** Opinion ”’ 67) 


By FRANCIS HEMMING, C.M.G., C.B.£. 


(Secretary to the International Commission on Zoological Nomenclature) 


In the present note I re-submit with additional considerations the 
problem whether the name Gallinago or the name Capella should be 
accepted as the generic name for the Common Snipe, a question 
which I submitted to the Commission in 1952 but on which no decision 
has as yet been promulgated. 


108 OPINIONS AND DECLARATIONS 


2. The main features of this case were set out in a paper of mine 
which was published in Triple Part 1/3 of volume 9 of the Bulletin of 
Zoological Nomenclature (Hemming, Bull. zool. Nomencl. 9 : 93—95), 
and it is necessary here to recapitulate only the following points :— 


(a) In 1916 by the Ruling given in Opinion 67 the generic name 
Gallinago Koch, 1816 (type species: Scolopax gallinago 
Linnaeus, 1758) was placed on the Official List of Generic 
Names in Zoology. 


(b) In the process of checking against recent monographs, catalogues, 
etc., the entries made in the Official List in the pre-Lisbon 
(1935) period, I observed that, whereas in 1916 Hartert in the 
main portion of his Végel pal. Faun. had used the generic name 
Gallinago Koch for the Common Snipe, he had replaced that 
name by the name Capella Frenzel, 1801 in the supplementary 
volume published in 1921. 


(c) The nominal genus Capella Frenzel, 1801, has as its type species 
by monotypy the nominal species Capella coelestis Frenzel, 
1801, which species is subjectively identified by ornithologists 
with Scolopax gallinago Linnaeus, 1758, the type species of 
Gallinago Koch, 1816. The latter generic name is therefore 
a junior subjective synonym of Capella Frenzel, 1801. 


(d) On becoming aware of the substitution by Hartert of the name 
Capella Frenzel, 1801, for the name Gallinago Koch, 1816, I 
made further enquiries as to current usage. This showed that 
Peters (in vol. 2 of his Checklist) has used the name Capella 
Frenzel and that he had been widely followed by later authors. 
Since my paper of 1952 I have ascertained, however, that 
several important authors have continued to use the name 
Gallinago Koch, despite the priority of Capella Frenzel. I 
attach as Appendix | to this note particulars of recent and 
comparatively recent usage of the names Capella and Gallinago 
respectively which have been gleaned from a search of standard 
works and from a study of the annual volumes of the Zoological 
Record undertaken by this Office. Though doubtless not 
complete, the list so given is, I believe, representative in the 
picture which it gives. 


(e) In 1951/52 I undertook for quite a different purpose a survey. 
of the names used in Brisson’s Ornithologia of 1760, a work 
which, considering its early date, its intrinsic importance and 
the large numbers of well-known names which it contains, 
has been surprisingly neglected by ornithologists in many 
respects. This survey disclosed amongst many other things 
the existence of a generic name Gallinago Brisson (type species 
by absolute tautonymy : Scolopax gallinago Linnaeus, 1758). 
The name Gallinago Brisson, 1760, is therefore a senior 
subjective synonym of Capella Frenzel, 1801, and a senior 


DIRECTION 39 109 


objective synonym of Gallinago Koch, 1816. Moreover, the 
name Gallinago Koch is not merely (as previously thought) a 
junior subjective synonym of Capella Frenzel but is also an 
objectively invalid name, being a junior homonym of 
Gallinago Brisson. 


3. In my paper of 1952 I suggested, after consultation with Colonel 
Richard Meinertzhagen, at that time Chairman of the Standing 
Committee of Ornithological Nomenclature established by the Tenth 
International Ornithological Congress, Uppsala, 1950, that the 
situation discussed in the preceding paragraph should be resolved 
by the use by the Commission of its Plenary Powers for the purpose 
of suppressing the name Gallinago Brisson, 1760, thereby validating 
the name Capella Frenzel. I submitted this recommendation because 
on the evidence then available it seemed to be the one best calculated to 
promote stability in the nomenclature of the group concerned. 


4. In the period following the publication in October 1952 of Triple- 
Part 1/3 of volume 9 of the Bulletin of Zoological Nomenclature, the 
Part which contained a large number of applications relating to 
ornithological nomenclature, including the application relating to the 
present case, communications intimating general support for the 
action proposed in all these applications were received from thirty-six 
institutions, groups of ornithologists and individual specialists. In 
addition, communications specifically concerned with the present 
problem were received from nine ornithologists, of whom eight were 
Americans and one was an Australian. Of these specialists six favoured 
the suppression of Gallinago Brisson and the consequent validation of 
Capella Frenzel, while three favoured the retention of the name 
Gallinago. 


5. The following are the names of the six American ornithologists 
who supported the proposal to validate the name Capella Frenzel, for 
which they expressed their “‘ strong approval’’ and intimated their 
hope that the Commission would “‘take favorable action on it” :— 
Josselyn Van Tyne (Ann Arbor, Michigan); Robert W. Storer (Ann 
Arbor); Andrew J. Berger (Ann Arbor); Olin Sewall Pettingill, Jr. 
(Northfield, Minnesota); Frank A. Pitelka (Berkeley, California) ; 
John Davis (Los Angeles, California). 


6. The three ornithologists who objected to the substitution of 
Capella for Gallinago on the Official List were: Ernst Mayr (then of 
New York); John T. Zimmer (New York); D. L. Serventy (Perth, 
Western Australia). All three of these specialists made the point 
that the name Gallinago had been placed on the Official List and that 
it would be wrong in principle to remove it therefrom (in favour of 
Capella), for the utility of the Official List would be seriously impaired 
if names were to be removed from it if later they were found to be 


110 OPINIONS AND DECLARATIONS 


technically defective in some nomenclatorial respect. Mayr pointed 
out that the change-over from Gallinago to Capella had not been so 
complete as had been suggested in my 1952 paper, and in this con- 
nection he wrote: “In the Birds of the Southwest Pacific and Birds 
of the Philippines { have used the name Gallinago, as have other authors 
working on the Australian, Papuan, Malayan and India region. For 
instance, it is the name used in Delacour’s Birds of Malaysia and in his 
recent publications on Indo-Chinese birds”’. Serventy wrote: “It 
must be confessed that in the Gallinago case, the majority of orni- 
thologists followed Gregory Mathews in defying the Commission in this 
matter and used Capella. The Royal Australasian Ornithologists’ 
Union was an exception and Australian publications have almost 
invariably continued to use Gallinago”’’. As regards the name Capella 
Mayr pointed out that it “‘ was published in an ornithologically 
worthless publication which has been wisely ignored for 150 years 
and of which, to my knowledge, only a single copy is extant—that in 
the Library of Cambridge University ’’. 


7. It is very unfortunate that, when the name Capella Frenzel, 1801, 
was unearthed as a senior subjective synonym of the name Gallinago 
Koch, 1816, the matter was not at once brought to the attention of the 
Commission, for there would have been the strongest grounds for the 
use by the Commission of its Plenary Powers for the purpose of 
promoting stability in the nomenclature of the group of birds con- 
cerned, having regard to the fact (1) that the generic name Gallinago 
Koch was extremely well-known and had been in use for well over 
one hundred years, (2) that this generic name had already been placed 
on the Official List of Generic Names in Zoology, (3) that the name 
Capella Frenzel, 1801, had been totally ignored for upwards of one 
hundred and twenty years, and (4) that it was published in a work of 
(as Mayr explains) no ornithological value of which only one copy 
survives and which therefore has not been seen and will never be seen 
by any but a very few ornithologists. The failure by ornithologists 
to bring this matter before the Commission at the time when the name 
Capella Frenzel first came to light has, however, created a difficult 
situation, for a return to the name Gallinago would mean a reversal 
of much modern practice. On the other hand, the use of Capella in 
place of Gallinago in recent decades has not been so unanimous as I 
had supposed in 1952, and from this point of view the substitution of 
Capella for Gallinago would be a hardship for those workers who have 
throughout continued to use the name Gallinago in the belief that this 
was the correct course, having regard to the fact that the name 
Gallinago had been placed on the Official List before the rival claims 
of Capella were advanced. If such a case were to arise today, the 
rejection of Gallinago Koch in favour of the older name Capella 
Frenzel by individual workers without resort to the Commission, 
would in fact be contrary to the Régles, for in 1948 (1950, Bull. zool. 
Nomencl. 4 : 268, Point (c)) the Thirteenth International Congress 
of Zoology adopted a provision protecting generic names which had 


DIRECTION 39 Let 


been placed on the Official List from attacks of this kind until the 
matter had been brought before, and decided by, the International 
Commission. 


8. It is essential that the Commission should now take a decision in 
this matter, for the entry on the Official List of the name Gallinago 
Koch must be regularised in one way or another before the Official 
List can be published in book-form. For the reasons explained above 
it seems certain that some inconvenience will be inevitable, for 
whatever the decision now taken by the Commission some ornithologists 
will need to change the generic name which they are currently using 
for the Common Snipe. When in 1954 the stage was reached for 
asking the Commission to vote on the present case, I prepared for its 
consideration a Voting Paper (V.P.(54)78) in which the Members of the 
Commission were invited to vote on the proposal submitted in 1952, 
namely that Gallinago Brisson, 1760, should be suppressed under the 
Plenary Powers and that its junior subjective synonym Capella Frenzel, 
1801, thus validated, should be placed on the Official List in place of 
the objectively invalid name Gallinago Koch. From communications 
received from certain of the Members of the Commission it was evident 
that, having regard to the fact that this was not a case where the only 
action possible was a straight choice between two opposing alternatives, 
some Commissioners had found it difficult to decide how to record 
their vote. Accordingly, at the close of the Prescribed Voting Period 
I took the view, as Secretary, that the best course would be to re-submit 
this case to the Commission for a fresh vote, the Revised Voting 
Paper so to be issued to be prepared in such a way that instead of being 
asked to vote affirmatively or negatively on a single proposal 
Commissioners would have before them alternative proposals of an 
affirmative character from which they could choose whichever seemed 
to them best calculated to serve the general interest. 


Alternative Proposals on which it is proposed 
that the Commission now be asked to vote 


9. There are two issues of substance on which it is now proposed 
that Members of the Commission should be asked to vote. The first 
issue Of substance is whether the generic name properly applicable 
to the Common Snipe shall be Gallinago or Capella. On this question 
two alternatives are now submitted for decision, namely Alternative 
*“ A” (acceptance of Gallinago) and Alternative “‘ B’’ (acceptance of 
Capella). The second issue of substance is whether, if Gallinago is to 
be accepted, that name shall rank from Brisson, 1760 (as it does at 
present) or whether it shall be credited to Koch, 1816, the author and 
date to which it has always been attributed in the literature. This 
question is submitted because it is clear that some specialists (e.g. 
Zimmer) are of the opinion not only that the name Gallinago should be 
accepted but also that it should be attributed to the author and date 
by which it has always been known and not to Brisson, 1760, whose 


riZ OPINIONS AND DECLARATIONS 


prior responsibility for this name has only come to light in the course 
of the last year or so. On this question also two alternatives are now 
submitted, namely Alternative “‘ X”’ (acceptance of Koch, 1816, as 
the author and date of the name Gallinago) and Alternative *“‘ Y ”’ 
(acceptance of Brisson, 1760, as the author and date of the name 
Gallinago). Itis particularly hoped that all members of the Commission 
will vote on the foregoing question and not merely those who favour 
the acceptance of the name Gallinago, for, if the proposal in favour 
of that name (A/ternative “‘A ’’ above) is adopted by a majority of the 
Commission, it is desired to have on record the views of all 
Commissioners on the question of the authorship and date to be 
attributed to it. 


10. While the simplified titles given above for the various 
alternatives now submitted will, it is hoped, be of convenience to the 
Members of the Commission in deciding how to cast their votes in the 
present case, I feel that it is desirable also to set out in full the Ruling 
which would be given by the Commission according to whichever of the 
alternative Rulings it decides to adopt. These particulars are given in 
Appendix 2 to the present paper and it is only necessary to note here 
that the broad effect of the adoption of each of the alternative decisions 
now submitted for consideration would be as follows :— 


(1) An affirmative vote on Alternative “‘A’’ (acceptance of the name 
Gallinago as against Capella Frenzel), on Part 1 of the annexed 
Voting Paper, if coupled with an affirmative vote on A/ternative 
**X ” (acceptance of Gallinago as from Koch, 1816) in Part 2 
of the annexed Voting Paper, would lead to the adoption of 
the Ruling given as Draft Ruling No. 1 in Appendix 2 to the 
present paper. 


(2) An affirmative vote on Alternative “‘A’’ (acceptance of the name 
Gallinago as against Capella Frenzel, 1801) on Part 1 of the 
annexed Voting Paper, if coupled with an affirmative vote on 
Alternative “‘ Y’’ (acceptance of Gallinago as from Brisson, 
1760) in Part 2 of the annexed Voting Paper would lead to the 
adoption of the Ruling given as Draft Ruling No. 2 in Appendix 
2 to the present paper. 


(3) An affirmative vote on Alternative ‘“‘ B”’ (acceptance of Capella 
Frenzel, 1801, as against the name Gallinago) on Part 1 of the 
annexed Voting Paper would lead to the adoption of the 
Ruling given as Draft Ruling No. 3 in Appendix 2 to the 
present paper. 


A Proposal on Procedure which Members of the 
Commission are invited to approve 


11. The method of voting by way of a series of alternative affirmative 
proposals, which has been adopted in the present case, is well adapted 


DIRECTION 39 113 


for securing decisions from the Commission when the issue involved is 
one where a direct vote on a single proposition would not necessarily 
exhaust the possible courses of action which might be taken. This 
procedure, however, does not fit in very readily with the procedure 
prescribed for dealing with cases where the proposal submitted involves 
the use of the Commission’s Plenary Powers. The required Public 
Notices have been duly given in the present case and no difficulty 
arises under that head. However, in order to secure a decision under 
the Plenary Powers procedure, it is now necessary not only that there 
should be a majority of votes in the Commission in favour of the 
action proposed, but also that that majority shall consist of not less 
than two out of every three votes cast (1950, Bull. zool. Nomencl. 
4:51, Point (b)). This creates a procedural problem in the present 
case which must be faced, for under the foregoing rule a vote by the 
majority of the Commission could be negatived by a vote of a minority 
of only just over one-third of the total number of votes cast. Moreover, 
the fact that some of the alternatives do not involve the use of the 
Plenary Powers (and therefore can be approved by a simple majority), 
while others do involve the use of those Powers (and therefore require 
for their adoption a two-thirds majority) might result in the adoption, 
on the separate Parts of the Voting Paper now submitted, of decisions 
which were irreconcilable with one another. Leaving aside the question 
whether the two-thirds rule for Plenary Powers cases represents the 
most suitable procedure for new applications, it will, I think, be 
agreed that it is not necessarily appropriate in a case such as the 
present which is concerned with the regularisation in one manner or 
another of an erroneous decision already taken. Although it is desirable 
that cases of all types should be handled as expeditiously as possible, 
there is an element of urgency attaching to the correction of former 
errors which does not normally arise in the case of new applications. 
For it is particularly important that errors of the past should be 
corrected as quickly as possible, so that specialists in the groups 
concerned may be relieved without delay of the uncertainties created 
by those errors. In cases such as the present which are concerned 
with correcting former errors in entries made in the Official List there 
is at the present time a further element of special urgency, for all such 
errors must be regularised by the Commission in one way or another 
before arrangements can be made for the publication in book-form 
of the instalments so far placed on the Official Lists. The preparations 
for the publication of this volume are now well advanced and it is of 
the highest importance that the completion of this task should not 
be impeded by any failure to provide decisions on outstanding 
questions. Accordingly I recommend that, in order to ensure the 
adoption of a definitive decision in the present case, a twofold 
procedure should be adopted, namely (i) that the Commission should 
vote first on the nomenclatorial issues involved in the present case, 
and (ii) that, having done so, it should vote on a procedure designed 
to ensure that the decisions so taken shall be effectively binding as 
decisions of the Commission as a body, irrespective of whether, if in 


114 OPINIONS AND DECLARATIONS 


the case of any of the proposals which involve the use of the Plenary 
Powers there is a majority in favour of the use of those Powers, the 
majority of votes cast is a two-thirds majority. The Voting Paper 
now submitted consists therefore of three separate Parts. As already 
explained (paragraph 10 above), Parts 1 and 2 each deal with alternative 
decisions on the nomenclatorial issues involved in the present case 
as between which the Commission is invited to make a choice. Part 3 
is concerned with a proposal designed to secure that decisions taken 
on Parts 1 and 2 shall not be upset by the operation of the technical 
conditions attaching to the use of the Plenary Powers procedure. In 
this Part a proposal is submitted that, if a majority of the Members of 
the Commission vote in favour of a course involving the use of the 
Plenary Powers either on Part 1 or on Part 2 of the annexed Voting 
Paper but the majority of votes so cast does not amount to a majority 
of two out of every three votes cast, this fact shall nevertheless not 
render ineffective the majority vote taken on the earlier Part or Parts 
of the Voting Paper. The procedure recommended above is in full 
conformity with the provisions prescribed by the International Congress 
of Zoology regarding the procedure to be observed in voting upon 
applications involving the use of the Plenary Powers, for the two- 
thirds majority rule prescribed by the Congress will apply to the vote 
on Part 3 of the Voting Paper. All that in effect the proposal submitted 
in Part 3 does is to transfer the application of the two-thirds majority 
rule from the votes on Parts 1 and 2 to the vote on Part 3, thus leaving 
the questions specified in Parts 1 and 2 to be dealt with by a simple 
majority if the proposal submitted in Part 3 is approved = at least 
two votes out of every three cast. 


APPENDIX 1 


An indication of the extent to which during the years 1916 to 1954 
authors used the generic names ‘‘ Capella ’’ and ‘‘ Gallinago ”’ 
respectively as the generic name for the Common Snipe 


Neither the supporters of the generic name Capella nor those of 
the generic name Gallinago as the name for the Common Snipe have 
furnished detailed analyses of the relative usage of these names in 
recent times such as were furnished in the case of the Decapod names 
Crangon Fabricius, 1798, Crangon Weber, 1795, and Crago Lamarck, 
1801, by Dr. L. B. Holthuis (Leiden) and Dr. Fenner A. Chace, Jr. 
(Washington, D.C.) (Opinion 334). It appeared to me, however, that 
it would be useful in the present case to lay before the Commission 
some particulars as to current usage, even though the list so compiled 
could not claim to do more than give a general indication of the 
practice of ornithologists in this matter. The following particulars 
have therefore been compiled in the Office of the Commission, mainly 
by consulting well-known recent works but partly also by reference 
to the annual volumes of the Zoological Record and from information 
supplied by ornithologists who have offered comments as to the action 


DIRECTION 39 115 


which it is desirable should be taken by the International Commission. 
The list which has been compiled has not been prepared in the form 
of a detailed bibliography, for it has been considered that for the 
present purposes it would be sufficient to record the name of each 
author, and the date of the work concerned and to give some indication 
of the nature of that work. 


2. Subject to the foregoing reservations the following particulars are 
submitted of usage during the period since the publication in 1916 of 
Opinion 67 (in which the name Gallinago Koch, 1816, was placed on 
the Official List of Generic Names in Zoology) up to the present time :— 


(a) Works in which the generic name *‘ Capella” Frenzel, 1801, 
has been used in place of “* Gallinago’’ Koch, 1816 
American Ornithologists’ Union Checklist, 1931 
Peters, Checklist of Birds of the World, 1934 
Handbook of British Birds, 1940, vol. 4 
Gregory Mathews, 1920, Australian Avian Record 4 : 131 
Jackson, 1926 (Kenya and Uganda) 
Belcher, 1930 (Nyasaland) 
Bates, 1930 (West Africa) 
Friedemann, 1930 (Ethiopia and Kenya) 
Wait, 1931 (Ceylon) 
Wetmore & Swales, 1931 (Haiti) 
Priest, 1934 (Rhodesia) 
Taverner, 1934 (Canada) 
Roberts, 1940 (South Africa) 
Woods, 1943 (Michigan birds) 
Moltoni, 1944 (Italian East Africa) 
Bond, 1945 (West Indies) 
Bailey, 1948 (Alaska) 
Bannerman, 1951 (Tropical West Africa) 
Glenister, 1951 (Malaya, Singapore and Penang) 
Bannerman, 1953 (West and Tropical Africa) 
Cave and McDonald, 1955 (Sudan) 
Phillips, 1953 (Ceylon) 7 
Salim Ali, 1953 (Travancore and Cochin) 
Macleod, 1954 (Key to names of British Birds) 
Meinertzhagen, 1954 (Arabia) 


116 OPINIONS AND DECLARATIONS 


(b) Works in which the generic name “‘ Gallinago’’ Koch 
1816, has been retained 


Royal Australasian Ornithologists’ Union 
Almost all Australian publications (letter from D. L. Serventy) 


Authors working in the Australian, Papuan, Malayan and India 
Regions (letter from E. Mayr) 


Priest, 1929 (Southern Rhodesia) 
Mayr, 1945 (Southwest Pacific) 
Delacour and Mayr, 1946 (Philippines) 


Delacour, 1947 (Malaysia; also publications by this author on 
Indo-Chinese birds) 


3. It must be emphasised again that the foregoing particulars could 
no doubt be greatly extended if the literature of the last thirty years 
were to be carefully examined by ornithologists for this purpose. It is, 
however, clear that, as stated in litt. by Serventy, one of the advocates 
of the name Gallinago, the weight of usage since the time of Gregory 
Mathews has been on the side of Capella rather than on that of 
Gallinago. Against this due regard must, however, be given to the 
fact that prior to the re-awakening of the name Capella some thirty 
years ago, the name Gallinago Koch possessed an undisputed 
supremacy in the literature, having been used by leading authors 
for over one hundred years. 


4. In judging the particulars of usage given in paragraph 2 above, 
it must be noted also that those authors who changed over from 
Gallinago Koch, 1816, to Capella Frenzel, 1801, did so in the belief 
that Capella was the oldest available generic name subjectively 
applicable to the Common Snipe and therefore that this was the 
correct name to be used for the genus containing this species. The 
authors concerned were not aware that the oldest available name 
objectively applicable to the Common Snipe was Gallinago Brisson, 
1760, a name having more than forty years’ priority over Capella 
Frenzel. Most of the authors cited in section (a) of paragraph 2 above 
are well known for their orthodoxy in matters of nomenclature and 
it cannot be doubted that, if they had been aware of the existence of the 
name Gallinago Brisson, 1760, they would not have taken up the 
name Capella Frenzel but would have continued to use the name 
Gallinago, attributing it, however, to Brisson, 1760, instead of to 
Koch, 1816. But it was not only those authors who substituted the 
name Capella for that of Gallinago who thought that in so doing 
they were following the strict letter of the law, for some of those who 
continued to use the name Gallinago Koch believed that thereby they 
also were acting in strict conformity with the Régles, having regard 
to the fact that the name Gallinago Koch had been placed on the 


DIRECTION 39 117 


Official List of Generic Names in Zoology. For the period since the 
Paris Congress of 1948 this belief was well-founded, for that Congress 
laid it down that a name which has been placed on the Official List, 
even if later found to be technically defective, is not to be discarded in 
favour of some other name without the prior consent of the Commission. 


APPENDIX 2 


Drafts of Alternative Rulings submitted in relation to the generic names 
** Gallinago ”’ and ‘‘ Capella ”’ 


Draft Ruling No. 1 


(validation of the generic name “ Gallinago’’ as from Koch, 1816 
(author and date customarily attributed to this name) and 
confirmation of the entry of this name on the “ Official 
List’ made in 1916) 


(1) Under the Plenary Powers (a) Gallinago Brisson, 1760, to be 
suppressed for the purposes both of the Law of Priority and of the 
Law of Homonymy and (b) Capella Frenzel, 1801, to be suppressed 
for the purposes of the Law of Priority but not for those of the Law of 
Homonymy. 


(2) The entry on the Official List of Generic Names in Zoology of the 
name Gallinago Koch, 1816 (gender: feminine) (type species, by 
absolute tautonymy: Scolopax gallinago Linnaeus, 1758) made by 
the Ruling given in Opinion 67 to be confirmed. 


(3) The specific name gallinago Linnaeus, 1758, as published in the 
combination Scolopax gallinago (specific name of type species of 
Gallinago Koch, 1816) to be placed on the Official List of Specific 
Names in Zoology. 


(4) The generic names specified in (1) above as there suppressed 
under the Plenary Powers, together with Capella Keyserling & Blasius, 
1840 (a junior homonym of Capella Frenzel, 1801), to be placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology. 


(5) The specific name media Koch, 1816, as published in the 
combination Gallinago media (a junior objective synonym of gallinago 
Linnaeus, 1758, as published in the combination Scolopax gallinago, to 
be placed on the Official Index of Rejected and Invalid Specific Names 
in Zoology. 


Draft Ruling No. 2 


(acceptance of the name “* Gallinago ’’ Brisson, 1760, and substitution 
on the “* Official List”? of that name in place of the name 
** Gallinago’’ Koch, 1816) 


(1) The following entry to be inserted in the Official List of Generic 
Names in Zoology in place of the entry thereon of the name Gallinago 
Koch, 1816, made by the Ruling given in Opinion 67: Gallinago 


118 OPINIONS AND DECLARATIONS 


Brisson, 1760 (gender : feminine) (type species, by absolute tautonomy : 
Scolopax gallinago Linnaeus, 1758). 


(2) The specific name gallinago Linnaeus, 1758, as published in the 
combination Scolopax gallinago (specific name of type species of 
Gallinago Brisson, 1760) to be placed on the Official List of Specific 
Names in Zoology. 


(3) The names Gallinago Koch, 1816 (a junior homonym, and a 
junior objective synonym, of Gallinago Brisson, 1760) and Capella 
Keyserling Blasius, 1840 (a junior homonym of Capella Frenzel, 1801) 
to be placed on the Official Index of Rejected and Invalid Generic Names 
in Zoology. 


(4) The name media Koch, 1816, as published in the combination 
Gallinago media (a junior objective synonym of gallinago Linnaeus, 
1758, as published in the combination Scolopax gallinago) to be 
placed on the Official Index of Rejected and Invalid Specific Names in 
Zoology. 


Draft Ruling No. 3 


(validation of the generic name ‘‘ Capella’? Frenzel, 1801, and 
substitution of that name for “ Gallinago”’ Koch, 1816, 
on the ‘ Official List of Generic Names in Zoology ’’) 


(1) Under the Plenary Powers, the generic name Gallinago Brisson, 
1760, to be suppressed for the purposes of the Law of Priority but not 
for those of the Law of Homonymy. 


(2) The entry on the Official List of Generic Names in Zoology of the 
name Gallinago Koch, 1816, to be replaced by the following entry : 
Capella Frenzel, 1801 (gender : feminine) (type species by monotypy : 
Capella coelestis Frenzel, 1801 (see Note below)). 


(3) The specific name gallinago Linnaeus, 1758, as published in the 
combination Scolopax gallinago to be placed on the Official List of 
Specific Names in Zoology. 


(4) The generic name (a) Gallinago Brisson, 1760, as suppressed 
under the Plenary Powers under (1) above, and (b) Gallinago Koch, 
1816 (a junior homonym of Gallinago Brisson, 1760), and (c) Capella 
Keyserling & Blasius, 1840 (a junior homonym of Capella Frenzel 
1801) to be placed on the Official Index of Rejected and Invalid Generic 
Names in Zoology. 


(5) The name media Koch, 1816, as published in the combination 
Gallinago media (a junior objective synonym of gallinago Linnaeus, 


DIRECTION 39 119 


1758, as published in the combination Scolopax gallinago), to be placed 
on the Official Index of Rejected and Invalid Specific Names in Zoology. 


NOTE (to all three Draft Rulings): I have not yet been able to obtain 
a photostat copy of the portion of Frenzel’s book containing his 
generic name Capella, but I hope to do so at an early date®. It 
seems likely that, just as Koch later gave the substitute name 
Gallinago media to Scolopax gallinago Linnaeus, so also earlier 
Frenzel gave the substitute name Capella coelestis to that species. 
If this proves to be the case, the name coelestis Frenzel, 1801 (like 
media Koch, 1816) is a junior objective synonym of gallinago 
Linnaeus, 1758, as published in the combination Scolopax gallinago, 
and the invalid name coelestis Frenzel, 1801, should now be placed 
on the Official Index of Rejected and Invalid Specific Names in 
Zoology, as well as media Koch, 1816. 


IJI.—THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


14. Issue of Voting Paper V.P.(O.M.)(55)36 : On 27th October 
1955 a Revised Voting Paper numbered Voting Paper V.P.(O.M.) 
(55)36 was issued to the Members of the Commission in regard 
to the present case. This Voting Paper was divided into three 
Parts on which the Members of the Commission were invited 
to vote separately. Parts 1 and 2 were each concerned directly 
with the subject matter of the present case. On each of these 
Parts the Members of the Commission were asked to vote affirm- 
atively on one or other of alternative affirmative courses. Part 3 
was concerned with a question of procedure on which the Members 
of the Commission were invited to vote either affirmatively or 
negatively on the proposition submitted. Details of the proposals 
so submitted in the several Parts of the foregoing Voting Paper 
are given in paragraphs 16, 17 and 18 below, in which paragraphs 
are given also particulars of the votes cast thereon by the Members 


8 For later developments see paragraph 21 of, and the Appendix to, the present 
Direction. 


120 OPINIONS AND DECLARATIONS 


of the Commission. The following explanatory note referring 
to all three Parts appeared at the end of the Voting Paper :— 


IMPORTANT NOTE :—tThis Voting Paper is divided into Three 
Parts, on each of which each Member of the Commission is invited 
to record his vote. For an explanation of the reason for the adoption 
of this procedure see paragraphs 9, 10 and 11 of the memorandum 
bearing the same Registered Number as the present Voting Paper 
submitted by the Secretary simultaneously therewith.°® 


15. The Prescribed Voting Period: As the foregoing Voting 
Paper was issued under the One-Month Rule, the Prescribed 
Voting Period closed on 27th November 1955. 


16. Particulars of the question voted upon in Part 1 of Voting 
Paper V.P.(O.M)(55)36 and of the votes cast thereon: In Part I 
of Voting Paper V.P.(O.M.(55)36 the Members of the Commission 
were invited to vote either “‘ for Alternative ‘A’ (acceptance of 
name Gallinago) or for Alternative ‘ B’ (acceptance of the name 
Capella)’. At the close of the Prescribed Voting Period the state 
of the voting on Part 1 of Voting Paper V.P.(O.M.)(55)36 was as 
follows :— 


(a) In favour of Alternative “‘ A” (acceptance of “‘ Gallinago ”’), 
twenty-two (22) votes : 


Riley ; Holthuis; Bonnet; Hering; Mertens; Stoll; 
Vokes; Boschma; Bodenheimer; Mayr; Esaki; 
Dymond ; Key; Bradley (J.C.) ; do Amaral ; Hank ; 
Cabrera ; Kuhnelt ; Hemming; Tortonese ; Sylvester- 
Bradley ; Lemche ; 


(b) In favour of Alternative “‘ B”’ (acceptance of ‘‘ Capella’’), 
three (3) votes : 


Jaczewski ; Prantl ; Miller ; 
(c) Voting Papers not returned : 


None. 


® The memorandum here referred to has been reproduced in paragraph 13 of 
the present Direction. 


DIRECTION 39 121 


17. Particulars of the question voted upon in Part 2 of Voting 
Paper V.P.(O.M.)(55)36 and of the votes cast thereon : In Part 2 
of Voting Paper V.P.(O.M.)(55)36 the Members of the Commission 
were invited to vote either ‘‘ for Alternative ‘ X’ (acceptance of 
Gallinago as from Koch, 1816) or for Alternative ‘ Y ’ (acceptance 
of Gallinago as from Brisson, 1760)”. At the close of the 
Prescribed Voting Period the state of the Voting on Part 2 of 
Voting Paper V.P.(O.M.)(55)36 was as follows :— 


(a) In favour of Alternative “‘ X”’ (acceptance of ** Gallinago ”’ as 
from Koch, 1816), four (4) votes : 


Hering ; Esaki; Key; Miller; 


(b) In favour of Alternative ““ Y”’ (acceptance of ** Gallinago ” as 
from Brisson, 1760), twenty-one (21) votes : 


Riley ; Holthuis; Bonnet; Mertens; Stoll; Vokes ; 
Boschma ; Bodenheimer; Mayr; Dymond; Bradley 
(J.C.); do Amaral; Hank6d; Cabrera; Jaczewski ; 
Kiihnelt ; Hemming; Tortonese; Prantl; Sylvester- 
Bradley ; Lemche ; 


(c) Voting Papers not returned : 


None. 


18. Particulars of the question voted upon in Part 3 of Voting 
Paper V.P.(O.M.)(55)36 and of the votes cast thereon : In Part 3 
of Voting Paper V.P.(O.M.)(55)36 the Members of the Commission 
were invited to vote either for, or against, “‘ the proposal set out 
in paragraph 11 of the paper submitted by the Secretary simul- 
taneously with the present Voting Paper [i.e. in the paragraph 
numbered as above in the paper reproduced in paragraph 13 of 
the present Direction], that for the reasons there explained, if 
in the vote either on Part 1 or on Part 2 of the present Voting 
Paper a majority of votes is cast in favour of the use of the Plenary 
Powers but that majority is not a two-thirds majority, this fact 


122 OPINIONS AND DECLARATIONS 


shall nevertheless not render ineffective the majority vote or 
votes so taken on the earlier Parts of the present Voting Paper ”’. 
At the close of the Prescribed Voting Period the state of the voting 
on Part 3 of Voting Paper V.P.(O.M.)(55)36 was as follows :— 


(a) Affirmative Votes had been given by the following twenty-one 
(21) Commissioners (arranged in the order in which Votes 
were received) : 


Riley; Bonnet; Hering; Mertens; Stoll; Vokes; 
Bodenheimer ; Mayr; Esaki; Dymond; do Amaral ; 
Hanko ; Cabrera; Jaczewski; Kuhnelt ; Hemming ; 
Tortonese; Prantl; §Sylvester-Bradley;  Miéiller; 
Lemche ; 


(b) Negative Votes, four (4) : 


Holthuis ; Boschma ; Key ; Bradley (J.C.) ; 


(c) Voting Papers not returned : 


None. 


19. Deferment by the Secretary in November 1955 of the 
Declaration of the Result of the Vote on Voting Paper V.P.(O.M.) 
(55)36 : On 28th November 1955, the day following the close of 
the Prescribed Voting Period, Mr. Hemming, as Secretary, placed 
a Minute on the Commission’s File Z.N.(S.) 575, directing that 
the Declaration of the Result of the Vote on Voting Paper V.P. 
(O.M.)(55)36 be deferred until such time as a photostat copy of the 
relevant portion of Frenzel’s Beschreibung of 1801 had been 
obtained and until in consequence it was possible to determine 
whether the name for the Snipe published in that work was an 
available name or whether it was invalid and in consequence 
required under the terms of the vote taken by the Commission 
on the foregoing Voting Paper to be placed on the appropriate 
Official Index of Rejected and Invalid Names. 


DIRECTION 39 123 


20. Addition of the generic name ‘‘ Capella ’’ Frenzel, 1801, to 
the ‘* Official Index of Rejected and Invalid Generic Names in 
Zoology ’? : On 8th February 1956 Mr. Hemming, as Secretary, 
placed on the Commission’s File Z.N.(S.) 575 the following 
Minute recording the receipt of a photostat copy of the relevant 
portion of Frenzel’s Beschreibung, directing that in accordance 
with the vote taken on Voting Paper V.P.(O.M.)(55)36 the 
objectively invalid name Capella Frenzel, 1801, be placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology, 
and confirming the decision that the generic name Capella 
Keyserling & Blasius, 1840, be entered on the foregoing Official 
Index, subject to an amendment of the explanatory note to be 
attached to the said entry :— 


Addition of the name ‘‘ Capella’’ Frenzel, 1801, to the ‘° Official 
Index of Rejected and Invalid Generic Names in Zoology ”’ 


By FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


At the time of the submission of the proposals in regard to the 
Capella/Gallinago problem, laid before the International Commission 
with Voting Paper V.P.(O.M.)(55)36, I reported that I was attempting 
to obtain a photostat copy of the portion of Frenzel’s Beschreibung 
der Végel und ihrer Eier in der Gegend von Wittenberg of 1801 in which 
the generic name Capella was published in order to make sure that 
Capella coelestis was in fact an available name and recommended that, 
if this proved not to be the case, an addition should be made to the 
appropriate Official Index of Rejected and Invalid Names. This 
proposal, together with the other proposals then submitted, was 
=i by the Commission in its vote on the foregoing Voting 

aper. 


_ 2, At the time of the submission of the above proposal I had been 
informed by Dr. Ernst Mayr that, so far as he knew, there was only 
one copy extant of Frenzel’s book and that this copy was at Cambridge. 
After some enquiry I ascertained that the copy in question was in the 
Balfour Library in the Department of Zoology at Cambridge 
University. Through the kindness of the Librarian of that Institution 
in depositing the copy of Frenzel’s book at the University Library in 
order that it might be photographed, I have now obtained photostats 
of the relevant pages (pp. 58—S9). The grateful thanks of the 
Commission are due to the Librarians of both these Institutions for the 
prompt and willing help rendered by them in this matter. 


124 OPINIONS AND DECLARATIONS 


3. The information required is found to be on page 58 of Frenzel’s 
book where his account of the Snipe starts, page 59 containing only a 
continuation of the description of that bird. Now that at last a copy 
of the relevant portion of Frenzel’s Beschreibung is available for study 
the situation is found to be quite different from that hitherto supposed. 
The chief points to be noted are :— 


(a) Contrary to the statements in modern works Frenzel did not 
introduce for the Snipe a new generic name consisting of the 
word Capella and he did not call that bird by the name Capella 
coelestis. 


(b) Frenzel placed the Snipe in the same genus—Scolopax—as that 
in which it had been placed by Linnaeus in 1758 when it was 
given the name Scolopax gallinago. 


(c) Frenzel did use for the Snipe the new specific name coelestis. In 
doing so, he gave no explanation of why he considered that a 
new name was needed for this species or why he considered the 
Linnean name gallinago unsuitable. Indeed, he made no 
reference whatever to Linnaeus. 


(d) In the early part of the discussion of this species Frenzel observed 
that the Snipe was the bird which sportsmen called the 
** Becasse ’’ and that by some writers it was known as Capella 
coelestis. His actual words were: ‘“‘Es ist die Schnepfe, 
welche die Jager eigentlich ‘ Becasse’ und einige Schriftsteiler 
Capella coelestis nennen’’. This is the only mention by 
Frenzel of the word ‘‘ Capella’’. 


4. Frenzel’s statement that the name or expression ‘‘ Capella 
coelestis’’ had been applied to the Snipe by some writers prompted 
me to undertake a further investigation of the literature with the 
object of determining, if possible, who were the authors referred to. 
So far as concerns the use of this expression as a scientific zoological 
name in the period from the starting point of zoology (1st January 
1758) to the appearance in 1801 of Frenzel’s Beschreibung, these 
investigations were completely negative in their results. These 
investigations did, however, throw some very interesting light on the 
origin and history of the expression ‘* Capella coelestis’’’ as a vernacular 
(Latin) denomination applied to the Snipe in the XVIIIth century 
and either to that bird or to some other marsh bird both in the Middle 
Ages and, in a Greek equivalent, as far back as classical times. Though 
only of indirect concern to the question of the scientific name to be 
used for the Snipe, the information so collected is of such antiquarian 
and historical interest that it should, in my opinion, be placed on 
permanent record. I have accordingly summarised this information 
in a separate note which I hereby direct be annexed as an Appendix 
to the Commission’s Direction on this case. I am happy to take this 
opportunity of expressing the grateful thanks of the International 


DIRECTION 39 125 


Commission to the two eminent scholars who have taken part in this 
interesting and unusual investigation, namely: Professor the Rev. 
L. W. Grensted, M.A., D.D., the Commission’s Consulting Classical 
Adviser, and Professor Charles Singer, M.A., D.Litt., M.D., D.Sc., 
F.R.C.P. (London), Professor Emeritus of the History of Science 
in the University of London. 


5. As regards the so-called generic name Capella Frenzel, 1801, the 
information provided by the photostat supplied by the Cambridge 
authorities which has been summarised in paragraph 3 above shows 
that Frenzel did not publish the above name as a new name nor did he 
adopt this name from some manuscript source. All that he did was to 
cite a binomen containing it—the binomen Capella coelestis—as a 
name or term which had been used by some writers to denote the 
Snipe. He did not suggest even that the writers to whom he was 
referring were binominal writers of works published subsequent to 
the appearance in 1758 of the Tenth Edition of the Systema Naturae 
and the investigations since undertaken (paragraph 4 above) confirm 
the view expressed in modern ornithological works that Frenzel was the 
first author subsequent to the starting point of zoological nomenclature 
to include the term “‘ Capella” in a work on ornithology. Thus, on 
the most favourable view of the status of the term Capella, that name, 
if it can be recognised as ranking as a name, could not be rated higher 
than a manuscript name published by Frenzel in synonymy without 
an independent description or indication of its own. On the less 
favourable of the two possible views as to its status—and the view 
which is almost certainly the correct view—the term ‘“‘ Capella” 
was not used by any binominal author even as a manuscript scientific 
name and that it owes its origin entirely to usages as an expression to 
denote the Snipe by authors prior to the starting point of zoological 
nomenclature, i.e. prior to the year 1758. In this connection we have 
to take particular note that the Fourteenth International Congress of 
Zoology, Copenhagen, 1953, decided that a name published in a 
synonymy and without an independent description, shall, as from a 
date to be specified, possess no status of availability under the Régles 
(1953, Copenhagen Decisions zool. Nomencl. : 63, Decision 115). 
The Congress decided also that none of the decisions on nomenclature 
taken by it should come into force until the first day of January of the 
year following that in which the revised text of the Régles was 
promulgated (ibid. : 103, Decision 196) but that in its day-to-day 
work the International Commission should guide itself by the decisions 
taken by that Congress (ibid. : 103, Decision 199). In these circum- 
stances the Commission is already bound to apply Decision 115 where 
cases of the type covered thereby come up for decision. Thus, even 
on the most favourable of the possible views which can be taken as to 
the status of the term Capella as published by Frenzel in 1801 (namely, 
that it is a manuscript generic name rejected by Frenzel and published 
by him in a synonymy without an independent description) the generic 


126 OPINIONS AND DECLARATIONS 


name Capella Frenzel, 1801, would be invalid under Decision 115 of the 
Copenhagen Congress. Accordingly, as Secretary to the International 
Commission, I hereby direct that, in accordance with the General 
Directive relating to the recording of objectively invalid names on 
the appropriate Official Indexes of Rejected and Invalid Names in 
Zoology issued to the Commission by the Thirteenth International 
Congress of Zoology, Paris, 1948, and in compliance also with the 
decision taken by the Commission provisionally in its vote on Voting 
Paper V.P.(O.M.)(55)36 in anticipation of the possibility of its later 
being found that the name Capella Frenzel, 1801, was invalid, the 
foregoing name, being a name published in synonymy and without 
an independent description, be placed on the Official Index of Rejected 
and Invalid Generic Names in Zoology in the Ruling to be given in the 
Direction embodying the decision taken by the Commission in its 
vote on the Voting Paper referred to above. 


6. The rejection of Capella Frenzel, 1801, as a name possessing no 
status of availability makes it necessary to re-examine the position 
of the generic name Capella Keyserling & Blasius, 1840, a name which 
by its vote on the Voting Paper specified above the Commission 
directed should be placed on the Official Index of Rejected and Invalid 
Generic Names in Zoology as being a junior homonym of Capella 
Frenzel, 1801. Now that Frenzel’s Capella is seen to possess no 
status of availability the name Capella Keyserling & Blasius, 1840, is 
no longer invalid as being a junior homonym of Frenzel’s name and, 
other things being equal, would become an available name nomen- 
clatorially even though it might not be required for taxonomic purposes. 
In fact, however, the name Capella Keyserling & Blasius is invalid 
for quite a different reason, for it is a junior objective synonym of 
Rupicapra Blainville, 1816 (Bull. Soc. Sci. philomat., Paris 1816 : 75), 
both genera having Capra rupicapra Linnaeus, 1758 (Syst. Nat. (ed. 10) 
1 : 68) as type species. Accordingly, the name Capella Keyserling & 
Blasius, 1840, being an objectively invalid name, remains eligible 
for admission to the Official Index of Rejected and Invalid Generic 
Names in Zoology and there is therefore no call to modify the decision 
taken on Voting Paper V.P.(O.M.)(55)36 that it be inscribed on the 
foregoing Index. It will be necessary, however, to revise the note of 
explanation to be added to this entry, so that instead of stating that 
this name is a junior homonym of Capella Frenzel, 1801, it shall state 
that it is a junior objective synonym of Rupicapra Blainville, 1816. 
As Secretary, I hereby direct that this change be made. 


21. Declaration of the Result of Vote : On 8th February 1956, 
Mr. Hemming, Secretary to the International Commission, 
acting as Returning Officer for the Votes taken on Voting Paper 
V.P.(O.M.)(55)36, signed a Certificate that the Votes cast were 
as set out in paragraphs 16, 17 and 18 above and declaring 


DIRECTION 39 127 


(i) that as regards Part I of the foregoing Voting Paper, the 
proposal there styled “* Alternative “A’ (acceptance of the 
name Gallinago)”’ had been duly adopted, that (ii) as regards 
Part 2 of the said Voting Paper, the proposal there styled 
“‘ Alternative ‘Y’ (acceptance of Gallinago as from Brisson, 
1760)’ had been similarly adopted, (iii) that the proposal 
submitted in Part 3 of the said Voting Paper had been likewise 
adopted, (iv) that the name Capella Frenzel, 1801, had been 
duly placed on the Official Index of Rejected and Invalid Generic 
Names in Zoology, (v) that the decisions so taken were the decisions 
of the Committee in the matters aforesaid but (vi), having 
regard to the fact that neither the decision taken on Part 1, 
nor the decision taken on Part 2 of the foregoing Voting Paper 
nor that relating to the name Capella Frenzel involved the use of 
the Commission’s Plenary Powers, no occasion arose for giving 
effect to the decision taken in Part 3 of the said Voting Paper, 
subject to the clarifications specified in the Minute executed by 
the Secretary on 8th February 1956 (reproduced in paragraph 20 
of the present Direction). 


22. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 8th February 1956 Mr. Hemming prepared the Ruling given 
in the present Direction and at the same time signed a Certificate 
that the terms of that Ruling were in full accord with the decision 
taken by the International Commission in its Votes on the 
several portions of Voting Paper V.P.(O.M.)(55)36. 


23. Original References: The following are the original 
references for the names placed on Official Lists and Official 
Indexes by the Ruling given in the present Direction :— 


Capella Frenzel, 1801, Beschr. Vogel und Eier Wittenberg : 56 
Capella Keyserling & Blasius, 1840, Wirbelth. Europas 1:9 
Gallinago Koch, 1816, Syst. baier. Zool. 1 : 312 

gallinago, Scolopax, Linnaeus, 1758, Syst. Nat. (ed. 10) 1 : 147 
media, Gallinago Koch, 1816, Syst. baier. Zool. 1 : 314 


24. Family-Group Names: No family-group-name problem 
arises in connection with the generic names dealt with in the 
present Direction. 


128 OPINIONS AND DECLARATIONS 


25. At the time of the submission of the present application 
the name applicable to the second portion of a binomen was 
‘trivial name’”’. This was altered to “specific name” by the 
Fourteenth International Congress of Zoology, Copenhagen, 
1953, which at the same time made corresponding changes in the 
titles of the Official List and Official Index of names of this 
category. These changes in terminology have been incorporated 
in the Ruling given in the present Direction. 


26. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in 
dealing with the present case, and the present Direction is 
accordingly hereby rendered in the name of the said International 
Commission by the under-signed Francis Hemming, Secretary 
to the International Commission on Zoological Nomenclature, 
in virtue of all and every the powers conferred upon him in that 
behalf. 


27. The present Direction shall be known as Direction Thirty- 
Nine (39) of the International Commission on Zoological 
Nomenclature. 


Done in London, this Eighth day of February, Nineteen 
Hundred and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


DIRECTION 39 129 


APPENDIX 


ON THE USE OF THE EXPRESSION ‘* CAPELLA 
COELESTIS ” TO DENOTE THE SNIPE 


by 


FRANCIS HEMMING, C.M.G., C.B.E. 


Secretary to the International Commission on Zoological 
Nomenclature 


‘- ai ¥ cael 


‘ia 


Ys ? 4. i =i ' ba 
‘ee 
is, . ; iy : : “Hh 


el eri, 
F ‘ , ; Se el ee vind 
7 oda noe Wi Ge ANAS ae 


FD ae wel 


DIRECTION 39 131 


On the use of the expression ‘‘ Capella coelestis ’’? to denote 
the Snipe 


By FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on 
Zoological Nomenclature) 


(1 plate) 


The object of the present note is to place on record certain 
information which has been collected regarding the origin of 
the use of the Latin expression “‘ Capella coelestis”’ to denote the 
Snipe. 


2. By way of introduction it is necessary to recall that for a 
number of years past an attempt has been made to replace as the 
generic name for the Snipe the name Gallinago Koch, 1816 by the 
name Capella Frenzel, 1801, on the ground that the latter had 
priority over the former. In an application (Z.N.(S.) 575) which 
I recently submitted to the Commission in connection with the 
preparations for the publication of the Official List of Generic 
Names in Zoology in book-form, I recommended that the entry 
thereon of the name Gallinago Koch, 1816, should be removed 
and that the name Gallinago Brisson, 1760, which was undoubtedly 
the oldest available generic name for the Snipe should be entered 
on the Official List in place of the name Gallinago Koch, 1816, 
which, whatever might be its subjective relationship to the name 
Capella Frenzel, 1801, was invalid as a junior homonym of 
Gallinago Brisson, 1760. This application involved, incidentally, 
consideration being given to the name Capella Frenzel, 1801, and 
to the binomen Capella coelestis which Frenzel was reputed 
to have published for the Snipe. The book by Frenzel in which 
these names were reputed to have been published is entitled 
Beschreibung der Vogel und ihrer Eier in der Gegend von 
Wittenberg. This little work, of which the original (and only) 
edition was no doubt very small is excessively scarce but 
ultimately with the assistance of Professor Ernst Mayr I ascer- 
tained that there was a copy in the Balfour Library of the 
Department of Zoology at Cambridge University. The Librarian 


132 OPINIONS AND DECLARATIONS 


of that Institution very kindly made available his Library’s copy 
of Frenzel’s book to the Librarian of the University Library who 
most obligingly had indicated his willingness to arrange for 
photographs to be made of the relevant pages. Very shortly 
after this the much desired photographs were received in the 
Office of the Commission. The warmest thanks of the Com- 
mission are due to the great assistance rendered by the Librarians 
of these Institutions. Examination of the photographs received 
at once showed that of the pages concerned the page which 
contained the information of which I was in search was page 58. 
A facsimile reproduction of this page is given in the plate annexed 
to the present paper. 


3. The receipt of the photograph of page 58 of Frenzel’s book 
made it possible at last to determine what in fact was the action 
in this matter taken by Frenzel in 1801. This proved to be 
totally different from that which had been attributed to him in all 
ornithological works in which his Beschreibung had found a 
notice. Frenzel, we now see, did apply the specific name coelestis 
to the Snipe but he did not introduce a new generic name con- 
sisting of the “‘ Capella’’, still less did he use a generic name 
consisting of this word as the generic name for this bird. On the 
contrary he placed the Snipe in the genus Scolopax as Scolopax 
coelestis, thus assigning it to the same genus as did Linnaeus 
when forty-three years earlier he gave it the name Scolopax 
gallinago in the Tenth Edition of the Systema Naturae of 1758. 
The word “* Capella’? occurs only once in Frenzel’s book, being 
used as a vernacular (Latin) name for the Snipe and cited as a 
name comparable with the vernacular (German) name “ Becasse ”’, 
which, as Frenzel explained, was the name applied to it by 
sportsmen (“die Jager’). The sentence in question reads as 
follows: “‘Es ist der Schnepfe, welche die Jager eigentlich 
* Becasse’’ und einige Schriftsteiler Capella coelestis nennen”’. 


4. The particulars given above dispose completely of the 
claims which have been advanced by which Frenzel has been 
credited with having introduced the generic name Capella in his 
little book of 1801 and with having used this name for the Snipe. 
This discovery, interesting as it is, has, however, been robbed of 
the practical significance which it would otherwise have possessed 
by the other discovery made in this case, namely that Brisson 


DIRECTION 39 158 


in 1760 established the nominal genus Gallinago for the Snipe 
and therefore that, even if there had been such a generic name as 
Capella Frenzel, 1801, that name would not have been required, 
as it would have been a junior subjective synonym of Gallinago 
Brisson, 1760. Nevertheless, it is very satisfactory that the 
reputed generic name Capella Frenzel, 1801, should have been 
shown up in its true colours as a mere phantom engendered of 
careless copying. 


5. While Frenzel’s book is now seen to have contained no new 
generic name consisting of the word ‘‘ Capella”’, a subject which 
proves to be of great antiquarian and historical interest is opened 
up by Frenzel’s statement that the Latin expression “*‘ Capella 
coelestis’’ had been used for the Snipe by several writers. This 
tantalising observation prompted me to undertake an investigation 
which, before it had run its course, was to give rise to the most 
surprising discoveries and was to show that the expression 
** Capella coelestis’”’ applied to the Snipe by Frenzel—or an 
equivalent Greek expression—was one of the greatest antiquity. 


6. As a start of the proposed investigation, I made a search 
in all likely places in the literature of ornithology in the period 
from 1758 to 1801 for the purpose of making sure that the 
binomen Capella coelestis or at least the generic name Capella 
had not been used by some obscure author as scientific names 
for the Snipe. In view of the extent to which the literature of this 
period had already been combed through by ornithologists, it 
was no matter for surprise that I was unable to find a trace 
of any such usage in the period subsequent to the publication in 
1758 of the Tenth Edition of the Systema Naturae of Linnaeus 
and prior to the publication of Frenzel’s Beschreibung in 1801. 
Concurrently with the above investigation I attempted also to 
ascertain whether during the same period either of the foregoing 
expressions had been applied to the Snipe not as scientific 
zoological names but as vernacular (Latin) designations. This 
search also proved entirely fruitless. It was just when I was on the 
point of abandoning the search that light was thrown on this 
subject from a most unexpected source. For, although no one 
subsequent to Linnaeus could be found to have used the elusive 
expression “* Capella coelestis’’, it suddenly came to light that in 


134 OPINIONS AND DECLARATIONS 


1758 this expression had been applied to the Snipe by no less a 
writer than Linnaeus himself (Syst. Nat. (ed. 10) 1 : 147). 
Linnaeus then gave the name Scolopax gallinago to the Snipe 
and, after describing this species, citing some bibliographical 
references and indicating its distribution, added a brief comment 
in which he applied the expression “‘ Capella coelestis” to this 
species. Linnaeus’s comment was as follows :— 


Haec Capella coelestis cujus mas tam alte celerrimeque 
volitat, ut audiri, non vero oculis attingi possit. 


7. Other works of Linnaeus were examined after the astonishing 
discovery described above but these threw no new light. on this 
subject. In the Twelfth Edition of the Systema Naturae published 
in 1766 (Syst. Nat. (ed. 12) 1(1) : 244) the comment quoted 
above was taken over from the Tenth Edition, though in a 
slightly amended form. It read as follows: “‘haec Capella 
coelestis, cujus mas tam alte celerrimeque volitat cadendo, ut 
audiri, vix vero oculis attingi possit, dum femina sedit humi’’. 


8. On making the surprising discovery described in paragraph 6 
above, I at once consulted Professor L. W. Grensted, Consulting 
Classical Adviser to the International Commission, who on 29th 
March 1956 replied as follows :— 


An intriguing problem—even after the primary question is settled. I 
think that the expression ‘‘ Capella coelestis ’’ was used by some author 
before Linnaeus and without a definitely binominal intention. The 
sentence you quote from Linnaeus i.e. the sentence quoted in paragraph 
4 above] can only mean :— 


** This [is] Capella coelestis, whose male flies so high and so 
swiftly that it can be heard but not reached by the eyes ”’. 


In other words Linnaeus is saying (presumably as against somebody 
else) that what some earlier writers have called Capella coelestis—the 
Heavenly Goat—is the Snipe and not some other bird, probably the 
Woodcock. 


The point would be that the Cock Snipe, when “ roding ’’, does go 
up to a great height (up to 400 ft. according to the books), while the 
Woodcock flies quite low—at tree-top height. 


_—"" 7" oe 


ee Te eT ae eT lee 


DIRECTION 39 135 


Is there a connection with the star called “‘ Capella ’’—also the 
Heavenly Goat? Some fanciful writer may easily have compared the 
roding of the Woodcock or the Snipe with bleating and linked up the 
bird with the star? 


9. In a further letter (dated 9th April 1956) Professor Grensted 
supplemented his earlier report as follows :— 


I did a little search round in our Oriel Library yesterday and found 
very little to help. The big Ducange Glossarium has nothing relevant. 
And the only thing I found in the early English books is that in 
Willughby’s Ornithology, page 228, the “‘ Lapwing ’’ or Green Plover 
is called Capella sive Vanellus. This book was edited or revised by 
John Ray, and is dated 1676. 


In mediaeval Latin “ capella’? seems to be entirely used for a 
chapel. It is derived from a very late Latin “‘ cappella ”’ (from “‘ cappa ’’, 
a cloak—originally the cloak cut in half by St. Martin of Tours). The 
classical meaning, a goat (more often “‘capra’’), survived in the 
star “‘ capella”’. How Willughby came to use it for a plover (probably 
from the star in some way) is quite obscure. 


10. At this point I consulted Professor Charles Singer, Professor 
Emeritus of the History of Science in the University of London, 
who, if anyone, I knew would be able to throw some light upon the 
mediaeval usage of the word “capella’’. The following is the 
interesting letter (dated 11th April 1956) received from Professor 
Singer in reply to this enquiry :— 

I cannot answer your question or explain “‘ capella”? but I can 
connect, from classical sources, the word “‘ capella” with a goat, a 
bird, and the constellation ‘‘ Capeila”’. This association may, at least, 
explain the term coelestis. 


Aristotle’s Historia Animalium, 593623 mentions. a marsh bird 
au€ (genitive aiyds) which D’Arcy Thompson conjecturally translates 
“horned grebe’’. The term “‘ goat like’’, ay-, is applied to other 
birds, e.g. aitye8os in 609a31 and 616b10. 


The early Greco-Latin and Latino-Greek glossaries before the 12th 
century have been collected and indexed. They yield an equation aié 
=“ capra”’ = “capella” and also the equation “‘ Capricornus”” = 
alyoképws = “signumincoelo’’. The star in Auriga that we name 
** Capella ’”’ is already so named by Pliny in Historia naturalis. 


_ Thus the classical association seems there, right enough, though 
its meaning is beyond me. Could the note of a snipe be compared to 
any noise made by a goat? 


136 OPINIONS AND DECLARATIONS 


11. The following is a copy of a letter (dated 14th April 1956) 
from Professor Grensted, to whom I had forwarded a copy of 
Professor Singer’s letter :— 


Queerer and queerer—and also nearer. Charles Singer’s letter is 
useful and revealing. I ought to have thought of looking at the Greek. 
Not only is there «’€é = “‘ capella’ (both as goat and as some sort of 
bird) but at& odpavios (= “capella coelestis’’) occurs more than 
once in a proverbial form in the Greek comic poets. ‘‘ The heavenly 
goat brings wealth’. What is wanted next is Maass on the Scholiasts 
on Aratus (the Greek astronomer-poet, who uses até of the star 
known, after him, as “‘ Capella’: see line 170 of his poem) and any 
evidence of the current astronomers or astrologers Linnaeus may have 
read. It will probably turn out, when we get at it, that some late 
mediaeval or sixteenth century writer makes the link—quite possibly 
with a knowledge of this Greek proverbial phrase—between the bird 
and the star, and gives the Latin form of the Greek. It might well have 
been in Pliny, but apparently isn’t. 


12. Further light was thrown on this matter by Professor 
Grensted in the following letter dated 19th June 1956 :— 


Capella coelestis: Y made one more effort when I was recently 
in Oxford and I got out ali the major works of Aldrovandus—which 
Linnaeus certainly knew. There is nothing directly relevant, and 
the one reference to Capella is a queer one under the heading 
** Stymphalides Aves’’. It reads: ‘‘ Hine author [sic] obscurus, qui 
de natura rerum scripsit, Vanellum avem vulgo dictam, seu Capellam 
Aristotelis, Stymphalidem Plinii esse arbitratus est’’. .(“‘ Hence an 
obscure author who wrote ‘On the Nature of Things’ thought that 
the bird commonly called Vanellus, or the Capella of Aristotle, is the 
Stymphalis of Pliny’”’.) This of course merely links Vanellus, Capella 
and a€ with some marsh bird, but it is interesting as showing that 
** Capella”’ and not the usual ‘* Capra’’ was used in some mediaeval 
version of Aristotle. But I got no further there. The reference to 
Aldrovandus is 1590, Ornithologia, Lib. X : 622. (The most curious 
thing about the passage quoted above is that Aldrovandus should 
have written “‘author’”’ for the ordinary “‘ auctor ’’.) 


13. It occurred to me on receiving the foregoing letter that it 
might be possible to obtain some further light on this subject 
by ascertaining whether Linnaeus had any astronomical or 
astrological works in his library. There is no separate catalogue 
of that library which for catalogue—though not for other— 
purposes has been incorporated in the general library of the 
Linnean Society of London. It is easy, however, to ascertain 


Opinions and Declarations, Vol. 1, Sect. D Plate | 


Betdclerdye, Atauda arborea. Dte Farbe tft der 
ibrigen siemlid) gleich), auch bat fle die Art, 
wo Gefabr su befitrdren, fb ntedersudritcten 5 
mur der Kopf it mehr weiblih. Cie Rlegt um 

‘Shige Haufenveife, und tabt tid auf Daumen 
nieder. Dad Cy gleichet den udvigen, jedod 
it ce clas weiser, und weniger vunteire. 

Beerfcuepfe SGimmrelssiege, Scolopax coelettis, 
eine befannte Art Sobnepfe, die von ihren Tb- 
nen, cic wie das Mecern dev Sregeit Eltn jen, 
den Namen ervalten har. Wenn Mefer Vogel 
das Meckern Hervorbringen will, fo erhcbt er 
fic) jederscit febr bed) in die Luft, und made 
algdann cinen Bogen unrerwarts nad dev red: 
ten Ceite, um mit feinem rechten Slugel ore 
nadabinender: Tine berverzubringen. Dtefes 
wiederholt cv fo fange, als er Wefabr gu be: 
firdten facy dann Pchiepe ex ta ctnem Sfuge 
febr febnell gu feinem Mete. Cott felattd 
fein Gefang des Vogels, oder Tone der Nehle, 
wie bey andern Vogein. Der Ktebiy bringe 
aud) ciacne Téne mic feinen Fiigqeta Hervor, 
fobaid man fich defjen Xeuce und erde nager. 

Es it die Sdhuepfe, weiche die Naaee ets 
gemtlicd) Becaive und erntae Zebriftfieder Ca- 
pella coelettis nennen. Cite leds ai den Cum: 
pfen ctufam init tore Weibder, und auper(t 
felten wird mai auch an grofen Seen iweb: 
rere yindct. 

Der Vogel hat bic Grove eines halowid: 

figen Nebhuhus; muv ue ex tanger ae ftrecte, ayer 

GHhnabel hat fat dre Lange des haiben Rorpers, 

tt vorne suaqecundet, oder vielncor bretr sul ments 

nen, der cbere cimwenig fanger, an dev Cptbe 
fchwars und mie decglcichen Warjden befebe, 
als of er mit Chagrin ubdcrjogen ware; dann 
wird der Dahuabes brauniich, unten geiblih, an 
ter Wurgel aber gruntlich; dte Mafenloder find 


Facsimile reproduction of page 58 of Frenzel, 1801, Beschreibung der Végel und 
ihrer Eier in der Gegend von Wittenberg : page containing the only use by 
Frenzel of the vernacular (Latin) expression ‘* Capella coelestis ’’. 


~ n p P LS 7 7 
P a? : a | 
am 6 z 
eu 


DIRECTION 39 137 


what were the books which Linnaeus actually possessed for 
every such book is indicated by a Gothic capital “L” in 
brackets (parentheses) in the Catalogue of the Printed Books 
and Pamphlets in the Society’s Library published in 1925. In 
hope of the clue suggested by Professor Grensted in his letter 
of 14th April the Society’s Catalogue was examined both by 
Professor Grensted and myself. Unfortunately, the results were 
negative. The library of Linnaeus is mainly medical and 
botanical. It contains no astronomical works—not even Tycho 
Brahe, or the much earlier Copernicus. There is an Aristotle— 
Aristoteles Stagarita de Historia Animalium—but this is only a 
translation published in 1504 (1514) and not likely to be of help 
in the present connection. There are two works on astrology in 
the library, namely Albinus Petrus Constantius, Magia astrologica, 
Hamburgi, 1716; Lemnius, De Astrologia, Antwerpiae, 1553. 
Neither of these seems very likely. Unfortunately, it was not 
practicable to examine either of these works, for Professor 
Grensted found on enquiry that neither is in the Bodleian 
Library. It looks therefore as though Linnaeus must have 
derived his information about the use of the expression “ Capella 
coelestis’’ from some book which was not in his own library. 


14. In communicating to me the results of his examination 
of the Linnean Society’s Catalogue (in litt., 4th July 1956) 
Professor Grensted added the following additional interesting 
ROE -— 


While in the Bodleian I looked up Aratus (the Greek astronomer, 
c. 270 B.c.) and found that he actually calls Capella Avé iepy “‘ the 
Sacred Goat’’, which is coming near. But the mediaeval commentaries 
which have been edited by E. Maass don’t help any further. There is 
no sign of the above expression having been used in connection with 
a bird. Still the phrase is interesting, coming on the way to the 
proverbial «i& odpavios and to Capella coelestis. 


15. While it has not been possible definitely to establish from 
whom was derived the expression “‘ Capella coelestis ’’, as used 
by Linnaeus, in 1758 as a vernacular (Latin) expression to denote 
the Snipe, the information supplied by Professor Grensted and 
by Professor Singer is of the greatest interest as showing that 
from the times of Classical antiquity down the centuries words 
meaning “ goat’ and “ goat-like”’ have been applied to some 


138 OPINIONS AND DECLARATIONS 


kind of marsh bird. There can be little doubt that this asso- 
ciation is based, as has been suggested by both the above authorities 
and as Colonel Richard Meinertzhagen (with whom I have dis- 
cussed this matter), agrees upon a real or fancied resemblance 
between the note of the birds concerned and the bleating of a goat. 


16. The grateful thanks of the Commission are due to Professor 
Grensted and to Professor Singer for the invaluable assistance 
which they have rendered in this interesting and unusual 
investigation. 


Printed in England by Metcatre & Cooper LiMiTED, 10-24 Scrutton St., London EC 2 


oe 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, cM.G., C.B.E. 


Secretary to the Commission 
VOLUME 1. SECTION D. Part D.5. Pp. 139—152 


DIRECTION 40 


Determination of the gender to be attributed to the 
names of fifty-five genera of the Phylum Arthropoda, 
exclusive of genera of Decapod Crustacea, placed on the 
Official List of Generic Names in Zoology in the period 


up to the end of 1936 
ww 
16 ess ) 


LONDON: 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
‘41, Queen’s Gate, London, S.W.7 
1956 


Price Nine Shillings 


(All rights reserved) 


Issued 17th September, 1956 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 40 


Ve | The Officers of the Commission 


Honorary Life President: Dr. Karl JORDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President : Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) 
(12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary: Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(Arranged in order of precedence by reference to date of election or of most recent re-election, 
as prescribed by the International Congress of Zoology) 


Professor H. BoscHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. pete LEMCHE (Universitetets Zoologiske Museum, Copenhagen, Denmark) (27th 
July 1948) 

Professor Teiso EsAKI (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEy (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Unstitute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (Sth July 1950) 

Professor Erich Martin HerInc (Zoologisches Museum der Humboldt-Universitdt zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMOND (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) (12th August 
1953) (President) 

Professor Harold E. Voxes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) (12th 
August 1953) 

Professor Béla HANKO (Mezdégazdasdgi Muzeum, Budapest, Hungary) (i2th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y., 
U.S.A.) (12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. Hottuuis (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (AS5th October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum V Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KiiHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

Professor F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 
1954) 

Professor Ernest MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico ToRTONESE (Museo di Storia Naturale ‘‘ G. Doria’’, Genova, Italy) 
(16th December 1954) 


DIRECTION 40 


DETERMINATION OF THE GENDER TO BE ATTRIBUTED 
TO THE NAMES OF FIFTY-FIVE GENERA OF THE 
PHYLUM ARTHROPODA, EXCLUSIVE OF 
GENERA OF DECAPOD CRUSTACEA, PLACED 
ON THE ‘“ OFFICIAL LIST OF GENERIC 
NAMES IN ZOOLOGY” IN THE 
PERIOD UP TO THE END OF 1936 


RULING :—(1) The gender to be attributed to each of 
the under-mentioned names of genera in the Phylum 
Arthropoda which were placed on the Official List of 
Generic Names in Zoology in the period up to the end 
of 1936 by the Rulings given in the Opinions severally 
specified below, is hereby determined as being the mascu- 
line gender :— 


(a) Class Insecta (12 names) : 


(i) Name included in Opinion 81 : 
Cimex Linnaeus, 1758 ; 
(it) Names included in Opinion 104: 


Anthocoris Fallén, 1814; Nabis Latreille, 
[1802—1803] ; Reduvius Fabricius, 1775; 
Ectobius Stephens, 1835 ; Gryllus Linnaeus, 
1758; Pediculus Linnaeus, 1758 ; Phthirus 
Leach, 1815 ; Pulex Linnaeus, 1758 ; 


(i11) Names included in Opinion 106 : 


Gasterophilus Leach, 1817; Oedemagena 
Latreille, 1818 ; Oestrus Linnaeus, 1758 ; 


(b) Class Diplopoda (one name) (Opinion 104) : 
Polydesmus Latreille, [1802—1803] ; 


ATT 717 1056 


142 OPINIONS AND DECLARATIONS: 


(c) Class Crustacea (4 names) (Opinion 104) : 


Diaptomus Westwood, 1836; Oniscus Lin- 
naeus, 17535" Porcelio“Latrcile., qie03— 
1804] : Gammarus Fabricius, 1775 ; 


(d) Class Merostomata (one name) (Opinion 104) : 
Limulus Miiller (O.F.), 1785 ; | 


(e) Class Arachnida (16 names) : 


(1) Names included in Opinion 73 : 


Argas Latreille, 1795 ; Dermacentor Koch, 
1844 ; Ixodes Latreille, 1795; Rhipicentor 
Nuttall & Warburton, 1908 ; Rhipicephalus 
Koch, 1844 ; 


(ii) Names included in Opinion 104 . 


Dendryphantes Koch, 1837; Latrodectus 
Walckenaer, 1805 ; Scorpio Linnaeus, 1758 ; 
Cheyletus Latreille, 1796 ; Chorioptes Ger- 
vais & van Beneden, 1859 ; Demodex Owen, 
1843 ; Dermanyssus Dugeés, 1834; Glycy- 
phagus Hering, 1838; Psoroptes Gervais, 
1841 ; Rhizoglyphus Claparéede, [1869] ; 


(ii1) Name included in Opinion 113: 
Sarcoptes Latreille, [1802—1803]. 


(2) The gender to be attributed to each of the under- 
mentioned names of genera in the Phylum Arthropoda 
which were placed on the Official List of Generic Names 
in Zoology in the period up to the end of 1936 by the 
Rulings given in the Opinions severally specified below, 1s 
hereby determined as the feminine gender :— 


(a) Class Insecta (9 names) : 


(i) Name included in Opinion 82 : 
Musca Linnaeus, 1758 ; 


DIRECTION 40 143 


(11) Names included in Opinion 104 : 


Podura Linnaeus, 1758 ; Notonecta Linnaeus, 
1758 ; Triatoma Laporte, [1832]; Forficula 
Linnaeus, 1758; Blatta Linnaeus, 1758 ; 
Periplaneta Burmeister, 1838 ; 


(iii) Name included in Opinion 106: 
Cephenemyia Latreille, 1818 ; 

(iv) Name included in Opinion 128 : 
Nycteribia Latreille, 1796 ; 


(b) Class Crustacea (one name) (Opinion 77) : 
Lepas Linnaeus, 1758 ; 


(c) Class Arachnida (5 names) : 
(1) Name included in Opinion 73 : 

Haemaphysalis Koch, 1844 ; 

(11) Names included in Opinion 104: 
Avicularia Lamarck, 1818 ; Dysdera Latreille, 
[1804]; Segestria Latreille, [1804] ; 

(iii) Name included in Opinion 128: 
Spinturnix Heyden, 1826. 


(3) The gender to be attributed to each of the under- 
mentioned names of genera in the Phylum Artropoda 
which were placed on the Official List of Generic Names in 
Zoology in the period up to the end of 1936 by the Rulings 
given in the Opinions severally specified below, is hereby 
determined as the neuter gender :— 


(a) Class Insecta (one name) (Opinion 106) : 
Hypoderma Latreille, 1818 ; 


(b) Class Crustacea (one name) (Opinion 104) : 
Armadillidium Brandt & Ratzeburg, [1831] ; 


144 OPINIONS AND DECLARATIONS 


(c) Class Trilobita (one name) (Opinion 88) : 
Otarion Zenker, 1833 ; 


(d) Class Arachnida (3 names) : 


(1) Names included in Opinion 73: 


Amblyomma Koch, 1844; Hyalomma Koch, 
1844 ; 


(11) Name included in Opinion 104 : 
Trombidium Fabricius, 1775. 


Il. THE SUBJECT MATTER OF THE PRESENT 
“DIRECTION 


The present Direction contains the fifth instalment of deter- 
minations of the gender attributable to the names of genera 
placed on the Official List of Generic Names in Zoology in the 
period up to the end of 1936. The following is the paper which 
formed the basis of the decisions taken by the Commission in 
the present case, which was submitted to the Commission on 
29th September 1955 :— 


Determination of the gender of names placed on the ‘* Official List 
of Generic Names in Zoology ’’ in the period up to the end of 
1936: Fifth Instalment (names of genera of Arthropods, 
excluding Decapod Crustacea) 


By FRANCIS HEMMING, C.M.G., C.B.E. 
(Secretary to the International Commission on Zoological Nomenclature) 


The present paper contains the fifth instalment of proposals for the 
determination of the gender to be attributed to generic names placed 
on the Official List of Generic Names in Zoology during the period up 
to the end of 1936 ; it is concerned with the names of genera of the 
Phylum Arthropoda other than those of Decapod Crustacea, which 
were dealt with in Voting Paper V.P.(O.M.)(55)6!. 


* The decision taken by the International Commission by its vote on the above 
Voting Paper has since been embodied in Direction 11 (1955, Ops. Decls. int. 
Comm. zool. Nomencl. 1(e) : 15—34). 


DIRECTION 40 145 


2. The pre-Lisbon Opinions contain Rulings placing the names of 
fifty-six genera of the above group of Arthropods on the Official List. 
The distribution of names by Opinions is as follows :—Opinion 73 : 
8 names ; Opinions 77, 81, 82, 88 and 113 : one name each ; Opinion 
104: 36 names; Opinion 106: 5 names; Opinion 128: 2 names. 


3. As in the case of the previous instalments of proposals relating 
to genders, the present recommendations are based upon the Report 
furnished by Mr. F. J. Lelievre (at that time of the Classical Languages 
Department of Bedford College, London University), on the gender 
to be assigned to each name placed on the Official List in the period 
prior to the Paris (1948) Congress. Also, current checklists, catalogues 
and monographs have been consulted to ascertain whether the genders 
now proposed to be assigned to the names in question are those 
currently used by specialists in the various groups. Finally, drafts 
of the relevant portions of the list forming Annexe 2 to this paper 
were sent to various specialists for comment. The Commission is 
much indebted to the following specialists for their co-operation in 
this matter :— 


Insecta : Dr. Theresa Clay, Mr. H. Oldroyd, and Dr. W. E. 
China (British Museum (Natural History), London). 


Crustacea : Dr. J. P. Harding (British Museum (Natural History), 
London). 


Prof. A. Vandel (Université de Toulouse, Toulouse, 
France). 


Myriapoda & Dr. G. O. Evans and Dr. A. C. Townsend (British 
Arachnida : Museum (Natural History), London). 


Trilobita : Dr. C. J. Stubblefield (Geological Survey and Museum, 
e*“POndon). 


In only three cases were comments made by specialists on the genders 
proposed in the list submitted by Mr. Lelievre. These cases are 
discussed in the immediately following paragraphs. 


4. Lepisma Linnaeus, 1758: This is a Greek neuter word, but it 
was treated as feminine by the original author and, Miss Clay informs 
me, has been consistently used as such ever since. One species 
(L. saccharina) is widely used in the non-specialist literature and the 
change to the correct gender would inevitably lead to confusion. 
A separate application (File Z.N.(S.) 988) is therefore being made 
for the use of the Plenary Powers to vary the gender correctly 
attributable to this name?. 


* For the separate application subsequently submitted in regard to the gender 
to be attributed to the generic name Lepisma Linnacus see Hemming, 1955, 
Bull. zool. Nomencl. 11 : 299—300. 


146 OPINIONS AND DECLARATIONS 


5. Oedemagena Latreille, 1818 : Mr. Lelievre reports that this word 
is “common, on the analogy of Trojugena ”’, and should therefore be 
treated as masculine. This view is in agreement with the Rules adopted 
by the Copenhagen Congress (1953, Copenhagen Decisions Zool. 
Nomencl., Decision 84(4)). Mr. Oldroyd informs me that it is a 
relatively obscure genus with a single known species (O. tarandi, 
the Warble Fly of Reindeer) and therefore une attribution of the 
correct gender will cause no difficulty. 


6. Nabis Latreille, [1802—1803]: Mr. Lelievre reports that this 
name is feminine. Dr. China, in a letter dated 6th January, 1955, 
reproduced in Annexe 1 to this paper, states that it has always been 
treated by hemipterists as masculine. Lewis & Short give a feminine 
usage of the word “nabis”’, meaning a cameleopard, and as a 
masculine word it was the name of a king of Sparta. Since it is un- 
certain which meaning was intended for the generic name Nabis by the 
author, Latreille, it ts proposed, on the basis of Dr. China’s letter, 
to treat it as masculine. 


7. No difficulties have been found as regards the remaining fifty- 
three names ; in each case there appears to be no doubt as to the 
gender correctly attributable, and this is also the gender commonly 
used. 


8. The genders attributable to the fifty-five names dealt with in this 
paper are given in Annexe 2, and for the reasons set out above, I 
recommend that the International Commission should render a 
Direction that the gender specified in that Annexe for each of the names 
concerned be now entered against that name in the Official List of 
Generic Names in Zoology. 


ANNEXE oi 


Gender proposed to be assigned to the generic name ‘‘ Nabis ”’ 
Latreille 


Extract from a letter, dated 6th January, 1955, from Dr. W. E. 
China Secs Museum (Natural History), London) 


Nabis Latreille, 1802, which you list as feminine, is always regarded 
by Hemipterists as masculine. 


Amyot, 1848, Rhynchotes, Méthode Mononymique (a work which 
is invalid) gives the meaning of Nabis as “‘ Brebis sauvage’’, a wild 


DIRECTION 40 147 


ewe, presumably with the-idea that the predatory Nabis is ‘“‘a wolf 
in sheep’s clothing’. If Nabis really was a classical word meaning 
ewe, it would, of course, be feminine, but I can trace no Latin or 
Greek word Nabis meaning a female sheep. Nabis was a King of 
Sparta about 200 B.c. which would make the genus masculine. Nabis 
is also an African word for Giraffe. 


BONN BOE: 2 


Gender proposed to be inscribed in the ‘‘ Official List of Generic Names 
in Zoology ’’ in respect of the names of genera of Arthropoda 
other than those of Decapod Crustacea, placed on that ‘‘ List ”’ 
in the period up to the end of 1936 


Gender ** Opinion” in 
proposed which name 
to be assigned specified in Col. | 
to name speci- was placed on 
Generic Name fredin Cols“ Official List” 


(1) (2) (3) 


Class Insecta 


Podura Linnaeus, 1758 Feminine 104 
Musca Linnaeus, 1758 Feminine 82 
Cephenemyia Latreille, 1818 Feminine 106 
Gasterophilus Leach, 1817 Masculine “ 
Hypoderma Latreille, 1818 Neuter i 
Oedemagena Latreille, 1818 Masculine ie 
Oestrus Linnaeus, 1758 Masculine - 
Nycteribia Latreille, 1796 Feminine 128 
Cimex Linnaeus, 1758 Masculine 81 
Anthocoris Fallén, 1814 Masculine 104 
Nabis Latreille, [1802—1803] Masculine “5 
Notonecta Linnaeus, 1758 Feminine ey 
Reduvius Fabricius, 1775 Masculine o 
Triatoma Laporte, [1832] Feminine ‘ 
Forficula Linnaeus, 1758 Feminine _ 
Blatta Linnaeus, 1758 Feminine s: 
Ectobius Stephens, 1835 Masculine a 
Gryllus Linnaeus, 1758 Masculine £ 
Periplaneta Burmeister, 1838 Feminine re 
Pediculus Linnaeus, 1758 Masculine ‘- 
Phthirus Leach, 1815 Masculine e 
Pulex Linnaeus, 1758 Masculine Ly 


148 OPINIONS AND DECLARATIONS 


Gender ** Opinion’? in 
proposed which name 
to be assigned specified in Col. | 
to name speci- was placed on 
Generic Name fied in Col. 1 “* Official List ”’ 


(1) (2) (3) 
Class Diplopoda 
Polydesmus Latreille, [1802—1803] Masculine 104 


Class Crustacea 


Diaptomus Westwood, 1836 Masculine 104 

Lepas Linnaeus, 1758 Feminine vi 

Armadillidium Brandt & Neuter 104 
Ratzeburg, [1831] 

Oniscus Linnaeus, 1758 Masculine - 

Porcellio Latreille, [1803—1804] Masculine * 

Gammarus Fabricius, 1775 Masculine 


Class Merostomata 


Limulus Miiller (O.F.), 1785 Masculine 104 


Class Trilobita 


Otarion Zenker, 1833 Neuter 88 
Class Arachnida 
Avicularia Lamarck, 1818 Feminine 104 
Dendryphantes Koch, 1837 ~ Masculine a 
Dysdera Latreille, [1804] Feminine " 
Latrodectus Walckenaer, 1805 Masculine 2 
Segestria Latreille, [1804] Feminine Ms 
Scorpio Linnaeus, 1758 Masculine es 
Amblyomma Koch, 1844 Neuter 73 
Argas Latreille, 1795 Masculine Es 
Dermacentor Koch, 1844 Masculine ¥ 


Haemaphysalis Koch, 1844 Feminine 


DIRECTION 40 


Gender 
proposed 


149 


** Opinion’ in 
which name 


to be assigned specified in Col. 1 


to name speci- 


Generic Name 


(1) 


Hyalomma Koch, 1844 

Ixodes Latreille, 1795 

Rhipicentor Nuttall & Warburton, 1908 
Rhipicephalus Koch, 1844 

Cheyletus Latreille, 1796 


Chorioptes Gervais & van Beneden, 
1859 


Demodex Owen, 1843 
Dermanyssus Dugés, 1834 
Glycyphagus Hering, 1838 
Psoroptes Gervais, 1841 
Rhizoglyphus Claparéde, [1869] 
Trombidium Fabrictus,® 1775 
Sarcoptes Latreille, [1802—1803] 
Spinturnix Heyden, 1826 


(2) 


Neuter 

Masculine 
Masculine 
Masculine 
Masculine 
Masculine 


Masculine 
Masculine 
Masculine 
Masculine 
Masculine 
Neuter 
Masculine 
Feminine 


was placed on 
jean: Cole +* Official List” 


(3) 


as 


2. Registration of the present application : Upon the receipt 
of Mr. Hemming’s paper, the question of the gender to be assigned 
to the names of genera of the Phylum Arthropoda dealt with 
therein was allotted the Registered Number Z.N.(S.) 989. 


II. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


3. Issue of Voting Paper V.P.(O.M.)(55)29 : On 29th September 
1955, a Voting Paper (V.P.(O.M.)(55)29) was issued in which 


150 OPINIONS AND DECLARATIONS 


each Member of the Commission was asked (1) to state whether 
he agreed “ that, in conformity with the General Directive relating 
to the recording in the Official List of Generic Names in Zoology 
of the gender of each name placed thereon prior to 1948, issued 
to the International Commission by the Thirteenth International 
Congress of Zoology, Paris, 1948, the gender of the fifty-five 
generic names in the Phylum Arthropoda specified in Column (2) 
of Annexe 2 to the paper by the Secretary bearing the Registered 
Number Z.N.(S.) 989, submitted simultaneously with the present 
Voting Paper [i.e. in the corresponding column in the Annexe 
numbered as above attached to the paper reproduced in the first 
paragraph of the present Direction] be entered in the Official 
List of Generic Names in Zoology in respect of the names enumer- 
ated in the first column of the said Annexe ”’, and (2), if he did not 
agree as regards any given item, to indicate that item. 


4. The Prescribed Voting Period: As the foregoing Voting 
Paper was issued under the One-Month Rule, the Prescribed 
Voting Period closed on 29th October 1955. 


5. Particulars of the Voting on Voting Paper V.P.(O.M.)(55)29 : 
At the close of the Prescribed Voting Period, the state of the voting 
on Voting Paper V.P.(O.M.)(55)29 was as follows :— 


(a) Affirmative Votes had been given by the following twenty- 
four (24) Commissioners (arranged in the order in which 
Votes were received) : 


Riley ; Boschma ; Holthuis; Vokes; Hering; Stoll ; 
Prantl; Lemche; Mayr; Key; do Amaral; Esaki ; 
Mertens ; Miller ; Bradley (J.C.) ; Kiihnelt ; Dymond ; 
Jaczewski; Hemming; Sylvester-Bradley ; Tortonese ; 
Bonnet ; Cabrera ; Hanko ; 


(b) Negative Votes: 


None ; 


DIRECTION 40 151 


(c) On Leave of Absence, one (1) : 


Bodenheimer ; 


(d) Voting Papers not returned : 


None. 


6. Declaration of Result of Vote: On 29th October 1955, 
Mr. Hemming, Secretary to the International Commission, acting 
as Returning Officer for the Vote taken on Voting Paper V.P. 
(O.M.)(55)29, signed a Certificate that the Votes cast were as set 
out in paragraph 5 above and declaring that the proposal sub- 
mitted in the foregoing Voting Paper had been duly adopted and 
that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


7. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 22nd February 1956, Mr. Hemming prepared the Ruling 
given in the present Direction and at the same time signed a 
Certificate that the terms of that Ruling were in complete accord 
with those of the proposal approved by the International Com- 
mission in its Vote on Voting Paper V.P.(O.M.)(55)29. 


8. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in dealing 
with the present case, and the present Direction is accordingly 
hereby rendered in the name of the said International Com- 
mission by the under-signed Francis Hemming, Secretary to the 
International Commission on Zoological Nomenclature, in virtue 
of all and every the powers conferred upon him in that behalf. 


£52 OPINIONS AND DECLARATIONS 


9. The present Direction shall be known as Direction Forty (40) 
of the International Commission on Zoological Nomenclature. 


Done in London, this Twenty-second day of February, Nine- 
teen Hundred and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


ee 


Printed in England by Metcatre & Cooper LimiTeED, 10-24 Scrutton St., London EC 2 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, c.M.G., C.B.E. 


Secretary to the Commission 
VOLUME 1. SECTION D. Part D.6. Pp. 153—170 


DIRECTION 42 


Determination of the gender to be attributed to the names 

of seventy-eight genera of various Classes of Invertebrates 

and of six genera of the Class Urochorda placed on the 

Official List of Generic Names in Zoology in the period 
up to the end of 1936 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956 


Price Twelve Shillings 
(All rights reserved) 


_—— oe oe eS 


AJ 


OCT 16 1955 
Nii. 


\ 
4 
1 
1 
1 
, 
) 
| 


S 


Issued 17th September, 1956 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 42 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JoRDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President : Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) 
(12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(Arranged in order of precedence by reference to date of election or of most recent re-election 
as prescribed by the International Congress of Zoology) 


Professor H. BoSCHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr; Ty ee (Universitetets Zoologiske Museum, Copenhagen, Denmark) (27th 
July 194 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEY (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Unstitute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (Sth July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitat zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMoND (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) (12th August 
1953) (President) 

Professor Harold E. Vokes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) (12th 
August 1953) 

Professor Béla HANKO (MezGgazdasdgi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y., 
U.S.A.) (12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. HoLtuuis (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (15th October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum vy Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KiHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

Seen S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 

Professor Ernest MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico TORTONESE (Museo di Storia Naturale ‘‘ G. Doria’’, Genova, Italy) 
(16th December 1954) 


le ee SS ee 


ee EE 


——_— ee 


DIRECTION 42 


DETERMINATION OF THE GENDER TO BE ATTRIBUTED 
TO THE NAMES OF SEVENTY-EIGHT GENERA OF 
VARIOUS CLASSES OF INVERTEBRATES AND OF 
SIX GENERA OF THE CLASS UROCHORDA PLACED 
ON THE ‘** OFFICIAL LIST OF GENERIC NAMES 
IN ZOOLOGY ” IN THE PERIOD UP TO THE 
END OF 1936 


RULING :—(1) The gender to be attributed to each 
of the under-mentioned names of genera of the Classes 
of Invertebrates severally specified below and of the 
Class Urochorda which were placed on the Official List 
of Generic Names in Zoology in the period up to the end 
of 1936 by the Rulings given in the Opinions specified 
in each case against the generic name concerned is hereby 
determined as being the masculine gender :— 


(a) Class Trematoda (3 names) : 
(i) Hemiurus Rudolphi, 1809 (Opinion 77) ; 
(11) Gastrodiscus Cobbold, 1877 ; Heterophyes Cob- 
bold, 1866 (both in Opinion 84) ; 
(b) Class Cestoda (1 name) : 
Echinococcus Rudolphi, 1801 (Opinion 84) ; 


(c) Class Nematoda (5 names) : 
(i) Dracunculus Reichard, 1759; Strongyloides 
Grassi, 1879 ; Trichostrongylus Looss, 1905 
(all in Opinion 66) ; 
(ii) Strongylus Miiller, 1780; Syngamus Siebold, 
1836 (both in Opinion 104) ; 
(d) Class Nematomorpha (2 names) : 
Gordius Linnaeus, 1758 ; Paragordius Camerano, 
1897 (both in Opinion 66) ; 
(e) Class Chaetopoda (2 names) : 
(i) Lumbricus Linnaeus, 1758 (Opinion 75) ; 
(ii) Enchytraeus Henle, 1837 (Opinion 104) ; 


156 OPINIONS AND DECLARATIONS 


(f) Class Gastropoda (1 name) : 
Limax Linnaeus, 1758 (Opinion 94) ; 


(g) Class Pelecypoda (1 name) : 
Mytilus Linnaeus, 1758 (Opinion 94) ; 
(h) Class Cephalopoda (1 name) : 
Argonauta Linnaeus, 1758 (Opinion 94) ; 


(1) Class Brachiopoda (1 name) : 
Spirifer Sowerby, 1816 (Opinion 100) ; 
(j) Class Crinoidea (4 names) : 

Bathycrinus Thomson, 1872 ; Holopus d’Orbigny, 
1837 3 Metacrinus Carpenter, (1 secu 
Rhizocrinus Sars, [1865] (all in Opinion 73) ; 

(k) Class Urochorda (1 name): 
Botryllus Gaertner, 1774 (Opinion 94). 


(2) The gender to be attributed to each of the under- 
mentioned names of genera of the Classes of Invertebrates 
severally specified below and of the Class Urochorda 
which were placed on the Official List of Generic Names 
in Zoology in the period up to the end of 1936 by the 
Rulings given in the Opinions specified in each case 
against the generic name concerned is hereby determined 
as being the feminine gender :— 


(a) Class Rhizopoda (2 names) : 
(i) Arcella Ehrenberg, [1830] (Opinion 77) ; 
(11) Lepidocyclina Giimbel, [1870] (Opinion 127) ; 


(b) Class Mastigophora (1 name) : 

Volvox Linnaeus, 1758 (Opinion 75) ; 
(c) Class Ciliata (1 name): 

Bursaria Miller, 1773 (Opinion 104) ; 
(d) Class Sporozoa (1 name) : 

Eimeria Schneider, 1875 (Opinion 104) ; 


DIRECTION 42 LS7 


(e) Class Neosporidia (1 name) : 
Sarcocystis Lankester, 1882 (Opinion 104) ; 


(f) Class Hydrozoa (2 names) : 
(i) Hydra Linnaeus, 1758 (Opinion 77) ; 
(ii) Physalia Lamarck, 1801 (Opinion 80) : 


(g) Class Trematoda (1 name) : 
Fasciola Linnaeus, 1758 (Opinion 84) ; 


(h) Class Cestoda (7 names) : 


(i) Anoplocephala Blanchard, 1848 ; Hymenolepis 
Weinland, 1858 ; Moniezia Blanchard, 1891 ; 
Stilesia Stiles & Hassall, 1893 (all in Opinion 
Woe 

(11) Davainea Blanchard & Railliet, 1891 ; Taenia 
Linnaeus, 1758 (both in Opinion 84) ; 


(ii) Ligula Bloch, 1782 (Opinion 104) ; 


(i) Class Nematoda (4 names) : 
(i) Ascaris Linnaeus, 1758 (Opinion 66) ; 
(ii) Filaria Miller, 1787; Heterodera Schmidt, 


1871; Rhabditis Dujardin, 1845 (all in 
Opinion 104) ; 


(j) Class Hirudinea (3 names) : 
(i) Hirudo Linnaeus, 1758 (Opinion 75) ; 


(11) Haemadipsa Tennent, 1859 ; Limnatis Moquin- 
Tandon, 1827 (both in Opinion 104) ; 


(k) Class Gastropoda (11 names) : 


(i) Calyptraea Lamarck, 1799 ; Columbella 
Lamarck, 1799; MHelix Linnaeus, 1758 ; 
Physa Draparnaud, [1801]; Succinea Drapar- 
naud, [1801] (all in Opinion 94) ; 


(ui) Leucochila von Martens, 1860 (Opinion 115) ; 


ArT 111956 


158 OPINIONS AND DECLARATIONS 


(iii) Oleacina Réding, 1798; Neritina Lamarck, 
1816 ; Clausilia Draparnaud, [1805] ; Vitrina 
Draparnaud, [1801]; Tornatellina Pfeiffer, 
1842 (all in Opinion 119) ; 

(1) Class Pelecypoda (4 names) : 
Anodonta Lamarck, 1799 ; Mya Linnaeus, 1758, 
Teredo Linnaeus, 1758; Mactra Linnaeus, 
1767 (all in Opinion 94) ; 
(m) Class Cephalopoda (1 name) : 
Sepia Linnaeus, 1758 (Opinion 94) ; 
(n) Class Brachiopoda (1 name) : 
Syringothyris Winchell, 1863 (Opinion 100) ; 
(o) Class Asteroidea (1 name) : 
Luidia Forbes, 1839 (Opinion 129) ; 
(p) Class Holothuroidea (1 name) : 
Holothuria Linnaeus, 1767 (Opinion 80) ; 
(q) Class Crinoidea (1 name) : 
Antedon Fréminville, 1811 (Opinion 73) ; 
(r) Class Urochorda (4 names) : 


Clavelina Savigny, 1816 ; Diazona Savigny, 1816 ; 
Distaplia Della Valle, 1881 ; Molgula Forbes, 
1848 (all in Opinion 94). 


(3) The gender to be attributed to each of the under- 


mentioned names of genera of the Classes of Inverte- 
brates severally specified below and of the Class Uro- 
chorda which were placed on the Official List of Generic 


Names in Zoology in the period up to the end of 1936 
by the Rulings given in the Opinions specified in each 
case against the generic name concerned is hereby 
determined as being the neuter gender :— 


(a) Class Mastigophora (1 name) : 
Trypanosoma Gruby, 1843 (Opinion 95) ; 


DIRECTION 42 159 


(b) Class Anthozoa (1 name) : 
Lithostrotion Fleming, 1828 (Opinion 117) ; 


(c) Class Trematoda (2 names) : 
(1) Schistosoma Weinland, 1858 (Opinion 77) ; 
(11) Dicrocoelium Dujardin, [1844] (Opinion 84) ; 
(d) Class Cestoda (2 names) : 
(i) Thysanosoma Diesing, 1835 (Opinion 77) ; 
(11) Dipylidium Leuckart, 1863 (Opinion 84) ; 
(e) Class Nematoda (2 names) : 
Ancylostoma Dubini, 1843 ; Gnathostoma Owen, 
1837 (both in Opinion 66) ; 
(f) Class Gastropoda (3 names) : 
(1) Buccinum Linnaeus, 1758 (Opinion 94) ; 
(11) Cerion Réding, 1798 (Opinion 119) ; 
(i1) Agasoma Gabb, [1869] (Opinion 12) ; 
(g) Class Scaphopoda (1 name) : 
Dentalium Linnaeus, 1758 (Opinion 94) ; 


(h) Class Pelecypoda (1 name) : 
Sphaerium Scopoli, 1777 (Opinion 94) ; 
(i) Class Cephalopoda (1 name) : | 
Lytoceras Suess, 1865 (Opinion 130) ; 


(j) Class Urochorda (1 name) : 
Pyrosoma Peron, 1804 (Opinion 77). 


I. THE SUBJECT MATTER OF THE PRESENT 
DIRECTION? 


The present Direction contains the sixth instalment of deter- 
minations of the gender attributable to the names of genera 
placed on the Official List of Generic Names in Zoology in the 
period up to the end of 1936. In all, eighty-four generic names 
are dealt with in the present Direction; of these, seventy-eight 
are the names of genera of Classes of invertebrates belonging to 


160 OPINIONS AND DECLARATIONS 


various Phyla, other than the Phylum Arthropoda, and six are 
the names of genera belonging to the Class Urochorda. The 
adoption of the present Direction marks the close of the survey 
of the gender to be attributed to each of the five hundred and 
sixty-three names placed on the Official List up to the end of 
1936 undertaken by the International Commission in compliance 
with a Directive issued to it by the Thirteenth International 
Congress of Zoology, Paris, 1948. Of the total number of names 
involved the gender attributable to five hundred and forty-six 
names has now been determined by the Commission, and proposals 
are now before it in the case of the remaining seventeen names. 
The paper on which the present Direction is based was submitted 
by the Secretary on 29th September 1955 and was as follows :— 


Determination of the gender of names placed on the ‘‘ Official List of 
Generic Names in Zoology ”’ in the period up to the end of 1936 : 
Sixth Instalment (names of genera of Invertebrates, other 
than Arthropods, and of genera of the Class Urochorda) 


By FRANCIS HEMMING, C.M.G., C.B.E. 
(Secretaty to the International Commission on Zoological Nomenclature) 


This paper contains the sixth and last instalment of proposals for 
the determination of the gender to be attributed to names placed on 
the Official List of Generic Names in Zoology during the period up to 
the end of 1936; it is concerned with genera belonging (a) to 
various Invertebrate Phyla, excluding the Arthropods (which have 
been dealt with in two previous Voting Papers, namely V.P.(O.M.)(55)6 
and V.P.(O.M.)(55)29! and (b) to the Class Urochorda. 


2. The pre-Lisbon Opinions contain Rulings placing the names of 
ninety genera of the foregoing groups on the Official List. The 
distribution of names by Opinions is as follows: Opinion 66: 9 
names ; Opinions 73: 5 names; Opinion 75: 3 names ; Opinion 77: 
10 names ; Opinion 84: 9 names ; Opinion 94: 22 names ; Opinion 
104: 14 names; Opinion 119: 7 names; Opinions 80, 95 and 100: 
2 names each ; Opinions 115, 117, 127, 129 and 130: one name each. 


3. As in the case of the previous instalments of proposals relating 
to the gender to be attributed to generic names on the Official List, 


1 The decisions taken by the International Commission by its votes on the fore- 
going Voting Papers have since been embodied in Direction 11 (1955, Ops. 
Decls. int. Comm. zool. Nomencl. 1(C) : 15—-34) and Direction 40 (pp. 139—152 
of the present Section (Section D) of Volume 1 of the present Series) 
respectively. 


DIRECTION 42 161 


the present recommendations are based upon the Report furnished by 
Mr. F. J. Lelievre (at that time of the Classical Language Department 
of Bedford College, London University). Also, current checklists, 
catalogues and monographs have been consulted to ascertain whether 
the genders now proposed to be assigned to the names in question are 
those currently used by specialists in the various groups. Finally, 
drafts of the relevant portions of the list now submitted as Annexe 2 
to this paper were sent to various specialists for comment. The 
Commission is much indebted to the following specialists for their 
co-operation in this matter :— 


Dr. Ellsworth C. Dougherty (University of California) (Protozoa, 
Trematoda, Cestoda and Nematoda) 


Dr. W. J. Rees, D.Sc. (British Museum (Natural History)) 
(Coelenterata) 


Mr. N. Tebble, B.Sc. (British Museum (Natural History)) (Hirudinea) 


Mr. R. V. Melville, M.Sc. (Geological Survey & Museum, London) 
(Cephalopoda) 


Dr. L. Bairstow, M.A., Ph.D. (British Museum (Natural History)) 
(Cephalopoda) 


4. Of the ninety names concerned, the gender attributable to five 
has already been dealt with. These are Necator Stiles, 1903 (Opinion 
201) ; Endamoeba Leidy, 1879 (Opinion 312); Plasmodium Marchiafava 
& Celli, 1885 and Laverania Feletti & Grassi, 1889 (Opinion 283) ; 
Ostrea Linnaeus, 1758 (Opinion 356). 


5. In the case of one further name, Gigantorhynchus Hamann, 1892, 
an application has been submitted for the use of the Plenary Powers 
to vary the gender correctly attributable to it. (File Z.N.(S.) 942 ; 
see Bull. Zool. Nomencl. 11 : 260—262.) 


6. Proposals regarding the remaining 84 names are now placed before 
the Commission. Only two of them call for special comment, namely 
Argonauta Linnaeus, 1758, and Antedon Fréminville, 1811. These are 
discussed in the immediately following paragraphs. 


7. Argonauta: This, according to the Latin dictionaries, is a 
masculine word, although it has an apparently feminine termination. 
Both Mr. Melville and Dr. Bairstow have been consulted in this case 
and it seems that, in spite of the fact that many authors have treated 
this name as being feminine, no serious inconvenience would result 
from attributing to it the correct (masculine) gender. The letters 
from these specialists are reproduced in Annexe 1. 


8. Antedon: This name is commonly treated as being of the 
feminine gender, but doubts have been expressed as to whether this is 
correct. Dr. L. W. Grensted, the Commission’s Consulting Classical 


162 OPINIONS AND DECLARATIONS 


Adviser, has reported (Annexe 2) that Antedon is probably an error for 
Anthedon, which is a feminine word. The likelihood that this is the 
correct explanation is strengthened by the fact that Agassiz (1846, 
Nomencl. Zool. Index Uniy. : 25) emended it to Anthedon. Fréminville 
himself did not give any information regarding the derivation of his 
new generic name, and the name of the single originally included 
species (gorgonia) is a feminine noun in apposition. In these circum- 
stances, and having regard to Dr. Grensted’s Report, I recommend 
that the Commission should assign the feminine gender to this name. 


9. As regards the remaining 82 names, in each case there appears 
to be no doubt as to the gender correctly attributable, and that gender 
is also the one commonly used. 


10. I accordingly recommend that, in compliance with the General 
Directive issued by the Thirteenth International Congress of Zoology, 
Paris, 1948, the Commission should now render a Direction that in 
respect of the seventy-eight generic names of Invertebrates and six 
generic names of Urochordates specified in Annexe 3 of the present 
paper, the gender to be inserted in the entry on the Official List of 
Generic Names in Zoology be that shown against each name in column 
(2) of the said Annexe. 


11. It will, I am confident, be a source of satisfaction to the 
Commission—as it is to myself as Secretary—that when it has taken 
decisions on the proposals now submitted, it will have completed the 
determination of the gender to be attributed to all the generic names 
placed on the Official List prior to the Paris Congress of 1948, with 
the exception of a small group of names as regards which special 
problems arise and on which separate proposals have been, or shortly 
will be, submitted for decision. The taking of these decisions marks 
an important stage in the preparations needed before the urgently 
awaited volume containing particulars of all names so far placed on the 
Official Lists and Official Indexes can be sent to the printer for 
publication. 


ANNEXE 1 


The gender attributable to the generic name ‘‘ Argonauta ’’ Linnaeus, 


(a) Extract from a letter from R. V. Melville, M.Sc. (Geological Survey 
and Museum, London), dated 10th January 1955 


Argonauta is obviously more difficult, since it is a masculine noun 
with an apparently feminine ending. I have looked it up in Sherborn’s 
Index Animalium and find (apart from nouns in apposition, e.g. argo, 
patronymics in the genitive, e.g. sismondai, and adjectives with 
indifferent endings, e.g. hians) five feminine and one masculine specific 
names in the 1758—1800 section, where ten names are listed ; and in the 


DIRECTION 42 163 


1801—1850 section, eleven feminine and five masculine out of 26 
specific names. On this basis usage is clearly in favour of a feminine 
gender. On balance, I consider that the correct (masculine) gender 
should be accepted for the name Argonauta. 


(b) Letter from L. Bairstow, M.A., Ph.D. (British Museum (Natural 
History), London), dated 14th January 1955 


I see from a Latin dictionary that nauta -ae and Argonautae -arum 
are masculine ; thus these entries confirm the claim that the correct 
gender of Argonauta is indeed masculine. 


Nevertheless, Argonauta has customarily been treated as feminine. 
Of specific names that have been used in combination with Argonauta, 
those that are adjectives have been employed with feminine terminations 
in the great majority of instances. Most of the comparatively few 
instances of their employment with masculine terminations occurred 
between the years 1790 and 1830. 


However, though the species of Argonauta have been known by many 
different names, the number of well-established species at present 
recognised is not great—perhaps half a dozen Recent species and a few 
fossil ones. Moreover, of the currently accepted names for these 
species, most would not be affected by correction of the gender of 
Argonauta : argo is a noun in apposition ; béttgeri, nowryi, sismondae 
and yoshiwarae are nouns in the genitive; johanneus is already 
masculine in termination; and hians would presumably remain 
unchanged. The only currently important names needing change of 
termination would be cornuta and nodosa, though certain other names 
might need alteration if certain species of uncertain status, or that 
are now considered synonyms, were valued more highly by some future 
reviser. 


I do not see that it need cause much confusion or inconvenience if 
the correct (masculine) gender were to be attributed to Argonauta 
on the Official List. My own view is that the correction should receive 
official endorsement. 


ANNEXE 2 
The gender attributable to the generic name ‘‘ Antedon ’’ Fréminville, 
1811 
Extract from a letter from L. W. Grensted dated 12th July 1955 


Antedon : As this name stands, in Latin, the gender is quite obscure. 
The word as it stands has no direct derivation, but being a name, with 
a Greek look about it, would probably be masculine and might be 


164 OPINIONS AND DECLARATIONS 


feminine. But I suspect that Fréminville wrote Antedon for Anthedon, 
which is found both in Greek and Latin in two meanings, as the name 
of a plant and as a town in Boeotia with mythological associations. In 
both senses Anthedon is feminine, and I suspect that this is the reason 
why Antedon has normally been regarded as feminine. I think this 
usage might stand, in view of this probable derivation—but probably 
it should be confirmed, and the reason given. I do not know whether 
Fréminville did anything to determine either the gender or the 
derivation. 


ANNEXE 3 


Gender proposed to be inscribed in the ‘* Official List of Generic Names 
in Zoology ’’ in respect of the names of seventy-seven genera 
belonging to various Invertebrate Phyla and of the names of 
six genera belonging to the Class Urochorda 


Gender ** Opinion”? in 
- proposed which name 
to be assigned specified in Col. | 
to name speci- was placed on 
Generic Name fied in Col. 1 * Official List 
(1) (2) (3) 
Class Rhizopoda 
Arcella Ehrenberg, [1830] Feminine ae 
Lepidocyclina Giimbel, [1870] Feminine 127 
Class Mastigophora 
Volvox Linnaeus, 1758 Feminine 75 
Trypanosoma Gruby, 1843 Neuter 95 


Class Ciliata 
Bursaria Miller, 1773 Feminine 104 


Class Sporozoa 
Eimeria Schneider, 1875 Feminine 104 


Class Neosporidia 
Sarcocystis Lankester, 1882 Feminine 104 


Class Anthozoa 
Lithostrotion Fleming, 1828 Neuter Ly 


Class Hydrozoa 


Hydra Linnaeus, 1758 Feminine yh 
Phyvsalia Lamarck, 1801 Feminine 80 


DIRECTION 42 165 
Gender 
proposed 
to be assigned 
to name speci- 


** Opinion” in 
which name 
specified in Col. 1 
was placed on 


Generic Name fied in Col. 1 - Ofrcial. List: 
(2) (3) 
Class Trematoda 

Hemiurus Rudolphi, 1809 Masculine eh 
Schistosoma Weinland, 1858 Neuter » 
Dicrocoelium Dujardin, [1844] Neuter 84 
Fasciola Linnaeus, 1758 Feminine fs 
Gastrodiscus Cobbold, 1877 Masculine i. 


Hezterophyes Cobbold, 1866 Masculine 35 


Class Cestoda 


Anoplocephala Blanchard, 1848 Feminine TY 
Hymenolepis Weinland, 1858 Feminine hs 
Moniezia Blanchard, 1891 Feminine 5 
Stilesia Stiles & Hassall, 1893 Feminine ¥ 
Thysanosoma Diesing, 1835 Neuter a 
Davainea Blanchard & Railliet, 1891 Feminine 84 
Dipylidium Leuckart, 1863 Neuter f 
Echinococcus Rudolphi, 1801 Masculine *. 
Taenia Linnaeus, 1758 Feminine at 
Ligula Bloch, 1782 Feminine 104 


Class Nematoda 


Ancylostoma Dubini, 1843 Neuter 66 
Ascaris Linnaeus, 1758 Feminine os 
Dracunculus Reichard, 1759 Masculine - 
Gnathostoma Owen, 1837 Neuter of 
Strongyloides Grassi, 1879 Masculine ef 
Trichostrongylus Looss, 1905 Masculine , 
Filaria Miller, 1787 Feminine 104 
Heterodera Schmidt, 1871 Feminine be 
Rhabditis Dujardin, 1845 Feminine e 
Strongylus Miiller, 1780 Masculine A 
Syngamus Siebold, 1836 Masculine , 


Class Hirudinea 


Hirudo Linnaeus, 1758 Feminine 75 

Haemadipsa Tennent, 1859 Feminine 104 

Limnatis Moquin-Tandon, 1827 Feminine “a 
Class Nematomorpha 

Gordius Linnaeus, 1758 Masculine 66 

Paragordius Camerano, 1897 Masculine he 


166 OPINIONS AND DECLARATIONS 


Gender 
proposed 
to be assigned 
to name speci- 


** Opinion” in 
which name 
specified in Col. 1 
was placed on 


Generic Name fied in Col. 1 ** Official List” 
(1) (2) (3) 
Class Chaetopoda 
Lumbricus Linnaeus, 1758 Masculine 2D 
Enchytraeus Henle, 1837 Masculine 104 
Class Gastropoda 
Buccinum Linnaeus, 1758 Neuter 94 
Calyptraea Lamarck, 1799 Feminine 2 
Columbella Lamarck, 1799 Feminine x 
Helix Linnaeus, 1758 Feminine 
Limax Linnaeus, 1758 Masculine oe 
Physa Draparnaud, [1801] Feminine ts 
Succinea Draparnaud, [1801] Feminine ms 
Leucochila von Martens, 1860 Feminine tr 
Cerion Réding, 1798 Neuter 119 
Oleacina Réding, 1798 Feminine pa 
Neritina Lamarck, 1816 Feminine + 
Clausilia Draparnaud, [1805] Feminine xs 
Vitrina Draparnaud, [1801] Feminine % 
Tornatellina Pfeiffer, 1842 Feminine Fs 
Agasoma Gabb, [1869] Neuter 121 
Class Scaphopoda 
Dentalium Linnaeus, 1758 Neuter 94 
Class Pelecypoda 
Anodonta Lamarck, 1799 Feminine 94 
Mya Linnaeus, 1758 Feminine Ne 
Mytilus Linnaeus, 1758 Masculine ms 
Sphaerium Scopoli, 1777 Neuter " 
Teredo Linnaeus, 1758 Feminine M: 
Mactra Linnaeus, 1767 Feminine * 
Class Cephalopoda 
Argonauta Linnaeus, 1758 Masculine 94 
Sepia Linnaeus, 1758 Feminine Hs 
Lytoceras Suess, 1865 Neuter 130 
Class Brachiopoda 
Spirifer Sowerby, 1816 Masculine 100 
Syringothyris Winchell, 1863 Feminine oz 


DIRECTION 42 167 


Gender ** Opinion ”’ in 
proposed which name 
to be assigned specified in Col. 1 
to name speci- was placed on 


Generic Name fied in Col.1  “ Official List ”’ 
I (2) (3) 
Class Asteroidea 
Luidia Forbes, 1839 Feminine 129 


Class Holothuroidea 
Holothuria Linnaeus, 1767 Feminine 80 


Class Crinoidea 


Antedon Fréminville, 1811 Feminine 73 
Bathycrinus Thomson, 1872 Masculine e 
Holopus d@’Orbigny, 1837 Masculine a 
Metacrinus Carpenter, 1882 Masculine ~ 
Rhizocrinus Sars, [1865] Masculine ¥: 


Class Urochorda 


Pyrosoma Peron, 1804 Neuter vt 
Botryllus Gaertner, 1774 Masculine 94 
Clavelina Savigny, 1816 Feminine es 
Diazona Savigny, 1816 Feminine ef 
Distaplia Della Valle, 1881 Feminine i 
Molgula Forbes, 1848 Feminine es 


2. Registration of the present application : Upon the receipt of 
Mr. Hemming’s paper, the question of the gender to be assigned 
to the names of genera of invertebrates and Urochorda previously 
placed on the Official List of Generic Names in Zoology was 
allotted the Registered Number Z.N.(S.) 991. 


Il. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


3. Issue of Voting Paper V.P.(O.M.)(55)30 : On 29th September 
1955, a Voting Paper (V.P.(O.M.)(55)30) was issued in which 


168 OPINIONS AND DECLARATIONS 


each Member of the Commission was asked (1) to state whether 
he agreed that, “‘in conformity with the General Directive 
relating to the recording in the Official List of Generic Names in 
Zoology of the gender of each name placed thereon prior to 
1948, issued to the International Commission on Zoological 
Nomenclature by the Thirteenth International Congress of 
Zoology, Paris, 1948, the gender of the seventy-eight generic names 
of invertebrates and of the six generic names in the Class 
Urochorda specified in Column (2) of Annexe 3 of the paper 
bearing the Registered Number Z.N.(S.) 991 by the Secretary 
submitted simultaneously with the present Voting Paper [i.e. 
in column (2) of Annexe 3 of the paper reproduced in the first 
paragraph of the present Direction] be entered in the foregoing 
Official List in respect of the names enumerated in the first 
column of the said Annexe ”’ and, if he did not so agree as regards 
any given item, to indicate that item. 


4. The Prescribed Voting Period: As the foregoing Voting 
Paper was issued under the One-Month Rule, the Prescribed 
Voting Period closed on 29th October 1955. 


5. Particulars of the Voting on Voting Paper V.P.(O.M.)(55)30 : 
At the close of the Prescribed Voting Period, the state of the 
voting on Voting Paper V.P.(O.M.)(55)30 was as follows :— 


(a) Affirmative Votes had been given by the following twenty-four 
(24) Commissioners (arranged in the order in which Votes 
were received) : 


Riley ; Boschma; Holthuis; Vokes; Hering; Stoll ; 
Prantl; Lemche; Mayr; Key; do Amaral; Esaki ; 
Mertens ; Miller ; Bradley (J.C.) ; Kiihnelt ; Dymond ; 
Jaczewski ; Hemming; Sylvester-Bradley ; Tortonese ; 
Bonnet ; Cabrera ; Hanko ; 


DIRECTION 42 169 


(b) Negative Votes : 


None ; 


(c) On Leave of Absence, one (1) : 


Bodenheimer ; 


(d) Voting Papers not returned : 


None. 


6. Declaration of Result of Vote: On 2lst November 1955, 
Mr. Hemming, Secretary to the International Commission, acting 
as Returning Officer for the Vote taken on Voting Paper 
V.P.(O.M.)(55)30, signed a Certificate that the Votes cast were as 
set out in paragraph 5 above and declaring that the proposal 
submitted in the foregoing Voting Paper had been duly adopted 
and that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


7. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 24th February 1956 Mr. Hemming prepared the Ruling given 
in the present Direction and at the same time signed a Certificate 
that the terms of that Ruling were in complete accord with those 
of the proposal approved by the International Commission in its 
Vote on Voting Paper V.P.(O.M.)(55)30. 


8. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in 
dealing with the present case, and the present Direction is 
accordingly hereby rendered in the name of the said International 
Commission by the under-signed Francis Hemming, Secretary 
to the International Commission on Zoological Nomenclature, 
in virtue of all and every the powers conferred upon him in that 
behalf. 


170 OPINIONS AND DECLARATIONS 


9. The present Direction shall be known as Direction Forty-Two 
(42) of the International Commission on Zoological Nomenclature. 


Done in London, this Twenty-Fourth day of February, Nineteen 
Hundred and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMM ING 


Note to Subscribers 


The reason why the present Part, containing Direction 42, follows 
immediately after a Part containing Direction 40, is that the inter- 
vening Direction (Direction 41) is concerned with the addition to the 
Official List of Family-Group Names in Zoology of names of that 
category based upon generic names dealt with in Opinions comprised 
in Volume 11 of the present Series. Direction 41 was published in the 
above volume on 17th May 1956 (Ops. Decls. int. Comm. Zool. Nomencl. 
11 : 431—452). 


Printed in England by MrercaLFe & CooprEer LimiTeED. 10-24 Scrutton St.. London E C 2 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, C.M.G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.7. Pp. 171—210 


DIRECTION 43 


Addition to the Official List of Specific Names in Zoology 

(a) of the specific names of the type species of seventy- 

three genera in the Class Aves, the names of which were 

placed on the Official List of Generic Names in Zoology 

in the period up to the end of 1936 and (b) of twelve 

specific names which are currently regarded as senior 
subjective synonyms of such names 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956 


Price Twenty-Six Shillings 
(All rights reserved) 


Issued 19th October, 1956 


f 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 43 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JoRDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President : Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) 
(12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(arranged in order of precedence by reference to date of election or of most recent re-election 
as prescribed by the International Congress of Zoology) 


Professor H. BoscHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(1st January 1947) ; 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr Sar pe waa (Universitetets Zoologiske Museum, Copenhagen, Denmark) (27th 
July 194 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RuiLey (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (Sth July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitdt zu 
Berlin, Germany) (5th July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMonD (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) (12th August 
1953) (President) 

Professor Harold E. VoKEs (Johns Hopkins University, Baltimore, Maryland, U.S.A.) (12th 
August 1953) 

Professor Béla HANK6 (Mezégazdasdgi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. SToL. (Rockefeller Institute for Medical Research, New York, N.Y., 
U.S.A.) (12th August 1953) a 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. HoxttHuis (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (15th October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferninand PRANTL (Ndrodni Museum V Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KiiHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

Professor F. §. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 

4 


) 
Professor Ernst MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 
Professor Enrico TORTONESE (Museo di Storia Naturale ‘“‘ G. Doria”’’, Genova, Italy) 
(16th December 1954) 


DIRECTION 43 


ADDITION TO THE ‘* OFFICIAL LIST OF SPECIFIC 
NAMES IN ZOOLOGY ”’ (a) OF THE SPECIFIC NAMES 
OF THE TYPE SPECIES OF SEVENTY-THREE 
GENERA IN THE CLASS AVES, THE NAMES OF 
WHICH WERE PLACED ON THE “ OFFICIAL 
LIST OF GENERIC NAMES IN ZOOLOGY ”’ 
IN THE PERIOD UP TO THE END OF 1936 
AND (b) OF TWELVE SPECIFIC NAMES 
WHICH ARE CURRENTLY REGARDED 
AS SENIOR SUBJECTIVE SYNONYMS 
OF SUCH NAMES 


RULING:—(1) The under-mentioned specific names 
of species in the Class Aves, each of which is the type 
species of a genus, the name of which was placed 
on the Official List of Generic Names in Zoology in the 
period up to the end of 1936 by the Ruling given in 
Opinion 67, are hereby placed on the Official List of 
Specific Names in Zoology with the Name Numbers 
severally specified against the names in question:— 


7) vulturina Hardwicke, 1834, as published in the com- 
bination Numida vulturina (specific name of type 
species of Acryllium Gray (G.R.), 1840) (Name 
No. 950); 


(2) occidentalis Lawrence, 1858, as published in the 
combination Podiceps occidentalis (specific name 
of type species of Aechmophorus Coues, 1862) 
(Name No. 951); 


(3) maximus Geoffroy, 1850, as published in the com- 
bination Aepyornis maximus (specific name of 
type species of Aepyornis Geoffroy, 1850) (Name 
No. 952); 


WNY | 


A iQtes& 


174 OPINIONS AND DECLARATIONS 


(4) sponsa Linnaeus, 1758, as published in the combina- 
tion Anas sponsa (specific name of type species 
of Aix Boie, 1828) (Name No. 953); 


(5) arvensis Linnaeus, 1758, as published in the com- 
bination Alauda arvensis (specific name of type 
species of Alauda Linnaeus, 1758) (Name No. 
954); 


(6) narina Stephens, 1815, as published in the com- 
bination Trogon narina (specific name of type 
species of Apaloderma Swainson, 1833) (Name 
No, 955); 


(7) patagonica Miller, 1778, as published in the com- 
bination Aptenodytes patagonica (specific name 
of type species of Aptenodytes Miller, 1778) 
(Name No. 956); 


(8) australis Shaw, 1813, as published in the com- 
bination Apteryx australis (specific name of type 
species of Apteryx Shaw, 1813) (Name No. 957); 


(9) scolopacea Gmelin, 1788, as published in the com- 
bination Ardea scolopacea (specific name of type 
species of Aramus Vieillot, 1816) (Name No. 
958); 


(10) cinerea Linnaeus, 1758, as published in the com- 
bination Ardea cinerea (specific name of type 
species of Ardea Linnaeus, 1758) (Name No. 
959); 


(11) nigra Gmelin, 1788, as published in the combina- 
tion Paradisea [sic] nigra (specific name of type 
species of Astrapia Vieillot, 1816) (Name No. 
960); 


(12) sulcatus Swainson, 1820, as published in the com- 
bination Pteroglossus sulcatus (specific name of 
type species of Aulacorhynchus Gould, 1834) 
(Name No. 961); 


DIRECTION 43 175 


(13) auritus Gray (J.E.), 1829, as published in the com- 
bination Podargus auritus (specific name of type 
species of Batrachostomus Gould, 1838) (Name 
No. 962); 


(14) pyrrhopterus Latham, 1801, as published in the 
combination Psittacus pyrrhopterus (specific name 
of type species of Brotogeris Vigors, 1825) (Name 
No. 963); 


(15) moschata Linnaeus, 1758, as published in the 
combination Anas moschata (specific name of 
type species of Cairina Fleming, 1822) (Name 
No. 964); 


(16) niger Miller (P.L.S.), [1776], as published in the 
combination Bucco niger (specific name of type 
species of Capito Vieillot, 1816) (Name No. 965); 


(17) aura Linnaeus, 1758, as published in the combina- 
tion Vultur aura (specific name of type species of 
Cathartes Illiger, 1811) (Name No. 966); 


(18) urophasianus Bonaparte, 1827, as published in the 
combination Tetrao urophasianus (specific name 
of type species of Centrocercus Swainson, 1832) 
(Name No. 967); 


(19) ornatus Geoffroy, 1809, as published in the com- 
bination Cephalopterus ornatus (specific name of 
type species of Cephalopterus Geoffroy, 1809) 
(Name No. 968); 


(20) novae-hollandiae Latham, 1801, as published in the 
combination Cereopsis novae-hollandiae (specific 
name of type species of Cereopsis Latham, 1801) 
(Name No. 969); 


(21) chavaria Linnaeus, 1766, as published in the com- 
bination Parra chavaria (specific name of type 
species of Chauna Illiger, 1811) (Name No. 970); 


176 OPINIONS AND DECLARATIONS 


(22) pictus Linnaeus, 1758, as published in the com- 
bination Phasianus pictus (specific name of type 
species of Chrysolophus Gray (J.E.), [1833—34]) 
(Name No. 971); 


(23) regia Linnaeus, 1758, as published in the combina- 
tion Paradisaea regia (specific name of type 
species of Cicinnurus Vieillot, 1816) (Name No. 
972); 


(24) glandarius Linnaeus, 1758, as published in the com- 
bination Cuculus glandarius (specific name of 
type species of Clamator Kaup, 1829) (Name No. 
973); 


b) 


(25) americanus Linnaeus, 1758, as published in the 
combination Cuculus americanus (specific name 
of type species of Coccyzus Vieillot, 1816) (Name 
No. 974); 


(26) flaveola Linnaeus, 1758, as published in the com- 
bination Certhia flaveola (specific name of type 
species of Coereba Vieillot, 1807) (Name No. 
OID); 


(27) auratus Linnaeus, 1758, as published in the com- 
bination Cuculus auratus (specific name of type 
species of Colaptes Swainson, 1825 (Name No. 
976); 


(28) ani Linnaeus, 1758, as published in the combina- 
tion Crotophaga ani (specific name of type species’ 
of Crotophaga Linnaeus, 1758) (Name No. 977); 


(29) exulans Linnaeus, 1758, as published in the com- 
bination Diomedea exulans (specific name of type 
species of Diomedea Linnaeus, 1758) (Name No. 
978); 


(30) ardeola Paykull, 1805, as published in the combina- 
tion Dromas ardeola (specific name of type species 
of Dromas Paykull, 1805) (Name No. 979); 


DIRECTION 43 Pay 


(31) garzetta Linnaeus, 1766, as published in the com- 
bination Ardea garzetta (specific name of type 
species of Egretta Forster, 1817) (Name No. 980); 


(32) javanicus Horsfield, 1821, as published in the com- 
bination Eurylaimus javanicus (specific name of 
type species of Eurylaimus Horsfield, 1821) 
(Name No. 981); 


(33) helias Pallas, 1781, as published in the combina- 
tion Ardea helias (specific name of type species of 
Eurypyga Illiger, 1811) (Name No. 982); 


(34) glacialis Linnaeus, 1761, as published in the com- 
bination Procellaria glacialis (specific name of 
type species of Fulmarus Stephens, 1826) (Name 
No. 983); 


(35) swainsonii Vigors, 1825, as published in the com- 
bination Gampsonyx swainsonii (specific name of 
type species of Gampsonyx Vigors, 1825) (Name 
No. 984); 


(36) ostralegus Linnaeus, 1758, as published in the com- 
bination Haematopus ostralegus (specific name 
of type species of Haematopus Linnaeus, 1758) 
(Name No. 985); 


(37) struthersii Vigors, [1832], as published in the com- 
bination /bidorhyncha struthersii (specific name 
of type species of Ibidorhyncha Vigors, [1832]) 
(Name No. 986); 


(38) torquilla Linnaeus, 1758, as published in the com- 
bination Jynx torquilla (specific name of type 
species of Jynx Linnaeus, 1758) (Name No. 987); 


(39) excubitor Linnaeus, 1758, as published in the com- 
bination Lanius excubitor (specific name of type 
Be). of Lanius Linnaeus, 1758) (Name No. 


178 OPINIONS AND DECLARATIONS 


(40) violacea Isert, 1789, as published in the combination 
Musophaga violacea (specific name of type species 
of Musophaga Isert, 1789) (Name No. 989); 


(41) mantellii Owen, 1848, as published in the combina- 
tion Notornis mantellii (specific name of type 
species of Notornis Owen, 1848) (Name No. 990); 


(42) meleagris Linnaeus, 1758, as published in the com- 
bination Phasianus meleagris (specific name of 
type species of Numida Linnaeus, 1764) (Name 
No. 991); 


(43) pectoralis Latham, 1801, as published in the com- 
bination Muscicapa pectoralis (specific name 
of type species of Pachycephala Vigors, 1825) 
(Name No. 992); 


(44) sefilata Forster, 1781, as published in the com- 
bination Paradisea [sic] sefilata (specific name of 
type species of Parotia Vieillot, 1816) (Name No. 
093); 


? 


(45) major Linnaeus, 1758, as published in the com- 
bination Parus major (specific name of type 
species of Parus Linnaeus, 1758) (Name No. 994); 


(46) aethereus Linnaeus, 1758, as published in the com- 
bination Phaethon aethereus (specific name of type 
species of Phaethon Linnaeus, 1758) (Name No. 
Doak 


(47) mocinno de la Llave, 1832, as published in the com- 
bination Pharomachrus mocinno (specific name 
of type species of Pharomachrus de la Llave, 1832) 
(Name No. 996); 


(48) ruber Linnaeus, 1758, as published in the combina- 
tion Phoenicopterus ruber (specific name of type 
species of Phoenicopterus Linnaeus, 1758) (Name 
No. 997); 


DIRECTION 43 179 


(49) leucorodia Linnaéus, 1758, as published in the com- 
bination Platalea leucorodia (specific name of type 
species of Platalea Linnaeus, 1758) (Name No. 
998); 


(50) porzana Linnaeus, 1766, as published in the com- 
bination Rallus porzana (specific name of type 
species of Porzana Vieillot, 1816) (Name No. 
999); 


(51) erithacus Linnaeus, 1758, as published in the com- 
bination Psittacus erithacus (specific name of 
type species of Psittacus Linnaeus, 1758) (Name 
No. 1000); 


(52) crepitans Linnaeus, 1758, as published in the com- 
bination Psophia crepitans (specific name of type 
species of Psophia Linnaeus, 1758) (Name No. 
1001); 


(53) aracari Linnaeus, 1758, as published in the com- 
bination Ramphastos aracari (specific name of 
type species of Pteroglossus Illiger, 1811) (Name 
No. 1002); 


(54) paradiseus Swainson, 1825, as published in the 
combination Ptiloris paradiseus (specific name of 
type species of Ptiloris Swainson, 1825) (Name 
No. 1003); 


(55) aquaticus Linnaeus, 1758, as published in the com- 
bination Rallus aquaticus (specific name of type 
species of Rallus Linnaeus, 1758) (Name No. 
1004); 


(56) avosetta Linnaeus, 1758, as published in the com- 
bination Recurvirostra avosetta (specific name of 
type species of Recurvirostra Linnaeus, 1758) 
(Name No. 1005); 


180 OPINIONS AND DECLARATIONS 


(57) chrysocephala Lewin, 1808, as published in the 
combination Meliphaga chrysocephala (specific 
name of type species of Sericulus Swainson, 1825) 
(Name No. 1006); 


(58) europaea Linnaeus, 1758, as published in the com- 
bination Sitta europeaea (specific name of type 
species of Sitta Linnaeus, 1758) (Name No. 
1007); 


(59) abdimii Lichtenstein, 1823, as published in the 
combination Ciconia abdimii (specific name of 
type species of Sphenorynchus Lichtenstein, 1823) 
(Name No. 1008); 


(60) habroptilus Gray (G.R.), 1845, as published in the 
combination Strigops habroptilus (specific name 
of type species of Strigops Gray (G.R.), 1845) 
(Name No. 1009); 


(61) magna Linnaeus, 1758, as published in the com- 
bination Alauda magna (specific name of type 
species of Sturnella Vieillot, 1816) (Name No. 
1010); 


(62) vulgaris Linnaeus, 1758, as published in the com- 
bination Sturnus vulgaris (specific name of type 
species of Sturnus Linnaeus, 1758) (Name No. 
1011); 


(63) paradoxa Pallas, 1773, as published in the com-. 
bination Tetrao paradoxa (specific name of type 
species of Syrrhaptes Uliger, 1811) (Name No. — 
1012); | 


(64) rufa Boddaert, 1783, as published in the combina- 
tion Tanagra rufa (specific name of type species 
of Tachyphonus Vieillot, 1816) (Name No. 1013); 


(65) doliatus Linnaeus, 1764,.as published in the com- 
bination Lanius doliatus (specific name of type 
species of T; bai Vieillot, 1816) (Name No. 
1014); 


DIRECTION 43 181 


(66) chimaera Rothschild, 1895, as published in the 
combination Uratelornis chimaera (specific name 
of type species of Uratelornis Rothschild, 1895) 
(Name No. 1015). 


(2) It is hereby ruled that under the provisions of 
Declaration 21 the type species of each of the under- 
mentioned nominal genera is the nominal species specified 
below, that nominal species being objectively identical 
with the later-established nominal species cited by the 
author of the generic name concerned :— 


(1) The type species of Coturnix Bonnaterre, 1790, is 
Tetrao coturnix Linnaeus, 1758, and not the 
later-established nominal species Coturnix com- 
munis Bonnaterre, 1791. 


(2) The type of Eurynorhynchus Nilsson, 1821, is 
Platalea pygmaea Linnaeus, 1758, and not the 
later-established nominal species Eurynorhynchus 
griseus Nilsson, 1821. 


(3) The type species of Heliornis Bonnaterre, 1790, is 
Colymbus fulica Boddaert, 1783, and not the 
later-established nominal species Heliornis fuli- 
carius Bonnaterre, 1790. 


(4) The type species of Manucodia Boddaert, 1783, is 

: Paradisea [sic] chalybata Forster, 1781, and not 

the later-established nominal species Manucodia 
chalybea Boddert, 1783. 


(5) The type species of Oriolus Linnaeus, 1766, is 
Coracias oriolus Linnaeus, 1758, and not the 
later-established nominal species Oriolus galbula 
Linnaeus, 1766. 


(6) The type species of Pandion Savigny, 1809, is Falco 
haliaetus Linnaeus, 1758, and not the later- 
established nominal species Pandion fluvialis 
Savigny, 1809. 


182 OPINIONS AND DECLARATIONS 


(7) The type species of Polyplectron Temminck, 1807, 
is Pavo bicalcaratus Linnaeus, 1758, and not 
the later-established nominal species Polyplectron 
argus Temminck, 1807. 


(3) The under-mentioned specific names, each of which 
is the name of a nominal species which, as specified in (2) 
above, is, under Declaration 21, the type species of a 
genus in the Class Aves, the name of which was placed 
on the Official List of Generic Names in Zoology in the 
period up to the end of 1936 by the Ruling given in 
Opinion 67, are hereby placed on the Official List of 
Specific Names in Zoology with the Name Numbers 
severally specified against the names in question:— 


(1) coturnix Linnaeus, 1758, as published in the com- 
bination Tetrao coturnix (specific name of type 
species of Coturnix Bonnaterre, 1790) (Name No. 
1016); 


(2) pygmaea Linnaeus, 1758, as published in the com- 
bination Platalea pygmaea (specific name of type 
species of Eurynorhynchus Nilsson, 1821) (Name 
No. 1017); 


(3) fulica Boddaert, 1783, as published in the com- 
bination Colymbus fulica (specific name of 
type species of Heliornis Bonnaterre, 1790) (Name 
No. 1018); 


(4) chalybata Forster, 1781, as published in the com- 
bination Paradisea [sic] chalybata (specific name 
of type species of Manucodia Boddaert, 1783) 
(Name No. 1019); 


(5) oriolus Linnaeus, 1758, as published in the com- 
bination Coracias oriolus (specific name of type 
species of Oriolus Linnaeus, 1766) (Name No. 
1020); 


(6) haliaetus Linnaeus, 1758, as published in the com- 
bination Falco haliaetus (specific name of type 
species of Pandion Savigny, 1809) (Name No. 
1021); 


DIRECTION 43 183 


(7) bicalcaratus Linnaeus, 1758, as published in the 
combination Pavo bicalcaratus (specific name of 
type species of Polyplectron Temminck, 1807) 
(Name No. 1022). 


(4) The under-mentioned specific names are hereby 
placed on the Official List of Specific Names in Zoology 
with the Name Numbers severally specified below:— 


(1) tiphia Linnaeus, 1758, as published in the combina- 
tion Motacilla tiphia (Name No. 1023); 


(2) cristatus Shaw in White, [August] 1790, as published 
in the combination Caprimulgus cristatus (Name 
No. 1024); 


(3) nitidus Latham, 1790, as published in the combina- 
tion Falco nitidus (Name No. 1025); 


(4) caeruleus Desfontaines, [1789], as published in the 
combination Falco caeruleus (Name No. 1026); 


(5) cristata Pallas, 1764, as published in the combina- 
tion Columba cristata (Name No. 1027); 


(6) albicilla Linnaeus, 1758, as published in the com- 
bination Falco albicilla (Name No. 1028); 


(7) indus Boddaert, 1783, as published in the com- 
bination Falco indus (Name No. 1029); 


(8) militaris Linnaeus, 1758, as published in the com- 
bination Emberiza militaris (Name No. 1030); 


(9) hoazin Miiller (P.L.S.), [1776], as published in the 
combination Phasianus hoazin (Name No. 1031); 


(10) wallicus Kerr, 1792, as published in the combina- 
tion Psittacus wallicus (Name No. 1032); 


184 OPINIONS AND: DECLARATIONS 


(11) elegans Gmelin (J.F.), 1788, as published in the 
combination Psittacus elegans (Name No. 1033); 


(12) nigricephala Jameson, 1835, as published in the com- 
bination Tanagra nigricephala (Name No. 1034). 


I THE SUBJECT MATTER OF THE PRESENT 
‘DIRECTION ” 


The present Direction has been prepared under a General 
Directive issued to the International Commission on Zoological 
Nomenclature by the Thirteenth International Congress of 
Zoology, Paris, 1948, that the specific name of the type species 
of each genus, the name of which was placed on the Official 
List of Generic Names in Zoology in the period up to the close of 
1936, be inscribed on the Official List of Specific Names in Zoology 
except where that name is not the oldest available name for the 
species concerned, in which case the oldest available name, 
whatever it may be, shall be so inscribed. The present Direction, 
which is the third dealing with this group of cases, is concerned 
with the specific names of the type genera of birds. The total 
number of such names placed on the Official List during the 
foregoing period amounted to one hundred and three. Of these 
the specific names of the type species of four genera (Campephaga ; 
Colluricincla ; Neophron; Struthio)' have been placed on the 
Official List in Opinions in which these names arose incidentally, 
the opportunity so presented being taken to comply with the 
General Directive referred to above. In five cases (Balaeniceps ; 
Burhinus ; Ectopistes ; Gallinago ; Oedicnemus)' it was found 
necessary to submit separate proposals to the Commission and 
in the Directions rendered by the Commission in these cases the 
specific names of the type species of the genera concerned were 
placed on the Official List. In addition there are cases pending 
regarding the following nine generic names placed on the Official 
List in the period up to the end of 1936: Anas ; Bubo ; Circaetus ; 
Grus! ; Gypaetus ; Nyctea; Surnia ; Trichoglossus ; Vireo)’. In 
these cases also it has been considered that the question of the 
i | ee) ee 


‘1 Particulars of the action already taken or now in hand in regard to these 
generic names are given in the Appendix attached to the present Direction. 


DIRECTION 43 185 


action to be taken to comply with the General Directive here under 
discussion can most conveniently be deferred until decisions 
are taken by the Commission in regard to the other problems. 
arising in connection with these names. When account is taken of 
the foregoing exceptions there remain eighty-five names which 
require to be considered in connection with the General Directive 
issued by the Paris Congress. An examination of these cases 
shows that in sixty-six cases the name cited for the type species 
of the genus concerned by the original author of the generic 
name is the oldest available name for that species. In seven other 
cases the name cited by the author of the generic name is a junior 
objective synonym of another name and in these cases it is the 
senior objective synonym which under Declaration 21 becomes 
the name of the type species and the name which, as the oldest 
available name, requires to be placed on the Official List of 
Specific Names in Zoology. In the remaining twelve cases the 
specific name of the type species of the genus concerned is a 
junior subjective synonym of some older name. In these cases 
it is the specific name which is the senior subjective synonym 
and not the specific name of the type species itself which requires 
to be placed on the Official List. The decisions taken by the 
Commission in regard to each of the foregoing classes of case 
are dealt with in the present Direction. 


2. The proposals which formed the basis of the decisions 
embodied in the present Direction were submitted to the 
Commission by the pecretanys in | the folllowing paper on 12th 
October 1955 :— 


Proposed addition to the ‘‘ Official List of Specific Names in Zoology ’* 
of the specific names of the type species of seventy-five genera in the 
Class Aves, the names of which were placed on the ‘‘ Official 
List of Generic Names in Zoology ’’ in the period up to 
the end of 1936 and of twelve specific names of which 
are currently treated as senior subjective synonyms 
of the names of type species of such genera 


By FRANCIS HEMMING, C.M.G., C.B.E. 
(Secretary to the International Commission on Zoological Nomenclature) 


The purpose of the present paper is to lay recommendations before 
the International Commission to enable it to comply, so far as concerns 
names of species in the Class Aves, with the General Directive issued 


186 OPINIONS AND DECLARATIONS 


to it by the Thirteenth International Congress of Zoology, Paris, 
1948. By this Directive the Commission is required to place on the 
Official List of Specific Names in Zoology the specific name of the 
type species of every genus, the name of which had been placed on the 
Official List of Generic Names in Zoology in the period up to the end 
of 1936, except where that name is not the oldest available name for 
the taxon concerned, in which case the oldest available name, whatever 
it may be, is to be placed on that Official List. The present is the third 
instalment of proposals submitted in compliance with the foregoing 
Directive, the previous instalments being concerned with the names 
of mammals and Decapod Crustacea respectively.? 


2. In compiling the present proposals, I have, as on the previous 
occasions referred to above, consulted modern catalogues, checklists 
and monographs for the purpose of ascertaining the present practice of 
specialists in regard to the names which have now to be considered. 
Fortunately, this task was less difficult that in some of the other 
groups which it has been necessary to consider, for, owing to the 
popularity of ornithology, the number of modern and reliable works of 
the kind required is relatively large. The bulk of the work involved 
in the preparation of the proposals now submitted was carried out a 
number of years ago and | was fortunate in being able to draw for 
advice on the wide knowledge and sound judgement of my friend and 
colleague the late Dr. James L. Peters. In a number of cases which we 
had discussed together in this way I later received most valuable 
confirmatory advice from Colonel Richard Meinertzhagen, at that 
time the Chairman of the Standing Committee on Ornithological 
Nomenclature established in 1950 by the International Congress of 
Ornithology. 


3. The total number of names of genera of birds placed on the 
Official List of Generic Names in Zoology in the period up to the end 
of 1936 was one hundred and three. As regards five of these names it 
has been found necessary for one reason or another to seek further 
directions from the Commission, and in each case the opportunity was 
taken to place the name of the type species of the genus concerned on 
the Official List of Specific Names in Zoology. The names were: 
(i) Ectopistes (Bull. zool. Nomencl. 9 : 80—84) ; (ii) Gallinago (Bull. — 
9 : 93); (ii) Balaeniceps (Bull. 9 : 85—86) ; (iv) and (v) Oedicnemus 
and Burhinus (Bull. 9 : 86—88). In the case of Ectopistes and 
Balaeniceps, the decisions by the Commission have already been 


2 The proposals here referred to were submitted to the Commission with Voting 
Papers V.P.(O.M.)(55)9 and V.P.(O.M.)(55)27 respectively. Decisions regarding 
these proposals have since been taken by the Commission and embodied in 
Direction 22 in the case of V.P.(O.M.)(55)9 and in Direction 36 in the case of 
V.P.(O.M.)(55)27. 


DIRECTION 43 187 


announced (in Directions 18 and 19 respectively), and decisions on the 
remaining names will be announced at an early date.? In three 
additional cases, applications submitted to the Commission included 
proposals for the addition to the Official List of Specific Names in 
Zoology of the specific names of the type species of genera, the names 
of which were placed on the Official List of Generic Names in Zoology 
in the period under review. The names concerned were: (a) 
Campephaga (Bull. 9 : 32—33); (b) Colluricincla (Bull. 9 : 50—S1) ; 
(c) Struthio (Bull. 9 : 96—97).3 Altogether, therefore, the question 
of the addition to the Official List of the specific names of the type 
species of the genera with which we are here concerned is being dealt 
with separately in eight cases. Thus, the number of names to be 
considered on the present occasion is reduced to ninety-five. 


4. We have first to note that as regards the names of the type species 
of eight of these genera it is not possible at the moment to submit 
recommendations to the Commission. The difficulties involved arise 
from a diversity of causes: (1) and (2) In the case of the nominal 
genera Bubo and Surnia it is not clear what was the first occasion 
on which nominal species were cited in connection with them; (3) 
Gypaetus is commonly stated to be monotypical but it is doubtful 
whether or not this is correct; (4) In the case of Trichoglossus the 
nominal species currently accepted as the type species was not included 
in the genus at the time when this name was first published ; (5) The 
name Grus is commonly attributed to the wrong author and date.® 
Moreover, as from the date from which this name is currently accepted 
the species treated as the type species was not an included species ; 
(6) In the case of Anas there is an unresolved doubt as to what is the 
correct name for its type species ; (7) The next name is Nyctea where 
it has been impossible so far to trace where in the literature a first 
reviser selected the name currently used for the type species in preference 
to another name published in the same book; (8) In the case of 
Vireo it has not yet been found possible to trace a valid selection for 
its type species, the currently accepted selection being invalid, as it was 
the selection of a nominal species which, though currently identified 
subjectively with one of the included species, was not itself an included 
species. In all these cases investigations are proceeding in conjunction 
with interested specialists and proposals will be submitted to the 
Commission as soon as the factual position in regard to these names 
has been established. The postponement of these cases reduces the 
number of names with which it is possible to deal on the present 
occasion to eighty-seven.* 


5. In sixty-eight cases the specific name of the type species of the 
genus concerned is not only an objectively available name but is also 


3 Particulars of the action since taken in regard to these names are given in the 
Appendix attached to the present Direction. 

4 The proposals submitted in regard to two names were later withdrawn by the 
Minute reproduced in paragraph 5 of the present Direction. It is for this 
reason that the number of names dealt with in the present Direction amounts 
only to eighty-five. 


188 OPINIONS AND DECLARATIONS 


the name which is accepted by all authorities as the oldest available 
name for the species concerned. These names are enumerated in 
Annexe 1. It is recommended that, in compliance with the General 
Directive relating to the maintenance of the Official Lists issued to 
the Commission by the International Congress of Zoology, these names 
be now placed on the Official List of Specific Names in Zoology. 


6. In the application submitted to the Commission which later 
became the basis of the Ruling given in its Opinion 67, the applicants 
added indications in a number of cases making it clear that, in their 
opinion, the specific name of the type species was a junior subjective 
synonym of some other species. In most of the cases concerned the 
synonymies so established have stood the test of time and are currently 
accepted by ornithologists. In two cases, however, the synonymies 
established by the applicants are no longer considered to be correct. 
The names concerned are: (1) Elanus Savigny, 1809: The type 
species of this genus is the nominal species Elanus caesius Savigny, 
1809. In Opinion 67 the name caesius was treated as a junior subjective 
synonym of melanopterus Daudin (Falco melanopterus Daudin, 1800). 
Currently, however, it is sunk as a junior subjective synonym of the 
older name caeruleus Desfontaines (Falco caeruleus Desfontaines, 
1789). (ai) Pezoporus Illiger, 1811: The type species of this genus is 
Psittacus formosus Latham, 1790, which is invalid, as being a junior 
homonym of Psittacus formosus Scopoli, 1769. In Opinion 67 the 
oldest name subjectively available for this species was stated to be 
Psittacus terrestris Shaw, 1793. It is now considered that the oldest 
available name is Psittacus wallicus Kerr, 1792. 


7. In twelve cases the specific name of the type species concerned is 
an objectively available name but is regarded by all authorities as being 
a junior subjective synonym of some other name. Particulars of these 
cases are given in Annexe 2. It is recommended that, in compliance 
with the General Directive relating to the maintenance of the Official 
Lists issued to the Commission by the International Congress of 
Zoology, the twelve senior subjective synonyms concerned be placed 
on the Official List of Specific Names in Zoology. 


8. A different problem is involved with the last seven of the names 
to be considered. In each of these cases the specific name used for 
the type species by the original author of the generic name is a junior 
objective (and therefore invalid) synonym of the name of a previously 
established nominal species. In consequence the nominal species to be 
accepted as the type species of each of these genera will depend upon 
the decision to be taken by the Commission on the proposal 
Z.N.(S.) 908, now before it, for the adoption of a Declaration pres- 
cribing the nominal species to be accepted as the type species of a 
genus in such a case (Hemming, 1955, Bull. zool. Nomencl. 11 : 112— 
113).5 Under the proposals so submitted, the nominal species to be 


5 The application here referred to has since been approved by the International 
Commission and has been embodied in Declaration 21 (1956, Ops. Decls. int. 
Comm. zool. Nomencl. 12 : i—viii). 


DIRECTION 43 189 


treated as the type species of the genus concerned would be not the 
species cited by the author of the generic name but the older nominal 
species which is objectively identical with it. The situation discussed 
above arises only where the nominal species, which, under the present 
rules, is the type species of a genus, was established as a direct sub- 
stitute for a previously established nominal species and where, in 
consequence, the two nominal species are objectively identical with 
one another, each having the same specimen as its type specimen. 
This situation was illustrated in the application referred to above by 
the case of Homarus Weber, 1795, the type species of which is, under the 
existing rules, Astacus marinus Fabricius, 1775. When publishing 
this name, Fabricius did not, however, describe a new species, but 
deliberately gave a new name for (nom. nov. pro) the nominal species 
Cancer gammarus Linnaeus, 1758. Thus, the nominal species bearing 
the names A. marinus Fabricius and C. gammarus Linnaeus are 
objectively identical with one another, the specific name gammarus 
Linnaeus being the valid name for the taxon concerned and marinus 
Fabricius being no more than an invalid (because junior) objective 
synonym of gammarus Linnaeus. The object of the suggested 
Declaration® referred to above is to weed out these junior objective 
synonyms in favour of the corresponding senior objective synonyms, 
thus making it possible to cite as the type species of the genus con- 
cerned the nominal species bearing the name which is the valid (senior) 
synonym (in the case cited above, the well-known name gammarus 
Linnaeus) instead of the invalid (junior) objective synonym (in the case 
cited above, the invalid name marinus Fabricius, which is totally 
unknown, except to specialists familiar with the old literature). 


9. The seven names here involved are the following: Coturnix 
Bonnaterre, 1790; Eurynorhynchus Nilsson, 1821 ; Heliornis Bonna- 
terre, 1790; Manucodia Boddaert, 1783; Oriolus Linnaeus, 1766 ; 
Pandion Savigny, 1809; Polyplectron Temminck, 1807. Full par- 
ticulars of these cases are given in Section A of Annexe 3. In dealing 
with these names for the purpose of preparing the present application, 
it has been necessary in each case to make a working assumption on the 
question as to which of the two objectively identical nominal species 
should be treated as the type species of the genus concerned. The 
assumption which has been adopted is that the Commission will 
approve the adoption of the Declaration discussed above’ and therefore 
that, for these seven genera, the type species will be the nominal 
species bearing the older of the two objectively synonymous names. 
In each case the senior objective synonym is an available name and is 
regarded by specialists as the oldest such name for the species con- 
cerned. The names concerned are shown in Column (2) of Section B 
of Annexe 3. In the same Section of that Annexe there are given in 
Column (3) the names which are junior objective synonyms of the 
names shown in Column (2). Ifthe Declaration discussed in paragraph 
8 above is adopted by the Commission, the names shown in Column 


6 See footnote 5. 


190 OPINIONS AND DECLARATIONS 


(2) will be the names of the type species of the genera concerned, while 
if the Commission decides against that course, the names shown in 
Column (3) will be the names of the type species of the genera con- 
cerned. Irrespective of the action to be taken by the Commission in 
the matter of the above Declaration,’ the senior synonyms referred to 
above will, under the General Directive mentioned in paragraphs 
5 and 7, need to be placed on the Official List of Specific Names in 
Zoology. If the above Declaration is adopted,’ these names will need 
to be associated with the names enumerated in Annexe 1 (specific 
names of type species), while, if that Declaration is not adopted, these 
names will need to be associated with the names enumerated in Annexe 
2 (specific names of nominal species either subjectively or objectively 
identified with later established nominal species which are type species). 


10. In the light of the considerations advanced in the preceding 
paragraphs, I recommend that, in compliance with the General 
Directive relating to the maintenance of the Official Lists issued by 
the Thirteenth International Congress of Zoology, the Commission 
should :— 


(a) place on the Official List of Specific Names in Zoology :— 


(i) the sixty-eight specific names enumerated in Col. (1) of 
Annexe 1, each of which is the type species of the genus 
shown in the corresponding entry in Col. (3) of the said 
Annexe (paragraph 5) ; 

(ii) the twelve specific names enumerated in Col. (1) of Annexe 
2, each of which is the specific name of a nominal species 
which is currently subjectively identified with the later 
established nominal species which is the type species of 
the genus specified in the corresponding entry in Col. (2) 
of the said Annexe (paragraph 7) ; 


(b) place on the foregoing Official List the seven specific names 
enumerated in Section C of Annexe 3, these names to be so 
placed :— 


(i) as the names of the type species of the nominal genera 
specified in the said Section, in the event of the prior 
adoption by the Commission of the Declaration recom- 
mended in Application Z.N.(S.) 908’ ; 

(ii) as the names of nominal species which are objectively 
identical with the later established nominal species which 
are severally the type species of the genera referred to 
in (i) above, in the event of the non-adoption by the 
Commission of the Declaration referred to above. 


en ee ee 


7 For particulars of the action subsequently taken by the International Com- 
mission in this matter see Footnote 5. 


DIRECTION 43 


ANNEXE 1 


191 


Proposed addition to the ‘‘ Official List of Specific Names in Zoology ”’ 
of the specific names of the type species of sixty-eight nominal 
genera, the names of which were placed on the ‘* Official 
List of Generic Names in Zoology ”’ in the period up to 


Specific name 
proposed to be placed 
on the “* Official List 
of Specific Names in 

Zoology” 


(1) 


vulturina Hardwicke, 
1834 


occidentalis 
Lawrence, 1858 


maximus Geoffroy, 
1850 


sponsa Linnaeus, 
1758 


arvensis Linnaeus, 
1758 


narina Stephens, 1815 


patagonica Miller, 
1778 


australis Shaw, 1813 


scolopacea Gmelin, 
1788 


cinerea Linnaeus, 
1758 


nigra Gmelin, 1788 

sulcatus Swainson, 
1820 

auritus Gray, 1829 

pyrrhopterus Latham, 
1801 


moschata Linnaeus, 
1758 


the end of 1936 


Combination in which 
name specified in 
Col. (1) was originally 
published 


(2) 
Opinion 67 
Numida vulturina 


Podiceps 
occidentalis 


Aepyornis maximus 
Anas sponsa 
Alauda arvensis 
Trogon narina 
Aptenodytes 
patagonica 


Apteryx australis 
Ardea scolopacea 


Ardea cinerea 
Paradisea [sic] nigra 
Pteroglossus 

sulcatus 
Podargus auritus 
Psittacus 


pyrrhopterus 
Anas moschaia 


Name of genus of 
which species 
specified in Col. (1) 
is the type species 


(3) 


Acryllium Gray, 1840 


Aechmophorus Coues, 
1862 


Aepyornis Geoffroy, 
1850 


Aix Boie, 1828 


Alauda Linnaeus, 
1758 


Apaloderma Swainson, 
1833 


Aptenodytes Miller, 
1778 


Apteryx Shaw, 1813 
Aramus Vieillot, 1816 


Ardea Linnaeus, 
1758 


Astrapia Vieillot, 
1816 


Aulacorhynchus 
Gould, 1834 


Batrachostomus 
Gould, 1838 


Brotogeris Vigors, 
1825 


Cairina Fleming, 
1822 


192 


Specific name 
proposed to be placed 
on the “* Official List 
of Specific Names in 

Zoology” 


(1) 


niger Miiller (P.L.S.), 
[1776] 


aura Linnaeus, 1758 


urophasianus 
Bonaparte, 1827 


ornatus Geoffroy, 
1809 


novaehollandiae 
Latham, 1801 


chavaria Linnaeus, 
1766 


pictus Linnaeus, 1758 
regia Linnaeus, 1758 
gallicus Gmelin, 1788 


glandarius Linnaeus, 
1758 


americanus Linnaeus, 
1758 


flaveola Linnaeus, 
1758 


auratus Linnaeus, 
1758 


ani Linnaeus, 1758 


exulans Linnaeus, 
1758 


ardeola Paykull, 1805 


garzetta Linnaeus, 
1766 


Combination in which 


name specified in 


Col. (1) was originally 


published 


(2) 


Bucco niger 
Vultur aura 
Tetrao 


urophasianus 


Cephalopterus 
ornatus 


Cereopsis 
novae-hollandiae 
Parra chavaria 
Phasianus pictus 
Paradisaea regia 
Falco gallicus 
Cuculus glandarius 
Cuculus americanus 
Certhia flaveola 
Cuculus auratus 
Crotophaga ani 
Diomedea exulans 


Dromas ardeola 


Ardea garzetta 


OPINIONS AND DECLARATIONS 


Name of genus of 
which species 
specified in Col. (1) 
is the type species 


(3) 
Capito Vieillot, 1816 


Cathartes Illiger, 
811 


Centrocercus 
Swainson, 1832 


Cephalopterus 
Geoffroy, 1809 


Cereopsis Latham, 
1801 


Chauna Illiger, 1811 


Chrysolophus Gray 
(J.E.), [1833-34] 


Cicinnurus Vieillot, 
1816 


Circaetus Vieillot, 
1816 


Clamator Kaup, 1829 


Coccyzus Vieillot, 
1816 


Coereba Vieillot, 
1807 


Colaptes Swainson, 
1825 


Crotophaga 
Linnaeus, 1758 


Diomedea Linnaeus, 
1758 


Dromas Paykull, 
1805 


Egretta Forster, 1817 


Specific name 
proposed to be placed 
on the “‘ Official List 
of Specific Names in 

Zoology” 


(1) 
javanicus Horsfield, 
1821 


helias Pallas, 1781 


glacialis Linnaeus, 
1761 


swainsonii Vigors, 
1825 


ostralegus Linnaeus, 
1758 


struthersii Vigors, 
[1832] 


torquilla Linnaeus, 
1758 


excubitor Linnaeus, 
1758 


violacea Isert, 1789 


percnopterus 
Linnaeus, 1758 


mantellii Owen, 1848 


meleagris Linnaeus, 
1758 


pectoralis Latham, 
1801 


sefilata Forster, 1781 


major Linnaeus, 1758 


aethereus Linnaeus, 
1758 


mocinno de la Llave, 
1832 


ruber Linnaeus, 1758 


DIRECTION 43 


Combination in which 
name specified in 
Col. (1) was originally 
published 


(2) 
Eurylaimus 


jJavanicus 
Ardea helias 


Procellaria glacialis 


Gampsonyx 
SWainsonil 


Haematopus 
ostralegus 


Ibidorhyncha 
struthersii 


Jynx torquilla 
Lanius excubitor 
Musophaga violacea 
Vultur perenopterus 


Notornis mantellii 
Phasianus meleagris 


Muscicapa 
pectoralis 


Paradisea [sic] 
sefilata 


Parus major 
Phaethon aethereus 


Pharomachrus 
mocinno. 


Phoenicopterus 
ruber 


193 


Name of genus of 
which species 
specified in Col. (1) 
is the type species 


(3) 


Eurylaimus 
Horsfield, 1821 


Eurypyga Illiger, 
1811 


Fulmarus Stephens, 
Gampsonyx Vigors, 


Haematopus 
Linnaeus, 1758 


Ibidorhyncha Vigors, 
[1832] 


Jynx Linnaeus, 1758 


Lanius Linnaeus, 
1758 


Musophaga Isert, 
1789 


Neophron Savigny, 
1809 


Notornis Owen, 1848 


Numida Linnaeus, 
1764 


Pachycephala 
Vigors, 1825 


Parotia Vieillot, 1816 


Parus Linnaeus, 1758 


Phaethon Linnaeus, 
1758 


Pharomachrus ; 
de la Llave, 1832 


Phoenicopterus 
Linnaeus, 1758 


194 


Specific name 


proposed to be placed 
on the ‘‘ Official List Col. (1) was originally 
of Specific Names in 


Zoology” 
(1) 


leucorodia Linnaeus, 
1758 


porzana Linnaeus, 
1766 


erithacus Linnaeus, 
1758 


crepitans Linnaeus, 
1758 


aracari Linnaeus, 
1758 


paradiseus Swainson, 
1825 


aquaticus Linnaeus, 
1758 


avosetta Linnaeus, 
1758 


chrysocephala Lewin, 
1808 


europaea Linnaeus, 
1758 


abdimii Lichtenstein, 
1823 


habroptilus Gray, 
1845 


magna Linnaeus, 
1758 


vulgaris Linnaeus, 
1758 


paradoxa Pallas, 
77S 


rufa Boddaert, 1783 


doliatus Linnaeus, 
1764 


chimaera 
Rothschild, 1895 


Combination in which 
name specified in 


published 


(2) 


Platalea leucorodia 
Rallus porzana 
Psittacus erithacus 
Psophia crepitans 
Ramphastos aracari 
Ptiloris 

paradiseus 
Rallus aquaticus 
Recurvirostra 


avosetta 
Meliphaga 


chrysocephala 
Sitta europaea 
Ciconia abdimii 
Strigops 

habroptilus 
Alauda magna 
Sturnus vulgaris 
Tetrao paradoxa 
Tanagra rufa 


Lanius doliatus 


Uratelornis 
chimaera 


OPINIONS AND DECLARATIONS 


Name of genus of 
which species 
specified in Col. (1) 
is the type species 


(3) 


Platalea Linnaeus, 
1758 


Porzana Vieillot, 
1816 


Psittacus Linnaeus, 
1758 


Psophia Linnaeus, 
1758 


Pteroglossus Illiger, 
1811 


Ptiloris Swainson, 
1825 


Rallus Linnaeus, 
1758 


Recurvirostra 
Linnaeus, 1758 


Sericulus Swainson, 
1825 


Sitta Linnaeus, 1758 


Sphenorynchus 
Lichtenstein, 1823 


Strigops Gray, 1845 


Sturnella Vieillot, 
1816 


Sturnus Linnaeus, 
1758 


Syrrhaptes Uliger, 
1811 


Tachyphonus Vieillot, 
1816 


Thamnophilus Vieillot, 
1816 


Uratelornis 
Rothschild, 1895 


DIRECTION 43 195 


ANNEXE 2 


Proposed addition to the ‘* Official List of Specific Names in Zoology ”’ 
of the specific names of twelve nominal species, subjectively 
identified with nominal species of later date which are the type 
species of genera of which the names were placed on the 
** Official List of Generic Names in Zoology ”’ in the 
period up to the end of 1936 


Specific name proposed to be 
Placed on the *‘ Official List of 
Specific Names in Zoology” 
(1) 


tiphia Linnaeus, 1758, as pub- 
lished in the combination 
Motacilla tiphia 


cristatus Shaw in White, 1790, as 
published in the combination 
Caprimulgus cristatus® 


nitidus Latham, 1790, as pub- 
lished in the combination Falco 
nitidus 


caeruleus Desfontaines, [1789], as 
published in the combination 
Falco caeruleus 


cristata Pallas, 1764, as published 
in the combination Columba 
cristata 


Remarks 
(not for inclusion in the entry 
to be made in the “* Official List’’) 
(2) 

A senior subjective synonym of 
leucoptera Vieillot, 1807, as 
published in the combination 
Sylvia leucoptera, the specific 
name of the type species of 
Aegithina Vieillot, 1816 

a senior subjective synonym of 
novaehollandiae Latham, 1790, 
as published in the com- 
bination Caprimulgus 
novae-hollandiae, the specific 
name of the type species of 
Aegotheles Vigors & Horsfield, 
1826 

a senior subjective synonym of 
cinerea Vieillot, 1816, as pub- 
lished in the combination 
Asturia [sic] cinerea, the 
specific name of the type 
species of Asturina Vieillot, 
1816 

a senior subjective synonym of 
caesius Savigny, 1809, as pub- 
lished in the combination 
Elanus caesius, the specific 
name of the type species of 
Elanus Savigny, 1809 

a senior subjective synonym of 
coronata Linnaeus, 1766, as 
published in the combination 
Columba coronata, the specific 
name of the type species of 
Goura Stephens, 1819 


8 For a discussion of the relative priority of the name Caprimulgus cristatus 
Shaw in White, 1790, and of its subjective synonym Caprimulgus novaehollandiae 
Latham, 1790, see the Report by the Secretary reproduced in paragraph 9 of the 


present Direction. 


196 


Specific name proposed to be 
placed on the “‘ Official List of 
Specific Names in Zoology ”’ 


(1) 
albicilla Linnaeus, 1758, as pub- 


lished in the combination Falco 
albicilla 


indus Boddaert, 1783, as published 
in the combination Falco indus 


militaris Linnaeus, 1758, as pub- 
lished in the combination 
Emberiza militaris 


hoazin Miiller, [1776], as pub- 
lished in the combination 
Phasianus hoazin 


wallicus Kerr, 1792, as published 
in the combination Psittacus 
wallicus 


elegans Gmelin, 1788, as pub- 
lished in the combination 
Psittacus elegans 


OPINIONS AND DECLARATIONS 


Remarks 


(not for inclusion in the entry 


to be made in the “‘ Official List’’) 


a 


ro) 


ran) 


rat) 


99 


(2) 
senior subjective synonym of 
nisus Savigny, 1809, as pub- 
lished in the combination 
Haliaeetus nisus, the specific 
name of the type species of 
Haliaeetus Savigny, 1809 


senior subjective synonym of 
pondicerianus Gmelin, 1788, 
as published in the combina- 
tion Falco pondicerianus, the 
specific name of the type 
species of Haliastur Selby, 
1840 


senior subjective synonym of 
americanus Gmelin, 1788, as 
published in the combination 
Oriolus americanus, the 
specific name of the type 
species of Leistes Vigors, 1825 


senior subjective synonym of 
cristatus Gmelin, 1788, as 
published in the combination 
Phasianus cristatus the specific 
name of the type species of 
Opisthocomus Illiger, 1811 


junior subjective synonym of 
formosus Latham, 1790, as 
published in the combination 
Psittacus formosus, the specific 
name of the type species of 
Pezoporus Illiger, 1811, which 
is invalid because a junior 
primary homonym of 
Psittacus formosus Scopoli, 
1769 


senior subjective synonym of 
pennantii Latham, 1790, as 
published in the combination 
Psittacus pennantii the specific 
name of the type species of 
Platycercus Vigors, 1825 


DIRECTION 43 197 


Specific name proposed to be Remarks 
placed on the ‘* Official List of (not for inclusion in the entry 
Specific Names in Zoology ”’ to be made in the “ Official List’) 
(1) (2) 
nigricephala Jameson, 1835, as a senior subjective synonym of 
published in the combination bilineatus Jardine & Selby, 
Tanagra nigricephala 1836, as published in the com- 


bination Spindalis bilineatus, 
the specific name of the type 
species of Spindalis Jardine & 
Selby, 1836 


ANNEXE 3 


Seven nominal genera of birds, the names of which were placed on the 
** Official List of Generic Names in Zoology ”’ in the period up to 
the end of 1936, where the author of the generic name cited the 
type species under a name which is a junior objective 
synonym of a previously published name 


Section A :—Particulars of the circumstances in which in seven 
cases an author when establishing a new nominal genus of 
birds either cited the type species under a previously published 
name which was a junior objective synonym of some older 
name or himself gratuitiously renamed the species in question 


(1) ‘* Coturnix ’’ Bonnaterre, 1790 


Bonnaterre placed in his genus Coturnix a nominal species to which 
he gave the new name Coturnix communis. This name was expressly 
stated to be a substitute for (nom. nov. pro) the name Tetrao coturnix 
Linnaeus, 1758. The above is therefore the type species by absolute 
tautonymy under Rule (d) in Article 30. Under the proposal now 
submitted Tetrao coturnix Linnaeus and not Coturnix communis 
Bonnaterre will be the type species of Coturnix Bonnaterre. 


(2) ‘* Eurynorhynchus ’’ Nilsson, 1821 


Nilsson treated this genus as being monotypical, placing in it only 
the nominal species to which he then gave the new name Eurynorhynchus 
griseus. Nilsson stated that this was only a substitute name for (nom. 
nov. pro) the name Platalea pygmaea Linnaeus, 1758. Under the 
present proposal Platalea pygmaea Linnaeus and not Eurynorhynchus 
griseus Nilsson will be the type species of the genus Eurynorhynchus 
Nilsson. 


198 OPINIONS AND DECLARATIONS 


(3) ‘* Heliornis ’’ Bonnaterre, 1790 


Bonnaterre treated this genus as monotypical, placing in it only 
the nominal species Heliornis fulicarius. In introducing this new 
nominal species, Bonnaterre cited two bibliographical references, of 
which the first was ** Buff. tom. 15.385’. When we turn to the passage 
in Boddaert’s Tabl. Pl. enlum. Hist. nat. Daubenton where the name 
Colymbus fulica first appears, we find that Boddaert based the nominal 
species so named on the same reference to Buffon (XV.385). Thus 
Boddaert’s Colymbus fulica and Bonnaterre’s later Heliornis fulicarius 
have the same passage in Buffon as their “‘ indication ’’ and are there- 
fore objective synonyms of one another. It may be noted that Buffon 
dealt with this species under the French (vernacular) name ‘‘ Grébe 
foulque ”’ and it may certainly be concluded that this name, which was 
quoted from Buffon by Boddaert, led the latter to select the Latin 
word “‘ fulica ’’ as the specific name for this species. Under the present 
proposals Colymbus fulica Boddaert and not Heliornis fulicarius 
Bonnaterre will be the type species of the genus Heliornis Bonnaterre. 


(4) ‘‘ Manucodia *’ Boddaert, 1783 


Boddaert treated this as a monotypical genus, placing in it only 
the nominal species to which he gave the new name Manucodia 
chalybea. In bringing forward this name, he cited a few bibliographical 
references but he did not cite by name the nominal species Paradisea 
[sic] chalybata Forster, 1781, with which his species is currently 
identified. Boddaert introduced the name Manucodia when dealing 
with plate 634 in Daubenton’s Histoire naturelle des Oiseaux. The 
same plate formed the basis of Forster’s P. chalybata. The name 
Manucodia chalybea Bonnaterre is therefore a substitute name for 
(nom. noy. pro) the name Paradisea [sic] chalybata Forster. Accordingly,’ 
the type species of Manucodia Bonnaterre will, under the present 
proposals, be Paradisea [sic] chalybata Forster and not Manucodia 
chalybea Bonnaterre. 


(5) °* Oriolus ’’ Linnaeus, 1766 


Linnaeus placed in this genus a nominal species to which he gave 
the new name Oriolus galbula. This name was expressly stated to be a 
substitute for (nom. nov. pro) the name Coracias oriolus Linnaeus, 1758. 
The above is therefore the type species by absolute tautonymy under 
Rule (d) in Article 30. Under the proposal now submitted,Coracias 
oriolus Linnaeus and not Oriolus galbula Linnaeus will be the type 
species of Oriolus Linnaeus, 1766. 


(6) ‘* Pandion ”’ Savigny, 1809 


Savigny treated this genus as being monotypical, placing in it only . 
the nominal species to which he gave the new name Pandion fluvialis. 
Savigny stated that this was a substitute name for (nom. nov. pro) 


DIRECTION 43 199 


the name Falco haliaetus Linnaeus, 1758. Under the present proposal, 
Falco haliaetus Linnaeus and not Pandion fluvialis Savigny will be the 
type species of the genus Pandion Savigny. 


(7) ‘* Polyplectron ’’ Temminck, 1807 


Temminck treated this genus as being monotypical, placing in it 
only the nominal species to which he then gave the new name 
Polyplectron argus. Temminck stated that this was a substitute name 
for (nom. nov. pro) the name Pavo bicalcaratus Linnaeus, 1758. Under 
the present proposal Pavo bicalcaratus Linnaeus and not Polyplectron 


argus will be the type species of the genus Polyplectron Temminck. 


Section B :—Seven pairs of objectively synonymous names 
of which in each case the later-published name is the one 
used by the original author of a generic name when citing 
the type species of the genus 


The older of a pair 
of objectively 
synonymous names 


for the type species of 


the genus specified 
in Col. (3) 


(1) 

Tetrao coturnix 
Linnaeus, 1758 
Syst. Nat. (ed. 10) 
i's 161 


Platalea pygmaea 
Linnaeus, 1758, 
Syst. Nat. (ed. 10) 
1 : 140 


Colymbus fulica 
Boddaert, 1783, 
Tabl. Pl. enlum. 
Hist. nat. : 54 


Paradisea {sic} 
chalybata Forster, 
1781 Indische 
Zool. : 40 


Coracias oriolus 
Linnaeus, 1758, 
Syst. Nat. (ed. 10) 
i: 107 


The later-published 
objective synonym of 


the name cited in 
Coll) 


(2) 
Coturnix communis 
Bonnaterre, 1791, 
Ency. méth., Orn. 
NEG 


Eurynorhynchus 
griseus Nilsson, 
1821, Ornith. svec. 
ZC ys /29 


Heliornis fulicarius 
Bonnaterre, 1790, 
Ency. méth., Orn. 
£265 


Manucodia chalybea 
Boddaert, 1783, 
Tabl. Pl. enlum. 
Hist. nat, #39 


Oriolus galbula 
Linnaeus, 1766, 
Syst. Nat. (ed. 12) 
1(1) : 160 


Name of genus, the 
type species of 
which was cited under 
the name specified in 
Col. (2) by the author 
of the generic name 


(3) 
Coturnix Bonnaterre 
1790, Ency. méth., 
Orn. 1s Ixxxvi, 216 


Eurynorhynchus 
Nilsson, 1821 
Ornith. svec. 2(1) : 
29 


Heliornis 
Bonnaterre, 1790 
Ency méth., Orn. 
1 : lxxxiv, 64 


Manucodia Boddaert 
(783; Dabl.PI. 
enlum. Hist. nat. : 


39 


Oriolus Linnaeus 
1766, Syst. Nat. 
(ed: 12) 11): 160 


200 OPINIONS AND DECLARATIONS 


The older of a pair The later-published Name of genus, the 


of objectively objective synonym of type species of 
synonymous names the name cited in which was cited under 
for the type species of Col. (1) the name specified in 
the genus specified Col. (2) by the author 
in Col. (3) of the generic name 
(1) (2) (3) 

Falco haliaetus Pandion fluvialis Pandion Savigny, 1809, 
Linnaeus, 1758, Savigny, 1809, Descr. Egypte 1(1) | 
Syst Nat. (ed. 10) Descr. Egypte 1(1) (Ois) : 69, 95 
i 9! (Ois) : 96 

Pavo bicalcaratus Polyplectron argus Polyplectron 
Linnaeus, 1758, Temminck, 1807, Temminck, 1807, 
Syst. Nat. (ed. 10) Cat. Syst. Cab. Cat. syst. Cab. 
11356 Ornith. : 149 Ornith. : 149 


Section C :—Entries recommended to be made on the 
Official List of Specific Names in Zoology® 


coturnix Linnaeus, 1758, as published in the combination Tetrao 
coturnix (specific name of type species of Coturnix Bonnaterre, 1790) 


pygmaea Linnaeus, 1758, as published in the combination Platalea 
pygmaea (specific name of type species of Eurynorhynchus Nilsson, 
21) 


fulica Boddaert, 1783, as published in the combination Colymbus 
fulica (specific name of type species of Heliornis Bonnaterre, 1790) 


chalybata Forster, 1781, as published in the combination Paradisea 
[sic] chalybata (specific name of type species of Manucodia Boddaert, 
1783) 


oriolus Linnaeus, 1758, as published in the combination Coracias 
oriolus (specific name of type species of Oriolus Linnaeus, 1766) 


haliaetus Linnaeus, 1758, as published in the combination Falco 
haliaetus (specific name of type species of Pandion Savigny, 1809) 


bicalcaratus Linnaeus, 1758, as published in the combination Pavo 
bicalcaratus (specific name of type species of Polyplectron Temminck, 
1807). 


® The names which, as shown in Column (2) of Section B of the present Annexe, 
are junior objective synonyms of the names enumerated in the present Annexe 
have been placed on the Official Index of Rejected and Invalid Specific Names in 
ese by the Ruling given in Direction 44 (pp. 219—220 of the present 
volume). 


DIRECTION 43 201 


3. Registration of the present application : Upon the receipt 
of Mr. Hemming’s paper, the question of placing on the Official 
List of Specific Names in Zoology the specific names discussed 
therein was allotted the Registered Number Z.N.(S.) 993. 


Il. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


4. Issue of Voting Paper V.P.(O.M.)(55)31 : On 12th October 
1955 a Voting Paper (V.P.(O.M.)(55)31) was issued in which each 
Member of the Commission was asked (1) to state whether he 
agreed that, “in conformity with the General Directives issued 
to the International Commission on Zoological Nomenclature 
by the Thirteenth International Congress of Zoology, Paris, 1948, 
regarding the placing of names on the appropriate Official Lists 
and Official Indexes of names dealt with in, or otherwise involved 
in, Opinions rendered prior to 1948, the names referred to in 
Sections (a) and (b) of paragraph 10 of the paper bearing the 
Registered Number Z.N.(S.) 993 submitted by the Secretary 
simultaneously with the present Voting Paper [i.e. in the para- 
graph numbered as above in the paper reproduced in paragraph 2 
of the present Direction] be placed on the Official List of Specific 
Names in Zoology as there recommended and that the other 
action recommended in the said paragraph be similarly 
approved ”’ and (2), if he did not so agree as regards any given 
item, to indicate that item. 


5. Withdrawal from the scope of Voting Paper V.P.(O.M.)(55) 
31 of proposals submitted in regard to two specific names: On 
30th October 1955 Mr. Hemming, as Secretary, placed the 


202 OPINIONS AND DECLARATIONS 


following Minute on the Commission’s File Z.N.(S.) 993 :— 


Withdrawal from the scope of Voting Paper V.P.(O.M.)(55)31 of 
proposals submitted in regard to two specific names 


MINUTE by FRANCIS HEMMING, C.M.G., C.B.E. 
(Secretary to the International Commission on Zoological Nomenclature) 


I have this day received from Professor Ernst Mayr a letter dated 
26th October 1955 in which he has drawn my attention to the fact that 
within the last twenty years efforts have been made to show that the 
name gallicus Gmelin (J.F.), 1788, as published in the combination 
Falco gallicus, applies to the same taxon as that to which is applicable 
the specific name ferox Gmelin (S.G.), 1771, as published in the com- 
bination Accipiter ferox and that by the authors who had taken this 
view the name gallicus had been treated as a junior subjective synonym 
of the name ferox. It appears, however, from Professor Mayr’s letter 
that for some years past the attempt to synonymise the above names 
has been abandoned. Nevertheless, this is a matter which ought, in 
my opinion, to be examined separately before a decision is taken by 
the Commission on the question of the admission of the name gallicus 
Gmelin to the Official List. This will involve the postponement of a 
decision in relation to the name for the type species of Circaetus 
Vieillot, 1816. 


2. I have further to place on record that one of the specific names 
which in paper Z.N.(S.) 993 was proposed for admission to the Official 
List has, I now find, already been placed on that List. This is the name 
of the type species of Neophron Savigny, 1809. This name was 
originally published in the misspelled form perenopterus (Vultur 
perenopterus Linnaeus, 1758) but was emended by later authors to the 
correct spelling percnopterus. On the proposal of the Standing 
Committee on Ornithological Nomenclature established by the Tenth 
International Ornithological Congress (1952, Bull. zool. Nomencl. 
9 : 52) this emendation has been accepted by the Commission in its 
Opinion 411 (now in the press)!° and in the same Opinion this name in its 
emended form percnopterus was placed on the Official List. 


3. For the reasons set forth above, I now, as Secretary to the 
International Commission, hereby direct as follows :— 


(a) that the proposals relating to the under-mentioned specific 
names which in the paper numbered Z.N.(S.) 993 were sub- 
mitted to the Commission concurrently with Voting Paper 
V.P.(O.M.)(55)31 be withdrawn from the scope of the foregoing 
Voting Paper :— 

(i) gallicus Gmelin (J.F.), 1788, as published in the combina- 
tion Falco gallicus ; 

(ii) percnopterus (emend. of perenopterus) Linnaeus, 1758, as 
published in the combination Vultur perenopterus ; 


10 The Opinion here referred to was published on 31st July 1956 (Ops. Decls. int. 
Comm. zool. Nomencl. 13 : 191—202). 


DIRECTION 43 203 


(b) that the question whether the specific name gallicus Gmelin, 
1788, should be placed on the Official List of Specific Names 
in Zoology be examined on Commission File Z.N.(S.) 1052, 
which has this day been opened for this purpose, with a view 
to the submission after consultation with interested specialists 
of proposals as to the action to be taken in this matter. 


6. The Prescribed Voting Period : As Voting Paper V.P.(O.M.) 
(55)31 was issued under the One-Month Rule, the Prescribed 
Voting Period closed on 12th November 1955. 


7. Particulars of the Voting on Voting Paper V.P.(O.M)(55)31 : 
At the close of the Prescribed Voting Period, the state of the 
voting on Voting Paper V.P.(O.M.)(55)31 was as follows :— 


(a) Affirmative Votes had been given by the following twenty-four 
(24) Commissioners (arranged in the order in which Votes 
were received) : 


Riley ; Hering; Holthuis; Lemche; Vokes; Stoll; 
Esaki; Boschma; Miller; Bradley (J.C.) ; Sylvester- 
Bradley; Prantl; Mayr; Jaczewski;  Tortonese ; 
do Amaral ; Dymond ; Hemming; Bonnet; Mertens ; 
Cabrera ; Key ; Kithnelt ; Hanko ; 


(b) Negative Votes : 


None ; 


(c) On Leave of Absence, one (1) : 


Bodenheimer ; 


204 OPINIONS AND DECLARATIONS 


(d) Voting Papers not returned : 


None. 


8. Declaration of Result of Vote : On 2lst November 1955, 
Mr. Hemming, Secretary to the International Commission, acting 
as Returning Officer for the Vote taken on Voting Paper V.P. 
(O.M.)(55)31, signed a Certificate that the Votes cast were as set 
out in paragraph 7 above and declaring that the proposal sub- 
mitted in the foregoing Voting Paper had been duly adopted and 
that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


9. Report by the Secretary on the relative dates of publication of 
the names ‘* Caprimulgus novaehollandiae ’? Latham and ‘‘ Capri- 
mulgus cristatus ’’ Shaw in White, both currently treated as having 
been published in 1790: On 3lst January 1956 Mr. Hemming as 
Secretary placed on the Commission’s File Z.N.(S.)993 the follow- 
ing Report on the relative priority of the names Caprimulgus 
novaehollandiae Latham, and Caprimulgus cristatus Shaw in 
White, which are currently treated as having been published in 
the year 1790 and are synonymised with one another, Shaw’s 
name cristatus being treated as having priority over Latham’s 
name novaehollandiae. 


Relative priority of the binomina ‘* Caprimulgus cristatus ’’ Shaw in 
White, 1790, and ‘‘ Caprimulgus novaehollandiae ’’ Latham, 1790 


REPORT by FRANCIS HEMMING, C.M.G., C.B.E. 
(Secretary to the International Commission on Zoological Nomenclature) 


In returning Voting Paper V.P.(O.M.)(55) 31 Professor Ernst Mayr 
drew attention to the fact that, as stated in the second entry in Annexe 2 
to the paper submitted concurrently with the foregoing Voting Paper,” 
the name of the type species of the genus Aegotheles Vigors & Horsfield, 
1826, was Caprimulgus novaehollandiae Latham, 1790, but that that 
name was currently treated as a junior subjective synonym of the name 
Caprimulgus cristatus Shaw in White published in the same year. 
While not suggesting that this treatment was incorrect, Professor 


11 See page 195 of the present Direction. 


DIRECTION 43 ; 205 


Mayr thought it desirable that, if possible, the relative priority of 
these two names should be placed on a firm footing. 


2. On Professor Mayr’s suggestion I have investigated the above 
matter on which I now submit the following Report :— 


(a) The relative priority of the names Caprimulgus novaehollandiae 
Latham and Caprimulgus cristatus Shaw was discussed by 
Gregory Mathews ({1918—1919], Birds Australia 7 : 51) who 
brought forward the following evidence :— 


(i) Latham’s Index Ornith. is dated ‘‘ 1790’ but its exact 
date of publication cannot be ascertained. Gregory 
Mathews pointed out, however, that this work had 
sometimes been treated by later authors as having 
appeared in 1791 and not in 1790. His conclusion 
was that it was probably published late in 1790. 


(ii) As regards the name Caprimulgus cristatus Shaw in White, 
Gregory Mathews pointed out that the plate in Shaw’s 
Journal containing a figure of this species is dated 
“Dec. 29th 1789’. He pointed out further that this 
work was reviewed in the Gentleman’s Magazine (55(2) : 
742) in its issue of August 1790. Mathews added that 
the above serial was known not to have published reviews 
of books immediately upon their publication. He con- 
cluded that this work was probably published early in 
1790. 


(b) An examination of the two books concerned carried out by 
Miss D. N. Noakes, B.Sc.,!* has brought to light one piece of 
information not elicited by Gregory Mathews. This is that, 
while, as stated by him, the title page bears the date “‘ 1790 ”’ 
(i) the dedication by the author is written from ‘* Sydney 
Cove, Port Jackson, N.S. Wales’? and is dated ‘*‘ November 
18th 1788” and (ii) all the plates bear (in very small print) 
the legend “‘ Published as the act directs, Dec. 29th, 1789 by 
J. Debrett ’’. The use in this legend of the word “‘ Published ”’, 
which was not mentioned by Gregory Mathews, gives much 
greater precision to the date which he quoted. 


(c) Under a decision taken by the Thirteenth International Congress 
of Zoology, Paris, 1948, which was then incorporated into 
the Régles (1950, Bull. zool. Nomencl. 4 : 223—225), a book 
or paper, the exact date of publication of which in a given year 
is not known, is to be treated for the purposes of the applica- 


12 Research Assistant, Office of the International Commission on Zoological 
Nomenclature. 


206 OPINIONS AND DECLARATIONS 


tion of the Law of Priority to new names included in it, as 
having been published not earlier than the earliest date in that 
year in which it is known to have been published and, where 
no such evidence is available, as having been published on the 
last day of the year in question. 


(d) When the foregoing decision by the Paris Congress is applied in 
the present case, it is seen :— 


(i) that the earliest date in 1790 on which White’s Journal, 
and therefore the name Caprimulgus cristatus Shaw, is 
definitely known to have been published is the date 
in the month of August in which the issue of ihe 
Gentleman’s Magazine containing a review of White’s 
Journal was published ; 


(ii) that, as nothing apart from the date “‘ 1790” on the title 
page is known regarding the date of publication of 
Latham’s Index ornithologicus, the earliest date which 
can be definitely assigned to the above work and there- 
fore to the name Caprimulgus novaehollandiae Latham 
is 3lst December 1790. 


(e) For the reasons set forth above, the name Caprimulgus cristatus 
Shaw in White has between four and five months priority 
under the Régles over the name Caprimulgus novaehollandiae 
Latham. 


3. In view of the evidence brought forward in the foregoing paragraph 
the proposals submitted to the Commission in regard to the relative 
priority of the names discussed above is seen to have been correct 
and therefore, under the General Directive issued to the Commission 
by the International Congress of Zoology regarding the placing of 
names on the Official List of Specific Names in Zoology, the specific 
name cristatus Shaw, 1790, and not the name novaehollandiae Latham, 
1790, should now be placed on the foregoing List, as proposed in the 
paper referred to in paragraph 1 of the present Report and as subse- 
quently approved by the Commission in its vote on Voting Paper 
V.P.(O.M.)(55)31. In order to obviate the risk of any future misunder- 
standing arising in regard to this matter it is desirable that, when in 
accordance with the decision of the Commission the name cristatus 
Shaw in White is placed on the Official List, the date ‘‘ August ” should 
be cited in square brackets, together with the date “‘ 1790” and, as 
Secretary to the Commission, I hereby direct that this citation be made. 


DIRECTION 43 207 


10. Determination of the status under Article 30 of seven nominal 
species objectively identical with later-established nominal species 
cited for the type species of genera by the original authors of the 
generic names concerned: On 25th February 1956 Mr. Hemming as 
Secretary placed on the Commission’s File Z.N.(S.)993 the follow- 
ing Minute clarifying the status under Article 30 or seven nominal 
species, each of which is objectively identical with a later-estab- 
lished nominal species which was cited for the type species of a 
genus in the Class Aves by the original author of the 
generic name in question:— 


Status under Article 30 of a nominal species which is objectively 
identical with the nominal species cited as the type species 
of a genus by the original author of the generic name concerned 


MINUTE by FRANCIS HEMMING, C.M.G., C.B.E. 
(Secretary to the International Commission on Zoological Nomenclature) 


At the time of the submission to the Commission of the paper 
bearing the Registered Number Z.N.(S.) 993 it was not clear what 
nominal species should be treated as being the type species of a genus 
when two or more objectively identical nominal species were involved. 
This problem arose in connection with the names to be used for the 
type species of seven genera in the Class Aves, the names of which were 
placed on the Official List of Generic Names in Zoology in the period 
up to the close of 1936, and therefore called for decision in the vote 
to be taken by the Commission on Voting Paper V.P.(O.M.)(55)31. 


2. A proposal for the adoption of a Declaration clarifying the inter- 
pretation of Article 30 in the above regard was under consideration 
at the time when the foregoing paper was submitted to the Commission 
and it was anticipated that a decision thereon would have been taken by 
the Commission before the time came for the preparation of a 
Direction embodying the Commission’s decision on the Voting Paper 
referred to above. Accordingly, in the paper submitted with the 
foregoing Voting Paper alternative proposals were placed before the 
Commission in paragraph 10(b),!° under the first of which, in the event 
of the adoption of the suggested Declaration, the seven senior objective 
synonyms listed in Section C of Annexe 3 would be treated as the 
specific names of the type species of the genera concerned and in the 
event of the non-adoption of that Declaration as the specific names 
of nominal species objectively identical with the nominal species 
which would in that event be the type species of the genera in question. 


3. The foregoing alternative proposals, together with the other 
proposals submitted with Paper Z.N.(S.) 993 were approved by the 
Commission by its Vote on Voting Paper V.P.(O.M.)(55)31. 


18 See page 190 of the present Direction. 


208 OPINIONS AND DECLARATIONS 


4. By its Vote taken on Voting Paper V.P.(55)30 the Commission 
on 22nd February 1956 adopted the suggested Declaration referred 
to in paragraph 2 above, under which it ruled that under Article 30 
of the Régles where there are two or more objectively identical nominal 
species, the designation, indication or selection of any one of those 
nominal species to be the type species of a genus is to be treated as 
the designation, indication or selection of whichever of the nominal 
species concerned has the oldest available name, irrespective of whether 
or not that nominal species was cited by the author of the generic 
name in question. The decision so taken has since been embodied in 
Declaration 21 (now in the press)"*. 


5. As Secretary to the Commission, I accordingly hereby direct as 
follows :— 


(1) that in the Ruling to be given in the Direction embodying the 
decision taken by the Commission on Voting Paper V.P.(O.M.) 
(55)31 a Section be included prescribing that in accordance 
with the provisions of Declaration 21 the seven nominal species 
specified in Column (1) of Section B of Annexe 3 to the Paper 
Z.N.(S.) 993 submitted with the foregoing Voting Paper, each 
of which is objectively identical with the later-established 
nominal species enumerated in Column (2) of the said Annexe 
which was cited by the author of the generic name concerned, 
be treated as the type species of the genera severally specified 
in Column (3) of the Annexe in question ; . 


(2) that consequential upon the foregoing decision, the seven specific 
names enumerated in Section C of the above Annexe be 
entered on the Official List of Specific Names in Zoology 
as the specific names of the type species of the genera referred 
to above. 


11. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 29th February 1956 Mr. Hemming prepared the Ruling given 
in the present Direction and at the same time signed a Certificate 
that the terms of that Ruling were in complete accord with those 
of the proposal approved by the International Commission in its 
Vote on Voting Paper V.P.(O.M.)(55)31, subject (i) to the with- 
drawal from the scope of the foregoing Voting Paper of the 
proposals in regard to the two specific names specified in the 
Secretary’s Minute of 30th October 1955 (reproduced in paragraph 
5 of the present Direction), (ii) to the insertion of particulars of 
the month as well as the year of publication of the specific name. 


14 See Footnote 5. 


DIRECTION 43 209 


cristatus Shaw, [August] 1790, as published in the combination 
Caprimulgus cristatus, when that name is entered on the Official 
List of Specific Names in Zoology, as directed in the Minute 
executed by the Secretary on 3lst January 1956 (paragraph 9 of 
the present Direction) and (iii) to the determination of the nature 
of the entry to be made on the Official List of Specific Names in 
Zoology in relation to seven other specific names as prescribed 
in the Secretary’s Minute of 25th February 1956 (reproduced in 
paragraph 10 of the present Direction). 


12. Original References: The original references for the names 
placed on the Official List of Specific Names in Zoology by the 
Ruling given in the present Direction have already been noted in 
connection with the Official List of Generic Names in Zoology, the 
nominal species bearing the names concerned being either the 
type species of genera, the names of which have been inscribed 
on that List or are older-established nominal species subjectively 
identified therewith. 


13. Family-Group Names: No question relating to family- 
group names arises in connection with the Ruling given in the 
present Direction. 


14. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in 
dealing with the present case, and the present Direction is accord- 
ingly hereby rendered in the name of the said International Com- 
mission by the under-signed Francis Hemming, Secretary to the 
International Commission on Zoological Nomenclature, in virtue 
of all and every the powers conferred upon him in that behalf. 


15. The present Direction shall be known as Direction Forty- 
Three (43) of the International Commission on Zoological 
Nomenclature. 


Done in London, this Twenty-Ninth day of February, Nineteen 
Hundred and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


210 OPINIONS AND DECLARATIONS 


APPENDIX 


Particulars of the action taken or now in progress in the case of the 
eighteen names of genera belonging to the Class Aves which were 
placed on the ‘‘ Official List of Generic Names in Zoology ”’ in the 
period up to the end of 1936, as regards which no action has been 
taken in the present ‘‘ Direction ’’ under the General Directive 
issued to the International Commission by the International 
Congress of Zoology relative to the placing on the 
** Official List of Specific Names in Zoology ”’ of the 
specific names of the type species of genera, the names 
of which were placed on the ‘* Official List of 
Generic Names in Zoology’’ in the foregoing 
period 


(A) Names of ten genera, the names of the type species of which, or 
senior subjective synonyms of such names, have now been placed 
on the “* Official List of Specific Names in Zoology” 


| Balaeniceps Direction 19 1955, Ops. Decls. int. Comm. Zool. 
Nomencl. 1(C) : 133—144 

2 Burhinus Direction 20 1955, ibid. 1(C) : 145—160 

3 Campephaga Opinion 407 1956, ibid. 13 : 131—146 

4 Colluricincla Opinion 410 1956, ibid. 13 : 173—190 


5 Ectopistes Direction 18 1955, ibid. 1(C) : 113—132 

6 Gallinago Direction 39 1956, ibid. 17D) : 95—138 

7 Grus Direction 55 (ibid. 1(D), Part D.16, now in the 
press) 

8 Neophron Opinion 411 1956, ibid. 13 : 191—202 

9 Oedicnemus (See Burhinus above.) 

10 Struthio Opinion 413 1956, ibid. 13 : 233—242 


(B) Name of one genus, for the addition to the *“‘ Official List of Specific 
Names in Zoology’ of the name of the type species of which a 
proposal has now been published in the “‘ Bulletin of Zoological 
Nomenclature ”’ 


11 Circaetus (Z.N.(S.) 1052) 1956, Bull. zool. Nomencl. 12 : 120— 
122 


(C) Names of seven genera, in respect of which investigations are 
proceeding on the Files noted below on questions relating to 
the type species of the genera concerned 


12 Anas 


13. Gypaetus oe 
14 Nyctea joke ZINAS:) 1050 
15 Vireo i 

16 Bubo bee 

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18 Trichoglossus File Z.N.(S.) 1055 


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OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, c.M.G., C.B.E. 


Secretary to the Commission 


DIRECTION 44 


Correction of certain entries relating to the names of 
genera in the Class Aves made on the Official List of 
Generic Names in Zoology in the period up to the close 
of 1936 and addition to the Official Indexes of Rejected 
and Invalid Names in Zoology of certain names given to 


VOLUME 1. SECTION D. Part D.8. Pp. 211—232 
genera and species of the same Class. 


LONDON : 
Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 


1956 
Price Fifteen Shillings So ee 
VACUO PN 
(All rights reserved) f 2 “VN 
TOV NY 49-rk 
= \\ NO\ 20 195 F 
Issued 19th October, 1956 8 Lipearey 7 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 44 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JoRDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President : Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A. 
(12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) © 
Secretary: Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(Arranged in order of precedence by reference to date of election or of most recent re-election 
as prescribed by the International Congress of Zoology) 


Professor H. BOSCHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. ee eee (Universitetets Zoologiske Museum, Copenhagen, Denmark) (27th ~ 
uly 1 ) 

Professor Teiso EsAkI (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEy (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (5th July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitat zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMoND (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) (12th August 
1953) (President) 

Professor Harold E. Voxes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) (12th 
August 1953) 

Professor Béla HANKO (Mezdgazdasdgi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y., 
U.S.A.) (12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. Hottuuts (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 

Canberra, A.C.T., Australia) (ASth October 1954): 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum V Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KtHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

neve. F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 
1954 

Professor Ernst MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico TORTONESE (Museo di Storia Naturale ‘‘ G. Doria”’, Genova, Italy) 
(16th December 1954) 


DIRECTION 44 


CORRECTION OF CERTAIN ENTRIES RELATING TO THE 
NAMES OF GENERA IN THE CLASS AVES MADE ON 
THE ‘ OFFICIAL LIST OF GENERIC NAMES IN 
ZOOLOGY ” IN THE PERIOD UP TO THE CLOSE 
OF 1936 AND ADDITION TO THE ‘° OFFICIAL 
INDEXES OF REJECTED AND INVALID NAMES 
IN ZOOLOGY ”’ OF CERTAIN NAMES GIVEN 
TO GENERA AND SPECIES OF THE SAME 
CLASS 


RULING :—(1) The entries on the Official List of 
Generic Names in Zoology in regard to the names of 
three genera in the Class Aves made by the Ruling 
given in Opinion 67 are hereby corrected in the manner 
specified below :— 


(1) Alauda Linnaeus, 1758: The nominal species 
Alauda arvensis Linnaeus, 1758, was first selected 
to be the type species of the genus Alauda 
Linnaeus, 1758, by Selby in 1825 (///. Brit. 
Ornith. 1 (Landbirds) : xxix, xxxiil) and this 
reference is accordingly hereby substituted for the 
fcorrect teference ~-Tsd.):Swamson,:: '82:7)7?. 


(2) Fulmarus Stephens, 1826: The nominal species 
Procellaria glacialis Linnaeus, 1761, was first 
selected to be the type species of the genus 
Fulmarus Stephens, 1826, by Gray (G.R.) in 
1855 (Cat. Gen. Sub-Gen. Birds Brit. Mus. : 129) 
and this reference is accordingly hereby sub- 


stituted for the incorrect reference ‘‘ Palmer, 
FS36y"’: 


(3) Heliornis Bonnaterre, 1790: The generic name 


Heliornis Bonnaterre, was published in Part 1 
of the ** Ornithologie ”’ Section of the Encyclopédie 


Bav 4A 1Q4h 


214 OPINIONS AND DECLARATIONS 


méthodique in 1790 and not in 1791 as incorrectly 
stated in Opinion 67 and appeared on page 64 
as well as on page Ixxxiv. The entry on the 
Official List is accordingly hereby corrected by 
the substitution of the date “1790” for the 
date “* 1791 ”? and completed by the addition of a 
reference to page 64. 


(2) The nominal species Manucodia chalybea Boddaert, 
1783, a junior objective synonym of Paradisea [sic] 
chalybata Forster, 1781, would have been the type 
species of Manucodia Boddaert, 1783, if it had not been 
for the provisions of Declaration 21+, under which the 
type species of this genus is Paradisea [sic] chalybata 
Forster, 1781, as the senior of the two objective synonyms 
concerned. The specific name chalybata Forster was 
misspelled as chalybeata in the Ruling given in Opinion 67. 
This spelling, as an Erroneous Subsequent Spelling, is 
placed on the Official Index of Rejected and Invalid 
Specific Names in Zoology by Ruling (4) in the present 
Direction. 


(3) The under-mentioned generic names are hereby 
placed on the Official Index of Rejected and Invalid 
Generic Names in Zoology with the Name Numbers 
severally specified below :— 


(1) Aegithia Blyth, 1860 (an Erroneous Subsequent 
Spelling for Aegithina Vieillot, 1816) (Name 
ING. 152 75 


(2) Aix Agassiz (J.L.R.), 1846 (An Invalid Emendation 
of Aia Eyton, 1838 and a junior homonym of 
Aix Boie, 1838) (Name No. 528) ; 


(3) Alauda Daudin, 1816 (a junior homonym of Alauda 
Linnaeus, 1758) (Name No. 529) ; 


1 Published in 1956 (Ops. Decls. int. Comm. zool. Nomencl. 12 : i—viii). 


DIRECTION 44 215 


(4) Aptenodita Bonnaterre, 1791 (an Erroneous Sub- 
sequent Spelling for Aptenodytes Miller, 1778) 
(Name No. 530) ; 


(5) Aptenodites Lesson, 1837 (an Erroneous Subse- 
quent Spelling for Aptenodytes Miller, 1778) 
(Name No. 531) ; 


(6) Aptenodyta Gmelin (J.F.), 1788 (an Erroneous 
Subsequent Spelling for Aptenodytes Miller, 1778) 
(Name No. 532) ; 


(7) Aptenodytes Forster (J.R.), 1777 (a nomen nudum) 
(Name No. 533) ; 


(8) Aulacorhynchus Dittmar, 1872 (a junior homonym 
of Aulacorhynchus Gould, 1834) (Name No. 534) ; 


(9) Brotogerys Agassiz (J.L.R.), 1846 (an Invalid | 
Emendation of Brotogeris Vigors, 1825) (Name 
IWO.°935) ; 


(10) Capito Gistl, 1848 (a junior homonym of Capito 
Vieillot, 1816) (Name No. 536) ; 


(11) Catharthes d’Orbigny, [1835] (an Erroneous Sub- 
sequent Spelling for Cathartes Illiger, 1811) 
(Name No. 537) ; 


(12) Centrocercus Fitzinger, 1843 (a junior homonym 
of ne ay Swainson, 1832) (Name No. 
538) ; 


(13) Cephalopterus Risso, 1810 (a junior homonym 
of Cephalopterus Geoffroy, 1809 (Name No. 539) ; 


(14) Cephalopterus Powrie, 1870 (a junior homonym of 
Cephalopterus Geoffroy, 1809) (Name No. 540) ; 


(15) Chauna Loew, 1847 (a junior homonym of Chauna 
Illiger, 1811) (Name No. 541) ; 


216 OPINIONS AND DECLARATIONS 


(16) Chrysolophus Swainson, 1837 (a junior homonym 
of Chrysolophus Gray (J.E.), [1833—1834]) (Name 
No. 542) ; 


(17) Clamator Blyth, 1852 (a junior homonym of 
Clamator Kaup, 1829) (Name No. 543) ; 


(18) Coerebra Boie, 1826 (an Erroneous Subsequent 
Spelling for Coereba Vieillot, 1816) (Name No. 
544) ; 


(19) Coturnix Cuvier, 1800 (a nomen nudum) (Name 
No. 545) ; 


(20) Eurynorhinchus Lafresnaye, 1842 (an Erroneous 
Subsequent Spelling for Eurynorhynchus Nilsson, 
1821) (Name No. 546) ; 


(21) Eurynorynchus Reichenbach, [1852] (an Erroneous 
Subsequent Spelling for Eurynorhynchus Nilsson, 
1821) (Name No. 547) ; 


(22) Eurylaemus Gloger, [1841] (an Erroneous Subse- 
quent Spelling for Eurylaimus Horsfield, 1821) 
(Name No. 548) ; 


(23) Eurypygia Swainson, 1837 (an Erroneous Subse- 
quent Spelling for Eurypyga Lliger, 1811) (Name 
No. 549) ; 


(24) Gampsonyx Jordan, 1847 (a junior homonym of 
Gampsonyx Vigors, 1825) (Name No. 550) ; 


(25) Haliaethus Lesson, 1831 (an Erroneous Subse- 
quent Spelling for Haliaeetus Savigny, 1809) 
(Name No. 551) ; 


(26) Haliaétos Bonaparte, 1826 (an Erroneous Subse- 
quent Spelling for Haliaeetus Savigny, 1809) 
(Name No. 552) ; 


DIRECTION 44 21¥ 


(27) Haliaetus Cuvier, 1816 (an Erroneous Subsequent 
Spelling for Haliaeetus Savigny, 1809) (Name 
Nor593)8 


(28) Heliornis Billberg, 1820 (a junior homonym of 
Heliornis Bonnaterre, 1790) (Name No. 554) ; 


(29) Ibidorhinchus David, 1875 (an Erroneous Sub- 
sequent Spelling for [bidorhyncha Vigors, 1832) 
(Name No. 555) ; 


(30) Ibidorhynchus Gray (G.R.), 1844 (an Erroneous 
Subsequent Spelling for Jbidorhyncha Vigors, 
1832) (Name No. 556) ; 


(31) Musophagus Bonnaterre, 1790 (an Invalid Emenda- 
tion of Musophaga Isert, 1789) (Name No. 557) ; 


(32) Numidia Forster, 1817 (an Erroneous Subsequent 
Spelling for Numida Linnaeus, 1764) (Name 
No. 558) ; 


(33) Pachycephala Klug, 1834 (a junior homonym of 
Pachycephala Vigors, 1825) (Name No. 559) ; 


(34) Pachycephala Doleschall, 1858 (a junior homonym 
of Pachycephala Vigors, 1825) (Name No. 560) ; 


(35) Pachycephala Lioy, 1864 (a junior homonym of 
Pachycephala Vigors, 1825) (Name No. 561); 


(36) Pachycephala Broun, 1881 (a junior homonym of 
Pachycephala Vigors, 1825) (Name No. 562) ; 


(37) Pachycephala Fairmaire, 1883 (a junior homonym 
of Pachycephala Vigors, 1825) (Name No. 563) ; 


(38) Pachycephalus Stephens, 1826 (an Invalid Emenda- 
tion of Pachycephala Vigors, 1825) (Name No. 
564) ; 


218 OPINIONS AND DECLARATIONS 


(39) Parotica Wagler, 1827 (an Erroneous Subsequent 
Spelling for Parotia Vieillot, 1816) (Name No. 
565) ; 


(40) Pezoporus Klug, [1842] (a junior homonym of 
Pezoporus Illiger, 1811) (Name No. 566) ; 


(41) Pezoporus Foerster, 1868 (a junior homonym of 
Pezoporus Illiger, 1811) (Name No. 567) ; 


(42) Psittacus Catesby, 1777 (a junior homonym of 
Psittacus Linnaeus, 1758) (Name No. 568) ; 


(43) Pteroglossus Chaudoir, 1843 (a junior homonym 
of Pteroglossus Illiger, 1811) (Name No. 569) ; 


(44) Ptilorhys Bonaparte, 1854 (an Erroneous Subse- 
quent Spelling for Priloris Swainson, 1825) (Name 
No. 570); 


(45) Ptilornis Gray (G.R.), 1869 (an Erroneous Sub- 
sequent Spelling for Ptiloris Swainson, 1825) 
(Name No. 571) ; 


(46) Strigopsis Bonaparte, 1849 (an Invalid Emendation 
of Strigops Gray (G.R.), 1845) (Name No. 572). 


(47) Stringops Finsch, 1867 (an Erroneous Subsequent 
Spelling for Strigops Gray (G.R.), 1845) (Name 
Noe. 3573); 


(48) Stringopsis van der Hoeven, 1855 (an Erroneous 
Subsequent Spelling for Strigops Gray (G.R.), 
1845) (Name No. 574) ; 


(49) Strygops Des Murs, [1853] (an Erroneous Sub- 
sequent Spelling for Strigops Gray (G.R.), 1845) 
(Name No. 575) ; 


(50) Ptilorhis Agassiz (J.L.R.), 1846 (an Invalid Emenda- 
tion of Priloris Swainson, 1825) (Name No. 576). 


DIRECTION 44 219 


(4) The under-mentioned specific names are hereby 
placed on the Official Index of Rejected and Invalid Specific 
Names in Zoology with the Name Numbers severally 
specified below :— 


(1) argus Temminck, 1807, as published in the com- 
bination Polyplectron argus (a junior objective 
synonym of bicalcaratus Linnaeus, 1758, as 
published in the combination Pavo bicalcaratus) 
(Name No. 343) ; 


(2) chalybea Boddaert, 1783, as published in the com- 
bination Manucodia chalybea (a junior objective 
synonym of chalybata Forster, 1781, as published 
in the combination Paradisea [sic] chalybata) 
(Name No. 344) ; 


(3) chalybeata Stiles, 1916, as published in the com- 
bination Paradisea chalybeata (an Erroneous 
Subsequent Spelling for chalybata Forster, 1781, 
as published in the combination Paradisea [sic] 
chalybata) (Name No. 345) ; 


(4) communis Bonnaterre, 1791, as published in the 
combination Coturnix communis (a junior objec- 
tive synonym of coturnix Linnaeus, 1758, as 
published in the combination Tetrao coturnix) 
(Name No. 346) ; 


(5) fluvialis Savigny, 1809, as published in the combina- 
tion Pandion fluvialis (a junior objective synonym 
of haliaetus Linnaeus, 1758, as published in the 
combination Falco haliaetus) (Name No. 347) ; 


(6) formosus Latham, 1790, as published in the com- 
bination Psittacus formosus (a junior primary 
homonym of formosus Scopoli, 1769, as published 
in the combination Psittacus formosus) (Name 
No. 348) ; 


220 OPINIONS AND DECLARATIONS 


(7) fulicarius Bonnaterre, 1790, as published in the 
combination Heliornis fulicarius (a junior objective 
synonym of fulica Boddaert, 1783, as published 
a He combination Colymbus fulica) (Name No. 


(8) galbula Linnaeus, 1766, as published in the com- 
bination Oriolus galbula (a junior objective 
synonym of oriolus Linnaeus, 1758, as published 
- me combination Coracias oriolus) (Name No. 

50): 


(9) griseus Nilsson, 1821, as published in the com- 
bination Eurynorhynchus griseus (a junior objective 
synonym of pygmaea Linnaeus, 1758, as published 
i _ combination Platalea pygmaea) (Name No. 

OP): 


J. THE SUBJECT MATTER OF THE PRESENT 
"DIRECTION, 


The present Direction is the third? of a series containing decisions 
supplementary to the Rulings in regard to the names of genera 
in the Class Aves placed on the Official List of Generic Names 
in Zoology in the period up to the close of 1936. These decisions 
have been taken by the International Commission on Zoological 
Nomenclature in conformity with General Directives issued 
by the International Congress of Zoology that the entries made 
on the Official List in that period be completed by the entry 
thereon in respect of each generic name of certain additional 
particulars currently required for new entries on that List and 
that the specific name of the type species of every genus, the 
name of which was entered on the Official List in the foregoing 
period, be entered upon the Official List of Specific Names in 


2 The earlier Directions here referred to are Direction 26 (1955, Ops. Decls. int. 
Comm. zool. Nomencl. 1(C) : 259—272) and Direction 43 published in 1956 
as the immediately preceding Part of the present Section (Section D) of volume 1. 


DIRECTION 44 221 


Zoology, if the oldest available name for the species concerned, 
and that in other cases the oldest available specific name for the 
species in question be so entered. The present Direction which 
is taken under another General Directive issued to the Com- 
mission by the Congress contains Rulings prescribing certain 
minor corrections to be made in the entries made on the Official 
List of Generic Names in Zoology in the above period and placing 
certain names connected with the cases referred to above on the 
Official Indexes of Rejected and Invalid Generic and Specific 
Names in Zoology. The proposals which formed the basis of the 
decisions by the Commission in the present case were submitted 
by the Secretary in the following paper on 12th October 1955 :— 


Proposals relating to various matters outstanding in connection with 
the names of genera in the Class Aves placed on the ‘‘ Official List 
of Generic Names in Zoology ”’ in the period up to the end of 
1936 


By FRANCIS HEMMING, C.M.G., C.B.E. 
(Secretary to the International Commission on Zoological Nomenclature) 


The present paper contains the third instalment of proposals for 
giving effect, so far as the names of genera of birds are concerned, to 
the General Directive, issued to the International Commission by the 
Thirteenth International Congress of Zoology, Paris, 1948, that the 
entries made on the Official List of Generic Names in Zoology in the 
period up to the end of 1936 should be completed by the addition 
thereto of all the particulars required by other Directives issued to the 
Commission by that Congress. The first instalment, which related to 
the assignment of appropriate genders to the names of genera of birds 
placed on the Official List in the foregoing period, formed the subject 
of the vote taken by the Commission on Voting Paper V.P.(O.M.)(55)16. 
The Commission’s decision on this matter has since been embodied in 
Direction 26 (now in the press).2 The second instalment, which was 
concerned with the placing on the Official List of Specific Names in 
Zoology of the specific names of the type species of the same genera, 
formed the subject of proposals submitted to the Commission with the 
Voting Paper V.P.(O.M.)(55)31 issued simultaneously with the present 
proposals. The instalment now submitted is concerned with various 
miscellaneous matters, such as the correction of minor errors in the 
entries made on the Official List by Rulings given in Opinion 67 and 
the addition to the Official Indexes of invalid generic and specific 
names involved in one way or another with the generic names placed 
on the Official List in the above Opinion. 


8 Direction 26 has since been published. See Footnote 2. 


222 OPINIONS AND DECLARATIONS 


2. First, it is necessary to take note of four errors of dating or 
citation which require now to be corrected. Particulars are given in 
Annexe 1. 


3. Second, particulars are given in Section A of Annexe 2 of certain 
invalid generic names which under Directives issued to the Commission 
by the International Congress of Zoology require now to be placed on 
the Official Index of Rejected and Invalid Generic Names in Zoology. 


4. Finally, particulars are given in Section B of Annexe 2 of certain 
invalid specific names which under the Directive referred to above 
require now to be placed on the Official Index of Rejected and Invalid 
Specific Names in Zoology. Of these names seven are names which 
were cited by the original authors as the names for type species of 
genera but which are invalid by reasons of being junior objective 
synonyms of older names. For the reasons explained in the paper 
submitted with Voting Paper V.P.(O.M.)(55)31 (paragraphs 8—9)* it 
is proposed to treat the senior objective synonym as the name of the 
type species in these cases.® The point involved is procedural only 
and is designed solely to promote clarity of presentation in the entries 
to be made on the Official List of Generic Names in Zoology. It does not 
affect in any way the names now proposed to be placed on the Official 
Index of Rejected and Invalid Specific Names, for the seven specific 
names here concerned, being objectively invalid names, would in any 
case require to be placed on that Index. The names concerned are 
marked with an asterisk in Annexe 2 (Section B). 


5. In the light of the particulars outlined above and given in full 
in the Annexes attached to the present paper, I recommend that, in 
compliance with the Directives issued to it by the International 
Congress of Zoology, the Commission should now render a Direction 
(a) correcting, in the manner recommended in Annexe 1, certain 
errors in entries made on the Official List of Generic Names in Zoology 
by the Ruling given in Opinion 67, and (b) placing on the Official 
Indexes of Rejected and Invalid Names the generic and specific names 
specified in Annexe 2. 


ANNEXE 1 


Cases where, when a generic names in the Class Aves was placed on the 
‘* Official List of Generic Names in Zoology’, (a) an incorrect 
date was cited for the generic name, (b) incorrect particulars 
were given as to the selection of the type species of the genus 
concerned, or (c) the name of the type species was 
incorrectly spelled 


(1) ‘* Alauda ’’ Linnaeus, 1758 : 
The entry in Opinion 67 regarding the type species of this genus 


4 The paper here referred to is reproduced in full in paragraph 2 of Direction 43 
(pp. 185—200 of the present volume). 

5 The proposal here referred was approved by the International Commission 
and has been embodied in Declaration 21 (1956, Ops. Decls. int. Comm. zool, 
Nomencl, 12 : i—viiti). 


DIRECTION 44 223 


was “* Alauda arvensis Linn. (Tsd. Swainson, 1827)”. This may be 
interpreted as meaning that the type species of this genus is Alauda 
arvensis Linnaeus, 1758 by selection under Rule (g) in Article 30 by 
Swainson in some book or paper published in 1827. The attribution 
of the type selection for this genus to ‘‘ Swainson, 1827 ’’ was a mistake, 
for it was anticipated by the selection of the same species to be the 
type species of this genus made by Selby two years earlier, i.e. in 1825 
Ul. Brit. Ornith. 1 (Landbirds) : xxix, xxxiii). This is the reference 
adopted for this type selection both in the A.O.U Checklist and in 
vol. 1 of the Handbook of British Birds (1938). 


(2) ‘* Fulmarus ”*’ Stephens, 1826 


The type species of this genus is Procellaria glacialis Linnaeus, 
1761, by selection by Gray (G.R.) in 1855 (Cat. Gen. Sub-Gen. Birds 
Brit. Mus. : 129). This is the reference accepted by all modern authors. 
For some reason which can only be attributed to inadvertence the 
selection of the above species as the type species of this genus was 
attributed to “‘ Palmer, 1836” in Opinion 67. This was clearly a 
mistake. (When this matter was being investigated by the late Dr. 
James L. Peters, at my request, he reported that he was not aware of any 
worker named Palmer in this field prior to 1840 and that the entry in 
Opinion 67 must be rejected as being incorrect.) 


(3) ‘* Heliornis ’’ Bonnaterre, 1790 


This generic name was published on pp. lxxxiv and 64 of the first 
part of the “‘ Ornithologie ’’ Section of the Encyclopédie methodique 
and the name applied by Bonnaterre to its type species—Heliornis 
fulicarius—was published on page 65 of the same part of the above 
work. The part concerned was published in 1790 and not in 1791, the 
date incorrectly cited in Opinion 67. 


(4) ‘* Manucodia ’’ Boddaert, 1783 


The type species of this genus was cited by Boddaert under the 
new name Manucodia chalybea. The oldest available name for the 
taxon so named is Paradisea [sic] chalybata Forster, 1781. An attempt 
was made in Opinion 67 to indicate this synonymy but unfortunately 
this specific name was there misspelled as “* chalybeata”’. 


In order to prevent anyone, following Sherborn, from falling into 
the error of concluding that the name Paradisea [sic] chalybata Forster, 
1781, is invalid by reason of being a nomen nudum, it should be noted 
that, in citing this name Forster provided it with an “‘ indication ”’ for 
the purposes of Article 25, by adding the entry ‘“‘ Pl. enlum. 634”’. 
This is a reference to the plate so numbered in the Histoire naturelle 
des Oiseaux of Daubenton (E.I.), the plates (Planches enluminées) in 
which are stated in the title to have been prepared by Martinet under 
the direction of Daubenton, 


224 OPINIONS AND DECLARATIONS 


ANNEXE 2 


Proposed addition of certain objectively invalid names to the ‘‘ Official 
Indexes of Rejected and Invalid Names ”’ 


Section A :—Proposed addition of forty-six generic names 
to the “‘ Official Index of Rejected and Invalid Generic 
Names in Zoology” 


1. Aegithia Blyth, 1860, J. asiat. Soc. Bengal 29 : 254 (an Erroneous 
Subsequent Spelling for Aegithina Vieillot, 1816, placed on the 
Official List in Opinion 67) 


2. Aix Agassiz, 1846, Nomencl. Zool. Index Univ. : 12 (emend. pro 
Aia Eyton, 1838) (a junior homonym of Aix Boie, 1828, placed 
on the Official List in Opinion 67) 


3. Alauda Daudin, 1816, Dict. Sci. nat. (Levrault) 1 : 513 (a junior 
homonym of Alauda Linnaeus, 1758, placed on the Official 
List in Opinion 67) 


4. The under-mentioned Erroneous Subsequent Spellings of 
Aptenodytes Miller, 1778, placed on the Official List in Opinion 
67 :— 


(a) Aptenodita Bonnaterre, 1791, Tabl. ency. méth. Orn. 1 
(1823) : xxxiv 
(b) Aptenodites Lesson, 1837, Compl. Buffon 9 : 542 


(c) Aptenodyta Gmelin, 1788, in Linnaeus, Syst. Nat. (ed. 
13) 1(1) : 242, 559 


5. Aptenodytes Forster (J.R.), 1777, in Forster (G.), Voy. World 
1 : 98 (a nomen nudum) 


6. Aulacorhynchus Dittmar, 1872, Verh. Russ. min. Ges. (2) 7:2 (a 
junior homonym of Aulacorhynchus Gould, 1834, placed on the 
Official List in Opinion 67) 


7. Brotogerys Agassiz, 1846, Nomencl. Zool. Index Univ. : 54 (an 
Invalid Emendation of Brotogeris Vigors, 1825, placed on the 
Official List in Opinion 67) 


8. Capito Gistl, 1848, Nat. Thierr. : xi (nom. nov. pro Ophistomis 
Dejean, 1835) (a junior homonym of Capito Vieillot, 1816, 
placed on the Official List on Opinion 67) 


9. Catharthes d’Orbigny, [1835], Voy. Amer. mérid.9:tab. 1 (an 
Erroneous Subsequent Spelling for Cathartes IUlliger, 1811, 
placed on the Official List in Opinion 67) 


10. Centrocercus Fitzinger, 1843, Syst. Rept. : 18, 86 (a junior homonym 
of Centrocercus Swainson, 1832, placed on the Official List in 
Opinion 67) 


BY. 


12. 


13. 


14. 


1d. 


16. 


ie 


18. 


19. 


20. 


et: 


DIRECTION 44 a5 


The under-mentioned junior homonyms of Cephalopterus Geoflroy 
1809, placed on the Official List in Opinion 67 :— 
(a) Cephalopterus Risso, 1810, Ichth. Nice : 14 


(b) Cephalopterus Powrie, 1870, Trans. Edinb. geol. Soc. 1 : 298 


Chauna Loew, 1847, Ent. Ztg., Stettin 8 : 370 (a junior homonym 
of Chauna Illiger, 1811, placed on the Official List in Opinion 67) 


Chrysolophus Swainson, 1837, Nat. Hist. Classif. Birds 2 : 225 (a 
junior homonym of Chrysolophus Gray, [1833—34], placed on 
the Official List in Opinion 67) 


Clamator Blyth, 1852, Cat. Birds Mus. Asiat. Soc. 1849 : 250 
(a junior homonym of Clamator Kaup, 1829, placed on the 
Official List in Opinion 67) 


Coerebra Boie, 1826, Isis (Oken) 19 : 971 (an Erroneous Subse- 
quent Spelling for Coereba Vieillot, 1816, placed on the Official 
List in Opinion 67) 


Coturnix Cuvier, 1800, Legons comp. Anat. 1:2 (a nomen nudum) 


The under-mentioned Erroneous Subsequent Spellings for 
Eurynorhynchus Nilsson, 1821, placed on the Official List in 
Opinion 67 :— 

(a) Eurynorhinchus Lafresnaye, 1842, Rev. Zool. 5 : 402 
(b) Eurynorynchus Reichenbach, [1852], Av. Syst. 24 : xiii 


Eurylaemus Gloger, [1841], Handb. Naturg. 1 (1842) : 333 (an 
Erroneous Subsequent Spelling for Eurylaimus Horsfield, 1821, 
placed on the Official List in Opinion 67) 


Eurypygia Swainson, 1837, Nat. Hist. Classif. Birds 2 : 359 (an 
Erroneous Subsequent Spelling for Eurypyga Illiger, 1811) 


Gampsonyx Jordan, 1847, Verh. nat. ver. pr. Rheinl. 4:91 (a 
junior homonym of Gampsonyx Vigors, 1825, placed on the 
Official List in Opinion 67) 


The under-mentioned Erroneous Subsequent Spellings for 
Haliaeetus Savigny, 1809, placed on the Official List in Opinion 
67 :— 

(a) Haliaethus Lesson, 1831, Traité Orn. (8) : 654 


226 


22. 


23. 


24. 


25% 


26. 


ZT 


28. 


29. 


30. 


OPINIONS AND DECLARATIONS 


(b) Haliaetos Bonaparte, 1826, Ann. Lyceum nat. Hist. N. York 
2 (1828) : 24, 25 

(c) Haliaetus Cuvier, 1816, Régn. Anim. 1 (1817) : 351; id., 
Vieillot, 1819, Nouy. Dict. Hist. nat. (ed. 2) 28 : 277 


Heliornis (Dalman MS) Billberg, 1820, Enum. Ins. Mus. Billberg : 79 
(a junior homonym of Heliornis Bonnaterre, 1790, placed on the 
Official List in Opinion 67) 

The under-mentioned Erroneous Subsequent Spellings for 
Ibidorhyncha Vigors, 1832, placed on the Official List in 
Opinion 67 :— 

(a) Ibidorhinchus David, 1875, J. trois. Voy. Chin. 1 : 307 
(b) Ibidorhynchus Gray, 1844, List Birds B.M.3 : 95, 201 
Musophagus Bonnaterre, 1790, Ency. méth., Orn. 1 : xcv (an 


Invalid Emendation of Musophaga Isert, 1789, placed on the 
Official List in Opinion 67) 


Numidia Forster, 1817, Syn. Cat. Brit. Birds : 64) (an Erroneous 
Subsequent Spelling for Numida Linnaeus, 1764, placed on the 
Official List in Opinion 67) 


The under-mentioned junior homonyms of Pachycephala Vigors, 
1825, placed on the Official List in Opinion 67 :— 


(a) Pachycephala Klug, 1834, Jahrb. Insektenk. 1 : 4 


(b) Pachycephala Doleschall, 1858, Natuurk. Tijdskr. Nederl. 
Ind. 17 : 115 


(c) Pachycephala Lioy, 1864, Atti Ist. Veneto (3) 9 : 1343 
(d) Pachycephala Broun, 1881, Man. N.Z. Col. 2 : 672 


(e) Pachycephala Fairmaire, 1883, Ann. Soc. ent. Belgique 
27(2) : 4 


Pachycephalus Stephens, 1826, in Shaw, Gen. Zool, (Aves) 13(2) : 
267 (an Invalid Emendation of Pachycephala Vigors, 1825, 
placed on the Official List in Opinion 67) 


Parotica Wagler, 1827, Syst. Av. 1 : 99 (an Erroneous Subsequent 
Spelling for Parotia Vieillot, 1816, placed on the Official List 
in Opinion 67) 


The under-mentioned junior homonyms of Pezoporus Illiger, 1811, 
placed on the Official List in Opinion 67 :— 
(a) Pezoporus Klug, [1842], Abh. preuss. Akad. Wiss. 1840 : 311 
(b) Pezoporus Foerster, 1868, Verh. naturh. Ver. Rheinlande 
= | 


Psittacus Catesby, 1777, Pisc.... Imagines : 29 (ajunior homonym of 
Psittacus Linnaeus, 1758, placed on the Official List in Opinion 67) 


af. 


a2. 


am. 


ON 


DIRECTION 44 22h 


Pteroglossus Chaudior, 1843, Bull. Soc. imp. Nat. Moscou 16(3) : 405 
(a junior homonym of Ptreroglossus Wlliger, 1811, placed on the 
Official List in Opinion 67) 


The under-mentioned Erroneous Subsequent Spellings of Priloris 
Swainson, 1825, placed on the Official List in Opinion 67 :— 
(a) Ptilorhys Bonaparte, 1854, C.R. Acad. Sci. Paris 38 : 260 
(b) Prilornis Gray (G.R.), 1869, Handlist Birds 1 : 104, 105 


Strigopsis Bonaparte, 1849, Consp. Gen. Avium:8 (an Invalid 
Emendation of Strigops Gray (G.R.), 1845, placed on the 
Official List in Opinion 67) 


Section B: Proposed addition of nine specific names to the 
** Official Index of Rejected and Invalid Specific Names 
in Zoology” 


. *argus Temminck, 1807 (Cat. Syst. Cab. Ornith. : 149) as published 


in the combination Polyplectron argus (a junior objective 
synonym of bicalcaratus Linnaeus, 1758, as published in the 
combination Pavo bicalcaratus) 


. *chalybea Boddaert, 1783 (Tabl. Pl. enl. Hist. nat. Daubenton : 39), 


as published in the combination Manucodia chalybea (a junior 
objective synonym of chalybata Forster, 1781, as published in 
the combination Paradisea [sic] chalybata) 


. chalybeata Stiles, 1916 (Smithson. misc. Publ. 2409 : 181), as 


published in the combination Paradisea chalybeata (an Erroneous 
Subsequent Spelling for chalybata Forster, 1781, as published 
in the combination Paradisea [sic] chalybata) 


. *communis Bonnaterre, 1791 (Ency. méth., Orn. 1: 217), as 


published in the combination Coturnix communis (a junior 
objective synonym of coturnix Linnaeus, 1758, as published 
in the combination Tetrao coturnix) 


. *fluvialis Savigny, 1809 (Descr. Egypte 1(1) (Ois.) : 96), as pub- 


lished in the combination Pandion fluvialis (a junior objective 
synonym of haliaetus Linnaeus, 1758, as published in the 
combination Falco haliaetus) 


. formosus Latham, 1790 (Index ornith. : 103), as published in the 


combination Psittacus formosus (a junior primary homonym of 
formosus Scopoli, 1769, as published in the combination 
Psittacus formosus) 


*See paragraph 4 of the present paper. 


228 OPINIONS AND DECLARATIONS 


7. *fulicarius Bonnaterre, 1790 (Ency. méth., Orn. 1 : 65), as pub- 
lished in the combination Heliornis fulicarius (a junior objective 
synonym of fulica Boddaert, 1783, as published in the com- 
bination Colymbus fulica) 


8. *galbula Linnaeus, 1766 (Syst. Nat. (ed. 12) 1(1) : 160), as pub- 
lished in the combination Oriolus galbula (a junior objective 
synonym of oriolus Linnaeus, 1758, as published in the com- 
bination Coracias oriolus) 


9. *griseus Nilsson, 1821 (Ornith. svec. 2(1) : 29), as published in the 
combination Eurynorhynchus griseus (a junior objective synonym 
of pygmaea Linnaeus, 1758, as published in the combination 
Platalea pygmaea). 


2. Registration of the present application : Upon the receipt of 
Mr. Hemming’s paper the question of the action required to 
settle the questions raised therein was allotted the Registered 
Number Z.N.(S.) 994. 


Il. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


3. Issue of Voting Paper V.P.(O.M.)(55)32 : On 12th October 
1955 a Voting Paper (V.P.(O.M.)(55)32) was issued in which 
each Member of the Commission was asked (1) to state whether 
he agreed that, “‘ in conformity with the General Directives issued 
to the International Commission on Zoological Nomenclature 
by the Thirteenth International Congress of Zoology, Paris, 1948, 
regarding the placing on the appropriate Official Lists and 
Official Indexes of names dealt with in, or otherwise involved 
in, Opinions rendered prior to 1948, and other matters on which 
action is required in order to complete, to clarify, or to correct 
Rulings given in Opinions rendered in the foregoing period, the 
action recommended in paragraph 5 of the paper numbered 
Z.N.(S.) 994, regarding names in the Class Aves submitted by the 
Secretary simultaneously with the present Voting Paper [l.e. 


* See paragraph 4 of the present paper. 


DIRECTION 44 229 


in the paragraph numbered as above in the application repro- 
duced in the first paragraph of the present Direction] be now 
taken as there recommended ”’ and (2), if he did not so agree as 
regards any given item, to indicate that item. 


4. The Prescribed Voting Period: As the foregoing Voting 
Paper was issued under the One-Month Rule, the Prescribed 
Voting Period closed on 12th November 1955. 


5. Particulars of the Voting on Voting Paper V.P.(O.M.)(55)32 : 
At the close of the Prescribed Voting Period, the state of the 
voting on Voting Paper V.P.(O.M.)(55)32 was as follows :— 


(a) Affirmative Votes had been given by the following twenty-four 
(24) Commissioners (arranged in the order in which Votes 
were received) : 


Hering ; Holthuis ; Lemche ; Vokes ; Esaki ; Boschma ; 
Riley; Miller; Bradley (J.C.); Sylvester-Bradley ; 
Prantl; Mayr; Jaczewski; Tortonese; do Amaral ; 
Dymond; Hemming; Bonnet; Mertens; Stoll; 
Cabrera; Key; Kihnelt ; Hanko ; 


(b) Negative Votes : 


None ; 


(c) On Leave of Absence, one (1) : 


Bodenheimer ; 


(d) Voting Papers not returned : 


None. 


230 OPINIONS AND DECLARATIONS 


6. Declaration of Result of Vote : On 21st November 1955, 
Mr. Hemming, Secretary to the International Commission, acting 
as Returning Officer for the Vote taken on Voting Paper V.P. 
(O.M.)(55)32, signed a Certificate that the Votes cast were as 
set out in paragraph 5 above and declaring that the proposal 
submitted in the foregoing Voting Paper had been duly adopted 
and that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


-7, Addition of four further names to the ‘‘ Official Index of 
Rejected and Invalid Generic Names in Zoology ”’ : On 2\st 
November 1955 Mr. Hemming, as Secretary, executed the 
following Minute directing the addition to the Official Index of 
Rejected and Invalid Generic Names in Zoology of three further 
Erroneous Subsequent Spellings and one further Invalid Emend- 
ation of generic names placed on the Official List of Generic 
Names in Zoology in the period up to the close of 1936 :-— 


Addition of four further invalid generic names to the ‘‘ Official Index 
of Rejected and Invalid Generic Names in Zoology fs 


MINUTE by FRANCIS HEMMING, C.M.G., C.B.E. 
(Secretary to the International Commission on Zoological Nomenclature) 


I have received from Professor Ernst Mayr a letter dated 26th 
October 1955 in which he draws attention to the fact that the three 
Erroneous Subsequent Spellings and the one Invalid Emendation 
specified below of the names of genera in the Class Aves placed on the 
Official List of Generic Names in Zoology in the period up to the end of 
1936 were inadvertently omitted from the list of invalid generic names 
which in paper Z.N.(S.) 994 submitted to the Commission with Voting 
Paper V.P.(O.M.)(55)32 were recommended for addition to the 
Official Index of Rejected and Invalid Generic Names in Zoology :— 


(1) Stringops Finsch, 1867, Papageien 1 : 233, 241 (an Erroneous 
Subsequent Spelling for Strigops Gray (G.R.), 1845, placed 
on the Official List by the Ruling given in Opinion 67) ; 


(2) Stringopsis van der Hoeven, 1855, Handb. Dierk. (ed. 2) 2 : 692 
(an Erroneous Subsequent Spelling for Strigops Gray (G.R.), 
1845) ; 


DIRECTION 44 231 


(3) Strygops Des Murs, [1853], in Chenu, Ency. Hist. nat., Ois. 
1: 202 (an Erroneous Subsequent Spelling for Strigops 
Gray (G.R.), 1845) ; 


(4) Ptilorhis Agassiz (J.L.R.), 1846, Nomencl. zool. Index univ. (an 
Invalid Emendation of Priloris Swainson, 1825, placed on the 
Official List by the Ruling given in Opinion 67). 


2. In the light of the additional information received in this matter 
I, as Secretary to the International Commission, hereby direct that 
the names specified above be placed on the Official Index of Rejected 
and Invalid Generic Names in Zoology at the same time that the names 
specified in the paper referred to in paragraph 1 above are so placed. 


8. Preparation of the Ruling given in the present °° Direction ”’ : 
On 7th February 1956, Mr. Hemming prepared the Ruling 
given in the present Direction and at the same time signed a 
Certificate that the terms of that Ruling were in complete accord 
with those of the proposal approved by the International Com- 
mission in its Vote on Voting Paper V.P.(O.M.)(55)32, subject to 
the adjustment specified in the Minute executed by the Secretary 
on 21st November 1955, the text of which has been reproduced in 
paragraph 7 of the present Direction. 


9. Original References : The original references for the names 
specified in Annexe 2 to the present application which have been 
placed on the Official Index of Rejected and Invalid Generic Names 
in Zoology or, as the case may be, on the Official Index of Rejected 
and Invalid Specific Names in Zoology by the Ruling given in the 
present Direction are as set out in the Annexe referred to above. 
The original references for the four generic names placed on the 
Official Index by the direction given in the Secretary’s Minute 
of 21st November 1955 are as set out in that Minute. 


10. Family-Group Names: No family-group-name problem 
arises in connection with any of the names dealt with in the Ruling 
given in the present Direction. 


11. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in dealing 


PE OPINIONS AND DECLARATIONS 


with the present case, and the present Direction is accordingly 
hereby rendered in the name of the said International Commission 
by the under-signed Francis Hemming, Secretary to the Inter- 
national Commission on Zoological Nomenclature, in virtue 
of all and every the powers conferred upon him in that behalf. 


12. The present Direction shall be known as Direction Forty- 
Four (44) of the International Commission on Zoological 
Nomenclature. 


DoNE in London, this Twenty-Seventh day of February, 
Nineteen Hundred and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


—— 


Printed in England by MercaLFe & Cooper LimiTeED, 10-24 Scrutton St., London EC 2 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 


NATIONAL COMMISSION ON 
LOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, cM.G., c.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.9. Pp. 233—242 


DIRECTION 45 


Substitution of Pinnotheres Bosc, [1801—1802] for 

Pinnotheres Latreille, [1802—1803] (Class Crustacea, 

Order Decapoda) on the Official List of Generic Names 

in Zoology (correction of an error in the Ruling given 
in Opinion 85) 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 


1956 ae 
Ps 7 i-4 & / [ —— 
Price Seven Shillings ZO a eed! y 
fa | 
(All rights reserved) \ NOV 20 1955 | 


Issued 19th October, 1956 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 45 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JoRDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President : Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) 
(12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(arranged in order of precedence by reference to date of election or of most recent re-election 
as prescribed by the International Congress of Zoology) 


Professor H. BOSCHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. ies LEMCHE (Universitetets Zoologiske Museum, Copenhagen, Denmark) (27th 
July 1948 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEy (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (Sth July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitadt zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMOND (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) (12th August 
1953) (President) 

Professor Harold E. Vokes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) (12th 
August 1953) 

Professor Béla HANKO (Mezdgazdasdgi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.-Y., 
U.S.A.) (12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. Hottuuis (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
oes August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (15th October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum V Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KUHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

Professor F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 


) 
Professor Ernest MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 
Professor Enrico TORTONESE (Museo di Storia Naturale ‘‘ G. Doria’’, Genova, Italy) 
(16th December 1954) 


DIRECTION 45 


SUBSTITUTION OF “‘ PINNOTHERES ”’ BOSC, [1801—1802] 
FOR ‘“PINNOTHERES ” LATREILLE, [1802—1803] 
(CLASS CRUSTACEA, ORDER DECAPODA) ON 
THE “ OFFICIAL LIST OF GENERIC NAMES 
IN ZOOLOGY ” (CORRECTION OF AN 
ERROR IN THE RULING GIVEN IN 
‘‘ OPINION ”° 85) 


RULING :—(1) The incorrect entry relating to the 
generic name VPinnotheres Latreille made on _ the 
Official List of Generic Names in Zoology by the Ruling 
given in Opinion 85, in which also the date [1801—1802] 
was erroneously attributed to the foregoing name, is 
hereby deleted and the following revised entry is hereby 
mserted in its place under the same Name Number 
(Name No. 352) :— 


Pinnotheres Bosc, [1801—1802], Hist. nat. Crust. 1 : 59, 
239 (gender : masculine) (type species, by selection 
by Latreille (1810, Consid. gén. Anim. Crust. Arachn. 
Ins. : 422) : Cancer pisum Linnaeus, 1767, Syst. Nat. 
fed ta) (2): = 1039). 


(2) The under-mentioned specific name is hereby placed 
on the Official List of Specific Names in Zoology with the 


NOV 7 4 1956 


236 OPINIONS AND DECLARATIONS 


Name No. 1035: pisum Linnaeus, 1767, as published in 
the combination Cancer pisum (specific name of type 
species of Pinnotheres Bosc, [1801—1802}). 


(3) The under-mentioned generic name is hereby placed 
on the Official Index of Rejected and Invalid Generic Names 
in Zoology with the Name No. 577 : Pinnotheres Latreille, 
[1802—1803] (a junior homonym of Pinnotheres Bosc, 
[1801—1802]). 


I. THE SUBJECT MATTER OF THE; PRESENT 
* DIRECTION 


On 19th September 1955, Mr. Hemming as Secretary, submitted 
to the International Commission on Zoological Nomenclature 
two papers, each dealing with a number of matters relating to 
entries of names of genera of the Order Decapoda (Class 
Crustacea) made on the Official List of Generic Names in Zoology 
in the period up to the end of 1936 as regards which it appeared 
that action was needed in order to enable the Commission to 
comply with the General Directive relating to the completion 
of entries made on the above List in the foregoing period issued 
by the Thirteenth International Congress of Zoology, Paris, 
1948. The first of these papers, with which was associated 
Voting Paper V.P.(O.M.)(55)27!, bore the Registered Number — 
Z.N.(S.) 985 ; the second, with which was associated Voting Paper 
V.P.(O.M.)(55)28!, bore the Registered Number Z.N.(S.) 987. 
During the Prescribed Voting Period for the foregoing Voting 
Papers information received from Commissioner L. B. Holthuis 
(Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
disclosed the existence of a previously undetected problem in 
relation to the generic name Pinnotheres which, attributed to 
Latreille and dated ‘“‘ 1801—1802”, had been placed on the 
Official List by the Ruling given in Opinion 85. In the light of 
the information so received, Mr. Hemming, on 7th October 1955, - 


1 The decisions taken by the International Commission on these two Voting 
Papers have since been embodied in Direction 36 (1956, Ops. Decls. int. Comm. 
zool. Nomencl. 1(D) : 1—46) and Direction 37 (1956, ibid. 1(D) : 47—82) 
respectively. 


DIRECTION 45 237 


executed Minutes on the Commission’s Files Z.N.(S.) 985 and 
Z.N.(S.) 987 respectively withdrawing from the scope of the 
foregoing Voting Papers the proposals in relation to the generic 
name Pinnotheres Latreille submitted therewith and directing 
that the situation be reviewed in the light of the additional 
information received with a view to the submission of revised 
proposals to the Commission as soon as possible?. On 27th 
October 1955 the following paper containing revised proposals 
was submitted to the Commission by the Secretary :— 


Proposed correction of the entry on the ‘‘ Official List of Generic 
Names in Zoology ”’’ relating to the generic name ‘‘ Pinnotheres ”’ 
Latreille, [1802—1803] (Class Crustacea, Order Decapoda) 
(correction of an error in ‘‘ Opinion ”’ 85) 


By FRANCIS HEMMING, C.M.G., C.B.E. 
(Secretary to the International Commission on Zoological Nomenclature) 


The object of the present paper is to call attention to an incorrect 
entry in relation to the generic name Pinnotheres Latreille, [1802—1803], 
made on the Official List of Generic Names in Zoology by the Ruling 
given in the Commission’s Opinion 85 published in 1925 (Smithson. 
misc. Coll. 73 (No. 3) : 13—18) and to submit proposals for correcting 
the entry so made. 


2. The problem now to be considered was brought to light about a 
month ago by Dr. L. B. Holthuis (Rijksmuseum van Natuurlijke 
Historie, Leiden, The Netherlands) by whom it was encountered 
incidentally when considering proposals which I had then submitted 
with Voting Papers V.P.(O.M.)(55)27 and 28 for completing the entries 
on the Official List of Generic Names in Zoology made in the pre-Lisbon 
(1935) period. These proposals had been prepared for the purpose 
of placing before the Commission the material necessary to enable it 
to comply with the General Directives issued by the International 
Congress of Zoology that such additions be made to entries made on the 
Official List in the foregoing period as might be necessary to bring 
those entries into line with those made in regard to names similarly 
placed on the above Official List in more recent periods. On receiving 
Dr. Holthuis’s communication of 20th September 1955, I at once 
executed a Minute withdrawing the name Pinnotheres Latreille from 
the scope of the above Voting Papers. 


2 The Minute placed by the Secretary on Commission File Z.N.(S.) 985 on 
7th October 1955 has been reproduced in paragraph 4 of Direction 36 (published 
as Part | of the present Section (SECTION D) of the present volume (Volume 
1), pp. 1—46). The corresponding Minute placed by the Secretary on Com- 
mission File Z.N.(S.) 987 on the same day has been reproduced in paragraph 4 
of Direction 37 (published as Part 2 of the present Section, pp. 47—82). 


238 OPINIONS AND DECLARATIONS 


3. In Opinion 85 the name Pinnotheres Latreille was stated to have 
been published in “‘ 1801-2 (An X)”’ in volume 3 of the Hist. nat. gén. 
partic. Crust. Ins. in Sonnini’s Buffon. It was further stated that 
Cancer pisum Linnaeus, 1767 (Syst. Nat. (ed. 12) 1(2) : 1039) was the 
type species of this genus. 


4. In the first of the Voting Papers cited above (V.P.(O.M.)(55)27) 
a recommendation was submitted that the specific name pisum 
Linnaeus, 1767, as published in the combination Cancer pisum, should, 
as the specific name of the type species of a genus (Pinnotheres Latreille), 
which had been placed on the Official List of Generic Names in Zoology, 
now be placed on the Official List of Specific Names in Zoology. 


5. Nothing was said in the Ruling given in Opinion 85 as to the 
manner in which Cancer pisum Linnaeus, 1767, had become the type 
species of the genus Pinnotheres Latreille. In order to repair this 
omission, it was recommended in Annexe | to the paper? submitted 
with the second of the Voting Papers referred to above (Voting Paper 
V.P.(O.M.)(55)28), that the entry “‘ type species, by monotypy’’ be 
inserted in the entry in the Official List relating to this generic name, 
reference to the work cited in Opinion 85 having shown that the above 
was the only nominal species cited by Latreille when in that work he 
established the genus Pinnotheres. Further, in Appendix 2 (para- 
graph 10) of the same paper® attention was drawn to the fact that, 
although, as stated in Opinion 85, the third volume of the Hist. nat. 
gén. partic. Crust. Ins. bore the date “‘An X ”’ calculated in accordance 
with the French Revolutionary Calendar of a hundred and fifty years 
ago, there is evidence to show that this volume cannot have been 
published until some time in “‘An X1”’, ie. in the period from 
September 1802 to September 1803 (Hemming, 1946, Bull. zool. 
Nomencl. 1 : 205, nota 7). 


6. In a letter dated 20th September 1955, Dr. Holthuis informed 
me that he had ascertained that the name Pinnotheres was first published 
not by Latreille but by Bosc in the latter author’s Hist. nat. Crust.1 : 59, 
239, published in “‘An X”’, 1.e. in the period September 1801 to Sep- 
tember 1802. Bosc included in his genus Pinnotheres the nominal 
species Cancer pisum Linnaeus, 1758, and that species was selected as 
the type species by Latreille himself in 1810 (Consid. gén. Anim. Crust. 
Arach. Ins. : 422). Dr. Holthuis added that Bosc attributed (: 239) 
the generic name Pinnotheres to Latreille who was undoubtedly the 
author of this name in manuscript and suggested that Bosc became 
aware of it through the action of Latreille who, as Bosc explained in 
his preface (loc. cit. 1 : 49) had given him permission to make use 
(‘‘... ila permis de faire usage ici ’’) of the text of the then unpublished 
volume 3 of Latreille’s Hist. nat. gén. partic. Crust. Ins., referred to 
by Bosc as the forthcoming ‘‘ nouvelle edition” of the Précis des 


8’ The document here referred to has been reproduced in full in paragraph 1 of 
Direction 37 (pp. 63—78 of the present volume). 


DIRECTION 45 239 


caracteres des géneriques [sic] des Insectes of the “‘ savant auteur... 
Vestimable Latreille ”’. 


7. In the light of the evidence now furnished by Dr. Holthuis it is 
evident that the entry relating to the name Pinnotheres Latreille, 
[1802—1803], must be removed from the Official List and replaced 
by an entry relating to the name Pinnotheres Bosc, [1801—1802]. 
Fortunately, no change of type species is involved, for Cancer pisum 
Linnaeus, 1767, is the type species alike of Pinnotheres Bosc and of the 
later name Pinnotheres Latreille, though in the first of these cases that 
species is type species by selection by Latreille, 1810, while in the 
second case it is type species by monotypy. 


8. In order to remedy the defect in Opinion 85 discovered by 
Dr. Holthuis, I recommend the International Commission :— 


(1) to delete from the Official List of Generic Names in Zoology 
the incorrect entry of the name Pinnotheres Latreille, [1802— 
1803], made therein, with the incorrect date ‘* 1801—1802’’, 
by the Ruling given in Opinion 85 and to substitute therefor 
the following revised entry under the same Name Number 
(Name No. 352) :— 

Pinnotheres Bosc, [1801—1802] (gender : masculine) (type 
species by selection by Latreille (1810): Cancer pisum 
Linnaeus, 1767) 

(2) to place the under-mentioned specific name on the Official List 
of Specific Names in Zoology :—pisum Linnaeus, 1767, as 
published in the combination Cancer pisum (specific name of 
type species of Pinnotheres Bosc, [1801—1802]) ; 

(3) to place the under-mentioned generic name on the Official Index 
of Rejected and Invalid Generic Names in Zoology :—Pinnotheres 
Latreille, [1802—1803] (a junior homonym of Pinnotheres 
Bosc, [1801—1802]). 


2. Registration of the present application :—In accordance 
with the directions given in the Minute placed by the Secretary 
on Commission File Z.N.(S.)985 on 7th October 1955, the 
question dealt with in the present Direction was investigated on 
Commission File Z.N.(S.) 1029, which was then opened for this 


purpose. 


Il. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


3. Issue of Voting Paper V.P.(O.M.)(55)37 : On 27th October 
1955, a Voting Paper (V.P.(O.M.)(55)37) was issued in which 


240 OPINIONS AND DECLARATIONS 


the Members of the Commission were invited to vote either for, 
or against, “‘ the proposed correction of the entry on the Official 
List of Generic Names in Zoology regardingthe name Pinnotheres 
Latreille made by the Ruling given in Opinion 85, as recommended 
in paragraph 8 of the paper bearing the Number Z.N.(S.) 1029 
submitted by the Secretary simultaneously with the present 
Voting Paper ”’ [i.e. in the paragraph numbered as above in the 
paper reproduced in the first paragraph of the present Direction]. 


4. The Prescribed Voting Period: As the foregoing Voting 
Paper was issued under the One-Month Rule, the Prescribed 
Voting Period closed on 27th November 1955. 


5. Particulars of the Voting on Voting Paper V.P.(O.M.)(55) 37 : 
At the close of the Prescribed Voting Period, the state of the 
voting on Voting Paper V.P.(O.M.)(55)37 was as follows :— 


(a) Affirmative Votes had been given by the following twenty- 
four (24) Commissioners (arranged in the order in which 
Votes were received) : 


Riley ; Holthuis; Bonnet; Hering; Mertens; Stoll ; 
Vokes; Boschma; Bodenheimer; Mayr; Esaki; 
Dymond ; Bradley (J.C.) ; do Amaral ; Hank6 ; Cabrera ; 
Jaczewski; Kiuhnelt ; Hemming; Tortonese ; Prantl ; 
Sylvester-Bradley ; Miller* ; Lemche ; 


(b) Negative Votes, one (1): 


Key ; 


(c) Voting Papers not returned : 


None. 


* Commissioner Miller exercised in this case the right conferred by the Thirteenth 
International Congress of Zoology in Paris, 1948, under which a Commissioner 
may, if he so desires, signify his willingness to support the view or the majority 
or of other members of the Commission (1950, Bull. zool. Nomencl. 4 : 50— 
52); 


DIRECTION 45 241 


6. Declaration of Result of Vote : On 17th December 1955, 
Mr. Hemming, Secretary to the International Commission, acting 
as Returning Officer for the Vote taken on Voting Paper 
V.P.(O.M.)(55)37, signed a Certificate that the Votes cast were 
as set out in paragraph 5 above and declaring that the proposal 
submitted in the foregoing Voting Paper had been duly adopted 
and that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


7. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 17th March 1956, Mr. Hemming prepared the Ruling given 
in the present Direction and at the same time signed a Certificate 
that the terms of that Ruling were in complete accord with those 
of the proposal approved by the International Commission in 
its Vote on Voting Paper V.P.(O.M.)(55)37. 


8. Original References : The following are the original refer- 
ences for the names placed on Official Lists and Official Indexes 
by the Ruling given in the present Direction :— 


Pinnotheres Bosc, [1801—1802], Hist. nat. Crust. 1 : 59, 239 

Pinnotheres Latreille, [1802—1803], Hist. nat. gén. partic. Crust. 
dase 3 25 

pisum, Cancer, Linnaeus, 1767, Syst. Nat. (ed. 12) 1(2) : 1039 


9. The reference for the selection of a type species for the 
genus Pinnotheres Bosc, [1801—1802] is as specified in Part (1) 
of the Ruling given in the present Direction. 


10. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in dealing 
with the present case, and the present Direction is accordingly 
hereby rendered in the name of the said International Commission 
by the under-signed Francis Hemming, Secretary to the Inter- 
national Commission on Zoological Nomenclature, in virtue of 
all and every the powers conferred upon him in that behalf. 


242 OPINIONS AND DECLARATIONS 
11. The present Direction shall be known as Direction Forty- 


Five (45) of the International Commission on Zoological 
Nomenclature. 


Done in London, this Seventeenth day of March, Nineteen 
Hundred and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


Printed in England by Metcatre & Cooper LimitTeD, 10-24 Scrutton St., London EC 2 


= 


Issued 26th October, 1956. 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, c.M.G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.10. Pp. 243—254 


DIRECTION 47 


Substitution on the Official List of Generic Names in 

Zoology of a revised entry relating to the generic name 

Pandalus Leach, 1815 (Class Crustacea, Order Decapoda) 
(revision of a Ruling given in Opinion 104) 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 


1956 : 
Price Eight Shillings “ e a \ \iqoOsy 
(All rights reserved) ( i Dp Pane 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 47 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JoRDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President: Professor James ‘Chester BRADLEY (Cornell University, Ithaca, N.Y., 
U.S.A.) (12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(Arranged in order of precedence by reference to date of election or of most recent 
re-election, as prescribed by the International Congress of Zoology) 


Professor H. BoscHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(1st January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. Henning LEMcHE (Universitetets Zoologiske Museum, Copenhagen, Denmark) 
(27th July 1948) 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEy (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZESWKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (Sth July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitdt zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMOND (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N. Y., U.S.A.) (12th August 1953) 
(President) 

Professor Harold E. Voxes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) 
(12th August 1953) 

Professor Béla HANKO (Mezdgazdasdgi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y., U.S.A.) 
(12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. Hottnurs (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (15th October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, California, 
U.S.A.) (29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum v Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KUHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

Professor F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 


Professor Ernst MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico TORTENESE (Museo di Storia Naturale “‘ G. Doria’’, Genova, Italy) 
(16th December 1954) 


DIRECTION 47 


SUBSTITUTION ON THE ‘“ OFFICIAL LIST OF GENERIC 
NAMES IN ZOOLOGY ” OF A REVISED ENTRY 
RELATING TO THE GENERIC NAME ‘‘ PANDALUS ”’ 
LEACH, 1815 (CLASS CRUSTACEA, ORDER 
DECAPODA) (REVISION OF A RULING GIVEN 
IN ** OPINION ” 104) 


RULING :—(1) The entry on the Official List of 
Generic Names in Zoology relating to the generic name 
Pandalus Leach, 1815 (Class Crustacea, Order Decapoda), 
made by the Ruling given in Opinion 104 is hereby deleted 
and the following revised entry inserted in its place :— 


497 Pandalus [Leach], [1814] (gender: masculine) 
(type species by monotypy : Pandalus montagui 
[Leach], [1814]). 


(2) The under-mentioned specific name is hereby 
placed on the Official List of Specific Names in Zoology 
with the Name Number 1039 : montagui [Leach], [1814], 
as published in the combination Pandalus montagui 
(specific name of type species of Pandalus |Leach], 
[1814]). 


(3) The under-mentioned generic names, each of which 
is either a junior homonym of Pandalus [Leach], [1814], or 
a later usage of that name, are hereby placed on the 
Official Index of Rejected and Invalid Generic Names in 


246 OPINIONS AND DECLARATIONS 


Zoology with the Name Numbers 730 and 731 re- 
spectively :— 


(a) Pandalus Leach, [1815], Malac. Podophth. Brit. 
(b) Pandalus Leach, [1815], Trans. linn. Soc. Lond. 


(4) The under-mentioned specific name is hereby 
placed on the Official Index of Rejected and Invalid 
Specific Names in Zoology with the Name Number 356 : 
annulicornis Leach, [1815], as published in the combination 
Pandalus annulicornis (a junior objective synonym of 
montagui [Leach], [1814], as published in the combination 
Pandalus montagui, through the lectotype selection by 
Holthuis (L.B.) made in the note attached as an Annexe 
to the paper reproduced in paragraph | of the present 
Direction). 


L.. THE SUBJECT MATTER OF THE PRESENT 
“DIRECTION 


The present Direction is concerned with the correction of an 
erroneous entry on the Official List of Generic Names in Zoology 
relating to the generic name Pandalus Leach there attributed to 
the year “1815” (Class Crustacea, Order Decapoda) made by 
the Ruling given in Opinion 104 (1928, Smithson. misc. Coll. 73 


DIRECTION 47 247 


(No. 5) : 25—28). The problem involved came to light in the 
course of the preparations for the publication of the foregoing’ 
Official List in book form. It was laid before the International 
Commission by the Secretary in the following Report on 13th 
December 1955 :— 


Revised proposals relating to the entry on the ‘* Official List of Generic 
Names in Zoology ’’ regarding the name ‘‘ Pandalus ’’ Leach, 
[1815] (Class Crustacea, Order Decapoda) (proposed correction 
of an erroneous Ruling given in ‘‘ Opinion ”’ 104) 


By FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


The object of the present note is to report to the Commission 
certain additional information regarding the generic name Pandalus 
Leach, [1815], in the Order Decapoda (Class Crustacea) which was 
placed on the Official List of Generic Names in Zoology in the pre- 
Lisbon (1935) period. In Voting Papers V.P.(O.M.)27 and 281 issued 
on 19th September 1955, I submitted recommendations for the 
completion of the entry made on the Official List in regard to this 
name and the problem now to be considered came to light when 
Commissioner L. B. Holthuis (Rijksmuseum van Natuurlijke Historie) 
returned his completed copies of the above Voting Papers. Upon 
receipt of his communication I executed a Minute withdrawing from 
the scope of those Voting Papers the recommendations submitted in 
connection with the foregoing name, in order to allow further study 
of the problems involved, with a view to the submission of revised 
proposals to the Commission as soon as possible. 


2. The name Pandalus Leach, [1815], was placed on the Official List 
of Generic Names in Zoology by the Ruling given in Opinion 104 


1 The decisions taken by the International Commission on Voting Papers 
V.P.(O.M.)(55)27 and 28 have since been embodied in Directions 36 and 37 
respectively. 


B§imir «a fn 4IAkeSe 


248 OPINIONS AND DECLARATIONS 


(published in 1928). The type species of this genus was there stated 
to be “ annulicornis ”’, i.e. Pandalus annulicornis Leach, [1815] (Malac. 
Podophth. Brit. (2) : expl. pl. 40) by monotypy. The investigations 
carried out in this Office had shown that the name annulicornis Leach, 
[1815], was apparently an available name and that it was the oldest 
such name for the species concerned. Accordingly in the paper 
submitted with Voting Paper V.P.(O.M.)27 it was recommended 
that in accordance with the General Directives issued to the 
Commission by the International Congress of Zoology the foregoing 
name, as the name of the type species of a genus, the name of which 
had already been placed on the Official List of Generic Names in Zoology, 
should in its turn be placed on the Official List of Specific Names in 
Zoology. In the paper submitted in connection with Voting Paper 
V.P.(O.M.)(55)28 attention was drawn to the fact that the name 
Pandalus was published twice by Leach in the year 1815, that the 
first of these occasions was (as noted above) in the Malac. Podophth. 
Brit., but that in Opinion 104 it was the second of these references 
(Trans. linn. Soc. Lond. 11(2) : 346) which was cited as the place 
where this name was first published. It was proposed in that paper 
that the reference in the Official List for the name Pandalus should be 
altered accordingly. 


3. Ina letter dated 20th September 1955 Dr. Holthuis, after referring 
to the citation in the paper submitted with V.P.(O.M.)(55)28 of the 
reference to Leach’s publication of the name Pandalus in 1815 in his 
Malac. Podophth. Brit., went on as follows :— 


I find that Leach, 1814 (Edinburgh. Ency.'7(2) : 432) had already 
published the name Pandalus. The type species of Pandalus 
Leach, 1814, by monotypy is Pandalus montagui Leach, 1814 
(ibid. 7(2) : 432). Pandalus montagui Leach, 1814, and Pandalus 
annulicornis Leach, 1815, are synonymous, the former being the 
name currently in use. 


4. It was on the receipt of the foregoing letter that I withdrew 
the proposals relating to the name Pandalus Leach which I had 
submitted with the Voting Papers referred to above. At the same 
time I asked Dr. Holthuis to inform me whether the names montagui 
[Leach], [1814], and annulicornis Leach, [1815], were objective or sub- 
jective synonyms of one another. In his reply dated 15th October 1955 
Dr. Holthuis provided the following interesting information :— 


(a) When in 1814 Leach published (in the Edinburgh Ency.) his 
description of Pandalus montagui, he stated that this species 
was “‘named in honour of the first discoverer, Montagu, by 
whom it was called [presumably in MS.] Astacus maculatus. 


The Rev. J. Fleming took this species in Zetland ”’. 


DIRECTION 47 249 


(b) In his description of Pandalus annulicornis in the Malac. Podophth. 
Brit. published a year later (in 1815) Leach wrote as follows : 
“This highly interesting species was discovered in Zetland, 
and in St. Andrew’s Bay, Scotland, by the Rev. Dr. Fleming, 
who most kindly gave me the specimens I originally described 
in the Edinburgh Encyclopaedia’. 


Dr. Holthuis went on to say that, so far as he was aware, no lectotype 
had ever been selected either for the nominal species Pandalus montagui 
[Leach], [1814], or for the nominal species Pandalus annulicornis Leach, 
[1815], and that, if a specimen collected by Fleming in Zetland were to 
be selected as the lectotype of both species, the foregoing names 
would become objective (instead of only subjective) synonyms of one 
another. Dr.Holthuis then noted that in White’s (1850) List of the 
Specimens of British Animals in the Collection of the British Museum, 
Part IV Crustacea, the last line on page 41 contained the following 
entry under the name “ Pandalus Montagui. Ringed-horned Shrimp ”’: 
“d. Scotland, Zetland. Presented by Dr. Fleming’’. Dr. Holthuis 
added that this was evidently the specimen mentioned by Leach. 


5. On the receipt of Dr. Holthuis’s letter I at once asked Dr. Isobel 
Gordon whether the specimen recorded in White’s List was still in the 
British Museum Collection ; at the same time I asked her views on the 
question whether the Zetland specimen, if still extant, could suitably 
be selected as the lectotype. In her reply dated Ist November 1955, 
after explaining that in the Museum’s dry collection there were now 
only three of the specimens listed by White in 1850 (: 41), of which 
the first two (Nos. 267a and 267c) came from Norfolk and Devon 
respectively and the third numbered “‘ 267d ”’ was labelled ‘* Zetland, 
Scotland, Dr. Fleming ’’, Dr. Gordon proceeded as follows :— 


Specimen 267d is a large almost complete specimen, very like 
Leach’s 1815 figure, Pl. XI, No. 1. In fact, it might be the figured 
specimen mentioned by Leach in 1814 (referring to his plate as 
“Pandalus A’’, since it was not yet numbered). No other 
Fleming specimens are extant. This 267d specimen I should 
certainly select as the lectotype since Leach’s figures are natural 
size and this specimen agrees so well with his figure ; another 
specimen was evidently dissected in order to provide the separate 
appendages illustrated as Plate XI, figs. 2—14, but of course it 
cannot be traced. 


6. On receipt of Dr. Gordon’s letter quoted above, I communicated 
a copy to Dr. Holthuis who on 15th November 1955 addressed to me a 
letter, of which a copy is given in the Annexe to the present paper, in 
which he formally selected the Fleming specimen numbered “‘ 267d ” 


250 OPINIONS AND DECLARATIONS 


now in the dry collection of the British Museum (Natural History) to 
be the lectotype (a) of Pandalus montagui [Leach], [1814], and (b) of 
Pandalus annulicornis Leach, [1815]. 


7. The foregoing action by Dr. Holthuis has been communicated 
by this Office to Dr. Gordon with a request that the specimen referred 
to above may be labelled as the lectotype of the two foregoing nominal 
species by selection by Dr. Holthuis, 15th November 1955, it being 
agreed that as soon as Dr. Holthuis’s letter containing the above 
lectotype selection is published by the International Trust, the relevant 
bibliographical reference will be communicated to Dr. Gordon, so 
that this also may be inscribed on the label belonging to the above 
specimen. 


8. The action taken by Dr. Holthuis on the basis of the information 
kindly furnished by Dr. Gordon now makes possible the submission to 
the International Commission of the following revised proposal 
relating to the generic name Pandalus Leach, namely :— 


(1) that the entry on the Official List of Generic Names in Zoology 
relating to the generic name Pandalus Leach, 1815, made by 
the Ruling given in Opinion 104, be deleted therefrom and the 
following revised entry inserted in its place :— 


497. Pandalus [Leach], [1814] (gender : masculine) (type species, 
by monotypy : Pandalus montagui [Leach], [1814]) 


(2) that the under-mentioned name be placed on the Official List of 
Specific Names in Zoology : montagui [Leach], [1814], as pub- 
lished in the combination Pandalus montagui (specific name 
of type species of Pandalus [Leach], [1814]) ; 


(3) that the under-mentioned names, each of which is a junior 
homonym of Pandalus [Leach], [1814], be placed on the Official 
Index of Rejected and Invalid Generic Names in Zoology : 
(a) Pandalus Leach, [1815] (Malac. Podohpth. Brit. pl. 40(2)) ; 
(b) Pandalus Leach, 1815 (Trans. linn. Soc. Lond. 11(2) : 346) ; 


(4) that the under-mentioned specific name be placed on the Official 
Index of Rejected and Invalid Specific Names in Zoology : 
annulicornis Leach, [1815] (Malac. Podophth. Brit. : p\. 40(2)), 
as published in the combination Pandalus annulicornis (a 
junior objective synonym of montagui [Leach], [1814], as pub- 
lished in the combination Pandalus montagui). 


DIRECTION 47 251 


ANNEXE 


Selection of a specimen preserved in the British Museum (Natural 
History), London, to be the lectotype of the nominal species 
‘*Pandalus montagui’’ [Leach], [1814], and ‘‘ Pandalus 
annulicornis ’’ Leach, [1815] (Class Crustacea, Order 
Decapoda) 


By -. Bo HOLTHUIS 
(Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 


(letter, dated 15th November 1955, to Francis Hemming, Secretary 
to the International Commission on Zoological Nomenclature) 


Thank you for your letter of 11th November and the information 
concerning the genus Pandalus Leach and its type material. On the 
basis of the data so kindly provided by Dr. Gordon, I now select as the 
lectotype of both the species Pandalus montagui [Leach], [1814] (Edinb. 
Encycl. 7(2) : 432) and Pandalus annulicornis Leach, [1815] (Malac. 
Podophth. Brit. (2) : expl. pl. 40), the dry specimen labelled ‘‘ Zetland, 
Scotland, Dr. Fleming’’, and registered in the Crustacea collection 
of the British Museum (Natural History) under the number “ 267d ”’. 


2. Registration of the present application : Upon the receipt 
of Mr. Hemming’s Report (paragraph 1 above) the question of 
the correction of the entry in the Official List of Generic Names in 
Zoology relating to the name Pandalus Leach was allotted the 
Registered Number Z.N.(S.) 1028. 


Il. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


3. Issue of Voting Paper V.P.(O.M.)(55)38 : On 13th December 
1955 a Voting Paper (V.P.(O.M.)(55)38) was issued in which the 
Members of the Commission were invited to vote either for, or 


252 OPINIONS AND DECLARATIONS 


against “‘the revised proposal in relation to the generic name 
Pandalus Leach, [1815] (Class Crustacea, Order Decapoda) 
specified in paragraph 8 of the paper by the Secretary numbered 
Z.N.(S.) 1028 submitted simultaneously with the present Voting 
Paper ” [i.e. in the paragraph numbered as above in the Report 
reproduced in the first paragraph of the present Direction]. 


4. The Prescribed Voting Period: As the foregoing Voting 
Paper was issued under the One-Month Rule, the Prescribed 
Voting Period would normally have closed on 13th January 
1956. In view, however, of the possibility of delays in overseas 
mails consequent upon the abnormally heavy traffic at Christmas 
time, the Secretary on 13th December 1955 executed a Minute 
extending the Prescribed Voting Period on the above Voting 
Period from one calendar month to six weeks. Under this 
direction the Prescribed Voting Period on Voting Paper 
V.P.(O.M.)(55)38 closed on 24th January 1956. 


5. Particulars of the Voting on Voting Paper V.P.(O.M.)(55)38 : 
At the close of the Prescribed Voting Period, the state of the 
voting on Voting Paper V.P.(O.M.)(55)38 was as follows :— 


(a) Affirmative Votes had been given by the following twenty-five 
(25) Commissioners (arranged in the order in which Votes 
were received) ; 


Mertens; Lemche; Hering; Prantl; Bodenheimer ; 
Holthuis ; Vokes ; do Amaral ; Mayr; Hanko; Key ; 
Esaki; Riley; Jaczewski; Boschma; Dymond ; 
Bradley (J.C.) ; Sylvester-Bradley ; Stoll; Cabrera ; 
Kihnelt ; Bonnet; Miller; Tortonese; Hemming ; 


(b) Negative Votes : 


None ; 


DIRECTION 47 ae 


(c) Voting Papers not returned : 


None. 


6. Declaration of Result of Vote: On 25th January 1956, 
Mr. Hemming, Secretary to the International Commission, 
acting as Returning Officer for the Vote taken on Voting Paper 
V.P.(O.M.)(55)38, signed a Certificate that the Votes cast were 
as set out in paragraph 5 above and declaring that the proposal 
submitted in the foregoing Voting Paper had been duly adopted 
and that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


7. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 2nd May 1956, Mr. Hemming prepared the Ruling given 
in the present Direction and at the same time signed a Certificate 
that the terms of that Ruling were in complete accord with those 
of the proposal approved. by the International Commission in its 
Vote on Voting Paper V.P.(O.M.)(55)38. 


8. Original References: The following are the original 
references for the names placed on Official Lists and Official 
Indexes by the Ruling given in the present Direction :— 


annulicornis, Pandalus, Leach, [1815], Malac. Podophth. Brit. (2) : 
expl. 40 


montagui, Pandalus, [Leach], [1814], Brewster’s Edinburgh Ency. 
7(2) : 432 


Pandalus Leach, [1815], Malac. Podophth. Brit. (2) : expl. pl. 40(2) 
Pandalus Leach, 1815 Trans. linn. Soc. Lond. 11(2) : 346 


9. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in 
dealing with the present case, and the present Direction is 


254 OPINIONS AND DECLARATIONS 


accordingly hereby rendered in the name of the said International 
Commission by the under-signed Francis Hemming, Secretary 
to the International Commission on Zoological Nomenclature, 
in virtue of all and every the powers conferred upon him in that 
behalf. 


10. The present Direction shall be known as Direction Forty- 
Seven (47) of the International Commission on Zoological 
Nomenclature. 


Done in London, this Second day of May, Nineteen Hundred 
and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


Printed in England by MEeTcALFE & CoopeER LIMITED, 10-24 Scrutton St., London EC 2 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, c.M.G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.11. Pp. 255—264 


DIRECTION 48 


Determination of the authorship to be attributed to the 

work published anonymously in the year 1798 under the 

title Museum Boltenianum (Direction supplementary to 
Opinion 96) 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956 


Price Six Shillings and Sixpence 
(All rights reserved) 


Issued 21st November, 1956 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 48 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JoRDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President: Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., 
U.S.A.) (12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


Arranged in order of precedence by reference to date of election or of most recent 
re-election, as prescribed by the International Congress of Zoology) 


Professor H. BOSCHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(1st January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. Henning LEMCHE (Universitetets Zoologiske Museum, Copenhagen, Denmark) 
(27th July 1948) 

Professor Teiso EsAkI (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEy (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Unstitute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a. M., Germany) (Sth July 1950) 

Professor Erich Martin HERING) (Zoologisches Museum der Humboldt-Universitadt zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMOND (University of Toronto, Toronto, Canada) (12th August 1953) 

eee if ae BRADLEY (Cornell University, Ithaca, N. Y., U.S.A.) (12th August 1953) 

resident 

Professor Harold E. Voxes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) 
(12th August 1953) 

Professor Béla HANKO (Mezdgazdasagi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y., U.S.A.) 
(12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. HoLtuuts (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (15th October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, Berkeley, 
California, U.S.A.) (29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum v Praze, Prague, Czechoslovakia) (30th 

_ October 1954) 

Professor Dr. Wilhelm KiiHNELT (Zoologisches Institut der Universitdt, Vienna, Austria) 
(6th November 1954) 

ores F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 
1954 

Professor Ernst MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 

Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico TORTONESE (Museo di Storia Naturale ‘‘ G. Doria’’, Genova, Italy) 

(16th December 1954) 


DIRECTION 48 


DETERMINATION OF THE AUTHORSHIP TO BE ATTRI- 
BUTED TO THE WORK PUBLISHED ANONYMOUSLY 
IN THE YEAR 1798 UNDER THE TITLE ** MUSEUM 
BOLTENIANUM ” (‘‘ DIRECTION ’”? SUPPLE- 
MENTARY TO ‘* OPINION ”’ 96) 


RULING :—(1) It is hereby ruled that the authorship 
of the work published anonymously in 1798 under the 
title Museum Boltenianum is to be attributed to Peter 
Friedrich Réding. 


(2) The title of the work specified in (1) above is hereby 
placed on the Official List of Works Approved as Available 
for Zoological Nomenclature with the Title Number 26, 
the entry so to be made to be endorsed as regards author- 
ship as prescribed in (1) above. 


I. THE STATEMENT OF THE CASE 
On 29th October 1954, Mr. Hemming, after consultation with 


Dr. L. R. Cox (British Museum (Natural History), London) 
(paragraph 2 below), submitted to the International Commission 


DEC : + 10Re 


258 OPINIONS AND DECLARATIONS 


on Zoological Nomenclature the following application in which, 
in order to obtain guidance as to the action to be taken when 
entering on the Official Lists or Official Indexes names first 
published in 1798 in the anonymous work Museum Boltenianum, 
he asked for a Ruling on the question of the authorship to be 
attributed to the foregoing work, this being a matter which had 
not been dealt with by the Commission when in Opinion 96 
(1926, Smithson. misc. Coll. 73 (No. 4) : 16—18) it had ruled 
in favour of the acceptability of names published in the foregoing 
work :— 


Proposed determination of the authorship to be attributed 
to the catalogue published anonymously in 1798 under 
the title ‘‘ Museum Boltenianum’”’ (problem 
supplementary to ‘‘ Opinion ’’ 96) 


By FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


The active steps now being taken to prepare the Official List of 
Generic Names in Zoology (and also the other Official Lists and the 
Official Indexes) for publication in book form have brought to light 
inconsistencies in the treatment accorded to various books in some of 
the older entries on the List. Most of these present no real difficulty, 
but some are of a nature which, in my view, calls for a further decision 
by the International Commission. The present note, which is con- 
cerned with the authorship of the anonymously published catalogue 
entitled the Museum Boltenianum which appeared in 1798, is one of 
these cases. 


2. As the title suggests, the Museum Boltenianum is a catalogue of the 
collection formed by J. F. Bolten. It was prepared for publication by 
P. F. Réding. This work contains many new names, and the question 
which calls for decision is whether those names should be attributed 
to Bolten or to Réding. 


3. The question whether the Museum Boltenianum should be 
accepted as having been duly published for the purposes of Article 25 
of the Régles was submitted to the Commission some thirty years ago 
by Dr. Tate Regan (British Museum (Natural History), London), and 
in Opinion 96 (published in 1926, Smithson. misc. Coll. 73 (No. 4) : 16— 


DIRECTION 48 259 


18) the Commission ruled in favour of the availability of this work. 
In that Opinion the Commission gave no indication, however, of its 
view as to the authorship to be attributed to this work. 


4. I discussed this question with Mr. R. Winckworth (London) some 
years before his death, knowing that he possessed an exceptionally 
wide and detailed knowledge of the older literature of his subject. 
The view which he took was that the new names in the Museum 
Boltenianum were Bolten’s names, that the “‘indications”’ given for 
those names were provided by Réding and therefore that Réding 
and not Bolten should, under the Régles, be regarded as the author 
of the new names in this book. Not long after this discussion, 
Mr. Winckworth published a short note setting out the above views 
(Winckworth, 1945, Proc. malac. Soc. Lond. 26 : 136). 


5. In anticipation of making the present application, I recently 
applied for advice to Dr. L. R. Cox (British Museum (Natural History), 
London), whose work also, as I knew, brought him into contact with 
the Museum Boltenianum. Dr. Cox at once agreed that a decision 
on the authorship to be attributed to this work was overdue, and 
kindly undertook to write to me as soon as he had examined the 
literature relating to this subject. In an interesting letter dated 
26th October 1954 (which is being published simultaneously with the 
present application)! Dr. Cox gives what I consider to be very con- 
vincing reasons for accepting Réding as the author, for the purposes 
of zoological nomenclature, of the new names in the Museum 
Boltenianum. 


6. Bolten was undoubtedly the author of the names in question, 
but it appears to me to be clear from the information furnished by 
Mr. Winckworth and Dr. Cox that it was Réding alone who supplied 
the ‘‘indications’’ for these names. On this view, these names 
should be regarded as manuscript names of Bolten’s validated by 
being published with indications by Réding. As such, these names 
are attributable to Rdding, whose action alone gave them the status 
of availability under the Régles (1950, Bull. zool. Nomencl. 4 : 259—260). 


7. I accordingly recommend that Réding should be accepted as the 
author of the Museum Boltenianum. recommend also that the present 
opportunity should be taken to place the title of the above work on the 
Official List of Works Approved as Available for Zoological Nomen- 
clature (1953, Copenhagen Decisions zool. Nomencl. : 24, Decision 24), 
a step which will in any case need to be taken at some time in pursuance 


1 The letter from Dr. Cox here referred to is reproduced in Baraerape 2 of the 
present Direction. 


260 OPINIONS AND DECLARATIONS 


of the General Directive issued by the Thirteenth International Congress 
of Zoology, Paris, 1948, that decisions taken in Opinions previously 
rendered be transferred to the appropriate Schedules to the Régles 
(1950, Bull. zool. Nomencl. 4 : 340). 


8. The proposals now submitted are therefore that the International 
Commission should :— 


(1) give a Ruling that the authorship of the work published anony- 
mously in 1798 under the title Museum Boltenianum be attributed 
to Peter Friedrich Réding ; 


(2) place the title of the foregoing work, with the authorship attri- 
buted as directed in (1) above, on the Official List of Works 
Approved as Available for Zoological Nomenclature (codification 
of Ruling given in Opinion 96). 


2. Support by Dr. L. R. Cox (British Museum (Natural History), 
London) for the acceptance of P. F. Rodding as the author of the 
‘*Museum Boltenianum’’: Following oral discussions with 
Mr. Hemming, Secretary to the International Commission, 
Dr. L. R. Cox (British Museum (Natural History), London) on 
26th October 1954 addressed a letter to the Office of the Com- 
mission setting out the evidence available regarding the authorship 
of the work Museum Boltenianum. Dr. Cox’s letter, which is 
referred to by Mr. Hemming in paragraph 5 of his application, 
was as follows :— 


The position seems to be that J. F. Bolten built up a large collection 
which he arranged systematically in accordance with a new scheme of 
classification of his own, assigning to each species a binominal Latin 
name and a vernacular German name. There is no evidence that he 
contemplated any sort of publication, but we are told (p. iti. of the 
Museum Boltenianum) that there was a “‘ systematic catalogue of the 
collection which had been prepared with the greatest accuracy and 
care by its late possessor ”’ [translation from the Latin]. On his death 
it was desired to sell the collection, and P. F. Réding took the catalogue 
and added to each species any references to figures in earlier literature 
which he could trace and a reference to the name assigned to the species 
in the 13th (Gmelin’s) edition of Linnaeus, if it was there listed. 
All this is explained by RéGding in his “‘ Vorrede”’ to the Museum 
Boltenianum (pp. vii—viii), an English translation of which has been 
published by W. H. Dall (1915, Smithson. Publ. 2360 : 9). 


DIRECTION 48 261 


R. Winckworth (1945, Proc. malac. Soc. Lond. 26 : 136) has discussed 
the question of the authorship of the work as follows : 


‘** The names are often attributed to Bolten, as the originator. 
But his manuscript catalogue contained only Latin names with 
German vernacular equivalents. It was not until the catalogue 
was published ‘ revised and enlarged by the addition of synonymy 
by Peter Friedrich Roeding, a man devoted to natural history 
and especially to conchology ’, as the Abbé Lichtenstein records 
in the preface, that the names became available. Roeding in the 
introduction says: ‘On account of my love for natural history 
I accepted the labour and have added the Latin names according 
to the 13th Edition of Gmelin’s Linnean System, as well as many 
references to figures of the specimens. In many cases this had 
to be left undone because no figures were found to be satisfactory.’ 
It seems to me clear that the author should be cited as Réding 
ex Bolten MS., or more concisely as Réding. Iredale, however, 
takes the view that ‘ Bolten was the author of the genera and 


IeFo: 


nominator of the species ’. 


The practice by the earlier authors (such as Dall) who accepted the 
Museum Boltenianum was to attribute the generic and specific names 
to Bolten. Nowadays, however, almost every author known to me 
attributes them either to R6ding or to “‘ Réding ex Bolten MS.’’. 


I]. THE SUBSEQUENT HISTORY OF THE CASE 


3. Registration of the present application: Upon receipt of 
Mr. Hemming’s paper, the question of the authorship to be 
attributed to the work entitled Museum Boltenianum was allotted 
the Registered Number Z.N.(S.) 866. 


4. Publication of the present application : The present applica- 
tion, together with Dr. Cox’s letter reproduced in paragraph 2 
above, was sent to the printer on 22nd November 1954 and was 
published on 31st January in the following year in Part 2 of 
Volume 11 of the Bulletin of Zoological Nomenclature (Hemming, 
1955, Bull. zool. Nomencl. 11 : 56—S7 ; Cox, ibid. 11 : 58). 


262 OPINIONS AND DECLARATIONS 


5. No objection received: The publication of the present 
application elicited no objection to the action proposed from 
any source. 


Ill. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


6. Issue of Voting Paper V.P.(55)17 : On 9th November 1955, 
a Voting Paper (V.P.(55)17) was issued in which the Members 
of the Commission were invited to vote either for, or against, 
*“ the proposal relating to the authorship of the catalogue entitled 
Museum Boltenianum and other matters set out in Points (1) and 
(2) in paragraph 8 on page 57 of Volume 11 of the Bulletin of 
Zoological Nomenclature ’’ [i.e. in the Points numbered as above 
in paragraph 8 of the application reproduced in the first para- 
graph of the present Direction]. 


7. The Prescribed Voting Period: As the foregoing Voting 
Paper was issued under the Three-Month Rule, the Prescribed 
Voting Period closed on 9th February 1956. 


8. Particulars of the Voting on Voting Paper V.P.(55)17 : At 
the close of the Prescribed Voting Period, the state of the voting 
on Voting Paper V.P.(55)17 was as follows :— 


(a) Affirmative Votes had been given by the following twenty- 
four (24) Commissioners (arranged in the order in which 
Votes were received) : 


Holthuis; Hering; Vokes; Bradley (J.C.); Mayr; 
Jaczewski ; Riley ; Dymond ; Prantl ; Sylvester-Bradley ; 


DIRECTION 48 263 


Lemche ; Esaki; Mertens ; do Amaral ; Hanko; Key ; 
Boschma ; Cabrera ; Stoll ; Bonnet ; Miller ; Tortonese ; 
Kihnelt ; Hemming ; 


(b) Negative Votes : 


None ; 


(c) Voting Papers not returned, one (1) : 


Bodenheimer. 


9. Declaration of Result of Vote : On 10th February, 1956, 
Mr. Hemming, Secretary to the International Commission, acting 
as Returning Officer for the Vote taken on Voting Paper V.P.(55)17, 
signed a Certificate that the Votes cast were as set out in para- 
graph 8 above and declaring that the proposal submitted in the 
foregoing Voting Paper had been duly adopted and that the 
decision so taken was the decision of the International Commission 
in the matter aforesaid. 


10. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 20th May 1956, Mr. Hemming prepared the Ruling given 
in the present Direction and at the same time signed a Certificate 
that the terms of that Ruling were in complete accord with those 
of the proposal approved by the International Commission in 
its Vote on Voting Paper V.P.(55)17. 


11. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in dealing 
with the present case, and the present Direction is accordingly 
hereby rendered in the name of the said International Com- 
mission by the under-signed Francis Hemming, Secretary to the 
International Commission on Zoological Nomenclature, in virtue 
of all and every the powers conferred upon him in that behalf. 


264 OPINIONS AND DECLARATIONS 


12. The present Direction shall be known as Direction Forty- 


Fight (48) of the International Commission on Zoological 
Nomenclature. 


Done in London, this Twentieth day of May, Nineteen Hundred 
and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


a oe! 


Printed in England by Mercatre & Cooper LimiteDp, 10-24 Scrutton St., London BC2 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, C.M.G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.12. Pp. 265—280 


DIRECTION 49 
Validation under the Plenary Powers of the generic name 
Iphis Leach, 1817 (Class Crustacea, Order Decapoda) 
(correction of an error in the Ruling given in Opinion 73) 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956 


Price Ten Shillings and Sixpence 
(All rights reserved) Ail 


Issued 21st November, 1956 \ iri go fv 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 49 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JorRDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President : Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) 
(12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary : Mr. Francis HemMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(Arranged in order of precedence by reference to date of election or of most recent re-election, 
as prescribed by the International Congress of Zoology) 


Professor H. BoscHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(Ist January 1947) 

Senor Dr. Angel CaBRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

DE: ne was (Universitetets Zoologiske Museum, Copenhagen, Denmark) (27th 
uly 194 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh Riey (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (5th July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitat zu 
Berlin, Germany) (5th July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. Dymonp (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N.¥., U.S.A.) (12th August 
1953) (President) 

Professor Harold E. Vokes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) (12th 
August 1953) 

Professor Béla HANK6 (Mez6gazdasdgi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y¥% 
U.S.A.) (12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. Hottuuis (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (15th October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferninand PRANTL (Ndrodni Museum V Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KUHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

ee eo F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 
19 


) 
Professor Ernst MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 
Professor Enrico ToRTONESE (Museo di Storia Naturale ‘* G. Doria ”’ Genova, Italy) 
(16th December 1954) 


DIRECTION 49 


VALIDATION UNDER THE PLENARY POWERS OF THE 
GENERIC NAME “IPHIS ” LEACH, 1817 (CLASS 
CRUSTACEA, ORDER DECAPODA) (CORRECTION 
OF AN ERROR IN THE RULING GIVEN IN 
‘* OPINION ”’ 73) 


RULING :—(1) Under the Plenary Powers the generic 
name Iphis Meigen, 1800 (Class Insecta, Order Diptera) 
is hereby suppressed for the purposes both of the Law of 
Priority and of the Law of Homonymy. 


(2) The following revised entry on the Official List of 
Generic Names in Zoology for Name No. 160 is hereby 
substituted for the entry in respect of that Name Number 
made by the Ruling given in Opinion 73 :— 


Iphis Leach, 1817, as validated under the Plenary 
Powers under (1) above (gender: masculine) (type 
species, by monotypy: Cancer  septemspinosus 
Fabricius (J.C.), 1787). 


(3) The under-mentioned generic name is hereby 
placed on the Official List of Generic Names in Zoology 
with the Name Number 1035 :—Dolichopus Latreille, 
1796 (gender : masculine) (type species, by selection by 
Latreille (1810) : Musca ungulata Linnaeus, 1758). 


268 OPINIONS AND DECLARATIONS 


(4) The under-mentioned specific names are hereby 
placed on the Official List of Specific Names in Zoology 
with the Name Numbers 1055 and 1056 respectively :— 


(a) septemspinosus Fabricius (J.C.), 1787, as published 
in the combination Cancer septemspinosus (specific 
name of type species of Jphis Leach, 1817) ; 


(b) ungulata Linnaeus, 1758, as published in the com- 
bination Musca ungulata (specific name of type 
species of Dolichopus Latreille, 1796). 


(5) The under-mentioned generic names are hereby 
placed on the Official Index of Rejected and Invalid Generic 
Names in Zoology with the Name Numbers 752 to 754 
respectively :— 


(a) Dolichopus Van der Hoeven, [1856] (a junior 
homonym of Dolichopus Latreille, 1796) ; 


(b) Dolychopus Samouelle, 1819 (an Erroneous Subse- 
quent Spelling for Dolichopus Latreille, 1796) ; 


(c) Iphis Meigen, 1800 (as suppressed under the Plenary 
Powers under (1) above). 


(6) The under-mentioned family-group name is hereby 
placed on the Official List of Family-Group Names in 
Zoology with the Name Number 98 :—DOLICHOPODIDAE 
(correction of DOLICHOPODES) Latreille, 1809 (type genus : 
Dolichopus Latreille, 1796) (first published in. correct 
form aS DOLICHOPODIDAE by Loew, 1862). 


(7) The under-mentioned family-group names are 
hereby placed on the Official Index of Rejected and Invalid 
Family-Group Names in Zoology with the Name Numbers 
85 to 87 respectively :— 


(a) DOLICHOPODES Latreille, 1809 (type genus: Doli- 
chopus Latreille, 1796) (an Invalid Original 
Spelling for DOLICHOPODIDAE) ; 


DIRECTION 49 269 


(b) DOLYCHOPODAE Samouelle, 1819 (type genus : Doli- 
chopus Latreille, 1796) (an Erroneous Subsequent 
Spelling for DOLICHOPODIDAE) ; 


(c) DOLICHOPIDAE Stephens (J.F.), 1829 (type genus : 
Dolichopus Latreille, 1796) (an Erroneous Subse- 
quent Spelling for DOLICHOPODIDAE). 


i) PAE STATEMENT OF THE: CASE 


On 4th December 1954, Mr. Francis Hemming, Secretary, 
submitted to the International Commission on Zoological 
Nomenclature the following application in which he drew attention 
to the fact that the generic name J/phis Leach, 1817 (Class Crustacea 
Order Decapoda), which had been placed on the Official List of 
Generic Names in Zoology by the Ruling given in Opinion 73, 
was invalid as a junior homonym of Jphis Meigen, 1800, a name 
not currently in use in the group concerned (Class Insecta, Order 
Diptera), and in which he recommended that the position of the 
name Iphis Leach on the Official List should be validated by the 
suppression of the unwanted name /phis Meigen :— 


Proposed suppression under the Plenary Powers of the generic name 
‘* Tphis ’? Meigen, 1800, for the purpose of validating the generic 
name ‘‘ Iphis ’’ Leach, 1817 (Class Crustacea, Order Deca- 
poda) (proposed correction of an error in ‘‘ Opinion ’’ 73) 


By FRANCIS HEMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


The purpose of the present application is to seek the assistance of the 
International Commission in rectifying an error in the Official List of 
Generic Names in Zoology in connection with the entry thereon of the 
generic name Iphis Leach, 1817 (Zool. Miscell. 3 : 25) made by the 
Ruling given in Opinion 73 (1922, Smithson. misc. Coll. 73 (No. 1) : 
~23—31). The above genus is monotypical with Cancer septemspinosus 
Fabricius, 1787 (Mantissa Ins. 1 : 325) as type species. 


270 OPINIONS AND DECLARATIONS 


2. The error in Opinion 73 with which the present application is 
concerned came to light in 1944 in the course of a preliminary survey 
which I then made of the entries on the Official List made in the pre-1936 
Opinions. I then discovered that the name Jphis Leach, 1817, is invalid 
by reason of being a junior homonym of the name Jphis Meigen, 1800 
(Nouy. Class. Mouches deux Ailes : 27) (Class Insecta, Order Diptera). 
The fact that this has passed unnoticed by workers in the Crustacea 
is due no doubt to the fact that the pamphlet in which the name Iphis 
Meigen was published was completely ignored for more than a hundred 
years after its publication. 


3. In a case of this kind clearly the first step to be taken is to 
ascertain whether the older name is in use in the group concerned and 
therefore whether any disturbance or other inconvenience would be 
suffered by workers in that group if the name in question were to be 
suppressed by the Commission under its Plenary Powers. In the 
present instance I applied for advice to Dr. Alan Stone (U.S. Depart- 
ment of Agriculture, Agricultural Research Administration, Bureau of 
Entomology and Plant Quarantine, Washington, D.C., U.S.A.), who 
in a letter dated 16th February 1944 informed me :— 


(1) that the type species of Iphis Meigen, 1800, is Musca ungulata 
Linnaeus, 1758 (Syst. Nat. (ed. 10) 1 : 598) by selection by 
Coquillet (1910, Proc. U.S. nat. Mus. 37 : 555) ; 


(2) that the nominal species Musca ungulata Linnaeus, 1758, is also 
the type species of the genus Dolichopus Latreille, 1796 (Précis 
Caract. gén. Ins. : 159) by selection by Latreille, 1810 (Consid. 
gén. Anim. Crust. Arach. Ins. : 443, 387) ; 


(3) that in view of (1) and (2) above, the generic name Jphis Meigen, 
1800, is a junior objective synonym of Dolichopus Latreille, 
1796. 


4. In my view the information supplied by Dr. Stone clears the 
ground for the validation of the generic name /phis Leach, 1817, by 
the suppression under the Plenary Powers of the unwanted—because 
objectively invalid—name Jphis Meigen, 1800, in the Order Diptera. 
I accordingly recommend that this action should be taken. 


5. Under the General Directive given by the International Congress 
of Zoology that in rendering Opinions the Commission is to cover the 
whole field involved, it will be necessary in any decision on the present 
case also to place on the Official List of Generic Names in Zoology 
the generic name Dolichopus Latreille, 1796, and on the Official List 
of Specific Names in Zoology the name ungulata Linnaeus, 1758, as 
published in the combination Musca ungulata (specific name of type 
species of Dolichopus Latreille, 1796). At the same time it will be 
necessary to place on the Official Index of Rejected and Invalid Generic 


DIRECTION 49 271 


Names in Zoology certain names which are junior homonyms of the 
generic names dealt with in the present application. 


6. The nominal genus [phis Leach, 1817 (Class Crustacea) is, I am 
informed by Dr. Isobel Gordon (British Museum (Natural History), 
London), currently referred to the family LEUCOSIIDAE, and in conse- 
quence no family-group-name question arises in this case. 


7. I am indebted to Dr. John Smart (University of Cambridge, 
Department of Zoology, Cambridge) for information regarding the 
family-group names based on the generic name Dolichopus Latreille, 
1796. Samouelle (1819, Entomologist’s useful Compendium : 294) was 
the first author who has been traced as having used a “‘ -DAE”’ termina- 
tion, but he made two spelling mistakes, the name appearing in his 
book as DOLYCHOPODAE. Moreover, Samouelle attributed this name 
to Leach, without giving a bibliographical reference. Possibly, it was 
only used by Leach in manuscript. 


8. The recommendations which I now submit are that the Inter- 
national Commission should :— 


(1) use its Plenary Powers for the purpose of suppressing the generic 
name Iphis Meigen, 1800 (Class Insecta, Order Diptera), for 
the purposes both of the Law of Priority and of the Law of 
Homonymy ; 


(2) substitute the following revised entry on the Official List of 
Generic Names in Zoology for Name No. 160 :— 


Iphis Leach, 1817, Zool. Miscell. 3 : 25 (gender : masculine) 
(type species, by monotypy : Cancer septemspinosus Fabricius, 
1787 (Mantissa Ins. 1 : 325)) ; 


(3) place the under-mentioned generic name on the Official List of 
Generic Names in Zoology :— 


Dolichopus Latreille, 1796 (gender : masculine) (type species, 
by selection by Latreille (1810) : Musca ungulata Linnaeus, 
1758) ; 


(4) place the under-mentioned specific names on the Official List of 
Specific Names in Zoology :— 


(a) septemspinosus Fabricius, 1787, as published in the com- 
bination Cancer septemspinosus (specific name of type 
species Iphis Leach, 1817) ; 


(b) ungulata Linnaeus, 1758, as published in the combination 
Musca ungulata (specific name of type species of Doli- 
chopus Latreille, 1796) ; 


2/2 OPINIONS AND DECLARATIONS 


(5) place the under-mentioned generic names on the Official Index 
of Rejected and Invalid Generic Names in Zoology :— 


(a) Dolichopus Van der Hoeven, [1856], Handb. Zool. 2 : 422, 
431 (a junior homonym of Dolichopus Latreille, 1796) ; 

(b) Iphis Meigen, 1800, as suppressed under the Plenary Powers 
under (1) above ; 

(c) the under-mentioned names which are junior homonyms of 
Iphis Leach, 1817 :— 
(i) Iphis Koch, 1835, Deutschl. Crust. (Heft 2) : Tab. 6 
(ii) Iphis Laporte, 1836, Rev. Entom. 4(1) :7 

(6) place the under-mentioned family-group name on the Official 
List of Family-Group Names in Zoology :—DOLICHOPIDAE 


(correction of DOLICHOPIDES) Latreille, 1807, Gen. Crust. 
Ins. 3 : 290! (type genus : Dolichopus Latreille, 1796). 


(7) place the under-mentioned family-group names on the Official 
Index of Rejected and Invalid Family-Group Names in Zoology :— 


(a) DoLIcHoPIDES Latreille, 1807 (type genus: Dolichopus 
Latreille, 1897) (an Invalid Original Spelling for DOLI- 
CHOPIDAE) ; 


(b) DOLYCHOPODAE Samouelle, 1819 (type genus Dolichopus 
(misspelt Dolychopus by Samouelle) Latreille, 1796) (an 
Erroneous Subsequent Spelling for DOLICHOPIDAE). 


Il. THE SUBSEQUENT HISTORY OF THE CASE 


2. Registration of the present application : Upon the receipt of 
Mr. Hemming’s application the question of the status of the 
generic name Iphis Leach, 1817, was allotted the Registered 
Number Z.N.(S.) 562. 


3. Publication of the present application : The present application 
was sent to the printer on 31st December 1954 and was published 
on 28th February in the following year in Part 3 of Volume 11 
of the Bulletin of Zoological Nomenclature (Hemming, 1955, Bull. 
zool. Nomencl. 11 : 79—81). 


~ 


On the final re-check of the references cited in the present case it was found that 
the reference here given for DOLICHOPIDES Latreille is incorrect, that name not 
having appeared until Volume 4 of the work cited, which was published in 1809. 
See paragraph 18 of the present Direction. ; 


DIRECTION 49 273 


4. Issue of Public Notices : Under the revised procedure pre- 
scribed by the Thirteenth Internationa] Congress of Zoology, 
Paris, 1948 (1950, Bull. zool. Nomencl. 4 : 51—56) Public Notice 
of the possible use by the International Commission on Zoological 
Nomenclature of its Plenary Powers in the present case was given 
on 28th February 1955 (a) in Part 3 of Volume 9 of the Bulletin 
of Zoological Nomenclature (the part in which Mr. Hemming’s 
application was published) and (b) to the other prescribed serial 
publications. In addition, such Notice was given also to certain 
general zoological serial publications and to certain entomological 
serials in Europe and America. 


5. Comments received : No objection to the application sub- 
mitted in the present case was received from any specialist in the 
Order Decapoda. On the subsidiary entomological issue involved 
communications were however received from three specialists. 
The first two of these were from :—(1) Dr. Alan Stone (United 
States National Museum, Washington, D.C., U.S.A.), who, while 
agreeing that the name Jphis Meigen was not required for taxo- 
nomic purposes, expressed opposition on general grounds to the 
use by the Commission of its Plenary Powers for the purpose of 
suppressing that name as proposed in the present application ; 
(2) Professor J. Chester Bradley (Cornell University, Ithaca, N.Y., 
U.S.A.), who rejected the arguments advanced by Dr. Stone 
against the present application. The communications so received 
are reproduced in the immediately following paragraphs. The 
third specialist who communicated with the Office of the Com- 
mission in regard to the present case was Dr. Curtis W. Sabrosky 
(U.S. Department of Agriculture, Agricultural Research Service, 
Entomology Research Branch, Washington, D.C., U.S.A.). 
Dr. Sabrosky’s communication was concerned to secure a 
correction in the portion of the application relating to the 
form to be adopted for the family-group name based on 
the generic name Dolichopus Latreille, 1796 (paragraph 8 
below), a subject which had also been raised by Professor 
Chester Bradley. Dr. Sabrosky did not comment on the subject 
matter of the present application, having already in a separate 
application (Z.N.(S.) 191) recommended that the work (Nouvelle 
Classification des Mouches a deux Ailes) in which the name Iphis 
Meigen was published should be rejected by the International 


2T4 OPINIONS AND DECLARATIONS 


Commission under its Plenary Powers (Sabrosky, 1952, Bull. 
zool. Nomencl. 6 : 131—141). 


6. Comment received from Dr. Alan Stone (United States 
National Museum, Washington, D.C., U.S.A.) : In the early part 
of 1944 Mr. Hemming consulted Dr. Alan Stone (United States 
National Museum, Washington, D.C., U.S.A.) in regard to the 
present case and on 16th February of that year Dr. Stone furnished 
a statement of his views. The statement so submitted, which was 
endorsed by a further letter dated 10th December 1954, was as 
follows (Stone, 1955, Bull. zool. Nomencl. 11 : 82) :— 


I am writing in reply to your letter concerning the generic name 
Iphis Meigen, 1800. This name is an isogenotypic synonym of 
Dolichopus Latreille, 1796. The bibliographical data are as follows : 


Dolichopus Latreille, 1796, Précis Caract. Ins. : 159. Genotype, 
Musca ungulata L., by designation of Latreille, 1810, Consid. gén. 
Anim. Crust. Arach, Ins. : 443 


Iphis Meigen, 1800, Nouvelle classification : 27. Genotype, Musca 
ungulata L., by designation of Coquillett, 1910, Proc. US. nae: 
Mus. 37 335) 


To the best of my knowledge, Jphis Meigen has always been treated, 
when its zoological position was mentioned, as a synonym of Dolichopus. 
There seems to be no possibility that it will ever come into use in the 
Diptera, since this would require either the discovery that Dolichopus 
was a homonym or that the genotype of Jphis is incorrect and that the 
true genotype is not congeneric with Dolichopus. Neither of these 
contingencies seems at all probable. 


In spite of the fact that Jphis Meigen is invalid, although available, 
I should not like to see the name suppressed by the Commission. 
The basic principles of the International Rules are weakened every time 
an exception is made by official action, and I think that the Plenary 
Powers should be invoked only in the very few cases where the name 
to be saved is of extreme importance. The easiest and surest way of 
maintaining the prestige of the International Rules is by strict im- 
partiality of application, and such prestige should not be endangered 
because of a relatively unimportant name in Crustacea. A number 
of names have been accepted as unavailable because they proved to be 
homonyms of earlier names that are not in use. If the Commission 
makes an exception here it opens the gate for more and more changes 
of this sort, until nomenclature will not be a matter of following rules, 
but of attempting to weigh rather unpredictable opinion. 


DIRECTION 49 ZPD 


I might add that the Commission is going to be subject to consider- 
able pressure to suspend various of the Meigen 1800 names in spite 
_ of Opinion 28 and the recent reaffirmation of this Opinion. If the 
Commission intends to adhere to these Opinions, an exception made 
with one name, even though seemingly fostering stability, will weaken 
its stand. It will be far easier to adhere strictly to the Rules than to 
justify partial adherence. 


The situation then is this. As far as [phis is concerned, it affects the 
dipterists not at all, but does affect workers with Crustacea. In saving 
the name for the Crustacea, all zoology will lose, since the principles 
of zoological nomenclature will be weakened, and every name that has 
been changed in similar circumstances will be subject to uncertainty. 


7. Comment received from Professor J. Chester Bradley (Cornell 
University, Ithaca, N.Y., U.S.A.) : On 2nd June 1955, Professor J. 
Chester Bradley (Cornell University, Ithaca, N.Y., U.S.A.) 
addressed a letter to the Office of the Commission in which he 
commented on a number of cases. The following is an extract 
which bears on the present case :— 


2. Iphis. 1 fully approve of this case. I totally disagree with 
Stone’s concluding paragraph on page 82%. Zoology (and the prestige 
of the Commission) would lose a lot by removing without good cause 
a name from the Official List that has been there for 33 years. 


8. The question of the form to be adopted for the family-group 
name based on the generic name ‘* Dolichopus ”’ Latreille, 1796 : 
Communications were received from two specialists drawing 
attention to the fact that the correct form for the family name 
based on the generic name Dolichopus Latreille, 1796, was 
DOLICHOPODIDAE and not DOLICHOPIDAE (as stated in the 
present application. The communications so received were the 
following :— 


(a) From Professor J. Chester Bradley (Cornell University, 
Ithaca, N.Y., U.S.A.) (statement enclosed with a letter 
dated 2nd June 1955) : 


** Dolichopus ’’ comes from the Greek doAryés, long, plus zoids, foot. 
As the genitive of ods is 7050S, the family name should be spelled 
DOLICHOPODIDAE and this is the form with which I am most familiar, 
though I have not examined literature to see how its use compares 


2 The page number cited by Professor Chester Bradley is to the Part of the Bulletin 
of Zoological Nomenclature containing the comment by Dr. Alan Stone repro- 
duced in paragraph 6 of the present Direction. 


276 OPINIONS AND DECLARATIONS 


with the incorrect DOLICHOPIDAE. See Essig, 1942, College Entomology : 
Tb: 


(b) From Dr. Curtis W. Sabrosky (U.S. Department of Agri- . 
culture, Agriculture Research Service, Entomology Re- 
search Branch, Washington, D.C., U.S.A.) (extract from 
a letter dated 24th August 1955) : 

I note that this application proposes to place on the Official List 
the family name DOLICHOPIDAE (correction of DOLICHOPIDES) Latreille, 
and to place on the Official List of Rejected and Invalid Family-Group 
Names in Zoology the spellings DOLICHOPIDES Latreille (1809) and 
DOLYCHOPODAE Samouelle (1819). I should like to make the following 
comments on the proposal :— 


(1) The correct family name should be DOLICHOPODIDAE rather than 
DOLICHOPIDAE (last component of the generic name Dolichopus from 
the Greek zrous, zod0s), There is much usage both ways, incidentally. 


(2) I believe it is an unwise precedent for the Commission to place 
variant subsequent spellings of family names on the Official Index of 
Rejected Names. 


9. Modification of proposal submitted in Point (7) in paragraph 8 
of the present application regarding the form to be adopted for 
the family name based on the generic name ‘‘ Dolichopus ”’ 
Latreille, 1796 : When on 3rd November 1955, Mr. Hemming, 
as Secretary, prepared the Voting Paper to be submitted to the 
Members of the Commission in the present case (as to which 
see paragraph 10 of the present Direction) he added a note 
(Note 3) drawing attention to the communications received from 
Professor J. Chester Bradley and Dr. Curtis W. Sabrosky (para- 
graph 8 above) on the subject of the form of the family name 
based on the generic name Dolichopus Latreille, 1796, and 
recommending that the proposal on this matter be amended as 
follows :—‘* DOLICHOPODIDAE (correction of DOLICHOPODES) 
Latreille, 1809, to be placed on the Official List, Latreille’s 
Invalid Original Spelling to be placed on the Official Index.” 


Il. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


10. Issue of Voting Paper V.P.(55)21 : On 9th November 1955, 
a Voting Paper (V.P.(55)21) was issued in which the Members 


DIRECTION 49 Zier 


of the Commission were invited to vote either for, or against, 
“the proposal relating to Iphis Meigen, 1800, and Jphis Leach, 
1817, as set out in Points (1) to (7) in paragraph 8 on pages 80 
and 81 of Volume 11 of the Bulletin of Zoological Nomenclature 
[in the paragraph numbered as above in the application repro- 
duced in the first paragraph of the present Direction], subject 
to the amendment of Point (7) as proposed in Note 3 overleaf ”’ 
[i.e. amended as proposed in the Note quoted in paragraph 9 
_of the present Direction]. 


11. The Prescribed Voting Period: As the foregoing Voting 
Paper was issued under the Three-Month Rule, the Prescribed 
Voting Period closed on 9th February 1956. 


12. Particulars of the Voting on Voting Paper V.P.(55)21 : At 
the close of the Prescribed Voting Period the state of the voting 
on Voting Paper V.P.(55)21 was as follows :— 


(a) Affirmative Votes had been given by the following twenty- 
four (24) Commissioners (arranged in the order in which 
Votes were received) : 


Holthuis ; Hering ; Vokes ; Bodenheimer ; Bradley (J.C.) ; 
Mayr ; Jaczewski ; Riley ; Dymond ; Prantl ; Lemche ; 
Esaki ; do Amaral ; Hank6o ; Key ; Boschma ; Sylvester- 
Bradley ; Mertens ; Cabrera ; Stoll ; Bonnet ; Tortonese ; 
Kuhnelt ; Hemming ; 


(b) Negative Votes, one (1): 


Miller ; 


(c) Voting Papers not returned : 


None. 


278 OPINIONS AND DECLARATIONS 


13. Declaration of Result of Vote : On 10th February 1956, 
Mr. Hemming, Secretary to the International Commission, acting 
as Returning Officer for the Vote taken on Voting Paper 
V.P.(55)21, signed a Certificate that the Votes cast were as set out 
in paragraph 12 above and declaring that the proposal submitted 
in the foregoing Voting Paper had been duly adopted and that 
the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


14. Withdrawal of two proposals on which, subsequent to the 
publication of the present application, action was taken by the 
International Commission by the Ruling given in ‘* Direction ”’ 37 : 
On 29th December 1955, Mr. Hemming, as Secretary, after 
signing Direction 37, in which, inter alia, the Commission placed 
the generic names Jphis Koch, 1835, and Jphis Laporte, 1836, 
on the Official Index of Rejected and Invalid Generic Names in 
Zoology, executed the following Minute withdrawing from the 
scope of Application Z.N.(S.) 562 the proposals there submitted 
in regard to the foregoing names :— 


Withdrawai of proposals relating to two junior 
homonyms of ‘‘ Iphis ’? Leach, 1817 


By FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


The routine proposals submitted in the present case included a 
recommendation (in Point (5) of paragraph 8) that the names Iphis 
Koch, 1835, and Jphis Laporte, 1836, as junior homonyms of Iphis 
Leach, 1817, should be placed on the Official Index of Rejected and 
Invalid Generic Names in Zoology. The same proposal was later 
included in a paper numbered Z.N.(S.) 987, which was concerned 
with clearing up outstanding questions relating to the names of genera 
belonging to the Order Decapoda of the Class Crustacea placed on the 
Official List of Generic Names in Zoology in the period up to the end of 
1936. This latter paper was submitted to the Commission under the 
One-Month Rule on 19th September 1955 with Voting Paper 
V.P.(O.M.)(55)28 and was approved by the Commission on Ist 
November 1955. 


2. In the light of the above decision the proposals relating to the 
disposal of the foregoing invalid generic names submitted with Paper 
Z.N.(S.) 987, together with the other proposals there submitted, 


DIRECTION 49 279 


incorporated on 29th December 1955 in the Ruling given in a Direction 
numbered Direction 37 (now in the press)?. 


3. In these circumstances no further action requires to be taken in 
connection with the foregoing generic names and the proposals in 
regard thereto in Application Z.N.(S.) 562 are accordingly hereby 
withdrawn. 


15. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 26th May 1956, Mr. Hemming prepared the Ruling given 
in the present Direction and at the same time signed a Certificate 
that the terms of that Ruling were in complete accord with those 
of the proposal approved by the International Commission in its 
Vote on Voting Paper V.P.(55)21, subject to the omission there- 
from of the two names specified in the Minute executed by the 
Secretary on 29th December 1955, the text of which has been 
reproduced in paragraph 14 of the present Direction. 


16. Original References : The following are the original refer- 
ences for the generic and specific names placed on Official Lists 
and Official Indexes by the Ruling given in the present 
Direction :— 


Dolichopus Latreille, 1796, Précis Caract. gén. Ins. : 159 
Dolichopus Van der Hoeven, [1856], Handb. Zool. 2 : 422, 431 


Dolychopus Samouelle, 1819, Entomologist’s useful Compendium : 
294 

Iphis Meigen, 1800, Nouv. Classif. Mouches deux Ailes : 27 

septemspinosus, Cancer, Fabricius (J.C.), 1787, Mantissa Ins. 
if 2325 


ungulata, Musca, Linnaeus, 1758, Syst. Nat. (ed. 10) 1 : 598 


17. The following is the reference for the selection of the type 
species for the genus Dolichopus Latreille, 1796, specified in the 
Ruling given in the present Direction :—Latreille, 1810, Consid. 
gén. Ordre nat. Anim. Class. Crust. Arachn. Ins. : 443, 387 


3 Direction 37 has since been published on Ist September 1956 as Part D.2 of 
Volume 1, Section D, in the Opinions and Declarations Series. 


280 OPINIONS AND DECLARATIONS 


18. The following are the references for the family-group names 
placed on the Official List and Official Index established for 
recording names for taxa belonging to that category :— 


DOLICHOPODES Latreille, 1809, Gen. Crust. Ins. 4 : 239, 290 
(type genus : Dolichopus Latreille, 1796) 


DOLYCHOPODAE Samouelle, 1819, Entomologist’s useful Com- 
pendium : 294 (type genus Dolichopus (misspelled Dolychopus by 
Samouelle) Latreille, 1796) 


DOLICHOPIDAE Stephens (J.F.), 1829, Syst. Cat. Brit. Ins. : 266 
DOLICHOPODIDAE Loew, 1862, Smithson. misc. Coll. 6 (No. 1/2) : 32 


19. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in dealing 
with the present case, and the present Direction is accordingly 
hereby rendered in the name of the said International Commission 
by the under-signed Francis Hemming, Secretary to the Inter- 
national Commission on Zoological Nomenclature, in virtue of 
all and every the powers conferred upon him in that behalf. 


20. The present Direction shall be known as Direction Forty- 
Nine (49) of the International Commission on Zoological 
Nomenclature. 


Done in London, this Twenty-Sixth day of May Nineteen 
Hundred and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


oy os oe ee ee 
Printed in England by MetcaLtFE & Cooper LimITED, 10-24 Scrutton St., London EC 2 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, cM.G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.13. Pp. 281—292 


DIRECTION 50 


Validation under the Plenary Powers of the generic name 
Homalaspis Milne Edwards (A.), 1863 (Class Crustacea, 
Order Decapoda) (correction of an error in Opinion 85) 


LONDON : 


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ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 50 


A. The Officers of the Commission 


Honorary Life President : Dr. Karl JORDAN (British Museum (Natural History), Zoological 
Museum, Tring, Herts, England) 


President : Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) 
(12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo,Brazil) (A2th August 1953) 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(Arranged in order of precedence by reference to date of election or of most recent re-election, 
as prescribed by the International Congress of Zoology) 


Professor H. BOSCHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr: ee or LEMCHE (Universitetets Zoologiske Museum, Copenhagen, Denmark) (27th 
July 1948) 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950 

Mr. Norman Denbigh RILEy (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (Sth July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitat zu Berlin, 
Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice President) 

Professor J. R. DyMOND (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) (12th August 
1953) (President) 

Professor Harold E. Voxes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) 
(12th August 1953) 

Professor Béla HANKO (Mezdgazdasagi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y., 
U.S.A.) (12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. HoLttuuts (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
ae August 1953) 

K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 

Tees A.C.T., Australia) (AS5th October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum v Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KUHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

Professor F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 


Professor Ernst MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico TORTONESE (Museo di Storia Naturale “‘ G. Doria,’ Genova, Italy) 
(16th December 1954) 


DIRECTION 50 


VALIDATION UNDER THE PLENARY POWERS OF THE 
GENERIC NAME “HOMALASPIS” MILNE 
EDWARDS (A.), 1863, (CLASS CRUSTACEA, 
ORDER DECAPODA) (CORRECTION 
OF AN ERROR IN 
** OPINION ”’ 85) 


RULING :—(1) Under the Plenary Powers the generic 
name Homalaspis Reinhard, 1860 (Class Insecta, Order 
Hymenoptera) is hereby suppressed for the purposes both 
of the Law of Priority and of the Law of Homonymy. 


(2) The following revised entry on the Official List of 
Generic Names in Zoology for the Name No. 320 is hereby 
substituted for the entry in respect of that Name Number 
made by the Ruling given in Opinion 85 :— 


Homalaspis Miine Edwards (A.), 1863, as validated 

under the Plenary Powers under (1) above (gender : 

feminine) (type species, by original designation : 
Xantho planus Milne Edwards (H.), 1834). 


APA . « wee 


284 OPINIONS AND DECLARATIONS 


. (3) The under-mentioned generic name is hereby placed 

on the Official List of Generic Names in Zoology with the 
Name No. 1036 :—Omalaspis Giraud, 1860 (gender : 
feminine) (type species, by monotypy : Omalaspis noricus 
Giraud, 1860) (Class Insecta, Order Hymenoptera). 


(4) The under-mentioned specific names are hereby 
placed on the Official List of Specific Names in Zoology 
with the Name Nos. 1057 and 1058 respectively :— 


(a) planus Milne Edwards (H.), 1834, as published in 
the combination Xantho planus (specific name of 
type species of Homalaspis Milne Edwards (A.), 
1863) (Class Crustacea, Order Decapoda) ; 


(b) niger Hartig, 1840, as published in the combination 
Figites niger (Class Insecta, Order Hymenoptera). 


(5) The under-mentioned generic name is _ hereby 
placed on the Official Index of Rejected and Invalid 
Generic Names in Zoology with the Name No. 755 :— 
Homalaspis Reinhard, 1860, as suppressed under the 
Plenary Powers under (1) above. 


I. THE STATEMENT OF THE CASE 


On 3rd December 1954, Mr. Francis Hemming, as Secretary, 
submitted to the International Commission on Zoological 
Nomenclature the following paper drawing attention to the 
fact that the generic name Homalaspis Milne Edwards (A.), 1863 
(Class Crustacea, Order Decapoda) which in 1925 (Smithson. 
misc. Coll. 73 (No. 3) : 13) was placed on the Official List of 


DIRECTION 50 285 


Generic Names in Zoology by the Ruling given in Opinion 85, 
was invalid by reason of being a junior homonym of Homalaspis 
Reinhard, 1860 (Class Insecta, Order Hymenoptera) and recom- 
mending that, as the name Homalaspis Reinhard was not in use 
in the Class Insecta, the entry on the Official List in relation to the 
name Homalaspis Milne Edwards in Crustacea shou!d be validated 
under the Plenary Powers :— 


Proposed validation, under the Plenary Powers, of the generic name 
** Homalaspis ’? Milne Edwards (A.), 1863 (Class Crustacea, 
Order Decapoda) (proposed validation of an error in 
** Opinion ’’ 85) 


By FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


__ The purpose of the present application is to seek the assistance of the 
International Commission on Zoological Nomenclature in validating 
the generic name Homalaspis Milne Edwards (A.), 1863 (Ann. Sci. nat., 
Paris (Zool.) (4) 20 : 279) (Class Crustacea, Order Decapoda), which 
was placed on the Official List of Generic Names in Zoology by the 
Ruling given in Opinion 85 (1925, Smithson. misc. Coll. 73 (No. 3) : 13) 
in the erroneous belief that it was a nomenclatorially available name. 


2. The present case came to light in 1944 during a preliminary survey 
of the pre-1936 entries on the Official List of Generic Names in Zoology 
which I carried out at that time. That survey showed that the name 
Homalaspis Milne Edwards, 1863, was invalid by reason of being a 
junior homonym of the name Homalaspis Reinhard, 1860 (Berlin. ent. 
Z. 4: 239), an emendation of the name Omalaspis Giraud, 1860 
(Verh. zool.-bot. Ges. Wien 10 (Abh.) : 155), the name of a genus 
in the Order Hymenoptera (Class Insecta). 


3. As a first step in the examination of the question of the action 
needed in the present case, I consulted Dr. R. B. Benson (British 
Museum (Natural History), London) for the purpose of ascertaining 
whether the suppression under the Plenary Powers of the name 
Homalaspis Reinhard for the purpose of validating the name Homalaspis 
Milne Edwards in the Class Crustacea would cause any disturbance or 
inconvenience in the nomenclature of the Hymenoptera. Dr. Benson 
replied on 21st April 1944 as follows :— 


I cannot see that any possible harm could come by suppressing 
Homalaspis Reinhard. 


286 OPINIONS AND DECLARATIONS 


Omalaspis Giraud is in use for a genus of the Cynipoid family 
ASPICERIDAE. ‘The three described species, all European, are rare 
or, at least, little known, and they rarely appear in the literature. 
Since Dalla-Torre & Kieffer’s (1910) Cynipidae in Das Tierreich, 
where “‘Omalaspis’’ was used, I am not aware that the form of 
the name “‘ Homalaspis”’ has ever been used. I only know of 
three references to “‘Omalaspis”’ (Hedicke, 1928, Bull. Soc. ent. 
France 1928 : 280, Weld, 1931, Proc. ent. Soc. Wash. 33 : 221 
and Rohwer & Fagan, 1917, “‘ The type species of the Genera 
of the Cynipidac.... .,.;. 75, Proce U.S. nat. Mus. 53 3357-807 
and in this type list the form of the name “‘ Homalaspis”’ is not 
even mentioned !). I cannot see any reason therefore against 
suppressing Homalaspis Reinhard as no confusion is likely to 
arise. Even if Omalaspis Giraud is later found to be invalid 
for any reason, there is a second string in Lambertiona Kieffer, 
with a conspecific type species. 


4. Dr. Benson’s letter made it quite clear that there could be no 
objection from the entomological side to the validation of the generic 
name Homalaspis Milne Edwards, 1863, in the Class Crustacea, and 
I recommend that this action should now be taken by the International 
Commission, in order that the defective entry of this name on the 


Official List of Generic Names in Zoology made by Opinion 85 may 
thereby be regularised. 


5. I further recommend that in pursuance of the General Directive 
issued to the Commission by the International Congress of Zoology 
that Rulings given in Opinions are in future to cover the whole field 
embraced in any given instance, the Commission should at the same 
time place the generic name Omalaspis Giraud, 1860, on the Official 
List of Generic Names in Zoology. As regards the type species of that 
genus, I am informed (in Jitt., 6th December 1954) by Dr. Benson 
that L. H. Weld (1952, Cynipoidea (Hym.) 1905—1950, Ann Arbor, 
Michigan : 16) has expressed the view that the name Omalaspis noricus 
Giraud, 1860, is a junior synonym of Omalaspis nigra (Hartig, 1840) 
(=Figites niger Hartig, 1840, Z. Ent. (Germar) 2 : 202). In these 
circumstances, the correct course under the standard procedure will 
be to place the name niger Hartig, 1840, but not its junior subjective 
synonym noricus Giraud, 1860, on the Official List of Specific Names 
in Zoology. At. the same time the Invalid Emendation Homalaspis 
Reinhard, 1860, will need to be placed on the Official Index of Rejected 
and Invalid Generic Names in Zoology. 


6. No question of family-group names arises in connection with 
either of the generic names involved in the present application, for the 
Decapod genus Homalaspis Milne Edwards is, I am informed by 
Dr. Isobel Gordon (British Museum (Natural History), London), 


DIRECTION 50 287 


currently referred to the family XANTHIDAE, while the Hymenoptera 
genus Omalaspis Giraud is currently referred to the family ASPICERIDAE. 


7. The recommendation which I now submit is that the International 
Commission should :— 


(1) use its Plenary Powers for the purpose of suppressing the generic 
name Homalaspis Reinhard, 1860 (Class Insecta, Order 
Hymenoptera) for the purposes both of the Law of Priority 
and of the Law of Homonymy : 


(2) substitute the following revised entry on the Official List of 
Generic Names in Zoology for Name No. 320 to 


Homalaspis Milne Edwards (A.), 1863, Ann. Sci. nat., Paris 
(Zool.) (4) 20 : 279 (gender : feminine) (type species, by 
original designation : Yantho planus Milne Edwards (H.), 
1834, Roret’s Suite a Buffon, Hist. nat. Crust. 1: 397): 


(3) place the under-mentioned generic name on the Official List of 
Generic Names in Zoology :— 


Omalaspis Giraud, 1860, Verh. zool.-bot. Ges. Wien 10(Abh.) : 
IS5 (gender: feminine) (type species, by monotypy : 
Omalaspis noricus Giraud, 1860, Ver. zool.-bot. Ges. Wien 
10(Abh.) : 155) (Class Insecta, Order Hymenoptera) ; 


(4) place the under-mentioned specific names on the Official List of 
Specific Names in Zoology :— 


(a) planus Milne Edwards (H.), 1834, as published in the 
combination Xantho planus (specific name of type species 
of Homalaspis Milne Edwards (A.), 1863) ; 


(b) niger Hartig, 1840, as published in the combination Figites 
niger ; 


(5) place the under-mentioned generic names on the Official Index of 
Rejected and Invalid Generic Names in Zoology :— 


(a) Homalaspis Reinhard, 1860, as suppressed under the 
Plenary Powers under (1) above ; 


(b) Homalaspis Brunner v. Wattenwyl, 1895, Monogr. Pseudo- 
Phylliden : 18, 217 (junior homonym of Homalaspis 
Milne Edwards (A.), 1863) ; é 


(c) Homalaspis Kiaer, 1932, Skr. Svalbard Ishavet Oslo 52: 14 
(junior homonym of Homalaspis Milne Edwards (A.), 
1863). 


288 OPINIONS AND DECLARATIONS 


Il. THE SUBSEQUENT HISTORY OF THE CASE 


2. Registration of the present application : Upon the receipt of 
Mr. Hemming’s application the question of the proposed valida- 
tion under the Plenary Powers of the generic name Homalaspis 
Milne Edwards, 1863, was allotted the Registered Number 
Z.N.(S.) 560. 


3. Publication of the present application : The present application 
was sent to the printer on 31st December 1954 and published on 
28th February in the following year in Part 3 of Volume 11 of the 
Bulletin of Zoological Nomenclature (Hemming, 1955, Bull. zool. 
Nomencl. 11 : 83—85). 


4. Issue of Public Notices : Under the revised procedure pre- 
scribed by the Thirteenth International Congress of Zoology, 
Paris, 1948 (1950, Bull. zool. Nomencl. 4 : 51—56), Public Notice 
of the possible use by the International Commission on Zoological 
Nomenclature of its Plenary Powers in the present case was given 
on 28th February 1955 (a) in Part 3 of Volume 11 of the Bulletin 
of Zoological Nomenclature (the Part in which Mr. Hemming’s 
application was published) and (b) to the other prescribed serial 
publications. In addition, such Notice was given also to certain 
general zoological serial publications and to a number of ento- 
mological serials in Europe and America. 


5. No objection received : Neither the publication of the present 
application in the Bulletin of Zoological Nomenclature nor the 
issue of Public Notices in regard thereto elicited any objection 
to the action proposed from any source. 


Ill. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


6. Issue of Voting Paper V.P.(55)22 : On 9th November, 1955 
a Voting Paper (V.P.(55)22) was issued in which the Members 


DIRECTION 50 289 


of the Commission were invited to vote either for, or against, 
“the proposal relating to the generic name Homalaspis Milne 
Edwards, 1863, as set out in Points (1) to (5) in paragraph 7 on 
page 84 and the top of page 85 of Volume 11 of the Bulletin of 
Zoological Nomenclature’ {i.e. in the Points numbered as above 
in paragraph 7 of the application reproduced in the first para- 
graph of the present Direction]. 


7. The Prescribed Voting Period: As the foregoing Voting 
Paper was issued under the Three-Month Rule, the Prescribed 
Voting Period closed on 9th February 1956. 


8. Particulars of the Voting on Voting Paper V.P.(55)22 : At 
the close of the Prescribed Voting Period, the state of the voting 
on Voting Paper V.P.(55)22 was as follows :— 


(a) Affirmative Votes had been given by the following twenty-five 
(25) Commissioners (arranged in the order in which Votes 
were received) : 


Holthuis ; Hering ; Vokes ; Bodenheimer ; Bradley (J.C.) ; 
Mayr ; Jaczewski; Riley ; Dymond ; Prantl ; Lemche ; 
Esaki ; Mertens ; do Amaral ; Hanko ; Key ; Boschma ; 
Sylvester-Bradley ; Cabrera; Stoll; Bonnet; Miller ; 
Tortonese ; Kiihnelt ; Hemming ; 


(b) Negative Votes : 


None ; 


(c) Voting Papers not returned : 


None. 


9. Declaration of Result of Vote: On 10th February 1956, 
Mr. Hemming, Secretary to the International Commission, 
acting as Returning Officer for the Vote taken on Voting Paper 
V.P.(55)22, signed a Certificate that the Votes cast were as set 
out in paragraph 8 above and declaring that the proposal sub- 


290 OPINIONS AND DECLARATIONS 


mitted in the foregoing Voting Paper had been duly adopted and 
that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


10. Withdrawal of two proposals on which, subsequent to the 
publication of the present application, action was taken by the 
International Commission by the Ruling given in ‘* Direction ’’ 37 : 
On 29th December 1955, Mr. Hemming, as Secretary, after signing 
Direction 371, in which, inter alia, the Commission placed the 
generic names Homalaspis Brunner v. Wattenwyl, 1895, and 
Homalaspis Kiaer, 1932, on the Official Index of Rejected and 
Invalid Generic Names in Zoology, executed the following Minute 
withdrawing from the scope of Application Z.N.(S.) 560 the 
proposals there submitted in regard to the foregoing names :— 


Withdrawal of proposals relating to two junior homonyms of 
‘* Homalaspis ’’ Milne Edwards (A.), 1863 


By FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


The routine proposals submitted in the present case included a 
recommendation (in Point (5) of paragraph 7) that the names 
Homalaspis Brunner v. Wattenwyl, 1895, and Homalaspis Kiaer, 1932, 
as junior homonyms of Homalaspis Milne Edwards (A.), 1863, should 
be placed on the Official Index of Rejected and Invalid Generic Names 
in Zoology. The same proposal was later included in a Paper Num- 
bered Z.N.(S.) 987, which was concerned with clearing up outstanding 
questions relating to the names of genera belonging to the Order 
Decapoda of the Class Crustacea placed on the Official List of Generic 
Names in Zoology in the period up to the end of 1936. This latter 
paper was submitted to the Commission under the One-Month Rule 
on 19th September 1955 with Voting Paper V.P.(O.M.)(55)28 and was 
approved by the Commission on Ist November 1955. 


2. In the light of the above decision the proposals relating to the 
disposal of the foregoing invalid generic names submitted with Paper 
Z.N.(S.) 987, together with the other proposals there submitted, have 
today been incorporated in the Ruling given in a Direction numbered 
Direction 37. 


1 Direction 37 was published on 1st September 1956 (Ops. Decls, int, Comm. 
zool..Nomencl. 1(D.) : 47—82). 


DIRECTION 50 291 


3. In these circumstances no further action requires to be taken in 
connection with the foregoing generic names, and the proposals in 
regard thereto in Application Z.N.(S.) 560 are accordingly withdrawn. 


11. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 27th May 1956, Mr. Hemming prepared the Ruling given 
in the present Direction and at the same time signed a Certificate 
that the terms of that Ruling were in complete accord with those 
of the proposal approved by the International Commission in its 
Vote on Voting Paper V.P.(55)22, subject to the omission 
therefrom of the two names specified in the Minute executed 
by the Secretary on 29th December 1955, the text of which 
has been reproduced in paragraph 10 of the present Direction. 


12. Original References : The following are the original refer- 
ences for the names placed on Official Lists and Official Indexes 
by the Ruling given in the present Direction :— 


Homalaspis Reinhard, 1860, Berlin. ent. Z. 4 : 239 
niger, Figites, Hartig, 1840, Z. Ent. (Germar) 2 : 202 
Omalaspis Giraud, 1860, Verh. zool.-bot. Ges. Wien 10(Abh.) : 155 


planus, Xantho, Milne Edwards (H.), 1834, (Roret’s Suite a Buffon), 
at nat. Crust. 1 : 397 


13. Family-Group-Name Aspects : No family-group-name pro- 
blems arise in the present case, the genus Homalaspis Milne 
Edwards, 1863 (Class Crustacea, Order Decapoda) being currently 
placed in the family XANTHIDAE and the genus Omalaspis Giraud, 
1860 (Class Insecta, Order Hymenoptera) in the family ASPICERIDAE. 


14. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in dealing 
with the present case, and the present Direction is accordingly 
hereby rendered in the name of the said International Com- 


292 OPINIONS AND DECLARATIONS 


mission by the under-signed Francis Hemming, Secretary to the 
International Commission on Zoological Nomenclature, in virtue 
of all and every the powers conferred upon him in that behalf. 


15. The present Direction shall be known as Direction Fifty (SO) 
of the International Commission on Zoological Nomenclature. 


DONE in London, this Twenty-Seventh day of May, Nineteen 
Hundred and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


Printed in England by Mretcatrx & Cooper LimiTep, 10-24 Scrutton St., London EC 2 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, c.M.c., CBE. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.14. Pp. 293—308 


DIRECTION 51 


Revision of the entry on the Official List of Generic 

Names in Zoology relating to the name Homarus Weber, 

1795 (Class Crustacea, Order Decapoda) (revision of a 
Ruling given in Opinion 104) 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956 


Price Ten Shillings and Sixpence 


Issued 21st November, 1956 


(All rights reserved) f ADD 9—Ve cor 
a) oe bw 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 51 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JorDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts, England). 


President : Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) 
(12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(Arranged in order of precedence by reference to date of election or of most recent re-election 
as prescribed by the International Congress of Zoology) 


Professor H. BoscHMA (Rijksmuseum van Natuurlijke Histoire,-Leiden, The Netherlands) 
(Ist January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. Sar mace (Universitetets Zoologiske Museum, Copenhagen, Denmark) (27th 
uly 194 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEy (British Museum (Natural History), oe (9th June 1950) 

Professor Tadeusz JACZEWSKI (/nstitute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, 
Frankfurt a.M., Germany) (Sth July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitadt zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMOND (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) (12th August 
1953) (President) 

Professor Harold E. Voxes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) 
(12th August 1953) 

Professor Béla HANKO (Mezégazdasagi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y., 
U.S.A.) (12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. Hottnuis (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (1Sth October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum v Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KiHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

ee S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 
1954 

Professor Ernest MAYR (Museum of Comparative Zoology at Harvard College, Cambridge 
Massachusetts, U.S.A.) (4th December 1954). 

Professor Enrico TORTONESE (Museo do Storia Naturale ‘“‘ G. Doria”, Genova, Italy, 
(16th December 1954) 


DIRECTION 51 


REVISION OF THE ENTRY ON THE “ OFFICIAL LIST OF 
GENERIC NAMES IN ZOOLOGY” RELATING TO 
THE NAME “HOMARUS” WEBER, 1795 (CLASS 
~ CRUSTACEA, ORDER DECAPODA) (REVISION OF 
A RULING GIVEN IN “ OPINION ”° 104) 


RULING :—(1) The following revised entry in regard 
to the generic name Homarus Weber, 1795, is hereby 
inserted in the Official List of Generic Names in Zoology 
in. lieu of the incomplete entry made by the Ruling given 
in Opinion 104 :— 


Homarus Weber, 1795 (gender: masculine) (type 
species, (1) by selection by Rathbun (1904) of Cancer 
gammarus Linnaeus, 1758, a nominal species object- 
ively identical with Astacus marinus Fabricius, 1775, 
one of the nominal species originally included in 
this genus by Weber, and (2) through Declarations 
21 and 26: Cancer gammarus Linnaeus, 1758). 


- (2) The under-mentioned specific name is hereby placed 

on the Official List of Specific Names in Zoology with the 
Name No. 1059 :—gammarus Linnaeus, 1758, as pub- 
lished in the combination Cancer gammarus (specific 
name of type species of Homarus Weber, 1795) (Class 
Crustacea, Order Decapoda). 


(3) The under-mentioned generic name is hereby placed 
on the Official Index of Rejected and Invalid Generic 
Names in Zoology with the Name No. 756 :—Homarus 
Milne Edwards (H.), 1837 (a junior homonym of Homarus 
Weber, 1795). 


TART. «2 


4ncee 


296 OPINIONS AND DECLARATIONS 


(4) The under-mentioned specific names are hereby 
placed on the Official Index of Rejected and Invalid Specific 
Names in Zoology with the Name Nos. 361 and 362 
respectively :— 


(a) marinus Fabricius, 1775, as published in the com- 
bination Astacus marinus (a junior objective 
synonym of gammarus Linnaeus, 1758, as 
published in the combination Cancer gammarus) ; 


(b) vulgaris Milne Edwards (H.), 1837, as published in 
the combination Homarus yulgaris (a junior 
objective synonym of gammarus Linnaeus, 1758, 
as published in the combination Cancer gam- 
marus). 


I. THE STATEMENT (OF THE CASE 


On 6th December 1954, Mr. Francis Hemming, Secretary, 
submitted to the International Commission on Zoological 
Nomenclature two papers on different aspects of the same 
question, the first being concerned with the elucidation of a 
question of principle, the second with an individual case in which 
the problem had arisen in the course of the preparation of the 
Official List of Generic Names in Zoology for publication in book- 
form. In the first of these papers, to which was allotted the 
Registered Number Z.N.(S.) 8781, Mr. Hemming invited the 
Commission to consider the adoption of a Declaration clarifying 
RuJe (g) in Article 30 of the Rég/es in relation to the selection 
of the type species for a genus in a case where the nominal species 
so selected, though not itself cited at the time of the establishment 
of the genus in question, was objectively identical with another 


1 For particulars of the action taken by the International Commission on this 
application, see paragraph 3 of the Minute by the Secretary dated 23rd Febru- 
ary 1956, reproduced in paragraph 12 of the present Direction. 


- DIRECTION 51 297 


nominal species which was so cited. The second of the two 
papers submitted by Mr. Hemming was concerned with the 
question of the determination under Article 30 of the type species 
of the genus Homarus Weber, 1795, a case where the situation 
raised in the first of the two papers submitted had arisen in 
practice. Mr. Hemming’s paper regarding the type species of 
the genus Homarus Weber was as follows :— 


Proposed revision of the entry on the ‘‘ Official List of Generic Names 
in Zoology ’’ regarding the generic name ‘‘ Homarus ’’ Weber, 
1795 (Class Crustacea, Order Decapoda) (proposed revision 
of an entry made by the Ruling given in ‘‘ Opinion ”’ 104) 


By FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


The object of the present application is to ask the approval of the 
International Commission for a revision of the defective entry relating 
to the name Homarus Weber, 1795 (Nomencl. ent. Fabr. : 94) made on 
the Official List of Generic Names in Zoology by the Ruling given in 
Opinion 104 (1928, Smithson. misc. Coll. 73 (No. 5) : 25—28). 


2. The entry in respect of the foregoing name made on the Official 
List by Opinion 104 was as follows :— 


Homarus Fabr. in Weber, 1795a, 94, tsd. gammarus = marinus, s. 
vulgaris. Same as Milne Edw., 1837, HnC, 329, 333 


3. The first question which calls for consideration is the method by 
which a type species was fixed for the nominal genus Homarus Weber, 
1795. As will be seen from the foregoing extract from Opinion 104, 
it was there stated that this was by subsequent selection (t[ype by] 
s[ubsequent] d[esignation]). Weber’s little book, as is well known, 
was completely overlooked or ignored until Miss Mary Rathbun 
disinterred it and brought the names in it into use in her paper 
published in 1904 (Proc. biol. Soc. Wash. 17 : 170) when she stated 
that ‘“‘ Homarus gammarus (Linnaeus)’’, [i.e Cancer gammarus 
Linnaeus, 1758] was the type species of this genus. The foregoing 
nominal species was not included by Weber in his genus Homarus. 
The first nominal species cited by him as belonging to Homarus was, 
however, Astacus marinus Fabricius, 1775 (Syst. Ent. : 413), which is 
objectively indentical with the nominal species Cancer gammarus 
Linnaeus, 1758, the name published by Fabricius being no more than 
a substitute name (nom. noy.) for the earlier name published by 


298 OPINIONS AND DECLARATIONS 


Linnaeus. In 1912, Fowler (Ann. Rep. New Jersey State Mus. 1911 : 
333) selected the nominal species Astacus marinus Fabricius, 1775, 
one of the original included nominal species, to be the type species of 
Homarus Weber, 1795. 


4. Inanapplication numbered Z.N.(S.) 878? submitted simultaneously 
with the present case, I have recommended the International Com- 
mission to adopt a Declaration under which in a case such as the present 
where two objectively identical nominal species are involved, the 
selection of either as the type species of a genus established prior to 
Ist January 1931 is to be accepted as a valid type selection under 
Rule (g) in Article 30, it being immaterial whether the nominal species 
so selected is that actually cited by the author of the genus or whether 
it is only the nominal species which is objectively identical with that 
species. Accordingly, I now ask that consequentially upon the grant, 
as I hope, of the application referred to above, the Commission should 
accept as a valid type selection for the genus Homarus Weber, 1795, 
the selection by Miss Rathbun of the nominal species Cancer gammarus 
Linnaeus, 1758, notwithstanding the fact that it was not this species 
but the objectively identical nominal species Astacus marinus Fabricius, 
1775, which was cited by Weber as belonging to his genus Homarus. 


5. At this point it is necessary to refer to the Application Z.N.(S.) 9088 
(Bull. zool. Nomencl. 11 : 112—113) that the Commission should render 
a Declaration that, where there are two or more identical nominal 
species (i.e. nominal species, the names of which are objective synonyms 
of one another), the designation, indication or selection of any one 
of these nominal species to be the type species of a genus is to be 
treated as the designation, indication or selection of whichever of the 
nominal species concerned has the oldest available name, irrespective 
of whether or not that nominal species was cited by the author of the 
name of the genus in question. Further, it was suggested that the 
decision so recommended should be illustrated as follows by the case 
of the generic name Homarus Weber, 1795 :—‘‘ Example: The 
nominal species Cancer gammarus Linnaeus, 1758, and Astacus 
marinus Fabricius, 1775, are objectively identical with one another. 
The second, but not the first, of these nominal species was placed by 
Weber in his genus Homarus in 1795. Astacus marinus Fabricius was 
the first of the originally included nominal species to be selected to be 
the type species of Homarus Weber. Since the name Cancer gammarus 
Linnaeus is (a) an available name and (b) a senior objective synonym 
of the name Astacus marinus Fabricius, the nominal species Cancer 


2 See Footnote 1. 


8 For particulars of the action taken by the International Commission on this 
application, see paragraph 3 of the Minute by the Secretary dated 23rd Febru- 
ary 1956 reproduced in paragraph 12 of the present Direction. 


DIRECTION 51 299 


gammarus Linnaeus is to be treated as the type species of the genus 
Homarus Weber.”’ If, as I hope, the foregoing proposals are adopted 
by the Commission, the genus Homarus Weber, 1795, will take, as its 
type species, Cancer gammarus Linnaeus, 1758, and not, as otherwise 
would be the case, Astacus marinus Fabricius, 1775 (which, as explained 
above, is an identical nominal species but one possessing a later and 
therefore invalid name). 


6. The last point to which attention must be drawn is the cryptic 
reference to Milne Edwards (1837) in the entry regarding the generic 
name Homarus Weber made in Opinion 104. In the passage in question 
Milne Edwards adopted a nominal genus Homarus to which he referred 
a nominal species Homarus vulgaris Milne Edwards, 1837 (: 334), 
which, like Astacus marinus Fabricius, 1775, is objectively identical 
with Cancer gammarus Linnaeus, 1758. Milne Edwards made no 
reference whatever to Weber and clearly regarded himself as the 
author of the generic name Homarus as then used in his book. 
Homarus Milne Edwards (H.), 1837 (in Roret’s Suite a Buffon, Hist. 
nat. Crust. 2 : 333) must therefore be regarded as an independently 
established nominal genus. As such, its name (Homarus Milne 
Edwards) is invalid by reason of being a junior homonym of Homarus 
Weber, 1795. 


7. In order to bring the entry on the Official List of Generic Names in 
Zoology relating to the generic name Homarus Weber, 1795, into a 
form suitable for inclusion in that List when published in book form 
and in order also to dispose of certain other minor matters outstanding 
in the present case, I ask the International Commission to take the 
following action, namely :— 


(1) to substitute the following revised entry for Name No. 494 on 
the Official List of Generic Names in Zoology for the incomplete 
entry made under the Ruling given in Opinion 104 :— 


Homarus Weber, 1795, Nomencl. ent. Fabr.:94 (gender : 
masculine) (type species by selection by Rathbun (1904, 
Proc. biol. Soc. Wash. 17:170) of Cancer gammarus 
Einnaeps, 1758 (Syst. Nat. (ed. 10) 1 : 631), a nominal 
species objectively identical with Astacus marinus Fabricius, 
1775, one of the nominal species originally included in this 
genus by Weber : Cancer gammarus Linnaeus, 1758 (under 
the decision taken on Application Z.N.(S.) 908). 


Note :—The above proposal has been drafted on the 
assumption that approval will be given by the Com- 
mission to the proposal for the adoption of the Declarations 
recommended in Applications Z.N.(S.) 878 and Z.N.(S.) 
908. Ifthe first of these proposals were not to be approved, 

- the reference to the selection of the type species by Rathbun 


300 OPINIONS AND DECLARATIONS 


(1904) would need to be replaced by the words ‘‘ by 
selection by Fowler, 1912, Ann. Rep. New Jersey State 
Mus. 1911 : 333”. If the second of these proposals 
were not to be adopted, it would be necessary to cite the 
type species as Astacus marinus Fabricius, 1775, instead 
of as Cancer gammarus Linnaeus, 1758. 


(2) to place the under-mentioned specific name on the Official List 
of Specific Names in Zoology :—gammarus Linnaeus, 1758, as 
published in the combination Cancer gammarus (the specific 
name of the type species of Homarus Weber, 1795) ; 


Note :—See Note to Proposal (1) above. If the 
Application Numbered Z.N.(S.) 908 were to be rejected 
by the Commission, the words in brackets at the end of 
Proposal (2) would cease to be applicable. 


(3) to place on the Official Index of Rejected and Invalid Generic 
Names in Zoology the under-mentioned generic names, each 
of which is a junior homonym of Homarus Weber, 1795 :— 


(a) Homarus Milne Edwards (H.), 1837 ; . 
(b) Homarus Broun, 1881, Manual N. Zealand Coleopt. (1)(2) : 
740. 


(4) to place on the Official Index of Rejected and Invalid Specific 
Names in Zoology the under-mentioned specific names, each 
of which is a junior objective synonym of the name gammarus 
Linnaeus, 1758, as published in the combination Cancer 
gammarus :— 


(a) marinus Fabricius, 1775, as published in the combination 
Astacus marinus ; 


(b) vulgaris Milne Edwards (H.), 1837, as published in the 
combination Homarus vulgaris. 


Il. - THE SUBSEQUENT. HISTORY “OF THE CASE 


2. Registration of the present application : Upon the receipt of 
Mr. Hemming’s application, the question of the method by which 


DIRECTION 51 301 


the type species of the genus Homavus Weber, 1795, had been 
determined under Article 30 of the Régles was allotted the 
Registered Number Z.N.(S.) 879. 


3. Receipt of an application for a ‘* Declaration ’’ as to the 
nominal species to be accepted as the type species of a genus in 
the case where that nominal species is one of two or more identical 
species : Prior to the publication of the present application 
Mr. Hemming submitted to the Commission a proposal for the 
adoption of a further Declaration on a question allied to that 
raised in Application Z.N.(S.) 878 referred to in paragraph 1 
of the present Direction. In this second application, to which 
was allotted the Registered Number Z.N.(S.) 908, Mr. Hemming 
asked that, where one of two or more objectively identical 
nominal species is designated, indicated or selected as the type 
species of a genus, that genus should be cited as having as its 
type species the oldest established of the nominal species con- 
cerned. 


4. Publication of the present application : The present applica- 
tion was sent to the printer on 3lst December 1954 and was 
published in Part 4 of Volume 11 of the Bulletin of Zoological 
Nomenclature on 28th February 1955 (Hemming, 1955, Bull. zool. 
Nomencl. 11 : 114—116). 


5. No objection received: The publication of the present 
application and of the associated requests for the adoption of 
Declarations on the questions of principle involved elicited no 
objection of the action proposed from any source. 


6. Note on procedure prepared in anticipation of the submission 
of the present application to the Commission for decision : On 
4th November 1955, Mr. Hemming prepared for the consideration 
of the Commission the following note drawing attention to the 
fact that the form of the decision asked for would depend upon 
whether the Commission decided in favour of the adoption of the 


302 OPINIONS AND DECLARATIONS 


Declarations asked for respectively in Application Z.N.(S.) 878 
(paragraph | above) and Z.N.(S.) 908 (paragraph 3 above). The 
note so in question, which was as foJlows, was annexed as Note 4 
to the Voting Paper then prepared for submission to the Com- 
mission (paragraph 7 below) :— 


4. Procedural proposals : The proposal now submitted is that which 
has been prepared on the assumption that the Declarations recommended 
in Z.N.(S.) 878 (Voting Paper V.P.(55)23) and in Z.N.(S.) 908 (Voting 
Paper V.P.(55)30) are adopted by the Commission. The notes given 
in the last paragraph of the present application show the modifications 
in the present proposal which would follow automatically if the 
Commission were to decide against either or both of the suggested 
Declarations referred to above. 


III. THE-DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE © 


7. Issue of Voting Paper V.P.(55)31 : On 22nd November 1955, 
a Voting Paper (V.P.(55)31) was issued in which the Members 
of the Commission were invited to vote either for, or against 
“the proposal relating to the generic name Homarus Weber, 1795, 
as set out in Points (1) to (4) in paragraph 7 on page 116 of 
Volume 11 of the Bulletin of Zoological Nomenclature, subject to 
the alternatives on certain points of detail specified in the Notes 
to Points (1) and (2) referred to above, the adoption of these 
alternatives to depend on the decision reached by the Commission 
on Voting Papers V.P.(55)23 and V.P.(55)30 respectively, as 
explained in Note 4 overleaf”’ [i.e. in the Note numbered as 
above quoted in paragraph 6 of the present Direction]. 


8. The Prescribed Voting Period: As the foregoing Voting 
Paper was issued under the Three-Month Rule, the Prescribed 
Voting Period closed on 22nd February 1956. 


DIRECTION 5l 303 


9. Particulars of the Voting on Voting Paper V.P.(55)31 : At 
the close of the Prescribed Voting Period, the state of the voting 
on Voting Paper V.P.(55)31 was as follows :— 


(a) Affirmative Votes had been given by the following twenty- 
five (25) Commissioners (arranged in the order in which 
Votes were received) : 


Holthuis ; Hering; Riley; Vokes; Mayr; Jaczewski ; 
Lemche ; Prantl; Dymond; Esaki; Bodenheimer ; 
Mertens; do Amaral; Key; Boschma; Sylvester- 
Bradicy; Cabrera ;. Bonnet; Miller;  Tortonese; 
Kiuhnelt ; Bradley (J.C.) ; Stoll ; Hanko ; Hemming ; 


(b) Negative Votes : 


None ; 


(c) Voting Papers not returned : 


None. 


10. Declaration of Result of Vote : On 23rd February 1956, 
Mr. Hemming, Secretary to the International Commission, 
acting as Returning Officer for the Vote taken on Voting Paper 
V.P.(55)31, signed a Certificate that the Votes cast were as set 
out in paragraph 9 above and declaring that the proposal sub- 
mitted in the foregoing Voting Paper had been duly adopted 
and that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


11. Withdrawal of the proposal submitted in relation to the 
generic name ‘‘ Homarus ”’ Broun, 1881, on which, subsequent to 
the publication of the present application, action was taken by the 
International Commission by the Ruling given in ‘‘ Direction *’ 37 : 
On 29th December 1955, Mr. Hemming, as Secretary, after signing 


304 OPINIONS AND DECLARATIONS 


Direction 37, in which, inter alia, the Commission placed the 
generic name Homarus Broun, 1881, on the Official Index of 
Rejected and Invalid Generic Names in Zoology, executed the 
following Minute withdrawing from the scope of Application 
Z.N.(S.) 879 the proposal there submitted in regard to the fore- 
going name :— 


Withdrawal of the proposal relating to the generic name 
‘* Homarus ’’ Broun, 1881, a junior homonym of 
‘* Homarus ’> Weber, 1795 


By FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


The routine proposals submitted in the present case included a 
recommendation (in Point (3) of paragraph 7) that the name Homarus 
Broun, 1881, a junior homonym of Homarus Weber, 1795, should 
be placed on the Official Index of Rejected and Invalid Generic Names 
in Zoology. The same proposal was later included in a Paper Num- 
bered Z.N.(S.) 987, which was concerned with clearing up outstanding 
questions relating to the names of genera belonging to the Order 
Decapoda of the Class Crustacea placed on the Official List of Generic 
Names in Zoology in the period up to the end of 1936. This latter 
paper was submitted to the Commission under the One-Month Rule 
on 19th September 1955 with Voting Paper V.P.(O.M.)(55)28 and was 
approved by the Commission on Ist November 1955. 


2. In the light of the above decision the proposal relating to the 
disposal of the invalid name Homarus Broun, 1881, submitted with 
Paper Z.N.(S.) 987, together with the other proposals there submitted, 
has today been incorporated in the Ruling given in a Direction 
numbered Direction 37. 


3. In these circumstances no further action requires to be taken 
in connection with the name Homarus Broun, and the proposal in 
regard thereto in Application Z.N.(S.) 879 is accordingly withdrawn. 


12. Minute relative to the form of the Ruling to be given in the 
present case executed by the Secretary on 23rd February 1956 : 
On 23rd February 1956, Mr. Hemming, as Secretary, executed 
the following Minute containing directions as to the form of the 


DIRECTION 5] 305 


Ruling to be given in the Direction embodying the decision of the 
Commission in the present case :— 


Form of the Ruling to be given in the ‘‘ Direction’? embodying the 
decision by the International Commission in relation to the 
generic name ‘‘ Homarus ’’ Weber, 1795 (Class 
Crustacea, Order Decapoda) 


MINUTE by FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


Now that by its Vote on Voting Paper V.P.(55)31 the International 
Commission has approved the proposals in relation to the generic 
name Homarus Weber, 1795, submitted in Application Z.N.(S.) 879 
(1955, Bull. zool. Nomencl. 11 : 114—116), it is necessary to give 
consideration to the question of the alternative forms of decision 
indicated in the Notes attached to Points (1) and (2) in paragraph 7 
of the foregoing application to which attention was drawn in Note 4 
submitted with the Voting Paper referred to above. 


2. It will be recalled that, as pointed out in the foregoing Notes, 
the form of the decision to be recorded in the present case depends 
on the action taken by the Commission on two proposals then before 
it for the adoption of Declarations on questions of principle involved 
in this case. The proposals in question were the following :— 


(a) Application Z.N.(S.) 878 (Bull. zool. Nomencl. 11 : 86—89) 


In this application the Commission was asked to render a 
Declaration that, where there are two or more objectively identical 
nominal species, the designation, indication or selection of any of 
those species is to be accepted as conforming with the requirements 
of Article 30 of the Régles, irrespective of whether the nominal 
species concerned was cited by the original author when publishing 
the generic name in question. The decision to be taken on this 
application affects the decision to be taken in the case of Homarus 
Weber, for the first type selection for this genus (made by Rathbun 
in 1904) was that of Cancer gammarus Linnaeus, 1758, a nominal 
species which was not cited by Weber but which is objectively 
identical with Astacus marinus Fabricius, 1775, a nominal species 
which was cited by Weber. Under the rules hitherto obtaining 
Miss Rathbun’s type selection for this genus would be invalid but 
under the proposed Declaration it would be valid. 


306 OPINIONS AND DECLARATIONS 


(b) Application Z.N.(S.) 908 (Bull. zool. Nomencl. 11 : 112—113) 


In this application the Commission was asked to render a 
Declaration that, where one of two or more objectively identical 
nominal species is designated, indicated or selected as the type 
species of a genus, that genus shall be cited as having as its type 
species the oldest established of the nominal species concerned. 
This Declaration would affect the present case for under it the genus 
Homarus Weber, 1795, would have as its type species the nominal 
species Cancer gammarus Linnaeus, 1758, whereas under the rules 
hitherto obtaining it would be necessary to cite as the type species 
of this genus the nominal species Astacus marinus Fabricius, 1775, 
which is objectively identical with, but possesses a later name than, 
the nominal species Cancer gammarus Linnaeus, 1758. 


3. Both the foregoing applications have now been the subject of a 
vote by the Commission ; Application Z.N.(S.) 878 on Voting Paper 
V.P.(55)23 ; Application Z.N.(S.) 908 on Voting Paper V.P.(55)30. 
In each case the proposal submitted was approved by the Commission. 
The decision on Z.N.(S.) 878 has now been embodied in Declaration 
25! ; that on Z.N.(S.) 908 in Declaration 21°. 


4. By the decisions so taken the alternatives conditionally submitted 
in the Notes to Points (1) and (2) in paragraph 7 of Application 
Z.N.(S.) 879 cease to be applicable and accordingly the decision 
taken by the Commission on the foregoing application by its vote on 
Voting Paper V.P.(55)31 is to be interpreted as though those alternatives 
had not been put forward. 


13. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 29th May 1956, Mr. Hemming prepared the Ruling given in 
the present Direction and at the same time signed a Certificate 
that the terms of that Ruling were in complete accord with those 
of the proposa] approved by the International Commission in 
its Vote on Voting Paper V.P.(55)31, subject (1) to the withdrawal 
from the scope of the foregoing vote of the proposal relating to 
the name Homarus Broun, 1881, as directed in the Minute 
executed by the Secretary on 29th December 1955 and (ii) to the 
clarification in the Minute executed by that Officer on 23rd 
February 1956 of the situation regarding certain alternative 


* Declaration 25 was published on 14th August 1956 (Ops. Decls. int. Comm. 
zool. Nomencl. 13 : xxvii-xxxvi). 


5 Declaration 21 was published on 19th June 1956 (Ops. Decls. int. Comm. 
zool. Nomencl. 12 : i-viii). 


DIRECTION 51 307 


proposals submitted with the above Voting Paper on matters 
depending upon the decision to be taken by the Commission 
when voting on Voting Paper V.P.(55)23 and Voting Paper 
V.P.(55)30 respectively. The Minutes referred to above have 
been reproduced in paragraphs 11 and 12 respectively of the 
present Direction. 


14. Original References : The following are the original refer- 
ences for the names placed on Official Lists and Official Indexes 
by the Ruling given in the present Direction :— 


gammarus, Cancer, Linnaeus, 1758, Syst. Nat. (ed. 10) 1 : 631 


Homarus Milne Edwards (H.), 1837 (Roret’s Suite a Buffon) Hist. 
mar crust. 2 : 333 


marinus, Astacus, Fabricius (J.C.), 1775, Syst. Ent. : 415 


vulgaris, Homarus, Milne Edwards (H.), 1837 (Roret’s Suite a 
Buffon), Hist. nat. Crust. 2 : 334 


15. The following is the reference for the selection of a type 
species for the genus Homarus Weber, 1795, specified in the 
Ruling given in the present Direction :—Rathbun (M.), 1904, 
Proc biol. Soc. Wash. 17 : 170. 


16. Family-group-name problems : As in the case of previous 
Directions containing amplifications or corrections of entries 
made in the Official List of Generic Names in Zoology by Rulings 
given in Opinions rendered in the period up to the end of 1936, 
it has been decided that the question of the family-group names 
involved in the present case be deferred until it is possible to deal 
comprehensively with all the family-group-name problems 
involved in connection with the names of genera of the group 
concerned placed on the Official List in the foregoing period. 


17. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in dealing 


308 OPINIONS AND DECLARATIONS 


with the present case, and the present Direction is accordingly 
hereby rendered in the name of the said International Commission 
by the under-signed Francis Hemming, Secretary to the Inter- 
national Commission on Zoological Nomenclature, in virtue of 
all and every the powers conferred upon him in that behalf. 


18. The present Direction shall be known as Direction Fifty- 
One (51) of the International Commission on Zoological 
Nomenclature. 


Done in London, this Twenty-Ninth day of May, Nineteen 
Hundred and Fifty-Six. 


Secretary to the International Commission 


on Zoological Nomenclature 
FRANCIS HEMMING 


Printed in England by MetcatFe & Cooper LIMITED, 10-24 Scrutton St., London EC 2 


——<—— 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, cM.G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.15. Pp. 309—318 


DIRECTION 52 


Determination of the gender to be attributed to the generic 

names Cryptobranchus Leuckart, 1821 (Class Amphibia) 

and Eremias Wiegmann, 1834 (Class Reptilia) placed on 

the Official List of Generic Names in Zoology by the 
Ruling given in Opinion 92 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956 


Price Six Shillings and Sixpence 


(All rights reserved) { AP i? 2G 


Issued 5th December, 1956 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 52 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JORDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England). 


President : Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) 
(12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary : Mr. Francis Hemming (London, England) (27th July 1948) 


B. The Members of the Commission 


(arranged in order of precedence by reference to date of election or of most recent re-election 
as prescribed by the International Congress of Zoology) 


Professor H. BOSCHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. tare Tails (Universitetets Zoologiske Museum, Copenhagen, Denmark) (27th 

uly 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEy (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEwSKI (Jnstitute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (Sth July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitdt zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DYMOND (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) (12th August 
1953) (President) 

Professor Harold E. Vokes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) (12th 
August 1953) 

Professor Béla HANKO (Mezdgazdasdgi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y., 
U.S.A.) (12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. HoLtuuts (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (ASth October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum vy Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KUHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

tas Ne . S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 
195 

Professor Ernest MAyR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico TORTONESE (Museo di Storia Naturale ‘“‘ G. Doria”’’, Genova, Italy) 
(16th December 1954) 


DIRECTION 532 


DETERMINATION OF THE GENDER TO BE ATTRIBUTED 
TO THE GENERIC NAMES ‘°* CRYPTOBRANCHUS ”’ 
LEUCKART, 1821 (CLASS AMPHIBIA) AND 
** EFREMIAS ”? WIEGMANN, 1834 (CLASS REPTILIA) 
PLACED ON THE “* OFFICIAL LIST OF GENERIC 
NAMES IN ZOOLOGY” BY THE RULING 
GIVEN IN ‘** OPINION ” 92 


RULING :—(1) The gender to be attributed to the 
generic name Cryptobranchus Leuckart, 1821 (Class 
Amphibia), placed on the Official List of Generic Names 
in Zoology by the Ruling given in Opinion 92 is hereby 
determined as being the masculine gender. 


(2) The gender to be attributed to the generic name 
Eremias Wiegmann, 1834 (Class Reptilia) placed on the 
foregoing Official List by the Ruling given in the Opinion 
specified in (1) above is hereby determined as being the 
feminine gender. 


Pave, oUbIECE MATTER OF THE.PRESENT 
DIRECTION’ * 


On 23rd January 1956, Mr. Francis Hemming, as Secretary, 
submitted to the International Commission on Zoological 
Nomenclature the following paper containing proposals for the 
determination of the gender to be attributed to the generic names 
Cryptobranchus Leuckart, 1821 (Class Amphibia) and Eremias 


JAN - 8 1957 


Sy OPINIONS AND DECLARATIONS 


Wiegmann, 1834 (Class Reptilia), both of which had been placed 
on the Official List of Generic Names in Zoology :— 


Proposed determination of the gender to be attributed to 
the generic names ‘‘ Cryptobranchus ’’ Leuckart, 1821 
(Class Amphibia) and ‘‘ Eremias ’? Wiegmann, 
1834 (Class Reptilia) 


By FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


The Commission will recall that on 4th May 1955, in a paper 
numbered Z.N.(S.) 939, I submitted proposals for the determination 
of the gender to be assigned to the names of certain genera of fishes, 
cyclostomes, reptiles and amphibia which had been placed on the 
Official List of Generic Names in Zoology by Rulings given in Opinions 
adopted in the pre-Lisbon (1935) period. In that paper I explained 
that the gender properly attributable to the name Cryptobranchus 
Leuckart, 1821 (Class Amphibia) and Eremias Wiegmann, 1834 (Class 
Reptilia) were still under investigation and that proposals on this 
subject would be submitted as soon as the investigations then in hand 
had been completed (see Direction 30, published on 16th November 
1955, Ops. Decls. int. Comm. zool. Nomencl. 1(C) : 287—298). In 
the present paper the result of the foregoing investigations is sub- 
mitted to the Commission for consideration. 


(a) ‘* Cryptobranchus ”’ Leuckart, 1821 


2. In the Report on the gender attributable to each of the names 
placed on the Official List of Generic Names in Zoology in the period 
up to the end of 1936, initially obtained by the International Trust from 
Mr. F. J. Lelievre (at that time on the Classical Languages staff of 
Bedford College, London University) the masculine gender was 
assigned to the generic name Cryptobranchus Leuckart, 1821. The 
Copenhagen Rules contain no special provision as to the gender of 
nouns having a “*-branchus ”’ ending and for this reason and because 
of the “‘ -us ’® termination the masculine gender was suggested when this 
case was referred to Dr. W. I. Follett (Chairman of the Committee on 
Nomenclature of the American Society of Ichthyologists and Herpeto- 
logists) for advice. Dr. Follett replied on 15th February 1955 giving 
his reasons for regarding this name as being of the neuter gender, 
but adding that usage had been predominantly masculine (see Docu- 
ment (1) in Annexe A to the present paper). It was thereupon decided 
to seek the views (i) of Professor the Rev. L. W. Grensted, Consulting 
Classical Adviser to the Commission and (ii) of Commissioner Robert 


DIRECTION 52 313 


Mertens (Frankfurt a.M.). Each of these authorities replied that the 
acceptance of the masculine gender for this name was to be preferred 
on grounds both of grammar and usage. The letters received from 
Professor Grensted and Commissioner Mertens are reproduced as 
Documents (2) and (3) respectively in Annexe A to the present paper. 
Since the receipt of the foregoing correspondence a letter dated 9th 
December 1955 has been received from Dr. Follett reporting, as 
Chairman, that the Committee on Nomenclature of the American 
Society of Ichthyologists and Herpetologists as at present constituted 
‘has voted unanimously in favour of treating as masculine every 
generic name having the final term -branchus obviously derived from 
the corresponding Greek word’. In the same letter Dr. Follett 
furnished extracts from letters received from the three members of the 
Committee, whose views had not previously been reported to the 
Commission, namely :—(a) Dr. Robert Rush Miller (University of 
Michigan, Ann Arbor), who considers that the action proposed is the 
‘* wisest course’ ; (b) Dr. Jay M. Savage (Pomona College, Claremont, 
California), who states that he is in ‘‘ whole-hearted agreement ”’ ; 
(c) Mr. Norman J. Wilimovsky (Stanford University, California), who 
states that he ‘‘ will vote in favor ”’ of the action recommended. 


3. In the light of the advice received the Commission is recom- 
mended to give a Ruling that the masculine gender be attributed to 
the generic name Cryptobranchus Leuckart, 1821, in the entry relating 
thereto made on the Official List of Generic Names in Zoology by the 
Ruling given in Opinion 92. 


(b) ‘‘ Eremias ’’ Wiegmann, 1834 


4. The gender attributed to the generic name Eremias Wiegmann, 
1834, in Mr. Lelievre’s Report was the masculine gender. When, 
however, I consulted Mr. Follett, he replied that there had been an 
extensive usage of the feminine gender for this name. Professor 
Grensted and Commissioner Mertens were thereupon consulted in 
this case also. Both took the view this was a word of common gender 
which could be treated equally correctly either as masculine or feminine. 
Both recommended that usage should be followed in this case and 
therefore that the feminine gender should be adopted for this name. 
The letters so received are reproduced as Documents (1) and (2) 
respectively in Annexe B to the present paper. 


5. It is accordingly recommended that the Commission give a Ruling 
that the feminine gender be attributed to the generic name Eremias 
Wiegmann, 1834, in the entry relating thereto made on the Official 
List of Generic Names in Zoology by the Ruling given in Opinion 92. 


6. Summary : The recommendation now submitted is therefore that 
the gender to be attributed to the under-mentioned generic names 


314 OPINIONS AND DECLARATIONS 


placed on the Official List of Generic Names in Zoology by the Ruling 
given in Opinion 92 be the following :—(a) Masculine gender : Crypto- 
branchus Leuckart, 1821 (paragraph 3) ; (b) Feminine gender : Eremias 
Wiegmann, 1834 (paragraph 5). 


ANNEXE A 


The gender of the generic name ‘‘ Cryptobranchus ”’ Leuckart, 
1821 (Class Amphibia) 


Document (1) 


Extract from a letter dated \5th February 1955, from 
W. I. Follett (California Academy of Sciences, San Francisco, 
California, U.S.A.) 


I am disposed to regard the generic name Cryptobranchus Leuckart, 
1821, as of neuter gender for the following reasons : 


1. The Century Dictionary and Cyclopedia, Vol. 2, 1906, page 1380, 
states the derivation of Cryptobranchus as *“‘ NL., Gr. kpumros, hidden, 
+ Bpayyxos, in pl. equiv. to Bpayxia« gills ”’. 


2. Liddell and Scott, A Greek-English Lexicon, Vol. 1, 1940 (1948 
reprint), page 327, gives [line 14] Bpayy - .. . [line 19] -vov, ra, 
fin... . IL. inypl., gills of fishes . . . [line 29] IL. Bpayyos; ro, in ply 
Bpayx.a, Opp. H. 1. 160 ; but Bpayyxos, 6, in Ptol. Alm. 8. 1. 


3. While the last half-dozen words in the preceding line apparently 
represent an exceptional masculine usage, an older edition of this 
work (Liddell and Scott, A Greek-English Lexicon, Based on the 
German Work of Francis Passow, 1852, page 276), definitely stated 
that Bpayxos is neuter, thus: Ppdyxos, -e0s, tO=Bpdyxix, Opp. 
(akin to 6 Bpayyos). 


4. Roland Wilbur Brown, ‘“‘ Composition of Scientific Words ” 
1954, page 329 (under “ fin ’’) stated, ‘* Gr. branchos, n. fin, gill... ,” 
and on page 367 (under gill”), “Gr branchos neti eee 
On page 6, ““n”’ is stated to be the abbreviation for “‘ neuter ”’. 


I must add that in the event the foregoing views should be found 
to be correct, this fact must assuredly cause no sense of embarrassment 
on the part of the exceedingly capable teacher of classical languages 
who services you have employed on this project, since another word 
Bpayxos (meaning ‘“‘ hoarseness ’’) is in fact masculine, as indicated 
by Liddell and Scott, 1940, page 327, and it would obviously be a 


DIRECTION 52 315 


matter of mere conjecture for a non-zoologist to determine which of 
these two Greek words, of identical spelling, was intended by the 
author of the generic name. 


Document (2) 


Extract from a letter dated 12th July 1955, from L. W. Grensted 
(Consulting Classical Adviser to the Commission) 


Cryptobranchus : Here Dr. Follett has given all the evidence. The 
situation is curious. As the new Liddell and Scott Lexicon shows, 
the word Bpayxos only occurs twice, in two late writers, Oppianus and 
Ptolemaeus—in not over-reliable texts—and one, Oppianus, makes it 
neuter and one, Ptolemaeus, makes it masculine. I can see no reason 
for preferring Oppianus, except that parts of the body ending in -os 
are, in Greek, sometimes neuter and sometimes feminine. The evidence 
of Ptolemaeus undercuts the opinion of R. W. Brown, cited by 
Dr. Follett. So far as Classical evidence goes, as attested by the latest 
and best Greek Lexicon, -branchus may be either masculine or neuter. 


I think the Commission should give a decision based on current 
usage, citing the above evidence, which leaves the matter completely 
open. 


Oppianus and Ptolemaeus are approximately contemporaries— 
late second century (or very early third) A.D. Neither has any 
precedence. 


Document (3) 


Extract from a letter dated 13th July 1955, from Robert Mertens 
(Forschungs-Institut und Natur-Museum, Frankfurt a.M., Germany) 


Cryptobranchus is derived from the Greek word Bpdyxos. Since 
Bpayxos is masculine and Cryptobranchus is a latinised masculine form 
of it, I would regard Cryptobranchus as masculine, as it has hitherto 
universally been regarded. 


ANNEXE B 

The gender of the generic name ‘‘ Eremias ’? Wiegmann, 1834 
(Class Reptilia) 
Document (1) 


Extract from a letter dated 12th July 1955, from L. W. Grensted 
(Consulting Classical Adviser to the Commission) 


Eremias : This is coined on the model of Latin adjectival nouns, 
based on the Greek. It could be masculine or feminine according to 
its reference. Thus Pelias (a daughter of Pelias, masc.) is feminine. 


316 OPINIONS AND DECLARATIONS 


Pelias (a spear with a shaft from Pelion) is feminine, because hasta, 
a spear, is feminine. Eremias means a desert-dweller (not classical) 
coined from eremia (é€pynui«), desert. It would take its gender from the 
type of animal involved, serpens is m. of f., draco m., scorpio m., etc. 
etc. Again I think it is open to the Commission to decide by usage. 


There is, of course, a masculine presumption unless there is some 
feminine flavour in the animal intended. 


Document (2) 


Extract from a letter dated 13th July 1955, from Robert Mertens 
(Forschungs-Institut und Natur-Museum, Frankfurt a.M., Germany) 


Eremias means “living in solitude’? and is of Greek origin. 
According to our philologist colleagues Eremias can be used as 
masculine or feminine, but since it has hitherto been regarded as 
feminine I am in favour of this view being retained in future. 


2. Registration of the present application : Upon the receipt of 
Mr. Hemming’s Report the question of the determination of 
the gender to be attributed to the generic names Cryptobranchus 
Leuckart, 1821, and Eremias Wiegmann, 1834, was allotted the 
Registered Number Z.N.(S.) 1021. 


Il. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


3. Issue of Voting Paper V.P.(O.M.)(56)1 : On 23rd January 
1956 a Voting Paper (V.P.(O.M.)(56)1) was issued in which 
each Member of the Commission was asked to state (1) whether 
he agreed that, “in conformity with the General Directive 
relating to the recording in the Official List of Generic Names in 
Zoology of the gender of each name placed thereon prior to 1948, 
issued to the International Commission by the Thirteenth Jnter- 
national Congress of Zoology, Paris, 1948, the genders severally 
specified in paragraph 6 of the paper bearing the Registered 
Number Z.N.(S.) 1021 submitted by the Secretary simultaneously 


DIRECTION 52 317 


with the present Voting Paper [i.e. in the paragraph numbered 
as above in the Report reproduced in the first paragraph of the 
present Direction] be entered in the foregoing Official List as 
the gender attributable to the generic names Cryptobranchus 
Leuckart, 1821, and Eremias Wiegmann, 1834, respectively ”’, 
and (2), if he did not agree as regards either of the items included 
in the recommendations specified above, to indicate that item. 


4. The Prescribed Voting Period: As the foregoing Voting 
Paper was issued under the One-Month Rule, the Prescribed 
Voting Period closed on 23rd February 1956. 


5. Particulars of the Voting on Voting Paper V.P.(O.M.)(56)1 : 
At the close of the Prescribed Voting Period, the state of the 
voting on Voting Paper V.P.(O.M.)(56)1 was as follows :— 


(a) Affirmative Votes had been given by the following twenty- 
four (24) Commissioners (arranged in the order in which 
Votes were received) : 


Riley ; Boschma; Holthuis; Hering; Bodenheimer 
(Cryptobranchus only) ; Mayr ; Vokes ; Sylvester-Bradley ; 
Mertens ; Jaczewski; Kihnelt ; Key; Bradley (J.C.) ; 
Stoll; do Amaral; Dymond; Lemche; Hanko; 


Tortonese ; Hemming; Prantl; Bonnet; Cabrera; 
Miller ; 


(b) Negative Votes, one (1) (in part) : 


Bodenheimer (Eremias only) ; 


(c) Voting Papers not returned, one (1) : 


Esaki. 


6. Declaration of Result of Vote: On 28th February 1956, 
Mr. Hemming, Secretary to the International Commission, 


318 OPINIONS AND DECLARATIONS 


acting as Returning Officer for the Vote taken on Voting Paper 
V.P.(O.M.)(56)1, signed a Certificate that the Votes cast were as 
set out in paragraph 5 above and declaring that the proposal 
submitted in the foregoing Voting Paper had been duly adopted 
and that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


7. Preparation of the Ruling given in the present ‘* Direction ”’ : 
On 12th June 1956, Mr. Hemming prepared the Ruling given 
in the present Direction and at the same time signed a Certificate 
that the terms of that Ruling were in complete accord with those 
of the proposal approved by the International Commission in its 
Vote on Voting Paper V.P.(O.M.)(56)1. 


8. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in dealing 
with the present case, and the present Direction is accordingly 
hereby rendered in the name of the said International Commission 
by the under-signed Francis Hemming, Secretary to the Inter- 
national Commission on Zoological Nomenclature, in virtue of 
all and every the powers conferred upon him in that behalf. 


9. The present Direction shail be known as Direction Fifty- 
Two (52) of the International Commission on Zoological 
Nomenclature. 


Done in London, this Twelfth day of June Nineteen Hundred 
and Fifty-Six. 


Secretary to the International Commission 


on Zoological Nomenclature 


FRANCIS HEMMING 


Printed in England by MercatFe & CooPEeR LimITED, 10-24 Scrutton St., London EC 2 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, c.M.G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.16. Pp. 319—336 


DIRECTION 55 


Insertion in the Official List of Generic Names in Zoology 

of an entry relating to the generic name Grus Brisson, 

1760 (Class Aves) (correction of an error in the Ruling 
given in Opinion 103) | 


LONDON: 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956 


Price Twelve Shillings 


(All rights reserved) 


Reed 20h December,1956. 42 2«@€ + «#8 | SEAM =al 


INTERNATIONAL COMMISSION ON > 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 55 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JORDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President: Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., 
U.S.A.) (12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August i953) 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(Arranged in order of precedence by reference to date of election or of most recent 
re-election, as prescribed by the International Congress of Zoology) 


Professor H. BOSCHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. Henning LEMcHE (Universitetets Zoologiske Museum, Copenhagen, Denmark) 
(27th July 1948) 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEy (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (Sth July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitat zu 
Berlin, Germany) (5th July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMOND (University of Toronto, Toronto, Canada) (12th August 1953) 

aes Js oa BRADLEY (Cornell University, Ithaca, N. Y., U.S.A.) (12th August 1953) 

resident 

Professor Harold E. Vokes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) 
(12th August 1953) 

Professor Béla HANK6 (Mezdégazdasagi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y., U.S.A.) 
(12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. Hoituuis (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (15th October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, Berkeley, 
California, U.S.A.) (29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum v Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KUHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

aa F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 

Professor Ernst MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico TORTONESE (Museo di Storia Naturale, ‘*‘ G. Doria,’’ Genova, Italy) 
(16th December 1954) 


DIRECTION 55 


INSERTION IN THE “OFFICIAL LIST OF GENERIC 
NAMES IN ZOOLOGY” OF AN ENTRY RELATING 
TO THE GENERIC NAME “ GRUS ” BRISSON, 1760 

(CLASS AVES) (CORRECTION OF AN ERROR IN 
THE RULING GIVEN IN “ OPINION” 103) 


RULING :—(1) The entry relating to the generic name 
Grus Pallas, 1767 [sic], made on the Official List of Generic 
Names in Zoology by the Ruling given in Opinion 103 
is hereby deleted as being incorrect and the following 
revised entry bearing the same Name Number (Name No. 
474) is hereby made in its place :— 


Grus Brisson, 1760 (gender: feminine) (type species, 
by absolute tautonymy : Ardea grus Linnaeus, 1758) 


(2) The under-mentioned specific name is hereby placed 
on the Official List of Specific Names in Zoology with the 
Name No. 1062 :—grus Linnaeus, 1758, as published in 
the combination Ardea grus (specific name of type species 
of Grus Brisson, 1760). 


(3) The under-mentioned generic name is hereby placed 
on the Official Index of Rejected and Invalid Generic Names 
in Zoology with the Name No. 764 :—Grus Pallas, 1766 
(a junior homonym of Grus Brisson, 1760). 


(4) The under-mentioned specific name is hereby 
placed on the Official Index of Rejected and Invalid 
Specific Names in Zoology with the Name No. 364 :— 
psophia Pallas, 1767, as published in the combination 
Grus psophia (a junior objective synonym of crepitans 
Linnaeus, 1758, as published in the combination Psophia 
crepitans). 


JAN - 8 1957 


B22 OPINIONS AND DECLARATIONS 


I. THE SUBJECT MATTER: OF THE PRESENT 
“DIRECTION ” 


On 19th December 1955, Mr. Francis Hemming, as Secretary 
to the International Commission on Zoological Nomenclature, 
communicated to twenty ornithologists in various countries a 
note regarding certain errors which had been detected in the 
Ruling in regard to the generic name Grus Pallas, 1767 [sic] 
(Class Aves) given in Opinion 103. In the letter covering this 
communication Mr. Hemming invited the specialists concerned 
to advise the Commission as to which of two alternative courses 
of action they would advise it to take for the purpose of regularis- 
ing the position as regards the foregoing generic name. The 
alternatives so submitted, which were set out in paragraph 8 
of the paper distributed by Mr. Hemming, were as follows :— 


8. From the particulars given above there are, it will be seen, two 
ways by which to secure that the nominal species Ardea grus Linnaeus, 
1758, shall be the type species of the genus Grus, though under neither 
of these could that name be attributed to Pallas, 1767, the attribution 
given in Opinion 103. The possible alternatives are :— 


(a) to apply the ordinary provisions of the Régles and in consequence 
to accept the name Grus Brisson, 1760, with Ardea grus 
Linnaeus, 1758, as type species by absolute tautonymy ; 


(b) to use the Plenary Powers (i) to suppress the name Grus Brisson, 
1760, and (ii) to designate Ardea grus Linnaeus, 1758, to be 
type species of Grus Pallas, 1766, thus retaining the attribution 
of this name to Pallas, the author cited in Opinion 103, though 
with a different date and with a reference to a different work. 


2. Registration of the present application : At the time of the 
discovery by Mr. Hemming of the defects in the ruling relating 
to the generic name Grus Pallas, 1767 [sic] given in Opinion 103 
the question of the action to be taken to remedy the errors so 


uncovered was allotted the Registered Number Z.N.(S.) 558. 


3. Summary of the replies received from specialists to the 
questionnaire issued in December 1955: Replies were received 
from fourteen (14) of the twenty-four (24) specialists to whom the 
questionnaire referred to in paragraph 1 above was issued in 
December 1955. Of the specialists who furnished statements of 


DIRECTION 55 323 


their views eleven (11) expressed themselves as being in favour 
of Alternative ‘““A’’ (adoption of the generic name Grus as from 
Brisson, 1760, the author by whom it had been first published) 
and three (3) as being in favour of Alternative “* B” (use of the 
Plenary Powers to retain as from 1766 the authorship of the 
name Grus for Pallas, the author to whom it had been credited 
in Opinion 103). Ten (10) of the specialists consulted did not 
reply to the questionnaire issued. 


4. Replies from the eleven specialists who favoured the acceptance 
of the generic name ‘* Grus ”’ as from Brisson, 1760 (Alternative 
**A’’): The replies received from the eleven specialists who 
favouted the acceptance of the generic name Grus as from 
Brisson, 1760 (Alternative “‘A’’) were as follows :— 


(1) Richard Meinertzhagen (London) (21st December 1955) 


I prefer your course “‘A’”’ accepting the name Grus Brisson, 1760, 
with Ardea grus Linneaus, 1758, as type species. 


(2) W. B. Alexander (Swanage, England: formerly of the 
Edward Grey Institute of Field Ornithology, Oxford, 
England) (2\st December 1955) 


My opinion is that the best procedure would be to accept the name 
Grus Brisson, 1760, with Ardea grus Linnaeus as type. 


(3) Noél Mayaud (Faculté des Sciences de Paris) (22nd 
December 1955) | 


Les noms de genre de Brisson étant valides, Grus Brisson, 1760, 
préoccupe Grus Pallas, 1766. 


En conséquence le nom générique des grues doit étre Grus Brisson, 
1760. Type pas tautonymie absolue Ardea grus Linné, 1758, d’apres 
*“La Grue” Brisson et la premiére référence donnée par Brisson : 
Grus linné Syst. Nat. ed. 10. gen. 76. sp. 4 et la planche coloriée n° 199 
de Martinet (Atlas des planches coloriées de Brisson). 


C’est donc Valternative “‘A”’ qui doit étre adoptée. Elle a l’avantage 
(a) de respecter strictement la loi de priorité et (b) de respecter le 
logique car Grus Brisson désigne sans conteste possible les Grues 
(Cranes) sensu stricto, tandis que Grus Pallas s’applique 4 un oiseau 
qui ne fait pas partie de ce genre. If faudrait donc outrepasser l’idée 
de Pallas pour appliquer Grus aux Grues (Cranes) sensu stricto chose 


$24 OPINIONS AND DECLARATIONS 


bien inutile quand on a le genre Grus Brisson, antérieur, qui s’applique 
parfaitement bien. 


(4) Jean Delacour (Los Angeles County Museum, Los Angeles, 
California) (27th December 1955) 


The main point is to preserve the generic name Grus and it seems 
to me that the first proposition (accepting the name Grus Brisson, 
1760) is the simplest. However, I would agree to suppress Grus 
Brisson and to support Grus Pallas, 1766, if it was the opinion of the 
majority. 


(5) Helmuth Wagner (Ubersee-Museum, Bremen, Germany) 
(31st December 1955) 


Having considered the question with other members of our staff 
we conclude that the point would be whether the regulation adopted 
in Opinion 103 is to be regarded as a definite settling of the status of 
the generic name Grus or not. Since we realize from your letter that 
Opinions are liable to corrections if evidence is given of their being 
apparently based on incomplete information we would tend to prefer 
a way similar to that you mention under “‘A”’ in your note. If the names 
of Brisson are to be regarded as available there seems to be no reason 
to discuss further the status of Grus Pallas as relevant to the establishing 
of that genus. The exact date of Grus Pallas and its type species has, 
however, to be determined too. The two alternatives seem to be :— 


(a) to adopt Grus Pallas, 1766, in correction of Grus Pallas, 1767, 
and to suppress Grus Brisson, 1760, which name has the unquestionable 
rank of priority, or 


(b) to consider Grus as being established by Brisson in 1760 in 
accordance with both the Law of Priority and the decision of 1950 
as to the status of Brisson’s names, and to place Grus Pallas into 
synonymy, while correcting the date of that name as necessary. 


As (a) would involve the suppression of a name otherwise valid 
but (b) would account for the Law of Priority and do no harm but 
placing Grus Pallas into synonymy, we would, other things being 
equal, suggest following the course as outlined by you under “A”’ 
provided that the date of Grus Pallas is corrected which would be 
necessary for either way to be adopted, i.e. a course as outlined above 
under (b). 


(6) G. C. A. Junge (Rijksmuseum van Natuurlijke Historie, 
Leiden) (2nd January 1956) 


I am of the opinion that alternative “‘A” is the most logical and 
correct one. Thus I vote: to apply the ordinary provisions of the 


DIRECTION 55 325 


Régles and in consequence to accept the name Grus Brisson, 1760, with 
Ardea grus Linnaeus, 1758, as type species by absolute tautonymy. 


(7) Finn Salomonsen (Universitetets Zoologiske Museum, 
Kobenhayn) (6th January 1956) 


I am in agreement with the first alternative, i.e. to accept Grus 
Brisson, 1760, with Ardea grus Linnaeus, 1758, as type species by 
absolute tautonymy. This solution has the following advantages :— 


(a) the ordinary provisions of the Régles can be applied ; 
(b) Brisson’s Grus has priority over Pallas’s Grus ; 


(c) Brisson’s Grus is undoubtedly a designation for the Cranes, 
while Pallas’s Grus contained other, non-related birds, and the accept- 
ance of his name might give rise to unnecessary discussions and 
conflicts. 


(8) K. H. Voous (Zodlogisch Museum, Amsterdam) (6th 
January 1956) 


I am much in favour of the solution mentioned by you under “‘A”’’, 
that is, to apply the ordinary provisions of the Rég/es and in conse- 
quence to accept the name Grus Brisson, 1760, with Ardea grus 
Linnaeus, 1758, as type species by absolute tautonymy. 


Whatever may happen I should have to protest severely if the generic 
name Grus, which has been in constant use for so long a period, should 
be proposed to be rejected and replaced by another genus name. 


(9) J. Steinbacher (Senckenbergische Naturforschende Gesell- 
schaft, Frankfurt a.M., Germany) (9th January 1956) 


I wish to inform you that I would like to give my consent to the 
first alternative, i.e. to apply the ordinary provisions of the Régles 
and in consequence to accept the name Grus Brisson, 1760, with 
Ardea grus Linnaeus, 1758, as type species by absolute tautonymy. 


(10) B. Rensch (Zoologisches Institut der Westfalischen 
Wilhelms-Universitat, Miinster, Germany) (10th Janu- 

ary 1956) 
Meines Erachtens ist eine eindeutige Entscheidung in diesem Falle 


ohnehin nicht méglich, da man fiir beide Entscheidungen gute Griinde 
angeben kann. Im Ganzen halte ich es aber doch fiir zutreffend die 


326 OPINIONS AND DECLARATIONS 


unter 8 (a) getroffene Entscheidung zu bevorzugen, d.h. die Namen 
Grus Brisson, 1760, anzunehmen mit Ardea grus Linnaeus, 1758, als 
typische Art. 


(11) Ernst Mayr (Museum of Comparative Zoology at Harvard 
College, Cambridge, Massachusetts, U.S.A.) (26th 
January 1956) 


For several reasons I would prefer to accept the name Grus of 
Brisson, 1760, which is not only what had been done by ornithologists 
for many years (until someone thought one had to ignore the Brisson 
names), but it also eliminates a number of difficulties associated with 
the name Grus Pallas, 1766. Such an acceptance of the Brisson name 
would be distinctly in line with the recent trend to revert to the tradi- 
tional acceptance of Brissonian generic names. It seems to me, 
therefore, that there are four independent reasons for entering Brisson 
in the Official List as the author of Grus :—(1) priority, (2) the traditional 
acceptance of Brisson as the author during most of the eighteenth and 
nineteenth centuries, (3) the avoidance of the difficulties associated 
with Grus Pallas, and (4) the recent trend to restore Brissonian names 
to their formerly held rank. 


5. Replies from the three specialists who favoured the use by 
the International Commission of its Plenary Powers to secure that 
the generic name ‘* Grus ’’ should rank from Pallas, 1766 (Alter- 
native ‘*‘ B ’’) : The replies received from the three specialists who 
favoured the use by the International Commission of its Plenary 
Powers for the purpose of securing that the generic name Grus 
should rank from Pallas, 1766, instead of from Brisson, 1760 
(Alternative “‘B’’ were as follows :— 


(1) G. Niethammer (Zoologisches Forschungsinstitut und 
Museum Alexander Koenig, Bonn, Germany) (22nd 
December 1955) 


I agree with your second alternative “* B’’, i.e. Grus Pallas, 1766. 


(2) Alden H. Miller (Museum of Vertebrate Zoology, University 
of California, Berkeley, California) (28th December 
1955) 


I would favor your alternative “‘B”’. It seems to me this pro- 
cedure would cause the least change in current usage and not re-open 
old questions concerning Brisson. The use of Grus of Pallas is in 
line with very thoroughly established handbooks like the British 
handbook, AOU and Peters. 


DIRECTION 55 327 


~ . (3) G. Diesselhorst (Zoologische Sammlung des Bayerischen 
. Staates, Miinchen, Germany) (Sth February 1956) 


Having studied the problem in question I should think it least 
troublesome for the stability of nomenclature to use the Plenary 
Powers for suppressing the name Grus Brisson, 1760, and to designate 
Ardea grus Linnaeus, 1758, to be the type species of Grus Pallas, 1766, 
as in Alternative ‘“‘ B”’ of your sheet. 


6. Submission to the International Commission on Zoological 
Nomenclature in April 1956 of proposals for rectifying the defects 
in the Ruling relating to the generic name ‘‘ Grus ’’ Pallas, 1767 
[sic] given in ‘* Opinion ”’ 103 : On 25th April 1956, Mr. Hemming 
submitted to the International Commission a paper 1n which, 
after drawing attention to the defects in the Ruling relating to the 
generic name Grus Pallas, 1767 [sic] given in Opinion 103 and 
giving particulars of the views expressed by the specialists who 
had been consulted in this matter (paragraph 3 above), he set 
out two alternative courses of action which the Commission, 
if it so desired, might take for the purpose of regularising the 
entry regarding the foregoing name made on the Official List of 
Generic Names in Zoology by the Ruling given in the Opinion 
referred to above. The alternatives so set out corresponded with 
those which Mr. Hemming had previously placed before the 
specialists whom he had consulted in this case, i.e. those quoted 
in the first paragraph of the present Direction. The paper sub- 
mitted to the Commission by Mr. Hemming was as follows :— 


Proposed correction of an erroneous entry on the ‘‘ Official List of 
Generic Names in Zoology’’ in regard to the generic name 
** Grus ’’ (Class Aves) made by the Ruling given in 
** Opinion ’’ 103 


By FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


Among the Opinions rendered by the International Commission in 
the pre-Lisbon (1935) period in which errors have been detected in 
course of the preparation of the Official List of Generic Names in 
Zoology for publication in book form is Opinion 103 in which the 
generic name Grus Pallas, 1767, was placed on the above List with 
Ardea grus Linnaeus, 1758 (a non-included species) as type species. 


328 OPINIONS AND DECLARATIONS 


The purpose of the present paper is to lay the relevant facts before 
the Commission and to seek a decision on the action to be taken to 
provide a valid entry on the Official List in regard to the foregoing name. 


2. The following are the relevant facts in the present case :— 


(1) The generic name Grus was first published by Brisson in 1860 
(Ornithologia 5 : 374—391). Brisson, whose Ornithologia has 
been validated by the Commission under its Plenary Powers 
by the Ruling given in Direction 16 (1955, Ops. Decls. int. 
Comm. zool. Nomencl. 1(C) : 81—88), used the name Grus 
for a subgenus of the genus Ciconia. He placed a number 
of species in Grus, one of these being Ardea grus Linnaeus, 
1758 (Syst. Nat. (ed. 10) 1: 141). Brisson did not designate 
or indicate a type species for Grus under Rules (a), (b) or (c) 
in Article 30 and in consequence under Rule (d) in that Article 
Ardea grus Linnaeus, 1758, is the type species of Grus Brisson, 
1760, by absolute tautonymy. 


(2) In 1766 the name Grus was published as an apparently new name 
by Pallas (Miscell. Zool. : 65) . The main purpose of Pallas’s 
paper, which was headed ‘‘ GRUS CREPITANS seu PSOPHIA 
LINNEI”’, was to describe a specimen of the species to which 
in 1758 (: 154) Linnaeus had given the name Psophia crepitans. 
This was the only nominal species which he included in his 
genus Grus and is therefore the type species by monotypy. 


(3) In 1767 (Spicil. Zool. 4:1) Pallas again used the name Grus. 
This paper was substantially the same as that which he had 
published a year earlier in the Miscell. Zool., being also con- 
cerned with the description of a specimen of the species which 
in 1758 Linnaeus had named Psophia crepitans. In this paper 
Pallas renamed this species Grus psophia (: 1), presumably 
because, although he was rejecting the word “‘ Psophia”’ 
as a generic name, he wished to perpetuate in some way the 
word used by Linnaeus for this species in 1758 and thought 
that this might conveniently be done by employing it as a 
specific name in place of the name crepitans. 


(4) On some date not now known the Commission adopted Opinion 
103 (published in 1928 (Smithson. misc. Coll. 73 No. 5) : 21— 
24), in which it placed the generic name Grus Pallas, 1767 
(Spicil. Zool.) on the Official List of Generic Names in Zoology 
with Ardea grus Linnaeus, 1758, as type species by absolute 
tautonymy. In the paper on which this Opinion was based 
no reference was made either to the earlier name Grus Brisson, 
1760, or to the earlier usage by Pallas himself of the name 
Grus in the Miscell. Zool. of 1766. These matters therefore 
were not considered by the Commission. It was however 


DIRECTION 55 329 


aware that the nominal species Ardea grus Linnaeus, 1758, 
was not mentioned by Pallas in his 1767 paper and therefore 
that it was not an originally included species of the genus 
Grus. An attempt was made to get over this difficulty by 
arguing that in establishing the genus Grus, Pallas must have 
been aware that in 1758 Linnaeus had used the term “*‘ Grues ”’ 
to denote four species (of which one was Ardea grus) placed 
by him in the genus Ardea and therefore that it would be 
reasonable to infer that Pallas looked upon Ardea grus Linnaeus 
as belonging to his (Pallas’s) genus Grus of 1767. This argu- 
ment is invalid since the only species which are eligible for 
selection as the type species of a genus established before 
1931 without a designated or indicated type species are the 
nominal species cited (including nominal species cited in 
synonymy) by the author of the generic name at the time when 
that name was first published. 


3. The question now to be considered is what action the Commission 
should take to regularise the situation created by the Ruling given 
in Opinion 103. Broadly speaking, there are two courses open to the 
Commission, the adoption of either of which would ensure the con- 
tinued use of the generic name Grus in its accustomed sense. First, 
the Commission could accept the name Grus as from Brisson, 1760, 
the first occasion on which it was validly published, the type species 
in this case being Ardea grus Linnaeus, 1758, the species currently 
accepted as the type species of the genus Grus. Second, the Com- 
mission could use its Plenary Powers to validate Grus as from Pallas, 
1766, with the above species as type species by (a) suppressing Grus 
Brisson, 1760, and (b) designating Ardea grus Linnaeus as the type 
species of Grus Pallas, so validated. This second course would 
substantially validate the incorrect Ruling in Opinion 103, the only 
difference being that Grus would rank from Pallas’s Miscell. Zool. 
of 1766 instead of from that author’s Spicil. Zool. of 1767. If it 
were desired to validate this further mistake in Opinion 103, it would 
be necessary for the Commission to suppress Grus Pallas, 1766, as 
well as Grus Brisson, 1760, a course which it can hardly be supposed 
would find any support. 


4. In order to test the views of specialists, a note summarising the 
unfortunate history of this case and setting out the two main alterna- 
tives outlined above, was addressed in December 1955 to twenty 
ornithologists (who on previous occasions had shown themselves 
interested in nomenclature by furnishing comments), under cover of 
a letter asking them to state for the information of the Commission 
which of the foregoing alternatives it was, in their opinion, desirable 
that the Commission should adopt. Replies have now been received 


330 OPINIONS AND DECLARATIONS 


from fourteen of the specialists consulted. These replies may be 
classified as follows :— 


(1) In favour of applying the normal provisions of the “ Régles”’ 
that is, in favour of accepting “* Grus”’ from Brisson, 1760, 
with ‘‘Ardea grus’”’ Linnaeus, 1758, as type species (eleven (11) 
specialists) : ; 
W. B. Alexander (Edward Grey Institute of Field Ornithology, Oxford, 
England) 
Jean Delacour (Los Angeles County Museum, Los Angeles, California, 
U.S.A.) 
G. C. A. Junge (Rijksmuseum van Natuurlijke Historie, Leiden, The 
Netherlands) 


Noél Mayaud (Faculté des Sciences de Paris) 


Ernst Mayr (Museum of Comparative Zoology at Harvard College, 
Cambridge, Massachusetts, U.S.A.) 


Richard Meinertzhagen (London) 


B. Rensch (Zoologisches Institut der Westfalischen Wilhelms- Universitat, 
Miinster, Westf., Germany) 


Finn Salomonsen (Universitetets Zoologiske Museum, Copenhagen) 


Joachim Steinbacher (Senckenbergische Naturforschende Gesellschaft, 
Frankfurt a.M., Germany) 


Helmuth Wagner (Ubersee-Museum, Bremen, Germany) 
K. H. Voous (Zoologisch Museum, Amsterdam, The Netherlands) 


(2) In favour of using the Plenary Powers (a) to validate the name 
‘*“Grus”’ as from Pallas, 1766, and (b) to designate ‘Ardea 
grus”’ Linnaeus, 1758, to be the type species of the foregoing 
genus (three (3) specialists)? 

G. Diesselhorst (Ornith. Abteilung, Sammlung des Baverischen Staates, 

Miinchen, Germany) 


Alden H. Miller (Museum of Vertebrate Zoology, University of California, 
Berkeley, U.S.A.) 


G. Niethammer (Zoologisches Forschungsinstitut und Museum Alexander 
Koenig, Bonn, Germany) 


5. It will be seen from the above particulars that by a large majority 
(11 to 3) the ornithologists who have advised on this case support 
the acceptance of the generic name Grus as from Brisson, 1760, the 
first author by whom it was published, and are opposed to the use of 
the Plenary Powers for the purpose of validating this name as from 
its later usage by Pallas in 1766. | 


' For the communications received from the eleven specialists here enumerated 
see paragraph 4 of the present Direction. 

2. For the communications received from the three specialists Het enumerated 
see paragraph 5 of the present Direction. 


DIRECTION 55. 33] 


6. The present is one of the cases concerning defective entries made 
in the Official List of Generic Names in Zoology by Rulings given in 
Opinions rendered in the pre-1935 (Lisbon) period on which a decision 
is needed before the volume containing particulars of the entries so 
far made in the Official List can be published. It is accordingly 
submitted to the Commission for consideration in the light of the views 
which have been collected from specialists as set out in paragraph 4 
above. 


7. The Commission is accordingly invited to choose between the 
following two alternatives :— 


(1) ALTERNATIVE “A”: Under this Alternative the normal 
provisions of the Régles would be applied in this case, the 
oldest available name (Grus Brisson, 1760) being adopted for 
the genus concerned. The Ruling which would result from the 
adoption by the Commission of this Alternative is shown in 
Section (1) of the Annexe attached to the present note. 


(2) ALTERNATIVE “ B” : Under this Alternative the Commission 
would use its Plenary Powers (a) to suppress the oldest available 
name (Grus Brisson, 1760) for the genus concerned and 
(b) to designate Ardea grus Linnaeus, 1758, to be the type species 
of Grus Pallas, 1766, which would then become the oldest 
available name for the genus in question. The Ruling which 
would result from the adoption by the Commission of this 
Alternative is shown in Section (2) of the Annexe attached 
to the present note. 


8. Supplementary action required: Under the General Directive 
issued to the International Commission by the Thirteenth International 
Congress of Zoology, Paris, 1948, that in its Opinions it is to deal 
with all names involved in cases submitted to it, the names Psophia 
Linnaeus, 1758, and crepitans Linnaeus, 1758, as published in the com- 
bination Psophia crepitans, should be placed on the appropriate 
Official Lists and the name psophia Pallas, 1767, as published in the 
combination Grus psophia (an invalid nom. nov. for crepitans Linnaeus, 
1758) should be placed on the Official Index of Rejected and Invalid 
Specific Names in Zoology. The required action has already been taken 
or is in process of being taken as regards the first two of the above 
names, for the name Psophia Linnaeus, 1758, was placed on the Official 
List of Generic Names in Zoology by the Ruling given in Opinion 67, 
while by a vote recently taken on Voting Paper V.P.(O.M.)(55)31 the 
Commission has decided to place the name crepitans Linnaeus, 1758, 
as published in the combination Psophia crepitans, on the Official List 
of Specific Names in Zoology. The decision so taken has been embodied 
in Direction 43, which it is expected will be published at an early date.® 
The only action now called for is therefore the addition of the invalid 
specific name psophia Pallas, 1767, to the Official Index. 


8 Direction 43 was published on 19th October 1956 (Ops. Decls. int. Comm. 
zool. Nomencl. 1(D) : 171—210). 


332 OPINIONS AND DECLARATIONS 
ANNEXE ' 


Rulings regarding the generic name ‘‘ Grus ’’ which would result from 
the adoption by the International Commission of Alternative ‘‘A’’ 
and of Alternative “ B’’ respectively 


Section (1) of Annexe 


ALTERNATIVE, “Ay 


Draft Ruling applying the normal provisions of the 
** Régles ”’ 


DRAFT RULING :—(1) The entry relating to the generic name 
Grus Pallas, 1767 [sic], made on the Official List of Generic Names 
in Zoology by the Ruling given in Opinion 103 is hereby deleted as 
being incorrect and the following revised entry is hereby made in its 
place :—Grus Brisson, 1760 (gender: feminine) (type species, by 
absolute tautonymy : Ardea grus Linnaeus, 1758). 


(2) The under-mentioned name is hereby placed on the Official 
List of Specific Names in Zoology :—grus Linnaeus, 1758, as published 
in the combination Ardea grus (specific name of type species of Grus 
Brisson, 1760). 


(3) The under-mentioned name is hereby placed on the Official 
Index of Rejected and Invalid Generic Names in Zoology :—Grus 
Pallas, 1766 (a junior homonym of Grus Brisson, 1760). 


(4) The under-mentioned specific name is hereby placed on the 
Official Index of Rejected and Invalid Specific Names in Zoology :— 
psophia Pallas, 1767, as published in the combination Grus psophia 
(a junior objective synonym of crepitans Linnaeus, 1758, as published 
in the conbination Psophia crepitans). 


Section (2) of Annexe 


ALTERNATIVE “BB” 


Draft Ruling involving the use of the Plenary Powers to suppress ‘* Grus ”’ 
Brisson, 1760, to validate ‘‘ Grus ’’ Pallas, 1766, and to vary the 
type species of the genus so named 


DRAFT RULING :—(1) The following action is hereby taken 
under the Plenary Powers :— 


(a) The generic name Grus Brisson, 1760, is hereby suppressed for 
the purposes both of the Law of Priority and of the Law of 
Homonymy. 


DIRECTION 55 $33 


(b) All designations and selections of a type species for the genus 
Grus Pallas, 1766, are hereby set aside and the nominal species 
Ardea grus Linnaeus, 1758, is hereby designated to be the 
type species of the foregoing genus. 


(2) The entry relating to the generic name Grus Pallas, 1767, [sic], 
made on the Official List of Generic Names in Zoology by the Ruling 
given in Opinion 103 is hereby deleted and the following revised entry 
is hereby made in its place :—Grus Pallas, 1766 (gender : feminine) 
(type species, by designation under the Plenary Powers under (1)(b) 
above : Ardea grus Linnaeus, 1758). 


(3) The under-mentioned specific name is hereby placed on the 
Official List of Specific Names in Zoology :—grus Linnaeus, 1758, as 
published in the combination Ardea grus (specific name of type species 
of Grus Brisson, 1760). 


(4) The generic name Grus Brisson, 1760, as suppressed under the 
Plenary Powers under (1)(a) above is hereby placed on the Official 
Index of Rejected and Invalid Generic Names in Zoology. 


(5) The under-mentioned specific name is hereby placed on the 
Official Index of Rejected and Invalid Specific Names in Zoology :— 
psophia Pallas, 1767, as published in the combination Grus psophia 
(a junior objective synonym of crepitans Linnaeus, 1758, as published 
in the combination Psophia crepitans). 


II. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


7. Issue of Voting Paper V.P.(O.M.)(56)3 : On 25th April 1956 
a Voting Paper (V.P.(O.M.)(56)3) was issued in which the Members 
of the Commission were invited to vote either (1) * for “Alternative 
‘A*” as set out in the first part of the Annexe to the paper 
numbered Z.N.(S.) 558 relating to the generic name Grus sub- 
mitted by the Secretary simultaneously with the present Voting 
Paper [i.e. in the first part of the Annexe to the paper reproduced 
in paragraph 6 of the present Direction] (proposal that the 
normal provisions in the Régles be applied in this case); or 
(2) for “ Alternative ‘ B’”’ as set out in the second part of the 


334 OPINIONS AND. DECLARATIONS 


Annexe to the paper numbered Z.N.(S.) 558 relating to the 
generic name Grus referred to above (proposal that the Plenary 
Powers be used to suppress Grus Brisson, 1760, to validate Grus 
Pallas, 1766, and to vary the type species of the latter genus)”’. 


8. The Prescribed Voting Period : As the foregoing Voting 
Paper was issued under the One-Month Rule, the Prescribed 
Voting Period closed on 25th May, 1956. 


9. Particulars of the Voting on Voting Paper V.P.(O.M.)(56)3 : 
At the close of the Prescribed Voting Period, the state of the 
voting on Voting Paper V.P.(O.M.)(56)3 was as follows :— 


(a) Affirmative Votes in favour of the proposal styled *Altern- 
ative ‘A’ had been received from the following twenty- 
four (24) Commissioners (arranged in the order in which 
Votes were received): 

Holthuis ; Boschma; Stoll; Vokes; Hering; Miller ; 
Prantl ; Bodenheimer ; Cabrera; Esaki; Mayr; do 
Amaral ; Lemche ; Hanko ; Dymond ; Mertens; Key ; 


Jaczewski; Bonnet; Hemming; Sylvester-Bradley ; 
Riley ; Kuhnelt ; Tortonese ; 


(b) Affirmative Votes in favour of the proposal styled 
‘Alternative *‘ B’”’: 


None ; 


(c) On Leave of Absence, one (1) : 


Bradley (J.C.) ; 


(d) Voting Papers not returned : 


None. 


DIRECTION 55 309 


10. Declaration of Result of Vote: On 7th June 1956, 
Mr. Hemming, Secretary to the International Commission, 
acting as Returning Officer for the Vote taken on Voting Paper 
V.P.(O.M.)(56)3, signed a Certificate that the Votes cast were as 
set out in paragraph 9 above and declaring that the proposal 
styled “‘Alternative ‘A’’’ submitted with the foregoing Voting 
Paper had been duly adopted and that the decision so taken was 
the decision of the International Commission in the matter 
aforesaid. 


11. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On 25th September 1956, Mr. Hemming prepared the Ruling 
given in the present Direction and at the same time signed a Certi- 
ficate that the terms of that Ruling were in complete accord with 
those of the proposal approved by the International Commission 
in its Vote on Voting Paper V.P.(O.M.)(56)3. 


12. Original References: The following are the Original 
References for the names placed on Official Lists and Official 
Indexes by the Ruling given in the present Direction :— 


Grus Brisson, 1760, Ornithologia 5 : 374—391 

Grus Pallas, 1766, Miscell. Zool. : 65 

grus (Ardea), Linnaeus, 1758, Syst. Nat. (ed. 10) 1 : 141 
psophia (Grus), Pallas, 1767, Spicil. Zool. 4 : 1 


13. Family-Group-Name Problems: The family-group-name 
problem involved in the present case will be dealt with on a later 
occasion when comprehensive proposals will be submitted to the 
International Commission in regard to the problems arising in 
connection with this category of name involved in the Rulings 
given prior to the end of 1936 in Opinions in which the names 
of genera in the Class Aves were placed on the Official List of 
Generic Names in Zoology. 


14. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in dealing 
with the present case, and the present Direction is accordingly 


336 OPINIONS AND DECLARATIONS 


hereby rendered in the name of the said International Com- 
mission by the under-signed Francis Hemming, Secretary to the 
International Commission on Zoological Nomenclature, in 


virtue of all and every the powers conferred upon him in that 
behalf. 


15. The present Direction shall be known as Direction Fifty- 


Five (55) of the Jnternational Commission on Zoological 
Nomenclature. 


Done in London, this Twenty-Fifth day of September, Nineteen 
Hundred and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


oe sae tea rae ee ote ee earner a ieee eo 
Printed in England by MretcaLFe & COOPER LimitED, 10-24 Scrutton St., London BiG 2 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
“NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, c.M.G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.17. Pp. 337—364 


DIRECTION 56 


Completion and in certain cases correction of entries 

relating to the names of genera belonging to the Classes 

Pisces, Amphibia and Reptilia made in the Official List 

of Generic Names in Zoology in the period up to the end 
of 1936 


LONDON: 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956 


Price Eighteen Shillings and Sixpence 


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Issued 20th December, 1956 APR AO To y, 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 56 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JorDAN (British Museum (Natural History), 
Zoological Museum, Tring, Herts., England) 


President : Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A. 
(12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(arranged in order of precedence by reference to date of election or of most recent re-election 
as prescribed by the International Congress of Zoology) 


Professor H. BoscHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. aragen pai (Universitetets Zoologiske Museum, Copenhagen, Denmark) (27th 
uly 194 

Professor Teiso Esaki (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEy (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (5th July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt-Universitat zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMoND (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) (12th August 
1953) (President) 

Professor Harold E. Voxes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) (12th 
August 1953) 

Professor Béla HANKO (Mezégazdasdgi Muzeum, Budapest, Hungary) (12th August ae 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y., 
U.S.A.) (12th August 1953) / 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. HottHuts (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
ae August 1953) 

Dr. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
ee A.C.T., Australia) (1Sth October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum v Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KtiHNELT (Zoologisches Institut der Universitdt, Vienna, Austria) 
(6th November 1954) 

Professor F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 
1954) 

Professor Ernst Mayr (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico ToRTONESE (Museo di Storia Naturale ** G. Doria’’, Genova, Italy) 
(16th December 1954) 


DIRECTION 56 


COMPLETION AND IN CERTAIN CASES CORRECTION 
OF ENTRIES RELATING TO THE NAMES OF GENERA 
BELONGING TO THE CLASSES PISCES, AMPHIBIA 
AND REPTILIA MADE IN THE ‘* OFFICIAL LIST 
OF GENERIC NAMES IN ZOOLOGY ” IN THE 
PERIOD UP TO THE END OF 1936 


RULING :—(1) In the case of each of the under- 
mentioned names of genera belonging to the Class 
Pisces the incorrect statement that the type species was 
determined by subsequent selection under Rule (g) in 
Article 30 entered in the Official List of Generic Names in 
Zoology by the Ruling given in Opinion 77 is hereby 
deleted and the entry “type species by Linnean tau- 
tonymy ” is hereby inserted 1n its place :— 


(i) Acipenser Linnaeus, 1758 (Name No. 249) ; 
(ii) Chimaera Linnaeus, 1758 (Name No. 251) ; 
(ii1) Cyclopterus Linnaeus, 1758 (Name No. 255) ; 
(iv) Silurus Linnaeus, 1758 (Name No. 270). 


(2) In the case of each of the under-mentioned names of 
genera belonging to the Class Pisces the entry made in the 
Official List of Generic Names in Zoology by the Rulings 
given in the Opinions severally specified below 1s hereby 
completed by the insertion of a statement that the type 
species of the genus so named was determined by selection 
under Rule (g) in Article 30 by the author and on the date 
specified in each case below :— 


(a) Names placed on the “ Official List of Generic 
Names in Zoology” by the Ruling given in 
Opinion 1): 

(i) Callionymus Linnaeus, 1758: by selection 
by Bleeker, (1879): 


1 The expression ‘* Linnean tautonomy ”’ was introduced by the International 
Commission in Declaration 29 (1956, Ops. Decls. int. Comm. zool. Nomencl. 
14 : xxv—xxxii). 


JAN - 8 1957 


340 


OPINIONS AND DECLARATIONS 


(11) Clupea Linnaeus, 1758: by selection by 
Gill (1862) ; 


(111) Coryphaena Linnaeus, 1758: by selection 
by Jordan & Gilbert (1882) ; 


(iv) Cyprinus Linnaeus, 1758: by selection by 
Jordan & Gilbert (1882) ; 


(v) Gadus Linnaeus, 1758: by selection by 
Jordan & Gilbert (1882) ; 


(vi) Gasterosteus Linnaeus, 1758: by selection 
by Jordan & Gilbert (1882) ; 


(vil) Gobius Linnaeus, 1758: by selection by 
Gill (1863) ; 


(vii) Lophius Linnaeus, 1758: by selection by 
Jordan & Gilbert (1882) ; 


(ix) Mormyrus Linnaeus, 1758 : by selection by 
Jordan & Evermann (1917) ; 


(x) Mullus Linnaeus, 1758: by selection by 
Bleeker (1876) ; 


(xi) Perca Linnaeus, 1758: by selection by 
Gill (1862) ; 


(xl) Salmo Linnaeus, 1758: by selection by 
Jordan & Gilbert (1882) ; 


(xiii) Scomber Linnaeus, 1758: by selection by 
Gill (1863) ; 


(xiv) Scorpaena Linnaeus, 1758: by selection by 
Bleeker (1876) ; 


(xv) Syngnathus Linnaeus, 1758 : by selection by 
Jordan (1912) ; 


(xvi) Zeus Linnaeus, 1758: by selection by Gill 
(1863) ; 


DIRECTION 56 341 


(b) Names placed on the “ Official List of Generic 
Names in Zoology” by the Ruling given in 
* Opinion” 92:: 
(i) Blennius Linnaeus, 1758: by selection by 
Jordan & Gilbert (1882) ; 


(ii) Esox Linnaeus, 1758: by -selection by 
Jordan & Gilbert (1882) ; 


(ii1) Ophidion Linnaeus, 1758: by selection by 
Gill (1864). 


(3) In the case of each of the under-mentioned names of 
genera belonging to the Class Reptilia the entry made on 
the Official List of Generic Names in Zoology by the Ruling 
given in Opinion 92 is hereby completed by the insertion 
of a statement that the type species of the genus so named 
was determined by selection under Rule (g) in Article 30 
by the author and on the date severally specified below :— 


(a) Crotalus Linnaeus, 1758: by selection by Fitzinger 
(1843) ; 


(b) Lacerta Linnaeus, 1758: by selection by Fitzinger 
(1843). 


(4) The date “ 1849” attributed to the generic name 
Desmognathus Baird (Class Amphibia) when that name 
was placed on the Official List of Generic Names in Zoology 
by the Ruling given in Opinion 92 is hereby deleted and 
the date “(1850)” is hereby inserted in its place. 


(5) The entry on the Official List of Generic Names in 
Zoology regarding the generic name Desmognathus Baird 
referred to in (4) is hereby completed by the insertion of a 
statement that the type species of the genus so named was 
determined by selection under Rule (g) in Article 30 by 
Steyneger & Barbour in 1917. 


(6) The entry on the Official List of Generic Names in 
Zoology of a generic name spelled Coryhaena and 


342 OPINIONS AND DECLARATIONS 


attributed to Linnaeus, 1758, inadvertently made by 
the Ruling given in Opinion 77 is hereby corrected to 
Coryphaena Linnaeus, 1758. 


(7) The incorrect entry of Eleotris gyrinus Cuvier & 
Valenciennes, 1837, as the name of the type species of 
the genus Eleotris Schneider, 1801, made on the Official 
List of Generic Names in Zoology by the Ruling given in 
Opinion 93 is hereby deleted and the name Gobius pisonis 
Gmelin (J.F.), [1789], is hereby inserted in its place. 


(8) The under-mentioned correction is hereby made in 
the entry relating to the generic name Calamaria Boie 
(Class Reptilia) made on the Official List of Generic 
Names in Zoology by the Ruling given in Opinion 92: 
The entry “ 1827” as the date of publication of the fore- 
going generic name is hereby deleted and the entry 
‘* 1826 ”’ is hereby inserted in its place. 


(9) The entry on the Official List of Generic Names in 
Zoology relating to the generic name Dermochelys 
Blainville, 1816 (Class Reptilia), made by the Ruling 
given in Opinion 92 is hereby completed by the insertion 
of the words “* by monotypy, through Cuvier (1829) ” as 
indicating the manner by which the type species of this 
genus was determined under Article 30 of the Régles. 


(10) The under-mentioned generic names are hereby 
placed on the Official Index of Rejected and Invalid 
Generic Names in Zoology with the Name Numbers 
severally specified below :— 


| (i) Acipenses Linck, 1790 (an Erroneous Subse- 
quent Spelling for Acipenser Linnaeus, 1758) 
(Name No. 765) ; 


(ii) Ammodytes Bonaparte, 1831 (ajunior homonym 
of Ammodytes Linnaeus, 1758) (Name No. 
766) ; | 


DIRECTION 56 343 


(iii) Anarhicas Latreille, 1804 (an Erroneous Subse- 
quent Spelling for Anarhichas Linnaeus, 1758 
(Name No. 767) ; 


(iv) Calliongmus Linck, 1790 (an Erroneous Subse- 
quent Spelling for Callionymus Linnaeus, 
1758) (Name No. 768) ; 


(v) Callionimus Gouan, 1770 (an Erroneous Sub- 
sequent Spelling for Ca/lionymus Linnaeus, 
1758) (Name No. 769) ; 


(vi) Chimaera Poli, 1791 (a junior homonym of 
Chimaera Linnaeus, 1758) (Name No. 770) ; 


(vii) Chimaera Ochsenheimer, 1808 (a junior homo- 
~ nym of Chimaera Linnaeus, 1758) (Name 
Noz771); 


(vill) Chimaera Hitchcock, 1858 (a junior homonym 
of Chimaera Linnaeus, 1758) (Name No. 
Rid): 


(ix) Chimaira Duméril, 1856 (an Erroneous Sub- 
sequent Spelling for Chimaera Linnaeus, 
1758) (Name No. 773) ; 


(x) Coregonus Lacépéde, 1803 (a junior homonym 
of Coregonus Linnaeus, 1758) (Name No. 
7TA) ; 


(xi) Coregonus Jarocki, 1822 (a junior homonym 
of Coregonus Linnaeus, 1758) (Name No. 
ee ie 


(xi) Coriudo Fleming, 1822 (a junior objective 
synonym of Dermochelys Blainville, 1816) 
(Name No. 776) ; 


(xi1) Coryhaena Stiles, 1922 (an Erroneous Sub- 
sequent Spelling for Coryphaena Linnaeus, 
1758) (Name No. 777) ; 


(xiv) Coryphaena Coues, 1889 (a nomen nudum) 
(Name No. 778) ; 


OPINIONS AND DECLARATIONS 


(xv) Cyclopterus Marseul, 1872 (a junior homonym 
of riaeaues Linnaeus, 1758) (Name No. 
ALO) 


(xvi) Dermatochelys Wagler, 1830 (a junior objective 
synonym of Dermochelys Blainville, 1816) 
(Name No. 780) ; 


(xvi) Dermochelis Lesueur, 1829 (an Erroneous Sub- 
sequent Spelling for Dermochelys Blainville, 
1816) (Name No. 781) ; 


(xvill) Diodon O’Reilly, 1818 (a junior homonym of 
Diodon Linnaeus, 1758) (Name No. 782) ; 


(xix) Diodon Lesson, 1828 (a junior homonym of 
Diodon Linnaeus, 1758) (Name No. 783) ; 


(xx) Diodon Lesson, 1830 (a junior homonym of 
Diodon Linnaeus, 1758) (Name No. 784) ; 


(xx1) Draco Gouan, 1770 (a junior homonym of 
Draco Linnaeus, 1758) (Name No. 785) ; 


(xxii) Draco Oken, 1816 (a junior homonym of Draco 
Linnaeus, 1758) (Name No. 786) ; 


(xxi) Eleotris Gronovius, 1763 (a name published 
in a work rejected for nomenclatorial pur- 
poses by the Ruling given in Opinion 261) 
(Name No. 787) ; 


(xxiv) Fistularia Forskal, 1775 (a junior homonym of 
Fistularia Linnaeus, 1758) (Name No. 788) ; 


(xxv) Fistularia Miller (O.F.), 1776 (a junior homo- 
nym of Fistularia Linnaeus, 1758) (Name 
No: 789): 


(xxvi) Fistularia Lamarck, 1816 (a junior homonym 
of Fistularia Linnaeus, 1758) (Name No. 
790) ; 


(xxvil) Fistularia Blainville, 1830 (a junior homonym 
of Fistularia Linnaeus, 1758) (Name No. 
Aol): 


DIRECTION 56 345 


(xxvill) Fistularia Bowerbank, 1841 (a junior homonym 
of Fistularia Linnaeus, 1758) (Name No. 
792):3 


(xxix) Gadus Dejean, 1821 (a junior homonym of 
Gadus Linnaeus, 1758) (Name No. 793) ; 


(xxx) Gadus Rang, 1829 (a junior homonym of 
Gadus Linnaeus, 1758) (Name No. 794) ; 


(xxxi) Gadus Deshayes, 1861 (a junior homonym of 
Gadus Linnaeus, 1758) (Name No. 795) ; 


(xxxil) Gymnothorax Cuvier, [1800] (a nomen nudum) 
(Name No. 796) ; 


(xxxiii) Gymnothorax Duda, 1923 (a junior homonym 
of Gymnothorax Bloch, 1795) (Name No. 


by) 


(xxx1v) Mabouia Cuvier, 1829 (an Erroneous Subse- 
quent Spelling for Mabuya (Rafinesque) 
Fitzinger, 1826) (Name No. 798) ; 


(xxxv) Mabouya Duméril & Bibron, 1839 (an 
Erroneous Subsequent Spelling for Mabuya 
(Rafinesque) Fitzinger, 1826) (Name No. 
799) ; 


(xxxv1) Mabuia Cuvier, 1829 (an Erroneous Subse- 
quent Spelling for Mabuya (Rafinesque) 
Fitzinger, 1826) (Name No. 809) ; 


(xxxvil) Mabuya Rafinesque, 1815 (a nomen nudum) 
(Name No. 801) ; 


(xxxvill) Malapterus Jarocki, 1822 (an Erroneous Sub- 
sequent Spelling for Malapterurus Lacépéde, 
1803) (Name No. 802) ; 


(xxxix) Malapturus Swainson, 1838 (an Erroneous 
Subsequent Spelling for Malapterurus 
Lacépede, 1803) (Name No. 803) ; 


(xl) Mugie Macklot, 1830 (an Erroneous Subse- 
quent Spelling for Mugil Linnaeus, 1758) 
(Name No. 804) ; 


346 OPINIONS AND DECLARATIONS 


(xli) Mustellus Fischer de Waldheim, 1813 (an 
Erroneous Subsequent Spelling for Mustelus 
Linck, 1790) (Name No. 805) ; 


(xlii) Ophidion Pomel, 1853 (a junior homonym of 
Ophidion Linnaeus, 1758) (Name No. 806) ; 


(xliil) Ophidium Linnaeus, 1766 (an Invalid Emenda- 
tion of Ophidion Linnaeus, 1758) (Name 
No. 807) ; 


(xliv) Siren Hermann, 1783 (a junior homonym of 
Siren Linnaeus, 1766) (Name No. 808) ; 


(xlv) Sirena Fischer de Waldheim, 1808 (an Invalid 
Emendation of Siren Linnaeus, 1766) (Name 
No. 809) ; 


(xlvi) Sirene Oken, 1816 (an Invalid Emendation of 
Siren Linnaeus, 1766; published in a work 
rejected for nomenclatorial purposes by the 
Ruling given in Opinion 417 (Name No. 810) ; 


(xlvii) Sphargis Merrem, 1820 (a junior objective 
synonym of Dermochelys Blainville, 1816) 
(Name No. 811) ; 


(xlvili) Syngnathus Rafinesque, 1810 (a junior 
homonym of Syngnathus Linnaeus, 1758) 
(Name No. 812) ; 


(xlix) Teuthis Schneider, 1784 (a junior homonym 
of Teuthis Linnaeus, 1766) (Name No 813) ; 


(11) The under-mentioned specific names are hereby 
placed on the Official Index of Rejected and Invalid 
Specific Names in Zoology with the Name Numbers 
severally specified below :— 


(i) mercurialis Merrem, 1820, as published in the 
combination Sphargis mercurialis (a junior 
objective synonym of coriacea Linnaeus, 
1766, as published in the combination 
Testudo coriacea) (Name No. 365) ; 


DIRECTION 56 347 


(11) alleghaniensis Harlan, 1825, as published in 
the combination Abranchus  alleghaniensis 
(an Invalid Emendation of alleganiensis 
Daudin, [1803], as published in the 
combination Salamandra alleganiensis) 
(Name No. 366) ; 


I. THE SUBJECT MATTER OF THE PRESENT 
DIRECTION” 


The present Direction embodies decisions taken by the Inter- 
national Commission on Zoological Nomenclature for the 
purpose of completing, and, where necessary, of correcting, the 
entries in regard to the names of genera belonging to the Classes 
Pisces, Amphibia and Reptilia placed on the Official List of 
Generic Names in Zoology in the period up to the end of 1936, 
this action being a necessary preliminary to the publication of the 
Official List in book-form. The proposals upon which the 
decisions taken in the present case were based were submitted 
to the International Commission by the Secretary on 25th April 
1956 in the following paper :— 


Proposed completion and correction of entries relating to the names of 
genera belonging to the Classes Pisces, Amphibia and Reptilia 
made in the ‘* Official List of Generic Names in Zoology ”’ 
in the period up to the end of 1936 


By FRANCIS HEMMING, C.M.G., C.B.E. 
(Secretary to the International Commission on Zoological Nomenclature) 


The present paper contains particulars of various minor errors 
and omissions in the entries relating to the names of genera of the 
Classes Pisces, Amphibia and Reptilia which were made on the Official 
List of Generic Names in Zoology in the period up to the end of 1936, 


348° OPINIONS AND DECLARATIONS 


Proposals are submitted for rectifying these defects. The action called 
for, which falls into various groups, is described in the following 
paragraphs. 


(1) Correction of erroneous entries on the ‘* Official List ”’ 
as to the Rule in Article 30 of the ‘‘ Régles ’’ under which 
the type species of four genera belonging to the Class 
Pisces were determined 


2. In Opinion 77 the names of four genera (Acipenser ; Chimaera ; 
Cyclopterus ;  Silurus) published by Linnaeus in 1758 and belonging 
to the Class Pisces, were placed on the Official List of Generic Names 
in Zoology with the intimation, through the use of the symbol “‘ tsd ”’ 
[= type by subsequent designation], that in each case the type species 
had been determined under Rule (g) (type species by subsequent 
selection) in Article 30. When making the foregoing entries, the 
Commission must have inadvertently failed to recall that the Rules in 
Article 30 operate successively and that for any given name Rule (g) 
becomes applicable only after it has been established that none of the 
earlier Rules in that Article apply. In the case of the generic names 
cited above no type species was designated by the original author 
under Rule (a) ; nor was a type species indicated under either Rule (b) 
(indication of type species through the use for an included species of 
the specific name “‘ typus’’ or “‘ typicus’’? as a new name) or under 
Rule (c) (indication of type species by monotypy). The position is, 
however, quite different when we come to Rule (d) (indication of type 
species by absolute tautonymy). In none of the cases under considera- 
tion was there included a species having a specific name consisting 
of the same word as that of which the generic name concerned was 
composed. But in each case a tautonymous pre-1757 universal species- 
name was cited in the synonymy of one, and only one, of the included 
species. Under the Ruling given in Opinion 16, later incorporated into 
the Régles by the Thirteenth International Congress of Zoology, 
Paris, 1948 (1950, Bull. zool. Nomencl. 4 : 155) the inclusion in a new 
genus established prior to Ist January 1931 of a nominal species in 
the synonymy of which such a pre-1757 species-name is cited, con- 
stitutes the indication of that species under Rule (d) in Article 30 
as the type species of the genus concerned. It may be noted as some- 
what curious that in Opinion 16 the Commission had actually cited 
each of the names with which we are concerned in a list of the names of 
genera, the type species of which were determinable under the inter- 
pretation of Rule (d) in Article 30 then propounded. 


3. In order to rectify the position disclosed above it is proposed 
that the incorrect notation “‘type by subsequent designation ’”’ be 
expunged from the entry on the Official List relating to each of the 
four generic names discussed above and that in its place there be 


DIRECTION 56 349 


inserted a note that the type species of the genus concerned was deter- 
mined under Rule (d) in Article 30 as interpreted by Opinion 16, 
i.e. under what has come to be known as Linnean tautonymy.? 


(2) Proposed insertion in the ‘* Official List ’’ of particulars as 
to how the type species of certain genera were determined 
under Article 30 of the ‘‘ Régles ”’ 


4. In Opinion 77 the names of the following sixteen genera belonging 
to the Class Pisces were placed on the Official List of Generic Names 
in Zoology with no information as to how the species there specified 
had become the type species of the genera concerned, apart from the 
entirely inadequate symbol ‘“‘tsd’’ [= type by subsequent 
designation] :— 


Callionymus; Clupea;  Coryphaena; Cyprinus; Gadus ; 
Gasterosteus ; Gobius; Lophius; Mormyrus; Mullus ; Perca ; 
Salmo ; Scomber ; Scorpaena ; Syngnathus ; Zeus. 


5. In Opinion 92 the names of six genera of the groups with which 
we are here concerned were placed on the Official List with no 
indication whatever as to how the type species of the genera concerned 
had been determined under Article 30. The names in question were 
the following :— 


Class Pisces : Blennius ; Esox ; Ophidion ; 
Class Reptilia: Crotalus ; Lacerta ; 
Class Amphibia : Desmognathus. 


6. For each of the sixteen names specified in paragraph 4 above 
and of the six names specified in paragraph 5 above, it has been 
necessary to undertake a search of the literature (i) to ascertain by 
whom, when and where one of the originally included species was 
selected to be the type species of the nominal genus concerned, (ii) 
to check whether that species is the species adopted as such in the 
Official List, (111) to determine whether the species concerned is 
currently accepted by specialists as the type species, and (iv) to establish 
whether the specific name of the nominal species which is the type 
species is an available name and is moreover accepted by specialists 
as the oldest such name for the species in question. The first of these 
tasks proved unusually laborious, so far as the names of fishes were 
concerned, owing to the paucity of reliable catalogues dealing with this 
group. However, the search of the literature of all three groups 
concerned ultimately established (a) that the species adopted in the 


“ See Footnote 1. 


350 OPINIONS AND DECLARATIONS 


Official List as the type species of the various genera are currently 
accepted as such (b) that each of these species has been selected as the 
type species in accordance with the requirements of Rule (g) in Article 
30 and (c) that there is no evidence to suggest as regards any of the 
nominal genera concerned that any other included species was selected 
as the type species prior to the selection of the species currently 
accepted as such. In some cases the earliest selection which it has been 
possible to trace is relatively recent but it is believed that nevertheless 
the reference now brought forward is the earliest in which a type 
selection was made under Rule (g) of Article 30, rigorously construed as 
required by the Régles, for the genera concerned. The references for 
the selection of type species for the twenty-two genera concerned which 
it is recommended should be inserted in the Official List to make good 
the omissions in Opinions 77 and 92 are set out in Annexe | to the 
present paper. 


(3) Proposed correction or amplification of entries in regard 
to six generic names made in the ‘* Official List ’’ by Rulings 
given in ‘* Opinions ’’ 77, 92 and 93 


7. ** Coryphaena ’’ Linnaeus, 1758 (Class Pisces) : When the generic 
name Coryphaena Linnaeus, 1758, was placed on the Official List by 
the Ruling given in Opinion 77, it was misspelled as ‘‘ Coryhaena’’. This 
error which was presumably due to faulty proof-reading must now be 
corrected, the Erroneous Subsequent Spelling Coryhaena being at the 
same time placed on the Official Index of Rejected and Invalid Generic 
Names in Zoology. 


8. ‘* Eleotris ’? Schneider, 1801 (Class Pisces) : In Opinion 93 the 
International Commission used its Plenary Powers to designate 
Gobius pisonis Gmelin (J.F.), [1789], to be the type species of Eleotris 
Schneider, 1801. However, it was stated in the so-called ‘“* Summary ”’ 
of this Opinion that the species so designated as type species was 
‘* gyrinus Cuv. & Val.”, i.e. Eleotris gyrinus Cuvier & Valenciennes, 
1837 (Hist. nat. Poiss. 12 : 220), a nominal species which had been 
mentioned by the applicant in this case as a junior subjective synonym of 
Gobius pisonis Gmelin. The foregoing entry can only have been made 
through an inexplicable oversight and must be corrected by the 
substitution of a revised entry prescribing that, in accordance with the 
decision taken by the Commission in this Opinion the type species of 
this genus is Gobius pisonis Gmelin, [1789]. 


9, ** Calamaria ’’ Boie, 1826 (Class Reptilia) : In Opinion 92 the 
generic name Calamaria Boie was placed on the Official List ; it was 
there attributed to the year “‘ 1827 ”, and no reference was given beyond 
the page number “‘ 236’. The type species was said to be “‘ Coluber | 
calamaria Linnaeus, 1758”, by absolute tautonymy. The name 


DIRECTION 56 351 


Calamaria was published by Boie both in 1826 and 1827. In the 
earlier year, it was published in a paper by Férussac (Bull. Sci. nat. 
Géol. 9 : 236) and it is clear from the page number cited in Opinion 92 
that this is the paper there referred to and not the later paper of 1827 
(Isis (Oken) 10 : 519). The name Calamaria Boie, as published in 
Férussac’s paper of 1826 was treated by Neave (Nomencl. zool.) as a 
nomen nudum, but this is incorrect, for, although Boie included six 
nomina nuda in the genus, he included also one new nominal species 
which was provided with an indication as required by Article 25 of 
the Régles. This nominal species was introduced as follows: 
*“Calamaria Boie . . . Linnaei B. (Col. calamaria Linn.)”’. This 
species is therefore the type species by monotypy. Under Declaration 
21 (embodying the decision taken by the Commission in its vote on 
Voting Paper V.P.(55)30*) the type species of this genus is to be cited 
under the earlier of the two objective synonyms applied to it by the 
author of the generic name. That is, the type species is to be cited as 
Coluber calamarius Linnaeus, 1758, and not as Calamaria linnaei Bote, 
1826. In order to regularise the position, it is necessary now that the 
date attributed to Calamaria Boie should be corrected from ‘“‘ 1827” 
to “* 1826’, and that the erroneous entry “‘tat’’ [type by absolute 
tautonymy] should be corrected to “‘type by monotypy”’. At the 
same time the objectively invalid specific name linnaei Boie, 1826, 
should be placed on the Official Index.’ | 


10. ‘*‘ Dermochelys ’’ Blainville, 1816 (Class Reptilia) : The generic 
name Dermochelys Blainville, 1816, was placed on the Official List 
with Testudo coriacea Linnaeus, 1766, as type species. So far as it 
goes, the entry so made was correct. It contained no particulars, 
however, as to how that species became the type species and a word 
of explanation is needed on this subject. This nominal genus, as 
established by Blainville in 1816 (J. Phys. Chim. Hist. nat., Paris 
83 : 259), contained no nominal species. In order to determine 
what species should be regarded as being originally included species 
in this genus, it is necessary to ascertain by whom nominal species 
were first assigned to it and what that species or those species were 
(1950, Bull. zool. Nomencl. 4 : 159—160, 346). Before doing this, we 
should, however, take note of two events which occurred before any 
author dealt again with the nominal genus Dermochelys. ‘The first 
of these events was the establishment by Merrem in 1820 (Tent. Syst. 
Amph. : 19) of the genus Sphargis. Merrem only recognised one species 
as belonging to this genus and to this he gave the name Sphargis 
mercurialis (: 19), the name mercurialis being a name taken over by 


3 Declaration 21 was published on 19th June 1956 (Ops. Decls. int. Comm. zool. 
Nomencl. 12 : i—viii.) 


4 As explained in paragraph 4 of the present Direction, unexpected developments 
in regard to the generic name Ca/amaria Boie led to the temporary withdrawal 
of certain of the proposals submitted in regard to it, in order to permit of the 
study of the new problems disclosed. 


352 OPINIONS AND DECLARATIONS 


him from the XVIth century author Rondeletius. In a footnote 
Merrem stated that his Sphargis mercurialis was the same species as the 
Testudo coriacea of Linnaeus. The latter species is therefore under 
Declaration 21 the type species of Sphargis Merrem by monotypy. 
The next event was the establishment by Fleming in 1822 (Phil. Zool. 
2 : 271) of the nominal genus Coriudo. This was introduced in a key 
to the genera of ‘‘ CHELONEA ”’ and the sole species cited, and therefore 
the type species by monotypy, was Testudo coriacea Linnaeus. The 
name Coriudo Fleming is thus a junior objective synonym of Sphargis 
Merrem. In 1829 (Régne Anim. (ed. 2) 2 : 14) Cuvier used the name 
Dermochelys which, however he spelled Dermochelis and attributed, 
to Lesueur (by whom it had presumably been proposed in manuscript 
and from whom Blainville may also be assumed to have obtained the 
name). Cuvier, after sinking the names Sphargis Merrem and Coriudo 
Fleming as synonyms of Dermochelis, cited Testudo coriacea Linnaeus 
as belonging to this genus. In a footnote he cited a second species 
as belonging to this genus, namely Dermochelis atlantica Lesueur. No 
such name had ever been published by that author and the name as 
cited by Cuvier was a nomen nudum. Accordingly the only species 
placed by Cuvier in the genus Dermochelys was Testudo coriacea 
Linnaeus which therefore becomes the type species of Blainville’s 
genus by monotypy. In order to avoid the risk of misunderstanding, 
it is desirable that, when the words * by monotypy ”’ are entered in the 
Official List to indicate how Testudo coriacea Linnaeus, 1766, became 
the type species of Dermochelys Blainville, 1816, they should be 
qualified by the words “‘ through Cuvier, 1829’. At the same time the 
various objectively invalid names discussed in the present paragraph 
should be placed on the appropriate Official Index. | 


11. ‘‘ Desmognathus ’’ Baird, (1850) (Class Amphibia) : When the 
name Desmognathus Baird was placed on the Official List, by the 
Ruling given in Opinion 92, the date “‘ 1849 ”’ was attributed to it. The 
date which appears on the title page of Volume 1 of the Second Series 
of the J. Acad. nat. Sci. Philad., in which Baird’s paper was published, 
is “‘ 1847—1850”’ but particulars of the dates of publication of the 
various parts are given in the table of contents and this shows that that 
paper, which appeared in Part 4, was published in January 1850. This 
date—cited in round brackets (parentheses)—should therefore be 
attributed to the name Desmognathus Baird. 


12. ‘* Cryptobranchus ’’ Leuckart, 1821 (Class Amphibia): As 
explained in Paper Z.N.(S.) 1014 now being submitted with Voting 
Paper V.P.(O.M)(56)5° ; the name of the type species of the genus 


5 The decision taken by the International Commission on the Voting Paper 
here referred to has since been embodied in Direction 57 (which is being 
published in the immediately following Part of the present Section of Volume 1 
of the present Series). 


DIRECTION 56 353 


Cryptobranchus Leuckart, 1821, is currently regarded as a junior 
subjective synonym of the name Sa/lamandra alleganiensis Daudin, 
[1803]. It must be noted here that the alleganiensis Daudin, 
[1803] was emended to alleghaniensis by Harlan in 1825 (Ann. 
Lyceum nat. Hist. New York 1(8) : 271), who cited this species as 
Abranchus alleghaniensis. In this action Harlan was followed by some 
later authors but the original spelling employed by Daudin is that 
currently in use and is the correct name for this nominal species. The 
Invalid Emendation alleghaniensis Harlan, 1825, should now be placed 
on the Official Index. 


(4) Proposed addition of certain names to the ‘* Official Indexes 
of Rejected and Invalid Names in Zoology ”’ 


13. The examination of the literature undertaken in preparation for 
the submission of the proposals set forth in the present paper has 
brought to light the existence of a number of names which are 
objectively invalid as junior homonyms, as Invalid Emendations or 
Erroneous Subsequent Spellings, or as junior objective synonyms. 
These names, which should now be placed on the appropriate Official 
Indexes, are set out in Annexe 2 to the present paper, the generic 
names being listed in Section A and the specific names in Section B. 


(5) Recommendations 


14. In the light of the considerations set out in the present paper 
it is recommended that, in discharge of the duty of completing and, 
where necessary, correcting the entries made on the Official List 
of Generic Names in Zoology in the period up to the end of 1936 
imposed upon it by the Thirteenth International Congress of Zoology, 
Paris, 1948, the International Commission on Zoological Nomen- 
clature should take the following action, so far as concerns the entries 
relating to the names of genera of the Classes Pisces, Amphibia and 
Reptilia made on the Official List during the period referred to above, 
namely that it should :— 


_ (1) in the case of each of the four generic names specified in para- 
graph 2 above, substitute in the Official List of Generic Names 
in Zoology the words “‘ type species by Linnean tautonymy ”’ for 
the words “‘ t[ype by] s[ubsequent] d[esignation] ”’ as indicating 
the manner in which the type species of the genera concerned 
were determined under Article 30 of the Régles ; 


354 OPINIONS AND DECLARATIONS 


(2) in the case of each of the generic names specified in Annexe 1, 
insert in the foregoing Official List the particulars specified 
in that Annexe regarding the selection of the type species of 
the genera concerned under Rule (g) in Article 30 of the 
Régles (paragraphs 4—6 above) ; 


(3) correct from Coryhaena to Coryphaena the entry on the Official 
List in regard to the generic name Coryphaena Linnaeus, 1758, 
inadvertently made by the Ruling given in Opinion 77 
(paragraph 7 above) ; 


(4) insert on the Official List the name of the nominal species Gobius 
pisonis Gmelin (J.F.), [1789], as the name of the type species 
of Eleotris Schneider, 1801, in place of the name Eleotris 
gyrinus Cuvier & Valenciennes, 1837, inadvertently entered as 
such by the Ruling given in Opinion 93 (paragraph 8 above) ; 


(5) substitute on the Official List the date “‘ 1826”’ for the date 
** 1827” as the date of publication of the generic name 
Calamaria Boie (paragraph 9 above) ; 


(6) substitute on the Official List the words ‘type species, by 
monotypy ”’ for the words “‘ t[ype by] a[bsolute] tjautonymy] ”’ 
as indicating the manner in which the type species of 
Calamaria Boie, 1826, was determined under Article 30 of the 
Régles (paragraph 9)§ ; 


(7) take note that under Declaration 21 the type species of Calamaria 
Boie, 1826, is the nominal species Coluber calamarius Linnaeus, 
1758, and not the later-established objectively identical nominal 
species Calamaria linnaei Boie, 1826, cited at the time of the 
establishment of the above nominal genus (paragraph 9)°; 


(8) insert the words “ by monotypy, through Cuvier, 1829” in the 
Official List as recording the means by which the type species 
of the genus Dermochelys Blainville, 1816, was determined 
under Article 30 (paragraph 10) ; 


(9) substitute on the Official List the date ‘‘ (1850) ”’ for the date 
“1849”? as the date of publication of the generic name 
Desmognathus Baird (paragraph 11) ; 


(10) place on the Official Index of Rejected and Invalid Generic 
Names in Zoology the objectively invalid names enumerated 
in Section A of Annexe 2 to the present paper (paragraph 13 
above). 


(11) place on the Official Index of Rejected and Invalid Specific Names 
in Zoology the objectively invalid names enumerated in Section 
B of Annexe 2 to the present paper (paragraph 13 above). 


6 This proposal was later withdrawn. See paragraph 4 of the preseut Direction, 


DIRECTION 56 355 


ANNEXE | 


Proposals for completing the entries on the ‘‘ Official List of Generic 
Names in Zoology ’’ regarding the manner in which the type 
species of certain genera were determined under Article 30 
of the ‘‘ Regles ”’ 


Name of Genus 


(1) 


Callionymus Linnaeus, 1758 
Clupea Linnaeus, 1758 
Coryphaena Linnaeus, 1758 
Cyprinus Linnaeus, 1758 
Gadus Linnaeus, 1758 
Gasterosteus Linnaeus, 1758 
Gobius Linnaeus, 1758 
Lophius Linnaeus, 1758 
Mormyrus Linnaeus, 1758 
Mullus Linnaeus, 1758 
Perca Linnaeus, 1758 
Salmo Linnaeus, 1758 
Scomber Linnaeus, 1758 
Scorpaena Linnaeus, 1758 


Syngnathus Linnaeus, 1758 


Entry relating to the determination 
of the type species proposed to be 
inserted in the “‘ Official List of 
Generic Names in Zoology ” 


(2) 


by selection by Bleeker, 1879, Vers. 
Acad. Amsterdam 1879 : 80 


by selection by Gill, 1862, Proc. 
Acad. nat. Sci. Philad. 1861 : 35 


by selection by Jordan & Gilbert, 
1882, Bull. U.S. nat. Mus. 16 : 454 


by selection by Jordan & Gilbert, 
1882, Bull. U.S. nat. Mus. 16 : 254 


by selection by Jordan & Gilbert, 
1882, Bull. U.S. nat. Mus. 16 : 802 


by selection by Jordan & Gilbert, 
1882, Bull. U.S. nat. Mus. 16 : 393 


by selection by Gill, 1864, Proc. 
Acad. nat. Sci. Philad. 1863 : 268 


by selection by Jordan & Gilbert, 
1882, Bull. U.S. nat. Mus. 16 : 844 


by selection by Jordan & Evermann, 
1917, Gen. Fish. : 15 


by selection by Bleeker, 1876, Arch. 
Neerl. Sci. nat. Haarlem 11 : 334 


by selection by Gill, 1862, Proc. 
Acad. nat. Sci. Philad. 1861 : 48 


by selection by Jordan & Gilbert, 
1882, Bull. U.S. nat. Mus. 16 : 309 


by selection by Gill, 1863, Proc. 
Acad. nat. Sci. Philad. 1862 : 125 


by selection by Bleeker, 1876, Versi. 
Acad. Amsterdam 1875 : 295 


by selection by Jordan, 1912, Smiths. 
misc. Publ. 2060 : 103 


356 OPINIONS AND DECLARATIONS 


Name of Genus 


(1) 


Zeus Linnaeus, 1758 
Blennius Linnaeus, 1758 
Esox Linnaeus, 1758 
Ophidion Linnaeus, 1758 
Crotalus Linnaeus, 1758 
Lacerta Linnaeus, 1758 


Desmognathus Baird, (1850) 


Entry relating to the determination 
of the type species proposed to be 
inserted in the *‘ Official List of 
Generic Names in Zoology” 


(2) 
by selection by Gill, 1863, Proc. 
Acad. nat. Sci. Philad. 1862 : 126 
by selection by Jordan & Gilbert, 
1882, Bull. U.S. nat. Mus. 16 : 759 
by selection by Jordan & Gilbert, 
1882, Bull. U.S. nat. Mus. 16 : 352 
by selection by Gill, 1864, Proc. 
Acad. nat. Sci. Philad. 1863 : 210 
by selection by Fitzinger, 1843, 
Syst. Rept. : 29 
by selection by Fitzinger, 1843, Syst. 
Rept. : 20 
by selection by Stejneger & Barbour 


1917, Checklist N. Amer. Amph & 
Rept. : 22 


ANNEXE 2 


Proposed addition of certain invalid names to the ‘* Official Indexes 
of Rejected and Invalid Names in Zoology ”’ 


Section A: Proposed addition of forty-nine generic names 
to the ‘‘ Official Index of Rejected and Invalid Generic 
Names in Zoology ”’ 


1. Acipenses Linck, 1790, Mag. N. Phys. Naturg. 6(3) : 37 (an Erro- 
neous Subsequent Spelling for Acipenser Linnaeus, 1758, 
placed on the Official List in Opinion 77) 


2. Ammodytes Bonaparte, 1831, Giorn. Arcad. 52 : 149 (a junior 
homonym of Ammodytes Linnaeus, 1758, placed on the Official 


List in Opinion 75) 


3. Anarhicas Latreille, 1804, Nouv. Dict. Hist. nat. Tabl. méth. : 103 
(an Erroneous Subsequent Spelling for Anarhichas Linnaeus, 
1758, placed on the Official List in Opinion 75) 


4. Calliongmus Linck, 1790, Mag. N. Phys. Naturg. 6(3) : 34 (an 
Erroneous Subsequent Spelling for Callionymus Linnaeus, 1758, 
placed on the Official List in Opinion 77) 


10. 


LF. 
12. 


13. 


14. 


15. 


DIRECTION 56 357 


. Callionimus Gouan, 1770, Hist. Pisc. : 121 (an Erroneous Subse- 


quent Spelling for Callionymus Linnaeus, 1758, placed on the 
Official List in Opinion 77) 


. The under-mentioned junior homonyms of Chimaera Linnaeus, 


1758, placed on the Official List in Opinion 77 :— 


(a) Chimaera Poli, 1791, Test. Sicil. 1, Introd. : 31 
(b) Chimaera Ochsenheimer, 1808, Schmett. Europa 2 : 2 
(c) Chimaera Hitchcock, 1858, Ichnol. N. England : 118 


. Chimaira Duméril, 1856, Mém. Acad. Sci. Inst. France 27 : 155 (an 


Erroneous Subsequent Spelling for Chimaera Linnaeus, 1758, 
placed on the Official List in Opinion 77) 


. The under-mentioned junior homonyms of Coregonus Linnaeus, 


1758, placed on the Official List in Opinion 93 :— 


') Coregonus Lacépede,. 1803, Hist.. nat. Poiss. 3: 239; Le 
Sueur, 1818, J. Acad. nat. Sci. Philad. 1(9) : 231 


(b) Coregonus Jarocki, 1822, Zoologiia 4 : 35 


. Coriudo Fleming, 1822, Phil. zool. 2: 271 (a junior objective 


synonym of Dermochelys Blainville, 1816, placed on the Official 
List in Opinion 92) 


Coryhaena Stiles, 1922, Smiths. misc. Coll. 73 (no. 1) : 73 (an 
Erroneous Subsequent Spelling for Coryphaena Linnaeus, 1758, 
placed on the Official List in Opinion 77) 


Coryphaena Coues, 1889, Cent. Dict. 2(5) : 1286 (a nomen nudum) 


Cyclopterus (Seidlitz MS) Marseul, 1872, Abeille 10 : 30 (a junior 
homonym of Cyclopterus Linnaeus, 1758, placed on the Official 
List in Opinion 77) 

Dermatochelys Wagler, 1830, Syst. Amph. : 133 (a junior objective 
synonym of Dermochelys Blainville, 1816, placed on the Official 
List in Opinion 92) 

Dermochelis Lesueur, 1829, in Cuvier, Régn. Anim. (ed. 2) 2: 14 


(an Erroneous Subsequent Spelling for Dermochelys Blainville, 
1816, placed on the Official List in Opinion 92) 


The under-mentioned junior homonyms of Diodon Linnaeus, 1758, 
placed on the Official List in Opinion 77 :— 


(a) Diodon O’Reilly, 1818, Greenland:105; Storr, 1780, 
Prodr. meth. Mamm. : 42, tab. C 


(b) Diodon Lesson, 1828, Compl. de Buffon, Hist. nat. Mamm. 
Ois. 1 : 124, 440 


(c) Diodon Lesson, 1830, Traité. Ornith. (2) : 95 


358 
16. 


20. 


zt 


22. 


5 Me 
24. 


OPINIONS AND DECLARATIONS 


The under-mentioned junior homonyms of Draco Linnaeus, 1758, 
placed on the Official List in Opinion 75 :— 


(a) Draco Gouan, 1770, Hist. Pisc. : 117 (a nomen nudum) 
(b) Draco Oken, 1816, Lehrb. Naturgesch. 3(2) : 273 


. Eleotris Gronovius, 1763, Zooph. gronoy.1 : 83 (a name published 


in a book rejected for nomenclatorial purposes) 


. The under-mentioned junior homonyms of Fistularia Linnaeus, 


1758, placed on the Official List in Opinion 75 :— 


(a) Fistularia Forskal, 1775, Descr. Anim. : 121 
(b) Fistularia Miller, 1776, Zool. dan. Prodr. : 254 


(c) Fistularia Lamarck, 1816, Hist. nat. Anim. sans Vertebr. 
3:74; de Blainville, 1830, Dict. Sci. nat. 60 : 173 


(d) Fistularia Bowerbank, 1841, Proc. geol. Soc. Lond. 3 (no. 
TS) 2433 


. The under-mentioned junior homonyms of Gadus Linnaeus, 1758, 


placed on the Official List in Opinion 77 :— 


(a) Gadus Dejean, 1821, Cat. Coléopt. : 96 (a nomen nudum) 

(b) Gadus Rang, 1829, Ann. Sci. nat. 16 : 498 

(c) Gadus Deshayes, 1861, Descr. Anim. s. Vertébr. Bassin Paris 
Ze 2)7 


Gymnothorax Cuvier, [1800], Legons. comp. Anat. 1: tab. 4 (a 
nomen nudum) 


Gymnothorax Duda, 1923, Arch. Naturgesch. 89 : A4, 39 (a junior 
homonym of Gymnothorax Bloch, 1795, placed on the Official 
List in Opinion 93) 


The under-mentioned Erroneous Subsequent Spellings for Mabuya 
Rafinesque, 1815 :— 
(a) Mabouia Cuvier, 1829, Régn. Anim. (ed. 2) 2 : 62 
(b) Mabouya Duméril & Bibron, 1839, Erpét, 5 : 663, 671 
(c) Mabuia Cuvier, 1829, Régn. Anim. (ed. 2) 2 : 64 


Mabuya Rafinesque, 1815, Analyse Nat. : 76 (a nomen nudum) 


The under-mentioned Erroneous Subsequent Spellings for 
Malapterurus Lacépéde, 1803, placed on the Official List in 
Opinion 93) :— 

(a) Malapterus Jarocki, 1822, Zoologiia 4 : 87 
(b) Malapturus Swainson, 1838, Hist. Classif. Fish 1 : 348, 353 


25: 


26. 


ah. 


28. 


22. 


30. 


SI. 


a2: 


35. 


34. 


DIRECTION 56 359 


Mugie Macklot, 1830, Bijd. Natuurk. Wetensch. 5:177 (an 
Erroneous Subsequent Spelling for Mugil Linnaeus, 1758, 
placed on the Official List in Opinion 75) 


Mustellus Fischer de Waldheim, 1813, Zoognosia (ed. 3) 1: 78 
(an Erroneous Subsequent Spelling for Mustelus Linck, 1790, 
placed on the Official List in Opinion 77) 


Ophidion Pomel, 1853, Cat. Vert. foss. Loire : 128 (a junior 
homonym of Ophidion Linnaeus, 1758, placed on the Official 
List in Opinion 92) 


Ophidium Linnaeus, 1766, Syst. Nat. (ed. 12) 1 : 431 (an Invalid 
Emendation of Ophidion Linnaeus, 1758, placed on the Official 
List in Opinion 92) 


Siren Hermann, 1783, Tab. Affin. Anim. : 120 (a junior homonym 
of Siren Linnaeus, 1766, placed on the Official List in Opinion 92) 


Sirena Fischer de Waldheim, 1808, Zoognosia (ed. 2): tab. ili 
(an Invalid Emendation of Siren Linnaeus, 1766) 


Sirene Oken, 1816, Lehrb. Naturgesch 3(2) : 187 (an Invalid 
Emendation of Siren Linnaeus, 1766) 


Sphargis Merrem, 1820, Tent. Syst. Amph. : 19 (a junior objective 
synonym of Dermochelys Blainville, 1816, placed on the Official 
List in Opinion 92) 


Syngnathus Rafinesque, 1810, Ind. Itt. Sic. : 57 (a junior homonym 
of Syngnathus Linnaeus, 1758, placed on the Official List in 
Opinion 77) 


Teuthis Schneider, 1784, Sammi. verm. Abh.: 113 (a junior 
homonym of Teuthis Linnaeus, 1766, placed on the Official 
List in Opinion 93) 


Section B: Proposed addition of two specific names to the 
** Official Index of Rejected and Invalid Specific Names 
in Zoology ”’ 


. linnaei Boie, 1826, Bull. Sci. nat. Geol. 9 : 236, as published in the 


combination Calamaria linnaei (nom. nov. pro calamarius 
Linnaeus, 1758, as published in the combination Coluber 
calamarius)* 


. mercurialis Merrem, 1820, Tent. Syst. Amph. : 19, as published 


in the combination Sphargis mercurialis (nom. noy. pro coriacea 
Linnaeus, 1766, as published in the combination Testudo 
coriacea) 


* This proposal was later withdrawn. See paragraph 4 of the present Direction. 


360 OPINIONS AND DECLARATIONS 


3. alleghaniensis Harlan, 1825, Ann. Lyceum nat. Hist. New York 
1(8) : 271, as published in the combination Abranchus 
alleghaniensis (an Invalid Emendation of alleganiensis Daudin, 
[1803] as published in the combination Salamandra 
alleganiensis. 


2. Registration of the present application : Upon the receipt 
of Mr. Hemming’s paper the question of completing and 
correcting in the manner proposed the entries relating to the names 
of genera belonging to the Classes Pisces, Amphibia and Reptilia 
made on the Official List of Generic Names in Zoology in the 
period up to the end of 1936 was allotted the Registered Number 
Z.N.(S.) 1015. 


I]. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


3. Issue of Voting Paper V.P.(O.M.)(56)4 : On 25th April 1956 
a Voting Paper (V.P.(O.M.)(56)4) was issued in which each 
Member of the Commission was asked (1) to state whether he 
agreed “that, in accordance with the General Directives issued 
to the International Commission on Zoological Nomenclature 
by the Thirteenth International Congress of Zoology, Paris, 1948, 
regarding the placing on the appropriate Official Lists and 
Official Indexes of names dealt with in, or otherwise involved 
in, Opinions rendered prior to 1948, and other matters on which 
action is required in order to complete, to clarify, or to correct 
Rulings given in Opinions rendered in the foregoing period, the 
action recommended in paragraph 14 of the paper numbered 
Z.N.(S.) 1015, regarding names in the Classes Pisces, Amphibia 
and Reptilia submitted by the Secretary simultaneously with the 
present Voting Paper [i.e. in the paragraph numbered as above 
in the paper reproduced in the first paragraph of the present 
Direction] be now taken as there recommended ’”’, and (2), if 
he did not so agree as regards any given item, to indicate that 
item. 


DIRECTION 56 361 


4. Withdrawal from the scope of Voting Paper V.P.(O.M)(56)4 
of certain proposals relating to the generic name ‘* Calamaria ”’ 
Boie (Class Reptilia): On 2nd May 1956 Mr. Hemming, as 
Secretary, executed the following Minute in which he withdrew 
from the scope of Voting Paper V.P.(O.M.)(56)4 certain of the 
proposals in regard to the generic name Calamaria Boie submitted 
therewith :— 


Remoyal from the scope of Voting Paper V.P.(O.M)(56)4 of certain of 
the proposals relating to the generic name ‘* Calamaria’’ Boie 
(Class Reptilia) submitted therewith 


By FRANCIS HEMMING, C.M.G., C.B.E. 
(Secretary to the International Commission on Zoological Nomenclature) 


In returning their completed copies of Voting Paper V.P.(O.M.)(56)4, 
Dr. H. Boschma, Director, Rijksmuseum van Natuurlijke Historie, 
Leiden, The Netherlands, and Dr. L. B. Holthuis of the same Museum 
have this day notified the Office of the Commission that Dr. L. D. 
Brongersma of the above Museum is planning to re-examine the 
status under the Régles of the generic name Calamaria Boie (Class 
Reptilia) a name which had been placed on the Official List of Generic 
Names in Zoology by the Ruling given in Opinion 92 and on which 
certain supplementary proposals had been submitted in the paper 
bearing the Number Z.N.(S.) 1015 issued to the Members of the 
Commission concurrently with the Voting Paper referred to above. It 
appeared from the communication so received that the current 
interpretation of the nominal species Coluber calamarius Linnaeus, 
1758, the type species of the genus Calamaria Boie, and/or of the 
objectively identical nominal species Calamaria linnaei Boie, 1826, 
was open to question and that there was at least a possibility that the 
species to which Linnaeus had given the former name was a species 
at present placed by specialists in a genus far removed from Calamaria 
Boie. 


2. In these circumstances, as Secretary to the International Com- 
mission, I hereby withdraw from the scope of the Voting Paper cited 
above (a) the proposals in regard to the generic name Calamaria 
Boie submitted in Points (6) and (7) of paragraph 14 of the paper 
Z.N.(S.) 1015 submitted therewith, and (b) the proposal which appears 
as Item I in Section B of Annexe 2 referred to in Point (11) in the 
paragraph numbered as above in the same paper in order to permit 
of the further study of the issues alluded to in the communications 
referred to in paragraph | above. I further direct that the Registered 
File Z.N.(S.) 1114 be opened for the study of the questions now 
reserved and that the said studies be prosecuted as vigorously as 


362 OPINIONS AND DECLARATIONS 


possible in order to ensure that the publication of the Official List of 
Generic Names in Zoology in book-form be not delayed by the 
existence of an outstanding problem in regard to the above generic 
name. 


5. The Prescribed Voting Period for V.P.(O.M)(56)4: As 
Voting Paper V.P.(O.M.)(56)4 was issued under the One-Month 
Rule, the Prescribed Voting Period closed on 25th May 1956. 


6. Particulars of the Voting on Voting Paper V.P.(O.M)(56)4 : 
At the close of the Prescribed Voting Period for Voting Paper 
V.P.(O.M.)(56)4, the state of the voting on the proposals sub- 
mitted therewith, other than on that part of those proposals 
which had been withdrawn from the scope of that Voting Paper 
by the Minute executed by the Secretary on 2nd May 1956 (the 
text of which has been reproduced in paragraph 4 of the present 
Direction) was as follows :— 


(a) Affirmative Votes had been given by the following twenty-four 
(24) Commissioners (arranged in the order in which Votes 
were received) : 


Holthuis ; Boschma ; Stoll; Vokes; Hering; Miller ; 
Prantl ; Bodenheimer; Cabrera; Esaki; Mayr; do 
Amaral ; Lemche ; Hanké ; Dymond ; Mertens; Key ; 
Jaczewski; Bonnet; Hemming; Sylvester-Bradley ; 
Riley ; Kuhnelt ; Tortonese ; 


(b) Negative Votes : 


None ; 


(c) On Leave of Absence, one (1): 


Bradley (J.C.) ; 


DIRECTION 56 363 


(d) Voting Papers not returned : 


None. 


7. Declaration of Result of Vote: On 7th June 1956, Mr. 
Hemming, Secretary to the International Commission, acting 
as Returning Officer for the Vote taken on Voting Paper V.P. 
(O.M.)(56)4, signed a Certificate that the Votes cast on that 
Voting Paper, other than on the proposals in regard to the name 
Calamaria Boie submitted therewith, which had been withdrawn 
from its scope by the Minute executed by the Secretary on 
2nd May 1956 (the text of which has been reproduced in para- 
graph 4 of the present Direction) were as set out in paragraph 6 
above and that the proposals submitted with that Voting Paper, 
other than the proposals so withdrawn, had been duly adopted and 
that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


8. Preparation of the Ruling given in the present ‘* Direction ”’ : 
On 30th September 1956, Mr. Hemming prepared the Ruling 
given in the present Direction and at the same time signed a 
Certificate that the terms of that Ruling were in complete accord 
with those of the proposal approved by the International Com- 
mission in its vote on Voting Paper V.P.(O.M.)(56)4, subject to 
the exclusion therefrom of the proposals in regard to the generic 
name Calamaria Boie specified in the Minute executed by the 
Secretary on 2nd May 1956 referred to in paragraph 7 above. 


9. Original References : The original references for the generic 
names specified in the under-mentioned Rulings in the present 
Direction have already been noted in connection with the Official 
List of Generic Names in Zoology: (1); (2); (3); (5), subject 
to the correction of the date of publication attributed to the 
name concerned as specified in Ruling (4) ; (6) ; (7); (8), subject 
to the correction of the date of publication of the name concerned 
as there specified. The references for the selection of typespecies for 
the genera specified in Rulings (2), (3) and (5) are given in Annexe | 
to the paper reproduced in paragraph 1 of the present Direction. 


364 OPINIONS AND DECLARATIONS 


The reference for the nominal species specified in Ruling (7) as 
the type species of Eleotris Schneider, 1801, is given in paragraph 8 
of the paper referred to above, while in paragraph 10 of that paper 
will be found the reference specified in Ruling (9). The references 
for the names specified in Rulings (10) and (11) are given in 
Section A and Section B respectively of Annexe 2 to the foregoing 
paper. 


10. The prescribed procedures were duly complied with by 
the International Commission on Zoological Nomenclature in 
dealing with the present case, and the present Direction is 
accordingly hereby rendered in the name of the said International 
Commission by the under-signed Francis Hemming, Secretary 
to the Jnternational Commission on Zoological Nomenclature, 


in virtue of all and every the powers conferred upon him in that 
behalf. 


11. The present Direction shall be known as Direction Fifty-Six — 
(56) of the International Commission on Zoological Nomenclature. 


Done in London, this Thirtieth day of September, Nineteen 
Hundred and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


Printed in England by MretcaLtFe & COOPER LimITED, 10-24 Scrutton St., London EC 2 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, c.M.G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.18. Pp. 365—388 


DIRECTION 57 


Addition to the Official List of Specific Names in Zoology 
(a) of the specific names of forty-seven species belonging 
to the Classes Cyclostomata, Pisces, Amphibia and 
Reptilia, each of which is the type species of a genus, the 
name of which was placed on the Official List of Generic 
Names in Zoology in the period up to the end of 1936 and 
(b) of the specific name of one species of the Class 
Amphibia which is currently treated as a senior subjective 
synonym of the name of such a species 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1956 


Price Sixteen Shillings bs ae 
(All rights reserved) i 


Issued 20th December, 1956 


INTERNATIONAL COMMISSION ON | 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
RULING GIVEN IN DIRECTION 57 


A. The Officers of the Commission 


Honorary Life President : Dr. Karl JorpAN (British Museum (Natural History), Zoological 
Museum, Tring, Herts, England) 


President : Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) 
(12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(Arranged in order of precedence by reference to-date of election or of most recent re-election, 
as prescribed by the International Congress of Zoology) 


Professor H. BOSCHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(1st January 1947) 

Senor Dr. Angel CABRERA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

~ tia LEMCHE (Universitetets Zoologiske Museum, Copenhagen, Denmark) (27th 
uly 1948) 

Professor Teiso EsAKi (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEY (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEWSKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a. M., Germany) (Sth July 1950) 

Professor Erich Martin HERING (Zoologisches Museum der Humboldt- Universitat zu Berlin, 
Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice President) 

Professor J. R. DyMOND (University of Toronto, Toronto, Canada) (12th August 1953) 

Professor J. Chester BRADLEY (Cornell University, Ithaca, N.Y., U.S.A.) (12th August 
1953) (President) 

Professor Harold E. Voxes (Johns Hopkins University, Baltimore, Maryland, U.S.A.) 
(12th August 1953) 

Professor Béla HANKO (Mezdgazdasdgi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. StToLt (Rockefeller Institute for Medical Research, New York, N.Y., 
U.S.A.) (12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. Hottuuts (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (15th October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum vy Praze, Prague, Czechosiovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KUHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

Professor F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 


Professor Ernst MAYR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico TORTONESE (Museo di Storia Naturale ‘“‘G. Doria,’’ Genova, Italy) 
(16th December 1954) 


DIRECTION 57 


ADDITION TO THE ‘“ OFFICIAL LIST OF SPECIFIC 
NAMES IN ZOOLOGY ” (a) OF THE SPECIFIC NAMES 
OF FORTY-SEVEN SPECIES BELONGING TO THE 

CLASSES CYCLOSTOMATA, PISCES, AMPHIBIA 
AND REPTILIA, EACH OF WHICH IS THE TYPE 
SPECIES OF A GENUS, THE NAME OF WHICH 

WAS PLACED ON THE “ OFFICIAL LIST 

OF GENERIC NAMES IN ZOOLOGY ” IN 
THE PERIOD UP TO THE END OF 1936 

AND (b) OF THE SPECIFIC NAME OF 

ONE SPECIES OF THE CLASS 

AMPHIBIA WHICH IS CURRENTLY 
TREATED AS A SENIOR SUBJEC- 
TIVE SYNONYM OF THE NAME 
OF SUCH A SPECIES 


RULING :—(1) The under-mentioned specific name 
of a species belonging to the Class Cyclostomata, which 
is the type species of a genus, the name of which was 
placed on the Official List of Generic Names in Zoology 
in the period up to the end of 1936 by the Ruling given in 
Opinion 75, is hereby placed on the Official List of Specific 
Names in Zoology with the Name Number 1063: 
glutinosa Linnaeus, 1758, as published in the combination 
Myxine glutinosa (specific name of type species of Myxine 
Linnaeus, 1758). 


(2) The under-mentioned specific names of species 
belonging to the Class Pisces, each of which is the type 
species of a genus, the name of which was placed on the 
Official List of Generic Names in Zoology in the period 
up to the end of 1936 by the Rulings given in the Opinions 
severally cited below, are hereby placed on the Official 


368 OPINIONS AND DECLARATIONS 


Lisi of Specific Names in Zoology with the Name Numbers 
specified in each case against the names in question :— 


(a) Specific Names of type species of genera, the names 
of which were placed on the ** Official List” by 
the Ruling given in“ Opinion” 713 :—_ ~ 


(i) tobianus Linnaeus, 1758, as published 
in the combination Ammodytes tobianus 
(specific name of type species of 
Ammodytes Linnaeus, 1758) (Name 
No. 1064) ; 


(it) /upus Linnaeus, 1758, as published in 
the combination Anarhichas lupus 
(specific name of type species of 
Anarhichas Linnaeus, 1758) (Name No. 
1065) ; 


(iii) Hepsetus Linnaeus, 1758, as published 
in the combination Atherina hepsetus 
(specific name of type species of 
Atherina Linnaeus, 1758) (Name No. 
1066) ; 


(iv) tabacaria Linnaeus, 1758, as published in 
the combination Fistularia tabacaria 
(specific, name lof type specressenr 
Fistularia Linnaeus, 1758) (Name No. 
1067) ; 


(v) cephalus Linnaeus, 1758, as published in 
the combination Mugil cephalus (specific 
name of type species of Mugil Linnaeus, 
1758) (Name No. 1068) ; 


(v1) draco Linnaeus, 1758, as published in the 
combination Jrachinus draco (specific 
name of type species of Trachinus 
Linnaeus, 1758) (Name No. 1069) ; 


(vil) scaber Linnaeus, 1758, as published in the 
combination Uranoscopus scaber (speci- 
fic name of type species of Uranoscopus 
Linnaeus, 1758) (Name No. 1070) ; 


DIRECTION 57 369 


(viii) gladius Linnaeus, 1758, as published in the 
combination Xiphias gladius Linnaeus, 
1758 (specific name of type species of 
Xiphias Linnaeus, 1758) (Name No. 
POTAY 


(ix) sturio Linnaeus, 1758, as published in the 
combination Acipenser sturio (specific 
name of type species of Acipenser 
Linnaeus, 1758) (Name No. 1072) ; 


(x) /yra Linnaeus, 1758, as published in the 
combination Callionymus lyra (specific 
name of type species of Callionymus 
Linnaeus, 1758) (Name No. 1073) ; 


(xi) monstrosa Linnaeus, 1758, as published 
in the combination Chimaera 
monstrosa (specific name of type species 
of Chimaera Linnaeus, 1758) (Name 
No. 1074) ; 


(xii) harengus Linnaeus, 1758, as published in 
the combination Clupea harengus 
(specific name of type species of Clupea 
Linnaeus, 1758) (Name. No. 1073). : 


(xil) hippurus Linnaeus, 1758, as published 
in the combination Coryphaena 
hippurus (specific name of type species 
of Coryphaena Linnaeus, 1758) (Name 
No. 1076) ; 


(xiv) /umpus Linnaeus, 1758, as published in the 
combination Cyclopterus lumpus (speci- 
fic name of type species of Cyclopterus 
Linnaeus, 1758) (Name No. 1077) ; 


(xv) carpio Linnaeus, 1758, as published in the 
combination Cyprinus carpio (specific 
name of type species of Cyprinus 
Linnaeus, 1758) (Name No. 1078) ; 


(xvi) morhua Linnaeus, 1758, as published in the 
combination Gadus morhua (specific 


370 OPINIONS AND DECLARATIONS 


name of type species of Gadus Linnaeus, 
1758) (Name No. 1079) ; 


(xvil) aculeatus Linnaeus, 1758, as published in 
the combination Gasterosteus aculeatus 
(specific name-of type species on 
Gasterosteus Linnaeus, 1758) (Name 
No. 1080) ; 


(xviii) niger Linnaeus, 1758, as published in the 
combination Gobius niger (specific name 
of type species of Gobius Linnaeus, 
1758) (Name No. 1081) ; 


(xix) piscatorius Linnaeus, 1758, as published 
in the combination Lophius piscatorius 
(specific name of type species of 
Lophius Linnaeus, 1758) (Name No. 
1082) ; 


(xx) cyprinoides Linnaeus, 1758, as published 
in the combination Mormyrus 
cyprinoides (specific name of type 
species of Mormyrus Linnaeus, 1758) 
(Name No. 1083) ; 


(xx1) barbatus Linnaeus, 1758, as published in 
the combination Mullus barbatus (speci- 
fic name of type species of Mullus 
Linnaeus, 1758) (Name No. 1084) ; 


(xxl) fluviatilis Linnaeus, 1758, as published in 
the combination Perca fluviatilis (speci- 
fic name of type species’ of Pere 
Linnaeus, 1758) (Name No. 1085) ; 


(xxl) salar Linnaeus, 1758, as published in the 
combination Salmo salar (specific name 
of type species of Salmo Linnaeus, 
1758) (Name No. 1086) ; 


(xxiv) scombrus Linnaeus, 1758, as published in 
the combination Scomber scombrus 
(specific name of type species of 


DIRECTION 57 371 


Scomber Linnaeus, 1758) (Name No. 
1087) ; 


(xxv) porcus Linnaeus, 1758, as published in 
the combination Scorpaena porcus (spe- 
cific name of type species of Scorpaena 
Linnaeus, 1758) (Name No. 1088) ; 


(xxv1) glanis Linnaeus, 1758, as published in 
the combination Si/urus glanis (specific 
name of type species of Silurus 
Linnaeus, 1758) (Name No. 1089) ; 


(xxvil) acus Linnaeus, 1758, as published in the 
combination Syngnathus acus (specific 
name of type species of Syngnathus 
Linnaeus, 1758) (Name No. 1090) ; 


(xxvil) faber Linnaeus, 1758, as published in the 
combination Zeus faber (specific name 
of type species of Zeus Linnaeus, 1758) 
(Name No. 1091) ; 


(b) Specific Names of type species of genera, the names 
of which were placed on the “ Official List” by 
the Ruling given in ** Opinion” 92 :—- 


(i) ocellaris Linnaeus, 1758, as published in 
the combination Blennius ocellaris (spe- 
cific name of type species of Blennius 
Linnaeus, 1758) (Name No. 1092) ; 


(11) /ucius Linnaeus, 1758, as published in the 
combination Esox lucius (specific name 
of type species of Esox Linnaeus, 1758) 
(Name No. 1093) ; 


(ii) barbatus Linnaeus, 1758, as published in 
the combination Ophidion barbatus 
(specific name of type species of 
Ophidion Linnaeus, 1758) (Name No. 
1094) ; 


at2 OPINIONS AND DECLARATIONS 


(c) Specific Names of type species of genera, the names of 
which were placed on the “ Official List”? by the 
Ruling given in “‘ Opinion”? 93 :— 


(1) /avaretus Linnaeus, 1758, as published 
in the combination Salmo lavaretus 
(specific name of type species of 
Coregonus Linnaeus, 1758) (Name No. 
1095) ; 


(11) pisonis Gmelin (J.F.), [1789], as published 
in the combination Gobius pisonis 
(specific name of type species of 
Eleotris Schneider, 1801) (Name No. 
1096) ; 


(i1) marginalis Bloch, 1793, as published in the 
combination Epinephelus marginalis, 
(specific name of type species of 
Epinephelus Bloch, 1793) (Name No. 
1097) ; 


(iv) reticularis Bloch, 1795, as published in 
the combination Gymnothorax 
reticularis (specific name of type species 
of Gymnothorax Bloch, 1795) (Name 
No. 1098) ; 


(v) electricus Gmelin (J.F.), [1789], as pub- 
lished in the combination Silurus 
electricus (specific name of type species 
of Malapterurus Lacépéde, 1803) (Name 
No. 1099) ; 


(vi) mustelus Linnaeus, 1758, as published in 
the combination Squalus mustelus (spe- 
cific name of type species of Mustelus 
Linck, 1790) (Name No.1100) ; 


(vii) paradiseus Linnaeus, 1758, as published 
in the combination Polynemus 
paradiseus (specific name of type species 
of Polynemus Linnaeus, 1758) (Name 
No. 1101) ; 


DIRECTION 57 6 


(viii) cabrilla Linnaeus, 1758, as published in 
the combination Perca cabrilla (specific 
name of type species of Serranus 
Cuvier, 1817) (Name No. 1102). 


(3) The under-mentioned specific names of species 
belonging to the Class Amphibia, each of which is the 
type species of a genus, the name of which was placed on 
the Official List of Generic Names in Zoology in the period 
up to the end of 1936 by the Ruling given in Opinion 92, 
are hereby placed on the Official List of Specific Names in 
Zoology with the Name Numbers severally specified 
below :— 


(a) fuscus Rafinesque, 1820, as published in the com- 
bination Triturus fuscus (specific name of type 
species of Desmognathus Baird, (1850)) (Name 
Ne, 1103):; 


(b) Jacertina Linnaeus, 1766, as published in the com- 
bination Siren lacertina (specific name of type 
species of Siren Linnaeus, 1766) (Name No. 
1104). 


(4) The under-mentioned specific names of species 
belonging to the Class Reptilia, each of which is the type 
species of a genus, the name of which was placed on the 
Official List of Generic Names in Zoology in the period 
up to the end of 1936 by the Rulings given in the Opinions 
severally cited below, are hereby placed on the Official 
List of Specific Names in Zoology with the Name Numbers 
specified in each case against the names in question :— 


(a) Specific Name of the type species of a genus, the 
name of which was placed on the * Official List” 
by the Ruling given in “ Opinion” 75: 


volans Linnaeus, 1758, as published in the 
combination Draco volans (specific name 


374 OPINIONS AND DECLARATIONS 


of type species of Draco Linnaeus, 1758) 
(Name No. 1105) ; 


(b) Specific Names of type species of genera, the names 
of which were placed on the “ Official List’? by 
the Ruling given in “* Opinion”? 92 :— 


(1) serpentina Linnaeus, 1758, as published in 
the combination Testudo serpentina 
(specific name of type species of 
Chelydra Schweigger, 1812) (Name No. 
1106) ; 


(ii) horridus Linnaeus, 1858, as published i in 
the combination Crotalus horridus (spe- 
cific name of type species of Crotalus 
Linnaeus, 1758) (Name No. 1107) ; 


(iii) coriacea Linnaeus, 1766, as published in 
the combination Testudo coriacea (spe- 
cific name of type Species 
Dermochelys Blainville, 1816) (Name 
No: 1 108) 5 


(iv) agilis Linnaeus, 1758, as published in the 
combination Lacerta agilis (specific 
name of type species of Lacerta 
Linnaeus, 1758) (Name No. 1109). 


(5) The under-mentioned specific name which is cur- 
rently regarded as a senior subjective synonym of the 
specific name of the type species of a genus belonging to 
the Class Amphibia, the name of which was placed on the 
Official List of Generic Names in Zoology in the Opinion 
cited against the name in question is hereby placed on the 
Official List of Specific Names in Zoology with the Name 
Number specified below :— 


alleganiensis Daudin, [1803], as published in the 
combination Salamandra alleganiensis (Opinion 92) 
(Name No. 1110). 


DIRECTION 57 a7 


iY THE SUBIECT MATTER OFTHE PRESENT 
* DERECTION ” 


The present Direction has been prepared under a General 
Directive issued to the International Commission on Zoological 
Nomenclature by the Thirteenth International Congress of 
Zoology, Paris, 1948, that the specific name of the species which 
are the type species of genera, the names of which had been 
placed on the Official List of Generic Names in Zoology prior to the 
opening of that Congress, should, if the oldest available names 
for the species concerned, be placed on the Official List of Specific 
Names in Zoology and that in other cases there should be placed 
on that List whatever is currently considered to be the oldest 
available names for the species concerned. The present which 
is the fourth in this series of Directions, is concerned with the names 
of the type species of genera belonging to the Classes Cyclostomata, 
Pisces, Amphibia and Reptilia, the names of which were placed 
on the Official List of Generic Names in Zoology in the period 
up to the end of 1936. The total number of such generic names is 
forty-eight. Of these all are dealt with in the present Direction 
with the exception of the generic name Calamaria Boie, 1826, the 
consideration of which has, for the reasons explained in paragraph 
4 of the present Direction, been postponed for further study. 
The proposals which formed the basis of the decisions taken by 
the Commission in the present case were submitted by the 
Secretary in the following paper on 25th April 1956 :— 


Proposed addition to the ‘*‘ Official List of Specific Names in Zoology ”’ 
(a) of the specific names of forty-eight species belonging to the Classes 
Cyclostomata, Pisces, Amphibia and Reptilia, each of which is the 
type species of a genus, the name of which was placed on the 
** Official List of Generic Names in Zoology ”’ in the period 
up to the end of 1936 and (b) of the specific name of one 
species of the Class Amphibia which is currently treated 
as a senior subjective synonym of the name of such a 
species 


By FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


The present paper contains the fourth instalment of proposals for 
giving effect to the General Directive issued to the International 


376 OPINIONS AND DECLARATIONS 


Commission on Zoological Nomenclature by the Thirteenth Inter- 
national Congress of Zoology, Paris, 1948, that in every case where the 
specific name of the type species of a genus, the name of which was 
placed on the Official List of Generic Names in Zoology in the period 
up to the end of 1936 is the oldest available name for the species 
concerned, that name shall be placed on the Official List of Specific 
Names in Zoology and that in other cases the specific name currently 
regarded as the oldest available subjective synonym of that name shall 
be similarly placed on that List. 


2. The proposals now submitted relate to the names of the type 
species of genera belonging to the Classes Cyclostomata, Pisces, 
Amphibia and Reptilia, which were placed on the Official List of 
Generic Names in Zoology in the period up to the end of 1936. In all, 
sixty-two generic names are involved. The genera so named are dis- 
tributed among the foregoing Classes as follows : Class Cyclostomata, 
one genus ; Class Pisces, 48 genera ; Class Amphibia, 3 genera ; Class 
Reptilia, 10 genera. Details regarding the position of the names 
belonging to each of these groups are given in the following 
paragraphs. 


3. Class Cyclostomata: No difficulty arises in connection with 
the specific name of the type species of the only genus belonging to 
this Class, the name of which has been placed on the Official List 
(Myxine Linnaeus, 1758) and the specific name of that species should 
therefore now be placed on the Official List. 


4. Class Pisces : In the case of one of the genera of the Class Pisces 
involved, the specific name of the type species has already been placed 
on the Official List. The generic name concerned is Echeneis Linnaeus, 
1758, which was dealt with by the Commission in Opinion 242. In 
another case—relating to the reputed generic name Osmerus Linnaeus, 
1758—proposals covering inter alia the proposed addition of the specific 
name of its type species to the Official List are already before the 
Commission in Application Z.N.(S.) 564 (Hemming, 1955, Bull. zool. 
Nomencl, 11 : 281—282). In two cases applications have been 
received from ichthyologists which will, it is hoped, be published in the 
Bulletin of Zoological Nomenclature at an early date. Three names are 
involved in these applications, namely (a) Sciaena Linnaeus, 1758 
(Application Z.N.(S.) 850); (b) Stolephorus Lacépéde, 1803, and 
Teuthis Linnaeus, 1766 (Application Z.N.(S.) 569). In these cases it is 
proposed that action regarding the addition to the Official List of the 
specific names of the type species of the genera concerned be deferred 
until decisions are taken on other outstanding matters concerned with 
these generic names. Finally, there are four generic names against 
which no objection has been raised by specialists but which, on the 
basis of the information so far collected by this Office, appear to call 
for action by the Commission before the entries in regard to them 


DIRECTION 57 371 


made on the Official List in the period up to the end of 1936 can be 
regarded as correct. The problems involved in these cases are of two 
kinds. In one case the name in question, Conger Cuvier, 1817, appears 
to be a phantom ; in the three other cases (Cottus Linnaeus, 1758 ; 
Diodon Linnaeus, 1758 ; Muraena Linnaeus, 1758) it appears that the 
species currently accepted as the type species and adopted as such 
in the Official List is not the first of the originally included species to 
have been selected as the type species of the genus in question. Con- 
sultations are proceeding with representative ichthyologists in regard 
to these four names (on Commission File Z.N.(S.) 556) and it is 
proposed in these cases to defer for the present the submission of 
proposals for the addition to the Official List of the specific names of 
the species currently accepted as the type species of the genera 
concerned. 


5. It will be seen from the particulars given above that the action 
required has already been taken in regard to the name of the type species 
of one of the genera involved, while in eight other cases it is con- 
sidered that at the present moment it would be premature or otherwise 
undesirable to place on the Official List the specific name of the type 
species in question. In the case of the remaining thirty-nine generic 
names in the Class Pisces investigations carried out by this Office 
show (a) that the species accepted in the Opinions concerned as the 
type species of the genera involved are currently accepted as such 
by specialists, (b) that the species concerned have been validly desig- 
nated or indicated as the type species of the genera in question and that 
where the determination of the type species has to be made under 
Rule (g) in Article 30 (type species by subsequent selection), the 
species have been validly selected to be the type species of the genera 
concerned, and (c) that the specific names of the nominal species 
which are the type species of the genera concerned are the oldest 
available names for the said species. In these circumstances it is now 
recommended that the thirty-nine specific names involved should be 
placed on the Official List. 


6. Class Amphibia: The names of the three genera of the Class 
Amphibia which have been placed on the Official List are all available 
names and are currently accepted by specialists as the oldest such 
names for the genera concerned. In each case also the type species 
has been correctly determined under Article 30 of the Réegles. In the 
case of two of the genera concerned (Desmognathus Baird, (1850) ; 
Siren Linnaeus, 1766) the specific name of the type species is the oldest 
available name for the species concerned and is currently accepted as 
such. These two specific names should therefore now be placed on the 
Official List. In the case of the third of the genera involved 
(Cryptobranchus Leuckart, 1821) the specific name of the type species 
(gigantea) is currently regarded as a junior subjective synonym of 
another name (alleganiensis). In this case it is the specific name which 
is the senior subjective synonym and not the specific name of the type 


378 OPINIONS AND DECLARATIONS 


species which under the General Directive issued to the Commission 
by the International Congress of Zoology should now be placed on the 
Official List. 


7. Class Reptilia : The specific name of the type species of one of the 
ten genera of the Class Reptilia which have to be considered has already 
been placed on the Official List of Specific Names in Zoology. The genus 
concerned is Mabuya Fitzinger, 1826, which was dealt with by the 
Commission in its Opinion 240. In the case of the name Phrynosoma 
Wiegmann, 1828, attention has been drawn by a specialist to the 
existence of certain names which appear to be senior synonyms 
(Application Z.N.(S.) 399) and it is not desirable that the specific 
name of the type species of this genus should be dealt with by the 
Commission until the more general issues raised can also be settled. 
In the case of a third name, Eremias Wiegmann, 1834, the selection 
as type species of the species accepted as such in the Official List and 
currently so accepted by specialists appears to have been anticipated 
by the selection of a different species. This question is now being 
investigated in conjunction with interested specialists (Z.N.(S.) 556) 
and it is not considered that any action is desirable until the present 
consultations have been concluded. In the case also of the name 
Alligator Cuvier, 1807, certain difficulties have been noted (Z.N.(S.) 551) 
and consultations are proceeding with specialists!. In this case also 
the question of placing the specific name of the type species on the 
Official List must be left over until comprehensive proposals can be 
laid before the Commission. 


8. No difficulty arises in connection with the remaining six generic 
names in the Class Reptilia. The names themselves are available 
names and are currently accepted by specialists as the oldest such names 
for the genera concerned ; the species currently accepted as the type 
species of these genera have been correctly determined as such under 
Article 30 of the Régles, and the specific names for those species are 
available names and are the oldest such names for the species in 
question. The specific names of the type species of these six genera 
should therefore now be placed on the Official List. 


9. Summarising the conclusions set forth in the present paper, I 
recommend that the International Commission on Zoological 
Nomenclature should :— 

(1) place on the Official List of Specific Names in Zoology the specific 
names specified below, each of which is the specific name of 
the type species of a genus, the name of which was placed 
on the Official List of Generic Names in Zoology in the period 
up to the close of 1936 :— 


(a) the specific name of a species of the Class Cyclostomata 
cited in Section A of Annexe 1 attached hereto ; 


1 Proposals for dealing with this question have now been submitted to the 
International Commission (Hemming, 1956, Bull. zool. Nomencl. 12 : 163-175). 


DIRECTION 57 379 


(b) the thirty-nine specific names of species of the Class Pisces 
cited in Section B of Annexe | attached hereto ; 


(c) the two specific names of species of the Class Amphibia 
cited in Section C of Annexe | attached hereto ; 


(d) the six specific names of species of the Class Reptilia cited 
in Section D of Annexe | attached hereto ; 


(2) place on the Official List of Specific Names in Zoology the specific 
name of a species of the Class Amphibia (being a name 
currently regarded as a senior subjective synonym of the name 
of a species which is the type species of a genus the name of 
which was placed on the Official List cf Generic Names in 
Zoology in the period up to the close of 1936) cited in Annexe 2 
attached hereto. 


ANNEXE | 


Proposed addition to the ‘‘ Official List of Specific Names in Zoology ”’ 
of forty-eight specific names of species belonging to the Classes 
Cyclostomata, Pisces, Amphibia and Reptilia, each of which is the 
specific name of the type species of a genus, the name of which 
was placed on the °° Official List of Generic Names in Zoology ”’ 

) in the period up to the end of 1936 


Specific Name Combination in which Name of genus of 
proposed to be placed name specified in which species 
on the *‘ Official List — Col. 1 was originally specified in Col. | 
of Specific Names in published is the type species 

Zoology” 
(1) (2) (3) 


Section A : Class Cyclostomata (one name) 


** Opinion ”’ 75 


glutinosa Linnaeus, Myxine glutinosa Myxine Linnaeus, 
1758 1758 


Section B : Class Pisces (thirty-nine names) 
** Opinion *’ 75 


tobianus Linnaeus, Ammodytes tobianus | Ammodytes Linnaeus, 
1758 | 1758 
lupus Linnaeus, 1758 = Anarhichas lupus Anarhichas Linnaeus, 
| | 1758 
hepsetus Linnaeus, Atherina hepsetus Atherina Linnaeus, 


1758 | 1758 


380 


Specific name 
proposed to be placed 
on the “* Official List 
of Specific Names in 

Zoology” 
(1) 
tabacaria Linnaeus, 
1758 
cephalus Linnaeus, 
1758 
draco Linnaeus, 1758. 


scaber Linnaeus, 1758 


gladius Linnaeus, 
1758 


sturio Linnaeus, 1758 
lyra Linnaeus, 1758 


monstrosa Linnaeus, 
1758 

harengus Linnaeus, 
1758 

hippurus Linnaeus, 
1758 

lumpus Linnaeus, 1758 


carpio Linnaeus, 1758 


morhua Linnaeus, 
1758 

aculeatus Linnaeus, 
1758 

niger Linnaeus, 1758 

piscatorius Linnaeus, 
1758 

cyprinoides Linnaeus, 
1758 

barbatus Linnaeus, 
1758 

fluviatilis Linnaeus, 
1758 

salar Linnaeus, 1758 


scombrus Linnaeus, 
1758 


Combination in which 


name specified in 
Col. 1 was originally 
published 


(2) 


Fistularia tabacaria 
Musgil cephalus 
Trachinus draco 
Uranoscopus scaber 
Xiphias gladius 

** Opinion *’ 77 
Acipenser sturio 
Callionymus lyra 
Chimaera monstrosa 
Clupea harengus 
Coryphaena hippurus 
Cyclopterus lumpus 
Cyprinus carpio 
Gadus morhua 
Gasterosteus 


aculeatus 
Gobius niger 


Lophius piscatorius 
Mormyrus cyprinoides 
Mullus barbatus 
Perca fluviatilis 


Salmo salar 
Scomber scombrus 


OPINIONS AND DECLARATIONS 


Name of genus of 
which species 
specified in Col. | 
is the type species 


(3) 
Fistularia Linnaeus, 
1758 
Mugil Linnaeus, 1758 


Trachinus Linnaeus, 
1758 

Uranoscopus Linnaeus, 
1758 

Xiphias Linnaeus, 1758 


Acipenser Linnaeus, 
1758 

Callionymus Linnaeus, 
1758 

Chimaera Linnaeus, 
1758 

Clupea Linnaeus, 
1758 

Coryphaena Linnaeus, 
1758 

Cyclopterus Linnaeus, 
1758 


Cyprinus Linnaeus, 
i 
Gadus Linnaeus, 1758 


Gasterosteus Linnaeus, 
1758 

Gobius Linnaeus, 1758 

Lophius Linnaeus, 
1758 

Mormyrus Linnaeus, 
1758 

Mullus Linnaeus, 
1758 


Perca Linnaeus, 1758 


Salmo Linnaeus, 1758 


Scomber Linnaeus, 
1758 


DIRECTION 57 


Combination in which 
name specified in 
Col. 1 was originally 


Specific name 
proposed to be placed 
on the *‘ Official List 


of Specific Names in published 
Zoology” 
(1) (2) 


porcus Linnaeus, 1758 Scorpaena porcus 
glanis Linnaeus, 1758  Silurus glanis 


acus Linnaeus, 1758  Syngnathus acus 


faber Linnaeus, 1758 Zeus faber 


‘© Opinion *? 92 


ocellaris Linnaeus, Blennius ocellaris 


1758 
lucius Linnaeus, 1758 Esox lucius 


barbatum Linnaeus, Ophidion barbatum 
1758 
** Opinion ”’ 93 


lavaretus Linnaeus, Salmo lavaretus 


1758 
pisonis Gmelin, [1789] Gobius pisonis 


marginalis Bloch, Epinephelus 
marginalis 

reticularis Bloch, Gymnothorax 
1795 reticularis 

electricus Gmelin, Silurus electricus 
[1789] 

mustelus Linnaeus, Squalus mustelus 
1758 

eto Linnaeus, Polynemus paradiseus 
17 

cabrilla Linnaeus, Perca cabrilla 
1758 


381 


Name of genus of 
which species 
specified in Col. | 
is the type species 


(3) 
Scorpaena Linnaeus, 
1758 


Silurus Linnaeus, 
1758 


Syngnathus Linnaeus, 
1758 


Zeus Linnaeus, 1758 


Blennius Linnaeus, 
1758 


Esox Linnaeus, 1758 


Ophidion Linnaeus, 
1758 


Coregonus Linnaeus, 
1758 


Eleotris Schneider, 
1801 

Epinephelus Bloch, 
1793 

Gymnothorax Bloch, 


Malapterurus 
Lacépéde, 1803 
Mustelus Linck, 1790 


Polynemus Linnaeus, 
1758 
Serranus Cuvier, 1817 


Section C : Class Amphibia (two names) 


** Opinion ’’ 92 


fuscus Rafinesque, Triturus fuscus 


lacertina Linnaeus, Siren lacertina 


1766 


Desmognathus Baird, 
(1850) 
Siren Linnaeus, 1766 


382 OPINIONS AND. DECLARATIONS 


Specific name Combination in which Name of genus of 
proposed to be placed name specified in which species 
on the “* Official List — Col. 1 was originally specified in Col. 1 
of Specific Names in published is the type species 

Zoology ”’ 
(1) (2) (3) 


Section D : Class Reptilia (six names) 
** Opinion ’’ 75 
volans Linnaeus, 1758 Draco volans Draco Linnaeus, 1758 


** Opinion ”’ 92 


calamarius Linnaeus, Coluber calamarius Calamaria Boie, 1826 
1758 
serpentina Linnaeus, Testudo serpentina Chelydra Schweigger 
1758 1812 
horridus Linnaeus, Crotalus horridus Crotalus Linnaeus, 
1758 1758 
coriacea Linnaeus, Testudo coriacea Dermochelys 
1766 Blainville, 1816 
agilis Linnaeus, 1758 Lacerta agilis Lacerta Linnaeus, 
1758 
ANNEXE 2 


Proposed addition to the ‘‘ Official List of Specific Names in Zoology ”’ 
of the specific name of a species of the Class Amphibia which is 
currently treated as a senior subjective synonym of the name of a 
species which is the type species of a genus the name of which 
was placed on the ‘‘ Official List of Generic Names in 
Zoology ’’ in the period up to the close of 1936 


Specific name proposed to be Remarks 
placed on the “‘ Official List (not for inclusion in the entry to 
of Specific Names in Zoology ”’. be made in the Official List) 
(1) (2) 
alleganiensis Daudin, [1803], as a senior subjective synonym of 
published in the combination gigantea Barton, 1808, as pub- 
Salamandra alleganiensis. lished in the combination 


Salamandra gigantea, the specific 
name of the type species of 
Cryptobranchus Leuckart, 1821 
(which was placed on the Official 
List of Generic Names in Zoology 
by the Ruling given in Opinion 
92). 


DIRECTION 57 383 


2. Registration of the present application : Upon the receipt 
of Mr. Hemming’s paper the question of the addition to the 
Official List of Specific Names in Zoology of the specific names of 
the type species of genera belonging to the Classes Cyclostomata, 
Pisces, Amphibia and Reptilia, the names of which had been 
placed on the Official List of Generic Names in Zoology in the 
period up to the end of 1936 was allotted the Registered Number 
Z.NA(S.) 1014. 


Il. THE DECISION TAKEN BY THE INTERNATIONAL 
COMMISSION ON ZOOLOGICAL NOMENCLATURE 


3. Issue of Voting Paper V.P.(O.M.)(56)5 : On 25th April 
1956 a Voting Paper (V.P.(O.M.(56)5) was issued in which each 
Member of the Commission was asked (1) to state whether he 
agreed “that, in conformity with the General Directives issued 
to the International Commission on Zoological Nomenclature 
by the Thirteenth International Congress of Zoology, Paris, 1948, 
regarding the placing on the appropriate Official Lists and 
Official Indexes of names dealt with in, or otherwise involved in, 
Opinions rendered prior to 1948, the names of species in the 
Classes Cyclostomata, Pisces, Amphibia and Reptilia referred to 
in Sections (1) and (2) of paragraph 9 of the paper bearing the 
Registered Number Z.N.(S.) 1014 by the Secretary submitted 
simultaneously with the present Voting Paper [i.e. in the paragraph 
numbered as above in the paper reproduced in the first paragraph 
of the present Direction] be placed on the Official List of Specific 
Names in Zoology as there recommended and that the other action 
recommended in the said paragraph be similarly approved ”’, 
and (2), if he did not so agree as regards any given item, to 
indicate that item. 


384 OPINIONS AND DECLARATIONS 


4. Withdrawal from the scope of Voting Paper V.P.(O.M.)(56)5 
of the proposal that the specific name ‘‘ calamarius ’’ Linnaeus, 
1758, as published in the combination ‘‘ Coluber calamarius ’’, be 
placed on the ‘* Official List of Generic Names in Zoology ”’ to 
permit of the further study of the interpretation of the nominal 
species so named : On 2nd May 1956 Mr. Hemming, as Secretary, 
executed the following Minute withdrawing from the scope of. 
Voting Paper V.P.(O.M)(56)5 the proposal submitted therewith 
for the addition to the Official List of Specific Names in Zoology 
of the specific name calamarius Linnaeus, 1758, as published in 
the combination Coluber calamarius, the specific name of the 
type species of the genus Ca/amaria Boie, 1826 :— 


Withdrawal from the scope of Voting Paper V.P.(O.M.)(56)5 of the 
proposal that the specific name ‘‘ calamarius *’ Linnaeus, 1758, 
as published in the combination ‘ Coluber calamarius ”’ 
(Class Reptilia) be placed on the ‘* Official List of 
Specific Names in Zoology ”’ 


By FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


With their completed copies of Voting Paper V.P.(O.M.)(56)4 
received in the Office of the Commission today Dr. H. Boschma 
(Director, Rijksmuseum van Natuurlijke Historie, Leiden, The Nether- 
lands) and Dr. L. B. Holthuis of the same Museum reported that they 
had been informed by Dr. L. D. Brongersma of the above Museum 
that there were certain hitherto unsuspected difficulties in the matter 
of the interpretation of the nominal genus Calamaria Boie, 1826, a 
genus, the name of which had been placed on the Official List of 
Generic Names in Zoology by the Ruling given in Opinion 92 and on 
which certain supplementary proposals had been submitted in the 
paper bearing the Number Z.N.(S.) 1015 issued to the Members of the 
Commission concurrently with the Voting Paper referred to above. 
It appeared from the communications so received that the current 
interpretation of the nominal species Coluber calamarius Linnaeus, 
1758, the type species of the foregoing genus and/or of the objectively 
identical nominal species Calamaria linnaei Boie, 1826, was open to 
question and that there was at least a possibility that the species to 
which Linnaeus had given that name was a species at present placed by 
specialists in a genus far removed from Calamaria Boie. 


DIRECTION 57 385 


2. On the receipt of the communications referred to above I at once 
executed a Minute? numbered Z.N.(S.) 1015 in which, as Secretary, I 
withdrew certain proposals relating to the generic name Calamaria 
Boie from the scope of Voting Paper V.P.(O.M.)(56)4, in order to 
permit of the further study of the issues alluded to in the foregoing 
communications and directed that a new File bearing the Registered 
Number Z.N.(S.) 1114 be opened for the prosecution of the study 
of the issues so reserved for further examination. The same problem 
arises in connection with the proposals submitted to the Commission 
with Voting Paper V.P.(O.M.)(56)5, which are closely linked with those 
submitted with Voting Paper V.P.(O.M.)(56)4, being concerned with the 
addition to the Official List of Specific Names in Zoology of the specific 
names of the type species of genera belonging to the Classes 
Cyclostomata, Pisces, Amphibia and Reptilia, the names of which 
were placed on the Official List of Generic Names in Zoology during 
the period up to the end of 1936, the group of names in respect of 
which certain preliminary proposals of a miscellaneous character had 
been submitted with Voting Paper V.P.(O.M.)(56)4. In these cir- 
cumstances, as Secretary, I hereby withdraw from the scope of Voting 
Paper V.P.(O.M.)(56)5 the proposals in regard to the specific name 
calamarius Linnaeus, 1758, as published in the combination Coluber 
calamarius, submitted therewith and direct that whatever may be found 
to be the appropriate recommendation on this matter be submitted 
at the close of the investigations on File Z.N.(S.) 1114 opened for the 
corresponding investigations arising out of the withdrawal from the 
scope of Voting Paper V.P.(O.M.)(56)4 of the proposals submitted 
therewith in connection with the generic name Ca/amaria Boie. 


5. The Prescribed Voting Period for V.P.(O.M.)(56)5 : As 
Voting Paper V.P.(O.M.)(56)5 was issued under the One-Month 
Rule, the Prescribed Voting Period therefore closed on 25th 
May 1956. 


6. Particulars of the Voting on Voting Paper V.P.(O.M.(56)5 : 
At the close of the Prescribed Voting Period for Voting Paper 
V.P.(O.M.)(56)5, the state of the voting on the proposals sub- 
mitted therewith, other than on that part of those proposals 
which had been withdrawn from the scope of that Voting Paper 
by the Minute executed by the Secretary on 2nd May 1956 (the 


* The text of the Minute here referred to has been reproduced in paragraph 5 
of Direction 56 (published in the immediately preceding Part of the present 
Section of Volume | of the *‘ Opinions and Declarations ”’ Series). 


386 OPINIONS AND DECLARATIONS 


text of which has been reproduced in paragraph 4 of the present 
Direction) was as follows :— 


(a) Affirmative Votes had been given by the following twenty-four 
(24) Commissioners (arranged in the order in which Votes 
were received : 


Holthuis ; Boschma; Stoll; Vokes; Hering; Miller ; 
Prantl ; Bodenheimer; Cabrera; Esaki; Mayr; do 
Amaral; Lemche; Hanko; Dymond; Mertens; 
Key; Jaczewski; Bonnet; Hemming;  Sylvester- 
Bradley ; Riley ; Kuhnelt ; Tortonese ; : 


(b) Negative Votes : 


None ; 


(c) On Leave of Absence, one (1) : 


Bradley (J.C.) ; 


(d) Voting Papers net returned : 


None. 


7. Declaration of Result of Vote: On 7th June 1956, Mr. 
Hemming, Secretary to the International Commission, acting as 
Returning Officer for the Vote taken on Voting Paper V.P.(O.M.) 
(56)5, signed a Certificate that the Votes cast on that Voting Paper, 
other than on the proposals in regard to the name calamarius 
Linnaeus, 1758, as published in the combination Coluber 
calamarius, submitted therewith, which had been withdrawn 
from its scope by the Minute executed by the Secretary on 2nd 


DIRECTION 57 387 


May 1956 (the text of which has been reproduced in paragraph 
4 of the present Direction) were as set out in paragraph 6 above 
and that the proposals submitted with that Voting Paper, other 
than the proposals so withdrawn, had been duly adopted and 
that the decision so taken was the decision of the International 
Commission in the matter aforesaid. 


8. Preparation of the Ruling given in the present ‘‘ Direction ”’ : 
On Ist October 1956, Mr. Hemming prepared the Ruling given 
in the present Direction and at the same time signed a Certificate 
that the terms of that Ruling were in complete accord with those 
of the proposal approved by the International Commission in its 
vote on Voting Paper V.P.(O.M.)(56)5, subject to the exclusion 
therefrom of the proposals in regard to the specific name 
calamarius Linnaeus specified in the Minute executed by the 
Secretary on 2nd May 1956 referred to in paragraph 7 above. 


9. Original References : The original references for the specific 
names placed on the Official List of Specific Names in Zoology 
by the Ruling given in the present Direction have already been 
noted in connection with the Official List of Generic Names in 
Zoology, the nominal species bearing the names concerned being 
either (1) the type species of genera, the names of which have been 
inscribed on that List or (ii), in one case, a nominal species 
which is currently subjectively identified as representing the 
same taxon as that represented by a nominal species which is 
the type species of such a genus. 


10. The prescribed procedures were duly complied with by the 
International Commission on Zoological Nomenclature in 
dealing with the present case, and the present Direction is 
accordingly hereby rendered in the name of the said International 
Commission by the under-signed Francis Hemming, Secretary 
to the International Commission on Zoological Nomenclature, in 
virtue of all and every the powers conferred upon him in that 
behalf. 


388 OPINIONS AND DECLARATIONS 


11. The present Direction shall be known as Direction Fifty- 
Seven (57) of the International Commission on Zoological 
Nomenclature. 


Done in London, this First day of October, Nineteen Hundred 
and Fifty-Six. 


Secretary to the International Commission 
on Zoological Nomenclature 


FRANCIS HEMMING 


Note to Subscribers 


The next following ‘‘ Direction ’”’ (‘* Direction ’’ 58), which is being 
published simultaneously with the present ‘‘ Direction ’’, is being issued 
as Part 21 of Volume 13 of the present Series. 


Printed in England by MercaLtFe & Cooper LimITED, 10-24 Scrutton St., London EC 2. 


OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 
FRANCIS HEMMING, c..G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.19. 
(Concluding Part) 


te . 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1957 


Price Three Guineas 
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OPINIONS AND DECLARATIONS 
RENDERED BY THE INTER- 
NATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


Edited by 


FRANCIS HEMMING, C.M.G., C.B.E. 


Secretary to the Commission 


VOLUME 1. SECTION D. Part D.19. Pp. 389—470 
(also published with this Part : T.P—XIV) 


CONTENTS 


Supplementary Directions in regard to two names; 

Corrigenda ; Subject Index ; Particulars of the dates of 

publication of the several Parts in which the present 
volume was published; Instructions to Binders. 


Also published with this Part: Title Page, Foreword ; 
Table of Contents. 


LONDON : 


Printed by Order of the International Trust for 
Zoological Nomenclature 
and 
Sold on behalf of the International Commission on Zoological 
Nomenclature by the International Trust at its Publications Office 
41, Queen’s Gate, London, S.W.7 
1957 


Price Three Guineas 


(All rights reserved) 


Issued 10th October, 1957 


INTERNATIONAL COMMISSION ON 
ZOOLOGICAL NOMENCLATURE 


COMPOSITION AT THE TIME OF THE ADOPTION OF THE 
DIRECTIONS PUBLISHED IN THE PRESENT SECTION OF 
VOLUME 1 


A. The Officers of the Commission 


Honorary Life President: Dr. Karl JORDAN (British Museum (Natural History) 
Zoological Museum, Tring, Herts., England) 


President: Professor James Chester BRADLEY (Cornell University, Ithaca, N.Y., 
U.S.A.) (12th August 1953) 


Vice-President : Senhor Dr. Afranio do AMARAL (Sao Paulo, Brazil) (12th August 1953) 
Secretary : Mr. Francis HEMMING (London, England) (27th July 1948) 


B. The Members of the Commission 


(Arranged in order of precedence by reference to date of election or of most recent 
re-election, as prescribed by the International Congress of Zoology) 


Professor H. BoscHMA (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(ist January 1947) 

Senor Dr. Angel CapreraA (La Plata, Argentina) (27th July 1948) 

Mr. Francis HEMMING (London, England) (27th July 1948) (Secretary) 

Dr. Henning LEMCHE (Universitetets Zoologiske Museum, Copenhagen, Denmark) 
(27th July 1948) 

Professor Teiso EsAKkI (Kyushu University, Fukuoka, Japan) (17th April 1950) 

Professor Pierre BONNET (Université de Toulouse, France) (9th June 1950) 

Mr. Norman Denbigh RILEy (British Museum (Natural History), London) (9th June 1950) 

Professor Tadeusz JACZEwSKI (Institute of Zoology, Polish Academy of Sciences, Warsaw, 
Poland) (15th June 1950) 

Professor Robert MERTENS (Natur-Museum u. Forschungs-Institut Senckenberg, Frankfurt 
a.M., Germany) (5th July 1950) 

Professor Erich Martin HerING (Zoologisches Museum der Humboldt-Universitdt zu 
Berlin, Germany) (Sth July 1950) 

Senhor Dr. Afranio do AMARAL (S. Paulo, Brazil) (12th August 1953) (Vice-President) 

Professor J. R. DyMonpD (University of Toronto, Toronto, Canada) (12th August 1953) 

eer a: a BRADLEY (Cornell University, Ithaca, N. Y., U.S.A.) (12th August 1953) 
President 

Professor Harold E. Voxes (University of Tulane, Department of Geology, New Orleans, 

Louisiana, U.S.A.) (A2th August 1953) 

Professor Béla HANKO (Mezdégazdasadgi Muzeum, Budapest, Hungary) (12th August 1953) 

Dr. Norman R. STOLL (Rockefeller Institute for Medical Research, New York, N.Y., U.S.A.) 
(12th August 1953) 

Mr. P. C. SYLVESTER-BRADLEY (Sheffield University, Sheffield, England) (12th August 1953) 

Dr. L. B. Hottuuis (Rijksmuseum van Natuurlijke Historie, Leiden, The Netherlands) 
(12th August 1953) 

Dr. K. H. L. Key (Commonwealth Scientific and Industrial Research Organisation, 
Canberra, A.C.T., Australia) (15th October 1954) 

Dr. Alden H. MILLER (Museum of Vertebrate Zoology, University of California, U.S.A.) 
(29th October 1954) 

Doc. Dr. Ferdinand PRANTL (Ndrodni Museum V Praze, Prague, Czechoslovakia) (30th 
October 1954) 

Professor Dr. Wilhelm KUHNELT (Zoologisches Institut der Universitat, Vienna, Austria) 
(6th November 1954) 

Professor F. S. BODENHEIMER (The Hebrew University, Jerusalem, Israel) (11th November 
1954) 

Professor Ernst MAyrR (Museum of Comparative Zoology at Harvard College, Cambridge, 
Massachusetts, U.S.A.) (4th December 1954) 

Professor Enrico TORTONESE (Museo di Storia Naturale ‘‘ G. Doria,’’ Genova, Italy) 
(16th December 1954) 


SUPPLEMENT TO DIRECTION 56 


Instructions regarding the generic name ‘‘ Teuthys ”? Linck, 1790 (Class 
Pisces) and Supplementary Instructions relating to the generic name 
** Teuthis ’’ Schneider, 1784 (Class Cephalopoda) 


MINUTE by FRANCIS HEMMING, C.M.G., C.B.E. 


(Secretary to the International Commission on Zoological Nomenclature) 


In the course of the routine checking of entries made on the Official 
Lists and Official Indexes by Rulings given by the International Com- 
mission on Zoological Nomenclature during the year now closing my 
attention has been drawn to two minor matters affecting Direction 56, 
published on 20th of the current month (Ops. Decls. int. Comm. zool. 
Nomencl. 1(D) : 337—364), which call for supplementary action by the 
International Commission. 


2. First, I note that by an oversight the name TJeuthys Linck, 1790 
(Class Pisces) (Mag. f. Neueste aus der Physik u. Naturgesch. 6(3) : 32) 
(an Erroneous Subsequent Spelling for Teuthis Linnaeus, 1776, a name 
placed on the Official List of Generic Names in Zoology by the Ruling 
given in Opinion 93) was omitted from the list of invalid generic names 
enumerated in Section A of Annexe 2 of the paper bearing the Registered 
Number Z.N.(S.) 1015 there recommended for addition to the Official 
Index of Rejected and Invalid Generic Names in Zoology, which formed 
the basis on which the International Commission took the decisions later 
embodied in Direction 56. 


3. Second, I regret to find that the reputed generic name Teuthis 
Schneider, 1784 (Class Cephalopoda) which was placed on the foregoing 
Official Index as Name No. 813 by the Ruling given in the above Direction 
had already been entered on that Official Index as Name No. 58 by the 
Ruling given in Opinion 233 (1954, Ops. Decls. int. Comm. zool. Nomencl. 
4 : 275—296). 


4. In order to make good the omission specified in paragraph 2 above 
and to correct the error specified in paragraph 3 above, I now, as Secretary 
to the International Commission on Zoological Nomenclature, hereby 
direct as follows, namely :— 


(a) that the duplicate entry of Teuthis Schneider, 1784, as Name No. 813 

on the Official Index of Rejected and Invalid Generic Names in 
: Zoology made by the Ruling given in Direction 56 be cancelled 
\ and withdrawn ; 


Le | 


(b) that the name Teuthys Linck, 1790 (an Erroneous Subsequent 
Spelling for Teuthis Linnaeus, 1766), a name inadvertently 
omitted from the names placed on the Official Index of Rejected 
and Invalid Generic Names in Zoology by the Ruling given in 
Direction 56 be treated as having been entered on the said Official 
Index by the Ruling given in the foregoing Direction, with the 
Name No. 813 (a Name Number rendered vacant by the 
instruction specified in (a) above). 


(signed) FRANCIS HEMMING 


Secretary to the International Commission 
on Zoological Nomenclature 


31st December 1956 


page 6. 

page 8. 

page 11. 
page 17. 
page 20. 
page 23. 
page 24. 
page 31. 
page 32. 
page 34. 
page 36. 
page 38. 
page 40. 


page 57. 


page 77. 


page 159. 
page 175. 
page 214. 
page 246. 
page 249. 
page 340. 
page 371. 
page 374. 


Corrigenda 


Ruling (1)(24), lines 1 and 3 : delete square brackets from around “ 1869 ”’. 
Ruling (1)(40), lines 1 and 2 : substitute “ cylindrus”’ for ‘* cylindricus ” 
Ruling (1)(68), lines 1 and 3 : substitute ‘“‘ Zehntner ”’ for “‘ Zehnter ”’. 

Ruling (1)(121), lines 1 and 4: delete square brackets from around “ 1860 ”’. 
Ruling (1)(146), first line : substitute ‘“* [1828] ’’ for “‘ 1825 ”’. 

Ruling (2)(3), first line : substitute ‘‘ 1758” for “ 1767” 

Ruling (4)(9), first line : delete ““Adams &”’. 

Line 9 from end, columns 1 and 3 : delete square brackets from around “‘ 1869 ”’. 
Line 14 from end, columns | and 2: substitute “‘ cylindrus ”’ for “‘ cylindricus ”’. 
Line 6, column | and line 7, column 3 : substitute ‘‘ Zehntner ”’ for ‘‘ Zehnter ”’. 
Third line from end, column 3 : delete square brackets from around ‘“ 1860’. 
Line 19, column 1 : substitute “‘ [1828] ”’ for ‘“‘ 1825 ”’. 


Second entry, column |, first line and column 2, second line : substitute ** 1758 ” 
for “1767 ”’. 


Ruling (3)(k), entries (a) (b) and (c): insert ‘“‘ Milne Edwards &”’’ before 
= Emeas **. 


Section B, 2, first line : substitute “‘ pennanti”’ for ‘“* ponnanti”’. 

Ruling (3)(f)(@ii) : substitute “121” for “12”. 

Ruling (1)(20), first line : substitute “‘ novaehollandiae ” for ** novae-hollandiae ”’ 
Line 3 from end : substitute ‘‘ 1828” for “‘ 1838 ”’. 

Ruling (3)(b) : delete square brackets from around “‘ 1815’. 

Paragraph 5, lines 12 and 19: substitute “‘ XL” for “‘ XI”’. 

Ruling (2)(vii), second line : substitute “ 1864” for “‘ 1863’. 

Ruling (2)(b)(iii), lines 1 and 2 : substitute “‘ barbatum”’ for “* barbatus ” 
Ruling (4)(b)(i1), line 1 : substitute “‘ 1758” for ‘** 1858”. 


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Volume 1, Section D 


SUBJECT INDEX 


abdimii Lichtenstein, 1823, as published in the combination Ciconia abdimii (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 1008 


Acanthocyclus Dybowski, 1873 (a junior homonym of Acanthocyclus Milne Edwards 
(H.) & Lucas, [1844]), placed on the Official Index of Rejected and Invalid Generic 
Names in Zoology with Name No. 461 t; Ge a es ib rb 


Acanthopus Oken, 1816 (a junior homonym of Acanthopus Klug, 1807), placed on 
the Official Index of Rejected and Invalid Generic Names in Zoology with Name 
No. 462 2 Bh eS * ee *, * a on A 


Acanthopus Dahl, 1823 (a junior homonym of Acanthopus Klug, 1807), placed on 
the Official Index of Rejected and Invalid Generic Names in Zoology with Name 
No... 463" «> He at * ts ie Be Bs ae an or 


Acanthopus Latreille, 1829 (a junior homonym of Acanthopus Klug, 1807) placed 
on the Official Index of Rejected and Invalid Generic Names in Zoology with Name 
No. 464 , wa i y a the BY bs a. + 


Acanthopus de Haan, [1835] (a junior homonym of Acanthopus Klug, 1807, placed 
on the Official Index of Rejected and Invalid Generic Names in Zoology with Name 
No. 465 s4 fe Me =. a: ce a Be Es 3 


Acanthopus Muenster, 1839 (a junior homonym of Acanthopus Klug, 1807), placed 
on the Official Index of Rejected and Invalid Generic Names in Zoology with Name 
No. 466 .. Be ae ee ee wi oa a5 ie Age es 


Acanthopus Giebel, 1872 (a junior homonym of Acanthopus Klug, 1807), placed on 
the Official Index of Rejected and Invalid Generic Names in Zoology with Name 
No. 467 .. =e ok ee 5. a - oy a ae a, 


Acanthopus Vernet, 1877 (a junior homonym of Acanthopus Klug, 1807), placed on 
the Official Index of Rejected and Invalid Generic Names in Zoology with Name 
No. 468 .. we 2 me ae ek a ap Re ~ ae 


x 


Acipenser Linnaeus, 1758 (Class Pisces), correction of entry relating to type species 
of (correction of Opinion 77) a me fy: ite Lg Me oe 


Acipenses Linck, 1790 (an Erroneous Subsequent Spelling for Acipenser Linnaeus, 
1758), placed on the Official Index of Rejected and Invalid Generic Names in Zoology 
with Name No. 765 A ie ie i aM oe iM a ie 


395 


Page 
180 


a7 


57 


Si 


58 


58 


58 


58 


58 


339 


342 


396 Opinions and Declarations 


Acmaeopleura Stimpson, 1858 (Class Crustacea, Order sears 0 awsety of 
Ruling given in Opinion 85 : ae Lf i : ; a 


Actaea de Haan, [1833] eer Crustacea, Order kc bi: sie cian pou of one 
given in Opinion a3 ‘ 


Actumnus Dana, 1851 (Class Crustacea, Order Pecenoda): completion of Se 
given in Opinion 73 , ne oe js : i ee é : 


aculeatus Linnaeus, 1758, as published in the combination Gasterosteus aculeatus 
(Class Pisces), placed on the ee List ss ee Names in ea. with Name 
No. 1080 


acus Linnaeus, 1758, as published in the combination Syngnathus acus (Class Pisces), 
placed on the Official List of Specific Names in Zoology with Name No. 1090 : 


admete Herbst, [1803], as published in the combination Cancer admete (Class 
Crustacea, Order Decapoda), sain on the Bis ion List na sil: Names in 
Zoology with Name No. 834 .. 


Aegithia Blyth, 1860 (an Erroneous Subsequent Spelling for Aegithina Vieillot, 1816), 
placed on the Official Index of cae and Invalid Generic Names in n Zoology with 
Name No. 3527 =. 


aeneus Linnaeus, 1758, as published in the combination Cancer aeneus (Class 
Crustacea, Order Decapoda), Bires on the , ee List a Spe Names in 
Zoology with Name No. 931 .. 


aethereus Linnaeus, 1758, as published in the combination Phaethon aethereus (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 995 


Agasoma Gabb, [1869] (Class Gastropoda), placed on the Official List ue Generic 
Names in Zoology i in Opinion 121, determination of gender of ; 


agassizii Milne Edwards (A.), 1880, as published in the combination Eucratodes 
agassizii (Class Crustacea, Order Decapoda), Saar on the ae List “ be 
Names in Zoology with Name No. 856 


agilis Linnaeus, 1758, as published in the combination Lacerta agilis (Class Reptilia), 
placed on the Official List of Specific Names in Zoology with Name No. 1109 _ .. 


agilis Milne Edwards (A.), 1872, as published in the combination Hydrothelphusa 
agilis (Class Crustacea, Order Decapoda), rae on the mee List S ore 
Names in Zoology with Name No. 801 


Aix Agassiz (J.L.R.), 1846 (an Invalid Emendation of Aia Eyton, 1838, and a junior 
homonym of Aix Boie, 1828), placed on the eee Index Ke Rejected and Invalid 
Generic Names in Zoology with Name No. 528 : 


Alauda Linnaeus, 1758 (Class Aves), correction of reference for yo a pee’ of 
(correction of Opinion 67) 46 Ai he, he : : 


Page 
50 


53 


a 


370 


371 


11 


214 


22 


178 


159 


14 


374 


214 


213 


Volume 1, Section D 


Alauda Daudin, 1816 (a junior homonym of Alauda Linnaeus, 1758), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 529 


albicilla Linnaeus, 1758, as published in the combination Falco albicilla (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 1028 


alcocki Wood-Mason, 1891, as published in the combination Parilia alcocki (Class 
Crustacea, Order Decapoda), epee on the eae List ee i a Names in 
Zoology with Name No. 820 .. ‘ : 


alleganiensis Daudin, [1803], as published in the combination Salamandra 
alleganiensis (Class Amphibia), placed on the ies cia iCist een each Names in 
Zoology with Name No. 1110 .. a 


alleghaniensis Harlan, 1825, as published in the combination Abranchus alleghaniensis 
(an Invalid Emendation of alleganiensis Daudin, [1803], as published in the 
combination Salamandra alleganiensis), placed on the Official Index of Rejected 
and Invalid Specific Names in Zoology with Name No. 366 a ; 


Amblyomma Koch, 1844 (Class Arachnida), placed on the Sleaaes List ee Generic 
Names in Zoology i in Opinion 73, determination of gender of. . 


americana Saussure, 1857, as published in the combination Pseudothelphusa 


americana (Class ‘Crustacea, Order Decapoda), peers on the Official List of 


Specific Names in Zoology with Name No. 829 


americanus Linnaeus, 1758, as published in the combination Cuculus americanus 
(Class Aves), placed on the ce List aoy SPER Names in n Paie? with Name 
No. 974 .. 


amica Smith (S.I.), as published in the combination Hepatella amica (Class Crustacea, 
_ Decapoda), correction of date of, from 1870 to 1869 oneeren of cid 


placed on the Official List of Specific Names in Zoology with Name No. 797 


Ampelis Nozeman & Vosmaer, 1758 (a name published in a work rejected for nomen- 
clatorial purposes), placed on the Official Index of Rejected and Invalid Generic 
Names in Zoology with Name No. 518 : 


Ammodytes Bonaparte, 1831 (a junior homonym of Ammodytes Linnaeus, 1758), 
placed on the Official Index me ae and Invalid Generic Names in n Zoology with 
Name No. 766 .. 


Anarhicas Latreille, 1804 (an Erroneous Subsequent Spelling for Anarhichas Linnaeus, 
1758), placed on the Official Index ack mee a and Invalid Generic Names in Z Zoology 
with Name No. 767 oe 


Ancylostoma Dubini, 1843 (Class Nematoda), placed on the eta List ae Generic 
Names in Zoology in Opinion 66, determination of gender of 


andreossyi Audouin, [1826], as published in the combination Pilumnus andreossyi 
(Class Crustacea, ‘Order Decapoda), Blaceta: on the ances List Z Specie Names 
in Zoology with Name No. 786.. 


374 


347 


176 


86 


342 


343 


398 Opinions and Declarations 


angulatus Dana, 1851, as published in the combination Cyrtograpsus angulatus (Class 
Crustacea, Order Decapoda), ea on the is ei List ~ aes Names in 
Zoology with Name No. 851 .. 


ani Linnaeus, 1758, as published in the combination Crotophaga ani (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 977 i 


annulicornis Leach, [1815], as published in the combination Pandalus annulicornis 
(a junior objective synonym of montagui [Leach], [1814], as published in the 
combination Pandalus montagui), placed on the Official Index a Rejected and 
Invalid Specific Names in Zoology with Name No. 356 : - - 


Anodonta Lamarck, 1799 (Class Pelecypoda), placed on the sey List ae Generic 
Names in Zoology in Opinion 94, determination of gender of , 


anomalipes Miers, 1881, as published in the combination Thaumastoplax anomalipes 
(Class Crustacea, Order Decapoda), pees on the ote’ List ag Pies ie Names 
in Zoology with Name No. 924.. 


Anoplocephala Blanchard, 1848 (Class Cestoda), placed on the Off cial List eo Generic 
Names in Zoology in Opinion 77, determination of gender of : 


Antedon Fréminville, 1811 (Class Crinoidea), placed on the Cis List a Generic 
Names in Zoology in Opinion 73, determination of gender of : 


Anthocoris Fallén, 1814 (Class Insecta), placed on the Official List se Generic Names 
in Zoology in Opinion 104, determination of gender of 


Aptenodita Bonnaterre, 1791 (an Erroneous Subsequent Spelling for Aptenodytes 
Miller, 1778), placed on the Official Index “~ inert and Invalid Generic Names 
in Zoology with Name No. 530.. 


Aptenodites Lesson, 1837 (an Erroneous Subsequent Spelling for Aptenodytes Miller, 
1778), placed on the Official Index an ee and Invalid Generic Names in 
Zoology with Name No. 531 : 


Aptenodyta Gmelin (J.F.), 1788 (an Erroneous Subsequent Spelling for Aptenodytes 
Miller, 1778), placed on the Ts Index ge Race and Invalid Generic Names 
in Zoology with Name No. 532. 


Aptenodytes Forster (J.R.), 1777 (a nomen nudum), placed on the Official Index a 
Rejected and Invalid Generic Names in Zoology with Name No. 533 so 


aquaticus Linnaeus, 1758, as published in the combination Rallus aquaticus (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 1004 


aracari Linnaeus, 1758, as published in the combination Ramphastos aracari (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 1002 


Arcella Ehrenberg, [1830] (Class Rhizopoda), placed on the sear: List et Generic 
Names in Zoology in Opinion 77, determination of gender of 


Page 


13 


176 


246 


158 


at 


157 


158 


141 


2tS 


215 


215 


213 


173 


173 


156 


Volume 1, Section D 


ardeola Paykull, 1805, as published in the combination Dromas ardeola (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 979 : 


Argas Latreille, 1795 (Class Arachnida), placed on the Official List of Generic Names 
in Zoology in Opinion 73, determination of gender of fg aye ar iS 


Argonauta Linnaeus, 1758 (Class Cephalopoda), placed on the Official List of Generic 
Names in Zoology in Opinion 94, determination of gender of * se bet 


argus Temminck, 1807, as published in the combination Polyplectron argus (a junior 
objective synonym of bicalcaratus Linnaeus, 1758, as published in the combination 
Pavo bicalcaratus), placed on the Official Index of Rejected and Invalid Specific 
Names in Zoology with Name No. 343 cee e 6 a ae ae 


Armadillidium Brandt & Ratzeburg, [1831] (Class Crustacea), placed on the Official 
List of Generic Names in Zoology in Opinion 104, determination of gender of 


armata Milne Edwards (A.), 1869, as published in the combination Banareia armata 
(Class Crustacea, Order Decapoda), placed on the Official List of Specific Names 
in Zoology with Name No. 773.. ts as ore fs an a r: 


armatus Milne Edwards (H.), 1853, as published in the combination Potamocarcinus 
armatus (Class Crustacea, Order Decapoda), placed on the Official List of Specific 
Names in Zoology with Name No. 826 ee ae a oe me Ee 


arvensis Linnaeus, 1758, as published in the combination Alauda arvensis (Class Aves), 
placed on the Official List oy Specific Names in Zoology with Name No. 954 


Ascaris Linnaeus, 1758 (Class Nematoda), placed on the Official List of Generic 
Names in Zoology in Opinion 66, determination of gender of Se ni a 


Asthenognathus Stimpson, 1858 (Class Crustacea, Order Decapoda), completion of 
Ruling given in Opinion 85 ; oe a - Be ad ze Abe 


Asthenognathus Bocourt, 1884 (a junior homonym of Asthenognathus Stimpson, 
1858), placed on the Official Index of Rejected and Invalid Generic Names in 
Zoology with Name No. 469... ee 1 fe va ish Ae - 


Asthenognathus Broom, 1915 (a junior homonym of Asthenognathus Stimpson, 
1858), placed on the Official Index of Rejected and Invalid Generic Names in 
Zoology with Name No. 470... ah aa a zr in 7 ay 


Atergatis de Haan, [1833] (Class Crustacea, Order Decapoda), completion of Ruling 
given in Opinion 73 ne te ie 40 th ah ra Me é4 


Atergatopsis Milne Edwards (A.), 1862 (Class Crustacea, Order Decapoda), correc- 
tion of date of type species of (correction of Opinion 73) .. ae e ae 


Aulacorhynchus Dittmar, 1872 (a junior homonym of Aulacorhynchus Gould, 1834), 
placed on the Official Index of Rejected and Invalid Generic Names in Z oology with 
Name No. 534 .. ue s, sa st ae a ey, ay “ 


399 


Page 
176 


142 


156 


219 


50 


58 


58 


53 


35 


Ales, 


400 Opinions and Declarations 


aura Linnaeus, 1758, as published in the combination Vultur aura (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 966 


auratus Linnaeus, 1758, as published in the combination Cuculus auratus (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 976 


auritus Gray (J.E.), 1829, as published in the combination Podargus auritus (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 962 


australis Shaw, 1813, as published in the combination Apteryx australis (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 957 


Avicularia Lamarck, 1818 (Class Arachnida), placed on the Official List of Generic 
Names in Zoology in Opinion 104, determination of gender of As 


avosetta Linnaeus, 1758, as published in the combination Recurvirostra avosetta 
(Class Aves), placed on the Official List of Specific Names in Zoology with Name 
No. 1005 ; . * ee: as + nF an < Se, 


balansae Milne Edwards (A.), 1873, as published in the combination Clistocoeloma 
balansae (Class Crustacea, Order Decapoda), placed on the Official List of Specific 
Names in Zoology with Name No. 850 ae 4. - eh i *: 


barbatum Linnaeus, 1758, as published in the combination Ophidion barbatum (Class 
Pisces), placed on the Official List of Specific Names in Zoology with Name No. 1094 


harbatus Linnaeus, 1758, as published in the combination Mullus barbatus (Class 
Pisces), placed on the Official List of Specific Names in Zoology with Name No. 
1084 ch oe oy + ae ae oe! bes sa Ae 


barbiger Poeppig, 1836, as published in the combination Gecarcinus barbiger (Class 
Crustacea, Order Decapoda), placed on the Official List of Specific Names in 
Zoology with Name No. 942 _ .. a: Hs oe oF =e sh a 


Bathycrinus Thomson, 1872 (Class Crinoidea), placed on the Official List of Generic 
Names in Zoology in Opinion 73, determination of gender of SF a 


Bathynectes Stimpson (Class Crustacea, Order Decapoda), correction of date of, 
from 1871 to [1871], and of entry relating to type species of (correction of Opinion 


Bathynectes Guenther, 1878 (a junior homonym of Bathynectes Stimpson, [1871)]), 
placed on the Official Index of Rejected and Invalid Generic Names in Zoology with 
Name No. 471 26 ai: e ud va ia Ye oe za 


Bathynectes Brinkman, 1913 (a junior homonym of Bathynectes Stimpson, [1871)]), 
placed on the Official Index of Rejected and Invalid Generic Names in Zoology with 
Name No. 472 oe = ae i ee he a a ae 


Bathyplax Milne Edwards (A.), 1880 (Class Crustacea, Order Decapoda), com- 
pletion of Ruling given in Opinion 85 * is yi 2 


Page 
175 


176 


175 


174 


143 


179 


13 


37 


370 


2a 


156 


54 


58 


58 


50 


Volume 1, Section D 


bedoti Zehntner, 1894, as published in the combination Sphaerocarcinus bedoti 
(Class Crustacea, Order Decapoda), aii on the soma’ List = pace hg Names 
in Zoology with Name No. 832.. 


bella Stimpson, 1860, as published in the combination Xantho bellas(Class Crustacea, 
Order Sigg is ean on the aging List — sd Names in sit with 
Name No. 880 


Bellia Bate, 1851 (a junior homonym of Bellia Milne Edwards (H.), 1848), placed on 
the Official Index eh aka and Invalid Generic Names in Cee Se with Name 
No. 473 


Bellia Gray (J.E.), 1869 (a junior homonym of Bellia Milne Edwards (H.), 1848), 
placed on the Official Index nae sik ik and Invalid Generic Names in eo with 
Name No. 474 .. 


Bellia Tutt, 1902 (a junior homonym of Bellia Milne Edwards (H.), 1848), placed on 
the Official Index ee eas and Invalid Generic Names in Oey with Name 
No. 475 ... 


bicalcaratus Linnaeus, 1758 (Pavo) (Class Aves) (a nominal species objectively 
identical with Polyplectron argus Temminck, lad: Pe peel of rilesiir 
Temminck, 1807, under Declaration 21 


placed on the Official List of Specific Names in Zoology with Name No. 1022 


bicornis Gray (J.E.), 1831, as published in the combination Gomeza bicornis (Class 
Crustacea, Order Decapoda), cae on the Peek List ee ee Names in 
Zoology with Name No. 866... 


bidentatus Milne Edwards (A.), 1880, as published in the combination Xanthodes 
bidentatus (Class Crustacea, Order Decapoda), Be on the eee List ca epee 
Names in Zoology with Name No. 922 


bispinosa Kinahan, 1856, as published in the combination Litocheira bispinosa (Class 
Crustacea, Order Decapoda), pisea on the ee List ee) au Names in 
Zoology with Name No. 879... ; 


bispinosus Herbst, [1783], as published in the combination Cancer bispinosus (Class 
Crustacea, Order Decapoda), bipses io on the sue List iid pee Names in 
Zoology with Name No. 862... ; 


Blatta Linnaeus, 1758 (Class Insecta), placed on the Official List ae Generic Names in 
Zoology in Opinion 104, determination of gender of 


Blennius Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 92 .. 


Bolten (J.F.), ruled not to be accepted as the author of the Museum Boltenianum, 
1798. See Roding (P.F.). 


Botryilus Gaertner, 1774 (Class Urochorda), placed on the cneent List co Generic 
Names in Zoology i in Opinion 94, determination of gender of : 


16 


58 


58 


58 


182 


jl | 


16 


14 


143 


341 


156 


402 Opinions and Declarations 


Brotogerys Agassiz (J.L.R.), 1846 (an Invalid Emendation of Brotogeris Vigors, 
1825), placed on the Official Index of Rejected and Invalid Generic Names in 
Zoology with Name No. 535... A ae =. ¥ a aa if 


Buccinum Linnaeus, 1758 (Class Gastropoda), placed on the Official List of Generic 
Names in Zoology in Opinion 94, determination of gender of 4 a ee 


Bursaria Miiller, 1773 (Class Ciliata), placed on the Official List of Generic Names 
in Zoology in Opinion 104, determination of gender of ‘ ue a. bh 


cabrilla Linnaeus, 1758, as published in the combination Perca cabrilla (Class Pisces), 
placed on the Official List of Specific Names in Zoology with Name No. 1102... 


caeruleus Desfontaines, [1789], as published in the combination Falco caeruleus 
(Class Aves), placed on the Official List of Specific Names in Zoology with Name 
No. 1026 fe Be is 9 a ce a ae oF is 


Calamaria Boie (Class Reptilia), correction of date of, from 1827 to 1826 (correctio 
of Opinion 92) .. = ae ~ te ae ie ch at i, 


Calliongmus Linck, 1790 (an Erroneous Subsequent Spelling for Callionymus 
Linnaeus, 1758, placed on the Official Index of Rejected and Invalid Generic Names 
in Zoology with Name No. 768.. ¢ s a, si hie ‘S Et 


Callionimus Gouan, 1770 (an Erroneous Subsequent Spelling for Callionymus 
Linnaeus, 1758), placed on the Official Index of Rejected and Invalid Generic Names 
in Zoology with Name No. 769.. ae *: ~ sn ay ae oS 


Callionymus Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 


Calyptraea Lamarck, 1799 (Class Gastropoda), placed on the Official List of Generic 
Names in Zoology in Opinion 94, determination of gender of . aH i 


Camptandrium Stimpson, 1858 (Class Crustacea, Order Decapoda), completion of 
Ruling given in Opinion 85 re 5 oe Re! ¥ be ae Ma 


Camptoplax Miers, 1884 (Class Crustacea, Order Decapoda), completion of Ruling 
given in Opinion 85 ae = Le ne as x ee Ee ar 


Capella Frenzel, 1801 (a name published in synonymy without an independent 
description), placed on the Official Index of Rejected and Invalid Generic Names 
in Zoology with Name No. 522.. 236 Re “s ae ap is te 


Capella Keyserling & Blasius, 1840 (a junior objective synonym of Rupicapra 
Blainville, 1816), placed on the Official Index of Rejected and Invalid Generic 
Names in Zoology with Name No. 523 tr: i bee ay ae De: 


Caphyra Guérin-Méneville (Class Crustacea, Order Decapoda), correction of date 
of, from 1830 to 1832, and of entry relating to type species of (correction of 
Opinion 73) 4 Eo a 5 is = “i a 


Page 


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343 


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54 


Volume 1, Section D 


Caphyra Barrande, 1846 (a junior homonym of Caphyra Guérin-Méneville, 1832), 
placed on the Official Index esi sea and Invalid Generic Names in n Zoology with 
Name No. 476 . 


Capito Gistl, 1848 (a junior homonym of Capito Vieillot, 1816), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 536 


carinata Ortmann, 1894, as published in the combination Pseudopinnixa carinata 
(Class Crustacea, Order Decapoda), eee on the cc cee List oe ae Names 
in Zoology with Name No. 909... 


carinipes Heller, 1861, as published in the combination Zozymodes carinipes (Class 
Crustacea, Order Decapoda), ee on the hase List sie ae Names in 
Zoology with Name No. 932... ; 


carinipes de Man, 1889, as published in the combination Durckheimia carinipes 
(Class Crustacea, Order eee oe on the aaa List 8 plate Names 
in Zoology with Name No. 853. 


carolinensis Stimpson, 1859, as published in the combination Speocarcinus carolinensis 
(Class Crustacea, Order Decapoda), Bees on the Ree List xe spree Names 
in Zoology with Name No. 920.. 


carpio Linnaeus, 1758, as published in the combination Cyprinus carpio (Class 
Pisces), placed on the Official List of Specific Names in Zoology with Name No. 
1078 - i A e, oF a Me s 5 os AP 


Catharthes d’Orbigny, [1835] (an Erroneous Subsequent Spelling for Cathartes 
Illiger, 1811), placed on the ee Index a leiden and Invalid Generic Names 
in Zoology with Name No. 537. 


Catoptrus Milne Edwards (A.), 1870 ran Crustacea, Order rr abe saa 
of Ruling given in Opinion 85 . 


Centrocerus Fitzinger, 1843 (a junior homonym of Centrocercus Swainson, 1832), 
placed on the Official Index eee pera and Invalid Generic Names in n Zoology with 
Name No. 538 he : 


Cephalopterus Risso, 1810 (a junior homonym of Cephalopterus Geoffroy, 1809), 
placed on the Official Index age estat and Invalid Generic Names in erica with 
Name No. 539 : 


Cephalopterus Powrie, 1870 (a junior homonym of Cephalopterus Geoffroy, 1809), 
placed on the Official Index ies eer and Invalid Generic Names in eee with 
Name No. 540 .. 


cephalus Linnaeus, 1758, as published in the combination Mugil cephalus (Class 
Pisces), placed on the Official List of Specific Names in Zoology with Name No. 
1068 ae ae = af ed a a: «8 a we ore 


Cephenemyia Latreille, 1818 (Class Insecta), placed on the Official List oe Generic 
Names in Zoology in Opinion 106, determination of gender of 


403 


Page 


58 


215 


20 


22 


13 


21 


369 


215 


50 


215 


215 


215 


368 


143 


404 Opinions and Declarations 


Ceratoplax Stimpson, 1858 (Class Crustacea, Order Decapoda), completion of 
Ruling given in Opinion 85 ; $ cae ts a 7 oe ke 


Cerion Roding, 1798 (Class Gastropoda), placed on the Official List of Generic 
Names in Zoology in Opinion 119, determination of gender of 2. ot a 


cessacii Milne Edwards (A.), 1878, as published in the combination Epimelus cessacit 
(Class Crustacea, Order Decapoda), placed on the Official List of Specific Names 
in Zoology with Name No. 792.. 4 a aa re. - ee ie 


chalybata Forster, 1781 (Paradisea |sic]) (Class Aves) (a nominal species objectively 
identical with Manucodia chalybea Boddaert, 1783), type species of Manucodia 
Boddaert, 1783, under Declaration 21 Ae ae Me is es A 


placed on the Official List of Specific Names in Zoology with Name No. 1019 


chalybea Boddaert, 1783, as published in the combination Manucodia chalybea 
(a junior objective synonym of chalybata Forster, 1781, as published in the com- 


bination Paradisea [sic] chalybata), placed on the Official Index of Rejected and 


Invalid Specific Names im Zoology with Name No. 344 ay :@ 


chalybeata Stiles, 1916, as published in the combination Paradisea chalybeata (an 
Erroneous Subsequent Spelling for chalybata Forster, 1781, as published in the 
combination Paradisea [sic] chalybata), placed on the Official Index of Rejected 
and Invalid Specific Names in Zoology with Name No. 345 58 Ad 2 


Chasmagnathus de Haan, [1833] (Class Crustacea, Order Decapoda), completion of 
Ruling given in Opinion 85 255 an a im rr 2 As i 


Chasmocarcinus Rathbun, 1898 (Class Crustacea, Order Decapoda), completion of 
Ruling given in Opinion 85 be at + = + a ak a 


Chauna Loew, 1847 (a junior homonym of Chauna Illiger, 1811), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 541 


chavaria Linnaeus, 1766, as published in the combination Parra chavaria (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 970 


cheiragonus Tilesius, 1815, as published in the combination Cancer cheiragonus (Class 
Crustacea, Order Decapoda), placed on the Official List of Specific Names in 
Zoology with Name No. 939. *: sis a oe se ae ahs 


Cheyletus Latreille, 1796 (Class Arachnida), placed on the Official List of Generic 
Names in Zoology in Opinion 104, determination of gender of De a ee 


chilensis Milne Edwards (H.) & Lucas, [1844], as published in the combination 
Corystoides chilensis (Class Crustacea, Order Decapoda), placed on the Official 
List of Specific Names in Zoology with Name No. 783 o Ne ae on 


Chimaera Linnaeus, 1758 (Class Pisces), correction of entry relating to type species 
of (correction of Opinion 77) .. ay Ry of 3 ae ay a 


Page 


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219 


50 


49 


215 


175 


23 


142 


339 


Volume 1, Section D 


Chimaera Poli, 1791 (a junior homonym of Chimaera Linnaeus, 1758), placed on 
the Official Index af PapoKe and Invalid Generic Names in in Zoology with Name 
No. 770 ; 


Chimaera Ochsenheimer, 1808 (a junior homonym of Chimaera Linnaeus, 1758), 
placed on the Official Index of ccm: and Invalid Generic Names in n Zoology with 
Name No. 771 


Chimaera Hitchcock, 1858 (a junior homonym of Chimaera Linnaeus, 1758), placed 
on the Official Index Zt Release and Invalid Generic Names in mae with 
Name No. 772 


chimaera Rothschild, 1895, as published in the combination Uratelornis chimaera 
(Class Aves), placed on the Be cite List acd aes Names in nae see with Name 
No. 1015 


Chimaira Duméril, 1856 (an Erroneous Subsequent Spelling for Chimaera Linnaeus, 
1758), placed on the Official Index ee eee and Invalid Generic Names in 
Zoology with Name No. 773... 


Chlorodopsis Milne Edwards (A.), 1863 (a nomen nudum), placed on the Official 
Index of Rejected and Invalid Generic Names in Zoology with Name No. 477... 


Chlorodopsis Milne Edwards (A.), 1873 (Class Crustacea, Order Decapoda), 
completion of Ruling given in Opinion 73... ae = Pe 


Chorioptes Gervais & van Beneden, 1859 (Class Arachnida), placed on the Official 
List of Generic Names in Zoology in Opinion 104, determination of gender of 


chrysocephala Lewin, 1808, as published in the combination Meliphaga chrysocephala 
(Class Aves), placed on the oe List sid gees, Names in a with Name 
No. 1006 


Chrysolophus Swainson, 1837 (a junior homonym of Chrysolophus Gray (J.E.), 
[1833—1834)), placed on the Official Index of Beles and Invalid Generic Names 
in Zoology with Name No. 542.. As 


ciliatus Stimpson, 1858, as published in the combination Ceratoplax ciliatus (Class 
Crustacea, Order Decapoda), Se a on the een List ve Pigg igi Names in 
Zoology with Name No. 848... , 


Cimex Linnaeus, 1758 (Class Insecta), placed on the unde List oh Generic Names 
in Zoology in Opinion 81, determination of gender of ; 


cinctimanus White, as published in the combination Carpilius cinctimanus (Class 
Crustacea, Order Decapoda), correction of date of, from 1848 to [1849] (correction 
of Opinion 85) a : a Ae re ¥ we ne eo 


placed on the Official List of Specific Names in Zoology with Name No. 877 


cinerea Linnaeus, 1758, as published in the combination Ardea cinerea (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 959 ; 


405 


Page 


343 


343 


343 


181 


343 


58 


53 


142 


180 


216 


13 


141 


24 
16 


174 


406 Opinions and Declarations 


Clamator Blyth, 1852 (a junior homonym of Clamator Kaup, 1829), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 543 


Clausilia Draparnaud, [1805] (Class Gastropoda), placed on the Official List of 
Generic Names in Zoology in Opinion 119, determination of gender of sh 


Clavelina Savigny, 1816 (Class Urochorda), placed on the Official List of Generic 
Names in Zoology in Opinion 94, determination of gender of a ® ts 


Clistocoeloma Milne Edwards (A.), 1873 (Class Crustacea, Order Decapoda), com- 
pletion of Ruling given in Opinion 85 Ne 30: = ae we vs 


Clupea Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 


Coerebra Boie, 1826 (an Erroneous Subsequent Spelling for Coereba Vieillot, 1816), 
placed on the Official Index of Rejected and Invalid Generic Names in Zoology with 
Name No. 544 =e ape ae te au ihe a is “2 


Columba Linnaeus, 1758 (Class Aves), placed on the Official List of Generic Names 
in Zoology with Name No. 1015, with Columba oenas Linnaeus, 1758, as type 
species Ee a a “3 a we wy oe ive aa ar 


gender of name 


Columbella Lamarck, 1799 (Class Gastropoda), placed on the Official List of Generic 
Names in Zoology in Opinion 94, determination of gender of By ee - 


Columbus Linnaeus, 1758 (an Invalid Original Spelling for Colymbus Linnaeus, 
1758), placed on the Official Index of Rejected and Invalid Generic Names in 
Zoology with Name No. 520 .. ae of 4 Lv a = a3 


Columbus ({ Tunstall], 1771, (an Erroneous Subsequent Spelling for Columba Linnaeus, 
1758), placed on the Official Index of Rejected and Invalid Generic Names in 
Zoology with Name No. 519... Se nt. ste ak ie Je fe 


communis Bonnaterre, 1791, as published in the combination Coturnix communis (a 
junior objective synonym of coturnix Linnaeus, 1758, as published in the com- 
bination on Jetrao coturnix), placed on the Official Index of Rejected and Invalid 
Specific Names in Zoology with Name No. 346 Se ae 3 ae 


convexus Milne Edwards (A.), 1867, as published in the combination Ruppeliioides 
convexus (Class Crustacea, Order Decapoda), placed on the Official List of Specific 
Names in Zoology with Name No. 916 fi if =n: - eC) a 


coppingeri Miers, 1884, as published in the combination Camptoplax coppingeri 
(Class Crustacea, Order Decapoda), placed on the Official List of Specific Names 
in Zoology with Name No. 846.. Ay sek As as aS a oy 


corallinus Faxon, 1893, as published in the combination Trachycarcinus corailinus 
(Class Crustacea, Order Decapoda), placed on the Official List of Specific Names 
in Zoology with Name No. 837.. RG a ee ae Ss a Hy 


Page 
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158 


50 


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216 


86 
86 


157 


86 


86 


219 


20 


12 


11 


Volume 1, Section D 


Coregonus Lacépéde, 1803 (a junior homonym of Coregonus Linnaeus, 1758), placed 
on the Official Index es asia and Invalid Generic Names in n Zoology with Name 
No. 774 


Coregonus Jarocki, 1822 (a junior homonym of Coregonus Linnaeus, 1758), placed 
on the Official Index oe oe and Invalid Generic Names in n Zoology with Name 
No. 775 


coriacea Linnaeus, 1766, as published in the combination Testudo coriacea (Class 
Reptilia), placed ¢ on the gee List a Se eaeoles Names in Seen with Name No. 
1108 


Coriudo Fleming, 1822 (a junior objective synonym of Dermochelys Blainville, 1816), 
placed on the Official Index gal Bee ae and Invalid Generic Names in n Zoology with 
Name No. 776 


coronata Alcock & Anderson, 1894, as published in the combination Randallia 
coronata (Class Crustacea, Order Decapoda), pads on the ene List oo ee 
Names in Zoology with Name No. 821 


Coryhaena, attributed to Linnaeus, 1758, and placed on the Official List of Generic 


407 
Page 


343 


343 


374 


343 


10 


Names in Zoology by the Ruling given in Opinion 77, correction of, to Coryphaena 341-342 


attribution of, to Stiles, 1922, and addition of, to the Official Index of Rejected and 
Invalid Generic Names in Zoology with Name No. 777 as an Erroneous Sub- 
sequent Spelling oe oe ee a 7. uC : a 


Coryphaena Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 


Coryphaena Coues, 1889 (a nomen nudum), placed on the ees Index oy ae 
and Invalid Generic Names in Zoology with Name No. 778. 


Coturnix Bonnaterre, 1790 (Class ae: Tetrao coturnix Linnaeus, 1758, BPE species 
of, under Declaration 21 ? de ane a : : Ay: 


Coturnix Cuvier, 1800 (a nomen nudum), placed on the Oigah Index ney cies and 
Invalid Generic Names in Zoology with Name No. 545 


coturnix Linnaeus, 1758 (Tetrao) (Class Aves) (a nominal species objectively 
identical with Coturnix communis Bonnaterre, ee type seers of Coturnix 
Bonnaterre, 1790, under Declaration 21 . = m5 


placed on the Official List of Specific Names in Zoology with Name No. 1016 


crassimanus Dana, 1851, as published in the combination Eucrate crassimanus (Class 
Crustacea, Order Decapoda), pee on the anaes List Sia picasiin Names in 
Zoology with Name No. 857... 


crassipes Bell, 1855, as published in the combination Phiyxia crassipes (Class 
Crustacea, Order Decapoda), eee on the oma Dist yee pEeCe S Names in 
Zoology with Name No. 822 .. : 


343 


340 


343 


181 


216 


181 
182 


14 


10 


408 Opinions and Declarations 


crassum Dana, 1851, as published in the combination Samartium crassum (Class 
Crustacea, Order Decapoda), placed on the Official List of Specific Names in 
Zoology with Name No. 917... me ae ie fs i ae 4 


crenatus de Haan, [1835], as published in the combination Cancer (Eucrate) crenatus 
(Class Crustacea, Order Decapoda), placed on the Official List of Specific Names 
in Zoology with Name No. 855.. on Me A - x2 es ye 


crepitans Linnaeus, 1758, as published in the combination Psophia crepitans (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 1001 


cribrarius Lamarck, 1818, as published in the combination Portunus cribrarius (Class 
Crustacea, Order Decapoda), placed on the Official List of Specific Names in 
Zoology with Name No. 770 .. ne Mee =i age - is ba 


cristata Pallas, 1764, as published in the combination Columba cristata (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 1027 


cristatus Shaw in White, [August] 1790, as published in the combination Caprimulgus 
cristatus (Class Aves), placed on the Official List of Specific Names in Zoology with 
Name No. 1024 .. SH ay a a me Re he ys bp? 


Crotalus Linnaeus, 1758 (Class Reptilia), completion of Ruling given in Opinion 92 


Cryptobranchus Leuckart, 1821 (Class Amphibia), placed on the Official List of 
Generic Names in Zoology in Opinion 92, determination of gender of ; a 


cubensis Stimpson, 1860, as published in the combination Epilobocera cubensis 
(Class Crustacea, Order Decapoda), placed on the Official List of Specific Names 
in Zoology with Name No. 791.. 4. pe si 2 a He Sy 


cumingii Bell, 1855, as published in the combination Lithadia cumingii (Class 
Crustacea, Order Decapoda), placed on the Official List of Specific Names in 
Zoology with Name No. 807 : uf AY ei Pre am 14 ie 


cupulifer Latreille, 1825, as published in the combination Pilumnus cupulifer (Class 
Crustacea, Order Decapoda), placed on the Official List of Specific Names in 
Zoology with Name No. 906 ‘ iA By o. se Pe and 


Cyclodius Dana, 1851 (Class Crustacea, Order Decapoda), completion of Ruling 
given in Opinion 73 E a Ee se ™ - a we xe 


v 


Cyclopterus Linnaeus, 1758 (Class Pisces), correction of entry relating to type species 
of (correction of Opinion 77) .. uf ha ete or oe te ae 


Cyclopterus Marseul, 1872 (a junior homonym of Cyclopterus Linnaeus, 1758), 
placed on the Official Index of Rejected and Invalid Generic Names in Zoology with 
Name No. 779 : Ne 8 A = a2 - a a 


cylindricum Say, 1818, as published in the combination Pinnotheres cylindricum 
(Class Crustacea, Order Decapoda), placed on the Official List of Specific Names 
in Zoology with Name No. 901.. Ee a a ae % a. Le 


Page 


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179 


183 


341 


By 


19 
53 


339 
344 


19 


Volume 1, Section D 


cylindrus Fabricius, 1777, as published in the combination Cancer cylindrus (Class 
Crustacea, Order Decapoda), eecaet on the occlu List ae sl Names in 
Zoology with Name No. 804 _ .. 


Cymo de Haan, [1833] (Class Crustacea, Order in taba ee correction of date of 
type species of (correction of Opinion 73) on 


cyprinoides Linnaeus, 1758, as published in the combination Mormyrus cyprinoides 
ag Pisces), placed on the Pie List pis ee Names in bo with Name 
o. 1083 


Cyprinus Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77.. 


Cyrtograpsus Dana, 1851 eon Crustacea, Order Sane aber eee of Rue 
given in Opinion 85 


Cyrtograpsus Carruthers, 1867 (a junior homonym of Cyrtograpsus Dana, 1851), 
placed on the Official Index a gta and Invalid Generic Names in Zoology 
with Name No. 478 pie 


danae Milne Edwards (A.), 1869, as published in the combination Goniosoma danae 
(Class Crustacea, Order Decapoda), Deen on the pears List os ae Names 
in Zoology with Name No. 836.. 


Davainea Blanchard & Railliet, 1891 (Class Cestoda), placed on the ies List e 
Generic Names in Zoology i in Opinion 84, determination of gender of 


deplanatus Cano, 1889, as published in the combination Euryetisus deplanatus (Class 
Crustacea, Order Decapoda), Lapa on the ie Ses List - 5 nie Names in 
Zoology with Name No. 858... F : 


depressa Stimpson, [1871], as published in the combination Panoplax depressa 
(Class Crustacea, Order Decapoda), ees on the Bi satus List os ee Names 
in Zoology with Name No. 892.. 


Demodex Owen, 1843 (Class Arachnida), placed on the Official List at Generic 
Names in Zoology i in Opinion 104, determination of gender of ; 


Dendryphantes Koch, 1837 (Class Arachnida), placed on the Official List - Generic 
Names in Zoology in Opinion 104, determination of gender of : 


Dentalium Linnaeus, 1758 (Class Scaphopoda), placed on the aed ate List oe Generic 
Names in Zoology i in Opinion 94, determination of gender of 


dentata Bell, 1855, as published in the combination Nursilia dentata (Class Crustacea, 
Order Decapoda), pees on the ee East ie ee Names in Trees with 
Name No. 813... 


denticulatus Montagu, 1808, as published in the combination Cancer denticulatus 
(Class Crustacea, Order Decapoda), Sad on the el ee List n pes Names 
in Zoology with Name No. 823. 


Page 


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59 


De 


Py, 


14 


18 


142 


142 


10 


410 Opinions and Declarations 


Dermacentor Koch, 1844 (Class Arachnida), placed on the Official list ay Generic 
Names in Zoology i in Opinion 73, determination of gender of : 


Dermanyssus Dugés, 1834 (Class Arachnida), placed on the Official List of Generic 
Names in Zoology in Opinion 104, determination of gender of 


Dermatochelys Wagler, 1830 (a junior objective synonym of Dermochelys Blainville, 
1816), placed on the Official Index vA Beecie and Invalid Generic Names in 
Zoology with Name No. 780 t : 


Dermochelis Lesueur, 1829 (an Erroneous Subsequent Spelling for Dermochelys, 
Blainville, 1816), placed on the Official Index of adc and Invalid Generic 
Names in "Zoology with Name No. 781 a 


Dermochelys Blainville, 1816 poigeae a dah pomEleee of Rue eet in CRE 
92) : 


Desmognathus Baird (Class Amphibia), correction of date of, from 1849 to (1850) 
(correction of Opinion 92) and completion of Ruhue a in that Opinion for 
type species of oe oe o i : ' ve ae a 


Diaptomus Westwood, 1836 (Class Crustacea), placed on the Official List ot Generic 
Names in Zoology in Opinion 104, determination of gender of 


Diazona Savigny, 1816 (Class Urochorda), placed on the Official List a Generic 
Names in Zoology in Opinion 94, determination of gender of : 


Dicrocoelium Dujardin, [1844] (Class Trematoda), placed on the Official List a 
Generic Names in Zoology in Opinion 84, determination of gender of ; 


dilatipes Adams & White, [1849], as published in the combination Pilumnus dilatipes 
(Class Crustacea, Order ae abel on the ihe List pains Names 
in Zoology with Name No. 881. 


Diodon O’ Reilly, 1818 (a junior homonym of Diodon Linnaeus, 1758), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 782 


Diodon Lesson, 1828 (a junior homonym of Diodon Linnaeus, 1758), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 783 


Diodon Lesson, 1830 (a junior homonym of Diodon Linnaeus, 1758), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 784 


Dipylidium Leuckart, 1863 (Class Cestoda), placed on the Lila List ee Generic 
Names in Zoology in Opinion 84, determination of gender of. . 


Dissodactylus Smith (S.1.), 1870 (Class Crustacea, Order 1 Deen comple of 
Ruling given in Opinion 85 as : 


Distaplia Della Valle, 1881 (Class Urochorda), placed on the SS List o Generic 
Names in Zoology in Opinion 94, determination of gender of.. 


Page 
142 


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344 


342 


341 


142 


158 


159 


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344 


344 


344 


159 


50 


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Volume 1, Section D 


doliatus Linnaeus, 1764, as published in the combination Lanius doliatus (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 1014 


DOLICHOPIDAE Stephens (J.F.), 1829 (an Erroneous Subsequent Spelling for 
DOLICHOPODIDAE, placed on the Official Index of Rejected and Invalid Pans 
Group Names in Zoology with Name No. 87. ie se ie: o , 


DOLICHOPODES Latreille, 1809 (an Invalid Original Spelling for DOLICHOPODIDAE), 
placed on the Official Index of he pp and ia a ae Names in 
Zoology with Name No. 85 


DOLICHOPODIDAE (correction of DOLICHOPODES) Latreille, 1809 (Class Insecta, 
Order Diptera), placed on the Official List of Family- Group Names in FD with 
Name No. 98, with Dolichopus Latreille, 1796, as type genus 


Dolichopus Latreille, 1796 (Class Insecta, Order Diptera), placed on the Official List 
of Generic Names in sia t with Name No. 1035, with Musca pemsaae as OE 
species é 


gender of name 


Dolichopus Van der Hoeven, [1856] (a junior homonym of Dolichopus Latreille, 
1796), placed on the Official Index ES las and Invalid Generic Names in 
Zoology with Name No. 752... 


DOLYCHOPODAE Samouelle, 1819 (an Erroneous Subsequent Spelling for DOLICHOPO- 
DIDAE), placed on the Official Index of Baeiee and Invalid ey: ee Names in 
Zoology with Name No. 86 a 


Dolychopus Samouelle, 1819 (an Erroneous Subsequent Spelling for Dolichopus 
Latreille, 1796), placed on the Gian Index eh Heseied and Invalid Generic Names 
in Zoology with Name No. 753. 


dovii Stimpson, 1860, as published in the combination Euphylax dovii (Class Crus- 
tacea, Order Decapoda), es on the ne dais List o Spe Names in ie 
with Name No. 795 Hee 


Draco Gouan, 1770 (a junior homonym of Draco Linnaeus, 1758), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 785 


Draco Oken, 1816 (a junior homonym of Draco Linnaeus, 1758), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 786 


draco Linnaeus, 1758, as published in the combination Trachinus draco (Class 
Pisces), placed on the Official List of Specific Names in Zoology with Name No. 
1069 a: ae of hs ee 4. a ate 33 ae en 


Dracunculus Reichard, 1759 (Class Nematoda), placed on the ae List ee Generic 
Names in Zoology in Opinion 66, determination of gender of F 


Durckheimia de Man, 1889 (Class Crustacea, Order corn eat a a a of 
Ruling given in Opinion 85 


411 


Page 


180 


269 


268 


268 


267 
267 


268 


269 


268 


344 


344 


368 


155 


49 


412 Opinions and Declarations 


Dysdera Latreille, [1804] (Class Arachnida), placed on the Official List cag Generic 
Names in Zoology i in Opinion 104, determination of gender of 


Ebalia Leach, [1817] Socios Crustacea, Order Drape, ne of ae 
given in Opinion 5) Cf 


Echinococcus Rudolphi, 1801 (Class Cestoda), placed on the aiciasd List of Generic 
Names in Zoology in Opinion 84, determination of gender of ; 


Ectobius Stephens, 1835 (Class Insecta), placed on the ee List se Generic Names 
in Zoology in Opinion 104, determination of gender of 


Eimeria Schneider, 1875 (Class Sporozoa), placed on the Official List ae Generic 
Names in Zoology i in Opinion 104, determination of gender of 


electricus Gmelin (J.F.), [1789], as published in the combination Silurus electricus 
(Class Pisces), 2 ate on the phe List eon phic Names in aes with Name 
No. 1099 


elegans Gmelin (J.F.), 1788, as published in the combination Psittacus elegans (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 1033 


Eleotris Gronovius, 1763 (a name published in a work rejected for nomenclatorial 
purposes), placed on the Official Index gies a tak and Invalid Generic Names in 
Zoology with Name No. 787 3 


Eleotris Schneider, 1801 (Class Pisces), correction of entry on Official List of Generic 
Names in Zoology relating to type species of (correction of Opinion 93) .. me 


Enchytraeus Henle, 1837 (Class Chaetopoda), placed on the Official List oe Generic 
Names in Zoology i in Opinion 104, determination of gender of 


Epixanthus Heller, 1861 aa Crustacea, Order eatecapeda COmR eee of Peon 
given in Opinion 85 : : 


Eremias Wiegmann, 1834 (Class Reptilia), placed on the FS desist East le Generic 
Names in Zoology in Opinion 92, determination of gender of 


eremita Nobili, 1906, as published in the combination Dacryopilumnus eremita 
(Class Crustacea, Order eer ate? | ede on the Se List bf oe Names 
in Zoology with Name No. 787. 


erinaceus Fabricius, 1787, as published in the combination Cancer erinaceus (Class 
Crustacea, Order Decapoda), bigced.£ on the Pi List oh Species Names in 
Zoology with Name No: 768"... : hes 


erithacus Linnaeus, 1758, as published in the combination Psittacus erithacus (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 1000 


Page 
143 


a2 


155 


141 


156 


372 


184 


344 


342 


155 


50 


S12 


179 


Volume 1, Section D 


erosa Miers, [1877], as published in the combination Actaeomorpha erosa (Class 
Crustacea, Order Decapoda), pein on the kia List sie eveniens Names in 
Zoology with Name No. 766... 2 


Esox Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 92 


euagora de Man, 1895, as published in the combination Parapanope euagora (Class 
Crustacea Order Decapoda), ples on the gree List oct eee) Names in 
Zoology with Name No. 894. 


Euchirograpsus Milne Edwards (H.), 1853 Soa ck Crustacea, Order pa ge com- 
pletion of Ruling given in Opinion 85 


Eucrate Rafinesque, 1815 (a nomen nudum), placed on the Official Index ie Re re 
and Invalid Generic Names in Zoology with Name No. 479 


Eucrate de Haan, [1835] eee ge nen Order ae mooie of Pula 
given in Opinion 85 : 


Eucratodes Milne Edwards (A.), 1880 Ce aioe Crustacea, Order ep. com- 
pletion of Ruling given in Opinion 85. 


Eucratopsis Smith (S.1.), 1869 (Class po chai Order Ee hetara he of 
Ruling given in Opinion 85 fe 


eudactylus Bell, 1855, as published in the combination Myrodes eudactylus (Class 
Crustacea, Order Decapoda), eae on the mee List nee species! Names in 
Zoology with Name No. 810... ; 


europaea Linnaeus, 1758, as published in the combination Sitta europaea (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 1007 


Euryetisus Cano, 1889 ees Crustacea, Order toes age of Weare 
given in Opinion 85 ae ret ‘ ‘ ; 


Eurylaemus Gloger, 1841 (an Erroneous Subsequent Spelling for Eurylaimus Hors- 
field, 1821), placed on the Official Index Hs aes and Invalid Generic Names in 
Zoology with Name No. 548... 


Eurynorhinchus Lafresnaye, 1842 (an Erroneous Subsequent Spelling for 
Eurynorhynchus Nilsson, 1821), placed on the Official Index es ta igh and 
Invalid Generic Names in Zoology with Name No. 546 ee ; 


Eurynorhynchus Nilsson, 1821 (Class ao Platalea Pee Linnaeus, 1758, HES 
species of, under Declaration 21 


Eurynorynchus Reichenbach, 1852 (an Erroneous Subsequent Spelling for 
Eurynorhynchus Nilsson, 1821), placed on the Official Index ee er and 
Invalid Generic Names in Zoology with Name No. 547 es 


Euryplax Stimpson, 1859 Seis Cee Order De aa conmpiruong of ee 
given in Opinion 85 


50 


59 


50 


50 


50 


180 


49 


216 


216 


181 


216 


50 


414 Opinions and Declarations 


Euryplax Klapalek, 1909 (a junior homonym of Euryplax Stimpson, 1859), placed 
on the Official Index ain bad and Invalid Generic Names in aes with Name 
No. 480 ‘ : 


Eurypygia Swainson, 1837 (an Erroneous Subsequent Spelling for Eurypyga Illiger, 
1811), placed on ‘the Official Index a = claus and Invalid Generic Names in 
Zoology with Name No. 549... 


Eurytium Stimpson, 1859 es Crustacea, Order saownimies: pommlenente of ee 
given in Opinion 85 3 


exaratus Milne Edwards (H.), 1834, as published in the combination Chlorodius 
exaratus (Class Crustacea, Order Decapoda), Daa on the Sail List - Spee 
Names in Zoology with Name No. 874 


excelsus Heller, 1862, as published in the combination Perigrapsus exceisus (Class 
Crustacea, Order Decapoda), peeery on the ae List As ake Names in 
Zoology with Name No. 897... 


excubitor Linnaeus, 1758, as published in the combination Lanius excubitor (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 988 


exulans Linnaeus, 1758, as published in the combination Diomedea exulans (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 978 


faber Linnaeus, 1758, as published in the combination Zeus faber (Class Pisces), 
placed on the Official List of Specific Names in Zoology with Name No. 1091 


Fabia Dana, 1851 (Class Crustacea, Order prom a eames of vena given 
in Opinion 85 : 3 


fallax Henderson, 1893, as published in the combination Ebalia fallax (Class 
Crustacea, Order Decapoda), Oe on the peer List oo cme Names in 
Zoology with Name No. 798... , Be 


Fasciola Linnaeus, 1758 (Class Trematoda), placed on the Hane List ah Generic 
Names in Zoology i in Opinion 84, determination of gender of : 


Filaria Miller, 1787 (Class Nematoda), placed on the Official List Zh Generic Names 
in Zoology in Opinion 104, determination of gender of 


Fistularia Forskal, 1775 (a junior homonym of Fistu/aria Linnaeus, 1758), placed 
on the Official Index pak es and Invalid Generic Names in n Zoology with Name 
No. 788 .. 


Fistularia Miller (O.F.), 1776 (a junior homonym of Fistularia Linnaeus, 1758), 
placed on the Official Index ia Reteatnd and Invalid Generic Names in ZOE, 
with Name No. 789 q: 


Fistularia Lamarck, 1816 (a junior homonym of Fistularia Linnaeus, 1758), placed 
on the Official Index we Releoted and Invalid Generic Names in Pekan: with 
Name No. 790 .. 


Page 


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49 


16 


18 


177 


176 


Sv 


157 


344 


344 


344 


Volume 1, Section D 


Fistularia Blainville, 1830 (a junior homonym of Fistularia Linnaeus, 1758), placed 
oe the adaere Index a Rice and ae Generic Names in ne Gomes with 
ame No os P 


Fistularia Bowerbank, 1841 (a junior homonym of Fistularia Linnaeus, 1758), 
placed on the Official Index s acike and Invalid Generic Names in Zoology 
with Name No. 792... 


flaveola Linnaeus, 1758, as published in the combination Certhia flaveola (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 975 


fluvialis Savigny, 1809, as published in the combination Pandion fluvialis (a junior 
objective synonym of haliaetus Linnaeus, 1758, as published in the combination 
Falco haliaetus), placed on the Official Index of Reece and Invalid Rees 
Names in Zoology with Name No. 347 ‘ 


fluviatilis Linnaeus, 1758, as published in the combination Perca fluviatilis (Class 
Pisces), placed on the Official 1 List of Bae Names in Bee ase with Name No. 
1085 fe 


fluviatilis Latreille, as published in the combination Trichodactylus fluviatilis (Class 
Crustacea, Order Pecenoee): correction of date of, from 1825 to ee en ection 
of Opinion 73) F as i. sy f oe 


placed on the Official List of Specific Names in Zoology with Name No. 839 


Forficula Linnaeus, 1758 (Class Insecta), placed on the Official List ee Generic 
Names in Zoology i in Opinion 104, determination of gender of , 


formosus Latham, 1790, as published in the combination Psittacus formosus (a 
junior primary homonym of formosus Scopoli, 1769, as published in the combina- 
tion Psittacus formosus), placed on the as Index oe eee and Invalid 
Names in Zoology with Name No. 348 é ~ 


fragifer de Haan, [1835], as published in the combination Cancer (Halimede) fragifer 
(Class Crustacea, Order age od Bee on the yale at EISt "3 ee Names 
in Zoology with Name No. 867. 


frontalis Milne Edwards (H.), 1834, as published in the combination Ozius frontalis 
(Class Crustacea, Order Decapoda), Bae on the Onett List of pregnG Names 
in Zoology with Name No. 940.. 


fulica Boddaert, 1783 (Colymbus) (Class Aves) (a nominal species objectively 
identical with Heliornis fulicarius Bonnaterre, NOR type species of Heliornis 
Bonnaterre, 1790, under Declaration 21 ; : F » , Ab 


placed on the Official List of Specific Names in Zoology with Name No. 1018 


fulicarius Bonnaterre, 1790, as published in the combination Heliornis fulicarius (a 
junior objective synonym of fulica Boddaert, 1783, as published in the combination 
Colymbus fulica), placed on the Official Index of patie and Invalid EAP ecane 
Names in Zoology with Name No. 349 


Fulmarus Stephens, 1826 (Class Aves), correction of reference for nee ae of 
(correction of Opinion 67) es ee a3 - ; : 


415 


Page 


344 


345 


176 


Pale 


370 


24 
12 


143 


219 


15 


23. 


181 
182 


220 


213 


416 Opinions and Declarations 


fuscus Rafinesque, 1820, as published in the combination Triturus fuscus (Class 
Amphibia), ana on the eas oe List Ses sai baa Names in ee with Name 
No. 1103 : 


Gadus Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 


Gadus Dejean, 1821 (a junior homonym of Gadus Linnaeus, 1758), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 793 


Gadus Rang, 1829 (a junior homonym of Gadus Linnaeus, 1758), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 794 


Gadus Deshayes, 1861 (a junior homonym of Gadus Linnaeus, 1758), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 795 


galbula Linnaeus, 1766, as published in the combination Oriolus galbula (a junior 
objective synonym of oriolus Linnaeus, 1758, as published in the combination 
Coracias oriolus), placed on the Offi cial Index of Re ae and Invalid es 
Names in Zoology with Name No. 350 : 


Galene de Haan, [1833] Goa: Crustacea, Order  Desaheda), comple of eae 
given in Opinion 85 ; , 


Gallinago Brisson, 1760 (Class Aves), substitution of, on the eile: ist s Generic 
Names in Zoology i in place of Gallinago Koch, 1816 ae 


Gallinago Koch, 1816, removal of, from the Official List of Generic Names in 
Zoology, on which placed by Opinion 67, and insertion of, on the Official Index 
of Rejected and Invalid Generic Names in Zoology as a junior homonym of, and a 
junior objective synonym of, Gallinago Brisson, 1760, with Name No. 521 


gallinago Linnaeus, 1758, as published in the combination Scolopax gallinago (Class 
Aves), placed on the Official List ef Specific Names in Zoology with Name No. 945 


Gammarus Fabricius, 1775 (Class Crustacea), placed on the Official List a Generic 
Names in Zoology in Opinion 104, determination of gender of : 


gammarus Linnaeus, 1758, as published in the combination Cancer gammarus (Class 
Crustacea, Order Decapoda), placeec on the gene List Bl Speen Names in 
Zoology with Name No. 1059 .. 


Gampsonyx Jordan, 1847 (a junior homonym of Gampsonyx Vigors, 1825), placed 
on the Official Index a vis pleas and Invalid Generic Names in pee with Name 
No. 550 ; ; 


garzetta Linnaeus, 1766, as published in the combination Ardea garzetta (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 980 


Gasterophilus Leach, 1817 (Class Insecta), placed on the Official List ae Generic 
Names in Zoology in Opinion 106, determination of gender of 


Gasterosteus Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 


Page 


373 


340 


345 


345 


345 


220 


50 


OF 


97 


oF 


142 


295 


216 


177 


141 


340 


Volume 1, Section D 


Gastrodiscus Cobbold, 1877 (Class Trematoda), placed on the ioe List ih Generic 
Names in Zoology in Opinion 84, determination of gender of : 


gaudichaudii Milne Edwards (H.), 1837, as published in the combination Platymera 
gaudichaudii (Class Crustacea, Order Decapoda), Bere on the ssi List He 
Specific Names in Zoology with Name No. 824 : 


gayi Milne Edwards (H.) & Lucas, [1844], as published in the combination 
Acanthocyclus gayi (Class Crustacea, Order Decapoda), re on the pa 
List of Specific Names in Zoology with Name No. 765 F : 


Gecarcinicus Dana, 1852 (an Erroneous Subsequent Spelling for Gecarcinucus Milne 
Edwards (H.), 1844), placed on the A ae Index a ee and Invalid Generic 
Names in Zoology with Name No. 481 


Geryon Kreyer, 1837 es Crustacea, Order peer Oe) has wena of f Rue 
given in Opinion 85 , 4 


gigas Lamarck, 1818, as published in the combination Cancer gigas (Class Crustacea, 
Order Decapoda), See on the Pe isi List me aa Names in siege with 
Name No. 908 _ .. : 


glaber Stimpson, 1858, as published in the combination Ptychognathus glaber (Class 
Crustacea, Order Decapoda), Sears on the Ags List a Se Names in 
Zoology with Name No. 912... a 


glacialis Linnaeus, 1761, as published in the combination Procellaria glacialis (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 983 


gladius Linnaeus, 1758, as published in the combination Xiphias gladius (Class 
Pisces), placed on the Official List of Specific Names in Zoology with Name No. 
1071 Me ey: a ee = nfs ms As a ce ae 


glandarius Linnaeus, 1758, as published in the combination Cuculus glandarius 
(Class Aves), placed on the Open List en See? Names in aoe with Name 
No. 973... 


glanis Linnaeus, 1758, as published in the combination Silurus glanis (Class Pisces), 
placed on the Official List of Specific Names in Zoology with Name No. 1089 _ . 


glutinosa Linnaeus, 1758, as published in the combination Myxine glutinosa (Class 
Cyclostomata), placed on the peat List of ea a Names in ie with Name 
No. 1063 


Glycyphagus Hering, 1838 (Class Arachnida), placed on the Official List oy Generic 
Names in Zoology in Opinion 104, determination of gender of ‘ 


Glyptograpsus Smith (S.1.), 1870 oat Crustacea, Order ie pee aii 
of Ruling given in Opinion 85 .. 


Glyptoplax Smith (S.1.), 1870 (Class Crustacea, Order arias a of 
Ruling given in Opinion 85 ie : ; 


a9 


49 


20 


20 


Wie 


369 


176 


i 


367 


142 


49 


50 


418 Opinions and Declarations 


Gnathostoma Owen, 1837 (Class Nematoda), placed on the Official List ee Generic 
Names in Zoology in Opinion 66, determination of gender of 


Gobius Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 


Gomeza Gray (J.E.), 1831 eects Crustacea, Order aii Sau aetna sa of Beat: 
given in Opinion 85 : 


Goneplat [Leach], [1814 ](an Invalid Original Spelling for Goneplax [Leach], [1814}), 
placed on the Official Index o, bay ere and Invalid Generic Names in a 
with Name No. 482 wv. 


Goneplax [Leach], [1814] as Crustacea, Order ais tae, bites as of ca, 
given in Opinion 85 : 


Gonoplax Leach, 1815 (an Invalid Emendation of Goneplax [Leach], [1814]), placed 
on the Official Index ay sac and Invalid Generic Names in Sfesihs ce: with Name 
No. 483... 


Gordius Linnaeus, 1758 (Class Nematomorpha), placed on the tor List ue Generic 
Names in Zoology i in Opinion 66, determination of gender of 


gracilipes White, 1847, as published in the combination Utica gracilipes (Class 
Crustacea, Order Decapoda), pees on the , List 4 anes Names in 
Zoology with Name No. 925... : . e 


gracilipes Stimpson, 1858, as published in the combination Rhizopa gracilipes (Class 
Crustacea, Order Decapoda), placed & on the pce List Ee Sree Names in 
Zoology with Name No. 915... 


gracilipes Alcock, 1894, as published in the combination Platypilumnus gracilipes 
(Class Crustacea, Order Decapoda), are on the peat List el etc Names 
in Zoology with Name No. 904.. 


granulatus de Haan, [1835], as published in the combination Cancer (Actaea) 
granulatus (a junior homonym of granulatus Linnaeus, 1758, as published in the 
combination Cancer granulatus), placed on the Official Index eb Rejected and 
Invalid Specific Names in Zoology with Name No. 336 E 23 


granulatus Rathbun, 1893, as published in the combination Oediplax granulatus 
(Class Crustacea, Order Decapoda), Des on the ikea I Last ot, SRC Names 
in Zoology with Name No. 888 


granulatus Rathbun, 1893, as published in the combination Scleroplax granulatus 


(Class Crustacea, Order Decapoda), ulin on the eicial List ud sais Names 
in Zoology with Name No. 919. 


granulosus Milne Edwards (A.), 1879, as published in the combination Pyxidognathus 
granulosus (Class Crustacea, Order Decapoda), bleeet on the OS List of 
Specific Names in Zoology with Name No. 913 


Page 
159 


340 


59 


49 


59 


[55 


24 


20 


Lie 


. 61-62 


Li. 


| 


20 


Volume 1, Section D 


griseus Nilsson, 1821, as published in the combination Eurynorhynchus griseus (a 
junior objective synonym of pygmaea Linnaeus, 1758, as published in the com- 
bination Platalea pygmaea), placed on the Official Index ep iee and Invalid 
Specific Names in Zoology with Name No. 351 : 


Grus Brisson, 1760 (Class Aves), substitution of, on the eee List aE Generic 
Names in Zoology i in place of Grus Pallas, 1767 [recte 1766] 


determination of gender of 


Grus Pallas, 1767 [recte 1766], removal of, from the Official List of Generic Names 
in Zoology, on which placed by Opinion 103, and insertion of, on the Official 
Index of Rejected and Invalid Generic Names in seats as a junior homonym of 
Grus Brisson, 1760, with Name No. 764 : : : 


grus Linnaeus, 1758, as published in the combination Ardea grus (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 1062 


Gryllus Linnaeus, 1758 (Class Insecta), placed on the Official List a Generic Names 
in Zoology in Opinion 104, determination of gender of 


Gymnothorax Cuvier, [1800] (a nomen nudum), placed on the eae e Index | saci ase 
and Invalid Generic Names in Zoology with Name No. 796 


Gymnothorax Duda, 1923 (a junior homonym of Gymnothorax Bloch, 1795), 
placed on the Official Index sa eet and Invalid Generic Names in Zoology 
with Name No. 797 Re é shy ae ae F 


gyrinus Cuvier & Valenciennes, 1837, deletion of, as specific name of type species 

of Eleotris Schneider, 1801 (Class Pisces), from entry on Official List of Generic 

Names in Zoology and substitution therefor of pisonis Gmelin ns Fs ed 
Gobius (correction of Opinion 93) oe 


habroptilus Gray (G.R.), 1845, as published in the combination Strigops habroptilus 
(Class Aves), biaces on the ace List neh Brera Names in pa eey with Name 
No. 1009 


Haemadipsa Tennent, 1859 (Class Hirudinea), placed on the Official List as Generic 
Names in Zoology in Opinion 104, determination of gender of 


Haemaphysalis Koch, 1844 (Class Arachnida), placed on the ee List ay Generic 
Names in Zoology in Opinion 73, determination of gender of ; 


haematocheir de Haan, [1835], as published in the combination Grapsus (Pachysoma) 
haematocheir (Class Crustacea, Order Decapoda), eas on the Bec List o. 
Specific Names in Zoology with Name No. 871 


Haliaethus Lesson, 1831 (an Erroneous Subsequent Spelling for Haliaeetus Savigny, 
1809), placed on the Official Index of mee and Invalid Generic Names in 
Zoology with Name No. 551 ts 


419 


Page 


220 


321 
321 


321 


321 


141 


345 


345 


342 


180 


{ey 


143 


15 


216 


420 Opinions and Declarations 


Haliaétos Bonaparte, 1826 (an Erroneous Subsequent Spelling for Haliaeetus 
Savigny, 1809), placed on the ose Index a peaics and Invalid Generic Names 
in Zoology with Name No. 552.. 


Haliaetus Cuvier, 1816 (an Erroneous Subsequent Spelling for Haliaeetus Savigny, 
1809), placed on the Official Index of Baected and Invalid Generic Names in 
Zoology with Name No. 553 é 


haliaetus Linnaeus, 1758 (Falco) (Class Aves) (a nominal species objectively identical 
with Pandion fluvialis PO: pete type asa of Pandion oes 1809, under 
Declaration 21... ; 


placed on the Official List of Specific Names in Zoology with Name No. 1021 


Halimede de Haan, [1835] abs Crustacea, Order sb rcge rte comp Nene of te 
given in Opinion 85 c 


Halimede Rathke, 1843 (a junior homonym of Halimede de Haan, [1835]), placed 
on the Official Index oe ae and Invalid Generic Names in nee with 
Name No. 484 ; : ‘ 


Halimede Oberthuer & Houlbert, 1922 (a junior homonym of Halimede de Haan, 
[1835]), placed on the Official Index is Roce and Invalid Generic Names in 
Zoology with Name No. 485 


harengus Linnaeus, 1758, as published in the combination Clupea harengus (Class 
Pisces), Sa at on the Official 1 List of pace Names in ze with Name No. 
1075 : 


hardwickii Leach, 1817, as published in the combination Nursia hardwickii (Class 
Crustacea, Order Decapoda), pace on the pains List ‘a BREE Names in 
Zoology with Name No. 812... 


harrisii Gould, 1841, as published in the combination Pilumnus harrisii (Class Crus- 
tacea, Order Decapoda), ee on the ae List e ae Names in eee 
with Name No. 914 ie 


helias Pallas, 1781, as published in the combination Ardea helias (Class pes), nae 
on the Official List of Specific Names in Zoology with Name No. 982 


Helice de Haan, [1833] es Crustacea, Order is Spaees Rae of er a 
given in Opinion 85 ; 


Helice Chambers, 1873 (a junior homonym of Helice de Haan [1833]), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 486 


Heliornis Bonnaterre (Class we Cres sgt Boddaert, 1783, nee species of, 
under Declaration 21 ; 


correction of date of, from 1791 to 1790 pan ohare of Opinion 67) 


Heliornis Billberg, 1820 (a junior homonym of Heliornis Bonnaterre, 1790), placed 
on the Official Index past sei pic and Invalid Generic Names in n Zoology with Name 
No:..554° x 3 


Page 


216 


217 


181 
182 


ai! 


a9 


59 


369 


KIT 


51 


59 


181 
213 


217 


Volume 1, Section D 


Helix Linnaeus, 1758 (Class Gastropoda), placed on the Oaieial,t List of Generic 
Names in Zoology i in Opinion 94, determination of gender of ‘ 


hemingi Alcock & Anderson, 1899, as published in the combination Benthochason 
hemingi (Class Crustacea, Order Decapoda), mest on the Ngai List _ ee 
Names in Zoology with Name No. 775 ‘ 


Hemiurus Rudolphi, 1809 (Class Trematoda), placed on the og List ae Generic 
Names in Zoology in Opinion 77, determination of gender of 


henslowii Leach, [1820], as published in the combination Polybius henslowii (Class 
Crustacea, Order Decapoda), plaee se on the anita List non areas Names in 
Zoology with Name No. 825... 


Hepatella Smith (Class Crustacea, Order Decapoda), correction of date of, from 
1870 to 1869, and of entry relating to type species of (correction of Opinion 73) .. 


Hephthopelta Alcock, 1899 oS Crustacea, Order eee rebea)s abies tases of as 
given in Opinion 85 \ 


hepsetus Linnaeus, 1758, as published in the combination Atherina hepsetus (Class 
Pisces), placed on the Official List of Specific Names in Zoology with Name No. 
1066 ie — - et: Be i te et ae re is 


Heteroda Schmidt, 1871 (Class Nematoda), placed on the Official List aire Generic 
Names in Zoology i in Opinion 104, determination of gender of 


Heterophyes Cobbold, 1866 (Class Trematoda), placed on the ee List an Generic 
Names in Zoology in Opinion 84, determination of gender of.. 


Hexapus de Haan, [1833] aes Crustacea, Order ‘asia ite ura re of bee: 
given in Opinion 85 ; 


hippurus Linnaeus, 1758, as published in the combination Coryphaena hippurus 
(Class Pisces), placed on the eed List es ees Names in ane, with Name 
No. 1076 


hirtellus Linnaeus, 1761, as published in the combination Cancer hirtellus (Class 
Crustacea, Order Decapoda), Bee ° on the OCI List ia Mins Names in 
Zoology with Name No. 899... 


hirtipes Heller, [1865], as published in the combination Pinnaxodes hirtipes (Class 
Crustacea, Order Decapoda), ae on the One List des eee Names in 
Zoology with Name No. 900... 


Hirudo Linnaeus, 1758 (Class Hirudinea), placed on the Ocal F List of: Generic 
Names in Zoology i in Opinion 75, determination of gender of 


hispida Eydoux & Souleyet, 1842, as published in the combination Domecia hispida 
(Class Crustacea, Order Decapoda), peed on the Ofiiclaks List of Specific Names 
in Zoology with Name No. 789.. 


155 


54 


51 


368 


7 


195 


51 


eM 


18 


19 


157 


422 Opinions and Declarations 


hoazin Miiller (P.L.S.), [1776], as published in the combination Phasianus hoazin 
es paces on the al List ni heat Names in sa ies with Name 
fe) 1 


Holometopus Milne Edwards (H.), 1853 eats Crustacea, Order ata ee com- 
pletion of Ruling given in Opinion 85 


Holometopus Angelin, 1854 (a junior homonym of Holometopus Milne Edwards 
(H.), 1853), placed on the Cue Index of ROG and Invalid Generic Names 
in Zoology with Name No. 487. 


Holopus d’Orbigny, 1837 (Class Crinoidea), placed on the as List cg Generic 
Names in Zoology in Opinion 73, determination of gender of. . 


Holothuria Linnaeus, 1767 (Class Holothuroidea), placed on the fee List gh 
Generic Names in Zoology i in Opinion 80, determination of gender of ‘ 


Holothuriophilus Nauck, 1880 (Class Crustacea, Order wheerrct Mora of 
Ruling given in Opinion 85 ais @ 


Homalaspis Reinhard, 1860 (Class Insecta, Order Hymenoptera), suppression of, 
under the Plenary Powers, for the purposes both of the Law of Priority and of the 
Law of Homonymy nae “2 an : : ee Be 


placed on the Official Index rg ae and Invalid Generic Names in oe 
with Name No. 755 .. a 2 : 


Homalaspis Milne Edwards (A.), 1863 (Class Crustacea, Order Decapoda), placed 
on the Official List of Generic Names in sy ne es Opinion 85, substitution of 
revised entry for 3 


determination of gender of 


Homalaspis Brunner von Wattenwyl, 1895 (a junior homonym of Homalaspis Milne 
Edwards (A.), 1863), placed on the osc Index an Ree and Invalid Generic 
Names in Zoology with Name No. 488 


Homalaspis Kaier, 1932 (a junior homonym of Homalaspis Milne Edwards (A.), 
1863), placed on the Official Index “A nee and Invalid Generic Names in 
Zoology with Name No. 489... 


Homarus Weber, 1795 (Class Crustacea, Order Decapoda), placed on the Official 
List of Generic Names in Zoology by Opinion 104, substitution of revised entry for 


determination of gender of 
Homarus Milne Edwards (H.), 1837 (a junior homonym of Homarus Weber, 1795), 


placed on the Official Index 2 pines and Invalid Generic Names in Zoology 
with Name No. 756 de : 


Homarus Broun, 1881 (a junior homonym of Homarus Weber, 1795), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 490 


Page 


183 


51 


59 


156 


158 


49 


283 


284 


283 
283 


59 


a2 


295 
295 


295 


59 


Volume 1, Section D 


horridus Linnaeus, 1758, as published in the combination Crotalus horridus (Class 
Reptilia), placed on the ba aie List i epee Names in shai Na with Name 
No. 1107 : 


horridus Riippell, 1830, as published in the combination Oreophorus horridus (Class 
Crustacea, Order Decapoda), Blaredue on the peal List eh Specie Names in 
Zoology with Name No. 815... 


Hyalomma Koch, 1844 (Class Arachnida), placed on the oe List ly Generic 
Names in Zoology i in Opinion 73, determination of gender of : 


Hydra Linnaeus, 1758 (Class Hydrozoa), placed on the Official List ats Generic 
Names in Zoology i in Opinion 77, determination of gender of. . : 


Hymenolepis Weinland, 1858 (Class Cestoda), placed on the nieiah: List a Generic 
Names in Zoology in Opinion 77, determination of gender of F 


Hypoderma Latreille, 1818 (Class Insecta), placed on the Official List of Generic 
Names in Zoology in Opinion 106, determination of gender of : 


Ibidorhinchus David, 1875 (an Erroneous Subsequent Spelling for Ibidorhyncha 
Vigors, 1832), placed on the Official Index a poe and Invalid Generic Names 
in Zoology with Name No. 555 uy 


Ibidorhynchus Gray (G.R.), 1844 (an Erroneous Subsequent Spelling for [bidorhyncha 
Vigors, 1832), placed on the Official Index = HEMET and Inyalid Generic Names 
in Zoology with Name No. 556.. oi 


Iliacantha Stimpson, [1871] (Class Crustacea, Order og aud a ec of 
Ruling given in Opinion 73 5 M Be : : 


imitatrix Hilgendorf, 1869, as published in the combinaion Deckenia imitatrix 
(Class Crustacea, Order Decapoda), act on the ee List of le Names 
in Zoology with Name No. 788. 


impressus Smith (S.I.), 1870, as published in the combination Glyptograpsus 
impressus (Class Crustacea, Order Decapoda), ps on the Paes List sd 
Specific Names in Zoology with Name No. 864 


inaequipes Stimpson, 1858, as published in the combination Asthenognathus 
inaequipes (Class Crustacea, Order Decapoda), en on the Coe List of 
Specific Names in Zoology with Name No. 843 j 


indus Boddaert, 1783, as published in the combination Falco indus (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 1029 .. 


integerrimus Lamarck, 1818, as published in the combination Cancer integerrimus 
(Class Crustacea, Order Decapoda), aaa te on the isles List on epee Names 
in Zoology with Name No. 771. 


Iphiculus White, 1847 (a nomen nudum), placed on the alas Index ae ere and 
Invalid Generic Names in Zoology with Name No. 491 4 


IS? 
LST 


143 


Za 


217 


14 


12 


183 


60 


424 Opinions and Declarations 


Page 
Iphiculus Adams & White (Class Crustacea, Order Gopatasens correction of date of, 
from 1848 to [1849] (correction of Opinion 73) ; 56 


Iphis Meigen, 1800 (Class Insecta, Order Diptera), suppression of, under the Plenary 
Powers, for the purposes both of the Law of Priority and of the Lawof Homonymy 268 


insertion of, on the Official Index of Rejected and Invalid Generic Names in 


Zoology with Name No. 754 “ aye e a ue as 267-268 
Iphis Leach, 1817 (Class Crustacea, Order Decapoda), placed on the Official List Re 

Generic Names in Zoology in Opinion 73, substitution of revised entry for.. “ 267 

determination of gender of si wii se = =" NG sfie os meee 


Iphis Koch (C.L.), 1835 (a junior homonym of Jphis Leach, 1817), placed on the 
Official Index of se tence and Invalid Generic Names in Zoology with Name No. 
492 60 


Iphis Laporte, 1836 (a junior homonym of Jphis Leach, 1817), placed on the Official 
Index of Rejected and Invalid Generic Names in Zoology with Name No. 493 .. 60 


isenbeckii Brandt, 1848, as published in the combination Platycorystes (Podacanthus) 
isenbeckii (Class Crustacea, Order Decapoda), tae on the ee List s 
Specific Names in Zoology with Name No. 793 ‘ 6 


Ixodes Latreille, 1795 (Class Arachnida), sine on the Cia List o Generic Names 
in Zoology in Opinion 73, determination of gender of. . 142 


jacquemontii Milne Edwards (H.), 1844, as published in the combination 
Gecarcinucus jacquemontii (Class Crustacea, Order Decapoda), placed on the 
Official List of Specific Names in Zoology with Name No. 796... Rs ae es 


javanicus Horsfield, 1821, as published in the combination Eurylaimus javanicus 
(Class Aves), placed on the Cae List bg sa ale Names in gs with Name 
No. 981 .. 177 


jurinei Saussure, 1853, as published in the combination Guaia (lia) jurinei (Class 
Crustacea, Order Decapoda), aka on the ee List an SPEER Names in 
Zoology with Name No. 805... 8 


jurinii Bell, 1855, as published in the combination Leucosilia jurinii (an Erroneous 
Subsequent Spelling for jurinei Saussure, 1853, as published in the combination 
Guaia (Ilia) jurinei), placed on the Official Index ts Schad and Invalid og 
Names in Zoology with Name No. 339 62 


Lacerta Linnaeus, 1758 (Class Reptilia), completion of Ruling given in Opinion 92 341 


lacertina Linnaeus, 1766, as published in the combination Siren lacertina (Class 
Amphibia), placed on the RS List ed pectic, Names in nuZanlony with Name 
No. 1104 373 


Volume 1, Section D 


Lachnopodus Stimpson, 1858 (Class Crustacea, Order Res rainy ee of 
Ruling given in Opinion 85 : : 


laevigata Milne Edwards (H.) & Lucas, as published in the combination 
Pinnotherelia laevigata (Class Crustacea, Order Decapoda), correction of date of, 
from 1843 to [1844] (correction of Opinion 85) a - , aq 


placed on the Official List of Specific Names in Zoology with Name No. 902 


lambriformis Milne Edwards (A.), 1873, as published in the combination Merocryptus 
lambriformis (Class Crustacea, Order Decapoda), placed 5 on the fee List of 
Specific Names in Zoology with Name No. 809 : 


lamelligera Stimpson, 1858, as published in the combination Onychomorpha 
lamelligera (Class Crustacea, Order Decapoda), placed on the ow List o 
Specific Names in Zoology with Name No. 814 : : 


Latrodectus Walckenaer, 1805 (Class Arachnida), placed on the Official List o 
Generic Names in Zoology i in Opinion 104, determination of gender of 


lavaretus Linnaeus, 1758, as published in the combination Salmo lavaretus (Class 
Pisces), placed on the Official List of Specific Names in Zoology with Name No. 
1095 oa ne Ar urs ee oe oe ae oe Se sy 


leeanus Rathbun, 1898, as published in the combination Lipaesthesius leeanus (Class 
Crustacea, Order Decapoda), peccar’ on the iar wast List of es Names in 
Zoology with Name No. 878 é ; : : 


Lepas Linnaeus, 1758 (Class Crustacea), placed on the eds List os Generic Names 
in Zoology in Opinion 77, determination of gender of 3 


Lepidocyclina Giimbel, [1870] (Class Rhizopoda), placed on the Official List oh 
Generic Names in Zoology i in Opinion 127, determination of gender of 


Leptodius Milne Edwards (A.), 1863 ab asian Order pecenora) coinpienge 
of waite given in Opinion 85 .. 


Leucochila von Martens, 1860 (Class Gastropoda), placed on the Official List mi 
Generic Names in Zoology in Opinion 115, determination of gender of 


leucorodia Linnaeus, 1758, as published in the combination Platalea leucorodia 
(Class Aves), pers on the eee List Ae Seige Names in ET oooey with Name 
No. 998 .. 


Leucosilia Bell, 1855 (Class Crustacea, Order ee correction of reference 
for (correction of Opinion 73) . ; aes cf ioe a 


Liagora Gistl, 1848 (an Erroneous Subsequent Spelling of Liagore de Haan, [1833],) 
placed on the Official Index of aa and Invalid Generic Names in aban 
with Name No. 494 : ne be ; 


Liagore de Haan, [1833] Sree Crustacea, Order Se wiaiten) Hy eae of one 
given in Opinion 85 : 


425 


Page 
51 


25 


372 


156 
py! 


ew! 


by9 


J 


60 


51 


426 Opinions and Declarations 


Libystes Milne Edwards (A.), 1867 Soi Crustacea, Order si case ei ere 
of Ruling given in Opinion 85 .. 


Ligula Bloch, 1782 (Class Cestoda), placed on the Official List ies Generic Names in 
Zoology in Opinion 104, determination of gender of.. 


liguricus Milne Edwards (H.), 1853, as published in the combination Euchirograpsus 
liguricus (Class Crustacea, Order Decapoda), Pre on the eae List a 
Specific Names in Zoology with Name No. 854 : 


Limax Linnaeus, 1758 (Class Gastropoda), placed on the os List os Generic 
Names in Zoology in Opinion 94, determination of gender of 


Limnatis Moquin-Tandon, 1827 (Class Hirudinea), placed on the Official List of 
Generic Names in Zoology in Opinion 104, determination of gender of 


limosa Say, 1818, as published in the combination Cancer limosa (Class Crustacea, 
Order Decapoda), aa on the neal List cal Specie Names in A EOES with 
Name No. 860... 


Limulus Miller (O. F.), 1785 (Class Merostomata), placed on the Official List a 
Generic Names in Zoology in Opinion 104, determination of gender of 


Liomera Dana, 1851 (Class Crustacea, Order De completoE of t Rute 
given in Opinion 85 ‘ - i ae a , ; 


Lipaesthesius Rathbun, 1898 (Class Crustacea, Order Baie paren es of 
Ruling given in Opinion 85 , 


Lissocarcinus Adams & White, 1847 (a nomen nudum), placed on the Official Index 
of Rejected and Invalid Generic Names in Zoology with Name No. 495 .. 


Lissocarcinus Adams & White (Class Crustacea, Order Vege eer correction of 
date of, from 1848 to [1849] (correction of Opinion 73) : of a 


Lithadia Bell, 1855 (Class Crustacea, Order peo correction of reference for 
(correction of Opinion 73) eA : ay ae xs rz Ay 


Lithostrotion Fleming, 1828 (Class Anthozoa), placed on the Official List is Generic 
Names in Zoology in Opinion 117, determination of gender of : 


Litocheira Kinahan, 1856 (Class Crustacea, Order tena basi Sea of RUNRE 
given in Opinion 85 d a2 


litteratus Fabricius, 1798, as published in the combination Cancer litteratus (Class 
Crustacea, Order Decapoda), ee on the parnnan List . SP ean Names in 
Zoology with Name No. 926... 


Lophius Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 . 


Lophopanopeus Rathbun, 1898 (Class Crustacea, Order rDecapen): eqriplesonis of 
Ruling given in Opinion 85 ; 


Page 
51 


1357 


13 


156 


157 


14 


142 


51 


51 


60 


56 


55 


139 


51 


22 


340 


49 


Volume 1, Section D 


Lophopilumnus Miers, 1886 (Class Crustacea, Order ae riparia henge He of 
Ruling given in Opinion 85 A ‘ 


lucius Linnaeus, 1758, as published in the combination Esox lucius (Class Pisces), 
placed on the Official List of Specific Names in Zoology with Name No. 1093... 


lugubris Alcock, 1899, as published in the combination Hephthopelta lugubris (Class 
Crustacea, Order Decapoda), pen ae on the pica List ag Specific Names in 
Zoology with Name No. 869... 


Luidia Forbes, 1839 (Class Asteroidea), placed on the pe i List a Generic Names 
in Zoology in Opinion 129, determination of gender of 


Lumbricus Linnaeus, 1758 (Class Chaetopoda), placed on the a List ef Generic 
Names in Zoology i in Opinion 75, determination of gender of : 


lumpus Linnaeus, 1758, as published in the combination Cyclopterus lumpus (Class 
Pisces), placed on the Official List of Specific Names in Zoology with Name No. 
1077 a as: = ae 3 = ne ie a ae Ae 


Lupocyclus White, 1847 (a nomen nudum), placed on the Official Index us Passi 
and Invalid Generic Names in Zoology with Name No. 496 


Lupocyclus Adams & White (Class Crustacea, Order Meigen correction of date 
of, from 1848 to [1849] (correction of Opinion 73) ‘ 


lupus Linnaeus, 1758, as published in the combination Anarhichas lupus (Class 
Pisces), placed on the Official List of Specific Names in Zoology with*Name No. 
1065 ee oe a we » on ae X fi ne i 


Lybia Milne Edwards (H.), 1834 sare Crustacea, Order frag ee poh ar of 
Ruling given in Opinion 85 A 


lyra Linnaeus, 1758, as published in the combination Callionymus lyra (Class Pisces), 
placed on the Official List of Specific Names in Zoology with Name No. 1073... 


Lytoceras Suess, 1865 (Class Cephalopoda), placed on the Official List phi Generic 
Names in Zoology i in Opinion 130, determination of gender of : 


Mabouia Cuvier, 1829 (an Erroneous Subsequent Spelling for Mabuya (Rafinesque) 
Fitzinger, 1826), placed on the Official Index of ea and Invalid Generic 
Names in Zoology with Name No. 798 ; 


Mabouya Duméril & Bibron, 1839 (an Erroneous Subsequent Spelling for Mabuya 
(Rafinesque) Fitzinger, 1826), placed on the Official Index a Bre and Invalid 
Generic Names in Zoology with Name No. 799 s 


Mabuia Cuvier, 1829 (an Erroneous Subsequent Spelling for Mabuya (Rafinesque) 
Fitzinger, 1826), placed on the Official Index of pa and Invalid Generic 
Names in Zoology with Name No. 800 , 


427 


Page 
49 


371 


15 
158 


iD 


“56 


368 


51 


369 


159 


345 


345 


345 


428 Opinions and Declarations 


Mabuya Rafinesque, 1815 (a nomen nudum), placed on the Official Index i area etit 
and Invalid Generic Names in Zoology with Name No. 801 


macgillivrayi White, 1852, as published in the combination Ommatocarcinus 
macgillivrayi (Class Crustacea, Order Decapoda), cae on the has ese List 2 
Specific Names in Zoology with Name No. 889 


Mactra Linnaeus, 1767 (Class Pelecypoda), placed on the ewan s List ot Generic 
Names in Zoology i in Opinion 94, determination of gender of : 


maculatus Linnaeus, 1758, as published in the combination Cancer maculatus (Class 
Crustacea, Order Decapoda), placed on the Dasciaerd List S ine paas Names in 
Zoology with Name No. 777... Bc 


magna Linnaeus, 1758, as published in the combination A/lauda magna (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 1010 3. 


major Linnaeus, 1758, as published in the combination Parus major (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 994... 


Malapterus Jarocki, 1822 (an Erroneous Subsequent Spelling for Malapterurus 
Laceépéde, 1803), placed on the Official Index of pny and Invalid Generic 
Names in Zoology with Name No. 802 . 


Malapturus Swainson, 1838 (an Erroneous Subsequent Spelling for Malapterurus 
Lacépéde, 1803), placed on the Official Index of kane and Invalid Generic 
Names in Zoology with Name No. 803 ‘ : 


mantellii Owen, 1848, as published in the combination Notornis mantellii (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 990 


Manucodia Boddaert, 1783 (Class Aves), Paradisea sel ere ih Forster, 1781, 
type species of, under Declaration 21 . 


marginalis Bloch, 1793, as published in the combination Epinephelus marginalis 
(Class Pisces), placed on the Oita Vist ia epscigci) Names in neevlesy with Name 
No. 1097 


marinus Fabricius, 1775, as published in the combination Astacus marinus (a junior 
objective synonym of gammarus Linnaeus, 1758, as published in the combination 
Cancer gammarus), placed on the Official Index ay Reecieg and Invalid RPE 
Names in Zoology with Name No. 361 


maximus Geoffroy, 1850, as published in the combination Aepyornis maximus (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 952 


media Koch, 1816, as published in the combination Gallinago media (a junior 
objective synonym of gallinago Linnaeus, 1758, as published in the combination 
Scolopax gallinago), placed on the Official Index “4 age and Invalid praeie” 
Names in Zoology with Name No. 341 


Page 
345 


17 


158 


180 


178 


345 


345 


178 


181 


Biz 


296 


175 


98 


Volume 1, Section D 


melanochirus Milne Edwards (A.), 1873, as published in the combination Chlorodopsis 
melanochirus (Class Crustacea, Order Decapoda), eajieeh on the eae Lisi a 
Specific Names in Zoology with Name No. 781 ; : : 


meleagris Linnaeus, 1758, as published in the combination Phasianus meleagris 
(Class Aves), oly on the ee List nee ae Names in pees) with Name 
Wo. 991 .. 


Melybia Stimpson, [1871] mete Crustacea, Order gman a at of Fag 
given in Opinion 85 g 


mercurialis Merrem, 1820, as published in the combination Sphargis mercurialis (a 
junior objective synonym of coriacea Linnaeus, 1766, as published in the com- 
bination Testudo coriacea), placed on the Official Index off Rejected and Invalid 
Specific Names in Zoology with Name No. 365 8 ; a, a3 ie 


Metacrinus Carpenter, 1882 (Class Crinoidea), placed on the oad List - Generic 
Names in Zoology in Opinion 73, determination of gender of ‘ 


Metasesarma Milne Edwards (H.), 1853 ena Crustacea, Order Deo) com- 
pletion of Ruling given in Opinion 85 


Metopocarcinus Stimpson, [1871], (Class Laubach Order Decapoda), PCPs 
of Ruling given in Opinion 85 .. ay : rf ne be 


Micropanope Stimpson, [1871] ae Crustacea, Order ae buen of 
Ruling given in Opinion 85 3 


microps Alcock & Anderson, 1894, as published in the combination Xanthodes 
microps (Class Crustacea, Order Decapoda), aed on the aaa List on en 
Names in Zoology with Name No. 891 


militaris Linnaeus, 1758, as published in the combination Emberiza militaris (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 1030 


milneedwardsii Miers, 1886, as published in the combination Paracyclois milne- 
edwardsii (Class Crustacea, Order caine Bina on the eg List a BREE 
Names in Zoology with Name No. 817 


minutus Linnaeus, 1758, as published in the combination Cancer minutus (Class 
Crustacea, Order Decapoda), sabiets on the OO List ney Seow Names in 
Zoology with Name No. 943... ‘ 


mocinno de la Llave, 1832, as published in the combination Pharomachrus mocinno 
(Class Fakes pers on ‘the cla List rey a Names in ELON with Name 
No. 996 .. 


moebii Richters, 1880, as published in the combination Xenophthalmodes moebii 
(Class Crustacea, Order oe air on the es List _ eee Names 
in Zoology with Name No. 929. 


Molgula Forbes, 1848 (Class Urochorda), placed on the Official List ie Generic 
Names in Zoology in Opinion 94, determination of gender of : 


429 


Page 


178 


od 


346 


156 


51 


ai 


51 


18 


183 


23 


178 


an 


158 


430 Opinions and Declarations 


Page 
Moniezia Blanchard, 1891 (Class Cestoda), placed on the Omelet List ai Generic 

Names in Zoology in Opinion 77, determination of gender of : LS 
monstrosa Linnaeus, 1758, as published in the combination Chimaera monstrosa 

(Class Pisces), placed on the oiierar List Lee aia Names in neo es with Name 

No. 1074 : 369 
montagui [Leach], [1814], as published in the combination Pandalus montagui (Class 

Crustacea, Order Decapoda), amen on the re Oees List al Speniiraie Names in 

Zoology with Name No. 1039 .. 4 245 
morhua Linnaeus, 1758, as published in the combination Gadus morhua (Class 

Pisces), kG on the cae List of ee Names in site ee with Name No. 

1079 ; 369-370 
Mormyrus Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 340 
moschata Linnaeus, 1758, as published in the combination Anas moschata (Class 

Aves), placed on the Official List of Specific Names in Zoology with Name No. 964 ~—-175 
Mugie Macklot, 1830 (an Erroneous Subsequent Spelling for Mugil Linnaeus, 1758), 

placed on the Official Index a ae and Invalid Generic Names in Zoology 

with Name No. 804 a : ia x. - ; 345 
Mullus Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 340 
muriger Adams & White, as published in the combination Tlos muriger (Class 

Crustacea, Order Decapoda), correction of date of, from 1848 to [1849] (correction 

of Opinion 73) ; aA : Be : : Bhs aie ei 24 

placed on the Official List of Specific Names in Zoology with Name No. 837 11 
murigera White, 1846, as published in the combination Xanthasia murigera (Class 

Crustacea, Order Decapoda), placed 2 on the Ve List oe Specie Names in 

Zoology with Name No. 927... : vas 
Musca Linnaeus, 1758 (Class Insecta), placed on the A pad List a Generic Names 

in Zoology in Opinion 82, determination of gender of : 142 
Museum Boltenianum, 1798, authorship of, to be attributed to R6ding (P.F.) ; 2x0 

addition of title of, to the ‘Official List a Works S quetaved as Available gies Zoological 

Nomenclature with Title No. 26 257 
Musophagus Bonnaterre, 1790 (an Invalid Emendation of Musophaga Isert, 1789), 

placed on the Official Index af Haaged and Invalid Generic Names in Zoology 

with Name No. 557 ee 210 
Mustellus Fischer de Waldheim, 1813 (an Erroneous Subsequent Spelling for 

Mustelus Linck, 1790), placed on the Sola & Index “ Reraes and Invalid Generic 

Names in Zoology with Name No. 805 346 


Volume 1, Section D 


mustelus Linnaeus, 1758, as published in the combination Squalus mustelus (Class 
ls paces on the a eases List of Brena: Names in OMG with Name No. 
1 


Mya Linnaeus, 1758 (Class Pelecypoda), placed on the annie List a Generic Names 
in Zoology in Opinion 94, determination of gender of 


Myrodes Bell, 1855 (Class Crustacea, Order ere Nea correction of reference for 
(correction. of Opinion 73) ta a vs 


Mytilus Linnaeus, 1758 (Class Pelecypoda), placed on the Bisel List ef Generic 
Names in Zoology i in Opinion 94, determination of gender of ‘ 


Nabis Latreille, [1802—1803] (Class Insecta), placed on the Official List “ Generic 
Names in Zoology in Opinion 104, determination of gender of ; 


narina Stephens, 1815, as published in the combination Trogon narina (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 955 


neapolitana Kossmann, 1878, as published in the combination Paragalene 
neapolitana (Class Crustacea, Order Decapoda), peeeaye on the Ratan List 2 
Specific Names in Zoology with Name No. 893 : 


Nephrops [Leach] (Class Crustacea, Order Peco) correction of date of, from 
1815 to [1814] (correction of Opinion 104) .. f af of et +e 


Neritina Lamarck, 1816 (Class Gastropoda), placed on the Official List be: Generic 
Names in Zoology i in Opinion 119, determination of gender of 


niger Linnaeus, 1758, as published in the combination Gobius niger (Class Pisces), 
placed on the Official List of Specific Names in Zoology with Name No. 1081 


niger Miller (P.L.S.), [1776], as published in the combination Bucco niger (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 965 


niger Hartig, 1840, as published in the combination Figites niger (Class Insecta, 
Order Hymenoptera), uaa on the ao List a Be Names in eee 
with Name No. 1058 


nigra Gmelin, 1788, as published in the combination Paradisea [sic] nigra (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 960 


nigricephala Jameson, 1835, as published in the combination Tanagra nigricephala 
(Class Aves), placed on the alee List ate ae Names in fae ee with Name 
No. 1034 


nitidus Latham, 1790, as published in the combination Falco nitidus (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 1025 


nitidus Stimpson, 1858, as published in the combination Sphaerozius nitidus (Class 
Crustacea, Order Decapoda), placed on the * Oaiecial List es eicaite Names in 
Zoology with Name No. 921... 


431 
Page 


372 


158 


55 


156 


14] 


174 


18 


x7 


158 


370 


175 


284 


174 


184 


183 


21 


432 Opinions and Declarations 


nitidus Stimpson, 1859, as published in the combination Euryplax nitidus (Class 
Crustacea, Order Decapoda), ty on the ee List bo ee Names in 
Zoology with Name No. 8359  ;. f 


nitidus Milne Edwards (A.), 1867, as published in the combination Libystes nitidus 
(Class Crustacea, Order Decapoda), shai on the a List a SPEER Names 
in Zoology with Name No. 876.. 


nitidus Milne Edwards (A.), 1870, as published in the combination Catoptrus nitidus 
(Class Crustacea, Order Decapoda), Reece on the eee List 4) Spa Names 
in Zoology with Name No. 847.. 


nitidus Smith (S.I.), 1870, as published in the combination Dissodactylus nitidus 
(Class Crustacea, Order Decapoda), ert ae on the pe List “a cree Names 
in Zoology with Name No. 852.. 


nitidus Milne Edwards (A.), 1873, as published in the combination Notonyx nitidus 
(Class Crustacea, Order Decapoda), pee on the Bue List of Specie: Names 
in Zoology with Name No. 887.. 


nobile Milne Edwards (A.), 1880, as published in the combination Trichopeltarion 
nobile (Class Crustacea, Order Decapoda), pie de on the Sian: List of SERGI 
Names in Zoology with ‘Name No. 840 


nodosus Bell, 1855, as published in the combination Oreophorus nodosus (Class 
Crustacea, Order Decapoda), isa on the Pi dias List re ee Names in 
Zoology with Name No. 831... 


norvegicus Linnaeus, 1758, as published in the combination Cancer norvegicus 
(Class Crustacea, Order Decapoda), ape on the eee go List e Retin Names 
in Zoology with Name No. 934.. 


Notonecta Linnaeus, 1758 (Class Insecta), placed on the Official List 4 Generic 
Names in Zoology i in Opinion 104, determination of gender of : 


Notonyx Milne Edwards (A.), 1873 a Crustacea, Order Pecnpoa comes 
of Ruling given in Opinion 85 .. 


novaehollandiae Latham, 1801, as published in the combination Cereopsis novae- 
hollandiae (Class Aves), placed ¢ on the Sane List ef pres Names in Ba 
with Name No. 969 : 


Numidia Forster, 1817 (an Erroneous Subsequent Spelling for Numida Linnaeus, 
1764), placed on the Official Index ot i cheed and Invalid Generic Names in 
Zoology with Name No. 558 3 


Nursilia Bell, 1855 (Class Crustacea, Order Pe correction of reference for 
(correction of Opinion 73) ce ? ae : a or x 


Nycteribia Latreille, 1796 (Class Insecta), placed on the Official List cy Generic 
Names in Zoology i in Opinion 128, determination of gender of ; 


Page 


14 


16 


12 


13 


17 


12 


14 


22 


143 


51 


175 


at7 


55 


143 


Volume 1, Section D 


occidentalis Lawrence, 1858, as published in the combination Podiceps occidentalis 
(Class ‘eke placed on the Cine: List my, elie Names in seas with Name 
Mo, 951... 


ocellaris Linnaeus, 1758, as published in the combination Blennius ocellaris (Class 
Pisces), placed on the Official List of Specific Names in Zoology with Name No. 
1092 ee oe Li a6 ~ sR a ae Ae ha th: 


Oedemagena Latreille, 1818 (Class Insecta), placed on the Giga! List oh Generic 
Names in Zoology in Opinion 106 determination of gender of 


Oediplax Rathbun, 1893 isa Crustacea, Order Poesia eee of etn >> 
given in Opinion 85 : 


oenas Linnaeus, 1758, as published in the combination Columba oenas (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 944 : 


Oestrus Linnaeus, 1758 (Class Insecta), placed on the Official List me Generic Names 
in Zoology in Opinion 106, determination of gender of é ; ; 


Official Index of Rejected and Invalid Family-Group Names in Zoology, names 
placed on: 


DOLICHOPIDAE Stephens (J.F.), 1809 
DOLICHOPODES Latreille, 1809 .. 
DOLYCHOPODAE Samouelle, 1819 


Official Index of Rejected and Invalid Generic Names in Zoology, names placed 
on: 


Acanthocyclus Dybowski, 1873 
Acanthopus Oken, 1816 
Acanthopus Dahl, 1823 .. 
Acanthopus Latreille, 1829 
Acanthopus de Haan, [1835] 
Acanthopus Muenster, 1839 
Acanthopus Giebel, 1872 
Acanthopus Vernet, 1877 
Acipenses Linck, 1790 

Aegithia Blyth, 1860 ‘ 

Aix Agassiz (J.L.R.), 1846 
Alauda Daudin, 1816... 
Ampelis Nozeman & Vosmaer, 1758 
Ammodytes Bonaparte, 1831 ¢ 
Anarhicas Latreille, 1804 
Aptenodita Bonnaterre, 1791 
Aptenodites Lesson, 1837 Bas 
Aptenodyta Gmelin (J.F.), 1788 
Aptenodytes Forster (J.R.), 1777 
Asthenognathus Bocourt, 1884 .. 
Asthenognathus Broom, 1915 
Aulacorhynchus Dittmar, 1872 
Bathynectes Guenther, 1878 
Bathynectes Brinkman, 1913 


433 
Page 


173 


371 


141 


51 


86 


141 


269 
269 


434 Opinions and Declarations 


Page 
Official Index of Rejected and Invalid Generic Names in Zoology, names placed 

on (contd.) : 
Bellia Bate, 1851 Pe ant - ae ie ae < 2s 58 
Bellia Gray (J.E.), 1869 . we ate aa ad ae a ae iv 58 
Bellia Tutt, 1902 . Ye >. ve Ea a¥ ae te 58 
Brotogerys Agassiz (J.L. R. we 1846 oe ee as ae a a5  UEDTS 
Calliongmus Linck, 1790 : a a ee ae 5 qs os 343 
Callionimus Gouan, 1770 Ne oie Me as mA me om «3° ~343 
Capella Frenzel, 1801 er os Lo ae y ‘ a as 97 
Capella Keyserling & Here 1840 =e a Kas As oa ays =F 97 
Caphyra Barrande, 1846 ie x ae a ae Bg a 58 
Capito Gistl, 1848 ; oe. A if Ke Me oe - on ots 
Cathartes d’Orbigny, [1835] avs = fl th ae a. 38 reid 
Centrocerus Fitzinger, 1843 a ae ot se oh oe a sy eelS 
Cephalopterus Risso, 1810 47. i * oa hee a S&S SLES 
Cephalopterus Powrie, 1870 a: i Le 5 # st ae ot pe 
Chauna Loew, 1847 ne - Ae os Ae bs a sh LenS 
Chimaera Poli, 1791 ae : we oe Ae es oe ve ae 343 
Chimaera Ochsenheimer, 1808 ue he se Oe ve = .3| 385 
Chimaera Hitchcock, 1858 8 ie ie ae e i: ee .. ‘343 
Chimaira Duméril, 1856 by at Bs HA Ho vee MBAS 
Chlorodopsis Milne Edwards (A. ), 1863 ie e a ee as ae 58 
Chrysolophus Swainson, 1837 .. & Le i se on ia) ele 
Clamator Blyth, 1852... fie see =e Wy, Re a he aa | BAIG 
Coerebra Boie, 1826 Et by fe s oe bs Ne bre .. V2L6 
Columbus Linnaeus, 1758 We he oe, =e we os Pe a3! 86 
Columbus [Tunstall], 1771 a Re mA oz a “ es e 86 
Coregonus Lacépéde, 1803 a Ss es Ae be A oe ig TAS 
Coregonus Jarocki, 1822.. a oe i mal > se oot 343 
Coriudo Fleming, 1822 .. ee of + gt cn on 2. "343 
Coryhaena Stiles, 1922 .. bm a a fe’ ah wi se wan 349 
Coryphaena Coues, 1889 < we ae ba Se ue as vs | ga 
Coturnix Cuvier, 1800 .. As af es ue ie ce J ry SG 
Cyclopterus Marseul, 1872 ve ki i 8 se ate i 8 344 
Cyrtograpsus Carruthers, 1867 .. - 7 ae a hs Bre Se 59 
Dermatochelys Wagler, 1830... a ee bye a = aE .. 344 
Dermochelis Lesueur, 1829 2 a a a Bis ee bic ies 344 
Diodon OReilly, 1818 .. of ey a ae oe - fe che 344 
Diodon Lesson, 1828 Ke Ad Es aD =h ae ae amt nS 344 
Diodon Lesson, 1830 : Be ier ae Fe es ae .. 344 
Dolichopus Van der Hoeven, [1856] * :3 fs a oe - 32 268 
Dolychopus Samouelle, 1819 ae Ae oh a we Mae 4A Set 26S 
Draco Gouan, 1770 ae a ae Pa ah ae as 2 le 344 
Draco Oken, 1816 ae i a os om a ay =; .. 344 
Eleotris Gronovius, 1763 58 at of, et 3 Ah oe 94d. 
Eucrate Rafinesque, 1815 7 a ran i sk me mi me 34) 
Eurylaemus Gloger, 1841 As “s % a 3 Aes fe aa 216 
Eurynorhinchus Lafresnaye, 1842 aif ae ae a3 oh me AG 
Eurynorynchus Reichenbach, 1852 ae Be 2 me ip on app phew AS 
Euryplax Klapalek, 1909 oe +h ie ap 4 ir) n, hee 59 
Eurypygia Swainson, 1837 Bas re se =a ie eu By a NG 
Fistularia Forskal, 1775 .. ee sp ue a BS a Ne .. 344 
Fistularia Miller (O.F.), 1776 .. a, gi ay ne abd Ri af AA 
Fistularia Lamarck, 1816 a sh 58 a Ms “a ch .. 344 
Fistularia Blainville, 1830 Y, - os mM ae M me .. 344 
Fistularia Bowerbank, 1841 ‘. s Hs zt be Me AP nadke | 2, 
Gadus Dejean, 1821 re ne ae 4 ze a ~ an es 345 
Gadus Rang, 1829 hy ik 5% a a ah =p =r PPS AS 
Gadus Deshayes, 1861 .. i a ae aN 4c Ab £2 Sie (2 


Gallinago Koch, 1816 .. Bax oe ze af m nis a Ae 97 


Volume 1, Section D 435 


Page 
Official Index of Rejected and Invalid Generic Names in Zoology, names placed 

on (contd.) : 
Gampsonyx Jordan, 1847 de 3% ae af — oe 7 PDS 
Gecarcinicus Dana, 1852 3 in s -— ot yi ar ie 59 
Goneplat | Leach], [1814] - au aa cd a “a 4 a 59 
Gonoplax Leach, 1815 .. ca ay Bi ats a o. - 7 59 
Grus Pallas, 1767 [recte 1766] .. be = 7 A Le re 9 321 
Gymnothorax Cuvier, [1800] .. * se a fe th e id 345 
Gymnothorax Duda, 1923 a an cd ae >. ae, y ohh 345 
Haliaethus Lesson, 1831 a aa Se aa ne eA i; a 216 
Haliaétos Bonaparte, 1826 ee ne - Se a a ie = 216 
Haliaetus Cuvier, 1816 .. in aie ic Jk i Ae . came | ie 
Halimede Rathke, 1843 7 eg os Se bs fe 59 
Halimede Oberthuer & Houlbert, 1922 a - ai 3% ee fri 59 
Helice Chambers, 1873 .. f se Bi, _ 40 st ae 59 
Heliornis Billberg, £820» . a of ie ie a Ae Be + See? 
Holometopus Angelin, 1854 - fe ue Bs Ht be an oe 59 
Homalaspis Reinhard, 1860 is oe te ad a me 2 ©6283 
Homalaspis Brunner von Wattenwyl, 1895. ri ae e am 3 59 
Homalaspis Kaier, 1932 oe aud a fe on og of 59 
Homarus Milne Edwards (H. ay ($37). ar . a A Ze 21 295 
Homarus Broun, 1881 .. ore aie ae ee he es a 59 
Ibidorhinchus David, 1875 2 ats 8 att # ae Le Oe 
Ibidorhynchus Gray es R:); 1844 a iP 8 s4 by Pe Pee ae 2), 
Iphiculus White, 1847... ws oa in ae by ? oF 60 
Iphis Meigen, 1800 ay ae F ws se 5A it 267-268 
Iphis Koch (C.L.), 1835 . we Ke ad ey <a ae oe Ay 60 
Iphis Laporte, 1836 te ie ie af ae Ay Q A +. 60 
Liagora Gistl, 1848 ’ oe wh a oe 2 ue * 60 
Lissocarcinus Adams & White, 1847... as at 3 AS 7 A 60 
Lupocyclus White, 1847 . ye es of a = of ie 4 60 
Mabouia Cuvier, 1829. A, 33 3 ee ay # x 345 
Mabouya Duméril & Bibron, 1839 fa ns sé Sa * Fas 345 
Mabuia Cuvier, 1829... se - we ie id me ee 345 
Mabuya Rafinesque, 1815 fe ea 4 Be ae As bes .. 345 
Malapterus Jarocki, 1822 Af: m we ae ae ae fe a 345 
Malapturus Swainson, 1838 is ayn sits - Se fe i me 345 
Mugie Macklot, 1830 wi f ie = oy Me oe oe. ne 345 
Musophagus Bonnaterre, 1790 . : ee ae fe 4 BE if 217 
Mustellus Fischer de Waldheim, 1813 ays MM Es AY ns 20'/ 0S46 
Numidia Forster, 1817 .. : ay a tes oF oy. a Wee a) 
Ophidion Pomel, 1853... as ye ne ie rh ie ae Py 346 
Ophidium Linnaeus, 1766 si ae ua “ ry LS: Bai ar 346 
Pachycephala Klug, 1834 ; Ei Me a = a ay Arie 3 
Pachycephala Doleschall, 1858 . ar oy Re *. oF ae Remi?) 
Pachycephala Lioy, 1864 be a7 oe sa ae Yr At eases: 
Pachycephala Broun, 1881 fe Be ae a Pa ¥: M: Ph AZ 
Pachycephala Fairmaire, 1883 .. cf ae os * ‘? re a! 
Pachycephalus Stephens, 1826 .. ne 4. ae a AM ALS 
Pandalus Leach, [1815] (Malac. Podophth. Brit.) J et ie oe 245-246 
Pandalus Leach, 1815 (Trans. linn. Soc. Lond. ) oe ce a e 245-246 
Parotica Wagler, 1827" ae fe fe of a Pre 55 UT 
Persephona Leach, 1852 aie a ies a3 22 ae * 8 60 
Pezoporus Klug, [1842] a Ws Bs “a es ee eh ze a 218 
Pezoporus Foerster, 1868 - ar wg a ae } a” Ag 218 
Pilumnus Rafinesque, 1815 tg it oy Ae aa ia es a 60 
Pilumnus Dejean, 1821 .. ps 33 v2 up 2 ee xe a 60 
Pilumnus Koch (C.L.), 1837... - ae iM: Efe 4h aM! ue 60 
Pilumnus Gistl, 1848 de wd 53 3 a ie a ar er 60 


Pilumnus Bonaparte, 1854 is. Be re as is. #. BH ss 60 


436 Opinions and Declarations 


Page 
Official Index of Rejected and Invalid Generic Names in Zoology, names placed 
on (contd.) : 
Pinnotheres Latreille, Samia a4 i ax At ae a. 235-236 
Planes Saussure, 1862 ry a ae oe as * * eae 60 
Planes Rondani, 1864 .. ee * ae ae ne rs a = 60 
Podacanthus Brandt, 1848 te a Mf ys te - ee Gs 60 
Podophtalmus Lamarck, 1801 .. oy as ae 3% sy ae se 61 
Polydectus Rafinesque, 1815... A fe! Me By: srs os a 61 
Pseudopinnixa Holmes, 1895... a: Ne e oa + “ti i 61 
Psittacus Catesby, 1777 ate ae se ae Pe cA bys sn ele 
Pteroglossus Chaudoir, 1843 .. ais vi ia a" on i tha ee 
Ptilorhis Agassiz (J.L.R.), 1846 xe a a te ce ee ci: Melee 
Ptilorhys Bonaparte, 1854 and BE ne me a es ~ andy Ag een 
Ptilornis Gray (G.R.), 1869 nh ae aa ny ~ he oe ate, speed 
Ptychognathus Owen, 1860 ae Ml me Ae ye Ae dis a 61 
Siren Hermann, 1783 sd a! Be 3 ngs a! ngs mt 346 
Sirena Fischer de Waldheim, 1808 oy. at m3 Se t: ae ei 346 
Sirene Oken, 1816 ; Vis By. ate is, ae Bx rv 346 
Sphargis Merrem, 1820 Ae aps mA a rs oe es a 346 
Strigopsis Bonaparte, 1849 ne mes oe ee By ae se Stein ees 
Stringops Finsch, 1867 ef = ie a3 ah oe <tieep ene 
Stringopsis van der Hoeven, 1855 45 ai oe = ro a cacent deli 
Strygops Des Murs, [1853] te bl u ‘4 Res st cc Shit sakes 
Syngnathus Rafinesque, TS1D Vit, oe Pe is ne Ne on be 346 
Telmessus Stal, 1866 ah & ay a uN a a8 33 io 61 
Teuthys Lamarck, 1790 sf ny ve te a ae aye Pome 
Tlos White, 1847 it ne oe yi! wa x4 “ic *e 61 
Trichodactylus Dufour, 1838 whe ae fis ate ue we cS 61 
Utica Hewitson, [1865] . ie wy a Sis i ae a? ) 61 
Valdivia Ragonot, 1888 .. ie me; Py a. se oe x ae 61 
Valdivia Shannon, 1927 . yy ave ae a he + 1% Na 61 
Zosimus Schaufuss, 1886 te be ne BG at an be 61 
Zozymus Milne Edwards (H.), is34° Aes abe he 2. ss id 61 
Official Index of Rejected and Invalid Specific Names in Zoology, names placed on : 

alleghaniensis Harlan, 1825, Abranchus * ae as te . 5 yang SLE 
annulicornis Leach, [1815], Pandalus .. oe ¥ ce tat as 2 a pe ee 
argus Temminck, 1807, Polyplectron .. Bf ie ie ar: es oa phat 
chalybea Boddaert, 1783, Manucodia .. 2 Ed, ~~ I ms Jina Lge 
chalybeata Stiles, 1916, Paradisea u a ai ~ = eo Skink ele 
communis Bonnaterre, 1791, Coturnix os aa spe oe oe pe 219 
fluvialis Savigny, 1809, Pandion. . ate sr ne gl ae a are ees 
formosus Latham, 1790, Psittacus oe he & ais a a Sot hehe 
fulicarius Bonnaterre, 1790, Heliornis PA “2 ie we i. eae et 
galbula Linnaeus, 1766, Oriolus : 3 a oh me 7 on: Meee 
granulatus de Haan, [1835], Cancer (Actaea) AS i 7 so ae .. 61-62 
griseus Nilsson, 1821, atau als Ab at zi * 5c eee 
Jurinii Bell, 1855, Leucosilia sia : + My che i - 3 62 
marinus Fabricius, 1775, Astacus ae ax af Ne Ae we pene 22) 
media Koch, 1816, Gallinago ro] Me Me: a Re ‘te ies. ee 98 
mercurialis Merrem, 1820, Sphargis  .. owe Eis aa LE pat ae 346 
pennanti Leach, [1817], Ebalia es nie ie a Mh ee be 62 
psophia Pallas, 1767, Grus : Sp oe At ow Ps Semen 6 | 
Savignii Milne Edwards (H.), 1834, Cancer .. as ae a aes ae 24 
tresselata Milne Edwards, (H.), 1834, Lybia .. ste ae ae ie mn 62 
vigil Fabricius, 1798, Portunus : Es =f Zz wes a 62 


vulgaris Milne Edwards (H.), 1837, Homarus’ ne hs ba oe os) eo 


Volume 1, Section D 437 
Page 
Official Index of Rejected and Invalid Works in Zoological Nomenclature, titles of 
works placed on : 
[Tunstall (M.)], 1771, Ornithologia britannica 85 
Official List of Family-Group Names in Zoology, names placed on: 
DOLICHOPODIDAE (correction of DOLICHOPODES) Latreille, 1809 268 
Official List of Generic Names in Zoology, names already placed on, corrections and 
additions to entries concerning, exclusive of determination of gender of names 
placed on: 
Acipenser Linnaeus, 1758 oe 339 
Acmaeopleura Stimpson, 1858 .. 50 
Actaea de Haan, [1833] .. =, 
Actumnus Dana, 1851 55 
Alauda Linnaeus, 1758 .. 213 
Asthenognathus Stimpson, 1858 50 
Atergatis de Haan, [1833] ao 
Atergatopsis Milne Edwards (A. ), 1862. 55 
Bathynectes Stimpson, [1871] .. re 54 
Bathyplax Milne Edwards oe L880 <. 50 
Blennius Linnaeus, 1758 P ve 341 
Calamaria Boie, 1826 342 
Callionymus Linnaeus, TPIS: | ss 339 
Camptandrium Stimpson, 1858 . 50 
Camptoplax Miers, 1884 50 
Caphyra Guérin-Meéneville, 1832 54 
Catoptrus Milne Edwards (A. ), 1870 50 
Ceratoplax Stimpson, 1858 ey 50 
Chasmagnathus de Haan, [1833] 50 
Chasmocarcinus Rathbun, 1898 49 
Chimaera Linnaeus, 1758 339 
Chlorodopsis Milne Edwards (A. y 1873. 53 
Clistocoeloma Milne Edwards (A.), 1873 50 
Clupea Linnaeus, 1758 a A Ls ses 340 
Coryhaena Linnaeus, 1758; correction of, to Coryphaena ht Bs a 341-342 
Coryphaena Linnaeus, 1758 : oe ae 340 
Coturnix Bonnaterre, 1790 181 
Crotalus Linnaeus, 1758 341 
Cyclodius Dana, 1851 .. 53 
Cyclopterus Linnaeus, 1758 339° 
Cymo de Haan, [1833] 56 
Cyprinus Linnaeus, 1758 340 
Cyrtograpsus Dana, 1851 50 
Dermochelys Blainville, 1816 342 
Desmognathus Baird, (1850)... 341 
Dissodactylus Smith (S.1.), 1870 50 
Durckheimia de Man, 1889 49 
Ebalia Leach, [1817] 53 
Eleotris Schneider, 1801 342 
Epixanthus Heller, 1861 50 
Esox Linnaeus, 1758; 341 
Euchirograpsus Milne Edwards (H. ); 1853 50 
Eucrate de Haan, [1835] ; 50 
Eucratodes Milne Edwards (A. ), 1880 50 
Eucratopsis Smith (S.1.), 1869. 50 


438 Opinions and Declarations 


Page 
Official List of Generic Names in Zoology, names already placed on, corrections and 

additions to entries concerning, exclusive of determination of gender of names 
placed on (contd.) : 
Euryetisus Cano, 1889 .. re as ch. ye at <% z rae 49 
Eurynorhynchus Nilsson, 1821 .. sie wh Eee ve Pi iy - 181 
Euryplax Stimpson, 1859 Ae x = hes ies a. do. LA 50 
Eurytium Stimpson, 1859 7; se ba ae ae os 4 a: 49 
Fabia Dana, 1851 Ae 2 = ee ef a =e ck a 50 
Fulmarus Stephens, 1826 on ae see oa oe nS OF ine ope 
Gadus Linnaeus, 1758 .. Ap Se ‘i + sf ae ee ee 340 
Galene de Haan, [1833] .. ae ee an ae 's x a ae 50 
Gasterosteus Linnaeus, 1758... aes ~~ ak + sf Ls “ 340 
Geryon Kr@yer, 1837... a avs ae 2 o ae Pe 49 
Glyptograpsus Smith (S.I.), 1870 £. Ris ies ve aA es si 49 
Glyptoplax Smith (S.I.), 1870 .. ‘a ie oh ve ” a a 50 
Gobius Linnaeus, 1758 ay ig “8 a ste a bt - 340 
Gomeza Gray (J.E.), 1831 es ae Bat Sd a a aa y 50 
Goneplax [Leach], [1814] ae fe a zt a es xe a 49 
Halimede de Haan, [1835] =e Ae “ a “a ‘y she ee 5k 
Helice de Haan, [1833] .. we et Pe Re - ay m eg 51 
Heliornis Bonnaterre, 1790 De me ats at ae - re 181, 213 
Hepatella Smith, 1869 .. as on ~ af a te ae oe 54 
Hephthopelta Alcock, 1899 #: she oe 4 ahd = ws av 
Hexapus de Haan, [1833] : a € fe - oe ae | 
Holometopus Milne Edwards (H. a 1853 iat aH EH wah Ad te 51 
Holothuriophilus Nauck, 1880 .. : at am je * a Ne 49 
Homalaspis Milne Edwards (A. )s 1863. Ae A na ae a% ery 8 
Homarus Weber, 1795 .. : mA me as bh oh Aare See 
Tliacantha Stimpson, [1871] ; ak op ax ne sg ¢ if 33 
Iphiculus Adams & pt [1849 He i She vs des ss zi 56 
Iphis Leach, 1817 ¥: te ae a at 3% uke em 
Lacerta Linnaeus, 17158... me Si os i2 as ne a2 ses St 
Lachnopodus Stimpson, 1858 ne ae a e 3 os oY. 51 
Leptodius Milne Edwards (A. ds 1863... oe a a sik y, ke af 
Leucosilia Bell, 1855 13 ee Af ee Le . me $y 55 
Liagore de Haan, [1833].. mi ae be x - oo 5 51 
Libystes Milne Edwards (A.), {S67 ae are a ue tb ae 51 
Liomera Dana, 1851 Be zp me Le ae ee we wh 51 
Lipaesthesius Rathbun, F898 2 y [2 y - J a ve Si 
Lissocarcinus Adams & White, (1849) oe ahs ie 9: sie bo 56 
Lithadia Bell, 1855 ; : ne Send a; ne Ee he 55 
Litocheira Kinahan, 1856 bt Be ae ae sa) as Le va 51 
Lophius Linnaeus, 1758 ss ce a wi Re 8 a oon 340 
Lophopanopeus Rathbun, 1898 .. an ee = an oa a a 49 
Lophopilumnus Miers, 1886 ae es s ea a. de si 49 
Lupocyclus Adams & White, [1849] .. eG or Ae — a a 56 
Lybia Milne Edwards (H.), 1834 i Ae ee ‘8 He As ci 51 
Manucodia Boddaert, 1783 xe, f. ce a s iy it ve 181 
Melybia Stimpson, [1871] : ee te a Ms % $3 2) | 
Metasesarma Milne Edwards (H. Y 1853 be ne * a3 ie ee 51 
Metopocarcinus Stimpson, [1871] nhs oe Lt ae es A td a1 
Micropanope Stimpson, [1871] .. Ze se a a "a ft a 51 
Mormyrus Linnaeus, 1758 av of a + by im ne Re ate 
Mullus Linnaeus, 1758 ie 5 ees ay a ae. Ins 340 
Myrodes Bell, 1855 Pi : et 7 ae Se Me nd wis 55 
Nephrops [1 each, [1814] st ays as fe bh $. we 57 
Notonyx Milne Edwards (A.), ce7gc Si an as e a S1 
Nursilia Bell, 1855 4 on i at me tk ft id 55 
Oediplax Rathbun, 1893 oe on = on $6 Le ne is 51 
Ommatocarcinus White, S52 ee at Ds a bs ae ce = 51 


Ophidion Linnaeus, 1758 a, at bs ee zs - ey bg 341 


Volume 1, Section D 439 


Page 
Official List of Generic Names in Zoology, names already placed on, corrections and 

additions to entries concerning, exclusive of determination of gender of names 
placed on (contd.) : 
Opisthopus Rathbun, 1893 ey ss y ; m Ss pe at 51 
Oriolus Linnaeus, 1766 .: Si a if “« e, %, ff Li 181 
Orphnoxanthus Alcock, 1898 .. ate ae a ny ne ve ar 51 
Pandion Savigny, 1809 .. uP aN a3 he a’ #9 ce mS 181 
Panoplax Stimpson, [1871] my Hes Ae a 3 a be ats 51 
Paragalene Kossmann, 1878 .. Fa iat af nu! ar by, a, 51 
Parapanope de Man, 1895 oh a2 Bsr Lge a fe a: +, 51 
Parapleurophrycoides Nobili, 1906 ; ne Me i st mit 51 
Paraxanthus Milne Edwards (H.) & Lucas, [1844] - ae “ig an .ic) SES 
Pariphiculus Alcock, 1896 ne rt Et ee He a4 53 
Perca Linnaeus, 1758 er oe ee site a oe ae. cs ae 340 
Percnon Gistl, 1848 Be af = te chs ch 7 me rl 52 
Perigrapsus Heller, 1862 AS as Oe on Fal a8 Rs at 52 
Persephona Leach, 1817. i 3 ae ae a Py a ty 53 
Phlyxia Bell, 1855 ng ats ad £Y, 2 Dao 
Pilumnoides Milne Edwards (H. ) & Lucas, [1844] ce a vp: ” oi SRST 
Pilumnus Leach, 1815... eee a ve st i 2 
Pinnaxodes Heller, [1865] he " oes Eis ae a 1 ve 50 
Pinnixia White, 1846 . oe Y sz Ea ae 50 
Pinnotherelia Milne Edwards (H. ) & Lucas, [1844] oe aa ms ee ac hoe eT 
Planes Bowditch, 1825 .. , DF = a; ‘ig Ae 52 
Platychirograpsus de Man, 1896 ay: ct “es ie 4, 4% Ay: 50 
Platypilumnus Alcock, 1894 ; be ae ae ot He a 50 
Platyxanthus Milne Edwards (A. ), 1863 as 2 rat ee on i 52 
Polydectus Milne Edwards (H.), 1837 . a) 7 ae Pcl se 32 
Polyplectron Temminck, 1807 .. at Bre! * re ar a 182 
Prionoplax Milne Edwards cE.) 1852 . oh a 4 eh - - 24 
Pseudocarcinus Milne Edwards os y ic) ee Ae - ae Bi: ah 3 
Pseudophilyra Miers, 1879 : oe an a = ats ee 53 
Pseudopinnixa Ortmann, 1894 .. i ae oe Be a ar 52 
Pseudorhombila Milne Edwards (TA.), 137 e We i it, $% 49 
Psopheticus Wood-Mason, 1892 a ae sc x x te 52 
Ptychognathus Stimpson, 1858 Ay pe of ye at = 52 
Pyxidognathus Milne Edwards (A.), 1879 ae oh. sf i Pe ae SZ 
Rhithropanopeus Rathbun, 1898 ae 2g ie - mn a4 a. 52 
Rhizopa Stimpson, 1858 a : a Rr. “e fe: 52 
Ruppellioides Milne Edwards (A. ), 1867 : 7 <i 22 + = 52 
Salmo Linnaeus, 1758 .. fe v re Bs 23140 
Samaritium Dana, tsb : i we a Se vi 33 a a 52 
Scalopidia Stimpson, 1858 at an ee ae: ie ae bag 52 
Scleroplax Rathbun, 1893 ce +. ve ay ‘a a 4. vs 52 
Scomber Linnaeus, 1758.. a es si av 7 we ie ea 340 
Scorpaena Linnaeus, 1758 eg te & ms ry se i, 7 340 
Scylla de Haan, [1833] .. < ye ae Ce Kt if 2 di 56 
Silurus Linnaeus, MISS) G: Re ae = be ee LS ig oo 
Speloeophorus Milne Edwards (A. Je 1865 20 is ol se . Mave DS 
Speocarcinus Stimpson, 1858  .. te a i <a 3 Lied 52 
Sphaerozius Stimpson, 1858 __.... ee a ts a se a aes 2 
Syngnathus Linnaeus, 1758 ae 3 if at is be ae a 340 
Tetraxanthus Rathbun, 1898 .. es ahs nt ote ae ae oe: a2 
Tetrias Rathbun, 1898 .. ba te a ~~ ae 7 52 
Thalamitoides Milne Edwards (A. Vs 1869 +f i a. oF se A 54 
Thalamonyx Milne Edwards (A.), 1873 we ah 1% HE ae Y: 54 
Thaumastoplax Miers, 1881 Se ; ed “ae Be ag 7 ae 50 
Tlos Adams & White, 1849 a) a ne oe be J Ar als 56 
Trichodactylus Latreille, ae i ee Ae j2 te a a8 56 


Utica White, 1847 a ; ~ iY i ae a ” t 52 


440 Opinions and Declarations 


Page 
Official List of Generic Names in Zoology, names already placed on, corrections and 
additions to entries concerning, exclusive of determination of gender of names 
placed on (contd.) : 
Varuna Milne Edwards (H.), 1830 a ov; ne af a a a 2 
Xanthasia White, 1846 .. ia ef x ae a ae we 52 
Xanthodius Stimpson, 1859 Ais tie es ee oe on A 53 
Xenophthalmodes Richters, 1880. ae ~ ue - Se - taf 50 
Xenophthalmus White, 1846 ie ae te ve ae a as of 53 
Zeus Linnaeus, 1758 ws af oe a sn ae ag a i 340 
Zosimus Leach, 1823 ae oe af a Be rs & ee in 53 
Zozymodes Heller, 1861 .. Se me Mie Ps om oe he ae 52 
Official List of Generic Names in Zoology, names already placed on, determination 
of gender of : 
Agasoma Gabb, [1869] .. Ma ny a sh se As i ah 159 
Amblyomma Koch, 1844 x a5 * re rhe a 2 at 144 
Ancylostoma Dubini, 1843 ae a A ae # ae = oe SS 
Anodonta Lamarck, 1799 we an xe We a a = uh 158 
Anoplocephala Blanchard, 1848.. os ze oF ra A ass as 157 
Antedon Fréminville, 1811 oe A i Ay 9: . a fe 158 
Anthocoris Fallén, 1814 .. a < ie a ue ie a an 141 
Arcella Ehrenberg, [1830] se ft ity ea wy ae a ros 156 
Argas Latreille, 1795 Pe ak = a a ns re bi ae 142 
Argonauta Linnaeus, 1758 4 A; 5% Aa # Ie 156 
Armadillidium Brandt & Ratzeburg, 1831] an A, fe i. a af 143 
Ascaris Linnaeus, 1758 .. .& A me a ad 1 1S7 
Avicularia Lamarck, 1818 * oa ah ee # aed as seg 143 
Bathycrinus Thomson, O72 He & * rt se te ut A; 156 
Blatta Linnaeus, 1758 .. a Sh £% Ms a my a ZY 143 
Botryllus Gaertner, 1774 ae fe iy re re Hi as oy 156 
Buccinum Linnaeus, 1758 e ¥ he 2: ad, if) eho ae 159 
Bursaria Miller, 1773... fos = a oe 4 Es # f, 156 
Calyptraea Lamarck, 1799 oe sy ss Le Lt it ii ae 157 
Cephenemyia Latreille, 808° ee: #4 " me As ay ef ae 143 
Cerion Réding, 1798 me me ot a fe a * th - 159 
Cheyletus Latreille, 1796 - in Hi ie. fe a 142 
Chorioptes Gervais & van Beneden, 1859 ie be id = ats a8 142 
Cimex Linnaeus, 1758 .. : ak a ae ig * a te 141 
Clausilia Draparnaud, [1805]. 4 at 3 - = Xs #e sf 158 
Clavelina Savigny, 1816 .. =f ch a5. aA a: ine 4 x 158 
Columbella Lamarck, 1799 % a os ae 4 H ae es: isi) 
Cryptobranchus Leuckart, 1821 = = ‘% te bt ae of 2s 
Davainea Milne Edwards ahh 1869 =. of = sid a as 4 LF 
Demodex Owen, 1843... sie fs ote i a 142 
Dendryphantes Koch, 1837 ss ay oe: af by kg As A 142 
Dentalium Linnaeus, 1758 oy “e is ns = ae on ea 159 
Dermacentor Koch, 1844 Cy we ite Ae ve as Ee 142 
Dermanyssus Dugeés, 1834 8 an 7 sa ie zd fe in 142 
Diaptomus Westwood, 1836 _.... Mi be ss :, ats Fi ae 142 
Diazona Savigny, 1816 .. e He at os nf ee ve an 158 
Dicrocoelium Dujardin, 1844 .. £4, .e - ¢ A ae jy 159 
Dipylidium Leuckart, 1863 = si a ae ‘- Ae om wi 159 
Distaplia Della Valle, 1881 Re: 5 ae ok of > <4 a 158 
Dracunculus Reichard, 1759... vs ie = A + yh: Py ps) 
Dysdera Latreille, [1804] - Y ee =e = at 2 i 143 
Echinococcus Rudolphi, 1801 .. “f ee, * fs aA aN A E55 


Ectobius Stephens, 1835 aN = oti ft “Me ie x a 141 


Volume 1, Section D 441 


Page 
Official List of Generic Names in Zoology, names already placed on, determination 

of gender of (contd.) : 
Eimeria Schneider, 1875 48 a. e ga wig oe we fee 156 
Enchytraeus Henle, 1837 xe i ie “¢ gf ue ae " 155 
Eremias Wiegmann, 1834 43 a3 ar <2 sk a kate sl: 
Fasciola Linnaeus, 1758 vd 3 oe 24 af a * oF 157 
Filaria Miller, 1787 oe ns ey £3 ae oF 3 ae e: $57 
Forficula Linnaeus, 1758 a #2 = ue’ os ie 7" - 143 
Gammarus Fabricius, 1775 oe Bs: oe se he ch. Ay uM 142 
Gasterophilus Leach, 1817 x ct Be a 5% 7, & e 141 
Gastrodiscus Cobbold, 1877 4. nf 2" 45 Ao a a ae {55 
Glycyphagus Hering, 1838 sod : of a ie fe Rs ¥ 142 
Gnathostoma Owen, 1837 i." ae a én ~ a3 be iv 159 
Gordius Linnaeus, 1758 .. Pe ne Bet me bi haf a a4 [55 
Grus Brisson, 1760 a ma 3 Be: i Ba et oe oe 521 
Gryllus Linnaeus, 1758 .. nS a a xa Be La a ae 141 
Haemadipsa Tennent, 1859 a Ps a a ee o a? uy 57 
Haemaphysalis Koch, 1844 ae fe ig a4 Fi Ae - 143 
Helix Linnaeus, 1758 __—... is i a a he Se & i iy 
Hemiurus Rudolphi, 1809 es - 2 ats A: oF me De 155 
Heteroda Schmidt, 1871 Ae ee “in a a, ar af . | Is i 
Heterophyes Cobbold, 1866... Ee He fi ie “ Ad e: £55 
Hirudo Linnaeus, 1758 .. oe Pa “A bs ve: ie re ilayy 
Holopus @Orbigny, 1837 ys oe at 38 ea hae Hi Be 156 
Holothuria Linnaeus, 1767 2 Bi ae - 24 be ae 158 
Homalaspis Milne Edwards (A), 1863. oe Z% ve ‘ae "0 285 
Homarus Weber, 1795 .. oa oe Be ~ es ne 295 
Hyalomma Koch, 1844 .. ef ae = a hgh * it fe 144 
Hydra Linnaeus, 1758 .. Ay Re ing 2s a be Sa % 157 
Aymenolepis Weinland, 1858 .. es a ay ie ., aA a 157 
Hypoderma Latreille, 1818 e ae ds 2 ie ~ 4 ik 143 
Iphis Leach, 1817 ee, 2.4 Pe: ae a af Pe ay 267 
ixoaes watreille, 1795 .. = a he oi ue Le ix ie 142 
Latrodectus Walckenaer, 1805 .. oe A a we a Me ge 142 
Lepas Linnaeus, 1758... e: i. me 98 am z. ae ~- 143 
Lepidocyclina Gimbel, [1870] .. a Pee si a as a % 156 
Leucochila von Martens, 1860 .. he a wie fe aS #4 » 157 
Ligula Bloch, 1782 a - a she oe 2 2 = BA 157 
Limax Linnaeus, M58). 3.3 a 2 a & a =e 156 
Limnatia Moquin-Tandon, 1827 ee i = ae an A ye 157 
Limulus Miller (O.F.), 1785... Par it ea a i i 142 
Lithostrotion Fleming, 1828 ee ae ye on : ae ct Stes 2) 
Luidia Forbes, 1839 wis Me aa ae oe a ae i: 158 
Lumbricus Linnaeus, 1758 ie Ae ee if Le * at m ip) 
Lytoceras Suess, 1865... a rs ae ee oe ws hs ve 159 
Mactra Linnaeus, 1767 .. a ae ot ae af Le es ae 158 
Metacrinus Carpenter, 1882... oe Be ui ak ae ms ae 156 
Molgula Forbes, 1848 .. oo oe “ae a oa oe ae ae 158 
Moniezia Blanchard, 1891 ae ay oe x ae a on ae 157 
Musca Linnaeus, 1758 .. ae ee ae a an af “tr ne 142 
Mya Linnaeus, 1758 a he He “ie Ps I a a ea 158 
Mytilus Linnaeus, 1758 a ee ae oe ne: ay =a We 156 
Nabis Latreille, [1802—1803] .. cig a by i! iz ae 141 
Neritina Lamarck, 1816 Ad ee ny oe nA ee ee Re 158 
Notonecta Linnaeus, 1758 - € ee obs oe ie ty: 343 
Nycteribia Latreille, 1796 ae os at ot ye e dt; es 143 
Oedemagena Latreille, 1818 ae a: aa ae ae ee ie vt 141 
Oestrus Linnaeus, 1758 eh a re Dt i ib Ag 141 
Oleacina RGding, 1798 .. = a ae i ve ey: ey: x 158 


Oniscus Linnaeus, 1758 .. <i of a ye at ES att a 142 


442 Opinions and Declarations 


Page 
Official List of Generic Names in Zoology, names already placed on, determination 
of gender of (contd.) : 
Otarion Zenker, 1833 .. an Ba ee ee bs, af = 144 
Paragordius Camerano, 1897 .. =a me ae ie 3 4: ‘e 155 
Pediculus Linnaeus, 1758 se rf a x = sg ~ a 141 
Periplaneta Burmeister, 1838  .. “e Be et be . ne i 143 
Phthirus Leach, 1815 ms a ms os ait at iss ie ome 141 
Physa Draparnaud, [1801] ae a ye c ss £: es a 157 
Physalia Lamarck, 1801 ae se 4 ie 3 A ae ae 157 
Podura Linnaeus, 1758 os a aa aA ext on st9 ote 143 
Polydesmus Latreille, [1802—1803] at ue ae og a me a 141 
Porcellio Latreille, (1802—1803] ait ne ay. ake ve - a 141 
Psoroptes Gervais, 1841 : or ay: ae Ze Ae 5 a 142 
Pulex Linnaeus, 1758 _... one 8 2% ae ae a es = 141 
Pyrosoma Peron, 1804 .. i Sa , oe a3, Be ay a 159 
Reduvius Fabricius, 1775 ak e% - xe ‘* = Aa 5% 141 
Rhabditis Dujardin, 1845 ae Ee -e ve on a3 157 
Rhipicentor Nuttall & Warburton, 1908 a sh .% she ae EA 142 
Rhipicephalus Koch, 1844 oe = ae by me sh oa 142 
Rhizocrinus Sars, [1865] a we 2 Be ste ve at 156 
Rhizoglyphus Claparéde, [1869] me He Aye J an a ae 142 
Sarcocystis Lankester, 1882 - ae es oe 24 ns oe 157 
Sarcoptes Latreille, [1802—1803] fe it ne >» re ahd oe 142 
Schistosoma Weinland, 1858"... sts ee se a i a Les 159 
Scorpio Linnaeus, 1758 ie a a Be oa ae “ a 142 
Segestria Latreille, [1804] . - a # i re Kf a 143 
Sepia Linnaeus, 1758... oe ae oh Be af ay i 2 158 
Sphaerium Scopoli, 1777 we oe re pa ae ia - nb 159 
Spinturnix Heyden, 1826 _ ze 7 fe it s _ as 143 
Spirifer Sowerby, 1816 .. ee - of Ae mA a Es as 156 
Stilesia Stiles & Hassall, 1893 .. 7s a =f ia uy 3 ae LF 
Strongyloides Grassi, 1879 ve Fi an Ue afte if 48 x 1595 
Strongylus Miller, 1780 ou i - oe Se igs 3 i 155 
Succinea Draparnaud, [1801] .. ae Ap “i Ae Ed ep iV: 157 
Syngamus Siebold, 1836 un i ee ae ate Se ae is £535 
Syringothyris Winchell, 1863... ee sa Ba, as me nee we 158 
Taenia Linnaeus, 1758 .. As ee xe ae 2 5903 ji ay 137 
Teredo Linnaeus, 1758 .. oe ae oe fe o go re shy 158 
Thysanosoma Diesing, 1835 we a oe e e- a oi Sie 159 
Tornatellina Pfeiffer, 1842 ae = af, ris es oe a oa 158 
Triatoma Laporte, [1832] a a oe 8 ye 25 me Re 143 
Trichostrongylus Looss, 1905... We st ls . Sis e a 155 
Trombidium Fabricius, 1775 _.... oF ae $2 te =e ian ae 144 
Trypanosoma Gruby, 1843 a Aa ne - e ae at sis 158 
Vitrina Draparnaud, [1801] xe ait Se es a 7 ts oh 158 
Volvox Linnaeus, 1758 .. = om as A u - se fo 156 
Official List of Generic Names in Zoology, names placed on: 

Columba Linnaeus, 1758 ae ae ae he 2 as ar #3 86 
Dolichopus Latreille, 1796 a a of * ae A om oo (ee 
Gallinago Brisson, 1760 .. x <2 ge She he Ae x ats 97 
Grus Brisson, 1760 Aes Ld a a ae AA en e a 321 
Omalaspis Giraud, 1860 ae a ke - ee oe a “0 
Pandalus Leach, [1814] ¥e oS - ae = ae sie ai 245 


Pinnotheres Bosc, [1801—1802].. ah ae we na NE os eK 235 


Volume 1, Section D 443 


Page 
Official List of Generic Names in Zoology, names removed from : 
Gallinago Koch, 1816 ; ate =k es i J, i zy 97 
Grus Pallas, 1767 [recte 1766] . zs ee a oF xs 321 
Pandalus Leach, 1815 (Trans. linn. Soc. Lond. ) ge e3 ahs ts 245-246 
Pinnotheres Latreille, [1802—1803] ae he Pm di he 235-236 
Official List of Specific Names in Zoology, names placed on: 

abdimii Lichtenstein, 1823, Ciconia .. ae io ne ie a ~ 180 
aculeatus Linnaeus, 1758, Gasterosteus ae He oe Ne ail 370 
acus Linnaeus, 1758, Syngnathus i mel wh Y an a sist ones 
admete Herbst, [1803], Cancer e sie) ae Ht Ly, ie ro 11 
aeneus Linnaeus, 1758, Cancer . a ne at ae dae se st 22 
aethereus Linnaeus, 1758, Phaethon . , ve ae x. ge re 178 
agassizii Milne Edwards (A. ), 1880, Eucratodes = 1s uy i 98 14 
agilis Linnaeus, 1758, Lacerta .. ais ot of uf oe pee 
agilis Milne Edwards (A.), 1872, Hydrothelphusa a ey S3 a 3 7 
albicilla Linnaeus, 1758, Falco : - a Me sa - 183 
alcocki Wood-Mason, 1891, Paria <3 ie 20 ft iz + de 9 
alleganiensis Daudin, [1803], Salamandra Nye ed a. ne et “eis 374 
americana Saussure, 1857, Pseudothelphusa .. Ke ie of a ve 10 
americanus Linnaeus, 1758, Cuculus .. Ne a te ee a ce 176 
amica Smith (S.1.), 1869, Hepatella .. ae at a3 “ ste - 7 
andreossyi Audouin, [1826], Pilumnus .. sa Re a ti aus ss 6 
angulatus Dana, 1851, Cyrtograpsus .. 5. he se As ae ae 13 
ani Linnaeus, 1758, Crotophaga ee a a 7 ees 7 176 
anomalipes Miers, 1881, Thaumastoplax * ne tt : eat ae 21 
aquaticus Linnaeus, 1758, Rallus or: i bl ia oh - ee i. 
aracari Linnaeus, 1758, Ramphastos PA su a" e ug = oe 179 
ardeola Paykull, 1805, Dromas .. ore ae * aT - bos 176 
armata Milne Edwards (A.), 1869, Banareia .. a, - ie: a a 4 
armatus Milne Edwards (H.), 1853, Potamocarcinus a af e 10 
arvensis Linnaeus, 1758, Alauda a fe we o a + ne 174 
aura Linnaeus, 1758, Vultur.. 5 Se 4 ee ~ ay ae LIS 
auratus Linnaeus, 1758, Cuculus oe my tid ms ge ie $e 176 
auritus Gray (J.E.), 1829, Podargus .. ifs at - te * ss 7s 
australis Shaw, 1813, Apteryx me : a nes Ne a - =A 174 
avosetta Linnaeus, 1758, Recurvirostra a Me sd si — 179 
balansae Milne Edwards (A.), 1873, Clistocoeloma ee v7 2 se 13 
barbatum Linnaeus, 1758, Ophidion ad fa of v ys cc een 
barbatus Linnaeus, 1758, °"Mullus oe _ ae as ¥ a an 370 
barbiger Poeppig, 1836, Gecarcinus 4s be, tye ae ee He me 23 
bedoti Zehntner, 1894, Sphaerocarcinus a ae mt Af Me? ie Na 
bella Stimpson, 1860, Xantho .. PN: ce a a ie sé e 16 
bicalcaratus Linnaeus, 1758, Pavo a me ais ae at te i 183 
bicornis Gray (J.E.), 1831, Gomeza te i a a: ais - 15 
bidentatus Milne Edwards (A.), 1880, Xanthodes x: uf if - ee 21 
bispinosa Kinahan, 1856, Litocheira .. re a fe fs af 16 
bispinosus Herbst, [1783], Cancer a aA ie m os a Ay 14 
cabrilla Linnaeus, 1/585 Perea. ..; ite ae he a5 s. ee Pe 373 
caeruleus Desfontaines, [1789], Falco .. “3 2 Pe “ ¥e cx 183 
carinata Ortmann, 1894, Pseudopinnixa a 4 bas a Be 5 20 
carinipes Heller, 1861, Zozymodes a en x Be ib si at 22 
carinipes de Man, 1889, Durckheimia es oe i cf -_ ty 13 
carolinensis Stimpson, 1859, Speocarcinus Ae =. . ae se ty: 21 
carpio Linnaeus, 1758, Cyprinus hte gi re * rs a4 ne 369 
cephalus Linnaeus, 1758, Mugil ne =) a af 4; Ee 368 
cessacii Milne Edwards (A.), “878, Epimelus as we oe ey se ds. 6 


chalybata Forster, 1781, Paradisea [sic] 7% " of hs .. 182 


444 Opinions and Declarations 


Official List of Specific Names in Zoology, names placed on (contd.) : 


chavaria Linnaeus, 1766, Parra 

cheiragonus Tilesius, 1815, Cancer ; 
chilensis Milne Edwards (H.) & Lucas, [1844], Conystoides: 
chimaera Rothschild, 1895, Uratelornis fe ; 
chrysocephala Lewin, 1808, Meliphaga 

ciliatus Stimpson, 1858, Ceratoplax 
cinctimanus White, 1849, Carpilius 

cinerea Linnaeus, 1758, Ardea .. 

convexus Milne Edwards (A.), 1867, Ruppellioides 
coppingeri Miers, 1884, Camptoplax 

corallinus Faxon, 1893, Trachycarcinus 
coriacea Linnaeus, 1766, Testudo 

coronata Alcock & Anderson, 1894, Randallia 
coturnix Linnaeus, 1758, Tetrao 

crassimanus Dana, 1851, Eucrate 

crassipes Bell, 1855, Phiyxia 

crassum Dana, 1851, Samartium 

crenatus de Haan, [1835], Cancer (Eucrate) 
crepitans Linnaeus, 1758, Psophia 

cribrarius Lamarck, 1818, Portunus 

cristata Pallas, 1764, Columba sty 

cristatus Shaw in White, [August] 1790, Caprimulgus. 
cubensis Stimpson, 1860, Epilobocera .. ‘ 
cumingii Bell, 1855, Lithadia ; 

cupulifer Latreille, 1825, Pilumnus 

cylindricum Say, 1818, Pinnotheres 

cylindrus Fabricius, 1777, Cancer 

cyprinoides Linnaeus, 1758, Mormyrus re 
danae Milne Edwards (A.), 1869, Goniosoma 
deplanatus Cano, 1889, Euryetisus — 9 
depressa Stimpson, [1871], Panoplax 

dentata Bell, 1855, Nursilia 

denticulatus Montagu, 1806, Cancer .. 
dilatipes Adams & White, [1849], Pilumnus 
doliatus Linnaeus, 1764, Lanius. . ‘ 

dovii Stimpson, 1860, Euphylax 

draco Linnaeus, 1758, Trachinus 

electricus Gmelin (UE F. L789 |: Silurus 

elegans Gmelin (J.F.), 1788, Psittacus . 

eremita Nobili, 1906, Dacryopilumnus 
erinaceus Fabricius, 1787, Cancer 

erithacus Linnaeus, 1758, Psittacus 

erosa Miers, [1877], Actaeomorpha 

euagora de Man, 1895, Parapanope 

eudactylus Bell, 1855, Myrodes 

europaea Linnaeus, 1758, Sitta 

exaratus Milne Edwards (H.), 1834, Chlorodius 
excelsus Heller, 1862, Perigrapsus 

excubitor Linnaeus, 1758, Lanius 

exulans Linnaeus, 1758, Diomedea 

faber Linnaeus, 1758, Zeus se 

fallax Henderson, 1893, Ebalia 

flaveola Linnaeus, 1758, Certhia 

fluviatilis Linnaeus, 1758, Perca 

fluviatilis Latreille, [1828], Trichodactylus 
fragifer de Haan, [1835], Cancer (Halimede) 
frontalis Milne Edwards (H.), 1834, Ozius 
fulica Boddaert, 1783, Colymbus 

fuscus Rafinesque, 1820, Triturus 


Volume 1, Section D 


Official List of Specific Names in Zoology, names placed on (contd.) : 


gallinago Linnaeus, 1758, Scolopax 

gammarus Linnaeus, 1758, Cancer 

garzetta Linnaeus, 1766, Ardea at 
gaudichaudii Milne Edwards (H.), 1837,  Platymera x re 
gayi Milne Edwards (H.) & Lucas, (1844), pi actin aan ae 
gigas Lamarck, 1818, Cancer nt 
glaber Stimpson, 1858, Ptychognathus .. 

glacialis Linnaeus, 1761, Procellaria 

gladius Linnaeus, 1758, Xiphias 

glandarius Linnaeus, 1758, Cuculus 

glanis Linnaeus, 1758, Silurus .. 

glutinosa Linnaeus, 1758, Myxine 

gracilipes White, 1847, Utica 

gracilipes Stimpson, 1858, Rhizopa 

gracilipes Alcock, 1894, Platypilumnus. . 

granulatus Rathbun, 1893, Oediplax 

granulatus Rathbun, 1893, Scleroplax .. ; 
granulosus Milne Edwards (A.), 1879, Pyxidognathus 
grus Linnaeus, 1758, Ardea Bi : 
habroptilus Gray iG: R.), 1845, Strigops : 
haematocheir de Haan, [ 1835], rhe is (Pachysoma) 
haliaetus Linnaeus, 1758, Falco : 
harengus Linnaeus, 1758, Clupea 

hardwickii Leach, 1817, Nursia 

harrisii Gould, 1841, Pilumnus 

helias Pallas, 1781, Ardea 

hemingi Alcock & "Anderson, 1899, Benthochason 
henslowii Leach, [1820], Polybius 

hepsetus Linnaeus, 1758, Atherina 

hippurus Linnaeus, 1758, Coryphaena 

hirtellus Linnaeus, 1761, Cancer 

hirtipes Heller, [1865], Pinnaxodes as 

hispida Eydoux & Souleyet, 1842, Domecia 

hoazin Miiller (P.L.S.), [1776], Phasianus 

horridus Linnaeus, 1758, Crotalus 

horridus Rippell, 1830, Oreophorus 

imitatrix Hilgendorf, 1869, Deckenia .. 

impressus Smith (S.1.), 1870, Glyptograpsus 
inaequipes Stimpson, 1858, Asthenognathus 

indus Boddaert, 1783, Falco “X 

integerrimus Lamarck, 1818, Cancer .. 

isenbeckii Brandt, 1848, Platycorystes (Podacanthus).. 
Jacquemontii Milne Edwards (H.), 1844, Gecarcinucus 
Javanicus Horsfield, 1821, Eurylaimus a 
jurinei Saussure, 1853, Guaia (llia) 

lacertina Linnaeus, 1766, Siren 

laevigata Milne Edwards (H.) & Lucas, [1844], Pinnotherelia 
lambriformis Milne Edwards (A.), 1873, Ree reeeyetes 
lamelligera Stimpson, 1858, Onychomorpha 

lavaretus Linnaeus, 1758, Salmo ae 

leeanus Rathbun, 1898, Lipaesthesius .. 

leucorodia Linnaeus, 1758, Platalea .. ne 
liguricus Milne Edwards (H. yr i853, Euchirigrapsus Ee 
limosa Say, 1818, Cancer i 
litteratus Fabricius, 1798, Cancer 

lucius Linnaeus, 1758, Esox ue 

lugubris Alcock, 1899, Hephthopelta 

lumpus Linnaeus, 1758, Cyclopterus 

lupus Linnaeus, 1768, Anarhichas 


446 Opinions and Declarations 


Page 
Official List of Specific Names in Zoology, names placed on (contd.) : 

lyra Linnaeus, 1758, Callionymus ae ay ie ea = igs ig 369 
macgillivrayi White, 1852, Ommatocarcinus .. B: b a A a 17 
maculatus Linnaeus, 1758, Cancer Bis ese hd me mt ois AS 5 
magna Linnaeus, 1758, Alauda .. ee Ly ~ - tay fas he 180 
major Linnaeus, 1758, Parus .. a He a a of a “ 178 
mantellii Owen, 1848, Notornis bed se et xe Pe at A 178 
marginalis Bloch, 1793, Epinephelus .. ae ve M4 e a ; eee 
maximus Geoffroy, 1850, Aepyornis . ike Je ie Le 173 
melanochirus Milne Edwards (A.), 1873, Chlorodopsis ie i La aa 5 
meleagris Linnaeus, 1758, Phasianus .. as ae ae i 178 
microps Alcock & Anderson, 1894, Xanthodes’ ie os ae ces L: 18 
militaris Linnaeus, 1758, Emberiza ses ; as iy Le uh oF, 183 
milneedwardsii Miers, 1886, Paracyclois es wt BY re = ae 9 
minutus Linnaeus, 1758, Cancer ' ad K diy sie Ae a 23 
mocinno de la Llave, 1832, Pharomachrus te ot i a so a 178 
moebii Richters, 1880, Xenophthalmodes Py se ae oe % a 22 
monstrosa Linnaeus, 1758, Chimaera .. a oh sf Ee! ts ae 369 
montagui [Leach], [1814], Pandalus  .. a8 a x. ay, - 245 
morhua Linnaeus, 1758, Gadus ee ae aN ov bie st 369-370 
moschata Linnaeus, 1758, Anas.. ; a ay hs a ae zs 175 
muriger Adams & White, [1849], Tlos” ag ie 2 ahs a ua i 
murigera White, 1846, Xanthasia Ne an as a a si a3 22 
mustelus Linnaeus, 1758, Squalus a ae be bc ‘f =e sp eye 
narina Stephens, 1815, Trogon ae Ss ie Ne rat = i 174 
neapolitana Kossmann, 1878, Paragalene Re oe az * aA i 18 
niger Linnaeus, 1758, Gobius ee - oe A Fe Hs, - we 370 
niger Miller (P.L.S.), [V7 7OleBuceo: a re "2 re a {9 175 
niger Hartig, 1840, Figites we oe a af i Ks 8 at 284 
nigra Gmelin, 1788, Paradisea [sic] .. ms Pe tes LA 2 be 174 
nigricephala Jameson, 1835, Tanagra .. a ca an ee ua = 184 
nitidus Latham, 1790, Falco ah Ye Hs oe a Se tae a3 183 
nitidus Stimpson, 1858, Sphaerozius .. bs a ya Ye ae iS 21 
nitidus Stimpson, 1859, Euryplax Se hd - oe use i a 14 
nitidus Milne Edwards (A.), 1867, Libystes  .. ae a: Le os - 16 
nitidus Milne Edwards (A.), 1870, Catoptrus .. * es a su e 12 
nitidus Smith (S.I.), 1870, Dissodactylus oe a ue) x mh = 13 
nitidus Milne Edwards (A.), 1873, Notonyx .. us << as fy a 17 
nobile Milne Edwards (A.), 1880, Dene Me + ae ie ys 12 
nodosus Bell, 1855, Oreophorus ‘ 3 - aa a otf 1 
norvegicus Linnaeus, 1758, Cancer ae oa an Ae Be he ee 22 
novaehollandiae Latham, 1801, Cereopsis ve we ns + te me Wis 
occidentalis Lawrence, 1858, Podiceps .. fe a ie Hh x A 173 
ocellaris Linnaeus, 1758, Blennius ne te ca Me i Hs < 371 
oenas Linnaeus, 1758, Columba ae a a re 86 
orbignyi Milne Edwards (H.) & Lucas, [1844], “Xantho a Ae LE * 19 
orientalis Miers, 1886, Lupocyclus (Parathranites) set oe ah oF AE 9 
oriolus Linnaeus, 1758, Coracias a Re <t 24 aS ae ah 182 
ornata Randall, 1840, Ilia ays tye a ye on a: a 11 
ornatus Geoffroy, 1809, Cephalopterus. . a an sus Bh: eh re 175 
ornatus Dana, 1852, Cyclodius ae a ne Ae de oA 6 
ostralegus Linnaeus, 1758, Haematopus ae Ne Ae ee es i 7a, 
pagurus Linnaeus, 1758, Cancer : bs ae ws a be x Dips 
papulosus Stimpson, [1871], Carpoporus ne ae Ba ae Be Lt 5 
paradiseus Linnaeus, 1758, Polynemus ats ne ee a bd a 372 
paradiseus Swainson, 1825, Prloriss —... “me £. we ae 10 sa) 179 
paradoxa Pallas, i778 Tetrao i , a ue he, ae "e 180 
parvula Stimpson, 1858, Acmaeopleura ae sa es ce of os AZ 
patagonica Miller, 1778, Aptenodytes .. i Ba oe ot a5 hy 174 


pectoralis Latham, 1801, Muscicapa .. . if fys oe As ms 178 


Volume 1, Section D 


Official List of Specific Names in Zoology, names placed on banal : 


pentagonus Stimpson, 1858, Cryptocnemus .. 
perlata Milne Edwards (H. ), 183% Ti (ieee (sie) . 
perlatus Poeppig, 1836, Hepatus : os 
picta Milne Edwards (H. ), 1848, Bellia 

pictus Linnaeus, 1758, Phasianus 

pinnotheroides White, 1846, Xenophthalmus 
piscatorius Linnaeus, 1758, Lophius 

pisonis Gmelin (J.F.), [1789], Gobius 

pisum Linnaeus, 1758, Cancer .. 

planissimus Herbst, [1804], Cancer - 

planus Milne Edwards (H.), 1834, Xantho 

polybioides Adams & White, [1849], Lissocarcinus 
porcus Linnaeus, 1758, Scorpaena 

porzana Linnaeus, 1766, Rallus ae 

pugnax Smith (S.1.), 1870, Glyptoplax 

punctatus Linnaeus, 1758, Cancer 

pygmaea Linnaeus, 1758, Platalea 

pPyrrhopterus Latham, 1801, Psittacus .. : 
quadridens Milne Edwards (Ry 1869, Thalamitoides 
quadridentata Latreille, [1828], Melia . : ; 
regia Linnaeus, 1758, Paradisaea 

reticularis Bloch, 1795, Gymnothorax 

rhomboides Linnaeus, 1758, Cancer 

rodgersii Stimpson, 1858, Lachnopodus 

roseus Nobili, 1906, Parapleurophrycoides on 
rotundatus Adams & White, [1849], Lupocyclus . 
rotundifrons Milne Edwards (A.), 1867, Heterozius .. 
rousseauxi Milne Edwards (H.), 1853, Metasesarma 
rouxii Guérin-Meéneville, 1832, Caphyra 

ruber Linnaeus, 1758, Phoenicopterus ae 
rubromaculatus de Haan, [1833], Cancer ’ (Liagore) 
rufa Boddaert, 1783, Tanagra 

salar Linnaeus, 1758, Salmo 


Savignyi (correction of savignii) Milne | Edwards a ds 1834, Cancer 


scaber Linnaeus, 1758, Uranoscopus 

scabripes Rathbun, 1898, Tetrias 

scolopacea Gmelin, 1788, Ardea 

scombrus Linnaeus, 1758, Scomber 

sculptipes Stimpson, [1871], Micropanope 
sefilata Forster, 1781, Paradisea [sic] ; 
septemspinosus Fabricius CC), 178i Cancer 
serpentina Linnaeus, 1758, Testudo 

serrata White, 1847, Valdivia 

serratus Forskal, lake Cancer .. ae 
sexdentatum Stimpson, 1858, Camptandrium 
sexdentatus Paulson, 1875, Archias 

sexpes Fabricius, 1798, Cancer 

signatus Adams & White, [1849], Carpilus 
speciosa Dana, 1852, Nucia é 

spectabilis de Man, 1896, Platychirograpsus va 
spinicarpus Milne Edwards (CH), 1852, EaCheri ar 
spinosipes Stimpson, 1858, Scalopidia 

spinosus Rathbun, 1894, Erimetopus d j 
spongiosus Adams & White, [1849], Iphiculus 
sponsa Linnaeus, 1758, Anas see : 
sternberghii Stimps on, 1859, Xanthodius 
stridulans Wood-Mason, 1892, Psopheticus 
struthersii Vigors, (18321, Ibidorhyncha 

sturio Linnaeus, 1758, Acipenser 


448 Opinions and Declarations 


Page 
Official List of Specific Names in Zoology, names placed on (contd.) : 
subglobosa Stimpson, [1871], Zliacantha be) ay ad i At = Jf 
subquadrata Dana, 1851, Fabia .. ey ave a g% ae we we 14 
sulcatus Swainson, 1820, Preroglossus .. Mn Ae is a aG Ay 174 
superba Costa, 1838, Portunus .. 3 hee fs Oi £4 Oe ae 25 
swainsonii Vigors, 1825, Gampsonyx .. a a bis se Ae Ws | i a 
tabacaria Linnaeus, 1758, Fistularia .. ik - on vA Lys Lk 368 
tenuipes Dana, 1852, Carupa .. aa SE ab & He Be ae 5 
tesselatus Latreille, 1812, Graspus ae he et 1 A Ay = 17 
thalamita Stimpson, [1871], Melybia .. se fee me oh " ee Ly 
tiphia Linnaeus, 1758, Motacilla e B a: ‘3 oe ak o 183 
tobianus Linnaeus, 1758, Ammodytes .. ee - a a Ba cid 368 
tomentosus Dana, 1852, Actumnus Ae x Ay a me a ans 3 
torquilla Linnaeus, 1758, Jynx .. Me me vs as he oF 2h ae 
transversus Rathbun, 1893. Opisthopus an nA as at, = ce Ly 
trapeziformis Nauck, 1880, Holothuriophilus .. nt “A ie Be oe 15 
tridens de Haan, [1835], Ocypode (Helice) .. ae sit A ns ah 15 
tridens Kr@yer, 1837, Geryon .. E ue as i, a 14 
tridentata Milne Edwards (Hi) 1853. Parathelphusa tds fe . ie 9 
tridentata Miers, 1879, Pseudophilyra 4 af a in na 10 
tridentatus Milne Edwards (A.), 1879, Coenophthalmus se A ie oe 5 
tristis Dana, 1852, Carpilodes - ai in bye 2 
truncatus Stimpson, 1860, Metopocarcinus eis 5 bi $s zt ee iF 
tuberosa Stimpson, [1871], Osachila. ~.. ve A mt ie oe ue 9 
tuberosus Pennant, 1777, Cancer : ut sad - at es 6 
typhlus Milne Edwards (A. ), 1880, Bathyplax ae it ws “st i 12 
typicus Rathbun, 1898, Chasmocarcinus my He Se ay a4 # 13 
ungulata Linnaeus, 1758, Musca ve ble Me AL he un 3) 26s 
urophasianus Bonaparte, 1827, Tetrao .. ae ih ss AS 4 at We: 
vesiculosa Alcock, 1896, Heteronucia .. Me cc Res v aS Ae 7 
vigil Weber, 1795, Portunus oe (3 me ot ne ni oe sf 23 
violacea Isert, 1789, Musophaga a a: Ese ae ae x ne 178 
volans Linnaeus, 1758, Draco .. - a o at je we 373-374 
vulgaris Linnaeus, 1758, Sturnus fe a ie Re a) ey bw 180 
vulturina Hardwicke, 1834, Numida .. ret a Ne ry “2 “is 173 
wallicus Kerr, 1792, Psittacus .. ae — Ms a es a a. 183 
Official List of Works Approved as Available for Zoological Nomenclature, titles of 
works placed on: 
Roding (P.F.), 1798, Museum Boltenianum .. As ts ane ob en 
Oleacina Réding, 1798 (Class Gastropoda), placed on the Official List 2s Generic 
Names in Zoology in Opinion 119, determination of gender of A 158 
Omalaspis Giraud, 1860 (Class Insecta, Order Hymenoptera), placed on the Official 
List of Generic Names in Zoology with Name No. 1036, with Omalaspis noricus 
Giraud, 1860, as type species .. ae a ee oe nid ae «piv REE 
gender of name .. ee a = Ae aie “ ne 2 3; 284 


Ommatocarcinus White, 1852 (Class Crustacea, Order pera completion of 
Ruling given in Opinion 85 Ne -% ey a ‘ ve a of a 


Volume 1, Section D 449 


Page 
Oniscus Linnaeus, 1758 (Class Crustacea), placed on the Official List of Generic 
Names in Zoology in Opinion 104, determination of gender of a cn - 142 


Ophidion Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 92.. 341 


Ophidion Pomel, 1853 (a junior homonym of Ophidion Linnaeus, 1758), placed on 
the Official Index ei pees & and Invalid Generic Names in 1 Zoology with Name 
No. 806 .. 346 


Ophidium Linnaeus, 1766 (an Invalid Emendation of Ophidion Linnaeus, 1758), 
placed on the Official Index of puree and Invalid Generic Names in Oe 
with Name No. 807 S3 ; 346 


Opinion 38 (status of the anonymous work published in 1771 entitled Ornithologia 
britannica), cancellation of, for all except historical purposes Be ze a 85 


Opisthopus Rathbun, 1893 ais Crustacea, Order pened: eompicuenis of by 
given in Opinion 85 : 51 


orbignyi Milne Edwards (H.) & Lucas, as published in the combination Xantho 
orbignyi (Class Crustacea, Order Decapoda), correction of date of, from 1843 to 
[1844] (correction of Opinion 85) ay: oa a ah ns be ee 25 


placed on the Official List of Specific Names in Zoology with Name No. 905 _ 19 


orientalis Miers, 1886, as published in the combination Lupocyclus (Parathranites) 
orientalis (Class Crustacea, Order Decapoda), Sa on the on List i 
Specific Names in Zoology with Name No. 819 ‘ 9 


Oriolus Linnaeus, 1766 (Class Aves), Coracias oriolus Linnaeus, 1758, type species 
of, under Declaration 21 ay Bs eo, re e “ de a 181 


oriolus Linnaeus, 1758 (Coracias) (Class Aves) (a nominal species objectively 
identical with Oriolus galbula Linnaeus, ee type species of Oriolus Linnaeus, 
1766, under Declaration 21 ne : ; A - ae she 181 


placed on the Official List of Specific Names in Zoology with Name No. 1020 .. 182 


ornata Randall, 1840, as published in the combination J/ia ornata (Class Crustacea, 
Order Decapoda), eee on the te ae List aos. ae Names in ee, with 
Name No. 830 Il 


ornatus Geoffroy, 1809, as published in the combination Cephalopterus ornatus 
(Class he DE on the OES List ee BaECte Names in RORY with Name 
No. 968 175 


ornatus Dana, 1852, as published in the combination Cyclodius ornatus (Class 
Crustacea, Order Decapoda), Blescine on the Sissel List "eh Seeaiie a Names in 
Zoology with Name No. 785 .. 6 


Orphnoxanthus Alcock, 1898 ee Crustacea, Order Hoc drt bien tac of 
Ruling given in Opinion 85 : A aL 


450 Opinions and Declarations 


Page 
ostralegus Linnaeus, 1758, as published in the combination Haematopus ostralegus 
ae placed on the Cae List i esha: Names in aca, with Name 

fe) ae mW, 


Otarion Zenker, 1833 (Class Trilobita), placed on the oes List ia Generic Names 
in Zoology in Opinion 88, determination of gender of 144 


Pachycephala Klug, 1834 (a junior homonym of Pachycephala Vigors, 1825), placed 
on the Official Index ee ieunce and Invalid Generic Names in aus with Name 
ING. F509 Le 217 


Pachycephala Doleschall, 1858 (a junior homonym of Pachycephala Vigors, 1825), 
placed on the Official Index oy, bara and Invalid Generic Names in FOR 
with Name No. 560 We PAR 


Pachycephala Lioy, 1864 (a junior homonym of Pachycephala Vigors, 1825), placed 
on the Official Index ees pe and Invalid Generic Names in iil with Name 
No. Soler: 217 


Pachycephala Broun, 1881 (a junior homonym of Pachycephala Vigors, 1825), placed 
on the Official Index a paces and Invalid Generic Names in Morante with Name 
No. 562° =. 247 


Pachycephala Fairmaire, 1883 (a junior homonym of Pachycephala Vigors, 1825), 
placed on the Official Index of eeiai and Invalid Generic Names in ey 
with Name No. 563... i : : : 217 


Pachycephalus Stephens, 1826 (an Invalid Emendation of Pachycephala Vigors, 
1825), placed on the Official Index oy sails ¢ and Invalid Generic Names in 
Zoology with Name No. 564... i : 207 


pagurus Linnaeus, 1758, as published in the combination Cancer pagurus (Class 
Crustacea, Order Decapoda), Saag: on the ea List vi per Names in 
Zoology with Name No. 933... Z wR 


Pandalus {Leach], [1814] (Class Crustacea, Order Decapoda), substitution of, on the 
Official List of Generic Names in Zoology in place of Pandalus Leach, 1815 ss AS 


Pandalus Leach, [1815] (Malac. Podophth. Brit.) (a junior homonym of Pandalus 
[Leach], [1814]), placed on the Official Index of ig and Invalid Generic 
Names in Zoology with Name No. 730 : nee 245-246 


Pandalus Leach, 1815 (Trans. linn. Soc. Lond.), removal of, from the Official List of 
Generic Names in Zoology, on which placed by Opinion 104, and insertion of, on 
the Official Index of Rejected and Invalid Generic Names in Le as a junior 
homonym of Pandalus {Leach], [1814], with Name No. 731 : 245-246 


Pandion Savigny, 1809 (Class ie Falco haliaetus Linnaeus, 1758, ‘ype species of, 
under Declaration 21 .. 181 


Panoplax Stimpson, [1871] (Class Crustacea, Order Decapoda), completion of 
Ruling given in Opinion 85 re an i fe ne wt Se =| 


Volume 1, Section D 


papulosus Stimpson, [1871], as published in the combination Carpoporus papulosus 
(Class Crustacea, Order Decapoda), placed on the Official List of Specific Names 
in Zoology with Name No. 779.. < ¥. a = a: — ey 


paradiseus Linnaeus, 1758, as published in the combination Polynemus paradiseus 
(Class Pisces), placed on the Official List of Specific Names in Zoology with Name 
No. 1101 ¥ ee os a at ‘% i: re a. a 


paradiseus Swainson, 1825, as published in the combination Priloris paradiseus 
(Class Aves), placed on the Official List of Specific Names in Zoology with Name 
No. 1003 be ay = a ae Sh — are i ae 


paradoxa Pallas, 1773, as published in the combination Tefrao paradoxa (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 1012 


Paragalene Kossmann, 1878 (Class Crustacea, Order Decapoda), completion of 
Ruling given in Opinion 85 i we y a a i : 


Paragordius Camerano, 1897 (Class Nematomorpha), placed on the Official List 
of Generic Names in Zoology in Opinion 66, determination of gender of a: 


Parapanope de Man, 1895 (Class Crustacea, Order Decapoda), completion of Ruling 
given in Opinion 85 ne “ 23 ae Poe ae as in ce 


Parapleurophrycoides Nobili, 1906 (Class Crustacea, Order Decapoda), completion 
of Ruling given in Opinion 85 .. Ae she ma ae xt &3 a 


Paraxanthus Milne Edwards (H.) & Lucas, [1844] (Class Crustacea, Order Deca- 
poda), completion of Ruling given in Opinion 85 .. Be a ah ibe 


correction of date of, from 1843 to [1844] 


Pariphiculus Alcock, 1896 (Class Crustacea, Order Decapoda), completion of Ruling 
given in Opinion 73 ite a $5 Si ae Ae: of Ass a 


Parotica Wagler, 1827 (an Erroneous Subsequent Spelling for Parotia Vieillot, 1816), 
placed on the Official Index of Rejected and Invalid Generic Names in Zoology 
with Name No. 565 Fx at ny Ne be ts ae oR im 


parvula Stimpson, 1858, as published in the combination Acmaeopleura parvula 
(Class Crustacea, Order Decapoda), placed on the Official List of Specific Names 
in Zoology with Name No. 842.. & Pgs ee & ae ze a 


patagonica Miller, 1778, as published in the combination Aptenodytes patagonica 
(Class Aves), placed on the Official List of Specific Names in Zoology with Name 
ING: 956 .. Ae ae “3 ‘i i; e. - as ae a 


pectoralis Latham, 1801, as published in the combination Muscicapa pectoralis 
(Class Aves), placed on the Official List of Specific Names in Zoology with Name 
No. 992 .. ie a La a. J he be 4 ~ ie 


Pediculus Linnaeus, 1758 (Class Insecta), placed on the Official List of Generic 
Names in Zoology in Opinion 104, determination of gender of ors 2: 


S72 


179 


180 


a 


Is5 


51 


51 


51 
aT 


53 


PF 


||P 


174 


178 


141 


452 Opinions and Declarations 


pennanti Leach, [1817], as published in the combination Ebalia pennanti (a junior 
objective synonym of tuberosus Pennant, 1777, as published in the combination 
Cancer tuberosus), placed on the Official Index of beeen and Invalid patie: 
Names in Zoology with Name No. 337 


pentagonus Stimpson, 1858, as published in the combination Cryptocnemus 
pentagonus (Class Crustacea, Order Decapoda), PE rnaO on the racial List of 
Specific Names in Zoology with Name No. 784 : : 


Perca Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 


Percnon Gistl, 1848 (Class Crustacea, Order eames aA ea of pas sie 
in Opinion 85 wt aes : 


Perigrapsus Heller, 1862 oe Crustacea, Order eva tas eau te of euer: 
given in Opinion 85 : 


Periplaneta Burmeister, 1838 (Class Insecta), placed on the Official List a Generic 
Names in Zoology in Opinion 104, determination of gender of : 


perlata Milne Edwards (H.), 1837, as published in the combination Thelpheusa [sic] 
perlata (Class Crustacea, Order Decapoda), cia on the aaa: List = Specie 
Names in Zoology with Name No. 827 


perlatus Poeppig, 1836, as published in the combination Hepatus perlatus (Class 
Crustacea, Order Decapoda), Place on the eee List hl eee Names in 
Zoology with Name No. 898 .. 


Persephona Leach, 1817 (Class Crustacea, Order a as souehuee of gees 
given in Opinion 73 ‘ vs ve os : es é 


Persephona Leach, 1852 (a junior homonym of Persephona Leach, 1817), placed on 
the Official Index es 7 ae and Invalid Generic Names in 1 Zoology with Name 
No. 497 2 ~ 


Pezoporus Klug, [1842] (a junior homonym of Pezoporus Illiger, 1811), placed on 
the Official Index a Eee and Invalid Generic Names in et with Name 
No. 566 


Pezoporus Foerster, 1868 (a junior homonym of Pezoporus Illiger, 1811), placed on 
the Official Index a a sbaa and Invalid Generic Names in Oa with Name 
No. 567 : 


Phlyxia Bell, 1855 (Class Crustacea, Order chaieieasiene eam ee of peo aie 
in Opinion 73 : : 


correction of reference for 


Phthirus Leach, 1815 (Class Insecta), placed on the Official List oe Generic Names 
in Zoology in Opinion 104, determination of gender of ‘ 


Physa Draparnaud, [1801] (Class Gastropoda), placed on the ae List ue Generic 
Names in Zoology in Opinion 94, determination of gender of ‘ 


Page 


62 


340 


143 


10 


18 


53 


60 


218 


218 


53 
aS 


141 


157 


Volume 1, Section D 453 


Page 
Physalia Lamarck, 1801 (Class Hydrozoa), placed on the oor List ~ Generic 
Names in Zoology i in Opinion 80, determination of gender of ; (oi 
picta Milne Edwards (H.), 1848, as published in the combination Bellia picta (Class 
Crustacea, Order Decapoda), Seager Mt on the ee List ae Names in 
Zoology with Name No. 774 ae ; 4 
pictus Linnaeus, 1758, as published in the combination Phasianus pictus (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 971 ee 176 
Pilumnoides Milne Edwards (H.) & Lucas, [1844] (Class Crustacea, Order Deca- 
poda), completion of Ruling given in Opinion So e: §2 
correction of date of, from 1843 to [1844] .. = 2 M: a nf a7 
Pilumnus Leach, 1815 ees: Crustacea, Order pees); bea of art) 
given in Opinion 85 d e 32 
Pilumnus Rafinesque, 1815 (a junior homonym of Pilumnus Leach, 1815), placed on 
the Official Index pe ee and Invalid Generic Names in : Zoology with Name 
No. 498 .. 60 


Pilumnus Dejean, 1821 (a junior homonym of Pilumnus Leach, 1815), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 499 60 


Pilumnus Koch (C.L.), 1837 (a junior homonym of Pilumnus Leach, 1815), placed 
on the Official Index see Sak and Invalid Generic Names in n Zoology with Name a 
mae, 500 > .. 


Pilumnus Gistl, 1848 (a junior homonym of Pilumnus Leach, 1815), placed on the 
Official Index up cha and Invalid Generic Names in ee”: with Name - 
No. 20... : 


Pilumnus Bonaparte, 1854 (a junior homonym of Pilumnus Leach, 1815), placed on 
the Official Index ao noes 2 and Invalid Generic Names in 1 Zoology with Name a 
ING: 502... . 


Pinnaxodes Heller, [1865] Selatan Crustacea, Order sean ea Soe ens of ee 


given in Opinion 85 5 50 
Pinnixa White, 1846 Gee Crustacea, Order ee ee. competion of Reoen: 

given in Opinion 85 : , 50 
Pinnotherelia Milne Edwards (H.) & Lucas, [1844] oe Crustacea, Order 

Decapoda), completion of Ruling given in Opinion 85 : at ae bs 52 

correction of date of, from 1843 to [1844] .. or 48 — iis Ls 57 


Pinnotheres Bosc, [1801—1802] (Class Crustacea, Order Decapoda), substitution of, 
on the Official List of Generic Names in Zoology i in place of Pinnotheres Latreille, 
[1802—1803 ] 7 cae 2: As ; ie ee ¥ a = 29235 


454 Opinions and Declarations 


Page 
Pinnotheres Latreille, [1802—1803], removal of, from the Official List of Generic 
Names in Zoology, on which placed by Opinion 85, and insertion of, on the Official 
Index of Rejected and Invalid Generic Names in Zoology as a sot homonym 
of Pinnotheres Bosc, [1801—1802], with Name No. 577 sc - 235-236 


pinnotheroides White, 1846, as published in the combination Xenophthalmus 
pinnotheroides (Class Crustacea, Order Decapoda), aaa on the Sisto: List - 
Specific Names in Zoology with Name No. 930 ee 22 


piscatorius Linnaeus, 1758, as published in the combination Lophius piscatorius 
(Class Pisces), placed on the ee List sd pelle Names in Sper) with Name 
No. 1082 : 370 


pisonis Gmelin (J.F.), [1789], Gobius (Class Pisces), insertion of, on the Official 
List of Generic Names in Zoology as name of type gana of Eleotris Schneider, 
1801 (correction of Opinion 93) ¥4 : ' sre at at 342 


placed on the Official List of Specific Names in Zoology with Name No. 1096 .. 372 


pisum Linnaeus, 1767, as published in the combination Cancer pisum (Class 
Crustacea, Order Decapoda), Seach on the ee List 1 liek Names in 
Zoology with Name No. 1035 .. 235-236 


Planes Bowditch, 1825 apes Se ee Order Hereage eek: between of goon? 
given in Opinion 85 : : : 52 


Planes Saussure, 1862 (a junior homonym of Planes Bowditch, 1825), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 503 60 


Planes Rondani, 1864 (a junior homonym of Planes Bowditch, 1825), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 504 60 


planissimus Herbst, [1804], as published in the combination Cancer planissimus 
(Class Crustacea, Order Decapoda), se a on the esieak List es pcos Names 
in Zoology with Name No. 896.. 18 


planus Milne Edwards (H.), 1834, as published in the combination Xantho planus 
(Class Crustacea, Order Decapoda), otis on the une c List a ADEEDIE Names 
in Zoology with Name No. 1057 ce 284 


Platychirograpsus de Man, 1896 (Class Crustacea, Order eee bee a of 
Ruling given in Opinion 85 gy : 50 


Platypilumnus Alcock, 1894 (Class au Order Sati cigs! SOP at of 
Ruling given in Opinion 85 sys , 50 


Platyxanthus Milne Edwards (A.), 1863 (Class Crustacea, Order Decapoda), 
completion of Ruling given in Opinion 85... ag ae ae 52 


Podacanthus Brandt, 1848 (a junior homonym of Podacanthus Gray (J. E.), 1833), 
placed on the Official Index y oe and Invalid Generic Names in feet 
with Name No. 505 _ : x? 60 


Volume 1, Section D 


Podophtalmus Lamarck, 1801 (an Invalid Original Spelling for Podophthalmus 
Lamarck, 1801), placed on the Official Index of tae and Invalid Generic 
Names in Zoology with Name No. 506 


Podura Linnaeus, 1758 (Class Insecta), placed on the oleae List sa Generic Names 
in Zoology in Opinion 104, determination of gender of 


polybioides Adams & White, as published in the combination Lissocarcinus 
polybioides (Class Crustacea, Order ede: correction of date of, from 1848 
to [1849] (correction of Opinion ye) abs ty au a va m2 


placed on the Official List of Specific Names in Zoology with Name No. 806 


Polydectus Rafinesque, 1815 (a nomen nudum), placed on the Se a Index ab. 
Rejected and Invalid Generic Names in Zoology with Name No. 507. 


Polydectus Milne Edwards (H.), 1837 ane Crustacea, Order Pera com- 
pletion of Ruling given in Opinion 85 


Polydesmus Latreille, [1802—1803] (Class Diplopoda), placed on the Official List 
of Generic Names in Zoology in Opinion 104, determination of gender of 


Polyplectron Temminck, 1807 (Class sna Pavo bicalcaratus Linnaeus, 1758, Bas 
species of, under Declaration 21 : at Be ‘ 


Porcellio Latreille, [1803—1804] (Class Crustacea), placed on the Official List of 
Generic Names in Zoology in Opinion 104, determination of gender of 


porcus Linnaeus, 1758, as published in the combination Scorpaena porcus (Class 
Pisces), placed on the as List e sia ie Names in ee with Name 
No. 1088 


porzana Linnaeus, 1766, as published in the combination Rallus porzana (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 999 


Prionoplax Milne Edwards (H.), 1852 (Class Crustacea, Order Decapoda), 
completion of Ruling given in Opinion 85... ms sa ia 


Pseudocarcinus Milne Edwards (H.), 1834 (Class Crustacea, Order eee 
completion of Ruling given in Opinion 85 : 


Pseudophilyra Miers, 1879 (Class Crustacea, Order pega ee of 
Ruling given in Opinion 73 : : 


Pseudopinnixa Ortmann, 1894 (Class Crustacea, Order Pe pou) ene of 
Ruling given in Opinion 85 a iy we ‘ : a 


Pseudopinnixa Holmes, 1895 (a junior homonym of Pseudopinnixa Ortmann, 1894), 
placed on the Official Index Hat Rpegee and Invalid Generic Names in Zoology 
with Name No. 508 - . 


Pseudorhombila Milne Edwards (H.), 1837 (Class Crustacea, Order Decapoda), 
completion of Ruling given in Opinion 85 : ae Be fs 


455 
Page 


61 


143 


24 
28 


6] 


ope 


14] 


182 


142 


371 


179 


52 


39) 


26, 


52 


61 


49 


456 Opinions and Declarations 


Psittacus Catesby, 1777 (a junior homonym of Psittacus Linnaeus, 1758), placed 
on the Official Index — Roce and Invalid Generic Names in n Zoology with Name 
No. 568 : : 


Psopheticus Wood-Mason, 1892 (Class Crustacea, Order riacanes ea deere of 
Ruling given in Opinion 85 ; 


psophia Pallas, 1767, as published in the combination Grus psophia (a junior objective 
synonym of crepitans Linnaeus, as published in the combination Psophia crepitans), 
placed on the Official Index of baa and Invalid ees Names in meme 
with Name No. 364 be ; 


Psoroptes Gervais, 1841 (Class Arachnida), placed on the Official List ue Generic 
Names in Zoology i in Opinion 104, determination of gender of 


Pteroglossus Chaudoir, 1843 (a junior homonym of Preroglossus Illiger, 1811), placed 
on the Official Index ge pe ge and Invalid Generic Names in eee with Name 
ING: "569" = 


Ptilorhis Agassiz (J.L.R.), 1846 (an Invalid Emendation of Pri/oris Swainson, 1825), 
placed on the Official Index of Hee and Invalid Generic Names in Zoology 
with Name No. 576 : 


Ptilorhys Bonaparte, 1854 (an Erroneous Subsequent Spelling for Ptiloris Swainson, 
1825), placed on the Official Index a lai and Invalid Generic Names in 
Zoology with Name No. 570 


Ptilornis Gray (G.R.), 1869 (an Erroneous Subsequent Spelling for Ptiloris Swainson, 
1825), placed on the Official Index of Reeaied and Invalid Generic Names in 
Zoology with Name No. 571 ; 


Prychognathus Stimpson, 1858 (Class Crustacea, Order eee hagas” of 
Ruling given in Opinion 85 : : - me f 


Ptychognathus Owen, 1860 (a junior homonym of Prychognathus Stimpson, 1858), 
placed on the Official Index a acuiiti and Invalid Generic Names in Coct 
with Name No. 509 : ; 


pugnax Smith (S.I.), 1870, as published in the combination Glyptoplax pugnax 
(Class Crustacea, Order Decapoda), Be on the Oba & List si Speer Names 
in Zoology with Name No. 865. 


Pulex Linnaeus, 1758 (Class Insecta), placed on the Official List ola Generic Names 
in Zoology in Opinion 104, determination of gender of ; 


punctatus Linnaeus, 1758, as published in the combination Cancer punctatus (Class 
Crustacea, Order Decapoda), Sonal on the ree List a pte Names in 
Zoology with Name No. 937 .: a 


pygmaea Linnaeus, 1758 (Platalea) (Class Aves) (a nominal species objectively 
identical with Eurynorhynchus griseus Nilsson, oe type ae of Eg aa 
Nilsson, 1821, under Declaration 21 oA af 


placed on the Official List of Specific Names in Zoology with Name No. 1017 .. 


Page 


218 


a2 


324 


142 


218 


218 


218 


218 


52 


61 


15 


141 


25 


181 
182 


Volume 1, Section D 


Pyrosoma Peron, 1804 (Class Urochorda), placed on the Official List _ Generic 
Names in Zoology i in Opinion 77, determination of gender of 


pyrrhopterus Latham, 1801, as published in the combination Psittacus pyrrhopterus 


Sor placed on the eas List a nage Names in paris with Name 
No. 963... 


Pyxidognathus Milne Edwards (A.), 1879 (Class Crustacea, Order Decapoda), 
completion of Ruling given in Opinion 85... C4 B 2 & 


quadridens Milne Edwards (A.), 1869, as published in the combination Thalamitoides 
quadridens (Class Crustacea, Order Decapoda), ies on the isan List = 
Specific Names in Zoology with Name No. 835 ; 


quadridentata Latreille, [1828], as published in the combination Melia quadridentata 


(Class Crustacea, Order Decapoda), aang on the ete List fe eta Names in 
Zoology with Name ING 2910! 1 ee 


Reduvius Fabricius, 1775 (Class Insecta), placed on the Official List oe, Generic 
Names in Zoology i in Opinion 104, determination of gender of 


regia Linnaeus, 1758, as published in the combination Paradisaea regia (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 972 


reticularis Bloch, 1795, as published in the combination Gymnothorax reticularis 


(Class Pisces), Placed on the ee List ss bagi Names in ae with Name 
No. 1098 


Rhabditis Dujardin, 1845 (Class Nematoda), placed on the ae List we Generic 
Names in Zoology in Opinion 104, determination of gender of 


Rhipicentor Nuttall & Warburton, 1908 (Class Arachnida), placed on the Official 
List of Generic Names in Zoology in Opinion 73, determination of gender of 


Rhipicephalus Koch, 1844 (Class Arachnida), placed on the ae List Be Generic 
Names in Zoology i in Opinion 73, determination of gender of 


Rhithropanopeus Rathbun, 1898 aes Crustacea, Order Decapoda), completion 
of Ruling given in Opinion Noe mp or £2 ms oe 


Rhizocrinus Sars, [1865] (Class Crinoidea), placed on the Official List ue Generic 
Names in Zoology i in Opinion 73, determination of gender of : 


Rhizoglyphus Claparede, [1869] (Class Arachnida), placed on the Official List of 
Generic Names in Zoology in Opinion 104, determination of gender of 


Rhizopa Stimpson, 1858 (Class Crustacea, Order vere ae a completene of eee 
given in Opinion 85 ; 


457 


Page 
159 


175 


20 


14] 


176 


2 


157 


142 


142 


a2 


156 


142 


52 


458 Opinions and Declarations 


rhomboides Linnaeus, 1758, as published in the combination Cancer rhomboides 
(Class Crustacea, Order Decapoda), icin on the se: Bane List = one Names 
in Zoology with Name No. 941. 


rodgersii Stimpson, 1858, as published in the combination Lachnopodus rodgersii 
(Class Crustacea, Order ee tage cat on the Sd es List A pe ie Names 
in Zoology with Name No. 873. 


Roding (P.F.), ruled to be treated as the author of the Museum Boltenianum, 1798 


roseus Nobili, 1906, as published in the combination Parapleurophrycoides roseus 
(Class Crustacea, Order pecaros), ea on the il List of Sree Names 
in Zoology with Name No. 895. 


rotundatus Adams & White, as published in the combination Lupocyclus rotundatus 
(Class Crustacea, Order Decapoda), correction of date of, from 1848 to Rise 
(correction of Opinion 73) ; : 


placed on the Official List of Specific Names in Zoology with Name No. 808 


rotundifrons Milne Edwards (A.), 1867, as published in the combination Heferozius 
rotundifrons (Class Crustacea, Order Decapoda), Bpecais on the pOmoee List a 
Specific Names in Zoology with Name No. 800 ‘ 


rousseauxi Milne Edwards (H.), 1853, as published in the combination Metfasesarma 
rousseauxi (Class Crustacea, Order Decapoda), Waa on the ee List “3 
Specific Names in Zoology with Name No. 884 , 


rouxii Guérin-Méneville, as published in the combination Caphyra rouxii (Class 
Crustacea, Order Decapoda), correction of date of, from 1830 to 1832 (correction 
of Opinion 73) + re a4 id ae ari oe * 


placed on the Official List of Specific Names in Zoology with Name No. 776 


ruber Linnaeus, 1758, as published in the combination Phoenicopterus ruber (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 997 


rubromaculatus de Haan, [1833], as published in the combination Cancer (Liagore) 
rubromaculatus (Class Crustacea, Order Decapoda), placed on the Official List 
of Specific Names in Zoology with Name No. 875 .. cf: a3 bts aa 


rufa Boddaert, 1783, as published in the combination Tanagra rufa (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 1013... 


Ruppellioides Milne Edwards (A.), 1867 (Class Crustacea, Order Decapoda), 
completion of Ruling given in Opinion 85 } oe oe Ar 


salar Linnaeus, 1758, as published in the combination Salmo salar (Class Pisces), 
placed on the Official List of Specific Names in Zoology with Name No. 1086 .. 


Salmo Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 


Page 


23 


16 


257 


18 


24 


| 


178 


16 


180 


52 


370 


340 


Volume 1, Section D 459 


Page 
Samartium Dana, 1851 (Class Crustacea, Order ee sioteS completion of Ruling 
given in Opinion 85 As i ~ .? d ie Se + ae 52 
Sarcocystis Lankester, 1882 (Class Neosporidia), placed on the Official List 
Generic Names in Zoology i in Opinion 104, determination of gender of .. Py, 
Sarcoptes Latreille, [1802—1803] (Class Arachnida), placed on the Official List uh 
Generic Names in Zoology in Opinion 113, determination of gender of .. 142 
savignii Milne Edwards (H.), 1834, as published in the combination Cancer savignii 
(an Invalid Original Spelling for savignyi), placed on the Official Index of. ASIECs 
and Invalid Specific Names in Zoology with Name No. 335 24 
savignyi (correction of savignii) Milne Edwards (H.), 1834, as published in the com- 
bination Cancer savignii (Class Crustacea, Order Decapoda), placed on the 
Official List of Specific Names in Zoology with Name No. 935 ee * Ka 23 
scaber Linnaeus, 1758, as published in the combination Uranoscopus scaber (Class 
Pisces), placed on the ope. List oF tad Names in peeey with Name No. 
1070 ae 368 
scabripes Rathbun, 1898, as published in the combination Tetrias scabripes (Class 
Crustacea, Order Decapoda), De on the lip List gt Bs Names in 
Zoology with Name No. 923... 21 
Scalopidia Stimpson, 1858 (Class Crustacea, Order Decapoda), completion of 
Ruling given in Opinion 85 yf tS ae ay a ao eye 32 
Schistosoma Weinland, 1858 (Class Trematoda), placed on the Official List of 
Generic Names in Zoology i in Opinion 77, determination of gender of _... 139 
Scleroplax Rathbun, 1893 (Class Crustacea, Order Decapoda), completion of 
Ruling given in Opinion 85 a : th ee a, ae Bt Re 34 


scolopacea Gmelin, 1788, as published in the combination Ardea scolopacea (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 958 174 


Scomber Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 .. 340 


scombrus Linnaeus, 1758, as published in the combination Scomber scombrus (Class 
Pisces), See on the egress List of Sag Names in ies ae with Name No. 
1087 , 370-371 


Scorpaena Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion77.. 340 


Scorpio Linnaeus, 1758 (Class Arachnida), placed on the Official List os Generic 
Names in Zoology i in Opinion 104, determination of gender of = 142 


sculptipes Stimpson, [1871], as published in the combination Micropanope sculptipes 
(Class Crustacea, Order Decapoda), Be oe on the eT Sst oe pees Names 
in Zoology with Name No. 886.. 17 


460 Opinions and Declarations 


Scylla de Haan, [1833] (Class Crustacea, Order Beeanedal: correction of oaks 
relating to type species of (correction of Opinion 73) 


sefilata Forster, 1781, as published in the combination Paradisea [sic] sefilata (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 993 


Segestria Latreille, [1804] (Class Arachnida), placed on the Official List of Generic 
Names in Zoology i in Opinion 104, determination of gender of : 


Sepia Linnaeus, 1758 (Class Cephalopoda), placed on the ie List ae Generic 
Names in Zoology i in Opinion 94, determination of gender of 


septemspinosus Fabricius (J.C.), 1787, as published in the combination Cancer 
septemspinosus (Class Crustacea, Order Decapoda), pa on the a List 4 
Specific Names in Zoology with Name No. 1055 oe ; x 


serpentina Linnaeus, 1758, as published in the combination Testudo serpentina 


(Class Reptilia), placed 0 on the per List be See Names in Gey with 
Name No. 1106. 


serrata White, 1847, as published in the combination Valdivia serrata (Class 


Crustacea, Order Decapoda), placed on the 9 OU List oh perc Names in 
Zoology with Name No. 841 : : : 


serratus Forskal, 1775, as published in the combination Cancer serratus (Class 


Crustacea, Order Decapoda), a i on the Se pe List sa Speci Names in 
Zoology with Name No. 833 .. 


sexdentatum Stimpson, 1858, as published in the combination Camptandrium 
sexdentatum (Class Crustacea, Order Decapoda), Brett on the es List ce 
Specific Names in Zoology with Name No. 845 3 


sexdentatus Paulson, 1875, as published in the combination Archias sexdentatus 


(Class Crustacea, Order Decapoda), Placed 2 on the ee List of Spee Names 
in Zoology with Name No. 769. 


sexpes Fabricius, 1798, as published in the combination Cancer sexpes (Class 
Crustacea, Order Decapoda), at on the nee List a ARES Names in 
Zoology with Name No. 870" 


signatus Adams & White, as published in the combination Carpilus signatus (Class 
Crustacea, Order Decapoda), correction of date of, from 1848 to [1849] (correction 
of Opinion 73) te ‘ i Ss ae 


placed on the Official List of Specific Names in Zoology with Name No. 772 


Silurus Linnaeus, 1758 (Class Pisces), correction of entry relating to Be species of 
(correction of Opinion 77) - : 


Siren Hermann, 1783 (a junior homonym of Siren Linnaeus, 1766), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 808 


Page 
56 


178 


143 


158 


268 


374 


12 


11 


12 


Volume 1, Section D 


Sirena Fischer de Waldheim, 1808 (an Invalid Emendation of Siren Linnaeus, 1766), 
placed on the Official Index an cau and Invalid Generic Names in Zoology 
with Name No. 809 of : 


Sirene Oken, 1816 (an Invalid Emendation of Siren Linnaeus, 1766), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 810 


speciosa Dana, 1852, as published in the combination Nucia speciosa (Class 
Crustacea, Order Decapoda), Barats on the Ge List Ph PPPS Names in 
Zoology with Name No. 811 


spectabilis de Man, 1896, as published in the combination Platychirograpsus 
spectabilis (Class Crustacea, Order Decapoda), sare on the gills So List = 
Specific Names in Zoology with Name No. 903 


Speloeophorus Milne Edwards (A.), 1865 (Class Crustacea, Order iia 
completion of Ruling given in Opinion 73 : 


correction of reference for type species of 


Speocarcinus Stimpson, 1858 (Class Crustacea, Order Decapoda), completion of 
Ruling given in Opinion 85 : an i He a ot 


Sphaerium Scopoli, 1777 (Class Pelecypoda), placed on the childs List ie Generic 
Names in Zoology in Opinion 94, determination of gender of 


Sphaerozius Stimpson, 1858 (Class Crustacea, Order Decapoda), completion of 
Ruling given in Opinion 85 Sn we aS se _ id a4 


Sphargis Merrem, 1820 (a junior objective synonym of Dermochelys Blainville, 
1816), placed on the Official Index y peeeled and Invalid Generic Names in 
Zoology with Name No. 811 F : 5 


spinicarpus Milne Edwards (H.), 1852, as published in the combination Prionoplax 
Spinicarpus (Class Crustacea, Order Decapoda), i eaearet on the pei ci List be 
Specific Names in Zoology with Name No. 907 


spinosipes Stimpson, 1858, as published in the combination Scalopidia spinosipes 
(Class Crustacea, Order aCe einibd ee on the Cas List a pens Names 
in Zoology with Name No. 918. 


spinosus Rathbun, 1894, as published in the combination Erimetopus spinosus 
(Class Crustacea, Order Decapoda), seca on the Fea List Z are Names 
in Zoology with Name No. 794.. 


Spinturnix Heyden, 1826 (Class Arachnida), placed on the Official List a Generic 
Names in Zoology in Opinion 104, determination of gender of : 


Spirifer Sowerby, 1816 (Class Brachiopoda), placed on the Official List hs Generic 
Names in Zoology in Opinion 100, determination of gender of : 


19 


53 
55 


a2 


£59 


52 


346 


ai 


143 


156 


462 Opinions and Declarations 


spongiosus Adams & White, as published in the combination Iphiculus spongiosus 
(Class Crustacea, Order Decapoda), ¢ correction of date of, from 1848 to eal See 
(correction of Opinion 73) , A 


placed on the Official List of Specific Names in Zoology with Name No. 803 


sponsa Linnaeus, 1758, as published in the combination Anas sponsa (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 953 


sternberghii Stimpson, 1859, as published in the combination Xanthodius sternberghii 
(Class Crustacea, Order Decapoda), Sa on the pa ltage List 2 oe Names 
in Zoology with Name No. 928. 


Stilesia Stiles & Hassall, 1893 (Class Cestoda), placed on the a List S Generic 
Names in Zoology in Opinion 77, determination of gender of ; 


stridulans Wood-Mason, 1892, as published in the combination Psopheticus 
stridulans (Class Crustacea, Order Decapoda), placed on the vasclit List ol 
Specific Names in Zoology with Name No. 911 ’ 


Strigopsis Bonaparte, 1849 (an Invalid Emendation of Strigops Gray (G. R.), 1845), 
placed on the Official Index as Aas and Invalid Generic Names in iss 
with Name No. 572 aye 


Stringops Finsch, 1867 (an Erroneous Subsequent Spelling for Strigops Gray (G.R. ), 
1845), placed on the Official Index 2 sacar and Invalid Generic Names in 
Zoology with Name No. 573 : 


Stringopsis van der Hoeven, 1855 (an Erroneous Subsequent Spelling for Strigops 
Gray (G. R.), 1845), placed on the Official Index of a aon and Invalid Generic 
Names in Zoology with Name No. 574 


Strongyloides Grassi, 1879 (Class Nematoda), placed on the mer List eh Generic 
Names in Zoology in Opinion 66, determination of gender of , 


Strongylus Miller, 1780 (Class Nematoda), placed on the Official List et Generic 
Names in Zoology i in Opinion 104, determination of gender of 


struthersii Vigors, [1832], as published in the combination Ibidorhyncha struthersii 
(Class Aves), Bee on the Omen List eh Bneeiiee! Names in pceeey with Name 
INo-986 43. 


Strygops Des Murs, [1853] (an Erroneous Subsequent Spelling for Strigops Gray 
(G. R.), 1845), placed on the Once Index a Reise and Invalid Generic Names 
in Zoology with Name No. 575. 


sturio Linnaeus, 1758, as published in the combination Acipenser sturio (Class 
Pisces), placed on the CaaS List of Bases Names in fo with Name No. 
1072 


subglobosa Stimpson, [1871], as published in the combination Jliacantha subglobosa 
(Class Crustacea, Order Decapoda), pie: on the aes ree List or Sie Names 
in Zoology with Name No. 802.. 


Page 


22 


157 


20 


218 


218 


218 
155 


155 


Wie 


218 


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Volume 1, Section D 


subquadrata Dana, 1851, as published in the combination Fabia subquadrata (Class 
Crustacea, Order Decapoda), a on the eaiaiaiad List Sia sissies Names in 
Zoology with Name No. 861... . 


Succinea Draparnaud, [1801] (Class Gastropoda), placed on the Official List of 
Generic Names in Zoology in Opinion 94, determination of gender of ; 


sulcatus Swainson, 1820, as published in the combination Pteroglossus sulcatus 
(Class Aves), placed on the an a List ie 5 a Names in sates with Name 
Mo: .961 .. 


superba Costa, 1838, as published in the combination Portunus superba (Class 
Crustacea, Order Decapoda), Sapa on the ose Bist ee nthe Names in 
Zoology with Name No. 936... ; 


swainsonii Vigors, 1825, as published in the combination Gampsonyx swainsonii 
(Class Aves), ores on the rsiias List pe Hide a Names in naaieey with Name 
No. S34) ©... 


Syngamus Siebold, 1836 (Class Nematoda), placed on the Official List a Generic 
Names in Zoology i in Opinion 104, determination of gender of : 


Syngnathus Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 


Syngathus Rafinesque, 1810 (a junior homonym of Syngnathus Linnaeus, 1758), 
placed on the Official Index of re ig and Invalid Generic Names in Zoey 
with Name No. 812 ; : 


Syringothyris Winchell, 1863 (Class Brachiopoda), placed on the Official List Ed 
Generic Names in Zoology i in Opinion 100, determination of gender of 


tabacaria Linnaeus, 1758, as published in the combination Fistularia tabacaria (Class 
Pisces), placed on the pas List of SPS Names in ZR ES with Name No. 
1067 an 


Taenia Linnaeus, 1758 (Class Cestoda), placed on the Ofiiciabd List oe Generic Names 
in Zoology in Opinion 84, determination of gender of 


Telmessus Stal, 1866 (a junior homonym of Te/messus White, 1846), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 510 


tenuipes Dana, 1852, as published in the combination Carupa tenuipes (Class 
Crustacea, Order Decapoda), ah on the Ones List | iptae Names in 
Zoology with Name No. 780 _.. 


Teredo Linnaeus, 1758 (Class Pelecypoda), placed on the Official List oh Generic 
Names in Zoology i in Opinion 94, determination of gender of f 


tesselatus Latreille, as published in the combination Graspus tesselatus (Class 
Crustacea, Order Decapoda), correction of date of, from 1818 to 1812 ea eeta 
of Opinion 85) ; ae yt a ie my bed : “¢ 


placed on the Official List of Specific Names in Zoology with Name No. 882 


463 
Page 


14 


157 


174 


23 


177 


[55 


340 


346 


158 


368 


BS 


464 Opinions and Declarations 


Page 
Tetraxanthus Rathbun, 1898 (Class Crustacea, Order igs eae 48 See of 
Ruling given in Opinion 85 a > is sé ee 38 52 


Tetrias Rathbun, 1898 Se Crustacea, Order eee fperrgiges of sabia 
given in Opinion 85 , : 


Teuthis Schneider, 1784, a name placed on the Official Index of Rejected and Invalid 
Generic Names in Zoology by the Ruling given in Opinion 233, deletion of 
inadvertent entry relating to, made by the Ruling given in Direction 56 .. 346, 391 


Teuthys Lamarck, 1790 (an Erroneous Subsequent Spelling for Teuthis Linnaeus, 
1766), insertion of, on the Official Index of Rejected and Invalid Generic Names in 
Zoology in the entry rendered vacant therefor by the removal of the duplicate 
entry relating to Teuthis Schneider, 1784, eee made Ha the nee cos 
in Direction 56 .. : : : 391 


thalamita Stimpson, [1871], as published in the combination Melybia thalamita 
(Class Crustacea, Order Deen pa on the Poa List of saat Names 
in Zoology with Name No. 883. i 


Thalamitoides Milne Edwards (A.), 1869 (Class Crustacea, Order Decarrey 
completion of Ruling given in Opinion 73... 54 


Thalamonyx Milne Edwards (A.), 1873 (Class Crustacea, Order Dectenes) 
completion of Ruling given in Opinion 73... 54 


Thaumastoplax Miers, 1881 (Class Crustacea, Order Decapoda), completion of 
Ruling given in Opinion 85 a ; ts ie ae aK vy. ti 50 


Thysanosoma Diesing, 1835 (Class Cestoda), placed on the eiaes List sci Generic 
Names in Zoology in Opinion 77, determination of gender of . 159 


tiphia Linnaeus, 1758, as published in the combination Motacilla tiphia (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 1023... 183 


Tlos White, 1847 (a nomen nudum), placed on the ae Index xh Helly and 
Invalid Generic Names in Zoology with Name No. $11 é 61 


Tlos Adams & White (Class Crustacea, Order PeCADETEn correction of date of, 
from 1848 to [1849] (correction of Opinion 73) ; am s x te 56 


tobianus Linnaeus, 1758, as published in the combination Ammodytes tobianus (Class 
Pisces), placed on the Official List of Specific Names in Zoology with Name No. 
1064 sat me 4s ap Le t. Dye x ae au) ® 


tomentosus Dana, 1852, as published in the combination Actumnus tomentosus 
(Class Crustacea, Order ee np ee) aie on the aiciah List af ge Names 
in Zoology with Name No. 767. 3 


Tornatellina Pfeiffer, 1842 (Class Gastropoda), placed on the Official List cf Generic 
Names in Zoology i in Opinion 119, determination of gender of a 158 


Volume 1, Section D 


torquilla Linnaeus, 1758, as published in the combination Jynx torquilla (Class Aves), 
placed on the Official List of Specific Names in Zoology with Name No. 987 i 


transversus Rathbun, 1893, as published in the combination Opisthopus transversus 
(Class Crustacea, Order Decapoda), as on the Oe ial List be ie apn Names in 
Zoology with Name No. 890... 


trapeziformis Nauck, 1880, as published in the combination Holothuriophilus 
trapeziformis (Class Crustacea, Order Decapoda), pee on the Sila List uF 
Specific Names in Zoology with Name No. 872 ; 


tresselata Milne Edwards (H.), 1834, as published in the combination Lybia 
tresselata (an Erroneous Subsequent Spelling for tesse/atus Latreille, 1812, as 
published in the combination Grapsus tesselatus), placed on the Official Index Zz 
Rejected and Invalid Specific Names in Zoology with Name No. 338 ~ ; 


Triatoma Laporte, [1832] (Class Insecta), placed on the Official List sa Generic 
Names in Zoology in Opinion 104, determination of gender of 


Trichodactylus Latreille (Class Crustacea, Order Decapoda), correction of date of, 
from 1825 to [1828] (correction of Opinion 73) _ os ox ae a: 


Trichodactylus Dufour, 1839 (a junior homonym of Trichodactylus Latreille, 1828), 
placed on the Official Index sd paae: and Invalid Generic Names in n Zoology with 
Name No. 512 .. 


Trichostrongylus Looss, 1905 (Class Nematoda), placed on the eoeut List a Generic 
Names in Zoology in Opinion 66, determination of gender of : 


tridens de Haan, [1835], as published in the combination Ocypode (Helice) tridens 
(Class Crustacea, Order rea ates on the ee List a eee Names 
in Zoology with Name No. 868. 


tridens Kr@yer, 1837, as published in the combination Geryon tridens (Class 
Crustacea, Order Decapoda), pee on the os List us PE Names in 
Zoology with Name No. 863°. 


tridentata Milne Edwards (H.), 1853, as published in the combination Parathelphusa 
tridentata (Class Crustacea, Order Decapoda), i on the my List us 
Specific Names in Zoology with Name No. 818 ; : ee 


tridentata Miers, 1879, as published in the combination Pseudophilyra tridentata 
(Class Crustacea, Order Decapoda), uae on the ann List of eres Names 
in Zoology with Name No. 828.. 


tridentatus Milne Edwards (A.), 1879, as published in the combination Coenophthalmus 
tridentatus (Class Crustacea, Order Decapoda), placed on the ar List e 
Specific Names in Zoology with Name No. 782 ; : 4 


tristis Dana, 1852, as published in the combination Carpilodes tristis (Class 
Crustacea, Order Decapoda), places\e on the ae seiat List of maa Names in 
Zoology with Name INO sch Oe..." & 


62 


143 


56 


61 


155 


15 


10 


466 Opinions and Declarations 


Trombidium Fabricius, 1775 (Class Arachnida), placed on the Official List i Generic 
Names in Zoology in Opinion 104, determination of gender of 4 


truncatus Stimpson, 1860, as published in the combination Metopocarcinus truncatus 
(Class Crustacea, Order gata geass: on the esas List - arpa Names 
in Zoology with Name No. 885. 


Trypanosoma Gruby, 1843 (Class Mastigophora), placed on the Official List a 
Generic Names in Zoology in Opinion 95, determination of gender of 


tuberosa Stimpson, [1871], as published in the combination Osachila tuberosa (Class 
Crustacea, Order Decapoda), nv gs on the pap List he ae a Names in 
Zoology with Name No. 816... : 


tuberosus Pennant, 1777, as published in the combination Cancer tuberosus (Class 
Crustacea, Order Decapoda), Se on the Oe List ie Dia Names in 
Zoology with Name No. 790. : : G 


[Tunstall (M.)], 1771, Ornithologia britannica, all new names in, ruled as possessing 
no availability for nomenclatorial purposes, except Pyrrhocorax [Tunstall], 1771, 
validated by the Ruling given in Opinion 404 a We : 


addition of title of, to the Official Index ease: and Invalid Works in ee 
Nomenclature with Title No. 48 st 2 


typhlus Milne Edwards (A.), 1880, as published in the combination Bathyplax 
typhlus (Class Crustacea, Order Decapoda), Oe on the ein i List Zs Sue 
Names in Zoology with Name No. 844 


typicus Rathbun, 1898, as published in the combination Chasmocarcinus typicus 
(Class Crustacea, Order Dera ble on the Seg List a preeee Names 
in Zoology with Name No. 849. 


ungulata Linnaeus, 1758, as published in the combination Musca ungulata (Class 
Insecta, Order Diptera), pe on the 5 Gee List Hh pts Names in  LOMeey 
with Name No. 1056... 


urophasianus Bonaparte, 1827, as published in the combination Tetrao urophasianus 
(Class Aves), es on the lias List i. Bids Names in basins bi with Name 
No. 9674s: 


Utica White, 1847 Cpa Crustacea, Order ae ore meee of Sige given 
in Opinion 85 4 


Utica Hewitson, [1865] (a junior homonym of Utica White, 1847), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 513 


Valdivia Ragonot, 1888 (a junior homonym of Valdivia White, 1847), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 514 


Valdivia Shannon, 1927 (a junior homonym of Valdivia White, 1847), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 515 


Page 
144 


17 


158 


85 


85 
12 


13 


268 


W7a 
52 


61 


61 


61 


Volume 1, Section D 467 
Page 
Varuna Milne Edwards (H.), 1830 eae? Crustacea, Order 5 banat coe 
of Ruling given in Opinion 85 .. 52 
vesiculosa Alcock, 1896, as published in the combination Heteronucia vesiculosa 
(Class Crustacea, Order Decapoda), pees on the at List of os see Names 
in Zoology with Name No. 799. . | 
vigil Weber, 1795, as published in the combination Portunus vigil (Class Crustacea, 
Order Decapoda), Seat on the aoe! List nen Snegific Names in AC? with 
Name No. 938 .. 23 
vigil Fabricius, 1798, as published in the combination Portunus vigil (a junior 
synonym of vigil Weber, 1795, as published in the combination Portunus vigil), 
placed on the Official Index of in and Invalid peciee Names in Pei 
with Name No. 340 Fe 62 
violacea Isert, 1789, as published in the combination Musophaga violacea (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 989 178 
Vitrina Draparnaud, [1801] (Class Gastropoda), placed on the Official List a Generic 
Names in Zoology in Opinion 119, determination of gender of ‘ 158 
volans Linnaeus, 1758, as published in the combination Draco volans (Class 
Reptilia), placed on the ag List fe oe Names in RS with Name 
No. 1105 es é 373-374 
Volvox Linnaeus, 1758 (Class Mastigophora), placed on the ce List oe Generic 
Names in Zoology i in Opinion 75, determination of gender of : 156 
vulgaris Linnaeus, 1758, as published in the combination Sturnus vulgaris (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 1011 180 
vulgaris Milne Edwards (H.), 1837, as published in the combination Homarus 
vulgaris (a junior objective synonym of gammarus Linnaeus, 1758, as published 
in the combination Cancer gammarus), placed on the Offical Index of ge 
and Invalid Specific Names in Zoology with Name No. 362. f idk 2D 
vulturina Hardwicke, 1834, as published in the combination Numida vulturina (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 950 173 
wallicus Kerr, 1792, as published in the combination Psittacus wallicus (Class 
Aves), placed on the Official List of Specific Names in Zoology with Name No. 1032 —-183 
Xanthasia White, 1846 oe Crustacea, Order Wedge res cemukegen of a eulpe 
given in Opinion 85 a2 
Xanthodius Stimpson, 1859 (Class Crustacea, Order hr ea coments of 
Ruling given in Opinion 85 ; 3 : : 53 
Xenophthalmodes Richters, 1880 Shee Crustacea, Order pee eae 
of Ruling given in Opinion 85 .. 50 


468 Opinions and Declarations 


Xenophthalmus White, 1846 (Class Crustacea, Order Decapoda), completion of 
Ruling given in Opinion 85 a ig 45 = ae - me £3 


Zeus Linnaeus, 1758 (Class Pisces), completion of Ruling given in Opinion 77 


Zosimus Leach, 1823 (Class Crustacea, Order Decapoda), completion of Ruling 
given in Opinion 85 re aye ie eR ae 2 - 4 a 


Zosimus Schaufuss, 1886 (a junior homonym of Zosimus Leach, 1825), placed on the 
Official Index of Rejected and Invalid Generic Names in Zoology with Name No. 516 


Zozymodes Heller, 1861 (Class Crustacea, Order Decapoda), completion of Ruling 
given in Opinion 85 i ae Us an ae: e. hp a he 


Zozymus Milne Edwards (H.), 1834 (an Erroneous Subsequent Spelling for Zosimus 
Leach, 1825), placed on the Official Index of Rejected and Invalid Generic Names 
in Zoology with Name No. 517.. fh 4 iA a e: ue a 


Page 
53 


340 


53 


61 


52 


61 


Volume 1, Section D 


469 


PARTICULARS OF DATES OF PUBLICATION OF THE SEVERAL 
PARTS IN WHICH THE PRESENT VOLUME WAS PUBLISHED 


Part No. 
Dl 


D.2 
D.3 
D.4 
|B Bas 
D.6 
Dy, 
D.8 
ES 
D.10 
DAE! 
12 
13 
D.14 
BAS 
D.16 
Det] 
D.18 
| ed he 


Page Nos. 
1—46 
47—82 
83—94 
95—138 
139—152 
153—170 
171—210 
211—232 
233—242 
243—254 
255—264 
265—280 
28 1—292 
293—308 
309—318 
319—336 
337—364 
365—388 


389-470 
T.P.—XIV 


Contents of Part 
Direction 36 


Direction 37 
Direction 38 
Direction 39 
Direction 40 
Direction 42 
Direction 43 
Direction 44 
Direction 45 
Direction 47 
Direction 48 
Direction 49 
Direction 50 
Direction 51 
Direction 52 
Direction 55 
Direction 56 
Direction 57 


( Corrigenda 

J and Index, 
Foreword, Table 

ie Contents 


Date of Publication 
Ist September 1956 


Ist September 1956 

Ist September 1956 
17th September 1956 
17th September 1956 
17th September 1956 
19th October 1956 
19th October 1956 
19th October 1956 
26th October 1956 
21st November 1956 
21st November 1956 
21st November 1956 
21st November 1956 

Sth December 1956 
20th December 1956 
20th December 1956 
20th December 1956 


: 
L 
( 

J 


10th October 1957 


470 Opinions and Declarations 


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