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VAK(eer)
800 THE UNIVERSITY
[PA
GEOLOGICAL SURVEY
KANSAS.
CONDUCTED UNDER AUTHORITY OF THE BOARD OF REGENTS
OF THE UNIVERSITY OF KANSAS.
) AG FES Ie
PAZ HON POLOGEY .
Part ILI.
CARBONIFEROUS AND CRETACEOUS.
SAMUEL W. WILLISTON,
Faleontologist.
TOPEKA:
W. Y. MORGAN, STATE PRINTER.
1909,
ae
“a4
ADVERTISEMENT.
The publications of the University Geological Survey of Kansas
are for free distribution, on receipt of proper postage, or they may be
sent by express, charges collect.
REPORTS PUBLISHED.
Volume I, 1896— Reconnaissance Report on General Stratigraphy of
Eastern Kansas. (Edition exhausted.) Postage.
‘* TI, 1897 General Geology of Western Kansas................ 24 cents.
‘* JII, 1898—Special Report on Coal....... fo 8 a Dees
** TV, 1898— Paleontology, Part I, (on the Upper Cretaceous).... 32 ‘
** —-V, 1899—Special Report on Gypsum and Gypsum Cement
EEABLOUS ie See ea cre hee ee Meee eine ol eh kA es lo =
** VI, 1900 — Paleontology, Part II, (Carboniferous Invertebrates
md. Cretaceous ishGs) sta scee ek ofteics s cee ne 28
Report on Mineral Resources of Kansas for 1897.... 4 ‘
Report on Mineral Resources of Kansas for 1898.... 8
Report on Mineral Resources of Kansas for 1899.... 4
REPORTS IN PREPARATION,
Volume VII, 1900—Special Report on Lead and Zinc.
** VIII, 1901—Special Report on Oil and Gas.
Address all applications to
THE CHANCELLOR OF THE UNIVERSITY OF KANSAS,
LAWRENCE, KAN.
MEMBERS OF THE UNIVERSITY GEOLOGICAL SURVEY.
FRANCIS H. SNOW,
Chancellor, and ex officio Director.
EDGAR H. S. BAILEY,
Department of Chemistry.
SAMUEL W. WILLISTON,
Department of Historical Geology and Paleontology.
ERASMUS HAWORTH,
Depaztment of Physical Geology and Mineralogy.
FOR VOLUME VI.
SAMUEL W. WILLISTON, Paleontologist.
JOSHUA W. BEEDE, Assistan’t Pa!eontologist.
ALBAN STEWART, Assistant Paleontologist.
SYDNEY PRENTICE, Artist.
Chancellor F. H. Snow,
Ex officio Director of the University Geological Survey:
SrrR—I have the honor herewith to submit to you for your ap-
proval the second part of my report upon the Paleontology of Kan-
sas, to constitute Volume VI of the University Geological Survey of
Kansas. It is with great pleasure that I again acknowledge to you
my appreciation of the uniform encouragement you have shown in
the prosecution of the work. Respectfully,
SAMUEL W. WILLISTON.
DEPARTMENT OF HISTORICAL GEOLOGY,
UNIVERSITY OF KANSAS, JUNE 1, 1900.
CONTENTS:
PAGE
UNIVERSITY, GEOLOGICAL SURVEY PUBLICATIONS. 55. 2 6 une eee iil
MEMBERS OF THE UNIVERSITY GHOLOGICAT, SURVEY «25.2000 eee eee iv
LETTER OR TRANSMITTAL A200) 15 yaks ia ehe weie 6 Leh ce Ee V
PREBAGH Coos 5 aes eS SAR ee coe even a tee aed PS IS SER ete rc a Vii
PART I.—CarpBoniFrerRous INVERTEBRATES. By Joshua W. Beede........ 1
Entroductions ..2. 2 ae ee Sco lee ae eee 3
Classification’of: Mossilsso..c.200 eek Oe eae ee ee eee 5
PTOCOZO «oie a nceedesh ate ww) ash oes pote Snes lek oN 9
Coelenterata... 2). <8 Sy Seah Bale ae Le > 11
Kichipoder ma ta i ee wees eae so Ie ee ene 26
Molluscoidea....... 2.50 Sets 2 oe eee eee oo eae ee 51
Molltise@aee nes oie oh a eee ek ee Nee ee 107
Explanation-of : Plates. ces. teva eee. Go ee eee 175
Plates. 1.t0%x11, nchisivess: 2.4500 See eo eee cee 189
PART .TI; CRETACEOUS FISHES: Ah. ace ee eee eee 235
FRONTISPIECE, Kansas Cretaceous Sea. Plate xxXIIJ.................... 234
SELACHIANS AND Pycnoponts. By S. W. Williston...... BU cs ea 237
Myliobatidver: ...)5 2. OS her Beene ore ee ee ee 237
Sy Tiree M8 cakes Soltero 244
iaminidseteas ah ees cht oO ee ae 246
Pycnodontidie.. oi... hohe sid dos sete SO ee 254
Lepidosterdee is) A5 os eSi ae Mo ee eee 256
TELEOSTS OF THE UPPER Cretaceous. By Alban Stewart...... de haan 257
Pntroductionac. 2 cee. 2 <6. eae de oS a 257
Teleostei:. 6b eects 6 LGR ee eee 259
Ichthyodeetidees ig 826.34 ae ok ee ee ee 262
Saurodon tides <.ic che c64 oi gehen eT ea 310
Stratodontidee.2) 2% 46600. ne eee eee eee ies are. tes 326
Osteorlossidae. witch lo ee eee ert ene ates MS 340
Salmon idee ool oi cc kali ey Le ote ee era ee 348
Pachycormidze 4... A 6s We (ah RRC RRS ER ae ee ee is SUG a 362
(©) Lvi of:1(6 iM ea es nn Gad sido e oe oudo Gace By
Hnchodontidae: yf. bok A. ene oe ree 373
Dercetidze. By S. W. Williston sejs5ee eee ee eee 380
Mugillide:.-.. 2. ER MPT nee ke, SN 8 we dua OSS ob ob ied Wao 66 5% 383
Range of American Cretaceous Teleosts 72. 42254-05-- 22 nee eae 385
73% 0) 012) 01 6h 5. Geen en neni Ars aS Rie tL oie cnet S ion 5 Bla aelons aip a0 ono 391
Explanation: of Plates. i/o. eae ae eee ee ee ee eee ee 394
Plates Xxrv towxxi1, inclusive... seo es eo eee ee 405
GENERAL: IN DEX®. 5 chaos; fcc) ne Osho lays ined oo SRE Dn ge eT en eee 509
PREFACE.
THE present volume on the paleontology of Kansas deals with
the Carboniferous Invertebrates and the Cretaceous Fishes.
The latter part has been made as complete as the present state
of the collections and knowledge will permit; the part on the
Carboniferous Invertebrates leaves yet several classes to be
treated, but which, it is hoped, will find a place in the next or
succeeding volumes of the series. The work on the fishes com-
pletes our present knowledge of the paleozoology of the Kansas
Cretaceous, with the exception of that of the pterodactyls and
plesiosaurs. It was the intention of the writer to make the dis-
cussion of these two groups of reptiles a part of the present vol-
ume, but the stress of duties in other directions has rendered
this impossible; he can only hope to treat them fully in the
early future.
It has been the aim to so treat the fossils of Kansas in the
preceding and present volumes of this series that the work will
be of immediate use to the people of Kansas. For this reason
ful] descriptions and figures of every species have been given,
so far as it is possible. Some of the descriptions and figures
might have been omitted, giving merely a catalogue, of use only
to the scientist, but such a plan would defeat the chief object
intended—the preparation of manuals or texts for the use of the
student in Kansas geology.
Messrs. Beede and Stewart, the authors of much the largest
portion of the present volume, have been students and assist-
ants in the paleontological laboratory of the University. While
(vii)
vii University Geological Survey of Kansas.
their work has been done under general directions and advice,
the merits or faults of their papers are theirown. The zeal,
intelligence and earnestness which they have devoted to the
work will be sure to have produced valuable results. Doctor
Beede, it is hoped, will be able in future papers to complete the
review of the paleozoic invertebrates.
At the present time, Mr. E. H. Sellards, assistant in the de-
partment of paleontology, has well along a work on the paleo-
zoic plants which, it is expected, will soon be published.
The writer desires to thank Mr. Sydney Prentice, who has
made all the drawings of this volume, for the care and faithful-
ness he has given to the work.
SAMUEL W. WILLISTON.
PART I.
CARBONIFEROUS INVERTEBRATES.
By JOSHUA W. BEEDE.
Plates I to XXII.
A thesis presented to the Faculty
of the University of Kansas for the
degree of Doctor of Philosophy,
June, 1899,
INTRODUCTION.
FORAMINIFERA TO PELECYPODA, INCLUSIVE.
this report the terms Lower and Upper Coal Measures are
only used relatively. The lowermost portion of the Coal
Measures is referred to as the Lower Coal Measures, and the
upper part as the Upper Coal Measures. As there is some
controversy concerning the boundary lines between these two
divisions, and as the material at hand is not of a decisive
nature, no attempt is made to draw any sharp division line, if
indeed one can be drawn otherwise than arbitrarily. The
collections studied, covering the entire Coal Measures area in
a very general way, show no very distinct faunal division
between the upper and lower rocks of the Coal Measures.
There are several species present in the lower portion of the
rocks that are absent in the upper, but for the most part the
species disappear gradually.
Cleiothyris roysii and Chonetes mesoloba are common to the
lowermost strata only, and apparently below what is called the
Erie limestone by the University geologists. Somewhat above
this limestone Lophophyllum westii fades out, and in the Garnett
limestone are found the last of Productus cora americanus and a
wide variety of the same species, and also Plewrotomaria missou-
riensis. However, this is considerably above what is generally
taken to be the limit of the Lower Coal Measures. For con-
venience, until a better and more complete study of the faunas
can be made, the limestone known in the Kansas reports as the
Erie limestone will be considered as the base of the Upper Coal
Measures.
In almost all cases the labeling of the fossils herein described
is not specific enough to tell whether or not they are from below
this limestone; consequently, their exact position with refer-
(3)
+ University of Kansas Geological Survey.
ence to it can only be guessed at. Nevertheless, it is distinct
enough to show that, in the Kansas area at least, there is no
abrupt change in the faunas of the upper and lower parts of
the Coal Measures, though the statement has been made that
‘‘the change in faunal features is striking.’’’
The upper limits of the Coal Measures, as defined by Prosser,,.
are followed in this report.
The fauna of the Coal Measures of Kansas presents some
very interesting and important features. The vertical range
of species is very great for both brachiopods and pelecypods.
The fauna also seems to be a young one in some respects,
though, as a whole, it is certainly that of the Coal Measures.
There are several species ranging through it which are charac-
teristic of the English Permian, some of them being even
restricted to the lower portion. This would not be so very
remarkable were they all brachiopods, but several of them are
pelecypods.
For the area and thickness of the Coal Measures, the reader
is referred to Volume III of these reports.
The writer wishes to gratefully acknowledge the kind assist-
ance of Dr. S. W. Williston in many ways in the preparation of
this report; also to thank Prof. Charles 8. Prosser, of the Uni-
versity of Ohio, and Mr. Charles Schuchert, of the United States.
National Museum, for the loan of many valuable specimens and
for offering many valuable suggestions and encouragement in
the work. To Mr. Austin I. Rogers, of the University, much
credit is due for assistance. The drawings for the illustrations.
accompanying this article were made by Mr. Sydney Prentice.
1. Keyes, Amer. Geol., xxiii, p. 303, 1899 (May).
CLASSIFICATION OF FOSSILS
DESCRIBED IN REPORT.
pee
Se re) age
jo ac S = on — =
ee oe ae eS
x lal n 0 ro) om &
PROTOZOA.
BAITZOPrODa.
HAGEOA NUT N TMA: Gacteeue. 2 loo. ccc kas p. 9
Fusulinidz.
Fusulina.
C@LENTERATA.
Porifera.
AS LOIS As ERR ee eee 2 eo bl
PHARETRONES.
Somphospongia.
SYCONES.
Amblysiphonella.
Cnidaria.
AMT OAs We, is ois a Patee baa p. 16
Zaphrentide.
Lophophyllum.
Cyathophyllide.
Campophyllum.
Axophyllum.
Favositide.
Michelinia.
Trachypora.
Auloporide.
Aulopora.
Cladochonus.
Syringoporide.
Syringopora.
Cheetetide.
(Chotetes.
{.))
Carboniferous Invertebrates.
CLASSIFICATION OF FOSSILS— Continued.
|
—| KINGDOM.
- q
s ° 3
Bee % S od s :
fo) 5 RQ > oa = x
She x A =
Be eS N 3 ae 5 5
ey M o n fo) em &
| | | | | | |
ECHINODERMATA.
Pelmatozoa.
GOREN OL DIBA ice Se SER oo Le eee p. 26
FISTULATA.
Poteriocrinide.
Scaphiocrinus.
Zeacrinus.
Ceriocrinus.
Phialocrinus.
Hrisocrinus.
Hupachycrinus.
Hydreionocrinus.
Agassizocrinide.
Echinozoa. A gassizocrinus.
LE OCHIN GOT DAE Ge se a eee ere p. 46
PERISCHOECHINOIDA.
Archeeocidaride.
Archeocidaris.
Melonitide.
MoLLUSCOIDEA. Oligoporus.
BREA CHTOPODA Se kcorus - py ak
ATREMATA.
Lingulide.
Lingula.
NEOTREMATA.
Discinide.
Orbiculoidea.
Craniide.
Crania.
PROTREMATA.
Strophomenide.
Derbya.
Meekella.
Productide.
Chonetes.
Productus.
Aulacorhyncus.
Orthide.
Rhipidomella.
Einteletes.
~I
BREDE. | Carboniferous Invertebrates.
CLASSIFICATION OF FOSSILS — Continued.
|
= zs 868 : é : S
Q rat = 2 S ee} = < :
s 15 5 2 > = 5 = s
Pee Oe 8S OR 8 E :
— Ns S N) ~) jae} Db oe &
=< Se mM o 2 (o) ™ fy ic)
| | | | |
MOoLLUSCOIDEA — continued.
BRAC HIOPODA — continued.
TELOTREMATA.
Rhynchonellide.
Pugnax.
Terebratulide.
Dielasma.
Spiriferide.
Spiriferina.
Spirifer.
Ambocelia.
Reticularia.
Athyride.
Hustedia.
Cleiothyris.
Seminula.
MOLLUSCA.
Pd YETTA, ee eee oa ie ee RSS Dy LOY
ASIPHONIDA.
MoNnNoMYARIA.
Anomiide.
Placunopsis.
Limide.
Lima.
Pectenide.
Entolium.
Aviculopecten.
Aviculide.
Pteria.
Limopteria.
Pseudomonotis.
Posidonomya.
Mytilide.
Modiola.
Myalina.
Pinnide.
Aviculopinna
Pinna.
Carboniferous Invertebrates.
CLASSIFICATION OF FOSSILS— Concluded.
s 3 a : ed
o) S) q : % ; a
ee ee ee tee ys 8
O ae 2 me SS} A 3 ig =
eet ees Se e : :
x n o n o) nn c S
| | | | | | |
ca
Mo.Liuusca — continued.
PELEC Y PODA— continued.
ASIPHONIDA— continued.
HOMOMYARIA.
Arcide.
Macrodon.
Nuculana.
Nuculide.
Nucula.
Yoldia.
Trigoniidez.
Schizodus.
SIPHONIDA.
INTEGRIPALLIATA.
Solemyide.
Solenomyda.
Astartide.
Pleurophorus.
Astartella.
Cardiide.
Conocardium.
Cypricardinia.
SINUPALLIATA.*
Cardiomorpha.
Hdmondia.
Allorisma.
Sedgwickia.
Chenomya.
* Of uncertain position in this order.
DESCRIPTION OF FOSSILS.
FORAMINIFERA.
The foraminifera are an order of that class of animals known
as rhizopods, the name meaning root-footed. The rhizopods
are very minute animals, resembling, under the microscope, a
mass of jelly full of little bubbles. They move about by push-
ing out portions of this jelly-like substance into thread-like pro-
trusions, which may be absorbed back into the animal. This
is how they get the name ‘“‘root-footed.’’ Food particles are
taken into the body anywhere, as there is no mouth, and digested
and the waste portions thrown out wherever they happen to
be, as there are no digestive organs. The animal can best be
understood as a minute bit of jelly-like substance, called sar-
code, capable of motion and the assimilation of food. Some
of these rhizopods have shells with minute holes in them
through which the thread-like feet are protruded. These feet
often interlace, forming a network about the animal. These
animals are called foraminifera on account of the little holes in
the shell. They live in both fresh and salt water, but are much
more numerous in the oceans, where, minute as they are, their
shells make up masses of deposits which become hardened
into limestone and extend over vast areas.
The only foraminifera which have been made known from
the rocks of eastern Kansas are those often called ‘‘ petrified
wheat.’’ Their scientific name is Fusulina secalica, the latter
name meaning arye grain. ‘They are very numerous; often
layers of limestone from two to ten feet thick, made up almost
entirely of these shells, extend across the state from north to
south. For further information, the reader is referred to Mr.
McClung’s article in Volume IV of this Survey, entitled
)
‘‘ Microscopic Organisms of the Upper Cretaceous,’’ on page
415.
2—vi (9)
10 University Geological Survey of Kansas.
FUSULINA.
Fischer, Oryct. Moscou, p. 126, pl. XVIII, ff. 1-5, (1837).
Fusulina secalica. Plate I, figs. 1, 1b.
Miliolites secalicus Say, Long’s Exped., p. 151, foot-note, (1823).
Fusulina cylindrica Fischer, ibid., (1837); Meek, Fin. Rep. U.S. Geol.
Surve (Neb. op. 140)" plicit. 2p), a. ofa 2 ile aleinesa pO berven tli orien lo
(1872).
For further synonomy, see Weller, Bull. U. S. Geol. Surv., 153, p. 280,
(1898).
Meek’s description: ‘‘Shell small, fusiform or subcylin-
drical, more or less ventricose in the middle, somewhat obtusely
pointed at the extremities, which generally have the appear-
ance of being a little twisted. Surface smooth excepting the
septal furrows, which are moderately distinct, more or less
regular, and a little curved as they approach the extremities.
Aperture apparently linear, and not visible as the specimens
are generally found. Volutions 6 to 8, closely coiled, the spaces
(near the middle) being rarely more than twice the thickness
of the shell walls. Septa about 20 to 33 in the last or outer
turn of adult specimens, counting around the middle; compara-
tively straight near the outer walls, but strongly undulated
laterally within; foramina passing through the walls, moder-
ately distinct in well-preserved specimens—as seen under a
high magnifying power, in transverse sections near the mid-
dle of the shell, somewhat radiating, and numbering in the
outer turn of a medium-sized specimen, from 12 to 20 between
each two of the septa. Varying considerably in size and form.’’
Range and distribution: Very abundant throughout the
Upper Coal Measures of the state.
There have been two species and three varieties of this little
shell described from the rocks of the United States. These varie-
ties were considered species by their author at one time and were
later reduced to varieties by him. Nearly all the forms that
have been described are found in the rocks of the state. They
are: Fusulina gracilis, robusta, and F’. ventricosa Meek, as well
as F. elongata Shumard. In dividing these foraminifera into
species, it is to be remembered the conditions for their growth
were extremely favorable here during Carboniferous time, for
BEEDE. | Carboniferous Invertebrates. 11
their skeletons make solid strata of limestone several feet in
thickness, extending over vast areas. Under such conditions
we always find a single species appearing in many forms, and
consequently great care should be taken in separating them into
species and varieties. So far as the external form and appear-
ance of the Kansas Fusulinas are concerned, I think it will be
impossible to divide the species, though a more careful study
of the minute structure of the interior of large numbers of in-
dividuals may reveal certain constant variations worthy of va-
rietal or even specific recognition. But even then, when the
fact is taken into account that they are but simple undifferen-
tiated animals, susceptible of such great variations from environ-
ment, great care would have to be taken in not making too
many divisions of them.
Say’s description of Miliolites secalicus is good enough, when
the locality from which his specimens were taken is brought
into account, so that it is practically impossible for it to be
missapplied, and consequently that specific term, together with
Fischer’s generic designation, will stand.
SPONGIZ.
The sponge is a rather loose collection of single cells variously
grouped into one mass, forming a compound organism. The
connection of the different cells with each other is slight. These
cells are arranged in a series, so that canals are formed running
through the sponge, the cells facing the canals. The inner ca-
nals are generally larger where they flow together for the water
to flow out, and this enlargement is called a ‘‘cloaca.’’ Water is
kept flowing through the canals by means of rapidly moving
cilia which some of the cells possess. These little whips also
aid in the capture of particles of food, which are taken from the
water as it flows through the canals.
The cells of the sponge are held together by horny, calcareous
_ or flinty needles called spicules. The sponge of commerce, as
we generally see it, is not the entire animal, but only its skele-
ton, made up of horny fibers so interlaced as to form a network.
12 University Geological Survey of Kansas.
The larger holes in it indicate where the larger canals were lo-
cated. In most of the fossil sponges these fibers of the skeleton
were made of lime or flint, and are known as calcisponges and
silicisponges, respectively. The spicules are often compound
and possess several points. Some are simple, like a needle
pointed at each end; some have three points, some four, and
some six; some are anchor-shaped and some are dagger-shaped.
Most sponges are attached to rocks or other objects during
their life, but some are free. They inhabit both fresh and, salt
water, but are, for the most part, found in the sea, in moder-
ately shallow water. The only fossil sponges known from the
Kansas rocks are calcisponges in which the spicules are rarely
preserved.
SOMPHOSPONGIA.
Beede, Kans. Univ. Quart., VIII, p. 128, (1899).
_A globular to mushroom-shaped calcisponge, attaining a large
size, and generally possessing a more or less spherical-shaped
cloaca near the base; the canals are all very irregular and
crooked, distributed over the entire surface, and moderately
large. A rather thick dermal layer is present. They were
free, apparently resting with the base in the mud in the adult
stage.
Somphospongia multiformis. Plate I, figs. 6-10; plate IT, figs. 1-5.
Somphospongia multiformis Beede, Kans. Univ. Quart., viz1, p. 128, pl.
XXXL, ff. 1-10, (1899).
A small to very large sponge, varying in form from globular
to mushroom-shaped, free, and gregarious. Connecting with
the cloaca there is an irregular, branching canal system, which
communicates with the exterior over the whole surface, though
in large individuals they are smaller and probably nearly use-
less at the base. These canals are.very irregular in shape, and,
when viewed on the,surface, appear to be labyrinthine; they
become smaller and less numerous as they proceed inward to-
ward the cloaca. When not worn the entire sponge is covered
with a moderately thick dermal layer, the folds in which form
the walls of the canals. There is no sign of attachment in any
of our specimens, and the young ones seem to have rolled about
BEEDE. | Carboniferous Invertebrates. 13
until they had gained some considerable size, as the pores are
about equally developed all over them; they are somewhat
globular in form. Where they were abundant, as is generally
the case wherever they are found, they soon come in contact
with each other and form a solid mass, sometimes appearing to
coalesce, but generally in breaking they part along the line of
contact, and neither specimen seems to be ruptured. As yet
spicules have not been positively made out. There are no sili-
ceous spicules, and several thin sections have failed to show
any calcareous ones. The absence of siliceous spicules and
chert in the specimens, and the absence of chert in the rock,
makes it practically certain that they are calcareous sponges.
There is on weathered specimens, where the dermal layer has
been removed, a peculiar, more or less haphazard arrangement
of pits, surrounded by elevations, which is probably caused by
an internal calcareous skeleton, composed of fused spicules.
The different individuals of this species vary from half an inch
to a foot in diameter, but seldom are more than six inches high.
They are found in abundance in the northwestern part of
Atchison, in western Doniphan and eastern Brown counties. It
is not uncommon to find them making up a stratum of lime-
stone six inches thick. They are confined to a single narrow
horizon in the Burlingame shales.
The cloaca is generally filled with limestone, which, except
at the center, is arranged in concentric layers as it was filtered
in, giving the cloaca and the parts immediately surrounding it
much the appearance of a concretion.
The sponge evidently belongs to the Pharetrones, and appears
most closely related to Corynella and Stellispongia. It differs
from the former in not having the cloaca funnel-shaped, and
the fact that the cloaca does not terminate below in vertical
branching tubes any more than it does above, and possesses no
distinct exhalent aperture. It is much more closely related to
the latter, but is simple, and appears quite different in its spic-
ules, while the cloaca is confined to the base.
14 University Geological Survey of Kansas.
AMBLYSIPHONELLA.
Amblysiphonella prosseri. Plate I, figs. 2-2f; plate II, fig. 6.
Amblysiphonella prosseri Clarke, Amer. Geol., xx, p. 387, pl. xx1t, ff.
1-9, (1897).
Clarke’s description : ‘‘ The bodies from Netawaka and Weep-
ing Water are simple subcylindrical individuals, straight or
gently curved, the largest fragment measuring 70 mm., and in-
dicating an entire length of not exceeding 100mm. The fossils
are from a calcareous shale and have, for the most part, been
somewhat compressed. Their interior cavities, the cloaca and
interseptal chambers, are filled with compact gray limestone,
distinctly and fine oolitic, and their exteriors are frequently
entangled with encrusting Bryozoa and the remains of other
fossils. The septate or annulate aspect of the exterior is always
shown, and when this external surface is free from other or-
ganic remains and cleansed from the attached matrix, it presents
the aspect of a Fistulipora or of a small-celled Alveolites; that
is, the meshwork of the skeleton is made up of polygonal cells,
all of a small size, not always opening directly outward, but in
places frequently presenting oblique apertures. So fine is this
superficial network, and so uniform the size of the cells, that
one might casually interpret the entire fossil as a macerated
Orthoceras, overgrown with an encrusting bryozoan.
‘One of the specimens is preserved with its aperture entire,
which shows it to have been a simple, narrow, circular cavity.
On cutting these bodies along their longer axis, we observe, first,
a continuous central cloaca, relatively much wider than would
be the siphon of an orthoceran of the same size, but slightly
constricted at intervals, where its walls meet the septa. This
cloaca is delimited by a well-defined circular wall, and thus has
no communication with the septal chambers or the cavities of
the annuli, except through the perforations in this wall. The
septa are at quite regular intervals and are convex upward. On
the gastral surface they project slightly inward, as observed,
into the cloaca. Each of these septa presents a former apertural
surface, and the sponge affords, thus, an interesting instance of
periodical intensity of growth force. The walls of the sponge
7~—"rr” EE ge ee
“2? 2, so at
: .
BEDE. | Carboniferous Invertebrates. 15
thus exposed in section are very thin, approximately uniform in
this respect, and are all perforated with straight, simple, rela-
tively large canals. Those traversing the gastral walls (ex-
halent pores) are larger than the rest and appear to be of
uniform size. The canals perforating the septal (that is, aper-
tural) and exterior walls are inhalent pores, and, with this
necessary interpretation, the septal cavities may properly be re-
garded either as chambers for the accumulation and discharge
into the cloaca, or as true ciliary chambers. We find that, for
the most part, these skeletal walls have been, perhaps by sec-
ondary changes, converted into crystalline calcite, and this
change has obliterated the spicular structure, and in some
places the perforate structure of the walls. Elsewhere, espe-
cially on the external walls, there has often been a deposit of ad-
yentitious calcite which, in sections, gives the wall an unusual
thickness. It is to be noted that there is no breach in the con-
tinuity of the external wall of any given annulation. The septal
or enclosed portions of the wall do not meet the outer or exposed
part as they meet the gastral wall, but the entire external wall
is arched from the peripheral base of the chamber beneath to
the apertural margin of the cloaca.’’ Thin vesicular tissue
often extends from the gastral to the dermal wall and some-
times between each other.
Range and distribution: Upper Coal Measures; Thayer,
Topeka.
I have a single specimen of this species from the Topeka
limestone. It agrees very well in all respects with the speci-
mens figured by Clarke. The smaller end was broken away
before fossilization, making it appear to have the aperture at
the smaller end of the specimen, and the septa sag downward in-
stead of curve upward. ‘There are several specimens probably
of this species from Thayer, which are too poorly preserved in-
teriorly to determine specifically, but the exterior resembles
very much this species, to which it is referred. Associated
with the above, at Thayer, is a smaller variety, very much more
constricted at the juncture of the septa and the outer wall, giv-
ing it very much the appearance of a row of large beads set
16 University Geological Survey of Kansas.
together. It has a very different external aspect from the
specimens of the form above mentioned from the same place,
and is probably a different species, but until specimens can be
secured showing the internal characters, it is referred provi-
sionally to this species.
ANTHOZOA.
Corals are exclusively marine animals. The reef-building
corals are found in comparatively shallow water, while some
other forms are found as deep as 1500 fathoms. The corals here
treated are all closely related to the reef-building forms, and
consequently indicate that the rocks in which they were formed
were laid down in shallow water. The coral is an animal with a
distinct digestive sac and body cavity. There are three layers of
the body walls—the outer layer or ectoderm, the middle layer
or mesoderm, and the inner layer.or endoderm. The hard parts
or skeleton are deposited by the inner layer between the inner
layer and the middle layer. This hard part forms a complete
layer around the lower part of the animal beneath the skin.
There are thin walls of the hard part projecting inward in the
folds of the mesentery. These walls are called septa. There
are often cross-plates or table- Looe
like platforms across the lower
part of the hard parts extend-
ing clear across the body ; these
are called tabule. Around the
mouth of the polyp, as the ani-
mal is called, there is a row of
tentacles, or feelers, which are
used to create currents of water
and assist in securing food.
Corals very often increase by
budding; that is, by a little
bud-like projection growing out
5 ° G.1. Transverse section of a simple coral
of one side of a coral and ina Eating NicholnGay: s, septum; c, columella.
wre
Pe a
. . ey ee
BgxEvgE. | Carboniferous Invertebrates. 17
short time becoming a polyp exactly like the first and connected
with it. Ina little while such a coral will form a cluster.
Some kinds bud much more abundantly than others, when the
cluster becomes a solid mass and is called a compound coral.
At Fort Scott there is a large limestone stratum almost en-
tirely made up of a coral called Chetetes milleporaceous. At the
time when the corals were living it was a small coral reef.
The other fossil corals in Kansas are much more rare, though
large masses of two other families are often found.
LOPHOPHYLLUM.
Milne-Edwards and Haime, Brit. Foss. Corals, pl. LX VI, (1850).
Lophophyllum profundum. Plate II, figs. 7-7b.
Cyathaxonia profunda Milne-Edwards and Haime, Mon. des Polyp. Foss.,
p. 323, (1851); etc.
Cyathaxonia prolifera McChesney, Descr. New Pal. Foss., p. 75, (1860);
ibid., pl. 11, ff. 1-3, (1865); ete.
Lophophyllum proliferum Meek, U. S. Geol. Surv. Neb., p. 149, pl. v,
ff. 4a, b, (1872): etc.
Lophophyllum profunda Foerste, Bull. Den. Univ., 111, p. 136, (1888);
Worthen, Geol. Surv. Ill., vir, p. 79, pl. x, ff. 14, 14a, (1890).
Meek’s description, in part: ‘‘Corallum elongate-conical,
more or less curved, or sometimes nearly straight, tapering to a
pointed base; epitheca very thin, with more or less distinct
encircling wrinkles and strice of growth, crossed by longitudinal
strie ; rarely sending off a few spines near the base. Calice
nearly, or quite circular, moderately deep; columella promi-
nent, compact in texture, compressed above, with its longer
axis coincident with the general curve of the corallum; septa
from about 30 to 50, every alternate one being generally con-
siderably shorter than the others, which latter are extended to the
columella, near which they are sometimes a little tortuous.’’
Columella formed by the enlarged prolongation of one of the
septa; tabule moderately remote, arching from the columella
outward and downward, sometimes a little upwards at first.
Septa striated obliquely upward. Columella striated by some-
what revolving longitudinal strie. Measurements: Length,
30 mm.; width of calyx, 9 mm.
Range and distribution: Upper and Lower Coal Measures ;
18 University Geological Survey of Nansas.
Fort Scott, Marmaton, Bourbon county, Thayer, Olathe, Kan-
sas City, Lawrence, Lecompton, Topeka, McFarland, Grand
Summit. Common throughout the Coal Measures.
In young specimens, the septa, except the cardinal, do not
reach quite to the columella.
Lophophyllum westi. Plate II, figs. 8, 8b; plate III, fig. 12.
Amplexus westi Beede, Kans. Univ. Quart., vir, p. 17, (1898).
Corallum simple, snbcylindrical to attenuate-conical, straight,
curved or geniculated, epitheca thin, longitudinal strize promi-
nent, and concentric lines and undulations of growth distinct.
The septa extend about half the distance to the center, 18 to 24
or more in number; the counterseptum extends to the center,
where it is somewhat enlarged ; others of about equal length ;
indications of secondary septa visible but very small. The
tabule are well developed 1 to 3 millimeters distant and
reaching from wall to wall; on leaving the walls they are di-
rected obliquely upward for a short distance, and then, slightly
arching and undulating, cross the center, occasionally branching
at or near the bend. Length, 60 mm. ; diameter in larger part,
9mm.
Range and distribution: Upper Coal Measures ; Kansas City,
Lecompton, Neosho county.
This species is very variable in size and shape as well as de-
gree of development of the septa. It was at first referred to the
genus Amplexus, as it seemed to have no columella, but more
material showed that many of the specimens had the cardinal
septum prolonged to or beyond the center and slightly enlarged,
which seems to make it agree more closely with Lophophyllum
than Amplexus. That the species is intermediate between the
two genera there can be but little doubt, as different specimens
of the same species seem to possess the characters of each genus
about equally.
It is the fossil usually identified by collectors as Cyathaxonia
distorta Worthen, and indeed there is no external difference
visible (judging from Worthen’s figures ) in the smaller, more
distorted forms. But if his species is a true Cyathazxonia it is
Breve. | Carboniferous Invertebrates. 19
of course specifically distinct from our coral. I have been un-
able to secure specimens of Worthen’s species for comparison,
and in his description he does not mention the generic charac-
ters nor does he figure the interior of the specimen so as to show
whether or not it had any tabule. If it turns out to be a Lo-
phophyllum, his name will, of course, have priority.
CAMPOPHYLLUM.
Milne-Edwards and Haime, Brit. Foss. Corals, pl. LX VIII, (1850).
Campophyllum torquium. Plate IV, fig. 1; plate V, figs. 1-4.
Cyathophyllum (vermiculare?) Owen, Geol. Rep. Wis., lowa, and Minn.,
tab. iv, fig. 3, (1852).
Cyathophyllum torquium Owen, ibid., tab. rv, f. 2.
Cyathophyllum flecuosum (?) Owen, ibid., tab. rv, ff. 3a, b.
Campophyllum torquium Meek, Fin. Rep. U. S. Geol. Surv. Neb., p.
145, pl. 1, ff. la—d, (1872); ete.
Meek’s description, in part: ‘‘Corallum simple, attaining a
rather large size, elongate conical, and often geniculated or bent
when two or three inches in length, but becoming nearly
straight, subcylindrical and considerably elongated in the larger
half of adult individuals. Epitheca thin, with small encircling
wrinkles and strong undulations of growth, showing no traces
of septal cost when unabraded, but, where even slightly worn,
exposing the regularly disposed septa and thin intervening dis-
sepiments distinctly. Calice circular or slightly oval, compara-
tively shallow, with thin margins, from which it slopes rather
steeply inward for some distance and then descends very
abruptly into a deeper, narrow, central depression ; provided
at the outside of the general curve of the corallum with a mod-
erately distinct septal fossula, formed by the shortening of one
of the primary septa, and the bending down of the tabule at
that point. Principal septa from 30 to 48, extending from
about one-half to two-thirds the distance from the exterior
toward the center, stout and usually straight inside of the
outer vesicular zone, but becoming distinctly more attenuated
(as seen in transverse sections) and somewhat curved or a little
flexuous in crossing the vesicular area, where they alternate
with an equal number of very short, thin ones; tabule very
wide, or occupying about two-thirds the entire breadth, as seen
20 University Geological Survey of Kansas.
in longitudinal sections, and passing nearly horizontally across
with a more or less upward arching; dissepiments thin and
forming numerous obliquely ascending, small vesicles, in trans-
verse sections seen to pass across between the costz with an
upward curve. Entire length unknown.”’
In young specimens the cardinal septum and all the other
septa on that side of the corallum are very greatly developed
laterally after passing inward from the vesicular zone; the in-
ner wall of the vesicular zone is also thickened on that side of
the corallum. As a result of this great thickening of the septa
the interspaces are small, producing a peculiar appearance in
cross-section. The septal development becomes less and less
marked as the specimen advances in age, until in old specimens
it is hardly noticeable, save in the cardinal septum, though a
close comparison generally shows them a little larger on the
outside of the general curve. These peculiarities may be seen
by sectioning the large and small ends of any adult specimen.
The thickness of the dissepimental zone and also the number of
tabular dissepiments are very variable. The young specimens
are either rather slender or quite turbinate.
Range and distribution: Kansas City, Jefferson, Douglas
and Chautauqua counties.
Some specimens in the University Museum are about nine
inches long and apparently incomplete at both ends. Such
specimens are usually a little crooked through the entire length.
AXOPHYLLUM.
Milne-Edwards and Haime, Brit. Foss. Corals, pl. XXI, (1850).
Axophyllum rudis. Plate II, figs. 9-9c.
Axophyllum rudis White and St. John, Trans. Chic. Acad. Sci., 1, p.
115, (1867); Meek and Worthen, Geol. Surv. Ill., vi, p. 525, pl. xxx, ff.
6a, b, (1875): White, 13th Ann. Rep. St. Geol. Ind., pt. 2, p. 118, pl.
Xxtll, ff. 8, 9, (1883); Keyes, Geol. Surv. Mo., 1v, p. 107, pl. x11, ff. 5a, b,.
(1894).
Corallum simple, occasionally clustered by gemmation at the
base, turbinate, curved or geniculated, generally attached at.
the base or side. ‘The outer portion of the calyx is shallow and
the central portion is rather deep; pseudocolumella flattened,
moderately prominent, and approaching subquadrate when seen
eS ee ee ee eS ee ee
_——_—_—, 2
BEEDE. | Carboniferous Invertebrates. 21
in a section below the calyx. The septa are alternately equal,
the larger nearly or quite joining the pseudocolumella. Sur-
face marked by irregular concentric wrinkles ; spines or rootlets
are sometimes present; septal furrows moderately distinct.
Range and distribution: Upper Coal Measures ; Kansas City,
Lawrence, Osage, Cowley and Elk counties.
This coral tapers very rapidly from the calyx to the apex,
being about as broad as long.
MICHELINIA.
de Koninck, Descr. Anim. Foss. Belg., p. 29, (1842).
Michelinia eugenex. Plate II, figs. 12-12b.
Michelinia eugenee White, 13th Rep. St. Geol. Ind., 1883, pt. 11, p. 119;
pl. xxiir, ff. 14-16, (1884).
Original description: ‘‘Corallum usually in the form of a
small globular or irregularly ovoid mass, higher than broad,
with the corallites usually opening upon all sides, except its
very small base, which is often concave and irregular; corallites
small, but very irregular in size and shape; calyces moderately
deep, their walls rather thin and margins narrow or even sharp.
Height on one of the larger masses in the collection, 26 mm. ;
diameter of the same, 17 mm. Diameter of the larger calyces,
3 mm.; of the smaller ones, 1 mm.”’
Range and distribution: Upper Coal Measures; Pomeroy,
Wyandotte county, Kansas.
He also remarks that ‘‘usually the calcyces cover the whole
outer surface of the corallum except the small base, which was
evidently attached to some foreign body; but occasionally a
considerable surface above the base is free from calyces, and is
covered with a wrinkled epitheca.’’ It may be interesting to
note that we have younger masses of but few corallites, one of
which is attached to a crinoid stem.
22 University Geological Survey of Kansas.
TRACHYPORA.
Milne-Edwards and Haime, Pol. Foss. d. Terr. Pal., p. 305, (1851).
Trachypora austini. Plate V. fig. 8: plate VI, figs. 7-7b.
Trachypora austini Worthen, Geol. Surv. Ill., vii, p. 81, pl. x1, ff. 1-1d,
(1890).
Worthen’s description: ‘‘Corallum dendroid, the branches
generally cylindrical, sometimes irregular, from 15 to 25 mm.
in diameter, and infrequently divided. Corallites conical,
diverging from an imaginary axial line to open on all parts
of the free surface. The calyces all oval or circular, very
variable in size, the larger ones 1.1 mm. in diameter, and
either level with the general surface, or with an elevated mar-
gin. Each orifice is adorned with small tubercles or short
ridges arranged in a radiate manner around the calyx margin.
Openings of the corallites separated by dense calcareous tissue
of variable thickness, but apparently always as wide as, or
wider than, the diameter of the tube orifices. Sections show
that the tubes are prismatic and in contact with each other,
that their walls are greatly thickened by a calcareous deposit
on the inner side of the tubes, and that the amount of thick-
ening increases toward the orifices. Mural pores of large size
are present, but apparently not numerous. Tabule are best
developed in the axial region, where they cross the tube cavity
at intervals equaling once or twice the diameter of the tubes.
Tangential sections show that the corallite cavities are sur-
rounded by blunt, thick, septal ridges. The walls are now com-
posed of fibro-crystalline calcite, and the change has destroyed
the finer details of structure.’’
Range and distribution: ‘‘Coal Measures; Labette county,
Kansas.”’
‘‘In all important respects the species described here re-
semble the Trachypora ornata Rominger sp., from the Hamilton
group, and I do not think any reasonable objection can be urged
against placing them in the same genus. Specifically they are
quite distinct, the corallum of Rominger’s species being much
smaller, the corallites much less variable in size and not so
prominently margined, nor are the interspaces so thick.”’
BREDE. | Carboniferous Invertebrates. 23
AULOPORA.
Goldfuss, Petref. Germ., p. 82, (1826).
Aulopora? prosseri. Plate III, fig. 2; plate IV, fig. 2.
Aulopora ? prosseri Beede, Kans. Univ. Quart., vil, p. 18, pl. —, f. 2,
(1898).
Corallum large, prostrate, bifurcating, the calyces rising ver-
tically or obliquely from 3 to 7 mm. or more; average diameter
2mm. orless; average diameter of prostrate portion a trifle
less. Calyces not campanulate, cylindrical,- openings nearly
circular; corallites wrinkled ; weathered specimens show longi-
tudinal striz indicating rudimentary septa; the corallites are
one to three diameters distant. In the lower portion of the
corallite the cavity is nearly or quite closed on account of the
internal thickening of the wall. Tabule generally wanting,
occasionally present in moderate numbers and pass from wall
to wall, sagging regularly near the center. Size of corallum
unknown. Colonies twenty-two inches across were recently
collected in the southeastern part of Douglas county, and still
appear to be incomplete. The base of this species broadens rap-
idly, and in good specimens appears somewhat like coarse lace.
Range and distribution: Upper Coal Measures; Lyndon,
Osage county, Lecompton and near Twin Mounds, Douglas
county.
In large slabs this species looks not a little like Syringopora,
but differs from it very markedly in the absence or sparsity of
tabule and the fact that the tubes are always rather short.
Aulopora? anna. Plate III, fig. 3.
Aulopora? anna Beede, Kans. Univ. Quart., vi, p. 18, pl. —, f. 3, (1898).
Corallum prostrate, diffusely branched, branches interlacing
and anastomosing at every contact; walls thin, save at the base
of the corallite; tubes very short, slightly subconical, immedi-
ate openings slightly flaring, circular to oval; no tabule dis-
tinguishable ; septa occasionally represented by a faint ridge in
the best-preserved corallites ; diameter of calyx opening, 2mm.,
contiguous to one or two diameters distant. Corallite low, a
little larger at upper extremity than at the base.
Range and distribution: Upper Coal Measures; Morehead.
24 University Geological Survey of Kansas.
This species is profusely branched baso-laterally and anasto-
moses to such an extent as often to form solid mats of coral.
It differs from Aulopora in having no tabule, in which respect
it agrees with Cladochonus, but it is prostrate, and does not seem
to reproduce by lateral gemmation as does the latter; hence it
is referred, provisionally, to Aulopora. At first it was thought
to be the base of a colony of Syringopora, as it was attached to
the top of a colony of that species, but, on examining the bases
of Syringopora, they were found all to contain very great num-
bers of tabule, which clearly separates the genera.
It differs from A.? prosseri in the proximity of the corallites
and the degree to which it anastomoses, and the corallites are
generally shorter and the mouths more flaring.
CLADOCHONUS.
Cladochonus? bennetti. Plate III, fig. 1; plate V, fig. 7.
Cladochonus? bennetti Beede, Kans. Univ. Quart., v1, p. 17, pl. —, f. 1,
(1898).
Corallum loosely fasciculate ; corallites one or two or more di-
ameters distant; erect corallites larger than the basal branches,
often five times as high as thick, upper portion budding and
sending off branches as at the base; epitheca strongly wrinkled,
upper portion of the wall of the calyx thin, opening circular,
deep, funnel-shaped by the thickening of the wall interiorly, in
the lower portion of which, as a rule, there is only a small cap-
illary opening, though sometimes larger. Average diameter of
corallite, 2 mm.; length, 6 to 18 mm.
Range and distribution: Upper and Lower Coal Measures ;
Fort Scott, Howard.
This species resembles Aulopora? prosseri more closely than
any other Kansas species. It differs from this, so far as known,
in having no tabule, and in the manner of budding profusely
at the upper part of the corallites. It resembles Romingeria
umbellifera Rom., but the absence of tabule removes it from
this genus. It agrees with Cladochonus McCoy, save that they
are only funnel-shaped when young, and very slightly so then.
The corallites are long and very stout, resembling Syringopora
in general appearance.
BEEDE. | Carboniferous Invertebrates. 25
SYRINGOPORA.
Goldfuss, Petref. Germ., p. 75, (1826).
Syringopora multattenuata. Plate II, figs. 10, 10b; plate V, fig. 6.
Syringopora multattenuata McChesney, Desc. New Pal. Foss., p. 75,
(1860); ibid., pl. 11, ff. 4a, b, (1865); Trans. Chic. Acad. Sci., 1., p. 2,
pl. u, f. 4, (1867); Meek, Fin. Rep. U. S. Geol. Surv. Neb., p. 144, pl. 1,
ff. 5a-d, (1872): White, U. S. Geogr. Surv. West 100 Mer., rv, p. 100,
(1877): Keyes, Geol. Surv. Mo., rv, p. 122, pl. xiv, f. 6b, (1894).
Corallum forming large masses; corallites subcylindrical,
somewhat tortuous, spaced one to two diameters distant; epi-
theca wrinkled, thick; septa obsolete; tabule very numerous,
somewhat funnel-shaped, often forming a tube down the center
of the corallite, running into each other laterally, forming vesi-
cles. Size of corallum unknown, specimens eighteen inches
in diameter appearing incomplete on all sides. Connecting
tubules numerous and irregularly placed. The diameter of the
corallites varies from 5 by 13 to 2 by 2 mm.
Range and distribution : Upper Coal Measures ; Atchison, Le-
compton, Lyndon, Osage county.
CHATETES.
Troost, manuscript (1849?). Milne-Edwards and Haime,. Monog. des Polyp. Foss.,
p. 272, (1851).
Chetetes milleporaceus. Plate II, figs. 11-11b.
Cheetetes milleporaceus Milne-Edwards and Haime, Monog. des Polyp.
Foss., p. 272, (1851): Hist. Nat. des Corr., 111, p. 271, (1860); White,
Powell’s Geol. Uinta Mts., p. 88, (1876); U.S. Geol. Surv. West 100 Mer.,
Iv, p. 98, pl. v1, f. 2a, (1877); Keyes, Geol. Surv. Mo., rv, p. 123, pl. xiv,
ff. 12a, b, (1894).
Corallum large, massive, somewhat globose; corallites very
closely arranged, very long, same size throughout the entire
length; walls thin, roughly five- to seven-sided, one diameter
usually longer than the other; average diameter about a third
of a millimeter. Tabule about a third to a seventh of a milli-
meter distant, horizontal, not always uniform throughout the
group.
Range and distribution: Coal Measures, Girard ; very abun-
dant in the Oswego limestone, in which they form large masses.
Specimens of this species often attain a very large size. Some
from the Oswego limestone are thirty inches in diameter, and
still appear to be incomplete.
3—Vi
26 University Geological Survey of Kansas.
CRINOIDEA.
Crinoids are those animals known as sea-lilies or stone-
lilies. They live in the sea at varying depths, from shallow
water to 3000 fathoms. They were very abundant during geo-
logical time, and probably reached their maximum development
during the Carboniferous period. They are not well adapted
for preservation as fossils. Their skeletons are made up of
angular pieces fitted together, but which usually fall apart
when the animals decay, so that the preservation of an entire
individual is a comparatively rare occurrence. It is not uncom-
mon to find masses of stone almost entirely made up of these
pieces, since the animals were gregarious, living in colonies as
they do at present. During the Carboniferous and Triassic
periods they were much more abundant than now.
Most of the crinoids are attached to the mud by stems which
have rootlets at the base to anchor them. However, some of the
forms are free during most, or all, of their lives. ‘These stems
had small canals running through their centers. They were
either round or three- to five-sided. They are often found
weathered out in the forms of little discs, which are generally
called ‘‘ Indian beads.’’ Upon the upper end of this stem rests
the calyx or cup-shaped base of the animal. The calyx is
generally composed of three circlets of plates. first, the in-
frabasals, or the plates which rest on the top of the stem.
There are usually five of them, and they are quite small, and
occasionally are united into one, or they may be entirely want-
ing. Above these are the basals, or second ring of plates, which
are longer and generally six-sided. Upon these rest the third
row, or the radials, which support the arms and the top part
of the animal. There are usually five of the radials, which
are flat on top and five-sided. On the posterior side of the ani-
mal there is often a series of plates supporting the ventral tube,
above which the anal opening is situated. These plates often
take part in the formation of the cup, and are called the anal
plates.
BEEDE. | Carboniferous Invertebrates. 27
Resting on the radials are the arm plates. The arms are
five or ten in number. They are composed of short plates laid
one on top of another, and
capable of motion. The A232
inner side of the arms has IY
usually a groove, along
each side of which is a Pee
row of smaller armlets or
pinnules. These grooves
connect at the base of the
arms with what are called
the ambulacral areas,
which in turn connect
with. the mouth, where
food particles which are
carried in the water are
selected out.
The ventral side or top :
. . . Fic. 2. Diagrammatic view of a crinoid with the plates
of the animal is often pro- separated, showing their relative form and posi-
= tions: The black spot in the center is the hole in
duced into a ventral Sac, the base of the calyx that communicates with the
aie stem. Jb, infrabasals; b, basals; r, radials; br,
or anal tube, as 1t 1s some- costals; a, anal. The remainder are arm plates.
times called. The anal
opening is either on top or in one side of this sac. The mouth
is almost always centrally located. The crinoids are more closely
related to the starfish and the sea-urchins or sea-cookies than
to any other animals.
In the accompanying figures the various parts of the crinoid
are shown and designated.
SCAPHIOCRINUS.
Hall, Geol. Surv. Iowa, p. 550, (1858).
Scaphiocrinus? washburni. Plate VI, figs. 2, 2a.
Scaphiocrinus? washburni Beede, Kans. Univ. Quart.,1, p. 21, pl. v, ff. 2,
2a, (1899).
Calyx broadly obconical, somewhat wider than high, rather
stout, and smooth. Infrabasals rather large, bent upward at
outer end, exterior outline pentagonal, plainly visible in te
side view of the calyx, well defined. Column large, circular,
28 University Geological Survey of Kansas.
composed of alternately thick and thin pieces, the outer half
of the sutures radially milled, the median canal of moderate
size. Basals large, three hexagonal, and the posterior and right
posterior ones heptagonal for the reception of the anals; the
plates are nearly all equal, and the sutures are placed in a
shallow, rounded furrow, which in branching at the angles of
the plates gives them a rounded appearance, so that the plates
appear subsemicircular in outline. The radials are the largest
plates of the calyx, all pentagonal, all about equal, save the
right posterior, which is smaller than the rest; the entire sur-
face truncated above, edge slightly beveled, wider than high,
entire upper surface faceted ; the transverse ridge is milled, the
lateral corners of the plates drawn in, apparently leaving a pore
which connected with the body cavity of the calyx; there is a
dorsal canal piercing the upper surface of the plates; the inner
edges of the plates are drawn in at the center and extend in the
form of a ridge to the canal. Costals 1x5, about half the size
of the radials, pentagonal, line of ‘articulation with the radial
gaping, lateral edges constricted and apparently not in con-
tact, about twice as wide as high. The first interradial is large
and situated well down in the cup, supported by the posterior
basal and the one at its right, on the sides by the special anal
and the right posterior radial, and supports another anal above
it on its truncated top. The special anal is moderately large
and hexagonal, bounded on the right by the two interradials,
below by the truncated upper surface of the posterior basal, on
the left by the left posterior radial and the space between it and
the costal; it is about half within the calyx. Above the first
interradial is a second, somewhat smaller, pentagonal plate
which is about, or a little less than, half within the calyx.
This plate, together with the special anal, supports the pos-
terior portion of the ventral sac (apparently two columns of
plates), which seems to be composed of rather stout five- to
seven-sided plates with no or very small pores at the angles ;
they have a botryoidal surface. At one side and on the top of
the sac there seems to be a madreporite plate pierced by large-
sized pores. The sac was apparently about the size of the calyx,
a ee ee eee. ea Te a = sa
BREDE. | Carboniferous Invertebrates. 29
or perhaps the calyx and costals together. Arm plates rounded
on the exterior, not at all or very slightly wedge-shaped at
the base, and moderately stout. Pinnule present and of mod-
erately large size.
Height. Width.
De EES a SURRY ptt a a 8 mm.
JOE SUG, Deals sh Se ee ae 3mm 5 fs
MISE etn im ae ly enn 6 uw oe ig hE aba eo eae i
RIUM Ce ec uec as bv cee lk sche Oy & 14 $5 i acre
i MR TUROCUAD ak) leek wees Sire QA * atari
SOS re) nr By ee 6 §§
PeMMPONERT TSUN edt a LG kde g cic ala Swe 2 a SF te
ENE Sil eck thd k ia s G. -* je
MUI EMM CD uhle Cole ka aKOK ain wis vee es 5 A a yi oe
Range and distribution: Upper Coal Measures; Topeka,
Kan., from the horizon of the Osage coal. Now in the collec-
tion of Washburn College, in honor of which it is named.
This species seems to belong to the Poteriocrinoidea, though
there is some difficulty in locating it generically, as it seems to
combine some of the characters of several genera. It agrees
with Homocrinus in having a round dorsal canal piercing the
first radials, but differs from it in that it has pinnules, a robust
calyx, and the entire top of the radials truncated. According
to the definition of Poteriocrinus, the presence of the round dor-
sal canal in the radials removes it from that genus, as would
also the fact that the facets of the radials face upward rather
than outward. It differs from Scaphiocrinus in having a circu-
lar column, and the fact that the transverse ridge does not oc-
cupy nearly the whole of the upper surface of the radials and
the brachials are not long. However, it agrees in other respects
with this genus better than any other, and it is provisionally
referred to it.
ZEACRINUS.
Troost, Cat. Foss., (1850).
Hall, Geol. Surv. Iowa, p. 541, (1858).
Zeacrinus? robustus. Plate VI, figs. 1, la.
Zeacrinus? 1 obustus Beede, Kans. Univ. Quart., 1, p. 21, pl. v, ff. 1, la.
Calyx shallow, saucer-shaped or nearly flat, unsymmetrical,
five or six times as broad as high, deeply concave at the base ;
plates tumid, and the sutures are in depression ; surface finely
granular. Infrabasals five, equal, half concealed by the col-
30 University Geological Survey of Kansas.
umn, deeply concave and superior to the basals, forming a
large elevation in the interior of the cup, nearly one-fourth its
entire width and fully one-half its height; column round, com-
posed of thin plates, which are carinated, and the sutures are
crenulated ; the canal is round and small. Basals five, large,
tumid, three hexagonal and two heptagonal, unequal, situated
below the infrabasals and forming a large part of the real body
cavity of the cup. The three anterior basals are equal, about
as broad as high, very convex, the posterior one compressed
laterally, superior lateral edges longer than the others; trun-
cated side for the support of interradial short, side next the
radianal long; the right posterior basal heptagonal very broad,
left posterior side very long, side adjacent for the reception of
the radianal very short, the latter being situated between the
posterior basal and the right posterior radial. Radials five,
large, very moderately ascending, placed superior to the basals,
five- to seven-sided, quite massive, more than twice as large as
the basals, convex, twice as wide as high, upper exterior por-
tion much beveled and concave; plates separated at the corners
and often along the line of union; at the upper union there
seems to be an opening that communicates with the calyx cav-
ity between the arms; right and left posterior radials forced
apart fully one-half the diameter of either by the interradial
and radianal; the upper surface is faceted the entire width of
the plate, two external ridges present and milled ; the remainder
of the surface is nearly flat or a little concave; the plane of
these surfaces is not horizontal but the inner side of each is
higher than the outside, so that if each were produced inward
they would form rather an obtuse cone; the inner notch occu-
pies about a third of the upper surface of the plate. NRadianal
long, coffin-shaped, curving upward above, about twice as long
as wide, pentagonal, side adjacent to the right radial much the
longer; the superior side supports an anal which is hexagonal,
very thick, six-faceted above, touching right radial for a short
distance, one and one-half times as high as wide, widest above,
mostly without the calyx. Anal large, heptagonal, very thick,
BEEDE. | Carboniferous Invertebrates. ol
one-third within the calyx, broadest above; height one and
one-half times the width.
A portion of the vault of this specimen remains, somewhat
crushed down into the calyx. It appears to have been in the
form of a pyramid, about as broad as high, composed of rather
heavy plates, which were rather rough. It appears to have
been rather angular, though it may have been conical. Several
rather irregular plates are preserved, three of which seem to be
from around the aperture; they are rather massive, five- or
six-sided, nearly smooth on the exterior; articular surfaces are
deeply faceted and roughened, while the articular surfaces of
the other plates of the sac are milled. |
Height. Width.
2 USSG 2 So 1 to 13 mm. 4} mm.
RTS eet et eu gig a 4d we ee we 2 rates on
BEHINOG SIGOFIOE DHSBIS ...... es ee eee ee iy dees een
ODS SPSS SU tig ts ey Coo) a a ne 9, 55 1g
NTI De Ot Sr ane De eo eae a Phin winnie amide Ta
REPENS flere ce NR oe soe pe Se 14S PP
EDS LW oh oS Se ee res ee 83 § y Se
i EE i er Ge ae a err ese Ts Ba.
NRE SIE id can WSF LT Cd whe s dia Sg Slave aoa es Se 4% ss
NE fer a ESS oe os ate odin bloikv wh aes nae. AB? tn,
Range and distribution: The specimen is labeled, ‘‘ From the
Upper Coal Measures; Kansas City.’’ University of Kansas
Museum.
It is impossible to locate the specimen generically with any
degree of certainty without having more of the specimen pre-
served. The ventral sac seems to have been angular and the
stemround. For this reason it is left with Zeacrinus. The up-
per extremities of the radials do not meet, but leave a small
aperture, which seems to communicate in life with the general
body cavity. I know of no Coal Measures crinoid with which
it is likely to be confounded.
32 University Geological Survey of Kansas.
CERIOCRINUS.
White, U. S. Geol. and Geogr. Surv. Wyo. and Idaho for 1878, p. 128, ( 1883).
Ceriocrinus craigi. Plate VI, figs. 9, 9b.
Eupachycrinus craigit Worthen, Geol. Surv. IIl., vi, p: O27, pl. xoxo
Feb clsia)):
Ceriocrinus craigii Wachsmuth and Springer, Proc. Acad. Nat. Sci.
Phil., p. 178, (1886); Rev. Paleocrin., 111, p. 254.
Calyx basin-shaped, smooth, concave at base, about three
times as broad as high, roundly pentagonal in outline when
viewed from below. Infrabasals hidden by the column, situ-
ated within the calyx. Basals probably pentagonal or hexa-
gonal, if they have two short sides at base, except the dorsal-
one, which is truncated for the support of the interradial ; height
about equal to the width, strongly incurved at the base to meet
the infrabasals. Radials pentagonal, nearlv equal, convex,
twice as wide as high, upper edge beveled, with a slight depres-
sion in the center, upper side faceted over the entire top of the
plate, ridges prominent, crenulated, the large one not notched
interiorly, furrows distinct, inner portion of the plate depressed.
Interradial small, hexagonal, compressed laterally, about half
within the calyx, upper portion curving strongly inward. Cos-
tals five, short, obtusely spinous, extending half their length
beyond the calyx; lower interior surface faceted to articulate
with the radials, upper surface faceted to the tips of the spines
for the reception of the two distichals, each of which supports
an arm. ‘There is a strong keel on the upper side of the plate
extending to its inner edge; it also gives rise to two lateral
ridges which diverge from the tip of the upper surface to the
corners of the plate; the ridges are crenulated ; between these
ridges are long depressions. The surface of the calyx is smooth.
Height. Width. Length of spine.
Basa lau iy. i. STEN eINN sek, ee 7; mm 7% mm
adials iy. bea eS Sen eee ae dep See
interradial yy. 20.52 34, een ieee 4h 6 Oe ase =
(Osta: Pe POM Ue) kel aa Gauss LOM uCE 9mm.
(Cai ele is nl Eee Sn Om ae 21 =‘ (slightly compressed ).
Range and distribution: Upper Coal Measures; Kansas City.
According to Worthen, C. craigi differs from C. fayettensis in
a es —_ —_ Ts. - —
—
BeEDE. | Carboniferous Invertebrates. 33
its ‘‘ larger size, more robust form, and less convex plates.’’ In
these respects, judging from the figures, our specimens agree
more closely with (C. craigi.
Ceriocrinus? monticulatus. Plate VII, fig. 2.
oy dei de monticulatus Beede, Kans. Univ. Quart., vi1I, p. 123, pl.
XXxII, f. 2, (1899):
Calyx basin-shaped, moderately deep, slightly concave at base ;
arms moderately stout, their number not known; pinnules mod-
erately long. Infrabasals not known, but probably small and
nearly covered by the stem. Basals large, hexagonal, ‘convex,
curved regularly inward at the base, about as high as wide,
upper lateral edges sometimes a little unequal on account of
the unequal radials, which makes some of the upper and lower
lateral edges unequal also. Radials a little unequal in size,
pentagonal, the upper edges beveled, about twice as wide as
high, upper surface faceted, the line of articulation with the
costal gaping. Costals five, pentagonal in outline, somewhat
produced exteriorly but not spinous, twice as wide as high, with
single facet below to meet the radial and two above for the ar-
ticulation with the distichals. Distichais 2x5, quadrangular
or pentagonal according to the number of arms supported, ap-
parently faceted above and below; those supporting two arms
very similar in shape to the costals. There are often one or two
palmars before the postpalmars are reached, at the base of the
arms. Arms long, rather stout, somewhere from fourteen to
eighteen in number, ten in view above three radials; they are
made up of two series of short, stout, interlocking, cuneiform
plates, each bearing a single, long pinnule. Pinnules com-
posed of at least eight long, slender joints, slightly grooved on
the inner side ; along each side of the groove is a row of closely
set nodes, or little plates. ‘The entire surface of the cup and
arms is covered with monticules and fine granulations.
Height. Width.
Se a nO hn 2 ty be eS 10 mm. 10 mm.
RRM re tao NS bond ao d Siepinhi ite ame 6 11 *§
SURMAMREE TE ere Re Pere ge ae be RA : ob gla
IE hee 5 ie Crs lal y' Svie dive cede a eee a 1 lh
ia Re ee ee eT el 53 he ue a
RMEREERS AUNTS Dio Crates abies Siof nc, 21k & tie: are om COM ahs th el - sé
Ea Oe kee eck yy he a ok vate omer. 1s rT
34 University Geological Survey of Kansas.
Range and distribution: Upper Coal Measures; Topeka,
horizon of the Osage coal.
On account of the general resemblance of this species to some
of the species of Cervocrinus, it is left there for the present, un-
til the azygous plates and infrabasals can be determined.
Ceriocrinus hemisphericus. Plate VI, figs. 5, 5b.
Poteriocrinus hemisphericus Shumard, Trans. St. L. Acad. Sei., 1, p. 221,
( 1858.)
Scaphiocrinus? hemisphericus Meek, Fin. Rep. U.S. Geol. Surv. Neb.,
p. 147, pl. v, ff. la, b, pl. vit, ff. la-c, (1872); Meek and Worthen, Geol.
Surve Ll vive py o6le plaexcxcan, aio, St ane
Ceriocrinus hemisphericus Keyes, Geol. Surv. Mo., tv, p. 220, pl. xxvim,
ff. 2), 5, (1894).
Meek’s description: ‘‘ Body below the summit of the radials
subhemispherical, with the under -side deeply concave. Base
very small, pushed or inverted entirely within the cavity of the
body, and nearly or quite hidden externally; column facet
round and deeply sunken. Radials of moderate size, very
nearly equal, having a general pentagonal form, excepting on
the anal side which is a little truncated above by the anal
piece so as to give it a general hexagonal outline; each, how-
ever, has an additional very obscure mesial angle at its connec-
tion with the base, and all are strongly incurved below to form
the concavity of the under side. Radials nearly twice as large
as the basals, twice as wide as high, and equally pentagonal,
the upper side being longer than any of the others and all
evenly truncated. Costals (at least the two on the anal side)
comparatively narrow, but still wider at the base than high,
rounded on the outer side, a little constricted in the middle, and
pentagonal in form, the two upper sloping sides supporting the
first divisions of the arms, which are composed (at least for the
first three ranges) each of a single series of wedge-formed pieces.
First anal piece comparatively small, a little concave, resting
upon one of the basals and connecting with the radials on each
side, above which it projects, supporting upon its slightly in-
curved upper edge a second piece, the form of which is unknown.
Surface smooth, or only with traces of minute granules.”’
Range and distribution: Upper Coal Measures; Jefferson
county. Now in the collection of Washburn College.
BEEDE. | Carboniferous Invertebrates. 35
Ceriocrinus missouriensis. Plate VI, fig. 6.
Delocrinus missouriensis Miller and Gurley, Jour. Cine. Soc. Nat. Hist.,
xu, p. 14, pl. u, ff. 11-13, (1890); 16th Rep. St. Geol. Ind., p. 336, pl.
11, ff. 11-13, (1890).
Delocrinus hemisphericus Miller and Gurley, ibid., p. 335, pl. 1, ff. 8-10,
pl. x, f.5; Jour. Cinc. Soc. Nat. Hist., x11, p. 12, pl. 11, ff. 8-10. These
species referred to the proper genus by Weller, Bull. U. S. Geol. Surv.,
153, p. 172, (1898).
Calyx basin-shaped, shallow, pentagonal in outline viewed
from below, nearly four times as broad as high, base quite con-
cave. Infrabasals small, not entirely hidden on the exterior.
Basals five, moderately large, pentagonal (the angle at the base,
if it exists, is so small that it is almost impossible to discover it)
except the posterior one, which is truncated for the reception of
the interradial, giving it a hexagonal outline, about as high as
wide ; supralateral edges about equal, basal edge very short ;
plates not very sharply convex in the center. Radials five,
nearly equal, about twice as large as the basals, and about
twice as wide as high, upper edge beveled and upper side
faceted, ridges of facet crenulated. Interradial small, laterally
compressed, the upper portion bent strongly inward. Costals
five, roughly triangular in outline, stoutly spinous, faceted be-
low for the articulation with the radials, entire surface of the
inner portion faceted above for the support of the two arms; the
spines are stout and blunt. The arms are composed of two
series of cuneiform interlocking plates which are rather convex
on the exterior on the base of the arms. ‘The surface is smooth.
Height. Width. Length of spine,
CS ee il ee tl ee Pas Se 8 mm 6 mm
ET AUTO a Sie SS eae aa nO a a tS fos
MSM er CASE tors oS venta pds wide wha Bast id We. 14 mm.
SRE EORTNER Von ath a LSS lis heres coe ee + ls
Calyx (another individual)............ ie ae ———.
Range and distribution: Upper Coal Measures; Kansas City,
Argentine.
It is impossible to distinguish this species from C. craigi by
the calyx alone (if it is distinct from that species) unless the
base is so preserved as to show whether or not the infrabasals
are concealed, unless the costals be present. ‘The only differ-
ence between the two species is that in C. missouriensis the
36 University Geological Survey of Kansas.
spines are much longer, the infrabasals not hidden by the col-
umn, and that the cup is perhaps a little shallower, as is also
the concavity of the base. |
If Meek’s determination of C. hemzsphericus is correct, it does
not have the costals developed into spines at all, and hence it
is distinct from the spined forms. The minute differences be-
tween the two spined forms, as figured by Miller and Gurley, are
of little value.
PHIALOCRINUS.
Trautschold, Mon. Kalkbrueche von Mjatschkowa, p. 122, (1879).
Phialocrinus magnificus. Plate VI, fig. 10.
Esiocrinus magnificus Miller and Gurley, Jour. Cine. Soc. Nat. Hist.,
.Gatig 05 IU, pole sue, 36 Ibs,
Phialocrinus magnificus Carpenter, Ann. and Mag. Nat. Hist., p. 96,
(1891); Keyes, Geol. Surv. Mo., tv, p. 220, pl. xxviii, f. 4, (1894).
Calyx bowl-shaped, base truncated, of medium size, surface
finely granulated. Infrabasals five, cuneiform, pentagonal,
half covered by the column, stem facet radiately furrowed.
Basals five, hexagonal, posterior one heptagonal ; the two sides
at the base rather indistinct, edges straight; plates convex,
curving inward at the base to meet the infrabasals, very
slightly wider than high. Radials convex, nearly equal, the
right and left posterior ones occasionally a trifle smaller than
the rest, pentagonal, width about one and one-half times the
height, upper outer edges beveled. Interradial resting on the
truncated upper surface of the posterior basal, large, pentago-
nal, within the calyx, wider above, greatest width equai to
the height, supporting two anals. First brachials large, quad-
rangular, the lower outer edge beveled. The second brachials
pentagonal, highest in the center, superior side faceted for the
support of the two arms, about twice as broad as high
Arms ten, slender, often 100 mm. long, pinnulate, pinnule
often 10 mm. long. The arms are composed of a single series
of slightly cuneate plates, the inner side of which is fur-
nished with a deep, broad groove. Above the interradial rest
at least two anal plates which curve moderately inward. The
proboscis is long, rather slender, subquadrate in transverse out-
line; in adult specimens, between the lobes of the tube, are
.- -y=?
- aS a ee
BReEDE. | Carboniferous Invertebrates. 37
deep reentrant angles, giving it the appearance of four vertical
tubes closely placed. In young specimens it is more round and
less rough than in the adult. The plates are so closely united
that it is difficult to make out their exact outline, but they are
very convex on the exterior. The upper plates in young speci-
mens are almost smooth, becoming more highly ornamented as
the base is approached, while on old specimens the plates are
highly tuberculated and the sides are ornamented by raised
ridges which seem to surround apertures in the plates.
Height. Width.
UES USS 9 (SS PM ee hielo athe: « oe 33 mm. 3mm.
CR En SRE ae ee an Ay oie
MME S Er EAS eh Ce Ses Sa i ats Beebe oe ak 5A 8 es
MER MRRIRMERIN ee ee. nt ee in gs se bee ene erie ce
be ER SS coe oS 2s & oS
Urea BETES ESRRE ee AG eet oo 20S ecg sca, LN bln a ols as ee Bk: Ta:
SIME RAE DF IANS Sark ik Bo a Sinmia, Rio bm La wis sees meu
EMODOSCIS ....... PERRET ERAS Se rie is Shed tng es sw 8; .-“ ch
Range and distribution: Upper Coal Measures; Kansas City
and Argentine.
ERISOCRINUS.
Meek and Worthen, Am, Jour. Sci., LXXX, p. 174, (1865).
Erisocrinus megalobrachius. Plate VII, figs. la, 1b.
FPrisocrinus megalobrachius Beede, Kans. Univ. Quart., vir, p. 124, pl.
xxxii, ff. 14,.b
Calyx basin-shaped, quite concave, ornamented by very
coarse granulations, which are sometimes arranged in wavy
rows. Infrabasals unknown, covered, or nearly so, by the
small column. Basals large, convex, the lower portion curved
upward to meet the infrabasals forming a deep cavity in the
base of the cup, the inner (or lower) end of the basals being in
about the same horizontal plane as the upper end, and hence
forming most of the base of the calyx and leaving the infra-
basals entirely within the body cavity; higher than wide, all
about equal, and apparently pentagonal, the lower side (or
sides, if two) short, superior and inferior lateral edges nearly
equal, the apical angle extending upward between the radials
to fully one-half the height of the latter. Radials five, equal,
pentagonal, twice as wide as high, massive, convex, consider-
ably beveled at the upper edge; upper surface faceted for the
38 University Geological Survey of Kansas.
reception of the costals, the line of articulation with which is
gaping, raised portions of the facet milled. Costals five, mas-
sive, stoutly spine-like, pentagonal in outline, lower inner sur-
face faceted to meet the radials, upper surface faceted, except
the portion protruding in the form of a stout spine to support
the large arms. Distichals either one or two to each arm, more
commonly two in our specimen, the lowermost quadrangular in
outline, massive, a little more than twice as wide as high; the
second distichals are very variable, from four to six times as
broad as high. There are ten arms which are very broad and
stout, each made up of two series of wedge-shaped, interlock- —
ing plates which are twice as long as high in the lower portion
of the arms and each supporting a single pinnule. The pin-
nul are not well preserved, but are stout, composed of rather
large, square plates near their junction with the arms, while
farther away they assume a cylindrical form. Where not worn
the entire specimen is covered with coarse granulations which
are usually a little prolonged.
Height. Width. Length.
Basalsice Bega sr Mey ong) Mauer 11 mm. 10 mm. =
Ravana eae Skier mae eit, sar eae er Siac Gy 8s
Costas ca eee tS lciee Meet coke Me ON np ateolenas 15S 8 9 mm
He Distichialg.. were ne. fants UNE Cs Ae ae 1-2 ‘* Sake
Pa, | DSI RAK) alee tl pa atipe he aay SN I a Dies Anis a
Average lower arm plate...... ..... Dies ene Ss a
Calliyex ir ites Sen Os er rarer LOr Pah, O ———
Range and distribution: Upper Coal Measures ; Topeka, from
the horizon of the Osage coal.
This species agrees in many respects with Cerzocrinus craigz
(Worthen) W. and S. and C. hemisphericus (Shumard ) W. and
S., but each of these possesses an anal plate; the specimen in
hand, though preserved in very perfect condition, shows no in-
dication of such a plate. It is also very much like Hrisocrinus
typus as figured and described by White (Contributions to In-
vertebrate Palaeontology No. 6, p. 126, p. 33, f. 5), but the
arms of our species are flatter, plates much less convex later-
ally, and granulated on the surface instead of smooth. The
brachials also are more spine-like in our specimen, and the
basals are not nearly so sharply convex as in that species, and
4
BEEDE. | Carboniferous Invertebrates. 39
the cavity in the base is deeper and the calyx is not so high.
It may belong to Stemmatocrinus, as the infrabasals are not
known, but as that genus is not yet known from this country,
it is more probable that it is an Frisocrinus.
Erisocrinus typus. Plate VI, figs. 4—4b.
FPrisocrinus typus Meek and Worthen, Amer. Jour. Sci., xx1x, p. 174,
(1865); Geol. Surv. [ll., 1, p. 317, f. 33; Meek, Fin. Rep. U.S. Geol.
Surv. Neb., p. 146, pl. 1, ff. 3, a, b, (1872); Meek and Worthen, Geol.
Surv. Lll., v, p. 561, pl. xxiv, f. 6. (1873): White, Geol. Uinta Mountains,
(Powell’s Rep.), p. 's9, 1876): Cont. Inver. Pal., No. 6, p. 126, pl. xxxr1r,
f. 5a, (1880).
Frisocrinus nebrascensis Meek and Worthen, Amer. Jour. Sci., Xxx1x,
p. 174, (1865).
+ ee pelvis Meek and Worthen, Amer. Jour. Sci., xxxrx, p. 350,
Meek’s description: “‘ Body below the summit of the first
radials [radials], basin-shaped, rounded below, and obscurely
subpentagonal in outline as seen from above or below; com-
posed of thick, smooth, slightly convex plates. Basal pieces
[infrabasals] small, occupying a shallow convexity of the under
side, about half hidden by the column, all pentagonal in exter-
nal form. Subradial |basal] considerably larger than the basal
[infrabasal], and all equally hexagonal in form. First radial
[radial] pieces four or five times as large as the subradials
[basals], wider than long, equal, and all pentagonal; support-
ing upon their broadly and evenly truncated superior sides the
second primary radials [costals], which are of nearly the same
size and form as the first, but have their sloping sides above
instead of below, while they each support above two first bra-
chials [distichals], or a series of secondary radials [distichals]|
yet unknown. Surface smooth. Breadth of body below the
first primary radials, 0.72 inch; height of same, 0.35 inch.’’
Range and distribution: The specimen here under discussion
is from the Upper Coal Measures, Jefferson county, and now in
the collection of Washburn College.
The specimen is a beautifully preserved calyx and a single
costal. It differs very materially from the type specimen
figured by Meek and Worthen. The calyx is much higher,
which is due largely to the very much larger basals, which, in-
40 University Geological Survey of Kansas.
stead of being one-fourth or one-fifth the size of the radials, are
nearly as large. The costals show a marked spinous develop-
ment on the upper side, but in one of the figures (Geol. Surv.
ll., V, pl. xxiv, fig. 6) of the Illinois forms the costal is shown
as being somewhat extended above. It is quite different from
the figure of the species by White above referred to, in being
quite plain and in having the plates less convex. It is also
more sharply pentagonal in outline than most of the members
of the species. However, this is a somewhat variable species,
and our specimen is probably a true HL. typus, but presenting
interesting variant features.
EUPACHYCRINUS.
Meek and Worthen, Proc. Acad. Nat. Sci. Phil., p. 169, (1865).
Eupachycrinus magister. Plate VI, figs. 3-3b.
Hupachycrinus magister Miller and Gurley, Jour. Cine. Soc. Nat. Hist.,
x1, p. 4, pl. 1, ff. 1, 2; d6th Rep: St. Geol: Ind-; p. 3283 plar. tease 9,
TOs ity Gtele (1890) ; Keyes, Geol. Surv. Mo., rv, p. 218, pl. XXVIT, ff.
Original description: ‘‘ This species is very large; calyx low
and broad, somewhat saucer-shaped, bulged a little on the
azygous side, height about half the width, sutures deep, ex-
cavation extending about half the thickness of the plates, plates
very strongly tuberculated, tubercles conical, elongated, and ir-
regular in form and distribution. The five basals [infrabasals|
are sunk in a cavity on the other side, projecting only half their
length beyond the column; even this projection is tubercular ;
they form in the interior of the calyx a pyramid, which is
pierced at the summit by a five-rayed opening connecting with
the canal in the center of the column; the points of the rays
are rounded. The basal plates are made pentagonal by the
truncation made at the points of the rays for the central canal.
The diagrammatic views which have been made of the basal
plates in this genus are incorrect, in so far as they indicate a
pentagonal opening with the angles directed toward the sutures,
instead of truncating the plates with the concave depression for
the five-rayed opening to the columnar canal. The two basals
on the azygous side of the species before us are larger than the
others, being nearly as large as the other three.
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BEEDE.| . Carboniferous Invertebrates. 41
‘‘The subradials | basals] are very large, extending into the
basal cavity, and curve gently upward; three hexagonal,
the two longer sides unite with the subradials; the two up-
per sloping sides, uniting with the first radials, are a little
shorter, and the two under sides, uniting with the basals
|infrabasals] are very short; two are heptagonal, the one on
the right of the first azygous plate being much larger, and, ex-
cept the two short sides uniting with the basals, the other sides
are of subequal length; the one upon the left has, in addition
to the two short sides uniting with the basals, a short side unit-
ing with the second azygous plate. Four of the first radials are
pentagonal, twice as wide as high; the upper face is the full
width of the plates, and projects over the interior of the calyx so
as to give the appearance of having great thickness when viewed
from above. The other first radial, upon the right of the azygous
plates, is quadrilateral, except a very slight truncation by the
second azygous plate below the depth of the suture. The first
radial is separated from the second or brachial piece, on the
outer face, by a wide suture, but within a crenated ridge ex-
tends from one angle of the plates to another, forming a penta-
gon, except as separated by the second azygous plate; the ridge
has a furrow upon the outer side in the central part of each
plate, and within this is a wide expansion which supports the
brachial arm pieces. The first azygous plate has four sides,
rests between the upper sloping sides of the two subradials and
along the under side of the first radial on the right, with the
shorter side abutting on the second azygous plate. The second
azygous plate is hexagonal, curves inward, and supports upon
its two short inner faces the third and fourth azygous plates,
side by side. The vault and other parts unknown.’’ In our
specimen the right posterior radial seems to be covered by one
of the plates of the ventral sac.
Range and distribution: Upper Coal Measures; Kansas City.
4—vi
42 University Geological Survey of Kansas.
HYDREIONOCRINUS.
de Koninck, Bull. Acad. Royale Belgique, VII, pt. 11, p. 18, (1858.)
Hydreionocrinus kansasensis. Plate VII, figs. 4-7.
Hydreionocrinus kansasensis Weller, Trans. N. Y. Acad. Sci., xvi, p.
372, (1898. ) |
Weller’s description: ‘‘ Dorsal cup depressed, nearly saucer-
shaped, more than twice as wide as high, truncated below.
The margins of all the plates, except the lateral margins of the
infrabasals, ornamented by narrow papillose bands parallel to
the sutures. Exclusive of the papillose bands the plates are
smooth. Infrabasals five, large, extending far beyond the
column, anchylosed into an irregularly hexagonal, nearly flat
disc, the sutures marked by shghtly elevated ridges; central
portion of the disc slightly excavated for the attachment of the
column, the excavation surrounded by a low ridge. Basals
five, four of them broader than high, spherical-triangular in
outline, in lateral contact only at the extreme lateral angles.
The posterior basal higher than wide, quadrangular in outline,
truncated distally for the reception of the special anal plate,
the lateral and proximal sides curved as in the four other
plates. adials large, twice as wide as high, the three anterior
ones heptagonal in outline, the two posterior ones hexagonal.
The proximal sides concavely curved to conform to the curved
sides of the basals, the proximal angles meeting the distal angles
of the infrabasals and the lateral angles of the basals. First
brachials twice as wide as high, pentagonal in outline, bearing
upon the two upper sloping sides of the two main divisions of
the arms. Arms uniserial, the component plates broader than
long, except near the tips, rectangular in outline except the
axials, which are pentagonal. In the right posterior arm, the
only one known, each main division bifurcates several times,
the most posterior one showing ten divisions at the tip. Radi-
anal nearly as large as the basals, pentagonal, higher than
wide, lying upon the truncated right posterior infrabasal and
between the posterior basal and the right posterior radial.
Special anal larger than the radianal, hexagonal, higher than
wide, lying upon the truncated posterior basal and between the
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BREDE. | Carboniferous Invertebrates. 43
radianal and the left posterior radial. First tube plate partially
included in the calyx, higher than wide, lying upon one of the
upper sides of the radianal and between the special anal and
the right posterior radial. Ventral sac very large, expanding
above and surrounded at its summit by arow of large, spatulate,
spinous plates which form a crown of diverging spines around
the summit of the sac.”’
Range and distribution: ‘‘From the Upper Coal Measures,
about 700 feet below the horizon of the Cottonwood limestone,
at Neal, Greenwood county, Kansas.”’
‘‘Remarks: The form of the plates in the dorsal cup of the
species differs materially from the other species of the genus,
but the ventral sac, which is the most remarkable feature of the
genus, and which is said by Wachsmuth and Springer’ to be
the best character for distinction, in all respects like that in the
remaining species. This organ is much crushed and only
partially preserved in the type specimen, but enough is present
to show its great expansion toward the summit and the crown
of large, spinous plates. Eight of these spatulate spinous
plates are recognizable in the specimen, all of them belonging
to one side, so that there must have been at least fifteen or
more altogether.
‘¢The dorsal cup is remarkable for the large size of the in-
frabasals which extend far beyond the column and do not rest
in a deep cavity, but are consolidated into an irregularly hex-
agonal flat disc. The spherical-triangular form of the basals
is different from any of the other species of the genus, and the
manner of meeting in one point of the distal angles of the in-
frabasals, the proximal angles of the radials and the lateral
angles of the basals is quite remarkable.”’
Hydreionocrinus subsinuatus. [late VII, fig. 14.
Hydreionocrinus subsinuatus Miller and Gurley, Bull. 3, Ill. State Mu-
seum Nat. Hist., pp. 40, 41, pl. v1, ff. 11-14, (1893).
Original description: ‘‘Calyx depressed, saucer-shaped,
slightly concave below, longitudinally concave on the ventral
side; sutures distinct; surface smooth. When viewed from
2. Revision of the Paleocrinoidea, I, p. 130.
44 University Geological Survey of Kansas.
above or below, hexagonal, in outline, by reason of the trun-
cated first radials, and the concave, wide azygous area. Column
round.
‘‘Basals form a pentagon one-half wider than the column,
with a central, columnar cavity surrounded by an external rim,
for the support of the attaching column. Subradials of moder-
ate size, three of them apparently pentagonal, though as each
one abuts upon two basals, where there is an obscure angle,
they are really hexagonal. The two adjoining the azygous area
are heptagonal. They curve very slightly down to the basals,
and upward, toward the acute angles between the first radials.
They are not uniform in size or shape; the heptagonal plate on
the right of the azygous area is the wider, and the one on the
left the longer one. First radials twice as wide as high, trun-
cated the entire width above, much thickened within, and sep-
arated from the second radials, on the outer face, by a gaping
suture, but immediately within a straight ridge extends from
one outer angle to the other, having a furrow on each side so
as to form a hinge on which the second plates articulate; be-
hind this hinge, in the middle part of each plate, there is a
socket for the reception of a tooth-like projection. Second ra-
dials short and heavy and produced externally in a moderately
strong spine.
‘‘Wirst azygous plate longer than wide, pentagonal, upper side
short, abutting below on a subradial and resting very slightly
oblique between another subradial and the under sloping side
of the first radial on the right. Second azygous plate longer
than wide, truncates a subradial, abuts upon the superior lat-
eral side of the first radial on the left, and the first and third
azygous plates on the right. Third azygous plate longer than
wide, truncates the first azygous plate, and abuts upon the su-
perior lateral side of the first radial on the right. The second
and third azygous plates are hexagonal, but the plates abutting
the superior side are not preserved in our specimens. A small
part of a proboscis is preserved in one of our specimens, that
shows some heavy, transversely furrowed plates. Other parts
not preserved.
— SS a oe ss
BEEDE. | Carboniferous Invertebrates. 45
‘‘This species is so different from any other referred to this
genus, that no comparison will serve further to distinguish it.
There is a possibility that it is a Zeacrinus, but we think it is
not.”’
Range and distribution: Upper Coal Measures; Cameron’s
Bluff, near Lawrence.
AGASSIZOCRINUS.
Troost, Jour. Amer. Asso. Ady. Sci. Camb. Meeting, p. 60, (1850).
Wachsmuth and Springer, Revis. Paleocrin., III, p. 262, (1885).
Agassizocrinus carbonarius. Plate VI, fig. 8
Agassizocrinus carbonarius Worthen, Geol. Surv. Ill., v, p. 566, pl. xxiv,
4, (1873).
Infrabasals large, completely fused, semielliptical. The up-
per faces of the infrabasals are quite concave, and consequently
angular at the suture. The cavity is moderately deep. Meas-
urements: Height of the infrabasals, 10 mm.; diameter at their
summit, 11 mm. The faces of all five plates are equal.
Range and distribution: Upper Coal Measures; Kansas City.
The infrabasals of this species were figured by Worthen, with-
out description, from Illinois. It is the only species of the genus
known in the Coal Measures of the United States. The radials,
basals, arms, etc., of this species are not yet known.
46 University Geological Survey of Kansas.
ECHINOIDEA.
The ‘‘sea-urchins,”’ ‘‘sea-eggs,’’ ‘‘sand-dollars,’’ and ‘‘ sea-
cookies,’’ as they are variously called, are animals varying in
form from spheres to discs. They live in moderately shallow
water, especially near oyster beds, while some bury themselves
in the sand, and one kind is known to make holes for itself in
the solid rock. They always inhabit the sea. The parts that
are generally found fossil are the spines and the plates, which
are here explained.
The body of the animal is encased in a kind of box made up
of calcareous plates, which are united at their edges. A more
or less long spine is attached to each of certain plates. The
spines project out from the body in all directions. They are
attached to the plates by a sort of ball-and-socket joint, and are
movable. There are two kinds of plates—those with, and those
without, spines. The latter are called the ambulacral plates,
or the plates through which the tube-feet project, while those
with spines are called the interambulacral plates. The am-
bulacral plates are arranged in five bands, or areas, between
which the interambulacral plates are arranged. Each band
generally extends from the top to the center of the bottom of
the animal.
The mouth is located at the center of the bottom side. In
some sea-urchins the mouth is provided with five long teeth
and a complicated set of calcareous supports. The whole ar-
rangement is known as ‘‘Aristotle’s lantern.’’ On the top
side of the animal near the center there is a sieve-like plate,
which connects with a long membranous tube beneath. This
tube runs downward to the base of the inner cavity and con-
nects with another tube which forms a ring around the esoph-
agus. ‘There are five branches to this latter tube, one extending
along the center of each ambulacral area. In the ambulacral
plates there are series of holes through which the numerous
branches of these tubes extend to the outside of the shell.
These branches, or tube-feet as they are called, extend out-
- _———_ =. ~~); /
BEEDE. | Carboniferous Invertebrates. 47
ward for quite a distance and end in little disc-like suckers.
This tube system is filled with water, and is known as the
‘‘water-vascular system.’’ By filling these tubes with water,
and dilating them, the animal is enabled to move along. How-
ever long the spines may be, the tube-feet may be extended
beyond them. ‘The figures illustrate the hard parts which are
generally found fossil. There are at least four kinds known in
the Kansas rocks.
ARCHAOCIDARIS.
McCoy, Carb. Foss. Ireland, p. 173, (1844).
Archeocidaris trudifer. Plate VIII, fig. 10.
Archeocidaris trudifer White, Prelim. Rep. Invert. Foss., p. 17, (1874);
Powell’s Rep. Geol. Uinta Mts., p. 89, (1876); U.S. Geog. Surv. West
100 Mer., Iv, p. 104, pl. vi, f. 8, (1877): Keyes, Proc. Iowa Acad. Sci., 11,
p. 191, (1895).
White’s description: ‘‘Interambulacral plates compara-
tively broad, rather thin, having an elevated border all around,
which is apparently composed of aseries of small tubercles ;
areolar surface apparently plain; central tubercle small, per-
forated at the center, surrounded at its base by a very slightly
raised ring, and immediately outside of that by another ring,
which is so much elevated as to form a little cup with its rim
somewhat expanded. Diameter of the largest plate in the col-
lection about 20 mm. Spines very long and slender, one of
these in the collection having been, when perfect, about
12 cm. in length, terete; diameter of the basal ring, which
expands from the shaft, greater than that of any portion of the
shaft; diameter of the shaft nearly uniform for more than half
its length above the basal ring, the upper portion gradually
tapering to a point. Greatest diameter of the shaft of the long
spine referred to, scarcely 5mm.; diameter of the basal ring,
7mm. Surface of the spine for a short distance above the basal
ring apparently smooth, but, from that portion to the distal
end, it is ornamented with numerous small points or incipient
spinules, which are often removed by weathering, but in well-
preserved specimens they are seen to be arranged around the
spine in imperfectly spiral lines. The very long, slender, terete
spine, having a basal ring often much greater in diameter than
48 University Geological Survey of Kansas.
any part of the shaft, together with the other characters de-
scribed, distinguish this species from all others.”’ |
Range and distribution : Upper Coal Measures ; Topeka lime-
stone, Topeka. 3
Our specimen is a single spine with the proximal end buried
in the matrix, the distal three-fourths being exposed. The ex-
posed part agrees more closely with this species than any other
yet described, and it is provisionally referred to it.
Archeocidaris agassizi. Plate VIII, figs. 6-Ge.
Archeocidaris agassizi Hall, Geol. Iowa, 1, pt. 11, p. 698, pl. xxv, ff.
la-d, (1858); Keyes, Geol. Surv. Mo., iv, p. 127, pl. xv, f. 5, (1894);
Jackson, Bull. Geol. Soc. Amer., vit, p. 213, (1896); Keyes, Proc. lowa
Acad. Sci., 11, p. 185, pl. xvii, f. 5, (1895).
Hall’s description: ‘‘ Body unknown; plates small, hexago-
nal, except those adjacent to the ambulacral area, which are a
little rounded on that side, becoming pentagonal; central tu-
bercle slender, elongated, tubuliform, and projecting above the
surrounding annulation, the latter abruptly elevated, and leay-
ing, between its inner face and the central tubercle, a deep
cavity. Surface of the plate, immediately around the annula-
tion, elevated in a distinct low, annular ridge, beyond which it
is depressed and again elevated towards the margin, which is
ornamented by a series of low elongated nodes. Spines elon-
gated, compressed, contracted below and swelling out above,
so that the greatest diameter is about one-third the length
above the base; from this point very gradually tapering to the
summit. Surface of the lower contracted portion smooth, mu-
cronate above, with small spiniform tubercles, which, on the
lower part, are arranged in somewhat distant curving annulat-
ing rows, becoming more curved above, or in oblique ascending
rows, giving a quincunx order. Point of attachment somewhat
elongate, the thickened annulation strongly striate.’’
Range and distribution: Upper Coal Measures; Topeka.
This species was described from the Burlington by Hall. We
would hardly expect Coal Measures specimens to be conspecific
with the Burlington, but I am unable to detect any differences
that could be called specific in our specimens, which consist of
ee ee ee ee ee
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BREDE. | Carboniferous Invertebrates. 49
spines and plates. Without doubt, if we had entire specimens
from both formations to compare, they would be found to be
different ; but until such differences are found it is best to leave
them in the same species.
Archzocidaris megastylus. Plate VIII, fig. 7.
Archeocidaris megastylus Shumard, Trans. St. L. Acad. Sci., 1, p.
225, (1858); Keyes, Geol. Surv. Mo., Iv, p. 189, pl. xv, ff. 2a, b, (1894);
Proc.. Iowa Acad. Sci. 11, p. 189, pl. xvuui1, ff. 2a, b, (1895).
Shumard’s description: ‘‘ The interambulacral plates of this
species in the collection are large, hexagonal, wider than long,
and rather thick. The areolar surface is very broad, nearly
circular, slightly concave at its exterior portion and rising
gently to the base of the central boss. It is encircled by a
single series of small, secondary tubercles. The boss is broad,
smooth, and the central tubercle deeply perforated. The pri-
mary spines are long, robust, cylindrico-fusiform, and the
transverse section circular. The surface is very finely striated
longitudinally; and studded either rather distinct granules or
minute short spines, arranged spirally or promiscuously. The
ring at the base is oblique to the axis, its border neatly crenu-
lated, and the diameter less than the greatest diameter of the
spine. ‘The socket is deep, rather wide, and its margins smooth.
The neck is marked with a slightly raised ring, which is finely
striated longitudinally.”’
Range and distribution: Upper Coal Measures ; Kansas City.
This species often attains a length of three inches or more.
Our specimens are but spines, and they are worn so that the
minute surface markings are all removed, but, from their size
and shape, there is little doubt of their identity.
OLIGOPORUS.
Meek and Worthen, Proc. Acad. Nat. Sci. Phil., p. 474, (1860).
Oligoporus? minutus. Plate VII, fig. 3. «
Oligoporus? minutus Beede, Kans. Univ. Quart., virt, p. 126, pl. xxxu,
f. 3, (1899).
Small, depressed globular, melinitic ridges not very distinct.
There are four columns of pores, each column consisting of two
rows, in each ambulacral area; each row is generally, though
not always, made up of two pores placed side by side and very
50 University Geological Survey of Kansas.
close together; each of the two series is apparently separated
by a row or two of imperforate plates at the ambitus. Both
series are in contact at the apex and near the mouth. Number
of columns of interambulacral plates unknown, but apparently
about three. The ambulacral area is very wide and the two
series are widely separated at the ambitus. Some of the inter-
ambulacral plates seem to be pierced by a single pore, or, some-
times, two. Indistinct elevations seem to be present in three
columns, one of large and two of small size.
Diameter‘of specimen... 0.00.00 os eee ee ee eee 23 mm.
Maximum diameter of ambulacral area.................... Guna
Maximum diameter of each series...............-.-+--2-0+0: Aye Go
Maximum diameter of interambulacral area............... he Hews
Pores distant in verticalirows). 4 2 a ae ee renee ead
Pores distant in: the’same series, 6)... 9 ae ee ee eee lee OY
‘Pores distantiin single parr oan ee ee ae eee fe 8
Range and distribution: Upper Coal Measures; collected
from the Deer Creek limestone, near Topeka. Ba
The specimen is badly worn and somewhat compressed ; the
surface markings are almost entirely removed. It agrees to
some extent with Oligoporus, but the ambulacra are divided into
two series with, apparently, two columns of imperforate plates
between them. However, this is not unquestionably shown by
this specimen. It will, in all probability, be found to belong to
an entirely different genus. It is referred to Oligoporus for con-
venience, until better material can be secured. It does not
seem to present the appearance of any other Carboniferous sea-
urchin with which I am acquainted.
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BEEDE. | Carboniferous Invertebrates.
BRACHIOPODA.
Brachiopods are small animals with a two-valved shell, re-
sembling the clams somewhat in external appearance, though
they are very different in internal structure. They are in some
respects much more closely allied to the bryozoans, or sea-mats,
and worms, than to the clams. They are strictly marine
animals, living, as a rule, in rather deep water. Their distri-
bution, in this respect, has been divided into five zones,’ which
may be described as follows: Shore zone, or the beach between
high and low tide-marks; the shallow-water zone, or water to
a depth of 90 feet; the moderately deep zone, or water from 90
to 300 feet deep; the deep zone, or water from 300 to 1668 feet
deep; and the very deep zone, or water from 1668 to 17,670 feet,
or three and one-half miles deep. In each of these zones there
are species which do not occur in any of the others, while some
are common to two or more zones. There are, according to
Hall and Clarke, about 147 species of living brachiopods known,
which are distributed over the Atlantic, Pacific and Arctic
oceans, ranging from the island of Spitzbergen, north of Nor-
way, to Cape Horn, but being most abundant in tropical and
temperate waters. As arule, they are distributed along coast-
lines and in the vicinity of islands.
The living brachiopods are but a remnant of what was once
one of the most abundant and varied classes of animals of their
size that ever inhabited the earth. While the known living
forms are only 147 in number, the fossil species known at pres-
ent probably reach the enormous number of 6000, and of these,
upwards of 2000 are represented in the American rocks. They
are among the earliest fossils of which we have any record, and
culminated in the earlier part of geologic time. In the Cam-
brian, or earliest period of which we have any definite knowl-
edge of life, there are about 125 species known; in the
3. This introduction is largely drawn from Eastman’s Translation of Zittel’s Handbook of
Paleontology, Schuchert’s Synopsis of American Fossil Brachiopoda (Bulletin of the U. S.
Geological Survey No. 87), Paleontology of New York, vol. VIII, and Annual Report of the State
Geologist of New York for 1891, to which the reader is referred for more detailed information.
52 University Geological Survey of Kansas.
Devonian they reached their highest point in respect to num-
bers, about 1400 species; while at the close of the Paleozoic era
they appear to have fallen back to less than 100 species, and to
have continued to the present comparatively few in number.
Before entering upon the description of the Kansas species, it
may be well to give an idea of what the animal was and what
the markings found on the shells mean. In the first place, there
are two shells, or valves, generally unequalin size. The hinge,
or the place of union of the two valves, is considered the pos-
terior, or hind end, and the opposite edge the front end, while
the two sides at right angles to the hinge are the lateral mar-
gins or sides. The animals were attached to some foreign object
during all or a part of their life, either by a pedicle, along mus-
cular projection from the shell, or by a portion of the shell being
cemented to the object. Those having a pedicle usually have
an opening in the beak on one of the valves, the pedicle valve,
sometimes called the ventral valve, near the hinge, while the
others simply have the pedicle extended between the posterior
ends of the valves. :
The front portion of the shell is lined by a membrane which
divides the cavity into two parts, the anterior or brachial cavity,
and the posterior or visceral cavity. In the latter are located the
visceral organs and the muscles which open and close the shell
and retract and protude the pedicle. ‘The nervous system con-
sists of a single ring around the esophagus, in which there are
located two ganglia which give off branches to the different
organs of the body. The digestive canal consists of but a single
convolution, and terminates blindly in the living forms, though
it probably communicated with the exterior in certain fossil
forms. ‘The pair of muscles which close the shell, the adductors,
are large muscles which extend directly from the postero-central
part of one valve to the other. By their contraction the shell
is closed. The impression left in the shell by these muscles is
usually quite prominent. The diductors, or muscles which open
the shell, are large muscles attached to the middle of the rear
end of the pedicle valve just outside of the adductors, while the
other end is attached to a prolongation of the other, brachial,
VN,
ww
=p,
Fic.3. A, Lingula murphiana (after Hall and Clarke), showing the pedicle, which is
greatly developed in this genus; B, Magellania flavescens (after Schuchert, in Eastman’s
Trans. of Zittel); C, Seminula argentia; D, Productus giganteus (after Schuchert, in East-
man’s translation of Zittel); E, Derbya robusta; F, Productus semireticulatus; a, adduc-
tors: ec, diductors; d, spiral brachia; 6s, same as seen in a fossil; dl, diverging lines; dif,
deltidium: as, adductor scar; ca, cardinal area; ep,cardinal process; /, foramen; /, fringe
on brachia; fm. hinge line; Ir, ridge diverging from base of cardinal process; 758, mesial
septum; /it, teeth; z, alimentary canal: vr, mouth; vm, brachial markings.
54 University Geological Survey of Kansas.
sometimes called dorsal valve. This process extends beyond the
hinge backward under the beak of the other valve, and when
the muscle contracts acts as a lever opening the shell.
In the front cavity are situated the brachia or arms, as they
are called. They are often of coiled or spiral form, supported
by horny or calcareous supports which are sometimes preserved
in the fossils. These brachia carry a long fringe, which by its
vibration keeps a current of water flowing through the shell,
from which the animal separates its food. There are no special
organs of respiration, which is performed probably by the
blood flowing through many thin membranes which are con-
stantly in contact with the current of water passing through the
shell. The figures on page 53 serve to illustrate the parts of
the animal and the muscular impressions found in the shells.
The full synonymy is given for the species herein described,
but not afull bibhography. Fora full bibliography, the reader
is referred to Mr. Charles Schuchert’s work on the North Ameri-
can Brachiopoda, Bulletin No. 87 of the United States Geological
Survey of 1897. 3
LINGULA.
Bruguiere, Encye. Metho.. I, p. 250, (1792).
Meek and Hayden, Pal. Upp. Mo., p. 68, (1861) ; ete.
Lingula mytiloides. Plate VIII, fig. 5.
Lingula mytiloides Sowerby, Min. Conch., 1, p. 55, pl. x1x, ff. 1, 2, (1813);
Meek and Hayden, Geol. Surv. IIll., v, p. 572, pl. xxv, f. 2, (1873).
? Lingula umbonata Keyes, Geol. Surv. Mo.,v, p. 38, pl. xxxv, f. 4, (1895).
Shell small, elliptical, posterior extremity slightly narrowed ;
beak extending to the hinge line, not prominent. Anterior
margin broadly rounded, lateral margins subparallel; the great-
est diameter about the middle of the shell. The surface is
marked by fine concentric lines of growth. Length, 7 mm.;
breadth, 5 mm.; larger specimen, length, 11 mm.; breadth,
1 ies 3 019 0.0 We
Range and distribution: Upper Coal Measures; Kansas City,
Topeka, Burlingame.
: . ; : ait
~ ie ee ee
a + eB i ~ 5 ——— EE eae ee ee
ee ee ee ee ee a he i ae ae | J sal - a
—_ oe
BEEDE. | Carboniferous Invertebrates. 55
ORBICULOIDEA.
d’Orbigny, Prodrome de Pal. Strat., I, p. 44, (1850).
Dall, Bull. Mus. Comp. Zool., III, p. 37, (1871); ete.
Orbiculoidea missouriensis. Plate VIII, figs. 1-lc.
Diseina missouriensis Shumard, Trans. St. L. Acad. Sci., 1, p. 221, (1858).
Diseina nitida? Meek and Worthen (non Phillips), Geol. Surv. Ill.. v,
p. 572, pl. xxv, f. 1, (1873): White, 13th Ann. Rep. St. Geol. Ind., p. 121,
pl. xxv, f. 10, (1884); etc.
Discina meekiana Whitfield, Ann. N.Y. Acad. Sci., 11, p. 288, (1882); ete.
Orbiculoidea missouriensis Schuchert, Bull. 87, U.S. Geol. Surv., p. 281,
(1897).
White’s description (in part): ‘‘ Shell small, subcircular, de-
pressed conical, the sides sloping nearly straight from the apex
to the margin; apex prominent, situated about one-third the
diameter of the shell from the posterior border ; lower valve flat
with the usual depression around the foramen; surface of both
valves marked by concentric lines and fine lamellations. Di-
ameter of an average example about 8 mm.”’
In this shell the height of the apex is about one-third the di-
ameter of the shell. The concentric lines are prominent in
well-preserved specimens and are sharply elevated, separated
by wider, shallow furrows. The sulcus extends two-thirds or
all the distance to the posterior margin. The inner lamine of
the dorsal valve show a radiate structure.
Range and distribution: Upper and Lower Coal Measures ;
Fort Scott; Rosedale, Wyandotte county; Lansing, Leaven-
worth county; Topeka.
Orbiculoidea convexa. Plate VIII, figs. 3, 3b.
Discina convexa Shumard, Trans. St. L. Acad. Sci., 1, p. 221, (1858);
White, 13th Ann. Rep. St. Geol. Ind., p. 121, pl. xxv, f. 9, (1884); etc.
Orbiculoidea convexa Schuchert, Bull. 87, U.S. Geol. Surv., p. 278, (1897).
White’s description: ‘‘ Upper valve broadly but somewhat
prominently convex; subcircular in marginal outline; the
height nearly equal to one-half the diameter; apex somewhat
obtuse, but moderately prominent, situated about one-third the
diameter of the sheil from its posterior margin; surface marked
by the usual distinct lines of growth. A small under valve was
found in the same locality as the upper valve above described,
and probably belongs to this species. It shows a similar sur-
56 University Geological Survey of Kansas.
face, which is nearly flat, but it is depressed above the foramen,
which is of the usual character; the foramen is situated just
beneath the beak of the upper valve.’’ The posterior portion
of the shell of one individual which has the outer portion of
the shell exfoliated shows distinct, discontinuous, alternating,
radiating ridges, from near the beak to the posterior margin.
‘¢Diameter of the upper valve just described, 27 mm.; height,
2 ac
Range and distribution: Upper Coal Measures ; Kansas City,
Topeka. 3
Orbiculoidea manhattanensis. Plate VIII, figs. 2-2b.
Discina manhattanensis Meek and Hayden, Proc. Acad. Nat. Phil.,
(1859), p. 25.
Orbiculoidea manhattanensis Hall and Clarke, Introduction to Study
of Brach., pt. 1, pl. v, £. 12, (1892); Pal. NovY., vim, pt: plana.
(1892).
Shell of medium size for this genus, nearly circular, dorsal
valve moderately elevated ; umbo situated about one-half of a
diameter from the posterior margin; surface of dorsal valve
marked with concentric lines, which are much heavier near the
margin and finer and more wavy near the apex. The anterior
slope is slightly convex, posterior a little concave, distinctly so
in casts. The ventral valve is flat, shell quite thick; sulcus in
distinct depression, extending about half way from the center
to the margin; opening linear-elliptical. There is a thick
callosity near the outer margin which is marked by radiating
ridges corresponding to those of the other valve. Diameter of
large specimen, 12 mm.; height, 25 mm. Outer surface
marked by concentric lines similar to the convex valve.
Range and distribution: Upper Coal Measures; Wabaunsee
formation; black shale in wagon-road cut east side of Blue
Mount, Manhattan, Kan. In collection of E. A. Popenoe.
This species differs from O. missouriensis in being more com-
pressed, thicker shelled, shorter sulcus, and in having more
distinct concentric lines; from QO. convexa, in being much less
convex and much smaller.
.
2
4
:
a
‘
4
i
¥
s
BREDE. | Carboniferous Invertebrates. 57
CRANIA.
Retzius, Schrift. Ges. Naturf. Freuende, Berlin, II, p. 72, (1781).
Hall and Clarke, Pal. N. Y., pt. I, p. 145, (1892) ; ete.
Crania modesta. Plate VIII, fig. 4.
Crania modesta White and St. John, Trans. Chic. Acad. Sci., 1, p. 118,
(1868); White, 13th Rep. Ind. St. Geol., p. 121, pl. xxxv, f. 9, pl. xxxv1,
f. 5, (1884).
Crania carbonaria Whitfield, Ann. N. Y. Acad. Sci., 11, p. 229, (1882);
Geol. Ohio, vii, p. 484, pl. x1, ff. 11, 12, (1895); ete.
Whitfield’s description: ‘‘ Shell small, none of the specimens
observed exceeding three-eighths of an inch in diameter ; subor-
bicular in outline, or varied in form by the outline of the objects
to which they are attached. Free valve depressed convex,
marked by a few concentric lines of growth; attached valve
thin, but witha slightly thickened margin. Posterior muscular
impressions large and submarginal, the others being nearly
central and forming a small elevation just posterior to the
middle of the valve.’’ Measurements: Diameter, 9 mm.;
height, 3 mm.
Range and distribution: Upper Coal Measures; Kansas City,
Eudora, Grand Summit.
The specimens are attached to Productus semireticulatus, and
the convex valves reproduce the markings of the host. One
near the front of the shell shows the large radiating costie quite
plainly, while on the reticulated area shows both the radiating
and concentric marks, while still another situated partly on the
reticulated area shows the reticulated markings on one side and
only the radiating on the other. Two specimens on worn shells
show none of the markings of the host. The attached valves
are thin, but show the muscular markings quite distinctly.
DERBYA.
Waagen, Pal. Indica, ser. XIII, I, pp. 576, 591, (1884).
Hall and Clarke, Pal. N. Y., VIII, pt. I, p. 261, (1892); 11th Ann. Rep. N. Y. St. Geol.,
p. 286, (1895), for 1894.
The genus Derbya has been divided by Waagen into groups,
but I see no sufficient reason for such divisions for our American
shells.
A consideable stress has been laid upon the nature of the
hinge area in the different American species of the genus. |
5—vi
58 University Geological Survey of Nansas.
find, upon examination of the Kansas Coal Measures forms,
that the length of the hinge and the height of the area, as well
as the relative convexity and bilobation of the brachial valve,
are quite variable. The diverging lines which pass from the
apex of the deltidium obliquely to the hinge line, nearly bisect-
ing the area, are common to all the well-preserved specimens
from the Coal Measures that I have examined. The same is
true of the vertical striation of the area between these lines.
The area between the diverging lines is composed of two
large dental plates not entirely united to the remainder of the
hinge area. This is clearly shown by specimens with the area
crushed. In such cases it almost always breaks along these
lines with smooth fracture.
They also present some striking peculiarities in the surface
markings of the shell. Upon a cursory examination there
seem to be two distinct kinds of surface markings — those shells
in which the strive are small, somewhat rugose, separated by
wider channels; and those with more rounded, crowded, rugose
ridges with narrow valleys. Upon examination of several
specimens of different species, the difference in these respects is
found to be due, either to weathering or to wearing previous to
fossilization. ‘The striae are rounded, thicker at the top of the
ridge than at the base, furrows and ridges crossed by numer-
ous fine, concentric lamelle, which are raised into ruge on
the ridges and nearly imperceptible in the furrows. In the
slightly worn specimens a portion of the top and sides of the
ridges are worn away, making them appear keel-shaped, with
wider furrows. ‘The wearing of the shell does not seem to di-
minish the roughness of the strive very perceptibly, while it
seems to bring out the concentric markings of the furrows more
distinctly.
ee ee ee ae nl
BREDE. | Carboniferous Invertebrates. 59
Derbya bennetti. Plate VIII, figs. 8—8e.
Derbya bennetti Hall and Clarke, Pal. N. Y., vir, pt. 1, pp. 263, 348, pl.
. x1-A, ff. 34-39, (1892); Ann. Rep. St. Geol. N. Y. 1894, p. 347, pl. v, ff. 3-8,
(1895).
Original description: ‘‘Shell subtrihedral in general aspect,
quite irregular in its growth. Hinge line short, its extremities
on both valves being auriculate. Pedicle valve much more
irregular in growth, sometimes retaining the scar of attachment
at its apex. Cardinal area usually high, narrow, erect or
slightly incurved, and frequently distorted ; delthyrium curved.
General surface of the valve depressed, convex in the middle,
sometimes rapidly sloping in all directions, at others concave in
the umbonal region; as a rule very unsymmetrical. The
brachial valve is deep, more regularly convex, and has a full
rounded umbo and a conspicuous median sinus. On the in-
terior the pedicle valve bears an extremely high median septum
which is united with the dental ridges near the apex. The
cardinal process is high, erect and deeply bilobed, each of its
apophyses being strongly grooved on its posterior face. Other
internal characters unknown. The surface of both valves is
covered by fine, elevated, thread-like strive, increasing very
slowly by intercalation. The edges of these strie bear numer-
ous minute asperities which may be due to the crossing of fine
concentric lines. Irregular lines and wrinkles of growth are
| abundant near the margins.’’
Range and distribution: Upper Coal Measures ; Lecompton.
Mr. Bennett informs me that the type specimen which was sent .
Professor Hall, and upon which the description was based, was
from near St. Joseph, Mo., instead of Kansas City.
Hall and Clarke described another species from Kansas City
(D. broadheadi ) with very much the same characteristics as the
species given above. Among the specimens before me I am
unable to separate the two satisfactorily, and believe that they
will have to be united under a single species. D. broadheadi, in
some of its forms, approaches JD. affinis so closely, being so
slightly bilobate, that the two forms seem to merge completely,
as well as the characters of the beaks, which also approach each
i
60 University Geological Survey of Kansas.
other closely, so that it would be better to leave D. bennetti for the
species, should the two be found identical. D. broadheadi may,
perhaps, be varietally distinct from D. bennetti, but our mate-
rial would hardly seem to indicate it.
Derbya cymbula. Plate XII, fig. 10.
eae NUT Hall and Clarke, Pal. N. Y., vit, pt. 1, p. 348, pl. x1-B,
ff. 2, 3, (1892); Rep. N. Y. St. Geol. 1894, p. 348, pl. v1, ff. 1,2 2, (1895):
Oieiasl description (in part): ‘‘Shell large ; marginal outline
transversely subelliptical. Hinge line straight, its length be-
ing about two-thirds the greatest diameter of the shell. On
the pedicie valve the cardinal area is high, its base being one-
third longer than its sides, and it may be somewhat unsym-
metrical from distortion. Its surface is finely striated both
longitudinally and transversely, and is divided into an outer
and inner portion by two lines diverging from the apex and
meeting the hinge line half way between its extremities and
the edges of the deltidium. Deltidium broad at the base, rap- ~
idly narrowing for one-third its length, thence tapering more
gradually to the apex; its surface is marked by a well-defined
median groove for its entire extent. The surface of the valve
is elevated in the umbonal region, and slopes irregularly to a
low depression over the pallial region and about the margins.
The brachial valve is broadly concave at the umbo, but rapidly
becomes regularly convex, the greatest convexity being in the
middle of the valve, whence it slopes almost equally in all di-
rections. There is no tendency to irregular growth in this
valve. Surface covered with numerous fine, sometimes irregu-
lar strize, increasing by implanation. Over the umbonal and
pallial regions these strie are of about equal size, but above
the margin the tendency to fasciculate arrangement is more
apparent.’’ The mesial septum of the pedicle valve very high,
attach to the teeth, which extend to the top of the deltidium
for about a third of its height, and extending about a third of
the distance to the front of the shell, highest at the anterior
end. Teeth of the brachial valve narrow, extending well into
the opposite valve, where they curve inward, locking around
j
BREDE. | Carboniferous Invertebrates. 61
a process from the cardinal process of the brachial valve.
Cardinal process of the brachial valve large, elongated, and
expanded laterally, divided by a deep sinus at the summit,
each lateral lobe having a very deep groove on its posterior
side extending nearly its entire length. The cardinal process
is yery strongly recurved backwards, fitting closely into the
deltidium of opposite valve, and backing the process on either
side are two high plates extending well back into the visceral
cavity, and attached to the brachial valve. Measurements:
Length (hinge to front), 45 mm.; width, 58 mm.; convexity
(maximum), 39 mm.; length of hinge, 35 mm.; height of
area, 13 mm.
Range and distribution: Upper Coal Measures, and base of
Permian? Kansas City, Cambridge, Cowley county.
This species can be easily recognized and separated from the
preceding by its larger size, relatively lower beak (as a rule),
and its non-bilobate brachial valve. Hall and Clarke have
described another species from Kansas City, D. affinis, which
is, to say the least, exceedingly closely related to D. cymbula,
the principal difference being in the location of the point of
greatest conyexity in the pedicle valve and smaller size. So
far as the topography of the pedicle valve is concerned in this
genus, except, perhaps, the relative length and height of the
hinge area, it seems to be governed largely by the immediate
surroundings of the individual. The location of the point of
greatest convexity, and, even as to whether or not there is a
concavity in the valve, vary greatly in those species which
have a convex or raised valve. In the specimens of D. cymbula
in our collection I find no two with the same topography of the
pedicle valve, and they vary from regularly convex from the
apex of the beak to the extreme front margin to irregularly
concave over the same area. Some are regular in their growth,
while others are very irregular. It seems to me that the speci-
mens described as D. affinis are young or stunted forms of D.
cymbula, for it is almost impossible to separate the two forms
at all, if we possess the specimens of the two species, and we
have specimens that answer the descriptions very closely.
62 University Geological Survey of Kansas.
Derbya crassa. Plate VIII, figs. 11, 11b.
Orthis arachnoides Roemer (non Phillips), Kreidebildung Texas, p. 89,
pl. x1, f. 9, (1852); ete.
Orthisina crassa Meek and Hayden, Proc. Acad. Nat. Sci. Phil. 1858, p. 261.
Orthis lasallensis McChesney, Desc. New Pal. Foss., p. 32, (1860); ibid.,
plod, £.°6) (sb)?
Orthis richmonda McChesney, ibid., p. 32; also pl. 1, f. 5
Hemipronites crassus Meek and Hayden, Pal. Upp. Mo., Smiths. Cont.
Knowl., xtv, 172, p. 26, ue 1, f. 7, (1864); Meek, Fin. Rep. U.S. Geol.
Surv. Neb., p. 174, pl. v, f. 10, pl. viz, f. 1, (1872); ete.
Orthis crenistria Geinitz (non Phillips), Carb. u. Dyas in Neb., p. 46,
pl. 111, ff. 20, 21, (1866).
Hemipronites lasallensis McChesney, Trans. Chic. Acad. eae Dapaeoe
plat f. O(elsos))e
Hemipronites richmonda McChesney, ibid., p. 28, pl. 1, f. 5.
Hemiproniies crenistria White, Wheeler’s Expl. Surv. West. 100 Mer.,
Iv, p. 124, pl. x, f. 9, (1875).
Streptorhynchus richmondi Hall, 2d Ann. Rep. N. Y. St. Geol., pl. x1,
i085 IO) IIL 5 (Wketess}))4
Derbya crassa Waagen, Pal. “teeta, ser. XIII, I, p. 592, (1884); ete.
Meek’s description (in part): ‘‘Shell varying from nearly cir-
cular to truncato-subcircular, or transversely suboblong, generally
wider than long, varying from compressed to distinctly convex ;
hinge margin equaling or shorter than the greatest breadth of
the valves, rectangular, or sometimes more or rather less than
rectangular, at the extremities ; anterior outline forming a more
or less regular semicircular curve. Dorsal valve always convex,
sometimes very distinctly so, the greatest convexity being near
the middle; beak not distinct from the cardinal margin. Ven-
tral valve varying in convexity at the umbo, sometimes very
prominent, and occasionally distorted there; less convex, flat-
tened, or not unfrequently a little concave, around near the
front; area varying in height in proportion to the elevation to
the beak, and either flat or with the beak a little arched, usually
rather distinctly striated ; its closed fissure varying in the pro-
portions of height and breadth with the greater or less eleva-
tion of the beak; interior always provided with a prominent
mesial septum extending from the beak forward to near the
middle of the valve; surface on both valves marked by numer-
ous strong, raised radiating strie of unequal size, there being
generally one or several smaller ones between each two of the
larger ; crossing the whole are also numerous fine, regular con-
j
ee
oo Se ee ee oe SS ae Se
BEEDE. | Carboniferous Invertebrates. 63
centric striz, more or less defined both between and upon the
radiating striw, to which latter they impart a neatly crenate ap-
pearance.’’ The dental lamelle are narrow and thin in younger
specimens, more rounded and heavier in old ones, extending
the entire length of the deltidium and projecting beyond the
hinge margin. Muscular impressious of this valve are usually
indistinct in young specimens, but are well impressed in the old,
thick-shelled individuals, varying from nearly circular to linear-
elliptical in outline, in some cases extending two-thirds the
distance to the front of the shell, though as a rule it only covers
the central portion of the posterior half of the valve. There is
a faint mesial ridge in the posterior portion of the brachial
valve, extending about half way to the front in old specimens.
The muscular scar is nearly circular and marked by longitudinal
ridges extending about half way to the front and sides of the
valve. Cardinal process moderately prominent, bifid atits apex,
and possessing a lateral lobe on each of the supporting plates
near their anterior portions. Measurements: Length, 25 mm. ;
width, 31 mm.; length of hinge, 20 mm.
Range and distribution: Upper and Lower Coal Measures ;
Fort Scott, Kansas City, Lawrence, Topeka.
This little depressed or moderately convex shell is very easily
distinguished from the other species by its smaller size, moder-
ately low hinge area, and thick, or sometimes thin, shell.
Derbya keokuk. Plate VIII, fig. 13; text fig. 3, E.
Orthis crenistria Yandalland Shumard, Cont. Geol. Ky., pp, 19, 21, (1847).
Orthis keokuk Hall, Geol. Surv. Iowa, 1, pt. 0, p. 640, pl. xrx, f. 5,
(1858); ete.
Streptorhynchus keokuk Hall, 2d Ann. Rep. N. Y. St. Geol., pl. x1, ff.
1-3, (1883).
Streptorhynchus erenistria Walcott, Mon. U.S. Geol. Surv., viii, p. 279,
pl. xviit, f. 14, (1884).
Derbya keokuk Hall and Clarke, Pal. N. Y., vir, pt. 1, p. 262, pl. xz, ff.
1-3, (1892).
2 Orthis umbraculum Owen (non Schlotheim ), Geol. Sury. Wis., lowa, and
Minn., pl. v, f. 11, (1852).
2 Orthis robusta Hall, Geol. Surv. Iowa, 1, pt. 1, p. 743, pl. xxvii, f. 5,
(1858).
? Streptorhynchus robusta Hall, 2d Ann. Rep. N. Y. St. Geol., pl. x1, ff.
12-17, (1883).
?Derbya robusta Waagen, Pal. Indica, ser. x11, 1, p. 592, (1884); Hall and
Clarke, Pal. N. Y., v111, pt. 1, p. 262, pl. x, ff. 12-17, pl. x1B, ff. 7, 8, (1892).
64 University Geological Survey of Kansas.
Hall’s description (in part): ‘‘Shell resupinate, somewhat
broadly semielliptical in outline, depressed hemispheric; car-
dinal extremities rounded. Ventral valve flat or slightly con-
cave; area low, extending to the hinge extremities; foramen
forming an equilateral triangle, closed by a pseudo-deltidium.
Dorsal valve broadly convex, the greatest convexity a little
above the middle, and often equal to one-third the width of the
shell. Surface marked by even rounded radiating strie, which
increase by bifurcation and interstitial addition, and are crossed
by fine concentric strie.’’ Interior of pedicle valve marked by
large, deep, semicircular impression, with raised irregular per-
iphery, moderately high mesial septum extending to the front of
the muscular impression, which is marked by irregular radiat-
ing ridges and furrows of variable size. Area of the valve out-
side of the impression finely pitted to the edge, where the
striations of a new layer of shell form a radiate periphery.
Hinge area possessing the usual diverging lines, within which
are vertical strie. Entire hinge area marked by horizontal
lines or lamelle. :
Range and distribution: Upper Coal Measures to Permian ;
Kansas City, Carbondale, Topeka, Cambridge, Cowley county.
This shell agrees in many respects with the description of D.
keokuk and in others with D. robusta. If the previous remarks
on surface markings of the valves and the hinge area hold good
for the genus, there is little doubt that the two will prove syn-
onymous. The fact that our Coal Measures shells have the out-
line and also the interior of the pedicle valve almost exactly as
D. robusta and, where the shells are not worn, the markings of
D. keokuk, would seem to indicate that they are the same.
Specimens broken from limestone always leave a portion of
the shell attached to the stone, which makes the specimens
look like those with the markings ascribed to D. robusta. Some
of our specimens differ from D. keokuk in that the hinge is
hardly as short as ascribed to that species, though some of
them approach it quite closely. However, the length of the
hinge is not a very constant character in any species of the genus
that I have examined. The front and lateral margins expand
BeEDE. | Carboniferous Invertebrates. 65
much more rapidly than does the hinge after the sheli reaches
middle age.
Professor Clarke writes me concerning these two species that
‘“There can be no question of the close similarity of these two
species, and D. robusta was founded upon a single specimen
remarkable for the convexity of its brachial valve and its sub-
circular outline, but I recollect distinctly that we had a com-
paratively meager representation of this species from the Coal
Measures. I have little doubt that one represents the continu-
ation of the other specific type, perhaps with some slight varia-
tion, and that the genus attained its culminant variability of
expression in the Coal Measures.”’
The relative convexity of the brachial valve varies to a con-
siderable degree, though the most of them are very convex. Con-
sidering these points as a whole, the only grounds left, so far as
I can see, for D. robusta to rest upon, are its relatively little
longer hinge and greater convexity of the pedicle valve, both of
which are variable characters, and the two species approach each
other very closely in this respect. However, it may be suff-
cient to distinguish them, though I think that they will prove
to be the same.
MEEKELLA.
White and St. John, Trans. Chic, Acad. Sci. I, p. 120, ff. 4-6, (1868).
Hall and Clarke, Pal. N. Y. VIII, pt. I, p. 264 (1892) ; ete.
Meekella striatocostata. Plate XII, figs. 9—-9c.
Plicatula striatocostata Cox, Geol. Surv. Ky., 111, pt. 1, p. 568, pl. viii,
f, 7; (1857).
Orthisina shumardianus Swallow, Trans. St. L. Acad. Sci., 1, p. 183,
(1858).
Orthisina missouriensis Swallow, ibid., p. 219: ete.
Orthis striatocostata Geinitz, Carb. u. Dyas in Neb., p. 48, pl. 111, ff. 22-
24, (1866).
Meekella striatocostata White and St. John, Trans. Chic. Acad. Sci., pp.
129, 122, ff. 4-6 (1868): Meek, Fin. Rep. U. S. Geol. Surv. Neb., p. 175,
pl. v, f. 12, (1872); ete.
Streptorhynchus (Meekelia) striatocostata Hall, 2d Ann. Rep. N. Y. St.
Geol., pl. x, ff. 18-23, (1883).
Meek’s description (in part): ‘‘ Shell trigonal-subglobose, be-
coming very convex with age, generally a little longer than
wide ; hinge line very much shorter than the breadth of the
valves. Dorsal valve convex, the greatest prominence near the
66 University Geological Survey of Kansas.
umbo, thence rounding over to the front, being usually some-
what flattened over the central and anterior regions, but with-
out any mesial sinus; beak strongly incurved, and with its
most prominent part sometimes projecting a little beyond the
hinge line, but in others flattened, and with its immediate
apex nearly always terminating at the margin of the hinge;
posterior lateral margins laterally compressed and converging
toward the umbo at nearly aright angle; surface ornamented
by about ten to thirteen large, radiating, more or less angular,
simple or rarely bifurcating plications, which are themselves
(as well as the furrows between) marked by fine but distinct
radiating strive, which, toward the front, instead of continuing
parallel to the furrows and plications, converge forward on each
side of the latter so as to intersect along the crests of the same
at acute angles. Crossing all of these, there are usually toward
the front and lateral margins a few strong zigzag marks of
growth. Ventral valve more convex than the other, the great-
est convexity being at or near the beak, which is elevated and
usually more or less distorted, being sometimes twisted to one
side, and in other examples straight or somewhat arched back-
ward; cardinal area narrow transversely, but proportionally
high, being often distinctly higher than wide, but well defined,
and usually finely striated transversely and vertically, either
flat or more or less arched backwards; false deltidium closing
the fissure, narrow, and provided with a slender, rounded,
prominent mesial ridge extending to the apex of the beak; sur-
face as in the other valve.’?’ The two dental lamelle of the
pedicle valve extend from the apex of thé beak to the hinge
and forward about half the distance to the front of the shell,
dividing the beak into three nearly equal compartments. The
cardinal process of the brachial valve is curved forward and
upward, long, and thickened at the extremity. Measurements
of average specimen: Length, 23mm.; width, 29mm. ; convex-
ity, 18 mm.; length of hinge, 14 mm. ; height of cardinal area,
ga danan\s
Range and distribution: Upper and Lower Coal Measures ;
Fort Scott, Olathe, Kansas City, Eudora, Lawrence, Lecomp-
BEDE. | Carboniferous Invertebrates. 67
ton, Topeka, Beaumont, Grand Summit. Widely distributed,
but moderately rare throughout the Coal Measures and base of
the Permian. Abundant near the base of the Permian.
The individual variation of this species is very great. In
most of the larger specimens the width is considerable greater
than the length as indicated by the above measurements. The
distinctness of the plications varies with age, being very faint
in young specimens and very distinct and sharp in old ones.
Young specimens are much less convex and comparatively
longer than the old ones. The comparative height of the car-
dinal area varies greatly, some-specimens having the beak only
moderately elevated.
CHONETES.
Fisher and Waldheim, Oryct. du Gouv. de Moscou, pt. II, p. 134, pl. XX VI, ff. 8, 9, (1837).
eee Pakage Lei,
Prior to 1844 species belonging to this genus were described
under the generic terms Pecten, Hysterolythes, Terebratulites,
Leptxna, Orthis, Spirifera, Strophomena, Productus, and Delthyris.
Since that time the genus has been better understood.
After giving the generic description (loc. cit.) , Hall and Clarke
say of the genus: ‘‘(’‘honetes is remarkable for the persistence
of its characters. [rom its appearance in the middle of the
Silurian to its disappearance in the Permian, this type of struc-
ture has been maintained with few essential modifications. On
account of this stability in its features it is difficult to establish
any satisfactory subdivision of its members, especially since the
genus has been left more compact by the recent elimination of
some of its more aberrant forms.’’
The following is asummary of the sections of the genus, as
compiled by them, now accepted by leading authorities ; these
sections are mechanical and arbitrary :
I.—Concentrice, those with concentric folds or undulations, like (’. con-
centrica.
I1.—Striate, those with more than thirty radiating strie.
IlIl.—Plicose, those with less than thirty striz.
IV.—Rugose, those with rugose radiating striz.
V.—Grandicostate, those with high, strong radiating costz.
V1I.—Leves, those with smooth shells.
68 University Geological Survey of Kansas.
All of our Kansas shells fall within two of these sections, the
second and the sixth. The Striatx include: C. mesolobus, ver-
nuilianus, and granulifer. The Leves include C. glaber Gein.
Chonetes glaber. Plate IX, fig. 2.
Chonetes glaber Geinitz, Carb. u. Dyas in Neb., p. 60, pl. Iv, ff. 15-18,
(1866); Meek, Fin. Rep. U.S. Geol. Surv. Neb., p. 171, pl. xiv, f. 10, pl.
vill, ff. 8-8e.
Chonetes geinitziana Waagen, Pal. Indica, ser. x1ir, vol. 1, p. 261, (1884).
Chonetes levis Keyes, Proc,. Acad. Nat. Sci. Phil., p. 229, pl. x11, f. 3:
Geol Surv. Mo. vp) 2d) pl) Xxx varie heron lS9a))r
Chonetes geinitzianus Miller, N. Amer. Geol. and Pal., p. 339, (1889).
Chonetes glaber Schuchert, Bull. No. 87, U. S. Geol. Surv., p. 74, (1897).
Meek’s description: ‘‘Shell thin; transversely subsemicir-
cular, length being more than half the breadth; hinge line a
little longer than the greatest breadth of the valves, at any
point farther forward ; lateral extremities abruptly pointed and
sometimes slightly recurved ; anterior and anterior lateral mar-
gins, forming a semicircular curve in outline, excepting that
the former is generally faintly sinuous in the middle; lateral
margins curving abruptly outwards just before intersecting the
hinge extremities. Pedicle or larger valve moderately convex,
the most gibbous part being in the form of two broad, rounded,
undefined prominences, which diverge from the beaks to the
anterior lateral regions, leaving a rather broad, rounded, deep
mesial sinus between them, extending nearly to the beak, but
widening and deepening rather rapidly to the front; outside of
these prominences the posterior lateral regions are more or less
compressed ; beak small, compressed, slightly arched, and
scarcely projecting beyond the cardinal margin; area narrow,
inclined obliquely backward; its fissure small, nearly semi-
circular and partially closed by the cardinal process of the other
valve ; cardinal margin armed on each side of the beak by four
or five slender, moderately long, oblique spines, with sometimes
remains of one or two much smaller rudimentary additional
ones near the beak; cardinal teeth compressed, their longer
diameter ranging nearly parallel to the hinge line—as seen
under a strong lens, finely striated on the outside, at right
angles to their length. Interior, excepting the regions of the
a
:
ln i ee ee le
Ss —
BEEDE. | Carboniferous Invertebrates. 69
muscular impressions, with numerous, rather distinct granules,
arranged in radiating rows—immediately within the fissure,
provided with a short, rather prominent, compressed ridge
ranging at right angles to the hinge; muscular impressions
very obscure.
‘* Dorsal or concave valve, following nearly the curve of the
other; area of about the same size as in the dorsal |ventral|
valve, inclined forward from the hinge; cardinal process small,
not yery prominent, and, as seen on the outer side, somewhat
trilobate, the middle lobe or ridge being divided by a linear
sulcus; socket ridges very oblique; interior granulated as in
the other yalve. A very small, obscure linear ridge occupies
the middle of the valve, without, however, extending up to the
hinge ; muscular impressions unknown.
‘‘Surface of both valves nearly smooth, but showing obscure,
concentric marks of growth. In some conditions of weathering
there is a faint appearance of radiating markings, but this is
due rather to the structure of the shell and not to proper ex-
ternal lines. When a single one of the thin valves is cleaned,
and examined by the aid of a good magnifier and a strong trans-
mitted light, very scattering punctures or pores are seen arranged
in quincunx, and passing obliquely through the shell. These
appear to have been connected with minute tubular spines,
arranged over the whole surface, during the life of the animal.’’
Range and distribution: Upper Coal Measures; Topeka.
Chonetes granulifer. Plate IX, figs. llc.
Chonetes granulifera Owen, Geol. Rep. Lowa, Wis., and Minn., p. 583,
tab. v, f. 12, (1852); Meek, Fin. Rep. U. S. Geol. Surv. Neb., p. 171, pl.
iv, f, 19, pl. vit, f..8, (1872 );. etc.
Chonetes smithii Norwood and Pratten, Jour. Acad. Nat. Sci. Phil., 111,
p. 24, pl. 11, f. 2, (1854); ete.
Chonetes mucronata Meek and Hayden, Proc. Acad. Nat. Sci. Phil., p.
262, (1858); Pal. Upp. Mo., p. 22, pl. 1, f. 5, (1864); ete.
Chonetes granuliferus Beecher, Am. Jour. Sci., 3d ser., xu1, p. 357, pl.
xvit, f. 15, (1891).
Chonetes granulifer Schuchert, Bull. 87, U.S. Geol. Surv., (1897).
Meek’s description: ‘‘Shell attaining a rather large size,
semicircular in outline, having its greatest breadth on the
hinge line, which often terminates in extended mucronate ears.
70 University Geological Survey of Kansas.
Larger or ventral valve moderately convex, the greatest con-
vexity being in the central region, or rather on each side of it,
as there is usually a broad, shallow, mesial depression; ears
and lateral regions compressed ; front somewhat straightened
along the middle; beak small, rather compressed, a little
arched, and scarcely projecting beyond the cardinal margin,
which is provided with from seven to eleven oblique spines on
each side of the beak; area rather narrow, ranging nearly
parallel with the general plane of the valves, its fissure broad,
partly closed by the arching deltidium ; hinge teeth well devel-
oped, compressed, and minutely striated ; interior with impres-
sions of cardinal muscles subovate, diverging, attenuate above ;
adductor muscular scars small, narrow-subelliptical; mesial
ridge prominent near the beak, much lower, and nearly extend-
ing forward to the central region ; most of the interior occupied
by granules, which are largest and most crowded on a narrow
space around and near the front and lateral margins; but
around the immediate margin they are much smaller, and ar-
ranged in distinct radiating rows. Dorsal or smaller valve
following nearly the curve of the other, the beak and central
regions being concave, and the ears flat; area well developed,
but narrower than in the other valve; bifid cardinal process
and mesial prominence, nearly or quite closing the fissure of
the other valve. From the base of this process there are ex-
tending, on the inner side of the valve five radiating ridges,
two of which pass obliquely outward, forming the inner mar-
gins of the dental sockets, while a third mesial one extends at
right angles to the hinge, a little more than half way to the
front; the other two are much shorter, oblique, and occupy in-
termediate positions between the middle and two latter ones;
granules of the interior as in the other valve. Surface of both
valves ornamented with a few subimbricating marks of growth,
crossed by very fine, obscure, regularly and closely arranged
radiating strie, of which about 150 can be counted around the
free border of a large individual, where about eight or nine of
them may be counted in the space of one line.’’
BEEDE. | Carboniferous Invertebrates. eo
Measurements: Length, 15mm.; width,28mm.; convexity,
5mm.
Range and distribution : Common throughout the Coal Meas-
ures; from Kansas City to Topeka and Manhattan, Grand Sum-
mit, etc. .
There are two specimens from Grand Summit, Kan., which
show a remarkable variation from the average specimens of this
species. In these the ears are elevated, distinctly and deeply
notched, the lateral margins curving inward, then, making an
acute angle, are directed outward and backward to the hinge
tip. In all other respects, so far as can be determined from the
exterior appearance, they are exactly like the above species.
Two other specimens from eastern Kansas also show this re-
markable characteristic, though in a less pronounced manner.
This species differs from the preceding in possessing radiat-
ing strie on the surface of both valves and a thicker shell; it
is more broad, and the ridges on either side of the mesial sinus
are less prominent.
Chonetes mesolobus. Plate IX, figs. 3, 3b.
Chonetes mesolobus Norwood and Pratten, Jour. Acad. Nat. Sci. Phil., 111,
p. 27, pl. 1, f. 7, (1854); White, Wheeler’s Geog. Surv. West 100 Mer.,
iv, p. 193, pl.. 1x, f.) 7, (1875); . ete.
Shell small, broader than long, moderately convex, trans-
versely subelliptical or subquadrate in outline; hinge line a
little more than equal to the greatest width of the shell. Mesial
sinus extending from near the beak, broadening and deepening
to the anterior margin, bifurcating near its origin and enclosing
a mesial fold. Anterior margin nearly straight or slightly sinu-
ate; antero-lateral margin sharply rounded, then passing nearly
in a slightly convex line directly backwards to the hinge line,
making a slight outward turn before joining it. Cardinal area
narrow, foramen broad and shallow, nearly filled by the small,
trifid process of the other valve. Surface ornamented with
fine radiating striw, rather coarsely and sparsely punctate.
Four or five small spines on eather side of the beak point
obliquely backward from the hinge line. The markings on the
interior of the ventral valve are rather similar to the preceding.
~]
2 University Geological Survey of Kansas.
The mesial septum is apparently short and thick. Two small,
sharp elevations mark the location of the double sinus on the
exterior. The adductor scars are located close beneath the
beak, oblique and ovate-elliptical in outline, and rather deeply
impressed. The pustules are arranged as in C. granulifer, ex-
cept that they are not always arranged in radiating rows
around the periphery, though they are in some specimens.
The interior of the dorsal valve is different from the rest of our
species. The sinus dividing the extremity of the deltidial proc-
ess splits into two, making the end of the process appear trifid.
The radiating ridges are nearly as in the preceding species, ex-
cept that the middle pair are recurved, forward and inward,
enclosing a depression resembling a muscular marking. The
mesial septum extends fully half the length of the shell, ending
in a slight enlargement. On either half of the valve, corre-
sponding to the folds of the other valve, are rather crescent-
shaped brachial areas, the outer edges of which are beset with
large, rather long, pustules. Pustules of the remainder of the
surface as in the other valve. In the anterior half of the shell
is a deep double sinus, enclosing a mesial fold, corresponding
to the sinus and fold of the ventral valve.
Range and distribution: Lower Coal Measures; Fort Scott
and Bronson, Bourbon county.
The features which distinguish this species from the re-
mainder herein described are the mesial fold in the ventral
valve and the short hinge line. Its range is restricted to the
lower part of the Coal Measures.
Chonetes verneuilianus. Plate IX, figs. 4—4c.
Chonetes verneuilianus Norwood and Pratten, Jour. Acad. Nat. Sci. Phil.,
111, p. 26, pl. u, f. 6, (1854); Newberry, Ives’ Rep. Col. Riv. of West, p.
128, (1861); Meek, Fin. Rep. U. S. Geol. Surv. Neb., p. 170, pl. 1, f. 10,
(1872); Hall, 2d Rep. N. Y. St. Geol. pl. xiv, ff. 20, 21, (1883); White,
13th Ann. Rep. St. Geol. Ind., p. 128, pl. xxv, ff. 7, 8, (1884); Hall and
Clarke, Pal...N.-Y3,. vi11y pti playin iy 20,021:
Meek’s description (in part): ‘‘Shell rather small, varying
from subsemicircular to suboblong; hinge line more or less
extended beyond the breadth of the valves at any other point ;
sometimes greatly produced: Ventral valve very convex, with
Se ee Se ee re ee ee =
BEEDE. | Carboniferous Invertebrates. 73
a deep rounded mesial sinus, starting near the beak and deepen-
ing and widening rapidly to the anterior margin, to which it
imparts a distinctly sinuous outline, thus dividing the gibbous
‘part of the valve into two prominent rounded lobes or diverg-
ing ridges, separate from each other by a broad rounded de-
pression; ears more or less angular, sometimes extended and
acutely pointed, slightly arching, and a little reflexed; beak
rather prominent and recurved; area moderately developed
and common to both valves, but widest in the ventral; fora-
* men wide; cardinal margin provided with four oblique spines
on each side of the beak. Dorsal valve following rather nearly
the curve of the other, and provided with a mesial ridge corre-
sponding to the sinus of the other valve.’’ The interior of the
ventral valve possesses a short, sharp mesial septum extending
beneath the umbo; each side of the septum, close under the
beak, there is an oval muscular depression, the broader end ex-
tending obliquely forward; near the posterior end is a small
elliptical scar. Immediately in front of the scars the shell is
smooth. Around the visceral cavity are large pustules, while
exterior to these, on the periphery of the shell, are smaller ones
arranged in radiating order. There is a small, distinct tooth
on each side of the foramen. The deltidial process of the dorsal
valve is reenforced by five radiating ridges on the interior of the
valve, the first pair of which are nearly parallel to the hinge
line, but pointing a little forward and disappearing before reach-
ing the margin. Immediately in front of the process, which is
somewhat eleyated and broad, is a small depression or cavity.
The second pair of ridges extend obliquely forward from in
front of this cavity. The mesial septum reaches about half
way to the front of the shell. On either side of it is the semi-
circular brachial area. Pustules on the surface of the valve
arranged as in the other valve. ‘‘ Surface [exterior] of each
valve ornamented with about 100 fine, bifurcating, radiating
strie, and sometimes near the front, by a few marks of growth.”’
Length, 9 mm.; width, 15 mm.; convexity, 4 mm.
Range and distribution: Lower and Upper Coal Measures ;
6—Vvi
74 University Geological Survey of Kansas.
Bronson, Bourbon county, Kansas City, Buffalo Mound, Wa-
baunsee county.
This species is easily distinguished by its deep mesial sinus
and strong bilobate appearance, together with a long hinge.
PRODUCTUS.
Sowerby, Min. Con., I, p. 153, (1814).
de Koninck, Recher. Anim. Foss., il p. 11, (1847).
Hall, 29th Rep. N. Y. St. Cab. Nat. Hist., p. 245 (1867) : Pal. N. Y., IV, p. 146, (1867)
Hall and Clarke, Pal. N. Y., VIII, pt. I, p. 321, (1892) ; 11th Ann. Rep. N. Y Y. St. Geol., p. 297, (1894).
This genus has been divided into groups, as follows
Group I. Lineatt Waagen. ‘‘ Surface covered with fine radi-
ating coste which are rarely spinous and are not crossed by
concentric plications or wrinkles. ‘The shells are greatly pro-
duced and sometimes the anterior margins of the shells are
modified by the development of a fold or sinus. The shells
were very fragile and have usually been subject to much dis-
tortion in fossilization.’’*
Group IL. Irregulares Waagen. ‘‘ Klongate shells very nar-
row at the beak, mytiliform in outline; mode of growth quite
irregular. Surface as in the Lineatv; spines grouped almost
wholly about the cardinal line.’’ * .
Group III. Semireticulatt de Verneuil. ‘‘The longitudinal
ribs are sparsely spinous; surface of visceral disc covered with
concentric wrinkles.’’ *
Group IV. Spinosi dé Verneuil. ‘‘ Surface strongly tubercu-
lose or spinous; not reticulated.’’ *
Group V. Fimbriati de Koninck. ‘‘Surface without radiate
strie or ribs; covered with concentric ridges or plications,
bearing rows of small thickly set spines.’’ *
Group VI. Horridi de Verneuil. ‘‘Surface without concen-
tric or radiating plications; pedicle valve with a deep sinus.”’ *
Group VII. Mesolobi de Koninck. ‘‘ Surface without radiat-
ing or concentric plications except a prominent median rib.’’ *
All of our Kansas species come in groups I, III, IV, and V.
I.—Lineati: P. cora and its varieties.
Ill.— Semireticulati: P. semireticulatus, costatus, pertenuis, longispinus.
IV.—Spinosi: P. nebrascensis.
V.—Fimbriati: P. punctatus, symmetricus.
4, Hall and Clarke, Pal. N. Y., VIII, I, p. 326.
BEEDE. | Carboniferous Invertebrates. 75
Concerning the synonymy of the American species of Productus
there is considerable difference of opinion. As far as our Kan-
sas forms are concerned, they have not heretofore been collected
and preserved in a manner that indicates the exact range of the
different forms, or, indeed, the exact location of the specimens
themselves ; in fact, if we are to judge from the localities given
in paleontological papers, there has been very little if any ac-
curate systematic collecting done in the Coal Measures west of
the Mississippi river, except that of Meek in southeastern Ne-
braska. As a natural result, the range of the different forms
is not known, or even the degrees of variation in the same hori-
zon. Under these circumstances, it seems best to be quite con-
servative in the number of species recognized until further and
better work is done; however, I am of the opinion that some
of the forms here referred to a single species will, on more care-
ful study of range restrictions, be separated. This is especially
true of P. cora, though the data at hand will hardly permit of
it now.
Productus cora. Plate XI, figs. 1-lf.
Productus cora @Orbigny, Voy. dans |’Amér. Mé¢r. Pal., p. 55, pl. v, ff.
8-10, (1842); Owen, Geol. Rep. Wis., Iowa, and Minn., pp. 103-106, pl.
v, f. 2, (1852); White, 13th Rep. St. Geol. Ind., p. 126, pl. xxv1, ff. 1-3,
(1884); ete.
Productus lyelli de Verneuil, Lyell’s Travels in North America, 11, p. 221,
(1845); Dawson, Acad. Geol., p. 219, f. g, (1855).
Productus semireticulatus Hall, Stanberry’s Expl. and Surv. Valley Gt.
Salt Lake, Utah, p. 411, pl. 11, ff. 3, 5.
Productus prattenianus Norwood, Jour. Acad. Nat. Sci. Phil., 11, p. 17,
f. 10, (1854); Meek, Fin. Rep. U.S. Geol. Surv. Neb., p. 163, pl. 11, f. 5,
pl. v, f. 13, pl. vir, f. 10, (1872); etc.
Productus flemmingi Geinitz (non de Verneuil), Carb. u. Dyas in Neb.,
p. 53, pl. iv, f. 5, (1866); ete.
Productus calhounianus Geinitz, ibid. (non Swallow).
Meek’s description of Productus prattenianus (in part): ‘‘ Shell
attaining a medium to large size; breadth generally exceeding
the length, especially when the ears are entire; cardinal mar-
gin usually somewhat longer than the transverse diameter of
the valves at any point farther forward; anterior and antero-
lateral outline regularly rounded. Ventral valve distinctly and
rather evenly convex, with or without a shallow mesial sinus ;
76 University Geological Survey of Kansas.
umbonal region gibbous; beak incurved, but scarcely passing
the hinge margin ; ears large, rather compressed, and provided
with a few large, strongly defined concentric folds, which as-
cend a little upon the sloping sides of the umbo and extend
more or less along the postero-lateral margins, but never cross
the shell; surface ornamented with rather small, regular,
rounded cost or stri#, and armed with stout, erect, long
spines, those along the hinge margin are more frequent, larger,
directed backward, with an inward curve. Some specimens
seem to be nearly destitute of spines. Dorsal valve concave,
sometimes a little flattened in the visceral region, and following
the curve of the other valve around the front and anterior
lateral margins ; ears with folds as in the other valve, and each
separated from the concave central region by an oblique ridge
or prominence; surface without spines, but with radiating
strize, as in the ventral valve, and usually crossed by very ob-
scure concentric wrinkles and a few imbricating marks of
growth, particularly near the front and sides; cardinal process
small but strongly prominent and bifid, while from its base a
slender mesial ridge extends forward to, or a little beyond, the
middle.’’ The internal markings of this valve are obscure. Pro-
boscis well developed, bifid, and well produced into the beak
of the other valve. Lateral ridges small, straight or curved
backward, soon disappearing; adductor scars dim, placed well
to the posterior, nearly semicircular in outline; brachial mark-
ings indistinct, enlarging and bending to the front at the ex-
tremities. A cast of the interior of the pedicle valve shows a
narrow, short beak, with a small depression on either side,
faint mesial ridge extending across the central portion of the
visceral area; adductor scars long and narrow, placed close be-
side the ridge; exterior to these are the linear markings of the
diductors. Anterior marked by fine pittings. Length of fair-
sized specimen, 50 mm.; width, 70mm.
Range and distribution: Upper Coal Measures; Kansas City,
Eudora, Lawrence, Lecompton, Topeka, Geary county, Melvern,
Osage county.
The size of the costz in this species is variable, those with
BEEDE. | Carboniferous Invertebrates. 77
the larger ones possessing the larger and more numerous spines,
while those with fine costie seem to be nearly destitute of spines.
The forms of this species with the front margin emarginate
are restricted, or at least seem to be, to the base of the Upper
Coal Measures, and are for this reason worthy of varietal dis-
tinction. There is also a very wide form that is very much less
gibbous than the ordinary form, which is likewise restricted to
the same rocks as the preceding, and will probably prove varie-
tally distinct from the species. Waagen” suggests that Pro-
ductus cora should be divided into two series of forms, one with,
and the other without a mesial sinus in the visceral area. This
will hardly hold for our western forms, for those of the form
described by Swallow as P. americanus and those without the
fold characteristic of that form both possess the sinus. The
form that is the most common in the Kansas Coal Measures is
that described by Norwood as P. prattenianus, found abundantly
at the Nebraska City, Neb., locality of Meek, and throughout
the Kansas Coal Measures.
Productus cora americanus. Plate XI, fig. 2.
Productus americanus Swallow, Trans. St. L. Acad. Sci., 11, p. 91, (1863).
XVII-A, ff. 22, 23, (1893).
Productus cequicostatus Hall (non Shumard), Pal. N. Y., vii, pt. 1, pl.
This shell differs from the preceding species in possessing a
fold in the anterior portion of the shell, which is less developed
anteriorly than the sides, forming a deep sinus in the anterior
margin. In the most distinct forms of the variety the shell
is sometimes less gibbous and more alate than is true of the
species. |
Range and distribution: Upper and Lower Coal Measures ;
Kansas City, Eudora, Anderson county. Seems to be confined
to the base of the Upper Coal Measures and to the Lower Coal
Measures.
5. Pal. Indica, Salt Range Foss., 676.
78 University Geological Survey of Kansas.
. Productus semireticulatus. Plate X, figs. 2-2d; text fig. 2, f.
Anomites semireticulatus Martin, Petref. Derb., p. 7, pl. xxx11, ff. 1, 2,
pl. xxx, f. 4, (1809).
Productus inea d’ Orbigny, Voy. dans |’ Amér. Mérid. Pal., p. 51, pl. tv,
ff. 1-3, (1842); Derby, Bull. Mus. Comp. Zool., 111, p. 280, (1876).
Productus semireticulatus Norwood and Pratten, Jour. Acad. Nat. Sci.
Phil., 111, p. 11, (1854); White, 13th Ann. Rep. St. Geol. Ind., p. 125,
-pl. xxtv, ff. 1-3, (1884); ete.
Productus setigerus Hall, Geol. Surv. Iowa, 1, pt. 11, p. 638, pl. x1x, f. 3.
Productus setigerus var. keokuk Hall, ibid.
Productus martint A. Winchell, Proc. Acad. Nat. Sci. Phil., p. 4, (1863).
Productus magnus Halland Clarke, Pal. N. Y., vii, pt. 1, pl. xvi1-a, f. 15,
(1892).
Shell large, very convex, broader than long ( measured from
the hinge to the front) ; hinge line straight, equal to the great-
est width of the shell; cardinal area very narrow but distinct ;
beak prominent, appressed, barely arching around the hinge
line. Pedicle valve much inflated, greatly prolonged anteriorly
in old individuals. Beak broadening rapidly from its point at
the hinge toward the anterior; ears nearly flat, well defined,
separated from inflated portion of the shell by a gentle curve ;
mesial sinus beginning well toward the beak and continuing
to the anterior margin. Surface marked by radiating coste,
and the visceral area is also marked by more or less regular,
concentric cost, giving the shell a semireticulated appearance.
Sometimes both sets of strie extend over the ears, the concen-
tric taking the form of wrinkles, which, together with the bases
of stout tubular spines, make the ears, appear quite rough.
Spines sparsely and irregularly distributed over the costi of
the shell but thickest on the ears; radiating cost increasing
by bifurcation and insertion. The interior of the valve is.
marked on the ears, anterior and lateral border by a pitted sur-
face. Adductor scars are elongate-elliptical, arborescent, hav-
ing their anterior extremities raised, situated centrally in the
shell. Longitudinal linear lines or folds are present on the
outside of these impressions. ‘The dorsal or brachial valve is
strongly concave, visceral region nearly flat, in front of which
it curves abruptly downward. A slight indistinct mesial fold
is present, corresponding to the sinus of the other valve. Kars
and surface markings same as in the other valve, except that
=
o _ ae — ° "
ls Le ae
BEEDE. | Carboniferous Invertebrates. 79
there are no spines.’ Beak extending back of the hinge, produc-
ing a concavity below. Proboscis not visible from exterior of
complete specimens, only a minute projection seen fitting into
a small notch in the hinge of the othervalve. The entire inner
surface of the valve coarsely punctate, except the portion oc-
cupied by the brachial and adductor markings. Proboscis pro-
jecting, obovate in outline, trifid, at the base of which there
are three radiating ridges; two high lateral ones, which roll
slightly backward, extend nearly to the ears, where they fade
out; the third ridge extends directly forward, smaller and
sharper than the others, to the anterior portion of the flat sur-
face, where it ends abruptly. Situated about one-third, or less,
the distance from the posterior to the front of the flat surface
and close to the mesial ridge are the arborescent adductor
markings, triangular to elliptical in outline. The brachial
markings extend nearly directly outward from the anterior end
of the adductors, then bending forward and expanding, ap-
proaching very close to the antero-lateral extremity of the flat
surface of the valve. The front inner portion of these mark-
ings extends forward to the anterior end of the mesial ridge,
the posterior portion connecting with the front of the adductor
scar. In some specimens the brachial area is finely crinkled,
as are some other parts of the surface back of the muscular
scars. There are sometimes short, sharp pustules on the front
curve of the valve. Length, hinge to front, 40 mm.; width,
58 mm.; convexity, 40 mm.
Range and distribution: Upper Coal Measures; Kansas City,
Lawrence, Lecompton, Topeka, Alma, Marysville, etc. Com-
mon throughout the Coal Measures of the United States and
the world.
Productus costatus. Plate IX, fig. 8; plate X, figs. 1-1le; plate XI, fig. 4.
Productus costatus Sowerby ? Min. Con., v1, p. 115, pl. pix, f. 1, (1827).
Productus costatus de Koninck, Recher. Anim. Foss., pt. 1, p. 92, pl. viii,
f. 3, pl. x, f. 3, pl. vi, f. 3, (1847); Meek, Fin. Rep. U.S. Geol. Surv.
Neb., p. 159, pl. v1, f. 6, (1852); Hall and Clarke, Pal. N. Y., vi111, pt. 1,
pl. xx, ff. 8-13, (1892); ete.
Productus portlockianus Norwood and Pratten, Jour. Acad. Nat. Sci.
Phil., 111, p. 15, pl. 1, f. 9, ( 1854).
Productus viminalis White, Proc. Boston Soc. Nat. Hist., 1x, p. 29, ( 1862).
80: University Geological Survey of Kansas.
Meek’s description (in part): “Shell of medium size, wider
than long, very convex; hinge margins about equaling the
greatest breadth of the valves. Pedicle valve exceedingly gib-
bous, and very strongly incurved, with a deep rounded sinus
extending from near the beak to the front, to which it imparts
a sinuate outline; umbo prominent and strongly incurved, so
as to pass somewhat within the hinge margin; ears well de-
fined, arched and rather distinct from the abrupt swell of the
umbo, from which they are sometimes separated by a small
ridge or fold. Brachial valve flattened in the visceral region,
and more or less abruptly curved or geniculated toward the
front and anterior lateral margins, the former of which usually
shows a small mesial ridge. Surface of both valves ornamented
with distinct, rather unequal, depressed and rounding radiat-
ing costee, which sometimes bifurcate, or, in other instances,
two or more of them coalesce in front of the visceral portion,
to form a larger one; crossing all of these, on the visceral re-
gion, are numerous, well-defined, concentric wrinkles, produc-
ing a distinct reticulated appearance, while the whole surface
of the ventral valve is sometimes provided with a few scatter-
ing, rather stout, erect spines, somewhat regularly arranged in
quincunx. Sometimes nearly all the spines, excepting those on
the lateral regions, apparently wanting.’’ The internal mark-
ings of the ventral valve consist of long, narrow, arborescent
adductor scars mesially situated along the vaulted part of the
shell, with, exterior to these, the long linear folds, or line-like
diductor scars. The beak, as shown in a cast, is quite pointed,
possessing a depression on either side running obliquely back-
ward, near the posterior end of which there is an elevation, in
front of which the pit is deeper than back of it. Above this de-
pression is a rounded ridge on each side of the mesial sinus.
The proboscis of the dorsal valve extends beyond the hinge
line, and is backed by three radiating ridges, one of which ex-
tends directly forward to the front of the flattened area,
rounded, not prominent; one extends antero-laterally on each
side of the mesian ridge, running nearly outward until the
edge is approached, when it curves forward, joining the border
Le aah oan ee
BEEDE. | Carboniferous Invertebrates. 81
of the flattened area, and fades out. Adductor scars moderately
small, elliptical in outline, placed longitudinally and close to the
median ridge, a little anterior to the lateral ridges. Brachial
markings not prominent and generally indistinct, extending
outward from the anterior end of the adductor scars, curving
forward and ending near the antero-lateral portion of the
flattened area. Other internal markings unknown. Length,
20 mm.; width, 35 mm.; convexity, 33 mm.
Range and distribution: Upper Coal Measures; Kansas
City, Turner, Wyandotte county, Lawrence, Topeka, Wabaun-
see county. Common throughout the Coal Measures and
lower portion of the Permian.
This species differs from the previous in being smaller, hay-
ing a more distinct mesial sinus, ears separated from the body
of the shell by a ridge which generally bears spines and un-
equal cost on the anterior slope.
The markings of this shell vary considerably in different in-
dividuals. They are not so distinct as those of a Russian ex-
ample in the collection, and the shell is smaller; though the
ridge separating the ear from the shell is present and spinous,
it is not nearly so pronounced as in the Russian specimen.
Productus longispinus? Plate IX, figs. 9-9d.
Productus longispinus Sowerby? Min. Con., 1, p. 154, pl. uxvin, f. 1,
(1814).
Productus longispinus Salter, Quart. Jour. Geol. Soc. Lond., xvii, p. 64,
pl. 1v, f. 2, (1861); Meek, Fin. Rep. U.S. Geol. Surv. Neb., p. 161, pl.
vi, f. 7, pl. vir, f. 6, (1872); ete.
Productus flemmingi Roemer (non deKoninck), Kreidebildung Texas,
p. 89, pl. x1, f. 8, (1852); etc.
Productus splendens Norwood and Pratten, Jour. Acad. Nat. Sci. Phil.,
111, p. 11, pl. 1, f. 5, (1854); ete.
Productus wabashensis Norwood and Pratten, ibid.
Productus orbignyanus Geinitz (?non de Koninck ), Carb. u. Dyasin Neb.,
p. 56, pl. rv, ff. 8-11, (1866).
Produ-tus horridus Geinitz, ibid., f. 7.
Meek’s description (in part): ‘‘ Shell small, thin, wider than
long; hinge line generally longer than the transverse diameter
of the valves at any point farther forward, and terminating in
more or less distinct, rather vaulted, and often reflexed ears;
anterior and anterior-lateral outlines approaching a semicircu-
82 University Geological Survey of Kansas.
lar curve, but the middle of the front is generally rather dis-
tinctly sinuous. Ventral valve gibbous, the greatest convexity
being usually behind the middle, and the curve to the beak
more rapid than to the front, provided with rather deep mesial
sinus; posterior lateral slopes descending nearly vertically to
the ears; umbonal region moderately prominent, and usually
projecting rather distinctly beyond the hinge, as seen in look-
ing down upon the shell when lying with the dorsal valve be-
neath ; beak small, strongly curved, but scarcely passing beyond
the cardinal margin; surface ornamented with generally rather
obscure, somewhat variable radiating costze, which are often
obsolete in the umbonal region, or, in some examples, over much
of the valve farther forward, in other specimens quite distinct
to the beak, sometimes bifurcating, and in other instances
coalescing to form larger, faintly defined ribs in front; fine, in-
distinct marks of growth are also sometimes seen, and occa-
sionally very obscure traces of small, concentric wrinkles may
be observed near the beak; spines stout, erect, long, scattering, |
and arranged in quincunx. Ventral valve distinctly concave,
or following nearly the curve of the other, and provided with a
small mesial ridge corresponding with the sinus of the latter ;
surface marked as in the other valve, but apparently always
without spines.’’ Proboscis short, trilobate; two ridges extend
laterally from its base nearly parallel to the hinge line, until
the border of the visceral region is reached, where they bend
abruptly forward, and in very old individuals form a ridge
entirely around the visceral and brachial cavity, but in many
adult specimens it fades out before it reaches the front side of
this cavity ; the lateral borders of the shell are often striated
from this ridge to the margin; mesial ridge faint, extending
nearly to the front of the viscero-brachial area. Adductor scars
small, not prominent, ovate, placed close beside the mesial
ridge, well to the back of the shell, directed obliquely for-
ward ; directly in front of these the brachial markings extend
toward the antero-lateral margin of the brachial area, near
which they enlarge; in front of these the surface of the valve
is marked by many or few (according to the age of the shell)
BEEDE. | | Carboniferous Invertebrates. ‘) 83
spinous projections; the remainder of the surface of the valve
issmooth. Length, 11mm. ; width, 16mm. ; convexity, 10 mm.
Range and distribution: Upper Coal Measures; Kansas City
and Turner, Wyandotte county, Eudora, Lawrence, Lecomwp-
ton, Topeka, Wabaunsee county. Common or abundant
throughout the Coal Measures of the state.
This species is easily distinguished from the preceding by its
smaller size, fainter surface markings, longer hinge, and in the
shell being more transverse in appearance, as well as having no
fold separating the ears from the umbo. There is also great
difference in the internal markings of the two, as can be seen
by referring to the plate.
This species is quite variable in the distinctness of its mark-
ings, spines, distinctness of the mesial sinus, and size. The
internal markings vary greatly with age, the very old speci-
mens presenting the marginal ridge surrounding the visceral
and brachial areas, which forms the generic characters of Mar-
ginifera; but I am inclined to think that characters that are
only developed in aged specimens are not of generic value in
the brachiopods.
Productus pertenttis. Plate IX, figs. 5-5c.
Productus cancrini Geinitz, Carb. u. Dyas in Neb., p. 54, tab. iv, ff. 6
a—d, (1866).
Productus pertenuis Meek, Fin. Rep. U. S. Geol. Surv. Neb., p. 164, pl.
1, ff. 14a—c, pl. vii, ff. 9a-d, (1872); Drake, Proc. Am. Phil. Soc.,
XXXVI, p. 405, pl. 1x, ff. 8-10, (1898).
Meek’s description: ‘‘ Shell small, very thin, truncato-hem-
ispherical; sides and front regularly rounded; hinge line
usually less than the greatest breadth of the valves. Ventral
valve without any traces of a mesial sinus, moderately gibbous,
the greatest convexity being slightly behind the middle, from
which point it rounds off in all directions, but most abruptly
toward the beak and ears, which latter are flattened and sub-
rectangular; beak small, slightly prominent, and but little in-
curved beyond the hinge line; surface with fine, regular,
radiating striw, crossed by small, rather distinct and regular,
concentric wrinkles, which latter are generally most strongly
84: University Geological Survey of Kansas.
defined on the ears; over the whole there are also regularly
arranged in quincunx, very slender spines, .20-.30 inch in
length, rising from the slight prominences or swellings of the
radiating strie. Dorsal valve distinctively concave, following
nearly the curvature of the other valve, its greatest convexity
being in the central region, while its ears are nearly flat; sur-
face with concentric wrinkles and radiating striz as in the
other valve, but apparently without spines, though a series of
rather distinct pits are arranged over it in the same order as in
the other valve.”’ Length, 8 mm.; width, 11 mm.; convexity,
5 mm. |
Range and distribution: Upper Coal Measures; Kansas City,
Eudora, Lawrence, Lecompton, Topeka.
This species differs from the preceding ones of the genus in
the absence of the mesial sinus, thin shell, very small size and
semicircular outline, as well as being much less gibbous. It is
1 ikely to be confounded with no other shell of the Coal Measures
of Kansas.
Productus nebrascensis. Plate IX, figs. 7-7f.
Productus nebrascensis Owen, Geol. Rep. Wis., Iowa, and Minn., p. 584,
pl. v, f. 4, (1852); Meek, Fin. Rep. U.S. Geol. Surv. Neb., p. 165, pl. 11,
f. 2, pl. iv, f. 6, pl. v, ff. lla—c, (1872); ete.
Productus rogersi Norwood and Pratten, Jour. Acad. Nat. Sci. Phil.,
2d ser., 111, p. 9, pl. 1, f. 3, (1854); ete.
Productus asperus McChesney, New Pal. Foss., p. 34, (1860); ete.
Productus wilberanus McChesney, ibid., p. 36; etc.
Stropholosia horrescens Geinitz (non Murchison, etc.), Carb. u. Dyas in
Neb., p. 49, (1866).
Meek’s description (in part): ‘‘ Shell of about medium size,
approaching subhemispherical; length most usually a little
less than the breadth ; hinge line nearly or quite equaling the
greatest transverse diameter; anterior outline nearly straight,
or a little sinuous near the middle, rounding into the lateral
margins, which are generally straight posteriorly, and ranging
at an angle of from 90 to about 100 degrees with the hinge;
ears nearly rectangular or a little rounded in outline at the
immediate extremities. Ventral valve rather convex, most
gibbous behind the middle, thence rounding regularly to the
: ————
ee eT
BEEDE. | Carboniferous Invertebrates. 85
front and more abruptly to the beak, generally with a moder-
ately distinct mesial sinus; posterior lateral slopes descending
almost vertically to the ears; umbonal region gibbous, and with
strong incurved beak projecting beyond the hinge line. Dorsal
valve somewhat flattened in the visceral region, but most con-
cave near the beak and near the anterior lateral regions, the
concavity widening rapidly forward, so as to leave a kind of a
broad, obscure, oblique ridge between it and the flattened ears,
and another in the middle; anterior and lateral margins fol-
lowing the curvature of the other valve; cardinal process
prominent, bifid, and rather narrow; interior with mesial
ridge, narrow, well defined, extending forward beyond the
middle,’’ bifid, at its connection with the cardinal process en-
closing a moderately deep pit. The two lateral ridges are well
defined and parallel with the hinge, fading out before the ex-
tremity is reached. The adductor scars are either subsemicir-
cular or spatulate in ouiline, placed well to the posterior, with
the smaller end forward, ending in a small elevated lobe. The
brachial markings of this species are very obscure; they extend
from the anterior extremity of the adductor scars outward and
a little forward, rounding off near the antero-lateral edge of
the flattened area. The markings of the interior of the other
valve are rather indistinct. The adductor scars are long, slen-
der, subspatulate, with the larger end toward the beak, situated
in the middle of the visceral area; the two are separated in the
extreme posterior portion by a low mesial ridge; exterior to
these are the diductor scars, occupying a rather large space, in-
dicated by longitudinal folds or nearly parallel lines. ‘‘ Sur-
face of the ventral valve with more or less defined, rather broad
concentric undulations, and obscure strive of growth, over the
whole of which are arranged two sets of spines, connected at
their bases with short interrupted ribs or elongated tubercles.
One of these sets consists of small, short, appressed spines, and
the other of stout, more erect, long ones. Surface of dorsal
valve with small concentric ridges and strive, with many little
pits ; spines nearly or quite all small, short, and appressed.’’
86 University Geological Survey of Kansas.
Measurements: Length, 26 mm.; width, 30 mm.; convex-
ity, 16 mm. 7 |
Range and distribution : Upper Coal Measures; Kansas City,
Turner, Eudora, Lawrence, Lecompton, Topeka, Manhattan,
and Grand Summit. Common throughout the Upper Coal
Measures of the state.
This shell, when found with the shell and spines well pre-
served, looks so very different from specimens which are ordi-
narily found with the spines removed that one could hardly
imagine that they were the same. Those without the spines
seem to be complete specimens, with regular, concentric zones
of elongated tubercles, betraying very little indication that they
are the bases of detached spines.
This shell is very readily distinguished from the foregoing
species by its narrow, prominent beak and its peculiar surface
markings. The concentric zones of spines are found on none
of them.
Productus symmetricus. Plate IX, figs. 6-6b.
Productus symmetricus McChesney, New Pal. Foss., p. 35. (1860); Trans.
Chic. Acad. Sci., 1, p. 25, pl. 1, f. 9, (1868); Meek, Fin. Rep. U. S.
Geol. Surv. Neb., p. 167, pl. v, f. 6, pl. vim, f. 13, (1872); ete.
Meek’s description (in part): ‘‘Shell of medium size, sub-
orbicular, or a little wider than long; hinge line somewhat less
than the greatest breadth; sides rounding regularly to the
front, which is rather broadly rounded in outline; ventral
valve somewhat compressed, or only moderately convex, with- |
out any traces of a mesial sinus; ears compressed but not
abruptly separated from the swell of the umbo, obtusely an-
gular or a little rounded at the extremities; beak moderately
large, incurved, but not curving much within the hinge margin.
Dorsal valve rather evenly, and only moderately concave, car-
dinal process slender, prominent, curved [not always}, trifid,
the middle division being more prominent than the others, and
emarginate at its extremity, the emargination being caused by
a distinct mesial furrow that extends the entire length of the |
process [ana sometimes two-thirds the length of the mesial
septum].’’ Lateral ridges elevated, extending outward parallel
BEEDE. | Carboniferous Invertebrates. 87
to and a little in front of the hinge nearly to its extremity,
where it turns abruptly forward, forming a margin to the vis-
~ ceral area, and disappearing about the middle. Mesial ridge
distinct, broad posteriorly, narrowing and rising to the end,
which is two-thirds the distance to the anterior margin. Ad-
ductor scars distinct, the two nearly forming a circle or broad
ellipse, ending in an elevated anterior lobe. Scars placed well
to the posterior. Brachial markings very indistinct, beginning
at anterior end of adductor scars and extending outward, as
nearly as can be made out, similarly to those of the preceding
shell. ‘*‘ The remainder of the interior covered with pustules.
Surface of both valves ornamented by small, rather obscure,
more or less regular concentric wrinkles, and covered by numer-
ous small, short, rather appressed spines, which are larger on
the ventral valve where they are often connected with little,
sometimes elongated tubercles.’’ Measurements: Length, 40
mm.; width, 59 mm.; convexity, 25 mm.
Range and distribution: Upper Coal Measures; Kansas City,
Lawrence, Topeka.
This species can be distinguished from the preceding by the
less convex pedicle valve, which has no sinus, narrower con-
centric wrinkles, and the fact that it has but a single set of
spines, which are small and appressed. In some respects this
species is intermediate between the one preceding and the fol-
lowing, and is sometimes hard to distinguish from them with-
out good specimens.
Productus punctatus. Plate X, figs. 3-3e; plate XI, fig. 3.
Anomites punctatus Martin, Petref. Derb., pl. xxxvit, f. 6, (1809).
Productus punctatus? Morton, Amer. Jour. Sci., xxrx, p. 153, pl. xxv,
f. 38, (1836).
Productus punctatus Shumard, Marcy’s Rep. U. S. Expl. Red Riv.
Louis., p. 201, pl. 1, f. 5, pl. 11, f. 1, (1853); Meek, Fin. Rep. U. S. Geol.
Surv. Neb., p. 169, pl. 11, f. 6, pl. 1v, f. 5, (1872); etc.
Productus semipunctatus Shephard, Amer. Jour. Sci., xxxtv, f. 9, (1853).
Productus tubulospinus McChesney, New Pal. Foss., p. 37, (1860).
Meek’s description (in part) : ‘‘ Shell attaining a rather large
size, thin, varying from rotund-subquadrate to longitudinally
suboyate, being sometimes wider than long, and in other ex-
88 University Geological Survey of Kansas.
amples longer than wide, with all intermediate forms; hinge
always shorter than the greatest breadth of the valves; anterior
outline regularly rounded, or faintly sinuous in the middle.
Ventral valve more or less gibbous, with a moderately distinct
mesial sinus extending from near the beak to the front; beak
incuryed a little beyond the cardinal margin [considerably so] ;
ears rather compressed, but not distinctly defined from the swell
of the umbo. Dorsal valve moderately concave with a small
mesial elevation. Surface of both valves ornamented with
numerous rather regular concentric ridges, increasing in size
from the beaks toward the front, but becoming again smaller
and more crowded in adult shells near the margin; in the
ventral valve these ridges are a little prominent at the margin,
separated from each other by smoother spaces, and support
numerous small appressed spines, those of the upper row of
which are larger and less crowded than the others; on internal
casts, or partly exfoliated specimens, the spines are represented
by small tubercles ; surface of the dorsal valve as in the other,
excepting that the ridges are represented by little furrows.’’
Interior of the pedicle value with a very small, thin, nearly
obsolete mesial septum ; adductor scars long, narrow, larger at
posterior extremity, situated well to the back part of the shell.
Diductors occupying a large space outside of the adductors, in-
dicated by longitudinal folding ridges or parallel lines. In
the brachial valve, the cardinal process is long, arched, bifid,
slightly enlarged at the extremity; mesial ridge prominent,
extending three-fifths the distance to the front margin, and is
slightly thickened at the end. Lateral ridges strong, situated
just in front of the hinge and parallel to it, but disappearing
before the edge of the shell is reached. Adductor scars promi-
nent, situated well to the front, varying in outline from nar-
rowly elliptical to nearly ovate, with heavy, narrow lobe at the
anterior end. Traces of brachial markings very hard to obtain
and nearly obsolete. They extend outward from the ante-
rior end of the adductor scars and make a broad, subelliptical
loop forward to a point which is back of the end of the mesial
ridge. Measurements: Moderately large, perfect specimen:
BReEDE. | Carboniferous Invertebrates. 89
Length, 58 mm. ; width, 75 mm.; convexity, 35mm. A smaller
specimen: Length, 36 mm.; width, 36 mm.; length, 36 mm. ;
convexity, 23 mm.
Range and distribution: Upper Coal Measures; Kansas City,
Turner, Lawrence, Lecompton, Topeka, Moline.
This species can be distinguished from the previous by its
mesial sinus and corresponding fold of the dorsal valve, more
recurving beak, longer, more arched cardinal process, more
slender and elongate adductor markings on dorsal valve, and
also by the lateral ridges, which do not turn forward.
There seem to be two forms of this species in Kansas:
One, a large, rather depressed shell, occurring in the lower part
of the Upper Coal Measures at Kansas City; and the other, the
more common in the upper part of the stage, more narrow, gib-
bous and smaller. They may possibly prove to be varietally
distinct, occupying different horizons.
AULACORHYNCHUS.
Dittmar, Verhand. Kais. Min. Gesellsch. St. Petersburg, 2d ser., VIII, p. Q, pl. I, ff. 1-13, (1871).
Hall and Clarke, Pal. N. Y., VIII, pt. II, p. 311, (1893).
Aulacorhynchus millipunctatus. Plate XI, fig. 6; plate XII, figs. 1, 1b.
Chonetes?? millipunctata Meek and Worthen, Proc. Acad. Nat. Sci.
Phil., p. 35, 1870; Geol. Surv. Ill., v, p. 566, pl. xxv, f. 3, (1873).
Tsogramma millipunctata Meek and Worthen, ibid., p. 568.
Aulacorhynchus millipunctatus Hall and Clarke, Pal. N. Y., vii, pt. 11,
p. 312, pl. txxxrit, ff. 14, 15, (1893).
Meek and Worthen’s description: ‘‘ Shell attaining a large
size, very thin, transversely subsemicircular, or more than
twice as wide as long, with lateral extremities rounded. Dor-
sal valve nearly flat, or but slightly and evenly concave; hinge
line a little less than the greatest transverse diameter; car-
dinal process rather stout, with an obscure linear ridge (or
sulcus) extending forward from its base nearly to the front;
cardinal edge slightly thickened within so as to form a faintly
defined ridge extending about half way from the cardinal proc-
ess toward each lateral margin, but apparently without any
sockets for the reception of teeth in the other yalye; muscular
and other internal markings unknown; surface ornamented.
7—vi
90 University Geological Survey of Kansas.
by numerous slender, exceedingly regular, closely arranged
concentric lines, extending parallel with each other and the
front and lateral margins. Ventral valve unknown. Length
of medium-sized specimen, 1.3 inches; breadth, 2.95 inches.’’
Range and distribution: Upper Coal Measures; Kansas City,
Lawrence. :
Two small specimens very similar in form and markings to
the one above described were collected by the writer at Topeka.
They were only about an inch in width and only the cast of the
shells remain. They are probably specifically different, but
too poorly preserved to admit of description. There is a very
large form in the University collection which is quite convex
and seems to possess an extra number of ‘‘platforms’’ from
the one figured by Hall and Clarke, and may belong to a dif-
ferent species. It is figured in plate XII, figure 1b.
Rhipidomella pecosi.
Orthis pecosi Marcou, Geol. N. Amer., D. 48, pl. vi, f. 14, (1858); ete.
Orthis carbonaria Swallow, Trans. St. L. Acad. Sci., 1, p. 215, (1858);
Meek, Fin. Rep. U.S. Geol. Surv. Neb., p. 131, pl. 1, f. 8, (1872); ete.
Meek’s description: ‘‘Shell small, suborbicular, slightly
wider than long, moderately convex in adult specimens ; lateral
margins rounded, or, in some examples, faintly straightened
posteriorly ; front more broadly rounded, but usually very
slightly sinuous in the middle; valves nearly equally convex; —
hinge line very short, or only equaling about half the breadth
of the valves. Ventral valve usually most convex in the um-
bonal region, sometimes a little flattened anteriorly, so as to
give the shell slightly the form unusually called ‘resupinate,’
though in gibbous specimens this character is nearly obsolete ;
beak moderately prominent, rather pointed and arched; area
small, well defined, and arching with the beak; foramen nar-
row. Dorsal valve usually most convex between the middle
and the beak, which is small, and nearly as prominent and
arched as that of the other valve, generally with a shallow
sinus extending from the middle to the front; area well de-
veloped but smaller than in the other valve, arched, and divided
BEEDE. | Carboniferous Invertebrates. 91
by a proportionally shorter foramen. Surface of both valves
ornamented with concentric marks of growth and fine radiat-
ing crowded striz, which increase mainly by intercalation, and,
as in many other species of the genus, show occasional perfora-
tions toward the front, apparently left by the removal of very
small tubular spines.’’ ‘‘ Length of a well-developed gibbous
specimen, rather above medium size: .38 inch; breadth, .43.
inch ; convexity, .27 inch.’’
Range and distribution: Upper and Lower Coal Measures ;
Fort Scott, Kansas City, Eudora, Lawrence, Lecompton.
ENTELETES.
de Waldheim, Oryct. Gouv. Moscou, p. 193, tab. XX VI, ff. 6, 7.
Waagen, Pal. Indica, ser. xiii, I, p. 550, (1884) ; ete.
Enteletes hemiplicata. Plate XII, figs. 6, 6b.
Spirifer hemiplicata Hall, Stansbury’s Expl. Gt. Salt Lake, p. 409, pl.
Iv, f. 3, (1852).
Rhynchonella angulata Geinitz (non Linné), Carb. u. Dyas in Neb., p. 37,.
pl. rir, ff. 1-4, (1866).
Syntriclasma hemiplicata Meek and Worthen, Geol. Surv. Ill., 11, p. 323,
f. 36, p. 324, f. 37, (1866); Meek, Fin. Rep. U.S. Geol. Surv. Neb., p. 177,
pl. vi, f. 1, pl. vi11, f. 12, (1872); etc.
Meek’s description: ‘‘Shell in young examples only moder-
ately convex, and having all the external appearances of a true
Orthis; in adult specimens, often globose, or even more con-
vex than long or wide. Hinge line very short, or not more
than one-third the greatest breadth of the valves, and, owing to
the gibbosity of the shell, imparting little or no angularity to
the outline of the lateral slopes. Dorsal valve more convex
than the other, and very strongly arched, particularly in ma-
ture shells; umbonal region gibbous, and often, in adult ex-
amples, projecting somewhat beyond the beak of the other
valve; beak strongly incurved, so as to bring its apex under
the beak, and nearly against the area of the other valve; area
rather narrow, and distinctly incurved with the beak. Ventral
valve convex, beak moderately prominent, and arched or more
or less incurved; area triangular, small, about one-third as
high as wide, and moderately well defined ; its triangular fora-
men scarcely as wide as high. Surface of both valves orna-
mented with rather fine, regular, crowded, radiating strie, and
92 University Geological Survey of Kansas.
a few very large, rounded or more or less angular, radiating
plications, which latter are never formed on the umbones, but
occupy the anterior half, and become more prominent toward
the front, where they often terminate in deeply interlocking
angular marginal projections. Of these plications there are two,
rarely three, on each side of the larger and more prominent one
forming the mesial fold on the dorsal valve, while on the ven-
tral valve there are three, rarely four, on each side of the mesial
sinus; a few zigzag marks of growth also traverse the anterior
and lateral margins of the valves, parallel to their deeply
notched edges.’’ The measurements of three specimens are
here given — one rather large, one medium, and one young speci-
men: Length, 23 mm., 16 mm., 8 mm.;: width, 29 mm, 28
mm., 9 mm.; convexity, 32 mm., 20 mm., 5 mm-
Range and distribution: Upper Coal Measures; Kansas City,
Independence, lola, Edwardsville, Eudora, Lawrence, Lecomp-
ton, Topeka.
Besides the variation in size and age, this shell often pre-
sents very much the appearance of a Pugnaz, which is caused
by the mesial plication being divided into two, and the presence
of a small fold to correspond in the mesial furrow of the other
valve. This gives the brachial valve quite a large mesial fold,
and a correspondingly large sinus in the pedicle valve. This is
hardly noticable in the majority of specimens, but in some it is
very marked.
PUGNAX.
Hall and Clarke, Pal. N. Y., VIII, pt. II, p. 202, (1893) ; Ann. Rep. N. Y. St. Geol., p. 208, for 1895.
Pugnax rockymontana. Plate XII, figs. 8, 8b.
Terebratula rockymontana Marcou, Geol. N. Amer., p. 59, pl. vi, f. 13,
(1858).
Rhynchonella eatonicwformis McChesney, New Pal. Foss., p. 49, (1869).
Rhynconella rockymontana White, Wheeler’s Expl.Surv. West 100 Mer.,
Iv, p. 131, pl. 1x, f. 1, (1875).
Pugnax eatoniiformis Hall and Clarke, Pal. N. Y., vit, pt. 11, p. 204,
(1893).
Pugnax rockymontana Schuchert, Bull. U. S. Geol. Surv. 87, p. 336,
(1897).
White’s description: ‘‘Shell rather large, inflated, subtri-
hedral in outline, broadest near the front ; sides somewhat regu-
larly rounded from the antero-lateral portions to the beaks.
. ne . re Py ey fe 7
BEEDE. | Carboniferous Invertebrates. 93
Ventral valve having its greatest convexity toward the beak;
sides sloping away from the middle with a slight convexity and
becoming flattened or sometimes a little concave near the lateral
margins; beak rather small, prominent, and closely incurved
over that of the other valve; mesial sinus very broad but not
deep, prolonged far upward at the front, becoming obsolete
about the middle of the valve and is entirely wanting upon its.
posterior portion; from two to four depressed, angular plica-
tions occupy the mesial sinus and disappear with it, the sides
and posterior portion being free from plications. Dorsal valve
more capacious than the ventral; mesial fold distinct at the
front, and, like the mesial sinus, becoming obsolete about the
middle of the valve; from three to five plications like those of
the other valve mark the fold, but the surface upon each side of
it is plain, like that of the posterior portions of both valves.
The whole surface marked by fine striw of growth, but no radi-
ating striw have been detected.’’ Measurements of two speci-
mens:—Length, 19 mm., 23 mm.; width, 19 mm., 23 mm.;
convexity, 15 mm., 16 mm.
Range and distribution: Lower Coal Measures ; Coffeyville,
Montgomery county, Kansas City.
Our specimens seem to agree very well with White’s figures
and description, the principal difference being that the lateral
angles at the beak seem to be a little smaller. While there are
no radiating stri# in our specimens, the shell is very fibrous
and where exfoliated might appear to be striated. It differs
from P. pugnus missouriensis in having no plications on the
sides of the shell, and also in outline.
Pugnax utah. Plate XII, figs. 7-7c.
Terebratula uta Marcou, Geol. N. Amer., p. 58, pl. v1, f. 12, (Feb. 1858).
Rhynchonella (Camarophoria) osagensis Swallow, Trans. St. L. Acad.
Sci., 1, p. 289, (June, 1858).
Rhynchonella utah Meek and Hayden, Proc. Acad. Nat. Sci. Phil., p. 27,
(1859): ete.
? Rhynchonella_ species Salter, Quart. Jour. Geol. Soc., London, xvii,
p. 64, pl. 1v, f. 5, (1861).
Camarophoria globulina Geinitz (non Phillips), Carb. u. Dyas in Neb.,
p. 38, pl. 111, f. 5, (1866).
Rhynchonella osagensis Meek, Fin. Rep. U. S. Geol. Surv. Neb., p. 179,
pl. 1, f. 9, pl. v1, f. 2, (1872); ete.
94 University Geological Survey of Kansas.
Pugnazx utah Hall and Clarke, Pal. N. Y., viii, pt. 11, p. 204, pl. Lx, ff. 39-
42, (1893).
Rhynchonella uta Keyes, Geol. Surv. Mo., v, p. 103, pl. x1, f. 7, (1895). .
Meek’s description: ‘‘Shell small, more or less variable in
form, often subtrigonal, generally wider than long, more or less
gibbous ; front truncated, or sometimes sinuous in outline; an-
terior lateral margins rounded in outline; posterior lateral mar-
vins convex, or nearly straight and converging toward the
beaks at an angle of from 90 deg. to 120 deg. Dorsal valve
more convex than the other, greatest convexity near the middle
or between it and the front, which has a broad, rather deep,
marginal sinus, for the reception of the corresponding projec-
tion of the front of the other valve; mesial fold somewhat flat-
tened, but slightly prominent, and rarely traceable back of the
middle of the valve; generally composed of three, but some-
times four—rarely more—plications; sides rounding down
rapidly on each side of the mesial fold, and each occupied by
about three or four simple plications; beak curving strongly
beneath that of the other valve; interior with a faint linear
mesial ridge, on each side of which is a raised curved line en-
closing an ovate space, occupied by the adductor muscular
impressions. Ventral valve distinctly less convex than the
other, with a broad, shallow, short sinus occupied by about two
or three short plications; anterior lateral margins on each side
of the sinus, with from two to four plications ; beak moderately
vrominent, and more or less arched, rather pointed; foramen
small.’’ Measurements: Length,8 mm.; width,9 mm.; con-
vexity, 7 mm. :
Range and distribution: Upper Coal Measures; Bronson,
Bourbon county, Kansas City, Iola, Olathe, Lawrence, Lecomp-
ton, Topeka, Beaumont, Grand Summit.
The young of this species are perfectly plain ana betray no
indication that they are of the same kind as the folded, plicated
adults. It can be readily distinguished from the previous spe-
cies by its smaller size, shallower.and more ill-defined sinus, and
the presence of plications on its sides.
BREDE. | Carboniferous Invertebrates. 95 |
DIELASMA.
King ines Ng eon ag Mar Mon. Perm. Foss. Eng. Pal. Soc., p. 46, (1850) ;. ete.
King, P ublin Univ. Zool. Bot. Asso., I, p. 260, (1859).
Beecher and Schuchert, Biol. Soc. Wash.. VIII, pp. 71-82, (1893).
Hall and Clarke, Pal. N. Wee VILL pt. II, p. 393, (1893) ; ete.
Dielasma bovidens.
Terebratula bovidens Morton, Amer. Jour. Sci., xx1x, p. 150, pl. 11, f. 4,
(1836): Meek, Fin. Rep. U. S. Geol. Surv. Neb., p. 187, pl. 1, f. 7, pl. 11,
f. 4, (1872); ete.
Terebratula millipunctata Hall, Expl. Surv. R. R. ‘Route Miss. R. to Pac.
Ocean, 111, p. 101, pl. 11, ff. 1, 2, (1856); etc.
Terebratula elongata Shumard (non Schloth.), Trans. St. L. Acad. Sci.,
I, p. 393, (1859).
Terebratula geniculosa McChesney, New Pal. Foss., p. 82, (1861); ibid.,
pl. 1, f. 2, (1865).
Dieiasma? bovidens White, Wheeler’s Expl. Surv. West 100 Mer., Prel.
Rep., p. 21, (1874).
Terebratula ( Dielasma) bovidens White, ibid., Fin. Rep.,1v, p. 144, pl.
x1, f. 10, (1875).
Terebratula hastata Mi a cat Sowerby), Mon. U. S. Surv., viii, p.
224, (1893): ete.
Dielasma bovidens Hall ad Clarke, Pal: N. Y., viii; pt. 11, pp. 295,. 296,
f. 213, pl. Lxxxt, ff. 29-35, (1893).
Meek’s description (in part): ‘‘Shell ovate, rounded and
rather compressed at the anterior and anterior lateral margins,
and the most convex a little behind the middle; valves nearly
equally convex; ventral valve strongly arcuate longitudinally,
and presenting a regularly increasing curve, from the front to
the beak, which is moderately prominent, and very strongly
and closely curved over and upon that of the other valve;
foramen a little oval and not truncating the immediate apex of
the beak, but situated directly outside of it; mesial sinus rather
wide, and rounded at the front, but narrowing and becoming
less deep further back, until it dies out near the curve of the
umbo, which is sometimes slightly flattened. Dorsal valve
often nearly straight, or but slightly convex along the middle,
from the beak to the front, where its margin is usually some-
what raised for the reception of the sinus; sides sloping from
the middle to the lateral margins along nearly the entire length
of the valve, but terminating directly under that of the other
valve, without any distinct curvature. Surface nearly smooth,
or showing moderately distinct marks of growth; and, by the
aid of the magnifier, exhibiting very distinctly the moderately
96 University Geological Survey of Kansas.
large regularly arranged. punctures.’’ The dental lamelle do
not extend much beyond the teeth, and are situated close to the
sides of the shell, so that the lateral space enclosed is very small.
There seems to be a rounded, indistinct ridge extending about
two-thirds the distance to the front om the interior of the ventral
valve. Loop situated a little posterior to the middle of the
shell, and is about one-third the length of the shell, the two
pointing obliquely forward. Measurements of two specimens
illustrating the variation: Length, 21 mm., 25 mm.; width,
14 mm., 19 mm. ; convexity, 10 mm., 16 mm.
Range and distribution: Upper Coal Measures: Kansas City,
Eudora, Lawrence, Lecompton, Topeka, Grand Summit.
SPIRIFERINA.
d’Orbigny, Paris Acad. Sci., Comptes Rendus, XXV, p. 268, (1847).
Hall and Clarke, Pal. N. Y., VIII, pt. II, p. 51, (1893); ete.
Spiriferina cristata.
Terebratulites cristatus Schlotheim, Beit. zur Naturg. der Verst., Akad.
der Wiss. zu Muenchen, pl. 1, f. 3, (1816).
Spirifer octoplicata? Hall (non Sowerby), Stansbury’s Expl. Gt. Salt
Lake, p. 409, pl. iv, f. 4, (1852).
Spirifer kentuckyensis Shumard, Geol. Surv. Mo., 1, p. 293, (1855); ete.
Spiriferina cristata Davidson, Quart. Jour. Geol. Soc. London, p. 170,
pl. 1x, f. 6, (1863); ete.
Spirifer laminosus Geinitz (non McCoy), Carb. u. Dyas in Neb., p. 45,
pl. 111, f. 19, (1866).
Spirifer kentuckyensis var. propatulus Swallow, Trans. St. L. Acad.
Sci., 11, p. 489, (1866).
Spiriferina kentuckyensis Meek, Fin. Rep. U. S. Geol. Surv. Neb., p.
185, pl. v1, f. 3a—d, pl. vir, f. lla, b, (1872); ete.
Spirifer (Spiriferina) kentuckyensis Hall, 2d Rep. N. Y. St. Geol., pl.
LxI, fi. 14-16, (1883).
Spiriferina cristata Schuchert, Bull. U. S. Geol. Surv. 87, p. 410, (1897).
Meek’s description: ‘‘Shell rather small, varying from sub-
globose to semicircular, or even subfusiform, always wider than
long; breadth sometimes twice or even three times the length ;
hinge line always equaling the greatest breadth of the valves,
occasionally greatly extended, and terminating in slender mu-
cronate ears; anterior and lateral margins generally forming
a nearly semicircular curve. Ventral valve somewhat more
convex than the other, the greatest convexity being between
the beak and the middle; beak moderately prominent and
BEEDE. | Carboniferous Invertebrates. 97
rather distinctly arched or incurved; area arched, usually of
moderate height, well defined, and extending nearly or quite
to the lateral extremities, while it increases rapidly in height,
with concave lateral margins toward the beak; foramen gen-
erally higher than wide, with a marginal furrow on each side,
and, so far as known, not closed by a deltidium; mesial sinus
narrow, rather deep, sometimes with a small obscure rib along
its middle, but more frequently without it; plications on each
side of the sinus about five to eight or nine, rather narrow,
simple, prominent, and a little rounded; mesial septum of the
interior moderately prominent. Dorsal valve with greatest con-
vexity near the middle; beak scarcely projecting beyond the
hinge margin, more or less incurved; area very narrow, and
incurved with the beak; mesial fold narrow, not very promi-
nent, nor greatly larger than’ the first plication on either side,
most generally rounded, but not infrequently with an obscure
sulcus along the middle, near the front; lateral plications as
in the other valve. Entire surface of both valves ornamented
with numerous closely crowded, very regularly arranged, sub-
imbricating lamelle of growth, strongly arched in passing over
the cost; over the whole may also be seen, by the aid of a
magnifier, numerous granules, apparently connected with the
punctures passing through the shell which are comparatively
large and distinct, though regularly arranged.’’ Measure-
ments of an average specimen: Entire length, 9 mm.; width,
15 mm.; convexity, 7 mm.
Range and distribution: Upper and Lower Coal Measures ;
Fort Scott, Bronson, Bourbon county, Thayer, Kansas City,
Lawrence, Lecompton, Topeka.
Not having foreign specimens for comparison, Schuchert is
followed in referring our specimens to S. cristata.
98 University Geological Survey of Kansas.
SPIRIFER.
Sowerby, Min. Con., II, P; 41 (1815).
Hall and Clarke, Pal. N. Y., VI ide pt. II, pp. 1-40, Wey ete.
Billings, Can. Jour., VI, p. 253, (1861) ; etc.
This genus is very grealty developed in the Paleozoic rocks of
America, though there are but two forms that have been recog-
nized with certainty in our Coal Measures. Hall (loc. cit.) di-
vides the genus into six groups, as follows:
I. Radiati. ‘‘Smooth, radially undulated or plicated; fold ~
and sinus smooth; entire surface covered with fine, filiform ra-
diating strie which may be minutely crenulated or granulose.’’
Il. Lamellosi. ‘* Radially plicated ; surface covered with nu-
merous concentric lamelle. In Silurian species the fold and
sinus are non-plicate ; the later forms usually bear a low median
depression on the fold accompanied by a corresponding median
ridge in the sinus.’’ He further divides this group into two
smaller ones —the Septati, those possessing a mesial septum in
the pedicle valve, and the Aseptat?, without this septum.
Ill. Fimbriati. ‘‘ Shells with a few low plications or none;
hinge line not greatly extended, often shorter than the greatest
diameter of the shell; dental lamell:e moderately, sometimes
notably, developed; a low median septum may exist in the
pedicle valve. Surface covered with concentric rows or fringes
of fine spines.’’ He also divides this group into two smaller
groups—the Unicispinei, those species in which the concentric
fimbriz are made up of short, simple, hollow spines (Delthyris
Dolman), and the Duplicispinei, those with larger, compound,
hollow spines (feticularia McCoy).
IV. Aperturatvy. Those having plications on the fold and
sinus. These are the typical Spirifers. They are divided into
several smaller sub-groups which are of little interest here.
V. Ostiolati. Median fold and sinus without plications.
VI. Glaberati. Surface smooth ; fold and sinus faintly devel-
oped. These he divides into two smaller groups, the Aseptati
and the Septati, according to the presence or absence of the
dental lamelle and mesial septa. These are equal to Martinea
of McCoy, Martinopsis Waagen, and Mentzelia Quenstedt.
BgEDE. | Carboniferous Invertebrates. 99
Our two species, Reticularia perplexa and Spirifer cameratus,
belong to the Fimbriati and Aperturati respectively. Spiriferina
is, in some respects, more closely related to the Lamellos?.
Spirifer cameratus. Plate XII, figs, 5—5e.
Spirifer cameratus Morton, Amer. Jour. Sci., xxrx, p. 150, pl. 1, f. 3,
(1836); ete.
Npirifer meusebachanus Roemer, Kreidebildung Texas, p. 88, pl. x1, f. 7
852).
Spirifer triplicatus Hall, Stanbury’s Expl. Surv. Gt. Salt Lake, p. 410,
pl. tv, f. 5, (1852).
Npirifer fasiger Owen (non Keyserling), Geol. Surv. Iowa, Wis., and
Minn., pl. v, f. 4, (1852).
Npirifer Peguidontats:' ? Owen, ibid., p. 586, pl. v, f. 6, (1852).
Npirifer striatus var. triplicatus {ae Geol. N. Amer., p. 49, pl. vil,
f. 3, (1858).
Npirifera camerata Newberry, Ives’ Rep. Colo. Riv. West, p. 127, (1861);
etc.
Npirifera camerata var. kansasensis Swallow, Trans. St. L. Acad. Sci.,
I1, p. 409, (1867).
Nptrifer (Trigonotreta) camerata Meek, King’s U.S. Expl. 40th ee
ive p. OL, pi; 1x, £..2,.(1817).
Shell medium to large in size, greatest convexity back of the
middle, variable in outline from subsemicircular to trigonal ;
anterior margin sharply rounded to truncate-sinuate ; lateral
margins slightly curved to nearly straight, pointing outward
and backward to the ears; hinge line equaling the greatest
width of the shell, sometimes prolonged into attenuate ears ;
cardinal area broad, extending to the extremity of the hinge
line; foramen broadly triangular and nearly equilateral, partly
closed in the upper part by a pseudo-deltidium ; beak high,
prominent, somewhat recurved over the cardinal area, which is
sometimes slightly arched; mesial sinus prominent, beginning
at the beak and broadening and deepening until the front mar-
gin is reached ; fold of the dorsal valve to correspond, and the
beak of the same moderately rounded beneath that of the other
valve. Interior of pedicle valve marked by a subelliptical
muscular impression in the vicinity of the beak, posterior end
of this impression extending to the hinge line or beyond,
bisected by an indistinct mesial ridge, radiating from which
are small indistinct ridges for the attachment of the muscles.
The cardinal area projects over a portion of the visceral cavity,
100 University Geological. Survey of Kansas.
leaving a large space beneath; the shell here is well pitted; a
small tooth and depression are developed on the inner corners
of the cardinal area. Shell of the brachial valve thin, muscu-
lar marking distinct ; hinge line at beak broadly and shallowly
arched, one prominent socket on each side of the arch, for the
teeth of the other valve ; two small elevations in the center for at-
tachments. The exterior markings of the shell consist of rather
large, bifurcating, radiating striz or cost, almost always fascicu-
lated, covering the entire shell to the tips of the ears, where
they are nearly parallel. There are, in unweathered, unworn
specimens, minute pustules arranged in somewhat radiating
order, as well as lines of growth visible on the front border. In
exfoliated specimens the fasciculation is less distinct. Meas-
urements of average specimen: Length, 32 mm.; breadth, 44
mm.; convexity, 21 mm.
Range and distribution: Upper Coal Measures; Kansas
City, Lawrence, Lecompton, Topeka, Grand Summit. Com-
mon throughout the Upper Coal Measures to the base of the
Permian.
The variation in outline, convexity and length of the hinge
line is great in this species. This species has been considered
by some as identical with S. striatus (Martin ) Davidson. Schu-
chert has pointed out characters which separate it from that
species.®° There is also a marked difference in the spires of the
two, those of S. striatus being long, loose, and acute, while those
of S. cameratus are short, compressed, and obtuse, and they en-
close a slightly smaller angle. The striw of S. cameratus are
nearly always fasciculated, while those of S. striatus are not.
There is also a marked variation in the interior of the pedicle
valve of this shell. In some specimens the muscular scar in
this valve is elliptical and extends well back beneath the del-
thyrium into the beak, the teeth are not supported by heavy
deposits of sheli or lamella, and the cavity formed by the jut-
ting of the cardinal area forms a general posterior cavity ex-
tending across the shell, only slight ridges being present below
6. Bull. U. S. Geol. Surv. 87, p. 384: “The latter species [S. striatus], however, is closely and
finely reticulated with concentric growth lines, while in S. cameratus the plications are covered
with small pustules, which are arranged in radiating lines.”’
we
BREDE. | Carboniferous Invertebrates. 101
and behind the teeth. In other specimens the posterior of the
shell is much thickened, the muscular impression not extending
beneath the beak or hinge line, that portion of the cavity being
filled with shell, which has encroached upon the muscular area
until it is small and nearly circular. This heavy deposit of
shell also forms strong supports for the teeth, dividing the pos-
terior cavity into a right and a left cavity.
AMBOCGLIA.
Hall, 13th Rep. N. Y. St. Cab. Nat. Hist., p. 71, ff. 1-3, p. 72, ff. 4 )
Meek and ‘Hayden, Pal. Upp. Mo., Smiths. Cont. Knowl., XIV, 17 2, p. 20, (1864); etc.
Ambocelia planoconvexa.
Spirifer planoconvexa Shumard, Geol. Rep. Mo., p. 202, (1855); ete.
Ambocelia gemmula McChesney, New Pal. Foss., p. 41, (1860); ibid.,
pl. 1, f. 3, (1865).
Spirifer (Martinia) planoconvexa Meek and Hayden, Pal. Upp. Mo.,
Smiths. Cont. Knowl., xtv, 172, p. 20, ff. a-e, (1864); Meek, Fin. Rep.
U.S. Geol. Surv. Neb. pe 188, pl.1¥, f. 4, ply-virt, £. 25: ete.
Martinia planoconvexa McChesney, Trans. Chic. Acad. Sci., 1, p. 34,
pl. 1, f. 3, (1868).
Ambocelia planoconvexa Hall and Clarke, Pal. N. Y., vii, pt. 11, p. 56,
pl. xxx1x, ff. 10-15, (1893). ;
Meek’s description: ‘‘ Shell very small, plano-convex, or very
rarely a little concavo-convex, sometimes wider than long, in
other examples slightly longer than wide; hinge margin always
shorter than the greatest transverse diameter of the valves, and
rounded at the extremities ; lateral margins and front regularly
rounded ; surface apparently smooth, excepting a few very ob-
scure concentric marks of growth, but, when examined by the
aid of a magnifier, showing the remains of the bases of minute
hair-like spines. Dorsal or smaller valve truncato-suborbicular
in outline, generally nearly flat, with faint longitudinal depres-
sion in front, sometimes slightly convex near the beak, and con-
cave around the anterior and lateral margins; beak scarcely
distinct from the cardinal margin; area narrow, but well de-
veloped, or about half as large as in the other valve; socket
plates projecting like diverging teeth on each side of the small
fissure. Ventral valve very gibbous, particularly in the umbonal
region, sometimes with obscure traces of a narrow longitudinal
depression along the middle, but without a proper mesial sinus ;
beak very prominent and strongly arched back over the hinge ;
102 University Geological Survey of Kansas.
area subtriangular, being moderately high under the beak, but
narrowing rapidly, with moderately defined concave margins,
to the extremities of the hinge, and arching with the beak;
fissure rather narrow, or higher than wide, apparently rounded
above under the beak, and spreading at the hinge.’’ Measure-
ments: Length,9 mm.; width, 10 mm.; convexity, 5 mm.
Range and distribution: Upper and Lower Coal Measures ;
Fort Scott, Fredonia, Eudora, Lawrence, Lecompton, Topeka,
Emporia.
RETICULARIA.
McCoy, Carb. Foss. Ireland, p. 142, (1844).
Waagen, Pal. Indica, ser. xiii, I, p. 538, (1893).
Reticularia perplexa. Plate XII, figs. 4—4c.
Spirifer lineatus Shumard, Geol. Surv. Mo., p. 216, (1855); ete.
Spirifer perplexus McChesney, New Pal. Foss., p. 43, (1860).
Spirifer lineatus var. perplexus Swallow, Trans. St. L. Acad. Sci., 11, p.
408, (1866).
Spirifera lineata Meek, Fin. Rep. U. S. Geol. Surv. Neb., pl. 11, f. 3,
(1872); etc.
Spirifer (Martinia) perplexa Derby, Bull. Cornell Univ., p. 16, pl. 11,
ff. 27, 39, 40, 45, 50, pl. vir, f. 13, (1874).
Spirifer (Martinia) lineata? White, Wheeler’s Expl. Surv. West 100°
Mer., 111, Appendix, (1881); later without query; etc.
Spirifera perplexa Keyes, Geol. Surv. Mo., v, p, 84, (1895).
Reticularia perplexa Schuchert, Bull. U.S. Geol. Surv. 87, p. 342, (1897).
White’s description: ‘‘Shell moderately gibbous, trans-
versely subelliptical in marginal outline, the front and sides
regularly rounded; hinge much shorter than the width of the
shell; cardinal extremities rounded; cardinal area distinct,
arched, and moderately high; ventral valve convex; umbonal
portion prominent; beak prominent, incurved; area small,
without median sinus, but there is a slight flattening of the
valve in front, which gives the front margin a slight sinuosity ;
dorsal valve regularly convex, both transversely and longitudi-
nally ; umbonal portion prominent, but not so much so as that
of the other valve; beak moderately prominent and projecting
a little beyond the hinge line; surface marked by very numer-
ous very faint radiating lines and somewhat stronger concentric
lines, the latter being impressed and finely crenulate, the
minute crenulations apparently marking the bases of hair-like
BEEDE. | Carboniferous Invertebrates. 103
spines when the surface of the shell is perfect.’’ Measure-
ments: Length, 19 mm.; width, 23 mm.; convexity, 15 mm.
Range and distribution: Upper and Lower Coal Measures ;
Fort Scott, lola, Lawrence, Topeka, and so on to the base of
the Permian.
The shell varies a great deal in marginal outline, some hay-
ing the beak prolonged and sides.somewhat compressed, giving
it a very different appearance from the normal form.
HUSTEDIA.
Hall and Clarke, Pal. N. Y., VIII, pt. II, p. 120, (1893) ; 13th Ann. Rep. N. Y. St. Geol.,
p. 197, (1895).
Hustedia mormoni. Plate IX, figs. 10-10d: plate X, fig. 3.
Terebratula mormoni Marcou, Geol. N. Amer., p. 51, pl. v1, f. 11, (Feb.
1858); etc. ,
Retzia punctulifera Shumard, Trans. St. L. Acad. Sci., 1, p. 220, (June,
1858); Meek, Fin. Rep. U.S. Geol. Surv. Neb., p. 181, pl. 1, f. 13, pl. v,
f. 7, (1872); ete.
Retzia mormoni Meek and Hayden, Proc. Acad. Nat. Sci. Phil. 1859, p.
27; etc.
PRetzia subglobosa McChesney, New Pal. Foss., p. 45, (1860): ibid., pl. 1,
f. 1, (1865).
Retzia compressa Meek, Geol. Surv. Cal., 1, p. 14, pl. 11, f. 7, (1864); ete.
Retzia radialis Walcott (non Phillips), Mon. U.S. Geol. Surv., vimr, p.
220, pl. 7, ff. 5d—h, (1893); etc.
FEumetria punctulifera Derby, Bull. Cornell Univ., 1, p. 4, pl. viit, ff. 4,
&, 1, 8, 10, pl. x, f.. 3, (1874).
Hustedia mormoni Hall and Clarke, Pal, N. Y., vit, pt. 11, p. 120, f. 196,
pl. x1, ff. 1-9, (1893).
Meek’s description: ‘‘Shell small, ovate; in mature speci-
mens, gibbous; hinge line short, or scarcely extended enough
to show distinctly the little ears at the extremities. Ventral
valve more convex than the other, the greatest convexity being
between the middle and the umbo, which is prominent, rounded,
more or less strongly arched, and provided with a moderately
large circular foramen; area well defined, triangular, and arch-
ing with the beak. Dorsal valve most convex near the middle ;
beak extending a little beyond the hinge margin, and distinctly
incurved. Surface of each valve ornamented by fourteen or fif-
teen (very rarely sixteen to seventeen) simple, rather prominent,
radiating cost, one or two of which are sometimes slightly
more depressed than the others, near the front of the ventral
104 University Geological Survey of Kansas.
valve, so as to cause some appearance of an obscure mesial sinus,
but without producing any corresponding mesial elevation on
the other valve, or visibly inturrupting the general straightness
of the uniting margins of the two valves; lines of growth ob-
scure; punctures visible under a good pocket lens, and very
regularly disposed.’’ Measurements: Length, 10 mm. ; width,
S$ mm.; convexity, 7 mm.
Range and distribution: Upper Coal Measures; Fort Scott,
Iola, Kansas City, Lawrence, Lecompton, Topeka, Beaumont.
CLEIOTHYRIS.
King (non Phillips), Mon. Perm. Foss., Pal. Soc., p. 137, (1850).
Hall and Clarke, Pal. N. Y., VIII, pt. II, p. 90, (1893) ; ete.
Cleiothyris roissyi. Plate XII, fig. 2; plate XI, figs. 5-5e.
Spirifer de roissyi L’Eveille, Mém. Soc. Geol. de France, 11, p. 39, pl. 11,
ff. 18-29, (1835).
Terbratula roysti Marcou, Geol. N. Amer., p. 51, pl. vi, f. 10, (1858).
Athyris sublamellosa Hall, Geol. Surv. Iowa, 1, pt. 11, p. 702, pl. xxviz,
f. 1, (1858); ete.
Athyris parvirostris Meek and Worthen, Proc. Acad. Nat. Sci. Phil.,
1860, p. 451.
Spirigera americana Swallow, Trans. St. L. Acad. Sci., 1, p. 89, (1863).
Spirigera pectenifera Swallow (non Sowerby), ibid., p. 88.
Athyris planosulcata Geinitz (non Phillips), Carb. u. Dyas in Neb., p. 42,
(1866); ete.
Spirigera planosulcata? White, Wheeler’s Rep. Geog. Expl. Surv. West
100 Mer., iv, p. 287, pl. rv, ff.-10, 11, (1877).
Athyris hirsuta Walcott, Mon. U.S. Geol. Surv., viii, p. 222, pl. xviut, f. 5,
(1884). ,
Cleiothyris royssii Hall and Clarke, Pal. N. Y., viii, pt. 1, p. 91, pl. xLv1,
ff. 23, 24, pl. Lxxxtv, f. 32, (1893).
Cleiothyris sublamellosa Hall and Clarke, ibid., p. 91.
Meek and Worthen’s description: ‘‘Shell of medium size,
quadrato-subcircular, moderately gibbous, length and breadth
nearly equal, sometimes a little wider than long (other ex-
amples a little longer than wide); greatest convexity at the
middle; valves equally gibbous ; lateral margins usually promi-
nent, and narrowly rounded in outline at the middle, thence
converging with a slightly convex outline to the faintly sub-
truncate front; postero-lateral margins a little inflected, so as
to form slight concavities, converging to the beaks at an angle
of about ninety-seven degrees. Both valves destitute of a mesial
et i t,o ee
BREDE. | Carboniferous Invertebrates. 105
fold or sinus, but each sometimes slightly flattened in the mid-
dle near the front, where they meet without the slightest sinu-
Osity in the margin of either. Beak of ventral valve small,
pointed, closely incurved upon that of the other valve, which is
but little less prominent ; foramen round and very small. Sur-
face with small, obscure, concentric marks of growth. Spiral
appendages each making about twelve turns. Surface ( proba-
bly exfoliated ) showing only small, obscure, concentric ridges.’’
Measurements, two specimens: Length, 10 mm., 10 mm. ; width,
11 mm, 13 mm.; convexity, 6 mm, 5 mm.
tange and distribution: Lower Coal Measures; Fort Scott,
Marmaton station, Bourbon county, Prescott, Linn county.
This rare little fossil is rather variable in form. Most of our
specimens are rather more straight along the hinge than those
usually figured from this country, approaching, in this respect,
the form figured by C. Nikitin from the Upper Carboniferous
and Lower Permian of Europe, one specimen in particular be-
ing quite straight and wide along the hinge. The shells, in un-
worn specimens, show the surface fimbrizw very well.
SEMINULA.
McCoy, Carb. Foss. Ireland, pp. 550, 558, (1844).
Hall and Clarke, Pal. N. Y., VIII, pt. II, p. 93, (1893) ; ete.
Seminula argentea. Text fig. 3, C.
Terebratula argentea Shephard, Amer. Jour. Sci., xxxtv, p. 152, f. 8, (1838).
Terebratula roisyi d’Orbigny [non L’Eveille], Voy. dans l’Amé¢r. Mér.
Pal., p. 46, (1842).
Terebratula antisicnsis d’Orbigny, ibid., p. 46, (non 36).
Terebratula peruviana @’Orbigny, ibid., pl. 11, ff. 17-19, (non p. 36).
Terebratula subtilita Hall, Stanbury’s Expl. Gt. Salt Lake, p. 409, pl. rv,
ody.2,-(1852)3 ete;
Terebratula? subtilita Davidson, Mon. Brit. Carb. Brach., p. 18, pl. 1, ff.
21, 22, (1857); etc.
Npirigera subtilita Meek and Hayden, Proc. Acad. Nat. Sci. Phil., p. 20,
(1859); ete.
Athyris differentis McChesney, New Pal. Foss., p. 47, (1860).
Athyris subtilita Newberry, Ives’s Rep. Colo. Riv. West, p. 126, (1861);
Meek, Fin. Rep. U. S. Geol. Surv. Neb., p. 180, pl. 1, f. 12, pl. v, f. 9, pl.
vill, f. 4, (1872); ete.
Seminula subtilita Hall and Clarke, Pal. N. Y., vii, pt. 1, p. 95, ff. 66,
67, p. 86, ff. 58, 59, pl. xxvii, ff. 17-31, (1893).
Athyris argentea Keyes, Geol. Surv. Mo., v, p. 92, pl. xxxrx, f. 11, (1895).
Neminula argentea Schuchert, Bull. U. S. Geol. Surv. 87, p. 377, (1897).
8—vi
106 Unwersity Geological Survey of Kansas.
Meek’s description (in part): ‘‘Shell ovoid, being usually
widest a little in advance of the middle, and nearly always
somewhat longer than wide, moderately convex, becoming
rather gibbous with age. Ventral valve usually a little more
convex than the other, its greatest convexity being generally
behind the middle; beak prominent, rounded, and distinctly
incurved upon that of the other valve; foramen round, of mod-
erate size, and truncating the immediate apex of the beak;
mesial sinus absent, or very shallow, in young or compressed
individuals, but well defined, and round, flattened or angular
in adult gibbous specimens, in which it rapidly increases in
size, from near the middle to the front, where it produces a
more or less prominent marginal projection, fitting into a cor-
responding sinuosity in the margin of the other opposite valve.
Dorsal valve moderately convex, the greatest convexity in small
or compressed specimens often near the middle or between it
and the umbo, but in large, gibbous individuals, with a well-
defined, prominent mesial fold, sometimes near the front; beak
rather distinctly incurved under that of the opposite valve.
Surface of both valves nearly smooth, or with mere lines of
growth, in young shells, but in large or mature specimens with
well-defined, imbricating marks of growth on the anterior half;
exfoliated surfaces also show, under a magnifier, obscure traces
of radiating strie.’’ The spires are large and generally acutely
pointed. Measurements of two specimens: Length, 36 mm.,
26 mm.; width, 37 mm., 23 mm.; convexity, 26 mm., 13 mm,
Range and distribution: Upper and Lower Coal Measures ;
Marmaton station, Bourbon county, lola, Kansas City, Eudora,
Lawrence, Lecompton, Topeka, Manhattan, Grand Summit.
Very abundant throughout the Coal Measures of the state.
Hall has figured this species as having the ascending edges of
the spire straight and forming an obtuse angle at the apex. In
all the specimens at hand the sides of the spires are concave
and the tip of the spire is acute. However, they vary a great
deal in this respect. Swallow has described another form, S.
caputserpentis,’ from the Kansas Carboniferous, but I am able
7. Trans. St. L. Acad. Sci., II, p. 90, 1863.
-_—— .= © A ee ae eee oe
ee i ee te iN el i ie).
:
;
;
BEEDE. | Carboniferous Invertebrates. 107
to recognize but the one species, and think there is but little
doubt that his Spirigera caputserpentis is the same as the form
above described. The shells vary enough to easily include
Swallow’s description. The microscopic indications of stri«w
mentioned in the various descriptions of this shell are probably
due to the fibrous structure of the shell, which is quite coarse.
PELECYPODA.
Pelecypods (sometimes called lamellibranchs) are a group
of animals known under the names of mussels, clams, and oys-
ters. They live in fresh and salt water, and are covered by a
shell which is made up of two halves, or valves. Brachiopods
haye one valve on top and one on the bottom of the animal, but
the halves of the clam shell are located one on the right side
and one on the left. The body is nearly enclosed in a mantle
or fleshy membrane, which nearly surrounds the soft part of
the body. It is divided into two parts or halves, called lobes,
which secrete the two halves of the shell. Sometimes this
mantle is somewhat grown together for a large part of the way
along the lower sides of the animal near the open edges of the
shell.
In those with the mantle edges fastened together there are
two openings in the rear end; these openings are sometimes
prolonged into tubes called siphons, which are used in breath-
ing. A current of water passes constantly in through one of
these and out through the other. In those which do not have
the mantle united there is generally no siphon.
The edge of the mantle is also attached to the shell, impress-
ing a line nearly parallel with the edge of the shell. When the
siphons are present there is an inward notch or angle in the
line in the back part of the shell, caused by the muscles used in
pulling the siphon into the shell. When the siphons are ab-
sent the notch is not present. The line formed by the edge of
the mantle is called the pallial line.
Near the front of the shell lies an organ called the foot, which
108 University Geological Survey of Kansas.
is generally a muscular organ, hatchet-shaped in most fresh-
water clams; it aids the animal in moving. Sometimes this
organ secretes a substance which hardens quickly on being ex-
posed, and by this means the animal cements its shell firmly
to some rock or other object. The foot generally becomes
small in such cases on account of disuse. There is an opening
in one of the valves for it, so that the attachment may be more
certain. This opening is called the byssal groove or notch, and
the foot, in this case, is called the byssus.
There are other muscles in the clam which are important.
One or two large muscles are used in closing the shells together,
and run nearly directly across from one shell to the other, and
when two are present one is located near the front of the shell
and the other near the rear of it, and are known as the anterior
and posterior adductors, respectively. ‘These muscles are firmly
attached to the shell, causing a depression in it, showing the
size, shape and position of the muscle.
The pelecypods have no true heads. In the front part of
the animal is an opening, or mouth, surrounded by a pair of
membranous flaps. In the oyster these flaps are called the
beard. This opening is the end of the esophagus, or gullet,
which leads to the stomach, which is generally surrounded by
the liver. From the stomach the intestine is somewhat convo-
luted, generally passing through a part of the heart, and finally
ending near the back part of the shell.
The circulatory system consists of a heart, having two or
three chambers and afew arteries. The heart propels the blood
from the gills through the body of the animal. The respira-
tory organs consist generally of two pairs of gills. The water,
in the siphoned clams, enters through one side of the siphon,
or one tube, flows over the gills, constantly bathing them with
fresh water, and then flows past the mouth, where the food
particles are selected out of it, when it passes out of the other
siphon tube.
The nervous system consists of two knots of nerves near the
mouth and one near the posterior adductor muscle.
Now, since we have a general idea of the anatomy of the
;
)
|
;
BEEDE. | Carboniferous Invertebrates. 109
pelecypod, let us turn our attention to the shell. The hard
parts of animals are the only ones preserved in the rocks. In
the clams the shell only is found, and the impressions on the
inside of it are all we have to judge from as to the exact nature
of the animal which inhabited it; consequently, it is of great
importance to know what these markings are and what they
represent.
The elevated portions of the shell, which project upward near
the hinge, or where the two halves are fastened together, are
called the beaks or umbones. These generally point toward
the front of the shell. If we hold the shell so that the beaks
point away from us, the half on the right side will be the right
valve and the other half the left valve. The edge of the shell
on which the beaks are situated is the dorsal edge or portion of
the shell. The side opposite ( below) this is called the ventral
portion, and its edge is called the ventral or pallial margin.
The line where the two halves are united is called the hinge
line. The hard, gristly material that forms the hinge is called
the ligament. The projections along the hinge line, which fit
into the corresponding sockets in the opposite valve, are called
the teeth. We have previously mentioned the adductor scars,
the pallial line, and the notch in the pallial line caused by the
muscle (retractor) which pulls in the siphon.
The pelecypods described in the following page were inhabi-
tants of salt water.
I wish to acknowledge the kind aid in this work given by Mr.
Austin F. Rogers. He has also found and worked out several
species in the following list since the work closed. For full
bibliography the reader is referred to Weller, Bull. U.S. Geol.
Surv. No. 153, (1898).
\n Ww
SII)
<0 Peles
—T—-AX
Fic. 4. 1. Anatomy of a bivalve mollusk, Mya arenaria (after Woodward) ; left valve and
mantle lobe, and half the siphons, are removed: ss, respiratory siphons (the arrows indicate
the direction of the currents) ; aa!, adductor muscles; 0, gills; h, heart; 0, mouth, surrounded
BEEDE.]| Carboniferous Invertebrates. i tat
PLACUNOPSIS.
Morris and Lycett, Mon. Foss. Great Oolite, (1853).
: Placunopsis carbonaria. Plate XX, fig. 18.
: Placunopsis carbonaria Meek and Worthen, Proc. Chic. Acad. Sci., 1, p.
; 13, (1866): Geol. Surv. Ill., v, p. 578, pl. xxvir, f. 2, (1873).
pe oe
Meek and Worthen’s description: ‘‘Shell orbicular, com-
pressed subhemispherical, extremely thin. Upper or right
valve irregularly convex, sometimes rather gibbous; hinge
margin straight, generally equaling about one-third to one-
half the diameter of the valves, and usually showing slight
disposition to develop small, obtuse ears at the extremities ;
beak very small, sometimes compressed and nearly obsolete,
marginal, but not projecting distinctly beyond the cardinal
border, located at the middle of the hinge, and showing a slight
forward curvature. Under valve flat or conforming to the in-
equalities of the surface to which it was attached; beak mar-
7 ginal, and very nearly obsolete. Surface of both valves marked
. with irregular, undulating or interrupted radiating lines, with
} broad, faint, irregular concentric wrinkles, and a set of obscure
strie of growth. Often there is also an entirely independent
; series of parallel ridges, crossing the umbonal region, or some-
times the whole surface obliquely and partly, or sometimes
almost entirely, obliterating the other markings. Diameter of
the largest specimen found, from the hinge to the pallial mar-
gin, 1.30 inches; oblique parallel ridges uniformly numbering
seven to eight in 0.2 inch.’’
Range and distribution: Upper Coal Measures; Topeka.
>
ee ee eee eS a Se
by (p) labial palpi; /, foot; +, anus; m, cut edge of the mantle (after Nicholson). 2. Left valve
of Cytheria chione (after Woodward): A, anterior margin; #, posterior margin; (, ventral
margin or base; u, umbo; h, ligament; c, cardinal tooth; //, lateral teeth; a, anterior adduc-
tor; a‘, posterior adductor; p, pallial line; s, pallial sinus caused by the retractor muscles of
the siphons (after Nicholson). 3. Diagrammatic representation of the anatomy of a siphonate
pelecypod; the left valve and the left mantle lobe are re:noved, and the siphons are cnt short:
u, umbo; ca, cartilage pit; 0, the mouth; /p, labial palpi; a, stomach, surrounded by liver; i,
intestine perforating the heart; r, rectum, terminating in the anus; ad, anterior adductors,
pd, posterior adductors; n, supraesophageal or cerebral ganglion (the mouth is a little
displaced upwards, so that the ganglion comes to lie below the gullet instead of above it); 7’,
' parieto-splanchnic or branchial ganglion; /, foot; rx, cut edge of the right mantle lobe; rs,
retractor muscles of the siphon; 47, branchi# of the left side; gy, renal organ; s, inhalant si-
phon; s', exhalant siphon (after Nicholson).
=". - a 2s
112 University Geological Survey of Kansas.
LIMA.
Brugueire, Encycl. Meth., (1791).
Deshayes, Desc. de Coq. Foss. des Iny. de Paris, (1824).
Lima retifera. Plate XIII, fig. 5.
Lima retifera Shumard, Trans. St. L. Acad. Sci., 1, p. 214, (1858); Meek,
Fin. Rep. U.S. Geol. Surv. Neb., p. 188, pl. 1x, f. 5, (1872). See Weller,
Bull. 153, U.S. Geol. Surv., p. 324, (1898). |
Meek’s description: ‘‘ Shell obliquely subovate, moderately
convex, apparently not gaping in front; hinge line short, or
between one-half and one-third the antero-posterior diameter
of the valves; base forming a nearly regular semicircular curve ;
anterior side extended obliquely forward, rather narrowly
rounded below, and straight or slightly concave in outline, with
a rather long, oblique slope to the hinge above; posterior side
distinctly shorter than the other, and rounding from near the
ear into the base; ears subequal, the front margin of the ante-
rior one forming an obtuse angle with the hinge line, rather
distinctly flattened from the swell of the umbo, and somewhat
extended along the anterior margin below; posterior ear a little
more convex than the other, with its upper margin incurved,
and its lower margin separated from the umbo by a faint ob-
lique furrow, sometimes faintly sinuous behind, and nearly
rectangular at its extremity ; umbones rather convex or mod-
erately compressed, extending very little above the cardinal
margin, and placed near the middle of the same; surface orna-
mented by about twenty-five slightly irregular, angular, radiat-
ing cost, about equaling the spaces between, and occasionally
bifurcating on the umbones and lateral margins of the body
part of the valves, where they become obsolete; crossing all of
these, as well as on the ears and lateral margins, are numerous
fine concentric stric. Height, 15 mm.; length, 18 mm.
Range and distribution: Upper Coal Measures; Lawrence,
Topeka.
—~— ee
BREDE. | Carboniferous Invertebrates. 113
ENTOLIUM.
Meek, Geol]. Sury. Cal. II.
Entolium aviculatum. Plate XIX, fig. 1.
Pecten aviculatus Swallow, Trans. St. L. Acad. Sci:, 1. p. 213, (1858).
Entolium aviculatum Meek, Fin. Rep. U. 8S. Geol. Surv. Neb., p. 189, pl.
Ix, ff. lla—g, (1872).
Meek’s description : ‘‘ Shell compressed lenticular, very thin,
nearly or quite equivalve, suborbicular, or broad subovate in
outline exclusive of the ears, the antero-posterior diameter
being often a little less than that at right angles to the same ;
sides and base more or less regularly rounded; lateral margins
above the middle apparently a little gaping, straight,.and con-
verging to the beaks at an angle of 115° to 125°; cardinal
margin very short, or less than one-third the transverse diam-
eter of the valves, and in the left valve generally concave, or
more or less sloping in outline from the extremities of the ears
to the beaks; straight or nearly so in the right valve; ears
small, flat, very nearly equal, obtusely angular at the extremi-
ties, and separated from the body of the valves by an impressed
line, not defined by any proper sinus in either valve, though
the broad obtuse notch separating the anterior ‘one from the
straight, sloping adjacent margin is slightly more defined than
the other; beaks small, rather compressed, equal, and not pro-
jecting beyond the cardinal margin. Each valve with two
shallow undefined impressions diverging from the beak nearly
to the anterior and posterior margins; that on the posterior
side being longer than the other. Surface with very fine close
concentric striw, scarcely visible without the aid of a magnifier ;
crossing these are sometimes seen traces of extremely minute
radiating strive, curving gracefully outward toward the lateral
margins.’’ Ina foot-note he says: ‘‘In most of these speci-
mens these radiating strizw are entirely obsolete, even as seen
under a magnifier; and it is generally only on specimens that
have been slightly weathered that they are most distinctly seen,
while even on these they seem to be more due to some pecul-
iarity of the shell structure than proper surface sculpturing, the
shell showing a disposition to crack along these curved lines.
114 University Geological Survey of Kansas.
Both these and the concentric strize are almost invisible to the
unassisted eye.’’
‘‘Anterio-posterior diameter of a specimen a little under me-
dium size, 0.85 inch; height, 0.89 inch; length of the hinge
line, 0.27 inch. Specimens are sometimes found of nearly
double these dimensions.’’ In some delicately preserved casts
there are traces of zigzag markings on the anterior of the ven-
tral margin of the left valve. 3
Range and distribution: Upper Coal Measures; Lawrence,
Topeka.
AVICULOPECTEN.
McCoy, Ann. Mag. Nat. Hist., 2d ser., VII, p. 171, (1851 ).
Aviculopecten occidentalis. Plate XIII, fig. 7.
Pecten occidentalis Shumard, Geol. Rep. Mo., p. 207, pl. c, f. 18, (1855) ;
Newberry, Ives’s Colo. Expl. Exped., p. 128, (1861).
Pecten cleavelandicus Swallow, Trans. St. L. Acad. Sci., 1, p. 184, (1858).
Aviculopecten—— ? Meek and Hayden, Pal. Upp. Mo. (Smiths. Cont.
Knowl., x1v), p. 50, pl. 11, f. 10, (1864).
Aviculopecten occidentalis Meek and Toren, Geol. Surv. Ill., 1, p.
331, pl. xxvit, ff. 4-5a, (1866); Meek, U.S. Geol. Surv. Neb., p. 191, pl.
1x, f. 10, (1872); etc.
Pecten missouriensis ? Geinitz, Carb. u. Dyas in Neb., p. 36, pl. 11, f. 18,
(1866).
Meek’s description: ‘‘Shell distinctly inequivalve, not ob-
lique; subovate exclusive of the ears; lateral and basal mar-
gins regularly rounded; hinge margin nearly or quite equaling
the greatest breadth of the valves; cardinal plate of moderate
breadth. Left valve convex, with ears subequal; anterior
one with distinct radiating costs, more convex, shorter,
and more obtuse than the posterior, as well as more defined
from the swell of the umbonal slope; posterior ear flattened
and more angular at the extremity than the other, some-
times without radiating costz, but in other instances having
them more or less developed, each separated from the mar-
gin below by a rounded, rather broad, more or less deep sinus.
Right valve nearly flat and having the general outline of the
other excepting that its beak is scarcely distinct from the car-
dinal margin, and its anterior ear much narrower and defined
by a deep, sharp angular sinus. Surface of valve ornamented
7"
7a
'<
ee eee eae
BEDE. | Carboniferous Invertebrates. 115
with rather depressed or flattened irregular radiating cost, of
which only about twelve or fourteen of the largest reach the
beak, others dying out at various distances between the mar-
gins and the umbo; in proportion to size, the larger of the
intercalated ones being longer than the smaller. Crossing all
of these are numerous fine concentric strive, some of which on
the ears, particularly the anterior one, often form little vaulted
scales; in well-preserved specimens these vaulted projections
are strongly developed on one of the posterior costz of the body
part of the valve. Surface of right valve generally with only
very obscure radiating costz and fine crowded lines of growth.’’
Measurements: Length, 24 mm.; height, 26 mm.; convexity,
about 5 mm.
Range and distribution: Upper Coal Measures; Turner,
Eudora, Lawrence, Topeka, Wabaunsee county.
Aviculopecten rectilaterarius.
Avicula rectilateraria Cox, Geol. Surv. Ky., m1, p. 578, p. 1x, f. 2, (1857).
Aviculopecten rectilaterarius Meek and Worthen, Geol. Surv. Ill., 0, p.
326, (1866).
Shell variable in outline, about as long as high; ventral mar-
gin rounded, nearly semicircular; posterior margin nearly
straight, meeting the hinge at nearly a right angle; anterior
margin slightly sinuate at the lower part of the ear, then, round-
ing upward, meets the hinge line; iower anterior margin more
abruptly rounded than the corresponding posterior one; hinge
line slightly shorter than the shell, the beak not extending
above the hinge; posterior ear not separated from the shell by
a sulcus, anterior ear separated from it by an indistinct furrow.
The anterior ear of the right valve contains a sharp byssal
notch. The shell is marked by fairly prominent, slightly sinu-
ous, radiating strive, which increase by implanation and bifur-
cation; on the posterior ear of the left valve these striw are
very dimly yisible but are more prominent on the anterior ear.
These strizw are crossed by very fine, close, concentric strive and
larger lines of growth. Measurements, two specimens: Length,
18 mm., 18 mm.; height, 19 mm., 15 mm.
116 University Geological Survey of Kansas.
Range and distribution: Upper Coal Measures; Leaven-
worth, Wyandotte county.
Specimens are abundant in the shales at Topeka which may
belong to this species or to A. whitec. The material in hand is
too poor to assign with certainty to either species. The proba-
bility is, however, that they belong to A. whitec.
This species can be easily distinguished from A. occzdentalis
by its smaller size, thinner shell, and the fact that the posterior
ear is not so distinct and the shell is less elongate vertically.
Aviculopecten interlineatus. Plate XIIJ, fig. 6.
Aviculopecten interlineatus Meek and Worthen, Proc. Acad. Nat. Sci.
Phil:,) p. 454, (1860)-5 Geol, Surv. Wear, p: 3295 ole xexvinw fied ommweras
fie. (1866); Whitfield, Ann: N: Y. Acad? Sci., v, p: 694, ply xvay ieee:
11, (1891); Keyes, Geol. Surv. Mo:, v, p. 112, (pl. xm, foeG (soe)
Whitfield, Geol. Sury. Ohio, vit, p. 489, pl. x11, ff. 10, 11.
Aviculopecten ? interlineatus White, U. S. Geog. Surv. West 100 Mer.,
Iv, p. 149, pl. xa, f..3, (1877); 13th Ann: Rep: Ind. St: Geolz; pp, 1455 gale
SO0.6, 115 s (( fetes),
Meek and Worthen’s description: ‘‘Shell (left valve) rather
small, compressed, broad ovate or subcircular exclusive of the
ears, not oblique; length and breadth nearly equal; hinge
straight, about equaling the greatest breadth of the valve
below, ranging at right angles to the vertical axis of the shell.
Base regularly rounded; -posterior and anterior margins
rounded from below the ears to the base. Anterior ear tri-
angular, flattened so as to be very distinct from the umbonal
slope; posterior ear somewhat larger than the other, com-
pressed, triangular, the hinge side being longer than either of
the others, rather acutely angular at the extremity. Beak com-
pressed, a little nearer the anterior than the posterior extremity
of the hinge; umbonal slopes diverging from the beaks at an
angle of about 78°; anterior one subangular. Surface orna-
mented by about fifteen regular, very prominent, slender, and
obscurely crenulated concentric cost, which are separated by
spaces from four to six times their own breadth, excepting on
and near the ears; spaces between the cost occupied by numer-
ous fine, regular, closely arranged concentric strive, which are
crossed by faint indications of radiating ribs. Diameter, from
BEDE. | Carboniferous Invertebrates. tT
the pallial border to the hinge, 15 mm.; breadth, from the
anterior to the posterior margin, 16 mm.; convexity, 3 mm.
Right valve unknown.’’ In the discussion they also add:
‘« This exceedingly beautiful species seems to have varied a
little in the details of its surface markings at different ages.
In young shells the regular concentric cost seem to be nearly
or quite smooth, but as the shell advanced in its growth, they
gradully assumed a regularly crenulated outline, and became
themselves ornamented with extremely fine longitudinal striw,
not visible without the aid of a lens. ‘These striw are very
much finer and less distinct than those occupying the spaces
between the coste. ‘The faint indications of radiating costa,
seen crossing the concentric striw in the depressions between
the concentric ribs, are also wanting in young shells.”’
Range and distribution: Upper Coal Measures; Kansas City.
The absence of radiating strie and costie and the strong con-
centric cost with the fine concentric strie between them easily
and cleariy distinguish this species from all others of the genus
Aviculopecten carboniferus. Plate XIII, fig. 9.
Pecten carboniferus Stevens, Amer. Jour. Sci. and Arts, xxv, p. 261,
(1858). .
Pecten broadheadii Swallow, Trans. St. L. Acad. Sci., 11, p. 97, (1862).
Pecten hawni Swallow and Geinitz, Carb. u. Dyas in Neb., p. 36, pl. n,
ff. 19a, b, (1866).
Aviculopecten carboniferus Meek, Fin. Rep. U. S. Geol. Surv. Neb.,
p. 193, pl. iv, f. 8, and pl. 1x, ff. 4a, b, (1872).
Meek’s description: ‘‘Shell rather small, slightly oblique,
moderately convex, length and breadth nearly equal; hinge
line nearly or quite straight, and somewhat less than the great-
est breadth of the valves, provided with a marginal ridge in
both valves; basal margin regularly rounded. Left valve more
convex than the other; posterior ear rather well defined from
the swell of the umbo, somewhat extended and terminating in
an acute point, separated from the margin below by a deep,
rounded sinus; anterior ear about two-thirds as long as the
other, and rather more distinct from the umbo and more ob-
tuse, but still rather acutely angular, defined by a moderately
distinct subangular sinus. Right valve nearly flat, or distinctly
118 University Geological Survey of Kansas.
less convex than the other; its anterior ear narrow, and de-
fined by a deep, rather sharp sinus; posterior ear of the same
size and form as in the left valve. Surface ornamented in the
left valve with about fifteen or sixteen regular, distinct, angu-
lar, radiating plications separated by furrows of the same size,
each one of which terminates at the free border in little spine-
like projections, with curved-up margins; lines of growth fine
on the body of the valve, but becoming more distinct and irreg-
ular on the ears, where there are rarely any defined radiating
coste. Ata few distantly separated intervals there are promi-
nent imbricating lamelle of growth, showing the same digitate
margins as the free borders of the shell. In the right valve
the surface markings are somewhat like those of the other valve,
but much more obscure, excepting on the anterior wing, where
there are a few more distinct radiating coste.’’
Measurements: Height, 29 mm.; length, 22 mm.; convex-
ity, about 4mm.
Range and distribution: Upper Coal Measures; Kansas City,
Turner, Eudora, Lawrence, Lecompton, Topeka.
The strong radiating cost with the vaulted and spine-like
projections of the free margins and larger lamelle of growth |
distinguish this species from all others of the genus. When
broken from limestone, the larger lamelle of growth are gen-
erally broken away.
Aviculopecten maccoyi. Plate XIII, fig. 10.
Aviculopecten Mecoyi Meek and Hayden, Pal. Upp. Mo., p. 50, pl. un, f.
9, (1864); White, U.S. Geog. Surv. West 100 Mer., tv, p. 150, pl. x1, f.
Peal (UST):
Shell below medium size, moderately convex, subovate in
outline exclusive of the ears; hinge nearly or quite equaling
the greatest length of the shell, slightly falling from the beak,
which extends a trifle above it; posterior ear larger and more
acute than the anterior. The ventral margin is broadly
rounded ; the posterior margin is more abruptly rounded than
the lower anterior margin; anterior ear separated from the
shell by a moderately deep, rounded sinus, and from the body
of the shell by a distinct, moderately deep, rounded depression ;
BgEDE. | Carboniferous Invertebrates. 119
anterior extremity of the ear somewhat rounded; posterior ear
separated from the margin by a larger and rounder sinus, and
from the shell by a nearly equal depression with that of the
anterior one; the sides of the beak converge at an angle of
about 85°. The surface of the left valve is ornamented with ra-
diating strizw, from three to six of which are marked at intervals
with vaulted, scale-like projections, give them a rough appear-
ance; between each of these are four to eight smaller, slightly
flexuous striz, increasing by implanation ; crossing all of these
are fine, closely set, concentric strize and occasionally larger
lines of growth; anterior ear marked by radiating and fine con-
centric strie ; the radiating strize on the posterior ear are less
distinct. Measurements: Height, 21 mm.; length, 29 mm.;
convexity, 4mm.; length of the hinge line, 19mm.
Range and distribution: Upper Coal Measures; Turner,
Lawrence, Lecompton, Topeka, Cowley county.
This species can almost always be distinguished by its larger
ribs, with the vaulted scale-like elevations and smaller strize
between. Care must be taken, however, not to confound it
with Pseudomonotis, which it resembles to a considerable degree,
in an outward way, but it is much more regular and has the
ears better developed than in Pseudomonotis.
Aviculopecten providencesis. Plate XIII, fig. 2.
Pecten providencesis Cox, Geol. Surv. Ky., 111, p. 556, pl. viii, f. 1, (1857).
Aviculopecten providencesis Worthen, Geol. Surv. Ill., viii, p. 116, pl.
xx1, ff. 4, 4a, (1890).
Aviculopecten fasciculatus Keyes, Geol. Surv. Mo., v, p. 113, pl. xxu, f.
7, (1894).
Shell large, broadly subovate in outline, somewhat oblique ;
margins, except the ears, broadly and regularly rounded, the
beak extending very slightly beyond the hinge line, which is
shorter than the length of the shell; height about equal the
length. Ears large, well defined, the posterior one much more
acute than the anterior, which is somewhat rounded, and sepa-
rated from the body of the shell by a somewhat shallow, distinct
sinus; posterior ear separated from the swell of the umbo by a
sharp depression and from the margin by adistinct sinus. Sur-
120 University Geological Survey of Kansas.
face of the shell ornamented by about twenty-three radiating
cost, each consisting of three to five striw, which extend nearly
to the beak ; these bundles are separated by rather broad depres-
sionsor grooves. There are fine crowded lines of growth visible ;
they are prominent on the margins of large specimens as lam-
ellar strie. The anterior ear is ornamented by ten to fifteen
simple radiating strie and concentric lines of growth; the
posterior ear is similarly marked, except that the radiating lines
are fewer and less distinct. Measurements: Height, 69 mm. ;
length, 64 mm.; length of hinge, 40 mm.; convexity, 12 mm.
Range and distribution: Upper Coal Measures; Kansas City,
Munner
The truncation of the posterior side, nature of the fascicles,
ears and the general appearance of this shell are so similar to A.
providencesis of Cox that I do not hesitate to refer it to that
species. Worthen evidently considered it as belonging to that
species, for he borrowed a fine specimen from Mr. W. J. Parrish,
of Kansas City, to figure, in the volume above referred to, for
comparison, with his new species which he named A. chesterensis.
Mr. Parrish, so he informs me, collected the specimens figured
by Worthen from the Kansas City rocks. Some of our speci-
mens are from the same locality. It is therefore probable that
they are the same as Keyes’s A. fasciculatus, which also is from
Kansas City. Keyes, in his diagnosis, gives no characters
which distinguish his species from that of Cox. It differs from
Worthen’s species in the larger fascicles and the truncated
upper posterior extremity of the outline, as well as having a
comparatively longer hinge. Our specimens are also much
more convex than those of A. chesterensis.
The species can be easily separated from the remainder of our
Carboniferous species by,its large size and the fasciculation
of the stric.
BEEDE. | Carboniferous Invertebrates. 121
Aviculopecten hertzeri. Plate XIII, figs. 8, 1, 1b.
Aviculopecten. (Strebloplteria?) hertzeri Meek, Proc. Acad. Nat. Sci.
Phil., 1871, p. 61; Pal. Ohio, 11, p. 330, pl. xrx, ff. 13a—c, (1875).
Aviculopecten hertzeri Herrick, Bull. Den. Univ., 11, p. 25, pl. 1, ff. 5, 10.
Meek’s description: ‘‘Shell usually under medium size,
higher than wide, rather compressed, the right valve being
nearly flat, and the left only moderately convex; subovate in
general outline (exclusive of the ears), with a slight backward
obliquity, caused by the greater prominence of the anterior
margin ; basal outline.semicircular and rounded regularly into
the rather prominently rounded anterior margin; posterior
margin less prominent than the anterior, and forming a longer
and more or less gentle curve from the posterior ear into the
base; hinge distinctly shorter than the antero-posterior di-
ameter of the valves; posterior ear in both valves very small,
flattened, very obtusely angular, and much shorter than the
margin below from which it is only separated by a slight sinu-
osity, though it is well defined from the umbo; anterior ear of
each valve distinctly larger than the posterior, though not
nearly so prominent as the anterior margin below, rather
strongly compressed or flattened, so as to be abruptly separated
from the umbo, and in both valves defined by a distinct sinus.
from the margin below, the sinus being deeper and more angu-
lar in the right valve; beaks compressed, scarcely projecting
above the cardinal margin, and placed a little behind the middle
of the hinge, as well as that of the valves. Surface of both
valves elegantly ornamented by numerous, sometimes sharply
elevated, nearly equal, very regularly arranged radiating and
concentric lines, which are larger and more strongly defined on
the anterior ear of the right valve, particularly the radiating
markings, which there sometimes assume the character of small
coste, while the concentric markings there in some examples
project as little lamelle above the hinge margin, so as to give
it a subtended appearance. Height of one of the largest speci-
mens seen, 1.32 inches; antero-posterior diameter, 1.20 inches ;
convexity, about 0.18 inch.’’
9—Vvi
122 University Geological Survey of Kansas.
Range and distribution: Upper Coal Measures; Topeka. -
From the horizon of the Osage coal.
The fine, regular, cancellated strie and the larger strive: on-
the anterior ear distinguish this species from any of the pre-
ceding. We have a single specimen from Topeka which seems
to agree very well with this species.
Aviculopecten sculptilis. Plate XIII, figs. 3, 3b.
Aviculopecten sculptilis Miller, 17th Ann. Rep. St. Geol. Ind., p. 702, pl.
Oia Dea The SPAN
Shell a little large, somewhat oblique, ovate in outline, taper-
ing to the beak at an angle of 90° or less, very moderately con-
vex; beak extending to or a trifle beyond the hinge line, and
located back of the middle of it. The ears are small, the
anterior flattened, and separated from the swell! of the umbo by
a deep depression, and considerably larger than the posterior
ear; the margin of the ear makes a deep sinus at its junction
with the margin of the shell; however, the anterior border of
the shell generally extends beyond the ear; posterior ear small,
well defined, margin not separated from the margin of the shell
by a very distinct sinus; the posterior is about two-thirds as
long as the anterior. Both ears (in the left valve) are marked
by small, well defined, concentric striz. Entire border below
the ears regulariy rounded and subsemicircular. The shell is
marked by numerous, closely set radiating strie, which are
crossed by about equally prominent closely set concentric striae,
giving the shell a beautifully cancellated appearance ; the radi-
ating striae increase by implanation. At the anterior border
the shell often becomes quite rough in appearance on account
of the enlarging of the striec. The anterior ear seems to be
crossed by a few indistinct radiating striz. According to Mil-
ler, the anterior (‘‘ posterior ’’ ) ear of the right valve is marked
by more prominent radiating strize which ‘‘ give it a strongly
pitted aspect.’’ Measurements: Height, 49mm.; length, 40
mm.; length of hinge, 23 mm.; convexity, about 8 mm.
Range and distribution: Upper Coal Measures; Kansas City.
Taken from the oolite.
>. af eee nae
BgEDE. | Carboniferous Invertebrates. 123
This species differs from A. hertzeri Meek in being much
larger, in having a much more (20°) acute beak, which makes
it amore slender shell. Aside from the species just mentioned,
it can easily be distinguished from our other Coal Measures
species by its strongly reticulated surface.
The specimen at hand agrees very well with Miller’s descrip-
tion and figure, though there are a few minor differences, but
both are from the same locality, so there can be but little doubt
of their identity.
Aviculopecten germanus. Plate XIII, fig. 4.
Aviculopecten germanus Miller and Faber, Jour. Cinn. Soc. Nat. Hist.,
Xv, pp. 81, 82, pl. 1, f. 9, (1892.)
Original description: ‘‘ Shell small, a little higher than long ;
inequilateral; oblique; base regularly rounded; antero-basal
and postero-basal margins rounded. Hinge oblique, nearly
straight, not quite equaling the greatest length of the valves
below. Posterior ear extends to the lateral border, with which
it forms nearly a right angle; it graduates into the shell below
without the presence of a sinus. Anterior ear rather shorter
than the posterior one, angular at the extremity and rounding
on the margin below into a notch, and arching from the wing
into a deep and distinct sinus that separates it from the poste-
rior margin of the shell. Umbo tumid and umbonal slopes
diverging to the margin. Beak high, pointed and projecting
beyond the cardinal margin a little forward of the middle of
the hinge line. There are two cost on the posterior ear, and
three or more finer ones on the anterior ear. There are about
twelve principal radiating ribs on the body of the shell and
about as many rudimentary and intercalated shorter ribs be-
tween them; they are separated by wider flattened spaces. It
is difficult to tell from our specimens whether or not there were
any concentric lines, though they are quite well preserved; if
there were such lines they were very fine.
‘«This species approaches more nearly to A. rectilaterarius than
to any other, but it may be readily distinguished by the scarcity
of the radiating ribs and the wide flattened spaces between
124 University Geological Survey of Kansas.
them; the beak, too, is higher and more pointed, as well as
having a more tumid umbo.”’
Range and distribution: Upper Coal Measures; Lawrence.
Aviculopecten coxanus. Plate XIX, fig. 2.
Aviculopecten coxanus Meek and Worthen, Proc. Acad. Nat. Sci. Phil.,
p. 453, (1860); Meek, U.S. Geol. Surv. Neb., p. 196, pl. rx, ff. 6a,b, (1872).
Meek’s description: ‘‘Shell very small; thin, compressed,
slightly oblique; broad subovate exclusive of the ears; basal
margin rounded; anterior margin more or less rounded, rather
straight and oblique above; posterior margin more prominent
than the anterior, often subangular at the point where the pos-
tero-basal margin rounds up to meet the obliquely sloping edge
above. Hinge generally a little less than the greatest breadth
of the valves below. Left valve with anterior ear of moderate
size, flat, triangular, with the extremity generally a little less
than a right angle, sometimes very slightly rounded, separated
from the margin below by.an abruptly rounded or subangular
sinus; posterior ear slightly larger and much more acutely
angular than the other, but shorter than the most prominent
part of the margin below, from which it is separated by a mod-
erately deep rather broadly rounded sinus; beak small, com-
pressed, scarcely projecting beyond the cardinal margin, and
placed a little in advance of the middle of the hinge; surface
ornamented with linear, simple, often more or less flexuous
cost, which alternate in size, the smaller ones dying out at
varying distances between the free margins and the umbo—
crossing all of these are numerous, extremely fine, regular,
closely arranged concentric strie, which, like the costz, are
more or less distinctly defined on the ears, as well as on the
body of the valve. Right valve unknown.”’
Range and distribution: Upper Coal Measures ; Lecompton.
A single specimen of a left valve of this species agreeing very
well indeed with Meek’s description and figure was collected in
the shale underlying the Upper Oread limestone at Lecompton.
MPR ad 2
wees pee |
\e ey
Ne ‘ay be Nt Shee ee OR, *, ‘Ve? . «
>. - 7 A E
BREDE. | Carboniferous Invertebrates. 125
PTERIA.
Pteria longa. Plate XVI, fig. 4.
Gervillia longa Geinitz, Carb. u. Dyas in Neb., p. 32, pl. 1, f, 15, (1866).
Avicula longa Meek, Fin. Rep. U.S. Geol. Surv. Neb., p 199, pl. rx, f. 8,
(1872); Meek and Worthen, Geol. Surv. Ill., v, p. 578, pl. xxv, f. 1,
(1873); ete.
Meek’s description: ‘‘Shell nearly or quite equivalve ; body
part obliquely elongated and more or less arcuate ; posterior
end narrow and abruptly rounded; base nearly straight and
parallel to the cardinal margin behind, but ascending obliquely
forward from near the middle of the valves; anterior side ob-
lique, and broadly and faintly sinuous under the ear. Hinge
line about three-fourths the length of the valves, and provided
with a marginal ridge, produced behind into a very narrow,
elongated ear, considerably shorter than the oblique portion of
the valves, from which it is separated by a deep sinus which
narrows to an abruptly rounded or subangular extremity close
under the ear; anterior ear shorter and much broader than the
other, in the left valve convex, with its extremity pointed, and
faintly sinuous just below the point—separated from the swell
of the umbo by an oblique sulcus extending from the anterior
side of the same to the back part of the broad, shallow marginal
sinus of the ear. Beaks of both valves convex, very oblique,
placed one-fourth to one-fifth the length of the hinge back of
the anterior extremity ; in the right valve, rising a little above
the hinge, but in the left somewhat more prominent, according
to Professor Geinitz’s figures. Length of medium-sized speci-
men, measuring obliquely from the extremity of the anterior
ear to the posterior end of the body part of the valves, 9.61
inch; height, measuring at right angles to the hinge, 9.33 inch ;
length of hinge, about 9.23 inch.’’
Range and distribution: Upper Coal Measures; Lawrence,
Topeka.
126 University Geological Survey of Kansas.
Pteria sulcata. Plate XVI, fig. 3.
Gervillia suleata Geinitz, Carb. u. Dyas in Neb., p. 33, pl. 11, f. 16, (1866).
Avicula? suleata Meek, Fin. Rep. U.S. Geol. Surv. Neb., p. 200, pl. rx, f.
9, (1872).
All the specimens before me having the ears either broken off
or not visible, Meek’s description of Geinitz’s figure is given,
with remarks on the specimens at hand. ‘‘ Rhombic oblique
shell, with short, compressed, triangular anterior ear, defined
by a faint, wide marginal sinus; and a larger compressed, some-
what elate, posterior ear, with a marginal ridge, showing a
tendency to be produced into a narrow appendage behind, sepa-
rated from the margin below by a rather deep, rounded sinus.
The posterior basal extremity is rather narrowly rounded, but
not much produced; while the outline of the base is broadly
semielliptical, and the umbo convex, and rising somewhat above
the hinge line, which is apparently shorter than the greatest
length of the valve. Surface ornamented by fine lines of growth,
and before by two or three sulci, extending from the anterior
side of the beak to the antero-basal margin, leaving ridges be-
tween which are more or less crenated by the crossing of the
marks of growth; just behind the posterior one of these sulci,
the margin of the latter is ornamented with regular, rather
strongly defined wrinkles, or little folds, some of which are pro-
longed backward parallel to the lines of growth.”’
Range and distribution: Upper Coal Measures; Kansas City,
Lawrence. | ,
Most of our specimens are younger than the one figured by
Meek and Geinitz, and, as a result, somewhat less ornamented.
The ornamentation is varied somewhat in different individuals. .
In some the ridge between the sulci is very crenate, and less so
in the others. It may easily be distinguished from the other
species by the ornamentation of this part of the shell.
BEEDE. | Carboniferous Invertebrates. 127
LIMOPTERIA.
Meek and Worthen, Proc. Chic. Acad. Sci., I, p. 29, (1866).
Limopteria longispina. Plate XVI, fig 6.
Gervillia longispina Cox, Geol. Rep. Ky., 111, p. 568, pl. viri, f. 6, (1857).
Monopteria longispina Keyes, Geol. Surv. Mo., v, p. 114, pl. xxii, f. 1,
(1894).
Shell elliptical-subquadrate in outline; antero-dorsal margin
nearly straight to the anterior margin, which is nearly circular,
the curve rapidly decreasing again to the extremity of the shell,
where it is sharply curved, almost angular; the posterior mar-
gin is a deep U-shaped sinus between the acute point of the ear
and the extremity of the shell. Beak placed well forward; shell
very oblique; umbonal ridge very prominent, extending to the
extremity of the shell, greatest curve about one-third the distance
from the beak, which protrudes slightly beyond the hinge line.
Anterior portion of the shell quite convex. The deflection of
the margin anterior to the beak, forming the lunule, is very
sharp, almost angular, and extends backward some distance.
The internal muscular impressions very faint, as excellently
preserved casts show practically no traces of them. Spine acute,
about three-fourths the length of the shell. The posterior ear
is separated from the shell by a moderately well-defined depres-
sion, which falls directly from the umbonal ridge, which is very
prominent. Surface marked by concentric strive of growth
which are nearly parallel to the outline of the shell.
Range and distribution: Upper Coal Measures ; Kansas City,
Turner, Lawrence.
Some specimens from Lawrence are extremely long when
measured from the tip of the ear to the anterior margin. The
great length of the ear, with the convexity of this portion of
the shell, are its principal features.
128 Uriversity Geological Survey of Kansas.
Limopteria marian. Plate XVI, figs. 5-5e.
Monopteria marian White, Prelim. Rep. Inv. Foss., p. 22, (1874); U.S
Geog. Surv. West 100 Mer., rv, p. 151, pl. x1, ff. 4a—c, (1877).
White’s, description (with additions): ‘‘ Shell of moderate
size, slender, nearly or quite equivalve, narrow and much ex-
tended posteriorly, the curvature being much the greater in
the anterior half of the shell, the posterior half being nearly
straight; body of the shell gradually tapering to near the pos-
terior end, which is abruptly rounded; a more or less promi-
nent ridge, which is sometimes in part raised as a distinct
carina, extends along the middle of the body of each valve
from the beak to the posterior end; from this carina, or angle,
the sides slope abruptly to both the inferior and upper borders,
so that a cross-section of the shell behind the ear would have a
rhomboidal outline; beaks moderately prominent, separate ;
hinge equal in length to about one-half the full length of the
shell, and its direction is nearly parallel with the posterior
half of the body; posterior wing well developed, not sharply
defined’ from the body by an auricular furrow; its cardinal
portion narrow and moderately extended; anterior ear obso-
lete; lunule moderately large and deep, the borders of which
are slightly prominent laterally, but its margins sharply
rounded inward. Surface smooth in aspect, but it is marked
by very numerous fine lines of growth. Length, from the front
to the posterior extremity, about 4cm.; height, from the base
to hinge margin, 18 mm.; average width of the body of the
shell) about 1 em, ~
We have a left valve, probably of a large specimen of this
species, showing part of the muscular impressions. The ad-
ductor is located behind the umbonal ridge, just above and in
front of the marginal sinus formed by the ear and the body of
the shell. From the lower side of the impressions a slight line
runs directly forward across the umbonal ridge, then turns
abruptly and runs toward the anterior end of the lunule, but
fades out at about two-thirds of the distance. In front of the
scar, and crossing the front edge of it, is a line at right angles
to the first, beginning a little above the scar and fading out
BgEDE. | Carboniferous Invertebrates. 129
about half way to the lower end; beyond the line appears to
be a very indistinct furrow near to the ventral margin. Ina
cast of what appears to be a right valve of another specimen
the adductor impression is shown, located nearly opposite to
that of the left as described above. It is situated mostly be-
hind the umbonal ridge but extends over it, is deeper and
roundly ovate. The pallial line extends from the lower side
of the impression a little forward and then upward, reaching to
the anterior edge of the beak. The umbonal ridge appears to
be much less sharp in this specimen than in the other.
Range and distribution: Upper Coal Measures; Turner,
Topeka.
This species may be easily distinguished from the foregoing
by its more slender form, more gently curving front, and less
spinous ear.
Limopteria gibbosa. Plate XVI, fig. 9.
Pterinia(Monopteria) gibbosa Meek and Worthen, Geol. Surv. IIl., 1,
p. 330, pl. xxvu, ff. 11-11b, (1866).
Monopteria gibhosa White, 13th Ann. Rep. Ind. St. Geol., p. 139, pl. xxx,
ff. 11, 12, (1884).
Monopteria ( Pterinia) gibbosa Heilprin, 2d Geol. Surv. Penn., Ann. Rep.
1885, p. 455, f. lla, p. 444, f. 11, (1886).
White’s description: ‘‘ Shell, exclusive of the wing and pos-
terior prominence, irregularly suborbicular in marginal out-
line; the valves moderately convex or a little gibbous; the
anterior and basal margins forming an almost regular semicir-
cular curve; posterior portion of the shell produced, narrow
and narrowly rounded, or subangular, at the extremity; wing
slender, compressed and extending backward as far as the nar-
row extremity, between which the posterior margin forms a
deep, broad notch, that is narrowly rounded at the bottom ;
umbonal ridge moderately distinct; beaks equal, not so far
forward as the front margin of the shell; anterior lunule deep ;
cardinal border not so long as the full diameter of the shell;
surface marked only by the ordinary lines of growth. Length,
from posterior extremity to front, 27 mm.; height, from base
to beaks, 23 mm.’’
130 University Geological Survey of Kansas.
Range and distribution : Upper Coal Measures; Kansas City,
Turner, Lawrence.
The short, gibbous shell with a moderately long spine sepa-
rates this species from the rest of the genus. :
Limopteria alata. Plate V, fig. 5.
Monopteria gibbosa alata Beede, Kans. Univ. Quart., vir, p. 189, f. 5,
(1898).
Shell small, extremely thin, laterally compressed; beak ex-
tending a trifle beyond the hinge line, prominent on account of
the lunule, but not much elevated, and placed well back for a
member of this genus ; umbonal ridge less prominent than in any
other species of the genus and less curved. Posterior ear very
greatly developed, about equaling the entire body of the shell
in area. Antero-dorsal margin sinuate on account of the turn-
ing down of the margin to form the lunule; anterior margin
circular nearly to the postero-ventral extremity of the shell,
which is acute; the posterior margin consists of a broad, shal-
low sinus, extending from the postero-ventral end to the point
of the ear, which is apparently rounded and obtuse. The ear is
not separated from the shell by a distinct depression, but slopes
gradually from the umbonal swell, save at the extreme lower
edge, where the depression is more abrupt. Very fine con-
centric lines of growth are visible, all of which pass around the
shell with a double curve to the ear, where they again curve
backward, and then forward to the hinge line. Length, 20
mm.; depth, 18 mm.; convexity of single valve, a trifle less
than 4 mm.
Range and distribution: Upper Coal Measures; Turner,
Lawrence. ;
This shell differs from L. gibbosa M. and W., to which it is
most closely related in some respects, in being much less gib-
bous, the ear much larger and more obtuse, antero dorsal out-
line more sinuous, umbonal ridge more nearly straight and less
prominent, beak placed farther back, and the depression sepa-
rating the ear from the umbonal ridge more shallow.
> By One ig Seal gl ee ee ae
B&EDE. | Carboniferous Invertebrates. 131
Limopteria subalata. Plate XXI, figs. 3a, 3b.
Monopteria ? subalata Beede and Rogers, Kans. Univ. Quart., virr, p.
133, pl. xxxtv, ff. 3a, 3b, (1899).
Shell moderateiy small, subcrescentic in outline, gibbous, a
little longer than high; valves subequal, but beak prominent,
slightly projecting, somewhat inflated, situated about one-fifth
the length of the shell from the anterior margin, which is trun-
cated, on account of the lunule which is formed by the turning
in of theshell. The anterior margin is convex below. Ventral
margin broadly rounded to the postero-ventral extremity, where
it is abruptly rounded to meet the concave posterior. Hinge
short and straight; posterior ear but slightly developed; um-
bonal ridge prominent, somewhat sickle-shaped, sloping abruptly
posteriorly, forming a cavity broadly convex anteriorly. Sur-
face marked by moderately distinct lines of growth. Length,
16 mm.; height, 14 mm.; convexity of single valve, 4 mm.
Range and distribution: Upper Coal Measures; Lawrence,
Topeka.
At two localities in the lower part of the Upper Coal Meas-
ures are found great numbers of Limopteria. Beside the three
. large species are found two smaller species, L. alata and L. sub-
alata; one with an exceedingly large wing and compressed shell,
and the other with a gibbous shell and the wing almost want-
ing. There are many variations of the two species and they
seem to grade into the other species to some extent, but differ
sufficiently, so far as we can observe, from even the young of
those, to be considered as distinct.
The size of the shell and the exceedingly small ear easily
separate L. subalata from any other member of the genus.
132 University Geological Survey of Kansas.
PSEUDOMONOTIS.
Beyrich, Zeitsch. der Deutsch. Geol. Gesellsch., XIV, (1862).
Pseudomonotis hawni. Plate XIII, figs. 11-llc; plate XV, figs. 1-1f, 2, 2a.
Monotis hawnit Meek and Hayden, Trans. Alb. Inst., rv, p. 76, (1858);
Proc. Acad. Nat. Sci. Phil. 1859, p. 28.
Eumicrotus hawni Meek and Hayden, Pal. Upp. Mo., p. 54, pl. 1, ff.
5a—-c, (1864); White, 13th Ann. Rep. Ind. St. Geol., p. 142, pl. xxx,
f. 10, (1884); ete.
Meek and MHayden’s description: ‘‘Shell subcircular, or
subovate; hinge straight, equaling about half the length of
the valves; beaks subcentral, short, not oblique; ears nearly
obsolete; base rounded; antero-ventral and postero-ventral
margins rounded, the latter being somewhat more prominently
rounded than the other. Left valve convex; anterior mar-
gin sometimes slightly sinuous near the hinge above; poste-
rior margin intersecting the hinge at an obtuse angle; beak
convex, extending but little beyond the hinge line. Right valve
nearly or quite flat; beak flat, not projecting beyond the hinge ;
byssal sinus narrow, deep, or extending back parallel to the hinge
to a point nearly under the beak. Surface of both valves, par-
ticularly the left one, ornamented by more or less distinct radi-
ating cost, which are usually separated by spaces three or four.
times their own breadth, and armed with regularly disposed
vaulted, spinelike prominences, formed apparently from the pro-
jecting laminez of growth. Between each two of the principal
radiating costae from one to three or four much smaller radiat-
ing ribs or lines are usually seen, crossed by obscure concen-
tric markings. Hinge and muscular impressions unknown.
Length, 1.47 inches; height, 1.42 inches; convexity, about 0.40
OGIO, : .
Range and distribution: Upper Coal Measures; Turner, Law-
rence, near Topeka.
To the above synonyms should be added, I think, P. radials
Meek, described in the Nebraska report. From his description
and figures, it seems to agree more closely with P. hawnz than
with any other form.
BEEDE. | Carboniferous Invertebrates. 138
Pseudomonotis kansasensis, nom. nov. Plate XIV, figs. 1-ld.
Pseudomonotis tenuistriata sp.? var.? Beede, Kans. Univ. Quart., vii,
p. 81, pl. xviii, ff. 1-1d, (1899).8
Shell large, ovate in outline, rather compressed ; beak moder-
ately prominent, projecting beyond the hinge, which is nearly
straight. Anterior ear small, rounded to meet the hinge, rather
flat, the shell rising rather abruptly to the swell of the umbo.
Anterior margin slightly sinuate ; antero-ventral margin broadly
rounded to the ventral portion of the shell, where it becomes
nearly straight, then rounding more abruptly to the posterior
ear, which is also rounded to the hinge. Greatest convexity a
trifle below the beak, but it is very slight. The surface is
marked by many fine, wavy, radiating strice of uniform size, ex-
tending from the beak to the ventral margin; occasionally one
striation will be a trifle larger than another on the central por-
tion of the shell, but it soon loses itself, and on old individuals
the striz on the ventral border are all about equal. They in-
crease by implantation and are rather sharply defined, sepa-
rated by troughs from one to three times their width, and are
generally crossed by fine concentric lines or lamine. Right
valve unknown. Height, 62 mm.; length, 69mm. ;” length of
hinge, 25 mm.; convexity, 10 mm.
Range and distribution: Upper Coal Measures; Turner, To-
peka, Auburn (Shawnee county). This species differs from
P. hawni in always having small, regular striz and shorter
hinge in the large individuals, as well as being a larger species.
Pseudomonotis? robusta. Plate XIV, figs. 2-2c.
Pseudomonotis? robusta sp.? var.? Beede, Kans. Univ. Quart., vri1, p.
82, pl. xviii, ff. 2-2c, (1899).
This shell differs from the preceding in being much more con-
vex and arcuate, in having a longer hinge, higher umbo, beak
very much more compressed and scarcely distinct from the
8. Through the kindness of Mr. Charles Schuchert, my attention was called to the description
of a 2 ies of this genus under the name tenuistriata, by Mr. Bittner. The paper was pub-
lished in the Jahrbuch der K. K. Geolog. Reichsanstalt, xlix, heftsiii and iv, (1899). The species
is from the Triassic of central Asia. As the paper probably appeared before my article, the
—— of the Kansas species will have to be changed. Aansasensis is proposed as the new name
of the species.
9. The specimen used as the type is somewhat crushed on the posterior end, making the shell
eee longer than it really is. The length given here is that of the specimen in its crushed
condition.
134 University Geological Survey of Kansas.
umbo, not projecting very sensibly above the hinge. The strize
are more regular and much fainter, and either very indistinct
or absent on at least the upper third of the shell. Both concen-
tric wrinkles and lamelle of growth are distinct. Length, 48
mm.; convexity, 18 mm.; height, 42 mm.; length of hinge
about 28 mm. !
Range and distribution : Upper Coal Measures; Turner.
This species differs from P. hawni in being very arcuate, hav-
ing a plain umbo, and full anterior and posterior marginal
outlines, and fine, even striz on the margins. It differs from
P. kansasensis in its broad, smooth umbo and indistinct beak,
long hinge, and more circular outline. |
Pseudomonotis hawni equistriata. Plate XIV, figs. 3-3b.
Pseudomonotis hawni equistriata Beede, Kans. Univ. Quart., O. 82. pl.
xvitl, ff. 3-3b, (1899).
Shell of medium size, ovate in outline, moderately to quite
gibbous, a little oblique with respect to the hinge; beak moder-
ately prominent, extending to or a little beyond the hinge, which
is about half the length of the shell and somewhat arcuate.
Umbo quite gibbous. Posterior ear very slightly developed,
merging into and forming a slight sinus in the posterior mar-
gin; ventral, antero- and postero-ventral margins regularly
rounded ; anterior margin sinuate in the upper portion on ac-
count of the anterior ear, which is small and round. ‘The sur-
face is marked by fine, somewhat regular, rather wavy strie,
which increase by intercalation, each fourth to tenth being
usually a little larger than the remainder, though not very con-
spicuously so. Small lamelle of growth sometimes distinguish-
able. Some of the striz extend nearly to the beak. The right
valve is flat, or a little concave; otherwise unknown. Measure-
ments, two specimens: Height, 51 mm., 34 mm.; length, 24
mm., 26 mm.; convexity, 7 mm., 18 mm.; length of hinge, 12
mm., 16mm. These two specimens represent the extremes of
convexity.
Range and distribution : Upper Coal Measures; Turner.
This variety differs from P. hawni in being a shorter shell and
awry he
ee 5
ar
BEEDE. | Carboniferous Invertebrates. 135
a little more convex, having regular striw, and in being a little
smaller. P.cf. hawni, in the article above referred to, should also
be considered as a true member of the species. The species here
separated are, I believe, distinct from P. hawni; and this variety
is worthy of varietal distinction, as often shells of these kinds are
found in localities where the others are absent and some method
of distinguishing the two forms of the species is necessary.
POSIDONOMYA.
Bronn, Leth. Geogn., (1837).
Posidonomya? recurva. Plate XIX, figs. 6-Ge.
Posidonomya ? recurva Beede, Kans. Univ. Quart., vi11, p. 126, pl. xxx,
f. 6, (1899).
Shell of medium size, lenticular, oblique, and thin. The hinge
line is nearly straight, about two-thirds the length of the shell.
The beak is moderately prominent, recurved, projecting very
slightly beyond the hinge. The greatest convexity is on the
upper half of the shell and constitutes the umbonal swell, which
is moderately prominent and curved backward, making the
shell oblique. The surface is marked by concentric undula-
tions of growth, and fine, closely set, concentric strie. Height,
23 mm.; length, 23 mm.; convexity of valve, about 4 mm.
Range and distribution: Upper Coal Measures; Lawrence.
A specimen from the dam, at Lawrence, and shown in figs.
6b, 6c, Plate XIX, is probably of the same species, though it has
a somewhat different appearance and outline. It shows the
beak projecting beyond the hinge, and the cast of the interior
shows the beak to have been hollow beyond the hinge. The
cast figured is of both valves in place, one of which is slightly
crushed. It may be a distinct species, though as it is from the
same horizon it is probable that the difference is individual
rather than specific.
The character of the hinge and the internal markings are too
poorly shown in our specimens to permit of accurate location
of the species. It resembles very much in appearance species of
the genus Posidonomya, to which it is provisionally referred, and
with which it agrees in its surface markings and in having a
136 University Geological Survey of Kansas.
very thin shell. It also agrees very well in these respects with
Paracyclas, but that genus is not at present known from the
Carboniferous.
Posidonomya? pertenuis. Plate XIX, fig. 5.
Posidonomya? pertenuis Beede, Kans. Univ. Quart., vr, p. 127, pl. xxx,
f. 5, (1899).
Shell a little larger than in the, previous species and less
oblique. Transversely ovate in outline, very thin, quite com-
pressed. Hinge line is nearly straight, about equaling half the
length of the shell. The posterior? margin is somewhat trun-
cate and nearly straight; anterior extension of the hinge longer
than the posterior, rather flat, not separated from the shell by
well-defined depression. ‘The front and ventral margins regu-
larly rounded. The shell is compressed, probably most convex
near the middle; beak obtuse, not very prominent, protruding
above the hinge line. The surface is marked by concentric un-
dulations of growth, and fine, close, concentric striz. Height,
40mm.; length, 36 mm.; convexity of single valve, 4 or 5mm.
Range and distribution: Upper Coal Measures; Lawrence.
This species belongs to the same genus as the preceding. It
has some resemblance to the figure of Keyes,” which he refers
to Placunopsis carbonaria, though it is very difficult to see why
it should be referred to that genus or species; his species may
be the same as the one here described.
MODIOLA.
Lamarck, Hist. Nat. les Anim. sans Vert., (1801).
Modiola subelliptica.
Clidophorus (Pleurophorus) occidentalis Geinitz, Carb. u. Dyas in
Neb%, p--28, pl. 11, £216.) ( 1866)).
Modiola? subelliptica Meek, Fin. Rep. U. S. Geol. Surv Neb., 211, pl.
Kshs oe MOMeye
Meek’s description: ‘‘ Shell narrow, subelliptical, rather con-
vex, extremely thin, usually a little more than twice as long as
high; basal margin nearly straight, or sometimes very slightly
convex or sinuous near the middle, rounding up at each ex-
10. Geol. Surv. Mo., V, p. 108, pl. xu111, fig. 9.
BEEDE. | | Carboniferous Invertebrates. 7 137
tremity ; anterior margin narrowly rounded ; posterior extremity
more compressed, and more broadly rounded, sometimes a little
oblique above; cardinal margin somewhat straightened along
the middle, but rounding imperceptibly into the anterior and
posterior extremities ; beaks much depressed; or scarcely dis-
tinct from the cardinal margin, moderately convex, and placed
very near the anterior margin, but not terminal; umbonal slopes
forming a very obscure narrow ridge, which extends, with a
slight curve from each umbo, to the posterior basal margin.
Surface marked with moderately distinct lines of growth, which
on the posterior dorsal region above the umbonal ridge are
crossed by very minute or microscopic radiating and rather dis-
tinctly divericating striz. Length of the largest specimen seen,
1.05 inches; height of the same, 1.43 inches; convexity, about
0.25 inch.”’
Range and distribution: Upper Coal Measures; Topeka.
MYALINA.
de Koninck, Dese. Anim. Foss. Carb. Belg., p. 125, (1844).
Myalina swallovi. Plate XVI, fig. 7.
Myalina swallovi McChesney, Desc. New Pal. Foss., p. 57, (1860);
Meek, Fin. Rep. U.S. Geol. Surv. Neb., p. 201, pl. rx, ff. 7a, b, (1872);
ete.
Aucella hausmani Geinitz, Carb. u. Dyas. in Neb., p. 25, pl. 1, f. 8,
(1866).
Meek’s description: ‘‘Shell rather small, nearly or quite
equivalve, modioliform or mytiloid, convex, or even subangu-
lar, along the umbonal slopes from the beak to the anterior
basal margin; posterior and postero-dorsal regions, cuneate ;
cardinal margin nearly straight, and about half the length of
the shell, passing almost imperceptibly or without any angu-
larity into the posterior margin, which rounds down with a
semicircular curve to the narrowly rounded basal extremity ;
antero-basal margin ascending obliquely forward, more or less
sinuous near the middle or sometimes a little above, usually
swelling out into kind of lobe or protuberance above the middle
in front of the umbonal slope, as in Modiola. This prominence
sometimes extends a little beyond the beaks, and varies more
10—vi
138 University Geological Survey of Kansas.
or less in breadth. Beaks small, very oblique, not projecting
beyond the cardinal margin, and located so near the anterior ©
extremity as often to appear very nearly terminal. Surface
rather smooth, but showing fine concentric lines, which in well-
preserved specimens are sometimes crossed by very fine, ob-
scure traces or radiating stric that curve upward on the
posterior dorsal region.’’
Range and distribution: Upper Coal Measures; Kansas City,
Topeka, Turner, Lawrence.
Myalina subquadrata. Plate XVI, figs. 10, 10b.
Myatlina subquadrata Shumard, Geol. Surv. Mo., p. 207, pl. C, f. 17,
(1855); Meek, Fin. Rep. U.S. Geol. Surv. Neb., 202, pl. iv, f. 12, pl.
Ix, f. 6 (1872); ete.
Meek’s description: ‘‘ Shell large and thick, oblong or sub-
quadrate, the height being greater than the antero-posterior
diameter; right valve nearly flat; left convex, both somewhat
compressed and alate above and behind the umbonal promi-
nence. Hinge line nearly straight, about equaling the greatest
breadth of the valve, and ranging at right angles to the vertical
axis; the basal margin regularly rounded; posterior margin
nearly vertical, rounding into the base below, a little sinuous
above the middle, and intersecting the hinge above at very
nearly right angles; anterior margin thickened within, round-
ing into base, thence rising nearly vertically with a broadly
rounded concavity mainly above the middle. Beaks terminal
and directed forward. Cardinal plate or area usually rather
broad, with cartilage furrows distinctly defined. Surface of
the left valve marked with fine concentric strize, and stronger
imbricating lamelle of growth. These markings are much less
distinct on the right valves.’’
Range and distribution: Kansas City, Melvern (Osage
county), Topeka, Lawrence.
This species can readily be distinguished from the previous
by its much larger size, by the hinge forming about aright angle
with the vertical axis, and by the sinuosity or at least straight
manner in which the margin approaches the posterior end of the
hinge.
BgEDE. | Carboniferous Invertebrates. 139
Myalina ampla. Plate XVI, figs. 1, 1b.
Myalina ampla Meek and Hayden, Pal. Upp.|Mo., p. 33, ff. A, B, (1864).
Myalina subquadrata Meek and Hayden, ibid., p. 32.
Meek and Hayden’s description: ‘‘ Shell attaining a large size,
compressed and somewhat alate in the postero-dorsal region, and
convex anteriorly—considerably higher than long; posterior
margin forming a broad gentle curve, being nearly straight and
ranging almost vertically near the middle, and curying forward
so as to intersect the hinge at an obtuse, undefined angle above,
while below it arches regularly forward into the rather narrowly
rounded base; anterior margin thickened within above, broadly
arcuate or concave in outline, its curvature being nearly paral-
lel to that of the posterior margin. Beaks terminal, directed
forward; umbonal ridge most prominent and oblique above,
and in adult shells curving downwards so as to range nearly
vertically near the middle. Hinge line straight, and ranging
nearly at right angles to the longer or vertical axis of the
valves ; cartilage furrows distinct, straight, and in mature shells
numbering about ten or twelve; area broad. Height, about 4
inches; antero-posterior diameter at the middle, 2.40 inches;
convexity of a left valve, 0.83 inch.’’
Range and distribution: Upper Coal Measures; the type
specimen was collected at Leavenworth.
There is one specimen in the collection from the type locality
which probably belongs to this species, the only difference
being that the lines of growth do not bend quite so sharply
forward as in the ones figured by Meek. The principal differ-
ences between this shell and M. subquadrata are: Very convex
posterior outline instead of a straight or concave outline, greater
size, and the curving forward of the lines of growth.
Meek was in doubt as to whether it was the same as JM. sub-
quadrata of Shumard ornot. He proposes the name at the close
of the description of the shell, and comments upon it, stating
that he is of the opinion that it is a distinct species. Later, in
the final report on Nebraska, he states, in the discussion of J.
subquadrata (page 203) : ‘‘The large species figured by Doctor
Hayden and the writer on page 53 of the Paleontology of the
140 University Geological Survey of Kansas.
Upper Missouri as WM. subquadrata is, as we then suspected, a
distinct species, differing in having the posterior margin round-
ing forward into the hinge above, and not sinuous or meeting
the hinge at right angles, as in I. subquadrata. It will have to
take the name MM. ampla, suggested by us for it, in case it should
be found distinct.”’
I have collected a large specimen of this species near Topeka
with both valves present, but separated.
Myalina kansasensis. Plate XVI, fig. 11.
Myalina kansasensis Shumard, Trans. St. L. Acad. Sci., 1, p. 213, (1858);
Keyes, Geol. Surv. Mo., v, p. 117, pl. xuii, f. 5, (1894).
Original description: ‘‘Shell sub-rhomboidal, sub-inequi-
valve, inequilateral, gibbous, the left valve more gibbous than
the right; height about double the length; in young specimens
the greatest length is at the cardinal border, but in the adult
toward the pallial margin; cardinal margin oblique, slightly
arched, and forming with the posterior border an angle of
about 120°; posterior margin rather strongly arched in adult
specimens, and very gently rounded in the young; pallial mar-
gin rounded; buccal margin concave ; umbones very prominent
anteriorly, and declining with a moderate slope to the posterior
margin; anterior slope very abrupt; beaks terminal, attenu-
ated, directed obliquely forward, incurved ; surface with strong,
imbricating, sub-equidistant, concentric lamellae, whose free
edges are often irregularly crenate; lamellae: most prominent
on the left valve. The ligament face is broad, marked with
equidistant, close, deeply impressed lines parallel to the cardi-
nal edge, the number varying with the age of the shell; be-
neath these is a rather broad, smooth space, which is continuous
with a similar space extending from the pallial region. Hach
valve exhibits a singular muscular impression, which is large,
ovate, and situated towards the posterior margin. Height, 23
inches; length, 1.17 inches; thickness, 0.88 inch.
Range and distribution: Upper Coal Measures; Argentine,
Cowley county, Neosho river near Council Grove.
This species may be easily recognized by the arched lamellez
BEEDE. | Carboniferous Invertebrates. 141
of the shell when these are not worn away, and also by the
convexity of the left valve and the small angle formed by the
hinge line and anterior margin (exclusive of the beaks).
Myalina? exasperata. Plate XIX, fig. 4.
Myalina? exasperata Beede, Kans. Univ. Quart., viri, p. 128, pl. xxx11,
f. 4, (1899).
Shell cuneate-ovate in outline; beaks pointed, terminal; shell
very thin, apparently composed of a single layer, compressed ;
valves nearly or quite equal. The anterior? margin nearly
straight above, and merging into the narrowly rounded ventral
region ; the posterior? region similar to the anterior, but more
oblique. The surface is granular and marked by indistinct,
rather broad, concentric strie. Height, 43 mm.; length, 28
mm.; convexity, 3mm.; length of hinge, about 29 mm.
Range and distribution: Upper Coal Measures; Topeka.
The hinge of this shell is not sufficiently well shown to per-
mit of its proper classification. It is left in Myalina for the
present, for want of better information concerning its beak and
muscularimpressions. The extreme thinness of the shell makes
it very probable that it does not belong to that genus.
Myalina perattenuata. Plate XVI, fig. 8.
Myalina perattenuata Meek and Hayden, Trans. Albany Inst., Iv, p. 77,
(1858); Pal. Upp. Mo., p. 32, pl. 1, ff. 12a, b, (1864).
Meek and Hayden’s description: ‘‘Shell very thin and frag-
ile, obliquely elongate, narrow and slightly arcuate; valves
convex anteriorly, and compressed behind. Beaks pointed,
terminal, and attenuate; hinge line equaling rather more than
half the entire length of the shell, and ranging at an angle of
about fifty degrees above the oblique anterior margin. Posterior
border sloping from the end of the hinge, nearly parallel to the
anterior side above, and rounding to the narrow antero-basal
extremity below; anterior margin of the valves a little arcuate,
and rather abruptly deflected inward from the umbonal ridge
above the middle, and in outline nearly straight below. Um-
bonal slopes prominent from the beaks down the anterior side.
Surface with obscure subimbricating marks of growth. Length,
142 University Geological Survey of Kansas.
from the beaks to the postero-basal extremity, 1.50 inches;
breadth, 0.65 inch; convexity, about 0.44 inch.”’
Range and distribution: Upper Coal Measures; Topeka.
This species can be distinguished at once from M. subquadrata
by the small angle at the beaks. It differs markedly from M.
swallovi in its attenuate beak, and the fact that the beak is not
recurved and the umbonal ridge is more nearly straight.
Myalina congeneris. Plate XVI, figs. 2, 2b.
Myalina congeneris Walcott, Pal. Eureka Dist., Mon. U.S. Geol. Surv.
NOMS; p.e201 ply xix fs) Osa pls xin flO:
Walcott’s description: ‘‘ Shell oblong, the height being twice
the width even in the broad examples; hinge line straight and
ranging at an angle of 60° with the nearly straight anterior
margin; base narrow, but rather broadly rounded, posterior
very broadly rounded, curving slightly inwards towards the
cardinal line. General surface of the valves strongly convex
towards the front border, and beaks becoming more depressed
on the posterior portion. Surface of both valves marked by
slightly imbricated lamelle of growth.’’
Range and distribution: Upper Coal Measures; Topeka.
Walcott does not mention the nature of the hinge line in his
description. In one of our specimens the beak is exposed,
showing it to be acute, extending a little beyond the hinge.
The hinge line is similar to that of J/. swallovi in having a sin-
gle groove, but there is no swell in front of the beak. Our
specimens agree more closely with the narrow form as figured
on! Plate XIX.
Our Kansas forms are from nearly the same horizon as M.
perattenuata, to which they are most closely related, but differ
in having their anterior and posterior margins parallel, and, as
a consequence, are not trigonal in outline and the anterior
margin is straight, while that of M. peratienuata is concave.
BREDE. | Carboniferous Invertebrates. 143
AVICULOPINNA.
Meek, Amer. Jour. Sci., X LIV, p. 282, (1867).
Aviculopinna americana. Plate XVIII, fig. 2.
Avicula pinneformis Geinitz, Carb. u. Dyas in Neb., p. 31, pl. 11, f..18,
(not S. pinnwformis Geinitz, 1848), (1866).
Aviculopinna americana Meek, Amer. Jour. Sci. ser. 11, XLIV, p. 282,
(1867); Fin. Rep. U. S. Geol. Surv. Neb., p. 197, pl. rx, ff. 12a—d, (1872);
Pal. Onio; a, p. dal, pl. xx,'f. Z,'( 1815); Herrick, Bull.. Den. Univ., 11,
p. 38, pl. 1, f. 20, (1887); Keyes, Geol. Surv. Mo., v, p. 115, (1894).
Meek’s description: ‘‘ Shell small, compressed, with the slen-
der elongated form of some of the Carboniferous species of Pinna ;
cardinal and ventral margins generally nearly straight (the
_ latter being more convex in outline) and converging gradually
from behind to the rather obtusely pointed anterior extremity ;
posterior side truncated, rounding into the base, and inter-
secting the posterior extremity of the hinge very nearly at
right angles —a little sinuous just below the extremity of the
hinge. Cardinal margin so slightly convex in outline as to ap-
pear quite straight, very nearly equaling the greatest length of
the valves, and provided with a well-developed marginal ridge,
which narrows to a mere line, or dies out before reaching the
beaks, and widens gradually to the posterior extremity. Beaks
nearly or quite obsolete, extremely oblique, and very slightly
behind the very narrow, obtusely pointed, anterior extremity.
Surface with two or three broad, nearly obsolete radiating
ridges on the pcsterior dorsal region, and ornamented by nu-
merous slender, very regularly disposed and abruptly elevated
lines or lamellz, much narrower. than the spaces between, and
curving gracefully parallel to the posterior border, while on
the basal half of the valves they are closely approximate and
curved forward.”’
Range and distribution: UpperCoal Measures; Kansas City,
Lansing (Leavenworth county), Topeka.
Aviculopinna illinoiensis. Plate XVII, figs. 1-lc.
Aviculopinna illinoiensis Worthen, Bull. No. 2 Ill. St. Mus. Nat. Hist.,
p. 13, (1884); Geol. Surv. IIl., viz, p. 128, pl. xx, ff. 5, 5a, (1890).
Shell of medium size, both valves convex; dorsal and ventral
margins moderately curved, converging at an angle of about
20°. Beak very small and indistinct but placed about as in
144 University Geological Survey of Kansas.
A. americana. Surface ornamented with concentric lines paral-
lel to the posterior end of the shell. They are directed from
the hinge line obliquely backward until they pass the central
part of the valve, where they curve rapidly forward, finally be-
coming nearly parallel to the ventral margin before reaching
it. These lines are sharp, thread-like elevations or lamelle,
abruptly elevated from the surface of the shell and separated
by broad, shallow spaces of several times their width. Length
of specimen (the posterior portion is shortened by compres-
sion), 55 mm.; height, at the posterior end, 19 mm.; con-
ERIN] 5) aoa -
Range and distribution : Upper Coal Measures; Kansas City,
head of Deer creek ( western portion of Douglas county ).
This specimen agrees with Worthen’s description and figures,
except that the angle formed by the dorsal and ventral margins
is a little smaller. Itdiffers from A. americana in being a little
more convex, larger, in having the lines farther apart and
larger.
PINNA.
Linneeus, Syst. Nat., 10th ed., (1758).
Pinna peracuta. Plate XVII, figs. 3, 3b.
Pinna peracuta Shumard, Trans. St. L. Acad. Sci.,1, p. 214, (1858); Meek,
Fin. Rep. U.S. Geol. Surv. Neb., p. 198, pl. v1, ff. lla, b, (1872); White,
U. S. Geog. Surv. West 100 Mer., iv, p. 151, pl. x1, f. 5a, (1877); ete.
See Weller, Bull. 153 U. S. Geol. Surv., p. 429, (1898).
Pinna adamsi, McChesney, New Pal. Foss., p. 74, (1860).
Meek’s description: ‘‘Shell thin, very narrow, elongated,
and tapering gradually and regularly from the larger to the
smaller extremity ; convex or almost subcylindrical, excepting
toward the posterior extremity, which is compressed and ob-
liquely rounded, or subtruncated. Hinge margin very long,
and almost perfectly straight—carinated in consequence of the
sudden erection of the dorsal edges of the valves; ventral mar-
gin equally as straight as the dorsal, and ranging at an angle of
about 12° with the latter. Surface nearly smooth, or showing
very obscure lines of growth.”’
Range and distribution: Upper Coal Measures; Kansas City,
Lawrence, Topeka.
BEEDE. | Carboniferous Invertebrates. 145
Pinna subspatulata. Plate XVII, fig. 2; plate XVIII, figs. 1, 1d, and 3.
Pinna subspatulata Worthen, Geol. Sury. Ill., v1, p. 524, pl. xxx, f. 4,
(1875).
Shell very large, triangular in outline, cardinal and ventral
margins diverging at an angle of about twenty degrees ; com-
pressed laterally. Dorsal and ventral margins nearly straight,
the dorsal slightly convex. Surface ornamented by very fine
lines of growth with occasionally larger ones; beginning at the
dorsal margin, which is ridged, the lines pass across the cen-
tral portion of the valve nearly transversely and curve rapidly
forward, finally reaching the ventral edge several centimeters
in front of their origin in the dorsal margin. The hinge fur-
row down the dorsal margin makes a corresponding ridge on
the outside of the shell. Length of specimen, incomplete at
both ends, 25 cm.; from broken posterior edge to the beaks, or
apex of the angle formed by the sides, 27 cm.; height at the
posterior end, 5.5 cm.; convexity (both valves), in the central
portion of the shell, about three-sevenths the height at that
place.
Range and distribution: Upper Coal Measures; Topeka.
This species differs from P. peracuta Shumard in being much
more compressed, and the angle of the dorsal and ventral mar-
gins being about twice as large. If the concentric lines of
growth cross the shell and reach the ventral margin nearly as
directly as the figure indicates, and the angle at the anterior
end of the shell is as small as represented, they will sharply dis-
tinguish our species from P. subspatulata Worthen. He makes
no statement concerning the convexity of the shell, but accord-
ing to the figure it appears nearly flat, even more so than our
specimen.
Some large specimens from near Kansas City, all of which
are either incomplete or crushed, may belong to this species,
or may be exceedingly large specimens of P. peracuta Shumard.
Near the anterior end they are very much more convex than
the Topeka specimens while they are nearly flat at the poste-
riorend. The angle of divergence of the anterior end is about
the same as in P. peracuta.
146 University Geological Survey of Kansas.
MACRODON.
Lycett, Murch. Geol. Chelt., (1845).
Macrodon sangamonensis? Plate XX, figs. 2-2b.
Macrodon sangamonensis Worthen, Geol. Surv. Ill., viii, p. 123, pl. xx1,
ff. 3, 3a, (1890).
Worthen’s description: ‘‘ Shell large, transversely elongated,
hinge line equal to about four-fifths of the entire length of the
shell. Posterior margin compressed and obliquely truncated,
so as to meet the hinge line at a rather acute angle; posterior
extremity quite narrow and rounding gently downward to the
basal margin. Anterior margin regularly rounded from the
anterior extremity to the basal margin, which is slightly sinu-
ous about the middle. Beak depressed, strongly incurved,
placed about one-fifth the length of the hinge line from the
anterior extremity. A gradually widening depression extends
from the beaks to the posterior extremity on the dorsal margin,
and on this flattened portion six or seven strong strive may be
seen, which extend from the beak to the posterior extremity.
Strong lines of growth extend around the basal margin, and
minute transverse strize are visible under a lens, especially on
the anterior portion of the shell. Length, 24 inches; length of
hinge, 14 inches; height, +3 inch; convexity of the valve, +,
ine apes
Range and distribution: Upper Coal Measures ; Turner (Wy-
andotte county). |
It is with much hesitation that I refer these specimens to
this species, even provisionally. The specimens from Turner
are very closely related on the one hand to M. sangamonensis
Worthen, and on the other to M. striatus (Schloth.). They differ
from the former in not possessing minute radiating striae on
the anterior portion of the shell, but strie larger than on any
other part of the shell except on and above the umbonal ridge,
and in possessing a large shallow sinus with obsolete coste in
the central portion of the shell. — It differs from MM. striatus also
in the obsolete area in the central or sinuate portion of the shell,
and very markedly in the nature of the teeth in the front of
the hinge; in M. striatus (as represented by King) there are
BEEDE. | Carboniferous Invertebrates. 147
three or four teeth on the anterior of the hinge which are but
slightly oblique, having a small angle with the long teeth of
the posterior part of the shell. The first three teeth are much
more oblique, and just back of them are six small, round teeth,
back of which are two or three teeth oblique in the opposite
direction to those on the front of the hinge. It differs from M.
carbonarius (Cox) in the area of obsolete striw, in having the
beak placed farther toward the rear, in having the coarse stri«
in front, and the posterior margin curving farther forward to
meet the hinge.
These specimens are from about the same locality as that
figured by Keyes as IM. sangamonensis? Keyes’s" figure agrees
exactly with neither J. sangamonensis as figured by Worthen,
nor entirely with our specimens, and, inasmuch as there is no
description given that will distinguish this from several other
species of the genus, I am in doubt as to whether his figure
represents a specimen more like J. sangamonensis or like ours ;
but he has the hinge line indicated as longer than the rest of
the shell and the umbonal ridge as nearly destitute of stria,
while our specimens possess large strie at this place.
It is with some difficulty that the young of these specimens
can be distinguished from MM. tenuistriatus Meek, but the striz
are much heavier on the anterior than on the central portions
of the shell, even in the young specimens. The outline and
general appearance aside from this are very similar.
Macrodon obsoletus. Plate XX, fig. 13.
Macrodon obsoletus Meek, List Carb. Foss. W. Va., v, (Ex. Rep. Reg.
Univ. W. Va.), (1871); Geol. Surv. Ohio, 11, Pal., p. 334, pl. xrx, f. 9.
Shell oblong, elliptical, of medium size; hinge line about
equaling the length of the shell; anterior margin making a
right angle, or sometimes less, with the hinge line, and round-
ing gradually into the ventral margin, which is a little sinuate
near the center and nearly parallel with the hinge line; pos-
terior margin abruptly but regularly rounded, curving forward
to meet the hinge line. Sinus across the shell from the umbo to
11. Gen. Surv. Mo., V, p. 121, pl. xuvr, f. 2, (1894).
148 University Geological Survey of Ixansas.
the ventral margin nearly obsolete; sinus along the hinge line
widening considerably at the posterior extremity, making the
posterior umbonal convexity narrow. Valves rather convex.
There are three long teeth on the posterior portion of the hinge,
nearly parallel, but apparently do not reach more than one-third
the distance from the posterior to the anterior end of the shell;
beneath the umbo there are four or more small, comb-like, trans-
verse teeth. In front of these there is a thickening of the shell;
anterior to the thickening of the shell, and near the front, are
one or two large, oblique teeth making an angle of about 45°
with the hinge line. Concentric lines of growth visible, dis-
tinct but not prominent. Length, 20 mm.; height, 7 mm.;
convexity, single valve, 5 mm.
Range and distribution: Upper Coal Measures; Olathe, Tur-
ner, Lawrence.
a
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eo.
ye
one
a:
a
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a
i
ee E ieee tn 2. — +
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I am able to find no trace of radiating strie, mentioned by
Meek, on these specimens. However, the specimens appear to
have been water-worn before being enclosed in the matrix, and
may have been abraded. The specimens seem to be identical
with some from Carbon Hill, Ohio, except that the latter are a.
little larger. The hinge markings are also very similar. The
anterior margin of the shell often meets the hinge line at an ‘
angle of less than 90°; otherwise it seems to correspond very
well to M. obsoletus.
PO ga ae
se Le al i an
NUCULANA.
Likn, Rost. Samml., III, p. 55, (1807).
Nuculana bellistriata. Plate XX, figs. 14, 14b.
Leda bellistriata Stevens, Amer. Jour. Sci., (2), xxv, p. 261, (1858).
Nucula (Leda) kazanensis Swallow, Trans. St. L. Acad. Sci., 1, p. 190, ;
(1858).
Nuculana bellistriata White, 13th Rep. Ind. St. Geol., p. 146, pl. xxxt,. ‘
ff. 8, 9, (1884). *
White’s description: ‘‘ Shell transversely elongate-subovate,
gibbous anteriorly and attenuate behind; basal margin broadly
convex, straightened in the middle; anterior margin narrowly
rounded; posterior margin very narrow; postero-dorsal mar-
gin nearly straight, sloping backward and a little downward
from the beaks; umbonal ridges well defined, situated near to.
BEEDE. | Carboniferous Invertebrates. 149
the postero-dorsal margin, their outline, as seen from above,
forming an elongate ellipse which has a concave surface on each
side of the median ridge, which is formed by the upflexed
margins of the valves there; umbones prominent; beaks in-
curved, situated about two-fifths the full length of the shell
from its front; surface marked by fine, regular, concentric,
raised striw, which are obsolete upon the umbonal ridges and
the space which they enclose.’’
Length of our specimen, 27 mm.; height, 14 mm.; con-
vexity, 10 mm.
Range and distribution: Upper Coal Measures; Kansas City,
Rosedale, Lawrence, Topeka.
Nuculana bellistriata attenuata.
Nuculana bellistriata var. attenuata Meek, Fin. Rep. U.S. Geol. Surv.
ING. p.200, Dl. x, io. dla, b, (1872),
Some are inclined to doubt the validity of this variety. There
are forms of NV. bellistriata which are more pointed, more finely
striated and considerably smaller than others. The striations
on this small shell are much smaller, and are separated by much
narrower spaces, than the corresponding strie of the umbonal
regions of the larger specimens. I believe they are varietally
distinct.
Range and distribution: Upper Coal Measures; Kansas City,
Topeka.
NUCULA.
Lamarck, Anim. Sans Vert., p. 87, (1801).
Nucula beyrichi. Plate XXII, fig. 8.
Nucula beyrichi Geinitz, Carb. u. Dyas in Neb., p. 21, pl. 1, ff. 36, 37,
(1856); Meek, Fin. Rep. U. S. Geol. Surv. Neb., p. 203, pl. x, f. 18,
(1872); ete.
Meek’s description: ‘‘ Shell very small, longitudinally sub-
ovate, moderately convex, widest posteriorly; anterior end
somewhat narrowly rounded; base forming a semiovate curve,
the most prominent part being near the shorter end; posterior
side comparatively wide, and subtruncated; beaks near the
posterior extremity ; hinge line nearly rectangular at the beaks ;
denticles comparatively large, about seven on the longer side,
150 University Geological Survey of Kansas.
and five or six on the shorter; surface marked with moderately
distinct regular concentric strie. Length, 0.16 inch; height,
Oh Osineht
Range and distribution: Upper Coal Measures; Grand Sum--
mit. :
A single worn specimen, probably of this species, is in the
University collection, from Grand Summit. The markings are
abraded, and most if not all of the shell is gone. However,
it agrees very well with the figures of the species given by Meek
in the base of the plate in the Nebraska report.
Nucula ventricosa. Plate XXII, figs. 9, 9b.
Nucula ventricosa Hall, Geol. Iowa, 1, pt. 11, p. 716, pl. xx1x, ff. 5a, b,
cee Meek, Fin. Rep. U.S. Geol. Surv. Neb., p. 204, pl. x, ff. 17a—e,
Meek’s desription: ‘‘Shell small, thick, subovate, very con-
vex; the greatest convexity slightly in advance of the middle
of the valve; posterior (shorter) end obliquely truncated from
the beaks to its narrowly rounded or subangular connection with
the base, rather deeply excavated just behind the beaks; ante-
rior (longer ) end rather narrowly rounded, its most prominent
part being near or slightly above the middle; dorsal outline de-
clining gently, with moderate convexity from the beak to the
anterior extremity ; basal margin forming a nearly semiovate
curve, being a little more prominent before than behind the
middle; beaks convex, rather prominent, and placed about half
way between the middle and the most projecting part of the
postero-ventral extremity. Surface with (at least near the base)
fine, regular, concentric strie. Length, 0.42 inch; height,
OOZineh.?? | |
Description of the cast: The posterior adductors are strongly
developed, ovate in outline, situated at the lower part of the
posterior extremity of the shell, nearly beneath the beaks; pal-
lial line very heavily impressed, forming a somewhat oval curve.
Anterior adductor scars also prominent, considerably larger
and more circular in outline than the posterior, situated on the
upper side of the extremity of the umbonal ridge. Immedi-
ately to the rear of this are a pair of small pyriform impressions.
BgEDE. | Carboniferous Invertebrates. 151
There is a depression on some casts on the anterior side of the
umbones extending somewhat below to about the elevation of
the upper part of the posterior scar, in front of which is a cor-
responding nodose ridge. This character varies greatly in dif-
ferent specimens, but the characters are generally sufficiently
developed to be detected. In the cast, the beaks are rather
widely separated and the top of the cast in front of them is
slightly concave, yet nearly flat.
Range and distribution : Upper Coal Measures; Kansas City,
Rosedale, Turner, Topeka, Grand Summit.
This species may be easily distinguished from the preceding
by its larger size, longer shell, and more robust appearance.
Nucula pulchella. Plate XXIJI, figs. 5a-c.
Nucula pulchella Beede and Rogers, Kans. Univ. Quart., viii, p. 132, pl.
XxXxIV, ff. 5a-c, (1899).
Original description: ‘‘ Shell very small, subtrigonal in out-
line, ventricose ; anterior border straight, rounding abruptly to
the ventral margin, which is rounded ; posterior abruptly trun-
cated nearly at right angles with the ventral margin. Beaks
prominent, incurved, situated nearly at the posterior end of
the shell; lunule not well defined. The greatest convexity is
at the umbo. The surface is ornamented by fine, elevated,
concentric strie and undulations of growth. Length, 43 mm. ;
height, 3) mm.; convexity, 8 mm.”’
Range and distribution: Upper Coal Measures; Lawrence.
This species differs from NV. beyricht in being more triangu-
lar, shorter, and the beaks more prominent. It differs from
N. ventricosa, also, in the above respects, as well as being very
much smaller and having very much more prominent surface
markings.
152 Uriversity Geological Survey of Kansas.
YOLDIA.
Moeller, Kroyer’s Nat. Tid., IV, p. 91, (1842).
Yoldia subscitula. Plate XX, fig. 8.
Leda subscitula Meek and Hayden, Trans. Alb. Inst., rv, p. 79, (1858).
Yoldia subscitula? Meek, Fin. Rep. U.S. Geol. Surv. Neb., p. 205, pl. x,
oy LOS (Sia) mene:
Meek’s description: ‘‘Shell longitudinally subovate or sub-
elliptic, compressed, the greatest convexity a little in advance
of the middle, about twice as long as high; anterior extremity
wider than the other but rather narrowly rounded, the most
prominent point being usually slightly above the middle; out-
line of base forming a broad semiovate curve, being more
prominent anteriorly than behind; posterior side narrowed, its
margin rounding up gradually from the base, so as to meet the
dorsal margin at nearly right angles, sometimes faintly truncate
at the immediate extremity ; posterior dorsal margin compressed
or cuneate, and declining gradually, with a nearly straight, or
slightly concave outline; anterior dorsal margin not cuneate,
sloping forward gradually, and a little convex in outline; beaks
rather depressed or subcentral, or very little in advance of the
middle; umbonal slopes without any defined ridge or angle.
Surface smooth, or only showing traces of very minute con-
centric strie. Length, 0.77 inch; height, 0.37 inch; comes
ity, 0.14 inch.
‘‘Tt is with considerable doubt that I have concluded to refer
this shell to Y. subscetula M. and H., because the specimens from
Nebraska City are distinctly more compressed than the type
upon which that species was founded, their convexity being
uniformly not more than half as great proportionally. The
five or six individuals in the collection are constant in this
character, and yet show no evidences whatever of accidental
compression. Otherwise the two forms are very similar in their
general outline, but we know nothing of the internal and hinge
characters of the form under consideration. I strongly suspect,
however, that it will be found to be a distinct species, in which
case I would propose to call it Yoldia propinqua, from its near
resemblance to Y. subscztula.’’
:
’
;
BEEDE. | Carboniferous Invertebrates, 153
Range and distribution: Upper Coal Measures; Topeka.
Only a single left valve of this species has been collected by
the writer, at Topeka, Kan. It agrees so well with this descrip-
tion that it is referred to it, though the lines of growth pass
forward on approaching the hinge and become nearly parallel
to it.
Yoldia glabra. Plate XXI, figs. 4a, b.
Yoldia glabra Beede and Rogers, Kans. Univ. Quart., viti, p. 133, pl.
XxxIVv, ff. 4a, b, (1899).
Shell of medium size for this genus, truncate-subelliptical in
outline, nearly flat, greatest convexity at the umbo, sloping
- gently in all directions from the central portion of the valve ;
anterior dorsal outline nearly straight, sloping gently, and
broadly rounded into the anterior ventral margin, which forms
an elliptical curve to the posterior end, where it is somewhat
truncate, meeting the hinge at an obtuse angle. Hinge line
back of the beak nearly straight. Beaks depressed, nearly
central. Surface nearly smooth, with traces of obscure, dis-
tant, concentric strive parallel to the ventral outline; lines of
growth visible on the postero-dorsal region. Measurements :
Length, 14; mm.; height, 7 mm.; convexity of single valve,
14+ mm. |
Range and distribution: Upper Coal Measures ; Lawrence.
This shell differs from Y. levistriata M. and W., from the St.
Louis group, in having its posterior border truncated, strix
distant rather than closely arranged, and in being much less
convex. It differs from Y. subscitula? Meek, or Y. propinqua
Meek, in being more depressed and the posterior end much
more broadly rounded.
11—vi
154 University Geological Survey of Kansas.
Yoldia knoxensis? Plate XX, fig. 4.
Leda polita McChesney, New Pal. Foss., p. 53, (1860).
Leda knoxensis McChesney, Ill. New Spec. Foss., pl. i, ff. 9a—c, (1865).
Yoldia knoxensis McChesney, Trans. Chic. Acad. Sci., 1, p. 39, pl. 11, ff.
9a-c, (1868).
Shell rather large for this genus, ovate in outline. The hinge
line in front of the beaks bends somewhat downward; the en-
tire anterior and ventral border from near the beaks around to
the antero-dorsal border forming a regular, perfectly ovate
curve, the broad end of which is placed foremost; postero-
dorsal border rounded more sharply but not angulated, turning
considerably forward to meet the hinge, in an almost straight
line. Hinge, back of the beak, nearly straight ; teeth numerous
and very fine; valves moderately convex ; umbones not promi-
nent; beaks closely approximated and incurved. Greatest
convexity in front of the middle of the shell; the greatest
vertical diameter near or a little in front of the center; beaks
placed about one-third the distance from the front end of the
hinge. ‘There is a faint depression extending from back of
the umbo, near the hinge line, obliquely backward over the
umbonal ridge but soon vanishing. This is so faint as to be
hardly noticeable, yet it is present in every specimen before me.
Surface ornamented by very fine, regular, concentric lines of
erowth. Length, 28 mm.; height, 16 mm.; convexity ( both
valves),8 mm. Specimen a little larger than the average.
Range and distribution : Upper Coal Measures ; Leavenworth,
Lansing coal-shaft.
This shell agrees very well with McChesney’s description of
Y. knoxensis, Proc: Chie, Acad: Nat. Sei. 1, plist 29> loutadane
fers from the figure of the species in being about as wide just
back of the beaks, near the center of the shell, as at any other
place, in being less attenuate behind, less abruptly rounded on
the posterior extremity, and meeting the hinge line at a very
much greater angle than is shown by his figure, which shows
the posterior extremity of the shell to be subtruncate.
BEEDE.| Carboniferous Invertebrates. 155
SCHIZODUS.
King, Ann. Mag. Nat. Hist., XIV, p. 313, (1844).
Schizodus wheeleri. Plate XXII, figs. 1-lc.
Cypricardia? wheeleri Swallow, Trans. St. L. Acad. Sci., 11, p. 96, (1862).
Schizodus obseurus Geinitz, Carb. u. Dyas in Neb., p. 20, pl. 1, ff. 30, 31,
(1866).
Schizodus wheeleri Meek, Fin. Rep. U.S. Geol. Surv. Neb., p. 209, pl. x,
ff. la—f, (1872). .
Meek’s description: ‘‘ Shell attaining a medium size, longi-
tudinally subovate, moderately convex; anterior side wider
than the other, and regularly rounded; posterior side nar-
rowed, and obliquely truncated; basal outline rather promi-
nently rounded anteriorly, and straightened, or slightly sinuous
between the middle, and sharply rounded or subangular pos-
terior basal extremity; dorsal margin straight, and sloping
from the beaks to the truncated posterior edge; beaks rather
depressed (for a species df this genus), incurved, placed about
half way between the middle and the front, or perhaps nearer
the middle; posterior umbonal slope rather prominent, or
usually forming a rather obtuse ridge near the posterior basal
extremity ; surface with merely fine lines and obscure marks of
growth.’’ :
Range and distribution: Upper Coal Measures; Kansas City.
Schizodus hari. Plate XXII, figs. 2-2d.
Schizodus harii Miller, 17th Ann. Rep. St. Geol. Ind., p. 701, pl. xx, ff.
1-3, (1892); Keyes, Geol. Surv. Mo., v, p. 123, pl. xiv, f. 4, (1894).
Shell moderately large, quite convex, rather obliquely sub-
ovate in outline, transverse outline ovate. The outline is regu-
larly rounded from near the beaks around the anterior end to
the postero-ventral margin, where it bends rather abruptly up-
ward and forward to meet the hinge line, which is somewhat
arcuate. Beaks placed well forward, moderately prominent,
somewhat incurved, pointing a little forward, and moderately
approximate. Valves very regularly convex for this genus.
Greatest gibbosity a little in front of the middle; umbonal
ridge very indistinct or almost wanting; the depression ex-
tending back from the beaks along the cardinal line very
156 University Geological Survey of Kansas.
indistinct and ill defined. Posterior portion of the shell some-
what wedge-shaped. The fine concentric strie of growth are
most apparent on the anterior and posterior ends and along the
ventral margin. The interior of the shell is marked by rather
deep adductor scars, the anterior being situated very near the
anterior dorsal margin, rather broadly elliptical in outline,
while the posterior is situated near the posterior end of the liga-
ment and is much larger, rather ovate in outline. The pallial
line is indistinct, but seems to pass rather closely to the ventral
margin around to the anterior end, where it curves rather regu-
larly upward to the anterior adductor. There is a ridge in
the interior of the shell extending from the umbo obliquely
backward about one-half.to two-thirds the distance to the pos-
tero-ventral margin. In the right valve the tooth is quite large
and extends obliquely forward and downward, and is largest at
its extremity ; in front of this is a depression or socket for the
smaller anterior tooth of the left valve, while back of it is a
larger cavity for the large central tooth of the left valve; back
of this cavity there is a slight sharp elevation extending ob-
liquely backward. In the left valve there is asmall, thin tooth
extending obliquely forward, back of which is a large socket for
the reception of the large tooth of the right valve. Posterior to
this there is a thin, deep depression, extending nearly parallel
to the hinge, for the reception of the thin posterior tooth of the
right valve. Length, 52 mm.; height, 40mm.; convexity of
single valve, 13 mm.
Range and distribution: Upper Coal Measures; Kansas City,
Lansing ( probably from the penitentiary coal-shaft ).
One specimen, somewhat below the average size, was worked
from the shale, showing the internal and external features of
both valves, including the hinge.
BEEDE. | Carboniferous Invertebrates. } 157
Schizodus subcircularis.
Schizodus subcircularis Herrick, Bull. Den. Univ., 11, p. 41, pl. ri, f. 24,
(1887).
Schizodus subcircularis Girty, U. 8. Geol. Surv., xrx, pt. 111, p, 582, pl.
LXxII, f. 8a, (1899).
Herrick’s description (in part): ‘‘. . . The anterior and
lower margins are continuous parts of almost a circle, which
is only a little produced posteriorly ; the posterior margin is in-
clined; beak submedian, posterior umbonal ridge not sharp.
The shell is moderately convex and is nearly smooth, except for
the distant lines of growth. Length, 0.63 inch; width, 0.59
inch; convexity (single valve), about 0.20 inch.”’
Range and distribution: Upper Coal Measures; Lawrence.
A single right valve from the dam rock at Lawrence agrees
well with Herrick’s figure and description. It isa little smaller
and, apparently, better preserved than Herrick’s specimen, for
it shows excedingly fine, inconspicuous concentric striz in addi-
tion to the distant lines of growth mentioned in his description.
Schizodus compressus,n. sp. Plate XXII, figs. 6, 6d.
?Schizodus wheeleri Herrick, Bull. Den. Univ., 11, p. 42, pl. 111, f. 15, (1887).
Shell medium size for this genus, not very convex, elongate
suboyate in outline, cuneate posteriorly. Antero-dorsal mar-
gin non-sinuate ; anterior margin broadly rounded ; ventral mar-
gin quite convex, terminating posteriorly in arather prominent
acute angle; posterior margin obliquely truncate; dorsal mar-
gin nearly straight, curving very slightly upward. Hinge line
straight, a little less than one-half the length of the shell.
Beaks depressed, incurved, inconspicuous, scarcely elevated
above the hinge line, located about one-third the length of the
shell from the anterior end. Umbonal slope subangular,
rather prominent, becoming obsolete towards the posterior
angle of the shell. Greatest convexity of the shell below and
slightly behind the umbo. Posterior to the umbonal slope
the shell is much compressed. Adductor scars small, sub-
elliptical in shape, anterior one situated near and a little above
the middle of the anterior margin; posterior one placed close
to the dorsal margin, a little behind the beak. Pallial line sub-
158 University Geological Survey of Kansas.
parallel to the ventral margin. Surface smooth, showing oc-
casional very faint undulations of growth. Length, 31 mm. ;
height at beak, 21 mm.; convexity of single valve, 5 mm.
Range and distribution: Upper Coal Measures; Lawrence.
There is but little doubt that our shells are specifically iden-
tical with Herrick’s specimen, as far as can be learned from his
figure and brief description. At any rate, our species is not S.
wheeler’! It is much more nearly related to S. chesterensis M.
and W., from the Chester group of Lllinois. The beak is less
elevated and placed farther back, the shell is less convex, and
the posterior adductor scar is placed farther forward than in
that species. It bears some resemblance to S. meekanus Girty,”
but the beak is a little less prominent, the shell less elongate,
and the ventral margin curves up more rapidly posteriorly, the
posterior angle being nearer the hinge line than in that species.
A large number of these shells have been coliected from the
dam rock at Lawrence, and from its equivalent at Cameron’s
Bluff. They all agree well with the above description and rep-
resent a distinct species.
SOLENOMYA.
Lamarck, Hist. Nat. Anim. sans Vert., V, p. 488, (1818).
Solenomya parallella. Plate XXI, fig. 1.
Solenomya parallella Beede and Rogers, Kans. Univ. Quart., viii, p. 131,
pln xoocivet. Inn CES aaie
Shell large, moderately convex, elongate-subquadrate in out-
line; anterior margin broadly rounded to nearly truncate,
meeting the hinge at a little more than aright angle; dorsal:
and ventral margins straight, nearly parallel; postero-dorsal
margin sloping obliquely downward, meeting the rounded pos-
terior margin at an obtuse angle. Beaks depressed, incurved,
situated about one-fourth the distance from the posterior to the
anterior end of the shell. The greatest convexity is at the
umbo, the shell gaping a little wider anteriorly than behind.
Surface marked by flat, radiating plications, which become
obsolete at either end of the shell, the whole surface of which
12. U. S. Geol. Surv., 19th Ann. Rep., pt. IIT, p. 588, pl. LX XIT, ff. 7a-c, (1899),
BEEDE.| | Carboniferous Invertebrates. 159
is apparently covered with radiating strive parallel to the plica-
tions. Neither the plications nor the striw radiate directly
from the beak, but from a point above the beak and a trifle in
front of it. These are crossed by fine obscure lines of growth.
The anterior and posterior adductors are prominent; the pos-
terior one is triangular, located between the beak and the pos-
terior margin, along its anterior side, and extending above it
is a ridge caused by the thickening of the shell, which produces
a slight oblique furrow in the cast. The anterior scar is lo-
cated on the upper side of the shell, near the anterior angle.
The pallial line is indistinct, parallel to the ventral margin,
curying backward to meet the anterior adductor scar. Length,
53 mm.; height, 18 mm.; convexity of single valve, about
5mm.
Range and distribution: Upper Coal Measures; Kansas City,
Lawrence.
Solenomya trapezoides. Plate XXI, figs. 2a, b.
Solenomya sp. Meek and Worthen, Geol. Surv. Ill., v, p. xxvut, ff. la, b,
(1873).
Solenomya trapezoides Meek, Amer. Jour. Jci., (3), vii, pp. 582, 583,
(1874): Beede and Rogers, Kans. Univ. Quart., vii1, p. 132, pl. xxxrv, ff.
2a, b, (1899).
Shell large, subelliptical in outline, length from two to two
and a half times the height, convex, open at both ends. Ven-
tral,margins slightly convex, curving abruptly upward on the
posterior extremity and more gently on the anterior; posterior
side obliquely truncated above; hinge straight in front of the
beaks, which are depressed, approximate, located about one-
fourth the length of the shell from the posterior extremity.
Surface marked by rather obscure concentric undulations of
growth, crossed by faint radiating strive, which seem to radiate
from the beak. Posterior adductor scar moderately prominent,
more or less irregularly subcircular; the ridge in front of the
scar is broad and its outline is rather indistinct, nearly perpen-
dicular to the hinge, curving backward below; anterior scar
indistinct, somewhat subcircular; pallial line extending back-
ward and downward from the lower side of the posterior scar,
160 University Geological Survey of Kansas.
then curving abruptly forward parallel to the ventral margin to
the middle of the shell, where it becomes too indistinct to trace
in our specimens. Measurements of a specimen a little below
the average size and a larger specimen: Length, 56 mm., 76
mm.; height, 24 mm., 33 mm.; convexity, 15 mm., 25 mm.,
respectively.
Range and distribution: Near the junction of the Upper and
Lower Coal Measures, at Porterville, Kan., and Westport, Mo.
Solenomya radiata. Plate XXII, figs. 5-5b.
Solenomya radiata Meek and Worthen, Proc. Acad. Nat. Sci. Phil., p.
457, (1860).
Solenomya radiata Meek and Worthen, Geol. Surv. IIl., 11x, p. 349, pl.
XxVI, ff. 10a, b, (1866).
Meek and Worthen’s description: ‘‘Shell thin, narrow ob-
long-oval or subelliptical, moderately convex, nearly closed at
the each end; pallial margin rather straight or very slightly
contracted along the middle, and rounding up more gradually
in front than behind; anterior (longer) side narrowly rounded,
its most prominent part being above the middle; posterior
(shorter) side narrowly rounded below and obliquely subtrun-
cate above; dorsal outline nearly parallel to the base; beaks
much depressed, located less than one-fourth the entire length
of the shell in advance of the posterior extremity. Surface
with obscure marks of growth, crossed by flat, nearly obsolete
radiating plications, which are sometimes separated, near the
middle of the valves, by spaces greater than their own breadth ;
plications very oblique and more closely arranged on the an-
terior side. Length, 1.17 inches; height, 0.47 inch; convex-
ity, about 0.33 inch.”’
Range and distribution: Upper Coal Measures; Topeka.
33,74 Ss sie 2 vee eee
BEEDE. | | Carboniferous Invertebrates. 161]
PLEUROPHORWUS.
King, Ann. Mag. Nat. Hist., XIV, p. 313, (1844).
Pleurophorus subcostatus. Plate XX, figs. 11-11b.
Pleurophorus subcostatus Meek and Worthen, Proc. Acad. Nat. Sci.
Phil., p. 246, (1865).
Pleurophorus subcostatus Meek and Worthen, Geol. Sury. I[ll., 11, pp.
347, 348, pl. xxvii, ff. 2, 2a, (1866).
Meek and Worthen’s description: ‘‘ Shell elongate-oblong,
moderately convex; umbonal ridges the most convex part of
the valves, and extending obliquely from the beaks toward the
postero-basal margin; anterior ventral region somewhat com-
pressed ; basal and cardinal margins very nearly straight and
subparallel, the former being usually somewhat sinuous or ar-
cuate along the middle; extremities rather narrowly rounded,
the posterior being generally a little wider than the other, and
sometimes faintly subtruncate obliquely. Hinge line long and
nearly straight; posterior lateral tooth of each valve elongated
parallel to the hinge margin, very remote from the cardinal
teeth, and extending back a little beyond the posterior muscular
impression. Beaks depressed upon a line with the dorsal mar-
gin, small, somewhat compressed, and placed about one-ninth
the entire length of the shell behind the anterior margin. Scar
of the anterior adductor muscle deep, trigonal-subovate, pointed
-above, and strongly defined by the prominent vertical ridge
just behind it; those of the pedal muscles small, nearly mar-
ginal, and located directly over the anterior adductors ; posterior
adductor scars larger and more shallow than the anterior, sub-
quadrate in outline, and placed close up:under the posterior
hinge teeth. Pallial impression well defined. Surface of casts
showing traces of a few obscure concentric markings, crossed’
on the postero-dorsal region by traces of about three equal ob-
scure radiating coste. Exterior surface and cardinal teeth un-
known. Length of a medium-sized specimen (internal cast) ,
0.88 inch; height of same, 0.37 inch; convexity, 0.26 inch.
Some larger specimens, of same proportions, measure 1.33
inches in length.’’
Range and distribution: Upper Coal Measures; Kansas
City, Mo.
162 University Geological Survey of Kansas.
Pleurophorus tropidophorus. Plate XX, fig. 7.
Pleurophorus tropidophorus Meek, Geol. Surv. Ohio, 1, Pal., p. 338,
pl. x1x, ff. 10a, b, (1875). -
Original description: ‘‘ Shell transversely oblong, much com-
pressed, with length a little greater than twice the height;
posterior margin flattened and bifurcated, the lower truncation
being nearly vertical, and the upper sloping obliquely down-
ward and backward from the hinder end of the hinge; cardinal
margin straight, equaling about two-thirds the length of the
valves ; anterior rounded below and sloping abruptly forward
from the beaks above; basal margin long, parallel to the hinge,
nearly straight for most of its length, or faintly sinuous near
the middle, rounding up anteriorly, and forming a more or less
defined angle at its connection with the lower part of the pos-
terior margin behind; posterior umbonal slope distinctly an-
gular from the beaks to the angular posterior basal extremity,
while a second carina passes obliquely backward and downward
along the middle of the dorsal space above the umbonal ridge
of each valve; beaks depressed to the line of the cardinal
margin, very little projecting, and placed one-fifth to one-fourth
the length of the valves from the anterior margin. Surface
marked by distinct concentric lines of growth, that become
strongly defined on the flanks and anterior parts of the valves,
but are less distinct on the space above and behind the umbonal
angles. Length, 1.10 inches; height, 0.52 inch; convexity,
about 0.20 inch.’’
Range and distribution: Upper Coal Measures; Kansas
City, Mo.
Pleurophorus costatus.
Arca costata Brown, Trans. Man. Geol. Soc., 1, p. 32, pl. vi, ff. 34, 35,
(1841).
Pleurophorus costatus Brown, King, Cat., p. 11, (1848): King, Mon.
Perm. Foss. Eng., p. 181, pl. xv, ff. 13, 14, (1850). acai
Pleurophorus costatiformis Meek and Worthen, Geol. Surv. IIl., 1, p.
535, pl. x1x, f. 8, and text fig., (1868).
Meek and Worthen’s description: ‘‘Shell elongate, suboval,
moderately convex, slightly arcuate; the dorsal and ventral
margins rather long, and more or less nearly parallel, the
BREDE. | Carboniferous Invertebrates. 163
former being a little concave in outline, and the latter convex ;
extremities narrowly rounded. Beaks small, depressed, or ris-
ing little above the hinge line, very oblique, somewhat com-
pressed, incurved, and placed very near the anterior end;
lunule apparently small and deep. Surface ornamented by
concentric strive of growth, and a few larger, obscure, concen-
tric wrinkles, crossed on the postero-dorsal region by five dis-
tinct, equidistant and radiating ridges, extending obliquely
from the beak to the posterior margin, the lower one being the
largest, and forming the umbonal ridge, while the upper one
runs parallel to the cardinal margin, and forms the edge of the
long corslet, or escutcheon. Length, about 1.10 inches; height,
0.46 inch ; convexity, near 0.43 inch.’’
Range and distribution: Upper Coal Measures; Kansas
City, Mo.
I see no reason for putting this shell in a distinct species.
Our Kansas City specimens agree well with King’s descrip-
tions and figures, being longer than Meek and Worthen’s fig-
ures. It seems that the principal reason for making a distinct
species of this shell was the fact that it came from a much
lower horizon than the European forms, but this is true of sev-
eral species of fossils which are common to the Coal Measures
of the Missouri valley and the Permian of Europe. The lines
of growth (or striw) on our specimens meet the hinge at a very
obtuse angle, while Meek’s figure represents them meeting
nearly at right angles. In this respect it is like costatus:
ASTARTELLA.
Hall, Geol. Iowa, pt. II, p. 715.
Astartella vera. Plate XXII, fig. 10.
Astartella vera Hall, Geol. lowa, pt. 11, p. 715, pl. xxrx, ff. la-c, (1858);
Keyes, Geol. Surv. Mo., v, p. 125, pl. xtv1, f. 6, (1895); ete.
Original description: ‘‘ Shell somewhat rhomboid-ovate, gib-
bous on the umbones; beaks subanterior, elevated, approxi-
mate ; anterior end slightly concave below the beak and rounded
below, posterior end obliquely truncate. An oblique undefined
ridge extends from the beak to the posterior basal margin, hav-
ing the space between it and the ligamental area flattened ;
1€4 University Geological Survey of Kansas.
lunule cordiform, strongly impressed; ligamental area deeply
marked, and extending to the posterior extremity. Anterior
and posterior muscular impressions distinct; hinge strong.
Teeth of right valve separated by a deep pit; the anterior tooth
with a longitudinal pit in the summit, and a callosity on the
inner margin at its base. Surface marked by strong concentric
furrows, which are separated by sharp angular ridges; the in-
termediate space finely striated.’’
Range and distribution: Upper Coal Measures; Grand Sum-
mit.
CONOCARDIUM.
Bronn, Leth. Geo., I, p. 92, (1885).
Conocardium parrishi. Plate XX, fig. 9.
Conocardium parrishi Worthen, Geol. Surv. Ill., vit, p. 112, pl. xx, f.
7, (1890); Keyes, Geol. Surv. Mo., v, p. 124, pl. xxv1, ff. Ga, b, (1894).
Worthen’s description: ‘‘Shell obliquely triangular, hinge
line straight, beaks depressed, umbonal ridge elevated into a
strong rounded fold, with faint traces of fine strie towards the
lower extremity. Anterior side evenly and rapidly sloping
from the umbonal fold to the extremity, traversed by radiating
costze, only four or five of which extend to the hinge line, but
increasing by division and implantation so that ten or more
may be counted on the margin of the shell, the one nearest the
umbonal ridge being somewhat stronger than the others. Pos-
terior side flattened towards the extremity, and marked by
eight or more flattened strive, nearly all of which extend to the
hinge line. Length, about ;; inch.”’
Range and distribution: Upper Coal Measures; Kansas
City.
CYPRICARDINIA.
Hall, Pal. N. Y., III, p. 266, (1860).
Cypricardinia? carbonaria. Plate XX, fig. 16.
Cypricardinia? carbonaria Meek, Proc. Acad. Nat. Sci. Phil., p.163, (1871).
Cypricardinia ? carbonaria Meek, Geol. Surv. Ohio, 11, Pal., p. 342,
pl xix, ff.'8a, bo, (i8ia):
Meek’s description: ‘‘Shell small, longitudinally oval, less
than twice as long as high, the widest (highest) part being
under the posterior extremity of the hinge; rather gibbous,
with usually a broad impression extending from the beaks ob-
——T-
Sa eo ere rt
ba awe ee ie ee a —s
BEEDE. | Carboniferous Invertebrates. 165
liquely backward and downward to the middle of the base of
each valve; anterior side extremely short, or nearly obsolete,
convex, and rounded ; posterior side broader, more compressed
or cuneate, with its upper edge straight and sloping obliquely
backward to the regularly rounded posterior margin; base
broadly and slightly sinuous in the middle, and rounding up-
ward at the extremities; hinge line straight, between one-half
and two-thirds as long as the valves, ranging at an angle of
about twenty-five degrees with the oblique, longer axis of the
shell, so as to meet the sloping upper edge of the posterior
margin at a very obtuse but moderately well-defined angle, thus
imparting to the somewhat compressed posterior dorsal region
a very faintly alate appearance; beaks extremely oblique, de-
pressed nearly to the dorsal margin, very nearly terminal, and
scarcely projecting beyond the rounded outline of the anterior
extremity. Surface ornamented by about fifteen to twenty ex-
ceedingly regular, well-defined, subimbricating, flattened, con-
centric ridges or undulations, that gradually become smaller
and more closely approximating on the umbones.’’
Range and distribution: Upper Coal Measures; in the oolite
at Rosedale and Turner.
CARDIOMORPHA.
de Koninck, Anim. Foss, Carb. Belg., p. 101, (1844).
Cardiomorpha missouriensis. Plate XX, fig. 17.
Cardiomorpha missouriensis Shumard, Trans. St. L. Acad. Sci., 1, p.
207, (1858); Meek and Worthen, Geol. Surv. Ill., v, p. 588, pl. xxvu, f. 8,
(1873).
Nucula mercerensis McChesney, Trans. Chic. Acad. Sci., 1, p. 40, pl. m1,
ff. 12a—c, (1868).
Nucula cylindrica McChesney, New Pal. Foss., p. 54, (1860).
Original description: ‘‘ Shell inequilateral, elliptico-subquad-
rate, very thin, length double the height; superior and infe-
rior borders subparallel ; cardinal margin long, slightly arcuate,
inferior border slightly arched ; anal and buccal margins strongly
rounded, the latter being very short ; umbonal region moderately
conyex in young specimens, and very gibbous in the old, great-
est convexity short distance below the beaks; beaks situated
' near the anterior margin, rounded, closely incurved and nearly
166 University Geological Survey of Kansas.
approximate ; surface marked with very fine crowded, concen-
tric strie, which are sometimes more or less flexuous. Length
of full-grown specimen, 1.30 | inches?] ; height, 0.64; thick-
ness, 0.62.”’
Range and distribution : Upper Coal Measures; Topeka.
EDMONDIA.
de Koninck, Anim. Foss. Carb. Belg., p. 66, (1844).
Edmondia nebrascensis. Plate XX, fig. 5.
Astarte nebrascensis Geinitz, Carb. u. Dyas in Neb., p. 16, pl. 1, f. 25,
(1866).
Edmondia nebrascensis Meek, Fin. Rep. U. S. Geol. Surv. Neb., p. 214,
pl. x, f. 8-8b, (1872).
Meek’s description: ‘‘ Shell subovate, Lo eee more or
less rounded at the extremities ; length nearly once anda half the
height; basal margin broadly semielliptic or semiovate in out-
line; dorsal margin sloping from the beaks, but more abruptly
in front than behind, rounding into the extremities; beaks
moderately prominent, and located somewhat in advance of the
middle. Surface marked by broad, rounded, rather regular
concentric furrows, separated by sharp, moderately prominent
concentric linear ridges, sometimes show under a magnifier in-
dications of being minutely crenate; impressions or furrows
between the ridges, showing concentric strice, which, by the aid
of a lens, in a cross-light appear to be crossed by fine, nearly
obsolete radiating markings. Length of the largest specimen
seen, oe inches; hereht, 0:95 amels oe about 0.30
inch,’
Range and distr iieiade Upper Coal Measures; Kansas City,
Turner, Topeka.
Edmondia aspinwallensis. Plate XXII, figs. 3-3b.
EHdmondia aspinwallensis Meek, Fin. Rep. U.S. Geol. Surv. Neb., p.
216, pl. iv, ff. 2-2c, (1872); ete.
Original description : ‘‘ Shell longitudinally subovate, moder-
ately convex, the greatest convexity being a little in advance of
and above the middle; base nearly semielliptic in outline; pos-
terior side rather narrowly rounded, or sometimes very faintly
subtruncate obliquely above; dorsal margin nearly straight just
BEEDE. | Carboniferous Invertebrates. 167
behind the beaks, but very gradually declining, with a slightly
convex outline posteriorly ; anterior side quite short, declining
very abruptly from the beaks above, and rounded below ; beaks
rather depressed, incurved, and located nearer the anterior end
than the middle. Surface of cast with moderately distinct, ir-
regular concentric undulations, showing behind the beaks dis-
tinct impressions of the cartilage fulera. Length, 1.45 inches ;
height, 1.05 inches; convexity, about 0.68 inch.”’
Range and distribution: Upper Coal Measures; Cherryvale,
Kansas City, Turner, Topeka.
This species may be distinguished from the foregoing by its
more undulate and less carinate concentric ridges, which are
never crenate, and the absence of the nearly obsolete radiating
Strice.
ALLORISMA.
King, Ann. Mag. Nat. Hist., XIV, p. 315 (1844).
Allorisma geinitzi. Plate XX, fig. 6.
Allorisma elegans Geinitz, Carb. u. Dyas in Neb., p. 13, pl. 1, f. 31,
(1866), (non King).
Allorisma (Sedqwickia) geinitzi Meek, Fin. Rep. U.S. Geol. Surv. Neb.,
p. 219, pl. x, ff. 16a, b, (1872).
Allorisma geinitzi Meek, Geol. Surv. Ill., v, p. 586, pl. xxv1, f. 23, (1873).
Meek’s description: ‘‘Shell small, rather compressed, longi-
tudinally subovate, abruptly narrowed from the beaks posteri-
orly ; umbonal slopes distinctly carinate from the beaks to the
posterior basal angle; anterior side subtruncate, with an ab-
rupt slope from the beaks obliquely forward above, and round-
ing into the base below; basal margin somewhat prominently
rounded anteriorly, and nearly straight or faintly sinuous be-
hind ; posterior end compressed, its margin abruptly truncated
vertically, so as to make its upper and lower parts nearly rec-
tangular; cardinal margin sloping, with a slightly concave out-
line, from the beaks to the truncated posterior end; beaks
elevated, incurved, and placed about half way between the
middle and the anterior extremity of the valve. Surface or-
namented with numerous minute, closely crowded granules,
which, on the umbones and other parts of the valves in front
of the angular umbonal slope, show a tendency to arrange
168 University Geological Survey of Kansas.
themselves in radiating lines, which are crossed by more or
less distinct lines of growth; on the compressed corselet, above
and behind the umbonal carina, there are usually two or more
obscure radiating ridges and furrows, crossed by moderately
distinct, granular lines of growth, parallel to the truncated
posterior margin. Length of the largest example seen, 0.50
inch; height of the umbones in the same, 0.30 inch; height of
the truncated posterior end, same, 0.16 inch; convexity, about
OMS inch:
Range and distribution: Upper Coal Measures; Topeka.
Allorisma granosum. Plate XX, fig. 10.
Leptodomus granosus Shumard, Trans. St. L. Acad. Sci., 1, p. 207.
Allorisma (Sedgwickia) granosa Meek, Fin. Rep. U. S. Geol. Surv.
Neb., p. 220, pl. 11, f. 8, (1872).
Allorisma granosum Keyes, Geol. Surv. Mo., v, p. 128, (1895).
Meek’s description: ‘‘Shell very thin, approaching an irregu-
lar, oblong form, the length being less than twice the height,
very convex, most gibbous part being near the middle of the
valves; beaks prominent, incurved, somewhat flattened on the
outer side, and placed about half way between the middle and
the front. Dorsal margin striz straight behind the beaks and
nearly parallel to the general outline of the base, inflected so
as to form a distinct, flattened, lanceolate, lunule-like area,
bounded on each side by a well-defined subangular ridge; pos-
terior side nearly or quite closed, obliquely truncated, with
sometimes a faint sinuosity near the middle; anterior side
rather abruptly sloping forward, and straightened above, and
rounding into the base below, near which it seems to be a little
gaping; base somewhat straightened, or even a little sinuous
in outline, just in front of the middle, at the termination of a
broad, very shallow cavity, extending a little obliquely down-
ward and backward from the umbonal region ; behind this rather
prominent, thence ascending obliquely, with a slightly convex
outline, to the truncated posterior margin. Posterior umbonal
slopes very prominently rounded above, and continued as a low
undefined ridge, obliquely backward and downward ; posterior
dorsal slope, above the umbonal ridge, with an oblique, shal-
Ts oe
eg ee +
BREDE. | Carboniferous Invertebrates. 169
low, rounded sulcus, extending from the back part of the beaks
to the middle of the truncated margin behind. Surface marked
with fine lines of growth and small irregular, concentric wrin-
kles, which latter are not defined on the posterior dorsal region
above the umbonal ridge; crossing these are the usual radiat-
ing rows of minute granules. Length, 2 inches; height, 1.15
inches; convexity, 1 inch.”’
Range and distribution: Upper Coal Measures; Kansas
City, Iola, Lawrence, Lecompton, Topeka.
This species may be very easily distinguished from the fore-
going by its very much larger size and less angular and more
robust form, and the sinuosity of the ventral margin.
Allorisma subcuneatum. Plate XX, figs. 1-1b.
Allorisma regularis? Owen, Geol. Rep. Wis., Iowa, and Minn., pl. v, f.
13, (1852).
Allorisma subcuneata Meek, Fin. Rep. U. S. Geol. Surv. Neb., p. 221,
pl. 11, ff. 10a, b, (1872); etc.
Meek’s description: ‘‘ Shell attaining a large size, longitudi-
nally elongated, or twice to three times as long as high, pro-
portional length increasing with age; greatest convexity a little
in advance of the middle and in the umbonal region; cuneate
and a little gaping behind, where the margin is more or less
narrowly rounded in outline. Basal and dorsal margins nearly
parallel, the latter being more or less concave in outline, or
nearly straight, and inflected so as to form a lanceolate kind of
false area, bounded by an obtuse ridge on each side, just out-
side of which there is a shallow undefined sulcus ; basal margin
slightly convex, or somewhat straightened along the middle,
and sometimes very faintly sinuous just under the beaks, round-
ing up more abruptly before than behind; anterior margin
very short, a little gaping and rather prominently rounded be-
low ; beaks convex, incurved, and placed near the anterior end,
rather depressed, but rising moderately above the dorsal margin.
Surface ornamented with fine strive of growth, and well-defined
concentric undulations usually more distinct and regular on the
beaks and umbonal region. Length of the largest specimen
12—vi
170 University Geological Survey of Kansas.
seen, 4.81 inches; height from the ventral to the Core mar-
gins, near middle, 1.76 inches; convexity, 1.57 inches.’
Range and distribution : peer Coal Measures ; Wesinore
(Mo.), Kansas City, Mont Ida (Anderson county), Lawrence,
Lecompton, Topeka, Elmont, Grand Summit.
This species is distinguished, on account of its larger and
more graceful form, from either of the preceding.
Allorisma costatum. Plate XX, fig. 12.
Allorisma costata Meek and Worthen, Proc. Acad. Nat. Sci. Phil. 1869,
p-. 171; Geol. Surv. IIl., v, p. 585, pl. xxvi, f. 15, (1873).
Meek and Worthen’s description : ‘‘Shell under medium size,
longitudinally oblong, the length being more than twice the
height, very thin, rather convex in the central umbonal regions ;
anterior margin rather short, closed and narrowly rounded ;
basal margin forming a long, rather semielliptic curve, with a
very slight sinuosity in front of the middle; posterior side com-
pressed, but apparently a little gaping and distinctly truncated
nearly vertically from the base about half way up, and thence
a little obliquely forward and upward to the dorsal margin ;
posterior dorsal region compressed above the umbonal ridge ;
cardinal margin equaling about two-thirds the entire length
of the shell, very nearly straight, and inflected so as to form a
narrow or lance-linear corselet, extending over its whole length ;
beaks convex, rising a little above the cardinal margin, and
placed slightly more than one-sixth the length of the valve be-
hind the anterior extremity; lunule well defined and lance-
ovatein form. Surface ornamented by about twenty-five very
regularly arranged, distinctly elevated, concentric costz, which
commence near the lunule and extend backward parallel to the
base, to the well-defined, angular umbonal ridge leading from
the beaks to the posterior basal extremity, at which ridge they
become suddenly obsolete, or very nearly so, being mainly rep-
resented on the more compressed posterior dorsal region by
distinct lines of growth, which are crossed on the middle of
this area by a second oblique linear ridge extending from the
beaks to the middie of the posterior margin. Some indications
ef.
— 9 _ Se
BEEDE. | Carboniferous Invertebrates. 171
of the usual minute surface granules appear to be vislble in
some of the molds left in the matrix. Length, about 1.20
inch ; height, 0.53 inch; convexity, 0.44 inch.”
Range and distribution : Upper Coal Measures; Kansas City,
Lawrence, Topeka.
This species may be easily distinguished by its large, even,
sharply elevated ribs and the radiating cost on the upper pos-
terior portion, as well as its long, graceful shape.
SEDGWICKIA.
McCoy, Synop. Carb. Foss, Ireland, p. 61 (i844).
Sedgwickia topekaensis. Plate XX, fig. 3.
Leptodomus topekaensis Shumard, Trans. St. L. Acad. Sci., 1, p. 208.
Sedgwickia topekaensis? Meek and Hayden, Pal. Upp. Mo., p. 40,
A, B, (1864).
Allorisma topekaensis Keyes, Geol. Surv. Mo., v, p. 128.
Meek and Hayden’s description: ‘‘ Shell depressed subovate,
about twice as long as high, extremely thin and fragile, gib-
bous in the region of the beaks and along the oblique umbonal
slopes. Sides flattened above, and becoming a little concave
towards the base in front of the middle. Dorsal border nearly
horizontal, and slightly concave in outline behind the beaks,
where its inflected edge is margined by a rather distinct ridge ;
ventral border presenting a broad semiovate outline, excepting
a very slight sinuosity just in advance of the middle — round-
ing up abruptly in front and more gradually behind; anterior
side prominent, gibbous, and narrowly rounded below, obliquely
truncated above; posterior side compressed, narrowed, and
apparently subtruncate and somewhat gaping at the extremity.
Lunular impression in front of the beaks (in casts) moderately
distinct, defined by a faintly impressed line. Beaks prominent,
gibbous, a little flattened, incurved, and placed between the
middle and the anterior extremity, but nearer the former. En-
tire surface, in well-preserved specimens, closely covered with
minute granules arranged in radiating rows; and ornamented
with small concentric ridges, which are almost regular and dis-
tinct on the umbones, and end abruptly along an impressed
line extending from the posterior side of each beak obliquely
172 University Geological Survey of Kansas.
towards the postero-basal margin, thus leaving the compressed
postero-dorsal region comparatively smooth. (Muscular and
pallial impressions unknown.) Length, about 2 inches; height,
1 inch; convexity, 0.75 inch.’’ |
Range and distribution: Upper Coal Measures; Topeka.
Our specimens differ somewhat from those figured in having
the sides less depressed near the beaks and ventral margin ; they
having shells a trifle thicker than the description calls for.
CHANOMYA.
Meek, Pal. Upp. Mo., p. 42, (1864).
Chenomya leavenworthensis. Plate XIX, figs. 3-3b.
Allorisma? leavenworthensis Meek and Hayden, Proc. Acad. Nat. Sci.
Phil. 1858, p. 263. |
Chenomya leavenworthensis Meek and Hayden, Pal. Upp. Mo., p. 43,
pl. 11, ff. la—c, (1864); ete.
Meek and MHayden’s description: ‘‘Shell subcylindrical ;
anterior side rounded, a little compressed and apparently
entirely closed ; posterior side long, truncated, and very widely
gaping, the margins being even a little reflexed. Base nearly
straight, or slightly convex in outline, rounding up gradually
in front, and very abruptly behind; dorsal side concave in out-
line from the beaks to its elevated posterior extremity, and
nearly parallel to the base. Beaks rather depressed, somewhat
flattened, incurved, nearly or quite touching, and located about
half way between the middle and the anterior end. Surface
marked by fine lines of growth, and a few irregular, nearly
obsolete concentric undulations, which curve up abruptly be-
hind, parallel to the truncated posterior margin. Crossing these,
the radiating rows of minute granules may be seen by the aid
of a good lens, on well-preserved specimens. Internal casts of
this species show quite distinctly the scar of the anterior ad-
ductor muscle, which is oval, and located near the buccal
margin, with its longer axis nearly at right angles to that of the
shell. At its upper extremity the small oval pedal scars are also
well defined in both valves. The posterior muscular impres-
sions broad, oval, and rather faintly marked; from near the
middle of the under side the pallial line descends with a gentle
BEEDE. | Carboniferous Invertebrates. 173
forward curve, so as to form a broad, rounded, very shallow
sinus. Length, 2.85 inches; height, from the ventral margin
to the middle of the dorsal side, 1.36 inches; length, from the
base to the line drawn from the beaks across to the most ele-
vated part of the posterior extremity, 1.50 inches; greatest
convexity, near the middle, 1.11 inches ; breadth of the posterior
hiatus, 1.07 inches ; height of the posterior hiatus, 1.44 inches.”’
Range and distribution: Upper Coal Measures; Kansas City,
Lawrence.
EXPLANATION OF PLATES.
Nore.— The page numbers printed in the following matter refer to where the description is
given. Numbers (1, 2, 3, etc.,) refer to figures on the plate, The plates will be found be-
yvinning on page 189.
PLATE I.
Fusulina secalica, (p. 10.)
1. Illustrates the different forms and sizes of these shells.
1). Longitudinal section of a specimen, showing the foramina in the inner
walls (/).
Amblysiphonella prosseri, (p. 14.)
2. Longitudinal section much enlarged, showing pores in the walls and the
thin gastral tissues. (After Clarke.)
2). Cross-section enlarged to show the structure. (After Clarke.)
2e. Longitudinal section about natural size, showing the cloaca in the center
and the apertural walls. (After Clarke.)
2d. Average-sized specimen. About natural size. (After Clarke.)
2e. Section of specimen as seen in matrix, from Topeka. Oblique longitu-
dinal section.
27. Specimen of different form, and possibly different species.
Somphospongia multiformis, (p. 12.) °
6. Drawing to illustrate the thickness of the dermal layer. Magnified.
7. Outline of larger specimen, showing mushroom form. One-half natural
size.
8. Tracing of a section through the top of one of the sponges, showing the
arrangement of the canals, which are stippled. The unstippled
portion is the body of the sponge. One-half natural size.
9. Semidiagrammatic section through the cloaca of a small individual, show-
ing canals, which are stippled, and the semiconcretionary nature of
the outer part of it.
10. Portion of the surface of a weathered specimen, showing what appears
to be the body skeleton of the sponge. Magnified.
PLATE II.
Nomphospongia multiformis, (p. 12.)
1 to 5. Photographs showing the various forms of the smaller specimens, the
last showing the manner in which the tops of the sponges weather,
bringing out the canals. Less than natural size.
Amblysiphonella prosseri, (p. 14.)
6. Specimen showing aperture. About natural size. (After Clarke.)
(175)
176 Uniwersity Geological Survey of Kansas.
Lophophyllum profundum, (p. 17.)
7. Side view of specimen, with part of the calyx broken away, showing the
protruding columella. Natural size.
7b. Section of a specimen, showing the tabulze. Natural size.
Lophophyllum westi, (p. 18.)
8. Side view of moderately large specimen. Incomplete.
8b. Cross-section of the same, showing the tendency to form a columella
from the counter septum.
Axophyllum rudis, (p. 20.)
9. Side view of the average-sized specimen. Natural size.
9b. View from above, showing the interior of the calyx and the pseudo-
columella.
9c. Cross-section of a specimen below the calyx, showing the structure of
the pseudocolumella in the center and the arrangement of the septa
and vesicles around it. Magnified. |
Syringopora multattenuata, (p. 25.)
10. Enlarged cross and oblique sections across the top of a specimen as it
appears when polished, showing the funnel-shaped tabule.
106. Longitudinal section, showing funnel-shaped tabule. Enlarged.
Cheetetes milleporaceus, (p. 25.)
11. Longitudinal section, showing tabule and arrangement of the corallites.
Magnified.
115. Cross-section of the same, showing shape of the corallites. Enlarged.
Michelinia eugene, (p. 21.)
12. Side view of average specimen. Natural size.
126. View of base of another specimen, showing its attachment to a crinoid
stem.
PLATE III.
Cladochonus bennetti, (p. 24.)
1. Side view of type. Enlarged two diameters.
Aulopora? prossert, (p. 23.)
2. View from above. Natural size.
Aulopora anna, (p. 23.)
3. Top view, showing the way in which it anastomoses. Natural size.
Lophophyllum westi, (p. 18.)
12. Side view of type. About natural size.
PLATE IV.
Campophyllum torquium, -(p. 19.)
1. Typical slab of these corals. Reduced in size.
Aulopora? prosseri, (p. 23.)
2. View of base, showing how the individual expands, and the absence of
tabule and septa. Reduced in size.
BREDE. |
Carboniferous Invertebrates. 177
PLATE V.
Campophyllum torquium, (p. 19.)
1. Cross-section of young individual, showing the thickening of the septa on
one side. Natural size.
2. Cross-section near the calyx of an older individual, showing the septa
nearly equal, except the counter septum, which is thickened.
Natural size.
3. Longitudinal section of a rather young individual, showing the compara-
tively simple tabule and a narrow vesicular zone.
4. Side view of young individual, showing a rapidly expanding form. Twice
natural size.
Limopteria alata, (p. 130.)
5. View of right valve. Twice natural size.
Syringopora multattenuata, (p. 25.)
- 6. Side of specimen. Nearly natural size.
Lophophyllum westi, (p. 18.) |
7. Longitudinal section of specimen, showing nature of the tabulex.
Enlarged.
Trachypora austini, (p. 22.)
3. Lateral view of specimen. (After Worthen.)
PLATE VI.
Zeacrinus?? robustus, (p. 29.)
1. Base of calyx of type. Natural size.
la. Side view of same (inverted).
Neaphiocrinus? washburni, (p. 27.)
2. Azygous side of calyx, showing base of arms. Natural size.
2a. Opposite side of type. Natural size.
Bupachycrinus magister, (p. 40.)
3. Side view of calyx, showing the anal plates on the right. Natural size.
3b. Base of calyx. Natural size.
Krisocrinus typus, (p. 39.)
4. Base of calyx. Natural size.
4b. Side view of calyx. Natural size.
Ceriocrinus hemisphericus, (p. 34.)
5. Base of calyx. Natural size.
5b. Side view of same.
Ceriocrinus missouriensis, (p. 35.)
6. View of the base of the calyx.
Trachypora austini, (p. 22.)
7. Cross-section. (After Worthen.)
7b. Section showing microscopic structure. (After Worthen.)
Agassizocrinus carbonarius, (p. 45.)
8. Side view of calyx.
178 University Geological Survey of Kansas.
Ceriocrinus craigi, (p. 32.)
9. Base of calyx. Natural size.
9b. Side view of same.
Phialocrinus magnificus, (p. 36.)
10. Side view of specimen, showing calyx, arms, and anal tube. Natural size.
PLATE VII.
Erisocrinus megalobrachius, (p. 37.)
la. View of base of type. Natural size.
16. Side view of same.
Ceriocrinus? monticulatus, (p. 33.)
2. Side view of type. Natural size.
Oligoporus? minutus, (p. 49.)
3. Apical? view. Natural size. Type.
FHydreionocrinus kansasensis, (p. 42.)
4 to 7. Showing structure and appearance of various parts. (After Weller.)
H. subsinuatus. See plate VIII.
PLATE VIII.
Orbiculoidea missouriensis, (p. 59d.)
land 1). Upper and lower aspects of the flat valve. Enlarged.
lc. Convex valve as seen from above. Enlarged.
Orbiculoidea manhattanensis. (p. 56.)
2. Internal appearance of the fiat valve, showing the peripheral ridge.
ural size.
2b. Convex valve. Enlarged.
Orbiculoidea convexa, (p. 5d.)
3. Convex valve. About natural size.
3b. Flat valve, perhaps of this species, internal view. Natural size.
Crania modesta, (p. 57.)
4, Upper view of specimen on a smooth shell, Natural size.
Lingula mytiloides, (p. 54.)
5. Specimen. Natural size.
Archeocidaris agassizi, (p. 48.)
6 to Gc. Illustrations of plates and spines.
6e cross-section of 6d. All natural size. (After Hall.)
Archeocidaris megastylus, (p. 49.)
7. Mass of spines and plates. (After Keyes.),
Derbya bennetti, (p. 59.)
8. View of brachial valve.
86. View of hinge area.
8c. Lateral view of specimen. (All after Hall.)
Archeocidaris trudifer, (p. 47.)
10. Spine. Natural size. (After White.)
Nat-
BEEDE. | Carboniferous Invertebrates. 179
Derbya crassa, (p. 62.)
11, 116. External] and internal views, aepEPoHvaly: Natural size.
Derbya biloba? :
12. Specimen, perhaps of this species, from Lecompton. (Rogers. )
Derbya keokuk, (p. 63.) .
13. Brachial valve. Natural size.
Hydreionocrinus subsinuatus, (p. 438.)
14. Basal view of calyx. Natural size.
PLATE IX.
Chonetes granulifer, (p. 69.)
1. External view of pedicle valve. Natural size.
1). Internal view of brachial valve, showing radiating ridges, indications of
the brachial areas, and the granular inner surface, |
le. External view of a peculiar specimen of this species, showing the auricu-
lation. Natural size.
Chonetes glaber, (p. 68.)
2. Exterior of pedicle valve. Natural size.
Chonetes mesolobus, (p. 71.)
3. Pedicle valve. Natural size.
3). Interior of brachial valve, showing the cardinal process, brachial and
other markings. Enlarged.
Chonetes verneuilianus, (p. 72.)
4. Pedicle valve of long-eared variety. (After Meek.)
4). Same of short-hinged specimen. Natural size.
4c, Interior of brachial valve, showing the markings. . Natural size.
de. Interior of pedicle valve, showing adductor and diductor sears. Natural
size.
Productus pertenuis, (p. 83.)
5. Pedicle valve. Natural size.
5b. Brachial valve, interior view, markings removed. Natural size.
5c. Longitudinal section, showing the relation of the valves in this species.
Productus symmetricus, (p. 86.)
6. Interior of brachial valve, showing adductor markings, lateral ridges, and
mesial ridge. Natural size.
6). Pedicle valve, partially covered by matrix. Natural size.
Productus nebrascensis, (p. 84.)
7. View showing the curvature of the beak and the relation of the pedicle
to the brachial (or flatter) valve. Natural size.
7 and 7d. Lateral and full view of pedicle valve of the same specimen.
7c. Cast of specimen, showing adductor and diductor sears. Natural size.
7e. Pedicle valve, showing the nature of the spines when not removed.
Natural size.
7f. Interior view of brachial valve, showing the cardinal process, large pit
immediately back of it, mesial septum, adductor markings, and a
dim outline of the brachial marks. Natural size.
180 University Geological Survey of Kansas.
Productus costatus, (p. 79.)
8. View of concave side, showing the relation of the two valves. Natural
size.
Productus longispinus, (p. 81.) 2
9. Interior of brachial valve of very old specimen. The lateral ridges are
well developed and extend entirely around the valve, forming a very
prominent feature. Natural size.
95. Brachial valve, showing relation of that valve to the pedicle. Se
Natural size.
9c. Pedicle valve, spines removed. Natural size.
9d. Interior of brachial valve of younger specimen (full grown), showing the
lateral ridges developed along the hinge only, with slight turn to
the front where they fade out. Natural size.
Hustedia mormoni, (p. 103.)
' 10. View of brachial valve.
106. Lateral view.
10c. View of pedicle valve. All natural size.
10d. Surface enlarged to show punctures in shell.
PLATE X.
Productus costatus, (p. 79.)
1. Pedicle valve, showing the ridge between the umbo and the ears on which
are spines. Natural size.
1b. Side view of same. |
le. Interior of brachial valve, showing adductor markings, cardinal process,
mesial ridge, lateral ridges, and brachial markings. The border in
front is matrix between the brachial and pedicle valve. Natural
size.
Productus semireticulatus, (p. 78.)
2. Exterior of brachial valve, showing hinge and beak. Natural size.
2b. Pedicle valve. The umbo of this specimen is narrower than most of
the Coal Measures forms, which are probably varietally distinct
from the one figured, which is from the Permian, but is figured here
on account of its being a much better specimen. Natural size.
2c. Interior of pedicle valve, showing adductor and diductor UE IE The
central ones are the adductors. Natural size.
2d. Interior of brachial valve, showing adductor, brachial and other mark-
‘ings. Natural size. (Small specimen.)
Productus punctatus, (p. 87.)
3. Pedicle valve. Natural size.
3b. Brachial valve, exterior view, showing the beak and hinge. Natural
size. ;
3c. Lateral view of the same specimen..
3d. Pedicle valve of young specimen. Natural size.
3e. Cast of interior of the pedicle valve, showing the adductor and diduetne
scars. Natural size.
BEEDE. | Carboniferous Invertebrates. 181]
‘ PLATE XI.
Productus coray (p. 75.)
1. Pedicle valve of the common form. Natural size.
1b. Cast of pedicle valve. (After Hall.)
le. Brachial valve, interior view, showing traces of brachial markings.
Natural size.
ld. Pedicle valve of large specimen. Natural size.
le. Spine as seen on a slab of shale, from a large specimen. Natural size.
17. Pedicle valve of broad, large variety. Natural size.
Productus cora americanus, (p. 77.)
2. Pedicle valve of typical form. Natural size.
Productus punctatus, (p. 87.)
3. Interior of brachial valve, showing adductor scars. Natural size.
Productus costatus, (p. 79.)
4. Cast of pedicle valve, view of beak. Natural size.
Cleiothyris royssii, p. 104.)
5. Outline of specimen with nearly straight hinge.
5b. Brachial valve and beak of pedicle valve of specimen.
5c. Lateral outline of specimen. All natural size.
Aulacorhynchus millepunctatus, (p. 89.)
6. External view of one of the valves. (After Meek and Worthen.)
PLATE XII.
Aulacorhynchus millepunctatus, (p. 89.)
1. Interior of brachial valve. (After Meek and Worthen.)
1b. View of a specimen from Kansas City, showing indications of extra plat-
forms, and probably a different species. Natural size. It is quite
convex for the genus.
Cleiothyris royssii, (p. 104.)
2. Side view of more robust specimen. Natural size.
Hustedia mormoni, (p. 103.)
3. Hinge apparatus. (After Hall.)
Reticularia perplexa, (p. 102.)
4, Pedicle valve. About natural size.
4b. Opposite valve of same.
4c. Side view of same.
4d. Surface enlarged. (After Grity.)
Spirifer cameratus, (p. 99.)
5. Interior of brachial valve, showing sockets for cardinal teeth. Natural
size.
5b. Pedicle valve of long-eared form.
5¢. Brachial valve and hinge area of robust form.
5d. Young specimen. All natural size.
5e. Spire of one of this species (broken). Natural size.
182 University Geological Survey of Kansas.
Enteletes hemiplicata, (p. 91.)
6.. Side view of specimen. Natural size.
6b. Front view of same.
Pugnax utah, (p. 93.)
7. Brachial side, showing beak of the opposite valve, with the pedicle open-
ing. Natural size.
7b. Front view of same.
7c. Side view of same.
Pugnax rockymontana, (p. 92.)
8. Side view of specimen. Natural size.
8b. Front view of same.
Meekella: striatocostata, (p. 65.)
9. Lateral view.
9b. Pedicle valve.
9c. Brachial valve and hinge area.. All natural size.
Derbya cymbula, (p. 60.)
10. Interior view of hinge apparatus.
PLATE XIII.
Aviculopecten hertzeri, (p. 121.)
1. Left valve.
1b. Surface markings enlarged.
Aviculopecten providencensis, (p. 119.)
2. Left valve. Natural size.
Aviculopecten sculptilis, (p. 122.)
3. Lateral view of left valve. Natural size.
3b. End view of same.
Aviculopecten germanus, (p. 123.) 2
4. Left valve. Natural size. (Rogers.)
Lima retifera, (p. 112.)
5. Left valve of shell. Natural size.
Aviculopecten interlineatus, (p. 116.)
6. Left valve. Natural size.
Aviculopecten occidentalis, (p. 114.)
7. Left valve. Natural size.
Aviculopecten hertzeri, yp. 121.)
8. Right valve. (After Meek and Worthen.)
Aviculopecten carboniferus, (p. 117.)
9. Left valve. Natural size.
Aviculopecten mecoyi, (p. 118.)
10. Left valve. Natural size.
Pseudomonotis hawni, (p. 132.)
11. Interior view of flat valve, showing byssal notch.
116. End view of convex valve.
lle. Lateral view of convex valve. (All after Meek.)
BEEDE. | Carboniferous Invertebrates. 183
PLATE XIV.
Pseudomonotis kansasensis, (p. 133.)
1. Left valve of type. Postero-ventral portion crushed, probably giving it
undue prominence. The striv are finer and more crinkly than rep-
resented in drawing. About natural size.
la. Outline showing convexity of the same.
1b. Outline showing the convexity of the adult long-hinged forms.
le and id. Young specimen of the long-hinged form, showing different de-
grees of development of the hinge and ears. About natural size.
Pseudomonotis robusta, (p. 133.)
2. Left valve of type. The striw are finer and more wavy than represented,
and the upper third is practically glabrous, only the faintest traces
of strix being present. About natural size.
2a. Anterior view of the same shell, showing gibbosity and arcuity of shell,
as well as concentric lines of growth. About natural size.
2 and 2e. Anterior and lateral views of the left valve of a young individual
of the above shell. About natural size.
Pseudomonotis hawni equistriata, (p. 134.)
3. Left valve of type. A little less than natural size.
3a and 3b. Anterior views of two left valves, showing variation in convexity
and indistinctness of beak. About natural size.
PLATE XV.
Pseudomonotis hawni. Lobed variety, (p. 132.)
1. Convex valve, showing the bending of the costw and the lobe on the end
of the shell. AlJl natural size.
la. Interior of left valve, showing retractor scars and adductor scar.
1). Outline showing convexity of same.
le. Hinge view of a specimen, showing the indistinctness of the beak and
the great convexity of the shell.
1d. Specimen showing the stronger lobing of the shell.
le. End of outline of a specimen, showing the great convexity of the beak
in some individuals.
1f. Hinge view of a specimen, showing the great convexity and the manner -
in which the shell flattens out on the end where the lobe is located.
2. Large, old specimen, showing umbonal region with only the finer striz.
2a. Flat valve of the same specimen, showing the place of attachment to
foreign object.
Pseudomonotis kansasensis, (p. 133.)
3. Enlargement of the surface of the type, showing the lines of growth ex-
tending backward up the furrows and down farther toward the
ventral margin on the ridges.
184 University Geological Survey of Kansas.
PLATE XVI.
Myalina ampla, (p. 139.)
1. External view of shell. (After Meek.)
1b. Interior of same. Both reduced one-half.
Myalina congeneris, (p. 142.)
2. Exterior of specimen. (After Walcott.)
26. Another specimen. (After Walcott.)
Pteria sulcata, (p. 126.)
3. .Left valve enlarged. (After Meek.)
Pteria longa, (p. 125.)
4, Left valve of specimen. Natural size.
Limopteria marian, (p. 128.)
5. Right valve. Natural size. (After White.)
5b. End view of specimen. (After White.)
5c. Surface markings enlarged.
Limopteria longispina, (p. 127.)
6. (After Keyes.)
Myalina swallovi, (p. 137.)
7. Left valve. Natural size.
Myalina perattenuata, (p. 141.)
8. Right valve of specimen. Natural size.
Limopteria gibbosa, (p. 129.)
9. Right valve. (After White.)
Myalina subquadrata, (p. 1388.)
10. Impression of left valve. (After Meek.)
10b. Part of left valve broken away, showing the interior of fae right valve
and outline of the specimen. (After Meek.)
Myalina kansasensis, (p. 140.)
11. Specimen showing left valve. (After Keyes.)
PLATE XVII.
Aviculopinna illinoiensis, (p. 143.)
1. Outline of cross-section.
16. Specimen. Natural size.
le. Surface markings. Enlarged.
Pinna subspatulata, (p. 145.)
2. Drawing of cast. Reduced one-half.
Pinna peracuta, (p. 144.)
3 and 3b. Lateral and dorsal views. (After Keyes.)
BEEDE. | Carboniferous Invertebrates. 185
PLATE XVIII.
Pinna subspatulata, (p. 145.)
1. Semidiagrammatic dorsal aspect. Does not show dorsal ridge.
1b. Aspect of broken right valve.
le. Same of left valve.
ld. Cross-section. All natural size.
Aviculopinna americana, (p. 148.)
2. Lateral view of shell. Natural size.
Pinna subspatulata, (p. 145.)
3. View of another cast from different locality.
PLATE XIX.
Entolium aviculatum, (p. 113.)
1. Left valve. (After Meek.)
Aviculopecten coxanus, (p. 124.)
2. View showing left valve. Natural size. (After Rogers.)
Chenomya leavenworthensis, (p. 172.)
3. Lateral view of specimen. (After Meek.)
3b. Dorsal aspect of same.
Myalina? exasperata, (p. 141.)
4. Left? view of type. Natural size.
Posidonomya? pertenuis, (p. 136.)
5. Left? valve of type.
Posidonomya? recurva, (p. 135.)
6a. Left? valve of type.
656. Right? valve of another specimen.
6c. Left? valve of the preceding, with a part of the shell adhering on one
side. The beak is somewhat crushed and twisted, so that it does
not seem to extend above the hinge. All natural size.
PLATE XX.
Allorisma subcuneatum, (p. 169.)
1. Lateral view of specimen. (After Keyes.)
14. Dorsal aspect of the same.
Macrodon sangamonensis? (p. 146.)
2. Dorsal view of left valve.
2b. Lateral view of same. Natural size.
Sedgcewickia topekaensis, (p. 171.)
3. Left valve of specimen.
Yoldia knoxensis? (p. 154.)
4. Specimen with valves distended. By mistake, this is not the specimen
described in text, and may possibly be a different species.
Edmondia nebrascensis, (p. 166.)
5. Left valve of specimen. Natural size.
13—Vvi
186 University Geological Survey of Kansas.
Allorisma geinitzi, (p. 167.)
6. Right valve of specimen. Natural size.
Pleurophorus tropidophorus, (p. 162.)
7. View of right valve. Natural size.
Yoldia subscitula, (p. 152.)
8. Right valve. Natural size.
Conocardium parrishi, (p. 164.)
9. Left valve. (After Keyes.)
Allorisma granosa, (p. 168.)
10. Left valve of specimen. Natural size.
Pleurophorus subcostatus, (p. 161.)
11. Cast of left valve. (After Meek and Worthen.)
116. Dorsal aspect of the same.
Allorisma costatum, (p. 170.)
12. Right valve of specimen. Natural] size.
Macrodon obsoletus, (p. 147.)
13. Right valve of specimen. (After Meek.)
Nuculana bellistriata, (p. 148.)
14. Dorsal view of specimen. Natural size.
146. Left valve of same.
Nucula ventricosa, (p. 150.)
15. Exterior view of specimen. Natural size.
Cypricardinia? carbonaria, (p. 164.)
16. Left valve. Natural size.
Cardiomorpha missouriensi?s, (p. 165.)
17. Right valve. Natural size.
Placunopsis carbonaria, (p. 111.)
18. Lateral view of specimen. (After Meek and Worthen.)
PLATE XXI.
Solenomya parallela, (p. 158.)
1. Side view of type.
Solenomya trapezoides, (p. 159.)
2a. Left valve. Anterior adductor scar and pallial line a little too strong.
3b. Dorsal view of another specimen.
Limopteria subalata, (p. 131.)
3a. Side view of left valve.
3b. Side view of right valve. Both natural size.
Yoldia glabra, (p. 153.)
4a. Side view of type. Natural size.
4. Surface markings. Enlarged.
Nucula pulchella, (p. 151.)
5a. Side view of type. Enlarged.
5b. End view of same.
5e. Surface markings. Greatly enlarged.
BEEDE. | Carboniferous Invertebrates. 187
PLATE XXII.
NSchizodus wheeleri, (p. 155.)
1. Dorsal view of cast. (After Keyes.)
1). Side view of specimen with shell on.
le. Side view of No. 1.
Nehizodus hari, (p. 155.)
2. End view of specimen. Natural size.
2b. Left hinge.
2e. External view of right valve.
2d. Internal view of same.
Edmondia aspenwallensis, (p. 166.)
3. Lateral view of right valve. (After Keyes.)
3). Dorsal aspect of same.
Solenomya radiata, (p. 160.)
5. Lateral view of specimen. (After Meek and Hayden.)
5b. Dorsal view of same.
Nchizodus compressus Rogers. N.sp., (p. 157.)
6. Internal apparatus of a worn right valve.
65. Same of left valve.
6c. Outline showing convexity of shell.
6d. Outer appearance of better specimen. All natural size.
Nuculana bellistriata attenuata, (p. 149.)
7. Lateral view. Enlarged.
7b. Surface markings. Enlarged.
Nucula beyrichi, (p. 149.)
8. Side view of specimen. (After Meek.)
Nucula ventricosa, (p. 150.)
9. Cast showing adductors and pallial line. Natural size.
95. External appearance of shell.
Astartella vera, (p. 163.)
10. View of right valve of specimen.
CORRECTIONS.
Page 18: Following Lophophyllum westi, after fig. 12, add ‘‘ plate V, fig. 7.”
Page 24: Following Cladochonus? bennctti, after fig. 1, erase ‘‘ plate V, fig. 7.”’
Page 43: After 17ydreionocrinus subsinuatus, instead of ‘‘ Plate VII, fig. 14,”’
read ‘* Plate VIII, fig. 14.”’
Page 78: Following Productus semirecticulatus, after figs. 2-2d, instead of
“text fig. 2, f,’’ read ‘text fig. 3, f.’’
Page 103: Following Hustedia mormoni, for “plate X, fig. 3,’’ read ‘‘ plate
XIT, fig. 3.”
Page 149: After Nucu/ana Lellistriata attenuata, add ‘*‘ Plate XXII, figs. 7, 7b.”
sie as
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University Geological Survey of Kansas. VOLUME VI. PLATE I,
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SOM PHOSPONGIA, 1-5. LOPHOPHYLLUM, 7, 8.
AMBLYSIPHONELLA, 6. AXOPHYLLUM, 9.
SYRINGOPORA, 10.
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CLADOCHONUS, 1. AULOPORA, 2, 3. LOPHOPHYLLUM, 12.
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University Geological Survey of Kansas. VOLUME VI. PLATE IY.
PLATE V
VoLuME VI.
University Geological Survey of Kansa
SYRINGOPORA, 6.
TRACHYPORA, &.
IMOPTERIA, 5.
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1-4.
CAMPOPHYLLUM
LOPHOPHYLLUM,
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University Geological
Survey of Kansas.
ZEACRINUS, 1. EUPACHYCRINUS, 3.
SCAPHIOCRINUS, 2. ERISOCRINUS, 4.
CERIOCRINUS, 5, 6.
VOLUME VI.
TRACHYPORA, 7.
PLATE VI.
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University Geological Survey of Kansas. VouuME VI. PLATE VII.
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DERBYA, 8, 11-13.
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VouLuME VI.
University Geological Survey of Kansas.
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SOLENOMYA, 1, 2. LIMOPTERIA, 3. YOLDIA, 4.
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PART ITI.
CRETACEOUS FISHES.
SELACHIANS AND PYCNODONTS,
By S. W. WILLISTON. » :
TELEOSTS,
AS ‘ ‘
By ALBAN STEWART.
—-
Plates XXIII to LXXIII. 4
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SELACHIANS AND PYCNODONTS.
By S. W. WILLISTON.
Tue following notes and descriptions of selachian and
pycnodont teeth are based upon the material that has accumu-
lated in the University of Kansas Museum during the past ten
or twelve years, supplemented by a collection kindly loaned for
study by Mr. T. W. Stanton, of the National Museum. The
material is by no means exhaustive, nor even sufficient to settle
several doubtful points, but I trust that, incomplete as it may
be, it will be of service in the determination of our numerous
forms.
‘*'The specific determination of the detached teeth of sharks
and skates is little more than guesswork, and to decide upon
their generic relationships with any approach to certainty is
also often very difficult.’’' Nevertheless, because such de-
tached teeth are so often found, and connected series so rarely,
an attempt at their determination is desirable. Fortunately,
in the present collection there are several forms represented by
such complete specimens that the positive addition they afford
to the knowledge of the species and genera is very welcome.
MYLIOBATID.
PTYCHODUS.
Teeth with the crown more or less elevated and overhanging,
ornamented with transverse or radiating ridges, and surrounded
by a larger or smaller, finely marked area. Surface of root
smooth.
This genus of Upper Cretaceous selachians was for a long
time placed among the cestracionts, but recent discoveries of
the nearly complete dentition render it more probable that its
1, Woodward: Proc. Geol. Assoc., XIII, p. 190.
(237)
238 University Geological Survey of Kansas.
proper location is with the Myliobatide. The living members
of the family, the sea-devils, are broad, flat fishes, allied to the
rays, with a disk-hike body. Many attain an enormous size,
fifteen or twenty feet in length, and weigh a thousand pounds
or more. In some the pectoral fins take on almost the charac-
ter of limbs, and are said to be used in scooping up their food
and transferring it tothe mouth. The teeth are flat and pave-
ment-like, and are used for crushing crabs and shell-fish. The
fish are viviparous, and for the most part live in tropical or
semitropical waters.
The teeth in Ptychodus are not less than 600 in number in
each jaw, at least in some species. They are arranged in par-
allel rows, decreasing in size from within outward, except that
in the supposedly upper jaws the median row is composed of
small, low and smooth teeth, very much unlike the immediately
adjacent ones. In P. mortona there are eight rows on either
side of this median row, or seventeen in all. The lateral teeth
become more transversely elongated, the surface markings less
conspicuous, and the form more unsymmetrical. About fifteen
species of the genus have so far been discovered, all from the
Upper Cretaceous. One or two species, including our most
common one, haye been discovered in both Europe and North
America, and it is not improbable that the identity of yet others
will be established when they are better known. The teeth vary
so much in size and shape in the same individual that the iden-
tification from single specimens is often impossible or a matter
of great uncertainty.
Ptychodus mortoni. Plate XXV; plate XXVI, fig. 1; plate XX VII.
Ptychodus mortoni (Mantell) Morton, Journ. Acad. Nat. Sci. Phil., vim,
p- 215, pl. x, f. 7; Agassiz, Poiss. Foss., 111, p. 158, pl. xxv, ff. 1-3; Leidy,
Proc. Acad. Nat. Sci. Phil., (1868), p. 205; Ext. Vert. Fauna, p. 295, pl.
xvill, ff. 1-14; Cope, Cret. Vert., p. 294; Woodward, Quart. Journ. Geol.
Soc., xLit, p. 130; Cat. Foss. Fishes Brit. Mus., 1, p. 159; Proc. Geol.
Assoc., xit1, p. 191, pl. v, f. 4; Williston, Kans. Univ. Quart., 1x, p. 30, pl.
a vin f. 1, pl. 1x— Alabama, Mississippi, Niobrara of Kansas, English
alk.
This species is the most common one of this genus in the
Kansas Cretaceous, occurring only in the Niobrara beds, so far
as I am aware, and, for the most part at least, in the lower part
WILLISTON. | Cretaceous Fishes. 239
of the beds. I have before me at the present time two excellent
series of teeth of this species ; one, including about eighty teeth,
obtained from the estate of the late Joseph Savage; the other
collected in the vicinity of Castle Rock, in Trego county, by
Prof. G. E. Rose—an exceedingly interesting specimen, be-
cause most of the teeth are in place in the matrix. A number
of the teeth of the Savage specimen have been arranged serially
and photographed in plate XX V. Of course the arrangement
is not the natural one, but the plate will show in an excellent
way many of the characters of the teeth better than they can
be described. In plates XXVI and XXVII are given three
views of portions of the Rose specimen; that of plate XX VI
(fig. 1) shows a little more than one-half of the upper view.
One end (the left of the figure) has been folded underneath
obliquely. This folded end is shown in plate XXVIII, fig. 1.
Figure 2 of the same plate gives a view of a transverse series,
as arranged from the loose teeth taken from the right end of the
specimen—the one that protruded from the chalk when discov-
ered. About 550 teeth, all told, have been obtained, and doubt-
less not a few had been lost before the specimen was discovered.
The set is referred to the upper jaw, on the supposition of
Woodward that the small median teeth belong in this jaw.
Not a trace of osseous substance is preserved in the specimen.
The cartilage of the sharks’ jaws is often preserved in a soft,
calcified condition, but it is evident that the material in which
the teeth of Ptychodus were lodged was of a more perishable
nature, accounting doubtless for the fact that Ptychodus teeth
are so rarely found associated.
The teeth of this species differ markedly from those of all
other known species, in having the center of the crown raised
into a conical apex, the summit of which is crossed by a short
transverse ridge from which other diverging ridges run. In
the smaller lateral teeth these ridges become less well marked
and occupy a relatively smaller space, becoming almost obsolete
in the fifth row. The marginal area is formed of fine reticula-
tions in many of the larger teeth, though in most of these and
in all the smaller teeth the markings are more like a fine punc-
240 University Geological Survey of Kansas.
tulation, clearly visible only with the aid of a lens, giving a
uniform, finely roughened appearance. The median upper row
is composed of low, flattened teeth, transversely oval or sub-
quadrate in shape, with a slight elevation in the middle, and
finely roughened throughout the whole coronal surface, there
being only the slightest trace of the divergent ridges on the
summit of the elevation. This does not quite agree with Wood-
ward’s description of these teeth, in which he states that they
are ‘‘not marked with the radiating ridges, but exhibit a mi-
nute smooth eminence in the middle of the crown.’’ Possibly
this effect is due to wear.
Ptychodus polygyrus. Plate XXIX, fig. 9; plate XXX, fig. 14.
Ptychodus polygyrus (Buckland) Agassiz, Poiss. Foss., 111, p. 156, pl. xxv,
ff. 4-11, pl. xxv-s, ff. 21-23; Gibbes, Journ. Acad. Nat. Sci. Phil.,1, p. 299,
pl. u, ff.5,6; Leidy, Proc. Acad. Nat. Sci. Phil. 1868, p. 208; Cope, Cret.
Vert., p. 294; Woodward, Cat. Foss. Fishes Brit. Mus.,1, p. 148, pl. v,
f. 7—Senonian, Turonian of Europe, Rotten Limestone of Alabama,
Niobrara of Kansas; Williston, cf. cit. 31.
? Ptychodus latissimus Agassiz, |. c., fig.8; Dixon, Foss. Sussex, pl. xxx,
1
A single tooth of very large size from the lower beds of the
Niobrara Cretaceous of the Smoky Hill river is referred to this
species provisionally. Until numerous specimens are examined
there can be no certainty of its correct location, though the re-
semblances are sufficiently great to render the determination
not improbable; at least with some of its varieties.
Ptychodus martini. Plate XXVIII.
Ptychodus martini Williston, ef. cit. 32.
A large series of teeth, 110 in number, found together in the
Niobrara chalk of the Smoky Hill river, and collected by Mr. H.
T. Martin, cannot be identified with any described species. I
have photographed them, arranged as symmetrically as possible,
but with no assurance that the arrangement is a natural one.
In fact, it is not improbable that the teeth belong to both upper
and lower jaws. The teeth apparently from the lower median
row are much elongate transversely, with a very flat crown,
wherein they differ from the teeth of other known species. The
ridges are nine or ten in number, and reach nearly to the lateral
a
WILLISTON. | Cretaceous Fishes. 241
margin. In some of the teeth several of the ridges form loops
near the extremities. The marginal area of granulations is
small, and presents scarcely any distinct vermiculations. The
teeth of the lateral rows are less elongated than those of the
middle one, though still more so than is usual. The granula-
tions become rather more extensive in area proportionally in
the small teeth, as is the case with other species. A _ series
(left vertical row of the plate) that may belong in the medio-
lateral rows of the upper jaws are more nearly square in‘shape,
and the crown has a distinct, though low, convexity extending
over nearly its whole area. Antero-posteriorly the surface is
nearly flat, with a moderate convexity of the margin. The
surface posterior to the large grooves on the upper part shows
small, radiating and branched ridges.
The largest teeth measure 45 by 20 mm.; the ones more
nearly square, 39 by 25 mm.
Ptychodus anonymus. Plate XXIX, figs. 5-8, 16-18, 20-22, 24.
Ptychodus anonymus Williston, ef. cit. 32.
Seven teeth of nearly uniform size, four of them united in
the matrix, from Walnut creek, Kansas, seem to belong to a
species distinct from any previously described (figs. 16-18).
They are of about the same size as those described as P. whip-
pleyi and P. occidentalis, but will be distinguished from the
former by the more broadly conical crowns. In the teeth of
this size of P. whippleyi the crown is much compressed, stand-
ing up, tooth-like; in the present specimens they are nearly
straight or gently concave from the apex to the rims. [rom
P. occidentalis the species will be distinguished by the very dis-
tinctly reticulate marginal areas, the transverse ridges not
reaching to the rims of the crown. Other specimens agreeing
in these characters are from the Niobrara. The horizon is
probably Benton.
242 Uriversity Geological Survey of Kansas.
Ptychodus occidentalis. Plate X XIX, fig. 4; plate XXX, fig. 13.
Ptuchodus occidentalis Leidy, Proc. Acad. Nat. Sci. Phil. 1868, p. 207;
Ext. Vert. Fauna West. Terr.. p. 308, pl. xvi, ff. 7, 8, pl. xv1it, ff. 15-18;
Cope, Cret. Vert., p. 244; Williston, cf. cit. 33—Niobrara, Benton of
Kansas.
Two teeth, one from the same conglomerate that yielded the
teeth referred to P. janewayz?, the other, without locality, from
Mr. Joseph Savage’s collection, I refer to this species. The
species differs from the following in having the transverse
ridges continued to the lateral rims, and not separated by an
area of fine reticulation. The anterior surface has finer, elon-
gated, nearly straight ridges and grooves in this species, while
in the others the markings are reticulate or vermiculate.
Ptychodus janewayii. Plate XXX, figs. 9, 10, 11.
Sporetodus janewayii Cope, Hayden’s Bull. U. S. Geol. Surv. No. 2,
(1874). pl. xivit.
Ptychodus janewayti Cope, Cret. Vert., p. 244; Williston, cf. cit. 33.
‘‘Surface irregularly convex, covered with a dense layer,
which does not exhibit pores, and is thrown into transverse or
oblique ridges. Surface with four folds, which traverse it
obliquely from border to border. At the base of the outer, at
one end, is a series of adherent tubercles; at the basis of that,
at the opposite end, is a broken fold, with tubercles at its outer
base. Length, 0.0045 m.; width, 0.0025 m. A portion of a
larger and more central tooth has the surface with an unsym-
metrical convexity, and is crossed transversely by five folds,
from border to border.’’? |
Three small teeth, shown enlarged in plate XXX, ff. 9-11,
from the conglomerate containing specimens of Corax curvatus,
appear to belong to this species. The horizon of the conglom-
erate is near the line of contact between the Dakota and Ben-
ton, in Ellsworth county. Cope’s type was from a bed of
conglomerate containing Lamna and Isurus teeth of small size
near Stockton. It is probable that the horizon is the same in
both.
2. Cope, l. c.
Penny. wae
WILLISTON. | Cretaceous Fishes. 243
Ptychodus whippleyi. Plate XXIX, figs. 10-15.
arp ion whippleyi Marcou, Geol. North Amer., p. 33, pl. 1x, f.4; Leidy,
Ext. Vert. saetbing p. 300, pl. xvitl, ff. 19, 20; J. S. Newberry, Rep.
Expl. Exp., p. 147, pl. 111, f. 2: Cope, Cret. Vert., p. 294: Williston, cf. cit.
34— Cretaceous, Texas | Marcou, Leidy): Kansas, Arkansas Valley (Cope):
Colorado, New Mexico.
Thirteen teeth from Dallas, Tex., and a number of others re-
ceived from Mr. Frank Springer, collected in the vicinity of
Las Vegas, in New Mexico, agree with the descriptions and fig-
ures of this species, as given by Leidy. The same species has
been referred to the Niobrara chalk of the Arkansas valley by
Cope. If his determination and locality are correct the species
must be referred to the Benton of Kansas, since the Niobrara
does not occur in the Arkansas yalley. A single specimen from
the Benton of Kansas in the museum, without definite locality,
seems to agree pretty well with the Texas specimens, but the
specimen is an uncharacteristic one and may pertain to some
other species.
Some of the teeth referred to this species show a marked re-
semblance to those figured by Woodward? (P. rugosus) , and by
Dixon (P. altior Dix.)
The European species is described as having the sides of the
median elevation of the crown smooth, which is not the case
with the present species, the grooves continuing midway into
the lateral granulations.
e Ptychodus, sp. Plate XXIX, figs. 2,3: plate XXXI, fig. 53.
Ptychodus, sp., Williston, ef. cit. 34.
Four teeth of moderately large size, from the Benton Creta-
ceous, of Salt creek, Russell county, and two others of smaller
size, also from the Benton, seem to belong to a species distinct
C from any hitherto known. The larger ones will be distinguished
from those referred to the upper series of P. martini, which are
of nearly the same size and shape, by the smaller area of trans-
verse ridges, and the much larger area of marginal reticula-
tions, which are coarser. The teeth are more nearly square and
the convexity of the crown is greater. The two teeth of smaller
ee
>
y
%
.
3. Cat. Foss, Fishes Brit. Mus., I, pl. V, fig. 2.
ee
244 University Geological Survey of Kansas.
size probably belong with the others. It is possible that some
of these teeth may belong with P. polygyrus.
The other described species of this genus are the following:
Ptychodus mammilaris Agassiz.—Senonian, Turonian, and Cenomanian,
Europe.
Ptychodus rugosus Dixon.—Senonian, England.
Ptychodus decurrens Agassiz.—Senonian, Turonian, and Cenomanian,
Europe.
Ptychodus multistriatus Woodward.— Senonian and Turonian, England.
Ptychodus latissimus Agassiz.—Turonian and Senonian, Europe.
Ptychodus papillosus Cope, Cret. Vert. 294.— Upper Cretaceous, Colorado.
Ptychodus triangularis Reuss.— Upper Cretaceous, Bohemia.
Ptychodus levis Woodward.— Lower Chalk of England.
SCYLLIIDA.
The family Scylliide comprises small sharks with sharp-
pointed cuspidate teeth, arranged in numerous series. The fol-
lowing genera are given by Woodward :* Paleoscyllium Wagner,
Lower Kimmeridgian of Bavaria; Scyllcorhinus Blainv., Turo-
nian and Senonian ; Pristvwrus Bonaparte, Lower Kimmeridgian
of Bavaria; Mesiteia Kramb., Senonian and Middle Eocene ;
Chiloscylliwm Miller and Henle, Molasse; Crossorhinus Miller
and Henle, Gault; Cantioscyllium Woodw., Turonian; Gingly-
mostoma Miller and Henle, Danaian, Eocene.
Numerous teeth from the Lower Cretaceous of Kansas seem
in all probability to belong in this family, and agree pretty well,
though rather large, with the teeth of Scylliorhinus, to which I
refer them provisionally.
Scylliorhinus rugosus. Plate XXIV, fig. 5.
Seyllium rugosum Williston, Kans. Univ. Quart., rx, p. 39.
Central cusp broad, pointed, nearly symmetrical, the cutting
edges nearly straight, one of them a little longer than the other
and slightly convex near the tip; a single pair of lateral den-
ticles, which are nearly equilaterally triangular in shape; prin-
cipal cusp with six or seven strong ridges on the basal two-fifths ;
denticles with four or five similar ridges reaching two-thirds of
4, Cat. Foss. Fishes Brit. Mus., I, p. 338.
WILLISTON. | Cretaceous Fishes. 245
the way to the apex; root narrow, apparently not at all pro-
duced at the angles; thinned and not at all tumid.
Type No. 1949, U.S. National Museum, Greenleaf sandstone
at Greenleaf ranch.
eae URED NADIR EEE AORSERID VaR al p oly icin ain, tyes osm v6 oe omg we os Se 7mm.
NUMER MENER TER SERRE tee ey ook vlc bic vale Sse cee eet A
Height of denticles ........ et et Se On eee ee er re oes
MRMREET RMN Ree POE ratte eo oe Sav Ach os bea ade eae 24 **
Scylliorhinus planidens. Plate XXIV, fig. 7.
Seyllium planidens Williston, Kans. Univ. Quart., rx, p. 35.
Central cusp broad, pointed, convex from side to side, with
sharp, non-crenulate edges; lateral cusps sharply pointed,
smooth, two in number; root thin, narrow, moderately pro-
duced below the posterior denticle, smooth.
C2 SS COP 2 0, en 4mm.
OES EE SEES ATO og 21S 2a SR re ie ae ee ra anes
REVS SESE SE) ea] ss OO ied ag Sn 2 eal
(SPR SDS) oe Seay re ee a ere 1} «
Type No. 1949, U. S. National Museum. From same horizon
as the preceding species.
I refer provisionally to this species numerous other speci-
mens from the same horizon and collection. They differ in the
relative size of the denticles, the more posterior direction of the
main cusp, and the size. One tooth seems to lack the anterior
denticle, which is always the smaller of the two; its absence
may be due to injury.
Scylliorhinus (Lamna?) gracilis. Plate XXIV, fig. 6.
Scyllium (Lamna?) gracilis Williston, Kans. Uniy. Quart., 1x, p. 35.
Main cusp elongate, slender; inner surface smooth, gently
convex longitudinally, more so transversely, with sharp, smooth
edges ; the interior edge nearly straight, the posterior somewhat
concave; denticles of nearly equal size, small, slender, acute ;
base narrow, prolonged into a slender root at each extremity.
Se MMRR ME se cE dle du d's as hud) cee oislbr gh GreRaweene neue 9mm.
Meee RT OMEs RERME MCRL UII 2, (coals eos a cla led du Rice a tidak dl ea
RE CMI AUIS 6 oo oor. ok ha'n se ws hE ek ee eee eh vie 5 ak
MUI CO CMPAURIOUNERI oo as Sos ch bee 'ck oh vot mtsd Pea aee tu ok mite
One specimen, No. 1949, U.S. National Museum, with the
preceding species.
246 University Geological Survey of Kansas.
LAMNID.
The Lamnide comprise the largest and most voracious of the
sharks, represented by a number of species in the oceans of the
present time. They are elongated fishes, the dorsal fin without
spine; there is no nictitating membrane to the eye, and the gill
Openings are wide. The teeth are solid in the adult, and are
300 or more in number. The teeth are found very commonly
in the Cretaceous deposits of Kansas, as elsewhere, usually
scattered singly, though occasionally found more or less con-
nected by the calcified cartilage of the jaws in several rows.
Owing to the great variation of size and shape of the teeth in
the same individual, it is often difficult or impossible to cor-
rectly determine the forms. Doctor Eastman has recently fig-
ured and described the nearly complete dentition of Isurus
manteli, the most common species of the family in Kansas.
Doubtless similar variations will be found in the different spe-
cies of the other genera of this family.
ISURUS.
This genus differs from Lanna only in the prevailing absence
of the lateral denticles of the teeth. The teeth are large. The
genus occurs from Jurassic to the present time.
Isurus mantelli. Plate XX XI, figs. 41-46; plate XXXII, figs. 2-2m.
Oxyrhina mantelli (Geinitz) Agassiz, Poiss. Foss., 111, p. 282, pl. xxx111,
ff. 1-5, 7-9; Eastman, Paleontographica, xi, pp. 149-192, pll. xvi—xviit
(where additional extensive synonymy will be found); Woodward, Proc.
Geol. Assoc., x1II, p. 196 Cenomanian, Senonian and Turonian of Eu-
rope; Kansas, Texas, New Jersey, Alabama, Colorado, etc.
Oxyrhina extenta Leidy, Ext. Vert. Fauna, p. 302, pl. xviii, ff. 21-25.
‘‘ Moderate-sized, stout, three-cornered teeth; the crown on
the outer side nearly flat, with one or more vertical wrinkles ;
on the inner side, lightly convex and smooth ; root long, thick,
low, moderately deeply furcate, usually obtuse at the ends, and
on both sides more or less flattened.’’®
This species is very common in the Kansas Niobrara, in fact,
5. Eastman, l.c.
WILLISTON. | Cretaceous Fishes. 247
the most common of all, and not infrequently it is represented
by many associated teeth. From the plates, and from East-
man’s figures, it will be readily identified in all its forms.
LAMNA.
Teeth, except some of the hindmost ones, with a narrow,
compressed, conical cusp, with one or two pairs of small, pointed
denticles.
Some of the following species may belong to Odontaspis,
which can hardly be distinguished by the teeth alone, differing
only in the relatively less high and less subulated character of
the anterior ones, and in the usually larger size of the lateral
denticles.
Lamna appendiculata. Plate XXVI, figs. 3-3c; plate XXXI, figs. 47-49.
Otodus appendiculatus (Roemer) Agassiz, Poiss. Foss., 111, p. 279, pl.
xxx, ff. 1-25; Davis, Trans. Roy. Dubl. Soc., rv, p. 402, pl. xu, ff. 1-11.
Lamna appendiculata Woodward, Cat. Foss. Fishes Brit. Mus., 1, p. 393;
Proc. Geol. Assoc. x11, p. 196: Williston, cf. cit. 37 —Senonian, Cenoma-
nian, Turonian (?), Danian of Europe, Niobrara of Kansas, and Green-
sand of New Jersey.
“Teeth robust, with a thick root, having a much flattened
postero-inferior face, the nutritive foramen not in a groove.
Outer face slightly convex or flat, often with a few indefinite
vertical folds on the basal half; inner side of crown markedly
convex, smooth; cutting edges prominent; a single pair of
lateral denticles, broad, but pointed. Anterior teeth narrow
and upright; lateral teeth much inclined backward, the anterior
teeth much more arcuate and longer than the posterior ones.’’ °
Several teeth from the Niobrara chalk agree sufficiently well
with the foregoing description, and especially with Woodward’s
figures, to permit their allocation here. They are somewhat
broader than the specimens figured by Woodward. Two of the
specimens differ markedly from the others in haying the base
flatter and the roots much less prolonged downward, the notch
of the base shallower and shorter. Another tooth from the
base of the Benton, in the conglomerate containing the speci-
mens of Corax curvatus and Ptychodus janewayii, agrees well
6. Woodward, l. c.
248 University Geological Survey of Kansas.
with these last specimens and apparently belongs to the same
species, if distinct. Their resemblance to Odontaspis kopingen-
sis Davis likewise cannot be denied, but the lateral denticles are
more triangular in shape.
Lamna sulcata. Plate XXIV, figs. 1-1b.
Otodus sulcatus Geinitz, Char. Schicht. u. Petriffact, saechs-boehm Krei-
Gelono, leet veehapo
Otodus divaricatus Leidy, Ext. Vert. Fauna, p. 305, vol. xviii, ff. 26-28;
Cope, Cret. Vert., p. 295.
Lamna suleata Woodward, Cat. Foss. Fishes Brit. Mus., 1, p. 398 (where
additional synonymy will be found); Proc. Geol. Assoc., x11, p. 197;
Williston, cf. cit. 37—Cenomanian and Turonian, England, France,
Belgium, Saxony. Bohemia; Senonian, England; Cretaceous, Texas,
(Leidy); Jewell county, Kansas, (Cope); Mississippi (Cope). ;
‘Teeth very robust, the crown sometimes attaining a height
of nearly 50 mm. Outer face of crown slightly convex, gener-
ally uneven; both the inner and the outer faces with more or
less prominent series of vertical wrinkles near the base, usually
irregular. A single pair of large, acuminate lateral denticles,
slightly divergent, often incompletely separated from the prin-
cipal cone. Root with a considerable inward prominence im-
mediately below the base of the crown.’’‘
“A name given to very large, robust teeth with vertically
wrinkled crown and slightly divergent acuminate lateral denti-
cles. There are specimens in the British Museum from un-
determined horizonsin the chalk of Kent, Surrey, and Sussex.’’ *
This species is unknown to me. It occurrence in Kansas is’
given on the authority of Cope. The horizon is evidently the
Benton.
Lamna mudgeli.
Lamna mudgei Cope, Cret. Vert., p. 207, pl. x11, ff. 11, 12; Williston, cf.
cit. 38— Niobrara of Kansas, Greensand of New Jersey.
‘‘ Indicated by three teeth from the Niobrara epoch of Kan-
sas and one from the Greensand No. 4, from New Jersey.
These teeth are rather stout, especially at the base, and the
crown is not very elongate. The root is excessively protuber-
7. Woodward, l. c.
8. Woodward.
WILLISTON. | Cretaceous Fishes. 249
ant, projecting horizontally beyond the convex side, and flat
or truncate below the protuberance. The enamel is entirely
smooth. Length, 14 mm.’’
This species is unknown to me, or unrecognizable from the
description and figures of the mutilated type specimens.
Lamna macrorhiza.
Lamna macrorhiza Cope, Cret. Vert., p. 297, pl. xii, ff. 9,10; Woodward,
Cat. Foss. Fishes, Brit. Mus., 1, p. 399; Williston, cf cit. 38— Niobrara of
Kansas: Albian, of England; Cenonian, of S. E. Russia (Woodward).
‘‘Teeth of small size, elevated though robust, the maximum
total height being about 25 mm. Outer coronal face flat, or
nearly so, with a faint median longitudinal elevation, and
often a few folds at the base; inner coronal face very convex,
smooth ; cutting edges sharp; a single pair of relatively large,
narrow, acuminate lateral denticles, divergent, also often
marked at the base by minute vertical folds; root with a
prominent inward projection below the base of the crown;
nutritive foramen in a groove.’’
The above description by Woodward is drawn from a Euro-
pean specimen, while the type described and figured by Cope
is from Ellis county, Kansas, probably Niobrara. I do not
know the species.
Lamna (Odontaspis?), sp. Plate XXX, fig. 5.
Lamna (Odontaspis?), sp., Williston, cf. cit. 38.
A single tooth from the Lower Cretaceous ( Kiowa shales,
Clark county), resembles the figure of Odontaspis kopingensis
Davis, as figured by that author’ except that it is smaller and
has the base rather more prominent, more triangular, and more
pointed. The tooth has alsoresemblance to Lamna appendiculata,
but the denticles are stouter (compare Woodward.) Height
of crown, 15 mm.; width of base, 18 mm.; width of base of
crown, 9 mm.; distance between points of denticles, 14 mm.
9. Trans. Roy. Dubl. Soc., [V, XXXVI, figs. 27, 2s.
10. Proc. Geol. Assoc., XIII, pl. VI, fig. 26.
lj—vi
250 University Geological Survey of Kansas.
Lamna, sp. Plate XXX, fig. 6.
Lamna, sp., Williston, cf. cit. 39.
A somewhat injured tooth, of larger size than the last, dif-
fers in having a larger and stouter base, the inner projection
in the middle of the latter stouter and broader, and the lateral
denticles smaller and more obtuse. Height of tooth (approxi-
mately), 32 mm.; width of base of crown, 12 mm.; width of
base of tooth, 25 mm. ‘
One specimen, Kiowa shales, Clark county.
Lamna quinquelateralis.
Lamna quinquelateralis Cragin, Colo. Coll. Studies, v, p.189; Williston,
cf. cit. 39.
‘The specific name quinquelateralis is applied to a species of
shark whose vertebre differ from all others of which I have any
knowledge. The type vertebra is short, much broader than
high, shallow-cupped, and more or less sharply pentagonal
ended.
‘‘Measurements: Height, 20mm.; length, 12 mm. ; breadth,
12 mm. The two upper angles measure each about 180 deg. ;
either lateral angles about 105 deg.; the lower angle is broad
and rounded.
‘‘Occurrence: A single vertebra of this form was found by
. the writer at Belvidere, Kan., with the above-described remains
of Plesiochelys, in the upper part of No. 4 of the Belvidere sec-
tion.’’
Possibly this vertebra belongs with one or the other of the
above-described teeth from these same deposits, but the corre-
lation cannot be made until the teeth and vertebre are found
associated, which may be long hence.
Scapanorhynchus.
Rhinognathus Davis, Trans. Roy. Dubl. Soc. (2), 111, p. 480.
Scapanorhynchus Woodward, Cat. Foss. Fishes Brit. Mus., 1, p. 351, (1889).
? Mitsukurina Jordan, Proc. Calif. Acad. Sci. Zool., 1, (1898); Amer. Nat.,
XXXIV, p. 234.
The genus Scapanorhynchus, first proposed by Davis under a
preoccupied name, has been more closely defined by Woodward.
The teeth themselves cannot in many cases be generically dis-
WILLISTON. | Cretaceous Fishes. 251
tinguished from those of Odontaspis, under which name some
were originally described.
Recently Doctor Woodward" has identified a modern genus
of sharks, from the deep sea off Yokahama, Japan, with this
supposedly extinct type —Mitsukurina Jordan.
Possibly the positive identification is premature, but there
seems to be no doubt of the close relationship of the two forms,
at least.
Scapanorhyncus rhaphiodon. Plate XXVI, fig. 4: plate XXXII, fig. 5.
Lamna (Odontaspis) raphiodon Agassiz, Poiss. Foss., 111, p. 296, pl.
xxxvil-a, ff. 12-16.
Scapanorhynchus rhaphiodon Woodward, Cat. Foss. Fishes Brit. Mus.,
I, p. 303 (where additional synonymy will be found); Proc. Geol. Assoc.
x11, p. 196 — Cenomanian, Russia and Galicia; Cenomanian and Turonian,
France, Saxony, and Bohemia; Cenomanian-Senonian, England; Upper
Cretaceous, §. India: Upper Cretaceous of Texas, Mississippi, New
Jersey: Benton Cretaceous of Kansas.
Lamna texana Roemer, Kreideb. von Texas, p. 29, pl. 1, ff. 7: Leidy, Rep.
U.S. Geol. Surv., 1, p. 304, pl. xviti, ff. 46-50; Cope, Cret. Vert., p. 296.
Teeth of considerable size, slender, the anterior ones without
lateral denticles ; inner coronal face conspicuously and finely
striate.
A number of teeth before me from the Cretaceous of New
Jersey and one from the Benton Cretaceous of Kansas agree
fairly well with the figures given by Leidy of specimens from
Mississippi, New Jersey, and ‘‘from near the mouth of Ver-
milion creek, in Kansas,’’ and which agree with those from
Texas called Lamna texana by Roemer.
The specimens agree so well with the European species,
especially as figured by Woodward (l.c.; I have no European
specimens for comparison), that I think there cannot be much
doubt of their identity, a conclusion suggested by Woodward.
The Kansas specimen described by Leidy was said to have
been obtained by Hayden from a ‘‘gray sandstone from near
the mouth of Vermilionriver.’’ The Vermilion in Kansas runs
its whole length through the Carboniferous in eastern Kansas ;
nor do I think there is any gray sandstone (necessarily Dakota
Cretaceous) in the state which will yield these teeth. In all
11. Amer. Mag. Nat. Hist., III, p. 487 (1899).
252 University Geological Survey of Kansas.
probability the specimens did not come from this state. How-
ever, a specimen in our collection agreeing with the species was
obtained in the state, and probably from the Benton, though
possibly from either the Niobrara or Fort Pierre.
CORAX.
The genus Corax is confined wholly to the Cretaceous, and
is yet incompletely known. Its distinction from Galeocerdo,
under which name some of its species have been described, is
based upon the solidity of the teeth— those of Galeocerdo have
a hollow cavity within. The teeth are small, compressed, more
or less triangular, with marginal serrations, though this charac-
ter may be more or less wanting in young individuals. They
vary not a little in shape in the same individual. In some the
crown is nearly bilaterally symmetrical, but they more usually
have the crown directed more or less obliquely backward, the
anterior margin convex, the posterior more or less straight and
angulated.
Three species of the genus are known in England—C. falcatus,
C'. pristodontus, which is hardly distinct, and C. affinis. In ad-
dition, C. antiquus Desl., C. incisus Egert., C. levis Gieb. and
C. pygymxus Munst. have been described from Europe, and C.
crassidens Cope and C. hartvelli Cope from the United States.
Corax falcatus. Plate XXXI, figs. 1-40; plate XXXII, figs. 1-11.
Corax falcatus Agassiz, Poiss. Foss., 111, p. 226, pl. xxvi, f. 14, xxv1-a, ff.
1-15; Woodward, Cat. Foss. Fishes Brit. Mus.,1, p. 424 (where additional
synonymy will befound); Proc. Geol. Assoc., x111, p. 198, pl. vi, ff. 13-15 —
Cenomanian and Turonian of England, France, Switzerland, Saxony,
Bohemia, Galicia, and Russia; Senonian of England and France; Cre-
taceous of Texas, New Jersey, and Mississippi; Niobrara of Kansas.
Galeocerdo falcatus Leidy, Ext. Vert. Fauna West. Terr., p. 301, pl. xv1t,
ff. 29-42.
The very variable shape of the teeth referred to this species
will be seen in plate XX XI, figs. 1-40. Possibly the specimens
there figured represent distinct species. C. (Galeocerdo) crassi-
dens Cope seems to be represented by fig. 24, and C. (Galeocerdo)
hartwelli Cope” by fig. 23. Possibly this species also includes
12. Cret. Vert., p. 244.
WILLISTON. | Cretaceous Fishes. 253
C. pristodontus and C. lindstromi, both of which seem to be im-
perfectly differentiated from C. falcatus at present.
In plate XX XIII, figs. 1-1/, are shown a number of teeth per-
taining to a single individual and found associated with many
others, by Mr. Martin, in the Niobrara Cretaceous of the Smoky
Hill valley. Isolated teeth of this species are the most abun-
dant of the selachian teeth in the Niobrara of Kansas. Only in
very few instances have many teeth been found associated, so
that it is yet impossible to fully understand the dentition. The
species occurs rarely, if at all, in the lower Niobrara horizons,
where those of /Jsurus and Ptychodus are the most abundant.
Corax curvatus, p. sp. Plate XXX, figs. 7, 8.
Two specimens from the same block which yielded those of
Ptychodus janewayit and Lamna species, antea, seem evidently
specifically distinct from the foregoing. These teeth, while not
differing much in outline from certain ones referred to C. falca-
tus, show a marked variance in structure. In C. falcatus the
outer surface of the tooth stands out but very slightly. In C.
curvatus the crown is attached to the root very obliquely, so that
when resting upon a plane the tooth forms a high arch, touch-
ing only by the extreme tips of the roots and crown. The inner
surface, also, is very much more uneven and convex, the crown
separated from the root by a marked, narrow, transverse ridge,
which is scarcely indicated in the teeth of (. falcatus. Altitude,
8 mm.; greatest width, 14 mm.; horizon, lower or lowermost
Benton of Ellsworth county.
LEPTOSTYRAX.
Williston, Kans. Univ. Quart., IX, p. 42, 1900,
Leptostyrax bicuspidatus. Plate XXIV, figs. 15, 15a: plate XXVI, fig. 7.
Leptostyrax bicuspidatus Williston, cf. cit. 42.
Principal cusp long and slender, flattened upon the outer
side, with sharp, smooth edges and a median convexity in the
middle of the flattened surface ; for the most part convex lon-
gitudinally, gently concave before the apex. Inner surface
strongly convex from side to side, concave on the lower half
254 University Geological Survey of Kansas.
longitudinally, gently convex on the upper part. A single
denticle present, slender, flattened cylindrical, with an anterior
and posterior carina; it arises below the base of the main cusp,
and is directed more outwards, its inner surface concave longi-
tudinally. Immediate base of crown of both main cusp and
denticle with short ridges. Base of tooth short, truncate (?)
below the main cusp, prolonged downward below the denticle.
Length of main cusp, 19 mm.; width of same at base, 5 mm. ;
length of denticle, 5 mm.; width of same at base, 2+ mm. ;
height of tooth, 26 mm.; width of base, 10 mm.
A small tooth of the same form found with this has a total
length of 14 mm. The base is deeply emarginate below, with
two slender roots; that below the denticle the longer. Mentor
beds, 4% miles southwest of Marquette, Kan.
U. 5. Nat. Mus. No. 1979.
PYCNODONTIDA.
The pycnodonts are a peculiar group of ganoid fishes, whose
remains have been found in the Jurassic, Cretaceous and EKo-
cene deposits of Europe, North America, Asia, and Australia.
They are all rather small fishes, very much flattened and oval
in shape, covered with rhomboidal scutes having close-lying
spines, which give aribbed appearance. The united palatine
and vomer of the upper jaws are provided with five rows of
round or oval, smooth-pavement teeth; the premaxillary with
two or four chisel-like teeth. The dentaries below have a like
number of teeth, similar to those of the premaxillary, while on
the splenial there are three, four or five rows of pavement teeth
similar to those of the vomer.
Celodus brownii. Plate XXIV, fig. 12.
Celodus brownii Cope, Journ. Acad. Nat. Sci. Phil., rx, p. 447, pl. xx,
f.19; Williston, cf. cit. 28.
A fragment of the left lower jaw, containing two rows of
teeth, the middle and the inner. There are four crowns pre-
served on the inner row, nearly corresponding in length with
WILLIsTON. | Cretaceous Fishes. 255
the six teeth of the middle row, of which only two have the
crowns preserved. Cope’s type had only the middle and ex-
ternal rows, and none of the teeth had well-preserved crowns.
The middle teeth seem to correspond exactly in size with the
types.
On the inner side the jaw projects as a rather broad trough,
with a thin edge, apparently broader posteriorly than anteriorly.
Its width here is nearly as great as the width of the inner row
of teeth. The inner teeth are large, their width equal to nearly
half their length. The surface of the crowns is smooth and
convex, more so antero-posteriorly than transversely. The
middle row has the teeth placed a little obliquely to the others,
and the surface is more flattened transversely in the middle.
The axes of the crowns of the two rows are placed at a distinct
angle with each other.
eeneen OF LOU LeGhhy Inner TOW . . 6. 2a. ew. ee ee te ee es 36 mm.
Transverse diameter of crowns, inner row.................. vay
henpth of five teeth, middie row... 25.6... 2. eee ees ol,
Transverse diameter of crowns, middle row................ 5 he
Thickness of jaw, at middle row of teeth.................. ya
The specimen was collected from the Kiowa shales, near Bel-
videre, by Mr. C. N. Gould.
Celodus stantoni, n. sp. Plate XXIV, fig. 12; plate XXVI, fig. 6.
Celodus stantoni Williston, ef. cit. 28.
A fragment of the right lower jaw, containing two perfect
crowns of the internal row, together with the bases of four
teeth of the middle row, evidently represents a species distinct
from the previous one. The teeth are much smaller in size,
more elongated and distintly kidney-shaped, the ends narrowed.
The surface is smooth, strongly convex antero-posteriorly, and
gently so from side to side. The jaw is much less robust than
in the preceding species.
Transverse diameter of tooth, internal row................ 14 mm.
Antero-posterior diameter of same...................0.006- a 8°
Peenrhiee TOME GOCE, SIGIGNO TOW. . «.. <.ciais sus't int oe sien xe thie i Ge
Transverse diameter of tooth, middle row.................. me Nee
The middle teeth seem to be larger in proportion to those of
the internal series than in the preceding species. Kiowa shales.
256 University Geological Survey of Kansas.
Mesodon abrasus.
Mesodon abrasus Cragin, Colorado College Studies, v, 1894; Williston,
Kans. Univ. Quart., 1x, p. 29.
‘“This name is proposed for certain pycnodont teeth of low,
rhomboidal form and feebiy convex upper surface which occur
in No. 38 of the Belvidere section, southwest of the Belvidere
railroad station, and seem to agree with the large mandibular
teeth of Mesodon. The specific name refers to the occurrence
in the type species of two small, oblique facets produced at one
end by attrition. The type has a height (above root) of 3mm.,
a length of 13 mm., and a breadth of 5 mm.
‘“To the vomerine set of the same species may belong the
rotund, oval or nearly hemispherical teeth of similar height
but smaller size, which occur not uncommonly at the same
locality and horizon, the largest now available example of
which measures about 6 and 7 mm. in major and minor hori-
zontal diameters.”’
In the National Museum collection there are several teeth,
occurring singly, corresponding to the vomerine teeth described
by Cragin. That they belong with the other teeth there de-
scribed is very doubtful — indeed it is doubtful whether the other
teeth belong with J/esodon, since it is impossible to locate the
genus from single teeth. It is not at all impossible that the
vomerine teeth are identical with Cope’s M. diastematicus. The
larger teeth may be the same as those of either the above-
described species of Cwilodus.
The largest of the specimens in the present collection meas-
ures 10 by 73 mm.; several smaller examples have diameters
of 6and5mm. (See plate XXX, fig. 4.)
LEPIDOSTEIDE.
Lepidotus, sp.
In the National Museum collection there is a single example
of a scute pertaining to some lepidotid fish ( No. 1063, Kiowa
shales). Cope has described Macrepistius of this family from a
stratum between the Upper and Lower Trinity Sands of Texas.
It seems very probable that the teeth referred to the vomer of
Mesodon abrasus really belong here.
Pat St
+. a. PF > _
TELEOSTS OF THE UPPER CRETACEOUS.
By ALBAN. STEWART.
INTRODUCTION.
AxBouT two years ago, through the kindness of Dr. 8. W.
Williston, I was given the privilege of working up for publica-
tion the excellent University collection of teleost fishes from the
Upper Cretaceous formations of western Kansas, the results of
which are given in the following pages. At the time the work
was begun the literature on the subject was poorly represented
in the University library, and, as some of it was very hard to
obtain, the work has been delayed on this account, causing
a later appearance of it than was first expected. However, it
is hoped by the author that it is fairly complete, and that it
will be a guide to students of the Cretaceous ichthyology of
North America, and especially that of Kansas.
For more than twenty years past the Kansas Cretaceous fishes
haye been almost entirely neglected by writers on ichthyology,
and not since the publication of Professor Cope’s ‘‘ Vertebrata
of the Cretaceous Formations of the West,’’ in 1875, has any
work appeared beyond a few scattering papers on some special
form or group; and, with the exception of those by Prof. O. P.
Hay and myself, these have all been written by students of Ger-
man universities, from collections which have been made in
Kansas and sent to the museums there. ‘Thus, it seems very
proper that there should be a more complete work undertaken
at home, where the collecting grounds are easy of access and
the collections are probably more complete than elsewhere."
In addition to the original collection made by Chancellor I’.
H. Snow, Dr. S. W. Williston, and others, the geological ex-
13. See also the Appendix to this part.—S. W. W.
( 257 )
258 University Geological Survey of Kansas.
pedition to western Kansas during the summer of 1898 ob-
tained some valuable material from the Niobrara and Fort
Pierre formations, among which several new types have ap-
peared. Before entering into the work in detail, I wish to ex-
press my indebtedness to Dr. S. W. Williston for advice in the
preparation and revision of the work, and also to Prof. O. P. Hay,
of the United States National Museum, of Washington, D. C.,
for the loan of his type specimen of Ichthyodectes cruentus for
comparison with some of our specimens, and also for informa-
tion on some of the points on which I was in doubt. I must
also express my sincere thanks to Mr. W. O. Bourne, of Scott,
and Mr. Travis Morse, of Iola, for the loan of material from
their collections of fossil vertebrates; and also to Mr. H. M.
McDowell, of this city, for his present of the fine skull of
Saurodon xiphirostris. The drawings from which the plates
were made were all done by Mr. Sydney Prentice, under the
writer’s direction.
LAWRENCE, KAN., May 15, 1899.
STEWART. | Cretaceous Fishes. 259
TELEOSTEI.
The fishes belonging to this order are usually characterized
by the margin of the upper jaw being formed of the maxilla
and premaxilla and by the absence of spinous rays in the pelvic
fin. Some of the Cretaceous forms are exceptions to this, as
the pelvic fin often is provided with spinousrays. The parietal
bones are united in the median line and the scales are usually
cycloid. This order comprises the most generalized types of
bony fishes, which are. closely related to the bony ganoids.
They are among the most abundant fossils obtained.from the
chalk of western Kansas, and are usually in an excellent state
of preservation, which makes them easy to collect and study.
They range in size from the gigantic Xiphactinus, which often
attains a length of nearly twenty feet, to Hnchodus and some
of the other small forms, which are often less than a foot in
length. They are found most abundantly in the Niobrara group,
not because they were more abundant in that period, but prob-
ably because the conditions which prevailed at that time were
more favorable for fossilization than in the Fort Pierre and Fox
Hill time which followed. They include several families, which
will be described and discussed below.
A. S. Woodward," of the British Museum, has divided
these fishes into six groups, which are given below.
“IT. Laterally compressed fishes with large and powerful maxille and pre-
maxillz, bearing teeth, the dentary being the only tooth-bearing bone of the lower
jaw, provided with a single series, the palatine and ectopterygoid toothless. The
teeth are placed in complete sockets. Vertebri, except near the head, deeply
two-grooved on each side in addition to possessing pits for insertion of neural and
hemal arches. Ex.: Por/theus(Xiphactinus), Ichthyodectes, Daptinus (Sau-
rodon), Saurocephalus (forming the family of Saurodontida@ of Cope).
‘‘TJ. Fishes somewhat less laterally compressed, provided with scales or bony
scutes, or both, and having the premaxille# and maxille large, bearing powerful
teeth in one or more series. The teeth not implanted in sockets, but anchylosed
14. Proc. Geol. Assoc., vol. X, 1888, pp. 309, 310.
260 University Geological Survey of Kansas.
to the jaw-bones. Abdominal vertebra, at least, without deep lateral pits, but
longitudinally striated. Ex.: Pachyrhizodus, Empo (?), Stratodus.
‘TTT. Fishes only moderately compressed from side to side, naked, or pro-
vided with scutes. Both maxilla and premaxilla long and slender, the former
about half excluded from the margin of the upper jaw by the latter; the maxil-
lary and premaxillary teeth small. Palatines and ectopterygoids powerful, and
bearing a single series of large teeth, upon expanded, anchylosed to the bone.
Dentary bone of lower jaw with one series of large teeth and one or more series
of small teeth similarly anchylosed. Vertebre with two deep lateral impressions
and pits for the neural and hemal arches. Hx.: Hnchodus, Furygnathus,
Eurypholis, Ischyrocephalus, Cimolichthys, Pomagnathus.
“TV. Elongated fishes, with powerful dentition, and the trunk armed with
several longitudinal series of bony scutes; not yet precisely defined and separated
from groups II and III. Ex.: Dercetis (Leptotrachelus), Pelargorhynchus,
Plinthophorus.
“WV. The Protosphyrenide (Hrisichtheidce Cope), with much elongated
snout (ethmoid bone), and long maxilla loosely connected with the premaxilla;
also believed to have an unusually complex mandible. Ex.: Protosphyrenda.
‘“VI, Clupeoids and salmonoids.”’
After making a careful study of the material at hand, I think
it advisable to make some changes in the above grouping for
the American genera, which are given below.
I. Fishes with skull laterally compressed; jaws powerful,
and bearing a single row of cylindric teeth, without nutrient
foramen or notches below the internal alveolar border; preden-
tary not present; vertebrze deeply grooved, with pits above and
below for the insertion of the neuro- and hemapophyses. Hm-
bracing the genera Viphactinus, Icthyodectes, Gillicus, and Clado-
cyclus; family, Icthyodectide.
Il. Fishes with skull laterally compressed; jaws powerful,
and bearing a single row of compressed, knife-like teeth, with
nutrient foramina or notches below the internal alveolar bor-
der; predentary present and without teeth. Including the
genera Saurodon and Saurocephalus; family, Saurodontide.
Ill. Fishes with skull depressed, the top of which is beauti-
fully sculptured ; jaws provided with one or more series of con-
ical teeth, which are firmly anchylosed to the bone; vertebre
STEWART. | Cretaceous Fishes. 261
striated and contracted medially. Including the genera Empo,
Stratodus,” and (?)Cimolichthys; family, Stratodontide.
IV. Fishes with an elongated compressed body; palatines
usually provided with a single large tooth; dentaries with an
internal row of large teeth, and usually an external fringe of
smaller ones. Including the genera HMnchodus, Tetheodus(?) ;
family, Enchodontide.
VY. Fishes with skull depressed, beautifully sculptured, and
probably covered with bony plates. Maxille, premaxille and
dentaries with many rows of minute teeth. Vertebree striated
and contracted but little medially. Including one genus,
Anogmius; family, Osteoglosside.
VI. Fishes with maxille and dentaries provided with a
single row of cylindric subequal teeth, all of which are partially
enclosed in alveoli and partially anchylosed to the bone.
Bones of the skull without ornamentation. Vertebrie longi-
tudinally striated. Including the genera Pachyrhizodus and
Oricardinus; family, Salmonide.
VII. Fishes with the ethmoid bone prolonged into a rostrum
anteriorly. ‘Teeth laterally compressed, lanciform in two series,
the largest of which are set in alveoli. Predental bones prob-
ably present bearing teeth. Including one genus, Protosphyrexna ;
family, Pachycormide."
VIIl. Body covered with large cycloid scales. Abdomen
frequently compressed intoaserrated edge. Dorsal fin elongate ;
caudal fin often deeply cleft. Including the American genera
Leptichthys, Sardinius, and Spaniodon; family, Clupeida.
IX. Body slender and covered with several rows of bony
scutes. Teeth in a single series. Including the American
genera Dercetis (Leptotrachelus), Triwnaspis, Ichthyotringa, and
Leptecodon ; family, Dercetidez.
_ 15. Dr. A. S. Woodward, of the British Museum, has since examined our specimen, and
thinks the large size of the parietals, their meeting medially and the general shape of the top
of the skull would indicate that Siratodus was closely related to Dercetis.
16. Vertebrate Paleontology (A. 8. Woodward), p. 111.
262 University Geological Survey of Kansas.
X. Body short and covered with ctenoid scales. Mouth
oblique. Jaws with many villiform teeth. Embracing the one
genus, Beryx; family, Berycidx. |
XI. Body somewhate elongate, compressed, and covered with
large or medium-sized cycloid scales. Lateral line doubtfully
present. Dental bones short and without teeth. Eye lateral.
Gill openings large. Dorsal fin often elongated. Embracing
the genera Sylemus, (?) Apsopelix, and (?) Pelecorapis; family,
Mugihde.
ICHTHYODECTIDZ Crook.
Saurodontide Cope; Saurocephalide Zittel.
The family name of Ichthyodectidx as characterized by Doctor
Crook” embraced the genera Xiphactinus, Ichthyodectes, and
Sauwrodon, and was intended by the author to be used instead of
the family name Saurodontidx, as given to this group of fishes
by Professor Cope.” The reasons given by Doctor Crook for
changing the name of this family are that the teeth are not like
those of saurians, and that the name Saurodontide was pre-
occupied by a family of ganoid fishes. Concerning this Pro-
fessor Cope says:' ‘In the first place, the author [Crook] has
not observed that I have on several occasions published the
fact that the name Daptinus Cope is a synonym of Saurodon
Lea,” which was proposed many years previously. It was from
this genus that I gave the family name first proposed of Sauro-
dontide. The fact that Professor Zittel.many years later gave
this name to a very distinct family does not authorize the giving
of a new name to the family first called by me, as is done by
Mr. Crook.’’ Doctor Crook also removes the genus Sauro-
cephalus from this group, and places it in the family Proto-
sphyrenidx (Pachycormidx), concerning which he says of the
maxilla, premaxilla, and dentary which he examined :™
“Sie ist so ginzlich verschieden von denjenigen der anderen Glieder
dieser Familie, dass dies Merkmal allein geniigt, um Sawrocephalus
17. Paleontographica, 1892, p. 120. 20. Should be Hays.
18. Proc. Am. Phil. Soc. 1870, p. 529. 21. 1. c., 120,
19. American Naturalist, vol. XXVI, p. 941.
eo.
STEWART. | Cretaceous Fishes. 263
einer anderen Gruppe zuzuweisen. Diese Annahme wird noch mehr
bestitigt durch den Charakter der Maxilla und Dentale, die Foramina
und die Art und Weise der Aufeinanderfolge der Zihne. Auf Grund
der Gleichartigkeit der Zihne und der aus erordentlichen Aehnlichkeit
der Preemaxilla mit derjenigen von Protosphyrewnan diirfen wir
Saurocephalus bis auf weiters in die Familie von Protosphyrana
einreihen.”
I have been unable to discover the similarity between the
premaxillie of the two genera mentioned above. In fact, they
are very widely different, and I am inclined to think that Doctor
Crook mistook the predentary for a premaxilla of Saurocephalus.
However, this genus is too closely related to Sawrodon to be far
separated from it, and if one of these gencra is removed from
this family the other would have to follow.
As it is evident that these genera cannot be placed in the
Pachycormidx, and as some new characters have come to light
since Doctor Crook’s paper appeared, I deem it advisable to
separate these two forms from the original group and place
them in a separate family, to be known as the Saurodontide,
and to use the name Ichthyodectidx to include the three genera
NXiphactinus, Ichthyodectes, and Gillicus.
Concerning the same Saurocephalidx Zittel, it can only be
said that even if the name Sauwrodontidx had been used by Doc-
tor Zittel to distinguish his family of ganoid fishes before Pro-
fessor Cope applied it to his group, it would have to be
abandoned in light of the fact that Cope’s family is founded
on the genus Saurodon, which would give it priority over another
family of the same name.
In this family the tooth-bearing elements are each provided
with a single row of teeth, and the upper border of the mouth
is formed by the maxilla and premaxilla. The supraoccipital
is raised into a prominent crest and the maxilla is bound to the
skull by means of the palatine.
This family embraced some of the largest physostomous fishes
of the Cretaceous period of North America, and from the size of
the jaws and the powerful dentition we may suppose that they
rivaled the Mosasaurs, the smaller ones at least, in strength
264 University Geological Survey of Kansas.
and ferocity. Below is the synopsis of the family Sawrodontide,
as given by Cope:
I. Jaws without foramina on the inner face below the alveolar margin:
a. Teeth cylindric:
Teeth of unequal lengths; some of them greatly de-
VOLO POG cua. el ale a ee Ao ee eee pe ee Re Portheus.
Teeth of equal length......... eR se eens A as Ichthyodectes.
aa. Teeth compressed, knife-like:
Teeth of unequal length; some of the anterior
preatly developed! 1 Jaa. kas oe ee eee Hrisicthe.
MPeeth veq wali ih Nee en he nen ape ean an et Daptinus.
II. Dentary bones pierced by foramina below the alveolar border:
Teeth with!subeylindriclerownsse see nie eee Saurodon.
Teeth with short, compressed crowns ................. Saurocephalus.
Professor Cope also says: ‘‘ There are some other forms to be
referred to this family, whose characters are not yet fully de-
termined. Thus Hypsodon Agass., from the European chalk, is
related to the two genera first above named, but, as left by its
author in the ‘ Poissons. Fossiles,’ includes apparently two
generic forms. The first figured and described has the man-
dibular teeth of equal length. In the second, they are unequal,
as in Portheus, to which genus this specimen ought, perhaps,
to be referred. Retaining the name fHypsodon for the genus
with equal mandibular teeth, its relations to [chthyodectes re-
main to be determined by further study of H. lewesiensis. The
view of the superior walls of the cranium given by Professor
Agassiz presents characters quite distinct from what I have ob-
served in Portheus.’’ Hypsodon has since been shown to be a
synonym of Pachyrhizodus, Portheus a synonym of Xiphactinus,
Hrisicthe a synonym of Protosphyrxna, and Daptinus a synonym
of Saurodon. Thus, itis seen that the old synopsis will no longer
hold, so I give the following revised one :
a. Teeth large and of unequal length...............2 Niphactinus.
6. Teeth medium, of equal lengthy 0 ne eoce Ichthyodectes.
e. Reeth small and iiringe-hkes. tee eee Gillicus.
The two large oval scales from the chalk of Lewes, figured by
Agassiz” as Cladocyclus, bear a close resemblance to those of
22, Poiss. Foss., vol. V, pl. XXVa, figs. 5, 6.
Led kd
Aa
STEWART. | Cretaceous Fishes. 265
Ichthyodectes, and it may later be found that the two genera are
identical. However, as Cladocyclus is so imperfectly known, it
would hardly be safe to say that the two genera are synonymous.
Xiphactinus.
Niphactinus Leidy, Proc. Acad. Nat. Sci. Phil. 1870, p.12; Rep. U.S.
Geol. Surv. Terr. 1873, p. 290, pl. xvir.
Portheus Cope, Proc. Am. Phil. Soc. 1871, p. 175: Cret. Vert. West, pp.
190-193.
The genus \iphactinus was first described by Leidy* from the
remains of a pectoral fin-spine from the Cretaceous deposits of
Kansas. Three years later the spine was more completely de-
scribed and figured by the same author in his Contributions to
the Extinct Fauna of the Western Territories, ( p. 290, pl. XVII.)
Professor Cope recognized the relationship of this spine to his
family Saurodontidae, in a paper published in Hayden’s Second
Annual Report of the Geological Survey of the Territories, 1871,
(p. 418,) but assigned it to the genus Saurocephalus, and always
claimed that Viphactinus was distinct from Portheus and Ichthyo-
dectes, although he never stated in what way they differed.
It has since been shown, without doubt, that Portheus is a syno-
nym of Xiphactinus, the latter name having the priority by
about a year.
This genus is represented in the museum by two specimens
which are more than twelve feet in length each, and these are
only medium-sized individuals; so there can be but little doubt
that the species attained a length of twenty or more feet in the
largestspecimens. Besides their large size, the genus is well char-
acterized by the tooth-bearing elements, which are all covered with
a powerful dentition. The premaxille are firmly united with
the maxillz and usually have one or more large, fang-like teeth
projecting downward from the alveolar border. The maxille
are large and provided with a single row of teeth which vary
greatly in size. The upper anterior portion of the bone is pro-
vided with two condyles which serve to bind the jaw to the
skull. The dentaries are deep and also provided with a single
23. Proc. Acad. Nat. Sci. Phil. 1870, p. 12.
18—vi
266 University Geological Survey of Kansas.
row of teeth, very irregular in size. The larger of these teeth
are very deeply set in the jaw and Professor Hay has recently
shown™ that those near the symphysis descend nearly to the
lower border. The pulp cavities are very large at the base and
the displacement takes place by the young crown rising within
the pulp cavity of the functional tooth and the absorption of
the old root.
In the early work upon this genus, a great deal of stress
was placed upon the number, size and arrangement of the teeth
in the determination of the species, which characters have since
been shown to be very inconstant.” The reason for the great
inconstancy is easily explained when we consider the fact that the
old teeth are being constantly shed and new ones taking their
places, thus having fully developed and young teeth on the same
jaw. The palatines and pterygoid bones are probably toothless ;
at least there are no teeth of any size upon them. The palatine
is peculiar in having a malleolar portion connecting the max-
illa with the skull. It is connected with the quadrate by means
of the pterygoid bones, thus forming a strong arch.
The skull is provided with a prominent supraoccipital crest
posteriorly. The orbital cavity is large and the orbit is sur-
rounded by a heavy sclerotic ring, and also a chain of supra-
and suborbital bones. The pterotics form prominent posterior
lateral angles of the skull and also furnish the principal sup-
port for the hyomandibular. The parasphenoid forms a strong
lower axis of the skull and has prominent transverse processes
just in front of the brain case.
The opercular bones are large and thin, and probably all
present. The pectoral fin is large and its rays powerful, prob-
ably forming weapons of offense and defense. ‘The pelvic fins
are supported by the pelvic actinosts, which are heavy, compact
bones, strongly united in the median line by means of suture.
The vertebre are mostly two-grooved. The known American
species are:
Xiphactinus audax Cope, Niobrara Cretaceous, western Kansas.
Xiphactinus lestrio Cope, Niobrara Cretaceous, western Kansas.
24. Zool. Bull., vol. II, p. 36.
25. Kans. Univ. Quart., vol. VII, pp. 115-119, pl. VII-X.
STEWART. | Cretaceous Fishes. 267
Niphactinus mudgei Cope, Niobrara Cretaceous, western Kansas.
Niphactinus lowiti Stewart, Fort Benton Cretaceous, central Kansas.
NXiphactinus brachygnathus Stewart, Niobrara Cretaceous, central Kansas.
In a paper recently published by Professor Hay, of the United
States National Museum,” the author recognizes two more spe-
cies, VY. molossus and X. thawmas, not mentioned in the list given
above, and says that it is quite probable that \. audaz is the
same as some one of Professor Cope’s species of Portheus. He
also recognized the great variation in the size and arrangement
of the teeth, but thought that the two species could be recog-
nized by the difference in form of the superior condyles of the
maxillz, the variation of which will be discussed later.
Now it seems to me that, if Y. molossus and .Y. thaumas are
distinct species, Professor Cope failed to designate any constant
characters by which they can be determined, and, until it is
shown that such characters exist, we can do nothing more than
call them synonymous. I have carefully compared the descrip-
tion and figure of Doctor Leidy’s specimen with specimens of
fin-spines in our collection that undoubtedly belong to one of
these species, and find them identical with the description.
This being the case, we must, for the present at least, consider
X. molossus and X. thaumas Cope as synonyms of Xiphactinus
audax Leidy.
Xiphactinus audax. Plates XXXIII to XLVII.
Saurocephalus thaumas Cope, Proc, Am. Phil. Soc. 1870, p. 533.
Saurocephalus audax Cope, Proc. Am. Phil. Soc. 1870, p. 533.
Portheus molossus Cope, Proc. Am. Phil. Soe. she p- 174; Cret. Vert.
West., pp. 194-196.
Portheus thaumas Cope, Cret. Vert. West., pp. 196- 201.
Xiphactinus audax Leidy, Proc. Acad. Nat. Sci. Phil. 1870, p. 12; Rep.
U. S. Geol. Surv. Terr. 1873, p. 290, pl. xvi.
This species is by far the best represented of the physostomous
fishes in the University collection, the material consisting of
several individuals, some of which are almost complete and
others remarkably well preserved in certain parts, so that from
this material a better idea of the anatomy can be gained than
from any of the other collections that have been described.
yp. a a Pe «7 Pe
268 University Geological Survey of Kansas.
The premaxilla is large and oval in outline, convex externally
and very irregular internally. The bone thickens toward the
lower border, where there are alveoli for the large teeth, which
are never more than three in number, and even three is an ex-
ception. The teeth are acutely pointed at the apex, non-striated,
and directed slightly forward. Considerably more than half of
the bone is supported on the anterior lamina of the maxilla,
and in the larger specimens the posterior union with this bone
is usually by a dentate suture. In the smaller specimens the
suture is usually an undulating line, which seems to indicate
that the dentate character of the suture becomes more evident
with the age of the animal. There was no sutural union with
its fellow on the opposite side, but, from the tubercular nature
of the anterior border in some of the specimens, it seems that
there may have been a cartilaginous or ligamentous union be-
tween the two. In some of the specimens there are a number
of tubercular exostoses just above the alveolar border, but this
character is as inconstant as the form of the suture mentioned
above.
The maxilla is a large lanciform bone, much thickened to-
ward the anterior end and thin posteriorly. Just back of the
premaxilla the alveolar border is rather thin, and contains
alveoli for several small teeth, usually four. Back of this the
bone swells considerably for the accommodation of several large
teeth, which are acutely pointed, non-striate, and vary in cross-
section from a cylinder to an oval. Back of this the border is
occupied by a varying number of small and medium-sized teeth,
the alveoli of which are probably not all filled with functional
teeth at once. The bases of all of these teeth are cylindric, or
nearly so.
On the next page is given a table showing the great variation
in the size and arrangement of the teeth on both the maxilla
and premaxilla. The numbers in the left-hand column are
the catalogue numbers of the specimens.
eee.
STEWART. | (retaceous Fishes. 269
MAXILLARY. PREMAXILLARY.
No. |—— —_——S_
Small. Medium. Large. | Small. | Total. | Small. | Large. | Small. | Total.
1 IG eae 5 | ae fo] Nae BA a dete 2
2 Be es ge ss 4 5 | 22 (Sey Mites wos Bae a liste avai ore 2
3 eee ee 7 ey | eR) live 7s a W Aos ted Ab hs 2
4 DS eee 5 27 ag Eee 7) gf (ae PEN 2
_ ta, ae os 4 5 27 at Ss My Sieh, 2
132 ey, Sec are eae 5 20 2 Og Eee ee oe On ae (ene ee
ey 1 Ayal RROS i ee 2 2 4
179 PS hye ee oe 4 26 3 eee RR? SIM Pa, 2
266 ey hee 6 191) 974-| 1 ee ee 3
275 ole OO ae 5 27 by ue aS eee eeu ed a a
279 EO arene een ae 4 18 Ls Cds 2h Sara elt 2 Yar i a
287 3 a 26 Cote Ol eee lel Oe De at Aer (eee
301 Pe cee ek a 15 EAN Re et retort eked PRs entaslite vias os
353 co OR (5 eae ag Oe 4 22? A a bs ee vida hy, ee 2
From the above table, it is seen that in not a single instance
do the size and arrangement of the teeth exactly agree with
either of Cope’s species, Y. molossus or Y. thaumas. The num-
ber of teeth is also as varied as is the size and arrangement.
Two more or less constant characters are observed in the above
table, viz.: The four smali teeth on the anterior portion of the
maxilla and the two large teeth on the premaxilla which are the
least variable. |
From the above, it is seen that the size and arrangement of
the teeth on the maxilla cannot be taken as specific in char-
acter. The teeth on the premaxilla show a diversity in size
and arrangement which is quite as marked as the size and
arrangement of those on the maxilla. While the two large
teeth are fairly constant, and in some specimens are about
equal in size, yet in other specimens the anterior is much the
larger of the two, while in still others the opposite is the case.
The outer surface of the maxilla is somewhat rugosely
marked just above the alveolar border, while the internal sur-
face is nearly smooth at this point. This surface curves regu-
larly into the superior with no sharp break between the two,
except near the posterior extremity, where the bone becomes
very thin, but the external surface ends abruptly above and
forms a slight shelf for the attachment of probably a jugal bone.
270 University Geological Survey of Kansas.
In some specimens this shelf is somewhat groove-like, while in
others it is broad and flat. Near the anterior end superiorly
there are two condyles, of which the posterior is large, irregular
in outline, and slightly convex. Professor Hay is inclined to
think that two forms can be determined from these condyles, as
he says :* ‘‘I believe that the two species may be identified from
the condyles of the maxillary. Atleast, these condyles are quite
different in the two species which I have been able to examine,
X.thaumas and X. molossus. Fig. 2 represents the maxillary of
the former species, fig. 3 that of XY. molossus. From these fig-
ures, it will be seen that in X. thawmas the posterior condyle is
notched behind, while that of P. molossus |should be X. molossus |
is excavated in front. It appears, too, that the condyle is more
extended longitudinally in X. thawmas, more transversely in X.
molossus.’’ Jam inclined to think that Professor Hay laid too
much stress on this and the anterior condyle in the determina-
tion of his species, as a glance at the series of drawings of these
parts on plate XLI will show that there was quite as much va-
riation in these condyles as there is in the teeth, as shown
above. J cannot be so sure about the variation of the anterior
condyle, as it is broken away in many of the specimens. How-
ever, the constancy or inconstancy of this one minor character
is of little specific value.
The anterior condyle, mentioned above, is oval in outline,
convex, and occupies the extreme anterior part of the laminar
portion of the maxilla, being thus well separated from the pos-
terior condyle. The two condyles are more in line with each
other antero-posteriorly than in Ichthyodectes. Just in front
of the posterior of these condyles, the thickness of the maxilla
becomes suddenly less and forms an abutment, against which
the posterior portion of the premaxilla rests. In front of this
the bone is laminar and supports more than one-half of the
premaxillary bone. The posterior extremity is very thin, and
in some specimens it is nearly straight, while in others it is
curved strongly upward.
27. Zool. Bull,, vol. II, No. 1, p. 34.
A>
STEWART. | Cretaceous Fishes. O71
Maxilla: Distance from anterior condyle to posterior extremity, 338 mm.
Depth of bone at posterior condyle .................. 92
Distance between the two anterior condyles.......... 44°
DER EIVOGIET WOKGOE 6 655. sen ace eee ete 245, *
Thickness at posterior condyle....................... ee
Peemelin. EOnal Ge IG WER DOTUOT, «o.oo. od owe 5 ee ie was Gi
ae ctl nie Fe ly at A Bese 9 Sk en ie aS Silty
A table of measurements showing the great variation in size
of the maxille in different individuals is given:
MAXILL.
Length | Depth at Length Depth at
No. from pre- | posterior No. from pre- | posterior
maxillary.| condyle. maxillary.| condyle.
Reece 102 179 183 68
2 | 9604+'| 111 Cae ie 84
a ge? Od Rae 975 | 9901-1 94
4 | 216.5] 78 279 | 263 90
SS 1 378.5 | Sa | Ane +6 ¥ f 220 80
1 in AR HE oe ep 88.5 SOE fiw al P teal 90
132 264 | 98 353 219+ 75
155 205+ 83.5 |
The dentary is deep, strongly built, and not incurved ante-
riorly. The alveolar border is slightly concave, and bears two
large teeth at the anterior extremity, which are usually sepa-
rated by a transverse groove from those following. Just back of
this groove there are three or four large teeth which are followed
by a number of medium-sized and small ones, the diversity in
size and arrangement of which is shown in the table following
this description. The teeth are all strongly implanted, with
straight conical crowns, non-striate except under the microscope,
and are cylindric in cross-section. The symphyseal portion
slopes strongly backward and is very thick, but the principal
articulation with the opposite jaw seems to be on the internal
side, where the bone is very rugose.. The lower border is rather
sharp, and usually has a row of large nutrient foramina leading
inward and forward just aboveit. The internal side is broadly
vertically concave in the posterior portion, but becomes much
less so anteriorly. At the end of the alveolar border the bone
is projected upward and slightly outward into a weak coronoid
process.
272 University Geological Survey of Kansas.
MANDIBLES.
vo (Ele | ee ee ee
= 03 2 a 2 e Se eee tes = =
= © = © = ® = iaal= Ss = =
a RAPS alia
Me ile Was PR RI ial Da sees Maa we Oya eerae 9 1 20
Be ll eyaenne® 2 i A invariant oe A 8 I> Wieiaiges eal eeteueaene 24
Banik we Seal een Sal ree a eau Brel i a 19-4
2 ON niet terse 7h eA 2 al ans ees a alla tan Macc 10 2 22
ood Meta tes 7 OR aie | ea rs |S 8 Dy el eae 10 4 22
LOT aes das OREO SD eek Uayetey learns Oa eee 10. Sh eee 21
TSS voce ey aN eae SS dla pect eal ee arene Wipe Ayn: 9 IL 22
LOO legoeee Peet | ee iL 2 ope arene cee BL etna ee 22 lees
DAs ae ete 2 1 AL pais, ea aes 8 TD sgh. Ae need 23
Zid |e 2 1 Do te ia etl Ra atl cle eae ele ale eh eel a eee et ee
Pale ce a dese Difal|. eat e a et Sane AS Om apUM mal Lee so oy
Aoi Wah Deter ine Pa Ws cs 3 HO ail ata AA) VOR clit 9 3 22
ale aie QBN Cedve eas Bet weep hal WU ih) Bittle wre cece 8 2 24
353 2 ene aM aan Pale ERASE Ne ehc toe nas te IY fe ete dialect 21
The articular portion as described by Hay* is composed of the
derm and antarticulars, but I am inclined to think that this
author was somewhat mistaken in the extent of the last of these
elements, as he says in the description of this part: ‘‘ Relying
on two good specimens of Xizphactinus and one of Ichthyodectes,
I am confident that the proximal end of the antarticular is con-
tinuous with the long, sword-shaped process described by Cope,
and that this process is entirely distinct from the dermarticu-
lar.’’? Professor Cope states” that the articular is distinct, short,
and irregularly wedge-shaped, and supports half of the cotylus.
In this Professor Cope was correct, as the antarticular is not con-
tinued forward in along, sword-shaped process, but is separated
from this portion by a suture, which is indistinct in many of
the specimens, and it is probably owing to this fact that Hay
was unable to find it.
The dermarticular is a large bone, and in most of the speci-
mens it supports less than one-half of the cotylus. On the ex-
ternal side it is exposed for some distance beyond the cotylus,
when it is covered by the dentary, and continues forward on
the internal side, as a long sword-shaped process, for nearly —
two-thirds the length of the jaw. The cotyloid process is promi-
28, Zool. Bull., vol. II, No. 1, p. 37.
29, Cret. Vert. West, p. 194.
STEWART. | Cretaceous Fishes. 273
nent and curved upward on the external side of the cotylus.
In addition to the above bones, there is another, which is not
mentioned by either Cope or Hay. It is asmall bone, which
lies in front of the antarticular, and is marked sp. in the figure
of the lower jaw on plate XXXV. It presents a small facet
above, which seems to have had attachment with something.
This bone occupies about the same position as the splenial in
some fishes, which name | will give it provisionally.
ETE Gs MAVEOINT DOLGED. iti i le. calc ce slsn nn ba Wed cee 260 mm.
Length of dentary from cotyloid cavity................... aan °F
LAS SEN 2107 I SE Ce a 94 *
Mee RE PE ULE, WCORCAUIAONIED UPIIGL 1s as as wa Gia siete sid a buchclp 6 xin slave was L3G: ..4
Length of articular from cotyloid cavity.................. 224 *
The above measurements are of a single specimen, No. 88 in
the following table, which shows the great variation in the size
of the mandible of this species :
| Length of Length of
No. | alveolar | coronoid. || No. | alveolar | coronoid.
1 242.5 145.5 179 185 94
2 251 144 PUT e dk eto te oee't bass
3 279 140 Tes eel cele oe
4 | 194 109 52 0S Rd eee
88 260 136 287 Gin Ging ot om,
127 beets 321 0 i Reet aa a
; 2a ER gat i Mave | Wee P| 242 130
135 10 tes he. 353 195 115
155 leg Be eee
The hyoid arch is represented by only one bone, the cerato-
hyal, and is the same bone figured by Cope” and described by
Crook” as an interopercular. The bone is elongated, and in out-
line is somewhat the shape of a parallelogram with one side
crushed in. The posterior end is the broader of the two and
bears an elongated, narrow and concave facet for the articula-
tion with the epihyal. The anterior end is more irregular in
outline and bears two facets for the hypo- and urohyals. The
bone is thin and finely striated throughout.
30. Cret. Vert. West., pl. XL, fig. 7.
31. Paleontographica, 1892, p. 117.
274 University Geological Survey of Kansas.
Ceratohyal:. Greatest Jemestly os noi) Ga) eels poten lan hen era 238 mm.
Width across) anterior endian... 2. ya. ee Bou ne
Width across posterior end..................06- Oe
The palato-quadrate arch is made up of six bones —the pala-
tine, pterygoid, metapterygoid,. mesopterygoid, symplectic, and
quadrate. The first and last of these mentioned form the ex-
tremities of the arch, the other four intervening. The palatine
presents a prominent malleolar portion, which has an articular
face above and below for the prefronto-palatine articulation for
binding the maxilla to the cranium, so characteristic of the
Saurodontide and Ichthyodectide. The lower of these faces is
the larger and is slightly concave, while the upper is somewhat
convex. The external surface presents no markings other than
a prominent tuberosity on the posterior side. Just back and
above this portion there are two prominent tubercles of bone,
one of which may give attachment for a nasal bone.
The pterygoid and mesopterygoid are joined to the palatine
posteriorly. The first of “hese is a long, thin bar of bone}y
which extends backward and downward nearly to the condyle
of the quadrate and forms most of the lower border of the arch.
The mesopterygoid is a somewhat triangular-shaped bone oc-
curring just above the pterygoid. It is evidently united to the
metapterygoid by means of a very broad squamose suture, as
the bone appears to be rather small when seen externally, while
the internal side, as figured by Professor Hay*® seems to be
nearly as large as the metapterygoid. I have been unable to
discover any teeth on the external side of this and the ptery-
goid, and none on the internal side of the pterygoid, as far as
can be examined, which seems to be beyond the point where
Professor Hay has shown them to be in his specimen. A sec-
ond specimen shows that in this region there was a great deal
of ossified cartilage covered with denticles, some of which is
adhering to the matrix on the internal side of the mesoptery-
goid, and I am inclined to think that it was some of this carti-
lage that Professor Hay mistook for teeth on the bone. The
32, Zool. Bull., vol. II, p. 39.
—
le |
ST EE ae eee a el ee ee
LS SCC Cle
|
STEWART. | Cretaceous Fishes. 275
metapterygoid is a broad, flat plate just above the quadrate,
and is covered with fine strizw on the lower portion.
The symplectic is a rather small element, occupying a groove
on the inner side of the quadrate. On its superior end it pre-
sents a small articular facet.
The quadrate is very thin above where it joins the metaptery-
goid, but in passing downward to the condyle it becomes more
robust and is contracted into a neck. ‘The condyle is somewhat
elliptical in outline, the ellipse being invaded by a deep notch
on the external side, and is strongly convex from before back-
ward. On the internal side there is a deep groove for the sym-
plectic, and just back of this groove, posteriorly, there is another
smaller groove into which the anterior border of the preoper-
culum fits. The anterior border articulates with the pterygoid
by means of a squamose suture. The bone is minutely striated,
both externally and internally.
aes RETIRE Uh ANON FE 6 So iim, Pater eh sinh xm ainda Veda ens es O42 MM.
PeREIUr MPistt OF HIMMOEOIAT DODGY. <.0 2 Yc odiecaleesinf slows ca suns 4) <,<*
Between malleolar body and condyle for the quadrate..... 261 ***
Antero-posterior length of palatine....................... PAS a jee
PePeNe teeth GME MECH? ks cn ees ecw de sie cis ches iol. =
The opercular bones are represented by the operculum, pre-
operculum, and asupposed suboperculum. There is still another
bone (plate XLIII, fig. 3), which may represent an interoper-
culum.
The operculum is a broad, flat plate of bone, near the anterior
superior portion of which there is a deep elliptical pit for articu-
lating the bone with the hyomandibular. This pit is projected
slightly outward from the rest of the bone, very closely resembling
the corresponding portion of Savrodon in this respect. Just above
this pit there is a sharp angular projection at the point where
the anterior and superior borders meet, but I am inclined to
think that this character will not be constant in all specimens.
From this point the superior border extends backward and down-
ward in an irregularly curved line to the lower border, which
is less curved and somewhat striated just above it. The ante-
rior border, extending from the articular portion just mentioned
downward, is nearly straight and somewhat thickened. Back
276 University Geological Survey of Kansas.
of this border the bone is very thin, thus making it difficult to
collect in perfect condition.
The preoperculum is a somewhat triangular-shaped bone with
a very thick and concave anterior border. The lower portion
of this border is directed forward at quite an-angle, and is re-
ceived in a slight groove on the back portion of the quadrate.
Extending backward from the extremity of this portion there is
a row of large, shallow foramina, the posterior ones of which .
have shallow grooves leading into them from behind. Below
this line of foramina the bone is not so thick, and is covered
with numerous fine strie. The superior portion has a rather
broad process extending upward, formed by the anterior and
superior borders. The posterior portion of the bone is thin and
striate on both sides of the upper portion.
There is still another opercular bone, which I regard as a
suboperculum. The bone is broad and flat, thin along the bor-
ders, and thickened toward the central portion. The posterior
extremity is the broader of the two and is somewhat rounded.
The anterior end is much more narrow and bears a small ellip-
tical facet on a slight elevation of the bone. It also presents a
beak-like process, separated from the above by a slight depres-
sion. Both the external and internal sides are striated, the
striz becoming very pronounced near the posterior end of one
of the sides. Measurements of the opercular bones are:
Operculum: Length of the anterior border.................. * 373 mm.
Length of the anterior border below facet for the
Hyomandibwlarece. ooh eects eee kee eee JOSnis
Length of facet for the hyomandibular......... 5a
Transverse width of facet). 2 seuss see eee DD, ves
Preoperculum: Greatest length............. Bd oe aa sein ea oe
Width near superior extremity .............. SOneSE
Width? ofanferion end eee ee LOS:
Subopercular:Wotalileme tine neta Soa 290)
Width across anterior extremity............. oO eae
Width across posterior extremity............ * 146
There are four other bones, some of which are fragmentary,
which are figured on plate XLIV. They were all found in con-
nection with the skull and evidently belong to it. The first of
these bones, figs. 1 and 5, is an elongated bone, expanded at
* Estimated,
ae
land
STEWART. | Cretaceous Fishes. 277
one end and contracted into an elongated, narrow and some-
what thickened process at the other, at the extremity of which
there is a roughened surface which probably gives attachment
for cartilage. On one side of this process there is a long, thin
lamina of bone extending toward the narrow extremity, and so
closely applied to it that at first sight it has the appearance of
being the border of a groove on the edge of the process. The
expanded end is thickened and bears a small articular surface,
the face of which is almost in line with the process mentioned
above. This bone was figured by Cope” as a ‘‘? hyomandibu-
lar,’’ which has since been shown to be incorrect.
The second of these, fig. 2, is represented by portions of two
bones, neither of which is complete. They are thin on one bor-
der, while the other is much thicker and is invaded by ashallow
groove. This thickened portion was probably continued out-
ward in a process beyond the end of the bone. The third, fig. 3,
is plate-like and more or less sculptured on both sides. On one
of the sides there is a broad, triangular-shaped depression, which
is invaded by a notch from the edge of the bone, and on the
other side there is a prominent ridge extending away from the
apex. I have found a bone almost identical with this in con-
nection with the supposed operculum of an unknown fish, which
will be discussed later. This was figured by Cope as an “‘ un-
certain bone.’’*
The fourth and last of these bones (fig. 4) I have described
as probably an infraopercular,” but it is probably a fragmentary
coracoid. It is the largest of the four under consideration, and
is very thin and flat, excepting near the posterior end, where it
suddenly thickens into a prominent ridge, which probably con-
tinues to the point where the bone articulates with the scapula.
The hyomandibular is a large and somewhat triangular-
shaped bone, with the base of the triangle above, where there is
a long, narrow articular face for articulating this bone with the
pit on the side of the skull forits reception. The face is convex
33. Cret. Vert. West., pl. XL.
34. l.c., pl. XL.
35. Kans, Univ. Quart,, vol. VIII, p. 21, pl. XI, fig. 4. —
278 University Geological Survey of Kansas.
transversely and slightly concave in the middle. Extending
downward .and backward from near the anterior end of this
facet there is a ridge, which becomes very prominent just in
front of the condyle, and, continuing downward, forms a deep
groove with the posterior border, into which the anterior border
of the preoperculum is received to some extent. The anterior
portion of the bone is thin and continued forward quite a dis-
tance. The condyle for the operculum is long, narrow, and
situated about 45 mm. below the superior condyle. Just in
front of this condyle, externally, there is a large and deep pit,
which has another smaller one just above it. On the internal
side there is a prominent crest, which ends opposite the lower
end of the condyle just mentioned, and has another prominent
excavation in front of it. The lower end of the bone is some-
what bluntly pointed, but it does not seem to possess a facet at
this point, as is found in Saurodon.
Hyomandibular: “Geno thy eye aoa es cease ree eet anaes 265 mm.
Length of superior condyle)..:. .... 22. ...%. Sis
Length of condyle for operculum....,...... Oy ee
Greatest width of bone (estimated) ........ 1607 Ss
The ethmoid is much thickened and narrow in front, where
it forms a small beak, and broad and thin behind, where it
joins the frontals by a somewhat dentate suture. The bone is
moderately deep, and on the median line below there is a well-
marked median ridge, on either side of which there is a small
facet for the anterior condyle of the maxilla. In addition to
this there is also a small pit on each side, which probably
accommodates a part of the superior edge of the premaxilla.
The upper anterior portion is often covered with numerous
tubercles of bone, which fade out posteriorly.
The frontals are broad, flat plates, and are the largest bones
entering into the formation of the top of the skull. Just back
of the ethmoid they join the prefrontals, and back of this form
the superior rims of the orbits. Posteriorly they unite with
the postfrontals, parietals, and supraoccipitals. The two
bones are united in the median line by a distinct suture, and
possess no characteristic markings other than an irregular
cto
STEWART | Cretaceous Fishes. 279
articular surface near the anterior extremity, which probably
gives attachment for a preorbital bone.
The prefrontals are stout bones, and are especially character-
ized by the presence of a large anterior facet for articulation
with the malleolar portion of the palatine. In the figure of the
skull of Niphactinus, as given by Crook,” the prefrontals are
represented as extending well backward and forming a portion
of the anterior and superior rim of the orbit. In this I am in-
clined to think that he was in error, for in several more or less
disarticulated skulls in the collection I find that this supposed
posterior portion of the prefrontal is a distinct bone which has
become separated, showing a well-marked suture where it
united with this bone and the frontal. This bone is the supra-
orbital mentioned above, and, is rather thick and lunate in
outline.
The parietal bones have been much in doubt by most authors,
probably owing to the fact that this portion of the skull is often
much distorted and crushed, thus obliterating many of the
sutures, or making them so indistinct as to render them un-
recognizable.
Cope first thought that the supraoccipital might represent
the conjoined parietals,” but concluded later” that this was not
the case, and that the bones he had called epiotics were the
parietals. Crook states that the parietals are completely sepa-
rated in the median line by the supraoccipital,” but seems to
have been somewhat in doubt about the exact extent of these
bones, as the anterior portion is but poorly defined in his figure
of the specimen on plate XVIII. Professor Hay has described
them“ as wedge-shaped narrow bones which lie between the
anterior ends of the pterotics and the posterior ends of the
frontals, probably meeting each other in the median line, and
including the elevated granular portion assigned by Crook to
the supraoccipital, in which I think he is right. If we ex-
amine a well-preserved skull of Jchthyodectes we will at once
36. Paleontographica, 1892, pl. XVII. 39. Paleontographica, 1892, p. 115.
37. Cret. Vert., p. 183. 40. Zool, Bull., vol. II, pp. 28, 29.
38. l.c., p. 188.
280 University Geological Survey of Kansas.
see that this portion is entirely distinct from the supraoccipital,
and that the bone he calls parietal is included in the pterotic
and is not separated from it by suture.
The epiotics are wedged in between the pterotics and supra-
occipital and form prominent inner processes of the skull.
They become narrow anteriorly and are provided with promi-
nent ridges along the superior borders, which end in slight
upward projections posteriorly which are thickened at the ex-
tremity.
The supraoccipital forms a prominent crest and is projected
well backward and upward. In some of the specimens having
the posterior end preserved, I find that there are a number of
tooth-like projections which closely resemble a suture, but they
probably gave attachment to muscles alone. The bone is
moderately thick and is somewhat rounded along the superior
border. I think that I can detect the suture which separates
this bone from the granular portion mentioned above. The bone
is somewhat irregularly striated on the sides, and posteriorly it
descends almost to the basioccipital, forming a prominent narrow
ridge in the median line.
The pterotics lie just external to the epiotics and back of the
parietals, and form prominent outer processes of the skull. The
bones are very robust, and have deep grooves along the external
borders, which articulate with the superior condyles of the hyo-
mandibulars. The posterior extremities are somewhat ex-
panded, and have small elliptical surfaces internally, which are
covered with coarse strize radiating from near the center. These
surfaces probably are connected with the post-temporal. The
sutures separating these bones from the parietals are often in-
distinct.
The postfrontals form prominent processes just back of the
orbital cavities, and anteriorly they form the posterior sides of
the conspicuous notches which accommodate the posterior su-
praorbitals mentioned above. Posteriorly, by their union with
the pterotics and prootics, they give a small support to the hyo-
mandibulars, and are excavated along the sides for this purpose.
Internally they unite with the parietals.
STEWART. | Cretaceous Fishes. 281
The prootics have been correctly interpreted by Professor Hay
as being the largest of the otic bones. They are irregular in
outline and extend from the groove for the hyomandibular
above to the parasphenoid below. Posteriorly they unite with
the opisthotic, but are not separated from the basioccipital by
the opisthotic, as Professor Hay has figured them in Tarpon at-
lanticus.** Just below the hyomandibular facets there is a large
foramen on each side, which, according to Professor Hay, trans-
mits the glossopharyngeal nerve. Just back of these foramina
there are large fossx, the boundaries of which are formed by
the pterotics, opisthotics, and prootics.
The sutures separating the lower portions of the opisthotics
from the basioccipital and the prootics are well defined, but the
ones separating them from the pterotics above are almost oblit-
erated, although I think they can be traced in one of our speci-
mens as extending downward just back of the articular face
for the hyomandibulars. They are small bones and do not
separate the prootics from the basioccipital, as mentioned above.
The basioccipital is deeply concave, and the exoccipitals are not
well defined in any.of our specimens.
The parasphenoid is triangular in section and forms a strong
lower axis of the skull. The posterior extremity of the bone is
deeply emarginate where it joins the basioccipital, a short dis-
tance in front of which it sends up a short process on each side
for union with the prootic. Extending outward from the sides
of the bone, just in front of the brain-case, there are two well-
developed transverse processes which are bluntly pointed and
directed slightly forward. At the base of each of these proc-
esses there are two foramina which, according to Crook, trans-
mit the facialis and trigeminus. Just above the transverse
processes there is a Y-shaped bone, called basisphenoid by Hay,”
which unites with the prootic above. The anterior end of the
parasphenoid is somewhat broadened and bifurcated for union
with the vomer. The latter bone is not well defined in any of
41. Zool. Bull., vol. II, No. 1, ». 28.
42. 1, Cus Ps See
19—vi
282 University Geological Survey of Kansas.
the specimens, owing to the crushed condition of this part, thus
making the line of separation indistinct between it and the
ethmoid. The orbito- and alisphenoids are also in a much dam-
aged condition, so as to render their boundaries only conjectural.
The orbital cavity was large, and bounded above by a chain
of thickened supraorbital bones, and below by a chain of thin
suborbitals, which seem to be quite large. Below this there
seems to be a sheet of thin membranous bone covering the hyo-
mandibular and the bones of the palato-quadrate arch. The
sclerotic ring is composed of two pieces, which are found in
place in one of our specimens. The outer borders of these are
very heavy, but become thin internally and form a slight ring,
which is extended inward.
Just below the suborbital ring there is a long, thin bar of
bone which rests on the superior border of the maxilla, and is
covered with minute canals leading inward. This I take to be
the same bone called a ‘‘ supernumerary bone’’ by Cope,” and
an ‘‘extraknochen’’ by Crook.* It seems to me that both of
these authors have been in error in regard to this bone, and
that it is the jugal. Crook thought that the jugal was found
just beneath this, and has it so marked in the figure of his skull
of Xiphactinus,® but I am unable to find any such bone in our
specimens.
PECTORAL GIRDLE AND FINS.
The pectoral girdle is represented by numerous specimens,
some of which are preserved almost complete, thus giving a
more correct idea of the form than any of the specimens hereto-
fore described. This part was not well understood by the early
writers on this form, and it was not until about a year ago that
Professor Hay demonstrated*® that this portion had been de-
scribed in an inverted position.
The girdle is composed of the cleithrum, scapula, coracoid,
and precoracoid, of which the first is the largest and the second
the smallest and most compact. The cleithrum is a long bar of
bone composed of two parts, which are bent almost at right
43. Cret. Vert. West., p. 194. 45. l. c., t=. XVIII.
44, Paleontographica, 92, s. 115, 116. 46. Zool. Bull., vol. II, p. 42.
STEWART. | Cretaceous Fishes. 283
angles with each other, and are separated by a slightly con-
stricted neck just in front of the scapula. The anterior portion
is directed slightly downward at the extremity and is very thin,
fragile, and it is probably owing to this fact that it is secured
in so few specimens. Just above the scapula the cleithrum be-
comes very broad and continues so to near the upper extremity,
when it suddenly contracts in width toward the anterior border,
and ends in a somewhat thickened process, below which there is
a well-marked dentate suture for the anterior end of the precora-
eoid. The whole of the anterior border is firmly united with the
precoracoid, and the external side is covered with coarse stria,
which radiate upward and backward from just over the scapula.
The scapula is a rather small bone, composed of a heavy com-
pact portion which bears the glenoid surface and a rather thin
lamina of bone which extends upward along the internal side
of thecleithrum. The glenoid portion is united to the cleithrum
above by means of a well-marked undulating suture, and bears
three condyles, of which the uppermost is the largest, and is
separated from the middle one by aslight ridge. The middle
condyle is the smallest and is separated from the lower one by
a slight groove. The condyles are all convex from before back-
ward, the superior one very much so. Just internal to the
glenoid surface there is a large irregular surface which gives a
strong attachment to the coracoid, and also gives a partial sup-
port to one of the basiosts posteriorly. Just above this portion
the precoracoid is united by a strong suture and extends up-
ward as a broad bar on the internal side of the cleithrum.
The coracoid has been figured by Hay” as a broad, thin plate
of bone which extends forward to probably the anterior end of
the lower arm of the cleithrum. It is not well preserved in
any of our specimens, but from the portions that are present I
am inclined to think that it is the same bone described and fig-
ured by myself as a probable infraopercular.* It is somewhat
thickened where it joins the scapula and has one deep pit anda
portion of another for the extremities of two of the basiosts.
47. 1. c., p. 43.
48. Kans. Univ. Quart., vol, VIII, p. 21, pl. XI.
284 University Geological Survey of Kansas.
There is a great variation in the size of the various specimens,
and the measurements given are below the average, this speci-
. men being selected on account of its completeness.
Cleithrum: Length from scapula superiorly.................. 233 mm.
Length from scapula anteriorly.................. 230
7 Precoracoid:)-heneth ieee eee yc 2 ean oP Ca GOR mS
Numerous fragments of fins and fin-spines are present, which
show that these formed powerful weapons of defense; they haye
been described by Crook as resembling ribs. The first pair of
these are closely applied to each other and are each provided
with articular surfaces at the proximal extremity, the first of
which articulates directly with the upper condyle of the scapula
while the second articulates with one of the basiosts. The first
of these is broad and thin, while the second is slightly narrower
but thicker toward the proximal end. The two opposing sur-
faces are much roughened while the outer surfaces are finely
striated. The following rays decrease greatly in width and are
much smaller from above downward. ‘They are slightly bent
at the proximal extremities and the cross-segmented character
has disappeared throughout. The basiosts are three in
number. The first of these is an irregular-shaped bone which
is provided with two facets internally for articulating it with
the two lower condyles of the scapula. These two are some-
what oval in outline, flat, and well separated from each other.
The external facets are also two in number, one of which is
nearly circular in outline and flat, while the other is irregular
and strongly convex from above downward. The two remain-
ing basiosts are somewhat thickened bars of bone which are
slightly expanded at the extremities and are provided. with con-
vex condyles, the proximal ones of which are received into the
pits on the coracoid and scapula for their reception. About
midway between the extremities of each of these basiosts there
is a thin transverse process extending outward.
The pelvic actinosts, called femora by Professor Cope,* con-
sist of two parts—a massive posterior portion, bearing the
facets for the attachment of the pectoral fin, and the thin,
49, Cret. Vert. West., p. 186.
ee eee
}
|
.
;
j
]
‘
STEWART. | Cretaceous Fishes. 285
somewhat wing-like, anterior portion, whose entire structure is
not known. In the median line the two halves are strongly
united together by suture, and at the point of union there is a
prominent swelling of the bone all around. There are four
facets on each for the fin attachment. The upper and lower
are large, flat facets, somewhat ovoid in outline, and support
the upper and lower halves of the first ventral fin-ray. When
seen in cross-section from behind they are obliquely set, but
hardly so much so as in Jchthyodectes. The lower one of these
facets is the larger, and has two more facets between this and
the upper, of a somewhat tubercular nature. The last two are
for succeeding rays or basiosts. In front of the articular por-
tion the bone becomes thin and somewhat expanded. On the
outer border of this portion there is a prominent crest both
above and below, of which the upper is the larger. It is likely
that this crest did not extend very far forward, but was prob-
ably succeeded in front by another smaller crest or tuberosity,
as something of the kind is shown in the figure of this part by
Professor Hay,*’ but not described in the text. On the internal
border there is a prominent ridge of bone extending forward on
both the superior and inferior surfaces. These two ridges
nearly coincide with each other, thus forming a rod of bone,
which extends forward, but how far is not known. These two
bars are separated by quite an interval, as is shown in the cut.
There seems to be some difference between this specimen and
the one figured by Professor Hay. In his specimen the distance
across the external crests was probably about 15 cm., and the
distance between the internal ridges about 4.8cm. In our speci-
men the first of these measurements is 12.3 cm.,”' while the last
is 5cm., making the distance between these ridges proportion-
ally greater in our specimen.
iene Herons BUperior {AGES Lee ab Ms 83 mm.
eee Rt PED RELVES COO 5, oo iad aru pina, i vin vie ¥ alecemen ade Way **
PIStAnCe BCIORS MIGGIC TAGS... 6... te cee eee eean ne enar 5.16
Vertical depth of articular portion.............c0cc0dvecece ae
There are none of the pelvic fin-rays present, but they were
50. Zool. Bull. 1898, p, 44.
51. Approximated.
286 University Geological Survey of Kansas.
probably eight or nine in number, of which all were very feeble
excepting the first. :
There are five vetebral columns of this genus preserved, two
of which are practically complete, and the other three only par- ©
tiallyso. There are also portions of several more columns, but,
as they have not been numbered in the positions they occupied
in life, I will not use them in the table given below. The most
complete column has 77 centra present, which I think is all
the animal had in life, with the exception of one of the terminal
caudals. ‘The other contains 73. Unfortunately, the neural
and hemal arches are gone, or so badly preserved that their
characters cannot be made out in most instances. The first
anterior vertebra is much shorter than those following, and the
anterior end is not so deeply concave. It presents deep pits
above for the neurapophyses, but none below for the pleura-
pophyses, the points where they should be being marked by scars.
This one and the two or three following are usually without
lateral grooves, but the second and third sometimes show them.
The pits for the neurapophyses are large in these, as in the first,
and the pits for the pleurapophyses become functional at about
the third or fourth.
Back of the vertebre just mentioned, the pits for the neura-
pophyses begin to assume the elongated form found farther back
in the column, and the number of lateral grooves on the sides
become somewhat varied.
To illustrate this last point, I have arranged the following
table to show the individual variation in the different speci-
mens. In this table the numbers at the heads of the columns
represent the catalogue numbers of the various specimens,
while those on the left are the numbers of the various vertebre
from the skull. A blank is left in some places where the ver-
tebra is badly injured. The grooves are represented by 1-1
when there are two well developed, 1-0 when only one is pres-
ent. When there is one well developed and another slightly
so, they are expressed by 1-1— or 1-0+, 1-1— expressing a
slightly greater development than 1-0-+.
Srewakt. | Cretaceous Fishes. 287
No. No 1 ss
Me i Poe. 160... 6-0: | 0-0 oP 402.0.. 0. as. Tea de
- a eee eed) 0-00 10-8 0-0" |Y) ALc fe ..ce..e. ton) 3
“ee CSS SESS Ee es | eam |
PE et ces. Eon og EUG gal ag he gS ea
See Grae WA) AA OA deo WE aA, ive dies 10+}11
ae cy TS NI Top i Ts Pa 0) ia
ee ee Peek Ger eee age rye: TeREey aba
re ae ee TESTES OSS ES Trig coe es | Se EH i tii
[ORS Grae aa ere ed te Wo . vB pa
_ ee eee Bi tet: Witt) Aad AD os. ces ie as
BE Sc 52. ox: CMs ee ee re
eee Fein i car a Wye te ok 1 ie es | Uys kai a gr
i ae i Somos ac. Pirie fot eG sy oi TOR ala
te oS (Ore dos ola ee eos |e he ae | Dane) Tl
Rete s! Ss. See Linoe Settee a da GA 522), 3. Ali h i)
ae NR STS AG Gir SN Wi 104/11
ee Up gc ye oe A yen gy a | POL ert
Meee el... CSAS ea be ie Clie RS a ee ee tlh i
ae Cg a Se ee Se ese re Lg Be
_ ee Te oh LVS a Sv hat Res gs eee | os aay
ee EA Se 0 ne is | 16h het
Meese... fo 19 RT eS or Ta a ri | Ry Pee a
2 ee iid ge ee aa ie en Gober, fs 10+] 14
es Bee ites ee Te a. 2, bo tie ae 1.0 R11
5... Es eS las Re a ed a Gia ces ie ih arG
Dee salle + ETE TET CE Teg ag Tae ee | bOI
a TM gg SiR Ss Om a | > 10+] 10
eee fey. Pen dO pa OA do HG mck. ee oie
_ AS aN ie ot a ae eee ee peer
ne ec TS be Seg 2 or SE es hs |: (ee
Ss 1 {11 1S NO a0 pa | yo 10+] 1-0
ee TA TR a ee eB a a ee 6 i 0+| 1-0
a 11 }11 more fs baat ES ig a ae 2. E08
EY is pg a ee Poo 5 Sch eee oan Oeaeae 00
a a thes io a MN MALTS ati fob he eh ee ae 00 |00
AE 5 I hos Sees Op: ORT ON ere FO: gars
_ aeaieieats ee eee ek Ba ie (eas te ee a meee
_ es BO Ltd mtn cc sind ce |r ad | Aa rae GO He
SES LIS)... ree ee Like |
The ribs are not joined directly to the vertebra, but articulate
with small masses of bone set in deep pits on the side of the
centrum. Each of these has a concavity for the head of the rib.
According’ to Professor Cope, there are six vertebre in the
terminal caudal series,” none of which, I am inclined to think,
bore lateral grooves. They all, with the exception of the last,
have broad surfaces below for the attachment of the hwema-
pophyses, which are modified into two surfaces, one behind the
other, in next to the last vertebra. The last is a small tubercle
52. Cret. Vert. West, p. 199.
288 University Geological Survey of Kansas.
of bone with a quadrilateral face in front and a small spine pro-
jecting upward and backward behind. On the internal side
there is a large, slightly concave facet, which has the appear-
ance of having been united with a similar bone on the opposite
side. All of these vertebree are curved slightly upward.
Four of the hemal spines from this region are present in one
specimen. ‘The two anterior of these are very similar, being
broad, flat, and in contact with each other throughout. At the
upper extremity of each there are two large, roughened, sutural
surfaces for uniting the spine to the centrum. Between these
two surfaces runs the small hemal canal. The one following
these two is not so flat but slightly thicker, and has the upper
end expanded into a knob-like mass of bone. The articular
surfaces have become conjoined and form a saddle-shaped ar-
ticulation with the centra, the hemal canal having become
somewhat depressed. The spines following this have become
modified into somewhat fan-shaped hypural bones. The upper
extremities of these present convex condyles, which are proba-
bly received into pits on the last two vertebrx mentioned above.
There is one hemal spine from somewhere in front of the
caudal region which is of a different form from those described
above. It is a Y-shaped bone with elongated and roughened
articular surfaces on each of the arms for uniting it to the cen-
trum. The hemal canal is very large and below it the bone is
rather thin.
The caudal fin-rays are represented by a mass of spines, only
one of which is complete. The anterior of these are rod-like,
but those following become expanded at the upper end, where
they are longitudinally striated. These are followed by rays
which are broader and probably shorter, on which the striew
just mentioned are more pronounced.
The neural spines are represented, in good condition, by only
three or four from just back of the skull, some of which I have
been able to fit to the vertebree to which they belong. They all
have knob-like expansions at the lower end which are received
in the pits on the top of the centra. The two halves are in con-
tact with each other above the neural canal and nearly or quite
STEWART. | Cretaceous Fishes. 289
so below. Extending backward from above the neural canal
there is a somewhat expanded mass of bone which has an artic-
ular surface on the front of the arch following. These have a
superficial resemblance to the zygapophyses of the higher verte-
brates. Just opposite to this on the external side there are two
more processes which extend outward and backward. These
are what Professor Hay calls epineurals.” They are largest at
the head and become much smaller at the fourth or fifth vertebra,
and probably become insignificant or entirely disappear further
back. The neural spines of these are directed upward and ob-
liquely backward.
As we have no well-preserved neural arches farther back than
those described above, I will quote Professor Hay’s excellent
description of the structure of these in the caudal region.
Copies of his figures are given on plate XLII.** The neural
arches here, as elsewhere in this fish, are. connected with the
centra by suture, and have usually fallen out before burial,
leaving long grooves where their bases were inserted. This was
the case with the third vertebra behind the right-hand one
shown in fig. 1. When we come to examine the arches more
closely, we discover that each lateral half is not a single piece,
but consists of two pieces, a basal piece (a, n, a) and the arch
proper (vn, a). That the proper arch is a distinct piece is
shown not only by the existence of a suture, but likewise by
the fact that in the vertebra on the left hand of the figure the
arch has fallen out of its place before fossilization. The basal
or accessory piece is inserted by a shallow gomphosis into the
centrum for nearly the whole length of the latter. It rises high
in front, and projects so far forward as to come in contact with
the basal piece of the next vertebra in front. Behind, the basal
piece is directed upward and backward in a rather slender
process, which abuts against the anterior edge of the basal
piece of the next vertebra behind. It is thus seen that these
basal pieces provide the anterior and posterior zygapophyses.
They remind us of the articulating processes of certain other
53. Zool. Bull., vol. II, No. 1, p. 51.
54. See, also, Zool. Bull., vol. II, No. 1, pp. 47-51.
290 University Geological Survey of Kansas.
fishes (Mugil, etc.) Between the anterior and posterior processes
the basal pieces are excavated to receive the bases of the neural
arch, as shown in the figure. The two basal pieces of each
vertebra are distinct. Together they seem to form a saddle in
which the neural arch rides.
I find this same structure of the neural arches in some of
the vertebree belonging to specimens in the United States Na-
tional Museum; but in one section of connected vertebrae an
arch like those above is succeeded in the next vertebra by an
arch in, which every trace of a suture between the arch and the
apparent basal piece is lost. ‘This vertebra is shown in fig. 2.
The form of the base of the arch is not greatly different from
that of the arch with accessory piece in fig. 11,” and we may
even convince ourselves that we can trace a part of the bound-
ary line between the two portions. There is evidently at this
point of the vertebral column a sudden change from neural
arches furnished with basal accessory pieces to arches without
these, or consolidated with. them. Further backward the form
of the arches becomes modified somewhat, so that they resemble
the one shown in fig. 3. A section fourteen inches long and
containing seven vertebre having arches of this kind is before
me. This condition shows us that the neural arches which are
provided with basal pieces are confined to the anterior or middle
portion of the tail region, while the hinder portion contains no
such vertebral structures. We are reminded that in Amia the
middle portion of the caudal vertebral column is composed of
two for each muscular segment, while the anterior and poste-
rior portions have the vertebral centra of the ordinary kind.
It seems as if the tail portion of the vertebral column of the
amioid fishes and of the Isospondyli retained primitive condi-
tions longer than the abdominal portion.
It is difficult to determine what explanation is to be given of
the presence of these basal pieces. The so-called zygapophyses
of fishes are regarded as being outgrowths of the neural arches
— exogenous and not autogenous processes. It might be said,
55. Fig. 11, in Professor Hay’s article, refers to the pelvic actinosts, and seems to have no
connection with the subject.
STEWART. | Cretaceous Fishes. 291
possibly, that the basal pieces are the proper arches, while the
pieces which are borne on them are the spinous processes. I
hold that there are two objections to this view. The firstis, that
what are sometimes called spinous processes are always un-
paired pieces. The second is, that when the lateral halves of
the arches remain distinct from each other and are prolonged
into spines, as they are in various fishes— Ama and Salmo, for
instance —the spinous portion is never, so far as we know, de-
yeloped in the embryo as pieces separate from the base of the
arcula. ‘This is true in the case of Amia, which I have investi-
gated. We must, therefore, seek some other explanation. The
key to the understanding of the problem is, it seems to me, to
be found in the vertebral column of that primitive fish, Ama.
We may call this fish to our assistance, since the Isospondyli are
believed to have ancestors not far removed from Amia.
In the middle region of the tail of Amza there are, for each
muscular segment, two vertebral rings, the one bearing the
arches, upper and lower, the other archless. If a transverse
section be taken through the arch-bearing ring, there will be
found an X of cartilage, the upper arms of which are continu-
ous with the cartilage of the neural arch. In like manner, the
lower arms will seem to be continuous with the cartilage of the
hemal arch. If a section is made similarly through the arch-
less disk, a similar X of cartilage will be found, but the arms
project beyond the outer surface of the disk but a short distance.
These archless disks are developed in Am7a from ossifications
arising in the intercalated cartilages, upper and lower, and the
arms of the X are the unossified portions of these cartilages.
There appears to be no reason why these intercalated cartilages
should not sometimes take on a hypertrophied growth. In the
sharks they often become considerably larger than the neural
‘arches themselves.
In case these intercalated cartilages should become thus en-
larged and arch-like, each might develop a bony investment
that would simulate the bony neural half-arch, and thus would
rest on the top its proper epicentrum.”
56. For figures illustrating the architecture of the vertebral column of Amia, see the May
number of the American Naturalist, 149s,
292 University Geological Survey of Kansas.
Coming now to the anterior region of the vertebral column
of Ama, we find that each vertebra is formed through the
suppression of certain of the elements which, in the tail region,
constitute the vertebral rings or disks, and the union of the re-
maining elements of each muscular segment into a single mass.
The lower intercalated cartilages are suppressed. The upper
intercalated cartilages hypertrophy, and their ossifications unite
with the bones developed in the bases of the lower arch, thus
giving origin to the centrum. The ossification that we might
expect to find developing in the base of the cartilaginous neural
arch, the epicentrum, is absorbed, while the ossification of the
enlarged intercalated cartilage, the pleurocentrum, pushes
itself into the place of the epicentrum, and thereafter supports
the neural arch.
Now, we have the choice of two suppositions, neither of which, -
however, may be the true one. We may hold that a distinct
bone was developed in the somewhat elongated and projecting
intercalated cartilage, and this, of course, rested on top of the
pleurocentrum; when the latter was pushed forward beneath
the neural arch to take the place of the aborted epicentrum,
this newly developed bone was carried along, and thus brought
between the pleurocentrum and the base of the neural arch.
Or, we may hold that the bone which I have found in Xzphac-
tinus supporting the neural arch is simply the epicentrum itself,
aborted, indeed, in Amia, nevertheless persisting in \iphactinus,
but crowded upward out of its original seat on the notocord.
Neither of the above suppositions presupposes that the upper
half of the vertebral centrum takes its origin from the pleuro-
centrum. Professor Cope held that the vertebre of fishes are
‘‘intercentra’’—that is, have originated in the suppression of
all the other elements through the excessive development of the
hypocentra. But the very existence, in many genera, of a
cartilaginous X in a transverse section of the centrum is proof
that its upper portion was derived from either the bases of the
upper arches or the pleurocentra. The deep gashes in the ver-
tebral centra of Xiphactinus, where the arches have fallen out,
furnish evidence that this cartilaginous X was present.
)
:
STEWART. | Cretaceous Fishes. 2938
Xiphactinus lowii. Plate XLVIII, fig. 2.
Portheus lowii Stewart., Kans. Univ. Quart., vol. vir—a, p. 24.
This species is based on the dentary bones of a single indi-
vidual found at Fairbury, Neb.,in the same horizon of the Fort
Benton Cretaceous with Desmatochelys lowii, and was sent to the
museum by Mr. M. A. Low, of Topeka, in whose honor the
species is named. While this specimen has not been reported
from Kansas up to the present time, yet it no doubt occurs
here, as the same horizon from which it was obtained is found
just across the line in this state. Special interest is attached
to this specimen, as it is the first of this genus to be reported
from so low a horizon as the Fort Benton.
The dentary is short, with a more oblique symphysis than in
any other specimen of Xiphactinus that 1 have examined. It is
also not so roughly marked at this point for the attachment of
ligaments as in XY. audax. The alveolar border is short, and
does not have the prominent swelling just back of the symphy-
sis which is found in the species just mentioned. The posterior
extremity is projected upward into a short coronoid process,
which is bent outward but slightly. The teeth are slightly
oval in cross-section, acutely pointed, directed slightly back-
ward at the extremities, and non-striate even under the micro-
scope. Their arrangement is as follows: One large, two small,
one large, ten or eleven medium and small. The number and
arrangement will no doubt vary with the individual.
RUS h OF BIVEOlAY POFOO? s).. 1) i clekic dd casei sh sd alae 177.0 mm
MEE SE, URMEMEV NISL ed ibe eve bie ba) wciik o's s ooo Stein's yay
Depth of bone just back of symphysis.................. Ga.0'
RID On CONMGATY Gh TIAGO 245 cia ki oo es mpd went see 64.5 *
Xiphactinus brachygnathus. Plate XL\, figs. 3, 4.
Niphactinus brachygnathus Stewart, Kans. Univ. Quart., vol. viii.
In addition to the forms described above, there is one speci-
men that I am unable to locate as X. lestrio or X. mudgei,
described by Professor Cope. As has been shown above, Cope
based most of the specific differences on very inconstant char-
acters in his description of Y. molossus and X. thawmas, and as
he has used much the’same characters in his description of the
294 University Geological Survey of Kansas.
two species mentioned above, I am inclined to think that these
will have but little specific value until the type specimens of
each are redescribed and figured in a way to render them recog-
nizable. |
The specimen, No. 155, consisting of the upper and lower
jaws almost complete, fragments of the skull, and numerous
vertebre, was collected by Prof. B. F. Mudge from the Niobrara
Cretaceous, four miles north of Gorham, Kan.
The premaxilla is somewhat oval in outline, and the anterior
portion is thickened instead of the central, as is usually the case
in X. audaz. The posterior border is thin, and is peculiar in
having the internal side of the bone beveled off to meet it, some-
what similar to that found in /chthyodectes and Saurodon. The
alveolar border is quite elongate, and supports three teeth on
one side and an empty alveolus for a fourth, all of which are
small, with the exception of one.
The maxilla is especially characterized by the manner in
which it unites with the premaxilla, for instead of having the
abrupt change to the laminar portion in front of the posterior
condyle, the bone is gradually beveled off to a sharp anterior
edge, similar to the condition found in Ichthyodectes. Both of
the superior condyles seem to be small, and the anterior one is
directed inward to a considerable extent. The two are well
separated from each other. The bone is quite deep at the pos-
terior condyle, and the superior border, back of it, presents a
conspicuous groove for the attachment of the jugal. There are
nineteen or twenty teeth on one side, the most of which are
small or medium in size.
Premaxilla: Length of alveolar border. .................. 62.0 mm.
Greatest depth of toome:)..0. =. eee eee 90.0 *
Maxilla: eneth of alveolar border...) .4... eee 240), 2C
Depth: of condyle. 00) 22.2220 ee Soke ae eee 83.5
The dentary bones are short and deep, thus giving them a
very robust appearance. The bones are much thickened at the
symphysis, and slope backward nearly as much as in X. lowi.
The alveolar border is short and is remarkable for the small
number and large size of the teeth toward the anterior extremity.
STEWART. | Cretaceous Fishes. 295
The arrangement is three large, two small, one large, and the
twelve or thirteen medium and small. Many of the alveoli are
empty, so the size of these teeth has to be estimated; the num-
ber of these, however, no doubt varies with the individual.
The coronoid process is but poorly developed. The dermar-
ticular invades the dentary but little externally, and the cotyloid
process of this bone is not so well developed as in \. audaz.
See Gk BIVOOMT HOLUET, 24 Sots). Sac hes swe sce sins 5s 215 mm.
Depth of coronoid process......... Per tee Cee es Lis
Mineo nna eae ESVATIPMIE VENA, 38 S00 Le Bcd Cl bs Ss dave h ee ck ae. 105; **
Depth of bone just back of symphysis................... TUS FF
Penne Te IONS FLOM) COPYIUS, oct 2 wie ee le eee ceeds A (a
There are fragments of several other bones, among which
are the ethmoid, a prefrontal and palatine, and several verte-
bre. The ethmoid is acutely pointed anteriorly, and the pos-
terior suture is very dentate. The prefrontals are small, as are
also the malleolar portions of the palatines. The external
tuberosities of the latter are not so prominent as in LY. audaz.
The vertebree do not seem to differ materially from the species
mentioned.
ICHTHYODECTES.
Cope, Proc. Am. Phil. Soc. 1870, p. 536.
This genus is so closely related to Xiphactinus that it does not
need to be characterized, beyond pointing out some of the prin-
cipal differences between the two forms. The teeth are more
regular in size and never reach the enormous development of
those in Xiphactinus. They are cylindrical in cross-section and
are usually directed slightly inward at the apices.
As indicated by the size and dentition, this genus was much
less ferocious than the form mentioned above, which no doubt
preyed upon them to a great extent. The known American
species, all from the Niobrara Cretaceous, Kansas, are:
Ichthyodectes anaides Cope. Ichthyodectes goodeanus Cope.
Ichthyodectes ctenodon Cope. Ichthyodectes acanthicus Cope.
Ichthyodectes hamatus Cope. Ichthyodectes pernicosus Cope.
Ichthyodectes prognathus Cope. Ichthyodectes cruentus Hay.
Ichthyodectes multidentatus Cope.
During the past summer, while collecting in the Fort Pierre
group, at Lisbon, Logan county, Kansas, I discovered a string
296 University Geological Survey of Kansas.
of five vertebrze which I am inclined to think belong to this
genus. They are about the same size as those of J. anaides,
but, as there is nothing about them that would characterize
them specifically, nothing more can be done with them until
more complete specimens are found.
Ichthyodectes anaides. Plate XLIX, figs. 1-3.
Ichthyodectes anaides Cope, Proc. Am. Phil. Soc. 1871, pp. 339, 340.
This species is represented by the remains of several indi-
viduals, including the jaws, portions of the vertebral column,
and one skull without the jaws, which probably belongs to this
form.
The maxilla is represented in one specimen in a compressed
and somewhat fragmentary condition, the posterior extremity
being absent. The bone, as a whole, is about the same size as
the maxilla of J. cruentus, but is much thinner in cross-section
than in thisform. The bone is moderately deep at the posterior
condyle, which is slightly convex. Back of the condyle the su-
perior border presents a sharp ridge which continues for some
distance. Just back of the premaxillary surface the alveolar
border is slightly concave and then slightly convex. The teeth
are small, about 4.5 occurring to the centimeter.
Maxilla: Depth-at pasteror condylec ea. eee eee een eee 33 mm.
Thickness Mack of condyle. 03). 0 Y oi cies cena LO a
I am inclined to think that the dentary bone figured by Pro-
fessor Hay” as that of J. cruentus belongs to this species. The
bones are short and deep, with moderately sloping symphyses.
There are aiso two conyexities in the alveolar border, the pos-
terior of which is not shown in Hay’s specimen owing to this
extremity of the border being absent. ‘The coronoid process is
not prominent, and the border seems to be very steep below and
back of it. The teeth are large, directed slightly inward at the
apices, and without strie. The lower border is thin and has a
number of large foramina just above it.
Wentary: Wength of alveolar borders. 3 2.2 eee ee eee ee 99 mm.
Depth of symphysis-i2.0-4+ 5 eee BEER Eis Py AR pt 7 a
Depth at coronoid process (estimated).............. Done
Number of teeth in one centimeter.................. 3
57. Amer. Jour. Sci., VI, p. 227.
|
.
|
|
STEWART. | Cretaceous Fishes. 297
The quadrate is thin and fan-shaped, and has the condyle
bent forward to a considerable extent.
The skull that I regard as belonging to this species is much
crushed, so that many of the sutures on the top cannot be made
out. The otic region, however, is well shown. The ethmoid
seems to have been rather blunt. The frontals are narrow, and
each has a broad and prominent ridge which extends outward
and forward from just in front of the supraoccipital and ends
just above the prefrontals. They seem to have had supraorbital
bones attached along the border. The prefontal condyles are
large, and the postfrontals are prominent and offer a consider-
able support for the hyomandibular. The pterotics are crushed
beyond recognition above, but laterally the groove for the hyo-
mandibular is very prominent. I am unable to make out the
parietals in this specimen, but in another and much smaller
skull I find no suture separating it from the pterotic, as figured
by Crook,* and am inclined to think that the expanded por-
tion of the supraoccipital represents the coalesced parietals, as
has been suggested by Professor Hay.” The supraoccipital
crest extends upward and backward at quite an angle, but the
extremity is broken away. The epiotics are prominent and do
not extend far forward. The prootics and opisthotics are very
similar to those of Xiphactinus. The basioccipital is deeply
concave and expanded in front of the condyle. The parasphe-
noid is bifurcated posteriorly, triangular in section, with rather
small transverse processes in front of the brain-case.
So far as can be determined the vertebrae are all two-grooved,
but near the posterior end of the column they become rather
faintly marked. ‘The neural spines are long and longitudinally
expanded at their proximal ends. The caudal fin was much
expanded and composed of large rays, which become flattened
and longitudinally split toward the distal ends. The scales are
large, with small grooves radiating outward from near the cen-
ter on one of the sides.
58. Paleontographica, p. 92, pl. XV.
59. L.c., p. 230.
20—vi
298 University Geological Survey of Kansas.
Ichthyodectes hamatus. Plate XLVII, fig. 2; plate L, figs. 1-7.
Ichthyodectes hamatus Cope, Proc. Am. Phil. Soc. 1872, p. 340.
This species is represerted by a fragmehtary skull and ante-
rior vertebree of one individual and a portion of a dentary and
a hyomandibular of another.
The maxilla is a long, slender bone, which is thickened
along the superior border and rather thin along the inferior.
The alveolar border is remarkable for the way in which it is
directed downward anteriorly just back of the premaxilla. This
border is concave from before backward, and supports alveoli
for nearly fifty teeth, about half of which are functional at once.
The surface for the premaxilla is directed inward and is coarsely
striated. It is similar in many respects to J. cruentus, and the
premaxilla was no doubt directed downward as in this species.
The posterior superior condyle is irregular in outline, and has
its internal portion continued downward on the inner side.
The anterior condyle is rather small, directed inward, and sepa-
rated from the posterior condyle by along, smooth space. In
front of this the anterior border descends almost vertically.
The posterior extremity is expanded and directed slightly
upward. ‘The external surface is finely striated above, and
covered with small pits just above the alveolar border. Un-
fortunately the premaxilla is not present.
Maxilla; Wenge thot alveolar borderee Sere ae eee 117 mm.
Depthioteposterioricondyles ss. 4 re eee ee eee 310) Ae
Distance between superior condyles................ DOR oe
Number of alveolse in one centimeter............... 4
The condyle of the quadrate is elliptical in outline, and is not
projected forward as much as in /. ctenodon. The groove for
the symplectic is broad, and reaches downward to within 22
mm. of the condyle, below and back of which the bone is much
thickened.
Only the upper portion of the dentary is preserved, which
shows this part to be thicker in cross-section than in J. anaides.
The anterior extremity of the alveolar border is produced up-
ward into a hook-like process, from which character the species
probably derives itsname. This portion bears two teeth and is
STEWART. | Cretaceous Fishes. 299
directed slightly inward. The symphysis is much thickened
and was about the same slope as that of J. anaides. Just back
of the hook-like process mentioned above the alveolar border
is somewhat excavated, and back of this slightly convex fora
distance of 55 mm., when it is directed slightly upward. The
teeth are closely set with elliptical crowns, the long axis of
which is placed obliquely to the tooth line.
Dentary: Length of alveolar border (estimated)............ 125.0 mm.
Length of long axis of cross-section of tooth...... Oe?
Length of short axis of cross-section of tooth..... paves
Number of teeth in one centimeter............... 3
The hyomandibular shows the anterior and posterior portions
of the superior condyle to be more robust and the central por-
tion to be more concave than in Xiphactinus. The condyle for
the operculum is also proportionally longer and not projected
backward so far as in the above. The groove for the preoper-
culum is deep.
Hyomandibular: Length of superior condyle................. 30 mm,
Length of condyle for the operculum ....... (0
Total length of bone (estimated) ........... Lat: ok
Length of groove for the preoperculum (est.). 93 ‘
The preoperculars are preserved in part. They are rather
thick on the anterior border and are probably projected forward
to the angle of the mandible below. Superiorly, there is a long,
slender process of bone extending upward, which is much more
prominent than in the corresponding portion of Xiphactinus.
This portion is somewhat roughly striated externally. The re-
maining portion of the external surface is nearly smooth, ex-
cepting along the superior border, where there are numerous
strie leading out to the border.
The malleolar portion of the palatine is rather thin in a ver-
tical direction, resembling J. anaides somewhat in this respect.
The inferior articular surface has the same general outline as
the posterior condyle of the maxilla. Anterior to this, the bone
is beveled off antero-superiorly, causing the prefrontal articular
surface to be produced forward beyond the inferior. This sur-
face is nearly flat. On the external side of the bone there is a
prominent tuberosity.
300 University Geological Survey of Kansas.
The ethmoid is rather broad posteriorly and pointed anteri-
orly. The prefrontals are heavy masses of bone with large
facets for the superior condyles of the palatines. ‘The frontals
are broad, and each has a prominent ridge extending from near
the median line posteriorly, and curving outward to a point over
the prefrontals. The parasphenoid is triangular, the base of
which is below. The transverse processes are long and not very
slender. ‘The orbit is surrounded by quite a heavy ring of scle- .
rotic bones.
The centra of several vertebree are preserved. They are
deeply concave and without lateral grooves in the anterior re-
gion. The more posterior vertebre are grooved, and have the
ribs attached to small processes of bone which fit into pits on
the sides of the centre.
Ichthyodectes cruentus. Plate L, figs. 8a, 8b, 9, 10a, 10b.
Ichthyodectes cruentus Hay, Amer. Jour. Sci., vol. vi, 1898, pp. 225-228.
This species of /chthyodectes has been recently established by
Professor Hay from a fragmentary maxilla and a portion of a
mandible supposed to belong with it, from the Butte creek re-
gion of western Kansas. Through the kindness of Professor
Hay, I have been allowed to examine his type and compare it
with a specimen of the same species in our collection, which
fortunately is somewhat more complete than his.
The premaxilla is especially remarkable for its great depth
and for the length of the alveolar border. The outer portion of
the bone is very convex from before backward, especially so
just above the alveolar border, where the bone is covered with
minute canals, which give it a somewhat rugose appearance.
The inferior third of the bone receives no support from the
maxilla, thus causing it to be projected downward to a consid-
erable extent. ‘The bone becomes contracted in width supe-
riorly, and bears an articular facet above, probably for the |
ethmoid. The lower portion of the anterior border is much
roughened, probably for igamentous union with its fellow on
the opposite side. Alveoli for nine teeth are found, all of
which are broken away.
STEWART. | Cretaceous Fishes. 301
The maxilla of this species is very similar in general form to
that of J. hamatus, with the exception of being less slender and
more robust. So far as can be seen, the superior border is
nearly straight and much thickened, with a groove along the
external side for the reception of the jugal. This border rounds
off regularly into the internal. The condyle for the palatine
is similar in form to that of J. hamatus, beneath which the bone
is very thick. The anterior condyle is oval in outline and situ-
ated quite a distance from the last. The surface for the pre-
maxilla is bent inward and is much roughened. The anterior
border is very sharp and the lower is separated from the alveolar
border by a prominent notch. This last border is very concave
in the anterior portion and has a prominent hook-like projection
extending downward in front, which is broken away in Professor
Hay’s specimen. The teeth are rounded in cross-section and
non-striate under the microscope. Both the external and inter-
nal surfaces of the bone are finely striated.
EDS REO GE NY IOP UM). Uh Sas ad see ae ge cles eae e ee 50 mm.
ener Gt alveolar DOrdGr. 5.000260 Sees (BQ
Mums: Depth at posterior condyle... 00. 6... ee on
Distance between superior condyles................. me ire
Number of alveoli in one centimeter................. 3
Fragments of the dentaries show these parts to be remarkably
thickened at the symphysis. At this point the alveolar border
is directed slightly upward, forming a slight hook-like pro-
jection, below which the symphysis descends almost vertically.
Dentary: Greatest lateral width at symphysis................ 11 mm.
Number of teeth in one centimeter.................
This specimen, No. 180, was obtained from the Niobrara
Cretaceous of Gove county by Mr. E. P. West some years ago.
&
Ichthyodectes acanthicus ?. Plate LI, figs. 1-11.
Ichthyodectes acanthicus Cope, Proc. Am. Phil. Soc. 1878, p. 177.
This species, as described by Professor Cope, is the smallest
member of the genus, but the description, as a whole, is so very
meager that the species cannot be identified with any degree of
certainty. There are two specimens in our collection which are
much smaller than any of the other species described by Cope,
302 University Geological Survey of Kansas.
but I am far from certain that these belong to J. acanthicus. If
they prove not to be this, | would propose the name of Jchthyo-
dectes parvus for them.
The maxilla is long and slender, with the superior border
moderately curved back of the posterior condyle. This condyle
is slightly convex, and is invaded by a notch from behind. The
surface for the premaxilla is bent inward to a considerable ex-
tent, and has a small tubercular condyle above and in front.
The alveolar border is somewhat concave anteriorly, thin, and
bears a row of teeth which do not seem to decrease much in size
toward the posterior extremity. The teeth are cylindrical in
cross-section, acutely pointed, and directed slightly inward at
the apices. The premaxilla is thin and plate-like, with seven
teeth on the alveolar border.
Premaxilla: Length of alveolar border........................ 18 mm.
Maxilla: Depth at posterior condyle...................... eee Oars
Length of alveolar border (estimated)............... Tile
Number of teeth in one centimeter.................. 8
Besides being smaller in size, the alveolar border does not
have the two convexities found in J. anaides, but is convex in
front and slightly concave in the middle and posterior portions.
The symphyses are but slightly sloping and not much thick-
ened. ‘The coronoid process is poorly developed and the border
back of it slopes gradually to the cotylus. The teeth are cyl-
indrical in cross-section and dirécted shghtly inward. The derm-
articular extends well forward in a long, sword-shaped process
which is very robust below. The antarticular is wedge-shaped
and similar in general form to that of X¢phactinus.
Dentary :. Wenethvofvalveolar border oan. eee eee eee ee 75 mm.
Deoth of symphysis. 2 51ers eh cee ccna einer lene meas Zoos
that: Ccoronoid, process.)3). 0) s ee eee Lee eee SD ene
Number of teeth in one centimeter................. 4.5
Only the anterior portions of the palatines are preserved,
which show the malleolar portions to be projected outward in a
manner similar to that found in J. hamatus, but differs from
this form in having them proportionally broader and not so
deep. The articular facet for the prefrontal is the smaller of
the two and is nearly flat, while the one for the posterior con-
STEWART. | Cretaceous Fishes. 303
dyle of the maxilla is slightly concave and much broader than
the condyle articulating with it, thus forming a rather loose
union.
There are numerous vertebrae present which show them to
be two-grooyved, and, with the exception of the difference in
size, very similar to those of J. anaides. The pectoral fin is
represented in a somewhat fragmentary condition, but enough
of the rays are present to show that the upper ones are sword-
like and but slightly less curved than in NXiphactinus. The
pelvic actinosts are very similar in many respects to those of
the genus just mentioned. The two halves are strongly sutured
together in the median line, where the bones are massive for
the support of the pelvic fin. The facets are obliquely set and
are four in number on each side. The upper and lower of these
are large and flat, while the two median ones are round and
somewhat tubercular. Extending forward, both above and be-
low, there are ridges of bone which are not so prominent as in
Xiphactinus, which may have formed bars of bone on each side.
External to these there are wing-like processes.
Ichthyodectes ctenodon. Plate XLIX, figs. 5-7; plate LI, figs. 12, 13.
Ichthyodectes ctenodon Cope, Proc. Am. Phil. Soc. 1870, p. 536.
There is one specimen, consisting of the mandibles, a portion
of a maxilla, and other bones, which do not seem to differ ma-
terially from those of the specimen described and figured by
Professor Cope. It evidently does not belong to any of the other
American species, and for the present at least I will leave it here.
The dentary is about the same size as that of J. anaides, de-
scribed above, but differs from this in the absence of the pos-
terior convexity of the alveolar border. This border unites
with the symphysis at an angle of about 75 deg., which is 10
deg. more than Cope describes of his specimen. ‘There seems
to be some discrepancy between Cope’s description and figure,
for in the latter the angle seems to be about the same as in our
specimen, which evidently shows that Cope was wrong in one of
the two. The teeth are large anteriorly but somewhat smaller
posteriorly, round in section, non-striate, and directed inward.
304 University Geological Survey of Kansas.
Cope describes his specimen with the posterior teeth slightly
larger than the anterior, which seems rather remarkable for
this family. The posterior extremity of the maxilla shows this
portion to be upturned, as Cope has indicated in his figure.”
The quadrate has its condyle twisted forward until the face
is almost in line with the anterior border. The upper border is
straight and the groove for the symplectic narrow. The cerato-
hyal is concave posteriorly, convex and contracted in width an-
teriorly.
ee eng thiofalveolar borden ee eee ee a en oe 101 mm.
Length of symphysis... 2... ...0- ed RRGANG NS hai SIR AS 8
Keng thiof crowns amtenion cee yee ieee pape
Number of teeth iMnvone Centimeter nes. ieee 3
This specimen was obtained from the Niobrara Cretaceous,
five miles northeast of Russell Springs, Logan county, Kansas,
by the geological expedition of the summer of 1898.
GILLICUS. |
Hay, Amer. Jour. Sci., vol. VI (1898), p. 320.
The genus Guillicus has recently been proposed by Professor
Hay, to include I[chthyodectes arcuatus Cope and I. polymicrodus
Crook, which differ materially from the other species of Ichthyo-
dectes, especially in the form of the teeth and tooth-bearing
elements. Before entering into the description of this interest-
ing form, it will be well to give a short reswme of the discus-
sion which has taken place over these two species, as is also
done at some length by Professor Hay.”
In 1875 Professor Cope first described his Portheus arcuatus
from some very imperfect remains from the yellow chalk of the
Solomon river, Kansas. It is difficult to say just why Professor
Cope first referred this to ‘Xiphactinus (Portheus), as the dental
and other characters are entirely different from that genus.
Cope evidently discovered his mistake soon afterward, as he
refers it to [chthyodectes in 1877.° Nothing more was done with
this species until 1892, when Doctor Crook published his paper
entitled ‘‘ Uber einige fossile Knochenfische aus der mitteleren
60. Cret. Vert. West, pl. XLVI, fig. 1. 62. Proc. Am. Phil. Soc., XVII, p. 177.
61. l. c., pp. 228, 229.
STEWART. | Cretaceous Fishes. 305
Kreide von Kansas,’’ in which he described his Jchthyodectes
polymicrodus. Shortly after this Professor Cope reviewed this
paper in the American Naturalist,” and claimed that Crook’s
species is the same as the one already described by him as J.
arcuatus. He further states that if it had not been for certain
conditions, figures of this species would have been published, a
statement which I think means that more characteristic parts
would have been figured and described.
Cope was evidently in doubt about the specimen figured as
Ichthyodectes (Portheus) arcuatus, as on plate XLVII of the
‘* Cretaceous Vertebrata,’’ he refers to it as ? Portheus arcuatus,
while on the opposite page, in the explanation of the plate, it is
Portheus ? arcuatus. Furthermore, on page 220 B of the same
work, he refers to these figures of an unknown species of
saurodont. Altogether I am inclined to think that Professor
Cope’s description and figures are for entirely different speci-
mens, but the description, so far as it goes, does not seem to
differ from the description and figures of J. polymicrodus of
Crook.
-When we take into consideration the fact that Professor Cope
was able to recognize his species in Crook’s description, and
also that this form is so abundant in the chalk of western Kan-
sas that it could hardly be missed by a party collecting fossil
fishes for any length of time in that locality, I do not think we
should hesitate to regard Gillicus (Ichthyodectes) polymicrodus
Crook as a synonym of Gillicus arcuatus Cope. The characters
in which Gillicus Hay differs from Jchthyodectes Cope are
enumerated as follows by Professor Hay :" ‘‘ While these forms
can by no means belong to Cope’s Portheus (Xiphactinus of
Leidy), they can hardly belong to the genus Ichthyodectes. In
the latter genus the maxilla is long, nearly equal to the distance
from the tip of the vomer to the occipital condyle. The gape of
the mouth must therefore have been large. In Doctor Crook’s
species and related forms the maxilla is short, between one-half
and two-thirds the distance referred to above; hence the gape of
63. XXVI, p. 942.
64. Amer. Jour. Sci., vol. VI, p. 229,
306 University Geological Survey of Kansas.
the mouth must have been rather small. The maxille of typ-
ical species of [chthyodectes are nearly straight along the tooth
line, or sinuous, or, in J. hamatus, strongly concave. In J.
polymicrodus the tooth line is strongly convex, except just be-
hind the palatine condyle. The teeth of I. polymicrodus are
numerous and feeble; in the other species, strong and in small
number.’’
Gillicus arcuatus. Plates LII, LIII, and LIV.
Portheus arcuatus Cope, Cret. Vert. West., pp. 204, 274, pl. xivit, figs. 7-9.
Ichthyodectes arcuatus Cope, Proc. Am. Phil. Soc. 1878, p. 177; Amer.
Nat., vol. xvi, p. 942.
Ichthyodectes polymicrodus Crook, Paleonto., 1892, p. 112, pl. xvt.
Ichthyodectes arcuatus Cope, and Ichthyodectes polymicrodus Crook
and Hay, Amer. Jour. Sci., vol. vi, p. 228.
This species is represented in the museum by the remains of
three individuals with the skull in good condition, and frag-
ments of several more.
The maxilla is a broad and thin bone, with the greater part
of the alveolar border very convex. Just back of the surface
for the premaxilla this border is directed sharply upward, form-
ing a rather deep concavity near the anterior end, back of which
it is gently curved. This border gives support to a single row
of minute teeth, which are bent slightly inward at the apices.
Doctor Crook says that twenty-four of these occur to the centi-
meter, in which I am inclined to think that he is mistaken, unless
he intended it to apply to the posterior end, where the number
will probably reach that many, but at the anterior extremity I
have been unable to make out more than from seventeen to
eighteen. The surface for the premaxilla is bent inward to a
considerable extent and is separated from the alveolar border
by a slight notch inferiorly. The condyle for the palatine is el-
liptical in outline, slightly convex, and elevated slightly above
the rest of the bone. Professor Hay was able to distinguish
some differences in the form of these condyles, as he says :® ‘‘ In
looking over my specimens it seems to me that I can observe
characters that indicate two species. In one maxilla I find that
65. It is doubtful if these figures refer to this species.
66, Amer. Jour. Sci., VI, p. 229.
STEWART. | Cretaceous Fishes. 307
the posterior or palatine condyle is comparatively short, and
has, in its hinder border, a distinct notch ; in other maxille the
condyle is longer and, apparently, without the posterior notch.
The distance between the condyles appears to be greater in
some cases than in others.’’
[have been unable to discover any such difference in any of
our specimens, but, if this difference does occur, I think that it
could not be called a specific character, as there is likely to be
almost as much individual variation in Gillicus and Ichthyodectes
as in Xiphactinus, in which the variation is remarkably great.
The superior border of the bone, back of the condyle, is slightly
concaye and presents a slight shelf, which probably accommo-
dates the jugal. Unfortunately the premaxilla is preserved with
none of the specimens, nor has it been found with any of the
specimens described by other authors. It was, no doubt, a
very small and light bone, thus easily washed away when the
soft parts disintegrated.
Maxilla: Length of alveolar border (estimated)............... 97 mm.
SPCGiM Ub POSUEKION COMCYIG © ook. ack a bielece.d sie e wise we 24 %
The dentaries are remarkable for their short length and great
depth. The symphysis is very deep, almost vertically directed,
and smooth. Just back of it on the internal side there is a
broad, shallow and somewhat elliptical-shaped pit, similar tothe
one found in Saurodon and Saurocephalus. The alveolar border
presents a slight convexity near the anterior end, back of which
it is slightly concave. The teeth are all very small and form a
fringe on the edge of the jaw. The coronoid process is not
developed at al], and the lower border is very sharp. Pos-
teriorly the bone descends very abruptly to the symphysis.
eH Y VOD Ai BYMADMYOIS.. codes eb cn cidenecaesvnavoekoas 45 mm.
Rett Ie AVERT MOTO i. Sods OS apes eee away OO on
Length of mandible from cotylus................... By
The ceratohyal is preserved in one specimen. Itis very thin,
much contracted at the center, and expanded at the ends, the
posterior of which is concave and the anterior convex.
The palato-quadrate arch is represented by the palatine,
mesopterygoid, pterygoid, and quadrate. The palatine consists
308 University Geological Survey of Kansas.
of a broad, thin posterior plate and the malleolar portion.
This latter is somewhat deeper than in Jchthyodectes acanthicus,
and does not have a tuberosity on the external side. The two
portions of the bone are separated by a constricted neck.
Nothing more can be said of the mesopterygoid than that it is
a very thin bone.
The quadrate is very deep, as might be expected from the
corresponding depth of the mandible. The upper portion is
very thin and has a slight groove on the internal side for the
symplectic. The condyle is very convex and directed well for-
ward. ‘The upper border seems to be nearly straight and prob-
ably unites with the metapterygoid in a manner similar to that
found.in Niphactinus. The preoperculum is very thin, broad
below, and narrow above.
The ethmoid is very similar to this bone in \iphactinus and
Ichthyodectes, being formed into a beak anteriorly, and thin, and
united with the frontals by a dentate suture posteriorly. The
frontals are separated by a well-marked non-dentate suture,
which is shown throughout its length in one of the specimens.
Over the orbital cavities the bones are somewhat thickened, and
evidently gave attachment to a chain of supraorbital bones, as
in X7iphactinus. Posteriorly they unite with the postfrontals,
parietals, and probably the pterotics.
As there has been some doubt as to the exact position of the
parietals and pterotics, | quote the following paragraph from
Professor Hay,” which seems to me to be the correct explana-
tion of these parts: :
‘‘Crook has interpreted the bones of the posterior upper re-
gion of the skull of his J. polymicrodus as he has those of Por-
theus (Xiphactinus) ; but, as in the case of the latter, | am
compelled to differ with him. However, many of the sutures
are very difficult to demonstrate. According to Crook, the su-
praoccipital is greatly expanded in front, while the parietals he
laterad to the epiotics. It seems to me that the supposed ex-
panded portion of the supraoccipital is really the area belong-
ing to the parietals. There is a very distinct fold running along
67. Amer. Jour. Sci., vol. VI, p. 230.
SrTewakrt. | Cretaceous Fishes. ~ 809
the hinder border of the expansion, and this, continuing up to
the midline, has the appearance of a suture. In the crushed
specimen before me there is, along the middle line of the ex-
pansion, a break in the bone, but whether due to a fracture or
to the parting of the bones along a suture I cannot determine.
Possibly the parietals were coossified along the midline.”’
As to Crook’s parietal, I cannot convince myself that there
is any suture cutting off the area assigned to it from that as-
signed to the pterotic. J regard as pterotic the whole area ex-
tending from the posterior external angle of the skull to the
lateral expansion considered by Crook as supraoccipital.™
In one of the specimens before me the portion under consid-
eration above is so well marked that there can be no more doubt
that the expanded portion called supraoccipital by Crook is a
separate and distinct bone, which is not separated by suture in the
middle line. The portion called parietal by the same author
often has numerous cracks running through it, which will easily
lead one to believe that they are sutures.
The prefrontals are small, triangular-shaped bones which le
below the frontals and give attachment to the palatine, as in
NXiphactinus. The postfrontals form prominent processes, and
are somewhat irregular in form. They form the posterior
boundaries of the orbits, and present grooves externally which
support about one-fourth of the upper faces of the hyomandibu-
lars.
The parasphenoid is completely preserved in three specimens.
It is more slender than in either Niphactinus or Ichthyodectes,
and is quadrilateral in cross-section, instead of triangular, as in
the forms mentioned. The transverse processes seem to be well
developed, just back of which there are very thin lateral proc-
esses which extend upward to the prootics, just beneath which
the bone is strongly bent at an angle of about 55 deg., thus giv-
ing a strong upward direction to the anterior portion of the skull.
The sclerotic ring is unusually large for the size of the speci-
men, the distance across the orbit being 37 mm., a distance
68. See Crook’s fig. 5, pl. XVI, Sq. and Pa.
310 University Geological Survey of Kansas.
which is even greater than in \iphactenus. The bones are very
thin.
One specimen has fifty-seven vertebre preserved, and there
are probably several more missing. ‘So far as can be deter-
mined, this specimen was at least four feet in length, and,
judging from the skull, it does not seem to be larger than the
other specimens.
SAURODONTIDA.
The characters that separate this family from the [chthyodec-
tide are found in the presence of a predentary in the mandible
and in the form and manner of succession of the teeth. In
other respects the skull is very similar to that of the Jchthyo-
dectide, except that it is likely that Saurodon possesses a tooth-
bearing element not found in this family. The predentary is a
long, triangular element, pointed at the extremity, and was used
as a weapon of offense. Contrary to what would be expected,
this bone is not paired, and is also edentulous in both genera
of this family, Sawrodon’and Saurocephalus. Doctor Crook® has
removed one of these genera, Sawrocephalus, to the family Pachy-
cormidx, the reasons for which are:
“Das Dentale hat andere Proportionen als dasjenige von /chthyo-
dectes und Portheus | Xiphactinus | indem es viel niedriger und gegen
die zahntragende Oberfliche dicker wird. Die maxilla ist mehr dick
als tief, die Preemaxilla dreieckig und lang, die langste Seite als zahn-
tragender Rand ausgebildet. Sie ist so ganzlich verschieden von
denjenigen der anderen Glieder dieser Familie, dass dies Merkmal
allein geniigt, um Sauvrocephalus einer anderen Gruppe zuzuweisen.
Diese Annahme wird noch mehr bestatigt durch den Charakter der
Maxilla und des Dentale, die Foramina und die Art und Weise der
Aufeinanderfolge der Zahn. Auf Grund der Gleichartigkeit der
Zahnea und der auserordentlichen Athnlichkeit der Premaxilla mit
derjenigen von Protosphyrena diirfen wir Saurocephalus bis auf
weiters in die Familie von Protosphyrenide einreihen.”
Concerning the above, I would say that I have been unable
to recognize the great similarity between the premaxille of the
69. Paleontographica, 1892, p. 120.
STEWART. | Cretaceous Fishes. 311
two genera from the specimens in the museum. Furthermore,
if Saurocephalus were remoyed to another family, Sawrodon
would have to follow, as the two genera are too closely allied
to be far separated from each other, and the absence of the long
rostrum, paired and toothed predentary, and the presence of
more than one row of teeth on the maxille and mandibles would
preclude any close relationship with the family to which he re-
fers it. I think the two genera should be placed in a distinct
family, as has been done above.
Revised synopsis of the Saurodontide :
Foramina below the alveolar border internally.
Teeth with short compressed crowns ..............-.2.-005: Saurocephalus.
Deep notches below the alveolar border internally.
OS be With SUDCYINGTIC CYOWDS . . ...c.06 cle viele s cone Nee oS Saurodon.
SAURODON.
Hays, Trans. Am. Phil. Soc., 1830, p. 476.
Cope, Proc. Acad. Nat. Sci. Phil., 1873, p. 339.
In the year 1830, Doctor Hays“ described the genus Saurodon
from a portion of a skull and jaws from the Marl of New
Jersey. He also examined the type specimen of Sauwrocephalus,
described by Doctor Harlan” six years before, and decided that
the two genera were synonymous, and, as Doctor Harlan’s genus
was founded upon erroneous characters, the name Sawrodon
should take precedence over it. In 1856 Doctor Leidy ” rede-
scribed both of the above specimens and decided that the name
Saurodon should be abandoned and Saurocephalus used instead.
Nothing further was done with either of these forms until 1873,
when Professor Cope added the genus Daptinus, which he later
recognized as a synonym of Saurodon.
The exact date of Professor Cope’s retraction I have been un-
able to exactly determine, but it was probably not until after
1878, as during this year Mr. E. T. Newton” described a fish
from the Lower Chalk of Dover and provisionally placed it in
Daptinus. A little later in the same year, Mr. Newton pub-
lished another paper“ in which he carefully goes over the
70. Trans. Am. Phil. Soc., vol. III, p. 471. 73. Trans. Am. Phil. Soc., vol. XI, p. 91.
71. Jour. Acad, Nat. Sci. Phil., vol. III, p. 331. 74. 1. c., No. 136, p. 786.
72. Quart. Jour. Geol. Soc., 1878, No. 135, p. 439.
312 University Geological Survey of Kansas.
ground, and finally concluded, as Doctor Leidy had already
done, that the name Saurodon should be no longer used.
It has since been shown that there are two distinct genera,
the differences being found mainly in the more slender mandi-
ble and the occurrence of deep notches instead of foramina at
the bases of the crowns of the teeth in Saurodon. There is also
some difference in the shape of the teeth in the two forms.
Below is given a list of the known American species :
Saurodon leanus Hays, Marl, New Jersey.
Saurodon phlebotomus Cope, Niobrara Cretaceous, western Kansas.
Saurodon broadheadi Stewart, Niobrara Cretaceous, western Kansas.
Saurodon xiphirostris Stewart, Niobrara Cretaceous, western Kansas.
Saurodon ferox Stewart, Niobrara Cretaceous, western Kansas.
Saurodon phlebotomus. Plate LVII, figs. 4 and 5.
Daptinus phlebotomus Cope, Cret. Vert. West, p. 213.
Saurocephalus phlebotomus Cope, Proc. Am. Phil. Soc. 1870, p. 530;
Hayden’s Ann. Rep. U.S. Geol. Surv. 1871, p. 416.
There are remains of several specimens of this species, in-
cluding the jaws, vertebre, and portion of one skull, in the mu-
seum.
The maxilla is more slender than in S. broadheadi, and the
surface for the premaxilla is bent inward very strongly, al-
though this may be due to distortion. The alveolar border is
nearly straight, and has alveoli for twenty-nine subcylindric
teeth, which have the characteristic notches for nutrient ves-
sels at the base of each. The posterior extremity of the border
is edentulous, curved slightly upward. The condyle for the
palatine is not elevated as in Xiphactinus, and is flatter than in
Saurocephalus. The internal side of this condyle is invaded by
a slight notch, which is absent in S. broadheadi and S. ferox.
The anterior condyle is broken away, but it seems to have been
elevated upon a slight pedestal, and was probably very small.
The upper portion of the bone is striated ; the lower portion, just
above the alveolar border, is covered with numerous punctations.
The dentary is slender, and the symphysis is not so straight
as in Saurocephalus dentatus, but is slightly convex in front, with-
out the prominent bony tubercles which are so pronounced on
Srewart. | Cretaceous Fishes. 313
the internal side of the symphysis of the form just mentioned.
Back of the symphysis there is a slight concavity in the alveolar
border, just over the large shallow pit at this point. This pit,
which is probably for the Mento-Meckelian ossicle, is much
smaller than in Saurocephalus and the swelling above it is less
pronounced. The alveolar border supports thirty-six teeth, the
anterior ones’ of which are very small and the posterior ones
about twice the size of those on the maxilla. The crowns are
subcylindric, knife-like, and extend nearly to the coronoid proc-
ess, which is rather weak, and not curved upward and outward
as in Xiphactinus. The lower border of the bone is very thin,
with numerous fine strive extending obliquely upward and for-
ward on both the external and internal sides.
The articular portion is divided into the dermarticular and
antarticular portions. Owing to the crushed condition of the
specimens these portions cannot be made out with certainty,
but seem to be very similar to the same parts of S. ferox, de-
scribed below. The predentary is not preserved in any of the
specimens.
Da MITRE Bb Ne ds, AI nh kd ee eaN Roe dds 96.0 mm.
BPephlh BE. CONGR, |. Soc. cease 8 DA ee Cease eae Zo **
Penpi OF palstme CONGYIG). . 6 oe fees os els dees Lisa
Number of teeth in one centimeter, 5.
La AAS i 1 ees ea 4.0 mm.
ESNCA CARR ESE ORISWUE si hike abla es tes vats Oe adhe ok ewe
Depth of bone at palatine pane Ree ett gS as cases, hs BOO
Mandible: Length of bone from cotylus.................... 116.0) *
POS PUNCOL AVEO VOI, 205: 60s 5d aree em BY Ears coe 26,00 £
DIB PER BL COTONOUS BNLIG. 3. ee wie cw ew ons ene 4150; **
Length of alveolar border...................... SAA
Number of teeth in one centimeter, 4.
Saurodon broadheadi.
Daptinus broadheadi Stewart, Kans. Univ. Quart., vol. vir, pp. 24, 25.
This species was established upon the left maxilla and one of
predentaries found in Wallace county, Kansas, collected by Mr.
Geo. W. Cooper. The catalogue number of the specimen is 212.
The maxilla is less elongate and ends more abruptly than in Sau-
rocephalus. The anterior border slopes forward more obliquely
than in Xiphactinus. The premaxillary surface is continuous
21—vi
314 University Geological Survey of Kansas.
with the outer surface of the maxilla and is provided with numer-
ous small protuberances, which probably fit into corresponding
depressions in the premaxilla. It is seen from the above that the
premaxilla is not so immovably fixed as in Xiphactinus, where
this bone fits into a deep depression on the maxilla and has a
thin lamina of bone extending forward nearly to the anterior
extremity, which gives it additional support. The ramus is
thin above and thickens but slightly at the alveolar border.
The bone does not materially thicken below the palatine condyle
as in AXiphactinus, and the anterior condyle is situated much
nearer to it than in this form. The palatine condyle is elon-
gated, elliptical, nearly flat, and does not have the internal
notch found in S. phlebotomus. The teeth are closely set, with
compressed, knife-like crowns and smooth enamel surface, ap-
pearing very slightly striated under the microscope. Alveoli
for thirty-one are found, below each of which there is a deep
notch, characteristic of this genus.
A predentary was found on the same slab with the above
which has the form of the predentary of Saurocephalus. It is
short and. triangular, the posterior surface of which is very
rough for cartilage, binding it to the symphysis. The upper bor-
der is edentulous, the lower thin and sharp, and the two meet
at quite an acute angle in front.
Maxillaewength (estimated) ja airy he tial i aune aed aha 122.0 mm.
Depthiat cenbeny as oys con xe ay cieer eae cnae ont meng a ale St Ona
Depth at palatine condylers shen.) sae eee J aca AAO atetts
Number of teeth in one centimeter, 3.5.
Predentary , Weneth(estimated)\e see. eatin ee ee 30.0 mm.
Depth atisyimplaysisy aye woe kee ca tencen arta 285)
Saurodon xiphirostris. Plate LV.
Saurodon xiphirostris Stewart, Kans. Univ. Quart., vol. vi, p. 178.
This specimen consists of a skull crushed obliquely, the centra
of several vertebre, and also a portion of a shoulder girdle in a
very bad state of preservation. The specimen is of nearly the
same size as that of the type of Saurodon (Daptinus) broadheadi”
described by myself.
75. Kans. Univ. Quart., vol. VII, pp. 21-29.
Srewarrt | Cretaceous Fishes. 315
The maxilla is short and deep, the depth not being as great
as in S. broadheadi. The alveolar border is nearly straight and
has alveoli for about thirty-one teeth, which are about the size
of those described in the above species. The posterior extrem-
ity cannot be examined, as there is a suborbital bone covering
this portion on each side of the skull, but it is probably very
similar to that of the figures of Sawrodon ferox described below.
The superior border is sharp, and gives attachment to some
bone, probably a suborbital or jugal. The palatine condyle
seems to be very similar to that of S. broadheadi already de-
scribed. The bone, just above the alveolar border, presents a
somewhat shagreened appearance. [Farther than this, there
seem to be no characteristic markings upon the external sur-
face. As the maxilla is firmly attached to the skull, the inter-
nal surface cannot be examined.
The premaxilla is plate-like, and nearly twice as aes as
broad. The superior border is irregular and presents no con-
dyle at this point. The external markings are very similar to
those found in Saurocephalus dentatus, and the bone is directed
more obliquely backward than in that form. The anterior bor-
der is directed sharply inward, giving the external surface of
the bone a very convex appearance. ‘This border is very rugose,
probably for ligamentous union with its fellow of the opposite
side. The alveolar border is very convex, and has alveoli for
twelve teeth, which are of about the same size as those on the
maxilla.
Maxillary: Length of alveolar border (estimated)............. 82 mm.
Depth at palatine condyle ......665.......0ci0e.s BB
Number of teeth in one centimeter, 4.
PUES = COPOALORE GPE, 6 occ cs cakes bce. cone ab ened 50 mm.
SPEEDS IDLO GE, xd. olive Ais sey Os 0 vw obo DENS ot ee
beng Of anterior DOGG? 2s oi. cae se 6 me, S
In the mandible is found one of the chief characters that
separate this genus from Saurocephalus; instead of the upper
and lower jaws terminating at about the same vertical plane as
in the other member of the Saurodontidx, the mandible projects
fully an inch beyond this point. The dentary is long and
slender throughout; in Sauwrocephalus it is short and deep.
316 University Geological Survey of Kansas.
This difference is well illustrated by comparing the types of
Saurocephalus dentatus and Saurodon ferox; the maxilla of the
first is considerably longer than that of the second, but with
the dentaries the reverse is the case. Only a small portion of
the dentary can be seen, as most of the external and superior
portions are hidden by the overlying maxille. The bones are
irregular and shallow at the symphysis and seem to have given
strong attachment for the predentary. The lower border is
thin and sharp. Only twenty-seven millimeters of the alveolar
border can be seen in the specimen, upon which the teeth are
small and twelve innumber. At the base of each tooth is found
the deep notch for the nutrient vessels, so characteristic of this
genus. As the articular portion does not seem to differ materi-
ally from that of S. ferox, its description may be deferred.
Contrary to anticipations, there is but one predentary, as is
proven by the discovery of all of the parts in place. It is long
and slender, triangular in outline, with a broad, elliptical articu-
lar surface at the posterior extremity. When this element was
first made known” in this genus, I was under the impression
that it was paired, which is not the case. This slender pro-
jection was no doubt used as a weapon of offense, analogous to
the rostrum of Protosphyrena. In connection with the descrip-
tion of Saurodon broadheadi I figured a predentary of an en-
tirely different form from the above, which was found on the same
slab with the maxilla described; the form is the same as that
found in Saurocephalus. Whether the two bones belonged to
the same individual or not only future discoveries can deter-
mine. After carefully comparing the type of S. dentatus with
that of the species under consideration and S. ferox, I am con-
vinced that there is but one predentary in the mandible of this
form, as one would expect from the great similarity of the two
genera.
Mandible: engthitromcotyloidicavity nee oe eee 155 mm.
Depth at symphysis 2:f ee eet tee may. | OF
Number of teeth in one centimeter, 4.5.
‘Predentary : Length (estimated ey. etree eee ee 73 mm.
Depth of symphyseal surface.................... 23. wes
Width of symphyseal suriacenn. ae ae eae NPA eG
76. Kans. Univ. Quart., vol, VII. p, 24. Ue dle @ag Tolle JU,
STewaRrt. | Cretaceous Fishes. 317
The ethmoid is broad and flat posteriorly, becoming thickened
and pointed at the anterior extremity. ‘The lower surface can-
not be seen, but it probably is not materially different from that
of Ichthyodectes. In Prof. E. T. Newton’s description of S. in-
termedius“ he says, concerning this part: ‘‘Anterior to the
frontals, upon the upper surface of the skull, there are two
bones (fig. 2) separated by a median longitudinal suture ;
towards the front of these an osseous band passes across at
right angles, obliterating the suture.’’ In our skull I am un-
able to detect any indication of a suture at this point such as is
shown in the figure referred to above. I have also examined
all of the specimens of NXiphactinus and IJchthyodectes in the
museum, and find no trace of such a suture in any of them.
It seems probable to me that the skull described by Mr. Newton
was of a younger individual than are any of ours.
The frontals are broad, flat bones extending from the ethmoid,
with which they are united by a squamose suture, back to the
parietals. lLaterally, they form the superior borders of the
orbits. In the median line they are separated by a suture.
The bones are probably in contact with the supraoccipital, but,
owing to the crushed condition of this region, this point cannot
be definitely determined.
The parietals (?) are small elements in contact with the pter-
otics and epiotics posteriorly. They are probably very similar
to this portion of Xiphactinus and Jchthodectes, although this re-
gion of the skull is so crushed as to render the determination of
this portion somewhat difficult. There seems to be a faint su-
ture between the pterotic and sphenotic. The epiotic does not
seem to be produced as far backward in this species as in S. in-
termedius, figured and described by Mr. Newton.” They are
heavy projections of bone, and form the inner lateral processes
of the skull as in other members of the Saurodontidx and Ichthyo-
dectide. Mr. Newton* seems somewhat in doubt about the
bones in this region and is unwilling to accept the bone called
parietal by Professor Cope, stating, for his reason, that the lines
78. Quart. Journ. Geol. Soc., vol. XXXIV, No. 135, p. 444, pl. XTX.
We, 136.5 ps 444, pl. XXIV. 80. J. c., pp. 444, 445,
318 University Geological Survey of Kansas.
indicative of the direction of growth were from the extreme pos-
terior angle of the skull, instead of from the anterior portion, as
we would expect if this bone were the parietal. In skulls of
Xiphactinus, Ichthyodectes, and Gillicus, before me, I find the
same condition, which seems to confirm Mr. Newton’s idea.
Recently Professor Hay*' has shown wherein Professor Cope was
wrong in his identification of this part of Yiphactinus and Ich-
thyodectes, and, as the top of the skull as a whole seems to be
very similar to that of the two genera mentioned, I have no
doubt but that the same explanation will apply to Sauwrodon.
The pterotics are large bones and seem to be very dense in
structure.
The supraoccipital is very much crushed and partially broken
away, but enough remains to show that the bone was raised
into quite a prominent crest. It extends backward beyond the
points of the epiotics, a condition different from that described
by Newton.” It probably does not join the frontals in front, the
parietals intervening.
The orbit is somewhat smaller than in Jchthyodectes and is
surrounded by a thin ring of sclerotic bones similar to that
found in the genus just mentioned. Just in front of the orbit
there is a small triangular bone attached to the frontal above,
which I take to be a preorbital. The same bone is figured by
Newton* but not named or described by the author. Justin
front of this there is a long slender bar of bone, which seems
to articulate somewhere in the palatine region. On one side
the anterior end is crushed down to near the posterior condyle
of the maxillary, but on the other side it fits in just back of the
superior condyle of the palatine, and as a palatine of another
specimen shows a sutural surface at this point, I think it not
unlikely that this is the correct position of the bone. This
may be the bone that Newton™ figured as a ‘‘nasal bone,’’ (?)
although it is of an entirely different shape from that shown in
the cut of his specimen. The bone Newton calls ‘‘jugal’’ (?)
I am inclined to think is one of the suborbital bones, as found
81. Zool. Bull., vol. II, p. 28. 83. 1. c., p. 444.
82. l.c., p. 444. 84. l. c., pl. XXXIV.
iy Secret aaa
i
y
3
E
Lal
STEWART. | Cretaceous Fishes. 319
in Xiphactinus, especially as there seems to be a suture indi-
cated between this and a bone just above which articulates with
the jugal ( probably a suborbital ) above.
Owing to the crushed condition, the prefrontals are almost
entirely covered by the ethmoid and frontal. The description
of the palatine will be given with the next species. Parts of
the operculum and preoperculum are present. The first is a
broad flat plate of bone which articulates with the hyomandibu-
lar in a manner similar to that found in Viphactinus and Ich-
thyodectes. The anterior border of the preoperculum is deeply
concave, the anterior inferior extremity reaching forward to the
angle of the mandible. The hyomandibular of this species is
not visible.
The vertebrie are deeply concave, with deep grooves closely
situated above for the neural arches. The ribs articulate with
small ossicles set into pits on the side of the centrum. Just
above these ossicles there is a deep pit on each side.
A part of the shoulder girdle, including a fragmentary fin,
is present. The fin seems rather small.
The skull as a whole is especially remarkable for the extreme
length of the mandible, and also the long predentary in front.
This portion probably had a dermal covering similar to that
covering the sword-fishes’ sword, and was no doubt used as a
weapon of offense. In an animal with such a weapon as this
we might expect to find powerful fins, but this is not the case
with this species. In other respects the skull does not mate-
rially differ, excepting in details, from the skull of other mem-
bers of the Saurodontide and Ichthyodectide.
The type of this species was found in Gove county, Kansas,
by Mr. H. M. McDowell, who presented the specimen to the
museum.
Saurodon ferox. Plates LVI and LVII.
Saurodon ferox Stewart, Kans. Univ. Quart., vol. vu, p. 183.
This species is represented by the jaws, including the pre-
dentary, and other disarticulated bones and vertebre.
The maxillary is larger than the one just described. The
320 University Geological Survey of Kansas.
posterior condyles above are somewhat elliptical in outline and
but slightly convex from before backwards. Just anterior to
this there is a large protuberance, which may support a condyle
above, and on the external side of this there is a small facet,
which probably gives articulation with the ethmoid. The sur-
face for the premaxillary is very irregular and is directed in-
ward, becoming thinner toward the anterior border, which is
sharp. The superior border is strongly concave and sharp, and
presents a sutural surface on the external side, probably for a
jugal. The alveolar border is convex and has alveoli for forty
or forty-one teeth, which are non-striate and knife-like. Hach
tooth has the characteristic notch at the base. The posterior
extremity at the bone is very shallow and turned slightly up-
ward. Aside from the shagreened surface of the bone above
the alveolar border the external surface has no characteristic
markings.
The premaxillary is very similar to that of the species de-
scribed above, except that there are ten instead of twelve teeth.
On the internal side, the bone is beveled off toward the pos-
terior border in order to fit the surface for its reception on the
maxillary. |
Maxillary: Length of alveolar border........................ 115 mm.
Depth at condyle for palatine ................... Aten
Greatest Teneth of bone. ie ee eae ees 1 Sie
Number of teeth in one centimeter, 3.5.
Premaxallary,:) Depths ihe ee ee eee ae ee . 60 mm.
1 Tey a 24] 4 ONSEN APEC Vere alee AU MRUT RL eS te fl uh By UE
The dentary is elongate and slender. The alveolar border is
slightly incurved at the symphysis and supports forty-six teeth,
similar in form but about twice as large as those upon the max-
ila. Just back of the last tooth there is a slight coronoid
process, somewhat similar to that found in Xiphactinus. The
symphysis is very similar to that found in the last species de-
scribed, and has a long, slender pit just back of it on the internal
side. It is more elliptical than the corresponding pit in Sauro-
cephalus. The lower border of the bone is sharp.
The dermarticular supports only a small portion of the cotylus,
in the lower portion of the cavity. It sends along, sword-shaped
F
STEWART. | Cretaceous Fishes. 321
process forward internally, but does not encroach much upon
the dentary externally. Posteriorly it sends a lamina of bone
backward, which I think would be well named the cotyloid
process, as in most of the genera of this family it is the only
portion of this bone which articulates directly with the quad-
rate. Just beneath the cotyloid process there is a prominent
angle.
The antarticular is a small element not extending forward be-
yond the sixth posterior tooth, and is fitted into a groove in the
dermarticular. It supports nearly the whole of the cotylus,
which is somewhat elliptical and concave from above downward.
The predentary is not so elongated, but is slightly deeper than
in the species described above.
Maaatiole: Lenein of alveolar border .............:......-.+. 149 mm.
Deneth from cotyloid Cavity... 6. 65. oe Se ce nes 174 *
RECOIL AL SVT DMV SIE. Orica ke cele de dees soc ok i ees
BSF ib SLOROIE Sola oc els ek Ga liods SE cao vel Xen eee
Number of teeth in one centimeter, 3.
et Oe 0) pais Bares? epee si Bec 55 mm.
Length of symphyseal surface .................. 24 *
The quadrate is a broad, fan-shaped expansion. The condyle
is elliptical in outline and convex. Extending upward from
the condyle on the external side there is a ridge, which ends
above in a deep notch, which accommodates a portion of the
symplectic. This groove continues downward on the internal
side for more than one-half the depth of the bone. The poste-
rior border has a very slight groove, and extends upward the
whole extent of the symplectic. The superior border probably
articulates with the pterygoid and metapterygoid, as in Xiphac-
tinus. ‘The anterior borderissharp. Both the external and in-
ternal surfaces are covered with minute striw radiating upward
from the condyle. The symplectic is a long, slender element.
The upper end presents an articular surface sim/lar to that
found on the superior border of the quadrate.
The whole of the palatine is preserved. It is an irregular-
shaped bone, presenting a ragged sutural surface above and be-
low for the pterygoid and mesopterygoid. The bone is especially
remarkable for the great depth of the malleolar portion, being
322 University Geological Survey of Kansas.
nearly half as deep as the corresponding part in Xzphactinus.
The superior articular surface is small and oval in outline,
while the lower is larger and more elliptical. The depth of the
malleolar portion is 20 mm. |
The hyomandibular is very similar to that found in Ichthyo-
dectes. The superior condyle is elongated, and depressed in the
central portion. In the skull of S. intermedius, figured by
Newton,” this condyle is shown to be regularly rounded from
before backward. As all the figures and descriptions of the
hyomandibular of this family and the Ichthyodectide show the
depression described above, I am inclined to think that this
portion may have been distorted in the specimen figured by Mr.
Newton. Extending downward from the anterior and posterior
angles there are two slight ridges, which converge toward the
center and form a much larger one, which extends downward to
nearly the lower extremity of the bone. It forms the outer
border of the groove for the preoperculum. There is also
another ridge on the internal side, just in front of the condyle
for the operculum, but it is not so prominent as the one just
mentioned. The condyle for the operculum is elongated, as in
Ichthyodectes.
The lower extremity of the bone presents an articular surface
similar in size and character to that found at the upper end of
the symplectic. I think it is very likely that these two bones
articulate at this point.
A small portion of a scapulais preserved. It shows only two
distinct articular condyles, one large and one small, instead of
three, asin Xiphactinus. Portions of several spines are pre-
served, of which one complete and a portion of another are
shown in the figure and need no further consideration.
The first anterior vertebra has the posterior end deeply con-
cave, but the anterior end is not so deep, and has a slight pro-
tuberance projecting forward above. On the superior surface
there are two deep, rounded pits for the neurapophyses; aside
from this there are no other grooves displayed upon the centra.
A small toothed element was found on the internal side of one
85. 1. c., pl. XXXIV.
eS ee a ee
»
STEWART. | Cretaceous Fishes. 328
of the maxille at about the point where the pterygoid should
lie, but it is too large to be a portion of this bone. There are
nine teeth upon it, which are about the size of those on the
anterior portion of the maxilla.
The type of the above was found in Gove county, Kansas, by
Mr. W. O. Bourn, of Scott, Kan., who loaned it to the Univer-
sity for study.
SAUROCEPHALUS.
Harlan, Jour. Acad. Nat. Sci. Phil., vol. III, p. 337.
Saurocephalus is similar in many respects to the genus just de-
scribed. Unfortunately the skull has never been found in either
this country or England, but from the great similarity of the
jaws to those of Saurodon a close relationship in general struc-
ture to this form can be expected. In Saurocephalus the mandi-
bles are not so slender as in Sawrodon, and do not project forward
as in this form. There are distinct foramina instead of deep
notches below the alveolar border internally, and the teeth have
very short and compressed crowns instead of the subcylindric,
knife-like crowns of Sauwrodon.
Thus far four species have been described from America, as
follows:
Saurocephalus lanciformis Harlan, Cretaceous, Missouri river.
Saurocephalus arapahovius Cope, Niobrara Cretaceous, western Kansas.
Saurocephalus dentatus Stewart, Niobrara Cretaceous, western Kansas.
Saurocephalus pamphagus Hay, Niobrara Cretaceous, western Kansas.
Thus it is seen that only three species have thus far been dis-
covered in this state, of which one, S. arapahovius, was founded
upon only a small portion of a maxilla which presents but few
characters that can be called specific. It is very hard to refer
specimens to this species with any degree of certainty, and I
am inclined to think that it may prove to be a synonym of S.
lanciformis.
Saurocephalus dentatus. Plate LVIII, 3a and b, 4a and b.
Saurocephalus dentatus Stewart, Kans. Univ. Quart. vol. vil, pp. 25-27.
This species differs from S. arapahovius in having the teeth
slightly striated and never overlapping on the maxilla. It is
also a slightly larger form. It was established upon the left
O24 University Geological Survey of Kansas.
maxilla, premaxilla and mandible of one individual and the
left mandible of another, and was collected from the Niobrara
Cretaceous of Wallace county, Kansas, by Mr. E. P. West.
The catalogue number of the specimen is 82.
The maxilla is much larger and more elongated than in S.
broadheadi, and the superior border is very thin and more ele-
vated just back of the condyle for the palatine than in this
form.
This condyle is very convex ; anterior to it there are two more
condyles, which are probably for the ethmoid and vomer. The
most posterior of these is broken away, but from the base it
appears to have been elevated, as in Saurodon. The anterior
condyle is large and triangular in outline, and is bounded in
front by a shallow pit not found in the form mentioned above.
There are alveoli for thirty-eight teeth, which decrease in size
toward the posterior extremity, where they rise but slightly
above the alveolar border.
The premaxilla is more or less plate-like ; externally it is con-
vex from before backward and the anterior border is quite ob-
lique. ‘There is probably no close connection with its fellow on
the opposite side, and the upper portion of the bone is covered
with fine lines radiating upward and backward from the ante-
rior inferior angle. The teeth seem to be somewhat smaller
than those on the maxilla; alveoli for nine are found.
The ramus of the mandible decreases more in depth toward
the symphysis than in Sawrodon. The lower border is very thin,
but becomes thicker toward the alveolar border, though it never
attains the robustness seen in Xiphactinus. Just back of the
symphysis and below the alveolar border there is a prominent
swelling, below which there is an elongated ovoid pit, near the
position of the Mento-Meckelian ossiclein Ama. The predental
surface descends in almost ‘a straight line from the alveolar bor-
der, and has numerous bony tubercles internally, probably for
ligaments attaching the two jaws. Just beneath the dentary
there is a long thin element which seems to be united to it by
suture and forms the lower border of the jaw. If this be true,
it may represent a new element in the mandible, although more
Srewart. | Cretaceous Fishes. 325
material will have to be examined before this can be accurately
determined. The teeth are compressed and appear minutely
striated under the microscope. [rom the center to the anterior
extremity of the dentary the teeth decrease in size, while those
on the posterior portion are nearly twice the size of those on the
maxilla. Spaces for about forty-two are present, but quite a
number of the teeth seem to have been shed and never replaced,
as the alveoli seem to be entirely closed in some instances.
The dermarticular sends a long, dagger-like process forward
internally nearly to the ovoid pit mentioned above; externally
it is soon covered by the dentary. The cotylus is somewhat
vertically directed, narrow laterally, and slightly convex from
above downward.
The predentary is a triangular element joined to the dentary
by avery irregular surface, broader above than below. The
superior border is finely rugose and edentulous, and the tip is
acute. This bone is unpaired, as in Saurodon.
Beneon or maxilla and promaxilla...... ..........65..0.00 161.5 mm.
Depth of bone posterior to condyle for the palatine......... 44.5 *&
Height of condyle for palatine above the alveolar border.... 48.5 *
emeene Gl DreMa dill, TUIOTIOF LY. 0... co eck oles bs oe Stan Be
ELAS g: a eo ao
Average anterior posterior length of crown................. Soo
Number of teeth in one inch, 9.
manewwie: Iueneibh from GOUglus ... i. cece eee eee ace eee 161.0 mm.
Leneth Of alveolar DOTGer. ) i... ee tee 140.0
Depth of predental surface (estimated) ......... Ba) &
Average height of crown, posterior.............. BOh
Average length of crown, posterior.............. , oe
(SS TURE ASE Do 29.5 *
| SR ae ne ee yo ne oe O: &
326 University Geological Survey of Kansas.
STRATODONTIDA.
The family Stratodontide, as accepted by Professor Cope in his
‘‘Cretaceous Vertebrata,’’ embraced certain genera of physosto-
mous fishes which bear a general relationship to Msoxz, and in-
cluded the genera Stratodus, Hmpo, Pachyrhizodus, Enchodus,
Tetheodus, and Anogmius, for which the following synopsis was
given :
J. Premaxillary with several rows of teeth:
Palatine teeth numerous, large; all with pulpcavity, Stratodus.
IJ. Premaxillary with two rows of teeth:
Maxillary bone short; dentary with equal large inner
teeth and outer rows en O7osse.................: LEMpo.
Maxillary bone very long; one row of equal dentaries, Pachyrhizodus.
III. Premaxillary with one or no row of teeth:
A large premaxillary fang; anterior maxillary and
dentary teeth enlarged; cutting edges not oppo-
Sites Unsymimetrical. oer eee ea ee ere Einchodus.
Premaxillary toothless; anterior maxillary and den-
tary enlargaed. ois gpe oe tye cok See me eee ete eee me eee Tetheodus.
The characters of these genera are so varied that I have
thought it best, in the present work, to divide the old family of
the Stratodontide into four separate families, and to use this
name for Stratodus, Empo, and Cimolichthys, which may seem
remarkable when the great difference in the form of the teeth
and tooth-bearing elements of the genera are taken into con-
sideration. In Stratodus the teeth are ali small and present
a great similarity throughout the whole dental series, while in
Empo they are of a different size and form on the several parts,
and all are different from those in Stratodus. There are also
marked differences in other parts, as the form of the palatine
and the different lengths of the premaxille.
Notwithstanding all these differences, when we examine the
top of the skull of each, a remarkable similarity between the
two is seen, from which resemblance of the two forms we must
conclude that there is a close relationship between the two,
notwithstanding the differences of habits indicated by the
dentition.
STEWART. | Cretaceous Fishes. 327
The top of the skull is very flat in both genera, and made up
for the most part of the elongate, triangular-shaped frontals,
which are united in the median line by means of a long,
straight suture. These bones are covered on the superior sur-
face by small ridges or striw#, which radiate from a point inter-
nal to the postorbitals and are most strongly marked in Lmpo.
The supraoccipitals are small and invade the top of the skull
but little. It is raised into a slight crest in Empo, and prob-
ably so in Stratodus.
Below is given a revised synopsis of the family :
I. Premaxilla short, with several rows of teeth:
Palatine and mandibular teeth numerous, in several
2 ee aay ee ete oe Stratodus.
II. Premaxilla elongate, with one row of small teeth:
Mandibular teeth in two series, of which the outer are
small and in several rows, while the inner are very
large.
Palatine teeth large and in two rows.................. Empo.
Ill. Premavxilla and maxilla elongate, with very small teeth:
Palatines and ectopterygoids provided with powerful
teeth, some of which are semibarbed in shape....... Cimolichthys.
, STRATODUS.
This genus, as characterized by Cope, is remarkable for the
small size, great number and peculiar form of the teeth. Un-
fortunately the material from which the descriptions have been
made was very fragmentary, from which no adequate conception
of the cranial characters could be obtained. The jaws as well
as most of, the bones are very fragile, and it is probably owing
to this fact that no perfect specimens have been collected. The
geological expedition of 1898 to western Kansas obtained a
specimen belonging to this genus from near T'win Buttes, Wal-
“lace county, that shows some points not known before. The
specimen is far from perfect, but it is much better than any
specimen heretofore described.
The premaxilla is small and has three or four rows of teeth
on the lower side, of which those on the inner are the largest.
The dentaries are very slender and are covered with numerous
rows of small conical teeth on the internal side which are all
‘7
(Sly)
28 University Geological Survey of Kansas.
provided with a large pulp cavity. The teeth are largest on the
external row, but toward the symphysis they are all reduced to
nearly the same size. The symphysis is very shallow. The
palatines (?) are long and slender plates of bone, which are coy-
ered with teeth probably similar in form and size to those found
on the dentary. Fortunately the top of the skull is preserved,
showing it to be slender, flat, with the different elements arranged
very much asin Hmpo. ‘The vertebre are very similar to those
of this genus.” ‘Their centra in both abdominal and caudal
regions are elongate and contracted medially. There is a shal-
low longitudinal groove at the bases of the neural and hemal
arches, which are divided vertically by a median rib-like but-
tress. ‘The median lateral portion is smooth or nearly so.’’
The known American species are :
Stratodus apicalis Cope, Niobrara Cretaceous, western Kansas.
Stratodus oxypogon Cope, Niobrara Cretaceous, western Kansas.
Stratodus apicalis. Plate LX; plate LXI, fig. 1.
Stratodus apicalis Cope, Proc. Am. Phil. Soc. 1872, p. 348; Cret. Vert.
West, p. 227.
This species is represented by portions of the skull and tooth-
bearing elements as mentioned above.
The dentary is elongate and covered with several rows of
teeth above and on the internal side. These teeth are largest
in the outer row, where they are seven millimeters or more in
height, but become very small toward the inner and lower row.
The teeth are round in section, acutely pointed, and with
slightly recurved crowns. They are entirely without striae even
under the microscope. Toward the symphysis the number of
rows diminish until only two remain at the extremity, with the
exception of three or four small teeth situated external to the
outerrow. ‘The symphysis is very shallow, in fact, there is only
a small articular facet at this point, which shows that the jaws
were very loosely united here. Throughout its whole length
the dentary is very shallow and covered with coarse longitudinal
strie on the external side. Unfortunately the articular portion
is not preserved. |
86. Proc. Am. Phil. Soc. 1878, p. 181.
®
STEWART | Cretaceous Fishes. 329
There is a bone which I think represents the premaxilla. The
teeth are arranged in three or four rows, the largest being in-
ternal. The external face of the bone is non-striate, and ex-
tends upward and backward, while the internal side is grooved
for the maxilla.
In the description of this part, as given by Cope,” he says
that the premaxillary teeth are in two series. What he means
by this statement I am unable to say, as four rows of teeth are
shown in the figure of his specimen, and there does not seem to
be any abrupt change in size transversely.
The palatine (?) is a long, slender plate of bone, contracted
at both extremities, and covered with teeth on the lower side.
The number of rows decrease toward both ends and the size
of the teeth also becomes less toward one of them. The up-
per surface is divided longitudinally by a ridge and the three
grooves mentioned by Cope,” but they are not so strongly
marked as to attract particular attention.
Dentary: Greatest width of dental band .................... 10 mm.
Length of dental band (estimated)................ sae -**
Premaatas weneth Of Gental band .... 4.0606... Seen wees ei wk
Greatest width of dental band.................. Oras
RENMEI A RMEMIBTRE Seer oa Pied. lavei sis vite de eTa cos Bin gee dae a’ Ga > i RAI ice
Sic hs eg Gn eo he
The top of the skull is preserved completely, or nearly so.
The bones represented are the frontals, parietals, exoccipitals,
and probably the epiotics and pterotics.
The frontals are very long bones, broad behind, and probably
tapering to a point in front where the extremities are covered
with matrix and other bones. On the upper surface the bones
are beautifully sculptured with long, coarse strive extending
forward from near the center of each bone, while the back por-
tion presents only an occasional stria. These bones are espe-
cially remarkable for their great length and fragility. The
suture between the parietals and the epiotics cannot be made
out, although this portion is entirely free from matrix. For
this reason | am inclined ta think that these two bones are co-
87. Cret. Vert. West, p. 226, pl. XLIX,
88. Cret. Vert. West, p. 227.
99— vi
330 University Geological Survey of Kansas.
ossified in the specimen. They are somewhat triangular in out-
line and meet each other in the median line, thus separating
the supraoccipitals from the frontals. They do not seem to
possess any very characteristic markings. The pterotics (?)
are prominent processes of bone that project outward from the
parietals and join the exoccipitals below. The supraoccipitals
do not seem to be present in this specimen, but are probably
not raised into acrest. The exoccipitals are rather large bones
projecting upward and outward from the basioccipital and form
a large portion of the back part of the skull. Each sends a fan-
like process of bone backward. ‘There is still another long, flat
bone not connected with the rest of the skull that may repre-
sent a portion of the hyomandibular or an opercular. It is
somewhat irregular in outline and presents a sharp longitudinal
ridge near one of the borders. Measurements of skull are:
Distance from supraoccipital to anterior extremity of frontals, *166 mm.
Transverse distance across spterociessaqe, sae ener eee 65
Greatest transverse distance across frontals................. 60
Just back of the skull there is a bone that is no doubt con-
nected in some way with the vertebral column. It somewhat
resembles a terminal caudal vertebra or urostyle. It is con-
tracted and convex at one extremity and expanded and deeply
concave at the other. Near the center of the convex end there
is a foramina leading inward which probably continues through
to the other side.
EMPO Cope.
This genus is represented in the museum by the remains of
several individuals, more or less perfect, from the Niobrara
Cretaceous and a portion of one specimen from the Fort Pierre
of Kansas. It is probable that they include four species.
The palatines are much elongated and bear two rows of teeth
on the anterior portion, of which the external are large and the
internal somewhat smaller. The ethmoid is contracted into a
rather sharp beak anteriorly and bears a number of small teeth
on the median line below, which are either arranged in one or
two rows or form a small cluster near the anterior extremity.
* Wstimated.
Se eS SS
STewakt. | Cretaceous Fishes. 331
The dentaries bear several rows of teeth, of which those on the
external side are minute and somewhat thickly placed, while
those on the internal side are large, set upon expanded bases,
and are well separated from each other. The outer surface of
the dentary is covered with numerous well-marked sulci. The
cotylus is bifurcated in the center, each half appearing asa
small hemispherical pit. There are several other tooth-bearing
elements, among which is a long, slender bone with a single
ap?)
row of small conical teeth on the lower side, which probably
represents a premaxilla.
The top of the skull is elongated and somewhat triangular in
outline. It is especially remarkable for the great extent and
delicate markings of the frontals, as well as of the other bones
of this region. The supraoccipital is small and enters slightly
into the formation of the top of the skull. The parietals are:
small and do not extend farther forward than the anterior ex--
tremity of the supraoccipital.
Throughout the whole extent of the top of the skull there is
a close resemblance to the corresponding portion of Stratodus,.
the principal difference being found in the larger size of the
parietals and their meeting along the median line.
There is one specimen at hand that has the vertebral column
preserved completely, and another one nearly so, from which
we see that there were altogether about fifty-three vertebra.
Those in the cervical and dorsal regions are finely striated,
while those near the end of the tail are deeply grooved on the
sides, the grooves commencing on the eighth or ninth vertebra
from the end of the series. The neural canal is bounded by a
lamina of bone on each side, outside of which the neuropophyses
rise and form a vertical ridge on the side. The hemal canal
is bounded by lamine similar to those described above. The
hemal arches are located outside of these, so that in a detached
vertebra it is often difficult to determine the two sides.
In Cope’s description of the vertebral column of L. nepxolica™
he says that up to the fourteenth vertebra the neural canals are
not bounded by vertical lamin; in this I cannot agree with
89. Cret. Vert. West, p. 230.
oor University Geological Survey of Kansas.
him. I have been able to examine some of the detached ver-
tebree from near the head, and so far as I can judge the sides of
the neural arches do not differ materially from those farther
back in the column.
The abdominal cavity is short, with well-developed ribs. The
caudal fin is not well preserved in any of the specimens, only
the bases of the rays being present. On one of the slabs there
are a number of small rays and other fragments just back of
the abdominal cavity, which is probably a portion of the pelvic
fin. . The rays are small and have no great length. The pec-
toral rays are also rather delicate and curved near the proximal
extremity, while near the middle portion and distal end they
are cross-segmented. Along the back there are numerous dermal
plates, the most of which have no characteristic markings.
The known American species are:
Empo nepeolica Cope, Niobrara Cretaceous, western Kansas.
Empo semianceps Cope, Niobrara Cretaceous, western Kansas.
Empo contracta Cope, Niobrara Cretaceous, western Kansas.
Empo merrilli Cope, Niobrara Cretaceous, western Kansas.
Empo lisbonensis Stewart, Fort Pierre Cretaceous, western Kansas.
Empo nepeolica. Plate LIX.
Empo nepceolica Cope, Proc. Am. Phil. Soc. 1872, p. 347; Cret. Vert.
West, p. 230.
Empo sulcata Cope, Hayden’s Bull. U. 8. Geol. Surv. No. 2, p. 46.
Cimolichthys sulcatus Cope, |. ¢., p. 351.
This species is represented in our collection by the remains
of several individuals, among which is a skull almost complete,
from which a fair idea of the anatomy of this portion can be
gained.
The bone described and figured by Cope as a premaxilla has
since been shown to be a palatine by specimens in the British
Museum which have the maxille and premaxille in place.
The bone is elongate and curved longitudinally, and at the an-
terior extremity there is a rostrum-shaped apex of dense bone
which projects forward beyond the last tooth. On the superior
border there is a slight crest of bone, which probably serves to
loosely connect this portion to the skull. The teeth are arranged
in two series. At the anterior extremity there are several large
‘
iP
ares
—— ee eae
STEWART. | Cretaceous Fishes. 333
teeth extending backward for three or four centimeters from the
extremity, which are followed by a series of much smaller ones
occupying the inner side of the bone and extending backward
to near the posterior extremity, where there are usually one or
more teeth much larger than any of the others. Near this point
those on the internal side become very small. The bone called
maxilla by Cope is probably a pterygoid. It is short, bar-like,
and bears a single row of teeth, which decrease in size poste-
riorly. The posterior extremity is edentulous and somewhat
expanded.
Palatine: Length to large posterior tooth..................... 86 mm.
MRM EE TE SSE BONER Toe, Grado sf bic te Willis vier uel c Sinin’ Ha oa wee Bosh
Transverse distance across alveolar border near cen-
Hae it Sere SOE GIS Be Rg, 5 ee a eg Re ee es
Dee ()): Iuenpin (CStiMated) . .. .. eee le e e e Gives.
The dentaries are very much elongated and contracted regu-
larly toward the symphysis, where the bones are very shallow.
The symphysis is not very deep, consisting, for the most part,
of a small flat facet on the internal side which meets its fellow
on the opposite side, and to which it is probably united in a
manner somewhat similar to that found in Pachyrhizodus.
Cope” describes this part as presenting a marked fossa which I
am unable to find in any of our specimens, and [ think it not
at all unlikely that this was an individual] peculiarity of his
specimen. ‘The alveolar border supports two series of teeth, of
which the internal are the larger of thetwo. There are usually
about ten teeth present in this series, and room for probably as
many more, which are often represented by the tooth scars of
the shed teeth. These teeth are set upon hemispheres of bone
firmly anchylosed to the jaw. The teeth are compressed, finely
striated, and directed inward. ‘The external series consists of
several rows of teeth of which those on the internal row are the
larger. They are all directed inward and seem to have both
anterior and posterior cutting edges. Toward the posterior
extremity both series fade out, leaving a considerable space
between the last tooth and the end of the bone. The external
surface of the dentary is covered with deep longitudinal sulci,
90, Cret. Vert. West, p. 231.
334 University Geological Survey of Kansas.
while the internal is nearly smooth and deeply grooved. The
articular portion sends a long, sword-shaped process forward,
but how far cannot be determined, as the anterior portion is
covered by the overlying ceratohval. The cotylus is very broad
transversely but has no great vertical extent. It is composed
of two concave facets separated from each other by a well-
marked ridge. The external of these facets seems to be slightly
the larger of the two.
Premaxillaauengthy wri oe. cael cain ony nee Raia oe tenes Ha 117 mm.
Maxilla: “eng th (estimated 22.15 os ee
Dentary: Length of alveolar border to last tooth............. 182 *
Heng thvof mandiblefromycotylus <2.) 55. ee eee 252 **
@ransvyerediameteriof cotylus a5 10 ek wee eee ee nee Pass
Length of articular from cotylus (estimated)................. jason al
The quadrate is compressed and fan-shaped ; extending down
its posterior border there is a deep groove, which probably
accommodates the preoperculum. Just in front of this
groove, on the internal side, there is still another groove
which extends over half way down to the condyle, in which the
symplectic probably fits. Cope*’’ was unable to discover a sym-
plectic in the specimen of EH. semianceps which he described,
and I am inclined to think he was of the impression that this
genus did not possess this bone, but from the presence of this
groove we can assume that it was proportionately broader than
in the Saurodontidx. The anterior border is very thin and is
projected upward, with a small portion of the superior border,
into quite a prominent process of bone. The condyle is bifur-
cate in the center and has a superficial resemblance to the distal
end of the mammalian femur. The external side cannot be
seen.
The preoperculum is long and very slendor in the superior
portion. Near the lower extremity the bone bends suddenly
forward and broadens out into a thin plate below. The upper
portion is only slightly curved and very roughly and irregularly
marked. A portion of one operculum is present, showing the
internal side. It is a broad, thin plate of bone, with a small
condyle for the hyomandibular. The condyle is not situated so
91. Cret. Vert. West, p. 229.
STEWART. | Cretaceous Fishes. * 885
near the superior border as in the /chthyodectidx, and has a
strong ridge extending backward from it on the internal side.
The hyomandibular presents two articulating surfaces above
for union with the skull. These are slightly grooved longi-
tudinally and are separated from each other by a deep notch.
The anterior of these is projected well forward. The condyle
for the operculum is situated probably more than one-half the
distance down the side of the bone, and is elliptical in outline
and very irregular. Just in front of the condyle there is a
prominent ridge which extends downward, but not to the lower
extremity of the bone. The inferior portion is rather narrow ;
the anterior very thin.
The cerato- and epilyals are preserved in one specimen, while
the urohyal is shown in another. The epihyal is very thin and
has the anterior extremity slightly convex for the ceratohyal.
Near the postero-superior end there is a deep notch, probably
for the accommodation of the interhyal. Both the external and
internal sides are covered with fine strie. The ceratohyal is
an elongate, slender bone, expanded at the extremities and con-
tracted slightly at the center. Both sides are covered with
striew, which are especially noticeable toward the extremities,
which are nearly flat. The urohyal is a small, triangular ele-
ment, badly crushed, with a small facet on the anterior end for
its fellow on the opposite side.
There is a long, thin plate of bone that I am inclined to
think represents the palatine. It is contracted to a blunt point
at both extremities, and covered with fine striz# on one side
which radiate from a point near the superior border and about
one-third the distance back from the anterior end. The central
portion of the lower border is covered with a single row of
small conical teeth, each of which seems to be firmly anchy-
losed to the bone, with the apex directed forward: The ex-
tremities of this border are edentulous.
The ethmoid is a symmetrical tongue-shaped bone covered
with teeth below. Professor Cope” says this bone is sometimes
symmetrical and sometimes unsymmetrical, and he was in-
92. Cret. Vert. West, p. 228.
336 University Geological Survey of Kansas.
clined to think that it represented a superior or inferior phar-
yngeal. In all of the specimens that I have examined it seems.
to be entirely symmetrical, and in one specimen it is in place
at the anterior extremity of the skull, so there can be no doubt
of its being an ethmoid. I am inclined to think that the
asymmetry of Cope’s specimen was due to distortion. The an-
terior end is extended forward into a beak of dense bone, which
is directed slightly downward. Just back of the beak, on the
lower side, there are a number of small teeth, which are ele-
vated considerably above the rest of the bone when it is re-
versed. There seems to be a great deal of irregularity in the
arrangement of these teeth. In some specimens they are
grouped together, with the largest near the center, while in
others there are a number of large teeth anteriorly, with a
single row of smaller ones extending back from them. ‘The
upper surface of the bone is covered with numerous striw radi-
ating from the apex.
There are fragments of other bones whose location cannot be
determined. One of these is a long bar of bone which has an
articular surface at the anterior end and is expanded at the
other ; another is a flat semicircular plate, finely striated ; while
others are irregular in outline.
The whole of the upper part of the skull is preserved, either
by the bones themselves or by their impression in the chalk.
It is very flat and bears a marked resemblance in general out-
line to the top of the skull of Stratodus. The frontals form the
greater part of the upper walls of the skull and the two are
separated by a well-marked suture in the median line. The
two bones together are somewhat triangular in outline and are
beautifully sculptured, on the upper surface, by radiating
ridges. These ridges are especially marked just internal to
the postorbitals, from which point they radiate toward the
front and back of the skull. They are in contact with the
supraoccipital posteriorly. The supraoccipital enters but little
into the top of the skull. It projects backward in a small
crest, which is probably not raised above the rest of the skull.
The postorbitals form beak-like processes on the sides and are
STEWART. | Cretaceous Fishes. 337
directed slightly downward. Inferiorly they form a prominent
ridge on each side which extends inward toward the median
line. The occipital condyle is rather flat in this specimen, but
in another specimen, where the back of the skull is shown, the
condyle is deeply concave. It is likely that this specimen may
be of another species, as it seems to be more narrow than the one
under consideration. It is crested below and united with the
exoccipitals by well-marked sutures. There are several other
bones at the base of the skull, but they are so badly crushed
and their sutures so obliterated that they cannot be made out
with any degree of certainty.
Ly BOR SSO OSGI na Ce ee 239 mm.
Distance across postorbitals (estimated).................. IZG~ **
A portion of the pectoral fin is present, with fragments of the
pectoral girdle. The rays are small in size and seem to be very
numerous. They are nearly straight, excepting near the proxi-
mal end where they are strongly bent. The distal end of each
ray is cross-segmented.
Empo lisbonensis. Plate LXI, figs. 10a and b.
Empo lisbonensis Stewart, Kans. Univ. Quart., vol. vit.
This species was established on the left palatine of a single
individual from the Lisbon shales, Fort Pierre Cretaceous.
The specimen was found by myself one mile northeast of Lisbon,
Logan county, Kansas, and is in a fair state of preservation.
It indicates a fish of about the size of EF. nepwolica Cope. The
catalogue number of the specimen is 528.
The palatine is much more depressed than in the species just
mentioned, but is broader across the alveolar portion, giving
the bone a very robust appearance when seen from below. ‘The
anterior extremity was probably not so acutely pointed as in
the other species of this genus. None of the teeth are preserved
complete, but there are alveoli present which show that there
was an outer row of large teeth and an inner row of small ones.
The outer row are reduced in size toward the distal extremity,
and may entirely disappear before the end is reached, as there
are no tooth scars on this portion. At the posterior end of the
338 University Geological Survey of Kansas.
internal row there is a very large tooth, back of which there is
a broad shallow pit.
Length to the large posterior tooth.................5./2.29- 82 mm.
Greatestitransverserwadthh 0202 oon tach ae ee Os
Empo semianceps. Plate LX1, figs. 6-9.
Cimolicthys semianceps Cope, Proc. Am. Phil. Soc. 1872, p. 351.
Empo semianceps Cope, Cret. Vert. West, p. 233.
The material, consisting of the palatines, fragmentary mandi-
bles, and portions of other parts, is referred to this species with
some doubt. It seems to correspond with the figure and de-
scription of this species as given by Professor Cope,” but it may
possibly be a specimen of FE. merrilli. The specimen, No. 278,
was collected from the Niobrara Cretaceous of Graham county
by Messrs. Overton and Martin during the summer of 1895.
The palatine is nearly straight on the internal side and de-
pressed above. The teeth are arranged in two series, as in
I. nepeolica. The most anterior one of these is rather small,
but is followed by two or three that are somewhat larger. The
teeth following are of about the size of those just mentioned,
excepting toward the posterior extremity, where they become
slightly reduced in size. There seems to be a considerable dif-
ference in the arrangement of the teeth on the internal side in
the two palatines of this specimen. On the bone of the right
side there is a continuous row which are directed strongly
backward, while on the left there are but two tooth scars pres-
ent on the internal side. The pterygoid is very similar to that
of ly. nepxolica described above.
Palatine sen ths airs sy sey ace eerie: Wp etevar Be AEA oe 166 mm.
Greatest transverse wid tins 2 ee ee epee OWNS
The dentaries are elongate and contracted in depth at the
symphysis, where the lower edge is beveled off to meet the su-
perior border, thus forming an acute apex. The external side
is covered with longitudinal strie which are not as strongly
marked as in L. nepxolica. These strive are more noticeable on
the external portion of the articular than on any other part.
The lower border is rather sharp but soon becomes thickened
93. Cret. Vert. West, p. 233, pl. LIIT, fig. 6.
STEWART. | Cretaceous Fishes. 339
above, especially in the posterior portion, where there is a strong
bar just below the articular. The internal side is finely striate
and deeply grooved. The small teeth on the external side are
in two rows, of which those on the external row form a minute
fringe, while. those on the internal are larger and are directed
slightly forward. Those of the internal series are large at the
center but decrease in size toward the extremities. Hach of
these is set upon a bony tubercle, as in lL. nepwolica. The ar-
ticular portion is very similar to that portion of the species
just mentioned.
PRET VOOIAT IOIUOL |. but act Ge. s ak OSG 6 Sia bdo ce eee 121 mm.
EMRE TG GET ETE Se cd hike tae fle oa sh od Gmc Glee eve ele lect oo.“
PMNVOLSS MIBMOLEr Of COLVIUS o5 oc ec oo ce we ee ee Lares +
There are numerous other bones, including the ceratohyals,
postorbitals, and a fragmentary ethmoid. The ceratohyals do
not differ materially from those of I. nepwolica excepting in
size. ‘The ethmoid is more blunt than in the species just men-
tioned and bears a single row of eight or nine teeth on the
median lihe below. The postorbital is somewhat hook-shaped
and has a roughened articulating surface internally.
Empo contracta.
Empo contracta Cope, Hayden’s Bull. U. S. Geol. Surv. Terr. No. 2, 1874,
p. 46: Cret. Vert. West, p. 232.
The specimen referred to this species consists of the greater
portion of palatine and dentaries, the hyoid arch, numerous
vertebre, and other bones. As the original description of this
species is rather vague and the figures are of fragments only, it
will always be difficult to identify this species with any degree
of certainty unless access is had to the type specimen for com-
parison or until it is more completely described and figured.
The specimen is much smaller than the one described above as
EI. semianceps.
Only the internal sides of the dentaries are shown. The ex-
ternal series of teeth do not seem to be as large as in FE. semian-
ceps described above. The teeth of the inner series are long,
acutely pointed, and directed slightly inward at the apices.
The maxille are present with probably all of the teeth, which
340 University Geological Survey of Kansas.
are elongate, with anterior and posterior cutting edges, and are
recurved. Their size seems to diminish very suddenly toward
the posterior extremity, which is expanded. ‘The epi-, cerato-
and urohyals are in place on one side but do not seem to differ
materially from those described above. A portion of the pec-
toral fin shows the rays to be cross-segmented as in HL. nepx-
olica, but the proximal ends are straighter than in this form.
The bones are for the most part in so fragmentary a condition
that only approximate measurements can be made; hence they
are omitted.
OSTEOGLOSSIDE.
The genus Anogmius, described by Professor Cope,” was left
by the author in his family Stratodontidx, to which, it seems evi-
dent, it does not belong.
After carefully studying the material at hand, it seems to bear
a close relation in many respects to the genus Plethodus, recently
described and figured by Dr. A. 5. Woodward,” which the
author is inclined to place in the family Osteoglosside. For the
present, at least, I think that Anogmius should be left in this
family.
ANOGMIUS.
Cope, Proc. Am. Phil. Soc. 1870, p. 170.
This genus was first described by Professor Cope” from ver-
tebree and fin remains of an individual from the Niobrara Cre-
taceous of western Kansas; the name Anogmius contractus was
applied to it, and, later, two more species,” A. favirostris and
A. evolutus, were added to the genus by the same author. The
first of these was a small individual, while the second was much
larger, and is probably the same form described as ? Beryx multi-
dentatus,”* described by myself. Professor Cope also mentions
another species, A. aratus, but so far I have been unable to find
any other reference to it. Iam inclined to think that this was
a slip of the pen on the part of Professor Cope, and that A. con-
94. Proc. Am. Phil. Soc. 1871, p. 170. 97. Proc. Am. Phil. Soc. 1878, pp. 178-180.
95. Proc. Am. Phil. Soe. 1870, p. 170. 98. Kans. Univ. Quart., vol. VII—A, p. 196.
96. Annals and Magazine of Natural History, ser. 7, vol. III, pp. 353-361.
STEWART. | Cretaceous Fishes. 341
tractus was intended to be expressed, as the author says:” ‘‘The
characters of the genus Anogmius Cope having up to the present
time rested upon but one species (A. aratus), it is satisfactory
to be able to confirm them by the study of new material,’’ etc.
This genus is well characterized by the tooth bearing elements,
all of which seem to be covered with villiform teeth. The pre-
maxille are triangular in outline and covered with a semiellip-
tical band of teeth below. The mavxille are elongated and
slightly concave on the lower border, which is covered with
several rows of small teeth. The jaws are bound directly to the
skull by means of a small condyle above, which presents a
roughened surface, probably for cartilage. The dentaries are
also covered with small teeth similar to those on the maxille
and premaxille, and are incurved and loosely united at the
symphysis. The angle is prominent, and the mouth as a whole
is quite oblique.
The top of the skull is flat, and beautifully sculptured with
coarse strize radiating from near the center of each of the prin-
cipal bones. The frontals are long, while the parietals are
rather small. The supraoccipital is depressed and invades the
top of the skull very little, if any. The orbital cavity is large
and the sclerotic ring very thin. In this region there are
numerous membranous bones which probably form a covering
for the top of the skull to a greater or less extent. The oper-
cular bones are thin and covered with coarse strix.
In one specimen of A. polymicrodus there are seventy-two ver-
tebrae exposed to view, and there are probably eight or ten
more hidden by the opercular bones just back of the skull,
making, in all, eighty vertebrae in the complete column. The
centra are all rather short and do not present the lateral grooves
found in the Saurodontidx and Ichthyodectide, but are striated
and somewhat resembling the vertebre of the genus Pachyrhi-
zodus Agassiz in this respect, and it is probably for this reason
that Zittel gives Anogmius as a synonym of this genus.” Just
in front of the caudal fin the vertebre are crowded together,
99. l. c., p. 178.
100. Zittel’s Handbuch der Paleontologie, b. III, s. 268. -—
342 University Geological Survey of Kansas.
and those supporting the fin are very much so. ‘The last
vertebra is succeeded by a fan-shaped expansion of bone or
urostyle, which presents a well-marked lateral ridge on the side
exposed. The neural arches fit into pits at their bases, and in
the caudal region they have laminz of bone projecting down
the sides, forming yoke-like articulations with the centra. The
arches are expanded at their bases and are largest in the anterior
region, where they are directed backward but slightly, while in
the region of the fiftieth posterior vertebra the arches are very
slender and directed strongly backward. ‘The manner of attach-
ment of the haemapophyses cannot be made out with certainty,
although they were probably inserted in gomphoses, as are the
neurapophyses. The ribs are long and slender and slightly ex-
panded at the proximal extremity. ‘They seem to be inserted
in pits on the sides of the centra. The caudal fin is homocer-
cal, and is composed of numerous rays which articulate with the
centra by means of the yoke-like articulation mentioned above.
Toward the distal extremity these rays gradually split up into
many small filaments, which makes this portion very difficult
to collect in perfect condition. One of the smaller specimens
of this genus shows an imperfect impression of the dorsal fin.
It seems to have been composed of short spines rather far
separated and extending along the greater extent of the back.
There are no remains of the other fins sufficiently well pre-
served to show their important characters.
The following species are known :
Anogmius contractus Cope, Niobrara Cretaceous, western Kansas.
Anogmius aratus (?) Cope, Niobrara Cretaceous, western Kansas.
Anoymius evolutus Cope, Niobrara Cretaceous, western Kansas.
Anogmius polymicrodus Stewart, Niobrara Cretaceous, western Kansas.
Anogmius polymicrodus. Plate LXIV; plate LXV, figs. 1-7; plate LX VI;
plate LXVII, fig. a.
Beryx polymicrodus Stewart, Kans. Univ. Quar., vol. vir, pp. 195, 196.
This species was first described by myself as Beryx polymv-
crodus from some fragments of jaws collected in western Kansas
several years ago, but the material was too fragmentary to give
any idea of the animal. During the past summer the geolog-
STEWART. | Cretaceous Fishes. 343
ical expedition to western Kansas was fortunate enough to find
several good specimens from the Butte creek region of Logan
county, and was also loaned an almost complete specimen, the
fins excepted, by Mr. Travis Morse, of lola, who secured it sey-
eral years ago while collecting vertebrate fossils in this part of
the state. From this specimen a good idea of the anatomy can
be gained.
The bone called premaxilla in the preliminary description is
evidently not a premaxilla, but some other bone, the location
‘of which I have been unable to determine. The premaxilla is
rather short, with a semiellipitical tooth band on the inferior
side covered with several rows of villiform teeth, all of which
seem to be directed inward. Above the tooth band, on the ex-
ternal side, there is a thin wall of bone extending upward,
which is covered with coarse longitudinal striz on the posterior
portion, while the anterior part, just behind the apex, is cov-
ered with minute tooth-like tubercles, which extend backward
for some little distance just over the tooth band. The internal
side of the bone is deeply concave, the width of the concavity
becoming very narrow at the anterior end. The union between
this bone and the maxilla was no doubt very loose, allowing
this bone to be moved very freely. The two bones were proba-
bly not united anteriorly. ;
The maxilla is moderately long and thin transversely toward
the posterior extremity. It bears a tooth band on the lower
border, which is slightly concave longitudinally in front, where
this band is the broadest. Posteriorly the tooth band gradu-
ally contracts in width until the two borders come together at
the posterior extremity. The surface for the premaxilla is
directed slightly inward, and in some of the specimens it is
slightly roughened for the ligaments binding the two bones to-
gether. Just back of this, on the superior border, there is an
elevated articular portion which serves to bind the maxilla to
the skull proper. It is rather rough above and does not allow
the free motion of the upper jaws found in some of the other
families of physostomous fishes. Just back of this there is a
thin crest of bone extending backward over one-half the length
344 University Geological Survey of Kansas.
of the jaw. Both the external and internal sides of the bone
are covered with coarse longitudinal strie, the intensity of
which varies in different individuals.
Maxilla) enethiof tooth band iene) sae ee eae 118.0 mm.
Heveht at posternoniexpremibye se senna eee 23a NOs
Prema xillae: Teme they ey Se pas eaten ae ee eet oe SN UCR nen SOs:
Greatest height i097 he Bae es ee ee ee O10 we
The dentary has a tooth band above, covered with teeth very
similar to those on the parts described above. It is nearly flat
in front, but soon becomes directed downward internally, giv-,
ing it a somewhat beveled appearance, which may be due to
compression, although it seems to occur in all of the specimens
examined. The band is broadest near the center and is over-
hanging externally and in front. The symphysis is composed
of two parts: an upper facet, which is directed inward and
comes in contact with a corresponding facet on the opposite
side ; and a lower one, which is directed forward and enters but
little, if any, into the articulation of the two jaws. At the sym-
physis the bones are shallow, but they soon broaden and be-
come very deep at the coronoid process. ‘There are very coarse
ridges and grooves on both the external and internal sides.
The articular extends well forward on the external side, and is
united with the emargination in the dentary by an indistinct
suture. The cotylusis supported principally by a thick shelf of
bone extending inward from the main portion of the bone. It
presents a small concave facet which looks slightly forward,
back of which the angle is slightly recurved, and has been de-
scribed by Professor Cope™ as resembling a boot with the toe
inverted.
enethrotmandiblestromicotvilusia. ere nae cere eee eee 204 mm.
Depth at coronoid process (estimated)...................-+-.. Gore
Depth iat, sym phy sissies othe ie oh eee ele ar en ee OW Geib
ene th of tooth bandi (estimated) cee eee ere Wy OF
The bone originally described as a premaxilla'’ may be a
pterygoid, as it cannot be a vomer or palatine from the descrip-
tion of these parts, as given by Professor Cope. It is slightly
curved longitudinally and has a broad tooth band on one sur-
101. Proc. Am. Phil. Soc. 1878, pp. 179, 180.
HR I Gan top ahi
STEWART. | Cretaceous Fishes. 345
face, which bears denticles slightly larger than those on the
parts described above. On the opposite surface from the tooth
band there is a ridge extending its full length, which is over-
hanging on one side, thus forming a groove, which is probably
for the reception of some other bone.
OP DS as cll eg RoR OE BE OO ee a 69 mm.
Greatest width of tooth band ............ Be hvac yates fy ae ats Tie. 368
There are two other bones which are covered with small pits,
and somewhat resemble those found on the tooth-bearing ele-
ments described above, although it is likely that these bones
never bore teeth. One of these is a cordiform bone, and is
probably a pharyngeal. The pitted surface is slightly de-
pressed in the middle, and the sides slope sharply toward the
edges. The other element is not so thick as the one just men-
tioned; it has a broad band of these pits on one side and a
median ridge on the other, somewhat similar to that found in
the supposed pterygoid described above. At one of the ends
there is a roughened articular surface. There is another long
and slender element that bears a superficial resemblance to the
palatine bone of Stratodus apicalis, which is no doubt the same
bone described as a palatine by Professor Cope.'” It is broader
at one end than at the other. The lower (?) surface is covered
with small pits similar to those described above, but there are
no teeth present in any of the specimens that I have examined.
The upper surface is somewhat striated at the anterior (?) ex-
tremity.
The ceratohyal is a broad and thin bone, concave at the pos-
terior end and somewhat irregular in outline at the anterior,
where there are two surfaces, for the hyo- and urohyals, re-
spectively. The bone seems to be striated, especially so toward
the extremities. Its length is 98 mm.
The quadrate is fan-shaped and very thin anteriorly. The
condyle is very convex and has a superficial resemblance to the
distal end of the mammalian femur. Extending upward from
this, along the posterior border externally, there is a prominent
103. Proc. Am. Phil. Soc. 1878, p, 179.
23—vi
346 University Geological Survey of Kansas.
ridge which continues upward to the superior border. The
groove for the symplectic seems to be very small.
One specimen shows the top of the skull with most of the
bones in place. The frontals are long, extending backward as
far as the pterotics externally. They meet in the median line
by a suture which is probably dentate, although this cannot be
determined with certainty. On each side there is a small post-
orbital process. The bones are beautifully sculptured above
with coarse sulci, which radiate from near the centers of each,
internal to the postorbital processes. The parietals are small,
meet each other in the median line, and are covered with mark-.
ings very similar to those on the frontals. The supraoccipital
is in a poor state of preservation, but probably entered but little
into the formation of the upper part of the skull. It seems to
have been projected backward for quite a distance and was de-
pressed, as is the rest of the top of the skull. The epiotics and
pterotics seem to be united on the side of that skull in which they
are preserved. The pterotics form prominent angles of the
skull, while the epiotic processes are not so prominent. The
two are covered with small pits and tubercles of bone. Just.
back of the skull there are portions of two other bones, which
may be parts of the hyomandibular and supratemporal. The
first of these presents two articular surfaces, one of which is.
extended outward from the rest of the bone, the two being sepa-
rated by a wide space somewhat similar to that found in the
hyomandibular bone of Hmpo. ‘The other is an irregularly
shaped bone and has an articular surface on the side next to
the skull. There are numerous other small bones, sculptured
in a manner similar to the bones of the top of the skull. Three
of these are joined together in a chain, and from their position
would indicate that they might form a part of the rim of the
orbital cavity. The remainder of these bones are small and
scattered along the top of the skull, indicating that this part
was covered with dermal plates. The orbital cavity is quite
large, and the orbit is surrounded by a very thin ring of bone,
the number of pieces composing which cannot be determined,
owing to their fragmentary condition. Just in front of the or-
STEWART. | Cretaceous Fishes. 347
bital cavity, and separated from it by the chain of bones just
mentioned, there is a bone that occupies about the position of
the ethmoid. It seems to be rather thin and crushed down on
the opposite side, so that its remaining characters cannot be
made out.
The opercular bones are thin and scale-like and are all in
place. They are all covered with fine striz and are so thin at.
the edges that the different directions that these strive take is
the only means of locating the boundary lines between some of
the different bones. The preoperculum is small, narrow above,.
and extended well forward below. The operculum is large and
is extended for some distance above its articulation with the
hyomandibular. The inter- and suboperculars extend back--
ward as far as the operculum.
Anogmius evolutus. Plate LXV, figs. 8,9, and 10; plate LXVII.
Anogmius evolutus Cope, Proc. Am. Phil. Soc. 1878, p. 179.
Beryx? multidentatus Stewart, Kans. Univ. Quart., vol. vir, p. 196.
This fish was first described by Professor Cope from an en-
tire left mandible, of which no figure is given, and I am in-
clined to think that it is the same form as was described by
myself as Beryx? multidentatus,'* the paper containing the de-
scription by Professor Cope*” having escaped my notice until
after my article was published. As Anogmius evolutus Cope has
priority, Anogmius multidentatus will have to remain a synonym,
unless there are other characters in Cope’s specimen not men-
tioned in his description that will separate the two forms.
About the only difference is in size, and I do not consider that
sufficient to separate them.
The principal points of difference between this species and
A. polymicrodus, just described, are found in the differences in
form of the symphyses, the extension of the teeth over the
external sides of the dentaries in this species, and the differ-
ence in form of the cotyloid cavities.
The dentary is elongated and slightly incurved at the sym-
348 University Geological Survey of Kansas.
physis. Thesymphysis presents but one articular surface, which
is roughened, and does not have an extension of the tooth band
overhanging it. In these respects the jaw bears a superficial re-
semblance to that of some of the recent siluroids. The symphysis
is slightly deeper than wide. The tooth band is broadest at about
the center and narrowest at the posterior extremity, where the two
borders of the band are suddenly drawn inward. The teeth
continue over the side externally, and the whole band is inclined
inward toward the center and posterior extremity. The dental
pits are small, and densely packed in somewhat irregular longi-
tudinal rows, there being forty or more of these rows at the
broadest portion of the band, which is ten more than Cope
found in his specimen. The articular is more robust through-
out than in A. polymicrodus, and the form of the cotylus is dif-
ferent, for, instead of being partially supported on a shelf-like
expansion of bone, as in this species, it rests directly across the
main part of the articular. Itis concave longitudinally, slightly
convex transversally, and is directed slightly backward instead
of forward, as in the species just mentioned.
There is a small portion of a maxilla present, and also a frag-
ment of another bone, which is probably a palatine, although
it differs somewhat from that of A. polymicrodus. The tooth
band of the maxilla seems to be more convex transversally than
in the species just mentioned, and there seems to be a total ab-
sence of the coarse striz which ornament the external and in-
ternal surfaces of the bone in this form. The supposed palatine
is a long and thin plate of bone which is covered with small
tooth pits on one side, which decrease in size toward one of the
extremities. At the end where the teeth are the largest there
is a projection which probably serves to bind the bone to some-
thing.
Below are given the principal measurements of the mandible
of our specimen and the one described by Professor Cope.
Cope. K.U.
Henethof tooth) band! ch. a oa en ae eee eee 150mm. 165.0mm.
Depth of symphysis: ... ac) ea1 eae aoe es 16 *§ 1957.0
Width-of sy mph ysis.2)..). 5.0. sunk oe erie eae. Diles Oy eee
STEWART. | Cretaceous Fishes. 349
SALMONIDE.
Pachyrhizodontide Cope, Proc. Am. Phil. Soc., vol. x11, p. 343.
In the year 1875 Professor Cope described his family Pachy-
rhizodontidx, to include the genus Pachyrhizodus, and which he
characterized as follows: ‘‘ This family of physostomons fishes
differs from the last, Sauwrodontide and Ichthyodectidx, in the na-
ture of its dentition. Instead of elongated conic fangs sunk in
deep alveoli, it has shorter and stouter fangs occupying alveoli,
of which the inner side and part of the antero-posterior walls
are incomplete. The teeth are, in fact, more or less pleurodont,
but the extremity of the root is received into the conic fundus
of the alveolus.
‘<The premaxillary bones are well developed, but the maxilla-
ries are more so, and enter largely into the composition of the
border of the mouth. There is a well-developed angle of the
mandible, but no coronoid bone is preserved in the specimens.
The coronoid region is, however, broken in all of our specimens.
The other characters of the family are not determinable from
our imperfect material.’’
In the ‘‘ Cretaceous Vertebrata,’’ published in 1875, Professor
Cope abandons the name Pachyrhizodontide and includes Pachy-
rhizodus in his family Stratodontidx, to which it evidently does
not belong. It seems to show some relation to the Salmonidz,
in which family it should probably be included, along with Or7-
cardinus.
PACHYRHIZODUS.
Dixon, Geol. of Sussex, p. 374, 1850.
The muzzle of this genus is flat and the bones of the skull
more fragile than in members of the /chthyodectide and Sauro-
dontide, consequently indicating a less powerful and rapacious
fish than those belonging to these families. The mandibles are
loosely united at the symphysis and bear a single row of teeth
which are somewhat pleurodont, sharply pointed, and bear a
superficial resemblance to some of the Mosasaurs. Their mode
106. 1. c., p. 343.
350 University Geological Survey of Kansas.
of succession, as described by Professor Cope, is as follows :'”
‘The crown of the young tooth was developed in a capsule at
the base of the crown, or on the inner side of the apex of the
thick root. The absorption which followed excavated both the
former and the latter; but the crown was evidently first shed.
Finally the old root disappeared and the new one occupied the
alveolus, leaving a free separation all round. Finally, on the
accomplishment of the full growth of the root, it became an-
chylosed all round.’’ The articular portion of the mandible
does not seem to be formed of the derm and antarticulars, as in
Saurodon, etc.
The premaxille are much more elongated and of an entirely
different form from those of the Stratodontide. They bear a
single series of teeth on the external side and two more larger
ones internal to the regular series. The maxille are rather
shallow and are not united with the palatine, as in the Ichthyo-
decide and Saurodontide mentioned above. There are several
other elements covered with villiform teeth whose position is
not known.
There is no complete skull preserved in the museum, but
Professor Cope says: ‘‘The cranium is flat and wide, and
pressure has probably somewhat increased the effect in this —
instance. Exoccipital, supraoccipital, epiotic, pterotic, parietal
and frontal bones are clearly distinguishable, but there are
points where the sutures are obscure. The best defined are the
epiotics, which are subtriangular bones presenting the apex
inward and bearing a small round facet for the supratemporal
on the posterior angle. The pterotics and postfrontals may be
easily distinguished from adjoining bones, but not so well from
each other. They have a thin outer margin, and their superior
surface is marked by bands of irregular small fosse and an
obtuse longitudinal ridge. The middle line of the skull is
occupied by the supraoccipital. Its proximal portion probably
separates the exoccipitals, but this is not certain. It extends
well forward, and the line of separation from the frontal is not
107. Cret. Vert. West, p. 220.
108. l. c., pp. 220a, 220b.
}
"
Srewart. | Cretaceous Fishes. 351
well defined. Its anterior part has a massive transverse eleva-
tion, which sends a short median process backward, producing
a T-shaped body; the frontal suture is probably in front of
this. The supraoccipital is contracted behind this body, and
its postero-exterior suture presents a remarkable peculiarity in
a straight and wide truncate articular face. This is opposed
by a corresponding face of the parietal bone. The latter is of
an irregular form, and carries on its outer portion next the
pterotic a stout protuberance. This is at the inner end of a
strong ridge, which disappears near the outer edge of the
pterotic. The protuberance looks as though adapted for an
articulation. The frontals send a process backward, between
the ‘supraoccipital’ and the pterotic or postfrontal, to the base
of the tuberosity of the parietal. The suture between the
exoccipital and parietal is not clear. A suture is distinct
enough, bounding the latter behind, but whether an expansion
of the supraoccipital intervenes or not is not certain. The ex-
occipitals appear to be flat and quadrant-shaped, having convex
antero-lateral borders. Each bears a strong condyle.’’
The vertebrie present no deep lateral grooves, but are longi-
tudinally striated and rather elongated back of the cervical
region.
The following species have been described from Kansas :
Pachyrhizodus kingi Cope, Niobrara Cretaceous, western Kansas.
Pachyrhizodus latimentum Cope, Niobrara Cretaceous, western Kansas.
Pachyrhizodus sheareri Cope, Niobrara Cretaceous, western Kansas.
Pauchyrhizodus caninus Cope, Niobrara Cretaceous, western Kansas.
Puchyrhizodus leplopsis Cope, Niobrara Cretaceous, western Kansas.
Pachyrhizodus leptognathus Stewart, Niobrara Cretaceous, western Kansas.
Puchyrhizodus velox Stewart, Niobrara Cretaceous, western Kansas.
Pachyrhizodus minimus Stewart, Niobrara Cretaceous, western Kansas.
Pachyrhizodus leptognathus., Plate LXIX, fig. 1.
Pachyrhizodus leptognathus Stewart, Kans. Univ. Quart., vol. vir, p. 193.
This species is represented by the left mandible, quadrate,
symolectic, preoperculum, and two broad, flat plates of bone,
which are probably the operculum and suboperculum. With
the exception of the anterior portion of the dentary, only the
internal sides of the bones can be examined, as they are too
302 University Geological Survey of Kansas.
fragile to be removed from the matrix. The catalogue number
of the specimen is 75.
The dentary is elongated, very slender, and bears eighteens
small, conical teeth upon its superior border, each of which is
set on a small, bony tubercle of bone, similar to those in P.
caninus. The teeth are directed backward and inward, decrease
in size from the center toward the extremities, and end pos-
teriorly within about an inch of the coronoid process. The
symphysis is of a more or less tubercular nature, similar in
many respects to that found in P. latimentum, and there are no
teeth arranged in a triangle here, asin P. caninus. The external
surface of the dentary does not seem to possess any very char-
acteristic markings.
The articular extends forward fully two-thirds the length of
the mandible, and is deeply grooved. The cotylus is concave
from before backward, and strongly convex laterally. The
quadrate is triangular in outline and thin above. Toward the
lower portion the bone thickens and assumes a somewhat
twisted appearance at the condyle. The condyle is somewhat
excavated at the center in order to fit the cotylus. On the in-
ternal side there is a well-marked ridge extending from the
condyle upward through the center of the bone. The superior
border presents a long, narrow sutural surface for uniting this
with the metapterygoid. Just back of the quadrate, and articu-
lating closely with it, there is a long, narrow element which
may be the symplectic. It is covered in part by the preoper-
culum, so that its characters cannot be made out.
The supposed preoperculum consists of a horizontal and a ver-
tical portion which meet each other at almost right angles.
The horizontal portion possesses no marked characters beyond
that it is the broader of the two parts. The vertical portion is
broad below, but soon becomes more narrow as the upper ex-
tremity is approached. Just back of the anterior border there
is a well-defined ridge extending downward to the junction of
the two parts. The two flat bones which may represent por-
tions of the operculum and suboperculum possess no characters
beyond those shown in the figure.
STEWART. | Cretaceous Fishes. 353
In addition to the above there is a tooth-bearing element that
may be a portion of the maxilla, but I think the teeth are too
small to be a part of this bone. The teeth are closely set upon
small, bony rostrums similar to those of the lower jaw. The
external surface is nearly smooth.
Mandible: Length of alveolar border...................... 110.5 mm.
menetin from :cotyloid: CAVIbY . 2... 2.6 is ee ec eee 135.0: §
Depth at coronoid angle (estimated)............ ZU) Ss
Number of teeth in one inch, 5.
2 OE ES ay OS SST ct ene ee 32.0 mm.
Benet Of SUpenOr POLGe? ck. Sees ee 2 AO a
Pachyrhizodus velox. Plate XLIX, fig. 2.
Pachyrhizodus velox Stewart, Kans. Univ. Quart., vol. vir, pp. 193-195.
This species is represented by a maxilla, both mandibles,
fragmentary quadrate, numerous portions of branchiostegal
rays, and a fragment of another bone that I first described as a
portion of one of the hyoids."” I have since reached the con-
clusion that it must be some other part, as the bone is too much
contracted back of the extremity to be a portion of a ceratohyal.
The catalogue number of the specimen is 316.
The maxillary is long and slender and of about equal depth
from the superior condyle backward. The condyle is elevated,
but how much cannot be determined, as the superior portion is
not preserved. Just beneath the condyle the bone thickens and
the outer surface contracts inward. External to the condyle
there is a broad shelf of bone which is very roughly striated.
The premaxillary surface is not preserved. The alveolar border
supports about forty-seven teeth, as near as can be estimated.
They are conical, directed slightly inward, and closely set. The
crowns present a smooth enameled surface. The whole of the
posterior portion of the bone is finely striated. Just above the
alveolar border in the anterior half of the bone there are many
small, nutrient foramina leading inward.
The mandible differs from that of P. latimentum Cope in not
haying a tooth on the symphysis within the anterior one, and
in haying a greater depth at the coronoid with reference to its
length.
109. Kans, Univ. Quart., vol. VII, p. 194.
304 University Geological Survey of Kansas.
The dentary is short and strongly incurved at the symphysis
The symphysis is divided by a groove into an external and in-
ternal portion. The external is small and tubercular in its
nature. The internal is probably the only part that is in con-
tact with its fellow on the opposite side, and it has a well-marked
ridge extending backward which becomes more indistinct toward
the posterior portion. This ridge causes the bone to be thickened
just below the alveolar border. The lower portion of the
dentary is thin and smooth externally, except on the lower
border, where there are short and deep strise extending back-
ward. The alveolar border supports probably thirty-eight or
forty teeth. ‘These are closely set, non-striated. and directed
inward. The external alveolar wall rises considerably above
the internal.
The character of the cotyloid cavity cannot be made out,
owing to the quadrate being firmly in place. The outer surface
of the articular is covered with striz, which become coarser
toward the lower portion.
The head of the quadrate seems to be broad and bifurcated,
as in P. leptognathus. Above the head the bone broadens ante-
riorly, and has a strong ridge extending upward along the pos-
terior border. Between the jaws,there are several pieces of
ossified cartilage covered with minute denticles, somewhat re-
sembling shagreen.
Maxillary: Length from premaxillary surface.............. 135.0 mm.
Depth of condyle (estimated)...........2...... 26:0 jars
Mandible: Length from cotyloid cavity. 4.2424 oo eee: ayes 26
Mensth of alveolar border) 44- he peer oe D200:
Depthyat coronoidtangle sees eee ee coe 48.3
Number of teeth in one inch, 8.
Hyoid (?): Distance across anterior end (estimated)........ 23.0 mm.
Pachyrhizodus leptopsis. Plate LXX, fig. 1.
Pochyrhizodus leptopsis Cope, Cret. Vert. West, p. 225, pl. 1, figs. 8a,
b, and c.
Represented by a portion of the right dentary bone of one in-
dividual and asmall portion of the left dentary of another.
The number of the specimen is 289.
The dentary is elongate and the upper portion forms a thick-
ee a. eee
.
|
|
4
eee
Srewakt. | Cretaceous Fishes. 355
ened bar with a broad superior surface. This bar is bounded
by a well-marked ridge antero-internally, from which the inner
wall descends almost vertically. The teeth are not closely set,
the interspaces being occupied by the sockets of the shed teeth ;
they are all set on stout bases and are larger than in P. leptogna-
thus. These bases vary from round to oval and the apices are
about on a level with the external alveolar border.'” ‘‘ The
teeth diminish in size from the middle of the dentary bone to
the symphysis; beside the latter are two teeth of reduced size,’’
but on the posterior portion the teeth are but slightly reduced.
Just back of the last tooth there is an edentulous portion of
about an inch in length, which is followed by a somewhat ele-
yated coronoid process. ‘This process is bent outward and covy-
ered with irregular longitudinal striz on the external and
internal sides. The lower portion of the bone, below the alveo-
lar border, is thin and laminiform, smooth externally, and cov-
ered with coarse striw internally." ‘‘ The symphyseal part of
the ramus is not incurved as in P. caninus and P. kingi but is
obliquely truncate, indicating that the chin had a compressed
form, and was not rounded, as in them.’’
MEE aE RHINE G NG DOOENS oo adh o!- wou eee a vne cae eee es 60 mm.
NEES OE TMNER SAS 6 Sg ins yl gia inly wie hd eia Ss «yas a wits io
Distance from tip of coronoid to last tooth.................... tos
Number of teeth in one inch, 2.5 to 3.
Pachyrhizodus caninus. Plate LXX, figs. 3, 4,5, and 6.
Pachurhizodus caninus Cope, Proc. Am. Phil. Soc., vol. x11, 1872, pp. 344
—346.
This species is represented by portions of the premaxilla and
mandibles of twoindividuals. There is also a flat plate of bone
accompanying the mandibles whose identity cannot be deter-
mined. The numbers of the specimens are 145 and 315,
The teeth on the dentary are supported on a shelf of bone
that becomes narrow toward the distal extremity. This por-
tion ends abruptly below and is continued into an acute lower
border. The jaw is probably shallow throughout its extent,
and toward the symphysis the bone is incurved and swollen.
110. Cret. Vert. West, p. 226.
111. l.c., p. 225.
356 University Geological Survey of Kansas.
The symphyseal surface is of an irregular form, slightly convex,
with an emargination entering from the internal side. The
jaws were probably loosely united by cartilage at this point.
The distal extremity is edentulous, and forms a rather thin
coronoid process, coarsely striated internally. There are spaces
for from twenty-nine to thirty-two teeth, which decrease in size
toward the extremities, and at the symphysis a triangle is
formed by three of these. The teeth are rather closely set in
places, with enlarged striate bases, while in others the teeth
are separated quite a distance from each other where the head
crown has not been replaced. The external alveolar wall does
not rise as high as in P. leptopsis, mentioned above.” ‘‘The ar-
ticular cotylus is composed more largely of the angular than
the articular. Its long diameter extends inward and backward,
and is strongly convex; in the transverse direction, slightly
concave. Below and in front of it the lower margin of the jaw
is acute. The angle is oval and rather small; it is prominent
on the middle line on theinner side; the edges are thin, the
upper curved outward, concealing part of the cotylus.”’
The premaxilla is elongated, and in the posterior portion of
the bone it is narrow transversally. Toward the anterior ex-
tremity it becomes much wider and presents alveoli for two
large teeth situated internal to the regular series. It is likely
that only one of these teeth is functional at a time, as in most
of the specimens that I have examined I find only one tooth,
and an empty alveolus for the other. The surface for its fellow
on the opposite side is roughened by small protuberances of
bone. The teeth are slightly larger than those upon the den-
tary, but this may be due to the difference in size of the two
specimens. There are four preserved on the outer row, but
there were probably several more, as Professor Cope describes
ten in his specimen. The external surface is finely striated.
The plate of bone mentioned above shows one surface to be
covered with small pits except at the central portion, which is
smooth. These pits at first remind one of the empty alveolus
on the tooth-bearing elements of Anogmius and some of the
112. Cert. Vert. West, p. 222.
STEWART. | Cretaceous Fishes. 357
recent Siluridx. The opposite surface is irregularly marked,
and one of the extremities is convex.
Length of alveolar border of dentary...................... 174.5 mm.
Seeeia RAT CRIN ET URIS ERI VEMET 2 OR Weta SSS orn ciate cg Kura s wrens RRS Ww alse EROe
Number of teeth in one inch, 5.
Pachyrhizodus latimentum ?. Plate LXVIII; plate LXX, figs. 9, 10.
Pachyrhizodus latimentum? Cope, Proc. Am. Phil. Soc. 1872, p. 346.
This species of Pachyrhizodus is the largest American member
of the genus, and probably outrivaled some of the largest forms
of Ichthyodectes. The specimen was found by myself in the
Butte creek region, in Logan county, Kansas, during the summer
of 1898, and consists of a portion of a disarticulated skull, the
glenoid portion of a scapula, anda number of vertebrie. The
catalogue number of the specimen is 316.
Both the extremities are broken away from the maxilla, so
that the exact length of this bone cannot be determined. It is
finely striated externally and internally, and is somewhat
coarsely striated on the superior surface. The teeth are large |
and closely set, but they are not all functional at once. The
bases of these are somewhat striated and hidden from view ex-
ternally by the downward extension of the outer alveolar wall.
The teeth probably end some distance from the posterior ex-
tremity, as there is an edentulous portion back of the last tooth
that continues as far as the alveolar border is preserved in this
direction. The greatest depth of the fragment is 35 mm.
The premaxilla is more slender and pointed anteriorly than
in P. caninus. There are seventeen teeth and tooth scars on the
outer border, and there are spaces for one or two more that
show no indication of ever having borne teeth. The teeth seem
to be very similar to those found on the maxilla, with the ex-
ception that they are slightly smaller. The external border is
nearly straight, while the internal is beveled off toward the
anterior end, giving the extremity a slightly blunted appear-
ance. Internal to the regular series there is a tooth that is
much larger than those in the external row which has a deep
a
358 University Geological Survey of Kansas.
pit in front of it, probably the empty alveolus for the second.
large tooth. Measurements of premaxilla are:
Length’ of alveolar bordeng 7, Mase as cen okies ee eee 107 mm.
Greatest: transverse diameter $c3 6 s5 oss Wiesel oa DAD ess
Greatest depth (estimated) os. oe), Mee ere ee ae BO We
The mandible seems to be slightly larger than the one de-
scribed by Professor Cope. Only the posterior portion is pre-
served, showing the articular and a portion of the dentary,
including several teeth. The dentary is coarsely striated ex-
ternally and has a rather indistinct groove just below the alve-
olar border. The teeth are probably not much reduced toward
the coronoid process and their bases are partially hidden on the
outside, as in P. leptopsis. The last tooth is separated from the
coronoid by a long, thin crest of bone which is somewhat ele-
vated. ‘The exact line of separation between the dentary and
articular cannot be traced, probably owing to the fact that the
outer portions of the bone are injured and broken away in some
places. The cotylus is large, deeply concave from before back-
ward, and convex laterally. On the external side posteriorly
there is a wall of bone projecting upward that forms a slight
groove with the external side of the cotylus and receives the
outer rim of the quadrate. Just above and in front of the
cotylus there is a shailow pit, which seems to be not so well
marked as in the specimen described by Professor Cope. The
angular is a small element, and does not enter into the forma-
tion of the cotylus, asin P.caninus. It is irregularly marked
and has the appearance of being partially anchylosed to the
articular. The posterior extremity shows a roughened surface,.
which is probably for muscular attachment.
Length of ramusi(estimated,)) o.2 ee ee ee eee 330 mm.
Distance from cotylus to last posterior tooth................. LS owes
Depth at coroneid angle (estimated)i.s. --4--2..--- ase ee Sane
Depth: at coty Lassa 5 sg oete ee Rae e artei aen c aon os
The quadrate is a rather long bone, which is probably not.
very greatly expanded above. ‘The condyle is deeply concave
transversally, with sharp edges both externally and internally.
The anterior portion is thin above the condyle, while the pos-
terior portion is much thickened and has a well-marked ridge
STEWART. | Cretaceous Fishes. 359
extending upward on the external side. The superior border
is badly preserved.
IEEE CAL RRENE EEL ee vctr ieee PG Oe a tule ee ok Lio wiw es oe 101 mm.
earsverse GIAMOter OF CONGYVIO .. . 5. os. ex cals ck eee dee eens 24
There are several other tooth-bearing elements, the exact
location of which cannot be determined. ‘Two of these are
long, slender bars of bone covered with denticles on one side,
somewhat resembling shagreen. Another is more flat than the
above and covered with somewhat larger denticles. Still an-
other is very thin and covered with various-sized denticles on
one side.
The basioccipital is separated from all of the surrounding
bones. The condyle is deeply concave and broader above than
below. Just in front of the condyle there are two lateral facets
for the attachment of the exoccipitals, which are much rough-
ened and look outward and forward. In front of these facets
the bone is narrow and has a median ridge, which is continued
downward on the anterior end. On the lower portion the bone
has a deep carina, which extends forward to near the extremity
and forms a sharp projection posteriorly. There is a slight
groove in the median line that Professor Cope'’’ thought repre-
sented the muscular tube.
Transverse diameter of condyle above.......................+. 39 mm,
0 SPE per OU ag 26 oo *
eeu GE CSEL IS AETONE AIEEE, fra Wins oe 5 2.5; gM mvs wv, dns 2 Rede oe eas we ay ee
There are several other bones present whose location cannot
be determined, although they are probably located in the pos-
terior part of the skull. Figures of these are shown on plate
LXX. There are a number of vertebre with the specimen,
which are larger than in the largest specimens of [chthyodectes
that I have examined. The sides are devoid of lateral grooves,
but are covered with numerous small ridges which inosculate
with each other, giving the outer surface of the vertebra a some-
what lace-like appearance. In this respect the vertebrie have
a superficial resemblance to those of Anogmius, but when they
are examined closely it is found that the ridges are more irregu-
113. Cret. Vert., p. 220a.
360 University Geological Survey of Kansas.
lar and more sharply defined, especially at the extremities of
the centrum. The neural arches probably fit into pits above,
and the ribs seem to have been attached to tubercular processes
of bone set into pits, as in the Ichthyodectidx. Those in the an-
terior region are very thin, and the pits for the neural arches
are large and widely separated.
Nearly the whole of the scapula is preserved. It shows a
large convex superior (?) facet with a smaller one just below it,
the two being separated by a sharp ridge. Just back (?) of the
larger facet there is a large foramen, the anterior border of
which is broken away. :
There is a second specimen of a shoulder-girdle and a mass
of fin-rays which probably belong to the same genus as the
above, but probably a different species. The chief difference is
found in the absence of the small facet on the articular portion
of the scapula. At the lower extremity there is a large rough-
ened surface that probably unites with the coracoid. The
coracoid (?) is a broad, flat plate of bone which is thickened
at the posterior extremity and presents a roughened articular
surface for the scapula. The cleithrum is too fragmentary to
determine its exact form, but it was probably concave on the.
internal (?) side and somewhat irregular and roughened on the
external.
The proximal portions of the fin-rays of both sides are pres-
ent. They are large and very numerous, fourteen being pres-
ent on one side, and this probably does not represent all of
them. The larger of these are slightly bent at the proximal
extremities, and the smaller ones strongly so.
Lower jaws of Pachyrhizodus minimus, one-half natural size.
pelt een
StTewakrt. | Cretaceous Fishes. 361
Pachyrhizodus minimus.
Pachyrhizodus minimus Stewart, Kans. Univ. Quart., vol. viir, pp. 37, 38.
This species is represented by the mandibles alone, which are
characterized as follows: The dentaries are slender and slightly
incurved at the symphysis, where they also seem to be bent
slightly downward. The symphysis is bifurcated by a deep
groove on the external side, but internally it is continuous from
above downward. On the external side there is a shallow groove
just below the alveolar border, which becomes indistinct toward
the symphysis ; the lower border of the bone is sharp and nearly
straight. There are eleven teeth preserved upon one of the
rami, and room for at least as many more. They are conical,
non-striated, and directed inward; the last is situated quite a
distance from the coronoid angle, below which there is quite a
depression.
Only a small portion of the articular is exposed on the ex-
ternal side, but internally it extends well forward. The cotylus
is very convex transversally and situated well up toward the
coronoid process, while below and extending backward from it
on the external side is the prominent hook-like, angular process
found in this and other genera of fishes.
Length of alveolar border from coronoid angle........ ..... 52.0 mm.
wo VEER DIG OR RG Se ae a pee 40.0" 2*¢
meurPtn or Done Irom Cotyloid Gavily..:........ 61.6. .ss..... B60 *
Number of teeth in one centimeter, 4.
The specimen, No. 327, was found by myself in the Butte
creek region of Logan county, Kansas, during the summer of
1898.
24—vi
362 University Geological Survey of Kansas.
PACHYCORMIDAE.
Pelecopteride Cope. Hrisichtheide Cope.
Protosphyrenidce Woodward.
This family is very peculiar in having the ethmoid bone pro-
longed into a long rostrum or beak, and includes the single
American genus Protosphyrena. For along time this form was
not well understood, as Professor Cope described the pectoral
fins as belonging to the genus Pelecopterus, and the cranial por-
tions to the genus Hrisichthe, from which arose the family names
of Pelecopteride and Hrisichtherdx. It was later discovered that
these two genera were one and the same, and also that they
were synonyms of the genus Protosphyrena, which was described
by Leidy several years before.
PROTOSPHYRANA.
Protosphyrena Leidy, Trans. Am. Phil. Soc., vol. XI, pp. 91-95.
Xiphias Leidy, Trans. Am. Phil. Soc., vol. XI, pp. 91-95.
Erisichthe Cope, Proc. Acad. Nat. Sci. Phil., 1872, p. 280.
Pelecopterus Cope, Cret. Vert. West, p. 244 C.
"<The teeth are laterally compressed and lanciform, with-
out marginal serrations, and, when found in connection with
the jaw, are seen to be fixed in distinct sockets. They were first
noticed in 1822\by Doctor Mantell,'” who described one example
as belonging to an unknown fish, and referred a second to an
undetermined species of shark. ‘Twenty-one years later Profes-
sor Agassiz studied the fossils, and thought that they were most
probably identical with certain teeth discovered by Doctor Har-
lan'*® in the Cretaceous strata of North America, which had been
described under the name of Saurocephalus lanciformis, and
erroneously placed among the Reptilia. They were accordingly
made known as such in the great work on the ‘‘ Poissons Fos-
siles,’’!” and the determination was subsequently adopted by
Dixon,"™*® in 1850, who figured more satisfactory specimens. At
114. A. S. Woodward, Proc. Geol. Assoc., vol. X, 1888, pp. 320, 321.
115. Foss. South Downs, p. 227, pl. XX XIII, figs. 7-9.
116. Jour. Acad. Nat. Sci., Phil., vol. III, 1824, pp. 331-337, pl. XII, figs. 1-5.
117. Poiss. Foss., vol. V, pt. I, p. 102, pl. XX Ve, figs. 21-29.
118. Foss. Sussex, p. 374, pl. XXX, fig. 21; pl. XXXI, fig. 12; pl. XXXII, fig.1; pl. XXXIV, fig. 2.
> {= ——_ —
.
STEwakrt. | Cretaceous Fishes. 363
this time, moreover, an important addition to our knowledge of
the species was rendered possible by a fine fossil in Sir Philip
Egerton’s collection, which showed that the fish had a remark-
ably elongate snout. Six years afterwards Professor Leidy'”
carefully reexamined Doctor Harlan’s original American speci-.
mens of Sauwrocephalus, and soon became convinced that Agassiz.
was mistaken in supposing that the English fossils were identi-
cal with these even generically. He thus proposed a new ge-.
neric and specific name—Photosphyrxna ferox—for the specimens
figured by Mantell, Agassiz, and Dixon; wrongly suggesting,.
however, that Sir Philip Egerton’s fossil rostral bone truly be-.
longed to a swordfish, which might be appropriately termed
Niphias dixoni. Between 1875 and 1877 remains similar to those
of the English Chalk were actually discovered in America, and
not only proved that Egerton and Dixon were right in deciding:
upon the reference of the long snout to Protosphyrena, but.
further added important anatomical details. Professor Cope,
however, who described these fossils,’ proposed the difficultly
pronounceable name of Lrisichthe, which happily becomes a
synonym.’’
The frontals of this genus are broad and flat, and are more
or less covered with short corrugations. Anteriorly they unite
with the ethmoids, which, with the prefrontals, vomer, and
parasphenoid, form the long and pointed rostrum mentioned
above. The frontals form the superior boundary of the orbits,
which are large and surrounded by aring of sclerotic bones.
The pterygoids are covered with small, conical teeth, thickly
set, and elevated upon small, bony hemispheres. The meso-
pterygoid is a broad and thin bone, covered with teeth similar
to those found on the pterygoid. The upper jaw is composed
of the maxilla and premaxilla. The first of these is slender
and bears two rows of teeth, the inner of which is composed of
large teeth with anterior and posterior cutting edges, each of
which is set in a separate alveolus. The outer row is composed
of small, conical teeth united directly with the bone. ‘The:
119. Trans. Am. Phil. Soc., vol. XI, 1860, pp. 91-95.
120. Cret. Vert. West, pp. 217, 218, and Bull, U.S. Geol. Surv. Terr., vol. III, 1877, pp. 821-823.
364 University Geological Survey of Kansas.
smaller of these teeth are mere tubercles of bone. The pre-
maxillaries are small, triangular elements, loosely united to the
maxilla. They bear both large and small teeth, of which the
large are sharply pointed, smooth, and have anterior and pos-
terior cutting edges. The predentary is paired, and bears both
large and small teeth.
The fins of this genus were first described as the dorsal fin-
spines of a shark, Ptychodus, by Agassiz,’' but were later recog-
nized by Cope’ as belonging to a teleost, and described by him
as a new genus, Pelecopterus, indicating a new order, Actinochiri, «
and family, Pelecopteride. These fins seem to form powerful
spines, and are composed of slender, parallel rods of bone,
closely placed. The anterior border is usually undulated more
or less, and, in the smaller species, with tooth-like processes
which are often covered with a substance which closely resem-
bles enamel. Remains of this genus are found in the Fort
Benton, Niobrara and Fort Pierre groups in Kansas.
There seems to be some doubt concerning the structure of the
Jower jaw, as the two authors, Cope and Felix, who have writ-
ten upon the subject, seem to entertain somewhat different
opinions. Cope™ says: ‘‘A remarkable feature of the genus
is displayed in the mandibles. Hach of these is compound in
the region usually composed of the simple dentary bone. It
there consists of three parallel elements—an internal and an
external, embracing amedian element. The inner bears a band
of teeth en brosse on its inner and superior aspects, and ‘the ex-
ternal a few teeth of similar character on its superior edge.
The large, lancet-shaped teeth are borne by the middle element,
excepting some of the largest near the symphysis. ‘Two of these
on the inner side of the ramus originate in the internal bone.”’
Concerning this point Felix™ says:
“Wenn wir auch den Unterkiefer, abgesehen von dem Articular
und Angular, aus vier Stiicken gefunden haben, dem Preedentale und
Dentale, sowie einem vorderen und einem hinteren Spleniale, so
k6nnen doch dies nicht die Elemente sein, welche Cope meint.
121. Poiss. Foss., vol. III, pp. 56-59. 123. Bull. U. S. Geol. Surv. Terr., p. 821.
122. Cret. Vert. West, pp. 244, A-F. 124, 1. c., 287.
Se
SrEwart. | Cretaceous Fishes. 365
Denn das hintere Spleniale reicht vorn nur bis an das hintere Ende
des vorderen Spleniale, nirgends liegen drei Elemente parallel neben
einander, keins yon ihnen kann ‘middle element’ genannt werden.
Vergleicht man mit der citirten Beschreibung der friiher von ihm
gegeben Abbildung”? des vorderen Mandible-Theils, so muss man es
auch befremdlich finden; dass Cope die grossen Fangzihne, mit
Ausnahme allerdings von zwei, welche er in dem inneren Knochen
entstehen lasst, von dem mittleren Theil getragen werden lisst, da
dieselben dem dusseren Rande so nahe stehen, dass zwischen ihnen
und jehnen nur eine sehr diinne Knochen-lamelle gedacht werden
kann. Was sollte diese aber in osteologischer Hinsicht darstellen ?”
Protosphyrena bentoniana.
Protosphyrena bentonia Stewart, Kans. Univ. Quart., vol. vit A, pp.
6
5 =U
This species was established upon the rostrum and numerous
fragments of other bones which were, for the most part, in too
poor a condition for identification. They were found by Dr.
S. W. Williston in the Lincoln marble, on Rock creek, in south-
ern Mitchell county, Kansas. The low horizon from which the
specimen was obtained attaches special interest to it, as it is the
first time that a specimen of this genus has been found below
the Niobrara Cretaceous in America.
The proximal portion of the rostrum, bearing the large teeth,
is not preserved. The base is broad, becoming more narrow
toward the distal extremity, where it suddenly contracts, forming
a rather blunt apex. The anterior portion is oval in outline in-
stead of semicircular or round. The lower surface contracts
more rapidly than the upper, causing the apex to be slightly
aboye the center of the shaft. The superior and inferior sur-
faces gradually grade into each other, and are not separated by
the angular ridge found in some of the species of this genus.
The outer surface, where preserved, shows the rostrum to be
covered with irregular longitudinal ridges which inosculate
125. Cret. Vert. West, pl. XLVII, fig. 6a.
126. Through a typographical error, this species first appeared as P. bentonia instead of P.
bentoniana, as was intended.
366 University Geological Survey of Kansas.
more or less with each other, and give the outer surface of the
bone a somewhat net-like appearance.
Meneth of rostral fragmenta .e asso ck ee ee eee eee ate oO ma
Transverse diameter 136 mm. from distal extremity .......... 34%
Transverse diameter 22 mm. from distal extremity ........... Pipa ie
Protosphyrena recurvirostris.
Protosphyrena recurvirostris Stewart, Kans. Univ. Quart., vol. vir 4,
pp. 191, 192.
The material upon which this species was based consists of a
complete rostrum, No. 373, with the adjoining portions of the
vomer and parasphenoid, and differs from P. nitida in the fol-
lowing characters: The superior distal surface is regularly
rounded and not flat, as in that species, and the cross-section at
this point is round instead of semicircular or oval. The speci-
men corresponds in some respects with the rostrum of P. pene-
trans, and is more closely related to this species than to any of
the other forms that have been described. I was inclined to
it call P. penetrans until I found a rostrum of that species, when
I discovered that it differed from it in a number of points which
were sufficiently great to be called specific. The rostrum is more
slender as a whole and is contracted to a more acute point than
in P. penetrans. The markings are more sharply defined and
the ridges inosculate with each other but rarely. In P. pene-
Rostrum of Protosphyrena recurvirostris, one-half actual size.
trans the markings are more or less reticulate. In the anterior
portion of the species under consideration the ridges are closely
placed to each other, while posteriorly they become scattered and
are not so well marked as in the anterior portion of the bone.
Their direction also becomes more varied in this region. On
the posterior half of the inferior surface, the ridges become less
eager je
STEWART. | Cretaceous Fishes. 367
numerous and are larger than those on the superior surface
and on the sides. In P. penetrans there is no difference of
markings on the superior and inferior surfaces. <A part of one
of the large teeth at the base of the rostrum is preserved, and
presents a smooth, enameled surface and probably anterior and
posterior cutting edges.
A point that I have noticed is, that all of the figures and
specimens of this portion show only one tooth, the alveolus for
the other seeming to be filled with bone or matrix. This would
lead to the belief that these teeth were alternately functional.
The rostrum, as a whole, is slightly recurved.
Range and distribution: Niobrara Cretaceous, Gove county,
Kansas.
Protosphyrena gigas.
Protosphyrena gigas Stewart, Kans. Univ. Quart., vol. viri A, p. —.
The material upon which this species is based was obtained
from the Lisbon shales, Fort Pierre Cretaceous, one mile north-
east of Lisbon, Kan., during the summer of 1898. The specimen
consists of the distal portion of a pectoral fin-spine, and is inter-
esting in showing that this genus persisted into the Fort Pierre
group. As only the distal portion of the fin is preserved the
length of the fin cannot be determined, but judging from the
width near the middle portion of the fragment it must have
been larger than P. gladius Cope, which makes it probably the
largest member of this family. The catalogue number of the
specimen is 338. ,
The spine is made up of parallel rods closely placed. Near
the center of the fragment there are about twenty-five of these,
and probably more in life, as the posterior border is broken
away and the exact number cannot be determined. ‘These are
broad in the central portion and become more narrow toward
the distal extremity, where the spine seems to be regularly
rounded on the end. Toward the anterior border the rods be-
come much thicker and the border forms a rather blunt cutting
edge, which differs from P. gladius, in which this margin is
acute. This border is somewhat irregularly notched, the notches
568 University Geological Survey of Kansas.
seeming to be more pronounced than in the species just men-
tioned, and are covered with an enamel-like substance.
P. gigas. P.gladius.*
Total lenethiof iracment yy ies eee nee AQ .Osmamayiah ve eeaeee
Width near the wmiddileri a wits eek. conienas 195.0 * 175 mm.
Thickness, posterior, near the middle............ tal on cee 2, see
Thickness, anterior, near middle................. Pa Oe OMNI Cees
Protosphyrena, 0. sp.?
Protosphyrena, sp. nov., Kans. Univ. Quart., vol. vir a.
The material upon which this description is based consists of
a right premaxilla from the Niobrara Cretaceous of western
Kansas. Locality and collector not known. This material |
consider too scant to justify a specific name being given until
more complete specimens are found. It may prove to be the
premaxilla of one of the other species of Protosphyrxna that are
at present known only from the rostral bones and fin remains.
At present it will have to be considered as a new species with
an interrogation. It is evidently different from the premaxilla ©
of P. nitida described by Felix.”
When seen from the side the bone is triangular in outline,
with a thin superior and posterior border. The anterior ex-
tremity is broken away, but it was probably acutely pointed, as
in P. nitida. The principal differences which characterize this.
from the premaxilla of the species just mentioned are found in
the size and arrangement of the teeth. Near the center of the
bone there are alveoli for four large teeth, the anterior three of
which are almost complete, while the posterior one is. broken off
* Measurements after Cope. Cret. Vert. West, p. 244 F.
127. Zeits. d. Deutsch. Geol. Ges. 1890, vol. XLII, p. 283.
eee
STEWART. | Cretaceous Fishes. 369
at the base, but enough remains to show that this and the an-
terior one were the smallest of the four. The two in the center
are of about the same size; they are all directed forward, the
anterior slightly more so than the rest. These teeth all have
broad lanciform crowns, with anterior and posterior cutting
edges and slightly striate enameled surfaces. Just back of the
most posterior of these large teeth there is a row of small teeth,
the anterior of which are scarcely more than bony tubercles,
but posteriorly they assume definite dental characters; ten of
these are preserved inthe specimen. There are other teeth like
these on the anterior portion of the border, but owing to the tip
being broken away, the exact number of these cannot be deter-
mined. The surface for the maxillary contracts toward the an-
terior end, and is bounded below by a narrow shelf of bone
extending inward.
Approximate length of the alveolar border .................... 82 mm.
Reem unG DACK OF IASh ATSC COOUR. oo cic etl es cee ees ile tes
MMA aE MATOS DOWN. lec ee Vis x tfete ns nk ce we ecw eevee bu eee fat ys
SERRE OEE HIIULCL AMEE, LODHI fone <.c. Wvie's ss 201s se eye eitiety ete eu lee oe We? ass
Protosphyrena penetrans. Plate LXIII.
Protosphyrena penetrans Cope, Bull. U.S. Geol. Surv. Terr. 1877, vol.
11, No. 4, p. 821.
This species is represented in the museum by only one speci-
men, No. 372, consisting of a rostrum with a small portion of
the apex, and the basal portion with the large teeth absent.
Sa
SOOO SSS
a
—
SSeS
SS
Pho) SR
Cd
Rostrum of Protosphyrena penetrans, one-half natural size.
The superior distal surface was probably rounded in life, but,
owing to depression, it has become slightly flattened, giving the
cross-section at this point a somewhat ovoid or elliptical outline.
In P. nitida this surface is finely rugose, with a strong angle
bounding the superior plane on each side, while in this species
there is no such angle, and the markings on this surface vary
370 University Geological Survey of Kansas.
but little if any from the markings on the front and sides of the
bone. The markings on the inferior surface are similar to those
on the sides, and do not become larger and more scattered, as
in P. recurvirostris. The bone is pointed at the apex, giving a
more acute shaft than in P. nitida, in which the rostrum con-
tracts suddenly to a tip, but less acute than P. recurvirostris,
where the shaft is more slender and the apex drawn to a more
slender point. The markings are not so sharply defined and
inosculate with each other more often than in the species just
mentioned, especially on the anterior portion and sides. Nio-
brara Cretaceous, western Kansas. Collector, E. P. West.
There are remains of several pectoral fin-spines, and an-
other which may be the pelvic fin, that I will refer to this
species with doubt. They are much smaller than those de-
scribed by Cope as ‘Pelecopterus,’ and also differ from these in
other minor details. Doctor Crook mentions in his paper on
the ‘‘ Kansas Cretaceous Fishes’’* that he had aright and a left
pectoral fin-spine of this species in his possession at the time of
preparing his paper, but failed to describe them, which makes
the determination of the remains before me very difficult.
The fin is composed of parallel rods of bone, which are very |
slender at the posterior side, but toward the anterior border
they become broader and slightly curved. The anterior
border is nearly straight for a distance of 45 mm., when it
begins to assume an undulated appearance, which becomes
less marked and farther separated beyond the center, and prob-
ably finally disappears before the extremity is reached. In P.
nitida these undulations assume a dentate appearance, and con-
tinue thus to the extremity, while in P. gladius and P. chirurgus
this border is devoid of such undulations. This portion is cov-
ered with a thin layer of enamel-like substance, which seems to
be thickest on the apices of the undulations. The spine is
moderately broad at the base, where there are forty or more
rods, but becomes more narrow distally. There are portions of
probably seven basiosts, the anterior ones of which are probably
paired. These are all provided with well-rounded condylar
128. Uber ein. Knoch. fis. a. d. mitt. Krei. Kans., Paleontographica, 1892, 110.
STEWART. | Cretaceous Fishes. 871
heads where they articulate with the fin, and they probably
have similar expansions at the other extremity.
P.agilis. P.nitida. P. gladius.
Length of spine fragment.............. 360mm. 705mm. 1,040 mm.
OE a ee eee ae er oe aa ss Gr”? ee *
MEMGRE WG DANO. sou ec cave hed was aE Set Bye EAs va
The spine mentioned above, and thought to be pelvic, is about
the size of the one described above, but differs from it in having
the anterior border nearly straight, without the undulations,
and but very little of the enamel-like deposit mentioned above.
At the base of the spine there is a mass of bone which presents
a large, flat articular surface, above and in front of which there
is a large, pointed process of bone extending upward from the
base of the anterior border. The spine is probably more narrow
than the one just described.
a a eR Os ee a 428 mm.
IRENE EROS cient wets ok ok eet Pea uivees etic ue ote f 45 *
EAR Rs PE CSLTEMREOEL) ocd reinc bs vx eae des kev see cea ad
ime tiemetor or Condyle at base. ..... 2... 2.2... eee eee es alee.
Puamre tuamoter Of GCondyle af base... 2.6... eee cee ce ale teen A
I am unable to say to what species the pelvic fin-spine, de-
scribed above, may belong. It is of a different individual from
the pectoral spine described as belonging to P. penetrans, but
for the present, at least, I will consider it under this head.
CLUPEIDA.
The family Halecidx’’ was made by Agassiz to include the
two families Salmonide and Clupeidx, fossil specimens often
being so closely related to both of the above families as to ren-
der their exact relation to either of the above two families some-
what doubtful.
The tendency among ichthyologists seems to be to revive the
two old family names; thus I use the name Clupeidx to include
the single genus Leptichthys, which seems to bear a close rela-
tionship to this family in many respects.
129. Am. Geol., vol. XXIV, pp. 78, 79.
372 University Geological Survey of Kansas.
LEPTICHTHYS.
Stewart, Am. Geol., vol. XXIV, pp. 78, 79.
There are several specimens of Cretaceous fishes in our col-
lection that seem to differ from any of the other genera that I
have seen described, and for which I have ventured to propose
the generic name of Leptichthys. This genus seems to be related
to Osmeroides Agassiz in some respects, but differs from it in the
presence of a short dorsal fin and non-striated scales. It also
differs from Apsopelix Cope in the anterior position of the dorsal
fin.
The body is ellipsiform, and covered with large, thin cycloid
scales, which are concentrically striated ventrally, but seem to
lose this character to some extent in the dorsal region, where
they are somewhat rugosely marked. The pelvic bones seem
to be formed of heavy transverse bars of bone, which are thick-
ened externally for the attachment of the pelvic fin. These
bones probably had some anterior extension, but how much
cannot be ascertained from the specimens at hand. The dorsal
fin is short and composed of numerous rays, which are situated
in front of the middle portion of the body. The pectorals are
broad and rather elongated. The pelvic fin is broad and short.
The caudal fin is deeply cleft. The skull seems to be rather
bluntly pointed in front. The dentaries are short with acutely
pointed teeth, and the lower borders of the bones are directed
upward in front. The vertebre are two-grooved, and are longer
than deep. Only one species is represented in our collection,
which is described below.
Leptichthys agilis, sp. nov. Plate LXXII, fig. 1.
Scales large; about ten or twelve series to be seen across the
body. The pectoral fins are composed of at least fourteen rays,
the first two or three of which are cross-segmented, while the
remainder are longitudinally split. The pelvic fins are situated
posteriorly and the cross-segmentation is absent from their rays.
The dorsal fin is composed of twelve or more short bony rays.
STEWART. | Cretaceous Fishes. 373
Length of body to cleft of caudal fin...... Ghee US ee Pes 290 mm.
Length from anterior extremity to base of dorsal fin.......... 112
Depth of body at dorsal fin....... Re RRNA Rae nic Oe ie og 6s
Been SNDE TRE STIIMANO SATE fc rc cies dak. vue wpe aw ee ew ae Gn wes 46°88
Menara OE DOOLGNAL TE) BU DHS. . 26 0s oon ees Suisse ne dees ews Tee.
MMMM CED AUOUG MOMMIES So Ge Ci phic nwo Meek VS Os ap cee ees pS
SC ces SA Gh: ag a 10%. *
Genet RSLS TIREAUICINISUAPELE TATE, G0 She aly hw 6 sh oa a eg ba Bebiens 8 FiWing OEE
ENCHODONTID&.
This family may be characterized as comprising fishes with
laterally compressed bodies, either naked or covered with scutes.
The maxille and premaxille are elongated and covered with
small teeth, which are firmly anchylosed tothe bone. The den-
taries are provided with a single row of large teeth, and usually
one or more rows of smaller ones on the external side. Pala-
tines and ectopterygoids powerful, and bearing a single series
of large teeth on expanded bases.
It is probable that the only representative of this family in
the American Cretaceous is the type genus Lnchodus. There
are several other European and Asiatic genera belonging to this
group of fishes, among which are Hurypholis, from the Lebanon
beds of Syria, Pomognathus, from the Chalk of Europe and Leb-
anon, and Jschyrocephalus, from the Upper Cretaceous of West-
phalia.
ENCHODUS.
Agassiz, Poiss. Foss., vol. V, pt. I, p. 64.
Leidy, Rep. U.S. Geol. Surv. Tae: Kp 1873. p. 289,
The remains of this genus are constantly met with through-
out the Chalk of Kansas, but unfortunately in very fragment-
ary condition, consisting for the most part of the palatines and
mandibles. The skull, as described by A.S. Woodward," is tri-
angular in form, with the roof flattened and slightly depressed,
with the posterior lateral portions sculptured. The premaxilla
is thin and deep, while the maxilla is long and slender, with
small teeth along the lower border. The palatines, called pre-
maxillee by Cope, are dense masses of bone, with a single large
fang extending downward from the anterior extremity. This,
130. Proc. Geol. Assoc., vol. X (1888), p. 315.
374 University Geological Survey of Kansas.
as well as the other teeth, is firmly anchylosed to the bone.
The ectopterygoid, which joins the palatine posteriorly, is a
long, thin bar of bone, with a number of teeth on the lower
border of an irregular size. The mandible is moderately deep,
and bears two series of teeth. Those on the internal side are
very large and situated far apart, while those on the external
side form a minute fringe.
The succession of the teeth, as described by Professor Cope,™
is very peculiar: ‘‘The first teeth appear on the alveolar sur-
face at a considerable distance apart. The second teeth appear
immediately in front of these, and by their presence create the
irritation which results in the absorption of the root and shed-
ding of the crown of the first. The teeth of the third series —
appear in advance of the second, occupying the space between
them and the empty space previously occupied by No.1. These
may coexist for some time with teeth No. 2, as the specimen
indicates, but the result is as before—the shedding of the
adjacent older teeth. In the case of the anterior long tooth of
each side, the movement is reversed. Here the successional
tooth appears behind the position of the functional, which is
consequently shed, and in the old fish this tooth occupies a
position behind a concave symphyseal portion, which is con-
cave and edentulous, or only provided with the small teeth of
the marginal row.”’
The vertebre are deeply grooved laterally.
There are several bones of a single specimen which I refer to
this genus with doubt. These are figured on plate LXV, figs.
8,9, and 10. The largest of these bones, fig. 8, is no doubt an
opercular. It is very thin, excepting near the anterior-superior
portion, where the bone is much thickened and presents an an-
terior and internal articular portion, for the hyomandibular,
and also another along the lower border, where there is a long
and narrow facet, probably for attachment of one of the other
opercular bones. This bone is finely striated both externally
and internally.
The second of these bones, fig. 9, presents an articular facet
131. Cret. Vert. West, p. 301.
STEWART. | Cretaceous Fishes. 375
at each extremity, one of which is much longer than the other.
Along one of the borders there is a deep groove which is bounded
on each side by a thin lamina of bone. The third, fig. 10, is a
thickened mass of bone of an irregular shape, with a roughened
articular surface at one end, which extends beyond the remainder
of the bone in a spine-like projection. The fourth and last of
these bones, fig. 10, is very peculiar, in that it is almost iden-
tical with that described and figured by myself as an opercular
bone of Niphactinus. I would be inclined to think that I was
mistaken in referring this to \iphactinus were it not for the fact
that Professor Cope has figured this in his ‘‘ Cretaceous Verte-
brata’’’* as an ‘‘ uncertain bone’’ of this genus. This bone is
similar in structure, color, and in the matrix surrounding it, to
those described above; so there is little probability of it having
become mixed in collecting. ‘The known American species are:
Enchodus ferox Morton, Leidy, below Greensand No. 5, New Jersey.
Enchodus pressidens Cope, Greensand, New Jersey.
Enchodus petrosus Cope, Niobrara Cretaceous, western Kansas.
Enchodus dolichus Cope, Niobrara Cretaceous, western Kansas.
Enchodus calliodon Cope, Niobrara Cretaceous, western Kansas.
Enchodus oxytomus Cope, Cretaceous, New Jersey.
Enchodus tetracus Cope, Cretaceous, Delaware and New Jersey.
Enchodus shumardi Leidy, Cretaceous, Dakota.
Enchodus (Phasganodus) dirus Leidy, Cretaceous, Dakota.
Enchodus (Phasganodus) gladiolus Cope, Niob. Cret., western Kansas.
Enchodus (Phasganodus) anceps Cope, Niob. Cret., western Kansas.
Enchodus (Phasganodus) carinatus Cope, Niob. Cret., western Kansas.
Enchodus (Phasganodus) semistriatus Marsh, Cretaceous, New Jersey.
Enchodus amicrodus Stewart, Niobrara Cretaceous, western Kansas.
Enchodus minimus Stewart, Niobrara Cretaceous, western Kansas.
Enchodus shumardli.
Enchodus shumardi Leidy, Proc. Acad. Nat. Sci. Phil., 1856, p. 257;
’ Rep. U.S. Geol. Surv. Terr., 1, 1873, p. 289; Cret. Vert. West, p. 238.
Represented by the dentary bones of several individuals col-
lected by Professor Mudge from the Niobrara Cretaceous of
Kansas, the exact locality of which is not known.
The dentaries are small and slender and are about 35 mm. in
length. The internal alveolar border supports six large teeth,
132. Kans, Univ. Quart., vol, VIII A,
133. 1. c., pl. XL, figs. 8, 9.
376 University Geological Survey of Kansas.
all of which are firmly anchylosed to the bone. The largest of
these, situated just back of the symphysis, has a slightly ex-
panded base and is directed inward at the apex. It is strongly
striated and stands 8 mm. above the alveolar border. The
teeth following are smaller than this one and all are probably
of about the same size, except the last, which seems to be
smaller than the ones in front of it. They are minutely
striated under the microscope. The symphysis is incuryed,
and seems to have a strong articulation with its fellow on the
opposite side by means of the prominent bony tubercles so char-
acteristic of this portion of Hnchodus. The external alveolar
border bears a single row of minute teeth, the exact number of
which cannot be determined on account of the greater part of
them being broken away.
Enchodus petrosus. Plate LXX, fig. 11.
Enchodus petrosus Cope, Hayden’s Bull. U.S. Geol. Surv. Terr. No. 2,
p. 44; Cret. Vert. West, p. 238.
This species is represented by the palatines of numerous in-
dividuals and a small portion of one ectopterygoid.
The palatine is dense in texture and supports a very large
tooth at the anterior extremity, which is directed downward
and. slightly forward. The anterior portion of this tooth pre-
sents a sharp cutting edge, which extends from the apex to
near the base, along the sides of which the enamel is nearly
smooth, while posteriorly it is covered with conspicuous vertical
strie. Posterior to this tooth there are a number of curved
lines or scars formed in the replacement of this tooth, back of
which the border is slightly carinate. The upper and posterior
portions are very irregular.
Enchodus dirus. Plate LXX, fig. 14.
Phasganodus dirus Leidy, Proc. Acad. Nat. Sci. Phil., 1857, p. 167.
This species was first described by Doctor Leidy,™ from a
mutilated dentary bone from the Cretaceous deposits of Dakota,
as Phasganodus dirus, making the specimen the type of a hitherto
——=
134. 1. c., p. 167.
at gi See
STEWART. | Cretaceous Fishes. 377
unknown genus, which was later shown by Professor Cope to
be a synonym of Enchodus.*” The specimen here represented
consists of a fragmentary mandible, No. 322, and was obtained
by Mr. E. P. West from the Niobrara Cretaceous deposits of
western Kansas, the exact locality of which is not known.
Only the internal side of the dentary is exposed, which shows
four large teeth on the alveolar border, which are probably fol-
lowed by one or two more. The most anterior of these is set
upon an expanded base and has a sharp cutting edge antero-
externally, while postero-internally it is rounded and covered
with well-marked striz. The cross-section is thus seen to be
pyriform. At the base of the tooth, anteriorly, there is a slight
pit, in front of which there are two small teeth near the sym-
physis. The second of the large teeth is the smallest preserved.
It is non-striate and has anterior and posterior cutting edges
directed slightly outward and inward. The third and fourth
are slightly striated near the base and have the anterior cutting
edges directed outward more than in the second. The sym-
physeal portion is not well preserved, but the bone probably
becomes very shallow at this point. A well-marked fossa for
the articular extends forward to a point about half way between
the second and third large teeth.
Length of first large tooth (estimated)....................... 32 mm,
Jn ESCO EI gy prise yg oy) oh G22 ae Or 1 ich
IaREeE Noe ICOMIEN SCREAMING ys con 8 eco cis vee sia soso deuce Eoid¥,cc uv ot 14 ‘§
ENPRAIIS) CR GAIEN HW IELIDIANISES Ocoee cares 88 Sele eh woot bs Oe bee 4i°*s
EIS GEE ONE MELEE SD MAN CUO UE oe oy 0 ac. 2k Gms ok as aia alvie iv busine whys = Ae Se
REECE RETIN ERIE YONG soir ake see oa os nin ete 24s coe ie wh cf eae
Length of fourth large tooth (estimated)..................... LOS
MIRE EEC IVT VDI isis crate bs 5 of saci wate ow ae c's Bee > pean Ue
Enchodus dolichus. Plate LXX, fig. 12.
Enchodus dolichus Cope, Cret. Vert. West, pp. 239, 240.
This species is represented by portions of several individuals,
consisting of the palatines, ectopterygoids, and fragments of
other tooth-bearing elements.
The palatine is much smaller than in £. petrosus, and presents
the long, fang-like tooth at the anterior extremity, which is non-
135, Cret. Vert. West, p. 301.
25—vi
378 University Geological Survey of Kansas.
striate externally and has a cutting edge in front. Back of this
tooth the bone is more slender than in the species just mentioned,
and is very irregular above and behind where it joins the ecto-
pterygoid. The ectopterygoid (?) is along, slender bar of bone,
called maxilla by Professor Cope, and is covered below with teeth
well separated from each other. These teeth are large in front
but become smaller toward the posterior extremity. There is
one other tooth-bearing element that I have been unable to lo-
cate with any degree of certainty. It is expanded at one end
and contracted at the other, long, very thin, and covered with
small teeth on one of the edges. There are fragments of sey-
eral mandibles which may belong to this species, but as none
of them are associated with the palatines their identity as such
cannot be ascertained.
Palatine: Lengthiycc 4. ke ee ee hee ee 30 mm.
Greatestidepiht. se cea eee PO pee ete Ried. i esi 28 Simms
Isene th ohanterion toothy ys ee eee lite) OC
Enchodus parvus.
Enchodus parvus Stewart, Kans. Univ. Quart., vol. vir 4, p. 192.
This species is based on the right mandible of a single in-
dividual from Gove county, Kansas, the catalogue number of
which is 321. Itis of about the size of H. shumardz but differs
from it in several characters.
The dentary is very thin and presents nine or ten large teeth
on the border above. The first of these is the largest, but this
is smaller than in the species just mentioned. The second of
these is the smallest, while those following are of about the
same size.
Left dentary of Enchodus parvus, Right dentary of Enchodus amicrodus,
natural size. natural size.
©
~]I
©
STEWART. | Cretaceous Fishes.
Enchodus amicrodus.
Enchodus amicrodus Stewart, Kans. Univ. Quart., vol. vir a, p. 193.
This species is represented by the type specimen only, which
consists of the right dentary, the external side of which is ex-
posed. The catalogue number of the specimen is 324.
The principal character that distinguishes this species from
the others belonging to this genus is found in the absence of a
row of minute teeth on the external alveolar border. The
dentary is shallow and supports nine or ten large teeth. The
anterior of these teeth is slightly recurved at the apex, and the
base does not seem to be expanded as in some of the other
species. The external surface of this tooth seems to be non-
striated. The following teeth are mostly broken away but their
bases show them to have been irregular in size.
ea RAREREE VESTER EMONACTS Sc cedJarc AOS ACERS eek his kee Lee 63 mm.
Height of anterior tooth above border...:.. ........... etna ee 10; 4
Mr eemeY At aaIeT ON TAAEID VOI le eee wo ok ee es 6 eID so <a ow eee Bie
Pea SUSE ES. a RG i's kc et ona ae od ilalee kee ewe es Gia SS
Enchodus, sp. Plate LXX, fig. 13.
This species is represented by the remains of the mandibles
of several specimens, which may belong to L. dolichus. As
Professor Cope did not characterize this portion in a way to
render it recognizable, it will have to remain in doubt until the
specimen mentioned by Cope is more fully described.
The dentary is rather elongated, and thin inferiorly. The
alveolar border is slightly thickened, and bears teeth, of
which those on the external side are very small, while those
on the internal side are larger, less numerous. There are
nine or ten of these large teeth, the anterior of which is the
largest and slightly striated. The remaining eight or nine are
non-striated, and extend farther back than do those on the ex-
ternal row. The posterior extremity of the dentary is toothless
and directed slightly upward, forming a slight coronoid process.
The depth of the bone decreases rapidly toward the symphysis,
where there are numerous bony tubercles for attaching the two
jaws together. The external surface is smooth, with the excep-
380 University Geological Survey of Kansas.
tion of several conspicuous striew, which extend backward along
the middle of the side.
The articular fits into a deep emargination in the dentary,
and rises almost vertically from the cotylus. The bone is thin
throughout, and is covered with rows of bony tubercles. The
cotylus is very small, indicating a very weak articulation with
the quadrate.
Length ofdemtanycaeie ts. Son fk cas deem cle See eee ne 76.0 mm.
Depth ‘at conomoidne rete tic, to. ek cet Co oe ee AD) 9°
eng th of anticulaieaqreia cai LPG eee ee eee C020; es
eng throtitirstalarce soot, saci yes poe ein ere 16.5585
DERCETIDA.*
Hoplopleuride Pictet and Humbert.
LEPTECODON.
Leptecodon rectus. Plate LXXIII.
Leptecodon rectus Williston, Kans. Univ. Quart., vol. viira, p. 113.
Slender and elongate; head elongate, the jaws slender, the
anterior extremity in the specimen wanting. The hind end of
the mandibles is represented by an impression situated below
the posterior end of the skull. Teeth numerous, small, pointed,
slender. The orbit is situated posteriorly ; is of moderate size
and round. Scapular arch strong, the large opercular space in
front showing indications of the thin opercular bones. Verte-
bre about forty-five in number, elongate, more than three times
as long as deep, much constricted in the middle. Pectoral fins
small, composed of seven or eight rays. Ventral fin very small,
situated about the middle of the vertebral column; caudal fin
small, the rays feeble, evidently cartilaginous, the outline in-
dicated on the stone; the shape of the fin is regular apparently,
the angles produced. Other fins wanting or not preserved.
The side of the body, as preserved, shows three longitudinal
rows of large, firmly united scutes, apparently of the same
number in each series as the vertebre. The scutes are in the
* By S, W. Williston.
WILLISTON. | Cretaceous Fishes. 381
form of a double trapezium, with the V posterior, the middle
raised into a well-marked carina, which runs from the head to
the tail. Apparently there are five rows of these scutes on the
body. At the front the topmost row is near the middle line,
the lateral row has its lower edge over the line of the vertebra,
while the lowest row has the carina just below the pectoral fin.
The scutes have a finely roughened appearance, due to minute
rounded and shallow pits. There are no indications of small
scutes on the body intermediate between the larger ones.
MP MMUMME GI. AS TILONCLVOG. Cobia sj fei wc cle as 0k Side ee se 240 mm,
ReremEM CL GABTAPUID, cere uh, « woimy 8 ager a oon, whee 28 “1 ek a, See ie ak 250)“
See Oe VELL ONAN COMIC t Arce kins bo 2) Sein he ee es ee *
Greatest width, just back of pectoral fin..................5.5. ARE BG
Benen OF Gadus) tn, Upperlobe. ys. 2... os eee ce ee 22s
See mT RCC tON a AEE SD Bec, tire ed ete leer sg Uesacixveles Sees ge ede’ Wire
WE SS BRST Fg lis Ter RO a er 5 on
The specimen lies on the shell of a large Inoceramus, explain-
ing its excellent preservation. Close by are the remains, as
seen in the illustration, of several examples of a small fish of
unknown affinities, hitherto undescribed. The horizon is the
Niobrara Cretaceous of the Smoky Hill river. The specimen
was collected by Mr. H. T. Martin in 1895.
‘The family Hoplopleuride was established by Pictet for
fishes which were devoid of scales properly so-called, but which
are protected on the back and sides by rows of scutes. The
head is long and the jaws are provided with pointed teeth of
unequal size. The bones of the head are frequently sculptured
or granulose. The genera associated in this family by M. Pic-
tet are: Dercetis Agassiz, Saurorhamphus Heckel, Leptotrachelus
Von der Marck, Plinthophorus Guenther, Euryophilus Pictet, Pe-
largorhynchus Von der Marck. The fishes included in the genus
Dercetis were considered by Agassiz to resemble the sturgeons in
the arrangement of the dermal scutes, and were grouped among
the ganoids. Heckel held the same opinion with respect to the
position of Sauwrorhamphus, and Von der Marck also places the
genera Pelargorhynchus and Leptotrachelus amongst the ganoids,
but regards Ischyrocephalus as a teleostean. A careful review
of the whole of the genera, assisted by additional specimens o
382 University Geological Survey of Kansas.
Leptotrachelus | Dercetis| and Kuryophilus discovered in the Chalk
of Mount Lebanon, convinced M. Pictet that they formed a
group naturally associated, especially by the great analogy af-
forded by the peculiar arrangement of the series of scutes, and
that they formed a family of the teleosteans, to which he gave
the above name.’’ *”
Cope long ago described three species and two genera of this
group of fishes from Dakota, which seem to have been over-
looked by subsequent writers.” Concerning the relationships,
he says: ‘‘The relationship of the family of dercetiform fishes
has been discussed by various authors, especially by Pictet and
Von der Marck. The former regards them as teleosts; the
latter as ‘ganoids.’ As Ido not adopt the division signified by
the last name, I find Professor Pictet’s view nearer to the point.
The specimens indicate, further, that the Dercetidx belong to the
Actinoptert, and probably to the order Hemibranchii. The only
alternative is the order Isospondyli, and the characters which
separate the two are not clearly shown in the specimens. Dis-
tinct bones below the pectoral fins may be interclavicles, which
belong to the Hemibranchii.’’
The genus Trixnaspis, from the Niobrara of Dakota, there de-
scribed, has the dorsal and ventral scutes triradiate, the median
branch of the three directed anteriorly, together with numerous
band-like scutes. Ichthyotringa Cope, from the same locality, has
the body covered with small, round scales. The third species is
Leptotrachelus longipennis Cope, in which the dermal scutes con-
sist of median, dorsal and ventral rows of tripodal form.
From all these, as well as other forms, the present genus seems
distinct, though evidently nearest allied to Aspidopleurus Pictet
and Humbert, from the Lebanon Cretaceous.”
137. Davis, On the Fossil Fish of the Cretaceous Formations of Scandinavia, Trans. Ro; al
Dubl. Soc., IV, p. 428
138. Bull. U.S. Geol: Surv. Terr., IV, 67.
139. Pictet and Humbert, Nouv. rech.s. les. Poissons fossiles du Mont Liban, p. 109, pl. X,
TeNVET AS oe the Fossils Fishes of an Chalk of Mount Lebanon, Trans. Roy al alt ill, pl.
g.
STEWART. | Cretaceous Fishes. 383
MUGILIDA.
The family Jugilidx is represented in our collection by only
one genus, Syllamus, and it is very doubtful if this belongs
to it. In fact, it presents some characters which evidently do
not belong to this family, which are the presence of the lat-
eral line and more than four spines in the anterior dorsal fin.
There are also more than twenty-four vertebre, but it seems
likely that this could hardly be called a family character. On
the other hand, the absence of teeth seems to point to the Mu-
gilide, and for the present, at least, I think that it should be
left in this family until more complete specimens are found
which will determine its exact position.
SYLLAMUS. Plate LXXII, fig. 2.
Syliemus Cope, Rep. U. 8S. Geol. Surv. Terr., vol. 11, p. 180.
Cope has characterized this family as follows: ‘‘A short,
spinous dorsal fin. Ventral fins abdominal, posterior to the spi-
nous dorsal. Pectoral fins subinferior in position. Coracoid
bones forming a compressed, keeled body. Scales large, cycloid ;
laterai line present, extending along the middle of the sides.
Parietal bones less than epiotics, entirely separated by the
supraoccipital. [Frontal bones large, wide, their common suture
distinct. |
‘‘The opercular apparatus extends obliquely backward, while
the mandible is produced forward. Hence the inferior part of
the hyomandibular and the symplectic are directed obliquely
forward. The opercular bones are, and their inferior borders
reach the median line of the inferior side of the head.’’
The body is covered with moderately thick scales, which are
covered with fine concentric strie. The mandible is short, and
the dentary apparently toothless. In some respects our speci-
mens do not agree with Cope’s, for there are evidently two
dorsal fins in one of the specimens, which are separated from
each other by a short space. The anterior of these is well de-
140. Rep. U. S. Geol. Surv. West 100th Mer., vol. IV, 1877, p. 26.
384 University Geological Survey of Kansas.
veloped, but the posterior was probably composed of only a few
feeble spines, which were probably not preserved in the speci-
men described by Professor Cope.
Syllemus latifrons. Plate LX XII, fig. 2.
Syllemus latifrons Cope, Rep. U.S. Geol. Surv. Terr., vol. II, p. 181.
This species is represented by only two specimens in our col-
lection, from the Fort Benton Cretaceous, the exact locality of
which is not known.
The body is elongate and slightly larger than that of Leptich-
thys. The head is broad, flat, and, so far as can be determined,
the frontals form the greater part of the roof of the skull. They
are separated medially by a well-marked suture and possess no
markings. The orbits are large and the skull is rather acutely
pointed in front. The dentaries are short and seem to be with-
out teeth. The opercular bones are large and are directed well
downward. The pectoral fin is situated rather inferiorly, and
is made up of numerous small rays, fifteen to twenty in num-
ber, which are strongly bent at the proximal ends. The ante-
rior dorsal fin is short, and is made up of thirteen or more rays.
The posterior dorsal is small, and with but few rays, which
seem to be well separated from each other. The pelvic fin is in
too poor a state of preservation to determine its characters, and
the caudal fin is broken away in both of our specimens. Verte-
bree longer than deep, about thirty-six present.
Length of specimen to base of caudal fin (estimated)......... 345 mm.
Length of specimen to base of dorsal fin...................... 1B} 66
Length of specimen to base of pectoral fin.................... QOS
Depthiof body at anteriorsdorsaliin.)- eee ee eeaeee 69 *§
length of mandibular rami cw cen ee eee ee eee By) OF
Length of skull ie aos fits ie eRe ee An AOS
Width:of skullposteriorly .2.9-0.2.5). 0 ee eee ao °°
STEWART. | Cretaceous Fishes. 385
ON THE RANGE OF AMERICAN CRETACEOUS TELEOST FISHES
IN COMPARISON WITH THOSE OF SOME OTHER LOCALITIES.
After having treated the teleost fishes of the Kansas Creta-
ceous to as full an extent as our material will allow, I think
it will be well now to devote a little space to tables show-
ing their range in geological time, as well as to compare the
fauna with those of some of the otherZprincipal localities that
have yielded Cretaceous fishes. This work has already been
done with the fish faunas of Syria and Westphalia by Doctor von
der Marck, in his able paper entitled ‘‘ Uber die Verwandschaft
der syrischen Fischschichten mit denen der obern Kreide West-
falens,’’ but as yet I believe no work of this nature has been
done with the fish faunas of America and England. In the
present attempt, the selachians‘and ganoids will be omitted, as
they are beyond the scope of the foregoing paper. The species
of each genus are also omitted, as the other localities are so
widely separated from America as to have no great similarity
in this respect. The tables of Syrian, Westphalian and Eng-
lish genera have been copied from the works of Davis, von der
Marck, and A. S. Woodward. Thejlist of American genera has
been collected from the various government reports and scien-
tific journals in which they have been described, and is thought
to be fairly complete, although there may be a few genera which
have escaped my notice.
Notwithstanding the fact that very little has been done on
Cretaceous ichthyology in America during the past two de-
cades, twenty-six genera have already been described from its
deposits, and from the rich fish faunas of some of the localities
we may expect that in after-years the number will be mate-
rially increased, when more carefully and systematic collections
are made with this end in view.
Following is a table showing the distribution of the American
genera in time, the most of which have been found in Kansas:
386 University Geological Survey of Kansas.
Cretaceous No. 3 4 5
Benton, | Niobrara.| pion, | Hille, | Hecent
ANCISETOGODS 2)... fc Ae ein ee Bellow wore alee | aie aes ee]| Gok Ua eee toatl |
AMOR TNS 3 ai hs Ss een reer ee
A PSOPelisee2 5 Sees ae eee eee
1 S12) ih POR ee MA atest et gid.) Sear A pene at ra Nhe ko ce 4 ———s |
Cimohich tyes sere eee ees ee he eee a
Cladocycluswy. 2A eae OL Oe ee eens ee a
OR COLIS uss Fs eat We ae ee | A aeons a
TEIN O\s 15, he are cog er a ee ee oe BoC oe |
inChoGuse ae sorts oe Hee oe ween ——
Gailli@uisy Jar ences ee Bee eve ee ary oa a
fehthyodecteshen: ea. sherras el aes —s |
eh Ghiyotrin@ aye ck ok cuca. la Sena aes _——
dU) oly Zl a2) ae a ree Ra PnRrR Pt RRO Se A. aaa ear A IS. BN Gg —
Me PtLCCOdOMAAs Ee koe Laue 2. ae ale ee Seer —
Weptreh thy systeick os eee Aa ==
OPIiCARGIAUS ek Ue sas A ees ee ———
Pachyrhizodus iiss ek. eo eee a
Relecorapisie ic. ee ee ===
iBrotosphy tenance |e ee a ee
SAUL OG ONY. 42) oieh shir ce cei cos DereseAe Suet hee SE aay oo ob pela
Saurocephalhus J is. coer leer cee _———
SEPT OUT EIU SVs pint se iy QE a soa border egy tach ig a
Spaniodonite sac) Hie ee ee ae ———
SETA TCOCMUS Isls Al Weatte Seiclepa cas cee etd are ee ee cxcnmenaas
rehi7l Keys 2) 001) bs] ieee Nimes IR wee eure uate eet scl UR Geet 8 om | ED
PTissHas pis. cys AGh Woes ne gaia ee ae
Petheodus eye A ea ee ees a
MiphacsmMUs see ei ie ee ms me
Pachyrhizodus is probably represented from the Fort Pierre by
a fragment of a maxilla or dentary found by myself near Lisbon,
Kan., during the summer of 1898; the specimen is in a very
poor state of preservation, but bears a close resemblance to this
genus. Anogmius is represented from the Fort Benton by two
poorly preserved specimens, which were identified by Professor
Cope as Anogmius aratus.
Next is given a table of genera of teleost fishes from the Up-
per Cretaceous deposits of Hakel and Sahel-Alma, in Syria,
Baumberge and Sendenhorst, in Westphalia, England, and the
United States.
387
Cretaceous Fishes.
Srewakrt. |
‘puwisydsoqoig
*snjyeqdso0ineg
‘muOpoinesg
*SOTTTD
*saqyoopo fq 490T
‘snurjoeydiy
‘aopoosyderyT
*BSUlIgOAqAYOT
"SIdSeUuaety,
*SIqo010q
‘uopoturds
‘sfqQyorjdey
‘SUIUTPARg
's07B4g pozlug
*puawiAydsojorg
‘snyeydsooinesg
*‘uOpoInes
‘saqoepoAyyygoy
‘snurjoeydry
‘sIqo010q
*SOPlolouls”Eg
(4) snangeg
‘pussy
*SIQOD.10.
‘snqoudys0s1e [Og
‘SHTTOUITY
*snyeydso0a1A YOST
‘soprlo1oqydosstauy J,
‘snyoAqdozeulloed
‘sn, {puodsdyorrg
*sIgooudtoRy,
*BITMIOIOT
*SNWOSOPIIBYD
*snutosoydory
*SNIUIPIRG
*‘SOplOUIpPLeg
MOSOGOTAIOVCT
*sTde[Oo[O FT
‘sIToydoyay,
*(xosny) snronry
*SnOATOMR |
*SNLO9ST
‘qs1oyuepueg
puv os10equineg
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*SIZOOING
‘snqTouryy
‘snqquRsAingy
‘stpoydoyue dg
"BISIMOT
‘sTpoydo Lng
‘uopotukedg
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‘snaneldopidsy
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‘uoporurdg
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‘pad npIO.Ig WONG
‘vodul9
‘stpoqdouey
*SOPLOPMOIOR HT
‘PAP
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TALSOLTAL 1°pso
University Geological Survey of Kansas.
088
“SnpOADOpKIO
“snmp AS
*sIde1oo9[og
*xIjedosd vy
*‘snpooyyoy,
‘snpoyoug
"x ALOG
“SsnIwWSOUY
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*LOISVSOSO
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*10JSVSOIDV
*SIdo[O.108 JL
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*BUIOSOWGO
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ely 1edes
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‘Te18#H
389
Cretaceous Fishes.
*
_
a
a
=
e-
N
*sIZo0.10q
‘uopotueds
“SuIUIpsleg
‘mOpoi4slouy
*snpoAo0pe[D
‘snpoyoug
*xAlOg
‘snpoziqid4yoeg
‘sAQIYOITOUWT/)
‘supoyedsys
*puwidqdsoyolg
‘snyeydso0ineg
“uoOpoinesg
*soqyoopocyygoy
‘snurgoeydry
‘soyeyg poytag,
*BZIYAAYOST
‘NOpoOlyslouy
‘soqyeqg poy
‘sIgoo10q
‘uOpoi4stouy
*snjasoopR[D
*snpoyqouny
*xAIOG
‘snpoziyqrAyqoeg
‘sAqZYOTOUL
“4 Snpoqeiyg
‘euwisdysojoig
‘snyeqdsooineg
— mOpoineg
*saqyoopoAyyyoy
‘snhurgoeydiy
*puelsuy
‘STJOOLOG ‘SIZOO10q ‘sIgQ010q7
‘aopoturds ‘uopotuedg
‘SnIUIpseg *SRIUIPIRG
‘snpoyouq
*xAlOg
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pue os10quineg ‘TeAPH
‘sn pooyyse|d
*SILYOUAYAOTO, )
‘uopoiysiouy
*(aopo.olyy ) stydesdAfog
‘puelsagy
‘snanpoqyouny
‘snyeydoooupiyoy
‘(dOpoOuoT py ) StydesdAjog
*snhojsorpuoyg
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pus osiequantg "BUT Y-[eqes ‘oxy
GO}ISOd UIL}1OOUN JRYMOTIOS JO OAV BIOUOT SUIMOT[OJ OY ‘SuloSo10J oy sepisog
390 University Geological Survey of Kansas.
In the foregoing tables, we see that there have been twenty-
eight genera of teleost fishes found in the Upper Cretaceous
deposits of North America, twenty-nine from England, twenty-
eight from Westphalia, twenty-six from Sahel-Alma, and twen-
ty-two from Hakel. Of these, we find that England has thirteen
genera which are found in the American Cretaceous deposits,
Westphalia two, Sahel-Alma four, and Hakel three.
The genus Stratodus has been reported from the Chalk of Eng-
land by A. S. Woodward," who has described one species, S.
anglicus, from Sussex. From the figures given in plate I, 1 am
inclined to think that it is rather doubtful if this is Stratodus, as.
the dentary (fig. 4) is entirely different from that portion of S.
apicalis, described in the foregoing pages. There is also hardly
enough of the palatine (fig. 3a) to determine its exact relation-
ship.
141. Proc. Geol. Assoc., vol. X, pp. 314, 315.
———
i i i i i i a eee
APPENDIX.
IN justice to Mr. Stewart, it should be said that his absence
in the field during the printing of the foregoing paper has
made it impossible for him to see the proof. Its correction,
therefore, has devolved wholly upon myself, and I cannot hope
to have done as well as would have the author. While in the
printer’s hands an important paper on the Kansas Cretaceous
fishes was received from its author, Mr. F. B. Loomis,* too
late to be recognized in the text. A brief review of this article
is given below by Mr. Stewart, and the present writer has added
thereto a list of the species and genera treated by Mr. Loomis
in systematic sequence.—S. W. WILLISTON.
A recent article on the Kansas Cretaceous fishes, by Mr.
F. B. Loomis,* reaches me while engaged in field-work, where
I do not have access to the literature or the manuscript of
the foregoing article on the Cretaceous fishes of Kansas, and
while the work itself was rapidly going through the press. I
can, therefore, make only the briefest and most general com-
ments upon the paper. As a whole, the paper is to be com-
mended, though I am inclined to think that the author has
erred in some instances, which I may here point out briefly,
reserving a more extended commentary upon the paper for a
more propitious time.
On page 229 of the cited work, the author describes, under
the name of Thryptodus, a large part of the skull of a fish which
I am confident belongs to the genus Anqgmius, as herein de-
scribed. The Kansas Museum specimen shows the top of the
skull well preserved with the exception of the ethmoid and
* Die Anatomie und die Verwandschaft der Ganoid und Knochenfische aus der Kreide For-
mation von Kansas, Paleontugraphica, Bd. XCIV.
(391)
392 University Geological Survey of Kansas.
nasals, and all of which are so closely associated with the
mandibles, vertebree and other bones belonging unquestionably
to Anogmius that there can be doubt of their generic identity.
The bone figured on plate XXII (fig. 5), as a third cerato-
branchial of the right side of the new genus Pseudothryptodus,
appears to be the anterior internal portion of a maxilla of Anog-
mius. ‘The bones shown in figures 4,6 and 7 I have seen among
our specimens in a fragmentary condition.
On page 252 and plate X XV, the author figures and described
a portion of the skuil of Sawrocephalus broadheadi Steward (sic) *.
I have described but one species of Saurocephalus (S. dentatus) ,
which, on the preceding page, the author places as a synonym
of S. lanciformis Harlan. It seems evident that the author has
in some way got my two species, Saurocephalus dentatus and
Saurodon broadheadi, badly mixed. Later I will show conclu-
sively wherein S. dentatus differs from 8S. lanciformis. Under
the name Osmeroides the author describes material which I have
placed in the genus Anogmius. Whether or not the author is
right in doing so, | cannot say. I may be permitted to state,
however, that Dr. A. S. Woodward, after an examination of the
material in the University Museum described in the foregoing
pages, expressed his opinion that the genus to which it belonged
was different from anything that he knew. Possibly a more
thorough examination of better material might induce Doctor
Woodward to reverse this opinion. Nevertheless, the opinion
of so able an ichyologist is worthy of much consideration.
On page 265 is described ‘‘ Pachyrhizodus ferox Steward.’’ Evi-
dently intended for Pachyrhizodus velox Stewart.
The name Enchodide is inadmissible on etymological grounds
—it should be Enchodontide. A. STEWART.
Kimmswick, Mo., November 3, 1900.
* Every where throughout this paper, and in numerous places, Mr. Stewart’s name is wrongly
spelled.—s. W. W.
i i. ae
Cretaceous Fishes. 393
List of Kansas Fishes Described and Figured by Mr. Loomis, cf. c.
Protospondyli.
Protosphyrena penetrans Cope, p. 224.
obliquidens Loomis, p. 225.
tenuis Loomis, p. 225.
nitida Cope, p. 227.
Plethodidez.
Thryptodus Loomis, p. 229.
zitteli, p. 234.
rotundus Loomis, p. 235.
Pseudothryptodus Loomis, p. 235.
intermedius Loomis, p. 236.
Chirocentridez.
Ichthyodectes occidentalis Leidy, p. 242.
hamatus Cope, p. 243.
multidentatus Cope, p. 243.
ctenodon Cope, 244.
anaides Cope, p. 244.
Portheus Cope, p. 246.
Saurodon Hays, p. 247.
phlebotomus Cope, p. 248.
pugmeus Loomis, p. 248.
Saurocephalus Harlan, p. 249.
lanciformis Harlan, p. 251.
broadheadi Stewart, p. 252.
Albulidee.
Syntegmodus Loomis, p. 252.
altus Loomis, p. 253.
Elopidz.
Osmeroides Agassiz, p. 255.
polymicrodus Stewart, p. 256.
evolutus Cope, p. 257.
Salmonidze,
Pachyrhizodus Dixon, p. 258.
caninus Cope, p. 262.
latimentum Cope, p. 263.
sheari Cope, p. 264.
lepitopsis Cope, p. 264.
leptognathus Stewart, p. 264.
ferozx (sic) Stewart, p. 265.
curvatus Loomis, p. 265.
Enchodide.
Cimolichthys Leidy, p. 267.
nepceolica Cope, p. 271.
merilli Cope, p. 272.
semianceps Cope, p. 273.
contracta Cope, p. 273.
Enchodus Cope, p. 278.
petrosus Cope, p. 278.
dolichus Cope, p. 279.
shumardi Leidy, p. 280.
amicrodus Stewart, p. 280.
26—vi
EXPLANATION OF PLATES, PART II.
PLATE XXIII.
Kansas Cretaceous Sea. (Page 234.)
PLATE XXIV.
Figs. 1, la, 16.—Lamna sulcata Geinitz, after Leidy, natural size.
Figs. 2, 2a.—Scapanorhynchus rhaphiodon (texana) Agassiz, natural size,
after Leidy.
Fig. 3.—Leptostyrax bicuspidatus Willist., from the side, natural size.
Fig. 6.—Scylliorhinus (Lamna?) gracilis Willist., enlarged.
Figs. 7, 8.—Scylliorhinus planidens Willist., enlarged.
Fig. 5.—Scylliorhinus rugosus Willist., enlarged.
Figs. 9, 10.—Fragments of undetermined teeth from Kiowa shales.
Fig. 11.—Celodus stantoni Willist., imperfect right splenial dentition, natural
size.
Fig. 12.—Ca@lodus brownii Cope, imperfect left splenial dentition, natural size.
Figs. 13, 13a.—Amphibian atlas from Laramie Cretaceous.
Fig. 14.— Undetermined shark tooth (Coraz?) from Benton of Colorado, en-
larged.
Figs. 15, 15a.—Leptostyraz bicuspidatus Willist., natural size.
Middle figure of plate—Selachian verbebra (Coraz?) from Niobrara Cretaceous.
PLATE XXV.
Ptychodus mortoni Mantell, natural size.
PLATE XXVI.
Fig. 1.—Ptychodus mortoni Mantell, part of dentition of upper jaw, as pre-
served in the matrix, four-sevenths natural size.
Figs. 2, 4.—Scapanorhynchus rhaphiodon Agassiz, natural size.
Fig. 3.—Lamna appendiculata Agassiz, natural size.
Fig. 5.—Jsurus mantelli Agassiz, natural size.
Fig. 6.—C@lodus stantoni Willist., crowns of two teeth from internal row,
lower jaw, much enlarged.
Fig. 7.—Leptostyrax bicuspidatus Willist., enlarged.
(394 )
STEWART. | Cretaceous Fishes. 395,
PLATE XXVII.
Fig. 1.—Ptychodus mortoni, end of superior dentition, continuous with left
end of fig. 1, plate XX VI, but less reduced.
Fig. 2.— Ptychodus mortoni, a transverse series of teeth, arranged more loosely,
from near the right extremity of fig. 1, plate XX VI, about two-thirds natural
size; the upper series belong one at each end of the lower series.
PLATE XXVIII.
Ptychodus martini Willist., three-fourths natural size; the teeth were discoy-
ered disassociated.
PLATE XXIX.
Figs. 1, 2, 3.— Ptychodus, sp. indet.
Fig. 4.—Ptychodus occidentalis Leidy, natural size.
Figs. 5-8.—Ptychodus anonymus Willist., natural size.
Fig. 9.—Ptychodus polygyrus Buckl., a little enlarged. —
Figs. 10-15.—Ptychodus whipplei Marcou, nearly natural size.
Figs. 16-18, 20-22, 24.—Ptychodus anonymus Willist., nearly natural size.
Figs. 19, 23, 25, 26, 27.—Ptychodus, spp.
PLATE XXX.
Figs. 1-3.—Undetermined lamnids from Kiowa shales, Lower Cretaceous, en-
larged one-half.
Fig. 4.—? Mesodon abrasus Cragin, enlarged one-half.
Figs. 5, 6.—Lamna, sp., from Kiowa shales, enlarged one-half.
Figs. 7, 8.—Coraz curvatus Willist., from without and within, enlarged nearly
two diameters.
Figs. 9-11.—? Ptychodus janewayii Cope, enlarged.
Fig. 12.—Lamna, sp., enlarged nearly two diameters. Kiowa shales.
Fig. 14.—Ptychodus polygyrus, from the side, natural size (the same tooth
figured on plate XXIX, fig. 9).
Fig. 13.—Ptychodus occidentalis, enlarged nearly two diameters.
PLATE XXXI.
Figs. 1-40.—Coraz falcatus Agassiz, about nine-tenths natural size; isolated
teeth from many individuals.
Figs. 41-46, 50-52.—Jsurus mantelli Agassiz, nearly natural size.
Figs. 47-49.—Lamna appendiculata Roemer, nearly natural size.
Fig. 53.—Ptychodus, sp., enlarged one-fourth,
396 University Geological Survey of Kansas.
PLATE XXXII.
Figs. 1-1l.—Corax falcatus Agassiz, from a single individual.
Figs. 2-2/.—Tsurus mantelli, from a single individual.
Figs. 3-3c.—Lamna appendiculata.
Figs. 4, 5.—Scapanorhynchus rhaphiodon Agassiz.
Figs. 6, 7.—Isurus mantelli.
All six-sevenths natural size.
PLATE XXXIII.
Skull of Xiphactinus audax Leidy, one-third natural size: pmax, premaxilla;
mx, maxilla; den, dentary; dar, dermarticular; eth, ethmoid; /7, frontal;
pa, parietal; soc, supraoccipital; epot, epiotic; ptot, pterotic; pa, parietal ?;
prf, prefontal; pfr, postfrontal; pal, palatine; sor, suborbital; swor, supra-
orbital; sc, sclerotic ring; ps, parasphenoid; su, jugal?; hm, hyomandibu-
lar; pop, preopercular; op, opercular; sop, subopercular; ¢, quadrate; 7, 2,
vertebre; n,n, 7, neural spines.
PLATE XXXIV.
Left mandible of Xiphactinus audax Leidy, external view, one-half natural
size: den, dentary; dar, dermarticular.
PLATE XXXV.
Left mandible of Xiphactinus audaz Leidy, internal view, one-half natural size:
den, dentary; dar, dermarticular; auar,autarticular; sp,supposed splenial.
PLATE XXXVI.
Xiphactinus audax Leidy, one-half natural size.
Fig. 1.—Sclerotic ring.
Fig. 2.—Premazxilla.
Fig. 3.— Maxilla.
PLATES XXXVII, XXXVIII, XXXIX.
Maxille, premaxille and mandibles of Xiphactinus audax Leidy, one-fourth
natural size, showing the great individual variation in these parts.
PLATE XL.
No. 155. Maxilla, premaxilla and mandible of XYiphactinus brachygnathus
Stewart.
PLATE XULI.
Superior maxillary condyles of Xiphactinus audax Leidy, one-third natural
size, showing variation in the same.
STEWART. | Cretaceous Fishes. 397
PLATE XLII.
Figs. 1, 2, 3.—Vertebre of Viphactinus audax Leidy (after Hay), one-sixth
natural size; ana, basal piece of neural arch; na, neural arch; cen, centra.
Fig. 4.—Palato-quadrate arch of Niphactinus audax Leidy, one-third natural
size: pal, palatine; mpg, mesopterygoid; mtpg, metapterygoid; pt, ptery-
goid; g, quadrate; sym, symplectic.
Fig. 5.—Hyomandibular of Viphactinus audax Leidy, one-third natural size.
Fig. 6.—Pelvic actinosts of Niphactinus audawx Leidy, one-third natural size.
Fig. 7.—Ceratohyal of Viphactinus audax Leidy, one-third natural size.
PLATE XLIITI.
Opercular bones of Xiphactinus audax Leidy, one-third natural size.
Fig. 1.— Operculum.
Fig. 2.— Preopercular.
Fig. 3.—Supposed suboperculum,
PLATE XLIV.
Cranial bones of Xiphactinus audax Leidy, one-half natural size.
Figs. 1, 2, 3, 5.— Bones of uncertain position.
Fig. 4.— Probably a fragmentary coracoid.
PLATES XLV-A AND XLV-B.
Shoulder-girdle of Viphactinus audax Leidy, one-fourth natural size.
Fig. 1.— External view: c/e, cleithrum; sc, scapula.
Fig. 2.— Internal view of upper end of cleithrum and precoracoid, peor.
Figs. 3, 4.—Xiphactinus brachygnathus Stewart: pm, premaxilla; maz,
maxilla; den, dentary: dar, dermarticular—one-third natural size.
PLATE XLVI.
Bones of Xiphactinus audax Leidy.
Figs. 1, 2, 3, 4, 5.— Fin-spines and rays, one fourth natural size.
Figs. 6, 7.—Hypural bones, one-half natural size.
Figs. 8, 9, 10.— Basiosts from pectoral fin, one-half natural size.
Figs. 11, 12.— External and internal view of neural arch from near the skull,
one-half natural size.
598 University Geological Survey of Kansas.
PLATE XLVII-A.
Bones of Xiphactinus audax Leidy, one-half natural size.
Figs. la, b, ec.—Superior, anterior and posterior views of first anterior verte-
bra.
Fig. 2.—Side view of second anterior vertebra.
Figs. 3, 4, 5.— Third, fourth and sixth anterior vertebrz, seen from the side.
Figs. 6, 8.— Hemal arches and spines.
Figs. 7a, 6.— Posterior and side views of vertebrz from near the extremity of
caudal fin: cen, centra; hs, hemal arch and spine.
Fig. 9.—Superior view of maxilla and premaxilla.
PLATE XLVII-B.
Fig. 1.—Xiphactinus audax Leidy, portion of caudal fin, showing the upturned
condition of the vertebral centre in this region, about one-half natural size.
Fig. 2.—Top view of skull of Jchthyodectes hamatus Cope.
PLATE XLVIII.
Figs. la, 6, c.—External and internal views of maxilla and side view of pre-
dentary of Saurodon broadheadi Stewart, two-thirds natural size.
Fig. 2.—Dentary of Xiphactinus lowii Stewart, two-thirds natural size.
PLATE XLIX.
Ichthyodectes anaides Cope, one-half natural size.
Fig. 1.—Skull: eth, ethmoid; jr, frontal; so, supraoccipital; epo, epiotic;
ptot, pterotic; pos, postfrontal; pfr, prefrontal; opt, opisthetic; pro,
prootic; bo, basioccipital; v, vertebra; bs, basisphenoid; par, parasphe-
noid.
Figs. 2a, 6b.—_Maxilla and dentary.
Fig. 3.—Hyomandibular, probably belonging to this species.
Fig. 4.—Skull of Jchthyodectes or Gillicus, one-half natural size: pa,
parietal; other lettering as in fig. 1.
Figs. 5, 6, 7.—Mandible, posterior end of maxilla and quadrate of Ichthyo-
dectes ctenodon ? Cope, one-half natural size.
PLATE L.
-Ichthyodectes hamatus Cope, one-half natural size.
Figs. la, b.—Side and top views of maxilla.
Figs. 2, 3.—Top and side views of palatine.
Fig. 4.—Preoperculum.
Fig. 5.— Quadrate.
StTEwaktT. | Cretaceous Fishes. 399
PLATE L—continued.
Figs. 6a, 6.—Posterior and top views of first anterior vertebra.
Figs. 7a, b.—Side and top views of maxilla and premaxilla /chthyodectes
eruentis Hay.
Figs. 8a, b.—External and internal views of premaxilla.
Fig. 9.—Front view of dentary bones.
Figs. 10a, b.—Top and side views of maxilla of type specimen from Professor
Hay’s collection.
| PLATE LI.
Ichthyodectes acanthicus (?) Cope, one-half natural size.
Figs. 1, 2.— Superior and side views of maxilla.
Figs. 3, 4.— External and internal views of mandible: den, dentary: aut,
autarticular; der, dermarticular.
Figs. 5a, 6.— External and internal views of premaxilla.
Fig. €.— Portion of scapula. _
Fig. 7.— Palatine seen from side.
Fig.'8.—Side view of vertebre.
Figs. 9a, b, ec, d.— Pectoral fin-spines and rays.
Fig. 10.— Pelvic actinosts.
Fig. 11.— Portion of caudal fin, probably of this species.
Fig. 12.—Jchthyodectes ctenodon Cope, ceratohyal.
Fig. 13.— Bone of uncertain position.
PLATE LII.
Gillicus arcuatus Cope, one-half natural size.
Fig. 1.—Skull: eth, ethmoid; fr, frontal; pa, parietal; so, supraoccipital;
epo, epiotic; ptot, pterotic; pofr, postfrontal; Amnd, hyomandibular;
pfr, prefrontal; sc, sclerotic ring.
Fig. 2.— Maxilla, external view.
Figs. 3, 4, 5.— External and internal views of mandibles.
Fig. 6.— Ceratohyal.
Fig. 7.— Quadrate.
Fig. 8.— Palatine and mesopterygoid.
Fig. 9.— Preoperculum.
Fig. 10.— Fragment of sclerotic ring.
PLATE LIII.
Palato-quadrate arch and opercular bones of Gillicus arcuatus Cope, about
seven-tenths natural size: g, quadrate; mfpg, metapterygoid; pg, ptery-
goid; pop, preoperculum; op, operculum.
400 University Geological Survey of Kansas.
PLATE LIV.
Gillicus arcuatus (?) Cope. Caudal fin, slightly under one-half natural size.
PLATE LV.
Saurodon xiphirostris Stewart, about three-fifths natural size: pden, pre-
dentary; den, dentary; dar, dermarticular; maz, maxilla; pmax, pre-
maxilla; eth, ethmoid; /7, frontal; pfr, prefrontal; pa and ptot, pterotic;
spot, sphenotic; hm, hyomandibular; qu, quadrate; na, nasal; prop, pre-
operculum; op, operculum; pal, palatine; sc, sclerotic ring; 1, 2, 3, ver-
tebree.
PLATE LVI.
Saurodon ferox Stewart.
Fig. 1.— Upper and lower jaws, one-half natural size: max, maxilla; pmaz,
premaxilla; den, dentary; pden, predentary; dar, dermarticular.
Fig. 2.—A small toothed element, the exact location of which is not known,
natural size.
Fig. 3.— Right quadrate, one-half natural size.
Fig. 4.— Ceratohyal, one-half natural size.
Fig. 5.— Centra of first anterior vertebra, natural size.
Fig. 6.—Glenoid portion of scapula, natural size.
Figs, 7, 8.— Portions of pectoral fin-spines.
PLATE LVII.
Saurodon ferox Stewart.
Fig. 1.— One-half natural size: max, maxilla; pma, premaxilla; den, den-
tary; pden, predentary; dar, dermarticular; auar, autarticular.
Figs. 2, 3.— Internal view of right quadrate and hyomandibular.
Figs. 4, 5.—Saurodon phlebotomus Cope, external view of right maxilla
and mandible.
PLATE LVIITI.
Figs. la, 6.— Premaxille of Protosphyrena occidentalis Stewart, natural size.
Figs. 2a, b.— Outline and external markings of rostrum of Protosphyrena ben-
toniana Stewart; the first natural size, the second one-fourth natural size.
Figs. 3a, 6.—Maxilla, premaxilla and dentary of Saurocephalus dentatus
Stewart, one-half natural size.
Figs. 4a, b.— Internal view of maxilla, premaxilla and dentary of same.
STEWART. | Cretaceous Fishes. 401
PLATE LIX.
Empo nepcolica Cope, one-third natural size.
Fig. 1.—Top view of skull: eth, ethmoid; jr, frontal; po, postorbital; so,
supraoccipital; eo, exoccipital.
Fig. la.— Pectoral fin-rays.
Fig. 2.-—Inferior view of skull.
Fig. 3a.— Palatine.
Fig. 3).— Mandible: den, dentary; art, articular.
Fig. 4.— External view of dentary.
Fig. 5.— Ethmoid seen from below.
Figs. 6a, b, e.—Hyoid bones.
Fig. 7.— Quadrate,
Fig. 8.—Preoperculum.
Fig. 9.—Operculum.
PLATE, LX.
Stratodus apicalis Cope, slightly over one-half natural size: sk, top of skull;
pmx, den, dentary; pal, palatine.
PLATE LXI.
Stratodus apicalis Cope.
Fig. 1.—Top view of skull, one-half natural size: jr, frontal; pa, parietal;
pt, pterotic; eo, exoccipital.
Empo nepcolica Cope, one-half natural size.
Fig. 2.— Vertebrz seen from the side.
Fig. 3.—Ethmoid seen from below.
Fig. 4.— Hyomandibular.
Fig. iss Premaxilla(?).
Empo semianceps Cope, one-half natural size.
Figs. 6a, b.— Palatine, inferior and side views.
Fig. 7.— Mandible.
Fig. 8.—Ceratohyal.
Fig. 9.— Posterior view of cotylus of articular.
Empo lisbonensis Stewart.
Figs. 10a, b.—Inferior and side views of palatine.
PLATE LXII.
Protosphyrena gigas Stewart, slightly under one-third natural size.
402 University Geological Survey of Kansas.
PLATE LXIII.
Fin-spines of Protosphyrena penetrans (?), slightly under one-half natural
size.
PLATE LXIV.
Anogmius polymicrodus Stewart, one-half natural size: den, dentary; art,
articular; mz, maxilla; pm2, premaxilla; gq, quadrate; ch, ceratohyal; sce,
sclerotic ring; a,a,a,qa, dermal plates; op, operculum; pop, preoperculum ;
iop, interoperculum; sop, suboperculum; o7, supposed orbital bones.
PLATE LXV.
Anogmius polymicrodus Stewart.
Fig. 1.— Top view of skull, one-half natural size: jr, frontal; »o, postorbital ;
pa, parietal; pt, pterotic: hmnd, hyomandibular.
Fig. 2a, 6.—First anterior vertebra, posterior and superior views.
Fig. 3a, 6.—Abdominal vertebrez, top and side views.
Fig. 4.—Premaxilla.
Fig. 5.—Condyle of quadrate.
Fig. 6.—Superior view of cotylus.
Anogmius evolutus Cope.—Superior view of cotylus.
Figs. 8, 9, 10.—Opercle and other bones, probably of Hnchcdus.
PLATE LXVI.
Anogmius polymicrodus Stewart.— Caudal fin, about one-half natural size.
PLATE LXVII.
Anogmius evolutus Cope, slightly under natural size: den, dentary; pal,
supposed palatine; ma, fragmentary maxilla; phar, supposed pharyngeal
bone of A. polymicrodus Stewart; a, bone of uncertain position.
PLATE LXVIII.
Pachyrhizodus latimentum? Cope, one-half natural size.
Fig. 1.—Right mandible: den, dentary; art, articular.
Fig. 2.—Premazxilla.
Fig. 3.—Mazxilla.
Fig. 4.—Quaadrate.
Fig. 5.—Condyle of quadrate, seen from below.
Figs. 6a, b, c.—Bones of uncertain position.
Fig. 7.—Vertebre.
Fig. 8.—Glenoid portion of scapula.
=—_—.
STEWART. | Cretaceous Fishes. 403
POAT EH) Taek:
Fig. 1.—Pachyrhizodus leptognathus Stewart, natural size: den, dentary;
art, articular; g, quadrate; pop, preoperculum; op, operculum?; a, bone
of uncertain position.
Fig. 2.—Pachyrhizodus velox Stewart, one-half natural size; mx, maxilla;
den, dentary; arf, articular; Ay, bone of uncertain position, probably a
ceratohyal, natural size.
PLATE LXX.
Fig. 1.—Pachyrhizodus leptopsis Cope.—Internal view of dentary, one-half
natural size.
Fig. 2.—Superior view of dentary of Pachyrhizodus caninus Cope.
Fig. 3.— Premaxilla.
Fig. 4.— Bone of uncertain position.
Figs. 5, 6.—Superior and posterior views of vertebra.
Figs. 7, 8.— Posterior and side views of basioccipital of Pachyrhizodus lati-
mentum ? Cope.
Figs. 9, 10.—Superior and posterior vertebrz of same.
Fig. 11.— Palatine of Enchodus petrosus Cope.
Fig. 12.—Palatine of Enchodus dolichus Cope.
Fig. 13.—Dentary of Enchodus, sp.
Fig. 14.— Internal view of dentary of Enchodus dirus Leidy.
PLATE LXXI.
Shoulder-girdle and pectoral fin of Pachyrhizodus, about one-half natural size.
PLATE LXXII.
Fig. 1.—Leptichthys agilis Stewart, about one-half natural size.
Fig. 2.—Sylemus latifrons Cope, slightly under one-half natural size.
PLATE LXXITI.
Leptecodon rectus Williston, about three-fifths natural size.
‘University Geological Survey of Kansas. VoLumME VI. PLATE XXIV.
LAMNA, 1. SCAPANORHYNCHUS, 2. LEPTOSTYRAX, 3, 15.
SCYLLIORHINUS, 5-8. CaELODUS, 11, 12.
University Geological Survey of Kansas, VOLUME VI. PLATE XXV.
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PTYCHODUS, 1. LAMNA, 3. CCELODUS, 6.
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University Geological Survey of Kansas. VoLuME VI, PLATH XXX.
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University Geological Survey of Kansas. VoutuME VI. PLATE XXXI.
CORAX, 1-40. ISURUS, 41-46, 50, 52. PTYCHODUS, 53.
University Geological Survey of Kansas. VOLUME VI. PLATE XXXII.
CORAX, 1. ISURUS, 2, 6, 7. LAMNA, 3. SCAPANORHYNCHUS, 4, 5.
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GENERAL
INDEX.
A: VPM POCIGAMGI 5 cc ose ce ee ee ws 6
PERE MUPINREIOT G2. 5. <i ca elsie ac eos ews He PU COTITOS, 5 sons wha ern w Soh w e's rae ee oes
/Msiocrinus magnificus ........... “8, GOS 02) | a a 8, 163
PUR MOCEUNIO « <6. .'65 Sa wg Mess ee 6 Ree tne hue) Senn MONE Ue E Nest cise 163
PePUMCEIEVA oo is iow Y de ss a Se | TEN ae (6 Sea 8
OTe | a ee ae SAME CULT VETOES ic ts chee of ary ales 6% 0's toe 7
RIMMING 2 ets Ee aus co Se eee GEV TIS BT EM bla sc... oak vate oso 105
(SEES EA GSS GO ae te 8, 167 MUBRenwIAr aE. ect Meee ss 105
RI oo etc oo ee Sos a 170 [sGbSS) OI. Tapes Ok capita Ene eecaee Same 104
SUSE SSNS ee ea eee eth ie 167 DALVILOSRICT. Cee oie aS 104
LE a are eres 167 PlABOSUI HbA ce Sos ess Se ae 104
TSU ee 168 Sino MTGIOSH lec ov sic Wien a es 104
leavenworthensis............. 172 SI) OLLULT Of: Gage areal ree ee Rema a 105
PELL LSE ES ere a ee hehe SL ee rr 6
BUNCHMOALUM oes. tees 169) Aucella hausmani.....:....05. 00. ey
SRI MNIEMEST feo 3c se kG et | SeearacOrnymenhug. .. 0c ce.... 0.5. 6,. 89
a ee ee 14 HAN MUGEG UALS. Se ko wo «cd ake Pee 89
Amblysiphonella .............. Bi phe AITO OE ope ass os) <arard'sc ok ne Dy iy OE
SESS Sr Ae ec eee ae 14 | PURE ars ei aat aie a?) Uo sce Sate 23
PURINA OEE Mig chs loo naa tee i, 10) PER Meter ci (i onhy Ave vieere use 23
CE Ne ee ne eee DE, CATA CHOICES Cn a ccc ove corel ande siscre 5)
PIDMEGIVER Ass oo czeee ye ss p's USO a Peat ANE, LOTS Pooch cvs anus Wesese wis wx ase 125
ORME NAMES ES AS ut eh EER SG s.s 18 PUNMSOVOTIOIS «he. de vas de ewe Lo
Ses i ee ee a ee 18 FOCHUIALCTATIA‘L. 6. heise env 125
SIDE? AGES itn oon oh a's oe 340, 391 COALS f(a Neg ats oe rae Se 126
Site UY Tt: ae OAs, a rs eae PR ACAMN CSET ates tt Bs. 5, c aheeas nee 7
MOU IICTOUNAN ES 0.5 Sn hain ws 7s Cae | MAVICMIOPEGLETS .. <i. ives toda Size 7, 114
Th ba ed ee a 7 carboniferus. ..... Boat eee rem AF
Anomites punctatus.............. 7 GRE RLERBGSI Sz oy a us cl ayy me ds 120
semireticulatus, ..............:. 78 coxanus..... Pee inom tye cis Lot
Ls a eed dD, 16 PAACIGUSAGUS So vn ja eesti wad se 119
PUI MOOR IMAM gre St new zs. scan die K Bias 98 jo) = 10.000) 0 EE 123
2 ES eee eee 391 SERIE a tengeloo L Wen OU os 1271,, 123
Lif SLE | ae SS Se eer eanies 9 (| MIGERINCAGIS co bk veka sry LLG
Arcidse ...... ae Te 8 SER CEAN MUM Cale oi ain sR ie ou 096 x. ovo: 5 118
Archzocidaris................. 6, 47 QGGIIGIDBIIS 2.) aka salen oss 114
i) ae nS 48 PIOVIGCHGEHIGs. sie iene ees 119
Cot od 49 POCtUALOTATIUS, 6605 6.6 cieraa ns 115
Ci SS ae 47 SIRS MEAIRRO Shes 8s v9. & thelie Sal's 122
510
Aviculopinniaye us oe salto
AMELICANAN. «| Hac). + soa 1438
DINMOLENISISN sync c cs oe eee 143
ACK hylan, ad ee 5, 20
PUGIS? ole vistas Lice Renee 20
By;
Beede, J. W., article by........... 3
Beryxmultidentatus:..cs9e eee 347
polymuicroduste..+..--e nee 342
Bourne, We One vee. oe ee ee 258
Brachtopodaw: io. ee Oy U5 ll
C:
Camerophoria globulina.......... 93
Camipoplay/lumape eee eee Sel
HEX UOS Mer aa ae 19
TOE G UTTAR eee ele es ee 19
vermiculare nds ©. tan rene 19
Candide: aetna Bae ate see DE 8
Cardiomorpiaraese es eee ee 8, 165
missouriensis...... She) IED
Cerlocninus seme te ee ae Grae
CRAIGS hence eel ae OS
LAYVEULONSISKe say | sense ees a2
hemisphericus.... 3H) Bid Bis
MISSOULIONSISH! 4.50) eee 30
MOOV MMOUNENGESS 5540 Goes onoe cans 33
@heenOmyaye sete sh eee ee oy
leavenworthensis............. 172
@hestets nace ry eee gen Dy LLG 2D
milleporaceus’..2... 7...) . 25
Cheete tid aa! ee oe tea epee ce sane 5)
Chonetesn ease ee eee GS, (zl
eranulifene! see ee 68, 69; 72
TEUMMMUGZAIA MS, 5 eed es een ee 68
Slaber waa eee aa wee 68
De VAS: Cera eee eu sed, Sakae 68
mesolobUismer aa seere: By sign Wal
millepunctatusraa see ne 89
IMUCTON AUS er we ater ee ee 69
Srraditate 2 he ee rca earn ere 69
VELNeSUlllan US a eee wee Gis,
Cimoliechthiyse.i. ae eee 393 |
SCMMANCE PS knee ae ee ees 338
SUlcatUsas avon eee eee 332
Cladochonvsi see. ee
Denne titi. toe nee 24
University of Kansas Geological Survey.
Classification of fossils ........... 9)
Cleidophorus occidentalis......... 136
Cleiothyris. i. 3.0 oo eee 7, 104
FOISSWAleegs ee Peres sel Ai ci, 3, 104
sublamellosay 2-52 tee 104
Clu peidee) cine Be 371
Cnidarias:5s35 eee eee 5
Contents: (500. be eee eee vi
Coelentératians 2:24.32 eee 9)
Coelodus*brownil <2) ee ee 254
Stantoni,...uoR yu oe, ee 255
Concentrics 2. ee 67
Conocardium: 3. pee eee 8, 164
Parrishy. 14's) ne eee 164
Corals) oo ose Ss Se eee 16
Corax. i), Bae eee 202
CUrVatuS. ot ee 253
faleatuse i. ah. os ee eee 252
Corynella: ce. ee eee 13
@raniia, Ue aes ee S, Br
carbonariaa) cs... eee oT
modestac: 05... oT
Cramindas (si Wis See ee eee 6
| Cretaceous fishes, described by
families. .2548 Vga nent eee 259
| Cretaceous; Upper eee. eee $
Crinoidea: 2222 os eee 6, 26
Crinoids sis aves 28 sk eee 26
Cyathaxonia-i 45.05 eee gS lis)
distortac. ) 6 a eee 18
Cyathophyilide. =. 8 an. eee )
Cyathophy lit 74. eee 19
Cypricardia wheeleri............. 155
Cy pricarcdiniay 240-6) cee eee 8, 164
Carbonariau. 0... eee 164
D.
Daptinusbroadheadit een ee 313
phlebotomus: eee 312
Delocrinus hemisphericus........ 35
TMISSOWUMIC NSIS eeu ea) ae neee 35
Delthytis) 24-208 ease ae eee 67
ID evel oy id: peeies Scie sls AON Meee hac 6, 57
PHAGE O01 ONG UR ren cea eL TB Mn RaD 59, 61
Dene tt MeN GAS ee ee 59
broadheadiys9 22. ol ase 59
CPASSD e200 octet ae ee 62
eymbulayiics ase ae scene 60
~ General Index
—Vol. VI. bit
PRABRGEMVCTIDUS Oo cscs oe cs 8 6, 40
EP TESS Ie CES RE em aria 32
WRMESIOCE Seiten isis ck de WU CaS 40
Explanation of plates........ 175, 394
iy
| Families of Cretaceous fishes..... 259
PYAGOL UNOS). SO, ae Gar vc a Sees i)
PPRURTERADANGE 0 fb h ga tare thn wea? ass KL 74, 98
PPR EAESUL ERs CMe es iotape ORGS 2 oa inl TR G
PRUE SPUR IOLE wis nh Wye Soh wer ees Gre a We 14
BREE Canee LL RENEL OTE «5 cote aly 8 soa » wo aA by «9
PVURULIDS nyu ds ies Cece oa ay 10s ad
AALS al oh ge ae Rea eae 10
GOURIRIIE TE EAL, MEL IN Eh vs ss Wik raga 10
rig ney | VR eh ts as ae 10
OLS UUUS EE Oe rod te ae 10
SEGRE Remy nee hee ye es 9, 10
VEIMPECUBRA Pte e ok. d kw wees 10
LERETSTR CIN) 159 (2 2) a os a 5
G.
REE SCELUIOL AINE fies ue csace paresis ae 252
Garvilba donee 25.0 evo Ons 125
JCI ofS Oe ee ee gets eR Ra 304
BERLE DUIS wii ickan. «a4 oer eee UO
GR a) eo Ae nS Se ae OL 98
longispina....... see 127
SVEULEIES (22S aera | ae ee na 126
ETO OS TA ee jens Ho wee Phe Rue 67
El.
ASRS ee alas os Reis win WR 371
1 LUN A oy aR eee ae 258
| Hemipronites crassus............ 62
eh eh 2 | fi br RON ae aes ea 62
HRSAOIISIN: (ches pene ce chee aS 62
CELT NATTA UG) CORO) arn SE ae 62
RMMNARICUI EY a fig 52.0 95,» nssidiers x ¥en. 29
PCARRD, oo Seco wails poe dies teleis, ® 8
[Evereile} is hale (i? i a 380
TEP Ne ee ee 74
Ler HE a Ne ite ee gs Oe ty. 105
rey C000) oy eS Saar de a 103
EXVORCIONOCKIBUG: . 35.05 xfea 262s 6, 42
MERINGUE: iin = 5 is, u ave ally 62s 42
SURSITMA DG) siete d dispx a's. e din «ue 43
Pouca Wal rr, «0
Derbya:
TE ee oe 2 63
OMAN RR ye scuyas s sb 5 eo
PURIENONNE Sl ed oo. 6 uc nc OS 389
Dercetis (Leptotrachelus) longi-
SMI N Te ihre iss s > ols eS 382
OEE ee re 7, 95
PIMC eg 8's cs. phen MARE 95
Museine meckjana.... . 26.5 «seen 55
ESS) nn eo 5d
SS ree ee 5D
E.
ou a, eee 2st G
RE IMIOLTNIA GA. 6 «5. anc. a.w aloo e isle Ses 6
ERENCE eS ce cesses av-bio wnee 6, 46
(OES SS gel ee PPE 5 6
0 SS Se eee 8, 166 |
aspenwallensis............... 166 |
OOS Soe) ee a 166
RS ee OK css fa ce Maca 393
EE eee ee Stn ao ee 8 330
RM MRCRDE ee 2 heats sie wr ales 339
ROMAIN os 8 osc. Wa asi 8 337
DEE TLS a ee eet Boy
Teo) on ee ee 338
Se NESRMINE RSS STS Reh cw 6 tela iaentin 332
PUMICE ooo ccs. w AAS ti 392, 393
POS ys) oh ne 373
SCREEN, ects. os eS Bio. aon
SENEBIOC TOMALES wid. tick sabe Ase «G2 379
EI wae tc aoa 376
CORIO oy raison ieee an Kisii ori
RE VU eine ss enews, ohihls 378
LEU | See a Bc ee 376
RRUATRMNIREE 20 eg ee x os Bere ss 375
SU nnn a 6,- 94
SASHA STRONRES Se yo. ci dicl oes ied. <aicots 91
UE Ee i ores iyo
Eph OU, 1) ain sna Tee ee LS
SERS IETIORLONG:. 5.5 <ie's.«s «-0.8eleualine 3
PUP V PME ROTOSS «5 oid vieis wv een es 361
Cu NS a er 362
[Mahi 6, (31, 38
IMOBMIOMTACHIUG:......s5 00 Jet 37
BNE Peas Woe chaise x mate 38-40
Eumetria punctulifera............ 103
Eumicrotus hawni............... 132
512
I,
Tehthyedectidea 22 -.0 Ae eee 262
Ichthyodectess=. +. 55 ..40r WIN, ash
ACAMtMIGUST Geiss oF . 301 |
anatdes inka vss... owen 296 |
ALCUATUSE coco 6 occ oe ee 306
CRUGDGUSS Ao Wer. 2c 0289 ae es 300
clenodonites a is eee 303
hamatus ts. S96 Jo Ae eee 298
Polymicroduss:..42- > se eee 306
fehthyotrimnean. sa. ae 382
Intesripalliata se eee 8 |
Irremulares. e6 bik 0 Sees 74
Isogramma millepunctatum ...... 89
TSUruUs ancocshos sok. : Ths eee 246
miambellly ac. ccs ets ee 246
Ke
TROY ays eos See Seer ee ent 4
10 |
Toe VOS:.ce 6 Sele se te Gece cee 67
Toadme lost: 05 gk er een. haar 98
Tamimidiaa acs ee eh te ce oe artes 246
Lamna appendiculata............ 247
IMNACLOMMIZA ee wee ONG
MUSSEL ee ee eee eas 248 |
quinguelateraliseea nase 250 |
raphiodoness tae Gee 251 |
Stilcaibasc. oe ie Sen ar eee 248
POX AMG te ea © ee eer tae 251
eda bellisttiatay: . reese ere 148
KNMOXEMSISE: 35 1520 a ees 154
DOlibay 2s tae eer nomen e foe 154
subsentulay ais nee eee ee 152
Bepttomdie sienna ee eae 67
Bepidosterdes ss. cee eee 956 |
epidotusime: o 2254.5 h2 eee eee 256
Leptecodon ea. Sa AA eee 380
FOCtUS Bnet ee eee 380
Ibe ptichthiyoyy, 5) te eee ee 312
ONS na ia tee ae Mey ace ee 372
Leptodomus granosus............ 168
POPeKAeNSis oiuhie vee eae lial
eptostyrax ss. 25.600 se eee 253
bicuspidatus cee ean, 953 |
eéptotrachelus: 6:2 thoes ee 382
Letter of transmittal. Vesa. Vv
Dim tele. os a hazy Gee eee 7
University Geological Survey of Kansas.
54
Damas Jo Fe Sei es ee ty 12,
retifera-s. 6.6 422s. ee 112
Limopteriay2: 23 ol a eee Tel
alata ss dc at ssndaee eee 130
Pibbosa: caver ye eee 129, 130
longispinara-2. 240) ee eee 127
Manan. 0. cece eee 128
subalata 0 04 oe, eee 131
ineatitaic <a Se eee 74
sing Wlaw .c.8 ee ee 6, 54
murphianay.. 07.245 ee 53
my tiloidés!3.. >.) 3 eee 54
umMbonatanc.c cee ae see eee
Ling ulidea. Ack eo eee 6, 54
ophop ky, limare eee nile 1s)
profundum 207) =e eee 7)
DEOMeT UN ae 17
WSU sr cjisia newline asa eee 18
Loomis, F’. B., paper, by....... 391, 39a
iow; Mii Ans ee eee 293
M.
| Macrodon 44.500 ee 8, 146
carbonarius®.. 5.75 422 -7ee eee 147
obsoletus 3. 30... eee 147
Say amoOnensisi i. eee 146
StrIAEUS y/o. one 146
TEMWIStLIA GUS ee eee 147
Magellania flavescens............. 53
Margimifera...5250.. -). cos eee 83
Martinia planoconvexa........... 101
MeClung ©. Bvt oe eee 2)
MeDowell. Ee Me. eee 258
Meckellaaiewsts. 3 eee 6; 65
striatocostata)..\5 nas - ose 65
Melonitidee: 2.3 a eee 6
Members of the Survey........... iv
Miesodonvabrasus).- sa sense eee 256
| Mesolobi.s:...5s- eee aes ee 74
| MEichiliniae 2. eee eases 552i
SUCHE 2). eee eee oe 21
Miliolites secalicus............ LO yaa
Mitsulkurindieeec aati ss aes en mae 250
Modiokdint soe ese nee shen nee i, 186
subellipticavs -e.- 5. face see 136
Molltis caja ieee an, beer LariaS
Mollsiscoidéase.t = 5.) Gy.
co AWS ag oe 1 SARE ee 7
General Index —Vol. VI. 513
Monopteria alata. ...............-. 127 | Orthis:
SepPRRaPI RS cs Sh cere vet SS Sue 129 ARR, 1. chit ta cae howetae” + OU
WRI IFIMIN sis nnn 5c ols oe ee 127 RRMMIOTIGL 5 eet ik se ce cee aie 62
REMBMMCU RIE as CLG. OSes. c.0 i ee 128 RMTOLS UD 550s Onis am «05 63, 64, 6b
PEE 2A aes Chews hee 131 BIT tOCOSEAEA s .40.5 0% ecu eres onc 65
MIDE WHI si. se Poa COLE MISHIR CrAGSA Vi... 6k ae ee 62
PRM ERUIS Ooi ka et i eee WRESOUMGENAIS 5 6 i... Os Les 65
RRR 6S dS UR ts eMC ee a 383 BUIARGIANE, Swsiae sk ocak ass 65
REEL soe hard 2 sen rai howe Ble Petar t OSbEOPIOSSIOW 8). sa bee es 340
EIT el a ink. 1 oe OE Hee) POISTMOPOIOER 4 ...c 6 See hoe oe 312, 393
UTEP ORORUN oot. Lk aS ova Nee SPA EL yn cc cere da wid ow hee we 98
BMMRER MG Ot sak ona ec oe 141 Otodus appendiculatus........... 247
MIANOHISIA Soh Ae wales GON OR 140 GIVATIGHTOR 42h. Ss ee ao 3 ie 248
PTLOTIOERGAL) cc se Osc hae 141 SUCRIIS Oc Site. 5 vee NS ee ents 248
subquadrata.............. i358, 400 | Oxyrhina mantelli................ 246
UPA Whe sp V.si2 cate She ase 137 P
MM MOMRIGISET SOc ops cak ois we cid dns 237 | nit
PRIMERS BRS Caw sca Wn cee 8 wae S ©) Pachyrhizodontide ..........:... 349
N tPachiva miguel se. Pek. 8 uss 2G 349, 393
MITT RST oe Sc acted es td 6 eee RG, ne ee aoe
ae TOLU es ce eee Lares bee ODE
_ VUES SS eee 8, 149 : ae
aR Ke IRTIMIOMEUEIy Sk dks eae s es OO
OTP LES ae at i ihe! 149 ied
tee otal lepiepnathusiec sii... ches ioe BOL
RMU AMMEREAO EE Sood Sis tS = a) sce wa Ss 165 : Se
: | 7G Ces 01S (eC ee as ea 354
CODES ADC a San 148 ie i
' ae x AYEEEMANIS ee 2s See ee i Bs 361
le ihe iiss hh 165 |
nella 151 WGleaRenvee Cee See lk ae oe eeoO
ne oe oc. 5s a) PACH YGOPMUG)... oUt. sk ub. PeOO
RSENS Ft eerie So Se wea os 150 eo
re PECL re Oe okie a. aes anes 67
ee ee ae 8, 148 ;
See AVUCUIBGUSE. few oe su fb iae eet Oe 113
CELESTE | GS SE OE aa 148 : 2
Sie PROMOS ACI com nen See ee oS 117
bellistriata attenuata......... 149 ; x
. CATOOMITGPUS on. vey tue Me wa eo 117
O. | eleavelandicus ............... 114
EE eS 249 SRV AT Mr uey chars Ais rua the See ote tie by
BEMHODGTUA, C5245... 6 cee) 6, 49, 50 | PLOVICHECHIS, . ee Se 119
WMS. Sari. Liao Ga ee 49 APES (OPES) 1181-1 Sen an ea 114
BRPWCUIOIGOR 234 e20 a ceks 6, 55, 56. | GCOMCM TANS nts su. tees oe 114
RNS Ws oc NG a ose the Ese oe LECCLOENUISS ce Osa ook Obs ed pe ok 7
manhattanensis.............. BAU PRCICCOULCLUS ys oko ie desde ticle de 362
missouriensis............. Die Men) seOlecCaDplerue oo 5. cl we es oe 362
Organisms, microscopic. ......... BT). PPIBGY POON. : <i asia cea ess iy by. 107
EAM ey Sic we 0 ofaldie's o's sae hs Gut) FONE DOZOM oF ii sklss ota vo sss Pea 6
ES re eee Gap. OF | SSrininn COGAN). Saajc ise. cea 4
BERMCUMOIMICH 60.64.20 000 0002s 62 | Perischoéchinoida................ 6
CRRIMMIMED Se iiss a8 een os es os Gin} PHBSPRNOGMSH. Fay icie ie ne oe oe 374
Sy a" a a GZ, ao IDS ee iwaeios ost Vea 376
OOM MMeb tar sec os ce Oe 8s oy | Geet mloetminus.. foo chee cet 6, 36
por To ene ane 62 SARIS wie scaeisass» 0
pc ic a 63 ee ogee 39
514
Pharetrones se. +407 os eee dD,
Pina 30s Ok, 1 eee 7%. 144
ACAMST Gs. atid cs ee ee 144
peracutars.j; 20. cn’ eee 144, 145
subspatulata.: : . sce aes 145
Binmidse eee ior... eee ee ee
Placunopsis {6 3... 3 eee evel
CabOOMATal i.) 515 see ILL, Wee |
Plethodidee .oi42 5.) ot eee 393
Pleurophorussc. 045 2 eee 8, 161
COStatiormish. 4. sone eee 162 |
COSPTALUS AS ecie. 5 dike ae a eee 162
SUbCOStatUS!.. 4: Sn. yeeies eee 161
froOpidophonuse chess. one ctor OZ
Pleurotomaria missouriensis...... 3
Plicatula striatocostata........... 65
PVC OSES shea Re arcuate eee 67
Porifetase.) aigcwhe ie a eee 5
Portlieuisys 2) he pea tie hence, ra 265, 393
ALCUAPUS Me hie ee ee ae eee 306
IO) SAN LAs Nha’ 8) ae gUnEH ROR SC 2D 293
NMOLOSHUS ye wee Ne! Tec oe eae 267
‘Gh akulinais ete cee yone seis enced Bhs. 267 |
Rosidonomy anviny rece. pen ae (ea ssi) =|
PeCTECMUISN Me vere er MN 136
FOCULVA tee oe Sete ee 135
Poleriocrinigdse wera). eee 6
IRotenocrnimnuUs..6. 046 he oe ae: 29, 34
POR AIC Cree RNC | ea AN Oe Wie Be ee a Vil
Prentice, Sydney waa) te 4, 259
Productidee tenuate venice 6
Produchus).).!.c sn ae Oy, Oy. eee
PO GuUlCOsbabusiyis sa entat ee 77
AIMERICATIUIGE Leone ee ere ar ee 77
ASSL US ciate ee ae ie ener 84
CalhouniamUs eee eae oe 75
CAMCrMM essen uote eee 83
COBAR ers tile Cees eo oe Sa ee Cos UE
COraaMmeniCanUush a pate OTe Ta
COSUEAMISE. Ao enki cre ron eater 719
NALS) cau awh oF STAMINA At cary, ere a Woy. oll
NOPTESECHS 2 ete cre pashan TM gn 84
NOPTIC USA 6. osc ee tee ae 85
AIDC Bais 4 thea ok ee ee 78
lOnGiISpINUG: eee hes see 81
Lilie ee utoaiels cyt ere ee AD 75
DAS TVS ee were naan ae ous
LOOPED, eo ec are naan Sore: 78
13 |
University Geological Survey of Kansas.
Productus:
Nebrascensis W, .").s.. . aoe 84
pertenuigia): s/s ae 83
portlockianus)..0 5 eee 79
prattemianusi i (eee fio eT
punctatuss .6 00.0 eee 87
TOGCLS «oo een) ee 84
semipunctatus .....) 20... een
semireticulatus..... 53, Ol, ona
setiperis., .3c)talieee eee 78
splendens..... 00.0 3. 09 eee 81
Symmetricus =. 24... 45 eee 89
tubulosprmusea:... aoe Beno)
viminalis..4 3. 0.5m ee ee 79
Wabashensis 4. 44.02 81
wilberanus)..... 024.0 306i 84
Prosser, Charles S........ BAER sso 4
| Protospondylin 32505) ae 393
Protozoa... 00. Ge en ee ee D
Protsphyrenay ays eee 362, 393
bentoniana,. (2. ae eee 365
bentoniay: issues) Oe 360
PIPAS 2 Mok oy ne a eee 367
penetranes. ... eae ee 369
FECULVITOSLTIS 2) 4.55 eee 366
Protosphyreenidez..... > ud 362
Protremaita (25.5% ks 60 6 ee 6
iP Seudomonotishn as. aes Ue By
Va WAR NSCs ey ie cae ene T3a2 pele
haw equistfiata sss. -2 ene 134
kansasensisieas we eee 133, 134
FaG@ialis : fsck Se 132
robustac’ 220.264 os eae eee 133
tenuistriata(j... o0)..0 ba ee 133
Pseudothry ptodus:.....: 4 eee 393
Plena iis. cake see ee 7, 125
1OWG A she. clon Wok ee 125
sulecatai.. 1. “ake meee 126
Pterinia)gibbosasx,. . ae ace noreee 129
Ptychod ts’... Omar mera 237, 364
AMODY.MUS. oe eh, ee ee 241
foreign speciesia... 3.655 ee 244
JANOWAY 1.2 jaoace eek eee 242
IAT GUM ese Us Roe eee 240
MOPTOMI a eS. eee 238
occidentalis irc see eee 242 |
DOLYSYLUS ja.) .. a oe eee 240
wihipplet.. 220) ae yo amen 243
General Index —Vol. VI. 515
Publications of the Survey........ MY |) SCANANOTHYNCHDUS:... 0.0... e608 eae 250
a a Say ee 7192 IOUT. Sci, «eons 0 sive rctes 251
BE COUILOL INI: 6.6166 S626 in " SSNS Cc ri a 8, 155
PAM VINO ALIGS foes. «sedis vedas 92 BRORLOTODSS stews 9 waters uv bo 158
AR rei gicthc = 4.5 Qatar 93 compressus 157
PVOHOMONGOR.. 6.5... 6.5 ances ae 254 | PRA Bea ee cea Petre is see tn Sse BON Te 155
ci Delig )aF ee ee 158
R. RSID CU a Merja) a: vse stance a uw 155
OS Se ae es 78 | SINC INC UA TIS Ma ns: cm acid uleod oa, 157
Retzia compressa.............. 103 WVITE MUI cy Camvoane's s 155, 257, 158
MeMeReET Tc nc 6d cas iw OS ie scnuchert, CHATIGS. .. 4.5 246.0000 4
WMAII ES UME COD POS 5 aie ci cite. 5 oy ince aod Want OR SGU INGES: 25 Gi owas ton peau aed 244
LE UNS ae iS ae GETS ics alllargtag ch bh: Pea ne 244
RINE ESBUGISOSID ova wh a Sb nod duel Sees 103 (hamng) STaciiss .. 24 cared 245
BMPUMIMAP IE tock ok ce hn weet 7, 102 | FARERUSS USS aN caale ea. oy sch 8 Seta ys 244
PRIMER ea oh ait ace a hicware 102 | PADS y erm risks Was ack v win aos acetals 245
PUPMOPNACAUB. 0.5 ek ee ea SAE) SES Loh 4111s ag a ea 244
MBMEMIMOOUA. .. wo. sa ce wwe yee | See UPOMING sei kde sc ce hawk a aes 46
ELS? Seren ee sem ON Mie (218 beh @ bel StF: ge lea an ee ive devil
SME re rb ss ee cc ee Oe oe O| TOPBRBOMBISK ec gies ealen Geer a
Panehonelia angulata........... 91 | Selachians..........:..........0: 237
SOT Un (0 rd ee SAN rss 2iie ate): 8S One Aer a ee Tas
2h SS Se re ere 93 EUR PPANG ELI MRe de hi cid Use. 5:9 «Ams pase) 105
PER VIHOMPANG, ... . ou. Oy ses oe 92 AIO UL aie eos eae vias oe emcee 105
on OE a eis G3 memirsnculatl ...2. cx... 0 sus news 74
SING? Meee ce Lo Les en css ake | SUPHODIGK. . 2 vinsalil. cscs eevee’ 8
(2 LEY E RNY. N) | RS ea fi OUI aG ace «ey Rs. 6. oss aevemlacens 8
RU ANETISENGD S94 ti /ry aks Sa we Sau 30's Se DSL TROETLY Us 6 iow ac nte wa Regie genie 8, 158
MBN NG ON bd tts 2 eoiine VO ives 67 | BVCRIEUMN ES Aeteyecc sey tics, sAWo = ila, uae 158
Ls d ES cee Gs ee 239 | CUE A a ee 160
|g?) 0/200) (6 (2) = ee ee te 159
S. fesie ltroedin Cates Nan ae he 8
CUETO Ce a si 2 ae 349, 393 | SOMMPNOSPONG IG... oss. os varasrde es 5, 12
MAUIFOCEDHSIUS: .. ...06 dha vos. 323, 393 | een PR RAG Of y 000 (: Bega eee ee aoe ere 12
CL sda Ole ee Ses, ee a a ieee DINO SE scent Ur Gi ose nak rela cigs 74
SFORCHOANT 200 fu vis ts elas citi oe she ea W MALO Lote ats leas idiots vce 7, 67, 98
MOHIMEUS. «24 es obs haa: 323, 392 | Game natlige eae ca 5 Adan 99
TRAUMAS 3s il y's Seas re die Ae 267 cameratus var. kansasensis... 99
SBAREMCORNIREMAO 54 2s viel. vy cee ss 262 ILS Oh etalk UNotai cies ures» 104
UMRRMAINED 2 ees. s!s aes oP OLY sOuo 114512) RE A 99
ee CTs | oa cian Ayr 313 PEC QRICOM ELIS ss. 015/45 sain yes «2 99
MIR a et Cs bose sy bs eee 319 MENUUCEVENBIS. | cris sans ee ay 96
AMMEN 002570 6 2 sid u.a!'xn vies) Oe kentuckyensisvar. propatulus. 96
PIER IMOTAMNS i 2 ss Sc en rele 312 PRI IMOSUB yee ed sls Vive wey sd we 96
Saurodontids............... 262, 310 ert ah ey Oe Oe 102
PP MIGPE MIO crs os nnn Hates 6, 21 meusebachanus.............. 99
BICTRAB AGRIC 5). ince a)s oat o's 34 ajeprey sities: |. Mee isa rae aes +
MMBIUNTNN OD oe ees eae bus 27 aan ios 41S 6 a 102
516
Spirifer:
planoconvexa. 4-6. see ee 101
BUTIAtUS ace. ee 100
striatus var. triplicatus....... 99
tripheatuss;: ars. | eee nee 99
Spirilenidees 4.4. 2st.) eee 77
S pipifiorimany 6.22) sao see eee 1,7 96
Cristabarciceiws | cake eee: 96
kentuchkyensis; 225.0 nee ee 96
Spinigera amenicamuss....se aoe ee 104
Caput-Serpentis:. 72). .4-. 106, 107
pechimIferaras)2 leek eee eens 104
planosulcatae. sso eee 104
Sumo billiitay Wie See oe eee 105
SPONGES Te NU iews Vinal Sean ener ila
Sponeieere Pee a ee ee Dy UL
Sporetodusyanewayl... secon eee 242
Springerw Mranikes, ees, eee 243
Staton Weel ee ee eee 237
Sitellisponmera wie cer eee 133
Stemmatocrimushs se) on teae 39 -
Stewart, Alban, article by........ 257 |
Sibratodvis\ yey nee rhein
PDICANTS a Eye sre a ar 328 |
Sina tocombiceewy ress een ee ee 326
Streptorhynchus crenistria........ 63 |
|ESCEX) LU Hc GSP cE yh stig men aS yt 63
ELCMMIOME hese ee es 62
FODUSTAMS hte ete oneaene! 63
Stmatocostataiee seen eee 65
SiGEbartee sat sok Se ee ce ne ne ee pee 67
Strophomenas.1. =) 2. = ae eee 67
Strophomenidceys. se ne 6
DY.COMEAS Aue Meise Laka iae Ungaate Mace 5 |
rei lies) G00 TSH iy OR amee Ine yar meu a hh ate 383
latiirOnsi: Ak. aise ae hee ere 384 |
Synteomodus, i) yey ew ee eee 393
Syntrialasma hemiplicata......... 91
SyFuCOpOha n=. ee Dae d 2
TANlitatheMiatasy ees ee: 25
SV LIN COPORIG sels mee een 5
RE
Tablevoticombemtsmae ace eee vi
Teleosts of the Upper Cretaceous.. 256
Peleosteris eee, eee ee 259
University Geological Survey of Kansas.
Teleost fishes, genera of .......... 386
Telotrematar. 4 82 GRe eee 7
Terebratula antisiensis............ 105
argented is) 2a Bape eee 105
bovidens:..22).)0. 1.2). sae ee
GOV TOY SSL N52 Wile ae ee ee 104
elongata 2. xan i Oe 95
genicula: fee ae ene 95
haStatai 2:55 730 es eee 95
MOTMONA 1 i3h ae eee 103
peruviana BR ee Ge: 105
LOCKY MON tana eee 92
LOVSS1 4 2 hs oo 105
subtilita: 0... eee 105
Utah f0e ek ee 93
Terebratulide........ ee ete es U
Terebratulites cristatus........... 96
Thry ptddus.s) 29 eee Bw oes
Prachyporay, Seek ee eee Dp) 224
BUSI 4h hs eee 22
OLMa tay hohe on ee 22
Trimnaspis oi. ck ee ee ee 382
Trigontidee,... oe... Ae eee 8
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Williston, S. W., paper by........ 380
Woodward, A. Su2..4 See eee 259
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Xiphactimus ]. 4558. 46. 265
BUGAK ce ee eee 267
lbrachyoma:t sea. eee eee 293
LOWaAIS S/o ee 293
DG oy alts tslee ae ee et ee VM oS 362
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Yoldiias 4.25.05 5S se eee 8, 152
Glabrae. yk. oe eee eel eee 153
KnOX@NSIS'.2.0 5a or: ore 154
subseitulas... iar eer eee 152
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Aaphrentidee ws aes} eee eee 5
ZOACTUBUS Mase itr noe ones 6, 29, 49
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